Arachniodes

Species

Arachniodes species include but are not limited to those in the following list. A species name preceded by (=) is considered by the referenced authority to be a synonym of the accepted species name listed above it.

1828 Blume

Carl Ludwig von Blume published a description of the first Arachniodes species in Enumeratio Plantarum Javae et Insularum Adjacentium (An Enumeration of the Plants of Java and Adjacent Islands) in 1828.

Arachniodes aspidioides Blume 1828

1961 Tindale

The next Arachniodes species descriptions were published by Mary Douglas Tindale in Contributions from the New South Wales National Herbarium, volume 3, number 1, in 1961. This transferred two species from Conrad Vernon Morton's one-year-old Byrsopteris genus to Blume's 133-year-old Arachniodes genus.

Arachniodes amabilis (Blume) Tindale 1961 (India, Japan, Nepal, Philippines, Sri Lanka, Guangdong, Sichuan, Taiwan, Yunnan)

Arachniodes aristata (G.Forst.) Tindale 1961 – East Indian holly fern
(India, Japan, South Korea, Sri Lanka)

1962 Ohwi

Jisaburo Ohwi published the following Arachniodes species in the Journal of Japanese Botany, volume 37, number 3, in 1962.

Arachniodes assamica (Kuhn) Ohwi 1962 (Tibet, Yunnan, India, Nepal)

Arachniodes cavalerii (H.Christ) Ohwi 1962

Arachniodes maximowiczii (Baker) Ohwi 1962

Arachniodes miqueliana (Maxim. ex Franch. & Sav.) Ohwi 1962

Arachniodes mutica (Franch. & Sav.) Ohwi 1962

Arachniodes nipponica (Rosenst.) Ohwi 1962

Arachniodes pseudoaristata (Tagawa) Ohwi 1962

Arachniodes simplicior (Makino) Ohwi 1962 – simpler holly fern

Arachniodes sinomiqueliana (Ching) Ohwi 1962

Arachniodes standishii (T.Moore) Ohwi 1962 – upside-down fern

1962 Ching

The following species were transferred into Arachniodes by Ren Chang Ching in an article entitled "On the Nomenclature of the Compound-leaved Polysticha", published in the journal Acta Botanica Sinica, volume 10, number 3, Peking, China, September, 1962. Besides those listed below, a number of his species had already been published by Ohwi a few months earlier, and so appear in the section above, listed as synonyms marked nom. illeg.

Arachniodes amoena (Ching) Ching 1962 (Fujian, Guangdong, Guangxi, Guizhou, Hunan, Jiangxi, Zhejiang)

Arachniodes arisanica (Rosenst.) Ching 1962

= Arachniodes globisora (Hayata) Ching 1964

Arachniodes carvifolia (Kunze) Ching 1962 – East Indian holly fern (Yunnan, India, Philippines, Sri Lanka, U.S.)

Arachniodes chinensis (Rosenst.) Ching 1962

Arachniodes coniifolia (T. Moore) Ching 1962 (Sichuan, Yunnan, Nepal)

Arachniodes denticulata (Sw.) Ching 1962

Arachniodes diffracta (Baker) Ching 1962

= Acrorumohra diffracta (Baker) H. Itô 1938 (Guangxi, Hainan, Yunnan, Japan, Vietnam)

Arachniodes dimorphophyllum (Hayata) Ching 1962

Arachniodes exilis (Hance) Ching 1962

Arachniodes festina (Hance) Ching 1962 (China)

Arachniodes grossa (Tardieu & C.Chr.) Ching 1962 (Guangdong, Hainan)

Arachniodes hasseltii (Blume) Ching 1962

= Acrorumohra hasseltii (Blume) Ching 1964 (Hainan, Taiwan, India, Nepal)

Arachniodes hainanensis (Ching) Ching 1962 (Hainan)

Arachniodes henryi (H.Christ) Ching 1962

Arachniodes leucostegioides (C.Chr.) Ching 1962

Arachniodes nigrospinosa (Ching) Ching 1962

Arachniodes obtusissima (Ching) Ching 1962

Arachniodes simulans (Mett. ex Kuhn) Ching 1962

Arachniodes speciosa (D.Don) Ching 1962

Arachniodes spectabilis (Ching) Ching 1962

Arachniodes subreflexipinna (Ogata) Ching 1962

= Acrorumohra subreflexipinna (Ogata) H. Itô in Nakai & Honda 1938

Arachniodes tonkinensis (Ching) Ching 1962

Arachniodes zeylanica (Ching) Ching 1962

1962 Sa.Kurata

In 1962, Satoru Kurata published descriptions of the following specie in the Science Report of the Yokosuka City Museum, Yokosuka, Japan.

Arachniodes × kenzo-satakei (Sa.Kurata) Sa.Kurata 1962

Arachniodes subamabilis Sa.Kurata 1962

1964 Ching & Chu H.Wang

Ren Chang Ching and Chu Hao Wang published the following species in Acta Phytotaxonomica Sinica, volume 9, number 4, 1964.

Arachniodes acuminata Ching & Chu H.Wang 1964

= Arachniodes sphaerosora (Tagawa) Ching 1965

Arachniodes caudata (Tagawa) Ching 1964

Arachniodes globisora (Hayata) Ching 1964

Arachniodes obtusiloba Ching & Chu H.Wang 1964

Arachniodes pseudocavalerii Ching & Chu H.Wang 1964

Arachniodes rhomboidea (Schott) Ching 1964

Arachniodes yoshinagae (Makino) Ching 1964

1965 Ching

Ren Chang Ching in Acta Phytotaxonomica Sinica, volume 10, number 3, 1965, published:

Arachniodes sphaerosora (Tagawa) Ching 1965

1965 Sa.Kurata

Satoru Kurata published the following Arachniodes species name in the Journal of Geobotany (Hokuriku Journal of Botany), volume 13, in 1965.

Arachniodes hekiana Sa.Kurata 1965

1965 Proctor

Published by George Richardson Proctor in the British Fern Gazette. volume 9, number 6, 1965

Arachniodes denticulata (Sw.) Proctor 1965 (nom. illeg.)

= Arachniodes denticulata (Sw.) Ching 1962

Arachniodes rigidissima (Hook.) Proctor 1965

1967 Schelpe

Edmund André Charles Louis Eloi Schelpe published the following species in Boletim da Sociedade Broteriana, sér. 2, vol. 41, 1967.

Arachniodes webbiana Schelpe 1967

Arachniodes foliosa Schelpe 1967

1968 Sa.Kurata

In 1968, Satoru Kurata published the following Arachniodes species names in the Journal of Geobotany (Hokuriku Journal of Botany), volume 16.

Arachniodes repens Sa.Kurata 1968

Arachniodes yasu-inouei Sa.Kurata 1968

Arachniodes × masakii Sa.Kurata 1968

Arachniodes × clivorum Sa.Kurata 1968

Arachniodes × kurosawae Shimura & Sa.Kurata in Sa.Kurata 1968

Arachniodes × pseudohekiana Sa.Kurata 1968

Arachniodes × sasamotoi Sa.Kurata 1968

1969 & 1970 Nakaike

Toshiyuki Nakaike published the following new species in Journal of Geobotany (Hokuriku Journal of Botany), volume 17, 1969, and volume 18, 1970.

Arachniodes okinawensis Nakaike 1969 (Japan)

Arachniodes pseudorepens Nakaike 1970 (Japan)

1974 Ching

In Flora Tsinlingensis, volume 2, 1974, Ren Chang Ching published:

Arachniodes centrochinensis Ching 1974

= Arachniodes simulans (Ching) Ching 1962 (China)

1975 R.M.Tryon & D.S.Conant

Rolla Milton Tryon and Davis S. Conant published the following species in Acta Amazonica, volume 5, number 1, in 1975.

Arachniodes macrostegia (Hook.) R.M.Tryon & D.S.Conant 1775 (Brazil, Venezuela)

1975 Nakaike

In 1975, Toshiyuki Nakaike published the following species names in his Enumeratio Pteridophytarum Japonicarum, Filicales.

Arachniodes sporadosora (Kunze) Nakaike 1775

Arachniodes japonica (Kurata) Nakaike 1775

1978 Sa.Kurata

Satoru Kurata published the following Arachniodes species names in the Journal of Geobotany (Hokuriku Journal of Botany), volume 25, in 1978.

Arachniodes cantilenae Sa.Kurata 1978

Arachniodes hiugana Sa.Kurata 1978

Arachniodes kenzo-satakei Sa.Kurata 1978

Arachniodes tomitae Sa.Kurata 1978

Arachniodes × minamitanii Sa.Kurata 1978 (A. hiugana × A. nipponica)

Arachniodes × mirabilis Sa.Kurata 1978 (A. assamica × A. rhomboidea)

Arachniodes × mitsuyoshiana Sa.Kurata 1978 (A. miqueliana × A. simplicior)

Arachniodes × respiciens Sa.Kurata 1978 (A. exilis × A. simplicior)

1982 Proctor

Arachniodes argillicola (Proctor) Proctor 1982

Arachniodes chaerophylloides (Poir.) Proctor 1982 – false holly fern

1983 Y.T.Hsieh

Yin Tang Hsieh in Acta Botanica Yunnanica, volume 5, number 1, 1983.

Arachniodes jinpingensis Y.T.Hsieh 1983 (Yunnan)

Arachniodes valida Y.T.Hsieh 1983 (Yunnan)

Arachniodes dayaoensis Y.T.Hsieh 1983 (Yunnan)

1984 Y.T.Hsieh, Ching, et al.

Yin Tang Hsieh in Bulletin of Botanical Research, volume 4, number 2, 1984.

Arachniodes caudifolia Ching & Y.T.Hsieh 1984 (Yunnan)

Arachniodes gansuensis (Ching) Y.T.Hsieh 1984 (Yunnan, Sichuan)

Arachniodes guangnanensis Y.T.Hsieh 1984 (Yunnan)

Arachniodes lushuiensis Y.T.Hsieh 1984 (Yunnan)

Arachniodes reducta Y.T.Hsieh & Y.P.Wu 1984 (Zhejiang)

Arachniodes subaristata Ching & Y.T.Hsieh 1984 (China)

Yin Tang Hsieh and Ren Chang Ching, in Acta Phytotaxonomica Sinica, volume 22, number 2, 1984.

Arachniodes ishingensis Ching & Y.T.Hsieh 1984

Arachniodes obtusipinnula Ching & Y.T.Xie 1984

= Arachniodes tonkinensis (Ching) Ching 1962

Arachniodes ziyunshanensis Y.T.Hsieh 1984

Ren Chang Ching and Zheng-yu Liu, in Acta Phytotaxonomica Sinica, volume 22, number 4, 1984.

Arachniodes nanchuanensis Ching & Z.Y.Liu 1984 (Sichuan)

1986 Ching

Ren Chang Ching published the following species names in the Bulletin of Botanical Research, volume 6, number 3, in 1986.

Arachniodes abrupta Ching 1986 (Guangxi, Zhejiang)

Arachniodes ailaoshanensis Ching 1986 (Yunnan)

Arachniodes anshunensis Ching & Y.T. Hsieh 1986 (Guizhou)

Arachniodes aristatissima Ching 1986 (Zhejiang)

Arachniodes attenuata Ching 1986 (Yunnan)

Arachniodes austro-yunnanensis Ching 1986 (Yunnan)

Arachniodes baiseensis Ching 1986 (Guangxi)

Arachniodes calcarata Ching 1986 (Guizhou, Hunan, Sichuan)

Arachniodes cyrtomifolia Ching 1986 (Sichuan)

Arachniodes cornopteris Ching 1986 (Sichuan)

Arachniodes costulisora Ching 1986 (Sichuan)

Arachniodes decomposita Ching 1986 (Guizhou)

Arachniodes elevatas Ching 1986 (Sichuan)

Arachniodes emeiensis Ching 1986 (Sichuan)

Arachniodes erythrosora Ching 1986

= Arachniodes festina (Hance) Ching 1962 (Guangdong)

Arachniodes falcata Ching 1986 (Sichuan)

Arachniodes fengii Ching 1986

Arachniodes foeniculacea Ching 1986

Arachniodes fujianensis Ching 1986

Arachniodes gigantea Ching 1986

Arachniodes gijiangensis Ching 1986

Arachniodes gizushanensis Ching 1986

Arachniodes gongshanensis Ching & Y.T.Hsieh 1986

Arachniodes gradata Ching 1986

Arachniodes guangtongensis Ching 1986

Arachniodes guangxiensis Ching 1986

Arachniodes guanxianensis Ching 1986

Arachniodes hekouensis Ching 1986

Arachniodes heyuanensis Ching 1986

Arachniodes huapingensis Ching 1986

Arachniodes hunanensis Ching 1986

Arachniodes jiangxiensis Ching 1986

Arachniodes jinfoshanensis Ching 1986

Arachniodes jingdongensis Ching 1986

Arachniodes leuconeura Ching 1986

Arachniodes liyangensis Ching 1986

Arachniodes longipinna Ching 1986

Arachniodes lushanensis Ching 1986

Arachniodes maoshanensis Ching 1986

Arachniodes mengziensis Ching 1986

Arachniodes multifida Ching 1986

Arachniodes nanqingensis Ching 1986

Arachniodes neoaristata Ching 1986

Arachniodes nitidula Ching 1986

Arachniodes pianmaensis Ching 1986

Arachniodes pseudo-assamica Ching 1986

Arachniodes pseudo-longipinna Ching 1986

Arachniodes pseudo-simplicior Ching 1986

Arachniodes semifertilis Ching 1986

Arachniodes setifera Ching 1986

Arachniodes sichuanensis Ching 1986

Arachniodes similis Ching 1986

Arachniodes sino-aristata Ching 1986

Arachniodes sino-rhomboidea Ching 1986

Arachniodes shuangbaiensis Ching 1986

Arachniodes sparsa Ching 1986

Arachniodes spino-serrulata Ching 1986

Arachniodes subamoena Ching 1986

= Arachniodes amoena (Ching) Ching 1962

Arachniodes triangularis Ching 1986

Arachniodes xinpingensis Ching 1986

Arachniodes yaomashanensis Ching 1986

Arachniodes yinjiangensis Ching 1986

Arachniodes yunnanensis Ching 1986

1986 Seriz.

Arachniodes borealis Seriz. 1986 (Japan, Korea)

= Leptorumohra borealis ( Seriz. ) Shmakov 2009

1986 Y.T.Hsieh

Yin Tang Hsieh published the following species names in the Bulletin of Botanical Research, volume 6, number 4, in 1986.

Arachniodes chingii Y.T.Hsieh 1986

Arachniodes damiaoshanensis Y.T.Hsieh 1986

Arachniodes futeshanensis Y.T.Hsieh 1986

Arachniodes nibashanensis Y.T.Hsieh 1986

Arachniodes yunqiensis Y.T.Hsieh 1986

1991 Ching & Hsieh

Ren Chan Ching and Yin Tang Hsieh, in the Bulletin of Botanical Research volume 11, number 2, 1991, published:

Arachniodes basipinnata (Ching) Ching ex Y.T.Hsieh 1991

Arachniodes lanceolata Y.T.Hsieh 1991

Arachniodes menglianensis Y.T.Hsieh 1991

Arachniodes parasimplicior Ching ex Y.T.Hsieh 1991

= Arachniodes simplicior (Makino) Ohwi 1962

Arachniodes fengyangshanensis Ching & C.F.Zhang ex Y.T.Hsieh 1991

= Arachniodes exilis (Hance) Ching 1962

1992 Nakaike

Arachniodes davalliaeformis (Christ) Nakaike 1992 – shiny bristle fern

1993 W.H.Wagner

Arachniodes insularis W.H.Wagner 1993 – Santa Cruz Island holly fern

1995 S.F.Wu

The following two species were published by Shi Fu Wu in Keys to the Vascular Plants of the Wuling Mountains Area (edited by Wang Wentsai) 572–573, pl. 5. Beijing 1995.

Arachniodes hupingshanensis S.F.Wu 1995 (China)

Arachniodes wulingshanensis S.F.Wu 1995 (China)

1998 R.C.Moran & B.Øllg.

Arachniodes squamulosa R.C.Moran & B.Øllg. 1998

2008 Fraser-Jenk.

In 2008, Christopher Roy Fraser-Jenkins published the following new Arachniodes species in his Taxonomic revision of three hundred Indian subcontinental pteridophytes with a revised census-list: a new picture of fern-taxonomy and nomenclature in the Indian subcontinent.

Arachniodes cornucervi (D.Don) Fraser-Jenk. 2008

Arachniodes palmipes (Kunze) Fraser-Jenk. 2008

Arachniodes sledgei Fraser-Jenk. 2008

Leptorumohra

The genus Leptorumohra was segregated from the genus Rumohra in 1938 by Hiroshi Itô. In 1962, Jisaburo Ohwi included at least three of the Leptorumohra species in Arachniodes, but these placements were not universally accepted. In 2006, Alan Reid Smith & al. in "A classification for extant ferns" included the genus Leptorumohra in Arachniodes. Leptorumohra species, which may be considered part of the Arachniodes genus include:

Leptorumohra amurensis (H.Christ) Tzvelev 1989 (Siberia, Poland, Japan, Heilongjiang, Jilin, Liaoning, Mongolia)

Arachniodes borealis Seriz. 1986

(=) Leptorumohra borealis ( Seriz.) Shmakov 2009

Arachniodes miqueliana (Maxim. ex Franch. & Sav.) Ohwi 1962

(=) Polystichopsis miqueliana (Maxim. ex Franch. & Sav.) Tagawa 1958(=) Leptorumohra miqueliana (Maxim. ex Franch. & Sav.) H. Itô 1938(=) Rumohra miqueliana (Maxim. ex Franch. & Sav.) Ching 1934(=) Dryopteris miqueliana (Maxim. ex Franch. & Sav.) C.Chr 1905(=) Nephrodium miquelianum (Maxim. ex Franch. & Sav.) Kom. 1901(=) Aspidium miquelianum Maxim. ex Franch. & Sav. 1876

Arachniodes mutica (Franch. & Sav.) Ohwi 1962

(=) Polystichopsis mutica (Franch. & Sav.) Tagawa 1958(=) Leptorumohra mutica (Franch. & Sav.) Czerep. 1981(=) Rumohra mutica (Franch. & Sav.) Ching 1934(=) Dryopteris mutica (Franch. & Sav.) C.Chr. 1905(=) Aspidium muticum Franch. & Sav. 1876

Arachniodes quadripinnata (Hayata) Seriz. 1986

(=) Leptorumohra quadripinnata (Hayata) H.Itô 1938(=) Microlepia quadripinnata Hayata 1911

Arachniodes sinomiqueliana (Ching) Ohwi 1962

(=) Polystichopsis sinomiqueliana (Ching) Tagawa 1958(=) Leptorumohra sinomiqueliana (Ching) Tagawa 1939(=) Rumohra sinomiqueliana Ching 1934