Protoblastenia incrustans (DC.) J. Steiner var. incrustans

in Rechinger, Verh. zool.-bot. Ges. Wien, 61: 47, 1911. Basionym: Patellaria incrustans DC. in Lamarck & de Candolle - Fl. Franç., éd. 3, 2: 361, 1805.
Synonyms: Biatora rupestris var. incrustans (DC.) A. Massal.; Blastenia incrustans (DC.) A. Massal.; Blastenia rupestris var. incrustans (DC.) Lettau; Caloplaca incrustans (DC.) Flagey; Lecidea rupestris var. incrustans (DC.) Schaer.; Placodium incrustans (DC.) A.L. Sm.; Placodium rupestre f. incrustans (DC.) A.L. Sm.; Protoblastenia rupestris var. incrustans (DC.) Zahlbr.
Distribution: N - VG (Cucchi & al. 2009), Frl (Nimis & Salvadori 1998), Ven (Pinna & al. 1998, Nascimbene & Caniglia 2003c, Nascimbene 2005c, Thor & Nascimbene 2007, Nascimbene & Marini 2007), TAA (Nascimbene 2003, 2008b, Spitale & Nascimbene 2012, Nascimbene & al. 2022), Lomb, Piem (Isocrono & al. 2004), VA (Piervittori & Isocrono 1999), Emil (Fariselli & al. 2020), Lig. C - Tosc (Benesperi 2006, 2007, 2007b, Tretiach & al. 2008), Marc (Nimis & Tretiach 1999), Umb (Nimis & Tretiach 1999, Ravera & al. 2006, Panfili 2007), Laz (Nimis & Tretiach 2004), Abr (Nimis & Tretiach 1999, Gheza & al. 2021), Mol (Nimis & Tretiach 1999, Caporale & al. 2008, Genovesi & Ravera 2014), Sar. S - Camp (Aprile & al. 2003, 2003b, Nimis & Tretiach 2004, Garofalo & al. 2010), Pugl (Nimis & Tretiach 1999), Bas (Nimis & Tretiach 1999), Cal (Puntillo 1996), Si (Nimis & al. 1994, 1996b).
Description: Thallus crustose, endosubstratic, white, hardly differing in colour from the rock. Apothecia orange to orange-yellow, 0.2-0.5 mm across, slightly to strongly constricted at base, totally immersed in pits or slightly protruding from the rock, at first flat, but soon moderately convex and immarginate, the disc smooth. Proper exciple poorly developed, of colourless to yellowish branched hyphae, patchily pigmented; epithecium orange-brown, 15-30 µm high, with granular crystals reacting K+ red; hymenium colourless, 45-55 µm high, hemiamyloid; paraphyses coherent, sparingly branched and anastomosing, 1.5-3.5 µm thick, the apical cells slightly swollen, 2-4.5 µm wide; hypothecium colourless to pale yellow. Asci 8-spored, elongate-clavate, with a thin, outer amyloid layer and a thickened tholus penetrated by a tube, the sides of which stain I/KI+ deep blue, Porpidia-type. Ascospores 1-celled, hyaline, ellipsoid to ovoid, 9-13 x 4-7 µm, without a distinct perispore. Photobiont chlorococcoid. Spot tests: thallus K-, C-, KC-, P-; apothecia K+ red. Chemistry: apothecia with anthraquinones, mainly parietin.
Note: a widespread temperate to circum-arctic lichen, one of the most common species on calcareous rocks in natural habitats throughout the country, with a very wide altitudinal range, reaching the nival belt in the Alps.
Growth form: Crustose endolithic
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: common
Subalpine belt: very common
Oromediterranean belt: very common
Montane belt: extremely common
Submediterranean belt: extremely common
Padanian area: very rare
Humid submediterranean belt: extremely common
Humid mediterranean belt: common
Dry mediterranean belt: rather rare

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples


Felix Schumm - CC BY-SA 4.0
[19678], Germany, Baden-Württemberg, Schwäbische Alb, westlich von Ochsenwang, auf Kalkfelsen im Randecker Maar, 48.57156° N, 9.52611° E, 708 m; TK 7423. Leg. et det. F. Schumm, 18.08.2017.



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (13539)
2001/11/29



Domenico Puntillo; Owner: Domenico Puntillo
Italy


Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php



Felix Schumm - CC BY-SA 4.0
[19678], Germany, Baden-Württemberg, Schwäbische Alb, westlich von Ochsenwang, auf Kalkfelsen im Randecker Maar, 48.57156° N, 9.52611° E, 708 m; TK 7423. Leg. et det. F. Schumm, 18.08.2017.



Felix Schumm - CC BY-SA 4.0
[19678], Germany, Baden-Württemberg, Schwäbische Alb, westlich von Ochsenwang, auf Kalkfelsen im Randecker Maar, 48.57156° N, 9.52611° E, 708 m; TK 7423. Leg. et det. F. Schumm, 18.08.2017.



Felix Schumm - CC BY-SA 4.0
[19678], Germany, Baden-Württemberg, Schwäbische Alb, westlich von Ochsenwang, auf Kalkfelsen im Randecker Maar, 48.57156° N, 9.52611° E, 708 m; TK 7423. Leg. et det. F. Schumm, 18.08.2017.



Felix Schumm - CC BY-SA 4.0
[19678], Germany, Baden-Württemberg, Schwäbische Alb, westlich von Ochsenwang, auf Kalkfelsen im Randecker Maar, 48.57156° N, 9.52611° E, 708 m; TK 7423. Leg. et det. F. Schumm, 18.08.2017.



Felix Schumm - CC BY-SA 4.0
[19678], Germany, Baden-Württemberg, Schwäbische Alb, westlich von Ochsenwang, auf Kalkfelsen im Randecker Maar, 48.57156° N, 9.52611° E, 708 m; TK 7423. Leg. et det. F. Schumm, 18.08.2017.



Felix Schumm - CC BY-SA 4.0
[19678], Germany, Baden-Württemberg, Schwäbische Alb, westlich von Ochsenwang, auf Kalkfelsen im Randecker Maar, 48.57156° N, 9.52611° E, 708 m; TK 7423. Leg. et det. F. Schumm, 18.08.2017.



Felix Schumm - CC BY-SA 4.0
[19678], Germany, Baden-Württemberg, Schwäbische Alb, westlich von Ochsenwang, auf Kalkfelsen im Randecker Maar, 48.57156° N, 9.52611° E, 708 m; TK 7423. Leg. et det. F. Schumm, 18.08.2017.



Felix Schumm - CC BY-SA 4.0
[19678], Germany, Baden-Württemberg, Schwäbische Alb, westlich von Ochsenwang, auf Kalkfelsen im Randecker Maar, 48.57156° N, 9.52611° E, 708 m; TK 7423. Leg. et det. F. Schumm, 18.08.2017.



Felix Schumm - CC BY-SA 4.0
[19678], Germany, Baden-Württemberg, Schwäbische Alb, westlich von Ochsenwang, auf Kalkfelsen im Randecker Maar, 48.57156° N, 9.52611° E, 708 m; TK 7423. Leg. et det. F. Schumm, 18.08.2017.



Curtis Randall Björk, - CC BY-SA 4.0
Sugarbowl-Grizzly Den Provincial Park, British Columbia, Canada
August 2017
as v. tilesii



Pier Luigi Nimis - CC BY-SA 4.0
TSB 41306



Pier Luigi Nimis - CC BY-SA 4.0
TSB 41306



Pier Luigi Nimis - CC BY-SA 4.0
TSB 42166



Pier Luigi Nimis - CC BY-SA 4.0
TSB 42166


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Protoblastenia_incrust_incr.htm
France, 18/9/2015 - Vercors


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Protoblastenia_incrust_incr.htm
France, 18/9/2015 - Vercors


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Protoblastenia_incrust_incr.htm
France, 18/9/2015 - Vercors