CANTHOCAMPTIDAE Sars, 1906, emend. Monard, 1927, Lang, 1948


BRYOCAMPTUS Chappuis, 1928


Kiefer (1978) and Dussart & Defaye (1990), splitted the genus Bryocamptus s. l. Chappuis, 1928 in the subgenera Bryocamptus s.str. Chappuis, 1928, Rheocamptus , Borutzky, 1948, Limocamptus Chappuis, 1928 and Arcticocamptus Chappuis, 1928. Wells (2007) do not report the subgenus Rheocamptus, including the related species in the subgenus Bryocamptus s. str. More recently Lee & Lee (2010) revalidated the subgenus Echinocamptus.


BRYOCAMPTUS Chappuis, 1928 (subgenus BRYOCAMPTUS Chappuis, 1928)


  • Bryocamptus (Bryocamptus) aberrans Apostolov & Pesce, 1991 [Bulgaria; cave waters]
  • Bryocamptus (Bryocamptus) abramovae Novikov et a., 2023 [Russia]
  • Bryocamptus (Bryocamptus) abyssicola Borutzky & Okuneva, 1972 [Baikal lake]
  • Bryocamptus (Bryocamptus) baikalensis Borutzky, 1930
  • Bryocamptus (Bryocamptus) bogoriensis Kiefer, 1933 [India; fresh waters?]
  • Bryocamptus (Bryocamptus) borutzkyi Petkovski, 1969 (Asia, Afghanistan; fresh waters?]
  • Bryocamptus (Bryocamptus) bulbochaetus Borutzky, 1931 [Baikal lake]
  • Bryocamptus (Bryocamptus) campaneri Reid, 1994
  • Bryocamptus (Bryocamptus) chappuisi Borutzky, 1948 [Baikal lake]
  • Bryocamptus (Bryocamptus) cokeri Borutzky, 1948 [Baikal lake]
  • Bryocamptus (Bryocamptus) elaphoides Borutzky & Okuneva, 1972 [Baikal lake]
  • Bryocamptus (Bryocamptus) hiatus hiatus (Willey, 1925) [north America; moss, marsh]
  • Bryocamptus (Bryocamptus) hiatus australis (Coker, 1934) [north America; fresh waters]
  • Bryocamptus (Bryocamptus) hutchinsoni Kiefer, 1929 [USA; fresh waters]
  • Bryocamptus (Bryocamptus) incertus Borutzky, 1931 [Baikal lake]
  • Bryocamptus (Bryocamptus) intercalaris Shen & Tai, 1973 [China; cave waters]
  • Bryocamptus (Bryocamptus) jejuensis Lee & Chang, 2016 [Korea]
  • Bryocamptus (Bryocamptus) kozhowi Borutzky & Okuneva, 1971 [Baikal lake]
  • Bryocamptus (Bryocamptus) longicaudatus Borutzky & Okuneva, 1972 [Baikal lake]
  • Bryocamptus (Bryocamptus) longifurcatus Borutzky, 1931 [Baikal lake]
  • Bryocamptus (Bryocamptus) longisetosus Borutzky, 1931 [Baikal lake]
  • Bryocamptus (Bryocamptus) minutus minutus (Claus, 1863) [cosmopolitan; fresh and brackish waters]
  • Bryocamptus (Bryocamptus) minutus var. bispinosa Lang, 1957 [Asia; fresh waters]
  • Bryocamptus (Bryocamptus) minutus schizodon (Mrazek, 1893)
  • Bryocamptus (Bryocamptus) minutus simplicidentata (Willey, 1934)
  • Bryocamptus (Bryocamptus) minutus vejodovskyformis Kiefer, 1934
  • Bryocamptus (Bryocamptus) mrazeki (Minkiewicz, 1916) [cenral Europe; fresh and brackish waters]
  • Bryocamptus (Bryocamptus) newyorkensis (Chappuis, 1927) [USA; fresh waters]
  • Bryocamptus (Bryocamptus) pilosus Flossner, 1989
  • Bryocamptus (Bryocamptus) putoranus Novikov et a., 2023 [Russia]
  • Bryocamptus (Rheocamptus) pygmaeus bulbochaetus Sterba, 1962 [Europe, Asia; fresh waters]
  • Bryocamptus (Bryocamptus) sinuatus Borutzky & Okuneva, 1972 [Baikal lake]
  • Bryocamptus (Bryocamptus) subarcticus (Wi8lley, 1925)
  • Bryocamptus (Bryocamptus) tarnogradskyi Borutzky, 1934 [Asia, Europe; caves, fresh waters]
  • Bryocamptus (Bryocamptus) tenuis Borutzky & Okuneva, 1971 [Baikal lake]
  • Bryocamptus (Bryocamptus) tuberculatus Borutzky, 1948 [Baikal lake]
  • Bryocamptus (Bryocamptus) umiatensis Wilson, 1958 [Alaska; fresh waters]
  • Bryocamptus (Bryocamptus) vejdovskyi vejdovskyi (Mrazek, 1893) [Europe, Asia, Japan ; fresh and brackish waters]
  • Bryocamptus (Bryocamptus) vejdovskyi minutiformis Kiefer, 1934 [north America; fresh waters}
  • Bryocamptus (Bryocamptus) yohteiensis Ishida, 1997
  • Bryocamptus (Bryocamptus) washingtonensis Wilson, 1958 [north America; fresh waters]


    BRYOCAMPTUS Chappuis, 1928 (subgenus RHEOCAMPTUS)


  • Bryocamptus (Rheocamptus) albidus Okuneva, 1983 [Baikal lake]
  • Bryocamptus (Rheocamptus) alosensis Apostolov, 1998 [France; hyporheic waters]
  • Bryocamptus (Rheocamptus) aquaeductus aquaeductus Borutzky, 1934 [Europe; aqueducts]
  • Bryocamptus (Rheocamptus) aquaeductus var. petkovski Apostolov, 1971 [Bulgaria; springs]
  • Bryocamptus (Rheocamptus) balcanicus balcanicus (Kiefer, 1933) [Europe; cave, phreatic waters]
  • Bryocamptus (Rheocamptus) balcanicus f. babunae Petkovski, 1956 [Yugoslavia; ground waters] **
  • Bryocamptus (Rheocamptus) birsteini Borutzky, 1940 [URSS, cave waters]
  • Bryocamptus (Rheocamptus) bispinosus Borutzky, 1940 [URSS; cave waters]
  • Bryocamptus (Rheocamptus) borus Karanovic & Bobic, 1998 [Serbia; cave waters]
  • Bryocamptus (Rheocamptus) brevipes Borutzky, 1948 [Baikal lake]
  • Bryocamptus (Rheocamptus) crassipes Borutzky & Okuneva, 1972 [Baikal lake]
  • Bryocamptus (Rheocamptus) cristatus Borutzky & Okuneva, 1972 [Baikal lake]
  • Bryocamptus (Rheocamptus) dentatus Chappuis, 1937 [Italy, Spain, France; cave waters, wells]
  • Bryocamptus (Rheocamptus) denticulatus Borutzky & Okuneva, 1972 [Baikal lake]
  • Bryocamptus (Rheocamptus) gauthieri (Roy, 1924) [north America; moss]
  • Bryocamptus (Rheocamptus) innominatus Borutzky, 1940 [URSS, Azerbaidjan; cave waters]
  • Bryocamptus (Rheocamptus) littoralis Borutzky & Okuneva, 1972 [Baikal lake]
  • Bryocamptus (Rheocamptus) madarensis Apostolov, 1969 [Bulgaria; fresh waters]
  • Bryocamptus (Rheocamptus) marinus Gurney, 1927 [Suez; brackish waters]
  • Bryocamptus (Rheocamptus) mirus Petkovski & Karanovic, 1997 [Balkan Peninsula; Ohrid lake]
  • Bryocamptus (Rheocamptus) pirgos Apostolov, 1969 [Bulgaria; fresh waters]
  • Bryocamptus (Rheocamptus) pygmaeus pygmaeus (Sars, 1863) [cosmopolitan, except Australia?; fresh waters, moss]
  • Bryocamptus (Rheocamptus) pyrenaicus (Chappuis, 1923) [France, Spain; cave waters]
  • Bryocamptus (Rheocamptus) reductus Borutzky, 1948 [URSS, Azerbaidjan; cave waters]
  • Bryocamptus (Rheocamptus) rylovi Borutzky, 1931 [Baikal lake]
  • Bryocamptus (Rheocamptus) saxicola Borutzky & Okuneva, 1972 [Baikal lake]
  • Bryocamptus (Rheocamptus) spinulosus spinulosus Borutzky, 1934 [Asia; fresh waters]
  • Bryocamptus (Rheocamptus) spinulosus f. occidentalis Sterba, 1961 [Europe; fresh waters]
  • Bryocamptus (Rheocamptus) spinulosus f. triarticulata Sterba, 1968 [Asia; fresh waters?]
  • Bryocamptus (Rheocamptus) stillae Cottarelli et al., 2011 [Italy; caves]
  • Bryocamptus (Rheocamptus) tauricus (Borutzky, 1930) [URSS; cave waters]
  • Bryocamptus (Rheocamptus) tatrensis (Minkiewicz, 1916) [Asia; fresh waters]
  • Bryocamptus (Rheocamptus) typhlops (Mrazek, 1893) [Europe; fresh waters,moss]
  • Bryocamptus (Rheocamptus) unisaetosus Kiefer, 1930 * [Europe; cave waters]
  • Bryocamptus (Rheocamptus) zschokkei zschokkei (Schmeil, 1893) [cosmpolitan, except Australia?; fresh waters]
  • Bryocamptus (Rheocamptus) zschokkei alleganiensis Coker, 1934 [USA; fresh waters]
  • Bryocamptus (Rheocamptus) zschokkei var. apilosus Sterba, 1955 [Europe; moss]
  • Bryocamptus (Rheocamptus) zschokkei caucasicus Kiefer, 1978
    [Asia; fresh waters] *
  • Bryocamptus (Rheocamptus) zschokkei f. kalinae Petkovski, 1956 [Yugoslavie;cave waters]
  • Bryocamptus (Rheocamptus) zschokkei komi Borutzky, 1962 [URSS; rivers]
  • Bryocamptus (Rheocamptus) zschokkei sinkiangensis Shen & Sung, 1965 [China; fresh waters?]
  • Bryocamptus (Rheocamptus) zschokkei triarticulata Kiefer, 1929 [France, URSS; ground waters] **
  • Bryocamptus (Rheocamptus) zschokkei yunnannsis Shen & Tai, 1964 [China; fresh waters]
  • Bryocamptus (Rheocamptus) weberi (Kessler, 1914) [Europe; fresh waters, wet moss]

    * syn. of B. (R.) tatrensis (Minkiewicz, 1916); according to Petkovski (1972) could be syn. of B. (R.) typhlops Mrazek, 1893)

    ** questionable taxa


    NO SUBGENERIC STATUS DEFINED

  • Bryocamptus lacustris Wells 2007
  • Bryocamptus pacificus Ishida, 1992 [Japan; streams]
  • Bryocamptus pilosus Flossner, 1989 [north America; lakes]
  • Bryocamptus stouti Herding, 1958 [New Zealand, fresh waters]
  • Bryocamptus yohteiensis Ishida, 1997 [Japan; stream puddles]




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