PLANT DIVERSITY OF THE IWOKRAMA FOREST, GUYANA

Transcription

1 SIDA,BOTANICAL MISCELLANY BOTANICAL RESEARCH INSTITUTE OF TEXAS 21 PLANT DIVERSITY OF THE IWOKRAMA FOREST, GUYANA H.D. CLARKE, V.A. FUNK AND T. HOLLOWELL Using Checklists and Collecting Data to Investigate Plant Diversity. I: A Comparative Checklist of the Plant Diversity of the Iwokrama Forest, Guyana

2 PLANT DIVERSITY OF THE IWOKRAMA FOREST, GUYANA H.D. CLARKE, V.A. FUNK AND T. HOLLOWELL

3 SIDA,BOTANICAL MISCELLANY BOTANICAL RESEARCH INSTITUTE OF TEXAS 21 Using checklists and collections data to investigate plant diversity. I: A comparative checklist of the PLANT DIVERSITY OF THE IWOKRAMA FOREST, GUYANA H.D. CLARKE, V.A. FUNK AND T. HOLLOWELL BRIT 2001

4 ISSN ISBN SIDA, BOTANICAL MISCELLANY NO. 21 FOUNDED BY wm. F. MAHLER AND BARNEY L. LIPSCOMB, 1987 EDITOR: BARNEY L. LIPSCOMB BOTANICAL RESEARCH INSTITUTE OF TEXAS 509 PECAN STREET FORT WORTH, TEXAS USA CONTRIBUTING SPANISH EDITOR: FÉLIX LLAMAS DPT. BIOLOGÍA VEGETAL, UNIVERSIDAD DE LEÓN CAMPUS DE VEGAZANA S/N, LEÓN, SPAIN COVER DESIGN: LINNY HEAGY USING CHECKLISTS AND COLLECTIONS DATA TO INVESTIGATE PLANT DIVERSITY. I: A COMPARATIVE CHECKLIST OF THE PLANT DIVERSITY OF THE IWOKRAMA FOREST,GUYANA COVER PHOTOGRAPHS:THE HIGHEST PEAK IN THE IWOKRAMA MOUNTAINS (UNNAMED), ELEVATION 1000 METERS. INSET PHOTOGRAPH: CATTLEYA VIOLACEA (KUNTH) ROLFE PLANT DIVERSITY OF THE IWOKRAMA FOREST, GUYANA IS PUBLISHED WITH THE SUPPORT OF: BOTANICAL RESEARCH INSTITUTE OF TEXAS AND NATIONAL MUSEUM OF NATURAL HISTORY DISTRIBUTION OF COPIES BY: BOTANICAL RESEARCH INSTITUTE OF TEXAS 509 PECAN STREET FORT WORTH, TEXAS TELEPHONE: FAX: SIDA@BRIT.ORG BRIT SIDA, BOTANICAL MISCELLANY, NO MAY 2001 COPYRIGHT 2001 BY BOTANICAL RESEARCH INSTITUTE OF TEXAS PRINTED IN THE UNITED STATES OF AMERICA

5 TABLE OF CONTENTS ABSTRACT / RESUMEN / INTRODUCTION / THE IWOKRAMA FOREST 1 PLANT COLLECTING 4 EXPEDITIONS 6 RESULTS 12 ECOLOGY AND HABITATS Regional Context / Mora Forest / Forests Along the Lower Essequibo 15 Forests Along the Upper Essequibo / Iwokrama Mountains 16 Forests North of the Kurupukari Annai Road 17 Pakutau Hills,Turtle Mountain, and Sandstone / CONCLUSION 18 ACKNOWLEDGMENTS / REFERENCES 19 APPENDIX 1 Checklist of Plant Species from Iwokrama and Four Other Sites in Northeast South America 20 APPENDIX 2 Iwokrama Plant Collections 61 APPENDIX 3 List of Ranks of Species Richness by Family for Iwokrama, Four Other Sites in NE S. America, and the Guianas Overall 79 APPENDIX 4 Specialists Contributing Determinations of Iwokrama Specimens 84

6 Top photo / Daniel Allicock collecting plants in the Gorge in the Iwokrama Mountains. Bottom photo / Victoria amazonica (Poepp.) Sowerby

7 USING CHECKLISTS AND COLLECTIONS DATA TO INVESTIGATE PLANT DIVERSITY. I: A COMPARATIVE CHECKLIST OF THE PLANT DIVERSITY OF THE IWOKRAMA FOREST, GUYANA H.D. CLARKE V.A. FUNK and T. HOLLOWELL Department of Biology Biological Diversity of the Guianas Program University of North Carolina at Asheville Department of Botany MRC 166 Asheville, NC 28804, U.S.A. National Museum of Natural History Smithsonian Institution Washington, DC 20560, U.S.A. ABSTRACT Results are presented from a floristic inventory of the 360,000-hectare Iwokrama Forest (formally, The Iwokrama International Centre for Rain Forest Conservation and Development) in central Guyana. The collecting efforts, which resulted in 3887 collections, are described as are the ecology and habitats of the study site. Descriptive data extracted from the results are presented regarding patterns of diversity of the 147 families, 579 genera, and 1251 species that constitute the known flora of the Iwokrama Forest in comparison to four other floras of areas of restricted size (10, ,000 hectares) in the Guianas and northeastern Brazil. The actual number of species in the flora is estimated to be perhaps 50% greater than the known flora based on comparisons to the other floras in the region and the fact that almost half of the species are known from only single collections. The combined checklist of 4659 species is presented as an appendix. RESUMEN Se presentan los resultados de un inventario florístico del Iwokrama Forest de Ha (previamente, The Iwokrama International Centre for Rain Forest Conservation and Development) en la Guyana central. Se describen los esfuerzos en el muestreo, que dieron como resultado 3887 muestras, así como la ecología y hábitats del lugar de estudio. Los datos descriptivos extraídos de los resultados se presentan siguiendo los patrones de diversidad en 147 familias, 579 géneros, y 1251 especies que constituyen la flora conocida del Iwokrama Forest en comparación con otras cuatro floras de áreas de pequeño tamaño (10, ,000 Ha) en las Guyanas y noreste de Brasil. El número actual de especies en la flora se estima que sea quizás un 50% mayor que la conocida, basándonos en comparaciones con otras floras de la región y en el hecho que casi la mitad de las especies se conocen sólo de una colección. El catálogo combinado de 4659 especies se presenta como un apéndice. INTRODUCTION Intensive plant collecting by the Smithsonian Institution as part of the inventory of the biological diversity of the Iwokrama Forest in central Guyana affords an opportunity to present floristic data for a diverse, low-elevation rainforest in a very poorly known area. Under the Basic Rainforest Science objective of the Iwokrama Site Resource Survey (Hawkes & Wall 1993) the Smithsonian Institution s Biological Diversity of the Guianas Program was contracted to do a floristic inventory of the 360,000 hectare Iwokrama Forest (formally, the Iwokrama International Centre for Rain Forest Conservation and Development). The present work constitutes a summary of the floristic inventory; the inclusion of associated data from Iwokrama itself as well as comparative data from four other similar floristic inventories in nearby areas of northeastern South America helps to place the study in a more meaningful context. A combined checklist of all five florulas will prove useful for those working with the plant diversity of the Guiana Shield and adjacent Amazonia. A quantitative analysis of the descriptive floristic analysis is presented as a separate paper (Clarke & Funk, submitted). THE IWOKRAMA FOREST The Forest is located in central Guyana within administrative Region No. 8 (Potaro-Siparuni; Fig. 1). It lies to the north and east of the Annai Amerindian District with the village of Surama lying immediately southwest of the boundary and is centered around 4º 30'N, 58º 50'W. The exact boundaries are set out in the Iwokrama Rain Forest Programme Synopsis (Guyana Inter- Agency Committee 1992). It is bordered on the northeast by the Essequibo River, on the northwest

8 2 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana FIG. 1. General location and geographic features of the Iwokrama Forest.

9 BRIT.ORG/SIDA 3 by the Siparuni River, on the west by the Sipariparu River and the Burro Burro River, and on the south by the southern border of the Iwokrama Mountains, more or less the Kaiparu River and Ladysmith Creek. Guyana as a whole has a tropical climate characterized by a relatively high mean annual temperature exceeding 25 C at sea level; a diurnal range of temperature exceeding the annual range; and an annual temperature range of less than 5 C. Guyana has two rainy seasons that correspond to the passage of the intertropical convergence zone as it moves north between May and August and south in December January. The heaviest rains begin in May and cause extensive flooding of the rivers and areas where slopes are insufficient to provide adequate drainage. The December January rains are shorter, less intense, and do not penetrate as far inland from the Atlantic Ocean, so that in southern Guyana the passage of the intertropical convergence zone at this time of year brings only a slight increase in the frequency of convection storms that also occur throughout the dry season. These isolated storms do occur, however, with sufficient frequency to maintain closed canopy evergreen tropical moist forest throughout most of southern Guyana. In the north, the dry seasons are also ameliorated by periodic storms and this, combined with the occurrence of two rainy seasons, also maintains evergreen tropical moist forest. Climatic data are for the most part lacking, but most of the country, Iwokrama included, probably receives between two and three meters annual precipitation. Data from the Mabura Hill area immediately north and east of Iwokrama indicate an annual rainfall of 2700 mm (Khan et al. 1980; ter Steege et al. 1996). The Rupununi Savanna, which borders Iwokrama on the southwest, has formed primarily as a result of edaphic, rather than climatic factors. This region has highly weathered white sands overlying a clay hard pan that is recalcitrant to penetration by tree roots and access to ground water in the dry season and promotes extensive flooding in the May August rainy season. Fragmentary meteorological data from Annai (Hawkes & Wall 1993) do indicate less precipitation there than Kurupukari, probably due to higher albedo in the savanna which disrupts convections storms and results in slightly less precipitation in savanna proper, particularly in the dry season. This is probably a very local phenomenon, however, as evidenced by the very sharp savanna-forest ecotone so probably only areas of the forest very near the ecotone experience diminished rainfall. Thus, although annual precipitation in the Rupununi is still more than adequate to support the growth of tropical moist forest, forest only occurs in upland areas on granitic or lateritic substrates; along rivers as gallery forest in alluvial sands; and in so-called bush islands where the clay pan has been penetrated, allowing for adequate drainage and access to soil moisture and the development of forest (Eden 1964, 1973; Frost 1968; Rutherford & Hills 1979; Sinha 1968). So, despite the lack of climatic data, there is little reason to expect there to be any significant moisture or other climatic gradients within Iwokrama. Where this might be suspected, such as in the Humiria/Pagamea scrub forest along the Kurupukari Annai road in the southwestern part of the Forest, one should instead look for factors related to substrate and drainage as determining the vegetation type. The prevailing winds are northeasterly and originate from the Atlantic Coast. Within the Forest, the slopes of the mountains which face the incoming trade winds may be expected to have increased rainfall due to orographic precipitation, although, since the Iwokrama Mountains are isolated and no more than 1000 m in elevation, probably neither orographic precipitation nor a rainshadow on the lee side of the mountains are pronounced effects. Collection of more data from the Iwokrama Mountains will be necessarily to document more thoroughly how it differs climatically from adjacent lowland areas. Forest types can be difficult to define on the ground because forests do not occur as the discrete entities seen on vegetation maps. The small-scale habitat heterogeneity (e.g.: Euterpe palm swamp) that is invariably found within a forest area frustrates simple definitions over large areas. Davis and Richards (1933, 1934) identified five intergrading types in their classic tropical ecology paper on central Guyana s lowland rainforests: 1) Mora (Mora excelsa Benth.); 2) Morabukea (Mora gonggrijpii Sandw.), 3) Mixed (no dominance), 4) Greenheart (Chlorocardium rodiei [Schomb.] Rohwer et al.) and 5) Wallaba (Eperua falcata Aublet). Mixed rain forest (sensu

10 4 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana Davis and Richards) exists in moderate drainage conditions, with a similar set of species in changing ratios of dominance. All of the forest types recognized by Davis and Richards occur within the Forest. Other authors (ter Steege et al. 1993; Ek 1997) considered soil drainage the most important factor in classifying the lowland forest habitats at Mabura Hill, north of the Iwokrama Forest. Huber et al. (1995), in a vegetation map of Guyana, represent the Iwokrama area as having mainly lowland, evergreen mixed forest and wallaba forest and a small area of premontane forest in the Iwokrama Mountains. The work to describe and map vegetation in diverse and complex tropical plant communities is one of successive approximation; none of the above systems will be found to be completely adequate for the Iwokrama Forest, but given the checklist presented here and consideration of soil and drainage a useful delineation of the forest types is possible following further surveys. Discussions of forest types and plant communities for the parts of the forest that were sampled in plant collecting are presented below in the section on the ecology and habitats of the forest and in the individual expedition reports. PLANT COLLECTING This report is based on the 3887 collections made in the Iwokrama Forest during the 18 expeditions undertaken by taxonomists associated with the Smithsonian Institution (Fig. 2). The very few historical collections that exist from this area (Ek 1990) were not used because of the poor quality of their locality and other associated data. Since the quality of this product is directly dependent on how these collections are made and identified, a discussion of these methods is appropriate. Tropical plant diversity is such that determination of a plant s identity to the species level is usually only possible following detailed examination of fertile specimens. Identification of sterile material from Iwokrama will be facilitated, however, by reference to the checklist presented here in conjunction with other key references including field guides such as Gentry (1993) which emphasizes vegetative characters, and floristic references such as Boggan et al. (1997), the Flora of the Guianas series (published by Kluwer) and the Flora of the Venezuelan Guayana (published by Timber Press) along with well determined reference collections. Collections for this study were made with the goal of collecting as many of the taxa that occur within the forest as possible, concentrating on the non-weedy species which comprise the vast majority of the diversity of the flora of the Iwokrama Forest. Factors that affect how thoroughly plants are collected include the methods for sampling of plants of diverse habits and phenology and the sampling of different areas and habitats. Tropical forests differ from temperate forests by having an increased number of plant habit types: In addition to the canopy tree, understory tree and shrub, and herbaceous elements found in temperate forests, tropical forests also have lianas, epiphytes, midstory trees, and canopy emergents. Plant collecting that does not aggressively pursue all of these elements will not accurately reflect the plant diversity of the study area. Collecting of plants of diverse habit types was pursued by using tree climbing spikes and extendible aluminum pole clippers to reach and collect in the canopy. Climbing is particularly important because the tree, epiphyte, and liana forest of Iwokrama is both very diverse and very poorly known. Sampling of different areas and habitat types was carried out by identifying and collecting in as many different areas of the forest and habitats as possible. Unfortunately, neither identifying the different habitats of Iwokrama nor gaining access to them is trivial. Habitats in Iwokrama have been identified as either a set of landsystems (Hawkes & Wall 1993) or in reference to Davis and Richards (1933, 1934) forest types. Although the landsystems of Hawkes and Wall (1993) maintains a useful distinction between forests on granitic- or doleritic-derived soils, application of Davis and Richards system allows for a finer scale characterization of the habitats of Iwokrama. Although surveys on the ground in the forest are still inadequate to accurately map the soils and associated vegetation types of Iwokrama, broad scale patterns are apparent that allowed for targeting areas based on topography, drainage, and soils. Access to these areas is gained through the rivers, the Kurupukari Annai road, and the limited number of transects that have been cut. The transects are particularly important for allowing sampling away from riparian and ruderal roadside habitats and for

11 BRIT.ORG/SIDA 5 FIG. 2. Summary of all collections from the Iwokrama Forest. allowing access to the mountainous areas of the Forest. Collecting was carried out on all of the transects in the forest and additional transects were cut in a number of instances. Transects of lengths from 3 10 kilometers were sampled at different times of the year with the expectation that varying phenologies would allow for sampling of additional fertile taxa. Because northern South America has the highest plant diversity in the world, even skilled field botanists working in the neotropics generally have an apprehension of this diversity only at the generic level. However, determining local species diversity is possible with knowledge of the genera because species may usually be distinguished, though not determined, by visual inspection. Identification of the collections listed here was carried out by distributing duplicate

12 6 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana specimens to specialists. The Biological Diversity of the Guianas Program has cultivated a longterm relationship with specialists in the groups of plants that occur in Guyana to insure that specimens are determined as accurately as possible (Appendix 4). These specialists produce monographs, revisions, and treatments for floras of these plant groups. Smithsonian benefits by getting the most accurate determinations and the specialists benefit by getting good, well-documented specimens to use in their research. Collecting is an important component of the overall work to produce revisions, monographs, and flora treatments for the Guianas because the plants from the Guianas are in general poorly known and very under-represented in herbaria. Many species, for instance, are only known from the original type collection; this specimen may have inadequate documentation or may not display enough distinguishing features (flowers or fruit, e.g.) and certainly more collections may be needed to understand that plant s morphology, phenology, distribution, ecology, variation, and delimitation with respect to closely related species. There is a major impediment, however, being faced by organizations trying to carry out this type of work, which is the lack of trained systematists to participate in all phases of this work. In developed countries, this situation has been precipitated by the retiring of many botanists with specialized knowledge and experience in specimen-based systematics research and their replacement by systematists oriented towards molecular and phylogenetic research. Developing countries face an even more acute problem since they contain only 6% of the world s systematists yet have 80% of the world s species (Systematics Agenda 2000, 1994). Countries that are as rich in biodiversity as Guyana have never been successful in understanding this diversity by relying on workers in developed countries to carry out this work and this situation is not likely to improve given the trends in systematics research away from floristic and monographic research. Fortunately, however, even a modest investment in Guyana s human resource potential and infrastructure could accomplish a great deal in this area. In contrast to other fields in biology, specimenand morphology-based plant systematics demands only modest resources including reference materials, dissecting microscopes, and a well-curated herbarium. With these resources and a group of trained botanists, much of the floristic and monographic work on the plants of Guyana now being carried out in North America and Europe could be very adequately done in Guyana. Efforts to search for Guyanese counterparts to accompany the collectors into the field concentrated on the students and faculty of the University of Guyana. Nearly all of our work has been done with extensive assistance from Guyanese Amerindians in the interior. They have an intimate knowledge and great affinity for the forest. The collaboration with the Amerindians included training in methods of plant collecting, including methods of documentation and plant identification; training with maps, gps, and compasses; and introduction to ecological principles. The training provided is useful both for further work in field botany and for work in related fields including forestry, wildlife management, faunistics and ecology, and ecotourism. Training was provided to members of the Amerindian communities from areas adjoining Iwokrama (B. Allicock, D. Allicock, S. Allicock, C. Captain, M. Captain, M. George, B. James, and A. Rodrigues) as well as those from other parts of the country (R. Williams (Bartica) and Claudius Perry (Maruanawa)). A number of the people listed above as well as several other members of the Amerindian communities surrounding Iwokrama participated in the two Amerindian Parataxonomy Courses offered by the Centre for the Study of Biological Diversity (University of Guyana), in collaboration with the Smithsonian Institution. EXPEDITIONS A. Collector: Pedro Acevedo Rodríguez (US) Date: 19 February 1 March 1990 General Location: Along new road and around Surama Village (Fig. 3). Number Series: PA Collected with: C. Feuillet (P), T. McDowell (US), S. Tiwari (BRG) This was the first recent expedition made into the Iwokrama area. Part of the time was spent following the road crew from the Brazilian gold mining company Paranapanema as it built a road through southwest Guyana that eventually bisected the Iwokrama forest.

13 BRIT.ORG/SIDA 7 FIG. 3. Collecting expeditions B. Collector: Tim McDowell (US) Date: 19 February 1 March 1990 General Location: Surama Village and along new road being built by Paranapanema (Fig. 3). Number Series: TM Collected with: P. Acevedo (US), C. Feuillet (P), S. Tiwari (BRG) Assisted by: Sidney Allicock Region around Surama. Collectors attempted to follow the road crew but found it very difficult because trees were being pushed down constantly and immediately pushed up to 40 meters to clear the roadway; the road advanced ca. 2 km per day. Trees were quickly broken and intermingled and duplicate specimens were difficult to obtain.

14 8 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana C. Collector: Tim McDowell (US) Date: September 1990 General Location: Along the Essequibo River and on islands near Kurupukari Village and along survey line for road (Fig. 3). Number Series: TM McDowell spent only a few days collecting, mostly along the Essequibo River and on the islands near Kurupukari Village with one expedition through the forest along the survey line of the road. D. Collector: Bruce Hoffman (US) Date: 16 April 5 May 1992 General Location: Kurupukari area, the Siparuni River at Pakutau Falls, the lower slopes of the Iwokrama Mountains, the Surama Village area, and at various locations along the Kurupukari Annai Road (Fig. 3). Number Series: BH Collected with: Toby Pennington (K) & G. Gharbarran (BRG) Field Assistants: C. Capellaro, D. Allicock, & T. Allicock This three week botanical collecting expedition was in collaboration with an international team of consultants doing a multi-disciplinary inventory. The mission of the Commonwealth Project was to study the site and develop methods of sustainable natural resource use, with some of the land being set aside as wilderness. A variety of soil types and vegetation classes occur within the boundaries including basic doleritic, acidic granitic, white sand, brown sand, sedimentary, mora forest, wallaba forest, commercial quality mixed-forest, acidic bog, and scrub. E. Collector: Patrice Mutchnick (US) Date: 27 November 14 December 1994 General Location: Essequibo River, Pakutau hill transect, Siparuni River, Kurupukari Falls (Fig. 3). Number Series: PM Collected with: Vicki Funk (US) Field Assistants: Bertrand Allicock, Erol McBirney, Raymond Edwards, Clive Captain, and Rodrigues Antone The majority of the collecting took place on the Siparuni River, a major tributary of the Essequibo River. The confluence is approximately 40 kilometers downstream (North) of the base camp at Kurupukari. Collecting was also done along several established transects and on the Essequibo directly north and south of the base camp. F. Collector: Patrice Mutchnick (US) Date: 10 February 3 March 1995 General Location: Pisham Pisham, Akromukro Falls, Ladysmith Creek, and Mount Daniel transects (Fig. 4). Number Series: PM Field Assistants: Bertrand Allicock and Daniel Allicock. The majority of the collecting took place on the Essequibo River, upstream from Kurupukari base camp to the border of the Forest at Lady Smith Creek. Additionally, a separate expedition was made to one of the two parts of the Iwokrama Mountains surveyed in this work, Mt. Daniel, located about 18 kilometers southwest of the base camp. G. Collector: David Clarke (US) Date: 17 September 9 October 1995 General Location: Upper Essequibo, Burro Burro River, and transects along Kurupukari Annai Road (Fig. 4). Number Series: Assistants: Daniel Allicock, Milner Captain, Clive Captain, and Baptist James The work to be done on this expedition was to extend and compliment that done by previous collectors, viz. Bruce Hoffman, Toby Pennington, and Patrice Mutchnick. Collecting was concentrated along rivers and machete cut transect lines with the goal of sampling across a number of different ostensible land systems described in the Iwokrama Phase 1 report. It was also expected that the collecting that would be conducted would help to achieve a better understanding of these land systems and the vegetation they support. During the course of this expedition, Iwokrama personnel abandoned mapping of the Forest on the basis of these land systems in favor of a map delineating more detailed vegetation types based on maps produced by the Guyana Forest Department and Food and Agriculture Organization of the United Nations (1970).

15 BRIT.ORG/SIDA 9 FIG. 4. Collecting expeditions H. Collector: David Clarke (US) Date: 4 November 2 December 1995 General Location: Pakutau Hills along the Siparuni River, the Iwokrama Mountains, and a transect along the lower Burro Burro River (Fig. 4). Number Series: DC Collected with: Bruce Hoffman (US) Assisted by: Daniel Allicock, Milner Captain, Baptist James, Antone Rodrigues, and Marcus George

16 10 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana Collecting on this expedition concentrated on areas not visited earlier by David Clarke viz., the Pakutau Hills along the Siparuni River, the Iwokrama Mountains, and a new transect along the Burro Burro River. A new vegetation map of the Forest was used to help direct collecting efforts and, in turn, it is hoped that the collection information could help to refine the map by providing information about previously unvisited areas. Throughout this expedition, Bruce Hoffman sampled 0.1 ha plots in an effort to assess alpha- and beta-levels of plant species diversity. I. Collector: David Clarke (US) Date: 14 March 6 April 1996 General Location: Akromukro Falls, Ladysmith Creek, Moco Moco II Creek, Iwokrama Mountains, Turtle Pond, Upper Burro Burro River, and southwest border of the Forest (Fig. 5). Number Series: DC Collected with: Bruce Hoffman (US) Assisted by: Daniel Allicock, Baptist James, Milner Captain, and Antone Rodrigues. This expedition was carried out together with Bruce Hoffman who was doing research on Heteropsis and its potential as a non-timber forest product. Collecting on this expedition in part emphasized recollecting in areas previously collected to document phenology. In some cases fruiting samples were collected from individual plants that had previously been collected in flower. Excursions were also made to previously uncollected areas such as an outlier of the Iwokrama Mountains, Sandstone on the upper Burro-Burro River, and floodplain areas adjacent to the Essequibo River near Turtle Pond. J. Collector: David Clarke (US) Date: 17 September 5 October 1996 General Location: Iwokrama Mountains, Turtle Mountain, and areas near base camp (Fig. 5). Number Series: DC Assisted by: Milner Captain, Baptist James, and Antone Rodrigues This expedition was carried out to collect in previously uncollected areas near the base camp and Turtle Mountain on the Essequibo River and to collect intensively at higher elevations in the Iwokrama Mountains. Two newly cut transects were sampled, one near the base camp on the Kurupukari Annai road in mixed forest on undulating terrain and one from the Essequibo River through forest prone to flooding and heavy siltation to an exposed area at the top of a cliff on Turtle Mountain. More effective collecting in the Iwokrama Mountains was carried out by establishing a camp at the top of the drainage in the gorge between the highest peaks in the mountains. Carrying ration and equipment from the road to the base of the mountains and up the gorge is difficult, but the time needed to climb the mountains from the camp at the bottom of the gorge is too long (3 hours each way) to allow for sufficient time to collect on the mountains. K. Collector: David Clarke (US) Date: March1997 General Location: Iwokrama Mountains (Mt. Daniel and gorge between the highest peaks in the mountains) and near base camp (Fig. 5). Number Series: Collected with: Scott Mori (NY) and Scott Heald (NY) Assistants: Romeo Williams, Claudius Perry, Baptist James, and Antone Rodrigues Scott Mori, a senior curator at the New York Botanical Garden, participated in this expedition as did his assistant, Scott Heald. Together, these two provided additional expertise and enhanced intensity of collecting efforts in the areas visited. Scott Mori s contribution was particularly valued as he has nearly completed a flora of Central French Guiana (Mori et al. 1997) and was able to provide important insights into the similarities and differences between that flora and Iwokrama s. L. Collector: Bruce Hoffman (US) Date: 15 May 21 May 1995 General Location: Surama area and upper Burro Burro River (Fig. 4). Number Series: BH Collected with: C. Ehringhaus (US) Field Assistant: Daniel Allicock This expedition was part of an effort to document the flora of the forest in a more quantitative fashion. Two 0.1 hectare transects were established during May, each located within one of the six different land systems recognized in the Phase 1, Site Resource Survey (Hawkes & Wall 1993).

17 BRIT.ORG/SIDA 11 FIG. 5. Collecting expeditions M. Collector: Bruce Hoffman (US) Date: 8 29 November 1995 General Location: Pakutau Hills, Burro Burro River, and Iwokrama Mountains (Fig. 4). Number Series: BH Collected with:david Clarke (used separate numbers see expedition report for details) Field Assistants: Daniel Allicock, Milner Captain, Antone Rodrigues and Baptist James This expedition was part of an effort to document the flora of the Forest in a more quantitative fashion. Four 0.1

18 12 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana hectare transects were established during November, each located within one of the six different land systems recognized in the Phase 1, Site Resource Survey (Hawkes & Wall 1993). N. Collector: Bruce Hoffman (US) Date: March 1996 General Location: Near Ladysmith Creek, SE border of Forest (Fig. 4). Number Series: BH Collected with: David Clarke (separate numbers), see Clarke report for details Field Assistants: Daniel Allicock The purpose of this expedition was to determine the density of Heteropsis flexuosa (Kunth) Bunting, commonly called nibbi. The results indicate that nibbi was as common in some areas of the forest as it is in the Pomeroon area of northern Guyana, the source of most commercial nibbi, but not as extensive. O. Collector: Christiane Ehringhaus (US) Date: May 1995 General Location: Annai and Surama area (Fig. 4). Number Series: CE Collected with: w/ B. Hoffman beginning on the 17th Field Assistants: Daniel Allicock Collecting expedition in and around Annai and Surama Villages and into upper Burro-Burro River area. P. Collector: Scott Mori (NY) Date: 3 27 March 1997 General Location: Kurupukari area and Iwokrama Mountains (Fig. 5). Number Series: SM Collected with: D. Clarke (used separate numbers) and Scott Heald (NY) For discussion of expedition see Section K, Clarke March 1997 expedition report. Q. Collector: Andre Chanderbali (MO) Date: July 1996 General Location: Along the Essequibo River, Iwokrama Mountains and Surama area (Fig. 5). Number Series: AC Collected with: D. Gopaul (BRG) Assisted by: Antone Rodrigues This expedition concentrated on documenting the Lauraceae of the Forest. Travel was by road and river to cut lines or trails that penetrate the forest for several kilometers. Ten of these trails and the vegetation along the Essequibo River boundary of the Forest were surveyed. R. Collector: Andre Chanderbali (MO) Date: 8 16 July 1997 General Location: Base camp at Kurupukari and revisited sites from 1996 expedition (Fig. 5). Number Series: AC Collected with: D. Gopaul (BRG) Assisted by: Bradford Allicock, Antone Rodrigues and Clive Captain RESULTS Since 1992, collectors associated with the Smithsonian Institution have visited the Iwokrama Forest 18 times and made 3887 collections of plants. Examination of the collections data shows that 81% of the total number of specimens have been identified to species and 91% of the 3323 fertile collections have been identified. Of the residual 9% of indeterminate specimens, approximately one third are in three large families (Melastomataceae, Bignoniaceae, and Flacourtiaceae) that proved recalcitrant due to the lack of an available specialist. The collections are distributed across 147 families, 579 genera, and 1251 species. This list was compiled together with checklists of four other florulas of areas on and adjacent to the Guiana Shield: Mabura Hill (Ek 1997) and Kaieteur Falls National Park (Kelloff & Funk 1998), both in Guyana; Saül, Central French

19 BRIT.ORG/SIDA 13 Guiana (Mori 1997); and Reserva Ducke near Manaus, Brazil (Ribeiro et al. 1999). The checklists were reconciled with respect to their synonymies using Boggan et al. (1997) and other references. Only native vascular plants were included in the combined checklist and only taxa determined to species level or below are shown here with the exception that taxa determined to genus were included as a unique taxon when there were no fully determined taxa for that genus at a given site. The combined checklist contains 4659 species and is provided in Appendix 1. Appendix 2 lists the plant collections that form the basis of the Iwokrama checklist. The identified collections show that the five most diverse families at the species level in Iwokrama are the Fabaceae (106 spp.), Rubiaceae (82 spp.), Orchidaceae (51 spp.), Cyperaceae (42 spp.), and the Myrtaceae (37spp.). A table comparing the rank order of species diversity of families inclusive to the twentieth most diverse family in each of the five florulas as well as the Guianas overall from Boggan et al. (1997) can be found in Appendix 3. These results confirm for Iwokrama what has been observed for other low-elevation rainforests sites, viz. that certain families such as the Fabaceae are over-represented while other families characterstic of either open habitats or higher elevation are under-represented (Lentibulariaceae, Bonnetiaceae, Bromeliaceae, Xyridaceae, and Eriocaulaceae, e.g.). The level of completeness of this checklist may be evaluated both in comparison to data for species diversity of the other four floras (Table 1) and by considering a histogram that compiles the number of specimens by which each of the 1251 species are represented in the collections made during the course of field work (Table 2). Iwokrama s 1251 species are low compared to Central French Guiana s 2069 or Reserva Ducke s The lower number for Iwokrama is almost certainly an indication of incomplete collecting rather than lower actual diversity; the flora of Central French Guiana is documented by 12,847 specimens compared to Iwokrama s 3887 and almost all of the collecting at Iwokrama has been carried out over the past several years whereas Reserva Ducke may be the best known area in Amazonia, having benefited from a wealth of historical collections and intensive recent collecting as part of the flora project. Similarly, Kaieteur Falls with an area of 58,000 ha and Mabura Hill with an area of 220,000 ha are probably also undersampled. Table 2 also indicates that over half (640) of the 1251 species from Iwokrama are known from only a single collection, strongly suggesting that continued collecting effort would yield a substantial number of new records. It is difficult to estimate the total plant diversity of Iwokrama with the data at hand, but a figure of approximately 2000 species may be reasonable given that Central French Guiana is a comparable, low elevation area of the Guiana Shield. Iwokrama s plant diversity may even exceed that of Central French Guiana s given its greater size (360,000 vs. 140,000 ha), greater elevational gradient (1000 m vs. 760 m), and higher habitat diversity (soils derived from both sandstone and granitic substrates vs. solely granitic based soils and riparian habitats associated with a major river, the Essequibo, vs. only riparian habitats associated with small- or medium-sized rivers). That Reserva Ducke, with an area of 10,000 ha, is smaller than any other of the four flora areas yet has the second highest diversity has been attributed to its position at the contact point among several Tertiary and Quaternary refugia indicating that its diversity may have been enhanced by the coalescence of the distributions of erstwhile allopatric species in this area (Oliveira & Daly 1999; Oliveira & Mori 1999). This assertion is further substantiated by noting that Reserva Ducke s generic diversity (658) is less than Central French Guiana s 781, comparable to Iwokrama s 579 and Mabura Hill s 573, and only somewhat more than Kaieteur Falls 492 genera, and arguing that, particularly in the case of late Tertiary and Quaternary refugia, most diversity was generated at the species rather than genus level. Inspection of the checklist in Appendix 1 allows for qualitative confirmation of this scenario because Reserva Ducke is seen to be particularly diverse in large, diverse genera such as Inga, Protium, Licania, Ocotea, etc. that may be expected to conform to this scenario. Data from Table 1 regarding the extent to which the diversity represented by the five checklists is shared among or is unique to each area has implications for both the usefulness of the combined checklist and conservation issues related to patterns of plant distributions. Overlap of

20 14 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana TABLE 1. Species diversity of the five florulas. Diagonal: Number of unique species/total number of species. Off diagonal: Number of uniquely shared species/number of shared species. Total = 4659 Iwokrama Mabura Kaieteur 312/ / / /383 63/ / / /627 73/ /2069 C. Fr. Ducke Guiana Iwokrama Mabura Kaieteur C. Fr. Guiana Ducke 40/370 51/422 41/ / /1922 TABLE 2. Number of collections representing species found in Iwokrama. species diversity is less than half in the case of Iwokrama and Mabura Hill and in comparisons involving Reserva Ducke is as low as one quarter. The combined checklist contains 4659 species or slightly more than half that of the 8507 species in the checklist of the Guianas (Boggan et al. 1997). So, although the total area of coverage of this checklist is but a small fraction of that of Boggan et al. (1997), because the flora areas encompass a broad expanse of the geographic and floristic gradients of the Guiana Shield and adjacent Amazonia, it still represents a good deal of the plant diversity of the entire region and should prove useful to those working with the plant diversity of a wide area of northeastern South America. That relatively little overlap is observed among the floras is an indication of a high degree of floristic heterogeneity among the areas. Even in the case of Iwokrama and Mabura Hill, which are contiguous and are remarkably similar with regard to climate, soils, geology, and habitats, the fact that they share only half of their floras is an indication of the patchiness of plant distributions which is apparently uncorrelated with ecological factors and perhaps corresponds better to historical factors. This finding has important implications for conservation policy and indicates that policy must be created with an understanding of plant distributions, hopefully on the basis of a good deal more field work,

21 BRIT.ORG/SIDA 15 rather than just a coarse understanding of ecological parameters. Thus the present work gives little assurance that conservation measures in Iwokrama, which has been planned for both development and conservation, would be effective in preserving the plant diversity of Mabura Hill which is being actively logged as part of a logging concession. The overlap of the Iwokrama and Mabura Hill and the other floras is also probably a conservative estimate given that the floras are undersampled and that widespread and common species have been preferentially sampled over uncommon, endemic, or patchily distributed species. ECOLOGY AND HABITATS Regional Context. The Iwokrama Forest is located in central Guyana and the composition of its forest are typical of lowland, evergreen forests found in much of northern and central Guyana. The soils that determine this composition are also characteristic of most lowland sites in northern and central Guyana, dominated by granitic and doleritic brown sands, laterite, and white sands resulting from either in situ erosion of sandstone substrates or deposition of sediments from the sandstones of the Pakaraimas Mountains. Although the Iwokrama Forest is not part of the Pakaraima Mountains, these mountains are found immediately across the Siparuni River from the Forest. Despite the elevational differences, certain affinities between the floras of the two regions are apparent such as between the Humiria scrub of Iwokrama and similar scrub areas in the Pakaraimas. Many tree species found in the forest that are typical of the northern and central lowland forests of Guyana are at or near the southern limits of their distributions: Chlorocardium rodiei (greenheart), Mora excelsa (mora), Peltogyne venosa (purpleheart), Caryocar nuciferum (sauri), and Heisteria cauliflora. This observation should have some bearing on the conservation plan of the forest, because, for example, the Iwokrama Forest is now the only area in Guyana that has stands of Chlorocardium that are not allocated as a timber concession. The forests of southern Guyana, as one moves southward from Iwokrama, experience successive replacement of typical guianan by amazonian elements so that, by the time one reaches the southernmost forests in the Acarai Mtns., there are very few species shared between the two areas. This turnover occurs despite the fact that there are very few gradients with regard to soils, climate, topography, elevation, or other abiotic factors. In the absence of such environmental gradients that may be used to explain this variation, historical factors from the Paleocene onwards may need to be invoked to explain how essentially different floras occupied areas that are otherwise so uniform. Observations on the ecology and plant communities within Iwokrama may best be presented by focusing on a number of sites and plant communities within the forest. Mora Forest. Areas along the major rivers and many other relatively flat, low lying areas experience a good deal of flooding during the rainy season. These areas are dominated by Mora excelsa, a canopy emergent with large buttresses, and a few other associates such as Eperua falcata, Catostemma spp., Aspidosperma excelsa, Tovomita spp., and Caraipa spp. These areas very often have basal areas that are strongly dominated by Mora excelsa and tend to be lower in overall diversity than upland terra firme forests. Mora forests along the Essequibo River may extend up to two km from the river itself and occupy a low lying area behind levees along the riverbank. Mora forests along the Siparuni and Burro Burro rivers occupy a much narrower strip no more than one km wide that is also usually a low lying area behind levees forming the riverbank. Soils in these forests are alluvial brown sands. Where periodically inundated forests occur away from the major rivers, soils are of varying composition. Mora forest will develop on brown sands in these areas, as at Moco Moco II creek along the Kurupukari Annai road and may also develop on white sands along blackwater creeks as is found in the first kilometer of the A B transect. Certain periodically inundated forests on white sand with a clay hard pan near the surface, however, do not have mora, but are dominated by a semi-open forest with Protium, Maximiliana, Euterpe, Dimorphandra, Moronobea, Pouteria, and Catostemma such as is found in the first two kilometers of the Mt. Daniel transect. Forests Along the Lower Essequibo. Above Kurupukari, the Essequibo passes over a number of rapids and has a braided channel with numerous islands. Below Kurupukari, the river takes on

22 16 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana more of the character of a typical lowland river with meanders, oxbow lakes, and a shifting channel with few rapids or significant drops. The forests along the lower Essequibo are dominated by the successional gradient created by the shifting of the river channel. Here, in addition to the flooding found in mora forests, siltation may also be a significant factor and in such conditions a low stature forest dominated by various Myrtaceae, Symmeria, Coccoloba, Bonafousia, and others occurs as is found along much of the turtle pond transect or the first 0.5 km of the transect that begins on the Essequibo and ends at the top of the cliff at the summit of Turtle Mtn. In areas that remain flooded during most of the year, as along the first 2 km of the Post Arinda Inlet transect, an open Mauritia/Clusia forest occurs. Forests Along the Upper Essequibo. The four transects along the upper Essequibo River (Kuratoka, Pisham Pisham, Akromukro, and Ladysmith) all begin in mora forest and end in either mixed forest on brown sand or, in the case of Kuratoka transect, Eperua (wallaba) forest on white sand. Mora forest and mixed forest intergrade extensively on these transects, with mora forest occurring well away from the river but also having a significant admixture of terra firme elements including Chlorocardium, Eschweilera, and Manilkara. The portion of the Kuratoka transect away from the river is dominated by an oligotrophic Eperua (wallaba) forest. This forest occurs on well-drained white sand and has trees of a lower stature (ca. 20 m) and smaller diameter than mixed forest (hence the name pole forest occasionally attributed to forests of this type). Here, Eperua and other caesalpinioid legumes dominate as does Protium, Clusia, Tovomita, and Citharexylum. That caesalpinioid legumes dominate these forests, both in terms of species diversity and numbers of individuals, as well as most forests of Guyana, for that matter, may perhaps be attributed to their mode of mineral nutrition that allows them to dominate highly weathered, nutrient-poor oxisols and ultisols. Unlike most legumes, which form a symbiotic relationship with nitrogen-fixing Rhizobium bacteria, these trees instead form an association with ectomycorrhizal fungi that are important in phosphorous uptake, in agreement with the general observation that phosphorous and not nitrogen is the limiting nutrient in many stable forest habitats (Raaimakers 1994). Fire also appears to have been an important ecological factor in this area. Anecdotal accounts of a large fire that occurred in the southern part of the Forest are substantiated by areas of semidisturbed liana forest on parts of the Ladysmith and Kuratoka transects. These fires are said to have occurred in the 1940 s. Although naturally caused fires are probably not a natural part of the ecology of Iwokrama and plants in the Forest have not evolved to respond to this type of disturbance, anthropogenic fires are now a very important factor in the Rupununi savanna south of the Forest and in degraded forests north of the Forest. That the forest in Iwokrama is recovering from a large scale fire, albeit slowly, has implications for determining the limits of perturbation from which forests of this type can recover. These limits have clearly been exceeded in forests north of the Iwokrama Forest, as along the Linden Mabura road, and in the forest-savanna ecotone at the edge of the Rupununi Savanna. Forests north of Iwokrama that were once closed canopy Eperua or mixed forest have now more or less permanently been degraded to a Dimorphandra/Humiria scrub forest as a result of logging, which opened up the understory and exposed it to drying effects, followed by repeated burning which eliminated all but the remaining fire-tolerant species. The scrub forest is very recalcitrant to reestablishment of the original closed canopy forest probably because of shifting of the microclimate towards hotter and drier conditions out of the range of tolerance for seedling establishment and formation of mycorrhizal symbioses. Because the forests of Iwokrama are very similar to these forests as they occurred originally, any plan for sustainable timber utilization must take into account both the direct effects of logging and the concomitant effect of increased risk of fire resulting from alteration of the microclimate. Iwokrama Mountains. Collecting efforts were concentrated in the Iwokrama Mountains because these mountains offer high local and overall diversity and because some plants that occur here are not found elsewhere in the Forest. These mountains offer nearly 1000 m of elevational gradient and a moisture gradient that is determined by slope and exposure as well as elevation.

23 BRIT.ORG/SIDA 17 Collections on the highest peaks of the Iwokrama Mountains and in the gorge separating these peaks were very diverse because of the large number of habitats in this area. The 4.5 km approach to these mountains from the Kurupukari Annai road begins in Humiria scrub forest on poorly drained white sand and quickly passes into periodically inundated mora forest and mixed forest on brown sand. Flooding in this area is probably responsible for a somewhat more open forest with a prominent Astrocaryum/Scheelea palm understory and abundant lianas. From the mouth of the gorge, the mountains rise abruptly and the creek in the gorge has a steep drop through the rocky streambed. Mora occurs only along the streambed and becomes progressively less abundant towards the top of the gorge. Chlorocardium is also common in the gorge and on the lower slopes of the mountains, but is not found on the upper slopes. The streambed has characteristic rheophytic elements that exploit higher light levels by being able to withstand inundation and the force of rushing water; such plants include Spigelia multispicata, Gonzalagunia dicocca, and Spathiphyllum cuspidatum. Frequent rockslides and treefalls in the gorge account for the presence of disturbance-associated elements such as Jacaratia, Cecropia, Psychotria poeppigiana, Gurania, and many lianas. Epiphyte diversity and abundance is also high in the gorge perhaps because wetter conditions are caused by reduced exposure to drying in the gorge and its ability to trap moisture. This is certainly the most mesic area of the Forest observed during the course of field work and has a corresponding abundance of epiphytic aroids, ferns, mosses, orchids, and bromeliads. Extensive collections were made on the three peaks that surround the gorge. The composition of the forest on the slopes is similar to that found in mixed forest in the lowland parts of the Forest with several differences in physiognomy (slightly more open forest, more understory palms, lower average diameter of trees) that may be a result of higher frequency of treefalls due to greater exposure to wind action and shallower soils. The summits of these mountains have forest with a relatively low canopy (10 m) and many epiphytes occurring at all levels in the trees. Like the Kanuku and Acarai Mountains in southern Guyana, the Iwokrama Mountains are apparently not high enough to have anything that could be called genuine cloud forest. However, Makarapan Mountain, a 1000 m mountain south and west of the Forest near Annai, was found to have at its summit scrub forest typical of inselbergs (Erythroxylum, Clusia, and Myrtaceae) and an epiphytic element typical of cloud forests (e.g. Utricularia), so it is apparent that local weather patterns and edaphic conditions may be sufficient to alter the composition of summit floras in mountains of this kind. More extensive collecting in the Iwokrama Mountains is needed to fully document the distribution and composition of its plant communities. Although the effort expended in the work described here was considerable, collections from these mountains are still inadequate since they had never been botanically surveyed before and were scarcely explored, as evidenced by the discovery of previously unknown petroglyphs in the Iwokrama gorge in the course of carrying out this work. Forests North of the Kurupukari Annai Road. Most of the forests accessible from the Kurupukari Annai Road, the Burro Burro River, and the Siparuni River are characterized by relatively flat topography. Most of the soils are either brown sands or clays or white sands and are either poorly drained in flat, low lying areas or well drained in areas with even very modest local relief. Transects such as the three mile transect (three miles SW of Kurupukari on the Kurupukari Annai road) have modest local relief that leads to the formation of Euterpe swamp forest in low lying areas and mixed forest in upland areas. Forests in these areas have a relatively high abundance of commercial timber species. This area is particularly valuable for the collection of baseline data concerning the ecology of these species in undisturbed forest. For instance, from what is known of the ecology of Chlorocardium and observations in these areas, it is clear that its patchy distribution in small, pure stands ( greenheart reefs ) is attributable to medium or long term equilibrium of these microsites. Growth of juveniles of greenheart can occur in the shade of mature individuals, but other mixed forest species are usually dependent on canopy gaps for recruitment to occur (Zagt 1997). Chlorocardium would be able to competitively exclude other species in areas that do not experience treefalls for an extended period. Treefalls may occur almost stochastically, but in some places may happen with decreased frequencies due to the

24 18 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana depth of soils, slope or other factors; Chlorocardium may be expected to become a local dominant in any area, whether for stochastic reasons or due to microsite conditions, that has not experienced a treefall for an extended period. Any sort of logging, even selective logging, could be expected to favor the recruitment of species other than greenheart indicating that it may be impossible to harvest Chlorocardium more than once from these forests and that Chlorocardium may soon, at least commercially, be an extinct species. Pakutau Hills, Turtle Mountain, and Sandstone. These three areas are grouped together because they are all low or moderate elevation hilly areas with doleritic substrates. They were investigated with the goal of determining whether they differ significantly in terms of floristic composition from forest on granitic substrates found in the Iwokrama Mountains. Although subtle differences do exist, there were no overwhelming differences found in the mixed, terra firme forests between these two areas. Forests in these areas, do however, provide an added measure of diversity over that found in lowland areas and, as in other areas in southern Guyana, the effort expended in cutting trails to gain access to mountains or any upland area is justified by the diversity of collections these areas offer. CONCLUSION This work is a critical component of baseline data needed to adequately monitor the effects of development activities on the plant community. Continued work is needed, however, to gain a more complete understanding of the flora of Iwokrama. Floristic work in a given area often proceeds by first producing a preliminary checklist, such as the one presented here, followed by an annotated checklist which would provide additional information about the species, and culminating in the production of a flora, a work which would be a complete treatment of all the plants known from that area along with keys to the species, distributions, descriptions, notes on ecology and other pertinent features, and, often, illustrations. Because of the difficulty in obtaining complete collections and accurate determinations and the complexity of resolving synonomies and other taxonomic problems, the production of a flora of a diverse tropical area such as Iwokrama is necessarily a long term project. The flora of La Selva Biological Station in Costa Rica, which at 1500 hectares is a small fraction of the area of the Iwokrama Forest s 360,000 ha, has been ongoing for twenty years and has still not been completed (Hammel 1990; Wilbur 1994). The flora of Central French Guiana (Mori et al. 1997), which at 140,000 ha is an area more comparable to that of Iwokrama, is nearing completion after nearly fifteen years effort. Despite the concentrated effort needed for such an endeavor, this work is essential since questions concerning the nature and dynamics of the plant community cannot be adequately addressed without a comprehensive understanding of the flora. Thus, a flora is useful not only for the community of plant taxonomists, but also for ecologists, foresters, ethnobotanists, pharmacognosists, anthropologists, zoologists, and land use planners. To achieve this goal, more collections will be needed. Collecting efforts should be concentrated in areas that have not been adequately sampled. Although all the existing transects were extensively sampled, these transects give access to but a small fraction of the total area of the Forest. Some suggestions include: areas such as the southern side of the Iwokrama Mtns. and the area between there and the Essequibo River; other portions of the Iworkrama massif accessible from the road other than the two areas reached through the Mt. Daniel transect and the gorge; the upper Siparuni River above Pakutau Falls and Ireng Creek which have not been visited by any research team and which show a distinctive vegetation type in satellite photos; and more comprehensive sampling of unique habitats found in the course of collecting such as, the inselberg vegetation on the summit of Turtle Mountain, the xeric and rheophytic vegetation of sand islands in the Essequibo above Kurupukari, the Mauritia swamp forest on the Post Arinda transect, and the natural and undisturbed Humiria scrub habitat at the terminus of the western extension of the C D transect. It is perhaps necessary to justify the not inconsiderable effort expended to date to document the plant diversity of Iwokrama and to substantiate the need for more work of this type. Plant

25 BRIT.ORG/SIDA 19 collecting in Iwokrama is time-consuming, arduous, demanding of a certain level of skill, both in the field and in follow-up herbarium work, expensive, and even dangerous. This work, however, is a prerequisite for any quantitative or qualitative work associated with the plant diversity of the forest. The ability to pursue floristic research in a pristine area such as the Iwokrama Forest is a rare and irreplaceable opportunity to collect baseline data prior to human disturbance. ACKNOWLEDGMENTS We wish to thank the Macushi, Wapisiana, and Arawak Amerindians of the communities surrounding Iwokrama, particularly Romeo Williams, Claudius Perry, Milner Captain, Daniel Allicock, and Rodrigues Antone, without whose help the field portion of this work would not have been possible. Staff and faculty of the University of Guyana and the Centre for the Study of Biological Diversity on the campus of the University of Guyana, particularly Marlene Cox, Indarjit Ramdass, John Caesar, Phillip da Silva, and Dyantie Naraine, provided assistance with permits. Special thanks are due to the curators and staff of numerous herbaria for the determination of specimens; without their many years of accumulated effort in working with the Guyana flora this work would not have been possible. Scott Mori gave permission to use the checklist of the Flora of Central French Guiana. Mike Hopkins and Ghillean Prance gave similar permission to use data from the Flora of Reserva Ducke as did Renske Ek for the checklist of the Mabura Hill area. Particular gratitude is extended to Carol Kelloff for many intercessions on behalf of the authors. Vijay Datadin of Iwokrama and Naseem Nasir of the University of Guyana provided GIS coverages for the base maps. Peter Jørgensen and Toby Pennington provided helpful comments on the manuscript. Funding was provided by the United Nations Development Programme through the Iwokrama International Centre for Rainforest Conservation and Development, the Biological Diversity of the Guianas Program of the Smithsonian Institution, and in the form of a postdoctoral fellowship to HDC from the Office of Biodiversity Programs of the Smithsonian Institution. This is publication No. 46 in the Biological Diversity of the Guianas publication series. REFERENCES BOGGAN, J., V. FUNK, C. KELLOFF, M. HOFF, G. CREMERS, and C. FEUILLET Checklist of the plants of the Guianas (Guyana, Suriname, French Guiana). 2nd Edition. Biological Diversity of the Guianas Program, Department of Botany, National Museum of Natural History, Smithsonian Institution, Washington, DC. CLARKE, H.D. and V. FUNK. Submitted. Using Checklists and Collections Data to Investigate Plant Diversity. II. An analysis of five florulas from northeastern South America. Ann. Missouri Bot. Gard. DAVIS, T.A.W. and P.W. RICHARDS The vegetation of Moraballi Creek, British Guiana. An ecological study of a limited area of tropical rain forest I. J. Ecol. 21: DAVIS, T.A.W. and P.W. RICHARDS The vegetation of Moraballi Creek, British Guiana. An ecological study of a limited area of tropical rain forest II. J. Ecol. 22: EDEN, M.J The savanna ecosystem northern Rupununi, British Guiana. McGill University Savanna Research Project, Savanna Research Series No. 1. Dept. of Geography, McGill University, Montreal, Canada. EDEN, M.J The savanna environment, Guyana, I. Some aspects of weathering and landforms. II. Savanna vegetation in the northern Rupununi. McGill University Savanna Research Project, Savanna Research Series No. 17. Dept. of Geography, McGill University, Montreal, Canada. EK, R.C Index of Guyana plant collectors. In: A.R.A. Görts-van Rijn, (ed.). Flora of the Guianas. Supplementary series 1. Koeltz Scientific Books, Koenigstein, Germany. EK, R.C. and H. TER STEEGE The flora of the Mabura Hill area, Guyana. In: R.C. Ek. Botanical diversity in the tropical rainforest of Guyana. Tropenbos-Guyana Series 4. Pp Tropenbos-Guyana Programme, Georgetown, Guyana. FROST, D.B The climate of the rupununi savannas. McGill University Savanna Research Project, Savanna Research Series No. 12 and Climatological Research Series No. 3. Dept. of Geography, McGill University, Montreal, Canada. GENTRY, A.H A field guide to the families and genera of woody plants of northwest South America (Colombia, Ecuador, Peru) with supplementary notes on herbaceous taxa. University of Chicago Press, Chicago, IL. GUYANA FOREST DEPARTMENT AND FOOD AND AGRICULTURE ORGANIZATION OF THE UNITED NATIONS Forest type map of Guyana. Government of Guyana and United Nations Development Programme. Georgetown, Guyana.

26 20 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana GUYANA INTER-AGENCY COMMITTEE Iwokrama Rainforest Programme synopsis. Guyana, Guyana Natural Resources Agency. HAMMEL, B The distribution of diversity among families, genera, and habit types in the La Selva Flora. In: A. H. Gentry, ed. Four neotropical rainforests. Yale University Press, New Haven, CT. HAWKES, M.D. and J.R.D. Wall The Commonwealth and Government of Guyana Iwokrama Rain Forest Programme, Phase I, site resource survey, main report. Natural Resources Institute, Overseas Development Administration, Chatham, UK. HUBER, O., G. GHARBARRAN, and V. FUNK Vegetation map of Guyana. Centre for the Study of Biological Diversity, University of Guyana, Georgetown, Guyana. KELLOFF, C.L. and V.A. Funk Preliminary checklist of the plants of Kaieteur Falls National Park, Guyana. Biological Diversity of the Guianas Program, Department of Botany, National Museum of Natural History, Smithsonian Institution, Washington, D.C. KHAN, Z., S. PAUL, and D. CUMMINGS Mabura Hill, upper Demerara forestry project. Soils investigation report no. 1. National Agricultural Research Institute, Mon Repos, Guyana. MORI, S., G. CREMERS, C. GRACIE, J.-J. DE GRANVILLE, M. HOFF, and J.D. MITCHELL Guide to the vascular plants of central French Guiana. Pt. 1. pteridophytes, gymnosperms, and monocotyledons. Mem. New York Bot. Gard. 76(1): OLIVEIRA, A.A. DE and D.C. DALY Geographic distribution of tree species occurring in the region of Manaus, Brazil: Implications for regional diversity and conservation. Biodiversity and Conservation 8: OLIVEIRA, A.A. DE and S.A. MORI A central Amazonian terra firme forest. I. High tree species richness on poor soils. Biodiversity and Conservation 8: RAAIMAKERS, D Growth of tropical rain forest trees as dependent on phosphorous supply. Tropenbos Series 11. The Tropenbos Foundation, Wageningen, The Netherlands. RIBEIRO, J.E.L. DA S., M.J.G. HOPKINS, A. VICENTINI, C.A. SOTHERS, M.A. DA S. COSTA, J.M. DE BRITO, M.A.D. DE SOUZA, L.H.P. MARTINS, L.G. LOHMANN, P.A.C.L. ASSUNÇÃO, E. DA C. Pereira, C.F. DA SILVA, M.R. MESQUITA, L.C. Procópio Flora da Reserva Ducke: Guia de identificação das plantas vasculares de uma floresta de terrafirme na Amazônia Central. Instituto Nacional de Pesquisas da Amazônia, Manaus, Brazil. RUTHERFORD, K.G. and T.L. HILLS Soils of the Rupununi savanna/forest ecotone southern Guyana. McGill University Savanna Research Project, Savanna Research Series No. 21. Dept. of Geography, McGill University, Montreal, Canada. SINHA, N.K.P Geomorphic evolution of the northern Rupununi basin, Guyana. McGill University Savanna Research Project, Savanna Research Series No. 11. Dept. of Geography, McGill University, Montreal, Canada. SYSTEMATICS AGENDA Systematics Agenda 2000: Charting the biosphere. Technical report. Produced by Systematics Agenda TER STEEGE, H., V.G. JETTEN, M. POLAK, and M.J.A. WERGER Tropical rainforest types and soils of a watershed in Guyana. J. Veg. Sci. 4: TER STEEGE, H., R.G.A. BOOT, L.C. BROUWER, J.C. CAESAR, R.C. EK, D.S. HAMMOND, P.P. HARIPERSAUD, P. VAN DER HOOT, V.G. JETTEN, A.J. VAN KEKEM, M.A. KELLMAN, Z. KHAN, A.M. POLAK, T.L. PONS, J. PULLES, D. RAAIMAKERS, S.A. ROSE, J.J. VAN DER SANDEN, and R. ZAGT Ecology and logging in a tropical rainforest in Guyana: with recommendations for forest management. The Tropenbos Foundation, Wageningen, The Netherlands. WILBUR, R Vascular plants: an interim checklist. In: L.A. McDade, S.B. Kamaljit, H.A. Hespenheide, and G.S. Hartshorn, eds. La Selva: Ecology and natural history of a neotropical rainforest. University of Chicago Press, Chicago. ZAGT, R.J Tree demography in the tropical rain forest of Guyana. Tropenbos-Guyana Series 3. Tropenbos- Guyana Programme, Georgetown, Guyana. APPENDIX 1 CHECKLIST OF PLANT SPECIES FROM IWOKRAMA AND FOUR OTHER SITES IN NORTHEAST SOUTH AMERICA Acanthaceae Aphelandra aurantiaca (Schweidw.) Lindl., S Beloperone violacea Planch. & Linden, M Blechum pyramidatum (Lam.) Urb., S Chaetochlamys ciliata Lindau, S Gynocraterium guianense Bremek., S Herpetacanthus rotundus (Lindau) Bremek., S Justicia calycina (Nees) V.A.W. Graham, I Justicia cayennensis (Nees) Lindau, S Justicia comata (L.) Lam., D Justicia pectoralis Jacq., I, S Justicia potarensis (Bremek.) Wassh., S Justicia schomburgkiana (Nees) V.A.W. Graham, I Justicia secunda Vahl, S Justicia sprucei V.A.W. Graham, S Lepidagathis alopecuroidea (Vahl) R. Br. ex Griseb., K, M, S, D Odontonema schomburgkianum (Nees) Kuntze, K, M Pachystachys coccinea (Aubl.) Nees, S Polylychnis fulgens Bremek., S Pulchranthus congestus (Lindau) V.M. Baum, Reveal & Nowicke, S, D I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

27 BRIT.ORG/SIDA 21 Pulchranthus variegatus (Aubl.) V.M. Baum, Reveal & Nowicke, S Rhacodiscus sp., S Ruellia geminiflora Kunth, I Ruellia rubra Aubl., M, S Ruellia saülensis Wassh., S Ruellia sprucei Lindau, D Trichanthera gigantea (Bonpl.) Nees, I Adiantaceae Adiantopsis monticola (Gardner) T. Moore, K Adiantopsis radiata (L.) Fée, I, S Adiantum adiantoides (J. Sm.) C. Chr., I, S Adiantum amazonicam A.R.Sm., D Adiantum cajennense Willd. ex Klotzsch, I, S, D Adiantum cordatum Maxon, S Adiantum dolosum Kunze, S Adiantum fructuosum Poepp. ex Spreng., S Adiantum fuliginosum Fée, S Adiantum glaucescens Klotzsch, S Adiantum latifolium Lam., I, S Adiantum latifolium x serratodentatum, I Adiantum leprieurii Hook., S Adiantum lucidum (Cav.) Sw., S Adiantum macrophyllum Sw., S Adiantum multisorum A. Samp., D Adiantum obliquum Willd., S Adiantum olivaceum Baker, I Adiantum petiolatum Desv., S Adiantum phyllitidis J. Sm., I Adiantum pulverulentum L., S Adiantum rondonii Samp., I Adiantum serratodentatum Humb. & Bonpl. ex Willd., S Adiantum terminatum Kunze ex Miq., I, K, S Adiantum tomentosum Klotzsch, I, S, D Adiantum sp., M Doryopteris sagittifolia (Raddi) J. Sm., S Hemionitis rufa (L.) Sw., S Hemionitis sp., S Pityrogramma calomelanos (L.) Link, I, M, S, D Alismataceae Echinodorus subalatus (Mart.) Griseb., M Amaranthaceae Alternanthera brasiliana (L.) Kuntze, S Alternanthera sessilis (L.) DC., S, D Alternanthera tenella Colla, S Pfaffia glomerata (Spreng.) Pedersen, S Anacardiaceae Anacardium giganteum W. Hancock ex Engl., K, M Anacardium occidentale L., M, S Anacardium parvifolium Ducke, D Anacardium spruceanum Benth. ex Engl., S, D Astronium lecointei Ducke, D Astronium ulei Mattick, S Campnosperma gummifera (Benth.) Marchand, D Loxopterygium sagotii Hook.f., M Spondias mombin L., I, S Tapirira bethamiana J.D. Mitch., S Tapirira guianensis Aubl., I, K, M, S, D Tapirira obtusa (Benth.) J.D. Mitch., M, S, D Tapirira retusa Ducke, D Thyrsodium guianense Sagot ex Marchand, I, S Thyrsodium puberulum J.D. Mitch. & Daly, S Thyrsodium spruceanum Salzm. ex Benth., S, D Anisophylleaceae Anisophyllea manausensis Pires & W.A.Rodrigues, D Annonaceae Anaxagorea brevipes Benth., D Anaxagorea dolichocarpa Sprague & Sandwith, I, M, S Anaxagorea manausensis Timmerman, D Anaxagorea phaeocarpa Mart., D Annona amazonica R.E. Fr., D Annona ambotay Aubl., S, D Annona foetida Mart., D Annona glabra L., K Annona haematantha Miq., I, M, S, D Annona hypoglauca Mart., I Annona impressivenia Saff. ex R.E. Fr., D Annona montana Macfad., I Annona muricata L., S Annona nitida Mart., D Annona paludosa Aubl., S Annona sericea Dunal, I, M Annona symphyocarpa Sandwith, I, K, M Annona sp. nov. aff. excellens R.E. Fr., D Bocageopsis mattogrossensis (R.E. Fr.) R.E. Fr., D Bocageopsis multiflora (Mart.) R.E. Fr., I, M, D Bocageopsis pleiosperma Maas, D Cardiopetalum surinamense R.E. Fr., S Cremastosperma brevipes (DC. ex Dunal) R.E. Fr., S Cymbopetalum brasiliense (Vell.) Benth., I, M, S Cymbopetalum euneurum N.A.Murray, D Diclinanona calycina (Diels) R.E. Fr., D Duguetia arenicola Maas, D Duguetia asterotricha (Diels) R.E. Fr., D Duguetia cadaverica Huber, M, S Duguetia calycina Benoist, I, K, M, S, D Duguetia eximia Diels, S Duguetia flagellaris Huber, D Duguetia inconspicua Sagot, I, M, S Duguetia insculpta R.E. Fr., D Duguetia megalocarpa Maas, D Duguetia neglecta Sandwith, I, M Duguetia obovata R.E. Fr., I Duguetia paraensis R.E. Fr., I Duguetia pycnastera Sandwith, K, M, S, D Duguetia quitarensis Benth., I Duguetia riparia Huber, S, D Duguetia stelechantha (Diels) R.E. Fr., D Duguetia surinamensis R.E. Fr., I, S, D Duguetia trunciflora Maas & A.H. Gentry, D Duguetia ulei (Diels) R.E. Fr., D Duguetia sp., S Ephedranthus amazonicus R.E. Fr., D Ephedranthus guianensis R.E. Fr., M Fusaea longifolia (Aubl.) Saff., I, M, D Guatteria atra Sandwith, M Guatteria blepharophylla Mart., S, D Guatteria citriodora Ducke, I, D Guatteria conspicua R.E. Fr., K Guatteria discolor R.E. Fr., D Guatteria foliosa Benth., D I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

28 22 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana Guatteria guianensis (Aubl.) R.E. Fr., D Guatteria megalophylla Diels, D Guatteria meliodora R.E. Fr., D Guatteria olivacea R.E. Fr., D Guatteria procera R.E. Fr., D Guatteria punctata (Aubl.) R.A. Howard, M, S Guatteria scandens Ducke, I Guatteria schomburgkiana Mart., I, M Guatteria scytophylla Diels, D Guatteria umbonata R.E. Fr., K Guatteriopsis friesiana W.A. Rodrigues, D Guatteriopsis hispida R.E.Fr., D Malmea manausensis Maas & Miralha, D Oxandra asbeckii (Pulle) R.E. Fr., M, S Oxandra guianensis R.E. Fr., I, M Oxandra xylopioides Diels, I Pseudoxandra coriacea R.E. Fr., D Pseudoxandra cuspidata Maas, S Pseudoxandra sp., I Rollinia cuspidata Mart., S Rollinia elliptica R.E. Fr., S Rollinia exsucca (DC. ex Dunal) A. DC., I, K, M, S Rollinia insignis R.E. Fr., D Tetrameranthus duckei R.E. Fr., D Trigynaea caudata (R.E. Fr.) R.E. Fr., I, M Trigynaea duckei (R.E. Fr.) R.E. Fr., I, M Unonopsis duckei R.E. Fr., D Unonopsis glaucopetala R.E. Fr., M Unonopsis guatterioides (A. DC.) R.E. Fr., I, M, D Unonopsis perrottetii (A. DC.) R.E. Fr., M, S Unonopsis rufescens (Baill.) R.E. Fr., I, M, S Unonopsis stipitata Diels, I, M, S, D Xylopia amazonica R.E. Fr., D Xylopia aromatica (Lam.) Mart., I, M Xylopia benthamii R.E. Fr., I, K, M, D Xylopia calophylla R.E. Fr., D Xylopia cayennensis Maas, S Xylopia crinita R.E. Fr., D Xylopia discreta (L.f.) Sprague & Hutch., I Xylopia emarginata Mart., D Xylopia frutescens Aubl., S Xylopia nitida Dunal, I, M, S, D Xylopia parviflora Spruce, D Xylopia polyantha R.E. Fr., D Xylopia pulcherrima Sandwith, M Xylopia sericea A. St.-Hil., D Xylopia spruceana Benth. ex Spruce, D Apiaceae Eryngium foetidum L., S Apocynaceae Allamanda cathartica L., I, M Ambelania acida Aubl., I, M, S, D Ambelania duckei Markgr., D Aspidosperma album (Vahl) Benoist ex Pichon, M, S Aspidosperma araracanga Marc. Ferr., D Aspidosperma carapanauba Pichon, S Aspidosperma cruentum Woodson, M, S Aspidosperma desmanthum Benth. ex Müll. Arg., D Aspidosperma excelsum Benth., I, M, S Aspidosperma macrophyllum Müll. Arg., S Aspidosperma marcgravianum Woodson, S, D Aspidosperma nitidum Benth. ex Müll. Arg., D Aspidosperma oblongum A. DC., S Aspidosperma parvifolium A. DC., D Aspidosperma sandwithianum Markgr., M, S, D Aspidosperma schultesii Woodson, S, D Aspidosperma spruceanum Benth. ex Müll. Arg., I, S, D Aspidosperma vargasii A. DC., I, M Condylocarpon amazonicum (Markgr.) Ducke, S Condylocarpon pubiflorum Müll. Arg., S Couma catingae Ducke, K Couma guianensis Aubl., I, D Couma rigida Müll. Arg., K Couma utilis (Mart.) Müll. Arg., D Forsteronia acouci (Aubl.) A. DC., M, S, D Forsteronia gracilis (Benth.) Müll. Arg., M Forsteronia guyanensis Müll. Arg., I, K, M, S Forsteronia schomburgkii (Benth.) Müll. Arg., K, M Geissospermum argenteum Woodson, S, D Geissospermum laevis (Thunb.) Miers, S Geissospermum sericeum Benth. & Hook.f. ex Miers, M, S Geissospermum urceolatum A.H. Gentry, D Himatanthus articulatus (Vahl) Woodson, K, M, S Himatanthus articulatus x bracteatus, M Himatanthus bracteatus (A. DC.) Woodson, K, D Himatanthus speciosus (Müll. Arg.) Plumel, S Himatanthus stenophyllus Plumel, D Himatanthus sucuuba (Spruce ex Müll. Arg.) Woodson, M, D Lacmellea aculeata (Ducke) Monach., I, K, S Lacmellea arborescens (Müll. Arg.) Markgr., D Lacmellea gracilis (Müll.Arg.) Markgr., D Lacmellea utilis (Arn.) Markgr., M Laxoplumeria baehniana Monach., S Macoubea guianensis Aubl., M, S Macoubea sprucei (Müll. Arg.) Markgr., D Macropharynx spectabilis (Stadelm.) Woodson, S Malouetia gracilis (Benth.) A. DC., I, M Malouetia tamaquarina (Aubl.) A. DC., I, M Mandevilla benthamii (A. DC.) K. Schum., K Mandevilla hirsuta (Rich.) K. Schum., I, K Mandevilla rugellosa (Rich.) L. Allorge, S Mandevilla scabra (Hoffmanns. ex Roem. & Schult.) K. Schum., I, D Mandevilla subcarnosa (Benth.) Woodson, K Mandevilla trianae Woodson, K Mandevilla vanheurckii (Müll. Arg.) Markgr., K Mesechites trifida (Jacq.) Müll. Arg., I, K Odontadenia geminata (Hoffmanns. ex Roem. & Schult.) Müll. Arg., M Odontadenia macrantha (Roem. & Schult.) Markgr., I Odontadenia nitida (Vahl) Müll. Arg., I, K Odontadenia perrottetii (A. DC.) Woodson, M, S, D Odontadenia puncticulosa (Rich.) Pulle, K, M, S, D Odontadenia verrucosa (Willd. ex Roem. & Schult.) K. Schum. ex Markgr., M, D Pacouria guianensis Aubl., S Parahancornia fasciculata (Poir.) Benoist, K, M, D Prestonia acutifolia (Benth. ex Müll. Arg.) K. Schum., I, K Prestonia annularis (L.f.) G. Don, K Prestonia cayennensis (A. DC.) Pichon, K Prestonia coalita (Vell.) Woodson, I, K I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

29 BRIT.ORG/SIDA 23 Prestonia lindleyana Woodson ex Gleason & A.C. Sm., K Prestonia megagros (Vell.) Woodson, D Prestonia perplexa Woodson, K Prestonia surinamensis Müll. Arg., M Prestonia tomentosa R. Br., I Rauvolfia sprucei Müll. Arg., D Rhigospira quadrangularis (Müll. Arg.) Miers, D Secondatia duckei Markgr., D Tabernaemontana angulata Mart. ex Müll. Arg., S, D Tabernaemontana attenuata (Miers) Urb., M Tabernaemontana cerea (Woodson) Leeuwenb., K, M Tabernaemontana disticha A. DC., K, M, S Tabernaemontana divaricata (L.) R. Br. ex Roem. & Schult., S Tabernaemontana flavicans Willd. ex Roem. & Schult., S, D Tabernaemontana heterophylla Vahl, I, D Tabernaemontana lorifera (Miers) Leeuwenb., I Tabernaemontana macrocalyx Müll. Arg., K, S, D Tabernaemontana muricata Link ex Roem. & Schult., D Tabernaemontana riedelii Müll. Arg., D Tabernaemontana rupicola Benth., K Tabernaemontana sananho Ruiz & Pav., S Tabernaemontana siphilitica (L.f.) Leeuwenb., I, S Tabernaemontana undulata Vahl, I, M, D Aquifoliaceae Ilex fanshawei Edwin, K Ilex jenmanii Loes., K Ilex martiniana D. Don, K, M Ilex soderstromii Edwin, K Ilex sp. 1, D Araceae Anaphyllopsis americana (Engl.) A. Hay, S Anthurium bonplandii G.S. Bunting, D Anthurium clavigerum Poepp., S Anthurium cowanii Croat, K Anthurium crassinervium (Jacq.) Schott, K, S Anthurium eminens Schott, D Anthurium expansum Gleason, I, K Anthurium gracile (Rudge) Schott, I, M, S, D Anthurium hookeri Kunth, K, M, S Anthurium jenmanii Engl., K, M, S Anthurium kunthii Poepp., K Anthurium pentaphyllum (Aubl.) G. Don, K, S, D Anthurium rubrinervium (Link) G. Don, S Anthurium scandens (Aubl.) Engl., S Anthurium sinuatum Benth. ex Schott, D Anthurium thrinax Madison, I, S Anthurium trinerve Miq., I, K, M, S, D Caladium bicolor (Aiton) Vent., I, S Dieffenbachia elegans A.M.E. Jonker & Jonker, S, D Dieffenbachia humilis Poepp., S Dieffenbachia paludicola N.E. Br. ex Gleason, M, S Dieffenbachia seguine (Jacq.) Schott, S Dracontium longipes Engl., D Dracontium polyphyllum L., S Heteropsis flexuosa (Kunth) G.S. Bunting, I, M, S, D Heteropsis macrophylla A.C. Sm., D Heteropsis melinonii (Engl.) A.M.E. Jonker & Jonker, I Heteropsis riedeliana Schott, D Heteropsis spruceana Schott, I, S, D Heteropsis steyermarkii G.S. Bunting, D Heteropsis tenuispadix G.S. Bunting, D Monstera adansonii Schott, I, S Monstera dubia (Kunth) Engl. & K. Krause, M, S Monstera obliqua Miq., I, S, D Monstera spruceana (Schott) Engl., M, S Monstera sp., S Montrichardia arborescens (L.) Schott, I, K, M, D Montrichardia linifera (Arruda) Schott, I Philodendron asplundii Croat ined., D Philodendron barrosoanum G.S. Bunting, D Philodendron billietiae Croat, S, D Philodendron brevispathum Schott, D Philodendron callosum K. Krause, K, M, S, D Philodendron camposportoanum G.M. Barroso, I Philodendron cremersii Croat & Grayum, S Philodendron cuneatum Engl., D Philodendron deflexum Poepp. ex Schott, I, S, D Philodendron distantilobum K. Krause, D Philodendron duckei Croat & Grayum, S Philodendron ecordatum Schott, M Philodendron elaphoglossoides Schott, D Philodendron fragrantissimum (Hook.) G. Don, I, K, M, S, D Philodendron goeldii G.M. Barroso, S, D Philodendron grandifolium (Jacq.) Schott, S Philodendron guianense Croat & Grayum, S Philodendron hederaceum (Jacq.) Schott, S Philodendron hylaeae G.S. Bunting, I, D Philodendron insigne Schott, I, K, M, S, D Philodendron jenmanii K. Krause, I Philodendron linnaei Kunth, I, K, M, S, D Philodendron melinonii Brongn. ex Regel, S, D Philodendron muricatum Willd. ex Schott, I Philodendron ornatum Schott, I, K, S, D Philodendron pedatum (Hook.) Kunth, I, K, M, S, D Philodendron placidum Schott, I, S Philodendron platypodum Gleason, D Philodendron pulchellum Engl., D Philodendron pulchrum G.M. Barroso, D Philodendron quinquelobum K. Krause, D Philodendron rudgeanum Schott, I, M, S Philodendron scandens K. Koch & Sello, M, S Philodendron solimoesense A.C. Sm., I, M, S Philodendron sphalerum Schott, S, D Philodendron squamiferum Poepp., S Philodendron surinamense (Miq.) Engl., I, M, S, D Philodendron wittianum Engl., S Pistia stratiotes L., K Rhodospatha brachypoda G.S. Bunting, S Rhodospatha latifolia Poepp., S Rhodospatha oblongata Poepp., I, S Rhodospatha venosa Gleason, M, S, D Schismatoglottis spruceana (Schott) G.S. Bunting, D Spathiphyllum cuspidatum Schott, I, K, M Spathiphyllum humboldtii Schott, S Spathiphyllum maguirei G.S. Bunting, D Stenospermation maguirei A.M.E. Jonker & Jonker, I, K, M Stenospermation multiovulatum (Engl.) N.E. Br., S Stenospermation spruceanum Schott, K I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

30 24 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana Syngonium podophyllum Schott, I, S Urospatha sagittifolia (Rudge) Schott, I, K, M, D Xanthosoma granvillei Croat & S.A. Thomps., S Xanthosoma striolatum Mart. ex Schott, S Xanthosoma violaceum Schott, S Araliaceae Dendropanax sp., M Oreopanax capitatus (Jacq.) Decne. & Planch., S Schefflera decaphylla (Seem.) Harms, I, M, S Schefflera morototoni (Aubl.) Maguire, Steyerm. & Frodin, S, D Schefflera umbrosa Frodin ined., D Arecaceae Astrocaryum aculeatum G. Mey., D Astrocaryum gynacanthum Mart., I, M, S, D Astrocaryum murumuru Mart., S, D Astrocaryum paramaca Mart., S Astrocaryum sciophilum (Miq.) Pulle, I, S, D Bactris acanthocarpa Mart., M, S, D Bactris acanthocarpoides Barb. Rodr., S, D Bactris aubletiana Trail, S Bactris campestris Poepp. ex Mart., K Bactris constanciae Barb. Rodr., D Bactris corossilla H. Karst., I, K Bactris elegans Barb. Rodr., I, S, D Bactris floccosa Spruce, S Bactris gasipaes Kunth, S Bactris gastoniana Barb. Rodr., S, D Bactris hirta Mart., K, S, D Bactris maraja Mart., S, D Bactris oligocarpa Barb. Rodr. & Trail ex Barb. Rodr., S Bactris oligoclada Burret, I, K, M Bactris pliniana Granv. & An. Hend., S Bactris ptariana Steyerm., I, K, M, D Bactris rhaphidacantha Wess. Boer, S Bactris simplicifrons Mart., I, K, S, D Bactris syagroides Trail, D Bactris tomentosa Mart., D Bactris trailiana Barb. Rodr., D Desmoncus macroacanthos Mart., K, M, S Desmoncus orthacanthos Mart., I Desmoncus phoenicocarpus Barb. Rodr., K Desmoncus polyacanthos Mart., D Dictyocaryum ptarianum (Steyerm.) H.E. Moore & Steyerm., K Euterpe catinga Wallace, D Euterpe oleracea Mart., I, M, S Euterpe precatoria Mart., S, D Geonoma aspidiifolia Spruce, D Geonoma baculifera (Poit.) Kunth, I, S Geonoma deversa (Poit.) Kunth, I, S, D Geonoma euspatha Burret, I, S Geonoma maxima (Poit.) Kunth, I, M, S, D Geonoma stricta (Poit.) Kunth, S, D Geonoma umbraculiformis Wess. Boer, S Geonoma sp., K Hyospathe elegans Mart., S, D Iriartella setigera (Mart.) H. Wendl., D Jessenia-Oenocarpus complex, K Manicaria saccifera Gaertn., D Mauritia flexuosa L.f., M, D Mauritiella aculeata (Kunth) Burret, K, D Mauritiella martiana (Spruce) Burret, I Maximiliana maripa (Aubl.) Drude, I, M, D Oenocarpus bacaba Mart., I, M, S, D Oenocarpus bataua Mart., I, M, S, D Oenocarpus minor Mart., D Oenocarpus sp., K Orbignya sagottii Trail ex Thurn, I, D Scheelea sp., S, D Socratea exorrhiza (Mart.) H. Wendl., I, S, D Syagrus inajai (Spruce) Becc., S, D Aristolochiaceae Aristolochia bukuti Poncy, S Aristolochia consimilis Mast., M Aristolochia cremersii Poncy, S Aristolochia daemoninoxia Mast., M Aristolochia didyma S. Moore, S Aristolochia iquitensis O.C. Schmidt, I, S Aristolochia rugosa Lam., M Aristolochia stahelii O.C. Schmidt, S Aristolochia surinamensis Willd., I Aristolochia trilobata L., S Aristolochia weddellii Duch., M Aristolochia sp., D Asclepiadaceae Asclepias curassavica L., K, S Blepharodon nitidus (Vell.) J.F. Macbr., K, M Matelea badilloi Morillo, D Matelea gracieae Morillo, S Matelea palustris Aubl., K, M, S Matelea stenopetala Sandwith, I, K, M Metalepis albiflora Urb., S Tassadia guianensis Decne., I Tassadia propinqua Decne., K, M Aspleniaceae Asplenium abscissum Willd., S Asplenium angustum Sw., I, M, S, D Asplenium auriculatum Sw., I Asplenium auritum Sw., M, S Asplenium cristatum Lam., I Asplenium cuneatum Lam., K Asplenium hostmannii Hieron., S Asplenium juglandifolium Lam., S, D Asplenium laetum Sw., S Asplenium pedicularifolium A. St.-Hil., S Asplenium rutaceum (Willd.) Mett., S Asplenium salicifolium L., K, D Asplenium serratum L., I, K, M, S, D Asplenium zamiifolium Willd., K Asteraceae Acanthospermum australe (Loefl.) Kuntze, K, M Acmella brachyglossa Cass., S Acmella uliginosa (Sw.) Cass., S Bidens cynapiifolia Kunth, K, M, S Bidens pilosa L., S Calea caleoides (DC.) H. Rob., I, M Calea lucidivenia Gleason & S. F. Blake, K Calea oliveri B.L. Rob. & Greenm., K Calea solidaginea Kunth, I Centratherum punctatum Cass., I I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

31 BRIT.ORG/SIDA 25 Centratherum sp. 1, D Chromolaena odorata (L.) R.M. King & H. Rob., M, S Chromolaena squalida (DC.) R.M. King & H. Rob., I Clibadium surinamense L., I, K Clibadium sylvestre (Aubl.) Baill., I, M, S Conyza bonariensis (L.) Cronquist, M Conyza floribunda Kunth, D Cyanthillium cinereum (L.) H. Rob., M, S Eclipta prostrata (L.) L., I, S Elephantopus mollis Kunth, K, S, D Eleutheranthera ruderalis (Sw.) Sch. Bip., I Emilia fosbergii Nicolson, K Emilia sonchifolia (L.) DC. ex Wight, K, M, S Erechtites hieracifolia (L.) Raf. ex DC., K, M, S, D Eupatorium sp. 1, D Gnaphalium polycaulon Pers., I Hebeclinium macrophyllum (L.) DC., S Lepidaploa cotoneaster (Willd. ex Spreng.) H. Rob., K Lepidaploa gracilis (Kunth) H. Rob., I, K, M Lepidaploa remotiflora (Rich.) H. Rob., K, M Mikania banisteriae DC., K Mikania congesta DC., I Mikania gleasonii B.L. Rob., I, M, S Mikania guaco Bonpl., S Mikania hookeriana DC., I, K, M Mikania lindleyana DC., I Mikania microptera DC., S Mikania parviflora (Aubl.) H. Karst., I, K, S Mikania psilostachya DC., I, K, M, D Mikania sprucei Baker, K Mikania trinitaria DC., I Piptocarpha triflora (Aubl.) Benn. ex Baker, I Porophyllum ruderale (Jacq.) Cass., M Pseudelephantopus funckii (Turcz.) Philipson, D Rolandra fruticosa (L.) Kuntze, I, K, S, D Sphagneticola trilobata (L.) Pruski, S Spilanthes acmella (L.) Murray, D Synedrella nodiflora (L.) Gaertn., S Trichospira verticillata (L.) S.F. Blake, I Unxia camphorata L.f., K, M Wedelia paludosa DC., D Wulffia baccata (L.) Kuntze, I, K, M, S Balanophoraceae Helosis cayennensis (Sw.) Spreng., I, K, S Begoniaceae Begonia fischeri Schrank, K Begonia glabra Aubl., S Begonia guaduensis Kunth, I Begonia heloisana Brade, I Begonia humilis Dryand., M, S Begonia jenmanii Tutin, K Begonia prieurii DC., K, S Bignoniaceae Adenocalymna impressum (Rusby) Sandwith, D Adenocalymna inundatum Mart. ex DC., I Adenocalymna saulense C.C. Berg & Kooy, S Adenocalymna subincanum Huber, D Amphilophium paniculatum (L.) Kunth, S Anemopaegma chrysoleucum (Kunth) Sandwith, I Anemopaegma floridum Mart. ex DC., D Anemopaegma foetidum Bureau & K. Schum., M, D Anemopaegma ionanthum A.H. Gentry, S, D Anemopaegma oligoneuron (Sprague & Sandwith) A.H. Gentry, M, S Anemopaegma parkeri Sprague, M Anemopaegma robustum Bureau & K. Schum., K, D Arrabidaea bilabiata (Sprague) Sandwith, I Arrabidaea candicans (Rich.) DC., I, K, M, S Arrabidaea chica (Bonpl.) B. Verl., D Arrabidaea cinnamomea (A. DC.) Sandwith, S, D Arrabidaea egensis Bureau & K. Schum., M Arrabidaea fanshawei Sandwith, M, S, D Arrabidaea florida DC., I, S Arrabidaea inaequalis (DC. ex Splitg.) K. Schum., I, S, D Arrabidaea japurensis (DC.) Bureau & K. Schum., S Arrabidaea mollis (Vahl) Bureau ex K. Schum., M, S Arrabidaea nigrescens Sandwith, S, D Arrabidaea oligantha Bureau & K. Schum., S Arrabidaea patellifera (Schltdl.) Sandwith, M, S Arrabidaea prancei A.H. Gentry, D Arrabidaea trailii Sprague, M, S, D Arrabidaea triplinervia (Mart. ex DC.) Baill. ex Bureau, S, D Callichlamys latifolia (Rich.) K. Schum., M, S, D Clytostoma binatum (Thunb.) Sandwith, I, K, M, S Clytostoma sciuripabulum Bureau & K. Schum., M Cydista aequinoctialis (L.) Miers, I, M, S, D Digomphia densicoma (Mart. ex DC.) Pilg., K Distictella elongata (Vahl) Urb., I, K, M, S, D Distictella magnoliifolia (Kunth) Sandwith, K, M Distictella parkeri (DC.) Sprague & Sandwith, K, M, D Distictis pulverulenta (Sandwith) A.H. Gentry, D Jacaranda copaia (Aubl.) D. Don, I, K, M, S, D Jacaranda obtusifolia Bonpl., I, K Leucocalantha aromatica Barb.Rodr., D Lundia corymbifera (Vahl) Sandwith, S Lundia densiflora DC., I, D Lundia erionema DC., M Macfadyena unguis-cati (L.) A.H. Gentry, S Manaosella cordifolia (DC.) A.H. Gentry, D Mansoa alliacea (Lam.) A.H. Gentry, S, D Mansoa standleyi (Steyerm.) A.H. Gentry, S Martinella iquitosensis A. Samp., D Martinella obovata (Kunth) Bureau & K. Schum., K, M, S, D Memora adenophora Sandwith, D Memora bracteosa (DC.) Bureau ex K. Schum., M, D Memora flavida (DC.) Bureau & K. Schum., M, S, D Memora flaviflora (Miq.) Pulle, D Memora longilinea A. Samp., D Memora moringifolia (DC.) Sandwith, M, S, D Memora racemosa A.H. Gentry, I, M, S, D Memora schomburgkii (DC.) Miers, I, K, M Memora tanaeciicarpa A.H. Gentry, S, D Mussatia hyacinthina (Standl.) Sandwith, I Mussatia prieuriei (DC.) Bureau ex K. Schum., M Parabignonia steyermarkii Sandwith, I, M, S Paragonia pyramidata (Rich.) Bureau, I, K, M, S Periarrabidaea truncata A. Samp., D Phryganocydia corymbosa (Vent.) Baill., K, M Pithecoctenium crucigerum (L.) A.H. Gentry, S Pleonotoma albiflora (Salzm. ex DC.) A.H. Gentry, M, S, D I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

32 26 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana Pleonotoma clematis (Kunth) Miers, I, M Pleonotoma dendrotricha Sandwith, S Pleonotoma echitidea Sprague & Sandwith, M Pleonotoma jasminifolia (Kunth) Miers, D Potamoganos microcalyx (G. Mey.) Sandwith, M Pyrostegia cinerea Bureau ex K. Schum., D Roentgenia sordida (Bureau & K. Schum.) Sprague & Sandwith, M Schlegelia fuscata A.H. Gentry, S Schlegelia paraensis Ducke, M, S, D Schlegelia spruceana Bureau & K. Schum., K, D Schlegelia violacea (Aubl.) Griseb., K, M Spathicalyx duckei (A. Samp.) A.H. Gentry, D Stizophyllum riparium (Kunth) Sandwith, S, D Tabebuia capitata (Bureau & K. Schum.) Sandwith, S Tabebuia incana A.H. Gentry, D Tabebuia insignis (Miq.) Sandwith, I, K, M Tabebuia serratifolia (Vahl) G. Nicholson, M, S, D Tabebuia stenocalyx Sprague & Stapf, I Tabebuia subtilis Sprague & Sandwith, K Tanaecium jaroba Sw., I Tanaecium nocturnum (Barb. Rodr.) Bureau & K. Schum., S Tynanthus panurensis (Bureau) K. Schum., D Tynanthus polyanthus (Bureau) Sandwith, S Tynanthus pubescens A.H. Gentry, S Tynanthus sp., I Bixaceae Bixa orellana L., I, K, S Cochlospermum orinocense (Kunth) Steud., I Blechnaceae Blechnum gracile Kaulf., S Blechnum serrulatum Rich., K, M Salpichlaena hookeriana (Kuntze) Alston, D Salpichlaena volubilis (Kaulf.) J. Sm., S, D Bombacaceae Bombacopsis macrocalyx (Ducke) A. Robyns, D Bombacopsis nervosa (Uittien) A. Robyns, D Carolinea minor Sims, D Catostemma albuquerquei Paula, D Catostemma commune Sandwith, I, K, M Catostemma fragrans Benth., I, K, M Catostemma milanezii Paula, D Catostemma sclerophyllum Ducke, D Catostemma sp., S Ceiba pentandra (L.) Gaertn., S Eriotheca globosa (Aubl.) A. Robyns, S, D Eriotheca longitubulosa A. Robyns, D Eriotheca macrophylla (K. Schum.) A. Robyns, D Huberodendron swietenioides (Gleason) Ducke, S, D Matisia ochrocalyx K. Schum., S Pachira aquatica Aubl., I, K Pachira insignis (Sw.) Sw. ex Savigny, I, S Quararibea duckei Huber, S Quararibea guianensis Aubl., S Quararibea lasiocalyx (K. Schum.) Vischer, S Quararibea spatulata Ducke, S Rhodognaphalopsis duckei A. Robyns, D Rhodognaphalopsis faroensis (Ducke) A. Robyns, D Rhodognaphalopsis flaviflora (Pulle) A. Robyns, K, M Rhodognaphalopsis minor (Sims) A. Robyns, K, D Scleronema micranthum (Ducke) Ducke, D Bonnetiaceae Archytaea triflora Mart., K Bonnetia sessilis Benth., K Boraginaceae Cordia alliodora (Ruiz & Pav.) Oken, I, S Cordia bicolor DC., M, S Cordia exaltata Lam., I, M, S Cordia fallax I.M. Johnst., I, M Cordia goeldiana Huber, S Cordia grandiflora (Desv.) Kunth, I Cordia hirta I.M. Johnst., S Cordia lomatoloba I.M. Johnst., S Cordia naidophilla I.M. Johnst., D Cordia nervosa Lam., K, M, S Cordia nodosa Lam., I, K, M, S, D Cordia sagotii I.M. Johnst., I, M, S, D Cordia schomburgkii DC., I, M, S Cordia silvestris Fresen., D Cordia sprucei Mez, D Cordia tetrandra Aubl., I Cordia trachyphylla C.Mart., D Heliotropium filiforme Lehm., I, M Heliotropium ternatum Vahl, I Tournefortia bicolor Sw., S Tournefortia cuspidata Kunth, I Tournefortia hirsutissima L., I Tournefortia maculata Jacq., S Tournefortia melanochaeta DC., I Tournefortia paniculata Cham., S Tournefortia ulei Vaupel, I, K, M, S Bromeliaceae Aechmea aquilega (Salisb.) Griseb., K, S Aechmea beeriana L.B. Sm. & M.A. Spencer, D Aechmea brassicoides Baker, K Aechmea bromeliifolia (Rudge) Baker, I, K, M, S, D Aechmea contracta (Mart. ex Schult.f.) Baker, D Aechmea fernandae (E. Morren) Baker, D Aechmea lingulata (L.) Baker, S Aechmea melinonii Hook., M, S Aechmea mertensii (G. Mey.) Schult. & Schult.f., I, K, M, S, D Aechmea nudicaulis (L.) Griseb., K Aechmea rodriguesiana (L.B.Sm) L.B.Sm., D Aechmea setigera Mart. ex Schult.f., D Aechmea tillandsioides (Mart. ex Schult.f.) Baker, I, K, M Aechmea tocantina Baker, K, S Ananas ananassoides (Baker) L.B. Sm., K Ananas parguazensis L.A. Camargo & L.B. Sm., I, K Ananas sp., M Araeococcus flagellifolius Harms, S Araeococcus micranthus Brongn., S, D Billbergia violacea Beer, S Billbergia sp., K Brocchinia micrantha (Baker) Mez, K Brocchinia reducta Baker, K Bromelia plumieri (E. Morren) L.B. Sm., M Bromelia tubulosa L.B. Sm., D I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

33 BRIT.ORG/SIDA 27 Catopsis berteroniana (Schult. & Schult.f.) Mez, K, M, S Catopsis sessiliflora (Ruiz & Pav.) Mez, S Disteganthus basi-lateralis Lem., S Disteganthus lateralis (L.B. Sm.) Gouda, S Gravisia rodriguesiana L.B. Sm., D Guzmania altsonii L.B. Sm., K, S Guzmania brasiliensis Ule, D Guzmania lingulata (L.) Mez, I, K, M, S, D Guzmania melinonis Regel, S Guzmania roezlii (E. Morren) Mez, I, M Guzmania squarrosa (Mez & Sodiro) L.B. Sm. & Pittendr., K Lindmania guianensis (Beer) Mez, K Navia angustifolia (Baker) Mez, K Navia sandwithii L.B. Sm., K Neoregelia eleutheropetala (Ule) L.B.Sm., D Pitcairnia caricifolia Mart. ex Schult.f., K, M Pitcairnia geyskesii L.B. Sm., S Pitcairnia leprieurii Baker, S Pitcairnia maidifolia (C. Morren) Decne. ex Planch. & Linden, K Pitcairnia nuda Baker, K Pitcairnia sastrei L.B. Sm. & Read, S Pitcairnia sprucei Baker, I, D Streptocalyx longifolius (Rudge) Baker, M, S, D Tillandsia adpressiflora Mez, K, S Tillandsia anceps Lodd., K, M, S Tillandsia bulbosa Hook., K, M, S Tillandsia fasciculata Sw., M, S Tillandsia fendleri Griseb., K Tillandsia jenmanii Baker, K Tillandsia kegeliana Mez, S Tillandsia monadelpha (E. Morren) Baker, M, S Tillandsia paraensis Mez, S Tillandsia spiculosa Griseb., K, M, S Tillandsia tenuifolia L., M, S Tillandsia sp., D Vriesea amazonica (Baker) Mez, M, S Vriesea gladioliflora (H. Wendl.) Antoine, K, M, S Vriesea pleiosticha (Griseb.) Gouda, K, S Vriesea rubra (Ruiz & Pav.) Beer, M Vriesea soderstromii L.B. Sm., K Vriesea splendens (Brongn.) Lem., K, S Vriesea splitgerberi (Mez) L.B. Sm. & Pittendr., D Vriesea viridiflora (Regel) Wittm. ex Mez, K Burmanniaceae Apteria aphylla (Nutt.) Barnhart ex Small, K, S Apteria sp. 1, D Burmannia bicolor Mart., K Burmannia capitata (Walter ex J.F. Gmel.) Mart., K Burmannia tenella Benth., K Campylosiphon purpurascens Benth., I, K, M, S, D Cymbocarpa refracta Miers, D Cymbocarpa saccata Sandwith, D Dictyostega orobanchoides (Hook.) Miers, M, S, D Gymnosiphon brachycephalus Snelders & Maas, D Gymnosiphon breviflorus Gleason, I, K, M, S, D Gymnosiphon divaricatus (Benth.) Benth. & Hook.f., I, M, S, D Gymnosiphon guianensis Gleason, K, M Gymnosiphon minutus Snelders & Maas, S, D Gymnosiphon recurvatus Snelders & Maas, K Gymnosiphon tenellus (Benth.) Urb., D Hexapterella gentianoides Urb., M, S, D Thismia saülensis H. Maas & Maas, S Burseraceae Bursera graveolens (Kunth) Triana & Planch., D Crepidospermum goudotianum (Tul.) Triana & Planch., I, S Crepidospermum rhoifolium (Benth.) Triana & Planch., I Dacryodes cuspidata (Cuatrec.) Daly, D Dacryodes nitens Cuatrec., D Dacryodes roraimensis Steyerm. & Maguire, S Dacryodes sp. nov., D Protium altsonii Sandwith, D Protium apiculatum Swart, S, D Protium aracouchini (Aubl.) Marchand, K, M, S, D Protium araguense Cuatrec., S Protium crassipetalum Cuatrec., D Protium cuneatum Swart, S Protium decandrum (Aubl.) Marchand, I, M, S, D Protium demerarense Swart, K, M, S Protium divaricatum Engl., S, D Protium elegans Engl., D Protium ferrugineum (Engl.) Engl., D Protium fimbriatum Swart, D Protium gallosum Daly, D Protium giganteum Engl., D Protium grandifolium Engl., D Protium guianense (Aubl.) Marchand, I, M, S Protium hebetatum Daly, D Protium heptaphyllum (Aubl.) Marchand, I, M, S, D Protium hostmannii (Miq.) Engl., D Protium insigne (Triana & Planch.) Engl., D Protium klugii J.F. Macbr., D Protium laxiflorum Engl., D Protium morii Daly, S Protium nitidifolium (Cuatrec.) Daly, D Protium nodulosum Swart, D Protium occultum Daly, S Protium opacum Swart, I, S, D Protium pallidum Cuatrec., S, D Protium paniculatum Engl., D Protium pedicellatum Swart, D Protium pilosissimum Engl., D Protium pilosum (Cuatrec.) Daly, S, D Protium plagiocarpium Benoist, S Protium poeppigianum Swart, S Protium polybotryum (Turcz.) Engl., D Protium robustum (Swart) D.M. Porter, S, D Protium rubrum Cuatrec., D Protium sagotianum Marchand, I, S Protium spruceanum (Benth.) Engl., D Protium strumosum Daly, D Protium subserratum (Engl.) Engl., K, S, D Protium tenuifolium (Engl.) Engl., M, S, D Protium trifoliolatum Engl., K, S, D Protium unifoliolatum Spruce ex Engl., D Protium sp. nov. ined., D Tetragastris altissima (Aubl.) Swart, I, M, S Tetragastris hostmannii (Engl.) Kuntze, I Tetragastris panamensis (Engl.) Kuntze, I, S, D I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

34 28 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana Trattinnickia burserifolia Mart., K, M, D Trattinnickia demerarae Sandwith, M Trattinnickia glaziovii Swart, D Trattinnickia peruviana Loes., D Trattinnickia rhoifolia Willd., M, S, D Cactaceae Epiphyllum phyllanthus (L.) Haw., I, S, D Epiphyllum sp., M Rhipsalis baccifera (J.S. Mill.) Stearn, S Rhipsalis sp., M Selenicereus sp., M Selenicereus sp., S Caesalpiniaceae Aldina heterophylla Spruce ex Benth., D Aldina insignis (Benth.) Endl., K Batesia floribunda Spruce ex Benth., S, D Bauhinia alata Ducke, D Bauhinia cupreonitens Ducke, D Bauhinia guianensis Aubl., I, K, M, S, D Bauhinia outimouta Aubl., S Bauhinia rutilans Spruce ex Benth., D Bauhinia scala-simiae Sandwith, K, M Bauhinia siqueiraei Ducke, M, S Bauhinia stenopetala Ducke, D Bauhinia surinamensis Amshoff, M, S Bocoa alterna (Benth.) R.S. Cowan, D Bocoa viridiflora (Ducke) R.S. Cowan, S, D Brownea sp., K Campsiandra comosa Benth., I, M Campsiandra laurifolia Benth., I Candolleodendron brachystachyum (DC.) R.S. Cowan, S Cassia cowanii H.S. Irwin & Barneby, I, S Cassia fastuosa Willd. ex Benth., S Cassia grandis L.f., S Cassia reticulata Willd., D Cassia rubriflora Ducke, D Cassia spruceana Benth., S, D Chamaecrista adiantifolia (Spruce ex Benth.) H.S. Irwin & Barneby, I, K, M, D Chamaecrista apoucouita (Aubl.) H.S. Irwin & Barneby, I, K, M Chamaecrista diphylla (L.) Greene, I Chamaecrista hispidula (Vahl) H.S. Irwin & Barneby, I Chamaecrista negrensis (H.S. Irwin) H.S. Irwin & Barneby, D Chamaecrista nictitans (L.) Moench, I Copaifera guyanensis Desf., K, S Copaifera multijuga Hayne, D Copaifera sp., I Crudia glaberrima (Steud.) J.F. Macbr., I, M Cynometra bauhiniifolia Benth., I Dialium guianense (Aubl.) Sandwith, I, S, D Dicorynia guianensis Amshoff, I, S Dicorynia paraensis Benth., D Dicymbe altsonii Sandwith, M Dicymbe jenmanii Sandwith, K Dicymbe pharangophila R.S. Cowan, K Dimorphandra coccinea Ducke, D Dimorphandra conjugata (Splitg.) Sandwith, M Dimorphandra cuprea Sprague & Sandwith, K Dimorphandra ignea Ducke, D Dimorphandra macrostachya Benth., I, K, S Dimorphandra multiflora Ducke, S Dimorphandra parviflora Spruce ex Benth., D Dimorphandra pennigera Tul., D Dimorphandra polyandra Benoist, M Dimorphandra rufa Sprague & Sandwith, K Dinizia excelsa Ducke, D Elizabetha coccinea M.R. Schomb. ex Benth., I, K, M Elizabetha durissima Ducke, M Elizabetha princeps M.R. Schomb. ex Benth., I, K, S Elizabetha speciosa Ducke, D Eperua duckeana R.S. Cowan, D Eperua falcata Aubl., I, K, M, S Eperua glabriflora (Ducke) R.S. Cowan, D Eperua grandiflora (Aubl.) Benth., K, M, S Eperua jenmanii Oliv., I, K, M Eperua rubiginosa Miq., I, K, M Eperua schomburgkiana Benth., I, K, M Heterostemon ellipticum Mart. ex Benth., I Hymenaea courbaril L., K, M, S Hymenaea intermedia Ducke, D Hymenaea parvifolia Huber, D Hymenaea reticulata Ducke, D Macrolobium acaciifolium (Benth.) Benth., I Macrolobium angustifolium (Benth.) R.S. Cowan, K Macrolobium arenarium Ducke, D Macrolobium bifolium (Aubl.) Pers., I, K, M, S Macrolobium huberianum Ducke, K Macrolobium jenmanii (Gleason) Sandwith, M Macrolobium limbatum Spruce ex Benth., D Macrolobium microcalyx Ducke, D Macrolobium montanum Ducke, M Macrolobium prancei R.S. Cowan, D Macrolobium suaveolens Spruce ex Benth., K, D Martiodendron excelsum (Benth.) Gleason, I Martiodendron parviflorum (Amshoff) R. Koeppen, S Mora excelsa Benth., I, M Mora gonggrijpii (Kleinhoonte) Sandwith, M Paloue guianensis Aubl., I, M Paloue riparia Pulle, M Peltogyne excelsa Ducke, D Peltogyne paniculata Benth., M, S, D Peltogyne venosa (Vahl) Benth., I, M Recordoxylon speciosum (Benoist) Gazel ex Barneby, K, S Sclerolobium chrysophyllum Poepp., D Sclerolobium eriopetalum Ducke, D Sclerolobium guianense Benth., M, D Sclerolobium melanocarpum Ducke, D Sclerolobium melinonii Harms, S Sclerolobium micropetalum Ducke, D Sclerolobium paraense Huber, S Sclerolobium setiferum Ducke, D Senna alata (L.) Roxb., S Senna chrysocarpa (Desv.) H.S. Irwin & Barneby, S Senna hirsuta (L.) H.S. Irwin & Barneby, S Senna latifolia (G. Mey.) H.S. Irwin & Barneby, I, K, M, S Senna lourteigiana H.S. Irwin & Barneby, S Senna multijuga (Rich.) H.S. Irwin & Barneby, K Senna obtusifolia (L.) H.S. Irwin & Barneby, S Senna occidentalis (L.) Link, K, S I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

35 BRIT.ORG/SIDA 29 Senna quinquangulata (Rich.) H.S. Irwin & Barneby, I, K, M, S, D Senna reticulata (Willd.) H.S. Irwin & Barneby, I, S Senna tapajozensis H.S. Irwin & Barneby, D Senna undulata (Benth.) H.S. Irwin & Barneby, I Swartzia amshoffiana R.S. Cowan, S Swartzia arborescens (Aubl.) Pittier, S, D Swartzia benthamiana Miq., I, M Swartzia brachyrachis Harms, D Swartzia corrugata Benth., D Swartzia cuspidata Spruce ex Benth., D Swartzia eriocarpa Benth., K Swartzia grandifolia Bong. ex Benth., K, M Swartzia guianensis (Aubl.) Urb., M Swartzia ingaefolia Ducke, D Swartzia jenmanii Sandwith, M Swartzia laevicarpa Amshoff, I, K, M Swartzia lamellata Ducke, I, K, D Swartzia leiocalycina Benth., M Swartzia leiogyne (Sandwith) R.S. Cowan, K, M Swartzia longipedicellata Sandwith, M Swartzia longistipitata Ducke, D Swartzia oblanceolata Sandwith, M, D Swartzia panacoco (Aubl.) R.S. Cowan, S, D Swartzia polyphylla DC., S, D Swartzia recurva Poepp., I, D Swartzia reticulata Ducke, D Swartzia roraimae Sandwith, M Swartzia schomburgkii Benth., M, D Swartzia sprucei Benth., M Swartzia tessmannii Harms, D Swartzia tillettii R.S. Cowan, I Swartzia tomentifera (Ducke) Ducke, D Swartzia ulei Harms, D Swartzia xanthopetala Sandwith, M Swartzia sp. nov., M Tachigali amplifolia (Ducke) Barneby, S Tachigali guianensis (Benth.) Zarucchi & Herend., I, S Tachigali mymecophila (Ducke) Ducke, D Tachigali paniculata Aubl., D Tachigali plumbea Ducke, D Tachigali pubiflora Benth., K, M Tachigali rusbyi Harms, M Vouacapoua americana Aubl., M, S Vouacapoua macropetala Sandwith, M Vouacapoua pallidior Ducke, D Campanulaceae Centropogon cornutus (L.) Druce, I, K, S Canellaceae Cinnamodendron tenuifolium Uittien, S Capparaceae Capparis leprieurii Briq., S Capparis macrophylla Kunth, D Capparis maroniensis Benoist, M Cleome aculeata L., D Cleome sp., I Crateva tapia L., I Caricaceae Jacaratia spinosa (Aubl.) A. DC., I, S Caryocaraceae Anthodiscus mazarunensis Gilly, K Anthodiscus trifoliatus G. Mey., I Caryocar glabrum (Aubl.) Pers., I, S, D Caryocar microcarpum Ducke, I, K Caryocar montanum Prance, I Caryocar nuciferum L., I, M Caryocar villosum (Aubl.) Pers., S, D Caryophyllaceae Drymaria cordata (L.) Willd. ex Schult., K, S, D Cecropiaceae Cecropia angulata I.W. Bailey, K, M Cecropia concolor Willd., D Cecropia distachya Huber, D Cecropia latiloba Miq., D Cecropia obtusa Trécul, M, S Cecropia purpurascens C.C. Berg, D Cecropia sciadophylla Mart., M, S, D Coussapoa angustifolia Aubl., S Coussapoa asperifolia Trécul, I, D Coussapoa latifolia Aubl., M, S, D Coussapoa microcephala Trécul, I, K, M Coussapoa orthoneura Standl., D Coussapoa sprucei Mildbr., D Coussapoa trinervia Spruce ex Mildbr., D Coussapoa viridifolia Cuatrec., I Pourouma bicolor Mart., S, D Pourouma cecropiifolia Mart., D Pourouma cucura Standl. & Cuatrec., D Pourouma cuspidata Mildbr., D Pourouma ferruginea Standl., D Pourouma guianensis Aubl., M, S, D Pourouma heterophylla Mart. ex Miq., D Pourouma melinonii Benoist, S, D Pourouma minor Benoist, S, D Pourouma mollis Trécul, D Pourouma myrmecophyla Ducke, D Pourouma ovata Trécul, D Pourouma saulensis C.C. Berg & Kooy, S Pourouma tomentosa Mart. ex Miq., S, D Pourouma velutina Mart. ex Miq., D Pourouma villosa Trécul, I, S, D Celastraceae Goupia glabra Aubl., I, M, S, D Maytenus ebenifolia Reissek, D Maytenus floribunda Reissek, S Maytenus guyanensis Klotzsch ex Reissek, I, K, D Maytenus myrsinoides Reissek, M, S Maytenus oblongata Reissek, K, S Chrysobalanaceae Couepia bracteosa Benth., I, M, D Couepia canomensis (Mart.) Benth. ex Hook.f., D Couepia caryophylloides Benoist, S Couepia cognata (Steud.) Fritsch, K, M Couepia comosa Benth., I, K, M Couepia elata Ducke, D Couepia excelsa Ducke, M Couepia guianensis Aubl., K, S, D Couepia habrantha Standl., K, S Couepia joaquinae Prance, S I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

36 30 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana Couepia longipendula Pilg., D Couepia magnoliifolia Benth. ex Hook.f., D Couepia multiflora Benth., I Couepia obovata Ducke, K, S, D Couepia paraensis (Mart. & Zucc.) Benth., I Couepia parillo DC., I, M, S, D Couepia racemosa Benth. ex Hook.f., D Couepia robusta Huber, D Couepia ulei Pilg., D Hirtella angustissima Sandwith, K Hirtella bicornis Mart. & Zucc., S, D Hirtella caduca Fanshawe & Maguire, K Hirtella davisii Sandwith, M, S Hirtella duckei Huber, D Hirtella fasciculata Prance, D Hirtella glandulosa Spreng., S Hirtella hispidula Miq., I, K, S Hirtella myrmecophila Pilg., D Hirtella paniculata Sw., I, K Hirtella physophora Mart. & Zucc., S, D Hirtella piresii Prance, D Hirtella racemosa Lam., I, K, M, S Hirtella rodriguesii Prance, D Hirtella silicea Griseb., I, K, M, S Hirtella suffulta Prance, S Hirtella tenuifolia Prance, S Licania adolphoduckei Prance, D Licania affinis Fritsch, K Licania alba (Bernoulli) Cuatrec., I, K, M, S Licania albiflora Fanshawe & Maguire, S Licania amapaensis Prance, S Licania apetala (E. Mey.) Fritsch, I, S, D Licania boyanii Tutin, I, M Licania bracteata Prance, D Licania buxifolia Sandwith, M Licania canescens Benoist, I, M, S, D Licania caudata Prance, M, S, D Licania coriacea Benth., I, M, D Licania cuprea Sandwith, M Licania densiflora Kleinhoonte, I, M Licania discolor Pilg., I, S Licania divaricata Benth., M Licania egleri Prance, D Licania fanshawei Prance, M, S Licania gracilipes Taub., D Licania granvillei Prance, S Licania guianensis (Aubl.) Griseb., M, S Licania heteromorpha Benth., I, K, M, S, D Licania hirsuta Prance, D Licania hypoleuca Benth., M, D Licania impressa Prance, D Licania incana Aubl., I, K, M Licania intrapetiolaris Spruce ex Hook.f., I, M Licania kunthiana Hook.f., S Licania laevigata Prance, M, S, D Licania lata J.F. Macbr., D Licania latifolia Benth. ex Hook.f., K, D Licania laxiflora Fritsch, K, M, S, D Licania leptostachya Benth., I, K, M Licania licaniiflora (Sagot) S.F. Blake, I Licania longistyla (Hook.f.) Fritsch, I, D Licania macrophylla Benth., D Licania majuscula Sagot, I, M, S Licania membranacea Sagot ex Laness., M, S Licania micrantha Miq., K, M, S, D Licania microphylla Fanshawe & Maguire, K Licania niloi Prance, D Licania oblongifolia Standl., D Licania octandra (Hoffmanns. ex Roem. & Schult.) Kuntze, S, D Licania pallida Spruce ex Sagot, D Licania parvifolia Huber, I Licania persaudii Fanshawe & Maguire, I, M Licania piresii Prance, M Licania polita Spruce ex Hook.f., I Licania prismatocarpa Spruce ex Hook.f., D Licania reticulata Prance, D Licania rodriguesii Prance, D Licania sandwithii Prance, D Licania sothersae Prance, D Licania sprucei (Hook.f.) Fritsch, I, D Licania unguiculata Prance, D Parinari campestris Aubl., I, M Parinari excelsa Sabine, M, S, D Parinari maguirei Prance, K Parinari montana Aubl. emend. Ducke, S Parinari parvifolia Sandwith, M, D Parinari rodolphii Huber, I, M Clusiaceae Calophyllum angulare A.C. Sm., D Calophyllum brasiliense Cambess., M, D Calophyllum sp., K Caraipa costata Spruce ex Benth., D Caraipa densifolia Mart., D Caraipa grandifolia Mart., I, D Caraipa heterocarpa Ducke, D Caraipa odorata Ducke, D Caraipa punctulata Ducke, I, D Caraipa rodriguesii Paula, D Caraipa sp., M Clusia amazonica Planch. & Triana, D Clusia columnaris Engl., I Clusia crassifolia Planch. & Triana, K Clusia cuneata Benth., K, M Clusia flavida (Benth.) Pipoly, I, S Clusia fockeana Miq., I, M Clusia grandiflora Splitg., I, K, M, S, D Clusia hammeliana Pipoly, K, M Clusia insignis Mart., D Clusia leprantha Mart., S, D Clusia maguireana Pipoly, K Clusia melchiorii Gleason, S Clusia mutica Maguire, K Clusia myriandra (Benth.) Planch. & Triana, I, K, M Clusia nemorosa G. Mey., I, K, M, S Clusia obovata (Spruce ex Planch. & Triana) Pipoly, S, D Clusia octandra (Poepp.) Pipoly, S, D Clusia palmicida Rich. ex Planch. & Triana, I, K, M, S Clusia panapanari (Aubl.) Choisy, I, K, M, S Clusia penduliflora Engl., D Clusia platystigma Eyma, S I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

37 BRIT.ORG/SIDA 31 Clusia renggerioides Planch. & Triana, D Clusia rosea Jacq., I Clusia savannarum Maguire, K Clusia schomburgkiana (Planch. & Triana) Benth. ex Engl., K, M Clusia scrobiculata Benoist, K, S, D Clusia sessilis Klotzsch ex Engl., K Clusia spathulaefolia Engl., D Clusia viscida Engl., I Clusiella axillaris (Engl.) Cuatrec., D Clusiella elegans Planch. & Triana, M Dystovomita brasiliensis D Arcy, D Haploclathra paniculata (Mart.) Benth., D Havetiopsis flavida (Benth.) Planch. & Triana, M, D Lorostemon bombaciflorum Ducke, D Lorostemon coelhoi Paula, D Mahurea exstipulata Benth., K Moronobea coccinea Aubl., K, S, D Moronobea jenmanii Engl., K Moronobea pulchra Ducke, D Platonia insignis Mart., M, S Rheedia acuminata (Ruiz & Pav.) Planch. & Triana, K Rheedia benthamiana Planch. & Triana, I, M Rheedia macrophylla (Mart.) Planch. & Triana, I Rheedia madruno (Kunth) Planch. & Triana, I, S, D Symphonia globulifera L.f., K, M, S, D Tovomita acutiflora M.S.Barros & G. Mariz, D Tovomita amazonica Poepp. & Endl. ex Walp, D Tovomita brasiliensis (Mart.) Walp., S, D Tovomita brevistaminea Engl., I, M, S Tovomita calodictyos Sandwith, M, S Tovomita caloneura A.C. Sm., D Tovomita carinata Eyma, I Tovomita cephalostigma Vesque, I, M Tovomita choisyana Planch. & Triana, D Tovomita eggersii Vesque, I Tovomita gracilipes Planch. & Triana, D Tovomita grata Sandwith, D Tovomita longifolia (Rich.) Hochr., I, S Tovomita macrophylla (Poepp.) Walp., I, S, D Tovomita martiana Engl., D Tovomita obovata Engl., S, D Tovomita pyrifolia Planch. & Triana, I Tovomita schomburgkii Planch. & Triana, I, D Tovomita tenuiflora Benth. ex Planch. & Triana, K, M, D Tovomita umbellata Benth., D Tovomita weddeliana Planch. & Triana, D Tovomitopsis membranacea (Planch. & Triana) D Arcy, S Vismia cauliflora A.C. Sm., D Vismia cayennensis (Jacq.) Pers., I, S, D Vismia glaziovii Ruhland, S Vismia gracilis Hieron, D Vismia guianensis (Aubl.) Choisy, I, K, M, S, D Vismia japurensis Reichardt, I, K Vismia latifolia (Aubl.) Choisy, I, K, S Vismia macrophylla Kunth, I, K, M, S Vismia ramuliflora Miq., S Vismia sandwithii Ewan, S, D Vismia sessilifolia (Aubl.) Choisy, I, M Combretaceae Buchenavia congesta Ducke, D Buchenavia fanshawei Exell & Maguire, K, M Buchenavia grandis Ducke, M, S, D Buchenavia guianensis (Aubl.) Alwan & Stace, D Buchenavia macrophylla Eichler, I, D Buchenavia megalophylla van Heurck & Müll. Arg., I Buchenavia nitidissima (Rich.) Alwan & Stace, S Buchenavia ochroprumna Eichler, D Buchenavia parvifolia Ducke, K, S, D Buchenavia reticulata Eichler, D Buchenavia sericocarpa Ducke, D Buchenavia tomentosa Eichler, D Combretum laurifolium Mart., K Combretum laxum Jacq., I, K, M, S, D Combretum rotundifolium Rich., I, M, S Terminalia amazonia (J.F. Gmel.) Exell, I, M Terminalia dichotoma G. Mey., M Terminalia guyanensis Eichler, M, S Terminalia quintalata Maguire, K Commelinaceae Buforrestia candolleana C.B. Clarke, S Commelina diffusa Burm.f., K, S Commelina erecta L., M Commelina rufipes Seub., I, S Commelina sp., D Dichorisandra hexandra (Aubl.) Standl., I, K, S Dichorisandra sp., D Floscopa peruviana Hassk. ex C.B. Clarke, S Tripogandra serrulata (Vahl) Handlos, K Connaraceae Cnestidium guianense (G. Schellenb.) G. Schellenb., I, M, S Connarus coriaceus G. Schellenb., M Connarus erianthus Benth. ex Baker, M Connarus fasciculatus (DC.) Planch., S Connarus lambertii (DC.) Sagot, I Connarus megacarpus S.F. Blake, I, M Connarus perrottetii (DC.) Planch., M, S, D Connarus punctatus Planch., M Pseudoconnarus macrophyllus (Poepp.) Radlk., M, D Pseudoconnarus rhynchosioides (Standl.) Prance, D Rourea accrescens Forero, D Rourea cuspidata Benth. ex Baker, D Rourea frutescens Aubl., M Rourea krukovii Steyerm., D Rourea ligulata Baker, M Rourea paraensis Forero, D Rourea pubescens (DC.) Radlk., M Rourea surinamensis Miq., S Convolvulaceae Calycobolus glaber (Kunth) House, I Dicranostyles ampla Ducke, I, S Dicranostyles densa Spruce ex Meisn., D Dicranostyles falconiana (Barroso) Ducke, D Dicranostyles globostigma D.F. Austin, D Dicranostyles guianensis Mennega, M, S Dicranostyles holostyla Ducke, I, D Dicranostyles integra Ducke, S, D Dicranostyles laxa Ducke, D I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

38 32 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana Dicranostyles longifolia Ducke, D Dicranostyles scandens Benth., D Evolvulus nummularius (L.) L., D Ipomoea batatoides Choisy, S Ipomoea phillomega (Vell.) House, I, M, S Ipomoea quamoclit L., S Ipomoea squamosa Choisy, I, S Ipomoea tiliacea (Willd.) Choisy, I, M Jacquemontia tamnifolia (L.) Griseb., M Lysiostyles scandens Benth., I, M Maripa glabra Choisy, M, S, D Maripa reticulata Ducke, D Maripa scandens Aubl., I, M, S, D Merremia aturensis (Kunth) Hallier f., I Merremia dissecta (Jacq.) Hallier f., S Merremia macrocalyx (Ruiz & Pav.) O Donell, I, K, S Merremia umbellata (L.) Hallier f., M Costaceae Costus arabicus L., I, D Costus claviger Benoist, M, S Costus congestiflorus Rich. ex Gagnep., S, D Costus erythrothyrsus Loes., S Costus lanceolatus Petersen, S Costus scaber Ruiz & Pav., I, S Costus spiralis (Jacq.) Roscoe, S Costus sprucei Maas, D Cucurbitaceae Cayaponia botryocarpa C.Jeffrey, D Cayaponia jenmanii C. Jeffrey, S Cayaponia ophthalmica R.E. Schult., I, M, S Cayaponia racemosa (Mill.) Cogn., M, S Cayaponia rigida (Cogn.) Cogn., S, D Cayaponia selysioides C. Jeffrey, M Cayaponia tessmannii Harms, S Gurania bignoniacea (Poepp. & Endl.) C. Jeffrey, I, M, S, D Gurania brevipedunculata Cogn., D Gurania huebneri Harms, D Gurania lobata (L.) Pruski, I Gurania reticulata Cogn., S Gurania spinulosa (Poepp. & Endl.) Cogn., I, M, S Gurania subumbellata (Miq.) Cogn., I, K, M, S Helmontia cardiophylla Harms, S Helmontia leptantha (Schltdl.) Cogn., M, S, D Melothria pendula L., S Psiguria triphylla (Miq.) C. Jeffrey, I, K, M, S Selysia prunifera (Poepp. & Endl.) Cogn., S Cyatheaceae Alsophila cuspidata (Kunze) D.S. Conant, S Alsophila imrayana (Hook.) D.S. Conant, S Cnemidaria roraimensis (Domin) R.M. Tryon, K Cnemidaria spectabilis (Kunze) R.M. Tryon, I Cyathea andina (H. Karst.) Domin, S Cyathea cyatheoides (Desv.) K.U. Kramer, I, S Cyathea macrocarpa (C. Presl) Domin, I, K Cyathea microdonta (Desv.) Domin, I, S Cyathea pungens (Willd.) Domin, I, K, S Cyathea spectabilis (Kunze) Domin, S Cyathea surinamensis (Miq.) Domin, I, M, S, D Trichipteris microdonta (Desv.) R.M. Tryon, D Cyclanthaceae Asplundia brachyphylla Harling, S Asplundia gleasonii Harling, K Asplundia heteranthera Harling, S Asplundia maguirei Harling, I, K Asplundia vaupesiana Harling, D Asplundia xiphophylla Harling, D Asplundia sp. nov., D Cyclanthus bipartitus Poit., K, M, S, D Dicranopygium angustissimum (Sandwith) Harling, K Dicranopygium pygmaeum (Gleason) Harling, K, S Evodianthus funifer (Poit.) Lindm., K, M, S, D Ludovia lancifolia Brongn., S Stelestylis stylaris (Gleason) Harling, K Thoracocarpus bissectus (Vell.) Harling, I, K, M, S Cyperaceae Becquerelia cymosa Brongn., I, K, M, S, D Bisboeckelera longifolia (Rudge) Kuntze, K Bisboeckelera microcephala (Boeck.) T. Koyama, K, S Bulbostylis junciformis (Kunth) C.B. Clarke, K Bulbostylis juncoides (Vahl) Kük. ex Osten, I Calyptrocarya bicolor (H. Pfeiff.) T. Koyama, I, S, D Calyptrocarya glomerulata (Brongn.) Urb., I, K, M, S Calyptrocarya poeppigiana Kunth, I, K, S Cephalocarpus rigidus Gilly, K Cyperus aggregatus (Willd.) Endl., I Cyperus diffusus Vahl, K, D Cyperus filifolius Willd. ex Kunth, I Cyperus guianensis (C.B. Clarke) Kük., K Cyperus haspan L., I Cyperus laxus Lam., K, S Cyperus ligularis L., I, K, M, D Cyperus luzulae (L.) Rottb. ex Retz., I, M, S, D Cyperus miliifolius Poepp. & Kunth, I Cyperus odoratus L., I, S Cyperus polystachyos Rottb., M Cyperus simplex Kunth, M, S Cyperus sphacelatus Rottb., K, M Cyperus surinamensis Rottb., M, D Cyperus trailii C.B. Clarke, I Didymiandrum stellatum (Boeck.) Gilly, K Diplacrum capitatum (Willd.) Boeck., I Diplacrum guianense (Nees) T. Koyama, K Diplasia karatifolia Rich., I, K, M, S, D Eleocharis debilis Kunth, K Eleocharis filiculmis Kunth, D Eleocharis geniculata (L.) Roem. & Schult., K Eleocharis maculosa (Vahl) Roem. & Schult., K Eleocharis nigrescens (Nees) Steud., D Eleocharis retroflexa (Poir.) Urb., S Eleocharis subfoliata C.B. Clarke, I Exochogyne amazonica C.B. Clarke, K Fimbristylis annua (All.) Roem. & Schult., K, M Fimbristylis aspera Boeck., K Fimbristylis complanata (Retz.) Link, I Fimbristylis dichotoma (L.) Vahl, K, S, D Fimbristylis miliacea (L.) Vahl, I, M Fimbristylis vahlii (Lam.) Link, I Fuirena umbellata Rottb., I, D Hypolytrum amplum Poepp. & Kunth, M Hypolytrum jenmanii C.B. Clarke, I, K, M, S I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

39 BRIT.ORG/SIDA 33 Hypolytrum longifolium (Rich.) Nees, I, K Hypolytrum pulchrum (Rudge) H. Pfeiff., I, K Hypolytrum schraderianum Nees, S, D Kyllinga brevifolia Rottb., D Kyllinga odorata Vahl, M Kyllinga pumila Michx., S, D Lagenocarpus guianensis Nees, I Lagenocarpus rigidus (Kunth) Nees, K Lagenocarpus verticillatus (Spreng.) T. Koyama & Maguire, M Mapania imeriensis (R. Gross) T. Koyama, K Mapania insignis Sandwith, K Mapania macrophylla (Boeck.) H. Pfeiff., K Mapania paradoxa J. Raynal, S Mapania pycnocephala (Benth.) Benth., K, S, D Mapania surinamensis Uittien, I Mapania sylvatica Aubl., I, K, M, S, D Mapaniopsis effusa C.B. Clarke, S Pleurostachys sparsiflora Kunth, D Rhynchospora albida (Nees) Boeck., K Rhynchospora albomarginata Kük., I Rhynchospora amazonica Poepp. & Kunth, I Rhynchospora arenicola Uittien, K Rhynchospora barbata (Vahl) Kunth, I, K Rhynchospora cephalotes (L.) Vahl, I, K, M, S Rhynchospora curvula Griseb., K Rhynchospora emaciata (Nees) Boeck., K Rhynchospora holoschoenoides (Rich.) Herter, I, K, M Rhynchospora nervosa (Vahl) Boeck., I Rhynchospora pubera (Vahl) Boeck., K, M, S, D Rhynchospora radicans (Schltdl. & Cham.) H. Pfeiff., S Rhynchospora reptans (Rich.) Boeck., I Rhynchospora riparia (Nees) Boeck., K Rhynchospora spruceana C.B. Clarke, K Rhynchospora tenella (Nees) Boeck., I, K Rhynchospora watsonii (Britton) Davidse, M, S Scleria baldwinii (Torr.) Steudel, D Scleria bracteata Cav., K Scleria cyperina Kunth, I, K, D Scleria flagellum-nigrorum Bergius, I, K Scleria latifolia Nees, I, S Scleria macrogyne C.B. Clarke, I, K Scleria melaleuca Rchb. ex Schltdl. & Cham., I, K, D Scleria microcarpa Nees ex Kunth, I, K Scleria mitis Bergius, S Scleria pterota J. Presl & C. Presl, K, M Scleria secans (L.) Urb., S Scleria stipitata Uittien, I Scleria stipularis Nees, I, M Scleria tepuiensis Core, S Cyrillaceae Cyrilla racemiflora L., K Dennstaedtiaceae Dennstaedtia obtusifolia (Willd.) T. Moore, S Hypolepis parallelogramma (Kunze) C. Presl, S Lindsaea divaricata Klotzsch, I, K, M, S, D Lindsaea dubia Spreng., S Lindsaea guianensis (Aubl.) Dryand., I, S, D Lindsaea lancea (L.) Bedd., I, K, M, S, D Lindsaea pallida Klotzsch, S Lindsaea parkeri (Hook.) Kuhn, K Lindsaea pendula Klotzsch, K Lindsaea portoricensis Desv., K, S Lindsaea quadrangularis Raddi, I, S Lindsaea sagittata (Aubl.) Dryand., K, M Lindsaea schomburgkii Klotzsch, K Lindsaea stricta (Sw.) Dryand., I, K, D Lindsaea surinamensis Posth., S Lindsaea ulei Hieron., M Pteridium aquilinum (L.) Kuhn, M, S, D Saccoloma elegans Kaulf., I Saccoloma inaequale (Kunze) Mett., I, S, D Dichapetalaceae Dichapetalum pedunculatum (DC.) Baill., I, M, S Dichapetalum rugosum (Vahl) Prance, M, S, D Dichapetalum spruceanum Baill., D Tapura amazonica Poepp., S, D Tapura capitulifera Spruce ex Baill., M Tapura guianensis Aubl., I, M, S, D Tapura lanceolata (Ducke) Rizzini, D Dilleniaceae Curatella americana L., I Davilla cuspidulata Mart. ex. Eichler, D Davilla kunthii A. St.-Hil., I, K, M, S, D Davilla nitida (Vahl) Kubitzki, I Davilla rugosa Poir., S, D Doliocarpus amazonicus Sleumer, D Doliocarpus brevipedicellatus Garcke, I, M, S, D Doliocarpus dentatus (Aubl.) Standl., S, D Doliocarpus guianensis (Aubl.) Gilg, I, K, M, S Doliocarpus macrocarpus Mart. ex Eichler, M, D Doliocarpus magnificus Sleumer, D Doliocarpus major J.F. Gmel., M Doliocarpus paraensis Sleumer, M, S Doliocarpus savannarum Sandwith, K Doliocarpus spraguei Cheesman, I, K, M, D Pinzona coriacea Mart. & Zucc., K, M, S, D Tetracera amazonica Kubitzki, D Tetracera asperula Miq., I, K, M Tetracera costata Mart. ex Eichler, D Tetracera tigarea DC., M Tetracera volubilis L., M Tetracera willdenowiana Steud., D Dioscoreaceae Dioscorea alata L., S Dioscorea amazonum Mart. ex Griseb., I, M Dioscorea cayennensis Lam., S Dioscorea crotalariifolia Uline, M Dioscorea dodecaneura Vell., M Dioscorea guianensis R. Knuth, K Dioscorea marginata Griseb., D Dioscorea megacarpa Gleason, M Dioscorea pachycarpa Kunth, I Dioscorea pilosiuscula Bertero ex Spreng., I, M Dioscorea potarensis R. Knuth, K Dioscorea samydea Griseb., M Dioscorea trichanthera Gleason, K Dioscorea trifida L.f., S Dioscorea truncata Miq., I Droseraceae Drosera capillaris Poir., I I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

40 34 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana Drosera cayennensis Sagot ex Diels, K Drosera kaieteurensis Brumm.-Ding., K Dryopteridaceae Arachniodes ochropteroides (Baker) Lellinger, K Cyclodium guianense (Klotzsch) van der Werff ex L.D. Gómez, K, S Cyclodium inerme (Fée) A.R. Sm., I, K, M, S Cyclodium meniscioides (Willd.) C. Presl, I, K, M, S, D Didymochlaena truncatula (Sw.) J. Sm., S Olfersia cervina (L.) Kunze, K, S Polybotrya caudata Kunze, S Polybotrya fractiserialis (Baker) Sm., S Polybotrya osmundacea Humb. & Bonpl. ex Willd., D Stigmatopteris rotundata (Willd.) C. Chr., K, S Ebenaceae Diospyros artantifolia Mart. ex Miq., S Diospyros bullata A.C. Sm., D Diospyros capreifolia Mart. ex Hiern, S, D Diospyros carbonaria Benoist, S, D Diospyros dichroa Sandwith, M, S Diospyros duckei Sandwith, D Diospyros guianensis (Aubl.) Gürke, K, M, S, D Diospyros ierensis Britton, I, K, M, S Diospyros lissocarpoides Sandwith, I Diospyros manausensis Cavalcante, D Diospyros martinii Benoist ex Amshoff, S Diospyros matheriana A.C. Sm., S Diospyros melinonii (Hiern) A.C. Sm., D Diospyros praetermissa Sandwith, D Diospyros pseudoxylopia Mildbr., D Diospyros vestita Benoist, S, D Elaeocarpaceae Sloanea brevipes Benth., S Sloanea echinocarpa Uittien, S, D Sloanea eichleri K. Schum., M Sloanea excelsa Ducke, D Sloanea fendleriana Benth., D Sloanea floribunda Spruce ex Benth., D Sloanea garckeana K. Schum., I Sloanea grandiflora Sm., I, M, S Sloanea guianensis (Aubl.) Benth., I, M, S, D Sloanea latifolia (Rich.) K. Schum., I, S, D Sloanea laurifolia (Benth.) Benth., M, S, D Sloanea laxiflora Spruce ex Benth., S, D Sloanea nitida G. Don, D Sloanea parviflora Planch. ex Benth., I Sloanea pubescens Benth., S, D Sloanea rufa Planch. ex Benth., S, D Sloanea schomburgkii Spruce ex Benth., D Sloanea sinemariensis Aubl., S Sloanea synandra Spruce ex Benth., D Sloanea tuerckheimii Donn. Sm., S Eremolepidaceae Antidaphne amazonensis Rizzini, D Ericaceae Bejaria sprucei Meisn., K Cavendishia callista Donn. Sm., S Notopora schomburgkii Hook.f., K Orthaea apophysata (Griseb.) A.C. Sm., K Orthaea merumensis Maguire, Steyerm. & Luteyn, K Psammisia guianensis Klotzsch, K, D Satyria cerander (Dunal) A.C. Sm., S Satyria panurensis (Benth. ex Meisn.) Benth. & Hook.f. ex Nied., K Sphyrospermum buxifolium Poepp. & Endl., K, S Sphyrospermum cordifolium Benth., K Thibaudia nutans Klotzsch ex Mansf., K Vaccinium puberulum Klotzsch ex Meisn., K Eriocaulaceae Eriocaulon heterodoxum Moldenke, M Paepalanthus bifidus (Schrad.) Kunth, I, M Paepalanthus dichotomus Klotzsch ex Körn., K Paepalanthus fasciculatus (Rottb.) Kunth, K, D Paepalanthus lamarckii Kunth, M Paepalanthus oyapockensis Herzog, K Paepalanthus pauper Moldenke, K Paepalanthus polytrichoides Kunth, K Rondonanthus capillaceus (Klotzsch ex Körn.) Hensold & Giul., K Syngonanthus gracilis (Bong.) Ruhland, K Syngonanthus jenmanii (Gleason) Giul. & Hensold, K Syngonanthus simplex (Miq.) Ruhland, K Syngonanthus umbellatus (Lam.) Ruhland, I, K, D Syngonanthus xeranthemoides (Bong.) Ruhland, K Tonina fluviatilis Aubl., I, K, S, D Erythroxylaceae Erythroxylum citrifolium A. St.-Hil., K, M, S, D Erythroxylum kapplerianum Peyr., S Erythroxylum macrophyllum Cav., M, S, D Erythroxylum mucronatum Benth., I, S, D Erythroxylum squamatum Sw., M Euphorbiaceae Acalypha diversifolia Jacq., S Alchornea schomburgkii Klotzsch, D Alchornea triplinervia (Spreng.) Müll. Arg., M, S Alchorneopsis floribunda (Benth.) Müll. Arg., M, S, D Amanoa gracillima W.J. Hayden, D Amanoa guianensis Aubl., I, M, D Amanoa sinuosa W.J. Hayden, D Anomalocalyx uleanus (Pax. & K. Hoffm.) Ducke, D Aparisthmium cordatum (A. Juss.) Baill., I, M, S, D Chaetocarpus schomburgkianus (Kuntze) Pax & K. Hoffm., K, M Chamaesyce hirta (L.) Millsp., M, S, D Chamaesyce hyssopifolia (L.) Small, S, D Chamaesyce prostrata (Aiton) Small, K, S Chamaesyce serpens (Kunth) Small, M Chamaesyce thymifolia (L.) Millsp., M, S Conceveiba guianensis Aubl., K, M, S, D Conceveiba martiana Baill., D Croton cuneatus Klotzsch, K Croton draconoides Müll. Arg., S Croton glandulosus L., D Croton hostmanni Miq., K Croton kaieteuri Jabl., K Croton lobatus L., D Croton matourensis Aubl., S, D Croton nuntians Croizat, K Croton palanostigma Klotzsch, S, D Croton schiedeanus Schltdl., S Croton trinitatis Millsp., I, M I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

41 BRIT.ORG/SIDA 35 Croton sp. nov.?, M Dalechampia affinis Müll. Arg., I Dalechampia brevicolumna Armbr., S Dalechampia dioscoreifolia Poepp., S Dalechampia fragrans Armbr., S Dalechampia heterobractea Armbr., S Dalechampia parvibracteata Lanj., I, M, D Dalechampia tiliifolia Lam., S Discocarpus essequiboensis Klotzsch, I Dodecastigma integrifolium (Lanj.) Lanj. & Sandwith, S Drypetes variabilis Uittien, S, D Euphorbia cyathophora Murray, S Glycydendron amazonicum Ducke, M, S, D Hevea guianensis Aubl., S, D Hevea pauciflora (Spruce ex Benth.) Müll. Arg., M Hevea sp., K Hura crepitans L., S Hyeronima alchorneoides Allemão, M, S Hyeronima oblonga (Tul.) Müll. Arg., I, M, S Jatropha gossypiifolia L., S Mabea angularis Hollander, D Mabea biglandulosa Baill., I Mabea montana Müll. Arg., I Mabea piriri Aubl., I, S, D Mabea rubicunda Jabl., K Mabea salicoides Esser, S Mabea speciosa Müll. Arg., I, M, S, D Mabea subsessilis Pax & K. Hoffm., S, D Mabea taquari Aubl., I, K Mabea uleana Pax & K.Hoffm., D Manihot sp. nov., S Maprounea guianensis Aubl., I, K, M, S, D Margaritaria nobilis L.f., I, S Micrandra elata (Didr.) Müll. Arg., I, M Micrandra siphonioides Benth., K, D Micrandra spruceana (Baill.) R.E. Schult., D Nealchornea japurensis Huber, D Omphalea diandra L., I, S Pausandra fordii Secco, S Pausandra macropetala Ducke, D Pausandra martinii Baill., I Pera bicolor (Klotzsch) Müll. Arg., K, M, D Pera glabrata (Schott) Baill., D Phyllanthus amarus Schumach. & Thonn., S Phyllanthus caroliniensis Walter, I Phyllanthus juglandifolius Willd., I Phyllanthus manausensis W.A. Rodrigues, D Phyllanthus niruri L., D Phyllanthus orbiculatus Rich., D Phyllanthus stipulatus (Raf.) G.L. Webster, M Phyllanthus urinaria L., I, S Phyllanthus vacciniifolius (Müll. Arg.) Müll. Arg., K Piranhea trifoliata Baill., K Plukenetia brachybotrya Müll. Arg., D Plukenetia polyadenia Müll. Arg., S Plukenetia verrucosa Sm., I Plukenetia volubilis L., M Podocalyx loranthoides Klotzsch, D Pogonophora schomburgkiana Miers ex Benth., D Richeria dressleri G.L. Webster, D Richeria grandis Vahl, S, D Ruprechtia brachysepala Meisn., I Sagotia racemosa Baill., I, S Sandwithia guyanensis Lanj., I, K, M, D Sapium glandulosum (L.) Morong, S Sapium jenmanii Hemsl., I Sapium montanum Lanj., I, S Sapium paucinervium Hemsl., S, D Sebastiania corniculata (Vahl) Müll. Arg., M, D Senefeldera macrophylla Ducke, D Tetrorchidium minor Ducke, D Tragia lessertiana (Baill.) Müll. Arg., M, S Fabaceae Acosmium nitens (Vogel) Yakovlev, I, M Acosmium praeclarum (Sandwith) Yakovlev, M Aeschynomene rudis Benth., I Aeschynomene sensitiva Sw., I, M Alexa imperatricis (R.H. Schomb.) Baill., I, K, M Andira coriacea Pulle, M Andira grandistipula Amshoff, K Andira micrantha Ducke, D Andira parviflora Ducke, D Andira surinamensis (Bondt) Splitg. ex Amshoff, I Andira unifoliolata Ducke, D Calopogonium mucunoides Desv., M Centrosema brasilianum (L.) Benth., D Centrosema triquetrum (Hoffmanns. ex Benth.) Benth., M Centrosema vexillatum Benth., I, S Centrosema sp., K Clathrotropis brachypetala (Tul.) Kleinhoonte, I, M Clathrotropis glaucophylla R.S. Cowan, I Clathrotropis macrocarpa Ducke, I, K, M Clathrotropis paradoxa Sandwith, K Clitoria kaieteurensis Fantz, K Clitoria leptostachya Benth., M, D Clitoria sagotii Fantz, K, M, S, D Dalbergia ecastaphyllum (L.) Taub., M Dalbergia monetaria L.f., I, D Dalbergia multiflora Heyne ex Wall., D Dalbergia riedelii (Benth.) Sandwith, I, K, M Dalbergia spruceana Benth., D Dalbergia sp., K Derris floribunda (Benth.) Ducke, D Desmodium adscendens (Sw.) DC., I, K, M, S, D Desmodium axillare (Sw.) DC., I, M, S Desmodium barbatum (L.) Benth., K, M, S, D Desmodium incanum DC., S Desmodium wydlerianum Urb., S Dioclea coriacea Benth., K Dioclea elliptica R.H. Maxwell, sp. nov. ined., M Dioclea guianensis Benth., I, K Dioclea macrantha Huber, I, K Dioclea macrocarpa Huber, K, S Dioclea malacocarpa Ducke, K Dioclea megacarpa Rolfe, I, D Dioclea reflexa Hook.f., K Dioclea scabra (Rich.) R.H. Maxwell, I, K, M Dioclea virgata (Rich.) Amshoff, I, K, S Dioclea wilsonii Standl., K Diplotropis martiusii Benth., D Diplotropis purpurea (Rich.) Amshoff, K, M, S I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

42 36 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana Diplotropis racemosa (Hoehne) Amshoff, K Diplotropis rigidocarpa H.C. Lima, K Diplotropis rodriguesii H.C. Lima, D Diplotropis triloba Gleason, D Dipteryx magnifica Ducke, D Dipteryx odorata (Aubl.) Willd., M, S, D Dipteryx polyphylla Huber, D Dipteryx punctata (S.F. Blake) Amshoff, D Dussia discolor (Benth.) Amshoff, S Eriosema simplicifolium (Kunth) G. Don, I Etaballia dubia (Kunth) Rudd, I Galactia sp., I Hymenolobium excelsum Ducke, D Hymenolobium flavum Kleinhoonte, I, M Hymenolobium heterocarpum Ducke, D Hymenolobium modestum Ducke, D Hymenolobium nitidum Benth., K, D Hymenolobium petraeum Ducke, M, S Hymenolobium pulcherrimum Ducke, D Hymenolobium sericeum Ducke, D Hymenolobium sp., I Hymenolobium sp. nov., M Lonchocarpus negrensis Benth., M, D Lonchocarpus scandens (Aubl.) Ducke, S Lonchocarpus sp., I, S Machaerium altiscandens Ducke, S Machaerium aureiflorum Ducke, M, S, D Machaerium cuspidatum Kuhlm. & Hoehne, D Machaerium ferox (Mart. ex Benth.) Ducke, I, M, D Machaerium floribundum Benth., I, K, S Machaerium hoehneanum Ducke, D Machaerium inundatum (Mart. ex Benth.) Ducke, I Machaerium kegelii Meisn., I Machaerium leiophyllum (DC.) Benth., I Machaerium madeirense Pittier, M Machaerium multifoliolatum Ducke, D Machaerium mutisii Killip ex Rudd, M Machaerium myrianthum Spruce ex Benth., K, M Machaerium oblongifolium Vogel, M Machaerium paraënse Ducke, S Machaerium quinata (Aubl.) Sandwith, I, K, M, D Macroptilium gracile (Poepp. ex Benth.) Urb., I Monopteryx inpae W.A. Rodrigues, S, D Mucuna urens (L.) Medik., I, S, D Mucuna sp., K Ormosia coarctata Jacks., I, M Ormosia coccinea (Aubl.) Jacks., I, M Ormosia costulata (Miq.) Kleinhoonte, M Ormosia coutinhoi Ducke, I, K, M Ormosia fastigiata Tul., I Ormosia flava (Ducke) Rudd, S Ormosia grandiflora (Tul.) Rudd, D Ormosia grossa Rudd, D Ormosia paraensis Ducke, M, D Paramachaerium schomburgkii (Benth.) Ducke, I Platymiscium duckei Huber, D Platymiscium ulei Harms ex Harms, S Poecilanthe effusa (Huber) Ducke, S Poecilanthe hostmannii (Benth.) Amshoff, K, M, S Pterocarpus officinalis Jacq., D Pterocarpus rohrii Vahl, K, M, S, D Pterocarpus santalinoides L Hér. ex DC., M Rhynchosia phaseoloides (Sw.) DC., S Taralea oppositifolia Aubl., S, D Vatairea guianensis Aubl., M, D Vatairea paraensis Ducke, S, D Vatairea sericea (Ducke) Ducke, D Vataireopsis surinamensis H.C. Lima, I, S Vigna lasiocarpa (Mart. ex Benth.) Verdc., I Vigna luteola (Jacq.) Benth., I, M Vigna vexillata (L.) A. Rich., S Zornia latifolia Sm., K, M, S, D Zornia sp., I Flacourtiaceae Banara guianensis Aubl., S Carpotroche crispidentata Ducke, S, D Carpotroche surinamensis Uittien, M Carpotroche sp., I Casearia acuminata DC., S Casearia arborea (Rich.) Urb., D Casearia combaymensis Tul., M, S, D Casearia commersoniana Cambess., I, M, S Casearia decandra Jacq., D Casearia duckeana Sleumer, D Casearia grandiflora Cambess., D Casearia javitensis Kunth, M, S, D Casearia macrophylla Vahl, D Casearia manausensis Sleumer, D Casearia negrensis Eichler, S, D Casearia pitumba Sleumer, M, S, D Casearia resinifera Spruce ex Eichler, D Casearia rusbyana Briq., S Casearia singularis Eichler, I, S Casearia sylvestris Sw., I, S, D Casearia ulmifolia Vahl ex Vent., S, D Hasseltia floribunda Kunth, S Homalium guianense (Aubl.) Oken, I, M Homalium racemosum Jacq., I, M Laetia procera (Poepp.) Eichler, M, S, D Mayna odorata Aubl., S Neosprucea sp., I Ryania pyrifera (Rich.) Uittien & Sleumer, M, D Ryania speciosa Vahl, M, S, D Xylosma benthamii (Tul.) Triana & Planch., S Xylosma tessmannii Sleumer, D Gentianaceae Coutoubea humilis Sandwith, K Coutoubea ramosa Aubl., I, K, M, S Coutoubea spicata Aubl., S Curtia tenuifolia (Aubl.) Knobl., K Irlbachia alata (Aubl.) Maas, I, K, M, S, D Irlbachia caerulescens (Aubl.) Griseb., I Irlbachia purpurascens (Aubl.) Maas, I, K, M, S Neurotheca loeselioides (Spruce ex Progel) Baill., K Tachia grandiflora Maguire & Weaver, S, D Tachia guianensis Aubl., K Voyria aphylla (Jacq.) Pers., I, K, M, S, D Voyria aurantiaca Splitg., S, D Voyria caerulea Aubl., I, M, S, D Voyria clavata Splitg., M Voyria corymbosa Splitg., I, M, S I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

43 BRIT.ORG/SIDA 37 Voyria flavescens Griseb., S Voyria rosea Aubl., I, M, S Voyria spruceana Benth., S, D Voyria tenella Hook., S, D Voyria tenuiflora Griseb., M, S Voyriella parviflora (Miq.) Miq., I, M, S, D Gesneriaceae Alloplectus savannarum C.V. Morton, K Besleria flavovirens Nees & Mart., I Besleria insolita C.V. Morton, S Besleria laxiflora Benth., I, M Besleria patrisii DC., S Besleria verecunda C.V. Morton, M Chrysothemis pulchella (Donn ex Sims) Decne., M, S Chrysothemis rupestris (Benth.) Leeuwenb., I Codonanthe calcarata (Miq.) Hanst., I, K, M, S, D Codonanthe crassifolia (H. Focke) C.V. Morton, I, K, M, S, D Codonanthopsis dissimulata (H.E. Moore) Wiehler, D Codonanthopsis ulei Mansf., D Columnea calotricha Donn. Sm., S Columnea guianensis C.V. Morton, K Columnea oerstediana Klotzsch ex Oerst., S Columnea sanguinea (Pers.) Hanst., S Drymonia coccinea (Aubl.) Wiehler, S, D Drymonia psilocalyx Leeuwenb., S Drymonia serrulata (Jacq.) Mart., M, S Episcia xantha Leeuwenb., I, S Episcia sp., K Gloxinia purpurascens (Rusby) Wiehler, K Kohleria hirsuta (Kunth) Regel, K Napeanthus macrostoma Leeuwenb., S Nautilocalyx adenosiphon (Leeuwenb.) Wiehler, S Nautilocalyx cordatus (Gleason) L.E. Skog, I, K Nautilocalyx mimuloides (Benth.) C.V. Morton, S Nautilocalyx pictus (Hook.) Sprague, K, M, S, D Paradrymonia campostyla (Leeuwenb.) Wiehler, S Paradrymonia ciliosa (Mart.) Wiehler, K, D Paradrymonia densa (C.H. Wright) Wiehler, I, K, M Paradrymonia maculata (Hook.f.) Wiehler, K, M Rhoogeton cyclophyllus Leeuwenb., K Rhoogeton viviparus Leeuwenb., K Tylopsacas cuneatum (Gleason) Leeuwenb., K Gleicheniaceae Dicranopteris flexuosa (Schrad.) Underw., I, K Dicranopteris pectinata (Willd.) Underw., K, S, D Sticherus remotus (Kaulf.) Chrysler, D Gnetaceae Gnetum nodiflorum Brongn., I, M, D Gnetum paniculatum Spruce ex Benth., I, D Gnetum schwackeanum Taub. ex Schenk, D Gnetum urens (Aubl.) Blume, I, M, S, D Grammitidaceae Cochlidium furcatum (Hook. & Grev.) C. Chr., I, K, M, S Cochlidium linearifolium (Desv.) Maxon ex C. Chr., I, K, S, D Cochlidium serrulatum (Sw.) L.E. Bishop, I, K, M, S, D Grammitis blanchetii (C. Chr.) A.R. Sm., M, S Grammitis bryophila (Maxon) F. Seym., K Grammitis flabelliformis (Poir.) C.V. Morton, K, M Grammitis kaieteura (Jenman) C.V. Morton, K Grammitis mollissima (Fée) Proctor, K Grammitis staheliana (Posth.) Lellinger, S Grammitis suspensa (L.) Proctor, K, S Grammitis taenifolia (Jenman) Proctor, K, S Grammitis taxifolia (L.) Proctor, S Haemodoraceae Schiekia orinocensis (Kunth) Meisn., S Xiphidium caeruleum Aubl., I, S Heliconiaceae Heliconia acuminata Rich., I, K, M, S, D Heliconia bihai (L.) L., I, K, M, S Heliconia chartacea Lane ex Barreiros, I, M Heliconia densiflora B. Verl., M, S Heliconia hirsuta L.f., I, S Heliconia lourteigiae Emygdio & E. Santos, I, S Heliconia pendula Wawra, S Heliconia psittacorum L.f., I, S, D Heliconia richardiana Miq., S Heliconia spathocircinata Aristeg., I, S Hernandiaceae Hernandia guianensis Aubl., S Sparattanthelium acreanum Pilg., D Sparattanthelium guianense Sandwith, S Sparattanthelium tarapotanum Meisn., D Sparattanthelium uncigerum (Meisn.) Kubitzki, S Sparattanthelium wonotoboense Kosterm., S Hippocrateaceae Anthodon decussatum Ruiz & Pav., D Cheiloclinium belizense (Standl.) A.C. Sm., K Cheiloclinium cognatum (Miers) A.C. Sm., M, S Cheiloclinium diffusiflorum (Miers) A.C. Sm., K, M Cheiloclinium hippocrateoides (Peyr.) A.C. Sm., K, M, S, D Cheiloclinium krukovii (A.C. Sm.) A.C. Sm., D Cuervea kappleriana (Miq.) A.C. Sm., D Elachyptera floribunda (Benth.) A.C. Sm., S Hippocratea volubilis L., I, K, M, S Peritassa compta Miers, I, M Peritassa laevigata (Hoffmanns.) A.C. Sm., I, M Prionostemma aspera (Lam.) Miers, I, M, S Pristimera nervosa (Miers) A.C. Sm., S Salacia elliptica (Mart. ex Roem. & Schult.) G. Don, I Salacia impressifolia (Miers) A.C. Sm., I, S, D Salacia insignis A.C. Sm., S Salacia juruana Loes., I, S Salacia maburensis Mennega, I, M Salacia multiflora (Lam.) DC., I, M, S, D Tontelea attenuata Miers, M, S Tontelea coriacea A.C. Sm., M Tontelea cylindrocarpa (A.C. Sm.) A.C. Sm., S Tontelea emarginata A.C. Sm., D Tontelea fluminensis (Peyr.) A.C. Sm., D Tontelea laxiflora (Benth.) A.C. Sm., K, M, S Tontelea nectandrifolia (A.C. Sm.) A.C. Sm., M Tontelea ovalifolia (Miers) A.C. Sm., I, M, S, D Tontelea sandwithii A.C. Sm., S Hugoniaceae Hebepetalum humiriifolium (Planch.) Benth., M, S, D Roucheria punctata (Ducke) Ducke, D I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

44 38 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana Roucheria schomburgkii Planch., M Roucheria sp., K Humiriaceae Duckesia verrucosa (Ducke) Cuatrec., D Endopleura uchi (Huber) Cuatrec., D Humiria balsamifera Aubl., I, K, M, D Humiria crassifolia Mart. ex Urb., K Humiriastrum excelsum (Ducke) Cuatrec., S Humiriastrum obovatum (Benth.) Cuatrec., M Sacoglottis ceratocarpa Ducke, D Sacoglottis guianensis Benth., M, S, D Sacoglottis matogrossensis Malme, D Schistostemon dichotomum (Urb.) Cuatrec., I Vantanea guianensis Aubl., I, S, D Vantanea macrocarpa Ducke, D Vantanea micrantha Ducke, D Vantanea parviflora Lam., S, D Hymenophyllaceae Hymenophyllum decurrens (Jacq.) Sw., K, S Hymenophyllum hirsutum (L.) Sw., K, S Hymenophyllum kaieteurum Jenman, K Hymenophyllum polyanthos (Sw.) Sw., I, K, S, D Hymenophyllum sp., M Trichomanes angustifrons (Fée) Wess. Boer, K, S Trichomanes ankersii C. Parker ex Hook. & Grev., I, M, S, D Trichomanes anomalum Maxon & C.V. Morton, K Trichomanes arbuscula Desv., I, K, M, D Trichomanes bancroftii Hook. & Grev., K Trichomanes botryoides Kaulf., M Trichomanes cellulosum Klotzsch, K, D Trichomanes crispum L., I, K, S Trichomanes cristatum Kaulf., K, S, D Trichomanes diaphanum Kunth, I, K, M, S Trichomanes diversifrons (Bory) Mett. ex Sadeb., I, S Trichomanes ekmanii Wess. Boer, S Trichomanes elegans Rich., K, S, D Trichomanes hostmannianum (Klotzsch) Kunze, K Trichomanes jenmanii Lellinger, M Trichomanes kapplerianum J.W. Sturm, K, M, S Trichomanes krausii Hook. & Grev., S Trichomanes martiusii C. Presl, I, M, D Trichomanes membranaceum L., S Trichomanes mougeotii Bosch, S Trichomanes osmundoides DC. ex Poir., S Trichomanes ovale (E. Fourn.) Wess. Boer, D Trichomanes pedicellatum Desv., I, K, M, S Trichomanes pilosum Raddi, D Trichomanes pinnatum Hedw., I, K, M, S, D Trichomanes polypodioides L., S Trichomanes punctatum Poir., M, S, D Trichomanes radicans Sw., S Trichomanes rigidum Sw., S Trichomanes robustum E. Fourn., S Trichomanes trollii Bergdolt, I, S Trichomanes tuerckheimii H. Christ, S Trichomanes vittaria DC. ex Poir., S Icacinaceae Casimirella rupestris (Ducke) R.A. Howard, D Dendrobangia boliviana Rusby, S, D Discophora guianensis Miers, M, S, D Emmotum acuminatum (Benth.) Miers, D Emmotum fagifolium Ham., I, M, S Emmotum glabrum Benth. ex Miers, K, D Leretia cordata Vell., M, S Pleurisanthes emarginata Tiegh., D Pleurisanthes flava Sandwith, K, M Pleurisanthes parviflora (Ducke) R.A. Howard, D Poraqueiba guianensis Aubl., S, D Poraqueiba sericea Tul., D Iridaceae Eleutherine bulbosa (Mill.) Urb., S Ixonanthaceae Ochthocosmus barrae Hallier f., K Ochthocosmus roraimae Benth., K Lacistemataceae Lacistema aggregatum (Bergius) Rusby, I, K, M, S, D Lacistema grandifolium W. Schnizl., S, D Lacistema polystachyum W. Schnizl., S, D Lamiaceae Hyptis atrorubens Poit., M, S, D Hyptis lanceolata Poir., I, S Hyptis lantanifolia Poit., D Hyptis mutabilis (Rich.) Briq., S Hyptis pachycephala Epling, S Hyptis parkeri Benth., I Hyptis pectinata (L.) Poit., K Leonotis nepetifolia (L.) W.T. Aiton, S Marsypianthes chamaedrys (Vahl) Kuntze, S, D Ocimum campechianum Mill., I, K, S Scutellaria uliginosa A. St.-Hil. ex Benth., S Lauraceae Aiouea grandifolia van der Werff, D Aiouea guianensis Aubl., I, M Aiouea laevis (Mart.) Kosterm., I Aiouea longipetiolata van der Werff, S Aiouea maguireana (C.K. Allen) S.S. Renner, D Aiouea myristicoides Mez, D Aiouea opaca van der Werff, S Aniba burchellii Kosterm., D Aniba canelilla (Kunth) Mez, M, D Aniba citrifolia (Nees) Mez, I, M, S Aniba excelsa Kosterm., I, M Aniba ferrea Kubitzki, D Aniba hostmanniana (Nees) Mez, I, D Aniba hypoglauca Sandwith, I, K, M Aniba jenmanii Mez, D Aniba kappleri Mez, M, S Aniba megaphylla Mez, I, D Aniba panurensis (Meisn.) Mez, I, M, D Aniba parviflora (Meisn.) Mez, S, D Aniba permollis (Nees) Mez, K Aniba riparia (Nees) Mez, D Aniba rosaeodora Ducke, K, M, D Aniba santalodora Ducke, D Aniba terminalis Ducke, D Aniba williamsii O.C. Schmidt, S, D Aniba sp. nov., S Beilschmiedia hexanthera van der Werff, S Cassytha filiformis L., I, K I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

45 BRIT.ORG/SIDA 39 Chlorocardium rodiei (R.H. Schomb.) Rohwer, H.G. Richt. & van der Werff, I, M Cinnamomum cinnamomifolium (Kunth) Kosterm., S Dicypellium manausense W.A. Rodrigues, D Endlicheria bracteata Mez, D Endlicheria bracteolata (Meisn.) C.K. Allen, D Endlicheria chalisea Chanderbali, I, M, D Endlicheria citriodora van der Werff, D Endlicheria longicaudata (Ducke) Kosterm., D Endlicheria macrophylla (Meisn.) Mez, D Endlicheria multiflora (Miq.) Mez, I, M, S Endlicheria punctulata (Mez) C.K. Allen, M, S Endlicheria pyriformis (Nees) Mez, S, D Endlicheria sericea Nees, S, D Endlicheria sprucei (Meisn.) Mez, D Endlicheria szyszylowiczii Mez, D Endlicheria wurdackiana C.K. Allen, S Endlicheria sp. nov., D Kubitzkia mezii (Kosterm.) van der Werff, S, D Licaria cannella (Meisn.) Kosterm., I, K, M, S, D Licaria chrysophylla (Meisn.) Kosterm., S, D Licaria debilis (Mez) Kosterm., I, K, S Licaria guianensis Aubl., I, S, D Licaria hirsuta van der Werff ined., D Licaria macrophylla (A.C. Sm.) Kosterm., D Licaria martiniana (Mez) Kosterm., I, S, D Licaria oppositifolia (Nees) Kosterm., D Licaria pachycarpa (Meisn.) Kosterm., D Licaria puchury-major (Mart.) Kosterm., D Licaria rodriguesii Kurz, D Licaria subbullata Kosterm., S, D Licaria vernicosa (Mez) Kosterm., S Mezilaurus duckei van der Werff, D Mezilaurus itauba (Meisn.) Taub. ex Mez, I, D Mezilaurus synandra (Mez) Kosterm., D Nectandra amazonum Nees, I, M Nectandra cissiflora Nees, S Nectandra cuspidata Nees, D Nectandra globosa (Aubl.) Mez, I, S Nectandra hihua (Ruiz & Pav.) Rohwer, S Nectandra purpurea (Ruiz & Pav.) Mez, S Nectandra reticulata (Ruiz & Pav.) Mez, S Ocotea aciphylla (Nees & Mart. ex Nees) Mez, M, S, D Ocotea adenotrachelium (Nees) Mez, D Ocotea amazonica (Meisn.) Mez, D Ocotea argyrophylla Ducke, D Ocotea boissieriana (Meisn.) Mez, D Ocotea canaliculata (Rich.) Mez, I, K, M, S Ocotea ceanothifolia (Nees) Mez, I, S, D Ocotea cernua (Nees) Mez, I, K, M, S, D Ocotea cinerea van der Werff, ined., S, D Ocotea commutata (Nees) Mez, S Ocotea cujumari aff. Mart., D Ocotea cymbarum Kunth, M Ocotea delicata Vicentini ined., D Ocotea discrepens C.K. Allen, M Ocotea dissimilis C.K. Allen, S Ocotea douradensis Vattimo-Gil, D Ocotea fendleri (Meisn.) Rohwer, S Ocotea floribunda (Sw.) Mez, M, S, D Ocotea glomerata (Nees) Mez, M, S Ocotea guianensis Aubl., I, M, D Ocotea immersa van der Werff ined., D Ocotea indirectinervia C.K. Allen, S Ocotea leucoxylon (Sw.) Laness., I, M, D Ocotea longifolia Kunth, S, D Ocotea matogrossensis Vattimo, D Ocotea minor Vicentini ined., D Ocotea myriantha (Meisn.) Mez, D Ocotea neblinae C.K. Allen, D Ocotea nigra Benoist, I Ocotea nigrescens Vicentini ined., D Ocotea nitida (Meisn.) Rohwer, D Ocotea obliqua Vicentini ined., D Ocotea oblonga (Meisn.) Mez, S, D Ocotea olivacea A.C. Sm., D Ocotea pauciflora (Nees) Mez, M, D Ocotea percurrens Vicentini ined., D Ocotea puberula (Rich.) Nees, M, S, D Ocotea rhodophylla Vicentini ined., D Ocotea rhynchophylla (Meisn.) Mez, D Ocotea rubra Mez, K, S Ocotea scabrella van der Werff ined., D Ocotea schomburgkiana (Nees) Mez, I, K, M Ocotea splendens (Meisn.) Baill., I, S, D Ocotea subterminalis van der Werff ined., D Ocotea tabacifolia (Meisn.) Rohwer, D Ocotea tomentella Sandwith, M, S Ocotea sp. nov., M Persea jenmanii Mez, K Pleurothyrium vasquezii van der Werff, D Rhodostemonodaphne crenaticupula Madriñán, I, D Rhodostemonodaphne elephantopus Madriñán, S Rhodostemonodaphne grandis (Mez) Rohwer, S, D Rhodostemonodaphne kunthiana (Nees) Rohwer, I, S Rhodostemonodaphne leptoclada Madriñán, S Rhodostemonodaphne morii Madriñán, I, S Rhodostemonodaphne negrensis Madriñán, D Rhodostemonodaphne parvifolia Madriñán, D Rhodostemonodaphne peneia Madriñán, D Rhodostemonodaphne praeclara (Sandwith) Madriñán, I Rhodostemonodaphne recurva van der Werff ined., D Rhodostemonodaphne rufovirgata Madriñán, S Rhodostemonodaphne saulensis Madriñán, S Rhodostemonodaphne scandens Madriñán, D Rhodostemonodaphne sordida Madriñán, D Rhodostemonodaphne sp. nov., I Sextonia rubra (Mez) van der Werff, D Lecythidaceae Allantoma lineata (Mart. ex O. Berg) Miers, D Bertholletia excelsa Bonpl., D Cariniana decandra Ducke, D Cariniana integrifolia Ducke, D Cariniana micrantha Ducke, D Corythophora alta R. Knuth, D Corythophora amapaensis Pires ex S.A. Mori & Prance, S Corythophora rimosa W.A. Rodrigues, S, D Couratari calycina Sandwith, K Couratari gloriosa Sandwith, M, S Couratari guianensis Aubl., M, S, D Couratari longipedicellata W.A. Rodrigues, D I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

46 40 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana Couratari multiflora (Sm.) Eyma, K, S, D Couratari oblongifolia Ducke & R. Knuth, S Couratari oligantha A.C. Sm., K Couratari riparia Sandwith, K Couratari stellata A.C. Sm., I, S, D Couratari tauari O. Berg, D Eschweilera alata A.C. Sm., K, M Eschweilera amazoniciformis S.A.Mori, D Eschweilera apiculata (Miers) A.C. Sm., S Eschweilera atropetiolata S.A.Mori, D Eschweilera bracteosa (Poepp. ex O. Berg) Miers, D Eschweilera carinata S.A.Mori, D Eschweilera chartaceifolia S.A. Mori, S Eschweilera collina Eyma, S, D Eschweilera congestiflora (Benoist) Eyma, I Eschweilera coriacea (DC.) S.A. Mori, I, K, M, S, D Eschweilera cyathiformis S.A.Mori, D Eschweilera decolorans Sandwith, I, K, M, S Eschweilera eperuetorum Sandwith, K Eschweilera grandiflora (Aubl.) Sandwith, S, D Eschweilera laevicarpa S.A. Mori, S Eschweilera micrantha (O. Berg) Miers, I, M, S, D Eschweilera parviflora (Aubl.) Miers, M, S Eschweilera pedicellata (Rich.) S.A. Mori, I, M, S, D Eschweilera piresii S.A. Mori, S Eschweilera pseudodecolorans S.A.Mori, D Eschweilera rhododendrifolia (R. Knuth) A.C. Sm., D Eschweilera romeu-cardosoi S.A.Mori, D Eschweilera sagotiana Miers, I, K, M, S Eschweilera simiorum (Benoist) Eyma, I, S Eschweilera squamata S.A. Mori, S Eschweilera subglandulosa (Steud. ex O. Berg) Miers, I, K, M Eschweilera tessmannii R. Knuth, D Eschweilera truncata A.C. Sm., D Eschweilera wachenheimii (Benoist) Sandwith, I, K, M, D Gustavia augusta L., I, M, S Gustavia elliptica S.A.Mori, D Gustavia hexapetala (Aubl.) Sm., I, S, D Lecythis alutacea (A.C. Sm.) S.A. Mori, I, M Lecythis amara Aubl., S Lecythis assuncaoi S.A.Mori ined., D Lecythis chartacea O. Berg, K, S Lecythis confertiflora (A.C. Sm.) S.A. Mori, I, M, S Lecythis corrugata Poit., I, K, M, S Lecythis gracieana S.A.Mori, D Lecythis holcogyne (Sandwith) S.A. Mori, I, M, S Lecythis idatimon Aubl., K, S Lecythis parvifructa S.A.Mori, D Lecythis persistens Sagot, S Lecythis pisonis Cambess., D Lecythis poiteaui O. Berg, S, D Lecythis prancei S.A.Mori, D Lecythis retusa Spruce ex ) O. Berg, D Lecythis zabucajo Aubl., K, M, S, D Lentibulariaceae Genlisea filiformis A. St.-Hil., K Genlisea pygmaea A. St.-Hil., K Genlisea repens Benj., K Utricularia adpressa Salzm. ex A. St.-Hil. & Girard, K Utricularia amethystina Salzm. ex A. St.-Hil. & Girard, K Utricularia calycifida Benj., K Utricularia foliosa L., I Utricularia guyanensis A. DC., K Utricularia hispida Lam., K Utricularia humboldtii R.H. Schomb., K Utricularia jamesoniana Oliv., K Utricularia juncea Vahl, I, K Utricularia longeciliata A. DC., K Utricularia nana A. St.-Hil. & Girard, K Utricularia pubescens Sm., K Utricularia pusilla Vahl, I, K, M Utricularia sandwithii P. Taylor, K Utricularia simulans Pilg., K Utricularia subulata L., K Utricularia tenuissima Tutin, K Utricularia trichophylla Spruce ex Oliv., K Utricularia sp., D Liliaceae Crinum erubescens L.f. ex Sol., M Curculigo scorzonerifolia (Lam.) Baker, I Hippeastrum puniceum (Lam.) Kuntze, M, S Hymenocallis tubiflora Salisb., I, K, S Lissocarpaceae Lissocarpa guianensis Gleason, M Loganiaceae Antonia ovata Pohl, M, S Bonyunia aquatica Ducke, D Mitreola petiolata (J.F. Gmel.) Torr. & A. Gray, S Potalia amara Aubl., S, D Spigelia anthelmia L., D Spigelia humilis Benth., M Spigelia multispica Steud., I, M, S Strychnos amazonica Krukoff, D Strychnos asperula Sprague & Sandwith, D Strychnos bredemeyeri (Schult.) Sprague & Sandwith, M Strychnos cayennensis Krukoff & Barneby, S Strychnos cogens Benth., I, S, D Strychnos diaboli Sandwith, M Strychnos erichsonii M.R. Schomb. ex Progel, M, S Strychnos froesii Ducke, D Strychnos glabra Sagot ex Progel, S, D Strychnos guianensis (Aubl.) Mart., I, K, M, S Strychnos hirsuta Spruce ex Benth., M Strychnos jobertiana Baill., D Strychnos medeola Sagot ex Progel, S Strychnos melinoniana Baill., M, S Strychnos mitscherlichii M.R. Schomb., K, M, D Strychnos oiapocensis Fróes, S Strychnos panurensis Sprague & Sandwith, S Strychnos parviflora Spruce ex Benth., D Strychnos peckii B.L. Rob., I, S, D Strychnos subcordata Spruce ex Benth., M, D Strychnos tomentosa Benth., M, S Strychnos toxifera R.H. Schomb. ex Benth., S Lomariopsidaceae Bolbitis nicotianifolia (Sw.) Alston, S Bolbitis semipinnatifida (Fée) Alston, I, K, S Bolbitis serratifolia (Mert. ex Kaulf.) Schott, S I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

47 BRIT.ORG/SIDA 41 Elaphoglossum apodum (Kaulf.) Schott ex J. Sm., S Elaphoglossum flaccidum (Fée) T. Moore, S Elaphoglossum glabellum J. Sm., K, M, S, D Elaphoglossum herminieri (Bory & Fée ex Fée) T. Moore, S Elaphoglossum laminarioides (Bory ex Fée) T. Moore, M, S Elaphoglossum longicaudatum Mickel, S Elaphoglossum luridum (Fée) H. Christ, M, S Elaphoglossum macrophyllum (Mett. ex Kuhn) H. Christ, K, D Elaphoglossum pteropodum (Sodiro) C. Chr., S Elaphoglossum pteropus C. Chr., S, D Elaphoglossum revolutum (Liebm.) T. Moore, S Elaphoglossum rigidum (Aubl.) Urb., S, D Elaphoglossum schomburgkii (Fée) T. Moore, D Elaphoglossum strictum (Raddi) T. Moore, S Lomagramma guianensis (Aubl.) Ching, S Lomariopsis japurensis (Mart.) J. Sm., I, S Lomariopsis sp., D Loranthaceae Cladocolea micrantha (Eichler) Kuijt, K, D Cladocolea sandwithii (Maguire) Kuijt, K Oryctanthus alveolatus (Kunth) Kuijt, I, S Oryctanthus florulentus (Rich.) Tiegh., I, M, S Phthirusa disjectifolia (Rizzini) Kuijt, M Phthirusa guyanensis Eichler, K, M Phthirusa ovata (DC.) Eichler, D Phthirusa pycnostachya Eichler, S Phthirusa pyrifolia (Kunth) Eichler, S Phthirusa retroflexa (Ruiz & Pav.) Kuijt, M Phthirusa rufa (Mart.) Eichler, I, K, M, D Phthirusa stelis (L.) Kuijt, I, M, S, D Psittacanthus cinctus Mart., D Psittacanthus corynocephalus Eichler, S, D Psittacanthus lasianthus Sandwith, K Psittacanthus peronopetalus Eichler, D Psittacanthus siphon, vel aff. Eichler, D Struthanthus dichotrianthus Eichler, S Struthanthus syringifolius (Mart.) Mart., S Struthanthus sp., M Lycopodiaceae Huperzia dichotoma (Jacq.) Trevis., M, S Huperzia funiformis (Cham. ex Spring) Trevis., K Huperzia linifolia (L.) Trevis., S Huperzia taxifolia (Sw.) Trevis., S Lycopodiella caroliniana (L.) Pic. Serm., I, K Lycopodiella cernua (L.) Pic. Serm., I, K, M, S, D Lycopodium sp., S Lygodiaceae Lygodium micans J.W. Sturm, I, K Lygodium venustum Sw., K, D Lygodium volubile Sw., I, K, M Lygodium sp., S Lythraceae Cuphea antisyphilitica Kunth, I, D Cuphea carthagenensis (Jacq.) J.F. Macbr., S Cuphea ramulosa Mart. ex Koehne, D Cuphea sp., K Malpighiaceae Banisteriopsis carolina W.R. Anderson, S Banisteriopsis lucida (Rich.) Small, K Banisteriopsis martiniana (A. Juss.) Cuatrec., I, K, M Banisteriopsis pulcherrima (Sandwith) B. Gates, K Banisteriopsis schwannioides (Griseb.) B. Gates, S Banisteriopsis wurdackii B. Gates, S Banisteriopsis sp., D Bunchosia argentea (Jacq.) DC., I Bunchosia decussiflora W.R. Anderson, S Bunchosia glandulifera (Jacq.) Kunth, S Burdachia sphaerocarpa A. Juss., M Byrsonima aerugo Sagot, I, M, S Byrsonima altissima DC., K Byrsonima christianeae W.R. Anderson, K Byrsonima chrysophylla Kunth, D Byrsonima concinna Benth., K Byrsonima crassifolia (L.) Kunth, I, K, M Byrsonima crispa A. Juss., D Byrsonima densa (Poir.) DC., S Byrsonima duckeana W.R. Anderson, D Byrsonima eugeniifolia Sandwith, I, K, M Byrsonima gymnocalycina A. Juss., K, M Byrsonima incarnata Sandwith, K, D Byrsonima poeppigiana A. Juss., D Byrsonima spicata (Cav.) DC., K Byrsonima stipulacea A. Juss., I, K, M, S Glandonia macrocarpa Griseb., D Heteropterys cristata Benth., I, M Heteropterys macradena (DC.) W.R. Anderson, M Heteropterys macrostachya A. Juss., M Heteropterys multiflora (DC.) Hochr., I, M Heteropterys nervosa A. Juss., I, M Heteropterys oligantha W.R. Anderson, S Heteropterys velutina W.R. Anderson, I Heteropterys sp., D Hiraea adenophora Sandwith, M Hiraea affinis Miq., M, S Hiraea fagifolia (DC.) A. Juss., M, S Hiraea faginea (Sw.) Nied., I, M Hiraea gracieana W.R. Anderson, I, S Hiraea morii W.R. Anderson, S Jubelina rosea (Miq.) Nied., S Malpighia emarginata DC., S Mascagnia bracteosa Griseb., D Mascagnia divaricata (Kunth) Nied., S Mascagnia guianensis W.R. Anderson, M Mascagnia sepium (A. Juss.) Griseb., I, M Mezia includens (Benth.) Cuatrec., M, S Pterandra arborea Ducke, D Spachea elegans (G. Mey.) A. Juss., I Stigmaphyllon sinuatum (DC.) A. Juss., I, K, M, S, D Tetrapterys acutifolia Cav., M, S Tetrapterys calophylla A. Juss., M Tetrapterys crispa A. Juss., I, M, S Tetrapterys discolor (G. Mey.) DC., I, S Tetrapterys fimbripetala A. Juss., I, M Tetrapterys glabrifolia (Griseb.) Small, S Tetrapterys megalantha W.R. Anderson, S Tetrapterys mucronata Cav., I, M, S, D Tetrapterys styloptera A. Juss., I, K, M I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

48 42 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana Malvaceae Hibiscus bifurcatus Cav., I Hibiscus furcellatus Desr., I Hibiscus sororius L.f., I Pavonia cancellata (L.) Cav., D Pavonia castanaefolia A. St.-Hil. & Naudin, I, S Peltaea trinervis (C. Presl) Krapov. & Cristóbal, I Sida cordifolia L., I, K Sida glomerata Cav., I, M Sida rhombifolia L., S, D Sida setosa Mart. ex Colla, S Urena lobata L., I, M Wissadula patens (A. St.-Hil.) Garcke, I Marantaceae Calathea altissima (Poepp. & Endl.) Körn., D Calathea cyclophora Baker, K Calathea cylindrica (Roscoe) K. Schum., S Calathea elliptica (Roscoe) K. Schum., I, S Calathea legrelleana (Linden) Regel, I Calathea lutea (Aubl.) Schult., I Calathea maasiorum H. Kenn., S Calathea mansonis Körn., S Calathea micans (Mathieu) Körn., S Calathea microcephala (Poepp. & Endl.) Körn., S Calathea propinqua (Poepp. & Endl.) Körn., S Calathea splendida (Verschaff. ex Lem.) Regel, S Calathea taeniosa Joriss., D Calathea zingiberina Körn., S, D Calathea sp. nov., D Hylaeanthe hexantha (Poepp. & Endl.) A.M.E. Jonker & Jonker, S Hylaeanthe unilateralis (Poepp. & Endl.) A.M.E. Jonker & Jonker, S Ischnosiphon arouma (Aubl.) Körn., I, K, S Ischnosiphon centricifolius L. Andersson, S Ischnosiphon enigmaticus L. Andersson, S Ischnosiphon foliosus Gleason, I, K, M Ischnosiphon gracilis (Rudge) Körn., I, K, M, S, D Ischnosiphon killipii J.F. Macbr., D Ischnosiphon martianus Eichler ex Petersen, D Ischnosiphon obliquus (Rudge) Körn., I, K, M, S Ischnosiphon petiolatus (Rudge) L. Andersson, K, S Ischnosiphon puberulus Loes., I, K, M, S, D Maranta amplifolia K. Schum., S Maranta friedrichsthaliana Körn., S Maranta gibba Sm., I Maranta humilis Aubl., S Maranta protracta Miq., I Maranta rupicola L. Andersson, S Monophyllanthe oligophylla K. Schum., S Monotagma angustissimum Loes, D Monotagma contractum Huber, S Monotagma densiflorum (Körn.) K.Schum., D Monotagma laxum (Poepp. & Endl.) K. Schum., D Monotagma plurispicatum (Körn.) K. Schum., S Monotagma secundum (Petersen) K. Schum., D Monotagma spicatum (Aubl.) J.F. Macbr., I, K, M, S, D Monotagma tomentosum K. Schum. ex. Loes, D Monotagma vaginatum Hagberg, D Stromanthe tonckat (Aubl.) Eichler, S Thalia geniculata L., K Thymocarpus cannoideus Steyerm., D Marattiaceae Danaea elliptica Sm., I, S, D Danaea nodosa (L.) Sm., S Danaea simplicifolia Rudge, I, K, S, D Marcgraviaceae Marcgravia coriacea Vahl, K, M Marcgravia crenata Poepp. ex Wittm., D Marcgravia parviflora Rich. ex Wittm., I, M, D Marcgravia sp., D Marcgravia sp., S Marcgraviastrum sp., D Norantea guianensis Aubl., I, K, M, S Norantea pendula Lanj. & Heerdt, K Norantea sp., D Souroubea guianensis Aubl., I, K, M, S Souroubea sp., D Mayacaceae Mayaca longipes Mart. ex Seub., K Mayaca sellowiana Kunth, S Melastomataceae Aciotis aequatorialis Cogn., S, D Aciotis annua (Mart ex. DC.) Triana, K Aciotis circaeifolia (Bonpl.) Triana, M, D Aciotis fragilis (DC.) Cogn., M Aciotis laxa (DC.) Cogn., I, K, M, S Aciotis ornata (Miq.) Gleason, M Aciotis purpurascens (Aubl.) Triana, I, K, M, S, D Aciotis rubricaulis (Mart. ex DC.) Triana, S Adelobotrys adscendens (Sw.) Triana, S, D Adelobotrys marginata Brade, D Adelobotrys scandens (Aubl.) DC., D Appendicularia thymifolia (Bonpl.) DC., K Bellucia dichotoma Cogn., D Bellucia grossularioides (L.) Triana, I, K, M, S, D Bellucia pentamera Naudin, I Blakea rosea (Ruiz & Pav.) D. Don, D Clidemia anisophylla DC., S Clidemia bullosa DC., M, D Clidemia capitata Benth., K Clidemia capitellata (Bonpl.) D. Don, K Clidemia charadrophylla Tutin, K Clidemia conglomerata DC., I, K, M, S Clidemia dentata D. Don, S Clidemia epibaterium DC., K, D Clidemia hirta (L.) D. Don, I, K, S, D Clidemia involucrata DC., K, M, S Clidemia japurensis DC., K, D Clidemia laevifolia Gleason, S Clidemia micrantha Sagot, K Clidemia minutiflora (Triana) Cogn., K, M Clidemia novemnervia (DC.) Triana, K, D Clidemia octona (Bonpl.) L.O. Williams, S Clidemia ostentata Wurdack, K Clidemia pycnaster Tutin, K Clidemia rubra (Aubl.) Mart., K, S, D Clidemia saulensis Wurdack, M, S Clidemia septuplinervia Cogn., S Clidemia sericea D. Don, I, K Clidemia silvicola Gleason, K Clidemia strigillosa (Sw.) DC., K I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

49 BRIT.ORG/SIDA 43 Comolia microphylla Benth., K Comolia vernicosa (Benth.) Triana, I, K, M Comolia villosa (Aubl.) Triana, I, K, S Desmoscelis villosa (Aubl.) Naudin, S Ernestia cataractae Tutin, K Ernestia glandulosa Gleason, S Ernestia granvillei Wurdack, S Graffenrieda caryophyllea Triana, K Graffenrieda irwinii Wurdack, K Graffenrieda weddellii Naudin, K Henriettea granulata Berg ex Triana, D Henriettea multiflora Naudin, I, K, M Henriettea ramiflora (Sw.) DC., K, M, S, D Henriettea stellaris O. Berg ex Triana, I, M Henriettea succosa (Aubl.) DC., S Henriettella caudata Gleason, K, S, D Henriettella duckeana Hoehne, S, D Leandra agrestis (Aubl.) Raddi, S Leandra candelabrum (J.F. Macbr.) Wurdack, D Leandra clidemioides (Naudin) Wurdack, S Leandra divaricata (Naudin) Cogn., K, M, S Leandra micropetala (Naudin) Cogn., K, S, D Leandra paleacea Wurdack, S Leandra purpurea Gleason, K, M Leandra rufescens (DC.) Cogn., K, S Leandra sanguinea Gleason, K Leandra secunda (D. Don) Cogn., D Leandra solenifera Cogn., I, K, S Loreya arborescens (Aubl.) DC., S Loreya mespiloides Miq., M, S Loreya riparia S.S.Renner, D Loreya spruceana Benth. ex Triana, D Loreya subrotundifolia (Wurdack) S.S. Renner, S Macairea pachyphylla Benth., I, K Macairea thyrsiflora DC., K Macrocentrum cristatum (DC.) Triana, I, K, M, S Macrocentrum droseroides Triana, K Macrocentrum fasciculatum (Rich. ex DC.) Triana, S Macrocentrum latifolium Wurdack, S Macrocentrum repens (Gleason) Wurdack, K Macrocentrum vestitum Sandwith, K Maieta guianensis Aubl., S, D Maieta poeppigii Mart. ex Cogn., I, K, D Meriania urceolata Triana, K Miconia acuminata (Steud.) Naudin, S Miconia affinis DC., S, D Miconia alata (Aubl.) DC., I, S, D Miconia aliquantula Wurdack, S Miconia ampla Triana, D Miconia aplostachya (Bonpl.) DC., I Miconia argyrophylla DC., I, M, S, D Miconia astroplocama Donn. Sm., D Miconia biglandulosa Gleason, D Miconia bracteata (DC.) Triana, I, K, M, S Miconia cacatin (Aubl.) S.S. Renner, S Miconia campestris (Benth.) Triana, K Miconia ceramicarpa (DC.) Cogn., I, M, S, D Miconia chrysophylla (Rich.) Urb., I, S Miconia ciliata (Rich.) DC., I, K, S Miconia cuspidata Mart. ex Naudin, D Miconia diaphanea Gleason, S Miconia dispar Benth., S, D Miconia dodecandra (Desr.) Cogn., K Miconia duckei Cogn., D Miconia egensis Cogn., K, D Miconia elaeagnoides Cogn., D Miconia eriodonta DC., S, D Miconia fallax DC., I Miconia gratissima Benth. ex Triana, I, K, M, S, D Miconia holosericea (L.) DC., S, D Miconia hypoleuca (Benth.) Triana, M Miconia ibaguensis (Bonpl.) Triana, D Miconia lateriflora Cogn., S Miconia lepidota DC., K, D Miconia longispicata Triana, D Miconia macrothyrsa Benth., I Miconia maguirei Gleason, K Miconia marginata Triana, K Miconia mariae Wurdack, I, K Miconia minutiflora (Bonpl.) DC., S, D Miconia mirabilis (Aubl.) L.O. Williams, M, S Miconia myriantha Benth., I, K, S Miconia nervosa (Sm.) Triana, M, S, D Miconia phanerostila Pilg., D Miconia platypoda Gleason, D Miconia plukenetii Naudin, K, S Miconia poeppigii Triana, D Miconia polita Gleason, K Miconia prasina (Sw.) DC., I, K, M, S Miconia pubipetala Miq., D Miconia punctata (Desr.) D. Don ex DC., M, D Miconia pyrifolia Naudin, D Miconia racemosa (Aubl.) DC., K, S Miconia regelii Cogn., D Miconia rubiginosa (Bonpl.) DC., M Miconia rufescens (Aubl.) DC., I Miconia sastrei Wurdack, S Miconia schwackei Cogn., D Miconia serrulata (DC.) Naudin, M Miconia spichigeri Wurdack, D Miconia splendens (Sw.) Griseb., D Miconia tetraspermoides Wurdack, D Miconia tillettii Wurdack, S Miconia tomentosa (Rich.) D. Don ex DC., S, D Miconia traillii Cogn., S, D Miconia trimera Wurdack, S Mouriri acutiflora Naudin, M Mouriri angulicosta Morley, D Mouriri brachyanthera Ducke, D Mouriri collocarpa Ducke, S, D Mouriri crassifolia Sagot, S Mouriri dimorphandra Morley, D Mouriri duckeana Morley, S, D Mouriri duckeanoides Morley, D Mouriri ficoides Morley, D Mouriri grandiflora DC., I, D Mouriri huberi Cogn., D Mouriri lunatanthera Morley, D Mouriri nervosa Pilg., D Mouriri nigra (DC.) Morley, D Mouriri oligantha Pilg., D Mouriri sagotiana Triana, S I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

50 44 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana Mouriri sideroxylon Sagot ex Triana, S Mouriri trunciflora Ducke, D Mouriri sp., K Myriaspora egensis Mart. ex DC., I, K Nepsera aquatica (Aubl.) Naudin, K, S, D Phainantha laxiflora (Triana) Gleason, K Pterolepis glomerata (Rottb.) Miq., K Rhynchanthera grandiflora (Aubl.) DC., S Siphanthera cordifolia (Benth.) Gleason, K Siphanthera hostmannii Cogn., K Tibouchina aspera Aubl., I, M, S Tibouchina fraterna N.E. Br., K Tibouchina multiflora (Gardner) Cogn., S Tococa aristata Benth., I, K, M Tococa bullifera Mart. & Schrank ex DC., D Tococa caryophyllaea (DC.) S.S. Renner, D Tococa desiliens Gleason, K Tococa guianensis Aubl., M, D Tococa nitens (Benth.) Triana, I, K Tococa subciliata (DC.) Triana, I, M Topobea parasitica Aubl., S Meliaceae Carapa guianensis Aubl., I, K, M, D Carapa procera A. DC., I, M, S, D Cedrela fissilis Vell., D Cedrela odorata L., M, S Guarea carinata Ducke, D Guarea cinnamomea Harms, D Guarea convergens T.D.Penn., D Guarea gomma Pulle, K, S Guarea grandifolia A. DC., S Guarea guidonia (L.) Sleumer, I Guarea humaitensis T.D.Penn., D Guarea kunthiana A. Juss., I, M, S Guarea macrophylla Vahl, K, D Guarea michel-moddei T.D. Penn. & S.A. Mori, S Guarea pubescens (Rich.) A. Juss., I, M, S, D Guarea purusana C. DC., D Guarea scabra A. Juss., S, D Guarea silvatica C. DC., S, D Guarea trunciflora C. DC., D Trichilia areolata T.D.Penn., D Trichilia cipo (A. Juss.) C. DC., I, S, D Trichilia euneura C. DC., S Trichilia martiana C. DC., I Trichilia micrantha Benth., S, D Trichilia micropetala T.D. Penn., D Trichilia pallida Sw., I, S, D Trichilia pleeana (A. Juss.) C. DC., D Trichilia quadrijuga Kunth, I, S Trichilia rubra C. DC., D Trichilia schomburgkii C. DC., I, S, D Trichilia septentrionalis C. DC., S, D Trichilia sp., M Mendonciaceae Mendoncia bivalvis (L.f.) Merr., S Mendoncia glabra (Poepp. & Endl.) Nees, S Mendoncia hoffmannseggiana Nees, I, S, D Mendoncia pedunculata Leonard, D Mendoncia squamuligera Nees, I, M, S Menispermaceae Abuta barbata Miers, M Abuta bullata Moldenke, K, M, S Abuta candollei Triana & Planch., M, D Abuta grandifolia (Mart.) Sandwith, M, S, D Abuta grisebachii Triana & Planch., D Abuta imene (Mart.) Eichler, K, M, S, D Abuta obovata Diels, M, D Abuta pahni (Mart.) Krukoff & Barneby, D Abuta panurensis Eichler, D Abuta rufescens Aubl., M, S, D Abuta sandwithiana Krukoff & Barneby, M, D Abuta solimoesensis Krukoff & Barneby, D Abuta velutina Gleason, M Abuta sp., I Anomospermum chloranthum Diels, S Anomospermum grandifolium Eichler, M Anomospermum solimoesanum (Moldenke) Krukoff & Barneby, D Anomospermum steyermarkii Krukoff & Barneby, S Caryomene olivascens Barneby & Krukoff, S Cissampelos andromorpha DC., I, M, D Curarea candicans (Rich. ex DC.) Barneby & Krukoff, I, M, S Disciphania lobata Eichler, D Disciphania moriorum Barneby, sp. nov. ined., S Elephantomene eburnea Barneby & Krukoff, S Odontocarya amazonum Barneby, D Odontocarya tamoides (DC.) Miers, M Odontocarya wullschlaegelii (Eichler) Barneby, D Orthomene hirsuta (Krukoff & Moldenke) Barneby & Krukoff, D Orthomene prancei Barneby & Krukoff, S Orthomene schomburgkii (Miers) Barneby & Krukoff, I Sciadotenia cayennensis Benth., I, M, S Sciadotenia duckei Moldenke, D Sciadotenia sprucei Diels, D Telitoxicum inopinatum Moldenke, M, S Telitoxicum krukovii Moldenke, M Telitoxicum minutiflorum (Diels) Moldenke, M, D Telitoxicum rodriguesii Krukoff, D Metaxyaceae Metaxya rostrata (Kunth) C. Presl, I, K, M, S, D Mimosaceae Abarema adenophora (Ducke) Barneby & J.W. Grimes, D Abarema barbouriana (Standl.) Barneby & J.W. Grimes, S Abarema campestris (Spruce ex Benth.) Barneby & J.W. Grimes, D Abarema cochleata (Willd.) Barneby & J.W. Grimes, D Abarema curvicarpa (H.S. Irwin) Barneby & J.W. Grimes, K, S, D Abarema floribunda (Spruce ex Benth.) Barneby & J.W. Grimes, D Abarema jupunba (Willd.) Britton & Killip, I, K, M, S, D Abarema laeta (Benth.) Barneby & J.W. Grimes, S, D Abarema mataybifolia (Sandwith) Barneby & J.W. Grimes, I, S, D Abarema microcalyx (Spruce ex Benth.) Barneby & J.W. Grimes, D I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

51 BRIT.ORG/SIDA 45 Abarema piresii Barneby & J.W. Grimes, D Acacia altiscandens Ducke, D Acacia multipinnata Ducke, D Acacia tenuifolia (L.) Willd., I, S Albizia subdimidiata (Splitg.) Barneby & J.W. Grimes, I Balizia elegans (Ducke) Barneby & J.W. Grimes, D Balizia pedicellaris (DC.) Barneby & J.W. Grimes, I, M, S Calliandra laxa (Willd.) Benth., K Calliandra surinamensis Benth., K Calliandra tenuiflora Benth., D Cedrelinga cateniformis (Ducke) Ducke, I, S, D Entada polyphylla Benth., S, D Enterolobium barnebianum A.L. Mesquita & M.F. Silva, M Enterolobium cyclocarpum (Jacq.) Griseb., M Enterolobium schomburgkii (Benth.) Benth., S, D Hydrochorea corymbosa (Rich.) Barneby & J.W. Grimes, I, K Hydrochorea gonggrijpii (Kleinhoonte) Barneby & J.W. Grimes, K Inga acreana Harms, S Inga acrocephala Steud., S Inga alata Benoist, I, S Inga alba (Sw.) Willd., I, K, M, S, D Inga albicoria Poncy, S Inga auristellae Harms, I, S Inga bicoloriflora Ducke, D Inga bourgonii (Aubl.) DC., S Inga brachystachys Ducke, S Inga bracteosa Benth., S Inga capitata Desv., I, S, D Inga cayennensis Sagot ex Benth., D Inga chrysantha Ducke, D Inga cordatoalata Ducke, D Inga disticha Benth., I, M Inga edulis Mart., S, D Inga fastuosa (Jacq.) Willd., S Inga glomeriflora Ducke, D Inga graciliflora Benth., S Inga gracilifolia Ducke, D Inga grandiflora Ducke, D Inga huberi Ducke, M, S, D Inga java Pittier, S Inga lateriflora Miq., M, D Inga laurina (Sw.) Willd., D Inga leiocalycina Benth., I, S, D Inga longiflora Spruce ex Benth., D Inga longipedunculata Ducke, S Inga macrophylla Humb. & Bonpl. ex Willd., D Inga meissneriana Miq., M Inga melinonis Sagot, D Inga nobilis Willd., I, K, M, S Inga nouragensis Poncy, S Inga nubium Poncy, S Inga nuda Salzm. ex Benth., S Inga obidensis Ducke, D Inga panurensis Spruce ex Benth., D Inga paraensis Ducke, S, D Inga pezizifera Benth., I, S, D Inga poeppigiana Benth., S Inga retinocarpa Poncy, S Inga rhynchocalyx Sandwith, S, D Inga rubiginosa (Rich.) DC., I, S, D Inga sarmentosa Glaz. ex Harms, S Inga semialata (Vell.) Mart., I, S, D Inga sertulifera DC., I, K, M Inga splendens Willd., S, D Inga stipularis DC., M, S, D Inga suberosa T.D.Penn., D Inga thibaudiana DC., I, K, M, S, D Inga umbellifera (Vahl) Steud., I, S, D Inga umbratica Poepp. & Endl., D Inga virgultosa (Vahl) Desv., S Macrosamanea discolor (Humb. & Bonpl. ex Willd.) Britton & Rose ex Britton & Killip, M Macrosamanea pubiramea (Steud.) Barneby & J.W. Grimes, I, M, D Macrosamanea sp., K Mimosa guilandinae (DC.) Barneby, S, D Mimosa myriadenia (Benth.) Benth., I, K, M, S Mimosa pellita Humb. & Bonpl. ex Willd., I Mimosa pigra L., I Mimosa polydactyla Humb. & Bonpl. ex Willd., S Mimosa pudica L., I, S, D Parkia decussata Ducke, S, D Parkia igneiflora Ducke, D Parkia multijuga Benth., D Parkia nitida Miq., M, S, D Parkia panurensis Spruce ex H.C.Hopkins, D Parkia pendula (Willd.) Benth. ex Walp., M, S, D Parkia reticulata Ducke, S Parkia ulei (Harms) Kuhlm., M, S Parkia velutina Benoist, S, D Pentaclethra macroloba (Willd.) Kuntze, I, K, M Piptadenia floribunda Kleinhoonte, S Piptadenia minutiflora Ducke, D Pseudopiptadenia psilostachya (DC.) G.P. Lewis & M.P. Lima, S, D Pseudopiptadenia suaveolens (Miq.) J.W. Grimes, M, S Stryphnodendron guianense (Aubl.) Benth., S, D Stryphnodendron moricolor Barneby & J.W. Grimes, S Stryphnodendron occhionianum E.M. Martins, D Stryphnodendron polystachyum (Miq.) Kleinhoonte, S Stryphnodendron pulcherrimum (Willd.) Hochr., D Stryphnodendron racemiferum (Ducke) W.A. Rodrigues, D Zygia basijuga (Ducke) Barneby & J.W. Grimes, K, D Zygia cataractae (Kunth) L. Rico, I Zygia latifolia (L.) Fawc. & Rendle, I, M, D Zygia potaroensis Barneby & J.W. Grimes, K Zygia racemosa (Ducke) Barneby & J.W. Grimes, M, S, D Zygia tetragona Barneby & J.W. Grimes, S Monimiaceae Mollinedia grazielae Peixoto, I Mollinedia laurina Tul., I, K, S Mollinedia ovata Ruiz & Pav., S, D Siparuna cristata (Poepp. & Endl.) A. DC., D Siparuna cuspidata (Tul.) A. DC., I, S, D Siparuna decipiens (Tul.) A. DC., I, M, S, D I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

52 46 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana Siparuna glycycarpa (Ducke) S.S. Renner, D Siparuna guianensis Aubl., I, K, M, S, D Siparuna monogyna Jang., D Siparuna pachyantha A.C. Sm., S Siparuna poeppigii (Tul.) A. DC., S, D Siparuna reginae (Tul.) A. DC., D Siparuna sarmentosa Perkins, D Siparuna sprucei A. DC., S Moraceae Bagassa guianensis Aubl., M, S Brosimum acutifolium Huber, S, D Brosimum guianense (Aubl.) Huber, I, K, M, S, D Brosimum lactescens (S. Moore) C.C. Berg, I, S, D Brosimum longifolium Ducke, D Brosimum parinarioides Ducke, S, D Brosimum potabile Ducke, D Brosimum rubescens Taub., M, S, D Brosimum utile (Kunth) Pittier, I, S, D Clarisia ilicifolia (Spreng.) Lanj. & Rossberg, I, M, S, D Clarisia racemosa Ruiz & Pav., D Ficus albert-smithii Standl., K, D Ficus amazonica (Miq.) Miq., S, D Ficus broadwayi Urb., K Ficus cremersii C.C. Berg, D Ficus duckeana C.C. Berg, D Ficus gomelleira Kunth & Bouché, M, S, D Ficus greiffiana Dugand, D Ficus guianensis Desv., K, M, S, D Ficus hebetifolia Dugand, D Ficus insipida Willd., S Ficus krukovii Standl., D Ficus leiophylla C.C. Berg, S Ficus malacocarpa Standl., I, K Ficus mathewsii (Miq.) Miq., K, D Ficus maxima Mill., I, S, D Ficus nymphaeifolia Mill., S Ficus obtusifolia Kunth, M, D Ficus pakkensis Standl., D Ficus paludica Standl., M Ficus panurensis Standl., K, M, S, D Ficus paraensis (Miq.) Miq., I, K, M, D Ficus pertusa L.f., S, D Ficus piresiana Vazq. Avila & C.C. Berg, D Ficus schumacheri (Liebm.) Griseb., S Ficus subapiculata (Miq.) Miq., D Ficus trigona L.f., I, D Helicostylis elegans (Macbr.) C.C. Berg, D Helicostylis pedunculata Benoist, S, D Helicostylis scabra (Macbr.) C.C. Berg, D Helicostylis tomentosa (Poepp. & Endl.) Rusby, I, K, M, S, D Helicostylis turbinata C.C. Berg, D Maquira calophylla (Poepp. & Endl.) C.C. Berg, D Maquira guianensis Aubl., S, D Maquira sclerophylla (Ducke) C.C. Berg, S, D Naucleopsis caloneura (Huber) Ducke, D Naucleopsis guianensis (Mildbr.) C.C. Berg, M, S Naucleopsis stipularis Ducke, D Naucleopsis ternstroemiiflora (Mildbr.) C.C. Berg, D Naucleopsis ulei (Warb.) Ducke, D Perebea guianensis Aubl., S Perebea mollis (Poepp. & Endl.) Huber, D Perebea rubra (Trécul) C.C. Berg, S Pseudolmedia laevigata Trécul, S, D Pseudolmedia laevis (Ruiz & Pav.) J.F. Macbr., I, S, D Sorocea guilleminiana Gaudich., D Sorocea muriculata Miq., D Sorocea pubivena Hemsley, I, D Trymatococcus amazonicus Poepp. & Endl., S, D Trymatococcus oligandrus (Benoist) Lanj., S Trymatococcus paraensis Ducke, M Myristicaceae Compsoneura ulei Warb., S, D Iryanthera coriacea Ducke, D Iryanthera dialyandra Ducke, D Iryanthera elliptica Ducke, D Iryanthera juruensis Warb., I, S, D Iryanthera laevis Markgr., D Iryanthera lancifolia Ducke, I, M, D Iryanthera macrophylla (Benth.) Warb., I, D Iryanthera paradoxa (Schwacke) Warb., D Iryanthera paraensis Huber, I, D Iryanthera polyneura Ducke, D Iryanthera sagotiana (Benth.) Warb., M, S, D Iryanthera tessmannii Markgr., S Iryanthera ulei Warb., D Osteophloeum platyspermum (Spruce ex A. DC.) Warb., S, D Otoba sp., M Virola caducifolia W.A. Rodrigues, D Virola calophylla (Spruce) Warb., D Virola divergens Ducke, D Virola elongata (Benth.) Warb., M Virola guggenheimii W.A. Rodrigues, D Virola kwatae Sabatier, S Virola michelii Heckel, M, S, D Virola mollissima (Poepp. ex A.DC.) Warb., D Virola multicostata Ducke, D Virola multinervia Ducke, D Virola pavonis (A. DC.) A.C. Sm., D Virola sebifera Aubl., M, S, D Virola surinamensis (Rol. ex Rottb.) Warb., I, M, S Virola venosa (Benth.) Warb., D Myrsinaceae Ardisia guianensis (Aubl.) Mez, I, S Cybianthus amplus (Mez) G. Agostini, K Cybianthus duidae (Gleason & Moldenke) G. Agostini, K Cybianthus fulvopulverulentus (Mez) G. Agostini, K, M Cybianthus fuscus Mart., S Cybianthus gracillimus Mez, M Cybianthus guyanensis (A. DC.) Miq., K, M, S, D Cybianthus longifolius Miq., D Cybianthus microbotrys A. DC., I, S Cybianthus pakaraimae Pipoly, sp. nov. ined., K Cybianthus potiaei (Mez) G. Agostini, K, S, D Cybianthus prieurii A. DC., K, M, S Cybianthus resinosus Mez, K, D Cybianthus surinamensis (Spreng.) G. Agostini, M, S Cybianthus venezuelanus Mez, K, M, S Stylogyne incognita Pipoly, S I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

53 BRIT.ORG/SIDA 47 Stylogyne micrantha (Kunth) Mez, D Stylogyne orinocensis Was Ardisia o.!, I Stylogyne rodriguesiana Pipoly, D Stylogyne schomburgkiana (A. DC.) Mez, I, M Myrtaceae Blepharocalyx eggersii (Kiaersk.) Landrum, D Calycolpus goetheanus (DC.) O. Berg, M, D Calycolpus revolutus (Schauer) O. Berg, M, D Calycolpus sp. 1, D Calycorectes bergii Sandwith, I Calyptranthes amshoffae McVaugh, K, S Calyptranthes bracteata M.L. Kawas. & B. Holst, S Calyptranthes creba McVaugh, D Calyptranthes cuspidata DC., D Calyptranthes forsteri O. Berg, M Calyptranthes macrophylla O. Berg, D Calyptranthes multiflora O. Berg, D Calyptranthes paniculata Ruiz & Pav., D Calyptranthes pullei Burret ex Amshoff, I Eugenia agathopoda Diels, D Eugenia albicans (O. Berg) Urb., I, S Eugenia anastomosans DC., I, K, D Eugenia arawakorum Sandwith, I Eugenia argyrophylla B. Holst & M.L. Kawas., S Eugenia armeniaca Sagot, S Eugenia biflora (L.) DC., I, D Eugenia citrifolia Poir., D Eugenia coffeifolia DC., I, M, S Eugenia conjuncta Amshoff, M Eugenia cribrata McVaugh, I Eugenia cucullata Amshoff, I, K, M Eugenia cupulata Amshoff, D Eugenia cuspidifolia DC., D Eugenia diplocampta Diels, D Eugenia egensis DC., I Eugenia excelsa O. Berg, M Eugenia feijoi O. Berg, S, D Eugenia ferreiraeana O. Berg, S, D Eugenia florida DC., I, D Eugenia gongylocarpa M.L. Kawas. & B. Holst, S Eugenia illepida McVaugh, D Eugenia kaieteurensis Amshoff, K Eugenia lambertiana DC., I Eugenia latifolia Aubl., I Eugenia ligustrina (Sw.) Willd., I Eugenia longiracemosa Kiaersk., D Eugenia luciae Amshoff, I Eugenia macrocalyx (Rusby) McVaugh, K, S Eugenia malaccensis L., D Eugenia mimus McVaugh, I, S Eugenia morii B. Holst & M.L. Kawas., S Eugenia muricata DC., S Eugenia omissa McVaugh, D Eugenia patrisii Vahl, I, M, S, D Eugenia prosoneura O. Berg, D Eugenia protenta McVaugh, D Eugenia pseudopsidium Jacq., S, D Eugenia punicifolia (Kunth) DC., K, D Eugenia ramiflora Desv., D Eugenia stylaris McVaugh, D Eugenia subterminalis DC., D Eugenia tafelbergica Amshoff, I Eugenia tapacumensis O. Berg, I, K, M, D Eugenia tetramera (McVaugh) M.L. Kawas. & B. Holst, S Eugenia trinervia Vahl, I, M Eugenia wullschlaegeliana Amshoff, S Eugenia sp. nov., D Marlierea buxifolia Amshoff, K Marlierea caudata McVaugh, D Marlierea montana (Aubl.) Amshoff, M Marlierea salticola Amshoff, K Marlierea schomburgkiana O. Berg, M Marlierea umbraticola (Kunth) O. Berg, D Myrcia albidotomentosa (Amshoff) McVaugh, K Myrcia aliena McVaugh, D Myrcia amazonica DC., D Myrcia bracteata (Rich.) DC., D Myrcia calycampa Amshoff, I Myrcia citrifolia (Aubl.) Urb., D Myrcia cuprea (O. Berg) Kiaersk., D Myrcia decorticans DC., I, S Myrcia extranea McVaugh, K Myrcia fallax (Rich.) DC., K, M, S, D Myrcia fenestrata DC., S, D Myrcia gigas McVaugh, S, D Myrcia graciliflora Sagot, S Myrcia grandis McVaugh, D Myrcia guianensis (Aubl.) DC., I, D Myrcia huallagae McVaugh, D Myrcia inaequiloba (DC.) Legrand, I Myrcia magnoliifolia DC., S, D Myrcia minutiflora Sagot, S, D Myrcia paivae O. Berg, D Myrcia platyclada DC., K, S Myrcia pyrifolia (Desv. ex Ham.) Nied., I Myrcia quitarensis (Benth.) Sagot, I Myrcia rotundata (Amshoff) McVaugh, K Myrcia rufipila McVaugh, D Myrcia rupta M.L. Kawas. & B. Holst, S Myrcia saxatilis (Amshoff) McVaugh, S Myrcia servata McVaugh, D Myrcia subobliqua (Benth.) Nied., I, M Myrcia subsericea A. Gray, D Myrcia subsessilis O. Berg, S Myrcia sylvatica (G. Mey.) DC., I, K, M, S, D Myrciaria amazonica O. Berd, I Myrciaria dubia (Kunth) McVaugh, I Myrciaria floribunda (West ex Willd.) O. Berg, I, M, D Myrciaria vismeifolia (Benth.) O. Berg, I, M Myrciaria sp., K Plinia pinnata L., I Psidium acutangulum DC., I Psidium guineense Sw., I Psidium persoonii McVaugh, I, M, S, D Psidium striatulum DC., I, M Nyctaginaceae Guapira eggersiana (Heimerl) Lundell, K, M Guapira fragrans (Dum.-Cours.) Little, M Guapira salicifolia (Heimerl) Lundell, S Neea constricta Spruce ex J.A. Schmidt, M, S Neea floribunda Poepp. & Endl., I, M, S, D I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

54 48 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana Neea laxa Poepp. & Endl., D Neea madeirana Standl., D Neea mollis Spruce ex J.A. Schmidt, S Neea obovata Spruce ex Heimerl, D Neea oppositifolia Ruiz & Pav., D Neea ovalifolia Spruce ex J.A. Schmidt, S, D Neea verticillata Ruiz & Pav., D Pisonia glabra Heimerl, M Pisonia tomentosa Casar., D Pisonia sp., S Nymphaeaceae Nymphaea glandulifera Rodschied, S Nymphaea rudgeana G. Mey., K Nympheaceae Victoria amazonica (Poepp.) Sowerby, I Ochnaceae Cespedesia spathulata (Ruiz & Pav.) Planch., M, D Elvasia essequibensis Engl., K Ouratea candollei (Planch.) Tiegh., S Ouratea cardiosperma (Lam.) Engl., M Ouratea cernuiflora Sandwith, K Ouratea coccinea (Mart.) Engl., D Ouratea discophora Ducke, D Ouratea erecta Sastre, S Ouratea ferruginea Engl., D Ouratea gillyana (Dwyer) Sandwith & Maguire, K Ouratea guianensis Aubl., M Ouratea leblondii (Tiegh.) Lemée, M, S Ouratea mazaruniensis A.C. Sm. & Dwyer, K Ouratea odora (Poepp.) Engl., D Ouratea paraensis Huber, D Ouratea pendula (Poepp.) Engl., S Ouratea roraimae Engl., K Ouratea rupununiensis (Klotzsch) Engl., I Ouratea saulensis Sastre, S Ouratea schomburgkii (Planch.) Engl., I Ouratea scottii Sastre, S Ouratea superba Engl., I Poecilandra retusa Tul., K Sauvagesia aliciae Sastre, S Sauvagesia elata Benth., D Sauvagesia erecta L., I, K, M, S, D Sauvagesia rubiginosa A. St.-Hil., I Sauvagesia sprengelii A. St.-Hil., K Sauvagesia tafelbergensis Sastre, S Olacaceae Aptandra tubicina (Poepp.) Benth. ex Miers, D Cathedra acuminata (Benth.) Miers, S Chaunochiton kappleri (Sagot ex Engl.) Ducke, M, S, D Dulacia candida (Poepp.) Kuntze, D Dulacia crassa (Monach.) Sleumer, K Dulacia guianensis (Engl.) Kuntze, I, K, M, S, D Heisteria barbata Cuatrec., D Heisteria cauliflora Sm., I, K, M, S Heisteria densifrons Engl., I, K, S, D Heisteria duckei Sleumer, D Heisteria laxiflora Engl., D Heisteria maguirei Sleumer, K Heisteria scandens Ducke, S Maburea trinervis Maas, I, M Minquartia guianensis Aubl., M, S, D Ptychopetalum olacoides Benth., D Oleandraceae Nephrolepis biserrata (Sw.) Schott, M, S, D Nephrolepis cordifolia (L.) C. Presl, K Nephrolepis multiflora (Roxb.) F.M. Jarrett ex C.V. Morton, M Nephrolepis pectinata (Willd.) Schott, D Nephrolepis rivularis (Vahl) Mett. ex Krug, K, S, D Oleandra articulata (Sw.) C. Presl, I, K, S, D Oleandra pilosa Hook., K Onagraceae Ludwigia affinis (DC.) H. Hara, S Ludwigia decurrens Walter, M Ludwigia dodecandra (DC.) Zardini & P.H. Raven, S Ludwigia erecta (L.) H. Hara, I, K, M, S Ludwigia hyssopifolia (G. Don) Exell, K, M, D Ludwigia leptocarpa (Nutt.) H. Hara, I, S Ludwigia octovalvis (Jacq.) P.H. Raven, K, S, D Ludwigia rigida (Miq.) Sandwith, I Ludwigia sericea (Cambess.) H. Hara, I Ophioglossaceae Ophioglossum palmatum L., D Opiliaceae Agonandra silvatica Ducke, M, D Orchidaceae Acacallis cyanea Lindl., D Aganisia pulchella Lindl., S Aspidogyne foliosa (Poepp. & Endl.) Garay, K, S Aspidogyne multifoliata (C. Schweinf.) Garay, K Batemannia colleyi Lindl., K, M, S, D Beloglottis sp., S Bifrenaria longicornis Lindl., I, M, D Bifrenaria rudolfii (Hoehne) Carnevali & Romero, D Bollea violacea (Lindl.) Rchb.f., I, K Brassavola martiana Lindl., I, M Brassia bidens Lindl., K Brassia caudata (L.) Lindl., S Brassia chloroleuca Barb. Rodr., S Brassia huebneri Schltr., S Brassia lanceana Lindl., S Brassia lawrenceana Lindl., I Brassia wageneri Rchb.f., K, M Bulbophyllum bracteolatum Lindl., S Caluera vulpina Dodson & Determann, S Campylocentrum fasciola (Lindl.) Cogn., I, M, D Campylocentrum micranthum (Lindl.) Rolfe, I, M, S, D Catasetum barbatum (Lindl.) Lindl., S, D Catasetum discolor (Lindl.) Lindl., K, M Catasetum gnomus Linden & Rchb.f., D Catasetum rivularium Barb.Rodr., D Catasetum tigrinum Rchb.f., D Cattleya violacea (Kunth) Rolfe, I Caularthron bicornutum (Hook.) Raf., M Caularthron bilamellatum (Rchb.f..) R.E. Schult., I Cheiradenia cuspidata Lindl., I, M, S Cleistes rosea Lindl., K, S Cochleanthes guianensis Lafontaine, G. Gerlach & Senghas, S Coryanthes sp., M I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

55 BRIT.ORG/SIDA 49 Cryptarrhena guatemalensis Schltr., S Cryptarrhena kegelii Rchb.f., I, S Cryptocentrum sp., K Cyclopogon olivaceus (Rolfe) Schltr., S Cycnoches haagii Barb. Rodr., S Degranvillea dermaptera Determann, S Dichaea ancoraelabia C. Schweinf., D Dichaea hookeri Garay & Sweet, K Dichaea kegelii Rchb.f., S Dichaea muricata (Sw.) Lindl., M, S Dichaea pendula (Aubl.) Cogn., S Dichaea picta Rchb.f., M Dichaea rendlei Gleason, K Dichaea splitgerberi Rchb.f., K Dichaea tenuis C. Schweinf., D Dichaea trulla Rchb.f., K, S Dichaea sp., I Dimerandra elegans (Focke) Siegerist, M, S Dimerandra sp., I Dryadella osmariniana (Braga) Garay & Dunsterv., D Elleanthus caravata (Aubl.) Rchb.f., I, K, M, S Elleanthus graminifolius (Barb. Rodr.) Lojtnant, K, M, S Encyclia aemula (Lindl.) Carnevali & I. Ramírez, M, S, D Encyclia calamaria (Lindl.) Pabst, M Encyclia chloroleuca (Hook.) Neumann, K Encyclia diurna (Jacq.) Schltr., M Encyclia granitica (Lindl.) Schltr., I, S Encyclia latipetala (C. Schweinf.) Pabst, K Encyclia pygmaea (Hook.) Dressler, M, S Encyclia tarumana Schltr., D Encyclia vespa (Vell.) Dressler, I, K, M Epidendrum anceps Jacq., M, S Epidendrum carpophorum Barb. Rodr., K, M, S Epidendrum compressum Griseb., K, M, D Epidendrum difforme Jacq., M Epidendrum diffusum Sw., M Epidendrum ecostatum Pabst ex Angley, K, S Epidendrum flexuosum G. Mey., I Epidendrum imitans Schltr., K Epidendrum longicolle Lindl., K Epidendrum macrocarpum (Rich.) Garay, comb. nov. ined., I, K, M Epidendrum microphyllum Lindl., M, S, D Epidendrum miserrimum Rchb.f., K Epidendrum nocturnum Jacq., I, K, M, S, D Epidendrum oldemanii Christenson, S Epidendrum orchidiflorum Salzm. ex Lindl., I, K Epidendrum purpurascens Focke, I, K, M, S Epidendrum ramosum Jacq., I, K, S Epidendrum rigidum Jacq., I, M, S, D Epidendrum schlechterianum Ames, M, D Epidendrum sculptum Rchb.f., M, D Epidendrum smaragdinum Lindl., K, D Epidendrum strobiliferum Rchb.f., I, K, M, S, D Epidendrum strobiloides Garay & Dunst., S Epidendrum tridens Poepp. & Endl., K Epidendrum tumuchumaciense (Veyret) Carnevali & Romero, K Epidendrum unguiculatum (C. Schweinf.) Garay & Dunst., S Epistephium amplexicaule (Ruiz. & Pav.) Poepp. & Endl., K Epistephium ellipticum R.O. Williams & Summerh., K Epistephium parviflorum Lindl., K Epistephium subrepens Hoehne, K Erythrodes stigmatoptera (Rchb.f.) Pabst, D Erythrodes sp., S Erythrodes sp., D Eulophia alta (L.) Fawc. & Rendle, K, M, D Gongora atropurpurea Hook., M Gongora pleiochroma Rchb.f., S Gongora sp., K Habenaria leprieurii Rchb.f., I, K Habenaria longicauda Hook., K Habenaria platydactyla Kraenzl., K Habenaria rodeiensis Barb. Rodr., S Habenaria sp., D Ionopsis satyrioides (Sw.) Rchb.f., S Ionopsis utricularioides (Sw.) Lindl., K Isochilus linearis (Jacq.) R. Br., S Jacquiniella globosa (Jacq.) Schltr., K, M, S Jacquiniella sp., D Kefersteinia lafontainei Senghas & G. Gerlach, S Kegeliella houtteana (Rchb.f.) L.O. Williams, S Koellensteinia carraoënsis Garay & Dunst., M, S Koellensteinia eburnea (Barb. Rodr.) Schltr., K Koellensteinia graminea (Lindl.) Rchb.f., M, S Koellensteinia tricolor (Lindl.) Rchb.f., K Lepanthes helicocephala Rchb.f., M, S Ligeophila peterana (Cogn.) Garay, S Ligeophila sp., K Lockhartia biserra (Rich.) Christenson & Garay, comb. nov. ined., S Lockhartia imbricata (Lam.) Hoehne, I, M, S Lycaste macrophylla (Poepp. & Endl.) Lindl., S Macroclinium mirabile (C. Schweinf.) Dodson, M Macroclinium wullschlaegelianum (Focke) Dodson, M Malaxis maguirei C. Schweinf., S Masdevallia cuprea Lindl., S Masdevallia minuta Lindl., I, M, S Masdevallia norae Luer, S Masdevallia sp., D Maxillaria acutifolia Lindl., S Maxillaria alba (Hook.) Lindl., M, S Maxillaria amazonica Schltr., D Maxillaria bolivarensis C. Schweinf., K Maxillaria brachybulbon Schltr., I, K, S Maxillaria brunnea Linden & Rchb.f., K Maxillaria caespitifica Rchb.f., K, M, S Maxillaria camaridii Rchb.f., I, K, M, S, D Maxillaria crassifolia (Lindl.) Rchb.f., I, S, D Maxillaria desvauxiana Rchb.f., M, S, D Maxillaria discolor (G. Lodd. ex Lindl.) Rchb.f., I, M, S, D Maxillaria huebneri Schltr., D Maxillaria imbricata Barb. Rodr., M Maxillaria mapiriensis (Kraenzl.) L.O. Williams, K Maxillaria melina Lindl., K Maxillaria parkeri Hook., I, M, D Maxillaria parviflora (Poepp. & Endl.) Garay, I, K, S Maxillaria pauciflora Barb. Rodr., D I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

56 50 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana Maxillaria pendens Pabst, D Maxillaria ponerantha Rchb.f., M, S Maxillaria porrecta Lindl., S Maxillaria pterocarpa Barb. Rodr., K Maxillaria ramosa Ruiz & Pav., K Maxillaria rufescens Lindl., M, S Maxillaria splendens Poepp. & Endl., M, S Maxillaria stenophylla Rchb.f., S Maxillaria superflua Rchb.f., M, S Maxillaria tenuis C. Schweinf., D Maxillaria trinitatis Ames, K, M Maxillaria uncata Lindl., I, K, M, S, D Maxillaria venezuelana (C. Schweinf.) Carnevali, I Maxillaria villosa (Barb. Rodr.) Cogn., K, S, D Maxillaria violaceopunctata Rchb.f., K, M, S Maxillaria xylobiiflora Schltr., D Mormodes buccinator Lindl., S Myoxanthus simplicicaulis (C. Schweinf.) Luer, K Myoxanthus trachychlamys (Schltr.) Luer, K Notylia peruviana (Schltr.) C. Schweinf., S Notylia sagittifera (Kunth) Link, Klotzsch, & Otto, M Notylia sp., D Octomeria amazonica Pabst, D Octomeria brevifolia Cogn., M, S, D Octomeria deltoglossa Garay, M Octomeria erosilabia C. Schweinf., K, D Octomeria exigua C. Schweinf., K Octomeria integrilabia C. Schweinf., K Octomeria minor C. Schweinf., K, M, S Octomeria sarthouae Luer, S Octomeria surinamensis Focke, M, S Octomeria taracuana Schltr., D Octomeria sp., I Oncidium lanceanum Lindl., M Oncidium orthostates Ridl., M Oncidium sp., S Orleansia amazonica Barb. Rodr., I, M Orleansia sp., D Ornithocephalus avicula Rchb.f. & Wullschl., D Ornithocephalus bicornis Lindl., M Ornithocephalus kruegeri Rchb.f., S Ornithocephalus sp., K, D Otostylis venusta (Ridl.) Schltr., K Palmorchis pabstii Veyret, S Palmorchis prospectorum Veyret, S Palmorchis pubescens Barb. Rodr., M Palmorchis sobralioides Barb.Rodr., D Palmorchis sp., K Palmorchis sp., I Paphinia grandiflora Barb.Rodr., D Paphinia sp., I Pelexia callifera (C. Schweinf.) Garay, S Phragmipedium lindleyanum (R.H. Schomb. ex Lindl.) Rolfe, K, S Platystele ovalifolia (H. Focke) Garay & Dunst., M, S, D Plectrophora cultrifolia (Barb. Rodr.) Cogn., S Plectrophora iridifolia H. Focke, M Pleurothallis archidiaconi Ames, S Pleurothallis aristata Hook., M, S Pleurothallis barbulata Lindl., M, S Pleurothallis biglandulosa Schltr., M Pleurothallis brevipes H. Focke, M, S Pleurothallis breviscapa C. Schweinf., M Pleurothallis ciliolata Schltr., M, S Pleurothallis consimilis Ames, S Pleurothallis determannii Luer, M Pleurothallis discoidea Lindl., S Pleurothallis ephemera Lindl., M Pleurothallis erythrogramma Luer & Carnevali, D Pleurothallis glandulosa Ames, M, S Pleurothallis grobyi Bateman ex Lindl., K Pleurothallis lanceana Lodd., I, S Pleurothallis mentosa Cogn., D Pleurothallis minima C. Schweinf., M Pleurothallis miqueliana (H. Focke) Lindl., M Pleurothallis monophylla (Hook.) Fawc. & Rendle, M Pleurothallis nanifolia Foldats, S Pleurothallis peperomioides Ames, D Pleurothallis picta Lindl., K, M, S, D Pleurothallis pluriflora (Barb. Rodr.) Cogn., D Pleurothallis polygonoides Griseb., M, S Pleurothallis pruinosa Lindl., M, S Pleurothallis pubescens Lindl., S Pleurothallis ruscifolia (Jacq.) R. Br., S Pleurothallis sclerophylla Lindl., K, S Pleurothallis semperflorens Lindl., M, S Pleurothallis seriata Lindl., M, S Pleurothallis spiculifera Lindl., M, S, D Pleurothallis stenocardium Schltr., K Pleurothallis suspensa Luer, K, S Pleurothallis uniflora Lindl., S Polycycnis surinamensis C. Schweinf., K Polycycnis vittata (Lindl.) Rchb.f., D Polystachya concreta (Jacq.) Garay & H.R. Sweet, S Polystachya foliosa (Hook.) Rchb.f., M, S Polystachya stenophylla Schltr., I, K, M, S, D Psychopsis papilio (Lindl.) H.G. Jones, S Psygmorchis glossomystax (Rchb.f.) Dodson & Dressler, S Psygmorchis pusilla (L.) Dodson & Dressler, K Psygmorchis sp., D Quekettia microscopica Lindl., I, M, S Quekettia vermeuleniana Determann, M Reichenbachanthus reflexus (Lindl.) Brade, I, K, M, S Rodriguezia candida (Lindl.) Christenson, M Rodriguezia lanceolata Ruiz & Pav., I, K, M, S, D Rudolfiella aurantiaca (Lindl.) Hoehne, I, M, D Sarcoglottis acaulis (Sm.) Schltr., K, S Sarcoglottis metallica (Rolfe) Schltr., K Scaphyglottis amazonica Schltr., D Scaphyglottis fusiformis (Griseb.) Schult., K, S Scaphyglottis graminifolia (Ruiz & Pav.) Poepp. & Endl., K Scaphyglottis grandiflora Ames & C. Schweinf., K Scaphyglottis modesta (Rchb.f.) Schltr., K, M, S Scaphyglottis ochroleuca Schltr., D Scaphyglottis prolifera Cogn., K, M, S Scaphyglottis sickii Pabst, I, K, M, D Scaphyglottis stellata Lodd. ex Lindl., K, M, S Scaphyglottis violacea Lindl., K, M Schomburgkia marginata Lindl., S Schomburgkia sp., I Scuticaria steelii Lindl., S I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

57 BRIT.ORG/SIDA 51 Selenipedium palmifolium (Lindl.) Rchb.f., M, S Sievekingia jenmanii Rchb.f., K Sigmatostalix amazonica Schltr., D Sobralia fragrans Lindl., S, D Sobralia liliastrum Lindl., K Sobralia macrophylla Rchb.f., D Sobralia oliva-estevae Carnevali & I. Ramírez, K Sobralia sessilis Lindl., S Sobralia suaveolens Rchb.f., S Sobralia valida Rolfe, K Sobralia yauaperyensis Barb. Rodr., S Stanhopea grandiflora (Lodd.) Lindl., S Stelis argentata Lindl., K, M, S Stelis endresii Rchb.f., D Stelis guianensis Rolfe, S Stelis intermedia Poepp. & Endl., K Stelis latisepala C. Schweinf., K Stelis santiagoensis Mansf., S Stelis sp., I Trichosalpinx ciliaris (Lindl.) Luer, S Trichosalpinx egleri (Pabst) Luer, K, S Trichosalpinx memor (Rchb.f.) Luer, M Trichosalpinx orbicularis (Lindl.) Luer, K, M, S, D Trigonidium acuminatum Bateman, K, M, S, D Trigonidium obtusum Lindl., I, K, M, D Triphora amazonica Schltr., D Triphora surinamensis (Lindl. ex Benth.) Britton, S Trisetella triglochin (Rchb.f.) Luer, S Trisetella sp., D Uleiorchis ulei (Cogn.) Handro, S, D Vanilla bicolor Lindl., K Vanilla cristato-callosa Hoehne, M Vanilla grandiflora Lindl., K, S Vanilla mexicana Mill., S Vanilla palmarum Lindl., D Vanilla planifolia Jacks. ex Andrews, S Vanilla sp., I Wullschlaegelia calcarata Benth., I, K, M, S, D Xerorchis trichorhiza (Kraenzl.) Garay, I Xerorchis sp., M Xylobium variegatum (Ruiz & Pav.) Garay & Dunst., S Oxalidaceae Oxalis barrelieri L., S, D Oxalis cytisoides C. Mart. ex Zucc., D Oxalis frutescens L., I, M Passifloraceae Dilkea johannesii Barb. Rodr., S, D Dilkea retusa Mast., D Dilkea wallisii Mast., M Passiflora acuminata DC., D Passiflora amoena L.K. Escobar, I, S Passiflora auriculata Kunth, I, K, M, D Passiflora candida (Poepp. & Endl.) Mast., S, D Passiflora capparidifolia Killip, I Passiflora cirrhiflora Juss., I, M Passiflora coccinea Aubl., I, M, S, D Passiflora edulis Sims, D Passiflora exura Feuillet, S Passiflora faroana Harms, D Passiflora foetida L., K, S Passiflora fuchsiiflora Hemsl., I, M, S Passiflora garckei Mast., K, M, S Passiflora glandulosa Cav., I, K, M, S Passiflora hexagonocarpa Barb.Rodr., D Passiflora laurifolia L., I, M, D Passiflora maguirei Killip, K Passiflora misera Kunth, M Passiflora nitida Kunth, K, D Passiflora riparia aff. Mart. ex Mast., D Passiflora rubra L., S Passiflora variolata Poepp. & Endl., S, D Passiflora vespertilio L., I, K, M, S Passiflora sp. nov., I Peridiscaceae Peridiscus lucidus Benth., D Phytolaccaceae Microtea debilis Sw., S Petiveria alliacea L., S Phytolacca rivinoides Kunth & Bouché, I, K, M, S Phytolacca sp., D Piperaceae Peperomia elongata Kunth, K, S Peperomia emarginella (Sw. ex Wikstr.) C. DC., S Peperomia glabella (Sw.) A. Dietr., M, S Peperomia gracieana Görts, S Peperomia macrostachya (Vahl) A. Dietr., I, K, M, S, D Peperomia magnoliifolia (Jacq.) A. Dietr., K Peperomia maguirei Yunck., M, S Peperomia obtusifolia (L.) A. Dietr., I, K, M, S Peperomia ouabianae C. DC., I, M, S Peperomia pellucida (L.) Kunth, K, M, S Peperomia pernambucensis Miq., S Peperomia quadrangularis (J.V. Thomps.) A. Dietr., I, M Peperomia rotundifolia (L.) Kunth, K, M, S Peperomia serpens (Sw.) Loudon, I, M, S Piper adenandrum (Miq.) C. DC., S Piper aduncum L., K, D Piper aequale Vahl, I, S Piper alatabaccum Trel. & Yunck., S, D Piper amalaqo L., S Piper amapense Yunck., K, D Piper amplectenticaule Trel. & Yunck., S Piper angustifolium Lam., S Piper anonifolium (Kunth) C. DC., I, S Piper arboreum Aubl., I, K, M, S, D Piper augustum Rudge, S Piper avellanum (Miq.) C. DC., I, S Piper baccans (Miq.) C. DC., D Piper bartlingianum (Miq.) C. DC., I, M, S, D Piper berbicense Miq., I, K Piper brachypetiolatum Yunck., D Piper brownsbergense Yunck., S Piper carniconnectivum C. DC., D Piper cernuum Vell., S Piper consanguineum (Kunth) Trel. & Yunck., S Piper crassinervium Kunth, S Piper cyrtopodon (Miq.) C. DC., S, D Piper dactylostigmum Yunck., D Piper demeraranum (Miq.) C. DC., I, M, S, D I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

58 52 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana Piper dilatatum Rich., I, S Piper divaricatum G. Mey., M, S Piper duckei C. DC., D Piper dumosum Rudge, S Piper durilignum C. DC., D Piper erectipilum Yunck., D Piper eucalyptifolium Rudge, S Piper froesii Yunck., D Piper guianense (Klotzsch) C. DC., D Piper gurupanum Yunck., D Piper hispidum Sw., I, M, S, D Piper hostmannianum (Miq.) C. DC., I, K, M, S, D Piper humaytanum Yunck., D Piper humillimum C. DC., D Piper humistratum Görts & K.U. Kramer, S Piper insipiens Trel. & Yunck., I Piper kegelianum (Miq.) C. DC., D Piper liesneri Steyerm., D Piper mastersianum C. DC., D Piper negroense C. DC., D Piper nematanthera (Miq.) C. DC., S Piper nigrispicum C. DC., M, S Piper obliquum Ruiz & Pav., I, S Piper paraense (Miq.) C. DC., D Piper paramaribense C. DC., S Piper piscatorum Trel. & Yunck., S Piper pleninervosum Trel., D Piper purusanum Yunck., D Piper renitens (Miq). Yuncker, D Piper reticulatum L., S Piper tectoniifolium Kunth, S Piper tortivenulosum Yunck., D Piper trichoneuron (Miq.) C. DC., I, M, S Pothomorphe peltata (L.) Miq., S Poaceae Andropogon bicornis L., K, M, S, D Andropogon leucostachyus Kunth, K, M, D Andropogon selloanus (Hack.) Hack., K, M Arthrostylidium scandens McClure, K Arundinella hispida (Humb. & Bonpl. ex Willd.) Kuntze, I Axonopus capillaris (Lam.) Chase, K, M Axonopus compressus (Sw.) P. Beauv., S Axonopus fissifolius non (Raddi) Kuhlm. sensu Amshoff & Henrard, K Axonopus flabelliformis Swallen, K Axonopus longispicus (Döll) Kuhlm., I, K Axonopus purpusii (Mez) Chase, K, S Cenchrus brownii Roem. & Schult., M Cenchrus echinatus L., M Cryptochloa capillata (Trin.) Soderstr., S Cynodon dactylon (L.) Pers., D Digitaria horizontalis Willd., I, K, D Digitaria lanuginosa (Nees) Henrard, M Digitaria violascens Link, K Eragrostis maypurensis (Kunth) Steud., K, M, S Guadua latifolia (Bonpl.) Kunth, S Guadua sp., I Gymnopogon foliosus (Willd.) Nees, M Gynerium sagittatum (Aubl.) P. Beauv., K, M Homolepis aturensis (Kunth) Chase, S, D Homolepis isocalycia (G. Mey.) Chase, K, M Ichnanthus breviscrobs Döll, S Ichnanthus calvescens (Nees) Döll, K Ichnanthus dasycoleus Tutin, K Ichnanthus leiocarpus (Spreng.) Kunth, I, S Ichnanthus nemoralis (Schrad.) Hitchc. & Chase, I, K, M, S Ichnanthus pallens (Sw.) Munro ex Benth., I, K, M, S Ichnanthus panicoides P. Beauv., I, S, D Ichnanthus tenuis (J. Presl) Hitchc. & Chase, I, K Ischaemum guianense Kunth ex Hack., K Ischaemum rugosum Salisb., K Lasiacis ligulata Hitchc. & Chase, I, K, M, S Lasiacis procerrima (Hack.) Hitchc., K Lasiacis sorghoidea (Desv.) Hitchc. & Chase, S Mesosetum loliiforme (Hochst. ex Steud.) Chase, K Olyra caudata Trin., S Olyra ciliatifolia Raddi, I Olyra ecaudata Doell, S Olyra latifolia L., I, S, D Olyra longifolia Kunth, I, K Olyra micrantha Kunth, I, K, M, D Olyra ramosissima Doell, D Oplismenus hirtellus (L.) P. Beauv., S Orthoclada laxa (Rich.) P. Beauv., I, M, S Panicum chnoodes Trin., K Panicum cyanescens Nees, I, K Panicum granuliferum Kunth, K Panicum hylaeicum Mez, I Panicum laxum Sw., I, K, S, D Panicum mertensii Roth, D Panicum micranthum Kunth, I, K Panicum parvifolium Lam., I, K Panicum pilosum Sw., I, K, M, S, D Panicum polycomum Trin., I, K Panicum pycnoclados Tutin, K Panicum rivale Swallen, I, K, S Panicum rudgei Roem. & Schult., I, K Panicum sciurotoides Zuloaga & Morrone, K, S Panicum stoloniferum Poir., I, K, M, S Pariana campestris Aubl., S, D Pariana radiciflora Sagot ex Doell, I, K, S, D Parodiolyra lateralis (J. Presl ex Nees) Soderstr. & Zuloaga, K Paspalum altsonii Chase, K Paspalum arenarium Schrad., K Paspalum bifidifolium Soderstr., K Paspalum carinatum Humb. & Bonpl. ex Flüggé, I Paspalum conjugatum Bergius, I, K, S, D Paspalum convexum Humb. & Bonpl. ex Flüggé, I Paspalum decumbens Sw., M Paspalum melanospermum Desv. ex Poir., I, K, M Paspalum paniculatum L., S Paspalum petilum Chase, K Paspalum pilosum Lam., I Paspalum plicatulum Michx., I Paspalum virgatum L., I, S Pharus lappulaceus Aubl., S Pharus latifolius L., S Pharus parvifolius Nash, S Pharus virescens Doell, S Piresia goeldii Swallen, S I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

59 BRIT.ORG/SIDA 53 Piresia macrophylla Soderstr., S Raddiella esenbeckii (Steud.) C.E. Calderón & Soderstr., I, K Raddiella kaieteurana Soderstr., K Raddiella potaroensis Soderstr., K Sporobolus jacquemontii Kunth, K, S, D Streptogyna americana C.E. Hubb., S Thrasya achlysophila Soderstr., K Thrasya scandens (Tutin) Soderstr. ex A.G. Burm., I, K Tripsacum andersonii J.R. Gray, S Podostemaceae Apinagia kochii (Engl.) P. Royen, S Apinagia richardiana (Tul.) P. Royen, S Jenmaniella ceratophylla Engl., K Jenmaniella isoetifolia P. Royen, K Marathrum sp., I, K Mourera fluviatilis Aubl., I Rhyncholacis apiculata P. Royen, K Rhyncholacis jenmanii Engl., K Polygalaceae Barnhartia floribunda Gleason, S Bredemeyera densiflora A.W. Benn., K, M Bredemeyera myrtifolia Spruce ex. A.W.Benn., D Moutabea guianensis Aubl., I, M, S, D Polygala adenophora DC., I, K Polygala appressa Benth., K Polygala spectabilis DC., S, D Polygala subtilis Kunth, I Polygala timoutou Aubl., I Securidaca diversifolia (L.) S.F. Blake, M Securidaca longifolia Poepp., K, D Securidaca marginata Benth., I Securidaca paniculata Rich., I, M Securidaca pubescens DC., S Securidaca pyramidalis Sprague ex Sandwith, K Securidaca retusa Benth., K Securidaca rivinaefolia A. St.-Hil., D Securidaca spinifex Sandwith, M, S Securidaca uniflora Oort, K, S, D Securidaca volubilis L., S, D Polygonaceae Coccoloba ascendens Duss ex Lindau, I, S Coccoloba confusa Howard, D Coccoloba densifrons Mart. ex Meisn., I Coccoloba excelsa Benth., S Coccoloba gymnorrhachis Sandwith, I, M Coccoloba lucidula Benth., M Coccoloba marginata Benth., K, M Coccoloba ovata Benth., I Coccoloba padiformis Meisn., I Coccoloba parimensis Benth., M, D Coccoloba striata Benth., M Polygonum acuminatum Kunth, I Symmeria paniculata Benth., I, M Polypodiaceae Campyloneurum fuscosquamatum Lellinger, S Campyloneurum phyllitidis (L.) C. Presl, I, S Campyloneurum repens (Aubl.) C. Presl, I, S, D Campyloneurum sphenodes (Kunze) Fée, S Dicranoglossum desvauxii (Klotzsch) Proctor, I, M, S, D Dicranoglossum furcatum (L.) J. Sm., S Microgramma baldwinii Brade, D Microgramma fuscopunctata (Hook.) Vareschi, I, K, S Microgramma loretensis? (Maxon), D Microgramma lycopodioides (L.) Copel., I, K, M, S Microgramma persicariifolia (Schrad.) C. Presl, I, S Microgramma reptans (Cav.) A.R. Sm., I, K, M, S Microgramma tecta (Kaulf.) Alston, I, S Microgramma thurnii (Baker) R.M. Tryon & Stolze, I, S, D Micropolypodium nanum (Fée) A.R. Sm., D Niphidium crassifolium (L.) Lellinger, I, K, S, D Pecluma consimilis (Mett.) M.G. Price, I Pecluma pectinata (L.) M.G. Price, S Pecluma plumula (Humb. & Bonpl. ex Willd.) M.G. Price, I, S Phlebodium aureum (L.) J. Sm., K Phlebodium decumanum (Willd.) J. Sm., S Phlebodium pseudoaureum (Cav.) Lellinger, K Pleopeltis percussa (Cav.) Hook. & Grev., I, M, S Polypodium adnatum Kunze ex Klotzsch, S Polypodium caceresii Sodiro, K, S Polypodium dulce Poir., I, S Polypodium lasiopus Klotzsch, S Polypodium loriceum L., K Polypodium panorense C. Chr., K Polypodium polypodioides (L.) Watt, I, S Polypodium triseriale Sw., K, M, S, D Pontederiaceae Eichhornia diversifolia (Vahl) Urb., I, K Portulacaceae Talinum fruticosum (L.) A. Juss., M Proteaceae Euplassa pinnata (Lam.) Johnst., S Panopsis rubescens (Pohl) Pittier, S Panopsis sessilifolia (Rich.) Sandwith, K, M, D Roupala montana Aubl., K, S Roupala schomburgkii Klotzsch, I Pteridaceae Polytaenium guayanense (Hieron.) Alston, D Pteris altissima Poir., S Pteris deflexa Link, S Pteris podophylla Sw., D Pteris pungens Willd., I, S Quiinaceae Lacunaria crenata (Tul.) A.C. Sm., I, S, D Lacunaria jenmanii (Oliv.) Ducke, S, D Lacunaria macrostachya (Tul.) A.C. Sm., D Quiina amazonica A.C. Sm., D Quiina guianensis Aubl., I, M, S Quiina indigofera Sandwith, M Quiina negrensis A.C. Sm., D Quiina obovata Tul., I, M, S, D Quiina oiapocensis Pires, K, S Quiina pteridophylla (Radlk.) Pires, D Quiina rhytidopus Tul., I Quiina sessilis Choisy, S Touroulia guianensis Aubl., M, S, D Rafflesiaceae Apodanthes caseariae Poit., S Apodanthes sp., D I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

60 54 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana Rapateaceae Potarophytum riparium Sandwith, K Rapatea fanshawei Maguire, K Rapatea paludosa Aubl., I, K, M, D Rapatea saulensis B.M. Boom, S Rapatea ulei Pilg., D Rapatea xiphoides Sandwith, K Saxofridericia regalis R.H. Schomb., K Saxofridericia subcordata Korn., D Spathanthus unilateralis (Rudge) Desv., I, K, S, D Stegolepis angustata Gleason, K Stegolepis ferruginea Baker, K Windsorina guianensis Gleason, K Rhabdodendraceae Rhabdodendron amazonicum (Spruce ex Benth.) Huber, I, M, S, D Rhabdodendron macrophyllum (Spruce ex Benth.) Huber, D Rhamnaceae Ampelozizyphus amazonicus Ducke, D Gouania blanchetiana Miq., I, M, S Gouania polygama (Jacq.) Urb., I Gouania velutina Reissek, I Ziziphus cinnamomum Triana & Planch., S Rhizophoraceae Cassipourea elliptica (Sw.) Poir., S Cassipourea guianensis Aubl., I, K, M, S, D Cassipourea lasiocalyx Alston, I, M Sterigmapetalum obovatum Kuhlm., D Rosaceae Prunus myrtifolia (L.) Urb., M, S, D Rubiaceae Alibertia edulis (Rich.) A. Rich. ex DC., D Alibertia hispida Ducke, D Alibertia myrciifolia K. Schum., I, S, D Alibertia surinamensis (Bremek.) Steyerm., S Alseis longifolia Ducke, S Amaioua brevidentata Steyerm., I Amaioua corymbosa Kunth, D Amaioua guianensis Aubl., I, M, S, D Bertiera guianensis Aubl., S Borojoa claviflora (K. Schum.) Cuatr., D Borreria alata (Aubl.) DC., S Borreria capitata (Ruiz & Pav.) DC., I, M Borreria latifolia (Aubl.) K. Schum., I, K, S Borreria prostrata (Aubl.) Miq., I Borreria verticillata (L.) G. Mey., I, K, S, D Bothriospora corymbosa (Benth.) Hook.f., S Botryarrhena pendula Ducke, D Capirona decorticans Spruce, S, D Chalepophyllum guianense Hook.f., K Chimarrhis barbata (Ducke) Bremek., D Chimarrhis duckeana Delprete, D Chimarrhis microcarpa Standl., I, S Chimarrhis turbinata DC., S Chiococca alba (L.) Hitchc., S Chiococca nitida Benth., S Chomelia estrellana Müll. Arg., D Chomelia malaneoides Müll. Arg., D Chomelia monachinoi Steyerm., I Chomelia tenuiflora Benth., I, S, D Coccocypselum aureum (Spreng.) Cham. & Schltdl., K Coccocypselum guianense (Aubl.) K. Schum., I, K, M, S Coccocypselum herbaceum P. Browne, I Coccocypselum hirsutum Bartl. ex DC., K, M Coccocypselum tontanea Kunth, I Cosmibuena grandiflora (Ruiz & Pav.) Rusby, K, M, D Cosmibuena triflora (Benth.) Klotzsch, D Coussarea amapaensis Steyerm., S Coussarea ampla Müll. Arg., D Coussarea fanshawei Steyerm., K Coussarea grandis Müll. Arg., D Coussarea granvillei Delprete & B. Boom, S Coussarea machadoana Standl., S Coussarea paniculata (Vahl) Standl., I, K, S Coussarea racemosa A. Rich., S Coussarea revoluta Steyerm., D Coussarea violacea Aubl., S Coutarea hexandra (Jacq.) K. Schum., S Didymochlamys connellii N.E. Br., K Diodia hyssopifolia (Willd. ex Roem. & Schult.) Cham. & Schltdl., I Diodia ocymifolia (Willd. ex Roem. & Schult.) Bremek., I, K, M, S Diodia spicata Miq., S Duroia amapana Steyerm., M Duroia aquatica (Aubl.) Bremek., S Duroia eriopila L.f., I, K, M, S Duroia fusifera (Spruce ex Benth. & Hook. f.) Hook. f. ex K. Schum., I Duroia genipoides (Spruce) Hook. f. ex K. Schum., I Duroia gransabanensis Steyerm., D Duroia longiflora Ducke, S, D Duroia macrophylla Huber, D Duroia micrantha (Ladbr.) Zarucchi & J.H. Kirkbr., I Duroia saccifera (Mart.) Hook.f. ex K. Schum., D Faramea capillipes Müll. Arg., D Faramea corymbosa Aubl., S, D Faramea egregia Sandwith, K Faramea guianensis (Aubl.) Bremek., S Faramea lourteigiana Steyerm., S Faramea multiflora A. Rich. ex DC., S Faramea occidentalis (L.) A. Rich., I, M Faramea platyneura Müll. Arg., D Faramea quadricostata Bremek., I, K, M, S Faramea sessilifolia (Kunth) DC., I, M Faramea spathacea Müll. Arg. ex Standl., K Faramea tinguana Müll. Arg., S Faramea torquata Müll. Arg., D Ferdinandusa elliptica Pohl, D Ferdinandusa goudotiana K. Schum., M Ferdinandusa guainiae Spruce ex K. Schum., K Ferdinandusa hirsuta Standl., D Ferdinandusa paraensis Ducke, S Ferdinandusa uaupensis Spruce ex K. Schum., I Genipa americana L., I, K, M Genipa spruceana Steyerm., I, K, S Geophila cordifolia Miq., K, M, S, D Geophila repens (L.) I.M. Johnst., I, K, S, D Geophila tenuis (Müll. Arg.) Standl., S Gonzalagunia dicocca Cham. & Schltdl., I, M, S I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

61 BRIT.ORG/SIDA 55 Guettarda acreana K. Krause, I, S Guettarda argentea Lam., I Guettarda spruceana Müll. Arg., I, S Hamelia axillaris Sw., S Henriquezia verticillata Spruce ex Benth., D Hillia illustris (Vell.) K. Schum., I, M, S Hillia parasitica Jacq., K, S Hillia ulei K. Krause, D Isertia coccinea (Aubl.) J.F. Gmel., S Isertia hypoleuca Benth., I, K, M, D Isertia parviflora Vahl, I Isertia rosea Spruce ex K. Schum., D Isertia spiciformis DC., S Ixora aluminicola Steyerm., M, S Ixora davisii Sandwith, M Ixora ferrea (Jacq.) Benth., I Ixora floribunda (A. Rich.) Griseb., M Ixora graciliflora Benth., I, M, S Ixora intensa K. Krause, D Ixora intropilosa Steyerm., M Ixora panurensis Müll. Arg., I, D Ixora piresii Steyerm., S Ixora pubescens Willd. ex Schult.. & Schult. f., S Ixora schomburgkiana Benth., K Ixora ulei K. Krause, K Kotchubaea insignis Fisch. ex DC., D Kotchubaea semisericea Ducke, D Kotchubaea sericantha Standl., D Ladenbergia amazonenis Ducke, D Malanea hypoleuca Steyerm., K, M, D Malanea macrophylla Bartl. ex Griseb., I, K, M, S, D Malanea obovata Hochr., K Malanea sarmentosa Aubl., M Manettia alba (Aubl.) Wernham, K, S Manettia coccinea (Aubl.) Willd., S Manettia reclinata L., S Mitracarpus hirtus (L.) DC., I Morinda brachycalyx (Bremek.) Steyerm., M, S Morinda calycina (Benth.) Steyerm., I Morinda debilis (Sandwith) Steyerm., M Morinda tenuiflora (Benth.) Steyerm., I, M Morinda triphylla (Ducke) Steyerm., D Oldenlandia corymbosa L., M Oldenlandia lancifolia (Schumach.) DC., S Pagamea capitata Benth., I, K, M Pagamea guianensis Aubl., K, M Pagamea macrophylla Spruce ex Benth., D Pagamea thyrsiflora Spruce ex Benth., K Palicourea amapaensis Steyerm., D Palicourea anisoloba (Müll. Arg.) B.M. Boom & M.T. Campos, D Palicourea brachyloba (Müll. Arg.) B. Boom, S Palicourea calophylla DC., S Palicourea crocea (Sw.) Roem. & Schult., K, M, S Palicourea croceoides Ham., I, K, M, S Palicourea guianensis Aubl., I, K, M, S, D Palicourea longiflora (Aubl.) A. Rich., S, D Palicourea longistipulata (Müll. Arg.) Standl., S, D Palicourea nitidella (Müll. Arg.) Standl., D Palicourea quadrifolia (Rudge) DC., S Palicourea triphylla DC., I, K Palicourea virens (Poepp. & Endl.) Standl., D Patima guianensis Aubl., S Perama dichotoma Poepp. & Endl., K Perama galioides (Kunth) Poir., I, K Perama hirsuta Aubl., I, K Posoqueria coriacea M. Martens & Galeotti, K Posoqueria latifolia (Rudge) Roem. & Schult., K, M, S Posoqueria longiflora Aubl., I, K Posoqueria panamensis (Walp. & Duchass.) Walp., I, K Psychotria acuminata Benth., K, S Psychotria alloantha Steyerm., S Psychotria anceps Kunth, K, M Psychotria aneurophylla (Standl.) Steyerm., K Psychotria apoda Steyerm., I, M, S, D Psychotria appuniana Steyerm., K Psychotria astrellantha Wernham, I, M, S, D Psychotria bahiensis DC., I, M, S, D Psychotria barbiflora DC., I, K Psychotria blepharophylla (Standl.) Steyerm., S Psychotria borjensis Kunth, S Psychotria bostrychothyrsus Sandwith, K Psychotria brachybotrya Müll. Arg., I, S Psychotria bracteocardia (DC.) Müll. Arg., I, K Psychotria bremekampiana Steyerm., S Psychotria callithrix (Miq.) Steyerm., S Psychotria capitata Ruiz & Pav., I, K, M, S Psychotria carthagenensis Jacq., K, S Psychotria chlorantha Benth., K Psychotria cincta Standl., D Psychotria colorata (Willd. ex Roem. & Schult.) Müll. Arg., S Psychotria crocochlamys Sandwith, K Psychotria cupularis (Müll. Arg.) Standl., I, S Psychotria deflexa DC., M, S Psychotria deinocalyx Sandwith, D Psychotria erecta (Aubl.) Standl. & Steyerm., S, D Psychotria fanshawei (Standl.) Steyerm., K Psychotria gracilenta Müll. Arg., D Psychotria granvillei Steyerm., S Psychotria guadalupensis (DC.) R.A. Howard, K, S Psychotria hemicephaelis Wernham, K Psychotria hoffmannseggiana (Willd. ex Roem. & Schult.) Müll. Arg., K, M Psychotria horizontalis Sw., I Psychotria humboldtiana (Cham.) Müll. Arg., D Psychotria iodotricha Müll. Arg., K, S Psychotria irwinii Steyerm., M Psychotria kaieteurensis Sandwith, K Psychotria kappleri (Miq.) Müll. Arg. ex Benoist, S Psychotria lateralis Steyerm., S Psychotria ligularis (Rudge) Steyerm., S Psychotria lupulina Benth., I, M Psychotria macrophylla Ruiz & Pav., S Psychotria manausensis Steyerm., D Psychotria mapourioides DC., I, K, M, S, D Psychotria maturacensis Steyerm., K Psychotria mazaruniensis Standl., K Psychotria medusula Müll. Arg., D Psychotria microbotrys Ruiz ex Standl., I, S Psychotria microbracteata Steyerm., S Psychotria microdon (DC.) Urb., I I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

62 56 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana Psychotria moroidea Steyerm., S Psychotria muscosa (Jacq.) Steyerm., K Psychotria oblonga (DC.) Steyerm., S Psychotria officinalis (Aubl.) Raeusch., I, M, S Psychotria pacimonica Müll. Arg., D Psychotria perferruginea Steyerm., S Psychotria platypoda DC., K, S, D Psychotria podocephala (Müll. Arg.) Standl., D Psychotria poeppigiana Müll. Arg., I, K, S, D Psychotria polycephala Benth., I, D Psychotria potaroensis (Sandwith) Steyerm., K Psychotria prancei Steyerm., D Psychotria psittacina Steyerm., K Psychotria puberulenta Steyerm., M Psychotria pullei Bremek., S Psychotria pungens Steyerm., S Psychotria racemosa (Aubl.) Raeusch., I, S Psychotria ramiflora Rusby, K Psychotria rhombibractea C.M. Taylor & M.T. Campos ined, D Psychotria rosea (Benth.) Müll. Arg., K, M Psychotria saulensis Steyerm., S Psychotria sciaphila S.Moore, D Psychotria sphaerocephala Müll. Arg., D Psychotria subundulata Benth., K, D Psychotria tapajozensis Standl., K Psychotria transiens Wernham, K Psychotria trichophoroides Müll. Arg., S Psychotria uliginosa Sw., I, K, M, S Psychotria ulviformis Steyerm., S Psychotria urceolata Steyerm., S Psychotria variegata Steyerm., S Psychotria viridibracteata Steyerm., S Randia armata (Sw.) DC., I Randia asperifolia (Sandwith) Sandwith, M, S Randia calycina Cham., I Randia nitida (Kunth) DC., S Remijia amazonica K. Schum., D Remijia ulei K. Krause, D Remijia sp., I Retiniphyllum schomburgkii (Benth.) Müll. Arg., I, K Rudgea ayangannensis Steyerm., K Rudgea bremekampiana Steyerm., S Rudgea cornifolia (Humb. & Bonpl. ex Roem. & Schult.) Standl., I, M Rudgea cornigera Bremek., M Rudgea crassiloba (Benth.) B.L. Rob., I Rudgea fissistipula Müll. Arg., I, D Rudgea graciliflora Standl., D Rudgea guyanensis (A. Rich.) Sandwith, S Rudgea hostmanniana Benth., I, M Rudgea lanceifolia Salisb., S Rudgea sandemannii Sandwith, S Rudgea standleyana Steyerm., S Rudgea stipulacea (DC.) Steyerm., I, S Sabicea amazonensis Wernham, D Sabicea aspera Aubl., M, S Sabicea glabrescens Benth., K, M Sabicea oblongifolia (Miq.) Steyerm., I Sabicea surinamensis Bremek., M Sabicea villosa Willd. ex Roem. & Schult., I, S Schradera polycephala DC., D Sipanea biflora (L.f.) Cham. & Schltdl., M Sipanea cowanii Steyerm., K Sipanea pratensis Aubl., I, K, M, S, D Sipanea wilson-brownei R.S. Cowan, I, S Spermacoce assurgens Ruiz & Pav., I Spermacoce capitata Ruiz & Pav., M, S, D Spermacoce exilis (L.O. Williams) C.D. Adams, D Spermacoce ocymoides Burm. f., S Sphinctanthus maculatus Spruce ex K. Schum., S Tocoyena longiflora Aubl., S, D Uncaria guianensis (Aubl.) J.F. Gmel., I, M, S Uncaria tomentosa (Willd. ex Roem. & Schult.) DC., S Warszewiczia schwackei K. Schum., D Rutaceae Adiscanthus fusciflorus Ducke, D Conchocarpus fanshawei (Sandwith) Kallunki & Pirani, I Conchocarpus guyanensis (Pulle) Kallunki & Pirani, S Conchocarpus toxicarius (Engl.) Kallunki & Pirani, M Ertela trifolia (L.) Kuntze, I, D Erythrochiton brasiliense Nees & Mart., S Esenbeckia cowanii Kaastra, S Hortia longifolia Benth. ex Engl., D Hortia regia Sandwith, M Hortia superba Ducke, D Nycticalanthus speciosus Ducke, D Raputia sp., D Ticorea foetida Aubl., I, S Ticorea pedicellata DC., M Zanthoxylum apiculatum (Sandwith) P.G. Waterman, M Zanthoxylum compactum (Huber ex Albuq.) P.G. Waterman, I Zanthoxylum djalma-batistae (Albuq.) P.G. Waterman, D Zanthoxylum ekmanii (Urb.) Alain, S Zanthoxylum huberi P.G. Waterman, D Zanthoxylum pentandrum (Aubl.) R.A. Howard, S Zanthoxylum rhoifolium Lam., S, D Sabiaceae Meliosma palustre Kuhlm., D Salviniaceae Salvinia auriculata Aubl., I Sapindaceae Allophylus angustatus (Triana & Planch.) Radlk., S Allophylus latifolius Huber, S, D Allophylus leucoclados Radlk., S Allophylus racemosus Sw., I Allophylus robustus Radlk., S Cupania hirsuta Radlk., I, K, M Cupania hispida Radlk., D Cupania rubiginosa (Poir.) Radlk., K, D Cupania scrobiculata Rich., I, M, S, D Matayba arborescens (Aubl.) Radlk., M, S, D Matayba camptoneura Radlk., M Matayba elegans Radlk., I Matayba grandis Radlk., D Matayba guianensis Aubl., D Matayba inelegans Spruce ex Radlk., M, D Matayba laevigata Radlk., M I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

63 BRIT.ORG/SIDA 57 Matayba macrolepis Radlk., I Matayba macrostylis Radlk., M, D Matayba oligandra Sandwith, K Matayba opaca Radlk., M, D Matayba peruviana Radlk., K, S, D Matayba purgans (Poepp.) Radlk., I, S, D Paullinia alata (Ruiz & Pav.) G. Don, S Paullinia capreolata (Aubl.) Radlk., M, S, D Paullinia castaneifolia Radlk., D Paullinia clathrata Radlk., S, D Paullinia cupana Kunth, D Paullinia fibulata Rich. ex Juss., S Paullinia grandifolia Benth. ex Radlk., M, D Paullinia ingifolia Rich. ex Juss., I, M Paullinia laeta Radlk., D Paullinia latifolia Benth. ex Radlk., I, S Paullinia leiocarpa Griseb., I Paullinia lingulata Acev.-Rodr., S Paullinia livescens Radlk., M Paullinia pachycarpa Benth., I, M, S Paullinia perlata Radlk., I Paullinia pinnata L., M Paullinia plagioptera Radlk., I, S Paullinia rubiginosa Cambess., S Paullinia rufescens Rich. ex Juss., I, M Paullinia rugosa Benth. ex Radlk., M, D Paullinia stellata Radlk., I, S Paullinia stenopetala Sagot, S Paullinia stipularis Benth. ex Radlk., D Paullinia tricornis Radlk., S Paullinia uloptera Radlk., D Paullinia vespertilio Sw., K Paullinia xestophylla Radlk., D Porocystis toulicioides Radlk., D Pseudima frutescens (Aubl.) Radlk., I, S Serjania caracasana (Jacq.) Willd., I Serjania circumvallata Radlk., D Serjania didymadenia Radlk., S Serjania exarata Radlk., K Serjania grandifolia Sagot ex Radlk., S Serjania membranacea Splitg., D Serjania oblongifolia Radlk., I, S Serjania paniculata Kunth, I Serjania paucidentata DC., I, M, S, D Serjania pyramidata Radlk., S Serjania setulosa Radlk., S Talisia allenii Croat, D Talisia carinata Radlk., I, S Talisia cerasina (Benth.) Radlk., D Talisia clathrata Radlk., S Talisia cupularis Radlk., D Talisia elephantipes Sandwith, M Talisia eximia K.U. Kramer, D Talisia firma Radlk., D Talisia furfuracea Sandwith, M Talisia guianensis Aubl., S, D Talisia hemidasya Radlk., S Talisia longifolia (Benth.) Radlk., S, D Talisia macrophylla (Mart.) Radlk., S, D Talisia megaphylla Sagot ex Radlk., I, S Talisia micrantha Radlk., S Talisia microphylla Uittien, M Talisia mollis Kunth ex Cambess., I, S, D Talisia pedicellaris Radlk., S Talisia praealta Radlk., S, D Talisia reticulata Radlk., S Talisia simaboides K.U. Kramer, S Talisia squarrosa Radlk., M Talisia veraluciana Guarim, D Thinouia myriantha Triana & Planch., S Toulicia elliptica Radlk., M Toulicia guianensis Aubl., I, M, D Toulicia pulvinata Radlk., I, M, D Urvillea ulmacea Kunth, I Vouarana guianensis Aubl., S, D Sapotaceae Chromolucuma rubriflora Ducke, D Chrysophyllum amazonicum T.D. Penn., D Chrysophyllum argenteum Jacq., S, D Chrysophyllum colombianum (Aubrév.) T.D. Penn., D Chrysophyllum cuneifolium (Rudge) A. DC., S Chrysophyllum lucentifolium Cronquist, M, S, D Chrysophyllum manaosense (Aubrév.) T.D. Penn., D Chrysophyllum pomiferum (Eyma) T.D. Penn., M, S, D Chrysophyllum prieurii A. DC., S, D Chrysophyllum sanguinolentum (Pierre) Baehni, M, S, D Chrysophyllum sparsiflorum Klotzsch ex Miq., I, M, D Chrysophyllum ucuquirana-branca (Aubrév. & Pellegr.) T.D. Penn., D Chrysophyllum venezuelanense (Pierre) T.D. Penn., S Diploön cuspidatum (Hoehne) Cronquist, S Ecclinusa guianensis Eyma, I, M, S, D Ecclinusa lanceolata (Mart. & Eichler) Pierre, S, D Ecclinusa psilophylla Sandwith, M Ecclinusa ramiflora Mart., S Elaeoluma glabrescens (Mart. & Eichler) Aubrév., D Elaeoluma nuda (Baehni) Aubrév., D Elaeoluma schomburgkiana (Miq.) Baill., K Manilkara bidentata (A. DC.) A. Chev., I, K, M, S, D Manilkara cavalcantei Pires & W.A.Rodrigues, D Manilkara huberi (Ducke) A. Chev., S Manilkara paraensis (Huber) Standl., S Micropholis acutangula (Ducke) Eyma, S, D Micropholis brochidodroma T.D.Penn., D Micropholis casiquiarensis Aubrév., D Micropholis cayennensis T.D. Penn., S Micropholis cylindrocarpa (Poepp.) Pierre, D Micropholis egensis (A. DC.) Pierre, D Micropholis guyanensis (A. DC.) Pierre, S, D Micropholis longipedicellata Aubrév., S Micropholis melinoniana Pierre, K, S Micropholis mensalis (Baehni) Aubrév., D Micropholis obscura T.D. Penn., S Micropholis porphyrocarpa (Baehni) Monach., S, D Micropholis splendens Gilly ex Aubrév., D Micropholis submarginalis Pires & T.D. Penn., S Micropholis trunciflora Ducke, D Micropholis venulosa (Mart. & Eichler) Pierre, I, K, M, S, D Micropholis williamii Aubrév. & Pellegrin, D Pouteria ambelaniifolia (Sandwith) T.D. Penn., M, S, D Pouteria anomala (Pires) T.D.Penn., D I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

64 58 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana Pouteria bilocularis (H. Winkl.) Baehni, S, D Pouteria caimito (Ruiz & Pav.) Radlk., I, M, S, D Pouteria campanulata Baehni, D Pouteria cayennensis (A. DC.) Eyma, S Pouteria cladantha Sandwith, M, D Pouteria congestifolia Pilz, D Pouteria coriacea (Pierre) Pierre, M, S Pouteria cuspidata (A. DC.) Baehni, I, K, M, S, D Pouteria decorticans T.D. Penn., S Pouteria deliciosa T.D. Penn., S Pouteria durlandii (Standl.) Baehni, M Pouteria egregia Sandwith, M, S Pouteria elegans (A. DC.) Baehni, D Pouteria engleri Eyma, M, S, D Pouteria erythrochrysa T.D.Penn., D Pouteria eugeniifolia (Pierre) Baehni, K, S, D Pouteria favensis Pires & T.D.Penn., D Pouteria filipes Eyma, M, S, D Pouteria fimbriata Baehni, M, D Pouteria flavilatex T.D.Penn., D Pouteria franciscana Baehni, D Pouteria freitasii T.D.Penn., D Pouteria gardneri (Mart. & Miq.) Baehni, D Pouteria glomerata (Miq.) Radlk., D Pouteria gongrijpii Eyma, S, D Pouteria guianensis Aubl., K, M, S, D Pouteria hispida Eyma, I, M, S, D Pouteria jariensis Pires & T.D. Penn., S Pouteria kaieteurensis T.D. Penn., K Pouteria laevigata (Mart.) Radlk., S, D Pouteria latianthera T.D. Penn., D Pouteria macrophylla (Lam.) Eyma, D Pouteria manaosensis (Aubrév. & Pellegr.) T.D. Penn., D Pouteria maxima T.D. Penn., S, D Pouteria melanopoda Eyma, S Pouteria minima T.D.Penn., D Pouteria multiflora (A. DC.) Eyma, D Pouteria oblanceolata Pires, I, S, D Pouteria opposita (Ducke) T.D. Penn., D Pouteria peruviensis (Aubrév.) Bernardi, D Pouteria petiolata T.D.Penn., D Pouteria platyphylla (A.C. Sm.) Baehni, D Pouteria putamen-ovi T.D. Penn., S Pouteria reticulata (Engl.) Eyma, M, S, D Pouteria retinervis T.D. Penn., S, D Pouteria rigidopsis (Monach.) T.D. Penn., I Pouteria rodriguesiana Pires & T.D. Penn., S Pouteria rostrata (Huber) Baehni, D Pouteria singularis T.D. Penn., S Pouteria speciosa (Ducke) Baehni, I, M, S Pouteria stipulifera T.D.Penn., D Pouteria surumuensis Baehni, I, M, D Pouteria tarumanensis Pires, D Pouteria torta (Mart.) Radlk., I, M, S, D Pouteria venosa (Mart.) Baehni, I, M, S, D Pouteria vernicosa T.D.Penn., D Pouteria virescens Baehni, S, D Pouteria williamii (Aubrév. & Pellegr.) T.D. Penn., D Pouteria sp. nov. aff. simulans, D Pradosia beardii (Monach.) T.D. Penn., K Pradosia cochlearia (Lecomte) T.D. Penn., D Pradosia decipiens Ducke, D Pradosia ptychandra (Eyma) T.D. Penn., I, M, S Pradosia schomburgkiana (A. DC.) Cronquist, I, M, D Pradosia surinamensis (Eyma) T.D. Penn., I, M Pradosia verticillata Ducke, S, D Sarcaulus brasiliensis (A. DC.) Eyma, S, D Schizaeaceae Actinostachys pennula (Sw.) Hook., K, M, D Anemia sp., S Schizaea elegans (Vahl) Sm.., K, S, D Schizaea fluminensis Miers ex J.W. Sturm, I, K, M, S, D Schizaea incurvata Schkuhr, K, M, D Scrophulariaceae Achetaria guianensis Pennell, K Angelonia sp., S Bacopa aquatica Aubl., I, M Benjaminia reflexa (Benth.) D Arcy, I Buchnera rosea Kunth, I Capraria biflora L., S Conobea aquatica Aubl., I, K Lindernia crustacea (L.) F. Muell., M, S, D Lindernia diffusa (L.) Wettst., M, S, D Scoparia dulcis L., K, M, S, D Stemodia pratensis (Aubl.) B. L. Turner, I Selaginellaceae Selaginella amazonica Spring, D Selaginella arthritica Alston, S Selaginella breynii Spring, D Selaginella epirrhizos Spring, K, S Selaginella falcata (P. Beauv.) Spring, S Selaginella flagellata Spring, K, S Selaginella mazaruniense Jenman, K Selaginella novae-hollandiae (Sw.) Spring, S Selaginella parkeri (Hook. & Grev.) Spring, I, M, S, D Selaginella porelloides (Lam.) Spring, K, M Selaginella potaroensis Jenman, K Selaginella producta Baker, K, M Selaginella radiata (Aubl.) Spring, S Selaginella revoluta Baker, S Selaginella rhodostachya Baker, K Selaginella sandwithii Alston, K Selaginella sobolifera A.R. Sm., K Selaginella sulcata (Desv. ex Poir.) Spring ex Mart., S Selaginella tuberculata Spruce ex Baker, K Selaginella valdepilosa Baker, K Simaroubaceae Picramnia guianensis (Aubl.) Jans.-Jac., I, M Picramnia juniniana J.F. Macbr., D Picramnia latifolia Tul., I, S Picramnia macrostachys Klotzsch ex Engl., S Picramnia magnifolia J.F. Macbr., I Picramnia sellowii Planch., S Picrolemma sprucei Hook. f., D Simaba arborea ined., D Simaba cedron Planch., I, M, D Simaba guianensis Aubl., I, M, S, D Simaba monophylla (Oliv.) Cronquist, K Simaba multiflora A. Juss., M Simaba polyphylla (Cavalcante) W.W. Thomas, D Simaba sp. nov., D Simarouba amara Aubl., K, M, S, D I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

65 BRIT.ORG/SIDA 59 Smilacaceae Smilax cumanensis Humb. & Bonpl. ex Willd., M Smilax elastica Griseb., D Smilax krukovii A.C. Sm., I Smilax lasseriana Steyerm., I, M Smilax latipes Gleason, I, M Smilax poeppigii Kunth, M Smilax pseudosyphilitica Kunth, D Smilax riedeliana A. DC., S Smilax santaremensis A. DC., M Smilax saülensis J.D. Mitch., S Smilax schomburgkiana Kunth, I, K, M, S Smilax siphilitica Humb. & Bonpl. ex Willd., I, K, M, S, D Solanaceae Brunfelsia guianensis Benth., S Brunfelsia martiana Plowman, S, D Capsicum frutescens L., I, S Cestrum megalophyllum Dunal, I Cestrum racemosum Ruiz & Pav., I Cyphomandra endopogon Bitter, S Duckeodendron cestroides Kuhlm., D Lycianthes pauciflora (Vahl) Bitter, I, K, S Markea camponoti Ducke, I, S Markea coccinea Rich., M, S, D Markea longiflora Miers, I Markea sessiliflora Ducke, I, K, M, S Physalis angulata L., S Physalis pubescens L., K, S Schwenckia americana L., D Schwenckia grandiflora Benth., I Solanum adhaerens Roem. & Schult., M Solanum americanum Mill., S Solanum anceps Ruiz & Pav., S Solanum asperum Rich., I, S Solanum coriaceum Dunal, I, M, S Solanum costatum M. Nee, sp. nov. ined., S Solanum crinitum Lam., I, M, D Solanum fulvidum Bitter, D Solanum jamaicense Mill., S Solanum leucocarpon Dunal, I, M, S, D Solanum monachophyllum Dunal, I Solanum morii S. Knapp, S Solanum oppositifolium Ruiz & Pav., S Solanum paludosum Moric., M Solanum pensile Sendtn., I, S Solanum rubiginosum Vahl, M, S Solanum rugosum Dunal, M, S, D Solanum schlechtendalianum Walp., I, K, S, D Solanum sendtnerianum van Heurck & Müll. Arg., D Solanum stramoniifolium Hort. ex Benth., I, K, M, D Solanum subinerme Jacq., M, D Solanum torvum Sw., S Solanum velutinum Dunal, S Solanum sp. nov., M Sterculiaceae Byttneria cordifolia Sagot, D Byttneria divaricata Benth., I Byttneria morii L.C. Barnett & Dorr, S Herrania kanukuensis R.E. Schult., S Herrania lemniscata (M.R. Schomb.) R.E. Schult., I Melochia arenosa Benth., I Melochia melissifolia Benth., K Sterculia duckeana da Silva & Coelho, D Sterculia excelsa Mart., D Sterculia frondosa Rich., S, D Sterculia guianensis Sandwith, I, M Sterculia pruriens (Aubl.) K. Schum., I, M, S, D Sterculia rugosa R. Br., I, M Sterculia villifera Steud., I, S Theobroma cacao L., K, S Theobroma grandiflorum (Willd. ex Spreng.) K. Schum., D Theobroma subincanum Mart., D Theobroma sylvestre Mart., D Theobroma velutinum Benoist, S Waltheria indica L., I, M Strelitziaceae Phenakospermum guyannense (Rich.) Endl. ex Miq., I, K, S Styracaceae Styrax discolor M.F.Silva, D Styrax pallidus A. DC., S Styrax sieberi Perkins, S Styrax squamulosa M.F.Silva, D Styrax tessmannii Perkins, S Symplocaceae Symplocos martinicensis Jacq., S Symplocos sp., M Tectariaceae Ctenitis refulgens (Klotzsch ex Mett.) C. Chr. ex Vareschi, S Cyclopeltis semicordata (Sw.) J. Sm., I, M Lastreopsis effusa (Sw.) Tindale, S Tectaria heracleifolia (Willd.) Underw., I, S Tectaria incisa Cav., S Tectaria plantaginea (Jacq.) Maxon, I, K, M, S Tectaria trifoliata (L.) Cav., I, S Triplophyllum acutilobum Holttum, M Triplophyllum dicksonioides (Fée) Holttum, K, S, D Triplophyllum funestum (Kunze) Holttum, I, K, S, D Theaceae Ternstroemia dentata (Aubl.) Sw., D Ternstroemia punctata (Aubl.) Sw., I, K, M Ternstroemia schomburgkiana Benth., K Ternstroemia urophora Kobuski, D Thelypteridaceae Thelypteris abrupta (Desv.) Proctor, M, S Thelypteris arborescens (Humb. & Bonpl. ex Willd.) C.V. Morton, D Thelypteris decussata (L.) Proctor, S Thelypteris gemmulifera (Hieron.) A.R. Sm., S Thelypteris glandulosa (Desv.) Proctor, S Thelypteris hispidula (Decne.) C.F. Reed, S Thelypteris juruensis (C. Chr.) R.M. Tryon & D.S. Conant, S Thelypteris leprieurii (Hook.) R.M. Tryon, S Thelypteris macrophylla (Kunze) C.V. Morton, S Thelypteris nephrodioides (Klotzsch) Proctor, S I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

66 60 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana Thelypteris opulenta (Kaulf.) Fosberg, I, M Thelypteris pennata (Poir.) C.V. Morton, I, S Thelypteris poiteana (Bory) Proctor, I, S Thelypteris serrata (Cav.) Alston, S Thelypteris tetragona (Sw.) Small, I Theophrastaceae Clavija lancifolia Desf., I, S Clavija sp., D Thurniaceae Thurnia jenmanii Hook.f., K Thurnia sphaerocephala (Rudge) Hook.f., I, K, M, S, D Thymelaeaceae Daphnopsis longifolia Taub., I Daphnopsis sp., S Goodallia guianensis Benth., I Schoenobiblus daphnoides Mart., S, D Tiliaceae Apeiba glabra Aubl., S Apeiba petoumo Aubl., I, M, S, D Apeiba tibourbou Aubl., I, S Corchorus hirtus L., I Corchorus orinocensis Kunth, I Luehea speciosa Willd., S Lueheopsis rosea (Ducke) Burret, M, D Lueheopsis rugosa (Pulle) Burret, M, S Mollia lepidota Spruce ex Benth., M Mollia ulei Burret, K Vasivaea alchorneoides Baill., I, M Trigoniaceae Trigonia hypoleuca Griseb., I Trigonia laevis Aubl., I, K, M, S Trigonia nivea Cambess., I Trigonia subcymosa Benth., I Trigonia villosa Aubl., I, S Triuridaceae Peltophyllum luteum Gardner, M Sciaphila albescens Benth., M, S Sciaphila oligantha Maas, D Sciaphila picta Miers, D Sciaphila purpurea Benth., D Sciaphila rubra Maas, D Soridium spruceanum Miers, S Triuris hyalina Miers, M Turneraceae Piriqueta cistoides (L.) Griseb., I, D Turnera aurantiaca Benth., I, M Turnera curassavica Urb., I Turnera glaziovii Urb., S Turnera rupestris Aubl., I, M Turnera scabra Millsp., I Turnera subulata Sm., D Ulmaceae Ampelocera edentula Kuhlm., S, D Celtis iguanaea (Jacq.) Sarg., I, S Celtis schippii Standl., I Trema micrantha (L.) Blume, I, K, M, S Trema sp., D Urticaceae Boehmeria sp., S Helianthostylis sprucei Baill., D Laportea aestuans (L.) Chew, M, S Pilea imparifolia Wedd., S Pilea microphylla (L.) Liebm., S, D Pilea pubescens Liebm., S Urera caracasana (Jacq.) Griseb., I, S Urera laciniata Goudot ex Wedd., S Verbenaceae Aegiphila bracteolosa Moldenke, D Aegiphila integrifolia (Jacq.) B.D. Jacks., K, S, D Aegiphila intermedia Moldenke, D Aegiphila laevis (Aubl.) J.F. Gmel., M, S Aegiphila macrantha Ducke, M Aegiphila membranacea Turcz., I, S, D Aegiphila microcalycina Moldenke, D Aegiphila racemosa Vell., S Aegiphila villosa (Aubl.) J.F. Gmel., I, M, S Amasonia campestris (Aubl.) Moldenke, I, K Amasonia lasiocaulos Mart. & Schauer ex Schauer, D Citharexylum macrophyllum Poir., I, S Clerodendrum sp., S Lantana camara L., I, S, D Lantana cujabensis Schauer, D Lippia alba (Mill.) N.E. Br., S Petrea bracteata Steud., I, S, D Petrea brevicalyx Ducke, D Petrea insignis Schauer, D Petrea macrostachya Benth., I, M Petrea volubilis L., I, M, S Priva lappulacea (L.) Pers., S Stachytarpheta cayennensis (Rich.) Vahl, I, K, S, D Stachytarpheta jamaicensis (L.) Vahl, K, M, S Vitex calothyrsa Sandwith, D Vitex capitata Vahl, I Vitex compressa Turcz., M Vitex duckei Huber, D Vitex guianensis Moldenke, S Vitex spongiocarpa Ducke, D Vitex sprucei Briq., D Vitex stahelii Moldenke, I, M Vitex triflora Vahl, S, D Violaceae Amphirrhox longifolia (A. St.-Hil.) Spreng., I, M Amphirrhox surinamensis Eichler, D Corynostylis arborea (L.) S.F. Blake, I, S Gloeospermum sphaerocarpum Triana & Planch., S Hybanthus sp., S Leonia cymosa Mart., D Leonia glycycarpa Ruiz & Pav., S, D Noisettia orchidiflora (Rudge) Ging., I, S Paypayrola confertiflora Tul., I Paypayrola grandiflora Tul., D Paypayrola guianensis Aubl., I, M Paypayrola longifolia Tul., I, M Paypayrola sp., S Rinorea amapensis Hekking, S, D Rinorea brevipes (Benth.) S.F. Blake, M Rinorea endotricha Sandwith, M Rinorea falcata (Mart. ex Eichler) Kuntze, D Rinorea flavescens (Aubl.) Kuntze, I, K, D Rinorea guianensis Aubl., M, D I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

67 BRIT.ORG/SIDA 61 Rinorea lindeniana (Tul.) Kuntze, I Rinorea macrocarpa (Mart. ex Eichler) Kuntze, I, D Rinorea neglecta Sandwith, S Rinorea pubiflora (Benth.) Sprague & Sandwith, I, S, D Rinorea racemosa (Martt. & Zucc.) Kuntze, D Rinorea riana (DC. ex Ging.) Kuntze, nom. illeg., I, M, S Viscaceae Phoradendron acinacifolium Mart. ex Eichler, K Phoradendron crassifolium (Pohl ex DC.) Eichler, I, K, M, S, D Phoradendron fanshawei Maguire, K Phoradendron inaequidentatum Rusby, K, D Phoradendron obtusissimum (Miq.) Eichler, K Phoradendron perrottetii (DC.) Eichler, I, M, S Phoradendron piperoides (Kunth) Trel., I, K, M, S Phoradendron racemosum (Aubl.) Krug & Urb., M, S Vitaceae Cissus erosa Rich., I, M, S, D Cissus guyanensis Desc., D Cissus subrhomboidea (Baker) Planch., I Cissus verticillata (L.) Nicolson & C.E. Jarvis, I, M, S Cissus sp., K Vittariaceae Anetium citrifolium (L.) Splitg., I, S Antrophyum cajenense (Desv.) Spreng., K, S Antrophyum guayanense Hieron., I, K, M, S, D Antrophyum jenmanii Benedict, I, K, S Antrophyum lanceolatum (L.) Kaulf., K, S Hecistopteris kaieteurensis Kelloff & G. McKee, K Hecistopteris pumila (Spreng.) J. Sm., K, M, S, D Vittaria costata Kunze, I, K, M, S Vittaria graminifolia Kaulf., I Vittaria lineata (L.) Sm., M, S, D Vittaria remota Fée, S Vittaria stipitata Kunze, S Vochysiaceae Erisma bicolor Ducke, D Erisma floribundum Rudge, S Erisma uncinatum Warm., S Qualea acuminata Spruce ex Warm., D Qualea brevipedicellata Stafleu, D Qualea coerulea Aubl., S Qualea mori-boomii Marc.-Berti, S Qualea paraensis Ducke, D Acanthaceae Justicia calycina Acevedo-Rdgz., P.: 3351; Clarke, H.D.: 609, 636, 2495, 2517, 2526, 4212; McDowell, T.: 3253; Mori, S.: Justicia pectoralis McDowell, T.: 3359 Justicia schomburgkiana Clarke, H.D.: 399; Hoffman, B.: 1377; Mutchnick, P.: 415 Ruellia geminiflora McDowell, T.: 2043 Trichanthera gigantea Acevedo-Rdgz., P.: 3389 Adiantaceae Adiantopsis radiata Mori, S.: APPENDIX 2 IWOKRAMA PLANT COLLECTIONS Qualea rosea Aubl., S Ruizterania albiflora (Warm.) Marc.-Berti, M, S, D Ruizterania cassiquiarensis (Spruce ex Warm.) Marc.- Berti, D Ruizterania clavata (Stafleu) Marc.-Berti, D Ruizterania ferruginea (Steyerm.) Marc.-Berti, K Vochysia biloba Ducke, D Vochysia complicata Ducke, D Vochysia grandis Mart., D Vochysia guianensis Aubl., S Vochysia neyratii Normand, S Vochysia rufescens W.A.Rodr., D Vochysia schomburgkii Warm., M Vochysia surinamensis Stafleu, I, M, S Vochysia tetraphylla (G. Mey.) DC., I Vochysia tomentosa (G. Mey.) DC., S Woodsiaceae Diplazium celtidifolium Kunze, S Diplazium cristatum (Desr.) Alston, S Diplazium expansum Willd., S Diplazium grandifolium (Sw.) Sw., S Hemidictyum marginatum (L.) C. Presl, S Xyridaceae Abolboda acaulis Maguire, K Abolboda americana (Aubl.) Lanj., K Abolboda grandis Griseb., I, K Orectanthe sp., K Xyris cyperoides Gleason, K Xyris fallax Malme, I, K Xyris guianensis Klotzsch, K Xyris involucrata Nees, I, K Xyris jupicai Rich., I, K, D Xyris laxifolia Mart., S Xyris malmeana L.B. Sm., K Xyris setigera F. Oliv., K Xyris subuniflora Malme, K Xyris surinamensis Spreng., I, K Xyris uleana Malme, K Zingiberaceae Renealmia floribunda K. Schum., I, K, S, D Renealmia guianensis Maas, S Renealmia monosperma Miq., I, S Renealmia orinocensis Rusby, I, K, M, S Renealmia thrysoidea (Ruiz & Pav.) Poepp. & Endl., I Adiantum rondonii Mori, S.: Adiantum adiantoides Mutchnick, P.: 485 Adiantum cajennense Clarke, H.D.: 2608, 2714 Adiantum latifolium Acevedo-Rdgz., P.: 3297; Hoffman, B.: 1409; McDowell, T.: 3371 Adiantum latifolium x serratodentatum Hoffman, B.: 1297 Adiantum olivaceum Hoffman, B.: 1458 Adiantum phyllitidis Clarke, H.D.: 4307 Adiantum terminatum Clarke, H.D.: 4280; Ehringhaus, C.: 116 I = Iwokrama, K = Kaieteur Fallas, M = Mabura Hills, S = Saül (Central French Guiana), D = Reserva Ducke

68 62 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana Adiantum tomentosum Clarke, H.D.: 2473; Mori, S.: Pityrogramma calomelanos Hoffman, B.: 1339, 1492; McDowell, T.: 2027 Anacardiaceae Spondias mombin Acevedo-Rdgz., P.: 3436 Tapirira guianensis McDowell, T.: 2032; Mori, S.: 24510, 24612, 24616; Mutchnick, P.: 495, 882 Annonaceae Anaxagorea dolichocarpa Acevedo-Rdgz., P.: 3426; Clarke, H.D.: 313, 554, 592, 1400, 1415, 2512, 2567, 2629, 4198, 4251; Hoffman, B.: 1401, 1475, 1487, 4582, 4872, 4876; McDowell, T.: 2110; Mori, S.: 24585; Mutchnick, P.: 384, 752, 933 Annona haematantha Clarke, H.D.: 2576 Annona hypoglauca Clarke, H.D.: 1542; Hoffman, B.: 1367, 1391, 1503; McDowell, T.: 1979; Mutchnick, P.: 483, 866 Annona montana Hoffman, B.: 1537 Annona sericea Acevedo-Rdgz., P.: 3381; Clarke, H.D.: 545 Annona symphyocarpa Clarke, H.D.: 583, 2644; Mori, S.: Bocageopsis multiflora Hoffman, B.: 4615, 4986; Mutchnick, P.: 785 Cymbopetalum brasiliense Acevedo-Rdgz., P.: 3330; Clarke, H.D.: 4152, 4265; Mutchnick, P.: 961 Duguetia calycina Clarke, H.D.: 192, 526, 2518, 4318; Hoffman, B.: 4520, 4995, 5038 Duguetia inconspicua Clarke, H.D.: 4324, 4329 Duguetia neglecta Clarke, H.D.: 187, 1416, 2480, 4316; Hoffman, B.: 4587, 5001; Mutchnick, P.: 391 Duguetia obovata Mutchnick, P.: 392 Duguetia paraensis Clarke, H.D.: 1448; Hoffman, B.: 1468; Mori, S.: 24371; Mutchnick, P.: 393 Duguetia quitarensis McDowell, T.: 1898 Duguetia surinamensis Clarke, H.D.: 2566 Duguetia yeshidan Clarke, H.D.: 161, 229, 1275, 1455; Hoffman, B.: 5003 Fusaea longifolia Clarke, H.D.: 1396, 4330; McDowell, T.: 2106; Mutchnick, P.: 744 Guatteria citriodora Clarke, H.D.: 670 Guatteria scandens Clarke, H.D.: 330, 401, 623, 1335; Hoffman, B.: 4619; Mutchnick, P.: 742 Guatteria schomburgkiana Chanderbali, A.: 182; McDowell, T.: 2026 Guatteria sp. Clarke, H.D.: 575, 1429, 2496, 2515, 4139, 4195; Hoffman, B.: 1400; Mori, S.: 24486, 24530, 24559, Oxandra guianensis Hoffman, B.: 1463 Pseudoxandra sp. Mori, S.: Rollinia exsucca Acevedo-Rdgz., P.: 3452 Trigynaea caudata Clarke, H.D.: 233, 412 Trigynaea guianensis Hoffman, B.: 1465 Unonopsis guatterioides Clarke, H.D.: 539, 1544; Mori, S.: 24369; Mutchnick, P.: 461 Unonopsis rufescens Clarke, H.D.: 4194 Unonopsis stipitata Clarke, H.D.: 1466, 1560, 2729, 4308 Unonopsis sp. McDowell, T.: 3375 Xylopia aromatica McDowell, T.: 1986 Xylopia benthamii Clarke, H.D.: 1417; Ehringhaus, C.: 127 Xylopia discreta Acevedo-Rdgz., P.: 3405; Clarke, H.D.: 1558; Hoffman, B.: 1559 Xylopia nitida McDowell, T.: 2072, 2091 Apocynaceae Allamanda cathartica Acevedo-Rdgz., P.: 3340 Aspidosperma excelsum Clarke, H.D.: 678, 1293; Hoffman, B.: 4591, 4878 Aspidosperma vargasii Clarke, H.D.: 420 Bonafousia lorifera Clarke, H.D.: 328, 1350, 2500 Bonafousia siphilitica Ehringhaus, C.: 75; Hoffman, B.: 1509; McDowell, T.: 1980 Bonafousia undulata Clarke, H.D.: 171, 206, 207, 321, 408, 601, 1288, 1458, 1487, 2545, 2573, 2574, 2630; Hoffman, B.: 1490, 4577; Mutchnick, P.: 576, 930 Couma guianensis Clarke, H.D.: 370 Forsteronia guyanensis Acevedo-Rdgz., P.: 3449 Lacmellea aculeata Mutchnick, P.: 750 Malouetia gracilis Clarke, H.D.: 2763; McDowell, T.: 3345; Mutchnick, P.: 561 Malouetia tamaquarina Clarke, H.D.: 162, 2730 Mandevilla hirsuta Acevedo-Rdgz., P.: 3344; Clarke, H.D.: 1600 Mandevilla scabra Acevedo-Rdgz., P.: 3343, 3387 Mandevilla sp. Mutchnick, P.: 413 Mesechites trifida Clarke, H.D.: 690, 1531; Hoffman, B.: 1287; McDowell, T.: 1962; Mutchnick, P.: 456, 527, 665 Odontadenia macrantha Clarke, H.D.: 1348; Hoffman, B.: 1372; Mutchnick, P.: 434 Odontadenia nitida Clarke, H.D.: 2656 Prestonia acutifolia Clarke, H.D.: 306 Prestonia coalita McDowell, T.: 1968b Prestonia tomentosa Mutchnick, P.: 439 Tabernaemontana heterophylla Acevedo-Rdgz., P.: 3320; Clarke, H.D.: 1300; Hoffman, B.: 4954; Mutchnick, P.: 786 Tabernaemontana undulata Acevedo-Rdgz., P.: 3428; Hoffman, B.: 1338, 1402, 1460; McDowell, T.: 3258, 3290 Araceae Anthurium expansum Clarke, H.D.: 641, 2510, 4153; Mutchnick, P.: 980 Anthurium gracile Clarke, H.D.: 630, 1547, 4085; Hoffman, B.: 1508; Mutchnick, P.: 496, 515 Anthurium thrinax Clarke, H.D.: 1430 Anthurium trinerve Clarke, H.D.: 2638, 4098 Caladium bicolor Ehringhaus, C.: 48 Heteropsis flexuosa Clarke, H.D.: 1336, 4163, 4217; Hoffman, B.: 4640, 4645, 4817, 5052, 5060, 5061, 5066, 5067, 5070, 5071 Heteropsis melinonii Hoffman, B.: 5069 Heteropsis spruceana Clarke, H.D.: 1489; Mutchnick, P.: 571 Monstera adansonii Acevedo-Rdgz., P.: 3373; Clarke, H.D.: 4264 Monstera obliqua Hoffman, B.: 1528 Montrichardia arborescens Clarke, H.D.: 263; Mutchnick, P.: 435, 673 Montrichardia linifera Mutchnick, P.: 805 Philodendron camposportoanum McDowell, T.: 2066 Philodendron fragrantissimum Acevedo-Rdgz., P.:

69 BRIT.ORG/SIDA ; Clarke, H.D.: 513, 1276, 1497; Ehringhaus, C.: 65; Hoffman, B.: 1574, 5062; Mutchnick, P.: 735 Philodendron hylaeae Clarke, H.D.: 1498; Mutchnick, P.: 731 Philodendron insigne Ehringhaus, C.: 134 Philodendron jenmanii Hoffman, B.: 1470 Philodendron linnaei Hoffman, B.: 5068; Mutchnick, P.: 736 Philodendron muricatum Clarke, H.D.: 1372; McDowell, T.: 2016; Mutchnick, P.: 436 Philodendron myrmecophilum Clarke, H.D.: 157 Philodendron ornatum Mutchnick, P.: 936 Philodendron pedatum Clarke, H.D.: 4315; Hoffman, B.: 5055 Philodendron placidum Clarke, H.D.: 629 Philodendron rudgeanum Clarke, H.D.: 1511 Philodendron solimoësense Clarke, H.D.: 4344; Mutchnick, P.: 716 Philodendron surinamense Mutchnick, P.: 623 Rhodospatha oblongata Clarke, H.D.: 2501 Spathiphyllum cuspidatum Clarke, H.D.: 633, 2491, 4243, 4269; Ehringhaus, C.: 136; Hoffman, B.: 1459; Mori, S.: Stenospermation maguirei Clarke, H.D.: 2587, 4099, 4125 Syngonium podophyllum Acevedo-Rdgz., P.: 3327; Clarke, H.D.: 4289; Hoffman, B.: 1407; McDowell, T.: 2082 Urospatha sagittifolia Clarke, H.D.: 555; McDowell, T.: 3307 Arecaceae Astrocaryum gynacanthum Hoffman, B.: 1551 Astrocaryum sciophilum Clarke, H.D.: 1344; Mutchnick, P.: 855 Attalea maripa Mutchnick, P.: 807 Attalea microcarpa Hoffman, B.: 4631 Bactris balanophora McDowell, T.: 3306; Mori, S.: Bactris corossilla Clarke, H.D.: 4262, 4337 Bactris elegans Mutchnick, P.: 823 Bactris oligoclada Clarke, H.D.: 333, 487, 527; Mori, S.: 24653; Mutchnick, P.: 822, 852, 854 Bactris ptariana Clarke, H.D.: 2597; Mori, S.: Bactris simplicifrons Acevedo-Rdgz., P.: 3369; Clarke, H.D.: 2631; McDowell, T.: 3302; Mori, S.: 24398; Mutchnick, P.: 611, 613 Bactris sp. Clarke, H.D.: 531, 1292, 1309 Desmoncus orthacanthos Mutchnick, P.: 829 Desmoncus sp. Acevedo-Rdgz., P.: 3354 Euterpe oleracea Clarke, H.D.: 1284, 1427 Geonoma baculifera Clarke, H.D.: 486; Mutchnick, P.: 844 Geonoma deversa Clarke, H.D.: 1298, 4336; Mutchnick, P.: 820 Geonoma euspatha Hoffman, B.: 1418 Geonoma maxima Clarke, H.D.: 648, 2532, 4167; Hoffman, B.: 1403, 1577; McDowell, T.: 3367; Mutchnick, P.: 959 Geonoma sp. Clarke, H.D.: 403, 1316, 1490 Mauritiella armata Hoffman, B.: 4633 Oenocarpus bataua Hoffman, B.: 4589 Socratea exorrhiza Clarke, H.D.: 1320, 4266; Mutchnick, P.: 845, 849 Aristolochiaceae Aristolochia iquitensis Clarke, H.D.: 519; Mutchnick, P.: 573 Aristolochia surinamensis Acevedo-Rdgz., P.: 3308 Asclepiadaceae Matelea stenopetala Clarke, H.D.: 659 Tassadia guianensis Hoffman, B.: 1376 Aspleniaceae Asplenium angustum Clarke, H.D.: 4083 Asplenium auriculatum Clarke, H.D.: 4257 Asplenium cristatum Ehringhaus, C.: 66 Asplenium serratum Clarke, H.D.: 373, 634, 4082, 4225; Hoffman, B.: 1322, 1406, 1562; Mori, S.: 24412; Mutchnick, P.: 787 Asteraceae Calea caleoides Ehringhaus, C.: 93; McDowell, T.: 2040, 3273; Mori, S.: Calea solidaginea Mutchnick, P.: 769 Centratherum punctatum McDowell, T.: 1885 Chromolaena squalida Clarke, H.D.: 557 Clibadium surinamense Acevedo-Rdgz., P.: 3386 Clibadium sylvestre Hoffman, B.: 1552; McDowell, T.: 3366 Eclipta prostrata Clarke, H.D.: 1355 Eleutheranthera ruderalis Clarke, H.D.: 500 Gnaphalium polycaulon Mutchnick, P.: 766 Lepidaploa gracilis Hoffman, B.: 1316; McDowell, T.: 3275; Mutchnick, P.: 556 Mikania congesta Clarke, H.D.: 262; Mutchnick, P.: 666 Mikania gleasonii Clarke, H.D.: 4206; McDowell, T.: 2104 Mikania hookeriana Mori, S.: Mikania lindleyana Mutchnick, P.: 411 Mikania parviflora Clarke, H.D.: 1328; Mutchnick, P.: 840 Mikania psilostachya Mori, S.: Mikania trinitaria Acevedo-Rdgz., P.: 3339 Piptocarpha triflora Acevedo-Rdgz., P.: 3304 Rolandra fruticosa Clarke, H.D.: 561 Trichospira verticillata Ehringhaus, C.: 61; Hoffman, B.: 1270; McDowell, T.: 1975; Mutchnick, P.: 765 Wulffia baccata Acevedo-Rdgz., P.: 3323; Clarke, H.D.: 1602, 2685; Hoffman, B.: 1553; Mori, S.: 24575; Mutchnick, P.: 907 Balanophoraceae Helosis cayennensis Acevedo-Rdgz., P.: 3295; Clarke, H.D.: 1516, 1574, 2588 Begoniaceae Begonia guaduensis Hoffman, B.: 1564 Begonia heloisana Hoffman, B.: 1564a Bignoniaceae Adenocalymna inundatum Acevedo-Rdgz., P.: 3326 Anemopaegma chrysoleucum Hoffman, B.: 1379 Arrabidaea bilabiata Acevedo-Rdgz., P.: 3315; McDowell, T.: 1963 Arrabidaea candicans Mori, S.: Arrabidaea florida Mutchnick, P.: 591 Arrabidaea inaequalis Acevedo-Rdgz., P.: 3441

70 64 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana Clytostoma binatum Clarke, H.D.: 1548; Mutchnick, P.: 421 Cydista aequinoctialis Acevedo-Rdgz., P.: 3318, 3443; Clarke, H.D.: 269; McDowell, T.: 3270, 3334, 3339; Mutchnick, P.: 447 Distictella elongata McDowell, T.: 3250; Mutchnick, P.: 602 Jacaranda copaia Hoffman, B.: 4569, 4609, 4841 Jacaranda obtusifolia Clarke, H.D.: 2744; Hoffman, B.: 4546; McDowell, T.: 2025; Mutchnick, P.: 394, 824 Lundia densiflora Acevedo-Rdgz., P.: 3400 Memora racemosa Mori, S.: Memora schomburgkii Mutchnick, P.: 414 Mussatia hyacinthina Hoffman, B.: 4875 Parabignonia steyermarkii Clarke, H.D.: 4165 Paragonia pyramidata McDowell, T.: 3282 Pleonotoma clematis Mutchnick, P.: 626, 657 Tabebuia insignis Mori, S.: Tabebuia stenocalyx Mori, S.: Tanaecium jaroba Mutchnick, P.: 873 Tynanthus sp. Ehringhaus, C.: 94 Bixaceae Bixa orellana Clarke, H.D.: 2752 Cochlospermum orinocense Clarke, H.D.: 1501; Mutchnick, P.: 803, 913 Bombacaceae Catostemma fragrans Hoffman, B.: 4525; Mutchnick, P.: 802 Pachira aquatica Clarke, H.D.: 1333 Pachira insignis Clarke, H.D.: 349, 1442; Mutchnick, P.: 553, 869 Boraginaceae Cordia alliodora Chanderbali, A.: 164 Cordia exaltata Clarke, H.D.: 234, 366, 546 Cordia fallax Hoffman, B.: 4606 Cordia grandiflora Clarke, H.D.: 217, 1376, 2691, 2748 Cordia nodosa Acevedo-Rdgz., P.: 3451; Clarke, H.D.: 173, 430, 1469, 1493, 2530, 2722, 4168; Hoffman, B.: 1368, 4578; Mori, S.: 24572; Mutchnick, P.: 848 Cordia sagotii Clarke, H.D.: 516 Cordia schomburgkii McDowell, T.: 1939 Cordia tetrandra Hoffman, B.: 1370 Cordia sp. Mutchnick, P.: 578 Heliotropium filiforme Hoffman, B.: 1392; Mori, S.: Heliotropium ternatum Mutchnick, P.: 595 Tournefortia cuspidata Acevedo-Rdgz., P.: 3294 Tournefortia hirsutissima Clarke, H.D.: 304 Tournefortia melanochaeta Mori, S.: 24422, Tournefortia ulei McDowell, T.: 3356 Tournefortia sp. Mutchnick, P.: 637 Bromeliaceae Aechmea bromeliifolia Clarke, H.D.: 155 Aechmea mertensii Acevedo-Rdgz., P.: 3411; Clarke, H.D.: 467, 1538, 2652; McDowell, T.: 3333; Mutchnick, P.: 480 Aechmea tillandsioides Clarke, H.D.: 1343, 1556 Ananas parguazensis Clarke, H.D.: 376, 1505, 2766; McDowell, T.: 2024, 3380; Mutchnick, P.: 559, 737 Guzmania lingulata Clarke, H.D.: 427, 4311; Ehringhaus, C.: 133 Guzmania roezlii Clarke, H.D.: 4302 Pitcairnia sprucei Mutchnick, P.: 734 Burmanniaceae Campylosiphon purpurascens Clarke, H.D.: 682 Gymnosiphon breviflorus Clarke, H.D.: 1421 Gymnosiphon divaricatus Clarke, H.D.: 194; Ehringhaus, C.: 129; Mori, S.: Burseraceae Crepidospermum goudotianum Hoffman, B.: 1570 Crepidospermum rhoifolium Mutchnick, P.: 582 Protium decandrum Clarke, H.D.: 2474, 4339; Mori, S.: 24474, Protium guianense Clarke, H.D.: 661; Mutchnick, P.: 917, 939 Protium heptaphyllum Acevedo-Rdgz., P.: 3409; Clarke, H.D.: 394, 464, 2739; Hoffman, B.: 1305, 1388, 4551; McDowell, T.: 1985, 3264; Mutchnick, P.: 417, 685, 887 Protium opacum Clarke, H.D.: 2524, 4133, 4271 Protium sagotianum Hoffman, B.: 1568 Protium sp. Hoffman, B.: 5004 Tetragastris altissima Clarke, H.D.: 1567 Tetragastris hostmannii Mutchnick, P.: 796 Tetragastris panamensis Mutchnick, P.: 725 Cactaceae Epiphyllum phyllanthus Clarke, H.D.: 1286; McDowell, T.: 1969; Mutchnick, P.: 524, 654, 794 Campanulaceae Centropogon cornutus Clarke, H.D.: 618, 4215 Cannaceae Canna indica Hoffman, B.: 1351 Capparaceae Cleome sp. Hoffman, B.: 1365 Crateva tapia Clarke, H.D.: 309; Mutchnick, P.: 708 Caricaceae Jacaratia spinosa Clarke, H.D.: 4237; Hoffman, B.: 4917 Caryocaraceae Anthodiscus trifoliatus Clarke, H.D.: 167, 2765; McDowell, T.: 3265 Caryocar glabrum Mori, S.: Caryocar microcarpum Mutchnick, P.: 514 Caryocar montanum Clarke, H.D.: 579 Caryocar nuciferum Clarke, H.D.: 327; Mutchnick, P.: 490, 781 Cecropiaceae Coussapoa asperifolia Clarke, H.D.: 341, 1377; Mutchnick, P.: 871 Coussapoa microcephala Clarke, H.D.: 666; McDowell, T.: 3256 Coussapoa viridifolia Clarke, H.D.: 2640 Pourouma villosa Mutchnick, P.: 850 Celastraceae Goupia glabra Acevedo-Rdgz., P.: 3430b ; Hoffman, B.: 4612; McDowell, T.: 2087; Mutchnick, P.: 387 Maytenus guyanensis Clarke, H.D.: 2769 Maytenus sp. Acevedo-Rdgz., P.: 3312; Hoffman, B.: 1265, 1312; Mutchnick, P.: 721

71 BRIT.ORG/SIDA 65 Chrysobalanaceae Couepia bracteosa Clarke, H.D.: 197, 198, 253, 254, 525; Mutchnick, P.: 920 Couepia comosa Clarke, H.D.: 397, 688; Hoffman, B.: 1268; Mutchnick, P.: 686 Couepia multiflora Mutchnick, P.: 562 Couepia paraensis Hoffman, B.: 1321; Mutchnick, P.: 690 Couepia parillo Clarke, H.D.: 2598; Mori, S.: 24542, 24589; Mutchnick, P.: 940 Hirtella hispidula Hoffman, B.: 4555, 4853 Hirtella paniculata Clarke, H.D.: 280, 2755; McDowell, T.: 2029; Mutchnick, P.: 519, 651, 879 Hirtella racemosa Clarke, H.D.: 446, 1337, 2770; Mutchnick, P.: 525, 530 Hirtella silicea Clarke, H.D.: 1464, 2610, 4127; Hoffman, B.: 4608 Hirtella sp. Clarke, H.D.: 219; Hoffman, B.: 5006, 5019; Mori, S.: Licania alba Clarke, H.D.: 2472, 2479; Hoffman, B.: 4581; Mutchnick, P.: 580a Licania apetala Clarke, H.D.: 1370; McDowell, T.: 1996 Licania boyanii Clarke, H.D.: 532 Licania canescens Mutchnick, P.: 492 Licania coriacea Clarke, H.D.: 2746; McDowell, T.: 3343; Mori, S.: 24391; Mutchnick, P.: 528, 747, 772, 774, 777 Licania densiflora Clarke, H.D.: 1340; Hoffman, B.: 1421; McDowell, T.: 2068; Mutchnick, P.: 433 Licania discolor Hoffman, B.: 4571 Licania heteromorpha Mutchnick, P.: 779 Licania incana Clarke, H.D.: 236, 2655 Licania intrapetiolaris Mutchnick, P.: 521 Licania leptostachya Clarke, H.D.: 1331; Hoffman, B.: 1308; Mutchnick, P.: 764, 867 Licania licaniiflora Clarke, H.D.: 4210 Licania longistyla Clarke, H.D.: 524, 1425 Licania majuscula Clarke, H.D.: 1570 Licania parvifolia Mutchnick, P.: 705 Licania persaudii Clarke, H.D.: 312; Hoffman, B.: 4559; Mori, S.: Licania polita Hoffman, B.: 1371; Mori, S.: 24373; Mutchnick, P.: 659 Licania sprucei Mutchnick, P.: 847 Licania sp. Hoffman, B.: 4910 Parinari campestris Clarke, H.D.: 282, 346 Parinari rodolphii Hoffman, B.: 4538 Clusiaceae Caraipa grandifolia Clarke, H.D.: 390, 1532 Caraipa punctulata Clarke, H.D.: 4089; Hoffman, B.: 4598 Clusia columnaris Clarke, H.D.: 2546 Clusia flavida Chanderbali, A.: 183; Clarke, H.D.: 424, 646, 2547 Clusia fockeana Clarke, H.D.: 216, 224, 533; Hoffman, B.: 1444 Clusia grandiflora Clarke, H.D.: 220, 627; Hoffman, B.: 1461, 5059; Mori, S.: 24489; Mutchnick, P.: 929 Clusia myriandra Mutchnick, P.: 568, 916 Clusia nemorosa Clarke, H.D.: 2678; Mutchnick, P.: 810 Clusia palmicida Clarke, H.D.: 4164; Mori, S.: 24506, Clusia panapanari Clarke, H.D.: 2764; Hoffman, B.: 1307; Mori, S.: Clusia viscida Hoffman, B.: 4636 Clusia sp. Mutchnick, P.: 569 Garcinia benthamiana Clarke, H.D.: 2580; Mutchnick, P.: 783 Garcinia macrophylla Clarke, H.D.: 2711; Hoffman, B.: 4513 Garcinia madruño Clarke, H.D.: 2771 Rheedia macrophylla Hoffman, B.: 1404, 5044 Tovomita brevistaminea Hoffman, B.: 5046; Mori, S.: Tovomita carinata Clarke, H.D.: 2568 Tovomita cephalostigma Mutchnick, P.: 431 Tovomita eggersii Mori, S.: Tovomita longifolia Hoffman, B.: 4597 Tovomita macrophylla Mori, S.: Tovomita pyrifolia Mutchnick, P.: 941 Tovomita schomburgkii Clarke, H.D.: 674, 1568; Mori, S.: Vismia cayennensis Clarke, H.D.: 1583; McDowell, T.: 1937; Mutchnick, P.: 883 Vismia guianensis Clarke, H.D.: 544; Mutchnick, P.: 589 Vismia japurensis Ehringhaus, C.: 103; Mori, S.: 24472; Mutchnick, P.: 570 Vismia latifolia Acevedo-Rdgz., P.: 3392, 3394 Vismia macrophylla Clarke, H.D.: 556; Hoffman, B.: 1332, 1419, 1491; McDowell, T.: 2062, 2099 Vismia sessilifolia Clarke, H.D.: 4286; Mori, S.: Combretaceae Buchenavia macrophylla Clarke, H.D.: 271 Buchenavia megalophylla Clarke, H.D.: 1380, 2757; Hoffman, B.: 1320; Mori, S.: 24347; Mutchnick, P.: 440, 770 Combretum laxum Acevedo-Rdgz., P.: 3302; McDowell, T.: 2071; Mutchnick, P.: 458, 565, 835 Combretum rotundifolium Acevedo-Rdgz., P.: 3319; Clarke, H.D.: 285, 357, 1378; Hoffman, B.: 1299; McDowell, T.: 3329; Mutchnick, P.: 506, 512 Terminalia amazonia Hoffman, B.: 1478, 4566 Commelinaceae Commelina rufipes McDowell, T.: 3354 Dichorisandra hexandra McDowell, T.: 3293 Connaraceae Cnestidium guianense Mori, S.: Connarus lambertii Clarke, H.D.: 1334; Hoffman, B.: 1361; Mutchnick, P.: 644, 763 Connarus megacarpus Clarke, H.D.: 4171 Connarus sp. Hoffman, B.: 4899 Convolvulaceae Calycobolus glaber McDowell, T.: 3260 Dicranostyles ampla Clarke, H.D.: 169, 389 Dicranostyles holostyla Hoffman, B.: 4556 Ipomoea phillomega Clarke, H.D.: 4347 Ipomoea squamosa Mutchnick, P.: 653 Ipomoea tiliacea Clarke, H.D.: 1382; Mutchnick, P.: 906 Lysiostyles scandens McDowell, T.: 2067; Mori, S.: 24507; Mutchnick, P.: 800

72 66 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana Maripa scandens Clarke, H.D.: 477; Hoffman, B.: 4930; McDowell, T.: 2065 Merremia aturensis Clarke, H.D.: 1579 Merremia macrocalyx Clarke, H.D.: 1599; McDowell, T.: 2060, 3351; Mutchnick, P.: 581, 903 Costaceae Costus arabicus Hoffman, B.: 1296; Mutchnick, P.: 454 Costus scaber Acevedo-Rdgz., P.: 3356; Clarke, H.D.: 163, 658, 1311, 2609; Ehringhaus, C.: 74; McDowell, T.: 1891, 3372 Cucurbitaceae Cayaponia ophthalmica Hoffman, B.: 1480 Gurania bignoniacea Clarke, H.D.: 4141, 4186, 4288; Hoffman, B.: 1331; Mori, S.: Gurania lobata Clarke, H.D.: 621 Gurania spinulosa Clarke, H.D.: 4282 Gurania subumbellata Mori, S.: Psiguria triphylla Mutchnick, P.: 575 Cyatheaceae Cnemidaria spectabilis Clarke, H.D.: 4241 Cnemidaria spectablilis Clarke, H.D.: 4131 Cyathea cyatheoides Hoffman, B.: 1486 Cyathea macrocarpa Mutchnick, P.: 609 Cyathea microdonta Mutchnick, P.: 928 Cyathea pungens Clarke, H.D.: 4120, 4278a Cyathea surinamensis McDowell, T.: 3312; Mori, S.: Cyclanthaceae Asplundia maguirei Clarke, H.D.: 4273 Thoracocarpus bissectus Clarke, H.D.: 2716; Hoffman, B.: 4536; Mutchnick, P.: 445 Cyperaceae Becquerelia cymosa Clarke, H.D.: 672; Mutchnick, P.: 922 Bulbostylis juncoides Clarke, H.D.: 564; Ehringhaus, C.: 79 Calyptrocarya bicolor Clarke, H.D.: 2728 Calyptrocarya glomerulata Clarke, H.D.: 222, 251, 371, 647a, 1476, 1477, 2476, 4248, 4297; Hoffman, B.: 1386; McDowell, T.: 3311b ; Mori, S.: 24412a ; Mutchnick, P.: 616 Calyptrocarya poeppigiana Clarke, H.D.: 1302 Cyperus aggregatus Mutchnick, P.: 636 Cyperus filifolius Hoffman, B.: 1315 Cyperus haspan Clarke, H.D.: 2666; McDowell, T.: 2028 Cyperus ligularis Hoffman, B.: 1329 Cyperus luzulae Clarke, H.D.: 491; Ehringhaus, C.: 115; McDowell, T.: 3342; Mori, S.: Cyperus miliifolius Clarke, H.D.: 472; Mutchnick, P.: 635 Cyperus odoratus Mutchnick, P.: 890 Cyperus trailii Mutchnick, P.: 768 Diplacrum capitatum Clarke, H.D.: 550; Mori, S.: Diplasia karataefolia Acevedo-Rdgz., P.: 3355; Clarke, H.D.: 1283; Ehringhaus, C.: 112; Mori, S.: Eleocharis subfoliata Hoffman, B.: 1514 Fimbristylis complanata Clarke, H.D.: 1398 Fimbristylis miliacea McDowell, T.: 1978 Fimbristylis vahlii Mutchnick, P.: 767 Fuirena umbellata Mori, S.: Hypolytrum jenmanii Clarke, H.D.: 1397; Mori, S.: 24445, Hypolytrum longifolium Clarke, H.D.: 361, 473, 480, 1474 Hypolytrum pulchrum Clarke, H.D.: 2663; Hoffman, B.: 1429 Lagenocarpus guianensis Hoffman, B.: 1442 Mapania surinamensis Clarke, H.D.: 2529 Mapania sylvatica Clarke, H.D.: 635, 2577 Rhynchospora albomarginata Clarke, H.D.: 694 Rhynchospora amazonica Clarke, H.D.: 1478 Rhynchospora barbata Clarke, H.D.: 563; Ehringhaus, C.: 111; Hoffman, B.: 1430; Mori, S.: Rhynchospora cephalotes McDowell, T.: 1899; Mutchnick, P.: 566 Rhynchospora holoschoenoides Ehringhaus, C.: 113; Hoffman, B.: 1330; Mori, S.: Rhynchospora nervosa Clarke, H.D.: 1585 Rhynchospora reptans Clarke, H.D.: 437 Rhynchospora tenella Clarke, H.D.: 2553 Scleria cyperina Clarke, H.D.: 567 Scleria flagellum-nigorum Clarke, H.D.: 218 Scleria latifolia Mori, S.: Scleria macrogyne Clarke, H.D.: 244 Scleria melaleuca Ehringhaus, C.: 114; Mori, S.: Scleria microcarpa Hoffman, B.: 1313, 1517; McDowell, T.: 1960; Mutchnick, P.: 691 Scleria stipitata McDowell, T.: 1961 Scleria stipularis Mori, S.: Dennstaedtiaceae Lindsaea divaricata Mori, S.: Lindsaea guianensis Mutchnick, P.: 610 Lindsaea lancea Clarke, H.D.: 4115, 4317; McDowell, T.: 3310; Mori, S.: 24394; Mutchnick, P.: 612 Lindsaea quadrangularis McDowell, T.: 3320 Lindsaea stricta Hoffman, B.: 1431 Saccoloma elegans Clarke, H.D.: 1434 Saccoloma inaequale Clarke, H.D.: 4132 Dichapetalaceae Dichapetalum pedunculatum Hoffman, B.: 4981; Mori, S.: Tapura guianensis Acevedo-Rdgz., P.: 3423; Clarke, H.D.: 522, 591, 1468, 2478, 2717, 2724, 4197; Ehringhaus, C.: 69; Hoffman, B.: 4544, 4879; Mutchnick, P.: 579, 615, 751 Dilleniaceae Curatella americana Mutchnick, P.: 881 Davilla kunthii Clarke, H.D.: 552; Ehringhaus, C.: 104; McDowell, T.: 1958 Davilla nitida Mutchnick, P.: 830 Doliocarpus brevipedicellatus McDowell, T.: 2049 Doliocarpus guianensis McDowell, T.: 2079 Doliocarpus spraguei Clarke, H.D.: 1409 Doliocarpus sp. Hoffman, B.: 4944 Dioscoreaceae Dioscorea amazonum Clarke, H.D.: 560 Dioscorea pachycarpa Mori, S.: 24509

73 BRIT.ORG/SIDA 67 Dioscorea pilosiuscula Mutchnick, P.: 441 Dioscorea truncata Hoffman, B.: 4638 Dioscorea sp. Clarke, H.D.: 1502; Mutchnick, P.: 627 Droseraceae Drosera capillaris Hoffman, B.: 1435 Dryopteridaceae Cyclodium inerme Clarke, H.D.: 375, 2569, 4126, 4207; Mutchnick, P.: 487 Cyclodium meniscioides Clarke, H.D.: 4116; McDowell, T.: 3321; Mori, S.: 24395, 24557; Mutchnick, P.: 943 Ebenaceae Diospyros ierensis Clarke, H.D.: 4267 Diospyros lissocarpoides Clarke, H.D.: 227, 388, 1352, 1450; Hoffman, B.: 1283, 1374, 4912; Mori, S.: 24368, 24570; Mutchnick, P.: 522 Elaeocarpaceae Sloanea garckeana Clarke, H.D.: 1457 Sloanea grandiflora Clarke, H.D.: 2604, 4270; Hoffman, B.: 5012; Mori, S.: 24576; Mutchnick, P.: 782 Sloanea guianensis Clarke, H.D.: 4095 Sloanea latifolia Hoffman, B.: 1453 Sloanea parviflora Hoffman, B.: 4928 Eriocaulaceae Syngonanthus umbellatus Clarke, H.D.: 2658 Tonina fluviatilis McDowell, T.: 2036; Mori, S.: Erythroxylaceae Erythroxylum mucronatum Acevedo-Rdgz., P.: 3404; Clarke, H.D.: 2498 Euphorbiaceae Amanoa guianensis Hoffman, B.: 1385, 1499; McDowell, T.: 1973, 3378; Mutchnick, P.: 508 Aparisthmium cordatum Clarke, H.D.: 4130; Mori, S.: Croton trinitatis Clarke, H.D.: 1590; Hoffman, B.: 1340; McDowell, T.: 1974, 3350 Dalechampia affinis Hoffman, B.: 1324; Mori, S.: Dalechampia parvibracteata Clarke, H.D.: 176 Discocarpus essequiboensis Clarke, H.D.: 414, 1445; Hoffman, B.: 1479; Mori, S.: 24359; Mutchnick, P.: 502, 538, 754 Hyeronima oblonga Clarke, H.D.: 679 Mabea biglandulosa Clarke, H.D.: 2538 Mabea piriri Clarke, H.D.: 577; Hoffman, B.: 4897 Mabea speciosa Clarke, H.D.: 2635 Mabea taquari McDowell, T.: 2057; Mutchnick, P.: 655, 706, 821 Maprounea guianensis Hoffman, B.: 4558; McDowell, T.: 1988 Margaritaria nobilis Clarke, H.D.: 231; Mori, S.: Micrandra elata Clarke, H.D.: 381 Omphalea diandra Acevedo-Rdgz., P.: 3440 Pausandra martinii Clarke, H.D.: 594, 2606, 2628, 2682, 4201; Hoffman, B.: 1397, 4594, 4614, 4701, 4873, 5065; Mori, S.: Phyllanthus caroliniensis Clarke, H.D.: 2768 Phyllanthus juglandifolius Ehringhaus, C.: 51 Phyllanthus urinaria McDowell, T.: 3362 Plukenetia verrucosa Clarke, H.D.: 622 Ruprechtia brachysepala Mutchnick, P.: 862 Sagotia racemosa Clarke, H.D.: 329 Sandwithia guyanensis Clarke, H.D.: 2619 Sapium jenmanii Hoffman, B.: 4869 Sapium montanum Clarke, H.D.: 4342 Fabaceae-Caesal. Bauhinia guianensis Mutchnick, P.: 410 Bauhinia sp. Clarke, H.D.: 2687, 4172; Hoffman, B.: 4588 Campsiandra comosa Hoffman, B.: 1319; Mori, S.: Campsiandra laurifolia Clarke, H.D.: 279 Campsiandra sp. Clarke, H.D.: 292, 363, 441, 2709; Mutchnick, P.: 420 Cassia cowanii Clarke, H.D.: 2620; Mori, S.: Chamaecrista adiantifolia Clarke, H.D.: 537 Chamaecrista apoucouita Hoffman, B.: 4519; Mutchnick, P.: 395 Chamaecrista diphylla Clarke, H.D.: 1587b Chamaecrista hispidula Clarke, H.D.: 1582; McDowell, T.: 1997 Chamaecrista nictitans Clarke, H.D.: 502, 1413; Ehringhaus, C.: 101 Copaifera sp. Clarke, H.D.: 350; Hoffman, B.: 1561 Crudia glaberrima Mutchnick, P.: 861 Crudia sp. Clarke, H.D.: 303 Cynometra bauhiniifolia Mutchnick, P.: 757, 838 Cynometra sp. Clarke, H.D.: 299, 345 Dialium guianense Clarke, H.D.: 455; Mutchnick, P.: 442 Dicorynia guianensis Clarke, H.D.: 1358; Hoffman, B.: 1366; Mori, S.: Dimorphandra macrostachya Mori, S.: Elizabetha coccinea Clarke, H.D.: 168, 1327; Hoffman, B.: 1317; Mori, S.: 24362; Mutchnick, P.: 650, 758 Elizabetha princeps Clarke, H.D.: 4202; Hoffman, B.: 4877 Elizabetha sp. Mutchnick, P.: 563, 672 Eperua falcata Clarke, H.D.: 255, 680, 2613, 2719, 4084, 4211; Hoffman, B.: 4599, 4788; McDowell, T.: 2061; Mutchnick, P.: 438, 753 Eperua jenmanii Mutchnick, P.: 760 Eperua rubiginosa Clarke, H.D.: 191; Mutchnick, P.: 674 Eperua schomburgkiana Clarke, H.D.: 1353, 2743; McDowell, T.: 3335; Mutchnick, P.: 446, 675, 728 Eperua sp. Clarke, H.D.: 393, 442, 450; Mutchnick, P.: 536, 676, 684, 726 Heterostemon ellipticum Mutchnick, P.: 520 Macrolobium acaciifolium Clarke, H.D.: 457, 1373, 2704; Hoffman, B.: 1363, 1513; McDowell, T.: 1964, 3287; Mutchnick, P.: 460, 554, 831 Macrolobium bifolium Clarke, H.D.: 440; McDowell, T.: 3255; Mutchnick, P.: 409 Martiodendron excelsum Mutchnick, P.: 543, 645, 834 Martiodendron sp. Clarke, H.D.: 685 Mora excelsa Hoffman, B.: 4595, 4989; Mori, S.: 24377; Mutchnick, P.: 791

74 68 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana Paloue guianensis Hoffman, B.: 1454 Peltogyne venosa Clarke, H.D.: 281, 468; Mutchnick, P.: 400, 403 Senna latifolia Acevedo-Rdgz., P.: 3402; Clarke, H.D.: 4236 Senna quinquangulata Clarke, H.D.: 433, 511, 639; Mutchnick, P.: 489 Senna reticulata Mutchnick, P.: 713 Senna undulata Mutchnick, P.: 908 Swartzia benthamiana Clarke, H.D.: 2616; Hoffman, B.: 4522, 4531, 4537, 4947, 4951, 4966; Mori, S.: Swartzia laevicarpa Clarke, H.D.: 2646 Swartzia lamellata Acevedo-Rdgz., P.: 3433, 3437 Swartzia recurva Mutchnick, P.: 724 Swartzia tillettii Clarke, H.D.: 2564, 2595 Swartzia sp. Acevedo-Rdgz., P.: 3434; Clarke, H.D.: 540, 2603, 2623; Hoffman, B.: 4576, 4641, 4789, 4883, 4884, 4896, 5056, 5057; Mutchnick, P.: 494, 540 Tachigali guianensis Hoffman, B.: 4625 Tachigali sp. Clarke, H.D.: 277, 1364; Mutchnick, P.: 509, 856 Fabaceae-Mimos. Abarema jupunba Clarke, H.D.: 2541; McDowell, T.: 2059; Mutchnick, P.: 739 Abarema mataybifolia Clarke, H.D.: 2681, 2720 Acacia tenuifolia Clarke, H.D.: 432 Albizia subdimidiata Mutchnick, P.: 634, 877 Balizia pedicellaris McDowell, T.: 2070 Cedrelinga cateniformis Hoffman, B.: 4548 Hydrochorea corymbosa Clarke, H.D.: 686; McDowell, T.: 1971; Mutchnick, P.: 885 Inga alata McDowell, T.: 2080 Inga auristellae Clarke, H.D.: 4334 Inga capitata Clarke, H.D.: 4305 Inga disticha Clarke, H.D.: 261, 2699; Hoffman, B.: 1369; McDowell, T.: 3268, 3328 Inga leiocalycina Clarke, H.D.: 4304 Inga marginata Hoffman, B.: 4624 Inga nobilis Clarke, H.D.: 462; Mutchnick, P.: 388, 429, 541 Inga pezizifera Acevedo-Rdgz., P.: 3450; Clarke, H.D.: 4296 Inga rubiginosa Clarke, H.D.: 210 Inga sertulifera Clarke, H.D.: 290; Mutchnick, P.: 804 Inga thibaudiana Acevedo-Rdgz., P.: 3366; Clarke, H.D.: 1279, 1342 Inga umbellifera Hoffman, B.: 4521 Macrosamanea pubiramea Clarke, H.D.: 1524; Mutchnick, P.: 428 Mimosa myriadenia Acevedo-Rdgz., P.: 3391; Clarke, H.D.: 445, 1401, 1414; Mori, S.: 24461; Mutchnick, P.: 516 Mimosa pellita Clarke, H.D.: 689, 1375; Hoffman, B.: 1276; Mutchnick, P.: 470 Mimosa pigra Mutchnick, P.: 891 Mimosa pudica Clarke, H.D.: 665; Hoffman, B.: 1350; Mutchnick, P.: 910 Pentaclethra macroloba Clarke, H.D.: 166, 453, 2735; Hoffman, B.: 1349, 4585; McDowell, T.: 3289; Mori, S.: 24366; Mutchnick, P.: 401 Zygia cataractae Clarke, H.D.: 265, 297, 1367; Hoffman, B.: 1285, 1359, 1511 Zygia latifolia Clarke, H.D.: 1527, 2738; Mori, S.: 24348; Mutchnick, P.: 398 Zygia sp. Clarke, H.D.: 1449 Fabaceae-Papil. Acosmium nitens Clarke, H.D.: 379, 1379; McDowell, T.: 1987; Mutchnick, P.: 547, 888 Aeschynomene rudis Mutchnick, P.: 837 Aeschynomene sensitiva Mutchnick, P.: 598 Aeschynomene sp. Clarke, H.D.: 569, 1389, 1399 Alexa imperatricis Clarke, H.D.: 1305 Alexa sp. Clarke, H.D.: 287 Centrosema vexillatum Ehringhaus, C.: 59 Centrosema sp. McDowell, T.: 1982 Clathrotropis brachypetala Acevedo-Rdgz., P.: 3328; Hoffman, B.: 4771; Mutchnick, P.: 743 Clathrotropis glaucophylla Hoffman, B.: 4596 Clathrotropis macrocarpa Clarke, H.D.: 4086; Hoffman, B.: 4635, 4852; Mori, S.: 24476, Clathrotropis sp. Hoffman, B.: 4563, 4845 Dalbergia monetaria McDowell, T.: 3330; Mutchnick, P.: 718 Dalbergia riedelii Clarke, H.D.: 456; Mori, S.: 24372; Mutchnick, P.: 482 Dalbergia sp. Clarke, H.D.: 1392, 1536 Desmodium adscendens Clarke, H.D.: 1577; McDowell, T.: 3364 Desmodium axillare Hoffman, B.: 1527 Desmodium sp. Clarke, H.D.: 497 Dioclea guianensis Acevedo-Rdgz., P.: 3313, 3388 Dioclea macrantha Hoffman, B.: 1272 Dioclea megacarpa Clarke, H.D.: 4216 Dioclea scabra Mutchnick, P.: 416 Dioclea virgata Hoffman, B.: 1271b Dioclea sp. Clarke, H.D.: 343, 603, 1523, 2750; Mutchnick, P.: 914 Eriosema simplicifolium Ehringhaus, C.: 76 Eriosema sp. Clarke, H.D.: 571 Etaballia dubia Clarke, H.D.: 1332, 1521; Hoffman, B.: 1291; Mutchnick, P.: 412 Galactia sp. Clarke, H.D.: 1580; Hoffman, B.: 1519 Hymenolobium sp. Hoffman, B.: 4552 Lonchocarpus sp. Clarke, H.D.: 687, 2694; Mutchnick, P.: 548, 649 Machaerium ferox Clarke, H.D.: 1351; Mutchnick, P.: 542, 697 Machaerium floribundum Clarke, H.D.: 451; Mutchnick, P.: 567, 868 Machaerium inundatum Hoffman, B.: 1357, 1560; McDowell, T.: 3281; Mutchnick, P.: 647, 720, 755 Machaerium kegelii Mutchnick, P.: 911 Machaerium leiophyllum Clarke, H.D.: 1533; Mutchnick, P.: 535 Machaerium quinata Hoffman, B.: 5037; Mutchnick, P.: 463 Machaerium sp. Clarke, H.D.: 340, 1349, 2753 Macroptilium gracile McDowell, T.: 1946 Mucuna urens Clarke, H.D.: 4351 Mucuna sp. Clarke, H.D.: 1551 Ormosia coarctata Hoffman, B.: 1364; Mutchnick, P.: 437

75 BRIT.ORG/SIDA 69 Ormosia coccinea Clarke, H.D.: 278, 352, 447 Ormosia coutinhoi Clarke, H.D.: 523 Ormosia fastigiata McDowell, T.: 2034 Ormosia sp. Clarke, H.D.: 2754 Paramachaerium schomburgkii Clarke, H.D.: 1368; Mutchnick, P.: 832 Vigna lasiocarpa McDowell, T.: 1981, 2012 Vigna repens McDowell, T.: 2039 Vigna sp. Clarke, H.D.: 344, 1356; Mutchnick, P.: 418 Zornia sp. Clarke, H.D.: 1587 Flacourtiaceae Casearia commersoniana Hoffman, B.: 1541 Casearia singularis Hoffman, B.: 4898 Casearia sylvestris Clarke, H.D.: 2651 Homalium guianense Mutchnick, P.: 427 Homalium racemosum Clarke, H.D.: 1520; Mutchnick, P.: 529 Neosprucea sp. McDowell, T.: 2103 Gentianaceae Coutoubea ramosa Clarke, H.D.: 499; Ehringhaus, C.: 97; Hoffman, B.: 1271a ; Mori, S.: 24349; Mutchnick, P.: 704, 899 Irlbachia alata Acevedo-Rdgz., P.: 3350; Clarke, H.D.: 505; Ehringhaus, C.: 107; Mori, S.: 24436; Mutchnick, P.: 593 Irlbachia caerulescens McDowell, T.: 2051 Irlbachia purpurascens Clarke, H.D.: 653, 2555; Hoffman, B.: 1440 Voyria aphylla Clarke, H.D.: 530, 2673; Ehringhaus, C.: 130; Mori, S.: 24386, Voyria caerulea Clarke, H.D.: 326 Voyria corymbosa Clarke, H.D.: 154, 256, 520, 2621; Ehringhaus, C.: 128; McDowell, T.: 3305a ; Mori, S.: 24446; Mutchnick, P.: 617 Voyria rosea Clarke, H.D.: 2622 Voyriella parviflora Clarke, H.D.: 338 Gesneriaceae Besleria flavo-virens Clarke, H.D.: 4143 Besleria laxiflora Clarke, H.D.: 615, 2525, 4258 Chrysothemis rupestris Hoffman, B.: 1395 Codonanthe calcarata Clarke, H.D.: 319, 444, 599, 645, 1495, 1515, 2596, 2653, 4213; Mutchnick, P.: 405, 544, 792 Codonanthe crassifolia Clarke, H.D.: 250, 443, 1322 Episcia xantha Clarke, H.D.: 4214, 4260; Mori, S.: Nautilocalyx cordatus Clarke, H.D.: 1439, 4112 Paradrymonia densa Clarke, H.D.: 638, 2520 Paradrymonia maculata Clarke, H.D.: 4181 Gleicheniaceae Dicranopteris flexuosa Mori, S.: Gnetaceae Gnetum nodiflorum Mutchnick, P.: 828 Gnetum paniculatum Hoffman, B.: 4643; Mutchnick, P.: 533 Gnetum urens Mori, S.: Gnetum sp. McDowell, T.: 2045 Grammitidaceae Cochlidium furcatum Clarke, H.D.: 2536; Mori, S.: Cochlidium linearifolium Clarke, H.D.: 4106 Cochlidium serrulatum Clarke, H.D.: 4287 Haemodoraceae Xiphidium caeruleum Hoffman, B.: 1573; McDowell, T.: 3355 Heliconiaceae Heliconia acuminata Clarke, H.D.: 180, 322, 1296, 1500, 2483, 4224; McDowell, T.: 3379; Mutchnick, P.: 584, 733 Heliconia bihai Clarke, H.D.: 4149; Hoffman, B.: 1550 Heliconia chartacea Clarke, H.D.: 474, 1594, 4345; Hoffman, B.: 1493, 1549; McDowell, T.: 1884 Heliconia hirsuta Acevedo-Rdgz., P.: 3337; McDowell, T.: 1954 Heliconia lourteigiae Mori, S.: Heliconia psittacorum Acevedo-Rdgz., P.: 3372 Heliconia spathocircinata Acevedo-Rdgz., P.: 3325 Hippocrateaceae Hippocratea volubilis Clarke, H.D.: 260, 298, 449, 2698; Hoffman, B.: 1266, 1354; McDowell, T.: 3262, 3331, 3332; Mutchnick, P.: 510, 875 Peritassa compta Clarke, H.D.: 482, 1357; Mutchnick, P.: 539, 801 Peritassa laevigata Hoffman, B.: 1352 Peritassa sp. Clarke, H.D.: 1488 Prionostemma aspera Clarke, H.D.: 1603 Salacia elliptica Mutchnick, P.: 876 Salacia impressifolia Mori, S.: Salacia juruana Clarke, H.D.: 1306, 4091 Salacia maburensis Clarke, H.D.: 320, 1443; Mutchnick, P.: 969 Salacia multiflora Mori, S.: Tontelea ovalifolia Clarke, H.D.: 514 Humiriaceae Humiria balsamifera Clarke, H.D.: 226, 656, 2654 Schistostemon dichotomum Hoffman, B.: 4564 Vantanea guianensis Hoffman, B.: 1447 Hymenophyllaceae Hymenophyllum polyanthos Clarke, H.D.: 2586 Trichomanes ankersii Clarke, H.D.: 4157; Mori, S.: 24399, Trichomanes arbuscula Clarke, H.D.: 535; Mori, S.: 24405; Mutchnick, P.: 603 Trichomanes crispum Mori, S.: Trichomanes diaphanum Mori, S.: Trichomanes diversifrons Clarke, H.D.: 4118 Trichomanes martiusii Mutchnick, P.: 618 Trichomanes pedicellatum Clarke, H.D.: 2726, 4101; Mori, S.: Trichomanes pinnatum Clarke, H.D.: 675, 1571, 2499, 2572; Mori, S.: 24397, Trichomanes trollii Clarke, H.D.: 652, 2570 Icacinaceae Emmotum fagifolium Clarke, H.D.: 668 Lacistemataceae Lacistema aggregatum Clarke, H.D.: 585; Hoffman, B.: 4543, 4843 Lacistema sp. McDowell, T.: 2050

76 70 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana Lamiaceae Hyptis lanceolata Clarke, H.D.: 510 Hyptis parkeri McDowell, T.: 3349 Ocimum campechianum McDowell, T.: 2030 Lauraceae Aiouea guianensis Chanderbali, A.: 176, 260 Aiouea laevis Clarke, H.D.: 2650 Aniba citrifolia McDowell, T.: 2074, 2086 Aniba excelsa Chanderbali, A.: 145, 156 Aniba hostmanniana Clarke, H.D.: 2477 Aniba hypoglauca Chanderbali, A.: 165 Aniba megaphylla Hoffman, B.: 4627 Aniba panurensis Chanderbali, A.: 180, 248, 266; Clarke, H.D.: 4309 Cassytha filiformis Clarke, H.D.: 703; McDowell, T.: 1950 Chlorocardium rodiei Chanderbali, A.: 142; Clarke, H.D.: 188, 538, 1303; Hoffman, B.: 4580; Mori, S.: 24487; Mutchnick, P.: 732 Endlicheria chalisea Chanderbali, A.: 141, 159; Clarke, H.D.: 1465 Endlicheria multiflora Chanderbali, A.: 147, 151, 152, 153, 269; Hoffman, B.: 1358; Mutchnick, P.: 687 Endlicheria sp. Acevedo-Rdgz., P.: 3432 Licaria cannella Chanderbali, A.: 174, 178, 261; Clarke, H.D.: 2642 Licaria debilis Chanderbali, A.: 272; Clarke, H.D.: 4314 Licaria guianensis Clarke, H.D.: 2579 Licaria martiniana Chanderbali, A.: 163 Licaria sp. McDowell, T.: 2014 Mezilaurus itauba Chanderbali, A.: 173 Nectandra amazonum Chanderbali, A.: 148, 150, 161; Clarke, H.D.: 270, 1546, 2695; Hoffman, B.: 1390, 1484; McDowell, T.: 3263; Mutchnick, P.: 532, 643 Nectandra globosa Chanderbali, A.: 143, 253; Clarke, H.D.: 483 Ocotea canaliculata Chanderbali, A.: 157 Ocotea ceanothifolia Chanderbali, A.: 154, 158, 168 Ocotea cernua Chanderbali, A.: 250 Ocotea guianensis Clarke, H.D.: 2684 Ocotea nigra Chanderbali, A.: 146, 162, 251, 268 Ocotea persulcata Chanderbali, A.: 181 Ocotea schomburgkiana Chanderbali, A.: 166, 167, 175, 259 Ocotea splendens Chanderbali, A.: 179 Ocotea sp. Clarke, H.D.: 4150; Mutchnick, P.: 965 Rhodostemonodaphne crenaticupula Chanderbali, A.: 160 Rhodostemonodaphne kunthiana Chanderbali, A.: 171, 172, 257; Clarke, H.D.: 4277 Rhodostemonodaphne morii Clarke, H.D.: 2689 Rhodostemonodaphne praeclara Chanderbali, A.: 144, 177 Rhodostemonodaphne scandens Clarke, H.D.: 647b, 2527, 4113; Mori, S.: 24529; Mutchnick, P.: 945 Lecythidaceae Couratari stellata Hoffman, B.: 4575 Eschweilera coriacea Clarke, H.D.: 2600, 2734, 4209, 4291 Eschweilera decolorans Clarke, H.D.: 2612; Mori, S.: Eschweilera pedicellata Clarke, H.D.: 230, 1573; Hoffman, B.: 4517, 4850, 4933, 5018, 5032; Mori, S.: 24381, 24516; Mutchnick, P.: 422 Eschweilera sagotiana Clarke, H.D.: 186, 572, 1299; Mori, S.: Eschweilera simiorum Clarke, H.D.: 4333 Eschweilera subglandulosa Acevedo-Rdgz., P.: 3446; Clarke, H.D.: 190, 273, 391; Mori, S.: 24380; Mutchnick, P.: 679, 778 Eschweilera wachenheimii Hoffman, B.: 4601; Mori, S.: 24444; Mutchnick, P.: 399, 419, 931 Eschweilera sp.a Hoffman, B.: 5035 Eschweilera sp. C Hoffman, B.: 4565 Eschweilera sp. Hoffman, B.: 4892, 4993; Mutchnick, P.: 678 Gustavia augusta Acevedo-Rdgz., P.: 3413; Clarke, H.D.: 1557; Hoffman, B.: 1284; McDowell, T.: 1893, 1900; Mutchnick, P.: 537, 759 Gustavia hexapetala Mori, S.: Lecythis alutacea Mori, S.: 24622; Mutchnick, P.: 555 Lecythis confertiflora Clarke, H.D.: 2471, 4087, 4088 Lecythis corrugata Acevedo-Rdgz., P.: 3438; Mori, S.: 24379; Mutchnick, P.: 799 Lecythis holcogyne Clarke, H.D.: 1459; Mori, S.: Lentibulariaceae Utricularia foliosa Acevedo-Rdgz., P.: 3408 Utricularia juncea Clarke, H.D.: 673 Utricularia pusilla McDowell, T.: 3296; Mori, S.: Utricularia sp. Hoffman, B.: 1443 Liliaceae-Amaryll. Hymenocallis tubiflora Hoffman, B.: 1529; Mori, S.: Liliaceae-Hypox. Curculigo scorzonerifolia McDowell, T.: 1949 Lissocarpaceae Lissocarpa guianensis Clarke, H.D.: 252 Loganiaceae Spigelia mulitspica Clarke, H.D.: 2487 Spigelia multispica Clarke, H.D.: 608, 4246; Mori, S.: Strychnos cogens Clarke, H.D.: 578; Hoffman, B.: 4867; Mori, S.: 24607; Mutchnick, P.: 468, 870 Strychnos guianensis Hoffman, B.: 4813 Strychnos peckii Mutchnick, P.: 863 Lomariopsidaceae Bolbitis semipinnatifida Clarke, H.D.: 4146, 4244, 4259; Mori, S.: 24553; Mutchnick, P.: 967 Lomariopsis japurensis Clarke, H.D.: 4341 Loranthaceae Oryctanthus alveolatus Clarke, H.D.: 311 Oryctanthus florulentus Hoffman, B.: 1378; Mutchnick, P.: 629, 658, 833 Phthirusa rufa Clarke, H.D.: 243 Phthirusa stelis McDowell, T.: 2083; Mutchnick, P.: 894 Phthirusa sp. Clarke, H.D.: 1519 Lycopodiaceae Lycopodiella caroliniana Clarke, H.D.: 2660 Lycopodiella cernua Clarke, H.D.: 558, 2665

77 BRIT.ORG/SIDA 71 Lygodiaceae Lygodium micans Acevedo-Rdgz., P.: 3383 Lygodium volubile Ehringhaus, C.: 72 Lythraceae Cuphea antisyphilitica Clarke, H.D.: 695 Malpighiaceae Banisteriopsis martiniana Clarke, H.D.: 655; Ehringhaus, C.: 98 Bunchosia argentea Hoffman, B.: 4533 Byrsonima aerugo Clarke, H.D.: 1403; Mori, S.: Byrsonima crassifolia Clarke, H.D.: 570; Ehringhaus, C.: 77; Mutchnick, P.: 880 Byrsonima eugeniifolia Clarke, H.D.: 2667 Byrsonima stipulacea Hoffman, B.: 1424; Mori, S.: 24601, Heteropterys cristata Mutchnick, P.: 406 Heteropterys multiflora Clarke, H.D.: 1387 Heteropterys nervosa Clarke, H.D.: 553 Heteropterys velutina Clarke, H.D.: 1541 Heteropterys sp. Mutchnick, P.: 811 Hiraea faginea Clarke, H.D.: 289, 305, 1325, 2701; Hoffman, B.: 1288; Mutchnick, P.: 648, 695 Hiraea gracieana Clarke, H.D.: 660 Mascagnia sepium Mutchnick, P.: 513 Spachea elegans Mutchnick, P.: 825 Stigmaphyllon sinuatum Acevedo-Rdgz., P.: 3309; Clarke, H.D.: 335, 405, 506, 1565; Ehringhaus, C.: 90; Hoffman, B.: 1446; Mori, S.: 24450; Mutchnick, P.: 585, 680, 912 Tetrapterys crispa McDowell, T.: 2101 Tetrapterys discolor Acevedo-Rdgz., P.: 3311; Clarke, H.D.: 461, 1529; McDowell, T.: 1886; Mutchnick, P.: 467, 517 Tetrapterys fimbripetala Mori, S.: Tetrapterys mucronata Clarke, H.D.: 2731; Mutchnick, P.: 631, 632, 664, 715 Tetrapterys styloptera McDowell, T.: 3376 Malvaceae Hibiscus bifurcatus Clarke, H.D.: 459; Mutchnick, P.: 614 Hibiscus furcellatus Acevedo-Rdgz., P.: 3406 Hibiscus sororius Mutchnick, P.: 552 Pavonia castanaefolia McDowell, T.: 1890 Peltaea trinervis McDowell, T.: 3272 Sida cordifolia Clarke, H.D.: 1588 Sida glomerata McDowell, T.: 3288 Urena lobata McDowell, T.: 1945, 3274 Wissadula patens McDowell, T.: 1895 Marantaceae Calathea elliptica Acevedo-Rdgz., P.: 3368; Clarke, H.D.: 1507, 1561, 4226; Ehringhaus, C.: 68; Hoffman, B.: 1411; Mori, S.: Calathea legrelleana McDowell, T.: 3373 Calathea lutea Acevedo-Rdgz., P.: 3359 Ischnosiphon arouma Clarke, H.D.: 4323; Hoffman, B.: 1373 Ischnosiphon foliosus Clarke, H.D.: 515, 643, 1423; Ehringhaus, C.: 117; Mori, S.: Ischnosiphon gracilis Mori, S.: Ischnosiphon obliquus Clarke, H.D.: 657; Hoffman, B.: 1540, 4528; Mori, S.: Ischnosiphon puberulus Mori, S.: Maranta gibba McDowell, T.: 1892 Maranta protracta Acevedo-Rdgz., P.: 3371; Hoffman, B.: 1539 Monotagma spicatum Clarke, H.D.: 368, 1295; Hoffman, B.: 1410; McDowell, T.: 3303; Mori, S.: 24496; Mutchnick, P.: 604, 605 Marattiaceae Danaea elliptica Clarke, H.D.: 4148 Danaea simplicifolia Clarke, H.D.: 4187 Marcgraviaceae Marcgravia parviflora Clarke, H.D.: 4119, 4275; Mori, S.: 24536, Norantea guianensis Mutchnick, P.: 748 Souroubea guianensis Clarke, H.D.: 448, 4301; Mutchnick, P.: 396 Melastomataceae Aciotis laxa Hoffman, B.: 1457; McDowell, T.: 3318 Aciotis purpurascens Acevedo-Rdgz., P.: 3380; Ehringhaus, C.: 95; Hoffman, B.: 1344; Mutchnick, P.: 572 Bellucia grossularioides Hoffman, B.: 1542; Mutchnick, P.: 594 Bellucia pentamera Mutchnick, P.: 511 Clidemia conglomerata Mori, S.: Clidemia hirta McDowell, T.: 1951, 3314 Clidemia sericea McDowell, T.: 2031 Comolia vernicosa Hoffman, B.: 1427 Comolia villosa Hoffman, B.: 1437 Henriettea multiflora Mutchnick, P.: 919 Henriettea stellaris Mutchnick, P.: 423 Leandra solenifera Hoffman, B.: 1497 Macrocentrum cristatum Hoffman, B.: 1456; Mutchnick, P.: 955 Maieta poeppigii Ehringhaus, C.: 138; McDowell, T.: 3304; Mutchnick, P.: 620 Miconia alata McDowell, T.: 1938 Miconia aplostachya Hoffman, B.: 1501; McDowell, T.: 1968a Miconia argyrophylla Acevedo-Rdgz., P.: 3376 Miconia bracteata McDowell, T.: 3294 Miconia ceramicarpa Mutchnick, P.: 474, 978 Miconia chrysophylla Hoffman, B.: 5031 Miconia ciliata Hoffman, B.: 1557; McDowell, T.: 2033 Miconia fallax Hoffman, B.: 1556 Miconia gratissima Hoffman, B.: 1423 Miconia macrothyrsa McDowell, T.: 1942; Mutchnick, P.: 902 Miconia mariae Mutchnick, P.: 975 Miconia myriantha Hoffman, B.: 1422 Miconia prasina Hoffman, B.: 1554 Miconia rufescens Ehringhaus, C.: 80; Hoffman, B.: 1555 Mouriri grandiflora Hoffman, B.: 1381 Myriaspora egensis Acevedo-Rdgz., P.: 3379; Hoffman, B.: 1524 Tibouchina aspera Ehringhaus, C.: 78; Mutchnick, P.: 905 Tococa aristata Ehringhaus, C.: 100, 135; McDowell, T.: 3308

78 72 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana Tococa nitens Hoffman, B.: 1428 Tococa subciliata Hoffman, B.: 1353 Meliaceae Carapa guianensis Acevedo-Rdgz., P.: 3431; Clarke, H.D.: 365, 1304; Hoffman, B.: 4593, 4882; Mutchnick, P.: 383, 843 Guarea guidonia Hoffman, B.: 1526 Guarea kunthiana Hoffman, B.: 4946 Guarea pubescens Clarke, H.D.: 606, 1460, 2594; Mori, S.: 24527; Mutchnick, P.: 946 Trichilia cipo Mutchnick, P.: 780, 817 Trichilia martiana Hoffman, B.: 4567, 4967 Trichilia pallida Ehringhaus, C.: 55; Hoffman, B.: 1572, 4509 Trichilia quadrijuga Hoffman, B.: 5045 Trichilia schomburgkii Acevedo-Rdgz., P.: 3298; Clarke, H.D.: 2589, 2647, 4219, 4328; Hoffman, B.: 4948 Trichilia sp. Clarke, H.D.: 2605 Mendonciaceae Mendoncia hoffmannseggiana McDowell, T.: 2085; Mori, S.: 24583, Mendoncia squamuligera Mori, S.: Menispermaceae Abuta sp. Clarke, H.D.: 4208 Cissampelos andromorpha Hoffman, B.: 1547 Curarea candicans Clarke, H.D.: 2723; Hoffman, B.: 4616 Orthomene schomburgkii Clarke, H.D.: 170, 2702; Hoffman, B.: 1387; McDowell, T.: 3337 Sciadotenia cayennensis Mutchnick, P.: 574 Metaxyaceae Metaxya rostrata Clarke, H.D.: 374; Ehringhaus, C.: 125; Hoffman, B.: 1455; McDowell, T.: 3322; Mori, S.: 24415; Mutchnick, P.: 486, 789, 925 Monimiaceae Mollinedia grazielae Clarke, H.D.: 582, 2583, 2645; Hoffman, B.: 4868; Mori, S.: 24587, 24603; Mutchnick, P.: 476 Mollinedia laurina Clarke, H.D.: 2544 Siparuna cuspidata Clarke, H.D.: 318; Mori, S.: Siparuna decipiens Clarke, H.D.: 372, 1291, 1461, 2727, 4346; Hoffman, B.: 4931; Mori, S.: Siparuna guianensis Acevedo-Rdgz., P.: 3338, 3396; Clarke, H.D.: 179, 212, 409, 548, 1595; Hoffman, B.: 1326, 1536; McDowell, T.: 3358; Mori, S.: Moraceae Brosimum guianense Mutchnick, P.: 797 Brosimum lactescens Clarke, H.D.: 307 Brosimum utile Mori, S.: Clarisia ilicifolia Clarke, H.D.: 165 Ficus malacocarpa McDowell, T.: 3252 Ficus maxima Clarke, H.D.: 610 Ficus paraensis Clarke, H.D.: 683, 1550; Mutchnick, P.: 390, 462 Ficus trigona McDowell, T.: 2089; Mutchnick, P.: 663 Helicostylis tomentosa Mori, S.: Pseudolmedia laevis Hoffman, B.: 4909 Sorocea pubivena Hoffman, B.: 1545 Myristicaceae Iryanthera juruensis Mutchnick, P.: 932 Iryanthera lancifolia Clarke, H.D.: 4177; Mori, S.: Iryanthera macrophylla Clarke, H.D.: 2615 Iryanthera paraensis Mutchnick, P.: 749 Virola surinamensis Acevedo-Rdgz., P.: 3424; Clarke, H.D.: 1562; Hoffman, B.: 4634 Myrsinaceae Ardisia guianensis Clarke, H.D.: 4254; Hoffman, B.: 1277, 1289 Cybianthus fulvopulverulentus Clarke, H.D.: 221, 2662; Hoffman, B.: 4637 Cybianthus microbotrys Clarke, H.D.: 4154; Mori, S.: 24526; Mutchnick, P.: 938, 970 Stylogyne orinocensis Clarke, H.D.: 611 Stylogyne sp. Mutchnick, P.: 756 Myrtaceae Calycorectes bergii Clarke, H.D.: 4121 Calyptranthes pullei Hoffman, B.: 1306; Mutchnick, P.: 564 Calyptranthes sp. Hoffman, B.: 4901 Eugenia albicans Mori, S.: Eugenia anastomosans Clarke, H.D.: 2618 Eugenia arawakorum Mutchnick, P.: 501 Eugenia biflora McDowell, T.: 3261 Eugenia coffeifolia Hoffman, B.: 4508, 5000 Eugenia cribrata Clarke, H.D.: 1384 Eugenia cucullata Clarke, H.D.: 172; Mori, S.: Eugenia egensis Hoffman, B.: 1356, 1360; Mutchnick, P.: 473 Eugenia florida Clarke, H.D.: 267 Eugenia lambertiana Clarke, H.D.: 308, 342, 1381; Hoffman, B.: 1398; Mutchnick, P.: 546 Eugenia latifolia Clarke, H.D.: 4340; Mori, S.: Eugenia ligustrina Acevedo-Rdgz., P.: 3301 Eugenia luciae Acevedo-Rdgz., P.: 3435 Eugenia mimus Clarke, H.D.: 2575; Mutchnick, P.: 944 Eugenia patrisii Acevedo-Rdgz., P.: 3416 Eugenia tafelbergica Hoffman, B.: 1525 Eugenia tapacumensis Clarke, H.D.: 377 Eugenia trinervia Hoffman, B.: 4500 Eugenia sp. Ehringhaus, C.: 67; Hoffman, B.: 4523, 4908, 5015 Myrcia calycampa Acevedo-Rdgz., P.: 3407; Clarke, H.D.: 2761; Ehringhaus, C.: 70; Hoffman, B.: 1318, 1498 Myrcia decorticans Clarke, H.D.: 1390 Myrcia guianensis Hoffman, B.: 4573; McDowell, T.: 2022, 3280 Myrcia inaequiloba Clarke, H.D.: 1537; Mutchnick, P.: 633 Myrcia pyrifolia Mutchnick, P.: 761, 858 Myrcia quitarensis Clarke, H.D.: 692, 1388 Myrcia subobliqua Clarke, H.D.: 476, 597, 1323, 1482; Hoffman, B.: 1282, 1311, 1380, 1416, 1533, 1576, 4620; McDowell, T.: 1941; Mori, S.: 24358; Mutchnick, P.: 771 Myrcia sylvatica Clarke, H.D.: 534, 669, 2659; McDowell, T.: 3257 Myrcia sp. Hoffman, B.: 4530

79 BRIT.ORG/SIDA 73 Myrciaria dubia Clarke, H.D.: 1485 Myrciaria floribunda Clarke, H.D.: 1504 Myrciaria vismeifolia Clarke, H.D.: 479, 1324, 2745; Hoffman, B.: 1309; McDowell, T.: 3267; Mutchnick, P.: 682 Plinia pinnata Clarke, H.D.: 1539 Psidium acutangulum Clarke, H.D.: 284, 293, 295, 478, 2707; Hoffman, B.: 1293; Mutchnick, P.: 464 Psidium guineense McDowell, T.: 1952 Psidium persoonii McDowell, T.: 3277 Psidium striatulum Mutchnick, P.: 452 Nyctaginaceae Neea floribunda Hoffman, B.: 4914 Nymphaeaceae Victoria amazonica Mutchnick, P.: 545 Ochnaceae Ouratea rupununiensis Clarke, H.D.: 364; Hoffman, B.: 1304 Ouratea schomburgkii Mutchnick, P.: 798 Ouratea superba Hoffman, B.: 1512 Ouratea sp. McDowell, T.: 1995 Sauvagesia erecta Clarke, H.D.: 496; Ehringhaus, C.: 108; Hoffman, B.: 1441; Mutchnick, P.: 450 Sauvagesia rubiginosa Acevedo-Rdgz., P.: 3374 Olacaceae Dulacia guianensis Mutchnick, P.: 864 Heisteria cauliflora Clarke, H.D.: 159, 378, 677, 1444, 2688, 2710; Hoffman, B.: 1278; McDowell, T.: 3368; Mori, S.: 24361; Mutchnick, P.: 389, 466 Heisteria densifrons Hoffman, B.: 4549, 4570; McDowell, T.: 2088 Oleandraceae Oleandra articulata Clarke, H.D.: 2560 Onagraceae Ludwigia erecta Hoffman, B.: 1301 Ludwigia leptocarpa Clarke, H.D.: 1354; Mutchnick, P.: 842 Ludwigia rigida Clarke, H.D.: 1406 Ludwigia sericea Mori, S.: Orchidaceae Bifrenaria longicornis Clarke, H.D.: 249 Bollea violacea Clarke, H.D.: 2506, 4284, 4310 Brassavola martiana Clarke, H.D.: 353, 359, 386, 1366 Brassavola sp. Clarke, H.D.: 2760 Brassia lawrenceana Mutchnick, P.: 872 Campylocentrum fasciola Clarke, H.D.: 1513 Campylocentrum micranthum Clarke, H.D.: 1503 Cattleya violacea Clarke, H.D.: 347, 351, 1365; Mutchnick, P.: 714, 719, 812 Caularthron bilamellatum Clarke, H.D.: 384 Cheiradenia cuspidata Clarke, H.D.: 642, 4155, 4240; Mori, S.: Cryptarrhena kegelii Clarke, H.D.: 4175 Dichaea sp. Clarke, H.D.: 2522, 4107, 4285; Ehringhaus, C.: 118, 132; Mutchnick, P.: 839 Dimerandra sp. Clarke, H.D.: 354; Hoffman, B.: 1505 Elleanthus caravata Clarke, H.D.: 4104 Elleanthus sp. Clarke, H.D.: 1440; Mutchnick, P.: 949 Encyclia granitica Mutchnick, P.: 826 Encyclia vespa McDowell, T.: 3344; Mutchnick, P.: 560 Encyclia sp. Clarke, H.D.: 356 Epidendrum flexuosum Mutchnick, P.: 453 Epidendrum macrocarpum Clarke, H.D.: 1545 Epidendrum nocturnum Clarke, H.D.: 1525, 2549, 2672, 2675, 2679 Epidendrum orchidiflorum Clarke, H.D.: 246 Epidendrum purpurascens Clarke, H.D.: 429 Epidendrum ramosum Clarke, H.D.: 651, 4100 Epidendrum rigidum Clarke, H.D.: 1362; Mutchnick, P.: 723 Epidendrum strobiliferum Mutchnick, P.: 819 Habenaria leprieurii Hoffman, B.: 1432 Lockhartia imbricata McDowell, T.: 2094 Masdevallia minuta Mori, S.: Maxillaria brachybulbon Clarke, H.D.: 422b Maxillaria camaridii Hoffman, B.: 1323, 1504 Maxillaria crassifolia Mori, S.: Maxillaria discolor Mori, S.: Maxillaria parkeri Clarke, H.D.: 1420 Maxillaria parviflora Clarke, H.D.: 421 Maxillaria uncata Clarke, H.D.: 422a ; McDowell, T.: 2095; Mutchnick, P.: 815 Maxillaria venezuelana Clarke, H.D.: 2639 Maxillaria sp. Acevedo-Rdgz., P.: 3453; Clarke, H.D.: 4105; Mori, S.: Octomeria sp. Mutchnick, P.: 503 Orleansia amazonica Clarke, H.D.: 360, 385, 395, 1383 Palmorchis sp. Clarke, H.D.: 4147, 4298, 4303; Mori, S.: 24387; Mutchnick, P.: 960, 981 Paphinia sp. Ehringhaus, C.: 119 Pleurothallis lanceana Mutchnick, P.: 606 Polystachya stenophylla Clarke, H.D.: 2550, 2767 Quekettia microscopica Hoffman, B.: 1467; Mutchnick, P.: 404 Quekettia sp. Clarke, H.D.: 1514 Reichenbachanthus reflexus Clarke, H.D.: 4102 Rodriguezia lanceolata Clarke, H.D.: 1307, 1371; Mori, S.: Rudolfiella aurantiaca Clarke, H.D.: 2637 Scaphyglottis sickii Clarke, H.D.: 2676, 4097, 4312; Ehringhaus, C.: 131; Mori, S.: Scaphyglottis sp. Clarke, H.D.: 415 Schomburgkia sp. Mutchnick, P.: 722, 897 Stelis sp. Clarke, H.D.: 589, 4204; Hoffman, B.: 1507 Trigonidium obtusum Mutchnick, P.: 921 Vanilla sp. Clarke, H.D.: 1575 Wullschlaegelia calcarata Clarke, H.D.: 1432, 1454 Xerorchis trichorhiza Clarke, H.D.: 4109 Oxalidaceae Oxalis frutescens Clarke, H.D.: 362, 684, 2759; Hoffman, B.: 1302; McDowell, T.: 3340; Mutchnick, P.: 638, 701 Passifloraceae Passiflora amoena Clarke, H.D.: 410, 602, 1282 Passiflora auriculata Clarke, H.D.: 562 Passiflora capparidifolia Clarke, H.D.: 259, 1359, 1522; Hoffman, B.: 1483; McDowell, T.: 3283

80 74 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana Passiflora cirrhiflora Clarke, H.D.: 4094 Passiflora coccinea Clarke, H.D.: 257, 431, 4191; Ehringhaus, C.: 89; Hoffman, B.: 1345, 1394; McDowell, T.: 3357; Mutchnick, P.: 580, 954 Passiflora fuchsiiflora Mutchnick, P.: 853 Passiflora glandulosa Clarke, H.D.: 209, 518, 1281, 2543; Hoffman, B.: 1334, 1566; Mutchnick, P.: 577, 795 Passiflora laurifolia Clarke, H.D.: 1598 Passiflora vespertilio Clarke, H.D.: 1518; Ehringhaus, C.: 88; McDowell, T.: 3284 Passiflora sp. nov. Hoffman, B.: 1530 Phytolaccaceae Phytolacca rivinoides Hoffman, B.: 1335 Piperaceae Peperomia macrostachya Acevedo-Rdgz., P.: 3415; Clarke, H.D.: 559, 595, 1346, 1494; Mori, S.: 24558; Mutchnick, P.: 385 Peperomia obtusifolia Clarke, H.D.: 4136, 4176 Peperomia ouabianae Clarke, H.D.: 4128, 4306 Peperomia quadrangularis Acevedo-Rdgz., P.: 3305; McDowell, T.: 2096 Peperomia serpens Clarke, H.D.: 4255; Mori, S.: Peperomia sp. Clarke, H.D.: 1437 Piper aequale McDowell, T.: 3315 Piper anonifolium Clarke, H.D.: 598 Piper arboreum Acevedo-Rdgz., P.: 3353; Clarke, H.D.: 2497, 2513; McDowell, T.: 3325; Mutchnick, P.: 478 Piper avellanum Acevedo-Rdgz., P.: 3382; Clarke, H.D.: 541; Mutchnick, P.: 900 Piper bartlingianum Clarke, H.D.: 316, 1289, 1314; Ehringhaus, C.: 52, 71; Hoffman, B.: 1415, 1489 Piper berbicense Clarke, H.D.: 178, 434; McDowell, T.: 3309, 3313, 3324 Piper demeraranum Clarke, H.D.: 1313, 4294 Piper dilatatum McDowell, T.: 1888 Piper emarginata McDowell, T.: 1903 Piper hispidum Acevedo-Rdgz., P.: 3370; Clarke, H.D.: 4283 Piper hostmannianum Clarke, H.D.: 232, 404; Hoffman, B.: 1535; McDowell, T.: 3323; Mutchnick, P.: 491 Piper insipiens Clarke, H.D.: 2489, 4144, 4179 Piper obliquum Clarke, H.D.: 4178 Piper trichoneuron McDowell, T.: 2047 Poaceae Arundinella hispida Hoffman, B.: 1346 Axonopus longispicus Mori, S.: Digitaria horizontalis Mori, S.: Guadua sp. Hoffman, B.: 1389 Ichnanthus leiocarpus Clarke, H.D.: 4274 Ichnanthus nemoralis Clarke, H.D.: 2470 Ichnanthus pallens Acevedo-Rdgz., P.: 3420; Clarke, H.D.: 4249, 4276; McDowell, T.: 3348; Mori, S.: Ichnanthus panicoides Clarke, H.D.: 1301 Ichnanthus tenuis McDowell, T.: 3352 Lasiacis ligulata McDowell, T.: 1889 Olyra latifolia Acevedo-Rdgz., P.: 3322, 3333; Clarke, H.D.: 4349; Hoffman, B.: 1482, 1494, 1543; McDowell, T.: 3353 Olyra longifolia Acevedo-Rdgz., P.: 3384; Clarke, H.D.: 1446; Ehringhaus, C.: 142; McDowell, T.: 1897; Mori, S.: Olyra micrantha Clarke, H.D.: 317, 2490; McDowell, T.: 2105; Mutchnick, P.: 972 Orthoclada laxa McDowell, T.: 3346 Panicum cyanescens Hoffman, B.: 1438 Panicum hylaeicum Hoffman, B.: 1275 Panicum laxum Hoffman, B.: 1522; McDowell, T.: 1977 Panicum micranthum McDowell, T.: 2018 Panicum parvifolium McDowell, T.: 1956 Panicum pilosum Clarke, H.D.: 543, 4245; Hoffman, B.: 1328, 1342; McDowell, T.: 1972; Mutchnick, P.: 681 Panicum polycomum Hoffman, B.: 1439 Panicum rivale Clarke, H.D.: 2552 Panicum rudgei McDowell, T.: 2035 Panicum stoloniferum Clarke, H.D.: 1475 Pariana radiciflora Hoffman, B.: 1393 Paspalum carinatum Hoffman, B.: 1520 Paspalum conjugatum McDowell, T.: 3360 Paspalum convexum McDowell, T.: 1943 Paspalum melanospermum Mori, S.: 24353; Mutchnick, P.: 879a Paspalum pilosum McDowell, T.: 3363 Paspalum plicatulum Hoffman, B.: 1314 Paspalum virgatum McDowell, T.: 3347 Raddiella esenbeckii Hoffman, B.: 1433 Thrasya scandens Clarke, H.D.: 2551 Podostemaceae Marathrum sp. Mutchnick, P.: 699 Mourera fluviatilis Clarke, H.D.: 396, 471; Mutchnick, P.: 557, 558, 700 Polygalaceae Moutabea guianensis Acevedo-Rdgz., P.: 3398; Clarke, H.D.: 392; Hoffman, B.: 4560; McDowell, T.: 2081; Mori, S.: Polygala adenophora Clarke, H.D.: 700; McDowell, T.: 1948 Polygala subtilis McDowell, T.: 2052 Polygala timoutou McDowell, T.: 2019 Securidaca marginata Clarke, H.D.: 400, 2762; Mutchnick, P.: 809 Securidaca paniculata Mutchnick, P.: 459 Polygonaceae Coccoloba ascendens Acevedo-Rdgz., P.: 3362; Clarke, H.D.: 663 Coccoloba densifrons Clarke, H.D.: 1517; Hoffman, B.: 4539 Coccoloba gymnorrhachis Clarke, H.D.: 4232 Coccoloba ovata Clarke, H.D.: 398 Coccoloba padiformis Clarke, H.D.: 469 Polygonum acuminatum Hoffman, B.: 1516; Mutchnick, P.: 896 Symmeria paniculata Clarke, H.D.: 1360, 1374, 1483; Hoffman, B.: 1292, 1362, 1515; McDowell, T.: 1992; Mutchnick, P.: 641, 895 Polypodiaceae Campyloneurum phyllitidis Clarke, H.D.: 4295; Mutchnick, P.: 806

81 BRIT.ORG/SIDA 75 Campyloneurum repens Clarke, H.D.: 1486 Dicranoglossum desvauxii Clarke, H.D.: 4092; Mori, S.: Microgramma fuscopunctata Mori, S.: 24406; Mutchnick, P.: 619 Microgramma lycopodioides Mutchnick, P.: 775 Microgramma persicariifolia Acevedo-Rdgz., P.: 3385 Microgramma reptans Clarke, H.D.: 1509, 4229, 4332; Mutchnick, P.: 788 Microgramma tecta McDowell, T.: 2090 Microgramma thurnii Clarke, H.D.: 183 Niphidium crassifolium Clarke, H.D.: 604, 4205, 4252 Pecluma consimilis Hoffman, B.: 1474; Mutchnick, P.: 504 Pecluma plumula McDowell, T.: 2097; Mutchnick, P.: 816 Pleopeltis percussa Clarke, H.D.: 1492, 4256; Mutchnick, P.: 814 Polypodium dulce Clarke, H.D.: 4325 Polypodium polypodioides Hoffman, B.: 1506; McDowell, T.: 2092 Pontederiaceae Eichhornia diversifolia Clarke, H.D.: 436 Proteaceae Roupala schomburgkii Clarke, H.D.: 2677 Pteridaceae Pteris pungens Clarke, H.D.: 4223; Mori, S.: Quiinaceae Lacunaria crenata Clarke, H.D.: 369 Quiina guianensis Mutchnick, P.: 884, 886 Quiina obovata Hoffman, B.: 1420, 4572, 4848, 4921; Mori, S.: Quiina rhytidopus Clarke, H.D.: 1473; Hoffman, B.: 1558; McDowell, T.: 1994 Ramalinaceae Ramalina anceps Clarke, H.D.: 2674 Rapateaceae Rapatea paludosa Clarke, H.D.: 1411, 2485; Ehringhaus, C.: 123; Hoffman, B.: 1412; McDowell, T.: 3299; Mori, S.: 24482; Mutchnick, P.: 923 Spathanthus unilateralis Ehringhaus, C.: 122; Mutchnick, P.: 738, 924 Rhabdodendraceae Rhabdodendron sylvestre McDowell, T.: 3251 Rhamnaceae Gouania blanchetiana Clarke, H.D.: 1564 Gouania polygama Acevedo-Rdgz., P.: 3363 Gouania velutina Hoffman, B.: 1382 Rhizophoraceae Cassipourea guianensis Clarke, H.D.: 417, 2599, 2601 Cassipourea lasiocalyx Clarke, H.D.: 235 Rubiaceae Alibertia myrciifolia Hoffman, B.: 1563 Amaioua brevidentata Hoffman, B.: 1303 Amaioua guianensis Clarke, H.D.: 175, 336, 1347, 4228 Borreria capitata Clarke, H.D.: 1601 Borreria latifolia Mori, S.: 24351; Mutchnick, P.: 596 Borreria prostrata Mori, S.: Borreria verticillata Clarke, H.D.: 696 Chimarrhis microcarpa Hoffman, B.: 1532 Chomelia monachinoi Clarke, H.D.: 1386 Chomelia tenuiflora Clarke, H.D.: 2747; Hoffman, B.: 5007 Chomelia sp. Mutchnick, P.: 457 Coccocypselum guianense Mori, S.: Coccocypselum herbaceum Acevedo-Rdgz., P.: 3375; Clarke, H.D.: 493 Coccocypselum tontanea Ehringhaus, C.: 91 Coussarea paniculata Hoffman, B.: 1396 Diodia hyssopifolia Hoffman, B.: 1274; McDowell, T.: 3341; Mori, S.: 24344, 24345, 24481; Mutchnick, P.: 549, 597, 639, 640 Diodia ocimifolia Hoffman, B.: 1544 Duroia eriopila Acevedo-Rdgz., P.: 3395; Clarke, H.D.: 189, 193; Hoffman, B.: 1414; Mori, S.: 24490; Mutchnick, P.: 818 Duroia fusifera McDowell, T.: 1970 Duroia genipoides Clarke, H.D.: 1394, 2641 Duroia micrantha Hoffman, B.: 1500; Mori, S.: 24540, Faramea occidentalis Clarke, H.D.: 4161, 4220; Hoffman, B.: 4924, 4927 Faramea sessilifolia Clarke, H.D.: 1471; Ehringhaus, C.: 57; Hoffman, B.: 1502; McDowell, T.: 1967, 1976 Ferdinandusa uaupensis Mori, S.: Genipa americana Mutchnick, P.: 688, 935 Genipa spruceana Clarke, H.D.: 275, 355, 470, 2690; Hoffman, B.: 1273; Mutchnick, P.: 550, 915 Geophila repens Acevedo-Rdgz., P.: 3314; McDowell, T.: 3311a Gonzalagunia dicocca Clarke, H.D.: 613, 2627, 4247; Hoffman, B.: 1481; Mori, S.: 24550; Mutchnick, P.: 479 Guettarda acreana Clarke, H.D.: 337 Guettarda argentea Mori, S.: Hillia illustris Clarke, H.D.: 1554 Isertia hypoleuca Clarke, H.D.: 492, 2648; Ehringhaus, C.: 110; McDowell, T.: 3319; Mori, S.: 24451; Mutchnick, P.: 586 Isertia parviflora Acevedo-Rdgz., P.: 3342; Clarke, H.D.: 1596; Mutchnick, P.: 813 Ixora ferrea Clarke, H.D.: 2537, 2611, 2715; Hoffman, B.: 1469 Ixora graciliflora Mutchnick, P.: 507 Ixora panurensis Clarke, H.D.: 228, 426, 1441 Malanea macrophylla Mori, S.: Mitracarpus hirtus Hoffman, B.: 1300 Morinda calycina Hoffman, B.: 1413, 1569; McDowell, T.: 2048 Morinda tenuiflora Clarke, H.D.: 215, 1484, 1496 Pagamea capitata Clarke, H.D.: 237, 2668; Hoffman, B.: 1445 Palicourea croceoides Clarke, H.D.: 413, 565, 628, 2481, 2482, 4081; Mori, S.: Palicourea guianensis Clarke, H.D.: 325, 581, 1310, 4137; Ehringhaus, C.: 109; Mori, S.: 24455; Mutchnick, P.: 601 Palicourea riparia Clarke, H.D.: 1535; Hoffman, B.: 1510; McDowell, T.: 1966; Mutchnick, P.: 608, 977

82 76 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana Palicourea triphylla Acevedo-Rdgz., P.: 3367; Hoffman, B.: 1333; Mori, S.: Perama galioides McDowell, T.: 2017 Perama hirsuta Hoffman, B.: 1434 Posoqueria longiflora Clarke, H.D.: 1534 Posoqueria panamensis Mutchnick, P.: 523 Psychotria apoda Clarke, H.D.: 484, 1422, 1451, 2519, 4111; Mutchnick, P.: 481 Psychotria astrellantha Hoffman, B.: 1466 Psychotria bahiensis Mutchnick, P.: 901 Psychotria barbiflora Clarke, H.D.: 2540, 4110; McDowell, T.: 3374; Mutchnick, P.: 952 Psychotria brachybotrya Clarke, H.D.: 177, 323 Psychotria bracteocardia Clarke, H.D.: 1447, 1508; Hoffman, B.: 1565; McDowell, T.: 1887, 1896 Psychotria capitata Clarke, H.D.: 2680 Psychotria cupularis Clarke, H.D.: 607, 637, 4160, 4231 Psychotria horizontalis Hoffman, B.: 1534 Psychotria lupulina Clarke, H.D.: 2732; Hoffman, B.: 1279; Mutchnick, P.: 469 Psychotria mapourioides Clarke, H.D.: 2508 Psychotria microbotrys Clarke, H.D.: 4138 Psychotria microdon Ehringhaus, C.: 50 Psychotria officinalis Mori, S.: Psychotria poeppigiana Clarke, H.D.: 614, 2505, 2521, 4253; Mori, S.: Psychotria polycephala Clarke, H.D.: 160, 488, 1452; Ehringhaus, C.: 56; Hoffman, B.: 1337, 1417; McDowell, T.: 3295 Psychotria racemosa Hoffman, B.: 1405 Psychotria uliginosa Clarke, H.D.: 605, 1433, 2523, 4140 Randia armata McDowell, T.: 2098; Mori, S.: Randia calycina Ehringhaus, C.: 54 Retiniphyllum schomburgkii Clarke, H.D.: 2669 Rudgea cornifolia Clarke, H.D.: 202, 1456, 1472; Hoffman, B.: 1280 Rudgea crassiloba Clarke, H.D.: 1326 Rudgea fissistipula Mori, S.: Rudgea hostmanniana Clarke, H.D.: 584, 590, 1431, 2492, 2516; Ehringhaus, C.: 126; Hoffman, B.: 1488, 1567, 4890; Mutchnick, P.: 607 Rudgea stipulacea Clarke, H.D.: 407 Sabicea oblongifolia Hoffman, B.: 1336; Mutchnick, P.: 424 Sabicea villosa Clarke, H.D.: 501; Ehringhaus, C.: 106; McDowell, T.: 3254; Mori, S.: Sipanea pratensis Clarke, H.D.: 1506; Ehringhaus, C.: 60 Sipanea wilson-brownei Clarke, H.D.: 2554, 4158 Spermacoce assurgens McDowell, T.: 3317 Spermacoce verticillata Hoffman, B.: 1523 Uncaria guianensis Clarke, H.D.: 274; Hoffman, B.: 4510; McDowell, T.: 2084 Rutaceae Conchocarpus fanshawei Clarke, H.D.: 402 Ertela trifolia Clarke, H.D.: 1512 Ticorea foetida Clarke, H.D.: 411; Mori, S.: Zanthoxylum compactum Clarke, H.D.: 1540, 2705 Salviniaceae Salvinia auriculata Clarke, H.D.: 1395, 1479 Sapindaceae Allophylus racemosus Acevedo-Rdgz., P.: 3457 Cupania hirsuta Acevedo-Rdgz., P.: 3316; Clarke, H.D.: 203, 334; Hoffman, B.: 5047 Cupania scrobiculata Hoffman, B.: 1298; Mutchnick, P.: 776 Matayba elegans Clarke, H.D.: 387, 1308; McDowell, T.: 3336; Mutchnick, P.: 860 Matayba macrolepis Acevedo-Rdgz., P.: 3346; Clarke, H.D.: 301, 1528, 1543, 2741 Matayba purgans Acevedo-Rdgz., P.: 3410 Matayba sp. Hoffman, B.: 4632 Paullinia ingaefolia Acevedo-Rdgz., P.: 3447, 3448 Paullinia latifolia Clarke, H.D.: 4203; Hoffman, B.: 1383; Mori, S.: 24561; Mutchnick, P.: 947 Paullinia leiocarpa Acevedo-Rdgz., P.: 3455 Paullinia pachycarpa Hoffman, B.: 5064 Paullinia perlata Clarke, H.D.: 616 Paullinia plagioptera Mori, S.: Paullinia rufescens Mori, S.: Paullinia stellata Clarke, H.D.: 4239 Pseudima frutescens Hoffman, B.: 4894 Serjania caracasana Acevedo-Rdgz., P.: 3296 Serjania oblongifolia Acevedo-Rdgz., P.: 3401, 3444 Serjania paniculata Hoffman, B.: 4900 Serjania paucidentata Acevedo-Rdgz., P.: 3361 Talisia carinata Clarke, H.D.: 2712; Mori, S.: Talisia mollis Clarke, H.D.: 4227 Toulicia guianensis Mutchnick, P.: 808 Urvillea ulmacea Acevedo-Rdgz., P.: 3456 Sapotaceae Chrysophyllum sparsiflorum Hoffman, B.: 4568 Ecclinusa guianensis Clarke, H.D.: 196, 213, 529, 1419 Manilkara bidentata Hoffman, B.: 4554 Micropholis venulosa Hoffman, B.: 5050 Pouteria caimito Mori, S.: Pouteria cuspidata Clarke, H.D.: 266; Mori, S.: 24484; Mutchnick, P.: 927 Pouteria hispida Clarke, H.D.: 4292 Pouteria oblanceolata Mori, S.: Pouteria rigidopsis Clarke, H.D.: 2713 Pouteria speciosa Clarke, H.D.: 2718 Pouteria surumuensis Clarke, H.D.: 4162 Pouteria torta Clarke, H.D.: 195, 1312, 4173; Mori, S.: 24560; Mutchnick, P.: 784 Pouteria venosa Clarke, H.D.: 2683; Mutchnick, P.: 451 Pouteria sp. Hoffman, B.: 4501, 4940 Pradosia ptychandra Clarke, H.D.: 331 Pradosia schomburgkiana Clarke, H.D.: 536; Hoffman, B.: 1290 Pradosia surinamensis Clarke, H.D.: 2758 Schizaeaceae Schizaea fluminensis Mori, S.: 24401, Scrophulariaceae Bacopa aquatica McDowell, T.: 3297 Benjaminia reflexa McDowell, T.: 2023 Buchnera rosea Clarke, H.D.: 1578 Conobea aquatica McDowell, T.: 2021 Stemodia pratensis Clarke, H.D.: 507

83 BRIT.ORG/SIDA 77 Selaginellaceae Selaginella parkeri Clarke, H.D.: 258, 315, 676, 1277; McDowell, T.: 3291; Mori, S.: 24402; Mutchnick, P.: 730 Simaroubaceae Picramnia guianensis Mutchnick, P.: 500 Picramnia latifolia Clarke, H.D.: 620, 625, 2539, 2721, 4278b ; Hoffman, B.: 4965; Mutchnick, P.: 937, 957 Picramnia magnifolia Clarke, H.D.: 4123 Simaba cedron Clarke, H.D.: 671; Mori, S.: Simaba guianensis Hoffman, B.: 4871; Mori, S.: Smilacaceae Smilax krukovii Clarke, H.D.: 580 Smilax lasseriana Hoffman, B.: 1472 Smilax latipes Clarke, H.D.: 2528, 4124; Mori, S.: 24590; Mutchnick, P.: 976 Smilax schomburgkiana Clarke, H.D.: 272, 452, 1319; Mutchnick, P.: 443, 471 Smilax syphilitica Clarke, H.D.: 681 Solanaceae Capsicum frutescens McDowell, T.: 1940 Cestrum megalophyllum Clarke, H.D.: 4182, 4221, 4300; Hoffman, B.: 4970 Cestrum racemosum Mutchnick, P.: 630b Lycianthes pauciflora Clarke, H.D.: 4188; Mutchnick, P.: 951 Markea camponoti Clarke, H.D.: 626 Markea longiflora Mori, S.: Markea sessiliflora Clarke, H.D.: 574 Solanum asperum Acevedo-Rdgz., P.: 3307; McDowell, T.: 1957 Solanum coriaceum Chanderbali, A.: 170; Clarke, H.D.: 512, 1408; Ehringhaus, C.: 105; Mori, S.: 24427; Mutchnick, P.: 588 Solanum crinitum Acevedo-Rdgz., P.: 3417; Clarke, H.D.: 1563; Hoffman, B.: 1348; Mutchnick, P.: 583 Solanum leucocarpon Clarke, H.D.: 617; Hoffman, B.: 1341, 1495; Mutchnick, P.: 587 Solanum monachophyllum Clarke, H.D.: 475, 1385, 2692; Hoffman, B.: 1294, 1327; McDowell, T.: 1990; Mutchnick, P.: 472, 646 Solanum pensile Clarke, H.D.: 288, 1552, 4235; McDowell, T.: 3338; Mutchnick, P.: 857 Solanum schlechtendalianum Clarke, H.D.: 624, 4185, 4299 Solanum stramoniifolium Acevedo-Rdgz., P.: 3352; Clarke, H.D.: 1586; McDowell, T.: 3316 Sterculiaceae Byttneria divaricata Acevedo-Rdgz., P.: 3310; Clarke, H.D.: 2751; Hoffman, B.: 1375; Mutchnick, P.: 625, 841 Herrania lemniscata Clarke, H.D.: 1555; Hoffman, B.: 4920; Mori, S.: Melochia arenosa McDowell, T.: 1991; Mutchnick, P.: 667 Sterculia guianensis Hoffman, B.: 1462 Sterculia rugosa Clarke, H.D.: 4134 Sterculia villifera Hoffman, B.: 1477 Waltheria indica Clarke, H.D.: 508, 1589; McDowell, T.: 1944 Strelitziaceae Phenakospermum guyannense Acevedo-Rdgz., P.: 3357; Clarke, H.D.: 1424 Tectariaceae Cyclopeltis semicordata Hoffman, B.: 1575 Tectaria heracleifolia Clarke, H.D.: 4180 Tectaria plantaginea Clarke, H.D.: 2503, 4156; Hoffman, B.: 1476; Mori, S.: 24545, Tectaria trifoliata Clarke, H.D.: 4261; Mori, S.: 24644; Mutchnick, P.: 968 Triplophyllum funestrum Mutchnick, P.: 971 Triplophyllum funestum Clarke, H.D.: 4117 Theaceae Ternstroemia punctata Clarke, H.D.: 242 Thelypteridaceae Thelypteris opulenta Clarke, H.D.: 4200 Thelypteris pennata Clarke, H.D.: 4222 Thelypteris poiteana McDowell, T.: 2102 Thelypteris tetragona Hoffman, B.: 1408 Theophrastaceae Clavija lancifolia Acevedo-Rdgz., P.: 3303; Clarke, H.D.: 1572, 4230; Hoffman, B.: 1571 Thurniaceae Thurnia sphaerocephala Clarke, H.D.: 201; Mori, S.: Thymelaeaceae Daphnopsis longifolia Mori, S.: 24537; Mutchnick, P.: 956 Goodallia guianensis Mutchnick, P.: 709, 712 Tiliaceae Apeiba albiflora Hoffman, B.: 1496 Apeiba petoumo Clarke, H.D.: 406 Corchorus hirtus Hoffman, B.: 1521 Corchorus orinocensis Ehringhaus, C.: 58; McDowell, T.: 1983 Vasivaea alchorneoides Clarke, H.D.: 1391; Hoffman, B.: 1286; Mutchnick, P.: 859 Trigoniaceae Trigonia hypoleuca Clarke, H.D.: 310; Hoffman, B.: 1355 Trigonia laevis Clarke, H.D.: 184, 573, 662; Mori, S.: Trigonia nivea Mutchnick, P.: 703 Trigonia subcymosa Mutchnick, P.: 531 Trigonia villosa Clarke, H.D.: 383; McDowell, T.: 3278, 3327; Mutchnick, P.: 622, 662 Turneraceae Piriqueta cistoides McDowell, T.: 1959; Mutchnick, P.: 702 Turnera aurantiaca Clarke, H.D.: 358, 1363, 2756; Ehringhaus, C.: 70a ; McDowell, T.: 3279; Mutchnick, P.: 599, 660, 661, 698 Turnera curassavica Mutchnick, P.: 630a Turnera rupestris Clarke, H.D.: 428; Hoffman, B.: 1471, 4763; Mutchnick, P.: 493 Turnera scabra Clarke, H.D.: 1591 Ulmaceae Celtis schippii Hoffman, B.: 4504 Trema micrantha Clarke, H.D.: 174; Hoffman, B.: 1325; McDowell, T.: 1894

84 78 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana Urticaceae Urera caracasana Clarke, H.D.: 593 Verbenaceae Aegiphila membranacea Clarke, H.D.: 4183, 4189, 4242; Mori, S.: Aegiphila villosa Clarke, H.D.: 664 Amasonia campestris Clarke, H.D.: 1593 Citharexylum macrophyllum Clarke, H.D.: 4331 Lantana camara McDowell, T.: 3365; Mutchnick, P.: 909 Petrea bracteata Clarke, H.D.: 2565 Petrea macrostachya Acevedo-Rdgz., P.: 3348; Clarke, H.D.: 208, 549, 1318, 1345, 1462, 2624, 4080; Mori, S.: 24504, 24569; Mutchnick, P.: 741 Petrea volubilis McDowell, T.: 3298 Stachytarpheta cayennensis Clarke, H.D.: 1576; McDowell, T.: 2054, 3361 Vitex capitata McDowell, T.: 2038 Vitex stahelii Clarke, H.D.: 1569; Hoffman, B.: 1546 Violaceae Amphirrhox longifolia Acevedo-Rdgz., P.: 3335; Clarke, H.D.: 158, 1491 Corynostylis arborea Clarke, H.D.: 380; Mutchnick, P.: 397 Noisettia orchidiflora Clarke, H.D.: 156 Paypayrola confertiflora Clarke, H.D.: 423 Paypayrola guianensis Mutchnick, P.: 484 Paypayrola longifolia Chanderbali, A.: 155; Clarke, H.D.: 199, 324, 367, 490, 517, 1290, 1338; Hoffman, B.: 4584; Mutchnick, P.: 727, 793, 934 Paypayrola sp. Clarke, H.D.: 418 Rinorea flavescens Clarke, H.D.: 1285; Mutchnick, P.: 740 Rinorea lindeniana Hoffman, B.: 1531 Rinorea macrocarpa Ehringhaus, C.: 53; Hoffman, B.: 4602, 4925; Mori, S.: 24389; Mutchnick, P.: 729 Rinorea pubiflora Acevedo-Rdgz., P.: 3445; Clarke, H.D.: 164, 314, 332, 588, 596, 1467, 1510, 2602, 2625, 4250; Hoffman, B.: 1281, 1485, 1538, 4503, 4893; McDowell, T.: 3370; Mori, S.: 24571; Mutchnick, P.: 382, 745 Rinorea riana Clarke, H.D.: 204, 1339, 4151; Hoffman, B.: 1399, 1464; Mutchnick, P.: 505 Viscaceae Phoradendron crassifolium Mori, S.: Phoradendron perrottetii Acevedo-Rdgz., P.: 3403 Phoradendron piperoides Clarke, H.D.: 1410, 4166 Vitaceae Cissus erosa Hoffman, B.: 1343; McDowell, T.: 3285; Mutchnick, P.: 624, 893 Cissus sicyoides Clarke, H.D.: 465; Mutchnick, P.: 642 Cissus subrhomboidea Clarke, H.D.: 300 Cissus verticillata Clarke, H.D.: 283, 1553; Mutchnick, P.: 534 Vittariaceae Anetium citrifolium Clarke, H.D.: 4234 Antrophyum guayanense Mori, S.: Antrophyum jenmanii Clarke, H.D.: 2632 Vittaria costata McDowell, T.: 2093 Vittaria graminifolia Clarke, H.D.: 2535 Vochysiaceae Vochysia surinamensis Clarke, H.D.: 2643 Vochysia tetraphylla Clarke, H.D.: 291, 439; McDowell, T.: 1989, 3381; Mutchnick, P.: 407, 865 Xyridaceae Abolboda grandis Clarke, H.D.: 225, 248, 2661 Xyris fallax Hoffman, B.: 1436 Xyris involucrata Clarke, H.D.: 245 Xyris jupicai Clarke, H.D.: 498, 1402; Ehringhaus, C.: 92; Mori, S.: Xyris surinamensis Clarke, H.D.: 239 Zingiberaceae Renealmia floribunda Clarke, H.D.: 211, 1341 Renealmia monosperma Clarke, H.D.: 4272 Renealmia orinocensis Clarke, H.D.: 644, 2488; Ehringhaus, C.: 140; McDowell, T.: 3300; Mutchnick, P.: 621, 974 Renealmia thyrsoidea Clarke, H.D.: 4145

85 BRIT.ORG/SIDA 79 APPENDIX 3 LIST OF RANKS OF SPECIES RICHNESS BY FAMILY FOR IWOKRAMA, FOUR OTHER SITES IN NE S. AMERICA, AND THE GUIANAS OVERALL Family Iwokrama I_rank Kaieteur K_rank Mabura M_rank CFG C_rank Ducke D_rank Guianas G_rank Fabaceae Rubiaceae Orchidaceae Cyperaceae Myrtaceae Melastomataceae Poaceae Lauraceae Annonaceae Araceae Chrysobalanaceae Clusiaceae Euphorbiaceae Sapindaceae Bignoniaceae Asteraceae Apocynaceae Malpighiaceae Arecaceae Piperaceae Sapotaceae Solanaceae Lecythidaceae Polypodiaceae Boraginaceae Adiantaceae Marantaceae Moraceae Verbenaceae Violaceae Burseraceae Convolvulaceae Hippocrateaceae Hymenophyllaceae Passifloraceae

86 80 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana APPENDIX 3 (continued) Family Iwokrama I_rank Kaieteur K_rank Mabura M_rank CFG C_rank Ducke D_rank Guianas G_rank Gentianaceae Malvaceae Meliaceae Gesneriaceae Bromeliaceae Dennstaedtiaceae Flacourtiaceae Heliconiaceae Polygonaceae Sterculiaceae Cucurbitaceae Cyatheaceae Dilleniaceae Polygalaceae Acanthaceae Caryocaraceae Combretaceae Elaeocarpaceae Menispermaceae Monimiaceae Myristicaceae Ochnaceae Scrophulariaceae Simaroubaceae Smilacaceae Tectariaceae Tiliaceae Trigoniaceae Turneraceae Vittariaceae Xyridaceae Aspleniaceae Cecropiaceae Dioscoreaceae Loganiaceae Loranthaceae Onagraceae Quiinaceae

87 BRIT.ORG/SIDA 81 Family Iwokrama I_rank Kaieteur K_rank Mabura M_rank CFG C_rank Ducke D_rank Guianas G_rank Rutaceae Thelypteridaceae Zingiberaceae Bombacaceae Burmanniaceae Connaraceae Eriocaulaceae Gnetaceae Grammitidaceae Humiriaceae Lamiaceae Lentibulariaceae Marcgraviaceae Myrsinaceae Olacaceae Rhamnaceae Ulmaceae Viscaceae Vitaceae Anacardiaceae Aristolochiaceae Asclepiadaceae Begoniaceae Bixaceae Capparaceae Celastraceae Commelinaceae Costaceae Cyclanthaceae Dichapetalaceae Dryopteridaceae Ebenaceae Liliaceae Lomariopsidaceae Lycopodiaceae Lygodiaceae Marattiaceae Mendonciaceae Podostemaceae

88 82 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana APPENDIX 3 (continued) Family Iwokrama I_rank Kaieteur K_rank Mabura M_rank CFG C_rank Ducke D_rank Guianas G_rank Rapateaceae Rhizophoraceae Thymelaeaceae Vochysiaceae Balanophoraceae Cactaceae Campanulaceae Caricaceae Droseraceae Erythroxylaceae Gleicheniaceae Haemodoraceae Icacinaceae Lacistemataceae Lythraceae Metaxyaceae Nyctaginaceae Oleandraceae Oxalidaceae Phytolaccaceae Pontederiaceae Proteaceae Pteridaceae Rhabdodendraceae Schizaeaceae Selaginellaceae Strelitziaceae Theaceae Theophrastaceae Thurniaceae Urticaceae Alismataceae Amaranthaceae Anisophylleaceae Apiaceae Aquifoliaceae Araliaceae Blechnaceae

89 BRIT.ORG/SIDA 83 Family Iwokrama I_rank Kaieteur K_rank Mabura M_rank CFG C_rank Ducke D_rank Guianas G_rank Bonnetiaceae Canellaceae Caryophyllaceae Cyrillaceae Eremolepidaceae Ericaceae Hernandiaceae Hugoniaceae Iridaceae Ixonanthaceae Lissocarpaceae Mayacaceae Nymphaeaceae Ophioglossaceae Opiliaceae Peridiscaceae Portulacaceae Rafflesiaceae Rosaceae Sabiaceae Styracaceae Symplocaceae Triuridaceae Woodsiaceae

90 84 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana APPENDIX 4 SPECIALISTS CONTRIBUTING DETERMINATIONS OF IWOKRAMA SPECIMENS Family Determiner(s) Acanthaceae D. Wasshausen (US) Adiantaceae D.B. Lellinger (US) Anacardiaceae J.D. Mitchell (NY) Annonaceae P.J.M. Maas (U) Apocynaceae L. Allorge (P) and A.J.M. Leeuwenberg (WAG) Araceae D.H. Nicolson (US) Araceae T.B. Croat (MO) Arecaceae A. Henderson (NY) Arecaceae F.W. Stauffer (VEN) Aristolochiaceae C. Feuillet (US) Asclepiadaceae G. Morillo (MERF) Asclepiadaceae J.C. Lindeman (U) Aspleniaceae G. McKee and D.B. Lellinger (US) Asteraceae V.A. Funk (US) Balanophoraceae H.D. Clarke (US) Begoniaceae D. Wasshausen (US) and J.C. Lindeman (U) Bignoniaceae A. Gentry (MO) and BDG Bixaceae H.D. Clarke (US) and R.L. Liesner (MO) Bombacaceae L.J. Dorr (US) Boraginaceae C. Feuillet (US) and J.S. Miller (MO) Bromeliaceae B.K.Holst (SEL) and E.J. Gouda (U) Burmanniaceae C.L. Kelloff (US) and P.J.M. Maas (U) Burseraceae D.Daly (NY) Cactaceae H.D. Clarke (US) and B.E. Leuenberger (B) Campanulaceae H.D. Clarke (US) Cannaceae P.J.M. Maas (U) Capparaceae J.C. Lindeman (U) Capparaceae M. Nee (NY) Caricaceae H.D. Clarke (US) and R.L. Liesner (MO) Caryocaraceae G.T. Prance (K) Cecropiaceae C.C. Berg (BG) Celastraceae A.R.A. Görts-van Rijn (U) Chrysobalanaceae G.T. Prance (K) Clusiaceae J.J. Pipoly (BRIT) Combretaceae C.A. Stace (LTR) Commelinaceae R.B. Faden (US) Connaraceae R.L. Liesner (MO) Connaraceae G. Aymard (PORT) Convolvulaceae D.F.Austin (FAU) Costaceae P.J.M. Maas (U) Cucurbitaceae M.Nee (NY) Cyatheaceae D.B. Lellinger and G. McKee (US) Cyclanthaceae R.L. Liesner (MO) Cyperaceae M.T. Strong (MO) Dennstaedtiaceae D.B. Lellinger and G. McKee (US) Dichapetalaceae G.T. Prance (K) Dilleniaceae C.L. Kelloff (US) and G. Aymard (PORT) Dioscoreaceae C.L. Kelloff (US) Droseraceae J.C. Lindeman (U) Dryopteridaceae D.B. Lellinger and G. McKee (US) Ebenaceae P. Berry (WIS) Elaeocarpaceae H.D. Clarke and C.L. Kelloff (US) Eriocaulaceae BDG Erythroxylaceae P. Berry (WIS) Euphorbiaceae L.J. Gillespie (CAN), R.L. Liesner (MO), and W. Punt (U) Fabaceae R.C. Barneby (NY), H.D. Clarke (US), R.L.Liesner (MO), and O. Poncy (P) Flacourtiaceae H.D. Clarke (US), R.L. Liesner (MO), and J.C. Lindeman (U) Gentianaceae C.L.Kelloff (US) and P.J.M. Maas (U) Gesneriaceae C. Feuillet and L.E. Skog (US) Gleicheniaceae G. McKee (US)

91 BRIT.ORG/SIDA 85 Family Gnetaceae Grammitidaceae Haemodoraceae Heliconiaceae Hippocrateaceae Humiriaceae Hymenophyllaceae Icacinaceae Lacistemataceae Lamiaceae Lauraceae Lecythidaceae Lentibulariaceae Liliaceae Lissocarpaceae Loganiaceae Lomariopsidaceae Loranthaceae Lycopodiaceae Lygodiaceae Lythraceae Malpighiaceae Malvaceae Marantaceae Marattiaceae Marcgraviaceae Melastomataceae Meliaceae Mendonciaceae Menispermaceae Metaxyaceae Monimiaceae Moraceae Myristicaceae Myrsinaceae Myrtaceae Nyctaginaceae Nymphaeaceae Ochnaceae Olacaceae Oleandraceae Onagraceae Orchidaceae Oxalidaceae Passifloraceae Phytolaccaceae Piperaceae Poaceae Podostemaceae Polygalaceae Polygonaceae Polypodiaceae Pontederiaceae Proteaceae Pteridaceae Quiinaceae Rapateaceae Rhabdodendraceae Rhamnaceae Rhizophoraceae Rubiaceae Rutaceae Determiner(s) BDG G. McKee (US) BDG W.J. Kress (US) and P.J.M. Maas (U) A.R.A. Görts-van Rijn and A.M. Mennega (U) H.D. Clarke (US), O. Huber (VEN), and R.L. Liesner (MO) D.B. Lellinger and G. McKee (US) A.C. deroon (U) R.L. Liesner (MO) R.M. Harley (K) A. Chanderbali and H. van der Werff (MO) S.A. Mori (NY) P.Taylor (K) BDG P. Berry (WIS) C.L. Kelloff (US) and R. L. Liesner (MO) G. McKee (US) J.W. Kuijt (UVIC) D.B. Lellinger and G. McKee (US) D.B. Lellinger and G. McKee (US) A. Lourteig (P) W.R. Anderson (MICH) L.J. Dorr (US) H. Kennedy (UBC) D.B. Lellinger and G. McKee (US) T. Hollowell (US) T. Hollowell and J.J. Wurdack (US) T.D. Pennington (K) D. Wasshausen (US) P. Hiepko (B) D.B. Lellinger and G. McKee (US) S.S. Renner (MO) C.C. Berg (BG) BDG J.J. Pipoly (BRIT) B.K. Holst (SEL) G. Aymard (VEN) T. Hollowell (US) C. Sastre (P) H.D. Clarke (US) and R.L. Liesner (MO) G. McKee (US) E. Zardini (MO) E.A. Christenson A. Lourteig (P) C. Feuillet (US) C.L. Kelloff (US) A.R.A. Görts-van Rijn (U) E.J. Judziewicz (WIS) H.D. Clarke (US) A. Jacobs-Brouwer (U) J. Brandbyge (AAU) D.B. Lellinger and G. McKee (US) C.N. Horn (NEWB) J.C. Lindeman (U) G. McKee (US) C.L. Kelloff (US) and R.L. Liesner (MO) P. Berry (WIS) A. Goldberg (US) H.D. Clarke (US) and J.C. Lindeman (U) H.D. Clarke (US) C.M. Taylor (MO) J. Kallunki (NY)

92 86 CLARKE ET AL., Plant diversity of the Iwokrama Forest, Guyana Family Salviniaceae Sapindaceae Sapotaceae Schizaeaceae Scrophulariaceae Selaginellaceae Simaroubaceae Smilacaceae Solanaceae Sterculiaceae Strelitziaceae Tectariaceae Theaceae Thelypteridaceae Theophrastaceae Thurniaceae Thymelaeaceae Tiliaceae Trigoniaceae Turneraceae Ulmaceae Urticaceae Verbenaceae Violaceae Viscaceae Vitaceae Vittariaceae Vochysiaceae Xyridaceae Zingiberaceae Determiner(s) G. McKee (US) P. Acevedo-Rdgz. (US) T.D. Pennington (K) C.L.Kelloff (US) H.D. Clarke (US) and N.H. Holmgren (NY) D.B. Lellinger and G. McKee (US) R.L. Liesner (MO) and W. Thomas (NY) R.L. Liesner (MO) M. Nee (NY) L.J. Dorr (US) P.J.M. Maas (U) D.B. Lellinger and G. McKee (US) A. Weitzman (US) D.B. Lellinger and G. McKee (US) J.C. Lindeman (U) M.T. Strong (US) A. Goldberg (US) and R.L. Liesner (MO) L.J. Dorr (US) C.L. Kelloff (US) R.L. Liesner (MO) H.D. Clarke (US) H.D. Clarke (US) M.J. Jansen-Jacobs (U) H.D. Clarke (US) and W.H.A. Hekking (CM) J. Kuijt (UVIC) H.D. Clarke (US) and R.L. Liesner (MO) D.B. Lellinger and G. McKee (US) C.L. Kelloff (US) R. Kral (BRIT) P.J.M. Maas (U)

93 PLANT DIVERSITY OF THE IWOKRAMA FOREST,GUYANA USING CHECKLISTS AND COLLECTING DATA TO INVESTIGATE PLANT DIVERSITY. I:A COMPARATIVE CHECKLIST OF THE PLANT DIVERSITY OF THE IWOKRAMA FOREST,GUYANA. Plant Diversity of the Iwokrama Forest, Guyana presents the results of floristic research conducted as part of the collection of baseline data for this 360,000 hectare area in central Guyana, an area that has been set aside for rainforest conservation and sustainable development. This work provides insight into the plant diversity of a pristine and previously undocumented lowland rainforest of the Guiana Shield. Extensive field work was carried out to make the plant collections that provide the basis for this flora and are detailed along with discussion of ecology and habitats, regional context, and comparison with four other recent floristic works of areas of comparable size on and adjacent to the Guiana Shield in northeastern South America. Appendices present lists of identified specimens, specialists that provided determinations for the various families, a synonymized checklist of the 1251 species of plants known from Iwokrama combined with those of the four other recent florulas, and a ranking of species diversity by family for these five areas. The combined checklist of 4659 species indicates that there is considerable floristic heterogeneity among even adjacent sites and will prove useful to botanists, conservation biologists, ecologists, and planners working in northeastern South America. H.D. Clarke is Assistant Professor of Botany at the University of North Carolina at Asheville. V.A. Funk is Research Scientist and Curator at the Smithsonian Institution. T. Hollowell is Data Manager, Biological Diversity of the Guianas Program at the Smithsonian Institution. BRIT BOTANICAL RESEARCH INSTITUTE OF TEXAS CONSERVATOR OF OUR BOTANICAL HERITAGE COVER DESIGN BY LINNY HEAGY ISBN