Mclaughlin check list anomura 2010Paguridae (2)

coenobitids, 5 is reduced and carried dorsally under the carapace. The six parapagurids and four pylochelids. That none of pleon is symmetrical, with well calcified tergites, weakly these investigations were able to recover a monophyletic bent under the cephalothorax; the second tergite clearly Paguroidea should not come as surprising. The monophyly visible in dorsal view. the authors refer to is that presented by McLaughlin (1983b) as a challenge to the polyphyly proposed by MacDonald et Adult males have paired first and second pleopods modified as al. (1957) based on their limited larval data. Both claims are gonopods, pleopods 3–5 and uropods are vestigial or entirely archaic and should be relegated to their places in the history of lacking; females have a pair of pleopods developed on each anomuran historical “side-steps.” Knowledge and understand of pleonal segments 2–5; the terminal pleomere has a pair of of morphological and developmental processes have been sexually dimorphically uropods. The telson is reduced to a advanced considerably in the ensuing quarter century. And small calcified plate in both sexes. One apomorphy unique to more recently our “toolbox” has been enhanced by molecular Lomis hirta (Lamarck, 1818) is this dimorphic development of methodologies. Unfortunately, the advancements on both uropods — well developed and elongate in females, vestigial fronts have not been incorporated into the analytical processes in males (McLaughlin et al., 2007). employed by either molecular or morphological practitioners. For example, Ahyong, et al. (2009) found their resulting Development. – Cormbie (1993) and McLaughlin et al. polyphyly of asymmetrical paguroids “difficult to reconcile (2004) provided limited information about development in with somatic morphology. ‘A priori’, the suite of associated the Lomisoidea. Lomis hirta larvae hatch in an advanced modifications required for gastropod shell habitation, present state, with the eyes at least partially stalked and buds in all asymmetrical hermit crabs is compelling evidence of pereopods 1–4 or 1–5 already present. There is some for monophyly.” Perhaps recognition that not all paguroids additional disagreement between Cormie’s (1993) account are asymmetrical, gastropod shell-inhabitants and that the 9 McLaughlin et al.: Checklist of world Lithodoidea, Lomisoidea and Paguroidea and that of McLaughlin et al. (2004) with the former author from either the right or the left gonopore or from both, or finding only a four-segmented pleon and poorly developed one or the other gonopore may be masked by a tuft of stiff pleopod buds. McLaughlin et al. found the pleon to consist setae. Adult females most commonly are provided with a of five pleomeres with fused telson but with the upcoming pair of gonopores on the coxae of the third pereopods, but separation already indicated in some specimens; pleopods occasionally only a single left gonopore is present. Females were moderately well developed and weakly biramous. rarely, if ever, are also provided with male gonopores. Males McLaughlin et al. were of the opinion that even if errors of certain genera develop paired first and second pleopods in observation occurred between the two reports, interstage modified as gonopods, or occasionally only second pleopods morphological variation was likely to occur in L. hirta as so modified. Males also may be provided with unpaired in other anomurans. pleopods on pleomeres 2–5 or reduced combinations thereof, usually occurring on the left side of the pleon, but occasionally Current status. – The phylogenetic relationship of the on either side; in certain genera, male pleopods are totally Lomisoidea with other major anomuran taxa is still lacking. Females not infrequently are provided paired and unresolved. modified first pleopods, occasionally asymmetrically paired second pleopods. Unpaired female pleopods usually occur PAGUROIDEA only on the left side of the pleon on pleomere 2–5, but in certain genera may develop on either side. Unpaired pleopods General morphology. – The dorsal surface of the are never entirely lost, but their number may decrease to 2 cephalothorax is covered by a chitinized to moderately or 3. well calcified carapace; the anterior portion of this carapace, the shield, is partially to completely delineated laterally The pleon terminates in paired symmetrical or asymmetrical, and posterolaterally by the cervical groove and posteriorly biramous uropods, almost always modified by propodal rasps. by the linea transversalis; near the anterolateral margin the The sixth pleonal somite and its uropodal appendages are linea anomurica is usually apparent. The dorsal surfaces most frequently contiguous with the telson; however, in a of the shield and posterior carapace may be pitted, weakly few species of two pagurid genera, the telson is separated grooved, spinulose or tuberculate, but not divided into regions from the sixth somite by a lobe of tissue. The reason for this corresponding to the internal organs of the cephalothorax. peculiarity has not yet been determined. A rostrum may or may not be developed, but rarely is it prominently produced; an intercalary rostral process is Development. – Larval development in the majority of occasionally developed; the anterior margin of the shield paguroids is similar to that of other anomurans in that may or may not show development of lateral projections. the developing embryos hatch from the parental eggs as The ocular peduncles are three-segmented, with the basal swimming larvae (zoeae) provided with carapace, rostrum segments fused in the midline and occasionally produced into and cephalothoracic appendages that include eyes, antennules, spinose projections; no ocular orbits are developed; corneas antennae, mandibles, maxillules, maxillae, and first and are well developed, reduced or rarely absent; ocular acicles second maxillipeds. The elongate pleon typically is composed are present. The antennular peduncles are innermost and of five pleomeres and a telson to which a sixth is added in a three-segmented; the antennal peduncles each are provided subsequent zoeal stage as is the third maxilliped, pereopodal with a supernumerary segment. Gills are bi- or quadriserial and pleopodal buds and uropods. Although four zoeal stages phylobranchiae, varying in number from 14 to eight pairs; are common before the molt to megalopa, this number may epipods occasionally are present. be reduced to one or two or extended to several. Rarely has direct development been documented. Chelipeds are equal, subequal or grossly unequal. Pereopods 2 and 3 are developed as ambulatory legs; pereopod 4 is reduced Developmental “clocks” are variable and phase predictability in size and modified, usually with a propodal rasp. Pereopod is challenging. Unlike brachyuran larvae where setal 5 is also reduced, usually provided with a terminal rasp, and variability is rare, paguroid setal variability is the norm. often carried within the carcinoecium. The paguroid pleons Despite the general developmental patterns paguroids share, may be elongate, straight or flexed, bulbous, or reduced; familial and/or generic specializations make zoeal, megalopal, entirely membranous, provided with calcified tergites on all and early juvenile stages potentially informative from six pleomeres, or with calcification reduced but not entirely evolutionary standpoints but caution is required. Development absent. Internally, space in the paguroid pleon is occupied by in one species is not necessarily going to be the same for the male or female reproductive organs and the cecal lobes potentially allied species. For example, the significant of the midgut gland (hepatopancreas). differences observed in the first zoeal stages of four pylochelid species (two species of Trizocheles Forest, 1987, one each Adult males may be provided with a pair of gonopores on of Pylocheles A. Milne-Edwards, 1880, and Pomatocheles the coxae of the fifth pereopods or only a single gonopore Miers 1879) substantially strengthened Lemaitre et al.’s may be present; rarely, males also may exhibit, presumably (2009) proposition of three distinct evolutionary pathways nonfunctional, female gonopores on the coxae of the third within the Pylochelidae. Clear differences also exist between pereopods. In addition to the male gonopores, a sexual diogenid and pagurid larval developmental patterns and tube of variable length and complexity may be extruded even among species of the polyphyletic genus Pagurus. Unfortunately, interest in the information available from 10 THE RAFFLES BULLETIN OF ZOOLOGY 2010 larval studies has waned, thus another phylogenetic resource Oedignathus Benedict, 1895 is being underutilized. = Oedignathus Benedict, 1895 Current status. – Most evolutionary hypotheses regarding the Paguroidea have focused on the presumed derivation of (type species Oedignathus gilli Benedict, 1895, lithodids from a Pagurus-like ancestor, or the more broadly explored the role of carcinization in paguroid evolution. by monotypy; gender masculine) {4} Only relatively recently have molecular techniques been applied and, as previously alluded to, with mixed results. Oedignathus inermis (Stimpson, 1860) [Hapalogaster] Relationships suggested by molecules often have been in direct conflict with those supported by morphological or = Hapalogaster brandtii Schalfeew, 1892 developmental data, or have surprised phylogenetists with information known from morphology for decades. However, = Oedignathus gilli Benedict, 1895 one recent exception is striking. Albeit limited in scope, developmental and morphological data for four species of the Placetron Schalfeew, 1892 paguroid family Pylochelidae are in accord with molecular analyses of three of the four same species. All three sources = Placetron Schalfeew, 1892 (type species Placetron confirm a broad separation of the subfamilies Pylochelinae and Trizochelinae. wosnessenskii Schalfeew, 1892, by monotypy; A different application of molecular methodology has gender masculine) provided the first thorough and in-depth review of speciation in a genus of the Paguroidea. Malay & Paulay (2010) = Lepeopus Benedict 1895 (type species Lepeopus constructed mitochondrial and nuclear gene phylogenies of the majority of species of Calcinus using replicates forcipatus Benedict, 1895, by monotypy; gender from multiple locations spanning the known ranges of the species. Their study demonstrated the practical usefulness masculine) {5} of sequence data in examining speciation, diversification and even the evolution of coloration. It can only be hoped Placetron wosnessenskii Schalfeew, 1892 that their technique will be applied to other elements of the paguroid puzzle. = Lepeopus forcipatus Benedict, 1895 CHECKLIST NOTES SUPERFAMILY LITHODOIDEA {1} McLaughlin et al. (2007) removed the family SAMOUELLE, 1819 {1, 2} Lithodidae from the superfamily Paguroidea and elevated the family to the rank of superfamily with Family Hapalogastridae Brandt, 1850 {1} families Hapalogastridae and Lithodidae. Acantholithodes Holmes, 1895 {2} {2} The status of Acantholithodes is controversial. In his original description of Dermaturus hispidus, Stimpson = Acantholithodes Holmes, 1895 (1860) commented that it might be necessary to separate it [the species] generically from Dermaturus, (type species Dermaturus hispidus Stimpson, 1860, but he had not seen the typical species of Brandt’s (1850) genus. Holmes (1895) established the genus by original designation; gender masculine) Acantholithodes for Stimpson’s (1860) D. hispidus, differentiating it from Dermaturus by presence of Acantholithodes hispidus (Stimpson, 1860) spines on the body and legs and by the large, spiny prolongation of the outer side of the second segment of [Dermaturus] the antenna, characters entirely lacking in Dermaturus. Although A. hispidus rarely is encountered, the genus Dermaturus Brandt, 1850 {3} has been recognized as a valid Northeastern Pacific taxon since its original description (e.g., Rathbun, = Dermaturus Brandt, 1850 (type species Dermaturus 1904; Schmitt, 1921; Hart, 1982; Williams et al., 1989; McLaughlin et al., 2005). mandtii Brandt, 1850, by monotypy; gender {3} Dawson & Yaldwyn (1985) and Dawson (1989) masculine) considered Acantholithodes a synonym of Dermaturus and apparently that opinion was shared by Macpherson Dermaturus mandtii Brandt, 1850 (1988) who listed only Dermaturus among the four genera of the subfamily Hapalogastrinae. Additionally, Hapalogaster Brandt, 1850 the date of publication given by De Grave et al. (2009) of 1894 is incorrect. Holmes’ paper was published in = Hapalogaster Brandt, 1850 1895. (type species Hapalogaster mertensii Brandt, 1850, {4} Makarov (1938) and Dawson (1989) both specifically cited Oedignathus inermis as the type species. by monotype; gender masculine) However, in his original description of the genus, Benedict (1895) included only his own new species Hapalogaster cavicauda Stimpson, 1859 O. gilli. Thus O. gilli is the type species of the genus Oedignathus by monotypy, despite it being a junior Hapalogaster dentata (De Haan, 1849) [Lomis] synonym of O. inermis. Hapalogaster grebnitzkii Schalfeew, 1892 Hapalogaster mertensii Brandt, 1850 11 McLaughlin et al.: Checklist of world Lithodoidea, Lomisoidea and Paguroidea Family Lithodidae Samouelle, 1819 {1, 2} = Lithodes Maia Samouelle, 1819 (misspelling of maja) Cryptolithodes Brandt, 1848 = Lithodes maia Smith, 1879 = Cryptolithodes Brandt, 1848 (type species (misspelling of maja) Cryptolithodes typicus Brandt, 1848, by monotypy; = Maia vulgaris Bosc, 1802 gender masculine) (suppressed generic name ICNZ Opinion 511) Cryptolithodes expansus Miers, 1879 Lithodes mamillifer Macpherson, 1988 Cryptolithodes sitchensis Brandt, 1853 Lithodes manningi Macpherson, 1988 = Cryptolithodes alta-fissura Bate, 1864 Lithodes megacantha Macpherson, 1991 Cryptolithodes typicus Brandt, 1848 Lithodes murrayi Henderson, 1888 = Cryptolithodes brevifrons Miers, 1879 Lithodes nintokuae Sakai, 1978 Glyptolithodes Faxon, 1895 Lithodes panamensis Faxon, 1893 = Glyptolithodes Faxon, 1895 Lithodes paulayi Macpherson & Chan, 2008 (type species Rhinolithodes cristatipes Faxon, Lithodes rachelae Ahyong, 2010 1893, by monotypy; gender masculine) Lithodes richeri Macpherson, 1990 Glyptolithodes cristatipes (Faxon, 1893) Lithodes robertsoni Ahyong, 2010 [Rhinolithodes] Lithodes santolla (Molina, 1782) [Cancer] {6} Lithodes Latreille, 1806 = Lithodes antarctica Hombron & Jacquinot, = Lithodes Latreille, 1806, 1842 {6} (type species Cancer maja Linnaeus, 1758, by = Pseudolithodes zenkevitschi Birstein & Vinogradov, monotypy; gender masculine) 1972 = Pseudolithodes Birstein & Vinogradov, 1972 Lithodes turkayi Macpherson, 1988 (type species Pseudolithodes zenkevitchi Birstein Lithodes turritus Ortmann, 1892 & Vinogradov, 1972, by original designation; Lithodes unicornis Macpherson, 1984 {7} gender masculine) Lithodes wiracocha Haig, 1974 Lithodes aequispinus Benedict, 1895 {2, 3} Lopholithodes Brandt, 1848 = Paralithodes longirostris Navozov-Lavrov, = Lopholithodes Brandt, 1848 (type species 1929 Lopholithodes mandtii Brandt, 1848 by monotypy; Lithodes aotearoa Ahyong, 2010 gender masculine) Lithodes australiensis Ahyong, 2010 = Ctenorhinus Gibbons, 1855 (type species Lithodes ceramensis Takeda & Nagai, 2004 Ctenorhinus setimanus Gibbons, 1855, by Lithodes chaddertoni Ahyong, 2010 monotypy; gender masculine) Lithodes confundens Macpherson, 1988 = Echinocerus White, 1848 (type species Echinocerus Lithodes couesi Benedict, 1895 {2} cibarius White, 1848, by monotypy; gender Lithodes ferox Filhol, 1885 {4} masculine) = Lithodes tropicalis A. Milne-Edwards, 1883 Lopholithodes foraminatus (Stimpson, 1859) (nomen nudum) [Echinocerus] = Lithodes tropicalis Bouvier, 1895 Lopholithodes mandtii Brandt, 1848 = Pseudolithodes pyriformis Birstein & Vinogradov, = Echinocerus cibarius White, 1848 1972 = Ctenorhinus setimanus Gibbons, 1855 Lithodes formosae Ahyong & Chan, 2010 Neolithodes A. Milne-Edwards & Bouvier, 1894 {8} Lithodes galapagensis Hall & Thatje, 2009 = Neolithodes A. Milne-Edwards & Bouvier, 1894 Lithodes jessica Ahyong, 2010 (type species Lithodes Grimaldii A. Milne- Lithodes longispina Sakai, 1971 Edwards & Bouvier 1894, by original designation; Lithodes macquariae Ahyong, 2010 gender masculine) Lithodes maja (Linnaeus, 1758) [Cancer] Neolithodes agassizii (Smith, 1882) [Lithodes] = Cancer horridus Pennant, 1777 Neolithodes asperrimus Barnard, 1947 = Cancer spinosus Ascanius, 1776 Neolithodes brodiei Dawson & Yaldwyn, 1970 = Cancer spinosus Amboinensis Seba, 1759 Neolithodes bronwynae Ahyong, 2010 = Cancer spinosus maximus, orientalis Seba, 1759 Neolithodes capensis Stebbing, 1905 (non binominal, not valid name) Neolithodes diomedeae (Benedict, 1895) = Parthenope Maja Fabricius, 1798 [Lithodes] {2} = Inachus Maja Fabricius, 1798 = Neolithodes martii Birstein & Vinogradov, 1972 = Maja eriocheles Lamarck, 1801 (invalid name Neolithodes duhameli Macpherson, 2004 ICZN 511) Neolithodes flindersi Ahyong, 2010 = Lithodes arctica Latreille, 1806 (invalid name Neolithodes grimaldii (A. Milne-Edwards & Bouvier, ICZN 511) 1894) [Lithodes] = ? Lithode douteuse H. Milne Edwards, 1837 (not = Lithodes goodei Benedict, 1895 {2} Latin, not valid name) {5} Neolithodes nipponensis Sakai, 1971 = ? Lithodes dubius Brandt, 1848 {5} Neolithodes vinogradovi Macpherson, 1988 12 THE RAFFLES BULLETIN OF ZOOLOGY 2010 Neolithodes yaldwyni Ahyong & Dawson, 2006 = Lithodes verrucosa Dana, 1852 Paralomis grossmani Macpherson, 1988 Paralithodes Brandt, 1848 Paralomis haigae Eldredge, 1976 Paralomis hirtella de Saint Laurent & Macpherson, = Paralithodes Brandt, 1848 (type species Lithodes 1997 Paralomis histrix (De Haan, 1849) [Lithodes] brevipes H. Milne Edwards & Lucas 1841, by Paralomis hystrixoides Sakai, 1980 Paralomis inca Haig, 1974 monotypy; gender masculine) Paralomis indica Alcock & Anderson, 1899 Paralomis investigatoris Alcock & Anderson, 1899 Paralithodes brevipes (H. Milne Edwards & Lucas, 1841) Paralomis jamsteci Takeda & Hashimoto, 1990 Paralomis japonicus Balss, 1911 [Lithodes] Paralomis kyushupalauensis Takeda, 1985 Paralomis longidactylus Birstein & Vinogradov, 1972 Paralithodes californiensis (Benedict, 1895) {14} [Lithodes] {2, 9} Paralomis longipes Faxon, 1893 Paralomis makarovi Hall & Thatje, 2009 Paralithodes camtschaticus (Tilesius, 1815) [Maja] Paralomis manningi Williams, Smith & Baco, 2000 Paralomis medipacifica Takeda, 1974 = Lithodes spinosissimus Brandt, 1848 Paralomis mendagnai Macpherson, 2003 Paralomis microps Filhol, 1884 = Lithodes Japonicus De Haan 1849 = Rhinolithodes biscayensis Bouvier, 1895 {15} = Paralithodes rostrofalcatus MacKay, 1932 Paralomis multispina (Benedict, 1895) [Leptolithodes] Paralithodes platypus (Brandt, 1850) [Lithodes] {2} Paralomis nivosa Hall & Thatje, 2009 Paralithodes rathbuni (Benedict, 1895) Paralomis ochthodes Macpherson, 1988 Paralomis odawarai (Sakai, 1980) [Lopholithodes] [Lithodes] {2, 9} Paralomis otsuae Wilson, 1990 Paralomis pacifica Sakai, 1978 Paralomis White, 1856 Paralomis papillata (Benedict, 1895) [Leptolithodes] = Paralomis White, 1856 (type species Lithodes {2} Paralomis pectinata Macpherson, 1988 granulosa (Hombron & Jacquinot, 1846, by Paralomis phrixa Macpherson, 1992 Paralomis poorei Ahyong, 2010 original designation; gender feminine) {10} Paralomis roeleveldae Kensley, 1981 Paralomis seagranti Eldredge, 1976 = Acantholithus Stimpson, 1858 (type species Paralomis serrata Macpherson, 1988 Paralomis spectabilis Hansen, 1908 Lithodes hystrix De Haan, 1849, by original Paralomis spinosissima Birstein & Vinogradov, 1972 Paralomis staplesi Ahyong, 2010 designation; gender masculine) Paralomis stella Macpherson, 1988 Paralomis stevensi Ahyong & Dawson 2006 = Leptolithodes Benedict, 1895 [type species Paralomis taylorae Ahyong, 2010 Paralomis truncatispinosa Takeda & Miyake, 1980 Paralomis aculeata Henderson, 1888 by = Paralomis heterotuberculata Tung, Yang & Li, subsequent designation by Ahyong et al. (2010); 1984 gender masculine] {2, 11} Paralomis tuberipes Macpherson, 1988 Paralomis verrilli (Benedict, 1895) [Pristopus] {2} = Pristopus Benedict, 1895 [type species Pristopus Paralomis webberi Ahyong, 2010 Paralomis zealandica Dawson & Yaldwyn, 1971 verrilli Benedict, 1895, by subsequent designation = Paralomis shinkaimaruae Takeda & Hatanaka, by Ahyong et al. (2010) designation; gender 1984 masculine] {2, 12} Phyllolithodes Brandt, 1848 = Phyllolithodes Brandt, 1848 (type species Paralomis aculeata Henderson, 1888 Phyllolithodes papillosus Brandt, 1848 by monotypy; gender masculine) Paralomis africana Macpherson, 1982 = Petaloceras White, 1856 (type species Petaloceras bellianus White, 1856, by monotypy; gender Paralomis alcockiana Hall & Thatje, 2009 masculine) Paralomis anamerae Macpherson, 1988 Paralomis arae Macpherson, 2001 Paralomis arethusa Macpherson, 1994 Paralomis aspera Faxon, 1893 Paralomis birsteini Macpherson, 1988 Paralomis bouvieri Hansen, 1908 Paralomis ceres Macpherson, 1989 Paralomis chilensis Andrade, 1980 Paralomis cristata Takeda & Ohta, 1979 Paralomis cristulata Macpherson, 1988 Paralomis cubensis Chace, 1939 Paralomis danida Takeda & Bussarawit, 2007 Paralomis dawsoni Macpherson, 2001 Paralomis diomedeae (Faxon, 1893) [Echinocerus] Paralomis dofleini Balss, 1911 Paralomis echidna Ahyong, 2010 Paralomis elongata Spiridonov, Türkay, Arntz & Thatje, 2006 Paralomis erinacea Macpherson, 1988 Paralomis formosa Henderson, 1888 Paralomis gowlettholmes Ahyong, 2010 {13} Paralomis granulosa (Hombron & Jacquinot, 1846) [Lithodes] {10} = Lithodes granulata Jacquinot & Lucas, 1854 {10} 13 McLaughlin et al.: Checklist of world Lithodoidea, Lomisoidea and Paguroidea Phyllolithodes papillosus Brandt, 1848 species to A. Milne-Edwards (1883), although in = Petaloceras bellianus White, 1856 his “References” Macpherson incorrectly cited the = Phyllolithodes bicornis Bate, 1866 publication date as 1884. Authorship of L. ferox correctly belongs to Filhol (1885). As pointed out Rhinolithodes Brandt, 1848 by Macpherson (1988b), A. Milne-Edwards (1883) = Rhinolithodes Brandt, 1848 (type species mentioned only that a distinct species from off West Rhinolithodes wosnessenskii Brandt, 1848, by Africa was designated Lithodes tropicalis. As Milne- monotypy; gender masculine) Edwards’ remark was not accompanied by either a description or an illustration, the name was a nomen Rhinolithodes wosnessenskii Brandt, 1848 nudum. The species was first formally described Sculptolithodes Makarov, 1934 by Bouvier (1895). However, in the interim, Filhol (1885) described Lithodes ferox from the same area. = Sculptolithodes Makarov, 1934 (type species Although Filhol’s specimens proved to be juveniles Sculptolithodes derjugini Makarov, 1934, by of L. tropicalis, Filhol’s name had priority and L. monotypy; gender masculine) tropicalis became a junior synonym. Sculptolithodes derjugini Makarov, 1934 {5} Among the plates of Seba’s (1734–1765; 1828–1831) Locupletissimi Rerum Naturalium Thesauri & NOTES Planches de Seba (Holthuis 1969), was a figure of a lithodid considered by H. Milne Edwards (1837: {1} In his table of gill elements in lithodids, Macpherson 186) to depict the same species as an unidentified (1988b: 18) indicated that pleurobranchs were entirely specimen in the collections of the Muséum national absent. This is incorrect. Only in a few genera of d’Histoire naturelle. Because the specimen was in the Paguridae are pleurobranchs lost completely; poor condition, and he was unsure of its conspecificity lithodoids all have a pleurobranch on the thoracic with the unidentified museum specimens H. Milne wall above each fourth pereopod (McLaughlin et al., Edwards gave it the name “Lithode douteuse” using 1997). a question mark to indicate his doubt. Brandt (1848) reported the species in its Latinized form as Lithodes {2} Although correctly cited by Dawson (1989), as noted dubius M. Edw. but although he repeated H. Milne by Shirley (2002), the date of publication of Benedict’s Edwards’ (1837) diagnosis, noted that he had not seen report on the lithodids collected by the U.S. Fish the species. Subsequently, Brandt (1851: 93, footnote) Commission steamer Albatross has been incorrectly suggested that “Lithode douteuse” was probably a cited by numerous authors including Hart (1982), specimen of Lithodes arcticus, but because of his Macpherson (1988b) and Williams et al. (1989). uncertainty, retained H. Milne Edwards’ question Benedict’s paper was issued as a “separate” (1016) mark. for volume 17 of the Proceedings of the U.S. National Museum. The separate’s cover carried the date 1894; {6} Despite being first described by Molina (1782) as however, according to the table of contents for the Cancer santolla, this species was reported under volume, Benedict’s paper was actually published on the name “Lithodes antarctica Jacquinot” for many 29 January 1895. years. As discussed by Macpherson (1988), the date of publication of Jacquinot’s species was uncertain, as {3} Also correctly cited by Dawson (1989), and noted the expedition results were published in several parts by Shirley (2002), the spelling of the specific name and the illustrations published separately. Based on of this taxon has been incorrectly cited by numerous the mention by H. Milne Edwards & Lucas (1842- authors including Macpherson (1988b) and Williams 1844: vol. 6(1): 32, pl. 7) of Lithodes antarctica, the et al. (1989). Benedict (1895) described the species correct date was taken to be 1844. However, Clark & as Lithodes aequispinus, incorrectly assigning a Crosnier (2000) painstakenly traced the accurate dates masculine ending to the feminine noun spina. It does of publication of the 28 livraisons that comprised the not appear that this is an inadvertent error on the Atlas of the “Voyage au pole …” and determined that author’s part; endings of Latin nouns are not changed the correct date of publication was 1842. Authorship to agree in gender with generic name. Nonetheless, was corrected by Holthuis (2002) from Jacquinot Bouvier (1896) corrected the spelling to the original in Hombron & Jacquinot to Hombron & Jacquinot, form of the noun, and for many years this species was October, 1842. reported under the name Lithodes aequispina. The current International Code of Zoological Nomenclature {7} Dawson & Yaldwyn (1985) and Dawson (1989) (ICZN1999: Art. 32.2) is unambiguous in stating that considered L. unicornis to be junior synonym of L. the original spelling is the correct spelling unless it is murrayi. While acknowledging the morphological shown to be an inadvertent error; thus L. aequispinus similarities between the two species, Macpherson as the original spelling is correct. (1988) believed that certain characters appeared to confirm their distinctness and we accept his {4} Macpherson (1988b) corrected the error in authorship judgment. of L. ferox made by both Dawson & Yaldwyn (1985) and Dawson (1989). Those authors attributed the 14 THE RAFFLES BULLETIN OF ZOOLOGY 2010 {8} Davie (2002) cited monotypy as the type designation results of the expedition of the “Astrolabe” and for Neolithodes, and the source as A. Milne-Edwards “Zélée”, Clark & Crosnier (2000) determined that & Bouvier (1894). Actually, the type was fixed by Paralomis granulosa (as Lithodes) was first published A. Milne-Edwards & Bouvier (1894a) by original in July of 1846 as part of Atlas plate 8. The text of designation. These authors included two species, volume 3 Crustacés was published by Jacquinot and N. grimaldii and N. agassizi (Smith, 1882) in their Lucas in 1854 where the species was cited as Lithodes remarks accompanying the original description, granulata. Holthuis (2002) subsequently confirmed both species initially having been included in the the correct authorship as Lithodes granulosa Hombron genus Lithodes. A. Milne-Edwards & Bouvier & Jacquinot, 1846. (1894a) described L. grimaldii on page 62 with accompanying plate 3. However, in an appendix to {11} Benedict (1895) established the genus Leptolithodes that report (A. Milne-Edwards & Bouvier, 1894a: for certain species previously assigned to Paralomis 91) the authors introduced the description of their that he believed differed from the designated type new genus, Neolithodes with the following statement. species, Lithodes granulosus. Although he included “En comparant Lithodes Grimaldii (voir ci-dessus, p. three previously described taxa and added two new 62) avec les autres représentants de la sous-famille species of his own, Benedict did not designate a des Lithodinés nous avons pu nous convaincre qu’il type species. In a review of the lithodids of Taiwan, diffère de ceux-ci par la structure de son abdomen Ahyong et al. (2010) designated Paralomis aculeatus et par la forme de son rostre; comme ces differences as the type species of Leptolithodes. sont d’ordre essentiellement générique dans la sous- famille, il y a lieu de ranger l’espèce de l’Hirondelle {12} Benedict (1895) established the genus Pristopus, dans un nouveau genre pour lequel nous proposons le which he considered similar to Leptolithodes. To nom de Neolithodes.” The description following was Pristopus, Benedict assigned Henderson’s Paralomis identical to the one published by A. Milne-Edwards & formosa and his own new species Pristopus verrilli. Bouvier (1894b), but the latter included two different Again, Benedict did not designate a type species. illustrations. Despite being titled “Neolithodes genre Ahyong et al. (2010) designated Paralomis verrilli nouveau de la sous-famillle des Lithodinés” the second as the type species of Pristopus. description was published on 10 July 1894, whereas the first appeared in print on 1 June 1894. {13} The specific epithet, gowlettholmes, is treated as a noun in apposition by the author, and is correct as {9} The original descriptions of Paralithodes californiensis written (Ahyong, per. comm.). and P. rathbuni were based on two female specimens of the former and one male of the latter, respectively. {14} Birstein & Vinogradov (1972) described their new Benedict (1895) emphasized the appreciable similarities species as Paralomis longidactylus, the “us” ending between the two taxa, but indicated that he was being correct for a masculine noun. However, Takeda hesitant to consider them a single species. Although et al. (1984) changed the specific ending to “a”, three additional females were subsequently reported presumably because the generic name Paralomis from off San Diego, California (Schmitt, 1921), Dr. is feminine. The original spelling has been retained Schmitt, curator of Crustacea the National Museum (ICZN Art. 32.2); dactylus is a noun and as such, need of Natural History for many years, expressed serious not agree in gender with its generic name (ICZN Art. doubts about their distinctiveness and suggested that 31.2.1). perhaps they simply represented males and females of a single taxon (personal communication to PMcL). {15} Authorship of Rhinolithodes biscayensis, a synonym Both species are still reported as part of the eastern of Paralomis microps, was credited to A. Milne- Pacific fauna (e.g., Anderson & Cailliet, 1974; Edwards & Bouvier (1894a) by Bouvier (1896), an Wicksten, 1987; Zaklan, 2002); however, their true error repeated by Dawson (1989), and to A. Milne- status is unclear. Edwards & Bouvier (1900) by Dawson & Yaldwyn (1985). As pointed out by Macpherson (1988b) the {10} Both authorship and publication date for this species original description of the species by the two authors have been sources of disagreement. Macpherson was never published. The first published description (1988b) reemphasized Haig’s (1955) contention that and illustrations were presented by Bouvier (1895); no authorship was established in Hombron & Jacquinot, mention was made of co-authorship, thus the species 1847, page 4, Section Crustacea, volume 3, where the is correctly attributed to Bouvier. species was attributed to Jacquinot. Although the text was not published until 1854, publication of the figures SUPERFAMILY LOMISOIDAE BOUVIER, 1895 preceded it, and while no actual date of publication was specified, White’s (1847) inclusion of the species Family Lomisidae Bouvier, 1895 in his list of Crustacea in the British Museum was = Lomisinés Bouvier 1894 (not Latin, not valid accepted as an indication of the correct date. However, name) {1} in their detailed account of the published zoological 15 McLaughlin et al.: Checklist of world Lithodoidea, Lomisoidea and Paguroidea = Lomisidae Bouvier 1895 Coenobita cavipes Stimpson, 1858 = Lomidae Glassner, 1969 (incorrect stem) = Cenobita cavipes Stimpson, 1858 (incorrect Lomis H. Milne Edwards, 1837 = Porcellana hirta (type species Porcellana hirta spelling of Coenobita) Lamarck, 1818, by monotypy; gender feminine) = Coenobita baltzeri Neumann, 1878 Lomis hirta (Lamarck, 1818) [Porcellana] Coenobita clypeatus (Fabricius, 1787) [Pagurus] {6}{8} = Cancellus Terrestris Bahamensis Catesby, 1743 (pre-Linnean name, not valid) {7} Coenobita compressus H. Milne Edwards, 1836 NOTES = Cenobita compressa H. Milne Edwards, 1836 {1} Lemaitre & McLaughlin (2009) erred when they (incorrect spelling of Coenobita) corrected the date for the Lomisoidea from 1895 to 1894. They were citing the date that Bouvier published = Cenobita intermedia Streets, 1871 (incorrect the family group name in the French vernacular, but that was not a valid name. Only when the name spelling of Coenobita) was Latinized by Bouvier in 1895 did it become an acceptable family-group name. = Cenobita panamensis Streets, 1871 (incorrect spelling of Coenobita) Coenobita compta White, 1847 (nomen nudum) {9} Coenobita laeviuscula White, 1847 (nomen nudum) {9} Coenobita longitarsis De Man, 1902 Coenobita olivieri Owen, 1839 SUPERFAMILY PAGUROIDEA LATREILLE, 1802 = Caenobita olivieri Owen, 1839 (incorrect spelling of Coenobita) Family Coenobitidae Dana, 1851 Coenobita perlatus H. Milne Edwards, 1837 = Cenobita perlata H. Milne Edwards, 1837 Birgus Leach, 1816 {1} (incorrect spelling of Coenobita) = Birgus Leach (type species Cancer latro Linnaeus, = Coenobita perlatus var. affinis Miers, 1880 1767, by monotypy; gender masculine) Coenobita purpureus Stimpson, 1858 Birgus latro (Linnaeus, 1767) [Cancer] = Cenobita purpurea Stimpson, 1858 (incorrect = Cancer crumenatus Rumphius, 1705 (pre-linnean spelling of Coenobita) name; not valid) Coenobita pseudorugosus Nakasone, 1988 = Cancer crumenatus orientalis Seba, 1759 {2} Coenobita rubescens Greeff, 1884 = Cancer (Astacus) latro Linnaeus, 1767 Coenobita rugosus H. Milne Edwards, 1837 = Birgus laticauda Latreille, 1829 = Cenobita rugosa H. Milne Edwards, 1837 Coenobita Latreille, 1829 {3} (incorrect spelling of Coenobita) = Coenobita Latreille, 1829 (type species Pagurus = Coenobita subrugosa Neumann, 1878 clypeatus Fabricius, 1787, by monotypy, gender = Coenobita rugosa var. wagneri Doflein, 1900 masculine) {4} Coenobita scaevola (Forskål, 1775) [Cancer] = Eremita Osbeck, 1765 (type species Eremita = Coenobita rugosa var. Jousseaumi Bouvier, javanica Osbeck, 1765, by monotypy) {4} 1890 = Carcinion Jarocki, 1825 (type species Pagurus = Coenobita rugosa var. granulata Bouvier, 1890 clypeatus Olivier = Coenobita olivieri Owen, Coenobita spinosus H. Milne Edwards, 1837 1839, by monotypy) {4} = Cenobita spinosa H. Milne Edwards, 1837 = Coénbite Latreille, 1825 (not Latin, unavailable (incorrect spelling of Coenobita) name) = Cenobita brunnea Dana, 1851 (incorrect spelling = Cenobites Berthold 1827 (type species Pagurus of Coenobita) clypeatus Fabricius, 1787, by monotypy) {4} = Birgus hirsutus Hess, 1865 = Cenobita H. Milne Edwards 1837 (incorrect Coenobita variabilis Mc Culloch, 1909 spelling of Coenobita) Coenobita violascens Heller, 1862 Coenobita brevimanus Dana, 1852 = Cenobita violascens Heller, 1862 (incorrect = Cenobita clypeata var. brevimanus Dana, 1852 spelling of Coenobita) (incorrect spelling of Coenobita) = Coenobita hilgendorfi Terao 1913 (unnecessary replacement name) {5} NOTES = Coenobita clypeata var. puerto-galerae Yap- {1} The date of publication usually given for Birgus is 1815. But as pointed out by Davie (2002) while the Chiongco, 1938 volume of the Transactions of the Linnean Society, London is dated 1815, the actual publication date is Coenobita carnescens Dana, 1851 24 January 1816. = Cenobita carnescens Dana, 1851 (incorrect spelling of Coenobita) 16 THE RAFFLES BULLETIN OF ZOOLOGY 2010 {2} The publication date for volume 3 of Seba’ Edwards (1837) credited Herbst (1791), whereas Locupletissimi rerum naturalium thesauri … Hilgendorf (1869) believed the author to be H. Milne routinely was given as 1761 by carcinologists and Edwards (1837) and Rathbun (1919) attributed the 1758 by ichthyologists. However, Holthuis (1969) species to Latreille, but incorrectly cited with date demonstrated conclusively that the correct date for the and reference to Olivier (1811). The correct author original publication was 1759. A title reprint edition is Fabricius (1787). was published in 1761. {7} Catesby’s initial publication is usually cited with the {3} Morgan & Holthuis (1989) pointed out that while some date 1754, which is the date of the second edition. authors routinely considered Coenobita masculine, Catesby’s (1743, pl. 33, figs. 1, 2) illustrations were a similar number thought it to be feminine. Thus reproduced by Latreille (1818) as Pagurus diogenes the literature is “sprinkled” with reports of the same Linnaeus. specific names ending in “us” or “a” depending upon the author(s). As explained by Morgan & Holthuis, {8} Not only has authorship of Coenobita clypeatus been a Coenobita is later ecclesiastical Latin, meaning “a problem, the actual identity of the species has been as monk”, and while the majority of Latin names ending well, although the problem went unrecognized for two in “a” are feminine in gender, Coenobita is masculine. hundred years. Fabricius (1787) based his description Consequently, the correct ending for species-group of the species on Herbst’s (1791), at the time names in Coenobita is the masculine suffix “us.” unpublished, illustrations of two specimens reported to be from the East Indies. Subsequent carcinologists {4} Morgan & Holthuis (1988), concerned that generic incorrectly assumed that both figures represented name Coenobita might be replaced by one of three the same specimen, one simply an enlargement. In existing senior, but obscure, synonyms, applied to a review Herbst’s collection in the Naturhistorisches the International Commission for conservation of the Forschungsinstitut Museum für Naturkunde zu name Coenobita. They argued: 1) Although Jarocki’s Berlin, Hilgendorf (1869) was convinced that the two (1825) Carcinion was a senior objective synonym, it specimens of Coenobita present in the collection did was an entirely forgotten name whose reintroduction not represent C. clypeatus. As a result of his inaccurate would cause considerable undesirable confusion; 2) interpretation of the taxon, Hilgendorf concluded that Berthold’s (1827) Cenobites was simply a translation Herbst (1791) had erred in citing the type locality. He of Latreille’s (1825) vernacular Cénobite and its “corrected” the type locality to West Indies. Although reintroduction into carcinological literature would the West Indian Coenobita had been known for many result in great confusion. Osbeck’s (1765) Eremita years as Coenobita diogenes (Linnaeus), Rathbun posed a somewhat different problem. Despite the (1919) pointed out that Linnaeus’ (1758) Pagurus generalities of Osbeck’s description, which could diogenes represented a species of the marine genus justify Eremita’s possible status as a nomen dubium, Petrochirus, and she adopted ‘Coenobita clypeatus the name is the oldest name ever applied to a Herbst’ for the terrestrial species. It was not until hermit crab thus posing a serious threat to paguroid Herbst’s (1791) specimens were critically reexamined nomenclature. In Opinion 1575, March 1990, the (McLaughlin & Holthuis, 2002) was it realized that Commission placed Coenobita and its type species, Herbst had confounded two species, C. rugosus C. clypeatus, on the Official List of Valid Names in H. Milne Edwards, 1837, and C violascens Heller, Zoology, and the names Carcinion, Cenobites and 1862, both Indo-Pacific taxa, under the specific name Eremita with its type species E. javanick Osbeck, clypeatus. An application was then submitted to the 1765, on the Official Index of Rejected and Invalid ICZN to replace the two existing East Indies syntypes Names in Zoology. with a West Indies neotype. By so doing, Coenobita clypeatus, as commonly used for many years would {5} Terao (1913) proposed the replacement name, C. be retained, as would the two common East Indies hilgendorfi for Coenobita clypeatus sensu Latreille species. In Opinion 2052 (2003), the ICZN set aside (1826), stating: “on Hilgendorf’s authority we learn all previous type fixations for the nominal species that Cancer clypeatus Herbst (1791) is not identical Pagurus clypeatus Fabricius, 1767, and designated the with Latreille’s Coenobita clypeata (1826) but is the Caribbean specimen USNM 126773 as the neotype. same as Coenobita diogenes M.–Edwards (1837). Additionally, Pagurus clypeatus Fabricius, 1787, … Since, now, this C. clypeatus (Herbst) is clearly was added to the Official List of Specific Names a species distinct from C. clypeata Latr., the latter in Zoology, with an endorsement recording that the must receive a new specific name.” However, Terao’s species is defined by its designated neotype. Cenobita (1913) interpretation of C. clypeatus was based on a rugosus H. Milne Edwards, 1837, and Cenobita misidentification of C. clypeatus (= C. brevimanus). violascens were similarly added to the Official List. {6} Until recently, considerable confusion has existed {9} In his list of Crustacea in the British Museum, Adam regarding the correct authorship of Coenobita White (1847: 59–62) listed as new species, Coenobita clypeatus. For example, Latreille (1826) and H. Milne compta from the collection of Mr. Broderip, and 17 McLaughlin et al.: Checklist of world Lithodoidea, Lomisoidea and Paguroidea Coenobita laeviuscula from Port Essington. Neither Areopaguristes perspicax (Nobili, 1906) [Paguristes] taxon was described nor illustrated and mention of neither could be found in any of White’s other = Paguristes jousseaumii var. perspicax Nobili, publications. 1906 Areopaguristes pilosus (H. Milne Edwards, 1836) [Pagurus] Areopaguristes rubrodiscus (Forest, 1952) [Paguristes] Family Diogenidae Ortmann, 1892 Areopaguristes setosus (H. Milne Edwards, 1848) [Pagurus] Allodardanus Haig & Provenzano, 1965 = Pagurus setosus Filhol, 1885 = Allodardanus Haig & Provenzano, 1965 Areopaguristes taenia (Komai, 1999) [Paguristes] (type species Allodardanus rugosus Haig & Areopaguristes tuberculatus (Whitelegge, 1900) Provenzano, 1965, by original designation; gender [Paguristes] masculine) Areopaguristes virilis (Forest, 1952) [Paguristes] Allodardanus bredini Haig & Provenzano, 1965 Bathynarius Forest, 1989 Allodardanus midas McLaughlin & Gore, 1985 = Bathynarius Forest, 1989 (type species Clibanarius Allodardanus rugosus Haig & Provenzano, 1965 anomalus A. Milne-Edwards & Bouvier, 1893, by Aniculus Dana, 1852 original designation; gender masculine) = Aniculus Dana, 1852 (type species Pagurus Bathynarius albicinctus (Alcock, 1905) [Clibanarius] aniculus Fabricius, 1787, by original designation; Bathynarius anomalus (A. Milne-Edwards & Bouvier, gender masculine) 1893) [Clibanarius] Aniculus aniculus (Fabricius, 1787) Bathynarius izuensis Komai & Takeda, 2004 = Aniculus typicus Dana, 1852 Bathynarius pacificus Forest, 1993 Aniculus elegans Stimpson, 1858 Bathynarius wolffi Forest, 1993 = Aniculus longitarsis Streets, 1871 Calcinus Dana, 1851 {3} Aniculus erythraeus Forest, 1984 = Calcinus Dana, 1851 (type species Cancer tibicen Aniculus hopperae McLaughlin & Hoover, 1996 Herbst, 1791, by original designation; gender Aniculus maximus Edmondson, 1952 masculine) Aniculus miyakei Forest, 1984 Calcinus albengai Poupin & Lemaitre, 2003 Aniculus retipes Lewinsohn, 1982 Calcinus anani Poupin & McLaughlin, 1998 Aniculus sibogae Forest, 1984 Calcinus argus Wooster, 1984 Aniculus ursus (Olivier, 1812) [Pagurus] {1} Calcinus californiensis Bouvier, 1898 Aeropaguristes Rahayu & McLaughlin, 2010 Calcinus chilensis (H. Milne Edwards, 1836) = Aeropaguristes Rahayu & McLaughlin, 2010 (type [Pagurus] {4} species Pagurus setosus Filhol, 1885 by original Calcinus dapsiles Morgan, 1989 designation; gender masculine) (replacement Calcinus elegans (H. Milne Edwards, 1836) [Pagurus] name) {2} = ? Pagurus fasciatus Bell, 1853 {5} = Stratiotes Thomson, 1899 (junior homonym of Calcinus explorator Boone, 1930 Stratiotes Putzeys, 1846, Coleoptera) Calcinus gaimardii (H. Milne Edwards, 1848) Areopaguristes abbreviatus (Dechancé, 1963) [Pagurus] [Paguristes] Calcinus gouti Poupin, 1997 Areopaguristes breviantennatus (Rahayu, 2005) Calcinus guamensis Wooster, 1984 [Stratiotes] Calcinus haigae Wooster, 1984 Areopaguristes cyanops (Forest, 1978) [Paguristes] Calcinus hakahau Poupin & McLaughlin, 1998 Areopaguristes difficilis (Forest, 1952) [Paguristes] Calcinus hazletti Haig & McLaughlin, 1984 Areopaguristes engyops (Barnard, 1947) [Paguristes] Calcinus imperialis Whitelegge, 1901 Areopaguristes hewatti (Wass, 1963) [Paguristes] Calcinus inconspicuus Morgan, 1991 Areopaguristes hispidus (A. Milne-Edwards & Bouvier, Calcinus isabellae Poupin, 1997 1892) [Paguristes] Calcinus kurozumii Asakura & Tachikawa, 2000 Areopaguristes hummi (Wass, 1955) [Paguristes] Calcinus laevimanus (Randall, 1840) [Pagurus] Areopaguristes iris (Forest & de Saint Laurent, 1968) = Pagurus tibicen H. Milne Edwards, 1836 [Paguristes] (preoccupied name) Areopaguristes japonicus (Miyake, 1961) [Paguristes] = Pagurus lividus H. Milne Edwards, 1848 Areopaguristes mclaughlinae (Ayón Parente & Hendrickx, = Calcinus herbstii De Man, 1887 2006) [Stratiotes] Calcinus latens (Randall, 1840) [Pagurus] Areopaguristes micheleae (Rahayu, 2005) [Stratiotes] = Pagurus cristimanus H. Milne Edwards, 1848 Areopaguristes ngochoae (Rahayu, 2005) [Stratiotes] = Calcinus intermedius De Man, 1881 Areopaguristes nigroapiculus (Komai, 2009) = Calcinus terrae-reginae Haswell, 1882 [Stratiotes] = Calcinus abrolhensis Morgan, 1988 {6} Areopaguristes orbis (Komai, 2009) [Stratiotes] Calcinus laurentae Haig & McLaughlin, 1984 Calcinus lineapropodus Morgan & Forest, 1991 18 THE RAFFLES BULLETIN OF ZOOLOGY 2010 Calcinus mclaughlinae Poupin & Bouchard, 2006 Ciliopagurus liui Forest, 1995 Calcinus minutus Buitendijk, 1937 Ciliopagurus macrolepis Forest, 1995 Calcinus morgani Rahayu & Forest, 1999 Ciliopagurus major Forest, 1995 = Calcinus areolatus Rahayu & Forest, 1999 Ciliopagurus pacificus Forest, 1995 Calcinus nitidus Heller, 1865 Ciliopagurus plessisi Forest, 1955 Calcinus obscurus Stimpson, 1859 {7} Ciliopagurus shebae (Lewinsohn, 1969) [Trizopagurus] Calcinus orchidae Poupin, 1997 Ciliopagurus strigatus (Herbst, 1804) [Cancer] Calcinus paradoxus Bouvier, 1922 {8} = Pagurus annulipes H. Milne Edwards, 1848 Calcinus pascuensis Haig, 1974 {10} Calcinus pulcher Forest, 1958 Ciliopagurus tenebrarum (Alcock, 1905) [Aniculus] Calcinus revi Poupin & McLaughlin, 1998 Ciliopagurus tricolor Forest, 1995 Calcinus rosaceus Heller, 1861 Ciliopagurus vakovako Poupin, 2001 Calcinus seurati Forest, 1951 Clibanarius Dana, 1852 Calcinus sirius Morgan, 1991 = Clibanarius Dana, 1852 (type species Cancer Calcinus spicatus Forest, 1951 clibanarius Herbst, 1791, by original designation; Calcinus talismani A. Milne-Edwards & Bouvier, 1892 gender masculine) Calcinus tibicen (Herbst, 1791) [Cancer] Clibanarius aequabilis (Dana, 1851) [Pagurus] = Calcinus sulcatus H. Milne Edwards, 1836 {9} Clibanarius africanus Aurivillus, 1898 Calcinus tropidomanus Lewinsohn, 1981 Clibanarius albidigitus Nobili, 1901 Calcinus tubularis (Linnaeus, 1767) [Cancer] Clibanarius ambonensis Rahayu & Forest, 1993 = Pagurus ornatus Roux, 1830 Clibanarius antennatus Rahayu & Forest, 1993 = Clibanarius rouxi Heller, 1863 Clibanarius antillensis Stimpson, 1859 {11} Calcinus urabaensis Campos & Lemaitre, 1994 = Clibanarius brasiliensis Dana, 1852 Calcinus vachoni Forest, 1958 Clibanarius arethusa De Man, 1888 Calcinus vanninii Gherardi & McLaughlin, 1994 Clibanarius astathes (Stebbing, 1924) [Calcinus] Calcinus verrillii (Rathbun, 1901) [Clibanarius] Clibanarius bimaculatus (De Haan, 1849) [Pagurus] Cancellus H. Milne Edwards, 1836 Clibanarius bistriatus Rahayu & Forest, 1993 = Cancellus H. Milne Edwards, 1836 (type species Clibanarius boschmai Buitendijk, 1937 Cancellus typus H. Milne Edwards, 1836, by Clibanarius carnifex Heller, 1861 original designation; gender masculine) Clibanarius chapini Schmitt, 1926 {12} Cancellus canaliculatus (Herbst, 1804) [Cancer] Clibanarius clibanarius (Herbst, 1791) [Cancer] Cancellus frontalis Forest & McLaughlin, 2000 = Clibanarius vulgaris Dana, 1852 Cancellus investigatoris Alcock, 1905 Clibanarius cooki Rathbun, 1900 Cancellus laticoxa Forest & McLaughlin, 2000 Clibanarius corallinus (H. Milne Edwards, 1848) Cancellus macrothrix Stebbing, 1924 [Pagurus] Cancellus mayoae Forest & McLaughlin, 1998 = Pagurus globosi-manus Dana, 1851 Cancellus ornatus Benedict, 1901 = Clibanarius corallinus var. spinatus Yap- Cancellus panglaoensis McLaughlin, 2008 Chiongco, 1938 Cancellus parfaiti A. Milne-Edwards & Bouvier, 1891 Clibanarius cruentatus (H. Milne Edwards, 1848) Cancellus quadraticoxa Morgan & Forest, 1991 [Pagurus] Cancellus rhynchogonus Forest & McLaughlin, 2000 Clibanarius danai Rahayu & Forest, 1993 Cancellus sphaerogonus Forest & McLaughlin, 2000 Clibanarius demani Buitendijk, 1937 Cancellus spongicola Benedict, 1901 Clibanarius digueti Bouvier, 1898 Cancellus tanneri Faxon, 1893 Clibanarius englaucus Ball & Haig, 1972 Cancellus typus H. Milne Edwards, 1836 Clibanarius elongatus (H. Milne Edwards, 1848) Cancellus viridis Mayo, 1973 [Pagurus] {13} Ciliopagurus Forest, 1995 Clibanarius erythropus (Latreille, 1818) [Pagurus] = Ciliopagurus Forest, 1995 (type species Cancer = Pagurus misanthropus Risso, 1827 strigatus Herbst, 1804, by original designation; Clibanarius eurysternus (Hilgendorf, 1879) [Pagurus] gender masculine) Clibanarius fonticola McLaughlin & Murray, 1990 Ciliopagurus albatrossi Forest, 1995 Clibanarius foresti Holthuis, 1959 Ciliopagurus alcocki Forest, 1995 Clibanarius harisi Rahayu, 2003 Ciliopagurus babai Forest, 1995 Clibanarius hirsutimanus Kobjakova, 1971{14} Ciliopagurus caparti (Forest, 1952) [Trizopagurus] Clibanarius humilis (Dana, 1851) [Pagurus] Ciliopagurus galzini Poupin & Malay, 2009 Clibanarius inaequalis (De Haan, 1849) [Pagurus] Ciliopagurus haigae Forest, 1995 Clibanarius infraspinatus (Hilgendorf, 1869) Ciliopagurus hawaiiensis (McLaughlin & Bailey-Brock, [Pagurus] 1975) [Trizopagurus] Clibanarius janethaigae Hendrickx & Esparza-Haro, Ciliopagurus krempfi (Forest, 1952) [Trizopagurus] 1997 19 McLaughlin et al.: Checklist of world Lithodoidea, Lomisoidea and Paguroidea Clibanarius laevimanus Buitendijk, 1937 Dardanus brachyops Forest, 1962 Clibanarius lineatus (H. Milne Edwards, 1848) Dardanus calidus (Risso, 1827) [Pagurus] [Pagurus] {15} = ? Cancer hungarus Herbst, 1791{22} = Clibanarius panamensis (Stimpson, 1859) Dardanus callichela Cook, 1989 = Clibanarius lordi Miers, 1877 Dardanus corrugatus Cook, 1989 Clibanarius longitarsus (De Haan, 1849) [Pagurus] Dardanus crassimanus (H. Milne Edwards, 1836) = ? Pagurus asper H. Milne Edwards, 1848 [Pagurus] Clibanarius merguiensis De Man, 1888 {16} = Pagurus sculptipes Stimpson, 1858 = Clibanarius aequabilis var. merguiensis De Man, = Pagurus pavimentatus Hilgendorf, 1879 1888 Dardanus dearmatus (Henderson, 1888) [Pagurus] = Clibanarius misanthropus var. merguiensis De Dardanus deformis (H. Milne Edwards, 1836) Man, 1888 [Pagurus] Clibanarius nathi Chopra & Das, 1940 = Pagurus cavipes White, 1847 (nomen nudum) Clibanarius olivaceus Henderson, 1915 = Pagurus cavipes White 1847 Clibanarius pacificus Stimpson, 1858 {17} = Pagurus cultratus White, 1847 (nomen nudum) Clibanarius padavensis De Man, 1888 = ? Glaucothoe carinata Henderson, 1888 {23} Clibanarius ransoni Forest, 1953 Dardanus fucosus Biffar & Provenzano, 1972 Clibanarius rhabdodactylus Forest, 1953 Dardanus gemmatus (H. Milne Edwards, 1848) = Clibanarius zebra var. rhabdodactylus Forest, [Pagurus] 1953 Dardanus guttatus (Olivier, 1812) [Pagurus] {1} Clibanarius rosewateri Manning & Chace, 1990 {18} = ? Pagurus catephractus Boone, 1935 {24} Clibanarius rubroviria Rahayu, 1999 Dardanus hessii (Miers, 1884) [Pagurus] Clibanarius rutilus Rahayu, 1999 Dardanus imbricatus (H. Milne Edwards, 1848) Clibanarius sachalinicus Kobjakova, 1955 {19} [Pagurus] Clibanarius sclopetarius (Herbst, 1796) [Cancer] Dardanus imperator (Miers, 1881) [Pagurus] = Pagurus tuberculosus H. Milne Edwards, 1836 Dardanus impressus (De Haan, 1849) [Pagurus] = Pagurus cubensis de Saussure, 1858 Dardanus insignis (de Saussure, 1858) [Pagurus] = Clibanarius carnescens Miers, 1877 = Aniculus arrosor var. petersi A. Milne-Edwards, = Clibanarius formosus Ives, 1892 1880 Clibanarius senegalensis Chevreux & Bouvier, 1892 Dardanus jacquesi Asakura & Hirayama, 2002 Clibanarius serenei Rahayu & Forest, 1993 Dardanus janethaigae Ayón Parente & Hendrickx, Clibanarius signatus Heller, 1861 2009 = Clibanarius semistriatus Heller, 1862 Dardanus lagopodes (Forskål, 1775) [Cancer] Clibanarius similis Rahayu & Forest, 1993 = Pagurus sanguinolentus Quoy & Gaimard, 1824 Clibanarius snelliusi Buitendijk, 1937 = Pagurus affinis H. Milne Edwards, 1836 Clibanarius striolatus Dana, 1852 = Pagurus euopsis Dana, 1852 Clibanarius taeniatus (H. Milne Edwards, 1848) = Pagurus depressus Heller, 1861 [Pagurus] = Dardanus helleri Paul’son, 1875 Clibanarius tricolor (Gibbes, 1850) [Pagurus] Dardanus longior Asakura, 2006 = Clibanarius hebes Verrill, 1908 Dardanus magdalenensis Ayón Parente & Hendrickx, = Clibanarius brachyops Bouvier, 1918 2009 Clibanarius virescens (Krauss, 1843) [Pagurus] Dardanus megistos (Herbst, 1804) [Cancer] = Clibanarius philippinensis Yap-Chiongco, in = Pagurus punctulatus Miers, 1879 Estampador, 1937 = Dardanus jordani Schmitt, 1921{25} Clibanarius vittatus (Bosc, 1802) [Pagurus] Dardanus nudus Ayón Parente & Hendrickx, 2009 = Pagurus symmetricus Randall, 1840 Dardanus pectinatus (Ortmann, 1892) [Pagurus] = Clibanarius cayennensis Miers, 1877 = Pagurus arrosor var. pectinata Ortmann, 1892 = Clibanarius speciosus Miers, 1877 Dardanus pedunculatus (Herbst, 1804) [Cancer] Clibanarius willeyi Southwell, 1910 {20} = Pagurus asper De Haan, 1849 Clibanarius zebra Dana, 1852 [Pagurus] = Dardanus haani Rathbun, 1903 Dardanus Paul’son, 1875 {21} = Neopagurus horai Kamalaveni, 1950 = Dardanus Paul’son, 1875 (type species Pagurus Dardanus pilosus Ayón Parente & Hendrickx, 2009 depressus Heller, 1861, by original designation) Dardanus robustus Asakura, 2006 = Pagurias Benedict, 1901 (unnecessary replacement Dardanus rufus Buitendijk, 1937 name) Dardanus sanguinocarpus Degener, 1925 Dardanus arrosor (Herbst, 1796) [Cancer] Dardanus scutellatus (H. Milne Edwards, 1848) = Aniculus chiltoni Thompson, 1930 [Pagurus] Dardanus aspersus (Berthold, 1846) [Pagurus] = Pagurus fabimanus Dana, 1852 = Pagurus diogenes De Haan, 1849 = Pagurus watasei Terao, 1913 Dardanus australis Forest & Morgan, 1991 20 THE RAFFLES BULLETIN OF ZOOLOGY 2010 Dardanus setifer (H. Milne Edwards, 1836) [Pagurus] Diogenes manaarensis (Henderson, 1893) Dardanus sinistripes (Stimpson, 1859) [Pagurus] [Troglopagurus] = Pagurus peruensis Balss, 1921 Diogenes merguiensis De Man, 1888 Dardanus squarrosus Cook, 1989 Diogenes mercatoris Forest, 1952 Dardanus stimpsoni Ayón Parente & Hendrickx, 2009 Diogenes miles (Fabricius, 1787) [Pagurus] Dardanus sulcatus Edmondson, 1925 = Pagurus diaphanus Fabricius, 1798 Dardanus tinctor (Forskål, 1775) [Cancer] Diogenes mixtus Lanchester, 1902 = Pagurus varipes Heller, 1861 = Diogenes intermedius De Man, 1892 (preoccupied Dardanus umbella Asakura, 2006 name) Dardanus undulatus (Balss, 1921) [Pagurus] = Diogenes hainanica Wang & Dong, 1977 Dardanus venosus (H. Milne Edwards, 1848) = Diogenes plavoeti McLaughlin & Clark, 1997 [Pagurus] Diogenes moosai Rahayu & Forest, 1995 = Pagurus arrosor var. divergens Moreira, 1905 Diogenes nitidimanus Terao, 1913 {26} Diogenes ovatus Miers, 1881 Dardanus vulnerans (Thallwitz, 1892) [Pagurus] = Diogenes pugilator var. ovata Miers, 1881 Dardanus woodmasoni (Alcock, 1905) [Pagurus] Diogenes pallescens Whitelegge, 1897 Diogenes Dana, 1851 = Diogenes gardineri Alcock, 1905 = Diogenes Dana, 1851 [type species Pagurus miles = Diogenes serenei Forest, 1956 Fabricius, 1787, by subsequent designation by Diogenes paracristimanus Wang & Dong, 1977 Dana (1852); gender masculine] {27} Diogenes patae Asakura & Godwin, 2006 Diogenes alias McLaughlin & Holthuis, 2001 {28} Diogenes penicillatus Stimpson, 1858 Diogenes avarus Heller, 1865 Diogenes persicus (Nobili, 1905) [Troglopagurus] Diogenes bicristimanus Alcock, 1905 Diogenes planimanus Henderson, 1893 Diogenes biramus Morgan, 1987 = Diogenes custos var. planimanus Henderson, Diogenes brevirostris Stimpson, 1858 1893 Diogenes capricorneus Grant & McCulloch, 1907 Diogenes pugilator Roux, 1828 Diogenes costatus Henderson, 1893 = Pagurus algarbiensis Brito Capello, 1875 Diogenes crosnieri Dechancé, 1964 = Pagurus arenarius Lucas, 1846 Diogenes custos (Fabricius, 1798) [Pagurus] = Pagurus Bocagii Brito Capello, 1875 = Diogenes affinis Henderson, 1893 = Pagurus curvimanus Clément, 1874 Diogenes deflectomanus Wang & Tung, 1980 = Pagurus dillwynii Bate, 1851 Diogenes denticulatus Chevreux & Bouvier, 1892 = Diogenes pugilator var. gracillima Balss 1921 = Diogenes pugilator var. cristata Balss, 1921 (misspelling of gracillimanuus) = ? Diogenes pugilator var. subcristata Balss, = Diogenes varians var. gracillimanus Miers, 1921 1881 Diogenes dorotheae Morgan & Forest, 1991 = Diogenes intermedius Bouvier, 1891 Diogenes dubius (Herbst, 1804) [Cancer] = Pagurus Lafonti Fischer, 1872 Diogenes edwardsii (De Haan, 1849) [Pagurus] = Pagurus ponticus Kessler, 1860 Diogenes extricatus Stebbing, 1910 = Diogenes pugilator orientalis Codreanu & Diogenes fasciatus Rahayu & Forest, 1995 Balcesco, 1968 Diogenes foresti Rahayu & Hortle, 2002 Diogenes rectimanus Miers, 1884 Diogenes goniochirus Forest, 1956 Diogenes senex Heller, 1865 Diogenes granulatus Miers, 1880 Diogenes serripes (Costa, 1838) {29} Diogenes guttatus Henderson, 1888 Diogenes spinicarpus Rahayu & Forest, 1995 Diogenes inglei McLaughlin & Clark, 1997 Diogenes spinifrons (De Haan, 1849) Diogenes investigatoris Alcock, 1905 Diogenes spinulimanus Miers, 1880 Diogenes izananiae Asakura, 2006 Diogenes tomentosus Wang & Tung, 1980 Diogenes jousseaumei (Bouvier, 1897) [Troglopagurus] Diogenes tirmiziae Siddiqui & McLaughlin, 2003 = Diogenes setocristatus Morgan & Forest, 1991 Diogenes tumidus Rahayu & Forest, 1995 = Diogenes stenops Morgan & Forest, 1991 Diogenes viridis Haig & Ball, 1988 Diogenes jubatus (Nobili, 1903) [Troglopagurus] Diogenes violaceus Henderson, 1893 = Diogenes platyops Rahayu & Forest, 1995 Diogenes waltairensis Kamalaveni, 1950 Diogenes karwarensis Nayak & Neelakantan, 1989 = Diogenes custos var. waltairensis Kamalaveni, Diogenes klaasi Rahayu & Forest, 1995 1950 Diogenes laevicarpus Rahayu, 1996 Isocheles Stimpson, 1858 Diogenes lanaris Yap-Chiongco, in Estampador, 1937 = Isocheles Stimpson, 1858 (type species Bernhardus Diogenes leptocerus Forest, 1956 aequimanus Dana, 1852, by original designation; Diogenes lophochir Morgan, 1989 gender masculine) Diogenes maclaughlinae Nayak & Neelakantan, 1985 21 McLaughlin et al.: Checklist of world Lithodoidea, Lomisoidea and Paguroidea = Holopagurus Holmes, 1900 (type species Paguristes eremita (Linnaeus, 1767) [Cancer] Holopagurus pilosus Holmes, 1900, by original = Pagurus oculatus Fabricius, 1775 designation; gender masculine) = Pagurus maculatus Risso, 1827 Isocheles aequimanus (Dana, 1852) [Bernhardus] Paguristes erythrops Holthuis, 1959 Isocheles pacificus Bouvier, 1907 Paguristes fagei Forest, 1952 Paguristes fecundus Faxon, 1893 = Isocheles Wurdemanni var. pacificus Bouvier, Paguristes foresti Scelzo, 1971 1907 Paguristes frontalis (H. Milne Edwards, 1836) [Pagurus] Isocheles pilosus (Holmes, 1900) [Holopagurus] Paguristes gamianus (H. Milne Edwards, 1836) Isocheles sawayai Forest & de Saint Laurent, 1968 [Pagurus] Isocheles wurdemanni Stimpson, 1859 Paguristes geminatus McLaughlin, 2008 Loxopagurus Forest, 1964 Paguristes gonagrus (H. Milne Edwards, 1836) [Pagurus] = Loxopagurus Forest, 1964 (type species Pagurus Paguristes grayi Benedict, 1901 loxochelis Moreira, 1901, by original designation; Paguristes hernancortezi McLaughlin & Provenzano, gender masculine) 1974 Paguristes holguinensis Manjón-Cabeza, García Raso & Loxopagurus loxochelis (Moreira, 1901) [Pagurus] Martínez Iglesias, 2002 Paguristes Dana, 1851 Paguristes incomitatus Alcock, 1905 Paguristes inconstans McLaughlin & Provenzano, = Paguristes Dana, 1851 (type species Paguristes 1975 hirtus Dana, 1851, by subsequent designation by Paguristes insularis Forest, 1966 Stimpson, 1858; gender masculine) Paguristes jalur Morgan, 1992 Paguristes jousseaumei Bouvier, 1892 = Pagurites Lörenthey & Beurlen, 1929 (misspelling of Paguristes) = Paguristes jousseaumei var. glabra Nobili, 1906 = Paguristes jousseaumei var. intermedia Nobili, Paguristes acanthomerus Ortmann, 1892 Paguristes aciculus Grant, 1905 1906 Paguristes agulhasensis Forest, 1954 Paguristes kimberleyensis Morgan & Forest, 1991 Paguristes albimaculatus Komai, 2001 Paguristes lapillatus McLaughlin & Provenzano, 1975 Paguristes alegrias Morgan, 1987 Paguristes laticlavus McLaughlin & Provenzano, 1975 Paguristes alcocki McLaughlin & Rahayu, 2005 Paguristes lauriei McLaughlin & Hogarth, 1998 Paguristes anahuacus Glassell, 1938 Paguristes lewinsohni McLaughlin & Rahayu, 2005 Paguristes anomalus Bouvier, 1918 Paguristes limonensis McLaughlin & Provenzano, 1975 = Paguristes anaryballus Provenzano, 1965 Paguristes longirostris Dana, 1851 Paguristes angustithecus McLaughlin & Provenzano, Paguristes longisetosus Morgan, 1987 1974 Paguristes lymani A. Milne-Edwards & Bouvier, 1893 Paguristes antennarius Rahayu, 2006 Paguristes maclaughlinae Martinez-Iglesias & Gómez, Paguristes arostratus Rahayu, 2006 1989 Paguristes aulacis Rahayu & Forest 2009 Paguristes macrops Rahayu & Forest, 2009 Paguristes aztatlanensis Glassell, 1937 Paguristes macrotrichus Forest, 1954 Paguristes bakeri Holmes, 1900 Paguristes markhami Sandberg, 1996 Paguristes marocanus A. Milne-Edwards & Bouvier, = ? Paguristes holmesi Glassell, 1937 {30} 1892 Paguristes balanophilus Alcock, 1905 Paguristes mauritanicus Bouvier, 1906 Paguristes barbatus (Heller, 1862) [Clibanarius] Paguristes meloi Nucci & Hebling, 2004 Paguristes barnardi Forest, 1954 Paguristes microphthalmus Forest, 1952 Paguristes brachyrostris Rahayu, 2006 Paguristes miyakei Forest & McLaughlin, 1998 Paguristes brevicornis (Guérin, 1830) [Pagurus] Paguristes moorei Benedict, 1901 Paguristes brevirostris Baker, 1905 Paguristes mundus Alcock, 1905 Paguristes ocellus Komai, 2010 = Paguristes brevirostrus Mower & Shepard, 1988 Paguristes oculiviolaceus Glassell, 1937 (misspelling of brevirostris) Paguristes ortmanni Miyake, 1978 Paguristes oxyacanthus Forest, 1952 Paguristes cadenati Forest, 1954 Paguristes oxyophthalmus Holthuis, 1959 Paguristes calvus Alcock, 1905 Paguristes palythophilus Ortmann, 1892 Paguristes ciliatus Heller, 1862 Paguristes paraguanensis McLaughlin & Provenzano, Paguristes crinitimanus McLaughlin, 2008 1975 Paguristes dampierensis McLaughlin, 2008 Paguristes parvus Holmes, 1900 Paguristes depressus Stimpson, 1859 = Paguristes depressus Stimpson, 1858 (nomen nudum) Paguristes digitalis Stimpson, 1858 = Paguristes kagoshimensis Ortmann, 1892 Paguristes digueti Bouvier, 1893 Paguristes doederleini Komai, 2001 22 THE RAFFLES BULLETIN OF ZOOLOGY 2010 Paguristes pauciparus Forest & de Saint Laurent, 1968 Paguristes weddellii (H. Milne Edwards, 1848) Paguristes petalodactylus Rahayu, 2007 [Pagurus] Paguristes perplexus McLaughlin & Provenzano, 1974 = Paguristes hirtus Dana, 1851 Paguristes perrieri Bouvier, 1895 Paguristes werdingi Campos & Sanchez, 1995 Paguristes planatus A. Milne-Edwards & Bouvier, Paguristes zebra Campos & Sanchez, 1995 1893 Paguristes zhejiangensis Wang & Tung, 1982 Paguristes praedator Glassell, 1937 = Paguristes pusillus zhejiangensis Wang & Tung, Paguristes pugil McCulloch, 1913 1982 Paguristes puncticeps Benedict, 1901 Paguropsis Henderson, 1888 Paguristes puniceus Henderson, 1896 = Paguropsis Henderson, 1888 (type species = Paguristes puniceus var. unispinosa Balss, 1912 Paguropsis typicus Henderson, 1888, by monotypy; Paguristes purpureantennatus Morgan, 1987 gender feminine) Paguristes pusillus Henderson, 1896 = Chlaenopagurus Alcock, 1899 (type species Paguristes robustus Forest & de Saint Laurent, 1968 Chlaenopagurus andersoni Alcock 1899, by Paguristes rostralis Forest & de Saint Laurent, 1968 monotypy; gender masculine) Paguristes rubropictus A. Milne-Edwards & Bouvier, Paguropsis typica Henderson, 1888 {32} 1892 = Paguropsis andersoni (Alcock, 1899) = Paguristes maculatus var. rubro-picta A. Milne- [Chlaenopagurus] {33} Edwards & Bouvier, 1892 Petrochirus Stimpson, 1858 = Paguristes oculatus var. brunneo-pictus A. Milne- = Petrochirus Stimpson, 1858 (type species Pagurus Edwards & Bouvier, 1900 granulatus Olivier, 1812, by original designation; Paguristes runyanae Haig & Ball, 1988 gender masculine) Paguristes sanguinimanus Glassell, 1938 Petrochirus californiensis Bouvier, 1895 Paguristes sayi A. Milne-Edwards & Bouvier, 1893 Petrochirus diogenes (Linnaeus, 1758) [Cancer] {34} Paguristes seminudus Stimpson, 1858 = Petrochirus bahamensis (Herbst, 1791) Paguristes sericeus A. Milne Edwards, 1880 [Cancer] = Paguristes rectifrons Benedict, 1901 = Petrochirus granulatus (Olivier, 1812) = Paguristes tenuirostris Benedict, 1901 [Pagurus] {1} Paguristes simplex Rahayu & McLaughlin, 2006 Petrochirus pustulatus (H. Milne Edwards, 1848) Paguristes sinensis Tung & Wang, 1966 {31} [Pagurus] Paguristes skoogi Odhner, 1923 = Isocheles gracilis Miers, 1881 Paguristes spectabilis McLaughlin & Provenzano, = Petrochirus cavitarius Osorio, 1887 1975 Pseudopaguristes McLaughlin, 2002 Paguristes spinipes A. Milne-Edwards, 1880 = Pseudopaguristes McLaughlin, 2002 (type species = Paguristes armatus Hay, 1917 Pseudopaguristes janetkae McLaughlin, 2002, by = Paguristes visor Henderson, 1888 original designation; gender masculine) Paguristes squamosus McCulloch, 1913 Pseudopaguristes araeos Rahayu, 2007 Paguristes starcki Provenzano, 1965 Pseudopaguristes asper Rahayu, 2005 Paguristes streaensis Pastore, 1984 Pseudopaguristes bicolor Asakura & Kosuge, 2004 Paguristes subpilosus Henderson, 1888 Pseudopaguristes bollandi Asakura & McLaughlin, Paguristes sulcatus Baker, 1905 2003 Paguristes syrtensis de Saint Laurent, 1971 Pseudopaguristes calliopsis (Forest & de Saint Laurent, Paguristes tomentosus (H. Milne Edwards, 1848) 1968) [Paguristes] [Pagurus] Pseudopaguristes hians (Henderson, 1888) Paguristes tortugae Schmitt, 1933 [Paguristes] Paguristes tosaensis Komai, 2010 = Diogenes desipiens Lanchester, 1902 Paguristes triangulatus A. Milne-Edwards & Bouvier, Pseudopaguristes invisisacculus (McLaughlin & 1893 Provenzano, 1974) [Paguristes] Paguristes triangulopsis Forest & de Saint Laurent, Pseudopaguristes janetkae McLaughlin, 2002 1968 Pseudopaguristes kuekenthali (De Man, 1902) Paguristes triton McLaughlin, 2008 [Paguristes] Paguristes turgidus (Stimpson, 1857) [Clibanarius] Pseudopaguristes laurentae (Morgan & Forest, 1991) = Paguristes turgides Gordan, 1956 (misspelling of [Paguristes] turgidus) = Paguristes brachytes Komai, 1999 Paguristes ulreyi Schmitt, 1921 = Pseudopaguristes gracilis Rahayu, 2005 = Paguristes occator Glassell, 1937 Pseudopaguristes monoporus (Morgan, 1987) Paguristes versus Komai, 2001 [Paguristes] Paguristes wassi Provenzano, 1961 Pseudopaguristes shidarai Asakura, 2004 23 McLaughlin et al.: Checklist of world Lithodoidea, Lomisoidea and Paguroidea Pseudopagurus Forest, 1952 was Forest & McLaughlin (2000) that finally placed = Pseudopagurus Forest, 1952 (type species the two taxa in synonymy. However, when Rahayu Pagurus granulimanus Miers, 1881, by original (2005) found the gill number of Paguristes species designation; masculine) in the collection of the Muséum national d’Histoire naturelle variable, she restricted Paguristes to species Pseudopagurus granulimanus (Miers, 1881) having 13 pairs of gills and reinstated Stratiotes for Pseudopagurus biafrensis (Monod, 1927) species with only 12 pairs. Unfortunately Stratiotes Strigopagurus Forest, 1995 Thomson was found to be a junior homonym of Stratiotes Putzeys, 1846, and has been replaced by = Strigopagurus Forest, 1995 (type species Pagurus Areopaguristes Rahayu & McLaughlin, 2010. strigimanus White, 1847, by original designation; gender masculine) {3} A newly published molecular analysis of the genus Calcinus (Malay & Paulay, 2010) has indicated Strigopagurus bilineatus Forest, 1995 that several previously unrecognized species are Strigopagurus boreonotus Forest, 1995 distinguishable, not only at the molecular level, but also by coloration. = Clibanarius multipunctatus Wang & Tung, 1986 {4) Poupin & Bouchard (2006) considered the status of C. chilensis uncertain. Strigopagurus elongatus Forest, 1995 Strigopagurus poupini Forest, 1995 {5} On the authority of L. B. Holthuis, Biffar & Strigopagurus strigimanus (White, 1847) [Pagurus] Provenzano (1972) reported Pagurus fasciatus Bell, 1853 (as Dardanus) to be a synonym of Calcinus = Pagurus strigimanus White, 1847 (nomen elegans. The enlarged left cheliped, setose ventral nudum) margins of the propodi of the ambulatory legs, and the blue banding of these legs would support this opinion. Tisea Forest & Morgan, 1991 However, its reported collection site of Falmouth, = Tisea Forest & Morgan, 1991 (type speciesTisea England may justify Ingle’s (1993) opinion that it grandis Forest & Morgan, 1991, by original should be regarded incerta sedis. It is entered in this designation; gender feminine) checklist as a questionable synonym. Tisea grandis Forest & Morgan, 1991 {6} Although Morgan’s (1988) Australian Calcinus Trizopagurus Forest, 1952 abrolhensis proved to be conspecific with C. latens, the Hawaiian population thought to represent this = Trizopagurus Forest, 1952 (type species Clibanarius latter taxon apparently represents a distinct species. melitai Chevreux & Bouvier, 1892, by original designation; gender masculine) {7} The date of publication for Calcinus obscurus has been given in the older literature as 1862 (e.g., Gordan, Trizopagurus melitai (Chevreux & Bouvier 1892) 1956). However, Stimpson’s description appeared in [Clibanarius] a separate published in 1859, and that is the correct Trizopagurus magnificus (Bouvier, 1898) [Clibanarius] date of publication. = Clibanarius chetyrkini Boone, 1932 {8} Calcinus paradoxus, represented by a single specimen Trizopagurus rubrocinctus Forest & Garcia Raso, 1990 was thought to represent a Calcinus species by Ingle (1993) but its placement was questioned by both Incerta sedis the Bouvier (1922) and by Joseph Poupin (personal communication). Pagurus lar Heller, 1862 {9} Calcinus sulcatus H. Milne Edwards, 1836, was put NOTES into synonymy with Calcinus tibicen by Hilgendorf (1869) because he thought that Herbst (1791) had erred {1} Volume 8 of Olivier’s (1811–1812) Encyclopédie in his type locality. It was shown by McLaughlin & Methodique (Dictionnaire Encyclopédique Holthuis (2002) that it was Hilgendorf, not Herbst Méthodique) was published in two parts. Although who was in error regarding Herbst’s type locality for his paguroid descriptions most frequently have been another presumed western Atlantic species, Coenobita cited with the 1811 date, as pointed out by Lewinsohn clypeatus Herbst. Thus is possible that further (1982a), Sherborn & Woodward (1906) showed that examination of Herbst’s material will show Calcinus the second part (pages 361–722 and containing the sulcatus is a species distinct from C. tibicen. “Pagures”) was not published until 1812. {2} Forest & McLaughlin (2000) described in detail the confusion that had existed regarding the New Zealand species Paguristes pilosus and P. setosus and the errors made by Filhol (1885b, d) and Thomson (1899) that led to the establishment, by Thomson, of the genus Stratiotes with Pagurus setosus Filhol, 1885, as its type. Although Alcock (1905b) questionably considered Thomson’s (1899) Stratiotes setosus a synonym of H. Milne Edwards’ Paguristes setosus, it 24 THE RAFFLES BULLETIN OF ZOOLOGY 2010 {10} Eupagurus annulipes Stimpson, 1860, became a acensionis; however, the etymology for this species secondary junior homonym of Pagurus annulipes H. clearly indicated that C. acensionis was a laspus Milne Edwards, 1836, when the ICZN (Opinion 472) calami on the parts of the authors. placed Eupagurus on the Officical Index of Rejected and Invalid Generic Names in Zoology. However, {19} As with C. hirsutimanus, the status and validity of Milne Edwards’ species had been transferred from Kobjakova’s (1955) Clibanarius sachalinicus are Pagurus to Aniculus, then Trizopagurus, and finally uncertain. to Ciliopagurus (Forest, 1995), and was never considered conspecific with Stimpson’s (1860) taxon. {20} Clibanarius willeyi Southwell, 1910 appears to have Consequently, no replacement name was required. been overlooked. It was not mentioned by Fize and Serène (1955) in their monographic treatment of {11} Although Clibanarius antillensis Stimpson, 1859, was Indo-Pacific species or by Gordan (1956) in her the acknowledged junior synonym of C. brasiliensis bibliographic account of pagurids not covered by Dana, 1852a, Forest & de Saint Laurent (1968: 99) Alcock (1905b). Whether it will prove to be a forgotten cited common usage as their reason for selecting senior synonym has yet to be determined. Stimpson’s name. {21} As previously mentioned, although Paul’son (1875) {12} Clibanarius chapini was considered a subspecies of proposed the genus Dardanus for one of the species C. tricolor by Forest & de Saint Laurent (1968), a assigned to Pagurus (sensu Dana), Paul’son’s ranking dismissed by Manning & Chace (1990). genus soon thereafter was placed in synonymy with Dana’s Pagurus. It was not until considerably later {13} The status and validity of Pagurus elongatus H. Milne that Benedict (1896) realized that Dana’s Pagurus Edwards, 1848, are uncertain. H. Milne Edwards contained none of the species originally assigned to the (1848) in his original description said that it was very genus by Fabricius (1775). As Bernhardus had been close to Pagurus tuberculosus H. Milne Edwards, shown to be a synonym of Eupagurus and both species 1836 [= Clibanarius sclopetarius (Herbst, 1791)]; had the same type species originally designated for the type locality was given as the Hogolu Islands, the Pagurus sensu Fabricius, Benedict (1896) urged former name of the Truk Islands of the western Pacific. carcinologists to return to using the name Pagurus However Fize & Serène (1955) list C. tuberculosus for species then assigned to Eupagurus. However, (H. Milne Edwards, 1848) among their Atlanto- while Benedict’s recommendation was accepted by Mediterranean species of Clibanarius. Clibanarius most American and Russian researchers the same was sclopetarius is a western Atlantic species. not true for most European and Asian workers. This nomenclatorial confusion was resolved only after the {14} The status and validity of Kobjakova’s (1971) question of the true interpretation of Pagurus was Clibanarius hirsutimanus are uncertain. brought before the ICZN (Forest & Holthuis, 1955). In its subsequent ruling (Opinion 472), Pagurus sensu {15} Rahayu & Forest (1992) determined that Clibanarius Fabricius and Dardanus Paul’son were placed on the lineatus was not the Indo-Pacific species that it had Official Index of Valid Names in Zoology. been thought to be, but rather H. Milne Edwards’ (1848) Pagurus lineatus collected by Gaudichaud {22} Sakai (1999) suggested that Herbst’s (1791) Cancer off the west coast of South America, and known hungarus might be conspecific with Dardanus regionally as Clibanarius panamensis (Stimpson, calidus; however, the type apparently is no longer 1859). extant, and Herbst’ brief description is inadequate for confirmation. It is questionably included as a {16} Originally described as a subspecies of Clibanarius synonym. aequabilis Dana, 1851, De Man’s Clibanarius aequabilis merguiensis was elevated to full specific {23} Initially described as a distinct species, Glaucothoe rank by Forest (1953) when he determined that C. carinata subsequently was shown to be the megalopal aequabilis was an Atlantic species. stage of a Dardanus species. Its true specific identity is not known with certainty, thus it is included {17} Clibanarius pacificus is another enigmatic species. only questionably in the synonymy of Dardanus According to McLaughlin et al. (2007) the species has deformis. not been accurately reported from anywhere except the type localities of Tanegashiima and Anami-Oshima, {24} McLaughlin et al. (2005: 303) pointed out that Japan. All recent investigations by one of the authors Boone’s species had not been found since the original (TK) failed to locate any specimens fitting Stimpson’s description and suggested that D. cataphractus most (1858) description. probably was actually a specimen of D. guttatus, a species that had been reported occasionally in the {18} In their discussion of Clibanarius rosewateri Manning Hawaiian Islands. and Chace (1990: 37) used the manuscript name C. 25 McLaughlin et al.: Checklist of world Lithodoidea, Lomisoidea and Paguroidea {25} In a study in progress, Manuel Ayon Parente has found {31} Studies in progress by one of the current authors (TK) evidence to indicate that Schmitt’s (1921) Dardanus indicate that Paguristes sinensis Tung & Wang, 1966, jordani is conspecific with Dardanus megistos. may be transferred to the genus Areopaguristes. {26} Gordan (1956) gave the date for Pagurus arrosor var. {32} Henderson (1888) incorrectly used the masculine divergens as 1905 and the pagination as 123–145. De ending “us” in his original description of Paguropsis Melo (1999) gave the date as 1905 and pagination as typica. 1–25; Forest & de Saint Laurent (1968) and Biffar & Provenzano (1972) gave the same pagination, {33} One of the authors (TK) has examined specimens from but a date of 1906. The correct date is 1905 and the Kerara, India and found that Paguropsis andersoni pagination 123–145. (Alcock, 1905b) is not a synonym of P. typica and should be reinstated. {27} As pointed out by McLaughlin & Holthuis (2001), although Dana (1851) specified Pagurus miles as the {34} Holthuis (1959) demonstrated that while this species type of the genus Diogenes, Dana’s species was not was best known as Petrochirus granulatus or P. P. miles of Fabricius (1787). The species had been bahamensis, the correct name was P. diogenes, misidentified by both H. Milne Edwards (1836) and described by Linnaeus (1758) as Cancer diogenes. Dana (1851, 1852a, c). Their P. miles was actually Linnaeus’ (1758) Cancer diogenes actually was Diogenes merguiensis De Man, 1888. While the a composite taxon, based on the mirror image third edition of the ICZN Code required that such a reproductions of Rumphius’ (1705) [1745] illustrations misidentification be referred to the Commission, the of an Indo-Pacific species of Coenobita and Catesby’s fourth edition (1999) gave the revisers the option to fix (1743) description and figure of Cancellus maximus the status of the type species. McLaughlin & Holthuis Bahamensis. Holthuis (1959) selected Catesby’s (2001) were of the opinion that the best interests of figured specimen as the lectotype of Cancer diogenes. stability in nomenclature were served by retaining Herbst’s (1791) Cancer bahamensis was based on Diogenes miles (Fabricius, 1787) [not Diogenes miles Catesby’s same figured specimen, thus the two species sensu H. Milne Edwards, 1836; Dana, 1851, 1852a, were objective synonyms. Holthuis (1959) noted that c] as the type species of the genus Diogenes. the specific name diogenes Linnaeus had not been in use for the past 40 years, so its reintroduction for {28} Holthuis (1959) determined that C. diogenes had the American Petrochirus species would not result been based on two distinct species, one the western in confusion. Herbst’s (1791) species, bahamensis, Atlantic Petrochirus and the other an Indo-Pacific although senior, at the time was better known by species of Coenobita. Holthuis designated the the specific name granulatus. Therefore Holthuis lectotype of Linnaeus’ (1758) composite taxon the (1959) found no reason not to strictly follow in Rule specimen figured by Catesby (1743) of his Cancellus of Priority. maximus Bahamensis, an invalid name for the taxon known as Petrochirus bahamensis (Herbst, 1791). Family Paguridae Latreille, 1802 The species reported in the literature as Pagurus diogenes Fabricius, 1787, Cancer diogenes Herbst, Acanthopagurus de Saint Laurent, 1968 1791 and Diogenes diogenes of subsequent authors was based on misidentifications of Linnaeus’ (1758) = Acanthopagurus de Saint Laurent, 1968 (type Cancer diogenes and was replaced by Diogenes alias McLaughlin & Holthuis, 2001. species Anapagurus? dubius A. Milne-Edwards {29} This species was credited to Costa (1838) by Hope & Bouvier, 1900, by original designation; gender (1851) who provided a description. No description could be found in the portion of Costa’s paper masculine) {1} available. Diogenes serripes (Costa, 1838) is the name and description also given by Carus (1885). The type Acanthopagurus dubius (A. Milne-Edwards & Bouvier, locality is reported as Naples. Although D. serripes has not been cited in any recent synonymy, it most 1900) [Anapagurus] probably is another name for D. pugilator. = Catapaguroides macrophthalmus Bouvier, 1922 {30} Haig et al. (1970) considered P. holmesi a synonym of P. bakeri. However, Glassell’s (1937) specimens Agaricochirus McLaughlin, 1981 can no longer be found, and Hendrickx & Harvey (1999) believe some question exists as to the accuracy = Agaricochirus McLaughlin, 1981 (type species of the synonymy. It is included only questionably in the synonymy of P. bakeri. Pylopagurus boletifer A. Milne-Edwards & Bouvier, 1893, by original designation; gender masculine) Agaricochirus acanthinus McLaughlin, 1982 Agaricochirus alexandri (A. Milne-Edwards & Bouvier, 1893) [Pylopagurus] Agaricochirus boletifer (A. Milne-Edwards & Bouvier, 1893) [Pylopagurus] Agaricochirus cavimanus (Chace, 1939) [Pylopagurus] Agaricochirus echinatus McLaughlin, 1982 26 THE RAFFLES BULLETIN OF ZOOLOGY 2010 Agaricochirus erosus (A. Milne-Edwards, 1880) Anapagurus hyndmanni (Bell, 1846) [Pagurus] {2} [Eupagurus] Agaricochirus gibbosimanus (A. Milne-Edwards, 1880) = Pagurus Hyndmanni Thompson, 1844 (nomen [Eupagurus] Agaricochirus hispidus (Benedict, 1892) [Eupagurus] nudum) Alainopaguroides McLaughlin, 1997 Anapagurus japonicus Ortmann, 1892 = Alainopaguroides McLaughlin, 1997 (type species Alainopaguroides lemaitrei McLaughlin, 1997, by = Anapagurus pusillus var. japonicus Ortmann, original designation; gender masculine) 1892 Alainopaguroides andamanensis McLaughlin, 2002 Alainopaguroides lemaitrei McLaughlin, 1997 Anapagurus laevis (Bell, 1846) [Pagurus] {2} Alainopaguroides megalophthalmus McLaughlin, 2006 Alainopagurus Lemaitre & McLaughlin, 1995 = Pagurus laevis Thompson, 1844 (nomen = Alainopagurus Lemaitre & McLaughlin, 1995 nudum) (type species Alainopagurus crosnieri Lemaitre & McLaughlin, 1995 by original designation; gender Anapagurus longispina A. Milne-Edwards & Bouvier, masculine) 1900 Alainopagurus crosnieri Lemaitre & McLaughlin, 1995 = Anapagurus laevis var. longispina A. Milne- Alloeopagurodes Komai, 1998 Edwards & Bouvier, 1900 = Alloeopagurodes Komai, 1998 (type species Alloeopagurodes spiniacicula Komai, 1998, by Anapagurus petiti Dechancé & Forest, 1962 original designation; gender masculine) Anapagurus smythi Ingle, 1993 Alloeopagurodes spiniacicula Komai, 1998 Anapagrides de Saint Laurent-Dechancé, 1966 Anapagurus vossi García-Gómez, 1994 = Anapagrides de Saint Laurent-Dechancé, 1966 Anapagurus wolffi Forest, 1961 [type species Eupagurus (Spiropagurus) facetus Melin, 1939, by original designation; gender Anisopagurus McLaughlin, 1981 masculine] = Anisopagurus McLaughlin, 1981 (type species = Nanopagurus McLaughlin, 1986 (type species Nanopagurus reesei McLaughlin, 1986, by Eupagurus bartletti A. Milne-Edwards, 1880, by original designation; gender masculine) original designation; gender masculine) Anapagrides aequalis Komai, 1999 Anapagrides facetus (Melin, 1939) [Eupagurus Anisopagurus actinophorus Lemaitre & McLaughlin, (Spiropagurus)] Anapagrides reesei (McLaughlin, 1986) 1996 [Nanopagurus] Anapagurus Henderson, 1886 Anisopagurus bartletti (A. Milne-Edwards, 1880) = Anapagurus Henderson, 1886 [type species Pagurus [Eupagurus] laevis Bell, 1846, by subsequent designation by Holthuis (1962); gender masculine] Anisopagurus hopkinsi Lemaitre & McLaughlin, 1996 Anapagurus adriaticus García-Gómez, 1994 Anisopagurus pygmaeus (Bouvier, 1918) [Eupagurus] Anapagurus alboranensis García-Gómez, 1994 Anapagurus atlantidii García-Gómez, 1994 Anisopagurus vossi Lemaitre & McLaughlin, 1996 Anapagurus bicorniger A. Milne-Edwards & Bouvier, 1892 Bathypaguropsis McLaughlin, 1994 Anapagurus bonnieri Nobili, 1905 Anapagurus breviaculeatus Fenizia, 1937 = Bathypaguropsis McLaughlin, 1994 (type species Anapagurus chiroacanthus (Lilljeborg, 1856) [Pagurus] Bathypaguropsis yaldwyni McLaughlin, 1994, by = Anapagurus brevicarpus A. Milne-Edwards & original designation; gender feminine) Bouvier, 1892 Bathypaguropsis carinatus Komai & Takeda, 2004 = Pagurus ferrugineus Norman, 1861 = Anapagurus chiroacanthus var. cristatus Fenizia, Bathypaguropsis cruentus de Saint Laurent & McLaughlin, 1937 2000 = Anapagurus chiroacanthus var. gracilis Fenizia, Bathypaguropsis foresti Komai & Lemaitre, 2002 1937 Anapagurus congolensis García-Gómez, 1994 Bathypaguropsis kuroshioensis (Miyake, 1978) Anapagurus curvidactylus Chevreux & Bouvier, 1892 Anapagurus hendersoni Barnard, 1947 [Pagurus] = Bathypaguropsis rahayuae McLaughlin, 1997 Bathypaguropsis marionensis McLaughlin, 1994 Bathypaguropsis microps (Balss, 1911) [Eupagurus] Bathypaguropsis yaldwyni McLaughlin, 1994 Boninpagurus Asakura & Tachikawa, 2004 = Boninpagurus Asakura & Tachikawa, 2004 (type species Boninpagurus acanthocheles Asakura & Tachikawa, 2004, by original designation; gender masculine) Boninpagurus acanthocheles Asakura & Tachikawa, 2004 {3} Bythiopagurus McLaughlin, 2003 = Bythiopagurus McLaughlin, 2003 (type species Bythiopagurus macrocolus McLaughlin, 2003 by original designation; gender masculine) Bythiopagurus macrocolus McLaughlin, 2003 Catapaguroides A. Milne-Edwards & Bouvier, 1892 = Catapaguroides A. Milne-Edwards & Bouvier, 1892 [type species Catapaguroides microps A. Milne-Edwards & Bouvier, 1892, by subsequent selection by Holthuis (1962); gender masculine] 27 McLaughlin et al.: Checklist of world Lithodoidea, Lomisoidea and Paguroidea Catapaguroides cristimanus de Saint Laurent, 1968 Catapagurus lewinsohni (Asakura, 2001) Catapaguroides declivis McLaughlin, 1997 [Hemipagurus] Catapaguroides foresti McLaughlin, 2002 Catapagurus maclaughlinae (Asakura, 2001) Catapaguroides fragilis (Melin, 1939) [Eupagurus [Hemipagurus] {9} (Catapagurus)] Catapagurus misakiensis Terao, 1914 Catapaguroides hooveri McLaughlin & Pittman, 2002 = Catapagurus japonicus Yokoya, 1933 Catapaguroides iejimensis Osawa & Takeda, 2004 Catapagurus oculocrassus McLaughlin, 1997 {11} Catapaguroides inermis de Saint Laurent, 1968 Catapagurus sharreri A. Milne-Edwards, 1880 {12} Catapaguroides iris Bouvier, 1922 = Hemipagurus socialis Smith, 1881 Catapaguroides japonicus de Saint Laurent, 1968 = Catapagurus socialis (Smith, 1882) Catapaguroides karubar McLaughlin, 1997 = Catapagurus Sparreri Verrill, 1883 (misspelling Catapaguroides kasei Osawa & Takeda, 2004 {4} of sharreri) Catapaguroides megalops A. Milne-Edwards & Bouvier, = Catapagurus sharrei Abele & Kim, 1986 1892 (misspelling of sharreri) Catapaguroides melini de Saint Laurent, 1968 Catapagurus spinicarpus de Saint Laurent & McLaughlin, Catapaguroides microps A. Milne-Edwards & Bouvier, 2000 1892 {5} Catapagurus tanimbarensis McLaughlin, 1997 Catapaguroides mortenseni de Saint Laurent, 1968 Catapagurus toyoshioae (Asakura, 2001) Catapaguroides olfaciens (Alcock, 1905) [Hemipagurus] [Cestopagurus] Catapagurus tuberculosus (Asakura, 1999) Catapaguroides pectinipes (Lewinsohn, 1969) [Icelopagurus] [Cestopagurus] Catapagurus sp. 1 {7} Catapaguroides setosus Edmondson, 1951 {6} Catapagurus sp. 2 {7} Catapaguroides spinulimanus de Saint Laurent, 1968 Catapagurus sp. 3 {7} Catapaguroides umbra Komai, 2009 Catapagurus sp. 4 {7} Catapaguropsis Lemaitre & McLaughlin, 2006 Catapagurus sp. 5 {7} = Catapaguropsis Lemaitre & McLaughlin, 2006 Ceratopagurus Yokoya, 1933 {13} (type species Catapaguropsis queenslandica = Ceratopagurus Yokoya, 1933 (type species Lemaitre & McLaughlin, 2006, by original Ceratopagurus pilosimanus Yokoya, 1933, by designation; gender feminine) monotypy; gender masculine) Catapaguropsis queenslandica Lemaitre & McLaughlin, Ceratopagurus pilosimanus Yokoya, 1933 2006 Cestopagurus Bouvier, 1897 Catapaguropsis brucei McLaughlin & Lemaitre, 2007 = Cestopagurus Bouvier, 1897 (type species Catapagurus A. Milne-Edwards, 1880 {7} Cestopagurus coutieri Bouvier, 1897, by = Catapagurus A. Milne-Edwards, 1880 (type monotypy; gender masculine) species Catapagurus sharreri A. Milne-Edwards, Cestopagurus coutieri Bouvier, 1897 1880, by monotypy; gender masculine) Cestopagurus puniceus Komai & Takeda, 2005 = Hemipagurus Smith1881 [type species Cestopagurus timidus (Roux, 1830) [Pagurus] Hemipagurus gracilis Smith, 1881, by subsequent = Catapaguroides acanthodes Fenizia, 1935 designation by Asakura (2001); gender = Catapaguroides acutifrons A. Milne-Edwards & masculine] {8} Bouvier, 1892 Catapagurus albatrossae (Asakura, 2001) = Catapaguroides crassipes Fenizia, 1935 [Hemipagurus] = Catapaguroides crassipes var. tuberculatus Catapagurus alcocki McLaughlin, in Hogarth et al., Fenizia, 1937 1998 {9} = Catapaguroides macrochirus Fenizia, 1937 Catapagurus danida McLaughlin, 2002 = Catapaguroides macrochirus var. platichelus Catapagurus ensifer Henderson, 1893 Fenizia, 1937 Catapagurus franklinae McLaughlin, 2004 = Catapaguroides timidus var. neapolitanus Fenizia, Catapagurus gracilis (Smith, 1881) [Hemipagurus] 1937 Catapagurus gracilis var. intermedius A. Milne-Edwards Chanopagurus Lemaitre, 2003 & Bouvier, 1893 {10} = Chanopagurus Lemaitre, 2003 (type species Catapagurus granulatus Edmondson, 1951 Chanopagurus atopos Lemaitre, 2003, by original Catapagurus haigae (Asakura, 2001) [Hemipagurus] designation; gender masculine) Catapagurus hirayamai (Asakura, 2001) Chanopagurus atopos Lemaitre, 2003 [Hemipagurus] Cycetopagurus McLaughlin, 2004 Catapagurus holthuisi McLaughlin, 1997 = Cycetopagurus McLaughlin, 2004 (type species Catapagurus imperialis (Asakura, 2001) Cycetopagurus morgani McLaughlin, 2004, by [Hemipagurus] {11} original designation; gender masculine) Catapagurus insolitus Komai & Osawa, 2009 Cycetopagurus morgani McLaughlin, 2004 Catapagurus kosugei (Asakura, 2001) [Hemipagurus] 28 THE RAFFLES BULLETIN OF ZOOLOGY 2010 Decaphyllus de Saint Laurent, 1968 Enallopaguropsis janetae McLaughlin, 1982 = Decaphyllus de Saint Laurent, 1968 (type species Enallopaguropsis williamsi Lemaitre & McLaughlin, Decaphyllus spinicornis de Saint Laurent, 1968, 2003 by original designation; gender feminine) Enallopagurus McLaughlin, 1981 Decaphyllus barunajaya McLaughlin, 1997 = Enallopagurus McLaughlin, 1981(type species Decaphyllus junquai de Saint Laurent, 1968 Pylopagurus spinicarpus Glassell, 1938, by Decaphyllus maci McLaughlin, 1997 original designation; gender masculine) Decaphyllus similis de Saint Laurent, 1968 Decaphyllus spinicornis de Saint Laurent, 1968 Enallopagurus affinis (Faxon, 1893) [Pylopagurus] Dentalopagurus McLaughlin, 2007 Enallopagurus coronatus (Benedict, 1892) [Eupagurus] = Dentalopagurus McLaughlin, 2007 (type species Enallopagurus provenzanoi Lemaitre & McLaughlin, Dentalopagurus levii McLaughlin, 2007, by 2003 original designation; gender masculine) Enallopagurus spinicarpus (Glassell, 1938) [Pylopagurus] Dentalopagurus levii McLaughlin, 2007 Enneobranchus García-Gómez, 1988 Dentalopagurus rutilocinctus Osawa & Chan, 2008 Diacanthurus McLaughlin & Forest, 1997 = Enneobranchus García-Gómez, 1988 (type species Enneobranchus flavioculatus García-Gómez, 1988, = Diacanthurus McLaughlin & Forest, 1997 (type by original designation; gender masculine) species Eupagurus spinulimanus Miers, 1876, by original designation; gender masculine) Enneobranchus bermudensis García-Gómez, 1988 Enneobranchus flavioculatus García-Gómez, 1988 Diacanthurus ecphyma McLaughlin & Forest, 1997 Enneobranchus markhami García-Gómez, 1988 Diacanthurus ophthalmicus (Ortmann, 1892) Enneopagurus McLaughlin, 1997 [Eupagurus] Diacanthurus richeri McLaughlin & Forest, 1997 = Enneopagurus McLaughlin, 1997 (type species Diacanthurus rubricatus (Henderson, 1888) Enneopagurus garciagomezi McLaughlin, 1997, [Eupagurus] by original designation; gender masculine) Diacanthurus spinulimanus (Miers, 1876) [Eupagurus] Enneopagurus garciagomezi McLaughlin, 1997 = Eupagurus edwardsii Filhol, 1883 Enneophyllus McLaughlin, 1997 = Eupagurus intermedius Lenz, 1901 = Eupagurus norae Chilton, 1911 = Enneophyllus McLaughlin, 1997 (type species = Eupagurus chiltoni E. F. Thompson, 1930 Enneophyllus spinirostris McLaughlin, 1997 by Discorsopagurus McLaughlin, 1974 original designation; gender masculine) = Discorsopagurus McLaughlin, 1974 (type species Enneophyllus spinirostris McLaughlin, 1997 Pylopagurus schmitti Stevens, 1925, by original Forestopagurus García-Gómez, 1995 designation; gender masculine) Discorsopagurus cavicola Komai & Takeda, 1996 = Forestopagurus García-Gómez, 1995 (type species Discorsopagurus maclaughlinae Komai, 1995 Anapagurus drachi Forest, 1966, by original Discorsopagurus schmitti (Stevens, 1925) designation; gender masculine) [Pylopagurus] = Phylopagurus schmitti Hart 1930 (misspelling of Forestopagurus drachi (Forest, 1966) [Anapagurus] Pylopagurus) Goreopagurus McLaughlin, 1988 Discorsopagurus tubicola Komai, 2003 Elassochirus Benedict, 1892 = Goreopagurus McLaughlin, 1988 (type species = Eupagurus (Elassochirus) Benedict, 1892 (type Pagurus piercei Wass, 1963 by original designation; species Bernhardus tenuimanus Dana, 1851, by gender masculine) original designation; gender masculine) Elassochirus cavimanus (Miers, 1879) [Eupagurus] Goreopagurus garthi McLaughlin & Haig, 1995 = Eupagurus gotoi Terao, 1913 Goreopagurus piercei (Wass, 1963) [Pagurus] = Eupagurus (Elassochirus) munitus Benedict, Goreopagurus poorei Lemaitre & McLaughlin, 2003 1892 Goreopagurus lemaitrei Nucci & de Melo, 2007 Elassochirus gilli (Benedict, 1892) [Eupagurus] Hachijopagurus Osawa & Okuno, 2003 = Eupagurus porcellanus Molander, 1914 Elassochirus tenuimanus (Dana, 1851) [Bernhardus] = Hachijopagurus Osawa & Okuno, 2003 (type Enallopaguropsis McLaughlin, 1981 species Hachijopagurus rubrimaculatus Osawa = Enallopaguropsis McLaughlin, 1981 (type species & Okuno, 2003, by original designation; gender Pylopagurus guatemoci Glassell, 1937, by original masculine) designation; gender feminine) Enallopaguropsis guatemoci (Glassell, 1937) Hachijopagurus rubrimaculatus Osawa & Okuno, [Pylopagurus] 2003 = Pylopagurus hancocki Walton, 1954 Haigiopagurus McLaughlin, 2005 = Haigia McLaughlin, 1981(type species Pylopagurus diegensis Scanland & Hopkins, 1969 by original designation; gender masculine) (name preoccupied by Haigia Steyskal, 1961, Insecta) = Haigiopagurus McLaughlin, 2005 (replacement name) Haigiopagurus diegensis (Scanland & Hopkins, 1969) [Pylopagurus] 29 McLaughlin et al.: Checklist of world Lithodoidea, Lomisoidea and Paguroidea Icelopagurus McLaughlin, 1997 Lophopagurus (Australeremus) stewarti (Filhol, 1883) = Icelopagurus McLaughlin, 1997 (type species [Eupagurus] Icelopagurus crosnieri McLaughlin, 1997, by Lophopagurus (Australeremus) triserratus (Ortmann, original designation; gender masculine) 1892) [Eupagurus] Icelopagurus crosnieri McLaughlin, 1997 = Pylopagurus serpulophilus Miyake, 1978 Icelopagurus undulatus McLaughlin, 2006 Lophopagurus (Lophopagurus) McLaughlin, 1981 Iridopagurus de Saint Laurent-Dechancé, 1966 = Lophopagurus McLaughlin, 1981 (type species = Iridopagurus de Saint Laurent-Dechancé, 1966 Eupagurus thompsoni Filhol, 1885, by original (type species Spiropagurus iris A. Milne- designation; gender masculine) Edwards, 1880, by original designation; gender Lophopagurus (Lophopagurus) foresti McLaughlin & masculine) Gunn, 1992 Iridopagurus caribbensis (A. Milne-Edwards & Bouvier, Lophopagurus (Lophopagurus) lacertosus (Henderson, 1893) [Spiropagurus] 1888) [Eupagurus] {15} Iridopagurus dispar (Stimpson, 1859) [Spiropagurus] = Eupagurus crenatus Borradaile, 1916 = Spiropagurus dispar Stimpson, 1858 (nomen Lophopagurus (Lophopagurus) nanus (Henderson, 1888) nudum) [Eupagurus] Iridopagurus globulus de Saint Laurent-Dechancé, Lophopagurus (Lophopagurus) nodulosus McLaughlin 1966 & Gunn, 1992 Iridopagurus haigae García-Gómez, 1983 Lophopagurus (Lophopagurus) pumilus de Saint Laurent Iridopagurus iris (A. Milne-Edwards, 1880) & McLaughlin, 2000 [Spiropagurus] Lophopagurus (Lophopagurus) thompsoni (Filhol, 1885) Iridopagurus margaritensis García-Gómez, 1983 [Eupagurus] Iridopagurus occidentalis (Faxon, 1893) = Eupagurus thomsoni Thomson, 1898 (misspelling [Spiropagurus] of thompsoni) Iridopagurus reticulatus García-Gómez, 1983 Manucomplanus McLaughlin, 1981 Iridopagurus violaceus de Saint Laurent-Dechancé, = Manucomplanus McLaughlin, 1981 (type species 1966 Eupagurus ungulatus Studer, 1883, by original Labidochirus Benedict, 1892 designation; gender masculine) = Eupagurus Labidochirus Benedict, 1892 (type Manucomplanus cervicornis (Benedict, 1892) species Pagurus splendescens Owen, 1839, by [Eupagurus] original designation; gender masculine) Manucomplanus longimanus (Faxon, 1893) = Libidochirus Benedict, 1892 (misspelling of [Pylopagurus] Labidochirus) Manucomplanus spinulosus (Holthuis, 1959) Labidochirus anomalus (Balss, 1913) [Eupagurus] [Pylopagurus] Labidochirus splendescens (Owen, 1839) [Pagurus] = Pagurus impressus zilchi Türkay, 1968 Lithopagurus Provenzano, 1968 Manucomplanus ungulatus (Studer, 1883) [Eupagurus] = Lithopagurus Provenzano, 1968 (type species = Eupagurus corallinus Benedict, 1892 Lithopagurus yucatanicus Provenzano, 1968, by Manucomplanus varians (Benedict, 1892) [Eupagurus] original designation; gender masculine) Michelopagurus McLaughlin, 1997 Lithopagurus boucheti McLaughlin & Lemaitre, 2004 = Michelopagurus McLaughlin, 1997 (type species Lithopagurus tribulomanus McLaughlin & Lemaitre, Pagurodes limatulus Henderson, 1888, by original 2004 designation; gender masculine) Lithopagurus yucatanicus Provenzano, 1968 Michelopagurus atlanticus (Bouvier, 1922) Lophopagurus (Australeremus) McLaughlin, 1981 [Pagurodes] = Australeremus McLaughlin, 1981 (type species Michelopagurus chacei McLaughlin, 1997 Eupagurus cookii Filhol, 1883, by original Michelopagurus limatulus (Henderson, 1888) designation; gender masculine) [Pagurodes] Lophopagurus (Australeremus) cookii (Filhol, 1883) Michelopagurus richardi (Bouvier, 1922) [Pagurodes] [Eupagurus] Micropagurus McLaughlin, 1986 Lophopagurus (Australeremus) cristatus (H. Milne = Micropagurus McLaughlin, 1986 (type species Edwards, 1836) [Pagurus] Micropagurus devaneyi McLaughlin, 1986, by Lophopagurus (Australeremus) eltaninae (McLaughlin original designation; gender masculine) & Gunn, 1992) [Australeremus] Micropagurus n. sp. Lemaitre, 2010 (in press) Lophopagurus (Australeremus) indonesiensis Micropagurus acantholepis (Stimpson, 1858) (McLaughlin, 1997) [Australeremus] {14} [Eupagurus] Lophopagurus (Australeremus) kirkii (Filhol, 1883) = Anapagurus australiensis Henderson, 1888 [Eupagurus] Micropagurus devaneyi McLaughlin, 1986 Lophopagurus (Australeremus) laurentae (McLaughlin Micropagurus polynesiensis (Nobili, 1906) & Gunn, 1992) [Australeremus] [Anapagurus] = Micropagurus vexatus Haig & Ball, 1988 30 THE RAFFLES BULLETIN OF ZOOLOGY 2010 Micropagurus propinquus Asakura, 2005 Nematopagurus vallatus (Melin, 1939) [Eupagurus Micropagurus spinimanus Asakura, 2005 Munidopagurus A. Milne-Edwards & Bouvier, 1893 (Catapagurus)] = Munidopagurus A. Milne Edwards & Bouvier, Orthopagurus Stevens, 1927 1893 (type species Eupagurus macrocheles A. Milne-Edwards, 1880, by monotypy; genus = Orthopagurus Stevens, 1927 [type species Pagurus masculine) minimus Holmes, 1900; by subsequent designation Munidopagurus macrocheles (A. Milne-Edwards, 1880) [Eupagurus] by Makarov (1938); gender masculine] Nematopaguroides Forest & de Saint Laurent, 1968 = Orthopaguropsis Serène, 1957 (type species = Nematopaguroides Forest & de Saint Laurent, 1968 (type species Nematopaguroides fagei Forest & Orthopagurus harmsi Gordon 1935, by original de Saint Laurent, 1968, by original designation; gender masculine) designation; gender feminine) Nematopaguroides fagei Forest & de Saint Laurent, Orthopagurus minimus (Holmes, 1900) [Pagurus] 1968 Nematopaguroides pusillus Forest & de Saint Laurent, Ostraconotus A. Milne-Edwards, 1880 1968 Nematopagurus A. Milne-Edwards & Bouvier, 1892 = Ostraconotus A. Milne-Edwards, 1880 (type = Nematopagurus A. Milne-Edwards & Bouvier, species Ostraconotus spatulipes A. Milne- 1892 (type species Nematopagurus longicornis A. Milne-Edwards & Bouvier, 1892, by monotypy; Edwards, 1880 by monotypy; gender masculine) gender masculine) Ostraconotus spatulipes A. Milne-Edwards, 1880 Nematopagurus alcocki McLaughlin, 1997 Nematopagurus australis (Henderson, 1888) Paguridium Forest, 1961 [Catapagurus] Nematopagurus chanani McLaughlin, 2004 = Paguridium Forest, 1961 (type species Eupagurus Nematopagurus chauseyensis McLaughlin, 1998 Nematopagurus crosnieri McLaughlin, 1998 ? minimum Chevreux & Bouvier, 1892, by original Nematopagurus diadema Lewinsohn, 1969 Nematopagurus gardineri Alcock, 1905 designation; gender neuter) {16} = Nematopagurus holthuisi McLaughlin & Hogarth, Paguridium minimum (Chevreux & Bouvier, 1892) 1998 [Eupagurus] = Nematopagurus pilosus Komai, 1999 Nematopagurus helleri (Balss, 1916) [Cestopagurus] Paguritta Melin, 1939 Nematopagurus indicus Alcock, 1905 Nematopagurus jacquesi McLaughlin, 2004 = Paguritta Melin, 1939 (type species Eupagurus Nematopagurus kosiensis McLaughlin, 1998 (Paguritta) gracilipes Melin, 1939, by monotypy; = Nematopagurus shinnyoae Komai, 1999 Nematopagurus lepidochirus (Doflein, 1902) gender feminine) [Eupagurus] Nematopagurus lewinsohni Türkay, 1986 Paguritta corallicola Lewinsohn, 1978 Nematopagurus sp. McLaughlin & Okuno, 2010 (in press) Paguritta gracilipes (Melin, 1939) [Eupagurus Nematopagurus longicornis A. Milne-Edwards & Bouvier, 1892 (Paguritta)] Nematopagurus meiringae McLaughlin, 1998 Nematopagurus muricatus (Henderson, 1896) Paguritta harmsi (Gordon, 1935) [Orthopagurus] [Catapagurus] Nematopagurus ostlingochirus McLaughlin, 1997 Paguritta kroppi McLaughlin & Lemaitre, 1993 Nematopagurus ricei McLaughlin, 2004 Nematopagurus richeri McLaughlin, 2004 Paguritta morgani McLaughlin & Lemaitre, 1993 Nematopagurus scutellichelis Alcock, 1905 Nematopagurus scutelliformis McLaughlin, 1997 Paguritta scottae McLaughlin & Lemaitre, 1993 Nematopagurus spinulosensoris McLaughlin & Brock, 1974 Paguritta vittata Komai & Nishi, 1996 Nematopagurus spongioparticeps McLaughlin, 2004 Nematopagurus squamichelis Alcock, 1905 Pagurixus Melin, 1939 Nematopagurus tricarinatus (Stimpson, 1858) [Eupagurus] = Pagurixus Melin, 1939 (type species Eupagurus (Pagurixus) boninensis Melin, 1939, by monotypy; gender masculine) Pagurixus acanthocarpus Komai & Okuno, 2009 Pagurixus amsa Morgan, 1993 Pagurixus anceps (Forest, 1957) [Eupagurus] Pagurixus aurantiaca Komai, 2010 Pagurixus boninensis (Melin, 1939) [Eupagurus (Pagurixus)] Pagurixus brachydactylus Komai & Osawa, 2006 Pagurixus carinimanus Komai & Osawa, 2006 Pagurixus cavicarpus Komai, 2010 Pagurixus concolor Komai & Osawa, 2006 Pagurixus crosnieri Komai, 2010 Pagurixus dissimilis Osawa & Komai, 2007 Pagurixus fasciatus Komai & Myorin, 2005 Pagurixus festinus McLaughlin & Haig, 1984 Pagurixus formosus Komai, 2010 Pagurixus granulimanus Morgan, 1993 Pagurixus haigae Komai & Osawa, 2007 Pagurixus handrecki Gunn & Morgan, 1992 Pagurixus hectori (Filhol, 1883) [Eupagurus] = Eupagurus campbelli Filhol, 1885 Pagurixus icelus Komai, 2010 Pagurixus jerviensis McLaughlin & Haig, 1984 31 McLaughlin et al.: Checklist of world Lithodoidea, Lomisoidea and Paguroidea Pagurixus kermadecensis de Saint Laurent & McLaughlin, Pagurus alaini Komai, 1998 2000 Pagurus alatus Fabricius, 1775 {18} Pagurixus laevimanus (Ortmann, 1892) [Eupagurus] = Pagurus tricarinatus Norman, 1869 Pagurixus longipes Osawa, Fujita & Okuno, 2006 = Eupagurus variabilis A. Milne-Edwards & Pagurixus maorus (Nobili, 1906) [Eupagurus] Bouvier, 1892 Pagurixus nomurai Komai & Asakura, 1995 Pagurus albidianthus de Saint Laurent & McLaughlin, Pagurixus patiae Komai, 2006 2000 Pagurixus paulayi Komai & Osawa, 2006 Pagurus albus (Benedict, 1892) [Eupagurus] Pagurixus pilosus Komai, 2010 = Pagurus fuscomaculatus Bouvier, 1895 Pagurixus pseliophorus Komai & Osawa, 2006 Pagurus alcocki (Balss, 1911) [Eupagurus] Pagurixus pulcher Osawa, Fujita & Okuno, 2006 Pagurus aleuticus (Benedict, 1892) [Eupagurus] Pagurixus purpureus Komai & Okuno, 2009 Pagurus anachoretoides Forest, 1966 Pagurixus ruber Komai & Osawa, 2006 Pagurus anachoretus Risso, 1827 Pagurixus rubrovittatus Komai, 2010 = Pagurus pictus H. Milne Edwards, 1836 Pagurixus sculptus Komai, 2010 = Clibanarius mediterraneus Kossmann, 1878 Pagurixus stenops Morgan, 1993 Pagurus angustus (Stimpson, 1858) [Eupagurus] Pagurixus tweediei (Forest, 1956) [Eupagurus] Pagurus annexus McLaughlin & Haig, 1993 Pagurodes Henderson, 1888 Pagurus annulipes (Stimpson, 1860) [Eupagurus] = Pagurodes Henderson, 1888 [type species Pagurus arcuatus Squires, 1964 Pagurodes inarmatus Henderson, 1888, by = Pagurus bankensis Nesis, 1964 subsequent designation by de Saint Laurent (1969); Pagurus arenisaxatilis Harvey & McLaughlin, 1991 gender masculine] Pagurus armatus (Dana, 1851) [Bernhardus] Pagurodes inarmatus Henderson, 1888 Pagurus benedicti (Bouvier, 1898) [Eupagurus] Pagurodofleinia Asakura 2005 = Eupagurus minutus Benedict, 1892 (preoccupied = Dofleinia McLaughlin & Asakura, 2004 (type name) species Catapagurus doederleini Doflein 1902, = Nympagurus galapagensis Boone, 1932 by original designation; gender feminine) (name Pagurus beringanus (Benedict, 1892) [Eupagurus] preoccupied by Dofleinia Wassifieff, 1908, = Eupagurus newcombei Benedict, 1892 Cnidaria) Pagurus bernhardus (Linnaeus, 1758) [Cancer] = Pagurodofleinia Asakura, 2005 (replacement = Bernhardus typicus Dana, 1851 name) = Pagurus ulidiae Thompson, 1844 (nomen Pagurodofleinia doederleini (Doflein, 1902) nudum) [Catapagurus] = Pagurus ulidanus Bell, 1846 {19} Pagurojacquesia de Saint Laurent & McLaughlin, 2000 = Pagurus eblaniensis Kinahan, 1860 = Jaquesia de Saint Laurent & McLaughlin, Pagurus boriaustraliensis Morgan, 1990 1999 (type species Jacquesia polymorpha de Pagurus bouvieri (Faxon, 1895) [Eupagurus] Saint Laurent & McLaughlin, 1999, by original = Eupagurus smithi A. Milne-Edwards & Bouvier, designation; gender feminine) (name preoccupied 1893 (preoccupied name) by Jacquesia Mendes, 1944, Mollusca) Pagurus brachiomastus (Thallwitz, 1892) [Eupagurus] = Pagurojacquesia de Saint Laurent & McLaughlin, Pagurus brandti (Benedict, 1892) [Eupagurus] 2000 (replacement name) Pagurus brevidactylus (Stimpson, 1859) [Eupagurus] Pagurojacquesia polymorpha (de Saint Laurent & = Pagurus miamensis Provenzano, 1959 McLaughlin, 1999) [Jacquesia] = Pagurus miamensis uncifer Forest & de Saint Pagurus Fabricius, 1775 {17} Laurent, 1968 = Pagurus Fabricius, 1775 [type species by Pagurus bullisi Wass, 1963 subsequent selection by Latreille (1810) Cancer Pagurus capillatus (Benedict, 1892) [Eupagurus] bernhardus Linnaeus, 1758 as defined by lectotype Pagurus capsularis McLaughlin, 1997 selection by Forest & Holthuis (1955): specimen Pagurus carneus (Pocock, 1889) [Eupagurus] figured by Swammerdam, 1737, ICZN Opinion Pagurus carolinensis McLaughlin, 1975 472] Pagurus carpoforaminatus (Alcock, 1905) [Eupagurus] = Pagures Ordinaires H. Milne Edwards, 1836 (not Pagurus caurinus Hart, 1971 in Latin, invalid name) Pagurus cavicarpus (Paul’son, 1875) [Eupagurus] = Pagures Ordinaires, Dextres H. Milne Edwards, = Eupagurus carpoforaminatus var. nephromma 1848 (not in Latin, invalid name) Alcock, 1905 = Eupagurus Brandt, 1851 [type species by Pagurus chevreuxi (Bouvier, 1896) [Eupagurus] subsequent selection by Stimpson (1858) Cancer Pagurus chinensis White, 1847 (nomen nudum) {20} bernhardus Linnaeus, 1758] Pagurus compressipes (Miers, 1884) [Eupagurus] = Bernhardus Dana, 1851 (type species Bernhardus Pagurus comptus White, 1847 {21} typicus Dana, 1851 by original designation) = Pagurus gayi Nicolet, 1849 Pagurus acadianus Benedict, 1901 32 THE RAFFLES BULLETIN OF ZOOLOGY 2010 Pagurus conformis De Haan, 1849 Pagurus ikedai Lemaitre & Watabe, 2005 Pagurus imafukui McLaughlin & Konishi, 1994 = Eupagurus megalops Stimpson, 1858 Pagurus imaii (Yokoya, 1939) [Eupagurus] Pagurus imarpe Haig, 1974 Pagurus confragosus (Benedict, 1892) [Eupagurus] Pagurus impressus (Benedict, 1892) [Eupagurus] Pagurus indicus Sarojini & Nagabhushanam, 1972 Pagurus confusus Komai & Yu, 1999 Pagurus inermis (Chevreux & Bouvier, 1892) [Eupagurus] Pagurus constans (Stimpson, 1858) [Eupagurus] Pagurus insulae Asakura, 1991 Pagurus investigatoris (Alcock, 1905) [Eupagurus] = Pagurus sagamiensis Miyake, 1978 Pagurus iridocarpus de Saint Laurent & McLaughlin, 2000 Pagurus cornutus (Benedict, 1892) [Eupagurus] Pagurus irregularis (A. Milne-Edwards & Bouvier, 1892) [Eupagurus] Pagurus criniticornis (Dana, 1852) [Bernhardus] Pagurus isochirus White, 1847 (nomen nudum) {20} Pagurus japonicus (Stimpson, 1858) [Eupagurus] Pagurus cuanensis Bell, 1846 {19} = Eupagurus barbatus Ortmann, 1892 = Pagurus Cuanensis Thompson, 1844 (nomen Pagurus kaiensis McLaughlin, 1997 Pagurus kennerlyi (Stimpson, 1864) [Eupagurus] nudum) Pagurus kulkarnii Sankolli, 1962 Pagurus lanuginosus De Haan, 1849 = Pagurus spinimanus Lucas, 1846 Pagurus laurentae Forest, 1978 Pagurus lepidus (Bouvier, 1898) [Eupagurus] = Pagurus lucasi Heller, 1863 Pagurus leptonyx Forest & de Saint Laurent, 1968 = Eupagurus chiereghini Nardo, 1868 = Pagurus leptonix Fausto-Filho, 1970 (misspelling of leptonyx) = Eupagurus placens Stebbing, 1924 Pagurus limatulus Fausto Filho, 1970 Pagurus curacaoensis (Benedict, 1892) [Eupagurus] Pagurus liochele (Barnard, 1947) [Pylopagurus] Pagurus dalli (Benedict, 1892) [Eupagurus] = Pagurus barnardi Forest, 1966 Pagurus longicarpus Say, 1817 Pagurus dartevellei (Forest, 1958) [Pylopagurus] = Pagurus truncatulus Rafinesque, 1817 Pagurus decimbranchiae Komai & Osawa, 2001 Pagurus longimanus Wass, 1963 Pagurus lophochela Komai, 1999 Pagurus defensus (Benedict, 1892) [Eupagurus] Pagurus luticola Komai & Chan, 2006 Pagurus macardlei (Alcock, 1905) [Eupagurus] Pagurus delsolari Haig, 1974 Pagurus maclaughlinae García-Gómez, 1982 Pagurus maculosus Komai & Imafuku, 1996 Pagurus dissimilis (A. Milne-Edwards & Bouvier, 1893) Pagurus marshi Benedict, 1901 [Eupagurus] = Pagurus marschi Forest & de Saint Laurent, 1968 (misspelling of marshi) Pagurus edwardsii (Dana, 1852) [Bernhardus] Pagurus mbizi (Forest, 1955) [Eupagurus] Pagurus emmersoni McLaughlin & Forest, 1999 Pagurus meloi Lemaitre & Cruz Castaño, 2004 Pagurus mertensii Brandt, 1851 Pagurus erythrogrammus Komai, 2003 Pagurus middendorffii Brandt, 1851 Pagurus minutus Hess, 1865 Pagurus excavatus (Herbst, 1791) [Cancer] {18} = Eupagurus dubius Ortmann, 1892 = Pagurus angulatus Risso, 1816 Pagurus moluccensis Haig & Ball, 1988 Pagurus nanodes Haig & Harvey, 1991 = Eupagurus meticulosus Roux, 1830 Pagurus nesiotes Haig & McLaughlin, 1991 Pagurus nigrivittatus Komai, 2003 Pagurus exiguus (Melin, 1939) [Eupagurus Pagurus nigrofascia Komai, 1996 Pagurus nipponensis (Yokoya 1933) [Eupagurus] (Pagurillus)] Pagurus nisari Siddiqui & Komai, 2008 Pagurus novizealandiae (Dana, 1852) [Bernhardus] Pagurus exilis (Benedict, 1892) [Eupagurus] Pagurus ochotensis Brandt, 1851 Pagurus filholi (De Man, 1887) [Eupagurus] = Pagurus (Eupagurus) bernhardus var. B. granulato-denticulata Brandt, 1851 = Pagurus geminus McLaughlin, 1976 = Eupagurus alaskensis Benedict, 1892 Pagurus fimbriatus Forest, 1966 = Eupagurus ortmanni Balss, 1911 Pagurus parvispina Komai, 1997 Pagurus forbesii Bell, 1846 {19} Pagurus forceps H. Milne Edwards, 1836 {21} = Eupagurus comptus var. jugosa Henderson, 1888 = Eupagurus comptus var. latimanus Miers, 1875 Pagurus fungiformis Komai & Rahayu, 2004 Pagurus fuscomaculatus (Bouvier, 1895) [Eupagurus] Pagurus gladius (Benedict, 1892) [Eupagurus] Pagurus gordonae (Forest, 1956) [Eupagurus] Pagurus gracilipes (Stimpson, 1858) [Eupagurus] Pagurus granosimanus (Stimpson, 1859) [Eupagurus] Pagurus gymnodactylus Lemaitre, 1982 Pagurus hartae (McLaughlin & Jensen, 1996) [Parapagurodes] Pagurus heblingi Nucci & de Melo, 2003 Pagurus hedleyi (Grant & McCulloch, 1906) [Eupagurus] (replacement name for Eupagurus kirkii Miers, 1884, preoccupied by Eupagurus kirkii Filhol 1883) Pagurus hemphilli (Benedict, 1892) [Eupagurus] Pagurus hirsutiusculus (Dana, 1851) [Bernhardus] Pagurus hirtimanus (Miers, 1880) [Eupagurus] = Pagurus hirtimanus White, 1847 (nomen nudum) = Eupagurus janitor Alcock, 1905 Pagurus holmi Ng & McLaughlin, 2009 33 McLaughlin et al.: Checklist of world Lithodoidea, Lomisoidea and Paguroidea Pagurus parvus (Benedict, 1892) [Eupagurus] Pagurus stimpsoni (A. Milne-Edwards & Bouvier, 1893) [Eupagurus] Pagurus pectinatus (Stimpson, 1858) [Eupagurus] = Pagurus bonairensis Schmitt, 1936 = Eupagurus seriespinosus Thallwitz, 1891 = Pagurus hendersoni Wass, 1963 Pagurus tanneri (Benedict, 1892) [Eupagurus] = Clibanarius japonicus Rathbun, 1903 Pagurus townsendi (Benedict, 1892) [Eupagurus] Pagurus traversi (Filhol, 1885) [Eupagurus] Pagurus pergranulatus (Henderson, 1896) [Eupagurus] Pagurus triangularis (Chevreux & Bouvier, 1892) [Eupagurus] Pagurus perlatus H. Milne Edwards, 1848 Pagurus trichocerus Forest & de Saint Laurent, 1968 Pagurus trigonocheirus (Stimpson, 1858) [Eupagurus] = Bernhardus obesocarpus Dana, 1852 Pagurus tristanensis (Henderson, 1888) [Eupagurus] Pagurus undosus (Benedict, 1892) [Eupagurus] Pagurus pilosimanus White, 1847 (nomen nudum) = Eupagurus trigonocheirus var. paulensis Balss, {20} 1913 Pagurus venturensis Coffin, 1957 Pagurus pilosipes (Stimpson, 1858) [Eupagurus] = Pagurus hirsutiusculus venturensis Coffin 1957 Pagurus pilosiusculus White, 1847 (nomen nudum) Pagurus vetaultae Harvey & McLaughlin, 1991 Pagurus villosus Nicolet, 1849 {20} Pagurus virgulatus Haig & Harvey, 1991 Parapagurodes McLaughlin & Haig, 1973 Pagurus pitagsaleei McLaughlin, 2002 = Parapagurodes McLaughlin & Haig, 1973 (type Pagurus politus (Smith, 1882) [Eupagurus] species Parapagurodes makarovi McLaughlin & Haig, 1973, by original designation; gender Pagurus pollexcavus Glassell, 1937 uncertain status masculine) [22] Parapagurodes laurentae McLaughlin & Haig, 1973 Parapagurodes makarovi McLaughlin & Haig, 1973 Pagurus pollicaris Say, 1817 Phimochirus McLaughlin, 1981 = Eupagurus floridanus Benedict, 1892 = Phimochirus McLaughlin, 1981 (type species Eupagurus operculatus Stimpson, 1859, by Pagurus prideaux Leach, 1815 original designation; gender masculine) Pagurus protuberocarpus McLaughlin, 1982 Phimochirus californiensis (Benedict, 1892) [Eupagurus] Pagurus provenzanoi Forest & de Saint Laurent, 1968 = Eupagurus mexicanus Benedict, 1892 Pagurus proximus Komai, 2000 Phimochirus holthuisi (Provenzano, 1961) [Pylopagurus] Pagurus pubescens Kröyer, 1838 Phimochirus leurocarpus McLaughlin, 1981 Phimochirus occlusus (Henderson, 1888) [Eupagurus] = Pagurus thompsoni Bell, 1853 {15} Phimochirus operculatus (Stimpson, 1859) [Eupagurus] = Eupagurus kroyeri Stimpson, 1859 Pylopagurus samariensis Sanchez, 1978 Phimochirus randalli (Provenzano, 1961) Pagurus pubescentulus (A. Milne-Edwards & Bouvier [Pylopagurus] Phimochirus roseus (Benedict, 1892) [Eupagurus] 1892) [Eupagurus] Phimochirus venustus (Bouvier, 1898) [Eupagurus] Porcellanopagurus Filhol, 1885 = Eupagurus variabilis var. charcoti Bouvier, = Porcellanopagurus Filhol, 1885 (type species 1914 Porcellanopagurus edwardsi Filhol, 1885 by monotypy; gender masculine) Pagurus pulchellus (A. Milne-Edwards & Bouvier, 1892) Porcellanopagurus adelocercus McLaughlin & Hogarth, [Eupagurus] 1998 Porcellanopagurus belauensis Suzuki & Takeda, 1987 Pagurus quaylei Hart, 1971 Porcellanopagurus chiltoni de Saint Laurent & McLaughlin, 2000 Pagurus quinquelineatus Komai, 2003 Porcellanopagurus edwardsi Filhol, 1885 Porcellanopagurus filholi de Saint Laurent & McLaughlin, Pagurus rathbuni (Benedict, 1892) [Eupagurus] 2000 Porcellanopagurus foresti Zarenkov, 1990 = Eupagurus (Trigonocheirus) polaris Sivertsen, Porcellanopagurus haptodactylus McLaughlin, 2000 1932 Pagurus redondoensis Wicksten, 1982 Pagurus retrorsimanus Wicksten & McLaughlin, 1998 Pagurus rhabdotus Haig & Harvey, 1991 Pagurus rotundimanus Wass, 1963 Pagurus ruber (A. Milne-Edwards & Bouvier, 1892) [Eupagurus] Pagurus rubrior Komai, 2003 Pagurus samoensis (Ortmann, 1892) [Eupagurus] Pagurus samuelis (Stimpson, 1857) [Eupagurus] Pagurus setosus (Benedict, 1892) [Eupagurus] Pagurus similimanus (Balss, 1921) [Eupagurus (?Anapagurus)] Pagurus similis (Ortmann, 1892) [Eupagurus] Pagurus simulans Komai, 2000 Pagurus sinuatus (Stimpson, 1858) [Eupagurus] Pagurus smithi (Benedict, 1892) [Eupagurus] Pagurus souriei (Forest, 1952) [Eupagurus] Pagurus spighti McLaughlin & Haig, 1993 Pagurus spilocarpus Haig, 1977 Pagurus spina Komai, 1994 Pagurus spinulentus (Henderson, 1888) [Eupagurus] Pagurus stevensae Hart, 1971 Pagurus sticticus McLaughlin, 2008 34 THE RAFFLES BULLETIN OF ZOOLOGY 2010 Porcellanopagurus jacquesi McLaughlin, 1997 Pylopaguridium markhami McLaughlin & Lemaitre, Porcellanopagurus japonicus Balss, 1913 2001 Porcellanopagurus nihonkaiensis Takeda, 1985 Pylopaguropsis Alcock, 1905 Porcellanopagurus platei Lenz, 1902 Porcellanopagurus tridentatus Whitelegge, 1900 = Pylopaguropsis Alcock, 1905 (type species Porcellanopagurus truncatifrons Takeda, 1981 Pylopagurus magnimanus Henderson, 1896, by Propagurus McLaughlin & de Saint Laurent, 1998 monotypy; gender feminine) = Propagurus McLaughlin & de Saint Laurent, 1998 = Galapagurus Boone, 1932 (type species (type species Pagurus gaudichaudii H. Milne Galapagurus teevana Boone, 1932, by monotypy; Edwards, 1836, by original designation; gender gender masculine) masculine) Pylopaguropsis atlantica Wass, 1963 Propagurus deprofundis (Stebbing, 1924) [Eupagurus] Pylopaguropsis bellula Osawa & Okuno, 2006 Propagurus gaudichaudii (H. Milne Edwards, 1836) Pylopaguropsis fimbriata McLaughlin & Haig, 1989 [Pagurus] Pylopaguropsis furusei Asakura, 2000 Pylopaguropsis garciai McLaughlin & Haig, 1989 = Bernhardus barbiger A. Milne-Edwards, 1891 Pylopaguropsis granulata Asakura, 2000 Propagurus haigae (McLaughlin, 1997) [Pagurus] Pylopaguropsis keijii McLaughlin & Haig, 1989 Propagurus miyakei (Baba, 1986) [Pagurus] Pylopaguropsis laevispinosa McLaughlin & Haig, Propagurus obtusifrons (Ortmann, 1892) [Eupagurus] 1989 Pylopaguropsis lemaitrei Asakura & Paulay, 2003 = Pagurus yokoyai Makarov, 1938 Pylopaguropsis lewinsohni McLaughlin & Haig, 1989 Protoniopagurus Lemaitre & McLaughlin, 1996 Pylopaguropsis magnimanus (Henderson, 1896) [Pylopagurus] = Protoniopagurus Lemaitre & McLaughlin, 1996 Pylopaguropsis pustulosa McLaughlin & Haig, 1989 (type species Protoniopagurus bioperculatus Pylopaguropsis speciosa McLaughlin & Haig, 1989 Lemaitre & McLaughlin, 1996, by original Pylopaguropsis teevana (Boone, 1932) [Galapagurus] designation; gender masculine) Pylopaguropsis vicina Komai & Osawa, 2004 Pylopaguropsis zebra (Henderson, 1893) [Eupagurus] Protoniopagurus bioperculatus Lemaitre & McLaughlin, Pylopagurus A. Milne-Edwards & Bouvier, 1893 1996 Pseudopagurodes McLaughlin, 1997 = Pylopagurus A. Milne-Edwards & Bouvier, 1893 [type species Eupagurus discoidalis A. Milne- = Pseudopagurodes McLaughlin, 1997 (type species Edwards, 1880, by subsequent designation by Pagurodes piliferus Henderson, 1888, by original Miyake (1978); gender masculine] designation; gender masculine) Pylopagurus discoidalis (A. Milne-Edwards, 1880) Pseudopagurodes piliferus (Henderson, 1888) [Eupagurus] [Pagurodes] Pylopagurus gorei McLaughlin & Lemaitre, 2001 Pseudopagurodes reconditus (Wang & McLaughlin, Pylopagurus holmesi Schmitt, 1921 2000) [Nematopaguroides] Pteropagurus McLaughlin & Rahayu, 2006 = Pylopagurus longicarpus Walton, 1954 Pylopagurus macgeorgei McLaughlin & Lemaitre, = Pteropagurus McLaughlin & Rahayu, 2006 (type 2001 species Pteropagurus inermis McLaughlin & Pylopagurus pattiae Lemaitre & Campos, 1993 Rahayu, 2006 by original designation; gender Rhodochirus McLaughlin, 1981 masculine) = Rhodochirus McLaughlin, 1981 (type species Pteropagurus inermis McLaughlin & Rahayu, 2006 Pylopagurus rosaceus A. Milne-Edwards & Pteropagurus spina McLaughlin & Rahayu, 2006 Bouvier, 1893, by original designation; gender Pteropagurus spinulocarpus McLaughlin, 2007 masculine) Pumilopagurus McLaughlin & Rahayu, 2008 Rhodochirus hirtimanus (Faxon, 1893) [Pylopagurus] = Pumilopagurus McLaughlin & Rahayu, 2008 Rhodochirus rosaceus (A. Milne-Edwards & Bouvier, (type species Pumilopagurus tuberculomanus 1893) [Pylopagurus] McLaughlin & Rahayu, 2008, by original designation; gender masculine) = Pylopagurus acutus Forest & de Saint Laurent, 1968 Pumilopagurus tuberculomanus McLaughlin & Rahayu, 2008 Scopaeopagurus McLaughlin & Hogarth, 1998 Pygmaeopagurus McLaughlin, 1986 = Scopaeopagurus McLaughlin & Hogarth, 1998 (type species Scopaeopagurus megalochirus = Pygmaeopagurus McLaughlin, 1986 (type species McLaughlin & Hogarth, 1998, by original Pygmaeopagurus hadrochirus McLaughlin, 1986 designation; gender masculine) by original designation; gender masculine) Scopaeopagurus megalochirus McLaughlin & Hogarth, Pygmaeopagurus hadrochirus McLaughlin, 1986 1998 Pylopaguridium McLaughlin & Lemaitre, 2001 = Pylopaguridium McLaughlin & Lemaitre, 2001 (type species Pylopaguridium markhami McLaughlin & Lemaitre, 2001 by original designation; gender neuter) 35 McLaughlin et al.: Checklist of world Lithodoidea, Lomisoidea and Paguroidea Solenopagurus de Saint Laurent, 1968 = Benthopagurus Wass, 1963 (type species Benthopagurus schmitti Wass, 1963, by original = Solenopagurus de Saint Laurent, 1968 (type designation; gender masculine) species Cestopagurus lineatus Wass, 1963, by Tomopagurus chacei (Wass, 1963) [Pylopagurus] Tomopagurus cokeri (Hay, 1917) [Pagurus] original designation; gender masculine) = Benthopagurus schmitti Wass, 1963 Solenopagurus diomedeae (Faxon, 1893) Tomopagurus cubensis (Wass, 1963) [Benthopagurus] Tomopagurus maclaughlinae Haig, 1976 [Catapagurus] Tomopagurus merimaculosus (Glassell, 1937) [Pagurus] Solenopagurus lineatus (Wass, 1963) [Cestopagurus] Tomopagurus purpuratus (Benedict, 1892) [Eupagurus] Solitariopagurus Türkay, 1986 = Pagurus bunomanus Glassell, 1937 (nomen nudum) = Solitariopagurus Türkay, 1986 (type species Tomopagurus rubropunctatus A. Milne-Edwards & Solitariopagurus profundus Türkay, 1986 by Bouvier, 1893 original designation; gender masculine) = Pagurus rubrolineatus Wass, 1963 Tomopagurus wassi McLaughlin, 1981 Solitariopagurus profundus Türkay, 1986 Trichopagurus de Saint Laurent, 1968 Solitariopagurus triprobolus Poupin & McLaughlin, = Trichopagurus de Saint Laurent, 1968 (type species Catapaguroides ? trichophthalmus Forest, 1954, 1996 by original designation; gender masculine) {24} Solitariopagurus trullirostris McLaughlin, 2000 Trichopagurus trichophthalmus (Forest, 1954) [Catapaguroides ?] Solitariopagurus tuerkayi McLaughlin, 1997 Trichopagurus macrochela Komai & Osawa, 2005 Turleania McLaughlin, 1997 Spiropagurus Stimpson, 1858 = Laurentia McLaughlin & Haig, 1996 (type species = Spiropagurus Stimpson, 1858 (type species Laurentia albatrossae McLaughlin & Haig, 1996, by original designation) (name preoccupied by Pagurus spiriger De Haan, 1849 by monotypy; Laurentia Ragonot, 1888, Insecta) gender masculine) = Turleania McLaughlin, 1997 (replacement name) Spiropagurus elegans Miers, 1881 Turleania albatrossae (McLaughlin & Haig, 1996) Spiropagurus fimbriatus Lewinsohn, 1982 [Laurentia] Turleania boucheti McLaughlin, 2007 Spiropagurus lophomeris Alcock, 1905 {23} Turleania balli (McLaughlin & Haig, 1996) [Laurentia] = Spiropagurus spiriger var. lophomeris Alcock, Turleania multispina McLaughlin, 1997 Turleania saliens Osawa & Fujita, 2008 1905 Turleania senticosa (McLaughlin & Haig, 1996) [Laurentia] Spiropagurus profundorum Alcock, 1905 {23} = Turleania similis Komai, 1999 = Spiropagurus spiriger var. profundorum Alcock, Turleania sibogae (McLaughlin & Haig, 1996) [Laurentia] 1905 Turleania spinimanus Komai, 1999 Turleania tenebrosa Osawa & Fujita, 2008 Spiropagurus spinosicarpis Alcock, 1905 {23} Xylopagurus A. Milne-Edwards, 1880 = Spiropagurus spiriger var. spinosicarpis Alcock, = Xylopagurus A. Milne-Edwards, 1880 (type species Xylopagurus rectus A. Milne-Edwards, 1905 1880, by monotypy; gender masculine) Spiropagurus spiriger (De Haan, 1849) [Pagurus] Xylopagurus anthonii Lemaitre, 1995 Xylopagurus caledonicus Forest, 1997 Tarrasopagurus McLaughlin, 1997 Xylopagurus cancellarius Walton, 1950 Xylopagurus philippinensis Forest, 1997 = Tarrasopagurus McLaughlin, 1997 (type Xylopagurus rectus A. Milne-Edwards, 1880 Xylopagurus tayrona Lemaitre & Campos, 1993 species Tarrasopagurus rostrodenticulatus Xylopagurus tenuis Lemaitre, 1995 McLaughlin, 1997, by original designation; gender Incertae sedis masculine) Anapagurus acutus A. Milne-Edwards& Bouvier, 1893 {25} Tarrasopagurus rostrodenticulatus McLaughlin, 1997 Tomopaguroides Balss, 1912 = Tomopaguroides Balss, 1912 (type species Parapagurus valdividae Balss, 1911, by original designation; gender masculine) Tomopaguroides valdividae (Balss, 1911) [Parapagurus] Tomopaguropsis Alcock, 1905 = Tomopaguropsis Alcock, 1905 [type species Tomopaguropsis lanata Alcock, 1905, by subsequent selection by McLaughlin (1997); gender feminine] Tomopaguropsis lanata Alcock, 1905 Tomopaguropsis miyakei McLaughlin, 1997 Tomopaguropsis problematica (A. Milne-Edwards & Bouvier, 1893) [Eupagurus] Tomopagurus A. Milne-Edwards & Bouvier, 1893 = Tomopagurus A. Milne-Edwards & Bouvier, 1893 (type species Tomopagurus rubropunctatus A. Milne-Edwards & Bouvier, 1893, by monotypy; gender masculine) 36 THE RAFFLES BULLETIN OF ZOOLOGY 2010 Anapagurus marginatus A. Milne-Edwards & Bouvier, Catapagurus; still no mention was made of a type species for Hemipagurus. After examining one of 1893 {25} the syntypes of Catapagurus sharreri A. Milne- Edwards, 1880, Smith (1883) concluded that his own NOTES C. socialis was conspecific with C. sharreri. It was only when Asakura (2001) resurrected Hemipagurus {1} To indicate their uncertainty regarding the generic was H. gracilis cited as the type species of Smith’s assignment of the new species to Anapagurus, A. (1881a) taxon. Although at the time of Asakura’s Milne-Edwards & Bouvier, 1900, separated the (2001) reinstatement, Hemipagurus contained only H. generic name from the specific name, dubius, with a gracilis, it is not the type species by monotypy (ICZN question mark. Art. 69.4) but rather by designation by Asakura. {2} Thomas Bell’s “A History of British Crustacea” was {9} Catapagurus maclaughlinae Asakura, 2001, was published in parts between 1844 and 1853. Part III, established on two females from the Seycheles and which contained the pagurid species Anapagurus distinguished from other species of Catapagurus hyndmanni and A. laevis was published in January by weak development of carpal and meral spines 1846 (Gordon, 1959). on the ambulatory legs. Subsequent discovery of a male of C. maclaughlinae from a French Seycheles {3} Pagurus pilosipes (Stimpson, 1858) will be transferred expedition and comparison with an expanded sample to Boninpagurus by Komai & Okuno (in preparation) of C. alcocki has shown that the two taxa are as the senior synonym of B. acanthocheles. conspecific. Catapagurus maclaughlinae will be put into synonymy with C. alcocki in an upcoming review {4} Catapaguroides kasei Osawa & Takeda, 2004 appears of the genus where justification for the synonymy will to be synonymous with C. foresti, but confirmation be presented. is still needed. {10} Although C. gracilis var. intermedius was described by {5} De Saint Laurent (1968a) felt certain that the western A. Milne-Edwards & Bouvier (1893), this subspecies Pacific species she identified as Catapaguroides was overlooked by Asakura (2001) in his generic microps was conspecific with the eastern Atlantic revision. In a reassessment currently in progress population; however recent collections suggest that McLaughlin has compared the two syntypes of this the former may actually represent a distinct taxon. subspecies with the nominal subspecies and found no justification for two distinct taxa. Catapagurus {6} Catapaguroides setosus Edmondson, 1951, was gracilis intermedius will be placed in synonymy in questionably synonymized with C. fragilis by de the upcoming reassessment. Saint Laurent (1968a) but reinstated by McLaughlin & Pittman (2002) after comparing representatives of {11} In her ongoing study of species of Catapagurus, the two species. McLaughlin has found C. imperalis to be conspecific with C. oculocrassus and will be placed in synonymy {7} Catapagurus A. Milne-Edwards, 1880, had long with this species. been considered the senior synonym of Hemipagurus Smith, 1881 until Asakura (2001) proposed the {12} Asakura (2001) inappropriately designated a lectotype resurrection of the latter. That reinstatement was for C. sharreri; the same male had previously been so rejected by McLaughlin (2004) who showed that designated by A. Milne-Edwards & Bouvier (1893: the characters used by Asakura (2001) to distinguish 130). Catapagurus from Hemipagurus reflected only intrageneric variability and growth related differences. {13} It is possible that the type series of this genus was McLaughlin returned Hemipagurus to synonymy. Five a mixture of more than one species of different more species of Catapagurus are now being described sexes, because such a misidentification is not by the first author. uncommon in Yokoya’s (1933) material. Despite the collecting efforts of one of the present authors {8} In Smith’s (1881a) initial description of Hemipagurus, (TK), no specimens that agree with Yokoya’s original no species were mentioned. Subsequently, Smith description have been encountered in Japan. (1881b) repeated his description of the genus and described the two species he assigned to it, H. {14} The possibility exists that this taxon may be a junior socialis Smith, 1881 and H. gracilis Smith, 1881; synonym of L. (A.) triserratus. he did not designate either species as the type for the genus. The following year Smith (1882) determined {15} Eupagurus thompsoni Filhol, 1885, became a that Hemipagurus was identical with A. Milne- secondary junior homonym of Pagurus thompsoni Edwards’ (1880) Catapagurus, placed Hemipagurus Bell, 1853 when the ICZN (Opinion 472) placed in synonymy and transferred its two species to Eupagurus on the Officical Index of Rejected and 37 McLaughlin et al.: Checklist of world Lithodoidea, Lomisoidea and Paguroidea Invalid Generic Names in Zoology. As discussed isochirus and Pagurus pilosimanus without localities, by McLaughlin & Gunn (1992), Forest (in Pike, and Pagurus pilosiusculus from Mr. Dring’s collection, 1961) suggested that Filhol’s (1885b) P. thompsoni Swan River. None of these taxa were provided with was synonymous with P. lacertosus (Henderson, even brief descriptions or illustrations, and no further 1888). Forest (unpublished) subsequently concluded mention of any of them could be found in White’s that those two taxa were distinct, and both species subsequent publications. were transferred to Pylopagurus by Forest & de Saint Laurent (1968) thus eliminating the need for a {21} The conspecificity of Pagurus comptus and P. forceps replacement name for Filhol’s taxon (ICZN, 2000: has been the subject of debate since Lagerberg (1905) Art 59.2). Both species were ultimately transferred to first placed P. comptus in synonymy with P. forceps. Lophopagurus McLaughlin, 1981, whereas Bell’s P. However, a recent study by Mantelatto et al. (2009), thompsoni was found to be synonymous with Pagurus using both morphological and genetic data, has pubescens Kröyer, 1838. demonstrated conclusively that the two taxa represent distinct species. {16} Apparently uncertain about the generic placement of their solitary juvenile specimen, Chevreux & Bouvier {22} Glassell’s (1937) Pagurus pollexcavus was not (1892) described the taxon as Eupagurus ? minimum. reported as occurring in the Gulf of California by Forest (1961), having obtained additional, and mature Hendrickx & Harvey (1999), nor was it included specimens, recognized the species as belonging to a the Lemaitre & Cruz Castaño’s (2004) checklist of genus distinct from other known pagurid genera, and eastern Pacific species of Pagurus. But to the best extablished the genus Paguridium Forest, 1961 for of our knowledge, this species has not been put it. into synonymy with any other regional species or considered incertae sedis for any reason. Therefore {17} As previously noted, when first established by we have listed its status simply as uncertain. Fabricius (1775) Pagurus included a number of Linnaeus’ (1758) non crab-like species of Cancer. It {23} McLaughlin (2002) elevated all of Alcock’s (1905) was 35 years later that Pagurus bernhardus (Linnaeus, subspecies of Spiropagurus spiriger to full generic 1758) was selected by Latreille (1810) as the type rank. species of the genus and another quarter century before the first detailed classification of Pagurus was {24} When Forest (1954) described Catapaguroides published (H. Milne Edwards 1836). As can be clearly ? trichophthalmus, the genus Catapaguroides seen in the brief accounts of subsequent carcinologists appeared to have considerable overlap with a from Brandt (1851) to Rathbun (1903) and beyond, similar genus, Cestopagurus. Not being certain of inaccurate interpretations and misidentifications its generic placement, Forest questionably assigned resulted in nomenclatorial chaos that prevailed until trichophthalmus to Catapaguroides. the matter was brought before the International Commission (Forest & Holthuis, 1955). In its ruling, {25} García-Gómez (1994) examined the two female the Commission validated the generic names Pagurus specimens upon which A. acutus and A. marginatus (sensu Fabricius) and Dardanus Paul’son (Opinion were based and concluded that they were inaccurately 472). assigned to Anapagurus. As they could not be assigned with confidence to any taxon, García-Gómez {18} Forest (1955) placed Pagurus excavatus (Herbst, considered them incertae sedis. 1791) in synonymy with Pagurus alatus Fabricius, 1775. However, Ingle (1985) presented evidence Family Parapaguridae Smith, 1882 supporting Selbie’s (1921) opinion that the two taxa were distinct, with P. excavatus restricted in its Bivalvopagurus Lemaitre 1993 northern distribution to the Bay of Biscay. Ingle (1985) reinstated P excavatus and placed the morphologically = Bivalvopagurus Lemaitre 1993 (type species similar P. variabilis (A. Milne-Edwards & Bouvier, 1892) in synonymy with the northern P. alatus. Parapagurus sinensis de Saint Laurent, 1972, by {19} Thomas Bell’s A History of British Crustacea was monotypy; gender masculine) published in parts between 1844 and 1853. Part III, which included the pagurid species Pagurus Bivalvopagurus sinensis (de Saint Laurent, 1972) cuanensis, ulidianus and P. Forbesii, was published in January 1846 (Gordon 1959). [Parapagurus] {20} In his list of the Crustacea in the British Museum, Oncopagurus Lemaitre, 1996 Adam White (1847: 59–62) listed four new species of Pagurus: Pagurus chinensis from China, Pagurus = Oncopagurus Lemaitre, 1996 (type species Eupagurus bicristatus A. Milne-Edwards, 1880, by original designation; gender masculine) Oncopagurus africanus (de Saint Laurent, 1972) [Parapagurus] Oncopagurus bicristatus (A. Milne-Edwards, 1880) [Eupagurus] {1} 38 THE RAFFLES BULLETIN OF ZOOLOGY 2010 Oncopagurus cidaris Lemaitre, 1996 Paragiopagurus hobbiti (Macpherson, 1983) Oncopagurus conicus Lemaitre, 2006 [Parapagurus] Oncopagurus curvispina (de Saint Laurent, 1974) Paragiopagurus insolitus Lemaitre, 1997 [Parapagurus] {2} Paragiopagurus macrocerus (Forest, 1955) Oncopagurus glebosus Lemaitre, 1997 [Parapagurus] Oncopagurus gracilis (Henderson, 1888) Paragiopagurus pacificus (Edmondson, 1925) [Parapagurus] [Sympagurus] = Parapagurus bicristatus gracilis Henderson, Paragiopagurus pilimanus (A. Milne-Edwards, 1880) 1888 {1} [Eupagurus] = Sympagurus arcuatus A. Milne-Edwards & Paragiopagurus rugosus (de Saint Laurent, 1972) Bouvier, 1893 [Parapagurus] = Pylopagurus exquisitus Boone, 1927 Paragiopagurus ruticheles (A. Milne-Edwards, 1891) Oncopagurus haigae (de Saint Laurent, 1972) [Eupagurus] [Parapagurus] Paragiopagurus schnauzer Lemaitre, 2006 Oncopagurus indicus (Alcock, 1905) [Sympagurus] Paragiopagurus tuberculosus (de Saint Laurent, 1972) = Sympagurus bicristatus var. indicus Alcock, [Parapagurus] 1905 {1} Paragiopagurus ventilatus Lemaitre, 2004 Oncopagurus minutus (Henderson, 1896) Paragiopagurus wallisi (Lemaitre, 1994) [Sympagurus] [Parapagurus] Paragiopagurus n. sp. 1 {5} Oncopagurus mironovi Zhadan, 1997 Paragiopagurus n. sp. 2 {5} Oncopagurus monstrosus (Alcock, 1894) Paragiopagurus n. sp. 3 {5} [Parapagurus] Paragiopagurus n. sp. 4 {5} = Sympagurus arcuatus var. monstrosus Alcock, Parapagurus Smith, 1879 1894 = Parapagurus Smith, 1879 (type species = ? Eupagurus brevimanus Yokoya, 1933 {3} Parapagurus pilosimanus Smith, 1879, by Oncopagurus oimos Lemaitre, 1998 monotypy; gender masculine) Oncopagurus orientalis (de Saint Laurent, 1972) Parapagurus abyssorum (Filhol, 1885) [Pagurus] {7} [Parapagurus] = Parapagurus abyssorum var. scabra Henderson, Oncopagurus stockmani Zhadan, 1997 1888 Oncopagurus tuamotu Lemaitre, 1994 Parapagurus alaminos Lemaitre, 1986 Oncopagurus sp. A Zhadan, 1997 {4} Parapagurus andreui Macpherson, 1984 Oncopagurus n. sp. 1 {5} Parapagurus benedicti de Saint Laurent, 1972 Oncopagurus n. sp. 2 {5} = Parapagurus pilosimanus benedicti de Saint Oncopagurus n. sp. 3 {5} Laurent, 1972 Oncopagurus n. sp. 4 {5} Parapagurus bouvieri Stebbing, 1910 Paragiopagurus Lemaitre, 1996 Parapagurus foraminosus Lemaitre, 1999 = Paragiopagurus Lemaitre, 1996 (type species Parapagurus furici Lemaitre, 1999 Sympagurus diogenes Whitelegge, 1900, by Parapagurus holthuisi Lemaitre, 1989 {8} original designation; gender: masculine) = Parapagurus abyssorum Henderson, 1888 Paragiopagurus acutus (de Saint Laurent, 1972) Parapagurus janetae Lemaitre, 1999 [Parapagurus] Parapagurus latimanus Henderson, 1888 = Parapagurus acutus acutus de Saint Laurent, Parapagurus microps de Saint Laurent, 1972 1972 {6} Parapagurus nudus (A. Milne-Edwards, 1891) Paragiopagurus bicarinatus (de Saint Laurent, 1972) [Sympagurus] [Parapagurus] Parapagurus pilosimanus Smith, 1879 {9} = Parapagurus acutus bicarinatus de Saint Laurent, = Eupagurus jacobii A. Milne-Edwards, 1880 1972 {6} = Sympagurus grimaldii A. Milne-Edwards & Paragiopagurus boletifer (de Saint Laurent, 1972) Bouvier, 1897 [Parapagurus] Parapagurus richeri Lemaitre, 1999 = Parapagurus sculptochela Zarenkov, 1990 Parapagurus saintlaurentae Lemaitre, 1999 Paragiopagurus bougainvillei (Lemaitre, 1994) Parapagurus stenorhinus Lemaitre, 1999 [Sympagurus] Parapagurus wolffi Lemaitre, 1999 Paragiopagurus diogenes (Whitelegge, 1900) Probeebei Boone, 1926 [Sympagurus] = Probeebei Boone, 1926 (type species Probeebei Paragiopagurus fasciatus Lemaitre & Poupin, 2003 mirabilis Boone, 1926, by monotypy; gender Paragiopagurus hirsutus (de Saint Laurent, 1972) masculine) [Parapagurus] = Planopagurus Wolff, 1960 (type species = Parapagurus acutus hirsutus de Saint Laurent, Planopagurus galathea Wolff, 1960, by monotypy; 1972 {6} gender masculine) 39 McLaughlin et al.: Checklist of world Lithodoidea, Lomisoidea and Paguroidea Probeebei mirabilis Boone, 1926 Typhlopagurus foresti de Saint Laurent, 1972 = Planopagurus galathea Wolff, 1960 NOTES Strobopagurus Lemaitre, 1989 {1} In her revision of Parapagurus, de Saint Laurent (1972) recognized three subspecies of A. Milne- = Strobopagurus Lemaitre, 1989 (type species Edwards’s bicristatus, Parapagurus bicristatus bicristatus (A. Milne-Edwards, 1880), P. b. gracilis Sympagurus gracilipes A. Milne-Edwards, 1891, Henderson, 1888, and P. b. indicus (Alcock, 1905b). All were elevated to specific status by Lemaitre by original designation; gender masculine) (1989). Strobopagurus breviacus Lemaitre, 2004 {2} This species was described by de Saint Laurent (1974) as Parapagurus curvispina and later placed Strobopagurus gracilipes (A. Milne-Edwards, 1891) in Paragiopagurus by Lemaitre (1996). Examination of type material, however, has revealed that it needs [Sympagurus] to be formally assigned to Oncopagurus as a new combination (Lemaitre, in prep.). Strobopagurus sibogae (de Saint Laurent, 1972) {3} According to Lemaitre (1996), de Saint Laurent (1972) [Parapagurus] questionably synonymized Eupagurus brevimanus with Alcock’s (1894) Parapagurus monstrosus. = Parapagurus kilburni Kensley, 1973 Lemaitre (1996) found insufficient information to confirm or deny the synonymy, and presumed that Sympagurus Smith, 1883 Yokoya’s (1933) specimens, like most of his other material, were no longer extant. = Sympagurus Smith, 1883 (type species Sympagurus {4} Zhadan (1997: 67, fig. 9) did not provide a specific pictus Smith, 1883, by monotypy; gender name, and only noted that his material was intermediate between O. tuamotu and O. cidaris masculine) {5} Formal descriptions are in preparation by one of the Sympagurus acinops Lemaitre, 1989 authors (RL). Sympagurus affinis (Henderson, 1888) [Parapagurus] {6} De Saint Laurent’s (1972) three subspecies, Parapagurus acutus acutus de Saint Laurent, 1972, Sympagurus andersoni (Henderson, 1896) P. a. bicarinatus de Saint Laurent, 1972, and P. a. hirsutus de Saint Laurent, 1972, were assigned [Parapagurus] to Sympagurus Smith, 1883 by Lemaitre (1989). Subsequently Lemaitre (1996) elevated these three = Parapagurus andersoni var. brevimanus Alcock, subspecies to specific status and assigned them to Paragiopagurus Lemaitre, 1996. 1901 {7} This is a senior homonym of Parapagurus abyssorum Sympagurus aurantium Lemaitre, 2004 Henderson, 1888. Sympagurus brevipes (de Saint Laurent, 1972) {8} Replacement name for Parapagurus abyssorum Henderson, 1888, a junior homonym of Parapagurus [Parapagurus] abyssorum (Filhol, 1885) Sympagurus burkenroadi Thompson, 1943 {9} All the subspecies of Parapagurus pilosimanus Smith, 1879 proposed by de Saint Laurent (1972) were = Sympagurus papposus Lemaitre, 1996 elevated to specific status by Lemaitre (1989). Sympagurus chani Lemaitre, 2004 {10} De Grave et al. (2009) incorrectly attributed authorship of the genus Tylaspis to Tizard, Mosely, Buchanan Sympagurus dimorphus (Studer, 1883) [Eupagurus] & Murray (1885), presumably because they were the editors of the narrative of the ‘Challenger’ cruise. = Parapagurus brevimanus Balss, 1911 However, the report on the Anomura (pp. 897–901) = ? Eupagurus modicellus Stebbing, 1914 = Sympagurus arcuatus var. johnstoni Hale, 1941 = Sympagurus arcuatus var. mawsoni Hale, 1941 Sympagurus dofleini (Balss, 1912) [Parapagurus] = Parapagurus ijimai Terao, 1913 = Parapagurus rectichela Zarenkov, 1990 Sympagurus pictus Smith, 1883 Sympagurus planimanus (de Saint Laurent, 1972) [Parapagurus] Sympagurus poupini Lemaitre, 1994 Sympagurus soela Lemaitre, 1996 Sympagurus spinimanus (Balss, 1911) [Parapagurus] Sympagurus symmetricus Lemaitre, 2004 Sympagurus trispinosus (Balss, 1911) [Parapagurus] = Parapagurus arcuatus var. trispinosa Balss, 1911 Sympagurus villosus Lemaitre, 1996 Tsunogaipagurus Osawa, 1995 = Tsunogaipagurus Osawa, 1995 (type species Parapagurus chuni Balss, 1911, by original designation; gender masculine) Tsunogaipagurus chuni (Balss, 1911) [Parapagurus] Tylaspis Henderson, 1885 {10} = Tylaspis Henderson, 1885 (type species Tylaspis anomala Henderson, 1885, by monotypy; gender feminine) Tylaspis anomala Henderson, 1885 Typhlopagurus de Saint Laurent, 1972 = Typhlopagurus de Saint Laurent, 1972 (type species Typhlopagurus foresti de Saint Laurent, 1972, by original designation; gender masculine) 40 THE RAFFLES BULLETIN OF ZOOLOGY 2010 actually was written by Henderson, as Tizard et al. = Pylodietes miersi McGinitie & McGinitie, 1949 acknowledged. The illustration and brief diagnosis of (misspelling of Pylocheles) “Tylaspis n. gen. et sp.” by Henderson (1885: 900–901, fig. 329) meet ICZN requirements for publication. The Subfamily Pomatochelinae Stebbing, 1914 correct author of this genus is therefore Henderson (1885). Pomatocheles Miers, 1879 = Pomatocheles Miers, 1879 (type species Family Pylochelidae Bate, 1888 Pomatocheles jeffreysii Miers, 1879, by monotypy; gender masculine) Subfamily Pylochelinae Bate, 1888 Pomatocheles gaillardi Forest, 1987 Pomatocheles jeffreysii Miers, 1879 Pomatocheles stridulans Forest, 1987 Bathycheles Forest, 1987 {1} = Pylocheles (Bathycheles) Forest, 1987 (type species Subfamily Trizochelinae Forest 1987 Pylocheles (Bathycheles) incisus Forest 1987, by original designation; gender masculine) Tribe Cancellochelini Forest, 1987 Cancellocheles Forest, 1987 Bathycheles crosnieri (Forest, 1987) [Pylocheles = Cancellocheles Forest, 1987 (type species (Bathycheles)] Pomatocheles sculptipes Miyake, 1978, by monotypy; gender masculine) Bathycheles cubensis (Ortiz & Gómez, 1986) Cancellocheles sculptipes (Miyake, 1978) [Pylocheles] [Pomatocheles] = Pylocheles (Bathycheles) chacei Forest, 1987 Bathycheles incisus (Forest, 1987) [Pylocheles (Bathycheles)] Bathycheles integer (Forest, 1987) [Pylocheles Tribe Mixtopagurini Bouvier, 1895 {3} (Bathycheles)] Mixtopagurus A. Milne-Edwards, 1880 = Pylocheles (Bathycheles) profundus Forest, = MixtopagurusA.Milne-Edwards,1880(typespecies 1987 Mixtopagurus paradoxus A. Milne-Edwards, Bathycheles macgilchristi (Alcock, 1905) [Chiroplatea] 1880, by monotypy; gender masculine) (misspelling of Cheiroplatea) Mixtopagurus paradoxus A. Milne-Edwards, 1880 Bathycheles phenax McLaughlin & Lemaitre, 2009 Cheiroplatea Bate, 1888 Tribe Parapylochelini Parapylocheles Alcock, 1901 = Cheiroplatea Bate, 1888 (type species Cheiroplatea = Parapylocheles Alcock, 1901 (type species cenobita Bate, 1888, by monotypy; gender Pylocheles scorpio Alcock, 1894, by monotypy; gender masculine) masculine) Parapylocheles scorpio (Alcock, 1894) = Chiroplatea Ortmann, 1892 (misspelling of Cheiroplatea) Cheiroplatea cenobita Bate, 1888 Cheiroplatea laticauda Boas, 1926 Tribe Trizochelini Forest, 1987 Forestocheles McLaughlin & Lemaitre, 2009 Cheiroplatea mitoi Miyake, 1978 = Forestocheles McLaughlin & Lemaitre, 2009 Cheiroplatea pumicicola Forest, 1987 (type species Trizocheles perplexus Forest, 1987, by original designation; gender masculine) Cheiroplatea scutata Ortmann, 1892 Forestocheles perplexus (Forest, 1987) [Trizocheles] = Chiroplatea scutata Ortmann, 1892 (misspelling Trizocheles Forest, 1987 of Cheiroplatea) = Trizocheles Forest, 1987 (type species Pylocheles spinosus Henderson, 1888, by original designation; Cheiroplatea stenurus Forest, 1987 gender masculine) Pylocheles A. Milne-Edwards, 1880 Trizocheles albatrossi Forest, 1987 Trizocheles balssi (Stebbing, 1914) [Pomatocheles] = Pylocheles A. Milne-Edwards, 1880 (type species Trizocheles boasi Forest, 1987 Pylocheles agassizi A. Milne-Edwards, 1880, by = Trizocheles gracilis Forest, 1987 Trizocheles brachyops Forest & de Saint Laurent, 1987 monotypy; gender masculine) Trizocheles brevicaulis (Boas, 1926) [Mixtopagurus] Trizocheles caledonicus Forest, 1987 Pylocheles agassizi A. Milne-Edwards, 1880 Trizocheles hoensonae McLaughlin & Lemaitre, 2009 Trizocheles laurentae Forest, 1987 = Pylocheles partitus Benedict, 1901 Trizocheles longicaulis (Boas, 1926) [Mixtopagurus] Trizocheles loquax Forest, 1987 Pylocheles mortensenii Boas, 1926 Trizocheles manningi Forest, 1987 Trizocheles mendanai McLaughlin & Lemaitre, 2009 = Pylocheles rigidus Yokoya 1933 Xylocheles Forest 1987 {2} = Xylocheles Forest, 1987 (type species Pylocheles (Xylocheles) macrops Forest, 1987, by original designation; gender masculine) Xylocheles macrops (Forest, 1987) [Pylocheles (Xylocheles)] Xylocheles miersi (Alcock & Anderson, 1899) [Pylocheles] 41 McLaughlin et al.: Checklist of world Lithodoidea, Lomisoidea and Paguroidea Trizocheles moosai Forest, 1987 Ahyong, S. T. & T.-Y. Chan, 2010. Lithodes formosae, a new species Trizocheles mutus Forest, 1987, of king crab from Taiwan (Crustacea: Decapoda: Lithodidae). Trizocheles pilgrimi Forest & McLaughlin, 2000 Zootaxa, 2332: 61–68. Trizocheles pulcher Forest, 1987 Trizocheles sakaii Forest, 1987 Ahyong, S. T. & E. W. Dawson, 2006. Lithodidae from the Ross Trizocheles spinosus (Henderson, 1888) Sea, Antarctica, with descriptions of two new species (Crustacea: Decapoda: Anomura). Zootaxa, 1303: 45–68. = Trizocheless spinosus bathamae Forest & de Saint Laurent, 1987 Ahyong, S. T. & D. O’Meally, 2004. Phylogeny of the Decapoda Reptantia: Resolution using three molecular loci and morphology. NOTES Raffles Bulletin of Zoology, 52: 673–693. {1} Authorship of Bathycheles was incorrectly credited Ahyong, S. T., E. Macpherson & T.Y. Chan, 2010. Lithodoidea. to McLaughlin & Lemaitre, 2009, by De Grave et al. In T.-Y. Chan (ed.), Crustacean Fauna of Taiwan: Crab-like (2009). Bathycheles was proposed by Forest (1987) Anomurans (Hippodea, Lithodoidea and Porcellanidae). as a subgenus of Pylocheles. In their reassessment National Taiwan Ocean University, Keelung. Pp. 42–66. of pylochelid classification, McLaughlin & Lemaitre (2009) elevated Bathycheles to full generic rank, but in Ahyong, S. T., K. E, Schnabel & E. W. Maas, 2009. Anomuran accordance with ICZN Article 43.1 (1999) the author Phylogeny: New insights from molecular data. In: Martin, J. and date for the taxon remain as Forest (1987). W., K. A. Crandall & D. L. Felder (eds.), Decapod Crustacean Phylogenetics. Crustacean Issues 18. CRC Press, Boca Raton, {2} Authorship of Xylocheles was incorrectly credited FL, New York, London. Pp. 399–414. to McLaughlin & Lemaitre, 2009 by De Grave et al. (2009). Xylocheles was proposed by Forest (1987) Alcock, A., 1894. Natural history notes from H.M. Indian Marine as a subgenus of Pylocheles. In their reassessment Survey Steamer ‘Investigator’, Commander R.F. Hoskyn, R.N., of pylochelid classification, McLaughlin & Lemaitre commanding.—Series II., No. 1. On the results of deep-sea (2009) elevated Xylocheles to full generic rank, but in dredging during the season 1890-91. Annals and Magazine of accordance with ICZN Article 43.1 (1999) the author Natural History, (6)13: 225–245; 321–334; 400–411. and date for the taxon remain as Forest ( 1987). Alcock, A., 1901. Catalogue of the Indian decapod Crustacea {3} Although the tribe name Mixtopaguriens was in the collection of the Indian Museum. Part 1. Brachyura. Latinized by A. Milne-Edwards & Bouvier (1899), Fascicle I. Introduction and dromides or Dromiacea (Brachyura Forest (1987) correctly assigned the authorship and Primigenia). Indian Museum, Calcutta. 80 pp. date to Bouvier (1895a) in accordance with Article 11f (iii) of the ICZN Code (1985). Alcock, A., 1905a. Marine Crustaceans. XIV. Paguridae. In: Gardiner, J. S. (ed.). The fauna and geography of the Maldive Family Pylojacquesidae McLaughlin & Lemaitre, 2001 and Laccadive Archipelagoes. Being an account of the work carried on and the collections made by an expedition during Pylojacquesia McLaughlin & Lemaitre, 2001 the years 1899 and 1900, 2: 827–835. University Press, = Pylojacquesia McLaughlin & Lemaitre, 2001 (type Cambridge. species Pylojacquesia colemani McLaughlin & Lemaitre, 2001, by original designation; gender Alcock, A., 1905b. Anomura. Fasc. I. Pagurides. Catalogue of feminine) the Indian decapod Crustacea in the collections of the Indian Museum, 2: 1–197. Indian Museum, Calcutta. Pylojacquesia colemani McLaughlin & Lemaitre, 2001 Lemaitreopsis McLaughlin, 2007 Alcock, A. & A. R. Anderson, 1899. Natural history notes from H.M. Indian marine survey steamer “Investigator”, Commander = Lemaitreopsis McLaughlin, 2007 (type species T.H. Heming, R.N. commanding. Series 3, no. 2. An account Lemaitreopsis holmi McLaughlin, 2007, by of the deep-sea Crustacea dredged during the surveying-season original designation; gender feminine) of 1897–98. Annals and Magazine of Natural History, (7)3: 1–27. Lemaitreopsis holmi McLaughlin, 2007 Anderson, E. M. & G. M. Cailliet, 1974. Crab and snailfish LITERATURE CITED commensalism in Monterey Bay. Underwater Naturalist, 8: 39–31. Ahyong, S.T., 2010a. Neolithodes flindersi, a new species of king crab from southeastern Australia (Crustacea: Decapoda: Andrade, H., 1980. Nueva especie de Paralomis en aguas de Chile: Lithodidae). Zootaxa, 2362: 55–62. Paralomis chilensis n. sp. (Crustacea, Anomura, Lithodidae). Boletin del Museo Nacional de Historia Natural, Chile, 37: Ahyong, S. T., 2010b. The Marine Fauna of New Zealand: King 269–273. Crabs of New Zealand, Australia and the Ross Sea (Crustacea: Decapoda: Lithodidae). NIWA Biodiversity Memoir, 123: Asakura, A., 1991. A new hermit crab (Decapoda: Paguridae) from 1–195. rocky shores of the Ogasawara (Bonin) Islands. Proceedings of the Biological Society of Washington, 104(4): 793–799. Asakura, A., 1999. Icelopagurus tuberculosus, a distinct new hermit crab species (Crustacea: Decapoda: Paguridae) from Japan. Proceedings of the Biological Society of Washington 112: 381–395. Asakura, A., 2000. A review of Japanese species of Pylopaguropsis Alcock, 1905 (Decapoda: Anomura: Paguridae). Crustacean Research, 29: 70–108. Asakura, A., 2001. A revision of the hermit crabs of the genera Catapagurus A. Milne-Edwards and Hemipagurus Smith 42 THE RAFFLES BULLETIN OF ZOOLOGY 2010 from the Indo-West Pacific (Crustacea: Decapoda: Anomura: Diogenidae) species complex with the description of five new Paguridae). Invertebrate Taxonomy, 15: 823–891. species from the Mexican Pacific. Zootaxa, 2323: 1–71. Asakura, A., 2004. Pseudopaguristes shidarai, new species, a Baba, K., 1986. In: Baba, K., K-I Hayashi & M. Toriyama, distinct hermit crab (Crustacea: Decapoda: Diogenidae) from Decapod crustaceans from continental shelf and slope around Japan. Proceedings of the Biological Society of Washington, Japan. The Intensive research of unexploited fishery resources 117: 153–168. on continental slopes. Japan Fisheries Resource Conservation Association, Tokyo. 336 pp. Asakura, A., 2005a. Pagurodofleinia, a replacement generic name for Dofleinia McLaughlin & Asakura, 2004 (Crustacea: Decapoda), Baker, W. H., 1905. Notes on South Australian decapod Crustacea. preoccupied by Dofleinia Wassilieff, 1908 (Cnidaria: Anthozoa). Part III. Transactions of the Royal Society of South Australia, Zootaxa, 1089: 64. 29: 252–269. Asakura, A., 2005b. A review of the genus Micropagurus (Crustacea Ball, E. E. & J. Haig, 1972. Hermit crabs from eastern New Guinea. Decapoda Anomura Paguridae). Zootaxa, 1090: 1–34. Pacific Science, 26: 87–107. Asakura, A., 2006a. Dardanus sanguinocarpus Degener, 1925 Balss, H., 1911. Neue Paguriden aus den Ausbeuten der deutschen (Crustacea: Decapoda: Anomura: Diogenidae) species complex: Tiefsee-Expedition “Valdivia” und der japanischen Expedition A redescription of D. sanguinocarpus and descriptions of three Prof. Dofleins. Zoologischer Anzeiger, 38(1): 1–9. new species. Zootaxa, 1230: 1–54. Balss, H., 1912. Paguriden. In: Chun, C. (ed.), Wissenschaftliche Asakura, A., 2006b. Shallow water hermit crabs of the families Ergebnisse der deutschen Tiefsee-Expedition “Valdivia” 1898- Pylochelidae, Diogenidae and Paguridae (Crustacea: Decapoda: 1899, 20(2): 85–124. Gustav Fischer, Jena. Anomura) from the Sea of Japan, with the description of a new species of Diogenes. Bulletin of the Toyama Science Museum, Balss, H., 1913. Ostasiatische Decapoden I. Die Galatheiden 29: 23–103. und Paguriden. In: Beiträge zur Naturgeschichte Ostasiens, herausgegeben von Dr. F. Doflein. Abhandlungen der Asakura, A. & S. Godwin, 2006. Diogenes patae n. sp., a new species mathematisch-physische Klasse der Königlich Bayerischen of hermit crab (Crustacea: Decapoda: Anomura: Diogenidae) Akademie der Wissenschaften, Supplement 2(9): 1–85. from American Samoa. Zoosystema, 28(2): 457–463. Balss, H., 1916. Die Decapoden des Roten Meeres. II. Anomuren, Asakura, A. & A. Hirayama, 2002. Dardanus jacquesi, a new Dromiaceen und Oxystomen. Expeditionen S.M. Schiff “Pola” in species of hermit crab from the Indo-west Pacific, with the das Rote Meer. Nördliche und südliche Häfte 1895/96–1897/98. redescription of a little known species, Dardanus dearmatus Zoologische Ergebnisse XXXI. Denkschriften der Kaiserlichen (Henderson, 1888). Crustaceana, 75(3–4): 213–230. Akademie der Wissenschaften in Wien Mathematisch- Naturwissenschaftliche Klasse, 92: 1–20. Asakura, A. & T. Kosuge, 2004. Pseudopaguristes bicolor, a new species of hermit crab (Crustacea: Decapoda: Diogenidae) Balss, H., 1921. Crustacea VI: Decapoda Anomura (Paguridea) from Japan, the third species of the genus. Proceedings of the und Brachyura (Dromiacea bis Brachygnatha I.). Beiträge zur Biological Society of Washington 117: 57–67. Kenntnis der Meeresfauna Westafrikas Herausgegeben von W. Michaelsen (Hamburg), 2(2): 37–67. Asakura, A. & P. A. McLaughlin, 2003. A new species of Pseudopaguristes McLaughlin, 2002 (Crustacea: Decapoda: Balss, H., 1957. Decapoda. 12. Systematik. In: Dr. H.G. Bronns’ Diogenidae) from Japan. Proceedings of the Biological Society Klassen und Ordnungen des Tierreichs. Vol. 5, part 1, Book 7: of Washington, 116(2): 453–463. 1505–1672. Akademische Verlagsgesellschaft, Geest & Portig K.-G., Leipzig. Asakura, A. & G. Paulay, 2003. Pylopaguropsis lemaitrei, a new species of hermit crab (Decapoda: Anomura: Paguridae) from Barnard, K. H., 1947. Descriptions of new species of South African French Polynesia. Crustacean Research, 32: 13–25. decapod Crustacea, with notes on synonymy and new records. Annals and Magazine of Natural History, (11)13: 361–392. Asakura, A. & H. Tachikawa, 2000. A new hermit crab of the genus Calcinus from Micronesia, including new information Bate, C. S., 1851. On a new genus and several new species of on C. revi Poupin and McLaughlin (Decapoda: Anomura: British Crustacea. Annals and Magazine of Natural History, Diogenidae). Journal of Crustacean Biology, 20 (Special (2)7: 318–321. Number 2): 266–280. Bate, C. S., 1864. Characters of new species of crustaceans discovered Asakura, A. & H. Tachikawa, 2004. Boninpagurus acanthocheles, by J.K. Lord on the coast of Vancouver Island. Proceedings of a new genus and species of hermit crab (Decapoda: Anomura: the Zoological Society of London, 1864: 661–668. Paguridae) from shallow waters of the Ogsawara (Bonin) Islands, Japan. Journal of Crustacean Biology, 24(1): 157–167. Bate, C. S., 1866. Departure from Colville—Vancouver Island crabs. In: Lord, J. K., Naturalist in Vancouver and British Columbia, Ascanius, P., 1776. Icones Rerum Naturalium, ou Figures enluminées 2: 262–285, Richard Bentley, London. d’Histoire Naturelle du Nord. Cahier, 4: 6 pp. [re-issued, 1806, Möller, Copenhagen] Bate, C. S., 1888. Report on the Crustacea Macrura dredged by H.M.S. Challenger during the years 1872–76. In: Report on the Aurivillius, C. W. S., 1898. Krustaceen aus dem Kamerun–Gebiete. scientific results of the voyage of H.M.S. Challenger during the Bihang till Kungliga Svenska Vetenskaps-Akademiens years 1873-76. Zoology, 24(52): 1–942. Handlingar, 24(4), 1: 1–31. Bell, T., 1844–1853. A history of the British stalk-eyed Crustacea. Ayón Parente, M. & M. E. Hendrickx, 2006, A new species of John van Voorst: London. 386 pp. (issued separately in Stratiotes Thomson, 1899 (Anomura, Paguroidea, Diogenidae) parts). from the eastern tropical Pacific. Zoosystema, 28(2): 487– 494. Benedict, J. E., 1892. Preliminary descriptions of thirty-seven new species of hermit crabs of the genus Eupagurus in the U.S. Ayón Parente, M. & M. E. Hendrickx, 2009. A review of the National Museum. Proceedings of the United States National Dardanus sinistripes (Stimpson, 1859) (Decapoda, Anomura, Museum, 15: 1–26. 43 McLaughlin et al.: Checklist of world Lithodoidea, Lomisoidea and Paguroidea Benedict, J. E., 1895. Descriptions of new genera and species of Borradaile, L. A., 1916. Crustacea. Part 1. Decapoda. British crabs of the family Lithodidae with notes on the young of Antarctic (“Terra Nova”) Expedition, 1910. Natural History Lithodes camtschaticus and Lithodes brevipes. Proceedings of Report. Zoology. 3(2): 75–110. the United States National Museum, 17: 479–488. Bosc, L. A. G., 1802. Histoire Naturelle des Crustacés, contenant Benedict, J. E., 1896. A question concerning a British pagurid. leur description et leurs meours; avec figures dessinés d’après Annals and Magazine of Natural History, (6)18: 99–100. nature. 2: 1–296. Benedict, J. E., 1901a. Four new symmetrical hermit crabs (Pagurids) Bouvier, E.-L., 1890. Révision des Cénobites du Muséum. Bulletin from the West India region. Proceedings of the United States de la Société Philomatique de Paris, (8)2(4): 143–150. National Museum, 23: 771–778. Bouvier, E.-L., 1891. Étude de quelques paguriens recueillis par M. Benedict, J. E., 1901b. The anomuran collections, made by the Fish Jules de Guerne sur les côtes de France et de Norvége. Mémoires Hawk Expedition to Porto Rico. Bulletin of the United States de la Société Zoologique de France, 4: 393–407. Fish Commission, (1900), 20: 131–148. Bouvier, E.-L., 1892. Étude sur les Paguriens recueillis par M. le Berthold, A. A., 1827. Latreille’s Natürliche Familien des Dr. Jousseaume sur les côtes de la Mer Rouge. Bulletin de la Thierreichs. Verlag des Gross-herzoglichen Staates privates Société Philomathique de Paris, (8)4(2): 50–55. Landes-Industrie Comptoirs, Weimar. 604 pp. Bouvier, E.-L., 1893. Paguriens recueillis par M. Diguet, sur Berthold, A. A., 1845. Ueber verschiedene neue oder seltene le littoral de la Basse Californie. Bulletin de la Société Reptilien aus New Granada und Crustaceen aus China. Philomathique de Paris, (8)5: 18–25. Nachrichten Gesellschaft Wissenschaftern Göttingen, 1845: 37–49. Bouvier, E.-L., 1894a. Sur les caractéres et l’évolution des Lomisinés, nouveau groupe de crustacés anomures. Comptes Rendus des Biffar, T. A. & A. J. Provenzano, Jr., 1972. A reexamination Séances de l’Académie des Sciences, 118: 1353–1355. of Dardanus venosus (H. Milne Edwards) and D. imperator (Miers), with a description of a new species from the western Bouvier, E.-L., 1894b. Sur la transformation des Paguriens en crabes Atlantic (Crustacea, Decapoda, Diogenidae. Bulletin of Marine anomures de la sous-famille des Lithodinés. Comptes Rendus Science, 22: 777–805. Hebdomadaires des Séances de l’Académie des Sciences, 119: 350–352. Birstein, Ya. A. & L. G. Vinogradov, 1972. Crustacea (Decapoda, Anomura, Lithodidae) of the Atlantic sector of the Antarctic, Bouvier, E.-L., 1895a. Recherches sur les affinités des Lithodes & South America and South Africa. Zoologicheskii Zhurnal, 51: des Lomis avec les Pagurides. Annales des Sciences Naturelles, 351–363. Zoologie et Paléontologie, (7)18:157–213. Boas, J. E. V., 1880a. II. Wissenschaftliche Mittheilungen. 1. Bouvier, E.-L., 1895b. Sur une collection de crustacés décapodes Lithodes und Pagurus. Zoologischer Anzeiger, 3: 349–352. recueillis en Basse-Californie par M. Diguet. Bulletin du Muséum d’Histoire Naturelle Paris, 1: 6–8. Boas, J. E. V., 1880b. Studier over decapodernes Slægtskabsforhold. Kongelige Danske Videnskabernes Selskabs Skrifter, Bouvier, E.-L., 1896. Sur un Pagurien nouveau (Eupagurus naturvidenskabelig og mathematisk Afdeling (5)6: 25–210. Chevreuxi) de la Mediterranée. Bulletin du Muséum d’Histoire Naturelle, 2: 95–100. Boas, J. E. V., 1924. Die verwandtschaftliche Stellung der Gattung Lithodes. Konelige Danske Videnskabernes Selskabs Skrifter, Bouvier, E.-L., 1897a. La transformation des Bernards l’Ermite en Biologiske Meddelelser, 4(4): 1–34. Lithodes. Naturaliste (2)19(230): 41–43. Boas, J. E. V., 1926. Zur Kenntnis symmetrischer Paguriden. Bouvier, E.-L., 1897b. Sur deux paguriens nouveaux trouvés par Konelige Danske Videnskabernes Selskabs Skrifter, Biologiske M. Coutière dans réclifs madréporiques à Djibouti. Bulletin du Meddelelser, 5(6): 1–52. Muséum d’Histoire naturelle Paris, 6: 229–233. Boero, F., 2010. The study of species in the era of biodiversity. A Bouvier, E.-L., 1898. Sur quelques Crustacés anomures et tale of stupidy. Biodiversity, 2: 115–126. brachyures recueillis par M. Diguet en Basse-Californie. Bulletin du Muséum d’Histoire Naturelle, Paris, 4: 371–384. Boone, L., 1926a. Unusual deep-sea Crustacea – some forms secured by the Arcturus Oceanographic Expedition. New York Bouvier, E.-L., 1907. Crustacés décapodes nouveaux recueillis à Zoological Society Bulletin, 29: 69–72. Paita (Pérou) par M. le D’Rivet. Bulletin du Muséum d’Histoire Naturelle Paris, 13(2):113–116. Boone, L., 1926b. A new family of Crustacea. Preliminary technical description. New York Zoological Society Bulletin, 29: 73. Bouvier, E.-L., 1914. Les Crustacés de profondeur et les Pygnogonides recueillis par le Pourquoi-Pas? sous la direction Boone, L., 1927. Scientific results of the first oceanographic de M. le Dr Jean Charcot, dans l’Atlantique septentrional, au expedition of the “Pawnee” 1925. Bingham Oceanographic cours de la campagne estivale de 1913. Bulletin du Muséum Collection, 1(2): 1–147. National d’Histoire Naturelle, 20: 215–221. Boone, L., 1930. Scientific results of the cruises of the yachts Bouvier, E.-L., 1918. Sur une petite collection de Crustacés de “Eagle” and “Ara”, 1921–1928, William K. Vanderbilt, Cuba offerte au Muséum par M. de Boury. Bulletin du Muséum commanding. Crustacea: Anomura, Macrura, Schizopoda, National d’Histoire Naturelle, 24: 6–15. Isopoda, Amphipoda, Mysidacea, Cirripedia, and Copepoda. Bulletin of the Vanderbilt Marine Museum, 3: 1–221. Bouvier, E.-L., 1922. Observations complémentaires sur les Crustacés décapodes (Abstraction faite des Caridés) provenant Boone, L., 1932. The littoral crustacean fauna of the Galapagos des Campagnes de S.A.S. le Prince de Monaco. Résultats des Islands. Part II. Anomura. Zoologica, New York, 14: 1–62. campagnes scientifiques accomplies sur son yacht par Albert 1st Prince souverain de Monaco. Fasc. 62: 1–106. Boone, L., 1935. Crustacea and Echinodermata. In: Scientific results of the world cruise of the yacht “Alva”, 1931, Wm. Brandt, J. F., 1848. Die Gattung Lithodes Latreille nebst vier neuer K. Vanderbilt commanding. Bulletin of the Vanderbilt Marine ihr verwandten von Wosnessenski entdeckten, als Typen einer Museum, 6: 1–264. besondern Unterabtheilung (Tribus Lithodea) der Edward’shen Anomuren. Bulletin de la Classe physico-mathmatique de 44 THE RAFFLES BULLETIN OF ZOOLOGY 2010 l’Académie Impériale des Sciences de Saint Pétersbourg, Zélée exécuté par ordre du roi pendant les années 1837–1838– 7(11): 171–176. 1839–1840 sous le commandement de M. Dumont-d’Urville (1842–1854): titles, volumes, plates, text, contents, proposed Brandt, J. F., 1850. Vorlaüfige Bemerkungen über eine aus dates and anecdotal history of the publication. Archives of zwei noch unbeschreibenen Gattungen und arten gebildette Natural History, 27: 407–435. Unterabtheilung (Hapalogastrica) der Teibus Lithodina begleitet von einer Charakteristik der eben genannten Tribus Clément, M. C., 1874. Description d’un pagure nouveau. Bulletin der Anomuren. Bulletin de la Classe physico-mathmatique de la Société d’Etude des Sciences Naturelles de Nimes, 2(4): de l’Académie Impériale des Sciences de Saint Pétersbourg, 155–157. 8(16–17): 268–269. Codreanu, R. & D. Balcesco, 1968. Étude biométrique comparée de Brandt, J. F., 1851. Krebse. In: Middendorff, A. Th. von, Reise in certains caractères dans deux populations du pagure Diogenes den äussersten Norden und Osten Sibiriens während der Jahre pugilator (Roux) de la Mer Noire et de l’Océan Atlantique. 1843 und 1844, 2(1) (Zoologie): 77–148. Bulletin Scientifique de la France et de la Belgique, Paris, 102: 369–383. Brandt, J. F., 1853. Ueber eine neue Art der Gattung Cryptolithodes (Cryptolithodes sitchensis). Bulletin de la Classe physico- Coffin, H. G. 1957. A new southern form of “Pagurus hirsutiusculus” mathmatique de l’Académie Impériale des Sciences de Saint (Dana) (Crustacea, Decapoda). Walla Walla College Publications Pétersbourg, 11(15–16): 653–654. of the Department of Biological Sciences and the Biological Station, 21: 1–8. Brito Capello, F. de, 1875. Appendice à lista dos Crustaceos Decapodios de Portugal. Jornal de Sciências Naturais (Lisboa), Cook, S. D., 1989. Dardanus imbricatus (H. Milne Edwards) and 18: 121–128. descriptions of three new species of Dardanus (Decapoda, Anomura, Diogenidae). Memoirs of the Queensland Museum, Buitendijk, A. M., 1937. Biological results of the Snellius expedition. 27(2): 111–122. IV. The Paguridea of the Snellius Expedition. Temminckia, 2: 251–280. Cormie, A. K., 1993. The morphology of the first zoea stage of Lomis hirta (Lamarck, 1818) (Decapoda, Lomisidae). Crustaceana, Campos, N. H. & R. Lemaitre, 1994. A new Calcinus (Decapoda: 64(2): 249–255. Anomura: Diogenidae) from the tropical western Atlantic, and a comparison with other species of the genus from the region. Costa, A., 1838–1871. Crostacei ed Aracnedi. In: Costa O. G. Proceedings of the Biological Society of Washington, 107(1): & A. Costa (1829-1886), Fauna del Regno di Napoli ossia 137–150. enumerazione di tutti gli Animali che abitano le diverse regioni di questo regno e le acque che le bagnano contenenta la Campos, N. H. & H. Sánchez, 1995. Los cangrejos ermitaños del descrizione de nuovi o poco esattaments conosciuti con figure género Paguristes Dana (Anomura: Diogenidae) de la costa ricavate de originali viventi e dipinte al naturele. Naples. norte colombiana, con la descripcion de dos nuevas espécies. Folia 1–43 (259 pp., 31 pls.), 1–4 (30 pp., 1 pl.), 1–5 (34 pp., Caldasia, 17 (82–85): 569–586. 5 pls.) Carus, J. V., 1885. Coelenterata, Echinodermata, Vermes, Crain, J. A. & P. A. McLaughlin, 2000a. Larval and early Arthropoda. Prodromus faunae Mediterraneae sive descriptio juvenile development in the Lithodidae (Decapoda: Anomura: animalium Maris Mediterranei Incolarum quam comparata silva Paguroidea) reared under laboratory conditions. 1. Subfamily rerum quatenus innotuit adiectis locis et nominibus vulgaribus Lithodinae: Lopholithodes mandtii Brandt, 1848. Invertebrate eorumque auctoribus in commodum zoologorum congessit. Part Reproduction and Development, 37(1): 43–59. 1: 1–524, addenda. Crain, J. A. & P. A. McLaughlin, 2000b. Larval and early Catesby, M., 1743. The Natural History of Carolina, Florida, and juvenile development in the Lithodidae (Decapoda: the Bahama Islands: Containing the Figures of Birds, Beasts, Anomura: Paguroidea) reared under laboratory conditions. 2. Fishes, Serpents, Insects and Plants: Particularly the Forest- Hapalogastrinae: Placetron wosnessenskii Schalfeew, 1892, with Trees, Shrubs and other Plants not hitherto described or very notes on comparative development within the subfamilies of incorrectly figured by Authors. Together with their Descriptions the Lithodidae. Invertebrate Reproduction and Development, in English and French. To which are added, Observations on 37(2): 113–127. the Air, Soil, and Water: With Remarks upon Agriculture, Grain, Pulse, Roots &c, 2: pl. 33, figs. 1, 2. Cunningham, C. W., N. W. Blackstone & L. W. Buss, 1992. Evolution of king crabs from hermit crab ancestors. Nature, Chace, F. A., Jr., 1939. Preliminary descriptions of one new genus London, 355: 539–542. and 17 new species of decapod and stomatopod Crustacea. In: Reports on the scientific results of the first Atlantis expedition Dana, J. D., 1851. Conspectus crustaceorum quae in orbis terrarum to the West Indies, under the joint auspices of the University circumnavigatione, Carolo Wilkes e classe reipublicae foederatae of Havana and Harvard University. Memorias de la Sociedad duce, lexit et descripsit. (Preprint from) Proceedings of the Cubana de Historia Natural, 13: 31–54. Academy of Natural Sciences, Philadelphia, 5: 267–272. Chevreux, E. & E.-L. Bouvier, 1892. Voyage de la goèlette Melita Dana, J. D., 1852a. Conspectus crustaceorum, etc., Conspectus of aux Canaries et au Sénégal. Note préliminaire sur les paguriens. the Crustacea of the Exploring Expedition under Capt. Wilkes, Bulletin de la Société Zoologique de France, 16: 252–256. U.S.N., including the Paguridea, continued, the Megalopidea, and the Macroura. Paguridea, continued, and subtribe Chilton, C., 1911. Crustacea. In: Scientific results of the New Megalopidea. (Preprint from) Proceedings of the Academy of Zealand government trawling expedition, 1907. Records of the Natural Sciences, Philadelphia, 6: 6–28, 1854. Canterbury Museum, 1(3): 285–312. Dana, J. D., 1852b. On the classification of the Corystoidea, Chopra, B. & K. N. Das, 1940. Futher notes on Crustacea Decapoda Paguridea, etc. 2. Conspectus crustaceorum, etc.—Conspectus of in the Indian Musuem. X. On two species of hermit crabs from the Crustacea of the Exploring Expedition under Capt. Wilkes, Karachi. Records of the Indian Museum, 42: 145–153. U.S.N. (Preprint from) American Journal of Science and Arts (2) 23: 119–124. Clark, P. F. & A. Crosnier, 2000. The zoology of the Voyage au pôle sud et dans l’Océanie sur les corvettes l’Astrolabe et la 45 McLaughlin et al.: Checklist of world Lithodoidea, Lomisoidea and Paguroidea Dana, J.D., 1852c. Crustacea, part I. United States Exploring Eldredge, L. G., 1976. Two new species of lithodid (Anomura, Expedition, during the years 1838, 1839, 1840, 1841, 1842, Paguridea, Lithodidae) crabs from Guam. Micronesica, 12(2): under the command of Charles Wilkes, U.S.N., 13: 1–685. C. 309–315. Sherman, Philadelphia (Reprinted Antiquariaat Junk, Lochem, Netherlands, 1972.) Fabricius, J. C. 1775. Systema Entomologiae, sistens insectorum classes, ordines, genera, species, adiectis synonymis, locis, Davie, P. J. F., 2002. Crustacea: Malacostraca: Eucarida (Part 2) descriptionibus , observationibus. Officina Libraria Kortii. Decapoda – Anomura, Brachyura. In: Wells, A.and W.W.K. 832 pp. Houston (eds.), Zoological Catalogue of Australia, 19.3B: 1–641. CSIRO Publishing, Melbourne, Australia. Fabricius, J. C., 1787. Mantissa insectorum sistens eorum species nuper detectas adjectis characteribus genericis, differentiis Dawson, E. W., 1989. King crabs of the world (Crustacea: specificis, emendationibus, observationibus. 1. Hafniae. 348 Lithodidae) and their fisheries. A comprehensive bibliography. pp. New Zealand Oceanographic Institute Miscellaneous Publication 101: 1–338. Fabricius, J. C., 1793. Entomologia systematica emendate et aucta. Secundum, Classes, Ordines, Genera Species, adjectis Dawson, E. W. & J. C. Yaldwyn, 1970. Diagnosis of a new species synonimis, loxcis, observationibus, descriptionibus. 2. Hafniae. of Neolithodes (Crustacea: Anomura: Lithodidae) from New 519 pp. Zealand. New Zealand Journal of Marine and Freshwater Research, 4(2): 227–228. Fabricius J. C., 1798. Supplementum Entomologiae systematicae. 572 pp. Dawson, E. W. & J. C. Yaldwyn, 1971. Diagnosis of a new species of Paralomis (Crustacea: Anomura: Lithodidae) from New Fausto Filho, J., 1970a. Quarta contribuiçáo ao invent rio dos Zealand. Records of the Dominion Museum [New Zealand], Crustáceos Decápodos marinhos do Nordeste Brasileiro. 7(7): 51–54. Arquivos Ciências do Mar, 10(1): 55–60. De Grave, S., N. D. Pentcheff, S. T. Ahyong, T.-Y. Chan, K. A. Fausto Filho, J., 1970b. Pagurus limatulus, nova espécie de Crandall, P. C. Dworschak, D. L. Felder, R. M. Feldmann, C. Crustáceo do Brasil (Crustacea Decapoda: Paguridae). Arquivos H. J. M. Fransen, L. Y. D. Goulding, R. Lemaitre, M. E. Y. Ciências do Mar, 10(1): 69–72. Low, J. W. Martin, P. K. L. Ng, C. E. Schweitzer, S. H. Tan, D. Tshudy & R. Wetzer, 2009. A classification of living and Faxon, W., 1893. Reports on the dredging operations off the west fossil genera of decapod crustaceans. Raffles Bulletin of Zoology, coast of Central America to the Galapagos, to the west coast of Supplement 21: 1–109. Mexico, and in the Gulf of California, in charge of Alexander Agassiz, carried out by the U.S. Fish Commission steamer Dechancé‚ M., 1963. Sur des Paguristes littoraux de l’ouest de “Albatross” during 1891, Lieut.-Commander Z.L. Tanner, U. S. l’océan Indien: P. jousseaumei Bouvier, P. perspicax Nobili N., commanding. VI. Preliminary descriptions of new species et P. abbreviatus sp. nov. (Crustacea Decapoda Paguridea). of Crustacea. Bulletin of the Museum of Comparative Zoology Bulletin du Muséum National d’Histoire Naturelle, (2)35(3): at Harvard College, 24(7): 149–220. 291–301. Faxon, W., 1895. Reports on an exploration off the west coasts of Dechancé‚ M. & J. Forest, 1962. Sur Anapagurus bicorniger A. Mexico, Central and South America, and off the Galapagos Milne Edwards et E. L. Bouvier et A. petiti sp. nov. Bulletin du Islands, in charge of Alexander Agassiz, by the U.S. Fish Muséum National d’Histoire Naturelle, (2)34(4): 293–307. Commission steamer “Albatross,” during 1891, Lieut.- Commander Z.L. Tanner, U. S. N., commanding. XV. The Dechancé, M., 1964. Sur une collection de Crustacés Pagurides stalk-eyed Crustacea. Memoirs of the Museum of Comparative de Madagascar et des Comores. Cahiers O.R.S.T.O.M., série Zoology at Harvard College, 18: 1–292. Océanographie [1964], 2(2): 27–45. Fenizia, G., 1935. I Paguridi del golfo di Napoli. 1. Contributo. Due Degener, O., 1925. Dardanus sanguinocarpus, new species. In: nuove specie di Catapaguroides: C. crassipes - C. acanthodes. Edmondson, C.H., Marine Zoology of Tropical Central Pacific, Annuario del Museo Zoologico della R. Universitá di Napoli Crustacea. Bernice P. Bishop Museum Bulletin, 27: 24–27. (Nuova Serie) 6(18): 1–19. Dixon, C. J., S. T. Ahyong, & F. R. Schram, 2003. A new hypothesis Fenizia, G., 1937. I Paguridi del golfo di Napoli. 2 Contributo. of decapod phylogeny. Crustaceana, 76: 935–975. Specie e varietá dei generi Catapaguroides M. Edwards e Bouvier, Anapagurus Henderson et Nematopagurus M. Edwards Doflein, F., 1900. Weiter Mitteilungen über dekopode Crustaceen e Bouvier. Annuario del Museo Zoologico della R. Universitá der k. bayerischen Staatssammlungen. Sitzungsberichte der di Napoli (Nuova Serie), 7(2): 1–40. Bayerischen Akademie der Wissenschaften, 30: 125–145. Filhol, H., 1883. Note sur quelques espèces nouvelles d’Eupagurus Doflein, F., 1902. Ostasiatische Dekapoden. Abhandlungen der recueillies en Nouvelle-Zélande. Bulletin de la Société Kgl. Bayerischen Akademie der Wissenschaften Math.-Phys. Philomatique de Paris, (7)8 (2): 66–68. Klassen, 21(3): 613–670. Filhol, H., 1884. Exploration sous-marines. Voyages du “Talisman.” Edmondson, C. H., 1925. Marine zoology of tropical central Pacific. La Nature, Paris, 12(568): 326–330. Crustacea. Bernice P. Bishop Museum, Bulletin, 27: 3–62. Filhol, H., 1885a. Description d’un nouveau genre de Crustacés Edmondson, C. H., 1951. Some central Pacific crustaceans. provenant de la Nouvelle-Zélande. Bulletin de la Société Occasional Papers of Bernice P. Bishop Museum, 20(13): Philomatique de Paris (7)9: 47–48. 185–243. Filhol, H., 1885b. Considérations relatives a la faune des Crustacés Edmondson, C. H., 1952. Additional central Pacific crustaceans. de la Nouvelle-Zélande. Bibliothéque de l’École des Hautes Occasional Papers of Bernice P. Bishop Museum, 21(6): Ètudes, Section des Sciences Naturelles, 30(2): 3–60. 67–86. Filhol, H., 1885c. La view au fond des mers. La Nature, Paris, Ejchel, F., 1965. Notes on crustacean decapods. Anais da Academia 13(2): 152–154. Brasileira de Ciências, 37, Supplemento: 325–326. 46 THE RAFFLES BULLETIN OF ZOOLOGY 2010 Filhol, H., 1885d. 1. Zoologie. 3. Crustacés. In: Recherches Forest, J., 1956a. Les Pagures du Viet-Nam. I. Le genre Diogenes zoologiques, botaniques et géologiques faites à l’île Campbell Dana. Bulletin du Muséum National d’Histoire Naturelle, et en Nouvelle-Zelande. Recueil de Mémoires, Rapports et (2)28(6): 524–532. Documents relatifs à l’Observation du Passage de Vénus sur le Soleil du 9 Décembre 1874. Mission de île Campbell. Forest, J., 1956b. Sur une collection de Paguridae de la côte de Zoologie, 3(2)1: 349–510. Institut de France, Académie des l’Or. Proceedings of the Zoological Society of London, 126(3): Sciences, Paris. 335–367. Filhol, H., 1885e. La view au fond des mers. Les explorations sous Forest, J., 1957. Crustacés Décapodes Paguridae d’oceanie française. marines et les voyages du “Travailleur” et du “Talisman”. In: Proceedings of the Eighth Pacific Science Congress, 3A, Bibliothèque de la Nature, Paris. viii, 301 pp. Oceanography and Zoology. Pp. 1053–1076. Fischer P., 1872. Crustacés Podophthalmaires et Cirrhipèdes du Forest, J., 1958a. Les Crustacés Anomoures du Musée royal du department de la Gironde et des côtes du Sub-Ouest de la France. Congo belge. Revue de Zoologie et de Botanique Africaines, Acetes de la Société Linnéenne de Bordeaux, 13: 405–438. 58: 144–168. Fize, A. & R. Serène, 1955. Les Pagures du Vietnam. Institut Forest, J., 1958b. Les Pagures du Viet-Nam. II. Sur quelques Océanographique Nhatrang, Note 45: ix, 1–228. espèces du genre Calcinus Dana. Bulletin du Muséum National d’Histoire Naturelle, (2)30(2): 184–190, 285–290. Forest, J., 1951. Remarques sur quelques Paguridae du genre Calcinus à propos de la description de deux espèces nouvelles Forest, J., 1961. Pagurides de l’Afrique occidentale. Scientific de Polynésie orientale: Calcinus seurati et Calcinus spicatus. Results of the Danish Expedition to the coasts of tropical West Bulletin de la Société Zoologique de France, 76: 83–99. Africa 1945–1946. Atlantide Report, 6: 203–250. Forest, J., 1952a. Remarques sur les genres Diogenes Dana et Forest, J., 1961. In: Pike, R. B., A new bopyrid parasite collected by Troglopagurus Henderson à propos de la description d’un the Chatham Islands 1954 expedition. In: Biological results of Paguridae nouveau de la côte occidentale d’Afrique, Diogenes the Chatham Islands 1954 Expedition. NewZealand Department mercatoris sp. nov. Institut Royal des Sciences Naturelle de of Scientific and Industrial Research, Bulletin, 139(5): 223. Belgique, Bulletin, 28(11): 1–15. Forest, J., 1962. Sur un Dardanus des Hawaii et de Madagascar, Forest, J., 1952b. Contributions a la revision des Crustacés Forest, D. brachyops sp. nov. (Crustacea Paguridea Diogenidae). J., 1952. Sur Trizopagurus caparti gen. et sp. nov., Paguride Bulletin du Muséum National d’Histoire Naturelle, (2)34(5): de la côte occidentale d’Afrique. Bulletin, Institut Royal des 365–370. Sciences Naturelles de Belgique, 28(39): 1–8. Forest, J., 1964. Sur un nouveau genre de Diogenidae (Crustacea Forest, J., 1952c. Notes préliminaires sur les Paguridae (Crustacés Paguridea) de l’Atlantique Sud-Américan, Loxopagurus gen. Décapodes) des côtes occidentales d’Afrique. I. Définition de nov., établi pour Pagurus loxochelis Moreira. Zoologische Pseudopagurus gen. nov. et de Trizopagurus gen. nov. Bulletin Mededelingen, 39: 279–296. du Muséum National d’Histoire Naturelle, (2)24: 254–256. Forest, J., 1966. Crustacés Décapodes: Pagurides. In: Campagne de la Forest, J., 1952d. Contributions a la revision des Crustacés Calypso dans le Golfe de Guinée et aux Iles Principe, São Tomé‚ Paguridae. Paguridae. I. Le genre Trizopagurus. Mémoires et Annobon (1956). Annales de l’Institut Océanographique de du Muséum National d’Histoire Naturelle (A, Zoologie), 5: Monaco, 44: 125–172. 1–40. Forest, J., 1978. Sur deux pagurides nouveaux de l’Atlantique Forest, J., 1952e. Notes préliminaires sur les Paguridae (Crustacés tropical africain: Pagurus laurentae et Paguristes cyanops spp. Décapodes) des côtes occidentales d’Afrique. II. Diagnose nov. Bulletin du Muséum National d’Histoire Naturelle, Paris, sommaire de 6 espèces nouvelles appartenant au genre (3) Zoologie, 356: 525–538. Paguristes Dana. Bulletin du Muséum National d’Histoire Naturelle, (2)24: 257–262. Forest, J., 1984. Révision du genre Aniculus. Crustaceana, Supplement 8: 1–91. Forest, J., 1952f. Notes préliminaires sur les Paguridae (Crust. Décap.) des côtes occidentales d’Afrique. III. Sur un Eupagurus Forest, J., 1987. Les Pylochelidae ou “Pagures symetriques” nouveau de la région de Dakar, E. souriei sp. nov. Bulletin du (Crustacea Coenobitoidea). In: Résultats des campagnes Muséum National d’Histoire Naturelle, (2)24: 355–359. MUSORSTOM. Mémoires du Muséum National d’Histoire Naturelle, Série A, Zoologie, 137: 1–254. Forest, J., 1953. Crustacés Décapodes Marcheurs des îles de Tahiti et des Tuamotu.—I. Paguridea. Bulletin du Muséum National Forest, J., 1989. Sur le genre Bathynarius gen. nov. (Decapoda, d’Histoire Naturelle, (2)25 (5): 441–450. Diogenidae). Bulletin du Muséum National d’Histoire Naturelle, Paris, (4)10A(4): 759–784. Forest, J., 1954a. Crustacés Décapodes Marcheurs des îles de Tahiti et des Tuamotu.—I. Paguridea. Cont. Bulletin du Muséum Forest, J., 1993. Présence du genre Bathynarius (Crustacea, National d’Histoire Naturelle, (2)26(1): 71–79. Decapoda, Diogenidae) en Indonésie et dans le Pacifique central, avec la description de deux espèces nouvelles. Bulletin Forest, J., 1954b. Les Paguristes des côtes occidentales et du Muséum National d’Histoire Naturelle, Paris [1992], méridionales d’Afrique. Annals of the South African Museum, (4)14(A)(2): 483–500. 41(4): 159–213. Forest, J., 1995. Crustacea Decapoda Anomura: Révision du genre Forest, J., 1954c. Sur un pagure littoral nouveau de la Martinique, Trizopagurus Forest, 1952 (Diogenidae), avec l’établissement Paguristes cadenati sp. nov. Bulletin du Muséum National de deux genres nouveaux. In: Crosnier, A. (ed.), Résultalts des d’Histoire Naturelle, (2)26(3): 353–357. Campagnes MUSORSTOM, 13. Mémoires du Muséum National d’Histoire Naturelle, 163: 9–149. Forest, J., 1955. Crustacés Décapodes, Pagurides. Expédition océanographique Belge dans les eaux côtiéres africaines de Forest, J. 1997. Présence du genre Xylopagurus A. Milne-Edwards, l’Atlantique Sud (1948-1949). Résultats scientifique, 3(4): 1880 (Crustacea, Decapoda, Paguridae) dans l’Indo-Ouest 23–147. Pacifique, avec la description de deux espèces nouvelles. Zoosystema, 19: 421–435. 47 McLaughlin et al.: Checklist of world Lithodoidea, Lomisoidea and Paguroidea Forest, J. & J. E. Garcia Raso, 1990. Trizopagurus rubrocinctus Glassell, S. A., 1937. The Templeton Crocker Expedition. XI. sp. nov. des Iles du Cap-Vert (Decapoda, Diogenidae). Bulletin Hermit crabs from the Gulf of California and the west coast du Muséum National d’Histoire Naturelle, Paris (4) 12(A)(1): of Lower California. Zoologica, 22(3): 241–263. 187–195. Glassell, S. A., 1938. New and obscure decapod Crustacea from the Forest, J. & L. B. Holthuis, 1955. Application for a decision west American coasts. Transactions of the San Diego Society regarding the status of the generic name “Pagurus” Fabricius, of Natural History, 8(33): 411–454. 1775 (Class Crustacea, Order Decapoda) and application for the use of the Plenary Powers in regard thereto in certain Glaessner, M. F., 1969. Decapoda. In: Moore, R. C. (ed.), Treatise circumstances. Bulletin for Zoological Nomenclature, 11(10): on Invertebrate Palaeontology. Part R Arthropoda. 2. The 307–321. Geological Society of America & University of Kansas. Pp R400–R651. Forest, J. & P. A. McLaughlin, 1998. Descriptions of two new Japanese hermit crabs (Decapoda: Paguridae: Diogenidae). Gordan, J., 1956. A bibliography of pagurid crabs, exclusive of Proceedings of the Biological Society of Washington, 111(1): Alcock, 1905. Bulletin of the American Museum of Natural 188–198. History, 108: 253–352. Forest, J. & P. A. McLaughlin, 2000. Superfamily Coenobitoidea. Gordon, I., 1935. On two new species of Crustacea from Christmas In: Forest, J., M. de Saint Laurent, P. A. McLaughlin, & Island. Annals and Magazine of Natural History, (1)16: R. Lemaitre, The marine fauna of New Zealand: Paguridea 629–637. (Decapoda: Anomura) exclusive of the Lithodidae. NIWA Biodiversity Memoir, 114: 31–103. Gordon, I., 1959. The dates of publication of parts I-VI of A History of British Crustacea, Thomas Bell. Annals & Magazine of Forest, J. & G. J. Morgan, 1991. A large new species of Dardanus Natural History (13)2: 191–192. (Crustacea, Decapoda, Diogenidae) from southwestern Australia. Bulletin du Muséum National d’Histoire Naturelle, Grant, F. E., 1905. Crustacea dredged of Port Jackson in deep Paris, (4)13(A)(1-2): 203–208. water. Proceedings of the Linnean Society of New South Wales, 1905(2): 312–324. Forest, J. & N. Ngoc-ho, 1992. Description de Pagurus dartevellei (Forest, 1958) (Crustacea, Decapoda, Paguridae). Bulletin du Grant, F. E. & A. R. McCulloch, 1906. On a collection of Crustacea Muséum National d’Histoire Naturelle, Paris, (4)14(A)(1): from the Port Curtis district, Queensland. Proceedings of the 217–227. Linnean Society of New South Wales, 1906: 1–53. Forest, J. & M. de Saint Laurent, 1968. Résultats scientifiques des Grant, F.E. & A.R. McCulloch, 1907. Decapod Crustacea from campagnes de la “Calypso”, Part VII. Campagne de la Calypso Norfolk Island. Proceedings of the Linnean Society of New au large des côtes Atlantiques de l’Amérique du Sud (1961- South Wales, 1907: 151–156. 1962). 6. Crustacés Décapodes: Pagurides. Annales de l’Institut Océanographique de Monaco, n.s. 45(2): 45–172. Greeff, R., 1884. Ueber die Fauna der Guinea-Inseln S. Thomé und Rolas. Sitzungsberichte der Gesellschaft zur Beförderung der Forskål, P., 1775. Descriptiones animalium avium, piscium, gesammten Naturwissenschaften zu Marburg, 2: 41–80. amphibiorum, insectorum , vermium; quae in itinere oriental observ avit. 1–19, 164 pp. Grobben, C. 1878. Beitrage zur Kenntniss der männlichen Geschlechtsorgane der Dekapoda. Arbeiten aus dem Zoologischen García-Gómez, J., 1982. The Provenzanoi group of hermit crabs Institute de Universität Wien und der Zoologischen Sation in (Crustacea, Decapoda, Paguridae) in the Western Atlantic. Part Triest, 1: 1–94. I. Pagurus maclaughlinae, a new species. Bulletin of Marine Science, 32(3): 647–655. Guérin-Méneville, F. E., 1830. Crustacés, arachnids et insects. Chapter 12, Crustacés et arachnids [F.-E.Guérin]. In: Duperrey, García-Gómez, J., 1983. Revision of Iridopagurus (Crustacea: M. L. I., Voyage autour du monde, execute par ordre du roi, sur Decapoda: Paguridae) with the descriptions of new species from la corvette de Sa Majesté, La Coquille, pendant les années 1822, American waters. Bulletin of Marine Science, 33(1): 10–34. 1823 et 1825. Zoologie, 2(2), first division. Pp. 1–319. García-Gómez, J., 1988. A new genus and three new species of Gunn, S. W. & G. J. Morgan, 1992. A new species of Pagurixus hermit crabs (Crustacea: Decapoda: Paguridae) from the western (Crustacea: Decapoda: Paguridae) from southern Australia. Atlantic Ocean. Bulletin of Marine Science, 42(1): 44–64. Memoirs of the Museum of Victoria, 53(1): 31–41. García-Gómez, J., 1994. The systematics of the genus Anapagurus Haan, W., De, 1833–1850. Crustacea. In: Siebold, P. F. von, Henderson, 1886, and a new genus for Anapagurus drachi Fauna Japonica sive Descriptio Animalium, quae in Itinere Forest, 1966 (Crustacea, Decapoda, Paguridae). Zoologische per Japoniam, Jussu et Auspiciis Superiorum, qui Summum Verhandelingen, 295: 1–131. in India Batava Imperium Tenent, Suscepto, Annis 1823-1830 Collegit, Notis, Observationibus et Adumbrationibus Illustravit. Gherardi, F. & P. A. McLaughlin, 1994. Shallow-water hermit crabs Lugduni- Batavorum, Leiden. 243 pp. (Crustacea: Decapoda: Anomura: Paguridea) from Mauritius and Rodrigues Islands, with the description of a new species of Haig, J., 1974a. Calcinus pascuensis, a new hermit crab from Calcinus. Raffles Bulletin of Zoology, 42(3): 613–656. Easter Island (Decapoda, Anomura, Diogenidae). Crustaceana, 27(1): 27–30. Gibbons, W. P., 1855. [no title] Proceedings of the California Academy of Natural Sciences, 1 [second edition]: 47–48. Haig, J., 1974b. Two new species of Pagurus from deep water off Peru and Chile (Decapoda, Anomura, Paguridae). Crustaceana, Gibbs, L. R., 1850. On the carcinological collections of the United 27(2): 119–130. States, and an enumeration of species contained in them, with notes on the most remarkable, and descriptions of new species. Haig, J., 1974c. Observations on the lithodid crabs of Peru, with Proceedings of the American Association for the Advancement description of two new species. Bulletin of the Southern of Science, 3: 167–201. California Academy of Sciences, 73(3): 152–164. Haig, J., 1976. Tomopagurus maclaughlinae, a new hermit crab from the eastern Pacific (Crustacea, Anomura, Paguridae). Bulletin of Marine Science, 26(1): 27–32. 48 THE RAFFLES BULLETIN OF ZOOLOGY 2010 Haig, J., 1977. Description of a new hermit crab (family Paguridae) Heller, C., 1863. Die Crustaceen des südlichen Europa. Crustacea from southern California and Mexico. Proceedings of the Podophthalmia. Mit einer übersicht über die horizontale Biological Society of Washington, 90: 648–657. Verbreitung sämmtlicher europäischer Arten. Wilhelm Braumüller, Wien. 336 pp. Haig, J. & E. E. Ball, 1988. Hermit crabs from northern Australian and eastern Indonesian waters (Crustacea Decapoda: Anomura: Heller, C. 1865. Crustaceen. In: Reise der osterreichischen Paguroidea) collected during the 1975 Alpha Helix Expedition. Fregatte Novara um die Erde in den Jahren 1857, 1858, Records of the Australian Museum, 40: 151–196. 1859 unter den Befehlen des Commodors B. von Wüllerstorf- Urbair. Zoologischer Theil. Kaiserlich-königlichen Hofund Haig, J. & A. W. Harvey, 1991. Three new species of the Pagurus Staatsdruckerei, Wien. 2(3): 1–280. lepidus complex (Decapoda, Anomura, Paguridae) from the eastern Pacific. Contributions in Science, 430: 1–11. Hemming, F. (Ed.), 1958. Official index of rejected and invalid names in zoology, 1: 1–132. London, International Trust for Haig, J. & P. A. McLaughlin, 1984. New Calcinus species Zoological Nomenclature. (Decapoda: Anomura: Diogenidae) from Hawaii, with a key to the local species. Micronesica, 19(1–2): 107–121. Henderson, J. R., 1885. Anomura. In: Tizard, T. H., H. N. Moseley, J. Y. Buchanan & J. Murray, 1885. Narrative of the cruise of Haig, J. & P. A. McLaughlin, 1991. The identity of Pagurus lepidus H.M.S. Challenger with a general account of the scientific (Bouvier) (Decapoda, Anomura, Paguridae) and description of results of the expedition. Report on the Scientific Results ofthe a new eastern Pacific insular species. Contributions in Science, Voyage of H.M.S. Challenger during the Years 1873–76. Vol. 425: 1–12. 1. Neill and Company, Edinburgh. Pp. 897–901. Haig, J. & A. J. Provenzano, Jr., 1965. A new genus and two Henderson, J. R., 1888. Report on the Anomura collected by H.M.S. new species of diogenid hermit crabs (Decapoda, Anomura). Challenger during the years 1873-76. Scientific Results of the Crustaceana, 9(2): 199–207. Exploratory Voyage of HMS Challenger, (Zoology) 27: i–xi, 1–221. Edinburgh etc., Her Majesty’s Stationary Office. Haig, J., T. S. Hopkins & T. B. Scanland, 1970. The shallow water anomuran crab fauna of southwestern Baja California, Mexico. Henderson, J. R., 1893. A contribution to Indian carcinology. Transactions of the San Diego Society of Natural History, 16: Transactions of the Linnean Society of London, 1888-94. (2)5, 13–31. Zoology: 325–458. Hale, H. M., 1941. Decapod Crustacea. In: B.A.N.Z. Antarctic Henderson. J. R., 1896. Natural history notes from H. M. Research Expedition 1929-1931 under the command of Douglas ‘Investigator’ Commander C.F. Oldham, R.N., commanding.— Mawson, Kt. O.B.E., B.E., D.Sc., F.R.S. Reports—Series B Series II., No. 24. Report on the Paguridae collected during (Zoology and Botany) 4(9): 257–286. the season 1893-94. Journal of the Asiatic Society of Bengal, 65(2): 516–536. Hall, S. & S. Thatje, 2009. Four new species of the family Lithodidae (Decapoda: Anomura) from the collections of the Henderson, J. R., 1915. II. Hermit-crabs from the Chilka Lake. National Museum of Natural History, Smithsoniian Institution. Records of the Indian Museum, 11: 25–29. Zootaxa, 2302: 32–47. Hendrickx, M. E. & J. A. Esparza-Haro, 1997. A new species Hansen, H. J., 1908. Crustacea Malacostraca I. In: The Danish of Clibanarius (Crustacea, Anomura, Diogenidaae) from the Ingolf-Expedition, 3(2): 3–120. Bianco Luno, Copenhagen. eastern tropical Pacific. Zoosystema, 19(1): 111–119. Hart, J.F. L., 1930. Some decapods from the south-eastern shores Hendrickx, M. E. & A. W. Harvey, 1999. Checklist of anomuran of Vancouver Island. The Canadian Field-Naturalist, 44(5): crabs from the eastern tropical Pacific. Belgian Journal of 101–109. Zoology, 129(2): 363–389. Hart, J. F. L., 1971. New distributional records of reptant decapod Herbst, J.F.W., 1791–96. Versuch einer Naturgeschichte der Crustacea, including descriptions of three new species of Krabben und Krebse nebst einer systematischen Beschreibung Pagurus, from the waters adjacent to British Columbia. Journal ihrer verschiedenen Arten. 2: 1– 226. Stralsund, Berlin. of the Fisheries Research Board of Canada, 28: 1527–1544. Herbst, J.F.W., 1799–1804. Versuch einer Naturgeschichte der Hart, J. F. L., 1982. Crabs and their relatives of British Columbia. Krabben und Krebse etc. etc., 3 (1): 1–46; 3(2): 1–46; 3(3): British Columbia Provincial Museum Handbook 40: 1–266. 1–54; 3(4): 1–49. Stralsund, Berlin. Harvey, A. W. & P. A. McLaughlin, 1991. Two new hermit crabs Hess, W., 1865. Beiträge zur Kenntnisse der Decapoden-Krebse of the genus Pagurus (provenzanoi group) (Crustacea, Anomura, Ost-Australiens. Druck von Carl Georgi, Bonn. 47 pp. Paguridae) from the eastern Pacific, with notes on their ecology. Contributions in Science, 425: 13–21. Hilgendorf, F., 1869. Crustaceen. In: van der Decken, C. C., Reisen in Ost-Afrika in dem Jahren 1859-1865, 3(1): 69–116. C.F. Haswell, W. A., 1882. Description of some new species of Australian Winter’sche Verlagshandlung, Leipzig, Heidelberg. Decapoda. Linnean Society of New South Wales, 6: 750–763. Hilgendorf, F., 1879. Die von Hrn. W. Peters in Moçambique Hay, W. P., 1917. Preliminary descriptions of five new species of gesammelten Crustaceen. Monatsbericht der Königlich crustaceans from the coast of North Carolina. Proceedings of Preussichen Akademie der Wissenschaften zu Berlin, 1878 the Biological Society of Washington, 30: 71–74. (1879): 782–851. Heller, C., 1861. Beiträge zur Crustaceen-Fauna des rothern Hilton, W. A., 1916. Crustacea from Laguna Beach. Journal of Meers, II. Teil. Sitzungs-Berichte der Mathematisch-Physikisch Entomology and Zoology, 8: 65–73. Klasse der Kaiserlichen Akademie der Wissenschafte, Wien, 44: 241–295. Hogarth, P. J., F. Gherardi & P. A. McLaughlin, 1998. Hermit crabs from the Maldives, with a new species of Catapagurus. Heller, C., 1862. Neue Crustaceen, gesammelt während der Tropical Zoology, 11: 149–175. Weltumseglung der k. k. Fregatte Novara. Zweiter vorläufiger Bericht. Verhandlungen der Kaiserlich-Königlichen Zoologisch- Holmes, S. J., 1895. Notes on west American Crustacea. Proceedings Botanischen Gesellschaft in Wien, 12: 519–528. of the California Academy of Sciences, 4(2): 563–588. 49 McLaughlin et al.: Checklist of world Lithodoidea, Lomisoidea and Paguroidea Holmes, S. J., 1900. Synopsis of the California stalk-eyed Crustacea. and designation under the same Powers of a type species for Occasional Papers of the California Academy of Science, 7: that genus in harmony with established practice. International 1–262. Trust for Zoological Nomenclature, 18(15): 257–273. Holthuis, L. B., 1959. The Crustacea Decapoda of Suriname (Dutch ICZN, 2003. [International Commission on Zoological Guiana). Zoologische Verhandelingen, 44: 1–296. Nomenclature]. Opinion 2052. Pagurus clypeatus Fabricius, 1787 (currently Coenobita clypeatus; Crustacea, Decapoda): Holthuis, L. B., 1961. Report on a collection of Crustacea Decapoda usage conserved by designation of a neotype. Bulletin of and Stomatopoda from Turkey and the Balkans. Zoologische Zoological Nomenclature, 60(3): 235–236. Verhandlingen, 47: 1–67. Ives, J. E., 1892. Crustacea from the northern coast of Yucatan, the Holthuis, L. B., 1962. Forty-seven genera of Decapoda (Crustacea); harbor of Vera Cruz, the west coast of Florida and the Bermuda proposed addition to the official list. Z. N.(S.) 1499. Bulletin Islands. Proceedings of the Academy of Natural Sciences of of Zoological Nomenclature, 19(4): 232–252. Philadelphia, 1892: 176–207. Holthuis, L. B., 1969. Albertus Seba’s “Locupletissimi rerum Jacquinot, H. & H. Lucas, 1854. In: Hombron, J. B. and H. Jacquinot, naturalium thesauri ….” (1734–1765) and the “Planches Crustacea. Zoology, 3 [1853]. Voyage au pole sud et dans de Seba” (1827-1831). Zoologische Mededelingen, 43(19): l’océnie sur les corvettes l’Astrolabe et la Zélée; exécuté de 239–252. ordre du roi pendant les années 1837-1838-1839-1840, sous le commandement De M. J. Dumont-d’Urville, Capitaine de Holthuis, L. B., 2002. A few notes on the authors and dates of Vaisseau, publié par ordre du Gouvernment et sous la direction the names of Crustacea collected by the “Voyage au Pôle Sud supérieure de M. Jacquinot. Gide et Baudry, Paris. 107 pp. et dans l’Océanie sur les corvettes l’Astrolabe et la Zélée. Crustaceana, 75(3–4): 413–422. Jarocki, F. P., von, 1825. Zoologia czyli zwierzetopismo Ogólne, podlug Náynowszego Systematu, 5: 108. Warszawa. Hombron, J. & H. Jacquinot, 1842. Voyage au pole sud et dans l’océnie sur les corvettes l’Astrolabe et la Zélée; exécuté de Kamalaveni, S., 1950. On hermit-crabs (Family Paguridae) in ordre du roi pendant les années 1837-1838-1839-1840, sous the collection of the Indian Museum. Records of the Indian le commandement De M. J. Dumont-d’Urville, Capitaine de Museum, 47: 77–85. Vaisseau, publié par ordre du Gouvernment et sous la direction supérieure de M. Jacquinot. Livraison 2, Crustacés pl. 7. Gide, Kensley, B., 1973. A new species of hermit crab from Natal, South Paris. Africa (Decapoda, Anomura, Paguridae). Durban Museum Novitates, 9(10): 285–290. Hombron, J. & H. Jacquinot, 1846. Voyage au pole sud et dans l’océnie sur les corvettes l’Astrolabe et la Zélée; exécuté de Kensley, B., 1981. The South African Museum’s Meiring Naude ordre du roi pendant les années 1837-1838-1839-1840, sous cruises. Part 12. Crustacea Decapoda of the 1977, 1978, 1979 le commandement De M. J. Dumont-d’Urville, Capitaine de cruises. Annals of the South African Museum, 83(4): 49–78. Vaisseau, publié par ordre du Gouvernment et sous la direction supérieure de M. Jacquinot. Livraison 19, Crustacés pls. 5, 8. Kessler K. T., 1860. Pyteshestvie s zoologhicheskoyu tzyel’yu k Gide et Cie, Paris. syevernomu bereghu Chernagho Morya I v Kruim v 1858 ghodu. (Voyage for zoological purposes to the northern shores of the Hope, G., 1851. Catalogo dei Crostacei italiani e di Molti Altri Black Sea and the Crimea in 1858.) Kiev. 248 pp. del Mediterraneo. Stabilimento Tifografico di Fr. Azzolino, Napoli. Pp. 1–48. Kinahan, J. R., 1857. Remarks on Crustacea collected in Peru, the high seas, and South Australia; with descriptions of undescribed Ingle, R. W., 1985. Northeastern Atlantic and Mediterranean hermit species. Journal of the Royal Dublin Society, 1856/1857: crabs (Crustacea: Anomura: Paguroidea: Paguridae). I. The 328–352. genus Pagurus Fabricius, 1775. Journal of Natural History, 19: 745–769. Kobjakova, S. I., 1955. Novye vidy desyatinogikh rakov (Crustacea, Decapoda) iz yuzhnoi chasti Kurilo-Saxalinskogo raiona. Ingle, R. W, 1993. Hermit crabs of the northeastern Atlantic Ocean Trudy Zoologicheskogo Instituta Akademii Nauk SSSR, 18: and Mediterranean Sea. An illustrated key. Chapman & Hall, 235–242. London, New York. 495 pp. Kobjakova, S. I., 1971. Clibanarius hirsutimanus sp. n., novyi vid. ICZN, 1985. International Code for Zoological Nomenclature, Desyatinogikh rakov (Crustacea, Decapoda) iz Zaliva Poseta Third Edition. Adopted by the XX General Assembly of the Yaponskogo Morya. [Clibanarius hirsutimanus sp. n. (A new International Union of Biological Sciences. International Trust Crustacea Decapoda from the Possjet Bay of the Sea of Japan.) for Zoological Nomenclature, in association of the British Issledovahiya Fauny Morei 8(14): 185–187. (Fauna i Flora Museum (Natural History) London. 338 pp. Zaliva Poseta Yaponskogo Morya.) ICZN, 1999. International Code for Zoological Nomenclature, Komai, T., 1994. Pagurus spina, a new species of hermit crab Fourth Edition. Adopted by the XXI General Assembly of the (Decapoda: Anomura: Paguridae) from Japan. Crustacean Internation Union of BiologicalSciences. International Trust for Research, 23: 23–41. Zoological Nomenclature, in association of the British Museum (Natural History) London. 306 pp. Komai, T., 1995. A new species of the genus Discorsopagurus (Crustacea: Decapoda: Paguridae) from Japan, previously known ICZN, 1957. [International Commission on Zoological Nomenclature]. as D. schmitti (Stevens). Proceedings of the Biological Society Opinion 472. Addition to the Official List of Generic Names in of Washington, 108(4): 617–628. Zoology of the generic name “Pagurus” Fabricius, 1775, with “Cancer bernhardus” Linnaeus, 1758, as types species (Class Komai, T., 1996. Pagurus nigrofascia, a new species of hermit Crustacea, Order Decapoda). International Trust for Zoological crab (Decapoda: Anomura: Paguridae) from Japan. Crustacean Nomenclature, 16(13): 213–276. Research, 25: 59–72. ICZN, 1958. [International Commission on Zoological Nomenclature]. Komai, T., 1997. Pagurus parvispina, a new species of hermit crab Opinion 511. Validation under the Plenary Powers of the generic (Decapoda: Anomura: Paguridae) from northern Japan. Natural name Maja Lamarck, 1801 (Class Crustacea Order Decapoda) History Research, 4(2): 113–123. 50 THE RAFFLES BULLETIN OF ZOOLOGY 2010 Komai, T. 1998a. The taxonomic position of Pagurus gracilipes of a new closely related species (Decapoda: Anomura: (Stimpson, 1858) and Pagurus nipponensis (Yokoya, 1933), and Paguridae). Journal of Crustacean Biology, 16(4): 782–796. description of a new species of Pagurus (Decapoda, Anomura, Paguridae) from Japan. Zoosystema, 20(2): 265–288. Komai, T. & R. Lemaitre, 2002. A new species of Bathypaguropsis McLaughlin, 1994 from Japan, and redescription of B. Komai, T., 1998b. Alloepagurodes spiniacicula, a new genus and kuroshioensis (Miyake, 1978) (Decapoda, Anomura, Paguridae). new species of hermit crab (Decapoda: Anomura: Paguridae) Crustaceana, 75(3–4): 423–441. from Japan. Crustacean Research, 27: 70–81. Komai, T. & E. Myorin, 2005. A new species of Pagurixus Komai, T., 1999. Hermit crabs of the families Diogenidae and (Crustacea: Decapoda: Anomura: Paguridae) from Southern Paguridae (Crustacea: Decapoda: Anomura) collected during the Japan. Zootaxa, 876: 1–12. Shin’yo-maru cruise to the Ogasawara Islands and Torishima Island, oceanic islands in Japan. Natural History Research, Komai, T. & E. Nishi, 1996. A new species of Paguritta (Crustacea: Special Issue, 6: 1–66. Decapoda: Anomura: Paguridae) from the western Pacific, previously confused with P. harmsi or P. gracilipes. Raffles Komai, T., 2000. The identity of Pagurus brachiomastus and Bulletin of Zoology, 44(2): 461–473. descriptions of two new species of Pagurus (Decapoda: Anomura: Paguridae) from the northwestern Pacific. Species Komai, T. & J. Okuno, 2009. Two new species of the hermit crab Diversity, 5: 229–265. genus Pagurixus (Decapoda: Anomura: Paguridae) from the western Pacific. Bulletin of the National Museum of Nature and Komai, T., 2001. A review of the north-western Pacific species Science, (A)(Zoology), Supplement 3: 137–156. of the genus Paguristes (Decapoda: Anomura: Diogenidae), I. Five species initially reported by Ortmann (1892) from Japan. Komai, T. & M. Osawa, 2001. A new distinctive species of pagurid Journal of Natural History, 35: 357–428. hermit crab (Crustacea: Decapoda: Anomura) from Japan. Zoological Science, 18: 1291–1301. Komai, T., 2003a. Identities of Pagurus japonicus (Stimpson, 1858), P. similis (Ortmann, 1892) and P. barbatus (Ortmann, Komai. T. & M. Osawa, 2004. A new hermit crab species of 1892), with description of a new species (Crustacea, Decapoda, Pylopaguropsis (Crustacea: Decapoda: Anomura: Paguridae) Anomura, Paguridae). Zoosystema, 25: 377–411. from the western Pacific and supplemental note on P. laevispinosa McLaughlin and Haig. Zoological Science, 21: Komai, T., 2003b. Reassessment of Pagurus pilosipes (Stimpson), 93–104. supplemental description of P. insulae Asakura, and descriptions of three new species of Pagurus from East Asian waters Komai, T. & M. Osawa, 2005. A new species of Trichopagurus (Crustacea: Decapoda: Anomura: Paguridae). Natural History de Saint Laurent (Crustacea: Decapoda: Anomura: Paguridae) Research, 7(2): 115–166. from the Ryukyu and Yap Islands, and redescription of T. trichophthalmus (Forest). Zootaxa, 801: 1–20. Komai, T., 2003c. A new species of the hermit crab genus Discorsopagurus McLaughlin (Crustacea: Decapoda: Anomura: Komai, T. & M. Osawa, 2006. A review of the Pagurixus boninensis Paguridae) from Japan. Natural History Research, 7(2): species group, with descriptions of six new species (Crustacea: 181–192. Decapoda: Anomura: Paguridae). Zootaxa, 1214: 1–107. Komai, T., 2009a. A review of the northwestern Pacific species of Komai, T. & M. Osawa, 2007. A new species of the hermit crab the genus Paguristes (Decapoda: Anomura: Diogenidae). II. genus Pagurixus Melin (Crustacea: Decapoda: Anomura: Species transferred to the genus Stratiotes, with descriptions of Paguridae) from the Indo-West Pacific. Raffles Bulletin of two new species. Natural History Research, 10(2): 59–92. Zoology, 55: 97–105. Komai, T., 2009b. A new species of the hermit crab genus Komai, T. & M. Osawa, 2009. A new hermit crab species of Catapaguroides (Crustacea: Decapoda: Anomura: Paguridae) the genus Catapagurus (Crustacea: Decapoda: Anomura: from shallow water in the Ryukuy Islands, Japan. Species Paguridae) from the Ryukyu Islands, southern Japan. Zootaxa, Diversity, 14: 197–205. 2211: 57–68. Komai, T., 2010a. A review of the northwestern Pacific species of Komai, T. & D. L. Rahayu, 2004. Redescription of Pagurus the genus Paguristes (Decapoda: Anomura: Diogenidae). III. moluccensis Haig & Ball, 1988, with description of a new Clarification of the identity of a species heretofore referred to species of Pagurus from Indonesia, and taxonomic notes on the Paguristes balanophilus Alcock and descriptions of two new Pagurus anachoretus group (Crustacea: Decapoda: Anomura: species from Japan. Natural History Research, 11(1): 9–33. Paguridae). Raffles Bulletin of Zoology, 52: 183–200. Komai, T., 2010b. New species and new records of the hermit crab Komai, T. & Y. Takada, 2006. A new species of the hermit crab genus Pagurixus Melin, 1939 (Crustacea: Decapoda: Anomura: genus Pagurixus (Crustacea: Decapoda: Anomura: Paguridae) Paguridae) from the Indo-West Pacific. Journal of Natural from shallow coreal reefs of Ishigahi Island, Ryukyu Islands. History, 44(21&22): 1269–1342. Species Diversity, 11(4): 327–337. Komai, T. & A. Asakura, 1995. Pagurixus nomurai, new species, and Komai, T. & M. Takeda, 1996. A new hermit crab of the genus additional record of Pagurixus maorus (Nobili, 1906), hermit Discorsopagurus (Crustacea: Anomura: Paguridae) from Japan. crabs from Kume-jima Island, the Ryukyus, Japan (Decapoda: Species Diversity, 1: 75–85. Anomura: Paguridae). Journal of Crustacean Biology, 15(2): 341–354. Komai, T. & M. Takeda, 2004. Two new deepwater species of hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from Komai, T. & T.-Y. Chan, 2006. A new species of the hermit crab Japan. Bulletin of the National Science Museum, (A)(Zoology), genus Pagurus (Crustacea: Decapoda: Anomura: Paguridae) 30(3): 113–127. from deep waters off Taiwan. Proceedings of the Biological Society of Washington, 119(2): 239–250. Komai, T. & M. Takeda, 2005. First report of Cestopagurus Bouvier (Crustacea: Decapoda: Anomura: Paguridae) from the Pacific Komai, T. & M. Imafuku, 1996. Redescription of Pagurus Ocean, and the description of a new species. Bulletin of the lanuginosus with the establishment of a neotype, and description National Science Museum, (A)(Zoology), 31(3): 93–104. 51 McLaughlin et al.: Checklist of world Lithodoidea, Lomisoidea and Paguroidea Komai, T. & H.-P. Yu, 1999. Identity of Pagurus obtusifrons, with Latreille, P. A. 1831. Cours d’entomologie ou de l’histoire naturelle description of a new species of Pagurus (Decapoda: Anomura: des Crustacés des Arachnides, des Myripodes et des Insectes. Paguridae). Journal of Crustacean Biology, 19(1): 188–205. Exposition méthodique des orders, des familles et des genres des trios premieres classes. Roret, Paris. 568 pp. Kossmann, R. 1878. Notizen über einige neue Crustaceen sowie über neue Fundorte einiger bereits beschrieben. Archiv für Leach, W. E., 1815. Malacostraca Podophthalmata Britanniae; or, Naturgeschichte, 44: 251–258. Descriptions of such British species of the Linnean genus Cancer as have their eyes elevated on footstalks. No. 4 (unpaginated). Kossmann, R., 1880. III Malacostraca (2. Theil: Anomura). London. Zoologische Ergebnisse einer im Auftrage der Königlichen Academie der Wissenschaften zu Berlin ausgeführten Reise in Leach, W. E., 1816. A tabular view of the external characters of die Küsterngebiete des Rothen Meeres. Verlag von Wilhelm four classes of animals, which Linné‚ arranged under Insecta; Engelmann, Leipzig. Pp. 67–140 with the distribution of the genera composing three of these classes into orders, &c., and descriptions of several new genera Krauss, F., 1843. Die Südafrikanischen Crustaceen. Eine and species. Transactions of the Linnean Society of London, Zusammenstellung aller bekannten Malacostraca. Bemerkungen 11: 306–400. über deren Lebensweise und geographische Verbreitung, nebst Beschreibung und Abbildung mehrerer neuen Arten. E. Lemaitre, R., 1982. The Provenzanoi group of hermit crabs Schweizerbart’sche Verlagsbuchhandlung, Stuttgart. 68 pp. (Crustacea, Decapoda, Paguridae) in the Western Atlantic. Part II. Pagurus gymnodactylus, a new species from the Gulf of Lamarck, J. B. P. A., 1801. Systême des animaux sans vertèbres, Mexico and a comparison with Pagurus annulipes (Stimpson). ou tableau général des classes, des ordres et des genres de Bulletin of Marine Science, 32(3): 656–663. ces animaux; présentant leurs caractères essentiels et leur distribution, d’après la considération de leurs rapportsnaturels Lemaitre, R., 1986. Western Atlantic species of the Parapagurus et de leur organisation, et suivant l’arrangemen tétablis dans les pilosimanus complex (Anomura: Paguroidea: Parapaguridae): galeries du Muséum d’Histoire Naturelle, parmi leur dépouilles Description of a new species and morphological variations. conservées; précédé du discours d’ouverture du cours de Journal of Crustacean Biology, 6(3): 525–542. zoologie, donné dans le Muséum National d’Histoire Naturelle l’an de la République. Vol. 11. Déterville, Paris. 432 pp. Lemaitre, R., 1989. Revision of the genus Parapagurus (Anomura: Paguroidea: Parapaguridae), including redescriptions of the Lamarck, J. B. P. A., 1818. Histoire naturelle des animaux western Atlantic species. Zoologische Verhandelingen, 253: sansvertèbres. Vol. 5. Paris. 612 pp. 1–106. Lanchester, W. F., 1902. On the Crustacea collected during the Lemaitre, R., 1993. A new genus of Parapaguridae (Decapoda: “Skeat Expedition” to the Malay Peninsula. Proceedings of the Anomura). Crustacean Research, 22: 11–20. Zoological Society of London, 1902: 363–381. Lemaitre, R., 1994. Crustacea Decapoda: Deep-water hermit crabs Latreille, P. A., 1802. Histoire naturelle, générale at particulière, (Parapaguridae) from French Polynesia with descriptions of four des Crustacés et des Insects, Vol. 3. Paris. 467 pp. new species. In: Crosnier, A. (ed.), Résultats des campagnes MUSORSTOM, 12. Mémoirs de Muséum National d’Histoire Latreille P. A., 1806. Genera Crustaceorum et Insectorum secundum Naturelle, Paris, 161: 375–419. Ordinem naturalem in Familias disposita, Iconibus Exemplisque plurimis explicate. Tomus Primus. Apud Loenig Bibliopola, Lemaitre, R., 1995. A review of the hermit crabs of the genus Parisiis et Argentorati, Paris and Strasbourg. 302 pp. Xylopagurus A. Milne Edwards, 1880 (Crustacea: Decapoda: Paguridae), including descriptions of two new species. Latreille, P. A., 1810. Considérations generales sur l’Ordre Smithsonian Contributions to Zoology,, 570: 1–27. naturel des Animaux composant les Classes des Crustacés, les Archinides et des Insectes: avec un tableau Méthodique de Lemaitre, R., 1996. Hermit crabs of the family Parapaguridae leurs genres disposes en familles. Paris, 144 pp. (Crustacea: Decapoda: Anomura) from Australia: Species of Strobopagurus Lemaitre, 1989, Sympagurus Smith, 1883 Latreille, P. A., 1816. Les Crustacés, les Arachnides et les and two new genera. Records of the Australian Museum, 48: Insectes. In: Cuvier, G., Le règne animal distribute d’après 163–221. son organization, pour server de base à l’histoire naturelle des animaux, et d’introduction à l’anatomie compare, (first Lemaitre, L., 1997. Crustacea Decapoda: Parapaguridae from the edition) 3: 1–653. KARUBAR cruise in Indonesia, with descriptions of two new species. In: Crosnier, A. & P. Bouchet (eds.), Résultats des Latreille P. A., 1817. Macroures, Macroura. Nouveau Dictionnaire Campagnes MUSORSTOM, 16. Mémoires du Muséum national d’Histoire naturelle, 18: 357–358. d’Histoire Naturelle, 172: 573–596. Latreille, P. A., 1818. Pagure. Nouveau Dictionnaire d’Histoire Lemaitre, R., 1998. A new species of hermit crab of the family naturelle, 24: 358–367. Parapaguridae (Decapoda, Anomura) from French Polynesia. Zoosystema, 20(1): 101–107. Latreille, P. A., 1825. Familles naturelles du règne animal, exposées succinctement et dans un ordre analytique, avec l’indication Lemaitre, R., 1999. Crustacea Decapoda: A review of the species of de leurs genres. J.-B. Baillière, Paris. 570 pp. the genus Parapagurus Smith, 1879 (Parapaguridae) from the Pacific and Indian Oceans. In: A. Crosnier (ed.), Résultats des Latreille, P. A., 1825–1828. Histoire Naturelle. Entomologie, ou Campagnes MUSORSTOM, 20. Mémoires du Muséum national Histoire naturelle des Crustacés, des Arachnides et des Insectes. d’Histoire naturelle, 180: 303–378. Encyclopédie Méthodique, 10. Agasse Imprimeur-Libraire, Paris. 832 pp. Lemaitre, R., 2003. A new genus and species of hermit crab (Crustacea: Anomura: Paguridae) from Taiwan. Memoirs of Latreille, P. A. 1829. Crustacés, Arachnides et partie des Insects. Museum Victoria, 60(1): 105–110. In: Cuvier, G., Le règne animal distribute d’après son organization, pour server de base à l’histoire naturelle des Lemaitre, R., 2004. A review of Strobopagurus Lemaitre, animaux, et d’introduction à l’anatomie compare. Deterville, 1989 (Crustacea: Decpoda: Paguroidea: Parapaguridae), Paris. Pp. 1–39. 52 THE RAFFLES BULLETIN OF ZOOLOGY 2010 with description of a new species. Scientia Marina, 68(3): Lenz, H., 1901. Ergebnisse einer Reise nach dem Pacific 355–372. (Schauinsland 1891–1897). Crustaceen. Zoologische Jahrbücher, 14(5): 429–482. Lemaitre, R., 2004. Crustacea Decapoda: A worldwide review of the hermit crab species of the genus Sympagurus Smith, 1883. Lenz, H., 1902. Die Crustaceen der Sammlung Plate. (Decapoda In: Marshall, B. and B. Richer de Forges (eds.), Tropical Deep- und Stomatopoda). Zoologischer Jahrbücher, Supplment, 5: Sea Benthos, 23. Mémoires du Muséum national d’Histoire 731–772. naturelle, 191: 85–149. Lewinsohn, Ch., 1969. Die Anomuren des Roten Meeres (Crustacea Lemaitre, R., 2004. Discovery of the first hermit crab (Crustacea: Decapoda: Paguridea, Galatheidea, Hippidea). Zoologische Decapoda: Parapaguridae) associated with hydrothermal vents. Verhandelingen, 104: 1–213. Cahiers de Biologie marine, 45: 325–334. Lewinsohn, Ch., 1978. Bemerkungen zur Taxonomie von Lemaitre, R., 2006. Two new species of Parapaguridae (Crustacea, Paguritta harmsi (Gordon) (Crustacea Decapoda, Anomura) Anomura, Paguroidea) with subconical corneas, and new data on und beschreibung einer neuen Art der gleichen Gattung aus biology of some rare species. Zoosystema, 28(2): 517–532. Australien. Zoologische Mededelingen, 53(22): 243–252. Lemaitre, R., 2010. A new species of Micropagurus McLaughlin, Lewinsohn, Ch., 1981. Researches on the coast of Somalia. Calcinus 1986 (Decapoda, Anomura, Paguridae) from French Polynesia. tropidomanus n. sp. a new hermit crab from Somalia. Monitore Crustaceana, in press. Zoologico Italiano, n.s. Supplement 14(10): 147–152. Lemaitre, R. & N. H. Campos, 1993. Two new hermit crabs Lewinsohn, Ch., 1982a. Researches on the coast of Somalia. The (Crustacea: Decapoda: Paguridae) from the Caribbean Sea. shore and the dune of Sar Uanle. 33. Diogenidae, Paguridae Proceedings of the Biological Society of Washington, 106(3): and Coenobitidae (Crustacea Decapoda Paguridea). Monitore 554–565. Zoologico Italiano, n.s. Supplment 16: 33–68. Lemaitre, R. & N. Cruz Castaño, 2004. A new species of Pagurus Lewinsohn, Ch., 1982b. Aniculus retipes n. sp. ein neuer Fabricius, 1775 from the Pacific coast of Colombia, with a Einsiedlerkrebs aus dem Roten Meer (Decapoda, Anomura). checklist of eastern Pacific species of the genus. Nauplius, Crustaceana, 42(1): 76–82. 12(2): 71–82. Lewinsohn, Ch., 1982c. Eine neue Art der Gattung Spiropagurus Lemaitre, R. & P. A. McLaughlin, 1995. Alainopagurus crosnieri n. Stimpson (Decapoda, Anomura) aus dem Roten Meer. gen., n. sp. (Decapoda: Anomura: Paguridae) from the western Crustaceana, 42(2): 212–218. Pacific. Bulletin du Muséum National d’Histoire Naturelle, (4)17A(3-4): 273–282. Lilljeborg, W., 1856. Om Hafs-Crustaceer vid Kullaberg i Skåne. (Crustacea marina ad Kullaberg in Scania mense Julii 1852 Lemaitre, R. & P. A. McLaughlin, 1996. Revision of Pylopagurus collecta.) Ofversigt af Kongliga Vetenskapsakademiens and Tomopagurus (Crustacea: Decapoda: Paguridae), with the Forhandlingar. Stockholm, 12: 117–138. descriptions of new genera and species. Part V. Manucomplanus McLaughlin, Anisopagurus McLaughlin, and Protoniopagurus Linnaeus, C., 1758. Systema naturae per regna tria naturae, new genus. Bulletin of Marine Science, 59(1): 89–141. secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis locis, (Edition 10) 1. Holmiae. 824 pp. Lemaitre, R. & P. A. McLaughlin, 2003. Revision of Pylopagurus and Tomopagurus (Crustacea: Decapoda: Paguridae) with Linnaeus, C., 1767. Systema naturae per regna tria naturae, descriptions of new genera and species. Addendum and secundum classes, ordines, genera, species cum characteribus, taxonomic summary. Proceedings of the Biological Society of differentiis, synonymis, locis, (Edition 12) 1. Stockholm. 1327 Washington, 116(2): 464–486. pp. Lemaitre, R. & P. A. McLaughlin, 2003. New species of Lörenthey, I. & K. Beurlen, 1926. Die fossilen Dekapodes Goreopagurus (Decapoda: Anomura: Paguridae) from Tasmania der Länder der Ungarischen Krone. Geologica Hungarica, and reevaluation of sexual tubes in hermit crab systematics. SeriesPalaeontologica, 3: 1–421. Memoirs of Museum Victoria, 60(2): 221–227. Lucas, H., 1846. Part 1. Crustacés, Arachnides, Myriopodes et Lemaitre, R. & P. A. McLaughlin, 2006. A new genus and species Hexapodes. In: Exploration scientifique de l’Agérie pendant of the hermit crab family Paguridae (Crustacea: Decapoda: les années 1840, 1841, 1842. Zoologie. 2. Histoire naturelle Anomura: Paguroidea) from Australia. Zootaxa, 1297: des animaux articulés par H. Lucas, Paris. 403 pp. 57–68. MacDonald, J. D., R. B. Pike & D. I. Williamson, 1957. Larvae Lemaitre, R. & P.A. McLaughlin, 2009. Recent advances and of the British species of Diogenes, Pagurus, Anapagurus and conflicts in concepts of anomuran phylogeny (Crustacea: Lithodes (Crustacea, Decapoda). Proceedings of the Zoological Malacostraca). Arthropod Systematics and Phylogeny, 67(2): Society of London, 128: 209–257. 119–135. MacGilchrist A. C. 1905. Natural history notes from R.I.M.S. Lemaitre, R. & J. Poupin, 2003. A strikingly coloured new species “Investigator” Capt. T.H. Hemming, R.N. (retired), commanding. of Paragiopagurus Lemaitre, 1996 (Crustacea: Decapoda: Series III, No. 6. An account of the new and some of the rarer Anomura: Parapaguridae) from French Polynesia. Zootaxa, decapod Crustacea obtained during the surveying seasons 386: 1–11. 1904-1904. Annals and Magazine of Natural History, (7)15: 233–268. Lemaitre, R. & H. Watabe, 2005. Pagurus ikedai (Crustacea: Anomura: Paguridae), a new hermit crab species of the MacKay, D. C. G., 1932. Description of a new species of crab of bernhardus group from Japanese waters. Zootaxa, 819:1–12. the genus Paralithodes. Contributions to Canadian Biology and Fisheries, 27(A, 24):335–340. Lemaitre, R., P. A. McLaughlin & U. Sorhannus, 2009. Phylogenetic relationships within the Pylochelidae (Decapoda: Anomura: MacLeay, W. S., 1838. On the brachyurous decapod Crustacea Paguridae): A cladistic analysis based on morphological brought from the Cape by Dr. Smith. In: Smith, A., Illustrations characters. Zootaxa, 2022: 1–14. of the Annulosa of South Africa; being a portion of the objects of natural history chiefly collected during an expedition into 53 McLaughlin et al.: Checklist of world Lithodoidea, Lomisoidea and Paguroidea the interior of South Africa, under the direction of Dr. Andrew Malay, M. C. D. & G. Paulay, 2010. Peripatric speciation drives Smith, in the years 1834, 1835. and 1836; fitted out by “The diversification and distributional pattern of reef hermit Cape of Good Hope Association for Exploring Central Africa”. crabs (Decapoda: Diogenidae: Calcinus). Evolution, 64(3): Smith, Elder, and Co., London. Pp. 53–71. 634–662. Macpherson, E., 1982. A new species of Paralomis (Decapoda Man, J. G., De, 1881. On a new collection of podophthalmous Anomura) from the southeasten Atlantic. Crustaceana, 45(2): Crustacea, presented by Mr. J.A. Kruyt, collected in the Red 142–146. Sea near the town of Djeddah. Notes from the Leyden Museum 3: 93–107. Macpherson, E., 1983. Parapagurus hobbiti, new species (Decapoda, Anomura, Parapaguridae), a hermit crab from Man, J. G., De 1887. Uebersicht der indo-pacifischen Arten der the Valdivia Bank, southeast Atlantic. Journal of Crustacean Gattung Sesarma Say, nebst einer Kritik der von W. Hess und E. Biology, 3(3): 472–476. Nauck in den Jahren 1865 und 1880 beschriebenen Decapoden. Zoologische Jahrbücher Systemischer, 2: 639–722. Macpherson, E., 1984. Crustáceos Decápodos del Banco Valdivia (Atlántico sudoriental). Resultados Expediciones Científicas, Man, J. G., De 1888. Bericht über die im indischen Archipel von 12: 39–105. Dr. J. Brock gesammelten Decapoden und Stomatopoden. Archiv für Naturgeschicte (1887): 53: 215–600. Macpherson, E., 1988a. Three new species of Paralomis (Crustacea, Decapoda, Anomura, Lithodidae) from the Pacific and Antarctic Man, J. G., De 1888. Report on the Podophthalmous Crustacea oceans. Zoologica Scripta, 17(1): 69–75. of the Mergui Archipelago, collected for the Trustees of the Indian Museum, Calcutta, by Dr. John Anderson, F.R.S., Macpherson, E., 1988b. Revision of the family Lithodidae Samouelle, Superintendent of the Museum, parts IV and V. Journal of the 1819 (Crustacea, Decapoda, Anomura) in the Atlantic Ocean. Linnean Society, London, 22: 177–240; 241–305. Monografías de Zoología Marina, 2: 9–153. Man, J. G., De, 1892. Decapoden des Indischen Archipels. In: Weber, Macpherson, E., 1988c. Lithodid crabs (Crustacea, Decapoda, M., Zoologischs Ergebnisse einer Reise in Niederlandisch Ost- Lithodidae) from Madagascar and La Réunion (SW Indian Indien. Pp. 265–527. Ocean) with descriptions of two new species. Bulletin de Muséum National Historie Naturelle, Paris, (4)10A(1): Man, J. G., De, 1902. Die von Herrn Professor Kükenthal im Indischen 117–133. Archipel gesammelten Dekapoden und Stomatopoden. In: Kükenthal, W., Ergebnisse einer zoologischen Forschungsreise in Macpherson, E., 1989. A new species of the genus Paralomis den Molukken und Borneo, im Auftrage der Senckenbergischen (Crustacea, Decapoda, Anomura, Lithodidae) from the Indian naturforschenden Gesellschaft. Abhandlungen herausgegeben Ocean. Scientia Marina, 53(1): 117–120. von der Senckenbergischen Naturforschenden Gesellschaft, 25: 465–929. Macpherson, E., 1990. Crustacea Decapoda: On some species of Lithodidae from the western Pacific. In: Crosnier, A. (ed.), Manjón-Cabeza, M. E., J. E. García Raso & J. C. Martínez-Iglesias, Résultats des Campagnes MUSORSTROM, 6. Mémoires de 2002. The genus Paguristes (Crustace: Decapoda: Diogenidae) Muséum National d’Histoire Naturelle (A)145: 217–226. from Cuba (Western Atlantic). A new record and a new species. Scientia Marina, 66(2): 135–143. Macpherson, E., 1991. A new species of the genus Lithodes (Crustacea, Decapoda, Lithodidae) from French Polynesia. Manning, R. B. & F. A. Chace, Jr., 1990. Decapod and stomatopod Bulletin du Muséum National d’Historie Naturelle, Paris (4)13 Crustacea from Ascension Island, South Atlantic Ocean. (A)(1-2): 153–158. Smithsonian Contributions to Zoology, 503: 1–91. Macpherson, E., 1992. Paralomis phrixa (Decapoda, Anomura, Mantelatto, F. L., L. M. Pardo, L. G. Pileggi & D. L. Felder, Lithodidae), a new species from northern Peru, and a key to 2009. Taxonomic re-examination of the hermit crab species the eastern Pacific species of the genus. Crustaceana, 63(3): Pagurus forceps and Pagurus comptus (Decapoda: Paguridae) 313–317. by molecular analysis. Zootaxa, 2133: 20–31. Macpherson, E., 1994. Occurrence of two lithodid crabs (Crustacea: Martens, E, von, 1872. Ueber Cubanische Ctustaceen nach den Decapoda: Lithodidae) in the cold seep zone of the south Sammlungen Dr. J. Grundlach’s. Archiv für Naturgeschichte, Barbados accretionary prism. Proceedings of the Biological Berlin, 38(1): 77–147. Society of Washington, 107(3): 465–468. Martin, J. W. & G. E. Davis, 2001. An updated classification of Macpherson, E., 2001. New species and new records of lithodoid the Recent Crustacea. Natural History Museum of Los Angeles crabs (Crustace, Decapoda) from the southwestern and central County, Science Series 39: 1–124. Pacific Ocean. Zoosystema, 23(4): 797–805. Martínez-Iglesias, J. C. & O. Gómez, 1989. Una especie nueva del Macpherson, E., 2003. Some lithodid crabs ) Crustacea: Decapoda: género Paguristes (Crustacea: Decapoda: Diogenidae) en las Lithodidae) from the Solomon Islands (SW Pacific Ocean), aguas de Cuba. Poeyana, 379: 1–30. with the description of a new species. Scientia Marina, 67(4): 413–418. Mayo, B. S., 1973. A review of the genus Cancellus (Crustacea: Diogenidae) with the description of a new species from the Macpherson, E., 2004. A new species and new records of lithodid Caribbean Sea. Smithsonian Contributions to Zoology, 150: crabs (Crustacea: Decapoda: Lithodidae) from the Crozet and 1–63. Kerguelen Islands area (Subantarctica). Polar Biology, 27: 418–422. McCulloch, A. R., 1909. Studies in Australian Crustacea. No. 2. Records of the Australian Museum, 7: 305–314. Macpherson, E. & T.-Y. Chan, 2008. Some lithodid crabs (Crustacea: Decapoda: Lithodidae) from Taiwan and adjacent McCulloch, A. R., 1913. Studies in Australian Crustacea. No. 3. waters, with the description of one new species from Guam. Records of the Australian Museum, 9: 321–353. Zootaxa, 1924: 43–52. McLaughlin, P. A., 1974. The hermit crabs (Crustacea Decapoda, Makarov, V.V., 1934. Eine neue Gattung der Familie Lithodidae Paguridea) of northwestern North America. Zoologische aus dem Japanischen Meer. Zoologischer Anzeiger, 108(9/10): Verhandelingen, 130: 1–396. 201–204. 54