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A revision of Licuala (Arecaceae, Coryphoideae) in Borneo

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Summary

This paper presents an account of Licuala Wurmb in Borneo based on herbarium studies and the author’s own fieldwork. The revision enumerates 46 species of Licuala found in the island of Borneo. Of these, 26 are new species and a further two are new varieties. Forty-two are Bornean endemics, reperesenting 91% endemism in Licuala on the island. A key to the species is provided and an index of specimens examined is included. Conservation assessments have also been made for all of the taxa found in Borneo. Twenty-eight taxa (57%) are threatened.

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Acknowledgements

I am grateful for support from the British High Commission’s Chevening Scholarships Scheme: Royal Society — Malaysian Fellowship, which supported the study between January and June 2000. Without the Fellowship, it would have been difficult to complete this revision. I am also very grateful to the Royal Botanic Gardens, Kew (K) for providing the necessary institutional help and support while I was there. Dr John Dransfield and his lovely wife Soejatmi gave constant help and encouragement during my stay at K. John further provided the translation for the Latin diagnoses. I wish also to express my appreciation to Dr William J. Baker for his help and good company while working in the Palm Room at Kew. I also wish to thank the curators of the following herbaria that provided loans and access to herbarium material during the study: AAU, BH, BM, BO, BR, BRUN, CAVA, CP, E, FI, K, L, PNH, SAN, SAR and SING. My sincere thanks to Lucy Smith and Joseph Pao for the illustration of the plates. I also wish to thank Wendy Yong who helped prepare the distribution maps for the conservation assessments and Dato’ Dr Abd. Latif Mohmod, the Director-General of the Forest Research Institute Malaysia for supporting botanical work in the institute. Finally, I would like to thank the two reviewers who have made constructive comments to improve the manuscript.

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  1. Forest Research Institute Malaysia, 52109, Kepong, Selangor, Malaysia

    L. G. Saw

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  1. L. G. Saw
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Correspondence to L. G. Saw.

Appendix 1

Appendix 1

Alphabetical list of collections studied

The symbols in round brackets (( )) are the codes for the herbaria (according to Index Herbariorum) from which the specimens have been seen. In the list below, only the main collector is listed. The number in square brackets ([ ]) indicates the taxon as in the following list: 1. atrovirens; 2. bidentata; 3. bintulensis; 4. borneensis; 5. bruneiana; 6. burretii; 7. campestris; 8. chaiana; 9. collina; 10. cordata var. cordata; 11. cordata var. ashtonii; 12. densiflora; 13. elliptica; 14. exigua; 15. flammula; 16. glabra; 17. hallieriana; 18. intermedia; 19. kuchingensis; 20. lambii; 21. lanata; 22. leopoldii; 23. leucocarpa; 24. maculata; 25. mattanensis var. mattanensis; 26. mattanensis var. paucisecta; 27. meijeri; 28. micholitzii var. micholitzii; 29. micholitzii var. leeana; 30. miriensis; 31. mukahensis; 32. orbicularis; 33. oryzoides; 34. paludosa; 35. petiolulata; 36. pilosa; 37. pseudovalida; 38. punctulata; 39. reptans; 40. rheophytica; 41. rubiginosa; 42. sabahana; 43. sarawakensis; 44. spathellifera; 45. spicata; 46. spinosa; 47. triphylla; 48. valida; 49. yiiana.

Aban SAN 95384 (K, L, SAN) [18], Ambriansyah AA 588 (K, L) [48], Amin SAN 115021 (K, KEP, L) [46], SAN 115121 (E, K, KEP) [48], Ampuria SAN 19087 (SAN) [27], SAN 32706 (K) [27], Andersen 167 (KEP, SAN) [41], 168 (KEP) [48], Anderson 55 (SING) [19], 201 (K) [16], 4318 (K) [9], 4558 (K, SAR) [9], 8998 (SAR) [19], 9101 (K, L) [25], 10196 (SAR, SING) [25], S 20807 (SAR) [26], S 27308 (SAR) [2], S 27309 (K, SAR) [35], S 27512 (BH, K, SAR) [26], Ariffin 16 (K, SING) [4], Arifin BRF 1808 (BO, L) [48], Ashton A 497 (K) [3], A 499 (K) [3], BRUN 5277 (K) [48], S 18062 (K, SAR) [48], S 18263 (K, L, SAR) [11], S 19431 (K, SAR) [35], S 19437 (K, L, SAR) [3], S 19442 (K, L, SAR, SING) [35], S 21571 (BH, K, L, SAR) [3], S 21575 (BH, K, SAR) [48], S 22021 (K, L, SAR) [48], Atar s.n. 26 July 1996 (K) [34], Au S 16751 (L, SAR) [30], S 16769 (K, SAR) [48], S 16783 (K, L, SAR) [3], S 16794 (K, L, SAR) [30], Awang SAN 68980 (K) [7], SAN 109786 (SAN) [6], SAN 111101 (K) [48], Awmack 26 (K) [2], 47 (K) [2], 111 (K) [17], 242 (K) [37], 327 (BO, K) [17], 328 (K) [17], 329 (K) [17], 330 (K) [17], 352 (BO, K) [2].

Bakar SAN 28356 (SAN) [42], SAN 32827 (SAN) [48], Baker 699 (SING) [2], 713 (K, KEP, SAR) [40], 719 (K, KEP, SAR) [24], 746 (K, KEP, SING) [26], Balajadia 2891 (K) [46], 3717 (K, SAN) [46], Beaman 11497 (K) [9], 11862 (K) [19], 11967 (K) [2], Beccari PB 24 (FI) [43], PB 404 (FI) [34], PB 580 (FI) [32], PB 1230 (FI) [43], PB 1379 (FI, K) [2], PB 1786 (FI) [32], PB 1834 (FI) [45], PB 2044 (FI) [25], PB 2133 (FI) [2], PB 2432 (FI) [16], PB 2568 (FI) [25], PB 3642 (FI) [35], PB 3646 (FI, K, P) [25], 3646 (FI, K) [25], PB 3699 (FI, K) [44], PB 3790 (FI) [10], PB 3985 (FI) [35], PB 4032 (FI) [3], PB 4033 (FI) [35], PB 4034 (FI) [48], PB 4035 (FI) [4], Bell 2043 (BM) [16], Bernstein JHB 60 (K) [34], JHB 61 (K) [5], JHB 80 (K) [34], JHB 168 (K) [34], Blicher S 57992 (AAU) [35], Bogner 1535 (K) [2], Borchsenius SAN 145532 (SAN) [18], Brooke 9252 (L) [11], Brunig 2 (SING) [12], 6 (SING) [48], 8 (SING) [35], 9 (SING) [45], 18 (SING) [45], 25 (SING) [45], 26 (SING) [48], 27 (SING) [48], 32 (SING) [35], 49 (SING) [48], 53 (SAR, SING) [48], S 2467 (SING) [32], S 2468 (SING) [2], S 4681 (SAR) [2], S 8715 (SAR) [1], S 8766 (K, SAR) [1], S 9911 (SAR) [2], S 9945 (SAR) [2], S 12383 (SAR, SING) [48], S 12399 (K, SAR, SING) [35], S 17503 (SAR) [2], S 17539 (K, SAR) [35], Bujang 13024 (K, SAR) [46], 13025 (K, SAR) [32], Burtt 5576 (E) [9].

Chai S 18526 (K, L, SAR) [3], S 18914 (K, L, SAR, SING) [35], S 19321 (K, SAR) [37], S 19328 (K, SAR) [35], S 31144 (K, L, SING) [41], S 32135 (L, SAR) [37], S 33197 (L, SAR) [35], S 34635 (BH, L, SAR) [2], S 36813 (K, SAR) [13], S 39527 (K, KEP, L, NY, SAN, SAR) [21], S 40051 (K, KEP, L, SAR) [8], S 40055 (K, SAR) [36], S 43401 (K, KEP, L, SAN, SAR) [3], Chew CWL 1111 (L, SING) [9], Chin 2875 (K) [48], Christensen 1094 (AAU) [35], 1410 (AAU) [43], Church 254 (BO) [48], 1529 (BO) [48], Clemens 20472 (B, K, NY, SAR) [38], Cockburn SAN 66001 (K) [48], SAN 66205 (SAN) [35], SAN 82504 (SAN) [46], SAN 85033 (SAN) [7], Coode 6412 (K) [48], 6831 (K) [5], 7329 (K) [5], Cowley 37 (K) [41], Creagh s.n. Aug. 1895 (K) [48], Cuadra A 229 (K, KEP, L, SING) [7].

Dayang Awa S 46353 (K, SAR) [16], S 51191 (K, L, SAR) [9], Dewol SAN 55975 (K) [7], SAN 93583 (K) [42], SAN 93689 (K, L) [42], SAN 97029 (K, SAN) [7], SAN 100209 (K) [48], SAN 108872 (K, L, SAN) [27], Dius DT 5 (K, KEP) [48], Doinis 208 (K) [48], Down s.n. July 1900 (SING) [32], Dransfield 761 (BH, K) [35], 762 (BH, K) [2], 775 (K) [48], 790 (K) [44], 794 (BH, K) [5], 798 (K) [35], 1573 (BO, L) [48], 2117 (BO, K, L) [48], 2978 (BO, L) [48], 4351 (BO, K) [48], 4388 (BO, K, L) [2], 4436 (BO, K) [48], 4731 (K, KEP) [2], 4740 (K, SAR) [2], 5302 (K) [9], 5311 (K, L, SAN, SAR) [21], 5324 (K, L) [9], 5620 (K) [48], 5632 (K) [48], 5758 (K, SAN) [47], 5759 (K, SAN) [18], 5808 (K, L) [42], 5852 (SAN) [18], 5865 (BH, K, SAR) [2], 5890 (K, L, SAR) [26], 5913 (BH, K, SAR) [2], 5952 (BH, K) [48], 5962 (K, SAR) [48], 6001 (BO, K, L, SAR) [38], 6008 (K, L) [16], 6020 (K, L) [3], 6022 (K) [25], 6023 (BH, K) [2], 6033 (BO, K, L, SAR) [26], 6038 (K, L, SAR) [35], 6048 (K, L, SAR) [37], 6079 (K, SAR) [2], 6094 (BH, K, SAR) [32], 6123 (K, NY, SAR) [13], 6134 (K, SAR) [2], 6350 (E, K) [48], 6533 (K) [5], 6541 (K) [5], 6596 (K) [48], 6598 (K) [5], 6605 (K) [48], 6629 (K, SING) [41], 6668 (K) [48], 6728 (K) [3], 6804 (K) [48], 6892 (K) [48], 6927 (K) [48], 6929 (K, KEP) [41], 7070 (K) [34], 7138 (K) [34], 7415 (K) [48], 7457 (K) [41].

Elmer 21635 (BM, BO, CAVA, CP, K, L, PNH, SING) [47], Endert 3784 (BO, L) [48], 4795 (BO, L) [48].

Fidilis SAN 82195 (K, KEP, L) [48], SAN 82195 (K) [48], SAN 119711 (K, KEP) [18], Forman 430 (BO) [48], 900 (K) [48], Fraser 131 (K) [46], Fuchs 21212 (K, L, SAR) [48].

Gale 1 (AAU) [35], 2 (AAU) [35], 3 (AAU) [35], 4 (AAU) [3], 4 (AAU) [3], 5 (AAU) [42], 6 (AAU) [48], 7 (AAU) [48], 8 (AAU) [5], 9 (AAU) [5], 10 (AAU) [3], 11 (AAU) [5], George SAN 117634 (K, KEP, L) [7], Gibot SAN 30647 (K, SAN) [37], Giesen 66 (L) [34].

Hallier 1650 (BO, FI) [35], 1992 (BO, FI, L) [39], 3374 (BO, FI) [17], Hansen 888 (SAR) [10], 1069 (SAR) [2], Haslani 33 (K) [5], Henderson SFN 38963 (K, SING) [7], 38963 (SING) [7], Hewitt s.n. 9 Nov. 1911 (K) [2], s.n. 1908 (BO, K) [2], s.n. Sept. 1901 (K) [48], 41 (K) [35], 42 (K) [16], 43 (K) [35], 542 (SAR) [26], 543 (SAR) [2].

Ilias S 16637 (K, L, SAR, SING) [32], S 16638 (K, L, SAR, SING) [35], S 16654 (K, L, SAR, SING) [35], S 35661 (BH, K, SAR) [35], S 39115 (SAR) [31], S 46096 (SAR) [26].

Jacobs 5656 (L) [3], Jaheri 1 (BO, FI) [48], 215 (BO) [16], 1436 (BO) [35], Jais S 12157 (SAR, SING) [35], James S 33472 (BH, SAR) [19], S 34530 (K, L, SAR) [35], S 42114 (L, SAR) [35], S 42118 (K, KEP, L, SAR) [2], S 42118 (KEP) [2], S 42133 (SAR) [26], Jarvie 5050 (BO) [35], 5129 (BO) [17], 6210 (BO) [17], Jemree S 73775 (KEP) [48], S 78635 (SAR) [9], Jibrin 86 (K, KEP) [6], JS 118 (KEP) [7], Jidwin JJ 86 (K, KEP) [48], Jugah S 23808 (BH, SAR) [35], S 34376 (K, SAR) [35], Julia SBC 1231 (SBC) [26].

Kato B 4766 (BO, L) [35], Keith 6714 (K, SING) [42], 9082 (K, SING) [35], 10241 (K) [34], Kessler 2079 (L) [16], 2474 (L) [35], 2570 (L) [16], Kiew 42 (KEP) [34], 46 (KEP) [45], 54 (KEP) [2], 56 (SAR) [2], 57 (KEP) [2], 80 (KEP) [48], 81 (KEP) [35], 85 (SAR) [35], 449 (KEP) [9], 851 (KEP) [35], 852 (KEP) [35], 872 (SAR) [29], 892 (KEP) [26], 906 (SAR) [3], 929 (KEP) [29], 931 (SAR) [29], 937 (KEP) [23], Kinsun 138 (K, KEP) [6], 253 (KEP) [7], Kloss SFN 18674 (SING) [18], SFN 18674 (SING) [42], Kohyama K 10597 (BO, L) [15], Kokawa 294 (SAN) [27], Kostermans 4088 (BO) [48], 4250 (BO) [47], 4740 (BO) [2], 7017 (BO, K, L) [16], 7768 (BO, L) [48], 10136 (BO) [16], Kulip SAN 141004 (SAN) [27], Kutuh 8-20 (K) [25], 29-4 (K) [12], 35-10 (K) [2], 40-12 (K) [3], 46-14 (K) [2], 49-4 (K) [12], 56-3 (K) [43], 57-13 (K) [3], 58-5 (K) [37], 59-7 (K) [35], 62-16 (K) [43], 63-32 (K) [2], 74-41 (K) [13], 74-17 (K) [13], 95-34 (K) [26], 105-49 (K) [26], 132-61 (K) [48], 135-67 (K) [26], 141-70 (K) [25].

Lagman TL 144 (K) [2], Lai S 69665 (KEP) [35], S 72699 (SAR) [30], S 74651 (SAR) [9], S 75323 (KEP, SAN, SAR) [35], S 76965 (KEP, SAR) [9], S 76966 (SAR) [9], S 77210 (KEP, SAR) [35], S 80842 (SAR) [30], S 83501 (SAR) [32], Lee S 41907 (K, L, SAR) [29], S 43205 (K, SAR) [2], S 45428 (SAR) [18], S 45523 (SAR) [10], S 53353 (K, SAR) [35], S 53844 (K, SAR) [35], S 54011 (K, L, SAR) [2], S 54117 (K, KEP, L, SAR) [2], Lim LSP 1261 (SAN) [7], SAN 145401 (KEP, SAN, SAR, SING) [6], Lobb s.n. 1853 (K) [2], s.n. 1853 (K) [2], s.n. 1853 (K) [2], s.n. 1853 (K) [25], s.n. 1853 (K) [43], s.n. 1857 (K) [43], s.n. 1857 (K) [43].

Madani SAN 82640 (K) [48], SAN 130688 (K, KEP, SAN) [22], Maidin 2338 (K) [7], Martin S 36981 (K, KEP, L, SAR) [32], S 36982 (BH, K, SAR) [2], S 38049 (K, KEP, L, SAN) [9], McDonald 3517 (BO) [48], Meekiong SBC 1962 (SBC) [26], Meijer 1060 (BO, L) [48], 1920 (BO) [34], SAR 14852 (SAN) [35], SAN 19588 (K, SAN) [27], SAN 21238 (K) [42], SAN 22583 (K, L) [48], SAN 25461 (SAN) [27], SAN 27610 (SAN) [7], SAN 28376 (K, L, SAN) [48], SAN 28454 (SAN) [42], SAN 43794 (SAN) [18], SAN 49844 (SAN) [27], SAN 50009 (SAN) [22], SAN 141453 (SAN) [18], Meliden 23 (K, KEP) [48], Micholitz s.n. Sept. 1903 (SING) [28], 11848 (FI, K) [2], Mitsuru 12519 (BRUN, SAR) [34], 12654 (BRUN, SAR) [5], 13450 (BRUN, SAR) [3], 15708 (SAR) [48], Mogea ITTO/BA 59 (SAR) [40], ITTO/BA 539 (SAR) [40], ITTO/BA 547 (SAR) [40], 775 (BO, K, KEP) [46], ITTO/BA 893 (SAR) [40], ITTO/BA 895 (SAR) [40], 1785 (BO, K) [48], 1826 (BO) [48], 1841 (BO, K) [48], 2578 (BO, K, L) [48], 3460 (BO) [29], 3461 (BO, K) [17], 3507 (BO, K, L) [48], 3616 (BO, K) [48], 3837 (BO) [17], 4079 (BO) [17], 4291 (BO) [17], 4292 (BO, L) [48], 4293 (BO) [48], 4419 (BO) [17], Mohidin S 21663 (K, SAR) [8], Mohtar S 49366 (K, L, SAR) [21], Mondi 249 (BO, L) [35], Moore 9079 (K) [25], 9081 (BH, K) [2], 9107 (BH) [2], 9132 (BH, K) [2], 9145 (BH, K) [3], 9146 (BH, K) [44], 9163 (BH, K) [42], 9176 (BH) [48], 9186 (BH, K) [34], 9192 (BH, K) [48], 9213 (BH) [48], SAN 32827 (SAN) [48], Motley 1203 (K) [34], Muas S 13003 (SAR) [35], Munting S 55604 (K, L, NY, SAR) [25].

Nais SNP 5539 (KEP) [48], Native collector 2253 (FI, K) [2], Nielson 1067 (AAU, K) [41], 1102 (AAU, K) [35], Nooteboom 4114 (L) [17], 4268 (L) [17]. Othman S 55797 (K) [1], S 61756 (K) [37].

Pearce S 78171 (SAR) [40], S 89248 (SAR) [4], Pereira SAN 43679 (SAN) [48], Pickles 2914 (BM, L, SAR, SING) [35], 3695 (BM, L, SAR) [37], Polak 422 (BO) [34], Poulsen 149 (AAU, K) [35], Puasa SAN 10136 (K, SING) [7], FMS 48910 (KEP) [7], Purseglove P 4575 (SING) [48], P 4746 (K, L, SING) [16], P 4800 (K, SING) [38], P 5054 (SING) [2].

Rantai S 70078 (SAR) [26], S 72992 (KEP, SAR) [48], S 74160 (SAR) [40], S 74229 (KEP) [2], S 74232 (KEP, SAR) [32], S 74236 (KEP, SAR) [2], S 90577 (SAR) [40], Reza s.n. 22 Sept. 1997 (SAN) [7], Richards 1821 (K) [9], Ridley s.n. July 1893 (K) [2], s.n. July 1893 (K) [2], s.n. July 1893 (K) [2], s.n. Jan. 1915 (K) [2], s.n. July 1893 (K, SING) [26], s.n. 1893 (SING) [32], s.n. Jan. 1915 (K) [35], 12398 (K, SING) [2], 12400 (K) [26], 12407 (K, SING) [12], Ridsdale 1938 (K, L) [48], Runi S 59694 (SAR) [14], S 71205 (K, KEP, SAR) [32].

Said BRUN 17368 (K) [48], BRUN 17369 (K) [48], Saikeh SAN 82374 (K, SAN) [6], Salleh BRUN 15340 (SING) [5], Sands 5435 (K) [34], 5483 (K) [35], Sani 33 (K) [48], SS 53 (K, KEP) [48], SS 175 (K, KEP) [48], SS 177 (KEP) [48], 181 (K, KEP) [48], 412 (K) [48], 627 (K) [48], Saw s.n. 18 March 2004 (KEP) [45], FRI 37491 (KEP) [2], FRI 37495 (KEP) [2], FRI 37497 (KEP) [28], FRI 37498 (KEP) [9], FRI 37499 (KEP) [9], FRI 37500 (KEP) [12], FRI 37522 (KEP) [2], FRI 37523 (KEP) [43], FRI 37691 (KEP) [42], FRI 37692 (KEP) [48], FRI 37693 (KEP) [42], FRI 39828 (SAR) [40], FRI 39832 (SAR) [35], FRI 39914 (KEP) [35], FRI 39981 (K, KEP, SAN) [42], FRI 40156 (KEP) [48], FRI 40161 (KEP) [10], FRI 40188 (KEP) [48], FRI 40208 (KEP) [10], FRI 40216 (KEP) [48], FRI 40283 (KEP, SAN) [48], FRI 44521 (KEP, SAN) [48], FRI 44566 (K, KEP, SAN) [42], FRI 44907 (KEP) [41], FRI 44908 (KEP) [48], FRI 48168 (KEP) [2], FRI 48169 (KEP) [43], FRI 48170 (KEP) [25], FRI 48172 (AAU, K, KEP, SAR) [2], FRI 48173 (AAU, K, KEP, SAR) [26], FRI 48174 (KEP) [4], FRI 48175 (AAU, K, KEP, SAR) [40], FRI 48176 (KEP, SAR) [19], FRI 48177 (KEP, SAR) [24], FRI 48178 (KEP, SAR) [28], FRI 48194 (KEP) [20], S 94701 (KEP, SAR) [2], S 94702 (K, KEP, SAR) [2], S 94703 (K, KEP, SAR) [2], S 94704 (K, KEP, SAR) [2], S 94705 (SAR) [25], S 94706 (KEP) [32], S 94707 (AAU, K, KEP, SAR) [2], S 94708 (K, KEP, SAR) [38], S 94709 (KEP, SAR) [15], S 94710 (KEP, SAR) [15], S 94711 (AAU, K, KEP, SAR) [2], S 94712 (AAU, K, KEP, SAR) [2], S 94713 (KEP, SAR) [35], S 94715 (KEP, SAR) [19], S 94716 (KEP, SAR) [26], Schatz 3267 (K) [35], Shaevy SBC 501 (SBC) [26], Shea SAN 77287 (BO) [48], Sibat S 23673 (BH, SAR) [48], Sie S 43891 (K, SAR) [26], Sigin SAN 99757 (SAN) [33], Simpson 2130 (K) [6], Singh SAN 31071 (K, SAN) [7], SAN 39294 (SAN) [27], Slooten 2239 (BO) [48], Soedjito 1695 (BO) [47], Suzuki K 14789 (KEP) [5].

Talib SAN 86134 (K, KEP, L, SAR) [48], Teo S 77309 (K, KEP, SAR) [31], S 77320 (SAR) [31], Teysmann 283 (BO) [39], Tomlinson 2 (SING) [35], 12 (SAR, SING) [2], Tong S 34176 (L, SAR) [2], S 34177 (BH, SAR) [35], S 34178 (SAR) [32].

Utteridge 496 (SAN) [42].

van Balgooy W 917 (BO, L) [48], 5384 (BO, K, L) [2], VB 6086 (K) [48], Veitch & Sons 370-1885 (KEP) [32].

Watson, 370-1885 (K) [32], Whitmore 3131 (K) [48], 3138 (K) [21], Winkler 2630 (FI) [48], 3323 (FI) [48], Wong 497 (K, SING) [3], 498 (K) [48], 521 (K, SING) [34], 527 (K, SING) [5], 553 (K) [3], 981 (K, SING) [3], 1124 (K, SING) [5], 1182 (K) [41], 1239 (K, SAN, SAR, SING) [35], 1310 (K) [41], 1642 (K, SAR, SING) [48], 1645 (K, KEP, L, SAR, SING) [34], 1683 (K, SING) [48], 1684 (K) [48], 2103 (K, L, SAR, SING) [5], 2187 (E, KEP, SAN, SAR) [27], 2309 (L) [48].

Yahud S 81985 (SAR) [9], Yeo S 38482 (K, SAR) [48], Yii S 42003 (K, L, SAR) [15], S 42276 (L, SAR) [2], S 43445 (K, L, SAR) [2], S 45901 (K, SAR) [2], S 45912 (K, L) [16], S 46000 (K, SAR) [35], S 48305 (K, SAR) [35], S 50311 (K, L, SAR) [26], S 53428 (K) [48], S 53601 (K, L, SAR) [35], S 55818 (K, SAR) [49], S 58681 (KEP, SAR) [9], S 58736 (KEP, SAR) [9], S 60523 (KEP, SAR) [48], S 67565 (KEP, SAR) [3], S 70901 (K, KEP, SAR) [48].

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Saw, L.G. A revision of Licuala (Arecaceae, Coryphoideae) in Borneo. Kew Bull 67, 577–654 (2012). https://doi.org/10.1007/s12225-012-9414-0

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