Inventory of Ophrys (Orchidaceae) hybrids of Italy

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1 J. Eur. Orch. 49 (3-4): Romieg Soca Inventory of Ophrys (Orchidaceae) hybrids of Italy For Rodolfo Pichi Sermolli «Suppongo che la tendenza a formare ibridi si trovi in varie specie di questo genere, e per questo vorrei raccomandare a quelli che visitano luoghi ove crescono promiscuamente varie specie di Ophrys, di esaminarne con attenzione il maggior numero possibile di esemplari.» (SOMMIER 1896: 254). Keywords Orchidaceae, Ophrys, hybrids, flora of Italy. Summary Soca, R. (2017): Inventory of Ophrys (Orchidaceae) hybrids of Italy.- J. Eur. Orch. 49 (3-4): The author makes the census of the hybrids of the Ophrys genus described from Italy. It is the first work on this subject in Italy which can be part of a larger project to identify the endemic plants of Italy. The binomials of the parents are indicated such as announced by the descriptors as well as according to ROMOLINI & SOUCHE (2012). Indication of the herbarium(a) in which the types are deposited is also given. Some statistics are presented: the list of the described Ophrys hybrids from Italy by region, the list of the herbaria and the presence of the types, the number of hybrids described by Italian and by foreigners. It should be noted that very few synonymous descriptions exist: only seventeen, which are mostly the result of individuals who do not have a comprehensive documentation correlated to journals without proper scientific advices. Only fifty three can be regarded as non strict endemic in Italy. And twenty-nine are absent from The International Plant Names Index (IPNI) list. Journal Europäischer Orchideen 49 (3-4):

2 Zusammenfassung Soca, R. (2017): Inventar der Ophrys-Hybriden von Italien.- J. Eur. Orch. 49 (3-4): Der Autor listet alle aus Italien beschriebenen Hybriden der Gattung Ophrys auf. Dies ist der erste Beitrag zu diesem Thema in Italien, die Ergebnisse können in ein erweitertes Projekt zur Erfassung der endemischen Pflanzen Italiens einfließen. Aufgeführt werden für alle die Hybridformel mit den Namen der Eltern, dem Hybridnamen soweit vorhanden, den im Protolog angegebenen Namen der Eltern sowie diese nach aktueller Nomenklatur, gefolgt von der Angabe des den Typus aufbewahrenden Herbariums. In einer Übersichtsstatistik wird die Herkunft der Hybriden nach Regionen und Provinzen Italiens aufgeschlüsselt, die Herbarien der Typen zusammengestellt sowie die Anzahl der Hybriden nach Herkunft der Beschreiber Italien oder Ausland ausgewertet. Bemerkenswert ist, dass für die beschriebenen Hybriden nur sehr wenig Synonyme existieren, insgesamt nur siebzehn. Diese gehen auf Beschreibungen in Publikationsorganen mit ungenügender Begutachtung und auf unvollständige Dokumentation zurück. Von knapp über 280 bisher füer Italien bekannten Ophrys-Hybriden sind lediglich 53 nicht endemisch für Italien. In der Liste The International Plant Names Index (IPNI) sind 29 Namen von Ophrys-Hybriden aus Italien nicht aufgeführt. Riassunto Soca, R. (2017): Inventario degli ibridi di Ophrys dell Italia.- J. Eur. Orch. 49 (3-4): L'autore fa il censimento di tutti gli ibridi del genere Ophrys descritti per l'italia. È il primo lavoro su questo argomento in Italia che può inserirsi in un vasto progetto di censimento di piante endemiche dell'italia. Sono indicati i binomi dei genitori, come annunciati dai descrittori e secondo la tassonomia aggiornata. Con indicazione dell'erbario(i) ove sono depositati i tipi. Sono presentate alcune statistiche: l'elenco degli ibridi di Ophrys descritti per l'italia per regione, l'elenco degli erbari e presenza dei tipi, il numero di ibridi descritti da italiani e da stranieri. È interessante notare che esistono pochissime descrizioni sinonimi: solamente diciassette, che sono per lo più il risultato di individui che non hanno una documentazione completa correlato a riviste senza un'adeguata consulenza scientifica. Soltanto cinquantatre possono essere considerati come non endemici stricti dell'italia. Ventinove sono assenti della lista The International Plant Names Index (IPNI). Résumé Soca, R. (2017): Inventaire des hybrides d Ophrys d Italie.- J. Eur. Orch. 49 (3-4): L auteur fait le recensement de tous les hybrides du genre Ophrys décrits 566 Journal Europäischer Orchideen 49 (3-4): 2017.

3 d Italie. C est le premier travail sur ce sujet en Italie qui s inscrit dans un vaste projet de recenser les plantes endémiques de ce pays. Sont indiqués les binômes des parents tels qu annoncés par les descripteurs ainsi que selon la taxinomie des auteurs, avec indication de l herbier dans lequel sont déposés les types. Quelques statistiques sont présentées : la liste des hybrides d Ophrys décrits d Italie par région, la liste des herbiers et la présence des types, le nombre d hybrides décrits par des italiens et par des étrangers. Il est à noter que très peu de descriptions synonymes existent : seulement dix-sept, qui sont pour la plupart le fruit de personnes isolées ne disposant pas d une documentation exhaustive corrélée aux revues sans véritable conseil scientifique. Seulement cinquante-trois hybrides peuvent être considérés comme non endémiques stricts d Italie. Enfin vingt-neuf sont absents de la liste «The International Plant Names Index» (IPNI). 1. Introduction * * * Since 2012 several articles were related to "endemic plants of Italy" (PERUZZI 2015, PERUZZI et al. 2012, 2012, 2014a, 2014b, 2015, WAGENSOMMER et al. 2014). This work is a logical continuation of "The checklist della flora vascolare italiana" published by CONTI et al. (2005, 2007). Many articles came to supplement this overall work to treat with one, several species or a genus. The family of Orchidaceae, difficult to assess for non-specialized botanists (see, e.g, WALLNÖFER 2008: 266 et seq., DIMOPOULOS et al. 2016: 320 et seq.), was never yet treated as a whole. The hybrids of the Ophrys genus described from Italy were never compiled before in an article. The present paper is an inventory of these italian taxa ranging from 1865, year of the first description, to 2017 pro parte. Three hundred and three descriptions or combinations are listed. The taxa considered can be classified into three categories: the described taxa, the undescribed ones and the doubtful taxa. Very few descriptions of synonymous have been performed: only seventeen, which are mostly the result of individuals who do not have a comprehensive documentation correlated to journals without proper scientific advices, see e.g. BARTOLO & PULVIRENTI 1998, PERILLI 2016, SILLETTI & MEDAGLI Only twenty-four can be regarded as non strict endemic from Italy. Five appendices to make an assessment of the taxa of the three categories according to their presence in the various regions, in the herbaria, not listed in IPNI as well as the authors who described them. Finally the list of the papers in which the hybrid descriptions were published is also given. Journal Europäischer Orchideen 49 (3-4):

4 2. Materials and methods Taxonomy used is that of ROMOLINI & SOUCHE (2012). The list is arranged according to the alphabetic order of the hybrid parent names. Taxa considered can be classified into three categories: the described taxa, the undescribed ones and the doubtful taxa. The studied hybrids are among 328 while 29 combinations are given. The number of valid descriptions is 283. We can note 51 non valid descriptions (26 sine descriptio, 13 design. typi omissa, 4 invalid, 3 doubtful, 2 sphalm., 2 nom. nud, 1 nom. prov.). Furthermore, 64 descriptions are contrary to Art. H.5.2, 3 contrary to Art and 7 pro species. 67 hybrids have the two parents endemic of Italy, 140 have one endemic parent, and 53 have the two parents non endemic, while 25 are still difficult to classify. The data provided are classified as follows: - Name of the parents according to ROMOLINI & SoUche (2012) - Name of the hybrid - Original name of the parents used by the author(s) of the description - Basionym - Synonyms - Terra typica, type, date, herbarium - Notes (eg. mistake in interpretation, not listed in IPNI, unknown herbarium, ecc.). The main error encountered is that of the taxonomic rank according to CIN Art. H.5.2.: If the postulated or known parent taxa are of unequal rank the appropriate rank of the nothotaxon is the lowest of these ranks. Hybrids quoted in a list without intention of description, as well as description of a hybrid of Ophrys with another genus, are not taken into account. E.g.: - BOURNERIAS, J., BOURNERIAS, M. & J. FLAMENT (1992): Voyage d études de la S.F.O. dans les Pouilles (16-23 avril 1991).- Orchidophile (Asnières) 23(103): Which quote in a simple list : «de nombreux hybrides d'ophrys scolopax subsp. cornuta avec O. tenthredinifera, O. bombyliflora, O. apulica et O. garganica.» - NELSON E., (1962): Gestalwandel und Artbildung erörtet am Beispiel der Orchidaceen Europas und Mittelmeerländer, inbesondere der Gattung Ophrys mit einer Monographie und Ikonographie der Gattung Ophrys pages + 66 pl. NELSON describes it a simple lusus: sub Ophrys mammosa Serapias lingua. Taf. XLII, Fig. 1, Fig. a. Grèce, Corfou. - SACCARDO, P. A. (1881): Nova species italica ex genere Ophrydum - Ophrys integra Sacc.- Nuovo Giorn. Bot. Ital. 13(4): SACCARDO describes it a simple lusus of Ophrys apifera, sub Ophrys apifera O. bertolonii. 568 Journal Europäischer Orchideen 49 (3-4): 2017.

5 These examples were explained in ROMOLINI & SOUCHE (2012: 8, 23) After this list, some statistics are provided: - List of the described Ophrys hybrids from Italy by region and province, - List of the hybrids by herbarium and types, - Hybrids not listed in The International Plant Names Index (IPNI), - Authors who described the hybrids as authors or co-authors, - List of articles in which a description is published. Author of all photographs Fig is the author itself. 3. List of the Ophrys hybrids reported from Italy Ophrys annae Devillers-Tersch. & Devillers Ophrys bombyliflora Link, Ophrys sulcitana Scrugli, Todde & Cogoni, J. Eur. Orch. 35(3): Type: Sardegna, Sulcis, S. Antioco, in loco dicto Maladroxia, altitudine 80 m ca. 29.III.2003, Scrugli et Todde (CAG). Paratypus: ibidem, 31.III.2003 (CAG). Ophrys annae Devillers-Tersch. & Devillers Ophrys chestermanii (J.J.Wood) Gölz & H.R.Reinhard, Ophrys holoserica (Burm.f.) Greuter subsp. annae (Devillers-Tersch. & Devillers) H.Baumann, Giotta, Künkele, R.Lorenz & Piccitto Ophrys holoserica (Burm.f.) Greuter subsp. chestermanii J.J.Wood. GIOTTA & PICCITTO in BAUMANN & al. 1995: 238. sine descriptio. Type: [Italia, Sardinia] Tertenia: Scua d'orgius, 430 m (C. Giotta & M. Piccitto, ). Ophrys annae Devillers-Tersch. & Devillers Ophrys exaltata Ten. subsp. morisii (Martelli) Del Prete, Ophrys maladroxiensis Scrugli, Todde & Cogoni, Fl. Medit. 2: Sub Ophrys holoserica (Burman fil.) Greuter Ophrys morisii (Martelli) Soó. Type: [Italia] Sardinia, S. Antioco, loco vulgo Maladroxia dicto, , Scrugli. Todde & Cogoni (CAG). Paratypus: ibidem, , Scrugli, Todde & Cogoni (CAG). Ophrys annae Devillers-Tersch. & Devillers Ophrys tenthredinifera Willd. subsp. neglecta (Parl.) E.G.Camus, Ophrys spanui P.Delforge, Naturalistes Belges 78(3): Sub Ophrys annae J. & P. Devillers-Terschuren Ophrys tenthredinifera Willdenow. Type: Italia, Sardinia, provincia Cagliari, apud San Antioco, alt. s.m. 50 m, Journal Europäischer Orchideen 49 (3-4):

6 17.IV In herb. Pierre Delforge sub n Ophrys apifera Huds. Ophrys appennina Romolini & Soca, Ophrys petruccii Romolini & Soca, GIROS Notiz. 55: 58-59, Syn.: Ophrys albertiana nothosubsp. petruccii (Romolini & Soca) T.Bruschi & Polverelli, GIROS Orch. Spont. Eur. 58(2): 215. sub Ophrys apifera Huds. Ophrys holosericea [] subsp. appennina (Romolini & Soca) Kreutz. (Author of the species not indicated). Type: Italia, Toscana, Firenze, Firenze, Loc. Montorsoli, Via Viuccia, alt. 223 m. 32T V Coll. Romolini & Soca. MPU. n RS Ophrys apifera Huds. Ophrys apulica (O.Danesch & E.Danesch) O.Danesch & E.Danesch, Ophrys albertiana E.G. Camus nothosubsp. grovesii Gennaio, Gargiulo & Medagli, GIROS Notiz. 54: Sub Ophrys apifera Huds. Ophrys holosericea [] subsp. apulica (O.Danesch & E.Danesch) Buttler. (Author of the species not indicated). Type: Bacino Rottacapozza nord (Ugento, LE) m 2 s.l.m., legit R. Gennaio, 16.IV.2013 (in LEC Herbarium Universitatis Lupiensis). Ophrys apifera Huds. Ophrys aranifera s.l., Ophrys pseudoapifera Caldesio, Nuovo Giorn. Bot. Ital. 12(4): 'pseudo-apifera'. Sub Ophrys apifera Huds. Ophrys aranifera Huds. Type: [Italia,] in sylvaticis di Pergola, Castelraniero. Floret Maio, Iunio. Herbarium not indicated. Not listed in IPNI. Ophrys apifera Huds. Ophrys bombyliflora Link, Ophrys circae W.Rossi & Prola, Atti Accad. Naz. Lincei, Rendiconti Cl. Sci. Fis. Mat. Nat., ser. 8, 74(4): Type: 2.V.1982, G. Prola legit, RO. [località denominata «Il Precipizio», sul versante meridionale del Promontorio del Circeo (Lazio, Italia centrale). Un esemplare con cinque fiori, designato come olotipo, è depositato nell'herbarium Romanum presso l'istituto Botanico di Roma (RO); uno dei fiori è conservato in formalina al 5%. Un altro esemplare con tre fiori (isotipo) è depositato nell' Herbarium Centrale Italicum presso l'istituto Botanico di Firenze (FI)]. Ophrys apifera Huds. Ophrys candica (E.Nelson ex Soó) H.Baumann & Künkele, Ophrys morellensis O.Danesch & E.Danesch, Orchid. Eur. Ophrys-Hybriden: 224, Sub Ophrys apifera Huds. Ophrys candica (Nelson) O. & E. Danesch. 570 Journal Europäischer Orchideen 49 (3-4): 2017.

7 Fig. 1: Ophrys apifera O. appennina, Firenze, Toscana (FI), Fig. 2: Ophrys apulica O. brutia, Martina Franca, Puglia (TA), Fig. 3: Ophrys argentaria O. crabronifera, Castagneto Carducci, Toscana (LI), Fig. 4: Ophrys argentaria O. exaltata subsp. montis-leonis, Castagneto Carducci, Toscana (LI), Journal Europäischer Orchideen 49 (3-4):

8 Fig. 5: Ophrys biscutella Ophrys exaltata subsp. archipelagi, Manfredonia, Puglia (FG), Fig. 6: Ophrys brutia Ophrys dinarica, Roccaraso, Abruzzo (AQ), Fig. 7: Ophrys brutia Ophrys parvimaculata, Martina Franca, Puglia (TA), Fig. 8: Ophrys brutia Ophrys passionis subsp. majellensis, Palena, Abruzzo (CH), Journal Europäischer Orchideen 49 (3-4): 2017.

9 Type: Italien (Apulien), bei Lecce, leg. O. et E. Danesch, (IB). Syn.: Ophrys albertiana E.G.Camus nothosubsp. morellensis (O.Danesch & E.Danesch) Del Prete, Webbia 37(2): pro synonyme. Sub Ophrys apifera Huds. subsp. apifera Ophrys holoserica (N.L. Burm.) Greuter subsp. candica (E.Nelson ex Soó) Renz et Taubenh. Ophrys apifera Huds. Ophrys conradiae Melki & Deschâtres, Ophrys minuticauda Duffort nothosubsp. donorensis Gulli et M.P.Grasso, Orchidophile (Asnières) 28(128): Sub Ophrys apifera Huds. Ophrys scolopax Cav. subsp. conradiae (Melki & Deschâtres) H.Baumann, Giotta, Lorenz, Künkele & Piccitto. Type: Sardaigne, Donori, localité «Acqua sa Murta», bord d'une pinède. Leg. V. Gulli et M. P. Grasso (FI). Ophrys apifera Huds. Ophrys exaltata Ten. subsp. morisii (Martelli) Del Prete, Ophrys domus-maria M.P.Grasso, Orchidee (Hamburg) 40(3): Sub Ophrys arachnitiformis auct. fl. sard. Ophrys apifera Huds. Type: [Italia, Sardegna] Sulcis, prope pagum, "Domus de Maria", in loco dicto "Sa murta bianca", 11/04/1987, leg. M. P. Grasso, n 1 (Cag.); Isotypi: Ibidem , leg. M. P. Grasso, n 2 e 3 (Cag.), n 4 (Fi); Paratypi: Ibidem , leg. M. P. Grasso, n' 5 (Cag.) n' 6 (FI). Ophrys apifera Huds. Ophrys romolinii Soca, Ophrys vespertilio W.Rossi & Contorni, Orchid Rev. 104(1209): Sub Ophrys apifera Huds. Ophrys bertolonii Moretti. Type: Italy, Mt. Amiata, 20 May 1995, W. Rossi & M. Contorni s.n. (FI). Ophrys appennina Romolini & Soca Ophrys argentaria Devillers-Tersch. & Devillers, Ophrys torrensis Dekker, Eurorchis 11: Sub Ophrys argentaria Devillers-Terschuren & Devillers Ophrys pseudoscolopax Moggridge. Type: Italia, Livorno, Castagneto Carducci, 200 m , leg. H. Dekker, in L. Ophrys appennina Romolini & Soca Ophrys bertolonii Moretti, Ophrys valparmensis J.J.Wood, Orchidee (Hamburg) 33(2): Sub Ophrys bertoloniiformis O. & E. Danesch Ophrys holoserica (Crantz) Greuter. Type: Italien: Provinz Parma, ein halb stabilisierter Erdrutsch unterhalb Curatico, in der Nähe der Brücke über den Fluß Parma, , D.D. Chesterman 27 (K, Holotyp!). Ophrys appennina Romolini & Soca Ophrys crabronifera Mauri, Ophrys monachorum O.Danesch & E.Danesch, Orchid. Eur. Ophrys- Journal Europäischer Orchideen 49 (3-4):

10 Hybriden: 225, contrary to Art. H.5.2. Sub Ophrys exaltata Ten. Ophrys fuciflora (Crantz) Moench subsp. fuciflora. Type: Italien, Mte Argentario, leg. O. et E. Danesch, (IB). Ophrys appennina Romolini & Soca Ophrys funerea Viv., Ophrys ferruginea W.Rossi & Liuti, Webbia 44(2): Sub Ophrys fusca Link Ophrys holoserica (Burm. fil.) W. Greuter. Type: [Italia] Lazio, M. Angiano, presso Bagni di Stigliano (RM), in area sottoposta a pascolo bovino, su terreno marnoso-calcareo, 300 m circa, 6.V.1989, W. Rossi et A. Liuti (FI). Ophrys appennina Romolini & Soca Ophrys romolinii Soca, Ophrys enobarbia Del Prete & G.Tosi, Atti Soc. Tosc. Sci. Nat. Mem. B, 88: (1981) contrary to Art. H.5.2. Sub Ophrys bertolonii Moretti Ophrys holoserica (N.L.Burm.) Greuter subsp. holoserica. Type: [Italia] Monte Argentario (Etruria), loco dicto "Croce", leg. G. Tosi, die 20.V.1981 [PI]. Syn.: Ophrys gumprechtii O.Danesch & E.Danesch nothosubsp. enobarbia (Del Prete & G.Tosi) Del Prete, Webbia 37(2): Ophrys appennina Romolini & Soca Ophrys tenthredinifera Willd. subsp. neglecta (Parl.) E.G.Camus, Ophrys maremmae O.Danesch & E.Danesch, Orchid. Eur. Ophrys-Hybriden: 225, contrary to Art. H.5.2. Sub Ophrys fuciflora (Crantz) Moench subsp. fuciflora Ophrys tenthredinifera Willd. Type: Italien (Toscana), P. dei Venti, leg. O. et E. Danesch, (IB). Ophrys appennina Romolini & Soca Ophrys tetraloniae W.P.Teschner, Ophrys cetonae Romolini & Soca, GIROS Orch. Spont. Eur. 58: Type: Italia. Toscana (SI). Sarteano, Monte Cetona, alt. 918 m s.l.m., 32T VI Coll. Romolini. FI 54/09. Ophrys apulica (O.Danesch & E.Danesch) O.Danesch & E.Danesch Ophrys bertolonii Moretti subsp. explanata (Lojac.) Soca, Ophrys valparmensis [] nothosubsp. hilleri Cristaudo, Galesi, R.Lorenz & Zelesny, J. Eur. Orch. 42(1): Sub Ophrys bertolonii subsp. explanata (Lojac.) Soca Ophrys holosericea subsp. apulica (O.Danesch & E.Danesch) Buttler. Type: [Italia] Sizilien, Ägadische Inseln, Favignana (TP), 0,35 ENE Punta Campana, 135 m, UTMED S TC , , leg. K. & R. Lorenz, FO Lo09.036b (STU). 574 Journal Europäischer Orchideen 49 (3-4): 2017.

11 Ophrys apulica (O.Danesch & E.Danesch) O.Danesch & E.Danesch Ophrys biscutella O.Danesch & E.Danesch, Ophrys rossiniae Medagli & Turco, GIROS Notiz. 47: contrary to Art. H.5.2. Sub Ophrys argolica Fleischm. subsp. biscutella O.Danesch & E.Danesch Ophrys holosericea (Burm.f.) Greuter subsp. apulica (O.Danesch & E.Danesch) Buttler. Type: [Italia] Monte Sacro, apud Mattinata (FG), legit P. Medagli, 22.IV.2009 (in LEC - Herbarium Universitatis Lupiensis). Ophrys apulica (O.Danesch & E.Danesch) O.Danesch & E.Danesch Ophrys bombyliflora Link, Ophrys resurrecta O.Danesch & E.Danesch, Orchid. Eur. Ophrys-Hybriden: 221, Type: Italien (Apulien), bei Lecce, leg. O. et E. Danesch, (IB). Ophrys apulica (O.Danesch & E.Danesch) O.Danesch & E.Danesch Ophrys brutia P.Delforge, Ophrys trazzonarae Romolini & Soca, GIROS Notiz. 55: 56, Type: Italia. Puglia. Taranto. Martina Franca. Buffaloria (Monte Trazzonara), alt. 404 m. 33T IV Coll. Romieg Soca. MPU. n RS Ophrys apulica (O.Danesch & E.Danesch) O.Danesch & E.Danesch Ophrys candica (E.Nelson ex Soó) H.Baumann & Künkele, Ophrys valdevariabilis O.Danesch & E.Danesch, Orchid. Eur. Ophrys- Hybriden: 221, Sub Ophrys apulica (Danesch) O. & E.Danesch Ophrys candica (Nelson) O. & E.Danesch. Type: Italien (Apulien), östlich Lecce, leg. O. et E. Danesch, (IB). Ophrys apulica (O.Danesch & E.Danesch) O.Danesch & E.Danesch Ophrys candica (E.Nelson ex Soó) H.Baumann & Künkele Ophrys tenthredinifera Willd. subsp. neglecta (Parl.) E.G.Camus, Ophrys hydruntensis O.Danesch & E.Danesch, Orchid. Eur. Ophrys- Hybriden: 221, Sub Ophrys apulica (Danesch) O. & E.Danesch Ophrys candica (Nelson) O. & E.Danesch Ophrys tenthredinifera Willd. Type: Italien (Apulien), bei Otranto, leg. O. et E. Danesch, (IB). Ophrys apulica (O.Danesch & E.Danesch) O.Danesch & E.Danesch Ophrys cinnabarina Romolini & Soca, Ophrys holosericea nothosubsp. robertiarum D Alonzo & Perilli, GIROS Orch. Spont. Eur. 60(1): as 'robertidarum'. Journal Europäischer Orchideen 49 (3-4):

12 sub O. holosericea subsp. apulica (O.Danesch & E.Danesch) Buttler O. holosericea subsp. cinnabarina (Romolini & Soca) Kreutz. Italia, Apulia, in Gargano ad Sanctum Marcum in Lamis, Foveanam provinciam), m 742 s.l.m., UTMWGS 84 33T E N ; leg. F. D Alonzo & M. Perilli, cons. in BI (Herbarium Horti Botanici Barensis), spec , data Ophrys apulica (O.Danesch & E.Danesch) O.Danesch & E.Danesch Ophrys classica Devillers-Tersch. & Devillers, Ophrys sanctimarci Perilli, GIROS Orch. Spont. Eur. 58(2): Type: [Italia, Puglia] località Valle della Torre, (San Marco in Lamis, Foggia), m 511 s.l.m., 33T E N. 03.IV.2015, M. Perilli, cons. in BI (Herbarium Horti Botanici Barensis), spec. n Ophrys apulica (O.Danesch & E.Danesch) O.Danesch & E.Danesch Ophrys conradiae Melki & Deschâtres, Ophrys difesagrandeana D Alonzo, GIROS Notiz. 57: Type: [Italia] Bosco Difesa Grande (Gravina in Puglia, Bari), m 426 s.l.m.; legit F. D Alonzo, 10.V.2014; in BI (Herbarium Horti Botanici Barensis) n campione Ophrys apulica (O.Danesch & E.Danesch) O.Danesch & E.Danesch Ophrys exaltata Ten., Ophrys turcoi Silletti & Medagli, Silvae: epublished, contrary to Art. H.5.2. Sub Ophrys exaltata Ten. subsp. mateolana (Medagli, D'Emerico, Bianco & Ruggiero) Paulus & Gack Ophrys holosericea (Burm.f.) Greuter subsp. apulica (O. et E. Danesch) Buttler. Type: [Puglia] Santeramo in Colle (Ba) località Pantarosa, legit G. N. Silletti in LEC. Photography of Ophrys exaltata Ten. Not listed in IPNI. Ophrys apulica (O.Danesch & E.Danesch) O.Danesch & E.Danesch Ophrys fuciflora s.l. (O. oxyrrhynchos subsp. celiensis?), Ophrys cosentiana H.Baumann & Künkele, Mitteilungsbl. Arbeitskreis Heimische Orchid. Baden-Württemberg 18(3): 406, Sub Ophrys apulica (O. & E. Danesch) O. & E. Danesch Ophrys holosericea (N.L.Burm.) W. Greuter. Type: Italia [Calabria] Cosenza: Cropalati, 350 m , leg. B.u. H. Baumann (STU). Ophrys apulica (O.Danesch & E.Danesch) O.Danesch & E.Danesch Ophrys incubacea Bianca, Ophrys franciniae Bianco, Medagli, d Emerico & Ruggiero, Webbia 42(2): 576 Journal Europäischer Orchideen 49 (3-4): 2017.

13 Type: [Italia] Apulia, Cassano Murge (Ba), in pascuis, m 400 ca., 22.IV.1986, P. Bianco, P. Medagli, S. D'Emerico et L. Ruggiero, n. 1 (FI). Isotypi: ibidem, n. 2 (BI). Syn.: Ophrys yvonnae Dekker, Eurorchis 4: pro synonyme. Italia, Foggia, Viëste, 15 m leg. H. Dekker, in L. Ophrys apulica (O.Danesch & E.Danesch) O.Danesch & E.Danesch Ophrys lacaitae Lojac., Ophrys delargeana P.Delforge nothosubsp. staraleana D Alonzo & Perilli, GIROS Orch. Spont. Eur. 59(1): Sub Ophrys holosericea subsp. apulica (O.Danesch & E.Danesch) Buttler O. lacaitae Lojac. Type: Italia, Apulia, Garganus (Fovea), loco dicto Starale prope Sanctum Marcium in Lamis, m 672 s.l.m., UTMWGS84 33T E N ; leg. F. D Alonzo & M. Perilli, cons. in BI (Herbarium Horti Botanici Barensis), spec , in data 26 maggio Ophrys apulica (O.Danesch & E.Danesch) O.Danesch & E.Danesch Ophrys oestrifera M.-Bieb. subsp. montis-gargani Van de Vijver & W.Looken, Ophrys turrica R.Lorenz & Gembardt, Mitteilungsbl. Arbeitskreis Heimische Orchid. Baden-Württemberg 19(3): contrary to Art. H.5.2. Sub Ophrys apulica (O.Danesch & E.Danesch) O.Danesch & E.Danesch Ophrys oestrifera M. Bieb. subsp. oestrifera. Type: Italia, Puglia, Gargano prope Manacore, Torre Usmai (UTM: WG , 60 m) , leg. C.Gembardt & R. Lorenz, Blütenanalyse Hb. R. Lorenz No (STU). Ophrys apulica (O.Danesch & E.Danesch) O.Danesch & E.Danesch Ophrys oxyrrhynchos Tod. subsp. celiensis (O.Danesch & E.Danesch) Del Prete, Ophrys caliandri O.Danesch & E.Danesch, Orchid. Eur. Ophrys-Hybriden: 222, contrary to Art. H.5.2. Sub Ophrys apulica (Danesch) O. & E.Danesch Ophrys fuciflora (Crantz) Moench subsp. celiensis O. & E.Danesch. Type: Italien (Apulien), bei Ostuni, leg. O. et E. Danesch, (IB). Ophrys apulica (O.Danesch & E.Danesch) O.Danesch & E.Danesch Ophrys parvimaculata (O.Danesch & E.Danesch) Paulus & Gack, Ophrys cosentiana H.Baumann & Künkele nothosubsp. nociana H.Baumann & Künkele, Mitteilungsbl. Arbeitskreis Heimische Orchid. Baden- Württemberg 18(3): 407, Sub Ophrys apulica (O. & E. Danesch) O. & E. Danesch Ophrys holosericea (N.L.Burm.) W. Greuter subsp. parvimaculata (O. & E. Danesch) O. & E. Danesch. Journal Europäischer Orchideen 49 (3-4):

14 Type: Italia, Puglia: Noci, 500 m , leg. B. &. H. Baumann (STU). Ophrys apulica (O.Danesch & E.Danesch) O.Danesch & E.Danesch Ophrys passionis Sennen, Ophrys coturii Romolini & Soca, GIROS Notiz. 55: 56-57, Type: Italia. Puglia. Taranto. Manduria. Bosco di Coturi, alt. 88 m. 33T IV Coll. Romieg Soca. MPU. n RS Ophrys apulica (O.Danesch & E.Danesch) O.Danesch & E.Danesch Ophrys romolinii Soca, Ophrys degiorgii Ruggiero, Bianco, Medagli & D Emerico, Atti Soc. Ital. Sci. Nat. Mus. Civico Storia Nat. Milano 129(4): Sub Ophrys apulica (O.Danesch & E.Danesch) O.Danesch & E.Danesch Ophrys bertolonii Moretti. Type: [Italia] «Apulia, Arnesano (Lecce), in pascuis, 20 m ca., 12.IV.86, legit: Ruggiero L. et Medagli P., n. 1 (in Herbario Sallentino «M. Marinosci» sub 305)». Isotypi: «Ibidem, 18.IV.86, Medagli P. et Ruggiero L. n. 2, 3, 4, 5 (ibidem sub 306, 307, 308, 309)». LEC? Not listed in IPNI. Ophrys apulica (O.Danesch & E.Danesch) O.Danesch & E.Danesch Ophrys tarentina Gölz & H.R.Reinhard, Ophrys marinoscii Ruggiero, Bianco, Medagli & D Emerico, Atti Soc. Ital. Sci. Nat. Mus. Civico Storia Nat. Milano 129(4): Sub Ophrys apulica (O.Danesch & E.Danesch) O.Danesch & E.Danesch Ophrys tarentina Gölz et Reinhard. Type: [Italia] «Apulia, Martina Franca (Taranto), m 400 ca., 24.III.87, legit: D'Emerico S. et Medagli P. (in Herbario Sallentino «M. Marinosci» sub 345)». LEC? Not listed in IPNI. Ophrys apulica (O.Danesch & E.Danesch) O.Danesch & E.Danesch Ophrys tenthredinifera Willd. subsp. neglecta (Parl.) E.G.Camus, Ophrys salentina O.Danesch & E.Danesch, Orchid. Eur. Ophrys-Hybriden: 222, Sub Ophrys apulica (Danesch) O. & E.Danesch Ophrys tenthredinifera Willd. Type: Italien (Apulien), bei Carpignano Salentino, leg. O. et E. Danesch, (IB). Ophrys aranifera s.l. Ophrys bertolonii Moretti, Ophrys pseudobertolonii Murr, Deutsche Bot. Monatsschr. 16(12): As «pseudo-bertolonii». Sub Ophrys aranifera Huds. Ophrys bertolonii Moretti. 578 Journal Europäischer Orchideen 49 (3-4): 2017.

15 Type: [Italia] bei Vigolo Vattaro nächst Trient. Mitte Mai 1898, leg. E. Gelmi. Syn.: Ophrys gelmii Murr, Deutsche Bot. Monatsschr. 16(12): [Italia] bei Vigolo Vattaro nächst Trient. Mitte Mai 1898, leg. E. Gelmi. sub Ophrys aranifera Huds. Ophrys bertolonii Moretti, Syn.: Ophrys disiecta Murr, Deutsche Bot. Monatsschr. 19(8): [Italia] Nago-Torbole. Mai Not listed in IPNI. Syn.: Ophrys saratoi E.G.Camus nothosubsp. gelmii (Murr) H.Baumann & Künkele, Mitteilungsbl. Arbeitskreis Heimische Orchid. Baden-Württemberg 18(3): Bas.: Ophrys gelmii Murr, Deutsche Bot. Monatsschr. 16(12): sub Ophrys pseudobertolonii Murr subsp. pseudobertolonii Ophrys sphegodes Mill. subsp. sphegodes. [Italia] bei Vigolo Vattaro nächst Trient , leg. E. Gelmi. Ophrys aranifera s.l. Ophrys bertolonii Moretti Ophrys insectifera L., Ophrys daneschiana W.J.Schrenk, Amer. Orchid Soc. Bull. 50(4): (413) Sub Ophrys bertoloniiformis O.Danesch & E.Danesch Ophrys insectifera L. Ophrys sphecodes Mill. Type: FR. leg. W. J. Schrenk 5/17/80, Albisano, Mte. Baldo, Lago di Garda, Italy. Ophrys aranifera s.l. Ophrys bertolonii Moretti subsp. bertoloniiformis (O.Danesch & E.Danesch) H.Sund., Sub Ophrys pseudobertolonii Murr subsp. bertoloniiformis (O.Danesch & E.Danesch) H.Baumann & Künkele Ophrys sphegodes Mill. LORENZ & GEMBARDT, Mitteilungsbl. Arbeitskreis Heimische Orchid. Baden- Württemberg 1987: 512, 580. sine descriptio. Type: [Italia, Puglia, Gargano] 1) 6,5 km WNW Monte San Angelo, UTM: 72.40, 700 m. H. R.& R. Reinhard ) 2,8 km SSE Rodi Garganico, UTM: , 200 m. C. Gembardt Ophrys aranifera s.l. Ophrys exaltata Ten. subsp. archipelagi (Gölz & H.R.Reinhard) Del Prete, Ophrys trombettensis Soca, Monde Pl. 92(460): Sub Ophrys archipelagi Gölz & H.R.Reinhard Ophrys sphegodes Miller. Type: Italia, Puglia (Foggia), Monte Gargano: inter Sanicandro et Cagnano, Coste di Trombetta, in septemtrionale latere Mt Pizzulo, alt. 190 m. 12. IV MPU n RS Ophrys aranifera s.l. Ophrys incubacea Bianca, Ophrys todaroana Macch., Nuovo Giorn. Bot. Ital. 13(4): nom. nud. Sub Ophrys aranifera L. Ophrys atrata Lindl. Bas.: Arachnites aranifera-panormitana Tod. Journal Europäischer Orchideen 49 (3-4):

16 Type: [Italia, Sardegna] Maggio e giugno. Ophrys aranifera s.l. Ophrys lunulata Parl., Ophrys frascae Galesi, J. Eur. Orch. 31(2): contrary to Art. H.5.2. Sub Ophrys lunulata Parl. Ophrys sphegodes Miller subsp. sphegodes. Type: [Italia, Sicilia, Vittoria (RG)] nei pressi di C.se Frasca (VA ). All'interno di un rimboschimento ad Eucalyptus camaldulensis, 130m, 26.III.1995, R. Galesi (CAT). Ophrys aranifera s.l. Ophrys oestrifera M.-Bieb. subsp. montis-gargani Van de Vijver & W.Looken, Ophrys philippei Gren. nothosubsp. calenae O.Danesch & E.Danesch, Orchid. Eur. Ophrys-Hybriden: 228, 97, 78e as "philippii". Sub Ophrys scolopax Cav. subsp. cornuta (Stev) E. G. Camus Ophrys sphecodes Mill. subsp. sphecodes. Type: Italien (Mte Gargano), westlich Vieste, leg. O. et E. Danesch, (IB). Syn.: Ophrys calenae (O.Danesch & E.Danesch) H.Baumann & Künkele, Mitteilungsbl. Arbeitskreis Heimische Orchid. Baden-Württemberg 18(3): Sub Ophrys oestrifera M. Bieb. subsp. oestrifera Ophrys sphegodes Mill. subsp. sphegodes. Ophrys aranifera s.l. Ophrys oxyrrhynchos Tod., Ophrys galesii Soca, Monde Pl. 91(458): Sub Ophrys oxyrrhynchos Tod. Ophrys sphegodes Miller. Type: Italia, Sicilia (Ragusa), loco dicto C.da Zotte, Valle Fiume Dirillo (S. Pietro). 16.IV MPU n RS Ophrys aranifera s.l. Ophrys passionis Sennen, Ophrys biancoae Medagli, d Emerico & Ruggiero, Orchidophile (Asnières) 20(85): contrary to Art. H.5.2. Sub Ophrys garganica E.Nelson ex O.Danesch & E.Danesch Ophrys sphegodes Mill. subsp. sphegodes. Type: Italia, Apulia, Castellaneta (TA), in pascuis. m 300 ca, 15.IV.1988, P. Medagli et S. d'emerico. ln herb. Gruppo Naturalisti Salentini di Lecce sub 351, fig. 2. Ophrys aranifera s.l. Ophrys speculum Link, Sub Ophrys aranifera speculum. MACCHIATI, Nuovo Giorn. Bot. Ital. 1881: Nom. nud. Type: [Italia, Sardegna] nel Sassarese a Baddimanna. Marzo ed aprile. Ophrys aranifera s.l. Ophrys tarentina Gölz & H.R.Reinhard, 580 Journal Europäischer Orchideen 49 (3-4): 2017.

17 Ophrys calabrica H.Baumann & Künkele, Mitteilungsbl. Arbeitskreis Heimische Orchid. Baden-Württemberg 18(3): Sub Ophrys sphegodes Mill. subsp. sphegodes Ophrys tarentina Gölz & H.R.Reinhard. Type: Italia, Puglia: Martina Franca, 500 m , leg. B. & H. Baumann (STU). Ophrys argentaria Devillers-Tersch. & Devillers Ophrys bombyliflora Link, Ophrys ansedoniensis Soca, Monde Pl. 90(452): Type: Italia, Toscana, apud Ansedonia. 16.IV MPU n RS Ophrys argentaria Devillers-Tersch. & Devillers Ophrys crabronifera Mauri, Ophrys rabautei Soca, Monde Pl. 90(452): Type: Italia, Toscana, Livorno, Marina di Castagnetto-Donoratico. 17.IV MPU n RS Ophrys argentaria Devillers-Tersch. & Devillers Ophrys exaltata Ten. subsp. montis-leonis (O.Danesch & E.Danesch) Soca, Ophrys crocii Soca, Monde Pl. 90 (452): Sub Ophrys argentaria Devillers-Tersch. & Devillers Ophrys tyrrhena Gölz & H.R.Reinhard. Type: Italia, Toscana, Livorno, Marina di Castagnetto-Donoratico. 17.IV MPU n RS Ophrys argentaria Devillers-Tersch. & Devillers Ophrys incubacea Bianca, Ophrys turchinae Gransinigh & Buono, GIROS Notiz. 47: 75 (68-75; figs.) Type: Italia, Lazio, Monteromano (provincia Viterbo), località Macchia della Turchina, q. 171 m UTM E N. 13 aprile 2009 in Erbario della Tuscia (UTV), Viterbo, legit. S. Buono & E. Gransinigh. Ophrys argentaria Devillers-Tersch. & Devillers Ophrys passionis Sennen, Ophrys gembardtii Soca, Monde Pl. 90(452): Sub Ophrys argentaria Devillers-Tersch. & Devillers Ophrys garganica O.Danesch & E.Danesch. Type: Italia, Toscana, Livorno, Marina di Castagnetto-Donoratico. 17.IV MPU n RS Ophrys argentaria Devillers-Tersch. & Devillers Ophrys tarquinia P.Delforge, Ophrys sassettae Romolini & Soca, GIROS Notiz. 38: Type: Italia, Toscana, Sassetta (Livorno Provincia), loco dicto Campo di Carlo. alt. 122 m. (UTM 32T PN37). 22.IV Coll. Romolini 40/04. FI. Journal Europäischer Orchideen 49 (3-4):

18 Ophrys argentaria Devillers-Tersch. & Devillers Ophrys tenthredinifera Willd. subsp. neglecta (Parl.) E.G.Camus, Ophrys teresae Soca, Monde Pl. 92(460): Sub Ophrys argentaria Devillers-Tersch. & Devillers Ophrys tenthredinifera Willd. Type: Italia, Toscana, Grosseto, Alberese, Parco della Maremma. 10.IV MPU n RS Syn.: Ophrys garganica E.Nelson ex O.Danesch & E.Danesch Ophrys sphecodes Mill. Ophrys tenthredinifera Willd. SCHRENK, Amer. Orchid Soc. Bull. 1981: 131, 134. nom. nud. sine descriptio. Ansedonia, Italy. 4/28/82. Fig. 12, photography of Ophrys argentaria Devillers-Tersch. & Devillers Ophrys tenthredinifera Willd. subsp. neglecta (Parl.) E.G.Camus [Ophrys teresae Soca]. Ophrys ausonia Devillers, Devillers-Tersch. & P.Delforge Ophrys lacaitae Lojac., Ophrys rasettii W.Rossi & Minutillo, Webbia 43(2): Sub Ophrys lacaitae Lojacono Ophrys sphegodes Miller. Type: [Italia] Lazio meridionale, M.ti Aurunci, M. Lapillo, 750 m, 12.VI.1988, W. Rossi et F. Minutillo (FI). Ophrys ausonia Devillers, Devillers-Tersch. & P.Delforge Ophrys promontorii O.Danesch & E.Danesch, Ophrys terrae-laboris W.Rossi & Minutillo, Webbia 43(2): Sub Ophrys sphegodes Miller Ophrys promontorii O. & E. Danesch. Type: [Italia] Lazio meridionale, M.ti Aurunci, M. Revole, versante SE, 800 m circa, su gariga calcarea, 1.V.1985, W. Rossi et F. Minutillo» (FI). Uno dei fiori è conservato in una soluzione idroalcolica contenente piccole quantità di glicerina e formaldeide. Ophrys bertolonii Moretti Ophrys incubacea Bianca, Ophrys pseudaranifera Murr, Deutsche Bot. Monatsschr. 16(12): as «pseud-aranifera». Sub Ophrys aranifera Huds. Ophrys bertolonii Moretti. Type: [Italia] bei Vigolo Vattaro nächst Trient. Mitte Mai 1898, leg. E. Gelmi. Syn.: Ophrys araniferiformis Dalla Torre & Sarnth., Fl. Tirol 6(1): [Italia] bei Nago-Torbole. Not listed in IPNI. Ophrys bertolonii Moretti Ophrys occitanica nom. prov., Ophrys baldensis P.Delforge, Naturalistes Belges 70(3): contrary to Art. H.5.2. Sub Ophrys benacensis (Reisigl) Danesch & Ehrendorfer Ophrys holoserica (Burm.) Greuter subsp. holoserica. Type: Italia,Veneto, provincia Verona, apud Albisano, supra montem Baldo, 582 Journal Europäischer Orchideen 49 (3-4): 2017.

19 In herb. Pierre Delforge sub n Ophrys bertolonii Moretti subsp. bertoloniiformis (O.Danesch & E.Danesch) H.Sund. Ophrys biscutella O.Danesch & E.Danesch, Ophrys permutata O.Danesch & E.Danesch, Orchid. Eur. Ophrys-Hybriden: 224, contrary to Art. H.5.2. Sub Ophrys biscutella O. & E. Danesch Ophrys bertoloniiformis O. & E. Danesch subsp. bertoloniiformis. Type: Italien (Mte Gargano), bei Monte S. Angleo, leg. O. et E. Danesch, (IB). Ophrys bertolonii Moretti subsp. bertoloniiformis (O.Danesch & E.Danesch) H.Sund. Ophrys bombyliflora Link, Ophrys manganaroi Soca nothosubsp. manganaroi, Caesiana 19(43): Type: Italia. Puglia. Foggia. Manfredonia. Posta Manganaro. Alt. 403 m s.l.m. 33T IV Coll. Romieg Soca. Cons. in herb. MPU. sub n RS Ophrys bertolonii Moretti subsp. bertoloniiformis (O.Danesch & E.Danesch) H.Sund. Ophrys exaltata Ten. subsp. archipelagi (Gölz & H.R.Reinhard) Del Prete, Ophrys neocamusii Godfery nothosubsp. lupoi Soca, Caesiana 19(43): Type: Italia. Puglia. Foggia. Mattinata. Alt. 214 m s.l.m. 33T IV Coll. Romieg Soca. Cons. in herb. MPU. sub n RS Ophrys bertolonii Moretti subsp. bertoloniiformis (O.Danesch & E.Danesch) H.Sund. Ophrys incubacea Bianca, Ophrys lorenzii Soca, Monde Pl. 92(460): Sub Ophrys bertoloniiformis O.Danesch & E.Danesch Ophrys incubacea Bianca. Type: Italia, Puglia (Foggia), Monte Gargano: inter Monte San Angelo et Vico, Monte Croce, alt. 800 m. 28.IV MPU n RS Ophrys bertolonii Moretti subsp. bertoloniiformis (O.Danesch & E.Danesch) H.Sund. Ophrys lojaconoi P.Delforge, Ophrys pagliae Perilli, GIROS Orch. Spont. Eur. 58(2): contrary to Art. H.5.2. Type: [Italia, Puglia] località Ponte Ciccalento (San Marco in Lamis, Foggia), m 101 s.l.m., 33T 33T553731E N. 03.IV.2015; M. Perilli, cons. in BI (Herbarium Horti Botanici Barensis), spec. n Ophrys bertolonii Moretti subsp. bertoloniiformis (O.Danesch & E.Danesch) H.Sund. Ophrys passionis Sennen, Journal Europäischer Orchideen 49 (3-4):

20 Ophrys cantalupoi Soca nothosubsp. cantalupoi, Caesiana 19(43): Type: Italia. Puglia. Foggia. Manfredonia. Posta Manganaro. Alt. 410 m s.l.m. 33T IV Coll. Romieg Soca. Cons. in herb. MPU. sub n RS Ophrys bertolonii Moretti subsp. bertoloniiformis (O.Danesch & E.Danesch) H.Sund. Ophrys promontorii O.Danesch & E.Danesch, Ophrys azurea nsubsp. azurea H.Baumann & Künkele, Mitteilungsbl. Arbeitskreis Heimische Orchid. Baden-Württemberg 18(3): 516, Sub Ophrys promontorii O. & E. Danesch Ophrys pseudobertolonii Murr subsp. pseudobertolonii. Type: Italia, Puglia: Mte. Gargano, Mte. San Angelo, 500 m , leg. B. & H. Baumann (STU). Ophrys bertolonii Moretti subsp. bertoloniiformis (O.Danesch & E.Danesch) H.Sund. Ophrys romolinii Soca, Ophrys marzensis Soca nothosubsp. zonnoi Soca, Caesiana 19(43): Type: Italia. Puglia. Foggia. San Marco in Lamis. Zonno. Alt. 615 m s.l.m. 33T IV Coll. Romieg Soca. Cons. in herb. MPU. sub n RS Ophrys bertolonii Moretti subsp. bertoloniiformis (O.Danesch & E.Danesch) H.Sund. Ophrys sipontensis (Gumprecht) O.Danesch & E.Danesch, Ophrys castellerana P.Delforge & C.Delforge, Naturalistes Belges 67(5/6): contrary to Art. H.5.2. Sub Ophrys sipontensis (Gumpr.) Danesch Ophrys bertoloniiformis subsp. bertoloniiformis Danesch. Type: Italia, provincia Foggia, San Giovanni Rotondo, super montem Cope di Castellera, alt. 550m In herb. P. Delforge sub n 86/6. Ophrys bertolonii Moretti subsp. bertoloniiformis (O.Danesch & E.Danesch) H.Sund. Ophrys tarentina Gölz & H.R.Reinhard, Ophrys motolesensis Soca, Monde Pl. 92(460): Sub Ophrys bertoloniiformis O.Danesch & E.Danesch Ophrys tarentina Gölz & H.R.Reinhard. Type: Italia, Puglia, Taranto, apud San Simone (Crispiano). 17.IV MPU n RS Ophrys bertolonii Moretti subsp. bertoloniiformis (O.Danesch & E.Danesch) H.Sund. Ophrys tenthredinifera Willd. subsp. neglecta (Parl.) E.G.Camus, Ophrys daneschiorum P.Delforge, Naturalistes Belges 69(2): contrary to Art. H.5.2. Sub Ophrys bertoloniiformis O. & E.Danesch subsp. bertoloniiformis Ophrys tenthredinifera Willd. 584 Journal Europäischer Orchideen 49 (3-4): 2017.

21 Type: Italia, Puglia, provincia Foggia, apud Monte Sant'Angelo, In herb. Pierre Delforge sub n 86/7. Ophrys bertolonii Moretti subsp. explanata (Lojac.) Soca Ophrys incubacea Bianca, Ophrys melkii Soca, Monde Pl. 91(458): Sub Ophrys explanata (Lojac.) P.Delforge Ophrys incubacea Bianca. Type: Italia, Sicilia (Enna), Pinède sur sable; inter Piazza Armerina et Barrafranca, in septemtrionale latere Montagna di Marzo. 15.IV MPU n RS Ophrys bertolonii Moretti subsp. explanata (Lojac.) Soca Ophrys lunulata Parl., Ophrys mazzolana Falci, S.A.Giardina & Serio, J. Eur. Orch. 40(3): 601 ( ; figs. 1-2) Type: Sicilia, Vittoria (Ragusa), Monti Iblei presso la Riserva Naturale Orientata "Pino d'aleppo" in Contrada Salina, (UTMED 1950 VA ), (leg. Falci, Giardina & Serio (PAL). Sub Ophrys explanata (Lojac.) Delforge Ophrys lunulata Parl. Syn.: Ophrys bertolonii subsp. explanata (Lojac.) Soca Ophrys lunulata Parl. CAMPO, GIROS Notiz. 2012: sine nomine, nom. inval., design. typi omissa, pro synonym. iter Pantanelle apud Gela (CL), legit V. Campo IGM fg. 272 U.T.M. VB , 100 m circa s.l.m. Ophrys bertolonii Moretti subsp. explanata (Lojac.) Soca Ophrys romolinii Soca, Ophrys marzensis Soca, Monde Pl. 92(460): Sub Ophrys bertolonii Moretti Ophrys explanata (Lojac.) P.Delforge. Type: Italia, Sicilia (Enna). Pinède sur sable; inter Piazza Armerina et Barrafranca, in septemtrionale latere Montagna di Marzo. 23.IV MPU n RS Ophrys bertolonii Moretti subsp. explanata (Lojac.) Soca Ophrys speculum Link, Ophrys denisiana nothosubsp. denisiana H.Baumann, Künkele & R.Lorenz, J. Eur. Orch. 36(2): Sub Ophrys bertolonii Moretti subsp. explanata (Lojac.) Soca Ophrys ciliata Biv.-Bern. Type: Sicilia, ad pagum Mondello, , leg B. & H. Baumann (STU). Ophrys bertolonii Moretti subsp. explanata (Lojac.) Soca Ophrys tenthredinifera Willd. subsp. grandiflora (Ten.) Kreutz, Ophrys insengae (Tod.) Nym., Syll. Fl. Eur., suppl: pro species. Bas.: Arachnites insengae Tod., Nuov. Gen. Sp Journal Europäischer Orchideen 49 (3-4):

22 Type: Sicilia (Palermo). Italia, Sicilia, Palermo: Mte Catalfano [Nelle colline calcaree e nelle falde dei monti attorno Palermo. Fiorisce nei mesi di Marzo ed Aprile]. Syn.: Ophrys insengae (Tod.) Ces., Pass. & Gibelli, Comp. Fl. Ital. [Cesati]: as inzengae. Bas.: Arachnites Tod. [Italia] Monte Catalfano in Sicilia. Not listed in IPNI. Ophrys bertolonii Moretti subsp. saratoi (E.G.Camus) Soca, Monde Pl. 96(472): , Ophrys saratoi E.G.Camus, J. Bot. (Morot) 7(8): Bas.: Ophrys araniferobertoloni Barla & Sarato, Fl. Ill. Nice, Orchid.: 70, Pl. 58, Fig. 16, 17, Syn.: Ophrys barlae E.G.Camus, J. Bot. (Morot) 7(8): pro synonym. Bas.: Ophrys Bertoloni hybride c. bilineata Barla, Fl. Ill. Nice, Orchid.: 70, Pl. 58, Fig. 19 à Syn.: Ophrys araniferobertoloni Barla & Sarato, Fl. Ill. Nice, Orchid.: 70, Pl. 58, Fig. 16, 17, 'aranifero-bertoloni'. au col de Villefranche, avril Leg. C. Sarato. sub Ophrys aranifera Huds. Ophrys bertolonii Moretti. Syn.: Ophrys Bertoloni hybride c. bilineata Barla, Fl. Ill. Nice, Orchid.: 70, Pl. 58, Fig. 19 à sub Ophrys aranifera Huds. Ophrys bertolonii Moretti. Montgros, avril Leg. C. Sarato. Not listed in IPNI. Syn.: Ophrys bilineata (Barla) Fiori, Fl. Italia 1(1): Bas.: Ophrys Bertoloni hybride c. bilineata Barla, Fl. Ill. Nice, Orchid.: 70, Pl. 58, Fig. 19 à sub Ophrys aranifero Bertolonii. Not listed in IPNI. Ophrys bertolonii s.l. Ophrys fuciflora s.l., Ophrys goiranii Cif. & Giacom., Nomenclator Fl. Ital. 1: nom. nud. Sub Ophrys Bertolonii fuciflora. Not listed in IPNI. Ophrys biscutella O.Danesch & E.Danesch Ophrys exaltata Ten. subsp. archipelagi (Gölz & H.R.Reinhard) Del Prete, Ophrys garganensis Soca, Caesiana 9: Sub Ophrys archipelagi Gölz & H.R.Reinhard Ophrys biscutella O.Danesch & E.Danesch. Type: Italia, Puglia (Foggia), Monte Gargano: inter Monte San Angelo et Vico, Monte Croce, inter Valle delle Rose et Valle degli Aceri, alt. 770 m. 27.IV MPU n RS Ophrys biscutella O.Danesch & E.Danesch Ophrys gracilis (Büel, O.Danesch & E.Danesch) Paulus, Sub Ophrys argolica [] biscutella (O. Danesch & E. Danesch) Kreutz 586 Journal Europäischer Orchideen 49 (3-4): 2017.

23 Ophrys holosericea [] gracilis (Büel, O. Danesch & E. Danesch) Büel, O. Danesch & E. Danesch. (Author of the species not indicated). Perilli, GIROS Notiz. 2012: 109. sine nomine, nom. inval., design. typi omissa Type: [Italia, Puglia] Monte Celano, San Marco in Lamis (FG). Ophrys biscutella O.Danesch & E.Danesch Ophrys minipassionis Romolini & Soca, Ophrys boscoquartensis O.Danesch & E.Danesch, Orchidee (Hamburg) 21(6): , contrary to Art. H.5.2. Sub Ophrys biscutella O. & E.Danesch Ophrys sphecodes Mill. subsp. sphecodes. Type: [Italia, Puglia] Mte. Gargano, 9 km westlich Vieste, leg. O. et E. Danesch Institut für botanische Systematik und Geobotanik der Universität Innsbruck (IB). Syn.: Ophrys boscoquartensis O.Danesch & E.Danesch nothosubsp. difesae Perilli, GIROS Orch. Spont. Eur. 59(1): sub Ophrys crabronifera [] subsp. biscutella (O.Danesch & E.Danesch) Klaver & Kreutz Ophrys sphegodes [] subsp. minipassionis (Romolini & Soca) Biagioli & Grünanger. Italia, Apulia, Garganus (Fovea), Saltu Difesae Sancti Matthaei, prope Sanctum Marcium in Lamis, m 867 s.l.m., UTMWGS84 33T E N ; leg. M. Perilli, cons. in BI (Herbarium Horti Botanici Barensis), spec , data 24 aprile Author of the species not indicated. Ophrys biscutella O.Danesch & E.Danesch Ophrys oestrifera M.-Bieb. subsp. montis-gargani Van de Vijver & W.Looken, Ophrys carpinensis O.Danesch & E.Danesch, Orchidee (Hamburg) 21(6): 361, contrary to Art. H.5.2. Sub Ophrys biscutella O. & E.Danesch Ophrys scolopax Cav. subsp. cornuta (Stev.) E. G. Cam. Type: [Italia, Puglia] Mte. Gargano, 4 km südlich Carpino, leg. O. et E. Danesch Institut für botanische Systematik und Geobotanik der Universität Innsbruck (IB). Ophrys biscutella O.Danesch & E.Danesch Ophrys parvimaculata (O.Danesch & E.Danesch) Paulus & Gack, Ophrys pizzulensis Soca, Monde Pl. 92(460): Type: Italia, Puglia (Foggia), Monte Gargano: inter Sanicandro et Cagnano, Coste di Trombetta, in septemtrionale latere Mt Pizzulo, alt. 190 m. 12.IV MPU n RS Ophrys biscutella O.Danesch & E.Danesch Ophrys promontorii O.Danesch & E.Danesch, Ophrys vernonensis O.Danesch & E.Danesch, Orchid. Eur. Ophrys- Journal Europäischer Orchideen 49 (3-4):

24 Hybriden: 224, Type: Italien (Mte Gargano), M. Vernone, leg. O. et E. Danesch, (IB). Ophrys biscutella O.Danesch & E.Danesch Ophrys romolinii Soca, Ophrys salvatoris O.Danesch & E.Danesch, Orchid. Eur. Ophrys-Hybriden: 223, Sub Ophrys biscutella O. & E. Danesch Ophrys bertolonii Mor. Type: Italien (Mte Gargano), S. Salvatore, leg. O. et E. Danesch, (IB). Ophrys biscutella O.Danesch & E.Danesch Ophrys sipontensis (Gumprecht) O.Danesch & E.Danesch, Ophrys cornelli O.Danesch & E.Danesch, Orchid. Eur. Ophrys-Hybriden: 224, Type: Italien (Mte Gargano), bei Giovanni Rotondo, leg. O. et E. Danesch, (IB). Ophrys biscutella O.Danesch & E.Danesch Ophrys tenthredinifera Willd. subsp. neglecta (Parl.) E.G.Camus, Ophrys montis-angeli O.Danesch & E.Danesch, Orchidee (Hamburg) 21(6): , Sub Ophrys biscutella O. & E.Danesch Ophrys tenthredinifera Willd. Type: [Italia, Puglia] Mte. Gargano, 7 km westlich Monte S. Angelo, leg. O. et E. Danesch, Institut für botanische Systematik und Geobotanik der Universität Innsbruck (IB). Ophrys bombyliflora Link Ophrys candica (E.Nelson ex Soó) H.Baumann & Künkele, Ophrys medaglii Turco, Ruggiero, Gennaio & D Emerico, Thalassia Sal. 34: 27. [epublished]. Type: [Italia] apud S. Cataldo (Le), m 20 circa s.l.m., legit L. Ruggiero et A. Turco, 12.V.2009 (in LEC - Herbarium Universitatis Lupiensis). Three photographs = Ophrys candica! Ophrys bombyliflora Link Ophrys classica? (or argentaria), Ophrys hoeppneri Ruppert, Repert. Spec. Nov. Regni Veg. 31: Sub Ophrys aranifera Huds. Ophrys bombyliflora Link. Type: [Italia, Toscana, Grosseto] Me. Argentario, bei einem alten Turm über Cap Lividonia Nro. 213 & Nro leg. Höppner. Ophrys bombyliflora Link Ophrys corsica Soleirol ex G.Foelsche & W.Foelsche, Ophrys domitia Del Prete, Atti Soc. Tosc. Sci. Nat. Pisa Mem. B, 85: Journal Europäischer Orchideen 49 (3-4): 2017.

25 (1978) contrary to Art. H.5.2. Sub Ophrys bombyliflora Link Ophrys lutea (Gouan) Cav. subsp. murbeckii (Fleischm.) Soó. Type: [Italia, Toscana, Grosseto] PI, Argentario prope Villa Domitia ad Domitianii Balnea. Leg. C. Del Prete et G. Tosi, 26 Majo in Del Prete C., Webbia 37(2): 254. parents = Ophrys bombyliflora Link Ophrys lutea Cav. subsp. minor (Tod.) Danesch O. et E. contrary to Art. H.5.2. Not listed in IPNI. Ophrys bombyliflora Link Ophrys crabronifera Mauri, Del Prete & al., Le orchidee spontanee della provincia di Grosseto, 1982: 65. sine nomine neque descriptione. Type: [Italia] Etruria. Ophrys bombyliflora Link Ophrys discors Bianca, Ophrys martinii [] nothosubsp. ragonesii Galesi & R.Lorenz, J. Eur. Orch. 42(1): Author of the species not indicated. Sub Ophrys bombyliflora Link Ophrys oxyrrhynchos Tod. subsp. biancae Galesi, Cristaudo & Maugeri. Type: [Italia, Sicilia] Costa Reitani (WA Vendicari, Noto), in una gariga a Corydothymus capitatus su litorale roccioso a circa 50 m di distanza dal mare, 2 m s.l.m., , R. Galesi (CAT). Ophrys bombyliflora Link Ophrys exaltata Ten., Ophrys ragusana H.Baumann & Künkele, Mitteilungsbl. Arbeitskreis Heimische Orchid. Baden-Württemberg 16(4): Type: [Italia] Insula Sicilia, ad pagum Marina di Ragusa , leg. H. Baumann, in Herbario Staatliches Museum für Naturkunde in Stuttgart (STU) conservatur. Ophrys bombyliflora Link Ophrys exaltata Ten. subsp. archipelagi (Gölz & H.R.Reinhard) Del Prete, Sub Ophrys bombyliflora Link Ophrys arachnitiformis Gren. et Phil. DANESCH & DANESCH, Orchideen Europas. Ophrys Hybriden 1972: 142. sub Ophrys semibombyliflora Bergon & Camus. sine descriptio. Italien (Mte Gargano). Ophrys bombyliflora Link Ophrys fusca s.l., Sub Ophrys bombyliflora Ophrys fusca. TICHY, Mitteilungsbl. Arbeitskreis Heimische Orchid. Baden-Württemberg 1977: 183. sine descriptio. Gavorrane (sic), Ophrys bombyliflora Link Ophrys incubacea Bianca, Ophrys cosana H.Baumann & Künkele, Mitteilungsbl. Arbeitskreis Heimische Orchid. Baden-Württemberg 18(3): 438, Journal Europäischer Orchideen 49 (3-4):

26 Type: Italia [Toscana] Grosseto: Mte. Argentario, Porto S. Stefano, 80 m , leg. B. & H. Baumann (STU). Ophrys bombyliflora Link Ophrys maritima Pacifico & Soca, Ophrys stivii Romolini & Soca, GIROS Orch. Spont. Eur. 58: Type: Italia. Toscana (LU). Lucca. Santa Maria del Giudice, Monte Cotrozzi, alt. 115 m s.l.m., 32T IV Coll. Romolini. FI. Romolini 15/2014. Ophrys bombyliflora Link Ophrys minipassionis Romolini & Soca, Ophrys lessonae Romolini & Soca, GIROS Orch. Spont. Eur. 58(2): Type: Italia. Toscana. Firenze. Fiesole. Caldine. alt. 182 m s.l.m. 32T IV Coll. Rolando Romolini. FI. n 24/2015. Ophrys bombyliflora Link Ophrys oxyrrhynchos Tod., Ophrys martinii O.Danesch & E.Danesch, Orchid. Eur. Ophrys-Hybriden: 227, as 'martini'. Type: Sizilien, bei Syracus, leg. O. et E. Danesch, (IB). Ophrys bombyliflora Link Ophrys parvimaculata (O.Danesch & E.Danesch) H.F.Paulus & Gack), Ophrys ozantina Gennaio, GIROS Notiz. 41: Type: [Italia] in loco dicto "Rottacapozza" apud urbem Ausentum (odie Ugento), 7.IV.2004, legit. R.Gennaio (LEC). Ophrys bombyliflora Link Ophrys passionis Sennen, Ophrys daunia H.Baumann & Künkele, Mitteilungsbl. Arbeitskreis Heimische Orchid. Baden-Württemberg 18(3): 436, Sub Ophrys bombyliflora Link Ophrys garganica E.Nelson ex O. & E. Danesch. Type: Italia, Puglia: Mte. Gargano, Vieste, 30 m , leg. B. & H. Baumann (STU). Ophrys bombyliflora Link Ophrys romolinii Soca, Ophrys cataldii Gölz, Orchidee (Hamburg) 22(4): 163, as 'cataldi'. nom. inval., design. typi omissa. Sub Ophrys bertolonii Mor. Ophrys bombyliflora Link. Type: [Italia, Puglia] in der apulischen Küstenebene östlich von Lecce. 11.IV Ophrys bombyliflora Link Ophrys tarentina Gölz & H.R.Reinhard, Ophrys mannarica P.Delforge & C.Delforge, Naturalistes Belges 67(5/6): Sub Ophrys bombyliflora Link Ophrys tarentina Gölz & Reinhard. 590 Journal Europäischer Orchideen 49 (3-4): 2017.

27 Type: Italia, provincia Taranto, apud urbem Grottaglie In herb. P. Delforge sub n 86/3. Ophrys bombyliflora Link Ophrys tenthredinifera Willd. subsp. neglecta (Parl.) E.G.Camus, Ophrys sommieri (Sommier) E.G.Camus ap Cortesi, Ann. Bot. (Roma) 1(5): 360, in adnot Sub Ophrys bombyliflora Link Ophrys tenthredinifera Willd. Bas.: Ophrys bombyliflora tenthredinifera Sommier, Nuovo Giorn. Bot. Ital. N.S. III: , tav. III Type: [Italia, Toscana, Grosseto] Monte Argentario (colle di Faccia alla diga). 21 aprile [FI]. Not listed in IPNI. Syn.: Ophrys bombyliflora tenthredinifera, ASCHERS. & GRAEBN., Syn. Mitteleur. Fl. [Ascherson & Graebner] vol. 3: pro synonyme. Bas.: Ophrys bombyliflora tenthredinifera Sommier, Nuovo Giorn. Bot. Ital. N.S. III: , tav. III Bisher nur in Toscana am Monte Argentario (Sommier). Not listed in IPNI. Ophrys brutia P.Delforge Ophrys dinarica Kranjčev & P.Delforge, Ophrys capracottae Romolini & Soca, GIROS Notiz. 55: 53, Type: Italia. Molise. Isernia. Capracotta. Lamatura, alt m. 33T V Coll. Romolini & Soca. MPU. n RS Ophrys brutia P.Delforge Ophrys exaltata Ten., Ophrys olearum Silletti & Medagli, Silvae: epublished, contrary to Art. H.5.2. Sub Ophrys exaltata Ten. subsp. mateolana (Medagli, D'Emerico, Bianco & Ruggiero) Paulus & Gack Ophrys incubacea Bianca subsp. brutia P. Delforge. Type: [Puglia] Santeramo in Colle (Ba), legit G. N. Silletti, in LEC. Photography of Ophrys exaltata Ten. Not listed in IPNI. Ophrys brutia P.Delforge Ophrys incubacea Bianca, Ophrys incubacea [] nothosubsp. ionica Gennaio, Gargiulo, Medagli & Chetta, GIROS Orch. Spont. Eur. 59(2): sub Ophrys incubacea [] subsp. brutia (P.Delforge) Kreutz Ophrys incubacea subsp. incubacea Bianca. (Author of the species not indicated). Type: Italia, Apulia, Sallentum, circa Callipolim, Lupiensi provincia (N E ) (DD N, E) , Li Foggi, Parco Naturale Regionale Isola di Sant Andrea e Litorale di Punta Pizzo (LE), leg. R. Gennaio, cons. in LEC. Journal Europäischer Orchideen 49 (3-4):

28 Ophrys brutia P.Delforge Ophrys parvimaculata (O.Danesch & E.Danesch) Paulus & Gack, Ophrys buffaloriae Romolini & Soca, GIROS Notiz. 55: 57-58, Type: Italia. Puglia. Taranto. Martina Franca. Buffaloria (Monte Trazzonara), alt. 396 m. 33T IV Coll. Romieg Soca. MPU. n RS A. Ophrys brutia P.Delforge Ophrys passionis Sennen, Ophrys celani O.Danesch & E.Danesch nothosubsp. hawkingi Gennaio, Gargiulo, Medagli & Chetta, GIROS Orch. Spont. Eur. 59(2): sphalm. sub Ophrys incubacea [] subsp. brutia (P.Delforge) Kreutz Ophrys passionis [] subsp. garganica E.Nelson ex H.Baumann & R. Lorenz. (Author of the species not indicated). Type: Italia, Apulia, Sallentum, circa Ausentum, Lupiensi provincia (N E ) (DD N, E) , Serra di Rottacapozza, Parco Naturale Regionale Litorale di Ugento (LE), leg. R. Gennaio, cons. in LEC. Ophrys palenae Soca nothosubsp. hawkingi Gennaio, Gargiulo, Medagli & Chetta. Ophrys brutia P.Delforge Ophrys passionis Sennen subsp. majellensis (Helga Daiss & Herm.Daiss) Romolini & Soca, Ophrys palenae Soca, J. Eur. Orch. 46(3/4): contrary to Art. H.5.2. Type: Italia, Abruzzo, Chieti, Palena, San Cataldo, alt. 900 m. 33T Coll. Romieg Soca. MPU n RS Ophrys brutia P.Delforge Ophrys romolinii Soca, Ophrys vernacchiae Soca, J. Eur. Orch. 46(3/4): Type: Italia, Abruzzo, Chieti, Gamberale, Colle Vernacchia, alt m. 33T Coll. Romieg Soca. MPU n RS Ophrys brutia P.Delforge Ophrys tenthredinifera Willd. subsp. neglecta (Parl.) E.G.Camus, Ophrys manfredoniae O.Danesch & E.Danesch nothosubsp. delilloi D Alonzo & Perilli, GIROS Orch. Spont. Eur. 59(2): sphalm. sub Ophrys incubacea [] subsp. brutia (P.Delforge) Kreutz Ophrys tenthredinifera [] subsp. neglecta (Parl.) E.G.Camus. (Author of the species not indicated). Type: [Italia] località Bosco Difesa Grande (Gravina in Puglia, Bari), m 397 s.l.m., UTMWGS 84 33T E N ; leg. F. D Alonzo & M. Perilli, cons. in BI (Herbarium Horti Botanici Barensis), spec Ophrys grampinii Cortesi nothosubsp. delilloi D Alonzo & Perilli. 592 Journal Europäischer Orchideen 49 (3-4): 2017.

29 Ophrys calliantha Bartolo & Pulv. Ophrys incubacea Bianca, Ophrys carnazzae Galesi, J. Eur. Orch. 31(2): Type: [Italia, Sicilia, Vittoria (RG)] Nei pressi di C.se Carnazza (VA ), all'interno di un carrubeto abbandonato, 50m, 25.IV.I992, R. Galesi (CAT). Ophrys calliantha Bartolo & Pulv. Ophrys lacaitae Lojac., Ophrys pantalicae Bartolo & Pulv., J. Eur. Orch. 30(3): Type: [Italia] Sicilia, Monti Iblei, Pantalica (SR), , lego Bartolo & Brullo, (CAT holotypus e isotypi). Ophrys calliantha Bartolo & Pulv. Ophrys oxyrrhynchos Tod., Ophrys ferliana Bartolo & Pulv., J. Eur. Orch. 30(3): Type: [Italia] Sicilia, Monti Iblei, Ferla (SR), , leg. Bartolo & Brullo, (CAT holotypus e isotypi). Ophrys candica (E.Nelson ex Soó) H.Baumann & Künkele Ophrys parvimaculata (O.Danesch & E.Danesch) Paulus & Gack, Ophrys sivana nothosubsp. renatafontae Gennaio, Gargiulo, Medagli & Chetta, GIROS Orch. Spont. Eur. 59(2): sub Ophrys candica (E. Nelson ex Soó) H. Baumann & Künkele Ophrys holosericea [] subsp. parvimaculata (O.Danesch & E.Danesch) O.Danesch & E.Danesch (Author of the species not indicated). Type: Italia, Apulia, Sallentum, Cesinae circa Vernulas, Lupiensi provincia; coordinate geografiche in DD N, E Riserva Naturale Statale Le Cesine, Vernole (LE), leg. R. Gennaio, cons. in LEC. Photograph = Ophrys candica! Ophrys candica (E.Nelson ex Soó) H.Baumann & Künkele Ophrys tenthredinifera Willd. subsp. neglecta (Parl.) E.G.Camus, Ophrys tardans O.Danesch & E.Danesch, Orchid. Eur. Ophrys-Hybriden: 225, [pro spec. hybr.]. Sub Ophrys candica (Nelson) O. & E.Danesch Ophrys tenthredinifera Willd. Type: Italien (Apulien), bei Lecce, leg. O. et E. Danesch, (IB). Syn.: Ophrys maremmae O.Danesch & E.Danesch nothosubsp. tardans (O.Danesch & E.Danesch) Del Prete, Webbia 37(2): pro synonyme. sub Ophrys holoserica (N.L. Burm.) Greuter subsp. candica (E.Nelson ex Soó) Renz et Taubenh. Ophrys tenthredinifera Willd. Ophrys chestermanii (J.J.Wood) Gölz & H.R.Reinhard Ophrys conradiae Melki & Deschâtres, Ophrys vicina Duffort nothosubsp. corriasiana H.Baumann, Giotta, Künkele, R.Lorenz & Piccitto, J. Eur. Orch. 27(2): Sub Ophrys holoserica (Burm.f.) Greuter subsp. chestermanii J.J.Wood Ophrys scolopax Cav. subsp. conradiae (Melki & Deschâtres) H.Baumann, Journal Europäischer Orchideen 49 (3-4):

30 Giotta, Lorenz, Künkele & Piccitto. Type: [Italia] Sardinia, Iglesiente, Domusnovas, Sa Duchessa, 400 m, UTM MJ , leg. B. & H. Baumann (STU). Ophrys chestermanii (J.J.Wood) Gölz & H.R.Reinhard Ophrys exaltata Ten. subsp. morisii (Martelli) Del Prete, Ophrys maladroxiensis Scrugli, Todde & Cogoni nothosubsp. daissiorum H.Baumann, Giotta, Künkele, R.Lorenz & Piccitto, J. Eur. Orch. 27(2): Sub Ophrys holoserica (Burm.f.) Greuter subsp. chestermanii J.J.Wood Ophrys morisii Martelli. Type: [Italia] Sardinia, Iglesiente, Grotta di San Giovanni, 220 m, MJ , leg. B. & H. Baumann (STU). Syn.: Ophrys daissiorum (H.Baumann, Giotta, Künkele, Lorenz & Piccitto) P.Delforge, Naturalistes Belges 78(3): sub Ophrys chestermanii (J.J.Wood) Gölz & H.R.Reinhard Ophrys morisii (Martelli) Soó. Ophrys chestermanii (J.J.Wood) Gölz & H.R.Reinhard Ophrys incubacea Bianca, Ophrys montis-lorae Giotta & Piccitto, J. Eur. Orch. 44(4): Type: Italia, Sardegna, Villasalto (CA), "Serra Madau" (UTM: NJ 37.90), macchia-foresta, m 94 s.l.m., , leg. M. Piccitto & C. Giotta (FI). Ophrys chestermanii (J.J.Wood) Gölz & H.R.Reinhard Ophrys tenthredinifera Willd. subsp. neglecta (Parl.) E.G.Camus, Ophrys normanii J.J.Wood, Orchid Rev. 91(1082): contrary to Art. H.5.2. Sub Ophrys holoserica (Burm.f.) Greuter subsp. chestermanii J.J.Wood Ophrys tenthredinifera Willd., Type: Italy. Sardinia. Cagliari Province. near Fluminimaggiore. c.80m T. Norman s n (K. holotypus. Kew Spirit Colln no 46312). Syn.: Ophrys maremmae O.Danesch & E.Danesch nothosubsp. woodii Corrias, Inform. Bot. Ital. 15(2/3): 178. (1983) nomen illegitime. [Italia] Sardegna. Fluminimaggiore: loc. «Candiazzu». (Il nome qui adottato per la nuova entità si basa (art. 36) sulla diagnosi pubblicata e sull'holotypus designato da WOOD (1983). Syn.: Ophrys maremmae O.Danesch & E.Danesch nothosubsp. normanii (J.J.Wood) H.Baumann & Künkele, Mitteilungsbl. Arbeitskreis Heimische Orchid. Baden-Württemberg 18(3): Ophrys cinnabarina Romolini & Soca Ophrys lacaitae Lojac., Ophrys montiscelanensis Perilli, GIROS Orch. Spont. Eur. 58(2): Type: [Italia, Puglia] località Monte Celano (San Marco in Lamis, Foggia), m 746 s.l.m., 33T E N ; M. Perilli, cons. in BI 594 Journal Europäischer Orchideen 49 (3-4): 2017.

31 (Herbarium Horti Botanici Barensis), spec. n Ophrys classica Devillers-Tersch. & Devillers Ophrys crabronifera Mauri, Ophrys camusii Cortesi, Ann. Bot. (Roma) 5(3): Sub Ophrys aranifera Huds. Ophrys exaltata Ten. Type: [Italia, Lazio, Roma] Habitat in arenosis maritimis alla pineta di Maccarese inter parentes. (Aprili ). Not find in RO. Not listed in IPNI. Ophrys classica Devillers-Tersch. & Devillers Ophrys exaltata Ten. subsp. montis-leonis (O.Danesch & E.Danesch) Soca, Ophrys dionysii P.Delforge, Naturalistes Belges 70(3): 90, contrary to Art. H.5.2. Sub Ophrys tyrrhena Gölz & Reinhard Ophrys sphegodes Miller subsp. sphegodes. Type: Italia, Toscana, provincia Grosseto, apud Manciano, Leg. Denis Delforge in herb. Pierre Delforge sub n Ophrys classica Devillers-Tersch. & Devillers Ophrys incubacea Bianca, Ophrys montalciniae nothosubsp. montalciniae Gennaio, Gargiulo, Medagli & Chetta, GIROS Orch. Spont. Eur. 59(2): sub Ophrys incubacea subsp. incubacea Bianca Ophrys sphegodes [] subsp. classica (Devillers-Tersch. & Devillers) Kreutz (Author of the species not indicated). Type: Italia, Apulia, Sallentum, circa Ausentum, Lupiensi provincia (DD N E) , Serra di Rottacapozza, Parco Naturale Regionale Litorale di Ugento (LE), leg. R. Gennaio, cons. in LEC (Herbarium Universitatis Lupiensis) n Ophrys classica Devillers-Tersch. & Devillers Ophrys lojaconoi P.Delforge, Ophrys sophiae nothosubsp. sophiae D Alonzo & Perilli, GIROS Orch. Spont. Eur. 60(1): sub O. iricolor subsp. lojaconoi (P. Delforge) Kreutz O. sphegodes subsp. classica (Devillers-Tersch. & Devillers) Kreutz. Italia, Apulia, in Gargano ad Matinatam, Foveanam provinciam, m 247 s.l.m., UTMWGS 84 33T E N ; leg. F. D Alonzo & M. Perilli, cons. in BI (Herbarium Horti Botanici Barensis), spec , data Ophrys classica Devillers-Tersch. & Devillers Ophrys tenthredinifera Willd. subsp. neglecta (Parl.) E.G.Camus, Ophrys etrusca (Sommier) Aschers. & Graebn., Syn. Mitteleur. Fl. [Ascherson & Graebner] vol. 3: Sub Ophrys aranifera Ophrys tenthredinifera Willd. Journal Europäischer Orchideen 49 (3-4):

32 Bas.: Ophrys tenthredinifera aranifera Sommier, Nuovo Giorn. Bot. Ital. (Bull. Soc. bot. It.) 24: Type: [Italia, Toscana, Grosseto] Nel bosco di Quercus suber, vicino alla strada maestra che conduce da Orbettello a Burano. 14 aprile [FI]. Not listed in IPNI. Ophrys conradiae Melki & Deschâtres, Orchidophile (Asnières) 24(107): , Sub Ophrys holosericea Ophrys scolopax (Ophrys vicina Duffort). GRASSO, Mitteilungsbl. Arbeitskreis Heimische Orchid. Baden-Württemberg 1988: Abb. 5, photography of Ophrys conradiae. sine descriptio. Type: [Italia, Sardegna] bei Aqua sa Murta, Donori, 200 m ü.d.m Ophrys corsica Soleirol ex G.Foelsche & W.Foelsche Ophrys exaltata Ten. subsp. morisii (Martelli) Del Prete, Ophrys damilanoi Romolini & Soca nothosubsp. damilanoi, Caesiana 21(45): 17, Type: Italia. Sardegna (OG). Ulassai. Monte Matzeu, loc. Escorga, 669 m s.l.m., 32S IV Coll. R. Romolini et R. Soca. Leg. FI. n RS Ophrys corsica Soleirol ex G.Foelsche & W.Foelsche Ophrys fusca s.l., Ophrys cesinensis Medagli, d'emerico & Ruggiero, Orchidophile (Asnières) 20(85): contrary to Art. H.5.2. Sub Ophrys lutea Cav. subsp. melena Renz Ophrys fusca Link. Type: Italia, Apulia, «Le Cesine» (LE), m 3 ca., 5. 1V.1988, P. Medagli et L. Ruggiero. In herb. Gruppo Naturalisti Salentini di Lecce sub 352 fig. 3. Ophrys corsica Soleirol ex G.Foelsche & W.Foelsche Ophrys incubacea Bianca, Ophrys piscinica P.Delforge & C.Delforge nothosubsp. mattinatellae R.Kohlmüller, Orchidee (Hamburg) 44(2): Sub Ophrys incubacea Bianca Ophrys lutea subsp. minor (Tod.) O. & E.Danesch. Type: Italia, Monte Gargano (Puglia), 6,4 km NO Mattinata, , leg. R. Kohlmüller, Botanische Staatssammlung München (M). Ophrys corsica Soleirol ex G.Foelsche & W.Foelsche Ophrys lutea Cav. [ Soleroil ], Ophrys sulphurea Gennaio & Medagli, GIROS Orch. Spont. Eur. 58(1): Sub Ophrys lutea Cav. Ophrys corsica Soleroil (sic) ex G.Foelsche & W.Foelsche. Type: Italia. Puglia (LE). Otranto, alt. 52 m s.l.m., lat. N 40, lon. 596 Journal Europäischer Orchideen 49 (3-4): 2017.

33 Fig. 9: Ophrys corsica Ophrys flammeola, Santo Stefano Quisquina, Sicilia (AG), Fig. 10: Ophrys corsica Ophrys sicula, Monte Sant Angelo, Puglia (FG), Fig. 11: Ophrys corsica Ophrys tenthredinifera subsp. neglecta, Monte Sant Angelo, Puglia (FG), Journal Europäischer Orchideen 49 (3-4):

34 Fig. 12: Ophrys corsica Ophrys exaltata subsp. morisii, Ulassai, Sardegna (OG), Fig. 13: Ophrys lojaconoi Ophrys passionis, San Giovanni Rotondo, Puglia (FG), Fig. 14: Ophrys dinarica Ophrys pinguis, Ateleta, Abruzzo (AQ), Fig. 15: Ophrys dinarica Ophrys romolinii, Borgorose, Lazio (RI), Journal Europäischer Orchideen 49 (3-4): 2017.

35 E 18, prati aridi presso la cava di bauxite nel Parco Naturale Regionale Costa Otranto - Santa Maria di Leuca e Bosco di Tricase, Otranto (LE), leg. R. Gennaio, 6.IV.2014, cons. in LEC - Herbarium Universitatis Lupiensis, sub n , 9.IV Ophrys corsica Soleirol ex G.Foelsche & W.Foelsche Ophrys romolinii Soca, Ophrys anxatina R.Congedo, GIROS Notiz. 36: contrary to Art. H.5.2. Sub Ophrys bertolonii Moretti Ophrys lutea Cav. subsp. minor []. Author of the subspecies not indicated. Type: [Italia] loc. Torre del Pizzo, lat ,26 N long ,98 E (UTM BK43), Gallipoli, Lecce, facente parte del SIC (coordinate geografiche IT E1759 N3958), esposizione SSO. altitudine 7 m s.l.m. Data13/04/2006, legit R.Congedo (LEC), campione conservato presso l Erbario del Di.S.Te.B.A. dell Università del Salento. Not listed in IPNI. Ophrys corsica Soleirol ex G.Foelsche & W.Foelsche Ophrys sicula Tineo, Ophrys calchasii Romolini & Soca, GIROS Notiz. 55: 54, Type: Italia. Puglia. Foggia. Monte Sant Angelo. Via Pulsano, alt. 734 m. 33T IV Coll. Romieg Soca. MPU. n RS Ophrys corsica Soleirol ex G.Foelsche & W.Foelsche Ophrys tarentina Gölz & H.R.Reinhard, Ophrys sansimonensis Soca, Monde Pl. 90(452): Sub Ophrys sicula Tineo Ophrys tarentina Gölz & H.R.Reinhard. Type: Italia, Puglia, Taranto, apud San Simone (Crispiano). 13.IV MPU n RS Ophrys corsica Soleirol ex G.Foelsche & W.Foelsche Ophrys tenthredinifera Willd. subsp. neglecta (Parl.) E.G.Camus, Ophrys angelii Soca nothosubsp. angelii, Caesiana 19(43): Type: Italia. Puglia. Foggia. Monte Sant Angelo. Gli Angeli. Alt. 741 m s.l.m. 33T V Coll. Romieg Soca. Cons. in herb. MPU. sub n RS Ophrys crabronifera Mauri Ophrys exaltata Ten. subsp. montis-leonis (O.Danesch & E.Danesch) Soca, Ophrys terschureniana P.Delforge, Naturalistes Belges 70(3): 90, Sub Ophrys tyrrhena Gölz & Reinhard Ophrys crabronifera Mauri. Type: Italia, Toscana, provincia Livorno, Marina di Castagneto-Donoratico, In herb. Pierre Delforge sub n Journal Europäischer Orchideen 49 (3-4):

36 Ophrys crabronifera Mauri Ophrys incubacea Bianca, Ophrys argentariensis Ruppert, Repert. Spec. Nov. Regni Veg. 31: Sub Ophrys atrata Lindl. Ophrys exaltata Ten. Type: [Italia, Toscana, Grosseto] über Porto S. Stefano. 9 April Unknown herbarium. Ophrys crabronifera Mauri Ophrys passionis Sennen, Sub Ophrys exaltata Ten. Ophrys garganica E.Nelson ex O.Danesch & E.Danesch. SCHRENK, Amer. Orchid Soc. Bull. 1981: 131, 133. nom. nud. sine descriptio. Monte Argentario, Italy. 4/26/82. Ophrys crabronifera Mauri Ophrys promontorii O.Danesch & E.Danesch, Sub Ophrys crabronifera Ophrys promontorii. MORALDO & al., Atti Accad. Naz. Lincei, Rendiconti Cl. Sci. Fis. Mat. Nat. 387(264): ; ROSSI & al., Acad. Nazl. Lincei Roma 387(264): sine descriptio. Italia, Lazio meridionale. Ophrys crabronifera Mauri Ophrys romolinii Soca, Ophrys liberationis Antonetti, Contorni & Bertolini, GIROS Orch. Spont. Eur. 59(1): contrary to Art. H.5.2. Sub Ophrys bertolonii subsp. bertolonii Moretti Ophrys crabronifera subsp. crabronifera Mauri. Type: Italia, Etruria (Grossetum), prope Alborensem (N ; E ). cons. in PI (Herbarium Horti Botanici Pisani) Nuove Acquisizioni, N Ophrys liberationis Antonetti, Contorni & Bertolini / 1 (holotypus). Ophrys crabronifera Mauri Ophrys tenthredinifera Willd. subsp. neglecta (Parl.) E.G.Camus, Ophrys tuscanica nsubsp. tuscanica H.Baumann & Künkele, Mitteilungsbl. Arbeitskreis Heimische Orchid. Baden-Württemberg 18(3): 450, Sub Ophrys crabronifera Mauri Ophrys tenthredinifera Willd. subsp. tenthredinifera. Type: Italia [Toscana] Grosseto: Mte Argentario, Orbetello, 20 m , leg. B. & H. Baumann (STU). Ophrys dinarica Kranjčev & P.Delforge Ophrys incubacea Bianca, Ophrys recchiae Soca, J. Eur. Orch. 46(3/4): Type: Italia, Abruzzo, L'Aquila, Collepietro, Monte Ospedalera, alt. 651 m, 33T Coll. Romieg Soca. MPU n RS Ophrys dinarica Kranjčev & P.Delforge Ophrys molisana P.Delforge, Ophrys milioniae Soca, J. Eur. Orch. 49(2): Italia. Abruzzo. L Aquila. Ortona dei Marsi. Pratolungo. 33T Journal Europäischer Orchideen 49 (3-4): 2017.

37 Alt. 941 m. 26.V Coll. R. Soca. Cons. in herb. FI. sub n RS Ophrys dinarica Kranjčev & P.Delforge Ophrys passionis Sennen subsp. majellensis (Helga Daiss & Herm.Daiss) Romolini & Soca, Ophrys brunamontei Soca, J. Eur. Orch. 46(3/4): contrary to Art. H.5.2. Type: Italia, Abruzzo, Chieti, Palena, San Cataldo, alt. 900 m, 33T Coll. Romieg Soca. MPU n RS Ophrys dinarica Kranjčev & P.Delforge Ophrys pinguis Romolini & Soca, Ophrys impresciae Soca, J. Eur. Orch. 46(3/4): Type: Italia, Abruzzo, L'Aquila, Capistrello, Selvetta, alt. 721 m, 33T Coll. Romieg Soca. MPU n RS Ophrys dinarica Kranjčev & P.Delforge Ophrys promontorii O.Danesch & E.Danesch, Ophrys aquilana H.Baumann & Künkele, Mitteilungsbl. Arbeitskreis Heimische Orchid. Baden-Württemberg 18(3): Sub Ophrys holosericea (N.L.Burm.) W. Greuter subsp. holosericea Ophrys promontorii O. & E. Danesch. Type: Italia [Abruzzo] Aquila: Villetta Barrea, 1100 m , leg. B. & H. Baumann (STU). Ophrys dinarica Kranjčev & P.Delforge Ophrys romolinii Soca, Ophrys piconei Soca, J. Eur. Orch. 46(3/4): Type: Italia, Abruzzo, Chieti, Palena, Girone, alt. 906 m, 33T Coll. Romieg Soca. MPU n RS Ophrys dinarica Kranjčev & P.Delforge Ophrys tetraloniae W.P.Teschner Ophrys capistrelloi Soca, J. Eur. Orch. 46(3/4): Type: Italia, Abruzzo, L'Aquila, Capistrello, Selvetta, alt. 754 m, 33T Coll. Romieg Soca. MPU n RS Ophrys discors Bianca Ophrys exaltata Ten., Ophrys mangiagessoi Cataldo & S.A.Giardina, J. Eur. Orch. 43(4): 792 ( ; figs.) Sub Ophrys biancae (Tod.) Macch. Ophrys exaltata Ten. Type: Sicilia, Scicli (RG), Monti Iblei presso l'area boscata di Mangiagesso, UTMED S E N, 355 m s.l.m., , leg. D. Cataldo & S.A. Giardina (PAL). Ophrys discors Bianca Ophrys incubacea Bianca, Ophrys martorinae Galesi, J. Eur. Orch. 31(2): Type: [Italia, Sicilia, Vittoria (RG)] C.da Martorina (VA ). In una gariga Journal Europäischer Orchideen 49 (3-4):

38 a Thymus capitatus, 130m, 06.IV.l992, R. Galesi (CAT). Ophrys discors Bianca Ophrys lacaitae Lojac., Ophrys bibbiensis Soca, Monde Pl. 92(460): Sub Ophrys biancae (Tod.) Macch. Ophrys lacaitae Lojac. Type: Italia, Sicilia (Siracusa), in septemtrionali Canicattini Bagni, loco dicto Villa Bibbia. 20.IV MPU n RS Ophrys discors Bianca Ophrys lunulata Parl., Ophrys toussaintiana P.Delforge, Naturalistes Belges 69(2): Sub Ophrys biancae (Tod.) Macchiati Ophrys lunulata Parlat. Type: Italia, Sicilia, provincia Siracusa, Necropoli di Pantalia (sic), In herb. Pierre Delforge sub n 87/5. Ophrys discors Bianca Ophrys oxyrrhynchos Tod. subsp. oxyrrhynchos, Ophrys necropolitana H.Baumann & Künkele, Mitteilungsbl. Arbeitskreis Heimische Orchid. Baden-Württemberg 18(3): 452, Type: Italia, Sicilia: Necropoli di Pantalia, 450 m , leg. H. Baumann M. Kalteisen (STU). Ophrys discors Bianca Ophrys tenthredinifera Willd. subsp. grandiflora (Ten.) Kreutz, Sub Ophrys discors Bianca Ophrys tenthredinifera Willd. PETEREK & VOLLMAR, Mitteilungsbl. Arbeitskreis Heimische Orchid. Baden- Württemberg 1992: 111, 113, 123. sine descriptio. [Italia, Sicilia] Nähe von Ferla, zw. Ferla u. Sortino, Nähe Necropoli di Pantalica, Weide, WB 0109, Ophrys exaltata Ten. Ophrys fusca s.l., Ophrys laureantis Galesi, J. Eur. Orch. 31(4): Sub Ophrys exaltata Tenore Ophrys fusca Link. Type: [Italia, Sicilia, Niscemi] C.da Banco (VB: 40.69), in un uliveto abbandonato, 280m, 15.III.1995, R. Galesi (CAT). Ophrys exaltata Ten. Ophrys incubacea Bianca, Ophrys nettii Silletti & Medagli, Silvae: epublished, contrary to Art. H.5.2. Sub Ophrys exaltata Ten. subsp. mateolana (Medagli, D'Emerico, Bianco & Ruggiero) Paulus & Gack Ophrys incubacea Bianca subsp. incubacea. Type: [Puglia] Santeramo in Colle (Ba), legit G. N. Silletti, in LEC. Photography of Ophrys exaltata Ten. Not listed in IPNI. Ophrys exaltata Ten. Ophrys incubacea Bianca Ophrys romolinii Soca, Ophrys santeramana Silletti & Medagli, Silvae: epublished, 602 Journal Europäischer Orchideen 49 (3-4): 2017.

39 contrary to Art. H.5.2. Sub Ophrys exaltata Ten. subsp. mateolana (Medagli, D'Emerico, Bianco & Ruggiero) Paulus & Gack Ophrys lyrata H.Fleischm. Type: [Puglia] Santeramo in Colle (Ba) località Pantarosa, legit G. N. Silletti in LEC. Photography of Ophrys exaltata Ten. Ophrys romolinii Soca. Not listed in IPNI. Ophrys exaltata Ten. Ophrys lunulata Parl., Ophrys goelziana H.Baumann & Künkele, Mitteilungsbl. Arbeitskreis Heimische Orchid. Baden-Württemberg 16(4): Type: [Italia] Insula Sicilia, ad pagum Acate , leg. H. Baumann, in Herbario Staatliches Museum für Naturkunde in Stuttgart (STU) conservatur. Syn.: Ophrys nemorosa Gölz, Orchidee (Hamburg) 22(4): 165, nom. inval., design. typi omissa. sub Ophrys arachnitiformis Gren. et Phil. Ophrys lunulata Parl. [Italia, Sicilia] im Hinterland südlich Palermo in einem lichten Hain von Ginsterbüschen und Obstbäumen. 4.IV Ophrys exaltata Ten. Ophrys oxyrrhynchos Tod., Ophrys reinhardiana H.Baumann & Künkele, Mitteilungsbl. Arbeitskreis Heimische Orchid. Baden-Württemberg 16(4): Type: [Italia] Insula Sicilia, 8 km E Gela , leg. H. Baumann, in Herbario Staatliches Museum für Naturkunde in Stuttgart (STU) conservatur. Ophrys exaltata Ten. Ophrys passionis Sennen, Ophrys giardinae Cristaudo, Galesi, R.Lorenz & Zelesny, J. Eur. Orch. 42(1): Sub Ophrys exaltata Ten. Ophrys garganica (E.Nelson) O.Danesch & E.Danesch. Type: [Sicilia] Giardino del Fico, Niscemi (CL), sandiger Boden mit Sandsteinagglomeraten in ehemaligem Eucaliptetum, Garrigue mit Corydothymus capitatus, 290 m ü.d.m., UTMEDl950 33S VB , , R. Galesi (CAT). Syn.: Ophrys durae Silletti & Medagli, Silvae: epublished, contrary to Art. H.5.2. [Puglia] Santeramo in Colle (Ba) località Pantarosa, legit G. N. Silletti in LEC. Sub Ophrys exaltata Ten. subsp. mateolana (Medagli, D'Emerico, Bianco & Ruggiero) Paulus & Gack Ophrys passionis Sennen ex Devillers-Tersch. et Devillers subsp. garganica O. Danesch et E. Danesch. Not listed in IPNI. Ophrys exaltata Ten. Ophrys pollinensis E.Nelson ex Devillers-Tersch. & Devillers, Ophrys breuerorum P.Delforge, Naturalistes Belges 83(Orchid. 15): Sub Ophrys cilentana Devillers-Tersch. & Devillers Ophrys pollinensis Journal Europäischer Orchideen 49 (3-4):

40 E.Nelson ex Devillers-Tersch. & Devillers. Type: Italia, Campania, provincia Salernum, Cilento, circa Pisciotta (UTM: 33TWE2041), alto s.m. 280 m, l.iv In herb. Pierre Delforge sub n Ophrys exaltata Ten. Ophrys romolinii Soca, Ophrys gambettae Silletti & Medagli, Silvae: epublished, contrary to Art. H.5.2. Sub Ophrys bertolonii Moretti subsp. bertolonii Ophrys exaltata Ten. subsp. mateolana (Medagli, D'Emerico, Bianco & Ruggiero) Paulus & Gack. Type: [Puglia] Santeramo in Colle (Ba), legit G. N. Silletti, in LEC. Not listed in IPNI. Ophrys exaltata Ten. Ophrys tarentina Gölz & H.R.Reinhard, Ophrys margheritae Silletti & Medagli, Silvae: epublished, contrary to Art. H.5.2. Sub Ophrys exaltata Ten. subsp. mateolana (Medagli, D'Emerico, Bianco & Ruggiero) Paulus & Gack Ophrys tarentina Gölz & H.R.Reinhard. Type: [Puglia] Santeramo in Colle (Ba), loc. Lama Lunga, , legit G. N. Silletti & Angelo Margherita in LEC. Photography of Ophrys exaltata Ten. Not listed in IPNI. Ophrys exaltata Ten. Ophrys tenthredinifera Willd. subsp. grandiflora (Ten.) Kreutz, Ophrys sanconoensis M.Peterek & J.Vollmar, Mitteilungsbl. Arbeitskreis Heimische Orchid. Baden-Württemberg 24(1): Sub Ophrys exaltata Ten. Ophrys tenthredinifera Willd. Type: Italia, Sicilia: inter Gela et Piazza Armerina in pratis apricis prope San Cono , leg. M. Peterek & J. Vollmar (STU). Ophrys exaltata Ten. Ophrys tenthredinifera Willd. subsp. neglecta (Parl.) E.G.Camus, Ophrys vanderspekiae P.Delforge, Naturalistes Belges 82(Orchid. 14): Sub Ophrys cilentana Devillers & Devillers-Terschuren Ophrys tenthredinifera Willdenow. Type: Italia, Campania, provincia Salernum, Cilento, circa Forla (UTM: 33TWE3038), alt. s.m. 90 m, 5.lV In herb. Pierre Delforge sub n Ophrys exaltata Ten. subsp. archipelagi (Gölz & H.R.Reinhard) Del Prete Ophrys fusca s.l. (lojaconoi?), Ophrys turrium R.Kohlmüller, Orchidee (Hamburg) 44(2): contrary to Art. H.5.2. Sub Ophrys archipelagi Gölz & Reinhard Ophrys fusca Link subsp. fusca. 604 Journal Europäischer Orchideen 49 (3-4): 2017.

41 Type: Italia, Monte Gargano (Puglia), 2 km NNO Mattinata, , leg. R. Kohlmüller, Botanische Staatssammlung München (M). Ophrys exaltata Ten. subsp. archipelagi (Gölz & H.R.Reinhard) Del Prete Ophrys incubacea Bianca, Ophrys geniezii Soca, Monde Pl. 91(458): Sub Ophrys archipelagi Gölz & H.R.Reinhard Ophrys incubacea Bianca. Type: Italia, Puglia (Foggia), N.-E. Mattinata, loco dicto Mattinatella, alt. 130 m. 6.IV MPU n RS Ophrys exaltata Ten. subsp. archipelagi (Gölz & H.R.Reinhard) Del Prete Ophrys lutea Cav., Sub Ophrys arachnitiformis Gren. et Phil. Ophrys lutea Cav. LORENZ & GEMBARDT, Mitteilungsbl. Arbeitskreis Heimische Orchid. Baden- Württemberg 1987: 505, 541. sine descriptio. [Italia, Puglia, Gargano] 6.6 km NNE Mattinata, UTM: , 580 m. W. Klopsch, W. Malkmus & H. Ort Ophrys exaltata Ten. subsp. archipelagi (Gölz & H.R.Reinhard) Del Prete Ophrys oestrifera M.-Bieb. subsp. montis-gargani Van de Vijver & W.Looken, Ophrys lewinii Soca, Monde Pl. 91(458): Sub Ophrys archipelagi Gölz & H.R.Reinhard Ophrys oestrifera Steven in M.Bieb. Type: Italia, Puglia (Foggia), Peschici, alt. 160 m. 11.IV MPU n RS Ophrys exaltata Ten. subsp. archipelagi (Gölz & H.R.Reinhard) Del Prete Ophrys parvimaculata (O.Danesch & E.Danesch) Paulus & Gack, Sub Ophrys archipelagi Gölz & H.R.Reinhard Ophrys parvimaculata (O.Danesch & E.Danesch) Paulus & Gack. Ophrys ingaranensis Soca, Monde Pl. 92(460): Type: Italia, Puglia (Foggia), Monte Gargano: inter Apricena et Sanicandro, Ingarano, in septemtrionale latere Mt Trattunale, alt. 280 m. 12.IV MPU n RS Ophrys exaltata Ten. subsp. archipelagi (Gölz & H.R.ReinhardDel Prete Ophrys passionis Sennen, Ophrys manacorensis H.Baumann & Künkele, Mitteilungsbl. Arbeitskreis Heimische Orchid. Baden-Württemberg 18(3): 410, Sub Ophrys arachnitiformis Gren. & Philippe Ophrys garganica E.Nelson ex O. & E. Danesch. Type: Italia, Puglia: Mte. Gargano, Manacore, 20 m , leg. B. & H. Baumann (STU). Journal Europäischer Orchideen 49 (3-4):

42 Ophrys exaltata Ten. subsp. archipelagi (Gölz & H.R.Reinhard) Del Prete Ophrys romolinii Soca, Sub Ophrys arachnitiformis Gren. et Phil. Ophrys bertolonii Moretti. LORENZ & GEMBARDT, Mitteilungsbl. Arbeitskreis Heimische Orchid. Baden-Württemberg 1987: 505, 541. sine descriptio. [Italia, Puglia, Gargano] sans locus, R. Gumprecht Ophrys exaltata Ten. subsp. archipelagi (Gölz & H.R.Reinhard) Del Prete Ophrys tenthredinifera Willd. subsp. neglecta (Parl.) E.G.Camus, Ophrys devillersiana P.Delforge, Naturalistes Belges 69(2): Sub Ophrys archipelagi Gölz et Reinhard Ophrys tenthredinifera Willd. Type: Italia, Puglia, provincia Foggia, super Montem Sacro, alt. 790 m In herb. Pierre Delforge sub n 86/4. Ophrys exaltata Ten. subsp. montis-leonis (O.Danesch & E.Danesch) Soca Ophrys tarquinia P.Delforge, Ophrys carduccii Romolini & Soca, GIROS Notiz. 38: contrary to Art. H.5.2. Type: Italia, Toscana, Castagneto Carducci (Livorno Provincia). alt. 203 m. (UTM 32T PN37). 22.IV FI Romolini 42/04. Ophrys exaltata Ten. subsp. montis-leonis (O.Danesch & E.Danesch) Soca Ophrys tenthredinifera Willd. subsp. neglecta (Parl.) E.G.Camus, Ophrys gillesii P.Delforge, Naturalistes Belges 70(3): 89, Sub Ophrys tyrrhena Gölz & Reinhard Ophrys tenthredinifera Willd. Type: Italia, Toscana, provincia Grosseto, apud Manciano, Leg. Gilles Delforge in herb. Pierre Delforge sub n Ophrys exaltata Ten. subsp. montis-leonis (O.Danesch & E.Danesch) Soca, Monde Pl. 97(475): , Ophrys montis-leonis O.Danesch & E.Danesch, Orchid. Eur. Ophrys- Hybriden: 225, contrary to Art. H.5.2. Sub Ophrys arachnitiformis Gren. et Phil. Ophrys fuciflora (Crantz) Moench subsp. fuciflora. Type: Italien (Toscana), M. Leoni, leg. O. et E. Danesch, (IB). Ophrys exaltata Ten. subsp. morisii (Martelli) Del Prete Ophrys incubacea Bianca, Sub Ophrys incubacea Ophrys morisii. GÖLZ & REINHARD, Mitteilungsbl. Arbeitskreis Heimische Orchid. Baden-Württemberg 1988: 127, 135. sine descriptio. Italia, Sardegna, Nuoro (SA): Siniscola, Capo Comino, Ophrys exaltata Ten. subsp. morisii (Martelli) Del Prete Ophrys iricolor Desf. subsp. eleonorae (Devillers-Tersch. & Devillers) Paulus & Gack, Ophrys pardui Giotta & Piccitto nothosubsp. pardui, J. Eur. Orch. 43(4): Journal Europäischer Orchideen 49 (3-4): 2017.

43 ( ; figs.) Sub Ophrys iricolor subsp. maxima (A.Terracciano) Paulus & Gack Ophrys morisii (Martelli) Soó. Type: Italia, Sardegna, Ulassai (OG), "Escorca" (UTM: NK 40.36), scarpata erbosa, m. 680 s.l.m., , M. Piccitto & C. Giotta (FI). Ophrys exaltata Ten. subsp. morisii (Martelli) Del Prete Ophrys panattensis Scrugli, Cogoni et Pessei, Ophrys pesseiae P.Delforge, Naturalistes Belges 78(3): Sub Ophrys morisii (Martelli) Soó Ophrys panattensis Scrugli, Cogoni & Pessei. Type: Italia, Sardinia, provincia Nuoro, apud Baunei, alt. s.m. 450 m, 13.IV.l996. In herb. Pierre Delforge sub n Ophrys exaltata Ten. subsp. morisii (Martelli) Del Prete Ophrys speculum Link, Ophrys barbaricina M.Allard & M.P.Grasso, Orchidophile (Asnières) 35(160): 59 (-62; figs.) Sub Ophrys ciliata Biv.Bern. Ophrys morisii (Martelli) Soó. Type: Sardinia centralis, loco dicto "Montixeddu" 5km NO Villanovatulo (Nu), Legit M. Allard. M.P.Grasso n.1 (FI). Ophrys exaltata Ten. subsp. morisii (Martelli) Del Prete Ophrys tenthredinifera Willd. subsp. neglecta (Parl.) E.G.Camus, Ophrys laconensis Scrugli & M.P.Grasso, Webbia 37(2): , Sub Ophrys arachnitiformis auct. fl. sard. Ophrys tenthredinifera Willd. Type: «Sardegna, Nuoro, in agro circa pagum Laconi, in loco dicto Malifattu, m 760 ca. 9.V.1981, Grasso M. P. et Scrugli A, n. 1» (CAG). Paratypi: Ibidem, 12.v.1981, Grasso MP. et Scrugli A, n. 4 et5 (CAG), n. 2 (FI), n. 3 (Z); ibidem, 15.V.1983, Grasso M. P. et Scrugli A, n. 7 (CAG); «Sardegna, Nuoro, in agro circa pagum Laconi, in loco dicto Genna su Porcu. m 750 ca. 9.V.1981, Grasso M.P. et Scrugli A, n. 6» (CAG, fig. 2). Ophrys funerea Viv. Ophrys romolinii Soca, Ophrys chimaera W.Rossi & Contorni, Webbia 44(2): Sub Ophrys fusca Link Ophrys bertolonii Moretti. Type: [Italia] «Monte Amiata, loc. La Forestale, presso Abbadia S. Salvatore, ai margini di un rimboschimento a pino nero, 850 m, 16.V.1989, W. Rossi et M. Contorni» (FI). Ophrys fusca s.l. (lojaconoi?) Ophrys passionis Sennen, Ophrys gumprechtiana Soó, Acta Bot. Acad. Sci. Hung. 25(3-4): 362. (1979) nom. inval., design. typi omissa. Sub Ophrys fusca Ophrys garganica. Journal Europäischer Orchideen 49 (3-4):

44 Bas.: Ophrys sphecodes subsp. garganica Ophrys fusca Gumpr., Orchidee (Hamburg) 21(6): Type: Italia, Monte Gargano: Sipontea; Herb. Gumprecht, Freiburg. Syn.: Ophrys sphecodes subsp. garganica Ophrys fusca. GUMPRECHT, Orchidee (Hamburg) 1970: sine descriptio. in der Nähe von Siponto (Italien) am Fuß des Monte Gargano, Frühjahr Ophrys fusca s.l. (lojaconoi?) Ophrys tenthredinifera Willd. subsp. neglecta (Parl.) E.G.Camus, Ophrys sancti-leonardi O.Danesch & E.Danesch, Orchid. Eur. Ophrys- Hybriden: 229, contrary to Art. H.5.2. Sub Ophrys fusca Link subsp. fusca Ophrys tenthredinifera Willd. Type: Italien (Mte Gargano), bei Manfredonia, leg. Wiefelsputz, Herbar Wiefelsputz. Syn.: Ophrys lievreae Maire nothosubsp. sanctis-leonis (O.Danesch & E.Danesch) Del Prete, Webbia 37(2): nom. inval. sphalm. sancti leonardi. Ophrys fusca s.l. (obaesa Lojac.?) Ophrys pallida Raf., Ophrys ficuzzana H.Baumann & Künkele, Mitteilungsbl. Arbeitskreis Heimische Orchid. Baden-Württemberg 18(3): 468, Sub Ophrys fusca Link Ophrys pallida Raf. Type: "Italia, Sicilia: Ficuzza, 750 m , leg. H. Baumann & Kalteisen (STU). Ophrys fusca s.l. Ophrys oxyrrhynchos Tod., Ophrys marsianii Galesi, J. Eur. Orch. 27(2): contrary to Art. H.5.2. Sub Ophrys fusca Link Ophrys oxyrrhynchos Tod. subsp. oxyrrhynchos. Type: [Italia, Sicilia, Niscemi] C.da Banco (VB: 40.69), all'interno di un uliveto abbandonato, 280m, 09.IV.1993, R. Galesi (CAT). Ophrys fusca s.l. Ophrys romolinii Soca, Ophrys spuria G.Keller ex Gölz & H.R.Reinhard, Orchidee (Hamburg) 27(5): nom. inval., design. typi omissa. contrary to Art. H.5.2. Sub Ophrys bertolonii Mor. Ophrys fusca Link subsp. fusca. Type: [Italia, Sicilia] bei Canicattini Bagni. Ophrys fusca s.l. Ophrys tarentina Gölz et H.R.Reinhard, Ophrys parenzani Medagli, d'emerico & Ruggiero, Orchidophile (Asnières) 20(85): Sub Ophrys fusca Link O. tarentina Gölz et Reinhard. Type: Italia, Apulia, Martina Franca (TA), m 300 ca., 10.lV.1988, P. Medagli et S. d'emerico. In herb. Gruppo Naturalisti Salentini di Lecce sub Journal Europäischer Orchideen 49 (3-4): 2017.

45 Ophrys gackiae P.Delforge Ophrys incubacea Bianca, Sub Ophrys gackiae P.Delforge Ophrys incubacea subsp. incubacea Bianca. CAMPO, GIROS Notiz. 2012: sine nomine, nom. inval., design. typi omissa. Type: locus Burlò, apud Noto Antica (SR), legit V. Campo IGM fg. 277 U.T.M. 33S WA020895, 450 m circa s.l.m., terreno terrazzato, esposizione a nord, boschetto rado. Foto: V. Campo, P. Nanè. Ophrys gackiae P.Delforge Ophrys laurensis Melki & Geniez, Sub Ophrys gackiae P.Delforge Ophrys subfusca [] subsp. laurensis (Geniez & Melki) Kreutz. CAMPO, GIROS Notiz. 2012: sine nomine, nom. inval., design. typi omissa. Author of the species not indicated. Type: Mons Lauro, apud Buccheri (SR), legit V. Campo, IGM fg. 273 U.T.M. S33 VB , 948 m s.l.m., terreno pianeggiante, bosco rado. Ophrys gracilis (Büel, O.Danesch & E.Danesch) Paulus Ophrys lacaitae Lojac., Ophrys delargeana P.Delforge, Naturalistes Belges 69(2): contrary to Art. H.5.2. Sub Ophrys holoserica (N.L. Burm.) Greuter subsp. holoserica Ophrys lacaitae Lojacono. Type: Italia, Lazio, provincia Latina, Monti Aurunci, apud Formia, In herb. Pierre Delforge sub n 84/3. Syn.: Ophrys fajarama Perilli, GIROS Orch. Spont. Eur. 58(2): pro synonyme. [Italia, Puglia] località Fajarama (San Marco in Lamis, Foggia), m 735 s.l.m., 33T E N ; M. Perilli, cons. in BI (Herbarium Horti Botanici Barensis), spec. n sub Ophrys gracilis (Büel, O.Danesch & E.Danesch) Paulus Ophrys lacaitae Lojac. Ophrys gracilis (Büel, O.Danesch & E.Danesch) Paulus Ophrys parvimaculata (O.Danesch & E.Danesch) Paulus & Gack, Sub Ophrys holosericea [] subsp. gracilis (Büel, O. Danesch & E. Danesch) Büel, O. Danesch & E. Danesch Ophrys holosericea [] subsp. parvimaculata (O. Danesch & E. Danesch) O. Danesch & E. Danesch. Author of the species not indicated. PERILLI, GIROS Notiz. 2012: sine nomine, nom. inval., design. typi omissa. Type: [Italia, Puglia] Fajarama, San Marco in Lamis (FG). Ophrys gracilis (Büel, O.Danesch & E.Danesch) Paulus Ophrys tenthredinifera Willd. subsp. neglecta (Parl.) E.G.Camus, Ophrys maremmae O.Danesch & E.Danesch nothosubsp. alburnica Büel, O.Danesch & E.Danesch, Orchid. Eur. Ophrys-Hybriden: 226, Sub Ophrys fuciflora (Crantz) Moench subsp. gracilis Büel & Danesch Ophrys tenthredinifera Willd. Journal Europäischer Orchideen 49 (3-4):

46 Type: Italien (Provinz Salerno), Vorland der Monti Alburni, leg. O. et E. Danesch, (IB). Syn.: Ophrys maremmae O.Danesch & E.Danesch nothosubsp. alburnica (Büel, O.Danesch & E.Danesch) Del Prete, Webbia 37(2): pro synonyme. sub Ophrys holoserica (N.L. Burm.) Greuter subsp. gracilis (Buel et Al.) Danesch O. et E. Ophrys tenthredinifera Willd. Ophrys gravinensis D Alonzo Ophrys incubacea Bianca, Ophrys pulicchiensis D Alonzo & Perilli, GIROS Orch. Spont. Eur. 59(2): Type: [Italia] località Pulicchio (Gravina in Puglia, Bari), m 578 s.l.m., UTMWGS 84 33T E N ; leg. F. D Alonzo & M. Perilli, cons. in BI (Herbarium Horti Botanici Barensis), spec Ophrys gravinensis D Alonzo Ophrys parvimaculata (O.Danesch & E.Danesch) Paulus & Gack, Ophrys difonzoi nothosubsp. difonzoi D Alonzo & Perilli, GIROS Orch. Spont. Eur. 59(2): sub Ophrys gravinensis D Alonzo Ophrys holosericea [] subsp. parvimaculata O.Danesch & E.Danesch (Author of the species not indicated). Type: [Italia] località Pulicchio (Gravina in Puglia, Bari), m 591 s.l.m., UTMWGS 84 33T E N ; leg. F. D Alonzo & M. Perilli, cons. in BI (Herbarium Horti Botanici Barensis), spec Ophrys gravinensis D Alonzo Ophrys romolinii Soca, Ophrys rosae D Alonzo, GIROS Orch. Spont. Eur. 58(2): Sub Ophrys bertolonii Moretti Ophrys gravinensis D Alonzo. Type: [Italia] località Pulicchio (Gravina in Puglia, Bari), m 535 s.l.m., UTMWGS 84 33T E N ; leg. Filippo D Alonzo, cons. in BI (Herbarium Horti Botanici Barensis), spec. n Ophrys gravinensis D Alonzo Ophrys sipontensis (Gumprecht) O.Danesch & E.Danesch, Ophrys filomenoi D Alonzo & Perilli, GIROS Orch. Spont. Eur. 59(2): sub Ophrys gravinensis D Alonzo Ophrys sipontensis R.Lorenz & Gembardt Type: [Italia] località Pulicchio (Gravina in Puglia, Bari), m 578 s.l.m., UTMWGS 84 33T E N ; leg. F. D Alonzo & M. Perilli, cons. in BI (Herbarium Horti Botanici Barensis), spec Ophrys gravinensis D Alonzo Ophrys tenthredinifera Willd. subsp. neglecta (Parl.) E.G.Camus, Ophrys mariana D Alonzo, GIROS Orch. Spont. Eur. 58(2): contrary to Art. H Journal Europäischer Orchideen 49 (3-4): 2017.

47 Type: [Italia] località Pulicchio (Gravina in Puglia, Bari), m 535 s.l.m., UTMWGS 84 33T E N Filippo D Alonzo, cons. in BI (Herbarium Horti Botanici Barensis), spec. n Ophrys holosericea [] subsp. paolina (V.Liverani & Romolini) Ophrys tenthredinifera [] subsp. neglecta (Parl.) E.G.Camus. Author of the species not indicated, Ophrys maremmae O.Danesch & E.Danesch nothosubsp. bankosii D Alonzo & Perilli, GIROS Orch. Spont. Eur. 59(1): Type: Italia, Apulia, terra garganica (Fovea), agro Sancti Nicandri, tricesimo uno dimidioque miliario viae SS 89, m 261 s.l.m., UTMWGS84 33T E N ; leg. F. D Alonzo & M. Perilli, cons. in BI (Herbarium Horti Botanici Barensis), spec , data 27 aprile Ophrys incubacea Bianca (= O. atrata Lindl.) Ophrys lutea Cav., Ophrys piscinica P.Delforge & C.Delforge, Naturalistes Belges 67(5/6): Type: Italia, provincia Foggia, ad locum Piscina di Camparano In herb. P. Delforge sub n 86/5. Ophrys incubacea Bianca Ophrys lunulata Parl., Ophrys camarinana H.Baumann & Künkele, Mitteilungsbl. Arbeitskreis Heimische Orchid. Baden-Württemberg 18(3): 490, Type: Italia, Sicilia, Vittoria, 150 m , leg. H. Baumann & M. Kalteisen (STU). Ophrys incubacea Bianca Ophrys oxyrrhynchos Tod. subsp. celiensis (O.Danesch & E.Danesch) Del Prete, Ophrys gelana H.Baumann & Künkele nothosubsp. murgettensis Soca, Monde Pl. 95(470): Type: Italia, Taranto, La Murgetta, Masseria Parco della Vigna, alt 440 m (UTM XF 90.52). 30.IV MPU n RS Not listed in IPNI. Ophrys incubacea Bianca Ophrys oxyrrhynchos Tod. subsp. oxyrrhynchos, Ophrys gelana nsubsp. gelana H.Baumann & Künkele, Mitteilungsbl. Arbeitskreis Heimische Orchid. Baden-Württemberg 18(3): 492, Type: Italia, Sicilia: Gela, 100 m , leg. H. Baumann & M. Kalteisen (STU). Ophrys incubacea Bianca Ophrys panormitana (Tod.) Soó, Ophrys morensis Giotta & Piccitto nothosubsp. morensis, J. Eur. Orch. 33(4): Sub Ophrys incubacea Bianca Ophrys sphegodes [] subsp. praecox Corrias. Author of the species not indicated. Journal Europäischer Orchideen 49 (3-4):

48 Type: Italia, Sardegna, Mòres (SS), "Istrampu" (UTM: MK 88.68), oliveto abbandonato, m 379, , M. Piccitto & C. Giotta (FI). Ophrys incubacea Bianca Ophrys parvimaculata (O.Danesch & E.Danesch) Paulus & Gack, Ophrys pasquinae Silletti & Medagli, Silvae: epublished, contrary to Art. H.5.2. Sub Ophrys incubacea Bianca subsp. incubacea Ophrys holosericea (Burm.f.) Greuter subsp. parvimaculata (O. et E. Danesch) O. et E. Danesch. Type: [Puglia] Santeramo in Colle (Ba), legit G. N. Silletti, in LEC. Photography of Ophrys parvimaculata (O.Danesch & E.Danesch) Paulus & Gack. Not listed in IPNI. Ophrys incubacea Bianca Ophrys passionis Sennen, Ophrys celani O.Danesch & E.Danesch, Orchid. Eur. Ophrys-Hybriden: 223, Sub Ophrys atrata Lindl. Ophrys garganica (Nelson) O. & E.Danesch. Type: Italien (Mte Gargano), M. ividori, leg. O. et E. Danesch, (IB). Ophrys incubacea Bianca Ophrys passionis Sennen Ophrys romolinii Soca, Ophrys pescolusae Gennaio & Medagli, GIROS Notiz. 54: Sub Ophrys lyrata H. Fleischm. (O. bertolonii Moretti O. incubacea Bianca) Ophrys passionis Sennen. Type: [Italia] Pescoluse, Salve (LE), m 1 s.l.m., legit R. Gennaio, 8.IV.2013 (in LEC Herbarium Universitatis Lupiensis). Ophrys incubacea Bianca Ophrys promontorii O.Danesch & E.Danesch, Ophrys angelensis H.Baumann & Künkele, Mitteilungsbl. Arbeitskreis Heimische Orchid. Baden-Württemberg 18(3): 493, Type: Italia, Puglia: Mte. Gargano, Mte. San Angelo, 480 m , leg. B. & H. Baumann (STU). Ophrys incubacea Bianca Ophrys romolinii Soca Ophrys tarentina Gölz & H.R.Reinhard, Ophrys amatoi Silletti & Medagli, Silvae: epublished. Sub Ophrys lyrata H.Fleischm. Ophrys tarentina Gölz & H.R.Reinhard. Type: [Puglia] Santeramo in Colle (Ba), legit G. N. Silletti in LEC. Photography of Ophrys tarentina Gölz & H.R.Reinhard. Not listed in IPNI. Ophrys incubacea Bianca Ophrys romolinii Soca Ophrys tenthredinifera Willd. subsp. neglecta (Parl.) E.G.Camus, Ophrys lupiae O.Danesch & E.Danesch, Orchid. Eur. Ophrys-Hybriden: , Journal Europäischer Orchideen 49 (3-4): 2017.

49 Sub Ophrys atrata Lindl. Ophrys bertolonii Mor. Ophrys tenthredinifera Willd. Type: Italien (Apulien), bei Lecce, leg. O. et E. Danesch, (IB). Ophrys incubacea Bianca Ophrys sipontensis (Gumprecht) O.Danesch & E.Danesch, Ophrys dinovellae Romolini & Soca, GIROS Notiz. 55: 52, 65, Type: Italia. Campania. Salerno. Teggiano. La Manca di San Donato, alt. 678 m. 33T IV Coll. Romieg Soca. MPU. n RS Ophrys incubacea Bianca Ophrys speculum Link, Ophrys pantaliciensis Kohlmüller, Riech. & Schöb., Ber. Arbeitskr. Heim. Orch. 8(1): Type: Italia, Sicilia, prope Ferla, 540 m s.m. Bot. Staatssammlung München, leg. R. Kohlmüller, A. Riechelmann & M. Schöbinger, Ophrys incubacea Bianca Ophrys tarentina Gölz & H.R.Reinhard, Ophrys alberobellensis H.Baumann & Künkele, Mitteilungsbl. Arbeitskreis Heimische Orchid. Baden-Württemberg 18(3): 496, Type: Italia, Puglia: Martina Franca, 300 m , leg. B. & H. Baumann (STU). Ophrys incubacea Bianca Ophrys tenthredinifera Willd. subsp. neglecta (Parl.) E.G.Camus, Ophrys grampinii Cortesi, Ann. Bot. (Roma) 1(5): Sub Ophrys aranifera Huds. Ophrys tenthredinifera Willd. Type: Italia, Lazio, Roma. Via Appia Antica, leg. Grampini, 14 aprile RO / (et proferrre Mte Testaccio). Syn.: Ophrys manfredoniae O.Danesch & E.Danesch, Orchid. Eur. Ophrys- Hybr.: 228, pro syn. Sub Ophrys atrata Lindl. Ophrys tenthredinifera Willd. Italien (Apulien) bei Manfredonia, leg. O. et E. Danesch, (IB). vide SOCA Ophrys insectifera L. Ophrys minipassionis Romolini & Soca, Ophrys barbieriarum Romolini & Soca, GIROS Orch. Spont. Eur. 58(2): 'barbierii'. Type: Italia. Toscana. Siena. Montepulciano. Monti di Poggiano. alt. 501 m s.l.m. 32T V Coll. Romieg Soca. Cons. FI. n RS Ophrys insectifera L. Ophrys molisana P.Delforge, Ophrys rasinii Soca, J. Eur. Orch. 49(2): Italia. Abruzzo. L Aquila. Roccaraso. Contrada Barone. 33T Alt m. 23.V Coll. R. Romolini et R. Soca. Cons. in herb. FI. sub n RS Journal Europäischer Orchideen 49 (3-4):

50 Ophrys iricolor Desf. subsp. eleonorae (Devillers-Tersch. & Devillers) Paulus & Gack Ophrys lojaconoi P.Delforge, Ophrys quitadamoi Medagli & Turco, GIROS Notiz. 47: contrary to Art. H.5.2. Sub Ophrys iricolor subsp. maxima (A. Terracc.) Paulus & Gack Ophrys lojaconoi P.Delforge. Type: [Italia] apud Mattinata (FG), m 130 circa s.l.m., legit Turco e Medagli, 14.III.2011 (in LEC - Herbarium Universitatis Lupiensis). Ophrys iricolor Desf. subsp. eleonorae (Devillers-Tersch. & Devillers) Paulus & Gack Ophrys speculum Link, Ophrys sanctae-sofiae M.P.Grasso & Liverani, Orchidophile (Asnières) 24(109): Sub Ophrys iricolor Desf. Ophrys ciliata Biv. Type: [Italia] «Su Taccu» (NU), Sardaigne centrale, 5 km NO S. Sofia, Leg. M.P. Grasso. Musée de Botanique de Florence (FI). Ophrys lacaitae Lojac. Ophrys oxyrrhynchos Tod., Ophrys campoi Soca, Monde Pl. 95(470): Type: Italia, Sicilia (Messina), Roccella Valdemone (UTM VB 99.96). 28.IV MPU n RS Ophrys lacaitae Lojac. Ophrys tetraloniae W.P.Teschner. MORALDO & al., Atti Accad. Naz. Lincei, Rendiconti Cl. Sci. Fis. Mat. Nat. 387(264): sine descriptio. Italia, Lazio meridionale. Ophrys lojaconoi P.Delforge Ophrys passionis Sennen, Ophrys palladinoi Soca, Caesiana 19(43): Type: Italia. Puglia. Foggia. San Giovanni Rotondo. Alt. 135 m s.l.m. 33T IV Coll. Romieg Soca. Cons. in herb. MPU. sub n RS Ophrys lojaconoi P.Delforge Ophrys sipontensis (Gumprecht) O.Danesch & E.Danesch, Ophrys friderici nothosubsp. friderici D Alonzo & Perilli, GIROS Orch. Spont. Eur. 60(1): sub O. iricolor [] subsp. lojaconoi (P.Delforge) Kreutz O. sipontensis (R. Lorenz & Gembardt). (Author of the species not indicated). Italia, Apulia, in Gargano ad Manfredoniam, Foveanam provinciam, m 52 s.l.m. (UTMWGS 84 33T E N) ; leg. F. D Alonzo & M. Perilli, cons. in BI (Herbarium Horti Botanici Barensis), spec , data Ophrys lucana P.Delforge Ophrys passionis Sennen subsp. majellensis 614 Journal Europäischer Orchideen 49 (3-4): 2017.

51 (Helga Daiss & Herm.Daiss) Romolini & Soca, Ophrys lociceroi Soca, J. Eur. Orch. 46(3/4): contrary to Art. H.5.2. Type: Italia, Abruzzo, Chieti, Palena, San Cataldo, alt. 932 m, 33T Coll. Romieg Soca. MPU n RS Ophrys lucifera Devillers-Tersch. & Devillers Ophrys speculum Link, Ophrys capoliverii Iamonico, Forbicioni & Frangini, Ann. Bot. Fenn. 48(5): 436 (-437; fig. 1) contrary to Art. H.5.2. Sub Ophrys fusca Link subsp. lucifera (Devillers-Tersch. & Devillers) Kreutz Ophrys speculum Link. Type: Italy. Tuscany region, Livorno Province, Elba Island, Capoliveri, Gabbiani coast calcareous cliff (aspect south), 120 m a.s.l., 42 43' 2.93" N, 10 24' 13.11" E, 30.III.2009 L. Forbicioni, G. Frangini & D.Iamonico (FI). Ophrys lunulata Parl. Ophrys oxyrrhynchos Tod. subsp. oxyrrhynchos, Ophrys sicana H.Baumann & Künkele, Mitteilungsbl. Arbeitskreis Heimische Orchid. Baden-Württemberg 18(3): 502, Type: Italia, Sicilia: Gela, 100 m , leg. H. Baumann & M. Kalteisen (STU). Ophrys lunulata Parl. Ophrys romolinii Soca, Ophrys vittoriana H.Baumann & Künkele, Mitteilungsbl. Arbeitskreis Heimische Orchid. Baden-Württemberg 18(3): , Sub Ophrys bertolonii Moretti Ophrys lunulata Parl. Type: Italia, Sicilia: Vittoria, 150 m , leg. H. Baumann & M. Kalteisen (STU). Syn.: Ophrys solitaria Forster, Orchidee (Hamburg) 22(4): , nom. inval., design. typi omissa. sub Ophrys bertolonii Mor. Ophrys lunulata Parl. [Italia, Sicilia] bei Sortino im Hinterland von Syracusa an der Ostküste Siziliens. 11.IV Ophrys lunulata Parl. Ophrys speculum Link subsp. speculum, Ophrys syracusana O.Danesch & E.Danesch, Orchid. Eur. Ophrys-Hybriden: 227, contrary to Art. H.5.2. Type: Sizilien, bei Syracus, leg. O. et E. Danesch, (IB). Syn.: Ophrys syracusana O.Danesch & E.Danesch, Orchidee (Hamburg) 22(1): without type. Ophrys lunulata Parl. Ophrys tenthredinifera Willd. subsp. grandiflora (Ten.) Kreutz, Ophrys lidbergii Mazzola, Webbia 36(1): Sub Ophrys lunulata Parl. Ophrys tenthredinifera Willd. Type: [Italia, Sicilia] in declivitatibus Herctae Montis ad septemtriones versis Journal Europäischer Orchideen 49 (3-4):

52 "Manca Addaura", prope aditum inferioris specus. 150 m, 18 aprili 1981, P. Mazzola (PAL). Syn.: Arachnites lunulata var. benoitiana Tod., Hort. Bot. Pan. 2: 24, t sub Arachnites fuciflora Hoffm. var. panormitana Tod. Arachnites Bertoloni Tod. Bas.: Ophrys Benoitii Tin. Ind. Nei luoghi erbosi delle falde delle montagne calcaree dell agro Palermitano esposte a Settentrione, precipuamente nelle colline di Gibilrossa. dubium; Not listed in IPNI. Ophrys lutea Cav. Ophrys oxyrrhynchos Tod., Ophrys monicae La Spina, GIROS Orch. Spont. Eur. 59(2): contrary to Art. H5.2. sub Ophrys lutea subsp. lutea Cav. Ophrys oxyrrhynchos subsp. oxyrrhynchos Tod. Type: Italia, Sicilia, Silva Sancti Petri, terra Calatina, Catinensi provincia (DD N E) , Contrada Renelle, Bosco di Santo Pietro, Caltagirone (CT), leg. A. La Spina, cons. in CAT (Herbarium Universitatis Catanensis) n Ophrys lutea Cav. Ophrys romolinii Soca, Ophrys cugniensis Soca, Monde Pl. 92(460): Sub Ophrys bertolonii Moretti Ophrys lutea Cav. Type: Italia, Sicilia (Siracusa). Sortino, Monte Cugni. 19.IV MPU n RS Syn.: Ophrys opaca G.Keller ex Gölz & H.R.Reinhard, Orchidee (Hamburg) 27(5): nom. inval., design. typi omissa. Bas.: Ophrys opaca G.Keller, Monogr. Iconogr. Orchid. Eur., Bd. 2: nomen nudum (artificiel F. Denis). [Italia, Sicilia] bei Canicattini Bagni. Sub Ophrys bertolonii Mor. Ophrys lutea Cav. var. lutea. Not listed in IPNI. Ophrys lutea Cav. Ophrys tarentina Gölz & H.R.Reinhard, Ophrys circlarium Pellegrino, Ann. Bot. Fenn. 45(1): 66 (64-65) Type: Italy, Along road to Cerchiara di Calabria, 39 51' N, 16 23' E. 18.IV.2005, Pellegrino & al. s.n. (CLU). Ophrys lutea Cav. Ophrys tenthredinifera Willd. subsp. neglecta (Parl.) E.G.Camus, Ophrys personei Cortesi ex E.G.Camus, Iconogr. Orchid. Europe. Texte 2: Bas.: Ophrys personei Cortesi, Ann. Bot. (Roma) 13(2): nomen invalide, nom. prov. hybr. nov. ad interim. Type: [Italia] Contrada Arnèo, Serra degli Angioli distante da Nardo circa 30 km. Coll. Federico Personè. Herbarium? Not listed in IPNI. 616 Journal Europäischer Orchideen 49 (3-4): 2017.

53 Fig. 16: Ophrys dinarica Ophrys passionis subsp. majellensis, Palena, Abruzzo (CH), Fig. 17: Ophrys bertolonii subsp. bertoloniiformis Ophrys passionis, Rignano Garganico, Puglia (FG), Fig. 18: Ophrys incubacea Ophrys sipontensis, Campania (SA), Teggiano, Fig. 19: Ophrys insectifera Ophrys minipassionis, Montepulciano, Toscana (SI), Journal Europäischer Orchideen 49 (3-4):

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