When restoration work began at Pleasant Valley Conservancy, there were a few fine prairie remnants and some degraded savannas, woodlands, and wetlands. However, grazing and other agricultural activities have caused a marked loss in plant species.

In addition to removing many invasive plants, we worked hard to increase the diversity of native plant species. This work has been quite successful. From an early estimate of about 250 native plant species, our list has grown to over 400 species.

The number 250 is derived from a merger of plant checklists made by botanists who visited Pleasant Valley Conservancy before any restoration work had been carried out. These lists, presented below, were edited to remove any duplicates. The total list includes some weeds and other invasive species, has 270 species. It is estimated that about 250 of these species are plants native to prairies, savannas, woodlands, and wetlands.

The early plant species checklists given below were done by different people at different times, and some of the Latin names used, even for the same species, may be different. Plant nomenclature is a complicated field, and different authorities use different names. Continuing research in plant taxonomy leads to altered nomenclature. Therefore, some of the names used in these checklists may no longer be valid.

Early Species List by Brian Pruka

The first detailed plant checklist for the land that was to become Pleasant Valley Conservancy was carried out by plant ecologist Brian Pruka in 1995. Pruka had just finished his Master’s thesis at the University of Wisconsin-Madison working on a nearby oak savanna, and was familiar with the area. His list was prepared for us as part of the early management plan that Pruka prepared for the property. Pruka’s list contains 170 species, which include prairie, savanna, woodland, and wetland species.

Even considering the nomenclature problems mentioned above, and the fact that surveys were made only in the summer, there are some noteworthy species on the Pruka list, including a number with relatively high Coefficients of Conservatism.

Species List by Paul West

In 1997, we hired plant ecologist Paul West, then working only part-time at the Nature Conservancy, to further develop management objectives for the property. As part of this work, West prepared another plant checklist. Although there was some overlap with Pruka’s list, West’s list, with 136 species, added some additional species.

Kathie Brock’s Species List for Goat Prairie (Unit 1)

Our best prairie remnant, Unit 1, was probably never grazed. In 2002 Kathie Brock prepared a species list for this prairie. At that time, this unit had been burned six times. Noteworthy prairie species on Kathie’s list of 37 species include sky-blue aster, side oats grama, purple prairie clover, showy and Illinois tick-trefoil, tall boneset, flowering spurge, false boneset, small yellow flax, fringed puccoon, violet wood sorrel, little bluestem, small skullcap, blue-eyed grass, gray goldenrod, Indian grass, prairie dropseed, spiderwort, and bird’s foot violet.

Unintroduced Species Missing From the Above Lists

There are a number of species that were originally present at Pleasant Valley Conservancy that are not given in any of these early lists. We know these species were present because they are present now and we did not introduce them. Of these, the most interesting are: White wild indigo, glade mallow, Canada anemone, wood anemone, poke milkweed, silky aster, white wild indigo, Bromus kalmii, large yellow lady-slipper orchid, shooting star, wood betony, bottlebrush grass, silky rye, Virginia wild rye, upland boneset, sweet Indian plantain, prairie alum-root, great St. John’s wort, yellow star grass, seneca snakeroot, lion’s foot, rosinweed, cup plant, showy goldenrod, elm-leafed goldenrod, yellow pimpernel, Culver’s root, purple meadow-rue, angelica, spiderwort, bellwort, prairie violet, golden Alexanders, and purple milkweed. Several of these species are endangered, threatened, or of special concern.

The presence of purple milkweed (Asclepias purpurascens), a State endangered species, is noteworthy. This beautful savanna species was first found flowering in Unit 12B after it had been cleared and burned. With further clearing and burning, additional sites developed. Seeds of this species were never brought in from elsewhere, although efforts have been made (partly successfully) to get it established in new sites on the Conservancy.

It turned out that a significant number of native species were present on our property along County Highway F. This north-facing slope had been periodically mowed by county highway crews, but this mowing had been infrequent enough that some important species were not eradicated. Species that were present in relatively large amounts included New Jersey tea, golden Alexanders, arrow-leafed aster, great St. John’s wort, great blue lobelia, fringed loosestrife, and Culver’s root.

Increasing Species Diversity

Originally, our goal was to collect seed from the remnants and transfer it to other sites on the property. This work went in tandem with the removal of invasive shrubs and trees. The focus initially was on the south-facing slope where large amounts of red cedar, planted red pines, elms, buckthorn, honeysuckle, and other woody plants were removed. Removal of these trees opened up “holes”, into which seeds collected elsewhere on the property were planted. This sort of activity is ongoing throughout the Conservancy.

As much as possible, we have used seeds that had been collected on our property. If no useful source was found there, we collected seeds from nearby town roads, or from other sites nearby. In only a few cases were seeds obtained from more distant sites, although always from southern Wisconsin sources.

In 1998, our first year of planting, we planted 20 species on the south slope that had been collected elsewhere on the property. Only the lower part of the slope, which lacked any significant savanna oaks, had been cleared. It was burned in early April and planted a few days later, with seeds that had been stored in the barn all winter. We had only limited amounts of grass species, so this seeding was primarily forbs. As it turned out, burning had stimulated the growth of pre-existing grasses, and little bluestem and Indian grass flourished without planting.

Controlled burns played a major role in increasing diversity, and the south slope has been burned almost every year since restoration began. The photo here is from a burn in March 2003.

First Planted Prairie At the same time, we were planting completely new prairies on the various small agricultural fields present at the site. These small fields, amounting to a total of less than 15 acres, had been entered into the Conservation Reserve Program (CRP) in 1987. When our 10-year contract came up for renewal in 1997, we opted to plant prairie on two of these fields, which gave us extra points in the evaluation of our CRP project. Instead of purchasing seeds, we decided, as much as possible, to collect our seeds. Because our supply of seeds was limited, the first year we planted only a single field of prairie (Toby’s Prairie). Although almost all the species planted came from elsewhere on the property, a few species that “should” have been in a tallgrass prairie were completely missing. These were obtained by trade from the Madison Audubon Society prairie restorations at Goose Pond. What we traded was volunteer work for seeds. Species that we obtained from this source were: wild onion, Canada milk vetch, white prairie clover, pale purple coneflower, ox-eye sunflower, compass plant, and stiff goldenrod. All the rest of the 70 species planted came from our own sources.

Indian grass, Sorghastrum nutans, is an important species in a tallgrass prairie. It produces a good prairie sod, and it provides great fuel for a prairie burn. All of our Indian grass has come from our own property. Initially, it was present in significant amounts only on a small prairie remnant high on the south slope (Unit 4), surrounded by cedars and buckthorn, This tiny prairie of less than 1/2 acre was first burned in April 1998 and as a result the Indian grass flourished. Seed was collected in the fall of 1998 and used as part of the prairie seed mix for Toby’s Prairie, planted in November 1998. It has done exceeding well there and has been the source of Indian grass seed for all of our subsequent planted prairies, as well as for the open savanna areas.

2007: What Eleven Years Have Wrought

Restoration work at Pleasant Valley Conservancy has proceeded at a steady pace. As degraded savannas have been cleared, they have been planted with seeds collected elsewhere on the site. The planted prairies have turned out to be excellent seed sources because of the large acreages of relatively level land where collecting is easy.

Only minimal work has been done in the wetlands, which were initially in fairly good shape. Minimal work has also been done on the north-facing oak woods, which have plentiful spring ephemerals and, except for a few areas, only small amounts of invasive shrubs.

As of the end of the 2007 season, over 400 native species are present at Pleasant Valley Conservancy. Even more significant, the floristic quality index (FQI) has a value of 91.3, which is an impressively high number.

Most of the species used for the FQI calculation were not introduced but were already here, although perhaps present in small numbers. The most important work that we have done is to remove the invasives, both woody and herbaceous, thus providing a habitat for pre-existing species to spread. Controlled burns also have played a major role.

We have increased diversity in another way, by learning to identify the plants and by exploring the Conservancy in great detail. Often a rare species will be present in only one or a few sites and will be missed in any casual walk-through. Further, restoration of an area will almost immediately result in the appearance of species that have not been seen in that site before. They might well have been present but were suppressed by the great weight of invasives.

As an example, shooting star, a very desirable prairie and savanna plant, can survive under the shade of exotic bush honeysuckle but is not readily visible. Removal of the honeysuckles leads almost immediately, usually in the first subsequent growing season, to a great increase in the growth of shooting stars, many of which will flower (see photo below). These were not planted but burst into view after their habitat was restored.

Brian Pruka’s Check List from 1995

LatinCommon
Acer negundoBox elder
Acer saccharumSugar maple
Achillea millefoliumYarrow
Actaea albaWhite baneberry
Actaea rubraRed baneberry
Adiantum pedatumNorthern maidenhair fern
Agalinis gattingeriRound-stemed false foxglove
Agalinis purpurea v. parvifloraSmooth false foxglove
Agrimonia gryposepalaTall agrimony
Ambrosia trifidaGiant ragweed
Amorpha canescensLead-plant
Amphicarpaea bracteataHog peanut
Andropogon gerardiiBig bluestem
Anemone canadensisMeadow anemone
Anemone cylindricaThimbleweed
Anemone quinquefoliaWood anemone
Anemone virginianaTall anemone
Antennaria plantaginifoliaPlantain-leaved pussytoes
Apocynum sibiricumClasping dogbane
Aquilegia canadensisWild columbine
Arabis lyrataSand cress
Aralia nudicaulisWild sarsaparilla
Aralia racemosaSpikenard
Aristida tuberculosaDune three-awn grass
Armoracia lacustrisLake cress
Artemisia campestris caudataBeach wormwood
Artemisia frigidaPrairie sagewort
Asclepias hirtellaTall green milkweed
Asclepias viridifloraShort green milkweed
Asparagus officinalisAsparagus
Asplenium trichomanesMaidenhair spleenwort
Aster sagittifoliusArrow-leaved aster
Bidens frondosaCommon beggar’s-tick
Botrychium dissectum obliquumOblique grape fern
Botrychium minganenseMingan moonwort
Botrychium multifidumLeather-leaved grape fern
Brassica spMustard
Cacalia plantagineaPrairie Indian plantain
Calamagrostis strictaBog reed-grass
Calamovilfa longifolia longifoliaSand-reed
Carex sppSedge
Carex torreyiTorrey sedge
Carex trisperma 
Celtis occidentalisHackberry
Conium maculatumPoison hemlock
Coptis trifolia groenlandicaGoldthread
Corydalis sppCorydalis
Crataegus chrysocarpaFireberry hawthorn
Cynosurus cristatusDogtail
Cyperus odoratusFragrant cyperus
Cyperus squarrosusAwned cyperus
Cystopteris fragilisNorthern fragile fern
Cystopteris protrusaCreeping fragile fern
Dasistoma macrophyllaMullein foxglove
Desmodium paniculatumPanicled tick-trefoil
Eleocharis ovataBlunt spike-rush
Eleocharis tenuis borealisNeat spike-rush
Equisetum palustreMarsh horsetail
Equisetum variegatum jesupiE. X TRACHYODON
Equisetum x litoraleE. ARVENSE X FLUVIATILE
Erigeron philadelphicusMarsh fleabane
Erigeron pulchellusRobin’s plantain
Erigeron spFleabane
Erigeron sppFleabane
Erigeron strigosusDaisy fleabane
Eriophorum spCotton-grass
Euthamia graminifoliaGrass-leaved goldenrod
Euthamia gymnospermoidesWestern flat-top goldenrod
Festuca brachyphyllaShort-leaf fescue
Festuca spFescue grass
Festuca sppFescue grass
Fraxinus americanaWhite ash
Fraxinus nigraBlack ash
Fumaria officinalisFumitory
Galium lanceolatumLance-leaved wild licorice
Gentiana spGentian
Goodyera pubescensRattlesnake plantain
Gymnocladus dioicaKentucky coffee tree
Hedeoma hispidaRough pennyroyal
Helianthus divaricatusWoodland sunflower
Helianthus sppSunflower
Helianthus tuberosusJerusalem artichoke
Hordeum sp 
Hudsonia tomentosaFalse heather
Hypochaeris radicataSpotted cat’s ear
Juncus alpinoarticulatusRichardson’s rush
Juncus gerardiBlack grass
Koeleria macranthaJune grass
Lechea pulchellaPinweed
Lepidium perfoliatumClasping cress
Liatris x gladewitziiHYBRID BLAZING STAR
Linum medium texanumSmall yellow flax
Linum rigidumStiff yellow flax
Lithospermum spPucoon
Lycopodium inundatumBog clubmoss
Menispermum canadenseMoonseed
Mentha aquatica crispaCurled mint
Microseris cuspidataPrairie dandelion
Myosotis scorpioidesCommon forget-me-not
Myriophyllum spWater milfoil
Myriophyllum tenellumDwarf milfoil
Neslia paniculataBall mustard
Oenothera perennisSmall sundrops
Oenothera pilosellaPrairie sundrops
Orchis spectabilisShowy orchis
Panax trifoliumDwarf ginseng
Panicum latifoliumBroad-leaved panic-grass
Panicum miliaceumBroom-corn millet
Panicum oligosanthesFew-flowered panic-grass
Panicum oligosanthes scribnerianumScribner’s panic-grass
Penstemon grandiflorusLarge-flowered beard-tongue
Penstemon sppBeard-tongue
Platanthera laceraRagged fringed orchid
Polygonum hydropiperoidesMild water
Potamogeton oakesianusOakes’ pondweed
Potentilla argutaPrairie cinquefoil
Potentilla norvegicaRough cinquefoil
Prenanthes spWhite lettuce
Ptelea trifoliataHop-tree
Pycnanthemum tenuifoliumSlender mountain mint
Pyrola chloranthaGreen shinleaf
Pyrola ellipticaLarge-leaved shinleaf
Pyrola spShinleaf
Quercus bicolorSwamp white oak
Quercus spOak
Ranunculus abortivusSmall-flowered buttercup
Ranunculus gmeliniiSmall yellow water-crowfoot
Ranunculus hispidusRough buttercup
Ranunculus spButtercup
Ribes hudsonianumCanadian black currant
Ribes uva-crispiGarden gooseberry
Rorippa sinuataSpreading yellow-cress
Rosa blandaEarly wild rose
Rubus flagellarisCommon dewberry
Rubus setosusBristly blackberry
Rumex acetosellaField sorrel
Salix cordataDune willow
Salix lucidaShining willow
Salix myricoidesBlue-leaved willow
Scirpus cyperinusWoolgrass
Scirpus validusGreat bulrush
Silene csereiGlaucous campion
Silene dichotomaForked catchfly
Silene latifoliaWhite campion
Silene vulgarisBladder campion
Sisymbrium officinaleHedge-mustard
Sisyrinchium sppBlue-eyed grass
Smilax herbacea herbaceaCarrion flower
Solanum carolinenseHorse nettle
Sonchus arvensisPerennial sow thistle
Sparganium fluctuansFloating-leaved bur-reed
Stellaria borealisNorthern starwort
Stellaria sppStarwort
Streptopus roseus longipesTwisted stalk
Symphoricarpos spSnowberry
Talinum rugospermumFame flower
Thelypteris phegopterisNarrow beach-fern
Triadenum virginicumMarsh St. John’s wort
Trientalis borealisStarflower
Trifolium campestreLow hop-clover
Trifolium repensWhite clover
Utricularia geminiscapaTwin-stemmed bladderwort
Utricularia spBladderwort
Verbena simplexNarrow-leaved vervain
Verbena spVervain
Vernonia fasciculataCommon ironweed
Vincetoxicum nigrumBlack swallow-wort
Viola conspersaDog violet
Viola sppViolet
Vitis sppWild grape

Paul West’s Check List From 1997-98

Latin nameCommon name
Acer rubrumRed maple
Achillea millefoliumYarrow
Actaea rubraRed baneberry
Adiantum pedatumNorthern maidenhair fern
Agalinis asperRough false foxglove
Agastache spGiant hyssop
Ambrosia artemisiifoliaCommon ragweed
Amorpha canescensLead-plant
Amphicarpaea bracteataHog peanut
Andropogon gerardiiBig bluestem
Anemone cylindricaThimbleweed
Anemone quinquefoliaWood anemone
Anemone virginianaTall anemone
Antennaria neglectaField pussytoes
Antennaria plantaginifoliaPlantain-leaved pussytoes
Apocynum androsaemifoliumSpreading dogbane
Aquilegia canadensisWild columbine
Aralia nudicaulisWild sarsaparilla
Aralia racemosaSpikenard
Aristida tuberculosaDune three-awn grass
Artemisia biennisBiennial wormwood
Asclepias ovalifoliaOval milkweed
Asclepias spMilkweed
Asclepias syriacaCommon milkweed
Asclepias tuberosaButterfly weed
Asclepias viridifloraShort green milkweed
Asplenium spSpleenwort
Asplenium trichomanesMaidenhair spleenwort
Aureolaria grandifloraYellow false foxglove
Bidens discoideaSwamp beggar’s-tick
Bidens frondosaCommon beggar’s-tick
Botrychium dissectum obliquumOblique grape fern
Calamovilfa longifolia longifoliaSand-reed
Calylophus serrulatusToothed evening-primrose
Cardamine spCress
Carex albolutescens 
Carex leptalea 
Carex meadiiMead’s sedge
Carex pauciflora 
Carex rostrata 
Carex sppSedge
Carex torreyiTorrey sedge
Carya ovataShagbark hickory
Celtis occidentalisHackberry
Chenopodium gigantospermumMaple-leaved goosefoot
Coptis trifolia groenlandicaGoldthread
Corydalis sppCorydalis
Cymbalaria muralisKenilworth ivy
Cyperus lupulinusSand cyperus
Cyperus odoratusFragrant cyperus
Cyperus squarrosusAwned cyperus
Cystopteris fragilis mackayiMackay’s frafile fern
Cystopteris protrusaCreeping fragile fern
Dasistoma macrophyllaMullein foxglove
Desmodium paniculatumPanicled tick-trefoil
Eleocharis flavescens olivaceaTufted spike-rush
Equisetum palustreMarsh horsetail
Equisetum variegatum nelsoniiE. X NELSONII
Eriophorum spCotton-grass
Euphorbia marginataSnow-on-the-mountain
Euthamia gymnospermoidesWestern flat-top goldenrod
Fagopyrum esculentumBuckwheat
Festuca pratensisMeadow fescue
Filipendula rubraQueen-of-the-prairie
Fraxinus nigraBlack ash
Fumaria officinalisFumitory
Galium lanceolatumLance-leaved wild licorice
Galium tinctoriumStiff bedstraw
Gentiana sppGentian
Goodyera repens ophioidesCreeping rattlesnake plantain
Halenia deflexaSpurred gentian
Hedeoma hispidaRough pennyroyal
Helianthus divaricatusWoodland sunflower
Helianthus mollisDowny sunflower
Hieracium spHawkweed
Hieracium sppHawkweed
Hypochaeris radicataSpotted cat’s ear
Isopyrum biternatumFalse rue anemone
Juncus gerardiBlack grass
Juncus squarrosus 
Lespedeza virginicaSlender bush clover
Liatris x gladewitziiHYBRID BLAZING STAR
Linum rigidumStiff yellow flax
Lithospermum latifoliumAmerican gromwell
Lycopodium inundatumBog clubmoss
Malaxis monophyllosWhite adder’s mouth
Menispermum canadenseMoonseed
Myosotis scorpioidesCommon forget-me-not
Orchis rotundifoliaSmall round-leafed orchis
Orchis spectabilisShowy orchis
Oryzopsis racemosaBlack-seeded ricegrass
Oxalis acetosellaNorthern wood-sorrel
Panicum latifoliumBroad-leaved panic-grass
Penstemon grandiflorusLarge-flowered beard-tongue
Pinus banksianaJack pine
Platanthera blephariglottisWhite fringed orchid
Platanthera laceraRagged fringed orchid
Poa nemoralis 
Polygonella articulataJointweed
Polygonum erectumErect knotweed
Potamogeton oakesianusOakes’ pondweed
Potentilla argutaPrairie cinquefoil
Potentilla norvegicaRough cinquefoil
Potentilla sppCinquefoil
Ptelea trifoliataHop-tree
Puccinellia distansAlkali grass
Pycnanthemum tenuifoliumSlender mountain mint
Pyrus malusApple
Ranunculus abortivusSmall-flowered buttercup
Ranunculus gmeliniiSmall yellow water-crowfoot
Rheum rhabarbicumRhubarb
Rumex obtusifoliusBitter dock
Sagittaria spArrowhead
Salix cordataDune willow
Salix lucidaShining willow
Scirpus supinus halliiSharp-scale bulrush
Senna hebecarpaWild senna
Silene vulgarisBladder campion
Silene vulgarisBladder campion
Sisymbrium officinaleHedge-mustard
Sisyrinchium fuscatumBlue-eyed grass
Solanum carolinenseHorse nettle
Stellaria sppStarwort
Streptopus roseus longipesTwisted stalk
Talinum rugospermumFame flower
Triadenum virginicumMarsh St. John’s wort
Trientalis borealisStarflower
Trifolium repensWhite clover
Triosteum perfoliatumEarly horse gentian
Utricularia spBladderwort
Verbena simplexNarrow-leaved vervain
Vernonia fasciculataCommon ironweed
Vicia americanaAmerican vetch
Vincetoxicum nigrumBlack swallow-wort
Viola sppViolet
Vitis sppWild grape

Kathie Brock’s 2002 Prairie List

Latin nameCommon name
Amorpha canescensLead-plant
Andropogon gerardiiBig bluestem
Anemone cylindricaThimbleweed
Antennaria neglectaField pussytoes
Aquilegia canadensisWild columbine
Asclepias verticillataWhorled milkweed
Aster oolentangiensisSky-blue aster
Bouteloua curtipendulaSide oats grama
Campanula rotundifoliaHarebell
Dalea purpureumPurple prairie clover
Desmodium canadenseShowy tick-trefoil
Desmodium illinoenseIllinois tick-trefoil
Erigeron pulchellusRobin’s plantain
Erigeron strigosusDaisy fleabane
Eupatorium altissimumTall (Upland) boneset
Euphorbia corollataFlowering spurge
Geranium maculatumWild geranium
Kuhnia eupatorioidesFalse boneset
Linum medium texanumSmall yellow flax
Lithospermum incisumFringed puccoon
Monarda fistulosaWild bergamot
Oxalis acetosellaNorthern wood-sorrel
Oxalis violaceaViolet wood-sorrel
Panicum latifoliumBroad-leaved panic-grass
Panicum sp.Small-seed panic grass
Ratibida pinnataYellow coneflower
Rudbeckia hirtaBlack-eyed Susan
Schizachyrium scopariumLittle bluestem
Scutellaria leonardiSmall skullcap
Sisyrinchium sppBlue-eyed grass
Solidago canadensisCommon goldenrod
Solidago nemoralisOld-field goldenrod (grey; dyer)
Sorghastrum nutansIndian grass
Sporobolus heterolepisPrairie dropseed
Tradescantia ohiensisCommon spiderwort
Verbena strictaHoary vervain
Viola pedataBird’s foot violet

Detailed picture of the oak savanna diversity at Pleasant Valley Conservancy

A summary of the species found in the various savanna units is given in a PDF that can be downloaded. The species in this file are sorted by C value (Coefficient of Conservatism), which puts the most interesting species on top.

Oak savanna diversity PDF