Tropical Marine Mollusc Programme (TMMP)
Lexical approach to CARDIACEA
Records of taxa
Illustrated and annotated records of living and fossil shells,
with emphasis on the families Cardiidae and Lymnocardiidae
(Mollusca: Bivalvia)
Jorgen Hylleberg
Part 3: N-Z
Editor of the TMMP Proceedings
Prof Dr Jorgen Hylleberg, Aarhus University, Denmark
Assistant Editor
Dr Anuwat Nateewathana, Thailand
Advisory Editorial Board
Director Oonchit Bhatiyasevi, PMBC, Phuket, Thailand
Prof. Dr. Barbara Brown, University of Newcastle, UK
Chief of Reference Collection Dr Somchai Bussarawit, Phuket, Thailand
Phuket Marine Biological Center Special Publication vol. 30 (2004)
RECORDS N-Z
naganoensis Hayami, 1959. - Unknown species. »
Regarding Cardium naganoensis: Cardiidae. Lower
Liassic. REcORDs. "1960 ZR: Cardium naganoensis
Hayami, 1960 sp. nov. Lias, Higashinagano, Japan.
"1973 Keen : Cardium naganoensis Hayami, 1959.
Incertae sedis.
nahali (subspecies) . - See under caspia nahali TadjalliPour, 1977.
nalivkini Kolesnikov, 1929. - Changed combination:
Cerastoderma nalivkini (Kolesnikov, 1929). Cardiidae.
Fossil. RECORDS. "1929 ZR : Cardium nalivkini
Kolesnikov, 1929 sp. n. Sarmatian. "1977 Popov:
Cerastoderma nalivkini. "1987 Taktakishvili: Cardium
nalivkini.
nalivkini Vassoevich, 1930 . - Didacna nalivkini
Vassoevich, 1930. Lyrnnocardiidae. Fossil. RECORDs.
"1930 ZR: Didacna nalivkini Vassoevich, 1930 sp. n.
fossil. "1989 Mikhailesku: Didacna nalivkini Wass.
nalivkini morosa Ko1esnikov, 1929 (variety) . Upgraded. Combination to be revised : "Cardium
nalivkini morosa Kolesnikov, 1929 . Cardiidae.
Sarmatian. REcoRDs. "1929 ZR: Cardium nalivkini var.
morosa Kolesnikov, 1929 nov. Sarmatian.
nanpei'lanense - Combination to be revised : "Cardium
nanpeizanense. Cardiidae . Pleistocene . RECORDS .
"1933 ZR: Cardium (Trachycardium) nanpeizanense
Pleistocene [Ref. Nomura 1933].
naphtalanicum Alizade. - Raricardium naphtalanicum
(Alizade). Cardiacea. Neogene. RECORDS. "1986
Nevesskaya et al.: Raricardium naphtalanicum
(A1izade) comb. nov. "1987 ZR : Raricardium
naphtalanicum (Alizade) comb. nov. Ref. to
Nevesskaya et at. p. 191.
napoieonense Winchell, 1862. - Combination to be
revised: "Cardium napoleonense Winchell , 1862.
Cardiidae. Fossil. RECORDs. "1980 Ruhoff: Cardium
napoleonense Winchell, 1862; p. 419.
nargiavagica Nevesskaja, 1986 . - Prosodacna
nargiavagica Nevesskaja, 1986. Lymnocardiidae.
Upper Miocene. TYPE LOCALITY. Azerbaijan. RECORDS.
"1986 Nevesskaya et al., p. 190, fig.: Prosodacna
(Zamphiridacna?) nargiavagica Nevesskaja, 1986 sp.
nov. "1987 ZR: Prosodacna (Zamphiridacna?)
nargiavagica Nevesskaja, 1986 sp. nov. Azerbaidjan,
Upper Miocene. Ref. to Nevesskaya et al. p. 190.
nargiavagicum Ebersin. - Changed combination:
Nargicardium
nargiavagicum
(Ebersin) .
Lymnocardiidae. Pliocene. RECORDS. "1958 ZR:
Limnocardium (Nargicardium) nargiavagicum Eberzin,
1958 sp. nov. Azerbaijan. Pliocene of the Ponto Caspian
basin, Russia. Subgenotype of (Nargicardium). "1987
Taktakishvili: Nargicardium nargiavagicum Ebers.
Nargicardium Eberzin, 1958 (genus). - Lyrnnocardiidae.
Pliocene. RECORDS. "1958 ZR: (Nargicardium)
Eberzin, 1958 subgen. nov. of Limnocardium Pliocene
of the Ponto Caspian basin, Russia. Subgenotype:
Limnocardium (Nargicardium) nargiavagicum Eberzin,
1958 . Azerbaijan . Pliocene . "1958 . ZR :
(Nargicardium) Eberzin, 1958 subgen. nov. of
Limnocardium q .v. "1980. Vokes: Nargicardium
Ebersin, 1947 in subfamily Lymnocardiinae.
Nargicardium (subgenus). RECORDS. "1987.
Taktakishvili: (Nargicardium) Ebersin, 1947.
naricum Vredenburg, 1928. - Discors naricum
645
naganoensis - natella
(Vredenburg, 1928). Cardiidae. Fossil. RECORDS. "1977
Popov: Discors naricum (Vredenburg, 1928).
narisawaense (error). - Printer 's mistake for narusawaense in Nomura, 1935 (Keen 1973).
narusawaense Nomura, 1935. - Changed combination:
Vasticardium narusawaense (Nomura, 1935) .
Cardiidae. Tertiary. RECORDS. "1935 Nomura: Cardium
narusawaense. ··1935 ZR: Cardium (Trachycardium)
narusawaense sp. n. Tertiary Kurikoma Japan. "1973
Keen: Vasticardium narusawaense (Nomura, 1935).
nasyrica - Lymnocardium nasyrica. Lyrnnocardiidae.
Fossil. RECORDS. "1987. Taktakishvili: Tauricardium
nasyrica.
nasyrica (subspecies and variety). - See under petersi
nasyrica and subsquamulosum nasyrica Eberzin, 1958.
natalense Krauss, 1848. [Figs . Part 3 p. 916]: - Changed
combination: Fulvia natalensis (Krauss, 1848). OTHER
JUDGEMENTS : Fulvia apertum (Bruguiere, 1789). Papyridea papyraceum (Chemnitz) Schroter [not
available]. » Regarding Cardium natalensis:
Cardiidae. Recent. DISTRIBUTION. South Africa Indo-Pacific (Hylleberg 1994). RECORDS. "1848 Krauss, Tab.
I. fig. 9: Cardium natalense Krauss. "1869 Romer:
Cardium natalense Krauss = Cardium papyraceum
Chemnitz. "1890 Paetel : Cardium natalense Krss. is
Papyridea. "1890 Paetel: Papyridea natalensis Krss.
= papyracea Ch. [not available]. "1904 Smith:
Cardium natalense Sowb. ··1915 Bartsch: Cardium
(Fulvia) natalensis Sowerby. ··1922-32 Sherborn, p.
4160: Cardium natalense Krauss, 1848. "1932 ZR:
Cardium natalensis [Ref. Turton 1932]. "1950
Barnard: Papyridea natalensis . ··1974 Barnard:
Cardium natalense Krauss 1848 = Papyridea
papyracea. "1977 Fischer-Piette: Cardium natalense
Krauss = Papyridea papyraceum (Chemnitz) Schroter.
"1977 Wilson & Stevenson : Cardium natalense
Krauss, 1848 = Fulvia apertum (Bruguiere, 1789).
'·1994 Vidal: Fulvia (Fulvia) natalensis (Krauss,
1848).
natalense Sowerby. - See under natalense Krauss, 1848.
natalense adjacens Turton, 1932 (variety). - Changed
combination: Fulvia natalensis (Krauss, 1848). OTHER
JUDGEMENTS: Fulvia aperta (Bruguiere, 1789). »
Regarding Cardium natalensis var. adjacens:
Cardiidae. Recent. RECORDS. "1932 ZR: Cardium
natalensis var. adjacens Turton, 1932 nov. Port Alfred
S. Africa [Ref. Turton 1932]. ··1977 Wilson &
Stevenson: Cardium natalense var. adjacens Turton,
1932 = Fulvia aperta (Bruguiere, 1789). ··1994 Vidal:
Cardium natalensis var. adjacens Turton, 1932 = Fulvia
(Fulvia) natalensis (Krauss, 1848).
Natella Ebersin, 1949 (genus). - Homonym. Replaced
by Pachydacna Ebersin. 1955. Lymnocardiidae.
Tertiary. RECORDS. ··1980 Vokes: Natella Ebersin, 1949
non Watson, 1934 (Mamm.), see Pachydacna Ebersin.
1955.
natella Ebersin, 1947. -Natella natella Ebersin, 1947.
Lymnocardiidae . Fossil. RECORDS. ··1987 Anaya :
Natella natella Ebersin, 1947 is type species of genus
Pachydacna Ebersin, 1955.
natella Ebersin, 1960. - Pachydacna natella (Ebersin,
1960). Lymnocardiidae. Tertiary. RECORDS. ··1960 ZR:
Pachydacna natella Eberzin, 1960 sp. nov. Duab river,
Abkhaziya, Georgia, USSR. ··1960 ZR: Pachydacna
646
Tropical Marin e Mollusc Programme (TMMP)
natella extenllis - negativllm
natella natella Eberzin, 1960 subsp. nov. Duab river,
Abkhaziya, Georgia, USSR.
flatella exteflllis Eberzin, 1960 (subspecies). - Pachydacna natella extenuis Eberzin, 1960. Lymnocardiidae,
Tertiary. RECORDS. --1960 ZR: Pachydacna natella
extenuis Eberzin, 1960 subsp. nov. Duab river,
Abkhaziya, Georgia, USSR.
natella pselldocilllmerica Eberzin, 1960 (subspecies) . Pachydacna natella pseudocimmerica Eberzin, 1960.
Lymnocardiidae. Tertiary. RECORDS. --1960 ZR:
Pachydacna natella pseudocimmerica Eberzin, 1960
subsp. nov. Duab river, Abkhaziya, Georgia, USSR.
flalltiloides Castelnau , 1843. - Combination to be
revised: "Cardium " nauti/oides Castelnau, 1843 .
Cardiidae. Si Iurian. TYPE SPECIES. Stoliczka (1871) stated
that Cardium nauti/oides is type species of genus
Goldfussia Castelnau, 1843. Cardium nauti/oides is
referred to Cardiidae by Philippi. Fossil. RECORDS.
"1871 Stoliczka: Cardium nauti/oides. "1922-32
Sherborn, p. 4263: Cardium nauti/oides Caste1nau,
1843.
*neblliosum . - Homonymy to be noticed. Cardium
nebulosum from 2 sources: Reeve, 1845. - Ireda1e,
1927.
fleblliosa (mode). - See under nebulosum.
flebulosuIII Iredale, 1927. - Vasticardium nebulosum
(Iredale, 1927). Cardiidae. Recent. TYPE SPECIES. Iredale
(1927) designated Cardium nebulosum, citing Martyn,
1786, type species of the genus Vasticardium.
Gladenkov & Sinelnikova (1990) stated that Cochlea
nebulosa (citing Martyn) is type species of genus
Vasticardium Iredale, 1927. TYPE LOCALITY. Unknown.
RECORDS.
--1927 Iredale : Trachycardium
(Vasticardium) nebulosum . --1969 Moore et al. :
Acrosterigma (Vasticardium) nebulosum (Iredale,
1927). - CHARACTERS AND OCCURRENCE REMARKS. --1936 Keen: Martyn' s Universal
Conchologist is not consistently binomial and hence is
to be rejected. HABITAT. --1971 Kuroda, Habe & Oyama:
18-110 m depth. DISTRIBUTION. "1971 Kuroda, Habe
& Oyama: Indo-Pacific; in Japan from Honshu to the
tropical Pacific.
nebulosum Martyn, 1784. Non binominal. - Not
available . Replaced by Vasticardium nebulosum
Iredale, 1927. OTHER JUDGEMENTS: Acrosterigma
elongatum (Bruguiere, 1789). » Regarding Cardium
nebulosum: Cardiidae. Recent. TYPE SPECIES . Cotton
(1961) stated that Cardium nebulosum, citing Martyn
1786, is type species of Vasticardium Iredale 1927.
Martyn rejected for nomenclatoria1 purposes. REMARKS.
The specimen figured by Martyn has 28 ribs.
DISTRJBUTION. China (Martyn, 1784). RECORDS. "1784
Martyn, p. 50, pI. 46, fig. I : Cardium nebulosa. --1880
Martens: Cardium nebulosum Martyn = Cardium
(Trachycardium) elongatum Bruguiere. --1961 Cotton:
Cardium nebulosum Martyn 1786. Type species of
Vasticardium Iredale 1927. "1971 Kuroda, Habe &
Oyama : Co chlea n ebulosa citing Martyn =
Vasticardium nebulosum Iredale, 1927. --1977 Popov:
Cochlea nebulosa Martyn , 1792. Type of
(Vasti cardium) Iredale , 1927 . .. 1977 Wilson &
Stevenson: Cochlea nebulosa citing Martyn, 1792 (=
Cardium elongatum Bruguiere, 1789). --1977 Wilson
& Stevenson: Vasticardium nebulosum citing Martyn,
1799. --1978 Maxwell: Vasticardium nebulosum citing
Martyn, 1784 in Iredale 1927 = Cardium elongatum
Bruguiere 1792. --1980 Keen: Cochlea nebulosa citing
Martyn = Cardium elongatum Bruguiere, 1789. --1990
G1adenkov & Sinelnikova: Cochlea nebulosa citing
Martyn . --1993 Vida\: ?Cochlea nebulosa citing
Martyn, 1786 = Acrosterigma elongatum (Bruguiere,
1789).
neblllOSlllll Reeve, 1845. [Figs. Part 1 p. 142. Part 3 p.
877, 927] . - Synonym of Acroslerigma simplex
(Spengler, 1799). OTHER JUDGEMENTS : Tra chycardium
unicolor (Sowerby, 1841). - Cardium enode Sowerby.
» Regarding Cardium nebulosum : Cardiidae. Recent.
TYPE LOCALITY. Unknown (Reeve 1845); the Maldives
(Vidal 1999). TYPE REPOSITORY. Holotype, BMNH reg.
no. 1900.2.13.22. Neg. No. 0386. A paired specimen
from the Maldives figured by Reeve (Vidal 1999).
RECORDS. "1845 Reeve: Cardium nebulosum . "1858
Adams & Adams, p. 455: Cardium (Trachy cardium)
nebulosum , Reeve. --1890 Paete1 : Cardium nebulosum
Rv. --1922-32 Sherborn, p. 4270: Cardium nebulosum
Reeve, 1845 . "1932 Prashad : Cardium nebulosum
Reeve, 1845 = Cardium unicolor Sowerby, 1841 .
"1977 Fischer-Piette: Cardium nebulosum Reeve =
Cardium enode Sowerby. --1988 Drivas & Jay:
Trachycardium nebulosum (Reeve, 1845) . "1993
Vidal: Cardium nebulosum not synonym of Cardium
enode as suggested by Fischer-Piette. --1999 Higo el
al.: Cardium nebulosum Reeve, 1845 = Acrosterigma
(Vasticardium) unicolor (Sowerby, 1841). "1999 Vidal :
Cardium nebulosum Reeve, 1845: Sp. 99 =
Acroslerigma simplex (Spengler, 1799). --1999 Vidal :
Drivas & Jay 1988, p. 17, fig. 15: Trachycardium
nebulosum (Reeve) = Acrosterigma mauritianum
(Deshayes, 1855) [Not Cardium nebulosum Reeve =
Acrosterigma Simplex (Spengler)]. --1999 Vidal: Drivas
& Jay 1988, p. 140, pI. 55: Trachycardium nebulosum
(Reeve) = Acroslerigma mauritianum (Desha yes, 1855)
[Not Cardium nebulosum Reeve = Acrosterigma
Simplex (Spengler)]. - CHARACTERS AND OCCURRENCE - DESCRIPTIONS. "1845 Reeve: shell ovate,
somewhat Pecten-shaped, radiately ribbed, 42-43
convex, smooth ribs, posteriorly very finely crenulated;
whitish, clouded with pale brown spots, ribs irregularly
painted here and there with red transverse lines; interior
white, stained with pink beneath the umbones,
ornamented near the margins with blotches of bright
purple violet. DIMENS IONS. --1988: Drivas & Jay: max .
ant.- post. 26.5 mm, umbo-ventr. 29.5 mm. REMARKS.
--1845 Reeve: the external painting ofthis shell is very
similar to that of the Pectunculus pectinatus. "1988 :
Drivas & Jay : Shape similar to C. mauritianum
(Deshayes) but has a lower number of ribs, and a more
rounded outline. HABITAT. ,,1988: Drivas & Jay:
Reunion, western Indian Ocean. DISTRIBUTION. "1845
Reeve: unknown ...Of authors: Central Indo-West
Pacific, Australia.
ne/anduIII Kolesnikov, 1935. - Combination to be
revised : "Cardium " nefandum Kolesnikov, 1935 .
Cardiidae. Sarmatian. RECORDS. --1945 ZR: Cardium
nefandum Kolesnikov, 1935 sp. nov. USSR Sarmatian.
ne/arius (variety). - See nikitini nef arius Uspenskaia,
1931.
negativum Andrussow, 1909. - Combination to be
Phuket Marine Biological Center Special Publication vol. 30 (2004)
647
neglectum - neozelandicum
revised : "Cardium" negativum Andrussow, 1909.
Cardiidae. Miocene. RECORDS. ··1910 ZR: Cardium
negativum Andrussow, 1909 sp. n. Fossil. ··1963 ZR :
Cardium negativum Andrusov, 1963 sp. nov. Miocene,
Russia.
neglectum Holzl, 1962. - Combination to be revised:
"Cardium" neglectum Holzl, 1962. Cardiidae.
Oligocene. RECORDs. ··1962 ZR: Cardium neglectum
Holzl, 1962 sp. nov. Sulz-Graben (Leitzachtal)
Chattian, Oligocene, Bavaria. ··1962 ZR: Cardium
neglectum Oligocene.
neglectum angustum Holzl, 1962 (variety). Combination to be revised: "Cardium" neglectum var.
angustum Holzl, 1962. Cardiidae. Oligocene. RECORDs.
··1962 ZR: Cardium neglectum var. angus tum Holzl,
1962 nov. Bad Tolz Chattian, Bavaria.
neglectum intersulcatum Baldi, 1975 (subspecies). New
combination: Trachycardium neglectum intersulcatum
(Baldi, 1975). Cardiidae. Upper Oligocene. RECORDS.
··1975 ZR : Cardium (?Trachycardium) negtectum
intersulcatum Baldi, 1975 subsp . nov. Hungary,
Borehole Solymar. Ref. to Baldi, p. 208.
nemo Fischer-Piette, 1977. [Figs . Plate 3 p. 930].
Dubious species. » Regarding Laevicardium nemo:
Cardiidae. Recent. TYPE REPOSITORY. Holotype, right
valve, Paris Museum. RECORDS. ··1977 Fischer-Piette:
Laevicardium nemo n. sp. ··1977 ZR: Laevicardium
nemo Fisher-Piette, 1977 sp. nov. Type locality not
given. Ref to Fischer-Piette, E. p. 24. ··1980 Keen:
species insufficiently known. ··1991 Voskuil &
Onverwagt: Laevicardium nemo Fischer-Piette, 1977.
- CHARACTERS AND OCCURRENCE - DESCRIPTIONS. ··1977 Fischer- Piette: Shell white, equilateral,
oblong, inside with marginal dentation, median tooth
small, lateral teeth well separated, different size. 55
ribs. [I have translated from French]. DIMENSIONS.
Umbo-vent. 31 mm, anterior-post. 22 mm, half
diameter 9 mrn. REMARKS. ··1980 Keen: a fragmentary
specimen resembling a fossil Clinocardium from the
North Pacific . ··1991 Voskuil & Onverwagt:
Laevicardium nemo Fischer-Piette, 1977 proposed as
a nomen novum for Cardium fragile Sowerby, but
Cardium fragile Sowerby, 1841 is considered a nomen
dubium . DISTRIBUTION . Unknown.
nemocardiforme Kafanov & Savizky, 1983 . Laevicardium nemocardiforme Kafanov & Savizky,
1983 . Cardiidae. Middle Miocene. RECORDS. ··1983
Kafanov & Savizky, p . 119: Laevicardium
nemocardiforme Kafanov et Savizky. ··1983 ZR:
Laevicardium nemocardiforme Kafanov et at., 1983 sp.
nov. Russian SFSR, Middle Miocene. Ref. to Kafanov
etal., p. 119.
nemocardiiformis Eames, 1951. - Discors nemocardiiformis Eames, 1951 . Cardiidae. Eocene. RECORDS.
··1951 ZR: Discors nemocardiiformis Eames, 1951 sp.
nov. Eocene Pakistan.
Nemocardium Meek, 1876 (genus). - Cardiidae. Fossil
and Recent. Type species: Sacco (1899) designated
Cardium semiasperum Deshayes, 1858 [Eocene,
France] type species of genus Nemocardium . RECORDS.
··1876 Meek: Nemocardium, Meek . Cardium
semiasperum Deshayes regarded as genotype. ··1921
ZR: Cardium (Nemocardium) [Ref. Brun & Vedel
1921]. ··1923 ZR: Nemocardium [Ref. Cossmann
1923]. ··1923 ZR: Nemocardium [Ref. Lissajous 1923].
··1946 ZR: Nemocardium shell structure [Ref. Wrigley
1946] . ··1947 ZR: Nemocardium Meek 1876, Rep. U.S.
Geol. Surv. Territories (Haydm. sic) 9 167, 173 type
by virtual monotypy and subseq. desig. Sacco 1899.
Cardium semiasperum Deshayes, Eocene, France [Ref.
Vokes 1946]. ··1950 ZR: Nemocardium history of the
genus and list of species with localities [Ref. Keen
1950]. ··1961 ZR: Nemocardium Meek: Tertiary
species [Ref. Hughes 1961]. ··1954 Keen:
Nemocardium Meek, 1876. ··1975 ZR: Nemocardium.
Key to species. Ref. to Merklin, p. 104. ··1977 Popov:
Nemocardium Meek, 1876. Genotype: Cardium
semiasperum Deshayes , 1858. ··1977 ZR:
Nemocardium . Key to subgenera. Ref to Popov p. 63 . /
California, Pliocene. Ref to Hertlein & Grant p. 262.
··1978 Maxwell: Genus Nemocardium Meek, 1876.
··1980 Vokes: Nemocardium Meek, 1876 in subfamily
Protocardiinae . ··1986 Abdel-Gawad : genus
Nemocardium . ··1986 Scott: Nemocardium Meek,
1876. ··1992 Schneider p. 155: Nemocardium. Type
species Nemocardium semiasperum (Deshayes). ··1995
Pacaud & Renard : Nemocardium Meek, 1876 in
subfamily Protocardiinae. ··1998 Lamprell & Healy:
Nemocardium Meek, 1876. ··2000 Co an et al.: Genus
Nemocardium Meek, 1876. ··2002 Schneider:
Nemocardium (Nemocardium) originated in the Aptian,
is the oldest living cardiid genus or subgenus. ··1999
Higo et al.: Nemocardium Meek, 1876. Type species
Cardium semiasperum Deshayes, 1858 (s. d. Sacco,
1899). ··200 I Vidal, p. 407, Group 2: Nemocardium. CHARACTERS AND OCCURRENCE - DESCRIPTIONS. ··1994 Hylleberg: sculpture radial throughout,
strong and ornamented on posterior slope. Central and
anterior slopes nearly smooth, marginal crenulations
changing abruptly in size at posterior- ventral junction.
··200 I Vidal: tentacles numerous. No eyes on tentacles.
··2002 Schneider: Nemocardium has labial palps with
length less than one-third the length ofthe entire animal
(minus the shell).
Nemocardium Meek, 1876 (section and subgenus). Cardiidae . Fossil and Recent. R EMARKS. The
designation "section" takes the value of subgenus.
RECORDS. ··1887 Fischer: Nemocardium, Meek. 1876.
Section of subgenus Laevicardium Swainson, 1840.
··1954 Keen: (Nemocardium) s.s . ··1986 AbdelGawad: subgenus Nemocardium Meek, 1876. ··1986
Scott: (Nemocardium) Meek, 1876. ··1999 Higo et al.:
Nemocardium (subgenus) s. str.
neoinflatum Miletic-Spajic, 1960. - New name. New
combination: Lymnocardium neoinjlatum MileticSpajic, 1960. Lymnocardiidae. Tertiary. RECORDS.
··1960 ZR: Limnocardium neoinjlatum Miletic-Spajic,
1960 nom. nov. pro. Limnocardium inflatum lekelius,
1944 (non Gorjanovic-Kramberger 1902)
Aleksandrovae, Tertiary.
neotropica Cowper-Reid, 1943. - Pa chycardium
neotropica Cowper-Reid, 1943 . Cardiidae. Trias .
RECORDS. ··1947 ZR: Pachycardium neotropica
Cowper-Reid, 1943 sp. nov. Trias, Brazil.
neozelandicum Deshayes, 1853 . - New name:
"Cardium" neozelandicum Deshayes . Cardiidae.
Recent. RECORDS . ··1857 Deshayes : Cardium
neozelandicum new name for Cardium striatulum
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Tropical Marin e Mollusc Programme (TMMP)
neozelanica - nigropunctatum
Sowerby, 1840. 001980 Ruhoff: Cardium
neozelandicum Deshayes, 1853; vol. 2: 46.
neozelanica Marshall & Murdoch. - Lahillia neozelanica
Marshall & Murdoch. Cardiacea. Fossil. RECORDS.
001937 ZR: Lahillia neozefanica Marshall & Murdoch,
Danian Wangaloa, New Zealand [Ref. Finlay &
Marwick 1937].
neptuni Goldfuss, 1837. Not a cardiid. REcORDs. 001857
Deshayes: Cardium nepluni Goldfuss. Not a cardiid:
in genus Pinna. 001922-32 Sherborn, p. 4293 : Cardium
neptuni Goldfuss, 1837.
nequam Healey, 1908. - Combination to be revised :
" Cardi u m " nequam Healey, 1908 . Cardiidae .
Mesozoic. RECORDs. 001976 Ma Qi-hong el al. p. 255 :
Cardium nequam Healey, 1908. 001977 ZR :
Vietnamieardium nequam (Healey) Comb. nov . .
Original combination not given. Ref. to Vu-Huc D. p.
679. 00 1995 Schneider: Cardium nequam Healey, 1908
= Tulongoeardium nequam (Healey, 1908). 001995
Schneider: Tulongocardium nequam (Healey, 1908).
nequam (aff.) Healey, 1908. - Combination to be revised:
"Cardium " aff. nequam Healey, 1908. RECORDs. 001976
Ma Qi-hong el al.: Cardium aff. nequam Healey.
neszmelyense Strausz, 1975 . - New combination :
Laevicardium neszmelyense (Strausz, 1975). Cardiidae.
Middle Eocene . RECORDS . 001975 ZR: Cardium
(Laevieardium) neszmelyense Strausz, 1975 sp. nov.
Hungary, Neszmely. Ref. to Strausz, p. 20.
neumayri Fuchs, 1873 . - Prosodaena neumayri (Fuchs,
1873). Lymnocardiidae. Fossil. TYPE LOCALITY. From
Matitza, north of Ploesti . RECORDS. 001873 Fuchs, p.
22, pI. III, fig. 7-10 : Cardium neumayri. 001886 Fischer:
Prosodaenaneumayri (Fuchs, 1873).
nevesskajae Ali-Zade & Ribina. - New combination:
Replidacna nevesskaja e (Ali-Zade & Ribina) .
Cardiacea . Fossil. RECORDS. 001977 ZR: Cardium
(Rep/idacna) nevesskajae Ali-Zade & Ribina. Ref to
Paramonova p.70 [Trans!. p. 320).
nevesskajae Para monova, 1977. - Cerasloderma
nevesskajae Paramonova, 1977 . Cardiidae. Upper
Miocene. RE COR DS. 001977 ZR : Cerasloderma
(Planaeardium) nevesskajae Paramonova, 1977 sp.
nov. Ukraine, Upper Miocene. Ref to Il'ina ,
Nevesskaya & Paramonova p. 208, fig'd.
newberryana Gabb. - Pr%cardia newberryana Gabb.
Cardiidae. Fossil. RECORDs. 001922 Olsson: Protoeardia
newberryana Gabb.
newtoni Eames, 1957. - Loxocardium newloni (Eames,
1957). Cardiidae. Eocene. TYPE SPECIES. Eames, 1957
designated Fragum (A canlhoeardia) newloni type of
the subgenus Afrieofragum, which is synonym with
Schedocardia Stewart, 1930 (Moore el al. 1969) .
RECORDs. 001957 ZR: Fragum (Afrieofragum) newtoni
Eames, 1957 sp. nov. Eocene, Nigeria, subgenotype of
(Africofragum) Eames, 1957 subgen. nov. 001969
Moore et al.: Acanlhoeardia (Africofragum) newtoni
(Eames, 1957). 001970 Glibert & Poel: Loxoeardium
newtoni (Eames, 1957).
nexile Branson, 1942. - New name: Conocardium nexile
Branson, 1942. Conocardiidae. Fossil. RECORDs. 001942
ZR: Conocardium nexile Branson, 1942 nom. nov. p.
389, for Conoeardium phillipsi Koninck 1885 non
Orbigny 1850.
nexilis Martyn, 1784. Non binominal. - Not available.
» Regarding Cardium nexilis: not a cardiid. RECORDS.
001784 Martyn: Cardium nexilis.
nexingense (subspecies) . - See under latisufeus
nexingense Papp, 1954.
nexum Humphrey, 1797. - Synonym of Fragumfragum
(Linne , 1758) . » Regarding Cardium nexum :
Cardiidae . Recent. RECORDS. 001797 Humphrey :
Cardium nexum. 001932 Prashad : Cardium nexum
Humphrey, 1797 = Fragum fragum (Linne, 1758).
001977 Fischer-Piette: Cardium nexum Humphrey =
Coreulum fragum Linne. 001977 Wilson & Stevenson:
Cardium nexum Humphrey, 1797 (non binomial) =
Fragum fragum (Linnaeus, 1758). 001999 Higo el al.:
Cardium nexum Humphrey, 1797 = Fragum fragum
(Linnaeus, 1758).
nwgarense Winchell & Marcy, 1865. - Conoeardium
niagarense Winchell & Marcy, 1865. Conocardiidae.
Fossil. REMARKS. Conocardiids have no connection with
the Cardiacea except analogy of form, which does not
offer any point of relationship (Dall 1895, p. 519).
RECORDS. 001980 Ruhoff: Conocardium niagarense
Winchell & Marcy, 1865; vo!. 1: 97, pI. 2, fig. 14.
nicense Bellardi, 1852. - Combination to be revised:
"Cardium" nieense Bellardi, 1852. Cardiidae. Fossil.
RECORDS. 001980 Ruhoff: Cardium nicense Bellardi,
1852; vol. 4: 241, pI. F (sic = 17) fig. 9.
niciense d'Orbigny, 1850. - Combination to be revised:
"Cardium" niciense d'Orbigny, 1850. Cardiidae.
Fossil. RECORDS. 001922-32 Sherborn, p. 4305: Cardium
niciense d'Orbigny, 1850.
nicklesi Adegoke, 1977. - Cardium nicklesi. Cardiidae.
Paleocene. RECORDS. 001977 ZR: Cardium (Cardium)
nicklesi sp. nov. Nigeria. Palaeocene. Ref. to Adegoke
p. 270.
nicol/elli Conrad, 1842 . - Changed combination:
Nemoeardium nicollelli (Conrad, 1842). Cardiidae.
Eocene. RECORDS . --1886 Dall : Cardium nicollelli
Conrad [Eocene]. 001922-3 2 Sherborn, p . 4307:
Cardium nicollelli Conrad, 1841. 001961 ZR:
Nemocardium nicollelli (Conrad) systematic
description and review ofN. American Paleocene forms
[Ref. Hughes 1961]. 001977 Popov: Nemocardium
(Nemocardium) nieol/elli (Conrad, 1842). 002001
Schneider & Carter: Nemocardium nicollelli (Conrad,
1841). Cardiidae. Early Cenozoic, Eocene, Mississippi,
USA.
nigamiensis (subspecies). - See under makiyami
nigamiensis Noda, 1962.
niger Zhizhchenko, 1934. - Combination to be revised:
"Cardium " niger Zhizhchenko, 1934 . Cardiidae.
Miocene . RECORDS . 001939 ZR: Cardium niger
Zhizhchenko, 1934 sp. n. Miocene Ciscaucasia. 001952
ZR : Cardium niger Miocene Caucasus [Ref.
Zhezhcenko 1934].
nigropunctatum Habe & Kosuge, 1966. - Synonym of
Vasticardium peeliniforme (Born, 1780). OTHER
JUDGEMENTS: Aeroslerigma rugosum (Lamarck, 1819).
- Aeroslerigmaflava (Linnaeus, 1758). » Regarding
Vaslicardium nigropunetatum: TYPE LOCALITY. Amami
Oshima (Higo et al. 1999). Cardiidae. Recent. RECORDS.
001966 Habe & Kosuge , p. 154 (pI. 59, fig . 9):
Vaslieardium nigropunetalum . 001966 Ga rrard:
Va s tieardium nigropunclalum . 001966 ZR :
Vaslieardium nigropunelatum Habe & Kosuge, 1966
Phuket Marine Biological Center Special Publication vol. 30 (2004)
649
nikitini - nivale
sp. nov. Amami Is. Japan. "1967 ZR: Vaslicardium
nigropunctatum, North Keppel Island, Queensland,
Australia [Ref. Garrard 1967]. "1974 Habe & Kosuge:
Vasticardium nigropunctatum Habe et Kosuge. "1977
Wilson & Stevenson: Vasticardium nigropunctatum
Habe & Kosuge, 1966 = Acrosterigma rugosum
Lamarck, 1819. "1992 Lamprell & Whitehead:
Vasticardium nigropunctatum Habe & Kosuge, 1966
= Acrosterigma flava (Linne, 1758). "1977 FischerPiette, p. 174: Vasticardium nigropunctatum Habe &
Kosuge = Laevicardium flavum Linne auctorum. "1977
Habe: Trachycardium (Vasticardium) nigropunctallIm
Habe & Kosuge, 1966. "1997 Vidal : Vasticardium
nigropunctatum Habe & Kosuge, 1966 = Vasticardium
pectiniforme (Born, 1780). "1998 Poutiers :
Vasticardium nigropunctatum Habe & Kosuge, 1966.
"1999 Vidal: Vasticardium nigropunctatum Habe &
Kosuge 1966 = Vasticardium pectiniforme (Born,
1780). "1999 Higo et al.: Acrosterigma (Regozara)
nigropunctatum (Habe & Kosuge, 1966). "1999 Higo
et al.: Vasticardium nigropunctallIm Habe & Kosuge,
1966. - CHARACTERS AND OCCURRENCE REMARKS. Trachycardium nigropunctatum Habe &
Kosuge, 1966 is not synonym with Trachycardium
flavum (Linne, 1758). HABITAT. "1999 Higo et al.:
subtidal to 20 m, sand. DISTRlBUTION. "1994 HylJeberg:
Indo-Pacific, Red Sea, Indo-West Pacific, Australia.
"1999 Higo et al.: Amami islands and southwards, East
China Sea, Philippines, northern Australia.
*nikitini - Homonymy to be noticed. Cardium nikitini
from 2 sources: Andrussow, 1902. - Andrusov, 1963.
- REMARKS. I do not know how Andrussow 1902
compares with Andrusov, 1963.
nikitini Andrusov, 1963. - Combination to be revised:
"Cardium" nikitini Andrussow, 1902. Cardiidae.
Neogene. RECORDS. "1961 ZR: Cardium nikitini
Andrusov, 1961 "sp. nov."[in a table] Miocene Caspian
region, Russia. "1963 ZR: Cardium nikitini Andrusov,
1963 sp. nov. Neogene, Russia [Ref. Andrusov 1963].
nikitini Andrussow, 1902. - Combination to be revised:
"Cardium" nikitini Andrussow, 1902. Cardiidae.
Miocene. RECORDS. "1905 Dollfus, commenting
Andrussow 1902: Cardium nikitini n. sp. tends towards
Cerastoderma. "1932 ZR: Cardium nikitini Andrus.
Miocene [Ref. Uspenskaia 1931].
nikitini Andrussow. -Avicardium nikitini (Andrussow).
Cardiacea. Fossil. RECORDS. "1977 Popov: Avicardium
nikitini (Andrussow).
nikitini nefarius Uspenskaia, 1931 (variety). Upgraded. "Cardium" nikitini nefarius Uspenskaia,
1931. Cardiidae . Miocene . RECORDS. "1932 ZR:
Cardium nikitini Andrus var. nefarius U spenskaia, 1931
nov. Miocene Daghestan and Transcaspian.
nikolaevae Merklin, 1975. - Parvicardium nikolaevae
Merklin, 1975. Cardiidae. Upper Oligocene. RECORDS.
"1975 ZR: Parvicardium nikolaevae Merklin, 1975
sp. nov. Kazakhstan, south west Mangyshlak. Ref. to
Merklin, p. 99.
nimbata Martyn, 1784. Non binominal. - Not available.
» Regarding Cardium nimbata: not a cardiid.
RECORDS. "1784 Martyn: Cardium nimbata.
nipponenseTashiro, 1976. - Granocardium nipponense
Tashiro, 1976. Cardiidae. Upper Cretaceous. RECORDS.
"1976 ZR: Granocardium nipponense sp. nov. Japan,
Kyushu, Miyanokawachi. Ref. to Tashiro, p. 65.
nipponica Yokoyama, 1925. - Changed combination:
Profulvia nipponica (Yokoyama, 1925). OTHER
JUDGEMENTS: Fulvia hariimani (Dall, 1904) . »
Regarding Papyridea nipponica: Cardiidae.
Oligocene-Miocene. TYPE LOCALITY. Honshu, Japan.
TYPE REPOSITORY. Type CM 22090 missing at the
University Museum, University of Tokyo (Kafanov et
al. 1994). RECORDS. "1924 Yokoyama, p. 17, p1.3, figs.
3, 4: Papyridea (Fulvia) nipponica (fide Kafanov et
al. 1994). "1924 ZR: Papyridea (Fulvia) nipponica
Yokoyama, 1924 sp . n. Tertiary Japan. "1925
Yokoyama: Papyridea (Fulvia) nipponica Yokoyama.
"1973 Keen: Papyridea nipponica Yokoyama, 1924
(fide Oyama, Mizuno, & Sakamoto, 1960) = Fulvia
hariimani (DalJ, 1904). "1976 ZR: Profolvia nipponica
(Yokoyama, 1924) Comb. nov. Transferred from
Papyridea. Ref. to Kafanov, p. 482 [orig. p. III]. "1980
ZR: Profulvia nipponica (Yokoyama 1924) is ajunior
synonym of P hariimani (Dall 1904). ZR Ref. to
Kafanov.
nipponicum Noda, 1991. - Vasticardium nipponicum
Noda, 1991. Cardiidae. Pleistocene. TYPE LOCALITY.
Ryukyu Islands, Japan. RECORDS. "1991 Noda, p. 27,
fig.: Vasticardium nipponicum sp. nov.
*nitens. - Homonymy to be noticed. Cardium nitens
from 2 sources: Sowerby, 1813. - Galeotti (?year).
nitens Galeotti. - Combination to be revised: "Cardium"
nitens Galeotti. Cardiidae. Fossil. TYPE LOCALITY.
Belgium. RECORDS. "1857 Deshayes: Cardium nitens
Galeotti.
nitens Sowerby, 1813. - Changed combination:
Nemocardium nitens (Sowerby, 1813). Cardiidae.
Fossil. RECORDS. "1922-32 Sherborn, p. 4369: Cardium
nitens Sowerby, 1813. "1970 Glibert & Poel:
Nemocardium (s.s.) nitens (Sowerby, 1813), compared
with N. turgidum (Solander). "1977 Popov :
Nemocardium (Nemocardium) nitens (Sowerby, 1813).
nitida Andrussov, 1924. - Monodacna nitida Andrussov,
1924. Lyrnnocardiidae. Fossil. RECORDS. "1924 ZR:
Monodacna nitida Andrussov, 1924 sp. n. Pleistocene.
"1963 ZR: Monodacna nitida Andrusov, 1963 sp. nov.
Pliocene, Russia.
nitidulum Tremlett, 1950. - Nemocardium nitidulum
Tremlett, 1950. Cardiidae. Fossil. RECORDS. "1950 ZR:
Nemocardium nitidulum Tremlett, 1950 sp. nov. ex
Edwards MS . London Basin. "1970 Glibert & Poel:
nitidulum Tremlett, 1950. "1977 Popov: Nemocardium
(Nemocardium) nitidulum Tremlett, 1950.
nitidum - Unknown. « Regarding Cardium
papyraceum : RECORDS. "1817 Dillwyn: under the name
Cardium nitidum, Mr. Humphrey sold Cardium
papyraceum Chernnitz.
nivale Reeve, 1845. [Figs. Part I p. 142. Part 3 p. 909,
931]. - Changed combination: Fragum nivale (Reeve,
1845). Cardiidae. Recent. RECORDS. "1845 Reeve:
Cardium nivale. "1858 Adams & Adams, p. 458:
Hemicardia (Fragum) nivalis Reeve. "1870
Macandrew: Hemicardia (Fragum) nivale. "1884
Grasset: Hemicardium nivale Rv. "1890 Paetel:
Hemicardium nivale Rv. ··1899 Sturany: Hemicardia
nivale. "1903 Hidalgo: Hemicardium nivale Reeve.
1845. "1905 Sturany: Hemicardium nivale (Rve.).
"1909 Melvill: Cardium (Ctenocardia) nivale, Reeve.
650
Tropical Marine Mollusc Programme (TMMP)
nivalis - nodosum
"1922-32 Sherborn, p. 4387 : Cardium nivale Reeve,
1845. "1926 Fischer & Fischer: Cardium
(Hemicardium) nivale Rve. "1928 Faustino: Cardium
(Hemicardium) nivale Reeve. "1931 Thiele & Jaeckel:
Cardium (Fragum) nivale Reeve. ··1956 ZR :
Hemicardia nivale Reeve new to Seychelles [Ref. Rost
& Soot-Ryen 1956]. "1965 Biggs: Hemicardia
(Hemicardia) nivale. "1977 Fischer-Piette: Corculum
(Hemicardia) nivale Reeve. "1992 Oliver: Fragum
nivale . "2001 Vidal, p. 408 : Fragum nivale. CHARACTERS AND
OCCURRENCE
DESCRIPTIONS. "1845 Reeve: shell triangularly cordate,
a little compressed towards the margins, angulated and
flatly depressed on the posterior side; radiately flatly
ribbed, 35 ribs, those of the posterior area smooth,
irregular, sometimes narrow, sometimes rather broad,
the anterior very finely nodulously crenated; transparent
white, profusely ornamented, on the posterior area
especially, with opaque snowy spots . .. 200 I Vidal: only
the exhalant aperture is tube-shaped. Inhalant siphon
open towards the foot aperture. No eyes. REMARKS.
"1845 Reeve: An interesting new species, intermediate
in form between Cardium fragum and Cardium
hemicardium. HABITAT. .. 1845 Reeve: dredged from
coral sand at the depth of 8 fathoms . DISTRIBUTION.
.. 1845 Reeve: Island ofCorrigidor, Philippines. "1884
Grasset: China ... 1909 Melvill and "1994 Hylleberg:
Red Sea, Kenya.
nivalis (mode). - See under nivale Reeve, 1845 .
nixicollis Stephenson, 1923 . - New name : Protocardia
nixicollis (Stephenson, 1923) nom. nov. Cardiidae.
Upper Cretaceous. RECORDS . .. 1936 ZR : Cardium
nixicollis Stephenson, 1923 nom . nov. for Cardium
(Protocardia) vaughani Stephenson, 1923 Invert. pos.
Upper Cretaceous N. Carol. Geol. & Econ. Surv. 5 1923
p. 287 non Shattuck.
njalindungense Martin, 1922. - Combination to be
revised: "Cardium" njalindungense Martin, 1922.
Cardiidae. Tertiary. RECORDS . "1922 ZR : Cardium
(Eucardium) njalindungense Martin, 1922 sp. n.
Tertiary.
noae Roding, 1798. - Synonym of Tridacna maxima
(Roding, 1798). Tridacnidae. Recent. RECORDS ... 1947
ZR: Tridacna noae (Roding) [Ref. McLean 1947).
.. 1999 Higo et al.: Tridacnes noae Roding, 1798 =
Tridacna (Flodacna) maxima (Roding, 1798).
nobile Sabba Stefanescu, 1896. - Changed combination:
Euxinicardium nobile (Sabba Stefanescu , 1896).
Lymnocardiidae. Tertiary. RECORDS. "1987 Anaya:
Limnocardium nobile Sabba, Stefanescu 1896. "1987
Taktakishvili: Limnocardium nobile Sabba. "1987
Taktakishvili: Limnocardium (Euxinicardium) nobile
Sabba. "1987 Taktakishvili: Euxinicardium nobile
(Sabba).
nobile (cf.) Sabba Stefanescu, 1896. - RECORDS ... 1987
Anaya: Limnocardium (Euxinicardium) cf. nobile
Sabba Stefanescu.
nobile amicorum Ebersin, 1947 (variety). - Upgraded.
Changed combination: Euxinicardium nobile amicorum
(Eberzine, 1947). Lymnocardiidae. Tertiary. TYPE
LO CALITY. Russia. RECORDS. "1987 Anaya:
Limnocardium nobile var. amicorum Eberzine, 1947 .
.. 1987 Taktakishvili: Limnocardium (Euxinicardium)
nobile Sabba vaT. amicorum Andrus. "1987
Taktakishvili: Euxinicardium nobile amicorum Ebers.
"1987 Taktakishvili: Limnocardium (Euxinicardium)
nobile amicorum Ebers.?
nobile amicorum (aff.) Ebersin, 1947. - RECORDS. "1987
Taktakishvili: Euxinicardium aff. nobile amicorum
Andrus.
nodai Kafanov, 1980. - Serripes nodai Kafanov, 1980.
Cardiidae. Fossil. RECORDS. "1980 Kafanov, p. 319:
Serripes nodai Kafanov. "1980 ZR: Serripes nodai
Kafanov, 1980 nom. nov. for Cardium pauperculum
Yokoyama 1923 nee Meek 1871. Ref. to Kafanov, p.
319.
nodosulum Wood, 1853 . - Parvicardium nodosulum
(Wood, 1853). Cardiidae. Fossil. RECORDS. "1970
Glibert & Poel, p. 46. Parvicardium nodosulum (Wood,
1853).
*nodosum. - Homonymy to be noticed. Cardium
nodosum from 4 sources: Montagu, 1803. - Turton,
1822. - Chiereghini in Nardo, 1847. - Mourlon, 1880.
nodosum Chiereghini in Nardo, 1847. - Combination
to be revised: "Cardium" nodosum Chiereghini in
Nardo, 1847. Cardiidae. Fossil. RECORDS. "1922-32
Sherborn, p. 4401 : Cardium nodosum Nardo (ex
Chiereghini MS), 1847.
nodosum Montagu, 1803 . [Figs. Part 1 p. 145, 171]. Identical with Parvicardium scabrum (Philippi, 1844) .
OTHER JUDGEMENTS: Acanthocardia tuberculata
(Linnaeus, 1758). » Regarding Cardium nodosum :
TYPE MATERIAL missing (Glibert & Poel, p. 43).
Cardiidae. Recent. RECORDS. "1803 Montagu, No.6:
Cardium nodosum n. sp. "1807 Maton & Rackett, No.
11: Cardium nodosum Montagu p. 81. "1813 Fleming:
Cardium nodosum Montagu ... 1815 Wood: Cardium
nodosum Mont. .. 1817 Dillwyn: C. nodosum Montagu
has too uncertain status to be retained. "1840 Sowerby,
catalogue p. 2, No. 23: Cardium nodosum Montagu =
Cardium nodosum Turton, 1822. "1845 Reeve:
Cardium nodosum Montagu. "1852 Leach: Cardium
nodosum Mont. = Testa junior Cardium tuberculatum.
"1856 Wood & Hanley No. 7: Cardium nodosum Mont.
T. B. (not Turt.). "1858 Adams & Adams, p. 458:
Hemicardia (Fragum) nodosa Mont. .. 1864 Mayer: C.
nodosum (comparison with Cardium papil/osum Poli.)
[Tertiary). "1868 ZR: Cardium nodosum (Mont.) is
perhaps not = roseum (Lam.), or punctatum (Brocchi),
orscabrum(Phil.) [Ref. Romer 1868]. "1871 Morch,
p. 215, No. 204: Cardium (Cerastoderma) nodosum
Mtg. "1888 Petersen: Cardium nodosum Mtg. "1890
Paetel: Cardium nodosum Mtg. "1907 ZR: Cardium
nodosum [Refs. Colgan, Hoel, 0yen]. "1922-32
Sherborn, p. 4401 : Cardium nodosum Montagu, 1803 .
"1932 Winckworth: Cardium nodosum = Cardium
(Parvicardium) scabrum Philippi, 1844 ... 1970 Glibert
& Poel: Parvicardium nodosum (Montagu, 1803).
"1977 Fischer-Piette: Cardium (Cerastoderma,
Parvicardium) nodosum Montagu. "1978 Janssen &
Van der Slik: Cardium nodosum Montagu, 1803 =
Cardium tuberculatum Linnaeus. .. 2000 Aartsen &
Goud: Cardium nodosum Montagu, 1803 not Gmelin,
1791 is a synonym of Parvicardium scabrum (Philippi,
1844) . .. 200 I Vidal, p. 408 : Parvicardium nodosum . CHARACTERS AND
OCCURRENCE
DESCRIPTIONS. "1845 Reeve: shell orbicular, rather
depressed, posteriorly slightly angulated ; radiately
Phuket Marine Biological Center Special Publication vol. 30 (2004)
651
nodosum - nodulosum
ribbed, 25-26 ribs, conspicuously obtusely noduled
whitish. 002001 Vidal : only the exhalant aperture is
tube-shaped. Inhalant siphon open towards the foot
aperture. No eyes. REMARKS. 001845 Reeve: each rib is
surmounted with a row of small blunt nodules. HABITAT.
001876 Duprey: Cardium nodosum Turton intertidal on
the coast of New Jersey with a tidal range of about 13
m . 001969 Nordsieck: 9-420 m depth, in the North Sea
from 42 m depth and down. DISTRIBUTION. 001807 Maton
& Racket!: Falmouth in Cornwall. 001845 Reeve:
Britain. 001856 Wood & Hanley: Britain. ooOf authors:
North Norway, Ireland, Canary Islands, Mediterranean.
nodosum Mourlon 1880. - Replaced by Parvicardium
straelini (Gilbert, 1945) . » Regarding Cardium
nodosum: Cardiidae. Miocene. REcORDs. 001947 ZR:
Cardium nodosum Mourlon 1880-1881 non Montagu,
1803, = Cardium (Parvicardium) straelini Gilbert, 1945
Miocene Anversian, Belgium.
nodosum Turton, 1822. - Identical with Parvicardium
scabrum (Philippi, 1844). OTHER JUDGEMENTS :
Cardium muricatulum jun., auctorurn. - Plagiocardium
papil/osum (Po Ii). - Cardium punctatum Broc. - juv.
Cardium tubercula tum Linnaeus. - Cardium roseum
Lamarck, 1818. » Regarding Cardium nodosum:
Cardiidae. Recent. REcORDs. 001822 Turton: Cardium
nodosum . 001855 Clark: C. nodosum Turton = C.
muricatulum jun., auctorum. 001856 Jeffreys: Cardium
nodosum (Turton) B.M (= "British Mollusca") =
Cardium papillosum (Poli) Ph . 001857 Deshayes :
Cardium nodosum Turton. 001859 Danielssen: Cardium
nodosum Turton = Cardium scabrum Philippi. 001859
Sowerby: C. nodosum Turt. = Cardium punctatum
Broc. 001867 ZR: Cardium nodosum (Turt. and other
authors) [Ref. Weinkauff 1867]. 001868 March:
Cardium (Cerastoderma) nodosum Turt. 001868 ZR:
Cardium nodosum (Mont.) is perhaps not = roseum
(Lam.), or punctatum (Brocchi), or scabrum (Phil.)
[Ref. Romer 1868]. 001882 Cooke: Cardium nodosum
Turt. = C. pygmaeum Don. = C. exiguum Gmel. 001884
Grasset: Cardium nodosum Turt. 001886 Locard:
Cardium nodosum Turton, 1822 = Cardium roseum de
Lamarck, 1818. 001888 Servain : Cardium nodosum
Turton, 1822 = Lamarck, 1818. 001892 Locard:
Cardium nodosum Turton, 1822. 001900 Bragger:
Cardium nodosum, Turt. 001900 Locard & Caziot:
Cardium nodosum Turton, 1822. 001906 Dautzenberg
& Durouchoux: Cardium (Parvicardium) nodosum
Turton. 001933 Coen: Cardium nodosum Turton =
Eucardium ( Cardium) echinatum var. roseum Lamarck.
001933 Coen & Vatova: Eucardium (Rudicardium)
nodosum Turton. 001937 Coen: Cerastoderma
(Rudicardium) nodosum Turton. 001947 ZR: Cardium
nodosum Turton, mode ofiocomotion, figs. [Ref. Stoll
1947]. 001962 Pasteur-Humbert: Cardium nodosum
Turton. 001969 Nordsieck: Parvicardium nodosum
(Turton, 1822). ooScariato & Starobogatov: Cardium
nodosum Turt. in Kobelt 1898 = Plagiocardium
papillosum (Poli, 1791). 001978 Janssen & Van der Slik:
Cardium nodosum Turton, 1822 = Parvicardium
scabrum (Philippi). 001998 Macedo et at.: Cardium
nodosum Turton, 1822 non Montagu, 1803 =
Parvicardium scabrum (Philippi, 1844). CHARACTERS AND OCCURRENCE - DISTRIBUTION. 001884 Grasset: Algeria.
nodosum ovata Jeffreys (variety). - Infrasubspecific. No
evidence of subspeci fic status. Parvicardium scabrum
(Philippi, 1844). » Regarding Cardium nodosum var.
ovata: Cardiidae. Recent. REcoRDs. 001897 Marshall :
Cardium nodosum var. ovata Jeff.
nodosum rosea Lamarck (variety). - Infrasubspecific.
No evidence of subspecific status. Parvicardium
scabrum (Philippi, 1844). » Regarding Cardium
nodosum var. rosea: Cardiidae. Recent. RECORDS.
001876 Duprey: Cardium nodosum var. rosea Lam.
001897 Marshall : Cardium nodosum var. rosea Lamk.
001901 Chaster et at.: Cardium nodosum var. rosea
Lamarck. - CHARACTERS AND OCCURRENCEDISTRIBUTION. Intertidal on the coast of New Jersey with
a tidal range of about 13 m (Duprey 1876).
nodulosum Lamarck . [Figs. part 3 p . 847]. Acanthocardia tuberculala. Lamarck collection MNHN
Paris. No published records encountered.
nodulosum Orbigny, 1850. - Conocardium nodulosum
d 'Orbigny, 1850. Conocardiidae. Fossil. RECORDS.
001922-32 Sherborn: Pleurorhynchus nodulosum
M'Coy. 001922-32 Sherborn, p. 4405: Conocardium
(M'Coy) nodulosum d'Orbigny, 1850. - Pleuro-
rhynchus.
nodulosum Spengler, 1799. [Figs. Part 1 p. 93 . Part 3 p.
906]. - ? Americardia sp. OTHER JUDGEMENTS: Cardium
medium Linnaeus, 1758. » Regarding Cardium
nodulosum: Cardiidae. Recent. TYPE LOCALITY. The East
Indies (Spengler 1799. TYPE REPOSITORY. ooZMUC: I
specimen (Spengler coiL). RECORDS. 001799 Spengler,
p. 57, No. 37, pI. I, figs. 4 a, b: Cardium nodulosum.
001808 Spengler, p . 131: Cardium nodulosum. 001870
Morch: Cardium nodulosum Spengler; good species
related to C. imbricatum Sow. 001902. Sherborn, p. 669:
Cardium nodulosum Spengler, 1799. 001977 FischerPiette: Cardium nodulosum Spengler = Corculum
medium Linne. 001999 Knudsen & HylJeberg: Cardium
nodulosum Spengler, 1799. - CHARACTERS AND
OCCURRENCE - CHARACTERS of diagnostic value: a
three-sided cockle, posterior slope marked, 10 very
smooth ribs, margin dentate. Shell surface with about
20 ribs, narrower than on the posterior slope, rounded
tops ornamented with nodules instead of scales, narrow
interstices. DESCRIPTIONS. 001799 Spengler: testa
subcordata aequivalvi, carinato-angulata, sulcis
nodulosis. 001799 Spengler translated by Knudsen &
Hylleberg 1999: this triangular cockle is a distinct and
new species, which is not published by any author. After
its heart-shaped front side, which is encased by an
elevated blunt crest, lengthwise broadest at the middle,
towards the side of the back hollowed, but in the middle
elevated . It is covered by ten glossy ribs, which on the
margin, where both valves closely interlock, are incised
with serrate teeth. Deeply incised ribs, which however
are narrower than those on the front side, and which
on their rounded backs are furnished with tubercles
instead of scales, cover the two inflated surfaces. The
backside of the bivalve, from the umbo to the lower
margin, is rather long, not very much shorter than the
anterior margin. The bivalve is white all over, only
inside is the front side yellow. DIMENSIONS . 001799
Spengler: length 33 mm (1.25 Tom.), width 26 mm (I
Tom.). REMARKS . It is not clear which specimen
Spengler (1799) has described. The type specimen at
652
Tropical Marine Mollusc Programme (TMMP)
nodulosum - norvegicum
ZMUC has dense chevrons on top of radial ribs (like
in C. medium Linne, from the West Indies). The figures
(4 a, b) by Spengler indicates that he apparently
described a white specimen of Fragum unedo. His figs.
do not show the characters of medium Linne so the
original specimen must have been lost. DISTRIBUTION.
--1799 Spengler: Its home is the East Indies [?error,
should be the W. Indies if the species is C. medium L.].
nodulosum Wood, 1853. - Not valid. » Regarding
Cardium nodulosum: Cardiidae. Tertiary. Synonym of
Parvicardium papil/osum (Po Ii, 1791). REcORDs. --1864
Mayer: C. nodulosum (comparison with Cardium
papillosum. Poli.). "1867 Weinkauff: Cardium
nodulosum Wood, (Crag Moll.) = Cardium papillosum
Poli. --1888 Kobelt: Cardium nodulosum Wood (fide
Weinkauff) = Cardium papillosum Poli. "1922-32
Sherborn, p. 4405: Cardium nodulosum Wood, 1840
[n.n.]. --1970 Glibert & Poel: Parvicardium nodulosum
(Wood, 1853).
noeggerathi 1. Muller, 1851. - Changed combination:
Pleuriocardia noeggerathi (J. Muller, 1851). Cardiidae.
Cretaceous . RECORDS. --1871 Stoliczka: Cardium
noeggerathii (Laevicardium). "1889 ZR: Cardium
noeggerathi with fig. (ref. Holzapfel). --1931 ZR:
Cardium noeggerathi Ref. to Rieder (1931). "1980
Ruhoff: Cardium noeggerathi J. Muller, 1851; part 2:
65 . "1986 Abdel-Gawad: Cardium noeggerathi J.
Miill. in Andert 1934. --1986 Abdel-Gawad, pI. 40,
fig. 10: Pleuriocardia (Pleuriocardia) noeggerathi (1.
Muller, 1851). "1999 Jacques Vidal (pers. com.),
MNHN, Paris: Fossils from the Paleontological
Collection as Cardium noeggerathi Muller. Lower
Senonien. Germany, B 18197.
nomurai (subspecies). - See under notabilis nomurai
Otuka,1943.
nomurai Hayasaka, 1957. - Combination to be revised:
"Clinocardium" nomurai Hayasaka, 1957. Cardiidae.
Pliocene. RECORDS. --1957 ZR: "Clinocardium"
nomurai Hayasaka, 1957 sp. nov. Pliocene, Futaba
district, Fukushima Prefecture, Japan.
nomurai Kuroda & Habe, 1951. - New name: Nemocardium nomurai Kuroda & Habe, 1951 . Cardiidae.
Recent. TYPE LOCALITY. Taiwan (fossil) (Higo et al.:
1999). GENOTYPE. Habe (1951) designated Cardium
nomurai Kuroda & Habe, 1951 type species of the
subgenus Trifaricardium Habe, 1951. RECORDS. --1951
Kuroda & Habe: Cardium nomurai. --1951 Habe:
Cardium nomurai Kuroda & Habe, 1951. --1952
Kuroda & Habe: Cardium (Trifaricardium) nomurai
Kuroda & Habe. --1961 ZR: Trifaricardium nomurai
Kuroda & Habe, 1961 nom. nov. pro Cardium
(Acanthocardia) cancellatum Nomura 1933 non
Gmelin 1791, Japan. "1962 ZR: Trifaricardium
nomurai Kuroda & Habe in Kuroda, 1962 nom. nov.
for Cardium cancel/ata Nomura 1933, non Gmelin
1791. --1973 Keen: Nemocardium (Trifaricardium)
nomurai Kuroda & Habe, 1951. --1977 Fischer-Piette:
Trifaricardium nomurai Kuroda & Habe. --1977 Habe:
Trifaricardium nomurai Kuroda & Habe, 1951 . --1977
Popov: Pratulum nomurai (Kuroda & Habe). "1981
Koyama et al.: Trifaricardium nomurai. --1995
Schneider: Nemocardium (Trifaricardium) nomurai
Habe, 1951. --1999 Higo et al.: Trifaricardium nomurai
Kuroda & Habe in Habe, 1951 . - CHARACTERS
AND OCCURRENCE - REMARKS. --1973: Keen stated
that Nemocardium (Trifaricardium) nomurai Kuroda
& Habe, 1951 is a new name for Cardium cancellatum
Nomura, 1933, preoccupied. HABITAT. "1999 Higo et
at.: 100-200 m, mud and sand. DISTRIBUTION . "1994
Hylleberg: Japan. --1999 Higo et al.: Mie prefecture
(offKii-Nagashima; offOwase Matsumoto, 1979) and
southwards, Kii peninsula (Kuroda & Habe, 1981),
Tosa bay; South China Sea.
noquam (error). - Printer's mistake for nequam in Ma
Qi-hong et al. 1976 .
nordenskioeldi (subspecies). - See under cilia tum
nordenskioeldi Kafanov, 1981.
nortonensis Cox, 1946. - Tutcheria nortonensis Cox,
1946. Cardiacea. Middle Lias. RECORDS. --1946 ZR:
Tutcheria nortonensis Cox, 1946 sp. nov. Middle Lias
of Chipping Norton, Oxon.
norvegicum Spengler, 1799. [Figs. Part I p. 94, 175.
176. Part 3 p. 895, 896] . - Synonym of Laevicardium
crassum (Gme\in, 1791). » Regarding Cardium
norvegicum: Cardiidae. Miocene and Recent. TYPE
SPECIES. Sacco (1899) stated that Laevicardium
norvegicum (Spengl.) is type species of subgenus
Laevicardium Swainson, 1840. TYPE LOCALITY. Bergen,
Norway (Spengler 1799). TYPE REPOSITORY. --ZMUC
(Spengler collection). RECORDS. --1799 Spengler, p. 42,
No. 26: Cardium norvegicum. --1808 Spengler:
Cardium norvegicum. --1840 Swainson : Cardium
norvegicum Spengler type of the genus Laevicardium
(in Nordsieck 1969, p. 97). "1846 Loven: Cardium
norvegicum Spengler. "1855 Clark: C. norvegicum
Spengler = C. laevigatum auctorum. --1856 Wood &
Hanley: C. norvegicum Speng. is to be preferred for
Cardium laevigatum Penn. --1857 Deshayes: Cardium
norvegicum Spengler. --1858 Adams & Adams, p. 457:
Laevicardium norvegicum, Spengl. .. 1867 ZR:
Cardium norvegicum (Spengler) = laevigatum of the
Mediterranean conchologists [Ref. Weinkauff 1867].
"1868 Colbeau: Laevicardium norvegicum Spgl.
"1868 ZR: Cardium norvegicum (Spengl.) = C.
laevigatum of Pennant not Linne = C. vitellinum [Ref.
Romer 1868]. "1868 ZR : Cardium norvegicum
(Spengl.). Living specimens observed in aquarium at
Arcachon; makes considerable leaps [Ref. Crosse
1868]. "1871 Morch, p. 215, No. 200: Cardium
norvegicum Spgl. = Cardium (Liocardium) crassum
Gm. --1872 Tryon: Laevicardium norvegicum Spengler.
"1884 Cundall: Cardium norvegica. "1884 Grasset:
Laevicardium norvegicum Spgl. "1885 Smart:
Cardium norvegicum Spengl. --1885 Smith: Cardium
norvegicum Spengler, Forbes and Hanley, pI. 31, figs.
1-2 = Cardium (Laevicardium) var. norvegicum
Spengler. --1886 Locard: Cardium norvegicum
Spengler, 1792 sic. --1887 Fischer: Laevicardium
norvegicum Spengler. --1888 Petersen: Cardium
norvegicum Spgl. --1888 Servain: Cardium norvegicum
Spengler, 1792 sic. --1890 Paetel: Cardium
norwegicum Spglr. "1890 Paetel: Laevicardium
norwegicum Spglr. "1892 Locard, p. 307: Cardium
norvegicum Spengler, 1792 sic. "1899 Sacco:
Laevicardium norvegicum (Spengl.) (an var. fragilis
Br.). --1899 Sacco: Laevicardium norvegicum
(Spengler 1790) [Miocene-Recent]. --1900 Locard &
Caziot: Cardium norvegicum Spengler, 1790 sic. --1902
Phuket Marine Biological Center Special Publication vol. 30 (2004)
653
norvegicum - norvegicum gibba
Dall: Cardium norvegicum Spengler, type of subgenus
Laevicardium Swainson. --1902 Sherborn, p. 670 :
Cardium norvegicum Spengler, 1799 . --1906
Dautzenberg & Durouchoux: Cardium (Laevicardium)
norvegicum Spengler. --1907 ZR: Cardium Ilorvegicum
[Refs. Christensen, Colgan]. "1912 Dautzenberg &
Fischer: Cardium (Laevicardium)norvegicum Spengler
1799. --1912 ZR: Cardium (Laevicardium) norvegicum
Speng. [Ref. Dautzenberg & Fischer 1912]. --1927
Dautzenberg: Cardium (Laevicardium) norvegicum
Spengler = Cardium (Laevicardium) crassum Gmelin.
--1931 ZR: Cardium norvegicum Ref. to Patrini (1931).
--1933 Coen & Vatova: Laevicardium norvegicum
Spengler var. gibba Jeffreys. --1962 Pasteur-Humbert:
Laevicardium norvegicum (Spengler) . --1969
Nordsieck : Laevicardium norvegicum norvegicum
Spengler, 1790 sic. "1969 Nordsieck: Cardium
norvegicum Spengler, 1790 sic = Laevicardium
crassum (Gmelin , 1791). "1977 Fischer-Piette:
Cardium norvegicum Schr., Spengler, 1799 =
Laevicardium crassum GmeIin. --1977 ZR :
Laevicardium norvegicum. Shell. Aberrant specimen.
Ref to Entrop, B. --1970 Glibert & Poel : Cardium
norvegicum Spengler, 1799 = Laevicardium (s.s.)
crassum crassum Gme1in, 1791. --1977 Fischer-Piette
p. 18: Cardium norvegicum in Jeffreys 1856 = Cardium
oblongum Gmelin. --1978 Janssen & Van der Slik:
Cardium Ilorvegicum Spengler = Laevicardium
(La evicardium) crassum (Gmelin, 1791). "1983
Anaya: Laevicardium (s. s.) norvegicum (Spengler,
1790 sic) [Pliocene] ... 1998 Macedo et al. : Cardium
norvegicum Spengler, 1799 = Laevicardium crassum
(Gmelin, 1791) . "1999 Knudsen & Hylleberg:
Cardium norvegicum Spengler, 1799 . .. 1999 Jacques
Vidal (pers. com.), MNHN, Paris: Fossils from the
Paleontological CoIlection as Laevicardium
norvegicum Spengler. Pliocene-Quaternary. Morocco,
B38093 to -95. B44385 . Italy, B40619, B36098,
B36099. Egypt, B42202 . - CHARACTERS AND
OCCURRENCE - OF COLLECTIONS . --ZMUC: 2
specimens from Bergen in Norway, the Spengler coil.
CHARACTERS of diagnostic value: shells thick, glossy,
not gaping, anterior part round like part of a circle,
posterior slope smooth, nymphs broad and deeply set,
ribs flat vanishing towards the umbo, 2 cardinals in
each shell, Colour yellow outside, white inside.
DESCRIPTIONS. --1799 Spengler: testa cordata
aequivalvi, laevi sulcis transversis antiquatis, cos tis
obsoletis. --1799 Spengler translated by Knudsen &
Hylleberg 1999: this Cardium is as long as it is broad.
Its actual length on the front side is reckoned from the
umbo down to the lower margin. The posterior end of
the bivalve is very short and round as a circular arc.
The front side is only slightly projecting and smooth
in the middle. Both valves have here only quite flattened
teeth and are closely connected. The nymphs are very
broad and placed deeply in the bivalve. Both surfaces,
on their lower part towards the middle, have flattened
elevated ribs, whose backs are straight and separated
by furrows . But these ribs become indistinct upwards
and leave only faint traces, which become almost
invisible due to the lustre by which the bivalve becomes
very shining. This cockle has two hinge teeth in each
valve, which is usual in this genus. The whole margin
of the bivalve is provided with small teeth by which
both valves interlock tightly. The valves are thick, light
yellow outside, and snow-white inside. DIMENSIONS.
--1799 Spengler: height and width 65 mm (2 1/2 Tom.).
HABITAT. --1969 N ordsieck: generally recorded from 7
to 180 m depth (maximum 2195 m) ... 1876 Duprey:
intertidal on the coast of New Jersey with a tidal range
of about 13 m. DISTRIBUTION. --1799 Spengler: Bergen
in Norway. --1856 Wood & Hanley: Britain &c. --1884
Grasset: England. --1969 Nordsieck: North Europe,
West Africa, Mediterranean; Atlantic Ocean from
Lofoten to the Ivory Coast.
norvegicum Jeffreys (variety) . - Infrasubspecific. »
Regarding Laevicardium norvegicum var: Cardiidae.
Recent. RECORDS. --1899 Sacco: Laevicardium
norvegicum var. Jeffr. --1969 Nordsieck: Laevicardium
norvegicum norvegicum (Spengler, 1790 sic) var.
Jeffreys.
.
norvegicum Schrater 1786 (variety). - Infrasubspecific.
» Regarding Cardium norvegicum var: Cardiidae.
Recent. TYPE LOCALITY. Mediterranean. TYPE REPOSITORY.
--ZMUC (Spengler coli.). OF COLLECTIONS. Spengler p.
42 . No. 26b, --ZMUC : I specimen from the Mediterranean in Spengler coli. RECORDS. --1784 Schrater,
p. 57, No. 12 (1786), pI. 7, fig. 12 (1784) : Cardium
var. norvegicum. --1799 Spengler, p. セR L@ No. 26:
Cardium var. norvegicum in Schrater. 3 r . vol. p.57,
no. 12. Tab. 7 fig. 12 ... 1870 March: Cardium var.
norvegicum Spengler, Schroter Einl. = Cardium
crassum Gmelin. --1885 Smith: Cardium norvegicum
Spengler, Forbes and Hanley, pI. 31, figs. 1-2 =
Cardium (Laevicardium) var. norvegicum, Spengler.
--1999 Knudsen & Hylleberg: Cardium norvegicum
var. Spengler, 1799 (the Mediterranean) . CHARACTERS AND OCCURRENCE - CHARACTERS
of diagnostic value: larger than C. norvegicum, more
inflated, but otherwise similar to C. norvegicum from
Norway. DESCRIPTIONS. --1799 Spengler translated by
Knudsen & Hylleberg 1999 : This bivalve is
considerably larger, thicker and has a more solid shell ,
more strongly inflated; otherwise very similar to the
preceding one [Cardium norvegicum]. It belongs to the
Mediterranean. DIMENSIONS. --1799 Spengler: height
and width 72 mm (2 3/4 Tom.).
norvegicum Spengler, 1799 (variety). [Figs. Plate 3 p.
895] . - See norvegicum var. Schrater.
norvegicum dertogibba Sacco, 1899 (variety). - Identical
with Laevicardium crassum (Gmelin, 1791). »
Regarding Laevicardium norvegicum var. dertogibba:
no evidence of subspecific status. Cardiidae. Recent.
RECORDS. "1899 Sacco, p. 52, pI. 11 , fig. 45:
Laevicardium norvegicum var. dertogibba Sacco
norvegicum devulIls Bucquoy, Dautzenberg & Dollfus,
1892 (variety). - Identical with Laevicardillm crassum
(Gmelin, 1791). » Regarding Cardium norvegicum
var. devians: Infrasubspecific. No evidence of
subspecific status. Cardiidae. Recent. RECORDS. --1892
Bucquoy, Dautzenberg & Dollfus, Cardium
norvegicum vaT. de vians. --1969 Nordsieck:
Laevicardium norvegicum norvegicum var. devians
Bucquoi sic, Dautzenberg & Dollfus . Unknown
distribution.
norvegicum gibba (mode). - See under norvegicum
gibbllm Jeffreys, 1863.
654
Tropical Marine Mollusc Programme (TMMP)
norvegicum gibbum - norvegicum senegalensis
norvegicum gibbum Jeffreys, 1863 (variety). - Identical
with Laevicardium crassum (Gmelin, 1791). »
Regarding Laevicardium norvegicum var. gibba:
norvegicum minor (variety) - Identical with
Laevicardium crass urn (Gmelin, 1791). » Regarding
Cardium norvegicum var. minor: infrasubspecific. No
Infrasubspecific. No evidence of subspecific status.
Cardiidae. Recent. RECORDS. ··1933 Coen : Laevicardium norvegicum Spengler var. gibba Jeffreys .
··1937 Coen: Laevicardium norvegicum gibbum
Jeffreys. Described as a variety. ··1856 Jeffreys:
Cardium norvegicum var gibbum . ··1897 Marshall :
Cardium norvegicum var. gibba Jeff... 1899 Sacco:
Laevicardium norvegicum var. gibba (Jeffr.). ··1899
Sacco: Cardium norvegicum var gibba Jeffrey I sic
1863. ··190 I Chaster et al.: Laevicardium norvegicum
var. gibba Jeffreys. ··1969 Nordsieck: norvegicum var.
gibbum Jeffreys = norvegicum var senegalensis
Dautzenberg. ··1980 Ruhoff: Cardium norvegicum
gibba Jeffreys, 1863; [1862-1869] vol. 2: 295. CHARACTERS AND OCCURRENCE - DESCRITIONS.
··1969 Nordsieck: more inflated, lighter weight, more
marked sculpture . REMARKS. ··1969 Nordsieck:
Laevicardium norvegicum Spengler, 1799 =
Laevicardium crassum (Gmelin, 1791). DISTRIBUTION.
Unknown.
norvegicum lineolatum Bucquoy, Dautzenberg &
Dollfus, 1892 (variety). - Identical with Laevicardium
crassum (Gmelin, 1791). » Regarding Cardium norvegicum var. lineo/atum: infrasubspecific. No evidence
of subspecific status . Cardiidae. Recent. RECORDS.
··1892 Bucquoy, Dautzenberg & Dollfus , 1892:
Cardium norvegicum var. lineolatum . ··1969 Nordsieck: Laevicardium norvegicum norvegicum (Spengler, 1790 sic) var. lineolatum Bucquoi sic, Dautzenberg
& Dollfus, 1889 sic. Unknown distribution .
norvegicum marmoratum Bucquoy, Dautzenberg &
Dollfus, 1892 (variety). - Identical with Laevicardium
crassum (Gmelin, 1791 ).» Regarding Cardium
norvegicum var. marmoratum: no evidence of sub spec ific status. Cardiidae. Recent. RECORDS. ··1892 Bucquoy,
Dautzenberg & Dollfus, 1892: Cardium norvegicum
var. marmoratum. ··1969 Nordsieck: Laevicardium
norvegicum norvegicum (Spengler, 1790 sic) var.
marmoratum Bucquoi sic, Dautzenberg & Dollfus .
Unknown distribution .
norvegicum mediterraneum Bucquoy, Dautzenberg &
Dollfus, 1892 (variety). - Identical with Laevicardium
crassum (Gmelin, 1791). » Regarding Cardium
norvegicum mediterranea: infrasubspecific. No
evidence of subspecific status. Cardiidae. Recent.
RECORDS. ··1892 Bucquoy, Dautzenberg & Dollfus,
evidence of subspecific status . Cardiidae. Recent.
RECORDS. ··1900 Locard & Caziot: Cardium
norvegicum var. minor. Mediterranean.
norvegicum pallidum Jeffreys, 1863 (variety). Identical with Laevicardium crassum (Gmelin, 1791).
» Regarding Cardium norvegicum var pallidum:
infrasubspecific. No evidence of subspecific status.
Cardiidae. Recent. RECORDS. ··1863 Jeffreys: Cardium
norvegicum var pallidum. ··1896 Tregelles: Cardium
norvegicum var. pallida Jeff. ··1897 Marshall: Cardium
norvegicum var. pallida Jeff. 1901 Chaster et al.:
Laevicardium norvegicum var. pallida Jeffreys. ··1969
Nordsieck: Laevicardium norvegicum var. pallidum
(Jeffreys). ··1980 Ruhoff: Cardium norvegicum pallida
Jeffreys , 1863 ; [1862-1869] vol. 2: 295. CBARACTERS AND OCCURRENCE - REMARKS.
··1969 Nordsieck : yellow colour. HABITAT . "1876
Duprey: Cardium norvegicum var. pal/ida intertidal on
the coast of New Jersey with a tidal range of about 13
m. DISTRIBUTION. "1969 Nordsieck: unknown.
norvegicum ponderosum Bucquoy, Dautzenberg &
Dollfus, 1892 (variety). - Identical with Laevicardium
crassum (Gme\in, 1791). » Regarding Cardium
norvegicum var. ponderosum: infrasubspecific. No
evidence of subspecific status. Recent. DESCRIPTIONS.
Shell thick, heavy. DISTRIBUTION. Unknown (Nordsieck
1969). RECORDS . ··1952 Lecointre : Cardium
norvegicum Spengler var. ponderosa de Bory. ··1892
Bucquoi, Dautzenberg & Dollfus: Cardium norvegicum
var. ponderosum . ··1969 Nordsieck: Laevicardium
norvegicum var. ponderosum Bucquoi sic, Dautzenberg
& Dollfus, 1889.
norvegicum rotundum Jeffreys (variety). - Identical with
Laevicardium crassum (Gmelin, 1791). » Regarding
Cardium norvegicum var. rotunda: infrasubspecific. No
evidence of subspecific status. Cardiidae. Recent.
REMARKS. Laevicardium norvegicum (Spengler, 1799)
= Laevicardium crassum (Gmelin, 1791). DESCRIPTIONS.
More round and flat (Nordsieck 1969). DISTRIBUTION.
Unknown (Nordsieck 1969). RECORDS . ··1896
Tregelles : Cardium norvegicum var. rotunda Jeff.
"1897 Marshall : Cardium norvegicum var. rotunda
Jeff. "1901 Chaster et at.: Laevicardium norvegicum
var. rotunda Jeffreys. ··1969 Nordsieck: Laevicardium
norvegicum norvegicum var. rotundum Jeffreys. ··1980
Ruhoff: Cardium norvegicum rotunda Jeffreys, 1863;
[1862-1869] vol. 2: 295.
norvegicum senegalensis Dautzenberg, 1891 (variety).
- Laevicardium crassum (Gmelin, 1791 ) .OTHER
JUDGEMENTS: Laevicardium norvegicum norvegicum var
gibbum Jeffreys . » Regarding Laevicardium
norvegicum var. senegalensis: Cardiidae. Recent.
Infrasubspecific. No evidence of subspecific status.
RECORDS. ··1891 Dautzenberg, p. 59: Laevicardium
norvegicum Spengler var. senegalensis nov. ··1969
Nordsieck: Laevicardium norvegicum norvegicum
(Spengler, 1790 sic) var senegalensis Dautzenberg.
··1969 Nordsieck: Laevicardium norvegicum
norvegicum var. senegalensis Dautzenberg =
Laevicardium norvegicum norvegicum var gibbum
Jeffreys. - CHARACTERS AND OCCURRENCE -
1892: Cardium norvegicum mediterranea . ··1899
Sacco : Laevicardium norvegicum var. mediterranea
B.D.D. ··1900 Locard & Caziot: Cardium norvegicum
Spengler, 1790 sic var. mediterranea . ··1906
Dautzenberg & Fischer: Cardium (L.) norvegicum var.
mediterranea, Bucquoy, Dautzenberg & G. Dollfus
1892. ··1937 Coen: mediterraneum Monter. =
Laevicardium norvegicum gibbum Jeffreys. ··1969
Nordsieck: Laevicardium norvegicum mediterraneum
Bucquoi sic, Dautzenberg & Dollfus, 1889. CHARACTERS AND OCCURRENCE - REMARKS.
··1969 Nordsieck : La e vicardium norvegicum
mediterraneum coexists with a typical Atlantic form ,
e.g. at Marseilles. DISTRIBUTION. ··1900 Locard &
Caziot: Mediterranean.
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655
norwegicum - nuciforme
DESCRIPTIONS. ··1969 Nordsieck: more inflated, light
weight, marked sculpture. REMARKS. Laevicardium
norvegicum Spengler, 1790 = Laevicardium crassum
(Gmelin, 1791). DISTRIBUTJON. ··1969 Nordsieck :
unknown.
norwegicum erroneous spelling of norvegicum.
notabile (mode). - See under notabilis.
notabilis Sowerby, 1915. [Figs. Part 3 p. 854]. - Serripes
notabilis (Sowerby, 1915). OTHER JUDGEMENTS:
Serripes notabilis nomurai Otuka, 1943. - Serripes
groenlandicus (Bruguiere, 1789). » Regarding
Serripes notabilis: Cardiidae. Recent. TYPE LOCALITY.
Japan. DISTRIBUTION. Japan (Hylleberg 1994). RECORDS.
"1915 Sowerby, p. 169, pI. 10, fig. 9: Cardium
(Serripes) notabile. ··1915 ZR: Cardium (Serripes)
nota bile Sowerby, 1915 sp. n. Japan. ··1952 Kuroda &
Habe: Serripes notabilis Sowerby. ··1954 Taki:
Serripes notabi/is Sowerby. ··1962 Habe: Serripes
notabilis Sowerby. ··1964 Habe: Serripes notabilis
Sowerby. ··1964 Shikama: Serripes notabilis
(Sowerby). ··1967 Habe: Serripes notabilis. ··1973
Keen: Serripes notabilis (Sowerby, 1915) = S. notabilis
nomurai Otuka, 1943 (fide Noda, 1962). ··1975 ZR:
Cardium (Serripes) notabile Saverby sic, 1915.
Designated type species for new genus Yagudinella,
Ref. to Kafanov. ··1977 Popov: Serripes notabilis
(Sowerby). ··1977 Fischer-Piette: Serripes notabilis
Sowerby. ··1977 Habe: Cardium notabilis Sowerby,
1915 = Serripes groenlandicus (Bruguiere, 1789).
··1978 Kafanov: Yagudinella notabilis (Sowerby,
1915). ··1980 ZR: Yagudinella notabilis (Sowerby
1915). Removal from synonymy. Y notabilis nomurai
(Otuka (943). Ref. to Kafanov. ··1995 Chung-Hung
& Hsi-len: Serripes notabilis (Sowerby). ··1998
Schneider (I 998b ): Yagudinella notabile (Sowerby,
1915). ··1999 Higo et at.: Cardium notabilis Sowerby,
1915 = Serripes groenlandicus (Bruguiere, 1789).
··2000 Coan et al.: Serripes (Yagudinella) notabilis
(Sowerby, 1915).
notabilis nomurai Otuka, 1943 (subspecies). - Synonym
of Serripes groenlandicus (Bruguiere, 1789). »
Regarding Serripes notabilis nomurai: Cardiidae.
Recent. RECORDS. ··1975 ZR : Yagudinella notabilis
nomurai Otuka, 1943 comb. nov. transferred from
Serripes. Ref. to Kafanov, p. 147. ··1978 Kafanov:
Serripes notabilis nomurai Otuka, 1943 = Yagudinella
notabi/is (Sowerby, J 915). ··1980 ZR: Yagudinella
notabilis nomurai (Otuka 1943). Removal from
synonymy with Y notabilis (Sowerby 1915). Ref. to
Kafanov. ··2000 Co an et al.: Serripes notabi/is nomurai
Otuka, 1943 = Serripes (Yagudinella) notabilis
(Sowerby, 1915)
novacaspica (subspecies). - See under protraeta
novaeaspiea Glazunova, 1971.
novaezelandiae Deshayes. - Unknown species. »
Regarding Cardium novaezelandiae : Cardiidae.
Recent. RECORDS. ··1885 Smith: Cardium novae
zelalldiae Deshayes, MS. in Mus. Cuming (?) =
Cardium (Bueardium) pulehellum Gray.
*novakovskii. - Homonymy to be noticed. Cardium
novakovskii from 2 sources: Andrussow, 1902. Andrusov, 1963. - REMARKS. I do not know how
Andrussow 1902 compares with Andrusov 1963. novakovski Andrussow, 1902. - Combination to be
revised: "Cardium" novakovski Andrussow, 1902.
Cardiac ea. Fossi I. RECORDS . ··1905 Dollfus,
commenting Andrussow 1902: Cardium Ilovakovski n.
sp. tends towards Cerastoderma.
novakovskii Andrusov, 1963. - Combination to be
revised: "Cardium " Ilovakovskii Andrusov, 1963.
Cardiacea. Miocene. RECORDS. ··1961 ZR: Cardium
novakovskyi Andrusov, 1961 "sp. nov."[in a table]
Miocene Caspian region, Russia. ··1963 ZR: Cardium
novakovskii Andrusov, 1963 sp. nov. Neogene, Russia
[on the pI. as novakovskyi].
novakovskyi (error). - Printer's mistake for novakovskii
in Andrusov, 1963.
novarossica Barbot de Marny, 1869 . - Changed
combination: Pontalmyra novarossiea (Barbot de
Mamy, 1869). Lymnocardiidae. Fossil. ··1899 Sacco:
Cardium novarossiea Barbot de Mamy 1869. RECORDS.
··1899 Sacco: Adaehna sic novarossica (Barb.). ··1899
Sacco: Pontalmyra novarossica (Barb.). ··1987
Taktakishvili: Didacna novorossiea (Barb.). ··1987
Taktakishvili: Pontalmyra novorossiea (Barb.).
novatum Deshayes, 1853. - New name. Combination
to be revised: "Cardium" novatum Deshayes, 1853.
Cardiidae. Fossil. RECORDS. ··1857 Deshayes: Cardium
novatum new name for Cardium cognatllm Phillips in
Goldfuss. ··1980 Ruhoff: Cardium novatum Deshayes,
1853; vol. 2: 47, 53.
novorossica (subspecies) . - See under incerta
novorossica Barbot de Mamy, 1869.
novum Coen , 1941. - Synonym of Acanthocardia
echinata (Linne, 1758). » Regarding Cardium novum:
Cardiidae. Recent. RECORDS. ··1941 Coen, p. 170 (pI.
I , figs. 12-14): Cardium (Acanthocardia) novum nov.
spec. ··1948 ZR: Cardium (A canthocardia) novum
Coen, 1941 sp. nov. Aci Frezza. ··1977 Fischer-Piette:
C. (Acanthocardia) novum = C. echinatum Linne.
novus Rio, 1994. - Dinocardium novus Rio, 1994.
Cardiidae. Middle Miocene. TYPE LOCALITY. Argentina.
RECORDS. ··1994 Rio, p. 106, pI. I, fig. 5: Dinocardium
novus sp. nov.
noyamiana Slodkewitsch, 1938. - Changed
combination: Fulvia noyamiana (Slodkewitsch, 1938).
Cardiidae. Miocene. TYPE LOCALITY. Sakhalin Island.
RECORDS . ··1945 ZR: Papyridea noyamiana 'Kogan'
sic Upper Miocene Sakhalin. ··1973 Keen: Fulvia
noyamiana (Slodkewitsch, 1938) [Papyridea).
Miocene, Sakhalin Is. ··1976 ZR: Profulvia noyamiana
(Slodkewitsch, 1938) Comb. nov. Transferred from
Papyridea. Ref. to Kafanov, p. 482 [orig. p. III].
Itucella. - Conoeardium nueella. Conocardiidae. Fossil.
RECORDS. ··1889 ZR: Conoeardium nucella with fig.
(ref. Barrois).
nuci lekelius, 1934 - Changed combination:
Euxinieardium nuei (Jekelius, 1934). Lymnocardiidae.
Pliocene. RECORDS. ··1935 ZR: Limnocardium nuci
lekelius, 1934 sp. n. Pliocene Rumania. ··1971 ZR:
Limnocardium nuei Jekelius, 1932 Combination
revalidated. Transferred back from Pseudoprosodacna
[Ref. Andreescu 1971]. ··1975 ZR: Limnocardium
(Eu.xinieardium) nuci lekelius. Emended diagnosis.
Ref. to Andreescu. 1. (139). ··1987 Taktakishvili:
Euxinieardium nuci (Jek.).
Ituciforme d'Orbigny, 1850. - Combination to be
revised: "Cardium" nuciforme d'Orbigny, 1850.
656
Tropical Marine Mollusc Programme (TMMP)
nuciformis - nuttallii
?Cardiidae. Fossil. RECORDs. "1922-32 Sherborn, p.
4433: Petricola nuciforme Sowerby, 1836. "1922-32
Sherborn, p. 4433: Cardium (Sow.) nuciforme
d'Orbigny, 1850. - Petricola.
nuciformis (variety). - See under edule nuciformis
Milachewitch, 1916.
nudum Miinster, 1840. - Combination to be revised:
"Cardium" nudum Miinster, 1840. ?Cardiidae. Fossil.
REcORDs. "1922-32 Sherborn, p. 4438: Cardium nudum
Muenster, 1840.
nummuliticum d' Archiac, 1850. - Not valid. »
Regarding Cardium nummuliticum: Combination to be
revised: "Cardium" nummuliticum d' Archiac, 1850.
Cardiidae. Lower Eocene. RECORDS. "1922-32
Sherborn, p. 4442: Cardium nummuliticum d' Archiac,
1850 [n.n.]. "1999 Jacques Vidal (pers. com.), MNHN,
Paris: Fossils from the Paleontological Collection.
Cardium nummuliticum d'Archiac. Turkey, B22343,
B22344.
nunninkae Lucas, 1984. - Cerastoderma nunninkae
Lucas, 1984. Cardiidae. Recent. DISTRIBUTION. Europe,
West Africa. RECORDS. "1984 Lucas: C. nunninkae.
"1984 ZR: Cerastoderma nunninkae sp. nov. Lucas,
1984. Bay of Biscay. Ref. to Lucas p. 24.
nutans Perkins, 1961. - Protocardia nutans Perkins,
1961. Cardiidae. Cretaceous. "1961 ZR: Protocardia
nutans Perkins, 1961 sp. nov. Cretaceous, Texas, Fort
Worth. "1986 Scott: Protocardia nutans Perkins, 1961.
nutaIlii (error). - Printer's mistake for nuttallii in the
present work Part I, p. 171, fig. 5.
nuttallianum Carpenter, 1857. - Synonym of
Clinocardium nuttallii (Conrad, 1837). »Regarding
Cardium nuttallianum: Cardiidae. Recent. RECORDS.
"1857 Carpenter (1856), p. 192: Cardium nuttallianum.
"1872 Carpenter: Cardium nuttallianum. "190 I Dall:
Cardium nuttallianum Carpenter, 1864 = Cardium
corbis Martyn, 1784. "1977 Fischer-Piette: Cardium
nuttallianum Carpenter = Corculum (Keenocardium)
nuttallii Conrad. "1980 Ruhoff: Cardium nuttallianum
Carpenter, 1857; p. 192.
nuttalliiConrad, 1837. [Figs. Part I p. 136, 171. Part 3
p. 852]. - Clinocardium nuttallii (Conrad, 1837).
Cardiidae. Miocene and Recent. TYPE SPECIES. Keen
(1936) designated Cardium nuttallii Conrad, 1837 as
type species of the genus Clinocardium Keen, 1936.
Gladenkov & Sinelnikova (1990) stated that Cardium
nuttallii Conrad, 1837 is type species of genus
Clinocardium Keen, 1936. Glibert & Poel (1970) stated
that Cardium nuttallii Conrad, 1837 is type species of
subgenus Clinocardium Keen, 1936. TYPE LOCALITY. A
few miles from the estuary of the Columbia River. TYPE
REPOSITORY. Academy of Natural Sciences, Philadelphia,
Cat. No. 54036. RECORDS. "1837 Conrad: Cardium
nuttallii. "1845 Reeve, sp. 66: Cardium nuttallii
Conrad, Journal Academy Nat. Sci. Philadelphia, p.
229. pI. 17. f. 3. "1858 Adams & Adams, p. 456:
Cardium (Cerastoderma) nuttallii Conr. "1860
Carpenter: Cardium nuttalli Conr. "1869 ZR: Cardium
nuttallii (Conrad) is distinct from corbis (Martyn) [Ref.
Conrad 1869]. "1871 Stoliczka: Cardium nuttallii
(shows a transition to Pectunculus). "1870 Conrad:
Cardium nuttallii Conrad ... 1872 Dall: Cardium
nuttallii, Conrad (= corbis Desh., Cpr., not Martyn).
"1878 Whiteaves: Cardium nuttallii Reeve. "1884
Grasset: Cardium nuttalli. Conr. "1890 Paetel:
Cardium nuttalli Conr. "1896 Rolle: Cardium nuttallii
Conrad = Cardium californianum Conrad. "1900 Dall:
Cardium (Cerastoderma) nuttallii Conrad, 1837,
section Cerastoderma(jide Schneider 2002). "190 I
Dall: Cardium nuttallii Conrad, 1838 = Cardium corbis
Martyn, 1784. "1922-32 Sherborn, p. 4444: Cardium
nuttallii Conrad, 1837. "1927 Yokoyama: Cardium
nuttallii Conr. [Miocene]. "1930 Stewart: Cardium
nuttallii Conrad is a typical American Cerastoderma.
"1936 Keen: Cardium nuttallii Conrad, 1837. "1936
ZR: Cardium nuttallii Conrad, 1837. Genotype of
Clinocardium Keen, 1936 [Ref. Keen 1936]. "1944
Keen & Bentson: Clinocardium nuttallii (Conrad).
"1953 La Rocque: Clinocardium nuttallii Conrad
1837. "1954 Keen: Cardium nuttallii Conrad, 1837.
[U. Miocene to Recent]. "1963 ZR: C/inocardium
nuttalli, comparative study of base composition of DNA
[Ref. Smith & Quayle 1963]. "1963 ZR: Clinocardium
nuttalli, orientation and fOITn [Ref. Stasek 1963]. "1964
Shikama: Cardium (Clinocardium) nuttallii (Conrad).
"1967 Habe: Clinocardium nuttallii. "1968 ZR:
Clinocardium nuttallii Effect of insecticide Sevin on
survival and growth [Ref. Butler et at. 1968]. "1968
ZR: Clinocardium nuttallii interganglionic regulation
of heart-beat [Ref. Silvey 1968]. "1968 ZR:
Clinocardium nuttallii Notes on biology and
commercial importance, N. American Pacific coast,
[Ref. Amos 1968]. "1871 Stoliczka: Cardium nuttallii.
"1972 ZR: Clinocardium nuttalli [Ref. Evans 1972].
"1975 ZR: Clinocardium nuttalli. Rate of growth.
Seasonal variation. Measurements. Ref. to Ham. /
Clinocardium nuttallii. Associations. Endozoic alga.
Ref. to Cooke. "1973 ZR: Clinocardium nuttallii.
Palaeoecology. Japan. Ref. to Kilmer / Symbiont green
alga. Ref. to Hartman. "1976 ZR: Clinocardium
nuttallii. Parasites. Algal parasites. Northern Pacific.
Ref. to Hartman. "1977 Fischer-Piette: Corculum
(Keenocardium) nuttallii Conrad. "1977 Habe:
Clinocardium (Clinocardium) nuttallii (Conrad, 1937).
"1977 Popov: Cardium nuttallii Conrad, 1837. Type
of Clinocardium Keen, 1936. "1981 ZR: Cerastoderma
nuttallii. California. Ref. to Mcpeak. "1982 ZR:
Clinocardium nuttallii (Conrad 1837). Diagnostic
characters. Ref. to Kafanov et al. p. 43. "1982 ZR:
Clinocardium nuttallii syn. nov. Cardium ?decoratum
Grewingk 1850. Ref. to Kafanov et at. p. 43. "1982
ZR: Clinocardium nuttallii. Miocene. Russian SFSR.
Kamchatka & Sakhalin, first record for Miocene &
range extension. Ref. to Kafanov et al. ··1982 ZR:
Clinocardium nuttallii. Ref. to Brenchley. / Gallucci et
al. / Smith. "1983 ZR: Clinocardium nuttallii. Oregon.
Ref. to Robinson et al. "1985 ZR: Clinocardium
nuttallii. Ref. to Meyhofer. "1987 ZR: Clinocardium
nuttallii. Ref. to Bayer. - Clinocardium nuttalli. Ref.
to Estes et al. "1990 Gladenkov & Sinelnikova:
Clinocardium nuttallii (Conrad), 1837. "1994
Schneider: Clinocardium nuttallii (Conrad, 1837).
"1998 Schneider (1998b): Clinocardium nuttallii
(Conrad, 1837) from Oregon and Washington harbour
Chlorella in the tissues. "1999 Higo et al.:
Clinocardium (Clinocardium) nuttallii (Conrad, 1837).
Type locality: California. Distribution: northeastern
Honshu (Iwate prefecture) and northwards, Hokkaido,
Phuket Marine Biological Center Special Publication vol. 30 (2004)
657
nux - obliquatum
Okhotsk Sea; Bering Sea, Alaska, northwestern USA
to California. Habitat: 10-50 m, sandy mud. --2000
Coan et af.: Clinocardium (Clinocardium) nUl/allii
(Conrad , 1837). --200 I Schneider & Carter:
Clinocardium nuttallii (Conrad , 1837). --2002
Schneider: Clinocardium nuttallii. - CHARACTERS
AND OCCURRENCE - OF COLLECTIONS. --ZMUC wet
collection: Friday Harbor, 15 m muddy silt 1. Hylleberg
leg. DESCRIPTIONS. --1845 Reeve: shell cordately ovate,
Venus-shaped, radiately ribbed, 32 ribs, the extreme
posterior ribs very close and almost obsolete, the rest
rounded, nodulously crenated in an arched manner
throughout; whitish, covered with a rather thick Iivebrown epidermis. --1870 Conrad : 28 large ribs; no
intermediate rib between large ribs; 6 close narrow ribs
on posterior slope. Ligament very wide. Narrow and
prominent beaks. "1896 Rolle : 34 ribs. REMARKS.
--1845 Reeve: this is a noble species though not
exhibiting any attractive display of colours; the ribs
are rounded, and regularly crenated, in a peculiar arched
style, from the margins to the summit of the umbones.
All the specimens that have come under my notice are
curiously characteristic of several distinct epochs of
growth. The first stage of growth is completed within
about the radius of an inch from the umbones, and
others succeed it at rather less intervals throughout.
--1870: Conrad compared C. corbis Meusch or of
Martyn, from Pooloo Condor [Vietnam] with C.
nuttallii Conrad from Upper California. They were
claimed to be synonyms by Carpenter, but Conrad
showed the two to be very different. --1871: Stoliczka
stated that Cardium nUllalli; is a true intermediate
between the genera (subgenera) Acanthocardia Gray,
1853 and Peetuneulus Adans., 1757 = Veprieardium.
"1927 Yokoyama p. 178 : Cardium shinjiense =
Cardium nuttallii (sic) Conr.; they resemble Cardium
Jastosum Yokoyama, 1927. HABITAT. Intertidal, shallow
water. DISTRIBUTION. --1844 Sitka, Reeve: Northwest
USA. --1884 Grassel: Vancouver. "Of authors: North
West Pacific, Bering Sea, California.
nux da Costa, 1778. - Not a cardiid. » Regarding
Cardium nux: RECORDS. --1778 da Costa, p. 173 (pI.
13, fig. 2. 2): Cardium nux. "1803 Montagu: Cardium
nux Da Costa. p. 173 . "1853 Forbes & Hanley:
Cardium nux not a cardiid. "1902 Sherborn, p. 676:
Cardium nux da Costa, 1778. --1977 Fischer-Piette:
Cardium nux da Costa = freshwater species.
nystianum Orbigny, 1852. - Combination to be revised:
"Cardium" nystianum Orbigny, 1852. Cardiidae.
Fossil. RECORDS. --1980 Ruhoff: Cardium nystianum
Orbigny, 1852; [1850-1852] vol. 3: 21.
nystii - Conocardium nystii. Conocardiidae. Fossil.
RECORDS. "1857 Deshayes: Cardium nystii. "1999
Jacques Vidal (pers. com.), MNHN, Paris: Fossils from
the Paleontological Collection as Conoeardium nysti
de Konink. Belgium (Tournai) B06485.
noggerathi (invalid spelling). - See under noeggerathi
J. Mi.iller, 1851.
o
0, Cardium Martyn, 1789. Non binominal. - Not
available. Cardiidae. Recent. RECORDS. --1789 Martyn:
Cardium "0 ". REMARKS. "1936 Keen: Martyn's
Universal Conchologist is not consistently binomial.
Martyn's identification is erroneous and confusion has
been created. The only available name is Clinoeardium
nuttallii Conrad, 1837. DISTRIBUTION . North West
Pacific, Bering Sea, California.
obesa Sowerby, 1899 . - Synonym of Tridacna
(Ffodacna) derasa (Roding, 1798) . Tridacnidae.
Recent. RECORDS. --1899 ZR: Tridaena obesa Sowerby,
1899 n. sp. Philippines. --1999 Higo et af.: Tridacna
obesa Sowerby, 1899 = Tridacna (Ffodaena) derasa
(Roding, 1798) Tridacnidae.
obesum Bartsch, 1947. - Synonym ofCorcufum eardissa
(Linne, 1758). » Regarding Corcufum obesum:
Cardiidae. Recent. TYPE LOCA LITY. Cebu, Philippines.
TYPE REPOSITORY. USNM 543559. RECORDS. "1947
Bartsch, p. 224-225, pI. I, fig. 2: Coreufum obesum
new species. --1947 ZR: Coreufum obesum Bartsch,
1947 sp. nov. Cebu. --1960 Kuroda: Coreufum obesum
Bartsch. --1964 Shikama: Coreufum obesum Bartsch.
--1977 Fischer-Piette: Cardium obesum Bartsch =
Cardium eardissa Linne . --1977 Habe: Coreulum
obesum Bartsch, 1947 = Corell fum eardissa (Linnaeus,
1758). --1989 Severin & Cooper: Corculum obesum.
--1999 Higo et at. : Corculum obesum Bartsch, 1947 =
Corculum cardissum (Linnaeus, 1758).
CHARACTERS AND OCCURRENCE - DESCRIPTIONS. Anterior side much more convex than posterior
side. DIMENSIONS. --1947 Bartsch: height 36. 1 mm;
length 18 mm, diameter 29.9 mm. REMARKS. "1989:
Severin & Cooper found that C. obesum is part of an
ecological continuum of Corculum represented by a
'cardissa'-morph and a 'monostroma'-morph at each
of the two ends.
obliqua (subspecies). - See longiuscula obliqua Eberzin,
1960. - samarangae obliqua Ozaki, 1956.
ob/iqua Ozaki, 1956. - Nemocardium obliqua Ozaki,
1956. Cardiidae. Fossil. RECORDS. --1980 Noda:
Nemocardium obliqua Ozaki, 1956. --1988 Noda:
Nemocardium obliqua.
obliquatal-um (variety). - See under rectidens obliquata.
- papil/osum obliquata Aradas 1846. - papillosum
obliquata Michelotti 1839.
obliquata Gillet, 1970. - Plagiodacna obliquata Gillet,
1970. Lyrnnocardiidae. Miocene. RECORDS. --1970 ZR:
Plagiodacna obliquata Gillet, 1970 sp. nov. 'Monte
dei Corvi' deposit, Ancona deposits, Italy.
obliquata Warren, 1932. - Protocardia obliquata Warren,
1932. Cardiidae. Jurassic . RECORDS. --1933 ZR :
Protocardia obliquata Warren, 1932 sp. n. Jurassic
Alberta.
*obliquatum. - Homonymy to be noticed. Cardium
obliquatum from 2 sources: Michelotti, 1839. - Aradas,
1846.
obliqltatum Aradas 1846. - Synonym of Parvicardium
papillosum (Poli, 1795). » Regarding Cardium
obliquatum: Cardiidae. Recent. DISTRIBUTION. C.
obliquatum: unknown (Nord sieck 1969) ; C.
papi/losum : Europe, North West Africa. RECORDS .
--1878 Monterosato: Cardium obliquatum Arad. =
Cardium papillosum Poli . --1884 Monterosato:
Cardium obliquatum. "1922-32 Sherborn, p. 4461 :
Cardium obliquatum Aradas, 1846. --1969 Nordsieck:
Papillicardium papillosum (Poli, 1791) var. obliquatum
(Monterosato, 1884).
658
Tropical Marine Mollusc Programme (TMMP)
obliquatum - obliquum
obliquatum Michelotti, 1839. - Synonym of Cerastoderma edule incerla (Brn.). » Regarding Cardium
obliquatum: Cardiidae. Fossil. RECORDS. "1899 Sacco:
Cardium obliquatum Michelotti? 1839 = Cerastoderma
edule var. incerla (Bm.). --1922-32 Sherborn, p. 4461:
Cardium obliquatum Michelotti, 1839.
oblique (error). - Printer's mistake for obliqua in Noda
1980.
obliquicosta - Budmania obliquicosta. Lymnocardiidae.
Fossil. RECORDS. --1987 Taktakishvili: Budmania
obliquicosta. --1987 Taktakishvili: Limnocardium
(Budmania) obliquicosta.
obli<Juicostata Marinescu, 1975. New combination: Lymnocardium obliquicostata (Marinescu , 1975).
Lymnocardiidae. Pliocene. RECORDS. "1975 ZR: Limnocardium (Budmania) obliquicostata Marinescu, 1975
sp. nov. Romania, Banal. Ref. to Marinescu, p. 25.
obliqum (error). - Printer's mistake for obliquum in
Glibert & Poe I 1970.
obliquoobsoletum Kolesnikov, 1929. - Combination to
be revised: "Cardium " obliquoobsoletum Kolesnikov,
1929. Cardiidae. Sarmatian. RECORDS. --1929 ZR :
Cardium obliquoobsoletum Kolesnikov, 1929 sp. n.
Sarmatian.
obliquoobsoletum armavirense Kolesnikov, 1929
(variety). - Combination to be revised: "Cardium "
obliquoobsoletum armavirense Kolesnikov, 1929 .
Cardiacea. Sarmatian. RECORDS. --1929 ZR: Cardium
obliquoobsoletum var. armavirense Kolesnikov, 1929
nov. Sarmatian [Ref. Kolesnikov 1929].
*ob/iquum . - Homonymy to be noticed. Cardium
obliquum from 5 sources: Spengler, 1799. - Lamarck,
1805. - Woodward, 1833 . - Andrzejoswky, 1836. Dufour, 1881.
obliquum Andrzejoswky, 1836 . - Replaced by
"Cardium " protractum Eichwald. Combination to be
revised. Cardiidae. Fossil. RECORDS. --1857 Deshayes:
Cardium obliquum Andrzejoswky, 1836 - C.
protractum Eichwald.
obliquum Dufour, 1881. - Replaced by "Cardium "
obliquum Lamarck. Cardiidae . Eocene. RECORDS .
--1886 Cossmann: Cardium obliquum Dufour, 1881 =
Cardium obliquum Lamarck.
obliquum Lamarck, 1805. [Figs. Part 3 p. 920]. Changed combination: Loxocardium obliquum
(Lamarck, 1805). OTHER JUDGEMENTS: Cerastoderma
edule (Linnaeus, 1758). - Loxocardium bouei
(Deshayes, 1858). » Regarding Cardium obliquum:
Cardiidae. Eocene. TYPE LOCALITY. Paris Basin, France.
REMARKS. Anton (1839): stated that Cardium obliquum
Lamarck [fossil] is a synonym of Cardium edule Linne.
Glibert & Poel (1970): stated that Cardium obliqum
[sic = obliquum] Lamarck, 1805 non Spengler, 1799 is
a synonym of Loxocardium bouei (Deshayes, 1858).
RECORDS. "1835 Lamarck: Cardium obliquum Lamk.
"1839 Anton: Cardium obliquum Lamarck. "1844
Potiez & Michaud: Cardium obliquum Lam. fossil of
France. --1857 Deshayes: Cardium obliquum Lamarck,
1809. --1859 Sowerby: C. obliquum = Cardium edule
Linn. --1886 Cossmann: Cardium obliquum Lamarck.
--1887 Fischer: Cardium obliquum Lamarck. --1899
Sacco: Type species Loxocardium obliquum Lamarck.
"1904 Cossmann & Pissarro: Cardium (Loxocardium)
obliquum Lamarck. "1912 Favre: Cardium obliquum
Lamarck. "1922-32 Sherborn, p. 4463: Cardium
obliquum Lamarck, 1805 ... 1936 ZR: Laevicardium
obliquum Vin. & Cout. non Lk. Glibert, 1933 sp. n.
Wemmel Sands, Eocene Belgium. --1950 ZR: Cardium
obliquum Lk. 1805, non Spengler 1799 = Cardium
(Loxocardium) bouei Desh. var. barlonensis Tremlett,
1950 nov. --1970 Glibert& Poel: = Loxocardium bouei
(Deshayes, 1858). "1977 Popov: Loxocardium
obliquum (Lamarck, 1805). --1998 Schneider:
Loxocardium obliquum (Lamarck, 1805). "1999
Jacques Vidal (pers. com.), MNHN, Paris: Fossils from
the Paleontological Collection as Cardium
(Loxocardium) obliquum Lamarck. Lutetien,
Bartonien. France; Turkey; Egypt; i.a. --200 I Schneider
& Carter: Loxocardium obliquum (Lamarck, 1805).
Eocene, France. --2002 Schneider: Loxocardium
obliquum (Lamarck, 1805).
ob/iquum Meek & Worthen, 1865 . - Conocardium
obliquum Meek & Worthen, 1865 . Conocardiidae.
Fossil. REMARKS. Conocardiids have no connection with
the Cardiacea except analogy of form, which does not
offer any point of relationship (Dall 1895, p. 519).
RECORDS. --1980 Ruhoff: Conocardium obliquum Meek
& Worthen, 1865 [1865-1866] ; p. 249.
ob/iquum Spengler, 1799. [Figs. Part 1 p. 93. Part 3 p.
856]. - Synonym of Dinocardium robus/um (Lightfoot,
1786). » Regarding Cardium obliqvum: Cardiidae.
Recent. TYPE LOCALITY. -- Carolina. ZMUC. -- Lister
quoted by Spengler 1799: Caroline lsI. TYPE REPOSITORY.
ZMUC Spengler coIl. RECORDS. --1687 Lister, pI. 328,
fig . 165: Cardium obliqvum (fide Spengler 1799) [not
available] . "1799 Spengler, p. 23-24, No. 12, p. 60,
figs . 3 a,b: Cardium obliqvum Lister. --1853 Forbes &
Hanley: Cardium obliquum . --1856 Wood & Hanley:
C. obliquum Speno Sk. Nat. v. pt. I, p. 23, pI. I, f. 3. =
Cardium ventricosum Brug. --1902 Sherborn, p. 678:
Cardium obliquum Spengler, 1799. --1974 Abbott:
Cardium obliquum Spengler, 1799 = Dinocardium
robustum (Lightfoot, 1786). --1977 Fischer-Piette:
Laevicardium obliquum Spengler. --1999 Knudsen &
Hylleberg: Cardium obliqvum Spengler, 1799. CHARACTERS AND OCCURRENCE - DESCRIPTIONS. "1799 Spengler, fig. 3: pectunculus magnus,
maculatus, intus leviter rufescens. Lister Tab. 328. fig.
165. Shell with triangular outline, posterior edge very
long, nearly straight, anterior edge very short, margin
rounded from anterior to posterior end, 32 ribs,
anteriorly and posteriorly broad, flat and smooth,
centrally rOl!nded, transversely ornamented with scalelike stripes, interstices narrow, umbones pointed, turned
in anterior direction, exterior white, yellow posteriorly,
with scattered red-brown spots, interior bright red,
brown towards the margin. DIMENSIONS. --1799
Spengler: length 83 mm (3 Tom 2 Linier), width 83
rrim (3 Tom 2 Linier) The latter measure must have
been taken diagonally. If the width is measured
perpendicular to length, it is about 66 mm. Thickness
is 60 mm, measured on fig. 3. REMARKS. Cardium
obliquum Spengler, 1799 is very similar to
Dinocardium robus/um (Lightfoot in Solander, 1786).
However, the latter species is a Caribbean species.
Unfortunately Spengler 's specimen (ZMUC) is without
locality. Hence, the reference to Lister (1687) is without
relevance. DISTRIBUTION . Unknown (? Caroline Is!,).
Phuket Marine Biological Center Special Publication vol. 30 (2004)
659
obliquum - oblongum
obliquum Woodward, 1833. - Synonym of
Cerastoderma edule (Linne, 1758). » Regarding C.
obliquum: Cardiidae. Fossil. REMARKS. Forbes (1846)
stated that C. obliquum Woodward = Cardium edule
Linne; first appearance during Red Crag period.
Deshayes (1857) and Dollfus & Dautzenberg (1912)
were of the same opinion. RECORDS ... 1846 Forbes: C.
obliquum Woodward. --1853 Forbes & Hanley:
Cardium obliquum Woodward. --1857 Deshayes:
Cardium obliquum Woodward. -CO edule. --1902
Dollfus & Dautzenberg: Cardium obliquum Woodward,
1833. --1922-32 Sherborn, p. 4463: Cardium obliquum
Woodward, 1833.
obliquum bouei Deshayes (variety). - Upgraded.
Changed combination: Loxocardium obliquum bouei
(Deshayes). TYPE LOCALITY. Paris Basin, France.
Cardiidae. Eocene. RECORDS. --1904 Cossmann &
Pissarro: Cardium (Loxocardium) obliquum var. bouei
Deshayes.
obliqvum Spengler, 1799. - See under obliquum
Spengler, 1799.
oblonga (subspecies and variety). - See under caucasica
oblonga Androussov, 1923. - privum oblonga Eberzin,
1958. - propinqua oblonga Androussov, 1923.
oblonga Sowerby. - Cardiomorpha oblonga, Sowerby.
Cardiacea. Fossil. RECORDS. --1859 Chenu:
Cardiomorpha oblonga Sowerby.
oblongatum Solander in Brander, 1766. - Unknown
species. --1902 Sherborn, p. 679: Cardium oblongatum
So lander in Brander, 1766.
oblongulum Sacco, 1899 (Rov. in schedis). Combination to be revised: "Cardium" oblongulum
Rov. Cardiidae. Miocene. RECORDS. --1899 Sacco:
Cardium oblongulum Rov. --1899 Sacco, p. 38, pI. 9,
fig. 5: Cardium oblongulum Rov. (in schedis).
Described by Sacco.
*oblongum. - Homonymy to be noticed. Cardium
oblongum from 3 sources: Gmelin, 1791. - Brown,
1825. - Alieva, 1967.
oblongum Alieva, 1967. - Combination to be revised:
"Cardium" oblongum Alieva, 1967. Cardiidae. Fossil.
RECORDS. --1967 ZR: Cardium oblongum Alieva, 1967
sp. nov. figs. Akchagil deposits, Naftalan, Azerbaidjan,
SSR.
oblongum Born 1780. [Figs. Part I p. 67]. - RECORDS.
--1780 Born as C. latum [err.]. ID by Gmelin 1791: C.
oblongum mus. Caes. Vind. test. t. 3. f. 8.
oblongum Brown, 1825. - Synonym of Laevicardium
crassum (Gmelin, 1791). » Regarding Cardium
oblongum: Cardiidae. Recent. REcoRDs. --1844 Brown:
Cardium oblongum. --1867 Weinkauff: Cardium
oblongum Brown (non Gmelin) = C. norvegicum
Spengler. --1886 Locard: Cardium oblongum Brown,
1825 (non Chern.) = Cardium norvegicum Spengler,
1792 sic. --1888 Kobelt: Cardium oblongum Brown =
Cardium norvegicum Spengler.
oblongum Bruguiere, 1789 replaces oblongum
Chemnitz, 1782. [Figs. Part 1 p. 76, 169. Part 3 p. 897,
898]. Chemnitz is non binominal. Not available. Laevicardium oblongum (Gmelin, 1791). OTHER
JUDGEMENTS: Cardium sulcatum Lamarck, 1819. Cardium obsoletum Spengler, 1799. - Cardium jlavum
Poli (non Lin.). » Regarding Cardium oblongum:
Cardiidae. Recent. RECORDS. --1782 Chemnitz, Tab. 19
Fig. 190: Cardium oblongum. --1782 Chemnitz. 10 by
Gmelin 1791: Cardium oblongum t. 19. f. 190. --1788
Schrater: Cardium oblongum. [Reference to Chemnitz
(1782) fig. 195]. --1789 Bruguiere: Cardium oblongum
Martini, tom. 6. pag. 195, tab. 19, fig. 190. --1817
Dillwyn: Cardium oblongum Chemnitz. --1819
Lamarck: Cardium oblongum Chemnitz 6. t. 19. fig.
190 = Cardium sulcatum Lamarck, 1819. --1832 Bronn:
Cardium oblongum Chemnitz. --1832 Deshayes:
Cardium oblongum Chemnitz. = Cardium sulcatum
Lamk. --1840 Pfeiffer: Chemnitz Fig. 190. Cardium
oblongum = Cardium sulcatum Lam. 24. --1856 Wood
& Hanley, No. 23: Cardium oblongum Ch. vi. 195, f.
190. --1857 Deshayes: Cardium oblongum Chemnitz.
--1858 Adams & Adams, p. 457: Laevicardium
oblongum, Chemn. --1858 Gray: Cardium oblongum
Chemnitz. --1867 Weinkauff: Cardium oblongum
Chemnitz. --1867 Hidalgo: Cardium oblongum
Chemnitz. --1867 ZR: Cardium oblongum (Chemn.) =
sulcatum (Lam.) (nearly allied, but may be kept distinct
[Ref. Weinkauff 1867). --1869 Appelius: Cardium
oblongum Chemnitz. --1869 Petit de la Saussaye:
Cardium oblongum Chemnitz. --1870 Aradas & Benoit:
Cardium oblongum Chemnitz. --1870 Hidalgo:
Cardium oblongum Chemnitz. --1870 March: c.
oblongum Chemnitz. junior synonym of Cardium
obsoletum Spgl. --1872 Monterosato: Cardium
oblongum Chemnitz. --1878 Monterosato: Cardium
oblongum Chemnitz. = c.jlavum Poli (non Lin.) = C.
sulcatum Lk. --1874 Jeffreys: Cardium oblongum
Chemnitz. --1875 Monterosato: Cardium oblongum
Chemnitz. --1875 Clement: Cardium oblongum
Chemnitz. --1878 Monterosato: Cardium oblongum
Chemnitz. --1879 Granger: Cardium oblongum
Chemnitz. --1880 Stossich: Cardium oblongum
Chemnitz. --1883 Dollfus: Cardium oblongum
Chemnitz. --1883 Marion: Cardium oblongum
Chemnitz. --1884 Grasset: Laevicardium oblongum
Chern. --1884 Monterosato: Laevicardium oblongum
Chemnitz. --1884 repratx: Cardium oblongum
Chemnitz. --1889 Carus: Cardium oblongum Chemnitz.
--1890 Paetel: Laevicardium oblongum Ch. = Cardium
oblongum. --1891 Brusina: Cardium oblongum
Chemnitz. --1891 Dautzenberg: Cardium oblongum
Chemnitz. --1892 Locard: Cardium oblongum
Chemnitz, 1789 sic. --1896 Elera: Cardium oblongum
Chemnitz. --1898 Locard: Cardium oblongum
Chemnitz. --1899 Locard: Cardium oblongum
Chemnitz. --1899 Granger: Cardium oblongum
Chemnitz. --1899 Sacco: Laevicardium oblongum
Chemnitz 1782 [Pliocene-Recent]. --1900 Locard &
Caziot: Cardium oblongum Chemnitz, 1789 sic. --1903
Font y Sague: Laevicardium oblongum Chemnitz.
--1913 Pallary: Cardium oblongum Chemnitz. --1927
Dautzenberg: Cardium (Laevicardium) oblongum
(Chemnitz) Gmelin. --1962 Pasteur-Humbert: Cardium
(Laevicardium) oblongum Chemnitz. --1965 Mars:
Laevicardium oblongum Chemnitz. --1969 Nordsieck:
Laevicardium oblongum (Chemnitz, 1782). --1999
Knudsen & Hylleberg: Spengler, 1799 Cardium
oblongum Chemnitz Tom. VI. Tab. 19. fig. 190. = No.
15 Cardium obsoletum in Spengler (The
Mediterranean). --1999 Jacques Vidal (pers. com.),
MNHN, Paris: Fossils from the Paleontological
660
Tropical Marine Mollusc Programme (TMMP)
oblongum - oblongum comitatensis
Collection. Cardium oblongum Chemnil2. Belgium (An
Vers). B35562.
oblongum Gmelin, 1791.[Figs.Part 1 p . 67,76,101,138
Part 3 p. 896- 98]. - Laevicardium oblongum (Gmelin,
1791). OTHER JUDGEMENTS: Laevicardium attenuatum
(Sowerby). - Cardium crassum Gmelin. - Cardium
laevigatum Linne - Cardium norvegicum Spengler,
1799. - Cardium sulcatum Lamarck. » Regarding
Cardium oblongum : Cardiidae. Pliocene and Recent.
Type species: Stoliczka (1870) designated Cardium
oblongum Gmelin, 1791 type species of genus
Laevicardium Swainson, 1840. - Glibert & Poe I: (1970)
designated Laevicardium oblongum (Gmelin, 1791) as
type species of subgenus (Laevicardium) Stoliczka.
RECORDS. 001780 Born. ID by Gmelin 1791 : Cardium
oblongum mus. Caes. Vind. test. t. 3. f. 8. 001782
Chemnitz. ID by Grnelin 1791: Cardium oblongum t.
19. f.190. 001791 Grnelin, 13,p. 3254, No. 34: Cardium
oblongum. 001798 Roding: Cardium oblongum Gmelin.
001803 Montagu: Cardium oblongum Gme!. 34? =
Cardium laevigatum Lin. 001806 Link: Cardium
oblongum Linne, Gmelin, p. 3254. 001815 Wood:
Cardium oblongum. 001828 Wood: Cardium oblongum .
001829 Costa: Cardium oblongum. Gm. = Cardium
flavum, Poli et Born. 001836 Philippi: Cardium
oblongum Gmelin p. 3254; citing Chemnitz VI. t. 19.
fig. 190 = Cardium sulcatum Lamk. 001840 Sowerby,
catalogue p. 5, No. 64: Cardium oblongum = Cardium
sulcatum Lam. 001844 Potiez & Michaud: Cardium
oblongum Gme!. = Cardium sulcatum Lam. 001845
Reeve, species 71 : Cardium oblongum Linnaeus, Syst.
Nat. (Gmelin's edit.), p. 3254. 00 1859 Sowerby: C.
oblongum = Cardium norvegicum, Spengler. 001866
Brusina: Laevicardium oblongum Gmelin. 001870
Stoliczka: Cardium oblongum Gmelin, 1791. 001890
Paetel: Cardium crassum Gm. = oblongum Ch. [not
available]. 001891 Danilo & Sandri: Cardium oblongum
Gme!. = Cardium sulcatum Lam. 001897 ZR: Cardium
oblongum, note on the siphonal papillae Ref. to Nagel
(1897). 001900 Pallary: Cardium (Laevicardium)
oblongum Gmelin. 001902 Pallary: Cardium
(Laevicardium) oblongum Gmelin. 001902 Sherborn,
p. 679: Cardium oblongum Grnelin, 1790 sic. 001907
ZR : Cardium oblongum [Ref. Zimmermann 1907].
001913 Dautzenberg: Cardium oblongum. 001916
Hidalgo: Cardium oblongum. 001920 Palary: Cardium
oblongum Gmelin. 001933 Coen: Laevicardium
oblongum Gmelin. 001933 Co en & Vatova:
Laevicardium oblongum Gmelin. 001940 Smith:
Cardium oblongum L. = Laevicardium attenuatum
(Sowerby) (cited by Fischer-Piette 1977, p. 26). 001954
ZR: Cardium oblongum, ecological and biological
notes, figs. [Ref. Braun 1954]. 001959 Devidts: Cardium
oblongum Gmelin. 001961 ZR: Laevicardium oblungum
sic; deep sea faunal associations off Provence [Ref.
Picard 1961]. 001963 Riedl: Laevicardium oblongum
Gmelin. 001967 Colantoni: Laevicardium oblongum.
001969 Moore et al.: Laevicardium (Laevicardium)
oblongum (Gmelin, 1791). 001970 Glibert & Poel :
Laevicardium (s.s.) oblongum Gmelin, 1790 sic. 001977
Fischer-Piette: Laevicardium oblongum Gmelin .
001977 Popov p. 50.: Genotype: Cardium oblongum
Gmelin, 1790. Type of Laevicardium Swainson, 1840.
001977 Wilson & Stevenson: Cardium oblongum
Gmelin, 1791. 001983 Anaya: Cardium oblongum
Gmelin, 1791. Type species of Laevicardium Swainson,
1840. Type designated by Stoliczka 1871. 001992
Schneider p. 155: Laevicardium oblongum (Grnelin).
Type species of Laevicardium. 001998 Macedo et al.:
Laevicardium oblongum (Gmelin, 1791).002001 Vidal,
p. 407: Laevicardium oblongum. - CHARACTERS
AND OCCURRENCE - OF COLLECTIONS. ooZMUC wet
collection, West Indies. It is not Laevicardium
oblongum Gmelin. DESCRIPTIONS. 001845 Reeve: shell
ovately oblong, ventricose, somewhat attenuated
towards the umbones, rather thin, radiated with grooved
striae, posterior area smooth; brownish-white, covered
with a yellowish brown epidermis. 002001 Vidal :
tentacles numerous. No eyes on tentacles. REMARKS.
001845 Reeve: Lamarck appears to have omitted all
notice of this species. HABITAT. 001845 Reeve: dredged
from mud at the depth of about twenty fathoms. 001969
Nordsieck: sand at 4 to 200 m depth. DISTRIBUTION.
001845 Reeve: Devonshire, Britain. 001884 Grasset:
Algeria. 001969 Nordsieck: Mediterranean, Canary Is!.
oblongum Linne. - See under Gmelin, 1791.
oblongum of Brown 1845. - Misidentification of
Laevicardium crassum (Gmelin, 1791). Cardiidae.
Recent. RECORDS . 001912 Dautzenberg & Fischer:
Cardium oblongum Brown 1845 (non Chemnitz) =
Cardium (Laevicardium) norvegicum Spengler 1799.
oblongum of Montagu 1803 . - Misidentification of
Parvicardium Jasciatum (Montagu, 1808). Cardiidae.
Recent. RECORDS. 001886 Locard: Cardium oblongum
Montagu, 1803 (non Brug.) = Cardium Jasciatum
Montagu, 1808. 00 I 888 Servain: Cardium oblongum
Montagu 1803 non Bruguiere = Cardium Jasciatum
Montagu, 1808.
oblongum of Reeve 1845. - Misidentification of
Laevicardium crassum (Gmelin, 179 I). OTHER
JUDGEMENTS: Cardium oblongum var. ovale. - Cardium
laevigatum Penn. » Regarding Cardium oblongum:
Cardiidae. Recent. RECORDS. 001856 Wood & Hanley:
C. oblongum Reeve (not. Ch.) I. Car. f. 71 = Cardium
laevigatum Penn. 001868 ZR: Cardium oblongum of
Reeve to be distinguished from the Mediterranean
oblongum (Chemn.) =flavum (Born) =sulcatum (Lam.)
[Ref. Romer 1868]. 001869. Romer: Cardium oblongum
Reeve fig. 71 = Cardium norvegicum Spengler. 001917.
Hidalgo: Cardium oblongum Reeve = Cardium
norvegicum Spengler. 001912. Daul2enberg & Fischer:
Reeve 1845, fig . 71 is C. oblongum var. ovale. 00 I 912.
Daul2enberg & Fischer: Reeve 1.845, fig. 79 represents
the true Cardium oblongum.
oblongum attenuata Locard & Caziot, 1900 (variety).Infrasubspecific. No evidence of subspecific status. «
Regarding Cardium oblongum var. attenuata:
Cardiidae. Recent. TYPE LOCALITY. Mediterranean.
RECORDS. 001900 Locard & Caziot: Cardium oblongum
Chemnil2, 1789 sic var. attenuata.
oblongum comitatensis Fontannes (variety). - No
evidence of subspecific status. OTHER JUDGEMENTS:
Laevicardium crassum Jragile (Brocchi, 1814). »
Regarding Laevicardium oblongum var. comitatensis:
Cardiidae. Recent. RECORDS. 001902 Dollfus &
Dautzenberg: Cardium oblongum var. comitatensis
Fontannes. 001899 Sacco: Laevicardium oblongum var.
comitatensis Fontannes. 001970 Glibert & Poel:
Phuket Marine Biological Center Special Publication vol. 30 (2004)
661
oblongum gibbum - obscurum
Laevicardium oblongum variety comitatensis =
Laevicardium (s.s.) crassumfragile (Brocchi, 1814).
oblongum gibbum Jeffreys (variety). - Infrasubspecific.
No evidence of subspecific status. » Regarding
Laevicardium oblongum var. gibbum: Cardiidae.
Recent. RECORDS. ··1969 Nordsieck: Laevicardium
oblongum var. gibbum (Jeffreys).
oblongum minor Locard & Caziot, 1900 (variety). Infrasubspecific. No evidence of subspecific status. »
Regarding Laevicardium oblongum var. minor:
Cardiidae. Recent. TYPE LOCALITY. The Mediterranean.
DISTRIBUTION. Mediterranean (Locard & Caziot 1900).
RECORDs. ··1900 Locard & Caziot: Cardium oblongum
Chemnitz, 1789 sic var. minor.
oblongum ovale (variety). - Infrasubspecific. No
evidence of subspecific status. » Regarding
Laevicardium oblongum var. ovale: Cardiidae. Recent.
REcORDs. ··1912 Dautzenberg & Fischer: Reeve 1845,
fig. 71 is C. oblongum var. ovale.
oblungum (error). - Printer's mistake for oblongum in
Picard 1961.
obovalis (mode). - See under obovale.
obovale Sowerby, 1833. [Figs. Part I p. 106, 144. Part 3
p. 856,931]. - Changed combination: Trigoniocardia
obovalis (Sowerby, 1833). OTHER JUDGEMENTS: Cardium
exiguum Gmelin. » Regarding Cardium obovale:
Cardiidae. Pliocene and Recent. TYPE SPECIES. Olsson
(1961) designated Trigoniocardia obovalis (Sowerby,
1833) type species of the subgenus (Apiocardia). TYPE
LOCALITY. Hab. ad oras Americae Meridionalis
(Xipixapi) (Sowerby 1833). Xipixapi (Jipijapa),
Ecuador, in II fathoms, sandy mud (Hertlein & Strong
1947). TYPE REPOSITORY. 2 syntypes BMNH London.
Plesiotype [fossil] No. 6069 (C.A.S. type collection)
Hertlein 1934. RECORDS. ··1833 S0:'therby, p. 84:
Cardium obovale. ··1834 Sowerby, 46t part, fig. 4:
Cardium obovale nobis. ··1840 Sowerby, catalogue p.
7, No. 90: Cardium obovale, Sow. ··1840 Sowerby,
corrected list p. 8: Cardium obovale, (Sowerby No. 90).
··1845 Reeve: Cardium obovale Broderip & Sowerby.
"1846 Orbigny: Cardium obovale, Sowerby, 1833.
··1852 Adams: 431. Cardium obovale Brod. et Sowb.
1833. ··1858 Adams & Adams, p. 459: Hemicardia
(Fragum) obovalis, Brod. and Sow. ··1860 Carpenter:
Cardium obovale Brod. & S. ··1884 Grasset: Cardium
obovale. Sow. ··1890 Paetel: Hemicardium obovale
Brod. et Sow. ··1901 Dall: Cardium (Trigoniocardia)
obovale Sowerby, 1833. "1922-32 Sherborn, p. 4478:
Cardium obovale Sowerby, 1833. ··1932 Pilsbry &
Lowe: Cardium (Trigoniocardium) obovale Sby. ··1934
Hertlein, p. 62 (pI. 21, fig. 14): Cardium obovale
Sowerby [fossil]. "1941 Pilsbry & Olsson: Cardium
(Trigoniocardia) obovale Sby. [PI iocene]. ··1944
Smith: Trigoniocardia obovale Sowerby. ··1947
Hertlein & Strong: Cardium (Trigoniocardia) obovale
Sowerby. ··1955 Hertlein & Strong: Cardium
(Trigoniocardia) obovale Sowerby. ··1958. Abbott:
Trigoniocardia obovalis Sowerby 1833 [Recent].
··1961. Olsson: Trigoniocardia obovalis (Broderip &
Sowerby, 1833). ··1961. Olsson: Cardium obovale
Sowerby, 1833. ··1961. ZR: Cardium obovale Sowerby
Sub genotype of (Apiocardia) Olsson, 1961. "1961.
ZR: Trigoniocardia (Apiocardia) obovale subgenotype
of (Apiocardia) Olsson, 1961. ··1964. Parker:
Trigoniocardia (Trigoniocardia) obovalis (Sowerby,
1833). ··1969. Moore et al.: Trigoniocardia
(Apiocardia) obovalis (Sowerby, 1833). "1977. Popov:
Trigoniocardia (Trigoniocardia) obovalis (Sowerby,
1833). "1977. Fischer-Piette: Corculum (Trigoniocardia, Apiocardia) obovale Sowerby. ··1977. FischerPiette, p. 174: Cardium obovale Sowerby = Cardium
exiguum Gmelin. ··1998 Schneider (1998b):
Trigoniocardia (Apiocardia) obovale (Sowerby, 1833).
- CHARACTERS AND OCCURRENCE - OF COLLECTIONS. ··ZMUC: Gulf of California. CHARACTERS of
diagnostic value: sub ovate, somewhat oblique shell
sculptured with about 21 finely noded ribs is typical of
this species. DESCRIPTIONS. ··1845 Reeve: shell oblong,
heart-shaped, oblique, gibbous, sides very short;
radiately ribbed, 22 ribs, side ribs depressly flattened,
anterior of which are finely crenulated on the posterior
side, middle ribs rounded, very closely crenated
throughout, interstices between all the ribs transversely
impressly cancellated; whitish, faintly tinged with
brown towards the umbones. ··1947 Hertlein & Strong:
shell decidedly higher than long; umbo abruptly
angulated posteriorly; posterior margin truncated; ribs
beaded; anterior laterals crowded against cardinals
posteriorly; posterior margin truncated. - Shell
subovate, oblique, fairly convex and peaked,
ornamented by about 2 I finely noded radial ribs, those
anteriorly somewhat broadly and asymmetrically
triangular, those along the posterior umbonal ridge
higher and rounded, those on the posterior area about
8 in number, flatter, narrower and closer together than
those on the anterior area, interspaces transversely
grooved. This is a small white form, distinguished by
the very different outline and more numerous ribs (20
to 22), which are roughened by numerous minute nodes.
Maximum size attained is about 14 mm. Length; 20
mm. height. Magdalena Bay and the Gulf of California
to Salinas, Ecuador. Sometimes the shells are found
on shore but mostly are taken by dredging. REMARKS.
··1845 Reeve: of a very singular oblong form, with a
peculiarity of sculpture, which I have not observed in
any other species. DrMENSIONS. About 20.5 mm in height
and 14 mm in length. HABITAT. ··1845 Reeve: found in
sandy mud at the depth of I I fathoms. ··ZMUC: sand,
shelly sand, 9-104 m. DISTRIBUTION . •• I 845 Reeve:
Xipixapi, West Columbia. ··1884 Grasset: China.
··1964 Parker: Gulf of California.
obscura (mode). - See under obscurum.
obscura Papaianopol, 1981. - Pontalmyra obscura
Papaianopol, 1981. Lymnocardiidae. Upper Miocene.
RECORDS. ··1981 ZR: Pontalmyra (Pontalmyra)
obscura Papaianopol, 1981 sp. nov. Romania. Ref. to
Papaianopol, p. 40.
obscurum Miinster, 1839 - Not valid. » Regarding
Cardium obscurum: Combination to be revised:
"Cardium" obscurum. - ?Cardiidae. Fossil. TYPE
LOCALITY. Germany. RECORDS. ··1857 Deshayes:
Cardium obscurum (Germany). ··1922-32 Sherborn,
p. 4485: Cardium obscurum Muenster, 1839 [no n.].
obscurum Papaianopol, 1982. - Changed combination:
Euxinicardium obscurum (Papaianopol, 1982).
Lymnocardiidae. Pliocene. TYPE LOCALITY. Pontien et
Dacien du Bassin Dacique, Romania. RECORDS. ··1982
Papaianopol, p. 204, fig.: Limnocardium
662
Tropical Marin e Mollusc Programme (TMMP)
obsoleta vindobonense - obunca
(Euxinicardium) obscurum sp. nov. --1987
Taktakishvili: Euxinicardium obscurum Papaian.
--1987 Taktakishvili: Pontalmyra obscura Papaian.
obsoleta I'indobonense - Cerastoderma obsoleta
vindobonense. Cardiidae. Fossil. RECORDS. "1987
Taktakishvili: Cerastoderma (Obsoletiforma) obsoleta
vindobonense.
Obsoletiforma Kolesnikov in Paramonova, 1971 (genus).
- Lymnocardiidae. Tertiary. RECORDS. --1980. Vokes:
Obsoletiforma Kolesnikov in Paramonova, 1971.
Genus pro Obsoletiformes Kolesnikov, 1948.
Subfamily uncertain. "1987 Kojumdgieva p. 12:
Obsoletiforma: three subgenera: Kubanocardium
Muskhelishwili , 1965. - Obsoletiforma Kolesnikov
1948 - Sarmaticardium Bagdasarjan 1978. Cardiidae.
Miocene, Sarmatian. Emended diagnosis. --2002
Schneider: Obsoletiforma. Sarmatian (late Middle
Miocene) brackish-water.
Obsoletiforma (subgenus). - RECORDS. --1977 ZR :
Cerastoderma (Obsoletiforma). Key to species. Ref to
Il'ina , Nevesskaya & Paramonova p. 72. "1987
Kojumdgieva p. 8: (Obsoletiforma) Kolesnikov, 1948.
Cardiidae. Miocene, Sarmatian. Emended diagnosis.
obsoletiforme Kolesnikov, 1929 - Changed combination:
Cerastoderma obsoletiforme (Kolesnikov, 1929).
Cardiidae. Fossil. RECORDS. --1929 ZR: Cardium obsoletiformis Kolesnikov, 1929 sp. n. Sarmatian. --1977
Popov: Cerastoderma (Obsoletiforma) obsoletiforme.
Obsoletiformes Kolesnikov, 1948 (genus). - Invalid.
RECORDS. --1980 Vokes: invalid vernacular see Obso-
letiforma.
*obsoletum. - Homonymy to be noticed. Cardium
oblongum from 2 sources: Spengler, 1799. - Eichwald,
1830.
obsoletum Eichwa1d, 1830. - Changed combination:
Cerastoderma obsoletum (Eichwald, 1830). Cardiidae.
Miocene. RECORDS. --1899 Sacco: Plagiocardium
obsolelum (Eichwald). --1922-32 Sherborn, p. 4497:
Cardium obsoletum Eichwald, 1830. --1931 ZR:
Cardium obsoletum Ref. to Klinger (1931). --1966 ZR:
Cardium obsoletum, from the Sarmatian of the USSR,
taxonomic position [Ref. Paramonova 1966] . "1977
Popov: Cerastoderma (Obsoletiforma) obsolelum
(Eichw.). "1977 ZR: Cerastoderma (Obsoletiforma)
obsolelum (Eichwald) Key to subspecies. Refto Il'ina,
Nevesskaya & Paramonova. p. 59. "1987 Taktakishviii: Cardium obsoletum Eichw. --1999 Jacques Vidal
(pers. com.), MNHN, Paris: Fossils from the Paleontological Collection. Cardium obsolelum Eichwald.
Austria (Bassin de Vienna), B31391.
obsoletum aff. Eichwald, 1830. - RECORDS. --1907 ZR :
Cardium aff. obsolelum [Ref. Bogatchew 1907]. --1987
Taktakishvili: Limnocardium aff. obsoletum Eichw.
obsoletum Spengler, 1799. [Figs. Part I p. 16 No. 190.
Part 3 p. 898] . - Synonym of Laevicardium oblongum
(Gmelin, 1791). » Regarding Cardium obsoletum:
Cardiidae. Recent. TYPE LOCALITY. --1799 Spengler: The
Mediterranean. RECORDS. --1782 Chemnitz, pI. 19, fig.
190: Cardium obsoletum (}ide Spengler 1799). --1799
Spengler, p. 28, No. 15: Cardium obsoletum von Born
p. 47, Tab. 3. fig. 7. --1808 Spengler: Cardium
obsoletum. --1856 Wood & Hanley : C. obsoletum
Speng. Sk. Nat. v. pt. I, p. 28 = Cardium oblongum
Ch. --1870 March: c. oblongum Chemn., synonym of
Cardium obsolelum Spgl. --1902 Sherborn, p. 682:
Cardium obsoletum Spengler, 1799. --1977 FischerPiette, p. 17: Laevicardium oblongum (Gme1in, 1791)
synonym of Cardium obsoletum Spengler, 1799. --1999
Knudsen & Hylleberg: No. 15: Cardium obsoletum in
Spengler 1799 (Comments to sp. No. II) = Cardium
jlavum Born, 1780 Tab. 3, fig. 8. --1999 Knudsen &
Hylleberg: No. 15: Cardium obsoletum in Spengler
1799 = Cardium oblongum Chemnitz Tom. VI. Tab.
19. fig. 190. - CHARACTERS AND OCCURRENCE
- Or COLLECTIONS. --ZMUC: 3 specimens in the
Spengler coil. DESCRIPTIONS. --1799 Spengler: testa
subovata, longitudinaliter obsolete sulcata, margine
crenalo. --1799 Spengler: shell oblong, egg-shaped ,
inflated, very large nymphs, posterior slope occupies
about 25% of the surface, very smooth, ribs low,
decreasing in distinctness towards the anterior end,
where only traces of ribs are seen, umbones narrow,
turned towards the anterior end. Outside pale yellow,
brown around the umbones , inside snow-white.
DIMENSIONS . --1799 Spengler: length 72 mm (2 3/4
Tom .), width 57 mm (2 Tom. 2 Lin.). REMARKS.
--ZMUC : Cardium obsoletum Spengler, 1799 =
Cardium oblongum Gmelin, 1791. DISTRIBUTION. --1856
Wood & Hanley: Mediterranean ... Of authors:
Mediterranean, Canary lsI.
obsoletum of Born, 1780. - Identical with Laevicardium
oblongum (Gmelin, 1791). » Regarding Cardium
obsoletum: Cardiidae: Recent. REMARKS. --1799
Spengler (p. 29) is puzzled why Born has mistaken this
species for c.jlavum Linne; probably because it has a
yellow colour. I don't know why Spengler referred to
Born since Born did not use the name C. obsoletum in
his description. He called it C. jlavum. RECORDS. --1780
Born, p. 47, pI. 3, fig. 7: Cardium obsoletum (}ide
Spengler 1799).
obstinatus Barinov, 1992. - Clinocardium obstinatus
Barinov, 1992. Cardiidae. Neogene. TYPE LOCALITY.
Kamtchatka, Russia. RECORDS. "1992 Barinov et al.,
p. 123, fig.: Clinocardium obstinatus Barinov, 1992;
sp. nov.
obtrita (mode). - See under oblritum Locard, 1886.
obtritum Locard, 1886. - Synonym of Cerastoderma
edule (Linne, 1758). OTHER JUDGEMENTS: Cardium var.
edule. » Regarding Cardium oblritum: Cardiidae.
Recent. HABITAT. Littoral zone (Locard & Caziot 1900).
DISTRIBUTION. Etangs de Biguglia and Diane (Locard
& Caziot 1900); Atlantic to the British Channel
(Nordsieck 1969). RECORDS. "1886 Locard: Cardium
obtritum Locard. --1886 ZR: Cardium obtritum Locard,
1886, n. sp. (= C. edule, pars), France. "1888 Kobelt:
Cardium obtritum Locard = Cardium var. edule. --1888
Servain: Cardium obtritum Locard = Cardium edule
Linnaeus, 1767. --1892 Locard: Cardium obtrilum
Locard, 1886. --1897 Dollfus: Cardium obtritum
Locard (1886). --1900 Locard & Caziot: Cardium
obtritum reference to figs. 14-15, Requien 1848. --1902
Dollfus & Dautzenberg: Cardium oblritum Locard ,
1886 = Cardium (Cerastoderma) edule Linne, 1758.
--1969 Nordsieck: Cerastoderma obtrita (Locard, 1889
sic). --1977 Fischer-Piette: Cardium obtrilum Locard
= Cardium edule Linne.
obunca (subspecies). - See under praetrigonoides
obunca Popov, 1983 .
Phuket Marine Biological Cenler Special Publication vol. 30 (2004)
663
occidentalis - okrugljaki
occidentalis Ebersin, 1962 . - Changed combination:
Pontalmyra occidenlalis (Ebersin, 1962).
oelicum - Combination to be revised: "Cardium"
oelicum. Cardiidae. Miocene. RECORDS. ··1999 Jacques
Lymnocardiidae. Pliocene. RECORDS. ··1962 ZR :
Vidal (pers. com.), MNHN, Paris: Fossils from the
Paleontological Collection. Cardium oelicum [no
author stated]. Aquitanien. France (Gironde), B28623 .
oenanum Gumbel, 1861. - "Cardium" oenanum
Gumbel, 1861. Cardiidae. Fossil. RECORDS. ··1980
Ruhoff: Cardium oenanum Gumbel, 1861; p. 673.
officiosa Papaianopol, 1989. - Pachydacna officiosa
Papaianopol, 1989. Lymnocardiidae. Pliocene. TYPE
LOCALITY. Bassin Dacique, Romania. RECORDS. ··1989
Papaianopol, p. 92, fig.: Pachydacna (Pachydacna)
officiosa sp. nov.
ogurai Otuka, 1938 . - Changed combination:
Vepricardium ogurai (Otuka, 1938) . Cardiidae.
Miocene. RECORDS. ··1938 Otuka: Cardium ogurai.
··1941 ZR: Cardium ogurai Otuka, 1939 sp. n. Miocene
Tyugoku, Japan. ··1949 ZR: Cardium (Bucardium)
ogurai Otuka, 1938 sp. nov. Miocene, Tyugoku, Japan.
··1973 Keen: Vepricardium ogurai (Otuka, 1938).
Miocene.
ohomachii (subspecies). - See under heredium
ohomachii Oinomikado, 1939.
okamotoi Taguchi, 1990. - Vepricardium okamotoi
Taguchi , 1990 . Cardiidae. Middle Miocene . TYPE
LOCALJTY. Miocene Katsuta group in Okayama
Prefecture, Japan. RECORDS. ··1990 Taguchi, p. 609,
fig. : Vepricardium (Vepricardium) okamotoi sp. nov.
okeziei Adegoke, 1977. - Cardium okeziei. Cardiidae.
Palaeocene. RECORDS. ··1977 ZR: Cardium (Cardium)
okeziei sp. nov. Nigeria, Palaeocene. Ref. to Adegoke
p. 271.
okinawense Kuroda, 1960. [Figs. Part 3 p. 867]. Synonym of Vasticardium elongatum (Bruguiere,
1789). OTHER JUDGEMENTS: Cardium enode Sowerby.
» Regarding Vasticardium okinawense: Cardiidae.
Recent. TYPE LOCALITY. Amami Islands, Japan. TYPE
REPOSITORY. Osaka City Museum (Natural History); not
available for loan (Vidal 1993. RECORDS. ··1951 Habe:
Vasticardium serricostatum okinawense Kuroda M. S.
··1955 Kira: Vasticardium serricoslatum Kuroda M.
S. ··1959 Kira: Vasticardium var. serricostatum Melvill
& Standen. ··1960 Kuroda : Vasticardium
(Acrosterigma) okinawense. ··1961 ZR: Trachycardium (Acrosterigma) [serricostatum Melvill &
Standen var. ?] okinawense Kuroda (nov.) Okinawa Is.
[Ref. Kuroda 1961] . ··1962 Kira : Vasticardium
(Acrosterigma) okinawense. ··1977 Fischer-Piette:
Cardium okinawense Kuroda = Cardium enode
Sowerby. ··1990 Bieler & Petit : Trachycardium
(Acrosterigma)serricostatum Melvill & Standen var.?
okinawaense Kuroda nov. ··1993 Vidal: Trachycardium
(Acrosterigma) okinawense Kuroda, 1960 =
Acrosterigma elongatum (Bruguiere, 1789). ··1999
Vidal: Trachycardium okinawaense Kuroda, 1960 =
Vasticardium elongatum (Bruguiere 1789). CHARACTERS AND OCCURRENCE - DISTRIBUTION. ··1994 Hylleberg: Japan, Australia.
okrugici Brusina. - New combination: Lymnocardium
okrugici (Brusina). Lymnocardiidae. Fossil. RECORDS.
··1987 Taktakishvili : Limnocardium okrugici Brus.
okrugljaki Basch, 1990. - New combination :
Lymnocardium
okrugljaki
Basch,
1990.
Lymnocardiidae. Miocene. TYPE LOCALITY. Yugoslavia.
Didacna (Ponlalmyra) occidentalis Eberzin, 1962 sp.
nov. Kerchenskii, Azov region; Pliocene, Russia .
··1987 Taktakishvili : Pontalmyra occidentalis (Ebers.).
ochetophorum Brusina, 1902. - Changed combination:
Euxinicardium ochetophorum (Brusina, 1902). Lymnocardiidae. Tertiary. RECORDS. ··1987 Anaya : Limnocardium ochetophorum Brusina, 1902. ··1987 Taktakishvili: Limnocardium ochetophonlm (Brus.). ··1987
Taktakishvili: Euxinicardium ochetophorum (Brus.)
ochilleallum Stephenson, 1956. - Trachycardium
ochilleanum (Stephenson , 1956). Cardi idae .
Cretaceous. RECORDS. ··1956 ZR: Combination to be
revised: "Cardium" (Trachycardium) ochilleanum
Stephenson, 1956 sp. nov. Cretaceous. Ochillee Creek,
Georgia, U.S.A.
ochotense (subspecies). - See under cilia tum ochotense
Kafanov, 1981.
ochotellse Ilyina, 1963. - Changed combination: Serripes
ochotensis (Ilyina, 1963). Cardiidae. Middle Pliocene.
TYPE LOCALITY. Kamtchatka. REMARKS. The species C.
ochotense is close to Clinocardium ciliatum (Fabricius,
1780). RECORDS. ··1973 Keen: ?Serripes ochotensis
I1yina , 1963. [Kamtchatka). ··1981 Kafanov :
Clinocardium ochotense. ··1990 Gladenkov &
Sinelnikova: Sempes ochotensis Ilyina.
ochoterellai Muellerried, 1933. - Combination to be
revised: "Cardium" ochoterenai Muellerried, 1933.
Cardiidae. Cretaceous. RECORDS. ··1933 ZR: Cardium
ochoterenai Muellerried, 1933 sp. n . Cretaceous
Mexico. ··1934 sic ZR: Cardium ochoterenai
Muellerried, 1933 sp. n. Cretaceous Mexico.
*odessae. - Homonymy to be noticed. Cardium odessae
from 2 sources: Brab. (?year). - Capellini (?year).
odessae Brab . - New combination: Lymnocardium
odessae (Brab.). Lymnocardiidae . Fossil. RECORDS.
··1987 Taktakishvili: Cardium odessae . ··1987
Taktakishvili: Limnocardium odessae (Brab.).
odessae Capellini. - Synonym of Cerastoderma edule
(Linnaeus, 1758). » Regarding C. odessae: Cardiidae.
Fossil. RECORDS. ··1902 Dollfus & Dautzenberg:
Cardium odessae Capellini = Cardium (Cerastoderma)
edule Linne, 1758.
odischiellsis Eberzin, 1960. - Pachydacna odischiensis
Eberzin, 1960. Lymnocardiidae. Pliocene. RECORDS.
··1960 ZR: Pachydacna odischiensis Eberzin, 1960 sp.
n. Pliocene Nodzhikhevi, Mingrelli, Western Georgia,
USSR.
oedalium (subspecies). - See under muricatum oedalium
Dall, 1900.
oedalium Dall, 1900. - Changed combination:
Trachycardium oedalium (Dall; 1900). Cardiidae.
Fossil. RECORDS. ··1953 Olsson & Harbison: Cardium
(Trachycardium) oedalium Dall, 1900. "1977 Popov:
Trachycardium (Dallocardia) oedalium (Dall, 1900).
oedalium harveyense Mansfield, 1932 (subspecies). Combination to be revised: "Cardium" oedalium
harveyense Mansfield, 1932. Cardiidae. Miocene.
RECORDS. ··1938 ZR: Cardium (Trachycardium)
oedalium harveyense Mansfield, 1932 subsp. n.
Harvey's Creek, Leon Co. Choctawhatchee formation
(Miocene), Florida.
664
Tropical Marine Mollusc Programme (TMMP)
olchovensis - omanense
RECORDS. "1990 Basch, p. 30, fig.: Limnocardium
(Zagrabicardium) okrugljaki sp. nov.
olchovensis Petrov, 1982. - Cilia/ocardium olchovensis
Petrov, 1982. Cardiidae. Pliocene/Pleistocene.
RECORDS . ··1982 ZR : Cilia/ocardium olchovensis
Petrov, 1982 sp. nov. Russian SFSR, Ref. to Petrov, p.
104.
oldroydi Weaver and Palmer, 1922. - Combination to
be revised: "Cardium" oldroydi Weaver and Palmer,
1922. Cardiidae. Fossil. RECORDS. ··1926 ZR: Cardium
oldroydi Weaver & Palmer, 1926 sp. n. Eocene U.S .A.
··1930 Stewart: Cardium oldroydi Weaver and Palmer,
1922.
olea Vokes, 1946. - Pr%cardia olea Vokes, 1946.
Cardiidae. Cretaceous. RECORDS. ··1946 ZR:
Pr%cardia olea Vokes, 1946 sp. nov. Cretaceous
(Aptian) Lebanon.
olequahensis (cf.) Weaver. - Combination to be revised:
"Cardium" cf. olequahensis Weaver. Cardiidae. Fossil.
RECORDS. ··1915 Dickerson: Cardium cf. olequahensis
Weaver.
oligocenica I-um (variety). - See under semistria/um
oligocenica Sacco, 1899. - grateloupi oligocenicum
Noszki, 1936.
oligoparvum Sacco, 1899. Loxocardium oligoparvum
Sacco, 1899. Cardiidae. Fossil. RECORDS. ··1899 Sacco,
p. 47, pI. II , fig . 23 : Loxocardium oligoparvum Sacco
olisiponense Sharpe, 1850. - Combination to be revised:
"Cardium " olisiponense. Cardiidae . Cretaceous.
RECORDS. ··1871 Stoliczka: Cardium olisiponense
(Acanthocardium sic). ··1922-32 Sherborn, p. 4558:
Cardium olisiponense Sharpe, 1850. ··1980 Ruhoff:
Cardium olisiponense Sharpe, 1850; vol. 6: 181, pI.
14, figs. 4 a,b.
olivetum Papaianopol, 1982. - Changed combination:
Euxini cardium olivetum (Papaianopol, 1982).
Lymnocardiidae. Pliocene. TYPE LOCALITY. Ponti en et
Dacien du Bassin Dacique, Romania. RECORDS. ··1982
Papaianopol, p . 202 , fig.: Limnocardium
(Euxinicardium) olivetum sp. nov. ··1987 Taktakishvili:
Euxinicardium olivetum Papa ian.
olivifer Iredale, 1936. [Figs, Part 3 p. 864]. - Synonym
of Vasticardium vertebra tum (Jonas, 1844). OTHER
JUDGEMENTS: Laevicardiumjlavum (Lilmaeus, 1758).Acrosterigma reeveanum (Dunker, 1852). ?Trachycardium rugosum (Lamarck). » Regarding
Regozara olivifer: Cardiidae. Recent. GENOTYPE. Iredale
(i 936) designated Regozara olivifer as type species of
the genus Regozara Iredale, 1936. TYPE LOCALITY.
Australia. TYPE REPOSITORY . Australian Museum,
Sydney. Holotype C060606. RECORDS. ··1936 IredaIe,
p. 275 (pI. 20, fig. 8): Regozara olivifer sp. nov. ··1936
Iredale: Regozara, gen. nov. ··1936 Iredale designated
olivifer as type species of genus Regozara Iredale.
··1950 Allan : Regozara olivifer (southern form of
jlavum). ··1952 Kuroda & Habe: olivifer Iredale = ?
Trachycardium rugosum (Lamarck). ··1962 Iredale &
MacMichael: Cardium olivifer Iredale. ··1969 Moore
et al.: Acrosterigma (Regozara) olivifer (Iredale, ) 936).
··1977 Fischer- Piette, p. 31 : Regozara olivifer Iredale
= Laevicardiumjlavum Linne, auctorum. ··1977 Wilson
& Stevenson: Regozara olivifer Iredale, 1936; junior
synonym of Acrosterigma reeveanum (Dunker, 1852).
··1978 ZR: Regozara olivifer Iredale, 1936 junior
synonym of Acrosterigma reeveanum (Dunker, 1852).
Ref. to Wilson et af. p. 82. ··1980 Keen: Acrosterigma
(Regozara) olivifer (lredale) . •• ) 992 Lamprell &
Whitehead : Regozara olivifer Iredale, 1936 =
Acroslerigma reeveanum (Dunker, ) 852). ··1997 Vidal :
Regozara olivifer Iredale, 1936 = Vasticardium
vertebra tum (Jonas, 1844). ··1999 Vidal: Regozara
olivifer Iredale, 1936 = Vaslicardium vertebra tum
(Jonas, 1844). - CHARACTERS AND OCCURRENCE - REMARKS: Cardium j7.avum Linne and
Cardium rugosa Lamarck are closely related to
Regozara olivifer. DISTRIBUTION. ··1994 Hylleberg:
Indo-Pacific.
olssoni Marwick, 1944. Hedecardium olssoni Marwick,
1944. Cardiidae. Tertiary. TYPE LOCALITY. Southland,
N .Z . RECORDS. ··1944 ZR: Hedecardium olssoni
Marwick, 1944 sp. nov. Shell Gully, Kelly's Farm,
Chattoa (Duntroonian). ··1977 Popov: Hedecardium
olssoni Marwick, 1914. ··1978 Maxwell: Hedecardium
(Hedecardium) olssoni Marwick , 1944. ··2002
Schneider: Hedecardium olssoni Marwick, 1944 New
Zealand. Chattian.
olteniae lon.-Arget. - Changed combination :
Tauricardium olteniae (Ion.-Arget). Lymnocardiidae.
Fossil. RECORDS. ··1987 Taktakishvili: Cardium
olteniae Ion .-Arget. ··1987 Taktakishvili:
Tauricardium olteniae.
olteniae Motas, 1974. - Prosodacna olteniae Motas,
1974. Lymnocardiidae. Pliocene. RECORDS. ··1974 ZR:
Prosodacna (Prosodacna) ol/eniae Motas, 1974 sp.
nov. Romania, Bengesti, Valea Mare. Ref. to Motas, p.
94.
oltinae Pana & Andreescu, 1980. - Fischeridacna oltinae
Pana & Andreescu, 1980. Lymnocardiidae . Lower
Pliocene. RECORDS. ··1980 ZR: Fischeridacna oltinae
Pana & Andreescu, 1980 sp. nov. Romania. Ref. to
Andreescu, p. 58.
omagariense Honda, 1981. Clinocardium omagariense
Honda, 1981. Cardiidae. Middle Oligocene. RECORDS.
··1981 ZR: Clinocardium omagariense sp. nov. Japan,
Middle Oligocene. Ref. to Honda, p. 130.
omanense Melvill in Melvill & Standen, 1906. [Figs.
Part 3 p. 915]. - Changed combination : Papillicardium
omanense (Melvill, 1906). Cardiidae. Recent. TYPE
LOCALITY. Persian Gulf, Gulf of Oman (Melvill &
Standen, 1906). RECORDS. ··1906 Melvill, p. 838, pI.
53, fig. 5: Cardium (Acanthocardia) omanense n. sp.
··1906 ZR: Cardium omanense Mel viil & Standen,
1906 sp. n. G. of Oman. ··1977 Fischer-Piette:
Cardium (Cerastoderma, Acanthocardia) omanense
Melvill. ··1995 Oliver: Papillicardium omanense
(Melvill & Standen, 1907). - CHARACTERS AND
OCCURRENCE - DESCRIPTIONS. ··1995 Oliver:
roundly subquadrate, posterior margin a little longer
and straighter than anterior. Sculpture of 25 radiili ribs
all but posterior few bearing large rounded nodules,
these widely spaced and not the full width of the ribs,
posterior ribs with sharper scales. White. DIMENSIONS.
··1995 Oliver: to 5 mm. REMARKS. ··1906 Melvill &
Standen: a small species, with ribs coarse
proportionately speaking to the size of the shell, each.
rib being ornamented with distinct scaly nodules, these
not being in the least imbricate. Only imperfect
specimens, but presumably adult, occurred. HABJTAT.
Phukel Marine Biological Cenler Special Publication vol. 30 (2004)
665
omnifera - orbicuLare
"1906 Me1vill & Standen: 156 fathoms, shell-sand.
"1995 Oliver: offshore. DISTRIBUTION. Oman.
omnifera Papaianopol, 1981 . - Pontalmyra omnifera
Papaianopol, 1981. Lymnocardiidae. Upper Miocene.
RECORDS. 001981 ZR: Pontalmyra (Pontalmyra)
omnifera Papaianopo1, 1981 sp. nov. Romania. Ref. to
Papaianopol, p. 33. 001987 Taktakishvili : Pontalmyra
omnifera Papaian.
omnivaga Papaianopol, 1978. - Paradacna omnivaga
Papaianopol, 1978. Lymnocardiidae. Pliocene.
RECORDS. 001978 ZR: Paradacna omnivaga
Papaianopo1, 1978 sp. nov. Romania. Ref. to Papaianopol, I. p. 285.
omnivagus Ebersin, 1967. - Pseudocatillus omnivagus
Ebersin, 1967. Lymnocardiidae. Pliocene. RECORDS.
001967 Ebersin: Pseudocatillus omnivagus sp. n. 001967
ZR: Pseudocatillus omnivagus Eberzin, 1967 sp. nov.
Pliocene, USSR. 001982 ZR: Pseudocatillus omnivagus
Ebersin, 1967. Description. Ref. to Papaianopol et at.
onachitense Harris, 1919. - Combination to be revised:
"Cardium " onachilense Harris, 1919. Cardiidae.
Eocene. RECORDS. 001919 ZR: Cardium onachilense
Harris, 1919 sp. n. Eocene U.S.A.
oneroaense (mode) . - See under oneroaensis.
oneroaensis Powell, 1938. - Maoricardium oneroaense
(Powell, 1938). Cardiidae. Fossil. RECORDs. 001939 ZR:
Cardium oneroaensis Powell, 1938 sp. n. Tertiary near
Oneroa, Waikeke Island, Auckland. 001977 Popov:
Maoricardium oneroaense (Powell, 1938).
onestae McLearn, 1931 . - Laevicardium onestae
McLearn, 1931. Cardiidae. Cretaceous. RECORDS.
001932 ZR: Laevicardium onestae McLearn, 1932 sp.
n. Cretaceous Canada. 001980 ZR: Onestia onestae
(McLearn, 1931). Ref. to Day. 001995 Schneider:
Onestia onestae (McLearn, 1931): GSC 6345
(ho10type).
Onestia McLearn, 1933 (genus). - Cardiidae. Fossil.
RECORDS. 001980 Vokes: Onestia McLearn, 1933 in
subfamily Protocardiinae . 001980 ZR: Onestia is
referred to Lahillidae from Cardiidae. Ref. to Day, p.
40. 001980 ZR: Onestia Emended diagnosis. Lower
Cretaceous Alberta & Queensland Aptian & Albian.
Ref. to Day.
Onestia McLearn, 1933 (subgenus). - Cardiidae. Fossil.
REcORDs. 001934 ZR: Onestia McLearn, 1933 subgen.
n. of lntegricardium RoIlier, type Laevicardium onestae
McLearn [Ref. McLearn 1933].001970 ZR: Onestia
etheridgei (Etheridge, 1892) first adequate description
[Ref. Fleming 1970].
onoi Hayami, 1959. - Prolocardia onoi Hayami, 1959.
Cardiidae. Lower Liassic. RECORDS. 001960 ZR:
Protocardia onoi Hayami, 1960 sp. nov. Lias,
Higashinagano Japan. 001973 Keen: Prolocardia onoi
Hayami, 1959. Lower Liassic. lncertae sedis.
onusta Papaianopol, 1992 . - Pachydacna onusla
Papaianopol, 1992. Lymnocardiidae. Pliocene. TYPE
LOCALITY. Dacien inferieur (Getien) du Bassin Dacique,
Romania. RECORDS . 00 [992 Papaianopol, p. 23, fig .:
Pachydacna (Pachydacna) onusta sp. nov.
onustum Papaianopol, 1982. - Changed combination:
Euxinicardium onuslum (Papaianopol, 1982) .
Lymnocardiidae. Pliocene. TYPE LOCALITY. Ponti en et
Dacien du Bassin Dacique, Romania. RECORDS. 001982
Papaianopol, p. 201, fig.: Limnocardium
(Euxinicardium) onuslum sp. nov. 001987 Taktakishvili:
Euxinicardium onus tum Papaian.
Onychocardium Whitfield, 1905 (genus). - Cardiacea.
Fossil. RECORDS. 001905 ZR: Onychocardium Whitfield,
[905 n. gen. for Onychocardium porllandicum Whitfield, 1905 n. sp. Portage Rocks of Lake Erie Whitfield.
Opisocardium Bayle, 1879 (genus). - Synonym of
Lunulicardia Gray, 1853.» Regarding Opisocardium:
Cardiidae. Recent. RECORDS. 001879 ZR: Opisocardium,
substituted for Lunulicardium (Gray, 1847), type
Cardium retusum (L.), nec Lunulicardium (Munster,
1840) [Ref. Bayle [879]. 00 1980 Vokes: Opisocardium
Bayle, 1879 [unnecessary n.n. pro Lunulicardia Gray,
1853, not pre-occ. by Lunulacardium Muenster, 1840
(Lunulacardiidae)]. 001999 Higo et al.: Opisocardium
Bayle, 1879 = Lunulicardia Gray, 1853.
Opisocardium Bayle, 1879 (section and subgenus). Cardiidae. Recent. REMARKS. The designation "section"
takes the value of subgenus. RECORDS. 001887 Fischer:
Opisocardium Bayle, 1879. Section of subgenus
Hemicardia Klein, 1755.
oppenheimi Cuvillier, 1935 . - Combination to be
revised : "Cardium" oppenheimi Cuvillier, 1935 .
Cardiidae. Lower Eocene . RECORDS . 001935 ZR :
Cardium oppenheimi Cuvillier, 1935 sp. n. Bartonian
(Eocene) Egypt. 001999 Jacques Vidal (pers. com.),
MNHN, Paris : Fossils from the Paleontological
Collection. Cardium oppenheimi Cuvillier. Bartonien.
Egypt, B43632.
oraphense Davidasvili, 1930. - Oraphocardium
oraphense (Davidasvili, 1930). Lymnocardiidae. Fossil.
RECORDS . 001932 ZR: Phyllicardium oraphense
Davidasvili, 1930 sp. n. Pliocene Transcaucasia. 001977
Popov: Oraphocardium oraphense. 001987 TaktakishviIi: Oraphocardium oraphense (Dav.).
oraphellse Taktakishvili, 1970. - Arcicardium oraphense
Taktakishvili, 1970. Lymnocardiidae. Pliocene.
RECORDS . 001970 ZR: Arcicardium oraphense
Taktakishvili, 1970 sp. nov. Pliocene Upper Cimmerian,
Orapo River Makharadze district, Georgia, USSR.
Oraphocardium Ebersin, 1949 (genus) .
Lymnocardiidae . Pliocene. RECORDS . 001962 ZR :
Oraphocardium Ebersin in subfam. Didacninae
Eberzin, 1962, family Cardiidae [Ref. Eberzin 1962].
001977 Popov: Oraphocardium Ebersin, [949. 001980
Vokes: Oraphocardium Ebersin, 1949 in subfamily
Didacninae . 001987 Taktakishvili: Oraphocardium
Ebersin, 1949. 00200 [ Schneider & Carter:
Oraphocardium (genus). Lymnocardiidae.
*orbicuLare. - Homonymy to be noticed. Cardium
orbiculare from 3 sources: Forbes, 1846. - Schafhautl,
[852. - d'Acosta (?year).
orbicuLare d' Acosta. - Combination to be revised:
"Cardium" orbiculare d' Acosta. Cardiidae. Fossil.
REMARKS. Chemnitz (1782) is not available. RECORDS.
00[788 Schroter [not available]: Cardium orbiculare
[Reference to Chemnitz 1782 fig. 166. Not available] .
001789 Bruguiere: Cardium orbiculare d' Acosta, pag.
[76, tab. [4, fig. 2.
orbicuLare Forbes, [846. - Cardita orbiculare Forbes,
1846. Not in Cardiidae. Fossil. RECORDS. 001922-32
Sherborn, p. 4596: Cardita orbicularis Forbes, 1846.
001922-32 Sherborn , p . 4596: Cardium (Forbes)
orbiculare d'Orbigny, 1850. - Cardita.
666
Tropical Marine Mollusc Programme (TMMP)
orbiculare - orbita
orbiculare M'Coy in Griffith, 1844. - Nomen nudum.
Not available. Fossil. REcoRDs. ··1922-32 Sherborn,
p. 4596: Cardium orbiculare M'Coy in Griffith, 1842
[n.n.].
orbiculare Orbigny, 1850. -See under orbiculare Forbes,
1846.
orbiculare Schatbiiutl, 1852. Combination to be revised:
"Cardium" orbiculare Schatbiiuti, 1852. Cardiidae.
Fossil. REcoRDs. ··1980 Ruhoff: Cardium orbiculare
Schafhiiutl, 1852; p. 159, 168, pI. 4, figs. 7a,b . .. 1980
Ruhoff: Cardium orbiculare Schatbiiutl, 1863; p. 166,
pI. 41, fig. 3; pI. 65a, fig. 8.
orbiculata Kvaliashvili, 1978. - Digressodacna
orbiculata K valiashvili, 1978. Lymnocardiidae.
Pliocene. RECORDS. ··1978 ZR: Digressodacna
orbiculata K valiashvili, 1978 sp. nov. Georgia SSR.
Ref. to Kvaliashvili p. 86.
orbiculata Papaianopol, 1989. - Pachydacna orbiculata
Papaianopol, 1989. Lymnocardiidae. Pliocene. TYPE
LOCALITY. Bassin Dacique, Romania. RECORDS ... 1989
Papaianopol, p. 94, fig.: Pachydacna (Pachydacna)
orbiculata sp. nov.
orbiculo Day, 1980. - Onestia orbiculo Day, 1980.
Cardiidae. Lower Cretaceous. RECORDS. ··1980 ZR:
Onestia orbiculo Day, 1980 sp. nov. Queensland. Ref.
to Day, p. 39.
orbignyanum d ' Archiac, 1846. - Combination to be
revised: "Cardium" orbignyanum d'Archiac, 1846.
Cardiidae. Fossil. RECORDS. ··1922-32 Sherborn, p.
4600: Cardium orbignyanum d'Archiac, 1846
Orbis Blainville, 1825 non Miiller, 1767 (genus). Objective synonym. » Regarding Orbis: Cardiidae.
RECORDS. ··1969 Moore et al.: Orb is = Acanthocardia
Gray, 1851. ··1980 Vokes: Orbis Blainville, 1825, [non
Miiller, 1767 (Pisces), etc. = Acanthocardia Gray,
1851].
orbita Broderip & Sowerby, 1833. [Figs. Part 1 p. 108,
140, 173. Part 3 p. 866]. - Changed combination:
Vasticardium orbita (Broderip & Sowerby, 1833).
Cardiidae. Recent. TYPE LOCALITY. lsI. Annaa, Pacific
Ocean (Sowerby 1841). RECORDS ... 1833 Broderip &
Sowerby, p. 85: Cardium orbita. ··1834 Sowerby, 48 th
part, fig. 13: Cardium orbita nobis. ··1840 Sowerby,
catalogue p. 4, No. 53: Cardium orbita Brod. & Sow.
··1840 Sowerby, corrected list, p. 8: Cardium orbita,
(Sowerby No. 53) ... 1841 Reeve: Cardium orbita
Sowerby. ··1845 Reeve, sp. 85: Cardium orbita Brod.
& Sow. ··1856 Hanley: Cardium orbita Broderip &
Sowerby. "1858 Adams & Adams, p. 455: Cardium
(lsocardia) orbita, Brod. and Sow. · ··1869 Romer:
Cardium orbita Sowerby. "1871 Stoliczka: Cardium
orbita is a Trachycardium. ··1872 Tryon: Cardium
orbita Sowerby... 1890 Paetel: Cardium orbita Sow.
.. 1901 Dall: Cardium orbita Sowerby, 1833 (appears
to be an Indo-Pacific species). ··1906 Lamy: Cardium
orbita Sowerby. ··1920 Pilsbry: Cardium orbita B. and
S. ··1922-32 Sherborn, p. 4604: Cardium orbita
Sowerby, 1833. ··1923 Dautzenberg: Cardium orbita
Rve. ··1926 Pallary: Cardium orbita Brod. & Sow. =
Cardium enode Sowerby in Fischer-Piette (1977).
··1929 Dautzenberg: Cardium orbita Rve. ··1932
Dautzenberg: Cardium orbita Sowerby. ··1933
Dautzenberg & Rouge: Cardium orbita Broderip &
Sowerby. ··1935 Oestergaard: Cardium orbita Broderip
& Sowerby. ··1955 Rost & Soot- Ryen: Trachycardium
orbita Sowerby, 1833. ··1956 ZR: Trachycardium
orbita Swby., new to Seychelles [Ref. Rost & SootRyen 1956]. ··1977 Fischer-Piette: Cardium orbita
Broderip & Sowerby, 1833 = Cardium jlavum Linne
auctorum. ··1977 Wilson & Stevenson: Cardium orbita
Broderip and Sowerby, 1833. ··1979 Kay:
Trachycardium orbita (Sowerby, 1833) ... 1979 Kay:
Cardium orbiter sic Reeve, Bryan, 1915 Gun. syn.) =
Trachycardium orbita (Sowerby, 1833). ··1990 Abbott
& Dance: Trachycardium orbila. ··1990 Wells et al.:
Trachycardium orbita (Sowerby, 1832) ... 1997 Vidal:
Vasticardium orbita orbita (Broderip & Sowerby,
1833). ··1997 Vidal: Cardium orbita Reeve sensu
Dautzenberg, 1929 (not Cardium orbila Broderip &
Sowerby, 1833) = Vasticardium luteomarginatum
insulare Vidal, 1997. ··1997 Vidal: Trachycardium
orbita (Sowerby): Wells et al. 1990 = Vasticardium
orbita philippinense (Hedley, 1899). ··1998 Poutiers:
Trachycardium orbita (Sowerby, 1833). ··1998
Schneider: Cardium orbita is a Trachycardium.
(Stoliczka 1871, according to Schneider 1998). ··1999
Vidal: Vasticardium orbita (Broderip & Sowerby,
1833). ··1999 Vidal , p. 357: Vasticardium orbita
orbita(Broderip & Sowerby, 1833). - CHARACTERS
AND OCCURRENCE - CHARACTERS of diagnostic
value: foundation of anterior lateral teeth is strong and
angular (hooked appearance) in orbita and its
subspecies. DESCRIPTIONS ... 1845 Reeve: shell oval,
gibbous, slightly angulated on the posterior side;
radiate1y ribbed, 42 ribs, of which the anterior are
rounded and crenated, the middle are obtusely scaled
on each side, the scales being small, alternate, and
oblique, and the posterior are scaled on the posterior
side, the scales being larger, but still short and obscure;
interstices between the ribs, especially those of the
middle ribs, deeply excavated or rutted; whitish, palely
variegated with fulvous yellow or purple, interior white,
margins richly stained with purple. ··1979 Kay: shell
ovate, solid, equivalve, inflated; beaks full and
prominent; with flatted spinous ribs; margins fluted and
denticulated; flesh-colored maculated with brown.
Hinge: broad and strong with a strong cardinal in each
valve; left anterior and right posterior cardinals well
developed, the left posterior and right anterior cardinals
obsolete; laterals strongly developed and projecting.
Sculpture: about 42 large, high, flattened ribs separated
by narrow, deep interspaces; ribs imbricated with
distantly spaced spinous scales. Color: exterior flesh,
tinged with spots of chestnut brown, edge becoming
purplish; interior with a purple border and irregular
purple zones within. ··1979 Kay: maximum
dimensions, length 53, width 65, breadth 40 mm .
REMARKS. ··1840 Sowerby cf. Mr. Cuming: more
oblique, more closely tuberculated and ribbed than C.
marmoreum. ··1845 Reeve: the sculpture of this species
is well characterized by the oblique alternately
diverging minute scales, which ornament the middle
ribs. HABITAT. ··1845 Reeve: found in fine coral sand
on the reefs. ··1979 Kay: commonly occurring species
in shallow waters shoreward of fringing reefs and to
depths of less than 20 m; occasional valves have been
dredged to depths of60 m. DISTRIBUTION. ··1845 Reeve:
lsI. of Annaa, Pacific Ocean. ··1979 Kay: C. orbita is
Phuket Marine Biological Center Special Publication vol. 30 (2004)
667
orblta - ornatum
a wide-ranging species in the Indo-West Pacific. --1994
Hylleberg: Hawaii, Australia.
orbita Reeve. - See under orbita Broderip & Sowerby,
1833.
orbita hawaiensis Dall, Bartsch & Rehder, 1938
(subspecies). - Upgraded. Vasticardium hawaiensis
(Dall, Bartsch & Rehder, 1938). Cardiidae. Recent.
REcORDs. --1997 Vidal: Vasticardium orbita hawaiensis
(Dall, Bartsch & Rehder, 1938) new combination.
--1999 Vidal: Vasticardium orbita hawaiensis.
orbita mendanense Sowerby, 1897 (subspecies). Upgraded. Vasticardium mendanense (Sowerby, 1897).
Cardiidae. Recent. RECORDS. --1997 Vidal: Vasticardium orbita mendanense (Sowerby, 1897) new
combination. --1999 Vidal: Vasticardium orbita
mendanense. --1999 Vidal, p. 357: Vasticardium orbita
mendanense (Sowerby, 1897). Endemic of Marquis.
orbita philippinense Hedley, 1899 (subspecies). Figs.
Part 3 p. 866] . - Upgraded. Vasticardium philippinense
(Hedley, 1899). Cardiidae. Recent. REcORDs. --1997
Vidal: Vasticardium orbita philippinense (Hedley,
1899) new combination. --1999 Hylleberg, p. 40 I :
Vasticardium orbita philippinense (Hedley, 1899). ID
1. Vidal. "1999 Vidal, p. 357: Vasticardium orbita
philippinense (Hedley, 1899). "2003 Hylleberg &
Kilburn, p. 185, Plate 4, figs . 14 a,b. Vasticardium
orbita philippinense (Hedley, 1899). - CHARACTERS
AND OCCURRENCE - DISTRIBUTION. --1999 Vidal:
Indonesia (Lombok, Surnbawa, Komodo. --2003
Hylleberg & Kilburn: Vietnam (range extension).
orbiter Bryan 1915. - Synonym of Trachycardium
hawaiensis Dall, Bartsch & Rehder 1938. » Regarding
Cardium orbiter: Cardiidae. Recent. RECORDS. --1939
ZR: Cardium orbiter in Bryan Nat. Hist. Hawaii (1915)
p. 446 pI. civ f. 19 = Cardium orbita in Ostergaard
Bull. 51 B.P. Mus. p. 8 = Trachycardium hawaiensis
sp. n. [Ref. Dall, Bartsch & Rehder 1938]. --1979 Kay:
printer's error of orbita (Sowerby, 1833).
orbus Uspenskaia, 1931. - Combination to be revised:
"Cardium" orbus Uspenskaia, 1931. Cardiidae.
Miocene. RECORDS. --1932 ZR : Cardium orbus
Uspenskaia, 1931 n. sp . Miocene Daghestan and
Transcaspian.
ordinatum Sowerby in Dixon, 1850. - Combination to
be revised: "Cardium " ordinatum Sowerby in Dixon,
1850. Cardiidae. Fossil. RECORDS. --1922-32 Sherborn,
p. 4606: Cardium ordinatum Sowerby in Dixon, 1850.
--1980 Ruhoff: Cardium ordinatum Sowerby in Dixon,
1850; p. 52, 169, pI. 3, fig. 17.
orfellensis (variety). - See under martellii orfellensis
Oliveri, 1944.
orgrinensis (subspecies). - See under sturi orgrinensis
Andreescu, 1980.
orheiensis Gillet, 1938. - Phyllicardium orheiensis
Gillet, 1938. Lymnocardiidae. Sarrnatian. RECORDS.
--1939 ZR: Phyllicardium orheiensis Gillet, 1938 sp.
n. Sarrnatian Roumania.
orientale Janicchewskil 1900. - Conocardium orientale
Janicchewskil 1900. Conocardiidae. Fossil. RECORDS.
--1942 ZR: Conocardium orientale Janicchewskil 1900
[Ref. Branson 1942] .
orlentatis (subspecies, race, variety). - See under
otiophora orientaUs Stef. - otiophorum orientalis Stef.
- peregrinorsa orientaUs Douville, 1917.
orientalis Papaianopol, 1982. - Luxuridacna orientatis
Papaianopol, 1982. Lymnocardiidae. Upper Miocene.
TYPE LOCALITY. Bassin Dacique, Romania . RECORDS.
--1982 Papaianopol, p. 62, fig.: Luxuridacna orientatis
sp. nov. --1987 ZR: Luxuridacna orientalis sp. nov.
Romania, Upper Miocene. Ref. to Papaianopol p. 62.
orlentaUs Papaianopol, 1989 . - Pachydacna
(Pachydacna) orientalis Papaianopol, 1989.
Lymnocardiidae. Pliocene. TYPE LOCALITY. Bassin
Dacique, Romania. RECORDS. --1989 Papaianopol, p.
86, fig.: Pachydacna (Pachydacna) orientalis sp. nov.
orlentaUs Stef.- Prosodacna orientalis Stef.
Lyrnnocardiidae. Pliocene. RECORDS. --1987 Lutskanov
& Tcholakov: Prosodacna (Stylodacna) orientaUs Stef.
oriovacense (mode). - See under oriovacensis.
orlovacensis Neum. & Paul. - Didacna oriovacense
Neum. & Paul. Lymnocardiidae. Miocene. RECORDS.
--1946 ZR: Didacna oriovacensis Neum. & Paul
Pontian, Miocene [Ref. Gillet 1943] . "1987
Taktakishvili : Cardium oriovacense Neum.
oriovacensls banaticum Gillet, 1943 (variety) . Upgraded. Didacna oriovacensis banaticum Gillet,
1943. Lyrnnocardiidae. Miocene. RECORDS. --1946 ZR:
Didacna oriovacensis Neum. & Paul var. banaticum
Gillet, 1943 nov. Ponti an, Miocene, Rumania.
ornata (mode). - See under ornatum.
ornata kostlci - Caladacna ornata kostici.
Lymnocardiidae. Fossil. RECORDS. --1987 Taktakishvili:
Caladacna ornata kostici.
*ornatum; Cardiidae. - Homonymy to be noticed.
Cardium ornatum from 3 sources: Orbigny, 1850. Eichwa1d, 1855. - Sowerby, 1876.
ornatum Eichwald, 1855.- Synonym of Cerastoderma
glaucum (Bruguiere). » Regarding Cardium ornatum:
Cardiidae. Recent. REMARKS. Cerastoderma lamarcki
(Reeve) = Cerastoderma glaucum (Bruguiere).
RECORDS. --1876 Grimm : Cardium ornatum. --1890
Paetel: Cardium ornatum Eichw. --1968 Logvinenko
& Starobogatov: Cardium ornatum Eichwald, 1855 =
Cerastoderma lamarcki (Reeve).
ornatum Orbigny, 1850. - Combination to be revised:
"Cardium " ornatum d'Orbigny, 1850. - ?Cardiidae.
Fossil. RECORDS. "1922-32 Sherborn. 4618: Cardium
(Hall) ornatum d'Orbigny, 1850. - Pinnopsis. --192232 Sherborn: Pinnopsis ornatum Hall.
ornatulII Sowerby, 1876. - New name Trachycardium
sowerbyorum Voskuil & Onverwagt, 1992 . OTHER
JUDGEMENTS: Vasticardium ornatum (Sowerby, 1877).
» Regarding Cardium ornatum: Cardiidae. Recent.
DISTRIBUTION. Hong Kong. RECORDS. --1876 Sowerby:
Cardium ornatum . --1876 ZR: Cardium ornatum
Sowerby, 1876 sp . n. Hongkong. "1890 Paetel:
Cardium ornatum Sow. --1992 Voskuil & Onverwagt,
p. 41: Acrosterigma ornatum (Sowerby, 1876) nec
Orbigny 1850 = Trachycardium sowerbyorum nom.
nov. "1997 Vidal : Vasticardium ornatum (Sowerby,
1877). --1999 Vidal: Vasticardium ornatum (Sowerby,
1877).
*ornatum; Conocardiidae. - Homonymy to be
noticed. Conocardium ornatum from 2 sources:
Winchell & Marcy, 1865. -Cleland, 1912.
ornatulII Cleland, 1911. - New name Conocardium
plinthinatus Branson, 1942. Conocardiidae. Devonian.
REMARKS. Conocardiids have no connection with the
668
Tropical Marine Mollusc Programme (TMMP)
ornatum - otiophora
Cardiacea except analogy of form, which does not offer
any point of relationship (DaIl1895 , p. 5\9). RECORDS.
··1912 ZR: Conocardium ornatum Cleland, 1912 sp.
n. Wisconsin Devonian. ··1942 ZR : Conacardium
ornatum Cleland 1911 non Winchell & Marcy, 1865 =
Conocardium plinthinatus nom. nov. Branson, 1942.
ornatum Winchell & Marcy, 1865. - Conocardium
ornatum Winchell & Marcy, 1865. Conocardiidae .
Fossil. REMARKS. Conocardiids have no connection with
the Cardiacea except analogy of form, which does not
offer any point of relationship (Dall 1895, p. 519).
REcORDs. ··1942 ZR: Conocardium ornatllm Winchell
& Marcy 1865 [Ref. Branson 1942]. ··1980 Ruhoff:
Conocardium ornatum Winchell & Marcy, 1865; vol.
I: Ill , pI. 2, fig. 15.
ornatus Hall, 1843.- Synonym of Lunulicardia ornatus
(Hall, 1843). Not in Cardiidae. Devonian. REcoRDs.
··1997 Vidal: Pinnopsis ornatus Hall, 1843 [Devonian]
= Lunulicardia ornatus (Hall, 1843).
Lymnocardiidae.
*ornatum; Lymnocardiidae - Homonymy to be
noticed. Lymnocardium ornatum from 2 sources:
Pavlovic, 1929. Papp (?year).
ornatum Papp. - New combination: Lymnocardium
ornatum Papp. Lymnocardiidae. Fossil. RECORDS .
··1987 Taktakishvili: Limnocardium ornatum ornatum
Papp.
ornatum Pavlovic, 1929 . - Changed combination :
Caladacna ornatum (Pavlovic, 1929). Lyrnnocardiidae.
Pontian . RECORDS. ··1928 ZR : Limnocardium
(Pontalmyra) ornatum [Ref. Pavlovitch 1928]. ··1929
ZR: Limnocardium (Pontalmyra) ornatum Pavlovic,
1929 sp. n. Pontian Serbia. ··1987 Taktakishvili:
Limnocardium (Pontalmyra) ornatum. ··1987
Taktakishvili: Caladacna ornata (Pavl.)
ornatum biseptum Papp, 1953 (subspecies). - New
combination: Lymnocardium ornatum biseptum (Papp,
1953). Lymnocardiidae. Fossil. RECORDs. ··1948 ZR:
Limnocardium ornatum biseptum march sic [n. sp.] sic
[nomen nud .] [Ref. Papp 1948] . セᄋQYUV@
ZR:
Laevicardillm ornatum biseptum Papp, 1953 subsp.
nov. "1987 Taktakishvili: Limnocardium ornatum
biseptum Papp.
ornithopetronica Chapman & Crespin, 1928. Protocardia ornithopetronica Chapman & Crespin,
1928 . Cardiidae. Tertiary. RECORDS . .. 1928 ZR:
Protocardia [vide etiam Protocardium] ornithopetronica Chapman & Crespin, 1928 sp. n. Tertiary
Victoria.
orolesi (subspecies). - See under munieri orolesi.
orolesi Papaianopol, 1982. - Changed combination:
Euxinicardium orolesi (Papaianopol, 1982). Lyrnnocardiidae. Pliocene. TYPE LOCALITY. Pontien et Dacien
du Bassin Dacique, Romania. REcORDs. ··1982 Papaianopol, p. 200, fig .: Limnocardium (Euxinicardium)
orolesi sp. nov. ··1987 Taktakishvili : Euxinicardium
orolesi Papaian.
orsa (variety). - See underfervidum orsa Eberzin, 1958.
Orthocardiinae Schneider, 2002 (subfamily). RECORDS.
··2002 Schneider: Orthocardiinae subfam. nov. Type
genus: Orthocardium Tremlett, 1950.
Orthocardium Tremlett, 1950 (genus) . - Cardiidae
Middle Eocene and Upper Eocene. RECORDS. ··1998
Schneider (1998b): Orthocardium placed in Fraginae
by Popov (1977) and Kafanov & Popov (1977). Placed
in Cardiinae by Schneider 1992. ··2002 Schneider:
Orthocardium: Cardiinae , not Fraginae. CHARACTERS AND OCCURRENCE - DESCRIPTIONS. ··2002 Schneider: Cross striae present in rib
interspaces. Radially ribs weakly expressed internally,
adductor muscles strongly impressed. Ridges on lunule
flap absent. Perisiphonal suture present. Very heavy
and imbricated spines. ··1969 Moore et al.: the shell
of Orthocardium has rib-sculptUre tending to form
vertical frill at rib crest; outline more quadrate than in
V. (Vepricardium), hinge line narrower and straighter.
Orthocardium Tremlett, 1950 (subgenus). - Cardiidae
Middle Eocene and Upper Eocene. GENOTYPE. Tremlett
(1950) designated Cardium poru{osum Solander, 1766
type species of subgenus Orthocardium of genus
Vepricardium. RECORDS. ··1950 Tremlett: Orthocardium is less globose than Vepricardium and has a
straight, thinner hinge line. ··1950 ZR: Orthocardium
Tremlett, 1950 subgen. nov. of Cardium. Subgenotype
Cardium porulosum Solander, Middle and Upper
Eocene. ··1969 Moore et al.: Orthocardium. Europe.
··1977 Popov: Orthocardium Tremlett, 1950.
Genotype: Cardium porulosum Solander, 1766. ··1980
Vokes: Orthocardium Tremlett, 1950 in subfamily
Cardiinae . .. 1992 Schneider p. 149: Orthocardium
Tremlett, 1950 . .. 1995 Pacaud & Renard : (Orthocardium) Tremlett, 1950 subgenus of Cardium Linne,
1758 in subfamily Cardiinae.
orthogonale Buvignier, 1852 . - Combination to be
revised: "Cardium" orthogonale Buvignier. Cardiidae.
Li as-Jurassic. RECORDS. ··1980 Ruhoff: Cardium
orthogonale Buvignier, 1852; atlas p. 16, pI. 15, figs.
4-6. ··1999 - Jacques Vidal (pers. com.), MNHN, Paris:
Fossils from the Paleontological Collection. Cardium
orthogonale Buvignier. Sequanien. France (Cher),
S05033.
orthogonalis lexoviensis Chavan, 1952 (variety) . Upgraded. New combination: Protocardia orthogonalis
lexoviensis Bigot MS (Chavan, 1952). Cardiidae.
Jurassic and Cretaceous. RECORDS. ··1985 Fiirsich &
Werner: Protocardia (Protacardia) orthogonalis var.
lexoviensis Bigot MS (Chavan, 1952).
orthoprymnos Scott, 1978. - Pleuriocardia orthoprymnos. Cardiidae. Cretaceous. RECORDS. ··1978 Scott, p.
895, pI. \, figs. 12-13: Pleuriocardia (Pleuriocardia)
orthoprymnos n. sp. ··1978 ZR: Pleuriocardia
(Pleuriocardia) orthoprymnos Scott, 1978 sp. nov.
Texas, Lower Cretaceous. Ref. to Scott p. 895.
osijeki Basch, 1990. - New name: Prosodacna osijeki
Basch, 1990. Lymnocardiidae. Fossil. RECORDS. ··1990
Basch, p. 65 : Prosodacna (Prosodacnomya) osijeki
nom. nov. for Limnocardium stenopleura Brusina &
Brusina, 1902, nec Stefanescu.
Osoinia Stevanovic, 1979 (subgenus). Lymnocardiidae.
Fossil. RECORDS. "1979 ZR: Pseudocatillus (Osoinia)
Stevanovic, 1979. Subgen. nov. Type species P
laticostatus Stevanovic 1951. Ref. to Stevanovic, p.
326.
ossoinae (subspecies and variety). - See under retowskii
ossoinae Stevanovic, 1951.
olarii (variety). - See praechinatum otarii Kharatishvili,
1958.
otiophora Brusina. - Changed combination: Pontalmyra
Phuket Marine Biological Center Special Publication vol. 30 (2004)
669
otiophora orientalis - ovata
otiophora (Brusina). Lymnocardiidae. Fossil. REcORDs.
•• I 987 Taktakishvi1i: Pontalmyra otiophora (Brus.).
otiophora orientalis Stevanovic, 195 I (variety) . Upgraded. Pontalmyra otiophora orienta tis
(Stevanovic, 195 I). Lymnocardiidae. Fossil. REcORDs.
. •• I 953 ZR: Didacna otiophorum var. orientatis
Stevanovic, 195 I nov. Ponti an Serbia. ··1987
Taktakishvili: Pontalmyra otiophora orienta lis (Stev.).
otiosa Papaianopol, 198 I. - Pontalmyra otiosa
Papaianopol, 198 I. Lymnocardiidae. Upper Miocene.
RECORDS . •• I 987 Taktakishvili: Pontalmyra otiosa
Papa ian . •• I 98 I ZR: Pontalmyra (Pontalmyra) otiosa
Papaianopol , 1981 sp. nov. Romania . Ref. to
Papaianopol, p. 2 I.
otto; Geinitz. - Synonym of Trachycardium incomptum
(Sowerby apud Forbes, 1846).» Regarding Cardium
ottoi : Cardiidae. Cretaceous. RECORDS . ·'1871
Stoliczka: Cardium ottoi (Cerastoderma) = Cardium
(Trachycardium) incomptum Sowerby apud Forbes,
1846, according to Zittel.
ottonis Geinitz, 1843. - Combination to be revised :
"Cardium " ottonis Geinitz, 1843. Cardiidae. Fossil.
REcORDs. ··1922-32 Sherborn, p. 4645: Cardium ottonis
Geinitz, 1843.
otukai Hatai & Nisiyama, 1952. - See under otukai
Oyama, Mizuno & Sakamoto, 1960.
otukai Oyama, Mizuno & Sakamoto, 1960. Invalid name.
OTHER JUDGEMENTS: Vasticardium otukai Oyama,
Mizuno & Sakamoto, 1960. » Regarding V. otukai:
Cardiidae. Upper Oligocene. REcORDs . •• I 973 Keen :
Vasticardium otukai Oyama, Mizuno & Sakamoto,
1960, ex Hatai & Nisiyama, MS. ··1982 ZR :
Vasticardium otukai Hatai & Nisiyama, 1952. Invalid
name. Ref. to Masuda et al. p. 7. ··1999 Vidal:
Vasticardium otukai Hatai &. Nisiyama, 1952: 35.
ouralicum Verneui1 in Murchison, 1845. - Conocardium
ouralicum, de Verneuil. Conocardiidae . Fossil.
REcORDs. ··1859 Chenu: Conocardium ouralicum de
Verneuil. •• I 922-32 Sherborn, p. 6763 : Cardium
ouralicum Verneuil in Murchison, 1845 .
ovale Montagu, 1803. - See under ovale Sowerby, 1840.
ovale Pcelinzev, 1928. - Proto cardium ovale Pcelinzev,
1928 . Cardiidae. Jura. RECORDS . •• 1928 ZR :
Protocardium ovale [vide etiam Protocardia] Pcelinzev,
1928 sp. n . Jura. Turcoman USSR
ovale Sowerby, 1840. [Figs. Part I p . 144. Part 3 p. 9 19].
- New name and changed combination : Parvicardium
ovale (Sowerby, 1840). OTHER JUDGEMENTS: Cardium
elongatum Montagu . - Cardium fascia tum Montagu.
- Parvicardium pinnulatum (Conrad , 183 I). »
Regarding Cardium ovale: Cardiidae. Recent. TYPE
LOCALITY. Britain. RECORDS . •• I 840 Sowerby, catalogue
p. 2, No. 24 : Cardium ovale nob., new name for C.
elongatum Mont. (preoccupied) . •• I 845 Reeve, sp. I 19:
Cardium ovale Sowerby, Cat. Card . Conch. lIIus., no.
24. ··1846 Loven: Cardium ovale Sow. = Cardium
elongatum Mont. "1856 Wood & Hanley: C. ovale
Sow. = Cardium fasciatum Mont. •• I 858 Adams &
Adams, p. 456: Cardium (Cerastoderma) ovale Sow.
··1859 Sowerby: C. ovale = Cardiumfasciatum Mont.
•• I 867 Weinkauff: Cardium ovale Brown, Sowerby =
Cardium fascia tum Montagu. ··1869 Romer: Cardium
ovale Sowerby fig. 24 = Cardiumfasciatum Montagu.
•• I 884 Grasset: Cardium ovale Sow. •• I 886 Locard:
Cardium ovale Sowerby = Cardium fasciatum
Montagu, 1808. ··1888 Kobelt: Cardium ovale Sow. =
Cardiumfasciatum Montagu . •• I 888 Servain: Cardium
ovale Sowerby, Reeve = Cardiumfasciatum Montagu,
1808 . •• I 890 Paetel : Cardium ovale Sow. = fasciatum
Mtg . •• I 9 I 7 Hidalgo: Cardium ovale Reeve sic =
Cardium fasciatum Montagu . •• I 922-32 Sherborn, p.
4647: Cardium ovale Sowerby, ? 1840. ··1932
Winckworth: Cardium (Parvicardium) ovale Sowerby,
1840. ··1962 Allen : Cardium (Parvicardium) ovale
Sowerby 1840 = Cardium fasciatum in Jeffreys 1863.
•• 1969 Nordsieck : Parvicardium ovale (Sowerby,
184 I). ··1973 ZR: Cardium ovale Sowerby. Ref. to
Rasmussen . •• I 974 Clarke: Parvicardium ovale
(Sowerby) . •• I 987 ZR: Parvicardium ovale (Sowerby,
1840). Senior synonym of Cardium fasciatum
Montagu, 1808. Ref. to Backeljau et al. p. 4 . •• I 998
Macedo et af. : Parvicardium ovale (Sowerby G.B. III,
1840). ··2000 Aartsen & Goud : Cardium ovale
Sowerby, 1840 = Cardium ovale Sowerby, 1841 (1840
is most probably the correct year). ··2000 Aartsen &
Goud: Cardium ovale Sowerby, 1840 is a synonym of
Parvicardium pinnulatum (Conrad, 1831). ··200 I
Vidal, p. 408 : Parvicardium ovale. - CHARACTERS
AND OCCURRENCE - DESCRIPTIONS. ··1845 Reeve :
shell thin, somewhat squarely ovate, rather depressed,
posteriorly slightly elongated; radiately ribbed, ribs,
the anterior especially, obscurely very finely muricated;
whitish, faintly stained and banded with brown. ··200 I
Vidal: only the exhalant aperture is tube-shaped.
Inhalant siphon open towards the foot aperture. No
eyes. REMARKS . •• I 969 Nordsieck: Parvicardium ovale
(Sowerby, 1840) = Parvicardium fasciatum (Montagu,
1803). HABITAT. ··1969 Nordsieck: 0 to 350 m depth.
DISTRJBUTION. ··1845 Reeve: Britain (Torbay).·· I 884
Grasset: Sicily. •• I 969 Nordsieck: Arctic, North
Norway to the Canary Islands, Mediterranean.
ovalis (mode). - See also under ovale.
ovalis Barinov, 1992. - Clinocardium ovalis Barinov,
1992. Cardiidae. Neogene. TYPE LOCALITY. Kamtchatka,
Russia. RECORDS . •• I 992 Barinov et al., p. 124, fig .:
Clinocardium ovalis Barinov, 1992 sp. nov.
ovalis Martinson, 1959. - Pseudocardinia ovalis
Martinson, 1959. Cardiacea. Fossil. RECORDS . •• I 959
ZR: Pseudocardinia ovalis Martinson, 1959 sp. nov.
Sary-Chelek, eastern Fergana, fossil.
ovalis Yanina, 1983 . - Didacna ovalis Yanina, 1983.
Lymnocardiidae. Middle Pleistocene. RECORDS. ··1983
ZR : Didacna ovatis Yanina, 1983 sp. nov. No
description given . Russian SFSR. Ref. to Yanina, p.
36.
ovaloides (variety). - See under bistriata ovaloides
Sadykov, 1962.
ovalum Alieva, 1967. - Combination to be revised:
"Cardium " ovalum Alieva, 1967. Cardiidae. Tertiary.
RECORDS. ··1967 ZR: Cardium ovalum Alieva, 1967
sp. nov. figs . Akchagil deposits, Naftalan, Azerbaidjan,
SSR.
ovata (mode, subspecies, variety). - See under ovatum.
-echinatum ovata Jeffreys, 1863 . - sulcatina ovata
Deshayes. - exiguum ovata Milachewitch, 19 I 6. exiguum f. ovata Milachewitch, 19 I 6.
ovata Papaianopol, 1975. - Chartoconcha ovata
Papaianopol, 1975. Cardiacea. Pliocene . RECORDS.
670
Tropical Marine Mollusc Programme (TMMP)
ovalu", - ovuloides
"1975 ZR: Chartoconcha ovata Papaianopol, 1975 sp.
nov. Romania. Prahova, Lacuri. Ref. to Papaianopol,
p. 136.
*ovatu",. - Homonymy to be noticed. Cardium ovatum
from 2 sources: Miinster, 1840. - Deshayes, 1838.
ovalu", Deshayes, 1838. - Changed combination:
Didacna ovata (Deshayes, 1838). Lymnocardiidae.
Tertiary. TYPE LOCALITY. The Crimea. RECORDS. "1838
Deshayes, p. 56, pI. I, figs. J9-21: Cardium ovatum.
··1857 Deshayes: Cardium ovatum Deshayes, 1837 sic.
"1877 Grimm: Cardium ovatum Deshayes. "1905
Dollfus, commenting Andrussow 1903: Cardium
ovatum. "1922-32 Sherborn, p. 4652: Cardium ovatum
Deshayes, 1838. "1953 Celidje: Didacna ovata Desh.
"1987 Anaya: Didacna (s. s.) the group ovata
(Deshayes, 1810 sic). "1987 Taktakishvili: Didacna
ovata Desh.
ovalu", (aff.) Deshayes, 1838 . - RECORDS. "1987
Taktakishvili: Pontalmyra aff. ovata (Desh.).
ovalu", Miinster, 1840. - Replaced by Cardium cornutum Deshayes. » Regarding Cardium ovatum: Cardiidae. Fossil. RECORDS. "1857 Deshayes: Cardium ovatum Miinster, 1840 - C. cornutum Desh. "1922-32
Sherborn, p. 4654: Lunulacardium ovatum Muenster,
1840.
Ovicardiu", Marwick, 1944 (subgenus). - Cardiidae.
Pliocene. Type species. Original designation . Trachycardium (Ovicardium) rossi Marwick, 1944. CHARACTERS. Shell ovate, intercostal spaces cross-striate, with
vertical sides; hinge with oblique groove behind 4b.
New Zealand-Australia (Moore et al. 1969). RECORDS.
"1944 ZR: Ovicardium Marwick, 1944 subgen. nov.
of Trachycardium Morch, type Trachycardium (Ovicardium) rossi Marwick, 1944 sp. nov. (Opoitian) Te
Reinga Falls, Hangarua R. "1969 Moore et al. : Acrosterigma (Ovicardium) rossi Marwick, 1944. "1980
Vokes: Ovicardium Marwick, 1944 in subfamily
Trachycardiinae.
ovicella Dujardin. - "Cardium" ovicella Dujardin.
Cardiidae. Miocene. RECORDS. "1999 Jacques Vidal
(pers. com.), MNHN, Paris: Fossils from the Paleontological Collection. Cardium ovicella Dujardin.
Helvetien. France (Touraine), B.27157.
ovifor",is - Tauricardium oviformis. Lyrnnocardiidae.
Fossil. RECORDS. "1987 Taktakishvili: Tauricardium
oviformis.
ovifor",is (subspecies and variety). - See under petersi
oviformis and subsquamulosum oVlformis Eberzin,
1958.
ovipula",en Reeve, 1844. [Figs. Part I p. 130. Part 3 p.
894). - Synonym of Laevicardium serra tum (Linne,
1758). OTHER JUDGEMENTS: Laevicardium brasilianum
(Lamarck, 1819). » Regarding C. oviputamen:
Cardiidae. Recent. TYPE LOCALITY. Unknown (Reeve
1844). TYPE MATERJAL. University Museum of Zoology,
Cambridge, coil. 1. Saul no [691 (1. Vidal pers. comm.).
RECORDS . ,,1844. Reeve, p. 168, sp. 36: Cardium
oviputamen . "1858. Adams & Adams, p. 457:
Laevicardium oviputamen, Reeve. "1864. Krebs :
Cardium oviputamen Reeve. "1868. ZR: Cardium
elenense (Sow.) and oviputamen (Reeve) shortly
characterized [Ref. Romer 1868). "1884. Grasset:
Laevicardium oviputamen. R v . •• [890. Paetel:
Laevicardium oviputamen Rv. "1902. Dall:? Cardium
oviputamen Reeve, 1844 = Cardium serra tum
Linnaeus, 1758. "1921. Maury: ovipulamen Reeve,
1844 = Cardium (Laevicardium) serratum Linnaeus,
1758 . "1922-32. Sherborn , p. 4658: Cardium
oviputamen Reeve, [844 . •• [925 Woodring, p. 145:
Cardium oviputamen Reeve, [844 is identical with
Cardium serratum Linne, [758 . •• [977. Fischer-Piette:
Cardium oviputamen Reeve, 1844 = Laevicardium
brasilianum Lamarck, 1819. - CHARACTERS AND
OCCURRENCE - CHARACTERS of diagnostic value:
concentrically waved pinkish brown marks, though
faintly indicated, are characteristic. DESCRIPTIONS .
"1844 Reeve: shell obliquely ovate, thin, ventricose,
radiately very finely striated; opaque snowy white,
concentrically clouded with faint waved pinkish-brown
spots or lines, covered with a pale epidermis towards
the margins; margins finely crenulated within.
REMARKS. "1844 Reeve: the general appearance is very
like that of the Cardium serra tum; however the form
is of Jess oblique, and the Cardium oviputamen under
consideration is more ventricose. The concentrically
waved pinkish brown marks above noticed, though
faintly indicated, are nevertheless characteristic, as
distinguished from those of a different pattern in
Cardium vitellinum Reeve. [Present generic classification is Laevicardium oviputamen). DISTRIBUTION.
··1844 Reeve: unknown. "1884 Grasset: Bahamas.
"Of authors: Florida, Panama, West Indies.
ovolu", - Combination to be revised: "Cardium"
ovolum. Unfamiliar species. » Regarding Cardium
ovolum: Cardiidae. Cretaceous. TYPE LOCALITY.
Palaestine. RECORDS. "187 [ Stoliczka: Cardium ovolum
(Stoliczka: unfamiliar species).
ovuloide (error). - Printer's mistake for ovuloides in Dall
1901.
ovuloides Reeve, 1845. [Figs. Part [ p. 145. Part 3 p.
856, 931) . - Changed combination: Synonym of
Trigoniocardia obovale (Sowerby, 1833). OTHER
JUDGEMENTS' Trigoniocardia ovuloides (Reeve 1845).
» Regarding Cardium ovuloides: Cardiidae. Recent.
TYPE REPOSITORY. 3 syntypes. 1978139. BMNH London.
TYPE LOCALITY. Reeve 1845: unknown. RECORDS. "1845
Reeve: Cardium ovuloides. "1858 Adams & Adams,
p. 459: Hemicardia (Fragum) ovuloides, Reeve. "1890
Paetel : Hemicardium ovuloides Rv. "1901 Dall:
Cardium (Trigoniocardia) ovuloide sic Reeve, 1845.
"1922-32 Sherborn, p. 4660: Cardium ovuloides
Reeve, 1845. "1932 Pilsbry & Lowe: Cardium
(Trigoniocardia) ovoluides Rve. "1958 Abbott :
Trigoniocardia ovuloides Reeve [Recent). "1961
Olsson: Cardium oVllloides Reeve, 1845 = Trigoniocardia (Apiocardia) obovale (Sowerby, 1833). "1977
Popov: Trigoniocardia (Trigoniocardia) ovuloides
(Reeve, 1845). "1977 Fischer-Piette: Cardium ovuloides Reeve = Corculum obovale Sowerby. - CHARACTERS AND OCCURRENCE - DESCRIPTIONS. "1845
Reeve: shell oblong-ovate, very gibbous, radiately
ribbed, about 17 ribs, the middle and anterior large,
strong, nodulously granulated, interstices transversely
impressly striated, posterior ribs narrow, smooth,
flattened; yellowish white within and without.
REMARKS. This little species exhibits the form of the
Cardium obovale with the sculpture of the Cardium
graniferum. DISTRIBUTION. "1845 Reeve: unknown.
Phukel Marine Biological Center Special Publication vol. 30 (2004)
671
ovum - oxypleura
··Of authors : California, Ecuador.
ovum Martyn, 1784. Non binominal. - Not available.
» Regarding Cochlea ovum: Cardiidae. Recent. TYPE
LOCALITY. Pulo-Condore (Martyn, 1784). RECORDS.
··1784 Martyn, p. 24, sp. 79, pI. 28, fig. 1: Cochlea
ovum. ··1789 Martyn: Cochlea ovum. Pulo Condore.
- CHARACTERS AND OCCURRENCE - OF
COLLECTIONS. ··ZMUC: I specimen M U ?Cardium
ovum. The label refers to "cf79 in fig. Martyni".
Oxydacna Davitaschvili, 1930 (genus). - Lymnocardiidae. Tertiary. RECORDS. ··1962 ZR: Oxydacna
Davidaschvili in subfam. Didacninae Eberzin, 1962,
family Cardiidae [Ref. Eberzin 1962]. ··1980 Vokes:
Oxydacna Davidaschvili, 1930 in subfamily
Didacninae. ··1987 Taktakishvili, p. 90: Oxydacna
Davitaschvili, 1930; referred to new tribe Pontalmyrini.
oxygonum Sowerby, 1834. [Figs. Part I p. 107, 139. Part
3 p. 866, 878] . - Changed combination: Acrosterigma
oxygonum (Sowerby, 1834). Cardiidae. Recent. TYPE
MATERlAL. Two shells considered as syntypes in BMNH,
Cuming collection, labeled Philippines. The dimensions
and rib number given by Sowerby could fit the
specimen figured by Vidal (1999, figs. 5D-E); but the
locality given by Sowerby (China Sea) does not agree.
TYPE LOCALITY. ad mare Sinense (Chiga) (Sowerby
1840). RECORDS. ··1834 Sowerby, 47t part, fig. 9:
Cardium oxygonum. ··1840 Sowerby, p. 107: Cardium
oxygonum Sow. ··1840 Sowerby, Proc. Z.S. London
p. 107: Cardium oxygonum. ··1840 Sowerby, catalogue
p. 4, No. 50: Cardium oxygonum Sow. ··1840 Sowerby,
corrected list, p. 8: Cardium oxygonum, (Sowerby No.
50). ··1841 Reeve, p. 99: Cardium oxygonum Sow.
··1843 Hanley: Cardium oxygonum Sow. ··1845 Reeve,
sp. 77 : Cardium oxygonum Sow. ··1858 Adams &
Adams, p. 455: Cardium (Trachycardium) oxygonum,
Sow. ··1867 Mitchell: Cardium oxygonum Sow. ··1869
Frauenfeld: Laevicardium oxygonum Sow. ··1870
March: c. oxygonum Sowb junior synonym of Cardium
leucostoma Born. ··1872 Tryon: Cardium oxygonum
Sow. ··1884 Grasset: Cardium oxygonum Sow. ··1890.
Paetel: Cardium oxygonum Sow. ··1903 Hidalgo:
Cardium oxygonum Sow. ··1905 Hidalgo: Cardium
oxygonum Sow. ··1906 Standen & Leicester: Cardium
oxygonum. ··1912 Shirley: Cardium oxygonum Sow.
··1922-32 Sherborn, p. 4665: Cardium oxygonum
Sowerby, 1841. ··1923 Hedley: Cardium oxygonum
Sow. (erroneous identification, see Vidal 1999). ··1928
Faustino : Cardium oxygonum Sowerby. ··1941
Gravely : Cardium oxygonum . ··1962 Iredale &
McMichael: Cardium oxygonum Sow. ··1977 FischerPiette: Laevicardium oxygonum Sowerby. ··1977
Wilson & Stevenson: Cardium oxygonum Sowerby,
1841. ··1978 ZR : Trachycardium oxygonum. Bay of
Bengal Burma, Maungmagan. First record for Burmese
waters. Ref. to Ray ··1992 Oliver: Trachycardium
oxygonum. ··1999 Vidal : A croslerigma oxygonum
(Sowerby, 1833). ··1999 Vidal: Cardium oxygonum
Sowerby, 1833 in Hedley, 1923: 304. = Acrosterigma.
··2002 Hylleberg & Kilburn, p. 63: Acrosterigma
oxygonum (Sowerby, 1834). - CHARACTERS AND
OCCURRENCE - OF COLLECTIONS. ··ZMUC: one
specimen identified by Cuming, sine loc. CHARACTERS
of diagnostic value: the elaborate and distinctive rib
morphology. DESCRlPTJONS. ··1845 Reeve: Shell oblong-
ovate, attenuated towards the umbones, radiately
ribbed, 35 ribs, peculiarly sharp angled , finely
crenulated at the base on the posterior side, extreme
posterior ribs prickly, anterior very finely crenated;
white, profusely stained and spotted with rose. ··1999
Vidal: Shell ovoid, and almost equilateral, sometimes
with posterior dorsal margin a little straightened; hardly
elongated and not appreciably compressed. Ribs
straight in projection. Lunule about equivalent on both
valves, well delineated, and appreciably hollowed on
both sides. External colour beige, variegated with rather
dark brown to purple in large irregular patches; interior
white, except for umbonal rays. Hinge appreciably
asymmetrical. A long, thin, weak umbonal sterigma in
about 20% of the specimens. Mean rib number 38.9 ±
lA, range 36-41. Posteriorly ribs of equivalent width,
with a weakly marked median furrow and scales that
are sometimes irregular; interstices well marked.
Elsewhere, ribs always triangular, except for the first
anterior ribs, which are rounded; on median posterior
part, ribs first retro-ridged , becoming retrotuberculated anteriorly, with numerous small tubercles
situated on lower part of flank; anterior flank smooth,
and a significant smooth crestal fold is present. On
anterior half, anterior flanks of ribs become at first protuberculated at base, then pro-ridged , with ridges
terminating in crestal tubercles, but not reaching
interstice; in contrast, posterior flanks of ribs become
and remain smooth throughout. Interstices triangular
and smooth. DIMENSIONS. ··1999 Vidal: length 37.0,
width 42.8 , thickness 26.0 mm . REMARKS. ··1840
Sowerby: this species resembles C. maculosum of
Wood in form, but in sculpture it is more like C.
angulatum of Lamarck, from which it is distinguished
by being narrower towards the umbo, less ventricose,
and having the ribs distinctly angulated. ··1845 Reeve:
this species exhibits a very peculiar plan of sculpture,
the ribs being sharply angled at the summit, and very
finely crenulated at the base, on the posterior side.
··1999 Vidal: A. oxygonum has very constant
characters, particularly an elaborate and distinctive rib
morphology. One lot of this species in BMNH (reg.
1910-12-13-15), from Sri Lanka, is labeled as a type
series of "Cardium sanguineotincta" Preston; another
lot AMS C034080, from Sri Lanka, is labeled as a
cotype lot of the same taxon. There is no verification
of this name in the literature. DISTRIBUTION. ··1845
Reeve China, Philippines. ··1884 Grasset: China.
··1923 Hedley: Australia, New South Wales,
Queensland. ··1994 Hylleberg: Indo-West Pacific.
1999 Vidal: exclusively in the northern hemisphere:
Gulf of MaJrnar, Andaman Sea and Malacca Strait, Gulf
of Thailand, (south?) China, Philippines. Published
records from Australia (South Queensland and New
South Wales) are erroneous and should be referred to
A. kerslakae. Specimens from Zanzibar need to be
confirmed. ··2002 Hylleberg & Kilburn: South India.
oxynoti Quensted, 1856. - Combination to be revised :
"Cardium" oxynoti Quensted, 1856. Cardiidae. Fossil.
RECORDS. ··1980 Ruhoff: Cardium oxynoti Quensted,
1856; [1856--1858] p. 110, pI. 13, fig. 46.
oxypleura Davidasvili, 1930 - Changed combination:
Pontalmyra oxypleura (Davidasvili, 1930). Lymnocardiidae. Fossil. RECORDS . ··1932 ZR: Didacna
672
Tropical Marine Mollusc Programme (TMMP)
ozurgeticum - pallidum
oxypleura Davidasvili, 1930 sp. n. 001987 Taktakishvili:
Pontalmyra oxypleura (Dav.).
ozurgeticum Davidasvili, 1930 - Changed combination:
Moquicardium ozurgeticum (Davidasvi1i, 1930) .
Lymnocardiidae. Fossil. REcORDs. 001932 ZR: Limnocardium ozurgeticum Davidasvili, 1930 sp. n. Pliocene
Transcaucasia. 001987 Taktakishvili: Limno-cardium
(Moquicardium) ozurgeticum Dav. (?) . 001987
Taktakishvili: Moquicardium ozurgeticum Dav. (?).
P
Pachycardium Conrad, 1869 (genus). - Cardiidae.
Cretaceous. Pachycardium spillmani Conrad, 1869, p.
96, type species of the genus Pachycardium. RECORDS.
001980 Vokes: Pachycardium Conrad, 1869 in
subfamily Protocardiinae. 001995 Schneider:
Protocardia (Pachycardium). Type species Protocardia
spillmani (Conrad, 1858).
.
Pachycardium Conrad, 1869 (subgenus). - Cardiidae.
Cretaceous. REMARKS. The designation "section" takes
the value of subgenus. RECORDS . 001876 Meek:
Pachycardium Conrad, 1870 synon. of Protocardia,
Beyrich, 1845. 00 1887 Fischer: Pachycardium Conrad,
1870. Section of subgenus Protocardia Beyrich.
Pachydacna Ebersin, 1955 (genus) . - New Name.
Lymnocardiidae. Tertiary. RECORDS. 001974 ZR:
Pachydacna Ebersin, 1955. Referred to new subfamily
Prosodacninae, p. 128. Ref. to Andreescu. 001980
Vokes : Pachydacna (genus) Ebersin, 1955. n.n. pro
Natella Ebersin, 1949. In. subfamily Lymnocardiinae.
001987 Anaya: Pachydacna Ebersin , 1955. Type
species: Natella natella Ebersin, 1947.
Pacbydaena (error). - Printer's mistake for genus
Pachydacna in ZR for 1970.
Pachyprionopleura Andreescu, 1974 (genus) .
Lymnocardiidae. Tertiary. RECORDS. 001974 ZR :
Pachyprionopleura Ahvlediani, 1970. Referred to new
subfamily Prosodacninae, p. 128. Ref. to Andreescu.
001980 Vokes: Pachyprionopleura Andreescu, 1974.
Pachyrismella (genus). - Transferred from
Megalodontacea. REcORDs. 001978 ZR: Pachyrismella
Cox 1964. Ref. to Morris p. 267.
pachyrrhinum Eichwald, 1868. - Combination to be
revised : "Cardium" pachyrrhinum Eichwald, 1868.
Cardiacea. Fossil. REcORDs. 001980 Ruhoff: Cardium
pachyrrhinum Eichwald, 1868; [1853-1868] vol. 2.
678, pI. 25, figs. 9-10.
padimeicum (subspecies). - See under cilia tum
padimeicum Merklin & Zarkhidze, 1979.
padimeicum Merklin & Zarkhidze 1979. - Synonym of
Cilia to cardium ciliatum (Fabricius 1780). Cardiidae.
Fossil. RECORDS. 001980 ZR: Ciliatocardium padimeicum Merklin & Zarkhidze 1979 is a junior synonym of
C. ciliatum (Fabricius 1780). Ref. to Kafanov.
paetrigonoides (error). - Printer's mistake for
praetrigonoides in ZR for 1916.
Palaeocardita (genus). - REcORDs. 001992 Schneider p.
146: Palaeocardita is usually placed with the
Carditidae. Type species Palaeocardita austriaca
(Hauer). 001995 Schneider: Palaeocardita. May not
belong to the cardiids.
palatum Martyn, 1784. Non binominal. - Not available.
» Regarding Cardium palatum: Not a cardiid .
RECORDs. 001784 Martyn: Cardium palatum.
pallasanum Basterot in Mayer, 1864. - Combination to
be revised: "Cardium" pallasanum Basterot in Mayer,
1864. Cardiacea. Fossil. RECORDS. 001980 Ruhoff:
Cardium pallasanum Basterot in Mayer, 1864; [18571869], vol. 12: 355.
*pal/asianum. - Homonymy to be noticed. Cardium
pallasianum from 2 sources: Basterot, 1825. Andrzejoswky, 1836.
pallasianum Andrzejoswky. 1836. - Replaced by Parvicardium exiguum (Gmel.). Cardiidae. Fossil. RECORDS.
001857 Deshayes: Cardium pallasianum Audrz sic.,
1836 [? = Andrzejoswky]- C. exiguum Gmel.?
pal/asianum Basterot, 1825. - Changed combination:
Loxocardium pallasianum (Basterot, 1825). Cardiidae.
Miocene. RECORDS. 001835 Lamarck: Cardium
pallasianum Bast. 001857 Deshayes : Cardium
pallasianum Basterot 1825. 001899 Sacco: Cardium
pallasianum Basterot 1825. 001899 Sacco:
Loxocardium pallasianum (Bast.). 001922-32 Sherborn,
p. 4689: Cardium pallassianum sic de Basterot, 1825.
001977 Popov: Loxocardium pallasianum (Basterot,
1825).001999 Jacques Vidal (pers. com.), MNHN,
Paris: Fossils from the Paleontological Collection.
Cardium pallas ian urn Bastero!. Helvetien . France,
B26227.
pallassianum (error). - Printer's mistake for pallasianum
in Sherborn 1922-32, p. 4689
pal/eus. - Label name for Laevicardium laevigatum
(Linne, 1758). Cardiidae. Recent. OF COLLECTIONS.
ooZMUC: 1 specimen in dry collection referring to
Reeve (1845) No. 69, palleus = Laevicardium
laevigatum Linne, 1758.
pallida I-um (subspecies, variety) - See under
norvegicum paUida Jeffreys, 1863. - norvegicum
pallidum Jeffreys.
pallidum Reeve, 1845. [Figs. Part 1 p. 141. Part 3 p.
928]. - Synonym of Fulvia papyracea (Bruguiere,
1789). OTHER JUDGEMENTS: Fulvia tenuicostata
(Lamarck, 1819). » Regarding Cardium pallidum:
Cardiidae. Recent. TYPE REPOSITORY. 2 possible
syntypes. 1978134. BMNH London. TYPE LOCALITY.
Manila Bay, Philippines . RECORDS. 001844 Reeve:
Cardium pallidum. 001884 Grasset: Cardium pallidum.
Rv. 001885 Smith: Cardium pal/idum (Reeve in Romer,
fig. 92) = Cardium (Bucardium) tenuicostatum
Lamarck. 001890 Paete1: Cardium pallidum Rv. 001903
Pritchard & Gatliff: Cardium pallidum Romer in Kuster
1869 = Cardium tenuicostatum Lamarck. 001906
Melvill & Standen, p. 839: Cardium (Acanthocardia)
pallidum Reeve. 001922-32 Sherborn, p. 4701: Cardium
pallidum Reeve, 1845 . 001928 Faustino: Cardium
pallidum Reeve. 001945 Smith: Cardium pallidum
Reeve 1843 syn. = Cardium tenuicostata Lamarck
1819. 001977 Fischer-Piette: Cardium pallidum Reeve
= Papyridea papyraceum (Chemnitz) Schroter. 001992
Lamprell & Whitehead: Cardium pallidurn Reeve, 1844
= Fulvia tenuicostata (Lamarck, 1819). 001994 Vidal:
? Cardium pallidum Reeve, 1845 = Fulvia (Fulvia)
papyracea (Bruguiere, 1789). - CHARACTERS AND
OCCURRENCE - DESCRIPTIONS 001845 Reeve: shell
somewhat squarely ovate, rather depressed, posteriorly
a little angularly produced; radiately finely ribbed, ribs
very numerous, sometimes 42-43, sometimes 52-53,
Phukel Marine Biological Center Special Publication vol. 30 (2004)
673
palm alum - panamense
smooth, somewhat keel-shaped; pale. REMARKS. ··1845
Reeve: This is another species, which has been probably
confounded with the Cardium tenuicostatum, an
examination of several specimens assures me, however,
that it is distinct. It is more depressed, has, not the
slightest indication of the waved flesh-colour spots,
which adorn that species, and is altogether different.
HABITAT. ··1845 Reeve: found in sandy mud at a depth
of from 4 to 5 fathoms . DISTRIBUTION. ··1845 Reeve:
Bay of Manila, the Philippines. ··1884 Grassel: Manila.
palmalum Goldfuss, 1837. - Combination to be revised:
"Cardium" palmalum Goldfuss, 1837. Cardiidae. PreMesozoic. RECORDS. "1922-32 Sherborn, p. 4714:
Cardium palmatum Goldfuss, 1837. "1999 Jacques
Vidal (pers. com.), MNHN, Paris: Fossils from the
Paleontological Collection. Cardium palma tum
Goldfuss. Germany, B03948.
palmerae Harbinson, 1944. - Combination to be revised:
"Cardium " palmerae Harbinson, 1944. Cardiidae.
Eocene. RECORDS. ··1944 ZR : Cardium palmerae
Harbinson, 1944 sp. p..ov. (Eocene) Santee-Cooper
Canal 17 m. N.- W. of Moncks Comer, South Carolina.
paiudosllm (variety). - See under glaucum paludosa
Bucquoy, Dautzenberg & Dollfus, 1892.
panamense Dall, 1908. - Nemocardium panamense
(Dall, 1908). OTHER JUDGEMENTS: suggested new name
Microcardium williami. » Regarding Protocardia
panamensis: Cardiidae. Recent. REcORDs. ··1908 Dall:
Protocardia panamensis. ··1961 Olsson: Microcardium
panamense. ··1971 Keen & McLean: Nemocardium
(Microcardium) panamense (Dall, 1908). "1977
Fischer-Piette: Protocardia panamensis Dall. ··1977
ZR : Microcardium panamensis (Dall, 1908)
Preoccupied name Replaced by Microcardium williami.
Ref. to Fischer-Piette, E. p . 4. ··1980 Keen .
Nemocardium panamense (Dall, 1908) . CHARACTERS AND OCCURRENCE - REMARKS.
··1977 Fischer-Piette, p. 4: Microcardium williami new
name for Microcardium panamense Dall, 1908.
However, Keen (1980) claims that Microcardium
williami is an unnecessary new name for a supposed
homonym: Protocardia panamensis Dall, 1908, versus
Cardium panamense Sowerby, 1833, species that now
fall not only in different genera but in different
subfamilies. HABITAT. ··1971 Keen & McLean: 333 m.
DISTRIBUTION. ··1971 Keen & McLean: Panama.
panamense Sowerby, 1833. [Figs. Part I p. 109, 134.
Part 3 p. 857, 924). - Changed combination: Trachycardium panamense (Sowerby, 1833). OTHER JUDGEMENTS: Trachycardium procerum (Sowerby, 1833). »
Regarding Cardium panamense: Cardiidae. Recent.
TYPE LOCALITY. Ad Panamam in sandy mud at a depth
of 10 fathoms (Sowerby 1833). TYPE REPOSITORY.
BMNH; Cardium panamense Sowerby Holotype reg.
no. 1991024. Neg. No. 0629. RECORDS. ··1833 S0tlierby,
p. 85: Cardium panamense. "1834 Sowerby, 49 t part,
fig. 21: Cardium panamense nobis. ··1840 Sowerby,
catalogue p. 5, No. 62: Cardium panamense, Sow.
"1840 Sowerby, corrected list, p. 8 : Cardium
panamense (Sowerby No. 62). ··1844 Reeve, sp. 56:
Cardium panamense Sowerby, 1843 sic. ··1858 Adams
& Adams, p . 455: Cardium (Trachy cardium)
Panamense Sow. ··1860 Carpenter: Cardium
panamense Sby. ··1869 Romer: Cardium Panamense
Sowerby fig. 21 Uuvenile) = Cardium procerum
Sowerby. "1890 Paetel: Cardium panamense Sow. =
procerum Sow. juv. ··1891 Steams: C. Panamense Sby.
(probably a growth stage of Cardium procerum Sby.
"190 I Dall: Cardium panamense Sowerby, 1833 =
Cardium procerum Sowerby, 1833. "1909 Lamy:
Cardium panamense Sowerby (1833) = Cardium
procerum Sowerby. ··1910 Dall: Cardium panamense
Sowerby = Cardium procerum Sowerby (p. 282).
··1922-32 Sherborn, p. 4727: Cardium panamense
Sowerby, 1833. "1936 Jordan: Cardiumpanamense
Sowerby, 1833 = Cardium procerum Sowerby, 1833.
"1947 Hertlein & Strong: Cardium panamense
Sowerby, 1833 = Cardium (Mexicardia) procerum
Sowerby. ··1959 Ranson, p. 71 : Cardium panamense
Sow = Cardium procerum Sowerby. ··1964 Parker:
Trachycardium (Mexicardia) panamense (Sowerby,
1833) ... 1971 Keen & McLean: Trachycardium
(Mexicardia) panamense (Sowerby, 1833). "1977
Fischer-Piette: Cardium panamense Sowerby =
Laevicardium procerum Sowerby. "1995 Poutiers:
Trachycardium panamense (Sowerby, 1833). "1999
Vidal: Trachycardium (Mexicardia) panamense
(Sowerby in Broderip & Sowerby, 1833) . CHARACTERS AND OCCURRENCE - OF COLLECTIONS. ··ZMUC : Gulf of California. ··ZMUC: I
specimen. Philippines. ··ZMUC: I specimen. Panama;
II shells from Realejo (Oersted coil. No. 326)
panamanense Sowerby, 1833 ··ZMUC : Gulf of
California 13 m, shelly sand. Sowerby, 1843 sic.
··ZMUC: wet collection: Gulf of California, 13 m
muddy sand. DESCRIPTIONS. ··1844 Reeve: shell rather
elongately ovate, ventricose, attenuated towards the
umbones, anterior side rounded , posterior rather
angulated; radiately ribbed , about 20 large,
conspicuous, ribs, first angulated, then rounded;
whitish, covered with an olive epidermis, posterior
margins purple within. ··1971 Keen: there seem so
many points of difference between this and the
generally more southern form, Trachycardium
procerum, that the two are here considered as separate
species. The shell is much more massive and may attain
a height of 90 mm, compared with 55 mm for the
southern one. The ribs are triangular in outline, with
fine nodes along the sharp crest, especially on the
umbones, and the rather wide spaces between ribs are
flat. The shell is white under a greenish, persistent
periostracum, except for spots of brown on the
umbones. The species is a common one in mud, from
low-tide line to a depth of several fathoms, ranging
from San Ignacio Lagoon, Lower California,
throughout the Gulf of California but, in spite of its
name, apparently no farther south than Costa Rica;
common. REMARKS. "1840 Sowerby: ribs fewer and
more angulated than in C. procerum. ··1844 Reeve:
this is a truly noble species, very closely allied to the
Cardium procerum, but easily distinguished by its more
elongated form, and by the marked development of the
ribs. HABITAT. ··1844 Reeve: found in sandy mud at
the depth of about 10 fathoms. ··ZMUC: sand, shelly
sand, silty sand, shelly mud. 7-67 m depth. "1964
Parker: occasionally to a depth of about 100 m.
DISTRIBUTION. "1844 Reeve: Panama. "1884 Grasset:
Panama . ··1971 Keen & McLean: San Ignacio Lagoon
674
Tropical Marine Mollusc Programme (TMMP)
panamensis - papillosum
in Baja California to Costa Rica, not reaching Panama.
··ZMUC Gulf of California, Realejo.
panamensis (mode). - See under panamense Dall, 1908.
pancicostatum (error). - Printer's mistake for
paucicostatum in ZR for 1937 with ref. to Keen [937.
pancicostatum eburnea (error). - Printer 's mistake for
paucicostatum eburnea in Coen, [914.
panciotermedia (error) . - Printer's mistake for paucintermedia.
paniselense - Combination to be revised: "Cardium "
paniselense Cardiidae. Fossil. RECORDS . •• [932 ZR:
Cardium paniselense [Ref. Vincent 1932].
panis-sacchari (invalid spelling). - See under
panissacchari.
panissacchari Altena, 1961. - Trigoniocardia
panissacchari Aitena, 1961. Cardiidae. Pleistocene.
RECORDS. ··1961 ZR: Trigoniocardia (Trigoniocardia)
panissacchari Aliena, 1961 sp. nov. St. Eustatius Sugar
Loaf, presumably Pleistocene, Leeward Islands.
pannanicum Vodasz, 1910. - Combination to be revised:
"Cardium" pannanicum (Vodasz, 1910). Cardiidae.
Eocene. RECORDS. ··1910. ZR: Cardium (Trachy cardium) pannanicum Vodasz, 1910 sp. n. Hungary
Eocene.
pannonica Stevanovic, 1979. - Paradacna pannonica
Stevanovic, 1979. Lymnocardiidae. Upper Miocene.
RECORDS. ··1979 ZR: Paradacna pannonica
Stevanovic, 1979. Sp. nov. Yugoslavia. Ref. to Stevanovic, p. 322.
Pannonicardium Stevanovic 1951 (genus) .
Lymnocardiidae. Tertiary and Recent. RECORDS. ··1980.
Vokes: Pannonicardium Stevanovic 1951 in subfamily
Lymnocardiinae.
Pannonicardium Stevanovic 1951 (subgenus) .
Lymnocardiidae. Pontian . RECORDS. ··1953 ZR :
Pannonicardium subgen. nov. of Limnocardium type
Limnocardium dumieici (Gorj. Kramb) Pontian Serbia
[Ref. Stevanovic 1951] . ··1987 Taktakishvili :
(Pannonicardium) Stevanovic, 1951.
pansatrum . - Combination to be revised: "Cardium "
(Cerastoderma) pansatrum. ··2002 Schneider citing
Dall (1900): Cardium (Cerastoderma)pansatrum sp.
nov. section Cerastoderma .
pantecolpatum Cossmann & Peyrot, 1911. - Changed
combination: Clinocardium pantecolpatum (Cossmann
& Peyrot, 1911). Cardiidae. Neogene. RECORDS. ··1911
ZR: Cardium (Laevicardium) pantecolpatum Cossmann & Peyrot, 1911 sp. n. France Neogene. ··1952
Lecointre: Cardium (Laevicardium) pantecolpatum
Cossmann et Peyrot, 1912. ··1970 G[ibert & Poel:
Cardium (Laevicardium) pantecolpatum Cossmann &
Peyrot, 1909-1912 . ··1977 Popov : Clinocardium
pantecolpatum (Cossmann & Peyrot, 1911).
Panticapaea Andrussow 1923 (genus).
Lymnocardiidae. Pliocene. RECORDS. ··1977 Popov:
Panticapaea Andrussow, 1923. ··1980 Vokes:
Panticapaea Andrussow, 1923 in subfamily Paradacninae . •• [987 Taktakishvili, p. [88: designated
Panticapaea Andrussow 1923 type genus of the new
tribe Panticapaeini . ··200 I Schneider & Carter:
Panticapaea (genus). Lymnocardiidae. Pliocene.
panticapaea R. Hoem. - Synonym of Didacna
tamanensis R. Hoem . » Regarding Didacna
panticapaea: Lymnocardiidae. Fossil. RECORDS. ··1987
Taktakishvili: Didacna panticapaea (R. Hoem.) (=
Didacna tamanensis R. Hoem.). ··1987 Taktakishvili:
Pontalmyra panticapaea (R. Hoern.).
panticapaea (cf.) R. Hoem. - RECORDS. ··1987
Taktakishvili: Pontalmyra cf. panticapaea (Andrus .).
panticapaea gurianthica Tschel. (subspecies) . Pontalmyra panticapaea gurianthica (Tschel.) .
Lymnocardiidae. Fossil. RECORDS. •• [987 Taktakishvi[i:
Pontalmyra panticapaea gurianthica (Tschel.).
Panticapaeini Taktakishvili , 1987 (tribus). Lyrnnocardiidae. REcORDS. •• [987 Taktakishvili, p. 188:
Panticapaeini; tribe nov. ofLirnnocardiinae. Type genus
Panticapaea Andrussow, 1923.
Papillicardium Monterosato in Sacco, [ 899 (genus). Cardiidae. Middle Miocene to Recent. Subgenus
Papillicardium upgraded to generic level by authors.
RECORDS . •• [969 Moore et al.: Genus Papillicardium .
•• [979 ZR: Papillicardium F. Sacco [899. New
synonym of Parvicardium Monterosato 1884. Ref. to
Lambiotte, p. 46. ··1980 Vokes: Papillicardium
(Monterosato MS) Sacco in Bellardi & Sacco, [899.
In subfamily Cardiinae . ··2002 Schneider:
Papillicardium is considered basal fragines. ··1998
Schneider (1998b): Papillicardium. Junior synonym of
Parvicardium (Glibert & van de Poel, 1970; Popov,
1977; Lambiotte, 1979). ··1998 Schneider (l998b):
Papillicardium. Junior synonym of Plagiocardium
(Spada & Della BelJa, 1987). ··1998 Schneider:
Papillicardium. Type species Papillicardium
papillosum (Po Ii, 1791). Cardiidae. Middle Miocene
to Recent. (The genus originates in the Eocene; Keen
1980). ··2002 Schneider: citing Kafanov and Popov
(1977) : Papillicardium subfamily Fraginae, tribe
Parvicardiini.
Papillicardium Monterosato in Sacco, 1899 (subgenus).
- Cardiidae . Fossil and Recent. Type species :
Monterosato in Sacco (1899) designated Cardium
papillosum Poli, 1795 type species of subgenus
Papillicardium of genus Plagiocardium Cossmann,
1886. The subgenus (Papillicardium) was subsequently
raised to generic level. According to Sacco (1899):
Subgenus Papillicardium Montrs. 1899 in lilt. RECORDS.
··1899 ZR: Papillicardium (Monts. MS.) Sacco, 1899
n. sg. of Cardium for Cardium papillosum Polio ··1899
ZR: Cardium papillosum in sg. Papillicardium [Ref.
Sacco 1899] . ··1998 Schneider (I 998b) : Papillicardium. Subgenus of Parvicardium (Keen, 1969a,
1980; Kafanov & Popov, 1977; Voskuil & Onverwagt,
1989). - CHARACTERS AND OCCURRENCE CHARACTERS of diagnostic value: shell small, resembling
Parvicardium but with heavier hinge. With beaded
rather than spinose sculpture on ribs; on posterior ribs
in the form of semispatulate nodules. (HyUeberg 1994).
PapiIlocardium (error). - Printer's mistake for
Papillicardium in Oliver 1992.
*papi/losum. - Homonymy to be noticed. Cardium
papillosum from 3 sources: Chiaje, 1830. - Poli, 1791.
- Goldfuss (?year).
papillosum Chiaje, 1830. - Combination to be revised:
"Cardium " papillosum d. Chiaje, 1830. Cardiidae.
Fossil. RECORDS. ··1922-32 Sherborn, p. 4739: Cardium
papillosum d. Chiaje, 1830.
papi/losum Poli, 1791. [Figs. Part 1 p. 81 , 94, 121, 143,
173 . Part 3 (comparisons) p. 915, 933] . Papillicardium
Phukel Marine Biological Cenler Special Publicalion vol. 30 (2004)
675
papillosum - papil/osum depressum
papillosum (Poli, 1791). OTHER JUDGEMENTS: Cardium
punctatum Brocchi. - Cardium planatum (Renieri). Cardium poW Payraudeau. » Regarding C.
papillosum: Cardiidae. Miocene to Recent. Sacco
(1899) designated Cardium papi/losum Poli, 1795 type
species of the subgenus (Papillicardium) Sacco, 1899.
Subsequently raised to generic status. TYPE LOCALITY.
The Mediterranean. RECORDS. "1791 Poli, I, p. 56-57
(pI. 16, figs. 2-4): Cardium papillosum. "1840
Sowerby, I 84 th part, fig. 69: Cardium papillosum Poli,
1795. "1840 Sowerby, catalogue p. 3, sp. 37: Cardium
papillosum Poli. "1840 Sowerby, corrected list, p. 8:
Cardium papillosum (Sowerby, No. 37). "1844 Forbes:
Cardium papil/osum Poli o "1845 Reeve, sp. Ill , fig.
105: Cardiumpapillosum. "1848 Requien : Cardium
Papillosum Poli = Cardium punetatum Brocchi. tab.
16, fig. 11. "1864 Mayer: Cardium papillosum . Poli .
'·1867 Weinkauff, p. 138: Cardium papillosum Poli o
Weinkauffhas a very long list of synonyms, both Recent
and fossil occurrences. "1878 Monterosato: Cardium
papillosum Poli = C. plana tum (Renieri). "1885 Smith:
Cardium (Acanthocardia) papi/losum Poli o "1888
Servain: Cardium papil/osum Poli , 1791 . "1888
Simroth: Cardium papillosum Polio "1889 Martens :
Cardium papillosum Polio "1890 Paetel : Cardium
papillosum Polio "1892 Locard: Cardium papillosum
Poli, 1791. "1894 Jousseaume : Cerastoderma
papillosum (Dollfus & Dautzenberg 1912). "1899
Sacco: Papillicardium papil/osum (Poli 1791). "1900
Locard & Caziot: Cardium papil/osum Poli, 1791 .
"190 I Chaster et al.: Cardium papillosum Pol i. "1902
Sherborn, p. 716: Cardium papil/osum Poli, 1795 .
"1907 ZR: Cardium (Papillicardium) papi/losum [Ref.
Martinelli 1907]. "1927 Dautzenberg: Cardium
(Parvicardium) papillosum Polio "1931 ZR: Cardium
papillosum Ref. to Patrini (1931). "1932 Winckworth:
Cardium (Parvicardium) papillosum Poli, 1795 . "1933
Coen: Eucardium (Papillicardium) papillosum Poli .
"1933 Coen & Vatova: Eucardium (Papillieardium)
papillosum Polio "1937 Coen : Papillicardium
papillosum Poli . "Scarlato & Starobogatov: Plagiocardium papillosum (Poli, 1791). "1956 Rasmussen:
Coreulum (Papillieardium) papillosum (Pol i). "1958
Holzl: Coreulum (Parvieardium) papillosum (Poli).
"1962 Pasteur-Humbert : Cardium (Corculum)
papillosum Polio "1969 Moore et al.: Plagiocardium
(Papillieardium) papillosum (Poli , 1795). "1969
Nordsieck: Papillieardium papillosum (Poli, 1791).
"1970 Glibert & Poel: Parvieardium papillosum (Poli,
1791). "1974 Barnard: Cardium papillosum in Jaeckel
1931 = Cardium turtoni . "1977 Fischer-Piette :
Coreulum (Papillieardium) papi/losum Polio "1978
Janssen & Van der Slik: Parvieardium papillosum (Poli,
1795). "1978 Janssen & Van der Slik: Plagioeardium
(Papillieardium) papillosum (Po Ii). "1983 SalviniPlawen: Coreulum papil/osum (Poli). "1992 Oliver:
Papilloeardium sic papillosum (Poli, 1795). "1998
Macedo et al.: Parvieardium papillosum (Poli, 1795).
"1998 Schneider (1998b): Papillieardium papillosum
(Poli, 1795). "1998 Schneider: Papillieardium. Type
species Papillieardium papillosum (Poli, 1791).
Cardiidae. Middle Miocene to Recent. "1999 Jacques
Vidal (pers . com.), MNHN, Paris : Fossils from the
Paleontological Collection as Parvieardium papillosum
(Poli). Helvetien. France; Italy; Turkey; Morocco.
.. 2000 Aartsen & Goud: Parvieardium papil/osum
(Poli, 1795). "2001 Vidal, p. 408: Parvieardium
papillosum. - CHARACTERS AND OCCURRENCE
- OF COLLECTIONS. "ZMUC: Atlantide Exp. St. 153,
145, 231,44, 147, 123,45. "ZMUC: wet collection:
Neapel, Mediterranean. "ZMUC: West Africa.
DESCRIPTIONS. "1845 Reeve: shell somewhat orbicular,
convex, a little oblique; radiately ribbed, 24 ribs,
echinated throughout with short papillae; interstices
between the ribs transversely impressly punctured;
whitish, stained and sparingly spotted with rusty brown,
interior white, reddish brown on the posterior side.
"1888 Simroth: "Bunt dunckelbraun gefleckt, einige
StUcke innen mit amarantrothen Flecken oder mit zwei
Strahlen". "2001 Vidal: only the exhalant aperture is
tube-shaped. Inhalant siphon open towards the foot
aperture. No eyes. REMARKS. "1845 Reeve: a wellknown species, which may be easily recognised by the
papilliferous sculpture of the ribs, and the peculiar
transverse groove-like punctures in the interstices .
HABITAT. "1844 Forbes: Aegaean Sea depth range 675 fathoms. "1889 Martens: South of Madeira, 109128 m, mixed bottom. DISTRIBUTION. "1845 Reeve:
Mediterranean. "1884 Grasset: Sicily. "1845 Reeve:
Mediterranean. "1969 Nordsieck, "1994 Hylleberg :
West Europe to Canary Islands, Mediterranean, Red
Sea ... ZMUC: Mediterranean, West Africa. "1992
Oliver: Suez.
papil/osum Goldfuss. - Replaced by combination to be
revised: "Cardium" seobinula Merian. » Regarding
Cardium papillosum: Cardiidae. Oligocene. RECORDS.
"1892 B. D. & D., p. 277: Cardium papillosum
Goldfuss differs from papillosum (Poli, 1791). The
papillosum of Gold fuss has become Cardium seobinula
Merian. "1902 Dollfus & Dautzenberg: Cardium
papillosum Goldfuss (non Poli).
papillosum Mac Andrew (variety). - Parvieardium
seabrum (Philippi, 1844). Cardiidae. Recent. RECORDS.
"1890 Hidalgo: Cardium papillosum var. of Mac
Andrew = Cardium nodosum Montagu.
papillosum A Scacchi, 1836 (variety). - Infrasubspecific.
» Regarding Papillieardium papillosum var. a:
Cardiidae. RECORDS. "1836 Scacchi : Cardium
papillosum var. a penitus papillosa.
papi/losum alba Locard & Caziot, 1900 (variety). Infrasubspecific. No evidence of sub specific status. »
Regarding Cardium papillosum var. alba : Cardiidae.
Recent. TYPE LOCALITY . Mediterranean. RECORDS. "1900
Locard & Caziot: Cardium papillosum alba (var.).
papillosum aurea Bucquoy, Dautzenberg & Dollfus,
1892 (variety). - Infrasubspecific. No evidence of
subspecific status. » Regarding Cardium papillosum
var. aurea: Cardiidae . Recent. RECORDS . "1892
Bucquoy, Dautzenberg & Dollfus, p. 277 : Cardium
papil/osum var. (ex colore I) aurea. "1900 Locard &
Caziot: Cardium papil/osum var. aurea (B.D. & D.).
papil/osum B Scacchi, 1836 (variety). - IntTasubspecific.
» Regarding Papillieardium papillosum var. b:
Cardiidae. RECORDS. "1836 Scacchi: Cardium
papillosum var. b semi- papillosa.
papi/losum depressum Locard, 1898 (variety) . Infrasubspecific. No evidence of sub specific status. »
Regarding Papillieardium papillosum vai:. depressum:
676
Tropical Marin e Mollusc Programme (TMMP)
papillosum dertonense - papillosum transversa
Cardiidae. Recent. REcORDs. ··1898 Locard, p. 262:
Cardium papil/osum var. depressum . ··1969 Nordsieck:
Papillicardium papillosum (Poli, 1791) var. depressum
(Locard). - CHARACTERS AND OCCURRENCE DESCRIPTIONS. ··1898 Locard: often somewhat oblique
outline, depressed; very small.
papil/osum dertonense Michelloti, 1847 (variety). Infrasubspecific. No evidence of subspecific status. »
Regarding Papillicardium papi/losum var. dertonensis:
Cardiidae. Recent. RE CORDS . . . 1899 Sacco:
Papillicardium papi/losum var. dertonensis Michelotti
1847(an species distinguenda). "1902 Dollfus &
Dautzenberg: Cardium papillosum var. dertonense
Michelloti .
papil/osum globulosum Locard, 1898 (variety). Infrasubspecific. No evidence of subspecific status. »
Regarding Cardium var. gfobulosum: Cardiidae.
Recent. RECORDS. "1898 Locard, p. 263 : Cardium
globulosum. ··1969 Nordsieck: Papillicardium
papillosum (Poli, 1791) var. globulosum (Locard).
papillosum lactea Jeffreys, 1863 (variety) . Infrasubspecific. No evidence of subspecific status. »
Regarding Cardium papillosum var. lactea: Cardiidae.
Recent. RECORDS. ··190 I Chaster et al.: Cardium
papillosum var. lactea Jeffreys. ··1980 Ruhoff:
Cardium papillosum lactea Jeffreys, 1863 ; [18621869] vol. 2: 276.
papi/losum maculatum Danilo & Sandri, 1891 (variety).
- Infrasubspecific. No evidence of subspecific status.
» Regarding Cardium papil/osum var. maculatum :
Cardiidae. Recent. RECORDS. ··1891 Danilo & Sandri:
Cardium papillosum Poli var. maculatum nob. ··1892
Bucquoy, Dautzenberg & Dollfus, p. 277: Cardium
papillosum var. (ex colore 2) maculata. ··1969
Nordsieck: Papillicardium papillosum (Poli, 1791) var.
maculatum (Brusina) . - CHARACTERS AND
OCCURRENCE - REMARKS. Brusina wrote the
introduction to Danilo & Sandri (1891). DISTRIBUTION.
Unknown (Nordsieck 1969).
papillosum maxima Bucquoy, Dautzenberg & Dollfus,
1892 (variety). - Infrasubspecific. No evidence of
subspecific status. Papillieardium papillosum (Poli,
1791). » Regarding Cardium papillosum var.
maximum: Cardiidae. Recent. RECORDS. ··1892
Bucquoy, Dautzenberg & Dollfus, p. 276 : Cardium
papillosum var. ex forma 2 maximum . ··1927
Dautzenberg: Cardium papi/losum var. maxima,
Bucquoy, Dautzenberg et Dol1fus = Cardium
(Parvicardium) papillosum Poli . "1969 Nordsieck:
Papillicardium papillosum (Poli, 1791) var. maximum.
- CHARACTERS AND OCCURRENCE - DESCRlPTlONS. ··1892 Bucquoy, Dautzenberg & Dollfus: very
thick shell, and large size compared with the
Mediterranean specimens. REMARKS. Described as a
variety. DIMENSIONS. Umbo-ventral 25 mm, anteriorposterior 25 mm. HABITAT. ··1969 Nordsieck: 40-1494
m depth. DISTRIBUTION. "1892 Bucquoy, Dautzenberg
& Dollfus: The Azores.
papillosum minor Locard , 1898 (variety). Infrasubspecific. No evidence of subspecific status. »
Regarding Cardium papillosum var. minor: Cardiidae.
Recent. RECORDS . .. 1898 Locard, p. 263 : Cardium
papillosum var. minor. ··1900 Locard & Caziot:
Cardium papillosum var. minor. ··1969 Nordsieck:
Papillicardium papillosum (Poli , 1791) var. minor
(Locard) Distribution unknown.
papillosum obliquatum Michelotti, 1839 (variety). Infrasubspecific. No evidence of subspecific status.
Cardiidae. Fossil. REC ORDS. "1902 Dollfus &
Dautzenberg: Cardium papil/osum var. obliquatum
Michelotti, 1839.
papillosum obliquatum Monterosato, 1872 (variety).
Infrasubspecific. No evidence of subspecific status.
OTHER JUDGEMENTS : Cardium laevigatum. » Regarding
C. papillosum var. obliquatum: Cardiidae. Recent.
RECORDS. ··1872 Monterosato (p. 39): C. papillosum,
var. obliquatum = Cardium laevigatum. ··1892
Bucquoy, Dautzenberg & Dollfus: Cardium papillosum
var. exforma I obliquata (Aradas) Monterosato. ··1893
Marshall: Cardium papi/losum var. obliquata Monterosato. ··190 I Chaster et at.: Cardium papilloswn var.
obliquata Monterosato. ··1902 Dollfus & Dautzenberg:
Cardium papi/losum var. obliquata Aradas 1846 (non
var obliquatum Michelotti, 1839) = Cardium aradasi
Crema. ··1926 McClelland: Cardium papillosum V.
obliquata Monts. ··1969 Nordsieck: Papillieardium
papillosum (Poli, 1791) var. obliqllatum (Monterosato,
1884).
papillosum pertransversa Sacco, 1899 (variety). Infrasubspecific. No evidence of subspecific status.
Cardiidae. Recent. RECORDS. "1899 Sacco, p. 45, pI.
11, figs. 6,7: Papillicardium papillosllm var.
pertransversa Sacc.(an species distinguenda). "1902
Dollfus & Dautzenberg: Cardium papi/losum var.
pertransversa Sacco.
papillosum roseum Pallary (variety). - Synonym of
Papillicardium papi/losum (Poli, 1791. Cardiidae .
Recent. RECORDS. ··1969 Nordsieck : Papillieardium
papillosum (Poli, 1791) var. ros eum (Pallary).
Distribution unknown.
papillosum rotundum Locard, 1898 (variety). Infrasubspecific. No evidence of subspecific status.
Cardiidae. Recent. DESCRIPTIONS. Rounded outline,
nearly symmetric (Locard 1898). RECORDS. ··1898
Locard, p. 263 : Cardium papillosum var. rotundllm.
"1969 Nordsieck: Papillicardium papillosum (Poli,
1791) var. rotundum (Locard). Distribution unknown.
papil/osum senegalensis Dollfus, 1911 (variety). Infrasubspecific. No evidence of subspecific status.
Cardiidae. Recent. RECORDS. ··1911 Dollfus, p 59:
Cardium papillosum var. senegalensis.
papillosum senegalensis Rosso, 1974. - ?Corculum
papillosum senegalensis Rosso, 1974. Cardiidae.
Quaternary. RECORDS. ··1974 ZR: Careufum sic
papillosum senegalensis Rosso, 1974 subsp . nov.
Senegal, Tongon. Quaternary. Rosso p. 811.
papil/osum simile Mi1aschewitch , 1909 (variety).
Identical with Plagiocardium simile (Milaschewitch,
1909). » Regarding Cardium papillosum var. simile:
Cardiidae. Recent. RECORDS. ··1969 Nordsieck:
Papillicardium papillosum (Poli, 1795) var. simile
Milaskewicz (= Milachewitch). Distribution Black Sea.
··Scarlato & Starobogatov: Cardium papillosum var.
simile Mil. = Plagiocardium simile (Milaschewitch,
1909).
papillosum transversa Cerulli Irelli (variety). - Identical
with Cerastoderma edule (Linne, 1758). » Regarding
Cardium papillosum var. transversa: Cardiidae. Fossil.
Phuket Marine Biological Center Special Publication vol. 30 (2004)
677
papilfosum zonata - papyraceurn
RECORDS. --1902 Dollfus & Dautzenberg: Cardium
papillosum var. transversa Cerulli Irelli = C. edule var.
papillosum zonata Locard & Caziot, 1900 (variety). Infrasubspecific. No evidence of subspecific status.
Cardiidae. Recent. RECORDS. --1900 Locard & Caziot,
p. 236: Cardium papillosum var. zonata. Mediterranean.
papioiensis (variety). - See under edule papiolensis
Almera & Bofill.
pappi Strausz, 1930. - Lymnocardium pappi Strausz,
1930. Lymnocardiidae. Miocene. RECORDS. --1941 ZR:
Limnocardium pappi Strausz, 1930 sp. nov. Miocene,
Hungary. --1987 Taktakishvili : Limnocardium pappi
Strausz.
papuanum Vidal, 1996 . [Figs. Part 3 p. 869]. Vasticardium papuanum Vidal, 1996. Cardiidae.
Recent. RECORDS. --1999. Vidal: Vasticardium
papuanum Vidal, 1996. - CHARACTERS AND
OCCURRENCE - DESCRIPTIONS. "1996 Vidal:
foundation of anterior lateral teeth is gently curved and
moderately strong (non-hooked appearance).
DISTRIBUTION. --1996 Vidal: Papua Sibuko Bay, Borneo
(USNM 239129); "1999 Vidal: Palau.
papyracea (mode). - See under papyraceum Bruguiere,
1789.
*papyraceum. - Homonymy to be noticed. Cardium
papyraceum from 2 sources: Bruguiere, 1789. - Sinz.
(?year).
papyraceum Bruguiere, 1789. [Figs. Part 1 p. 118. Part
3 p. 889, 890, 893]. = Fulvia laevigatum (Linnaeus,
1758). OTHER JUDGEMENTS: Fulvia aperta (Bruguiere,
1789). - Cardium deshayesianum Bianconi 1856. »
Regarding Cardium papyraceum: Cardiidae. Recent.
TYPE MATERlAL. Two syntypes of Cardium laevigatum
Linne present in The Linnean Society, are Fulvia
papyracea (Bruguiere, 1789) [see Vidal 1994: 99] The
largest of the two bivalved shells [46.2 x 46.5 x 31.1],
hand marked 88 by Linne (his list number in 1767) has
been selected by Vidal (1999, p. 327) as lectotype of
Cardium laevigatum Linne, 1758, senior synonym of
Cardium papyraceum Bruguiere, 1789. - Higo et at.
(1999) note: 'dans les mers des grandes Indes' ('seas
of the East Indies' ; Bruguiere took this locality from
Chemnitz in Martini & Chemnitz - he noted that he
had never personally seen a specimen. RECORDS. --1789
Bruguiere, p. 231, No. 29. Cardium papyraceum;
Chemnitz, Martini, tom. 6. pag. 190, tab. 18, fig. 184.
"1836 Bosc: Cardium papyraceum. "1840 Sowerby,
Part 181, figs. 55 and 56: Cardium papyraceum
Bruguiere. "1840 Sowerby, catalogue p. 2, No. 13:
Cardium papyraceum Brug. --1902 Sherborn, p. 716:
Cardium papyraceum Bruguiere, 1789. "1960 Biggs
& Grantier: Cardium (Papyridea) papyraceum Gmelin.
--1973 Biggs: Laevicardium papyraceum (Bruguiere,
1789). ··1977 Habe: Fulvia papyracea (Bruguiere,
1798). ·-1977 Wilson & Stevenson: Cardium
papyraceum Bruguiere, 1789; probably only one
species: Fulvia aperta (Bruguiere, 1789) which has
page priority. -·1982 Kilburn & Rippey: Fulvia
papyracea (Bruguiere, 1789). --1984 Taylor & Reid:
Laevicardium papyraceum papyracea Bruguiere, 1789.
·-1989 Bosch & Bosch: Laevicardium papyraceum
(Bruguiere, 1789). ·-1994 Bonfitto, Sabelli, Tommasini
& Herbert: Fulvia papyracea (Bruguiere, 1789); senior
synonym of Cardium deshayesianum Bianconi 1856.
"1994 Vidal: Fulvia (Fulvia) papyracea (Bruguiere,
1789). --1998 Poutiers: Fulvia papyracea (Bruguiere,
1789). "1999 Hylleberg, p. 401: Fulvia papyracea
(Bruguiere, 1789). ·-1999 Higo et at.: Fulvia papyracea
(Bruguiere, 1789). --2002 Hylleberg & Kilburn, p. 63:
Fulvia papyracea (Bruguiere, 1789). - CHARACTERS AND OCCURRENCE - DISTRIBU-TION. --1928
Kuroda, --1977 Habe: Amami Oshima. --1993 Bernard
et at.: South China Sea, Taiwan. "1999 Higo et at.:
Red Sea, Mediterranean. ·-1999 Hylleberg: Indonesia.
.. 2003 Hylleberg & Kilburn: Vietnam.
papyraceurn Chemnitz, 1782. [Figs. Part 1 p. 75 No.
184, p. 101 , 125, 165, 174. Part 3 p. 889]. Non
binominal. Not available. = Fulvia laevigatum
(Linnaeus, 1758). OTHER JUDGEMENTS : Fulvia
papyraceum (Bruguiere, 1789). - Fulvia papyracea
(Grnelin, 1791). - Cardium muticum Reeve. - Cardium
tenuicostatum Lrnk. juv. » Regarding Cardium
papyraceum : Cardiidae. Recent. TYPE LOCALITY. East
Indies (Chernnitz 1782); Japan (Sowerby 1841).
RECORDS. "1781 Gronovius, fasc. 3, pI. I, fig. 5:
Cardium papyraceum (fide Spengler 1799). --1782
Chemnitz, Tab. 18 Fig. 184: Cardium papyraceum, ex
museo Spengleriano. --1782 Chemnitz. ID by Gmelin
1791 : Cardium papyraceum t. 18. f. 184. "1788
Schroter: Cardium papyraceum . [Reference to
Chemnitz (1782) fig. 190).--1791 Gmelin: p. 3254 No.
32: Cardiumpapyraceum. --1799 Spengler: p. 41, No.
25: Cardium papyraceum Chemnitz Tom. 6. Tab. 18.
fig. 184 . --1817 Dillwyn: Cardium papyraceum
Chemnitz. -·1819 Lamarck: p. 6 No . 9: Cardium
papyraceum. -·1839 . Anton: Cardium papyraceum
Chemnitz ... 1840 Sowerby, 181 st part, figs. 55-56:
Cardium papyraceum. --1840 Sowerby, corrected list,
p. 8: Cardium papyraceum (Sowerby, No. 13). "1844
Reeve, sp. 9: Cardium papyraceum Chemnitz, Conch .
Cab. vol. vi. p. 190. pI. 18. f. 184 "1858 Adams &
Adams, p. 457: Cardium (Fulvia) papyracea Chern.
··1867 Angas: Cardium (Papyridea) papyraceum
Chern. "1867 ZR: Cardium papyraceum (Chemn.) =
muticum (Reeve) Hakodate [Ref. Schrenck 1867].
--1869 ZR: Section, subtypical-7. Papyraceae: bullata
(L.), papyracea (Chemn.), &c. [Ref. Graham-Ponton
1869]. ·-1880 Martens: Cardium (Papyridea)
papyraceum Chemnitz = C. laevigatum Linne
according to Hanley. --1886 Cooke p. 97: Cardium
papyraceum Chemn. --1890 Paetel: Papyridea
papyracea Ch. ··1891 Romer: Fulvia papyracea Ch.
--1897 Sowerby: Cardium papyraceum Chemnitz.
··1902 Sherborn, p. 716: Cardium papyraceum Gmelin,
1790 sic. --1815 Wood, p. 230: Cardium papyraceum
Linn, Gmelin . p. 3254. --1903 Hidalgo: Papyridea
papyraceum . --1906 Melvill & Standen, p. 839:
Cardium (Papyridea) papyraceum Chemn. --1909
Lynge: Cardium (Papyridea) papyraceum. -·1933 ZR:
Cardium papyraceum Gm. [Ref. Gruvel 1933]. "1937
Viader: Cardium papyraceum Chemnitz = C.
laevigatum Linne. --1945 ZR: Cardium papyraceum
Chern. [Ref. Lamy 1945] . --1960 Biggs & Grantier:
Cardium (Papyridea) papyraceum Gmelin. "1977
Fischer-Piette: Papyridea (Fulvia) papyraceum
(Chemnitz) Schroter. --1984 Sharabati: Laevicardium
(Fulvia) papyraceum. "1992 Oliver: Fulvia papyracea.
--1992 Oliver: Cardium papyraceum Chemn. in Cooke
678
Tropical Marine Mollusc Programme (TMMP)
papyraceum - Papyridea
1886 = tenuicostatum Lmk. juv. - CHARACTERS
AND OCCURRENCE - OF COLLECTIONS. ··ZMUC:
Gulf ofThailand, I fathom, sand. ··ZMUC: I specimen
(the true one) in the Spengler coIl. sine loco Possibly
syntype from Chemnitz [not available] according to
note written by Vidal 1994. ZMUC wet coil. sine loc.
DESCRIPTIONS. ··1799 Spengler: testa cordata, jragili,
membranacea, longitudinaliter obsolete striata,
cinerea, maculis purpureis internae cavitatis extus
pellucentibus margine subtilissime crenulato. My
translation: Shells thin, umbo-ventral distance longer
than anterior-posterior distance, central umbones,
posterior area not expanded as a wing, shells only
slightly gaping, ribs raised posterior and anterior, but
low centrally. ··1844 Reeve: Shell depressly heartshaped, thin, fragile, anteriorly very minutely
granulated, radiately impressly striated, striae rudely
engraved, giving root to an epidermis; whitish or
yellowish, faintly tinged here and there with purple,
umbones reddish purple, interior reddish purple.
DIMENSIONS. ··1799 Spengler: height 44 mm (I Tom. 8
Lin.), width 39 mm (1.5 Tom.). ··1819 Lamarck: length
34 mm. REMARKS. ··1799 Spengler, p. 41 : the figure in
Gronovius (1781, fasc. 3, pI. I, fig. 5) should illustrate
Cardium hians, but it does not resemble that species at
all. However, it is identical with Cardium papyraceum
according to Spengler. ··1844 Reeve: The
accompanying figure, and Fig. 56. of Mr. Sowerby's
'Conchological Illustrations,' represent varieties of the
true Cardium papyraceum, and are quite distinct from
a fine new species, which has been confounded with it
in the same work, at Fig. 55. This shell, for which see
Cardium muticum, Plate VI. is of much larger
dimensions, of a more transversely ovate form, and
exhibits no indication of that curious minute granulation
which characterizes the posterior side of the Cardium
papyraceum, after the manner ofthe Cardium sinense.
HABITAT. ··1844 Reeve: found in sandy mud at the depth
of seven fathoms. ··1906 Melvill & Standen: 2-15
fathoms, mud. ··1960: Biggs & Grantier collected live
specimens buried in a mudflat, the Persian Gulf.
··ZMUC: sand 2 m depth. DISTRIBUTION. ··1844 Reeve:
East end of the Island of Luzon, Philippines. ··1856
Wood & Hanley: Indian Ocean. ··1884 Grasset; China.
··1891 Romer; East Indies. ··1994 Hylleberg; IndoPacific, Gulf of Thailand.
papyraceum Schrenck, 1867. - Misidentification of
Fulvia muticum (Reeve, 1844). » Regarding Cardium
papyraceum: Cardiidae. Recent. RECORDS. ··1867
Schrenck: Cardium papyraceum Chemn. ··1869
Lischke: Cardium papyraceum Chemnitz. in Schrenck
= Cardium muticum Reeve. ··1869 ZR: papyraceum
(Schrenck not Chemn.) = Cardium muticum (Reeve) =
japonicum (Dunker), both from Japan [Ref. Lischke
1869]. ··1890 Paetel: Cardium papyraceum Schrenk.
= muticum Rv.
papyraceum Sinz. - "Cardium" papyraceum Sinz.
Cardiidae. Sarmatian. RECORDS. ··1929 ZR: Cardium
papyraceum Sinz. Sarmatian [Ref. Kolesnikov 1929].
papyraceum Sowerby 1840 (variety). [Figs. Part I p.
118]. - Identical with Fulvia mutica (Reeve, 1844).
» Regarding Cardium papyraceum: Cardiidae.
Recent. TYPE LOCALITY. Unknown (Reeve 1844).
RECORDS. ··]840 Sowerby, 1840: Cardium papyraceum.
··1844 Reeve: Cardium papyraceum var. Sowerby,
Conch. lIIus. f. 55. = Cardium muticum = Fulvia mutica
(Reeve, 1844).
papyraceum Wood. - Identical with Fulvia tenuicostata
(Lamarck). Cardiidae. Recent. RECORDS. ··1856 Wood
& Hanley: Cardium papyraceum Wood = ?c.
tenuicostatum Lam. (Australia). ··1856 Wood &
Hanley, No. 33: Cardium papyraceum Wood (as of
Ch.).
papyraceum brevior Schrenck, 1867 (forma). Infrasubspecific. No evidence of subspecific status.
Cardiidae. Recent. RECORDS. ··1867 Schrenck: Cardium
papyraceum, auct. brevior. = Cardium papyraceum
Chernnitz forma elatior.
papyraceum depressior Schrenck, 1867 (forma). Infrasubspecific. No evidence of subspecific status.
Cardiidae. Recent. RECORDS. ··1867 Schrenck: Cardium
papyraceum Chemnitz forma depressior.
papyraceum elatior Schrenck, 1867 (forma). Infrasubspecific. No evidence of subspecific status.
Cardiidae. Recent. RECORDS. ··1867 Schrenck: Cardium
papyraceum Chemnitz forma elatior.
papyraceum aviculaeformis Kolesnikov, 1929. Combination to be revised: "Cardium" papyraceum
aviculaeJormis Kolesnikov, 1929. Cardiidae.
Sarmatian. RECORDS. ··1929 ZR: Cardiacea p. 93:
Cardium papyraceum Sinz. var. aviculaeJormis
Kolesnikov, 1929 nov. Sarmatian [Ref. Kolesnikov
1929].
Papyridea Swainson, 1840 (genus). - Cardiidae.
Miocene to Recent. Type species: Gray, 1847
designated Cardium soleniforme Bruguiere, 1789 type
of the genus Papyridea Swainson, 1840. RECORDS.
··1836 Bosc: Cardium soleniforme. ··1840 Swainson,
p. 374: Subgenus Papyridea Sw. ··1847 Gray, No. 560:
Papyridea Swainson. Type species Cardium
soleniforme. ··1857 Deshayes: Papyridea Swainson,
in Gray: syn. of Cardium. ··1858 Adams & Adams:
Genus Papyridea Swainson. ··1871 Stoliczka:
Papyridea, Swainson, 1840. Type not identified by
Stoliczka. ··1886 Cossmann: Genus Papyridea
Swainson, 1840. ··1890 Paetel, 992: Gen. Papyridea
Sw. ··1934 Johnson: Papyridea Swainson 1840. ··1953
Olsson & Harbison: Genus Papyridea Swains on, 1840.
··1961 Olsson: Papyridea Swainson, 1840. ··1977
Popov p. 49: Papyridea Swainson, 1840. Genotype:
Cardium soleniforme Bruguiere, 1789. ··1980 Vokes:
Papyridea Swainson, 1840 in subfamily Trachycardiinae. ··1995 Pacaud & Renard: Papyridea Swainson,
1840 in subfamily Trachycardiinae. ··1997 Kafanov
et al.: Cardium soleniforme Bruguiere, 1789 = Cardium
latum Born, 1780. ··1998 Schneider: Papyridea. All
alleged Eocene Papyridea are species of Parvicardium.
··1998 Schneider: Papyridea. Keen (J 969, 1980) states
that the stratigraphic range of Papyridea is Miocene to
Recent. - CHARACTERS AND OCCURRENCE DESCRIPTIONS. ··1840 Swainson: shell heart-shaped, or
transversely oval; inequilateral; the anterior side almost
always gaping representing the Pholidae. ··1997
Kafanov citing Clench & Smith 1944, p. 16: Shell
transversely oval, inequilateral, the anterior and
posterior sides always gaping. Radially and obliquely
ribbed, the ribs more or less spiny. Posterior portion of
shell longer than anterior, with the ribs of the posterior
Phuket Marine Biological Center Special Publication vol. 30 (2004)
679
Papyridea - parile
slope extending beyond the margins of the valves, so
that the ribs of opposing valves form a series of
interlocking teeth. ··1969 Moore et af. , emend.
Hy11eberg 1994: Shell longer than high, gaping at both
ends, ribs spinose; hinge short; posterior margin
notched (serrated). West Indies.
Papyridea Swainson, 1840 (section and subgenus). Cardiidae. Miocene to Recent. Raised to generic level
by authors. REMARKS. Section takes the value of
subgenus. RECORDS. ··1840 Swainson, p. 374: Subgenus
Papyridea Sw. of genus Cardium Linn. ··1869 ZR:
subgenus Papyridea (Swains.), type Cardium bulla tum
(Chernn.) [Ref. Romer 1869]. ··1887 Fischer: Subgenus
Papyridea Swainson, 1840 of genus Cardium Linne,
1758. ··1889 Dal1 : (Papyridea) Swainson. ··1 890
Paetel: 1. Sect. Papyridea Sw. ··1891 Romer: I. Sect.
Papyridea of genus Papyridea Sw. ··1922 Olsson :
(Papyridea) Swainson. ··1928 Kuroda: (Papyridea)
Swainson, 1840.
Papyridea sp. Swainson. - Invalid taxon. Cardiidae.
RECORDS. ··1847 Gray, No. 560: Papyridea sp. Swains.
Type species Cardium ringens.
Papyrocardium Gabunia 1953 (genu s ) .
Lymnocardiidae . Tertiary. RECORDS. ··1980. Vokes :
Papyrocardium Gabunia 1953 in subfamily Paradacninae.
Paradacna Andrussov, 1909 (genus). - Lyrnnocardiidae.
Pliocene and Recent. Cardium abichi Homes, 1874 is
type species of genus Paradacna Andrussov, 1909.
RECORDS. ··1927 ZR: Paradacna fossil [Ref. Krejci &
Wenz 1927]. ··1968 ZR: Paradacna from Pliocene
deposits Carpathian mts. [Ref. Pana 1968]. ··1977
Popov: Paradacna Andrussow, 1901. ··1978 ZR :
Paradacna. Pliocene. Romania. Dacian Basin, faunal
assemblages, systematic accounts. Ref. to Papaianopol,
I. ··1980 Vokes: Paradacna Andrussow, 1909 in
subfamily Paradacninae. ··2001 Schneider & Carter:
Paradacna (genus). Lymnocardiidae. Pliocene.
paradacniforme Tscheltzov, 1965 . - Avicardium
paradacniforme Tscheltzov, 1965. Lymnocardiidae.
Pliocene. REC ORDS. ··1965 ZR: Avicardium
paradacniforme Tscheltzov, 1965 sp. nov. Pliocene,
Turkmenia, USSR.
Paradacninae Ebersin, 1964 (genus) . Lymnocardiidae.
Fossil. RECORDS. ··1970 ZR: Paradacninae, Ebersin,
1964, brackish water fauna, Pontian, Jugoslavia [Ref.
Stevanovic 1970].
Paradacninae Ebersin, 1964 (subfamily) . Lymnocardiidae. Tertiary and Recent. RECORDS. ··1980
Vokes: Subfamily Paradacninae Ebersin, 1964.
*paradoxum. - Homonymy to be noticed. Cardium
paradoxum from 2 sources: Miinster, 1842. Buvignier, 1843.
paradoxum Buvignier, 1843 . - Transferred to Opis
paradoxa. Deshayes. Cardiacea. Fossil. RECORDS .
··1857 Deshayes: Cardium paradoxum Buvignier,
1843. - Opis paradoxa. Desh. ··1922-32 Sherborn, p.
4746: Cardium paradoxum Buvignier, 1843.
paradoxum Miinster, 1842. - Combination to be revised:
"Cardium" paradoxum Miinster, 1842. Cardiidae.
Fossil. RECORDS. ··1857 Deshayes: Cardium
paradoxum . Munster, 1842. ··1922-32 Sherborn, p.
4746: Cardium paradoxum Muenster, 1842.
parahillana Wade, 1926. - Protocardia parahillana
Wade, 1926. Cardiidae. Cretaceous. RECORDS. ··1926
ZR: Protocardia parahillana Wade, 1926 sp. n.
Cretaceous U.S.A. ··1963 ZR : Protocardia parahillana, first record from Delaware, Upper Cretaceous
[Ref. Richards & Shapiro 1963].
parallella Bogatschev, 1932. - Didacna parallella
Bogatschev, 1932. Lymnocardiidae . Fossil. TYPE
LOCALITY . Caspian Sea. RECORDS. ··1968 Logvinenko
& Starobogatov: Didacna paral/ella Bogatschev, 1932.
Parapachydacna Andreescu , 1974 (genus) .
Lymnocardiidae. Tertiary. RECORDS. ··1974 ZR:
Parapachyda cna Andreescu, 1974 gen . nov. of
Lymnocardiidae Type species Ly mnocardium
cobalcescui Fontannes, 1886. Ref. to Andreescu, p. 141.
··1980 Vokes: Parapachydacna Ebersin, 1959, nom.
nud. Lymnocardiidae. Tertiary.
Parapscheronia Ebersin , 1955 (genus) .
Lyrnnocardiidae. Tertiary and Recent. RECORDS. ··1980
Vokes: Parapscheronia Ebersin, 1955 in subfamily
Adacninae.
parasiticum O.G . Costa, 1839 . - Synonym of
Par vicardium exiguum Gmelin , 1791. OTHER
JUDGEMENTS: Parvicardium exiguum commutata B.D.O.
» Regarding Cardium parasiticum: Cardiidae. Recent.
DISTRIBUTION. Mediterranean. RECORDS. ··1839 O. G.
Costa: Cardium parasiticum. ··1844 Philippi: Cardium
parasitici sic Costa = Cardium exiguum Gm. ··1867
Hidalgo: Cardium parasiticum Costa = Cardium
exiguum , Gmelin . ··1867 Weinkauff: Cardium
parasiticum = Cardium exiguum Gmelin. ··1869
Romer: Cardium parasiticum Costa = Cardium
exiguum Gmelin. ··1870 Aradas & Benoit: Cardium
parasiticum Costa = Cardium exiguum, Gmelin. ··1888
Kobelt: Cardium parasiticum Costa = Cardium
exiguum Gmelin. ··1890 Hidalgo: Cardium parasitica
Costa = Cardium exiguum Gmelin. ··1890 Paetel:
Cardium parasiticum Cost. = exiguum Gm. ··1902
Bellini: C. parasiticum O.G. Costa = Cardium
(Parvicardium) exiguum Gm. ··1917 Hidalgo: Cardium
parasiticum Costa = Cardium exiguum Gmelin. ··192232 Sherborn, p. 4757: Cardium parasitum sic O.G.
Costa, 1839. ··1933 Coen : Cardium parasiticum Costa
= Eucardium (Parvicardium) exiguum var. commutata
B .D.D. ··1937 Coen: parasiticum Costa =
Cerastoderma (Parvicardium) exiguum Gmelin. ··1962
Grossu: Cardium parasiticum O.G. Costa, 1839 =
Cardium (Parvicardium) exiguum Gmelin, 1790.
··2000 Aartsen & Goud: Cardium parassitum O.G.
Costa, 1839 = C. parasitum of authors, which is a
synonym of Parvicardium exiguum (Gmelin, 1791).
parasitum (error). - Printer 's mistake for parasiticum.
parazujovici Stevanovic, 1951 - New combination:
Lymnocardium parazujovici Stevanovic, 1951.
Lymnocardiidae . Fossil. RE CORDS. ··1953 ZR :
Limnocardium parazujovici Stevanovic, 1951 sp. nov.
Ponti an Serbia. ··1987 Taktakishvili: Limnocardium
parazujovici Stev.
*parile. - Homonymy to be noticed. Cardium parile
from 2 sources: Deshayes, 1858. - Dall, 1900.
parile Dall, 1900 . - Changed combination :
Trachycardium parile (Dall , 1900). Cardiidae.
Oligocene . R ECORDS. ··1898 Dall: Cardium
(Trachycardium) parile Dall, 1898 n. sp. ··1977 Popov:
Trachycardium (Trachycardium) parile (Da11, 1900).
680
Tropical Marine Mollusc Programme (TMMP)
parile - Parvieardium
--1977 Vokes: "Trachycardium" parile (Dall) not
Cardium (Trachycardium) parile in Dall 1915.
parile Deshayes, 1858 . - Changed combination:
Nemocardium parile (Deshayes, 1858). Cardiidae.
Eocene. TYPE LOCALITY. Paris Basin, France. RECORDS.
--1871 Stoliczka: Cardium parile. --1886 Cossmann:
Protocardium parile (Deshayes). --1886 Dall: Cardium
parile Deshayes. --1899 Sacco: Nemocardium parile
(Deshayes). --1904 Co ssm ann & Pissarro: Nemocardium parile (Deshayes) . --1970 Glibert & Poel:
Nemocardium (s.s.) parile (Deshayes, 1858). --1977
Popov : Nemocardium (Nemocardium) parile
(Deshayes, 1858). --1980 Ruhoff: Cardium parile
Deshayes, 1858; [1856-1865] vol. I: 573, pI. 54, figs.
1-3 . --1985 Strougo: Nemocardium (Nemocardium)
parile (Deshayes, 1858). --1995 Le Renard et al.:
Nemocardium (s. str.) parile [Cardium parile]
Deshayes, 1858. --1995 Pacaud & Renard: Nemocardium (Nemocardium) pari/e. --1999 Jacques Vidal
(pers. comm.) MNHN, Paris: Fossils from the Paleontological Collection as Nemocardium parile
(Deshayes). Bartonien. France (Bassin de Paris),
B52884, B69480, B70579.
parisiense Orbigny, 1850. - Changed combination:
Discors parisiense (Orbigny, 1850). OTHER JUDGEMENTS:
Discors discors Lamarck, 1805. » Regarding Cardium
parisiense: Cardiidae. Lower Eocene. TYPE LOCALITY.
Paris Basin, France. RECORDS. --1886 Cossmann:
Divaricardium parisiense (d'Orbigny). --1899 Sacco:
Discors parisiensis (d'Orb.). --1904 Cossmann &
Pissarro: Laevicardium parisiense (d 'Orbigny). --192232 Sherborn, p. 4762: Cardium parisiense d'Orbigny,
1850. --1970 Glibert & Poe1: Discors parisiensis
(Orbigny, 1850). --1970 Glibert & Poel : Cardium
parisiense Orbigny = Cardium discors Lamarck, 1805.
--1977 Popov: Discors parisiensis. --1977 Popov:
Discors parisiense (Orb.). --1995 Le Renard et al.:
Nemocardium (Discors) parisiense [Cardium
parisiense] d 'Orbigny, 1850 (= discors). --1995 Pacaud
& Renard: Nemocardium (Discors) parisiense. --1995
Schneider: Discors parisiense (Orbigny, 1850). --1999
Higo et al.: Cardium parisiense Orbigny, 1850 =
Cardium diseors Lamarck, 1805 non Montagu, 1803.
--1999 Jacques Vidal (pers. com .), MNHN, Paris:
Fossils from the Paleontological Collection as Discors
(or Laevieardium) parisiense d'Orbigny. Bartonien.
France (Bassin de Paris). Egypt.
parisiensis (mode). - See under parisiense Orbigny,
1850.
parki Marwick, 1944. - Changed combination:
Ovicardium parki (Marwick, 1944). Cardiidae. Tertiary.
RECORDS. --1944 ZR: Trachyeardium (Ovicardium )
parki Marwick, 1944 sp. nov. (basal Castlec1iffian)
mouth of Butler's (Ototoka) Creek Niekirrnaru Surv.
Dist. --1973 Beu, p. 318: Ovicardium parki Marwick,
1944 = Ovicardium rossi Marwick, 1944 (Maxwell
1978). --1973 ZR: Aerosterigma (Ovicardium) parki
(Marwick 1944). New synonym of A. (0.) rossi
(Marwick 1944), fossiL). Ref. to Beu. --1977 Popov:
Trachyeardium (Vastieardium) parki Marwick, 1944.
--1978 Maxwell: Ovicardium parki Marwick, 1944.
parkinsoni Sowerby, 1814. - Changed combination:
Laevicardium parkinsoni (Sowerby, 1814). Cardiidae.
Pliocene. RECORDS. --1844 Potiez & Michaud: Cardium
parkinsoni Sow. fossil of England and France, Crag.
--1857 Deshayes : Cardium parkinsoni. --1922-32
Sherborn, p. 4763: Cardium parkinsoni Sowerby, 1814.
"1970 Glibert & Poel : Laevicardium (Dinocardium)
parkinsoni (Sowerby, 1818). --1999 Jacques Vidal
(pers. com.), MNHN , Paris: Fossils from the Paleontological Collection as Cardium parkinsoni Sowerby.
Red Crag. U.K. (Essex and Suffolk), B39920 to -22.
Belgium (Anvers), B43215.
parmata Papaianopol & Lubenescu, 1985. - Prosodacna
parmata. Lymnocardiidae. Pliocene. TYPE LOCALITY.
Ponti en et Ie Dacien du Bassin Dacique, Romania.
RECORDS. --1985 Papaianopol & Lubenescu, p. 140,
fig.: Prosodaella (Prosodacna) parmata sp. nov.
Conocardiidae
partsehii d'Orbigny, 1850. - Changed combination:
Conoeardium partschii d'Orbigny,
1850 .
Conocardiidae. Fossil. RECORDS. --1922-32 Sherborn,
p. 4769: Lunulaeardium partschii Muenster, 1840.
--1922-32 Sherborn, p. 4768: COllocardium (Muenst.)
partschii d 'Orbigny, 1850. - Lunuloeardium.
Lymnocardi idae
partschi Mayer. - Changed combination: Didacna
partschi (Mayer, 1871). Lymnocardiidae. Pliocene.
RECORDS. --1886 Fischer: Cardium partschi Mayer.
--1899 Sacco: Cardium partschi Mayer, 1871. --1899
Sacco: Cardium parlschi Fontannes , 1881 =
Ponlalmyra partschi (May.). --1899 Sacco: Adachna
sic partsehi (May.). --1899 Sacco: Pontalmyra partschi
(May.). --1987 Anaya: Didacna (s. s.)partschi (Mayer,
1871).
partschi castrensis Almera & Bofill, 1894 (variety). Upgraded. Changed combination: Didacna partschi
castrensis (Almera & Bofill, 1894). Lymnocardiidae.
Pliocene. RECOR DS. --1987 Anaya: Didacna (s. s.)
partschi caslrensis (Almera & Bofill, 1894). --1987
Anaya: Cardium partschi var. castrensis.
partschi miopleura Fontannes, 1879-1882 (subspecies).
- Changed combination: Didacna partschi miopleura
(Fontannes, 1879-1882). Lymnocardiidae. Pliocene.
RECORDS. --1987 Anaya: Didacna (s. s.) partsehi
miopleura (Fontannes, 1879-1882).
partsehi subrostrata Almera & Botill, 1894 (variety). Upgraded. Changed combination : Didacna partschi
subrostrata (Almera & Bofill, 1894). Lymnocardiidae.
Pliocene. RECORDS. --1987 Anaya: Didacna (s. s.)
partschi subrostrata (Almera & Bofill, 1894). --1987
Anaya: Cardium partschi var. subrostrata.
partsehi tyopleura Fontannes, 1879 (subspecies) . Changed combination: Didacna partsehi tyopleura
(Fontannes, 1879-1882). Lymnocardiidae. Pliocene.
RECOR DS. --1987 Anaya: Didaena (s. s.) partsehi
tyopleura (Fontannes, 1879-1882).
Parvicardiini Kafanov & Starobogatov, 1977 (tribus).
- Cardiidae. Fossil and Recent. --1977 Kafanov et
Starobogatov: Parvicardiini. --1977 ZR: Parvicardiini
Kafanov & Starobogatoy Tribe nov. ofCardiidae, type
genus Parvicardium Monterosato, 1884. Ref. to
Kafanov & Popov p. 63 [Trans!. p. 313]. --2002
Schneider: citing Kafanov and Popov (1977) subfamily
Fraginae, tribe Parvicardiini.
Parvicardium Monterosato, 1884 (genus). - Cardiidae.
Eocene and Recent. Type species: Crosse (1885)
designated Cardium parvum Philippi, 1844 (non da
Phuket Marine Biological Center Special Publication vol. 30 (2004)
681
Parvicardium - parvulum
Costa, 1778) type species of the genus Parvicardium.
C. parvum Philippi, 1844 has been replaced by C.
exiguum commutatum B.D.D., 1892. RECORDS. ··1900
ZR: Parvicardium Monterosato, 1900 n. sect. of
Cardium for Cardium parvum Phil., etc. ··1969 ZR:
Parvicardium Monterosato, 1884 in subfamily
Cerastodermatinae Nordsieck, F. 1969 ... 1975 ZR:
Parvicardium. Key to species. USSR. Oligocene. Ref.
to Merklin p. 99. ··1977 Popov: Parvicardium
Monterosato, 1884. Genotype: Cardium exiguum
Gmelin, 1790. ··1977 ZR: Parvicardium Monterosato
1884 Designated type genus for new tribe Parvicardiini
Kafanov & Starobogatov. Ref. to Kafanov & Popov p.
63 [Transl. p. 313]. ··1979 ZR: Parvicardium
Monterosato 1884. Ref. to Lambiotte . .. 1980 Vokes:
Parvicardium Monterosato , 1884 in subfamily
Cardiinae. ··1982 Kilburn & Rippey: Parvicardium
Monterosato , 1884. ··1983 Anaya: Parvicardium
Monterosato, 1884 (= Plagiocardium Cossmann, 1886;
= Papillicardium Sacco, 1896). Type: (Crosse, 1885)
Cardium parvum Philippi, 1844, non Da Costa. ··1995
Pacaud & Renard: Parvicardium Monterosato. 1884
in subfamily Cardiinae. ··1998 Schneider:
Parvicardium. Type species Parvicardium siculum
(Sowerby, 1841). Cardiidae. Recent. ··1998 Schneider
(l998b): Parvicardium. The type species of is
uncertain. ··1998 Schneider (1998b): Parvicardium in
Fraginae. The genus originates in the Eocene (Keen
1980). ··2000 Aartsen & Goud: Parvicardium
Monterosato, 1884. ··2001 Vidal, p. 408: Parvicardium.
··2002 Schneider: citing Kafanov and Popov (1977):
Parvicardium in subfamily Fraginae, tribe
Parvicardiini. ··2002 Schneider: Parvicardium is
considered basal fragines. - CHARACTERS AND
OCCURRENCE - DESCRIPTIONS. ··1969 Moore et al.,
emend. Hylleberg 1994. Small shell. Outline roundedsubquadrate; posterior margin less curved than anterior;
ribs sculptured with nodules, blunt spines or raised
ridges (heavy cross-threads) arising from the midline
or covering the complete rib. Hinge line gently curved;
hinge weak, cardinal teeth minute, lower cardinal much
larger and than upper cardinal; fused at base in LV.
Inner margin crenulate. Resembling Plagiocardium
(Papillicardium). ··2001 Vidal: only the exhalant
aperture is tube-shaped. Inhalant siphon open towards
the foot aperture. No eyes. DISTRIBUTION. Europe-North
America.
Parvicardium Monterosato, 1884 (subgenus and
section). - Cardiidae. Recent and Eocene. REMARKS.
The designation "section" takes the value of subgenus.
RECORDS. ··1887 Fischer: Parvicardium Monterosato,
1884. Section of genus Cardium Linne, 1758. "1912
Dautzenberg & Fischer: Subgenus Parvicardium
Monterosato 1884.
Parvidacna Stevanovic, 1951 (genus). Lymnocardiidae.
Pontian. RECORDs. ··1951 ZR: Parvidacna Stevanovic,
1950 gen. nov. Cardiidae genotype Parvidacna
planicostata Stevanovic, 1950 sp. nov. Upper Pontian
Serbia. ··1953 ZR: Parvidacna Stevanovic, 1951 gen.
nov. Parvidacna planicostata Stevanovic, 1951 sp. nov.
Ponti an Serbia. ··1980 Vokes: Parvidacna (genus)
Stevanovic, 1950 [Ebersin & Stevanovic, 1949 n.n.]
in subfamily Paradacninae. ··1987 ZR: Parvidacna
Stevanovic, 1951 . Referred to new tribe. Phylocardiini.
Ref. to Nevesskaya et al. p. 189.
parvissimum Svagrovsky, 1960. - Parvicardium
parvissimum (Svagrovsky, 1960). Cardiidae. Miocene.
RE COR DS . ··1960 ZR: Cardium (Parvicardium)
parvissimum Svagrovsky, 1960 sp. nov. Upper
Tortonian (Miocene) Kirzan massive "Slanske hory"
mts.
parvula Nalivkin, 1916. - Didacna parvula Nalivkin,
1916. Lymnocardiidae. Fossil. RECORDS. "1916 ZR:
Didacna parvula Nalivkin, 1916 sp. n. Berg der
Bakustufe.
parvula anomalodonta Nalivkin, 1916 (variety). Upgraded. Didacna parvula anomalodonta Nalivkin,
1916. Lymnocardiidae. Fossil. RECORDS. "1916 ZR:
Didacna parvula var. anomalodonta Nalivkin, 1916
var. n. Berg der Bakustufe.
parvula catillus Nalivkin, 1916 (variety) Upgraded. Didacna parvula catillus NaJivkin , 1916 .
Lymnocardiidae. Fossil. RECORDS. ··1916 ZR: Didacna
parvula var. catillus Nalivkin, 1916 var. n. Berg der
Bakustufe.
parvula planissima Nalivkin, 1916 (variety) . Upgraded. Didacna parvula planissima Nalivkin, 1916.
Lymnocardiidae. Fossil. REcORDs. ··1916 ZR: Didacna
parvttla var. planissima Nalivkin, 1916 var. n. Berg
der Bakustufe.
parvlllinllm Locard, 1900. - Synonym of Parvicardium
parvum (Philippi, 1844). » Regarding Cardium
parvulinum: Cardiidae. Recent. DISTRIBUTION.
Mediterranean, 50-60 m depth. RECORDS. ··1900
Locard & Caziot: Cardium parvulinum Locard, 1900.
··1972 Petersen & Russell: Cardium parvulinum
Locard & Caziot, 1900 = Parvicardium parvum
(Philippi, 1844). ··1977 Fischer-Piette: Cardium
exiguum Gmelin. ··2001 Vidal, p. 408: Parvicardium
parvulinum. - CHARACTERS AND OCCURRENCE
- DESCRIPTIONS. ··200 I Vidal: only the exhalant aperture
is tube-shaped. Inhalant siphon open towards the foot
aperture. No eyes.
parvuloides Popov, 1955. - Didacna parvuloides Popov,
1955. Lymnocardiidae. Quaternary. RECORDS. ··1956
ZR: Didacna parvuloides Popov, 1955 sp. nov.
Premanich part of Pricaspian Quaternary. Russia.
*parvulum. - Homonymy to be noticed. Cardium
parvulum from 4 sources: Dunker, 1861. - Martin,
1879. - Li , 1930. - Tavani, 1948.
parvllium Dunker, 1861. - Not a cardiid. » Regarding
Cardium parvulum: of genus Cardita or Carditella.
RECORDS. ··1862 Dunker, p. 36: Cardium parvulum.
··1890 Paetel: Cardium parvulum Dkr. = edule L.
"1951 Carcelles & Williamson : Cardium parvulum
Dunker 1862. ··1959 Soot-Ryen, p. 54: Cardium
parvu/um Dkr. = Cardita or Carditella. ··1977 FischerPiette: Cardium parvulum Dunker = Cardita or
Carditella. ··1980 Ruhoff: Cardium parvulum Dunker,
1861; vol. 8: 36.
parvulum Li, 1930. - Synonym of Trachycardium
procerum (Sowerby, 1833). » Regarding Cardium
parvulum: Cardiidae. Recent. RECORDS. ··1930 Li:
Cardium parvulum. ··1931 Pilsbry: Cardium parvulum
Li (unmistakable C. procerum Sowerby.). ··1931 ZR:
Cardum sic [Cardium] parvulum Li, 1931 sp. n.
Panama. ··1947 Hertlein & Strong: Cardium parvulum
Li = Cardium (Mexicardia) procerum Sowerby. ··1968
682
Tropical Marine Mollusc Programme (TMMP)
parvulum - parvum
Velez: Cardium parvulum = Trachycardium procerum
(Sowerby, 1833). ··1971 Keen & McLean: Cardium
parvulum Li, 1930 = Trachycardium (Mexicardia)
procerum (Sowerby, 1833). Distribution as for T
procerum: occasional in the Gulf of California, South
West Mexico to Chile. ··1974 Abbott: Cardium
parvulum Li, 1930 = Trachycardium (Mexicardia)
procerum (Sowerby, 1833). ··1977 Fischer-Piette:
Cardium parvulum Li = Laevicardium procerum
Sowerby.
parvulum Martin, 1879. - Cardium parvulum Martin,
1879. Cardiidae. Upper Miocene. RECORDS. ··1919
Martin: Cardium parvulum Mart. (U. Miocene).
parvulum Tavani, 1948. - "Cardium" parvulum Tavani,
1948. Cardiidae. Cretaceous. RECORDS. ··1948 ZR:
Cardium parvulum Tavani, 1948 sp. nov. Cretaceous,
Somali land.
parvulum Dunker (variety). - RECORDs. ··1890 Paetel:
Cardium var. parvulum Dkr.
Lymnocardiidae
parvulllm Fuchs. - Changed combination:
Lymnocardium parvulum (Fuchs). Lymnocardiidae.
Fossil. RECORDs. ··1987 Taktakishvili: Limnocardium
parvulum Fuchs.
parvllillm Morris & Lycett, 1853. - Unicardium
parvulum Morris & Lycett, 1853. Lymnocardiidae.
Fossil. REcORDs. ··1980 Ruhoff: Unicardium parvulum
Morris & Lycett, 1853; [1853-1854] vol. 2: 74; pI. 8,
fig. 6a.
parvllills (variety). - See under dahestanicum parvulus
Uspenskaia, 1931.
parvllm (subspecies and variety). - See under haueri
parvum Stevanovic, 1979. - glaucum rectidens var.
parvum Brusina, 1870.
*parvllm. - Homonymy to be noticed. Cardium parvum
from 6 sources: da Costa, 1778. - Mawe, 1823. Sowerby, 1838. - Philippi, 1844. - Adams in
MacAndrew, 1870. - Deshayes, (?year).
parvllm Adams in M' Andrew, 1870. - Unknown species.
» Regarding Cardium parvum: TYPE LOCALITY. Gulf
of Suez. REMARKS. No description, no figures (FischerPiette 1977). RECORDS. ··1870 Adams in MacAndrew,
p. 447 : Cardium parvum n. sp. ··1886 Cooke: Cardium
parvum Ad. ··1977 Fischer-Piette: Cardium parvum
(H. Adams) McAndrew = species insufficiently known.
··1980 Ruhoff: Cardium parvum H. Adams in
M'Andrew, 1870; vol. 6: 447.
parvllm Brusina, 1870 (variety) . - Synonym of
Parvicardium exiguum Gmelin. » Regarding Cardium
var. parvum: Cardiidae. Recent. RECORDS. ··1880
Stossich: Cardium var. parvum = Cardium exiguum
Gmelin.
parvllm da Costa, 1778 . - Unknown species. OTHER
JUDGEMENTS: Acanthocardia acuLeala (Linnaeus, 1758).
- Cardium ciliare Linnaeus. - Acanthocardia echinata
(Linnaeus, 1758). - Parvicardium exiguum var? Cerastoderma glaucum (Bruguiere, 1789). »
Regarding Cardium parvum: Cardiidae. Recent.
RECORDS. ··1778 da Costa: Cardium parvum. ··1789
Bruguiere: Cardium parvum d' Acosta pag. 177, num.
17. ··1803 Montagu: Cardium parvum Da Costa. p.
177. = Cardium ciliare Lin. ··1817 Dillwyn: Cardium
parvum Da Costa = Cardium echinalum Linnaeus.
··1828 Mawe: Cardium parvum da Costa, 1778 =
Cerastoderma glaucum (Bruguiere, 1789). ··1852
Leach: Cardium parvum Da Costa = Testa junior
Cardium eehinatum. ··1856 Wood & Hanley: C.
parvum Da Costa, 177 - Is the young of C. aeu/eatum.
··1890 Paetel : Cardium parvum Cost. = aculeatum L.
juv. ··1898 Bucquoy, Dautzenberg & Dollfus: Cardium
parvum da Costa, 1778 = juv. Acanthoeardia arnleatum
or A. eehinatum. ··1899 Sacco: Cardium parvum Da
Costa 1778 = Parvieardium exiguum var? ··1902
Sherborn, p. 720: Cardium parvum da Costa, 1778.
··1972 Petersen & Russell: Cardium parvum da Costa,
1778 = unknown cockle according to Sowerby (1840).
··1977 Fischer-Piette: Cardium parvum da Costa.
parvum Deshayes. - Combination to be revised :
"Cardium" parvum Deshayes. Cardiidae. Fossil.
RECORDs. ··1857 Deshayes: C. parvum Desh.
parvum Loven. - Misidentification of Parvicardium
exiguum (Gmelin, 1791). » Regarding Cardium
parvum: Cardiidae. Recent. REcoRDs. ··1871 March,
p. 215, No. 205 Cardium parvum Loven = Cardium
exiguum Gm.
parvum Mawe, 1823. - Cardium parvum Mawe, 1823 .
- OTHER JUDGEMENTS: Parvieardium exiguum (Gmelin,
1791). » Regarding Cardium parvum: Cardiidae.
Recent. REcORDs. ··1823 Mawe, p. 29, 32, Figs. 4- 5:
Cardium parvum. ··1922-32 Sherborn, p. 4777:
Cardium parvum Mawe, 1823 [n. et f] = nomen nudum
accompanied by figure. ··1977 Fischer-Piette: Cardium
parvum Mawe = Cardium exiguum Gmelin.
parvlIm Paramonova, 1977. - Cerastoderma parvum
Paramonova, 1977. Cardiidae . Upper Miocene.
RECORDS. ··1977 ZR: Cerastoderma (Planacardium)
parvum Paramo nova sp. nov. Russian SFSR. Ref to
II'ina, Nevesskaya & Paramonova p. 209, fig'd.
parvlIm Philippi, 1844. - Synonym of Parvicardium
exiguum (Gmelin, 1791). OTHER JUDGEMENTS: Cardium
aculeatum Linnaeus, 1758. - Parvieardium eommutalum (Bucquoy, Dautzenberg & Dollfus, 1892). Parvieardium exiguum (Gmelin, 1791). - Removal
from synonymy with C. exiguum Gmelin, 1791. Cardium fascialum Montagu. - Cardium parvulinum
Locard, 1900. - Parvieardium parvum (Philippi, 1844).
» Regarding Cardium parvum: Cardiidae. Recent.
Type species Cardium parvum Philippi, 1844, non da
Costa (Crosse, 1886) is type species of genus
Parvieardium Monterosato, 1884 (Glibert & Poe I
1970). Cardium parvum Philippi 1844 is type species
of subgenus Parvieardium Montrs. 1884 (Sacco 1899).
RECORDS. ··1844 Philippi, p. 39, pI. 14, figs. 17 a,b:
Cardium parvum n. sp. ··1849 Middendorff: Cardium
parvum . ··1856 Jeffreys: (Cardium parvum, Ph.) =
Cardiumfasciatum, B. M. (Note: Ph. & B. M. = Philippi
and "British Mollusca", Forbes & Hanley 1849
respectively). ··1859 Sowerby: C. parvum = Cardium
fasciatum Mont. ··1859 Sowerby: C. parvum (young)
= Cardium aeulealum, Linn. ··1869 Romer: Cardium
parvum Philippi (a variety of Cardium exiguum
Gmelin). ··1884 Monterosato: Parvicardium parvum ,
Ph. (Cardium) . ··1885 Crosse designated Cardium
parvum Philippi, 1844 as type species of the genus
Parvicardium Monterosato, 1884 . ··1886 Locard:
Cardium? parvum Philippi, 1844 = Cardium fasciatum
Montagu, 1808. ··1887 Fischer: Cardium parvum
Philippi. ··1888 Kobelt: Cardium parvum Philippi =
Phuket Marine Biological Center Special Publication vol. 30 (2004)
683
parvum - patrulii
Cardium exiguum Gmelin. "1890 Paete1: Cardium
parvum PhI. = exiguum Gm. "1891 Danilo & Sandri:
Cardium parvum Phil. ··1892 Locard: Cardium
parvum, Philippi, 1844. ··1900 Locard & Caziot:
Cardium parvum Philippi, 1844 = Cardium parvulinum
Locard, 1900. ··1900 ZR: Cardium parvum Phil. [Ref.
Monterosato 1900). ··1917 Hidalgo: Cardium parvum
Philippi = Cardium exiguum Gmelin. ··1922-32
Sherborn, p. 4777: Cardium parvum Philippi, 1844.
··1933 Coen: Cardium parvum Philippi = Eucardium
(Parvicardium) exiguum var. scripta B.D.D. "1933
Coen & Vatova: Cardium parvum Phil. (Stossich 1865)
= Eucardium (Parvicardium) exiguum Gmelin. ··1937
Coen: parvum Philippi = Cerastoderma (Parvicardium)
exiguum scriptum B.D.D. ··1962 Grossu: Cardium
parvum Philippi, 1844 = Cardium (Parvicardium)
exiguum Gmelin, 1790 sic. ··1969 Moore et al.:
Cardium parvum Philippi, 1844 = Cardium exiguum
commutatum Bucquoy, Dautzenberg & Dollfus, 1892.
··1969 Nordsieck: Parvicardium parvum (Philippi,
1844) = Parvicardium exiguum (exiguum) (Gmelin,
1791). "1972 ZR: Cardium parvum Philippi, 1844.
Removal from synonymy With C. exiguum Gmelin,
1791 [Ref. Petersen 1972). ··1977 Fischer-Piette, p.
174: Cardium parvum Philippi = Cardium exiguum
Gmelin. ··1978 ZR: Cardium parvum Philippi, 1844.
Valid name. Ref. to Janssen et at. p. 56. ··1983 Anaya:
Cardium parvum Philippi, 1844, non Da Costa. ··1991
van Aartsen & Giannuzzi-Savelli: Cardium parvum
Phi lippi, 1844; preoccupied name replaced by
Parvicardium commutatum (B.D.D., 1892). ··2000
Aartsen & Goud: Cardium parvum Philippi, 1844 not
Da Costa 1778, not Mawe 1823, not Sowerby 1840 is
a synonym of Parvicardium exiguum (Gmelin, 1791).
- CHARACTERS AND OCCURRENCE - OF COLLECTIONS. ··ZMUC: Lake Fusaro, near Naples, Italy.
DISTRIBUTION. ··1969 N ordsieck: Mediterranean.
REMARKS. Petersen & Russell (1972) Cardium
parvulinum Locard & Caziot, 1900 sic Locard 1900 =
Parvicardium parvum (Philippi, 1844); Nordsieck
(1969) stated the following synonymy: Parvicardium
exiguum (exiguum) (Gmelin, 1791) = pygmaeum
(Donovan, 1799) = siculum (Sowerby, 1839 sic. =
1834) = stellatum (Reeve, 1844).
parvum Philippi (variety).- Infrasubspecific. RECORDS.
··1890 Paetel: Cardium var. parvum PhI.
parvum Sowerby, 1838. [Figs. Part I p. 112). Synonym
of Cardium exiguum Gmelin. » Regarding Cardium
parvum: Cardiidae. Recent. TYPE LOCALITY. Ungown
(Sowerby 1840). RECORDS. ··1838 Sowerby, 149t part,
fig. 33: Cardiumparvum nobis. "1840 Sowerbyp. 110:
Cardium parvum. ··1840 Sowerby, catalogue p. 6, No.
80: Cardium parvum Sow. ··1840 Sowerby, corrected
list, p. 8: Cardium parvum (Sowerby No. 80). ··192232 Sherborn, p. 4777: Cardium parvum Sowerby, 1841.
··1977 Fischer-Piette: Cardium parvum Sowerby =
Cardium exiguum Gmelin. - CHARACTERS AND
OCCURRENCE - DESCRIPTIONS. ··1840 Sowerby: shell
oval, subquadrate, posterior subangulate, anterior
rounded, pale yellowish, dark reddish spotted along
margin, numerous ribs, somewhat flattened, narrow
inters paces [I have translated from Latin). DIMENSIONS.
··1840 Sowerby: long. 0.50 poll. (length 13 mm), lat.
0.40 poll. (width 10 mm), alt. 0.43 poll (height 11 mm).
parvum commutata Bucquoy, Dautzenberg & Dollfus,
1892 (variety). - Infrasubspecific. No evidence of
subspecific status. » Regarding Parvicardium parvum
var. commutata: Cardiidae. Recent. RECORDS. ··1899
Sacco: Cardium parvum var. commutata B.D.D.
parvum flavida (variety). - Cardium parvum var.flavida.
No evidence of subspecific status. » Regarding
Cardium parvum var. jlavida: Cardiidae. Recent.
RECORDS. ··1878 Monterosato: Cardium parvum, Ph.
var.jlavida.
Pascoella (genus) Cox, 1949. - Cardiidae. Fossil.
RECORDS. ··1980. Vokes: Pascoella Cox, 1949 in
subfamily Cardiinae.
pasinii Schauroth, 1865. - "Combination to be revised:
"Cardium" pasinii Schauroth, 1865. Cardiidae. Fossil.
RECORDS. ··1980 Ruhoff: Cardium pasinii Schauroth,
1865; p. 210, pI. 20, figs. 1-3.
passyi Deshayes, 1858. - Synonym of Parvicardium
granulosum (Lamarck, 1805). » Regarding Cardium
passyi: Cardiidae. Eocene. TYPE LOCALITY. Paris Basin,
France. RECORDS. ··1886 Cossmann: Cardium passyi
Deshayes. ··1904 Cossmann & Pissarro: Cardium
(Plagiocardium) passyi Deshayes. "1970 Glibert &
Poel: Cardium passyi Deshayes, 1856 = Parvicardium
granulosum (Lamarck, 1805). ··1977 Popov:
Plagiocardium (Plagiocardium) passyi (Deshayes).
··1980 Ruhoff: Cardium passyi Deshayes, 1858;
[1856-1865) vol. 1: 557, pI. 71, figs. 12-14 (sic = 1416). ··1995 Le Renard et al.: Plagiocardium
granulosum passyi [Cardium passyi) Deshayes, 1858.
··1995 Pacaud & Renard: Plagiocardium passyi. ··1999
Jacques Vidal (pers. comm.) MNHN, Paris: Fossils
from the Paleontological Collection as Plagiocardium
passyi (Deshayes). Lutetien. France (Bassin de Paris),
B52724, B52725.
patagonicum Ihering, 1907. - Hedecardium
patagonicum (Ihering, 1907). Cardiidae. Oligocene.
RECORDS. ··1907 Ihering: Cardium patagonicum (fide
Schneider 2002). ··2002 Schneider: Hedecardium
(lheringicardium) patagonicum Cardiidae. Oligocene.
Patrocardium (genus). - RECORDS. ··1926 ZR:
Patrocardium [Ref. Heller 1926).
patruelinum Deshayes, 1858. - Changed combination:
Loxocardium patruelinum (Deshayes, 1858). Cardiidae.
Eocene. TYPE LOCALITY. Paris Basin, France. RECORDS.
··1886 Cossmann: Cardium patruelinum Deshayes.
··1904 Cossmann & Pissarro: Cardium (Loxocardium)
patruelinum Deshayes. ··1970 Glibert & Poel:
Loxocardium patruelinum (Deshayes, 1858). ··1977
Popov: Loxocardium patntelinum (Desha yes, 1858).
··1980 Ruhoff: Cardium patruelinum Deshayes, 1858;
[1856-1865) vol. I: 565, pI. 55, figs. 8-10. "1995 Le
Renard et al.: Loxocardium patruelinum [Cardium
patruelinum) Deshayes, 1858. ··1995 Pacaud &
Renard: Loxocardium patruelinum. ··1999 Jacques
Vidal (pers. comm.) MNHN, Paris: Fossils from the
Paleontological Collection. Loxocardium patruelinum
(Deshayes). Ypresien. France (Bassin de Paris), B52818
to -25, B72593.
patrulii (subspecies). - See under penslii patrulii.
patrulii Marinescu, 1975. - New combination: Lymnocardium patrulii (Marinescu, 1975). Lymnocardiidae.
Pliocene. RECORDS. ··1975 ZR: Limnocardium
(Pannonicardium) patrulii Marinescu, 1975 sp. nov.
684
Tropical Marine Mollusc Programme (TMMP)
patruliusi - paucicostatum
Romania, Banat. Ref. to Marinescu, p. 19.
patruliusi Papaianopol, 1992. - Pseudocatillus patruliusi
Papaianopol, 1992. Lymnocardiidae. Pliocene. TYPE
LOCALITY. Dacien inferieur(Getien) du Bassin Dacique,
Romania. RECORDS. ··1992 Papaianopol, p. 26, fig.:
Pseudocatillus patn/liusi sp. nov.
patulum Hutton, 1873. - Nemocardium patulum (Hutton,
1873). Cardiidae. Fossil. TYPE SPECIES: Cardium
patulum Hutton, 1873. Type of (Varicardium) Marwick,
1944 (Popov 1977). RECORDS. ··1944 ZR:
Protocardium patulum Hutton 1873 [Ref. Marwick
1944]. ··1944 ZR: Varicardium patulum (Hutton, 1873)
[Ref. Marwick 1944]. ··1965 ZR: Varicardium patulum
revised, Tertiary, New Zealand [Ref. Boreham 1965].
.. 1977 Popov: Nemocardium (Varicardium) patulum
(Hutton, 1873). ··1995 Schneider: Nemocardium
(Varicardium) patulum (Hutton, 1873).
paucicostatal-um (mode, variety). - See under
paucicostatum. - exiguum paucicostatum Milascewicz,
1909.
*paucicostatum. - Homonymy to be noticed. Cardium
paucicostatum from 4 sources: Murchison, 1831. Sowerby, 1834. - Deshayes, 1838. - Munster, 1840.
paucicostatum Deshayes, 1838. - Pontalmyra
paucicostatum (Desha yes, 1837). Lymnocardiidae.
Fossil. TYPE LOCALITY. The Crimea. RECORDS. ··1838
Deshayes, p. 52, pI. 2, figs. 14-15: Cardium
paucicostatum. ··1857 Deshayes: Cardium
paucicostatum Deshayes, 1837 sic. "1886 Fischer:
Cardium paucicostatum Deshayes. ··1922-32
Sherborn, p. 4792: Cardium paucicostatum Deshayes,
1838. ··1970 Glibert & Poel: Cardium paucicostatum
non Sowerby. ··1980 ZR: Cardium paucicostatum.
Black Sea. Ref. to Gomoiu et at. ··1980 ZR: Didacna
(Pontalmyra) kamyshburunensis Kafanov, 1980 nom.
nov. for Cardium paucicostatum Deshayes 1838 nee
Sowerby 1834. Ref. to Kafanov, p. 623. ··1987
Taktakishvili: Pontalmyra paucicostata (Desh.). ··1987
Taktakishvili: Didacna paucicostata (Desh.). CHARACTERS AND OCCURRENCE - DESCRIPTIONS. ··1903 Andrussoff: Cardium pauci-costatum has
well developed cardinal teeth.
paucicostatum Murchison, 1831. - Combination to be
revised: "Cardium" paucicostatum Murchison, 1831.
Cardiidae. Recent. RECORDS. ··1857 Deshayes:
Cardium paucicostatum Murchison, 1831.
paucicostatum Munster, 1840. - Replaced. Combination
to be revised: "Cardium" raricostatum Deshayes. »
Regarding Cardium paucicostatum: ?Cardiacea. Fossil.
RECORDS. ··1857 Deshayes: Cardium paucicostatum
Munster = C. raricostatum Desh. ··1922-32 Sherborn,
p. 4792: Cardium paucicostatum Muenster, 1840.
··1922-32 Sherborn, p. 4792: Conocardium (Muenst.)
paucicostatum d'Orbigny, 1850. - Cardium.
paucicostatum Orbigny, 1850. - See under paucicostatum Munster, 1840.
paucicostatum Sowerby, 1834. [Figs. Part I p. 90, 109,
127]. - Changed combination: Acanthocardia
paucicostata (Sowerby, 1834). OTHER JUDGEMENTS:
Cardium ciliare Lamarck. » Regarding Cardium
paucicostatum: Cardiidae. Miocene and Recent. Keen
1937 designated Cardium paucicostatum Sowerby,
1841 type species of the genus Sphaerocardium Coen,
1933. TYPE LOCALITY. Adriatic Sea (Malta), Sowerby
(1840). TYPE セpositryN@
Unknown. RECORDS. "1834
Sowerby, 49 t part, fig. 20: Cardium paucicostatum.
··1840 Sowerby, p.l06: Cardium paucicostatum.
··1840 Sowerby, p. 2, No. 29: Cardium paucicostatum.
··1840 Sowerby, corrected list, p. 8: Cardium
paucicostatum (Sowerby No. 29). "1844 Philippi:
Cardium paucicostatum Sow. = Cardium ciliare. ··1844
Reeve: Cardium paucicostatum Sowerby, 1840. ··1857
Deshayes: Cardium paucicostatum Sowerby, 1833.
"1857 Deshayes: Cardium paucicostatum Sowerby,
1839 sic - C. ciliare. Lamk. ··1857 Deshayes: Cardium
paucicostatum Sowerby, 1839 - C. ciliare Lamk.
··1858 Adams & Adams, p. 455: Cardium (Isocardia)
paucicostatum Sow. ··1867 Weinkauff: Cardium
paucicostatum Sowerby Guv.) = Cardium echinatum
Linne. "1867 ZR: Cardium paucicostatum (Sow.) =
Cardium deshayesii (Payr.), which alone is found in
the Mediterranean, and not on the Atlantic shores of
Europe [Ref. Weinkauff 1867]. ··1868 ZR:
paucicostatum of Sowerby and Reeve = Cardium
ciliare (Linne, not Reeve) [Ref. Romer 1868]. ··1878
Monterosato: Cardium paucicostatum Sow. = C. ciliare,
auctt. (non Lin.). ··1884 Grasset: Cardium
paucicostatum Sow. "1890 Paetel: Cardium
paucicostatum Sow. = echinatum juv. ··1892 Locard:
Cardium paucicostatum Sowerby. 1859 sic ... 1899
Sacco: Cardium paucicostatum Sowerby 1839
[Miocene-Recent]. ··1900 Locard & Caziot: Cardium
paucicostatum Sowerby, 1859 sic. ··1907 ZR: Cardium
(s. strict.) paucicostatum [Ref. Martinelli 1907]. ··1907
ZR: Cardium paucicostatum [Refs. Roule 1907.
Zimmermann 1907]. ··1916 Milaschewitch: Cardium
paucicostatum Sowerby, 1893 sic = Cardium
paucicostatum Sowerby var. impedita Mil. (1909).
··1922-32 Sherborn, p. 4792: Cardium paucicostatum
Sowerby, 1841. ··1931 ZR: Cardium paucicostatum
Ref. to Borcea (1931). ··1933 Coen: Eucardium
(Sphaerocardium) paucicostatum Sowerby. ··1933
Coen & Vatova: Eucardium (Sphaerocardium)
paucicostatum Sowerby. "1937 Coen: Cerastoderma
(Sphaerocardium) paucicostatum Sowerby. ··1937 ZR:
Cardium paucicostatum Sowerby, 1841 type of
Sphaerocardium, probably = Acanthocardia Gray, 1851
[Ref. Keen 1937] ... 1939 Steuer: Cardium (Rudicardium) paucicostatum Sowerby. ··1962 PasteurHumbert: Cardium paucicostatum Sowerby. ··1969
Nordsieck: Sphaerocardium paucicostatum (Sowerby,
1839). ··1977 Fischer-Piette: Cardium (Cerastoderma,
Acanthocardia) paucicostatum Sowerby. ··1977
Popov: Acanthocardia (Acanthocardia) paucicostata
(Sowerby, 1839). "1978 Janssen & Van der Slik:
Sphaerocardium paucicostatum (Sowerby, 1839).
··1980 ZR: Cardium paucicostatum Sowerby 1834.
Replaced by C. ciliare Linnaeus 1758 Ref. to Kafanov,
p. 625 ... 1983 Salvini-Plawen: Sphaerocardium
paucicostatum (Sowerby). ··1991 Krzyminska &
Wolowicz: Cardium paucicostatum Sowerby. ··1998
Macedo et al.: Acanthocardia paucicostata (Sowerby
G.B. III, 1841). "1999 Vidal (pers. comm.). Fossils
from the Paleontological Collection as Acanthocardia
paucicostatum. Italy; Morocco; Bulgaria. ··200 I Vidal,
p. 407: Acanthocardia paucicostatum.
CHARACTERS AND OCCURRENCE - OF
COLLECTIONS. ··ZMUC: 45 m (Ashdod Israel). ··ZMUC:
Phukel Marine Biological Center Special Publication vol. 30 (2004)
685
paucieostalum alba - paucieostatum rarepartita
3-7 m (La Gorellette, Gulf of Tunesia). ··ZMUC: 16
ribs (range 14-19), n = 15. ··ZMUC wet collection:
Black Sea. DESCRIPTIONS. ··1844 Reeve: shell rounded,
thin, rather ventricose, nearly equilateral, radiately
ribbed, 16-17 flatly convex ribs, keeled in the middle,
keel armed with scale-like spines, interstices concave,
wide, crisply wrinkled zoned transversely in a waved
manner with white and reddish brown. ··200 I Vidal:
with a few additional larger tentacles outside siphons.
No eyes on tentacles. DIMENSIONS. ··1840 Sowerby:
long. 1.3 poll. (length 33 mm), lat. 1.3 poll. (width 33
mm), alt. I. poll (height 25 mm). REMARKS. ··1840
Sowerby: this species differs from C. eehinatum in
being comparatively smooth, and in having few ribs,
with wide interstices. The genus Aeanthoeardia is used
instead of the genus Sphaeroeardium. ··1844: Reeve
found that Mr. Sowerby has acted very judiciously in
promoting this shell to the rank of a new species. The
Cardium eiliare, with which it has been confounded,
has certainly as small a numberofribs, and is moreover
the only other aculeated cockle with so few, but the
spines of these two species vary most importantly in
their character. The Cardium eiliare has a solid laterally
compressed spine like that of the young Cardium
aeuleatum, noticed above; the Cardium paueieosta/um
has a folded scale-like spine, papilliform at the base,
like that of the Cardium erinaceum. HABITAT. ··1900
Locard & Caziot: 40-60 m. ··ZMUC: 3-45 m depth.
DISTRIBUTION. ··1844 Reeve: Adriatic, South Europe.
··1884 Grasset: Algeria. ··Of authors: West Europe,
Mediterranean, Black Sea, West Africa; Ashdod Israel,
La Gorellette, Gulf ofTunesia, Black Sea.
paucieostatum alba Bucquoy, Dautzenberg & Dollfus,
1892 (variety). - New combination: Acanthocardia
paucicostala var. alba (B.D.D., 1892). » Regarding
Cardium paucicoslalum var. alba: Infrasubspecific. No
evidence of subspecific status. Cardiidae. Recent.
RECORDS. ··1892 Bucquoy, Dautzenberg & Dol1fus, p.
270: Cardium paucicostatum var. (ex colore) alba
B.D.D. ··1969 Nordsieck: Sphaerocardium paucicoslalum (Sowerby, 1839) var. album (Bucquoi sic,
Dautzenberg & Dollfus). - CHARACTERS AND
OCCURRENCE - DISTRIBUTION. ··1969 Nordsieck:
unknown.
paucieostatum bianeoniana Cocconi 1873 (variety). Upgraded. New combination : Acanlhocardia
paucicostata bianconiana (Cocconi 1873). Cardiidae.
Fossil. RECORDS. ··1899 Sacco: Cardium paucicostatum
var. bianconiana Cocconi 1873 (an species
distinguenda). "1912 Favre: Cardium paucicostatum
Sowerby var. bianconiana Cocco ID suggested by
Favre.
paucieostatum impeditum Milaschewitsch, 1909
(variety). - Replaced by Acanlhocardia ciliaris
milaschewitehi. » Regarding Cardium paucicostata
var. impedila : Cardiidae. Recent. RECORDS. ··1916
Milaschewitch : Cardium paucieostatum Sowerby var.
impedita Mil. (J 909). ··1969 Nordsieck : Sphaerocardium paucicostatum (Sowerby, 1839) var.
impeditum (Milaskewicz) . ··1980 ZR : Cardium
paucicostatum var. impedita Milaschewitsch 1909.
Preoccupied name , replaced by Acanthocardia
(Sphaerocardium) ciliaris milaschewitchi. Kafanov,
1980. Ref. to Kafanov, p. 625. - CHARACTERS AND
OCCURRENCE - DISTRIBUTION . ··1969 Nordsieck:
Black Sea.
paucieostatum milasehewitsehi Kafanov, 1980 . Replacement name for Cardium paucieostatum var.
impedita Mi1aschewitsch 1909.
paueieostatum piaeenziana (variety). - New
combination: Aeanthocardia paucicostatum var.
piacenziana . » Regarding Cardium paueicostatum
var. piacenziana: Cardiidae. Fossil. RECORDS. ··1899
Sacco: Cardium paucicostatum var. piacenziana.
paueieostatum planieosta Coen, 1933 (variety). Infrasubspecific. No evidence of subspecific status. »
Regarding Eucardium paucicostatum var. planicosta:
new combination : Acanthocardia paucicostata
(Sowerby, 1834). Cardiidae. Recent. RECORDS. ··1933
Coen: Eueardium (Sphaerocardium) paucicostatum
var. planicosta Monterosato mss. ··1933 ZR:
Sphaerocardium pallcicostatum var. planieosta Coen,
1933 nov. Adriatic [Ref. Coen 1933]. ··1937 Coen, pI.
10, fig. 90, Note 263 : Cerastoderma (Sphaerocardium)
paucicostalum Sowerby planicosta Monterosato, Coen
n. var. ··1937 ZR : Cardium (Sphaerocardium)
paucicostatum var. planicosta (Monts. mss.) Coen,
1937 nov. Adriatic . ··1937 ZR : Sphaerocardium
paucicostatum var. planicosta Coen, 1933 nov. Venice.
paucieostatum p/anieosta eburnea Coen, 1914 (variety).
- Infrasubspecific . No evidence of subspecific status.
» Regarding Eucardium paucicostatum planicosta
var. eburnea: new combination: Acanthocardia
paucicostata (Sowerby, 1834). DISTRIBUTION. The
Mediterranean. Cardiidae. Recent. RECORDS. ··1914
ZR: Eucardium (Tropicardium) paucicostalum eburnea
var. n. Coen, 1914. ··1933 Coen: Eucardium (Sphaerocardium) paucicostatum var. eburnea Coen. ··1937
Coen, Note 263: Ceras/oderma (Sphaerocardium)
paucicos/atum Sowerby var. eburnea Coen n. var.
··1937 ZR : Eucardium (Sphoerocardium sic)
paucicostatum Sow planicos/a (Monts. mss.) var.
eburnea Coen, 1933 nov. Adriatic.
paucieostatum produetum Bucquoy, Dautzenberg &
Dollfus, 1892 (variety). - Infrasubspecific. No evidence
of subspecific status. » Regarding Cardium
paucicosta/a produc/um: new combination:
Acan/hocardia paucicosta/a (Sowerby, 1834).
Cardiidae. Recent. RECORDS. ··1892 Bucquoy,
Dautzenberg & Dollfus: Cardium paucicostatum var.
productum. ··1899 Sacco: Cardium paucicos/a/um var.
producta B.D.D. 1892. ··1933 Coen: Eucardium
(Sphaerocardium) paucicostalum var. producta B.D.D.
··1937 Coen: Cerastoderma (Sphaerocardium)
paucicoslalum productum. ··1969 Nordsieck :
Sphaerocardium paucicosta/um (Sowerby, 1839 sic)
var. productum (Bucquoi sic, Dautzenberg & Dollfus).
Unknown distribution.
paucieostatum rarepartita Coen, 1933 (variety). Infrasubspecific. No evidence of sub specific status.»
Regarding Eucardium paucicos/alum var. anomala
rarepartita: new combination: Aeanthocardia
paucicostata (Sowerby, 1834). Cardiidae. Recent.
RECORDS. ··1933 Coen: Eucardium (Sphaerocardium)
paucicostatum var. anomala rareparti/a Coen. ··1933
ZR: Sphaeroeardium paueicostalum var. rarepartita
Coen, 1933 nov. Adriatic [Ref. Coen 1933]. ··1937
Coen, Note 264, pI. 10, fig. 91: Cerastoderma
686
Tropical Marine Mollusc Programme (TMMP)
paucicostatum rotundicosta -pazianum
(Sphaerocardium) paucicostatum Sowerby var. (anom.)
rarepartila Coen n. var. "1937 ZR: Eucardium
(Sphoerocardium sic) paucicostalum Sow planicosla
(Monts. mss.) rarepartita (v. anom.) var. Coen, 1937
nov. Adriatic . "1937 ZR: Sphaerocardium paucicostatum var. rarepartita Coen, 1933 nov. Venice. CHARACTERS AND OCCURRENCE - LOCALITY.
The Mediterranean.
paucicostatum rotundicosta Sacco, 1899 (variety). New combination : Acanthocardia paucicostata var.
rotundicosla (Sacco, 1899). Cardiidae . Recent.
RECORDS. "1899 Sacco, pI. 8, fig. 18: Cardium
paucicostalum var. rOlundicosla Sacco
*paucintermedia. - Homonymy to be noticed. Didacna
paucintermedi from 2 sources: Andrussov, 1924. Andrusov, 1963. - REMARKS . I do not know how
Andrusov 1963 compares with Andrussov 1924.
paucintermedia Andrusov, 1963. - Didacna paucintermedia Andrusov, 1963. Lymnocardiidae. Pliocene.
RECORDS . "1963 ZR: Didacna paucintermedia
Andrusov, 1963 sp. nov. Apsheron layers, Pliocene,
Russia . "1977 Popov: Hyrcania (Hyrcania)
paucintermedia (?) (Andrussow).
paucintermedia Andrussov, 1924. - Didacna
paucintermedia (Andrussov, 1924). Lymnocardiidae.
Pleistocene. RECORD S. "1924 ZR: Didacna
paucintermedia Andrussov, 1924 sp. n. Pleistocene.
paueri Gumbel, 1861. - Combination to be revised:
"Cardium" paueri Gumbel, 1861. Cardiidae. Fossil.
RECORDS. "1980 Ruhoff: Cardium paueri Gumbel,
1861 ; p. 662.
paulayi Vidal, 1999. - Acrosterigma paulayi Vidal, 1999.
Cardiidae. Neogene fossil. TYPE LOCALITY. Holotype,
Niue Island, Neogene fossil reef. TYPE REPOSITORY.
Holotype, USNM. TYPE MATERIAL. All material from
Niue Island, Neogene fossil reef, will be stored in
USNM, Paulay Collection Ref. FNIUE 2B . HoJotype:
An incomplete left valve. Paratype I: An incomplete
right valve. Para type 2: An incomplete right valve.
Paratype 3: an incomplete right valve. RECORDS. "1999
Vidal, p. 299, Table 23, Figs. 8G-I: Acrosterigma
paulayi n. sp. - CHARACTERS AND OCCURRENCE - DIMENSIONS . "1999 Vidal: Length 25 .5,
height 29.5 , width (calculated) 20.0 mm. REMARKS.
Acrosterigma paulayi is close to A. profundum; it differs
in having a more depressed lunule, shorter oblique
scales on posterior slope not encroaching upon the
anterior smooth zone, no squared ribs elsewhere, no or
less marked anterior serrations or scales on the ribs of
the median part, more pronounced herringbone
structure on the central shell surface. DISTRIBUTION. Only
known from Neogene fossil reef of Niue Island.
pauli Coquand, 1862. - Changed combination: Protocardia pauli (Coquand, 1862). Cardiidae. Cretaceous.
TYPE LOCALITY. Constantine. RECORDS. "1871 Stoliczka:
Cardium pauli Coquand, 1862 (species insufficiently
known). "1980 Ruhoff: Cardium pauli Coquand, 1862;
vol. 2: 204, pI. 10, figs . 5-6. "1934 ZR: Cardium pauli
Cretaceous [Ref. Blanckenhom 1934]. "1999 Jacques
Vidal (pers. com.), MNHN, Paris: Fossils from the
Paleontological Collection. Protocardia pauli
Coquand. Senonien-Cenomanien. Gabon (Gorges No.
204), R51370. Algeria, B19813, B19814.
pauli aequilaterale Blanckenhorn, 1934 (variety). -
Upgraded. Combination to be revised: "Cardium"
pauli aequilaterale Blanckenhom, 1934. Cardiidae.
Cretaceous. RECORDS. "1934 ZR: Cardium pauli var.
aequilaterale Blanckenhorn, 1934 nov. Cretaceous
Syria [Ref. Blanckenhom 1934].
paululum Eberzin, 1958. - Changed combination:
Euxinicardium paululum (Eberzin, 1958). Lymnocardiidae. Pliocene. RECORDS. "1958 ZR : Limnocardium (Euxinicardium) paululum Eberzin, 1958 sp.
nov. Kerchenski Peninsula, Crimea. Pliocene of the
Ponto-Caspian basin, Russia. "1987 Taktakishvili:
Limnocardium (Euxinicardium) paululum Ebers .
"1987 Taktakishvili: Euxinicardium paululum.
paupera Papaianopol, 1989. - Pachydacna paupera
Papaianopol, 1989. Lymnocardi idae. Pliocene. TYPE
LOCALITY. Bassin Dacique, Romania. RECORDS. "1984
ZR: Prosodacna (Prosodacna) paupera sp. nov.
Romania, Pliocene. Ref. to Papaianopol p. 98 . "1989
Papaianopol, p. 78, fig.: Pachydacna (Pachydacna)
paupera sp. nov.
pauperculum Meek, 1871. - Pleuriocardia pauperculum
(Meek, 1871). Cardiidae. Fossil. RECORDS. "1978 ZR:
Cardium pauperculum Meek, 1871 designated type
species for new subgenus Pleuriocardia
(Dochmocardia) . Ref. to Scott p. 894. "1987 Geary:
The transformation from Pleuriocardia subcurtum to
Pleuriocardia pauperculum is fairly abrupt. "1995
Schneider: P/euriocardia (Dochmocardia) paupercula
(Meek , 1871). Type species of Pleuriocardia
(Dochmocardia).
pauperculum Yokoyama, 1923. - Serripes nodai
Kafanov, 1980 new name. OTHER JUDGEMENTS: Serripes
groen/andicus (Bruguiere, 1789). » Regarding
Cardium pauperculum : Cardiidae. Miocene, Pliocene
and Recent. TYPE LOCALITY. Japan, Lower Miocene to
Upper Pliocene. RECORDS. "1923 Yokohama, p. 6, pI.
I, fig. 2: Cardium paupercu/um n. sp. "1923 ZR:
Cardium paupercu/um Yokohama, 1923 sp. n. Neogene
Japan . "1925 Yokoyama : Cardium pauperculum,
Yokoyama. "1973 Keen: C. pauperculum Yokoyama,
1923, not Meek, 1872 (fide Noda 1962) = Serripes
groenlandicus (Bruguiere, 1789). "1980 ZR: Cardium
pauperculum Yokoyama 1923 . Removal from
synonymy with Serripes groenlandicus (Bruguiere
1789) . Ref. to Kafanov. "1980 ZR: Cardium
pauperculum Yokoyama 1923. Preoccupied name,
replaced by Serripes nodai. Ref. to Kafanov, p. 319.
pauperculum jacksoni (subspecies). - Pleuriocardia
pauperculum jacksoni. Cardiidae. Fossil. RECORDS.
"1987 Geary: Pleuriocardia pauperculum jacksoni .
pausicostatum (error). - Printer's mistake for
paucicostatum in Glibert & Poel 1970.
pavlinovi (variety). - See under andnlssavi pavlinovi
Zhizhchenko, 1961.
paxfordensis Cox, 1946. - Tutcheria paxfordensis Cox,
1946. Cardiacea. Lower Lias . RECORDS. "194. ZR:
Tutcheria paxfordensis Cox, 1946. sp. nov. Lower Lias
of Pax ford brickyard 3 1/2 m, N.N.W. of Moreton-inthe-Marsh, Glos.
paziana (mode). - See under pazianum Dall, 1917.
pazianum Dall, 1917. - Changed combination : Nemocardium pazianum (Dall, 1917). OTHER JUDGEMENTS:
?Protocardia panamensis Dall, 1908. » Regarding
Protacardia paziana: Cardiidae. Recent. TYPE LOCALITY.
Phuket Marine Biological Center Special Publication vol. 30 (2004)
687
pecteniforme - pectinatum
··1917 Dall: Off la Paz, Gulf of California in 10
fathoms. TYPE REPOSITORY. Hypotype USNM Cat. No.
211618. Calf. Acad. Sci. Paleo. Type Col.!. ··1947
Hertlein & Strong: left & right valves, Santa Inez Bay,
east coast of Lower California, in 10-15 fathoms (1827 m), Plate 1, figs. 9, 12, 15. RECORDS. ··1916 ZR:
Protoeardia paziana Dall, 1916 sp. n. N.W. America.
··1917 Dall, p. 412: Protoeardia paziana n. sp.
resembling Protoeardia panamensis Dall. ··1947
Hertlein & Strong: Cardium (Microeardium) pazianum
= 7 Protoeardia panamensis Dall, 1908. ··1957
Emerson & Puffer: Cardium (Mieroeardium) pazianum
Dall, 1916. ··1958 Keen: Nemoeardium pazianum.
··1960 Shasky: Nemoeardium pazianum Dall. ··1961
Olsson: Mieroeardium pazianum. ··1964 Parker:
Nemoeardium (Mieroeardium)pazianum (DaIl, 1916).
··1967 Dushane & Poorman: Nemoeardium pazianum
Dall. ··1971 Keen: Nemoeardium pazianum Dall.
··1971 Keen & McLean: Nemoeardium (Mierocardium) pazianum (Dall, 1916). ··1974 Abbott:
Nemoeardium (Mieroeardium) pazianum (Dall, 1916).
··1977 Fischer-Piette: Nemoeardium pazianum Dall.
··1977 Popov: Pratulum pazianum (Dall, 1916). CHARACTERS AND OCCURRENCE - OF COLLECTIONS. ··ZMUC: Gulf of California, 64 m sand, 61 m
sand, 73 m shelly sand, 61 m shelly sand, 92-119 m
silty clay, 46-57 m sand, 33 m sand. CHARACTERS of
diagnostic value: shell about as high as long; various
ribs on posterior area with strong scales only.
DESCRIPTIONS. ··1917 Dal1: shell small and delicate,
resembling P panamensis Dall, but smaller, more
delicate, more elongated, and with the spinose posterior
sculpture much more prominent when preserved intact;
sculpture of about 40 anterior minutely nodulous ribs,
each giving the effect of a string of beads, and to the
interspaces of punctuation; there are about 22 ribs on
the posterior area, which, when intact, carry close-set
spinules; the inner margins are sharply serrate; the hinge
margin near the umbones has a tinge of crimson,
otherwise the shell is yellowish white. ··1947 Hertlein
& Strong: shell thin, rounded, posteriorly somewhat
obliquely produced and subtruncated; the anterior area
ornamented by about 40 fine ribs which are finely
nodulous where the concentric crosses the radial
sculpture; posterior area with about 22 ribs which are
minutely scaled, but interspersed in this series there
are 6 with comparatively large scales; the colour of the
shell is yellowish-white, some specimens entirely so,
but are ornamented by vivid red and orange interiorly
toward the hinge and occasionally there is exteriorly a
narrow concentric band which is tinged with this colour.
DIMENSIONS. ··1917 Dall: length 10, height 8.6, diameter .
6 nun. ··1947 Hertlein & Strong: about 15 mm, both
in height and length. ··1971 Keen: fairly abundant in
deep water but has never been taken ashore. It is
sculptured with some 60 fine ribs, the posterior ribs
bearing fine scales. The color is yellowish white to red
or orange, uniformly or in bands. Length and height,
about 15 mm. REMARKS. This species is very close to
Protoeardia panamensis Dall, 1908, a Panamanian
species which was described as possessing a shorter
shell with about 33 ribs and the interior white and
polished. ··1947: Hertlein & Strong doubted whether
C. pazianum is really distinct from Protoeardia
panamensis. DISTRIBUTION. ··1971 Keen & McLean:
Cedros Island, Gulf of California to Panama. HABITAT.
··ZMUC: Sand, shelly sand, silty clay 33-119 m depth.
DISTRIBUTION. Gulf of California.
pecteniforme (error). - Printer's mistake for pectin iforme in Wood and other authors.
*pectinatum. - Homonymy to be noticed. Cardium
peetinatum from 8 sources: Linne, 1758. - Born 1778.
- Lamarck, 1819. - Kloden, 1834. - Tausch, 1884. Yales & Bean (7year). - Goldfuss in Decken (7year).Hoernes (7year).
pectinatum Born 1778. - Identical with Cardium
peetiniforme Born, 1780.» Regarding C. peetinatum:
Cardiidae. Recent. RECORDS. ··1778 Born, G 17.
Cardium peetinatum Ref to Mus. Caes. Vindobon.
··1780 Born: Cardium peetinatum in Born 1778 =
Cardium peetiniforme Born, 1780.
pectinatum Bruguiere. - Identical with Lyroeardium
peetinatum (Linne, 1758). » Regarding Cardium
peetinatum: RECORDS. ··1844 Potiez & Michaud:
Cardium peetinatum Brug. = Cardium aeolieum Born.
pectinatum Goldfuss in Decken. - Unknown species.
» Regarding Cardium peetinatum: Fossil. RECORDS.
··1857 Deshayes: Cardium peetinatum Goldf. in
Decken. Spec. incogn.
pectinatum Hoernes. - Synonym of Divarieardium
aquitanieum Mayer. » Regarding Cardium
peetinatum: Cardiidae. Fossil. RECORDS. ··1886 Dollfus
& Dautzenberg: C. peetinatum Hoernes = Cardium
(Divarieardium) aquitanieum Mayer.
pectinatum Kloden, 1834. - Name replaced: "Cardium
kloedeni" Deshayes. » Regarding Cardium
peetinatum: Cardiidae. Fossil. RECORDS. ··1857
Deshayes: Cardium peetinatum Kloden - C. klodeni
Desh. ··1922-32 Sherborn, p. 4802: Cardium
peetinatum Kloeden, 1834.
pectinatum Lamarck, 1819. [Figs. Part 3 p. 850]. Synonym of Cerastoderma edule (Linne, 1758). OTHER
JUDGEMENTS: Lyroeardium lyratum (Sowerby, 1834).Erroneous ID of Lyrocardium lyratum by Melvill &
Sykes (1898). Their material was collected at the
Andaman Islands. - Cardium peetiniforme Born. »
Regarding Cardium peetinatum: Cardiidae. Recent.
RECORDS. ··1819 Lamarck, p. 12: Cardium peetinatum.
··1836 Philippi: Cardium pee tina tum Lamk. 7. ··1853
Doublier: Cardium peetinatum Lamarck. ··1839 Anton:
Cardium pectinatum 7 Lam (non Linn.) = 7 C.
peetiniforme Born. ··1856 Wood & Hanley: C.
peetinatum Lam. = Cardium edule Lin. ··1857
Deshayes: Cardium peetinatum Lamarck, 1818 sieC. edule7. ··1866 Brusina: Cardium peetinatum
Lamarck. ··1867 Weinkauff: Cardium peetinatum
Lamarck = Cardium edule Linne. ··1867 Weinkauff:
Cardium peetinatum sic Philippi = Cardium edule
Linne. ··1868 ZR: Cardium peetinatum (L.) probably
not = aeolieum (Born) [Ref. Romer 1868]. ··1869 ZR:
Section, subtypical - 3. Lyrasculptae: peetinata (L.)
and lyrata (Sow.) [Ref. Graham-Ponton 1869]. ··1869
Romer: Cardium peetinatum Lamarck = Cardium edule
L. ··1880 Stossich: Cardium peetinatum, in Philippi.
1836 = Cardium edule Linne. ··1886 Locard: Cardium
peetinatum, de Lamarck, 1818 = Cardium edule, Linne,
1767. ··1887 Hoyle: Cardium peetinatum Lamarck.
··1888 Kobelt: Cardium peetinatum Lam. = Cardium
688
Tropical Marine Mollusc Programme (TMMP)
peetinatum - peetiniforme
edule Linne. --1890 Paete1: Cardium pee/ina/um Lm.
edule L. --1891 Dando & Sandri: Cardium
peelina/um Lam. --1898 Me1vill & Sykes:
Laevieardium pee/ina/um Lamarck = Lyroeardium
lyra/um [cited from Fischer-Piette 1977]. --1902
Dollfus & Dautzenberg: Cardium pee/ina/um Lamarck,
1819 (non Linne) = Cardium (Cerasloderma) edule
Linne, 1758. --1945 ZR: Cardium pee/inalum Lk. (non
L) [Ref. Lamy 1945]. --1977 Fischer-Piette: Cardium
peelinalum Lamarck = Cardium edule Linne.
peetblatum Linne, 1758. [Figs. Part 1 p. 75, 125. Part 3
p. 899]. - Lyroeardium p ee/inalum (Linne, 1758).
OTHER JUDGEMENTS: Lyroeardium aeolieum (Born,
1780). - Nomen dubium . - Valid species. » Regarding
C pee/inalum: Cardiidae. Tertiary and Recent. TYPE
LOCALITY. Mediterranean (Linne 1758), Guinea
(Spengler 1799); unknown (Reeve 1844). RECORDS.
--1742 Gualtieri . 10 by Linnaeus 1758: Cardium
pee/inalum t. 75. f. A. --1758 Linne, p. 681, No. 79:
Cardium peelinalum. --1764 Linne, p. 492, No. 47 :
Cardium peelinalum. --1767 Linne, p. 1124, No. 92:
Cardium peelinalum. --1771 Knorr: Cardium
peelina/um (fide Spengler 1799). --1782 Chemnitz:
Cardium peelinalum . --1786 Schroter, No . XX:
Cardium peelinalum Linn. XII . pag. 1124. Spec. 92.
--1788 Schroter: Cardium pee/in alum [Reference to
Chemnitz (1782) vol. VII, fig. 30, and fig. 188]. --1789
Bruguiere, No. 18. Cardium pee/inalum Linn. -- I 791
Gmelin No. 24: Cardium peelina/um. --1799 Spengler:
Cardium pee/in alum Linnei Sp. 92. p.1124 (from the
coast of Guinea). --1802 Bosc: Cardium peelinalum.
--1807 Fischer: Cardium pee/ina/um. --1815 Wood:
Cardium peelina/um Linn. Gmel. --1817 Dillwyn:
Cardium pee/ina/um Linnaeus. --1817 Dillwyn:
Linnaeus, for Cardium peelinalum, has erroneously
referred to Gualter. t. 75, f. A. which is Venus peelina/a.
--1823 Dillwyn, Tab. 314 fig. 150: Cardium
peelina/um. --1836 Bosc: Cardium pee/ina/um . --1844
Philippi: Cardium pee/inalum = Cardium edule L
[fossil]. --1844 Reeve, species 14: Cardium pee/ina/um
Linnaeus (not of Lamarck, nor Philippi), Mus. Ulric.
Deshayes, note in new edition of Lamarck, vol. vi. p.
404. --1853 Menke: Cardium pee/inalum L = C
aeolieum Born, 1780. --1857 Deshayes : Cardium
pee/ina/um Linne, 1780 sic. --1858 Adams & Adams,
p. 457: Laevieardium peelina/um Linn. --1864 Mayer:
Cardium pee/ina/um. Linne. [Tertiary] . --1877 Marrat:
Cardium (Laevieardium) peelinalum L --1884 Grasset:
Laevieardium pee/ina/um L --1899 Sacco: Diseors
peelina/us (L.) [Recent]. --1902 Dollfus &
Dautzenberg: C pee/inatum Linne = C. aeolicum Born.
--1902 Sherborn , p. 724: Cardium peelinalum
Linnaeus, 1758. --1906 Dautzenberg & Fischer:
Cardium pee/ina/um Linne. --1952 Dawydoff:
Laevieardium peelinalum Linne. --1952 Dodge :
Cardium pee/inalum = species insufficiently known.
--1977 Fischer-Piette: Cardium pee/inalum L = C
aeolieum Born, 1780. --1977 Wilson & Stevenson:
Lyroeardia pee/inalum (Linnaeus, 1758) (= aeolieum
Born, 1778). --1989 Voskuil & Onverwagt: Cardium
pee/inalum Linnaeus, 1758; nomen dubium. --1998
Poutiers: Cardium peeliniforme Born, 1780. --1999
Higo el al.: Cardium peeliniforme Born, 1780 =
Aeros/erigma (Regozara) jlava (Linnaeus, 1758).
=
--2003 Hylleberg & Kilburn, p. 183 : Lyroeardium
pee/ina/um (Linnaeus, 1758). West African species. CHARACTERS AND OCCURRENCE - DESCRIPTIONS . --1799 Spengler: /esla subeorda/a pee/ina/a.
--1799 Spengler: posteriorly, one half of the ribs set
with elevated stripes. The other half transversally
covered with folds which, like the scales, are arranged
from below upwards, i.e. an angular pattern of stripes
or folds . --1844 Reeve: shell gibbously globose,
radiately striated, striae numerous , close-set, the
posterior elevated, the anterior somewhat obsolete and
obliquely crossed with narrow slightly wrinkled plaits,
the extreme posterior area of the shell smooth; white,
everywhere painted with waved pink spots. DIMENSIONS.
--1799 Spengler: length : 47 mm (1.75 Tom.), width 44
mm (I Tom. 8 Lin.). REMARKS. Linnaeus has created
confusion because he gave reference to Venus peelina/a
in Gualtieri (1742), pI. 72, fig. A. However, the shell
of Cardium peelina/a is so characteristic that mistake
is impossible (Spengler 1799). Furthermore, following
Deshayes, Reeve 1844 restored the Linnaean name
Cardium pee/inalum as a senior synonym of Cardium
aeolieum Born, 1780. This restoration was correct, but
has not been observed by most modem authors. --1884
Tauch: Cardium pee/ina/um Linne is also known from
the "Wiener Becken" [Tertiary]. DISTRIBUTION. --1799
Spengler: Guinea [West Africa]. --1865 Reibisch:
Mediterranean. --1884 Grasset: Gabon. --1906
Dautzenberg & Fischer: Indian Ocean.
peetinatum Menke 1853. - Misidentification of
Lyroeardium pee/in alum (Linne, 1758). Cardiidae.
Recent. RECORDS. --1901 Dall: Cardium pee/in alum
(Linnaeus) Menke from St. Vincent of the Cap Verde
Islands. --1906 Dautzenberg & Fischer: Cardium
pee/inalum Menke 1853, not Linne 1758 = Cardium
(Diseors) aeolieum Born.
peetinatum of Philippi. - Misidentification of
Cerasloderma edule (Linne) . Cardiidae . Recent.
RECORDS. --1917 Hidalgo : Cardium peelina/um in
Philippi = Cardium edule Linne.
peetinatum Tausch, 1884. - Misidentification of
Lyroeardium pee/inalum (Linne, 1758). Cardiidae.
Recent. RECORDS. --1906 Dautzenberg & Fischer:
Cardium peelina/um Tausch, 1884, not Linne 1758 =
Cardium (Diseors) aeolieum Bom.
peetinatum Wood . - Not a cardiid. » Regarding
Cytherea pee/ina/um: identical with C gibba, Lam.;
Circe g. Sow. Red Sea; Philippines (Wood & Hanley
1856). RECORDS. --1856 Wood & Hanley, No. 30:
Cytherea pee/ina/um Wood (not Lin. nor Lam.).
peetinatum Yales & Bean. - Replaced by Cardium
. /runea/um Sowerby. » Regarding Cardium
pee/ina/um: Cardiidae . Fossil. RECORDS. --1857
Deshayes : Cardium pee/ina/um Yales et Bean - C
/runca/um Sow.
*peetiniforme. - Homonymy to be noticed. Cardium
pee/iniforme from 5 sources: Born, 1780. - Bosc in
Buffon, 180 I. - Megerle, 1811. - Wood, 1815. - Muller,
1859.
peetiniforme Born, 1780. [Figs. Part 1 p. 48, 67,89. Part
3 p. 863, and inside of front cover] . - Vas/ieardium
pee/iniforme (Born , 1780). OTHER JUDGEMENTS:
Cardium magnum vaL gamma Gmelin, 1791. Cardium jlavum Linnaeus, 1758. » Regarding C
Phl/kel Marine Biological Center Special Publication vol. 30 (2004)
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pecliniforme - pede/ayense
peetinijorme: Cardiidae. Recent. REcORDs. ··1780 Born,
p. 49, pI. 3, fig. 9: Cardium peelinijorme. ··1782
Chemnitz, Tab. 17 Fig. 180: Cardium peclinijorme
Bornii . ··1788 Schroter: Cardium pecliniforme.
[Reference to Chernnitz (1782) fig. 188, sic should be
fig . 178]. ··1789 Bruguiere, No. 23. Cardium
pectinijorme; Born. ··1789 Bruguiere: Cardium
peclinijorme Bornii; Martini, tom. 6. pag. 188, tab. 17,
fig. 180. "1815 Wood: Cardium peclinijorme Born.
··1817 Dillwyn: Cardium peclinijorme Born. ··1828
Wood: Cardium pectenijorme sic Born. ··1840 Pfeiffer:
Chemnitz Fig. 180. Cardium pectinijorme Bornii =
Cardium magnum gamma Gm. 16. "1856 Wood &
Hanley, No. 37: Cardium peclini/orme Born, T. Caes.
49, pI. 3, f. 10. "1869 Romer: Cardium peclinijorme
Bornii Chemnitz (juvenile) = C. jlavum L. "1878
Brauer: Cardium pecliniforme Born. "1889 ZR:
Cardium pectinijorme with fig. [ref. Holzapfel]. ··1891
Fischer: Cardium pectinijorme Born. ··1903 Hidalgo:
Cardium peclinijorme Born = Cardium jlavum Linne,
1758 . ··1929 Dautzenberg: Cardium peclinijorme
Born. ··1932 Dautzenberg: Cardium peclinijorme
Born. ··1932 Prashad: Cardium peclinijorme Born,
1780. ··1937 Serene: Cardium p eelinijorme Born.
··1939 Adam & Leloup: Cardium peclinijorme Bornii
Chemnitz = Laevicardium (Trachycardium) jlavum
(Linne 1758). ··1955 Rost & Soot-Ryen: Trachycardium peclinijorme Born, 1780. ··1956 ZR: Trachycardium pectinijorm sic Born, new to Seychelles [Ref.
Rost & Soot-Ryen 1956]. ··1961 Salvat: Cardium
pecliniforme Born. ··1974 Barnard: Cardium
pectinijorme in Dautzenberg 1929 = Cardium jlavum.
··1977 Fischer-Piette: Trachycardium pectinijorme
(Born, 1780) = Laevicardium jlavum Linne auctorum.
··1990 Abbott & Dance: Trachycardium pectinijorme.
"1991 Voskuil & Onverwagt p. 58: Trachycardium
peclinijorme (Born, 1780). ··1997 Vidal : Vasticardium
peclinijorme (Born, 1780). ··1999 Vidal, p. 354:
Cardium peclinijorme Born, 1780 is senior syn. of
Vasticardium rugosum (Lamarck, 1819). ··2000
Hylleberg, p. 532: Vaslicardium pectinijorme (Born,
1780). ··200 I Vidal, p. 407: Vasticardium pectinijorme.
··2002 Hylleberg & Kilburn, p. 64: Vasticardium
pectil1lforme (Born, 1778). ··2003 Hylleberg &
Kilburn, p. 185, Plate 4, figs. 15 a,b,c: Vasticardium
pectinijorme (Born, 1778). - CHARACTERS AND
OCCURRENCE - DESCRIPTIONS . ··200 I Vidal:
tentacles numerous. No eyes on tentacles. REMARKS.
The synonymy suggested by Prashad (1932) is
erroneous, maybe caused by a printer'S error in
Explicatio Tabularum (Table of Contents), Born (1780).
Table III fig. 9 is listed as Cardium pectinijorme, which
is correct on the Pl.ate. However, Cardiumjlavum has
No.7 in the Table, but No. lOon the Plate. DISTRJBUTION.
··1856 Wood & Hanley: unknown. "1991 Voskuil &
Onverwagt, "1994 Hylleberg: Red Sea, Indo-West
Pacific. ··2000 Hylleberg: Vietnam and Cambodia.
··2002 Hylleberg & Kilburn : South India . ··2003
Hylleberg & Kilburn: Vietnam.
pectiniforme Bose in Buffon, 1801. - Combination to
be revised: "Cardium" peclinijorme Bosc in Buffon,
1801. Cardiidae. Fossil. RECORDS. ··1922-32 Sherborn,
p. 4805: Cardium pectinijorme Bosc in Buffon, 1801.
pectiniforme Megerie, 1811. - Combination to be
revised: "Cardium" pectiniforme Megerie, 1811 .
Cardiidae. Fossil. RECORDS. ··1922-32 Sherborn, p.
4805: Cardium pectinijorme Megerle, 1811.
pectiniforme Miiller, 1859. - Combination to be revised:
"Cardium" pectinijorme J. Miiller, 1859. Cardiidae.
Cretaceous. RECORDS. ··1871 Stoliczka: Cardium
pectinijorme Miiller (Stoliczka : unfamiliar species).
··1980 Ruhoff: Cardium pectinijorme J. Miiller, 1859;
p.29 .
pectiniforme Wood, 1815. - Combination to be revised :
"Cardium " pectinijorme Wood , 1815. Cardiidae.
Fossil. REcORDs. ··1922-32 Sherborn, p. 4805: Cardium
pectinijorme Wood, 1815.
pectinioides Link, 1807. - Combination to be revised:
"Cardium" pectinioides Link, 1807. Cardiidae. ?Fossil.
RECORDS. ··1922-32 Sherborn, p. 4805: Cardium
pectinioides Link, 1807.
pectinoideum O.G. Costa, 1846. - Combination to be
revised: "Cardium" pectinoideum O.G. Costa, 1846.
Cardiidae. Fossil. RECORDS. ··1922-32 Sherborn, p.
4806: Cardium pectinoideum O.G. Costa, 1846.
pectuncu/oides d ' Archiac & Verneuil, 1842. Combination to be revised: "Cardium "pectuncu/oides
d ' Archiac & Verneuil, 1842. Cardiidae. Fossil.
RECORDS . ··1922-32 Sherborn, p. 4810: Cardium
pectunculoides d'Archiac & Verneuil, 1842.
Pectunculus Adanson (genus). Non binominal. - Not
available.» Regarding Pectunculus: Cardiidae. Fossil
and Recent. REcORDs. ··1857 Deshayes: Pectunculus
in Belon, Lister, Petiver, Rumphius, Adanson, Martini
[authors not available]: syn. of Cardium . ··1876 Meek:
Cardium asiaticum Lam. regarded as genotype of
Pectunculus(Adanson), Stoliczka. ··1847 Gray, No.
557: Pectunculus, Adans. [synonym of Cardium, Linn].
··1858 Adams & Adams: Sub-gen . Peclunculus
Adanson. ··1869 ZR: subgenus Pectunculus (Adanson,
March), type Cardium asiaticum (Brug.) [Ref. Romer
1869]. ··1876 Meek: Pectunculus (Adanson),
Stoliczka, 1870 (as a subgenus of Cardium). Synonym
ofCardiidae. ··1980 Vokes: Pectunculus (ex Adanson)
March, 1853 [non Da Costa, 1778 (Veneridae)]. ··1980
Vokes: Peetunculus (genus) "Adanson" Stoliczka, 1870
[non Da Costa, 1778 (Veneridae)].
Pectunculus Adanson (subgenus). Non binominal. - Not
available. RECORDS. ··1871 Stoliczka: (Peetunculus)
Adanson. Type species Cardium asialicum .
Pectunculus Mart. (genus). Non binominal. Not
available. - Synonym of Trachyeardium. » Regarding
Pectunculus: Cardiidae. Fossil and Recent. RECORDS.
··1858 Adams & Adams: subgenus Pectuneulus Mart.,
not Adanson is a synonym = subgenus Trachycardillm,
March.
Pectuncu/us Paetel, 1875(genus). - Nomen nudum.
RECORDS. ··1980. Vokes: Pectunculus "Romer" Paetel,
1875 [n.n.] . In subfamily Cardiinae.
peculiare - Conocardium peeutiare. Conocardiidae.
Fossil. RECORDS. ··1960 ZR: Conocardium pecutiare
[Ref. Branson 1960a].
pede/ayense Cossmann, 1921. - Changed combination:
Fragum pedelayense (Cossmann, 1921). Cardiidae.
Eocene. RECORDS. ··1922 ZR: Cardium (Parvicardium)
pedelayense Cossmann, 1922 sp. n. Eocene. ··1977
Popov: Fragum (Fragum) pedelayense (Cossmann,
1921).
690
Tropical Marine Mollusc Programme (TMMP)
pedernale - peraltaensis
pedernale Bonn, 1852. - Changed combination:
Protocardium pedernale (Bonn, 1852). Cardiidae.
Fossil. RECORDS. '-1871 Stoliczka: Cardium pedernale
Bonn sic, 1852 (Protocardium).
pedernalense Pilsbry & Olsson, 1941. - Synonym of
Laevicardium substriatum (Conrad, 1837). »
Regarding Laevicardium pedernalense: Cardiidae.
Pliocene. RECORDS. "1941 Pilsbry & Olsson, pp. 10,
60 (pI. 14, fig. 2): Laevicardium pedernalense new
species. --1941 ZR: Laevicardium pedernalense Pilsbry
& Olsson, 1941 sp. n. Pliocene, Jama formation, Puerto
Jama, Western Ecuador. --1977 Popov: Laevicardium
pedernalense (Pilsbry & Olsson, 1941). --2000 Coan
et al.: Laevicardium pedernalense Pilsbry & Olsson,
1941 = Laevicardium substriatum (Conrad, 1837).
pedernalis Roemer, 1849. - Nomina oblita. »
Regarding Cardium pedernalis: Cardiidae. Cretaceous.
TYPE LOCALITY . Cretaceous of Texas. RECORDS. "192232 Sherborn, p. 4811 : Cardium pedernalis C. F. Roemer,
1849. --1986 Scott: Cardium pedernalis Roemer (1849)
considered nomina oblita.
peetinatum (error). - Printer's mistake for pectinatum
in Weinkauff 1867.
pelagi - Combination to be revised: "Cardium" pelagi.
Cardiidae. Cretaceous. RECORDS. --1871 Stol iczka:
Cardium pelagi (Laevicardium) .
pellati Loriol in Loriol & Pellat, 1867. - Combination
to be revised: "Cardium" pellati Loriol in Loriol &
Pellat, 1867. ?Cardiidae. Lias and Jurassic. RECORDS.
--1980 Ruhoff: Cardium pellati Loriol in Loriol &
Pellat, 1867; vol. 19: 58, pI. 6, figs . 1-2. "1999 Jacques
Vidal (pers. com.), MNHN, Paris: Fossils from the
Paleontological Collection. Cardium pellati de Loriol.
Portlandien. France (Pas de Calais), S05323.
pellucidum Schroeter, 1802 . - Combination to be
revised: "Cardium" pellucidum Schroeter, 1802.
?Cardiidae. Fossil. RECORDS. "1922-32 Sherborn, p.
4824: Cardium pellucidum Schroeter, 1802.
pelonatense Cossmann & Peyrot, 1911. Changed combination: Acanthocardia pelonatense (Cossmann &
Peyrot, 1911). Cardiidae. Fossil. RECORDS. "1902
Dollfus & Dautzenberg: Cardium pelonatense
Cossmann & Peyrot. --1911 ZR: Cardium pelonatense
Cossmann & Peyrot, 1911 sp. n. France Neogene.
--1977 Popov: Acanthocardia (Acanthocardia)
pelouatensis sic (Cossmann & Peyrot, 1911).
pelouatensis (error). - Printer's mistake for pelonatense
in Popov 1977.
pelzelni Brusina. - New combination: Lymnocardium
pe/zelni (Brus.). Lymnocardiidae. Fossil. RECORDS.
--1987 Taktakishvili: Limnocardium pelzelni (Brus.)
pendens Cragin, 1893. - Changed combination: Granocardium pendens (Cragin, 1893). Cardiidae. Cretaceous. RECORDS. --1978 Scott: Protocardia pendens
Cragin, 1893. "1978 Scott: Granocardium (Granocardium) pendens (Cragin, 1893). "1978 ZR:
Granocardium (Granocardium) pendens (Cragin 1893)
Description. Ref. to Scott p. 897 .
penderense Stephenson, 1923 . - Combination to be
revised: "Cardium" penderense Stephenson, 1923.
Cardiidae. Cretaceous. RECORDS. --1923 ZR: Cardium
penderense Stephenson, J923 sp. n. Cretaceous U.S.A.
pennantii Beck, 1842. [Figs. Part I p. 132]. - Synonym
of Laevicardium crass urn (Gmelin, 1791). OTHER
JUDGEMENTS: Laevicardium laevigatum (Pennant). Laevicardium norvegicum (Spengler 1799), both
synonyms of Laevicardium crassum. » Regarding
Cardium pennantii: Cardiidae. Recent. TYPE REPOSITORY.
3 syntypes. 1978130. BMNH London. TYPE LOCALITY.
Exmouth, Devon, Britain (Reeve 1845). RECORDS.
--1842 Beck: Cardium pennantii. --1844 Reeve, species
48: Cardium pennantii Beck , MSS., H. M. The King
of Denmark's Museum. --1846 Loven: "Cardium
pennanti Beck", Reeve = Cardium norvegicum Spengl.
-'1856 Wood & Hanley: C. pennantii Reeve 1. Car. f.
48 = Cardium laevigatum Penn. --1858 Adams &
Adams, p. 457: Laevicardium pennantii Beck. "1859
Sowerby: C. Pennantii = Cardium norvegicum
Spengler. --1867 Mitchell: Cardium pennantii Beck =
Laevicardium crassum Gmelin in Fischer-Piette (1977,
p.12). "1867 Hidalgo: Cardium pennantii, Reeve =
Cardium norvegicum Spengler. --1867 Weinkauff:
Cardium Pennanti sic Reeve = Cardium norwegicum
sic Spengler. --1868 ZR: Cardium pennanti and
oblongum of Reeve to be distinguished from the
Mediterranean oblongum (Chemn.) =flavum (Born) =
sulcatum (Lam.) [Ref. Romer 1868]. "1869 Romer:
Cardium Pennanti Reeve fig. 48 = Cardium
norvegicum Spengler. --1888 Kobel!: Cardium
Pennanti Reeve = Cardium norvegicum Spengler.
--1890 Hidalgo: Cardium pennantii Reeve = Cardium
norvegicum Spengler. --1890 Paetel: Cardium pennanti
Bk. = norwegicum Spglr. "1912 Dautzenberg &
Fischer: Cardium penn anti Beck, Reeve 1844 =
Cardium (Laevicardium) norvegicum Spengler 1799.
--1917 Hidalgo: Cardium pennanti Reeve = Cardium
norvegicum Spengler. --1922-32 Sherborn, p. 4833:
Cardium pennantii Reeve (ex Beck MS), 1844. --1977
Fischer-Piette, p. 12: Cardium pennantii (Beck ms.) =
Laevicardium crass urn Gmelin. - CHARACTERS
AND OCCURRENCE - DESCRIPTION. "1844 Reeve:
shell depressly cordate, rather oblique, very
inequilateral, radiately very faintly grooved, posterior
area smooth; dirty yellowish colour covered with a thin
smooth earthy epidermis, interior white. REMARKS.
"1844 Reeve: The grooves of this species, which has
been confounded with the C. laevigatum, are almost
obsolete. HABITAT. -'1844 Reeve: found in sandy mud
in deep water. DISTRIBUTION . '-1844 Reeve: Hab.
Exmouth, Devon.
pennantii Reeve 1844. [Figs. Part 3 p . 930]. - See under
pennantii Beck, 1842.
pellslii Fuchs, 1870. - Changed combination:
Pannonicardium pens/ii (Fuchs, 1870). Lymnocardiidae . Fossil. RECORDS. --1970 Glibert & Poel:
. Limnocardium penslii (Fuchs, 1870). --1987
Taktakishvili: Limnocardium penslii (Fuchs). "1987
Taktakishvili: Pannonicardium penslii.
penslii patrulii (subspecies). - Pannonicardium penslii
patrulii. Lymnocardiidae. Fossil. RECORDS. --1987
Taktakishvili: Pannonicardium penslii patruW.
pentagonalis (subspecies) . - See under planum
pentagonalis Huica, 1977.
peraltaellsis Fiirsich & Werner, 1985. - Protocardia
peralta ens is Fiirsich & Werner, 1985. Cardiidae. Upper
Jurassic. TYPE LOCALITY. Portugal. RECORDS. "1985
Fiirsich & Werner, p. 440, figs. I c-e, 3, 4d: Protocardia
(Protocardia) peraltaensis n. sp . --1985 ZR:
Phuket Marine Biological Center Special Publication vol. 30 (2004)
691
peramabile - peregrinum
Protocardia (Protocardia) peraltaensis Fursich et al.,
1985 sp. nov. Portugal , Upper Jurassic. Ref. to Fursich
et al. p. 440.
peramabile (mode ). - See under peramabiles.
peramabilis DaB, 1881. - Microcardium peramabilis
(Dall, 1881). Cardiidae. Recent. Type species of
(Microcardium) Thiele, 1934 raised to generic level.
TVPE LOCALlTV. Off Yucatan. RECORDS. "1881 Dall, p.
132-133: Cardium (Fulvia) peramabilis n. s. "1881
ZR : Cardium (Fulvia) peramabilis, sp. n., Dall, 1881.
Florida and Barbados, 54-III fath. ··1882 Verrill:
Cardium peramabilis Dall. ··1884 Verrill: Cardium
peramabilis DaB. ··1886 Dall: Cardium (Fulvia)
peramabilis Dall. ··1889 Dall: Cardium peramabilis
Dall. ··1899 Sacco: Cardium peramabile Dal1. ··1900
Dall: Protocardia peramabilis Dall . ··1902 Dall:
Protocardia peramabilis DaB, 1886. ··1902 Dall &
Simpson: Protocardia peramabilis Dall. = Cardium
peramabile Dall, 1881. ··1915 Johnson: Cardium
peramabilis Dall. ··1921 Maury: Protocardia
peramabilis Dall, 1881 (as Fulvia). ··1937 Keen:
designated Cardium peramabile Dall, 1881 as type
species of the subgenus (Microcardium) Thiele, 1934.
Recognised at the generic level by authors. ··1937
Smith, M: Protocardia peramabilis Dall. ··1937 ZR:
Cardium (Fulvia) peramabilis Dall 1881 (= Cardium
peramabile Dall) here amended" [Ref. Keen 1937] .
··1939 ZR: Microcardium peramabile fig. [Ref.
McLean 1939]. ··1947 Hertlein & Strong: Cardium
(Micro cardium ) peramabile Dall, 1881. ··1958 Abbott:
Nemocardium (Microcardium) peramabilis Dall.
··1962 Warmke & Abbott: Microcardium peramabile
Dall, 1881. ··1974 Abbott: Nemocardium peramabile
(Dall, 1881). ··1977 Popov: Pratulum peramabilis
(Dall, 1886). ··1944 ZR: Microcardium peramabile
Dall fig. [Ref. Clench & Smith 1944]. ··1995
Schneider: Nemocardium (Microcardium) peramibilis
sic (Dall, 1881). ··1997 Shelton: Nemocardium
peramabile (Dall, 1881). ··200 I Schneider & Carter:
Nemocardium (Microcardium) peramibilis sic (Dall,
1881). Cardiidae. Recent. - CHARACTERS AND
OCCURRENCE - DESCRIPTIONS. ··1902 Dall &
Simpson: shell small, slightly inequilateral, rather thin
and fragile, inflated; posterior area sculptured with a
great number of fine, radiating, nearly smooth ribs, the
spaces between them being slightly pitted, the whole
being covered with a fine epidermis, which is scaly on
the ribs; the anterior 3/5 of the shell is sculptured with
equally fine, radiating ribs, and these are crossed by
very numerous somewhat fainter, concentric ridges,
leaving the surface minutely beaded ; borders of the
shell delicately and sharply serrate, not gaping; within
the shell the outside patterns of sculpture may be plainly
seen. Colour whitish Soft body: The foot is keeled
below but not serrate. DIMENSIONS. ··1902 Dall &
Simpson: 10 x II x 9 mm (McLean 1939); length 10,
height II, diameter 9 mm . REMARKS. ··1921 Maury:
the recent species is Protocardia peramabilis Dall , 1881
(as Fulvia). The ancestral form appears to be
Protocardia islahispaniolae Maury, Miocene. HABITAT.
··Of authors: 150 m depth. DISTRIBUTION . ··Of authors:
Rhode Island, West Indies, Mexico, Brazil.
peramabilis tinctum Dall, 1881 (subspecies). - Changed
combination: Microcardium tinctum (Dall, 1886).
OTHER JUDGEMENTS: Microcardium peramabilis tinctum
(Dall, 1881). » Regarding C. peramabilis var. tinctum:
Cardiidae. Recent. RECORDS . ··1881 Dall: Cardium
peramabilis var. tinctum . ··1886 Dall: Cardium (Fulvia)
peramabile var. tinctum. ··1889 Dall : Cardium
peramabilis var. tinctum Dall . "190 I ZR: Cardium
peramabile var. tinctum Dall is a good species [Ref.
Dall & Simpson 1900]. ··1902 Dall & Simpson :
Cardium peramabile var. tinctum Dall, 1886 =
Protocardia tincta Dall. ··1947 Hertlein & Strong:
Cardium peramabilis tinctum DaB = tinctum Dall,
1886. ··1951 McLean: Microcardium peramabile var.
tinctum (Dall, 1886) = Microcardium tinctum (Dall).
peramibilis (error). - Printer 's mistake for peramabile
in Schneider 1995 .
perarensis Gillet, 1970. - Euxinicardium ? perarensis
Gillet, 1970. Lymnocardiidae. Miocene. RECORDS.
··1970 ZR: Euxinicardium ? perarensis Gillet, 1970
sp . nov. ' Montei dei Corvi' deposits, Ancona district,
Italy
. .
perdita Papaia nopol, 1981 . - Pontalmyra perdita
Papaianopol, 1981. Lymnocardiidae. Upper Miocene
and Recent. RECORDS. ··1981 ZR: Pontalmyra
(Pontalmyra) perdita Papaianopol, 1981 sp . nov.
Romania. Ref. to Papaianopol, p. 36. ··1987
Taktakishvili: Pontalmyra perdita Papaian.
peregrina Papaianopo1, 1980. - Luxuridacna peregrina
Papaianopol, 1980. Lymnocardiidae. Pliocene .
RE CORDS. ··1980 ZR: Luxuridacna peregrina
Papaianopol, 1980 sp. nov. Romania. Ref. to
Papaianopol, p. 53 .
peregrinorsa Orbigny, 1843. - Changed combination:
Protocardia peregrinorsa (d'Orbigny, 1843).
Cardiidae. Cretaceous. RECORDS. ··1842 d'Orbigny, p.
81, lam . 18, figs. 6-8: Cardium peregrinorsa (jide
Arango & Gonzalez 1983). ··1857 Deshayes: Cardium
peregrinosum sic d'Orbigny close to Cardium hillanum
[fossil]. ··1917 ZR: Protocardia peregrinorsa [Ref.
Douville 1917] Cretaceous. ··1922-32 Sherborn, p.
4847 : Cardium peregrinorsum d 'Orbigny, 1842. ··1983
Arango & Gonzalez: Protocardia peregrinorsa
d'Orbigny, 1842. ··1983 Botero & Gonzalez: Protocardia peregrinorsa d'Orbigny, 1842. ··1986 Scott:
Protocardia peregrinorsa (d 'Orbigny, 1843).
peregrinorsa orientatis Douville, 1917 (variety). Upgraded. Protocardia peregrinorsa orientalis
Douville, 1917. Cardiidae. Cretaceous. RECORDS.
··1917 ZR: Protocardia peregrinorsa orien/alis race
n. Douville, 1917. Cretaceous Moghara.
peregrinorsum (mode). - See under peregrinorsa
Orbigny.
peregrinosum (error). - Printer's mistake for
peregrinorsa in Deshayes 1857.
peregrinum d'Orbigny. - Combination to be revised :
" Cardium" peregrinum. Cardiidae. Cretaceous.
RECORDS . ··1871 Stoliczka: Cardium peregrinum
(genus Protocardium). ··1876 Meek: Cardium
peregrinum d ' Orbigny.
peregrinum Eberzin, 1958. - Changed combination:
Arpadicardium peregrinum (Eberzin, ? ) 958) .
Lymnocardiidae. Pliocene. RECORDS. ··1953 Celidje:
Arpadicardium peregrinum Ebers. ··1958 ZR :
Limnocardium (Arpadicardium) peregrinum Eberzin,
1958 sp. nov. SUbgenotype. Pliocene of the Ponto-
692
Tropical Marine Mollusc Programme (TMMP)
peregrinum - perornatum
Caspian basin, Russia. ··1987 Ta ktakishvili :
Limnocardium peregrinum Ebers. ··1987 Taktakishvili:
Arpadicardium peregrinum.
peregrinum Jousseaume, 1888. [Figs. Part 3 p. 864). Synonym of Vasticardium pectiniforme (Born, 1780)
OTHER JUDGEMENTS: Trachycardiumflavum (Linnaeus,
1758). - Cardium dupuchense Reeve. » Regarding
Trachycardium peregrinum : Cardiidae Recent.
DISTRIBUTION. Indo-Pacific (Hylleberg 1994). REcORDs.
··1888 Jousseaume: Tra chy cardium p eregrinum .
··1888 ZR: Trachycardium peregrinum Jousseaume,
1888 n. sp. Red Sea. ··1893 Dautzenberg : Cardium
peregrinum Jousseaume. ··1905 Anthony: Cardium
peregrinum Jousseaume. ··1909 Lamy: Cardium
peregrinum. ··1919 Odhner: Cardium p eregrinum
Jousseaume (1888) = C. dupuchense Reeve. "1923
Dautzenberg: Cardium peregrinum. ··1927 Lamy :
Cardium peregrinum. ··1932 Prashad : Trachycardium
peregrinum (Jousseaume, 1888) = Trachycardium
flavum (Linne, 1758). ··1939 Adam & Leloup: Cardium
peregrinum Jousseaume = Laevicardium (Trachycardium )flavum (Linne 1758) ... 1977 Fischer-Piette,
p. 174: Cardium peregrinum Jousseaume = Cardium
flavum Linne. ··1992 Oliver: Trachycardium
peregrinum (Jousseaume, 1888) = Trachycardium
flavum (Linne, 1758). ··1997 Vidal: Trachycardium
peregrinum Jousseaume, 1888 = Vasticardium pectin iform e (Born, 1780). ··1999 Vidal: Trachycardium peregr'inum Jousseaume, 1888 = Vasticardium pectin iform e
(Born, 1780). ··1999 Higo et al.: Trachycardium peregrinum Jousseaume, 1888 = Acrosterigma (Regozara)
flava (Linnaeus, 1758).
peregrinum Papaianopol, 1982. - Changed combination:
Euxinicardium peregrinum. Lymnocardiidae. Pliocene.
TYPE LOCALITY. Ponti en et Dacien du Bassin Dacique,
Romania. RECORDS. ··1982 Papaianopol, p. 207, fig.:
LimfJocardium (Euxinicardium) peregrinum sp. nov.
··1987 Taktakishvili : Euxinicardium peregrinum
Papaian.
perelongata (variety). - See under deshayesi perelongata
Kutassy, 1929.
peres;; d 'Orbigny, 1850. - Combination to be revised:
"Cardium" peresii d'Orbigny, 1850. Cardiidae. Fossil.
RECORDs. ··1922-32 Sherborn, p. 4849: Cardium peresii
d'Orbigny, 1850.
perezi Bellardi, 1852. - Combination to be revised:
"Cardium" perezi Bellardi, 1852. Cardiidae. Fossil.
RECORDS. ··1980 Ruhoff: Cardium perezi Bellardi,
1852; vol. 4: 240, pI. H (sic = 19), figs. 2-5.
perezijorme Schauroth, 1865. - Combination to be
revised : "Cardium" pereziforme Schauroth, 1865.
Cardiidae. Fossil. RECORDS. "1980 Ruhoff: Cardium
pereziforme Schauroth, 1865; p. 209, pI. 18, fig. 9.
perfecta Chelidze, 1957. - Changed combination:
Pontalmyra perfecta (Chelidze, 1957). Lymnocardiidae. Pliocene . RECORDS. ··1957 ZR: Didacna
perfecta Chelidze, 1957 sp. nov. Pliocene, River
Zhuzhi, Makharadze town, Georgia, Russia. ··1987
Taktakishvili: Pontalmyra perfecta (Tschel.).
perflda (variety) . - See under ingratum perfida
Kolesnikov, 1929.
periimaris Maury, 1925 . - Trigoniocardia periimaris
(Maury, 1925). Cardiidae . Fossil. RECORDS. ··1977
Popov : Trigonio cardia (Americardia) periimaris
(Maury, 1925).
perincertum Gregorio , 1930. - Combination to be
revised : "Cardium " perincertum Gregorio, 1930.
Cardiidae. Lias . RECORDS. ··1936 ZR: Cardium
perincertum Gregorio, 1930 sp. n. Lias Palermo.
perlonga Kvaliashvili, 1978. - Digressodacna perlonga
Kvaliashvili , 1978. Lymnocardiidae. Pliocene.
RE CORDS. ··1978 ZR : Digressodacna perlonga
Kvaliashvili, 1978 sp. nov. Georgia SSR. Ref. to
Kvaliashvili p. 50.
permica. - Cypricardinia? Permica. Cardiacea. Upper
Permian. RECORDs. ··1931 ZR: Cypricardinia? permica
sp. n. Upper Permian Northern Province Russia.
permulta Celidje, 1953 . - Monodacna permulta Celidje,
1953. Lymnocardiidae. Tertiary. RECORDS ... 1953
Celidje, p. 157: Monodacna permulta nov. sp.
pernimia Kvaliashvili, 1978. - Digressodacna pernimia
Kvaliashvili, 1978. Lymnocardiidae. Pliocene.
RECORDS. ··1978 ZR: Digressodacna pernimia
Kvaliashvili , 1978 sp. nov. Georgia SSR. Ref. to
K valiashvili p. 76.
perobliquulII Koenen . - Fragum perobliquum Koenen.
Cardiidae. Cretaceous. RECORDS. ··1999 Jacques Vidal
(pers. com .) , MNHN , Paris: Fossils from the
Paleontological Collection. Fragum perobliquum
Koenen. Senonien. Cameroun, R51368 , R51369,
R51552.
perobliquum Martin. - Combination to be revised:
"Cardium " perobliquum Martin. Cardiidae. Miocene.
RECORDS. ··1919 Martin: Cardium perobliquum Mart.
perornata (mode). - See under perornatum Iredale, 1929.
perornatum Iredale, 1929. [Part 1. Fig. p. 144. Part 3 p.
903). - Synonym of Ctenocardia virgo (Reeve, 1845).
OTHER JUDGEMENTS: replaced by Fragum (Ctenocardia)
perornatum nom . nov. for Cardium imbricatum
Sowerby. » Regarding Ctenocardia perornatum :
Cardiidae. Recent. RECORDS. ··1929 Iredale, p. 264 (pI.
30, fig. 14): Fragum (Ctenocardia) perornatum nom .
nov. for Cardium imbricatum Sowerby,
1841(imbricatum preoccupied by Born, 1780). ··1929
ZR: Frag um perornatum nom. n. for Cardium
imbricatum Sow. [Ref. Iredale 1929b] . ··1977 FischerPiette : Corculum (Ct enocardia) p erornata Iredale.
··1977 Wilson & Stevenson : Fragum (Ctenocardium)
perornata Iredale, 1929; new name for Cardium
imbricatum Sowerby, 1841 . ··1977 Wilson &
Stevenson: Ctenocardia perornata (Iredale, 1929).
··1992 Lamprel.l & Whitehead: Ctenocardia perornata
Iredale, 1929 = Ctenocardia virgo Reeve, 1845 . ··1994
Chung-Hung & Hsi-Jen: Fragum perornatum Ireda1e.
··1995 Chung-Hung & Hsi-Jen: Fragum perornatum
Iredale. ··1999 Higo et af.: Ctenocardia p erornata
Iredale, 1929 = Ctenocardia virgo (Reeve, 1845).
··1999 Higo et af.: Fragum perornatum Iredale, 1929
sensu Bernard et al., 1993 = Ctenocardia virgo (Reeve,
1845). ··2003 Hylleberg & Kilburn , p. 177, PI. 2, fig.
2. Ctenocardia perornatum (Iredale , 1929) . CHARACTERS AND OCCURRENCE - DESCRIPTION.
··1977 Wilson & Stevenson: shell solid, inflated; umbo
central, prosogyrate; shell height slightly greater than
shell length. Lunule well defined, short and broad,
similar in left and right valves; escutcheon poorly
defined, elongate. Radial ribs strong, interstices wide
and deep, each rib with a central row of prominent
Phuket Marine Biological Center Special Publication vol. 30 (2004)
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perrieri - pesolinum
spines, crescent-shaped in section, concavity facing
ventrally; posterior slope with numerous smaller spines,
interstices anterior to umbonal keel smooth or crossed
by minute crowded transverse lamellae; interstices
behind keel crossed by stronger transverse lamellae
which thicken and extend up over posterior ribs as
closely packed V-shaped scales or ridges. Lateral,
delicate, compressed, pointed scales or spinelets
sometimes present on ribs. HABITAT. ··1977 Wilson &
Stevenson: mud, shelly sand with sponge, sand at 290 m depth. ··1994 Hylleberg: Indo-Pacific.
DISTRIBUTION. ··Of authors: Western Australia,
Queensland, Sulu Archipelago, Philippines. ··2003
Hylleberg & Kilburn: C. perornatum from Quinhon,
Vietnam is deposited in the ColI. Saurin at MNHN,
Paris (J. Vidal).
perrieri Dautzenberg & Fischer, 1897. - Synonym of
Parvicardium minimum Philippi, 1836. »Regarding
Cardium perrieri: Cardiidae. Recent. DISTRIBUTION.
Europe. RECORDS. ··1897 Dautzenberg & Fischer:
Cardium perrieri. ··1897 ZR: Cardium perrieri
Dautzenberg & Fischer, 1897 n. sp. N. Atlantic. ··1927
Dautzenberg: Cardium perrieri Dautzenberg & Fischer
= Cardium (Parvicardium) minimum Philippi. ··1977
Fischer-Piette: Cardium perrieri Dautzenberg &
Fischer = Cardium minimum Philippi. ··2001 Aartsen
& Goud: Cardium perrieri Dautzenberg & Fischer,
1897 = Parvicardium minimum (Philippi, 1836).
Persikima Iredale, 1937 (genus). - Tridacnidae. REcORDs.
··1937 ZR: Persikima Iredale, 1937 gen. n. Tridacnidae
type Persikima whitleyi Iredale, 1937 sp. n. Middleton
Reef, S. Pacific.
Persikima (subgenus). - REcORDs. ··1998. Schneider:
(Persikima) subgenus of Tridacna. ··1999 Higo et at.:
Persikima (subgenus) Iredale, 1937. Type species:
Persikima whitleyi Iredale, 1937 (0. d.).
persistans Schwetz, 1913. - Changed combination:
Moquicardium persistans (Schwetz, 1913).
Lymnocardiidae. Tertiary. RECORDS. ··1913 ZR:
Cardium persis tans Schwez, 1913 sp. n. Tertiary,
Kertsch. ··1987 Taktakishvili: Cardium persistans
Schwetz (= Prosodacna ampe/akiensis Andrus.).
··1987 Taktakishvili: Moquicardium persis tans.
personata Martyn, 1784. Non binominal. - Not
available. Not a cardiid. RECORDS. ··1784 Martyn:
Cardium personata.
perstriatum Kuroda, 1928. - Synonym of Acrosterigma
transcendens (Melvill & Standen, 1899). OTHER
JUDGEMENTS: Trachycardium "variegatum "(Sowerby).
» Regarding Trachycardium perstriatum: Cardiidae.
Fossil. TYPE MATERIAL. No data on types (Vidal 1999).
DISTRIBUTION. Indo-Pacific (Hylleberg 1994). RECORDS.
··1928 Kuroda, p. II, No. 117: Trachycardium
perstriatum Kuroda (nov.) Short description in
Japanese. ··1952 Kuroda & Habe: perstriatum Kuroda
= Trachycardium "variegatum (Sowerby). ··1977
Fischer-Piette: Cardium perstriatum Kuroda =
Laevicardium variegatum Sowerby. ··1981 Koyama et
at.: Cardium perstriatum Kuroda, 1928 = Cardium
(Trachycardium) transcendens Melvill & Standen,
1899. ··1999 Higo et at.: Cardium perstriatum Kuroda,
1928 = Acrosterigma (Vasticardium) transcendens
(Melvill & Standen, 1899). ··1999 Vidal: Kuroda 1928,
p.11 [Fide Habe, 1981: III]: Cardiumperstriatum=
Acrosterigma transcendens (Melvill & Standen, 1899).
*pertenue. - Homonymy to be noticed. Cardium
pertenue from 2 sources: Meek & Hayden, 1861. Gabb,1869.
pertenue Gabb, 1869. - Combination to be revised:
"Cardium" pertenue Gabb, 1870. Cardiidae. Tertiary.
TYPE LOCALITY. Payata, Peru. RECORDS. ··1870 Gabb p.
30: Cardium (Laevicardium) pertenue Gabb. ··1980
Ruhoff: Cardium (Laevicardium) pertenue Gabb, 1869;
vol. 5: 30.
pertenue Meek & Hayden, 1861. - Changed
combination: Protocardium perte/we. Cardiidae.
Cretaceous. TYPE LOCALITY. From North America.
RECORDS. ··1871 Stoliczka: Cardium pertenue
(Protocardium) (Referred to by Meek 1864). ··1876
Meek: Cardium pertenue Meek & Hayden type of (b)
Leptocardia Meek, 1876. ··1876 Meek: Protocardia
(Leptocardia) pertenuis Meek & Hayden, 1861. ··1970
Speden: Cardium pertenue Meek & Hayden, 1861, p.
442.
pertenuis (mode). - See under pertenue.
pertumidum Koenen, 1893. - Discors pertumidum
(Koenen, 1893). Cardiidae. Oligocene. RECORDS.
··1899 Sacco: Discors pertumidus (Koen.). ··1977
Popov: Discors pertumidum (Koenen, 1893).
Perucardia Olsson, 1944 (genus). - Cardiidae.
Maastrichtian. RECORDS. ··1980. Vokes: Peru cardia
Olsson, 1944 in subfamily Cardiinae. ··1998 Schneider
(1998b): Perucardia. Maastrichtian.
Peru cardia Olsson, 1944 (subgenus). - Cardiidae. Upper
Senonian. GENOTYPE. Olsson (1944) designated
Cardium brueggeni type species of subgenus
Peru cardia of genus Vepricardium. The shell of has
ribs with granular nodes. Peru. (Moore et al. 1969).
RECORDS. ··1944 ZR: Perucardia Olsson, 1944 subgen.
nov. of Cardium type Cardium (Peru cardia ) brueggeni
Olsson, 1944b sp. nov. Paita District Peninsula, Upper
Senonian. ··2002 Schneider: Peru cardia was found to
be a 'transitional fossil' between Profraginae and all
other eucardiids.
peruvianum Spieker, 1922. - Combination to be revised:
"Cardium" peruvianum (Spieker, 1922). Cardiidae.
Tertiary. RECORDS.
··1922 ZR:
Cardium
(Trachycardium) peruvianum Spieker, 1922 Tertiary,
Peru.
pervinquieri Freneix. - Protocardia pervinquieri
Freneix. Cardiidae. Cretaceous. RECORDS. ··1980 ZR:
Protocardia pervinquieri Freneix, 1980 nom. nov. for
Protocardia biseriata Pervinquiere 1912 nec Conrad
1852. Ref. to Freneix p. 172. ··1999 Jacques Vidal
(pers. com.) MNHN, Paris: Fossils from the
Paleontological Collection. Protocardia pervinquieri
Freneix. Santonien. Algeria, B 19932.
pesbovis d' Archiac, 1843- Combination to be revised:
"Cardium" pesbovis d' Archiac, 1843. Cardiidae.
Fossil. RECORDS. ··1857 Deshayes: Cardium pes bovis.
··1922-32 Sherborn, p. 4879: Cardium pesbovis
d' Archiac, 1843.
pesolinum Contejean, 1859. - Combination to be
revised: "Cardium" peso/inum Contejean. Cardiidae.
Lias-Jurassic. RECORDS. ··1980 Ruhoff: Cardium
pesolinum Contejean, 1859; p. 277, pI. 15, figs. 6-7.
··1980 Ruhoff: Cardium pesolinum Contejean, 1859;
quoted by Loriol & Cotteau 1867; vol. 21: 554, pI. 7,
694
Tropical Marine Mollusc Programme (TMMP)
pessoae - philippianum
fig. 13. "1999 Jacques Vidal (pers. com.), MNHN,
Paris: Fossils from the Paleontological Collection.
Cardium pesolinum Contejean. Portlandien. France
(Yonne), S05432, R50201.
pessoae Maury, 1924. - Changed combination:
Agnocardia pessoae (Maury, 1924). Cardiidae. Tertiary.
RECORDS. ··1984 ZR: Agnocardia pessoae (Maury,
1925) Comb. nov. Transferred from Cardium
(Trachycardium) pessoae Maury, 1925. Ref. to Vokes
p. 42. ··1998 Schneider: Agnocardia pessoae (Maury,
1924). ··2002 Schneider: Cardium (Trachycardium)
pessoae Maury, 1925. ··2002 Schneider: Agnocardia
pessoae (Maury, 1925). Cardiidae.
pestis Zhizhchenko, 1934. - Combination to be revised:
"Cardium" pestis Zhizhchenko, 1934. [Miocene,
Cardiidae], RECORDS. ··1939 ZR: Cardium pestis
Zhizhchenko, 1934 sp. n. Miocene Ciscaucasia. ··1952
ZR: Cardium pestis Miocene Caucasus [Ref.
Zhezhcenko 1934].
pesurae Mariani, 1919. - "Cardium" pesurae Mariani,
1919. Cardiidae. Rhaetic. RECORDS. ··1919 ZR:
Cardium pesurae Mariani, 1919 sp. n. Rhaetic
Lombardy.
*petersi. - Homonymy to be noticed. Cardium petersi
from 2 sources: Homes, 1862. - Zittel, 1865.
petersi Homes, 1862. - Synonym of Tauricardium
subsquamulosum.» Regarding Limnocardium petersi:
Lymnocardiidae. Miocene. RECORDS. ··1961 ZR:
Limnocardium (Tauricardium) petersi Pontian
Miocene, Kadar, Bosnia, Yugoslavia [Ref. Stevanovic
1961]. ··1980 Ruhoff: Cardium petersi Homes, 1862;
[1851-1870] vol. 4: 199, pI. 19, fig. 3. ··1987
Taktakishvili: Limnocardium petersi (M. Hoem.).
··1987 Taktakishvili: Tauricardium petersi =
subsquamulosum.
petersi Zittel, 1865. - Protocardium petersi, Zittel.
Cardiidae. Cretaceous. RECORDS. "1871 Stoliczka:
Protocardium petersi, Zittel. ··1980 Ruhoff: Cardium
(Protocardia) petersi Zittel, 1865 [1865-1866] vol.
24(2): 145, pI. 6, fig. 5.
petersi nasyrica (subspecies). - Tauricardium petersi
nasyrica. Lymnocardiidae. Fossil. RECORDS. ··1987
Taktakishvili: Tauricardium petersi nasyrica.
peters; oviformis (subspecies). - Tauricardium petersi
o viform is. Lymnocardiidae. Fossil. REcORDs. ··1987
Taktakishvili: Tauricardium petersi oviformis.
petilum Loriol, 1868. - Protocardium petilum (Loriol,
1868). Cardiidae. Cretaceous. TYPE LOCALITY. Valangien
beds, Suisse. RECORDS. ··1871 Stoliczka: Cardium
petilum Lorio!, 1868 (genus Protocardium). "1980
Ruhoff: Cardium petilum Loriol, 1868: ser. 4: 26, pI.
2, figs. 3-4.
petitianum Orbigny, 1853. - Synonym of Papyridea
semisulcata (Gray, 1825). OTHER JUDGEMENTS: close to
Cardium bullatum. » Regarding Cardium petitianum:
Cardiidae. Recent. RECORDS. ··1853 Orbigny, p. 309,
pI. 27, fig. 50-52: Cardium petitianum d'Orb., 1846
sic. ··1853 Orbigny: Cardium petitianum d'Orb., 1846
close to Cardium bullatum. ··1864 Krebs: Cardium
ringiculum Sowb. (syn.) = Cardium petitianum d'Orb.
··1884 Grasset: Cardium petitianum. Orb. = C.
bullatum. ··1885 Smith: Cardium petitianum d 'Orbigny
= Cardium (Papyridea) semisulcatum, Gray. 1825.
··1889 Dall: Papyridea petitiana Orbigny. ··1890
Paetel: Cardium petitianum Orb. = Papyridea bullata.
··1902 Dall: Cardium petianum Orbigny, 1845 =
Cardium semisulcatum Gray, 1825. ··1902 Dall &
Simpson: Cardium petitianum d'Orbigny, 1845 =
Cardium semisulcatum Gray 1825. ··1939 McLean:
Cardium petitianum Orbigny, 1853 = Papyridea
semisulcata (Gray, 1825). ··1951 McLean: Cardium
petianum d 'Orbigny (1846) = Papyridea semisulcata
(Gray). ··1958 Abbott: Papyridea petitianum Orbigny
1846 = Papyridea semisulcata Gray 1825. ··1970 Rios:
Cardium petitianum Orbigny = Papyridea semisulcata
(Grey, 1825). ··1974 Abbott: Cardium petitianum
Orbigny, 1846 (orange form) = Papyridea semisulcata
(Gray, 1825). - CHARACTERS AND OCCURRENCE - REMARKS. ··1958 Abbott: Papyridea
petitianum Orbigny 1846, is the orange form of
Papyridea semisulcata Gray 1825. DISTRIBUTION.
··1884 Grasset: Guadeloupe. ··Of authors: South
Florida, West Indies, Brazil.
petkovici Pavlovic, 1929. - New combination:
Lymnocardium petkovici (Pavlovic, 1929).
Lymnocardiidae. Ponti an. RECORDS. ··1928 ZR:
Limnocardium (Pseudocatillus) petkovici [Ref.
Pavlovitch 1928]. ··1929 ZR: Limnocardium
(Pseudocatillus) petkovici Pavlovic, 1929 sp. n. Ponti an
Serbia. ··1987 Taktakishvili: Limnocardium petkovici
Pavl.
pharaonica Abbass, 1966. - Lunulicardia pharaonica
Abbass, 1966. Probably not in Cardiidae. Cretaceous.
REcORDs. ··1966 ZR: Lunulicardia pharaonica Abbass,
1966 sp. nov. Cretaceous, Egypt.
pharaonum Cuvillier. - Cardium pharaonum Cuvillier.
Cardiidae. Lower Eocene. RECORDS. ··1999 Jacques
Vidal (pers. com.), MNHN, Paris: Fossils from the
Paleontological Collection. Cardium pharaonum
Cuvillier. Ypresien. Egypt (Thebes), B43626.
pharisiana (variety). - See tuberculatum pharisiana
Pallary, 1924.
pharnaci Schwez, 1913. - Pseudocatillus pharnaci
(Schwez, 1913). Lymnocardiidae. Tertiary. RECORDs.
··1913 ZR: Monodacna pharnaci Schwez, 1913 sp. n.
Tertiary, Kertsch. ··1967 Ebersin: Pseudocatillus
pharnaci (Schwets). ··1984 ZR: Pseudocatillus
pharnaci Schwetz. Referred to new subgenus P
(Donacicatillus). Ref. to Akhvlediani p. 71.
phasiaca Davidachvili, 1933. - Changed combination:
Didacnomya phasiaca Davidachvili, 1933.
Lymnocardiidae. Upper Pliocene. REcORDs. ··1934 ZR:
Didacnomya phasiaca Davidachvili, 1933 sp. n. Upper
Pliocene Georgia, Caucasus. ··1939 ZR: Didacnomya
phasiaca Bogatschev, 1938 "sp. n." Baku. ··1967
Ebersin: Didacnomya phasiaca (David.).
phataetisis Chiploukar, 1975. - Combination to be
revised: "Cardium" phataetisis Chiploukar, 1975.
Cardiidae. Upper Cretaceous. RECORDS. ··1975 ZR:
Cardium phataetisis Chiploukar, 1975 sp. nov.
Kashmir, Phata. Ref. to Chiploukar, p. 94.
philippianum Dunker, 1847. - Nemocardium
philippianum (Dunker, 1847). Cardiidae. Lias and
Jurassic. RECORDS. ··1922-32 Sherborn, p. 4909:
Cardium philippianum Dunker, 1847. ··1999 Jacques
Vidal (pers. com.), MNHN, Paris: Fossils from the
Paleontological Collection. Nemocardium philippianum Dunker. Heltangien. France (Vendee), R05122.
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695
philipp;; - Phyllocardillm
*philippii. - Homonymy to be noticed. Cardium phi/ippii from 2 sources: Deshayes, 1853. -Ihering, 1897.
philippii Ihering, 1897. - Hedecardium philippi
Cardiidae. Oligocene. ?Lower Miocene. RECORDS.
"1897 Ihering: Cardium philippi (fide Schneider 2002).
··2002 Schneider: Hedecardium (Iheringicardium)
philippi Cardiidae.
philipp;; Deshayes, 1853. - New name. Combination to
be revised: "Cardium" philippii Deshayes, 1853.
Cardiidae. Fossil. RECORDs. "1857 Deshayes: Cardium
philippii Desh. new name for Cardium pulchellum
Philippi, 1844. "1980 Ruhoff: Cardium philippii
Deshayes, 1853; vol. 2: 46.
*philippinense. - Homonymy to be noticed. Cardium
philippinense from 2 sources: Hedley, 1899. - Oeshayes
in Shirley, 1912.
philippinense Oeshayes in Shirley, 1912. Nomen nudum.
OTHER JUDGEMENTS: Vasticardium orbita philippinense
(Hedley, 1899). - Cardium angulatum Lamarck. Trachycardium orbita (Broderip & Sowerby, 1833). »
Regarding Cardium philippinense: Cardiidae. Recent.
TYPE LOCALITY. Philippines. TYPE REPOSITORY. Paris
Museum. RECORDS. "1899 Hedley: Cardium
philippinense Deshayes = Cardium angulatum
Lamarck. "1912 Shirley: Cardium philippinensis Desh.
"1934 Tomlin: Cardium (Trachycardium)
philippinense Deshayes. "1977 Fischer-Piette:
Laevicardium (Trachycardium) philippinense
(Deshayes) Shirley. "1980 Keen: Cardium
philippinense = Vasticardium orbita (Sowerby, 1833).
"1991 Voskuil & Onverwagt: Laevicardium
(Trachycardium) philippinense (Deshayes) Shirley,
1912 = Trachycardium orbita (Broderip & Sowerby,
1833). "1997 Vidal: Cardium philippinense Deshayes:
Hedley 1899 = Vasticardium orbita philippinense
(Hedley, 1899). "1997 Vidal: Laevicardium
(Trachycardium) philippinense (Deshayes) Shirley
sensu Fischer-Piette, 1977 (not Vasticardium orbita
philippinense (Hedley). - CHARACTERS AND
OCCURRENCE - REMARKS. "1899: Hedley was
unable to trace the species Cardium philippinense in
the literature. "1980 Keen: declared it nomen nudum
as of Shirley, 1912 and Tomlin, 1934; specimens
evidently labeled by reference to a lot in the British
Museum Collection that carried only Deshayes'
manuscript name. Cardium philippinensis Desh. is only
a name in a check list by Shirley 1912. DISTRIBUTION.
1994 Hylleberg: Indo-Pacific. Hawaii, Australia.
philippinense Hedley, 1899 [Figs. Part 3 p . 866]. Vasticardium philippinense (Hedley, 1899). OTHER
JUDGEMENTS: Vasticardium orbita (Broderip & Sowerby,
1833). RECORDS. "1999 Vidal: Cardium philippinense
Hedley, 1899 = Vasticardium orbita (Broderip &
Sowerby, 1833).
philippinense. - See also under orbita philippinense
Hedley, 1899.
phillipsi Koninck 1885. - Replaced by Conocardium
nexile Branson, 1942. Conocardiidae. Fossil. RECORDS.
'-1942 ZR : Conocardium phillipsi Koninck 1885 non
Orbigny 1850 replaced by Conocardium nexile
Branson, 1942 nom. nov.
phillipsi Orbigny 1850. - Conocardium phillipsi Orbigny
1850. Conocardiidae. Fossil. RECORDS. --1922-32
Sherborn, p. 4911: Conocardium phillipsii d'Orbigny,
1850. '-1942 ZR: Conocardium phillipsi Orbigny 1850
[Ref. Branson 1942].
Phlogocardia Stewart, 1930 (genus). - Cardiidae. Fossil
and Recent. Subgenus Phlogocardia upgraded to
generic level by authors. RECORDS. "1977 Popov p. 48:
Phlogocardia Stewart, 1930. Genotype: Cardium
belcheri Broderip & Sowerby, 1829 . .. 1980 Vokes:
Phlogocardia Stewart , 1930 in subfamily
Trachycardiinae.
Phlogocardia Stewart, 1930 (section, subgenus). Cardiidae. Fossil and Recent. Type species: Stewart
(1930) designated Cardium belcheri Broderip &
Sowerby, 1829 type species of subgenus Phlogocardia
of genus Trachycardium March, 1853. RECORDS. "1930
ZR: (Phlogocardia) Stewart, 1930 fossil sec. n. of
Trachycardium type Cardium belcheri Brod. and Sow.
[Ref. Stewart 1930]. "1961 Olsson: (Phlogocardia)
Stewart, 1930. "1999 Vidal : Subgenus (Phlogocardia)
Stewart, 1930 of the genus Trachycardium. CHARACTERS AND OCCURRENCE - CHARACTERS
of diagnostic value: central ribs with frills along
posterior sides, anterior ribs with knobby spines
(Hylleberg 1994).
phlyctaena Dall, 1900. - Combination to be revised:
"Cardium " phlyctaena Dal\, 1900. Cardiidae. Fossil.
RECORDS . "2002 Schneider: citing Dal1 (1900):
Cardium (Cerastoderma) phlyctaena sp. nov. section
Dinocardium.
pholadiforme Sowerby, 1847. - Combination to be
revised: "Cardium" pholadiforme Sowerby, 1847.
?Cardiidae. Fossil. RECORDS. "]922-32 Sherborn, p.
4922: Cardium pholadiforme Sowerby, 1847.
Pholadomya Agassiz, 1842 (genus). - Synonym of
Adacna Eichwald . » Regarding Pholadomya:
Lymnocardiidae RECORDS. "1858 Adams & Adams:
Syn. Pholadomya Agass. and Midd., not Sow. = Genus
Adacna Eichwald. "1890 Paetel: Pholadomya Ang.
syn. of Adacna. "1903 Andrussoff: Pholadomya
Agassiz, 1842.
Phylicardium (error). - Printer's mistake for
Phyllicardium in Pavlovic 1928 (Vokes 1980).
phyllicardiforme Jekelius, 1944. - Lymnocardium
phyllicardiforme Jekelius, 1944. Lymnocardiidae.
Miocene . RECORDS . --1946 ZR: Limnocardium
phyllicardiforme Jekelius, 1944 sp. nov. Miocene
Soceni Banat.
Phyllicardium (error). - Printer's mistake for
Phyllocardium in Andrussow 1903 (Vokes 1980).
Phyllocardiini (tribe) . - Cardiidae. MiocenelPliocene.
RECORDS. "1987 ZR: Phylocardiini Nevesskaja, 1986
Tribe nov. of Cardiidae. Type genus Phyllocardium
Fischer, 1887. Ref. to Nevesskaya et al. p. 189.
Phyllocardillm Fischer, 1887 (genus). - Lymnocardiidae.
Tertiary. RECORDS. -'1903 Andrussoff: Phyllicardium
Fischer, 1887. "1905 Dollfus, commenting Andrussow
1903: Phyllicardium Fischer. "1962 ZR:
Phyllocardium Fischer provisionally in subfam.
Didacninae Eberzin, 1962, family Cardiidae [Ref.
Eberzin 1962]. "1970 Glibert & Poel: Phyllicardium.
"1980 Vokes: Phyllocardium Fischer, 1887 in
subfamily Didacninae. "1987 ZR : Phyllocardium
Fischer, 1887. Designated type genus for new tribe
Phylocardiini. Ref. to Nevesskaya et al. p. 189.
Phyllocardium Fischer, 1887 (subgenus) . -
696
Tropical Marine Mollusc Programme (TMMP)
picardi - pinnulatum
Lymnocardiidae. Tertiary. GENOTYPE. Cardium planum
Deshayes (Dollfus 1905, commenting Andrussow
1903). RECORDS. ··1887 Fischer: Subgenus
Phyllocardium of genus Limnocardium, Stoliczka, em.
1870.
picardi Blanckenhorn, 1934. - Protocardia picardi
Blanckenhorn, 1934. Cardiidae. Cretaceous. RECORDS.
··1934 ZR: Pr%cardia picardi Blanckenhorn, 1934
sp. n. Cretaceous Syria.
picta (variety). - See under edule piela Milachewitch,
1916. - tuberculatum var. pic/a.
pictaviense d'Orbigny, 1850. - Combination to be
revised: "Cardium " pic/aviense d'Orbigny, 1850.
Cardiidae. Lias and Jurass ic . RECORDS . ··1922-32
Sherborn, p. 4949: Cardium piclaviense d ' Orbigny,
1850. ··1999 Jacques Vidal (pers. com.) MNHN, Paris:
Fossils from the Paleontological Collection. Cardium
pictaviense d ' Orbigny. Callovien. France (Sarthe),
S02743.
*picleti. - Homonymy to be noticed. Cardium picteti
from 2 sources: Rouault, 185 I. - d' Arch (?year).
pictet; d' Arch. - Combination to be revised: "Cardium"
picleli d' Arch. Cardiidae. Eocene. RECORDS. "1886
Cossmann: Cardium picteli d' Arch.
picteti Rouault, 185 I. - Combination to be revised:
"Cardium" picteli M. Rouault, 185 I. Cardiidae. Fossil.
REcORDs. ··1980 Ruhoff: Cardium picteti M. Rouault,
1851; vol. 8: 388.
pictum (subspecies). - See under edufe pictum.
pictum Dunker, 1861 . - Synonym of Cardium
papi/losum Polio OTHER JUDGEMENTS: unknown species.
» Regarding Cardium pictum: Cardiidae. Recent.
RECORDS. "1861 Dunker, p. 37: Cardium piclum (fide
Fischer-Piette 1977). "1872 Tryon: Cardium pictum
Dkr. ··190 I Dall: Cardium pictum Dunker, April, 1861
is probably a variety of Cardium papillosum Polio
··1944 Clench & Smith: Cardium pictum Rav. in
Dunker 1862 = unknown species. ··1980 Ruhoff:
Cardium pictum Dunker 1861; vol. 8: 37.
pictum Ravenel, 1861. - Unknown species. OTHER
JUDGEMENTS: Laevicardium pictum (Ravenel, 1861). Laevicardium serratum (Linnaeus, 1758). - Cardium
serratum var. sybariticum Dall, 1886. » Regarding
C. pictum: Cardiidae. Recent. RECORDS. "1861
Ravenel, vol. 13, p. 44: Liocardium pic/um . "1901
Dall: Cardium pictum Ravenel, February, 1861. ··1902
Dall: ?Cardium pic/um Ravenel (type lost) = Cardium
serra tum var. sybari/icum Dall, 1886. ··1902 Dall :
Cardium pic/um Ravenel, 1861 (not of Dunker, 1861)
= Cardium serra/um Linnaeus, 1758. "1921 Maury :
pic tum Ravenel, 1861 = Cardium (Laevicardium)
serratum Linnaeus, 1758. "1925 Woodring, p. 145:
Cardium pictum Ravenel, 1862 is identical with
Cardium serratum Linne, 1758 . ··1939 McLean :
Cardium pictum Ravenel , 1861 = Laevicardium
serralum (serra/um) (Linne, 1758). "1944 ZR :
Laevicardium pictum Ravenel [Ref. Clench & Smith
1944]. ··1962 Warmke & Abbott: Laevicardium pictum
Ravenel, 1861. ··1968 Jong & Kristensen: Americardia
pictum Ravenel. ··1970 Rios: Laevicardium pictum
(Ravenel, 1861). ··1974 Abbott: Laevicardium pictum
(Ravenel, 1861). ··1977 Popov: Laevicardium pic tum
(Ravenel). ··1977 Fischer-Piette: Laevicardium pic/um
Ravenel. ··1980 Ruhoff: Liocardium pic tum Ravenel,
1861; p. 44. ··1997 Shelton: Laevicardium pictum
(Ravenel, 1861). - CHARACTERS AND OCCURRENCE - REMARKS. ··1977: Fischer-Piette, p. 22 failed
to locate references given by Dunker (1862 (1861), VII,
p. 37), including references in Tryon and Dall. The
species is insufficiently known. DISTRIBUTION. ··1939
McLean: North Carolina to Brazil.
pictus Tryon, 1870. - Hemidonax pictus (Tryon, 1870).
Cardiidae, ?Hemidonacidae. Recent. RECORDS. ··1950
Allan: Hemidonax pictus (Anatomical structure places
Hemidonax in the family Cardiidae). ··2001 Schneider
& Carter: Hemidonax pic/us (Tryon, 1870).
Hemidonacidae.
piettei J.e. Fischer. - Pr%cardia piettei J.e. Fischer.
Cardiidae. Lias and Jurassic. RECORDS. ··1999 Jacques
Vidal (pers. com.), MNHN , Paris: Fossils from the
Paleontological Collection. Protocardia piette; J.e.
Fischer. Bathonien. France (Aisne, Ardennes), R05125,
B47036, B47038 .
pigmae Shu to, 1960. - Laevicardium pigmae Shuto,
1960. Cardiidae. Middle Miocene. RECORDS. ··1960
ZR: Laevicardium pigmae Shu to, 1960 sp. nov. Tano
member, Miyazaki Group, Middle Miocene, Japan.
··1973 Keen: P. (sic = Laevicardium) pigmae Shuto,
1960. Middle Miocene.
pi/atum Stoliczka, 1871. - Combination to be revised:
"Cardium" pitatum (Stoliczka, 1871). Cardiidae.
Cretaceous. TYPE LOCALITY. Ninnyoor, South India.
RECORDS. "1871 Stoliczka, p. 218, pI. 13, figs. 6-7:
Cardium (Cerastoderma) pitatum Stoliczka. "1980
Ruhoff: Cardium (Cerastoderma) pita tum Stoliczka,
1870; [1870-1871] vol. 3: 218, pI. 13, figs. 6-7.
pi/ide; Tournouer, mss. Fischer, 1886. - Prosodacna
pilidei Fischer, 1886. Lyrnnocardiidae. Pliocene. TYPE
LOCALITY. Valea N igovani, Roumania. RECORDS. ··1886
Fischer: Prosodacna pi/idei Tournouer, mss.
Description p. 223, PI. 9, fig. 7. ··1999 Jacques Vidal
(pers. com.), MNHN, Paris: Fossils from the
Paleontological Collection. Prosodacna pilidei
Tournouer. Rumania, B35815.
pinellasense Mansfield, 1937. - Combination to be
revised: "Cardium" pinellasense (Mansfield, 1937).
Cardiidae. Lower Miocene. RECORDS. "1937 ZR:
Cardium (Dinocardium?) pinellasense Mansfield, 1937
sp. n. Tampa limestone Florida (Lower Miocene).
Pingecardium (error) . - Printer's mistake for
Ringicardium in Clark & Woodford, 1927 (Stewart
1930; Vokes 1980).
pinnulatum Conrad, 1831. [Figs . Part 1 p. 145, 178].Changed combination: Parvicardium pinnulatum
(Conrad, 1831). Cardiidae. Pliocene, Pleistocene and
Recent. TYPE LOCALITY . Unknown (Reeve 1845);
Massachusetts (McLean 1939) . RECORDS. "1831
Conrad, p. 260 (pI. II , fig. 8): Cardium pinnulatum .
"1845 Reeve: Cardium pinnulatum Conrad. "1860
Stimpson: Cardium pinnulatum Con. ··1873 Tryon:
Cardium pinnulatum Conrad, 1836. ··1884 Grasset :
Cardium pinnulatum Conf. "1889 Dall : Cardium
pinnulatum Conrad [Po PI iocene] . "1890 Paetel :
Cardium pinnulatum Conf. ··1902 Dall : Cardium
(Cerastoderma) pinnulatum Conrad, 1831
[Pleistocene]. "1922-32 Sherborn, p. 4985: Cardium
pinnulatum Conrad, 1831. ··1934 Johnson: Cardium
pinnulatum Conrad 1831 . ··1939 McLean : Cardium
Phuket Marine Biological Center Special Publication vol. 30 (2004)
697
Pinzonela - Plagiocardium
(Cerastoderma) pinnulatum Conrad. "1939 ZR:
Cardium (Cerastoderma) pinnulatum fig. [Ref.
McLean 1939]. "1944 ZR: Cerastoderma pinnulatum
Conrad figs. [Ref. Clench & Smith 1944a]. "1953 La
Rocque: Cerastoderma pinnulatum Conrad 1831.
"1962 Dexter: Cerastoderma pinnulatum(Conrad).
"1971 ZR: Cerastoderma pinnulata Crawling. Veliger
larva. Photomicrography [Ref. Culliney 1971].001974
Abbott: Cerastoderma pinnulatum (Conrad, 1831).
001977 Fischer-Piette: Cardium (Cerastoderma)
pinnulatum Conrad. 001980 Keen: Parvicardium
pinnulatum (Conrad, 1831). 002000 Aartsen & Goud:
Parvicardium pinnulatum (Conrad, 1831).002001
Vidal, p. 408: Parvicardium pinnulatum. CHARACTERS AND OCCURRENCE - DESCRIPTIONS. 001845 Reeve: shell ovate, rather oblique,
radiately ribbed, 26 ribs, irregularly sculptured with
numerous short obtuse scales, posterior ribs the more
conspicuous; whitish. 00200 I Vidal: only the exhalant
aperture is tube-shaped. Inhalant siphon open towards
the foot aperture. No eyes. DIMENSIONS. 001939 McLean:
12 x 10 x 6.5 mm; ribs 25-28. REMARKS. 001845 Reeve:
although some doubt has been attached to the value of
this species, the shell here represented, is certainly
distinct from any of the small Cardium that I have had
an opportunity of examining. DISTRIBUTION. 001845
Reeve: unknown. 001884 Grasset: Portland. ooOf
authors: Labrador to North Carolina.
Pinzonela Cowper-Reid, 1943 (genus). Cardiacea. Trias.
RECORDS. 001947 ZR: Pinzonela [Pinzonella in Index]
Cowper-Reid, 1943 nov. gen. Trias, Brazil [Cardiacea]
Genotype Pinzonela illusa Cowper-Reid, 1943 sp. nov.
pironai.- Apricardia pironai. Cardiacea. RECORDS.
001933 ZR: Apricardia pironai [Ref. Parona 1933].
pirsagatica Andrusov, 1963. - Changed combination:
Pontalmyra pirsagatica (Andrusov, 1963).
Lymnocardiidae. Miocene. RECORDS. 001963 ZR:
Didacna pirsagatica Andrusov, 1963 sp. nov. Miocene,
Russia. 001987 Taktakishvili: Pontalmyra pirsagatica
(Andrus.).
pisiformis Gmelin. - Synonym of Cardium discors
Gmel. » Regarding Tellina pisiformis: Cardiidae.
Recent. RECORDS. 001803 Montagu: Tellina pisiformis
Gmel. Syst. p. 3241? = Cardium discors Gmel.
pisolithicum d'Orbigny, 1850 - Combination to be
revised: "Cardium" pisolithicum d'Orbigny, 1850.
Cardiidae. Fossil. RECORDS. 001871 Stoliczka: Cardium
pisolithicum (? Acanthocardium sic). 001922-32
Sherborn, p. 4994: Cardium pisolithicum d'Orbigny,
1850.
pissaroi (error). - Printer's mistake for pissarroi in ZR
for 1908 with ref. to Cossmann 1908.
pissarroi Cossmann, 1905. - Changed combination:
Fragum pissarroi (Cossmann, 1905). Cardiidae. Fossil.
RECORDS. 001908 ZR: Cardium pissaroi sic Cossmann,
1908b sp. n. Loire inferieure. 001970 Glibert & Poel:
Fragum pissarroi (Cossmann, 1905).
pium Zhizhchenko, 1934. - Cerastoderma pium (Zhizhchenko, 1934). Cardiidae. Miocene. RECORDS. 001939
ZR: Cardium pium Zhizhchenko, 1934 sp. n. Miocene
Ciscaucasia. 001952 ZR: Cardiumpium Miocene Caucasus [Ref. Zhezhcenko 1934]. 001977 ZR:
Cerastoderma (Inaequicostata) pium Key to subspecies. Ref to Il'ina, Nevesskaya & Paramo nova p. 77.
pium convexum Paramonova, 1977 (subspecies). Cerastoderma pium con vexum Paramonova, 1977.
Cardiidae. Upper Miocene. RECORDS. 001977 ZR:
Cerastoderma (Inaequicostata) pium convexum
Paramonova, subsp. nov. Russian SFSR. Ref to Il'ina,
Nevesskaya & Paramonova. p. 207, fig'd.
placerense Gabb, 1864. Combination to be revised:
"Cardium" placerense. Gabb, 1864. Cardiidae.
Cretaceous. TYPE LOCALITY. From California. RECORDS.
001871 Stoliczka: Cardium (Protocardium) placerense.
(Recorded by Gabb). 001980 Ruhoff: Cardium
(Protocardium) placerensis Gabb, 1864; vol. I: 173,
pI. 24, fig. 156.
placida Stefanescu, 1896. - Changed combination:
Pontalmyra placida (Stefanescu, 1896). Lymnocardiidae. Miocene. Type species of subgenus
Pontalmyra Stefanescu, 1896 in family Limnocardiidae
(Glibert & Poe I 1970). Type species of subgenus
Pontalmyra Sabba [= Stefanescu], 1896 (Sacco, 1899).
RECORDS. 001896 ZR: Pontalmyra placida Miocene,
Roumania [Ref. Stefanescu]. 001970 Glibert & Poel:
Pontalmyra placida Stefanescu, 1896. 001987
Taktakishvili: Didacna placida (Sabba). 001987
Taktakishvili: Pontalmyra placida.
plaekovalevskii Goretzki, 1956. Caspia plaekovalevskii
Goretzki, 1956. Lymnocardiidae. Quaternary. RECORDS.
001956 ZR: Caspia plaekovalevskii sic Goretzki, 1956
sp. nov. Lower Kamar basin, Quatemllry Russia.
Plagiadacna (error). - Printer's mistake for Plagiodacna
in Dollfus 1905 (Vokes 1980).
Plagiocardium Cossmann, 1886 (genus). - Cardiidae.
Tertiary and Recent. Type species: Crosse (1887)
designated Cardium granulosum Lamarck, 1805 type
species of genus Plagiocardium Cossmann, 1886.
RECORDS. 001930 Stewart: Plagiocardium Cossmann
1886 (p. 254-255). 001977 Kafanov and Popov:
Plagiocardium subfamily Fraginae, tribe Parvicardiini
(jide Schneider 2002). 001977 Popov: Plagiocardium
Cossmann, 1886. Genotype: Cardium granulosum
Lamarck, 1805. 001980 Vokes: Plagiocardium
Cossmann 1886 in subfamily Cardiinae. 001983 Anaya:
Plagiocardium Cossmann, 1886. Synonym of
Parvicardium Monterosato, 1884. 001995 Pacaud &
Renard: Plagiocardium Cossmann, 1886 in subfamily
Cardiinae. 001998 Schneider (1998b): Plagiocardium
in Fraginae. The genus appears in the Paleocene (Keen
1980). - CHARACTERS AND OCCURRENCE DESCRIPTIONS. 001969 Moore et al.: the shell of
Plagiocardium is elliptical-oblique, hinge arched; rib
sculpture of bead-like nodes. Europe-Africa-East
Indies-New Zealand. 002002 Schneider: the right
anterior cardinal tooth is subconical in shape, above
(or sometimes slightly anterior) to the right anterior
cardinal socket. The tooth does not obscure the socket
face. The right anterior cardinal tooth has a rounded
tip. The right posterior cardinal is lacking a hook.
Plagiocardium has symmetrical or nearly symmetrical
knob-like spines. Plagiocardium was found to have
composite spines.
.
Plagiocardium Cossmann, 1886 (subgenus). Cardiidae. Tertiary and Recent. RECORDS. 001886
Cossmann: Section Plagiocardium nov. sect. of genus
Cardium Linne, 1758. 001887 Fischer: Plagiocardium
Cossmann, 1887. Section of genus Cardium Linne,
698
Tropical Marine Mollusc Programme (TMMP)
Plagiodacna - plancarinata
1758. - CHARACTERS AND OCCURRENCE DESCRIPTIONS. ··1969 Moore et at.: Shell medium in
size; cardinal teeth nearly equidistant between lateral
teeth. Europe-East Indies. REMARKS. The designation
"section" takes the value of subgenus.
Plagiodacna Andrussow, 1903 (genus). Lyrnnocardiidae.
Fossil. GENOTYPE . Cardium carinatum Deshayes
(Dollfus 1905, commenting Andrussow 1903).
RECORDS . ··1905 Dollfus, commenting Andrussow
1903: Plagiodacna n. g. ··1962 ZR: Plagiodacna
Andrussov provisionally in subfam. Didacninae
Eberzin, 1962, family Cardiidae [Ref. Eberzin 1962].
··1977 Popov: Plagiodacna Andrussow, 1903. ··1977
ZR: Plagiodacna Pliocene. Romania, systematic
accounts. Ref. to Papaianopol, I. ··1980 Vokes:
Plagiodacna Andrussow, 1903 in subfamily
Didacninae. ··1987 Taktakishvili, p. 93: Plagiodacna
Andrussow, 1903; referred to new tribe Pontalmyrini.
··1987 ZR: Plagiodacna Andrussov, 1903. Referred
to new tribe Arcicardiini. Ref. to Nevesskaya et at. p.
190. ··200 I Schneider & Carter: Plagiodacna (genus).
Lymnocardiidae. Pliocene.
Plagiodacnopsis Andrussow, 1923 (genus). Lymnocardiidae. Tertiary and Recent. RECORDS. ··1980
Vokes: Plagiodacnopsis Andrussow, 1923 in subfamily
Adacninae.
Plagiodacnopsis Andrusov, 1963 (section and subgenus).
Lymnocardiidae. Pliocene. RECORDS. ··1963 ZR:
(Plagiodacnopsis) Andrusov, 1963 sect. nov. of
Monodacna Apsheron beds, Pliocene, Russia. ··1963
ZR: Plagiodacnopsis Andrusov, 1963 sect. nov. of
Monodacna q.v.
Plagiodaena (error). - Printer's mistake for Plagiodacna
in Salisbury & Edwards, 1960 (Vokes 1980).
*plagiomorphum.- Homonymy to be noticed. Cardium
plagiomorphum from 3 sources: Cossmann, 1886. Bois-Gouet (?year). - Laub (?year).
plagiomorphum Bois-Gouet. - Changed combination:
Parvicardium plagiomorphum (Bois-Gouet).
Cardiidae. Fossil. RECORDS. ··1970 Glibert & Poel:
Cardium plagiomorphum Bois-Gouet. ··1970 Glibert
& Poe!: Plagiocardiumplagiomorphum (Bois-Gouet).
··1970 Glibert & Poel: Parvicardium plagiomorphum
(Bois-Gouet).
plagiomorphum Cossmann , 1886. - Changed
combination: Parvicardium plagiomorphum (Cossmann, 1886). Cardiidae. Eocene. TYPE LOCALITY. Paris
Basin, France. RECORDS. ··1886 Cossmann, p. 170, PI.
VIII, fig. 21-22: Cardium plagiomorphum nov. sp.
··1899 Sacco: Papillicardium plagiomorphum
(Cossmann). ··1970 Glibert & Poel: Parvicardium
plagiomorphum (Cossmann, 1886). ··1995 Le Renard
et al.: Parvicardium plagiomorphum [Cardium
plagiomorphum] Cossmann, 1886. ··1995 Pacaud &
Renard: Parvicardium plagiomorphum.
plagiomorphum Laub. - Changed combination:
Plagiocardium plagiomorphum (de Laub). REMARKS. I
have only seen the below reference to de Laub.
RECORDS. ··1904 Cossmann & Pissarro: Cardium
(Plagiocardium) plagiomorphum de Laub.
*plana. - Homonymy to be noticed. Didacna plana
from 2 sources: Ebersin, 1959. - Nevesskaya, 1956.
plana Ebersin, 1959. - Didacna plana Ebersin, 1959.
Lyrnnocardiidae. Fossil. RECORDS. ··1959 ZR: Didacna
plana Ebersin, 1959 sp. nov. fossil.
plana Nevesskaya, 1956. - Didacna plana Nevesskaya,
1956. Lymnocardiidae. Quaternary. RECORDS. ··1956
ZR: Didacna plana Nevesskaya, 1956 sp. nov. Khazar
deposits, Quaternary. Cheleken, Russia, Turkmenia .
"1957 ZR: Didacnaplana Neveskaya, 1957 sp. nov.
Khazar deposits, Chelekhan Quaternary, Turkmenia,
Russia.
Planacardium Paramonova, 1971 (genus). Lymnocardiidae. Miocene. RECORDS. ··1980. Vokes:
Planacardium Paramonova, 1971. Subfamily
uncertain. ··1977 Popov: Planacardium Paramonova,
1971. ··1977 ZR: Cerastaderma (sic = Cerastoderma)
(Planacardium). Key to species. Ref to I1'ina, Nevesskaya & Paramonova. p. 78. ··200 I Schneider & Carter:
Planacardium (genus). Lymnocardiidae. Miocene.
planacardo Andrussow, 1907. - Arcicardium planacardo
Andrussow, 1907. Lymnocardi idae. F ossi I. TYPE
LOCALITY. Crimean iron ore deposits. RECORDS. ··1907
Andrussow, p. 114, Taf. IV, Fig. 10-12: Arcicardium
planacardo nov. sp. ··1964 ZR: Arcicardium
planacardo Andrusov, 1964 sp. nov. Tertiary.
planaecarinata. Nomen nudum. RECORDS. ··1949 ZR:
Replidacna planaecarinata 'no sp.' [nomen nud.]
Maestricktian, Sarmat. [Ref. Papp 1949].
planata (variety). - See under irgisicum planata var.
Yarkin, 1963.
planatum Brocchi. [Figs. Part 1 p. 94]. - Synonym of
Parvicardium papillosum (Poli, 1791). Cardiidae.
Quaternary and Recent. RECORDS. ··1886 Dollfus &
Dautzenberg: C. planatum Bf. = Cardium (s. g.?)
papillosum Poli. ··1869 Romer: Cardium planatum
Brocchi = Cardium papi/losum Polio ··1878
Monterosato: C. plana/um (Ren.) Broce. = C. Polii
Payr. = C. papillosum Polio ··1922-32 Sherborn, p.
5005: Cardium planatum Brocchi, 1814 (ex Renier,
1804). ··1999 Jacques Vidal (pers. com.), MNHN,
Paris: Fossils from the Paleontological Collection.
Cardium planatum Brocchi. Sicily, B40617, B40618.
planatum Renier, 1804. Nomen nudum. Not valid. OTHER
JUDGEMENTS: Parvicardium papi/losum (Poli, 1791). »
Regarding Cardium planatum: Cardiidae. Recent.
DISTRIBUTION. Europe, North West Africa. RECORDS.
··1804 Renier: Cardium plana/um. ··1836 Philippi:
Cardium plana/um Ren. Broc. = Cardium papi/losum
Poli o ··1836 Scacchi: Cardium planatum Ren. Broc.
··1862 Weinkauff: Cardium plana tum Ren. = Cardium
papil/osum Poli. ··1867 Weinkauff: Cardium planatum
Renieri = Cardium papil/osum Polio ··1886 Locard:
Cardium planatum Renieri, 1804 = Cardium
papillosum Poli, 1791. ··1888 Kobelt: Cardium
planatum Renier = Cardium papillosum Poli. ··1888
Servain: Cardium plana tum of Renieri = Cardium
papillosum Poli, 1791. ··1890 Paetel: Cardium
planatum Ren. = papillosum Polio ··1902 Dollfus &
Dautzenberg: Cardium planatum Renier, 1804 =
Cardium (Parvicardium) papillosum Poli, 1791 .
··1922-32 Sherborn, p. 5005: Cardium planatum
Renier, 1804 [n.n.]. ··1977 Fischer-Piette: Cardium
planatum Renier = Plagiocardium papillosum Poli.
plancarinata Papp, 1954. - Replidacna plancarinata
Papp, 1954. Lymnocardiidae. Sarmatian. RECORDS.
··1954 ZR: Cardium? (Replidacna) plancarinata Papp,
1954 sp. nov. Sarmatian, Vienna.
Phuket Marine Biological Center Special Publication vol. 30 (2004)
699
Planicardium - planicostatum
Planicardium Olsson, 1967 (genus). - Cardiidae. Fossil.
RECORDS . "1980 Vokes: Olsson, 1967. - CHARACTERS AND OCCURRENCE - DESCRIP-TIONS. "2002
Schneider: Clinocardium (Planicardium): Large shells
externally resembling Dinocardium, but with a hinge
resembling Cerastoderma (Campbell 1993).The right
anterior cardinal teeth are small, blunt, longer than high,
anterior and dorsal to the right anterior cardinal socket,
and do not obscure the face of the socket. The right
posterior cardinal tooth is twisted, curved, and lacking
a hook. REMARKS. "2002 Schneider: Olsson (1967)
erected Planicardium as a monotypic genus. Only
species was the Neogene Cardium virginianum Conrad,
1839. Keen (1980) classified Planicardium as a
subgenus of Clinocardium because the "beaks [are]
prosogyrate, [with] the initial part or tip lying above
anterior cardinal teeth." Planicardium, often considered
closely related to Clinocardium, is a cardiine.
planicostata Dockery, 1982. - Trachycardium plan icos/ata Dockery, 1982. Cardiidae. Lower Oligocene.
RECORDS . '·1982 ZR: Trachycardium planicostata
Dockery, 1982 sp. nov. Mississippi. Ref. to Dockery,
p.70.
planicostata Stevanovic, 1950. - Parvidacna
planicostata Stevanovic, 1950. Lymnocardiidae. Upper
Pontian. RECORDS. "1951 ZR: Parvidacna planicostata
Stevanovic, 1950 sp. nov. Upper Pontian Serbia,
genotype of Parvidacna Stevanovic, 1950. Cardiidae.
"1953 ZR: Parvidacna planicostata Stevanovic, 1951
sp. nov. genotype of Parvidacna Stevanovic, 1951 gen.
nov. Pontian Serbia.
planicosta (variety) . - See under paucicostatum
planicosta Monterosato.
planicostata kiptschakensis Andrusov, 1964 (variety).
- Infrasubspecific . Didacna planicostata var.
kiptschakensis Andrusov, 1964. Lymnocardiidae.
Miocene. RECORDS. "1964 ZR : Didacna planicostata
var. kiptschakensis Andrusov, 1964 nov. Miocene
USSR.
planicostata latecarinata Andrusov, 1964 (variety). Infrasubspecific. Didacna planicostata var.
latecarinata Andrusov, 1964 . Lymnocardiidae.
Miocene. RECORDS. "1964 ZR : Didacna planicostata
var. latecarinata Andrusov, 1964 nov. Miocene USSR.
planicostata pluricostata Andrusov, 1964 (variety). Infrasubspecific. Didacna planicostata var.
pluricostata Andrusov, 1964 . Lymnocardiidae.
Miocene. RECORDS. "1964 ZR: Didacna planicostata
var. pluricostata Andrusov, 1964 nov. Miocene USSR.
planicostata tenuicosUlta Andrusov, 1964 (variety). Infrasubspecific. Didacna planicostata var.
tenuicostata Andrusov, 1964. Lymnocardiidae.
Miocene. RECORDS. "1964 ZR: Didacna planicostata
var. tenuicostata Andrusov, 1964 nov. Miocene USSR.
planicostata verneuili Deshayes, 1838. - Pontalmyra
planicostata verneuili (Deshayes, 1838). Lymnocardiidae. Fossil. RECORDS. "1987 Taktakishvili:
Pontalmyra planicostata verneuili.
planicostata zuzica Chelidze, 1957 (variety). Upgraded. Didacna planicostata zuzica Chelidze, 1957.
Lymnocardiidae. Pliocene . RECORDS . "1957 ZR:
Didacna planicostata var. zuzica Chelidze, 1957 nov.
Pliocene, River Zhuzhi, Makharadze town, Georgia,
Russia.
planicostatum . - Homonymy to be noticed. Cardium
planicostatum from 4 sources: Sedgwick & Murchison,
1829. - Sowerby, 1833. - Deshayes, 1838. - Braunn
in Munster, 1842.
planicostatum Braun in Munster, 1842. - Replaced by
"Cardium " braunni Deshayes. Combination to be
revised. Cardiidae. Fossil. RECORDS. "1857 Deshayes:
Cardium planicostatum Braunn, 1842 - C. braunni
Desh . "1922-32 Sherborn, p. 5009: Cardium
planicostatum Braun in Muenster, 1842.
planicostatum Deshayes, 1838. - Pontalmyra
planicostata (Deshayes, 1838). Lymnocardiidae. Fossil.
TYPE LOCALITY. The Crimea. RECORDS. ··1838 Deshayes,
p. 53, pI. 2, figs . 7-8 : Cardium planicostatum. '·1857
Deshayes: Cardium planicostatum Deshayes, 1837 sic
- C. huoti Desh. ··1886 Fischer: Cardium planicosta tum Deshayes . ··1905 Dollfus, commenting
Andrussow 1903 : Cardium planicostatum Deshayes is
type species of sub-form Pontalmyra. ··1922-32
Sherborn, p. 5009: Cardium planicostatum Deshayes,
1838. ··1957 ZR: Didacna planicostata Pliocene [Ref.
Chelidze 1957]. ··1964 ZR: Didacna planicostata
Miocene [Ref. Andrusov 1964]. ·'1987 Taktakishvili:
Didacna planicostata (Desh.). "1987 Taktakishvili:
Pontalmyra planicostata (Desh.).
planicostatum Sedgwick & Murchison, 1829. Ctenocardia planicostatum (Sedgwick & Murchison,
1829). Cardiidae. Fossil. RECORDS. ··1829 Sedgwick
& Murchison: Cardium planicostatum. ··1857
Deshayes: Cardium plenicostatum sic Sedgw. et Murch.
1831 sic. "1947 Hertlein & Strong: Cardium
planicostatum.
planicostatumSowerby, 1833. [Figs. Part I p. 110,129.
Part 3 p. 904). - Americardia planicostata (Sowerby,
1833). OTHER JUDGEMENTS : Fragum magnijicum
(Carpenter, 1857). - Americardia guanacastensis
(Hertlein & Strong, 1947). » Regarding Cardium
planicostatum : Cardiidae. Recent. TYPE REPOSITORY.
Holotype. BMNH London. TYPE LOCALITY. Ad oras
Americae Central is (Guacomayo) in fine sand, at a
depth of 13 fathoms (Sowerby 1833). RECORDS. ··1833
Sowerby, p. 83: Cardium planicostatum. "1834
Sowerby, 50 th part, fig. 25 : Cardium planicostatum.
"1840 Sowerby, p. 7, No. 83 : Cardium planicostatum
Sow. "1840 Sowerby, corrected list, p. 8: Cardium
planicostatum (SowerbyNo. 83). ··1841 Reeve, pI. 77,
fig. 25: Cardium planicostatum Sow. ··1843 Hanley:
Cardium planicostatum Sow. ··1844 Reeve: Cardium
planicostatum Sowerby. ·'1852 Adams: 432. Cardium
planicostatum Sowb. 1833. "1858 Adams & Adams,
p. 459: Hemicardia (Fragum) planicostata Sow. "1885
Smith: Cardium planicostatum (on the west side of
America). '·1889 Dall p. 282 : Cardium planicostatum
Sowerby 1833 not Sedgwick & Murchison, 1829 = C.
magnificum Deshayes. "1890 Paetel: Hemicardium
planicostatum Sow. '·1891 Stearns: Cardium
(Hemicardium) planicostatum Sby. "190 I Dal1 :
Cardium planicostatum Sowerby, 1833 (not of
Sedgwick & Murchison, 1829) = Cardium (Fragum)
magnificum (Deshayes MS) in Carpenter, 1857. '·]92232 Sherborn, p. 5009: Cardium planicostatum Sowerby,
1833. "1947 Hertlein & Strong: Cardium planicostatum Sowerby, 1833 = Cardium (Americardia)
guanacastensis. "1947 ZR: Cardium planicostatum
700
Tropical Marine Mollusc Programme (TMMP)
planior - plenicostatum
Sow. P.Z.S. 1833 p. 83 not Sedgwick & Murchison
1829 p. 140, replaced by Cardium (Americardia)
guanacastense Hertlein & Strong, 1947 [nom. nov.].
··1961 Olsson: Cardium planicostatum Sowerby, 1833
=
Trigoniocardia (Americardia) guanacastense
(Hertlein & Strong, 1947). "1971 Keen: Cardium
planicostatum Sowerby, 1833 = Trigoniocardia
(Americardia) guanacastensis (Hertlein and Strong,
1947) . .. 1974 Abbott: Cardium planicostatum
Sowerby, 1833, not Sedgwick and Murchison, 1829 =
Americardia guanacastensis (Hertlein & Strong, 1947).
··1977 Fischer-Piette: Cardium planicostatum
Sowerby = Corculum guanacastensis Hertlein &
Strong. - CHARACTERS AND OCCURRENCE DESCRIPTIONS. ··1844 Reeve: shell somewhat squarely
cordate, attenuated towards the umbones, rounded
anteriorly, very short, concavely angulated posteriorly,
middle of the posterior area prominent, radiately ribbed,
30 flat, sharp-edged ribs, the anterior crenated in a
somewhat wrinkled manner, posterior finely serrated
along the posterior edge , the whole very closely
sculptured with arched impressed striae; interstices
between the ribs deeply excavated; warm pinkish fleshcolour, clouded and spotted with brown; interior white .
.. 1891 Steams: Cardium planicostatum Sby. is very
close to C. biangulatum Sby. Since Sowerby's specific
name had been used earlier by Sedgwick & Murchison,
1829, a new name was required for Cardium
planicostatum of Sowerby. Hence, Hertlein & Strong
(1947) proposed the name Cardium guanacastense.
REMARKS. "1844 Reeve: one of the most remarkable
features in this species is the extremely delicate manner
in which the flattened surface of the ribs is very
minutely engraved throughout with close-set arched
impressed striae. The specimen here figured, from the
collection of Mr. Cuming, and one in the British
Museum, are all that have as yet been discovered.
HABITAT. ··1844 Reeve: found in fine sand at the depth
of 13 fathoms . DISTRIBUTION. ··1844 Reeve: Guacomayo, Central America . ··1947 Hertlein & Strong:
California, Costa Rica.
planior (variety). - See under edule planior Stimpson
and vulgaris planior Krestovnikov, 1929.
planiovatum Tscheltzov, 1965. - Avicardium
planiovatum Tscheltzov, 1965. Cardiacea. Pliocene.
RECORDS. ··1965 ZR : Avicardium planiovatum
Tscheltzov, 1965 sp. nov. Pliocene, Turkmenia, USSR.
planissima (variety). - See under parvula var.
planissima.
planum Deshayes, 1838. - Changed combination:
Phyllocardium planum planum (Deshayes, 1838).
Lymnocardiidae . Tertiary. Type species. Cardium
planum Deshayes, type species of genus Phyllocardium
Fischer, 1887 (Dollfus 1905, commenting Andrussow
1903). Cardium planum Deshayes, 1838 (mono type)
type species of genus Phyllocardium sic Fischer, 1887
in family Limnocardiidae (Glibert & Poe I 1970). TYPE
LOCALITY. The Crimea. RECORDS ... 1838 Deshayes, p.
46, pI. 2, figs. 24-30: Cardium planum . --1857
Deshayes : Cardium planum Verneuil sic . ··1887
Fischer: Phyllocardium planum Deshayes. ··1907
Andrussow: Phyllicardium planum Desh. ··1922-32
Sherborn, p. 5017: Cardium planum Deshayes, 1838.
'·1953 Celidje: Phyllocardium planum Desh. ··1970
Glibert & Poel: Phyllocardium planum (Deshayes,
1838). ··1977 ZR: Phyllocardium planum planum
Pliocene. Romania, morphological Variation. Ref. to
Papaianopol & Popescu. ··1988 Lyutzkanov & Ziatev
Phyllocardium planum (Desh.).
planum giganteum Wenz, 1942 (subspecies) . Phyllicardium planum giganteum Wenz, 1942.
Lymnocardiidae. Pliocene. RECORDS. "1945 ZR:
Phyllicardium (Phyllocardium) planum giganteum
Wenz, 1942 subsp. nov. Pliocene Valea Salcii.
planum pentagonalis Huica , 1977 (subspecies). Phyllocardium planum pentagonalis Huica, 1977.
Lymnocardiidae . Pliocene . RECORDS. ··1977 ZR:
Phy llocardium planum pentagonalis Huica , 1977
subsp. nov. Romania, Pliocene. Ref. to Huica, I. p. 109,
fig'd .
planum rumanum Wenz , 1942 (subspecies) . Phy llicardium planum rumanum Wenz, 1942.
Lymnocardiidae . Pliocene . RECORDS. "1945 ZR:
Phyllicardium (Phyllocardium) planum rumanum
Wenz, 1942 subsp. nov.
platense Orbigny, 1842. - Changed combination:
Dinocardium platense (d'Orbigny 1842). Cardiidae.
Tertiary. RECORDS. ··1922-32 Sherborn, p. 5021:
Cardium pia tense d'Orbigny, 1842. "1930 Stewart:
Cardium platense d 'Orbigny. ··1994 Rio: Dinocardium
platense (d'Orbigny 1842).
platipleurum Brusina. - New combination:
Lymnocardium platipleurum (Brusina). Lymnocardiidae. Fossil. RECORDS. ··1987 Taktakishvili:
Limnocardium platipleurum Brus.
platonis Coquand, 1862 . - Unknown species. »
Regarding Cardium platonis : Cretaceous. TYPE
LOCALITY . Constantine. RECORDS. "1871 Stoliczka:
Cardium platonis Coquand, 1862 (species insufficiently
known). ··1980 Ruhoff: Cardium platonis Coquand,
1862; vol. 2: 205, pI. 10, figs. 11-12.
plebeium (spelling). - See under plebejum Solander,
1786.
plebejum Solander. - Not valid . OTHER JUDGEMENTS :
Cardium oblongum. Cardiidae . Recent. RECORDS.
··1817 Dillwyn : Cardium plebejum Solander's MSS =
Cardium oblongum Chemnitz [not available]. ··192232 Sherborn, p. 5033 : Cardium plebeium Solander,
1786 In. n.].
plectopleura Gardner, 1926. - Changed combination:
Trachycardium plectopleura (Gardner, 1926).
Cardiidae. Tertiary. RECORDS. ··1926 ZR: Cardium
(Trachycardium) plectopleura Gardner, 1926 sp. n.
Tertiary U.S.A. ··1970 Glibert & Poel: Trachycardium
(s.s.) plectopleura (Gardner, 1926).
pleistopleura Davidasvili 1932 Didacna pleistopleura
Davidasvili 1932. Lymnocardiidae. Fossil. RECORDS.
··1932 ZR: Didacna pleistopleura [Ref. Davidasvili
1932] ... 1964 Davidaschvili & Kitovani : Didacna
pleistopleura Dav. type species of genus Submonodacna Livental, 1931. ··1977 Popov: Didacna
(Pontalmyra) pleistopleura .
pleistopleura pselldomultistriata Dav (subspecies) .
Submonodacna pleistopleura pseudomultistriata Dav.
Lymnocardiidae. Fossil. RECORDS. ··1964 Davidaschvili
&
Kitovani: Submonoda cna pleistopleura
pseudomultistriata Dav.
plenicostatum (error). - Printer's mistake for
Phuket Marine Biological Center Special Publication vol. 30 (2004)
701
plenum - plicata
planicostatum in Deshayes 1857.
plenum Papaianopol, 1992. - Dacicardium plenum
Papaianopol, 1992. Lymnocardiidae. Pliocene. TYPE
LOCALITY. Dacien inferieur (Getien) du Bassin Dacique,
Romania. REcORDs. "1992 Papaianopol, p. 25, fig.:
Dacicardium plenum sp. nov.
pleonexia Davidachvili, 1933 - Changed combination:
Pseudocatillus pleonexia (Davidachvili, 1933).
Lymnocardiidae. Upper Pliocene. RECORDS. "1934 ZR:
Monodacna pleonexia Davidachvili, 1933 sp. n. Upper
Pliocene Georgia, Caucasus. "1939 ZR: Monodacna
pleonexia [? Bogatschev, 1938a] "sp. n." Baku. "1964
Davidaschvili & Kitovani: Monodacna pleonexia Dav.
"1967 Ebersin: Pseudocatillus pleonexia (David.).
plesiochora Dav.- Didacna plesiochora Dav.
Lymnocardiidae. Fossil. RECORDS. "1964 Davidaschvili
& Kitovani: Didacna plesiochora Dav. "1964
Davidaschvili & Kitovani: Submonodacna plesiochora
Dav.
Pleuriocardia Scott, 1978 (genus). - Cardiidae. Fossil.
GENOTYPE. Cardium (?) kansasense Meek, 1871. OD.
RECORDS. "1978 Scott: Pleuriocardia Scott, 1978 n.
gen. (Cretaceous). "1978 ZR: Pleuriocardia Scott,
1978 gen. nov. of Cardiidae. Type species Cardium
kansasense Meek, 1871. Lower Cretaceous. Ref. to
Scott p. 893. "1978 ZR: Pleuriocardia (Pleuriocardia)
Diagnosis. Ref. to Scott p. 894. "1980 Vokes:
Pleuriocardia Scott, 1978. Fossil. ··1986 AbdelGawad: genus Pleuriocardia Scott, 1978. "1995
Schneider: Pleuriocardia (Pleuriocardia) Scott, 1978
Cretaceous. '·1995 Schneider: Pleuriocardia
(Dochmocardia) Scott, 1978 Cretaceous. "1998
Schneider: Pleuriocardia. Type species Pleuriocardia
kansasense (Meek, 1871).
Pleuriocardia Scott, 1978 (subgenus). - Cardiidae.
Fossil. RECORDS. "1978 Scott p. 894, pI. 1, figs. 1-13:
(Pleuriocardia) Scott, 1978 n. subgen. (Cretaceous).
"1986 Abdel-Gawad: subgenus (Pleuriocardia) Scott,
1978.
Pleuriocardiinae (subfamily). - Cardiidae. RECORDS.
"1995 Schneider: Pleuriocardiinae Schneider, 1995
subfam. n. Type genus: Pleuriocardia Scott, 1978.
"2002 Schneider: Incacardium has been transferred
to Pleuriocardiinae. "200 I Schneider & Carter:
Pleuriocardiinae (subfamily) Schneider, 1995. CHARACTERS AND OCCURRENCE - DESCRIPTIONS. "2000 Schneider: Cardiidae with sculpture of
radial ribs on entirety of shell, cross-striae in rib
interspaces. Unlike most cardiids, the right cardinal
teeth are of almost equal size: anterior right cardinal
tooth large, wide, blunt, arching over anterior cardinal
socket. Posterior right cardinal tooth shaped as in
Nemocardium and Granocardium, but rotated
counterclockwise so that the tooth and the right
posterior cardinal socket are at less than a 35° angle
from the horizontal. Spines of concave-down triangles
emanating from top of ribs (may be microscopic).
OCCURRENCE. "2000 Schneider: Albian to
Maastrichtian, United States, Mexico, Jamaica, Peru,
France, England, Algeria, Libya, Egypt.
Pleurorhynchus Phillips, 1836 genus). - Synonym of
Conocardium Bronn, 1835. »
Regarding
Pleurorhynchus. Conocardiidae: Fossil. RECORDS.
"1847 Gray, No. 561: Pleurorhynchus, sp. Swains.
1840. Synonym of Cardium, sp. Lam.; type species
Card. avicularia. "1857 Deshayes: Pleurorhynchus
Phillips, in Swainson, Morris: syn. of Cardium. "1887
Fischer: Pleurorhynchus Phillips, 1836. Jun. syn. of
Conocardium Bronn, 1835.
plicata (mode). - See also under plicatum Eichwald,
1829 & 1840.
plicata Eichwald, 1829. [Figs. Part 1 p. 122, 181]. Hypanis plicata (Eichwald, 1829). Lymnocardiidae.
Miocene and Recent. TYPE LOCALITY. Caspian Sea.
RECORDS. "1829 Eichwald: Glycimeris plicata. ··1838
Eichwald: Adacna plicata m. "1838 Eichwald:
Hypanis plicata Pando Menetr. "1838 Eichwald:
Glycymeris plicata Zoo I. special. "1841 Eichwald:
Adacna plicata. "1849 Middendorff: Pholadomya
plicata Eichw. (fide Andrussoff 1903). "1858 Adams
& Adams, p. 460: Adacna plicata, Eichw. "1866 Issei:
Adacna plicata, Eichwald, 1842. "1877 Grimm:
Adacna plicata Eichwald. "1877 ZR: Adacna plicata
(Eichw.) Caspian Sea [Ref. Grimm 1877]. "1890
Paetel: Adacna plicata Eichw. "1890 Woodward:
Adacna plicata Eich. - Caspian, Black Sea. "1884
Grasset: Adacna plicata. Eich. "1903 Andrussoff:
Glycimeris plicata EichwaJd, 1829. "1903 Andrussoff:
Eichwald, 1838 Glycimeris plicata (moved to genus
Adacna). "1917 ZR: Adacna plicata [Ref. Andrusov
1914]. .. 1922-32 Sherborn, p. 5045: Glycimeris plicata
Eichwald, 1829, Fauna Caspio-Caucasia, p. 224. "1935
ZR: Limnocardium plicatum Eichw. [Ref. Friedberg
1934]. "1935 ZR: Limnocardium plicatum Miocene
Poland [Ref. Friedberg 1934]. "1968 Logvinenko &
Starobogatov: Hypanis plicata plicata (Eichwald).
"1968 Logvinenko & Starobogatov: Glycimeris plicata
Eichwald, 1829 = Hypanis plicata plicata Eichwald.
"1968 Logvinenko & Starobogatov: Mya edentula
Pallas, 1771 = Hypanis plicata plicata Eichwald.
"1969 Nordsieck: Hypanis plicatus (Eichwald, 1829).
"1977 Popov,: A. plicata (Eichw.). "1977 Popov:
Hypanis plicatus. "1977 Popov: Plicatiforma plicatum
[fossil] = ?Eichwald's species. "1992 Schneiderp. 155:
Hypanis plicatum (Eichwald). Type species of Hypanis.
- CHARACTERS AND OCCURRENCE - DISTRIBUTION. "1884 Grasset: Caspian Sea. "1969 Nordsieck:
Black Sea.
plicata Eichwald, 1829. - See also under plicata Pander
in Menetries, 1832.
plicata Eichwald, 1838. - Hypanis plicata Eichwald.
Cardiacea. Fossil. RECORDS. "1857 Deshayes: Cardium
plica tum Eichwald, 1840. "1882. Menetries designated
Cardium plicatum Eichwald, 1829 sic as type species
of the genus Hypanis Eichwald, 1838 [Remarks:
Eichwald did not describe a species called Cardium
plicatum in 1829]. "1888 Kobelt: Cardium plicatum
Eichw. (Adacna). "1893. Ostroumoff: Cardium
plicatum (Eichw.). "1922-32 Sherborn, p. 5045:
Adacne sic (Pand.) plicata Eichwald, 1838 ... 1922-32
Sherborn, p. 5045: Hypanis ? plicata Eichwald, 1838.
"1987. Taktakishvili: Cardium plica tum Eichw. "1999
Jacques Vidal (pers. com.), MNHN, Paris: Fossils from
the Paleontological Collection as Cardium plica tum
Eichwald. Austria, B31397.
plicata Eichwald, 1841. - Adacna plicata Eichwald.
Cardiacea. Fossil. RECORDS. "1922-32 Sherborn, p.
5045: Adacna plicata Eichwald, 1841 N. Mem. Soc.
702
Tropical Marine Mol/usc Programme (TMMP)
plicata - plicatum
Imp. Nat. Moscou, VII, p. 280. - Glycimeris plicata
Eichwald, 1829, Fauna Caspio-Caucasia, p. 224.
plicata Grimm. - Changed combination: Adacna plicata
Grimm non Eichwald , 1877. Replaced by Adacna
(Hypanis) grimmi Andrusov, 1963. Lymnocardiidae.
Pliocene. RECORDS. ··1963 ZR: Adacna plicata Grimm
non Eichwald, 1877 replaced by Adacna (Hypanis)
grimmi Andrusov, 1963 Pliocene , Russia [Ref.
Andrusov 1963].
plicata Pander in Menetries, 1832. - Changed
combination: Adacna plicata (Pander in Menetries,
1832). Lymnocardiidae . Fossil. RECORDS. ··1837
Krynicki: Hypanis plicata Pander. Menetries =
Glicimeris plicata Eichw. Zoo I. spec. I. 279. V. 2.
However, Krynicki doubts the proposed synonymy.
··1903 Andrussoff: Hypanis plicata Pando in Menetries
1832 = Amph idesm a caspia Krynicki, 1837. ··1905
Dollfus, commenting Andrussow 1903: Adacna plicata
type species of group Hypanis Pander.
plicata golbargae Tadjalli-Pour, 1977 (subspecies). Hypanis plicata golbargae Tadjalli-Pour, 1977 .
Lymnocardiidae. Recent. RECORDS. ··1977 ZR: Hypanis
plica/a golbarg ae Tadjalli-Pour, 1977 subsp . nov.
Caspian Sea. Ref to Tadjalli-Pour, M. p. 99, fig ' d.
plicata luciae Borcea, 1926 (form). - Infrasubspecific.
No evidence of subspecific status. » Regarding
Adacna plicata plicata forma luciae: Changed
combination: Hypanis luciae (Borcea , 1926) .
Lymnocardiidae. Recent. DISTRIBUTION. Caspian Sea.
RECORDS. ··1962 Grossu: Adacna plicata plicata forma
luciae Borcea.
plicata minima - Adacna plicata minima.
Lymnocardiidae. Recent. RECORDS. ··1963 ZR: Adacna
plicata minima, food and feeding mechanism [Ref.
Romanova 1963].
plicata relicta Milaschewitch, 1916 (variety). - Changed
combination: Hypanis plicata relicta (Milaschewitch,
1916). Lymnocardiidae. Recent. RECORDS. ··1962
Grossu: Adacna plicata var. relicta Mil. ··1963 ZR:
Adacna plicata reliCla from Quaternary deposits of the
Danube delta [Ref. Grossu & Baltac 1963] . ··1969
Nordsieck: Hypanis plica/us relictus (Milaskewicz).
··Scarlato & Starobogatov: Hypanis plicata relicta
(Milaschewitch , 1916). - CHARACTERS AND
OCCURRENCE - REMARKS. The name of the author
is spelled in different ways: Milaskewicz,
Milaschewitch, and Milasevic. DISTRIBUTION. ··1969
Nordsieck: Black Sea.
plicata relicta dolosmiana Borcea, 1926 (form). Hypanis dolosmiana Borcea, 1926. » Regarding
Adacna plicata relicta dolosmiana : Lymnocardiidae.
Fossil. RECORDS. ··1962 Grossu: Adacna plicata relicta
forma dolosmiana Borc. ··Scarlato & Starobogatov:
Hypanis dolosmiana Borcea, 1926.
plicataeformis Gorj.-Kramb. - New combination:
Lymno cardium plicataeformis (Gorj .-Kramb.).
Lymnocardiidae. Fossil. RECORDS. ··1987 Taktakishvili:
Limnocardium plicataeformis (Gorj.-Kramb.).
plicatella (variety). - See under plicatum plicatella
Friedberg, 1934.
plicatieostatum Tscheltzov, 1965. - Combination to be
revised: "Cardium " plicatieostatum sic Tscheltzov,
1965 . Cardiacea. Pliocene. RECORDS. ··1965 ZR:
Cardium plicatieostatum sic Tscheltzov, 1965 sp. nov.
Akchagylian (Pliocene), Turkmenia, USSR.
Plicatiforma Paramanova, 1971 (genus). - Synonym of
Plicatlformes Kolesnikov, 1948. Lymnocardiidae.
Tertiary. RECORDS. ··1977 ZR: Cerastoderma
(Plicatiforma). Key to species. Ref to II'ina,
Nevesskaya & Paramonova p . 70 . ··1980 Vokes:
Plicatiforma Paramanova, 1971 pro Plicatiformes
Kolesnikov, 1948. ··200 I Schneider & Carter:
Plicatiforma (genus). Lymnocardiidae. Miocene.
Plicatiforma Tournouer, 1882 (genus). - RECORDS.
··1977 Popov: Plicatiforma Tournouer, 1882.
Plicatiformes Kolesnikov, 1948 (genus) . Lymnocardiidae. Tertiary and Recent. RECORDS. ··1980
Vokes: Plicatiformes Kolesnikov, 1948.
plicatofittoni - Combination to be revised: "Cardium"
plicatofittoni. Cardiacea. Sarmatian. RECORDS. ··1965
ZR: Cardiumplicatofittoni figd. Sarmatian, Yugoslavia
[Ref. Spajic 1965].
plicatoforme Senes, 1955. - Combination to be revised:
"Cardium"
plicatoform e
Senes,
1955 .
Lymnocardiidae. Miocene. RECORDS. ··1955 ZR:
Cardium (Limnocardium) plicatoforme Senes, 1955 sp.
nov. Miocene, Czechoslovakia.
plicatoides chisinau Mateuti-Soroca . - Upgraded.
Combination to be revised : "Cardium" plicatoides
chisinau Mateuti-Soroca. Cardiacea . Sarmatian.
RECORDS. ··1946 ZR: Cardium plicatoides MateutiSoroca n. f. chisinau, Sarmatian fossil, Rumania.
plicatum Ius (mode). - See also under plicata Eichwald,
1829 & 1840.
*plicatum. - Homonymy to be noticed. Cardium
plica tum from 2 sources: Eichwald, 1830. - Munster
in Goldfuss, 1837.
plicatum Eichwald , 1830. - RECORDS. ··1922-32
Sherborn, p. 5046: Cardium plicatum Eichwald, 1830.
Nat. Lithauen, 209.
plicatum Munster in Goldfuss, 1837. - Replaced by
"Cardium " munsteri Deshayes. Combination to be
revised » Regarding "c. " plicatum : Cardiidae. Fossil.
RECORDS. ··1857 Deshayes: Cardium plica tum Munster,
1840 - C. munsteri Desh. ··1922-32 Sherborn, p. 5046:
Cardium plicatum Muenster in Goldfuss, 1837, p. 215 .
plicatum jammense (subspecies). - Plicatiforma
plicatumjammense. Lymnocardiidae. Fossil. RECORDS.
··1977 Popov: Plicatiforma plica tum jammense.
plicatum plicatella Friedberg, 1934 (variety). - New
combination: Lymnocardium plicatum plicatella
(Friedberg, 1934). Lymnocardiidae. Miocene. RECORDS.
··1935 ZR : Limnocardium plicatum var. plicatella
Friedberg, 1934 nov. Miocene Poland.
plicatum pseudoplicatum Friedberg, 1934 (variety) . New combination: Lymnocardium plica tum pseudoplicatum (Friedberg, 1934). Lymnocardiidae. Miocene.
RECORDS . ··1935 ZR: Limnocardium plica/um var.
pseudoplicatum Friedberg, 1934 nov. Miocene Poland.
plicatulII rybnicense Grishkevitch & Nevesskaya, 1959
(subspecies). - Nomen nudum. »Regarding Cardium
plica/um rybnicense: Cardiacea. Miocene. RECORDS.
··1959 ZR: Cardium plicatum rybnicense Grishkevitch
& Nevesskaya, 1959 subsp. nov. as nom. nud. in list p.
90, Northern shore of Kara-Bogaz, Turkmeniya,
Sarmatian, Miocene.
plicatum . - Cerastoderma plicatum. Cardiidae. ? fossil.
RECORDS. ··1977 ZR: Cerastoclerma sic (Plicatiforma)
703
Phuket Marine Biological Center Special Publication vol. 30 (2004)
plinthinatus - politum
plicatum. Key to subspecies. Ref to Il'ina, Nevesskaya
& Paramonova p. 70.
plinthinatus Branson, 1942. - New name: Conocardium
plinthinatus Branson, 1942. Conocardiidae. Fossil.
RECORDS. ··1942 ZR: Conocardium plinthinatus
Branson, 1942 nom. nov. for Conocardium ornatum
Cleland 1911 non Winchell & Marcy 1865. "1943 ZR:
Conocardium plinthinatum Branson, 1942 nom. nov.
for C. ornatum Cleland, 1911 not Winchell & Marcy.
plumsteadianum (error). - Printer's mistake for
plumstedianum in Deshayes 1857, Le Renard et al.
1995, and Pacaud & Renard 1995.
plumstedianum Sowerby, 1813. - Changed combination:
Nemocardium plumstedianum (Sowerby, 1813).
Cardiidae. Oligocene. RECORDS. ··1835 Lamarck:
Cardium plumstedianum Sow. = Cardium semigranulosum Sow. ··1857 Deshayes: Cardium
plumsteadianum sic Sowerby. ··1922-32 Sherborn, p.
5061: Cardium plumstedianum Sowerby, 1813. ··1970
Glibert & Poel: Nemocardium (s.s.) plumstedianum
(Sowerby, 1813). "1995 Le Renard et al.:
Nemocardium (s. str.) plumsteadianum [Cardium
plumsteadianum 1Sowerby 1., 1813. ··1995 Pacaud &
Renard: Nemocardium (Nemocardium) plumsteadianum. "1999 Jacques Vidal (pers. com.), MNHN,
Paris: Fossils from the Paleontological Collection.
Cardium plumstedianum Sowerby. Germany, B24909.
pluricostata - Hyrcania pluricostata. Lymnocardiidae.
Fossil. RECORDS. "1977 Popov: Hyrcania (Hyrcania)
pluricostata.
pluricostata (variety). - See under caucasica pluricostata
Androussov, 1923. - planicostata pluricostata
Andrusov, 1964.
plurintermedia Andrussov, 1924. - Didacna plurintermediaAndrussov, 1924. Lymnocardiidae. Pliocene
and Pleistocene. RECORDS. ··1924 ZR: Didacna
plurintermedia Andrussov, 1924 sp. n. Pleistocene.
··1963 ZR: Didacna plurintermedia Andrusov, 1963
sp. nov. Apsheron layers, Pliocene, Russia.
pluriradiatum Chen et al. - Tulongocardium
pluriradiatum (Chen, Chen & Zhang, 1976). Cardiacea.
Upper Triassic. RECORDS. ··1980 ZR: Cardium
(Tulongocardium) pluriradiatum Chen, Chen & Zhang,
1976 sp. nov. China, Upper Triassic. Type species of
subgenus (Tulongocardium). Ref. to Wen et al., p. 32.
··1995 Schneider: Cardium (Tulongocardium)
pluriradiatum Chen, Chen & Zhang in Lan & Chen,
1976. Type species of Tulongocardium Chen, Chen &
Zhang in Lan and Chen, 1976.
plurisuperjactata (variety). - See under lamarck;;
plurisuperjactata Coen, 1937.
podolicum Deshayes, 1853. - Combination to be revised:
"Cardium" podolicum Deshayes. Cardiacea. Fossil.
RECORDS. ··1857 Deshayes: Cardium podolicum Desh.
"1980 Ruhoff: Cardium podolicum Deshayes, 1853;
vol. 2: 49.
pokveschia Akhvlediani, 1966. - Plagiodacna
pokveschia Akhvlediani, 1966. Lymnocardiidae.
Middle Pliocene. RECORDS. "1966 ZR: Plagiodacna
pokveschia Akhvlediani, 1966 sp. nov. Mid-Pliocene,
River Galizqa, Kerchensk Peninsula. SSR.
poleanum Schauroth, 1865. - Combination to be revised:
"Cardium" poleanum Schauroth, 1865. Cardiidae.
Fossil. RECORDS. ··1980 Ruhoff: Cardium poleanum
Schauroth, 1865; p. 209, pI. 18, fig 10
polemonis Schwets, 1913. - Changed combination:
Pseudocatillus polemonis (Schwets, 1913).
Lymnocardiidae. Fossil. RECORDS. "1913 ZR:
Monodacna polemonis Schwez, 1913 sp. n. Tertiary,
Kertsch. ··1967 Ebersin: Pseudocatillus polemonis
(Schwets). "1984 ZR: Pseudocatillus polemonis
Schwetz. Referred to new subgenus P (Donacicatillus).
Ref. to Akhvlediani p. 71.
polii Payraudeau, 1826. [Figs. Part I p. 143. Part 3 p.
933]. - Synonym of Parvicardium papillosum (Poli,
1791). OTHER JUDGEMENTS: Cardium planatum Renier.
- Cardium scabrum Phil. » Regarding Cardium poW:
Cardiidae. Recent. DISTRIBUTION. Mediterranean (Reeve
1845). RECORDS. "1826 Payraudeau, N. 96: Cardium
polii Nob. (new name for C. papil/osum Poli;
papillosum preoccupied). "1832 Bronn: ? Cardium
polii Pa. 57 = Cardium planatum Brch. ··1836 Philippi:
Cardium poW Payr. p. 57. = Cardium papillosum Polio
··1845 Reeve: Cardium polii Payraudeau = Cardium
papillosum Poli. ··1848 Requien: Cardium polii. Payr.
= Cardium scabrum. Phil. tom. 2. p. 38. tab. 14. fig.
16. "1850 Jay: Cardium poW Payr. = Cardium
papillosum Poli. "1857 Deshayes: Cardium poW
Peyraudeau sic. ··1862 Weinkauff: Cardium poW, Payr.
= Cardium papillosum, Poli. ··1867 Hidalgo: Cardium
poW, Payraudeau = Cardium papillosum, Polio ··1867
Weinkauff: Cardium poW Payraudeau = Cardium
papillosum Polio ··1886 Locard: Cardium poW,
Payraudeau, 1826 = Cardium papillosum, Poli, 1791.
··1888 Kobelt: Cardium poW Payraudeau = Cardium
papillosum Poli. "1888 Servain: Cardium poW
Payraudeau = Cardium papillosum Poli, 1791. "1890
Hidalgo: Cardium poli Payraudeau = Cardium
papillosum Polio ··1890 Paetel: Cardium poW Payr. =
papil/osum Poli. ··1900 Locard & Caziot: Cardium
poW = Cardium papillosum Poli, 1791. ··1902 Dollfus
& Dautzenberg: Cardium po Iii Payraudeau, 1826 =
Cardium (Parvicardium) papillosum Poli, 1791.
··1922-32 Sherborn, p. 5074: Cardium polii
Payraudeau, 1826. ··1977 Fischer-Piette: Cardium
poW Payraudeau = Cardium papil/osum Polio
polii Payraudeau (variety). - Identical with Parvicardium
papillosum (Po Ii, 1791). Cardiidae. Recent. RECORDS.
··1878 Monterosato: Cardium var. poW Payr. C.
obliquatum Arad. = Cardium papillosum Polio
politioanei Jekelius, 1944. - Cerastoderma politioanei
(Jekelius, 1944). Cardiidae. Miocene. RECORDS. "1946
ZR: Cardium politioanei Jekelius, 1944 sp. nov.
Miocene Soceni Banat. ··1965 ZR: Cardium politioanei
figd. Sarmatian, Yugoslavia [Ref. Spajic 1965]. "1977
ZR: Cerastoderma (Inaequicostata) politioanei
(Jekelius) Key to subspecies. Ref to Il'ina, Nevesskaya
& Paramonova p. 77.
politioanei suessiformis Jekelius, 1944 (subspecies). Combination to be revised: "Cardium" politioanei
suessiformis Jekelius 1944. Cardiidae. Miocene.
RECORDS. ··1946 ZR: Cardium politioanei var.
suessiformis Jekelius, 1944 nov. (type f. 15). Miocene
Soceni Bana!. ··1971 ZR: Cardium politioanei
suessiformis Jekelius 1944 Neotype Designated p. 57
Morphological variation [Ref. Hinculov 1971].
*politum. - Homonymy to be noticed. Cardium politum
from 2 sources: Spengler, 1799. - Risso, 1826.
704
Tropical Marine Mollusc Programme (TMMP)
poUtum - pondicheriense
politum Risso, 1826. - Unknown species. » Regarding
Cardium poUtum: Cardiidae. Recent . TYPE LOCALITY.
South Europe (Fischer-Piette, 1977). REMARKS. No
description, no figure. REcORDs. "1826 Poli, p. 333:
Cardium politum. --1922-32 Sherborn, p. 5077:
Cardium poUtum Risso, 1826. --1977 Fischer-Piette:
Cardium politum Risso = species insufficiently known.
politum Spengler, 1799. [Figs. Part 3 p. 896]. - Unknown
species. » Regarding Cardium politum: Cardiidae.
Recent. TYPE LOCALITY. South Seas (Pacific Ocean)
(Spengler, 1799). TYPE REPOSITORY. ZMUC. RECORDS.
--1799 Spengler, p. 46, No. 30: Cardium politum.
"1808 Spengler, p. 126: Cardium politum . "1870
Morch: Cardium politum Spgl.; resembling C.
norvegicum Spengler. --1902 Sherborn , p. 765:
Cardium politum Spengler, 1799. "1977 FischerPiette : Cardium politum Spengler. Species
insufficiently known. --1999 Knudsen & Hylleberg:
Cardium politum Spengler, 1799. - CHARACTERS
AND OCCURRENCE - DESCRIPTIONS. --1799
Spengler: shell circular, compressed, not inflated,
smooth, very glossy, inner margin finely dentate,
extending inside shell, colour outside even light yellow,
blue-grey lines imbedded in the shell, extend from
umbo to margin, inside dark yellow. DESCRIPTIONS.
--1799 Spengler: Testa cordata aequivalvi, laevigatostriata, margine intus dentato. --1799 Spengler
translated by Knudsen & Hylleberg 1999: trus Cardium ,
which also comes from the South Sea, is still smaller
than the preceding one [Cardium striatum]. Like this,
it is circular in outl ine, not vaulted, but flatly
compressed . The whole surface is smooth, shining as
polished agate. The whole internal margin of the cockle
is provided with numerous fine teeth, which reach
rather far up internally. The outside has a uniform
beautiful light yellow colour, provided with grey-blue
lines extending longitudinally under the glaze and
shining through it. Inside the colour is dark-yellow.
DIMENSIONS. Height and breadth 3/4 Tom. (20 mm).
REMARKS. --1977: Fischer-Piette considers the species
insufficiently known because it has not been illustrated
or recorded since its description by Spengler. Vidal
(1996) identified the shell as Laevicardium crassum
(label at ZMUC) . HABITAT. --1799 Spengler: South
Seas.
polonicum Alth, 1850. - Synonym of Nemocardium
fenestratum (Kner 1850). » Regarding Cardium
polonicum: Cardiidae. Fossil. RECORDS . "1922-32
Sherborn, p. 5083 : Cardium polonicum Alth, 1850.
--1986 Abdel-Gawad: Cardium polonicum Alth, 1850
= Nemocardium (Nemocardium) fenestratum (Kner
1850). --1980 Ruhoff: Cardium polonicum Alth, 1850;
vol. 3(2): 227, pI. 12, fig. 3.
polycolpatum Cossmann & Peyrot, 1911. - Changed
combination: Europicardium polycolpatum (Cossmann
& Peyrot 1911). Cardiidae. Tertiary. RECORDS. --1902
Dollfus & Dautzenberg: Cardium polycolpatum
Cossmann & Peyrot, 1912 = Cardium (Trachycardium)
multicostatum Brocchi, 1814. "1911 ZR: Cardium
(Trachycardium) polycolpatum Cossmann & Peyrot,
1911 sp. n . France Neogene. "1959 CsepreghyMeznerics : Cardium (Trachycardium) polycolpatum
Cossmann et Peyrot. --1977 Popov: Bucardium
(Europicardium) polycolpatum (Cossmann & Peyrot,
1911). "1994 Marcomini: Cardium (Trachycardium)
polycolpatum Cossmann & Peyrot, 1912. "2002
Schneider: Europicardium polycolpatum (Cossmann &
Peyrot 1911). France. Aquitanian-Burdigalian.
polymorpha (subspecies). - See under angusticostata
polymorpha Logvinenko & Starobogatov, 1967.
polyodon Solander, 1786. - Cardium polyodon Solander,
1786. RECORDS. --1786 Solander, p. 17 no. 382:
Cardium polyodon.
polypleurum Brusina. - New combination: Lymnocardium polypleurum (Brusina). Lymnocardiidae.
Fossil. RECORDS. --1987 Taktakishvili: Limnocardium
polypleurum Brus.
polyptyctus Bayan, 1870. - Changed combination:
Discors polyptyctus (Bay.). Cardiidae. Eocene and
Oligocene. RECORDS. --1899 Sacco: Discors polyptyctus
(Bay.). "1980 Ruhoff: Cardium polyptyctum Bayan,
1870; vol. 27: 461, 485 . "1999 Jacques Vidal (pers.
com.), MNHN, Paris: Fossils from the Paleontological
Collection. Cardium po/yptyctum Bayan. Upper
Lutetien. Egypt, B43628.
pomeyroliKeen, 1954. - Granocardium pomeyroli Keen,
1954. Cardiidae. Upper Cretaceous. RECORDS. --1954
Keen: Granocardium (Ethmocardium) pomeyroli Keen,
n. sp. "1954 ZR: Granocardium (Ethmocardium)
pomeyroli Keen, 1954 sp. nov. Upper Cretaceous, New
Caledonia. --1956 ZR: Granocardium (Ethmocardium)
pomeyroli from New Caledonia, now thought to be
Cretaceous [Ref. Pomeyrol 1956]. --1978 Freneix &
Mackie: Granocardium (Ethmocardium) pomeyroli
Keen, 1954.
pomeyroli pomeyroli Freneix, 1957 (form). Infrasubspecific level. Granocardium (Ethmocardium)
pomeyroli pomeyroli Freneix, 1957. Cardiidae.
Cretaceous. RECORDS. "1957 ZR: Granocardium
(Ethmocardium) pomeyroli pomeyroli Freneix, 1957
forma nov. Cretaceous, New Caledonia [Ref. Freneix
1957].
pomeyroli transversa Freneix, 1957 (form). Granocardium pomeyroli transversa Freneix, 1957.
Cardiidae. Cretaceous. RECORDS. --1957 ZR:
Granocardium (Ethmocardium) pomeyroli transversa
Freneix, 1957 forma nov. New Caledonia [Ref. Freneix
1957].
pomeyrote Keen , 1954. - Granocardium pomeyrote
Keen, 1954. Cardiidae. Cretaceous. RECORDS. --1954
ZR: Granocardium (Ethmocardium) pomeyrote Keen,
1954 sp. nov. Upper Cretaceous, New Caledonia.
ponacoides (error). - Printer's mistake for donacoides
in Andrussow 1907, p. 108.
ponchibaense Amano, 1983. - Clinocardium
ponchibaense Amano, 1983. Cardiidae. Miocene.
RECORDS. "1983 ZR: Clinocardium (Keenocardium)
ponchibaense Amano, 1983 sp. nov. Japan. Ref. to
Amano, p. 47.
ponderosum (variety). - See under norvegicum
ponderosum Bucquoy, Dautzenberg & Dollfus, 1889.
pondicheriense Orbigny, 1847. - Protocardium
pondicheriense Orbigny, 1847. Cardiidae. Cretaceous.
REMARKS. Species intennediate between Protocardium
hillanum and Protocardium altum. Cardium pondicheriense considered syn . with Cardium altum by
d'Orbigny but Stoliczka finds the two species distinct.
RECORDS. --1871 Stoliczka: Protocardium
Phuket Marine Biological Center Special Publication vol. 30 (2004)
705
Pontalmyra - portaferrica
pondicheriense d'Orbigny, 1847.
Pontalmyra Sabba Stefanescu, 1896 (genus). Lymnocardiidae. Tertiary. Erected to accommodate
Roumanian Ponti an [Tertiary basin] cardiids. RECORDs.
"1896 ZR : Pontalmyra Stefanescu, 1896 n. g.
Miocene, Roumania. "1903 Andrussoff: Pontalmyra
Sabba Stefanescu, 1896. "1905 Dollfus, commenting
Andrussow 1903: Pontalmyra. ··1980 Vokes:
Pontalmyra Stefanescu, 1896 in subfamily Didacninae.
"1981 ZR: Pontalmyra Sabba Stefanescu, 1896.
Miocene, Romania. Ref. to Papaianopol, p. 7. "1987
Taktakishvili: Pontalmyra Sabba Stefanescu, 1896.
Pontalmyra Sabba Stefanescu, 1896 (subgenus). Lymnocardiidae. Tertiary. RECORDS. ··1905 Dollfus,
commenting Andrussow 1903: Cardium planicostatum
Deshayes is type species of sub-form Pontalmyra.
Pontalmyrini Taktakishvili , 1985 (tribe) . Lyrnnocardiidae. Fossil. RECORDS. "1987 Taktakishvili :
Pontalmyrini tribe nov. ofLimnocardiinae. Type genus
Pontalmyra Sabba Stefanescu, 1896. "1987
Taktakishvili, p.85: Oraphocardium Ebersin, 1949;
referred to new tribe Pontalmyrini.
ponteleviense Mayer, 1868. - Papyridea ponteleviense
(Mayer, 1868). Cardiidae. Fossil. RECORDS. "1886
Dollfus & Dautzenberg: Cardium (Papyridea)
ponteleviense Mayer. "1902 Dollfus & Dautzenberg,
p. 330: Cardium (Papyridea) pontileviense sic Mayr,
1868. "1970 Glibert & Poel: Papyridea ponteleviense
(Mayer, 1868). "1980 Ruhoff: Cardium ponteleviense
Mayer, 1868; [1857-1869] vol. 16: 189, pI. 7, fig. 5.
pontica (variety).- See under caspia pontica Eichwald
1838.
pontica Celidje, 1953. - Chartadacna pontica Celidje,
1953. Lymnocardiidae. Fossil. RECORDS. "1953 Celidje,
p. 157: Chartadacna pontica gen. et sp. nov.
pontica Eichwald, 1838. - Synonym of Hypanis caspia
(Eichwald, 1841). OTHER JUDGEMENTS: Monodacna
pseudocardium Deshayes, 1838. - Monodacna pontica
Eichwald, 1841. » Regarding Monodacna pontica:
Lymnocardiidae. Recent. TYPE LOCALITY. Caspian Sea .
DISTRJBUTION. Caspian Sea. RECORDS. "1838 Eichwald,
p. 168-169: Monodacna pontica m. "1841 Eichwald,
p. 219: Monodacna pontica m. ··1857 Deshayes:
Cardium ponticum. "1888 Kobelt: Cardium ponticum
Eichw. = Monodacna pseudocardium. "1890
Woodward: Monodacna pontica, Eich . - Black Sea.
"1893 Ostroumoff: Cardium ponticum . "1903
Andrussoff: Eichwald, 1838 Monodacna pontica n. sp.
sic. "1908 Milaschewitch : pontica Eichw. syn. =
Monodacna pseudocardium (Deshayes) 1838. '·192232 Sherborn, p. 5 \07: Monodacna pontica Eichwald,
1838. "1922-32 Sherborn, p . 5107 : Monodacna
pontica Eichwald , 1841 . "1962 ZR: Monodacna
pontica in the etangs ofCrapina-Jijila, Roumania [Ref.
Botnariuc 1962]. "1963 ZR: Monodacna pontica from
deposits of the Danube delta [Ref. Grossu & Baltac
1963]. "1974 ZR: Monodacna pontica. Artificial pond.
Acclimatization. Ref. to Ioffe. "1975 ZR: Monodacna
pontica. Distribution. Lake USSR. Ref. to Zhuravel.
pontica carinata Grossu & Baltac, 1963 (subspecies).Monodacnapontica carinata Grossu & Baltac, 1963.
Lymnocardiidae. Quaternary. RECORDS. ··1963 ZR:
Monodacna pontica carinata Grossu & Baltac, 1963
subsp. nov. Quaternary deposits of the Danube Delta,
Rumania.
ponticllm Deshayes, 1853. - Combination to be revised:
"Cardium"
ponticum
Deshayes,
1853.
Lymnocardiidae. Fossil. "1980 Ruhoff: Cardium
ponticum Deshayes, 1853; vol. 2: 49.
ponticllm (aff.) Hal. - New combination: Lymnocardium
aff. ponticum (Hal.). Lymnocardiidae. Fossil. RECORDS.
"1987 Taktakishvili: Limnocardium aff. ponticum
(Hal.).
ponticus Ebersin, 1955 . - Valenciennius ponticus
Ebersin, 1955. Cardiacea. Fossil. RECORDS. "1955
Ebersin : Valenciennius ponticus sp. nov.
pontileviense (error). - Printer's mistake for
ponteleviense in Dollfus & Dautzenberg 1902
pontocaspia Pavlov, 1925. - Changed combination:
Didacna pontocaspia (Pavlov, 1925). Lymnocardiidae.
Pleistocene and Quaternary. RECORDS. "1926 ZR:
Cardium (Didacna)pontocaspium Pavlov, 1926 sp. n.
Tertiary S.E. Europe. "1929 ZR: Cardium (Didacna)
pontocaspium Pavlov. 1925 sp. n. Neogene &
Quaternary ofS .E. Europe. "1932 Zoological Record,
VIII. Moll. Cardiacea p. 111: Didacna pontocaspia
[Ref. Petrbok 1932]. "1963 ZR: Didacna pontocaspia,
from Quaternary deposits of the Danube delta [Ref.
Grossu & Baltac 1963]. "1985 Mikhailescu: Didacna
pontocaspia Pavlov [1925] . "1989 Mikhailesku:
Didacna pontocaspia Pavl. "1989 Klimanov,
Mikhailesku & Chepalyga: Didacna pontocaspia Pavl.
pontocaspia tanaitica Popov, 1983 (subspecies). Didacna pontocaspia tanaitica Popov, 1983. Lymnocardiidae. Pleistocene. TYPE LOCALITY. Russian SFSR.
RECORDS . "1983 Popov, p. 191, fig.: Didacna
pontocaspia tanaitica subsp. nov.
pontocrassa (subspecies). - See under crassa pontocrassa Pavlov, 1925.
Pontolamyna (error). - Printer's mistake for Pontalmyra
in Dybowski & Grochmalicki 1917 (Vokes 1980)
populosum (error). - ?Printer's mistake for porulosum
in Potiez & Michaud 1844.
poratica Mih. - Didacna poratica Mih. Lymnocardiidae.
Fossil. RECORDS. "1989 Mikhailesku: Didacna poratica
Mih. "1989 Klimanov, Mikhailesku & Chepalyga:
Didacna poratica Mih.
porcellanlls Rosewater, 1982. - Hippopus porcellanus
Rosewater, 1982. Tridacnidae. Recent. RECORDS. "1982
ZR: Hippopus porcellanus Rosewater, 1982 sp. nov.
Philippines. Ref. to Rosewater, p. 3. "1982 ZR:
Hippopus porcel/anus. Refto D' Attilio et al. / Okutani.
"1998 Poutiers : Hippoplls porcellanliS Rosewater,
1982.
*porsllgelica. - Homonymy to be noticed. Didacna
porsugelica from 2 sources: Nevesskaya, 1956. Ebersin, 1959.
porsugelica Ebersin, 1959. - Didacna porsugelica
Ebersin, 1959. Lymnocardiidae . Fossil. RECORDS.
"1959 ZR: Didacna porsugelica Ebersin, 1959 sp. nov.
porsllgelica Nevesskaya, 1956. - Didacna porsugelica
Nevesskaya , 1956. Lymnocardiidae . Quaternary.
RECORDS. "1956 ZR: Didacna porsugelica Nevesskaya,
1956 sp. nov. Khazar deposits, Quaternary. Cheleken,
Russia, Turkmenia. "1957 ZR: Didacna porsugelica
Neveskaya, 1957 sp. nov. Khazar deposits, Chelekhan
Quaternary, Turkmenia, Russia.
portaferrica Papaian. & Marin. - Changed combination:
706
Tropical Marine Mollusc Programme (TMMP)
portaferriea - postdonaeoides
Pontalmyra portaferrica (Papaian. et Marin .)
Lymnocardiidae. Fossil. REcORDs. ··1987 Taktakishvili:
Pontalmyra portaferrica (Papaian. et Marin.)
portaferriea Papaianopol, 1975. - Didacna portaferrica
Papaianopol, 1975. Lymnocardiidae. PI iocene.
RECORDS. ··1975 ZR : Didacna (Crassadacna)
portaferrica Papaianopol, 1975 sp. nov. Romania,
Mehedinti, Susita. Ref. to Papaianopol, p. 153.
portaferriea Papaianopol, 1978. - Zamphiridacna
portaferrica Papaianopol, 1978. Lymnocardiidae.
Upper Miocene. RECORDS. ··1978 ZR: Zamphiridacna
portaferrica Papaianopol, 1978 sp. nov. Romania. Ref.
to Papaianopol, 1. p. 145.
portlandieum Whitfield, 1905. - Onychocardium
portlandicum Whitfield, 1905 . Cardiacea. Fossil.
RECORDS. ··1905 ZR: Onychocardium portlandicum
Whitfield, 1905 n. sp. Portage Rocks of Lake Erie
Whitfield.
portorieoensis (variety). - See under sambaicum
portoricoensis Hubbard, 1920.
porulosllm. - Homonymy. Cardium porulosum from 2
sources: Solander in Brander, 1766. Lamarck (?year).
porulosum Lamarck. [Figs. Part 3 p. 920]. - "Cardium"
porulosum Lamarck. Cardiidae. Fossil. REMARKS. J do
not know how the authorship of Solander, 1766
compares with Lamarck. RECORDS. ··1839 Anton:
Cardium porulosum Lamarck. ··1844 Potiez &
Michaud: Cardium porulosum Lam. Tertiary of France.
porulosum Solander in Brander, 1766. [Figs. Part 3 p.
920] . - Changed combination: Orthocardium
porulosum (So lander in Brander, 1776). Cardiidae.
Lower Eocene. Tremlett (\ 950) designated Cardium
porulosum Solander, 1766 type species of subgenus
Orthocardium of genus Vepricardium. TYPE LOCALITY.
Paris Basin, France. RECORDS. ··1766 Solander in
Brander p. 39, pI. 8, fig. 99: Cardium porulosum. ··1857
Deshayes, pI. 26, figs. 1-3 : Cardium porulosum
Brander. ··1886 Cossmann: Cardium porulosum
Solander in Brander, 1776. ··1886 Vincent: Cardium
porulosum Solander. ··1902 Sherborn, p. 773: Cardium
porulosum Solander in Brander, 1766. ··1904
Cossmann & Pissarro : Cardium (Trachycardium)
porulosum Sol. ··1912 Favre: Cardium porulosum
Solander. ··1930 Stewart:. "e." porulosum. "1950
Tremlett: Cardium (Orthocardium) porulosum
Solander, 1766. ··1950 ZR: Cardium (Orthocardium)
porulosum Sol. Eocene and Oligocene Cardiidae [Ref.
Tremlett 1950] . ··1950 ZR: Cardium porulosum
Solander, Middle and Upper Eocene subgenotype of
Orthocardium Tremlett, 1950 subgen. nov. Cardium.
··1969 Moore et al.: Vepricardium (Orthocardium)
porulosum (Solander, 1766). ··1970 Glibert & Poel:
Cardium (Orthocardium) porulosum porufosum
Solander in Brander, 1766. ··1970 ZR: Trachycardium
porulosum identified from Eocene and Lower
Oligocene sands in Zeeland, [Ref. Gittenberger 1970].
··1977 Popov: Orthocardium porulosum (Solander,
1766). ··1977 Popov: Cardium porulosum Solander,
1766. Type of Orthocardium Tremlett, 1950. ··1987
Cuif, Denis, laillard & Zhu: Vepricardium porulosum.
··1987 ZR : Vepricardium porulosum. Ref. to Cuif et
al. ··1995 Le Renard et af.: Cardium (Orthocardium)
porulosum Solander in Brander, 1766. ··1995 Pacaud
& Renard: Cardium (Orthocardium) porulosum. ··1998
Schneider: Orthocardium porulosum (Solander, 1766).
··1999 Jacques Vidal (pers . com.), MNHN, Paris:
Fossils from the Paleontological Collection as
Trachycardium porulosum Solander (Lamarck?).
Lutetien. France (Bassin de Paris), i.a. ··2002
Schneider: Orthocardium porulosum (Solander, 1766).
One of the very first species of any sort of fossil to be
given a Linnean binomen.
porulosum asiatica Kakhanova, 1957 (variety) . Nomen
nudum. » Regarding Cardium porulosum var.
asiatica: RECORDS. ··1957 ZR: Cardium porulosum var.
asiatica Kakhanova, 1957 nov. as nom. nud. Eocene,
Tadgic depression, Middle Asia; Tadzhiktistan
Republic, USSR?
porulosum belgiellm Glibert, 1933 (variety) . Upgraded . New combination: Orthocardium
porulosum belgicum (Glibert, 1933). Cardiidae.
Bruxellien. RECORDS. "1933 ZR : Cardium
(Trachycardium) porulosum var. belgicum Glibert,
1933 nov. Bruxellien Belgium.
porlliosum highcliffensis Tremlett, 1950 (variety). Upgraded . Orthocardium porulosum highclifJensis
(Tremlett, 1950). Cardiidae . Oligocene. RECORDS .
··1950 ZR: Cardium (Orthocardium) porulosum Sol.
var. highclifJensis Tremlett, 1950 nov. Hampshire
Basin, Highcliffp. 130 pI. 19 f. 21 English Eocene and
Oligocene Cardiidae.
porulosllm hopneri Glibert & Poel, 1970 (subspecies).
- New combination: Orthocardium porulosum hopneri
Glibert & Poe I, 1970. Cardiidae. Fossil. RECORDS.
··1970 Glibert & Poel, p. 52: Cardium (Orthocardium)
porulosum hopneri nov. subsp. ··1970 ZR: Cardium
(Orthocardium) porulosum hopneri Glibert & Van De
Poel, 1970 subsp. nov. Lutetian, Eocene, Parnes, Paris
Basin, France. ··1995 Le Renard et al. : Cardium
(Orthocardium) porulosum hopneri Glibert & Van De
Poel,1970.
porulosum keeni Glibert, 1933 (subspecies). - Nom.
mut. New combination: Orthocardium porufosum
belgicum Glibert, 1933. Cardiidae. Eocene. RECORDS.
··1936 ZR: Laevicardium (Trachycardium) porulosum
keeni nom. mut. = belgicum Glibert, 1933 Wemmel
Sands, Eocene Belgium, [Ref. Glibert 1933] . .. 1970
Glibert & Poel: Cardium (Orthocardium) porulosum
keeni Ghbert, 1933 .
porulosum subporulosum d'Orbigny 1850 (subspecies).
- New combination: Orthocardium porulosum
subporulosum d' Orbigny 1850. Cardiidae. Fossil.
RECORDS. ··1995 Le Renard et af.: Cardium
(Orthocardium) porulosum subporufosum d'Orbigny
1850.
postearditoides Nalivkin & Anisimov. - Didacna
postcarditoides Nalivkin & Anisimov, 1916 .
Lymnocardiidae . Pliocene. RECORDS. ··1916 ZR:
Didacna postcarditoides Nalivkin & Anisimov, 1916
sp. n. Pliocene Russia.
postcimmerica Davidachvili, 1933. - Chartoconcha
postcimmerica Davidachvili, 1933. Cardiacea. Upper
Pliocene . RECORDS. ··1934 ZR: Chartoconcha
postcimmerica Davidachvili, 1933 sp. n. Upper
Pliocene Georgia, Caucasus . ··1977 Popov :
Chartoconcha postcimmerica.
postdonacoides (variety). - See under donacoides
postdonacoides Davidachvili, 1933.
Phuket Marine Biological Center Special Publication vol. 30 (2004)
707
postdonacoides - praeechinata
postdonacoides Davitashvili. - Pseudocatillus
postdonacoides (Davitashvili). Lymnocardiidae. Fossil.
RECORDS. ··1967 Ebersin: Pseudocatillus postdonacoides (David.). ··1984 ZR: Pseudocatillus postdonacoides Davitashvili. Referred to new subgenus.
P (Donacicatillus). Ref. to Akhvlediani p. 71.
postduabica Davidachvili, 1933. - Prosodacna
postduabica Davidachvili, 1933. Lymnocardiidae.
Upper Pliocene. RECORDS. ··1934 ZR: Prosodacna
postduabica Davidachvili, 1933 sp. n. Upper Pliocene
Caucasus.
postmoutonianum Poyarkova, 1978. - New
combination: Trachycardium postmoutonianum
(Poyarkova, 1978). Cardiidae. Upper Cretaceous.
RECORDS. ··1978 ZR: Cardium (Trachycardium)
postmoutonianum Poyarkova, 1978 sp. nov.
Kazakhstan. Ref. to Poyarkova p. 112.
postoxypleura Akhvlediani, 1984. - Pontalmyra
postoxypleura Akhvlediani, 1984. Lymnocardiidae.
Pliocene. RECORDS. ··1984 ZR: Pontalmyra
postoxypleura sp. nov. Georgia SSR. Ref. to
Akhvlediani p. 55.
pracedentula Eberzin, 1956. - Pteradacna pracedentula
Eberzin, 1956. Lymnocardiidae. Fossil. RECORDS.
··1956 ZR: Pteradacna pracedentula Eberzin, 1956
sp. nov. Kerchenski, Crimea, Russia.
pracipitatumn Slodkewitsch, 1932. - Combination to
be revised: "Cardium" pracipitatumn Slodkewitsch,
1932. Cardiidae. Paleogene. RECORDS. ··1932 ZR:
Cardium
(Trachycardium?) pracipitatumn
Slodkewitsch, 1932b sp. n. Paleogene S. Russia.
praeaculeatum Holzl, 1958. - New combination:
Acanthocardia praeaculeatum (Holzl, 1958).
Cardiidae. Miocene. RECORDS. ··1958 ZR: Cardium
(Acanthocardia) praeculeatum sic Holzl, 1958 sp. nov.
Burdigalian, Miocene. Kaltenbach excavation, Upper
Bavaria, Germany.
praeangusticostata Eberzin, 1956 - Stenodacna
praeangusticostata Eberzin, 1956. Lymnocardiidae.
Pliocene. RECORDS. ··1956 ZR: Stenodacna
praeangusticostata Eberzin, 1956 sp. nov. Kuban.
Kerchenski, Crimea, Russia. Pliocene. ··1977 Popov:
Stenodacna praeangusticostata.
praeannulatus (subspecies). - See under annulatus
praeannulatus Ebersin, 1955.
praearenicola - Combination to be revised: "Cardium"
praearenicola. Cardiidae. Fossil. RECORDS. "1946
Abrard: Cardium praearenicola.
praeblandum Keen, 1954. - Clinocardium praeblandum
Keen, 1954. Cardiidae. Upper Miocene. RECORDS.
"1954 ZR: Clinocardium praeblandum Keen, 1954b
sp. nov. Briones formation, Upper Miocene Contra
Costa Co. California. ··1977 Popov: Clinocardium
praeblandum Keen, 1954.
Praecardium (genus). - Not a cardiid genus. »
RegardingPraecardium: Silurian. REcoRDs. ··1926 ZR:
Praecardium latecostatum Heller, 1926 Silurian
Germany.
praecedens Mayer, 1858. - Combination to be revised:
"Cardium" praecedens Mayer. Cardiidae. Miocene.
REcORDs. ··1980 Ruhoff: Cardium praecedens Mayer,
1858; [1857-1869] vol. 7: 187. "1999 Jacques Vidal
(pers. com.), MNHN, Paris: Fossils from the
Paleontological Collection. Cardium praecedens
Mayer. Helvetien. Suisse, B30825, B30826.
praecipitatum Slodkewitsch, 1932. - Combination to be
revised: "Cardium" praecipitatum (Slodkewitsch,
1932). Cardiidae. Fossil. REcORDs. ··1932 ZR: Cardium
(Trachycardium) praecipitatum Slodkewitsch, 1932a
"sp. nova".
praechinatum otarii Kharatishvili, 1958 (variety). Upgraded. Combination to be revised: "Cardium"
praechinatum otarii Kharatishvili, 1958. - Cardiidae.
Miocene. RECORDS. "1958 ZR: Cardium (Cardium)
praechinatum var. otarii Kharatishvili, 1958 nov. Kaspi
Sakar Aul horizon, Miocene. Georgia USSR
praecoloratus Eberzin, 1967. - Pseudocatillus
praecoloratus Eberzin, 1967. Lymnocardiidae.
Pliocene. RECORDS. ··1967 ZR: Pseudocatillus
praecoloratus Eberzin, 1967 sp. nov. Pliocene, USSR.
praecrenulata Eberzin, 1962. - Didacna praecrenulata
Eberzin, 1962. Lymnocardiidae. Pliocene. RECORDS.
··1962 ZR: Didacna (Pontalmyra) praecrenulata
Eberzin, 1962 sp. nov. Kerchenskii; Azov region;
Pliocene, Russia. ··1987 Taktakishvili: Ponta/myra
praecrenulata (Ebers.).
praeculeatum (error). - ?Printer's mistake for
praeaculeatum in ZR for 1958.
praecurrens Stoliczka, 1871. - Changed combination:
Profragum praecurrens (Stoliczka, 1871). Cardiidae.
Late Cretaceous. TYPE LOCALITY. North of Odium and
west of Anapaudy, South India. REMARKS. Resembles
a recent sp. living on coral reefs in the Nicobar islands
(Stoliczka 1871). REcoRDs. "1871 Stoliczka, p. 222,
pI. II, figs. 1-2: Fragum praecurrens Stoliczka. "1980
Ruhoff: Fragum praecurrens Stoliczka, 1870; [18701871] vol. 3: 222, pI. 11, figs. 1-2. "1998 Schneider:
Profragum praecurrens (Stoliczka, 1871). ··2001.
Schneider & Carter: Profragum praecurrens (Stoliczka,
1871). Cardiidae. Late Cretaceous, India.
praecygnorum Ludbrook, 1955. - Changed
combination: Vasticardium praecygnorum (Ludbrook,
1955). Cardiidae. Pliocene. TYPE LOCALITY. Dry Creek
Sands (Pliocene), Adelaide, South Australia. REcORDs.
··1955 ZR: Vasticardium (Regozara) praecygnorum
Ludbrook, 1954 sp. nov. nom. nud. Pliocene, Australia.
··1977 Popov: Trachycardium (Vasticardium)
praecygnorum (Ludbrook, 1955). "1999 Vidal:
Vasticardium (Regozara) praecygnorum Ludbrook,
1955. - CHARACTERS AND OCCURRENCE DESCRIPTIONS. ··1999 Vidal: this species has 48 ribs
(Ludbrook 1955: 61)
praedesertum Strausz, 1930. - New combination:
Lymnocardium praedesertum (Strausz, 1930).
Lymnocardiidae. Miocene. RECORDS. ··1941 ZR:
Limnocardium praedesertum Strausz, 1930 sp. nov.
[Miocene], Hungary.
praeduboisi - Panticapaea praeduboisi. Cardiacea.
Pliocene. RECORDS. ··1956 ZR: Panticapaea
praeduboisi sp. nov Pliocene. Kerchenski, Crimea
Peninsula, and Kuban Russia. ··1977 Popov:
Panticapaea praeduboisi.
praeechinata Hilber, 1882. - Synonym of Acanthocardia
turonica (Homes, 1861). Cardiidae. Middle Miocene.
RECORDS. ··1902 Dollfus & Dautzenberg: Cardium
praeechinatum Hilber. ··1958 ZR: Cardium (Cardium)
praechinatum Miocene. [Ref. Kharatishvili 1958].
··1970 Glibert & Poel: Cardium (Acanthocardia)
708
Tropical Marine Mollusc Programme (TMMP)
praeechinatum - praepermicus
praeechinatum Hilber, 1882. ··1977 Popov: Acanthocardia (Acanthocardia) praeechinata (Hilber, 1882).
··1986 Studencka, p. 69: Cardium praeechinatum
Hitber, 1882; syn. nov.; of Acanthocardia
(Acanthocardia) turonica (Homes, 1861). ··1987 ZR:
Cardium praeechinatum Hilber, 1882. New synonym
of Acanthocardia (Acanthocardia) turonica (Homes,
1861). Ref. to Studencka p. 69.
praeechinatum (mode). - See under praeechinata.
praeechinatum Mayer mss. - Combination to be revised:
"Cardium" praeechinatum Mayer. Cardiidae. Fossil.
RECORDS . ··1902 Dollfus & Dautzenberg: Cardium
praeechinatum Mayer mss.
praeextensa (variety). - See under propinqua
praeextensa Androussov, 1923.
praeflScherianum Kolesnikov, 1929. - Protractiforma
praejischerianum (Kolesnikov, 1929). Cardiidae.
Sarmatian . RECORDS . ··1929 ZR: Cardium
praejischerianum Kolesnikov, 1929 sp. n. Sarmatian.
··1979 ZR: Protractiforma praefischerianum
(Kolesnikov). Referred to new subgenus P.
(Fischericardium) Ref. to Bagdasaryan, p. 44.
praeformis (subspecies). - See doenginki praeformis
Papp,1954.
praehellesponticus Eberzin, 1967. - Pseudocatillus
praehellesponticus Eberzin, 1967. Lymnocardiidae.
Pliocene. RECORDS. ··1967 ZR: Pseudocatillus
praehellesponticus Eberzin, 1967 sp. nov. Pliocene,
USSR.
praehispidijorme Bagdasaryan, 1970. - Combination to
be revised: "Cardium" praehispidiforme Bagdasaryan,
1970. Cardiidae. Miocene. RECORDS. ··1970 ZR :
Cardium (Papillicardium) praehispidiforme
Bagdasaryan, 1970 sp. nov. Miocene, Chkumi, western
Georgia, USSR.
praehyrcania Tscheltzov, 1965. - Changed combination:
Cerastoderma praehyrcania Tscheltzov, 1965 .
Cardiidae. Pliocene. RECORDS. ··1965 ZR: Cardium
praehyrcania Tscheltzov, 1965 sp. nov. Akchagylian
(Pliocene), Turkmenia, USSR . ··1977 Popov :
Cerastoderma praehyrcania.
praehyrcanicllm Baba-Zade 1967. - Combination to be
revised: "Cardium " praehyrcanicum Baba-Zade 1967.
Cardiidae. Cenozoic. RECORDS. ··1967 ZR : Cardium
praehyrcanicum Akchagyl deposits (Cenozoic),
Azerbaidjan, SSR [Ref. Baba-Zade 1967).
praeinflatum Papp, 1953 - New combination:
Lymnocardium praeinflatum Papp , 1953.
Lymnocardi idae. Fossil. RECORDS. ··1948 ZR :
Limnocardium praeinjlatum Papp, 1948 'no sp.' [nomen
nud.] [Ref. Papp 1948]. "1956 ZR: Laevicardium
praeinflatum Papp, 1953 sp. nov. ··1987 Taktakishvili:
Limnocardium praeinjlatum Papp.
praekarpinskyi Ebersin, 1962. - Pontalmyra
praekarpinskyi (Ebersin, 1962). Lymnocardiidae .
Fossil. RECORDS. "1962 ZR: Didacna (Pontalmyra)
praekarpinskyi Eberzin, 1962 sp. nov. N .W.
Kerchenskogo region; Pliocene, Russia. ··1987
Taktakishvili: Pontalmyra praekarpinskyi (Ebers.).
praelaeiviuscula Andrussov, 1924. - Adacna
praelaeiviuscula Andrussov, 1924. Lymnocardiidae.
Pleistocene. RECORDS. ··1924 ZR: Adacna
praelaeiviuscula sic Andrussov, 1924 Pleistocene.
praelaeviuscula Andrusov, 1963. - Adacna
praelaeviuscula Andrusov, 1963. Lymnocardiidae.
Pliocene. RECORDS. ··1963 ZR: Adacna (Adacna )
praelaeviuscula Andrusov, 1963 sp. nov. Apsheron
layers, Pliocene, Russia [Ref. Andrusov 1963].
praemodiolaris Chelidze, 1975. Plagiodacna
praemvdiolaris Chelidze, 1975. Lymnocardi idae.
Lower Pliocene. RECORDS. ··1975 ZR: Plagiodacna
praemodiolaris Chelidze, 1975 sp. nov. Georgia SSR,
Zugdid. Ref. to Chelidze, p. 192.
praemoquicum Davidasvili, 1930 - Changed
combination: Moquicardium praemoquicum
(Davidasvili, 1930). Lymnocardiidae. Fossil. REcORDs.
··1932 ZR: Limnocardium praemoquicum Davidasvili,
1930 sp. n. Pliocene Transcaucasia. ··1957 ZR:
Limnocardium praemoquicum Pliocene [Ref. Chelidize
1957]. ··1958 ZR: Limnocardium (Moquicardium)
praemoquicum Pliocene [Ref. Eberzin 1958]. ··1987
Taktakishvili: Limnocardium (Moquicardium) praemoquicum Dav. ··1987 Taktakishvili: Moquicardium
praemoquicum Dav.
praemoquicum alta Ebersin, 1958 (variety). - Upgraded.
New combination: Moquicardium praemoquicum alta
(Ebersin, 1958). Lymnocardiidae. Fossil. RECORDS.
··1958 ZR: Limnocardium (Moquicardium)
praemoquicum var. alta Eberzin, 1958 nov. Abkhazia,
Mokvi, east coast of Black Sea. Pliocene of the Ponto
Caspian basin, Russia. ··1987 Taktakishvil i:
Limnocardium (Moquicardium) praemoquicum Dav.
var. alta Ebers.
praemoquicum suhmoquicum Ebersin (variety). Upgraded . New combination : Moquicardium
praemoquicum
submoquicum
(Ebersin).
Lymnocardiidae. Fossil. REcoRDs. ··1987 Taktakishvili:
Limnocardium praemoquicum var. submoquicum
Ebers.
praemoquicum subpraemoquicum Chelidize, 1957
(variety). - Upgraded. New combination:
Moquicardium praemoquicum subpraemoquicum
(Chelidize, 1957). Lymnocardiidae. Pliocene. REcORDs.
··1957 ZR: Limnocardium praemoquicum var.
subpraemoquicum Chelidize, 1957 nov. Makharadze
river Zhuzhi, Pliocene.
praeobtusa Finlay & Marwick, 1937. - Lahillia
praeobtusa Finlay & Marwick, 1937. Cardiacea.
Danian. RECORDS. ··1937 ZR: Lahillia (?Lahilleona)
praeobtusa Finlay & Marwick, 1937 sp. n. Danian
Wangaloa, New Zealand.
*praepapillosum. - Homonymy to be noticed. Cardium
praepapillosum from 2 sources: Baldi, 1966. - ?author
before 1891.
praepapil/osum - Combination to be revised:
"Cardium" praepapi//osum. Cardiidae. Pliocene.
RECORDS. ··1891 ZR: Cardium praepapillosum
Pliocene of Altavilla [Ref. Gregorio 1891].
praepapillosum Baldi, 1966. - Combination to be
revised: "Cardium" praepapil/osum (Baldi, 1966).
Cardiidae. Oligocene. RECORDS. ··1966 ZR: Cardium
(Parvicardium) praepapillosum Baldi, 1966 sp. nov.
Oligocene, Eger, Hungary.
praepermicus Fredericks, 1916. Conocardium
praepermicus Fredericks, 1916. Conocardiidae.
Carbon. RECORDS. ··1916 ZR: Conocardium
praepermicus Fredericks, 1916. sp. n. Ural Upper
Carboniferous. ··1926 ZR: Conocardium praepermicus
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praeplicatum - prigorovskii
Fredericks, 1926 sp. n. Carbon.
praeplicatum Hilber 1882. - Cerastoderma
praeplicatum (Hilber 1882). Cardiidae. Miocene.
RECORDS. "1974 ZR: Cerastoderma praeplicatum.
Interspecific variation. Miocene. Ref. to Baghdasariani.
"1977 Popov: Plicatiforma praeplicatum. "1977 ZR:
Cerastoderma (Plicatiforma) praeplicatum. Key to
subspecies. Ref to Il'ina, Nevesskaya & Paramonova
p. 70. "1979 ZR: Cerastoderma (Plicatiforma)
praeplicatum (Hilber 1882). Neotype designated. Ref.
to Paramonova, p. 45. "1979 ZR: Cerastoderma
(Plicatiforma) praeplicatum praeplicatum (Hilber
1882). Ref. to Paramonova, p. 46.
praeplicatum pseudoplicatum Friedberg 1934
(subspecies). - Cerastoderma praeplicalum
pseudoplicatum (Friedberg 1934). Cardiidae. Fossil.
RECORDS. "1977 Popov: Plicatiforma praeplicatum
pseudoplicalum . "1979 ZR: Cerastoderma
(Plicatiforma) praeplicatum pseudoplicalum
(Friedberg 1934). Ref. to Paramonova, p. 48.
praesandbergeri Holzl, 1962. - Combination to be
revised : "Cardium" praesandbergeri Holzl, 1962.
Cardiidae. Oligocene. RECORDS. "1962 ZR: Cardium
praesandbergeri Holzl, 1962 sp. nov. N. of T61z
Chattian, Oligocene, Bavaria.
praesquamu/osa (mode). - See under praesquamulosum.
praesquamulosum - Tauricardium praesquamulosum.
Lymnocardiidae. Fossil. REcORDs. "1987 Taktakishvili:
Tauricardium praesquamulosum. "1987 Taktakishvili:
Tauricardium praesquamulosa.
praesubsquamulosum (variety). - See under subsquamulosum praesubsquamulosum Eberzin, 1958.
praesquamu/osum minor. - Tauricardium praesquamulosum minor. Lyrnnocardiidae. Fossil. REcORDs.
"1987 Taktakishvili: Tauricardium praesquamulosum
minor.
praetenue Mayer. - Combination to be revised:
"Cardium" praetenue Mayer. Cardiidae. Pliocene.
RECORDS. "1886 Fischer: Cardium praetenue Mayer.
"1999 Jacques Vidal (pers. com.), MNHN, Paris:
Fossils from the Paleontological Collection as Cardium
praetenue Mayer. Plio. inf. France, B35022 .
praetrigonoides Nalivkin & Anisimov, 1916. - Didacna
praelrigonoides Nalivkin & Anisimov, 1916 .
Lymnocardiidae. Pliocene. RECORDS . "1916 ZR:
Didacna praetrigonoides sic Nalivkin & Anisimov,
1916 sp. n. Pliocene Russia. "1964 ZR: Didacna
praetrigonoides from the Caspian, biometrical study
[Ref. Asadullaev 1964]. "1968 Logvinenko &
Starobogatov: Didacna praelrigonoides Nat. & Anis.,
1945 = Didacna trigonoides praetrigonoides Nat. et
Anis. "1977 Popov: Didacna praelrigonoides.
praetrigonoides abescunica Popov, 1983 (subspecies).
- Didacna praelrigonoides abescunica Popov, 1983.
Lymnocardiidae. Pleistocene. TYPE LOCALITY. Russian
SFSR. RECORDS. "1983 Popov, p. 197, fig.: Didacna
praetrigonoides abescunica subsp. nov.
praetrigonoides inderana Svitoc, 1967 (subspecies). Didacna praetrigonoides inderana Svitoc, 1967.
Lymnocardiidae. Quaternary. RECOR DS. "1967 ZR:
Didacna praetrigonoides inderana Svitoc, 1967 subsp.
nov. figs.; Quaternary, central Caspian region [Ref.
Svitoc 1967].
praetrigonoides obunca Popov, 1983 (subspecies). -
Didacna praelrigonoides obunca Popov, 1983 .
Lymnocardiidae. Pleistocene. TYPE LOCALITY. Russian
SFSR. RECORDS. "1983 Popov, p. 198, fig.: Didacna
praetrigonoides obunca subsp. nov.
prahovensis (subspecies). - See under subodessae
prahovensis Papaianopol, 1974.
prahovensis Papaian. - Euxinicardium prahovensis
Papaian . Lymnocardiidae. Fossil. RECORDS. "1987
Taktakishvili: Euxinicardium prahovensis Papaian.
prahovensis Papaianopol, 1989. - Pachydacna
prahovensis Papaianopol , 1989. Lymnocardiidae.
Pliocene. TYPE LOCALITY. Bassin Dacique, Romania.
RECORDS. "1989 Papaianopol, p. 85, fig.: Pachydacna
(Pachydacna) prahovensis sp. nov.
prashadi Vidal, 1994. - Fulvia prashadi Vidal, 1994.
Cardiidae. Recent. TYPE LOCALITY. Indonesia. RECORDS.
"1994 Vidal, p. III , pI. 3, figs. 8 a,b : Fulvia
(Laevifulvia) prashadi sp. nov.
pratenue Meyer in Capellini, 1879. - Combination to
be revised: "Cardium" prate/1Ue (Meyer in Capellini,
1879). Cardiidae. Tertiary. RECORDS. "1987 Anaya:
Cardium pratenue Mayer.
pratellue (aff.) Meyer in Capellini, 1879. - RECORDS.
"1970 ZR: Cardium aff. Cardium praetenue Meyer in
Capellini, 1879, see under Monodacna tenuicostala sp.
nov. sic [?Ref. ?Gillet 1970].
pratenue cata/aunica Almera et Bofill, 1894 (variety).
- Upgraded. New combination: Lymnocardium
pratenue catalaunica (Almera et Bofill, 1894). Lymnocardiidae. Pliocene. RECORDS. "1987 Anaya: Cardium
pratenue Mayer var. catalaunica Almera et Bofill,
1894. "1987 Anaya: Limnocardium (Euxinicardium)
pratenue calalaunica (Almera et Bofill , 1894).
prattenanum Hall, 1858. - Conocardium prattenanum
Hall, 1858. Conocardiidae. Fossil. REMARKS. Conocardiids have no connection with the Cardiacea except
analogy of form, which does not offer any point of
relationship (Dall 1895, p. 519). RECORDS. "1980
Ruhoff: Conocardium prattenanum Hall, 1858; vol. 4:
15.
Pratu/um Iredale, 1924 (genus). - Cardiidae. Recent.
Subgenus (Pratulum) upgraded to generic level by
authors. RECORDS. "1924 ZR: Pratulum Iredale, 1924b
gen nov. for Cardium thetidis Hed . "1938 Cotton &
Godfrey: Pratulum Iredale 1924. "1977 Popov:
Pratulum Iredale, 1924. Genotype: Cardium thetidis
Hedley, 1902. "1978 Maxwell : Genus Pratulum
Iredale, 1924. "1978 ZR: Pratulum Iredale 1924,
Status. Ref. to Wilson et al. p. 69. "1980 Vokes:
Pratulum Ireda1e, 1924 in subfamily Protocardiinae.
"1992 Poutiers: Pratulum JredaJe 1924.
Pratulum Iredale, 1924 (subgenus). - Cardiidae. Recent.
GENOTYPE. Type species: Iredale (1924) designated
Cardium thetidis Hedley, 1902 type species of subgenus
(Prall/lum) of genus Nemocardium Meek , 1876.
Subsequently raised to generic level. RECORDS. "1954
Keen: (Pratl/lum) Iredale, 1924. - CHARACTERS
AND OCCURRENCE - DESCRIPTIONS. "1994
HylJeberg: more equilateral than Nemocardium
(Nemocardium); sculpture of fine radial ribs but coarser
on anterior slope; with very fine concentric ridges.
prigorovskii Boghachev, 1938 . - New combination:
Lymnocardium prigorovskii Boghachev, 1938. Lymnocardiidae. Fossil. RECORDS . "1939 ZR: Limnocardium
710
Tropical Marine Mollusc Programme (TMMP)
primigenillm - pristipleura
prigorovskii Boghachev, 1938 sp. n. ··1981 ZR:
Cardium prigorovskii Bogatschev is a senior synonym
of C helmerseni Ilyina, 1960. and C levinae Merklin,
1960. Ref. to Vialov et af. p. 45.
primigenillm Eberzin & Stevanovic, 1949. Arcicardium primigenium Eberzin & Stevanovic, 1949.
Lymnocardiidae. Pontic. RECORDS. ··1949 ZR:
Arcicardium primigenium Eberzin & Stevanovic, 1949
[sp. n.?] Pontic of Servia. "1949 ZR: Arcicardium
primigenium as the oldest representative of the genus.
[Ref. Eberzin & Stevanovic 1949].
primuillm Barrande. - Praecardium primulum Barrande.
Cardiacea. Fossil. RECORDS. ··1999 Jacques Vidal (pers.
com.), MNHN, Paris: Fossils from the Paleontological
Co llection. Praecardium primulum Barrande.
Bohemia, B02612.
prinopleura Andrus. - Sp. inedita. » Regarding
Prosodacna prinopleura: Lymnocardiidae. Pliocene.
RECORDS. ··1932 ZR: Prosodacna prinopleura Andrus.
sp. inedita p. 184 pI. xiii f. 1-5 Pliocene Transcaucasia,
[Ref. Davidasvili 1930a].
prionophorum Brusina. - New combination:
Lymnocardium
prionophorum
(Brus.).
Lymnocardiidae. Fossil. RECORDS. ··1987 Taktakishvili:
Limnocardium prionophorum (Brus.).
Prionopleura Ebersin, 1949 (genus). - Lymnocardiidae.
RECORDS. "1974 ZR: Prionopleura Ebersin, 1949.
Referred to new subfamily Prosodacninae. Ref. to
Andreescu, p. 128. ··1980 Vokes: Prionopleura
Ebersin, 1949 in subfamily Lymnocardiinae. ··1987
Taktakishvili: p. 162 designated Prionopleura Ebersin,
1949 type genus of new tribe Prionopleurini.
prionoplellra Wassoievic & Eberzin, 1931. - Prosodacna
prionopleura Wassoievic & Eberzin, 1931. Lymnocardiidae. Pliocene. RECORDS. ··1931 ZR: Prosodacna
prionopleura Wassoievic & Eberzin, 1931 sp. n.
prionopleura davilasrilii Wassoievic & Eberzin, 1931
(subspecies). - Prosodacna prionopleura davilasrilii
Wassoievic & Eberzin, 1931. Lymnocardiidae.
Pliocene. RECORDS. ··193 I ZR: Prosodacna prionopleura davilasrilii Wassoievic & Eberzin, 1931 sp. n.
prionopleura krestovnikoviWassoievic & Eberzin, 1931
(subspecies). - Prosodacna prionopleura krestovnikovi
Wassoievic & Eberzin, 1931. Lymnocardiidae.
Pliocene. RECORDS. ··1931 ZR: Prosodacna
prionopleura krestovnikovi Wassoievic & Eberzin,
1931 sp. n.
prisca (subspecies) Douville, 1917. - See under hillana
prisca Douville, 1916.
prisca Douville 1916. - Pr%cardia prisca Double 1916.
Cardiidae. Fossil. RECORDS. ··1980 ZR: Protocardia
prisca Douville 1916 raised from Protocardia hillana
prisca. Type designation p. 173. Ref. to Freneix.
priscae Strausz , 1942. - New combination :
Lymnocardium priscae Strausz, 1942. Lymnocardiidae.
Pre. Pliocene. RECORDS. ··1947 ZR: Limnocardium
(Pontalmyra) priscae Strausz, 1942 sp. nov. Somlojeno
(Upper Pannon) [Pre. Pliocene]. ··1987 Taktakishvili:
Limnocardium priscae Strausz.
pristinum Keen, 1954. - Clinocardium pristinum Keen.
Cardiidae. Middle and Upper Miocene. RECORDS.
··1954. Keen, p. 322: Clinocardium pristinum Keen n.
sp. n. ··1954 ZR: Clinocardium pristinum Keen, 1954
sp. nov. Neroly formation, Upper Miocene, Contra
Costa Co. California. ··1977 Popov, p. 52:
Clinocardium pristonum sic Keen, 1954.
pristipleura (mode). - See under pristipleurum Dall,
1901.
pristipieura Dall, 1901. - Changed combination:
Acrosterigma pristipleurum(Dall, 190 I ).Cardiidae.
Recent. TYPE MATERIAL. Types of Cardium pristipleura
have not been traced but Cardium maculosum Sowerby,
1833 is a synonym, and type material of this species is
available (see maculosum Sowerby). TYPE LOCALITY.
Tres Marias Islands, west Mexico, on the sands
(Hertlein & Strong 1947). RECORDS. ··1900 Dall, p. 389:
Cardium (Trachycardium) pristipleura Dall, 1900 cited
from Vidal (1999). ··190 I Dall: Cardium
(Trachycardium) pristipleura Dall, 1900. "1901 Dall:
Cardium pristipleurum a new name for C maculosum
Sowerby, 1833 (not C maculosum Wood, 1815). ··1932
Pilsbry & Lowe : Cardium (Trachycardium)
pris/ipleura Dall 1900 = (C maculosum Sby. 1833,
non Wood 1815). "1947 Hertlein & Strong: Cardium
(Acroslerigma) pristipleura Dall = Cardium
maculosum Sowerby. ··1953 Olsson & Harbison:
Cardium (Acrosterigrna) pristipieura Dall. ··1971 Keen
& Mclean: Trachycardium (Acrosterigma) pristipleura
(DaJ1, 1901). ··1977 Popov: Acrosterigm a pristipleura
(Dall, 190 I). "1977 Fischer-Piette: Laevicardium
pristipleura Dall. ··1977 Vokes: Cardium (Trachycardium) pristipleura Dall, 190 I = Cardium
maculosum Sowerby, 1833 (non Wood, 1815). "1977
Vokes: Cardium (Trachycardium) pristipleura Dall,
190 I = Cardium maculatum Sowerby, 1840 (non
Gmelin, 1791) = Cardium maculosum Sowerby, 1833.
"1991 Matsukuma, Okutani & Habe: Trachycardium
pristipleura (Dal1, 190 I). ··1999 Vidal: Acrosterigma
pristipleura (Dall, 1900). - CHARACTERS AND
OCCURRENCE - DESCRIPTIONS. "1971 Keen: Shell
elongately obliquely ovate, gently convex; ornamented
by 34-39 flattened, laterally imbricated ribs; coloration
whitish-yellow and reddish-brown exteriorly, the
interior white. The ribs, numbering from 34 to 39, are
flattened, with very narrow interspaces, and the anterior
ribs show a file-like roughening along each edge. The
color of the exterior is yellowish white, with reddishbrown markings and often with pinkish-purple
concentric bands or blotches. The interior is white.
There is a thin light-brown periostracum that persists,
mainly toward the ventral margin. Height, about 65
rnm. ··1999 Vidal : shells ovoid to 'pear-shaped", rarely
elliptical, and equilateral with possible weak truncation
on posterior margin. Always elongated in adult stage,
width rather variable. Ribs straight or slightly curved
in projection . Lunular area rather large. Exterior
coloured orange, pink, brown and/or purple; interior
white, with posterior margin generally orange and
ventral and anterior margins purple. Hinge line
moderately arched. No sterigma observed. Rib number
mean 35.0 ± 2.\' (range 31-41). Posterior slope with
wide ribs anteriorly, ribs reduced and with very small
scales on posterior part of slope. Median part of shell
with low, flat ribs overhanging thin interstices; regularly
notched bottom of interstices on juvenile shells
(elongated or crescent-shaped notches, which continue
on lower sides of ribs); contact between interstices and
ribs marked, on both sides, by a very thin longitudinal
Phukel Marine Biological Cenler Special Publication vol. 30 (2004)
711
pristis - proboscideum
depressed line. The last rib of MPQ, mainly on right
valve, can bear a longitudinal gash on its anterior part.
DIMENSIONS. "1999 Vidal: length 66.3, width 87.3,
breadth (calculated) 47.4 mm. REMARKS. --1999 Vidal:
Acrosterigma pristipleura conforms with the speciesgroup diagnosis as far as rib morphology is concerned,
and is very close to the genotype A. dalli, in that its
ribs are "flattened and peculiarly approximated,
interstices. deep narrow cuts" (Reeve 1844, Sp. 58).
Acrosterigma pristipleura, when adult, is easily
distinguishable from A. magnum , the other living
representative of the species-group in the Americas,
mainly by its flat low ribs and very thin interstices. It
is a rather uncommon species, little cited in the
literature. HABITAT. "1947 Hertlein & Strong: shallow
water at depth of 3-6 fathoms, sand, algae, crushed
shell. "1999 Vidal : recorded from 18 to 33 m.
DISTRIBUTION. "1971 Keen & McLean: this is not a
common species although it may be found by shore
collectors anywhere between the southern portion of
the Gulf of California and Guayaquil, Ecuador. --1999
Vidal: a stretch of about 3800 Ian from the southern
portion of Gulf of California to Ecuador. Dubious and
wrong localities. - Vidal (1999): Monterey, California
(MNHN du Petit Thouars 1839) - Philippines (LACM
50904 Burch).
pristis Valenciennes, 1827. [Figs. Part 1 p. 91. Part 3 p.
895]. - Laevicardium pristis (Valenciennes, 1827).
OTHER JUDGEMENTS: Cardium serratum Linne. Cardium laevigatum Linne. » Regarding Cardium
pristis: Cardiidae. Recent. TYPE MATERlAL. Holotype at
MNHN, Paris. RECORDS. --1798 Bruguiere. ID by
Lamarck, pI. 299. fig. 3. Cardium pristis Val. "1827
Bory St. Vincent: annotations to Bruguiere 1798. PI.
299 fig. 3 Cardium pristis Valenciennes. --1839 Anton:
Cardium pristis Val. = Cardium laevigatum Linne.
--1845 Reeve, sp. 69: Cardium laevigatum Linne =
Cardium pristis Valenciennes (1. Vidal pers. comm.).
--1846 Valenciennes, Voyage de la Venus: Cardium
pristis. --1864 Krebs: Cardium pristis Valene sic (syn.)
pristis
= Cardium laevigatum L. --1870 March :
Valenc. = C. laevigatum Reeve = C. var. citrinum
Chernnitz junior synonyms of Cardium laevigatum L.
"1900 Dall, p. 387,: Cardium pristis Valenciennes =
Cardium serratum Linne. --1902 Dall: Cardium pristis
Valenciennes, 1846 = Cardium serratum Linnaeus,
1758. --1977 Fischer-Piette: Cardium pristis
Valenciennes = species insufficiently known . CHARACTERS AND OCCURRENCE - REMARKS.
--1977 Fischer-Piette: no figure in Valenciennes 1846.
--2004 Vidal pers . comm .: Laevicardium pristis
(Valenciennes, 1827) is a good species. Summary of
ID's: American Atlantic species close to C. serratum
Linne, 1758 but larger and with different specific
characters. It has 3 synonyms C. sublineatum Conrad,
1841. C. glabratum Romer, 1869, and C. multilineatum
DaB & Simpson, 1901.
pristonum (error). - Printer's mistake for pristinum in
Popov 1977.
privum Ebersin. - Moquicardium privum (Ebersin) .
Lymnocardiidae. Fossil. RECORDS . --1958 ZR:
Limnocardium (Moquicardium) privum [Ref. Eberzin
1958]. --1987 Taktakishvili : Limnocardium
(Moquicardium) privum Ebers. (?). --1987
c.
Taktakishvi1i: Moquicardium privum. "1987
Taktakishvili: Moquicardium privum Ebers. (?).
privum oblonga Ebersin, 1958 (variety). - Upgraded.
Moquicardium privum oblonga (Ebersin, 1958). Fossil.
RECORDS. --1958 ZR: Limnocardium (Moquicardium)
privum var. oblonga Eberzin, 1958 nov. Abkhazia, river
Mokvi, east coast of Black Sea. --1987 Taktakishvili:
Limnocardium (Moquicardium) privum Ebers. var.
oblonga Ebers. --1987 Taktakishvili: Moquicardium
privum oblonga.
probatum Iredale, 1927. [Figs. Part 3 p. 900] . - Synonym
of Nemocardium bechei (Reeve, 1847). OTHER
JUDGEMENTS : Pratulum probatum. » Regarding
Pratulum probatum : Cardiidae. Recent. TYPE LOCALITY.
Australia . TYPE REPOSITORY. Australian Museum,
Sydney. Syntypes C028434 . DISTRIBUTION. Japan,
Australia (Hylleberg 1994). RECORDS. "1927 Iredale,
p. 333-334 (pI. 46, fig. 8): Pratulum probatum. --1927
ZR: Pratulum probatum Iredale, 1927 sp . n . E .
Australia. --1950 Allan: Pratulum probatum (somewhat
different from bechei Reeve) . "1950 Keen:
Nemocardium (Pratulum) probatum (Iredale, 1927).
"1951 Habe: Nemocardium probatum Rve. "1958
Powell : Nemocardium probatum Ired . --1961
Rippingale & MacMichae1: Pratulum probatum
Iredale. "1962 Iredale & MacMichae1: Cardium
probatum Ired . --1977 Fischer-Piette: Pratulum
probatum Iredale = Cardium bechei Reeve. --1977
Wilson & Stevenson: Pratulum probatum Iredale, 1927;
junior synonym of Nemocardium bechei (Reeve, 1840).
--1978 ZR: Pratulum probatum Iredale, 1927 New
synonym of Nemocardium bechei (Reeve, 1840). Ref.
to Wilson et al. p . 63. --1990 Abbott & Dance:
Nemocardium probatum (Iredale, 1927) = Cardium
bechei Reeve, 1847.
problematica. - See under problematicum Munster,
1840.
problematieum d'Orbigny, 1850. - Cardium
problematicum d'Orbigny, 1850. ?Cardiidae. Fossil.
RECORDS. --1922-32 Sherborn, p. 5156: Avicula
problematica Muenster, 1840. --1922-32 Sherborn, p.
5156: Cardium problematicum Muenster, 1842. "192232 Sherborn, p. 5156: Cardium (Muenster)
problematicum d'Orbigny, 1850. - Avicula.
*proboseidellm - Homonymy to be noticed. Cardium
proboscideum from 2 sources: Sowerby, 1817. Hoening, 1839.
proboscidellm Hoening, 1839. - Unknown species.»
Regarding Cardiumproboscideum: ?Cardiacea. Fossil.
RECORDS. --1857 Deshayes: Cardium proboscideum
Hoening, 1839 - spec. incog.
proboscideum Sowerby, 1815. - Granocardium
proboscideum (Sowerby, 1815). Cardiidae. Cretaceous.
RECORDS. "1815 Sowerby, Conch. no. 27. t. 156. fig.
1: Cardium proboscideum. --1819 Lamarck : Cardium
proboscideum close to Cardium echinatum. --1857
Deshayes: Cardium proboscideum Sowerby, 1815.
"1871 Stoliczka: Cardium proboscideum (Acanthocardium sic). "1899 Sacco: Cardium proboscideum
Sowerby [Cretaceous]. "1922-32 Sherborn, p. 5157:
Cardiumproboscideum Sowerby, 1817. --1931 ZR:
Cardium proboscideum Ref. to Rieder (1931). --1978
Scott: Granocardium proboscideum (Sowerby in
Woods, 1908). "1999 Jacques Vidal (pers. com.),
712
Tropical Marine Mollusc Programme (TMMP)
procarpatina - procumbens
MNHN, Paris: Fossils from the Paleontological
Collection as Cardium proboscideum d'Orbigny.
Albien-Cenom. France (Yonne, Nievu), S07059,
SOn81.
procarpatina Jekelius, 1944. - Replidacna procarpatina
Jekelius, 1944. Lymnocardiidae. Miocene. REcORDs.
··1946 ZR: Replidacna procarpatina lekelius, 1944 sp.
nov. Miocene Soceni Banat. ··1946 ZR: Replidacna
procarpatina Miocene [Ref. Jekelius 1944]. ··1977 ZR:
Cerastoderma (Obsoletiforma) procarpatinum
(Jekelius). Syn. nov. Cardium (Replidacna)
procarpatinum lekelius. Ref to Paramonova p. 70
[Trans!. p. 320].
procarpatina spinosa lekelius, 1944 (variety). Upgraded. Replidacna procarpatina spinosa lekelius,
1944. RECORDS. ··1946 ZR: Replidacna procarpatina
var. spinosa lekelius, 1944 nov. Miocene, Soceni Banat.
procera (mode). - See under procerum Sowerby, 1833.
procerum Sowerby in Broderip & Sowerby, 1833. [Figs.
Part 1 p. 110, 133, In. Part 3 p. 924]. - Changed
combination: Trachycardium procerum (Sowerby,
1833). Cardiidae. Pliocene and Recent. Type species:
Cardium procerum Sowerby, 1833 (DO) is type species
of subgenus Mexicardia Stewart, 1930 (Glibert & Poel
1970). TYPE LOCALlTY. In America Centrali (Real Llejos,
near Corinto, Nicaragua) in coarse sand from 4 to 6
fathoms (Sowerby 1833). TYPE REPOSITORY. BMNH;
Cardium procerum Sowerby. Three syntypes. reg. no.
1989118. Neg. No. 0630. RECORDS. ··1833 S0t'erby,
p. 83: Cardium procerum. ··1834 Sowerby, 50 t part,
fig. 23: Cardium procerum nobis. ··1840 Sowerby, p.
5, No. 61: Cardium procerum Sow. ··1840 Sowerby,
corrected list, p. 8: Cardium procerum (Sowerby No.
61). ··1844 Reeve, sp. 51: Cardium procerum Sowerby,
1833. Pro. Zoo I. Soc. ··1846 Orbigny: Cardium
procerum Sowerby, 1833. ··1852 Adams: 433. Cardium
procerum Sowerby, 1833. ··1858 Adams & Adams, p.
455: Cardium (Trachycardium)procerum, Sow. ··1860
Carpenter: Cardium procerum Sby. ··1868 ZR:
Cardium procerum (Sow.), a variety of it is laticostatum
(Sow.) [Ref. Romer 1868]. ··1890 Paetel: Cardium
procerum Sow. ··1895 Steams: Cardium procerum
Sowerby. ··190 I Dall: Cardium (Ringicardium)
procerum Sowerby, 1833 [Pleistocene]. ··1909 Lamy:
Cardium (Ringicardium) procerum Sowerby. ··192232 Sherborn, p. 5160: Cardium procerum Sowerby,
1833. ··1930 Stewart designated Cardium procerum
Sowerby, 1833 type species of the subgenus
(Mexicardia) Stewart, 1930. ··1932 Pilsbry & Lowe:
Cardium (Bingicardium sic)procerum Brod. and Sby.
··1934 Hertlein: Cardium procerum Sowerby [fossil].
··1941 Pilsbry & Olsson: Cardium (Mexicardia)
procerum Sby. [Pliocene]. ··1947 Hertlein & Strong:
Cardium (Mexicardia) procerum Sowerby. ··1959
Ranson, p. 71: Cardium procerum Sowerby. ··1964
Parker: Trachycardium (Mexicardia) procerum
(Sowerby, 1833). ··1969 Moore et at.: Trachycardium
(Mexicardia) procerum (Sowerby, 1833). ··1969 ZR:
Trachycardium procerum range extension, Peru [Ref.
Vegas Velez 1969]. ··1970 G libert & Poel:
Trachycardium (Mexicardia) procerum (Sowerby,
1833). ··1971 Keen & McLean: Trachycardium
(Mexicardia) procerum (Sowerby, 1833). ··1977
Fischer-Piette: Laevicardium (Mexicardia) procerum
Sowerby. ··1977 Popov: Cardium procerum Sowerby,
1833. Type of Mexicardia Stewart, 1930. ··1977 Wilson
& Stevenson: Cardium procerum Sowerby, 1833.
··1995 Poutiers: Trachycardium procerum (Sowerby,
1833). ··1999 Vidal: Trachycardium (Mexicardia)
procerum (Sowerby in Broderip & Sowerby, 1833).
··200 I Schneider & Carter: Trachycardium procerum.
··2001 Vidal, p. 407: Trachycardium procerum. CHARACTERS AND OCCURRENCE - CHARACTERS
of diagnostic value: shell decidedly higher than long;
umbo rounded or roundly angled posteriorly; posterior
area not set off with strong concentric sculpture; very
convex; ribs without scales. OF COLLECTIONS. ··ZMUC:
I specimen from Bai de la Magdalaine, Venus
Expedition, 1839. ··ZMUC Gulf of California.
DESCRIPTIONS. ··1844 Reeve: shell oblong-ovate,
swollen, anterior side rounded, posterior somewhat
squarely angulated, serrated, radiately ribbed, 25
flattened ribs, arcuately striated, the 4 extreme anterior
ribs rather obsolete, the rest posteriorly angulated;
brownish-white, ribs faintly stained with a very few
purple-brown spots, covered with a thin earthy
epidermis, posterior margins purple-brown within.
··1947 Hertlein & Strong: shell elongately trigonal,
somewhat oblique posteriorly, 22-25 smooth, flattishtopped radial ribs, these are not strongly developed on
the anterior slope of the shel.1 but the posterior margin
is serrated due to projecting ends of the ribs which
interlock with the opposite valve. Shells < 35 mm in
length are longer in proportion to the height. Shells
about 10-15 mm in length have rounded ribs which
gradually become broader, subtriangular and flatter and
are separated by narrow and fairly deep interspaces.
Colour externally brownish with a tendency toward
yellowish concentric areas. ··2001 Vidal: tentacles
numerous. No eyes on tentacles. REMARKS. ··1971
Keen: some authors have placed this species in the
subgenus Ringicardium, but that group is essentially
European and African in distribution. T procerum is
of moderate size (about 2 inches in height), with
smooth, rounded ribs that are separated by linear
interspaces. The shell is only moderately thick and the
color is white, extensively mottled and spotted with
brown. Although a few specimens have been found as
far north as the Gulf of California, the geographic range
seems mostly to be southward from Oaxaca, Mexico,
to Chile; not uncommon. DIMENSIONS. ··1947 Hertlein
& Strong: height more than 80 mm. REMARKS. ··1947
Hertlein & Strong: The rather considerable variation
in shape during growth has led to description of this
species under several different names: Cardium
laticostatum Sowerby = a young stage of C. procerum.
HABrrAT. ··1844 Reeve: found in coarse sand at a depth
of from 4 to 6 fathoms. ··ZMUC: sand at 22-70 m
depth. DISTRIBUTION. ··1844 Reeve: Real Llejos, Central
America. ··1884 Grasset: Panama. ··1971 Keen &
McLean: the Gulf of California, South West Mexico to
Chile. ··ZMUC: Gulf of California.
procrescens Munster, 1840. - Lunulacardium
procrescens Muenster, 1840. Not in Cardiidae. Fossil.
RECORDS. ··1922-32 Sherborn, p. 5161: Lunulacardium
procrescens Muenster, 1840.
procumbens - Not valid: Conocardium procumbens.
RECORDS. ··1922-32 Sherborn, p. 5162: Conocardium
Phuket Marine Biological Center Special Publication vol. 30 (2004)
713
productum - profundum
procumbens without authorship In. n.].
productum (variety). - See under paucicostatum
productum Bucquoy, Dautzenberg & Dollfus, 1892.
*productum. - Homonymy to be noticed. Cardium
productum from 3 sources: Sowerby, 1832. - Deshayes,
1854. - Orbigny (?year).
productum d'Orbigny. - Granocardium productum
(d ' Orbigny). Cardiidae. Cretaceous. RECORDS. "1999
Jacques Vidal (pers. com.), MNHN, Paris: Fossils from
the Paleontological Collection as Cardium.
Granocardium. productum d ' Orbigny. France; Austria,
S10015, S10016. Cameroun, R51365.
productum (aff.) - Cardiidae. Cenomanian. REcoRDs.
"1965 ZR: Cardium aff. productum synonymy,
Cenomanian, Egypt [Ref. Fawzi 1965].
productum Deshayes, 1854. [Figs. Part 3 p. 915]. ?Synonym of Corculum cardissa (Linne, 1758). OTHER
JUDGEMENTS: Corculum dionaeum (Sowerby) 1855.»
Regarding Cardium productum: Cardiidae. Recent.
TYPE LOCALITY. Torres Strait. TYPE REPOSITORY. 2
syntypes. BMNH. REcoRDs. "1855 Deshayes: Cardium
productum. "1890 Paetel: Hemicardium productum
Dsh. "1903 Hidalgo: Cardium productum Deshayes,
1854 sic; doubtful occurrence in the Philippines. "1913
ZR: Cardiumproductum Desh. fig. [Ref. Hedley 1913].
"1914 Hedley, p. 268 (pI. 16 figs. 20-22): Cardium
productum Deshayes = Cardium dionaeum Sowerby.
"1928 Faustino: Cardium (Hemicardium) productum
Deshayes. "1977 Fischer-Piette: Cardium productum
Deshayes = Cardium cardissa Linne. "1980 Ruhoff:
Cardium productum Deshayes, 1854; p. 333 . CHARACTERS AND OCCURRENCE - DIMENSIONS.
"1914 Hedley: 15 mm long 23 mm high, and 20 mrn
across the conjoined valves. REMARKS. "1914 Hedley
p. 268 (pI. 16 figs. 20-22): the accompanying figure is
taken from a Cumingian specimen, probably the type.
DISTRJBUTION ... Of authors: Indo-West Pacific.
productum J.Sowerby, 1832. - Granocardium
productum (Sowerby, 1832). Cardiidae. Upper
Cretaceous. Cardium productum Sowerby, 1832 is type
species of subgenus Granocardium Gabb. REMARKS.
Cardium productum (Trachycardium) identical with
European species (Stoliczka 1871). According to
Sherborn (1922-32) there is no description, but it is
figured [n. etf]. - RECORDS. "1832 Sowerby: Cardium
productum. "1857 Deshayes: Cardium productum
Sowerby. "1871 Stoliczka: Cardium productum
(Acanthocardium sic). "1871 Stoliczka: Cardium
productum Sowerby, 1832 . type of subgenus
Granocardium Gabb. "1887 Fischer: Cardium
productum Sowerby. "1889 ZR : Granocardium
productum with fig . (ref. Holzapfel) . "1922-32
Sherborn, p. 5164: Cardium productum Sowerby, 1832
[n. etf]. "1933 ZR: Cardium productum [Ref. Parona
1933]. "1935 ZR: Granocardium productum Sow. fig.
[Ref. Bakalov & Tzankov 1935] . "1963 ZR : Cardium
productum Cretaceous [Ref. Bobkova \963) . "1986
Abdel-Gawad, pI. 40, figs . 6,7 : Grano cardium
(Criocardium) productum (Sowerby, 1832). "1994 E1Nakhal & EI-Naggar: Cardium productum . "1998
Schneider: Cardium productum Sowerby, 1832.
Transferred to genus Granocardium Gabb, 1869.
productum daganakiikensis Bobkova, 1963 (variety).
- Infrasubspecific. » Regarding Cardium productum.
var. daganakiik.ensis: Cardiidae. Cretaceous. RECORDS.
"1963 ZR: Cardium productum var. daganakiikensis
Bobkova, 1963 nov. Cretaceous, Tadzhik depression,
USSR.
productum subtriangulare Cottreau, 1924. - Upgraded.
"Cardium " productum subtriangulare Sow. Cardiidae.
Cretaceous. RECORDS. "1924 ZR: Cardium productum
Sow. subtriangulare Cottreau, 1924 mut. n.
proechinatum Hilber. - Combination to be revised:
"Cardium" proechinatum Hilber. Cardiidae. Miocene.
RECORDS. "1899 Sacco: Cardium proechinatum Hilber
[Miocene).
Profraginae Badve, 1977 (subfamily). - RECORDS.
"2001 Schneider & Carter: Profraginae (subfamily)
Badve, 1977. "2002 Schneider: Criocardium ,
Ethmocardium, and Granocardium has been transferred
to Profraginae.
Profulvia Kafanov, 1976 (genus) . - Cardiidae. Fossil.
Subgenus (Profulvia) Kafanov, 1976 upgraded to
generic level by authors. RECORDS. "1976 ZR: Profulvia
gen. nov. of Cardiidae Type species Papyridea
harrimani Dal\, 1904. Ref. to Kafanov, p. 482 [org. p
III J. "1977 ZR: Profulvia Kafanov 1976. Designated
type genus for new tribe Profulviini. Ref. to Kafanov
& Popov p. 62 [Trans!. p. 312). "1980 Vokes: Profulvia
Kafanov, 1976. Fossil. "1980 ZR: Profulvia Kafanov
1976. Key to species. Ref. to Kafanov. "1992 Amano
& Tanaka: Profulvia Kafanov, 1976.
Profulvia Kafanov, 1976 (subgenus). - Cardiidae. Fossil.
Type species by original designation: Papyridea
harrimani Dall, 1904. RECORDS. "1976 Kafanov:
Subgenus Profitlvia Kafanov, 1976. - CHARACTERS
AND OCCURRENCE - DESCRIPTIONS. Shell elongateovate, truncated, variable inequilateral, moderately
convex, with gape at posterior or both ends. Beaks
relatively low, obtuse, nearly orthogyrate, opistogyrate
or weakly prosogyrate. Ribs about 30-65 in number;
ribs straight, narrow and low on the anterior valve
surfaces, more curved posteriorly. Ridges of ribs with
numerous commarginallines and wrinkles; weak nodes
occur where growth lines cross costal crests. Paired
cardinal teeth smal.l and straight in both valves; lateral
teeth usual.ly single.
Profulviini Kafanov in Kafanov and Popov, 1977
(tribus). - Cardiidae. Fossil. RECORDS. "1977 Kafanov:
Profulviini. "1999 Kafanov: Kafanov in Kafanov and
Popov, 1977. "1977 ZR: Profulviini Tribe nov. of
Cardiidae, type genus Produlvia Kafanov 1976.
Tertiary. Ref. to Kafanov & Popov p. 62. [Transl. p.
312] .
profundicola Logvinenko & Starobogatov, 1966. Didacna profundicola Logvinenko & Starobogatov,
1966. Lymnocardiidae. Recent. TYPE LOCALITY. Caspian
Sea. RECORDS. "1968 Logvinenko & Starobogatov:
Didacna profimdicola Logvinenko & Starobogatov,
1966.
profundum Vidal, 1999 . [Figs . Part 3 p. 878) . Acrosterigma profundum Vidal, 1999. Cardiidae .
Recent. TYPE LocALrrv. Holotype, New Caledonia, 245268m. TYPE REPOSITORY. Holotype and paratypes :
Museum national d 'Histoire naturelle, Paris. Paratypes,
National Museum of Natural History, Washington DC.
TYPE MATERIAL. Holotype: A right valve. (MNHN,
Bouchet, Metivier, Richer de Forges). Paratype 1: A
714
Tropical Marine Mollusc Programme (TMMP)
promethei - Prophyllicardium
left valve, New Caledonia, 105-110 m (MNHN, idem).
Celidje, p. 157: Chartadacna promethei gen. et sp. nov.
Paratype 2: A left valve, north of New Caledonia, 315prominens Popov & Nevesskaya, 2000. - Lymnocardium
350m (MNHN idem). Paratype 3: A left valve, broken
prominens Popov & Nevesskaya, 2000. Cardiidae. Late
Miocene. RECORDS. ··2000 Popov & Nevesskaya,
and incomplete, east coast of New Caledonia, 227250m (MNHN, idem). Para type 4: A very small left
downloaded abstract: Lymnocardium (Euxinicardium)
valve, from same locality as paratype 1. Paratype 5: A
prominens n. sp.
promultistriata lekelius, 1944. - Changed combination:
right valve, seaward (N.) of west end of Bikini Atoll,
Pontalmyra promultistriata (Jekel ius, 1944).
Marshall Islands, 177-244m (USNM 598697, R. W.
Russell Coil) . Paratype 6: A left valve, same lot as
Lymnocardiidae. Miocene. ··1946 ZR: Limnocardium
paratype 5. Paratype 7: A left valve, same lot as
promultistriatum Jekelius, 1944 sp. nov. Miocene
paratypes 5 and 6. RECORDS. ··1999 Vidal, p. 296, Table
Soceni Banat. RECORDS. ··1987 Taktakishvili: Didacna
promultistriata lekelius. ··1987 Taktakishvili:
20, Figs. 71-M, 8C: Acrosterigma profundum n. sp. Pontalmyra promultistriata (Jek.).
CHARACTERS AND OCCURRENCE - CHARACTERS
propebanneianum Dietrich, 1933. - Tendagurium
of diagnostic value: very elaborate and specific rib
morphology that cannot be confused with any other
propebanneianum (Dietrich, 1933). Cardiidae.
Cretaceous. RECORDS. ··1933 ZR: Cardium
cardiid. DESCRIPTIONS. "1999 Vidal: shell ovoid and
nearly equilateral with no posterior truncation,
propebanneianum Dietrich, 1933 sp. n. Type of
moderately elongated and globose. Lunule small, rather .
Tendagurium Dietrich, 1933 subgen. n. of Cardium.
Cretaceous Africa. ··1937 ZR: Cardium (Tendagurium)
imperfectly delineated and nearly the same on both
propebanneianum Dietrich 1933 type of Tendagurium
valves; umbonal margin raised, forming a double wall.
[Ref. Keen 1937]. ··1995 Schneider: Tendagurium
External colour pattern is unique: on adult part,
posteriorly entirely pink to orange, middle surface bears
Dietrich, 1933. Mesozoic subgenus of Cardium.
Salisbury
(1934)
designated
Cardium
large, pink to orange, irregularly concentric splashes,
propebanneainum (sic = propebanneianum) Dietrich,
and anterior half of the shell beige, with widely
scattered small splashes. Interior white with some
1933, as the type species. ··1995 Schneider:
Tendagurium propebanneianum (Dietrich, 1933).
exterior splashes showing through; no umbonal rays
propeciliare Dall, 1898. - Changed combination:
visible. Hinge symmetrical and moderately angled. On
right valve, cardinals show an unusual degree of fusion
Acanthocardia propeci/iare (Dall, 1898). Cardiidae.
at their base. Mean rib number 56.9 ± 3.1, range 55Fossil. RECORDS. "1898 Dall : Cardium propeciliare
63. Posteriorly, interstices thin; ribs flat with a central
Dall, 1898 n. sp. ··1977 Vokes Acanthocardia
propeciliare (Dall).
longitudinal furrow, anterior smooth part as wide as or
narrower than posterior. Elongated, thin, straight,
propeleve Ebersin, 1958. - Changed combination:
Euxinicardium propeleve (Ebersin, 1958).
oblique scales develop on posterior part, straddling
axial furrow and encroaching slightly upon anterior
Lymnocardiidae. Fossil. RECORDS. ··1958 ZR:
Limnocardium (Euxinicardium) propeleve Eberzin,
zone. On posterior half of middle surface, the same
features appear, but posterior zone slightly raised and
1958 sp. nov. Kerchenski Peninsula. Crimea; Kuban
Province. Pliocene of the Ponto-Caspian basin, Russia.
ribs become flatly triangular; longitudinal furrow
··1987 Taktakishvili : Limnocardium (Euxinicardium)
progressively disappears and anterior smooth zone
becomes anterior flank of the rib. On anterior half of
propeleve Ebers. ··1987 Taktakishvi Ii: Euxinicardium
middle surface, ribs become more rounded, scales
propeleve.
become progressively shorter and more numerous and
propeleve Ebersin (variety). - Infrasubspecific. »
Regarding Euxinicardium propeleve var:
change into posterior ridges, rib tops becoming smooth;
Lymnocardiidae. Fossil. RECORDS. ··1987 Taktakishvili:
at same time, numerous very thin oblique serrations
Euxinicardium propeleve Ebers. var.
appear on anterior flanks of ribs, changing anteriorly
propeleve inleve Ebersin (subspecies). - Euxinicardium
into small ridges or tubercles. Anteriorly, on middle
propeleve inleve. Lymnocardiidae. Fossil. RECORDS.
surface, this sculptural evolution leads to ribs which
are slightly square-sided, serrated on both sides, and
··1987 Taktakishvili: Euxinicardium propeleve inleve.
propesemicostatllm Gregorio, 1930. - Combination to
slightly overhanging interstices. Anteriorly, some lateral
be revised: "Cardium" propesemicostatum Gregorio,
serrations join on top of ribs, forming thin, free topridges, which are not imbricated; the first ribs of this
1930. Cardiidae. Lias. RECORDS. ··1936 ZR: Cardium
quarter slightly degenerate, their ornamentation
propesemicostatum Gregorio, 1930 sp. n. Lias Palermo
[Ref. Gregorio 1930].
becoming less regular. DIMENSIONS. ··1999 Vidal: length
28.2, height 36.7, width (calculated) 23.6 mm .
propexllm Monterosato, 1891. - Synonym of
REMARKS. "1999 Vidal: Acrosterigma profundum is
Acanthocardia echinata (Linne, 1758). » Regarding
separated from A. uniornatum by its larger size, higher
Cardium propexum: Cardiidae. Recent. RECORDS.
rib number, and particular rib ornamentation on the
"1891 Monterosato: Cardium propexum. "1892
median part of the shell, i.e. the small dense serrations
B.D.D. Cardium propexum = Cardium echinatum .
on the anterior flanks of the ribs. It cannot be confused
"1912 Dautzenberg & Fischer: Cardium propexum
with other species. DISTRJBUTION. North and east of New
Monterosato 1891 = Cardium (Acanthocardia)
Caledonia, at relatively great water depths for Cardiidae
echinatum, Linne.
other than Protocardiinae; present also in Bikini Atoll,
Prophyl/icardillm lekelius, 1944 (genus).
Marshall Islands.
Lymnocardiidae . Pliocene. RECORDS. ··1946 ZR :
promethei Celidje, 1953. - Chartadacna promethei
Prophyllicardium lekelius, 1944 gen. nov. Cardiidae,
Celidje, 1953. Lymnocardiidae. Fossil. RECORDS. ··1953
type Prophyllicardium soceni lekelius, 1944 sp. nov.
Phuicet Marine Biological Center Special Publication vol. 30 (2004)
715
Prophyllicardium - Prosodacnidae
Pliocene Rumania (Lower Congerien) Soceni. Banat.
Prophyllicardium lekelius, 1944 (genus). Lymnocardiidae. Tertiary. RECORDS. 001980 Vokes:
Prophyllicardium Jekelius, 1944. Subfamily uncertain.
propinqua Eichwald, 1841. [Figs. Part 1 p. 123] . Changed combination: Apscheronia propinqua
(Eichwald, 1841) . Lymnocardiidae. Tertiary. TYPE
LOCALITY. Caspian Sea. REcORDs. 001841 Eichwald, p.
220, pI. 40, figs. 3-4: Monodacna propinqua. 001866
Issei: Monodacna propinqua, Eichwald, 1842. 001903
Andrussoff: Monodacna propinqua Eichwald, 1841 .
001922-32 Sherborn, p. 5173: Monodacna propinqua
Eichwald, 1841. 0'1930 ZR: Apscheronia propinqua
[Ref. Androussov 1923]. 001977 Popov: Apscheronia
propinqua (Eichw.). - CHARACTERS AND OCCURRENCE - DESCRIPTION. 001903 Andrussoff: Monodacna
propinqua Eichwald has a smooth shell surface. See
also under the proposed replacement name: Cardium
proximum Deshayes.
propinqua Reed, 1944. - Plethocardia propinqua Reed,
1944. Cardiacea. Ordovician. RECORDS. 001944 ZR:
Plethocardia propinqua Reed, 1944 sp. nov. Balclatchie
Beds, Balclatchie, Girvan (Ordovician).
propinqua brevior Androussov, 1923 (variety). Upgraded. Apscheronia propinqua brevior Androussov,
1923. Lymnocardiidae. Tertiary. RECORDS. 001930 ZR:
Apscheronia propinqua var. brevior Androussov, 1923
nov. [Ref. Androussov 1923].
propinqua mactriformis Androussov, 1923 (variety). Upgraded. Apscheronia propinqua mactrijormis
Androussov, 1923. Lymnocardiidae. Tertiary. REcORDs.
001930 ZR: Apscheronia prop in qua var. mactriformis
Androussov, 1923 nov. [Ref. Androussov 1923].
propinqua oblonga Androussov, 1923 (variety). Upgraded. Apscheronia propinqua oblonga
Androussov, 1923. Lymnocardiidae. Tertiary. REcORDs.
001930 ZR: Apscheronia propinqua var. oblonga
Androussov, 1923 nov. [Ref. Androussov 1923].
propinqua praeextensa Androussov, 1923 (variety). Upgraded. Apscheronia propinqua praeextensa
Androussov, 1923. Lymnocardiidae. Tertiary. REcORDs.
001930 ZR: Apscheronia propinqua var. praeextensa
Androussov, 1923 nov. [Ref. Androussov 1923] .
propinqua typica Androussov, 1923 (variety). Upgraded. Apscheronia propinqua typica Androussov,
1923. Lymnocardiidae. Tertiary. RECORDS. 001930 ZR:
Apscheronia propinqua var. typica Androussov, 1923
nov. [Ref. Androussov 1923].
propinquum (mode). - See also under propinqua
Eichwald, 1841 .
propinquum. - Homonymy. Cardium propinquum from
3 sources: Munster in Goldfuss, 1837. Munster, 1839.
Eichwald, 1841.
propinquum Eichwald, 1841. - Replaced by Cardium
proximum Deshayes. Cardiidae. Fossil. RECORDS.
001857 Deshayes: Cardium propinquum Eichwald,
1841 - [C.] proximum Desh.
propinquwn Munster in Goldfuss, 1837. - Replaced by
"Cardium " bavaricum Deshayes. » Regarding
Cardium propinquum: Cardiacea. Cretaceous. REcORDs.
001857 Deshayes: Cardium propinquum Munster, 1840
- [C] bavaricum Desh. 001922-32 Sherborn, p. 5172:
Cardium propinquum Muenster in Goldfuss, 1837.
propinquum Munster, 1839 . - Combination to be
revised: "Cardium " propinquum Munster, 1839.
Cardiidae. Cretaceous. RECORDS. 001857 Deshayes:
Cardium propinquum Munster, 1839 .• 01871 Stoliczka:
Cardium propinquum (Cerasioderma) .
propinquum Orbigny, 1850. - Conocardium propinquum
d'Orbigny, 1850. Conocardiidae. Fossil. RECORDS.
001922-32 Sherborn, p. 5172: Conocardium (Muenst.)
propinquum d'Orbigny, 1850. - Cardium.
propria (subspecies). - See under corbuloides propria
Taktakishvili, 1978
proprium - Cerastoderma proprium . Cardiidae. Fossil.
REcORDs. 001977 Popov: Cerastoderma proprium .
proprium Alizade. - Acobaecardium proprium (AI izade).
Lymnocardiidae . Neogene. RECORDS. ··1986
Nevesskaya et al.: Acobaecardium proprium (Alizade)
comb. nov . • 01987 ZR: Acobaecardium proprium
(Alizade) comb. nov. Ref. to Nevesskaya et al. p. 192.
Prosadacna (error). - Printer's mistake for Prosodacna
in Fischer (Vokes 1980).
Prosochiasta Ebersin, 1959 (genus). - Lymnocardiidae.
Tertiary. REcORDs. 001974 ZR: Prosochiasta Ebersin,
1959. Referred to new subfamily Prosodacninae, Ref.
to Andreescu, p. 128 . 001980 Vokes : Prosochiasla
Ebersin, 1959 in subfamily Lymnocardiinae.
Prosochiasta Eberzin , 1960 (subgenus).
Lymnocardiidae. Pliocene. RECORDS. 001960 ZR:
(Proso c hiasta) Eberzin, 1960 subgen. nov. of
Prosodacna. SUbgenotype: Prosodacna prosochiasla
Andrus., Pliocene Galizgi river, Abkhaziya, Georgia
USSR. 001960 ZR: Prosochiasla Eberzin, 1960 subgen.
nov. of Prosodacna q.v.
prosochiasta Andrus. - Prosodacna prosochiasla
Andrus. Lymnocardiidae. Pliocene. REcORDs. 001960
ZR: Prosodacna prosochiasta Andrus., Pliocene
Galizgi river, Abkhaziya, Georgia USSR. [Ref. Eberzin
1960].
Prosodacna Toumouer, 1882 (genus). - Lymnocardiidae.
Pliocene. GENOTYPE. Cardium macrodon Deshayes,
1838. TYPE LOCALITY. Crimean fossil. REMARKS. Divided
into 3 sections by Dollfus 1905, commenting
Andrussow 1903; (I) type species Prosodacna
semisulcala Rouss. (II) type species P. rumana Sabba,
(III) type species P cobalcescui Fontannes. RECORDS.
0'1903 Andrussoff: Prosodacna Tournouer, 1882.
001905 Dollfus, commenting Andrussow 1903:
Prosodacna Tournouer. 001974 ZR: Prosodacna
Tournouer, 1882. Designated type genus for new
subfamily Prosodacninae. Ref. to Andreescu, p. 130.
0'1975 ZR: Prosodacna (Psilodon) Cobalcescu, 1883.
Diagnostic characters. Palaeoecology. Systematic
accounts of species. Pliocene. Romania. Ref. to Motas.
001977 Popov: Prosodacna Toumouer, 1882. 001977
ZR: Prosodacna Systematic accounts of species. Ref.
to Papaianopol, I. p. 25. 001980 Vokes: Prosodacna
Tournouer, 1882. 001980 ZR: Prosodacna Toumouer
1882. Key to species. Ref. to Andreescu. 001987 Anaya:
Prosodacna Tourquer, 1882. 002001 Schneider &
Carter: Prosodacna (genus). Lymnocardiidae. Pliocene.
Prosodacna Tournouer, 1882 (subgenus) . Lymnocardiidae. Tertiary. RECORDS. 001887 Fischer:
Subgenus (Prosodacna) Tournouer, 1882. 001987
Anaya: (Prosodacna) S.s.
Prosodacnidae - RECORDS. 001937 ZR: Prosodacnidae
(?) not a cardiid unit [Ref. Keen 1937].
716
Tropical Marine Mol/usc Programme (TMMP)
prosodacnoides - Protocardiinae
prosodacnoides Andreescu, 1972. - Euxinicardium
prosodacnoides (Andreescu, 1972). Lymnocardiidae.
Fossil. R ECORDS. "1972 ZR: Limnocardium
(Euxinocardium sic) prosodacnoides Andreescu, 1972
sp. nov . Romania. Pleistocene . "1975 ZR:
Limnocardium (Euxinicardium) prosodacnoides
Andreescu, 1975 sp. nov. Romania, Vrancea district.
Pleistocene . Ref. to Andreescu , p . 112 . "1987
Taktakishvili: Euxinicardium prosodacnoides
(Andreescu).
Prosodacnomya Eberzin, 1960 (genus). Lymnocardiidae. Upper Miocene !Pliocene. RECORDS. "1974
ZR: Prosodacnomya Ebersin, 1959. Referred to new
subfamily Prosodacninae. Ref. to Andreescu, p. 128.
"1980 Vokes: Prosodacnomya Ebersin, 1959 in
subfamily ProsodacninaeAndreescu, 1974. "1980 ZR:
Prosodacnomya Ebersin 1959. Key to species. Ref. to
Andreescu. "1984 ZR: Prosodacna (Prosodacnomya)
Upper Miocene. Dacian Basin. Ref. to Papaianopol.
Prosodacnomya Eberzin , 1960 (subgenus) .
Lymnocardiidae. Upper Miocene !Pliocene. RECORDS.
"1960 ZR: (Prosodacnomya) Eberzin, 1960 subgen .
nov. of Prosodacna. SUbgenotype: Prosodacna rostrata
(Sinz.) Pliocene Ukraine, Odessa USSR. "1978 ZR:
Prosodacna (Prosodacnomya) Upper Miocene
Romania, Boteni. List & stratigraphy. Ref. to
Pavnotescu e/ al.
prosodactoides Andreescu, 1971. - New combination:
Lymnocardium prosodactoides (Andreescu, 1971).
Lymnocardiidae. Pleistocene. RECORDS. "1971 ZR:
Limnocardium (Euxinicardium) prosodac/oides
Andreescu, 1971 sp. nov. Pleistocene Rimnicu- Sarat
Valley, Dumitresti, Romania.
prosogyrata Kafanov, 1994 - Profulvia prosogyrata
Kafanov, 1994. Cardiidae. Fossil. RECORDS. "1994
Kafanov in Arkhipova et aZ.: Profulvia prosogyrata
Kafanov.
prosogyrata Perkins, 1961. - Protocardia prosogyrata
Perkins, 1961. Cardiidae. Cretaceous. RECORDS. "1961
ZR: Prolocardia prosogyrata Perkins, 1961 sp. nov.
Cretaceous, Texas, Fort Worth.
prosogyrata Perkins, 1961. - Protocardia prosogyrata
Perkins, 1961 . Cardiidae. Cretaceous. RECORDS. "1986
Scott: Protocardia prosogyrata Perkins, 1961.
protei Brongniart, 1821. - Pholadomya protei
(Brongniart, 1821). Lymnocardiidae. Fossil. RECORDS.
"1857 Deshayes: Cardium protei Brongniart. In genus
Pholadomya. "1922-32 Sherborn, p. 5177: Cardium
protei Brongniart, 1821.
proterothetidis Ludbrook, 1955. Nomen nudum.
RECORDS. Cardiidae. Pliocene. "1955 ZR:
Nemocardium (Pratulum) pro/erothe/idis Ludbrook,
1954 sp. nov. nom. nud. Pliocene, Australia. "1977
Popov: Pra/ulum pro/ero/he/idis (Ludbrook, 1955).
proteua (error). - Printer 's mistake for proteus in Ma
Qi-hong e/ al. 1976.
proteus Krumbeck, 1914 . - Protocardia proteus
Krumbeck, 1914. Cardiidae. Trias. RECORDS. "1914
ZR: Protocardia proteus Krumbeck, 1914 sp. n. Trias
Buru. "1976 Ma Qi-hong et al.: Protocardia proteua
sic Krumbeck, 1913.
protextum Conrad, 1853. - Combination to be revised:
"Cardium" pro/extum Conrad, 1853. Cardiidae. Fossil.
RECORDS. "1980 Ruhoff: Cardium pro/ex/um Conrad,
1853; vol. 2: 275, pI. 24, fig . 12.
protoaliculum Richards in Richards & Palmer, 1953. Sawkinsia protoaliculum (Richards in Richards &
Palmer, 1953). Cardiidae. Eocene. RECORDS. "1998
Schneider: Cardium (Trigoniocardia) protoaliculum
Richards in Richards & Palmer, 1953 in genus
Sawkinsia. "1998 Schneider: Cardium (Trigoniocardium sic) protoaliculum Richards, in Richards &
Palmer, 1953. Sawkinsia pr%aliculum (Richards, in
Richards & Palmer, 1953).
Protocardia Beyrich, 1845 (genus). - Cardiidae. Eocene
and Recent. GENOTYPE. Herrmannsen (1847) designated
Cardium hillanum Sowerby, 1818 type species of genus
Pr%cardia Beyrich, 1845 (Speden 1970). RECORDS.
"1845 Beyrich, designated the genus Pr%cardia.
"1857 Deshayes: Proto cardia Beyrich, in Bronn:syn.
of Cardium. "1876 Meek: Protocardia , Beyrich, 1845.
··1876 Meek: Protocardium = Pr%cardia. '·1886
Cossmann: GenusPr%cardium Beyrich, 1845 ... 1906
Suter: Protocardia Beyrich, 1845 . "1920 ZR:
Protocardia [Refs. Henning . Lange . Roman &
Mazeran). "1922 Olsson: Pro/acardia Beyrich. "192232 Sherborn, p. 5180: Protocardia Beyrich, 1845.
"1924 Oldroyd: Genus Pr%cardia Beyrich, 1845.
"1926 ZR: Protocardia [Ref. Schmidtill 1926]. "1928
ZR: Pr%cardia [vide etiam Protocardium] [Ref. Patte
1928]. "1929 ZR: Protocardia [vide etiam
Pr%cardium] [Refs. Cox. Rennie. Parona]. "1934
Johnson : Pr%cardia Beyrich 1845 . "1961 ZR :
Protocardia, all Tertiary species considered to belong
to Nemocardium Meek [Ref. Hughes 1961). "1980
Vokes : Pr% cardia Beyrich, 1845 in subfamily
Protocardiinae . •• I 980 ZR: Protocardia India Rann
Islands, Jurassic, first record. Ref. to Singh et al. "1986
Scott: Protocardia von Beyrich, 1845 (p. 1192).
Protocardia Conrad, 1860 (genus). Invalid. RECORDS.
"1980 Vokes: Protocardia Conrad, 1860 [non Beyrich,
1845].
Protocardia Beyrich, 1845 (subgenus). - Cardiidae.
Eocene and Recent. RECORDS. "1887 Fischer: Subgenus
Pr%cardia, Beyrich. 1845 of genus Cardium Linne,
1758.
Protocardiinae Keen, 1951 (subfamily). - Cardiidae.
Fossil and Recent. RECORDS. "1978 Maxwell:
Subfamily Protocardiinae. "1980 Vokes: Subfamily
Protocardiinae Keen, 1951. "1980 ZR: Protocardiinae
Keen 1951. Key to genera. Ref. to Keen. "1980 ZR:
Systematics. Generic composition & position. Ref. to
Andreescu.! Origin of taxon. Ref. to Andreescu. "1986
Abdel-Gawad: subfamily Protocardiinae Keen. 1951.
"1987 ZR: Protocardiinae. Lower Cretaceous. Mexico
& USA. Caribbean Province paJaeozoogeography.
Stratigraphic distribution, palaeobiology &
Systematics . Ref. 10 Scot!. ··1992 Poutiers :
Protocardiinae. "1995 Pacaud & Renard: Subfamily
Protocardiinae. ··1999 Higo et al.: Protocardiinae
(subfamily) Keen, 1951. "2001 Schneider & Carter:
Prolocardiinae (subfamily) Keen, 1951. CHARACTERS AND OCCURRENCE - DESCRIPTIONS. "1994 Hylleberg: rounded quadrate, beaks
nearly central; posterior slope well defined by umbonal
ridge, its radial ribs tending to be spinose; hinge long,
slightly arched, cardinals 2 and 3b larger, peglike,
curved upward, sinuating lower border of hinge plate;
Phukel Marine Biological Center Special Publication vol. 30 (2004)
717
Protocardium - proximum
pallial line mostly entire, some with small sinus near
posterior adductor scar.
Protocardium (error). - Invalid emendation of genus
Protoeardia. RECORDS. 001871 Stoliczka: Proto cardium
Beyrich, 1817. 001980. Vokes: Protoeardium (emend.
pro -cardia Beyrich, 1845).
Protocardum (error). - Printer's mistake for Protoeardia
in Meek & Hayden, 1860 (Vokes 1980).
Protocerdia (error). - Printer's mistake for Protoeardia
in Preston, 1921 (Vokes 1980).
Protodaena (genus). - RECORDS. 001968 Logvinenko &
Starobogatov: Genus Protodaena Logvinenko &
Starobogatov.
Protodacninae (subfamily) . - RECORDS. 001980 ZR:
Protodacninae. Origin of taxon. Ref. to Andreescu .
001980 ZR : Systematics. Generic composition &
position. Ref. to Andreescu.
Protodidaena Logvinenko & Starobogatov, 1966
(genus). - Lymnocardiidae. Tertiary and Recent.
RECORDS. 001980 Vokes: Protodidaena Logvinenko &
Starobogatov, 1966.
Protoplagiodaena (genus). - Lymnocardiidae. REcORDs.
001987 ZR: Protoplagiodaena Stevanovic, 1978.
Referred to new tribe Arcicardiini. Ref. to Nevesskaya
et al. p. 190.
protosllbrllgosllm Noetling, 190 I. - Vastieardium
protosubrugosum (N oetl ing, 190 I). Cardiidae.
Miocene. RECORDS. 001977 Popov: Traehyeardium
(Vastieardium) protosubrugosum (Noetling, 1901).
001999 Vidal: Cardium protosubrugosum Noetling,
190 I : 179, p I. 10, figs 10-11 Miocene of Burma .
Compared by Pannekoek (1936: 71) with spolongense
and sedanense.
protraeta (mode). - See under protraelum.
protraeta Eichwald, 1841. [Figs. Part I p. 123, 181]. Changed combination : Didaena protraeta (Eichwald,
1841) . OTHE R JUDGEMENTS: Monodaena ea tillus
Eichwald . » Regarding Cardium protraela:
Lymnocardiidae. Pliocene and Recent. TYPE LOCALITY.
Caspian Sea. RECORDS. 001841 Eichwald, p. 223, pI.
40, figs. 9-10: Adaena protraeta m. 001875 ZR: Adaena
protraeta (Eichw.), exhibits vestiges of cardinal teeth
[Ref. Martens 1875]. 001877 Grimm: Adaena protraeta
Eichwald (syn.) = Monodaena eatillus Eichwald.
001903 Andrussoff: Adaena protraeta Eichwald, 1841.
001922-32 Sherborn, p. 5182: Adaena protraeta
Eichwald, 1841. 001968 Logvinenko & Starobogatov:
Didaena protraeta protraeta (Eichwald) . 001977
Popov : Didaena prolraeta. 001987 Taktakishvili :
Limnoeardium protraetum (Eichw.). 001987
Taktakishvili : Didaena protraeta.
protraeta grimmi Andrusov (variety). - Changed
combination . Identical with Didaena protraeta
submedia Andrusov, 1911. » Regarding Didaena
protraela submedia: Lymnocardiidae. Fossil. REcoRDs.
001968 Logvinenko & Starobogatov: Didaena protraeta
var. grimmi Andrusov = Didaena protraeta submedia
Andr.
protraeta novaeaspiea G lazunova, 1971. - Didaena
protraeta novaeaspiea Glazunova, 1971. Lymnocardiidae. Pleistocene. RECORDS. 001971 ZR: Didaena
protraeta novaeaspiea Glazunova, 1971 subsp. nov.
Pleistocene Cape Peschanyy area, Middle Caspian,
USSR.
protraeta submedia Andrusov, 1911 (variety) . Upgraded . Didaena protraeta submedia Andrusov,
1911. Lymnocardiidae . Fossil. RECORDS. 001968
Logvinenko & Starobogatov: Didaena protraeta
submedia Andr. 001968 Logvinenko & Starobogatov:
Didaena protraeta var. submedia Andrusov, 1911 =
Didaena protraeta submedia Andr.
Protraetiforma Bagdassarian, 1978 (genus). - Cardiidae.
Fossil. RECORDS. 001979 ZR: Protraetiforma. Ref. to
Bagdasaryan, p. 44 . 001980 Vokes: Protraetiforma
Bagdassarian, 1978.
protraetiformis (variety). - See under quadripartitum
protraetiformis Andrus.
*protractum. - Homonymy to be noticed. Cardium
protraetum from 2 sources: Eichwald, 1829. - Dall,
1900.
protractum Eichwald , 1829 . - Combination to be
revised: "Cardium " protractum Eichwald, 1829. Cardiidae. Fossil. RECORDS. 001829 Eichwald, p. 283, Tab.
5, f. 9: Cardium protraetum n. 001857 Deshayes:
Cardium protraetum Eichwald. 001922-32 Sherborn,
p. 5182: Cardium protraetum Eichwald, 1829. 00 1999
Jacques Vidal (pers. com.), MNHN, Paris: Fossils from
the Paleontological Collection as Cardium protraetum
Eichwald. Greece, 837539. Turkey, B41471.
protraetus Dall, 1900. - Synonym of Serripes groenlandieus (Bruguiere, 1789). » Regarding Cardium
protraetus. Cardiidae: Recent. DISTRIBUTION. Arctic
(McLean 1939). RECOR DS. 001900 Dall : Cardium
protraetus. 001974 Abbott: Serripes protaetus Dall.
1900 = Serripes groenlandieus (Bruguiere, 1789).
protractus (subspecies). - See under groenlandieus
prolraetus Dall, 1900.
protrusum Solander, 1786 - Synonym of Coreulum
impressum (Lightfoot, 1786). OTH ER JUDGEMENTS:
Coreu/um eardissa (Linnaeus, 1758). » Regarding
Cardium protrusum: Cardiidae. Recent. RECORDS.
001687 Lister, pI. 319, fig . 156: pectunculus dorso sine
margine laevi. 001786 Solander, p. 178, ref. to Lister:
Cardium protrusum. 001817 Dillwyn: Cardium
protrusum. Solander's MSS, lot 3325 = Cardium
hum anum Chemnitz. 001916 Iredale, p. 86: Cardium
protrusum Solander. 001922-32 Sherborn, p. 5182:
Cardium protrusum Solander, 1786. 001967 Rehder:
Portland Catalogue No. 3825 . A very fine specimen of
Cardium protrusum , S. a curious variety of the Venus
heart, from China, very rare. Lister, 319. 156. 001967
Rehder : Cardium prolrusum (Lightfoot, 1786) =
Coreu/um impress um (Lightfoot, 1786). 001977
Fischer- Piette, p. 153 : Cardium protrusum So lander =
species insufficiently known. - CHARACTERS AND
OCCURRENCE-REMARKS. Lister (1687, pI. 319, fig .
156) is most likely Cardium humanum Chemnitz, 1782
= Coreulum eardissa (Linne, 1758).
*proximllm. - Homonymy to be noticed. Cardium
proximum from 2 sources: Deshayes, 1853 . - ? possibly
Dufour, 1881.
proximum Deshayes, 1853. - Combination to be revised:
"Cardium" proximum Deshayes. Cardiidae. Fossil.
RECORDS. 001857 Deshayes: Cardium proximum Desh.
001980 Ruhoff: Cardium proximum Deshayes, 1853 ;
vol. 2: 48 .
proximum Dufour, 1881. - Loxoeardillm proximum
(Dufour, 1881). Lymnocardiidae. Fossil. RECORDS.
718
Tropical Marine Mollusc Programme (TMMP)
proximum - Pseudocatillus
··1970 Glibert & Poel, p. 50: Loxocardium proximum
(Dufour, 1881).
proximum Fuchs. - Lymnocardium proximum (Fuchs).
Lymnocardiidae . Fossil. R ECORDS. ··1928 ZR:
Limnocardium proximum Fuchs Ponti an [Ref.
Pavlov itch 1928]. ··1929 ZR: Limnocardium proximum
Fuchs Pontian [Ref. Pavlovic 1929]. ··1987
Taktakishvili: Limnocardium proximum (Fuchs).
proximum bone; Pavlovitch, 1928 (variety). - Upgraded.
New combination: Lymnocardium proximum bonei
Pavlovitch, 1928. Lymnocardiidae. Pontian. RECORDS.
··1928 ZR: Limnocardium proximum Fuchs var. bonei
Pavlovitch, 1928 nov. Pontian Serbia [Ref. Pavlovitch
1928]. ··1929 ZR: Limnocardium proximum Fuchs var.
bouei Pavlovic, 1929 nov. Pontian Serbia [Ref. Pavlovic
1929].
prunum Meuschen, 1778. Non binominal. - Not
available. RECORDS. ··1902 Sherborn, p. 780: Cardium
prunum Meuschen, 1778. ··1902 Sherborn, p. 780:
Cardium prunus Meuschen, 1778.
pseudacardo Andrussow, 1907. - Arcicardium
pseudacardo Andrussow, 1907 . Lymnocardiidae .
Fossil. TYPE LOCALITY . Crimean iron ore deposits.
RECORDS. ··1907 Andrussow, p. 113, Taf. III, Fig. 5-9,
Taf. IV, Fig. 14: Arcicardium pseudacardo nov. sp.
··1964 ZR: Arcicardium pseudacardo Andrusov, 1964
sp. nov. Tertiary.
pseudoangulata (mode) . - See under pseudoangulatum.
pseudoangulatum BUlow, 1905 . - Synonym of
Vasticardium philippinense = Vasticardium orbita
phi/ippinense (Hedley, 1899). » Regarding Cardium
pseudoangulatum : Cardiidae. Recent. TYPE LOCALITY.
None designated (Higo el al. 1999). RECORDS. ··1905
BUlow : Cardium pseudoangulalum . ··1905 ZR:
Cardium (Trachycardium) pseudoangulatum BUlow,
1905b sp . n. Hab . incert. ··1991 Vidal : Cardium
pseudoangulatum Bulow, 1905 . ··1991 Voskuil &
Onverwagt: Trachycardium pseudoangulatum . ··1997
Vidal: Cardium (Trachycardium) pseudoangulatllm
BUlow, 1905 = Vasticardium orbita philippinense
(Hedley, 1899) . ··1999 Vidal : Cardium
pseudoangulatum BUlow, 1905 = Vasticardium orbita
(Broderip & Sowerby, 1833. ··1999 Higo et af. :
Acrosterigma (Vasticardium) pseudoangulata (BUlow,
1905). ··1999 Vidal , p. 357: C. pseudoangulatum
Bulow, 1905 identical with Va sticardium orbita
philippinense. - CHARACTERS AND OCCURRENCE - HABITAT. ··1999 Higo et af.: subtidal to 20
m, sand. DISTRJBUTION. ··1994 Hylleberg: Central IndoWest Pacific. ··1999 Higo et af. : Amami islands and
southwards, Philippines, South China Sea, China,
Indonesia, IndoPacific.
Pseudoarcicardium Akhvlediani, 1974 (genus). Lymnocardiidae. Fossil. RE CORDS. ··1974 ZR :
Pseudoarcicardium Akhvlediani, 1974 gen. nov. of
Cardiidae . Type species Arcicardium kubanicum
Andrusov. Ref. to Akhvlediani , p. 486. ··1980 Vokes:
Pseudoarcicardium Akhvlediani, 1974. ··1987
Taktakishvili, p. 92: Pseudoarcicardium Akhvlediani,
1974; referred to new tribe Pontalmyrini.
pselldoaxinus Thurmann in Thurmann & Etallon, 1862.
- Combination to be revised: "Cardium" pseudoaxinus
Thurmann in Thurmann & Etallon, 1862. Cardiidae.
Fossil. RECORDS. ··1980 Ruhoff: Cardium pseudoaxinus
Thurmann in Thurmann & Etallon, 1862; p. 182, pI.
21, fig Ie.
pseudobogatchevi (variety). - See under gracile
pselldobogatchevi Gillet, 1938.
Pseudo cardia Conrad, 1866 (genus).- Synonym. »
Regarding Pseudocardia: Lymnocardiidae. Fossil.
RECORDS. ··1887 Fischer: Pseudocardia pars Conrad,
1866. Jun. syn. of Genus Limnocardium Stoliczka, em.
1870. ··1903 Andrussoff: Pseudo cardia Conrad.
pseudo cardia (mode). - See under pseudocardium.
pseudocardia Eichwald. RECORDS. ··1890 Paetel: Adacna
pseudocardia Eichw.
Pseudocardita Oppenheim, 1919 (genus). - Carditidae.
Tertiary and Recent. RECORDS . ··1980 Vokes:
Pseudocardita Oppenheim, 1919 in subfamily
Pseudocarditinae.
Pseudocarditinae Ebersin, 1964 (subfamily). - RECORDS.
··1980 Vokes: Subfamily Pseudocarditinae Ebersin,
1964. ··1977 ZR: Pseudocarditinae Ebersin 1964.
Referred to Carditidae from Cardiidae. Ref. to Kafanov
& Popov p. 56 [Trans I. p. 308].
pselldocardium Deshayes , 1838. - Synonym of
Monodacna pontica (Eichwald, 1838). » Regarding
Cardium pseudocardium: Lymnocardiidae. Tertiary.
TYPE LOCALITY. The Crimea. DISTRJBUTION. Caspian Sea.
RECORDS. ··1838 Deshayes, p. 59, pI. I, figs. 1-2:
Cardium pseudo cardillm . ··1849 Middendorff:
Cardium pseudocardium = Cardium? ponticum Eichw.
(fide Andrussoff 1903). ··1858 Adams & Adams, p.
460: Adacna (Monodacna) pseudocardia Desh. ··1888
Kobelt : Monodacna p seudocardium Desh.
(Monodacna) . ··1890 Woodward : Monodacna
pseudocardium Desh. - Black Sea. ··1903 Andrussoff:
Cardium pseudocardium Deshayes, 1838. ··1903
Andrussoff: Cardium pseudocardium Deshayes, 1838
= Cardium ponticum Eichw. ··1908 Milaschewitch :
Monodacna pseudocardium . ··1908 Milaschewitch :
Monodacna pseudocardium (Deshayes) 1838 = pontica
Eichw. ··1916 Milaschewitch : Monodacna
pseudocardium (Deshayes), 1838. ··1922-32 Sherborn,
p. 5194: Cardium pseudocardium Deshayes, 1838.
··2001 Vidal, p. 407: Monodacna pseudocardium. CHARACTERS AND OCCURRENCE - DESCRIPTIONS. ··200 I Vidal : no tentacles. No eyes.
Pseudocatillini Taktakishvi Ii, 1987 (tribus). Lymnocardiidae. RECORDS. ··1987 Taktakishvili, p. 102:
Pseudocatillini; tribe nov. of Limnocardiinae; Type
genus for new tribe Pseudocatillus Andrussow, 1903.
Pseudocatillus Andrussow, 1903 (genus). Lymnocardiidae. Tertiary and Recent. RECORDS. ··1980
Vokes : Pseudocatillus Andrussow, 1903 in subfamily
Didacninae. ··1981 Taktakishvili: Pseudocatilllls.
··1982 ZR: Pseudocatillus. Romania, Bucarest region.
Upper Miocene. Ref. to Papaianopol et al. ··1984 ZR:
genus Pseudocatillus. Ref. to Akhvlediani p. 70. ··1987
Anaya: Genus Pseudocatillus Andrusov, 1923. Type
species Cardium pseudocatillus Barbot de Marny, 1869.
Pseudocatillus Andrussow, 1903 (subgenus). Lymnocardiidae. Fossil. GENOTYPE. Monodacna
pseudocatillus (Abich) Barbot de Marny (Dollfus 1905,
commenting Andrussow 1903). RECORDS. ··1905
Dollfus, commenting Andrussow 1903 : Pseudocatillus
n.s .g. ··1970 ZR: Pseudocatillus Pliocene fauna of
region between Teleajen and Cricov valleys, Romania
Phuket Marine Biological Center Special Publication vol. 30 (2004)
719
pseudocatillus - pseudofossile
Taktakishvili:
(Prosodacna) pseudocucestiensis Eberzin, 1960 sp. n.
(Pseudocatillus) subgenus of Monodacna.
pseudocatillus Abich. - Combination to be revised:
"Cardium" pseudocatillus Abich. Lymnocardiidae.
Pliocene Kamyshburun, Kerchensky, Crimea USSR.
--1987 Anaya : Prosodacna pseudocucestiensis
Eberzine, 1959.
pseudocucestiensis (cf.) Ebersin, 1959.- RECORDS.
-'1987 Anaya : Prosodacna (s . s.)
cf.
pseudocuscestiensis sic Eberzine, 1959.
pseudocuscestiensis (error). - Printer's mistake for
pseudocucestiensis in Anaya 1987.
pseudodeserta. - Nomen nudum. RECORDS. --1949 ZR:
Replidacna pseudodeserta 'no sp.' [nomen nud.]
Maestricktian, Sannat. [Ref. Papp 1949].
pseudodoengingki Barnabas & Strausz, 1989. - New
combination: Ly mnocardium pseudodoengingki
Barnabas & Strausz, 1989. Lymnocardiidae. Miocene.
TYPE LOCALITY. Pannonian in SW Transdanubia
(Hungary). RECORDS. --1989 Barnabas & Strausz p. 234
& 305, fig.: Limnocardium pseudodoengingki sp. nov.
pseudodonax Dunker. - Combination to be revised:
"Hemicardium" ps eudodonax Dkr. Cardiidae .
RECORDS. "1890 PaeteJ: Hemicardium pseudodonax
Dkr.
pseudoduplicatum - Combination to be revised:
"Cardium" pseudoduplicatum. Cardiidae. Sannatian.
RECORDS. --1965 ZR: Cardium pseudoduplicatum figd .
Sannatian, Yugoslavia [Ref. Spaj ic 1965].
pseudoedule Andrussow, 1902. - Changed combination:
[Ref.
Pau liuc
1970].
"1981
Recent. TYPE LOCALITY . Caspian Sea. DISTRIBUTION.
Caspian Sea . RECORDS . --1877 Grimm: Cardium
pseudocatillus Abich. ? "1877 ZR : Cardium
pseudocatillus (Abich), also recent in the Caspian Sea
[Ref. Grimm 1877].
pseudocatillus Barbot de Marny, 1869. - Pseudocatillus
pseudocatillus (Barbot de Marny, 1869).
Lymnocardiidae. Pliocene. Cardium pseudocatillus
Barbot de Marny, 1869 (tautotype) is type species of
genus Pseudocatillus Andrussov, 1923 in family
Limnocardiidae sic (G1ibert & Poel 1970). RECORDS.
--1905 Dollfus, commenting Andrussow 1903:
Monodacna pseudocatillus (Abich) Barbot de Marny.
--1953 Celidje: Pseudocatillus pseudocatillus (Barb.).
--1967 Ebersin: Pseudocatillus pseudocatillus (Barbot).
--1982 ZR: Pseudocatillus pseudocatillus (Barbot,
1869). Description. Ref. to Papaianopol et al. --1987
Lutskanov
&
Tcholakov:
Pseudocatillus
pseudocatillus. --1987 Anaya: Cardium pseudocatillus
Barbot de Marny, 1869. --1987 Anaya : Pseudocatillus
pseudocatillus Barbot , 1869 . --1987 Anaya:
Monodacna pseudocatillus in Andrussov sic 1917.
--1987 Lutskanov: Pseudocatillus pseudocatillus Barb.
"1987 Lutskanov & Tcholakov : Pseudocatillus
pseudocatillus Barb.
pseudocatillus larga Taktakishvili , 1966 Changed
combination:. - Monodacna pseudocatillus larga
Taktakishvili , 1966 . Lymnocardiidae . Pliocene.
REcORDs. --1966 ZR: MOllodacna pseudocatillus larga
Taktakishvili, 1966 subsp. nov. western Gruzia, SSR,
Pliocene.
pseudocimmerica (subspecies) . - See under natella
pseudocimmerica Eberzin, 1960.
Pseudoconocardium Zavodovskii , 1962 (genus).
Conocardiidae. Permian. RE CORDS. --1962 ZR:
Pseudoconocardium Zavodovskii, 1962 gen. nov. of
Conocardiidae. Type species Pseudoconocardium
licharewi Zavodovskii, 1962 sp . nov. Permian R.
Khivach, N.E. Russia.
pseudocrassa Pay!. - Didacna pseudocrassa Pay!.
Lymnocardiidae. Fossil. RECORDs. -'1989 Mikhailesku:
Didacna pseudocrassa Pay!. "1989 Klimanov,
Mikhailesku & Chepalyga : Didacna pseudocrassa
Pay!.
pseudocrassatellata Ebersin, 1958 . - Limnodacna
pseudocrassatellata Ebersin, 1958. Lymnocardiidae.
Pliocene. RECORDS. --1958 ZR: Limnodacna
pseudocrassatellata Eberzin, 1958 sp. nov. Abkhazia,
Galizga river. Pliocene. "1987 Taktakishvili:
Limnodacna pseudocrassatellata Ebers.
pseudocrassum Pavlov, 1926. - Changed combination:
Didacna pseudocrassum (Pavlov, 1926). [Tertiary and
Quaternary, Lymnocardiidae]. RECORDS. "1926 ZR:
Cardium (Didacna) pseudocrassum Pavlov, 1926 sp.
n. Tr;:rtiary S.E. Europe. -'1929 ZR : Cardium
pseudocrassum Pavlov, 1929 sp. n. Neogene &
Quaternary of S.E. Europe.
pseudocucestiensis Ebersin, 1959 . - Prosodacna
pseudocucestiensis Ebersin, 1959. Lymnocardiidae.
PI iocene. RECORDS. "1960 ZR: Prosodacna
Cerastoderma pseudoedule(Andrussow, 1902) .
Cardi idae. Fossil. RE CORD S. "1905 Dollfus,
commenting Andrussow 1902 : Cardium pseudoedule
n. sp. tend s towards Cerastoderma. "1977 Popov:
Cerastoderma pseudoedule.
pseudoesperanzae Ebersin, 1967. - Macradacna
pseudoesperanzae Ebersin, 1967. Lymnocardiidae.
Pliocene. RECORDS . -·1967 Ebersin: Macradacna
pseudoesperallzae sp. n. --1967 ZR: Macradacna
pseudoesperanzae Eberzin, 1967 sp. nov. Pliocene,
USSR.
pseudofastosum Nomura, 1937. - Changed combination:
Clinocardium pseudofastosum (Nomura) , 1937.
Cardiidae. Pliocene. RECORDS. "1937 ZR: Cardium
(Clinocardium)pseudofastosum Nomura, 1937 sp. n.
Neogene Japan . --1954 Keen: Clinocardium
pseudofastosum (Nomura), 1937 [Pliocene] . "1973
Keen: Clinocardium pseudofastosum (Nomura, 1937)
[Lower Pliocene].
pseudofischerianum Kolesnikov. - Protracti/orma
pseudojischerianum (Kolesnikov). Lymnocardiidae.
Fossil. RECORDS . "1979 ZR : Protractiforma
pseudojischerianum (Kolesnikov). Referred to new
subgenus P (Fischericardium) . Ref. to Bagdasaryan,
p.44.
pseudofossile Reeve, 1844. [Figs. Part I p. 133 . Part 3
p. 926]. - Synonym of Clinocardium cali/orniense
(Deshayes, 1839). OTHER JUDGEMENTS: Clinocardium
uchidai Habe. » Regarding Cardium pseudofossile:
Cardiidae. Recent. TYPE LOCALITY. Unknown (Reeve
1844). TYPE REPOSITORY. BMNH; Cardium
pseudofossile Reeve Syntypes reg. no. 1975617. Neg.
No. 0227. RECORDS. "1844 Reeve, sp. 52: Cardium
pseudofossile. -'1858 Adams & Adams, p. 454 :
Cardium (Pectunculus) pseudofossile, Reeve. --1864
Carpenter: Cardium pseudofossile, Rve [Asiatic] .
·-1867 ZR: Cardillm pseudofossile (Reeve) Hakodate
720
Tropical Marine Mollusc Programme (TMMP)
pseudogigas - pseudolima
and Bay of Castries (Lischke 1867) ... 1869 Lischke:
Cardium pseudofossile Reeve = Cardium californiense
Deshayes. ··1869 Romer: Cardium pseudofossile
Reeve. ··1890 Paetel: Cardium pseudofossile Rv. =
californiense Dsh. ··1896 Rolle: Cardium pseudofossile Reeve. ··190 I Dall: Cardium pseudofossile
Reeve, 1844 = Cardium californiense Deshayes, 1839.
··1922-32 Sherborn, p. 5195: Cardium pseudofossile
Reeve, 1844 [as pseudo-fossileJ. ··1952 Kuroda &
Habe: Clinocardium pseudofossile Reeve . ··1977
Fischer-Piette: Cardium pseudo-fossile in Rolle (1896)
= Corculum uchidai Habe. ··1977 Fischer-Piette:
Cardium pseudo-fossile Reeve 1844 = Corculum
californiense Deshayes. ··1980 ZR : Keenocardium
pseudofossile (Reeve 1845) is a junior synonym of
Keenocardium californiense (Deshayes 1839). Ref. to
Kafanov. ··2000 Coan et al.: Cardium pseudofossile
Reeve, 1844 = Clinocardium (Keenocardium)
californiense (Deshayes, 1839). ··2002 Schneider:
citing Dall (1900): Cardium (Cerastoderma)
pseudofossile Reeve, 1844, section Cerastoderma. CHARACTERS AND OCCURRENCE - DESCRIPTION.
··1844 Reeve: shell depressly ovate, a little produced
on each side, radiately ribbed, about 50 narrow, smooth
ribs very numerous, a little swollen here and there; rusty
white. DIMENSIONS. ··1896 Rolle: length 70 mm, width
60 mm, diameter 40 mm. REMARKS ··1844 Reeve: this
is a remarkable species and has so much the appearance
of being in a fossil state, that I only felt satisfied to the
contrary upon discovering a large portion of fresh horny
ligament attached to the hinge of one of Mr. Cuming's
specimens. ··1869 Romer: Cardium pseudofossile
Reeve is closely related to Cardium ciliatum. ··1896
Rolle found that Cardium pseudofossile is closely
related to Clinocardium callforniense (Deshayes,
1839). They can hardly be separated based on the shell
form. HABITAT ··1844 Reeve: unknown. DISTRIBUTION.
North Japan, Alaska.
pseudogigas Eames, 1951. - Trachycardium pseudogigas Eames, 1951. Cardiidae. Eocene. RECORDS.
··1951 ZR: Trachycardium pseudogigas Eames, 1951
sp. nov Eocene Pakistan.
pseudohillanum de Ryckh. in Dew. - Synonym of
Protocardium edwardsi (Deshayes). » Regarding
Cardium pseudohillanum: Cardiidae. Eocene. RECORDS.
··1886 Cossmann: Cardium pseudohillanum de Ryckh.
in Dew. = Protocardium edwardsi (Deshayes).
pselldohispida (variety). - See under suessi
pseudohispida Simionescu & Barbu, 1940.
pseudo[at;costatus Chelidze, 1975 . - Pseudocatillus
pseudolaticostatus Chelidze, 1975. Lymnocardiidae.
Lower Pliocene. RECORDS. ··1975 ZR: Pseudocatillus
pseudolaticostatus Chelidze, 1975 sp. nov. Georgia
SSR, Khobs region. Ref. to Chelidze, p. 188.
pseudo/atum Voskuil & Onverwagt, 1991.
Maoricardium pseudolatum (Voskuil & Onverwagt,
1991). Cardiidae. Recent. TYPE LOCALITY. Padang,
western Sumatra, Indonesia (Higo et af. 1999).
RECORDS. ··1991 Voskuil & Onverwagt, p. 31-32, figs.
1-3 , 12): Maoricardium pseudolatum n. sp. ··1992
Lamprell & Whitehead: Plagiocardium (Maoricardium) pseudolalum. ··1998 Schneider: Maoricardium pseudolatum Voskuil & Onverwagt, 1991.
··1998 Poutiers: Plagiocardium pseudolatum (Voskuil
& Onverwagt, 1991). ··1998 Schneider (l998b):
Maoricardium pseudolatum Voskuil & Onverwagt,
1991 . ··1999 Higo et al.: Maoricardium pseudalatum
Voskuil & Onverwagt, 1991 . ··2002 Hylleberg &
Kilburn, p. 63: Plagiacardium pseudolalum (Voskuil
& Onverwagt, 1991). - CHARACTERS AND
OCCURRENCE - DISTRIBUTION. ··1994 Hylleberg:
Indo-West Pacific. ··1999 Higo et af.: apparently not
distributed in Japan. REMARKS. ··2002 Hylleberg &
Kilburn: the Indian species is probably Plagiacardium
pseudolatum (Voskuil & Onverwagt, 1991) but the
distinguishing characters between Plagiacardium
setosum and Plagiacardium pseudolatum are not clear.
pseudolima Lamarck, 1819. [Figs. Part I p. 120, 124,
171 . Part 3 p. 902]. - Changed combination: Plagiacardium pseudalima (Lamarck, 1819). Cardiidae.
Recent. Type species: Cardium pseudolima Lamarck,
1819. Type species of Plagiacardium Cossmann, 1886.
TYPE LOCA LITY. Unknown (Lamarck 1819). TYPE
REPOSITORY. Cabinet of Mr. Dufresne, (Lamarck 1819).
Unknown availability. DESCRIPTIONS. Oliver (1995): No
transverse ridges in the interstices (Lamarck 1819).
Subquadrate, distinctly longer than high, umbones high,
posterior margin almost straight. Sculpture of 36-40
flat radial ribs these bearing small cup--shaped spines
(nodules if worn) arising from the mid-line of each
rib. In fresh specimens periostracal bristles arise from
the base of each spine. Pink to reddish brown with
contrasting white spines. DIMENSIONS. 110 mm length.
To 150 mm (Oliver 1995). HABITAT. Muddy sand
(Oliver 1995). DISTRIBUTION. Red Sea (Sowerby 1841);
Zanzibar, E. Africa (Reeve 1844); Zanzibar (Grasset
1884); East Africa, Persian Gulf (Hylleberg 1994);
Arabian Sea (Oliver 1995). RECORDS. ··1819 Lamarck,
p. 7, No. 13 : Cardium pseudalima. ··1840 Sowerby,
183th part, fig. 67: Cardium pseudalima, Lam. ··1840
Sowerby, catalogue p. 2, No. L8 : Cardiumpseudolima,
Lam. ··1840 Sowerby, corrected list, p. 8: Cardium
pseudolima (Sowerby, No. 18). ··1844 Reeve: Cardium
pseudolima Lamarck Anim. sans vert. vol. vi. p. 396.
··1858 Adams & Adams, p. 455: Cardium (Isocardia)
pseudolima , Lam. ··1868 ZR: Cardium pseudolima
(Lam.), much more rarely white than pink. [Ref. Romer
1868]. ··1890 Paetel: Cardiumpseudolima Lm.·· j 92232 Sherborn, p. 5195: Cardium pseudolima Lamarck,
1819. ··1927 Lamy: Cardium (Acanthocardia)
pseudolima. ··1930 Stewart: "e." pseudo lima (Recent,
Indian Ocean). ··1964 Spry : Cardium pseudalima
(Lamarck) . ··1977 Fischer-Piette : Cardium
(Cerastoderma) pseudo lima Lamarck. ··1977 Wilson
& Stevenson: Maaricardium pseudalima (Lamarck,
1819). ··1982 Bosch & Bosch: Cardium pseudalima
Lamarck, 1819. ··1990 Abbott & Dance:
Plagiacardium pseudalima. ··199 L Voskuil &
Onverwagt : Maoricardium pseudalima (Lamarck,
1819). ··1992 Oliver: Rudicardia pseudalima. ··1995
OLiver, fig. 1079: Plagiacardium pseudalima (Lamarck,
1819). - CHARACTERS AND OCCURRENCE DESCRIPTION . •• L844 Reeve: shell somewhat heartshaped , ventricose, radiately grooved, about 38
grooves, intermediate ridges rather flat, sometimes the
whole, sometimes the lateraL ridges only, armed with
rows of very minute sharp-scaLed tubercles; pink fleshcolour, ornamented with pink transverse rays, tubercles
Phuket Marine Biological Center Special Publication vol. 30 (2004)
721
pseudolima beta - pseudosubdentata
white. HABITAT. "1844 Reeve: found in deep water.
DISTRIBUTION . --1844 Reeve: Zanzibar, east coast of
Africa.
pseudolima beta (variety). - Cardium pseudolima
Variety beta. Cardiidae. Recent. RECORDS. --1844
Reeve: Shell entirely snowy white. Mr. Cuming
possesses the variety beta of this well-known species,
of a pure snowy white, in fine fresh condition.
pseudomagnum McCoy, 1877. Hedecardium
pselldomagnum (McCoy, 1877). Cardiidae. Oligocene.
RECORDS. --1877 McCoy: Cardium pseudomagnum
McCoy, 1877 (fide Schneider 2002). --1970 Darragh
assigned Cardium pseudomagnum to Trachycardium
(fide Schneider 2002). --2002 Schneider: Hedecardium
pseudomagnum (McCoy, 1877). Victoria. Oligocene.
pseudomaxima Davidasvili , 1930. - Monodacna
pseudomaxima Davidasvili, 1930. Lymnocardiidae.
Pliocene . RECORDS. --1932 ZR: Monodacna
pseudomaxima Davidasvili, 1930 sp. n. Pliocene
Transcaucasia.
pseudomichailov (variety) . - See under gatuevi
pseudomichailov KOlesnikov, 1929.
pseudomulicostatum (error). - Printer's mistake for
pseudomulticostatum in Popov 1977.
pseudomulticostatum Zhizchenko, 1934. - Changed
combination: Europicardillm pseudomulticostatum
(Zhizchenko, 1934). Cardiidae. Miocene. RECORDS.
--1934. Zhizhchenko: Cardium pseudomulticostatum
Zhizhchenko, 1934 (fide Schneider 2002). --1937 ZR:
Cardium pseudomulticostatum Zhizchenko, 1936 sp.
n. --1939 ZR : Cardium pseudomulticostatum
Zhizhchenko, 1934 sp. n. Miocene Ciscaucasia. --1961
ZR: Cardium pselldomulticostatum ?Miocene. --1977
Popov: Bucardium (Europicardium) pseudomulticostatum (Zhizchenko, 1934). --1977 Popov, Tab.
I, pp. 20-21 : Bucardium (Europicardium) pseudomulicostatum sic. --1977 ZR: Bucardium (Europicardium) pseudomulticoslalum (Zhizhchenko, 1934)
Comb. nov. Transferred from Trachycardium . Ref. to
Popov p. 44 . --1989 Gontsharova, p. 123 : Europicardium pseudomulticostatum (Zhizhchenko, 1934).
--2002 Schneider : Europicardium pseudomulticostatum (Zhizhchenko, 1934). Ukraine.
pseudomulticostatum elongata Baghdasaryan, 1961
(variety). - Upgraded. New combination:
Europicardium pseudomulticostatum (Zhizchenko,
1934). RECORDS. --1961 ZR: Cardium pseudomulticostatum var. elongata Baghdasaryan, 1961 nov.
Chokraksk deposits, ?Miocene, Georgia, Russia.
pseudoobsoletum Fuchs, 1873. - New combination:
Lymnocardium pseudoobsoletum (Fuchs, 1873).
Lymnocardiidae. Pliocene . RECORDS. --1987 Anaya:
Cardium pseudo-obsoletum Fuchs, 1873. --1987
Anaya: Limnocardium sic (s. s.) the group pseudoobsoletum Fuchs, 1873. --1987 Taktakishvili:
Limnocardium pseudoobsoletum (Fuchs).
pseudoobsoletum Kolesnikov, 1929. - Combination to
be revised: "Cardium " pseudoobsoletllm Kolesnikov,
1929. Lymnocardiidae. Sarrnatian. RECORDS. --1929
ZR: Cardium pseudoobsoletum Kolesnikov, 1929 sp.
n. Sarrnatian.
pseudoottonis Frolenkova, 1983. - Combination to be
revised : "Cardium " pseudoottonis Frolenkova, 1983 .
Cardiidae. Upper Cretaceous. RECORDS . --1983 ZR :
Cardium pseudoottonis Frolenkova, 1983 sp. nov.
Tadzhikistan, Upper Cretaceous. Ref. to Frolenkova,
p.162.
pseudoovata Gillet, 1965. - Didacna pseudoovata Gillet,
1965. Lymnocardiidae. Pliocene. RECORDS. --1987
Anaya : Didacna (s. s.) pseudo-ovata Gillet, 1965.
pseudo-petersi (invalid spelling). - See under
pseudopetersi.
pseudopetersi Stevanovic, 1951 . - New combination:
Lymnocardium pseudopetersi Stevanovic, 1951 .
Lymnocardiidae . Ponti an . RECORDS. ·-1953 ZR :
Limnocardium pseudopetersi Stevanovic, 1951 sp. nov.
Pontian Serbia. --1987 Taktakishvili: Limnocardium
pseudopetersi Stev.
Pseudopis Cox, 1946 (genus). - Cardiidae. Fossil.
REcORDs . --1946 ZR : Pselldopis Cox, 1946 gen nov.
Cardiidae genotype Pseudopis astonensis Cox, 1946
sp. nov. Lower Aston Magna Wores.
pseudoplanicostata Gillet, 1965. - Didacna
pseudoplanicostata Gillet, 1965. Lymnocardiidae.
Pliocene . RECORDS. --1987 Anaya: Didacna (s. s.)
pseudo-planicostata Gillet, 1965.
pseudoplicatum (subspecies and variety). - See under
praeplicatum pseudoplicatum and plicatum pseudoplicatum Friedberg, 1934.
pseudopoiitionanci. - Nomen nudum. RECORDS. --1949
ZR: Replidacna pseudopoiitionanci 'no sp.' [nomen
nud.] Maestricktian, Sarmat. [Ref. Papp 1949].
pseudoproductum Peth6, 1906. - Combination to be
revised: "Cardium " pseudoproductum Peth6, 1906.
Cardiidae . Cretaceous. TYPE LOCALITY. Hungary.
REcORDs. --1906 Peth6 : Cardium pseudoproductum sp.
n. --1906 ZR: Cardium pseudoproductum Peth6, 1906
sp. n. Cretaceous, Hungary.
Pseudoprosodacna Gi lIet, 1943 (genus) .
Lymnocardiidae . Tertiary. RECORDS. --1946 ZR:
Pseudoprosodacna Gillet, 1943 gen. nov. between
Limnocardium and Prosodacna first species Cardium
littorale Eich. --1980 Vokes: Pseudoprosodacna Gillet,
1943.
pseudoschemaclrinica Gillet & Vicente, 1961. - Didacna
pseudos chema chinica Gillet & Vicente, 1961.
Lymnocardiidae. Pliocene . RECORDS. --1961 ZR:
Didacna pseudoschemachinica Gillet & Vicente, 1961
sp. nov. Pliocene, Barcelona, Spain. --1987 Anaya:
Didacna (s. s.)pseudoschemachinica Gillet, 1961.
pseudosecans Andrusov, 1917. - Changed combination:
Euxinicardium pseudosecans (Andrusov, 1917).
Lymnocardiidae. Tertiary. RECORDS. --1987 Anaya:
Limnocardium pseudosecans Andrusov, 1917. --1987
Anaya: Limnocardium (Euxinicardium) the group
pseudosecans Andrusov, 1917. --1987 Taktakishvili:
Limnocardium (Euxinicardium) pseudosecans Andrus.
--1987 Taktakishvili: Euxinicardium pseudosecans.
pseudosecans (aff.) Andrusov, 1917. - RECORDS. --1987
Taktakishvili: Limnocardium aff. pseudosecans Andrus.
pseudosoceni Jekelius, 1944. - Combination to be
revised: "Cardium " pseudosoceni lekelius, 1944.
Cardiidae. Miocene. RECORDS. --1946 ZR : Cardium
pseudosoceni Jekelius, 1944 sp. nov. (type f. II) .
Miocene Soceni Banat.
pseudosubdentata Celidje, 1953 . - Monodacna
pseudosubdentata Celidje, 1953 . Lymnocardiidae.
Fossil. REcORDs. --1953 Celidje, p. 157: Monodacna
722
Tropical Marine Mollusc Programme (TMMP)
pseudosubdentatus - pulchella
pseudosubdentata nov. sp.
pseudosubdentatus Chelidze, 1975. - Pseudocatillus
pseudosubdentatus Chelidze, 1975. Lymnocardiidae.
Lower Pliocene. REcORDs. --1975 ZR: Pseudocatillus
pseudosubdentatus Che1idze, 1975 sp. nov. Georgia
SSR, Khobs region. Ref. to Chelidze, p. 185.
pseudosuessi Hal. - New combination: Lymnocardium
pseudosuessi (Hal.). Lymnocardiidae. Fossil. REcORDs.
"1987 Taktakishvi li: Limnocardium pseudosuessi
(Hal.).
pseudosulcatina Gillet, 1965 .
Didacna
pseudosulcatina Gillet, 1965. Lymnocardiidae.
Pliocene. RECORDS. --1987 Anaya: Didacna (s. s.)
pseudosulcatina Gillet, 1965 [Pliocene].
Pseudotrapezium (genus). - Cardiacea. Fossil. REcORDs.
--1916 ZR: Pseudotrapezium (fossil) Ref. to Cossmann
(1916).
Psilodon Cobalcescu, 1883 (genus). - Synonym of
Prosodacna Tournouer, 1882. » Regarding Psilodon :
Lymnocardiidae. Tertiary and Recent. Type species:
Cardium macrodon Deshayes type of the genus
Psilodon Cobalcescu, 1882 (Andrussoff 1903).
REcORDS. --1887 Fischer: Psilodon, Cobalcescu. 1882?
lun. syn of subgenus Prosodacna Tournouer, 1882.
--1896 ZR: Stylodacna n. g. for Psilodon herbeti
Cobalcescu [Ref. Stefanescu 1896]. .. 1975 ZR:
Psilodon Cobalcescu, 1883. Emendation. Ref. to
Motas, p. 60. "1978 ZR : Prosodacna (Psilodon)
Pliocene. Romania. Dacian, systematics & phylogeny.
Ref. to PapaianopoJ, I. --1980 Vokes: Psilodon
Coba\cescu, 1883 in subfamily Lymnocardiinae.
Psilodonini Taktakishvili, 1987 (tribus). Lymnocardiidae. RECORDS. --1987 Taktakishvili (p.
159) designated Psilodon Cobalcescu 1883 type genus
of the new tribe Psilodonini.
Pteradacna Andrussow 1907 (genus). - Pteradacna
Andrussow 1907. Lymnocardiidae. Pliocene. RECORDS.
--1951 ZR: Pteradacna Andrussov 1907 and synonymy
of Pteradacna pterophora Brus. [Ref. Stevanovic
1951]. "1962 ZR: Pteradacna Andrussov in subfam.
Didacninae Eberzin, 1962, family Cardiidae [Ref.
Eberzin 1962]. --1977 Popov: Pteradacna Andrussow,
1907. --1980 Vokes: Pteradacna Andrussow, 1907 in
subfamily Didacninae. "1987 Taktakishvili p. 100,
Pteradacna Andrussow 1907; referred to new tribe
Pontalmyrini. --2001 Schneider & Carter: Pteradacna
(genus). Lymnocardiidae. Pliocene.
Pterocardia Bayan, 1874 (genus). - Cardiidae. Jurassic.
RECORDS. --1887 Fischer: Subgenus Pterocardia
(Agassiz) Bayan, 1874 of genus Cardium Linne, 1758.
"1931 ZR: Pterocardium Ref. to Douville (1931).
--1964 ZR : Pterocardia eparcyensis [Fischer, 1964.
Jurassic]. "1978 ZR: Pterocardia Favre 1834 sic.
Transferred from Megalodontacea. Ref. to Morris p.
267.
Pterocardium (error). - Printer's mistake for genus
Pterocardia in Deckaseaux 1945 and Douville 1931.
pteropernoida Repman, 1961. - Arkitella pteropernoida
Repman, 1961. Cardiidae. Jurassic. RECORDS. "1961
ZR: Arkitella pteropernoida Repman, 1961 sp. nov.
Arkit, USSR Jurassic. Genotype of Arkitella Repman,
1961.
pterophora Brusina. - Changed combination :
Pteradacna pterophora (Brusina). Lymnocardiidae.
Fossil. RECORDS . --1907 Andrussow : Adacna
pterophora Brusina compared with Prosodacna
semisulcata Rouss. --1951 ZR: Pteradacna pterophora
Brus. synonymy [Ref. Stevanovic 1951].
pubescens Couthouy, 1839. - Synonym of
Ciliatocardium ciliatum (Fabricius, 1780). »
Regarding Cardium pubescens: Cardiidae. Recent.
DISTRIBUTION. Iceland (Reeve 1844); Iceland, Greenland
(Wood & Hanley 1856); circumpolar. RECORDS. --1839
Couthouy, p. 60, pI. 3, fig. 6: Cardium pubescens.
--1844 Reeve: Cardium pubescens = juvenile Cardium
icelandicum Chemnitz, 1782. --1850 Jay: Cardium
pubescens Couth. = Cardium icelandicum Linn. sic.
--1856 Wood & Hanley: C. pubescens Couthouy =
Cardium icelandicum Ch. --1869 Romer: Cardium
pubescens Couthouy = Cardium cilia tum Fabricius.
"1870 Gould: Cardium pubescens Couthy. (syn.) =
Cardium islandicum Linn. --1873 Tryon: Cardium
pubescens Couthouy = Cardium islandicum, Linnaeus,
1767. --1888 Kobelt: Cardium pubescens Couthouy =
Cardium islandicum Linne. --1902 Dall: Cardium
pubescens Couthouy, 1838 = Cardium cilia tum
Fabricius, 1780. --1912 Dautzenberg & Fischer:
Cardium pubescens Ch. Couthouy 1838 = Cardium
(Cerastoderma) islandicum, Chemnitz 1782. "1912
Jensen: Cardium pubescens Couthouy, 1838 = Cardium
ciliatum Fabricius ... 1922-32 Sherborn, p. 5214:
Cardium pubescens Couthouy, 1838. --1924 Schlesch:
Cardium pubescens Couthouy = Cardium
(Cerastoderma) ciliatum (Fabricius). --1939 McLean :
Cardium pubescens = Clinocardium cilia/um Fabricius,
1780. --1974 Abbott: Cardium pubescens Couthouy,
1838 = Clinocardium cilia tum (Fabricius, 1780). --1977
Fischer-Piette: Cardium pubescens Couthouy =
Corculum cilia tum Fabricius. --2000 Coan et al. : C.
pubescens Couthouy, 1838 = Clinocardium
(Ciliatocardium) ciliatum (Fabricius, 1780).
pubescens (variety). - Infrasubspecific. RECORDS. --1890
Paetel: Cardium var. pubescens Couth.
puebloensis Anderson, 1929. - Trachycardium
puebloensis Anderson, 1929. Cardiidae. Miocene.
RECORDS. --1929 ZR: Cardium (Trachycardium)
puebloensis Anderson, 1929 spp. n. Miocene N.
Carolina.
puelchum Sowerby in Darwin, 1846. - Combination to
be revised: "Cardium" puelchum Sowerby in Darwin,
1846. Cardiidae. Tertiary. RECORDS . --1922-32
Sherborn, p. 5219: Cardium puelchum Sowerby in
Darwin, 1846. --2002 Schneider: Cardium puelchum
Sowerby, 1846. Tertiary sediments in Patagonia.
pulcellum White & Whitfield, 1862. - Conocardium
pulcellum White & Whitfield, 1862. Conocardiidae.
Fossil. REMARKS. Conocardiids have no connection with
the Cardiacea except analogy of form, which does not
offer any point of relationship (Dall 1895, p. 519).
RECORDS . --1980 Ruhoff: Conocardium pulcellum
White & Whitfield, 1862; vol. 8: 299.
pulchella (variety). - See under rectidens pulchella.
pulchella Chip10nkar & Tapaswi, 1976. - Pr%cardia
pulchella Chiplonkar & Tapaswi, 1976. Cardiidae.
Upper Cretaceous. RECORDS. --1983 ZR: Protocardia
(Tamilicardia) pulchella Chiplonkar et al. , 1983 sp. nov.
India. Upper Cretaceous. Ref. to Chiplonkar et al. p.
153 ... 1995 Schneider: Protocardia (Tamilicardia)
Phukel Marine Biological Center Special Publication vol. 30 (2004)
723
pulchellum - pulcherrimum
pulchella Chiplonkar & Tapaswi, 1976.
*pulchellum. - Homonymy to be noticed. Cardium
pulchellum from 3 sources: Gray in Dieffenbach, 1843.
- Philippi, 1844. - Reeve, 1844.
pulchellum Gray in Dieffenbach, 1843. - Changed
combination: Microcardium pulchellum (Gray, 1843)
OTHER JUDGEMENTS: in genus Nemocardium.
Nemocardium thetidis (Hedley, 1902). - Cardium
striatulum Sowerby. » Regarding Cardium
pulchellum: Cardiidae. Recent. TYPE LOCALITY. New
Zealand (pulchellum Gray, 1843); Australia
(pulchellum Reeve, 1844). RECORDS. --1843 Gray in
Dieffenbach: Cardium pulchellum. "1885 Smith:
Cardium pulchellum Gray. --1887 Tate: Cardium
pulchellum Gray. --1889 Whitelegge: Cardium
pulchellum Gray. --1903 Pritchard & Gatliff: Cardium
pulchellum, Gray = Cardium striatulum , Sowerby.
--1904 Hutton: Cardium pufchellum Gray. --1905
Hedley: Cardium pulchellum Gray. --1906 Suter:
Protocardia (Nemocardium) pulchella (Gray). --1907
Suter: Protocardia pulchella Gray. --1907 ZR: Cardium
pulchellum [Ref. Hedley 1907]. --1907 ZR: Cardium
pulchellum Gray ref. to Protocardia (q.v.) subgen.
Nemocardium [Ref. Suter 1907]. --1912 Suter:
Protocardia pulchella Gray. --1913 Suter: Protocardia
pulchella Gray. --1915 Oliver: Protocardia pulchella
Gray. --1915 Smith: Cardium pulchellum Gray. --1918
Hedley: Cardium pulchellum Gray. --1922-32
Sherborn, p. 5224: Cardium pulchellum Gray in
Dieffenbach, 1843. --1924 Iredale: Cardium pu{chellum
Gray, 1843. "1939 Powell, p. 213: Nemocardium
(Pratulum) pu{chellum (Gray 1843). "1950 Keen:
Nemocardium (Pratulum) pulchellum (Gray, 1843).
--1953 Macpherson & Chapple: Cardium pulchellum
Gray. --1955 Powell: Nemocardium pulchellum (Gray).
--1956 Gabriel: Cardium pulchellum Gray. --1962
Powell: Nemocardium pulchellum (Gray). "1970
Glibert & Poel: Nemocardium (Pratufum) pulchellum
(Gray, 1843). "1975 ZR: Nemocardium (Pratulum)
pulchellum. Depth distribution. Habitat substrate. New
Zealand zone. Ref. to Luckens. --1977 Fischer-Piette:
Cardium pulchellum Gray = Pratulum striatulum
Sowerby, 1834 . --1977 Fischer-Piette: Cardium
pulchellum Gray = Cardium thetidis Hedley, 1902.
"1977 Popov : Pratulum pulchellum (Gray, 1843).
--1979 Powell: Nemocardium (Pratulum) pulchellum
(Gray, 1843). - CHARACTERS AND OCCURRENCE - OF COLLECTIONS. --ZMUC Colville Channel,
New Zealand: 30, 70, 123, 128, 268 , 293 m, West
Pacific, Milford Sound. Campbell Island. --ZMUC: I
large specimen from Queen Charlotte Sound, New
Zealand, coli . Mortensen, det. Odhner. --ZMUC:
Albatross point New Zealand, 25 fathoms Twofold Bay,
NS Wales . --ZMUC: wet collection New Zealand.
DESCRIPTIONS. Fine radial riblets. Two divergent radial
streaks of dark pink. DIMENSIONS. 29 x 26 x 18 mm .
REMARKS. Closely related to thetidis Hedley, 1902
(Powell 1979). "1844: Reeve ウエ。・セ@
that Cardium
striatulum jun. , Sowerby, 1840 (l77 t part, fig. 45)
cannot surely be the young of the Cardium striatulum.
Therefore, Reeve coined the new name Cardium
pulchellum which, however, also was used by Gray,
1843, probably for the same species. Fischer-Piette had
the opinion that C. pulchellum Rve. = C. striatulum
Sowerby. --1979 Powell: the species is closely allied
to the New South Wales thetidis. HABITAT. --ZMUC:
sandy mud, sandy clay, mud, fine sand, mud at 18-293
m depth. DISTRIBUTION. "1950 Keen, "1979 Powell:
New Zealand. --ZMUC: New Zealand, Campbell
Island Twofold Bay, NS Wales.
pulchellum Philippi, 1844. - Replaced by "Cardium "
philippii Deshayes. » Regarding Cardium pulchellum:
Cardiidae. Fossil. RECORDS. --1857 Deshayes: Cardium
pulchellum Philippi, 1844. - C. philippii Desh.
pu/chellum Reeve, 1844. [Figs. Part I p. 131. Part 3 p.
928] . - Synonym of Nemocardium pulchellum (Gray,
1843). OTHER JUDGEMENTS: Cardium thetidis Hedley,
1902. - Pratulum striatulum (Sowerby, 1834).
Cardiidae. Recent. TYPE REPOSITORY. 3 syntypes .
1978127. BMNH London. TYPE LOCALITY. Australia
(Reeve, 1844). RECORDS. --1844 Reeve, sp. 42: Cardium
pulchellum. --1857 Deshayes: Living . Cardium
pulchellum Reeve , 1844. --1867 Angas : Cardium
pulchellum Reeve. "1869 Dunker: Laevicardium
pulchellum Reeve. --1872 Tryon: Cardium pulchellum
Reeve. --1878 Tenison-Woods: Cardium pulchellum
Reeve. --1879 Martens: Cardium pulchellum Reeve.
--1884 Grasse!: Cardium pulchellum. Rv. --1890 Paetel :
Cardium pulchellum Rv. --1901 Tate & May: Cardium
pulchellum Reeve. --1962 Graham: Nemocardium
pulchellum Rev. --1977 Fischer-Piette: Cardium
pulchellum Reeve 1844 = Cardium pulchellum Gray
1843 = Cardium thetidis Hedley, 1902 = Pratulum
striatulum Sowerby, 1834. - CHARACTERS AND
OCCURRENCE - DESCRIPTIONS. "1844 Reeve: shell
orbicular, thin, somewhat depressed, nearly equilateral,
radiately closely striated, striae solid, raised, regular,
the posterior striae covered with very minute scales;
white, striae some few white, most of them bright
scarlet; interior yellowish, two scarlet rays diverging
within the umbone in each valve. REMARKS. This very
pretty species cannot surely be the young of the
Cardium striatulum; it is of a different form, being truly
orbicular, and offers no indication of the wrinkled striae ·
with which that species is so peculiarly cancellated.
DISTRIBUTION. "1844 Reeve: Australia.
pu/cherrima (variety). - See under victor pulcherrima
Melvill, 1909.
pulcherrimum Sakurai & Habe, 1966. - Synonym of
Acrosterigma transcendens (Melvill & Standen, 1899
OTHER JUDGEMENTS: Laevicardium biradiatum
(Bruguiere , 1789). » Regarding Laevicardium
pulcherrimum: Cardiidae. Recent. TYPE LOCALITY.
Holotype, Kakeroma Jima, Amami Is. Japan. TYPE
REPOSITORY. Holotype, National Science Museum,
Japan. TYPE MATERIAL. Holotype and two paratypes, in
National Science Museum, Japan ref. NSMT-MO
48405 and MO 70S79 (Vidal 1999). RECORDS. --1966
Habe et Kosuge, p. 295-296 : Laevicardium
pulcherrimum sp. nov. --1966 ZR: Laevicardium
pulcherrimum Sakurai & Habe, 1966 sp. nov.
Kakeroma-jima, Amami Is . Japan. --1973 Keen :
Laevicardium pulcherrimum Sakurai & Habe, 1966.
Recent. --1977 Fischer-Piette: Laevicardium
pulcherrimum Sakurai & Habe. --1977 Habe:
Laevicardium pulcherrimum Sakurai & Habe, 1966.
--1981 Koyama et al.: Laevicardium pulcherrimum
Sakurai & Habe, 1966 = Trachycardium transcendens
724
Tropical Marine Mollusc Programme (TMMP)
pulchricostatum - pulchrum
(Melvill & Standen, 1899). "1999 Higo et af.:
Laevicardium pulcherrimum Sakurai & Habe, 1966 =
Acrosterigma (Vasticardium) transcendens (Melvill &
Standen, 1899). "1999 Vidal: Sakurai & Habe 1966,
p. 293: Laevicardium pulcherrimum = Acrosterigma
transcendens (Melvill & Standen , 1899) . CHARACTERS AND OCCURRENCE - REMARK S.
Fischer-Piette (1977, p. 26) refers to the authors stating
the Laevicardium pulcherrimum Sakurai & Habe, 1966
is close to Laevicardium biradiatum (Bruguiere, 1789)
(= Laevicardium rubropictum), and also to
Vasticardium arenicola Reeve; pulcherrimum is ill
defined, and, with some hesitation, considered ajunior
synonym of Laevicardium biradiatum by FischerPiette (1977). DISTRIBUTION. Indo-Pacific (Hyl\eberg
1994).
pulchricostatum Iredale, 1929. [Figs. Part 3 p. 885] . Synonym of Vepricardium multispinosum (Sowerby,
1838). » Regarding Vepricardiwn pulchricostatum :
Cardiidae. Recent. Vepricardium pulchricostatum
Iredale, 1929 (DO) is type species of subgenus
Vepricardium Iredale, 1929 (Glibert & PoeI1970). TYPE
LOCALITY. Australia. TYPE REPOSITORY. Australian
Museum, Sydney. Holotype C057839. RECORDS. "1929
Iredale: Vepricardium pulchricostatum. "1929 Iredale
designated pulchricostatum genotype. "1929 ZR:
Vepricardium pulchricostatum [type] sp. n. Iredale,
1929a E. Australia. "1950 Allan: Vepricardium
pulchricostatum = Cardium multispinosum. "1961
Rippingale & MacMichael: Vepricardium pulchricosta tum Iredale. "1964 Shikama: Cardium
(Vepricardium) pulchricostatum (Iredale). "1969
Moore et af.: Vepricardium (Vepricardium) multispinosum Iredale, 1929. "1974 Habe & Kosuge:
Vepricardium pulchricostatum Iredale, 1927. "1977
Fischer-Piette: Cardium pulchricostatum Iredale =
Laevicardium multispinosum Sowerby. "1977 Popov :
Bucardium (Vepricardium) pulchricostatum (Iredale,
1929). "1977 Wilson & Stevenson: Vepricardium
pulchricostatum Iredale, 1929; junior synonym of
Cardium multispinosum Sowerby, 1838. "1990 Abbott
& Dance: Vepricardium pulchricostatum Iredale, 1929
= Trachycardium multispinosum. "1992 Lamprel\ &
Whitehead : Vepricardium pulchricostatum Iredale,
1929 = Vepricardium mullispinosum (Sowerby, 1838).
"1999 H igo et al.: V. pulchricostatum Iredale, 1929 =
Vepricardium multispinosllm (Sowerby, 1841). "2000
Vidal, p. 458: Vepricardium pulchricoslatum Iredale,
1929, from Australia, is probably = Vepricardium
multispinosum (Sowerby, 1841). - CHARACTERS
AND OCCURRENCE - DISTRIBUTION. Solomon
Islands, Australia, Indo-Pacific (Hylleberg 1994).
*pulchrum. - Homonymy to be noticed. Cardium
pulchntm from 4 sources: Reeve, 1845. - Deshayes,
1858. - Briiggen, 1910. - Hinds (?year).
pulchrum Briiggen, 1910. - Combination to be revised:
"Cardium" pulchrum Briiggen, 1910. Cardiidae.
Senon. "1910 ZR: Cardium pulchrum Briiggen, 1910
sp. n. Peru Senon.
pulchrum Deshayes, 1858. - Synonym. Combination
to be revised: "Cardium"" bouei Deshayes. OTHER
JUDGEMENTS: Cardium obliquum Lamarck. »
Regarding Cardium pulchrum: Cardiidae. Eocene.
RECORDS. "1886 Cossmann: Cardium bouei Desh. =
C. pulchrum = Cardium obliquum Lamarck. "1980
Ruhoff: Cardium pulchntm Deshayes, 1858; [18561865] atlas I, pI. 55, figs . 25-28. "1995 Le Renard et
af.: Loxocardium bouei [Cardium bouei] Deshayes,
1858 (pulchntm).
pulchrum Hinds. - Combination to be revised :
"Cardium" pulchrum(Hinds.). Cardiidae. Recent.
DISTRIBUTION . Japan (Grasset 1884); Persian Gulf
(Melvill & Standen 1906). RECORDS. "1884 Grassel:
Cardium pulchntm Hinds. ··1906 Melvill & Standen,
p. 839: Cardium (Cerastoderma) pulchntm Hinds.
pulchrum Reeve, 1845. [Figs. Part I p. 141. Part 3 p.
929]. - Synonym of Fulvia australis (Sowerby, 1834).
OTHER JUDGEMENTS: Cardium striatum Spengler, 1799.
- Papyridea papyraceum (Chemnitz) Schroter [not
available]. » Regarding Cardium pulchrum:
Cardiidae. Recent. TYPE REPOSITORY. 3 syntypes.
1978136. BMNH London. RECORDS. "1845 Reeve, sp.
98: Cardium pulchrum. "1858 Adams & Adams, p.
457: Laevicardium pulchntm, Reeve. "1870 Morch:
C. pulchrum Reeve = C. australe Sowb.; junior
synonyms of Cardium striatum Spgl. "1878 Smith:
Cardium pulchrum Reeve = a variety of Cardium
(La evicardium) australe Sowerby. "1869 Romer:
Cardium pulchrum Reeve = Cardium australe
Sowerby. "1878 ZR: Cardium australe (Sow.) =
pulchrum (Reeve), Andaman examples [Ref. Smith
1878]. "1880 Martens: Cardium (Papyridea) pulchrum
Reeve (from China). "1882 Rochebrune: Laevicardium
pulchrum Reeve. "1884 Grasset: Cardium pulchntm
R v. (Papyridea). "1885 Smith: Cardium pulchrum,
Reeve , fig. 98 = Cardium (Bl/cardium) australe,
Sowerby, 1840. "1889 Martens: Cardium (Papyridea)
pulchrum Sow., Reeve. "1890 Paetel: Cardium
pulchrum Hinds. "1891 Fischer: Cardium pulchrum
Reeve. "1909 Lynge : Cardium pulchnlm Reeve, 1845
= Cardium australe Sowerby, 1840. "1909 Melvill:
pulchntm Reeve = Cardium (Papyridea) australe,
Sowb. '·1922-32 Sherborn, p. 5232: Cardium pulchntm
Reeve, 1845 . "1926 Fischer & Fischer: Cardium
pulchntm Rve. = Cardium (Papyridea) australe. "1928
Kuroda : pulchrum Reeve, 1845 syn. = Papyridea
papyracea (Bruguiere, 1789). "1937 Viader: Cardium
pulchntm Reeve = C. australe Sowerby. "1939 Adam
& Leloup: Cardium pulchrum Reeve = Laevicardium
(Laevicardium) australe (Sowerby 1840). "1952
Kuroda & Habe: pulchntm Reeve = Fulvia australis
(Sowerby). "1977 Fischer-Piette: Cardium pulchntm
Reeve = Papyridea papyraceum (Chemnitz) Schroter.
"1981 Koyama et al.: Cardium pulchrum Reeve, 1845
= Fulvia australe (Sowerby, 1840). "1994 Vidal: ?
Cardium pulchrum Reeve, 1845 = Fulvia (Fulvia)
australis (Sowerby, 1834). "1999 Higo et al.: Cardium
pulchrum Reeve, 1845 = Fulvia australis (Sowerby,
1841). ··2003 Hylleberg & Kilburn, p. 178: Cardium
pulchrum Reeve. Synonym of Fulvia australis
(Sowerby, 1834). - CHARACTERS AND OCCURRENCE - OF COLLECTIONS. "ZMUC: one individual =
australe according to H. Lynge. DESCRIPTIONS. "1845
Reeve: shell ovate, somewhat attenuated towards the
umbones; radiately very finely grooved; fleshy brown,
white towards the margins, prettily ornamented with
square rose spots , umbones purple, interior
concentrically variegated with rose. REMARKS. "1845
Phuket Marine Biological Center Special Publication vol. 30 (2004)
725
pulchrum - punctatum
Reeve: this interesting species is not only distinct from
Cardium australe in form, but in sculpture and
colouring. DISTRIBUTION. China (Reeve 1845); China
(Grasset 1884).
pulchrum Martens, 1869 (variety). - Infrasubspecific.
» Regarding Cardium var. pulchrum: Cardiidae.
Recent. TYPE LOCALITY. Seychelles. RECORDS. "1869
Martens, pI. 3, figs . 2a--c: Cardium var. pulchrum (in
between pulchrum and rugatum). ··1869 ZR: Cardium
pulchrum var., allied to rugatum (Gronov.) from
Zanzibar [Ref. Martens 1869).
pulicarium Reeve, 1845. [Figs. Part I p. 142] . Synonym of Acrosterigma elongalUm (Bruguiere,
(789). OTHER JUDGEMENTS: Trachycardium pulicarium
Reeve, 1845. » Regarding Cardium pulicarium:
Cardiidae. Recent. TYPE LOCALITY. Unknown (Reeve
1845). TYPE REPOSITORY. Unknown (Vidal 1993).
RECORDS. ··1845 Reeve, fig. 102: Cardium pulicarium.
··1858 Adams & Adams, p. 455: Cardium
(Trachycardium) pulicarium, Reeve. ··1890 Paetel:
Cardium pulicarium R v. ··1922-32 Sherborn, p. 5238:
Cardium pulicarium Reeve, 1845. ··1972 Cernohorsky:
Vasticardium pulicarium (Reeve, 1845). ··1977
Fischer-Piette: Laevicardium (Vasticardium)
pulicarium Reeve. ··1993 Vidal: ? Cardium pulicarium
Reeve, 1845 = Acrosterigma elongatum (Bruguiere,
1789). ··1993 Vidal: Cardium pulicarium Reeve, 1845
=
Acrosterigma elongata = Trachycardium elongatum
(Bruguiere, 1789). - CHARACTERS AND OCCURRENCE - DESCRIPTIONS. ··1845 Reeve, shell somewhat
orbicular, Pecten-shaped, radiately ribbed, 34 ribs, the
anterior prickly, the middle crenulated on both sides,
and the posterior crenated; yellowish or fleshy white,
stained with scattered blackish brown spots, interior
pinkish white, margins rose-purple towards the hinge.
··1972 Cernohorsky: 35-37 ribs, prominently crenulate
on each side, spinose on the posterior 5-6 ribs.
DIMENSIONS. 25 mm. REMARKS. ··1845 Reeve: a
characteristic little shell, which I cannot refer to any
described species. ··1993: Vidal suggested that the shell
described by Reeve could be a juvenile Acrosterigma
elongata (Bruguiere, 1789). DISTRIBUTION. ··1845
Reeve: unknown. ··1994 Hylleberg: Indo-Pacific.
pullatum Stoliczka, 1871. - Combination to be revised:
"Cardium" pullatum (Stoliczka, 1871). Cardiidae.
Cretaceous. TYPE LOCALITY. Vylapaudy and Anapaudy,
South India. RECORDS. ··1871 Stoliczka, p. 218, pI. 11,
figs. 8-10: Cardium (Acanthocardium sic) pullatum
Stoliczka. ··1980 Ruhoff: Cardium (Acanthocardium
sic)pullatum Stoliczka, J 870; [1870-1871] vol. 3: 218,
pI. 11, figs. 8-) O. •• )983 Gliozzi & Malatesta: Cardium
pullatum Stoliczka, 1871.
pulvinatum Lam . - Combination to be revised:
"Cardium" pulvinatum Lam. Cardiidae. Fossil. ··1844
Potiez & Michaud: Cardium pulvinatum Lam. ex fide
Mich.
pumila (variety). - See under lamarckii pumila Friedel,
1870.
punctatostriatmn Lycett, 1850. - Combination to be
revised: "Cardium " punctatostriatum .. Lycett, 1850.
Cardiidae. Fossil. RECORDS . •• ) 922-32 Sherborn, p.
5255: Cardium punctatostriatum Lycett, 1850 [as
punctato-striatum]. ··1980 Ruhoff: Cardium
punctatostriatum Lycett, 1850; vol. 6 : 423.
punctatum Brocchi, 1814. - Identical with Parvicardium
papillosum (Poli, 1791). OTHER JUDGEMENTS:
Parvicardium minimum (Philippi). - Cardium planatum
var. - Cardium roseum Lamarck. - Parvicardium
scab rum (Philippi, 1844). » Regarding C. punctatum:
Cardiidae. Fossil and Recent. RECORDS. ··1814.
Brocchi: Cardium punctatum. ··1832 Bronn: Cardium
puncta tum Brch. 666 = ?variety of Cardium planatum.
··1836 Philippi : Cardium punctatum Broc. p. 666. t.
16. fig. II. = Cardium papillosum Polio ··1844 Philippi:
Cardium punctatum Brocchi. ··1848 Requien: Cardium
punctalum Brocchi. ··1857 Deshayes: Cardium
puncta tum Brocchi . ··1866 Brusina: Cardium
punctalUm Brocchi . ··1867 Weinkauff: Cardium
punctatum Brocchi = Cardium papillosum Poli. ··1868
Colbeau : Cardium punctatum Brocchi = Cardium
nodosum. ··1869 Romer: Cardium puncta tum Brocchi,
an varietas ? (a good species, Romer, p. 34). "1880
Stossich : Cardium punctatum Brocchi. 1814 = Cardium
nodosum Montagu . ··1888 Kobelt : ?Cardium
punctatum Brocchi = Cardium minimum Philippi.
··1888 Kobel!: Cardium punc/a/um Brocchi = Cardium
papillosum Poli. ··1891 Danilo & Sandri: Cardium
puncta/um Broce. ··1899 Sacco: Cardium punc/atum
Brocchi 1814 = Papillicardium papilloswri (Poli 1791).
··1902 Dollfus & Dautzenberg: Cardium punc/atum
Brocchi, 1814 = Cardium (Parvicardium) papi/losum
Poli, 1791. ··1912 Dautzenberg & Fischer: C.
punctatum Brocchi is a synonym ofC. papillosum Poli.
··1922-32 Sherborn, p. 5259: Cardium punctatum
Brocchi, 1814 ··1977 Fischer-Piette: Cardium
punc/alum Brocchi = Corculum papi/losum Poli ref.
"1983 Anaya: Cardium punc/a/um Brocchi, 1814 =
Parvicardium papillosum (Poli, 1791). ··2000 Aartsen
& Goud: Cardium punctatum Brocchi, 1814 (?
misidentified). - CHARACTERS AND OCCURRENCE - REMARKS. According to Aartsen & Goud
(2000), Cardium punctatllln is probably a
misidentification of a worn Parvicardium papi/losum
(Poli, 1795). HABITAT. Aegaean Sea depth range 12
fathoms (Forbes 1844). DISTRIBUTION. Guernsey
(Grasset 1884); Mediterranean (Locard & Caziot 1900);
Europe, North West Africa (auct.).
punctatum of Forbes. - Identical with Cardium roseum
Lamarck. » Regarding C. punctatum: RECORDS. ··1867
Weinkauff: ? Cardium puncta/um Forbes = Cardium
roseum Lamarck.
punctatum of Monterosato 1891.- Identical with
Parvicardium minimum (Philippi 1836). » Regarding
C. punctatum: RECORDS. ··1891 Monterosato: Cardium
puncta/um. ··1912 Dautzenberg & Fischer: Cardium
puncta/um Monterosato 1891 (non Brocchi) =
Parvicardium minimum (Philippi 1836).
punctatum of Philippi, 1844. - Identical with
Parvicardium scabrum (Philippi, 1844) . OTHER
JUDGEMENTS: Cardium nodosum Turton , 1822. Parvicardium papi/losum (Poli). » Regarding C.
punc/atum: RECORDS . ··1856 Jeffreys : Cardium
punctatum and variety Cardium scabrum Ph . =
Cardium papillosum (Poli). ··1884 Grasset: Cardium
punctalum Phil. ··1890 Paetel: Cardium punc/atum PhI.
= nodosum Mtg. ··1933 Coen: Cardium punc/a/um
Philippi = Eucardium (Cardium) echinatum var. roseum
Lamarck. ··1937 Coen: punc/atum Phil ippi =
726
Tropical Marine Mollusc Programme (TMMP)
punctatum - pllsillum
Ceras/oderma (Rudicardium) nodosum Turton.
punctatum of Renier. - Identical with Parvicardium
scabrnm (Philippi, 1844).» Regarding C. punc/atum:
RECORDS. --1844 Forbes: Cardium punctatum Ren.
punctatum of Requien. - Identical with Parvicardium
scabnlm (Philippi, 1844). OTHER JUDGEMENTS: Cardium
nodosum Turton, 1822. - Parvicardium scabrum
(Philippi, 1844).» Regarding C. punctatum: REcORDs.
--1886 Locard: Cardium puncta/um Requien, 1848. =
Cardium roseum de Lamarck, 1818. --1888 Kobelt:
Cardium puncta/um Requien = Cardium nodosum
Turton. --1900 Locard & Caziot: Cardium puncta/um
Requien = Cardium nodosum Turton, 1822.
punctatum of Sowerby. - Identical with "Cardium "
nodosum Turton , 1822. RECORDS. --1868 March:
Cardium puncta tum " Brocchi" Sow. = Cardium
(Cerastoderma) nodosum Turt.
punctatum ofTapparone-Canefri 1869. - Identical with
Parvicardium minimum (Philippi 1836). » Regarding
C. punctatum: RECORDS. --1869 Tapparone-Canefri:
Cardium puncta/um. --1912 Dautzenberg & Fischer:
Cardiumpunc/atum Tapparone-Canefri (non Brocchi)
= Parvicardium minimum (Philippi 1836).
puncto!il/eatum Healy & Lamprell, 1992. Acrosterigma punctolineatum Healy & Lamprell, 1992.
Cardiidae. Recent. TYPE LOCAL/TY. Holotype, Little
Trunk Reef, N. Queensland, Australia. TYPE REPOSITORY.
Holotype, Queensland Museum, Brisbane . TYPE
MATERIAL. Holotype, a paired specimen from Little
Trunk Reef, N. Queensland, QM ref. M032905 .
Paratypes: Four paired specimens from N. Queensland,
in AMS. RECORDS. --1992 Healy & Lamprell :
Acros/erigma punc/olineatum . --1999 Vidal :
Acros/erigma punctolineata Healy & Lamprell , 1992 :
87, pI. 3, figs. e-h. --200 I Vidal, p. 407: Acrosterigma
punctolineatum. - CHARACTERS AND OCCURRENCE - CHARACTERS of diagnostic value: the coloured
spots, riblets in interstices. DESCRIPTIONS. --1999 Vidal:
Shell ovoid, equilaterally (only anterior dorsal margin
slightly raised) to transversally (posterior margin
expanded) but not posteriorly truncated, moderately
elongated, and variably compressed. Lunule of medium
size on right valve, with its umbonal margin raised and
encroaching on left valve, where lunule is almost non
existent. External colour beige with small, slightly
elongated brown spots, limited to smooth anterior part
of ribs on posterior slope, and to rib tops on other parts
of shell; on posterior slope these spots are longer and
darker than elsewhere. Interior shows the same features
by transparency. Hinge symmetrical and moderately
angled. Mean rib number 48.9 ± 3.3, range 42-56. On
posterior slope, ribs flat with thin interstices; anterior
smooth part of ribs narrower than posterior part and
terminating abruptly against interstice. Axial furrow
relatively wide and readily visible; oblique scales
slightly elongated, regular in shape and regularly
disposed in axial furrow, encroaching upon posterior
part of rib, but usually not upon anterior part. On median
part of shell, ribs roundly triangular to rounded, slightly
asymmetrical, and generally smooth, except for those
closest to posterior slope, which sometimes bear
numerous long ridges on posterior face; on some young
shells, small rounded tubercular pustules randomly
scattered on rib tops; interstices narrower than ribs,
bearing a well-delineated small riblet in bottom, and
finely notched in places on adult parts. Anteriorly, ribs
become retr04:renulated, then top-ridged (wide ridges,
somewhat imbricated, becoming even wider and
irregular approaching lunule). --2001 Vidal: tentacles
numerous. No eyes on tentacles. DIMENSIONS. --1999
Vidal: length 29.9, height 35.7, width 24.0 mm.
REMARKS. Vidal (1999) : Acrosterigma puncto!ineatum
has typical characters and cannot be confused; it is close
to the next species A. hobbsae, but can easily be
separated from it. HABITAT. Recorded from 10 to 38 m
depth . DISTRIBUTION. --1992 Healy & Lamprell:
Philippines , North East Australia . --1999 Vidal :
recorded only in the eastern part of the western Pacific,
in littoral , reef environment. The species has been cited
and figured from Taiwan by Kuroda (1941: 161, pI. 5,
fig. 74,) and from Okinawa by Kubo & Kurozumi
(1995: 18J , fig. 3).
Plll/ctulaium Broderip & Sowerby, 1829. [Figs. Part I
p. 107]. - Combination to be revised: "Cardium"
punctulatum Broderip & Sowerby, J829. Cardiidae.
Recent. RECORDS. --1829 Broderip & Sowergy, p. 366:
Cardium punctulatum. --1834 Sowerby, 47t part, fig.
7: Cardium punc/ula/um . --1840 Sowerby, catalogue
p. 2, No. 20: Cardium punc/ulatum Brad. & Sow.
-- 1840 Sowerby, corrected list , p. 8: Cardium
punc/ula/um (Sowerby No. 20). --1841 Reeve: Cardium
punctula/um Brod. & Sow. --1843 Hanley: Cardium
punc/ulatum Brod. --1851 MacAndrew : Cardium
punctulatum Broderip & Sowerby. --1922-32 Sherborn ,
p. 5282: Cardium punctulatum Broderip & Sowerby,
1829. --1977 Fischer-Piette: Cardium punctula/um
Broderip & Sowerby. - CHARACTERS AND
OCCURRENCE - DISTRJBUTION . --1841 Sowerby: not
known. --1851 MacAndrew: Gibraltar.
punicum Coquand, 1862. - Unknown species. »
Regarding Cardium punicum: Cretaceous. TYPE
LOCAL/TY. Constantine. RECORDS. --1871 Stoliczka:
Cardium punicllm Coquand, 1862 (species insufficiently known). --1980 Ruhoff: Cardium punicum
Coquand, 1862; vol. 2: 208, pI. II , figs. 10-11.
punjabensis Eames, 1951 . - Discors punjabensis Eames,
1951. Cardiidae. Eocene. RECORDS. -- J 951 ZR: Discors
punjabensis Eames, 1951 sp. nov. Eocene Pakistan.
pllrbeckense Loriol & Jaccard , 1865 . - Cardium
purbeckense Loriol & Jaccard , 1865 . Cardiidae.
Jurassic and Cretaceous. RECORDS. --1985 Fiirsich &
Werner: Cardium purbeckense Loriol & Jaccard, 1865.
purpurea Martyn, 1784. Non binominal. - Not available.
« Regarding Cardium purpurea: Not a cardiid.
RECORDS. --1784 Martyn: Cardium purpurea.
pusil/a Chiplonker, 1939. - Pr%cardia pusilla
Chiplonker, 1939. Cardiidae. Cretaceous. --1939 ZR:
Protocardia pusilla Chiplonker, 1939 sp. n. Bagh Beds
Cretaceous S. India.
*pusillum . - Homonymy to be noticed. Cardium
pusillum from 2 sources: Gregorio, 1930 . Anrezjowski (?year).
pusillulII Anrezjowski. Combination to be revised:
"Cardium " pus ilium Anrezjowski. Cardiidae. Fossil.
R ECOR DS. --1857 Deshayes: Cardium pusi/lum
Anrezjowski.
pusi/lum Gregorio, 1930. - Combination to be revised:
"Cardium" pusillum Gregorio, 1930. Cardiidae. Lias.
727
Phuket Marine Biological Center Special Publication vol. 30 (2004)
pustulosum - quadragenarium
··1936 ZR: Cardium pusillum Gregorio, 1930 sp. n.
Lias Palermo.
puslulosum (afT.) Munster, 1836. - Granocardium afT.
pustulosum (Munster, 1836). Cardiidae. Cretaceous.
RECORDS. ··1986 Abdel-Gawad, pI. 40, fig. 8:
Granocardium (Granocardium) aff. pustulosum (v.
Munster, 1836).
pustulosum Munster in Goldfuss, 1837. - Combination
to be revised: "Cardium" pustulosum Munster in
Goldfuss, 1837. Cardiidae. Cretaceous. RECORDS.
··1871 Stoliczka: Cardium puslulosum (Acantho cardium sic). "1922-32 Sherborn, p. 5317: Cardium
pustulosum Muenster in Goldfuss, 1837.
putcaradze Gadzhiev, 1968 . - Didacna putcaradze
Gadzhiev, 1968. Lymnocardiidae. Tertiary. ··1968 ZR:
Didacna putcaradze Gadzhiev, 1968 sp. nov. Caspian,
Baku archipelago.
pulzeysi Glibert, 1933. - Laevicardium pulzeysi Glibert,
1933. Cardiidae. Upper Eocene. RECORDS. ··1936 ZR:
Laevicardium pUlzeysi (E. Vincent MSS) Glibert, 1933
sp. n. Wemmel Sands, Eocene Belgium. ··1970 Glibert
& Poel: Laevicardium putzeysi (E. Vincent). ··1970
Glibert & Poel: Pratulum putzeysi.
pycnosora Kol. - Hyrcan ia PYCl1osora (Ko!.).
Lymnocardiidae. Fossil. RECO RDS. ··1977 Popov :
Hyrcania Pycl1osora (Kol.).
pygmaeum Donovan, 1799. - Synonym of Parvicardium
exiguum (Gmelin, 1791). OTHER JUDGEMENTS: Cardium
siculum Sowerby, 1839. - Cardium stellatum Reeve,
1844. » Regarding Cardium pygmaeum: Cardiidae.
Recent. REMARKS. Parvicardium exiguum (exiguum)
(Gmelin, 1791) = pygmaeum (Donovan, 1799) =
siculum (Sowerby, 1839) = stellatum (Reeve, 1844) in
Nordsieck (1969). Habitat: C. exiguum: littoral and
algal zone, Orkney Isles. (Nordsieck 1969).
DISTRIBUTION. C. exiguum: North Europe,
Mediterranean, Black Sea (Nordsieck 1969). RECORDS.
"1800 Donovan, PI. 9, fig. 4a Cardium pygmaeum.
"1803 Montagu: Cardiumpygmaeum Don. Br. Shells.
i. t. 32. f. 3. = Cardium exiguum Gmel. 37. ··1817
Dillwyn: Cardium pygmaeum Solander 's MSS =
Cardium exiguum Gmelin. ··1844 Brow n: Cardium
pygmaeum of Donovan = Cardium exiguum. ··1846
Forbes: Cardium pygmaeum Donovan = Cardium
exiguum Gmelin. ··1850 Jay: Cardium pygmaeum
Donov. = Cardium exiguum. ··1852 Leach: Cardium
pygmaeum Donov. = Cardium exiguum. ··1856 Wood
& Hanley: C. pygmaeum Don. pI. 32, f. 3. = Cardium
exiguum Mont. ··1867 Hidalgo: Cardium pygmaeum,
Donovan = Cardium exiguum Gmelin. ··1867
Weinkauff: Cardium pygmaeum = Cardium exiguum
Gmelin. ··1868 Colbeau: Cardium pygmaeum Donov.
··1869 Romer: Cardium pygmaeum Donovan =
Cardium exiguum Gmelin. "1871 March, p. 215, No.
205 Cardium pygmaeum Don. = Cardium exiguum Gm.
··1882 Cooke: Cardium pygmaeum Don . ··1882
Cooke : Syn. Cardium l1odosum Turt. labeled C.
pygmaeum Don. (= C. exiguum Gmel.). ··1886
Chevreux: Cardium pygmaeum Donov. ··1886 Locard:
Syn. Cardium pygmaeum, Donovan, 1804 = Cardium
exiguum, Gmelin, 1789 sic. ··1888 Kobelt: Syn.
Cardium pygmaeum Donovan = Cardium exiguum
Gmelin. ··1888 Servain: Cardium pygmaeum Donovan,
Forbes & Hanley, Sowerby = Cardium exiguum. ··1890
Hidalgo: Cardium pygmaeum Donovan = Cardium
exiguum Gmelin. ··1890 Paetel : Cardium pygmaeum
Donov. = exiguum Gm. ··1902 Sherborn, p. 80 I:
Cardiumpygmaeum Donovan, 1800. "1917 Hidalgo:
Syn. Cardium pygmaeum Donovan = Cardium exiguum
Gmelin. ··1932 Winckworth : Syn. Cardium pygmaeum
= Cardium (Parvicardium) exiguwn Gmelin, 1791.
··1962 Grossu: Syn. Cardium pygmaeum Donovan =
Cardium (Parvicardium) exigllum Gmelin, 1790.
··1977 Fischer-Piette: Syn. Cardium pygmaeum
Donovan = Cardium exiguum Gmelin. ··2000 Aartsen
& Goud: Cardium pygmaeum Donovan, 1800 is a
synonym of Parvicardium exiguum (Gmelin, 1791).
pygmaeum Philippi , 1860. -Combination to be revised:
"Cardium" pygmaeum Philippi, 1860. Cardiidae.
Fossil. RECORDS. ··1980 RuhofT: Cardium pygmaeum
Philippi, 1860; p. 176.
*pygmaeul1l; Conocardiidae . - Homonymy to be
noticed. COl1ocardium pygmaeum from 2 sources:
Hisinger, 1837. - Isberg, 1934.
pygmaeul1l Hisinger, 1837. - Conocardium pygmaeum
Hisinger 1837. Conocardiidae. Fossil. ··1942 ZR:
Conocardium pygmaeum Hisinger 1837 [Ref. Branson
1942].
pygmaeum Isberg 1934. - Replaced by COl1ocardium
isbergi Branson, 1942. Conocardiidae. Fossil. RECORDs.
··1942 ZR: Conocardium pygmaeum ]sberg 1934
replaced by Conocardium isbergi Branson, 1942.
pyramidatum Grimm, 1877. [Figs. Part I p. 180]. Changed combination: Didacna pyramidatum (Grimm,
1877). Lymnocardiidae. Fossil. TYPE LOCALITY. Caspian
Sea. RECORDS. ··1877 Grimm, p. 46, pI. 8, figs. I a-d:
Cardium pyramidalum sp. n. ··1877 ZR: Cardium
pyramidatum Grimm, 1877 sp. n. Caspian Sea. ··1968
Logvinenko & Starobogatov: Didacna pyramidata
(Grimm). ··1987 Taktakishvili : Didacna pyramidata
(Gr.).
pyramidum Abbass, 1978. - Cardium pyramidum
Abbass, 1978. Cardiidae. Upper Eocene. RECORDS.
··1978 ZR: Cardium (Cardium) pyramidllm Abbass,
1978 sp. nov. Egypt. Upper Eocene. Ref. to Abbass p.
155.
pyrijorme d'Orbigny, 1850. - Conocardium pyriforme
d 'Orbigny, 1850. Conocardiidae. Fossil. RECORDS.
··1922-32 Sherborn, p. 4990: Lunulacardium pyriforme
Muenster, 1840. "1922-32 Sherborn: Conocardium
(Muenster) pyriforme d'Orbigny, 1850.
LUllulacardium.
pyrocostatul1l (aff.) Hal. - New combination:
Lymnocardium aff. pyrocostatum Hal. Lymnocardiidae.
Fossil. RECORDS. ··1987 Taktakishvili : Limnocardium
aff. pyrocostatum Hal.
pyrophila (variety). - See under sjoegreni pyrophila
Andrussov, 1924.
Q
quadragenariul1l Conrad, 1837. - Changed combination:
Trachycardium quadragenarium (Conrad, 1837).
Cardiidae. Fossil and Recent. Type species: Stewart
(1930) designated Cardium quadragenariwn Conrad,
1837 type species of the subgenus (Dal/ocardia)
Stewart, 1930. Subsequently raised to generic status.
RECORDS. "1837 Conrad: Cardium quadragenarium.
728
Tropical Marine Mollusc Programme (TMMP)
quadragenarium - quadripartitum
--1860 Carpenter: Cardium quadragenarium Conr.
--1864 Carpenter: Cardium quadragenarium Conr. =
luteolabrum (= xanthocheilum) Gld. --1890 Paetel:
Cardium quadragenarium Conr. --190 I Dall: Cardium
(Trachycardium) quadragenarium Conrad, 1837.
--1918 Chase p. 103: Cardium quadragenarum [err.
pro quadragenarium] Conr. --1921 Dall:
Trachycardium quadragenarium Conrad, 1837. --192232 Sherborn, p. 5353: Cardium quadragenarium
Conrad, 1837 . "1924 Oldroyd: Cardium quadragenarium Conrad, 1837. "1930 Stewart: Cardium
quadragenarium Conrad, 1837. --1930 ZR:
Dal/ocardia Stewart, 1930 subgen. n. of Trachycardium
type Cardium quadrigenerium Conrad, fossil. "1937
Keen: Trachycardium quadragenarium. --1944 Keen
& Bentson: Cardium quadragenarium Conrad. --1954
Keen: Trachycardium (Dal/ocardia) quadragenarium.
--1960 Morris : Trachycardium quadragenarium
(Con.). --1969 Moore et at.: Trachycardium
(Dallocardia) quadragenarium (Conrad, 1837). --1977
Popov: Trachycardium (Dal/ocardia) quadragenarium
(Conrad, 1837). "1977 Fischer-Piette: Laevicardium
quadragenarium Conrad. --1977 p. 46. Popov:
Cardium quadragena rium Conrad, 1837. Type of
(Dallocardia) Stewart, 1930. ··1977 Wilson &
Stevenson: Cardium quadragenarium Conrad, 1837.
-·1983 Anaya: Type: Cardium quadragenarium
Conrad, 1837. Type species (D.O.) of (Dallocardia)
Stewart, 1930. ··1995 Poutiers: Trachycardium
quadragenarium (Conrad , 1837). "1999 Vidal:
Trachycardium quadragenarium (Conrad , 1837).
"1999 Vidal: "Cardium "quadragenarium. ··2000
Coan et al.: Trachycardium(Dallocardia) quadragenarium (Conrad, 1837) ... 200 I Schneider & Carter:
Trachycardium quadragenarium (Conrad, 1837). CHARACTERS AND OCCURRENCE - OF COLLECTIONS. --ZMUC: one old shell from Catalina Island,
California. REMARKS. --1999 : Vidal cites Conrad 1837,
p. 230: Cardium quadragenarium being "allied to
Cardium asiaticum". DISTRIBUTION. --ZMUC
California.
quadragetlariulII (aff.) Conrad, 1837. - RECORDS. "1907
ZR: Cardium aff. quadrigenarium sic [Ref. Arnold
1907b].
quadragenariumfernandoensis Arnold, 1907 (variety).
- Synonym of Trachycardium quadragenarium
(Conrad, 1837). »
Regarding Cardium
quadragenarium var.fernandoensis: Cardiidae. Fossil.
RECORDS. --1907 ZR: Cardium quadrigenarium sic var.
fernandoensis Arnold, 1907 var. n. Middle Fernando
formn. S. California [Ref. Arnold 1907a]. "1944 Keen
& Bentson: Cardium quadragenarium Conrad var.
fernandoensis Arnold, 1907 (n. var.). ·-2000 Coan et
al.: C. quadragenarium jernandoensis Arnold, 1907 =
Trachycardium (Dallocardia) quadragenarium
(Conrad,1837).
quadragenarum (error). - Printer's mistake for
quadragenarium in Chase 1918.
quadrans Cragin, 1894. - Synonym of Protocardiafilosa
(Conrad, 1857). » Regarding ROt/dairia quadrans:
Cardiidae . Cretaceous. RE CORDS. --1986 Scott:
Roudairia quadrans Cragin, 1894 = Protocardia filosa
(Conrad, 1857). --1987 ZR: Protocardia quadrans
(Cragin, 1894) Junior synonym of P. filosa (Conrad,
1857). Ref. to Scott p. 1203.
quadralls W.B. & H.D. Rogers, 1839. - Combination to
be revised: "Cardium" quadrans W.B. & H.D. Rogers,
1839. ?Cardiidae. Fossil. RECORDS. --1922-32 Sherborn,
p. 5354: Cardium quadrans W.B. & H.D. Rogers, 1839.
quadrans. - Conocardium quadrans. Conocardiidae.
Fossil. RECORDS. "1889 ZR: Conocardium quadrans
with fig. (ref. Barrois).
quadrariuIII Reeve, 1845. [Figs. Part I p. 145. Part 3 p.
933]. - Parvicardium quadrarium (Reeve, 1845).
OTHER JUDGEMENTS: Cardium exiguum Gmelin, 1791.
» Regarding Cardium quadrarium: Recent, Cardiidae.
TYPE LOCALITY. Unknown (Reeve 1845). TYPE
REPOSITORY. BMNH. holotype reg. no. 1978141. Neg.
No. 0392. DISTRIBUTION. Unknown (Reeve 1845).
RECORDS ... 1845 Reeve: Cardium quadrarium ... 192232 Sherborn, p. 5354: Cardium quadrarium Reeve,
1845. "1977 Fischer-Piette: Cardium quadrarium
Reeve = Cardium exiguum Gmelin. - CHARACTERS
AND OCCURRENCE - --1845 Reeve: Shell thin,
squarely ovate, posteriorly slightly concavely
angulated; radiately ribbed, ribs flattened, here and
there prickly scaled, those of the posterior area smooth,
interstices between the ribs rather deeply cut; whitish,
the concave posterior area pale rusty brown. REMARKS.
·-1845 Reeve: the chief peculiarities of this shell are
its flattened ribs, the interstices between which are
narrowly and deeply cut, and its posterior concavity.
DISTRIBUTION. -·1845 Reeve: unknown.
quadrata K valiashvili, 1978. - Digressodacna quadrala
Kvaliashvili, 1978. Lymnocardiidae. Pliocene.
RECORDS . --1978 ZR: Digressodacna quadrata
Kvaliashvili, 1978 sp. nov. Georgia SSR. Ref. to
Kvaliashvili p. 73.
quadrata (subspecies). - See under edule quadratum
Bucquoy, Dautzenberg & Dollfus, 1892.
*quadratlllll. - Homonymy to be noticed. Cardium
quadratum from 2 sources: Deshayes, 1853. Doncieux, 1912.
quadratum Chen, 1974. - Changed combination :
Myophoricardium quadratum (Chen, 1974). Cardiidae.
Fossil. RECORDS. ··1976 Ma et al. p. 260: Myophoriopis
quadralum Chen, 1974. --1976 Ma et al. p. 260:
Myophoricardium quadratum (Chen, 1974).
qlladratulII Deshayes, 1853. - Combination to be
revised: "Cardium" quadratum Deshayes, 1853.
Cardiidae. Fossil. RECORDS. "1857. Deshayes, p. 53:
Cardium quadratum (Germany) new name for Cardium
multicostatum in Goldfuss. --1980 Ruhoff: Cardium
quadratwn Deshayes, 1853; vol. 2: 48.
quadratuIII Doncieux, 1912. - Combination to be
revised: "Cardium" quadratum Doncieux, 1912.
Cardiidae. Eocene. RECORDS. "1912 ZR : "Cardium"
quadratum Doncieux, 1912 sp. n. France Eocene.
qlladricristatum Petho, 1906. - Combination to be
revised: Combination to be revised: "Cardium"
quadricristatum Petho, 1906. Cardiidae. Cretaceous.
TYPE LOCALITY. Hungary. RECOR DS. "1906 Petho :
Cardium quadricrislatum sp. n. -·1906 ZR: Cardium
quadricristatum Petho , 1906 sp. n. Cretaceous,
Hungary.
quadrigenarium (error).- Printer 's mistake for
quadragenarium in Arnold 1907.
quadripartitum Kolesnikov, 1929. - Cerastoderma
Phuket Marine Biological Center Special Publication vol. 30 (2004)
729
quadripartitum protractijormis - radiatil
quadripartitum (Kolesnikov, 1929). Cardiidae.
Sarmatian. RECORDS. ··1929 ZR: Cardium
quadriparlitum Kolesnikov, 1929 sp. n. Sarmatian.
··1977 ZR : Cerasloderma (Obsoletijorma) quadripartitum Key to subspecies. Ref to ll'ina, Nevesskaya
& Paramonova p. 76.
quadripartitum protractiformis Andrus (variety). Upgraded.
Cerastoderma
quadripartitum
profractiformis (Andrus.) Cardiidae. Sarmatian.
RECORDS. ··1929 ZR: Cardium quadripartitum var.
prolracliformis Andrus., Sarmatian [Ref. Kolesnikov
1929].
quadripartitum squamata Andrus (subspecies) . Cerasloderma quadripartifum squamala (Andrus.)
Cardiidae. Sarmatian. RECORDS. ··1929 ZR : Cardium
quadripartitum squamata Andrus. , Sarmatian [Ref.
Kolesnikov 1929].
quedongensi Sherrard , 1960. - Lunulicardium
quedongensi Sherrard, 1960. Not in Cardiacea. Silurian.
RECORDS. ··1960 ZR: Lunulicardium quedongensi
Sherrard, 1960 sp. nov. Quedong near Bombala, N.S.
Wales, Silurian.
quilonense Dey, 1962. - Combination to be revised :
"Cardium" quilonense Dey, 1962. Cardiidae. Miocene.
RECORDS. ··1962 ZR: Cardium (s.l.) quilonense Dey,
1962 sp. nov. Indo-Pacific Province, Miocene.
quinarium Marwick, 1944. - Changed combination:
Pratulum quinarium (Marwick, 1944). Cardiidae.
Opoitian. RECORDS. ··1944 ZR: Nemocardium
(Pratulum) quinarium Marwick, 1944 sp. nov. roadside
just E. of Mangatarchu Stream, Waikohu Surv. Dist.
(Opoitian). ··1977 Popov: Pralulum quinarium
(Marwick,1944).
quinquecostatum Douville, 1917. - Granocardium
quinquecostatum Douville, 1917. Cardiidae. Cretaceous. RECORDS. ··1917 ZR: Granocardium quinquecostalum Douville, 1917 n. sp. Cretaceous Moghara.
quinquecostatum Miinster, 1840. - Combination to be
revised: "Cardium" quinquecoslatum Munster, 1840.
Cardiidae. Fossil. RECORDS . ··1922-32 Sherborn, p.
5398: Cardium quinquecostatum Muenster, 1840.
R
rachitis Deshayes, 1829. - Changed combination:
Goniocardium rachitis (Deshayes, 1829) .
Aviculariidae. Eocene. TYPE LOCALITY. Paris Basin.
RECORDS. ··1835 Lamarck : Cardium rachitis Desh.
··1839 Anton: Cardium rachites Deshayes. ··1886
Cossmann: Fragum rachitis (Deshayes). ··1887
Fischer: Goniocardium rachitis Deshayes. ··1922-32
Sherborn, p. 5407: Cardium rachitis Deshayes, 1829.
··1937 ZR: Cardium rachites Deshayes, 1829 type of
Goniocardium Vasseur, 1880 Prosodacnidae (?) not a
cardiid unit [Ref. Keen 1937]. ··1995 Le Renard et al.:
Goniocardium rachitis [Cardium rachitis] Deshayes,
1829. ··1995 Pacaud & Renard : Goniocardium rachilis
Superfamily Tridacnoidea . Family Aviculariidae.
··1998 Schneider (\ 998b): Goniocardium rachitis
(Deshayes, 1829). ··1999 Jacques Vidal (1999, pers.
com.) MNHN, Paris: Fossils from the Paleontological
Collection as both rachite Deshayes & rachitis
Deshayes. Bartonien. France (Bassin de Paris).
racketti Donovan, 1825. - Synonym of Fulvia lenuicosta
(Lamarck, 1819). OTHER JUDGEMENTS: Papyridea
papyraceum Schroter [not available]. » Regarding
Cardium rackelti: Cardiidae. Recent. TYPE LOCALITY.
New South Wales. RECORDS. ··1825 Donovan: Cardium
racketti. ··1922-32 Sherborn, p. 5407: Cardium
rackeltii Donovan, 1825. ··1938 Cotton & Godfrey:
Cardium racketti Donovan 1826. ··1945 Smith:
Cardium rackelt;; Donovan, 1825 = Cardium
tenuicostata Lamarck 1819. ··1945 Smith: Cardium
ra ckeltii ' Donovan' Hedley 1916 = Cardium
tenuicostata Lamarck 1819. ··1950 Allan: Cardium
rackelti. ··1961 Cotton: Fulvia rackelti Donovan 1826
= Fulvia fenuicostata Lamarck 1819; Pliocene, South
Australia . ··1961 Rippingale & MacMichael : Fulvia
racketti Donova n. ··1977 Fischer-Piette: Cardium
racketti Donovan = Papyridea papyraceum (Chemnitz)
Schroter. ··1977 Wilson & Stevenson: Cardium rackelti
Donovan , 1825, junior synonym of Fulvia
tenuicostatum (Lamarck, 1819). ··1992 Lamprell &
Whitehead: Cardium rackelti Donovan, 1925 = Fulvia
tenuicostala (Lamarck, 1819). ··1994 Vidal: Cardium
rackelti Donovan, 1826 = Fulvia (Fulvia) tenuicosta
(Lamarck, 1819). - CHARACTERS AND OCCURRENCE - REMA RKS. ··1961: Cotton stated that Fulvia
rackelti Donovan, 1826 is a living form of the Pliocene
Fulvia lenuicostala Lamarck 1819. DISTRIBUTION.
··1994 Hylleberg: South Australia.
radians Mayer, 1857. - Combination to be revised:
"Cardium" radians Mayer, 1857. Cardiidae. Fossil.
RECOR DS . ··1980 Ruhoff: Cardium radians Mayer,
1857; [1856-1857] vol. 5: 363.
radiata (mode). - See under radiatum Reeve, 1845
radiata Donovan, 1799. - Synonym of Serripes
groenlandicum (Bruguiere, 1789). OTHER JUDGEMENTS:
Cardium eden/ula. - Not syn. of Cardium tenuicostata
Lamarck, 1819. » Regarding Mactra radiata:
Cardiidae. Recent. RECORDS. ··1799 Donovan: Mactra
radiata. ··1800 Donovan, PI. 44. figs. I, 2 Mactra
radiata. ··1813 Fleming: Mactra radiata Brit. Shells.
tab. 161. = Cardium edentula. ··1850 Jay: Cardium
radialum Donov. = Cardium groenlandicum. ··1856
Wood & Hanley: MaClra radiata Donov. = Cardium
Groenlandicum Ch. [not available]. ··1868 Morch, no.
120 : Mactra radiata Don. = Cardium (Serripes)
gronlandicum Chemn. var solida Morch, 1868. ··1869
Romer: Mactra radiala Donovan = Cardium
Gronlandicum Chemnitz [not available]. ··1870 Gould:
Mactra radiata Donovan (syn.) = Aphrodite
groenlandica. ··1888 Kobelt: Mactra radiata Donovan
= Aphrodite groenlandicus. ··1890 Paetel : Cardium
radiatum Donov. = groenlandicum Ch . [not available].
··1902 Dall: Mactra radiata of Donovan, 1799 =
Serripes gronlandicus Gmelin, 1792. ··1912
Dautzenberg & Fischer: Mactra radiala Donovan 1800
= Cardium (Serripes) groenlandicum, (Chemnitz)
Bruguiere, Gmelin. ··1939 McLean: Cardium radiatllm
(Donovan, 1799) = Serripes groenlandicum (Bruguiere,
1789). ··1945 Smith: radiatum ' Donovan' Gray 1824
not synonym of Cardium tenuicostata Lamarck, 1819.
··1977 Fischer-Piette: Mactra radiata Donova n =
Serripes groenlandicwn (Chemnitz) Schroter. ··1980
ZR: Serripes radiala Donovan 1803 is a junior
synonym of S. groenlandicus groenlandicus (Bruguiere
1789). Ref. to Kafanov. ··2000 Coan et al.: Mactra
730
Tropical Marine Mollusc Programme (TMMP)
radiata - radula
radiata Donovan, 1803 = Serripes(Serripes)
groenlandicus (Mohr, 1786). - CHARACTERS AND
OCCURRENCE - DISTRIBUTION. Arctic regions.
radiata Martyn, 1789. Non binominal. - Not available.
» Regarding Cochlea radiata: not a cardiid (?Mactra).
RECORDS. --1789 Martyn: Cochlea radiata.
radiata Meek & Worthen. - ?Cardiopsis radiata Meek
et Worthen. ?Cardiacea. Devonian-Carboniferous.
RECORDS. --1887 Fischer: ?Cardiopsis radiata Meek
et Worthen.
radiata Stevanovic, 1951 . - Paradacna radiata
Stevanovic, 1951. Lymnocardiidae. Pontian. REcORDs.
--1953 ZR: Paradacna radiata Stevanovic, 1951 sp.
nov. Pontian Serbia.
*radiatum. - Homonymy to be noticed. Cardium
radiatum from 4 sources: Dujardin, 1837. - Koninck,
1842. - Reeve, 1845. - Svarovsky, 1971.
radiatum Dujardin, 1837. - Combination to be revised:
"Cardium" radiatum Dujardin, 1837. Cardiidae. Fossil.
RECORDS. "1857 Deshayes: Cardium radiatum. Duj.
1836 sic . "1922-32 Sherborn, p. 5411: Cardium
radiatum Dujardin, 1837. "1945 Smith: Cardium
radiatum Dujardin, 1837 is not a synonym of Cardium
tenuicostata Lamarck, 1819.
radiatum Gray, 1824 . - Synonym of Serripes
groenlandicum (Bruguieree 1789). RECORDS. --1922-32
Sherborn: Ma ctra radiata Donovan. --1922-32
Sherborn, p. 5411: Cardium (Don.) radiatum Gray,
1824. - Mactra.
radiatum Koninck, 1842. Replaced by "Cardium "
modiolaeforme. Deshayes. ?Cardiacea. Si lurian .
RE CORDS . --1857 Deshayes: Cardium radiatum
Koninck. 1842 (cardiomorpha). - C. modiolaeforme.
Desh.
radiatum Reeve, 1845 . [Figs. Part I p. 14 t. Part 3. p.
929] . - Synonym of Fulvia tenuicosta (Lamarck, 1819).
OTHER JUDGEMENTS: Fulviafagea Voskuil & Onverwagt,
1992. - Papyridea papyracea (Chemnitz) Schroter. Cardium tenuicostatum var. » Regarding Cardium
radiatum: Cardiidae. Recent. TYPE LOCALITY. Unknown
(Reeve 1845). TYPE REPOSITORY. BMNH; Cardium
radiatum Reeve Holotype reg. no. 1912.6.4.17. Neg.
No. 0382. RECORDS. --1845 Reeve: Cardium radiatum.
--1868 ZR: Cardium radiatum (Reeve) seems to be a
variety of tenuicostatum (Lam.) [Ref. Romer 1868].
--1869 Romer: Cardium radiatwn Reeve fig . 89 =
Cardium tenuicostatum Lamarck. --1877 Marrat :
Cardium (Fulvia) radiatum Reeve. "1885 Smith:
Cardium radiatum , (Reeve in Romer, fig. 89) =
Cardium (Bucardium) tenuicostatum Lamarck. --1889
Martens: Cardium radiatum Reeve (p. 26 : off Liberia,
68 m, black mud). --1889 Martens: Cardium radiatum
Reeve (p. 29: off Liberia, 179 m, dark brown sand).
"1890 Paetel: Cardium radiatum Rv. "1890 Paetel:
Papyridea radiata Rv. "1903 Pritchard & Gatliff:
Cardium radiatum Romer in Kuster 1869 = Cardium
tenuicostatum Lamarck. "1903 Hidalgo: Cardium
radiatum Reeve. 1845 . "1922-32 Sherborn, p. 5411:
Cardium radiatum Reeve , 1845. --1928 Faustino:
Cardium radiatum Reeve. "1945 Smith: Cardium
radiatum Reeve, 1843 syn. = Cardium tenuicostata
Lamarck , 1819. "1977 Fischer-Piette: Cardium
radiatum Reeve = Papyridea papyracea (Chemnitz)
Schroter. --1992 Voskuil & Onverwagt, p. 42: Cardium
radiatum Reeve, 1845 = Fulviafagea (nom. nov.). Nec
Gray 1820, nor Dujardin 1837, nor Svagrovsky 1971 .
--1994 Vidal: Cardium radiatum Reeve, 1845 = Fulvia
(Fulvia) tenuicosta (Lamarck, 1819). - CHARACTERS AND OCCURRENCE - OF COLLECTIONS.
--ZMUC: I specimen. Quartz sediment inside the shell
contained the foraminiferan Amphistegina, spine of sea
urchin, crab claw, small gastropods. No remains of
corals. (label in box: N. S. Wales coil. Comeliussen
1814). DESCRIPTIONS. --1845 Reeve: shell somewhat
squarely ovate, radiately rather obscurely ribbed, 52
ribs, nearly obsolete at the umbones; fleshy white,
concentrically banded with pale brown, stained within
and without at each umbone with a conspicuous purple
ray. REMARKS. "1845 Reeve: This species has been
probably confounded hitherto with the Cardium
tenuicostatum, from which it differs somewhat in form,
besides being of a more solid structure. DISTRIBUTION.
"1845 Reeve: unknown. --ZMUC: North Australia,
Solomon Islands; N. S. Wales.
radiatum Svarovsky, 1971. - Combination to be revised :
"Cardium" radiatum. Cardiidae . Upper Miocene.
RECORDS. --1971 ZR: Cardium radiatum Svagrovsky,
1971 sp. nov. Upper Miocene Ostslowakei,
Czechoslovakia.
*radiiferum. - Homonymy to be noticed. Cardium
radiiferum from 2 sources: Andrusov, 1902. Andrusov, 1963. REMARKS. I do not know how
Andrusov, 1902 compares with Andrusov, 1963.
radiiferum Andrusov, 1902. - Changed combination:
Andrusovicardium radiiferum (Andrusov, 1902) .
Lymnocardiidae. Pliocene. GENOTYPE. Nevesskaya et
al. (1986 p. 192) designated Cardium radiiferum
Andrusov, 1902 type species of genus Andrusovicardium Paramonova , 1986. Dollfus (1905)
commenting Andrussow 1902: Cardium radiifentm n.
sp. tends towards Cerastoderma. RECORDS. --1902
Andrusov: Cardium radiiferum. --1905 Dollfus:
Cardium radiiferum Andrussow, 1902. --1977 Popov:
Avicardium radiiferum. --1986 Nevesskaya et al.:
Andrusovicardium radiiferum (Andrusov, 1902).
--1987 ZR: Cardium radiiferum Andrusov, 1902.
Designated type species for new genus Andrusovicardium. Ref. to Nevesskaya et af. p. 192.
radiiferuin Andrusov, 1963. - Cardium radiiferum
Andrusov, 1963 . Lymnocardiidae. Miocene. RECORDS.
"1961 ZR: Cardium radiiferum Andrusov, 1961 "sp.
nov."[in a table] Miocene Caspian region, Russia.
--1963 ZR: Cardium radiifentm Andrusov, 1963 sp .
nov. Neogene, Russia.
radmanesti (variety). - See under chartacea var.
radmanesti Gillet, 1943 .
radobojanllin Anic, 1952. - Combination to be revised:
" Cardium" radobojanum Anic, 1952. Cardiidae.
Oligocene . RECORDS. --1952 ZR: Cardium
radobojanum Anic, 1952 sp. nov. Upper Oligocene,
Croatia.
*radula. - Homonymy to be noticed. Cardium radula
from 3 sources: Broderip & Sowerby, 1829. Deshayes, 1832. - Thiele & Jaeckel, 1931.
radula Broderip & Sowerby, 1829. - Changed
combination: Trachycardium radula (Broderip &
Sowerby, 1829). Cardiidae. Recent. RECORDS. "1829
Broderip & Sowerby, p. 366: Cardium radula. "1840
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731
radula - rarum
Sowerby, catalogue p. 3, No. 43: Cardium radula .
--1857 Carpenter: Cardium radula. --1872 Carpenter:
Cardium radula. --1922-32 Sherborn, p. 5419: Cardium
radula Broderip & Sowerby, 1829. --1977 FischerPiette: Cardium radula Broderip & Sowerby. CHARACTERS AND OCCURRENCE - REMARKS. In
general appearance C. radula resembles C. muricatum;
but it differs in shape, and in the ribs being angular at
both edges (Brode rip & Sowerby 1829). Sowerby
(1841) noted: We have not seen this shell. It is in Mr.
Bland's collection. Fischer- Piette (1977) found that the
description of radula would fit C. senticosum, but
refrained from specifying synonymy because Cardium
radula is so poorly known. DISTRIBUTION. California,
Peru.
radula Deshayes, 1832. [Figs. Part 3 p. 921]. - Synonym
of Cerastoderma eduie (Linnaeus, 1758). OTHER
JUDGEMENTS: Cardium nodosum Montagu. »
Regarding Cardium raduia: Cardiidae. Fossil. REcORDs.
--1832 Deshayes, p. 106, pI. 18, figs. 15- 17: Cardium
radula Desh. --1890 Paetel: Cardium raduia Dsh. =
nodosum Mtg. --1922-32 Sherborn, p. 5419: Cardium
radula Deshayes in Bory de St. v., 1833 . --1977
Fischer-Piette: Cardium radula Deshayes = Cardium
edule Linne.
radula Thiele & Jaeckel, 1931. - Synonym of
Afrocardium infantile (Nomura & Zinbo, 1834). OTHER
JUDGEMENTS: New name Laevicardium thielei FischerPiette, 1977.» Regarding Cardium raduia: Cardiidae.
Recent. TYPE LOCALITY. Dar es Salaam, E. Africa.
REMARKS. Laevicardium thielei referred to Ctenocardia
(Afrocardium) in Keen (1980). DISTRIBUTION. E. Africa.
RECORDS. --1931 Thiele & Jaeckel, p. 227, pI. 9, no.
99: Cardium raduia n. sp. --1931 ZR: Cardium radula
Thiele & Jaeckel, 1931 sp. n. --1977 Fischer-Piette:
Laevicardium thielei new name for Cardium radula
Thiele & Jaeckel, 1931. --1977 ZR: Cardium radula
Thiele & Jaeckel 1931 . Preoccupied name, replaced
by Laevicardium (Trachycardium) thielei. Ref to
Fischer-Piette, E. p. 55. --1981 Koyama et al.: Cardium
raduia Thiele & Jaeckel, 1931 = Cardium
(Trachycardium) infantile Nomura & Zinbo, 1834.
--1999 Higo et af.: Cardium raduia Thiele & Jaeckel,
1831 sic non Broderip & Sowerby, 1829 = Afrocardium
infantile (Nomura & Zinbo, 1934).
rahmuensis Cox, 1965 . - Protocardia rahmuensis Cox,
1965. Cardiidae. Oxfordian. RECORDS. --1965 ZR:
Protocardia rahmuensis Cox, 1965 sp. nov. Rahmu,
S.E. Kenya, Oxfordian.
rammaurtii Singh et af. - Protocardia rammaurtii Singh
et al., 1980. Cardiidae. Jurassic. RECORDS. --1980 ZR:
Protocardia (Protocardia) rammvrtii sic sp. nov. India.
Spelling of species also as P. (P.) rammaurtii. Ref. to
Singh et aI., p. 73.
rammvrtii (spelling) . - See under rammaurtii.
ranvillensis Palmer, 1973 . - Cryptocardia ranvillensis
Palmer, 1973 . Cardiidae. Upper Jurassic. RECORDS.
--1973 ZR: Cryptocardia ranvillensis Palmer, 1973 sp.
nov. France, Calvados, Ranville. Ref. to Palmer p. 170.
ranvil/ianum d'Orbigny, 1850. - Unicardium
ranvillianum d' Orbigny, 1850. Cardiidae. Fossil.
RECORDS. --1922-32 Sherborn, p. 5429: Unicardium
ranvillianum d'Orbigny, 1850.
rara (mode). - See also under rarum Evans & Shumard.
rara Kvaliashvili, 1978 . - Digressodacna rara
Kvaliashvili, 1978 . Lymnocardiidae. Pliocene.
RECORDS. --1978 ZR: Digressodacna rara Kvaliashvili,
1978 sp. nov. Georgia SSR. Ref. to Kvaliashvili p. 54.
rarepartita (variety). - See under paucicostatum
planicosta var. rarepartita Coen, 1933. - paucicostalum
var. eburnea Coen, 1937 . - paucicostatum var.
rarepartita Coen, 1933 .
Raricardium (genus). - Paramonova, 1986. Cardiidae.
Pliocene. RECORDS . --1987 ZR: Raricardium
Paramonova, 1986 gen. nov. of Cerastoderrnini. Type
species Cardium konjuschkevskii Alizade, 1932.
Pliocene. Ref. to Nevesskaya et al. p. 191.
raricostata Papaianopol, 1992. - Prosodacna raricostata
Papaianopol, 1992. Lymnocardiidae. Pliocene. TYPE
LOCALITY. Dacien inferieur (Getien) du Basin Dacique,
Romania. RECORDS. --1992 Papaianopol, p. 28, fig.:
Prosodacna (Prosodacna) raricostata sp. nov.
raricostata Popov, 1983 . - Didacna raricostata Popov,
1983 . Lymnocardiidae. Pleistocene. TYPE LOCALITY .
Russian SFSR. RECORDS. --1983 Popov, p. 200, fig.:
Didacna raricostata sp. nov.
*raricostatum - Homonymy to be noticed. Cardium
raricostatum from 2 sources: Deshayes, 1853 . - AliZade & Ribina (?year).
raricostatum Ali-Zade & Ribina. - New combination:
Replidacna raricostatum (Ali-Zade & Ribina).
Lymnocardiidae. Fossil. RECORDS. --1977 ZR: Cardium
(Replidacna) raricostalum Ali-Zade & Ribina. Ref to
Paramonova p.70 [Transl. p. 320].
raricostatum Deshayes, 1853. - Combination to be
revised: "Cardium" raricostatum Deshayes, 1853.
?Cardiidae. Fossil. RECORDS. --1980 Ruhoff: Cardium
raricostatum Deshayes, 1853; vol. 2: 45.
raricostatum Kafanov, 1994. - Keenocardium
raricostatum Kafanov, 1994. Cardiidae. Fossil.
RECORDS . --1994 Kafanov in Arkhipova et ai.:
Keenocardium raricostatum Kafanov.
rarissima Stevanovic, 1951. - Didacna rarissima
Stevanovic, 1951 . Lymnocardiidae. Ponti an. RECORDS.
--1953 ZR: Didacna rarissima Stevanovic, 1951 sp.
nov. Pontian Serbia [Ref. Stevanovic 1951].
raristriatum Bellardi, 1852. - Combination to be revised :
"Cardium " raristriatum Bellardi, 1852. ?Cardiidae.
Fossil. RECORDS. --1980 Ruhoff: Cardium raristriatum
Bellardi, 1852; vol. 4: 242, pI. H (sic = 19), fig. 10.
raritanensis Richards, 1943. - Combination to be
revised : "Cardium" raritanensis Richards, 1943 .
Cardiidae. Fossil. --1943 ZR: Cardium (Granocardium)
raritanensis Richards, 1943 sp. nov. New Jersey.
Raritan.
rarum Evans & Shumard, 1857. - Changed combination:
Protocardia rarum Evans & Shumard. Cardiidae.
Cretaceous. RECORDS. --1860 Meek & Hayden:
Cardium rarum Evans & Shumard. --1876 Meek:
Cardium (Protocardia?) rarum Evans & Shumard.
"1876 Meek: Protocardia (Leptocardia) rara Evans
& Shumard. --1876 Meek: Cardium (Leptocardia) rara
Evans & Shumard. --1970 Speden: Cardium rarum
Evans & Shumard, 1857, p. 39 . --1980 Ruhoff:
Cardium rarum Evans & Shumard, 1857; vol. 1: 39.
rarum Paramonova, 1977. - Cerastoderma rarum
Paramonova , 1977. Cardiidae. Upper Miocene .
RECORDS. --1977 ZR: Cerastoderma (Pianacardium)
732
Tropical Marine Mollusc Programme (TMMP)
rasmys!ovae - reetidens minor
rarum Paramo nova, 1977 sp. nov. Ukraine. Ref to
Il'ina, Nevesskaya & Paramonova p. 208, fig'd.
rasmyslovae. - Nemocardium rasmyslovae. Cardiidae.
Fossil. RECORDS. "1977 Popov: Nemocardium
(Nemocardiwn) rasmyslovae.
rastrum Reeve, 1845. [Figs. Part 1 p. 139. Part 3 p. 923).
- Synonym of Traehycardium senticosum (Sowerby,
1833). » Regarding Cardium rastrum: Cardiidae.
Recent. TYPE REPOSITORY. 2 syntypes. 1978131 . BMNH
London. TYPE LOCALITY. (Reeve 1845 : unknown), Santa
Elena, Ecuador, in 6-12 fathoms, sandy mud (Hertlein
& Strong 1947). RECORDS. "1845 Reeve, sp. 82:
Cardium rastrum Reeve, 1845 = Cardium senticosum
Sowerby, 1841 (in the errata). "1852 Adams: Cardium
rastrum Reeve 1845 = Cardium senticosum Sowb.
1833. "1858 Adams & Adams , p . 455: Cardium
(lsocardia) rastrum Reeve. "1869 Romer: Cardium
ras/rum Reeve Fig. 82 = Cardium senticosum Sowerby.
"1886 Dall : Cardium ras/rum Reeve = C. senticosllm
Sowerby, "1890 Paetel: Cardium rastrum Rv. =
senticosum Sow. "1891 Stearns: C. rastrum Rve. =
Cardium senticosum Sby. "190 I Dall: Cardium
rastrum Reeve, 1845= Cardium senticosum Sowerby,
1833. "1910 Dall: Cardium rastrum Reeve = Cardium
sen/icosum Sowerby. "1922-32 Sherborn, p. 5436:
Cardium ras/rum Reeve, 1845. "1947 Hertlein &
Strong: Cardium rastrum Reeve = Cardium
(Dal/ocardia) senticosum Sowerby, 1833 . "1971 Keen
& McLean : Cardium rastrum Reeve , 1845 =
Trachycardium (Dal/ocardia) sen/icosum (Sowerby,
1833). - CHARACTERS AND OCCURRENCE DESCRIPTIONS, "1845 Reeve: shell nearly orbicular,
rather thin, radiately ribbed, 39 ribs, the first 13 anterior
ribs prickly on the anterior side, then 2, sometimes 3,
prickly on both sides, the rest prickly on the posterior
side; brownish white, banded and spotted with reddishbrown. REMARKS. "1845 Reeve: the sculpture of this
shell is arranged very much after the style of the
Cardium muricatum. HABITAT. "ZMUC: clay-sand at
11-29 m depth. DISTRIBUTION. "1845 Reeve: St. Elena,
West Columbia, "1971 Keen & McLean : Gulf of
California to Peru.
raulineanum Orbigny, 1844. - New combination:
Criocardium raulineanum (Orbigny, 1844). Cardiidae.
Fossil. RE CORDS. "1998 Schneider: Cardium
raulineanum Orbigny, 1844. Cardium (Crio cardium)
raulineanum Orbigny, 1844.
*raulini. - Homonymy to be noticed. Cardium raulini
from 4 sources: Coquand, 1859. - Gumbel (?year). Orbigny, (?year). - Heb. & Bosquet (?year).
raulini Coquand, 1859. - Combination to be revised:
"Cardium" raulini Coquand, 1859. Cardiidae. Fossil.
RECORDS. "1980 Ruhoff: Cardium raulini Coquand,
1859; vol. 16: 1003 .
raulini Gumbel. - New name. Combination to be revised:
"Cardium" teisenb ergerense Schlosser, 1926. »
Regarding Cardium raulini: Cardiidae. Fossil. RECORDS.
"1926 ZR: Cardium raulini Gumbel replaced by
Cardium teisenbergerense Schlosser, 1926 nom. nov.
raulini Hebert & Bosquet, 1849. - Synonym of
"Cardium" kochi Semp. Combination to be revised.
» Regarding Cardium raulini: Cardiidae . Fossil.
RECORDS . "1886 Vincent : Cardium raulini Heb . et
Bosquet in Dewalque = Cardium kochi Semp. "1922-
32 Sherborn, p. 5437: Cardium raulini Hebert, 1849.
raulini Orbigny. - Combination to be revised:
"Cardium" rartlini d'Orbigny. Cardiidae. Cretaceous.
RECORDS. "1999 Jacques Vidal (pers. com.), MNHN,
Paris : Fossils from the Paleontological Collection.
Cardium raulini d'Orbigny. Albien . France (Yonne),
S07060.
raulinianum Orbigny, 1844 . - Pleuriocardia
raulinianum (d'Orbigny, 1844). Cardiidae. Cretaceous.
RECORDS. "1922-32 Sherborn , p. 5438: Cardium
raulinianum d'Orbigny, 1844. "1978 Scott: Cardium
raulinianum d'Orbigny. "1978 Scott: Pleuriocardia
(Plellriocardia) raulinianwn (d'Orbigny) .
razelmiana (variety). - See under eolorata razelmiana
Borcea, 1926.
reatrieosum Hall. - Conocardillm rea/rieosum Hall.
Conocardiidae. Fossil. "1943 ZR : Conocardium
reatrieosum Hall (revised descriptions) Ref. to Branson
( 1942),
reetangulare Roemer, 1860. - Combination to be
revised: "Cardium" reetangulare Fr. Roemer, 1860.
Cardiidae. Fossil. RECORDS. "1980 Ruhoff: Cardium
reetangulare Fr. Roemer, 1860; vol. 9: 12, pI. 4, fig. 5.
reetangulare Schlosser, 1926. - Nemocardium
reclangulare Schlosser, 1926. Cardiidae. Eocene.
RECORDS. "1926 ZR: Nemoeardium ree/angulare
Schlosser, 1926 sp. n. Eocene Bavaria.
reetidens Coen, 1917. - Cerastoderma reetidens (Coen,
1917). OTHER JUDGEMENTS: ?Cerasloderma glaueum
(Bruguiere, 1789). » Regarding Cerastoderma
rectidens: Cardiidae. Recent. RECORDS ... 1917 ZR:
Cardium (Cerasloderma) reetidens Coen, 1917 sp. n.
Mediterra nean.
..1933
Coen:
Eucardium
(Cerastoderma) reelidens. "1937 Coen: Cerastoderma
rectidens Coen.
reetidens adusta Monts. Mss. (subspecies). Infrasubspecific. : Cerastoderma reetidens var. adusta.
RECORDS. "1917 ZR: Cardium (Cerastoderma)
reelidens adusta Monts. Mss. [Ref. Coen 1917].
reetidens alba Monts. Mss. (subspecies). Infrasubspecific . Cerastoderma reetidens var. alba
RECORDS . "1917 ZR: Cardium (Cerastoderma)
reetidens alba Monts. Mss. [Ref. Coen 1917).
reetidens clodiense Renier (variety). - Infrasubspecific.
No evidence of subspecific status. Cerastoderma
rectidens var. clodiense Renier. (Coen, 1917). OTHER
JUDGEMENTS: Cerastoderma rectidens var. clodiense
Renier. - ?Cerastoderma glaucum (Bruguiere, 1789).
» Regarding Cerastoderma reelidens var. clodiense:
Cardiidae. Recent. RECORDS. "1933 Coen: Eucardium
(Cerastoderma) reetidens var. clodiensis Renier. "1937
Coen: Cerastoderma rectidens clodiense Renier.
rectidens drepanensis Coen, 1917 (subspecies). Infrasubspecific. No evidence of subspecific status.
RECORDS. "1917 ZR: Cardium (Ceras toderma)
rectidens drepanensis Coen, 1917 n. var.
reetidens Jragi/is Coen, 1917 (subspecies). Infrasubspecific. No evidence of subspecific status.
RECORDS. "1917 ZR: Cardium (Ceras/oderma)
reelidens fragilis Monts. Mss. [Ref. Coen 1917].
reetidens minor Coen, 1917 (variety).- No evidence of
subspecific status. » Regarding Cardium reetidens
minor: See also Cardium glaueum rectidens var. minor
Coen, 1915. RECORDS. "1917 ZR: Cardium
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rectidens obliquata - regulare
(Cerastoderma) rectidens minor Coen, 1917 n. var.
rectidells obliquata Coen, 1917 (subspecies). Infrasubspecific. No evidence of subspecific status.
Cerastoderma rectidens var. obliquata RECORDS. ··1917
ZR: Cardium (Cerastoderma) rectidens obliquata
Monts. Mss. [Ref. Coen 1917] .
rectidells pulchefla Coen, 1917 (subspecies). Infrasubspecific. No evidence of subspecific status.
Cerastoderma rectidens var. pulchella. RECORDS.
··1917 ZR: Cardium (Cerastoderma) rectidens
pulchella Monts. Mss. [Ref. Coen 1917].
rectidells supilla Coen, 1917 (subspecies). Infrasubspecific. No evidence of subspecific status.
Cerastoderma rectidens var. supina RECORDS. ··1917
ZR: Cardium (Cerastoderma) rectidens sup ina Monts.
Mss. [Ref. Coen 1917].
rectidells varnellsis Coen, 1917 (variety) . Infrasubspecific. No evidence of subspecific status.
Cerastoderma rectidens var. varnensis REcORDs. ··1917
ZR : Cardium (Cerastoderma) rectidens varnensis
Coen, 1917 n. var.
rectispilla Koenen, 1893. - Changed combination:
Agnocardia rectispina (Koenen, 1893). Cardiidae.
Fossil. RECORDS. ··1893 Koenen: Cardium rectispina
(fide Schneider 2002). ··1984 ZR: Agnocardia
rectispina (Koenen, 1893). Taxonomic position. Ref.
to Vokes p. 43. ··1998 Schneider: Agnocardia rectispina
(Koenen, 1893) Cardiidae. Early Oligocene, Gennany.
··2002 Schneider: Agnocardia rectispina (Koenen,
1893). Cardiidae. Rupelian.
rectum Taltasse, 1950. - Synonym of Cardium hians
var. recta Dollfus. » Regarding Bucardium rectum:
Cardiidae. Miocene. TYPE LOCALITY. Portugal. TYPE
REPOSITORY. Unknown. RECORDS. ··1952 Lecointre:
Bucardium rectum Taltasse, 1950 = Cardium hians var.
recta Dollfus.
reductum Alizade. - Changed combination:
Acobaecardium reductum (Alizade) . Lymnocardiidae.
Neogene. R ECOR DS. ··1986 Nevesskaya et al.:
Acobaecardium reductum (Alizade) comb. nov. ··1987
ZR: Acobaecardium reductum (Alizade) comb. nov.
Ref. to Nevesskaya et af. p. 192.
reedi Loel & Corey, 1932. - Combination to be revised:
"Cardium " reedi Loel & Corey, 1932. Cardiidae.
Miocene. R ECOR DS. ··1933 ZR: Cardium
(Acanthocardia) reedi Loel & Corey, 1932 Miocene
California . ··1944 Keen & Bentson: Cardium
(Acanthocardia) reedi Loel & Corey, 1932 (n. sp.).
*reeveallum . - Homonymy to be noticed. Cardium
reeveanum from 2 sources: Dunker, 1852. - Deshayes,
1853 .
reeveanum Deshayes, 1853. - Combination to be
revised : "Cardium" reeveanum Deshayes, 1853.
Cardiidae. ?Recent. RECORDS. ··1980. Ruhoff: Cardium
reeveanum Deshayes, 1853; vol. 2: 46.
reeveanum Dunker, 1852. - Synonym of Vasticardium
vertebratum (Jonas, 1844). OTHER JUDGEMENTS:
Cardium fla vum Linnaeus of authors. » Regarding
Cardium reeveanum: Cardiidae. Recent. RECORDS.
··1852 Dunker: Cardium reeveanum. ··1858 Dunker:
Cardium reevea num. ··1869 Dunker: Cardium
reeveanum. ··1872 Tryon: Cardium reeveanum Dkr.
··1884 Grasset: Cardium reeveanum Desh. ··1884
Smith : Cardium ree veanum Dkr. ··1890 Paetel :
Cardium reeveanum Dkr. ··1906 Hedley: Cardium
reevianum sic Dunker. ··1910 Hedley: Cardium
reeveanum Dkr. ··1923 Shirley: Cardium reeveanum
Dkr. ··1977 Fischer-Piette, p. 30: Cardium reeveanum
Dunker = Laevicardium flavum Linne auctorum. ··1977
Wilson & Stevenson: Cardium reeveanum Dunker,
1852. "1977 Wilson & Stevenson: Acrosterigma
reeveanum (Dunker, 1852). ··1978 ZR: Acrosterigma
reeveanum (Dunker 1852). Senior synonym of
Regozara olivifer Iredale, 1936. Ref. to Wilson et al.
p. 82. ··1980 Ruhoff: Cardium reeveanum Dunker,
1852; [1852-1853] vol. 9: 54. ··1980 Ruhoff: Cardium
reeveanum Dunker, 1858; [1858-1870] vol. 2: 22, pI.
6, figs. 6-8 . ··1990 Abbott & Dance: Trachycardium
reeveanum. ··1922 Shirley: Cardium reeveanum
Dunker. ··1992 Lamprel\ & Whitehead : Acrosterigma
reeveanum . ··1997 Vidal : Cardium reeveanum Dunker,
1852 = Vasticardium vertebra tum (Jonas, 1844). ··1999
Vidal : Cardium reeveanum Dunker, 1852 = Vasticardium vertebra tum (Jonas, 1844). - CHARACTERS
AND OCCURRENCE - DISTRIBUTION. ··1884 Grasset:
Australia. ··1994 Hylleberg: Red Sea, Australia.
reevei Hidalgo, 1903. - New name Tridacna reevei
Hidalgo, 1903. Tridacnidae. Recent. RECORDS. ··1903
ZR: Tridacna reevei Hidalgo, 1903 n.n. for Tridacna
elongata Rve. nec Lamarck.
reevianum (error). - Printer's mistake for reeveanum in
Hedley 1906.
reflexum - Conocardium rejlexum. Conocardiidae.
Fossil. RECORDS. "1889 ZR : Conocardium rejlexum
with fig. (ref. Barrois).
Regozara Iredale, 1936 (genus). - Cardiidae. Neogene
and Recent. Type of genus Regozara olivifer Iredale,
1936. Original designation. RECORDS. ··1980 Vokes :
Regozara Iredale, 1936 in subfamily Trachycardiinae.
- CHARACTERS AND OCCURRENCE - DESCRIPTIONS. ··1969 Moore et af. : hinge less angulate than in
A. (Acrosterigma), sculpture of small scales or crossthreads on posterior sides of ribs. East Indies-Australia.
Regozara Iredale, 1936 (subgenus). - Cardiidae.
Neogene and Recent. RECORDS. ··1969 Moore et al.
classified Regozara as a subgenus of the genus
Acrosterigma Dall, 1900. "1978 Maxwell: Subgenus
Regozara Iredale, 1936. ··1999 Higo et af.: Regozara
(subgenus) Iredale, 1936. Type species: Regozara
olivifer Iredale, 1936 (o.d.).
*regulare. - Homonymy to be noticed. Cardium
regulare from 2 sources : Bruguiere, 1789. - Coquand,
1862.
regulare Bruguiere, 1789. - Synonym of Vasticardium
p ectiniforme (Born, 1780) . OTHER JUD GEME NTS:
Trachycardium jlavum(Linnaeus, 1758). - Variety of
Cardium rugosum. - Cardium leucostomum Born,
1780. » Regarding Cardium regulare: Cardiidae.
Recent. DISTRIBUTION. C. regulare St. Domingo (Wood
& Hanley 1856); Indo-Pacific (Hylleberg 1994). C.
pectiniforme: Red Sea, Indo-West Pacific . RECORDS.
··1789 Bruguiere, No. 24. Cardium regulare; Nob.
"1815 Wood: Cardium regulare Brug. ··1817 Dillwyn:
Cardium regulare Bruguiere. ··1819 Lamarck :
Cardium regulare, Brug. Dict. no. 24 . (American
variety of Cardium rugosum). ··1836 Bosc: Cardium
regttlare. ··1856 Wood & Hanley, No. 29 : Cardium
regulare Brug. E. M. Verso 227 . ··1902 Sherborn, p.
734
Tropical Marine Mol/usc Programme (TMMP)
regulare - replicalllm
824: Cardium regulare Bruguiere, 1789. --1932
Prashad: Cardium reguLare Bruguiere, 1789 =
Trachycardiumjlavum (Linne, 1758). --1977 FischerPiette : Cardium regulare Bruguiere = Cardium
Leucostomum Born. --1999 Higo et at.: Cardium
regulare Bruguiere, 1789 = Acrosterigma (Regozara)
jlava (Linnaeus, 1758). --1997 Vidal: Cardium regulare
Bruguiere, 1789 = Vasticardium pecliniforme (Born,
1780). --1999 Vidal: Cardium regulare Bruguiere, 1789
= Vasticardium pectiniforme (Born, 1780).
regulare Coquand, 1862 . - "Cardium" regulare
Coquand, 1862. Cardiidae. Fossil. RECORDS. -- I 980
Ruhoff: Cardium reguLare Coquand, 1862; vol. 2: 205,
pI. 10, figs. 13-14.
regularis (subspecies). - See under edule reguLaris
Pallary, 1900.
reissi Bronn in Reiss & Bronn, 1862. - Combination to
be revised : "Cardium" reissi Bronn in Reiss & Bronn,
1862. Cardiidae. Fossil. RECORDS. --1980 Ruhoff:
Cardium reissi Bronn in Reiss & Bronn, 1862; p. 40,
pI. I, fig. 17.
rekensis Ebersin, 1967. - Macradacna rekensis Ebersin,
1967. Lymnocardiidae. Pliocene. RECORDS . "1967
Ebersin: Macradacna rekensis sp. n. "1967 ZR:
Macradacna rekensis Eberzin, 1967 sp. nov. Pliocene,
USSR.
relicta (subspecies). - See under plicata relicta.
relicta Milaskewicz , 1916. - Adacna relicta
Milaskewicz, 1916. OTHER JUDGEMENTS: Hypanis
plicatus relictus (Milaskewicz).» Regarding Adacna
relicta: Lymnocardiidae. Recent. RECORDS. --1916
Milaschewitch, p. 274 (pI. 8, figs. 10-12): Adacna
relicta sp. n. --1923 ZR: Adacna relicta Milachewitch,
1916. "1927 ZR: Adacna relicta [Ref. Borcea 1927b).
--200 I Vidal, p. 407: Adacna relicta. - CHARACTERS
AND OCCURRENCE - DESCRIPTIONS. --2001 Vidal:
No tentacles. No eyes. DISTRIBU TION. Black Sea
(Nordsieck 1969).
relicta dolosmiana Borcea, 1926 (variety). - Upgraded.
Hypanis dolosmiana Borcea, 1926. Lymnocardiidae.
Recent. REMARKS. Upgraded to species. RECORDS .
--1927 ZR: Adacna relicta var. dolosmiana Rumania,
[Ref. Borcea 1927b). "1928 ZR: Adacna relicla var.
dolosmiana Borcea, 1928 nov. Roumania [Ref. Borcea
1928b J. --1962 Grossu: Adacna relicta var. dolosmiana
Borcea, 1926 = Hypanis dolosmiana BOIcea, 1926 in
Scarlato & Starobogatov. --1962 Grossu : Adacna
plicata relicta forma dolosmiana Borc. = Hypanis
dolosmiana Borcea, 1926 in Scarlato & Starobogatov.
remondianum Gabb, 1864. - Combination to be revised:
"Cardium " remondianum Gabb, 1864. Cardiidae.
Fossil. RECORDS. "1871 Stoliczka, 143 : C.
remondianum . --1980 Ruhoff: Cardium remondianum
Gabb, 1864; vol. I; 172, pI. 23, fig. 153.
renevieri - Combination to be revised: "Cardium"
renevieri. Cardiidae. Eocene. RECORDS . --1886
Cossmann: Cardium renevieri.
reniforme Koenen, 1893 . - Changed combination:
Goniocardium reniforme (Koenen, 1893). Cardiacea.
Oligocene . RECO RDS. "1998b Schneider, p. 353 :
Cardium (Fragum) reniforme Koenen, 1893. It is a
Goniocardium.
renngarteni Wassoievic & Eberzin, 1931. - Prosodacna
renngarteni Wassoievic & Eberzin , 1931.
Lymnocardiidae. Fossil. "1931 ZR: Prosodacna
renngarleni Wassoievic & Eberzin, 1931 sp. n.
renticosum (error). - Printer's mistake for ventricosum
in Zoological Record 1945.
repens Taktakishvili, 1966. - Didacna repens
Taktakishvili, 1966. Lymnocardiidae. Pliocene.
RECORDS. "1966 ZR: Didacna repens Taktakishvili,
1966 sp. nov. Pliocene, western Gruzia, SSR.
repentinus Grishkevitch, 1962. - Combination to be
revised: "Cardium " repentinus Grishkevitch, 1962.
Cardiidae. Miocene. RECORDS. --1959 ZR: Cardium
repentinus Grishkevitch & Nevesskaya, 1959 sp. nov.
as nom . nud. in list Trans-Carpathians, Sarmatian,
Miocene. --1962 ZR: Cardium repentinus Grishkevitch,
1962 sp. nov. Sarrnatian, Trans-Carpathians, USSR
replicatum Spengler 1799 . [Figs. Part 3 p. 914).
Synonym of Corculum monstrosum (Gmelin 1791).
OTHER JUOGEMENTS: Corculum cardissa (Linnaeus,
1758). » Regarding Cardium replicatum : Cardiidae.
Recent. TYP E LOCALITY. "1782 Chemnitz Nicobar
Islands. "1799 Spengler: unknown. TYPE REPOSITORY.
ZMUC, Spengler's coIl. RECORDS. --1799 Spengler, p.
52-53 , No. 33: Cardium replicatum Chemnitz Tom .
VI. Tab. 14. fig. 149.150. "1799 Spengler: replicatum
= monstrosum (reference to Chernnitz Tom. 6. Tab. 14.
fig . 149. 150). "1870 March: Cardium replica tum
Spengler = C. monstrosum Chemn. --1902 Sherborn,
p. 826: Cardium replicatum Spengler, 1799. --1977
Fischer- Piette: Cardium replicatum Chemnitz =
Cardillm cardissa Linne. --1999 Knudsen & Hylleberg:
Spengler, 1799 No. 33: Cardium replica tum in Spengler
= Cardium monstrosum Chemnitz Tom. VI. Tab. Tab.
14 Fig. 149-150. - CHARACTERS AND OCCURRENCE - OF COLLECTIONS. --ZMUC I specimen,
Spengler's collection. CHARACTERS of diagnostic value:
the anterior side very inflated, round, the shell almost
half-spherical, posterior side impressed into the curved
anterior, margin bent over, creating a sharp keel, ribs
rounded , broad interstices , irregularly nodulose.
Anterior surface, alternating narrow and broad stripes.
The bent in margin spiny, shell not gaping, colour white
throughout. DESCRIPTIONS. --1799 Spengler: testa
cordata aequivalvi, valvulis dentato-carinatis, poslice
e.xcavato-depressa, carina rejlexa. --1799 Spengler
translated by Knudsen & Hylleberg 1999: The back
side of this Venus heart is so strongly elevated and
roundly vaulted that the bivalve thereby gets the
appearance of half a sphere. The front side lies partly
very deep internally in the globular posterior part, on
which its margin is bent strongly forwards ; this front
side is rather flattened, only towards the middle elevated
with a sharp crest. The ribs on this part are rounded
and separated from each other by rounded furrows and
irregularly set with small tubercles. The surface of the
backside is covered by alternately narrow and broad
stripes, as is common in most Venus hearts. The anus
is a comparatively large and smooth heart-shaped
depression. The hymen is very small, the introvert
margin is provided with pointed spines. Both valves
interlock closely at the margins by means of small teeth.
Inside the margin of the anterior part is snow-white
and strongly enameled; on the inner and outer sides
the bivalve is white. Its height and breadth are I inch I
line. Another smaller specimen exists, which on the
Phuket Marine Biological Center Special Publication vol. 30 (2004)
735
Replidacna - retusum
posterior vaulted side is densely set with light-brown
spots on a white background, up to an elevated, heartshaped casing of the groove or Anus, which is white.
The enameled anterior side, after the external ribs to
below the umbo, is inside incised with narrow furrows,
which are serrate on the margin and interlock both
valves. DIMENSIONS. "1799 Spengler: length and width
28 nun (1 Tom. I Lin.). REMARKS. "1799: Spengler
did not give reasons for changing the name Cardium
monslrosum given by Chemnitz, 1782. DISTRIBUTION.
"1782 Chemnitz: Nicobar Islands.
RepJidacna Jekelius, 1944 (invalid genus).
Lymnocardiidae . Miocene. RECORDS. "1960 ZR:
Rep/idacna Jekelius, note on this genus created for
small Limnocardium with exaggerated hinge line
development. [Ref. Gillet 1960b]. "1960 ZR:
Replidacna used as a genus for young Iymnocardiids
[Ref. Gillet I960a). "1980 Vokes: Replidacna Jekelius,
1944. Subfamily uncertain.
RepJidacna Jekelius, 1944 (invalid subgenus).
Lymnocardiidae. Miocene. RECORDS. "1946 ZR:
Replidacna Jekelius, 1944 'subgen. nov.' [given as a
new genus on p. 153 and in Index p. 167 and used as
such in headings of species] Cardiidae, type Replidacna
carasi Jekelius, 1944. Miocene Soceni Bana!. "1977
ZR: Cardium (Replidacna) lekelius. 1944. Invalidation.
Ref to Paramonova p. 69 [Transl. p. 318).
requienianum Matheron, 1843. - Combination to be
revised: "Cardium" requienianum Matheron, 1843.
Cardiidae. Fossil. RECORDS. "1922-32 Sherborn, p.
5470: Cardium requienianum Matheron, 1843.
restinense Olsson, 1928. - Combination to be revised:
"Cardium" restinense Olsson, 1928. Cardiidae. Fossil.
RECORDS . "1928 ZR: Cardium (Ringicardium)
reslinense Olsson, 1928 sp. n.
reticulatum Fischer, 1807. - Combination to be revised:
"Cardium " reticulalum Fischer, 1807. Cardiidae.
Fossil. RECORDS. "1922-32 Sherborn, p. 5476: Cardium
reticulatum Fischer, 1807.
reticulatum Meuschen, 1778. Non binominal. - Not
available. RECORDS. "1902 Sherborn: Venus reticula tum
Linnaeus . "1902 Sherborn, p. 827: Cardium (L.)
reticulalum Meuschen, 1778. Venus reticulatum.
reticulatum Muromtzeva in Khalfina , 1962. Conocardium reticulatum Muromtzeva in Khalfina,
1962. Conocardiidae. Carboniferous. RECORDS. "1962
ZR: Conocardium reticulatum Muromtzeva in
Khalfina, 1962 sp. nov. Carboniferous, Sayano-Altai
Mts. USSR
retowskii Andrus. - Paradacna retowskii Andrus.
Lymnoc a rdiidae. Fossil. RE COR DS . "1953 ZR:
Paradacna relovskii. Ponti an [Ref. to Stevanovic 1951].
retowskii (cf.) Andrus. - RECORDS. "1953 Celidje:
Paradacna cf. retowskii Andrus.
retowskii controversa Papaianopol, 1978 (subspecies).
- Paradacna relowskii conlroversa Papaianopol, 1978.
Lymnocardiidae. Pliocene. RECORDS . "1978 ZR :
Paradacna relowskii conlroversa Papaianopol, 1978
subsp. nov. Romania. Ref. to Papaianopol, I. p. 289.
retowski ossoinae Stevanovic, 1951 (variety). Upgraded. Paradacna retowski ossoinae Stevanovic,
1951. Lymnocardiidae. Fossil. RECORDS. ··1953 ZR:
Paradacna retovskii var. ossoinae Stevanovic, 1951
nov. Pontian Serbia. "1988 Lyutzkanov & Ziatev:
Paradacna retowski ossoinae Sev.
retrostrialum Buch, 1832. - Combination to be revised:
"Cardium " relrOSlrialum d'Orbigny, 1850. Not in
Cardiidae. Fossil. RECORDS. "1857. Deshayes, p. 51:
Cardium retrostriatum Buch [Silurian-Devonian]
"1922-32 Sherborn, p. 5486: Venericardium
retrostrialum Buch, 1832. "1922-32 Sherborn, p. 5486:
Cardium (Buch) retrastriatum d'Orbigny, 1850. Venericardium.
retrostriatum Sandberger, 1845. - Lunulocardium
retrostriatum Sandberger, 1845. Not in Cardiidae.
Fossil. RE CORDS. "1922-32 Sherborn, p. 5486:
Lunulocardium retroslrialum Sandberger, 1845.
retusa (mode). - See under relusum Linne, 1758.
relusum Linnaeus, 1758. [Figs. Part I p. 66,71 , 86, 103,
168. Part 3 p. 910,911 , 932]. - Lunulicardia relusum
(Linne, 1758). OTHER JUDGEMENTS: Lunulicardia
auricula (Niebuhr in Forsskaal, 1775). - Cardium
subretusum (Sow., Reeve) is only a variety of retusum
(L.), but auricula (F orsk.) is a distinct species [Ref.
Romer in ZR for 1868]. » Regarding Cardium
retusum: Cardiidae. Recent. GENOTVPE. Gray (1853)
designated Cardium retusum Linne, 1758 type species
of the subgenus (Lunulicardia) Gray, recognised at the
generic level by authors. TVPE LOCALlTV. Unknown.
Spengler (1799) refers his specimens to Tranquebar,
the Coromandel coast; Reeve (1844) to China; Sowerby
(1841 No. 92) refers to Suez, Arabia. RECORDS. "1758
Regenfuss, Vol. 2, pI. 9, fig. 20: Cardium retusum (fide
Spengler 1799). "1767 Linne, p. 1121 , sp. 75: Cardium
retusum. Habitat in India. Sprengler sic. "1778 Born,
G3. Cardium retusum Linn. S.N. Sp. 75. "1780 Born:
Cardium retusum Linn. S. N. Sp. 75. "1780 Born. ID
by Gmelin 1791: Cardium retusum mus. Caes. Vind.
test. t. 3. f. I. 2. "1782 Chemnitz, Tab. 14 Fig. 139142: Cardium retusum Linnaei. "1782 Chemnitz. ID
by Gmelin 1791 : Cardium relusum t. 14. f. 139-142.
"1786 Schroter, No. III: Cardium retusum. Linn. XII.
pag. 1121 . Spec. 75. "1788 Schroter: Cardium retusum.
[Reference to Chemnitz 1782: fig . 147]. "1789
Bruguiere, No . 2: Cardium retusum Linn. "1791
Gmelin, p. 3245, No.4: Cardium relusum. "1798
Bruguiere ID by Lamarck, pI. 294. fig. 3 a, b, c, d.
Cardium retusum. ··1799 Spengler: Cardium retusum
Linnei Sp. 75. p. 1121. "1799 Spengler: Cardium
retusum in Regenfus sic Tom. 2. Tab. 9. fig. 20. "1815
Wood p. 243, No. 52: Cardium relusum Linn. Gme!. p.
3245. "1817 Dillwyn: Cardium relusum Linnaeus.
"1836 Bose: Cardium retusum . "1840 Sowerby,
catalogue p. 7, No. 92: Cardium relusum Linn. "1844
Potiez & Michaud: Cardium retusum Linn. Gme!.
.. 1844 Reeve, sp. 103: Cardium retusum. "1850
Cuvier, pI. 99: Cardium relusum Lin. "1853 Gray:
Cardium retusum Linne, 1758. "1853 Gray:
Lunulicardia relusum. "1858 Adams & Adams, p. 459:
Hemicardia (Lunulicardia) reillsa, Linn. "1869 Romer:
Cardium (Lunulicardia) relusum . ··1879 ZR: Cardium
relusum (L.), nec Lunulicardium (Munster, 1840): Type
of Opisocardium Bayle , 1879 . "1884 Grasset:
Hemicardium retusum . L. "1889 Martens: Cardium
relusum (L.?) = Cardium (Hemicardium) subrelusum
Sow. "1890 Paetel: Hemicardium retusum L. "1899
Sturany: Hemicardia relusum. "1901 Sturany:
Hemicardium relusum (L.) (subretusum Sow.). "1902
736
Tropical Marine Mollusc Programme (TMMP)
retusum - retusum var.
DalJ : Cardium retusum Linnaeus, type of genus
Lunulicardia Gray. ,·1902 Sherborn, p. 829: Cardium
retusum Linnaeus, 1758. "1909 Lynge : Hemicardium
(Lunulicardia) retusum. "1928 Kuroda : Lunulieardia
re tusa (Linne, 1767). "1939 Adam & Leloup:
Coreulum (Lunulieardia) rehlSum (Linne 1767). "1945
ZR: Cardium retusum L. [Ref. Lamy 1945] ... 1952
Dawydoff: Hemicardium retusum Linne. "1952
Kuroda & Habe: Coreulum retusum (Linne). "1958
Abbott: Hemicardia (Lunulicardia) retl/sum Linne.
"1969 Moore et al.: Fragum (Lunulicardia) retusum.
"1960 Kuroda: Hemicardia (Lunulieardia) retusa
(Linne). "1970 ZR: Corculum (Lunulicardia) relusum
new record for Mozambique [Ref. Moura 1970). "1977
Popov: Corcllium (Lunulicardia) relusum (Linne,
) 758). "1977 Fischer-Piette: Corculum (Lunulicardia)
retusum Linne. "1977 Popov : Cardium retusum Linne,
1758 . Type of (Lunulicardia) Gray, 1853 . .. 1977
Wilson & Stevenson: Fragum (Lunulicardia) relusum
(Linnaeus, 1758). "1978 ZR : Fragum (Lunulicardia)
retusa Bay of Bengal. Burma, Maungmagan. First
record. Ref. to Ray. "1988 Drivas & Jay: Lunulicardia
retusua (Linne, 1758). "1990 Abbott & Dance:
Lunulicardia retusum. "1991 Matsukuma, Okutani &
Habe: Lunulicardia retusa (Linnaeus, 1758) . .. 1992
Oliver: Cardium ret£lSum Linne, ) 758 = Lunulicardia
auricula Niebuhr in Forsskaal, 1775 . "1992 Oliver:
Cardium retusum Linne, 1758 in Sturany (1899) =
Lunulicardia auricula Niebuhr in Forsskaa l, 1775.
"1998 Schneider (1998b): Lunulicardia relusa (Linne,
1767) . Harbours photo symbiotic dinofl age llates.
"1999 Higo et al.: Lunulicardia retusa (Linnaeus,
1767). "1999 Aungtonya et al. , PMBC-SP 19(2): 377:
Lunulicardia retusa (Linnaeus, 1758) . .. 200 I
Schneider & Carter: Lunulicardia retusa (Linne, 1758).
.. 200 I Vidal, p. 408: Lunulieardia relusum ... 2002
Hylleberg & Kilburn, p. 63: Lunulicardia retusa (Linne,
1758). "2003 Hylleberg & Kilburn, p. 183: Plate 2,
fig. 5. Lunulicardia relusa (Linnaeus , 1758) . CHARACTERS AND OCCURRENCE - OF COLLECTIONS. "ZMUC: Tranquebar, Ceylon, Gulf ofThailand.
··ZMUC: wet collection. CHARACTERS of diagnostic
value: Shell gibbously cordate, somewhat oblique,
posteriorly concavely depressed; radiately ribbed, ribs
convex, smooth, interstices strongly latticed, lunule
deeply excavated, with a conspicuous callosity, white
within and without. DESCRIPTIONS. "1799 Spengler:
testa cordala, valvulis stria tis crenulalis subcarinatis,
ana lunalo cordiformi inlruso. "1799 Spengler
translated by Knudsen & Hylleberg 1999: I have
deliberately let this cockle follow after the preceding
one, because it is completely similar to this except on
the posterior side. On this posterior side, right below
the umbo or the anus, is placed a heart-shaped, deep,
and, towards the innermost side, strongly widened
cavity. In this cavity, a free, elevated mussel- shaped
ornamenta tion is suspended from the umbo tip,
resembling an adornment for decoration of the upper
part of a niche. A very large specimen, which Prof.
Forskiil sent home from Arabia, is kept in the
"Grevelige Moltkiske Naturaliesamling". Today this
rare bivalve, though of a smaller size, is still found at
Tranquebar. "1815 Wood: shell heart-shaped, valves
striated, crenated, and somewhat keeled; a lunate heart-
shaped hollow behind the beak [Linne's description
translated from Latin by Wood). "1845 Reeve: shell
gibbously cordate, somewhat oblique, posteriorly
concavely depressed; radiately ribbed, ribs convex,
smooth, interstices strongly latticed, lunule deeply
excavated, with a conspicuous callosity intruding; white
within and without. .. 2001 Vidal: only the exhalant
aperture is tube-shaped . Inhalant siphon open towards
the foot aperture. No eyes. DIMENSIONS. "1988 Drivas
& Jay: ant.-post. 20 mm , umbo-ventT. 28 mm.
REMARKS. "1969 Moore et al.: Hemicardia is used by
Spengler 1799 as a subdivision, not as a genus. HABITAT
"1988 Drivas & Jay: Reunion, western Indian Ocean
to 12 m depth. "1999 Higo et al.: subtidal to 50 m,
sand. "ZMUC : sand, gravel at 6- 18 m depth .
DISTRIBUTION . "1799 Spengler: Tranquebar the
Coromandel coast. "1845 Reeve: China. "1884
Grasset: China. "ZMUC: Ceylon, Gulf of Thailand .
"1994 Hylleberg: Indo- West Pacific. "1999
Aungtonya el al.: Andaman Sea & Gulf of Thailand.
"1999 Higo el al.: Kii peninsula (Kuroda & Habe,
1981) and southwards, East China Sea, Philippines,
South China Sea, Hainan, Beibu gulf, southeast Asia,
northern Australia, Indo-Pacific. "2002 Hylleberg &
Kilburn: South India. "2003 Hylleberg & Kilburn:
Vietnam.
retusum of Reeve, 1845 . - Misidentification = Cardium
subrelusum Sow. RECORDS. "1880 Martens: Cardium
relusum Reeve fig. 103 = Cardium (Hemicardium)
subretusum Sow.
retusum of Swainson, 1840. - Hemicardium relusum
Swainson, 1840. Cardiidae. Fossil. RECORDS. "192232 Sherborn, p. 5487: Hemieardium (Wood) relusum
Swainson, 1840. -? Cardium L. 1767.
retusum of Wood, 1815 . - Lunulicardia auricula
(Niebuhr in Forsskaal, 1775). » Regarding Cardium
retusum: Cardiidae. Recent. Wood did not describe C
relusum. RECORDS. "1856 Wood & Hanley, No. 52:
Cardium relusum Wood (not Lin .) G.c. [General
Conchology 1815] pI. 58, f. 4, 5. = no. 31 = Cardium
auricula Gm. 3253, from Forsk. Arab. Red Sea.
retusum Chemnitz, 1782 . Non binominal. - Not
available. »Regarding Cardium retusum: Cardiidae.
Recent. Chemnitz did not describe .C relusum.
RECORDS. "1863 Deshayes: Cardium retusum Chernnitz
(non Linne) = Cardium auricula Forsk.
retusum var. Spengler, 1799 (variety). [Figs. Part 3 p.
911] . Infrasubspecifi c. RECO RDS. » Regarding
Cardium var. relus um : Cardiidae . Recent. T YP E
LOCALITY. Coromandel coast, India. TYPE REPOSITORY.
ZMUC, Spengler 's colI. RECORDS. "1799 Spengler, p.
55, No. 35: Cardium var. retusum . "1999 Knudsen &
Hylleberg: Cardium retusum var. Spengler, 1799. CHARACTERS AND OCCURRENCE - OF
COLLECTIONS. "ZMUC: C relusum var. Spengler's coil.,
I specimen. Coromandel, South India. CHARACTERS of
diagnostic value: Shell broad, thick, anteriorly rounded,
surface with few broad folds, granulated on the
posterior side, lunule strongly excavated, round like
part of a circl<:, very broad. DESCRIPTIONS. "1799
Spengler translated by Knudsen & Hylleberg 1999:
This Cardium is equally beautifully heart-shaped from
the front side as the preceding one [Cardium retusum] ,
but somewhat broader, consequently thicker, and
Phuket Marine Biological Center Special Publica/ion vol. 30 (2004)
737
retusum a - richardson;;
strongly rounded posteriorly. The heart-shaped casing
of the anterior part is neither so high nor so sharp as
that of the other one, but more flattened and more
rounded, its surface is very different from that of the
usual Cardium retusum ; since, in the middle, where
both valves are joined, it is not hollowed as the others,
but on the contrary inflated, and the heart-shaped folds
are broader, more deeply incised and smoothly rounded
above. Moreover, the total surface of the bivalve is set
with only a few, but very broad folds, of which those
on the back are granulated at their front side. The
unusually strongly hollowed depression at the anus is
not heart-shaped, but rounded as a half-circular arc,
and very broad. The whole bivalve is snow-white .
DISTRIBUTION. Coromandel coast, South India.
retusum a Chemnitz, 1782. Non binominal (variety). Not available. RECORDS. ··1839 Anton: Cardium
retusum Linne var. a. Chemnitz.
reussi Zittel, 1865. - Combination to be revised:
"Cardium" reussi Zittel, 1865. Cardiidae. Fossil.
RECORDS. ··1980 Ruhoff: Cardium reussi Zittel, 1865;
[1865-1866] vol. 24(2): 144, pI. 6, fig. 3.
rewaense Ladd, 1934. - Combination to be revised :
" Cardium " rewaense Ladd , 1934. Cardiidae .
Quaternary. RECORDS. ··1934 ZR: Cardium rewaense
Ladd, 1934 sp. n. Quaternary Fiji [Ref. Ladd 1934).
reynoldsi Rennie, 1929. - Combination to be revised :
"Cardium" reynoldsi (Rennie, 1929). Cardiidae.
Cretaceous. RECORDS. ··1929 ZR: Cardium
(Trachycardium) reynoldsi Rennie, 1929 sp. n.
Cretaceous Angola.
rhegminum Oliver & Chesney, 1997. - Vasticardium
rhegminum (Oliver & Chesney, 1997). Cardiidae.
Recent. REcORDs. ··1997 Oliver & Chesney, Journal
of Conchology 1997 Vol. 36: Trachycardium
rhegminum . ··1998 Vidal: Vaslicardium rhegminum
(Oliver & Chesney, 1997). ··1999 Vidal: Vasticardium
rhegminum (Oliver & Chesney, 1997).
Rhipidocardium (section). - Conocardiidae. REMARKS.
The designation "section" takes the value of subgenus.
Rhipidocardium have no connection with the Cardiacea
except analogy of form, which does not offer any point
of relationship (Dall 1895, p. 519). RECORDS ··1887
Fischer: Rhipidocardium Fischer, J 887. Section of
genus Conocardium.
rholesi Papaianopol, 1982. - New combination:
Lymnocardium rholesi Lymnocardiidae. Pliocene. TYPE
LOCALITY. Pontien et Dacien. Basin Dacique, Romania.
RECORDS . ··1982 Papaianopol , p. 206, fig.:
Limnocardium (Euxinicardium) rholesi sp. nov.
rhombeum d'Orbigny, 1850. - Combination to be
revised: "Cardium " rhombeum d'Orbigny, 1850.
?Cardiidae. Fossil. RECORDS. ··1922-32 Sherborn, p.
5506: Cardium (Hall) rhombeum d'Orbigny, 1850. Cypricardia. ··1922-32 Sherborn, p. 5506: Cypricardia
rhombeum Hall, 1843.
rhomboides Lamarck, 1819. [Figs. Part 3 p. 921].
Synonym of Cerastoderma edule (Linne, 1758). »
Regarding Cardium rhomboides: Cardiidae. Pliocene.
RECORDS. ··1819 Lamarck: Cardium rhomboides
Lamark sic. Note: considered an unimportant variety
of fossil Cardium edule abundant in Italy. ··1832
Deshayes : Cardium rhomboides Lamk . Fossilis =
Cardium edule L. "1867 Weinkauff: Cardium
rhomboides in Lamarck (J 818-1822) = C. edule Linne.
··1888 Kobelt: Cardium rhomboides Lam. = Cardium
var. edule. ··1922-32 Sherborn, p. 5509: Cardium
rhomboides Lamarck, 1819. ··1999 Jacques Vidal (pers.
com.), MNHN, Paris: Fossils from the Paleontological
Collection. Cardium rhomboides Lamarck. Algeria
(Oran), B35905, B41580. Tunesia (Gabes), B42004.
rhomboideum Khomenko, 1934. - Clinocardium
rhomboideum (Khomenko, 1934). Cardiidae. Miocene.
TYPE LOCALITY. Sakhalin . RECORDS. ··1954 Keen:
Clinocardium rhomboideum (Khomenko), 1934. [L.
Miocene]. ··1973 Keen: Clinocardium rhomboideum
(Khomenko, 1934) [Cardium] . Miocene, Sakhalin.
··1977 Popov: Clinocardium rhomboideum
(Khomenko,1934).
richardi Audouin in Savigny, 1827. [Figs. Part 3 p. 918].
Afrocardium richardi (Audouin in Savigny, 1827).
Cardiidae. Recent. TYPE LOCA LITY. Egypt. TYPE
REPOSITORY. Type in the Savigny Collection at the
MNHN, Paris (Bouchet, & Danrigal 1982). RECORDS.
··1809 Audouin, 51, pI. 9, fig . 14: Cardium richardi
new. ··1827 Audouin: Cardium richardi. ··1922-32
Sherborn, p. 5515: Cardium richardi Audouin in
Savigny, 1826. ··1926 Pallary: Cardium
(Parvicardium) richardi . ··1977 Fischer-Piette:
Cardium richardi Audouin. ··1992 Oliver: Afrocardium
richardi. ··1995 Oliver, fig. 1080: Afrocardium richardi
(Audouin, 1826). ··2000 Aartsen & Goud: Afrocardiwn
richardi (Audouin, 1826) . ··200 I Vidal, p. 408:
Afrocardium richardi. ··2002 Schneider: Afrocardium
richardi (Audouin, 1827). Holocene. ··2003 Hylleberg
& Kilburn, p. 176: Afrocardium richardi (Audouin,
1827). - CHARACTERS AND OCCURRENCE DESCRIPTIONS. ··1995 Oliver: beaks well in front of
midline. Obliquely oval, almost modioli form; posterior
not keeled but roundly angulate; posterior expanded,
margin long, almost straight. Sculpture of35-38 narrow
ribs, all with thin, imbricate, projecting scales, these
becoming longer on posterior ribs. White, yellow,
orange, red, some with reddish brown markings. ··200 I
Vidal: two siphons . No eyes. DIMENSIONS. ··1995
Oliver: to II mm. HABITAT. ··1995 Oliver: offshore.
DISTRIBUTION. ··1994 Hylleberg: Red Sea, Indo-West
Pacific. ··2003 Hylleberg & Kilburn: Vietnam.
richardi hiulca (mode) . - See under richardi hiulcum
Reeve 1845.
richardi hiulcum Reeve 1845 (subspecies). - Not a
subspecies. Changed combination: Afrocardium
richardi (Audouin, 1826). Cardiidae. Recent. RECORDS.
··1845 Reeve: Cardium hiulcum. ··1977 FischerPiette: Papyridea richardi hiulea Reeve . •• J992 Oliver:
Papyridea richardi hiulca Reeve = Cardium richardi
Audouin, 1826. - CHARACTERS AND OCCURRENCE - DI STR IBUTION. ··1994 Hylleberg: IndoPacific.
richardsoni Cox, 1946. - TUlcheria richardsoni Cox,
1946. Cardiacea. Middle Lias. RECORDS. ··1946 ZR:
Tutcheria richardsoni Cox, 1946 sp. nov. Middle Lias,
Chipping Norton, Oxon.
richardson;; Whiteaves, 1878. - Synonym of
Nemocardium centifilosum (Carpenter, 1864). »
Regarding Cardium richardsoni: Cardiidae. Recent.
TYPE LOCALITY. Strait of Georgia, between Rhode Island
Lighthouse and Victoria Harbor, in 30 to 50 fathoms .
738
Tropical Marine Mollusc Programme (TMMP)
richmondensis - ringens
TYPE REPOSITORY. Unknown. RECOR DS. "1878
Whiteaves, p. 468, foot note: Cardium richardsoni n.
sp. "1880 ZR: Cardium richardsoni sp. n. Whiteaves,
1880. British Columbia . "1921 ZR: Protocardia
richardsonii [Ref. Dall 1921] . .. 1937 Keen:
Nemocardium richardsoni. "1947 Hertlein & Strong:
Cardium richardsoni Whiteaves = Cardium
centi/ilosum Carpenter, 1864. "1974 Abbott:
Nemocardium richardsoni (Whiteaves, 1878) =
Nemocardium centifilosum (Carpenter, 1864). "1977
Fischer-Piette: Nemocardium richardsoni Whiteaves.
"2000 Coan et al.: C. richardsoni Whiteaves, 1878 =
Nemocardium (Keenaea) centifilosum (Carpenter,
1864). - CHARACTERS AND OCCURRENCE DESCRIPTIONS. "1878 Whiteaves : shell inflated, but not
quite as thick as high, inequilateral ; outline transversely
and ovately subcircular, length slightly exceeding the
height; posterior side rather long, and more narrowly
rounded than the anterior: beaks large incurved and
approximating; placed a little in advance of the middle.
Surface of the anterior and central portions of the shell
marked by flattened and comparatively broad, radiating
ribs, separated by narrower, deeply impressed lines,
both of which are crossed by faint, concentric striae or
lines growth. On the posterior area, the radiating ribs
are thin, prominent, and much narrower than the
flattened interspaces, and the concentric striations are
developed into elevated, thin and crenulated laminar
ridges. Hinge with two small cardinal and two remote
lateral teeth in each valve, one of which is sublunular.
Pallial line entire; inferior margin of the valves
denticulated all round within. Valves touching at all
points when closed; not open at either extremity.
DIMENSIONS. "1878 Whiteaves: length 8 lines and a
quarter: height seven lines and a half; thickness through
the closed valves, not quite six lines. REMARKS. "1947
Hertlein & Strong: Cardium centifilosum richardsoni
Whiteaves, a northern form of Cardium centifilosum,
is generally a little larger and more coarsely sculptured.
DI STRIBUTION . "1947 Hertlein & Strong : Alaska,
California. "1937 Keen: British Columbia, between
38 and 55 degrees north.
richmondensis Foerste, 1914. - Conocardium
richmondensis Foerste, 1914. Conocardiidae .
Ordovician. RECORDS. "1914 ZR: Conocardium
richmondensis Foerste, 1914 sp. n. Ordovician U.S.A.
riege/i Homes, 1862. - New combination: Lymnocardium riegeli (Homes, 1870). Lymnocardiidae .
Fossil. RECORDS. "1980 Ruhoff: Cardium riege/i
Homes, 1862; [1851 - 1870] vol. 4: 195, pI. 28. Fig. 4.
"1990 Basch (p. 26): designated Cardium riege/i
Hornes, 1870 type species of subgenus
(Zagrabicardium) of genus Limnocardium.
*rigidum - Homonymy to be noticed. Cardium rigidum
from 2 sources: Wood, 1815. - Reeve. 1845.
rigidum of Reeve. 1845 non Wood. - New name
Ctenocardia translatum Prashad, 1932. » Regarding
Cardium rigidum: Cardiidae. Recent. RECORDS. "1845
Reeve,sp. 105: Cardiumrigidum. "1932ZR: Cardium
(Ctenocardia) translatum nom. n. for Cardium rigidum
Rve. non Wood [Ref. Prashad 1932]. "1977 FischerPiette: Reeve, sp. 105: Cardium rigidum = Corculum
translatum Prashad.
rigidum Wood, 1815. [Figs. Part I p. 102, p. 143
erroneous, see below] . - Combination to be revised:
"Cardium" rigidum Wood 1815. OTHER JUDGEMENTS:
Cardium (Ctenocardia) translatum. - Cardium edule.
» Regarding Cardium rigidum: Cardiidae. Recent.
RECORDS. "1815 Wood, p. 236, pI. 57, figs. 2-3:
Cardium rigidum. "1817 Dillwyn: Cardium rigidum
Solander's MSS, Wood fig. "1828 Wood: Cardium
rigidum Wood. "1843 Hanley: Cardium rigidum Wood
"1856 Wood & Hanley, No . 41: Cardium rigidum
Wood, G.c. pI. 57, f. 2, 3. [G.c. = General Conchology
1815]. "1890 Paetel: Cardium rigidum Wood. "192232 Sherborn, p. 5519: Cardium rigidum Wood, 1815.
"1932 Prashad : Cardium rigidum Wood, 1815. "1977
Fischer-Piette: Cardium rigidum Wood. CHARACTERS AND OCCURRENCE - REMARKS.
"1845 Reeve: error regarding Fig. 105 in Part I p. 143.
In legend to the figures , Reeve gave reference to
rigidum Wood Plate XX species 105. However, in the
text he wrote: As the shell here represented proves to
be merely an obscure specimen of the Cardium
papillosum, I take the opportunity of inserting a rare
and very distinct species omitted in the illustration. In
consequence, my explanation to Fig. 105 should read
Cardium papillosum and not Cardium rigidum. "1977
Fischer-Piette: the name rigidum Wood has been used
by Reeve, but for a completely different species (=
Cardium (Ctenocardia) translatum Prashad) . The
original species of Wood was probably Cardium edule.
"My remarks: The species figured by Wood is
definitely not Cardium edule. DISTRIBUTION. "1845
Reeve, "1856 Wood & Hanley: unknown. "1994
Hylleberg: Indo-Pacific.
rimula Meuschen, 1778. Non binominal. - Not available.
RECORDS. "1902 Sherborn, p. 833: Cardium rimula
Meuschen, 1778.
*ringens - Homonymy to be noticed. Cardium ringens
from 4 sources: Bruguiere, 1789. - Bronn, 1848. Wood, 1815. - Defrance (?year).
ringens Bronn, 1848 non Chemnitz. - Synonym of
Cardium hians Brocchi, 1814. » Regarding Cardium
ringens: Cardiidae. Fossil. RECORDS. "1877 Locard:
Cardium ringens 1848 Bronn (non Chemn.) = Cardium
hians Brocchi, 1814.
ringens Bruguiere, 1789. [Figs. Part I p. 73, 88. 99,105,
124, 166, 174. Part 3 p. 843]. - Ringicardium ringens
(Bruguiere, 1789). Cardiidae. Recent. Type species:
Gray (1847) designated Cardium ringens type species
of the invalid genus Papyridea sp. Swainson. West
(1875) designated Cardium ringens Bruguiere, 1789
type species of the subgenus (Bucardium) Gray, 1853.
Moore et al. (1969) quote Fischer (1887) who
designated Cardium ringens (Chemnitz, 1782) type
species of the genus Ringicardium Fischer, 1887 (obj.)
TYPE LOCALITY. Guinea (Spengler 1799); Gambia, Africa
(Sowerby 1841). RECORDS. "1836 Bosc: Cardium
ringens. "1875 von West: Cardium ringens Bruguiere,
1789. "1902 Sherborn, p. 833: Cardium ringens
Bruguiere, 1789. "1930 Stewart: Cardium ringens,
mono typic type species of (Ringicardium) . "1969
Moore et af.,' Cardium (Bucardium) ringens (Bruguiere,
1789) . "1969 Nordsieck: Ringicardium ringens
(Bruguiere, 1789). "1983 Anaya : Cardium ringens
Chemnitz (Bruguiere, 1789) . Type species of
(Bucardium) Gray, 1853. Type designated by von Vest
Phuket Marine Biological Center Special Publication vol. 30 (2004)
739
ringens - Rillgicardium
1875. ··1991 Matsukuma, Okutani & Habe:
Ringicardium ringens (Bruguiere, 1789). ringens
··2001 Vidal, p. 407: Cardium ringens. ··2002
Schneider: Bucardium ringens (Burger, 1789). CHARACTERS AND OCCURRENCE - OF COLLECTIONS. ··ZMUC: Accra tidal zone, Gambia (Cuming's
specimen), Guinea Morck's specimen (Lamarck no. 3),
Angola, off Monrovia, Coast of Congo, off Freetown
15 m, Port Marshall, mangrove, offBrathurst 18 m very
fine sand. ··ZMUC: I specimen and I shells. Spengler
p. 10 Guinea, coil. ··ZMUC: wet collection off
Brathurst, Gambia, 18 m, very fine sand, Atlantide St.
161. dorso-Iateral keels, Manrovia, Liberia 10m,
Atlantide St. 52. DESCRIPTIONS. ··1799 Spengler:
pectunculus orbicularis, ex altero lalere praelongis,
latisque denlibus conspicuus. ··1799 Spengler: shell
orbicular, 26 ribs; posteriorly with 8 ribs, sharp keellike, widely separated, interspaces broad centrally with
nodules or scales transversely extending, centrally with
12 ribs, broad but not rounded, interspaces broad with
a central line, anteriorly with 7 small, low, fine ribs.
Small nymphs. Large and strong lateral teeth, exterior
colour red posteriorly, the rest white. ··200 I Vidal: with
a few additional larger tentacles outside siphons. No
eyes on tentacles. DIMENSIONS. ··1799 Spengler: largest
specimen length and width 44 mm (I inch 4 lines).
HABITAT. ··ZMUC: tidal zone, mangrove, very fine
sand, shells at 0-18 m depth. DISTRIBUTION. ··1819
Lamarck: African coasts and American Seas. ··1844
Reeve: Mouth of the Gambia. ··1856 Wood & Hanley:
Mouth of the Gambia, Africa. ··1884 Grasset: Gabon.
··1891 Romer: Gambia. ··ZMUC: West Africa off
Liberia, Gambia, Guinea, Angola, Congo.
ringens Chemnitz, 1782. [Figs. Part I p. 68,73]. - Non
binominal, to be replaced by Bruguiere, 1789. »
Regarding Cardium ringens: Ringicardium ringens
(Bruguiere, 1789). Cardiidae. Recent. RECORDS. ··1687
Lister, pI. 330, fig. 167: Cardium ringens 10 by
Spengler 1799. ··1757 Adanson, pI. 18, fig. I: Cardium
ringens (fide Spengler 1799) [not available]. •• I 771
Knorr, pI. 14, fig. 3: Cardium ringens 10 by Spengler
1799. ··1780 Argenville: Cardium ringens ID by
Chemnitz . •• I 782 Chemnitz, Tab. 16 Fig. 170: Cardium
ringens. ··1782 Chemnitz. 10 by Gmelin 1791:
Cardium ringens t. 16. f. 170. ··1786 Schroter, No.5:
Cardium ringens citing Chemnitz Choncy!. Th. vi. tab.
16. fig. 170. ··1788 Schroter: Cardium ringens.
[Reference to Chemnitz (1782) fig. 178]. ··1789
Bruguiere, No. 21. Cardium ringens citing Martini.
··1799 Spengler refers Cardium ringens to Lister.
•• I 799 Spengler, p. 10-12, No.4: Cardium ringens
citing Chemnitz Tom. 6. Tab. 16. fig. 170. ··1817
Dillwyn: Cardium ringens Chemnitz. ··1823 Dillwyn,
Tab. 330 fig. 167 Cardium ringens. ··1844 Reeve:
Cardium ringens. ··1856 Wood & Hanley, No. 18:
Cardium ringens citing Ch. vi. 176, f. 170. ··1858
Adams & Adams, p. 454: Cardium (Peclunculus)
ringens citing Chemn. ··1877 Marrat: Cardium
(Peclunculus) ringens citing Chern. ··1884 Grasset:
Cardium ringens Chern. ··1887 Fischer: Cardium
ringens citing Chemnitz (in s. g. Ringicardium, Fischer.
1887). ··1889 Martens: Cardium ringens citing Chemn.
(p. 27: off Liberia, 11-18 m, sand). ··1889 Martens:
Cardium ringens citing Chemn. (p. 29: off Liberia, 108
m, mixed sand). ··1890 Paetel: Cardium ringens Ch.
··1891 Romer: Bucardium ringens Ch. ··1911 Dollfus:
Cardium (Ringicardium) ringens citing Chemnitz.
··1923 Lamy: Cardium (Ringicardium) ringens citing
Chemnitz. ··1927 Lamy: Cardium (Ringicardium)
ringens (citing Chemnitz) Gmelin. ··1945 ZR: Cardium
ringens Chern. [Ref. Lamy 1945]. ··1977 FischerPiette: Cardium (Ringicardium) ringens (citing
Chemnitz) Schroter. ··1999 Knudsen & Hylleberg:
Spengler, 1799 sp. 4: Cardium ringens. CHARACTERS AND OCCURRENCE - CHARACTERS
of diagnostic value: this species is singularly
characterized by the curious manner in which the
interstices between the ends of the posterior ribs remain
open (Reeve 1844). DESCRIPTION. ··1844 Reeve: shell
rounded, sometimes rather depressed, a little gaping at
the posterior side radiately ribbed, about 26 flatly
convex, smooth ribs, posterior interstices peculiarly cut
open near the margins; yellowish-white, partially
stained with pink. HABITAT. ··1844 Reeve: found in
sandy mud. DISTRIBUTION: ··1844 Reeve: Mouth of the
Gambia, Africa
rillgells Defrance. - Synonym of "Cardium"
burdigalinum Lamarck. » Regarding Cardium
ringens: Cardiidae. Fossil. RECORDS. ··1844 Potiez &
Michaud: Cardium ring ens Defr. = Cardium
burdigalinum Lam.
rillgells Gmelin, 1791. [Figs. Part I p. 42, 63, 65]. Changed combination: Rillgicardium ringens
(Bruguiere, 1789). Cardiidae. Recent. RECORDS. ··1687
Lister. ID by Gmelin 179 1: Cardium ringens t. 330. f.
167. ··1769 Knorr. 10 by Gmelin 1791: Cardium
ringens Vergn. 4. t. 14. f. 3. ··1772 Knorr. ID by Gmelin
1791: Cardium ringens Vergn. 6. t. 3 f. 4. ··1782
Chemnitz. ID by Gmelin 1791: Cardium ringens t. 16.
f. 170. ··1791 Gmelin No. 31: Cardium ringens. ··1902
Dall: Cardium ringens Gmelin, type of subgenus
Ringicardium Fischer. ··1902 Sherborn, p. 833:
Cardium ringens Gmelin, 1790 sic. ··1915 Tomlin &
Shackleford: Cardium ringens Gmel., 1791. CHARACTERS AND OCCURRENCE - REMARKS.
Gmelin did not provide the first description. He referred
to Lister, Adanson, Knorr, and Chemnitz. None of these
are available today. - See ringens Bruguiere, 1789.
rillgells Lamarck, 1798. - Cardium ringens Lamarck,
1798. Cardiidae. Recent. RECORDS. ··1798 Bruguiere.
ID by Lamarck, pI. 296. fig. 3. Cardium rillgens. ··1840
Sowerby, catalogue p. I: Cardium ringens Lamarck.
Gambia, Africa. ··1844 Potiez & Michaud: Cardium
ringens Lam. African and American coasts.
rillgells Wood, 1815. - Cardium ringens Wood 1815 non
Gmelin, 1791. Cardiidae. Recent. RECORDS. ··1815
Wood: Cardium ringens Linn. Gmel. ··1922-32
Sherborn, p. 5522: Cardium ringens Wood [non Grn.J,
1815.
Rillgicardium Fischer, 1887 (genus). - Objective
synonym of Bucardium Gray, 1853. Cardiidae. Recent.
RECORDS. ··1969 Moore et al.: Ringicardium =
Bucardium Gray, 1853 . •• I 978 Freneix & Mackie:
Ringicardium Fischer, 1887 = subgenus Bucardium
Gray, 1853. ··1980 Vokes: Ringicardium Fischer, 1887
in subfamily Cardiinae. ··1980. Vokes: Ringicardium
Fischer, 1887 [=Bucardium Gray. 1853]. ··1983 Anaya:
Ringicardium Fischer, 1887. Synonym of(Bucardium)
740
Trop ical Marine Mollusc Programme (TMMP)
Ringieardium - robillardi
Gray, 1853 . Type: Cardium ring ens Chemnitz
(Bruguiere, 1789). Type designated by von Vest 1875 .
Ringieardium Fischer, 1887 (section , subgenus).
Objective synonym of Bucardium Gray, 1853.
Cardiidae. Recent. REMARKS. The designation "section"
takes the value of subgenus. RECORDS. ··1887 Fischer:
(Ringicardium) Fischer, 1887 . Section of genus
Cardium Linne, 1758. ··1930 Stewart: (Ringicardium)
Fischer, 1887
ringiculum Sowerby, 1834. [Figs. Part I p. 108, 144,
174]. - Synonym of Papyridea semisulcata (Gray,
1825). OTHER JUDGEMENTS: Cardium petitianum
Orbigny.» Regarding Cardium ringiculum : Cardiidae.
Recent. TYPE LOCALITY. Hab. ad insulam cセケャッョ@
(Sowerby 1840). RECORDS. ··1834 Sowerby, 48t part,
fig. II: Cardium ringiculum nobis. ··1840 Sowerby,
Proc. Z.S. London p. 106: Cardium ringiculum. ··1840
Sowerby, catalogue p. 2, No. 16: Cardium ringiculum
Sow. "1840 Sowerby, corrected list, p. 8: Cardium
ringiculum (Sowerby No. 16). ··1845 Reeve : Cardium
ringiculum. ··1858 Adams & Adams, p. 456: Cardium
(Papyridea) ringicula Sow. ··1863 Swift: Cardium
ringiculum Sow., 1840 Conch. Icon. sp. 115. Sowerby's
name should be retained as it was published long before
d 'Orbigny's species Cardium petitianum. ··1864 Krebs:
Cardium ringiculum Sowb. = Cardium petitianum
d'Orb. ··1884 Grasset: Cardium ringiculum . Sow.
"1868 ZR: Cardium ringiculum (Sow.) comes from
the West Indies [Ref. Romer 1868]. ··1885 Smith:
Cardium ringiculum Sowerby. 1848 sic = Cardium
(Papyridea) semisulcatum, Gray. 1825. ··1890 Paetel:
Cardium ringiculum Sow. = Papyridea ringicula Sow.
.. 1902 DalJ: Cardium ringiculum Sowerby, 1840 =
Cardium semisulcatum Gray, 1825 . .. 1922-32
Sherborn, p. 5522: Cardium ringiculum Sowerby, 1841 .
"1939 McLean: Cardium ringiculum Sowerby, 1841
= Papyridea semisulcata (Gray, 1825). ··1951 McLean:
Cardium ringiculum Sowerby (1841) = Papyridea
semisulcata (Gray). ··1953 Olsson & Harbison :
Cardium ringiculum Sby. , 1840 = Papy ridea
semisulcatum (Gray, 1825). "1977 Fischer-Piette:
Cardium ringiculum Sowerby = Papyridea semisulcata
Gray. - CHARACTERS AND OCCURRENCE CHARACTERS of diagnostic value : strongly toothed
posterior area (Sowerby 1840). OF COLLECTIONS .
··ZMUC: many shells from St Martin, Krebs colI. no.
699, ··ZMUC : St Thomas. ··ZMUC : label C.
ringiculum Sowerby = C. petitianum d'Orb.
DESCRJPTIONS. ··1845 Reeve: Shell transversely ovate,
sides gaping, anterior very short, posterior elongated;
radiately ribbed, 24-25 narrow, rude, rather
inconspicuous ribs, anterior ribs finely crenulated,
posterior interstices cut near the margins; yellowishwhite, interior sometimes brightly stained with rose on
the posterior side. DIMENSIONS. ··1840 Sowerby: a pretty
little species. REMARKS. Mention of Ceylon as the type
locality by Sowerby (1840) may be erroneous. I have
only seen material from the West Indies in accordance
with Reeve (1845). "1845 Reeve: There seems to be
an error in the locality quoted by Mr. Sowerby for this
interesting little species. HABITAT. ··1845 Reeve: found
in sandy mud. DISTRIBUTION. ··1845 Reeve: Islands of
Nevis and St.Vincent, West Indies ··1884 Grasset:
Singapore. ··ZMUC: South Florida, West Indies,
Brazil; St Martin, St Thomas the West Indies.
ringriseni Jekelius, 1944. - Combination to be revised :
"Cardium" ringriseni Jekelius, 1944. Cardiidae .
Miocene. ··1946 ZR: Cardium ringriseni Jekelius, 1944
sp. nov. (type f. 6) Miocene Soceni Banat.
riogramamense Maury, 1930. - Combination to be
revised : "Cardium " riogramamense Maury, 1930.
Cardiidae. Cretaceous. RECORDS. ··1933 ZR: Cardium
riogramamense Maury, 1930 sp. n. Cretaceous Brazil.
ripleyanum Conrad, 1869. - Changed combination:
Pleuriocardia (Pleuriocardia) ripleyanum (Conrad,
1869). Cardiidae. Cretaceous. TYPE LOCALITY.
Haddonfield, New York. RECORDS. "1869 Conrad, p.
96, PI. 9, fig. 6: Cardium ripleyanum Conrad, 1869.
··1870 Conrad , Cardium ripleyanum Conrad
(Haddonfield, N. J.). ··1978 Scott: Cardium ripleyanum
Conrad. ··1978 Scott: Pleuriocardia (Pleuriocardia)
ripleyanum (Conrad) . ··1980 Ruhoff: Cardium
ripleyanum Conrad, 1869; vol. 5: 96, pI. 9, fig. 6.
ripleyense Conrad, 1858. - Combination to be revised :
"Cardium " ripleyense Conrad, 1858. Cardiidae. Fossil.
RECORDS. ··1980 Ruhoff: Cardium ripleyense Conrad,
1858; vol. 3: 326.
ritter-gulderi (in v alid spelling) . - See under
rittergulderi Steininger, 1963 .
rittergulderi Steininger, 1963. - Cardium rittergulderi
Steininger, 1963. Cardiidae. Miocene. RECORDS. "1963
ZR: Cardium (Cardium) rittergulderi Steininger, 1963
sp. nov. Miocene, Austria.
rivalis Muller. Not a cardiid. « Regarding Tellina
rivalis: RECORDS. ··1803 Montagu : Tellina rivalis
Muller verm. 387. "1803 Montagu : Tellina rivalis. Lin .
Transactions = Cardium amnicum .
roanokensis Perkins, 1961. - Protocardia roanokensis
Perkins, 1961. Cardiidae. Cretaceous. RECORDS. ··1986
Scott: Protocardia roanokensis Perkins, 1961 . ··1961
ZR: Protocardia roanokensis Perkins, 1961 sp. nov.
Cretaceous, Texas, Denton Co .
robid Brusina . - Combination to be changed:
"Cardium" robici Brus. Cardiacea. Fossil. RECORDS.
··1987 Taktakishvili: Cardium robici Brus.
robie; Brusina. - New combination : Lymnocardium
robici (Brus.). Lymnocardiidae . Fossil. RECORDS.
"1987 Taktakishvili: Limnocardium robici Brus.
robillardi Sowerby, 1894. - Changed combination:
Ctenocardia robillardi (Sowerby, 1894). Cardiidae.
Recent. RECORDS. ··1894 Sowerby, p. 47, pI. 4, fig. 20:
Cardium robillardi. "1894 ZR: Cardium robillardi
Sowerby, 1894 n. sp. Mauritius. ··1926 McClelland:
Cardium robillardi Sowerby, 1894. "1937 Viader:
Cardium robillardi Sowerby. ··1977 Fischer-Piette:
Laevicardium robillardi Sowerby. - CHARACTERS
AND OCCURRENCE - DESCRIPTIONS . Shell
subcircular, somewhat solid, medium convex, rounded
anteriorly, subangulate posteriorly, white, irregularly
distributed dark spots, ca. 33 rounded medium elevated
ribs with conspicuous nodose scales, interstices
concentric corrugated. Normal heart shape . With
ligament. White side with coloured stripes, dark redyellow in three stripes [I have translated from Latin].
DIMENSIONS. Antero-post 13 mm (length); umbo-margo
14 mm (width) . REMARKS. A pretty little species, with
numerous raised ribs covered with nodular scales, the
interstices between the ribs being transversely wrinkled.
Phuket Marine Biological Center Special Publication vol. 30 (2004)
741
robusta - robustum vanhyningi
DISTRIBUTION. Mauritius (Sowerby 1897).
robusta (subspecies). - See under brewerii robusta
Weaver & Kleinpell, 1963.
*robustum. - Homonymy to be noticed. Cardium
robustum from 2 sources: Lightfoot in Solander, 1786.
- Shikama, 1964.
robustum Lightfoot in Solander, 1786. [Figs. Part 1 p.
91. Part 3. p. 856]. - Changed combination:
Dinocardium robustum (Lightfoot in Solander, 1786).
OTHER JUDGEMENTS: Cardium robustum Solander, 1786
is a nomen nudum. »Regarding Cardium robustum:
Cardiidae. Upper Miocene and Recent. Dall (1900)
designated Cardium robustum Lightfoot in Solander,
1786 type species of the subgenus Dinocardium Dall,
1900. Rio (1994) Cardium robustum (Lightfoot, 1786)
is type species of genus Dinocardium Dall, 1900. TYPE
LOCALITY. Florida. RECORDS. "1786 Lightfoot in
Solander: Cardium robustum. ··1817 Dillwyn: Cardium
robustum Solander's MSS = Cardium maculatum
Gmelin. ··1900 Dall: Cardium robustum Lightfoot in
Solander, 1786. "1902 Dall: Cardium (Dinocardium)
robustum Solander, 1786 [0. Miocene. Recent]. ··1907
ZR: Cardium robustum [Ref. Hinkley 1907]. "1921
Maury: Cardium (Cerastoderma) robustum Solander,
1786 [U. Miocene. Recent]. "1922-32 Sherborn, p.
5533: Cardium robustum Solander, 1786 [no n.]. "1926
Weisbord: Cardium (Cerastoderma) robustum
So lander. ··1930 Stewart: Cardium robustum
"So lander". "1932 ZR: Cardium robustum [Ref.
Vincent 1932]. "1939 McLean: Cardium
(Dinocardium) robustum (Lightfoot in Solander, 1786).
··1939 ZR: Cardium (Dinocardium) robustum fig. [Ref.
McLean 1939]. "1942 Harry: Cardium robustum
Solander = Cardium muricatum Linne. ··1944 ZR:
Dinocardium robustum Sol. fig. [Ref. Clench & Smith
1944a]. "1945 Smith, Maxwell: Cardium
(Dinocardium )robustum Solander. "1947 Hertlein &
Strong: Cardium (Dinocardium) robustum Solander.
"1953 Olsson & Harbison: Cardium (Dinocardium)
robustum Solander, 1786. "1954 Abbott, p. 40 I:
Dinocardium robustum Solander (jide Rehder 1967).
··1958 Abbott: Laevicardium (Dinocardium) robustum
Solander. ··1960 ZR: Dinocardium robustum internal
volume measurement [Ref. Kornicker 1960]. "1960
ZR: Dinocardium robustum mobility when partially
submerged [Ref. Kornicker & Armstrong 1960]. "1963
ZR: Dinocardium robustum, seasonal distribution and
population statistics on valves deposited on Mustang
Island, Texas [Ref. Kornicker, Wise & Wise 1963].
··1965 ZR: Dinocardium robustum distribution of
opposing valves on Texas beach [Ref. Wise &
Kornicker 1965]. ··1967 Rehder: Portland Catalogue
No. 1358. Cardium robustum, S. (Lister, 328. 186).
··1967 Rehder: Dinocardium robustum (Lightfoot,
1786) valid name, from Florida . ··1967 Rehder:
Cardium robustum (Lightfoot, 1786) = Dinocardium
robustum (Lightfoot, 1786). ··1969 Moore et al.:
Laevicardium (Dinocardium) robust urn (Lightfoot,
1786). ··1970 ZR: Dinocardium robustum first record
in southwestern Texas, USA, marine sp., archaeological
site [Ref. Hester 1970]. ··1971 Andrews : Dinocardium
robustum (Solander, 1786). ··1972 ZR: Dinocardium
robustum [Ref. Tiffany 1972] . ··1974 Abbott :
Dinocardium robustum (Lightfoot, 1786). ··1977
Fischer-Piette: Laevicardium robustum Solander.
··1977 Popov : Cardium robustum Lightfoot, 1786.
Type of Dinocardium Dall, 1900. ··1979 ZR:
Laevicardium (Dinocardium) robustum. Ref. to
Lawrence. "1987 ZR: Laevicardium (Dinocardium)
robustum. Ref. to Wicksten. ··1994 Rio: Dinocardium
robustum S. str. (Lightfoot) [Miocene and Pliocene].
··1997 Shelton: Dinocardium robustum (Lightfoot,
1786). ··2002 Schneider: citing Dall (1900): Cardium
(Cerastoderma) robustum Solander, 1786, section
Dinocardium . .. 200 I Schneider & Carter: Dinocardium
robustum . Cardiidae . Recent . "2002 Schneider:
Dinocardium robustum (Lightfoot, 1786). CHARACTERS AND OCCURRENCE - OF COLLECTIONS. ··ZMUC: 33 shells from shore on Sapelo Island,
Georgia; 2 shells from shore, Florida, GulfofMexico.
DIMENSIONS. ··1939 McLean: 104 x 119 x 87 mm; ribs
about 35. REMARKS. ··1930 Stewart: Cardium robustum
"Solander", apparently the first valid name for
Dinocardium chipolanum (Dall) [Miocene].
DISTRIBUTION . "1947 Hertlein & Strong: it is a
Caribbean species and has not been recorded as
occurring in the Gulf of California. Virginia East USA
to Belize, British Honduras, and in the West Indies
south to Jamaica.
robustum Shikama, 1964. - Synonym of Laevicardium
obliquum (Spengler, 1799). » Regarding Cardium
robustum: Cardiidae. Recent. RECORDS. ··1964
Shikama, p. 71, pI. 43, fig. 6: Cardium robustum. ··1977
Fischer-Piette: Cardium robustum Shikama =
Laevicardium (Dinocardium) obliquum (Spengler,
1799).
robustum hazeli Ward & Blackwelder, 1987
(subspecies). - Dinocardium robustum hazeli Ward &
Blackwelder, 1987. Cardiidae. Pleistocene. TYPE
LOCALITY. North Carolina. RECORDS. "1987 Ward &
Blackwelder, p. 153, fig.: Dinocardium robustum hazeli
subsp . nov.
robustum vanhyningi Clench & Smith, 1944
(subspecies) . [Figs. Part I p. 91]. - Dinocardium
robustum vanhyningi Clench & Smith, 1944. OTHER
JUDGEMENTS: Laevicardium obliquum. » Regarding
Cardium robustum vanhyningi: Cardiidae. Recent. OF
COLLECTIONS. ··ZMUC: 4 specimens, Sanibel Island,
Florida. RECORDS. ··1944 Clench & Smith, p. II (pI.
7): Dinocardium robustum vanhyningi n. subsp. ··1944
ZR: Dinocardium robustum vanhyningi Clench &
Smith, 1944b subsp. nov. Sanibel Island, Florida.
··1948 Aguayo & Jaume: Cardium robustum
vanhyningi Clench & Smith. ··1950 Baker: Cardium
robustum vanhyningi Clench & Smith. ··1954 Abbott:
Dinocardium robustum vanhyningi Clench & Smith.
··1953 Olsson & Harbison: Dinocardium robustum
vanhyningi Clench & Smith, 1944 = Cardium
(Dinocardium) robustum Sol ander, 1786. ··1955 Perry
& Schwengel: Dinocardium robustum vanhyningi
Clench & Smith . ··1968 Abbott: Dinocardium
robustum vanhyningi Clench & Smith . ··1972
Shoemaker: Dinocardium robustum vanhyningi Clench
& Smith . ··1974 Abbott: Dinocardium robustum
vanhyningi Clench & L. C. Smith, 1944. ··1977
Fischer-Piette, p.175: Cardium robustum vanhyningi
Clench & Smith = Laevicardium obliquum Solander
sic. "1994 Rio: Dinocardium robustum vanhyningi
742
Tropical Marine Mollusc Programme (TMMP)
roehebruni - roseum
Clench and Smith 1944.
roehebruni Coquand, 1859. - Combination to be revised:
"Cardium" roehebruni Coquand , 1859. Cardiidae.
Fossil. RECORDS. "1980 Ruhoff: Cardium roehebruni
Coquand, 1859; vol. 16: 983.
rogata (error). - Printer's mistake for rugala in Vidal
1994.
rogenhoferi Brusina & Bru s ina , 1884. - New
combination: Lymnoeardium rogenhoferi (Brusina &
Brusina, 1884). Lymnocardiidae. Fossil. RECORDS .
"1990 Basch: Limnoeardium (Zagrabieardium)
rogenhoferi (Brusina & Brusina, 1884).
rogersi Rennie, 1936. - Combination to be revised:
"Cardium" rogersi Rennie, 1936. Cardiidae. Lower
Cretaceous. RECORDS. "1936 ZR: Cardium rogersi
Rennie, 1936 sp. n. Lower Cretaceous. Northern
Zululand.
rosea (variety). - See nodosum var. rosea Lamarck.
rosemariensis Wilson & Stevenson, 1977. [Figs. Part 3
p. 878]. - Synonym of Acrosl erigma impolitum
(Sowerby, 1833) . » Regarding Aeroslerigma
rosemariensis : Cardiidae . Recent. TYPE LOC ALITY .
Dampier Archipelago, Western Australia . TYP E
REPOSITORY. Holotype, Western Australian Museum,
650-66. TYPE MATERIAL. Holotype in WAM, a specimen
from Rosemary Island, Western Australia . Ten
para types, all from the northern coast of Western
Australia, from Point Cloates to Dampier Archipelago
(Vidal 1999). RECORDS. "1977 Wilson & Stevenson,
p. 95-97 (pI. 6 , figs . 23-24): Aeroslerigma
rosemariensis sp. nov. "1978 ZR: Aeroslerigma
rosemariensis sp. nov. Western Australia. Ref. to Wilson
et al. p. 95. fig'd. '·1999 Vidal, p. 305: Wilson &
Stevenson 1977, p. 95, pI. 6, figs 23-24: Aeroslerigma
rosemariensis = Aeroslerigma impolitum (Sowerby,
1833). - CHARACTERS AND OCCURRENCE DIMENSIONS. Length 17.0 mm, height 20.2 mm, width
(calculated) 11 .8 mm (Vidal 1999). DISTRIBUTION. West
Australia.
roseolum Melvill, 1909. [Figs. Part 3 p. 907]. - Fragum
roseolum (MeJvill , 1909). Recent, Cardiidae . TYPE
REPOSITORY. Holotype. BMNH London. TYPE LOCA LITY.
Cargados Carajos, western Indian Ocean. RECORDS.
"1909 Melvill , p . 134 (pI. 5, fig. 14): Cardium
(Fragum) roseolum sp. nov. "1909 ZR: Cardium
(Fragum) roseolum Melvill, 1909 sp. n. Cargados
Carajos. "1937 Viader: Cardium roseolum, Melvill.
"1977 Fischer-Piette: Coreulum (Afroeardium)
roseo/um Melvill. "2002 Schneider: Afroeardium
roseo/um (Melvill, 1909). Holocene. - CHARACTERS
AND OCCURRENCE - DESCRIPTIONS. "1909 Melvill :
shell equivalve, inequilateral, inflated, squarrose
contour, ventral and dorsal margins almost parallel, the
posterior margin much shortened. The whole surface
is closely scaly-ribbed, these ribs are 40 in number. A
clear rose colour. DIMENSIONS. Alt. 7, lat. 6, diam. 5
mm (Melvill 1909). REMARKS. Ribs are of much the
same character as exists in C. erenelloides, save that
they appear more persistent and imbricate, not being
of so delicate a substance. The margin of the valves
also is more deeply serrated or pectinate than in the
allied species, of which, indeed, it might be considered
a variety, but the great difference of form and colour, a
clear rose, have prompted the differentiation (Melvill
1909). DISTRIBUTION. Red Sea, Indo-West Pacific .
*roseum . - Homonymy to be noticed. Cardium roseum
from 2 sources: Gmelin, 1791. - Lamarck, 1819.
roseum Chemnitz, 1782. [Figs. Part 1 p. 45, 71, 103,
126]. Non binomina1 to be replaced by Gmelin, 1791 .Synonym of Coreu/um eardissa (Linne, 1758). OTHER
JUDGEMENTS: Coreulum humanum Roding, 1798. Coreulum impressum (Lightfoot, 1786). - Cardium
junoniae Lamarck . « Regarding Cardium roseum:
Cardiidae. Recent. RECORDS. "1742 Argenville. ID by
Gmelin 1791: Cardium roseum conch. t. 23. f. D. "1782
Chemnitz, Tab. 14 Fig. 147-148 : Cardium roseum.
"1782 Chemnitz. ID by Gmelin 1791 : Cardium roseum
t. 14 . f. 147. "1788 Schroter: Cardium roseum
[Reference to Chemnitz (1782) fig . 154]. "1789
Bruguiere: Cardium roseum. Martini tom. 6. pag. 154,
tab . 14, fig. 147, 148. "1791 Gmelin p. 3245, No.3:
Cardium roseum (reference to Chemnitz fig. 147).
"1815 Wood p. 239, No 48: Cardium roseum Linn.
Gme!. (reference to Chemnitz fig . 147) . "1817
Di IIwyn: Cardium roseum. Chemnitz. "1823 Di IIwyn,
Tab. 320 fig. 157 Cardium roseum . .. 1825 Wood p. 27
No. 48 : Cardium roseum (reference to Chemnitz fig.
147). "1840 Pfeiffer: Chemnitz Fig. 147-148. Cardium
roseum = Cardium Junoniae 2. Lam. 47 . .. 1844 Reeve:
Cardium roseum Chemnitz = Cardium eardissa
Linnaeus var. delta. "1856 Wood & Hanley, No. 48:
Cardium roseum Wood (as of Ch.) G.C. pI. 57, f. 6.
[G.c. = General Conchology 1815]. "1902 Sherborn,
p. 836: Cardium roseum Gmelin, 1790 sic. "1947
Bartsch: Coreulum ros eum (Chemnitz, 1782) =
Coreulum humanum R6ding, 1798. "1977 FischerPiette: Cardium roseum Gmelin = Cardium eardissa
Linne. "1977 Habe: Cardium roseum Gmelin, 1791 =
Coreulum impressum (Lightfoot, 1786)... 1999 Higo
el al.: Cardium rosellm Gmelin, 1791 = Coreu/um
impressum (Lightfoot, 1786). - CHARACTERS AND
OCCURRENCE - OF COLLECTI ONS. "ZMUC:
Chemnitz 6, 147-148, = junoniae, beta Lamarck sine
loco REMARKS. Chemnitz (1782) is not binominal.
DISTRIBUTION. China, Eastern Seas (Reeve 1844); Gulf
of Bengal (Wood & Hanley 1856).
roseum Chemnitz, 1782. (variety). Non binominal- Not
available. RECORDS. "1890 Paetel : Hemieardium var.
rosellm Ch.
roseum Lamarck, 1819. [Figs. Part 3 p. 915]. - Synonym
of Parvieardium seabrum (Philippi, 1836). OTHER
JUDGEMENTS: dubious species. - Cardium nodosum
Turton, 1822. - Plagioeardium papillosum (Poli, 1791).
» Regarding C. roseum: Cardiidae. Recent. TYPE
LOCALITY . Manche, close to Cherbourg. TYPE REPOSITORY.
Cabinet of de France's, unknown availability. RECORDS.
.. 1819 Lamarck, No . 37: Cardium roseum . .. 1844
Potiez & Michaud : Cardium roseum Lam. French
Atlantic coast. .. 1867 ZR: Cardium roseum (Lam) =
nodosum (Turt. and other authors) [Ref. Weinkauff
1867] ... 1869 Romer: Cardium roseum Lamarck; a
dubious species ... 1878 Monterosato: Cardium roseum ,
Lk. = C. nodosum, Turt. = C. seabrum, Ph. "1880
Stossich: Cardium roseum in Weinkauff 1867 =
Cardium nodosum Montagu . "1888 Servain: Cardium
roseum Lamarck, 1818. "1890 Paetel: Cardium roseum
Lm. = nodosum Mtg. "1922-32 Sherborn, p. 5548:
Cardium roseum Lamarck, 1819. "1937 Coen: roseum
Phukel Marine Biological Center Special Publication vol. 30 (2004)
743
roseum - rotundatum
Lamarck = Cerastoderma (Rudicardium) nodosum
Turton. "1969 Nordsieck: Parvicardium roseum
(Lamarck, 1819). "1977 Fischer-Piette: Cardium
roseum Lamarck = Cardium nodosum Montagu.
"Scarlato & Starobogatov: Cardium roseum (Lam) in
Ostroumoff 1893 = Plagiocardium papil/osum (Poli,
1791). ··2000 Aartsen & Goud: Cardium roseum
Lamarck, 1819 is a junior homonym of Cardium
roseum Gmelin, 1791. - CHARACTERS AND
OCCURRENCE - REMARKS. Cardium roseum
Lamarck, 1819 = Parvicardium nodosum (Turton,
1822) = Parvicardium scabrum (Philippi, 1836)
(Nordsieck 1969). HABITAT. The species C. nodosum
9--420 m depth, in the North Sea from 42 m depth and
down (Nordsieck 1969). DISTRIBUTION. C. nodosum
North Norway, Ireland, Canary Islands, Mediterranean
(Nordsieck 1969).
roseum Lamarck (variety). - Synonym of Cardium
scabrum Phil. RECORDS. "1867 ZR: a variety of
Cardium roseum (Lam) = scabrum (PhiL) [Ref.
Weinkauff 1867].
roseum (variety). - See under papillosum var. roseum
Pallary.
Rosphoricardium (error) - Printer's mistake for
subgenus Bosphoricardium in CeJidje 1953.
rossae Stephenson, 1941. - Combination to be revised:
"Cardium" rossae (Stephenson, 1941). Cardiidae.
Cretaceous. RECORDS. ,·1946 ZR: Cardium
(Granocardium) rossae Stephenson, 1941 sp. nov.
Navarro fonnations, Cretaceous, Texas, Neylandville
marl Tex Bu. "1947 ZR: Cardium (Granocardium)
rossae Stephenson, 1941 sp. nov. Upper Cretaceous,
Navarro Group Texas.
rossi Marwick, 1944. - Changed combination:
Trachycardium rossi (Marwick, 1944). Cardiidae.
Tertiary. RECORDS. "1944 Marwick: Ovicardium rossi.
"1944 ZR: Trachycardium (Ovicardium ) rossi
Marwick, 1944a sp. nov. genotype of (Ovicardium )
Marwick, 1944 (Opoitian) Te Reinga Falls, Hangarua
R. "1973 ZR: Acrosterigma (Ovicardium). rossi
(Marwick 1944), fossiL Ref. to Beu . "1977 Popov:
Trachycardium (Vasticardium) rossi Marwick, 1944.
"1977 Wilson & Stevenson: Trachycardium
(Ovicardium) rossi Marwick, 1944. "1978 Maxwell:
Trachycardium (Ovicardium) rossi Marwick, 1944.
ross;; Abbass, 1978. - Cardium rossii Abbass, 1978.
Cardiidae. Upper Eocene . RECORDS . "1978 ZR:
Cardium (Cardium) rossii Abbass, 1978 sp. nov. Egypt.
Ref. to Abbass p. 56.
rossolum (error) . - Printer's mistake for roseo/um
Melvill in Cotton 1961, p. 241.
rostrata Sinz. - Prosodacna rostrata (Sinz.). Lymnocardiidae. Pliocene. RECORDS . "1960 ZR: Prosodacna
rostrata (Sinz.) Pliocene Ukraine, Odessa USSR. [Ref.
Eberzin 1960].
*rostratum. - Homonymy to be noticed. Conocardium
rostratum from 2 sources : Bronn, 1848. - Orbigny,
1850.
rostratum Bronn, 1848. - Conocardium rostratum
Bronn, 1848. Conocardiidae. FossiL RECORDS. "192232 Sherborn, p. 5555: Arcites rostratus Martin, 1809.
"1922-32 Sherborn, p. 5556: Conocardium (Martin)
rostratum Bronn, 1848. - Arciles.
rostratum Deshayes, 1853. - Combination to be revised:
"Cardium" rostra tum Deshayes, 1853. ?Cardiacea.
Fossil. RECORDS. "1980. Ruhoff: Cardium rostratum
Deshayes, 1853; voL 2: 48.
rostratum Orbigny, 1850. - Conocardium rostratum
d'Orbigny, 1850. Conocardiidae. FossiL RECORDS.
"1922-32 Sherborn, p. 5555: Cardium (Mart.) rostratus
de Koninck, 1841. - Arcites. "1922-32 Sherborn, p.
5556: Conocardium (Koninck) rostratum d'Orbigny,
1850. - Cardium.
rothi Halavats, 1887. - New combination:
Lymnoeardium rothi (Halavats, 1887). Lymnocardiidae.
Pannonian. RECORDS. "1966 ZR: Limnocardium rothi
Pannonian, Mecsek Mts. Yugoslavia, pIs. [Ref. Bartha
1966]. "1990 Basch: Limnocardium (Zagrabicardium)rothi (Halavats, 1887).
rotulum Linnaeus (error). - Corculum rotulum
(Linnaeus) in Nguyen et al. (1997). Probably misprint
of C. roseum. RECORDS. ··2003 Hylleberg & Kilburn,
p. 177: Corculum rotulum (Linnaeus). This name does
not exist.
rotunda Harper, 1947. - Plethocardia rotunda Harper,
1947. Cardiacea. FossiL RECORDS. "1947 ZR: Plethocardia rotunda Harper, 1947 sp. n. "Caradoc" ofY nys
Gabel, Caernarvonshire.
rotunda Kvaliashvili, 1978. - Digressodacna rotunda
Kvaliashvili, 1978. Lymnocardiidae. Pliocene.
RECORDS . "1978 ZR: Digressodacna rotunda
Kvaliashvili, 1978 sp. nov. Georgia SSR. Ref. to
Kvaliashvili p. 63.
rotunda. - See under norvegicum rotunda Jeffreys, 1863.
*rotundatum. - Homonymy to be noticed. Cardium
rotundatum from 2 sources: Dujardin, 1837. Carpenter, 1857.
rotundatum Carpenter, 1857. - Synonym of
Trachycardium panamense (Sowerby, 1833). OTHER
JUDGEMENTS: Trachycardium procerum (Sowerby). »
Regarding Cardium rotundatum: Recent, Cardiidae.
RECORDS. "1857 Carpenter: Cardium rotundatum.
"1860 Carpenter: Cardium rotunda tum Cpr. "1886
Dall: Cardium rotundatum Carpenter = C. proeerum
juv. ··1909 Lamy: Cardium rotunda tum Carpenter
(1855-57) = Cardium procerum Sowerby. "1901 Dall:
Cardium rotundatum Carpenter, 1857 (testa juv.) =
Cardium procerum Sowerby, 1833. "1910 Dall :
Cardium rotundatum Carpenter = Cardium procerum
junior. "1947 Hertlein & Strong: Cardium rotundatum
Carpenter = Cardium (Mexicardia) procerum Sowerby.
"1951 Winkle: Cardium rotundatum Carpenter. "1961
Olsson: Cardium rotundatum Carpenter, 1857 =
Mexicardia procera (Sowerby, 1833). "1968 Keen:
Cardium rotundatum Carpenter, 1857 = Trachycardium
(Mexicardium) panamense (Sowerby, 1833). "1968
Velez: Cardium rotundatum = Trachycardium
procerum (Sowerby, 1833). "1971 Keen & McLean:
Cardium rotundatum Carpenter, 1857 = Traehycardium
(Mexicardia) panamense (Sowerby, 1833). "1977
Fischer-Piette: Cardium rotundatum Carpenter =
Laevicardillm procerum Sowerby). "1980 Ruhoff:
Cardium rotundatum Carpenter, 1857 [1855-1857] p.
531. - CHARACTERS AND OCCURRENCE DISTRI-BUTION. As for C. panamense San Ignacio
Lagoon in Baja California to Costa Rica, not reaching
Panama (Keen & McLean 1971.
rotundatum Dujardin, 1837. - Combination to be
744
Tropical Marine Mollusc Programme (TMMP)
rotundum - rubicundum
revised: "Cardium" rotundatum Dujardin, 1837.
Cardiidae. Miocene. REcORDs. --1877 Locard: Cardium
rotundatum Dujardin, 1837. --1922-32 Sherborn, p.
5564: Cardium rotundatum Dujardin, 1837. --1999
Jacques Vidal (pers. com.) MNHN, Paris: Fossils from
the Paleontological Collection. Cardium rotunda tum
Dujardin. Pontilevien. France (Indre and Loire),
B45417.
rotundum Senes, 1955. - Lymnocardium rotundum
(Senes, 1955). Lymnocardiidae. Miocene. RECORDS.
--1955 ZR: Cardium (Limnocardium) rotundum Senes,
1955 sp. nov. Miocene, Czechoslovakia.
rotundum (variety). - See under norvegicum rotundum
Jeffreys. - papil/osum var. rotundum Locard.
rouau/ti Bellardi, 1852. - Combination to be revised:
"Cardium" rouaulti Bellardi, 1852. Cardiidae. Fossil.
REcoRDs. --1980 Ruhoff: Cardium rouaulti Bellardi,
1852; vol. 4: 240 pI. H (sic = 19), fig. I.
rousseaui Mayer Combination to be revised: "Cardium"
rousseaui Mayer. Lymnocardiidae. Fossil. REcORDs.
--1886 Fischer: Cardium rousseaui Mayer.
rouyandum Orbigny, 1847. - Changed combination:
Vepricardium rouyandum (Orbigny, 1847). Cardiidae.
Fossil. RECORDS. --1977 Popov: Bucardium
(Vepricardium) rouyandum sic (Orbigny, 1847).
rouyanum Orbigny, 1850. - Combination to be revised:
"Cardium" rouyanum Orbigny, 1850. Cardiidae.
Eocene. RECORDS. --1886 Cossmann: Cardium
rouyanum d'Orbigny. --1922-32 Sherborn, p. 5572:
Cardium rouyanum d'Orbigny, 1850.
rubel/um Chiereghini in Nardo, 1847. - Combination
to be revised: "Cardium" rubellum Chiereghini in
Nardo, 1847. Cardiidae. Fossil. RECORDS. --1922-32
Sherborn, p. 5575: Cardium rubellum Nardo (ex
Chiereghini MS), 1847.
rubescens Smith, 1911. [Figs. Part 3 p. 917]. - Changed
combination: Afrocardium rubescens (Smith, 1911).
OTHER JUDGEMENTS: Afrocardium skeeti (Hedley, 1906).
- Afrocardium richardi (Audouin, 1827»> Regarding
Cardium rubescens: Cardiidae. Recent. TYPE
REPOSITORY. 2 syntypes. BMNH London. TYPE LOCALITY.
Christmas Islands, Flying Fish Cove, in 30 fathoms.
RECORDS. --1911 Smith: Cardium (Fragum) rubescens.
--1911 ZR: Cardium (Fragum) rubescens Smith, 1911
sp. n. Christmas Island Indian Ocean. --1926
McClelland: Cardium rubescens Smith, 1911 n. sp.
--1934 Tomlin: Cardium (Fragum) rubescens Smith.
--1977 Fischer-Piette: Corculum (Afrocardium)
rubescens Smith. --1992 Lamprell & Whitehead:
Cardium rubescens Smith, 1911 = Afrocardium skeeti
(Hedley, 1906). --2002 Schneider: Afrocardium
rubescens (Smith, 1911). Holocene. --2003 Hylleberg
& Kilburn, p. 176: C. rubescens Smith, 1911. Syn of
Afrocardium richardi (Audouin, 1827) according to J.
Vidal. - CHARACTERS AND OCCURRENCE DESCRIPTIONS. --1911 Smith: Shell small, thin,
inequivalve, triangular--cordate, short in front, posterior
produced, somewhat pointed, reddish; sometimes
completely reddish, about 38 radiating ribs, little
elevated around the umbo, delicately squamose, close
set until separated by furrows; anterior and posterior
ribs narrow, densely squamose, intercostal areas with
transverse sculpture; thin valves, slightly dentate along
central part of ventral margin, stronger denticulation
posteriorly; interior of valves white, posteriorly reddish,
shells have reddish colour in rays (I have translated
from Latin]. DIMENSIONS. --1911 Smith: length 8 mm,
width 6.5 mm, diameter 5.5 mm. REMARKS. --1911
Smith: one specimen entirely rosy-red, the other with
the front half ofthe valves almost white, and the hinder
portion stained rosaceous, the colour in two more or
less suffused rays. The radiating riblets upon the middle
ofthe valves are faintly convex, and separated by mere
impressed lines. These become more distinct grooves
both anteriorly and behind. The lines of growth are
distinct in the grooves, and in the impressed striae,
produce an almost punctuate appearance. Two
specimens were studied by Smith 1911. Allied to C.
crenelloides Melvill, and carditaeformis Reeve, both
Indian Ocean species. However, carditaeformis is
longer, the ribs are finer and more numerous, and the
is colour different; carditaeformis is narrower in front,
and less acuminate behind, has more ribs, and is devoid
of colour. (Smith 1911). HABITAT. In 30 fathoms.
DISTRIBUTION. East Indian Ocean (authors).
rubicundum Reeve, 1844. [Figs. Part I p. 132, 173. Part
3 p. 865, 926]. - Trachycardium rubicundum (Reeve,
1844). Cardiidae. Recent. TYPE LOCALITY. Zanzibar, east
coast of Africa (Reeve 1844). TYPE REPOSITORY. 3
syntypes. 1978128. BMNH London. RECORDS. --1844
Reeve, p.169: Cardium rubicundum. --1844 Reeve,
Conch. Icon. sp. 44: Cardium rubicundum. --1884
Grasset: Cardium rubicundum. Rv. --1890 Paetel:
Cardium rubicundum Rv. --1897 Sowerby: Cardium
rubicundum Reeve. --190 I Sturany: Cardium
rubicundum Rve. (possible synonym of Cardium
rugosum). --1906 Melvill & Standen, p. 837: Cardium
(Trachycardium) rubicundum Reeve. --1922-32
Sherborn, p. 5587: Cardium rubicundum Reeve, 1844.
--1955 Rost & Soot-Ryen: Trachycardium rubicundum
Reeve, 1844. --1956 ZR: Trachycardium rubicundum
Reeve, new to Seychelles [Ref. Rost & Soot-Ryen
1956]. --1964 Spry: Cardium (Vasticardium)
(Regozara) rubicundum (Reeve). --1974 Habe &
Kosuge: Vasticardium (Regozara) rubicundum (Reeve,
1844). --1977 Fischer-Piette: Laevicardium
rubicundum Reeve. --1977 Wilson & Stevenson:
Cardium rubicundum Reeve, 1844. --1981 Koyama et
at..' Cardium rubicundum Reeve, 1844 = Laevicardium
biradiatum (Bruguiere, 1789). --1982 Kilburn &
Rippey: Trachycardium rubicundum (Reeve, 1844).
--1992 Oliver: Trachycardium rubicundum. --1992
Oliver: Cardium rubicundum Reeve, 1844 in Shopland
(1896) = Trachycardium lacunosum (Reeve, 1845).
--1992 Oliver: Cardium rubicundum Reeve, 1844 in
Lamy (1927) = Trachycardium lacunosum (Reeve,
1844). --1998 Vidal: Vasticardium rubicundum (Reeve,
1844). --1999 Aungtonya et al., PMBC-SP 19(2): 378:
Trachycardium rubicundum (Reeve, 1845). --1999
Vidal: Vasticardium rubicundum (Reeve, 1844). --2000
Hylleberg, p. 532: Vasticardium rubicundum (Reeve,
1845). Det. 1. Vidal. --2003 Hylleberg & Kilburn, p.
185, Plate 4, fig. 16: Vasticardium rubicundum (Reeve,
1845). - CHARACTERS AND OCCURRENCE - OF
COLLECTIONS. Madagascar H. Cuming Coli. 7 specimens
Acc. No. 1829 BMNH reg. no. 1844.6.3. 105-110.
CHARACTERS of diagnostic value: 37 ribs. DESCRIPTIONS.
--1844 Reeve: Shell oblong-ovate, very slightly
Phuke/ Marine Biological Cen/er Special Publica/ion vol. 30 (2004)
745
rubicundus - rubrum
oblique, radiately ribbed, 37 sharply convex ribs, the
anterior third portion squamosely crenated, the middle
portion obtusely scaled on both sides, the posterior
portion obliquely tubercled; bright red, whitish towards
the umbones spotted with reddish brown, margins
edged with bright pinkish red interiorly. (Description
in Latin as well as in English). Cuming' specimen has
6-7 ornamented posterior ribs. REMARKS. --1844 Reeve:
An extremely pretty species, remarkable for its vivid
colouring and for the elaborate character of its
sculpture. -- Koyama et al. These authors indicate
erroneous synonymy. Trachycardium rubicundum
(Reeve, 1844) is a valid species. HABITAT. --1906
Melvill & Standen: at 3-10 fathoms depth, sandy mud.
DISTRIBUTION. --1844 Reeve: Zanzibar, E. Africa. --1884
Grasset: Natal, Zanzibar. --1897 Sowerby: Indian
Ocean. --1999 Aungtonya et al.: Thailand, Andaman
Sea . "1999 Vidal: Indonesia (Lombok). --2000
Hylleberg and --2003 Hylleberg & Kilburn: Vietnam.
rubicundus. - Not in Cardiacea. Glossus nlbicundus.
RECORDS. "1815 Oken, p. 235: genus Glossus with
Chama cor, Glossus rubicundus [Oken is not available].
--1818 Schmidt, p. 158: Glossus rubicundus (Chama
cor).
rubiginosum Gmelin 1791. [Figs. Part I p. 50]. - Nomen
dubium. » Regarding Cardium rubiginosum : not a
cardiid. RECORDS. --1742 Gualtieri, pI. 43 fig. D:
Concha cordiforrnis D. rubiginosa. "1742 Gualtieri .
ID by Gmelin 1791: Cardium rubiginosum t. 83. f. D.
"1791 Gmelin p . 3256 (fide Wood 1815). "1817
Dillwyn: Cardium rubiginosum Gmelin should be
ignored. --1828 Wood: Cardium rubiginosum. --1856
Wood & Hanley, No. 44: Cardium rubiginosum Gmel.
3256, from Gualt. 83, f. D. irrecognisable. --1856 Wood
& Hanley: Cardium rubiginosum Gmel. = C. edule var.
= irrecognisable. --1902 Sherborn, p. 844: Cardium
rubiginosum Gmelin, 1790 sic. --1977 Fischer-Piette:
Cardium rubiginosum Wood = not a Cardium . CHARACTERS AND OCCURRENCE - DISTRIBUTION. Mediterranean (Wood & Hanley 1856).
rubiginosum Wood, 1815. - See under rubiginosum
Gmelin 1791.
rubraquale Tscheltzov, 1965. - Avicardium rubraquale
Tscheltzov, 1965. Cardiacea. Pliocene. RECORDS. --1965
ZR : Avicardium rubraquale Tscheltzov, 1965 sp. nov.
Pliocene, Turkmenia, USSR.
rubricatti Almera y Bofill, 1894. - Combination to be
revised: "Cardium" rubricatti Almera y Bofill, 1894.
Cardiacea . Tertiary. RECORDS. --1983 Anaya:
"Cardium" rubricatti Almera y Bofill, 1894.
rubrohamatum Voskuil & Onverwagt, 1988. [Figs. Part
3 p. 881]. - Vepricardium rubrohamatum Voskuil &
Onverwagt, 1988. Cardiidae. Recent. DISTRIBUTION.
Philippines. RECORDS. --1988 Voskuil & Onverwagt:
Vepricardium rubrohamatum .
rubropictum Habe & Kosuge, 1966. - Synonym of
Acrosterigma biradiatum (Bruguiere , 1789). »
Regarding Laevicardium rubropictum: Cardiidae.
Recent. TYPE MATERIAL. --2000 Vidal: holotype, a
bivalved specimen from Bentayan Island, Philippines,
in ZBM No 53798; seven bivalved paratypes, all from
the type locality, in various collections. TYPE REPOSITORY.
Holotype and two paratypes in National Science
Museum (Tokyo). RECORDS. "1966 Habe & Kosuge,
p. 339 (pI. 29, fig . 20): Laevicardium rubropictum.
--1966 ZR: Laevicardium rubropictum Habe &
Kosuge, 1966 sp. nov. Samboanga , Mindanao,
Philippines. --1974 Habe & Kosuge: Laevicardium
rubropictum Habe et Kosuge. --1977 Fischer-Piette:
Laevicardium rubropictum Habe & Kosuge =
Laevicardium biradiatum Bruguiere. --1977 Habe:
Laevicardium rubropictum Habe & Kosuge, 1966.
--1999 Higo et al. : Laevicardium rubropictum Habe &
Kosuge, 1966 . "1999 Vidal: La e vicardium
rubropictum Habe & Kosuge, 1966: 153, pI. 59, fig.
2: = Acrosterigma biradiatum (Bruguiere, 1789).
--2000 Vidal, p . 460, Fig. 7: Vepricardium
rubrohamatum Voskuil & Onverwagt, 1988: 86, pl. 1,
fig . 1-3. - CHARACTERS AND OCCURRENCEDESCRIPTIONS. --2000 Vidal: V rubrohamatum is
characterised by its very oval shape , strongly
prosogyrate umbos, thinner shell and hinge, and higher
rib number with very elaborate ornamentation.
Distribution: - In addition to the type series from the
Philippines, only one shell from Visayas Islands,
Philippines is known in the MNHN, collection Vidal.
REMARKS. 1966. Habe et Kosuge: Laevicardium
rubropictum Habe et Kosuge, 1960, ally of
Laevicardium pulcherrimum Sakurai & Habe, 1966.
Habe & Kosuge state that Laevicardium rubropictum
differs from Laevicardium biradiatum which has many
fleshy reddish brown patches of various sizes and lacks
the large reddish brown patch on the posterior part of
the surface. Fischer-Piette (1977, p. 25) maintains that
colour variation is a poor reason to erect a new species,
because this character is too variable. HABITAT: intertidal
to 20 m, sand and mud (Higo et al. 1999). DISTRIBUTION.
Indo-Pacific, Red Sea, Indo-West Pacific (Hylleberg
1994); Kii peninsula (Kuroda & Habe, 1981) and
southwards, East China Sea, South China Sea,
Philippines, Indo-Pacific (Higo et al. 1999).
rubrum Chemnitz, 1782 [Figs. Part 1 p. 73 No. 168169]. Non binominal. - Not available. » Regarding
Fragum rubrum: Cardiidae. Recent. RECORDS. --1782
Chemnitz, Tab. 16 Fig. 168-169: Fragum rubrum.
Cardium unedo Lin. --1789 Bruguiere : Fragum
rubrum , Cardium unedo Linnaei; Martini tom. 6. tab.
16, fig. 168, 169. "1799 Spengler: Fragum rubrum
Chemnitz Tom. VI. Tab. 16 Fig. 168-169 = Cardium
unedo Lin. --1932 Prashad: Fragum rubrum (Chemnitz,
1782) = Fragum unedo (Linne, 1758). --1977 Wilson
& Stevenson: Fragum rubrum Chemnitz, 1782; junior
synonym of Fragum unedo (Linnaeus, 1758). --1999
Knudsen & Hylleberg: Spengler, 1799 Fragum rubrum
Chemnitz Tom. VI. Tab . 16 Fig. 168-169 = Cardium
unedo Lin. - CHARACTERS AND OCCURRENCE
- DISTRIBUTION. Indo-Pacific (HyUeberg 1994).
rubrum Montagu, 1803. - Unknown species. OTHER
JUDGEMENTS: Tellina r., Turt. C.D. 186; Syn. - Kellia r.
Turt. D. B. 57, pI. 11, f. 7,8; Syn. - Bomia seminulum,
Phil. Sic. i. 14, pI. 1, f. 16; Syn. - Poronia r. Recluz.
Rev. Z. 1843, p. 175.; - Syn. Lasaea r. Brown. »
Regarding Cardium rubrum: RECORDS. --1803
Montagu, p. 83 (pI. 27, fig 4): Cardium rubrum . "1803
Montagu: Cardium rubrum Walker's Minute Shells.
86? "1807 Maton & Rackett, No. 10: Cardium rubrum
Montagu p. 9. "1813 Fleming: Cardium rubrum
Montagu. "1815 Wood: Cardium rubrum Montagu.
746
Tropical Marine Mollusc Programme (TMMP)
rubrum - rudis catillusrudis
"1817 Dillwyn: Cardium rubnlm Montagu . "1819
Turton: Tellina robra Montagu. ··1822 Turton: Kellia
rubra Montagu. ··1825 Gerville: Cardium rubrum
Montagu. ··1828 Wood: Cardium nlbrom Montagu .
··1844 Brown: Lasaea rubra Montagu. ··1845 Reeve,
No. 124: Cardium rubrum Montagu. ··1850 Jay:
Cardium nlbrom Mont. = Kellia rubra Turton. ··1856
Wood & Hanley, No.8: Kellia robrom Mont. T. B. 83
(not Reeve). ··1859 Sowerby: C. robrom (young) =
Cardium fasciatum Montagu. ··1871 Lischke: Cardium
rubrom Montagu, non Reeve Fig. 124 = Lasaea robra.
··1892 Bucquoy, Dautzenberg & Dollfus: Lasaea robra
Montagu according to Fischer-Piette 1977. ··1922-32
Sherborn, p. 5578: Cardium robrom Montagu, 1803.
··1942 Fischer-Piette, Fischer, Germain & Pal\ary:
Cardium rubf1lm Montagu 1803 = Lasaea adansoni
Gmel. = Lasaea robra Montagu. ··1977 Fischer-Piette:
1845 Reeve, No. 124: Cardium robrom Montagu =
Cardium exiguum Gmelin. - CHARACTERS AND
OCCURRENCE - DISTRIBUTION. Kentish coast,
Falmouth, Kingsbridge, Devon (Maton & Rackett
1807); Britain &c. (Wood & Hanley 1856).
rubrum Reeve, 1845. [Figs. Part 1 p. 145]. - Unknown
species. OTHER JUDGEMENTS: Parvicardium ovale
(Sowerby, 1833). - Cardium fascialum Montagu =
Parvicardium ovale. » Regarding Cardium rubnlm:
Cardiidae. Recent. RECORDS. ··1845 Reeve, No. 124:
Cardium robnlm Montagu, Testacea Britannica, p. 83.
··1856 Wood & Hanley: C. rubrom Reeve f. 124 =
Cardiumfasciatum Mont. ··1868 ZR: Cardium robrom
(Reeve) = fasciatum (Mont.); sueeieum (Loven) is
distinct from minimum (Phil.) [Ref. Romer 1868].
··1867 Weinkauff: Cardium rubrom Reeve = Cardium
fasciatum Montagu. ··1869 Romer: Cardium robnlm
Reeve sic fig. 124 = Cardium fasciatum Montagu.
··1880 Stossich: Cardium ntbrum Reeve = Cardium
ovale in Reeve. ··1886 Locard: Cardium rubrum
Reeve, 1845 = Cardium fascialum, Montagu, 1808.
··1888 Kobelt: Cardium nlbrom Reeve = Cardium
fasciatum Montagu. ··1890 Paetel: Cardium robnlm
Rv. = fasciatum Mtg. ··1917 Hidalgo: Cardium rubnlm
Reeve = Cardium fascia tum Montagu. CHARACTERS AND OCCURRENCE - DESCRIPTIONS. ··1845 Reeve: shell globosely cordate, posteriorly slightly angulated; radiately ribbed, ribs convex,
here and there prickly; rose-colour, somewhat transparent. DISTRlBUTION . ··1845 Reeve: Britain.
rubus Meuschen, 1778. Non binominal. - Not available.
RECORDS. ··1902 Sherborn, p. 845: Cardium rubus
Meuschen, 1778.
rucali Uspenskaia, 1931 . - Combination to be revised:
"Cardium" rucali Uspenskaia, 1931. Cardiidae.
Miocene . RE COR DS. ··1932 ZR: Cardium rueali
Uspenskaia, 1931 n. sp. Miocene Daghestan and
Transcaspian.
rudentis Fischer-Piette, 1977. [Figs. Part 3 p. 884]. Mistake: a fossi l. » Regarding Laevicardium rodentis:
Cardiidae. Eocene. TYPE LOCALITY. West Africa. TYPE
REPOSITORY. Paris Museum. RECORDS. ··1977 FischerPiette: Laevicardium (Vepricardium) rodentis n. sp.
··1977. ZR: Laevicardium (Vepricardium) rodentis
Fisher-Piette, 1977 sp. nov. West Africa. Refto FischerPiette, E. p. 41, fig' d. ··1991 Voskuil & Onverwagt:
Laevicardium (Vepricardium) nldentis Fischer-Piette,
1977 . ··2000 Vidal, p. 448: Laevicardium (Vepricardium) rodentis Fischer-Piette, 1977: 41, pI. 3, figs
1-2; the holotype and paratype (in MNHN) are Cardium
porolosum Lamarck, an Eocene fossil.- CHARACTERS AND OCCURRENCE - REMARKS. Referred to
Cardium (Bucardium) in Keen (1980). DISTRIBUTION.
Indo-Pacific (Hylleberg 1994).
Rudicardium Coen, 1915 (genus). -Cardiidae. Miocene
and Recent. RECORDS. ··1937 ZR: Rudicardium Coen
1915 "type here designated Cardium tuberculatum
Linne, 1758" [Ref. Keen 1937]. ··1980 Vokes:
Rudicardium (Monterosato MS) Coen, 1914 in
subfamily Cardiinae.
Rudicardium Coen, 1915 (section, subgenus). Cardiidae. Miocene and Recent. Type species: Keen
(1937) designated Cardium tuberculatum Linne, 1758
type species of subgenus Rudicardium of the genus
Acanthocardia Gray, 1851 . Subsequently raised to
generic level by authors. The designation "section"
takes the value of subgenus. RECORDS. "1914 Coen:
Section Rudicardium Monterosato Mss. CHARACTERS AND OCCURRENCE - DESCRIPTIONS. Medium to large shells. Outline rounded,
subquadrate; posterior margin subtruncate. Sculpture
of prominent ribs with narrow cup shaped spines
(noded) arising from midlines of ribs . Hinge gently
curved, lower cardinal in RV much the larger. Internal
margin crenulate. Mediterranean (Moore et af. 1969,
emend. Hylleberg 1994). Characters compiled from
Schneider (2002): the right anterior cardinal tooth does
not extend ventrally to the hinge line, nor do the right
anterior cardinal teeth connect dorsally to the right
posterior cardinal teeth. The right posterior cardinal
teeth longer than high with a weaker hook. Notches on
the posterodorsal portion of the tuberculate spines.
Acanthoeardia (Rudicardium) has labial palps with
length less than one-third the length of the entire animal
(minus the shell).
rudis Camacho, 1960. - "Cardium" rudis Camacho,
1960. Cardiidae. Paleocene. RECORDS. "1960 ZR:
Cardium nldis Camacho, 1960 sp. nov. Paleocene River
Ewan, Tierra del Fuego .
*rudis. - Homonymy to be noticed. Didacna rudis from
2 sources: NaJivkin, 1916. - Ebersin (?year).
rudis Ebersin. - Didacna rodis Ebersin. Lymnocardiidae.
Quaternary. RECORDS. '·1943 ZR: Didacna rudis Ref.
to 10ukove (1943). ··1963 ZR: Didacna rodis Ebersin
Quaternary.
rudis NaJivkin, 1916. - Didacna rodis NaJivkin, 1916.
Lymnocardiidae. Fossil. "1916 ZR: Didacna rodis
Nalivkin, 1916 sp. n. Berg der Bakustufe.
rudis Reeve, 1862 . - Tridacna rudis Reeve, 1862.
Tridacnidae. RECORDS. ··1980 Ruhoff: Tridacna f1ldis
Reeve, 1862; [1851-1870] vol. 14 (Tridacna) pI. 5, sp.
4. ··1999 Higo et af.: Tridacna nldis Reeve, 1862 =
Tridacna (Flodacna) maxima (Roding, 1798). ·'2003
Hylleberg & Kilburn, p. 186: Tridacna rodis Reeve,
1862. Synonym of Tridacna maxima (Roding, 1798).
rudis (variety). - See tuberculatum rudis var. Coen, 1914.
rudis catillusrudis Nalivkin, 1916 (variety). - Upgraded.
Didaena rodis calillusrodis Nalivkin, 1916. Lymnocardiidae. Fossil. RECORDS. ··1916 ZR: Didacna rudis
var. eatillusrudis Nalivkin, 1916 var. n. Berg der
Bakustufe.
Phuket Marine Biological Center Special Publication vol. 30 (2004)
747
rudis euxinica - rugosum
rudis euxinica Popov, 1983 (subspecies). - Didacna
rudis euxinica Popov, 1983. Lymnocardiidae .
Pleistocene. TYPE LOCALITY. Russian SFSR. RECORDS.
"1983 Popov, p. 170, fig.: Didacna rudis euxinica
subsp. nov.
rudis sub vulgaris Nevesskaya, 1963 (subspecies) . Didacna rudis subvulgaris Nevesskaya, 1963. Lymnocardiidae. Quaternary. RECORDS. ··1963 ZR: Didacna
rudis Ebersin sub vulgaris Nevesskaya, 1963 subsp.
nov. Quaternary, Black Sea basin.
rudis varia Nalivkin, 1916 (variety). - Upgraded.
Didacna rudis varia Nalivkin. Lymnocardiidae. Fossil.
REcORDs. ··1916 ZR: Didacna nldis var. varia Nalivkin,
1916 var. n. Berg der Bakustufe.
Rudisles. - Not in Cardiacea. RECORDS . ··1894 ZR :
Rudis/es with new species [Ref. Douville 1894]. ··1896
ZR: Rudis/es in the Urgonia [Ref. Paquier 1896].
rufovariegalum (variety). - See under edule
rufovariega/um var.
rugala (mode). - See ruga/um Gronovius, 1781.
rugalum Gronovius, 1781 authorship replaced by
Dillwyn, 1817. [Figs. Part I p. 135, 165]. - Identical
with Fulvia aper/a (Bruguiere , 1789). PREVIOUS
JUDGEMENTS: species not sufficiently known (FischerPiette 1977). - Fulvia bullata (Meuschen) [not
available]. » Regarding Cardium ruga/um : Not
available. Cardiidae. Recent. RE CORDS . ··1781
Gronovius, pI. 18, fig. 5: Cardium rugatum (fide
Fischer-Piette 1977). ··1817 Dillwyn: Cardium
rugatum Gronovius. ··1844 Reeve, sp. 63: Cardium
ruga/um. ··1867 Frauenfeld: Papyridea rugatum.
··1868 ZR: Cardium ruga tum (Gronov.) = apertum
(Chemn.) [Ref. Romer 1868]. ··1869 ZR: C. ruga tum
(Gronov.) allied to Cardium pulchrum var. [Ref.
Martens 1869]. ··1874 Lischke: Cardium ruga/um
Meuschen, 1781. ··1884 Grasset: Cardium rugatum.
Gron. ··1889 Martens: Cardium (Papyridea) rugatum
Gronov., Reeve. ··1890 Paetel : C. rugata Gronov. =
Papyridea aperta Ch. [not available]. ··1891 Romer:
Papyridea rugata Gronov. Philippines. ··1903 Hidalgo:
Papyridea rugata Gronovius. 1781 . ··1928 Faustino:
Cardium (Papyridea) ruga tum Gronovius . ··1932
Prashad: Cardium ruga tum Gronovius, 1781 = Cardium
apertum Bruguiere, 1789. ··1952 Kuroda & Habe:
rugatum (Meuschen) = Papyridea ruga tum = Fulvia
bullata (Meuschen). ··1977 Fischer-Piette: Cardium
rugatum Gronovius = species insufficiently known
(Fischer-Piette 1977). ··1977 Wilson & Stevenson:
Cardium rugatum Gronovius, 1871 sic (non binomial)
= Fulvia aperta (Bruguiere, 1789). ··1991 Rao et al.:
Cardium (Fulvia) rugatum in Pres ton (1915) =
Laevicardium (Fulvia) apertum (Bruguiere, 1789).
··1991 Rao et al.: Cardium (Fulvia) ruga tum in
Annandale & Kemp (1916) = Laevicardium (Fulvia)
apertwn (Bruguiere, 1789). ··1994 Vidal : Cardium
rogata sic Gronovius, 1781 = Fulvia (Fulvia) aperta
(Bruguiere , 1789). ··1999 Higo et al.: Cardium
rugatum Reeve, 1844 non Dillwyn, 1817 = Fulvia
aperta (Bruguiere, 1789). ··2003 Hylleberg & Kilburn,
p. 178: Cardium rugatum Dillwyn, 1817. Synonym of
Fulvia aperta (Bruguiere, 1789). - CHARACTERS
AND OCCURRENCE - OF COLLECTIONS. C. rugatum
Gro. Gulf of Thailand, 5 fv. Gravel. DESCRIPTION. ··1844
Reeve: shel.l orbicular, very thin, fragile, posterior side
produced, gaping open; radiately ribbed, about 43-44
thin, rather superficial ribs, nearly obsolete towards the
umbones; sometimes whitish variegated with pink,
sometimes purple or yellowish bay, umbones pinkish
purple. REMARKS. ··1844 Reeve: This very delicate and
elegant species varies singularly in colour, some
examples being white mottled with pink, whilst others
are of a clear orange bay or purple. HABITAT. ··1844
Reeve: found in sandy mud at the depth of about 7
fathoms . DISTRIBUTION. ··1844 Reeve: Eastern coast of
the Island of Luzon, Philippines. ··1884 Grasset:
Luzon. ··1891 Romer: Philippines.
rugatum of Dillwyn, 1817. - Identical with Fulvia aperta
(Bruguiere, 1789). OTHER JUDGEMENTS: Fulvia bullata
(Linnaeus, 1758). - Species insufficiently known
(Fischer-Piette 1977). » Regarding C. apertum:
Cardiidae. Recent. TYPE LOCALITY. Jamaica and East
Indies (Smith 1945). TYPE REPOSITORY . Unknown .
RECORDS. ··1817 Dillwyn, I, p . 125, No. 31: Cardium
rugatum. ··1871 Lischke: Cardium rugatum in Reeve
= Cardium apertum Chemnitz. ··1909 Lynge: Cardium
(Papyridea) nlgatum. ··1922-32 Sherborn, p. 5668:
Cardium rugatum Dillwy n, 1817. ··1945 Smith :
Cardium rugatum Dillwyn 1817 syn. = Fulvia bullata
Linne. ··1977 Fischer-Piette: Cardium ruga tum
Dillwyn = species insufficiently known. ··1979
Richardson et al.: Cardium rugatum in Dillwyn, 1817
= Cardium apertum Bruguiere, 1789. ··1992 Lamprell
& Whitehead: Cardium ruga tum Dillwyn, 1817 =
Cardium apertum Bruguiere, 1789. ··1994 Vidal :
Cardium ruga tum "Gronovius" in Dillwyn, 1817 =
Fulvia (Fulvia) aperta (Bruguiere, 1789). ··1997
Nguyen et al.: ?Vepricardium rugatum (Dillwyn). CHARACTERS AND OCCURRENCE - DESCRIPTIONS. ··1844 Reeve: shell orbicular, very thin, fragile,
posterior side produced, gaping open; radiately ribbed,
ribs about three and forty in number, thin, rather
superficial, nearly obsolete towards the umbo;
sometimes whitish variegated with pink, sometimes
purple or yellowish bay, umbo pinkish purple. REMARKS.
··1844 Reeve: varies singularly in colour, some
examples being white mottled with pink, whilst others
are of a clear orange bay or purple. HABITAT. ··1844
Reeve: Island of Luzon, Philippines. ··ZMUC: in sandy
mud at a depth of about 7 fathoms, Cuming coil.
DISTRIBUTION. ··1844 Reeve: C. apertum Luzon, the
Philippines. ··1994 Hylleberg: C. apertum Gulf of
Thailand, Japan, East Indies, Australia.
rugalum of Reeve 1844. - See under rugatum
Gronovius, 1781.
rugiferullt Roverto, 1898. - Hemidiscors rugiferum
Roverto, 1898. Cardiidae. Fossil. RECORDS. ··1898 ZR:
Hemidiscors rugiferum Roverto, 1898 n. sp. Tongrian
of Liguria. ··1899 Sacco, p. 54: Hemidiscors rugiferus
Rovereto, 1898.
rugosicoslatum Schaffer, 1910. - Combination to be
revised: "Cardium " rugosicostatum Schaffer, 1910.
Cardiidae. Miocene. RECORDS. ··1911 ZR : Cardium
rugosicostatum Schaffer, 1910 sp. n. Austria Miocene.
rugosum Deshayes (variety) . - Synonym of Trachycardium angulatum (Lamarck). » Regarding Cardium
var. rugosum: Cardiidae . Recent. RECORDS. ··1839
Anton: Cardium var. rugosum Desh . = Cardium
angulatum Lamarck.
748
Tropical Marine Mollusc Programme (TMMP)
rugosum - rugosum gortanii
rugosum Lamarck, 1819 (variety). [Figs. Part 3 p. 870].
- Infrasubspecific. RECORDs. "1819 Lamarck: Cardium
rugosum var. testa minore, subaequilaterata.
rugosum Lamarck, 1819. [Figs. Part I p. 59 No.7, p. 76
No. 191,79 No. 11 a,b, p. 89,113,115,137. Part 3 p.
863]. - Synonym of Vasticardium pectiniforme (Born,
1780). OTHER JUDGEMENTS : Cardium angulatum
Lamarck 1819. - Not a synonym of Cardium angulatum
Lamarck 1819. - Cardiumjlavum Linnaeus, 1758. Cardium magnum Chemnitz cited. - Cardium
reeveanum Dunker, 1852. - Cardium rugosum Lamarck
1819. « Regarding Cardium rugosum : Cardiidae.
Recent. RECORDS. "1798 Bruguiere. ID by Lamarck,
pI. 297. fig. 2. Cardium rugosum. "1819 Lamarck, p.
10, No . 23: Cardium rugosum. "1827 Bory de St.
Vincent: Cardium rugosum Lamarck. "1835 Deshaye
Cardium rugosum Lamarck. "1838 Sowerby, 149 t
part, fig. 39: Cardium rugosum Lam. "1839 Anton:
Cardium rugosum Lamarck = Cahdium magnum
Chemnitz. "1840 Sowerby, 178 t part, fig . 41:
Cardium rugosum Lam. "1840 Sowerby, p. 5, No. 58:
Cardium rugosum Lam. "1841 Sganzin: Cardium
rugosum Lamarck. "1843 Hanley: Cardium rugosum
Lamarck. "1844 Potiez & Michaud: Cardium rugosum
Lam. Indian Ocean and American? "1845 Reeve, sp.
68: Cardium rugosum Lamarck. "1867 Mitchell:
Cardium rugosum Lamarck. "1867 Frauenfeld:
Trachycardium rugosum. "1868 ZR: C. rugosum
(Lam .) = Cardium jlavum (L.) [Ref. Romer 1868].
"1869 Frauenfeld: Cardium rugosum Lamarck. "1869
Issei: Cardium rugosum, Reeve, Conch. Icon. pI. 14,
fig. 86 (sic = 68) = Cardium magnum citing Chemnitz.
"1869 Romer: Cardium rugosum Lamarck = Cardium
jlavum L. "1870 Blanford: Cardium nlgosum Lamarck.
"1874 Martens: Cardium nlgosum Lamarck. "1877
Angas: Cardium rugosum Lam. "1879 Martens:
Cardium rugosum Lamarck. "1880 Martens in Mobius:
Cardium rugosum Lamarck. "1880 Martens: Cardium
(Trachycardium) rugosum Lamarck. "1884 Grasset:
Cardium rugosum. Lam. "1889 Martens: Cardium
(Trachycardium) rugosum Lam., citing Chemnitz.
"1890 Paetel: Cardium rugosum Lm. = jlavum L.
"1891 Smith: Cardium rugosum Lamarck. "1894
Sowerby : Cardium rugosum Lamarck. "1895 Martens
& Smith in Thurston: Cardium rugosum Lamarck.
··1896 Shop land : Cardium rugosum Lamarck. "1897
Sowerby: Cardium rugosum Lamarck. "1898 Melvill
& Sykes: Cardium rugosum Lamarck. "190 I Sturany:
Cardium rugosum Lamarck. "190 I Sturany: Cardium
rugosum Lm. (= magnum citing Chemn.). "1902
Shopland: Cardium rugosum Lamarck. "1905 Sturany:
Cardium rugosum Lm. (= magnum citing Chemn.).
"1906 Melvill & Standen: Cardium rugosum Lamk . =
jlavum L. in pt. "1909 Lynge: Cardium rugosum
Lamarck, 1819 = Cardium (Trachycardium) jlavum
(Linne, 1758) illustrated by Spengler (1799). "1917
Odhner: Cardium rugosum Lamarck. "1919 Martin:
Cardium rugosum Lamk. "1922-32 Sherborn, p. 5674:
Cardium rugosum Lamarck, 1819 . ··1923
Dautzenberg : Cardium rugosum Lamarck. "1927
Winckworth: Cardium rugosum Lamarck. "1931
Thiele & Jaeckel: Cardium (Trachycardium) rugosum
Lamarck = Cardium jlavum Linne. "1932 Prashad:
Cardium rugosum Lamarck, 1819 in Reeve (1845) =
b
Cardium reeveanum Dunker, 1852. "1939 Adam &
Le10up: Cardium rugosum Lamarck = Laevicardium
(Trachycardium)jlavum (Linne 1758). "1942 Abrard:
Cardium rugosum Lm. in Nardini 1933 is probably
leucostoma Reeve. "1942 Lamy: Cardium rugosum
Lamarck = Cardium jlavum Linne. "1945 ZR: Cardium
rugosum Lk. [Ref. Lamy 1945]. "1948 Ray: Cardium
rugosum Lamarck, 1819 = Cardium (Trachycardium)
jZavum Linnaeus, 1758. "1952 Braga : Cardium
rugosum Lamarck. "1954 Taki: Cardium rugosum
Lamarck. "1958 MacNae & Kalk: Cardium rugosum
Lamarck. "1960 Kuroda : Trachycardium (Acrosterigma) rugosum (Lamarck = (? jlavum Linne) .
"1960 Paes da Franca: Cardium rugosum Lamarck.
"1962 MacNae & Kalk: Cardium rugosum Lamarck.
"1974 Barnard: Cardium rugosum Lamarck 1819 in
Sowerby 1897 = C. jlava. "1977 Fischer-Piette:
Cardium rugosum Lamarck = Laevicardium jlavum
Linne auctorum. "1977 Wilson & Stevenson: Cardium
rugosum Lamarck, 1819. "1991 Vidal : Acrosterigma
rugosum (Lamarck, 1819); removal from synonymy
with A. angulatum (Lamarck 1819). "1991 Vidal:
Acrosterigma rugosum (Lamarck 1819) diagnostic
characters to distinguish from A. angulatum (Lamarck
1819). "1992 Dharma: Trachycardium rugosum .
"1997 Vidal: Cardium rugosum Lamarck, 1819 =
Vasticardium pectiniforme (Born, 1780). "1997 Vidal:
Cardium rugosum Lamarck sensu Reeve, 1845 sp. 68
(not Cardium rugosum Lamarck, 1819) = Vasticardium
luteomarginatum marerubrum (Voskuil & Onverwagt,
1991). "1998 Poutiers : Tra chycardium rugosum
(Lamarck, 1819). "1999 Higo et al.: Cardium rugosum
Lamarck, 1819 non Munster, 1835 = Acrosterigma
(Regozara)jlava (Linnaeus, 1758). "1999 Hylleberg,
p. 401: Vasticardium rugosum . ID J. Vidal. "1999
Vidal: Cardium rugosum Lamarck , 1819 =
Vasticardium pectiniforme (Born, 1780). CHARACTERS AND OCCURRENCE - DESCRIPTIONS. "1845 Reeve: shell oblong-ovate, rather thick,
radiately ribbed, 28-30 ribs, interstices deeply
excavated extreme posterior ribs prickly scaled, middle
ribs smooth, posteriorly crenulated, interior rugosely
crenated; white, banded with faint red spots. REMARKS.
"1819 Lamarck: Cardium rugosum close to Cardium
jlavum. Lin. ?Schroet. 2. t. 7. f. II. a, b. "1845 Reeve:
the deep interstices between the ribs of this species give
them an exceedingly prominent character. DISTRIBUTION.
"1845 Reeve: Ceylon, Nicobar, Madagascar, &c.
"1884 Grasset: Australia. "1994 Hylleberg IndoPacific.
rugosu", Orbigny, 1850. - Unicardium rugosum
d'Orbigny, 1850. Cardiidae. Fossil. RECORDS. "192232 Sherborn, p. 5682: Unicardium rugosum d'Orbigny,
1850.
rugosum Sowerby, 1840 (variety). [Figs. Part I p. \13 ,
115]. - Vasticardium pectiniforme (Born» > Regarding
Cardium rugosum: Cardiidae. Recent. RECORDS. "1840
Sowerby, corrected list, p. 8, No. 58: Figs. 39 and 41 =
Cardium var. rugosum Sowerby. - CHARACTERS
AND OCCURRENCE - REMARKS. Sowerby (1840) fig.
39, dark spotted; fig. 41, pink spotted.
rugosu", gortanii Nardini, 1937 (variety). - Changed
combination: Vasticardium gortanii (Nardini, 1937).
Cardiidae. Pleistocene. RECORDS. "1937 ZR: Cardium
749
Phuket Marine Biological Center Special Publication vol. 30 (2004)
rumana bassarabica - rusticum
rugosum var. gortanii Nardini, 1937 nov. Dahalac (Coil.
Ragazzi). 001997 Vidal : Cardium rugosum var. gortanii
Nardini, 1937 = Vasticardium gortanii (Nardini, 1937).
rumana bassarabica Macarovici, 1961 (variety). Upgraded . Prosodacna rumana Fischer, 1886 .
Lymnocardiidae. Miocene. RECOR DS. 001961 ZR:
Prosodacna rumana var. bassarabica Macarovici, 1961
nov. Odessa; Miocene, Bessarabia, Roumania.
rumana Motas, 1972. - New combination: Limnodacna
rumana (Motas, 1972). Lymnocardiidae. Pliocene.
RECORDS. 001972 ZR: Limnodacna rumana Motas, 1972
sp. nov. Romania. Pliocene. 001975 ZR: Limnodacna
rumana Motas, 1975 sp. nov. Romania, Prahova region
Pliocene. Ref. to Motas, p . 178. 001976 ZR:
Limnodacna rumana Motas & Papaianopol, 1971.
Redescription Lymnocardiidae. Ref. to Motas, p. 132.
rumana Papaianopol , 1989. - Pachydacna rumana
Papaianopol, 1989. Lymnocardiidae. Pliocene. TYPE
LOCALITY. Basin Dacique, Romania. RECORDS. 001989
Papaianopo[, p. 66, fig.: Pachydacna (Pachydacna)
rumana sp. nov. 00198 [ ZR: Pontalmyra (Pontalmyra)
rumana Papaianopo[, 198 [ sp. nov. Romania, Upper
Miocene. Ref. to Papaianopo[, p. 32.
rumana Pauliuc, [974. - Oxydacna rumana Pauliuc,
[974. Lymnocardiidae. Pliocene. RECORDS. [974 ZR:
Oxydacna rumana Pau[iuc, [974 sp. nov. Romania, La
Vallee de Lopatna. Pliocene. Ref. to Pauliuc, p. 74.
rumana Stefanescu, [896 - Horiodacna rumana
Stefanescu, [896. Lymnocardiidae. Miocene. RECORDS.
001896 ZR: Horiodacna rumana Stefanescu, 1896 n.
sp. Miocene, Roumania. [905 Dollfus, commenting
Andrussow [903: P rumana Sabba.
[96 [ ZR:
Prosodacna rumana Miocene [Ref. Macarovici 1961].
001987 Lutskanov: Horiodacna rumana Stef.
rumana Tournouer, mss. Fischer, 1886. - Prosodacna
rumana Fischer, 1886. Lymnocardiidae. Pliocene. TYPE
LOCALITY. Va[ea Nigovani, Roumania. RECORDS. [886
Fischer: Prosodacna rumana Toumouer, mss. 001886
Fischer: Description p. 223, PI. 9, fig. 6. 001999 Jacques
Vidal (pers. com.) MNHN, Paris: Fossils from the
Paleontological Collection. Protodacna rumana
Toumouer. Rumania, 835813.
rumanum (subspecies) . - See under planum rumanum
Wenz, 1942.
rumanum Fontannes, 1886. - Dacicardium rumanum
(F ontannes 1886). Lymnocardiidae. Fossil. RECORDS.
001975 ZR: Limnocardium rumanum Fontannes, [886
designated type species for new genus Dacicardium
Ref. to Papaianopo[, p. 255. 001978 ZR: Dacicardium
rumanum (Fontannes [886) . Description. Ref. to
Papaianopol et al. p. 304.
rupisa/bae. - Combination to be revised: "Cardium "
(?) rupisalba e Osswald, [930 . Cardiidae . Fossil.
RECORDS. [930 ZR: Cardium (?) rupisalbae Osswald,
[930 sp. n., fossil.
rustica (mode). - See under rusticum Linne, 1758.
rustici Sch[f. - Synonym of "Cardium " incertum Bm.
Cardiidae. Fossil. RECORDS. 001832 Bronn: Bucardites
rustici Sch[f. 176. = Cardium incertum Bm. n. 20 I.
*rusticum . - Homonymy to be noticed. Cardium
rusticum from 3 sources: Linnaeus, 1758. - Brocchi,
1814. - Eichwal.d, 1842.
rusticum Brocchi [814. - Synonym of "Cardium"
incertum 8m . Cardiidae. Fossil. RECORDS. 001832
00
00
00
00
00
Bronn: Cardium rusticum Brch. 500 (non Lmk.) =
Cardium incertum Bm. n. 20 [ .
rusticum Eichwa[d [841. [Figs. Part I p. [22, [42] . Synonym of Cerastoderma glaucum (Bruguiere) .
OTHER JUDGEMENTS: Cardium edule Linnaeus, [758 . Cardium eichwaldi Reeve. - Cardium lamarcki Reeve.
- Cerastoderma umbonatum (Wood, 1850) . »
Regarding Cardium rusticum : Cardiidae. Recent.
DISTRIBUTION. Caspian Sea (Reeve 1845). RECORDS.
00184 [ Eichwald, figs. 26-27: Cardium rus ticum .
001845 Reeve: Cardium rusticum, Eichwald (not of
Linnaeus, nor Lamarck) Fauna Caspio-Caucasia. pI.
38. f. 24-27 = Cardium eichwaldi. 001871 Morch, p.
215 , No . 202 : Cardium rusticum Eichw. (1842) =
Cardium eichwaldi Reeve = Cardium (Cerastoderma)
edule L. [871 ZR: C. rusticum (Eichwald) = Cardium
edule (L.) var. injlatum [Ref. Morch 1871] . 0019 [6
Milachewitch: Cardium rusticum in Eichwald (1829);
Krynicki( [837) ; Siemaschko ([ 847); Middendorff
(1849); Uljanin ([ 872); Weinkauff (1880) = Cardium
edule var. lamarcki Reeve (= Cardium glaucum
Bruguiere) . 001968 Logvinenko & Starobogatov :
Cardium rusticum Eichwa[d, [829 (non Linne, [758)
= Ceras toderma lamarcki (Reeve) . 001974
Starobogatov: Cardium rusticum Eichwald, 1829 (non
Linne, [758) = Cerastoderma umbonatum (Wood,
1850).
rusticum Linnaeus, 1758 / Gme\in, 1791. [Figs. Part I
p. 43,48, 55, 76. 8[,101,126. Part 3 p. 847, 848, 849].
- Synonym of Acanthocardia tuberculata (Linne,
1758). OTHER JUDGEMENTS: Cardium edule Linnaeus,
1758. » Regarding Cardium rusticum : Cardiidae.
Recent. TYPE LOCALITY. The Mediterranean (Speng[er
[799). TYPE REPOSITORY. Museum of the Linnaean
Society, London (Reeve 1844, in Note) . RECORDS.
001705 Rumphius. ID by Linnaeus [758: Cardium
rusticum mus. t. 78. f. K. 001742 Gualtieri . ID by
Linnaeus 1758 and by Gmelin [791j Cardium rusticum
t. 71. f. M. 001758 Linnaeus (10tl edition) No . 78 .
Cardium rusticum . [758 Regenfuss. ID by Gme\in
[791: Cardium rusticum Conch. I. t. [2. f. 77. 78.
001758 Regenfuss, I, pI. [2, figs . 77-78 : Cardium
rusticum (fide Spengler [799). 001767 Linne, p. 1124,
sp. 91: Cardium rusticum . 001778 Born, G 18. Cardium
rusticum Linn. S.N . Sp. 91. 001780 Born: Cardium
rusticum Linn. S. N. Sp. 91. 001782 Chemnitz, VI, p.
20 [-202, pI. 19, fig. 197: Cardium rusticum Linne (ex
museo Spengleriano). 001782 Chemnitz. ID by Gme[in
[ 791: Cardium rusticum t. [9 . f. 197. 001786 Schroter,
No. XIX: Cardium rusticum Linn. XII. pag. 1124. Spec.
91. 001788 Schroter: Cardium rusticum. [Reference to
Chemnitz (1782) fig. 201] . 001789 Bruguiere, p. 222,
No. [5: Cardium rusticum Linn. 00[791 Gmelin, p.
3252, No. 23: Cardium rusticum. 00179[ Poli: Cardium
rusticum Linn. pag. 1124 (p. 51-56) (pI. 16, fig. 5[9). [799 Spengler, p. 34, No. 20: Cardium rusticum
Linnei Sp. 91. p . 1124. 00 1800 Donovan, PI. 32, fig. 6
Cardium rusticum. 001802 Bosc, III, p. 102: Cardium
rusticum . 001815 Wood, p. 225, No . 24 : Cardium
rusticum Linne. 001817 Dillwyn, I, p. 128, No. 38 :
Cardium rusticum Linnaeus. [8 [9 Lamarck, VI, p.
12, No. 30: Cardium rusticum. 00[823 Di[lwyn, Tab.
333 fig. 170 Cardium rusticum. 001825 Wood, p. 26,
No. 25: Cardium rusticum . 001832 Deshayes: Cardium
00
00
00
00
750
Tropical Marine Mollusc Programme (TMMP)
rusticum
rusticum L.= Cardium edule L. ··1832 Deshayes:
Cardium rusticum in Brocc. Fossilis = Cardium edule
L. ··1836 Bosc: Cardium rusticum. ··1839 Anton, p.
II, No. 403: Cardium rusticum Linne. ··1840 Sowerby,
p. 2, No. 21: Cardium rusticum Linn. ··1841 Eichwald:
Cardium rusticum L . ··1844 Philippi: Cardium
nlsticum = Cardium edule L. [fossil). ··1844 Reeve,
sp. 16: Cardium rusticum of Polio ··1844 Reeve :
Cardium tubercula tum Chemnitz, Sowerby, (not of
Linnaeus) = Cardium rusticum. ··1856 Wood & Hanley,
No. 25: Cardium rusticum Lin. Syst. ··\858 Adams &
Adams, p. 455: Cardium (lsocardia) nlsticum, Linn.
··1859 Sowerby: C. rusticum = Cardium edule , Linn.
··1867 Hidalgo: Cardium nlsticum, Linne (in Reeve
1844 fig. 16) = Cardium tuberculatum Linne. ··1867
Weinkauff: Cardium rusticum Linne, Chemnitz =
Cardium tuberculatum Linne. ··1868 ZR: Cardium
tuberculatum (L.), this name to be restored for rusticum
(Linne, Reeve) [Ref. Romer 1868). ··1869 Romer:
Cardium rusticum Linne = Cardium tubercula tum L.
··1870 March: c. rusticum L. junior synonym of
Cardium tubercula tum L. ··1886 Locard: Cardium
rusticum Linne, 1767, Poli , 1791 = Cardium
tuberculatum , Linne, 1767. ··1888 Kobelt: Cardium
rusticum Linne, p. 1124 = Cardium tuberculatum Linne.
··\888 Servain: Cardium rusticum of Linnaeus, Po Ii,
Forbes & Hanley. Not of Chemnitz = Cardium
tuberculatum Linnaeus, 1767. ··1890 Paetel : Cardium
rusticum L. = tuberculatum L. juv. ··1891 Romer:
Acan/hocardia rusticum L. ··1892 B. D. & D. p. 256:
Cardium rusticum in Linne, 1767; Born 1780; Gmelin
1790; Poli 1791; Scacchi 1836; Reeve 1844; Hanley
1856; Sowerby 1859; Chenu 1862 = tuberculatum.
··1902 Sherborn, p. 859: Cardium rusticum Linnaeus,
1758. ··1914 Coen: Cardium rusticum L., non
Chemnitz nec Lamk ., nec auct. = Eucardium
(Rudicardium) tubercula tum L. forma typica. ··1932
Winckworth : Cardium rusticum = Cardium
(Cerastoderma) edule lamarcki. ··1939 Steuer:
Cardium (Rudicardium) rusticum L. ··1967 ZR:
Cardium rusticum, conchology, effect of environmental
salinity on size, palaeogeographical significance [Ref.
Kovalevski i 1967]. ··1969 Nordsieck: Cardium
rusticum Linne, 1758 = Rudicardium tubercula tum
tubercula tum (Linne, 1758). ··1977 Fischer- Piette:
Cardium rusticum Linne = Cardium tuberculatum
Linne. - CHARACTERS AND OCCURRENCE - OF
COLLECTIONS. ··ZMUC : 3 specimens Spengler coil.
Mediterranean. (?appears to be brown glaucum) One
specimen Mittellandmeere. DESCRIPTIO NS. ··1799
Spengler: Testa antiquata sulcis 20. remotis interstiis
rugosis. Linnei Sp. 91. p. 1124. Shell with 20 ribs.
Umbones rather close to the anterior end. Outside dull,
calcareous, colour variable; blue, yellow, brown and
red transverse stripes. ··1844 Reeve: shell somewhat
heart-shaped, gibbous, posterior side slightly angulated,
radiately ribbed, about 20 rounded ribs, interstices
peculiarly crisply wrinkled; fulvous white, ornamented
with rather broad rusty-brown transverse bands.
REMARKS. ··1819 Lamarck: Cardium rusticum; close
to Cardium edule Lin. but different from edule in Poli .
I. tab. 16. f. 5- 7. ··1844 Reeve: varieties of the common
Cardium edule appear to have been confounded by
several authors with Cardium rusticum. The richly
painted specimen here represented answers with
remarkable accuracy to Linnaeus' description of his
Cardium rusticum, which may be easily distinguished
by the curious crisped wrinkled sculpture in the
interstices between the ribs. ··1844 Reeve: Cardium
rusticum of Poli, but not of Chemnitz, Lamarck,
Sowerby and others. Linnaeus' specimen is present in
the collection of the Linnaean Society of London
marked "9 1", in Linnaeus' handwriting, referring to
the 12th edition of the 'Syst. Nat.' "1844 Reeve, in
Note: the original specimen of Cardium rustAcum No.
91, described by Linnaeus in Syst. natur. 12t edition,
is deposited in the Museum of the Linnaean Society of
London. Reeve concluded that Cardium tuberculatum
Chemnitz, Sowerby, (not of Linnaeus) = Cardium
rusticum . However, Cardium tubercula/urn of
Linnaeus, considered identical with Cardium rusticum
by most authors, is an obscure specimen of Cardium
echinatum. DIMENSIONS. ··1799 Spengler: length 39 mm
(1.5 Tom.) width 46 mm (1.75 Tom.) . HABITAT. ··1844
Forbes: Aegaean Sea depth range littoral. DISTRIBUTION.
··1844 Reeve: Adriatic and other seas of southern
Europe. ··1856 Wood & Hanley : Britain & C. ··1891
Romer: Rugen, the Baltic.
rusticum Chemnitz, 1782. Non binominal. - Not
available. OTHER JUDGEM ENTS: Cerastoderma edule
(Linnaeus, 1758). - Cerastoderma glaucum (Bruguiere,
1789). - Cardium lamarcki Reeve, 1845. » Regarding
Cardium rusticum : Cardiidae. Recent. RECORDS. ··1844
Forbes: Cardium rusticum Chemn . ··1856 Wood &
Hanley: C. rusticum Ch. (not Lin .) = Cardium glaucum
Brug. ··1867 Weinkauff: Cardium rusticum Chemnitz
non Linne = Cardium edule Linne. ··1868 ZR: C.
rusticum (Chemn. not L.) variety of Cardium edule (L.)
[Ref. Romer 1868]. "1869 Romer: Cardium rusticum
Chemn. non L. = Cardium edule L. ··1886 Locard:
Cardium rusticum (n. Linne, 1845), Chemnitz, 1782 =
Cardium lamarcki, Reeve, 1845 . "1888 Kobelt :
Cardium rusticum Chemnitz = Cardium edule var.
inaequilatera. ··1888 Servain: Cardium rusticum ,
Chemnitz, non Linnaeus = Cardium lamarcki Reeve,
1845. ··1890 Hidalgo: Var. Cardium nlSticum Chemnitz
and Lamarck, non Linne = Cardium edule Linne.
··1890 Paetel: Cardium rusticum Ch. = C. edule L.
··1902 Bellini: C. rusticum, Chemn., Lam. = Cardium
(Cerastoderma) Lamarcki, Reeve.
rusticum of Donovan - Synonym of Cerastoderma edule
(Linne, 1758). » Regarding Cardium rusticum:
Cardiidae. Recent. RECORDS. ··1817 Dillwyn: Cardium
rusticum Donovan = Cardium edule Linnaeus. ··1844
Brown : Cardium rusticum of Donovan = Cardium
edule.
rusticum of Lamarck. [Figs. Part 1 p. 141] . - Synonym
of Cerastoderma edule (Linne , 1758). OTHER
JUDGEMENTS: Cardium lamarckii . » Regarding
Cardium rusticum: Cardiidae. Recent. DISTRIBUTION.
Brest (Grasset 1884). RECORDS. ··1836 Philippi:
Cardium rusticum Lamk. = Cardium edule. ··1845
Reeve: Cardium rusticum Lamarck (not of Linnaeus)
= Cardium lamarckii. ··1850 Jay: Cardium ruslicum
Lam. = Cardium lamarckii. "1866 Brusina: Cardium
rusticum Lamarck. ··1870 ZR: Cardium rusticum Lam.,
non Linne, in brackish water on the shores of the in
Eastern Holstein [Ref. Friedel 1870]. ··1878
751
Phuket Marine Biological Center Special Publication vol. 30 (2004)
rusticum - sakura;;
Monterosato: Cardium rusticum, Lk. (non Lin.) = C.
Clodiense, Ren. (non Brocc.) = C. Lamarcki, Reeve =
C. glaucum Bruguiere = C. edule, auctt. (non L.). ··1884
Grasset: Cardium rusticum. Lam. Brest (Grasset 1884).
rusticum of Philippi, 1836. - Synonym of Cerastoderma
edt/Ie (Linne, 1758).» Regarding Cardium nlsticum:
Cardiidae. Recent. DISTRIBUTION. Mediterranean (Wood
& Hanley 1856). RECORDS. ··1856 Jeffreys: (Cardium
rusticum, Ph.) = Cardium var. edule, B. M. (Note: Ph.
& B. M. = Philippi and "British Mollusca", Forbes &
Hanley 1849 respectively). ··1880 Stossich: Cardium
rusticum, in Philippi. 1836 = Cardium edule Linne.
··1900 Locard & Caziot: Cardium rusticum (according
to Philippi 1844) in Requien, p. 27 = Cardium edule
Linne, 1767.
rusticum of Poli. - Synonym of Acanthocardia
tuberculata (Linne, 1758). » Regarding Cardium
rusticum: Cardiidae. Recent. RECORDS. ··1829 Costa:
C. rusticum Poli = Cardium tuberculatum L. ··1836
Philippi: Cardium rusticum Poli t. 16. f. 5 etc. =
Cardium tuberculatum L. ··1844 Potiez & Michaud:
Cardium nlsticum Poli = Cardium tuberculatum Linn.
··1902 Sherborn, p. 859: Cerastoderma (L.) rusticum
Poli, 1795. - Cardium rusticum.
rusticum alpha Born, 1778 (variety). - Infrasubspecific.
Unknown. RECORDS. ··1778 Born, G 18. rusticum var.
alpha: alba, facies ferrugineis.
rusticum beta Born, 1778 (variety). - Infrasubspecific.
Unknown. RECORDS. ··1778 Born, G 18. rusticum var.
bela: nivea.
rusticum gamma Eichwald (variety). - Infrasubspecific.
Probably = Cardium rusticum Eichwald =
Cerastodermaglaucum (Bruguiere). Cardiidae. Recent.
RECORDS . ··1876 Grimm: Cardium rusticum var.
gamma Eichwald.
rusticum minor Phil. (variety). - Infrasubspecific. »
Regarding Cardium rusticum minor: probably =
Acanthocardia tuberculatum. Cardiidae. Recent.
RECORDS. ··1891 Danilo & Sandri: Cardium rusticum
Chern. var. minor Phil. tav. 4. f. 14.
rusticum (variety). - See under edule rusticum var.
Lamarck.
ruthenicum. - Cardium ruthenicum. Cardiidae. Middle
Miocene. RECORDS. ··1976 ZR: Cardium ruthenicum.
Middle Miocene. Monastyrz. First record for Poland.
Ref. to Musial.
rutteni Beets, 1983. - Laevicardium nltleni Beets, 1983.
Cardiidae . Upper Miocene. RECORDS. ··1983 ZR:
Laevicardium (Discors) rutteni Beets, 1983 sp. nov.
Indonesia. Ref. to Beets, p. 57.
ruvarzi Stevanovic, 1951. - Plagiodaclla ruvarzi
Stevanovic, 1951. Lymnocardiidae. Pontian. RECORDS.
··1953 ZR: Plagiodacna ruvarzi Stevanovic, 1951
"(Pav!. in coIl.) sp. nov. Pontian Serbia.
rybnicense (subspecies). - See under plica tum
rybnicense Grishkevitch & Nevesskaya, 1959.
rzehaki Ctyroky, 1972. - Limnopagetia rzehaki Ctyroky,
1972. Cardiacea . Miocene. RECORDS . ··1972 ZR:
Limnopagetia rzehaki Ctyroky, 1972. Czechoslovakia.
Miocene.
Rzehakia (genus) Korobkov, 1954. - Cardiacea.
Miocene. RECORDS. ··1972 ZR: Rzehakia Korobkov,
1954 [Ref. Ctyroky 1972].
S
sabbae Papaianopol, 1981. - Pontalmyra sabbae
Papaianopol, 1981. Lymnocardiidae. Upper Miocene.
RECORDS. ··1981 ZR: Pontalmyra (Pontalmyra) sabbae
Papaianopol, 1981 sp. nov. Romania. Ref. to
Papaianopol, p. 26.
sabulosum Gabb, 1869. - Changed combination:
Granocardium sabulosllm (Gabb, 1869). Cardiidae.
Cretaceous. RECORDS. ··1978 Scott: Cardium
(Granocardium)sabulosum Gabb, 1869. ··1978 Scott:
Granocardium (Granocardium) sabulosum (Gabb,
1869).
saccelluni Papaianopol, 1982. - New combination:
Lymnocardium saccellum (Papaianopo1, 1982).
Lymnocardiidae. Upper Miocene. TYPE LOCALITY.
Pontien et Dacien. Basin Dacique, Romania. RECORDS.
··1982 Papaianopo1, p. 185, fig.: Limnocardium
(Euxinicardium) saccel/um sp. nov.
sachalinense Khramova , 1962 . - Clinocardium
sachalinense Khramova, 1962. Cardiidae . Fossil.
RECORDS. ··1977 Popov: Clinocardium sachalinense
Khramova, 1962.
sacrum Papaianopol, 1982. - New combination:
Lymnocardium sacrum (Papaianopol, 1982).
Lymnocardiidae. Upper Miocene. TYPE LOCALITY.
Pontien et Dacien. Basin Dacique, Romania. RECORDS.
··1982 Papaianopol, p. 196, fig .: Limnocardium
(Euxinicardium) sacrum sp. nov.
sagaseri Adegoke, 1969. - Trachycardium sagaseri
Adegoke, 1969. Cardiidae. Pliocene. RECORDS. ··1969
ZR: Trachycardium sagaseri Adegoke, 1969 sp. nov.
Early Pliocene, Coalinga region, California, U.S.A.
sagawai Yokoyama, 1927. - Combination to be revised:
"Cardium" sagawai Yokoyama, 1927 . Cardiidae.
Tertiary. RECORDS. ··1927 ZR: Cardium sagawai
Yokoyama, 1927 sp. n. Tertiary Japan.
sagittarium Papaianopol, 1982. - New combination:
Lymnocardium sagitlarium (Papaianopol, 1982).
Lymnocardiidae. Pliocene. TYPE LOCALITY . Pontien et
Dacien. Basin Dacique, Romania. RECORDS. ··1982
Papaianopol , p. 209, fig.: Limnocardium
(Euxinicardium) sagittarium sp. nov.
sakaraulense. - nomen nudum. RECORDS. ··197. ZR:
Cardium (Papillicardium) sakaraufense nom. nud. p.
505, Miocene, Georgia, USSR, Bagdasaryan .
sakhalinensis Slodkewitsch, 1938. - Changed
combination: Profulvia sakhalinensis (Slodkewitsch,
1938). Cardiidae. Miocene & Pliocene. TYPE LOCALITY.
Sakhalin Island. RECORDS. ··1945 ZR: Papyridea
sakhalinensis 'Kogan' sic Upper Miocene. Sakhalin.
··1973 Keen: Fulvia sakhalin ens is (Slodkewitsch,
1938) (Papyridea). ··1976 ZR: Profulvia sakhalinensis
(Slodkewitsch, 1938) Comb. nov. Transferred from
Papyridea. Ref. to Kafanov, p. 482 [orig. p.lll].
sakura;; Habe, 1961. - Nemocardium sakuraii Habe,
1961. Cardiidae . Recent. RECORDS. ··1961 Habe:
Nemocardium sakuraii Habe, 1961. ··1962 Habe:
Nernocardium sakuraii Habe. ··1964 Habe:
Nemocardium (Microcardium )sakuraii Habe. ··1977
Fischer-Piette: Nemocardiurn sakuraii Habe. ··1977
Habe: Microcardium sakuraii Habe, 1961. ··1981
Koyama et af.: Microcardium sakuraii. ··1991
Matsukuma, Okutani & Habe: Nemocardium sakuraii
752
Tropical Marine Mollusc Programme (TMMP)
salinaensis - samodurovi
Habe, 1961 . - CHARACTERS AND OCCURRENCE
- REMARKS. Resembles C. modestum Adams & Reeve
(non Philippi) but is more rounded posteriorly (FischerPiette 1977). DISTRIBUTION. Japan (Hylleberg 1994).
salinaensis Meek, 1871. - Protocardia salinaensis
Meek, 1872. Cardiidae. Cretaceous. RECORDS. --1876
Meek: Protocardia salinaensis. --1876 Meek :
Protocardia salinnaensis sic Meek, 1871 . --1986 Scott:
Protocardia salinaensis Meek, 1872.
salinnaensis (error). - Printer's mistake for salinaensis
in Meek 1876.
sallomacense Cossmann & Peyrot, 1911. - Synonym of
Acanthocardia turonica (Homes, 1861).» Regarding
Cardiumsallomacense: Cardiidae. Miocene. RECORDS.
--1911 ZR: Cardium sallomacense Cossmann &
Peyrot, 1911 sp. n. France Neogene. --1977 Popov:
Acanthocardia sallomacensis (Cossmann & Peyrot,
1911). --1987 ZR: Cardium sallomacense Cossmann
& Peyrot 1912. New synonym of Acanthocardia
(Acanthocardia) turonica (Homes 1861). Ref. to
Studencka p. 69. "1986 Studencka, p. 69 : Cardium
sallomacense Cossmann & Peyrot, 1912; syn. nov. ; of
Acanthocardia (Acanthocardia) turonica (Homes,
1861). --1999 Jacques Vidal (pers. com.) MNHN, Paris:
Fossils from the Paleontological Collection as Cardium
sallomacense Cossmann & Peyrot. Burdig . France
(Landes), B28704.
sallomacensis (mode). - See under sallomacense
Co ssm ann & Peyrot.
salrivalis Harris, 1919. - Proto cardia salrivalis Harris,
1919. Cardiidae. Eocene. RECORDS ... 1919 ZR:
Protocardia salrivalis Harris, 1919 sp. n. Eocene
U.S.A.
saltieri d' Archiac. - Combination to be revised :
"Cardium" saltieri d' Archiac . Cardiidae. Lower
Eocene. RECORDS. --1999 Jacques Vidal (pers. com.)
MNHN, Paris: Fossils from the Paleontological
Collection. Cardium saltieri d' Archiac. B22948 .
salvationemense Lautenschlager, 1962. - Synonym of
Ciliatocardium asagaiense (Makiyama 1934) . »
Regarding salvationemense: Cardiidae. Fossil.
RECORDS. --1974 ZR: Ciliatocardium salvationemense
Lautenschlager, 1962. Comb. nov. Ref. to Kafanov, p.
1469. "1980 ZR: Ciliatocardium salvationemense
(Lautenschlager 1962) is a junior synonym of C.
asagaiense (Makiyama 1934). Ref. to Kafanov.
samanicllm Olsson, 1928. - Combination to be revised :
"Cardium" samanicum (Olsson, 1928). Cardiidae.
Tertiary. RECORDS . --1928 ZR : Cardium
(Trigoniocardia) samanicum Olsson, 1928 sp. n.
Tertiary N. Peru.
samarangae Makiyama, 1934 . - Nemocardium
samarangae (Makiyama, 1934). Cardiidae. Recent.
Type species: Habe (1951) designated Nemocardium
samarangae Makiyama, 1934 type species of the
subgenus (Keenaea) Habe, 1951. Recognised at the
generic level by authors . Keen (1954) Cardium
samarangae Makiyama, 1934 is type species of
Keenaea Habe, 1951. TYPE LOCALITY. Eastern Seas.
RECORDS. "1850Adams & Reeve, p. 77, pI. 22, fig. 6:
Cardium modestum. "1872 Carpenter: Cardium
modestum Adams & Reeve. --1882 Dunker: Cardium
modestum Adams & Reeve. "1895 Pilsbry: Cardium
modestum Adams & Reeve. --1934 Makiyama:
Cardium samarangae; new name for Cardium
modestum Adams & Reeve, 1850. --1940 Nomura &
Niino: Cardium samarangae). "1940 Nomura:
Nemocardium samaranga e . --1950 Keen:
Nemocardium samarangae Makiyama, 1934. --1951
Habe: Nemocardium (Keenaea) samarangae. "1952
Kuroda & Habe: Nemocardium samarangae. --1952
ZR : Cardium samarangae Makiyama subgenotype of
Keenaea Habe, 1951 subgen. nov. of Nemocardium .
--1954 Keen: Cardium samarangae Makiyama, 1934.
--1952 Kuroda & Habe : samarangae Makiyama =
Nemocardium adamis sic. --1954 Keen: Nemocardium
(Keenaea) samarangae (Makiyama), 1934. "1955
Kira : Nemo cardium samarangae . --1958 Habe:
Keenaea samarangae Makiyama . --1962 Kira:
Nemocardium (Keenaea) samarangae . --1964
Shikama: Nemocardium samarangae. --1968 ZR:
Nemocardium samarangae first record from off
Rikuchu coast, Iwate prefecture, Japan [Ref. Habe &
Kosuge 1968]. "1969 Moore el al.: Nemocardium
(Keenaea) samarangae Makiyama, 1934. --1971
Kuroda , Habe & Oyama : Nemocardium (Keenaea)
samarangae. --1977 Fischer-Piette: Nemocardium
samarangae Makiyama . --1977 Popov : Pratulum
samarangae (Makiyama). "1981 Koyama et at. :
Keenaea samarangae . --1990 Gladenkov &
Sinelnikova: Nemocardium samarangae (Makiyama,
1934). --1991 Matsukuma, Okutani & Habe: Keenaea
samarangae (Makiyama, 1934). - CHARACTERS
AND OCCURRENCE - DIMENSIONS. 28 x 26 x 18 mm
(Kuroda et al., 1971). HABITAT. Fine sand, mud at 50300 m depth (Kuroda et al. 1971). DISTRIBUTION. East
China Sea (Hylleberg 1994); Japan, Honshu to Kyushu
(Kuroda et al. 1971).
samarangae ob/iqua Ozaki, 1957 (subspecies). Nemocardium samarangae obliqua Ozaki, 1957.
Cardiidae. Miocene. RECORDS. --1957 ZR:
Nemocardium (Keenaea) samarangae obliqua Ozaki,
1957 subsp. nov. Miocene Koti Prefecture, Japan.
samarangae taiwanensis Masuda & Huang, 1990
(subspecies). - Nemocardium samarangae taiwanensis
Masuda & Huang, 1990. Cardiidae. Miocene. TYPE
LOCALITY. Western Foothills of northern Taiwan.
RECORDS. "1990 Masuda & Huang, p. 153, fig.:
Nemocardium (Keenaea) samarangae taiwanensis
subsp. nov.
sambaicum Maury 1917. - Combination to be revised:
"Cardium" sambaicum (Maury 1917). Cardiidae.
Neogene. TYPE MATERIAL. Lectotype designated Vokes
1989 , p . 115. RECORDS . "1919 ZR: Cardium
(Trigoniocardia) sambaicum Maury, 1919 sp. n .
Tertiary S. Domingo. --1989 Vokes, p. liS : Cardium
(Trigoniocardia) sambaicum Maury, 1917.
sambaicum portoricoensis Hubbard, 1920 (variety). Upgraded . Combination to be revised: "Cardium "
sambaicum portoricoensis Hubbard, 1920. Cardiidae.
Tertiary. RECORDS. --1920 ZR : Cardium
(Trigoniocardia) sambaicum portoricoensis Hubbard,
1920 var. n. Tertiary Porto Rico.
samodurovi Merklin, 1960. - Cerastoderma samodurovi
Merklin, 1960. Cardiidae. Fossil. RECORDS. --1975 ZR:
Cerastoderma (Cerastoderma) samodurovi Merklin,
1975 sp. nov. USSR, north Aral region, Koktumak
Peninsula. Middle Oligocene. Ref. to Merklin, p. 96.
Phuket Marine Biological Center Special Publication vol. 30 (2004)
753
sancatsense - scabrum
··1977 Popov: Cerastoderma samodurovi Merklin,
1960.
sancatsense. - Changed combination: Acanthocardia
sancatsense. Cardiidae . Upper Miocene. RECORDS .
··1973 ZR: Cardium (A canlhocardia) sancatsense.
First record. Czechoslovakia. Ref. to Ondrejickova.
sanctdavidis Maury, 1925. - Trachycardium sanctdavidis
(Maury, 1925). Cardiidae. Fossil. RECORDS. ··1977
Popov: Trachycardium (Dallocardia) sancldavidis
(Maury, 1925).
sanctisabae c.F. Roemer, 1849. - Combination to be
revised: "Cardium " sanctisabae c.F. Roemer, 1849.
Cardiidae. Fossil. RECORDS. ··1922-32 Sherborn, p.
5732: Cardium sanclisabae C.F. Roemer, 1849 [as
sancti Sabael .
sanctisebastiani Maury, 1920. - Combination to be
revised: "Cardium " Trigoniocardia sanctisebasliani
(Maury, 1920). Cardiidae. Tertiary. RECORDS. ··1920
ZR: Cardium (Trigoniocardia) sanctisebastiani Maury,
1920 sp. n. Tertiary Porto Rico.
sandbergri Gumbel , 1861 . - Combination to be revised :
"Cardium" sandbergri Gumbel , 1861. Cardiidae.
Fossil. RECORDS. ··1980 Ruhoff: Cardium sandbergri
Gumbel, 1861 ; p. 743 .
sandigii Caster, 1938. - Combination to be revised :
"Cardium " sandigii Caster, 1938. Cardiidae. Eocene.
RECORDs. ··1938 ZR: Cardium sandigii sp. n. Eocene,
Luculo, R., Angola [without auct. 1938]. ··1939 ZR:
Cardium sandigii Caster, 1938 sp. n. Eocene at month
of Lucolo River. Angola.
sanguineotincta Preston (invalid). - Nomen nudum. One
lot (identified as Cardium oxygonum Sowerby, 1833
by Vidal 1999) at the BMNH (reg. 1910-12-13-15),
from Sri Lanka, is labeled as a type series of "Cardium
sanguineolincta " Preston; another lot AMS C034080,
from Sri Lanka, is labeled as a cotype lot of the same
taxon (Vidal 1999). Since there is no description or
reference to Cardium sanguineolincla in the literature,
it is a nomen nudum . J. Hylleberg located one specimen
of "Cardium sanguineotincla" at the Natural History
Museum in Vienna (labeled co-type) and one specimen
labeled type in the Dautzenberg collection, Natural
History Museum, Brussels. RECORDS. ··1999 Vidal, p.
289: Vastic ardium sanguineotinctum . See under
oxygonum .
*santacruzense. - Homonymy to be noticed. Cardium
sanlacruzense from 2 sources: Ihering, 1914. Feruglio, 1936.
santacruzense Feruglio, 1936 . Combination to be
revised : "Cardium " sanlacruzense (Feruglio, 1936).
Cardiidae. Cretaceous. RECORDS. ··1937 ZR: Cardium
(Hemicardium ?) sanlacruzense FerugJio, 1936 sp. n.
Cretaceous Argentina . ··1938 ZR : Cardium
(Hemicardium ?) sanlacruzense n. f. Upper Cretaceous,
Calafate [Ref. Ferruglio 1936].
santacruzense Ihering, 1914. - Combination to be
revised: "Cardium" sanlacruzense Ihering, 1914.
Cardiidae. Patagoniano RECORDS. ··1935 ZR: Cardium
santacruzense Ihering, 1914 sp. n. Upper Patagonian
Santa Cruz.
saportae Coquand, 1862. - Combination to be revised:
"Cardium " saporlae Coquand, 1862. Cardiidae. Fossil.
RECORDS. ··1871 Stoliczka, 113: C. saporlae Coquand,
1862. ··1980 Ruhoff: Cardium saportae Coquand,
1862; vol. 2: 207, pI. 11 , figs . 8-9.
sarbiense (variety). - See under schedelianum var.
sarbiense Gillet, 1943.
Sarmaticardium Bagdas sarian, 1978 (genus). Cardiidae. Fossil. RECORDS. ··1979 ZR: Sarmaticardium Bagdasaryan, 1979. Gen. nov. Type species
S. vigeni, Upper Miocene. Ref. to Bagdasaryan, p. 40.
··1980 Vokes: Sarmaticardium Bagdassarian, 1978.
Fossil.
Sarmoticardium (subgenus). - RECORDS. ··1987 p. 9.
Kojumdgieva: (Sarmaticardium) Bagdasarjan, 1978.
Cardiidae. Miocene, Sarmatian. Emended diagnosis.
sartasense Merklin & Nevesskaya, 1956. - Combination
to be revised : "Cardium " sartasense Merklin &
Nevesskaya, 1956. Cardiidae. Miocene. RECORDS.
··1956 ZR: Cardium sartasense Merklin &
Nevesskaya, 1956 sp. nov. Miocene. Turkmenstan and
w. Kazakhstan. ··1959 ZR : Cardium sartasense
Merklin & Nevesskaya, 1959 sp. nov. Turkmenistan,
Miocene.
sauciatum Beck. - Nomen nudum . REMARKS. After
publication of the name C. australe, Sowerby (1840)
met the name C. sauciatum in Mr. Cuming's collection
which had been studied by Dr. Beck, but the name was
never published. RECORDS. ··1840 Sowerby, Proc. Z.S.
London p. 105: Cardium saucialum (nomen nudum).
··1840 Sowerby, catalogue p. 1, re sp. 10: C. saucialum
Beck = Cardium auslrafe Sowerby 1840.
sauerzopji (subspecies). - See under kuiperi sauerzopji.
sautcotsense Mayer, 1866. - Combination to be revised :
"Cardium" saucalense Mayer, 1866. Cardiacea. Fossil.
RECORDS. ··1902 Dollfus & Dautzenberg p. 314 :
Cardium saulcatsense Mayer. ··1970 Glibert & Poe1,
p. 56: Cardium (A canthocardia) saucatsense Mayer,
1866. ··1980 Ruhoff: Cardium sal/lcatsense Mayer,
1866; [1857-1869] vol. 14: 75, pI. 2, fig-2.
Sawkinsia (genus). - Tridacnacea. Fossil. CHARACTERS.
Sawkinsia has symmetrical or nearl y symmetrical
knob-like spines (Schneider 2002). ··1976 ZR :
Sawkinsia Cox 1941. Redescription. p. 744 . Ref. to
Jung. ··1992 Schneider p. 149: Sawkinsia ; Cox 1941.
Referred to Cardiinae (Cardiacea) from Tridacninae.
··1998 Schneider: Sawkinsia.
sayre; Elias, 1957. - Conocardium sayrei Elias, 1957.
Conocardiidae. Carboniferous. RECORDS. ··1957 ZR:
Conocardium sayrei Elias, 1957 sp. nov. Mississippian,
Carboniferous, Southern Oklahoma, U.S .A.
sayri Richards, 1943 . - Combination to be revised :
"Cardium " sayri Richards, 1943. Cardiidae. Fossil.
RECORDS. ··1943 ZR: Cardium (Granocardium) sayri
Richards, 1943 sp. nov. New Jersey. Raritan .
scabriuscuillm Fuchs . - New combination: Lymnocardium scabriusculum (Fuchs). Lymnocardiidae.
Fossil. RECORDS. ··1987 Taktakishvili: Limnocardium
scabriuscufum (Fuchs).
*scabrum . - Homonymy to be noticed. Cardium
scabrum from 2 sources: Philippi, 1844. - Reuss, 1844.
scabrum ofUljanin (1872). - Misidentification. Changed
combination : Parvicardium simile (Milaschevich).
Cardiidae. Recent. RECORDS. ··1916 Milachewitch:
Cardium scabrum in Uljanin (\ 872) = Cardium simile
Milachewitch . ··1962 Grossu: Cardium scabrum
Philippi in Uljanin (\ 872) = Cardium (Parvicardium)
simile Milaschevich sic.
754
Tropical Marine Mollusc Programme (TMMP)
scabrum - schemachinica
scabrum Philippi, 1844. - Parvicardium scabrum
(Philippi, 1844). Cardiidae. Recent. RECORDS. "1844
Philippi , p. 38, pI. 14, fig. 16: Cardium scabrum n. sp.
"1859 Sowerby: C. scabrum Ph. = Cardium punctalum
Broc. "1867 Weinkauff: Cardium scabrum Philippi =
variety of Cardium roseum Lamarck. "1868 ZR:
Cardium nodosum (Mont.) is perhaps not = roseum
(Lam.), or punclalum (Brocchi), or scabrum (PhiL)
[Ref. Romer 1868]. "1869 Romer: Cardium scabrum
Philippi, an varietas? (not close to nodosum). "1880
Stossich: Cardium scabrum, Philippi. 1844 = Cardium
nodosum Montagu. "1886 Locard: Cardium scabrum
Philippi, 1844. = Cardium roseum, de Lamarck, 1818.
"1888 Kobelt: Cardium scabrum Philippi = Cardium
nodosum Turton. "1890 Paetel: Cardium scabrum PhI.
= nodosum MIg. "1900 Locard & Caziot: Cardium
scabrum (Phil) in Requien = Cardium nodosum Turton,
1822. "1917 Hidalgo: Cardium scabrum Philippi =
Cardium nodosum Turton. "1922-32 Sherborn, p.
5765 : Cardium scabrum Philippi, 1844 . "1932
Winckworth: Cardium (Parvicardium) scabrum
Philippi, 1844. "1938 ZR : Cardium scabnlm larvae,
figs . [Ref. Lebour 1938] . "1962 Allen: Cardium
(Parvicardium) scabrum Philippi 1844 = Cardium
nodosum in Jeffreys 1863. "1969 Nordsieck:
Parvicardium scabrum Philippi, 1836. "1971 ZR:
Cardium scabrum New record. West of British Isles
Glamorgan-Pembrokeshire coastal region [Ref. Taylor
1971]. "1973 ZR: Cardium scabrum Philippi. New
synonym of C. ovale Sowerby, p. 282. Ref. to
Rasmussen. "1974 Clarke: Parvicardium scabrum
(Philippi). "1977 Fischer-Piette: Cardium scabrum
Philippi = Cardium nodosum Montagu. "1978 Janssen
& Van der Slik: Parvicardium scabrum (Philippi, 1844).
"1991 van Aartsen & Giannuzzi-Savelli: Parvicardium
scabrum (Philippi, 1844) valid name. "1998 Macedo
et al. : Parvicardium scabrum (Philippi, 1844). CHARACTERS AND OCCURRENCE - HABITAT. The
species nodosum at 9-420 m depth, in the North Sea
from 42 m depth and down (Nordsieck 1969).
DISTRIBUTION . C. nodosum North Norway, Ireland,
Canary Islands, Mediterranean (Nordsieck 1969).
scab rum Reuss, 1844. - Combination to be revised:
"Cardium" scabrum Reuss, 1844. Cardiidae. Fossil.
RECOR DS. "1922-32 Sherborn, p. 5765: Cardium
scabrum Reuss, 1844.
scacchii Deshayes, 1853. - Combination to be revised:
"Cardium" scacchii Deshayes, 1853. Cardiacea. Fossil.
RECORDS. "1980 Ruhoff: Cardium scacchii Deshayes,
1853; vol. 2: 70.
scalata Vidal, 1994. [Figs. Part 3 p. 890] . - Fulvia scalata
Vidal, 1994. Cardiidae. Recent. TYPE LOCALITY. New
Caledonia. RECORDS. "1994 Vidal, p. 108, pI. 1, figs.
3,4; pI. 3, figs 5 a,b, 7: Fulvia (Fulvia) scalala sp. nov.
"1998 Lamprell & Healy : Fulvia (Fulvia) scalala
Vidal, 1994. "2001 Vidal , p. 407: Fulvia scalala.
'·2003 Hylleberg & Kilburn, p. 183: Fulvia scalala
(Vidal, 1994). - CHARACTERS AND OCCURRENCE - DESCRIPTIONS. ··200 I Vidal: tentacles
numerous. Eyes on tip of tentacles. DISTRIBUTION.
"2003 Hylleberg & Kilburn: 3 valves from "Sainson"?
are deposited in the Collection Saurin at MNHN, Paris
(J . Vidal).
scapoosense Clark, 1925 . - Changed combination:
Clinocardium scapoosense (Clark, 1925). Cardiidae.
Oligocene. RECORDS. "1927 ZR: Cardium (Cerasloderma) scapoosense Clark, 1927 sp. n. Oligocene W.
N. America. "1977 Popov : Clinocardium scapoosense
(Clark, 1925).
scarsellai Tavani, 1948. - Combination to be revised :
"Cardium" Tavani , 1948 . Cardiidae. Cretaceous.
REcORDs ... 1948 ZR: Cardium (Prolocardia) scarsellai
Tavani, 1948 sp. nov. Cretaceous Somaliland.
schafferi harrandei Kautsky, 1925 (variety). - Upgraded.
Acanlhocardia schafferi barrandei Kautsky, 1925.
Cardiidae. Miocene. RECORDS. "1999 Jacques Vidal
(pers. com.) MNHN, Paris : Fossils from the
Paleontological Collection. Acanthocardia schafferi
var. barrandei Kautsky, 1925. Bulgaria (Dobroucha),
B44409.
schakonensis Wassoievic & Eberzin, 1930. Macrodacna schakonensis Wassoievic & Eberzin,
1930. Lymnocardiidae. Fossil. RECORDS. ··1931 ZR:
Monodacna schakonensis Wassoievic & Eberzin, 1930
sp. n., m. URSS Pliocene. "1967 Ebersin: Macradacna
schakonensis Ebers. et Wass ... 1967 ZR: Macradacna
schakonensis (Ebers. & Wass., 1930) pro Monodacna
schakonensis Ebers. & Wass., 1930 [Ref. Eberzin
1967]. "1977 Popov: Macrodacna schakonensis.
schatskii Ruzencev, 1929. - Monodacna schalskii
Ruzencev, 1929. Lymnocardiidae. Apscheronian.
RECORDS. "1929 ZR: Monodacna schalskii Ruzencev,
1929 sp. n. Apscheronian, Transcaucasia.
schedeliallum Partsch. - New combination.
Lymnocardium schedelianum (Partsch). Lymnocardiidae . Miocene. RECORDS. "1946 ZR :
Limnocardium schedelianum Partch Miocene [Ref.
Gillet 1943]. "1998 Schultz: Limno cardium
schedelianum (Partsch) .
schedeliallum sarbiense Gillet, 1943 (variety). Upgraded. New combination. Ly mnocardium
schedelianum sarbiense Gillet, 1943 . Lymnocardiidae.
Miocene . RECORDS. "1946 ZR: Limnocardium
schedelianum Partch var. sarbiense Gillet, 1943 nov.
Miocene, Rumania.
Schedocardia Stewart, 1930 (genus). - Cardiidae. Fossil.
RECORDS. "1978 Freneix & Mackie: Genus
Schedocardia Stewart , 1930. "1980 Vokes:
Schedocardia Stewart, 1930 in subfamily Cardiinae.
··2002 Schneider: Schedocardia : Cardiinae, not
Fraginae. - CHARACTERS AND OCCURRENCE DESCRIPTION ... 2002 Schneider: Schedocardia has
symmetrical or nearly symmetrical knob-like spines.
Schedocardia Stewart, 1930 (subgenus). - Cardiidae.
Paleocene-Eocene. GENOTYPE. Stewart (1930)
designated Cardium halchetigbeense Alrich sic, 1886
type species of subgenus Schedocardia of genus
Acanlhocardia. RECORDS. ··1930 ZR: (Schedocardia)
Stewart, 1930 subgen. n. of Plagiocardium type
Cardium halchetigbeense A1rich sic. - CHARACTERS
AND OCCURRENCE - DESCRJPTlONS. Schedocardia
is less oblique than A. (Acanthocardia); spines on ribs
weak to obsolete, especially anteriorly. SE . AsiaAfrica-Europe-North and South America (Moore el
al. 1969).
schemachinica Andrusov, 1963. - Didacna
schemachinica Andrusov, 1963. Lymnocardiidae.
Miocene. REcORDs. "1963 ZR: Didacna schemachinica
Phuket Marine Biological Center Special Publication vol. 30 (2004)
755
schencki - scobinula
Andrusov, 1963 sp. nov. Miocene, Russia.
schencki Wiedey, 1928. - Changed combination:
Trachycardium schencki (Wiedey). Cardiidae.
Miocene. RECORDS. ··1928 ZR: Cardium schencki,
Wiedey, 1928 sp. n. Miocene California. ··1944 Keen
& Bentson: Cardium schencki Wiedey, 1928 (n. sp.).
··1944 Keen & Bentson: Trachycardium schencki
(Wiedey). ··1954 Keen: Trachycardium schencki
(Wiedey), 1928 (in unpublished thesis).
schirvanica (variety). - See under ampelakiensis var.
schirvanica Andrusov, 1963.
schirvanica A1i-Zade & A1eskerov, 1969. - Hyrcania
schirvanica A1i-Zade & A1eskerov, 1969 .
Lymnocardiidae. Apsheron. RECORDS. ··1969 ZR :
Hyrcania schirvanica Ali-Zade & Aleskerov, 1969 sp .
nov. Upper Apsheron, Mali Kharam, Azerbaidjan.
schirvanica Andrusov, 1909. - Prosodacna schirvanica
Andrusov, 1963. Lymnocardiidae. Miocene. RECORDs.
··1963 ZR: Prosodacna schirvanica Andrusov, 1963
sp. nov. Miocene, Russia. ··1981 ZR: Prosodacna
schrvanica sic Andrusov, 1909 Designated type species
for new genus Schirvanicardium. Ref. to Andreescu,
p.129.
schirvanica major Andrusov & Ebersin, 1959
(subspecies). - Replaced by Prosodacna semisulcata
babadjanica Andreescu, 1980. » Regarding P
schirvanica major: Lymnocardiidae. Fossil. RECORDS.
··1980 ZR: Prosodacna schirvanica major Andrusov
& Ebersin, 1959. Replaced by Prosodacna semisulcata
babadjanica Andreescu, 1980. Ref. to Andreescu.
Schirvanicardium Andreescu, 1974 (genus). Lymnocardiidae. Tertiary. RECORDS . ··1980. Vokes:
Schirvanicardium Andreescu, 1974. Subfamily
uncertain. ··1981 ZR: Schirvanicardium Andreescu,
1981 gen. nov. of Lymnocardiidae, type species
Prosodacna schrvanica sic Andrusov 1909, fossil. Ref.
to Andreescu, p. 129.
schmidti Homes, 1862. - New combination .
Lymnocardium schmidti (Homes,
1862).
Lymnocardiidae. Pannonian. RECORDS. ··1966 ZR:
Limnocardium schmidti Pannonian, Mecsek Mts.
Yugoslavia, pis. [Ref. Bartha 1966). ··1980 Ruhoff:
Cardium schmidti Homes, 1862; [1851-1870] vol. 4:
193, pI. 28. Fig. 1.
schmidti Khramova, 1962. - Ciliatocardium schmidti
Khramova, 1962. Cardiidae. Fossil. REcORDs. ··1974
ZR: Ciliatocardium schmidti Khramova, 1962 Comb.
nov. Ref. to Kafanov, p. 1469.
schmiereri Schlicum, 1964. Limnopagelia schmiereri
Schlicum, 1964. Cardiacea. Miocene. RECORDs. ··1964
ZR: Limnopagetia schmiereri Sch1icum, 1964 sp. nov.
Miocene, Brombach.
schoelleri Magna, 1950. - Ringicardium schoelleri
Magna, 1950. Cardiidae. Neogene. REcORDs. ··1950
ZR: Ringicardium schoelleri Magna, 1950 sp. nov.
Neogene Aquitaine.
schreteri Strausz, 1942. - New combination. Lymnocardium schreleri (Strausz, 1942). Lymnocardiidae.
Pre. Pliocene. RECORDS. ··1947 ZR: Limnocardium
(Pontalmyra) schreteri Strausz, 1942 sp. nov.
Som10jeno (Upper Pannon) [Pre. Pliocene).
schrvanica (error). - Printer's mistake for schirvanica
in ZR for 1981 with ref. to Andreescu.
schucherti McLearn, 1927. - Protocardia schucherti
McLearn, 1927. Cardiidae. Jura. REcORDs. ··1927 ZR:
Protocardia schucherti McLearn, 1927 sp. n. Jura.
Canada.
schuetti. - Limnopappia schuetti. Cardiacea. Fossil.
REcORDs. ··1962 ZR: Limnopappia schuett;' Viviparus
horizon. Genotype of Limnopappia Sch1ickum, 1962
of Cardiidae.
schulziYamani, 1975. -lurassicardium schulzi Yamani,
1975. Cardiidae. Upper Jurassic. RECORDS. ··1975 ZR:
lurassicardium schulzi Yamani, 1975 sp. nov. West
Germany, Bavaria, Ingo1stadt. Ref. to Yamani, p. 22.
sciloticum Sokolov, 1899. - Changed combination:
Parvicardium sciloticum (Sokolov, 1899). Cardiidae.
Fossil. RECORDS. ··1977 Popov: Parvicardium
sciloticum (Soko10v, 1899).
scitulum Meek, 1858. - Combination to be revised:
"Cardium" scitulum Meek, 1858. Cardiidae. Fossil.
RECORDS. ··1980 Ruhoff: Cardium scitulum Meek,
1858; vol. 4: 40.
scobinatum Lamarck, 1819. - Synonym of Parvicardium
papillosum (Poli, 1791). » Regarding Cardium
scobinatum: Cardiidae. Recent. TYPE LOCALJTY.
European Seas 7 (Lamarck). TYPE REPOSITORY. Museum
de Paris, but lost (Lamy 1942). DESCRIPTIONS. Ribs
close, echinate-squamose (Latin in Lamarck, translated
by lH.). DISTR1BUTION. Bonifacio (Locard & Caziot
1900). RECORDS. ··1819 Lamarck, p. 14, No. 38:
Cardium scobinatum. ··1829 Costa: Cardium
scobinatum = C. papillosum, Polio ··1835 Deshayes,
p. 408: Cardium scobinatum Lamarck. ··1836 Philippi:
Cardium scobinatum Lamarck = Cardium papillosum
Poli. ··1869 Romer: 7Cardium scobinatum Lamarck =
Cardium papillosum Po1i. ··1862 Weinkauff: Cardium
scobinatum Lamarck = Cardium papi/losum Poli.
··1867 Weinkauff: Cardium scobinatum Lamarck =
Cardium papi/losum Poli o ··1886 Locard: Cardium
scobinatum, de Lamarck, 18 J8 = Cardium papi/losum ,
Poli , 1791. ··1888 Kobelt: Cardium scobinatum Lam.
= Cardium papil/osum Poli o•• J888 Servain: Cardium
scobinatum Lamarck = Cardium papil/osum Poli, 1791 .
··1890 Paetel: Cardium scobinatum Lm. = papillosum
Poli . ··1892 Bucquoy; Dautzenberg & Dollfus:
Cardium scobinatum Lamarck = Cardium papillosum.
•• J900 Locard & Caziot: Cardium scobinatum in
Requien = Cardium papi/losum Poli, J791. •• J911
Dollfus : Cardium scobinatum Lamarck [junior
synonym] of Cardium (Parvicardium) papillosum Poli.
"1922-32 Sherborn, p. 5809: Cardium scobinatum
Lamarck, J 819. "1942 Lamy: Cardium scobinatum
Lamarck 7. ··1977 Fischer-Piette: Cardium scobinatum
= Cardium papillosum Poli 7
scobinella Deshayes, 1858. - Synonym of "Cardium"
sublima d 'Orbigny. » Regarding Cardium scobinella:
Cardiidae. Eocene . RECORDS . ··1886 Cossmann:
Cardium scobinella Deshayes = Cardium sublima
d'Orbigny. ··1980 Ruhoff: Cardium scobinella
Deshayes, 1858; [1856-1865] vol. 1: 564.
*scobinula. - Homonymy to be noticed. Cardium
scobinula from 2 sources: Merian in Deshayes, 1858.
- Cossmann & Lamb. (7year).
scobinula Cossmann & Lamb. - Synonym of "Cardium"
kochi Semp. » Regarding Cardium scobinula:
Cardiidae. Oligocene. RECORDS. ··1886 Cossmann:
Cardium scobinula. ··1886 Vincent: Cardium scobinula
756
Tropical Marine Mollusc Programme (TMMP)
scobinula - sedanense
Cossmann & Lamb . = Cardium kochi Semp. ··1936
ZR: Laevicardium scobinula figs . Authorship not
shown. [Ref. Gillet & Theobald 1936].
scobinula Merian in Deshayes, 1858. - Combination to
be revised: "Cardium" scobinula Merian . Cardiidae.
Fossil. RECORDS. "1892 B .D.D. p. 277: Cardium
scobinula Merian is related to Cardium papillosum
Goldfuss. ··1980 Ruhoff: Cardium scobinula Merian
in Deshayes, 1858; [1856--1865] vol. I : 562, pI. 56,
figs. 29-32. ··1999 Jacques Vidal (pers. com.) MNHN,
Paris: Fossils from the Paleontological Collection.
Stampien. Cardium scobinula Menan. France (Bassin
de Paris). Germany (Bade), B24907.
scripta (mode and variety) . - See under scriptum
Bucquoy, Dautzenberg & Dollfus, 1892 and exiguum
var. scripta Bucquoy, Dautzenberg & Dollfus, 1892.
scriptum Bucquoy, Dautzenberg & Dollfus, 1892. Changed combination: ParVicardium scriptum (B.D.D,
1892). OTHER JUDGEMENTS: Downgraded to variety
Parvicardium exiguum vai. scriptum. » Regarding
Cardium scriptum: Cardiidae. Recent. DISTRlBUnoN .
Mediterranean. RECORDS. ··1892 Bucquoy,
Dautzenberg & Dollfus: Cardium scriptum. "1969
Nordsieck: Parvicardium scriptum (Bucquoy,
Dautzenberg & Dollfus, 1892) = Parvicardium exiguum
(exiguum) (Gmelin, 1791) var. scriptum Bucquoi (sic
= Bucquoy), Dautzenberg & Dollfus. ··1998 Macedo
et af.: Parvicardium scriptum (B.D.D, 1892).
scrivetlori Kobayashi & Tamura, 1968. - Combination
to be revised : "Cardium" scrivenori Kobayashi &
Tamura, 1968. Cardiidae. Triassic. RECORDS. "1968
ZR: Cardium (s.l.) scrivenori Kobayashi & Tamura,
1968 sp. nov. Upper Triassic, Jurong Industrial Estate,
Singapore.
scrobiculatum Stoliczka, 1870. - Combination to be
revised: "Cardium" scrobiculatum (Pectunculus) Stoliczka, 1870. Cardiacea. Fossil. RECORDS. ··1980 Ruhoff:
Cardium scrobiculatum (Pectunculus) Stoliczka, 1870;
[1870-1871], vol. 3: 219, pI. II , fig. 4.
scruposum Deshayes, 1854. [Figs. Part 3 p. 907]. Changed combination: Fragum scruposum (Deshayes,
1855). OTHER JUDGEMENTS: Fragum fragum (Linnaeus).
» Regarding Cardium scruposum: Cardiidae. Recent.
TYPE LOCALITY . Malacca (syntype, coil. Cuming). TYPE
REPOSITORY. 3 syntypes (valves). BMNH London.
DISTRIBUTION . Malacca, Andaman Islands. RECORDS.
"1855 Deshayes (1854) p . 333: Cardium scruposum .
"1890 Paetel: Hemicardium scruposum Dsh. ··1977
Fischer-Piette: Cardium scruposum Deshayes =
Corculum fragum Linne. ··1980 Ruhoff: Cardium
scruposum Deshayes, 1854; p. 333. ··1999 Hylleberg,
p. 40 I: Fi-agum scruposum (Deshayes, 1855). ID 1.
Vidal. ··2000 HyUeberg, p. 532: Fragum scruposum
(Deshayes, 1855). Det. J. Vidal. ··2001 Vidal, p. 408:
Fragum scruposum. ··2003 Hylleberg & Kilburn, p.
178: Fragum scruposum (Deshayes, 1855) . CHARACTERS
AND
OCCURRENCE
DESCRIPTIONS. ··2001 Vidal: only the exhalant aperture
is tube-shaped. Inhalant siphon open towards the foot
aperture. No eyes. DISTRlBUTION. ··2000 Hylleberg:
Vietnam and Cambodia. ··2003 Hylleberg & Kilburn:
Vietnam.
scutellatum Chiereghini in Nardo, 1847. - Combination
to be revised: "Cardium" scutellatum Chiereghini in
Nardo, 1847. ?Cardiidae. Fossil. RECORDS. "1922-32
Sherborn, p. 5831 : Cardium scutellatum Nardo (ex
Chiereghini MS), 1847.
scutila Papaianopol & Lubenescu, 1985. Pseudoprosodacna scutila Papaianopol & Lubenescu,
1985. Lyrnnocardiidae. Upper Miocene. TYPE LOCALITY.
Ie Ponti en et Ie Dacien. Basin Dacique, Romania.
RECORDS. "1985 Papaianopol & Lubenescu, p. 138,
fig.: Pseudoprosodacna scutila sp. nov.
scutila Papaianopol , 1989. - Pachydacna scutila
Papaianopol, 1989. Lymnocardiidae. Pliocene. TYPE
LOCALITY. Basin Dacique, Romania. RECORDS. "1989
Papaianopo1, p. 83, fig.: Pachydacna (Pachydacna)
scutila sp. nov.
secans Fuchs, 1870. - New combination: Lymnocardium
secans (Fuchs, 1870). Lymnocardiidae . Fossil.
GENOTYPE. DoHfus (1905) Limnocardium secans
Fischer is type of Myocardia v. Vest. RECORDS. ··1899
Sacco, p. 58: Limnocardium secans (Fuchs, 1870).
"1905 Dollfus, commenting Andrussow 1903:
Limnocardium secans Fischer.
secundum Petiver. - Unknown status. Cardiidae .
RECORDS. ··1789 Bruguiere: Fragum secundum Petiver.
securiformis Cragin, 1894. - Synonym of Protocardia
filosa (Conrad, 1857) . » Regarding Trigonia
securiformis: Cardiidae. Cretaceous. RECORDS. ··1986
Scott: Trigonia securiformis Cragin , 1894 =
Protocardia filosa (Conrad, 1857) . ··1986 Scott:
Proto cardia securiformis Cragin, 1895 = Protocardia
filosa (Conrad, 1857). "1986 Scott: Roudairia
securiformis Cragin, 1895 = Protocardia filosa
(Conrad , 1857). ··1987 ZR: Protocardia securiformis
(Cragin, 1893) Junior synonym of jilosa (Conrad,
1857). Ref. to Scott p. 1203.
securiformis Slodkewitsch, 1938 . - Changed
combination: Fulvia securiformis (Slodkewitsch,
1938). Cardiidae. Pliocene. TYPE LOCALI1Y. K.amtchatka.
RECORDS. ··1938 Siodkewitsch: Papyridea securiformis
(fide Keen 1973). ··1973 Keen: Fulvia securiformis
(Slodkewitsch, 1938). "1976 ZR: Profulvia
securiformis (Slodkewitsch, 1938) Comb . nov.
Transferred from Papyridea. Ref. to Kafanov, p. 482
[orig. p. III]. ··1990 Gladenkov & Sinelnikova:
Papyridea sacuriformis sic Slod.
securigera Talent, 1963 . - Cypricardinia securigera
Talent, 1963. Cardiacea. Devonian. RECORDS. "1963
ZR: Cypricardinia securigera Talent, 1963 sp. nov.
Kilgower Member, Tabberabbera Formation, Victoria,
Australia, Devonian.
securus Papaianopol, 1982. - Pseudocatillus securus
Papaianopol, 1982. Lymnocardiidae. Upper Miocene.
RECORD S. ··1982 ZR : Ps eudocatillus securus
Papaianop'ol, 1982 sp . nov. Romania . Ref. to
Papaianopol et af. p. 145.
secutus Ebersin, 1967. - Pseudocatillus secutus Ebersin,
1967. Lymnocardiidae. Pliocene. RECORDS. "1967
Ebersin: Pseudocatillus secutus sp. n. ··1967 ZR:
Pseudocatillus seculus Eberzin, 1967 sp. nov. Pliocene,
USSR.
sedanense Pannekoek, 1936. - Combination to be
revised : "Cardium" sedanense (Pannekoek, 1936).
Cardiidae. Lower Miocene . TYPE LOCALITY. Java .
REMARKS. Elongated shape, strong rib sculpture;
compared with sp%ngense by its author. RECORDS.
Phuket Marine Biological Center Special Publication vol. 30 (2004)
757
sedgwichi - semiasperum
··1999 Vidal: Cardium (Trachycardium) sedanense
Pannekoek, 1936: 72, pI. 4, figs 51-5Ia.
sedgwichi Deshayes, 1853. - Combination to be revised:
"Cardium" sedgwichi Deshayes, 1853 . Cardiacea.
Fossil. RECORDS. ··1980 Ruhoff: Cardium sedgwichi
Deshayes, 1853; vol. 2: 45.
seikaiana Amano, Ogata, & Nire, 1958. - Changed
combination: ?Protocardia seikaiana Amano, Ogata,
& Nire, 1958. Cardiidae. Cretaceous. RECORDS. ··1958
Amano, Ogata, & Nire: Tendagurium seikaiana (jide
Keen 1973). ··1973 Keen: ?Protocardia (Tendagurium)
seikaiana (Amano, Ogata, & Nire, 1958).
selene Vidal, 1999. [Figs. Part 3 p. 877]. -Acrosterigma
selene Vidal, 1999. Cardiidae. Recent. TYPE MATERIAL.
All shells from several dredging campaigns in the
vicinity of New Caledonia, MNHN, Richer de Forges
ORSTOM collection. Holotype: A bivalved shell (Figs
5G, 5H, 51), S zone of Lagoon, 40-50 m. Paratype I: A
left valve (Fig. 5J),Lansdowne-Fairway Plateau, 6770 m. Para type 2: A left valve, south-east of New
Caledonia ridge, 200m . Paratype 3: A right valve,
Chesterfield-Bellona plateau, 40 m. Paratype 4: A left
valve, Lansdowne-Fairway Plateau, 69m. Paratype 5:
A bivalved shell, Grand Recif Sud, 70m. Paratypes 6
to 10: Six smaller valves or bivalved shells, from
Chesterfield and Landsdowne-Fairway areas. TYPE
LOCALITY. New Caledonia. RECORDS. ··1999 Vidal, p.
289, Table 16, Figs. 5G-J: Acrosterigma selene sp. nov.
- CHARACTERS AND OCCURRENCE - CHARACTERS of diagnostic value. It differs from all Acrosterigma
species by its extremely hollowed lunule (Vidal 1999).
DESCRIPTION. Vidal (1999): Shell almost equilateral,
neither elongated nor inflated, and very slightly
truncated on posterior margin. Lunule well delineated
and extremely hollowed on both sides, the right being
wider. External colour yellowish-beige, with irregular
darker zones and pink to orange spots. Interior whitish
with traces of external spots, and two pink umbonal
rays which are also visible exteriorly. Hinge moderately
angled, and strongly asymmetrical in contrast to the
shell itself, which is almost perfectly equilateral. Mean
rib number 36.6 ± 0.8 (range 35-38). On posterior shell,
the anterior smooth part of ribs much wider than
posterior part; intennediate furrow very thin and not
always visible in adult region . Scales elongated,
sometimes prismatic, straddling the longitudinal
furrow, and often almost longitudinally disposed rather
than oblique. In median-posterior area, ribs triangular
and slightly asymmetrical with posterior flank flat,
shorter, and more abrupt, and anterior flank slightly
convex. Ribs themselves practically smooth (rarely
slightly retro-ridged), with a crestal fold marked by
regular, short undulations. On anterior half, ribs become
progressively less triangular, then rounded, first retroridged or crenulated, then top-ridged. Interstices
rounded, smooth, and as wide as ribs. DIMENSIONS.
Length 32.9, width 35.3, thickness (calculated) 23.6
mm . REMARKS . Vidal (1999): Acrosterigma selene
superficially resembles A. mauritianum in shape and
colour, but has fewer ribs and its rib morphology is
different (ribs more triangular) . It differs from all
Acroslerigma species by its unique lunule.
DISTRIBUTION . South east of New Caledonia ,
Chesterfield Bellona Plateau and Landsdowne-F airway
banks.
sellardsi Gardner, 1926. - Combination to be revised:
"Cardium " sellardsi Gardner, 1926. Cardiidae. Fossil.
RECORDS. ··1926 Gardner, p. 140, pI. 23, figs . 11-12:
Cardium [(Fragum) (Trigoniocardia)] sellardsi. ··1926
ZR: [Fragum (Trigoniocardia)] sellardsi Gardner, 1926
sp. n. Tertiary U.S.A. ··1977 Vokes: Cardium
[(Fragum) (Trigoniocardia)] sellardsi Gardner, 1926.
se/lii Gillet, 1970. - Plagiodacna sellii Gillet, 1970.
Lymnocardiidae. Miocene. RE CORDS. ··1970 ZR:
Plagiodacna sellii Gillet, 1970 sp. nov. Miocene,
'Monte dei Corvi' deposits, Ancona district, Italy.
semialatum Andrzejovski, 1832. - Nomen nudum. Not
available. RECORDS. ··1857 Deshayes: Cardium
semialatum. Audrz. sic. 1836. ··1922-32 Sherborn, p.
5831: CardiumsemialatumAndrzejovski, 1832 [no n.].
semialatum Munster, 1840. - Cardium semialatum
Muenster, 1840. Conocardiidae. Fossil. RECORDS.
··1922-32 Sherborn, p. 5831: Cardium semialatum
Muenster, 1840.
semialatum Orbigny, 1850. - Transferred to
Conocardium semialatum d'Orbigny, 1850.
Conocardiidae. Fossil. RECORDS. ··1922-32 Sherborn,
p. 5831: Conocardium (Muenster) semialatum
d'Orbigny, 1850. - Cardium .
semiasperum Deshayes, 1858. - Changed combination:
Nemocardium semiasperum (Deshayes, 1858).
Cardiidae. Eocene. Type species: Sacco (1899)
designated Cardium semiasperum Deshayes, 1858 type
species of the genus Nemocardium Meek, 1876. Dall
(1902) Cardium semiasperum Deshayes is type of
section Nemocardium Meek. Keen (1954) and Scott
(1986): Cardium semiasperum Deshayes, 1858 is type
species of Nemocardium Meek, 1876. TYPE LOCALITY.
France, Paris Basin. RECORDS. ··1858 Deshayes:
Cardium semiasperum. ··1886 Cossmann:
Protocardium semiasperum (Deshayes). ··1886 Dall:
Cardium semiasperum Deshayes. ··1887 Fischer:
Lyrocardium semiasperum Deshayes. ··1899 Sacco:
Cardium semiasperum Deshayes, 1858. ··1902 Dall:
Cardium semiasperum Deshayes. ··1925 Woodring, p.
146: Cardium semiasperum Deshayes. Eocene, Paris
Basin . ··1947 ZR: Nemocardium semiasperum
(Desha yes ). Cardium semiasperum Deshayes, Eocene,
France, type of Nemocardium Meek 1876, subseq.
desig. Sacco 1899. [Ref. Vokes 1946]. ··1950 Keen :
Nemocardium semiasperum (Deshayes, 1860 sic) .
··1954 Keen: Cardium semiasperum Deshayes, 1858.
··1986 Scott: Cardium semiasperum Deshayes, 1858.
··1969 Moore et al.: Nemocardium (Nemocardium)
semiasperum (Deshayes, 1858). ··1977 Popov:
Nemocardium (Nemocardium) semiasperum. ··1977
Popov : Cardium semiasperum Deshayes, 1858. Type
of Nemocardium Meek, 1876. ··1977 Wilson &
Stevenson: Cardium semiasperum Deshayes, 1858
[Eocene]. ··1980 Ruhoff: Cardium semiasperum
Deshayes, 1858; [1856--1865] vol. 1: 573, pI. 55, figs.
1-2. ··1992 Schneider p. 155: Nemocardium
semiasperum (Deshayes). Type species of
Nemocardium. ··1995 Le Renard et al.: Nemocardium
(s. Sir.) semiasperum [Cardium semiasperum]
Deshayes, 1858. ··1995 Pacaud & Renard:
Nemocardium (Nemocardium) semiasperum . ··1999
Jacques Vidal (pers. com.) MNHN, Paris: Fossils from
758
Tropical Marin e Mollusc Programme (TMMP)
semicinetum - semis/riatum
the Paleontological Collection as Nemocardium
semiasperum Deshayes. Ypresien. France (Bassin de
Paris), B700580.
semicinetum Mi.inster, 1840. - Combination to be
revised: "Cardium " semicinetum Mi.inster, 1840.
Cardiidae. Fossil. RECORDS. 001922-32 Sherborn, p.
5853: Cardium semicinetum Muenster, 1840.
semicostata. - Pholadomya semicostata. Cardiacea.
Fossil. RECORDS . 001842 Agassiz : Pholadomya
semicostata (comparison with Pholadomya crispaAg.).
semicostatum Lycett, 1850. - Combination to be revised:
"Cardium" semicosta/um Lycett, 1850. Cardiidae.
Fossil. REcoRDs. 001922-32 Sherborn, p. 5854: Cardium
semicosta/um Lycett, 1850. 001980 Ruhoff: Cardium
semicostatum Lycett, 1850; vol. 6: 422. 001980 Ruhoff:
Cardium semicostatum Morris & Lycett, 1853; [18531854] vol. 2: 63, pI. 7, figs. 6 a,b.
*semiglabrum . - Homonymy to be noticed. Cardium
semiglabrum from 2 sources: Phill ips, 1829. Goldfuss, 1840.
semiglabrum Goldfuss , 1840. - Transferred to
lsocardia? semiglabrum (Goldfuss, 1840). ?Isocardiidae. Fossil. RECORDS. 001922-32 Sherborn, p.
5857: Cardium semiglabrum Muenster in Goldfuss,
1837. 001857 Deshayes: Cardium semiglabrum
Goldfuss, 1840. - lsocardia?
semiglabrum Phillips, 1829. - Nomen nudum
accompanied by figure. » Regarding Cardium
semiglabrum: Cardiidae. Fossil. RECORDS . 001857
Deshayes: Cardium semiglabrum Philips , 1829.
001922-32 Sherborn, p. 5857: Cardium semiglabrum
Phillips , 1829 [n o et f.] . 001947 ZR: Cardium
semiglabrum Phillips [Ref. Melmore 1947].
semiglobosum Matheron. - Combination to be revised :
"Cardium" semiglobosum Matheron. Isocardiidae.
Fossil. RECORDS. 001857 Deshayes: Cardium
semiglobosum Matheron. Not a cardiid: in genus
lsocardia.
semigranosum Sowerby. - Identical with Nemocardium
semislrialum oligocenica Sacco. » Regarding C.
semigranosum: Cardiidae. Oligocene. RECORDS. 001835
Lamarck: Cardium semigranulosum Sow. 001899
Sacco, p. 57 : Cardium semigranosum Sow. No. 1535,
Sacco 1889 . = Nemocardium semistriatum var.
oligocenica Sacco.
*semigranulatum . - Homonymy to be noticed.
Cardium semigranulatum from 2 sources: Sowerby,
1816. - Sowerby 1847.
semigranulatum Sowerby, 1816. - New combination.
Nemocardium semigranulatum (Sowerby, 1816).
Cardiidae. Fossil. RECORDS. 001922-32 Sherborn, p.
5858 : Cardium semigranulatum Sowerby, 1816. 001977
Popov: Nemocardium (Nemocardium) semigranulatum
(Sowerby, 1816).
semigranulatum of Sowerby 1847. - Replaced by
Laevicardium taurovalum Sacco, 1899. » Regarding
Cardium semigranulatum: Cardiidae. Fossil. RECORDS.
001899 Sacco: Cardium semigranulatum Sowerby 1847
= Laevicardium taurovatum Sacco, 1899. 001922-32
Sherborn, p. 5858: Cardium semigranulatum Sowerby
in Dixon, 1850.
semilineatum Koenen, 1864. - Nemocardium
semiliniatum (Koenen, 1864). Cardiidae. Oligocene.
RECORDS. 001899 Sacco, p. 54: Laevicardium
semilineatum (Koen.). 001977 Popov: Nemocardium
(Habecardium) semilineatum (Koenen, 1864).
seminulum Fr. Roemer, 1852. - Combination to be
revised: "Cardium " seminulum Fr. Roemer, 1852.
Cardiidae. Fossil. RECORDS. 001980 Ruhoff: Cardium
seminulum Fr. Roemer, 1852; [1850-1852] vol. 3: 79,
pI. 12, fig . 12.
semipapillatum Reuss, 1844. - Combination to be
revised: "Cardium " semipapillatum Reuss, 1844.
Cardiidae. Fossil. RECORDS. 001922-32 Sherborn, p.
5864: Cardium semipapillatum Reuss, 1844.
semipe/lucida Logvinenko & Starobogatov, 1967. Hypanis semipellucida Logvinenko & Starobogatov,
1967. Lymnocardiidae. Recent. RECORDS . 001968
Logvinenko & Starobogatov: Hypanis semipellucida
Logvinenko & Starobogatov, 1967.
semiplenum White. - Conocardium semiplenum White.
Conocardiidae . Fossil. RECORDS. 001943 ZR:
Conocardium semiplenum White Ref. to Branson
(\ 942).
sem;punctatum Mi.inster in Goldfuss, 1837. Combination to be revised : "Cardium" semipunctatum
Mi.inster in Go1dfuss, 1837. Cardiidae. Fossil. RECORDS.
001922-32 Sherborn, p. 5865 : Cardium semipunctatum
Muenster in Goldfuss, 1837.
semipustulosum - Combination to be revised:
"Cardium " semipustulosum. Cardiidae. Fossil.
RECORDS. 001889 ZR: Cardium semipustulosum with
fig. (ref. Holzapfel). 001922-32 Sherborn, p. 5866:
Cardium semipustulosum J. Mi.iller, 1846.
semiseptiferum d'Orbigny, 1850. - Combination to be
revised : "Cardium" semiseptiferum d'Orbigny, 1850.
Cardiidae. Lias-Jurassic . RECORDS. 001922-32
Sherborn, p. 5868 : Cardium semiseptiferum d'Orbigny,
1850. 001999 Jacques Vidal (pers. com.) MNHN, Paris:
Fossils from the Paleontological Collection. Cardium
semiseptiferum d'Orbigny. Rauracien. France (Yonne),
S04051.
semistriatulum Deshayes . - RECORDS . 001922-32
Sherborn, p. 5868: Cardium semistriatulum Deshayes;
err. Brown pro semis/riatum q.v.
*semistriatum. - Homonymy to be noticed. Cardium
semistriatum from 3 sources: Bean in Young & Bird,
1828. - Deshayes, 1829. Mi.inster, 1840.
semistriatum Bean in Young & Bird, 1828. Combination to be revised: "Cardium " semistriatum
Bean in Young & Bird, 1828. Cardiidae. Fossil.
RECORDS . 001922-32 Sherborn, p. 5868: Cardium
semistriatum Bean in Young & Bird, 1828. 001994
Renard, J. Ie: not Cardium semistriatum Deshayes,
1829.
semis tria tum Deshayes, 1829. - Replaced by
Nemocardium (Nemocardium) keenae Renard, 1994.
» Regarding Cardium semistriatum : Cardiidae.
Eocene. RECORDS. 001829 Deshayes: Cardium
semis/riatum. 001835 Lamarck: Cardium semistriatum
Desh. 001857 Deshayes: Cardium semis/ria/um Desh.
1830 sic. 001886 Cossmann: Protocardium
semis/ria/um (Deshayes). 001922-32 Sherborn, p. 5868:
Cardium semistria/um Deshayes, 1829. 001977 Popov:
Nemocardium (Nemocardium) semis/ria/um
(Deshayes, 1858). 001994 Renard, J. Ie, p. 39: Cardium
semistriatum Deshayes 1829; preoccupied name
replaced by Nemocardium (Nemocardium) keenae.
Phuket Marine Biological Center Special Publication vol. 30 (2004)
759
semistriatum - senticosum
··1995 Le Renard et at.: Nemocardium (s. str.) keenae
Le Renard, 1994 (= semistriatum Desh. non Bean).
··1999 Jacques Vidal (pers. com.) MNHN, Paris:
Fossils from the Paleontological Collection as Cardium
(and Nemocardium) semistriatum Deshayes. Lutetien.
Lower Eocene. France (Bassin de Paris). U.K. (Hants),
B21200. Turkey, B2233 I.
semistriatum Munster, 1840. - Conocardium
semistriatum (Munster, 1840). OTHER JUDGEMENTS:
Cardium lunatum Deshayes. » Regarding C.
semistriatum: Conocardiidae. Silurian-Devonian.
REMARKS . Lunulicardium Munster, 1840 not in
Cardiidae (fide Fischer 1887). RECORDS . "1857
Deshayes: Cardium semistriatum Munster. 1840. - C
lunatum. Desh. ··1887 Fischer: Lunulicardium
semis tria tum Munster, 1840. "1922-32 Sherborn, p.
5869: Lunulacardium semis/ria/um Muenster, 1840.
··1922-32 Sherborn, p. 5869: Conocardium (Muenster)
semistriatum d'Orbigny, 1850. - Lunulacardium.
semistriatum oligocenica (cf.) Sacco, 1899 (variety).Nemocardium cf. semis tria tum oligocenica Sacco,
1899. Cardiidae. Oligocene. RECORDS. "1899 Sacco,
p. 57, pI. 12, fig. 24: Nemocardium cf. semistriatum
var. oligocenica Sacco.
semisulcata (mode). - See under semisulcatum Gray,
1825.
*semisulcata. - Homonymy to be noticed. Cardium
semisulcata from 2 sources: Gray, 1825. - Rousseau
in Demidoff, 1842.
semisulcata Gray, 1825. [Figs. Part 3 p. 916]. - Changed
combination: Papyridea semisulcata (Gray, 1825).
Cardiidae. Pliocene and Recent. TYPE LOCALITY. Cuba .
RECORDS. "1825 Gray, p. 137: Cardium semisulcatum.
"1885 Smith: Cardium (Papyridea) semisulcatum,
Gray. 1825. ··1902 Dall: Cardium (Papyridea)
semisulcatum Gray, 1825 [Pliocene to Recent) . ··1902
Dall & Simpson: Cardium semisulcatum Gray. ··1934
Johnson: Papyridea semisulcatum Gray 1825. (C
petiana Orbigny 1846). ··1936 McLean: Papyridea
semisulcatum Gray. ··1939 McLean: Papyridea
semisulcata (Gray, 1825) . "1939 ZR: Papyridea
semisulcatum [Ref. McLean 1939]. ··1953 Olsson &
Harbison: Papyridea semisulcatum (Gray, 1825).
"1957 Nowell-Usticke: Papyridea semisulcata
Sowerby. ··1958 Abbott: Papyridea semisulcata Gray
1825 senior synonym of Papyridea petitianum Orbigny
1846, which is the orange form . ··1962 Warmke &
Abbott: Papyridea semisulcata Gray, 1825 . "1963
Coomans: Papyridea semisulcata (Gray). ··1968 Jong
& Kristensen: Papyridea semisulcata Gray. ··1968
Kempf & Matthews: Papyridea semisulcata Gray,
1825 . •• I 970 Rios: Papyridea semisulcata (Grey,
1825) . .. 1973 Waller: Papyridea semisulcata (Gray,
1825). ··1977 Fischer-Piette: Papyridea semisulcatum
Gray. - CHARACTERS AND OCCURRENCE DESCRIPTIONS. Dall & Simpson (1902): Shell small,
elliptical, thin, very inequilateral, moderately inflated,
with numerous low ribs, and with more or less perfectly
developed smaller ribs between the larger ones, the
margin at the posterior end sharply toothed; surface
covered with fine concentric growth lines, which are
stronger on the anterior part of the shell; interior
showing the sculpture; hinge teeth compressed. Colour
whitish or yellowish, with faint brown blotches and
spots. McLean (1939): 24-25 narrow ribs. DIMENSIONS.
13 x 9 x 7 mm (McLean 1939); length 10, height 6.5,
diameter 5 mm (Dall & Simpson 1902). DISTRIBUTION.
South Florida, West Indies, Brazil.
semisulcata Rousseau in Demidoff, 1842. - Changed
combination: Prosodacna semisulcatum (Rousseau in
Demidoff, 1842) . Lymnocardiidae. Fossil. TYPE
LOCALITY. Crimean. RECORDS. ··1842 Rousseau, vol. II,
p. 810, pI. IX, fig . I : Cardium semisulcatum . .. I 886
Fischer: Cardium semisulcatum Rousseau. ··1886
Fischer: Prosodacna semisulca/um Rousseau, 1842.
.. I 905 Dollfus, commenting Andrussow 1903:
Prosodacna semisulcata Rouss. ··1907 Andrussow:
Prosodacna semisulcata Rouss. ··1922-32 Sherborn,
p. 5870: Cardium semisulca/um Rousseau in Demidoff,
1842.
semisulcata angustata Eberzin, 1960 (subspecies). Prosodacna semisulcata angustata Eberzin, 1960.
Lymnocardiidae . Pliocene. RECORDS. ··1960 ZR:
Prosodacna (Prosodacna) semisulcata angustata
Eberzin , 1960 subsp . n. Pliocene Galizgi river,
Abkhaziya, Georgia USSR.
semisllicata antiqlla Eberzin, 1960 (subspecies). Prosodacna semisulcata antiqua Eberzin, 1960.
Lymnocardiidae. Pliocene. RECORDS. ··1960 ZR:
Prosodacna (Prosodacna) semisulcata antiqua
Eberzin, 1960 subsp. n. Pliocene Kamyshburun
Kerchensky, Crimea USSR.
semisulcata babadjanica Andreescu, 1980. Replacement name: Prosodacna semisulcata
babadjanica. Lymnocardiidae. Tertiary. RECORDS.
··1980 ZR: Prosodacna semisulcata babadjanica.
Replacement name for P schirvanica major Andrusov
& Eberzin. [Ref. to Andreescu).
semisllicatoides (subspecies). - See under littoralis
semisulcatoides Eberzin, 1960.
semisulcatum (mode and subspecies). - See under
semisulcata Rousseau, 1842 and hiatus semisulcatum
Gray.
semisllicatus Sacco, 1899. Discors semisulcatus (Sacco,
1899). Cardiidae. Fossil. RECORDS. "1970 Glibert &
Poel, p. 33: Discors semisulcatus (Sacco, 1899) [Raised
from Discors discrepans semisulcata Sacco, 1899, p.
54].
semitectllm Marwick, 1926. - Nemocardium semi tectum
Marwick , 1926. Cardiidae. Cretaceous. RECORDS.
··1926 ZR: Nemocardium semitectum Marwick, 1926
sp. n. Cretaceous N. Zealand.
semseyi Hal. - Changed combination: Budmania semseyi
(Hal.). Lymnocardiidae. Fossil. RECORDS ... I 987
Taktakishvili: Limnocardium semseyi Hal. - Budmania
semseyi.
senegalensis (variety) . - See under norvegicum var.
senegalensis Dautzenberg.
seninskii Andrus. - Changed combination : Euxinicardium seninskii (Andrus.). Lymnocardiidae. Fossil.
RECORDS . •• I 987 Taktakishvili: Limnocardium
(Euxinicardium) seninskii Andrus. - Euxinicardium
seninskii.
senticosllm Sowerby, 1833. [Figs. Part I p. 107, 173.
Part 3 p. 923]. - Trachycardium senticosum (Sowerby,
1833). OTHER JUDGEMENTS: Trachycardium radula
(Broderip & Sowerby, 1833). - Trachycardium
quadragenarium (Conrad , 1837). » Regarding
760
Tropical Marine Mollusc Programme (TMMP)
Sepidacna - Septocardiidae
Cardium senticosum: Cardiidae. Fossil and Recent.
TYPE LOCALITY. Hab. ad Sanctam Elenam, Americae
Meridionalis, in sandy mud at from 6-12 fathoms depth
(Sowerby 1833). RECORDS. --1834 Sowerby, 47th part,
fig. 10: Cardium senticosum nobis. "1840 Sowerby,
catalogue p. 3, No. 43: Cardium senticosum, Sow.
.. 1840 Sowerby, corrected list p. 8: Cardium
senticosum, (Sowerby No. 43). --1845 Reeve: C. radula
Reeve = C. senticosum Sowerby. --1852 Adams: 434.
Cardium senticosum Sowb. 1833. "1860 Carpenter:
Cardium senticosum Sby. --1884 Grasset: Cardium
senticosum. Sow. --1890 Paetel: Cardium senticosum
Sow. --1891 Steams: Cardium senticosum Sby. --1895
Steams: Cardium senticosum Sowerby. .. 190 I Dall:
Cardium (Trachycardium) senticosum Sowerby, 1833.
"190 I Dall : Cardium senticosum Pacific coast
analogue of Cardium muricatum Linnaeus. --1922-32
Sherborn, p. 5882: Cardium senticosum Sowerby, 1833.
"1928 Tomlin: Cardium (Trachycardium)senticosum
(SOw. , 1833). --1934 Hertlein: Cardium senticosum
Sowerby [fossil]. "1947 Hertlein & Strong: Cardium
(Dal/ocardia) senticosum Sowerby, 1833. "1961
Olsson: Trachycardium (Dal/ocardia) senticosum
(Sowerby, 1833) . "1964 Parker : Trachycardium
(Dallocardia) senticosum (Sowerby, 1833). "1971
Keen & McLean: Trachycardium (Dallocardia)
senticosum (Sowerby, . 1833) . "1977 Popov :
Trachycardium (Dal/ocardia) senticosum (Sowerby,
1833). --1977 Fischer-Piette: Laevicardium senticosum
Sowerby. --1991 Voskuil & Onverwagt: Laevicardium
(Trachycardium) senticosum Sowerby; junior synonym
of Trachycardium quadragenarium (Conrad, 1837).
--1999 Vidal: Trachycardium senticosum (Sowerby in
Broderip & Sowerby, 1833). - CHARACTERS AND
OCCURRENCE - OF COLLECTIONS. ZMUC from the
Gulf of California; Rulejo (Oersted coil. No. 325).
DESCRlPTIONS. Hertlein & Strong (1947): Shell rounded,
gently convex; in total 36-39 ribs, which in their earlier
stages are subtriangular but later become subrounded;
ribs bear obliquely projecting scale-like imbrications
which toward the ventral margin develop into oblique
nodes; nodes are strongly developed on the posterior
side of central ribs, may also present, but to a lesser
degree, on the anterior side; nodes may be present on
top or on the anterior side of anterior ribs; nodes
strongly developed on the posterior side of posterior
ribs . Exterior brownish-white or purple, interiorly the
portion under the umbo is usually reddish-purple; the
remainder, and sometimes all of the shell, is white.
Variation : specimens on a muddy bottom may be
thinner and less coloured than the other specimens.
Keen (1971): Compared with Trachycardium consors,
this is smaller, lighter in color, with more ribs (35 to
40), which bear only short, thornlike scales, most
prominent on the anterior part of the shell. The coloring
is basically white, inside and out, with reddish-brown
or purple bands and spots exteriorly. It attains a height
of about 40 mrn. The recorded range includes the whole
of the Gulf of California, southward to Paita, Peru.
Really good specimens can ordinarily be obtained only
by dredging on shallow muddy bottoms. The very
similar Atlantic analogue of this species is
Trachycardium (Dallocardia) muricatum (Linnaeus,
1758). DIMENSIONS. Altitude of 40 mm (Hertlein &
Strong 1947). REMARKS. Cardium muricatum Linnaeus
from the western Atlantic is a similar species. (Hertlein
& Strong 1947), but C. muricatum is rounder than C.
muricatum, and tubercles not so oblique (Sowerby
1841). Fischer-Piette (1977) identified a particular
cockle as Laevicardium (Tra chycardium) senticosum
(Sowerby, 1833) labeled as from Monterey, California .
According to Keen (1980) it should be Trachycardium
(Dallocardia) quadragenarium (Conrad, 1837).
HABITAT. In sandy mud, at from 6 to 12 fathoms depth
(Adams 1852); on clay-sand at 11-29 m depth
(ZMUC) ; occasionally to a depth of about 100 m
(Parker 1964) mostly on muddy bottoms (Hertlein &
Strong 1947). DISTRIBUTION . C. senticosum is known
from Columbia Occ. (Gras set 1884); Gulf of California
to Peru (Keen & McLean 1971). C. radula is known
from California to Peru.
Sepidacna (genus). - Tridacnidae. RECORDS. --1937 ZR:
Sepidacna Iredale, ) 937 gen. n. Tridacnidae type
Tridacna trough toni.
septemputeallus Ebersin, 1967. - Pseudocatillus
septemputeanus Ebersin, 1967. Lymnocardiidae .
Pliocene. RECORDS. --1967 Ebersin: Pseudocatil/us
septemputeanus sp. n. --1967 ZR: Pseudocatillus
septemputeanus Eberzin, 1967 sp . nov. Pliocene,
USSR. ··1984 ZR: Pseudocatillus septemputeanus
Ebersin. Referred to new subgenus P (Donacicatillus).
Ref. to Akhvlediani p. 71 .
septiferum Buvignier, 1843 . - Combination to be
revised: "Cardium" septiferum Buvignier. Cardiidae.
Lias-Jurassic. RECORDS. ··1922-32 Sherborn, p. 5889:
Cardium septiferum Buvignier, 1843. "1999 Jacques
Vidal (pers. com.) MNHN, Paris: Fossils from the
Paleontological Collection. Cardium septiferum
Buvignier. Rauracien . France (Yonne). S04455,
B 12335, S04686.
Septocardia Hall & Whitfield, 1877 (genus). - Cardiidae.
Upper Trias. GENOTYPE. Hall & Whitfield (1877)
designated Septocardia typica type species of genus
Septocardia . RECORDS. ··1980 Vokes: Septocardia Hall
& Whitfield, 1877 in subfamily Cardiinae. ··1992
Schneider p. 149: Septocardia Hall & Whitfield, 1877.
Referred to Septocardiinae from Cardiinae,
Tridacninae . Downgraded from Tridacnidae
(Tridacnacea) . "2002 Schneider: Septocardia has been
tentatively rejected from Cardiidae. - CHARACTERS
AND OCCURRENCE - DESCRIPTIONS. ··1969 Moore
et al.: thick-shelled, ribs beaded, incremental lines
conspicuous in interspaces as cross-striae; ligament
inserted in broad and slightly oblique furrow bordered
by nymph with pointed summit; hinge plate wide,
anteriorly forming deep cavity for reception of anterior
adductor muscle; anterior lateral teeth small, placed at
junction of muscle cavity flange and hinge plate proper,
cardinal teeth with 2 salient, 4b lamellar, sloping
obliquely backward from beak, 3a-3b joined above;
posterior lateral teeth well developed; pallial line entire,
posterior muscle scar ovate, not impinging on hinge
plate; inner margin strongly crenulate. REMARKS. ··1969
Moore et al. : Septocardia seems to be ancestral stock
of Cardiacea , but with strong affinities to
Palaeolcardita. North and South America.
Septocardiidae Kafanov & Starobogatov, 1977 (tribus).
- Cardiidae. RECORDS. ··1977 Kafanov et Starobogatov:
Phuket Marine Biological Center Special Publication vol. 30 (2004)
761
septuagenarium - serra tum
Septocardiidae.
septuagenarium Tate, 1887. - Synonym of Hedeeardium
monileteetum. Changed combination. Cardiidae. Fossil.
REcoRDs. --2002 Schneider: Cardium septuagenarium
is considered a junior synonym of Cardium
monileteetum by Schneider (2002). --2002 Schneider:
Darragh (1970) assigned Cardium septuagenarium
Tate, 1887 to Hedeeardium.
sera. - Protoeardia sera. Cardiidae. REcORDs. --1914 ZR:
Protoeardia sera [Ref. Suter 1914].
serbica Brusina 1902. - Prosodaena serbiea Brusina
1902. Lymnocardiidae. Fossil. RECORDS. --1979 ZR:
Prosodaena serbica Brusina 1902. Topotype Ref. to
Stevanovic, p. 331 .
serbica (aff.) Brusina 1902. - RECORDs. "1991 Muller
& Magyar, p. 356: Lymnoeardium aff. serbicum
(Brusina 1902).
serena Papaianopol, 1980. - Luxuridaena serena
Papaianopo1, 1980. Lymnocardiidae. Fossil. RECORDS.
--1980 ZR: Luxuridaena serena Papaianopol, 1980 sp .
nov. Romania, Pliocene. Ref. to Papaianopo1, p. 51.
Seripes (error). - Printer's mistake for Serripes in
Nomura, 1933 (Vokes 1980).
serogosicum exile Popov, 1982. - Cerastoderma
serogosieum exile Popov, 1982. Cardiidae. Oligocene.
RECORDS. --1982. ZR: Cerastoderma serogosieum exile
Popov, 1982 subsp. nov. Georgia SSR. Ref. to Popov
et at. p. 42.
serogosicum Nossovsky, 1962. - Changed combination:
Cerastoderma serogosieum (Nossovsky, 1962) .
Cardiidae. Oligocene. RECORDS. --1962 ZR: Cardium
(Cerastoderma) serogosieum Nossovskii, 1962 sp. nov.
Oligocene, Khersonsk province, USSR --1977 Popov:
Cerastoderma serogosicum (Nossovsky, 1962).
serratulum (variety) . - See under buekkianum var.
serratulum Noszki, 1936.
*serratum. - Homonymy to be noticed. Cardium
serratum from 2 sources: Linnaeus, 1758. - Bruguiere,
1789.
*serratum. - Summary of identifications: C. serratum
are American Atlantic species, rather variable in shape
and colour, but always identifiable by some other
constant specific characters (Vidal pers. comm.). C.
serra tum Linnaeus, 1758. has two synonyms: C.
oviputamen Reeve, 1844 and C. citrinum serratum
Chemnitz, 1782 [not available]. Many European and
American authors were right about the ID of C.
serratum before the regrettable intervention of Clench
& Smith, 1944, who named this species C. laeviga/um.
Unfortunately this erroneous interpretation was
immediately adopted by the majority of authors until
today. It has often been misidentified by authors. The
name Cardium serra tum has erroneously been used for
[I] Cardium biradia/um Bruguiere , 1789. "1784
Schroter, p. 46: Cardium serratum . "1799 Spengler,
p. 43: Cardium serra/um. [2] Cardium oblongumerassum complex. Bruguiere, 1789, p. 229: Cardium
serra/um Linn. Cardium serra/um Nob. [3] Cardium
erassum Gmelin, 1791. --1819 Lamarck, p. 11:
Cardium serra/um . .. 1835 Deshayes et al. p. 401:
Cardium serra/um. "1841 Sowerby, p. 5: Cardium
serra/um . [4] Cardium pristis Valenciennes, 1827.
--1970 Vilas & Vilas p, 39: Laevicardium serra/um var.
laeviga/um . The name Cardium serratllm has correctly
been used to name the American species by: --1822
Turton [erroneous locality: British Isles]. - "1828
Fleming [erroneous locality] . - --1844 Reeve. - --1847,
"1853 Orbigny. - "1869 Romer. - "1886, "1900,
"1901 Dalt. - "1855 Hanley. - "1900 Dautzenberg.
- --1922 Pi1sbry. - --1925 Woodring. - --1926
Weisbord. - --1929 Tomlin. - --1939 MacLean.
serratum Bolten, 1798. - Synonym of Laevieardium
serratum (Linne, 1758). RECORDS. --1798 Bolten:
Cardium serra/um = Cardium laevigatum Gmelin sp.
18. --1902 Sherborn, p. 893 : Cardium serralum Bolten,
1798.
serratum Bruguiere / Lamarck. [Figs. Part 1 p. 76. Part
3 p. 896]. - Identical with Laevieardium erassum
Gmelin. OTHER JUDGEMENTS : Cardium norvegieum
Spengler, 1799 = Laevicardium erassum (Gmelin,
1791) = Cardium oblongum = Cardium sulcatum
Lamarck, 1819. » Regarding Cardium serratum:
Cardiidae. Recent. RECORDS. --1789 Bruguiere, p. 229,
No. 27: Cardium serra/um Linn. & Cardium serra tum
Nob. --1798 Bruguiere. ID by Lamarck, pI. 299. fig. 2.
Cardium serratum . --1815 Wood: Cardium serra tum
of Bruguiere = Cardium oblongum Linn. "1817
Dillwyn: Cardium serra tum Bruguiere = Cardium
oblongum Chemnitz. --1827 Bory de St. Vincent :
Cardium serratum Lmk . = Laevieardium crassum
Gmelin (fide Fischer-Piette 1977). --1832 Deshayes:
Cardium serratum Brug. = Cardium suleatum Lamk.
--1835 Deshayes : Cardium serratum Lmk . =
Laevicardium erassum Gmelin (fide Fischer-Piette
1977). "1837 Deshayes: Cardium serratum Lmk. =
Laevieardium erassum Gmelin (fide Fischer-Piette
1977). --1839 Anton: Cardium serra tum Lam. (non
Linne). --1845 Reeve: Cardium serratum Bruguiere =
Cardium sulcatum Lamarck. "1846 Loven: Cardium
serratum Lamk., Desh., Thorpe, Turt. (non L., nee
Chernn., nee Brug.) = Cardium norvegieum Spengl.
"1848 Requien: Cardium serra/um Brug. = Cardium
oblongum Chernn. --1850 Jay: Cardium serratum Brug.
= Cardium sulea/um . --1851 Petit de la Saussaye:
Cardium serra/um Bruguiere (European record). --1856
Wood & Hanley: C. serra/um Lam. (not Lin.). --1860
Mace: Cardium serratum Bruguiere (European record).
--1867 Weinkauff: Cardium serra tum Lamarck (non
Linne) = C. norvegieum Spengler. --1869 Romer:
Cardium serra/um Bruguiere, Lamarck = Cardium
norvegieum Spengler. --1888 Kobelt: Cardium
serratum Lam . = Cardium norvegieum Spengler.
--1850 Jay: Cardium serra/um Lam. = Cardium
pennantii. --1890 Paetel: Cardium serratum Brug. =
norwegieum Spglr. "1912 Dautzenberg & Fischer:
Cardium serra/um Bruguiere 1789 (non Linne) =
Cardium (Laevicardium) norvegieum Spengler 1799.
--1942 Lamy: Cardium serratum Lmk. = Laevicardium
erassum Gmelin (fide Fischer-Piette 1977). --1945 ZR:
Cardium serratum Lk (non L.) [Ref. Lamy 1945]. CHARACTERS AND OCCURRENCE - DESCRJPTION.
--1789 Bruguiere: Cardium testa oblonga ovala, sulcis
longitudinalibus obsoletis, antice laevis; Nob.
REMARKS. The suggested synonyms are all European
species. They do not occur in the Caribbean (Antilles)
or the Atlantic Ocean of North America (Carolina
coast).
serratum Chemnitz, 1782. Non binominal. - Not
762
Tropical Marine Mollusc Programme (TMMP)
serraturn
available. » Regarding Cardium serratum: RECORDS.
--1788 Schroter: Cardium serratum [Reference to
Chemnitz (1782) figs. 193 . 194?]. --1864 Krebs :
Cardium serratum Ch., non L. = Cardium citrinum Ch.
serraturn Dall & Simpson 190 I (variety).
Infrasubspecific. Cardiidae. REcORDs. --190 I ZR: Cardium serratum n. var. described in Dall & Simpson 1901.
serraturn Linnaeus, 1758/ Gmelin, 1791. [Figs. Part 1
p. 75 No. 185-186, p. 91. p. 100, 124, 165, 179. Part 3
p. 879, 893, 894, 896]. - Laevicardium serra tum
(Linne, 1758). OTHER JUDGEMENTS: Cardium laevigatum
Linne sensu Clench & Smith 1944. (See overview of
ID). - Laevicardium oblongum (Gmelin, 1791). Laevicardium crassum (Gmelin, 1791). » Regarding
Cardium serratum: Cardiidae. Miocene and Recent.
T YPE LOCALITY. Matanzas Bay, Cuba. Unknown
(McLean 1939). TYPE MATERIAL. Vidal (1999): The
syntypes of C. serratum in Linne's personal collection
at The Linnean Society in London, encompass three
single valves that belong undoubtedly to the American
Atlantic Laevicardium laevigatum (Linne) sensu
Clench & Smith, 1944. The largest, a left valve [37.7 x
31.5 x (24.0»), with Linne's hand marks 73 and 89 (his
list numbers in 1758 and 1767) still discernible, has
been selected by Vidal (1999, p.327) as lectotype of
Cardium serratum Linne. RECORDS. --1758 Linne, p.
680, No. 73 : Cardium serratum. --1764 Linnaeus (Mus.
Lud. Ulr. Reg.) No. 45: Cardium serratum. --1767
Linne, p. 1123, sp. 89: Cardium serratum. --1782
Chernnitz. ID by Grnelin 1791: Cardium serratum t.
18. f. 185. 186. --1786 Schroter, No . XVII: Cardium
serra tum Linn. XII. pag. 1123. Spec. 89. "1791 Gmelin
No. 19: Cardium serratum. "1799 Spengler, No. 27:
Cardium serra tum Linnei Sp. 89. pag. 1123. "1817
Dillwyn: Cardium serratum Linnaeus. --1819 Lamarck,
p. ll , No. 25: Cardium serra tum Linne. --1819 Lamarck
(pro parte) : Cardium serra/um L. = Laevicardium
crassum Gmelin (fide Fischer-Piette 1977). --1836
Bosc: Cardium serratum. --1840 Sowerby, catalogue
p. 5, No. 66: Cardium sen'atum Linn. = Laevicardium
crassum Gmelin (fide Fischer-Piette 1977). --1843
Hanley: Cardium serra tum L. = Laevicardium
brasilianum Lamarck, 1819 (fide Fischer-Piette 1977).
"1844 Potiez & Michaud: Cardium serra tum Linn.
Coasts of Eng land and France. --1844 Thorpe: Cardium
serratum Lin. 1123. "1844 Reeve PI. 1 fig. I Cardium
serratum Linnaeus (not ofGmelin), Syst. Nat. 12th edit.
No . 89 . p. 1123 . "1859 Sowerby: C. serra tum =
Cardium norvegicum, Spengler. "1860 Stimpson:
Liocardium serratum Lin. --1868 Col beau : Cardium
serratum = Laevicardium norwegicum Spgl. --1884
Grasse!: Laevicardium serratum Linne. --1890 Paetel:
Laevicardium serratum L. --1902 Dall:Cardium
(Laevicardium) serratum Linnaeus, 1758. --1902 Dall:
Cardium serra tum Linnaeus, 1758 (not of Pennant,
1778). --1902 Dall & Simpson : Cardium
(Laevicardium) serra tum L. = La evicardium
brasilianum Lamarck, 1819 (fide Fischer-Piette 1977).
--1902 Sherborn, p. 893: Cardium serratum Linnaeus,
1758. --1921 Maury: Cardium (Laevicardium)
serratum Linnaeus, 1758 [Miocene to Recent] . --1925
Woodring, p. 145 , PI. 19, Figs. 14-16 : Cardium
(Laevicardium) sen'atum Linne, 1758 . Bowden,
Jamaica. --1926 Pei Ie: Liocardium serratum, L. = ? P
laevigatum , L., Dall . 1929 Tomlin: Laevicardium
serra tum (L.). --1932 Prashad : Cardium serratum
Gmelin, 1790 = Cardium biradiatum Bruguiere, 1789.
--1934 Johnson: Laevicardium serratum Linne 1758
(laevigatum Linne). --1936 Pearse: Trigoniocardia
serrata (L.) = Laevicardium brasilianum Lamarck,
1819 (fide Fischer-Piette 1977). "1939 McLean:
Laevicardium serratum serra tum (Linne, 1758). --1939
ZR : Laevicardium serratum serra tum fig. [Ref.
McLean 1939). --1942 Harry: Cardium serratum Linne
= Cardium muricatum Linne. --1977 Fischer-Piette:
Liocardium serratum Linne in Dall 1889 =
Laevicardium brasilianum Lamarck, 1819. --1977
Wilson & Stevenson: Cardium serratum Gmelin, 1791;
junior synonym of Laevicardium biradiatum
(Bruguiere, 1789). "1992 Lamprell & Whitehead:
Cardium serra tum Gmelin, 1790 = Laevicardium
biradiatum (Bruguiere, 1789). --1999 Vidal: Cardium
serra tum Linne, 1758 = Laevicardium laevigatum
(Linne) sensu Clench & Smith, 1944: 22. "1999
Knudsen & Hylleberg: Spengler, 1799 No. 27: Cardium
serratum in Spengler (without locality) = Cardium
laeviga/um Lin. Chemnitz Tom. VI. Tab. 18 Fig. 185186 ... 2001 Vidal, p. 407: Laevicardium serratum. CHARACTERS AND OCCURRENCE - OF COLLECTIONS. ZMUC from West Indies, 4 specimens; ZMUC
4 shells: laevigatum Lam. = serra/um L., Dansk
Vestindien, Steenberg coil. ZMUC label: Type, but
together with biradiatum no locality. ZMUC 2
specimens. Florida Keys. - 6 specimens. from West
Indies, - 4 112 specimens from Antilles, - I specimen
= laevigatum Lamarck (no. 26) & Chemnitz 6. fig. 189:
same species as labeled laevigatum from the Danish
West Indies and Tortola (Moellers samling); - ZMUC
2 specimens in Spengler colI. DESCRIPTIONS. --1799
Spengler: Testa obovata laevi striis obsoletis, margine
interiore serrato. [My translation: Shell inverted eggshaped, smooth, obsolete ribs, inner margin serrated ].
--1799 Spengler translated by Knudsen & Hylleberg
1999: Cardium serratum in Chemnitz Tom. VI. Tab.
18. fig . 185. 186. (is also called laevigatum and thus
confounded with serrato). Since Linne in his Syst. Nat.,
but notably in the Queen's Museum, described this
Cardium in great detail, it needs no further description.
Page 44 : Margins finely serrated from anterior to
posterior ends (contrary to C. laevigatum) . "1844
Reeve: shell ovate, rather gibbous towards the
umbones, smooth, shining, anteriorly rather obsoletely
striated; whitish yellow, posterior side bright citron
yellow or pinkish purple, palely stained with pinkishwaved spots around the umbones, interior yellowish
under the umbones; margins very minutely serrated
within. --1902 Dall & Simpson: Shell subsolid, suboval
or subquadrate, the posterior edge being less curved
than the anterior; the low, rounded, posterior ridge being
slightly produced at the base of the shell; surface nearly
smooth, shining and polished, with faint vestiges of
ribs, which appear on the border of the shell which
scarcely gapes and has delicate serrations within;
laterals high and strong; cardinals small; interior of the
shell polished; in a fresh state the surface is usually
covered with a thin, brownish epidermis; the shell is
beautifully polished; whitish, straw-coloured, lemonyellow, purplish or reddish, and it is slightly clouded
Phuket Marine Biological Center Special Publication vol. 30 (2004)
763
serratum - serratum multilineatum
with brown near the beaks; within it has much the same
range of colour, and it is one of the most delicate and
lovely shells of the Antilles. ··1939 McLean: Obsolete
radial striae. ··200 I Vidal: tentacles numerous. No eyes
on tentacles. DIMENSIONS. ··1939 McLean: 38 x 45 x
30 mm. REMARKS. ··1799 Spengler: stated that the
species illustrated by Chemnitz (1782 pI. 18, figs . 185186) as Cardium laevigatum has been confused with
Cardium serratum. The reason is that Chemnitz refers
to Linne (1769, p. 1123, No. 88), but species No. 88 is
Cardium laevigatum (a printer's error). Spengler (1799)
mentions the description of Cardium serra/um by Linne
(1764) as being very clear and in no need of further
comments . •• I 844 Reeve: An odd valve of the Cardium
serratum is still preserved in the Linnaean Society of
London marked "86" in Linnaeus' handwriting,
referring to the 12th edition of the ' Syst. Nat.'
Regarding erroneous ID of Cardium laevigatum by
Gmelin, Lamarck, Sowerby. The opinions of Lamarck
and Sowerby were induced probably by Gmelin (an
error), followed by Bruguiere, in the Encyclopedie
Methodique. ··1867 Weinkauff: mentioned that Linne
described 3 similar species: C.flavum, C. laevigatum ,
and C. serratum. Chemnitz illustrated the 3 species but
confused the figure numbers with the Linnean
descriptions. He stated that the figures 185 and 186
showed C. serratum but it was actually C. laevigatum.
The figure 189 illustrated C. laevigatum but actually
referred to C. serratum. Weinkauff maintained that figs.
185 and 186 did not illustrate either of the species.
Hanley (1855) discovered that C. laevigatum Linne =
C. papyraceum Chemnitz. C. serra tum Linne = C.
laevigatum Lamarck (West Indian species) (discussion
p. 147-148). HABITAT. ··1844 Reeve: found in sandy
mud at the depth of from 6 to 10 fathoms. DISTRIBUTION.
··1758 Linnaeus: Mediterranean . •• I 790 Gmelin: C.
serratum Mediterranean & Indian Ocean . •• I 844
Reeve: Islands of Nevis and St. Vincent, West Indies.
"1884 Grasset: Guadeloupe. "1939 McLean: C.
serra tum (serratum) North Carolina to Brazil.
serratum of authors. - Laevicardium la eviga tum
(Linnaeus). Cardiidae . Recent. RECORDS ... I 886
Locard: Cardium serra tum (non Linne) = Cardium
norvegicum Spengler, 1792 sic. ··1951 . McLean :
Cardium serralUm of authors non Linnaeus =
Laevicardium laevigatum (Linnaeus).
serratum of Reeve (1843). - Identical with Laevicardium
laevigatum (Linnaeus, 1758). » Regarding Cardium
serratum: Cardiidae. Recent. RECORDS. ··1969 Work:
Cardium serra tum Reeve , 1843 = Laevicardium
laevigatum (Linnaeus, 1758). - CHARACTERS AND
OCCURRENCE - DISTRIBUTIO N. ··1844 Reeve:
Mediterranean.
serratum ofRequien (1848). - Misidentification of Cardium norvegicum Spengler = Laevicardium crassum
(Gmelin, 1791). Cardiidae. Recent. RECORDS. ··1900
Locard & Caziot: Cardium serra/um Requien (1848)
p. 26 = Cardium norvegicum Spengler, 1790 sic.
serratum of Schroter (1788). - Identical with Cardium
linea/um Gmelin. Cardiidae. Recent. RECORDS. ··1817
Dillwyn: Cardium serratum Schroter = Cardium
linea tum Gmelin.
serratum of Thorpe (1844). - Identical with
Laevicardium laeviga/um (Linnaeus, 1758). Cardiidae.
Recent. RECORDS. ··1844 Thorpe: Cardium serra/um
D. p. 124. = Cardium levigatum Lin. 1123.
serratum of Turton (1822). - Identical with Laevicardium
crassum (Gmelin, 1791). Cardiidae. Recent. RECORDS.
··1917 Hidalgo: Cardium serratum Turton = Cardium
norvegicum Spengler. ··1977 Fischer-Piette: Cardium
serra/um L. in Turton 1822 = Laevicardium crassum
Gmelin.
serratum of Weinkauff (1862). - Identical with
Laevicardium norvegicum (Spengler) = Laevicardium
crassum (Gmelin, 1791). Cardiidae. Recent. RECORDS.
··1880 Stossich: Cardium serratum in Weinkauff. 1862
= Laevicardium norvegicum (Spengler).
serratum brasilianum Lamarck, 1819 (variety). Upgraded. Changed combination: Laevicardium
serra tum brasilianum (Lamarck, 1819). Cardiidae.
Recent. TYPE LOCALITY . Rio de Janeiro, Brazil. RECORDS.
··1908 Cary & Spaulding: Liocardium serratum var.
brasilianum. ··1939 McLean: Laevicardium serratum
brasilianum (Lamarck, 1819). ··1939 ZR:
Laevicardium serra/um brasilianum [Ref. McLean
1939]. - CHARACTERS AND OCCURRENCE DESCRIPTIONS. ··1939 McLean: obsolete radial striae.
DISTRIBUTION. Southern West Indies to Brazil.
serratum laevigatum Lamarck , 1819 (variety). Upgraded. Changed combination : Laevicardium
serratum (Linnaeus, 1758). Cardiidae. Recent. TYPE
LOCALITY. Rio de Janeiro, Brazil. RECORDS. ··1819
Lamarck, p. II, No. 26: Cardium laevigatum (quelques
variete dans la forme et les couleurs). ··1902 Dall &
Simpson, p. 489: Cardium (Laevicardium) serra/um
var. laevigatum Lamarck. ··1926 Weisbord: Cardium
(Laevicardium) serra/um var. laeviga/um Lamarck.
··1945 Smith, Maxwell, p. 50: Laevicardium serratum
laevigatum Lamarck. ··1970 Vilas & Vilas p. 38:
Laevicardium serra/um var. laevigatum. CHARACTERS AND OCCURRENCE - DESCRIPTIONS. Shell rather large, subsolid, moderately inflated,
subquadrate, usually, though not always, faintly ribbed,
ribs numerous when developed, strongest around the
outer part of the shell; there is an area at the anterior
and posterior parts of the shell which is generally
smoother than the rest of it, which is outlined by a
shallow groove, forming a sort of lunule and
escutcheon; laterals rather strong, especially the anterior
ones; edges of the shell serrated within. Colour
generally whitish, cream or pale buff, with a faint
subconcentric c10udings of brown (Dall & Simpson
1902). DIMENSIONS. Length 50, height 63, diameter 40
mm (Dall & Simpson , 1902). REMARKS . Generally
larger, more quadrate, duller coloured, and more painted
with brownish, nebulous bands than typical serra tum,
yet there are intermediate specimens which can be
assigned to one form about as well as the other. This
form is one of the characteristic species of West Florida
(Dall & Simpson 1902). DISTRIBUTION. West Florida,
Playa de Ponce, Porto Rico (Dall & Simpson 1902).
serratum multilineatum Dall & Simpson 1902 (variety).
- Synonym of Laevicardium pristiS (Valenciennes,
1827). OTHER JUDGEMENTS: Cardium leucostomum Born.
» Regarding Laevicardium serratum multilineatum:
Cardiidae. Pleistocene and Recent. TYPE LOCALITY.
Mayaguez Harbor, Puerto Rico. RECORDS. ··1902 Dall
& Simpson, p. 489-490: Cardium serra/um var.
764
Tropical Marine Mollusc Programme (TMMP)
serratum sybariticum - Serripedini
multilineatum, n. var. 001939 McLean: Laevicardium
serratum multilineatum (Dall & Simpson 1902).001939
ZR: Laevicardium serratum multilineatum fig. [Ref.
McLean 1939]. 001953 Olsson & Harbison: Cardium
serratum mllitilineatum Dall & Simpson, 190 I =
Laevicardium laevigatum laevigatum (Linne, 1758).
001977 Fischer-Piette: Laevicardium serratum multilineatum Dall & Simpson = Cardium leucostomum
Born. - CHARACTERS AND OCCURRENCE DESCRIPTIONS. 001902 DaB & Simpson: shell more
nearly round than the type. The smoother anterior and
posterior areas are distinctly marked out and are tawny
or buff coloured; the rest of the sheH is dirty straw
coloured, and the narrow spaces between the scarcely
developed ribs are pale brown; the beaks are purple,
clouded. The interior is much the colour of the yolk of
an egg; with purple c10udings at the cavities of the
beaks. 001939 McLean: obsolete radial striae. DIMENSIONS. 001939 McLean: 40 x 45 x 32 mm. 001902 Dall
& Simpson: length 40, height 45, diameter 32 mm.
DISTRIBUTION. 001939 McLean: Santo Domingo, Puerto
Rico.
serratum sybariticum Dall, 1886 (variety). - Upgraded.
Laevicardium serra tum sybariticum (Da\l, 1886).
Cardiidae. Oligocene and Recent. TYPE LOCALITY.
Barbados, West Indies. RECORDS. 001886 Dall, p. 271:
Cardium serratllm var. sybariticlIm. 001902 Dall:
Cardium serratum var. sybariticllm Dall, 1886. 001902
Dall & Simpson: Cardium (Laevicardillm) serratum
var. ?sybariticum. 001902 Dall & Simpson: Cardium
serratum var. sybariticum Dall, 1886. 001939 McLean:
Laevicardium serraillm sybariticum (Dall, 1886).
.01939 ZR: Laevicardium sen'alum sybariticum [Ref.
McLean 1939] . - CHARACTERS AND OCCURRENCE - DESCRIPTIONS. Shell much smaller and more
delicate than the type, more compressed, pale faintly
flecked with pink or purple; beaks often deep pink (Dall
& Simpson 1902); obsolete radial striae (McLean
1939). DIMENSIONS. 11.5 x 13.5 x 8 mrn (McLean 1939);
length II mm, height 14 mm, diameter 7 mm (Dall &
Simpson 1902). DISTRIBUTION. West of Florida to
Barbados.
serresense Popov & Nevesskaya, 2000. - Lymnocardillm
serresense Popov & Nevesskaya, 2000. Cardiidae. Late
Miocene . RECORDS. 002000 Popov & Nevesskaya,
downloaded abstract: Lymnocardillm (Euxinicardium)
serresense n. sp.
serricostatum Kira, 1955. Synonym of Acrosterigma
elongatllm (Bruguiere, 1789). » Regarding
Vasticardium serricostatum: Cardiidae. Recent. TYPE
LOCALITY & REPOSITORY. See okinawense Kuroda, 1960.
DISTRIBUTION . Japan. RECORDS. 001954 Kira :
Vasticardillm serricostatum Kuroda, MS. 001955 Kira:
Vasticardium serricostatum Kuroda, MS.
serricostatum Melvill & Standen, 1899. - Synonym of
Vasticardium elongatum (Bruguiere, 1789). OTHER
JUDGEMENTS: Cardium flavllm Linnaeus, 1758 . Trachycardium luteomarginata Voskuil & Onverwagt,
1991. » Regarding Cardium (Trachycardium)
serricostatum: Cardiidae. Recent. TYPE LOCALITY. Torres
Strait. TYPE REPOSITORY. BMNH reg. nr. 1899-2-2310 (Vidal 1993). RECORDS. 001899 Melvill & Standen,
p. 191, pI. 11, fig. 20: Cardium (Trachycardium)
serricostatum. 001899 ZR: Cardium (Trachycardium)
serricostatum Melvill & Standen, 1899 n. sp. 001909
Melvill: Cardium (Trachycardium) serricostatum, M.
& S. 001951 Habe: Vasticardium serricostatum. 001952
Kuroda & Habe: Trachycardium serricostatum. 001955
Kira: Vasticardium serricostatum . 001977 FischerPiette: Cardium serricostatum Melvill & Standen =
Laevicardium flavum Linne. 001990 Abbott & Dance:
Trachycardium serricostatum . 001993 Vidal p. 44:
Cardium serricostatum Melvill & Standen, 1899
probably = Acrosterigma elongatum (Bruguiere, 1789).
001993 Vidal : Trachycardium serricostatum in Abbott
& Dance, 1982 = ? Trachycardium luteomarginata
Voskuil & Onverwagt, 1991. 001999 Vidal: Cardium
serricostatum Melvill & Standen, 1899 = Vasticardium
elongatum (Bruguiere 1789). - CHARACTERS AND
OCCURRENCE - CHARACTERS of diagnostic value: the
deeply excavate interstices and squarely set serrated
ribs are noticeable (MelvillI909). DISTRIBUTION. North
Indian Ocean, Japan (Hylleberg 1994); elongatum West
Indies (Reeve J844); Philippines, Australia. Murray
Island, Torres Straits (Melvill 1909).
serricostatum Melvill & Standen, 1899 (variety). Synonym of Vasticardium elongatum (Bruguiere,
1789). RECORDS. 001952 Kuroda & Habe: serricostatum
Melvill & Standen var. = Trachycardium serricostatum
Melvill & Standen var. 001959 Kira : Vasticardium var.
serricoslatum Melvill & Standen. 001962 Kira, p. 139,
pI. 55, No. I J: Vasticardium var. serricoslalum. 001993
Vidal: Vaslicardillm var. serricoslatum (Melvill &
Standen, J899) = Acroslerigma okinawense = small
Acrosterigma elongatum.
serricostatum okinawense Kuroda, MS. - Synonym of
Vasticardium elongalum (Bruguiere, 1789). »
Regarding Vasticardium serricostatum okinawense:
Cardiidae. Recent. RECORDS. 001951 Habe:
Vaslicardium serricostatum okinawense Kuroda MS.
001960 Kuroda: Trachycardium (A crosterigma)
serricostalum okinawense Kuroda.
serrifera Lamarck. [Figs. Part I p. 152]. - Tridacnidae.
RECORDS. 001845 Chenu, pI. 8 f. 4: Tridacna serrifera
Lamarck. 001868 ZR: Tridacna serrifera [Ref. KUster
1868]. 001932 ZR: Tridacna serrifera [Ref. Lamy
1932]. 001947 ZR: Tridacna serrifere sic Lamarck,
1819. 00 1956 ZR: Tridacna serrifera spawning habits
[Ref. Wada 1956].
*serrigerum. - Homonymy to be noticed. Cardium
serrigerum from 2 sources: Lamarck, 1819. - Basterot
(?year).
serrigerum Basterot. - Combination to be revised :
"Cardium" serrigerum Basterot. Cardiidae. Miocene.
RECORDS . 001844 Potiez & Michaud: Cardillm
serrigerum Bast. Fossil of France. 001999 Jacques Vidal
(pers. com.) MNHN, Paris : Fossils from the
Paleontological Collection. As Cardium serrigerum
Basterot. Burdigalien. France (Gironde), S I 0999.
serrigerum Lamarck, 1819. - Combination to be revised :
"Cardium" serrigerum Lamk. Cardiidae. Fossil.
RECORDS. 001819 Lamarck: Cardium serrigerum Lamk.
001922-32 Sherborn, p. 5914: Cardium serrigerum
Lamarck, 1819.
Serripedini Kafanov, 1975 (tribus). - Cardiidae.
RECORDS. 001975 Kafanov : Serripedini. 001975 ZR:
Serripedini Tribe nov. of Cardiidae. Type genus
Serripes Gould 1841 , p. 146. Ref. to Kafanov.
Phukel Marine Biological Center Special Publication vol. 30 (2004)
765
Serripes - setosum
Serripes Gould, 1841 (genus). - Cardiidae. Fossil and
Recent. Type species: Gould, 1841 designated Cardium
groenlandicum Bruguiere, 1789 type species of the
genus Serripes Beck. RECORDS. "1847 Gray, No. 559:
Serripes, Beck MSS. [synonym of Aphrodite, Lea; 183?
not Linn.]. "1924 Oldroyd: Genus Serripes. "1934
Johnson: Serripes Beck (Aphrodite Lea 1834, not
Hubner 1816). "1975 ZR: Serripes Gould, 1841
designated type genus for new tribe Serripedini, p. 146.
Ref. to Kafanov. "1977 Popov: Serripes Beck in Gould,
1841. Genotype: Cardium groenlandicum. "1980
Vokes: Serripes Gould, 1841 in subfamily
Laevicardiinae. "1980 ZR: Serripes Gould, 1841. Key
to species & subspecies. ZR Ref. to Kafanov. "1981
ZR: Serripes Gould, 1841. Western Pacific. temperate
zone. Ref. to Skarlato, p. 347. "2000 Coan et af.: Genus
Serripes Gould, 1841. ··200 I Vidal, p. 406: SerripesCHARACTERS AND OCCURRENCE - CHARACTERS
of diagnostic value. "1994 Hylleberg: sculpture almost
obsolete; shell medium sized to large, inequilateral;
cardinal teeth weak in adults or absent, laterals also
evanescent. ··200 I Vidal: Siphons relatively long, close
to each other, often fused at their bases. No eyes on
tentacles. REMARKS. 1876. Meek: mentioned Serripes
as allied to Laevicardium.
Serripes Gould, 1841 (section & subgenus). - Cardiidae.
Fossil and Recent. RECORDS. "1858 Adams & Adams:
Subgenus Serripes Beck. "1869 ZR: subgenus Serripes
(Beck), type Cardium groenlandicum (Chemn.) [Ref.
Romer 1869]. "1887 Fischer: Subgenus Serripes Beck,
fide Gould 1841 of genus Cardium Linne, 1758. "1890
Paetel: 6. Sect. Serripes Bk. of Cardium. "1912
Dautzenberg & Fischer: Subgenus Serripes (Beck)
Gould, 1841 of genus Cardium, Linne 1758.
serrula Cox, 1927. - Identical with Cardium burnupi
Sowerby, 1897. » Regarding Cardium serrula:
Cardiidae. Miocene. RECORDS. "1927 ZR: Cardium
(Acanthocardia) serrula Cox, 1927 sp. n. Miocene
Pemba Isd. "2000 Vidal, p. 448: Cardium
(Acanthocardia) serrula Cox, 1927, a Neogene and
Quaternary fossil from Zanzibar. Synonym of Cardium
burnupi Sowerby, 1897.
serrulatum Deshayes, 1854. - Changed combination:
Europicardium serrulatum (Deshayes, 1855).
Cardiidae. Recent. TYPE LOCALITY. Guinea. TYPE
REPOSITORY. Possible holotype BMNH. REMARKS.
Referred to Vepricardium in Keen (1980). Syn. kobelti
v. Maltzan, 1885 (Vidal, pers. com.). DISTRIBUTION. West
Africa (Vidal 1999). RECORDS. "1855 Deshayes:
Cardium serrulatum. "1890 Paetel: Cardium
serrulatum Dsh. "1977 Fischer-Piette: Laevicardium
(Trachycardium) serrulatum Deshayes. "1980 Ruhoff:
Cardium serrulatum Deshayes, 1854; p. 330 ... 1999
Vidal: Cardium serrulatum Deshayes, 1855. "1999
Vidal: Europicardium serrulatum (Deshayes, 1855).
sertunayana Slodkewitsch, 1938. - Changed
combination: Fulvia sertunayana (Slodkewitsch,
1938). Cardiidae. Miocene. TYPE LOCALITY. Kamtchatka.
REcoRDS. "1945 ZR: Papyridea sertunayana 'Kogan'
sic Siodkewitsch, 1938 sp. nov. Upper Miocene E. coast
Kamchatka. "1973 Keen: Fulvia sertunayana
(Slodkewitsch, 1938) [Papyridea]. Miocene,
Kamtchatka.
sertunayana Siodkewitsch, 1938. - Profulvia
sertunayana (Slodkewitsch, 1938). Cardiidae. Fossil.
RECORDS. "1976 ZR: Papyridea sertunayana
Slodkewitsch, 1938 Referred to Profulvia. Ref. to
Kafanov, p. 482 [orig. p. III].
serum Hutton, 1873. - Changed combination:
Nemocardium serum (Hutton, 1873). Cardiidae. Fossil.
RECORDS. "1897 Harris: Protocardium serum Hutton.
1873. "1944 ZR: Protocardium serum Hutton Catal.
Tert. Moll. 1873, p. 23 [Ref. Marwick 1944]. "1944
ZR: Varicardium serum (Hutton, 1873) genotype of
Varicardium Marwick, 1944. "1965 ZR: Varicardium
serum revised, Tertiary, New Zealand [Ref. Boreham
1965]. "1977 Popov: Nemocardium (Varicardium)
serum (Hutton, 1873).
Serupes (error). - Printer's mistake for Serripes in
Gould, 1841 (Vokes 1980).
sessa Nelli, 1915. - New combination: Lymnocardium
sessa Nelli, 1915. Lymnocardiidae. Ponti an. RECORDS.
"1915 ZR: Limnocardium sessa Nelli, 1915 sp. n.
Pontian, Greece.
setigerum Dunker. - Combination to be revised:
"Cardium" setigerum Dkr. Cardiidae. REcORDs. "1890
Paetel: Cardium setigerum Dkr.
setosum Linne. (error). - Printer's mistake for retusum
in Shikama 1964.
setosum Redfield, 1848. [Figs. Part I p. 88. Part 3 p.
902]. - Changed combination: Plagiocardium setosum
(Redfield, 1848). Cardiidae. Recent. TYPE LOCALITY.
Seas of China. RECORDS. "1848 Redfield: Cardium
setosum. "1858 Adams & Adams, p. 456: Cardium
(Cerastoderma) setosum, Redf. "1884 Grasset:
Cardium setosum. Redf. "1885 Smith: Cardium
(Acanthocardia) setosum Redfield. "1890 Paetel:
Cardium setosum Rdfld. ··1909 Lynge: Cardium
(Acanthocardia) setosum. "1922-32 Sherborn, p. 5924:
Cardium setosum Redfield, 1846. "1952 Kuroda &
Habe: Papyridea setosa (Redfield). "1960 Kuroda:
Maoricardium setosum (Redfield). "1964 Shikama:
Papyridea (Maoricardium) setosa Redfield. "1967 ZR:
Cardium setosum, effect of myotropic substances on
the heart [Ref. Kratzing & Ladd 1967]. "1977 FischerPiette: Laevicardium (Vepricardium) setosum Redfield.
··1977 Wilson & Stevenson: Plagiocardium
(Maoricardium) setosum (Redfield, 1848). "1977
Popov: Plagiocardium (Plagiocardium) setosum
(Redfield). "1988 Drivas & Jay: Plagiocardium
(Maoricardium) setosum (Redfield, 1848). "1990
Abbott & Dance: Plagiocardium setosum. "1991 Rao,
Rao & Maitra: Cardium setosum in Satyamurti (1956)
= Acanthocardia lata (Born, 1778). "1991 Voskuil &
Onverwagt: Maoricardium setosum (Redfield, 1846).
"1992 Lamprell & Whitehead: Plagiocardium
(Maoricardium) setosum. ··1994 Chung-Hung & HsiJen: Plagiocardium (Maoricardium) setosum
(Redfield). "1994 Schneider: Maoricardium setosum
(Redfield, 1846). "1995 Chung-Hung & Hsi-Jen:
Maoricardium setosum (Redfield). "1997 Nguyen et
al.: Vepricardium setosum (Redfield). "1999
Aungtonya et al., PMBC-SP 19(2): 378: Vepricardium
setosum (Redfield, 1846). "1999 Hylleberg, p. 40 I:
Maoricardium setosum (Redfield, 1846). ID J. Vidal.
"2000 Hylleberg, p. 532: Plagiocardium setosum
(Redfield, 1848). "2003 Hylleberg & Kilburn, p. 184:
Plate 2, fig. 8: Plagiocardium setosum (Redfield, 1848).
766
Tropical Marine Mollusc Programme (TMMP)
setosum - shepstonense
- CHARACTERS AND OCCURRENCE - OF COLLECTIONS. ··ZMUC: I specimen from China. DESCRIPTIONS.
··1848 Redfield: Shell a little obliquely rounded ovate,
somewhat tumid ; anterior side shortest , beaks
prominent, inflected and nearly in contact; in front of
them is a rather narrow ovate depression. About 40
flattened ribs, twice as wide as the interstices. rib edges
very minutely denticulate, small semi-tubular spines
or scales upon the central line of each rib, spines
terminate in blackish bristles, more developed on the
posterior margin than elsewhere. The ribs which cross
the anterior lunular depression are not bristly, but
tubercular. Colour white, periostracum dirty yellow,
which becomes thicker and much wrinkled towards the
margin, especially upon the posterior end, where the
colour is nearly black. Interior white. ··1988 Drivas &
Jay: interior shell with a large brown blotch.
DIM ENSIONS. ··1848 Redfield : length: 2.75 inches ,
height 2.75 inches, breadth 2.1 inches. ··1988 Drivas
& Jay: max. ant.-post. 47.5 mm, umbo-ventr. 34.2 mm.
REMARKS. ··2003 Hylleberg & Kilburn: maintained that
several synonyms of P setosum (Redfield, 1848) have
been proposed. Maoricardium latum is a synonym of
Maoricardium setosum according to J. Vidal.
DISTRIBUTION. ··1884 Grasset: China. ··1994 Hylleberg:
South West Pacific, Australia. ··1988 Drivas & Jay:
Reunion, western Indian Ocean at 12 m depth. ··1999
Aungtonya et al.: Thailand, Andaman Sea. ··2000
Hylleberg: Vietnam and Cambodia. ··2003 Hylleberg
& Kilburn: Vietnam. ··ZMUC: China.
setosum Tryon , 1872. - Synonym of Cardium
quadragenarium Conrad, 1837. » Regarding Cardium
setosum: Cardiidae. Recent. RECORDS. ··1902 Dall:
Cardium setosum Tryon, 1872 (not Redfield, 1846) =
Cardium quadragenarium Conrad, 1837.
sellrati Vidal, 1999. [Figs. Part 3 p. 880]. - Acrasterigma
seurati Vidal, 1999. Cardiidae. Recent. TYPE LOCALITY.
Holotype, Tuamotu archipelago, south central-Pacific.
TYPE REPOSITORY. Holotype, Museum national d 'Histoire
naturelle, Paris. TYPE MATERlAL. All the existing material,
nineteen single valves, collected by L. G. Seurat,
probably on beaches, in the Marutea du Sud Atoll,
Tuamotu Archipelago (1902-1905), is selected as type
series, stored in MNHN . Holotype: A right valve.
Paratypes I to 18: Ten right valves and eight left valves.
RECORDS. ··1999 Vidal, p. 312, Table 28, Figs. 14 E-F:
Acrosterigma seurati n. sp. - CHARACTERS AND
OCCURRENCE - DESCRIPTION. ··1999 Vidal: Shell
almost perfectly ovoid and equilateral (anterior dorsal
margin sometimes very slightly raised), rarely slightly
truncated posteriorly, weakly elongated and rather
globose. Lunule narrow, almost equal on both valves,
and poorly differentiated, with its umbonal margins
slightly raised. External colour almost entirely white,
except for a few pink spots or small splashes and part
of posterior slope sometimes yellow; interior white,
sometimes with very pale yellow umbonal rays. Hinge
nearly symmetrical and variably angled. Mean rib
number 62 .8 ± 3.6, range 56-68. On posterior slope,
interstices very narrow and both parts of ribs about
equivalent in width; scales in·egular, tubercular. On
posterior part of median surface, ribs become more or
less flatly rounded, sometimes still bearing an axial
furrow; entire rib or wider posterior part can bear
elongated ridges, analogous to the scales of posterior
slope, but more numerous. On anterior part of median
part, ribs ornamented only with posterior crenulations;
interstices become wider and often very finely notched.
On anterior half of shell, ribs at first remain rounded
and retro-crenulated, and then become finely topridged ; on first anterior ribs, top ridges become
tubercular and irregular, tending to be arranged in
concentric alignments which can replace the
longitudinal ribbing. DIMENSIONS. ··1999 Vidal: Length
37.9, height 43.6, width (calculated) 30.0 mm.
REMARKS. A. seurati has several characters in common
with A. transcendens, but has a much larger, thicker,
heavier shells and slightly less elongated shell than A.
transcendens). It is also noticeably less colorful and
the hinge is a little less angled.
sevieriellse Hill , 1888. - Changed combination:
Pleuriocardia sevieriense (Hill , 1888). Cardiidae.
Cretaceous . RECORDS. ··1978 Scott: Cardium ?
sevieriense Hill, 1888. "1978 Scott: Pleuriocardia
(Pleuriocardia) sevieriense (Hill, 1888).
seJl!elli Prashad, 1932. [Figs . Part 3 p. 865]. - Changed
combination. Vasticardium sewelli (Prashad, 1932).
Cardiidae. Recent. TYPE LOCALITY. Indonesian Island.
RECORDS. ··1932 Prashad: Cardium (Trachycardium)
sewelli. ··1932 ZR: Cardium (Trachycardium) sewelli
Prashad, 1932 sp. n. ··1977 Fischer-Piette:
Laevicardium (Vasticardium) sewelli. ··1997 Poorten:
Acrosterigma sewelli (Prashad, 1932). ··1999 Vidal:
Vasticardium sewelli (Prashad , 1932).
CHARACTERS AND OCCURRENCE - DISTRIBUTION. Indonesia (Hylleberg 1994).
sexcostatllm Fr. Roemer, 1852. - Combination to be
revised: "Cardium " sexcostatum Fr. Roemer, 1852.
Cardiidae. Fossil. RECORDS. ··1980 Ruhoff: Cardium
sexcastatum Fr. Roemer, 1852; [1850-1852] vol. 3: 79,
pI. 12, fig . 13.
shabrawitensis Abbass, 1966 . - Protacardia
shabrawitensis Abbass, 1966. Cardiidae. Cretaceous.
RECORDS. ··1966 ZR: Protocardia shabrawitensis
Abbass, 1966 sp. nov. Cretaceous, Egypt.
Shanina Reed, 1915 (genus). - Cardiacea. Ordovician.
RECORDS. ··1915 ZR: Shanina Reed, 1915 gen. nov.
Ordovician, Shan States.
shehllenensis Ferugl io, 1936 . - Protocardia
shehuenensis Feruglio, 1936. Cardiidae. Cretaceous.
RECORD S. ··1936 ZR: Pr% cardia shehuenensis
Feruglio, 1936 sp. n. Cretaceous Patagonia.
shepstonense Tomlin, 1931. [Figs. Part 3 p. 918]. Changed combination : Afrocardium shepstonense
(Tomlin, 1931) . OTHER JUDGEMENTS : Afrocardium
richardi (Audouin, 1827). » Regarding Afrocardium
shepstonense: Cardiidae. Recent. Type species: Tomlin
(1931) designated Fragum (Afrocardium) shepstonense
type species of subgenus (Afrocardium) Tomlin, 1931.
Upgraded to generic level by authors. TYPE LOCALITY.
South Africa. RECORDS. ··1931 Tomlin: Fragum
(AJrocardium) shepstonense. ··1931 ZR: Fragum
(AJrocardium) shepstonense Tomlin, 1931 subg. et sp.
n. S. Africa. ··196 I Cotton: Fragum shepstonense
Tomlin 1931. Type species of Afrocardium Tomlin
1931 . ··1969 Moore et al.: Ctenocardia (Afrocardium)
shepstonellse (Tomlin, 1931). ··1977 Fischer- Piette:
Corculum (AJrocardium) shepstonense Tomlin. ··1977
Phuket Marine Biological Center Special Publication vol. 30 (2004)
767
shimotokuraense - silvacollina
Wilson & Stevenson: Fragum (Afrocardium)
shepstonense Tomlin, 1931. ··1982 Kilburn & Rippey:
Afrocardium shepstonense (Tomlin, 1932). ··2002
Schneider: Afrocardium shepstonense (Tomlin, 1931).
Holocene. ··2003 Hy\leberg & Kilburn, p. 176: C.
shepstonense Tomlin, 1931. Syn of Afrocardium
richardi (Audouin, 1827) according to 1. Vidal.
shimotokuraellse Akutsu, 1964. - Changed combination:
Clinocardium shimotokuraense (Akutsu, 1964).
Cardiidae. Miocene. RECORDS . ··1964 Akutsu:
Vasticardium shimotokuraense. ··1964 ZR:
Vasticardium shimotokuraensis Akutsu, 1964 sp. nov.
Miocene, Shimotokurazawa, Tochigi Pref. Japan.
··1973 Keen: Clinocardium shimotokuraense (Akutsu,
1964).
shimotokuraensis (mode). - See under shimotokuraense
Akutsu, 1964.
shinjiensis Yokoyama, 1923 . - Changed combination:
Clinocardium shinjiensis (Yokoyama, 1923). OTHER
JUDGEMENTS: Cardium nllttallii Conrad. - Cardium
californianum Conrad. » Regarding C. shinjiensis:
Cardiidae. Miocene. REMARKS. The fossil species is
most likely to be classified as Cerastoderma shinjiensis
(Yokoyama, 1923). RECORDS. ··1923 Yokohama, p. 7,
pI. 2, fig. 6 : Cardium shinjiensis, n. sp. ··1923 ZR:
Cardium shinjiense Yokohama, 1923 sp. n. Neogene
Japan. ··1926 Yokoyama: Cardium shinjiense
Yokoyama, 1923 = Cardium californianum Conrad.
··1926 Yokoyama: Cardium shinjiense Yokoyama,
1923 = Cardium nuttallii Conrad. ··1927 Yokoyama:
Cardium shinjense = Cardium nullallii Conr.; the shells
resemble Cardiumfastosum Yokoyama, 1927. ··1937
ZR : Cardium (Cerastoderma) shinjiense Yokoyama
[Ref. Kanehara 1937a]. ··1954 Keen: Clinocardium
shinjiense (Yokoyama, 1923). [Miocene]. ··1973 Keen:
Clinocardium shinjiense (Yokoyama, 1923) [Cardium].
Neogene. ··1974 ZR: Ciliatocardium shinjiense
(Yokoyama, 1923) Comb. nov. Ref. to Kafanov, p.
1469. ··1977 Popov: Clinocardium shinjense
(Yokoyama, 1923).
shiobaraellsis Noda 1962. - Changed combination:
Serripes shiobaraensis Noda, 1962. Cardiidae. Middle
Miocene. RECORDS. ··1962 ZR: Serripes shiobaraensis
Noda, 1962 sp. nov. Hoki River, Sekiya, Shiobaramachi, Shioya- gun, Tochigi Pref. Kanomatazawa
formation. ··1973 Keen: Serripes shiobaraensis Noda,
1962. ··1975 ZR: Yagudinella shiobaraensis (Noda,
1962) comb. nov. transferred from Serripes. Ref. to
Kafanov, p. 147. ··1978 Kafanov: Yagudine/la
shiobaraensis (Noda 1962) . ··1982 ZR: Serripes
shiobaraensis Noda, 1962. Diagnostic characters.
Combination revalidated. Transferred back from
Yagudinella. Middle Miocene. Russian SFSR. Ref. to
Kafanov et al. p. 44 . ··1982 ZR: Yagudinella
shiobaraensis (Noda, 1962). Referred to Serripes . Ref.
to Kafanov et al. p. 44.
shiobarense Yokoyama, 1926. - Changed combination:
Clinocardium shiobarense (Yokoyama, 1926).
Cardiidae. Miocene. RECORDS. ··1926 Yokoyama:
Cardium shiobarense, Yokoyama. ··1954. Keen:
Clinocardium shiobarense (Yokoyama), 1926. ··1973
Keen: Laevicardium shiobarense (Yokoyama, 1926).
shumardi Meek & Hayden, 1860. - Combination to be
revised: "Cardium " shumardi Meek & Hayden, 1860.
Cardiidae. Jurassic. RECORDS. ··1860 Meek & Hayden:
Cardium [Protocardum? sic] shumardi Meek &
Hayden, 1860. ··1980 Ruhoff: Cardium shumardi
Meek & Hayden, 1860; p. 182.
sicu/um Sowerby, 1834. [Figs. Part I p. 111]. - Synonym
of Parvicardium exiguum (Gmelin, 1791). »
Regarding Cardium siculum: Cardiidae. Recent. TYPE
LOCALITY. Ad mare Siculum (= Sicily), Sowerby (1840) .
RECORDS. ··1834 Sowerby, 51 th part, fig. 31: Cardium
siculum. ··1840 Sowerby, Proc. Z.S. London p.1 06:
Cardium siculum. ··1840 Sowerby, catalogue p. 2, No.
26: Cardium siculum Sow. ··1840 Sowerby, corrected
list, p. 8: Cardium sicu/um (Sowerby No. 26). ··1844
Philippi: Cardium sicu/um Sowerb. = Cardium exiguum
Gm. ··1862 Weinkauff: Cardium siculum Sow. =
Cardium subangulatum Scacchi. ··1858 Adams &
Adams, p. 457: Laevicardium sicu/um Sow. ··1867
Weinkauff: Cardium siculum = Cardium exiguum
Gmelin. ··1869 Romer: Cardium siculum Sowerby =
Cardium exiguum Gme\in. ··1878 Monterosato:
Cardium sicu/um Sow. = C. stellatum Reeve. ··1878
ZR: Cardium sicu/um (Sow.) = stellatum (Reeve) [Ref.
Monterosato 1878]. "1888 Kobelt: Cardium siculum
Sow. = Cardium exiguum Gmelin. ··1890 Paetel:
Cardium siculum Sow. = exiguum Gm. ··1917 Hidalgo:
Cardium siculum Sowerby = Cardium exiguum Gmelin.
··1922-32 Sherborn, p . 5944: Cardium siculum
Sowerby, 1841. ··1933 Coen : Cardium siculum
Sowerby = Eucardium (Parvicardium) exiguum var.
sllbangu/ata Scacchi. "1937 Coen: sicu/um Sowerby
Cerastoderma (Parvicardium) exiguum
subangulatum Scacchi. ··1962 Grossu : Cardium
siculum Sowerby, 1839 sic = Cardium (Parvicardium)
eXiguum Gmelin, 1790 sic. ··1969 Nordsieck:
Parvicardium siculum (Sowerby, 1839 sic). ··1977
Fischer-Piette: Cardium siculum Sowerby = Cardium
exiguum Gme\in. ··1989 VoskuiI & Onverwagt:
Parvicardium (Parvicardium) siculum (Sowerby,
1841). - CHARACTERS AND OCCURRENCE REMARKS. Cardium sicu/um was first figured by
Sowerby in 1834. DIMENSIONS. ··1840 Sowerby: long.
0 .5 poll. (length 13 mm), lat. 0.4 poll. (width 10 mm),
all. 0.45 poll (height II mm). ··1969 Nordsieck: stated
the following synonymy: Parvicardium exiguum
(exiguum) (Gmelin, 1791) = pygmaeum (Donovan,
1799) = sicu/um (Sowerby, 1839) = stella tum (Reeve,
1844) = parvum (Philippi, 1844). DISTRIBUTION. ··1969
Nordsieck: C. siculum: Mediterranean.
sidorovi Starobogatov 1974. - Hypanis minima sidorovi
Starobogatov. Lymnocardiidae. RECORDS. ··1974
Starobogatov, p. 246, fig. 213 : Hypanis minima sidorovi
Starobogatov subsp. nov.
silicatum Mansfield, 1937. - Combination to be revised:
"Cardium" silicatum Mansfield, 1937. Cardiidae. L.
Miocene. RECORDS. "1937 ZR: Cardium (Trachycardium) silicatum Mansfield, 1937 sp. n. Tampa
limestone, Florida (Lower Miocene).
silicea Blanckenhorn, 1934 . - Protocardia silicea
Blanckenhorn, 1934. Cardiidae. Cretaceous. RECORDS.
··1934 ZR: Protocardia silicea Blanckenhorn, 1934 sp.
n. Cretaceous Syria.
silvacollilla Dockery, 1982. - Trigoniocardia silvacollina
Dockery, 1982. Cardiidae. Lower Oligocene. RECORDS.
··1982 ZR: Trigoniocardia (Americardia) silvacollina
768
Tropical Marine Mol/usc Programme (TMMP)
simile - simplex
Dockery, 1982 sp. nov. Mississippi. Ref. to Dockery p.
72.
simile (variety). - See papillosum simile Milaschewitch,
1909.
simile Milachewitch, 1909. - Synonym of Parvicardium
papillosum (Poli, 1791). OTHER JUDGEMENTS: Cardium
exiguum Gmelin, 1791. » Regarding Cardium simile:
Cardiidae. Recent. RECORDS. ··1909 Milachewitch p.
318 : Cardium simile. ··1916 Milaschewitch: Cardium
simile Milaschewitsch, 1909. ··1962 Grossu: Cardium
(Parvicardium) simile Milaschevich (= Milachewitch)
1909. ··1977 Fischer-Piette: Cardium simile Milaschevich = Cardium exiguum GmeJin. - CHARACTERS
AND OCCURRENCE - OF COLLECTIONS. ··ZMUC: wet
collection from Bosporus IV. DISTRIBUTION. ··1969
Nordsieck: Black Sea.
similifittoni. - Nomen nudum . RECORDS. ··1949 ZR:
Replidacna similifittoni 'n o sp.' [nomen nud .]
Maestricktian, Sarmat. [Ref. Papp 1949].
simi/is Cowper-Reid, 1943. - Pinzonela similis CowperReid, 1943. Cardiacea. Trias. RECORDS. ··1947 ZR:
Pinzonela simi/is Cowper-Reid, 1943 sp. nov. Trias,
Brazil.
simi/is Kafanov, 1994. - Serripes similis Kafanov, 1994.
Cardiidae. RECORDS. ··1994 Kafanov in Arkhipova et
al.: Serripes similis Kafanov.
similis Papaianopol , 1989. - Pa chydaena similis
Papaianopol, 1989. Lymnocardiidae. Pliocene. TYPE
LOCALITY. Basin Dacique, Romania. REcORDs. ··1989
Papaianopol, p . 75, fig.: Paehydaena (Paehydacna)
similis sp. nov.
simillimum Smith, 1896. - New combination. Fragum
simillimum (Smith, 1896). Cardiidae. Recent. TYPE
LOCALITY. Off Colombo Lighthouse, in 142-400
fathoms. RECORDS. ··1896 Smith: Cardium (Fragum)
simillimum. ··1896 ZR: Cardium (Fragum) simillimum
Smith, 1896 n. sp. off Colombo . ··1909 Melvill :
Cardium (Fragum) simillimum, Smith. ··1937 Viader:
Cardium simillimum, Smith. ··1940 Winckworth:
Corculum (Fragum) simillimum. ··1948 Ray: Corculum
(Fragum) simillimum. ··1977 Fischer-Piette: Coreulum
(Fragum) simillimum Smith. - CHARACTERS AND
OCCURRENCE - CHARACTERS of diagnostic value.
The posterior portion of the surface occupies nearly
half the valve; it is divided off by two approximated
ribs. DESCRIPTIONS. ··1896 Smith: Description based on
a single left valve. The sculpture is similar to that of C.
peramabilis Dall, from 50-125 fathoms in the WestIndian region. However, the posterior end is distinctly
truncate, not curved as in C. peramabilis. ··1948 Ray:
nine complete shells and four valves were examined .
They agree in their essential characters with Smith's
description and figure. Its occurrence on the
Coromandel coast is, therefore of interest. The fine
radiating striae are absent in the lunular region, which
consequently is smooth. The granules, arranged in
radiating series, appear to be restricted to the hinder
part of the shell. Rounded anteriorly, truncate
posteriorly. The posterior portion of the surface is
somewhat more finely cancellate than the anterior, and
is divided off by two approximated ribs, but occupies
nearly half the valve. DIMENSIONS. ··1896 Smith: length
21 mm, width 21 mm, diameter 17 mm. HABITAT. ··1909
Melvill: Deep sea, 123-150 fathoms , on broken shell
and rough material; recorded to a depth of 400 fathoms.
DISTRIBUTION. ··1896 Smith: Indian Ocean. ··1948 Ray:
Bay of Bengal, the Coromandel coast.
simionesclli. - Combination to be revised: "Cardium"
simionescui. Cardiacea. Fossil. RECORDS. ··1930 ZR:
Cardium simioneseui, fossil [Ref. Vascautanu 1930] .
*simkewici. - Homonymy to be noticed. Cardium
simkewiei from 2 sources: Andrussow, 1902. Andrussow, 1963. REMARKS. I do not know how
Andrussow 1902 compares with Andrusov, 1963 .
Andrussow and Andrusov probably reflect different
transcriptions of the Russian alphabet.
simkewici Andrussow, 1902. - Changed combination:
Rarieardium simkewiei (Andrussow, 1902) .
Lymnocardiidae. Neogene. RECORDS. ··1905 Dollfus,
commenting Andrussow 1902: Cardium simkewici n.
sp. tends towards Cerastoderma . ··1986 Nevesskaya
et al.: Raricardium simkevici (Andrus) comb . nov.
··1987 ZR: Rarieardium simkevici (Andrus) comb. nov.
Ref. to Nevesskaya et al. p. 191.
simkewici Andrussow, 1963. - Combination to be
changed: "Cardium " simkewiei Andrusov, 1963 .
Lymnocardiidae . Neogene. RECORDS. ··1963 ZR :
Cardium simkewiei Andrusov, 1963 sp. nov. Russia
[spelled simkevici on plate].
simlaensis Vokes, 1937. - Discors simlaensis Vokes,
1937. Cardiidae. Eocene. REcORDs. ··1937 ZR: Diseors
simlaensis Vokes, 1937 sp. n. Lutetian, Eocene; Simla
Hills State, India.
simmetrica Ali-Zade & Aleskerov, 1969. - Monodacna
simmetrica Ali-Zade & Aleskerov, 1969.
Lymnocardiidae. Apsheron. RECORDS. ··1969 ZR :
Monodacna simmetrica Ali-Zade & Aleskerov, 1969
sp. nov. Apsheron deposits, Apsheron Peninsula,
Chemberkend, Azerbaidjan.
simollensis (variety). - See under echinatum var.
simouensis Cowper Reed, 1935.
simplex Douville, 1917. - Granoeardium simplex
Douville, 1917 . Cardiidae. Cretaceous. RECORDS.
··1917 ZR: Granocardium simplex Douville, 1917 n.
sp. Cretaceous Moghara.
simp/ex Spengler, 1799. [Figs. Part 3 p. 879]. - Changed
combination : Aerosterigma simplex (Spengler, 1799).
Cardiidae. Recent. TYPE LOCALITY. Holotype, Coast of
Guinea (Spengler 1799), without locality (Vidal 1999).
TYPE REPOSITORY. A paired specimen in ZMUC, Spengler
collection, Zoological Museum University of
Copenhagen (Vidal 1999). TYPE MATERIAL. Cardium
unieolor: Six paired syntypes from Ticao (Philippines),
in BMNH, Cuming coil., the largest of which fits the
specimen cited and figured by Sowerby, and was
selected as lectotype by Voskuil & Onverwagt (1991 :
116). Cardium nebulosum: Holotype, BMNH reg. no.
1900.2. 13.22. Neg. No. 0386. A paired specimen from
the Maldives figured by Reeve (Vidal 1999).
Laevicardium soyeri: Holotype and three paratypes, all
paired specimens, labeled (incorrectly) "Corsica", in
MNHN. RECORDS. ··1799 Spengler: Cardium simplex.
··1870 Morch : Cardium simplex Spengler; senior
synonym of C. maculosum Sow? non Wood = C.
arenieolum Reeve. ··1902 Sherborn, p. 904 : Cardium
simplex Spengler, 1799 . ··1977 Fischer-Piette:
Cardium simplex Spengler. ··1999 Knudsen &
Hylleberg: Cardium simplex Spengler, 1799. ··1999
Phuket Marine Biological Center Special Publication vol. 30 (2004)
769
simrothi - sinense
Vidal: Acrosterigma simplex (Spengler, 1799). ··200 I
Vidal, p. 407: Acrosterigma simplex. - CHARACTERS
AND OCCURRENCE - DESCRIPTIONS. Shell inflated,
round, 40 ribs, low, flat and smooth, interstices low
and very narrow. Outside white dotted with pale red
spots, inside white, but pale red under the umbones.
DESCRIPTIONS. ··1799 Spengler: Testa cordata
aequivalvi, obsolete sulcata, margine antico dentala.
··1799 Spengler, translated by Knudsen & Hylleberg
(1999): In external shape this bivalve very much
resembles the preceding species. The difference lies
alone in the shape, colour and number of ribs. Despite
its smallness it has no less than 40 of these; they are
not separated by furrows, but are situated very close to
each other; they are also low and flattened, without
scales and teeth, and consequently quite smooth. On
the external side it is strewn with pale red spots on a
white background. Internally, below the umbo, the same
pale colour prevails; the rest is white. It is slightly
smaller than the preceding one and comes from the
same place. ··1999 Vidal : Medium-sized; adult shells
generally asymmetrical on dorsal margin with a
receding posterior side and a raised, inflated anterior
one, and with an obtuse angle on dorsal margin at the
level of anterior laterals; ventral part also asymmetrical,
with an expansion of posterior side and ribs generally
curved backwards in projection. Generally slightly
truncated at posterior-median limit; a weak furrow
sometimes separates these two zones, producing an
obtuse notch on margin. Moderately but variably
elongated, and rather inflated. Lunule rather large and
flat on left valve, very large and appreciably depressed
on right valve. An additional thick dark-brown layer
of lamellated periostracum present on posterior slope
and part of median shell of the adult part. Exterior of
the young shells generally marked with irregular
concentric light purple stains; on adults, which
generally remain uniform pale white to yellow, these
aligned stains can persist on median part. Interiorly
white to somewhat purple by transparency; posterior
slope always white . Hinge slightly asymmetrical,
slightly angled. Mean rib number 47.6 ± 3.8, range 4259. On posterior slope, smooth anterior part of ribs flat,
with an abrupt anterior flank, and much wider than
posterior part, represented by a thin, low ridge between
axial furrow and interstice; this ridge, in tum, bears a
very thin top-furrow in which very small irregular
scales occur, often connected by a thin ridge. These
features are only present on juvenile part of shell; on
the adult part (except for some oflast ribs), the posterior
ridge and scales disappear and posterior part of ribs
becomes a "pseudo-interstice," simultaneously anterior
part of ribs lowers, shell surface becoming almost
smooth on the most adult margin of large shells. On
median part of shell there are low ribs of slightly
rounded profile, at first smooth beside posterior slope,
then becoming finely and regularly retro-ridged.
Interstices rounded, smooth, and half width of ribs.
Anteriorly, posterior thin ridges of ribs disappear or
change into scarce, irregular, very thin top-ridges; ribs
beside lunule become very low, sometimes with
somewhat wide, tubercular top-ridges, sometimes
entirely smooth and almost indistinct on fully adult part
of shell. ··200 I Vidal : tentacles numerous. No eyes on
tentacles. DIMENSIONS. ··1999 Vidal : length 39.4, height
52.8, width 36.0 mm. ··1799 Spengler: Slightly smaller
than C. Jucatum. REMARKS. ··1799 Spengler: outline
very similar to C. Jucatum. Major differences are the
number of ribs, their form and colour. ··1999 Vidal:
Acrosterigma simplex can be separated from the other
species of the species-group mainly by its large lunule,
the lack of bright colours (though exceptions exist),
unique character of posterior slope rib morphology
(particularly with ontogenetic disappearance of scales
and smoothing of surface and the weak ornamentation
of the ribs elsewhere. Junior synonyms according to
Vidal (1999): Cardium nebulosum Reeve, 1845;
Laevicardium soyeri Fischer-Piette, 1977; Cardium
unicolor Sowerby, 1840. HABITAT. "1999 Vidal:
recorded from 0 to 100 m depth. DISTRJBUTION. ··1799
Spengler: Guinea [West Africa, Hylleberg 200 I).
··1999 Vidal: Acrosterigma simplex has a large but
patchy distribution in the Indo-Pacific; it seems
inexplicably absent from the north-western Indian
Ocean and from the south--eastern west Pacific (eastern
Melanesia and Polynesia).
simrothi Dall 1898 . - Changed combination:
Trigoniocardia simrothi (Dall 1900). Cardiidae.
Tertiary. TYPE LOCALITY. China. RECORDS. ··1898 Dall:
Cardium (Trigoniocardia) Simrothi Dall, 1898 n. sp.
··1958 Abbott: Trigoniocardia simrothi Dall 1900.
··1977 Popov: Trigoniocardia (Trigoniocardia)
simzothi sic (Dall, 1900). ··1977 Vokes: Trigoniocardia
(Trigoniocardia) simrothi (Dall).
simulans Partsch . - Combination to be changed:
"Cardium" simulans Partsch. Cardiacea. Fossil.
RECORDS. ··1857 Deshayes: Cardium simulans Partsch.
sirnzothi err. spelling of simrothi Dall in Popov 1977.
sinense Sowerby, 1838. [Figs. Part 1 p. 112, 124, 174.
Part 3 p. 886). - Changed combination: Vepricardium
sinense (Sowerby, 1838). Cardiidae. Pliocene(?) and
Recent. Iredale (1929) Cardium sinense Sowerby is
type species of the genus Vepricardium. TYPE MATERJAL.
··2000 Vidal: type specimen Sowerby 1838: fig. 35.
Typfifas not been located. RECORDS. ··1838 Sowerby,
149 t part, fig . 35 : Cardium sinense nobis. ··1840.
Sowerby, Proc. Z.S. London p.1 05: Cardium sinense.
"1840 Sowerby, catalogue No.4: Cardium sinense.
··1840 Sowerby, corrected list, p. 8: Cardium sinense
(Sowerby No . 4). "1844 Reeve: Cardium sinense
Sowerby, Pro . Zool. Soc., 1840. Conch. IIIus. f. 35.
"1858 Adams & Adams, p. 454: Cardium
(Pectunculus) sinense Sow. ··1884 Grassel: Cardium
sinense. Sow. "1890 Paetel: Cardium sinense Sow.
··1891 Fischer: Cardium sinense Sowerby. ··1907 ZR :
Cardium sinense [Ref. Dautzenberg & Fischer 1907).
··1909 Lynge: Cardium (Bucardium) sinense. ··1919
Martin: Cardium sinense Sow. [Pliocene(?) to Recent).
··1922-32 Sherborn, p. 5979: Cardium sinense
Sowerby, 1841. "1929 Iredale: sinense (Sowerby).
··1962 Kira: Vepricardium sinense. ··1974 Habe &
Kosuge: Vepricardium sinense (Sowerby, 1844). ··1977
Fischer-Piette: Laevicardium sinense Sowerby. ··1995
Chung-Hung & Hsi-Jen: Cardium sinense Sowerby.
.. 1999 Aungtonya et al ., PMBC-SP 19(2) : 378:
Vepricardium sinense (Sowerby, 1841). ··2000
Hylleberg, p. 532: Vepricardium sinense (Sowerby,
1840). ··2000 Lutaenko, p. 372: Vepricardium sinense
770
Tropical Marine Mol/usc Programme (TMMP)
sinuatus - soceni
(Sowerby, 1841). ··2000 Vidal, p. 454, Fig. 4: Cardium
sinense Sowerby, 1841: 105. ··2001 Vidal, p. 407:
Vepricardium sinense. "2003 Hy\leberg & Kilburn, p.
186, Plate 5, fig. 21: Vepricardium sinense (Sowerby,
1840). - CHARACTERS AND OCCURRENCE CHARACTERS of diagnostic value: ··1844 Reeve: the ribs
of this species are peculiarly narrowed along the
summit; and the granulated surface of the shell is of an
extremely curious character. ··2000 Vidal: V sinense
is so characteristic, with its few triangular ribs, that it
has never been misidentified. OF COLLECTIONS.
··ZMUC: Siam 3-5 fv, soft clay, 9 fv mud, shells, 5-9
fv sand, 30 fv shells. ··ZMUC: Gulf of Thailand 5 fv
soft clay, 8 fv, mud, 10 fv mud China from Ludvigsen.
··ZMUC: wet collection: sine loc. DESCRlPTIONS. ··1844
Reeve: shell somewhat depressly orbicular, posteriorly
slightly beaked, everywhere minutely granulated,
especially on the anterior side; radiately ribbed, 24 ribs,
a little compressed, posterior narrower; fulvous white,
margins very strongly crenated. ··200 I Vidal: tentacles
numerous. Eyes on tip of tentacles. ··ZMUC: 23 ribs,
8 of which are posterior. DIMENSIONS. ··1840 Sowerby:
long. 1.55 poll. (length 39 mm), lat. I poll. (width 25
mm), alt. 1.40 poll (height 36 mm). REMARKS. ··1840
Sowerby: Slightly resembling C. asiaticum, from which
it is distinguished by having larger and fewer ribs, and
a small fringe on the posterior ribs. HABITAT. Found in
sand. DISTRIBUTION. ··1844 Reeve: China and Java.
··1884 Grasset: China. ··1994 Hylleberg: Gulf of
Thailand, Philippines. ··1999 Aungtonya et al.:
Thailand, Andaman Sea. ··2000 Hylleberg: Vietnam
and Cambodia. ··2000 Lutaenko: Vietnam. "2000
Vidal: Never recorded in the Indian Ocean, probably
only west Pacific: Java, Philippines, Cambodia, Gulf
of Thailand, Vietnam, south China, Japan. ··2003
Hylleberg & Kilburn: Vietnam, 10-20 m, near shore.
sinuatus Bog. - Valenciennius sinuatus Bog. Cardiacea.
Fossil. RECORDS. ··1955 Ebersin: Valenciennius
sinuatus Bog.
sinuosum d'Orbigny, 1850. - Cardium sinuosum
d'Orbigny, 1850. ?Cardiidae. Fossil. RECORDS. ··192232. Sherborn, p. 5990: Cardiola sinuosa Muenster,
1840. ··1922-32. Sherborn, p. 5990: Cardium
(Muenster) sinuosum d 'Orbigny, 1850. - Cardiola.
Sinupontalmyra (subgenus). - RECORDS. ··1981 ZR:
Pontalmyra (Sinupontalmyra) Papaianopol, 1981
subgen. nov. Type species Cardium corbuloides
Deshayes, 1838, Upper Miocene. Ref. to Papaianopol,
p. 41. ··1987 Taktakishvili pp. 58-85: (Sinupontalmyra)
Taktakishvili, 1987 subgen. nov. of Crassadacna
Ebersin.
sinzovi Sinz. - Monodacna sinzovi Sinz. Lymnocardiidae. Fossil. RECORDS. ··1907 Andrussow:
Monodacna sinzovi Sinz.
siphonophorum Andrussow, 1902. - Combination to be
revised: "Cardium" siphonophorum (Andrussow,
1902). Cardiidae. Miocene. RECORDS. ··1905 Dollfus,
commenting
Andrussow
1902:
Cardium
siphonophorum n. sp. tends towards Cerastoderma.
··1961 ZR: Cardium siphonophorum Andrusov, 1961
"sp. nov."[in a tabl.e] Miocene Caspian region, Russia.
··1972 ZR: Cardium siphonophorum [Ref.
Archambault 1972].
sjoegreni. - Monodacna sjoegreni. Lymnocardiidae.
Pliocene. RECORDS. ··1916 ZR: Monodacna sjoegreni
Ref. to Kalickij (1916). ··1963 ZR: Monodacna
sjoegreni. Pliocene. ··1977 Popov: Monodacna
sjoegreni.
sjoegreni minor Andrussov, 1924 (variety). - Upgraded.
Monodacna sjoegreni minor Andrussov, 1924.
Lymnocardiidae. Pliocene and Pleistocene. RECORDS.
··1924 ZR: Monodacna sjoegreni minor Andrussov,
1924 var. n. Pleistocene. ··1963 ZR: Monodacna
sjoegreni var. minor Andrusov, 1963 nov. Pliocene,
Russia.
sjoegreni pyrophila Andrussov, 1924 (variety). Upgraded. Monodacna sjoegreni pyrophila Andrussov,
1924. Lymnocardiidae. Pliocene and Pleistocene.
RECORDS. ··1924 ZR: Monodacna sjoegreni pyrophila
Andrussov, 1924 var. n. Pleistocene. ··1963 ZR:
Monodacna sjoegreni var. pyrophila Andrusov, 1963
nov. Pliocene, Russia.
skadovskense Eberzin, 1958. - New combination:
Lymnocardium skadovskense (Eberzin, 1958).
Lymnocardiidae. Pliocene. RECORDS. ··1958 ZR:
Limnocardium (Euxinicardium) skadovskense Eberzin,
1958 sp. nov. Ukraine, Skadovsk north coast of Black
Sea. Pliocene of the Ponto-Caspian basin, Russia.
skeeti Hedley, 1906. [Figs. Part 3 p. 918]. - Changed
combination: Afrocardium skeeti (Hedley, 1906). OTHER
JUDGEMENTS: Afrocardium richardi (Audouin, 1827).
Cardiidae. Holocene and Recent. TYPE LOCALITY.
Australia. TYPE REPOSITORY. Australian Museum Sydney.
Syntypes C018858. RECORDS. ··1906 Hedley, p. 476,
pI. 38, fig. 25: Cardium skeeti Hedley n. sp. ··1906
ZR: Cardium skeeti Hedley, 1906 sp. n. Mast Head
Reef Q. ··1977 Fischer-Piette: Cardium (Ringicardium) skeeti Hedley. ··1992 Lamprell & Whitehead:
Afrocardium skeeti. ··2002 Schneider: Afrocardium
skeeti (Hedley, 1906). Holocene. ··2003 Hylleberg &
Kilburn, p. 176: C. skeeti Hedley, 1906. Syn of
Afrocardium richardi (Audouin, 1827) according to J.
Vidal. - CHARACTERS AND OCCURRENCE DISTRIBUTION. ··1994 Hylleberg: East Australia.
slatteri Cox, 1946. - Tutcheria slatteri Cox, 1946.
Cardiacea. Middle Lias. RECORDS. ··1946. ZR:
Tutcheria slatteri Cox, 1946. sp. nov. Middle Lias,
Aston Magna, Wores.
slavonica (variety). - See under abichi var. slavonica
Jenko, 1961.
smekhovi Kafanov & Savizky, 1982. - Ciliatocardium
smekhovi Kafanov & Savizky, 1982. Cardiidae.
RECORDS. ··1982 Kafanov & Savizky: Ciliatocardium
smekhovi Kafanov & Savizky. ··1982 ZR:
Ciliatocardium smekhovi sp. nov. Russian SFSR,
Lower Miocene. Ref. to p. 58, fig'd Kafanov.
snideri Morgan. - Conocardium snideri Morgan.
Conocardiidae. Fossil. RECORDS. ··1960 ZR:
Conocardium snideri synonymy, description and
photographs of the ecotypes of Morgan's species [Ref.
Branson I 960b].
snjatkovi Fedotor, 1932. - Conocardium snjatkovi
Fedotor, 1932. Conocardiidae. Carboniferous. RECORDS.
··1932 ZR: Conocardium snjatkovi Fedotor, 1932 sp.
n. Carboniferous Donetz Basin. ··1933 ZR:
Conocardium snjatkovi Fedotov, 1932 sp. n.
Carboniferous.
soceni lekelius, 1944. - Prophyllicardium soceni
PhI/kef Marine Biological Center Special Publication vol. 30 (2004)
771
soceni -solidum
Jekelius, 1944. Lymnocardiidae. Pliocene. RECORDS.
--1946 ZR: Prophyllicardium soceni Jekelius, 1944 sp.
nov. type of Prophyllicardium Jekelius, 1944 gen. nov.
Cardiidae, Pliocene Rumania (Lower Congerien)
Soceni. Banat.
soceni Jekelius, 1944. - Replidacna soceni Jekelius,
1944. Lymnocardiidae. Miocene. REcORDs. --1946 ZR:
Replidacna soceni Jekelius, 1944 sp. nov. Miocene
Soceni Banat.
sociale Krauss, 1852. - Changed combination:
Cerastoderma sociale (Krauss, 1852). Cardiidae.
Fossil. REcORDs. --1977 Popov: Cerastoderma sociale.
"1980 Ruhoff: Cardium sociale Krauss, 1852; vol. 8:
154, pI. 3, fig. 7.
socialis Papaianopol, 1989. - Pachydacna socialis
Papaianopol, 1989. Lymnocardiidae. Pliocene. TYPE
LOCALITY. Basin Dacique, Romania. RECORDS. "1989
Papaianopol, p. 69, fig. : Pachydacna (Pachydacna)
socialis sp. nov.
socrivenori Tamura, 1968 . - Cardium socrivenori
Tamura, 1968. Cardiidae. Fossil. RECORDS. "1976 Ma
et al. p. 255: Cardium socrivenori Tamura, 1968.
sokkohense Martin, 1917. - Combination to be revised:
"Cardium " sokkohense Martin, 1917 . Cardiidae .
Lower Miocene. TYPE LOCALITY. West-Progogeb. Java,
Indonesia. TYPE REPOSITORY. ?Geologischen ReichsMuseum, Leiden, the Netherlands. REcORDs. "1917
ZR: Cardium sokkohense Martin, 1917 n. sp. Miocene
Java. --1919 Martin: Cardium sokkohense Martin
1916-1917 [L. Miocene].
sokolovi Andrussow, 1930. - Changed combination:
Macradacna sokolovi(Andrussow, 1930). Lymnocardiidae. Fossil. REMARKS. I do not know how
Andrussow, 1930 compares with Wassoievic &
Eberzin, 1930 (see below). REcORDs ... 1967 Ebersin:
Macradacna sokolovi (Andrus.). --1967 ZR:
Macradacna sokolovi (Andrus., 1930) pro Monodacna
sokolovi (Andrus., 1930) [Ref. Eberzin 1967].
sokolovi Wassoievic & Eberzin, 1930. - Monodacna
sokolovi Wassoievic & Eberzin, 1930. Lymnocardiidae.
Pliocene. RECORDS. --1931 ZR: Monodacna sokolovi
Wassoievic & Eberzin, 1930 sp. n. URSS
sola (variety). - See under subsquamulosum var. sola
Eberzin, 1958.
Solen (genus) Linne, 1758. Not in Cardiidae. REcORDs.
"1857 Deshayes: Solen (spec.) Linne, in Schroeter,
Spengler, Gmelin, Dillwyn: syn. of Cardium.
soleniforme (mode). - See under solen iform is Bruguiere,
1789.
soleniforme Bruguiere, 1789. [Figs. Part I p. 70, 88,
102. Part 3 p . 917]. - Changed combination: Papyridea
soleniforme (Bruguiere, 1789). OTHER JUDGEMENTS:
Cardium bulla/urn Lam. - Papyridea spinosum
(Meuschen, 1778) [not available]. » Regarding
Cardium soleniforme: Cardiidae. Recent. Gray (1847)
designated Cardium soleniforme Bruguiere, 1789 type
species of the genus Papyridea Swainson, 1840.
REcORDs. --1789 Bruguiere, p . 235 , No. 34: Cardium
soleniforme Nob. Compared with Lister, tab. 342, fig.
179. --1836 Bosc: Cardium soleniforme. --1840
Sowerby, catalogue p. 2, No. 14: Cardium soleniforme
Brug. = Cardium bulla tum Lam. --1844 Potiez &
Michaud : Cardium soleniforme Brug. = Cardium
bullatum Linn . Gmel. .. 1847 Gray: Cardium
soleniforme Bruguiere, 1789. "1850 Jay: Cardium
soleniforme Brug. = Cardium bullatum. --1856 Wood
& Hanley, No. 36: Cardium soleniforme Brug. --1864
Krebs: Cardium soleniforme Brug. (syn.) = Cardium
spinosum Meusch . --1869 Romer: Cardium
soleniforme Bruguiere N . 34 = Cardium bullatum (L.)
Chemnitz . "1885 Smith : Cardium soleniforme,
Bruguiere = Cardium (Papyridea) bulla/urn ([Linn. ?]
Chemnitz). --1902 Dall : Cardium soleniforme
Bruguiere, 1789 = Cardium spinosum Meuschen, 1778.
--1902 Sherborn , p. 911: Cardium soleniforme
Bruguiere, 1789 . "1915 Tomlin & Shackleford:
Papyridea soleniformis (Brug., 1789). "1921 Maury:
soleniforme Bruguiere, 1789 = Cardium (Papyridea)
spinosum Meuschen, 1778. --1939 McLean: Cardium
soleniforme Bruguiere, 1789 = Papyridea spinosum
(Meuschen, 1778). --1953 Olsson & Harbison:
Cardium soleniforme Bruguiere = Cardium hiatus
Meuschen. --1963 Coomans: Papyridea soleniformis
(Bruguiere) = Papyridea hiatus (Meuschen). --1968
Jong & Kristensen: Papyridea soleniformis Bruguiere
= P hiatus Meuschen. --1969 Moore et af.: Papyridea
soleniforme (Bruguiere, 1789). --1969 Nordsieck :
Papyridea soleniforme (Bruguiere, 1792). --1970 Rios :
Papyridea soleniformis (Bruguiere, 1789). --1971 Keen
& McLean : Papyridea soleniforme (Bruguiere, 1789).
--1977 Fischer-Piette: Papyridea soleniforme
Bruguiere. --1977 Popov p. 49: Cardium soleniforme
Bruguiere, 1789. Type of Papyridea Swainson, 1840.
--1977 Wilson & Stevenson: Cardium soleniforme
Bruguiere , 1789. --1997 Shelton: Papyridea
soleniformis (Bruguiere, 1789). - CHARACTERS
AND OCCURRENCE - OF COLLECTIONS. --ZMUC:
soleniforme, 3 specimens from Tortula, Brug. Encycl.
pI. 296 f. 6 a,b. --ZMUC: 2 specimens. sine loc.
--ZMUC: 2 specimens labeled West Indies. --ZMUC:
GulfofMexico, 64 m, rock. 1 old shell. Ornamentation
differ somewhat from the west Indian specimens in the
collection, but it is close to bullata from St Helena,
Sea Cow cove, 15 fv, identi fied by Odhner.
DESCRIPTIONS . --1819 Lamarck: Reddish spines .
HABITAT. --ZMUC Rocky bottom at 64 m depth. --1969
Nordsieck : 0-37 m depth. DISTRIBUTIO N. --1969
Nordsieck: Brazil, Florida-West Indies, West Africa,
Cape Verde Islands, Lusitanean Seas ... 1971 Keen &
McLean : South East USA, Brazil , West Africa;
Caribbean.
soleniformis aspera Sowerby, 1833 (subspecies) . Papyridea soleniformis aspera (Sowerby, 1833) .
Cardiidae. Recent. RECORDS. --1961 Olsson: Papyridea
soleniformis aspera (Sowerby, 1833).
solense Senes, 1955. - Combination to be revised:
"Cardium" solense Senes, 1955. Lymnocardiidae.
Miocene. RECORDS. --1955 ZR: Cardium (Limnocardium) solense Senes, 1955 sp. nov. Miocene,
Czechoslovakia
solida (variety). - See under groenlandicum var. solida
Morch, 1868.
solidum Linnaeus, 1758. [Figs. Part 1 p. 37]. - Not in
Cardiidae. Recent. RECORDS. --1678 Lister. ID by
Linnaeus 1758: Cardium solidum angl. t. 4. f. 24. "1684
Bonanno. ID by Linnaeus 1758: Cardium solidum recr.
3 sic. t. 51. .. 1758 Linnaeus (loth edition) No. 76:
Cardium solidum. "1786 Solander, p . 17 no. 382 :
772
Tropical Marine Mollusc Programme (TMMP)
solimani - sorrentoense
Cardium solidum. ··1788 Schroter: Cardium solidum
[Reference to Chemnitz (1782) fig . 231]. ··1902
Sherborn, p. 911: Cardium solidum Linnaeus, 1758
[Mactra, ed. 12] .
solimani Not in Cardiidae. "1966 ZR: lsocardia
(lsocardia) solimani Abbass, 1966 sp. nov. Cretaceous,
Egypt [Ref. Abbass 1966] REMARKS. Probably placed
in Cardiacea of ZR by mistake. Should be in Family
Isocardiidae.
solitaria Papaianopol, 1989. - Pachydacna solitaria
Papaianopol, 1989. Lymnocardiidae. Pliocene. TYPE
LOCALJTY. Basin Dacique, Romania. RECORDS. "1989
Papaianopol, p. 76, 88, fig.: Pachydacna (Pachydacna)
solitaria sp. nov.
solitarium Krauss, 1852. - Combination to be revised :
"Cardium" solitarium Krauss, 1852. Cardiidae. Fossil.
RECORDS . ··1980 Ruhoff: Cardium solitarium Krauss,
1852; vol. 8: ISS, pI. 3, fig. 8.
solitarius Papaianopol, 1992. - Pseudocatillus solitarius
Papaianopol, 1992. Lymnocardiidae. Pliocene. TYPE
LOCALJTY. Dacien inferieur (Getien). Basin Dacique,
Romania. RECORDS. "1992 Papaianopol, p. 25, fig .:
Pseudocatillus solitarius sp. nov.
solutum Uspenskaia, 1931. - Combination to be revised:
"Cardium" solutum Uspenskaia, 1931. Cardiidae.
Miocene. RECORDS. ··1932 ZR: Cardium solutum
Uspenskaia, 1931 n. sp . Miocene Daghestan and
Transcaspian.
Somalidacna Azzaroli, 1958 (genus) . - Cardiacea.
RECORDS. ··1958 ZR: Somalidacna Azzaroli, 1958 gen.
nov. Shows affinity to the Tridacnidi with pronounced
convergence towards the Osteridai. Guban Series.
Piano Dubar, Oligocene.
somaliensis Cox, 1935. - Protocardia somaliensis Cox,
1935 . Cardiidae. Jurassic. RECORDS. ··1935 ZR:
Protocardia somaliensis Cox, 1935 sp. n. Jurassic
British Somali land.
sonense Cossmann, 1895. - Changed combination:
Parvicardium sonense (Cossmann, 1895). Cardiidae.
Fossil. RECORDS. ··1977 Popov: Parvicardium sonense
(Cossmimn, 1895). ··1994 Marcomini: Cardium
(Parvicardium) sonense Cossmann, 1895.
sookense Clark & Arnold, 1923 . - Changed combination:
Clinocardium sookense (Clark & Arnold, 1923).
Cardiidae. Fossil. ··1923 ZR: Cardium sookensis Clark
& Arnold, 1923 Tertiary Vancouver Isd. ··1977 Popov:
Clinocardium sookense (Clark & Arnold, 1923).
sookensis (mode). - See under sookense Clark & Arnold,
1923.
.
soosi Bartha, 1954. - New combination: Lymnocardium
soosi Bartha, 1954. Lymnocardiidae. Pliocene.
RECORDS. ··1954 ZR: Limnocardium soosi Bartha, 1954
sp. nov. Pliocene. bcs.
soproniense Vital is, 1936. - New combination:
Lymnocardium soproniense Vital is, 1936.
Lymnocardiidae. Pontian. RECORDS. ··1937 ZR:
Limnocardium soproniense Vitalis, 1936 sp. n. Pontian
Hungary.
sorenseni Powell, 1958 . [Figs. Part 3 p . 865]. Trachycardium sorenseni Powell, 1958. Cardiidae.
Recent. TYPE MATERIAL. Holotype: A right valve. Two
other smaller valves in Auckland Museum from
Galathea 1952, stn 674 off Raoul Is, 75-85m, paratypes
were cited but not qualified by the author (Vidal 1999).
TYPE LOCALITY. Denham Bay, Raoul Island, Kermadec
Islands, 0 m. TYPE REPOSITORY . Auckland Museum, New
Zealand AK701236. RECORDS. "1958 Powell, p. 76,
pI. II, figs 6--7: Trachycardium sorenseni. ··1958 ZR:
Trachycardium (Vasticardium) sorenseni sp. nov.
Kermadec Islands, S. Pacific. ··1977 Fischer-Piette:
Laevicardium sorenseni Powell. ··1999 Vidal:
Trachycardium (Vasticardium) sorenseni Powell, 1958.
- CHARACTERS AND OCCURRENCE - CHARACTERS of diagnostic value. Anterior surface area with thin
interstices, high ribs which are square-sided rather than
rounded and, on the ribs, wide, imbricated ridges which
almost touch DESCRIPTIONS. ··1999 Vidal: Shell rounded
in outline. Posterior slope area very slightly flattened.
Lunule non-existent, ribs practically reaching margins.
Colour of fresh shells whitish heavily blotched with
orange-pink; interior with external colour pattern
showing through; holotype, a faded beach shell, shows
a sparsely speckled pattern on a pale ground. Hinge
variably asymmetrical and rather moderately angled.
Mean rib number 51 .2 ± 1.4 (range 48-54). On posterior
slope, ribs with irregular, somewhat elongated oblique
tubercles on posterior half of ribs. In central area ribs
retro-ridged, almost square-sided, but with rounded
tops, and a slight forward bending; interstices about II
3 of ribs width, and loosely striated. On anterior part
of shell, ribs become higher, square-sided, flat-topped
and top-ridged with wide touching ridges which are
sometimes imbricated. Interstices become progressively narrower. DIMENSIONS. Length 27.5, width 29.4,
breadth (calculated) 15.2 illill. REMARKS . ··1999 Vidal:
Acrosterigma sorenseni is close to A. cygnorum. The
former mainly differs in: (1) higher rib number (48-54
against 35-47 in cygnorum); (2) presence on PQ of a
discontinuous axial furrow, formed by successive holes
between the scales; (3) different rib morphology on
AQ, i.e. thinner interstices, higher ribs which are
square-sided rather than rounded and, on the ribs,
wider, more imbricated ridges which almost touch
(instead of the thin, free ridges of A. cygnorum). The
two valves in New Zealand Oceanographic Institute
cited by Powell (1958: 76) from E of Philip lsI., Norfolk
lsI. "similar to the Kermadec species in sculpture but
of oblique-ovate outline, and with a rib count 48-50"
are very probably A. maculosum howense, abundant in
Norfolk lsI. HABITAT. Recorded from 22 to 47 m depth.
DISTRIBUTION. Raoul Island, Kermadec Islands, New
Zealand.
soriensis Eames, 1951. - Discors soriensis Eames, 195 I .
Cardiidae. Eocene. RECORDS . ··1951 ZR: Discors
soriensis Eames, 1951 sp. nov. Eocene Pakistan.
sorokini Androussov, 1923 . - Apscheronia sorokini
Androussov, 1923. Lymnocardiidae. Pliocene.
RECORDS. "1930 ZR: Apscheronia sorokini
Androussov, 1923 sp. n. ··1963 ZR: Apscheronia
sorokini Andrusov, 1963 sp. nov. sic Apsheron layers,
Pliocene, Russia.
sorrentoense Hanna, 1927. - Changed combination :
Agnocardia sorrentoense (Hanna, 1927). Cardiidae.
Eocene. RECORDS. ··1927 ZR: Cardium sorrentoensis
Hanna, 1927 sp. n. Eocene California. ··1944 Keen &
Bentson: Cardium sorrentoense Hanna, 1927 (n. sp.).
··1944 Keen & Bentson: Trachycardium (Agnocardia)
sorrentoense (Hanna) . ··1984 ZR: Agnocardia
773
Phukel Marine Biological Center Special Publication vol. 30 (2004)
sorrentoensis - speciosum
sorrentoensis (Hanna, 1927). Taxonomic position. Ref.
to Vokes p . 43. --2002 Schneider: Agnocardia
sorrentoensis (Hanna, 1927). Cardiidae.
sorrentoensis (mode). - See under sorrenloense Hanna,
1927.
sotter;; Michelotti, 1839. - Combination to be revised:
"Cardium " solterii Michelotti, 1839. Cardiidae. Fossil.
RECORDS . "1922-32 Sherborn, p. 6033: Cardium
sotterii Michelotti, 1839.
sougraignense Dechaseaux, 1938. - Combination to be
revised: "Cardium" sougraignense Dechaseaux, 1938.
Cardiidae. Santonian. RECORDS. --1948 ZR: Cardium
sougraignense Dechaseaux, 1938 sp. nov. Santonian
Corbiere.
sowerbianum d ' Orbigny, 1850. - Combination to be
revised: "Cardium " sowerbianum d'Orbigny, 1850.
Cardiidae. Fossil. RECORDS. "1922-32 Sherborn, p.
6033: Cardium sowerbianum d'Orbigny, 1850.
sowerbyorum Voskuil & Onverwagt, 1992. - New name
reversed to original name: Vasticardium ornatum
(Sowerby, 1877). « Regarding Trachycardium
sowerbyorum: Cardiidae. Recent. DISTRIBUTION. Hong
Kong. RECORDS. --1992 Voskuil & Onverwagt, p. 41 :
Trachycardium sowerbyorum nom. nov.; for Cardium
ornalum Sowerby, 1876 nec Orbigny 1850. --1997
Vidal: Acrosterigma sowerbyorum Voskuil and
Onverwagt, 1992 (nomen novum for Cardium ornalum
erroneously considered unavailable) = Vasticardium
ornatum (Sowerby, 1877). --1999 Vidal: Acrosterigma
sowerbyorum Voskuil & Onverwagt, 1992 =
Vasticardium ornatum (Sowerby, 1877).
soyeri Fischer-Piette, 1977. [Figs. Part 3 p. 880] . Synonym of Acrosterigma simplex (Spengler, 1799).
OTHER JUDGEMENTS: Cardium unicolor Sowerby, 1834.
» Regarding Laevicardium soyeri: Cardiidae. Recent.
TYPE LOCALITY. ? Mediterranean area. TYPE REPOSITORY.
Museum national d'Histoire naturelle, Paris, Musee des
Colonies, holotype and 3 paratypes. TYPE MATERIAL.
Holotype and three paratypes, all paired specimens,
labeled (incorrectly) "Corsica", in MNHN. RECORDS.
"1977 Fischer-Piette, No. 22, p. 19, pI. 1, figs. 4-7:
Laevicardium soyeri n. sp. --1977 Fischer-Piette:
Cardium oblongum L., Gmel. sp. 71 in Reeve [non
Gmelin] 1845 = Laevicardium soyeri n. sp . --1977
Fischer-Piette: Cardium oblongum Chemnitz in
Granger 1886 = Laevicardium sayeri n. sp. --1991
Voskuil & Onverwagt: Laevicardium soyeri FischerPiette, 1977, junior synonym of Cardium unicolor
Sowerby, 1834. --1999 Vidal : Laevicardium soyeri
Fischer-Piette, 1977: 19, pI. I, figs 4-7 = Acrosterigma
simplex (Spengler, 1799). - CHARACTERS AND
OCCURRENCE - CHARACTERS of diagnostic value.
Sculpture on the entire shell surface ; 42-57 ribs.
DESCRIPTIONS. --1977 Fischer-Piette: shells slightly
inequilateral, tightly fitting; sculpture of 49 ribs on right
valve, 52 on the left valve, flat ribs with concentric
costulation, very fine and serrated, but visible with
naked eye, lunule and area not developed, ligament 9
mm long, white-yellow coloration, with remains of
olive coloured periostracum, white inside, but around
the cardinal plate is a confuse purple coloration divided
by a white median stripe [I have translated from
French]. DIMENSIONS. Umbo-ventral 48 mrn, anteriorpost. 36 mm, diameter 32 mm. REMARKS. --1980 Keen:
Probably a Vasticardium with incorrect locality data.
The species is close to Laevicardium oblongum
(Gmelin, 1791), which, however, only has 24-30 ribs.
DISTRIBUTION. Mediterranean, Canary lsI.
spadicea Meuschen. Non binominal. - Not available.
Not in Cardiidae . Meuschen, in the Museum
Geversianum, No . 1621, it is quoted for Cardium
ciliare. RECORDS. --1823 Dillwyn, Tab. 340 fig. 177:
Venus spadicea.
spatiosum Hutton, 1873. - Changed combination: Plagiocardium spatiosum (Hutton, 1873). Cardiidae.
Miocene and Pliocene. Type species: Hutton (1873)
designated Cardium spatiosum type species of the
subgenus (Maoricardium) Hutton, 1873 . Recognised
at the generic level by authors. DISTRIBUTION . New
Zealand. RECORDS. --1873 Hutton: Cardium spatiosum .
--1897 Harris : Cardium spatiosum , Hutton. 1873 .
[Miocene] . --1914 ZR: Cardium spatiosum [Ref. Suter
1914]. --1944 ZR: Cardiumspatiosum Hutton, Pliocene
(Catal. Tert. Moll. 1873, 23) genotype of Maoricardium
Marwick, I 944a. --1965 ZR: Maoricardium spatiosum
revised, Tertiary, New Zealand [Ref. Boreham 1965].
"1969 Moore et al.: Plagiocardium (Maoricardium)
spatiosum (Hutton, 1873). --1977 Popov: Cardium
spatiosum Hutton, 1873. Type of Maoricardium
Marwick, 1944. --1977 Wilson & Stevenson:
Maoricardium spatiosum (Hutton, 1873). "1977
Wilson & Stevenson: Cardium spatiosum Hutton, 1873
(tertiary fossil) .
speciosa (mode). - See under speciosum Adams &
Reeve, 1850.
*speciosum. - Homonymy to be noticed. Cardium
speciosum from 2 sources: Adams & Reeve, 1848. Meek & Hayden, 1856.
speciosum Adams & Reeve, 1848. [Figs. Part 3 p. 904].
- Changed combination: Americardia speciosa (Adams
& Reeve, 1848). Cardiidae. Recent. TYPE LOCALITY.
China Sea. TYPE REPOSITORY. 3 syntypes, BMNH,
London. RECORDS. "1848 Adams & Reeve, p. 77, pI.
22, fig. 9: Cardium speciosum. --1858 Adams &
Adams, p. 459: Hemicardia (Fragum) speciosa Adams
and Reeve. --1884 Grasset: Hemicardium speciosum.
Ad. --1890 Paetel: Hemicardium speciosum Ad. et Rv.
--1900 Dall: Cardium speciosum Adams and Reeve,
1850 sic is not a synonym of the Cretaceous Cardium
whitei Dal1. "1922-32 Sherborn, p. 6046: Cardium
speciosum Adams & Reeve, 1850. --1977 FischerPiette: Corculum (Hemicardia) speciosum Adams &
Reeve, 1850. - CHARACTERS AND OCCURRENCE
- DESCRIPTIONS. --1848 Adams & Reeve: Card. testa
subcordata, crassiuscu/a, tumida. postice peculiariter
conr:avo-truncata, a/bo rufoque tesse/ata, radiatim
costata, costis ad sex et viginti, convexis, squama togranu/atis, interstitiis transversim striatis. REMARKS.
--1977 Fischer-Piette: Cardium speciosum is allied in
form to C. fornicatum Sowb. (Melvill & Standen 1906,
p. 840) and C. medium. DISTRIBUTION. --1884 Grasset:
China.
speciosum Meek & Hayden, 1856. - Changed combination: Granocardium speciosum (Meek & Hayden,
1856). Cardiidae. Cretaceous. White (1880) designated
Cardium speciosum Meek & Hayden, 1857 type species
of subgenus Ethmocardium of genus Granocardium.
Dall (1900) Cardium whitei Dall is type species for
774
Tropical Marine Mollusc Programme (TMMP)
speluncarium - spinosum
the subgenus Ethmocardium. TYPE LOCALITY. Mouth of
Judith River, Montana. TYPE REPOSITORY. Lectotype
USNM 190, Hypotype YPM 24180 held at YPM
(Speden 1970). RECORDS. ··1860 Meek & Hayden:
Cardium speciosum Meek & Hayden, 1856 . •• I 876
Meek: Cardium (Criocardium) speciosum Meek &
Hayden, 1856. ··1880 White: Cardium speciosum
Meek & Hayden, 1857 ... 1900 Dall: Cardium whitei
Dall. ··1954 Keen: Cardium speciosum Meek and
Hayden (1857) = C. whitei Dall, 1900 ... 1969 Moore
et al.: Granocardium (Ethmocardium) speciosum
(Meek & Hayden, 1857). ··1970 Speden: Cardium
speciosum Adams & Reeve = Cardium (Ethmocardium)
whitei Dall, 1900 . •• I 980 Ruhoff: Cardium speciosum
Meek & Hayden, 1856; vol. 8: 274.
speluncarium Neum. - Combination to be changed:
"Cardium" speluncarium Neum. Cardiacea. Fossil.
RECORDS . •• I 987 Taktakishvi Ii: Cardium speluncarium
Neum.
speluncense Almera y Bofill, 1894. - Combination to
be revised: "Cardium" speluncense Almera y Bofill,
1894. Cardiacea. Miocene. RECORDS. ··1902 Dollfus
& Dautzenberg, p. 318: Cardium speluncense Almera
& Bofil. Syn of Cardium multicostatum. ··1983 Anaya:
"Cardium" speluncense Almera y Bofill, 1894.
Sphaerocardium Coen, 1933 (genus). - nomen null. =
Acanthocardia Gray, 1851. Cardiidae. Recent. Type
species: Cardium paucicostatum Sowerby, 1841 (non
Deshayes, 1838, SD Keen, 1937). REcORDs. ··1937 ZR:
Sphaerocardium (subgen. sp. Encardium sic) Coen,
1933. Type here designated Cardium pancicostatum
sic Sowerby, 1841 Probably = Acanthocardia Gray,
1851 [Ref. Keen 1937]. ··1969 Moore et al.:
Sphaerocardium = Acanthocardia Gray, 185 I. ··1980
Vokes: Sphaerocardium (genus) Coen, 1933 in
subfamily Cardiinae.
sphaeroidea Forbes, 1845. - Changed combination:
Protocardia sphaeroidea (Forbes, 1845). Cardiidae.
Lower Cretaceous. TYPE LOCALITY. In Europe. REcORDs.
··1871
Stoliczka, 13: C. sphaeroideum
(? Laevicardium). ··1922-32 Sherborn, p. 606 I:
Cardium sphaeroideum Forbes, 1845 ... I 965 ZR:
Cardium sphaeroideum Forbes, 1845, Aptian, W.
Europe and Japan. Subgenotype of (Globocardium)
[Ref. Hayami 1965]. ··1965 ZR: Protocardia
(Globocardium) sphaeroideum Forbes, 1845. "1973
Keen: Protocardia (Globocardium) sphaeroidea
(Forbes, 1845). Lower Cretaceous in Japan; type
locality in Europe.
sphaeroideum (mode). - See under sphaeroidea.
spiekeri Hanna & Israelsky, 1925. - Changed
combination: Trigoniocardia spiekeri Hanna and
Israelsky, 1925. Cardiidae. Pliocene. RECORDS . •• I 925
Hanna & Israelsky: Cardium (Trigoniocardia) spiekeri
(new name for affinis Nelson). ··1941 Pilsbry &
Olsson: Cardium (Trigoniocardia) spiekeri Hanna &
Israelsky, 1925 (new name for affinis Nelson)
[Pliocene]. ··1958 Abbott: Trigoniocardia spiekeri
Hanna and Israelsky.
spillmani Conrad, 1858. - Changed combination:
Protocardia spillmani (Conrad, 1858). Cardiidae.
Cretaceous. GENOTYPE of Pachycardium Conrad. TYPE
LOCALITY. Mississippi N. America. RECORDS. ··1869
Conrad: Pachycardium spillmani Conrad. ··1870
Conrad: Pachycardium spillmani Conrad (Mississippi).
•• I 871 Stoliczka: Card. spillmani, Conr. •• I 876 Meek:
Cardium (Laevicardium)spillmani Conrad is type of
Pachycardium Conrad . •• I 887 Fischer: Pachycardium
spillmani Conrad. ··1902 Dall: Cardium spillmani
Conrad; Cretaceous, type of section Pachycardium
Conrad, 1870. ··1980 Ruhoff: Cardium (Laevicardium)
spillmani Conrad, 1858; vol. 3: 326, pI. 34, fig. 3.
··1980 Ruhoff: Pachycardium spillmani Conrad, 1869:
vo 1. 5: 96. ··1998 Rindsberg: Protocardia
(Pachycardium) spillmani (Conrad, 1858).
spinalalum Rowley. - Conocardium spinalatum Rowley.
Conocardiidae. Fossil. RECORDS . •• I 960 ZR:
Conocardium spinalatum Rowley description and
figures of the holotype, metatype and paratype. [Ref.
Branson I 960a].
spinicosta Mayer, 1857. - Combination to be revised:
"Cardium" spinicosta Mayer, 1857. Cardiacea. Fossil.
RECORDS. ··1980 Ruhoff: Cardium spinicosta Mayer,
1857; [1856-1857] vol. 5; 361.
spinosa (mode and variety). - See under spinosum
Lightfoot in Solander, 1786. - procarpatina var.
spinosa Jekelius, 1944. - tuberculatum var. spinosa.
spinosifrons Vokes, 1984. - Agnocardia spinosifrons
Vokes, 1984. TYPE MATERIAL. Holotype 377405: United
States National Museum. Washington, DC, USA
(Schneider 2002). Cardiidae. Fossil. RECORDS. ··1984
ZR: Agnocardia spinosifrons sp. nov. Florida, Middle
Pliocene. Ref. to Vokes p. 43 ... 2002 Schneider:
Agnocardia spinosifrons Vokes, 1984.
*spinosum. - Homonymy to be noticed. Cardium
spinosum from 3 sources: Lightfoot in Solander, 1786.
- Nardo in Chiereghini, 1847. - Sowerby, 1804.
spinosum Nardo in Chiereghini, 1847. - Combination
to be revised: "Cardium" spinosum Nardo in
Chiereghini, 1847. ··1922-32 Sherborn, p. 6083:
Cardium spinosum Nardo (ex Chiereghini), 1847.
spinosum Lightfoot in Solander, 1786. [Figs. Part 1 p.
58 No.3, p. 68, 89]. - Changed combination:
Acanthocardia spinosa (Lightfoot in Solander, 1786).
OTHER JUDGEMENTS: Acanthocardia aculeata (Linne,
1758). - Acanthocardia erinacea (Lamarck, 18 I 9). »
Regarding Cardium spinosum: Cardiidae. Recent.
RECORDS. ··1786 Solander, p. 16 no. 344. Cardium
spinosum S. ··1786 Lightfoot in Solander: Cardium
spinosum . •• I 8 I 7 Dillwyn: Cardium spinosum
Solander's MSS . •• I 922-32 Sherborn, p. 6083:
Cardium spinosum Solander, 1786. ··1945 Smith:
Acanthocardia spinosa Solander = Cardium spinosa
Solander 1786 . •• I 967 Rehder: Portland Catalogue No.
2297. Cardium spinosum S. Figured in Favanne 52.
A2. ··1967 Rehder: Cardium spinosum (Lightfoot,
1786), p. 105 = Acanthocardia echinatum (Linne,
1758) ... 1967 Rehder: Cardium spinosum Dillwyn,
1817, p. 115 = Acanthocardia echinatum (Linne, 1758) .
•• I 969 Nordsieck: Acanthocardia spinosa (Sol ander,
1786). •• I 969 Nordsieck: Cardium spinosum (Solander,
1786) = Acanthocardia erinaceum (Lamarck, 1819).
··1977 Fischer-Piette: Cardium (Cerastoderma,
A canthocardia) spinosum So lander. ··1979 ZR:
Acanthocardia spinosa (Lightfoot 1786). Ref. to
Mienis, p. 912. ··1998 Macedo et at.: Acanthocardia
spinosa (Solander, 1786). ··200 I Vidal, p. 407:
Acanthocardia spinosum. - CHARACTERS AND
Phuket Marine Biological Center Special Publication vol. 30 (2004)
775
spinosum - spiridionis
OCCURRENCE - DESCRIPTIONS. "2001 Vidal: With a
few additional larger tentacles outside siphons. No eyes
on tentacles. REMARKS. "1969 Nordsieck: the species
erinaceum Lamarck, 1819 is a junior synonym of
spinosum Solander, 1786. HABITAT. "1969 Nordsieck:
Zone of laminarians. DISTRIBUTION. "1969 Nordsieck:
C. spinosum: Mediterranean, Lusitanean area. C.
aculeata: Portugal, West Europe, Mediterranean, West
Africa.
spinosum Meuschen, 1778. Non binominal. - Not
available. » Regarding Coeur spinosum: replaced by
Papyridea soleniformis (Bruguiere, 1789). Cardiidae.
Pleistocene and Recent. Swainson (1840) designated
spinosum (citing Meuschen, 1778) type species of the
genus Papyridea. TYPE LOCALITY. Jamaica. RECORDS.
"1778 Meuschen, p. 442: Coeur spinosum. "1840
Swainson: spinosum (citing Meuschen). "1858 Adams
& Adams, p. 456: Cardium (Papyridea) spinosa citing
Meusch. "1890 Paetel: Papyridea spinosa Meusch.
"1902 Dall: Cardium (Papyridea) spinosum citing
Meuschen, 1778 [Pleistocene]. "1902 Dall: Cardium
spinosum citing Meuschen, type of subgenus Papyridea
Swainson. "1902 Dall & Simpson: Cardium spinosum
citing Meuschen. "1902 Sherborn, p. 920: Cardium
spinosum Meuschen, 1778. "1921 Maury: Cardium
(Papyridea) spinosum citing Meuschen, 1778
[Pleistocene to present]. "1934 Johnson: Papyridea
spinosum citing Meuschen 1778. (c. bullata of
authors). "1936 McLean: Papyridea spinosum citing
Meusch. '·1939 McLean: Papyridea spinosum (citing
Meuschen, 1778). "1939 ZR: Papyridea spinosum
Meuschen [non binorn.] [Ref. McLean 1939]. "1951
McLean: Cardium spinosum citing Meuschen =
Papyridea hiatus (citing Meuschen). "1961 Olsson:
P hiatus citing Meuschen. '·1974 Abbott: Cardium
spinosum citing Meuschen, 1778 = Papyridea
soleniformis (Bruguiere, 1789). "1977 Fischer-Piette:
Cardium spinosum citing Meuschen = Laevicardium
soleniforme Bruguiere . - CHARACTERS AND
OCCURRENCE - OF COLLECTIONS. ··ZMUC: 5 shells
from Florida Keys coli. Wolff 1873. "ZMUC No. 676 :
corresponds with bullata Chemnitz, Lam No. 10, 50
leviforme Bruguiere. Yo1di coil. 40 I, the Antilles,
ZMUC I specimen from Tortola, Antilles. "ZMUC: 8
shells from the Antilles, label indicates bullatum =
soleniforme = spinosum; "ZMUC wet collection :
Antilles; Nah Trang, Vietnam, very strong spines, looks
different. DESCRIPTIONS. "1939 McLean: Ribs 46-47.
DIMENSIONS . .. 1939 McLean: 41 x 31 x 20 mm.
DISTRIBUTION. North Carolina to Brazil.
spinosum Sowerby, 1804. - Misidentification:
Acanthocardia aculeata (Linnaeus, 1758). OTHER
JUDGEMENTS : Cardium echinatum Linnaeus . »
Regarding Cardium spinosum of Sower by: Cardiidae.
Recent. RECORDS . "1804 Sowerby, p. 65, pI. 32:
Cardium spinosum. "1807 Maton & Rackett, No.4:
Cardium spinosum Sowerby. Coast of Devonshire.
"1856 Wood & Hanley: C. spinosum Sow. Brit. Misc.
PI. 32 = Cardium echinatum Lin. "1896 Elera: Cardium
spinosum Sowerby. "1922-32 Sherborn, p. 6083:
Cardium spinosum Sowerby, 1805. "1997 Kafanov:
Cardium spinosum in J. Sowerby, 1804 p. 65, pI. 32not syn. of Papyridea (Papyridea) aspersa Sowerby
in Broderip et Sowerby, 1833.
spinosum of authors. - Misidentification: Papyridea
aspersa Sowerby. Cardiidae. Recent. RECORDS. "1997
Kafanov: Cardium spinoswn auctt. - in part syn. of
Papyridea (Papyridea) aspersa Sowerby in Broderip
et Sowerby, 1833.
spinosum Chemnitz (1782). Non binominal. - Not
available. Misidentification. Cardium echinatum Linne.
Cardiidae. Recent. RECORDS. "1867 Weinkauff:
Cardium spinosum Chemnitz = Cardium eehinatum
Linne.
spinosum of Dillwyn 1817. - Misidentification: Cardium
aeulealum Wood. Cardiidae. Recent. RECORDS. "1856
Wood & Hanley: Cardium spinosum Dillw. 115 =
Cardium aculeatum Wood (not of authors). "1922-32
Sherborn, p. 6083: Cardium spinosum Dillwyn (ex
So lander MS), 1817. "1997 Kafanov: Cardium
spinosum in Dillwyn, 1817 p. 115, pI. 32 - not syn. of
Papyridea (Papyridea) aspersa Sowerby in Broderip
et Sowerby, 1833.
spinosum of Link (1806). - Misidentification:
Acanthocardia aculeata (Linne, 1758). Cardiidae.
Recent. RECORDS. "1806 Link: Cardium spinosum =
Cardium aculeatum [Chemnitz. figs . 155, 156]. "1807
Link: Cardium spinosum Sowerby, 1805 =
Acanthocardia aculeatum (Linne, 1758) [fide Rehder
1967]. "1922-32 Sherborn, p. 6083 : Cardium spinosum
Link, 1807. "1979 Richardson et al.: Cardium
spinosum in Link (1807) pt. 3, p. 151 = Acanthocardia
aculeata (Linne, 1758).
spinosum of Maton & Rackett (1807). Misidentification: Acanthocardia echinata (Linne,
1758). OTHER JUDGEMENTS: Acanthocardia aculeata
(Linnaeus, 1758). » Regarding Cardium spinosum:
Cardiidae. Recent. RECORDS. "1807 Maton & Rackett:
Cardium spinosum Sowerby. "1890 Paetel: Cardium
spinosum Mat. = echinatum L. "1903 Hidalgo, p. 343:
Cardium spinosum Maton = C. echinatum Linne.
"1905 Hidalgo: Cardium spinosum Maton. "1928
Faustino: Cardium spinosum Maton. "1977 FischerPiette: Cardium spinosum Maton = Cardium aculeatum
Linne.
spinosum of Sowerby (1859) . - Misidentification of
Acanthocardia echinata (Linne, 1758). Cardiidae.
Recent. RECORDS. "1859 Sowerby: C. spinosum =
Cardium echinatum Linn. "1977 Fischer-Piette:
Cardium spinosum in Sowerby 1859 = C. echinatum
Linne.
spinosum aspersum Sowerby, 1833 (variety). Upgraded. Changed combination: Papyridea
soleniformis aspera (Sowerby, 1833). Cardiidae.
Recent. RECORDS. "1901 Dall: Cardium (Papyridea)
spinosum Meuschen, 1797 var. aspersum Sowerby,
1833 . .. 1902 Dall: Cardium spinosum var. asperum
Sowerby. "1909 Lamy: Cardium (Papyridea)
spinosum Meuschen var. asperum Sowerby, Dall .
"1961 Olsson : Cardium spinosum var. aspersum
Sowerby, Dall, 1900 = Papyridea soleniformis aspera
(Sowerby, 1833).
spinosum turtoni (variety) Dall. - Synonym of
Papyridea soleniformis (Bruguiere, 1789). »
Regarding Cardium spinosum var. turtoni: Cardiidae.
Recent. RECORDS. "1902 Dall: Cardium spinosum var.
turtoni Dall = Cardium spinosum Meuschen, 1778.
spiridionis Pavlovic, 1929. - New combination :
776
Tropical Marine Mollusc Programme (TMMP)
spolongense - squamosum
Lymnocardium spiridionis Pavlovic, 1928.
Lymnocardiidae. Pontian. RECORDS. --1928 ZR :
Limnocardium spiridionis [Ref. Pavlov itch 1928].
--1929 ZR: Limnocardium spiridionis Pavlovic, 1929
sp. n. Pontian Serbia .
spolongense Martin, 1916. - Combination to be revised:
"Cardium" spolongense (Martin, 1916). Cardiidae.
Lower Miocene. TYPE LOCALITY. Lower Miocene of
West-Progogeb. Java, Indonesia. TYPE REPOSITORY .
?Geologischen Reichs-Museum, Leiden , the
Netherlands. RECORDS. --1917 ZR: Cardium
sp%ngense Martin, 1917 n. sp. Miocene Java. --1919
Martin: Cardium spolongense Martin, J 916. "1999
Vidal: cites Martin 1916, 266, pI. 4, figs 107-109:
Cardium (Trachycardium) sp%ngense.
spondyloides Hauer, 1847. - Changed combination :
Discors spondy loides (Hauer, 1847). Cardiidae .
Miocene. RECORDS. "1922-32 Sherborn, p. 6106 :
Cardium spondyloides Hauer, 1847. "1999 Jacques
Vidal (pers. com.) MNHN, Paris: Fossils from the
Paleontological Collection. Discors spondyloides.
Helvetien. France (Touraine), B27156.
spondyloides. - Identical with Discors semisulcatus
(Sacco, 1899, p. 54). RECORDS. --1970 Glibert & Poel :
Cardium spondyloides, not Hauer in Dollfus &
Dautzenberg 1902-1920 = Discors semisulcatus
(Sacco, 1899, p. 54).
spondyloides (cf.) - Combination to be revised:
"Cardium" cf. spondy/oides. Cardiidae. Miocene.
RECORDS. --1938 ZR : Cardium cf. spondyloides
Miocene, Lybia figs. [Ref. Alberichi & Mauroner
1938].
spondylopsis Almera et Bofill, J 894 . - Didacna
spondylopsis (Almera et Bofill, J 894). Lymnocardiidae.
Fossil. RECORDS. --1987 Anaya: Cardium spondylopsis
Almera et Bofill, 1894. --1987 Anaya: Didacna (s. s.)
spondylopsis (Almera et Bofill, 1894).
spratti Fuchs in Capellini J879. Lymnocardiidae. Fossil.
RECORDS. --1970 ZR: Cardium spratti Fuchs in
Capellini 1879, see Pontalmyra equilateralis.
spurium d 'Orbigny, 1850. - Combination to be revised :
"Cardium" spurium d'Orbigny, 1850. ?Cardiidae.
Fossil. RECORDS. --1922-32 Sherborn, p. 6110: Cardiola
spurius Muenster, 1840. "1922-32 Sherborn, p. 6110:
Cardium (Muenster) spurium d'Orbigny, 1850. Cardiola.
squalidum (mode). - See under squalidus Yokoyama,
1924.
squalidus Yokoyama, 1924. - Changed combination:
Serripes squalidus (Yokoyama, 1924). Cardiidae.
Upper Oligocene. RECORDS . --1924 Yokoyama :
Cardium squalidus (fide Keen 1973). --1924 ZR :
Cardium (Laevicardium) squalidum Yokoyama, 1924
sp. n. Tertiary Japan. --1973 Keen: Serripes squalidus
(Yokoyama, 1924).
squamata (subspecies). - See under quadripartitum
squamata Andrus.
squamosa Lamarck. [Figs. Part I p. 148]. - Tridacna
squamosa Lamarck. Tridacnidae. Recent. RECORDS.
--1845 Chenu, pis. 3, 6: Tridacna squamosa Lamarck.
"1868 ZR: Tridacna squamosa [Ref. Kiister 1868].
--1869 ZR: Tridacna squamosa. Specimens from Japan,
[Ref. Lischke 1869]. --1880 ZR: Tridacna squamosa
(Lam.) common on the reefs of North-east Australia,
attached to or imbedded in the coral rock, variable in
the color of the mantle [Ref. Woods 1880]. --1932 ZR:
Tridacna squamosa [Ref. Lamy 1932]. "1935 ZR:
Tridacna squamosa fig. [Ref. Nomura & Zinbo 1935].
"1947 ZR: Tridacna squamosa Lk. [Ref. McLean
1947]. --1950 ZR: Tridacna squamosa morphological
and histological characteristics [Ref. Mansour 1950].
--1956 ZR: Tridacna squamosa spawning habits [Ref.
Wada 1956]. --1965 ZR: Tridacna squamosa aging and
duration of life [Ref. Haranghy et al. 1965]. "1965
ZR: Tridacna squamosa described and figd. range,
habitat and synonymy discussed [Ref. Rosewater
1965]. "1937 ZR : Tridacna squamosa auct. of
Flodacna "group" [Ref. Ireda1e 1937]. "1974 ZR:
Tridacna squamosa. Shell. Calcification. Trochosphere.
Metamorphosis. Ref. to LaBarbera. "1975 ZR :
Tridacna squamosa Gametes, spawning, development,
lava. Postlarva. Ref. to La Bareera. --1981 ZR:
Tridacna squamosa. Ref. to Beckvar. Caroline Islands.
/ Fitt et al. Pacific Ocean islands. --1976 ZR: Tridacna
squamosa. Crustacean parasites. Anthessius alalus.
Molucca Strait. Ref. to Humes. --1979 ZR: Tridacna
squamosa Ref. to Khristoforova et af. / Uhlenbruck et
al. --1983 ZR: Tridacna squamosa. Ref. to
Khristoforova et aI. , Mangum et al. --1999 Aungtonya
et al., PMBC-SP 19(2): 378: Tridacna (Chametra chea)
squamosa Lamarck, 1819. --2000 Lutaenko, p. 372:
Tridacna squamosa Lamarck, 1819. --2002 Hylleberg
& Kilburn, p. 64: Tridacna squamosa Lamarck, 1819.
--2003 Hylleberg & Kilburn , p. 186: Tridacna
squamosa Lamarck, 1819. - CHARACTERS AND
OCCURRENCE - DISTRIBUTION. --1999 Aungtonya et
al.: Thailand, Andaman Sea. --2000 Lutaenko:
Vietnam. --2002 Hylleberg & Kilbum: South India.
--2003 Hylleberg & Kilburn: Vietnam.
squamosum Gmelin, 1791 . [Figs. Part 1 p. 48 . Part 3 p.
922 (comparison)]. - Uncertain status. » Regarding
Cardium squamosum: ? = Trachycardium isocardia
(Linne, 1758). Cardiidae. Recent. TYPE LOCALITY. --1845
Reeve: unknown. RECORDS. --1742 Gualtieri. ID by
Gme1in 1791: Cardium squamosum t. 71. f. N . "1791
Gmelin, 3256, No. 44: Cardium squamosum . .. 1815
Wood : Cardium squamosum Gmelin p. 3256 is a young
C isocardia. --1817 Dillwyn: Cardium squamosum
Gmelin = Cardium isocardia Linnaeus. --1844 Potiez
& Michaud: Cardium squamosum Gmel. = Cardium
isocardia Linn. "1902 Sherborn, p. 925 : Cardium
squamosum Gmelin, 1790 sic. --1930 Stewart: Cardium
squamosum Gmelin. --1944 Clench & Smith: Cardium
squamosum. --1977 Fischer-Piette: Cardium
squamosum Gme1in = species insufficiently known. CHARACTERS AND OCCURRENCE - REMARKS.
"1742 Gualtieri, pI. 71, fig. N shows 2 species, both
labeled N. The one to the left is Cfragum Linne (1758,
No. 67). The one to the right could be either Cardium
isocardia Linne (1758, No. 66) or Cardium consors
Sowerby. Clench & Smith (1944) states that ifit should
be decided that figure N is that of C consors Sowerby,
the name squamosum would take priority and would
replace consors.
squamosum Sandberger, 1842. - Conocardium
squamosum Sandberger, 1842. Conocardiidae. Fossil.
RECORDS. --1922-32 Sherborn: Conocardium
squamosum Sandberger, 1842.
Phuket Marine Biological Center Special Publication vol. 30 (2004)
777
squamulosa - stenoplura
squamulosa Stevanovic, 1951. - Didacna squamulosa
Stevanovic, 1951. Lymnocardiidae. Pontian. RECORDS.
--1953 ZR: Didacna squamulosa Stevanovic, 1951 sp.
nov. Pontian Serbia.
squamulosum Deshayes, 1838. - New combination:
Lymnocardium squamulosum (Deshayes, 1838).
Lymnocardiidae. Fossil. RECORDS. --1886 Fischer:
Cardium squamulosum Deshayes. --1907 Andrussow:
Limnocardium squamulosum Desh. --1922-32
Sherborn, p. 6122: Cardium squamulosum Deshayes,
1838.
stampinense Stanislas-Meunier, 1880 . - Fragum
stampinense (Stanislas-Meunier, 1880). Cardiidae .
Oligocene. RECORDS. --1986 Gitton et al.: Fragum
stampinense Meun. Oligocene. --1970 Glibert & Poel,
p. 59: Fragum stampinense (Stanislas-Meunier, \880).
"1999 Jacques Vidal (pers . com.) MNHN , Paris :
Fossils from the Paleontological Collection. Cardium
stampiense S. Meunier. Stampien. France (Bassin de
Paris), B52836.
stanae Brusina, 1893. - New combination :
Lymnocardium stanae Brusina, 1893. Lymnocardiidae.
Fossil. RECORDS. --1979 ZR : Limnocardium stanae
Brusina, 1893. Neotype. Ref. to Stevanovic, p. 337.
stantoni Wade, 1926. - Changed combination :
Protocardia stantoni (Wade, 1926). Cardiidae.
Cretaceous. RECORDS. --1926 ZR: Cardium stantoni
Wade, 1926 sp. n. Cretaceous U.S.A. --1998 Rindsberg:
Proto cardia (Pachycardium) stan toni (Wade, 1926).
starobogatovi Kafanov & Savizky, 1982 . Keenocardium starobogatovi Kafanov & Savizky,
1982. Cardiidae. Oligocene. RECORDS. --1982 . Kafanov
et Savizky: Keenocardium starobogatovi Kafanov &
Savizky. --1982 ZR: Keenocardium starobogatovi
Kafanov, 1982 sp. nov. Russian SPSR. Ref. to Kafanov
et al. p. 40.
statonis. - Paradacna slatonis. Lymnocardiidae. Fossil.
"1977. Popov: Paradacna statonis.
stefafanescoi (error). - Printer's mistake for stefanescoi
in Fischer 1886 p. 218.
stefanescoi Tournouer. - Changed combination:
Prosodacna stefanescoi (Tournouer, 1879).
Lymnocardiidae. Miocene. TYPE LOCALITY . Josseni,
Roumania. RECORDS. --1879. Toumouer. vol. 28, p. 263:
Cardium stefanescoi. "1886. Fischer: Prosodacna
stefanescoi Toumouer, 1879. "1999. Jacques Vidal
(pers. com.) MNHN , Paris : Fossils from the
Paleontological Collection as Prosodacna stefanescoi
Tournouer. Meotien. Rumania, B35686, B3581 \.
steindachneri Brusina. - Caladacna steindachneri
(Brusina). Lymnocardiidae. Fossil. RECORDS. "1953.
Celidje: Caladacna steindachneri Brus. --1977. Popov:
Caladacna steindachneri (Brus.).
steindachneri (cf.) Brusina. - RECORDS. --1953. Celidje:
Caladacna cf. steindachneri Brus.
steindachneri euxinica Eberzin, 1956. - Caladacna
steindachneri euxinica Eberzin, 1956. Lymnocardiidae.
Pliocene. RECORDS . --1956. ZR: Caladacna
steindachneri euxinica Eberzin, 1956 subsp. nov.
Pliocene, Kerchenski, Crimea Peninsula, Russia.
stellatum Reeve, 1845. [Figs. Part 1 p. 143. Part 3 p.
916]. - Parvicardium stella tum (Reeve, 1845). OTHER
JUDGEMENTS: Cardium exiguum Gmelin, 1791. Eucardium exiguum var. subangulata Scacchi. -
Cerastoderma exiguum subangulatum Scacchi. Parvicardium siculum (Sowerby, 1839). - Cardium
pygmaeum Donovan, 1799. « Regarding Cardium
stellatum: Cardiidae. Recent. TYPE LOCALITY. "1845
Reeve: unknown. TYPE REPOSITORY. BMNH: Cardium
stellatum Reeve 1 of3 syntypes (Reeve's Figure) reg.
no. 1975620. Neg. No. 0233. RECORDS. "1845. Reeve:
Cardium stellatum. "1867. Weinkauff: Cardium
stellatum = Cardium exiguum Gmelin. --1868. ZR:
Cardium stellatum (Reeve) = Cardium exiguum
(Gmel.) [Ref. Romer 1868]. "1888. Kobelt: Cardium
stellatum Reeve = Cardium exiguum Gmelin. "1890.
Paetel: Hemicardium stella tum Rv. "1892. Bucquoy,
Dautzenberg & Dollfus: Cardium stella tum Reeve =
Cardium exiguum Gmelin. --1962. Grossu : Cardium
stellatum Reeve 1844 = Cardium (Parvicardium)
exiguum Gmelin, 1790 ... 1922-32. Sherborn, p. 6133:
Cardium stella tum Reeve, 1845. --1933 . Coen:
Cardium stellatum Reeve = Eucardium (Parvicardium)
exiguum var. subangulata Scacchi. "1937. Coen :
stella tum Reeve = Cerastoderma (Parvicardium)
exiguum subangulatum Scacchi . --1965 . Mars :
Cardium stella tum Reeve = Cardium exiguum Gmelin
"1969. Nordsieck: Cardium stellatum Reeve, 1845 =
Parvicardium exiguum exiguum (Gmelin , 1791)
identical with C. pygmaeum Donovan, 1799 and C.
siculum Sowerby, 1839. ,,1977. Fischer-Piette :
Cardium stellatum Reeve. - CHARACTERS AND
OCCURRENCE - DESCRIPTIONS. --1845 Reeve: shell
obliquely quadrate, gibbous, very inequilateral, anterior
side very short, posterior angulated; radiately flatly
ribbed, 23-24 ribs, finely crenulated on each side,
anterior ribs noduliferous; rusty brown, starred with a
few white spots. REMARKS. --1845 Reeve: a square
gibbous shell of which the anterior side is remarkably
short and contracted. HABITAT. --1845 Reeve: unknown.
Stenodacna Andrussow
1923
(genus).
Lymnocardiidae. Fossil. RECORDS. --1962 . ZR:
Stenodacna Andrussov in subfam. Didacninae Eberzin,
1962, family Cardiidae [Ref. Eberzin 1962]. "1963 .
ZR: Stenodacna Andrusov, historical survey of genus
and related species [Ref. Akhvlediani 1963] . "1977.
Popov: Stenodacna Andrussow, 1923 . "1980. Vokes:
Stenodacna Andrussow, 1923 in subfamily Didacninae.
"1987. Taktakishvili, p. 87: Stenodacna Andrussow
1923; referred to new tribe Pontalmyrini .
stenopleura Brusina & Brusina, 1902 . - New name
Prosodacna osijeki Basch, 1990. Lymnocardiidae.
Pliocene. RECORDS. "1961. ZR : Prosodacna
stenopleura Dacian [Ref. Macarovici 1961]. "1987.
Lutskanov & Tcholakov: Prosodacna (Prosodacnomya) stenopleura. "1987. Lutskanov & Tcholakov:
Prosodacna (Prosodacnomya) stenoplura sic. --1990.
Basch, p. 65 : Limnocardium stenopleura Brusina &
Brusina, 1902; preoccupied name replaced by
Prosodacna (Prosodacnomya) osijeki.
stenopleura stllriensis (variety) Macarovici, 196\. Upgraded. Prosodacna stenopleura Brusina & Brusina,
1902. Lymnocardiidae. Pliocene. RECORDS. --1961. ZR:
Prosodacna stenopleura var. sturiensis Macarovici,
1961 nov. Dacian, Fersampenuazul Mare (Nou)
Cetatea-Alba.
stenoplura (error). - Printer's mistake for stenopleura
in Lutskanov & Tcholakov 1987.
778
Tropical Marine Mollusc Programme (TMMP)
stevanovici - striatulum
stevanovici Basch, 1990. - New combination: Lymnocardium stevanovici Basch, 1990. Lymnocardiidae.
Miocene. TYPE LOCALITY. Tal des Flusses Krapina
(Gebiet von Hrvatsko Zagorje) Nordwestkroatien,
Yugoslavia. RECORDS. 001990. Basch, p. 7, fig.:
Limnocardium (Zagrabicardium) stevanovici sp. nov.
stigmaticum (subspecies). - See under hungerfordi
stigmaticum Pilsbry, 1904.
sti/pnaulax Cossmann, 1886. - Changed combination:
Bucardium stilpnaulax (Cossmann, 1886). Cardiidae.
Eocene. RECORDS. 001886. Cossmann, p. 166, pI. VIII,
fig. 5-6: Cardium stilpnaulax nov. sp. 001977. Popov:
Bucardium (Europicardium) stilpnaulax (Cossmann,
1886). 001995. Le Renard et at.: Cardium (Orthocardium) stilpnaulax Cossmann, 1886. 00 1995. Pacaud
& Renard: Cardium (Orthocardium) stilpnaulax.
stiriacum Rolle, 1860. - Combination to be revised:
"Cardium" stiriacum Rolle, 1860. Cardiidae. Fossil.
RECORDS. 001980. Ruhoff: Cardium stiriacum Rolle,
1860; vol. 42: 275, fig. 6.
stiriatum Brown & Pilsbry, 1911. - Combination to be
revised: "Cardium" stiriatum Brown & Pilsbry, 1911.
Cardiidae. Oligocene. REcORDs. 001911. ZR: Cardium
(Trachycardium) stiriatum Brown & Pilsbry, 1911 sp.
n. Panama Oligocene.
stojadinovici Pavlovic, 1928. - New combination:
Lymnocardium stojadinovici Pavlovic, 1928
Lymnocardiidae. Pontian. RECORDS. 001928. ZR:
Limnocardium stojadinovici [Ref. Pavlovitch 1928].
001929. ZR: Limnocardium stojadinovici Pavlovic,
1929 sp. n. Pontian Serbia.
stonei Cragin, 1893. - Synonym of Protocardiajilosa
(Conrad, 1857). » Regarding Protocardia stonei:
Cardiidae. Cretaceous. RECORDS. 001986. Scott:
Protocardia stonei Cragin, 1893 = Protocardia jilosa
(Conrad, 1857). 001987 ZR: Protocardia stonei Cragin,
1893. Junior synonym of P.jilosa (Conrad 1857). Ref.
to Scott p. 1203.
stoosi Brusina. - New combination: Lymnocardium
stoosi (Brus.). Lymnocardiidae. Fossil. REcORDs. 001987
Taktakishvili: Limnocardium stoosi Brus.
straeleni Glibert 1945. - Synonym of Parvicardium
papillosum (Poli). OTHER JUDGEMENTS: Cardium
nodosum Mourlon 1880-1881. » Regarding C.
straeleni: Cardiidae. Miocene. REcORDs. 001945 Glibert:
Cardium (Parvicardium) straeleni. 001947. ZR:
Cardium (Parvicardium) straelini Gilbert, 1945 sp. nov.
= Cardium nodosum Mourlon 1880-1881 non Montagu,
1803, Miocene Anversian, Eocene, Belgium. 001958.
Sorgenfrei: Cardium straeleni Glibert. 001968.
Rasmussen: Cardium straeleni Glibert 1945 =
Parvicardium papillosum (Poli).
strangi Laws, 1930. - Changed combination:
Maoricardium strangi (Laws, 1930). Cardiidae. Fossil.
RECORDS. 001931. ZR: Cardium strangi Laws, 1931 sp.
n. Tertiary N.Z. 001977. Popov: Maoricardium strangi
(Laws, 1930).
strangulatum d'Orbigny, 1850. - Conocardium
strangulatum d'Orbigny, 1850. Conocardiidae. Fossil.
RECORDS. 001922-32. Sherborn, p. 6163: Cardium
strangulatum Koninck, 1841. 001922-32. Sherborn:
Conocardium (Koninck) strangulatum d'Orbigny,
1850. - Cardium.
stratonis Andrussow, 1907. - Abichia stratonis
Andrussow, 1907. Lymnocardiidae. Fossil. RECORDs.
001907. Andrussow: Abichia stratonis nov. sp. (no
description).
stratonis Vassoievitch, 1929. - Paradacna stratonis
Vassoievitch, 1929. Lymnocardiidae. Pliocene.
RECORDS. 001929. ZR: Paradacna stratonis
Vassoievitch, 1929 sp. n. Pliocene or Post Tertiary N.
Caucasus.
strauchi Glibert & Van De Poel, 1970. - Parvicardium
strauchi Glibert & Van De Poel, 1970. Cardiidae.
Eocene. RECORDS. 001970. ZR: Parvicardium strauchi
Glibert & Van De Poel, 1970 sp. nov. Auversian,
Eocene, Mary, Seine-et-Marne, France. 001995.
Pacaud & Renard: Parvicardium strauchi.
striata Martyn, 1784. Non binominal. - Not available.
» Regarding Cardium striata: identical with Cardium
costatum Linne, 1758. Cardiidae. Recent. REMARKS.
The figures by Martyn 1784 are excellent. However,
the locality China is wrong. The distribution of C.
costatum is limited to West Africa. RECORDS. 001784.
Martyn, p. 50, pI. 48, fig. 3, pI. 49. fig. 4: Cardium
striata.
striatissimum Kutassy, 1934. - Combination to be
revised: "Cardium" striatissimum Kutassy, 1934.
Cardiidae. Miocene. RECORDS. 001935. ZR: Cardium
striatissimum Kutassy, 1934 sp. n. Miocene Celebes.
striatopunctatum Munster, 1835. - Combination to be
revised: "Cardium" striatopunctatum Muenster, 1835.
?Cardiidae. Fossil. RECORDS. 001922-32. Sherborn, p.
6170: Cardium striatopunctatum Muenster, 1835.
striatula Sowerby 1829. - Protocardia striatula
(Sowerby 1829). Cardiidae. Upper Jurassic. RECORDS.
001982. Fursich, p. 78: Protocardia (Protocardia)
striatula (Sowerby 1829); senior synonym of
Protocardia concinna (von Buch 1844).
*striatulum. - Homonymy to be noticed. Cardium
striatulum from 5 sources: Brocchi, 1814. - Chiaje,
1830. - Sowerby, 1827. - Sowerby, 1834. - Phillips
(?year).
striatulum Beyrich, 1845. - Protocardia striatulum
Beyrich, 1845. Cardiidae. Fossil. RECORDS. 001922-32.
Sherborn, p. 6174: Protocardia (Sow.) striatulum
Beyrich, 1845. - Cardium striatulum.
striatulum Brocchi, 1814. [Figs. Part I p. 94]. -Changed
combination: Nemocardium striatulum (Brocchi,
1814). Cardiidae. Mio-Pliocene. TYPE LOCALITY. Italy,
Mio-Pliocene deposit. RECORDS. 001814 Brocchi:
Cardium striatulum. 001832 Bronn: Cardium striatulum
Brch. 507 (fide Keen 1950). 001857 Deshayes: Cardium
striatulum Brocchi, 1813 sic. 001899 Sacco, p. 56, PI.
12, figs. 18-23: Nemocardium striatulum (Brocchi,
1814).001922-32 Sherborn, p. 6172: Cardium
striatulum Brocchi, 1814. 001999 Jacques Vidal (pers.
com.) MNHN, Paris: Fossils from the Paleontological
Collection as Nemocardium striatulum Brocchi.
Oligocene. Germany, B31365, B36100.
striatulum Chiaje, 1830. - Combination to be revised:
"Cardium" striatulum d. Chiaje, 1830. Cardiidae.
Fossil. REcORDs. 001922-32 Sherborn, p. 6172: Cardium
striatulum d. Chiaje, 1830.
striatulum Phillips. - Combination to be revised:
"Cardium" striatulum Phillips. Cardiidae. Fossil.
REcORDs. 001947 ZR: Cardium striatulum Phillips [Ref.
Melmore 1947].
Phukel Marine Biological Center Special Publication vol. 30 (2004)
779
striatulum - striatum
striatulum Sowerby, 1827. - Combination to be revised:
"Cardium" striatulum Sowerby, 1827. Cardiidae.
Fossil. RECORDS. "1922-32. Sherborn, p. 6172:
Cardium striatulum J. De C. Sowerby, 1827. (Min.
Conch. VI, 101).
striatulum Sowerby, 1834. [Figs. Part I p. 109, 114, 135].
- Synonym of Nemocardium pulchellllm (Reeve, 1844).
OTHER JUDGEMENTS: Cardium substriatulum Orbigny.
» Regarding Cardium striatulum: Cardiidae. Recent.
TYPE LOCALITY. Australia, New z・。ャセョ、@
(Sowerby
1840). RECORDS. "1834 Sowerby, 49 t part, セGᄃ@
16:
Cardium striatulum nobis. "1840 Sowerby, l77t part,
fig. 45: Cardium striatulum jun. "1840 Sowerby, Proc.
Z.S. London p.105: Cardium striatulum. "1840
Sowerby, catalogue p. I, No.9: Cardium striatulum
Sow. "1840 Sowerby, corrected list, p. 8: Cardium
striatulum (Sowerby, No.9). "1841 Reeve: Cardium
striatulum Sowerby. "1843 Hanley: Cardium
striatulum Sowerby. "1844 Reeve, sp. 60: Cardium
striatulum Sowerby, 1840 (juvenile) = Cardium
pulchellum Reeve, 1844. "1857 Deshayes: Cardium
striatulum Sow. - C. substriatulum d'Orb. "1858
Adams & Adams, p. 457: Laevicardium striatulum
Sow. "1873 Martens: Cardium striatulum Sowerby.
"1873 Hutton: Cardium striatulum Sowerby. "1880
Hutton: Cardium striatulum Sowerby. "1884 Grasset:
Cardium striatulum Sow. "1884 Hutton: Cardium
striatulum Sowerby. "1885 Smith: Cardium striatulum
Sowerby. 1840. = Cardium (Eucardium) pulchellum
Gray. "1890 Paetel: Cardium striatulum Sow. "1901
Suter: Cardium striatulum Sow., 1840 (not ofBrocchi)
= Cardium pulchellum Gray. "1902 Hedley: C.
striatulum Sowerby 1841, fig. 45 = C. pulchellum Gray.
"1904 Hutton: Cardium striatulum Sowerby (non
Brocchi) = Cardium pulchellum Gray (non Reeve).
"1922-32 Sherborn, p. 6172: Cardium striatuillm G.
B. Sowerby, 1841. "1933 David: Protocardia
(Nemocardium) striatula Sow. "1977 Fischer-Piette:
Pratulum striatulum Sowerby. "1977 Wilson &
Stevenson: Nemocardium striatulum (Sowerby, 1841)
non Brocchi = pulchellum Gray, 1843. "1999 Jacques
Vidal (pers. com.) MNHN, Paris: Fossils from the
Paleontological Collection as Protocardium striatulum
Sowerby. Mesozoic. Madagascar, BIO064, BI0065.CHARACTERS AND OCCURRENCE - DESCRIPTIONS. "1840 Sowerby: found that the pink-striped
bands, which gave so much brilliancy to the young
shells, was scarcely to be traced in the older specimens.
"1844 Reeve: shell transversely ovate, rather thin,
posterior side a little gaping, radiately closely striated,
striae conspicuous, elevated, peculiarly cancellated
with very fine waved wrinkles, posterior striae covered
towards the margins with minute short spines; pale rusty
white, two orange-red rays diverging from within the
umbones of each valve. REMARKS. "1844 Reeve : a thin
delicate shell, slightly produced at the posterior side,
and having its entire surface cancellated with fine
irregularly waved wrinkles. "1844 Reeve: stated that
striatulum (Reeve 1844) differed significantly from
plllchellum (Reeve 1844). However, this difference has
been questioned by authors. "1977 Fischer-Piette:
claimed that Cardium striatulum Sowerby, 1834 =
Cardium pulchellum Gray, 1843 = Cardium thetidis
Hedley, 1902. DIMENSIONS. "1840 Sowerby: long. I.
poll. (length 25 mm),lat. 0.6 poll. (width 15 mm), all.
0.9 poll (height 23 mm). DISTRIBUTION. "1844 Reeve:
New Zealand. "1884 Grasset: N. Zealand.
striatulum thetidis Hedley 1902 (variety). [Figs. Part 3
p. 900]. - Upgraded. New combination: Nemocardium
striatulum thetidis (Hedley, 1902). » Regarding
Cardium striatulum Sow., var. thetidis: Cardiidae.
Recent. TyPE LOCALITY . Australia , New Zealand
(Sowerby 1840). RECORDS. "1902 Hedley: Cardium
striatllium Sow., var. thetidis nov. Hedley 1902. "1902
ZR: Cardium striatulum n. var. described by Hedley
1902. "1977 Wilson & Stevenson: Cardium striatulum
thetidis Hedley, 1902.
*striatum. - Homonymy to be noticed. Cardium
striatum from 9 sources: Wulff, 1790. - Wuleus, 1793.
- Spengler, 1799. - Schlotheim, 1813 . - Defrance,
1817. - Hoeninghaus, 1830. - Sowerby in Murchison,
1839. - Buvignier, J 843 . - Buckman in Murchison,
1844.
striatum Buckman in Murchison, J844. - Combination
to be revised: "Cardium " striatum Buckman in
Murchison, 1844. Cardiidae. Fossil. RECORDS. "1857.
Deshayes: Cardium striatum. Backmann. Spec. incog.
.. J922-32 Sherborn, p. 6178: Cardium striatum
Buckman in Murchison, 1844.
striatum Buvignier, 1843. - Replaced by "Cardium"
buvignieri Deshayes . » Regarding C. buvignieri:
Cardiidae. Fossil. RECORDS. ,,1857. Deshayes: Cardium
striatum Buvignier. - C. buvignieri Desh. "1922-32
Sherborn, p. 6178: Cardium striatum Buvignier, 1843.
striatum Defrance, 1817. - Identical with Cardium
multicostatum Brocchi. Cardiidae. Fossil. RECORDS.
"1844 Potiez & Michaud: Cardium striatum Defr. =
Cardium multicostatum Brocc. "1899 Sacco, p. 42: C.
striatum Defrance = Trachycardium multicostatum var.
miorotundata Sacco "1922-32 Sherborn, p. 6178:
Cardium striatum Defrance, 1817.
striatum Hoeninghaus, 1830. - See also under striatum
Schlotheim, 1813. RECORDS. "1922-32 Sherborn, p.
6178: Cardium (Schl.) striatum Hoeninghaus, 1830.Chamites .
striatum Koninck, 1842. - Combination to be revised:
"Cardium" striatum. Koninck 1842. Not in Cardiacea
(cardiomorpha). Silurian. RECORDS. "1857. Deshayes:
Cardium striatum Koninck (cardiomorpha). 1842.
striatum Munster, 1830. - Replaced by C. substriatum?
Miinster. Cardiacea. Fossil. RECORDS. "]857. Deshayes:
Cardium striatum Munster. 1830. -c. substriatum?
Munster.
striatum Munster, 1840. - Unknown species. ?Cardiacea.
Fossil. RECORDS. "1857. Deshayes: Cardium striatum
Munster, 1840. Spec. incog.
striatum Schlotheim, 1813. Not a cardiid. RECORDS.
"1857 Deshayes: Cardium striatum Schlotheim. Lima striata. "1922-32 Sherborn, p. 6178: Chamites
striatum Schlotheim, 1813.
striatum Sowerby in Murchison, 1839. - Combination
to be revised: "Cardium" striatum Sowerby in
Murchison, 1839. ?Cardiidae. Lias-Jurassic. RECORDS.
"1922-32 Sherborn, p. 6178: Cardium striatum
Sowerby in Murchison, 1839 [Silurian]. "1999 Jacques
Vidal (pers. com.) MNHN, Paris: Fossils from the
Paleontological Collection. Cardium striatum Sowerby.
Ludlow, Grande Bretagne, BO 1930. Rauracien. Meuse
780
Tropical Marine Mol/usc Programme (TMMP)
striatum - Styladacna
(France), S04454.
striatum Sowerby, 1836. - Replaced by "Cardium "
sowerbyanum d'Orb. » Regarding Cardium striatum :
Cardiacea. Fossil. REcoRDs. "1857 . Deshayes: Cardium
striatum. Sowerby. 1836. - C. sowerbyanum. d 'O rb.
striatum Spengler, 1799. [Figs. Part 3 p. 880]. - Synonym
of Fu/via australis (Sowerby, 1834). » Regarding
Cardium striatum: Cardiidae. Recent. TYPE LOCALITY.
South Seas [Pacific Ocean] (Spengler 1799). T YPE
REPOSITORY. ZMUC in Spengler coil. RECORDS. "1799
Spengler, p. 45, No . 29: Cardium striatum . "1902
Sherborn, p. 933 : Cardium striatum Spengler, 1799.
"1977 Fischer-Piette: Cardium striatum Spengler.
"1994 Vidal: Cardium striatum Spengler, 1799 =
Fu/via (Fu/via) australis (Sowerby, 1834). "1999
Knudsen & Hylleberg: Cardium striatum Spengler,
1799. - CHARACTERS AND OCCURRENCE - OF
COLLECTIONS. "ZMUC 2 specimens in Spengler coIl.
DESCRIPTIONS. "1799 Spengler: Testa cordata
aequiva/vi /ongitudinaliter striata, natibus /aevigatis
apice purpureis. "1799 Spengler: shell circular,
inflated, umbo central, posterior slope sharply keeled,
ribs terminate in teeth, closely fitting, shells not gaping,
central and anterior parts, very low ribs on the lower
half, upper half smooth, glossy. Colour outside white,
lower part dotted with red, upper part olive-green with
red-brown spots, umbo covered with dark red spot,
inside white. "1999 Spengler (1799) translated by
Knudsen & Hylleberg: this small cockle from the South
Sea is circular in outline, the umbo thus is placed in
the middle. From the middle to above the tip of the
umbo the bivalve is strongly vaulted. In the middle,
the front side is bent outward in a sharp crest and
covered by stripes. On the anterior end these stripes
have incised teeth, which interlock and tightly close
both valves with the remaining margin, which is neatly
set with small fine teeth. Both surfaces are set with
narrow, flattened stripes from the lower margin to the
middle of the valve. The rest of the bivalve is shining
smooth. In the lower part, it is white outside and strewn
with red spots. The upper part, together with the umbo,
is olive-green, ornamented with red-brown spots. Both
umbo tips are half covered by a dark spot. Inside the
cockle is white. Size: 1 inch in height and in breadth.
DIMENSIONS. "1799 Spengler: height and width 26 mm
(I Tom). REMARKS. Cardium striatum Spengler = Fu/via
australis identified by Vidal 1994.
striatum Wuleus, 1793. - Cardium striatum. Wuleus.
1793. Cardiidae. Living species, Recent. RECORDS.
"1857. Deshayes: Cardium striatum Wuleus, 1793.
especes vivantes.
striatum Wulff, 1790. - Cardium striatum Wulff, 1790.
Cardiidae. Fossil. RECORDS. "1899 Sacco, p. 42:
Cardium striatum Wulff, 1790 (oldest application of
the name C. striatum).
striatum beta Defrance, 1817 (variety). - Identical with
Cardium multicostatum Brocchi. Cardiidae. Fossil.
RECORDS. "1832 Bronn: Cardium striatum beta Df. =
Cardium mu/ticostatum Brch. 506.
striatum leonense Mansfield, 1932 (subspecies). Combination to be revised: "Cardium" striatum
/eonense Mansfield, 1932. Cardi idae. Miocene.
RECORDS ... 1938 ZR: Cardium (Trachycardium)
striatum /eonense Mansfield, 1932 subsp. n. Borrow
pit, nr. Jackson Bluff, Leon Co. Choctawhatchee
formation (Miocene) Florida.
stricklandi Morris & Lycett, 1852. - Combination to be
revised: "Cardium" strick/andi Morris & Lycett.
Cardiidae. Lias-Jurassic. RECORDS. "1980 Ruhoff:
Cardium stricklandi Morris & Lycett, 1852 [quoted
from Laube 1867, vol. 27(2): 41]. "1999 Jacques Vidal
(pers. com.) MNHN, Paris: Fossils from the
Paleontological Collection. Cardium strick/andi Morris
& Lycett. Bathonien. France (Pas de Calais), B 13069.
France (Pas de Calais), S01916.
strigilliferum Wood, 1856. - Combination to be revised:
"Cardium" strigilliferum Wood, 1856. Cardiidae.
Fossil. RECORDS. "1980 Ruhoff: Cardium strigilliferum
Wood, 1856 [1850-1856] vol. 2: 154, pI. 13, fig. 5a-d
strimonense Popov & Nevesskaya, 2000. Lymnocardium strimonense Popov & Nevesskaya,
2000. Cardiidae. Late Miocene. RECORDS. "2000 Popov
& Nevesskaya, downloaded abstract: Lymnocardium
(Hellenicardium) strimonense n. sp.
*strio/atum. - Homonymy to be noticed. Cardium
striolatum from 2 sources: Link, 1807. - Calcara, 1841.
strio/atum Calcara, 1841. - Combination to be revised:
"Cardium" strio/atum Calcara, 1841. Cardiidae. Fossil.
RECORDS . "1922-32 Sherborn, p. 6203: Cardium
strio/atum Calcara, 1841.
striolatum Link, 1807. - Combination to be revised:
"Cardium " strio/atum Link, 1807. Cardiidae. Fossil.
RECORDS. "1922-32 Sherborn, p. 6203: Cardium
strio/atum Link, 1807.
striolatum Orbigny, 1850. - Unicardium striolatum
d'Orbigny, 1850. Cardiidae. Fossil. RECORDS. "192232 Sherborn: Ma ctromya strio/atum Ag. "1922-32
Sherborn, p . 6204: Unicardium (Ag.) strio/atum
d 'Orbigny, 1850. - Mactromya striolatum.
stu/torum Linnaeus, 1758. [Figs. Part 1 p. 47]. - Not a
cardiid. » Regarding Cardium stu/torum: identical
with Mactra stu/torum. RECORDS. "1742 Gualtieri . ID
by Linnaeus 1758: Cardium stu/tontm t. 71 . f. C. "1758
Linnaeus (loth edition) No. 80: Cardium stultorum.
"1902 Sherborn, p. 940: Cardium stu/torum Linnaeus,
1758 [Mactra, ed. 12]. "1853 Forbes & Hanley, p. 362:
Cardium stultontm = Mactra stu/tontm.
stultorum ofSchroter, 1788. Not a cardiid. » Regarding
Cardium stultorum: identical with Mactra stultorum.
RECORDS. "1788 Schroter: Cardium stu/torum.
[Reference to Chemnitz (1782) fig . 226].
sturi Cobalcescu. - Prosodacnomya sturi (Cobalcescu).
Lymnocardiidae. Fossil. RECORDS . "1980 ZR:
Prosodacnomya sturi (Cobalcescu) Key to SUbspecies.
Ref. to Andreescu.
sturi orgrinensis Andreescu, 1980. - Prosodacnomya
sturi orgrinensis Andreescu, 1980. Lymnocardiidae.
Lower Pliocene. RECORDS. "1980 ZR: Prosodacnomya
sturi orgrinensis Andreescu, 1980 subsp. nov. Romania.
Ref. to Andreescu, p. 30.
sturiensis (variety). - See under stenopleura var.
sturiensis Macarovici, 1961.
sturzaiGillet, 1943. - New combination: Lymnocardium
sturzai Gillet, 1943. Lymnocardiidae . Pliocene.
RECORDS. "1946 ZR : Limnocardium sturzai Gillet,
1943 sp. nov. Pliocene Rumania.
Styladacna (error). - Printer 's mistake for Sty/odacna
in Dollfus, 1905 (Vokes 1980).
nl
Phuket Marine Biolog ical Center Special Publication vol. 30 (2004)
Stylodacna - subcarinata arcae/ormis
Stylodacna Sabba Stefanescu, 1896 (genus).
Lymnocardiidae. Tertiary. GENOTYPE. Styloda cna
heberti Cobal. (Dollfus 1905, commenting Andrussow
1903). Erected to accommodate Rumanian Pontian
[Tertiary b as in] cardiids. RECORDS . --1896 ZR:
Stylodacna Stefanescu, 1896 n. g. for Psilodon herbeti
Cobalcescu [Ref. Stefanescu 1896]. --1903 Andrussoff:
Styloda cna Sabba Stefanescu , 1896 ... 1974 ZR :
Stylodacna Sabba Stefanescu, 1896 referred to new
subfamily Prosodacninae. Ref. to Andreescu, p. 128.
--1980 Vokes : Stylodacna Stefanescu , 1896 in
subfamily Lymnocardiinae.
subacardo Andrussow, 1907. - Arcicardium subacardo
Andrussow, 1907. Lymnocardiidae. Tertiary. TYPE
LOCALITY. Crimean iron ore deposits. RECORDS. --1907
Andrussow, p. 114, Taf. IV, Fig. 4-9: Arcicardium
subacardo nov. sp. --1964 ZR: Arcicardium subacardo
Andrusov, 1964 sp. nov. sic Tertiary.
subaequilatera (variety). - See under muricatum var.
subaequilatera Roemer, 1849.
subalatum Andrzejovski , 1833. - Combination to be
revised : "Cardium" subalatum Andrzejovski, 1833 .
?Cardiidae. Fossil. RECORDS. --1922-32 Sherborn, p.
6219: Cardium subalatum Andrzejovski, 1833.
subalpina Orbigny, 1850. - Unicardium subalpina
d 'Orbigny, 1850. ?Cardiidae. Fossil. RECORDS. --192232 Sherborn : Tellina subalpina Munster. --1922-32
Sherborn, p. 6219: Unicardium (Muenst.) subalpina
d'Orbigny, 1850. - Tellina subalpina.
subalpinum Gumbel, 1861. - Combination to be revised:
"Cardium " subalpinum Gumbel, 1861. Cardiidae.
Fossil. RECORDS . --1980 Ruhoff: Cardium subalpinum
Gumbel, 1861 ; p. 743.
*subalternatum. - Homonymy to be noticed. Cardium
subalternatum from 2 sources: Orbigny, 1850. Jenkins, 1864.
subalternatum Jenkins, 1864. - Combination to be
revised : "Cardium " subalternatum (Jenkins, 1864).
OTHER JUDGEMENTS: Vasticardium flavum (Linnaeus,
1758) . » R ega rding Cardium subalternatum :
Cardiidae. Miocene. TYPE LOCA LITY. Miocene of Mount
Sela, on the Island of Java (Jenkins 1864). RECORDS.
--1864 Jenkins, p. 60, pI. 7, figs . 7a, b : Cardium
subalternatum spec. nov. --1925 Oostingh, p. 274:
Cardium subalternatum = Cardiumflavum Linne, 1758.
--1939 Adam & Lel oup: Cardium subalternatum
Jenkins = Laevicardium (Trachycardium) jZavum
(Linne 1758). --1980 Ruhoff: Cardium subalternatum
Jenkins, 1864; vol. 20 : 60, pI. 7, figs. 7a,b. --1999 Vidal
citing Jenkins 1864 : Cardium subalternatum.
Compared by its author with C. alternatum.
subalternatum Orbigny, 1850. - Combination to be
revised: "Cardium" subalternatum d ' Orbigny, 1850.
Cardiidae. Cretaceous. RECORDS. --1922-32. Sherborn,
p. 6219: Cardium subalternatum d ' Orbigny, 1850.
--1999. Jacques Vidal (pers . com .) MNHN, Paris:
Fossils from the Paleontological Collection. Cardium
subalternatum d ' Orbigny. Turonien. France (Loir and
Cher), S08378.
subalutaceum Falckenstein, 1911. - Combination to be
revised: "Cardium " subalutaceum Falckenstein, 1911.
Cardiidae. Senon . RECOR DS. --1911 ZR: Cardium
subalutaceum Falckenstein . 1911 sp. n. Senon
Germany. --1986 Abdel-Gaw a d: "Cardium"
subalutaceum Vogel von Falckenstein, 1911.
*subangulatum. - Homonymy to be noticed. Cardium
subangulatum from 2 sources: Scacchi, 1836. Orbigny, 1850.
subangulatum Orbigny, 1850. - Combination to be
revised: "Cardium " subangulatum d ' Orbigny, 1850.
Cardiidae. Fossil. RECORDS . --1922-32 Sherborn, p.
6220: Cardium subangulatum d ' Orbigny, 1850.
subangulatum Scacchi, 1836. [Figs. Part 3 p. 916]. No men nudum . OTHER JUDGEMENTS: Parvicardium
exiguum (GmeJin, 1791) . » Regarding Cardium
subangulatum: Cardiidae. Fossil. RECORDS . --1836
Scacchi, p. 7 (8): Cardium subangulatum. --1844
Philippi: Cardium subangulatum Scac . Uun.) =
Cardium exiguum Gm. --1851 Petit de la Saussaye:
Cardium subangulatum Scacchi. --1853 Doublier:
Cardium subangulatum (Scacchi). --1858 Gay:
Cardium subangulatum Scacchi. --1862 Weinkauff:
Cardium subangulatum Scacchi. --1867 Weinkauff:
Cardium subangulatum = Cardium exiguum Gmelin.
--1869 Romer: Cardium subangulatum Scacchi =
Cardium exiguum Gmelin. --1870 Aradas & Benoit:
Cardium subangulatum, Scacchi = Cardium exiguum,
Gmelin. --1880 Stossich : Cardium subangulatum in
Weinkauff. 1862 = Cardium exiguum GmeJin. "1886
Locard: Cardium subangulatum Scacchi, 1836 =
Cardium exiguum Gmelin, 1789 sic. --1888 Kobelt:
Cardium subangulatum Scacchi = Cardium exiguum
Gmelin. --1890 Paetel: Cardium subangulatum Scach.
= exiguum Gm. --1892 Bucquoy, Dautzenberg &
Dollfus: Cardium subangulatum Scacchi = Cardium
exiguum Gmelin (fide Fischer-Piette 1977). --1902
Be llini: C. subangulatum, in Scacchi = Cardium
(Parvicardium) exiguum, Gm. --1917 Pallary : Cardium
subangulatum Scacchi. --1962 Grossu: Ca rdium
subangulatum Scacchi = Cardium (Parvicardium)
exiguum Gmelin, 1790. --1969 Nordsieck:
Parvicardium exiguum subangulatum (Scacchi, 1836).
--1977 Fischer-Piette: Cardium subangulatum Scacchi
= nomen nudum. --1989 Voskuil, p. 263 : Cardium
subangulatum Scacchi , 1836 = nomen nudum. CHARACTERS AND OCCURRENCE - REMA RKS.
-·1977 Fischer-Piette: Cardium subangulatum Scacchi
= nomen nudum (there is no description, no figure).
HABITAT. --1969 Nordsieck: littoral and algal zone.
DISTRIBUTION. --1969 Nordsieck: Mediterranean.
subapertum Orbigny, 1852. - Combination to be revised:
"Cardium" subapertum Orbigny, 1852. Cardiidae.
Fossil. RECORDS. --1980 Ruhoff: Cardium subapertum
Orbigny, 1852; [1850-1852] vol. 3: 118.
sllbarqllatllm Munster, 1840. - Combination to be
re vised : "Cardium " subarquatum Muenster, 1840.
?Cardiidae. Fossil. RECORDS. --1922-32 Sherborn, p.
6222 : Cardium subarquatum Muenster, 1840.
sllbasperulllm d 'Orbigny, 1850. - Combination to be
revised: "Cardium " subasperulum d'Orbigny, 1850.
?Cardiidae. Fossil. RECORDS. --1922-32 Sherborn, p.
6223: Cardium subasperulum d ' Orbigny, 1850.
subassimile Vidal, 2003. [Figs. Part 3 p. 868]. Vasticardium subassimile Vidal, 2003. TYPE REPOSITORY.
MNHN. TYPE LOCALITY. Unknown. Cardiidae. Recent.
RECORDS. --2003 Vidal p. 57, figs. 1-3,6: Vasticardium
subassimile sp. nov.
subcarinata arcae/ormis Wenz, 1942 . - Didacna
782
Tropical Marine Mollusc Programme (TMMP)
subcarinata luxuriosa - subdeforme
subcarinata arcaeformis Wenz, 1942. Lynmocardiidae.
?Pliocene. REcORDs. ··1945 ZR: Didacna subcarinata
arcaeformis Wenz, 1942 subsp . nov. M. Pont [?
Pliocene].
subcarinata luxuriosa Wenz, 1942. - Didacna
subcarinata luxuriosa Wenz, 1942. Lymnocardiidae.
Pliocene. RECORDS. ··1945 ZR: Didacna subcarinata
luxuriosa Wenz, 1942 subsp. nov. O. Pout-A. Day
Valea Salcii. Pliocene. ··1980 ZR: Didacna subcarinata
luxuriosa Wenz 1942. Type species of Luxuridacna
Papaianopol, 1980. Pliocene. Ref. to Papaianopol.
subcarinatum Deshayes, 1838. - Combination to be
revised: "Cardium" subcarinatum Deshayes, 1838.
Lymnocardiidae. Fossil. REcORDs. ··1922-32 Sherborn,
p. 6226: Cardium subcarinatum Deshayes, 1838.
subcarinifera Ruzencev, 1929. - Monodacna subcarinifera Ruzencev, 1929. Lynmocardiidae. Apscheronian.
RECORDS . ··1929 ZR: Monodacna subcarinifera
Ruzencev, 1929 sp. n. Apscheronian Transcaucasia.
*subcatillus. - Homonymy to be noticed. Didacna
subcatillus from 2 sources: Andrusov, 1910. Pravoslavleff, 1913.
subcatillus Andrusov, 1910. - Didacna subcatillus
Andrusov, 1910. Lymnocardiidae. Fossil. RECORDS.
··1970 Glibert & Poel, p. 62: Didacna (s.s.) subcatillus
Andrusov, 1910.
subcatillus Pravoslav leff, 1913. - Didacna subcatillus
Pravoslavleff, 1913. Lymnocardiidae. Fossil. RECORDS.
"1930 ZR: Didacna subcatillus Pravoslavleff, 1913
sic sp. n.
subcatiJlus angusta Popov, 1983 (subspecies). - Didacna
subcalillus angusla Popov, 1983. Lymnocardiidae.
Pleistocene. TYPE LOCALITY. Russian SFSR. RECORDS.
··1983 Popov, p. 156, fig.: Didacna subcatillus angusta
subsp. nov.
subcatillus elongatoplana Popov, 1983 (subspecies). Didacna subcatillus elongatoplana Popov, 1983.
Lymnocardiidae. Pleistocene. TYPE LOCALITY. Russian
SFSR. RECORDS. ··1983 Popov, p. 158, fig.: Didacna
subcatillus elongatoplana subsp. nov.
subcatillus itilica Popov, 1983 (subspecies). - Didacna
subcatillus iti/ica Popov, 1983. Lymnocardiidae.
Pleistocene. TYPE LOCALITY. Russian SFSR. RECORDS.
··1983 Popov, p. 158, fig.: Didacna subcatillus iti/ica
subsp. nov.
subcolarata Andrusov, 1965. - Nomen nudum. RECORDS.
··1965 ZR: Monodacna subcolarala Andrusov, 1965
sp. nov. [nom. nud.], S. Russian seas.
subcolorata Vassoievitch, 1929. - Monodacna subcolorala Vassoievitch, 1929. Lymnocardiidae. Fossil.
RECORDS. ··1929 ZR : Monodacna subcolorata
Vassoievitch, 1929 sp. n. Pliocene or Post Tertiary N.
Caucasus. ··1935 ZR: Monodacna subcolorata [Ref.
Eberzin 1935]. ··1977 Popov: Monodacna subcolorata.
subcongestum Bose, 1910. - Combination to be revised:
"Cardium" subcongestum Bose, 1910. Cardiidae.
Cretaceous. RECORDS. ··1911 ZR: Cardium
subcongeslum Bose, 1910 sp. n. Mexico Cretaceous.
subcordatum Suter, 1917. - Changed combination:
Hedecardium subcordalum (Suter, 1917). Cardiidae.
Tertiary. TYPE LOCALITY. North Canterbury N. Z.
RECORDS. ··1917 Suter: Cardium subcordalum . ··1977
Popov: Hedecardium subcordalum (Suter, 1917).
··1978 Maxwell : Hedecardium (Hedecardium)
subcordatum (Suter, 1919). ··2002 Schneider:
Hedecardium subcordatum (Suter, 1917). New
Zealand. Chattian.
subcostale (mode). - See under subcostalis.
subcostalis Petrov, 1982. - Clinocardium subcostalis
Petrov, 1982. Cardiidae. Pleistocene. RECORDS. ··1982
ZR: Clinocardium subcostalis Petrov, 1982 sp. nov.
Russian SFSR. Ref. to Petrov, p. 103. ··1998 Kafanov:
Laevicardium subcostale (Petrov, 1982).
subcrassatellata Andrusov, 1964. - Didacna
subcrassatellata Andrusov, 1964. Lymnocardiidae.
Tertiary. RECORDS. ··1964 ZR: Didacna
subcrassatellata Andrusov, 1964 sp. nov. Tertiary,
USSR.
subcrassatellatoides Andrus. - Changed combination:
Pontalmyra subcrassatellatoides (Andrus.).
Lymnocardiidae. Fossil. RECORDS. ··1987 Taktakishvili:
Pontalmyra subcrassatellatoides (Andrus.)
subcrassidens (subspecies) . - See macrodon
subcrassidens Eberzin, 1960.
subcrenula. - Pontalmyra subcrenula. Lymnocardiidae.
Fossil. RECORDS. ··1987 Taktakishvili: Pontalmyra
subcrenula sic.
*subcrenulata. - Homonymy to be noticed. Didacna
subcrenulata from 2 sources: Andrusov, 1964. Possibly Andrussow (?year).
subcrenulata Andrusov, 1964. - Didacna subcrenulata
Andrusov, 1964. Lymnocardiidae. Tertiary. RECORDS.
··1964 ZR: Didacna subcrenulata Andrusov, 1964 sp.
nov. Tertiary, USSR.
subcrenlliata Andrussow. - Didacna subcrenulata
Andrussow. Lymnocardiidae. Fossil. RECORDS. ··1953
Celidje: Didacna subcrenulata Andrus.
subcurtum. - Pleuriocardia subcurtum. Cardiidae.
Fossil. RECORDS. "1987 Geary: The transformation
from Pleuriocardia subcurtum to Pleuriocardia
pauperculum is fairly abrupt.
subcylindricum Mayer, 1857. - Combination to be
revised: "Cardium" subcylindricum Mayer, 1857.
Cardiacea. Fossil. RECORDS. ··1980 Ruhoff: Cardium
subcylindricum Mayer, 1857; [1856-1857] vol. 5; 362.
subdecorticatum Bell, 1913. - Combination to be
revised: "Cardium" subdecorticatum Bell, 1913.
Cardiidae. Crag. RECORDS. "1913 ZR: Cardium
subdecorticatum Bell, 1913 sp. n. Suffolk Crag.
subdecussata Miinster, 1840. - See under subdecussatum
Orbigny, 1850.
subdecussatum Orbigny, 1850. - Combination to be
revised: "Cardium" subdecussatum d'Orbigny, 1850.
Cardiidae. Fossil. RECORDS. ··1922-32 Sherborn, p.
6235 : Cardiola subdecussata Muenster, 1840. "192232 Sherborn, p. 6235: Cardium (Muenster)
subdecussatum d'Orbigny, 1850. - Cardiola.
subdecussatum Shuto, 1960. - Clinocardium
subdecussatum Shuto, 1960. Cardiidae. Upper
Miocene. RECORDS. ··1960. ZR: Clinocardium
subdecussatum Shuto, 1960 sp. nov. Upper Miocene,
Tsuma member Miyazaki Group, Japan. ··1973 Keen:
Clinocardium subdecussatum Shu to, 1960. Upper
Miocene. ··1977 Popov: Clinocardium subdecussatum
Shuto, 1960.
subdeforme Falckenstein. 1911. - Combination to be
revised : "Cardium " subdeforme Falckenstein. 1911 .
Cardiidae. Senon. RECORDS. ··1911 ZR: Cardium
Phuket Marine Biological Center Special Publication vol. 30 (2004)
783
subdeitoidellin - subelongatllm
subdeforme Fa1ckenstein. 1911 sp. n. Senon Gennany.
subdeltoideum Orbigny, 1850. - Combination to be
revised: "Cardium" subdeltoideum d'Orbigny, 1850.
Cardiidae. Fossil. RECORDs. 001922-32. Sherborn, p.
6235: Cardium subdeltoideum d'Orbigny, 1850.
subdentatllm Deshayes, 1838. [Figs. Part I p. 181].Changed combination: Pselldocatillus subdentatum
(Desha yes, 1838). Lymnocardiidae. Fossil. RECORDS.
001877 Grimm: Cardium subdentatum Deshayes.
001922-32 Sherborn, p. 6236: Cardium subdentatum
Deshayes, 1838. 001967 Ebersin: Pseudocatillus
subdentatus (Desh.).
subdepressa Andrusov, 1964. - Didacna subdepressa
Andrusov, 1964. Lymnocardiidae. Tertiary. RECORDS.
001964 ZR: Didacna subdepressa Andrusov, 1964 sp.
nov. Tertiary, USSR.
subdeserta (afL) Loer. - Changed combination :
Pontalmyra aff. subdeserta (Loer.). Lymnocardiidae.
FossiL RECORDS. 001987 Taktakishvili : Didacna
subdeserla Loer. - Pontalmyra aff. subdeserta (Loer.).
subdichotomum d'Orbigny, 1850. - Cardium subdichotomum d'Orbigny, 1850. Cardiidae . FossiL
RECORDS. 001922-32 Sherborn, p. 6237: Cardium
subdicholomum d'Orbigny, 1850
sllbdinnense d'Orbigny, 1844. - Cardium subdinnense
d'Orbigny, 1844. Cardiidae. Cretaceous. RECORDS.
001922-32 Sherborn, p. 6237: Cardium subdinnense
d'Orbigny, 1844 [Cretaceous].
subdiprosopum (nomen nudum) Stevanovic, 1982. Laevicardium subdiprosopum Stevanovic, 1982.
Cardiidae. Miocene. RECORDS. 001956 ZR :
Laevicardium subdiprosopum Stevanovic, 1956 sp.
nov. nom. nlld. Miocene. Yugoslavia.
subdiprosopum Stevanovic, 1982. - New combination:
Lymnocardium subdiprosopum Stevanovic, 1982.
Lymnocardiidae. Upper Miocene. REcORDs. 001982 ZR:
Limnocardium subdiprosopum Stevanovic, 1982 sp.
nov. Yugoslavia. Ref. to Stevanovic, p. 123 .
subdiscors Orbigny 1850. - Changed combination:
Discors subdiscors (Orbigny, 1850). Cardiidae. Eocene.
GENOTYPE. Dall (1902) stated that Cardium subdiscors
d 'Orbigny is type of subgenus Discors Deshayes, 1858.
Subsequently raised to generic level by auct. TYPE
LOCALITY. Paris Basin. R ECORDS . 001850 Orbigny:
Cardium subdiscors. 001886 Cossmann : Divaricardium
subdiscors (d'Orbigny). 001902 DalJ : Cardium
subdiscors d'Orbigny. 001922-32 Sherborn, p. 6237:
Cardium subdiscors d'Orbigny, 1850 . 001969
Nordsieck : Discors subdiscors (d 'Orbigny). 001977
Popov: Discors subdiscors (Orbigny, 1850). Type of
Discors Deshayes, 1858. 001995 Le Renard et af.:
Nemocardium (Dis cors) subdiscors [Cardium
subdiscors] d 'Orbigny, 1850. 001995 Pacaud & Renard:
Nemocardium (Discors) subdiscors. 001999 Jacques
Vidal (pers. com.) MNHN, Paris : Fossils from the
Paleontological Collection as Discors subdiscors
d'Orbigny. Ypretien. France (Aisne, Oise), B52855 to
-58, B68916, B72596.
sllbdissimile Orbigny, 1850. - Combination to be revised :
"Cardium " subdissimile d ' Orbigny, 1850. Cardiidae.
Fossil. RECORDS. 001922-32 Sherborn, p. 6237: Cardium
subdissimile d 'Orbigny, 1850.
sllbdubium Orbigny, 1850. - Combination to be revised:
"Cardium " subdllbium d 'Orbigny, 1850. Cardiidae.
Fossil. REcORDs. 001922-32 Sherborn, p. 6238: Cardium
subdubium d ' Orbigny, 1850.
sllbedentuillm Orbigny, 1852. - Synonym of Pteradacna
subedentulum d' Orbigny, 1852. Lymnocardiidae.
Fossil. RECORDS . 001969 Moore et al.: Pteradacna
edentulum (Deshayes, 1838) syn. of Cardium
subedentulum d'Orbigny, 1852. 001980 Ruhoff:
Cardium subendentulum Orbigny, 1852; [1850-1852]
vol. 3: 120.
*subedllie. - Homonymy to be noticed. Cardium subedule from 2 sources: Orbigny, 1852. - Raulin, 1897.
subedule Orbigny, 1852. - Synonym of Cerastoderma
edule (Linnaeus, 1758). » Regarding C. sllbedule:
Cardiidae . FossiL RECORDS. 001902 Dollfus &
Dautzenberg, p. 324: Cardium subedule d'Orbigny,
1852. Syn . of Cardium (Cerastoderma) edule Linne,
1758.
subedule Raulin, 1897. - Synonym of Cerastoderma
edule (Linnaeus, 1758). » Regarding C. subedule:
Cardiidae . Fossil. RECORDS . 001902 Dollfus &
Dautzenberg, p. 325: Cardium subedule Raulin, 1897.
Syn. of Cardium (Cerastoderma) edule Linne, 1758.
subelangatllm "Reeve" of Valenciennes. - Err. pro
subelongatum. Misidentification: Mexicardia procerum
(Sowerby, 1833). Cardiidae. Recent. REMARKS. Reeve
has not described a species named Cardium
subelangatum. RECORDS. 00 1846 Valenciennes, pI. 17,
fig. 2: Cardium subelangatum "Reeve". 001886 Dall:
Cardium subelongatum Valenciennes, 1846 not of
Sowerby, 1840. 001947 Hertlein & Strong: Cardium
subelangatum "Reeve" = Cardium (Mexicardia)
procerum Sowerby.
subelegans Orbigny, 1852. - Combination to be revised:
"Cardium" subelegans Orbigny, 1852. Cardiidae.
Recent. RECORDS. 001980 Ruhoff: Cardium subelegans
Orbigny, 1852; [1850-1852] vol. 3: 21.
sllbellipticllm Alekseev, 1963. - Combination to be
revised: "Cardium" subellipticum Alekseev, 1963.
Cardiacea. Palaeogene. RECORDS. 001963 ZR: Cardium
subellipticum Alekseev, 1963 sp. nov. Palaeogene, PreArals, Russia.
*subelongatum. - Homonymy to be noticed. Cardium
subelongatum from 2 sources: Sowerby, 1840. Valenciennes, 1846.
sllbelongatum Sowerby, 1840. [Figs. Part I p. 119, 134,
172. Part 3 p. 867]. - Synonym of Acrosterigma
magnum (Linne, 1758). OTHER JUDGEMENTS:
Trachycardium leucostoma (Born, 1780). » Regarding
C. subelongatum: Cardiidae. Recent. TYPE MATERIAL.
001999 Vidal: the types of Sowerby's taxon are located
in BMNH. There are two uncatalogued specimens from
Cuming collection from the type locality (St. Thomas
Island, West Indies) which could be considered as
syntypes. TYPE LOCALITY. ad Sancta Thomae insulam
[St. Thomas, the West iョ、ゥセャ@
(Sowerby 1840) .
RECORDS. 001840 Sowerby, 182 n part, fig. 61: Cardium
subelongatum . • 01840 Sowerby, Proc. Z.S. London p.
108: Cardium subelongalum . 001840 Sowerby,
catalogue p. 4, No. 55: Cardium subelongatum , Sow.
001840 Sowerby, corrected list, p. 8 : Cardium
subelongatum (Sowerby, No. 55). 001858 Adams &
Adams, p. 455: Cardium (Trachycardium) subelongatum Sow. 001863 Swift: Cardium subelongatum
Sow. 001864 Krebs: Cardium subelongatum Sow.
784
Tropical Marin e Mollusc Programme (TMMP)
subelollgatum - subgracile
··1869 Romer: Cardium subelongatum Sowerby.
··1884 Grasset: Cardium subelongatum. Sow. "1890
Paetel: Cardium subelongatum Sow. ··1902 Dall:
Cardium (Trachycardium) subelongatum Sowerby,
1840. "1902 DaJl : Cardium subelongatum Sowerby,
1840 not Cardium angulatum Lamarck of authors.
··1922-32 Sherborn, p. 6238 : Cardium subelongatum
Sowerby, 1841. ··1939 McLean: Cardium
subelongatum Sowerby, 1840 = Trachycardium
leucostoma (Born, 1780). ··1944 Clench & Smith in
Abbott 1958: subelongatum Sowerby, 1840 junior
synonym of magnum Linne 1758. ··1946 Abrard:
Cardium subelongatum Sow. ··1951 McLean: Cardium
subelongatum Sowerby = Cardium (Trachycardium)
magnum Linnaeus. ··1977 Fischer-Piette: Cardium
subelongatum Sowerby = Trachycardium leucostoma
(Born , 1780). ··1977 Fischer-Piette: Cardium
subelongatum in Reeve 1844 = Laevicardium procentm
Sowerby. ··1999 Vidal: Cardium subelongatum
Sowerby, 1840: fig. 61; 1841: 108 = Acrosterigma
magnum (Linne, 1758). - CHARACTERS AND
OCCURRENCE - CHARACTERS of diagnostic value:
The deep-ploughed interstices between the ribs are very
characteristic in this species (Reeve, 1944). OF
COLLECTIONS. ··ZMUC I specimen. AntiJles DESCRIPTIONS. ··1844 Reeve: shell rather elongately ovate,
oblique, radiately ribbed, about 32-33 sharply angled
ribs, interstices between them deeply excavated, the
extreme posterior ribs armed with small tubercles, the
middle ribs smooth, the anterior crenated; whitish or
yeJlowish, stained and spotted with red. ··1999 Vidal:
shells medium (very rarely large), almost perfectly
ellipsoidal and equilateral, exceptionally with
truncation ofthe posterior part. Ribs show pronounced
backwards curvature, but posterior part very rarely
expanded. Adult shells always elongated; width also
relatively constant. Lunular area very small. Glossy
exterior surface brightly coloured with orange, white,
yeJlow, pink and purple. Interior often colored only in
umbonal area and on ventral margin. Hinge typical of
genus, . No sterigma. Ribs in median and anterior parts
a gash very
are higher than usual and intersticeihキゥ、・セ@
frequently occurs along posterior 8t or 9 t rib ofMPQ
in right valve. Interstices regularly notched by
transverse, elongated, sometimes crescent-shaped
notches. DIMENSIONS. ··1840 Sowerby: long. 1.85 poll.
(length 47 mm), lat. 1.40 poll. (width 36 mm), alt. 2.35
poJl (height 60 mm). ··1999 Vidal: the largest specimen
(73 x 55 x 44 mm) is probably the one figured by Reeve
(1844, fig. 57). REMARKS ... 1840 Sowerby: C. subelongatum resembles some of the figures to which
Lamarck refers for his C. marmoreum , and which
Bruguiere quotes for C. elongatum. It is much longer
and smoother than the former, and does not agree with
the description. The true C. elongatum is described by
Bruguiere, from a specimen of M. de Lamarck, as an
elongated, ventricose shell 0[39-40 ribs, and attaining
a large size. It seems to have been a matter of dispute
between the two conchologists, whether the abovenamed species were identical. We are unable to meet
with a sheJl agreeing with Bruguiere's description, until
the arrival of Mr. Cuming with fine specimens
sufficiently characteristic to set the matter at rest. The
present species has fewer ribs, wider apart, and less
ventricose than in C. elongatum Brug. DISTRIBUTION.
··1844 Reeve : lsI. of St. Thomas, West Indies. ··1884
Grasset: Guadeloupe. ··1999 Vidal : West Indies,
Jamaica (authors). South Florida and Bahamas, south
through the West Indies and Gulf of Mexico to Brazil
(Abrolhos Archipelago).
subelollgatum Valenciennes, 1846. - Identical with
Trachycardillm procentm (Sowerby). » Regarding
Cardium subelongatllm: Cardiidae. Recent. RECORDS.
··1901 DaJl : Cardium subelongatum Valenciennes,
1846 (not of Sowerby, 1840) = Cardium procerum
Sowerby, 1833 . ··1909 Lamy: Cardium subelongatum
Valenciennes, 1846 (non Sowerby) = Cardium
procerum Sowerby.
subemarg;lIatum Orbigny, 1852. - Combination to be
revised: "Cardium " subemarginatum Orbigny, 1852.
Cardiidae. Fossil. RECORDS. ··1980 Ruhoff: Cardium
subemarginatum Orbigny, 1852; [1850-1852] vol. 3:
119.
subesperallzae Eberzin, 1958. - New combination:
Lymnocardium subesperanzae Eberzin, 1958. Lymnocardiidae. Pliocene. RECORDS. ··1958 ZR: Limnocardium (Euxinicardium)subesperanzae Eberzin, 1958
sp . nov. Kerchenski Peninsula. Crimea Pliocene of the
Ponto-Caspian basin, Russia.
sUbjlavum Sowerby, 1840. - Trachycardium subjlavum
(Sowerby, 1840). Cardiidae. Fossil. RECORDS. ··1977
Habe: Trachycardium (Regozara)subjlavum (Sowerby,
1840).
*sub/ragile. - Homonymy to be noticed. Cardium
sub/ragUe from 3 sources : Orbigny, 1850. - Boettger,
1875. - Melville (?year).
sub/ragile Boettger, 1875. - Synonym of Discors
kyetubokiensis Eames, 1950. » Regarding Cardium
sub/ragile: Cardiidae. Upper Eocene. TYPE LOCALITY.
Nanggulansch, Java, Indonesia . RECORDS. ··1919
Martin: Cardium sub/ragile Boettger. ··1950 ZR:
Cardium sub/ragile Boettger, 1875 non Orbigny, 1850
= Discors kyetubokiensis Eames, 1950 nom. nov.
sub/ragile Melville . - Combination to be revised :
"Cardium" sub/ragile Melville. Cardiidae. ?Fossi1.
RECORDS. ··1886 Cossmann: Cardium sub/ragile
MelviJle.
sub/ragile Orbigny, 1850. - Changed combination:
Bucardium sub/ragile (Orbigny, 1850). Cardiidae.
Eocene . RECORDS. ··1922-32 Sherborn, p. 6243:
Cardium subfragile d'Orbigny, 1850. "1977 Popov :
Bucardium (Vepricardium) sub/ragile (Orbigny).
"1995 Le Renard et al.: Cardium (Bucardium)
sub/ragile d 'Orbigny, 1850 (= /ragile) [Authorship
needed. There are a number of Cardium fragile] . ··1995
Pacaud & Renard: Cardium (Bucardium) subfragile.
sub/raterculus Dufour, 1881 . - Nemocardium
sub/raterculus (Dufour, 1881). Cardiidae. Fossil.
RECORDS. "1970 Glibert & Poel: Nemocardium (s.s.)
sub/raterculus (Dufour, 1881).
sub/utile Celidje, 1953. - New combination :
Lymnocardium sub/utile Celidje, 1953. Lymnocardiidae. Fossil. RECORDS. ··1953 Celidje, p. 157:
Limnocardium (Tauricardium) sub/utile nov. sp.
subgraci/e Orbigny, 1850. - Combination to be revised:
"Cardium" subgracile d'Orbigny, 1850. ?Cardiidae.
Fossil. RECORDS. ··1922-32 Sherborn, p. 6245 : Cardium
subgracile d 'Orbigny, 1850.
Phukel Marine Biological Cenler Special Publica lion vol. 30 (2004)
785
subgrallosum - subleve
subgranosum (variety). » Regarding Cardium var.
subgranosum: I have not been able to trace a reference
to this taxon. Cardiidae. Recent. OF COLLECTIONS.
··ZMUC: H.C. [H. Cuming] No. 117 Philippines.
Cardium var. subgranosum. It could be a catalogue
name.
sllbgranulatum Orbigny, 1850. - Combination to be
revised: "Cardium" subgranulatum d'Orbigny, 1850.
?Cardiidae. Fossil. RECORDS. "1922-1932 Sherborn, p.
6246. Sherborn: Cardium subgranulatum d'Orbigny,
1850.
subgratum Dufour, 1881. - Loxocardium subgratum
(Dufour, 1881). Cardiidae. Fossil. RECORDS. "1970
Glibert & Poel, p. 50: Loxocardium subgratum (Dufour,
1881).
subguttiferum Archiac, 1854. - Combination to be
revised: "Cardium " subguttiferum Archiac, 1854.
?Cardiidae. Fossil. RECORDS. ··1980 Ruhoff: Cardium
subguttiferum AIchiac, 1854; vol. II : 211 , pI. 3, figs. 3
a,b.
subguttula Eberzin, 1958. - New combination: Lymnocardium subguttula Eberzin, 1958. Lymnocardiidae.
Pliocene. RECORDS. "1958 ZR: Limnocardium
(Ecericardium) subguttula Eberzin, 1958 sp. nov.
Esheri, Guriya region. Pliocene of the Ponto Caspian
basin, Russia.
subhians d' Archiac, 1850. Nomen nudum. Not available.
RECORDS. ··1922-32 Sherborn, p . 6246: Cardium
subhians d'Archiac, 1850 [no n].
subhians Fischer, 1869. - Combination to be revised:
"Cardium" subhians Fischer, 1869. Cardiidae.
Miocene. RECORDS. ··1877 Locard: Cardium subhians
Fischer, 1869. "1999 Jacques Vidal (pers. com.)
MNHN, Paris: Fossils from the Paleontological
Collection as Cardium subhians Fischer. Turkey,
B32955,B32956,B32978.
*subhillanum. - Homonymy to be noticed. Cardium
subhillanum from 3 sources: Munster, 1835. Leymerie, 1842. - d'Orbigny (?year).
subhillanum d'Orbigny. - Combination to be revised:
" Cardium" subhillanum d'Orbigny. Cardiidae.
Cretaceous. RECORDS. ··1999 Jacques Vidal (pers. com.)
MNHN , Paris: Fossils from the Paleontological
Collection as Cardium subhillanum d'Orbigny.
Cretaceous. Neocomien. France (Yonne), S060043.
subhillanum Leymerie, 1842. - Changed combination:
Nemocardium subhillanum (Leymerie , 1842).
Cardiidae. Cretaceous. RECORDS . ··1876 Meek:
Cardium subhillal1um Leymerie. ··1922-32 Sherborn,
p. 6246: Cardium subhillal1um de Leymerie, 1841
[n.n.]. "1922-32 Sherborn, p. 6246: Cardium
subhillanum de Leymerie, 1842. ··1965 ZR: Cardium
subhillanum described and figd. U. Barremian,
Keldzke, Tuarkir, Turkmenia [Ref. Purliev 1965].
··1986 Scott: Nemocardium subhillanum (Leymerie,
1842).
subhillanum Munster, 1835. - Combination to be
revised: "Cardium" subhillanum Munster, 1835.
?Cardiidae. Fossil. RECORDS. ··1922-32 Sherborn, p.
6246: Cardium subhillanum Muenster, 1835.
sUbhispidum Hilber, 1882. - Changed combination:
Parvicardium subhispidllln (Hilber, 1882). Cardiidae.
Fossil. RECORDs. "1913 ZR: Cardium subhispidum =
hispidum var. Tertiary, Kertsch [Ref. Schwetz 1913 b].
··1977 Popov: Parvicardium subhispidum (Hilber,
1882).
subillcerta Andrussow. - Didacna subil1certa
Andrussow. Lymnocardiidae. Tertiary. REcORDs. ··1953
Celidje: Didacna subincerta Andrussow. ··1964 ZR:
Didacna subincerta Andrusov, 1964 sp. nov. sic
Tertiary, USSR
subincertum Orbigny, 1850. - Combination to be
revised: "Cardium·' subincertum d'Orbigny, 1850.
?Cardiidae. Silurian-Devonian. RECORDS. ··1922-32
Sherborn, p. 6247: Cardium subil1certum d'Orbigny,
1850.
subilltermedia Andrussov, 1924 . - Didacna
subintermedia Andrussov, 1924. Lymnocardiidae .
Pliocene. RECORDs. ··1924 ZR: Didacna subintermedia
Andrussov, 1924 sp. n. Pleistocene. "1963 ZR:
Didacna subintermedia Andrusov, 1963 sp. nov. sic
Apsheron layers, Pliocene, Russia.
subilltermedium Orbigny, 1850. - Combination to be
revised: "Cardium" subintermedium d'Orbigny, 1850.
?Cardiidae. Fossil. RECORDS. ··1922-32 Sherborn, p.
6248: Cardium subinlermedium d 'Orbigny, 1850.
subkujalnicellsis Krestovnikov, 1929. - Prosodacna
subkujalnicensis Krestovnikov, 1929. Lymnocardiidae.
Pliocene. RECORDS. ··1929 ZR: Prosodacna
subkujalnicensis Krestovnikov, 1929 sp. n. Pliocene
S. Russia.
sublaeve Orbigny, 1850. - Unicardium sublaeve
d'Orbigny, 1850. ?Cardiidae. Fossil. RECORDS. ··192232 Sherborn: Corbis sublaeve Keys. ··1922-32
Sherborn, p. 6250: Unicardium (Keys.) sublaeve
d'Orbigny, 1850. - Corbis sublaeve.
sublaevigata Andrussov, 1924 . - Monodacna
sublaevigata Andrussov, 1924. Lymnocardiidae.
Pleistocene. RECORDS. ··1924 ZR: Monodacna
sublaevigata Andrussov, 1924 sp. n. Pleistocene. ··1963
ZR: Monodacna sublaevigata Andrusov, 1963 sp. nov.
Pliocene, Russia.
sublaevigatum. - Combination to be revised: "Cardium"
sublaevigatum. Cardiidae. Miocene. ··1999 Jacques
Vidal (pers. com.) MNHN, Paris: Fossils from the
Paleontological Collection. Cardium sublaevigatum.
Gironde, B27482.
slIblamellosllm Orbigny, 1850. - Combination to be
revised: "Cardium" sublamellosum d'Orbigny, 1850.
Cardiidae. Lias- Jurassic . RECORDS. ··1922-32
Sherborn, p. 6250: Cardium sublamellosum d 'Orbigny,
1850. ··1999 Jacques Vidal (pers. com.) MNHN, Paris:
Fossils from the Paleontological Collection. Cardium
sublamellosum d ' Orbigny. Rauracien. France (Yonne),
S04050.
sublatisulcatum Orbigny, 1852. - Combination to be
revised: "Cardium" sublatisulcatum Orbigny, 1852.
Cardiidae. Fossil. RECORDS. "1980 Ruhoff: Cardium
sublatisulcatum Orbigny, 1852; [1850-1852] vol. 3:
118.
sublatum Orbigny, 1850. - Combination to be revised:
"Cardium" sublatum d'Orbigny, 1850. ?Cardiidae.
Fossil. RECORDs. ··1922-32 Sherborn, p. 6251: Cardium
sublalum d'Orbigny, 1850.
sublebedillzevi (variety). - See lebedinzevi var.
sublebedinzevi Davidachvili, 1933.
subleve Tscheltzov, 1965. - Avicardium subleve
Tscheltzov, 1965. Cardiacea. Pliocene. RECORDs. ··1965
786
Tropical Marin e Mol/lise Programme (TMMP)
sublima - suboxypleura
ZR: Avicardium subleve Tscheltzov, 1965 sp. nov.
Pliocene, Turkmenia, USSR.
sublima Orbigny, 1850. - Changed combination:
Loxocardium sublima (Orbigny, 1850). Cardiidae .
Eocene. RECORDS. "1886 Cossmann: Cardium sublima
d'Orbigny, nom. mut. "1922-32 Sherborn, p. 6251:
Cardium sublima d 'Orbigny, 1850. ··1977 Popov :
Loxocardium sublima (Orbigny, 1850). "1995 Le
Renard et al.: Loxocardium sublima [Cardium sublima1
d 'Orbigny, 1850 (= lima). "1995 Pacaud & Renard:
Loxocardium sublima . "1999 Jacques Vidal (pers.
com.) MNHN, Paris: Fossils from the Paleontological
Collection as Loxocardium sublimum d'Orbigny.
Lutetien. France (Bassin de Paris), B52740 to -51,
B691 03, B70552 to -54.
sublimoides Cossmann, 1923. - Changed combination:
Loxocardium sublimoides. Cardiidae. Fossil. RECORDS.
"1995 Le Renard et al.: Loxocardium sublimoides
[Cardium sublimoides 1 Cossmann, 1923. "1995
Pacaud & Renard : Loxocardium sublimoides.
sublineatum Conrad, 1841. -Synonym of Laevicardium
pristis (Valenciennes, 1827). » Regarding Cardium
sublineatum : Cardiidae. Miocene. RECORDS. "1922-32
Sherborn, p. 6251: Cardium sublineatum Conrad, 1841 .
"1999 Jacques Vidal (pers. com .) MNHN, Paris:
Fossils from the Paleontological Collection. Cardium
sublineatum Conrad . USA (Wilmington), B33363.
(Syn. of Laevieardium pristis).
sublingulata Chen, 1976. - Protoeardia sublingulata
Chen , 1976. Cardiidae. Mesozoic fossil. RECORDS.
"1976 Ma et al. p. 258: Protocardia subUnguiata Chen,
1976.
submaeulosum Ludbrook, 1954. - Nomen nudum.
RECORDS. "1955 ZR: Vastieardium (Vastieardium)
submaeulosum Ludbrook, 1954 sp. nov. nom. nud.
Pliocene, Australia.
submagna Martinson, 1959. - Pseudocardinia
submagna Martinson, 1959. Cardiacea. Fossil.
RECORDS . "1959 ZR: Pseudoeardinia submagna
Martinson, 1959 sp. nov. Tyan-Shan, Atbaya region,
Fergana, fossil.
submartini Chen, 1976. - Combination to be revised:
"Cardium" submartini Chen, 1976. Cardiidae. Upper
Triassic. TYPE LOCALITY. Yunnan, China. RECORDS.
"1976 Ma et at. p. 256, pI. 23, figs . 1-4: Cardium
submartini Chen, 1976 sp. nov.
submedius Popov & Nevesskaya, 2000. - Pseudocatillus
submedius Popov & Nevesskaya, 2000. Cardiidae. Late
Miocene. RECORDS . ··2000 Popov & Nevesskaya,
downloaded abstract: Pseudoeatillus submedius n. sp.
sllbminlltllm Orbigny, 1850. - Combination to be
revised: "Cardium " subminutum d 'Orbigny, 1850.
?Cardiidae. Fossil. RECORDS. "1922-32 Sherborn, p.
6254: Cardium subminutum d 'Orbigny, 1850.
SlIbmonodaena Livental, 1931 (genus). - Nomen
nudum. Tertiary, Lymnocardiidae. Fossil. RECORDS.
"1964 Davidaschvili & Kitovani: Submonodaena
Livental, 1931. Genotype: Didaena pleistopleura Dav.
.. 1980 Vokes: Submonodaena Livental, 1931, nom.
nud.
sllbmoquicum Eberzin, 1958. - New combination:
Lymnocardium submoquicum Eberzin, 1958.
Lymnocardiidae. Pliocene. RECORDS. "1958 ZR :
Limnocardium (Moquieardium) submoquicum Eberzin,
1958 sp. nov. Gudauta, Abkhazia, east coast of Black
Sea. Pliocene of the Ponto Caspian basin, Russia.
submultieostatum Orbigny, 1850 . - Changed
combination: Tuteheria submulticostatum (d'Orbigny,
1850). Cardiacea. Fossil. RECORDS. "1922-32 Sherborn,
p. 6254 : Cardium submuiticostatum d'Orbigny, 1850.
"1946 ZR : Tutcheria submultieostatum (Orb.) type of
Tuteheria Cox, 1946 gen. nov. Cardiidae.
submuticum (variety). - See under mucronatum var.
submuticum Coen, 1941.
suboblongum Orbigny, 1852. - Combination to be
revised: "Cardium" suboblongum Orbigny, 1852.
Cardiidae. Fossil. RECORDS. "1980 Ruhoff: Cardium
suboblongum Orbigny, 1852; (1850-1852] vol. 3: 119.
subodenae Gillet, 1943. - New combination :
Lymnoeardium subodenae (Gillet, 1943).
Lymnocardiidae. Fossil. RECORDS . "1987 Anaya:
Limnoeardium subodenae sic Gillet, 1943 .
subodessae Sinzov, 1877.- New combination:
Lymnoeardium subodessae (Sinzov, 1877).
Lymnocardiidae. Fossil. RECORDS ... 1958 ZR:
Limnocardium (Euxinicardium) subodessae Pliocene
[Ref. Eberzin 1958]. "1987 Anaya: Limnocardium
(Euxinicardium) the group subodessae (Sinzov, 1877).
subodessae munteniae Papaianopol, 1974 (subspecies).
- New combination : Ly mnoeardium subodessae
munteniae (Papaianopol , 1974). Lymnocardiidae.
Pliocene. RECORDS. "1974 ZR: Limnoeardium
(Ellxinieardium) subodessae mllnteniae Papaianopol,
1974 subsp. nov. Romania. Prahova District, Dealul
Belciu Mare. Pliocene. Ref. to Papaianopol, p. 77.
subodessae prahovensis Papaianopol, 1974 (subspecies).
- New combination: Lymnocardium subodessae
prahovensis (Papaianopol, 1974). Lymnocardiidae.
Pliocene. RECORDS . "1974 ZR: Limnoeardium
(Euxinieardium) subodessae prahovensis Papaianopol,
1974 subsp. nov. Romania. Prahova District, Dealul
Belciu Mare. Pliocene. Ref. to Papaianopol, p. 80.
subodessae ursina Ebersin 1947 (variety). - Upgraded.
New combination: Lymnoeardium subodessae ursina
(Ebersin, 1947). Lymnocardiidae. Pliocene. REMARKS.
Raised to subspecies by Papaianopol 1982; from
Limnoeardium (Euxinieardium) subodessae var. ursina.
R ECORDS. "1958 ZR : Limnocardium (Euxinieardium)subodessae var. ursina Eberzin, 1958 nov.
Northern Caucasus, Stavropol. Pliocene of the Ponto-Caspian basin, Russia. "1982 Papaianopol :
Limnocardium (Euxinicardium) subodessae ursina
Ebersin, 1947. "1987 Taktakishvili : Limnoeardium
(Euxinieardium) subodessae (Sinzov) var. ursinae
Ebers.
suborbicula (variety). - See under tuberculatum var.
suborbicula Marsh.
suborbicularis Fan, 1963. - Proto cardia suborbieularis
Fan, 1963. Cardiidae. Lias. RECORDS ... 1963 ZR :
Protoeardia suborbicularis Fan, 1963 sp. nov.
Xiaoshui, Guangdong, China, Lias.
subovale Broderip & Sowerby. Cardiidae . Fossil.
RECORDS. "1850 Sowerby, p. 52: Cardium sllbovale
Brod. and Sow. [compared with Cardium haitense n.
sp.].
suboxypleura Gab. - Changed combination: Pontalmyra
suboxypleura (Gab.). Lymnocardiidae. Fossil. RECORDS.
"1987 Taktakishvili: Pontalmyra suboxypleura (Gab.).
787
Phuket Marine Biological Center Special Publication vol. 30 (2004)
subparallella - subretllsum
subparallella Bogachev & Gadzhiyev, 1968. - Didacna
subparallella Bogachev & Gadzhiyev, 1968.
Lymnocardiidae. Quaternary. RECORDS. ··1968 ZR:
Didacna subparallella sp. nov. p . 37, figs. Kbazarian,
Quaternary, Azerbaijan, USSR.
subparvocostata (subspecies) . - See under orbicularis
subparvocostata Baldi, 1963.
subpassyi Dufour, 1881 . - Parvicardium subpassyi
(Dufour, 1881). Cardiidae. Fossil. RECORDS . ··1970
Glibert & Poel, p. 48: Parvicardium subpassyi (Dufour,
1881).
sllbpallcicostata (cf.) R. Horn . - Didacna cf.
subpaucicostata R. Horn. Lymnocardiidae. Fossil.
RECORDs. ··1953 Celidje: Didacna cf. subpaucicostata
R.Horn.
subpetersi (variety and subspecies). - See under
subsquamosum subpetersi Eberzin, 1958.
subplanicostatum Orbigny, 1852. - Combination to be
revised: "Cardium " subplanicostatum Orbigny, 1852.
Cardiacea. Fossil. REcORDs. ··1980 Ruhoff: Cardium
subplanicostatum Orbigny, 1852; [1850-1852] vol. 3:
119.
subplicatojittoni Senes, 1955 . - Combination to be
revised : "Cardium" subplicatojittoni Senes, 1955 .
Lymnocardiidae. Miocene. RECORDS. ··1955 ZR :
Cardium (Limnocardium) subplicatojittoni Senes, 1955
sp. nov. Miocene, Czechoslovakia.
subpolemonis Ebersin, 1967. - Pseudocatillus
subpolemonis Ebersin, 1967. Lymnocardiidae.
Pliocene. RECORDS. "1967 Ebersin: Pseudocatillus
subpolemonis sp. n. ··1967 ZR: Pseudocatillus
subpolemonis Eberzin, 1967 sp. nov. Pliocene, USSR.
··1982 Papaianopol: Pseudocatillus subpolemonis
Eberzin, 1967. ··1987 ZR: Pseudocatillus
subpolemonis Ebersin, 1967. Ref. to Papaianopol.
subporulosum Orbigny, 1850. - Changed combination:
Orthocardium subporulosum (Orbigny, 1850).
Cardiidae. Eocene. RECORDS. ··1886 Cossmann:
Cardium subporulosum d'Orbigny. ··1922-32
Sherborn, p. 6261: Cardium subporulosum d'Orbigny,
1850. ··1977 Popov: Orthocardium subporulosum
(Orbigny, 1850). ··1995 Pacaud & Renard: Cardium
(Orthocardium) subporulosum. ··1999 Jacques Vidal
(pers. com.) MNHN, Paris: Fossils from the
Paleontological Collection as Cardium, Trachycardium,
subporulosum d'Orbigny. Ypresien, France (Oise),
B52657 to -67. B68876, B68890, B68972, B68999,
69038.
subporulosum (subspecies). - See under porulosum
subporulosum Orbigny 1850.
subpraemoqllicum (variety). - See under praemoquicum
var. subpraemoquicum Chelidize, 1957.
subprodllctum Peron, 1891 . - Combination to be revised:
"Cardium" subproductum Peron, 1891. Cardiidae.
Cretaceous. RECORDS. ··1891 ZR: Cardium
subproductum Cretaceous of Tunis [Ref. Peron 1891].
"1999 Jacques Vidal (pers. com.) MNHN, Paris:
Fossi[s from the Paleontological Collection. Cardium
subproductum Thomas Peron. Turonien. Tunesia,
R51451.
subprotracta Popov, 1983. - Didacna subprotracta
Popov, 1983. Lymnocardiidae. Pleistocene. TYPE
LOCALITY. Russian SFSR. RECORDS. ··1983 Popov, p.
167, fig. : Didacna subprotracta sp. nov.
sllbpunctatllm Munster, 1835. - Combination to be
revised: "Cardium" subpunctatum Muenster, 1835 .
?Cardiidae. Fossil. RECORDS.
ᄋQYセRMS@
Sherborn, p.
6262: Cardium subpunctatum Muenster, 1835 .
sllbpygmaeum d'Orbigny, 1850. - Combination to be
revised: "Cardium" subpygmaeum d 'Orbigny, 1850.
?Cardiidae. Fossil. RECORDS. "1922-32 Sherborn, p.
6263: Cardium subpygmaeum d'Orbigny, 1850.
subpyramidata Pravoslavleff, 19 [3 . - Didacna
subpyramidata Pravoslavleff, 19[3. Lymnocardiidae.
Fossil. RECORDS. ··1930 ZR: Didacna subpyramidata
Pravoslav[eff, 1913 sic sp. n.
subqlladrata (mode and subspecies) . - See under
subquadratum Evans & Shumard, 1857 and exiguum
subquadra/a Jeffreys, 1863.
subquadrata Sibiryakova, 1963. - Protocardia
subquadrata Sibiryakova, 1963. Cardiidae. Jurassic.
R ECORDS. ··1963 ZR: Protocardia subquadrata
Sibiryakova, 1963 sp. nov. Jurassic, Bol 'shoi Balkhans.
subquadratum (variety). - See under lamarckii tenue
subquadratum Coen 1937. - exiguum subquadratum
Jeffreys.
subquadratum Evans & Shumard, 1857. - Changed
combination: Protocardia subquadrata (Evans &
Shumard, 1857). Cardiidae. Cretaceous. REMARKS.
Meek (1876) stated that Cardium subquadratum Evans
& Shumard is type of (a) Leptocardia Meek, 1876.
Speden (1970) stated that adductor and pedal
musculature of P subquadrata is closely analogous to
that of Cardium edule Linnaeus and Corculum
hemicardium (Linnaeus). RECORDS. ··1860 Meek &
Hayden: Cardium subquadratum Evans & Shumard.
··1876 Meek: Protocardia (Leptocardia) subquadrata
Evans & Shumard, 1857. "1887 Fischer: Pachycardium subquadrata Evans et Shumard (in section
Leptocardia Meek, 1876). ··1902 DaB: Cardium
subquadratum Evans and Shumard . Type of section
Leptocardia Meek, 1876. ··1970 Speden: Protocardia
subquadrata (Evans & Shumard, 1857, p. 39). ··1980
Ruhoff: Cardium subquadratum Evans & Shumard,
1857; vol. I : 38. ··1999 Jacques Vidal (pers. com.)
MNHN, Paris: Fossils from the Paleontological
Collection as Cardium subquadratum [without author].
Cretaceous sup. USA (Dakota), B20348.
subradiatum d'Orbigny, 1850. - Combination to be
revised: "Cardium" subradiatum d'Orbigny, 1850.
?Cardiidae. Fossil. RECORDS. ··1922-32 Sherborn, p.
6264: Cardium subradiatum d'Orbigny, 1850.
subregulare d'Orbigny, 1850. - Unicardium subregulare
d' Orbigny, 1850. ?Cardiidae. Fossil. RECORDS. ··192232 Sherborn, p . 6265 : Unicardium subregulare
d'Orbigny, 1850.
subretusa (mode). - See under subretusum Sowerby,
1840.
subretusum Sowerby, 1834. [Figs. Part I p. 110, 142.
Part 3 p. 912]. - Changed combination: Lunulicardia
subretusa (Linne, 1758) . OTHER JUDGEMENTS:
Lunulicardia auricula (Niebhur in Forsskaal, 1775).
»Regarding Cardium subretusum: Cardiidae. Recent.
TypELOCALITY. Unknown (Sowerby 1840; Reeve 1845).
TYPE REPOSITORY . BMNH; Cardium subretusum
Sowerby. 3 syntypes. 197129. Neg. No. 54612/A-C.
RECORDS. ··1834 Sowerby, 50 th part, fig. 24: Cardium
subretusum nobis. ··1840 Sowerby, Proc. Z.S. London
788
Tropical Marine Mollusc Programme (TMMP)
subretusum - subrugosum
p. 110: Cardium subrelusum . ··1840 Sowerby,
catalogue p. 7, No. 91: Cardium subrelusum , Sow.
··1840 Sowerby, corrected list , p. 8: Cardium
subrelusum, (Sowerby No. 91). ··1845 Reeve, sp. 100:
Cardium subretusum Sowerby, 1840. ··1858 Adams
& Adams, p . 459: Hemi cardia (Lunulicardia)
subretusa , Sow. "1868 ZR : Cardium subretusum
(Sow., Reeve) is only a variety of retusum (L.), but
auricufa (Forsk.) is a distinct species [Ref. Romer
1868]. "1869 Martens: Cardium subretusum Sow.
(hardly to distinguish from relusum L.). ··1869 Romer:
C. subrelusum Sowerby, synonym of Cardium retusum
L., hardly a variety. ··1870 McAndrew : Hemicardia
(Lunulicardia) subretusum Sow. "1886 Cooke:
Cardium subretusum Sow. ··1884 Grasset:
Hemicardium subretusum. Sow. ··1889 Martens :
Cardium (Hemicardium) subretusum Sow. "1890
Paetel: Hemicardium subrelusum Sow. = retusum L.
"1898 Melvi I! & Standen: Cardium (Lunulicardia)
subrelusum. ··1903 Hidalgo: Hemicardium subretusum
Sowerby. 1840; considered a variety of Hemicardium
retusum Linne by authors. ··1906 Standen & Leicester:
Cardissa (Lunulicardia) subretusum . .. 1907 Hal! &
Standen: Cardissa (Lunulicardia) subretusa Sowerby.
··1907 ZR: Cardium (Lunulicardia) subretusa raised
beach, Red Sea [Ref. Hall & Standen 1907]. ··1909
Lynge : Cardium subretusum Sowerby, 1834 =
Lunulicardia retusum (Linne, 1758). ··1909 Melvill:
Hemicardium (Lunuficardia) subretusum, Sowb.
··1910 Hedley: Opisocardium subrelusum . ··1918 ZR:
Opisocardium subretusum [Ref. Hedley 1918b] .
"1922-32 Sherborn, p. 6265: Cardium subretusum
Sowerby, 1841. ··1931 Thiele & Jaeckel : Cardium
(Lunulicardia) subretusum Sowerby. ··1950 Allan:
Hemicardium subretusum. "1952 Kuroda & Habe :
Corcufum subretusum (Sowerby). ··1960 Kuroda :
Hemicardia (Lunulicardia) subretusa (Sowerby) .
··1962 Kira: Fragum subretus um . ··1964 Spry :
Lunulicardia (Hemicardium) subretusa (Sowerby).
•• I 977 Popov: Corcufum (Lunulicardia) subrelusa
(Linne, 1758). ··1977 Fischer-Piette: Cardium
subretusum Sowerby = Corcufum retusum Linne.
··1977 Wilson & Stevenson: Cardium subreillsum
Sowerby, 1834; junior synonym of Fragum
(Lunulicardium) retusum (Linnaeus, 1758). "1990
Abbott & Dance: Lunulicardia subretusum . "1991
Matsukuma, Okutani & Habe: Lunulicardia subretusa
(Sowerby, 1843 sic) = 1834. ··1992 Oliver: Cardium
subretusum in Macandrew (1870); Cooke (1886); Hall
& Standen (1907) = Cardium auricufa Niebuhr in
Forsskaal, 1775. "1999 Aungtonya el af., PMBC-SP
19(2): 377: Fragum subretusa (Sowerby, 1841). ··2000
Hylleberg, p. 532: Lunulicardia subretusum (Sowerby,
1834). ··2000 Lutaenko, p. 372: Lunuficardia subretusa
(Sowerby, 1841). ··2003 Hylleberg & Kilburn, p. 183:
Lunulicardia subretusum (Sowerby, 1834) . CHARACTERS AND OCCURRENCE - OF COLLECTIONS . ··ZMUC: wet collection. CHARACTERS of
diagnostic value : middle ribs noduled, interstices
merely finely punctured. DESCRIPTIONS. ··1845 Reeve:
Shell oblong-<:ordate, very much swollen, posteriorly
concavely depressed; radiately flatly ribbed, middle ribs
nodulous, interstices between all ribs very closely
punctured; lunule slightly excavated, with a somewhat
obsolete callosity intruding; white within and without.
DIMENSIONS. "1840 Sowerby long. 0.90 poll. (length
23 mm), lat. 1.20 poll. (width 30 mm), all. 1.30 poll
(height 33 mm) . REMARKS . "1840 Sowerby:
distinguished from the Cardium relusum by a more
elongated form and smoother behind the angle, the
cavity is not so deep, and the callosity is more strongly
marked. ··1845 Reeve: chiefly distinguished from the
Cardium relusum by the middle ribs being noduled,
and the interstices between the whole being merely
finely punctured. ··2000 Lutaenko: according to
Fischer-Piette (1977), this species is a synonym of
Corcufum (Lunulicardia) relusum (L., 1767). "1999:
Higo el af. regarded it as a synonym of Lunulicardia
hemicardium (L., 1758). ··1977 Habe, ··1993 Bernard
et al. and ··1997 Xu regarded this species as separate
individual taxon . ··2003 Hylleberg & Kilburn:
maintains that the status of this species is still debated .
Some malacologists consider it a junior synonym of
Lunulicardia relusa (Linnaeus, 1758. Known from
Vietnam. HABITAT. "1909 Melvill: Providence at 2580 fathoms , on hard rough ground. DISTRIBUTION. ··1845
Reeve: unknown. ··1884 Grasset: Ceylon, Singapore.
"1994 Hylleberg : Red Sea, Indo-Pacific. ··1999
Aungtonya el at.: Thailand, Andaman Sea. ··2000
Hylleberg and ··2000 Lutaenko: Vietnam. ··2003
Hylleberg & Kilburn: Vietnam.
subretusum Sowerby (variety). - Infrasubspecific.
Card iidae. Recent. RECORDS. ··1890 Paetel:
Hemicardium var. subretusum Sow.
subrhaetica Krumbeck, 1923 . - Protocardia subrhaetica
Krumbeck, 1923. Cardiidae. Fossil. RECORDS. "1976
Ma et al. p. 257: Protocardia subrhaetica Krumbeck,
1923.
subriegeli Sinz., 1875 . - Changed combination .
Macradacna subriegeli (Sinz., 1875). Lymnocardiidae.
Fossi l. RECORDS. "1967 Ebersin : Macradacna
subriegeli (Sinz.). ··1967 ZR: Macradacna subriegeli
pro Monodacna subriegeli (Sinz., 1875) [Ref. Eberzin
1967] .
subrotundatum Szots, 1953. - Corcufum subrotundatum
Szots, 1953. Cardiidae. Eocene. RECORDS. ··1954 ZR:
Corcufum subrotundatum Szots, 1953 sp. nov. Eocene
Hungary.
subrugosum Sowerby, 1838. [Figs. Part I p. 112, 121,
134. Part 3 p. 861]. - Downgraded to subspecies:
Vasticardium jlavum subrugosum (Sowerby). OTHER
JUDGEMENTS: Cardium jlavum Linnaeus , 1758. Vasticardium futeomarginata (Voskuil & Onverwagt,
1991). - Cardium rugosllm Lamarck. - Trachycardillm
vertebra tum (Jonas, 1844). » Regarding Cardium
subrugosllm: Cardiidae. Quaternary-Recent. TYPE
LOCALITY. Ad insulam Ceylon (Sri La[hka) (Sowerby
1840). RECORDS. "1838 Sowerby, 149 t part, fig. 3ffi
Cardium subrugosum nobis. ··1838 Sowerby, 184t
part, fig. 71: Cardium subrugosum . "1840 Sowerby,
Proc. Z.S. London p. 108: Cardium subrugosum . •• 1840
Sowerby, catalogue p. 5, No. 58: Cardium subrugosum
Sowerby, 1838 (fig. 32) = Cardium rugosum Lam.
··1840 Sowerby, catalogue p. 5, No. 59: Cardium
subrugosum Sow. ··1840 Sowerby, corrected list, p. 8:
Cardium subrugosum (Sowerby No. 59). ··1843
Hanley: Cardium subrugosum Sow. ··1844 Reeve, sp.
55: Cardium subrugosum Sow. "1857 Adams &
Phuket Marine Biological Center Special Publication vol. 30 (2004)
789
subrugosum - subsquamulosum
Adams: Cardium (Trachycardium) subrugosum . ··1858
Adams & Adams, p. 455 : Cardium (Trachycardium)
subrugosum Sow.··1858 Fischer, P: Cardium
subrugosum Sow. ··1869 Dunker: Cardium
subrugosum Sow. ··1869 Frauenfeld: Cardium
subrugosum Sow. ··1869 Romer: Cardium subrugosum
Sowerby, oblique, related to C. leucostomum Born.
"1872 Tryon: Cardium subrugosum Sow. ··1884
Grasset: Cardium subrugosum Sow. ··1890 Paetel:
Cardium subrugosum Sow. ··1896 Elera: Cardium
subrugosum Sow. ··1903 Hidalgo : Cardium
subrugosum Sow. ··1905 Hidalgo : Cardium
subrugosum Sow. ··1907 ZR : Cardium (Trachycardium)subrugosum raised beach, Red Sea [Ref. Hall
& Standen 1907]. "1919 Martin : Cardium subrugosum
Sow. [Quaternary to Recent]. "1922-32 Sherborn, p.
6268: Cardium subrugosum Sowerby, 1841. "1928
Faustino: Cardium subrugosum Sow. ··1932 Prashad:
Cardium subrugosum Sowerby, 1834 = Cardium
flavum Linne, 1758. "1942 Abrard: Cardium
subrugosum Sow. ··1945 Keen : Vasticardium
subrugosum Sow. ··1956 Franc : Laevicardium
subrugosum Sow. ··1964 Shikama: Trachycardium
(Vasticardium) subrugosum (Sowerby). "1974 Habe
& Kosuge: Vasticardium (Regozara) subrugosum
(Sowerby, 1840). "1977 Fischer-Piette: Cardium
subrugosum Sowerby = Laevicardium jlavum Linne
auctorum . ··1977 Wilson & Stevenson: Cardium
subrugosum Sowerby. ··1990 Abbott & Dance:
Trachycardium subrugosum. ··1992 Dharma:
Trachycardium subrugosum. ··1992 Oliver: Cardium
subrugosum Sowerby, 1834 = Trachycardium
vertebratum (Jonas, 1844). "1999 Aungtonya et al.,
PMBC-SP 19(2): 378 : Trachycardium subrugosum
(Sowerby, 1840). "1999 Vidal: Cardium subrugosum
Sowerby, 1838 = Vasticardium jlavum (Linne, 1758).
- CHARACTERS AND OCCURRENCE - OF COLLECTIONS. ··ZMUC 2 specimens from the Philippines.
DESCRIPTIONS. "1840 Sowerby: 33 ribs, of which the
25 anterior ones are rounded. ··1844 Reeve: shell rather
elongately ovate, attenuated towards the umbones, very
oblique, anterior side peculiarly rounded; radiately
ribbed, about 30 ribs, more or less, the extreme posterior
ribs smooth, the rest rugosely crenated; yellowish white
faintly tinged with rose, sometimes covered towards
the margins with a brown epidermis . DIMENSIONS.
"1840 Sowerby: long. 2.30 poll. (length 60 mm), lat.
1.70 poll. (width 43 mm), alt. 2.40 poll (height 61 mm).
REMARKS. ··1840 Sowerby: shell less spread, ribs less
elevated, the 8 posterior ones are so little raised as to
leave the surface nearly smooth, and anterior side
smoother than in C. rugosum. ··1844 Reeve: the
extreme posterior ribs of this species are smooth and
nearly obsolete, whilst the middle ribs are most
delicately wrinkled; and the wrinkles , increasing
gradually in size and distance from each other, impart
to the anterior portion of the shell, a distinctly crenated
character. "1997 Vidal: accepted subrugosum at the
subspecificJevel. DISTRIBUTION. ··1844 Reeve: Ceylon,
Philippine Islands, and Zanzibar. ··1994 Hylleberg:
Ceylon, Philippines, Zanzibar(Reeve 1844). IndoPacific. "1999 Aungtonya et al.: Thailand, Andaman
Sea.
subrugosum of Abrard 1942. - Identical with
Vasticardium pectiniforme. RECORDS . "1997 Vidal:
Cardium subrugosum in Abrard 1942 = V. pectiniforme.
subrugosum of Lamy 1927. - Identical with
Vasticardium luteomarginata marerubrum. RECORDS.
··1997 Vidal: Cardium subrugosum Sowerby sensu
Lamy 1927 (not Cardium subrugosum Sowerby, 1838)
= Vasticardium luteomarginatum marerubrum (Voskuil
& Onverwagt, 1991).
subrugosum of Reeve 1845. - Identical with Vasticardium luteomarginata. RECORDS. "1997 Vidal :
Cardium subrugosum in Reeve 1845 sp. 55 = V.
luteomarginata (Voskuil & Onverwagt, 1991).
subrugosum (subspecies). - See under flavum
subrugosum Sowerby, 1838.
subrugosum Sowerby, 1838 (variety) . [Figs. Part 1 p.
112). - Synonym of "Cardium " Jucatum Spgl. »
Regarding Cardium var. subrugosum: Cardiidae.
Recent. RECORDS. "1838 Sowerby, 149 th part, fig. 34:
Cardium var. subrugosum. ··1840 Sowerby, corrected
list p. 8: Cardium subrugosum. ··1870 Morch: Cardium
var. subrugosum Sow. = C. implicatum sic Sow. junior
synonym of Cardium Jucatum Spgl.
subschakonensis Akhvlediani, 1984. - Macradacna
subschakonensis Akhvlediani, 1984. Lymnocardiidae.
Pliocene. RECORDS. ··1984 ZR : Macradacna subschakonensis sp. nov. Georgia SSR. Ref. to Akhvlediani
p.80.
subserrigerum Orbigny, 1852. - Combination to be
revised: "Cardium" subserrigerum d'Orbigny. Cardiidae. Miocene. RECORDS. ··1980 Ruhoff: Cardium
subserrigerum Orbigny, 1852; [1850-1852) vol. 3: 118.
"1999 Jacques Vidal (pers. com .) MNHN, Paris:
Fossils from the Paleontological Collection. Cardium
subserrigerum d'Orbigny. Burdigalien. France
(Bordeaux), B27481 .
subsetosum Sowerby (error). - Printer's mistake for
subretusum (Sowerby). Sowerby never described a
species named subsetosum. RECORDS. ··1964 Shikama:
Corculum (Lunulicardia) subsetosum. ··1977 FischerPiette: Cardium subsetosum Sowerby = species
insufficiently known.
subsimile Munster, 1840. - Combination to be revised:
"Cardium" subsimile Muenster, 1840. ?Cardiidae.
Fossil. RECORDS. ··1922-32 Sherborn, p. 6271 : Cardium
subsimile Muenster, 1840
subspinigera (subspecies). - See under texana
subspinigera Cragin, 1893.
subsquamosum Andrussow. - New combination:
Lymnocardium subsquamosum Andrussow. Lymnocardiidae. Fossil. RECORDS. "1953 Ce1idje: Limnocardium (Tauricardium) subsquamosum Andrussow.
subsquamosum subpetersi Ebersin (variety) . Upgraded. New combination: Lymnocardium
subsquamosum subpetersi. Lymnocardiidae. Fossil.
RECORDS.
··1953
Celidje:
Limnocardium
(Tauricardium) subsquamosum Andrus var. subpetersi
Ebers.
subsquamulosum Andrusssov, 1903 . - New
combination : Lymnocardium subsquamulosum
Andrusssov, 1903 . Lymnocardiidae. Miocene and
Pliocene. RECORDS . ··1958 ZR: Limllocardium
(Tauricardium) subsquamulosum Andrusssov, 1903 .
PI iocene [Ref. Eberzin 1958). "1958 ZR:
Limnocardium subsquamulosum Andrusssov, 1903
790
Tropical Marine Mollusc Programme (TMMP)
subsquamulosllm bosphorica -subsllicatina
subgenotypeof(Tauricardium) Eberzin, 1958. Pliocene
of the Ponto-Caspian basin, Russia. "1964 ZR :
Limnocardium subsquamulosum Andrusov, 1964 sp.
nov. sic Rumania, Miocene.
subsqllamulosum bosphorica Eberzin, 1958 (variety).
- Upgraded . New combination: Lymnocardium
subsquamulosum bosphorica Eberzin, 1958.
Lymnocardiidae. Pliocene . RECORDS. "1958 ZR:
Limnocardium (Tauricardium) subsquamulosum var.
bosphorica Eberzin, 1958 nov. Kerchenski, Crimea,
Black Sea. Pliocene of the Ponto-Caspian basin,
Russia.
sllbsquamulosum jinitima Eberzin, 1958 (variety). Upgraded. New combination: Lymnocardium
subsquamufosum jinitima Eberzin , 1958 . Lymnocardiidae. Pliocene. RECORDS. "1958 ZR: Limnocardium (Tauricardium) subsquamulosum var.finitima
Eberzin, 1958 nov. Kerchenski, Crimea; Kipchak Ama
Darya river, south of the Aral Sea. Pliocene of the
Ponto-Caspian basin, Russia.
sllbsquamulosllm nasyrica Eberzin, 1958 (variety). Upgraded. New combination: Lymno cardium
subsquamulosum nasyrica Eberzin, 1958. Lymnocardiidae. Pliocene. RECORDS. "1958 ZR: Limnocardium (Tauricardium) subsquamulosum var. nasyrica
Eberzin, 1958 nov. Kerchenski, Sea of Azov. Pliocene
of the Ponto-Caspian basin, Russia.
sllbsquamulosum oviformis Eberzin, 1958 (variety) . Upgraded. New combination: Lymnocardium
s ubsquamulosum oviformis Eberzin , 1958.
Lymnocardiidae . Pliocene. RECORDS. "1958 ZR:
Limnocardium (Tauricardium) subsquamulosum var.
oviformis Eberzin, 1958 nov. Kerchenski, Crimea.
Pliocene of the Ponto-Caspian basin, Russia.
subsquamulosum praesubsquamulosum Eberzin, 1958
(variety). - Upgraded . New combination: Lymnocardium subsquamufosum praesubsquamulosum
Eberzin, 1958. Lymnocardiidae. Pliocene. RECORDs.
"1958 ZR : Limnocardium (Tauricardium) subsquamufosum var. praesubsquamulosum Eberzin, 1958
nov. Tamansky, Azov Sea-Black Sea. Pliocene of the
Ponto--Caspian basin, Russia.
subsquamulosum sola Eberzin, 1958 (variety) . Upgraded. New combination: Lymnocardium
subsquamulosum sola Eberzin, 1958. Lyrrmocardiidae.
Pliocene . RECORDS. "1958 ZR : Limnocardium
(Tauricardium) subsquamulosum var. sola Eberzin,
1958 nov. Kerchenski, Crimea. Pliocene of the PontoCaspian basin, Russia.
subsquamulosllm sllbpetersi Eberzin, 1958. - New
combination: Lymnocardium subsquamulosum
subpetersi Eberzin, 1958. Lymnocardiidae. Pliocene.
RECORDs. "1958 ZR: Limnocardium (Tauricardium)
subsquamulosum subpe/ersi Eberzin, 1958 subsp. nov.
Kerchenski Peninsula, and Crimea and Kuban. Pliocene
of the Ponto--Caspian basin, Russia.
substratonis Celidje, 1953. - Paradacna substratonis
Celidje, 1953. Lyrnnocardiidae. Fossil. RECORDS. "1953
Celidje, p. 157: Paradacna substratonis nov. sp... 1975
ZR: Paradacna substra/onis Chelidze, 1975 sp. nov.
Georgia SSR, Khobs region Lower Pliocene. Ref. to
Chelidze, p. 198.
substriatulum Orbigny, 1850. - Combination to be
revised: "CardiHln" substria/ulum d 'Orbigny, 1850.
?Cardiidae. Fossil. RECORDS. "1922-32 Sherborn, p.
6273: Cardium substria/ulum d 'Orbigny, 1850.
*substriatum . - Homonymy to be noticed. Cardium
substriatum from 2 sources: Conrad, 1837. - Orbigny,
1850).
substriatum Conrad, 1837. - Changed combination:
Laevicardium substria/um (Conrad, 1837). Cardiidae.
Pleistocene. TYPE LOCALITY. California. RECORDS. "1837
Conrad: Cardium substria/um Conr. "1843 Hanley:
Cardium substriatum Conr. "1856 Carpenter: Cardium
substriatum Conr. "1858 Adams & Adams, p. 456:
Cardium (Cerastoderma) subs/riatum , Conr. "1857
Carpenter: Cardium substria/um Conr. "1860
Carpenter: Cardium substria/um Conr. "1860
Stimpson: Cardium subs/riatum Conr. "1864
Carpenter: Liocardium subs/ria/um Conr. almost
identical with C. elenense [Peru). "1864 Carpenter:
Liocardium subs/riatum Conr. = cruen/a/um Gld.
"1872 Tryon : Laevicardium subs/ria/um Conr. "1872
Carpenter: Laevicardium substria/um (Conrad, 1837).
"1890 Paetel: Cardium subs tria tum Conr. is
Laevicardium. "1901 Dall: Cardium (Laevicardium)
substriatum Conrad, 1838. "1907 Kelsey: Liocardium
subs/ria tum Conr. "1907 Smith, M: Cardium
substriatum Conr. "1907 ZR: Cardium substriatum
[Ref. Smith 1907] . "1916 Dall : Laevicardium
substriatum Conr. "1918 Chace: Cardium subs/ria/um
Conr. "1921 Dall : Laevicardium substria/um Conr.
"1923 Strong: Cardium substriatum Conr. "1924
Oldroyd: Cardium subs/riatum Conrad, 1837. "1931
Grant & Gale: Cardium subs/riatum Conr. "1934
Hertlein: C. subs/ria/um Conrad, by Jordan = Cardium
elenense Sowerby . .. 1937 Keen: Laevicardium
subs/riatum. "1945 Burch: Laevicardium substriatum
Conr. "1948 Smith & Gordon: Cardium substriatum
Conr. "1954 Abbott: Laevicardium substriatum Conr.
"1958 Stohler: Laevicardium substria/um Conr. "1960
Morris: Laevicardium substriatum Conr. "1961
Stohler: Laevicardium substriatum Conr... 1965 Coan:
Laevicardium substria/um Conr. "1966 Keen:
Laevicardium substriatum (Conrad, 1837). "1966
Morris: Laevicardium substriatum Conr. "1968 Abbott:
Laevicardium subs/ria/um Conr. "1969 MacLean:
Laevicardium substria/um Conr. "1974 Abbott :
Laevicardium substria/um (Conrad, 1837). "1977
Popov : Laevicardium substriatum (Conrad, 1837).
"1977 Fischer-Piette: Laevicardium substria/um
Conrad. "2000 Coan e/ af.: Laevicardium subs/riatum
(Conrad, 1837) ... 2001 Vidal, p. 407: Laevicardium
substriatum. - CHARACTERS AND OCCURRENCE
- OF COLLECTIONS. "ZMUC from San Diego, USA.
DESCRIPTIONS ... 2001 Vidal : tentacles numerous. No
eyes on tentacles. DISTRIBUTION. "ZMUC: San Diego,
USA.
substria/um Orbigny, 1850. - Combination to be revised :
"Cardium" substriatum d'Orbigny, 1850. Cardiidae.
Fossil. RECORDS. "1922-32 Sherborn, p. 6274: Cardium
subs/ria/um d'Orbigny, 1850.
*subsulcatina. - Homonymy to be noticed. Didacna
subsulca/ina from 2 sources: Andrusov, 1910. Andrusov, 1964.
sllbsulcatina Andrusov, 1964. - Didacna subsulca/ina
Andrusov, 1964. Lymnocardiidae. Miocene. RECORDS.
"1964 ZR : Didacna subsulca/ina Andrusov, 1964 sp.
791
Phuket Marine Biological Center Special Publication vol. 30 (2004)
sllbslllcaltina - sllbzlatarski
nov. Miocene USSR.
subsulcaltina (cf.) Andrusov, 1910. - Didacna cfr.
subsulcaltina Andrusov, 1910. Lyrrmocardiidae. Fossil.
RECORDS. ··1987 Anaya: Didacna (s . s.) cfr.
subsuscaltina [err. pro subsulcaltina] Andrusov, 1910.
subsulcatum Link, 1807. - Combination to be revised:
"Cardium" subsulcatum Link, 1807. ?Cardiidae.
Fossil. RECORDs. ··1922-32 Sherborn, p. 6276: Cardium
subsulcatum Link, 1807.
subsyrmiense Andrussow. - Changed combination:
Lymnocardium subsyrmiense Andrussow. Lymnocardiidae. Pliocene. RECORDS. ··1907 Andrussow:
Limnocardium subsyrmiense Andrus. ··1958 ZR:
Limnocardium (ElIXinicardium) subsyrmiense Pliocene
[Ref. Eberzin 1958]. "1958 ZR: Limnocardium
subsyrmiense Andrus. sub genotype of (ElIXinicardium)
Eberzin, 1958. Pliocene of the Ponto-Caspian basin,
Russia. ··1987 Anaya: Lymnocardium subsyrmiense
Andrussov, 1903 . Type species (D.O .) of (Euxinicardium) Ebersin, 1947.
subsyrmiense lacta Eberzin, 1958 (variety). - Upgraded.
New combination: Lymnocardium subsyrmiense lacta
Eberzin, 1958. Lymnocardiidae. Pliocene. RECORDS.
··1958 ZR: Limnocardium (Euxinicardium)
subsyrmiense var. lacta Eberzin, 1958 nov. Kerchenski
Peninsula, Black Sea. Pliocene of the Ponto-Caspian
basin, Russia.
sllbsyrmiense lecta (variety). - Euxinicardium
subsyrmiense lecta. Lymnocardiidae. Fossil. REcORDs.
··1987 Taktakishvili : Euxinicardium subsyrmiense
lecta . - Limnocardium (ElIXinicardium) subsyrmiense
Andrus. var. lecta Ebers.
sllbsyrmiense turbida Eberzin, 1958 (variety) . Upgraded . New combination: Lymnocardium subsyrmiense turbida Eberzin, 1958. Lymnocardiidae.
Pliocene. REcORDs. ··1958 ZR: Limnocardium (ElIXinicardium) subsyrmiense var. turbida Eberzin, 1958 nov.
Kerchenski Peninsula, Black Sea. Pliocene of the
Ponto-Caspian basin, Russia.
subtentum. - Combination to be revised: "Cardium "
subtentum. Cardiidae. Lower Eocene. REcORDs. ··1871
Stoliczka, 150: C. subtentum.
subtenue integricosta Almera & Bofill (variety) . Combination to be revised:: "Cardium" subtenue var.
integricosta Almera et Bofill. Lyrrmocardiidae. Fossil.
RECO RDS. ··1987 Anaya: Cardium subtenue var.
integricosta Almera et Bofill.
subtenllisulcatum Orbigny, 1852 . - "Cardium "
subtenuisulcatum Orbigny, 1852. Cardiidae. Fossil.
RECORDs. ··1980 Ruhoff: Cardium subtenuisulcatum
Orbigny, 1852; [1850-1852] vol. 3: 21.
subtile. - "Cardium " subtile. Cardiacea. Fossil. REcORDs.
"1886 Fischer: Cardium subtile.
subtrapezinum Tscheltzov, 1965. - Avicardium
subtrapezinum Tscheltzov, 1965. Cardiacea. Pliocene.
RECORDS. ··1965 ZR: Avicardium subtrapezinum
Tscheltzov, 1965 sp. nov. Pliocene, Turkmenia, USSR.
subtrianglliare mut. n. - See under productum Sow.
subtriangulare Cottreau, 1924.
sllbtrigonale d'Orbigny, 1850. - Conocardium
subtrigonale d'Orbigny, 1850. Conocardiidae. Fossil.
RECORDs. ··1922-32 Sherborn, p. 6281: Conocardium
subtrigonale d'Orbigny, 1850.
*sllbtrigonllm . - Homonymy to be noticed. Cardium
subtrigonum from 2 sources: Orbigny, 1850. - Morris
& Lycett, 1853 .
sllbtrigonllm Morris & Lycett, 1853. - Changed
combination: Protocardium subtrigonum. Cardiidae.
Lias-Jurassic. RECORDS. ··1980 Ruhoff: Combination
to be revised: "Cardium" subtrigonum Morris &
Lycett, 1853; [1853-1854] vol. 2: 64, pI. 7, fig. 3.
··1999 Jacques Vidal (pers. com.) MNHN, Paris:
Fossils from the Paleontological Collection.
Protocardium subtrigonum . Bathonien. France (Pas de
Calais), S01914.
sllbtrigonum Orbigny, 1850. - Cardium subtrigonum
d'Orbigny, 1850. Cardiidae. Fossil. RECORDs. "192232 Sherborn, p. 6281 : Cardium subtrigonum d 'Orbigny,
1850.
subtrigonum Orbigny, 1850. - Unicardium subtrigonum
d'Orbigny, 1850. ?Cardiidae. Fossil. RECORDS. ··192232 Sherborn, p . 6281: Unicardium subtrigonum
d'OrbignY,1850.
subtruncata (variety). - See under martelli var.
subtruncata Viola, 1949.
subtruncatum Orbigny, 1850. - Combination to be
revised: "Cardium" subtruncatum d'Orbigny, 1850.
?Cardiidae. Fossil. REcORDs. ··1922-32 Sherborn, p.
6282: Cardium subtruncatum d'Orbigny, 1850.
subturgidum millasiense Fontannes, 1882. Laevicardium subturgidum millasiense (Fontannes,
1882). Cardiidae. Fossil. REcORDs. ··1970 Glibert &
Poel, p. 37: Laevicardium (Habecardium) subturgidum
millasiense (Fontannes, 1882). "1989 Gontsharova:
Laevicardium (Habecardium) subturgidum millasiense
in Glibert & Pool, 1970.
subturgidum Orbigny, 1852. - Changed combination:
Nemocardium subturgidum (d'Orbigny, 1852).
Cardiidae.
Pliocene.
RECORDS.
··1970
ZR:
Laevicardium (Habecardium) subturgidum (Orbigny,
1852) [Ref. Glibert & Van De Poe I 1970]. ··1980
Ruhoff: Cardium subturgidum Orbigny, 1852; [18501852] vol. 3: 118. ··1989 Gontsharova: Nemocardium
(Habecardium) subturgidum (Orbigny, 1852). "1989
Gontsharova: Laevicardium (Laevicardium) subturgidum Glibert, 1945. ··1999 Jacques Vidal (pers.
com.) MNHN, Paris: Fossils from the Paleontological
Collection. Cardium subturgidum d ' Orbigny. Belgium
(Anvers), B35561.
subumbonatum Orbigny, 1852. - Combination to be
, revised: "Cardium " subumbonatum Orbigny, 1852.
Cardiidae. Fossil. RECORDS. "1980 Ruhoff: Cardium
subumbonatum Orbigny, 1852; [1850-1852] vol. 3:
118.
subventricosum Orbigny, 1850. - Combination to be
revised: "Cardium" subventricosum d 'Orbigny, 1850.
?Cardiidae. Fossil. RECORDS. ··1922-32 Sherborn, p.
6289: Cardium subventricosum d'Orbigny, 1850.
subvitrea Andrussov, 1924. - Adacna subvitrea
Andrussov, 1924. Lymnocardiidae. Pliocene and
Pleistocene. RECORDS. "1924 ZR: Adacna subvitrea
Andrussov, 1924 Pleistocene. ··1963 ZR: Adacna
(Adacna) subvitrea Andrusov, 1963 sp. nov. Apsheron
layers, Pliocene, Russia.
subvulgaris (subspecies). - See under rudis subvulgaris
Nevesskaya, 1963.
Sllbzlatarski Ebersin, 1967. - Pseudocatillus
subzlatarskii Ebersin, 1967. Lymnocardiidae. Pliocene.
792
Tropical Marine Mollusc Programme (TMMP)
Succuridacna - sueziensis
RECORDs. ··1967 Ebersin: Pseudocatillus subzlatarskii
sp. n. ··1967 ZR: Pseudocatillus subzlatarskii Eberzin,
1967 sp. nov. Pliocene, USSR.
Succuridacna Korobkov, 1958 (subgenus) . Lymnocardiidae . Miocene . RECORDS . ··1958 ZR:
Succuridacna Korobkov, 1958 subgen . nov. of
Eoprosodacna , subgenotype Cardium goriense
Davidaschvili, Georgia, USSR Kotsakhursk horizon,
Miocene [Ref. Korobkov 1958] . ··1980 Vokes:
Succuridacna Korobkov, 1954 in subfamily
Lymnocardiinae.
suculum in Monterosato 1878, printer's error of C.
siculum Sowerby.
suecicum (spelling). - See under suediense Reeve, 1845.
suecicum Loven, 1846. - Synonym of Parvicardium
minimum (Philippi, 1836). OTHER JUDGEMENTS: not a
synonym of Parvicardium minimum (Philippi, 1836).
» Regarding Cardium suecicum : Cardiidae. Recent.
REcORDs. ··1867 Weinkauff: Cardium suecicum Forbes,
Loven = Cardium minimum Philippi. ··1867 ZR:
Cardium minimum (Phil.) identified on Jeffreys's
authority with Cardium suecicum (Loven) [this
identification is not correct, according to a communication from Mr. Jeffreys, Ref. to Weinkauff 1867].
··1886 Locard: Cardium suecicum Loven, 1846 =
Cardium minimum Philippi , 1845 . ··1890 Paetel:
Cardium suecicum Loven. ··190 I Friele & Grieg:
Cardium suecicum Loven, 1846 = Cardium minimum
Philippi, 1836. ··1912 Jensen: Cardium suecicum
Loven, 1846 = Cardium minimum Philippi . ··1969
Nordsieck: Parvicardillm minimum suecicum (Loven,
1846). ··2001 Aartsen & Goud : Cardium suecicum
Loven, 1846 emend. = Parvicardillm minimum
(Philippi, 1836).
suecicum Reeve (spelling) - See under suediense Reeve.
suedense (error). - Printer's mistake for suediense in
Kobelt 1888.
suediense Reeve, 1845. [Figs. Part I p. 145. Part 3 p.
915]. - Synonym of Parvicardium minimum (Phi lippi,
1836). » Regarding Cardium suediense: Cardiidae.
Recent. TYPE REPOSITORY. BMNH ; Cardium suediense
Reeve Syntypes 2 of 3 valves reg. no. 197628. Neg.
No. 0261. DISTRIBUTION. West coast of Sweden (Reeve
1845). RECORDS. ··1845 Reeve: Cardium suediense.
··1846 Loven: Cardium suecicum Reeve sp. 132.
··1851 Sars: Cardium svecicum Reeve. ··1856 Jeffreys:
(Cardium suecicum B.M.) = Cardium minimum Ph.
(Note: Ph. & B.M. = Philippi and "British Mollusca",
Forbes & Hanley 1849 respectively). ··1858 Adams &
Adams, p. 456: Cardium (Cerastoderma) suecicum
Reeve. ··1859 Danie1ssen: Cardium svecicum Reeve
= Cardium loveniThompson. ··1867 Weinkauff, p. 143:
Cardium suedense [err. pro suediense] Reeve =
Cardium minimum Philippi. ··1869 Romer: Cardium
suediense Reeve fig. 132 = Cardium suecicum Loven.
··1871 March, p. 215, No. 206 Cardium svecicum
Reeve = Cardium minimum Phi 1. ··1872 ZR: Cardium
minimum (Phil.) = suecicum (Reeve), ribs vary from
28-33 [Ref. Bragger 1872]. ··1886 Locard: Cardium
suldiense sic Reeve, 1836 = Cardium minimum,
Philippi, 1845. ··1888 Kobe1t: Cardium suedense sic
Reeve = Cardium suecicum Loven. ··1912 Dautzenberg
& Fischer: Cardium suediense Reeve 1843 = Cardium
(Parvicardium) minimum, Philippi 1836. ··1917
Hidalgo: Cardium suedense sic Reeve = Cardium
Suecicum Loven . ··1922-32 Sherborn, p. 6294:
Cardium suediense Reeve, 1845. ··1932 Winckworth:
Cardium suecicum = Cardium (Parvicardium)
minimum Philippi , 1836. ··1977 Fischer-Piette:
Cardium suediense Reeve, 1845 = Cardium minimum
Philippi. ··2001 Aartsen & Goud: Cardium suediense
Reeve, 1845 = Parvicardium minimum (Philippi, 1836).
- CHARACTERS AND OCCURRENCE - DESCRIPTIONS. ··1845 Reeve: shell minute, orbicular, thin,
fragile, radiately ribbed, 28 convex, prickly ribs; white,
covered with a thin straw-coloured epidermis. REMARKS.
··1845 Reeve: this is the smallest species of Cardium I
have met with; it is not much unlike the figure of an
odd valve discovered by M. Philippi, at Palermo, and
figured in his second volume of the Enum. Moll.
Siciliae, pI. 14. f. 18, under the name of C. minimum;
his description, however, by no means agrees with the
northern species under consideration. DISTRIBUTION.
··1845 Reeve: West coast of Sweden.
suense Sidorova, 1961. - Combination to be revised:
"Cardium" suense Sidorova, 1961. Cardiacea.
Miocene. RECORDS. ··1961 ZR: Cardium suense
Sidorova, 1961 sp. nov. Sue, Sarmatian, Miocene;
Mangyshlak, Turkmeniya, Russia . DISTRIBUTION.
Europe (Winckworth 1932); Arctic, Atlantic, Pacific,
Gulf of Suez (Nordsieck 1969).
suess; Barbot, 1869. - Combination to be revised:
"Cardium" suessi Barbot, 1869. Cardiacea. Sarmatian.
RECORDS. ··1929 ZR: Cardium suessi Barb. Sarmatian
[Ref Ko1esnikov 1929]. ··1946 ZR: Cardium suessi
Barbot. Sarmatian fossil [Ref. Simionescu & Barbu
1940]. ··1970 ZR: Cardium suessi Barbot, 1869 type
sp. for Maequicostates group nov. q.v. [?Ref.].
suessi. - Inaequicostata suessi. Cardiacea. Fossil.
RECORDS. ··1977 Popov: Inaequicostata suessi.
suessi imbecilla Kolesnikov, 1929 (variety). - Upgraded.
Combination to be revised: "Cardium" suessi imbecilla
Kolesnikov, 1929. Cardiacea. Sarmatian. RECORDS.
··1929 ZR: Cardium suessi Barb. var. imbecilla
Kolesnikov, 1929 nov. Sarmatian [Ref. Kolesnikov
1929].
suessi pseudohispida Simionescu & Barbu, 1940
(variety). - Upgraded. "Cardium " suessi pseudohispida Simionescu & Barbu, 1940. Cardiacea.
Sarmatian. REcORDs. ··1946 ZR: Cardium suessi Barbot
var. pseudohispida Simionescu & Barbu, 1940 nov.
Chisinau, Sarmatian fossil, Rumania.
suessiformis (subspecies and variety). - See under
politioanei suessiformis and politioanei var.
suessiformis Jekelius, 1944.
sueziense (mode). - See under sueziensis Issei, 1869.
sueziensis Issei, 1869. [Figs. Part 3 p. 852]. - Changed
combination: Parvicardium sueziensis (Issei, 1869).
Cardiidae. Recent. TYPE LOCALITY. Bay of Suez. RECORDS. ··1869 Issei, p. 76, PI. 3, fig. 4 : Cardium
sueziensis. ··1869 ZR: Cardium sueziense Issei, 1869
sp. n. from Suez. ··1870 Macandrew: Cardium
(Cerastoderma) sueziensis. ··1885 Smith, p. 158-159,
pI. 8, figs. 2-2b: Cardium (Acanthocardia) sueziense
IsseI. ··1899 Hedley: Cardium fragrum [sic. = fragum]
var. sueziense IsseI. ··1906 Melvill & Standen, p. 839:
Cardium (?Acanthocardia) sueziense IsseI. ··1907 ZR:
Cardium sueziense raised beach, Red Sea [Ref. Hall &
Phuket Marine Biological Center Special Publication vol. 30 (2004)
793
suicum - sulcatum
Standen 1907]. ··1909 Lynge: Cardium
(Acanthocardia) sueziensis. ··1938 Lamy: Cardium
(Cerastoderma) sueziensis IsseI. ··1973 Biggs:
Parvicardium sueziensis (Issei, 1869). ··1977 FischerPiette: Cardium (Cerastoderma, Parvicardium)
sueziense IsseI. ··1977 Popov: Loxocardium sueziense
(Issei, 1869). ··1980 Ruhoff: Cardium sueziensis Issei,
1869; p. 76, pI. 3, fig. 4. ··1992 Oliver: Parvicardium
sueziensis. ··1995 Oliver, fig. 1078: Parvicardium
sueziense (Issei, 1869). ··2000 Hylleberg, p. 532:
Parvicardium sueziense (Issei, 1869). Det. 1. Vidal.
··2001 Vidal, p. 408: Fragum sueziense. ··2002
Hylleberg & Kilburn, p. 63: Parvicardium sueziense
(Issei, 1869). ··2003 Hylleberg & Kilburn, p. 178:
Fragum sueziense (Issei, 1869). - CHARACTERS
AND OCCURRENCE - Of COLLECTIONS. ··ZMUC
from the Gulf of Thailand. DESCRlPTIONS. ··1869 Issei:
shell small, equivalve, inequilateral, suborbicular,
somewhat oblique, anterior rounded, posterior obtuse,
subangulate, somewhat inflated, thin, white, small
brown spots, about 23-24 ribs, numerous transverse
scales, small umbo, denticulated margin [I have
translated from Latin]. Roundly subquadrate, posterior
margin slightly longer and less rounded than anterior.
Sculpture of 20-24, low rather broad, radial ribs, all
but the posterior few bearing large, slightly curved cross
bars; posterior ribs with blunt spines. ··200 I Vidal: only
the exhalant aperture is tube-shaped. Inhalant siphon
open towards the foot aperture. No eyes. DIMENSIONS.
··1869 Issei: length 7.5 mm, width 6 mm, diameter 5
mm. ··1995 Oliver: to 8 mm. REMARKS. ··1869 Issei:
C. sueziensis resembles C. minimum Philippi, but
differs in number of ribs and their ornamentation. The dozen odd valves studied by Hedley (1899) passed
gradually into typical C. fragum. The species was
redescribed and reillustrated by Smith (1885, p. 158
and pI. 8) because he condemned the original figure
by Issei (Hedley 1899). Unfortunately C. minimum and
C. fragum are very different, so it is difficult to judge
the status of C. sueziensis, based on information in the
literature. HABITAT. ··1906 Melvill & Standen: sand 3
fathoms, mud, 10 fathoms, coral. ··ZMUC mud 30 m
depth. ··1995 Oliver: muddy sand, lower shore and
below. DISTRIBUTION. ··1899 Hedley: Fiji, Torres Strait.
··1994 Hylleberg: Suez, Red Sea. ··1995 Oliver: Red
Sea. ··ZMUC Arabian Sea. ··2000 HyJleberg: Vietnam
and Cambodia. ··2002 Hylleberg & Kilburn: South
India. ··2003 Hylleberg & Kilburn: Vietnam.
suicum Iqbal, 1970. - "Cardium" suicum Iqbal, 1970.
Cardiidae. Early Eocene. RECORDS. ··1970 ZR:
Cardium suicum Iqbal, 1970 sp. nov. Early Eocene,
Ghazij Formation, Shahrig, Pakistan.
sukaraii Habe, 1961. - Nemocardium sukaraii Habe,
1961. Cardiidae. Recent. RECORDS. ··1961 ZR:
Nemocardium (Microcardium) sukaraii Habe, 1961 sp.
nov. Cardiidae, Tosa Bay, Kochi Pref. Shikoku, Japan.
sulacense Andrusov, 1963. - Combination to be revised:
"Cardium" sulacense Andrusov, 1963. Cardiacea.
Neogene. RECORDS. ··1963 ZR: Cardium sulacense
Andrusov, 1963 sp. nov. Neogene, Russia.
sulcata (variety). - See under archiaciana var. sulcata
Acquaotta, 1929.
*sulcatinum. - Homonymy to be noticed. Cardium
sulcatinum from 2 sources: Lamarck, 1819. - Deshayes,
1838.
sulcatinum Deshayes 1838. - New name: Didacna
andrusovi Gude, 1913. OTHER JUDGEMENTS: Cardium
ponticum Deshayes. Lymnocardiidae. Fossil. RECORDS.
··1857. Deshayes: Cardium sulcatinum. Desh. 1837 sic.
- C. ponticum Desh. ··1913 ZR: "Cardium" sulcatinum
Desh. 1838 non Lamarck, 1819 replaced by Didacna
andrusovi Gude, 1913 nom. nov. 001922-32 Sherborn,
p. 6298: Cardium sulcatinum Deshayes, 1838.
sulcatinum Lamarck, 1819. [Figs. Part 3 p. 921]. Combination to be revised: "Cardium" sulcatinum
Lamarck, 1819. ?Cardiidae. Fossil. RECORDS. ··1819
Lamarck: "Cardium" sulcatinum Lamk. ··1886
Fischer: "Cardium" sulcatinum. ··1857. Deshayes:
Cardium sulcatinum. Lamk. 1818 sic. ··1922-32
Sherborn, p. 6298: "Cardium" sulcatinum Lamarck,
1819.
*sulcatum. - Homonymy to be noticed. Cardium
sulcatum from 3 sources: Lamarck, 1819. - Michelotti,
1839. - Koninck (?year).
sulcatum Koninck. - Replaced by "Cardium"
sulciferum. Deshayes. » Regarding Cardium
sulcatum: Not a cardiid (cardiomorpha). Silurian.
RECORDS. ··1857. Deshayes: "Cardium" sulcatum
Koninck (cardiomorpha). - C. sulciferum. Desh.
sulcatum Lamarck, 1819. [Figs. Part I p. 76 No. 190, p.
79 No. 12, p. 82 No.9. p. 90. Part 3 p. 898]. - Identical
with Laevicardium oblongum (Gmelin, 1791). OTHER
JUDGEMENTS: Laevicardium crassum (Gmelin, 1791). Cardiumflavum Poli - Cardium norvegicum Spengler,
1799. - Cardium serratum Bruguiere. » Regarding
Cardium sulcatum: Cardiidae. Pliocene and Recent.
TYPE LOCALITY. Mediterranean (Reeve 1845). TYPE
REPOSITORY. Lamarck's cabinet, unknown availability.
RECORDS. ··1798 Bruguiere. ID by Lamarck, pI. 298.
fig. 5. Cardium sulcatum. ··1819 Lamarck, p. 10, No.
24: Cardium sulcatum new name for Cardium
oblongum Chemnitz, t. 19, fig. 190. ··1832 Bronn:
Cardium sulcatum Lamarck. = Cardium oblongum
(Chemn.) Brch. 503. ··1835 Deshayes: Cardium
sulcatum Lamarck. ··1836 Philippi: Cardium sulcatum
Lamarck. = Cardium crassum Gm. p. 3254. ··1838
Maravigna: Cardium sulcatum Lamarck. ··1839 Anton:
Cardium sulcatum Lamarck = Cardium oblongum
Chemnitz = Cardium serra tum Bruguiere. ··1840
Sowerby, catalogue p. 5, No. 64: Cardium sulcatum
Lamarck. ··1844 Brown: Cardium sulcatum Lamarck
= Cardium oblongum. ··1844 Philippi: Cardium
sulcatum Lamarck. ··1844 Potiez & Michaud: Cardium
sulcatum Lamarck. ··1846 Verany: Cardium sulcatum
Lamarck. ··1848 Requien: Cardium Sulcatum
Lamarck. = Cardium oblongum Chemn. ··1856 Wood
& Hanley: C. sulcatum Lam. = Cardium oblongum Ch.
··1857. Deshayes: Cardiumsulcatum. Lamk 1818 sic.
··1862 Weinkauff: Cardium sulcatum Lamarck. ··1867
Hidalgo: Cardium sulcatum Lamarck = Cardium
oblongum, Chemnitz. ··1867 Weinkauff: Cardium
sulcatum Lamarck, Deshayes, Philippi = C. oblongum
Chemnitz. ··1867 ZR: Cardium sulcatum (Lam.) and
Cardium oblongum (Chemn.) nearly allied, but may
be kept distinct [Ref. Weinkauff 1867]. ··1868 ZR: The
Mediterranean Cardium sulcatum (Lam.) = oblongum
(Chemn.) = flavum (Born) to be distinguished from
Cardium pennanti and oblongum of Reeve [Ref. Romer
794
Tropical Marine Mollusc Programme (TMMP)
sulcatum - suluanum
1868]. --1869 Romer: Cardium sulcatum Lamarck =
Cardium oblongum Chemnitz. --1872 Monterosato:
Cardium sulcatum = C norvegicum (foss. recente).
--1878 Monterosato : C sulcatum Lamarck = Cflavum
Poli (non Lin.) = Cardium oblongum Chemn. --1884
Monterosato: C sulcatum Lamarck = C flavum Poli
(non Lin.) = Loevicardium sic oblongum. --1886
Locard : Cardium su/catum de Lamarck, 1818 =
Cardium oblongllm Chemnitz, 1782. --1888 Kobelt:
Cardium sulcatum Lamarck = Cardium oblongum
Chemnitz . --1899 Granger: Cardium sulcatum
Lamarck. --1843 Hanley: Cardium sulcatum Lamarck.
--1890 Hidalgo: Cardium su/calum Lamarck =
Cardium oblongum Chemnitz. --1890 Paetel: Cardium
silica tum Lm. = ob/ongum Ch. [not available]. --1900
Locard & Caziot: Cardium sulcatum Lamarck, 1819 =
Cardium ob/ongum Chemnitz, 1789 sic. "1922-32
Sherborn, p. 6300 : Cardium sulcatum Lamarck, 1819.
"1930 Stewart: Laevicardium sllicatum. "1945 ZR:
Cardium sulcatum Lk. [Ref. Lamy 1945]. "1969
Nordsieck: Cardium sulcatum Lamarck, 1819 =
Laevicardium oblongum (Chemnitz, 1782). --1977
Fischer-Piette, p. 176: Cardium sulcatum Lamarck =
Laevicardium oblongum Lamarck sic. --1999 Jacques
Vidal (pers. com.) MNHN, Paris: Fossils from the
Paleontological Collection as Cardium sulcatum
Lamarck. Pliocene. Italy; Algeria; Greece; B36096,
B38899, B40342, B41270. - CHARACTERS AND
OCCURRENCE - REMARKS. --1841 Sowerby, No. 64:
a shell [C sulcatum] has been collected at Plymouth
by Mr. Cuming, which is distinct from C serra tum ,
and may probably tum out to be a British variety of
this species. -- Label at ZMUC : "Lamarck No. 24 =
flavum Born t 3/8, and oblongum Ch fig. 190, E. pI.
298 f. 5 bon, andflavum Poli t 17 f. 9. Gulf d Torento =
oblongum Gmelin, 1791 Mediterranean, Canary lsI."
"1969 Nordsieck: Cardium flavum Born, 1780 =
Cardium sulcatum Lamarck, 1819 = Laevicardium
oblongum (Chemnitz, 1782) . HABITAT. --1969
Nordsieck: the species oblongum: sand at 4-200 m
depth. DISTRIBUTION. "1845 Reeve : the species
sulcatum: Mediterranean. --1969 Nordsieck:
oblongum: Mediterranean, Canary lsI.
sulcatum Leckenby, 1858. - Unicardium sulcatum
Leckenby, 1858. Cardiidae. Fossil. RECORDS. "1980
Ruhoff: Unicardium su/catum Leckenby, 1858; vol. 15 :
14, pI. 3, fig. II.
sulcatum of authors. - RECORDS. --1870 Aradas & Benoit:
Cardium sulcatum auctt. = Cardium ob/ongum
Chemnitz.
sulcatum of Blainville - RECORDS. "1917 Hidalgo :
Cardium sulcatum Blainville = Cardium oblongum.
sulcatum of Gmelin (1791). [Figs . Part I p . 139]. Synonym of Laevicardium oblongum (Gmelin, 1791).
» Regarding Cardium sulcatum: Cardiidae. Recent.
TYPE LOCALITY . Mediterranean (Reeve 1845) .
DISTRIBUTION. Mediterranean (Romer 1891). RECORDS.
"1791 Gmelin: did not refer to Cardiumsulcatum , only
to Cardium oblongum. --1891 Romer: Trachycardium
su/catum Gm. --1845 Reeve, fig. 79: Cardium sulcatum
Gmelin. "1977 Fischer-Piette, p . 18 : Cardium
sulcatum Gmelin = Cardium oblongum Gmelin. CHARACTERS
AND
OCCURRENCE
DESCRIPTIONS. "1845 Reeve: shell elongately ovate,
attenuated towards the umbones, very ventricose,
radiately grooved, posterior area smooth, depressed;
rusty white, covered with a yellowish olive epidermis.
REMARKS. --1845 Reeve: the sculpture of this species
is of such a very similar character to that of the Cardium
oblongum. that I (Reeve) at first doubted whether they
be truly distinct. Upon subsequent examination I
(Reeve) find they differ constantly in form, the Cardium
sulcatum being more elongated and so much more full
and ventricose about the umbones. DISTRIBUTION. --1845
Reeve: Mediterranean .
sulcatum of Payraudeau (1826). - RECORDS. --1826
Payraudeau: Cardium sufcatum Lamarck. n. 24. --1880
Stossich: Cardium sulcatum Payraudeau, 1862 sic =
Laevicardium oblongum Chemnitz.
sulcatum of Philippi . - RECORDS. --1856 Jeffreys:
(Cardium sulcatum Ph.) = Cardium norvegicum B.M.
(Note: Ph. & B.M . = Philippi and "British Mollusca
Forbes & Hanley 1849" respectively).
sulcatus Michelotti, 1839. - Combination to be revised:
"Cardium" sulcatus Michelotti, 1839. Cardiidae.
Fossil. RECORDS. --1922-32 Sherborn, p. 6300: Cardium
sulcatus Michelotti, 1839.
*sulciferum . - Homonymy to be noticed. Cardium
sulciferum from 2 sources : Bayle in Fournel, 1849. Deshayes, 1853 .
sulciferum Bayle in Fournel, 1849. - Combination to
be revised: "Cardium " sulciferum Bayle in Fournel,
1849. ?Cardiidae. Fossil. RECORDS. --1922-32 Sherborn,
p. 6311: Cardium sulciferum Bayle in Fournel, 1849.
sulciferum Deshayes, 1853. - Combination to be revised:
"Cardium" sulciferum Deshayes, 1853 . ?Cardiidae.
Fossil. RECORDS. --1980 Ruhoff: Cardium sulciferum
Deshayes, 1853; vol. 2: 49.
suldiense (error). - Printer's mistake for suediense in
Locard 1886.
sulini Alizade. - Raricardium? sulini (Alizade). Lymnocardiidae. Neogene. RECORDS. "1986 Nevesskaya et
af.: Raricardium? sulini (Alizade) comb. nov. "1987
ZR: Raricardium? sulini (Alizade) comb. nov. Ref. to
Nevesskaya et al. p. 191.
sulini magnum Tscheltzov, 1965 (subspecies). Combination to be revised : "Cardium " sulini magnum
Tscheltzov, 1965. Lymnocardiidae. Pliocene. RECORDS.
--1965 ZR: Cardium sulini magnum Tscheltzov, 1965
subsp. nov. Akchagylian (Pliocene), Turkmenia, USSR.
suluanum Vidal , 1999 . [Figs . Part 3 p . 880]. Acrosterigma suluanum Vidal, 1999. Cardiidae. Recent.
TYPE LOCALITY . Holotype, Tawitawi Island , Sulu
Archipelago, Philippines, 439 m. T YPE REPOSITORY.
Holotype, National Museum of Natural History,
Washington DC . Paratypes, Zoological Museum
University of Copenhagen. TYPE MATERlAL. Holotype,
a right valve (USNM 299435). Paratypes 1,2,3, a left
valve and two right valves from Sunasaki, Japan, 15035 m (ZMUC, coIl. Mortensen). Paratypes 4 and 5 two
right valves, Japan, 113 m, same repository and
collection. RECORDS. --1999 Vidal, p. 298, Table 22,
Figs 8D-F: Acrosterigma suluanum n. sp. CHARACTERS AND OCCURRENCE - CHARACTERS
of diagnostic value. Ribs are rounded and the fine lateral
ornaments are situated at the base of the ribs, touching
the interstice. DESCRIPTIONS. --1999 Vidal: shell ovoid,
not truncated posteriorly and almost equilateral,
Phuket Marine Biological Center Special Publication vol. 30 (2004)
795
sumsarica - symbo/icum
moderately elongated and tumid. Lunule small, rather
imperfectly delineated, its umbonal margin raised.
Colour yellow with orange splashes; interior partially
pink. Hinge slightly asymmetrical and moderately
angled. On right valve, cardinals slightly more than
usually fused at base. Rib number 43.8 ± 1.3, range
41-45. Posteriorly, interstices very thin, ribs very flat
with an axial furrow separating two zones of similar
width. Slightly elongated, tubercular, oblique scales on
posterior part encroaching upon anterior zone,
straddling axial furrow. on middle part of surface, ribs
are flatly triangular and asymmetrical posteriorly, then
rounded. Scales disappear and are replaced further
posteriorly with numerous very thin short ridges or
tubercles, situated at base of rib, touching interstice.
Anteriorly, large scales reappear on posterior flanks,
imposed on the latter fine ornamentation: a similar fine
beading but thinner and more tubercular appears also
at base of anterior flank of ribs. On anterior slope, the
fine beading of both sides disappear and posterior large
scales proceed to top and anterior flank of ribs, forming
free top-ridges which are not imbricated. DIMENSIONS.
Length 34.0, height 43.6, width (calculated) 30.0 mm.
REMARKS. ··1999 Vidal: Acrosterigma suluanum shares
with A. profundum the character of having fine
ornaments on both sides of the ribs on median surface
(difference with A. uniornatum and A. amirante that
have ornaments only on the posterior side). On A.
suluanum, however, the ribs are rounded and the fine
lateral ornaments are situated at the base of the ribs,
touching the interstice, whilst on A. profundum the ribs
are squared and the fine ornaments are separated from
the interstice and slightly overhang it. In addition, the
rib number of profundum is higher, with an average of
58.4, against 43.8 on suluanum. DISTRIBUTION. "1999
Vidal: Known from the Philippines and Japan.
sumsarica (variety). -See under beshbaluki var.
sumsarica Mirkamalova, 1958.
sundica Andrusov, 1963. - Didacna sundica Andrusov,
1963. Lymnocardiidae. Miocene. RECORDS. ··1963 ZR:
Didacna sundica Andrusov, 1963 sp. nov. Miocene,
Russia.
sundicum Vekilov, 1959. - Combination to be revised:
"Cardium" sundicum Vekilov, 1959. Lymnocardiidae.
Pliocene. RECORDS. "1959 ZR: Cardium sundicum
Vekilov, 1959 sp. nov. Pontic beds, Lower Pliocene,
Maraza region, Azerbaidjan.
superbum Tremlett, 1950. - Nemocardium superbum
Tremlett, 1950. Cardiidae. Cretaceous. RECORDS. ··1950
ZR: Nemocardium superbum Tremlett, 1950 sp. nov.
Hampshire Basin, Bracklesham Bay. ··1977 Popov:
Nemocardium (Nemocardium) superlum sic Tremlett,
1950.
superlum (error). - Printer's mistake for superbum in
Popov 1977.
superob/iquum Riedel, 1932. - Combination to be
revised: "Cardium" superobliquum Riedel, 1932.
Cardiidae. Cretaceous. RECORDS. ··1938 ZR: Cardium
superobliquum Riedel, 1932 sp. n. Cretaceous
Cameroons.
supina (variety). - See under rectidens sup ina.
supracimmerica Eberzin, 1960. - Pachydacna supracimmerica Eberzin, 1960. Lyrnnocardiidae. Pliocene.
RECORDS. ··1960 ZR: Pachydacna supracimmerica
Eberzin, 1960 sp. n. Pliocene Tamansk, Bugazski
region, Crimea, USSR.
suprae/egans Gregorio, 1930. - Combination to be
revised: "Cardium" supraelegans Gregorio, 1930.
Cardiidae. Lias. RECORDS. ··1936 ZR: Cardium
supraelegans Gregorio, 1930 sp. n. Lias Palermo.
supsae (subspecies). - See under crassa supsae
Nevesskaya, 1963.
susannae Bagdasaryan, 1979. - Parvicardium susannae
Bagdasaryan, 1979. Cardiidae. Lower Miocene.
RECORDS. ··1979 ZR: Parvicardium susannae Bagdasaryan, 1979. Sp. nov. Ukraine. Ref. to Bagdasaryan,
p.39.
suwannense Mansfield, 1937. - Combination to be
revised: "Cardium" suwannense Mansfield, 1937.
Cardiidae. Oligocene. RECORDS. "1937 ZR: Cardium
(Acanthocardia) suwannense Mansfield, 1937 sp. n.
Suwannee limestone (Oligocene) Florida.
svecicum (error). - Printer's mistake for suediense in
Sars 1851, Danielssen 1859 and Morch 1871.
swediense (error). - Printer's mistake for suediense in
Sowerby 1859.
sybariticum (subspecies). - See under serratum
sybariticum.
sybariticum Dall, 1886. - Changed combination:
Laevicardium sybariticum (Dall, 1886). Cardiidae.
Recent. OF COLLECTIONS. ZMUC from St Thomas, West
Indies. Antilles. RECORDS. "1886 Dall, 12, p. 171:
Cardium serratum var. sybariticum. "1900 Dall:
Cardium serratum var. sybariticum. ··1902 Dall &
Simpson, pI. 58, fig. II: Cardium (serratum var.?)
sybariticum Dall. "1939 McLean: Cardium
sybariticum Dall, 1886 = Laevicardium serra tum
sybariticum (Dall, 1886). "1944 Clench & Smith:
Laevicardium sybariticum Dall. ··1944 ZR:
Laevicardium sybariticum Dall [Ref. Clench & Smith
1944]. "1947 Aguayo & Jaume: Laevicardium
sybariticum Dall. ··1951 McLean: Laevicardium
sybariticum Dall. "1961 Weber: Laevicardium
sybariticum. ··1962 Warmke & Abbott: Laevicardium
sybariticum Dall, 1886. ··1968 Jong & Kristensen:
Americardia sybariticum Dall. "1971 Porter & Wolfe:
Laevicardium sybariticum (DaU) ? "1974 Abbott:
Laevicardium sybariticum (Dall, 1886). "1977
Fischer-Piette: Laevicardium sybariticum Dall &
Simpson. ··1997 Shelton: Laevicardium sybariticum
(Dall, 1886). ··200 I Vidal, p. 407: Laevicardium
sybariticum. - CHARACTERS AND OCCURRENCE
- DESCRlPTIONS. ··1902 Dall & Simpson: shell much
smaller and more delicate than the type, more
compressed, pale or faintly flecked with pink or purple;
beaks often deep pink. ··200 I Vidal: tentacles
numerous. No eyes on tentacles. DIMENSIONS. ··1902
Dall & Simpson: length 11, height 14, diameter 7 mm.
DISTRIBUTION. ··ZMUC St Thomas, West Indies;
Antilles. ··1977 Fischer-Piette Brazil. ··1902 Dall &
Simpson: Porto Rico.
sycophanta Boghachev, 1938. - New combination:
Lymnocardium sycophanta Boghachev, 1938. Lymnocardiidae. Fossil. RECORDS. ··1939 ZR: Limnocardium
sycophanta Boghachev, 1938 sp. n.
symbolicum Iredale, 1929. [Figs. Part 3 p. 903]. - New
name. Ctenocardia symbolicum (Ired ale, 1929). Cardiidae. Recent. Type species of Ctenocardia H. & A.
796
Tropical Marine Mollusc Programme (TMMP)
symmetrica - Tauricardium
Adams, 1857. DISTRIBUTION. central Indo-West Pacific
(Hylleberg 1994). RECORDS. "1929 Iredale, p. 264:
Fragum (Ctenocardia) symbolicum nom. nov. for
Ctenocardia hystrix (Reeve, 1844) (type of genus
named), hystrix preoccupied by Solander 1786. --1929
ZR: Fragum symbolicum nom. n. for Cardium hystrix
Rve. [Ref. lredale 1929bJ. --1932 Prashad:
Cardium(Ctenocardia) symbolicum. Nomen novum
replacing hystrix Reeve, 1844. --1977 Fischer-Piette:
Corculum (Ctenocardia) symbolicum Iredale. --1977
Wilson & Stevenson: Fragum (Ctenocardium)
symbolicum Iredale, 1929; new name for Cardium
hystrix.
symmetrica Popov, 1983. - Didacna symmetrica Popov.
1983. Lymnocardiidae. Pleistocene. TYPE LOCALITY.
Russian SFSR. REcORDs. "1983 Popov, p. 154, fig.:
Didacna symmetrica sp. nov.
syriacum (cf.) Conr. - Trachycardium cf. syriacum Conr.
Cardiidae. Fossil. REcORDs. --1917 ZR: Trachycardium
cf. syriacum Conr. [Ref. Newton 1917].
syrmiensis Homes, 1874. - Paradacna syrmiensis
(Homes, 1874). Lymnocardiidae. Fossil. RECORDS.
"1970 Glibert & Poel, p. 66: Paradacna syrmiensis
(Homes, 1874). "1987 Taktakishvili: Limnocardium
syrmiense R. Hoem.
syrticum (subspecies). - See under glaucum syrticum
Coen,1915.
szaboi Loer. - New combination: Lymnocardium szaboi
(Loer.). Lymnocardiidae. Fossil. RECORDS. --1987
Taktakishvili : Limnocardium szaboi Loer.
T
tabacoensis Myers, 1968. - New combination:
Granocardium tabacoensis Myers, 1969. Cardiidae.
Fossil. "1968 ZR: Cardium (Granocardium)
tabacoensis Myers, 1968 sp. nov. Cardenas formation,
Mexico. --1969 ZR: Cardium (Granocardium)
tabacoensis Myers, 1969 sp. nov. Arroyo del Tabaco,
Cirdenas formation, Arroyo del Aguaje, Mexico.
Tachycardium (error). - Printer's mistake for
Trachycardium in Csepreghy-Meznerics 1959 (Vokes
1980).
taciturna Papaianopol, 1989. - Pachydacna taciturna
Papaianopol, 1989. Lymnocardiidae. Pliocene. TYPE
LOCALITY. Basin Dacique, Romania. RECORDS. "1989
Papaianopo1, p. 71 , fig .: Pachydacna (Pachydacna)
taciturna sp. nov.
taeniopleura DaJl, 1900. - Changed combination:
Planicardium taeniopleura (DaJl, 1900). Cardiidae.
Fossil. --2002 Schneider: citing Dall (1900): Cardium
(Cerastoderma) taeniopleura sp. nov. section
Cerastoderma. --2002 Schneider: citing Stanley
(1986): Planicardium taeniopleura.
taidonense Tolmatchoff, 1925 . - Conocardium
taidonense Tolmatchoff, 1925. Conocardiidae.
Carboniferous . RECORDS. --1925 ZR: Conocardium
taidonense Tolmatchoff, 1925 sp. n. Carbon.
taiwanensis (subspecies). - See under samarangae
taiwanensis Masuda & Huang, 1990.
taiwanensis Masuda & Huang, 1990. - Vepricardium
taiwanensis Masuda & Huang, 1990. Cardiidae.
Miocene. TYPE LOCALITY . Western Foothills of northern
Taiwan. RECORDS. --1990 Masuda & Huang, p. 153,
fig.: Vepricardium taiwanensis sp. nov.
takoyellsis Kafanov & Savizky, 1982, - Yagudinella
takoyensis Kafanov & Savizky, 1982. Cardiidae. Upper
Miocene. RECORDS. --1982 Kafanov & Savizky:
Yagudinel/a takoyensis Kafanov & Savizky. --1982 ZR:
Yagudine/la takoyensis Kafanov, 1982 sp. nov. Russian
SFSR, Ref. to Kafanov, et al. p. 41.
talahabense Martin, 1922. - Combination to be revised:
"Cardium" talahabense (Martin, 1922). Cardiidae.
Lower Miocene. TYPE LOCALITY. Lower Miocene of Java
(Vidal 1999) . RECORDS. --1922 ZR: Cardium
(Eucardium) talahabense Martin, 1922 sp. n. Tertiary.
"1999 Vidal, p. 270: cites Martin 1922, 484, pI. 61,
figs. 105-105a: Cardium (Eucardium) talahabense.
tamallica (subspecies) . - See under caspia tamanica
Nevesskava, 1963.
Tamilicardia (subgenus). - Cardiidae. Upper Cretaceous.
RECORDS . --1983 ZR: (Tamilicardia) subgenus of
Protocardia. India.
tanaica Ebersin, 1967. - Didacnomya tanaica Ebersin,
1967. Lymnocardiidae. Pliocene. RECORDS. "1967
Ebersin : Didacllomya tanaica sp. n. --1967 ZR: Didacnomya tanaica Eberzin, 1967 sp. nov. figs. Pliocene,
USSR.
tallaitica (subspecies) . - See pontocaspia tanaitica
Popov, 1983.
tallaitica Popov, 1955. - Didacna tanaitica Popov, 1955.
Lymnocardiidae. Quaternary. RECORDS. --1956 ZR:
Didacna tanaitica Popov, 1955 sp. nov. Don and
Warsaw Manich Quaternary. Russia.
taphrium Dall, 1900. - Dinocardium taphrium (Dall,
1900). Cardiidae. Fossil. --2002 Schneider: citing Dall
(1900): Cardium (Cerastoderma) taphrium sp. nov.
section Dinocardium.
taracaicum Yokoyama, 1930. - Changed combination:
Laevicardium· taracaicum (Yokoyama , 1930).
Cardiidae. Upper Oligocene-Miocene. RECORDS. --1930
Yokoyama: Cardium taracaicum (fide Keen 1973).
--1930 ZR: Cardium taracaicum Yokoyama, 1930 sp.
n., fossil. --1973 Keen : Laevicardium taracaicum
(Yokoyama, 1930).
taramelli Gortani, 1906. - Conocardium taramelli
Gortani, 1906. Conocardiidae. Carboniferous. RECORDS.
--1906 ZR: COllocardium taramelli Gortani, 1906 sp.
n. Carboniferous, Carnian Alps.
tarchanicum (subspecies). - See under liverovskayae
tarchanicum .
tascharvaticu Sibiryakova , 1962 . - Pr%cardia
tascharvatica Sibiryakova, 1962. Cardiidae. Jurassic.
RECORDS. --1962 ZR: Protocardia tascharvatica
Sibiryakova , 1962. sp . nov. Tascharvat, Western
Turkrneniya, Russia, Jurassic.
tataricu Kafanov ex Lautenschlager, 1997. - Papyridea
tatarica Kafanov ex Lautenschlager, 1997. Cardiidae.
Fossil. RECORDS. --1997 Kafanov, p. 6, pI. 8, fig. 3.:
Papyridea (Pro/lllvia) tatarica Kafanov ex
Lautenschlager.
Tauricurdium Ebersin, 1947 (genus). - Lymnocardiidae.
Tertiary. RECORDS. --1980 Vokes: Tauricardium Ebersin,
1947 in subfamily Lymnocardiinae.
Tuuricurdium Eberzin, 1958 (subgenus) . - Lymnocardiidae. Pliocene. RECORDS. --1958 ZR: Tauricardium
Eberzin, 1958 subgen. nov. of Limnocardium ;
sUbgenotype: Limnocardium subsquamu/osum
Phuket Marine Biological Center Special Publication vol. 30 (2004)
797
tauricum - tenerum
Andrusssov, 1903 Pliocene of the Ponto-Caspian basin,
Russia. --] 958 ZR: Tauricardium Eberzin, 1958
subgen. nov. of Limnocardium q .v. "1983 ZR:
(Tauricardium) subgenus of Lymnocardium Romania.
Dacique Basin. Neogene. Ref. to Papaianopol et al.
"1987 Taktakishvili: (Tauricardium) Ebersin, 1947.
tauricum Sobetskii, 1977. - Granocardium tauricum
Sobetskii, 1977. Cardiidae. Upper Cretaceous.
RECORDs. --1977 ZR: Granocardium tauricum sp. nov.
Ukraine. Ref to Sobetskii, p. 184, fig'd. "1986 AbdelGawad: "Granocardium" tauricum Sobetski, 1977.
taurina Michelotti, 1847. - Acanthocardia taurina
(Michelotti, 1847). Cardiidae. Fossil. RECORDS. "1977
Popov: Acanthocardia (Acanthocardia) taurina
(Michelotti, 1847).
taurinium Michelotti, 1847. - Combination to be revised
"Cardium " taurinium Michelotti, 1847. ?Cardiidae.
Fossil. RECORDs. --1922-32 Sherborn, p. 6393 : Cardium
taurinium Michelotti, 1847.
taurovatum Sacco, 1899. - Laevicardium taurovatum
Sacco, 1899. Cardiidae. Tertiary. RECORDS . --1899
Sacco, p. 53, pI. 12, fig. 6: Laevicardium taurovatum
Sacco--1977 Popov: Laevicardium taurovatum Sacco,
1899.
tcheliense Debeaux in Pilsbry, 1904. - Synonym of
Fulvia mutica (Reeve, 1844). » Regarding Cardium
tcheliense: Cardiidae. Recent. RECORDS. --1994 Vidal:
Cardium tcheliense Debeaux in Pilsbry, 1904 = Fulvia
(Fulvia) mutica (Reeve, 1844).
tchuktchense (subspecies). - See under ciliatum
tchuktchense Kafanov, 1981.
tegalense Oostingh, 1934. - New combination :
Laevicardium tegalense Oostingh, 1934. Cardiidae.
Tertiary. RECORDS. --1934 ZR: Laevicardium
(Laevicardium) tegalense Oostingh, 1934 sp. n. Tertiary
Java.
tegulata Munster, 1840. - See under tegulatum Orbigny,
1850.
tegulatum Dautzenberg, 1900. [Figs. Part 3 p. 907] .
Synonym of Fragum unedo (Linne, 1758). »
Regarding Hemicardium tegulatum: Cardiidae. Recent.
TYPE REPOSITORY. Museum d'Histoire naturelle de Paris.
RECORDs. --1900 Dautzenberg, p. 5-8, pI. I, figs 3-6:
Hemicardium tegulatum n. sp. --1900. ZR :
Hemicardium tegulatum Dautzenberg, 1900 n. sp. hab.
unknown. --1909 Lynge: Hemicardium tegulatum
Dautzenberg, 1900 = Fragum unedo (Linne, 1758).
--1932 Prashad: Hemicardium tegulatum Dautzenberg,
1900 = Fragum unedo (Linne, 1758). --1970 Glibert
& Poel, p. 48: "Hemicardium" tegulatum Dautzenberg,
1900. --1977 Fischer-Piette, p. 144: Corculum
(Hemicardia) tegulatum Dautzenberg. --1981 ZR:
Fragum tegulatum (Dautzenberg j 900) Junior
synonym of F. unedo Linnaeus. Ref. to Germain, p.
166. "1982 ZR: Fragum (Fragum) tegulatum
(Dautzenberg 1900) Diagnostic characters. Taxonomic
position. Ref. to Germain. - CHARACTERS AND
OCCURRENCE - REMARKS. "1977 Fisher-Piette: reestablished tegulatum Dautzenberg, 1900 but the action
was erroneous . The synonymy (=Fragum unedo) is
correct according to Luc Germain who studied the
specimen of Dautzenberg. DISTRIBUTION. "1994
HylJeberg : Indo-Pacific.
tegulatum Hal. - New combination: Lymnocardium
tegulatum (HaL) . Lymnocardiidae. Fossil. RECORDS.
--1987 Taktakishvili : Limnocardium tegulatum (HaL).
tegulatum Orbigny, 1850. [Figs. Part 3 p. 907]. Cardium tegulatum d'Orbigny, 1850. Conocardiidae.
Fossil. REcORDs. --1922-32 Sherborn, p. 6397: Cardiola
tegulata Muenster, 1840. --1922-32 Sherborn, p. 6397:
Cardium (Muenster) tegulatum d'Orbigny, 1850. Cardiola.
teisenbergerense Schlosser, 1926. - New name.
"Cardium" teisenbergerense Schlosser, 1926.
Combination to be revised. Cardiidae. Fossil. RECORDs.
--1926 ZR: Cardium teisenbergerense Schlosser, 1926
nom. nov. for C. raulini Gumbel.
telluris Hoeninghaus, 1831 . - Hemicardium telluris
Hoeninghaus, 1831. - Cardiidae. Fossil. RECORDS.
"1922-32 Sherborn, p. 6401: Hemicardium tel/uris
Hoeninghaus, 1831. --1922-32 Sherborn : Hemicardium
telluris.
telluris Lamarck, 1819. - Not a cardiid. »Regarding
Cardium tel/uris: Cardiacea. Fossil. RECORDs. --1819
Lamarck: Cardium telluris Lamk. "1912 Favre:
Cardium tel/uris Lamarck. [Not a cardiid = Ceromya
elegans Deshayes]. "1922-32 Sherborn, p. 6401:
Cardium tel/uris Lamarck, 1819.
Tendagurium Dietrich, 1933 (genus). - Cardiidae.
Fossil. RECORDS. --1980 Vokes : Tendagurium Dietrich,
1933 in subfamily Protocardiinae.
Tendagurium Dietrich, 1933 (subgenus). - Cardiidae.
Cretaceous. RECORDS. --1933 ZR: Tendagurium
Dietrich, 1933 subgen. n. of Cardium type Cardium
propebanneianum Dietrich, 1933 sp. n. Cretaceous
Africa. --1937 ZR: Tendagurium Dietrich 1933 "type
here designated Cardium (Tendagurium) propebanneianum Dietrich 1933" [Ref. Keen 1937]. --1977
ZR: Protocardia (Tendagurium) . Lower Cretaceous.
Argentina, Patagonian Spring hill Formation, first
record for Patagonia. Ref. to Riccardi. --1995
Schneider: Tendagurium Dietrich, 1933. Mesozoic
subgenus of Cardium. Salisbury (1934) designated
Cardium propebanneainum [err. pro propebanneianum] Dietrich, 1933, as the type species. --1995
Schneider: Hayami (1958) considered the name
Tendagurium unavailable. - Tendagurium is valid, and
the taxon is considered to have been erected by
Salisbury (1934).
tenellum Wood, 1840. - Combination to be revised:
"Cardium " tenellum Wood, 1840. Cardiidae. Fossil.
RECORDS. --1922-32 Sherborn, p. 6409 : Cardium
tenellum Wood, 1840.
tenera (subspecies). - See under corbuloides tenera
Taktakishvili, 1978.
tenerum Solander, 1786. - Not valid. » Regarding
Cardium tenerum: Cardiidae. Recent. Synonym of
Fulvia aperta (Bruguiere, 1789). OTHER JUDGEMENTS:
Fulvia bullata (Linnaeus, 1758). - Fulvia aperta
(Bruguiere, 1789). - Nomen nudum. REMARKS. Cardium
tenerum was listed by Dillwyn (1817) with reference
to Cardium ruga tum Solander's MSS. No details are
available about the species. RECORDS. --1817 Dillwyn:
Cardium tenerum Solander's MSS = Cardium rugatum
Gronovius. "1922-32 Sherborn, p. 6412: Cardium
tenerum Solander, 1786 [n.n.]. "1945 Smith: Cardium
tenerum 'Solander'Diliwyn 1817 syn. = Fulvia bullata
Linne . --1977 Fischer-Piette: Cardium tenerum
798
Tropical Marine Mollusc Programme (TMMP)
tenue - tenuicostatum
Dillwyn = nomen nudum . "1992 Lamprell &
Whitehead: Cardium tenerum Dillwyn, 1817 = Fulvia
aperta (Bruguiere, 1789).
*tenlle. - Homonymy to be noticed. Cardium tenue
from 4 sources: Brown, 1827. - Sowerby, 1838. Orbigny, 1850. - Leach, 1852.
tenue Brown, 1827. - Nomen nudum but accompanied
by figure. » Regarding Cardium tenue: Cardiidae.
Recent. Synonym of Cerastoderma edule (Linne,
1758). REcORDs. "1852 Leach : Cardium tenue. "1867
Weinkauff: Cardium tenue Brown = Cardium edule
Linne. "1888 Kobelt: Cardium tenue Brown =
Cardium var. edule. "1922-32 Sherborn, p. 6422:
Cardium tenuis Brown, 1827 [n. et f.].
tenue Jekelius, 1944. - New combination:
Lymnocardium tenue Jekelius, 1944. Lymnocardiidae.
Miocene. RECORDS. "1946 ZR: Limnocardium tenue
Jekelius, 1944 sp. nov. Miocene Soceni Banat.
tenue Leach, 1852. - Insufficiently known species. »
Regarding Cardium tenue: ?Cardiidae. Recent. TYPE
LOCALITY. Seas of Great Britain. RECORDS. "1852 Leach,
p. 319: Cardium tenue. "1977 Fischer-Piette: Cardium
tenue Leach = species insufficiently known. "1980
Ruhoff: Cardium tenue Leach in J. Gray, 1852; p. 319.
- CHARACTERS AND OCCURRENCE - DIMENSIONS. Length I inch and three-eighths. REMARKS. The
specific name indicates that the species has a thin shell.
Fischer-Piette (1977) referred to numerous ribs, but
since Lea did not compare with other cardiids, the exact
nature of Lea's species is cryptic.
tenue Orbigny, 1850. - Combination to be revised:
"Cardium" tenue d'Orbigny, 1850. Cardiidae. Fossil.
RECORDS. "1922-32 Sherborn, p. 6422 : Cardita tenuis
Klipstein, 1845. "1922-32 Sherborn, p. 6422 : Cardium
(Klip.) tenue d'Orbigny, 1850. - Cardita tenue.
tenlle Sowerby, 1838. [Figs. Part I p. 112] . - Synonym
of Fulvia tenuicostata (Lamarck, 1819). OTHER
JUDGEMENTS: Fischer-Piette (1977) suggested Cardium
tenue Sowerby = Cardium tenuicostatum Lamarck =
Papyridea papyraceum (Chemnitz) Schroter [not
available] . » Regarding Cardium エ ・ ョセ ・ Z@ Cardiidae.
Recent. RECORDS. "1838 Sowerby, 149t part, No. 36:
Cardium tenue nobis. "1840 Sowerby, corrected list
p. 8: Cardium tenue nobis = Cardium tenuicostatum
Lam. 001840 Sowerby, corrected list, p. 8: Cardium
tenuicostatum (Sowerby No.8). "1977 Fischer-Piette:
Cardium tenue Sowerby = Papyridea papyraceum
(Chemnitz) Schroter.
tenue Stephenson, 1941. - Brevicardium tenue Stephenson, 1941. Cardiidae. Cretaceous. RECORDS. "1946 ZR:
Brevicardium tenue Cretaceous, Texas, Neylandsville
marl 2 1/2 m N. of Corbet. 001947 ZR: Brevicardium
tenue Stephenson, 1941 sp. nov. Navarro Group, Upper
Cretaceous, Texas.
tenue Vu Khuc 1991. - Vietnamicardium tenue Vu Khuc
1991. Cardiidae. Triassic. TYPE LOCALITY . Vietnam.
RE CORD S. .. 1991 Vu Khuc et. al. p . 10 I , fig :
Vietnamicardium tenue Vu Khuc 1991; sp. nov.
tenue beta Leach, 1852 (variety). - Insufficiently known
species . » Regarding Cardium tenu e var. beta :
?Cardiidae. RECORDS. 001852 Leach: Cardium tenue var.
beta Leach, 1852.
tenuicostata (mode and variety). - See under
tenuicostatum Lamarck, 1819 and planicostata var.
tenuicostata Andrusov, 1964.
tenuicostata Gillet, 1970. - Monodacna tenuicostata
Gillet, 1970. Lymnocardiidae. Miocene. RECORDS .
001970 ZR: Monodacna tenuicostata Gillet, 1970 sp.
nov. Miocene, ' Monte dei Corvi' deposits, Ancona
region, Italy.
tenuicostata Gillet, 1970. - Pontalmyra tenuicostata
Gillet, 1970. Lymnocardiidae. Miocene. RECORDS.
001970 ZR : Pontalmyra tenuicostata Gillet, 1970 sp.
nov. Miocene, 'Monte dei Corvi' deposits, Ancona
district, Italy.
*tenuicostatum. - Homonymy to be noticed. Cardium
tenuicostatum from 2 sources: Lamarck, 1819. Gumbel, 1861 .
tenuicostatum Gumbel, 1861. - Combination to be
revised: "Cardium " tenuicostatum Giimbel, 1861.
Cardiidae. Fossil. RECORDS. 001980 Ruhoff: Cardium
tenuicostatum Gumbel , 1861 ; p. 743 .
tenuicostatum Lamarck, 1819. [Figs. Part I p. 119, 133 ,
174. Part 3 p. 891 , 929]. - Changed combination: Fulvia
tenuicosta (Lamarck, 1819). OTHER JUDGEMENTS: Fulvia
australe. - Papy ridea papyra ceum. « Regarding
Cardium tenuicostatum : Cardiidae. Pliocene,
Quaternary and Recent. TYPE LOCA LITY . Timor and
Australia. TYPE REPOSITORY. National Museum in Paris,
unknown availability. Lectotype designated by Vidal
1994, p. 101. RECORDS. 001841 Lamarck, pI. II, fig. 6
a, b, c: Cardium tenuicostatum. 001844 Reeve, species
50. Cardium tenuicostatum Lamarck, Anim. sans vert.
(Deshayes' edit.), vol. vi . p . 372. 001865 Angas:
Papyridea tenuicosta Lam. 001867 Angas: Cardium
(Papyridium sic) tenuicostatum Lam. "1868 ZR:
Cardium tenuicostatum (Lam.). Cardium radiatum
(Reeve) seems to be a variety [Ref. Romer 1868].
001869 Issei: Cardium tenuicostatum ? Lamarck. 001871
Fischer: Cardium tenuicostatum Lamarck. 001885
Smith: Cardium (Bucardium) tenuicostatum Lamarck.
001890 Paetel: Cardium tenuicostatum Lm. 001898
Bavay: Cardium tenuicostatum Lmk. probably Fulvia
australe. 001899 Sturany : Cardium tenuicostatum Lm.
001905 Sturany: Cardium tenuicostatum Lm. 00192232 Sherborn, p. 6418: Cardium tenuicostatum Lamarck,
1819. "1932 Lamy: Cardium (Papyridea)
tenuicostatum Lk. 001932 Prashad: Cardium (Laevicardium) tenuicostatum . .. 1942 ZR: Cardium
tenuicostatum sterol fraction content [Ref. Fantl 1942].
001945 Smith: Fulvia tenuicostata Lamarck, 1819.
001961 Cotton: Fulvia tenuicostata Lamarck 1819;
Pliocene, South Australia. 001974 Habe & Kosuge:
Fulvia tenuicostata (Lamarck, 1822). 001977 FischerPiette: Cardium tenuicostatum Lamarck = Papyridea
papyraceum (Chemnitz) SchrNer. 001977 Wilson &
Stevenson: Fulvia tenuicostatum (Lamarck, 1819).
001992 Oliver: Cardium tenuicostatum ? Lamarck in
IsseI (1869) = Fulvia australe. 001992 Oliver: Cardium
tenuicostatum Lamarck in Fischer 1871 = Fulvia
australe. 001992 Oliver: Cardium tenuicostatum Lm.
in Sturany 1899 = Fulvia australe. 001992 Oliver:
Cardium tenuicostatum Lm. in Sturany 1905 = Fulvia
australe . 001992 Lamprell & Whitehead : Fulvia
tenuicostatum . 001994 Vidal : Cardium tenuicostatum
Lamarck, 1819; lectotype designated, p. 10 I. 00 I 994
Vidal: Fulvia (Fulvia) tenuicosta (Lamarck, 1819).
001999 Jacques Vidal (pers. com.) MNHN, Paris:
Phuket Marine Biological Center Special Publication vol. 30 (2004)
799
tenuicostatum - tenuo
Fossils from the Paleontological Collection as Cardium
tenuicostatum Lamarck. Quaternary. Egypt (Suez),
B42200. ··2001 Vidal, p. 408: Fulvia tenuicostata. CHARACTERS AND OCCURRENCE - DESCRIPTIONS. ··1844 Reeve: shell transversely ovate, rather
oblique, somewhat thin, ventricose, radiately very
finely ribbed, sometimes 40 smooth ribs, sometimes
about 52-53 ; whitish, ribs variegated in a waved manner
with small flesh-colour spots, umbones rose-purple.
"1972 Cernohorsky: 45-55 feeble radial ribs. "1819
Lamarck: 48 ribs. "2001 Vidal: tentacles numerous.
Eyes on tip of tentacles. DIMENSIONS. To 60 mm in
length. HABITAT. Coral sand, 10-20 m. DISTRIBUTION.
··1819 Lamarck: gives reference to Timor (Indonesia)
and New Holland. Reference to Timor is erroneous
according to Stephen & Wilson. Lamarck specifies that
specimens from Australia are slightly more
inequilateral, so he must have had specimens from
Timor to compare with. "1819 Lamarck p. 5, No.5:
Cardium tenuicostatum . "1844 Reeve: New Holland
= Western Australia. '·1897 Sowerby: Natal South
Africa. ··1994 Hylleberg: Western Australia.
tenuicostatum of Reeve. - Misidentification. Cardiidae.
Recent. RECORDS. "1844 Reeve, sp. 50: Cardium
tenuicostatum. '·1897 Sowerby: Cardium
tenuicostatum Lamarck with reference to Reeve fig.
50. "1903 Hidalgo: Cardium lenuicostatum in Reeve,
fig. 50 does not show tenuicostatum Lamarck.
tenuicostatum of Sowerby. - Misidentification of
Papyridea papyracea . OTHER JUDGEMENTS: Fulvia
australe. » Regarding Cardium tenuicostatum:
Cardiidae. Recent. DISTRIBUTION. Australia (Grasset
1884). REcoRDs. ·'1840. Sowerby, 182nd part, fig. 62:
Cardium lenuicostalum Lam. ··1840 Sowerby,
catalogue p. I, No. 8: Cardium tenui-costatum, Lam.
"1840 Sowerby, corrected list, p. 8: Cardium
tenuicostatum (Sowerby No.8). "1858 Adams &
Adams, p. 457: Cardium (Fulvia) tenuicostala, Sow.
··1870 Macandrew: Papyridea (Fulvia) lenuicostatum
Sow. "1884 Grasset: Cardium lenuicoslatum. Sow.
·'1974 Barnard: Cardium tenuicoslatum in Sowerby
1897 = Papyridea papyracea. '·1903 Hidalgo: Cardium
tenuicostatum in Sowerby, figs . 19 and 36 do not show
tenuicostatum Lamarck. "1992 Oliver: Papyridea
(Fulvia) tenuicostatum Sow. in Macandrew 1870 =
Fu/via australe.
tenuilamellosum Poutiers, 1981. [Figs. Part 3 p. 90 I] . Microcardium tenuilamellosum Poutiers, 1981.
Cardiidae. Recent. TYPE REPOSITORY. Holotype at
MNHN . TYPE LOCALITY. North of Lubang Island,
Philippines, 186-193 m. RECORDS ... 1981 Poutiers:
Microcardium tenuilamellosum. ·'1981 ZR:
Microcardium tenuilamellosum Poutiers, 1981 sp. nov.
Philippines. Ref. to Poutiers p. 338.
tenuis (mode). - See under tenue.
tenuisquamosum Jekelius, 1944. - New combination:
Lymnocardium tenuisquamosum Jekelius, 1944.
Lymnocardiidae. Miocene. RECORDS. ' ''1946 ZR:
Limnocardium tenuisquamosum Jekelius, 1944 sp. nov.
Miocene Soceni Banat.
tenuissima Ebersin, 1956. - Chartoconcha tenuissima
Ebersin, 1956. Cardiacea. Pliocene. RECORDS. ·'1956
ZR: Chartoconcha tenuissima Ebersin, 1956 sp. nov.
Pliocene. Kerchenski, Crimea Peninsula, Russia.
tenuissimum (subspecies). - See under barzaviae
tenuissimum Andreescu, 1975.
tenuissubquadratum Coen, 1915. - Infrasubspecific. No
evidence for subspecific status. Cardiidae. Recent.
DISTRIBUTION. Unknown (Nordsieck 1969). RECORDS.
·'1915 Coen: Cerastoderma glaucum tenuissubquadratum. ··1969 Nordsieck: Cerastoderma
glaucum tenuis-subquadratum (Coen, 1915).
*tenuistriatum . - Homonymy to be noticed. Cardium
tenuistriatum from 2 sources: Munster in Goldfuss,
1837. -Anton 1839.
tenuistriatum Anton, 1839. - Combination to be revised:
"Cardium" tenuistriatum. Cardiidae. Tertiary. RECORDS.
"1839 Anton: No . 389: Cardium tenuistriatum mihi.
"1922-32 Sherborn, p. 6427: Cardium tenuistriatum
Anton, 1839.
tenuistriatum Munster in Goldfuss, 1837. -Combination
to be revised: "Cardium" tenuistriatum Munster in
Goldfuss, 1837. "1922-32 Sherborn, p. 6427: Cardium
tenuistriatum Muenster in Go1dfuss, 1837.
tenuisulcatum Lamarck. - See under tenuisu/catum
Nyst, 1836.
*tenuisulcatum. - Homonymy to be noticed. Cardium
tenuisulcatum from 2 sources: Nyst, 1836. - Munster,
1840.
tenuisulcatum Munster, 1840. - Replaced by "Cardium"
comptum. Deshayes. ?Cardiidae. Fossil. RECORDS.
"1857. Deshayes: Cardium tenuisulcatum Munster,
1840. - C. comptum. Desh. '·1922-32 Sherborn, p.
6429 : Cardium tenuisu/catum Muenster, 1840.
tenuisulcatum Nyst, 1836. - Changed combination:
Laevicardium tenuisulcatum (Nyst, 1836). OTHER
JUDGEMENTS: Cardium cingulatum Goldfuss, 1840. »
Regarding Cardium tenuisulcatum: Cardiidae .
Oligocene . RECORDS. ··1844 Potiez & Michaud :
Cardium tenuisulcatum Nyst. fossil of Belgium. ·'1857.
Deshayes: Cardium tenuisulcatum Nyst, 1835 sic.
"1886. Vincent: Cardium tenuisulcatum Nyst =
Cardium cingula tum Goldfuss, 1840. '·1922-32
Sherborn, p. 6429 : Cardium tenuisulcatum Nyst, 1836.
··1945 ZR: Cardium tenuisulcatum Lk. [Ref. Lamy
1945]. ·'1970 ZR: Laevicardium tenuisulcatum (Nyst,
1836) designated type sp. of Laevicardium (Habecardium) subgen. nov. Glibert & Van De Poel, 1970.
··1977 Popov: Nemocardium (Habecardium)
tenuisulcatum (Nyst, 1836). "1986 Gitton et al.:
Laevicardium (Habecardium) tenuisulcalum Nyst.
Oligocene. ··1995 Schneider: Habecardium
tenuisulcatum (Nyst, 1836). ··1999 Jacques Vidal (pers.
com.) MNHN, Paris: Fossils from the Paleontological
Collection as Cardium tenuisulcatum Nyst. Tongrien.
Germany, 24904 to -6 . France (Bassin de Paris),
B52833 to -43.
tenuitesla Cossmann, 1908. - Combination to be revised:
"Cardium" tenuitesta Cossmann, 1908. Cardiidae.
Tertiary. RECORDS. "1909 ZR: Cardium
(Plagiocardium) lenuilesta Cossmann, 1908 sp. n.
Belgium Tertiary. ·'1975 ZR: Cardium (Plagiocardium) tenuitesta Cossmann, 1908. Lectotype
designated . p. 34. Ref. to Glibert.
tenuitesta. - Parvicardium tenuitesta. Cardiidae .
RECORDS. ·'1979 ZR: Parvicardium tenuitesta. Lower
Palaeocene. Ukraine. Ref. to Makarenko.
tenuo Meuschen, 1778. Non binominal. - Not available.
800
Tropical Marine Mollusc Programme (TMMP)
terquemi - thaumastum
"1902 Sherborn, p. 963: Cardium tenuo Meuschen,
1778.
terquemi Martin, 1863. - Combination to be revised:
"Cardium" terquemi 1. Martin, 1863. Cardiidae. Fossil.
RECORDs. "1980 Ruhoff: Cardium terquemi 1. Martin,
1863; vol. 7: 86, pI. 5, figs. 16-20.
tertiana Eberzin, 1956. - Oxydacna tertiana Eberzin,
1956. Lyrnnocardiidae. Pliocene. REcORDs. ··1956 ZR:
Oxydacna tertiana Eberzin, 1956 sp. nov. Pliocene
Russia Kuban Province.
terticostales (subspecies). - See under grande
terticostales Steininger, 1963.
tertium Fischer-Piette, 1977. - New name Laevicardium
tertium Fischer-Piette, 1977. Cardiidae. Recent.
DISTRIBUTION. Unknown. REMARKS. Fischer-Piette
(1977) states that Cardium fragile is preoccupied by
Brocchi, 1814, (Conchiologia fossile subappennina, II:
505 (pI. 13, Figs. 4a, b). There is a previous record of
Cardium fragile R6ding (Mus. Bolt.), but this is a
nomen nudum. RECORDS. "1977 Fischer-Piette, p. 28:
Laevicardium tertium new name for Cardium fragile
Sowerby, 1834. "1977 Fischer-Piette, p. 176 Laevicardium tertius nom. nov. "1977 ZR : Laevicardium
tertium Fisher-Piette, 1977 nom. nov. for Cardium
fragile Sowerby, 1834 nee Brocchi, 1814. Ref to
Fischer-Piette, E. p. 28. "1980 Keen: Laevicardium
(Fulvia) tertius Fischer-Piette, 1977,
teshimaense Saito, Bando & Noda, 1970. - Vasticardium
teshimaense Saito, Bando & Noda, 1970. Cardiidae.
Middle Miocene. RECORDS. "1970 ZR: Vasticardium
teshimaense Saito, Bando & Noda, 1970 sp. nov.
Middle Miocene Teshima 1., Shodo-Gun, Kagawa
Prefecture, Japan.
testata Kvaliashvili, 1978. - Digressodacna testata
Kvaliashvili, 1978. Lymnocardiidae. Pliocene .
RECORDS. "1978 ZR: Digressodacna testata
Kvaliashvili , 1978 sp. nov. Georgia SSR. Ref. to
Kvaliashvili p. 67.
*tetragonum. - Homonymy to be noticed. Cardium
tetragonum from 3 sources: Voigt in Cuvier, 1834. Michelin, 1838. - Munster, 1842.
tetragonum Michelin, 1838. - Replaced by Cardita
tenuicostata. d 'Orbigny. Not a cardiid. Fossil. RECORDS.
"1857 Oeshayes: Cardium tetragonum Michel. Cardita tenuicostata. d 'Orb. "1922-32 Sherborn, p.
6466: Cardium tetragonum Michelin, 1838.
tetragonum Munster, 1842. "Cardium" tetragonum
Munster, 1842. Cardiacea. Fossil. RECORDS. "1857.
Oeshayes: "Cardium " tetragonum Munster, 1842.
Ietragonum Orbigny, 1850. - Conocardium tetragonum
d'Orbigny, 1850. Conocardiidae. Fossil. RECORDS.
"1922-32 Sherborn, p. 6467 : Lunulacardium
tetragonum Muenster, 1840. ··1922-32 Sherborn, p.
6466: Conocardium (Muenster) tetragonum d'Orbigny,
1850. - Lunu/acardium.
tetragonum Voigt in Cuvier, 1834. - Synonym of
Fragum hemicardium (Linne, 1758). » Regarding
Cardium tetragonum: Cardiidae . Recent. OF
COLLECTIONS. One specimen at ZMUC . REMARKS.
Cardium tetragonum Voigt = Fragum hemicardium
(Linne , 1758) according to label in box with
hemicardium from Amboina at ZMUC. DISTRlBUTION.
East Indies (Romer 1891). RECORDS. '·1839 Anton:
Cardium tetragonum Voigt = Cardium hemicardium
Linne. "1890 Paetel: Hemicardium tetragonum Voigt.
"1891 Romer: Ctenocardia tetragonum Voigt. "192232 Sherborn, p . 6466: Cardium (Hemicardium)
tetragonum Voigt in Cuvier, 1834.
tetrameroide Ryckholt, 1860. - Conocardium tetrameroide Ryckholt, 1860. Conocardiidae. Fossil.
REMARKS. Previously in Cardiidae but Conocardiids
have no connection with the Cardiacea except analogy
of form, which does not offer any point of relationship
(Dall 1895, p. 519). RECORDS ... 1980 Ruhoff: Conocardium tetrameroide Ryckholt, 1860; [1852-1862] part
2: plate II, figs. 13-15.
tetrica Feruglio, 1937. - Lahillia (?) te/rica Feruglio,
1937. Cardiacea. Cretaceous. RECORDS. "1937. ZR:
Lahillia (?) telrica Feruglio, 1937 sp. n. Cretaceous
Argentina.
tevoroa. - Tridacna tevoroa. Tridacnidae. Fossil and
Recent. RECORDS. ··2002 Schneider: Tridacna tevoroa,
Tridacnidae . Fossil and Recent. The fossil record
extends back to the Tertiary (?Late Miocene). Hawaii .
tewartii Mathur, 1975. - Discors tewartii Mathur, 1975 .
Cardiidae. Lower Eocene . RECORDS. "1975 ZR:
Discors tewartii Mathur, 1975 sp. nov. Himachal
Pradesh, Simla Hills. Ref. to Mathur, p. 25.
texana Conrad, 1857. - Changed combination:
Protocardia lexana texan a (Conrad, 1857). Cardiidae.
Cretaceous. RECORDS. "1871 Stoliczka, 141: C.
texanum (genus Protocardium). "1980 Ruhoff:
Cardium (Protocardia) texanum Conrad, 1857; vol. 1:
150, pI. 6, figs. 6a-c. "1986 Scott: Cardium
(Protocardia) texana Conrad, 1857. "1986 Scott:
Protocardia lexana texana (Conrad, 1857).
texana of authors.- Misidentification of Protocardia
multistriata (Shumard, 1854). RECORDS. "1986 Scott:
Protocardia texana (Conrad, 1857) in Adkins & Winton
1920, Adkins 1928; Perkins 1961 = Protocardia
multistriata (Shumard, 1854).
texana salinaensis Meek, 1872 (subspecies). Protocardia texana salinaensis Meek, 1872. Cardiidae.
Cretaceous. RECORDS. "1986 Scott: Prolocardia texana
salinaensis Meek, 1872.
texana subspinigera Cragin, 1893 (subspecies). Protocardia lexana subspinigera Cragin, 1893.
Cardiidae. Cretaceous. RECORDS. "1986 Scott:
Protocardia texana subspinigera Cragin, 1893. "1987
ZR: Protocardia texana subspinigera Cragin, 1893.
Neotype designated. Ref. to Scott p. 1200.
textum Bronn, 1832. - Combination to be revised:
"Cardium" textum Bronn, 1832. Cardiidae. Fossil.
RECORDS. "1832 Bronn, No. 589, p. 610: Cardium
texlum n. sp. [fossil]. "1922-32 Sherborn, p. 6475:
Cardium textum Bronn, 183 I.
texturatum Munster, 1840. - Combination to be revised:
"Cardium" texturatum Munster, 1840. ?Cardiidae.
Fossil. RECORDS. "1922-32 Sherborn, p. 6475: Cardium
texturatum Muenster, 1840.
tha/assium Maury, 1925. - Changed combination:
Agnocardia tha/assium (Maury, 1925). Cardiidae.
Fossil. .. 2002 Schneider: Cardium (Trachycardium)
tha/assium Maury, 1925.
thaumasta (mode). - See under thaumastum Woodring,
1925.
thaumastum Woodring, 1925. - Changed combination:
Trigoniocardia thaumasta (Woodring, 1925) .
Phukel Marine Biological Center Special Publication vol. 30 (2004)
801
theidus - thurstoni
Cardiidae. Fossil. RECORDS. ··1925 Woodring, p. 144,
pI. 19, figs. 12-13: Cardium (Trigoniocardia)
thaumastum new species. ··1977 Popov: Trigoniocardia (Trigoniocardia) thaumasta (Woodring, 1925).
theidus (error). - Printer's mistake for thetidis in Wilson
& Stevenson 1977, Text Fig. 10.
thetidis Hedley, 1902. - Changed combination:
Pratulum thetidis (Hedley, 1902).
Cardiidae.
Recent. Type species : Iredale (1924) designated
Cardium thetidis Hedley, 1902 type species of the genus
Pratulum Iredale, 1924. Recognised only at the
subgeneric level by some authors. Keen (1954):
Cardium thetidis Hedley, 1902 is type species of
Pratulum Iredale, 1924. TYPE LOCALITY. A considerable
series from off Port Kembla in 63-75 fathoms, and off
Cape Three Points in 41-50 fathoms (Hedley 1902).
TYPE REPOSITORY. Australian Museum, Sydney. Syntype
COl3261. RECORDS. ··1902 Hedley, p. 322 : Cardium
striatulum var. thetidis Hedley var. nov. ··1924 Iredale
designated Cardium thetidis Hedley, 1902 . ··1924
lredale, p. 182 : Pratulum thetidis (Hedley, 1902).
··1924 ZR: Cardium thetidis replaced by Pratulum
thetidis lredale, 1924. ··1938 Cotton & Godfrey:
Pratulum thetidis Hedley 1902. ··1950 Allan: Pratulum
thetidis. ··1950 Keen: Nemocardium (Pratulum)
thetidis (Hedley, 1902). ··1954 Keen: Cardium thetidis
Hedley, 1902. ··1954 Keen: Nemocardillm (Pratulum)
thetidis (Hedley) , 1902. ··1961 Cotton: Cardium
thetidis Hedley 1902. ··1962 Macpherson & Gabriel :
Pratulum thetidis Hedl. ··1962 Iredale & MacMichael :
Cardium thetidis Hedl. ··1969 Moore et al.:
Nemocardium (Pratulum) thetidis. ··1977 FischerPiette: Pratulum thetidis Hedley = Pratulum striatulum
Sowerby. ··1977 Popov: Pratulum thetidis (Hedley,
1902). Type of Pratulum Iredale, 1924. ··1977 Wilson
& Stevenson : Nemocardium (Pratulum) thetidis
(Hedley, 1902) [text fig. 10 theidus err. pro thetidis].
··1977 Popov: Pratulum thetidis (Hedley, 1902). ··1979
Powell: Nemocardium thetidis . ··1992 Lamprell &
Whitehead : Nemocardium (Pratuillm) thelidis. ··1995
Schneider: Nemocardium (Pratulum) thetidis (Hedley,
1902). - CHARACTERS AND OCCURRENCE - OF
COLLECTIONS. ··ZMUC Australia, C. thetidis 36 S 150
E, 65 fv, sandy mud. DESCRIPTIONS. ··1979 Powell:
delicate radial ribs. DIMENSIONS. ··1902 Hedley: to 12.5
mm in length and 11.5 mm in height. REMARKS. ··1902
Hedley: stated that "thetidis" appears specifically
inseparable from C. striatulum Sowerby. But the new
variety is diminutive, thin, delicate, the concentric
sculpture fainter, and the angle limiting the posterior
prickly area is less pronounced. lredale (\ 924) found
the difference described by Hedley (1902) to be quite
constant and accepted their value as specific, i.e. raised
the variety thetidis to the specific level. The new genus
was not defined . Iredale only stated: "The difference
between this style and typical Cardium is very great,
and I, therefore propose the new genus Pratulum,
naming Cardium thetidis Hedley as type. HABITAT.
··ZMUC: sandy mud bottom 130 m depth. ··1902
Hedley: 41-75 fathoms . DISTRIBUTION. Australia, New
Zealand.
thetironiae Lange, 1920. - Combination to be revised :
"Cardium" thetironiae Lange, 1920. Cardiidae. Jura.
RECORDS. ··1920 ZR : Cardium thetironiae Lange, 1920
Jura E. Africa.
thetkegyiense Cotter, 1923. - Combination to be revised:
"Cardium" thetkegyiense Cotter, 1923. Cardiidae.
Eocene. TYPE LOCALITY. Eocene ofBurrna (Cotter 1923).
RECORDS. ··1923 ZR: Cardium th etkegyinense sic
Cotter, 1923 sp. n. Eocene Assam. ··1930 Stewart, p.
258 : "Cardium" thetkegyiense sic Cotter, 1923.
thetkegyinense (spelling) . - See under thetkegyiense
Cotter, 1923.
thielei Fischer-Piette, 1977. - Synonym of Afrocardium
exochum (Melvill, 1906). OTHER JUDGEMENTS: New
name for Cardium radula Thiele & Jaeckel, 1931. Trachyeardium infantile (Nomura & Zinbo, 1834).»
Regarding Laevicardium thielei: Cardiidae. Recent.
TYPE LOCALITY . Dar es Salam, E. Africa. DISTRIBUTION.
£. Africa, Indo-Pacific. RECORDS. ··1977 Fischer-Piette:
Laevieardium (Traehyeardium) thielei new name for
Cardium radula Thiele & Jaeckel, 1931. ··1977 ZR:
Laevieardium (Trachycardium) thielei nom. nov. for
Cardium radula Thiele & Jaeckel, 1931 nec Deshayes,
Broderip & Sowerby. Ref to Fischer-Piette, E. p. 55.
··1980 Keen: Ctenocardia (Afroeardium) thielei
(Fischer-Piette, 1977). ··1981 Koyama et al.:
Laevicardium (Trachycardium) thielei Fischer-Piette,
1977 = Cardium (Trachycardium) infantile Nomura &
Zinbo, 1834 . ··1991 Voskuil & Onverwagt:
Laevicardium (Trachyeardium) thielei Fischer-Piette,
1977 ; junior synonym of Afroeardium exoehum
(Melvill, 1906). ··1999 Higo et al.: Laevicardium
thielei Fischer-Piette, 1977 = Afrocardium infantile
(Nomura & Zinbo, 1934).
tholi Stephenson, 1941. - Combination to be revised:
"Cardium " thoU Stephenson, 1941 . Cardiidae. Upper
Cretaceous. RECORDS. ··1946 ZR: Cardium (Granocardium) tholi Stephenson, 1941 sp. nov. Navarro
formations, Cretaceous, Texas, Nacatoch sand , vicinity
of Chatfield . ··1947 ZR: Cardium (Granoeardium)
thoU Stephenson, 1941 sp. nov. Upper Cretaceous,
Navarro Group Texas.
thomassini Vidal, 1998 . [Figs. Part 3 p . 868] . Vasticardium thomassini Vidal, 1998 . Cardiidae.
Recent. DISTRIBUTION. South Africa (Zulu1and, 45-78
m). RECORDS. ··1998 Vidal, p. 123-124: Vastieardium
thomassini sp. nov. ··1999 Vidal, p. 269: Vastieardium
thomassini Vidal, 1998.
thunense Mayer, 1866. - Combination to be revised:
"Cardium " thunense Mayer, 1866 . Cardiidae.
Miocene. RECORDS. ··1902 Dollfus & Dautzenberg, p.
327: Cardium thunense Mayer, 1866.
thurstoni Dall, Bartsch & Rehder, 1938. [Figs. Part I p.
145]. - Synonym of Fragum mundum (Reeve, 1845).
OTHER JUDGEMENTS: Fragumfragum (Linnaeus, 1758).
» Regarding Fragum thurstoni: Cardiidae. Recent.
RECORDS . ··1938 Dall, Bartsch & Rehder, p. 154, pI.
41, figs. 9-10: Fragum thurstoni new species. ··1939
ZR: Fragum thurstoni Dan, Bartsch & Rehder, 1938.
sp. n. Keokia. Hilo, Hawaii. ··1977 Fischer-Piette:
Cardium thurstoni DaU, Bartsch & Rehder = Coreulum
fragum Linne. ··1977 Habe: Fragum thurstoni Dall,
Bartsch & Rehder, 1938 = Fragum mundum (Reeve,
1865). ··1979 Kay: Fragum thurstoni Dall, Bartsch,
and Rehder, J 938 Uun. syn.) = Fragum (Fragum)
mundum (Reeve, J845). ··1999 Higo et al.: Fragum
thurstoni Dall, Bartsch & Rehder, 1938 = Fragum
802
Tropical Marine Mollusc Programme (TMMP)
thaanumi - tintinnabularum
mundum (Reeve, 1845). - CHARACTERS AND
OCCURRENCE - OF COLLECTIONS. ··ZMUC: one
specimen F. thurstoni: Beach collection on Oahu,
Hawaii. Related to Fragum mundum Reeve, 1845 and
Fragum carinata Lynge, 1909. DISTRIBUTION. ··ZMUC:
Fragum thurstoni: Hawaii; Fragum mundum: West
Pacific, Hawaii.
thaanumi Pilsbry, 1920. - Changed combination:
Nemocardium thaanumi (Pilsbry, 1920). Cardiidae.
Recent RECORDS ... 1920 Pilsbry, p. 323-324, pI. 12,
fig. 24: Cardium thaanumi n. sp. ··1920 ZR: Cardium
thaanumi Pilsbry, 1920 sp. n. Hawaii. "1938 Dall,
Bartsch & Rehder: Microcardium thaanumi Pilsbry
1921. ··1964 ZR: Microcardium thaanumi photo. [Ref.
Weaver 1964]. "1977 Fischer-Piette: Microcardium
thaanumi Pilsbry. ··1977 Popov: Pratulum thaanumi
(Pilsbry, 1921). ··1977 Wilson & Stevenson:
Nemocardium (Microcardium) thaanumi Pilsbry, 1921.
"1979 Kay: Nemocardium thaanumi (Pilsbry, 1921).
- CHARACTERS AND OCCURRENCE - DESCRIPTIONS. ··1979 Kay: shell hemispherical, thin, equivalve;
anterior margin a little more strongly curved than
posterior; with slender radiating threads crossed by
concentric threads; white. Hinge: narrow; left valve
with two erect cardinals and elongate laterals; right
valve with the ventral cardinal larger than the dorsa!.
Sculpture: numerous (135 or more) slender radiating
ribs separated by impressed interspaces crossed by
regularly spaced concentric threads with elevated spines
at the junctions. Color: white, the external sculpture
showing through. DIMENSIONS. ··1979 Kay: Height, 7.5
mm; length, 7 mrn; diameter, 6 mrn. REMARKS. ··1979
Kay: N. thaanumi was described from the Hawaiian
Islands; shells similar to the Hawaiian shells have been
dredged in the Marquesas (Rehder cited by Kay 1979).
HABITAT. ··1938 Dall, Bartsch, and Rehder: these
cardiids occur at depths of from 16 m to 200 m, with
one report of valves at depths of 500 m. ··1979 Kay:
they are not common; occurring in fewer than one
percent of the dredge hauls of both the Pele and the
Albatross, but appear to be most frequent at depths of
25 to 50 m. DISTRIBUTION. Tropical Pacific, Hawaii.
tibisii Jekelius, 1944. - Replidacna tibisii Jekelius, 1944.
Lymnocardiidae. Miocene. RECORDS. ··1946 ZR:
Replidacna tibisii Jekelius, 1944 sp. nov. Soceni Banat
"1977 ZR: Cardium (Replidacna) tibisii lekelius. Ref
to Paramonova p. 70 [Trans!. p. 320].
tigilense Siodkewitsch, 1938. - Keenocardium tigilense
(Slodkewitsch, 1938). Cardiidae. Fossi!. RECORDS.
"1990 Gladenkov & Sinelnikova: Keenocardium
tigilense (Slodkewitsch), 1938. Laevicardium tigilense
Siodkewitsch, 1938.
tigrina Martyn, 1784. Non binominal. - Not available.
Not a cardiid. RECORDS. ··1784 Martyn: Cardium
tigrina.
tilimbergense Doderlein 1862. Identical with
Lymnocardium secans (Fuchs, 1870). » Regarding
Hemicardium tilimbergense: Lymnocardiidae. Tertiary.
RECORDS. ··1899 Sacco, p. 58: Hemicardium
tilimbergense Doderlein 1862 = Lymnocardium secans
(Fuchs, 1870).
timisense Jekelius, 1944. - New combination:
Lymnocardium timisense Jekelius,
1944.
Lymnocardiidae. Miocene. RECORDS. "1946 ZR:
Limnocardium timisense Jekelius, 1944 sp. nov.
Miocene Soceni Banat. ··1987 Taktakishvili: Limnocardium timisense Jek.
timorensis Krumbeck, 1924. - Protocardia timorensis
Krumbeck, 1924. Cardiidae. Trias. RECORDS. "1924
ZR: Protocardia (?lsocyprina) timorensis Krumbeck,
1924 sp. n. Trias Timor.
tinctum (mode and variety). - See under tincta Dall, 1881
and peramabile var. tinctum Dall.
tinctum Dall, 1886. - Changed combination:
Nemocardium tinctum (Dall, 1886). Cardiidae. Recent.
TYPE LOCALITY. St. Croix, West Indies. RECORDS. ··1881
Dall: Cardium peramabilis var. tinctum. ··1886 Dall,
p. 270: Cardium peramabilis var. tinctum. ··1900 Dall:
Protocardia tincta . .. 1902 Dall: Protocardia tincta
Dall, 1886 (quite distinct, not a variety) ... 1902 Dall
& Simpson, pI. 58, fig. 4: Protocardia tincta ... 1934
Johnson: Protocardia tincta Dall 1886. ··1939 McLean:
Microcardium tinctum. "1939 ZR: Microcardium
tinctum fig. [Ref. McLean 1939]. ··1944 ZR:
Microcardium tinctum Dall, fig. [Ref. Clench & Smith
1944]. "1968 Kempf & Matthews: Microcardium
tinctum (Dall, 1881). ··1970 Rios: Microcardium
tinctum (Dall, 1881). "1971 Altena: Nemocardium
tinctum . .. 1974 Abbott: Nemocardium tinctum (Dall,
1881) ... 1977 Popov: Pratulum tincta (Dall, 1886).
"1997 Shelton: Nemocardium tinctum (Dall, 1881).CHARACTERS AND OCCURRENCE - OF COLLECTIONS. ··ZMUC: S.w. of Mississippi delta, Gulf of
Mexico 88 m. CHARACTERS of diagnostic value: minute
radiating ribs. DESCRIPTIONS. "1902 Dall & Simpson:
shell small, fragile, nearly equilateral, considerably
inflated; posterior area sculptured with minute,
radiating ribs; every second, third, fourth and fifth one
bears a row of sharp, elevated spines or scales; the
spaces between theses ribs are crossed by delicate
scales, making them pitted; the remainder of the shell
is sculptured with very finely beaded ribs, as in P
peramabilis; borders of the shell finely and sharply
serrate, not gaping; outside sculpture showing faintly
within; anterior laterals showing as long arched ridges
within, entering far within the cavity of the beaks.
Colour whitish, beautifully tinted and clouded with pink
or red. DIMENSIONS. ··1902 Dall & Simpson: length 13,
height 12, diameter 9 mm. "1939 McLean: 13 x 12 x
9 mrn. REMARKS. ··1902 Dall & Simpson: P tincta is
certainly different from P peramabilis. It is not so high
in proportion to its width, is a little less inflated, the
line between the two areas of sculpture is not so
distinctly marked out, and when perfect it has the scaly
spines on the posterior area finely developed. The
anterior lateral teeth are different from those of P
peramabilis. HABITAT. ··ZMUC: 70-88 m depth.
DISTRIBUTION. ··1939 McLean: Key West, Florida, the
West Indies; "Of authors: Brazil; Mississippi Delta,
Gulf of Mexico.
tinrhertense Collignon, 1971. - Trachycardium
tinrhertense Collignon, 1971. Cardiidae. Upper
Cretaceous. RECORDS. "1971 ZR: Trachycardium
tinrhertense Collignon, 1971 sp. nov. Upper Cretaceous
Sahara desert.
tintinnabularum Maury, 1917. - Combination to be
revised: "Cardium" tintinnabularum (Maury, 1917).
Cardiidae. Fossil. TYPE LOCALITY. Restriction of type
Phuket Marine Biological Center Special Publication vol. 30 (2004)
803
tinyeanum - torii
locality Vokes 1989, p.l08. RECORDS. "1919 ZR:
Cardium (Trachycardium) tintinnabularum Maury,
1919 sp. n. Tertiary S. Domingo. "1922 Pilsbry:
Cardium tintinnabularum Maury. "1989 Vokes, p. 108:
Cardium (Trachycardium) tintinnabularum Maury,
1917.
tinyeanllm Loer. - New combination: Lymnocardium
tinyeanum (Loer.). Lymnocardiidae. Fossil. RECORDS.
"1987 Taktakishvili: Limnocardium tinyeanum Loer.
tippanllm Conrad, 1858. - Combination to be revised:
"Cardium" tippanum Conrad, 1858. Cardiidae. Fossil.
RECORDS. "1871 Stoliczka, 126: C. tippanum (subgenus
Acanthocardium sic). "1980 Ruhoff: Cardium
tippanum Conrad, 1858; vol. 3: 326, pI. 34, fig. 8b.
"1998 Schneider: Cardium tippanum Conrad, 1858.
Granocardium tippanum (Conrad, 1858).
tirolense Mayer & Gumbel in Gumbel, 1861. Combination to be revised: "Cardium " tirolense Mayer
& Gumbel in Gumbel, 1861. Cardiidae. Fossil.
RECORDS. "1980 Ruhoff: Cardium tirolense Mayer &
Gumbel in Gumbel, 1861; p. 673.
Titanocardium Maxwell, 1978 (subgenus). - Cardiidae.
Tertiary. GENOTYPE. Cardium (Trachycardium)
cantuariense Laws, 1933, original designation.
RECORDS. "1978 Maxwell, p. 18: Subgenus Titanocardium n. sg. "1978 ZR: Titanocardium New
subgenus. Hedecardium (Titanocardium). Ref. to
Maxwell. "1980 Vokes: Maxwell, 1978. Fossil. ··2002
Schneider: Titanocardium Maxwell, 1978 subgenus of
Hedecardium Marwick, 1944. Spines absent.
titthus Krishtofovich 1969. - Synonym of S.
groenlandicus (Bruguiere 1789). » Regarding
Serripes tiuhus: Cardiidae. Fossil. RECORDS. "1980 ZR:
Serripes tiUhus Krishtofovich 1969 is a junior synonym
of S. groenlandicus (Bruguiere 1789). Ref. to Kafanov.
tkalad Brus. - New combination: Lymnocardium tkalaci
(Brus.). Lymnocardiidae. Fossil. RECORDS. "1987
Taktakishvili : Limnocardium tkalaci Brus.
tobaruense Noda, 1988. - Frigidocardium tobaruense
Noda, 1988. Cardiidae . Pliocene. Noda (1988)
Frigidocardium tobaruensis Noda, 1988 is type species
of subgenus (Tobarum) Noda , 1988 of genus
Frigidocardium. TYPE LOCALITY. The Ryukyu Islands,
southwest Japan. REMARKS. Also spelt F (Tobarum)
tobanlensis. RECORDS. "1988 Noda p. 74, pI. 19, fig.
12a,b: Frigidcardium sic (Tobarum) tobaruense n. sp.
"1988 Noda: Frigidcardium sic (Tobarum)
tobaruensis. Type species of (Tobarum) . "1988 Noda:
Frigidocardium (Tobarum) tobaruense sp. nov. "1995
Schneider: Microcardium tobaruense (Noda, 1988).
"1995 Schneider: Tobarum Noda, 1988. Type species
Frigidocardium (Tobarum) tobaruense Noda, 1988.
Pliocene, Japan.
tobarllensis (mode). - See under tobaruense Noda 1988.
Tobarum Noda, 1988 (subgenus). - Cardiidae. Pliocene.
RECORDS. "1988 Noda: (Tobarum) Noda, 1988 n.
subgen. "1995 Schneider: Tobarum Noda, 1988. Type
species Frigidocardium (Tobarum) tobaruense Noda,
1988. Pliocene, Japan.
tocatum Nalivkin, 1934. - Conocardium tocatum
Nalivkin, 1934. Conocardiidae. Devonian. RECORDS.
"1935 ZR: Conocardium toeatum Nalivkin, 1934 sp.
n. Devonian S. Ural.
tochigiensis Hirayama, 1954. - Changed combination:
Vastieardium tochigiensis (Hirayama, 1954). Cardiidae.
Miocene. RECORDS. "1955 ZR: Vasticardium
tochigiensis Hirayama, 1954 sp. nov. Miocene, Tochigi
Prefecture, Japan . •• 1973 Keen: Vasticardium
tochigiense Hirayama, 1954. Miocene. [Cardium].
toerteliense (subspecies). - See under abichi toerteliense
Szeles, 1962.
tohanensis Papaianopol, 1974. - Pteradacna tohanensis
Papaianopol, 1974. Lymnocardiidae. Pliocene.
RECORDS . "1974 ZR: Pteradacna tohanensis sp. nov.
Romania, Prahova, Tohaneasa. Ref. to Papaianopol, p.
J 31.
tohanensis Papaianopol, 1977. - Plagiodacna tohanensis
Papaianopol, 1977. Lymnocardiidae. Pliocene.
RECORDS. "1977 ZR: Plagiodacna tohanensis Papaianopol , 1977 sp . nov. Romania, Pliocene. Ref. to
Papaianopol, I. p. 45, fig'd.
tohanensis Papaianopol, 1981. - Pontalmyra tohanensis
Papaianopol , 1981. Lymnocardiidae. Upper Miocene.
RECOROS . ··1981 ZR: Pontalmyra (Pontalmyra)
tohanensis Papaianopol, 1981 sp. nov. Romania. Ref.
to Papaianopol , p. 23.
tokunagai Yokoyama, 1922. - Changed combination:
Clinocardium tokunagai (Yokoyama, 1920). Cardiidae.
Pleistocene. REcORDs. "1922 ZR: Cardium tokunagai
Yokoyama, 1922 sp. n. Pleistocene Japan. "1927
Yokoyama: Cardium tokunagaiYok. (Up. Musashino).
"1973 Keen: Clinocardium tokunagai (Yokoyama,
1920) [Pleistocene?).
tokyoense Tokunaga, 1906. - Changed combination:
Vasticardium tokyoense (Tokunaga, 1906). Cardiidae.
Pliocene. TYPE LOCALITY. Pliocene of Tokyo (Vidal
1999). REMARKS. Tokunaga (1906) compared Cardium
tokyoensis with Acrosterigma burchardi but stated that
it was closer to Acrosterigma ul1icolor (Vidal 1999).
RECORDS. "1906 Tokunaga: Cardium tokyoensis n. sp.
post tertiary. "1906 ZR: Cardium tokyoensis Tokunaga,
1906 sp. n. Post Tertiary, Japan. "1973 Keen:
Vasticardium tokyoel1se (Tokunaga, 1906). "1990
Gladenkov & Sinelnikova: Vasticardium tokyoense
(Tokunaga). "1999 Vidal, p. 272 : Cardium tokyoensis
Tokunaga, 1906: 51 , pI. 3, figs 12a-a'.
tombecki Loriol. - Ul1icardium tombecki de Lorio!'
Cardiidae. Lias and Jurassic. RECORDS. "1999 Jacques
Vidal (pers. com.) MNHN, Paris: Fossils from the
Paleontological Collection. Ul1icardium tombecki de
Lorio!. Kimmeridgian. France (Bourges), S05175.
tomlini Cox, 1930. - Fragum tomlini Cox, 1930.
Cardiidae. Fossil. RECORDs. "1930 ZR: Fragum tomlini
Cox, 1930 sp. n. fossil, Mombassa.
toneanum Yokoyama, 1922. - New combination:
Cerastoderma toneanum (Yokoyama, 1922). Cardiidae.
Pleistocene. RECORDS. "1922 Yokoyama, pI. 13, figs.
6-7: Vel1ericardia toneanum. "1933 Nomura: Cardium
(7 Cerastoderma) toneanum (Yokoyama, 1922).
tongricum Bayan. - Synonym of Cardium kochi Semp.
» Regarding Cardium tongrieum: Cardiidae. Fossil.
RECORDs. "1886 Vincent: Cardium tongricum Bayan
= Cardium kochi Semp.
torii Nomura, 1933. - Changed combination:
Nemocardium torii (Nomura, 1933). Cardiidae. Mioand Pliocene. RECORDS. "1933 Nomura, p!. 2, figs. 6a,
b: Cardium (Nemocardium) torii n. sp. "1954 Keen:
Nemocardium (Nemocardium) torii (Nomura, 1933
804
Tropical Marine Mollusc Programme (TMMP)
toronglynicum - Trachycardilllll
sic). Japan . --1973 Keen: NemocardiulII (Nemocardium) torii (Nomura, 1933).
toronglynicum. -Combination to be revised: "Cardium"
toronglynicum. Cardiidae. Cenozoic. RECORDS. "1967
ZR: Cardium toronglynicum, Akchagy1 deposits
(Cenozoic), Azerbaidjan, SSR [Ref. Baba-Zade 1967].
torquota (variety). - See under tuberculatum var.
torquota .
torresi Smith, 1885. [Figs. Part 3 p. 900]. - Changed
combi. Frigidocardium torres[ (Smith, 1885). TYPE
LOCALITY. 'Challenger' sta. 188, south of New Guinea
09 59' 5, 139 42' E., 28 fms (Higo et at. 1999).
Cardiidae. Recent. RECORDS. --1885 Smith, p. 164-165,
pI. 8, figs. 4-4b: Cardium (Fragum) torresi, n. sp.
--1885 ZR: Cardium (Fragum) torresi Smith, 1885 sp.
n., S. of New Guinea. "1890 Paete1: Hemicardium
torresi Smith. --1931 Thiele & Jaeckel: Cardium
(Fragum) torresi E. Smith. "1932 Prashad: Cardium
(Pratttlum) torresi . .. 1977 Fischer-Piette: Microcardium torresi Smith. --1977 Wilson & Steven-son:
Nemocardium (Microcardium) torresi (Smith, 1885).
--1977 Wilson & Stevenson: Cardium (Fragum) torresi
Smith, 1885 . "1988 Drivas & Jay : Nemo-cardium
(Microcardium) torresi (Smith, 1885). --1992 Lamprell
& Whitehead: Frigidocardium torresi. --1999 Higo et
al.: Frigidocardium torresi (E. A. Smith, 1885). --2003
Hylleberg & Kilburn, p. 183: Nemocardium torresi
(Smith, 1885). - CHARACTERS AND OCCURRENCE - DESCRIPTIONS. Resembles Fulvia dulcis
(Deshayes, 1863) sensu Drivas & Jay (1988) but the
posterior end is more produced, and the coloration does
not include a violet umbonal area. DIMENSIONS. --1988
Drivas & Jay: ant.-post. 9.2 mm, umbo-ventr. 10 mm .
HABITAT. --1988 Drivas & Jay: Reunion, western Indian
Ocean at 54 m depth. --1999 Higo et at.: 30-200 m,
sandy mud. DISTRIBUTION. "1999 Higo et at. : North
Indian Ocean, West Pacific, Australia; Boso peninsula
(Tsuchida & Kurozumi 1986) and southwards, Japan
Sea (off central Niigata prefecture); East China Sea,
South China Sea, Australia, Persian Gulf, Indo-Pacific.
--2003 HylJeberg & Kilburn: Vietnam.
tortuoSllm Hermann, 1781. - Combination to be revised:
"Cardium" tortuosum Hermann, 1781. Cardiidae.
Fossil. RECORDS. "1902 Sherborn, p. 983 : Cardium
tortuosum Hermann, 1781 .
torui. - Changed combination: Nemocardium torui.
Cardiidae. Pliocene. RECORDS. --1933 ZR: Cardium
(Nemocardium) tonti Pliocene [Ref. Nomura 1933].
tosensis Kimura, 1956. - Protocardia tosensis Kimura,
1956. Cardiidae. Jurassic . RECORDS. --1973 Keen:
Protocardia (Protocardia) tosensis Kimura, 1956.
tOllcasianulII d'Orbigny, 1850. - Combination to be
revised: "Cardium " toucasianum d'Orbigny, 1850.
Cardiidae. Fossil. RECORDS ... 1871 Stoliczka, 46: C.
toucasianum (7 Acanthocardium sic). "1922-32
Sherborn, p. 6549: Cardium toucasianum d'Orbigny,
1850.
Touricardium (error). - Printer 's mistake for subgenus
Tauricardium in Celidje 1953.
tournaisense Branson, 1942. - New name. Conocardium
tournaisense Branson, 1942. Conocardiidae. Fossil.
RECORDS. --1942 ZR: Conocardium tournaisense
Branson, 1942 nom. nov. p. 389 for Conocardium
antiquum Koninck 1885.
touzini Cossmann, 1922. - Combination to be revised:
"Cardium" touzini Cossmann , 1922. Cardiidae.
Eocene. RECORDS. --1922 ZR : Cardium (Plagiocardium) touzini Cossmann, 1922 sp. n. Eocene.
Trachicardium (error). - Printer's mistake for
Tra chycardium in Romer, 1869 (Vokes 1980).
Trachycardiinae Stewart, 1930 (subfamily). Cardiidae. Eocene and Recent. RECORDS. --1936 ZR:
subfamily Trachycardiinae in Cardiidae [Ref. Keen
1936a] . "1980 Vokes: Subfamily Trachycardiinae
Stewart, 1930. --1980 ZR: Trachycardiinae Stewart
1930. Key to genera. Ref. to Keen. --1983 Anaya:
subfamily Trachycardiinae Stewart, 1930. "1994
Schneider: the subfamilies Trachycardiinae Stewart,
1930 and Cardiinae are synonyms. "1995 . Pacaud &
Renard: Subfamily Trachycardiinae. --1999 Higo et at.:
Trachycardiinae (subfamily) Stewart, 1930. --2000
Coan et al.: Subfamily Trachycardiinae Stewart, 1930.
--2001 Schneider & Carter: Trachycardiinae (subfamily) Stewart, 1930. --2002 Schneider: Trachycardiinae originate in the Palaeogene, not Cretaceous.
The monophyly of Trachycardiinae is not in doubt. CHARACTERS AND OCCURRENCE - DESCRIPTION .
"1994 Hylleberg : ovate but not oblique; rib
ornamentation normally of spines or imbricating scales
along posterior sides of ribs; hinge relatively short,
either nearly straight or sharply bent; cardinal teeth
unequal in size, A II with socket above; posterior margin
notched to digitate.
TrachycardiulII Morch, 1853 (genus). Cardiidae. Eocene
and Recent. GENOTYPE. Martens (1870) designated
Cardium isocardia Linne, 1758 type species of
subgenus Trachycardium Morch, 1853 of genus
Cardium Linne, 1758. Upgraded to generic level by
authors. RECORDS. --1876 Meek: Trachycardium,
Mbrch. Cardium allernatum , Sow. regarded as
genotype . "1876 Meek: Trachicardium [err. pro
Trachycardium] Morch, 1853 . Synon. - Cardiidae.
"1930 Stewart: Trachycardium Morch 1853. "1961
Olsson: Trachycardium Morch, 1853. --1977 Popov :
Trachycardium Morch, 1853. Genotype : Cardium
isocardia Linne, 1758. --1977 ZR: Trachycardium. Key
to subgenera. Ref. to Popov p. 46. -·1978 Maxwell:
Genus Trachycardium . ·-1980 Vokes: Trachycardium
Morch, 1853 in subfamily Trachycardiinae. --1983
Anaya: Trachycardium Morch, 1835 (sic = 1853). Type
species: Cardium isocardia Linne, 1758, designated by
von Martens, 1870. ·-1999 Vidal: Genus
Trachycardium Morch, 1853: 34. --2000 Coan et al.:
Genus Trachycardium Morch, 1853. -·2001 Vidal, p.
407, Trachycardium - CHARACTERS AND OCCURRENCE - DESCRIPTIONS. "1969 Moore et al. , emend.
Hylleberg 1994: medium to large shells. Outline ovate;
nearly equilateral, height greater than length; hinge
heavy, short, nearly straight, hinge line strongly arched.
Sculpture of prominent ribs or riblets ornamented with
spines, posteriorly arising from posterior edges of the
ribs, remainder smooth or with raised scales. Inner
margin crenulate. --2001 Vidal: tentacles numerous.
DISTRIBUTION. Tropical America.
Trachycardium Morch, 1853 (section and subgenus).Cardiidae. Eocene, Oligocene and Recent. GENOTYPE.
Martens (1870) designated Cardium isocardia Linne,
1758 type species of subgenus Trachycardium Morch,
Phukel Marine Biological Center Special Publication vol. 30 (2004)
805
transcarpaticum - transcendens
1853. RECORDS. "1858 Adams & Adams: subgenus
(Trachycardium) March. ··1869 ZR: subgenus
(Trachycardium) (March), type Cardium isocardia (L.)
[Ref. Romer 1869]. "1871 Stoliczka: (Trachycardium)
March, 1852 (?). Type not identified by Stoliczka.
"1886 Cossmann: Section (Trachycardium) Morch,
1853 of genus Cardium Linne, 1758. ··1887 Fischer:
(Trachycardium) March, 1853. Section of genus
Cardium Linne, 1758. "1890 Paetel: 3. Sect.
(Trachycardium) March of Cardium. "1891 Romer:
III. Sect. (Trachycardium) March of genus Cardium
L. "1899 Sacco: subgenus (Trachycardium) . "1922
Olsson: (Trachycardium) Moerch. "1924 Oldroyd :
Subgenus (Trachycardium) Morch, 1853. "1928
Kuroda: (Trachycardillm) March, 1853. "1934
Johnson: (Trachycardium) March 1853. ··1953 Olsson
& Harbison: subgenus (Trachycardium) March, 1853.
"1970 Glibert & Poel: subgenus (Trachycardium) .
··1999 Vidal: Subgenus (Trachycardium) March, 1853
of the genus Trachycardium with 4 living species and
at least 4 fossil nominal species. - CHARACTERS
AND OCCURRENCE - DESCRIPTION. "1969 Moore
et at.: sculpture of imbricating scales over entire shell,
reduced to beads anteriorly. West Indies-East and West
Central America-South America. REMARKS . The
designation "section" takes the value of subgenus.
*transcarpaticum. - Homonymy to be noticed.
Cardium transcarpaticum from 2 sources:
Grischkevitsch, 1955. - Merklin & Nevesskaya, 1959.
transcarpaticllm Grischkevitsch, 1955. - Combination
to be revised: "Cardium" transcarpaticum
Grischkevitsch, 1955. Cardiacea. Fossil. RECORDS.
"1985 Ionesi : Cardium transcarpaticum Grischkevitsch, 1955 . "1985 ZR: Cardium transcarpaticum
Grischkevitsch, 1955. Status. Ref. to Ionesi p. 15.
transcarpaticum Merklin & Nevesskaya, 1959. Combination to be revised: "Cardium" transcarpaticum Merklin & Nevesskaya, 1959. Cardiidae.
Miocene . RE CORDS. "1959 ZR: Cardium
transcarpaticum Merklin & Nevesskaya, 1959 sp. nov.
Turkmenistan, Miocene. "1965 ZR: Cardium
transcarpaticum figd. Sarmatian, Yugoslavia [Ref.
Spajic 1965] .
transcaspia Ebersin. - Changed combination:
Pontalmyra transcaspia (Ebersin). Lymnocardiidae.
Fossil. RECORDS. "1987 Taktakishvili: Pontalmyra
transcaspia (Ebers.).
transcaspica (subspecies). - See under catil/us transcaspica Nevesskaya, 1959.
transcaspica Eberzin, 1962. - Changed combination:
Hyrcania transcaspica Eberzin, 1962. LYJ1U1ocardiidae.
Pliocene. RECORDS. "1962 ZR: Didacna (Pontalmyra)
transcaspica Eberzin, 1962 sp. nov. Manghyshlak,
Azerbaidzhan, Azov region; Pliocene, Russia. "1977
Popov: Hyrcania (Hyrcanomya) transcaspica.
transcaucasiclIm Bobkova, 1956. - Combination to be
revised: "Cardium" transcaucasicum Bobkova, 1956.
Cardiidae. Cretaceous. "1956 ZR: Cardium transcaucasicum Bobkova, 1956 sp. nov. Armenia
Cretaceous. Caucasus, Russia.
transcaucasium (subspecies). - See under alatoplanum
transcaucasium Eberzin, 1956.
transcendens Melvill & Standen, 1899. - Changed
combination: Acrosterigma transcendens (Melvill &
Standen, 1899). Cardiidae. Recent. TYPE LOCALITY.
Holotype, Torres Strait. TYPE REPOSITORY. Holotype,
Natural History Museum, London. TYPE MATERIAL.
Holotype, a left valve, BMNH 1899.2.23.6 (Wilson &
Stevenson 1977). Paratype: A specimen from same
locality, NMW 55.158.696 Melvill colI. RECORDS.
"1899 Melvill & Standen, p. 191, pI. 11, fig. 21 :
Cardium transcendens . .. 1899 ZR: Cardium
(Trachycardium) transcendens Melvill & Standen,
1899 n. sp. "1909 Lynge: Cardium (Trachycardium)
transcendens.
"1909
Melvill:
Cardium
(Trachycardium) transcendens M. & St. "1960
Kuroda: Trachycardium (Acrosterigma) transcendens
(Melvill & Standen). "1977 Fischer-Piette: Cardium
transcendens Melvill & Standen = Cardium enode
Sowerby. "1977 Habe: Trachycardium (Vasticardium)
transcendens (Melvill & Standen, 1899). "1977 Wilson
& Stevenson: Acrosterigma transcendens (Melvill &
Standen, 1899). "1978 ZR: Trachycardium
transcendens Bay of Bengal Burma, Maungmagan.
First record for Burmese waters. Ref. to Ray, H.C.
"1999 Higo et al. : Acrosterigma (Vasticardium)
transcendens (Melvi II & Standen, 1899). "1993 Vida\:
Cardium transcendens not synonym of Cardium en ode
as suggested by Fischer-Piette. "1999 Vidal : Cardium
(Trachycardium) transcendens Melvill & Standen.
,,2001 Vidal, p. 407 : Acrosterigma transcendens. CHARACTERS AND OCCURRENCE - CHARACTERS
of diagnostic value. "1909 Melvill: distinguished by
its very fine longitudinal ribs, nearly 60 in number, and
bright coloration. DESCRIPTIONS. "1999 Vidal: Shell
medium to small, almost perfectly ovoid and equilateral
(anterior dorsal margin sometimes very slightly raised),
and never truncated; also weakly elongated and rather
globose. Lunule narrow, almost equal on both valves,
and poorly differentiated, with its umbonal margins
slightly raised . External colour is variable, often vivid
and bright, light beige-yellow variously spotted and
splashed with darker pink, orange, or brown. Interior
variable in colour, white to pink or orange, with two
umbonal rays which are often wide and well developed.
Hinge nearly symmetrical and moderately angled.
Mean rib number 64.4 ± 3.9, range 56-74. On posterior
slope, interstices narrow and both parts of ribs about
equivalent in width, with a variable axial furrow; scales
irregular, mainly tubercular, sometimes becoming very
small, or absent altogether. On posterior part of median
shell, ribs become more or less flatly triangular and
asymmetrical (the anterior flank shorter and steeper),
sometimes still bearing an axial furrow; wider posterior
flank smooth or bears elongated ridges which are
analogous to the scales of posterior slope, but more
numerous. On anterior part of median shell, axial
furrow and posterior long ridges eventually disappear
and ribs become ーイッァ・ウゥセャケ@
more symmetrical and
slightly rounded, ornamented only with posterior
crenulations; interstices become wider and often very
finely notched. On anterior half of shell, ribs at first
remain rounded and retro crenu1ated, and then become
somewhat top-ridged; on first anterior ribs, top ridges
become tubercular and irregular, tending to be arranged
in concentric alignments which can replace the
longitudinal ribbing. "200 I Vida\: tentacles numerous.
No eyes on tentacles. DIMENSIONS ... J999 Vidal : length
806
Tropical Marin e Mollusc Programme (TMMP)
transclathratum - transversum
27.1 mm, height 30.2 mm, width 18.9 mm. REMARKS.
A. transcendens is very close to A. maculosum ,
particularly as far as rib morphology is concerned. A.
transcendens can be separated from maculosum by a
more vividly coloured, thinner and often smaller shell,
less elongated, more globular and with more ribs (range
56-74 as opposed to 43-61 in maculosum) . A.
transcendens is generally rather consistent as far as size,
colours and thickness of the shell are concerned.
According to Vidal (1999) junior synonyms are :
Cardium perstriatum Kuroda 1928: II (Fide Habe,
1981: 111). Laevicardium pulcherrimum Sakurai &
Habe, 1966: 293. HABITAT. "1909 Melvill: 30 fathoms,
among quantities of Halimeda- and Lithothamniarubble. "1999 Vidal: recorded from 9 to 295 m depth,
mostly from 30-80 m. "1999 Higo et af.: intertidal to
50 m, sand and mud . DISTRIBUTION. "1909 Melvill:
Amirantes. "1994 Hyl1eberg : Indo-West Pacific .
"1999 Vidal : it is probably present in the main part of
the tropical Indian Ocean, and in many central Pacific
archipelagoes. "1981 Kuroda & Habe; "1999 Higo et
al.: Kii peninsula (and southwards, East China Sea,
Philippines, South China Sea, tropical Indo-Pacific,
Australia.
transclathratum Viader, 1951. - Parvicardium
transclathratum Viader, 1951. Cardiidae. Recent.
RECORDS . "1951 Viader, p . 143 , pI. III, fig. 7:
Parvicardium transclathratum . "1951 ZR: Cardium
transclathratum Viader, 1951 sp. nov. Stray valves
abundant in shell sand between Grand Bay and Cap
Malheureux Mauritius . "1988 Drivas & Jay :
Parvicardium transclathratum (Viader, 1951). "1977
Fischer-Piette: Cardium (Cerastoderma, Parvicardium)
transclathratum Viader, 1951. - CHARACTERS AND
OCCURRENCE - "1988 Drivas & Jay: tuberculated
ribs on the shell surface. DIMENSIONS. "1988 Drivas &
Jay: Max. ant.-post. 6.4 mm, umbo-ventr. 6.2 mm.
DISTRIBUTION. "1988 Drivas & Jay: Reunion, western
Indian Ocean at 40 m depth. "Of authors: Western
Indian Ocean, South Africa.
transilis Eberzin, 1960. - Pachydacna transilis Eberzin,
1960. Lymnocardiidae. Pliocene. RECORDS. "1960 ZR:
Pachydacna transilis Eberzin, 1960 sp. n . Pliocene
Duab river, Abkhaziya, Georgia, USSR.
transkaspica Andrussov, 1924 . - Monodacna
transkaspica Andrussov, 1924 . Lymnocardiidae.
Pliocene and Pleistocene. RECORDS. "1924 ZR:
Monodacna transkaspica Andrussov, 1924 sp. n.
Pleistocene. "1963 ZR: Monoda cna transkaspica
Andrusov, 1963 sp. nov. Pliocene, Russia.
translalum Prashad, 1932. [Figs. Part 3 p. 870]. Changed combination; new name : Ctenocardia
tran slatum (Prashad, 1932) . Cardiidae . Recent.
RECORDS. "1932 Prashad: Cardium (Ctenocardia)
translatum (n. nov.). "1932 ZR : Cardium
(Ctenocardia) translatum Prashad, 1932 nom. n. for
Cardium rigidum Rve . non Wood. "1977 FischerPiette: Corculum (Ctenocardium) translatum . "1977
Wilson & Stevenson: Cardium translatum Prashad,
1932. "1977 Wilson & Stevenson : Cardium
(Ctenocardia) translatum Prashad, 1932. "200 I Vidal,
p. 408: Ctenocardia translatum. ··2003 Hylleberg &
Kilburn, p. 178: Ctenocardia translatum (Prashad,
1932). - CHARACTERS AND OCCURRENCE -
DESCRIPTIONS. "2001 Vidal : only the exhalant aperture
is tube-shaped. Inhalant siphon open towards the foot
aperture. No eyes. DISTRIBUTION. "1994 Hylleberg:
Central Indo-West Pacific. ··2003 HylJeberg & Kilburn:
I valve from Quinhon (Vietnam) is deposited in the
Collection Saurin at MNHN, Paris. (1. Vidal).
translucidum. - Combination to be revised: "Cardium "
translucidum. Cardiidae . Cretaceous of California.
RECORDS. "1871 Stoliczka, 149: C. translucidum.
transpositum Deshayes, 1853 . - Combination to be
revised: "Cardium" transpos itum Deshayes.
Cardiacea. Fossil. RECORDS. "1980 Ruhoff: Cardium
transpositum Deshayes, 1853 ; vol. 2: 49, 53
transversa (subspecies) . - See under pomeyro/i
transversa Freneix, 1957.
transversa Fan, 1963. - Protocardia transversa Fan,
1963 . Cardiidae . Lias. RE CORD S. "1963 ZR :
Protocardia transversa Fan, 1963 sp. nov. Lechang,
Guangdong, China, Lias.
transversa Sowerby. - Papyridea transversa Sow. Cardiidae. RECORDS. "1890 Paetel: Papyridea transversa
Sow.
*transversale. - Homonymy to be noticed. Cardium
transversale from 2 sources: Romer, 1849. - Deshayes,
1854.
transversale Deshayes, 1854. [Figs. Part 3 p. 917]. Synonym of Parvicardium exiguum (Gmelin, 1791).
» Regarding Cardium transversale: Cardiidae.
Recent. REMARKS. RECORDS. "1854 Deshayes: Cardium
transversale . "1885 Smith: Cardium (Papyridea)
transversale, Deshayes. 1854. "1888 Kobelt: Cardium
transversale Deshayes. "1890 Paetel : Cardium
transversale Dsh. "1917 Hidalgo: Cardium
transversale Deshayes. "1927 Dautzenberg: Cardium
(Parvicardium) transversale Deshayes. "1962 Clarke:
Parvicardium transversale (Deshayes) 1854. "1969
Nordsieck: Papyridea transversale (Deshayes, 1854).
"1977 Fischer-Piette: Cardium transversale Deshayes
= Cardium exiguum Gmelin. "1980 Ruhoff: Cardium
transversale Deshayes, 1854; p. 333 . "1984 ZR:
Cardium transversale Deshayes, 1854. Preoccupied
name, replaced by Parvicardium vroomi. Ref. to van
Aart sen et al. p. 67 . ··2000 Aartsen & Goud:
Parvicardium transversale Deshayes. Type in BMNH.
- CHARACTERS AND OCCURRENCE - REMARKS.
"1962 Myra Keen in Clarke, p. 66: states that Cardium
transversale Deshayes is preoccupied by Cardium
transversale Romer 1849 (fide Fischer-Piette 1977).
However, a new name is not needed because C.
transversale Deshayes = C. exiguum Gmelin (FischerPiette 1977). HABITAT. "1969 Nordsieck: 126-1846 m
depth. DISTRIBUTION. ··1969 Nordsieck: Mediterranean,
Canary Islands, Algeria.
transversale Romer, 1849. - Nomina ob/ita . Cardiidae.
Cretaceous. TYPE LOCALITY. Friedrichsburg, Germany.
RECORDS. '·1849 Romer: Cardium transversale. "192232 Sherborn, p. 6560: Cardium transversale C.F.
Roemer, 1849. "1977 Fischer-Piette: Cardium
transversale Romer. "1986 Scott: Cardium
transversale Roemer (1849) considered nomina oblita.
transversum Rehder & Abbott, 1951 . - Changed
combination: Nemocardium transversum (Rehder &
Abbott, 1951). Cardiidae. Recent. RECORDS. "1951
Rehder & Abbott, p. 56, pI. 9, figs. 9-10: Microcardium
Phuket Marine Biological Center Special Publication vol. 30 (2004)
807
transversum - Tridachna
transversum. "1951 ZR: Mieroeardium transversum
Rehder & Abbott, 1951 sp. nov. 50 m. SSw. of Marsh
Island, Iberia Co. Louisiana 29 fms. "1964 Parker:
Mieroeardium transversum Rehder & Abbott. "1974
Abbott: Nemoeardium transversum Rehder & Abbott,
1951. "1977 Fischer-Piette: Mieroeardium
transversum Rehder & Abbott. "1979 ZR:
Mieroeardium transversum. Ref. to Treece. "1997
Shelton: Nemoeardium transversum Rehder & Abbott,
1951. - CHARACTERS AND OCCURRENCE - OF
COLLECTIONS. "ZMUC: I valve, west of Mississippi
delta, Gu1fofMexico. HABITAT. "ZMUC: clayey sand
at 60 m depth. DISTRIBUTION. "Of authors: Louisiana;
Mississippi Delta, Gulf of Mexico.
transversum Sowerby. - Insufficiently known. RECORDS.
"1840 Swainson: Papyridea transversum Sow. Conch.
f. 4 asterisk. "1858 Adams & Adams, p. 456: Cardium
(Papyridea) transversa Sow. "1922-32 Sherborn, p.
6562: Cardium transversum Sowerby, 1832 [no etf].
*trapezinum. - Homonymy to be noticed. Cardium
trapezinum from 3 sources : Andrusov, 1923.
Andrussov, 1924. Andrusov, 1963. REMARKS. Andrusov,
1923 and Andrussov 1924 may be identical with
Andrusov 1963 (see below).
trapezinum Andrusov, 1923. Changed combination:
Caspieardium trapezinum (Andrusov, 1923).
Lymnocardiidae. Pliocene. RECORDS. ··1958 ZR:
Cardium trapezinum Andrusov, 1923. Genotype of
Caspieardium Astafieva, 1958. Cardiidae. Pliocene.
Azerbaijan and Turkmenia, USSR ··1958 ZR:
Caspieardium trapezinum (Andrusov, 1923) [Ref.
Astafieva 1958].
trapezium Andrussov, 1924. - Combination to be
revised: "Cardium" trapezium Andrussov, 1924.
?Lymnocardiidae. Pleistocene. RECORDS. "1924 ZR:
Cardium trapezium Andrussov, 1924 sp. n.
trapezinum Andrusov, 1963. - Combination to be
revised: "Cardium" trapezium Andrusov, 1963.
?Lymnocardiidae. Pliocene. RECORDS. "1963 ZR:
Cardium trapezinum Andrusov, 1963 sp. nov. Apsheron
layers, Pliocene, Russia.
trapezium Cecalupo & Quadri, 1996. - Parvieardium
trapezium Cecalupo & Quadri, 1996, Cardiidae. Fossil.
RECORDS. "2000 Aartsen & Goud: Parvieardium
trapezium Cecalupo & Quadri, 1996.
trapezoida/e Orbigny, 1850. - Conocardium trapezoidale
Orbigny, 1850. Conocardiidae. Fossil. RECORDS.
"1922-32 Sherborn, p. 6566: Conocardium (Roemer)
trapezoidale d'Orbigny, 1850. - Pleurorhynchus
trapezoidalis. "1922-32 Sherborn, p. 6566:
Pleurorhynehus trapezoidalis FA. Roemer, 1843.
traubi Schlickum, 1970. - Cerastoderma traubi
Schlickum, 1970. Cardiidae. Upper Bavaria. RECORDS.
"1970 ZR: Cerastoderma traubi Schlickum, 1970 sp.
nov. Borehole SW. 13 Harmoning I, Teufe, Upper
Bavaria, West Germany. "1977 Popov: Cerastoderma
traubi Schlickum, 1970.
triangu/a G. Schneid. - Changed combination:
Pontalmyra triangula (G . Schneid.). Lymnocardiidae.
Fossil. RECORDS. "1987 Taktakishvili: Pontalmyra
triangula (G. Schneid.).
*triangulare. - Homonymy to be noticed. Cardium
triangulare from 2 sources: Young & Bird, 1828. Coquand, 1862.
trumgu!are Coquand, 1862, - Combination to be revised:
"Cardium" triangulare Coquand, 1862. Cardiidae.
Fossil. RECORDS. "1871 Stoliczka, 109: C triangulare
Coquand, 1862. "1980 Ruhoff: Cardium triangulare
Coquand, 1862; vol. 2: 206, pI. II, figs. 1,2.
triangu/are Young & Bird, 1828. - Combination to be
revised: "Cardium" triangulare Young & Bird, 1828.
?Cardiidae. Fossil. RECORDS. "1922-32 Sherborn, p.
6572: Cardium triangulare Young & Bird, 1828.
triangularis Martyn, 1784. Non binominaL - Not
available. Not a cardiid. RECORDS. "1784 Martyn:
Cardium triangularis.
triangularis Noda, 1962. - Serripes triangularis Noda,
1962. Cardiidae. Upper Miocene. RECORDS ... 1962 ZR:
Serripes triangularis Noda, 1962 sp. nov. Itanoki-sawa,
Araki-mura, Mogami-gun, Yamagata Pref. Mitsumori
formation, Miocene, Japan. "1973 Keen: Serripes
triangularis Noda, 1962.
triangulata Campbell & Engel, 1963. - Chaenoeardiola
triangulata Campbell & Engel, 1963 . Cardiacea.
Tournaisian. RECORDS. "1963 ZR : Chaenocardiola
triangulata Campbell & Engel, 1963 sp. nov. New
South Wales, Toumaisian.
triangulatocostatum (Hal.) . - New combination:
Lymnoeardium
triangulatoeostatum
(Hal.)
Lymnocardiidae. Fossil. RECORDS. "1987 Taktakishvili>
Limnoeardium triangulatoeostatum (Hal.).
triangulatum Laubriere, 1881. - Changed combination:
Parvieardium triangulatum (Laubriere, 1881).
Cardiidae. Eocene. RECORDS. "1886 Cossmann:
Cardium triangulatum Laub. "1904-06 Cossmann &
Pissarro, 69-20: Cardium (Plagiocardium)
triangulatum de Laub. "1970 Glibert & Poel, p. 48:
Parvicardium triangulatum (de Laubriere, 1881).
"1995 Le Renard et al.: Parvicardium triangulatum
[Cardium triangulatum] De Laubriere, 1881. "1995
Pacaud & Renard: Parvicardium triangulatum. "1998
Schneider: Parvicardium (Parvicardium) triangulatum
(Laubriere, 1881). "1998 Schneider: Parvicardium
triangulatum (Laubriere, 1881) Cardiidae. Eocene.
"1999 Jacques Vidal (pers . com.) MNHN, Paris:
Fossils from the Paleontological Collection as
Plagioeardium triangulatum de Laubriere. Lutetien.
France (Bassin de Paris), B52735 . -6.
triangulatum Sowerby. - Nomen nudum. Not available.
RECORDS. "1857 Carpenter, p. 94: Cardium
triangulatum Sow. (nomen nudum). "1872 Carpenter,
p. 30: Cardium triangulatum Sow. "190 I Dall :
Cardium triangulatum "Sowerby" Carpenter, 1857
(perhaps a MS name). "1977 Fischer-Piette, p. 123:
Cardium triangulatum Sowerby, Orbigny, 1847. Syn.
of Coreulum (Trigoniocardia) biangulatum Sowerby,
1829. "1977 Fischer-Piette, p. 155: Cardium
triangulatum Sowerby. Insufficiently known species.
triangulum Muenster in Goldfuss, 1837. - See under
triangulum Orbigny.
triangulum Orbigny, 1850. - Conoeardium triangulum
d'Orbigny, 1850. Conocardiidae. Fossil. RECORDS .
.. 1922-32 Sherborn, p. 6575: Cardium triangulum
Muenster in Goldfuss, 1837. "1922-32 Sherborn, p.
6575: Conocardium (Muenster) triangulum d'Orbigny,
1850. - Cardium.
Tridachna (error) . - Printer's mistake for Tridacna in
ZR for 1937 with ref to Iredale 1937.
808
Tropical Marine Mollusc Programme (TMMP)
Tridachnes - Trifaricardium
Tridachnes (genus). - Synonym of Trida cna Bruguiere,
1797. » Regarding Tridachnes: Tridacnidae. Recent.
RECORDS. --1937 ZR: Trida chnes Bolten [R6ding]
Logo-type here selected Chama gigas L. [Ref. Iredale
1937]. "1998 Schneider (l998b): Tridachnes maxima
R6ding, 1798. --1999 Higo et at.: Trida chnes R6ding,
1798 = Tridacna Bruguiere, 1797. Tridacnidae.
Tridacna. - (genus). Tridacnidae. RECORDS. --1884 ZR:
Tridacna [Ref. Sowerby 1884] ... 1891 ZR: Tridacna,
anatomy of [Ref. Menegaux 1891] ... 1899 ZR:
Tridacna its anatomy. [Ref. Grobben 1899] . --1904 ZR:
Tridacna morphology and anatomy [Ref. Anthony
1904] ... 1920 ZR: Tridacna [Ref. Anthony 1920].
.. 1927 ZR: Tridacna fossil [Ref. Cox 1927]. "1931
ZR: Tridacna (digestion) Ref. to Yonge (1931). --1932
ZR: Tridacna, crystalline style, [Ref. Yonge 1932b].
.. 1934 ZR : Tridacna fig. [Ref. Boutan 1934]. .. 1937
ZR : Trida chna [err. pro Trida cna] Humph . Mus.
CaloJU1. "Logotype here selected" Chama gigas L. [Ref.
Iredale 1937] ... 1938 ZR: Tridacna Biology [Ref.
Tomlin 1936]. --1939 ZR: Tridacna Pearls [Ref. Cobb
1939]. --1946 ZR: Tridacna communication between
mantle and stomach [Ref. Mansour 1946]. --1947 ZR:
Tridacna anatomy [Ref. Mansour 1947] ... 1957 ZR:
Trida cna nature and occurrence of pearls [Ref. Haas
1957]. --1962 ZR: Tridacna records oflarge specimens
[Ref. Roscoe 1962] ... 1963 ZR: Tridacna collecting
on Eniwetok [Ref. Roberts 1963]. --1969 ZR: Tridacna
spp. ecology in Palau, Western Caroline Is. [Ref. Hardy
& Hardy 1969]. --1970 ZR: Tridacna Biology, ecology.
[Ref. Cowen, R.] . .. 1971 ZR: Tridacna [Ref. Salvat
1971]. --1971 ZR: Tridacna Biology and ecology [Ref.
Fankboner 1971]. "1975 ZR: Trida cna . Steroid
hormones . Ref. to Teshima. / Distribution. Ref. to
Yonge. --1976 ZR: Tridacna. Heavy metal content.
Strontium partition coefficient between sea water &
aragonite. Western Pacific. Ref. to Konishi. / Toxin
toxicity. Ref. to Kanno. --1977 ZR: Tridacna. Care in
captivity. Popular account. Ref. to Weingarten &
Tuxson. / Coral Sea. Great Barrier Reef. Impact of
poaching on population dynamics. Fishing & fisheries .
Ref. to Pearson. / Holocene. Tasman Sea, Gilbert &
Ellice Islands, stratigraphy. Ref. to Schofield . .. 1981
ZR: Trida cna. Ref. to Benson et al. Great Barrier Reef.
/ Fankboner. / Trench et al. --1982 ZR: Tridacna . Ref.
to de Couet. / Harada el al. / Romano. / Ueda el al. /
Yonge. --1983 ZR: Tridacna. Ref. to Benson / Bruce /
Munro et at. / Poulicek / Wells. --1984 ZR Tridacna :
Eye. Retina cell hyperpolarization . Ref. to Wilkens.
.. 1992 Schneider p. 155: Tridacna. Type species
Tridacna gigas (Linnaeus, 1758). --1998 Schneider:
Trida cna with three subgenera: (Tridacna),
(Cham etra chea ), (Persikima) . .. 1999 Higo el at.:
Tridacna (subgenus) s. Sir. Tridacnidae ... 1999 Higo
el at.: Trida cna Bruguiere, 1797. Type species: Chama
gigas Linnaeus, 1758 (s. m. Lamarck, 1799).
Tridacnacea (superfamily). - RECORDS ... 1974 ZR:
Tridacnacea. Population censuses. Coral reef. West
Caroline Basin. Ref. to Hester. --1976 ZR: Tridacnacea.
Crustacean parasites. Cyclopoida. Parasite systematics.
Ref. to Humes. --1977 ZR: Tridacnacea. Mutualism.
Symbiosis, algae, lectin system defensi ve mechanism
or digestive function . Ref. to Uhlenbruck &
Steinhausen. --1979 ZR: Tridacnacea. General account.
Coral reefs, Australia. Ref. to Summerhays. --1980 ZR:
Tridacnacea, western Pacific. Ref. to Bishop, D.G. et
al. --1983 ZR: Tridacnacea. Ref. to Mangum:
comparative review. --1992 Schneider: superfamily
Tridacnacea of Keen (fide Schneider 1992) ... 1998
Schneider: superfamily Tridacnacea, Family
Tridacnidae.
Tridacnidae (family). - REcORDs. "1865 ZR: Tridacnidae [Ref. Vaillant 1865] . "1940 ZR: Tridacnidace
[err. pro Tridacnidae] in Mus. hist. nat. Paris [Ref. Lamy
1940]. --1947 ZR: Tridacnidae food and feeding habits
[Ref. Mansour 1947]. --1957 ZR: Tridacnidae pallial
attachment, nervous system and evolution with regard
to the monomyarian condition [Ref. Yonge 1957].
--1964 ZR: Tridacnidae, species of Dar es Salaam [Ref.
Spry 1964]. --1966 ZR : Tridacnidae ofthe Great Barrier
Reef, Australia, [Ref. Rosewater 1966] ... 1953 ZR :
Tridacnidae, mantle and water circulation, figs. [Ref.
Yonge 1953]. "1970 ZR : Tridacnidae sic symbiotic
zooxanthellae in mantle edge [Ref. Read 1970]. --1974
ZR: Tridacnidae . Particulate feeding. Gastric
diverticula. Functional morphology. Posture. Ref. to
Fankboner. "1975 ZR: Tridacnidae. Systematic
accounts of species. Ref. to Lambiotte. --1976 ZR:
Tridacnidae. Community structure. Population size.
West Caroline Basin. Ref. to Bryan . --1979 ZR:
Tridacnidae. Ref. to Heslinga. --1981 ZR: Tridacnidae.
Ref. to Gwyther et al. / Heslinga. / Yonge. Indo Pacific.
--1999 Higo el al.: Tridacnidae (family) Lamarck, 1819
(Refs.: Rosewater, 1965). --1983 ZR : Tridacnidae. Ref.
to D ' Attilio et al. - CHARACTERS AND OCCURRENCE - REMA RKS. --1998b Schneider: all species of
tridacnines harbor photosymbiotic dinoflagellate algae
(zooxanthellae) in their tissues. Extinct tridacnine taxa
are known from the Middle Eocene. Extant tridacnine
taxa are known from Miocene fossils .
Tridacninae (subfamily). - RECORDS. --1998 Schneider
(1998b): Tridacninae (subfamily). --2002 Schneider:
Tridacninae originate in the Pa laeogene , not
Cretaceous. --200 I Schneider & Carter: Tridacninae
(subfamily) Lamarck, 1819.
Tridacnodites KrUger, 1823. - Synonym of Trida cna
Bruguiere, 1797 . » Regarding Tridacnodiles:
Tridacnidae. Recent. RECORDS. --1999 Higo el al. :
Tridacnodites KrUger, 1823 = Tridacna Bruguiere,
1797. Tridacnidae.
Tridacnoidea (Superfamily). - RECORDS. --1995 Pacaud
& Renard : Superfamily Tridacnoidea.
Trifaricardium Kuroda & Habe, 1951. - ?Valid genus .
No description (Vokes 1980). GENOTY PE. Habe (1951)
designated Cardium nomurai Kuroda & Habe, 195 as
type species of subgenus Trifaricardium of the genus
Nemocardium Meek, 1876. RECORDS. --1961 ZR :
Trifaricardium Kuroda & Habe, 1961 gen . nov.
Cardiidae genotype Cardium (Acanthoca rdia)
cancel/alum Nomura 1933 = nomurai p. 34, nom . nov.
pro cancellalum Nomura 1933 non Gmelin 1791, Japan
[published after preparation of the manuscript of this
work in Illust. Cat. Jap. Shells No. 13 p. 86, 1951
Kuroda & Habe]. --1962 ZR: Trifaricardium gen . nov.
nomurai nom. nov. for Cardium cancel/ata Nomura
1933, non Gmelin 1791. --1980 Vokes : Trifaricardium
Kuroda & Habe, 1951 . --1992 Poutiers: Trifaricardium.
--1999 Higo el al.: Trifaricardium Kuroda & Habe in
Phuket Marine Biological Center Special Publication vol. 30 (2004)
809
trijidum - trigonoides
Babe, 1951. Type species : Trifarlcardium nomurai
Kuroda & Habe, 1951 (0. d.). -CHARACTERS AND
OCCURRENCE - REMARKS. --1994 Hylleberg: near
Nemocardium (Microcardium) but secondary
concentric sculpture on anterior slope only.
trifldum Deshayes, 1858 . - Changed combination:
Orthocardium trifidum (Deshayes, 1858). Cardiidae.
Eocene. RECORDS. "1886 Cossmann: Cardium trljuJum
Deshayes. --1904-06 Cossmann & Pissarro, 69-6:
Cardium (Trachycardium) trifidum Desh . "1930
Stewart, p, 254: "Cardium" trifidum. Ref to Cossmann
& Pissarro 1906, pI. 17, fig. 69-6. (Thanetien) "1970
Glibert & Poel, p. 53: Cardium (Orthocardium) trifidum
Deshayes, 1858. "1977 Popov: Orthocardium trifidum
(Deshayes, 1858). "1980 Ruhoff: Cardium trifidum
Deshayes, 1858; [1856-1865] vol. I: 556, pI. 56, figs.
5-7. "1995 Le Renard et al.: Cardium (Orthocardium)
trifidum Deshayes, 1858. "1995 Pacaud & Renard:
Cardium (Orthocardium) trifidum . "1999 Vidal (pers.
com.) MNHN, Paris: Fossils from the Paleontological
Collection. Trachycardium trifidum Deshayes.
Thanetien. France (Bassin de Paris), 52668 to -71 ,
B69974.
trijkovici (subspecies). - See under conjugens trifkovici
Brusina 1902.
trijkovici (cf.). - New combination: Lymnocardium cf.
trifkovici. Lymnocardiidae. Pliocene. RECORDS. --1961
ZR: Limnocardium sp. nov. Gillet & Vicente, 1961 cf.
trifkovici Pliocene, Barcelona, Spain.
triforme Sowerby, 1840. - Changed combination:
Discors triforme (Sowerby, 1840). Cardiidae. Fossil.
RECORDS. "1922-32 Sherborn, p . 6602: Cardium
triforme Sowerby, 1840. --1977 Popov: Discors
triforme (Sowerby, 1840).
trigona Fan, 1963. - Protocardia trigona Fan, 1963.
Cardiidae. Lias. RECORDS. --1963 ZR: Protocardia
trigona Fan, 1963 sp. nov. Xaioshui , Lechang,
Guangdong, China, Lias.
trigona. - Not in Cardiidae. » Regarding Cardium
trigona: RECORDS. "1839 Anton, no. 405 . Cardium
trigona = Donax cardioides Lamarck.
trigonale d'Orbigny, 1850 . - Conocardium (Ph .)
trigonale d'Orbigny, 1850. Conocardiidae. Fossil.
RECORDS. --1922-32 Sherborn: Pleurorhynchus
trigonale Ph . --1922-32 Sherborn, p. 6603 :
Conocardium (Ph.) trigonale d' Orbigny, 1850. Pleurorhynchus.
trigonalis. - Cardium trigonalis. ?Cardiacea. Fossil.
RECORDS. --1857 Deshayes: Cardiumtrigonalis
(England).
trigonellare Buvignier, 1852. - Combination to be
revised: "Cardium" trigonellare Buvignier, 1852.
Cardiidae. Fossil. RECORDS. --1980 Ruhoff: Cardium
trigonellare Buvignier, 1852; atlas p. 15, pI. 13, figs.
32-33.
*trigonellum. - Homonymy to be noticed. Cardium
trigonellum from 2 sources: Orbigny, 1852. - Bronn,
1848.
trigonellum Bronn, 1848. - Synonym of Parvicardium
papillosum (Poli, 1791).» Regarding C. trigonellum :
Cardiidae. Fossil. RECORDS. --1867 Weinkauff: Cardium
trigonellum Bronn, 1848 = Cardium papillosum Poli.
trigonellum Orbigny, 1852. - Synonym of Parvicardium
papillosum (Poli, 1791). » Regarding Cardium
trigonellum: Cardiidae. Fossil. RECORDS. --1888 Kobelt:
Cardium trigonellum d ' Orb. (fide Weinkauf£) =
Cardium papil/osum Poli o --1980 Ruhoff: Cardium
trigonellum Orbigny, 1852; [1850-1852] vol. 3: 118.
Trigoniocardia Dall, 1900 (genus). - Cardiidae. Recent.
GENOTYPE. Dall (1900) designated Cardium granifera
Broderip & Sowerby, 1829 type species of genus
Trigoniocardia. RECORDS. --1953 Olsson & Harbison:
Genus Trigoniocardia Dall, 1900. --1980 Vokes :
Trigoniocardia Dall, 1900 in subfamily Fraginae.
.. 1934 Johnson: Trigoniocardia Dall 1900. "1961
Olsson: Trigoniocardia Dall, 1900. --1977 Popov:
Trigoniocardia Dall, 1900. Genotype: Cardium
graniferum Broderip & Sowerby, 1829. "1977 ZR:
Trigonocardia [err. pro Trigoniocardia] Key to
subgenera. Ref. to Popov p. 55. --1992 Schneider p.
155: Trigoniocardia. Type species Trigoniocardia
granifera (Brode rip & Sowerby) . "1998 Schneider:
Trigoniocardia. A number of Eocene species have been
variously classified as Trigoniocardia. "1999 Higo et
at.: Trigoniocardia Dall, 1900. Type species: Cardium
graniferum Broderip & Sowerby, 1829 (0. d.). "2001
Vidal, p. 408 : Trigoniocardia . - CHARACTERS AND
OCCURRENCE- DESCRlPTION. --1994 Hylleberg: shell
ovate to quadrate; hinge asymmetric, anterior section
shorter than posterior. .. 2001 Vidal: only the exhalant
aperture is tube-shaped . Inhalant siphon open towards
the foot aperture. No eyes.
Trigoniocardia Dall, 1900 (section and subgenus). Cardiidae. Recent. RECORDS. --1900 ZR: Trigoniocardia
Dall, 1900a n. sect. of Fragum type Cardium
graniferum Sby. --1922 Olsson: section Trigoniocardia
Dall. --1924 Oldroyd: Subgenus (Trigoniocardia) Dall,
1900.
Trigonocardia (genus) . - Printer 's error. RECORDS.
--1980 Vokes: Trigonocardia (err. pro Trigonio- Dall,
1900) Cossmann, 1901.
trigonoides Eichwald, 1838. [Figs. Part I p. 123] . Identical with Didacna trigonoides Pallas, 1771.
Lymnocardiidae. Pliocene and Recent. RECORDS. "1838
Eichwald: Didacna trigonoides n. "1841 Eichwald:
Didacna trigonoides . --1844 Reeve : Didacna
trigonoides. Eichwald = Cardium trigonoides Pallas.
--1876 ZR: Cardium [Didacna] crassum (Eichw.) and
trigonoides (Pall.), in the Caspian Sea, connected by
intermediate forms [Ref. Grimm 1876]. "1922-32.
Sherborn, p. 6606 : Didacna (Pall.) trigonoides
Eichwald, 1838. - Cardium trigonoides Pallas, 1771.
--1999 Jacques Vidal (pers. com .) MNHN, Paris:
Fossils from the Paleontological Collection. Didacna
trigonoides Eichwald. Caspian Sea, B37642.
trigonoides Pallas, 1771. [Figs. Part I p. 128]. - Didacna
trigonoides (Pallas, 1771). Lymnocardiidae. Pliocene
and Recent. Type species: Stoliczka (1870) designated
Cardium trigonoides Pallas, 1771 type species of the
genus Didacna Eichwald, 1838. TYPE LOCALITY . Caspian
Sea. RECORDS. --1771 Pallas: Cardium trigonoides (fide
Eichwald 1841). --1776 Pallas [Anhang p. 26, No. 86]:
Cardium trigonoides . "1788 Schriiter: Cardium
trigonoides [Reference to Chemnitz (1782) fig. 203] .
.. 1841 Eichwald : Cardium trigonoides Pallas, 1771.
.. 1838 Eichwald: Cardium trigonoides Pall. "1844
Reeve species 26 : Cardium trigonoides Pallas, 1776.
'Reise durch verschied. Provinzen des Russischen
810
Tropical Marine Mollusc Programme (TMMP)
trigonoides ehasariea- trilaterum
Reichs,' vol. i.; Anhang (Appendix) , p. 26, no.
80. "1849 Middendorff: Cardium trigonoides (fide
Andrussoff 1903). 001858 Adams & Adams, p. 460:
Adaena (Didacna) trigonoides Pallas. 001866 Issei:
Didacna trigonoides Pallas. "1870 Stoliczka: Cardium
trigonoides Pallas, 1771. '·1877 Grimm: Cardium
trigonoides Pallas. "1884 Grasset: Adacna trigonoides.
Pal1. "1887 Fischer: Didacna trigonoides Pallas (in s.
g. Didacna Eichwald, 1838). 001890 Paetel: Adacna
trigonoides Pallas. ··1890 Woodward : Didacna
trigonoides Pal. - Caspian. 001902 Sherborn, p. 993:
Cardium trigonoides Pallas, 1771. "1903 Andrussoff:
Cardium trigonoides Pallas. 001903 Andrussoff:
Cardium trigonoides. 001916 ZR: Didacna trigonoides
Ref. to KaJickij (1916). "1930 ZR: Didacna
trigonoides [Ref. Pravoslavleff 1913 sic]. 001947 ZR:
Didacna trigonoides effects of salinity in the Caspian
Sea, [Ref. Karparich 1947]. 001963 ZR: Didacna
trigonoides, food and feeding mechanism [Ref.
Rosssanova 1963]. "1964 Shikama: Didacna
(Didacna) trigonoides (Pallas). 001968 Logvinenko &
Starobogatov: Didacna trigonoides trigonoides. 001969
Moore et al.: Didacna (Didacna) trigonoides (Pallas).
001977 Popov : Didacna trigonoides (Pal1.). 001980 ZR:
Didacna trigonoides. Caspian Sea. Ref. to Galperina.
001984 ZR: Didacna trigonoides. Ref. to Galperina.
001998 Schneider (1998b): Didacna trigonoides (Pallas,
1771). - CHARACTERS AND OCCURRENCEDESCRIPTIONS . . . 1776 Pallas, original: Testa
magnitudine fere rustici, subimbricata, valde gibba,
inaequilatera, subtriangularis. Valvulae aequales,
vesus nates inflexas angulatae, altero latere
planiusculae, altero lato, convexo. Striae in planiore
latere exiles circiter sex, area vulvae laevigata, in
convexa parte 14-18. latae, complanatae. Copiosissima
testa Maris Caspii, viva tamen mihi non visa. 001844
Reeve: shell triangularly cordate, elongated and slightly
angulated posteriorly with the umbones obtusely
keeled, shortened and compressed anteriorly; very
smooth, whitish, ornamented with slightly elevated
brown rays, very like worn flattened ribs, diverging
with great regularity from the umbones. REMARKS.
001844 Reeve: The Figure, Part I, p. 128, No. 26b, has
an interesting history. Reeve described the fate of the
shell as being one of three shells described by Pallas in
'Journey through different Provinces of the Russian
Empire,' published at Leipzic in 1776. Although a
species of some rarity in our collections, it must be
tolerably abundant in the Caspian Sea according to the
testimony of both Pallas and Eichwald. The locality
given by Lamarck, "Seas of America," is extremely
vague, and most probably incorrect, as it appears to be
founded merely upon the fact of his specimen, from
the cabinet of M. Defrance, having been presented to
that naturalist by M. Richard after his return from
Guyana. It will perhaps be interesting to know that the
shell before me, belonging to Miss Saul, is from the
same source; the chief portion of M. Richard's
conchological rarities fell into the possession of Dr.
Goodhall of Eton, and, at the death of the venerable
Provost, the specimen in question passed into the
collection of the Rev. Mr. Stain forth, to whom its
present possessor is indebted for it. 001903: Andrussoff
stated that Eichwald, 1838 moved Cardium trigonoides
to genus Didacna. DISTRIBUTION. 001844 Reeve, 001884
Grasset: Caspian Sea.
trigonoides ehasariea Svitoc, 1967. - Didacna
trigonoides chasarica Svitoc, 1967. Lymnocardiidae.
Quaternary. RECORDS. 001967 ZR: Didacna trigonoides
chasarica Svitoc, 1967 subsp. nov. figs.; Quaternary,
central Caspian region.
trigonoides ehvalyniea Svitoc, 1967. - Didacna
trigonoides chvalynica Svitoc, 1967. Lymnocardiidae.
Quaternary. RECORDS. 001967 ZR: Didacna trigonoides
chvalynica Svitoc, 1967 subsp. nov. figs.; Quaternary,
central Caspian region.
trigonoides erassum Grimm, 1876. - Synonym of
Didacna baeri (Grimm, 1877). » Regarding Cardium
trigonoides crassum: Lymnocardiidae. TYPE LOCALITY.
Caspian Sea. RECORDS. 001876 Grimm, p. 140: Cardium
trigonoides crassum m. 001877 Grimm: Cardium
trigonoides crassum m. (syn.) = Cardium baeri sp. n.
trigonoides praetrigonoides Nat. et Anis. - Didacna
trigonoides praetrigonoides Nat. et Anis. Lymnocardiidae. TYPE LOCALITY. Caspian Sea. RECORDS. 001968
Logvinenko & Starobogatov: Didacna trigonoides
praetrigonoides Nat. et Anis.
trigonoides tuzetae Tadjalli-Pour, 1977. - Didacna
trigonoides tuzetae Tadjalli-Pour, 1977. Lymnocardiidae. RECORDS. 001977 ZR: Didacna /rigonoides
/uze/ae Tadjalli-Pour, 1977 subsp. nov. Caspian Sea.
Ref to Tadjalli-Pour, M . p. 97, fig'd.
trigonoides elongata Pravoslavleff, 1913 (variety). Upgraded. Didacna trigonoides elongata Pravoslavleff,
1913. Lymnocardiidae. Fossil. RECORDS. 001930 ZR:
Didacna trigonoides var. elongata Pravoslavleff, 1913
sic nov.
*trigonum. - Homonymy to be noticed. Cardium
trigonum from 3 sources: Mi.inster in Goldfuss, 1837.
- Sismonda, 1842. - Michelotti, 1847.
trigonum Michelotti, 1847. - Replaced by Cardium
michelottii Deshayes. » Regarding Cardium trigonum:
Cardiidae. Fossil. RECORDS. 001857. Deshayes: Cardium
trigonum Michelotti, 1847. - C. michelottii. Desh.
trigonum Mi.inster in Goldfuss, 1837. - Cardium
trigonum Muenster in Goldfuss, 1837. Cardiidae.
Fossil. RECORDS. 001857. Deshayes: Cardium trigonum
Munster, 1840. 00 1922-32 Sherborn, p. 6607: Cardium
trigonum Muenster in Goldfuss, 1837. [Mi.inster in
Goldfuss 1837 is assumed to be identical with Mi.inster
1840].
trigonum Sismonda, 1842. - Synonym of Parvicardium
papillosum (Poli, 1791). » Regarding Cardium
trigonum: Cardiidae. Fossil. RECORDS. 001888 Kobelt:
Cardium trigonum Sism. (fide Weinkauff) = Cardium
papillosum Poli [Weinkauff 1867, p. 138]. 001922-32
Sherborn, p. 6607: Cardium trigonum Sismonda, 1842.
"1999 Jacques Vidal (pers. com.) MNHN, Paris:
Fossils from the Paleontological Collection. Cardium
trigonum Sismonda. The Crimea, 837640.
trilaterum Gmelin, 1791. - Unknown species. RECORDS.
001791. Gmelin No. 26: Cardium trilaterum. 001815.
Wood: Cardium trilaterum Gmelin (p. 3253) is not
specific. 001817 Dillwyn : Cardium trilaterum Gmelin
should be ignored. 001849 Middendorff: Cardium
trilaterum Gm. (fide Andrussoff \903). "1902
Sherborn, p. 993: Cardium trilaterum J.F. Gmelin,
1790.
Phuket Marine Biological Center Special Publication vol. 30 (2004)
811
trilaterum - truncatum
trilaterum Gmelins, 1774. - Unknown species. RECORDs.
001774 Gmelins, p. 248: Cardium trilaterum. 001849
Middendorff p. 29-42: Cardium trilaterum. 001902
Sherborn, p. 993: Cardium trilaterum S.G. Gmelinsie,
1774.
*trinacria. - Homonymy to be noticed. Cardium
trinaeria from 3 sources: Andrussow 1902. - Andrusov,
1963. - REMARKS. I do not know how Andrusov 1963
compares with Andrussow 1902.
trinacria Andrusov, 1963. - Combination to be changed :
"Cardium" trinaeria Andrusov, 1963. ?Lymnocardiidae. Neogene. REcORDs. 001963 ZR: Cardium trinaeria
Andrusov, 1963 sp. nov. Russia. 001986. Nevesskaya
et al.: Andrusovieardium trinaeria (Andrusov) comb.
nov. 001987 ZR: Andrusovieardium trinaeria (Andrusov) comb. nov. Ref. to Nevesskaya et al. p. 192.
trinacria Andrussow, 1902. - Combination to be
changed: "Cardium " trinaeria (Andrussow, 1902).
?Lymnocardiidae. Fossil. RECORDS. 001905 Dollfus,
commenting Andrussow 1902: Cardium trinaeria n.
sp. tends towards Cerastoderma.
trinitatis Tomlin, 1929. - Laevieardium trinitatis Tomlin,
1929. Cardiidae. Recent. DISTRIBUTION. Trinidad.
REcORDs. 001929 Tomlin : Laevieardium trinitatis n. sp.
001929 ZR: Laevieardium trinitatis Tomlin, 1929 sp.
n. Indies.
*tripartitum. - Homonymy to be noticed. Conoeardium
tripartitum from 2 sources: Muenster in Goldfuss, 1837.
- Branson, 1942.
tripartitum Branson, 1942. - New name. Conoeardium
tripartitum Branson, 1942. Conocardiidae. Fossil.
RECORDS. 001942 ZR: Conoeardium tripartitum
Branson, 1942 nom. nov. for Conoeardium truneatum
Pohll929 non Koninck 1885 [Ref. Branson 1942] .
tripartitum Muenster in Goldfuss, 1837. - Cardium
tripartitum Muenster in Goldfuss, 1837.
?Conocardiidae. Fossil. RECORDS. 001922-32 Sherborn,
p. 6620: Cardium tripartitllm Muenster in Goldfuss,
1837.
tripolitanum Blanckenhorn & Offenheim, 1927. Combination to be revised. "Cardium" tripolitanum
Blanckenhorn & Offenheim, 1927. Cardiidae. Miocene.
RECORDS. 001927 ZR: Cardium tripolitanum
Blanckenhorn & Offenheim, 1927 sp. n. Miocene Syria.
triquetrum Matheron. - Not a cardiid. 001857 Deshayes:
Cardium triquetrum Matheron In genus lsoeardia.
triste Gmelin, 1791. - See under triste Linnaeus, 1758.
triste Linnaeus, 1758. - Dubious セー・」ゥウN@
Cardiidae.
RECORDS . 001758 Linnaeus (lot edition) No. 74:
Cardium triste. 001764 Linnaeus (Mus. Lud . Ulr. Reg.)
No. 46: Cardium triste. 001767 Linnaeus : sp . 74 .
Cardium triste removed in the 12th edition 1767. 00 1789
Bruguiere, dubious species, p. 235. Cardium triste Linn.
pag. 680, num. 74. 001791 Gmelin No. 28: Cardium
triste. 001815 Wood: Cardium triste Gmelin is doubtful
and omitted. 001817 Dillwyn: Cardium triste Gmelin
should be ignored. 001864 Krebs: Cardium triste L. ?
(syn.) = Cardium eitrinum Ch. 001902 Sherborn, p. 997:
Cardium triste Linnaeus, 1758.
tristicu/um Yokoyama, 1924. - Changed combination:
Nemoeardium tristieulum (Yokoyama) . Cardiidae.
Upper Oligocene. RECORDS. 001924 ZR: Cardium
(Laevicardium) tristieulum Yokoyama, 1924 sp. n.
Tertiary Japan. 001954 Keen : Nemocardium
(Aretopratulum) tristieulum (Yokoyama), 1924 [U
Oligocene). 001999 Kafanov et al.: Laevieardium
tristieulum (Yokoyama) (= Cardium esutoruensis).
trombosum Gregorio, 1930. - Dieeroeardium trombosum Gregorio, 1930. Cardiidae. Lias. RECORDS.
001936 ZR: Dieeroeardium trombosum Gregorio, 1930
sp. n. Lias Palenno.
Tropicardium. - Printer's error of genus
Tropidoeardium. RECORDS. 001980 Vokes: (err. pro
Tropido- Romer, 1868) Preston, 1916. Synonym of
Cardium Linnaeus, 1758.
Tropidocardium Romer, 1869 (genus). - Synonym of
Cardium. Cardiidae. RECORDS. 001869 ZR: subgenus
Tropidoeardium, type Cardium costa tum (L.) [Ref.
Romer 1869). 001876 Meek : Tropidoeardium Roemer
(Cardium proper of most authors). Cardium eostatum
Linn. regarded as genotype. 001876 Meek:
Tropidoeardium Roemer, 1868. Synon. - Cardiidae.
001930 Stewart: Tropidocardium Romer in Kiister, 1869
(objective synonym of Cardium Linne). CHARACTERS AND OCCURRENCE - DESCRIPTIONS. 001869 Romer: shells very inflated, nearly
equilateral, thin, with broad, flat, longitudinal ribs, with
thin edges or spines and lamellae. Posterior end broadly
gaping, with strongly serrated edges. Ligament short,
thick, projecting. Edge of closure more or less straight
and long; Cardinal teeth small, two in the right valve,
one in the left valve; lateral teeth lamellar, prolonged,
little developed.
Tropidocardium Romer, 1869 (section and subgenus).
- Synonym of Cardium. Cardiidae. REMARKS. The
designation "section" takes the value of subgenus .
RECORDS. 001869 Romer: Sectio Tropidoeardium
Romer. 001887 Fischer: Tropidoeardium Romer, 1868.
Section of genus Cardium Linne, 1758.
troughtoni Iredale, 1927. - Synonym of Tridacna
maxima (Roding, 1798). » Regarding Tridacna
troughtoni: Tridacnidae. Recent. RECORDS. 001927 ZR:
Tridacna troughtoni Iredale, 1927 sp. n. Vanikoro.
001937. ZR: Tridacna trough toni Tridacnidae type of
Sepidaena [Ref. Iredale 1937]. 001999 Higo et af. :
Tridacna troughtoni Iredale, 1927 = Tridacna
(Flodacna) maxima (Roding, 1798).
truncata Vest. - Myocardia truncata von West.
Adacnidae (= Lyrnnocardiidae). Tertiary TYPE LOCALITY.
Tihany, Hungary. REMARKS. This Hungarian fossil
species bridges the gap between Adaena and Cardium.
It has a hinge like in Cardium and a siphonal sinus like
in Adaena (Andrussoff 1903). RECORDS. 001877 Grimm:
Myocardia truncata von West. 001903 Andrussoff:
Myocardia truncata Vest.
*truncatum; Cardiidae - Homonymy to be noticed.
Cardium truncatum from 2 sources: Sowerby, 1827.Goldfuss, 1840.
truncatum Beyrich, 1845. - Protoeardia truneatum
Beyrich, 1845 . Cardiidae. Fossil. RECORDS. 001922-32
Sherborn, p. 6658: Protoeardia (Sow.) truneatum
Beyrich, 1845. - Cardium truneatum.
truncatum Goldfuss, 1840. - Replaced by "Cardium"
transpositum Deshayes. Combination to be revised: »
Regarding Cardium truneatum: Cardiacea. Fossil.
RECORDS . 001857. Deshayes: Cardium truneatum.
Goldfuss. 1840. - C. transpositum . Desh.
truncatum Sowerby, 1827. - Combination to be revised :
812
Tropical Marine Mollusc Programme (TMMP)
truncatum - tuberculatum
"Cardium" truncatum Sowerby, 1827. Cardiidae. Lias.
RECORDS. ··1857. Deshayes: Cardium truncatum
Sowerby, 1827. ··1922-32 Sherborn, p. 6654: Cardium
truncatum Sowerby, 1827.
*truncatum; Conocardiidae - Homonymy to be
noticed. Conocardium truncatum from 2 sources:
Koninck, 1885. - Pohl, 1929.
truncatum Koninck 1885. Conocardium truncatum
Koninck 1885. Conocardiidae. Fossil. RECORDS. ··1942
ZR: Conocardium truncatum Koninck 1885 [Ref.
Branson 1942].
truncatum Poh1 1929. - Replaced by Conocardium
tripartitum. Conocardiidae. Fossil. » Regarding
Conocardium truncatum: RECORDS. ··1942 ZR:
Conocardium truncatum Pohl 1929 non Koninck 1885
replaced by Conocardium tripartitum Branson, 1942
nom. nov.
tschandae (error). - Printer's mistake for tschaudae in
Davidaschvili & Kitovani 1964.
tschandyricllm Andrusov. - Changed combination :
Andrusovicardium tschandyricum (Andrusov).
Lymnocardiidae. Neogene. RECORDS. ··1986
Nevesskaya et al.: Andrusovicardium tschandyricum
(Andrusov) comb. nov. ··1987 ZR: Andrusovicardium
tschandyricum (Andrusov) comb. nov. Ref. to
Nevesskaya et al. p. 192.
tschangeticum Poyarkova, 1978. - New combination:
Trachycardium tschangeticum (Poyarkova, 1978).
Cardiidae. Upper Cretaceous. RECORDS. ··1978 ZR:
Cardium (Trachycardium) tschangeticum Poyarkova,
1978 sp. nov. Uzbekistan. Upper Cretaceous. Ref. to
Poyarkova p. 110.
tschaudae Andrussow, 1889 - Didacna tschaudae
Andrussow, 1889. Lyrnnocardiidae. Fossil. Didacna
tschaudae Andrussow is type species of genus
Tschaudia Davitaschvili, 1956. (Davidaschvili &
Kitovani 1964) RECORDS . ··1889. ZR: Cardium
tschaudae Andrusson sic, 1889. n. sp. with fig. ··1964
Davidaschvili & Kitovani: Didacna tschaudae Andrus.
··1964 Davidaschvili & Kitovani: Didacna tschandae
sic Andrus. ··1965 ZR : Cardium tschaudae Andrusov,
1965 sp. nov. S. Russian seas. ··1970 Glibert & Poel,
p. 63: Didacna (s.s.) tschaudae tschaudae Andrusov,
1890. ··1977 Popov: Didacna (Pontalmyra) tschaudae.
Tschaudia Davidaschvili 1956 (genus).
Lymnocardiidae. Fossil. RECORDS. ··1964 Davidaschvili
& Kitovani : Tschaudia Davitaschvili, 1956. ··1980
Vokes: Tschaudia Davidaschvili, 1956 in subfamily
Didacninae. ··1987 Taktakishvi Ii: Tschaudia
Davitaschvili 1956 ; referred to new tribe
Digressodacnini.
tschelidzei Gab. - Changed combination: Macradacna
tschelidzei Gab. Lymnocardiidae. Fossil. RECORDS.
•• I 967 Ebersin: Macradacna tschelidzei Gab . •• I 967
ZR: Macradacna tschelidzei pro Monodacna
tschelidzei Gab. 1953 [Ref. Eberzin 1967].
tschepalygae Mih. - Didacna tschepalygae Mih .
Lymnocardiidae. Fossil. RECORDS. ··1989 Mikhailesku:
Didacna tschepalygae Mih.
tschernyschewi Licharew, 1931. - Conocardium
tschernyschewi Licharew, 1931. Conocardiidae. Upper
Permian . RECORDS. ··1931 ZR: Conocardillm
tschernyschewi Licharew, 1931 sp. n. Russia. Upper
Permian.
tschikojensis Maslennikov, 1961. - Cypricardinia
tschikojensis Mas1ennikov, 1961. Cardiacea. Permian.
RECORDS. ··1961. ZR: Cypricardinia tschikojensis
Maslennikov, 1961 "sp. nov.", Permian, Transbaika1ia,
Russia.
tshepalygai Mikhailescu, 1985. - Didacna tshepalygai
Mikhailescu, 1985. Lymnocardiidae. Pleistocene. TYPE
LOCALITY. The lower reaches of the Danube, Moldavia.
RECORDS. ··1985 Mikhailescu, p. 108, pI. 3, figs. 1-2:
Didacna tshepalygai sp. nov. ··1987 ZR: Didacna
tshepalygai sp. nov. Moldavia. Ref. to Mikhailescu p.
105.
tuba Camacho, 1960. - Combination to be revised :
"Cardium " tuba Camacho, 1960. Cardiidae .
Paleocene. RECORDS. ··1960 ZR: Cardium tuba
Camacho, 1960 sp. nov. Paleocene River Ewan, Tierra
del Fuego.
tuberclllare Sowerby 1820. - Synonym of
Acanthocardia tuberculata (Linne, 1758). »
Regarding Cardium tuberculare: Cardiidae. Recent.
RECORDS. ··182 I - 1834 Sowerby, fig. 3: Cardium
tuberculare (only figure). ··1850 Jay: Cardium
tuberculare Sowb. = Cardium rusticum . ··1856 Wood
& Hanley: C. tuberculare Sow. G. f. 3 = no . 25 =
Cardium tubercula tum Lin . •• I 867 Weinkauff: Cardium
tuberculare Sowerby = Cardium tuberculatum Linne.
··1869. Romer: Cardium tuberculare Sowerby =
Cardium tuberculatum L. ··1888 Kobelt: Cardium
tuberculare Sowerby = Cardium tubercula tum Linne.
··1890 Paetel : Cardium tuberculare Sow. =
tuberculatum L.
tuberculata Chemnitz 1782. Non binominal. - Not
available . •• I 845 Reeve in the index: Cardium
tuberculatum in Chernnitz 1782 = Cardium rusticum
Linne, 1767. ··1850 Jay: Cardium tubercula tum
Chernn. = Cardium rusticum.
tuberculata of Pult. - Misidentification. Acanthocardia
tuberculata (Linnaeus, 1758). ··1803 Montagu :
Cardium tuberculatum Pult. Cat. Dorset. p. 30? =
Cardium aeuleatum Lin.
tuberculatum (mode) . - See under tuberculata Linne,
1758.
tuberculatum Brongniart , 1821. - Hemicardium
tuberculatum Brongniart, 1821. Cardiidae. Fossil.
RECORDS. ··1922-32 Sherborn, p. 6667: Hemicardium
tubereulatum Brongniart, 1821.
tuberculatum Lamarck (variety). [Figs. Part 3 p. 847].Lamarck collection, MNHN Paris . No published
records seen.
tuberculatum Linnaeus, 1758 / Gmelin, 1791 . (Figs. Part
I p. 39, 42,44,46, 53, 55, 58,64, 68, 74,92,96, 126,
167, 171. Part 3 p. 846,847, 849]. - Changed
combination: Acanthocardia tuberculata (Linne, 1758).
Cardiidae. Pliocene and Recent. Type species: Keen
(1937) designated Cardium tuberculatum Linne, 1758
as type species of the subgenus Rudicardium Coen,
1915. TYPE LOCALITY. Unknown (Linne, 1758); Britain
and the Mediterranean (Sowerby 1841). RECORDS.
··1684 Bonanno. ID by Gmelin 1791: Cardium
tubercula tum recf. 2. f. 100. ··1687 Lister. ID by Gmelin
1791: Cardium tubercula tum t. 329. f. 166. ··1705
Rurnphius. ID by Linnaeus 1758 and Gmelin 1791:
Cardium tuberculatum mus. t. 48 . f. I I. ··1742
Argenville. ID by Linnaeus 1758: Cardium
Phuket Marine Biological Center Special Publication vol. 30 (2004)
813
tuberculatum
tuberculatum conch. t. 26. f. L. --1742 Gualtieri, pI.
71, fig. M: = Cardium tuberculatum (fide Spengler
1799). --1758 Linne, p . 679, No . 65: Cardium
tuberculatum. --1758 Regenfuss. ID by Gme1in 1791:
Cardium tuberculatum Conch. I . t. 8. f. 23. 24. --1758
Seba. ID by Gmelin 1791: Cardium tuberculatum mus.
3. t. 86. f. 7. --1764 Linnaeus (Mus. Lud. Ulr. Reg.)
No. 37. --1767 Linne, p. 1122, sp. 81: Cardium
tuberculatum. --1771 Knorr. ID by Gme1in 1791:
Cardium tuberculatum Vergn. 5. t. 30. f. 2. --1771
Knorr, pI. 30, fig. 2: Cardium tuberculatum (fide
Spengler 1799). --1772 ArgenviUe. ID by Gme1in 1791:
Cardium tuberculatum zoom. t. 6. f. C. D. --1772 Knorr.
ID by Gmelin 1791: Cardium tubercula tum Vergn. 6.
t. 8. f. 3. (Same ref. as to C. Jasciatum) . --1778 Born,
G8 . Cardium tuberculatum Linn. S.N. Sp. 81. "1780
Born: Cardium tuberculatum Linn. S. N . Sp. 81. --1782
Bonannio. ID by Gmelin 1791 : Cardium tubercula tum
mus Kirch. 2. f. 100 ... 1782 Chemnitz, Tab. 17 Fig.
173: Cardium tuberculatum Lin. --1782 Chemnitz. ID
by Gmelin 1791: Cardium tubercula tum t. 17. f. 173.
--1786 Schroter, No. IX: Cardium tuberculatum Linn.
XII . pag. 1122. Spec. 81. --1788 Schroter: Cardium
tuberculatum. [citing Chemnitz (1782) figs. 179. 180].
--1789 Bruguiere, No. 12. Cardium tuberculatum; Linn.
--1791 Gmelin No. II: Cardium tuberculatum. --1798
Bruguiere. ID by Lamarck, pI. 300. fig. 1. Cardium
tuberculatum. --1799 Spengler, p. 19, No. 8: Cardium
tuberculatum Linnei Sp. 81 . p. 1122. --1800 Donovan,
PI. 27, fig. 9 Cardium tuberculatum . "1803 Montagu:
Cardium tuberculatum . .. 1807 Maton & Rackett, No.
7 : Cardium tuberculatum Linn . --1817 Dillwyn:
Cardium tuberculatum. Linnaeus. --1823 Dillwyn, Tab.
329 fig. 166 Cardium tuberculatum . --1829 Costa: Poli
considered C. tuberculatum a variety of C. rusticum.
--1832 Deshayes: Cardium tuberculatum, L. = Cardium
echinatum L. "1836 Bose: Cardium tuberculatum.
--1840 Sowerby, catalogue p. 2, No. 28 : Cardium
tuberculatum, Linn. --1844 Potiez & Michaud:
Cardium tuberculatum Linn . --1850 Jay: Cardium
tubercula tum Linn. = Cardium echinatum, --1856
Wood & Hanley, No, 4: Cardium tuberculatum Lin.
--1858 Adams & Adams : Cardium (lsocardia)
tuberculatum Linn. --1859 Sowerby: C. tubercula tum,
Linn, ? = Cardium rusticum, Linn. --1868 ZR: Cardium
tuberculatum (L.), this name to be restored for rusticum
(L., Reeve) [Ref. Romer 1868]. --1884 Grasset:
Cardium tuberculatum. L. --1888 Servain: Cardium
tuberculatum Linnaeus, 1767. --1890 Paetel: Cardium
tuberculatum L. --1891 Romer : Acanthocardia
tuberculatum L. --1892 B. D. & D. p. 256-261: Cardium
tuberculatum , Linne ... 1902 Sherborn, p. 1003 :
Cardium tuberculatum Linnaeus, 1758. --1907 ZR:
Cardium tuberculatum [Refs. Roule; Zimmermann].
--1914 Coen: Eucardium (Rudicardium) tuberculatum
L. forma typica . --1932 Winckworth: Cardium
(Acanthocardia) tuberculatum Linne, 1758. --1933
Coen : Eucardium (Rudicardium) tubercula tum Linneo.
"1933 Coen & Vatova: Eucardium (Rudicardium)
tubercula tum Linnaeus. --1937 Coen: Cerastoderma
(Rudicardium) tubercula tum Linneo (Note 259: 8
varieties described by Coen in 1915). "1937 Keen
designated Cardium tuberculatum Linne, 1758. --1937
ZR: Cardium tuberculatum [Ref. Avnimelech 1937].
--1937 ZR: Cardium tuberculatum L. , 1758 type of
Rudicardium Coen 1915 [Ref. Keen 1937]. "1937 ZR:
Cerastoderma (Rudicardium) tuberculatum L. [Ref.
Coen 1937], ,,1938, ZR: Cardium tuberculatum fig .
luist [Ref. Girscher 1938a]. "1939 Adam & Leloup:
Cardium (Rudicardium) tuberculatum Linne 1758.
--1953 ZR: Cardium tubercula tum L. structure of shell
[Ref. Saban 1953]. "1959. Devidts: Cerastoderma
(Rudicardium) tuberculatum Linne. --1959 ZR:
Cardium tuberculatum enzyme system activated by
magnesium ions [Ref. Roche, et at. 1959]. --1962
Pasteur-Humbert: Cardium tubercula tum Linne. --1962
ZR: Cardium tuberculatum photo., shell shape and
co10uration [Ref. Pfeiffer 1962]. --1963 ZR: Cardium
tuberculatum [Santarelli 1963: from Italy] . --1963. ZR:
Cardium tuberculatum [Kjolberg et at. 1963 : properties
of some glycogens found in tissues]. "1966 ZR:
Cardium tuberculatum [Salanki 1966: experiments on
the rhythmic activity]. ..1966 ZR: Cardium
tuberculatum [Ref. Salanki & Zs.-Nagy 1966:
investigation of the adductor muscle]. "1969
Nordsieck: Rudicardium tuberculatum tuberculatum
(Linne, 1758). --1977 Fischer-Piette: Cardium
(Cerastoderma, Rudicardium) tuberculatum Linne.
--1978 Janssen & Van der Slik : Acanthocardia
(Rudicardium) tuberculata (Linnaeus, 1758). --1980
ZR : Acanthocardia (Rudicardium) tuberculatum.
Adriatic Sea, Ref. to Marano. et al. --1980 ZR: Cardium
tuberculatum, Ref. to Gade . --1981 ZR: Cardium
tuberculatum. Ref. to Ansell et ai , --1982 ZR:
Acanthocardia tuberculata. Ref. to Savazzi. --1983 ZR:
Acanthocardia tuberculata. Ref. to Ivell, Meinardus et
at. - Cerastoderma tuberculatum. Ref. to Cano Perez.
--1984 ZR: Acanthocardia (Rudicardium) tuberculata.
Ref. to Marano et at. --1985 ZR Cardium tuberculatum.
Ref. to Chrispeel et at. / Meinardus et al. --1987 ZR:
Acanthocardia tuberculata. Ref. to Vigna Ii et at. --1987
ZR: Cardium tuberculatum. Ref. Meinhardus-Hager et
al. --1993 Poutiers: Acanthocardia tuberculata (Linne,
1758). --1994 Schneider: Acanthocardia (Rudicardium)
tuberculata (Linne, 1758). --1998 Macedo et at.:
Acanthocardia tuberculata (Linnaeus, 1758), --1999
Jacques Vidal (pers. com.) MNHN, Paris: Fossils from
the Paleontological Collection as Cardium
tuberculatum Linne. Pliocene. France; Turkey; Tunesia.
--200 1 Vidal, p. 407: Acanthocardia tuberculatum.
--2002 Schneider: Acanthocardia (Rudicardium)
tuberculata (Linne, 1758). - CHARACTERS AND
OCCURRENCE - OF COLLECTIONS. --ZMUC: 2
specimens Spengler coil. (Mediterraneum. p. 19 no.
8). ZMUC wet collection. DESCRIPTIONS. --1758
Linnaeus : shell subcordate, interstices transversely
striated, obtuse nodes [I have translated from Latin].
--1799 Spengler: Testa subcordata, sulcis obtusis
nodosis transversim striatis. Linnei Sp. 81, p, 1122.
Colour very variable. Completely white or white with
brown or yellow transverse bands ... 200 1 Vidal:
Tentacles around siphons only. No eyes on tentacles.
REMARKS. The following authorship are not available:
Gualtieri 1742, Knorr 1771, Chernnitz 1782. Requien
made iconographic reference to pI. 4, fig. II in Philippi
(1844) but that figure shows Tapes decussatus
according to Locard & Caziot (1900). According to
Dillwyn (l817) Mr. Montagu doubted whether
814
Tropical Marine Mollusc Programme (TMMP)
tubereulatum - tubereulatum eoaleseens
Cardium tubereulatum Linnaeus should be considered
a distinct species. HABITAT. 001876 Duprey: Cardium
tubereulatum, L. intertidal on the coast of New Jersey
with a tidal range of about 13 m. 001969 Nordsieck:
among laminarians and in sand, 15-100 m depth.
DISTRIBUTION. J807 Maton & Rackett: Dorset, Devon.
001884 Grasset: Algeria. 001891 Romer: Mediterranean.
001969 Nordsieck ; West Europe, Mediterranean,
Canary Islands, Madeira . ooZMUC West Europe,
Mediterranean.
tubereulatum Pusch, 1836. - Cardium tubereulatum
Pusch, 1836. ?Cardiidae. Fossil. RECORDS. 001922-32
Sherborn, p. 6664: Cardium (Br.) tuberculatum Pusch,
1836. - Hemieardium.
tubereulatum aeutieostis Coen, 1937 (variety). Infrasubspecific. No evidence of subspecific status.
Aeanthoeardia tubereulata (Linne, 1758). »
Regarding A. tuberculata var. aeutieostis: Cardiidae.
Recent. RECORDS. 001933 Coen: Eueardium
(Rudicardium) lubereulatum var. aeuticostis Sta1io mss.
The Mediterranean . 001937 Coen, Note 260 :
Cerastoderma (Rudicardium) lubereulatum var.
aeutieostis (Stalio mss.) Co en n. var. 001937 ZR:
Cerastoderma (Rudicardium) tubereulatum L. var.
aeulieostis (Stalio mss .) Coen, 1937 nov. Adriatic.
001937 ZR: Eueardium (Rudieardium) tubereulatum L.
var. aeutieostis (Stalio mss. var. anom.) Coen, 1933
nov.
tubereu/atum a/bidum Danilo & Sandri, 1891 (variety).
- Infrasubspecific. No evidence of subspecific status.
Aeanthoeardia tub erculata (Linne, 1758). »
Regarding A. lubereulata var. albida: Cardiidae.
Recent. RECORDS. 001789 Karsten: Cardium
tubereulatum testa extus et intus albida. 001891 Danilo
& Sandri: Cardium tuberculalum Linn. var. albidum
nob. 001892 B.D. & D. p. 261: var. albidum Brusina =
Cardium tuberculatum var. ex colore alba Monterosato.
001914 Coen: Cardium tubercula tum vaT. albida
Brusina = Eueardium (Rudicardium) lubereulatum var.
alba Monts. - CHARACTERS AND OCCURRENCE
- DESCRIPTIONS. White, milkish, uniform colour.
REMARKS. Brusina wrote the introduction to Danilo &
Sandri (1891).
tubereulatum a/bum Monterosato, 1878 (variety). Infrasubspecific. No evidence of subspecific status.
Acanlhoeardia tuberc ulata (Linne , 1758). »
RegardingA . tubereulata var. alba: Cardiidae. Recent.
RECORDS. 001878 Monterosato: Cardium tubereulatum,
Lin . var. ex col. alba. 001892 B. D. & D. p. 261:
Cardium tubercula tum var. ex colore alba Monterosato.
001900 Locard & Caziot: var. alba (Mtr.) = Cardium
tubereulatum Linne. 001914 Coen: Eueardium
(Rudieardium) tubereulatum var. alba Monts. 001933
Coen: Eueardium (Rudieardium) tubereulalum var.
alba Monterosato. 001937 Coen: Cerasloderma
(Rudieardium) tubereulatum album Monterosato.
001969 Nordsieck: Rudieardium tubercula tum
tubercula tum (Linne, 1758) var. album (Monterosato).
- CHARACTERS AND OCCURRENCE - OF
COLLECTIONS. ooZMUC: two specimens labeled "90. B.
C. tb . 26 album , Sardinien". DESCRIPTION. White,
milkish, uniform colour. DI STRIBUTION. The
Mediterranean.
tubereulatum alta Pallary, 1904 (variety). 00
Infrasubspecific. No evidence of subspecific status.
Aeanthoeardia tubereulata (Linne, 1758). »
RegardingA. tubercula tum var. alta: Cardiidae. Recent.
RECORDS. 001904 Pallary, p. 246: Cardium tubereulatum
var. alta Pallary.
tuberculatum angulatum Coen, 1933 (variety). Infrasubspecific. No evidence of subspecific status.
Aeanthoeardia tub ereu lata (Linne, 1758). »
Regarding A. tubereulatum var. angulata: Cardiidae.
Recent. RECORDS. 001933 Coen: Eueardium
(Rudieardium) tubereulatum var. angulata Coen. The
Mediterranean. 001937 Coen, Note 260: Cerastoderma
(Rudieardium) tubereulatum Linn. var. angulatum
Coen n. var. 001937 ZR: Cerastoderma (Rudieardium)
tubereulatum L. vaT. angulata (Stalio mss.) Coen, 1937
nov. Adriatic. 001937 ZR: Eueardium (Rudieardium)
tubereulatum L. var. angulata (Stalio mss. var. anom.)
var. Coen, 1933 nov.
tubereulatum asperula Coen , 1914 (variety). Infrasubspecific. No evidence of subspecific status.
Aeanthoeardia tubereulata (Linne, 1758). »
Regarding A. tubereulata var. asperula: Cardiidae.
Recent. RECORDS. 001914 Coen: Eueardium
(Rudieardium) tubereulatum n. var. asperula Coen.
001915 ZR: Cardium tubercula tum var. asperula Coen,
1915 var. n. Adriatic. 001933 Coen: Eueardium
(Rudieardium) tubereulatum var. asperula Coen.
tubereulatum beta Gmelin, 1791 (variety). [Figs. Part I
p. 58]. - Unknown status. RECORDS. 001758 Seba. ID
by Gmelin 1791 : Cardium tubereulatum var. beta, mus.
3. t. 86. f. 2. 001791 Gme1in No . 11. Cardium
tubercula tum var. beta.
tubereulatum citrinum Danilo & Sandri, 1891 mss
(variety). - Infrasubspecific. No evidence of subspecific
status. Aeanthoeardia tubereulata (Linne, 1758). »
Regarding A. tubereulata var. citrina: Cardiidae.
Recent. RECORDS. 001891 Danilo & Sandri: Cardium
tubereulalum var. eitrinum nob. 001892 B. D. & D. p.
261: Cardium tubercula tum var. ex forma et colore
citrina Brusina. 001914 Coen: Eucardium
(Rudicardium) tuberculatum var. ex forma et colore
citrina Brusina. 001933 Coen: Eucardium
(Rudicardium) tubereulatum var. citrina Brusina.
001937 Coen: Cerastoderma (Rudicardium)
tuberculatum citrinum Brusina . 001969 Nordsieck:
Rudicardium tubercula tum citrinum (Brusina, 1865).
- CHARACTERS AND OCCURRENCE DESCRIPTIONS. With 19, angled ribs (normally 24 ribs);
colour yellow with dark brown bands. HABITAT. Sandy
bottom at 8-73 m depth (Nord sieck 1969). DISTRIBUTION.
001892 B.D. & D.: Adriatic Sea off Albany. 001969
Nordsieck: unknown.
tuberculatum coalescens Coen, 1937 (variety). Infrasubspecific. No evidence of subspecific status.
Aeanth oea rdia tubereulata (Linne, 1758). »
Regarding A. tubereulatum var. eoaleseens: Cardiidae.
Recent . RECORDS. 001933 Coen: Eu ea rdium
(Rudieardium) tubereulalum var. eoaleseens Coen. The
Mediterranean. 001933 ZR: Rudicardium tubereulatum
var. eoaleseens Coen, 1933 nov. Venice [Ref. Coen
1933] . 001937 Coen, Note 261, pI. 10, fig. 92:
Cerastoderma (Rudieardium) tuberculatum Linn. var.
anomlia eoaleseens Coen n. var. 001937 ZR:
Cerastoderma (Rudieardium) tubereulatum L. var.
Phuket Marin e Biological Center Special Publication vol. 30 (2004)
815
tubereulatum dauizellbergi - tuberculatum multifasciata
coalescens (Spinelli mss.) Coen, 1937 nov. Adriatic.
"1937 ZR: Rudicardium tuberculatum var. coalescens
Coen, 1933 nov. Venice. "1937 ZR: Eucardium
(Rudicardium) tuberculatum L. var. coalescens (var.
anom.) Coen, 1933 nov.
(uberculatum dautzellbergi Coen, 1914 (variety) . Infrasubspecific. No evidence of subspecific status.
Acantho cardia tuberculata (Linne, 1758) . »
Regarding A. tuberculatum var. dautzenbergi :
Cardiidae. Recent. RECORDS. "1914 Coen: Eucardium
(Rudicardium) tuberculatum var. dautzenbergi Coen n.
var. .. 1915 ZR: Cardium tuberculatum var.
dautzenbergi Coen, 1915 var. n. Adriatic. "1933 Coen:
Eucardium (Rudicardium) tuberculatum var.
dautzenbergi Coen. "1937 Coen: Cerastoderma
(Rudicardium) tubercula tum dautzenbergi Coen.
tubereulatum delta Gmelin, 1791 (variety). [Figs. Part
I p. 60). - Unknown status. RECORDS. "1758 Seba. ID
by Gmelin 1791 : Cardium tubereulatum var. delta mus.
3. t. 86. f. 14. "1791 Gmelin No. II: Cardium
tubercula tum var. delta.
tubereulatum depauperatum Coen, 1937 (variety). Infrasubspecific. No evidence of subspecific status.
Acanthocardia tuberculata (Linne , 1758). »
Regarding A. tuberculatum var. depaup erata :
Cardiidae. Recent. RECORDS. "1933 Coen: Eucardium
(Rudicardium) tuberculatum var. depauperata Coen.
The Mediterranean. "1937 Coen , Note 260:
Cerastoderma (Rudicardium) tubercula tum var.
depauperatum Coen n. var. "1937 ZR: Cerastoderma
(Rudicardium) tuberculatum L. var. depauperata
(Stalio mss.) Coen, 1937 nov. Adriatic. "1937 ZR:
Eucardium (Rudicardium) tuberculatum L. var.
depauperata (Stalio mss. var. anom.) var. Coen, 1933
nov.
tubereulatum elegalls Danilo & Sandri, 1891 mss
(variety). - Infrasubspecific. No evidence of sub specific
status. Acanthocardia tuberculata (Linne, 1758). »
Regarding A. tuberculatum var. elegans: Cardiidae.
Recent. RECORDS. "1891 Danilo & Sandri: Cardium
tuberculatum var. elegans nob. "1892 B. D. & D. p.
261: Cardium tubercula tum var. ex forma elegans
Brusina. "1914 Coen: Eucardium (Rudicardium)
tuberculatum var. ex forma elegans Brusina. "1933
Coen: Eucardium (Rudicardium) tubercula tum var.
elegans Brusina. "1937 Coen: Cerastode rma
(Rudicardium) tubercula tum elegans Brusina. "1969
Nordsieck: Rudicardium tuberculatum tuberculatum
(Linne, 1758) var. elegans (Brusina). - CHARACTERS
AND OCCURRENCE - DESCRIPTIONS. Elevated ribs
with tubercles; interstices with undulating transverse
stripes. Lunule deep . REM ARKS. Brusina wrote the
introduction to Danilo & Sandri (1891) . DISTRlBUTION.
"1969 Nordsieck : unknown.
tubereulatum epsiloll Gmelin, 1791 (variety) . Unknown
status. RECORDS. "1764 Knorr. ID by Gmelin 1791:
Cardium luberculatum var. epsilon, Vergn. 2. t. 29. f.
4. "1791 Gmelin No. II : Cardium tubercula tum var.
epsilon .
tubereulatum fuseapepratx (variety). - Infrasubspecific.
No evidence of subspecific status. Acanthocardia
tuberculata (Linne , 1758) . » Regarding A.
tuberculatum var. fuscapepratx : Cardiidae . Recent.
RECORDS ... 1914 Coen : Eucardium (Rudicardium)
tubercula tum var. ex col.fuscapepratx.
tubereulatum fuscum Pepratx , 1884 (variety) . Infrasubspecific . No evidence of subspecific status.
Acanthocardia tub erculata (Linne, 1758). »
Regarding A. tuberculatum var. fusca : Cardiidae .
Recent. RECO RDS. "1884 Pepratx : Cardium
tuberculatum var. fuscum . "1892 B. D. & D. p. 261:
Cardium tuberculatum var. ex colore fus ca pepratx.
"1969 N ordsieck : Rudicardium tuberculatum
tuberculatum (Linne, 1758) var.fuscum (Petratx sic).
tuberculatum fuscum Philippi (variety).
Infrasubspecific. No evidence of subspecific status.
Acanthocardia tuberculata (Linne , 1758). »
Regarding A. tubercula tum var. fusca: Cardiidae .
Recent. RECORDS. "1933 Coen : Eu cardium
(Rudicardium) tuberculatum var.fusca Philippi. "1937
Coen : Cerastoderma (Rudicardium) tuberculatum
fuscum Philippi.
tuberculatum gamma Gmelin, 1791 (variety). [Figs. Part
I p . 59). - Unknown status . RECORDS. "1758 Seba. ID
by Gmelin 1791 : Cardium tubercula tum var. gamma,
mus. 3. t. 86. f. 6. "179 J Gmelin No. II : Cardium
tubercula tum var. gamma.
tuberculatum lactea Clement, 1879 (variety). Infrasubspecific. No evidence of subspecific status.
Acanthocardia tuberculata (Linne , 1758). »
Regarding A. tuberculatum var. lactea: Cardiidae.
Recent . RECO RDS. "1879 Clement: Cardium
tuberculatum var. lactea. "1892 B. D. & D. p. 261:
var. lactea Clement = Cardium tuberculatum var. ex
colore alba Monterosato. "1914 Coen: Eucardium
(Rudicardium) tuberculatum var. lactea Coen. "1914
Coen: Cardium tuberculatum var. lactea Clement (non
Coen) = Eucardium (Rudicardium) tuberculatum var.
alba Monts. "1914 ZR: Eucardium (Cerastoderma)
tuberculatum lactea var. n. Coen, 1914. "1933 Coen:
Eucardium (Rudicardium) tubercula tum var. lactea
Coen. "1937 Coen: Cerastoderma (Rudicardium)
tuberculatum lacteum Coen. - CHARACTERS AND
OCCURRENCE - DESCRIPTIONS. White, milkish,
uniform colour.
tuberculatum millor Monterosato, 1878 (variety). Infrasubspecific . No evidence of subspecific status.
Acanthocardia tuberculata (Linne, 1758). »
Regarding A. tuberculatum var. minor: Cardiidae.
Recent. RECORDS. "1789 Karsten: Cardium tuberculati
varietas minores. "1878 Monterosato: Cardium
tuberculatum, Lin. var. ex forma minor. "1892 B. D.
& D. p. 261: Cardium tuberculatum var. ex forma minor
Monterosato. "1900 Locard & Caziot: var. minor (MIr.)
= Cardium tuberculatum Linne. "1914 Coen:
Eucardium (Rudicardium) tuberculatum var. ex forma
minor Monterosato. "1969 Nordsieck: Rudicardium
tuberculatum tuberculatum (Linne, 1758) var. minor
(Monterosato ).
tuberculatum millus Karsten , 1789 (variety). Infrasubspecific. No evidence of subspecific status.
Acanthocardia tuberculata (Linne, 1758). RECORDs. »
Regarding A. tuberculatum var. minus : Cardiidae .
Recent. "1789 Karsten: Cardium tuberculatum testa
minus.
tuberculatum multifasciata Danilo & Sandri, 1891 mss
(variety). - Infrasubspecific. No evidence of subspecific
status. Acanthocardia tuberculata (Linne, 1758). »
816
Tropical Marine Mollusc Programme (TMMP)
tuberculatum multiplicatum - tuberculatum torquata
Regarding A. tuberculatum var. multifasciatum:
Cardiidae. Recent. REMARKS. Brusina wrote the
introduction to Danilo & Sandri (1891). RECORDS.
··1891 Danilo & Sandri: Cardium tuberculatum var.
multifasciatum nob. ··1892 B. D. & D. p. 261: var.
multifasciata Brusina = Cardium tuberculata var. ex
colore zonata Monterosato. ··1914 Coen: Cardium
tuberculatum var. multifasciata Brusina = Eucardium
(Rudicardium) tubercula tum var. ex col. zonata
Monterosato. - CHARACTERS AND OCCURRENCE - DESCRIPTIONS. White with several bands of
marked brown colour.
tuberculatum multiplicatum Coen, 1933 (variety). Infrasubspecific. No evidence of subspecific status.
Acanthocardia tuberculata (Linne, 1758). »
Regarding A. tuberculatum var. multiplicata:
Cardiidae. Recent. RECORDS. ··1933 Coen: Eucardium
(Rudicardium) tuberculatum var. multiplicata Coen.
··1937 Coen, Note 260: Cerastoderma (Rudicardium)
tuberculatum var. multiplicatum Coen n. var. ··1937
ZR: Cerastoderma (Rudicardium) tubercula tum L. var.
multiplicata (Stalio mss.) Coen, 1937 nov. Adriatic.
··1937 ZR: Eucardium (Rudicardium) tuberculatum L.
var. multiplicata (Statio mss. var. anom.) Coen, 1933
nov.
tuberculatum muticum Bucquoy, Dautzenberg &
Dollfus, 1892 (variety). - InfTasubspecific. No evidence
of subspecific status. Acanthocardia tuberculata
(Linne, 1758). » Regarding A. tuberculatum var.
mutica: Cardiidae. Recent. RECORDS. ··1892 Bucquoy,
Dautzenberg & Dollfus, p. 261: Cardium tuberculatum
var. ex forma mutica B.D.D. ··1900 Locard & Caziot:
var mutica (B.D. & D.) = Cardium tuberculatum Linne.
··1914 Coen: Eucardium (Rudicardium) tuberculatum
var. mutica B.D.D. ··1914 ZR: Eucardium
(Cerastoderma) tuberculatum mutica var. n. Coen,
1914. ··1933 Coen: Eucardium (Rudicardium)
tuberculatum var. mutica B.D.D. ··1937 Coen:
Cerastoderma (Rudicardium) tuberculatum muticum
B.D.D. ··1969 Nordsieck: Rudicardium tuberculatum
tuberculatum (Linne, 1758) var. muticum (Bucquoi (sic
Bucquoy), Dautzenberg & Dollfus).
CHARACTERS AND
OCCURRENCE
DESCRIPTIONS. Without tubercles on the ribs.
DISTRIBUTION. The Mediterranean.
tuberculatum palaeomutica Coen, 1914 (variety). Infrasubspecific. No evidence of subspecific status.
Acanthocardia tuberculata (Linne, 1758). »
Regarding A. tuberculatum var. palaeomutica:
Cardiidae. New name for fossil varieties (Coen 1914).
RECORDS. ··1914 Coen: Eucardium (Rudicardium)
tuberculatum var. palaeomutica Coen nom. nov.
suggested for the fossil records of tubercula tum var.
mutica.
tuberculatum pharisiana Pallary, 1924 (variety). Infrasubspecific. No evidence of subspecific status.
Acanthocardia tuberculata (Linne, 1758). »
Regarding A. tuberculatum var. pharisiana: Cardiidae.
Recent. RECORDS. ··1924 ZR: Cardium tubercula tum
pharisiana Pallary, 1924 var. n. Tunis.
tuberculatum picta Coen (variety). - Infrasubspecific.
No evidence of subspecific status. Acanthocardia
tuberculata (Linne, 1758). » Regarding A.
tuberculatum var. picta: Cardiidae. Recent. RECORDS.
··1914 Coen: Eucardium (Rudicardium) tuberculatum
var. picta n. var. ··1915 ZR: Cardium tuberculatum var.
picta Coen, 1915 var. n. Adriatic. ··1933 Coen:
Eucardium (Rudicardium) tuberculatum var. picta
Coen. ··1937 Coen: Cerastoderma (Rudicardium)
tuberculatum pictum Coen.
tuberculatum potens Monterosato Mss (variety). Infrasubspecific. No evidence of subspecific status.
Acanthocardia tuberculata (Linne, 1758). »
Regarding A. tuberculatum var. potens: Cardiidae.
Recent. RECORDS. ··1914 Coen: Eucardium (Rudicardium) tuberculatum var. potens Monterosato Mss.
tuberculatum ruderata Monterosato (variety). Infrasubspecific. No evidence of sub specific status.
Acanthocardia tuberculata (Linne, 1758). »
Regarding A. tuberculatum var. ruderata: Cardiidae.
Recent. RECORDS. ··1914 Coen: Eucardium
(Rudicardium) tuberculatum var. ex forma ruderata
Mss. The Mediterranean. ··1933 Coen: Eucardium
(Rudicardium) tubercula tum var. ruderata Monterosato
mss. ··1937 Coen: Cerastoderma (Rudicardium)
tuberculatum ruderatum Monter. mss.
tuberculatum rudis Coen (variety). - Infrasubspecific.
No evidence of subspecific status. Acanthocardia
tuberculata (Linne, 1758). » Regarding A.
tuberculatum var. ntdis: Cardiidae. Recent. REcORDs.
··1914 Coen: Eucardium (Rudicardium) tuberculatum
var. ex forma rudis Coen. ··1914 ZR: Eucardium
(Cerastoderma) tuberculatum rudis var. n. Coen, 1914.
··1933 Coen: Eucardium (Rudicardium) tuberculatum
var. rudis Coen. ··1937 Coen: Cerastoderma (Rudicardium) tuberculatum rude Coen.
tuberculatum spinosa Coen, 1914 (variety). InfTasubspecific. No evidence of specific or subspecific
status. Acanthocardia tuberculata (Linne, 1758). »
Regarding A. tuberculatum var. spinosa: Cardiidae.
Recent. RECORDS. ··1914 Coen: Eucardium
(Rudicardium) tubercula tum var. ex forma spinosa
Coen n. var. The Mediterranean. ··1915 ZR: Cardium
tuberculatum var. spinosa Coen, 1915 var. n. Adriatic.
··1933 Coen: Eucardium (Rudicardium) tubercula tum
var. spinosa Coen. = tuberculatum unifasciata Danilo
& Sandri mss. ··1937 Coen: Cerastoderma
(Rudicardium) spinosum Coen.
tuberculatum suborbicula Marshall, 1893 (variety). Infrasubspecific. No evidence of specific or subspecific
status. Acanthocardia tuberculata (Linne, 1758). »
RegardingA. tuberculatum var. suborbicula: Cardiidae.
Recent. RECORDS. ··1893 Marshall, p. 246: Cardium
tuberculatum var. suborbicula Marsh. (n. var.) Britain.
··1901 Chaster et al.: Cardium tuberculatum v.
suborbicula Marshall. ··1926 McClelland: Cardium
tubercula tum suborbicula Marshall, 1893 n. sp.
tuberculatum torquata Coen, 1914 (variety). Infrasubspecific. No evidence of subspecific status.
Acanthocardia tuberculata (Linne, 1758). »
Regarding A. tuberculatum var. torquata: Cardiidae.
Recent. RECORDS. ··1914 Coen: Eucardium
(Rudicardium) tuberculatum var. torquata Coen n. var.
The Mediterranean. ··1915 ZR: Cardium tuberculatum
var. torquata Coen, 1915 var. n. Adriatic. ··1933 Coen:
Eucardium (Rudicardium) tuberculatum var. torquata
Coen. ··1937 Coen: Cerastoderma (Rudicardium)
tuberculatum torquatum Coen.
Phuket Marine Biological Center Special Publication vol. 30 (2004)
817
tuberculatum unifasciata - tumidum
tuberculatum unifasciata Danilo & Sandri, 1891 mss
(variety). - Infrasubspecific. No evidence of sub specific
status. Acanthocardia tuberculata (Linne, 1758). »
Regarding A. tuberculatum var. unifasciatum:
Cardiidae. Recent. RECORDS. "1891 Danilo & Sandri:
Cardium tuberculatum var. unifasciatum nob. --1892
B. D. & D. p. 261: Cardium tuberculata var. ex colore
unifasciata Brusina. - CHARACTERS AND OCCURRENCE - DESCRIPTIONS. White, with a single transverse
band of brown colour. REMARKS. Brusina wrote the
introduction to Danilo & Sandri (1891).
tuberculatum vittatum Danilo & Sandri, 1891 mss
(variety). - Infrasubspecific. No evidence of subspecific
status. Acanthocardia tuberculata (Linne, 1758). »
Regarding A. tuberculatum var. vittatum: Cardiidae.
Recent. RECORDS. "1891 Danilo & Sandri: Cardium
tuberculatum var. vittatum nob. --1892 B. D. & D. p.
261: Cardium tuberculatum var. ex colore vittata
Brusina. "1914 Coen: Eucardium (Rudicardium)
tuberculatum var. vittata Brusina. "1933 Coen:
Eucardium (Rudicardium) tuberculatum var. vittata
Brusina. "1937 Coen: Cerastoderma (Rudicardium)
tubercula tum vittatum Brusina. --1969 Nordsieck:
Rudicardium tuberculatum tuberculatum (Linne, 1758)
var. vittatum (Brusina). - CHARACTERS AND
OCCURRENCE - DESCRIPTIONS. Characterized by a
white rayon rib 17 and the adjoining two interspaces.
REMARKS. Brusina wrote the introduction to Danilo &
Sandri (1891). DISTRIBUTION. "1969 Nordsieck:
unknown.
tuberculatum zeta Gmelin, 1791 (variety). [Figs. Part I
p. 62]. - Unknown status. RECORDS. --1768 Knorr. ID
by Gmelin 1791: Cardium tubercula tum var. zeta,
Vergn. 3. t. 4. f. 5. "1791 Gmelin No. 11: Cardium
tuberculatum var. zeta.
tuberculatum zonata Monterosato, 1878 (variety). Infrasubspecific. No evidence of subspecific status.
Acanthocardia tuberculata (Linne, 1758). »
Regarding A. tubercula tum var. zonata: Cardiidae.
Recent. RECORDS. --1878 Monterosato: Cardium
tuberculatum, Lin. var. ex col. zonata. "1892 B. D. &
D. p. 261: Cardium tuberculata var. ex colore zonata
Monterosato. --1900 Locard & Caziot: var. zonata
(Mtr.) = Cardium tuberculatum Linne. "1914 Coen:
Eucardium (Rudicardium) tuberculatum var. ex col.
zonata Monterosato. --1933 Coen: Eucardium
(Rudicardium) tuberculatum var. zonata Monterosato.
--1937 Coen: Cerastoderma (Rudicardium)
tubercula tum zonatum Monterosato. - CHARACTERS
AND OCCURRENCE - DESCRIPTIONS. White with
several bands of marked brown colour. DISTRIBUTION.
Mediterranean. --1900 Locard & Caziot: unknown.
tuberosum Jekelius, 1944. - New combination:
Lymnocardium tuberosum JekeJius, 1944.
Lymnocardiidae. Miocene. RECORDS. --1946 ZR:
Limnocardium tuberosum Jekelius, 1944 sp. nov.
Miocene Soceni Banat.
tubuliferum Goldfuss, 1837. - Combination to be
revised: "Cardium" tubuliferum Goldfuss, 1837.
?Cardiidae. Cretaceous. REcoRDs. --1871 Stoliczka, 60:
C. tubuliferum (Acanthocardium sic). "1922-32
Sherborn, p. 6677: Cardium tubuliferum Goldfuss,
1837. "1999 Jacques Vidal (pers. com.) MNHN, Paris:
Fossils from the Paleontological Collection. Cardium
tubuliferum Goldfuss. Senonien. Holland (Vaels),
B18337.
tubulosum Eichwald, 1830. - Combination to be revised:
"Cardium" tubulosum Eichwald, 1830. ?Cardiidae.
Fossil. REcoRDs. --1922-32 Sherborn, p. 6678: Cardium
tubulosum Eichwald, 1830.
tucani Pavlovic, 1929. - Changed combination:
Bosphoricardium tucani (Pavlovic, 1929). Lymnocardiidae. Pontian. RECORDS. --1928 ZR: Limnocardium
lucani [Ref. Pavlovitch 1928]. --1929 ZR:
Limnocardium lucani Pavlovic, 1929 sp. n. Pontian
Serbia. --1987 Taktakishvili: Limnocardium lucani
Pavl. - Bosphoricardium tucani.
Tulongocardiinae Schneider, 1995 (subfamily). RECORDS. --1995 Schneider: Tulongocardiinae
Schneider, 1995 subfam. n. Type genus: Tulongocardium Chen, Chen & Zhang in Lan & Chen, 1976. CHARACTERS AND OCCURRENCE - DESCRIPTIONS. --1995 Schneider: Cardiidae with shell sculpture
ofradial ribs on entirety of shell, shell quadrate, longer
than high, posterior margin crenulate. Spines absent.
Hinge as in Protocardia, but right anterior cardinal
smaller (absent in some specimens). OCCURRENCE Norian and Rhaetian ofIndonesia, Peoples' Republic
of China, northern Vietnam and Burma.
Tulongocardium Chen, J. Chen & Zhang, 1976 (genus).
- Cardiidae. Late Triassic. RECORDS. --1980 Vokes:
Chen, J. Chen & Zhang, 1976. Fossil. "1980 ZR:
Cardium (Tulongocardium) Chen, Chen & Zhang 1976.
Subgen. nov. Type species C. (T.) pluriradiatum,
Triassic. Ref. to Wen et al. p. 31. --1995 Schneider:
Tulongocardium Chen, Chen & Zhang in Lan and Chen,
1976. Type species Cardium (Tulongocardium)
pluriradiatum Chen, Chen & Zhang in Lan & Chen,
1976. --1995 Schneider: Tulongocardium is not a
cardiid genus according to Keen (1980).
tulskajensis (variety). - See under andrussovi var.
tulskajensis Pavlinova-Ilyina, 1961.
*tumidum. - Homonymy to be noticed. Cardium
tumidum from 3 sources: Kloeden, 1834. - Deshayes,
1854. - Ali-Zade & Ribina (?year).
tumidum Ali-Zade & Ribina. - Synonym of
Cerastoderma (lnaequicostala) politioanei (Jekelius).
» Regarding Cardium tumidum: Cardiidae. Fossil.
RECORDS ... 1977 ZR: Cardium (Replidacna) tumidum
Ali-Zade & Ribina. Ref to Paramo nova p. 70 [Transl.
p.320].
tumidum Deshayes, 1854. [Figs. Part 3 p. 871]. Synonym of Vasticardiumjlavumjlavum (Linne, 1758).
OTHER JUDGEMENTS: Cardium enode Sowerby. Acrosterigma (Vaslicardium) enode (Sowerby, 1841).
- Not Vasticardium enode. - Possibly A. (R.) gratiosum.
» Regarding Cardium tumidum: Cardiidae. Recent.
TYPE LOCALITY. Moluccas. TYPE REPOSITORY. Syntype
BMNH. REcORDS. --1855 Deshayes: Cardium tumidum.
--1884 Grasset: Cardium tumidum Desh. --1890 Paetel:
Cardium tumidum Dsh. --1980 Keen: Cardium
tumidum not a synonym of Vasticardium enode but it
may fall within the range of variation of A. (R.)
graliosum. --1980 Ruhoff: Cardium lumidum
Deshayes, 1854; p. 331. "1993 Vidal: Cardium
tumidum not synonym of Cardium enode as suggested
by Fischer-Piette 1977. --1997 Vidal: Cardium tumidum
Deshayes, 1854 = Vasticardiumjlavumjlavum (Linne,
818
Tropical Marine Mollusc Programme (TMMP)
tumidum - turislavicum
1758). ··1999 Higo et at.: Cardium tllmidum Deshayes,
1854 = Acrosterigma (Vasticardium) enode (Sowerby,
1841). ··1999 Vidal: Cardium tllmidum Deshayes, 1855
= Vasticardium jlavum (Linne, 1758). - CHARACTERS AND OCCURRENCE - REMARKS. ··1980 Keen:
referred to Acrosterigma (Regozara) sp. DISTRIBUTION.
··1884 Grasset: Australia. ··Of authors: Moluccas.
tumidum Kloeden, 1834. - Combination to be revised :
"Cardium " tumidum Kloeden, 1834. Cardiidae. Fossil.
RECORDS . ··1922-32 Sherborn , p. 6681 : Cardium
tumidum Kloeden, 1834.
tumidum Meuschen, 1778. Non binominal. - Not
available. RECORDS. ··1902 Sherborn, p. 1006: Cardium
tumidum Meuschen, 1778.
tumorifera (mode). - See under tumoriferum Lamarck,
1819.
tumoriferum Lamarck, 1819. (Figs. Part I p. 110, 135 .
Part 3 p. 910]. - Synonym of Fragum hemicardium
(Linne, 1758). OTHER JUDGEMENTS : A dubious species.
» Regarding Cardium tumoriferum: Cardiidae .
Recent. TYPE LOCALITY. (Australia, Bay of ChiensMarins) (New Holland in Sowerby 1841 No. 89). TY PE
REPOSITORY. Mus. no. and in Lamarck's cabinet;
availability unknown RECORDS. ··1819 Lamarck, p. Ili
No. 43: Cardium tumoriferum. ··1834 Sowerby, 50 t
part, fig. 22: Cardium tumoriferum Lam. ··1841
Lamarck in Delessert, pI. II, fig. 7 a,b,c: Cardium
tumoriferum. ··1840 Sowerby, catalogue p. 7, No. 89:
Cardium tumoriferum. ··1840 Sowerby, corrected list,
p. 8: Cardium tumoriferum (Sowerby No. 89). ··1844
Reeve, pI. 12, sp. 64: Cardium tumoriferum. ··1945
ZR: Cardium tumoriferum Lk. (Ref. Lamy 1945].
··1858 Adams & Adams, p. 459: Hemicardia (Fragum)
tumorifera Lam. ··1869 Romer: Cardium tumoriferum
Lamarck, a dubious species. ··1884 Grasset :
Hemicardium tumoriferum Lam. ··1890 Paetel :
Hemicardium tumoriferum Lm. ··1922-32 Sherborn,
p. 6683: Cardium tumoriferum Lamarck, 1819. ··1950
Allan: Hemicardium tumoriforum sic. ··1964 Shikama:
Corrolum (Lunulicardia) tumoriferum. ··1977 FischerPiette: Cardium tumoriferum Lamarck = Corcllium
hemicardium Linne. ··1977 Wilson & Stevenson :
Cardium tumoriferum Lamarck, 1819; junior synonym
of Fragum (Lunulicardia ) hemicardium (Linnaeus,
1758). ··1986 Abbott & Dance: Lunulicardia
lumorifera (Lamarck, 1819). ··1992 Lamprell &
Whitehead: Cardium lumoriferum Lamarck, 1819 =
Fragum (Lunulicardia) hemicardium (Linne, 1758).CHARACTERS AND OCCURRENCE - DESCRIPTIONS. ··1840 Sowerby No. 89: C. lumorifentm with
ribs narrower and tubercles smaller than in C.
hemicardium. ··1844 Reeve: shell cordate, swollen,
obliquely quadrilateral, posteriorly compressed, with
a prominent keel running down from the umbones to
the margins; radiately ribbed, ribs flattened, almost
obsolete, interstices closely punctured; anterior area
large, smooth; snowy white. ··1844 Reeve: at first
Reeve hesitated to consider this species anything more
than a variety of the Cardium hemicardium. But, he
found that it appears to differ in having the anterior
area perfectly smooth, whilst the ribs are much less
prominently developed. The name given to this shell
is suggested by the circumstance, which Lamarck
relates of its interior being often blistered, and which
is certainly very conspicuous in the example Reeve
studied. DISTRIBUTION. ··1844 Reeve: New Holland =
Australia. ··1884 Grasset: Australia. ··1994 Hyl\eberg:
Red Sea, West Pacific.
tumoriferum of Reeve. - Suggested misidentification.
··1868 ZR: Cardium tumoriferum of Reeve is not that
of Lamarck (Ref. Romer 1868].
tumoriforum (error). - Printer ' s mistake for
lumoriferum inAJlan 1950 and in Shikama 1964.
tunicula Vokes, 1977. - Fragum tunicula Vokes, 1977.
Cardiidae. Oligocene. RECORDS. ··1977 Vokes, p. 172,
pI. 6, fig. 5, pl. 7, figs. 1-3 : Fragum tunicula H. E.
Vokes n. sp. ··1977 ZR: Fragum lunicula Vokes, 1977
sp. nov. Florida. Ref to Vokes p. 172, fig'd .
tuomeyi Aldrich. - Acanthocardia tuomeyi (Aldrich).
Cardiidae . Fossil. RECORDS. ··1977 Popov:
Acanlhocardia (Schedocardia) tuomeyi (Aldrich).
turbida (variety). - See under subsyrmiense var. turbida
Eberzin, 1958.
turcmena Andrussow. - Hyrcania turcmena
(Andrussow). Lymnocardiidae. Fossil. RECORDS. ··1977
Popov: Hyrcania (Hyrcania) lurcmena (Andrussow).
turfanensis Martinson, 1959. - Pseudocardinia
turfan ensis Martinson, 1959. Cardiacea. Jurassic.
RECORDS. ··1959 ZR: Pseudocardinia turfanensis
Martinson, 1959 sp. nov. Turfan Depression, Sinkiang,
Western China, Jurassic.
*turgidum. - Homonymy to be noticed. Cardium
turgidum from 4 sources: Solander in Brander, 1766.
Brand, 1837. Brown, 1845-49. Deshayes, 1854.
turgidum Brand, 1837. - Synonym of "Cardium"
comalufum Bronn, 1863. Combination to be revised .
Cardiidae. M.-U. Oligocene. RECORDS. ··1837 Brand,
p. 222, Tav. 145, fig. 3: Cardium turgidum (fide Ravn
1907). ··1907 Ravn: Cardium lurgidum Brand, 1837 =
Cardium comalulum Bronn, 1863 .
turgidum Brown, 1845-49. - Combination to be revised:
"Cardium " lurgidum Brown, 1845-49. ?Cardiidae.
Fossil. RECORDS. ··1922-32 Sherborn, p. 6691 : Cardium
turgidum Brown, 1845-49.
turgidum Deshayes, 1854. - Not a cardiid. » Regarding
Cardium lurgidum : printer's error. Should read
Caecella turgida . RECORDS. "1890 Paetel : Cardium
lurgidum Dsh. ··1903 Hidalgo : Cardium lurgidum in
Deshayes 1854; printer 's error = Caecella lurgida
Deshayes . ··1928 Faustino: Cardium turgidum
Deshayes. ··1977 Fischer-Piette: erroneous citation of
Caecella turgida Deshayes .
turgidum Solander in Brander, 1766. - Changed
combination : Nemocardium lurgidum (So lander in
Brander, 1766). Cardiidae. Fossil. RECORDS. ··1902
Sherborn, p. 1007: Cardium lurgidum So lander in
Brander, 1766 . ··1977 Popov: Nemocardium
(Nemocardium) turgidum (Solander, 1766). ··2001
Schneider & Carter: Nemocardium turgidum (Sol ander,
1766). Cardiidae. Eocene. England.
turislavicum Jek. - New combination: Lymnocardium
turislavicum (Jek.). Lymnocardiidae. Fossil. RECORDS.
··1987 Taktakishvili : Limnocardium turislavicum Jek.
turislavicum Jekelius , 1944. - New combination.
Lymnocardium turislavicum Jekelius, 1944. Lymnocardiidae . Miocene . RE CORDS . ··1946 ZR:
Limnocardium turislavicum Jekelius, 1944 sp. nov.
Miocene Soceni Banat.
Phuket Marine Biological Center Special Publication vol. 30 (2004)
819
Tllrkmena - tOrteliense
Turkmena Popov, 1956 (genus). - Lymnocardiidae.
Tertiary and Recent. REcORDs. ··1980. Vokes: Turkmena
Popov, 1956 in subfamily Adacninae.
turkmena Andrussov, 1924 . - Didaena turkmena
Andrussov, 1924. Lymnocardiidae. Pliocene and
Pleistocene. ··1924 ZR: Didaena turkmena Andrussov,
1924 sp. n. Pleistocene. ··1929 ZR: Didaena turkmena
Apscheronian Transcaucasia [Ref. Ruzencev I 929b].
··1963 ZR: Didaena turkm ena Andrusov, 1963 sp. nov.
Apsheron layers, Pliocene, Russia. ··1963 ZR: Didaena
turkmena Pliocene [Ref. Andrusov 1963].
turkmena latecostata Ruzencev, 1929 (variety). Upgraded. Didaena turkmena lateeostata Ruzencev,
1929. Lymnocardiidae. Apscheronian. RECORDs. ··1929
ZR: Didaena turkmena var.latecos/ata Ruzencev, 1929
nov. Apscheronian Transcaucasia [Ref. Ruzencev
I 929b].
turkmena major Andrusov, 1963 (variety). - Upgraded.
Didaena turkmena major Andrusov, 1963.
Lymnocardiidae. Pliocene. RECORDS. "1963 ZR:
Didaena turkmena var. major Andrusov, 1963 nov.
Apsheron layers, Pliocene, Russia.
turonense Cossmann, Peyrot, 1911. - Synonym. »
Regarding C. turonense: Cardiidae. Miocene. RECORDS.
··1989 Gontsharova: Cardium turonense Cossmann,
Peyrot, 1911 = Aean/hoeardia grundensis Ivolas et
Peyrot, 1900.
turonense Mayer emend. Synonym. Cardiidae. Miocene.
RECORDS. "1902 Dollfus & Dautzenberg: Cardium
turonense May. emend. Cossmann & Peyrot, 1912.
Synonym of Cardium turonieum Mayer in Hoemes,
1863.
tllronica (mode). - See under turonieum.
*tllronicllm. - Homonymy to be noticed. Cardium
turonieum from 3 sources: Homes, 1861. - Mayer in
Hoemes, 1863. - Possibly Mayer, 1870.
turonicum Homes, 1861. - Synonym of Cardium
sallomaeense Cossmann & Peyrot 1912 . OTHER
JUDGEMENTS: Cardium broeehi Mayer, 1866. - Cardium
praeeehinatum Hilber, 1882. » Regarding Cardium
turonieum : Cardiidae. Miocene. TYPE LOCALITY. Europe.
RECORDS. ··1986 Studencka, p. 69 : Aeanthoeardia
(Acanthoeardia) turoniea (Homes 1861); syn. nov. =
Cardium sallomaeense Cossmann & Peyrot 1912.
··1987 ZR: Aeanthoeardia (Aeanthocardia) turoniea
(Homes, 1861). New synonyms: Cardium broeehi
Mayer, 1866, Cardium praeeehinatum Hilber, 1882,
Cardium sallomaeense Cossmann & Peyrot, 1912. Ref.
to Studencka p. 69.
tllronicum Mayer in Hoemes, 1863 I Mayer, 1870. Aeanthoeardia turoniea (Mayer, 1870). Cardiidae.
Tertiary. RECORDS . ··1886 Dollfus & Dautzenberg:
Cardium turonieum Mayer. ··1952 Lecointre: Cardium
turonieum Mayer in Hoemes, 1863 . "1977 Popov:
Aeanthoeardia (Aeanthoeardia) turoniea (Mayer,
1870). ··1999 Jacques Vidal (pers. com.) MNHN, Paris:
Fossils from the Paleontological Collection. Cardium
(Aeanthoeardia) turonieum Mayer. Helvetien. France;
Austria ; Rumania; Hungary; Turkey; former
Yugoslavia.
tllronicllm grllndense Ivolas & Peyrot, 1900 (variety).
- Changed combination: Aeanthoeardia turonieum
grundense (Ivolas & Peyrot, 1900). Cardiidae. Fossil.
RECORDS.
··1989
Gontsharova :
Cardium
(Aeanthoeardia) turonieum grundense in Sieber, 1956
Acanthoeardia grrmdensis Ivolas et Peyrot, 1900.
··1989 Gontsharova : Cardium turonieum var.
grundensis Ivolas et Peyrot, 1900 = Aeanthoeardia
grundensis Ivolas et Peyrot, 1900.
tIIroniCllm vidali Cossmann et Peyrot (variety). Upgraded. New combination: Aeanthoeardia turoniea
vidali Cossmann & Peyrot. "1912 Favre: Cardium
turonieum Mayer var. vida Ii Cos sm. et Peyr. ID
suggested by Favre.
turoniense H. Woods. - Changed combination:
Pleurioeardia turoniense (H . Woods). Cardiidae.
Cretaceous. REcORDs. ··1978 Scott: Cardium turoniense
H. Woods. ··1978 Scott: Pleurioeardia (Doehmocardia) turoniense (H. Woods). ··1999 Jacques Vidal
(pers . com.) MNHN, Paris : Fossils from the
Paleontological Collection. Cardium turoniense Woods.
Turonien. France (du Rhone), B 17570.
turtoni (variety). - See under spinosum var. turtoni Dall.
turtoni Sowerby, 1894. - Synonym of Parvieardium
papillosum (Po Ii, 1791). » Regarding Cardium
turtoni: Cardiidae. Recent. DISTRIBUTION. South Africa
(Sowerby 1894). RECORDS. ··1894 Sowerby, p. 377378, pI. vii, fig . 4: Cardium tllrtoni Sowerby, n. sp.
··1894 ZR: Cardium turtoni Sowerby, 1894a n. sp. Port
Elizabeth. ··1897 Sowerby: Cardium turtoni. ··1904
Smith: Cardium turtoni Sowb. "1926 McClelland:
Cardium turtoni, Sowerby, 1894 n. sp. ··1950 Barnard:
Cardium turtoni . ··1974 Barnard: Cardium turtoni
Sowerby. ··1977 Fischer-Piette: Cardium turtoni
Sowerby = Coreulum papillosum Poli. ··1982 Kilburn
& Rippey: Parvieardium turtoni (Sowerby, 1894).
··1991 Voskuil & Onverwagt: Cardium turtoni
Sowerby, 1894 = Parvieardium papillosum (Poli,
1795).
tllteheri Palmer, 1973. - Cryptoeardia(?) tutcheri
Palmer, 1973 . ?Cardiidae. Lower Jurassic. REcORDs.
··1973 ZR: Cryptoeardia(?) tuteheri Palmer, 1973 sp.
nov. England, Somerset, Ilminster. Ref. to Palmer p.
168.
Tllteheria (genus) . - Not valid for Cardiacea. Fossil.
RECORDS. ··1946 ZR: Tuteheria Cox, 1946 gen. nov.
Cardiidae type Cardium submultieostatum Orb. ··1995
Schneider: Tuteheria is an astartid genus.
tllz/anum (subspecies). - See under lithopodolicum
tuzlanum Soklic, 1958.
Tychoeardia [Fig. Part 1, p. 163] (section & subgenus).
- Now in family Isocardiidae. RECORDs. ··1869 Ed.
Romer: I. Sectio Tyehoeardia Romer. Genus [socardia
Lamarck. ··1869 ZR: subgenus I. Tychoeardia, type
[soeardia cor (L.) [Ref. Romer 1869].
typica (variety). - See under isseli var. typiea Androussov,
1923. - propinqua var. typiea Androussov, 1923.
tzankovi Kojumdgieva, 1970. - Combination to be
revised: "Cardium" tzankovi Kojumdgieva, 1970.
Cardiidae. Upper Miocene. RECORDS. "1970 ZR:
Cardium (Cerastoderma) tzankovi Kojumdgieva, 1970
sp. nov. Sarmatian, Upper Miocene, Gaitantzi ,
Vidinsko, Bulgaria.
torteliense (invalid spelling). - See under toerteliense.
=
820
Tropical Marine Mollusc Programme (TMMP)
uaylandi - undatopictum
U
uaylandi Woodring, 1925.- Acrosterigma uaylandi
(Woodring, 1925). Cardiidae. Fossil. RECORDS. ··1977
Popov: Acrosterigma uaylandi (Woodring, 1925).
uchidai Habe, 1955. - Synonym of Clinocardium
californiense (Deshayes, 1839). » Regarding
Clinocardium uchidai: Cardiidae. Recent. RECORDS.
··1954 Kira: Clinocardium uchidai Habe, MS. ··1955
Habe, p. 11-12, pI. 2, figs. 5-6: Clinocardium uchidai
sp. nov. "1955 Habe: Clinocardium californiense in
Rolle 1896, p. 114, fig. b, non Deshayes = Clinocardium
uchidai sp. nov. "1955 ZR: Clinocardium uchidai
Habe, 1955 sp. nov Akkeshi Bay, Japan. ··1962 Kira:
Clinocardium uchidai. ··1973 Keen : Clinocardium
uchidai Kira, 1954, ex Habe, MS . Recent. ··1977
Fischer-Piette: Corculum (Keenocardium) uchidai
Habe. ··1977 Habe: Clinocardium uchidai Habe, 1955
= Clinocardium (Keenocardium) californiense
(Oeshayes, 1839). ··1980 Keen : Clinocardium uchidai
= Clinocardium californiense (Deshayes, 1839). ··1980
ZR: Clinocardium uchidai Kira 1954 Nomen nudum .
Ref. to Kafanov, p. 310 ... 1999 Higo et al. :
Clinocardium uchidai Habe, 1955 = Clinocardium
(Keenocardium) californiense (Deshayes, 1839) .
"2000 Coan et al.: Clinocardium uchidai Habe, 1955
= Clinocardium (Keenocardium) californiense
(Deshayes, 1839). - CHARACTERS AND OCCURRENCE - REMARKS. ··1980 Keen: probable that
Laevicardium interrogatorium Fischer-Piette, 1977 =
Clinocardium uchidai (Habe , 1955) from the
northwestern Pacific (not California) . DISTRIBUTION.
··1994 Hylleberg: Japan.
uiratamense Kolesnikov, 1929. - Combination to be
revised: "Cardium" uiratamense Kolesnikov, 1929.
Cardiacea. Sarrnatian. RECORDS. ··1929 ZR: Cardium
uiratamense Kolesnikov, 1929 sp. n. Sarrnatian.
uiratamense inepta Kolesnikov, 1929 (variety). Upgraded . Combination to be revised : "Cardium "
uiratamense inepta Kolesnikov, 1929. Cardiacea .
Sarrnatian. RECORDS. ··1929 ZR: Cardium uiratamense
var. inepta Kolesnikov, 1929 nov. Sarrnatian [Ref.
Kolesnikov 1929].
ukrainica Ebersin, 1967. - Macradacna ukrainica
Ebersin, 1967. Lymnocardiidae. Fossil. RECORDS.
··1967 Ebersin: Macradacna ukrainica sp. n. ··1967
ZR: Macradacna ukrainica Eberzin, 1967 sp. nov.
Lymnocardiidae. Pleistocene. TYPE LOCALITY. Russian
SFSR. RECORDS. "1983 Popov, p. 160, fig.: Didacna
ultima sp. nov.
IImbonare Lamarck, 1819. - Combination to be revised:
"Cardium " umbonare Lamarck, 1819. Cardiacea .
FossiL RECORDS. ··1819 Lamarck: Cardium umbonare
Lamk ... 1922-32 Sherborn, p. 6715: Cardium
umbonare Lamarck, 1819.
umbonata Ebersin, 1959. - Didacna umbonata Ebersin,
1959. Lymnocardiidae. Quaternary. RECORDS. "1957
ZR: Didacna umbonata Ebersin. Nizhnekhvalin
deposits, Chelekan and Pribal Khan. Quaternary,
Turkmenia, Russia [Ref. Neveskaya 1957]. ··1959 ZR:
Didacna umbonata Ebersin, 1959 sp. nov. fossil.
umbonatum (subspecies) . - See under glaucum
umbonatum Wood, 1850.
*umbonatum. - Homonymy to be noticed. Cardium
umbonatum from 5 sources: Sowerby, 1817. - Bell,
1913. - Wood, 1850. - Cossmann, 1921. - Goldfuss
(?year).
IImbonatum Bell, 1913 . -Combination to be revised :
"Cardium " umbonatum Bell, 1913. Cardiidae. Crag.
RECORDS. ··1913 ZR : Cardium umbonatum Bell, 1913
sp. n. Suffolk Crag.
IImbonatlim Cossmann, 1921 . - Fragum umbonatum
(Cossmann, 1921). Cardiidae. Eocene. RECORDS. ··1922
ZR: Cardium (Trachycardium) umbonatum Cossmann,
1922 sp. n. Eocene. "1930 Stewart: "c." umbonatum
Cossmann, 1921. ··1977 Popov: Fragum umbonatum.
IImbonatlim Goldfuss. - Replaced by "Cardium"
austriacum Desh. » Regarding C. umbonatum:
Cardiidae. Fossil. RECORDS. "1857. Oeshayes: Cardium
umbonatum. Goldf. - C. austriacum . Desh.
umbonatum Sowerby, 1817. - Not a cardiid. RECORDS.
··1857 Deshayes: Cardium umbonatum Sowerby. In
genus Glycymeris. "1922-32 Sherborn, p . 6715 :
Cardium umbonatum Sowerby, 1817.
umbonatum Wood, 1850. - Changed combination:
Cerastoderma umbonatum (Wood, 1850). Cardiidae.
ukrainica Eberzin & Semenenko, 1967. - Limnodacna
ukrainica Eberzin & Semenenko, 1967 .
Fossil. TYPE LOCALITY. Caspian Sea. DISTRJBUTION. North
Africa, Adriatic Sea (Nordsieck 1969). RECORDS. ··1850
Wood: Cardium umbonatum. "1974 Starobogatov:
Cerastoderma umbonatum (Wood, 1850).
umbricatum (error). - Printer's mistake for imbricatum
in Higo et al. 1999.
umkae Stevanovic, 1982. Paradacna umkae Stevanovic,
1982. Lymnocardiidae. Upper Miocene. RECORDS.
··1982 ZR: Paradacna umkae Stevanovic, 1982 sp.
nov. Yugoslavia. Ref. to Stevanovic p. 126.
undatopicta (mode). - See under undatopictum Pilsbry,
Lymnocardiidae. Lower Kimmeridgian. RECORDS.
··1967 ZR : Limnodacna ukrainica Eberzin &
Semenenko, 1967 sp. nov. Ukraine, USSR. ··1987
Taktakishvili: Limnodacna ukrainica Ebers. et Sem.
ukrainicum (variety). - See under mithridatis
ukrainicum var. Ossaulenko, 1936.
ukrainicum Bagdasaryan, 1979. - Sarmaticardium
ukrainicum Bagdasaryan, 1979. Cardiacea. Upper
Miocene . RECORDS. "1979 ZR: Sarmaticardium
ukrainicum Bagdasaryan, 1979 sp. nov. Ukraine. Ref.
to Bagdasaryan, p. 42.
ultima (variety). - See under inlongaevum ultima var.
Eberzin, 1958.
ultima Popov, 1983.- Didacna ultima Popov, 1983.
1904.
undatopictum Pilsbry, 1904. - Changed combination:
Fulvia undatopictum (Pilsbry, 1904). OTHER
JUDGEMENTS: Fulvia hungerfordi (Sowerby, 1901). Laevicardium hungerfordi undatopictum (Pilsbry,
1904). - Papyridea papyraceum (Chemnitz) Schrcter
[not available] . » Regarding Cardium hungerfordi
undatopictum: Cardiidae. Recent. TYPE LOCALITY.
Hirado, Nagasaki Pref. Kyushu (Kuroda et al. 1971).
RECORDS. "1904 Pilsbry, p. 556, pI. 40, figs . 14-15:
Cardium hungerfordi undatopictum . "1940 Nomura
& Niino: Cardium undatopictum Pilsbry. ··1951 Habe:
Microfragum undatopictum Pilsbry. ··1952 Kuroda &
Habe: Fulvia undatopicta (Pilsbry). ··1954 Taki:
Pliocene, USSR.
Phuket Marine Biological Center Special Publication vol. 30 (2 004)
821
undatum - unedo
Cardium undalopietum Pilsbry. "1954 Kira :
Laevieardium undalopietum (Pilsbry). --1955 Kira :
Laevieardium undatopietum (Pilsbry). --1958 Habe :
Laevieardium undatopietum (Pilsbry). --1962 Kira :
Laevieardium undatopietum (Pilsbry) . --1971 Kuroda ,
Habe & Oyama: Laevieardium undatopielum (Pilsbry).
--1977 Popov : Laevieardium undatopietum (Pilsbry).
--1977 Fischer-Piette: Cardium undatopietum Pilsbry
= Papyridea papyraeeum (Chemnitz) Schrater. --1977
Habe: Laevieardium undatopietum (Pilsbry, 1904).
--1981 Koyama et al.: Fulvia undatopietum. --1981
Koyama et al.: Laevieardium hungerfordi
undatopietum (Pilsbry, 1904) = Fulvia hungerfordi
(Sowerby, 1901). --1994 Vidal: Fulvia (Laevifulvia)
undatopiela (Pilsbry, 1904). --1994 Vidal: Fulvia
undatopieta (Pilsbry). Type of(Laevifulvia) Vidal, 1994
new subgen. --1998 Lamprell & Healy: Fulvia (Fulvia)
undatopiela (Pilsbry, 1904). --1999 Higo et al.: Fulvia
undatopieta (Pilsbry, 1904). --2000 Hylleberg, p. 532:
Fulvia undalopiela (Pilsbry, 1904). Det. 1. Vidal. "2000
Lutaenko : Fulvia undatopiela (Pilsbry, 1904). --2000
Lutaenko, p. 372: Fulvia undatopieta (Pilsbry, 1904).
--2001 Vidal, p. 408 : Fulvia undatopieta. --2003
HyUeberg & Kilburn, p . 183 : Fulvia undatopieta
(Pilsbry, 1904). - CHARACTERS AND OCCURRENCE - DESCRIPTIONS . --200 I Vidal : tentacles
numerous. Eyes on tip of tentacles. DIMENSIONS. -'1971
Kuroda et al.: 17.1 x 17.4 x 10.8 mm. HABITAT. --1971
Kuroda et al. : fine sandy, gravely bottoms, 10-100 m
depth . --1999 Higo et al.: subtidal to 100 m, sandy mud
DISTRIBUTION. --1971 Kuroda et al. : Japan, Honshu,
Shikoku, Kyushu . --1994 Hylleberg: Tropical Pacific,
Solomon lsI. --1999 Higo et at.: Boso peninsula
(Kuroda & Habe, 1981) and southwards, Seto island
sea, Japan Sea (Oga peninsula), Philippines, South
China Sea, Indo-Pacific, Australia (Gulf of Carpentaria
Lamprell & Healy 1998). --2000 Hyl\eberg: Vietnam
and Cambodia. --2003 Hylleberg & Kilburn: Vietnam.
*undawm. - Homonymy to be noticed. Cardium
undatum from 2 sources: Sismonda, 1846. - Reuss,
1868.
undatum Reuss, 1868. - Combination to be revised:
"Cardium " unda/um Reuss, 1868. ?Cardiidae. Fossil.
RECORDS. --1980 Ruhoff: Cardium undatum Reuss,
1868; vol. 58 : 87, pI. 2, figs. 3-4.
undatum Sismonda, 1846. - Combination to be revised:
"Cardium " undatum Sismonda, 1846. ?Cardiidae.
Fossil. RECORDS. --1922-32 Sherborn, p. 6726: Cardium
undatum Sismonda, 1846.
undatum Martyn, 1784. Non binominal. - Not available.
Not a cardiid . RECORDS. --1784 Martyn : Cardium
undalum.
unedo Linne, 1758 / Gmelin, 1791. [Figs. Part 1 p. 39,
40,43,46,50, 60, 61 , 68, 73,87, 103 , 125, 166.Part2
inside of back cover. Part 3 p. 905, 907] . - Changed
combination: Fragum unedo (Linne, 1758). Cardiidae.
Recent. Gray (1847, No. 557) Cardium unedo is type
species of Hemieardium, Swainson 1840. DaJl (1902)
Cardium unedo Linnaeus is type of subgenus Fragum
s.s. TYPE LOCALITY. Ceylon (Sowerby 1841 No. 86) .
None designated (Higo et al. 1999). RECORDS. --1684
Bonanno. ID by Gmelin 1791 : Cardium unedo recr. 3.
f. 375 . --1687 Lister. ID by Gmelin 1791: Cardium
unedo t. 315. f. 151 . --1705 Rumphius. ID by Linnaeus
1758 and Gmelin 1791: Cardium unedo mus. t. 44 . f.
F. --1742 Argenville. 1D by Linnaeus 1758 & Gmelin
1791 : Cardium unedo conch. 1. 26. f. N. --1742
Gualtieri. ID by Gmelin 1791: Cardium unedo 1. 83 . f.
A . --1758 Linne, p . 680, No. 68: Cardium unedo.
--1758 Regenfuss. ID by Gmelin 1791: Cardium unedo
Conch. 1. 1. 3. f. 25 . --1758 Regenfuss, fig. 25: Cardium
unedo (fide Linne 1758; Spengler 1799). --1758 Seba.
ID by Gmelin 1791: Cardium unedo mus. 3. t. 86. f.
12. --1758 Seba, Vol. 3, pI. 86, fig. 12: Cardium unedo
(fide Spengler 1799). --1764 Knorr. ID by Gmelin 1791:
Cardium unedo Vergn. 2. 1. 29. f. 2. --1764 Linnaeus
(Mus. Lud . Uir. Reg.) No. 40. --1767 Linne, p. 1123,
No. 84: Cardium unedo. --1769 Knorr (author not
available) : C. unedo . --1778 Born, G 10. Cardium unedo
Linn. S.N. Sp. 84. --1780 Born: Cardium unedo Linn.
S. N . Sp. 84. --1782 Bonannio. ID by Gmelin 1791:
Cardium unedo mus. Kirch. 2. f. 109. --1782 Chemnitz,
pI. 16, figs . 168-169: Cardium unedo. "1782 Chernnitz.
ID by Gmelin 1791 : Cardium unedo t. 16. f. 168. 169.
--1786 Schrater, No. XII: Cardium unedo Linn. XII.
pag. 1123 . Spec. 84. --1789 Bruguiere, No.7. Cardium
unedo Linn . --1789 Karsten : Cardium unedo testa
solida sulcata . "1791 Gmelin , p . 3250 , No. 14:
Cardium unedo. --1798 Bruguiere. ID by Lamarck, pI.
295 . fig. 4 . Cardium unedo. --1798 Rading: Fragum
unedo. --1799 Spengler, p. 59, No. 40: Cardium unedo
Linnei Sp. 84 . p. 1123. --1817 Dillwyn: Cardium unedo
Linnaeus . Born appears to have confounded C. ringens
with C. unedo. --1823 Dillwyn, Tab. 315 fig. 150
Cardium unedo. --1836 Bose: Cardium unedo. --1840
Sowerby, catalogue p. 7, No. 86: Cardium unedo Linn.
--1844 Potiez & Michaud : I . Cardium unedo Linn.
Gmel. --1844 Reeve, sp. 13: Cardium unedo. --1847
Gray, No . 557 : C. unedo . "1853 Gray: Bueardium
unedo. --1853 March: Fragum unedo. --1856 Wood &
Hanley, No. 50: Cardium unedo Lin. Syst. --1858
Adams & Adams, p. 459: Hemicardia (Fragum) unedo
Linn. --1869 ZR: Section, subtypical - 4.
Semicordiformes: fragum (L.), unedo (L.),
donaeiformis (Spengl.), &e. [Ref. Graham-Ponton
1869]. --1880 Martens: Cardium (Hemieardium) unedo
Linne. --1884 Grasset: Hemieardium unedo. L. --1887
Fischer: Hemieardium unedo Linne. --1889 Martens:
Cardium (Hemieardium) unedo L. , citing Chernnitz.
--1890 Paetel : Hemieardium unedo L. --1891 Fischer:
Cardium unedo Linne. --1891 Ramer: Fragum unedo
L. --1902 Dall : Cardium unedo Linnaeus . --1902
Sherborn, p. 1016: Cardium unedo Linnaeus, 1758.
--1905 Hidalgo : Hemieardium unedo Linne. --1895
Melvill & Standen : Hemieardium (Fragum) unedo
Linne. --1909 Lynge: Hemieardium (Fragum) unedo .
--1939 Adam & Leloup: Coreulum (Fragum) unedo
(Linne 1758). --1977 Wilson & Stevenson : Fragum
(Fragum) unedo (Linnaeus, 1758). --1932 Prashad:
Cardium (Fragum) unedo. --1945 ZR: Cardium unedo
L. [Ref. Lamy 1945]. --1960 Kuroda : Fragum unedo
(Linne). --1967 ZR: Fragum unedo, effect ofmyotropic
substances on the heart [Ref. Kratzing & Ladd 1967].
--1977 Fischer-Piette: Coreulum (Fragum) unedo
Linne. --1977 Wilson & Stevenson: Fragum (Fragum)
unedo (Linnaeus, 1758). Hemieardium unedo Linnaeus,
1758. --1990 Abbott & Dance: Fragum unedo. --1994
Chung-Hung & Hsi-Jen: Fragum unedo (Linne).
822
Tropical Marine Mollusc Programme (TMMP)
unedo - unicolor
··1995 Chung-Hung & Hsi-Jen: Fragum unedo
(Lilme) . ··1981 ZR: Fragum unedo Linnaeus Senior
synonym of F. tegulatum (Dautzenberg 1900). Ref. to
Germain, p. 166. ··1982 ZR: Fragum (Fragum) unedo
(Linnaeus 1758). Indian Ocean and Western Pacific.
Ref. to Germain. ··1983 ZR: Fragum unedo. Japan.
Ref. to Kawaguti. ··1985 ZR: Fragum unedo. Ref. to
Ohno. ··1998 Poutiers: Fragum unedo (Linnaeus,
1758) . ··1998 Schneider (l998b): Fragum unedo
(Linne,
1758) . Harbours photosymbiotic
dinoflagellates. ··1999 Aungtonya e/ al. , PMBC-SP
19(2): 377: Fragum unedo (Linnaeus, (758). ··1999
Higo el al.: Fragum unedo (Linnaeus, 1758). ··1999
Hylleberg, p. 40 I: Fragum unedo (Linne, 1758). ··1999
Knudsen & Hylleberg: Spengler, 1799 No. 40: Cardium
unedo in Spengler (East Indies) = Fragum rubrum
citing Chemnitz Tom. VI. Tab. 16 Fig. 168-169. ··2000
Hylleberg, p. 532: Fragum unedo (Linne, 1758··2000
Lutaenko, p. 372: Fragum unedo (L., 1758). ··200 I
Vidal, p. 408: Fragum unedo. ··2003 Hylleberg &
Kilburn, p. 178: Fragum unedo (Linnaeus, 1758). CHARACTERS AND OCCURRENCE - OF COLLECTIONS. ··ZMUC: Nicobar Islands, Fiji Islands, Japan,
Philippines. ··ZMUC 2 specimens, Spengler coil. East
Indies. ··ZMUC wet collection: Eilat, Lontor Banda,
Tonga Islands intertidal, Banda , Kei Islands .
DESCRIPTIONS . ··1758 Linne: Shell subcordate:
interstices crescent-shaped coloured [I have translated
from Latin]. ··1799 Spengler: Testa subcordata, sulcis
lunulis colora/is. ··1844 Reeve: shell somewhat heartshaped, posterior side angulated; radiately ribbed, ribs
about thirty in number, close-set, interstices on the
posterior side slightly open at the margins; yellowish,
ribs ornamented with bright pinkish-red, elevated cross
ridges or small shields. The bright red ornamental
sculpture of the ribs varies in its character and size from
minute cross ridges to small oblong square elevated
shields. ··200 I Vidal : only the exhalant aperture is tubeshaped. Inhalant siphon open towards the foot aperture.
No eyes. ··My rib counts: Material at ZMUC: 27,27,
27, 28 , 28, 28, 29 ribs. DIMENSIONS. ··1799 Spengler:
length 61 mm (2 Tom. 4 Lin .), width 52 mm (2 Tom.).
··1844 Reeve: Shell somewhat heart-shaped, posterior
side angulated; radiately ribbed, about 30 close-set ribs,
interstices on the posterior side slightly open at the
margins; yellowish, ribs ornamented with bright
pinkish-red, elevated cross ridges or small shields.
··1972 Cernohorsky: 60 mm. REMARKS. ··1844 Reeve:
The bright red ornamental sculpture of the ribs varies
in its character and size from minute cross ridges to
small oblong square elevated shields, as in the specimen
selected for illustration. HABITAT. ··1844 Reeve: found
in sandy mud at low water. ··1999 Higo et al.: intertidal
to 60 m, sand and mud. ··ZMUC: intertidal coral sand.
D ISTRIBUTION. ··1844 Reeve : Ceylon, Philippine
Islands, &C. ··1856 Wood & Hanley: Ceylon, &c.
··1884 Grasset: Ceylon. ··1891 Romer: East Indies.
··1994 Hylleberg: Indo-Pacific. ··1999 Aungtonya e/
al.: Thailand, Andaman Sea. ··1999 Higo et al.: Tosa
bay (southern Shikoku) and southwards, Philippines,
South China Sea, Beibu gulf, Indonesia, Indo-Pacific.
··2000 Hylleberg: Vietnam and Cambodia. ··2003
Hylleberg & Kilburn: Vietnam. ··ZMUC: Nicobar
Islands, Fiji Islands, Japan, Philippines, East Indies,
Banda, Tonga Islands, Kei Islands.
unedo Roding in Bolten, 1798. - The same as Fragum
unedo (Linnaeus, 1758). ··1902 Sherborn, p. 1016:
Fragum (Gm.) unedo Bolten, 1798. - Cardium unedo.
unedo Swainson, 1840. - The same as Fragum unedo
(Linnaeus, 1758). RECORDS. ··1922-32 Sherborn, p.
6738: Hemicardium (Wood) unedo Swainson, 1840.Cardium unedo.
unedo beta Gmelin, 1791 (variety). - Unknown status.
RECORDS. ··1764 Knorr. ID by Gmelin 1791 : Cardium
unedo var. beta, Vergn. 2. t. 29 . f. 3? ··1791 Gmelin
No. 14: Cardium unedo var. be/a.
ungheniensis. - Combination to be revised: "Cardium"
ungheniensis. Cardiacea. Fossil. RECORDS. ··1930 ZR:
Cardium ungheniensis, fossil [Ref. Vascautanu 1930].
ungulata Roding, 1798. - Synonym of Hippopus
hippopus (Linnaeus, 1758). » Regarding Tridacnes
ungulata: Tridacnidae. Recent. RECORDS. ··1999 Higo
et at.: Tridacnes ungulata Roding, 1798 = Hippopus
hippopus (Linnaeus, 1758).
Unicardium d'Orbigny, 1850 (genus). - Cardiacea.
Fossil. RECORDS. ··1922-32 Sherborn, p. 6742:
Unicardium d 'Orbigny, 1850. ··1924 ZR: Unicardium
[Ref. Cossmann 1924, p. 47] .
unicolor Sowerby, 1840. [Figs. Part I p. III, 115, 141.
Part 3 p. 881]. - Synonym of Acroslerigma simplex
(Spengler, 1799). OTHER JUDGEMENTS: Cardium enode
Sowerby. - Not a synonym of Cardium enode. »
Regarding Cardium unicolor: Cardiidae. Neogene and
Recent. TYPE LOCALITY. lsI. of Ticao, the Philippines
(Sowerby, 1840). TYPE REPOSITORY. Natural History
Museum, London. TYPE MATERIAL. Holotype missing.
Six paired syntypes from Ticao (Philippines), in
BMNH, Cuming coil., the largest of which fits the
specimen cited and figured by Sowerby, and was
selected as lectotype by Voskuil & Onherwagt (1991:
116). RECORDS. ··1840 Sowerby, 178 t part, fig . 42:
Cardium unicolor nobis . ··1840 Sowerby, p. 107:
Cardium unicolor. ··1840 Sowerby catalogue p. 4, No.
46 : Cardium unicolor Sow. ··1840 Sowerby, corrected
list, p. 8: Cardium unicolor (Sowerby No. 46) . ··1845
Reeve: Cardium unicolor. ··1858 Adams & Adams, p.
455: Cardium (Trachycardium) unicolor Sow. ··1884
Grasset: Cardium unicolor Sow ... 1890 Paetel:
Cardium unicolor Sow. ··1907 ZR: Cardium
(Trachycardium) unicolor raised beach, Red Sea [Ref.
Hall & Standen 1907]. ··1922-32 Sherborn, p. 6744:
Cardium unicolor Sowerby, 1841. "1928 Faustino:
Cardium unicolor Sowerby. ··1931 ZR : Cardium
unicolor with ref. to WD. Smith (1931). ··1932
Prashad: Cardium (Trachycardium) unicolor. ··1960
Kuroda: Trachycardium (Acrosterigma) unicolor
(Sowerby). ··1964 Spry: Cardium (Vasticardium)
(Regozara) unicolor (Sowerby) . ··1974 Habe &
Kosuge : Vasticardium unicolor (Sowerby, 1840).
··1977 Fischer-Piette: Cardium unicolor Sowerby =
Cardium enode Sowerby. ··1977 Habe: Trachycardium
(Vasticardium) unicolor (Sowerby, 1840). ··1987 Cuif,
Denis, JaiJlard & Zhu: Acrosterigma unicolor . •• J 987
ZR with ref. to Cuif et al.: Acrosterigma unicolor.
··1991 Abbott: Trachycardium unicolor. ··1992
Lamprell & Whitehead: Acrosterigma unicolor. ··1993
Vidal: Cardium unicolor not synonym of Cardium
enode as suggested by Fischer-Piette. ··1995 Chung-
Phuket Marine Biological Center Special Publication vol. 30 (2004)
823
unifasciatum - uniornatum
Hung & Hsi-Jen: Vasticardium unicolor (Sowerby).
"1999 Higo et al.: Acrosterigma (Vasticardium)
unicolor (Sowerby, 1841). "1999 Vidal:
Trachycardium unicolor Neogene fossils from Taiwan
(Gu Zhi-wei et al., 1976: 70). "1999 Vidal: Cardium
unicolor Sowerby, 1834 in Ranson 1967: 125 =
Acrosterigma seurati Vidal. "1999 Vidal: Cardium
unicolor Sowerby, 1834: fig. 29: = Acrosterigma
simplex (Spengler, 1799). "1999 Vidal: Cardium
unicolor Sowerby, 1834 in Sowerby 1841, p.107 =
Acrosterigma simplex (Spengler, 1799). CHARACTERS AND OCCURRENCE - OF COLLECTIONS. "ZMUC: I specimen. labeled Philippines colI.
Cuming. DESCRIPTIONS "1845 Reeve: shell oblongovate, somewhat attenuated towards the umbones,
radiately ribbed, 48 rounded, smooth ribs, the anterior
very finely crenul.ated; whitish, very obscurely spotted
with purple, interior tinged with pink. DIMENSIONS.
"1840 Sowerby: long. 1.50 poll. (length 38 mm), lat.
1.10 poll. (width 28 mm), alt. 1.70 poll (height 43 mm).
REMARKS. "1840 Sowerby: a slightly mottled form is
brought from the Brazils. "1845 Reeve: The ribs of
this species are very numerous considering the
distinctness with which they are developed; and the
interior of the shell is characteristically tinged in the
interior with rose pink. HABITAT. "1840 Sowerby:
Sandy mud at 5 fathoms. "1845 Reeve: found in sandy
mud at low water. "1999 Higo et at.: subtidal to 30 m,
sand and mud . DISTRIBUTION. "1845 Reeve: lsI. of
Ticao, Philippines. "1884 Grasse!: Philippines. "1994
Hylleberg: Central Indo-West Pacific, Australia. ··1999
Higo et al.: Tosa bay (southern Shikoku) and
southwards, Philippines, South China Sea, Micronesia,
Indo-Pacific . "ZMUC: Philippines.
unifasciatum Danilo & Sandri mss. - Synonym of
Acanthocardia tuberculata. » Regarding Cardium
unifasciata: Cardiidae. Recent. RECORDS. "1933 Coen:
Cardium unifasciata Danilo & Sandri mss = Eucardium
(Rudicardium) tuberculatum var. spinosa Coen. "1937
Coen: unifasciatum Danilo & Sandri mss =
Cerastoderma (Rudicardium) tuberculatum torquatum
Coen.
uniforme. - See under uniformis Phillips, 1835.
uniformis Phillips, 1835. - Nomen nudum accompanied
by figure. Combination to be revised: "Cardium "
uniforme Phillips, 1835. ?Cardiacea. Pre-Mesozoic.
RECORDS. '·1922-32 Sherborn, p. 6755: Corbis
uniformis Phillips, 1835 In. elf]. "1922-32 Sherborn,
p. 6755: Unicardium (Ph.) uniforme d'Orbigny, 1850.
- Corbis uniforme. "1999 Jacques Vidal (pers. com.)
MNHN, Paris: Fossils from the Paleontological
Collection. Cardium (Unicardium) uniforme Phillips.
Tourcien. Lombardy, B09996.
unimaculatum Broderip & Sowerby, 1833. [Figs. Part
I p. 106,144. Part 3 p. 914]. - ?Synonym ofCorculum
cardissa (Linne, 1758). OTHER SUGGESTIONS. Corculum
dionaeum (Broderip & Sowerby, 1829). » Regarding
Cardium unimaculatum : Cardiidae. Recent. TYPE
REPOSITORY. 3 syntypes. BMNH London. TYPE LOCALITY.
S. Pacific Ocean (Sowerby 1841). RECORDS. "1833
Broderip & sセキ・イ「ケZ@
Cardium unimaculatum. "1834
Sowerby, 46 t part, fig. I double asterisk: Cardium
unimaculatum nobis. "1840 Sowerby, p. 7, No. 94:
Cardium unimaculatum Sow., fig. I double asterisk =
Cardium Cardissa Linn. "1845 Reeve, species 114:
Cardium unimaculatum Broderip and Sowerby, 1838.
Pro. Zoo!. Soc. "1850 Jay: Cardium unimaculatum
Brod. & Sowb. = Cardium cardissa. 001858 Adams &
Adams, p. 458: Hemicardia unimaculata Sow. 001869
Romer: Cardium unimaculatum Sowerby, the west
coast of Central America. 001890 Paetel: Hemicardium
unimaculatum Sow. et Broderip. 001922-32 Sherborn,
p. 6757: Cardium unimaculatum Sowerby, 1833 .
001947 Bartsch: Cardium unimaculatum Broderip and
Sowerby, 1836 = Corculum dionaeum (Broderip and
Sowerby, 1829). "1977 Fischer-Piette: Cardium
unimaculatum Sowerby = Cardium cardissa Linne.
001977 Habe: Corcu/um unimaculatum Broderip &
Sowerby, 1836 sic = Corcutum cardissa (Linnaeus,
1758). "1992 Lamprell & Whitehead: Cardium
unimaculatum Broderip & Sowerby, 1833 = Corculum
dionaeum (Broderip & Sowerby, 1829).001999 Higo
et af.: Cardium unimaculatum Sowerby, 1833 =
Corculum cardissum (Linnaeus, 1758). - CHARACTERS AND OCCURRENCE - DESCRIPTIONS. 001845
Reeve: Shell laterally heart-shaped, very convex in the
middle, keeled, keel spinosely dentated, sides flatly
compressed, anterior radiately grooved, posterior
radiately flatly ridged, ridges armed down the middle
with minute raised dots; whitish, sprinkled with opaque
snowy spots, vividly stained around the posterior lunule
with pink. REMARKS. 001845 Reeve: Cardium
unimaculatum Sowerby, Conch . IIlus. f. 1* *, was
corrected to show the young of the Cardium cardissa;
it is, however, maturely grown shells. HABITAT. 001845
Reeve: found in coral sand on the reefs. DISTRlBUTlON.
001845 Reeve: lsI. of Annaa, Pacific Ocean. 001994
Hylleberg: Indo-Pacific.
Uniocardium Capellini, 1880 (genus).
Lymnocardiidae. Tertiary. RECORDS. 001980. Vokes:
Uniocardium Capellini, 1880. Subfamily uncertain.
Uniocardium Capellini, 1880 (subgenus). - Lymnocardiidae. Tertiary. RECORDS . 001887 Fischer:
Uniocardium Capellini, 1880 subgenus of genus
Limnocardium, Stoliczka, em. 1870. 001903
Andrussoff: Subgenus in genus Didacna to
accommodate species from the Ponti an basin. 001905
Dollfus, commenting Andrussow 1903: Uniocardium
Capellini, species from Italy.
uniornatum Vidal, 1999. [Figs . Part 3 p . 881]. Acrosterigma uniornatum Vidal, 1999. Cardiidae.
Recent. TYPE MATERIAL. Holotype: A right valve,
Waterwitch Bank, 500 m. Paratype 1: A left valve from
the same locality. Paratype 2: A right valve from the
same locality. Paratype 3: A right valve from the same
locality. Paratypes 4 to 25: Odd valves, same locality.
Paratypes 26-27: Two valves, Waterwitch Bank, 580600m. Paratypes 28 to 36: Odd valves, Waterwitch
Bank, 275-295m. Paratype 37: One valve, Combe Bank
370m. Paratypes 38-39: Two odd valves, Combe Bank,
550-552m. Paratypes 40 to 44: Odd valves, Field Bank,
490-500 m. (Vidal 1999). TYPE LOCALITY. Three
relatively shallow banks on the Melanesian Border
Plateau, within and in the vicinity of Wallis and Futuna
Territory, the south-western Pacific. TYPE REPOSITORY.
Museum National d'Histoire Naturelle, Bouchet,
Metivier, Richer de Forges col!. RECORDS. 001999 Vidal,
p. 293 , Table 19, Figs. 7C-H, 8A: Acrosterigma
824
Tropical Marine Mollusc Programme (TMMP)
uralicum - valahica
uniornatum sp. nov. - CHARACTERS AND OCCURRENCE - CHARACTERS of diagnostic value: ornamentation of the ribs uniform on all the parts of the
shell. On right valve cardinals show a great degree of
fusion. DESCRIPTION. ··1999 Vidal: Shell ovoid,
practically equilateral, never truncated posteriorly,
moderately elongated and rather globose. Lunule well
delineated, nearly similar on both valves, small, and
slightly hollowed. External colour uniform: white on
juvenile part, pale yellow on adult part. Interior white,
very rarely with traces of pink; no umbonal rays visible.
Hinge symmetrical and moderately angled. On right
valve cardinals show a greater degree of fusion at their
base than is typical for the genus. Mean rib nwnber
38.2 ± 1.4, range 36-41. In posterior area, interstices
thin, ribs flat with a central longitudinal furrow. Anterior
smooth part narrower than posterior part, which bears
elongated, thin , straight, oblique scales, often
encroaching upon axial furrow and even anterior part.
In median-posterior area, same features occur but
posterior zone is slightly raised and ribs become slightly
flattened triangles. Longitudinal furrow progressively
becomes less distinct, and anterior smooth part of rib
becomes its anterior flank. Scales remain about the
same as on posterior slope. On anterior half, ribs
become more rounded; scales become more numerous
and change, first into posterior flank-ridges, then into
moderately imbricated transverse top-ridges. MAXIMUM
DIMENSIONS. ··1999 Vidal: Length 18.1, width 22.4,
thickness (calculated) 15.2 mm. REMARKS. It is unclear
whether this species lives at 500-600 m where the shells
were dredged, or have rafted down from shallower
depths. Anyhow, its depth range is probably greater
than is usual for the genus. DISTRIBUTION. Known only
from the vicinity of Wallis and Futuna Islands.
uralicum. - Conocardium uralicum . Conocardiidae.
Fossil. RECORDS. ··1922-32 Sherborn, p. 6763:
Conocardium (Vern.) uralicum Keyserling, 1846. Cardium ouralicum sic. ··1931 ZR: Conocardium
uralicllm with ref. to Heritsch (1931).
ursaniense Landes, 1940. - Changed combination:
Granocardium ursaniense (Landes, 1940). Cardiidae.
Fossil. RECORDS. ··1954 Keen: Cardium (Ethmocardium) ursaniense Landes, 1940. ··1954 Keen:
Granocardium (Ethmocardium) ursaniense (Landes,
1940). ··1970 Speden: Ethmocardium ursaniense
Landes, 1940 and Granocardium (Ethmocardium) aff.
whitei (Dall) may be conspecific.
ursicimun. - Combination to be revised: "Cardium"
ursicinum. Cardiidae. Fossil. RECORDS. ··1891 ZR:
Cardium ursicinum, Lower Corallian of the Bernois
Jura [Ref. Loriol 1891).
ursina (subspecies and variety). - See under sllbodessae
ursina and subodessae ursina.
ursina Humphrey, 1797. - Synonym of Hippoplls
hippopus (Linnaeus, 1758). » Regarding Tridacna
ursina: Tridacnidae, Recent. RECORDS. ··1999 Higo et
al.: Tridacna ursina Humphrey, 1797 = Hippopus
hippopus (Linnaeus, 1758).
urthensis Chelidze, 1975. - Paradacna urthensis
Chelidze, 1975 . Lymnocardiidae. Lower Pliocene.
RECORDS. ··1975 ZR: Paradacna urthensis Chelidze,
1975 sp. nov. Georgia SSR, Zugdid region. Ref. to
Chelidze, p. 199.
urupellSe (variety). - See under bajarunasi urupense var.
Kolesnikov, 1929.
uspenica (subspecies). - See under matschigarica uspenica
uspenskaiae Koles. - Changed combination: Andrusovicardium uspenskaiae (Koles.). Lymnocardiidae.
Neogene. RECORDS. ··1986 N evesskaya et al.:
Andrusovicardium uspenskaiae (Koles.) comb. nov.
··1987 ZR with ref. to Nevesskaya et al. p.I92:
Andrusovicardium uspenskaiae (Koles.) comb. nov.
ustrix Gardner, 1926. - Combination to be revised:
"Cardium" ustrix Gardner, 1926. Cardiidae. Tertiary.
RECORDS. ··1926 ZR: Cardium (Trachycardium) ustrix
Gardner, 1926 sp. n. Tertiary U.S.A.
ustulatum Schroter. - Unknown species. ZMUC: label
in box with muricatum Schroter, 7: Cardium ustulatum
= muricatum. It is probably a catalogue name.
utcholokensis Slodkewitsch, 1938. - Changed
combination: Pro/ulvia utcholokensis (Slodkewitsch,
1938). Cardiidae. Pliocene. TYPE LOCALITY. Kamtchatka.
RECORDS. ··1973 Keen: Fulvia utcholokensis
(Slodkewitsch , 1938) (Papyridea). ··1976 ZR :
Pro/ulvia utcholokensis (Slodkewitsch, 1938) Comb.
nov. Transferred from Papyridea. Ref. to Kafanov, p.
482 [orig. p. 111]. ··1976 ZR: Papyridea utcholokensis
Slodkewitsch, 1938 Referred to Pro/ulvia. Ref. to
Kafanov, p. 482 [orig. p. 111).
uvutschensis Iiyina, 1963. - Synonym of Serripes
groenlandicus (Mohr, 1786). » Regarding Serripes
uvutschensis: Cardiidae. Upper Miocene. TYPE
LOCALITY. Kamtchatka. RECORDS. ··1973 Keen: ?
Serripes uvutschensis llyina, 1963. ··2000 Coan et at.:
? Serripes (?) uvutschensis Il'ina, 1963 = Serripes
(Serripes) groenlandicus (Mohr, 1786).
uyemurai Kanehara, 1937. - Changed combination:
Cilia to cardium uyemurai (Kanehara, 1937). Cardiidae.
Pliocene. TYPE LOCALITY. Sakhalin Island. RECORDS.
··1937 Kanehara: Cardium uyemurai. ··1937 ZR:
Cardium (Cerastoderma) uyemurai Kanehara, 1937b
sp. n. Congi Series (Pliocene) Japan. ··1973 Keen:
Clinocardium uyemurai (Kanehara, 1937). ··1974 ZR:
Ciliatocardium uyemurai Kanehara, 1937 Comb. nov.
Ref. to Kafanov, p. 1469.
uzunlarica Tschep. & Mih. - Didacna uzunlarica
Tschep. & Mih. Lymnocardiidae. Fossil. RECORDS.
··1989 Mikhailesku: Didacna uzunlarica Tschep. &
Mih.
V
valachica Papaianopol. - Pontalmyra valachica Papaian.
Lymnocardiidae. Fossil. RECORDS. ··1987 Taktakishvili,
p. 67: Pontalmyra valachica sic Papaian. Pontalmyra
valahica sic Papa ian.
valahica (spelling). - See under valachica Papaianopol.
valahica Papaianopo1, 1981. - Pontalmyra valahica
Papaianopo1, 1981. Lymnocardiidae. Upper Miocene.
RECORDS. ··1981 ZR: Pontalmyra (Pontalmyra)
valahica Papaianopol, 1981 sp. nov. Romania. Ref. to
Papaianopol, p. 17.
valahica Papaianopol, 1989. - Pachydacna valahica
Papaianopol, 1989. Lymnocardiidae. Pliocene. TYPE
LOCALITY. Basin Dacique, Romania. RECORDS. ··1989
Papaianopol, p. 96, fig.: Pachydacna (Pachydacna)
Phukel Marine Biological Center Special Publication vol. 30 (2004)
825
valahicus - variegatum
valahica sp. nov.
valahicus Papaianopol, 1975. - Dacicardium valahicus
sp. nov. Papaianopol, 1975. Lymnocardiidae. Pliocene.
REcORDs. ··1975 ZR: Dacicardium valahicus sp. nov.
Papaianopol, 1975 Romania. Dealul Virful Coastei.
Ref. to Papaianopol, p. 257. ··1978 ZR: Dacicardium
valahicus Papaianopol, 1975. Description. Ref. to
Papaianopol et af. p. 308.
valdense Pictet & Campiche, 1866. - Combination to
be revised: "Cardium" valdense Pictet & Campiche,
1866. Cardiidae. Fossil. RECORDS. ··1980 Ruhoff:
Cardium valdense Pictet & Campiche, 1866; [18611871] seL 4, p. 253, pI. 120, fig. 3
Valenciennius (genus). - Lymnocardiidae. Pliocene.
RECORDS. "1955 Ebersin: Valenciennius. "1975 ZR:
Valenciennius. Systematic accounts of species.
Romania. Ref. to Hanganu.
valense Boghachev, 1938. - New combination:
Lymnocardium valense Boghachev, 1938. Lymnocardiidae. Fossil. REcoRDs. ··1939 ZR: Limnocardium
valense Boghachev, 1938 sp. n.
valida Papaianopol, 1981. - Pontalmyra valida
Papaianopol, 1981. Lymnocardiidae. Upper Miocene.
RECORDS. ··1981 ZR: Pontalmyra (Pontalmyra) valida
Papaianopol, 1981 sp. nov. Romania. Ref. to
Papaianopol, p. 31.
validus Papaianopol, 1975. - Dacicardium validus
Papaianopol, 1975. Lymnocardiidae. Pliocene.
RECORDS. ··1978 ZR: Dacicardium validus Papaianopo1, 1975. sp. nov. Romania. Ref. to Papaianopol et
al. p. 311.
vandeneckii d'Orbigny, 1850. - Combination to be
revised: "Cardium" vandeneckii d'Orbigny, 1850.
RECORDS. ··1922-32 Sherborn, p. 6791: Cardium
vandeneckii d'Orbigny, 1850
vanhyningi (subspecies). - See under robustum
vanhyningi Clench & Smith, 1944.
vaquerosensis Arnold, 1908. - Changed combination:
Trachycardium vaquerosense (Arnold, 1908).
Cardiidae. Lower Miocene. RECORDS. ··1908 ZR:
Cardium (Trachycardium) vaquerosensis Arnold, 1908
sp. n. Lower Miocene (Vaqueros formn.), California
(San Mateo Co.) ... 1944 Keen & Bentson: Cardium
(Trachycardium) vaquerosensis Arnold, 1908 (n. sp.).
··1944 Keen & Bentson: Trachycardium vaquerosense
(Arnold).
varia (variety). - See under rudis vaL varia.
Varicardium (genus) Marwick, 1944. - Cardiidae. Fossil.
RECORDS. ··1944 ZR: Varicardium gen. nov. Cardiidae
type Protocardium serum Hutton Catal. Tert. Moll.
1873, p. 23 , gen. also includes Cardium patulum Hutton
tom. cit. p. 23 Ref. to Marwick 1944. ··1980 Vokes:
Varicardium Marwick, 1944 in subfamily
Protocardiinae.
Varicardium (subgenus). - RECORDS. ··1977 Popov:
(Varicardium) Marwick, 1944. SUbgenotype: Cardium
patulum Hutton, 1873.
varicostata Branson, 1938. - Dexiobia varicostata
Branson, 1938. Previously in Cardiacea. Carboniferous.
RECORDS. ··1940 ZR: Dexiobia varicostata Branson,
1938 sp. n. Northview Siltstone King Butte, Missouri
Mississippian (Carboniferous) U.S.A.
varicosum d'Orbigny, 1850. - Not a cardiid. RECORDS.
··1922-32 Sherborn, p. 6805 : Unicardium (Sow.)
varicosum d'Orbigny, 1850. - Venus varicosum
Sowerby, 1821.
variegatum Solander MS . - Synonym of Vasticardium
leucostomum (Born). » Regarding Cardium
variegatum: Cardi idae. Recent. RECORDS. ··1817
Dillwyn: Cardium variegatum Solander's MSS =
Cardium leucostomum Born. ··1922-32 Sherborn, p.
6807: Cardium variegatum Solander, 1786 [n on.] .
variegatum Sowerby, 1840. [Figs. Part I p. 118, 138,
173 . Part 3 p . 870]. - Changed combination:
Acrosterigma variegatum (Sowerby, 1840). Cardiidae.
Recent. RECORDS . TYPE MATERIAL . Three shells
considered as syntypes in BMNH, not catalogued,
Cuming collection, labeled from Ticao (Philippines)
and another location (label illegible). The dimensions
given by Sowerby for his figured specimen could fit
the smallest specimen, but the colour pattern, the rib
number (48 given), and the locality (Leyte given) do
not match. The specimen figured by Vidal (1999, figs .
5A-C) better fits the figure, description and locality
given by Reeve (1845: Sp. 75) . TYPE LOCALITY. Ad
insulam Leyte, the Philippines (Sowerby 1840).
RECORDS. ··1840 Sowerby, 181 th part, fig. 57: Cardium
variegatum nobis. ··1840 Sowerby, Proc. Z.S. London
p . 107: Cardium variegatum. ··1840 Sowerby,
catalogue p. 4, No. 45 : Cardium variegatum Sow.
··1841 . Sowerby, corrected list, p. 8: Cardium
variegatum (Sowerby, No. 45). ··1845 Reeve: Cardium
variegatum. ··1858 Adams & Adams, p. 455: Cardium
(Trachycardium) variegatum Sow. ··1884 Grasset:
Cardium variegatum Sow. ··1890 Paetel: Cardium
variegatum Sow. ··1903 Hidalgo: Cardium variegatum
Sowerby, 1840. "1922-32 Sherborn, p. 6807: Cardium
variegatum Sowerby, 1841. ··1928 Faustino: Cardium
variegatum Sowerby. ··1952 Kuroda & Habe:
"variegatum (Sowerby)" (non Solander) =
Trachycardium "variegatum (Sowerby)". ··1960
Kuroda: Trachycardium (Acrosterigma) variegatum
(Sowerby). ··1977 Fischer-Piette: Laevicardium
variegatum Sowerby. ··1977 Habe: Trachycardium
(Vasticardium) variegatum (Sowerby, 1841). ··1991
Rao, Rao & Maitra: Trachycardium variegatum
(Sowerby, 1840). ··1997 Kafanov: Cardium
variegatum in Sowerby, 1834 p. 4, 8, fig. 57 - not syn.
of Papyridea (Papyridea) aspersa Sowerby in Broderip
et Sowerby, 1833. ··1999 Higo et al.: Acrosterigma
(Vasticardium) variegata (Sowerby, 1841). ··1999
Vidal: Acrosterigma variegatltm (Sowerby, 1840).
··2002 Hylleberg & Kilburn, p. 63: Acrosterigma
variegatum (Sowerby, 1840). ID. by J. Vidal. .. 2003
Hylleberg & Kilburn, p. 177: Acrosterigma variegatum
(Sowerby, 1840). - CHARACTERS AND OCCURRENCE - CHARACTERS of diagnostic value. ··1999
Vidal: it cannot be confused with any other species
because of its unique lunule: practically absent on left
valve, wide on right valve, projecting from the margin.
DESCRIPTIONS. ··1840 Sowerby: shell oval,
subventricose, red-brown blotches, 48 ribs, anterior
ones rounded, crenulated; posterior ones much
angulated, the most posterior ribs somewhat flattened,
with oblique tubercles [I have translated from Latin).
··1845 Reeve: shell somewhat squarely ovate ,
depressed, posterior slightly angulated, radiately ribbed,
38 ribs, the extreme posterior ribs flatly obsolete,
826
Tropical Marine Mollusc Programme (TMMP)
variegatum - vattoni
slightly prickly, middle ribs peculiarly sharply angled,
anterior ribs rounded, very finely crenated; white,
profusely variegated with bright rose spots. --1999
Vidal: shell ovoid, almost equilateral (anterior dorsal
margin slightly angled), not posteriorly truncated, nor
elongated and moderately compressed . Lunule wide
on right valve, projecting from the margin against left
valve, and composed of two parts: a strongly hollowed
part near margin, forming an elongated socket, and an
elevated part behind that, forming a large rib-like ridge,
both parts being concentrically finely ridged; practically
no lunule on left valve. Exterior beige variegated with
large irregular patches of orange, brown, and purple,
which are visible interiorly. Hinge markedly
asynunetrical and moderately angled . Mean rib number
40.2, ± 2.2, range 36-43 [never as many as 48, as
indicated by Sowerby] . Anterior ribs of posterior shell
much wider, with oblique scales straddling a very thin
intermediate furrow. Ribs on median posterior part
triangular with a slightly concave posterior side, and a
small, entirely smooth crestal fold. Anteriorly ribs
become progressively rounded, first a little retro-ridged
then top-ridged on anterior half. Interstices always
rounded, excepted in posterior part, and slightly
narrower than ribs. DIMENSIONS. --1840 Sowerby: long.
1.70 poll. (length 43 mm), lat. 1.80 poll. (width 46 nun),
alt. 1.20 poll (breadth 31 mm). --1999 Vidal: length
52.6 mm, width 58 .1 mm, breadth (calculated) 37.0
mm. HABITAT. "1999 Higo et al.: recorded from I to
500 m depth; intertidal to 20 m, sand REMARKS. --1840
Sowerby: Ribs much closer than in C. muricatum
(Sowerby 1840). --1845 Reeve: there is a peculiarity
in the growth of this shell of which we find a similar
indication in the Cardium angulatum; - a depressed
square-like form, with the above described variation
in the structure of the ribs. --1999 Vidal: C. radula is
described as having the ribs angular on both sides,
which is not the case with this species. A. variegatum
is very constant in characters. It cannot be confused
with any other species, particularly because of its
unique lunule. DISTRIBUTION. --1845 Reeve: lsI. of
Ticao, Philippines. --1884 Grasset: Philippines. --1999
Vidal: tropical western Pacific as far east as Wallis &
Futuna; very abundant in the Philippines. -- I 999 Higo
et al.: Okinawa islands and southwards, Philippines.
South China Sea, Southeast Asia. "2002 Hylleberg &
Kilburn: South India. --2003 HyUeberg & Kilburn:
Vietnam. "Of authors: Thailand, Philippines, Japan.
variegatum of authors. - Misidenti fication of Papyridea
aspersa . RECORDS. --1997 Kafanov: Cardium
variegatum auctt. - in part syn. of Papyridea (Papyridea) aspersa Sowerby in Broderip et Sowerby, 1833 .
variegatum of Mabille 1895. - Misidentification of
Papyridea spinosum var. asperum Sowerby. RECORDS.
--1909 Lamy : Cardium variegatum in Mabille
(1895)(non Sow.) = Cardium (Papyridea) spinosum
Meuschen var. asperum Sowerby, Dall.
variegatum of Roemer. - Misidentification of Cardium
maculosum Wood. RECORDS. --1903 Hidalgo: Cardium
variegatum in Roemer, fig. 7 = Cardium maculosum
Wood.
variegatum h . Solander, 1786 (variety) . Infrasubspecific. RECORDS. --1786 Solander, p. 17 no.
382: Cardium variegatum, var. b. S.
*varium. - Homonymy to be noticed. Cardium varium
from 2 sources: Sowerby, 1834. - Tscheltzov, 1965.
varium Sowerby, 1834. - [Fig. Part I p. 109]. Synonym
of Fulvia tenuicostata (Lamarck) . OTHER JUDGEMENTS :
Fulvia australis (Sowerby, 1834). - Papyridea
papy raceum Schroter [not available]. - Fulvia
lenuicostata (Lamarck). » Regarding Cardium
varium: Cardiidae. Recent. TYPE LOCALITY. Swan River,
Australia. RECORDS. --1834 Sowerby, 49 th part, No. 19:
Cardium varium nobis. --1840 Sowerby, catalogue p.
I, No.8 : Cardium varium = Cardium tenuicostata Lam.
--1977 Fischer-Piette: Cardium varium Sowerby =
Papyridea papyraceum (Chemnitz) Schroter. --1994
Vidal: ? Cardium varium Sowerby, 1834 = Fulvia
(Fulvia) australis (Sowerby, 1834).
varium Tscheltzov, 1965. Combination to be revised:
"Cardium" varium Tscheltzov, 1965 . Card iacea.
Pliocene. RECORDS . -- I 965 ZR: Cardium varium
Tscheltzov, 1965 sp. nov. Akchagylian (Pliocene),
Turkmenia, USSR.
varnensis (variety). - See under rectidens var. varnensis
Coen, 1917.
vasoevitschi (spelling) . - See under vasoevitshi Kolesnikov.
vasoevitshi Kolesnikov, 1929. - Synonym of Replidacna
tumidum (Ali-Zade & Ribina). » Regarding C.
vasoevitshi: Cardiacea. Sarmatian. RECORDS. --1929
ZR : Cardium vasoevitshi Kolesnikov, 1929 sp. n.
Sarmatian. --1977 ZR: Cerastoderma (lnaequicostata)
vasoevitshi (Kolesnikov); syn. nov. Cardium
(R eplidacna) tumidum Ali-Zade & Ribina . Ref to
Paramonova p. 70. [Transl. p. 320]. --1979 ZR:
Protractiforma vasoevilschi (Kolesnikov). Referred to
new subgenus P. (Fischericardium) . Ref. to
Bagdasaryan, p. 44.
Vasticardium Iredale, 1927 (genus). - Cardiidae.
Oligocene and Recent . RE CORDS. --1927 ZR:
Vasticardium Iredale, 1927b [gen. n.] type Cochlea
nebulosa Martyn. --1978 Maxwell, p. 21: Vasticardium
is a synonym of Acrosterigma. --1980 Vokes :
Vasticardium Iredale, 1927 in subfamily
Trachycardiinae. --1999 Vidal: Vasticardium Iredale,
1927: 75 . --2001 Vidal, p. 407, Vasticardium. CHARACTERS
AND
OCCURRENCE
DESCRIPTIONS. --200 I Vidal: tentacles numerous.
Vasticardium Iredale, 1927 (subgenus). - Cardiidae.
Oligocene and Recent. GENOTYPE. Iredale (1927)
designated C. nebulosum ex Martyn type species of
subgenus Vasticardium of genus Acrosterigma Dali,
1900. Upgraded to generic level by authors. RECORDS.
"1977 Popov : (Vasticardium) Iredale, 1927 .
SUbgenotype: Cochlea nebulosa Martyn, 1792. --1999
Rigo et al.: Vasticardium (subgenus) Iredale, 1927.
Type species: Vasticardium nebulosum Iredale, 1927
(0. d.) . - CHARACTERS AND OCCURRENCE CHARACTERS of diagnostic value: resembling A.
(Acrosterigma) but eaves more strongly sculptured;
posterior margin notched. Indo-Pacific-North and
Central America-Europe (Moore et al. 1969).
vattoni Coquand, 1862. - Changed combination:
Proto cardia vattoni (Coquand) . Cardiidae. Cretaceous.
RECORDS. --1980 Ruhoff: Cardium vattoni Coquand,
1862 vol. 2: 207, pI. II, fig. 5. "1999 Jacques Vidal
(pers. com.) MNHN , Paris: Fossils from the
Phuket Marine Biological Center Special Publication vol. 30 (2004)
827
vaughani - venustum
Paleontological Collection. Protocardia vattoni
Coquand. Cenomanian. Algeria, B 19034.
vaughani Stephenson, 1923. - New name. Combination
to be revised: "Cardium" nixicollis Stephenson, 1923.
Cardiidae. Cretaceous. RECORDS. --1923 ZR: Cardium
vaughani Stephenson, 1923 sp. n. Cretaceous U.S .A.
--1936 ZR: Cardium (Protocardia) vaughani
Stephenson, 1923; see Cardium nixicollis Stephenson,
1923 nom. nov. --2001 Schneider & Carter:
Pleuriocardia
(Dochmocardia?)
vaughani
(Stephenson, 1923). Cardiidae. Upper Cretaceous,
North Carolina USA.
vaughaniana (subspecies). - See under gurabica
vaughaniana Mansfield, 1932.
venestum Kolesnikov, 1929. - Changed combination:
Sarmaticardium venestum (Kolesnikov). Cardiidae .
RECORDS. --1929 ZR: Cardium venestum Kolesnikov,
1929 sp. n. Sarmatian. --1977 Popov: Cerastoderma
(Obsoletiforma) venestum. --1979 ZR: Sarmaticardium
venestum (Kolesnikov) Comb. nov. Ref. to
Bagdasaryan, p. 41.
ventricosa (mode). - See under ventricosum Bruguiere,
1789.
*ventricosum. - Homonymy to be noticed. Cardium
ventricosum from 3 sources: Bruguiere , 1789 . Orbigny, 1843. - Hall, 1847.
ventricosum Bruguiere, 1789. [Figs. Part I p. 66, 91,
93. Part 3 p. 856]. - Synonym of Trachycardium
robustum (Lightfoot in Solander, 1786) . OTHER
JUDGEMENTS: Cardium macula tum Gmelin, 1791. Cardium magnum Born, 1780. - Cardium obliquum
Spengler, 1799. » Regarding Cardium ventricosum:
Cardiidae. Recent. RECORDS. --1789 Bruguiere, p. 228,
No. 25: Cardium ventricosum Nob . --1798 Bruguiere.
ID by Lamarck, pI. 299. fig. I. Cardium ventricosum.
"1815 Wood: Cardium ventricosum Brug. "1817
Dillwyn: Cardium ventricosum Bruguiere = Cardium
maculatum Gme1in. --1836 Bosc: Cardium
ventricosum. --1839 Anton : Cardium ventricosum
Bruguiere = Cardium maculatum Gmelin. --1840
Sowerby, catalogue p. 5, No. 63 : Cardium ventricosum,
Brug. --1844 Reeve, sp. 20 : Cardium ventricosum
Bruguiere in Lamarck [1819] = Cardium magnum
Born, 1780. --1849 Roemer: Cardium ventricosum
Brug. --1850 Jay : Cardium ventricosum Brug. =
Cardium magnum Born. "1856 Wood & Hanley, No.
19: Cardium ventricosum Brug. --1869 Romer:
Cardium ventricosum Bruguiere = Cardium magnum
80m. --1870 Morch: C. ventricosum 8rug. Jonas,
junior synonym of Cardium obliquum Spgl. = C.
magnum Born. --1890 Paetel: Cardium venlricosum
Brug. = magnum Born . --1902 Dall : Cardium
venlricosum Bruguiere, 1789 = Cardium robustum
Solander, 1786. --1921 Maury : ventricosum Bruguiere,
1789 = Cardium (Cerastoderma) robustum Solander,
1786. "1902 Sherborn, p. 1035 : Cardium ventricosum
Bruguiere, 1789. --1945 ZR: Cardium ventricosum
8rug. [Ref. Lamy 1945]. "1953 Olsson & Harbison:
Cardium ventricosum Bruguiere, 1789 = Cardium
(Dinocardium) robustum Solander, 1786. "1967
Rehder: Cardium ventricosum Bruguiere, 1789, p. 228
= Dinocardium robustum (Lightfoot, 1786). --1974
Abbott: Cardium ventricosa Bruguiere, 1789 =
Dinocardium robustum (Lightfoot, 1786). --1977
Fischer-Piette: Cardium ventricosum Bruguiere =
Laevicardium robustum Solander. - CHARACTERS
AND OCCURRENCE - OF COLLECTIONS. --ZMUC: 2
specimens from Galveston, Texas; 8 shells from St.
Anna East Mexico coil. Caspersen 1870 ZMUC: 1
specimen labeled Lond. 1835. REMARKS. --1939.
McLean: Cardium magnum 80m, 1780 in Reeve (1844
sp. 20) = Cardium robuslum. DISTRIBUTION. "1844
Reeve: C. magnum: Gulf of Mexico. --1939 McLean:
C. robuslum: East coast of USA, Mexico. --ZMUC: C.
ventricosum: Galveston, Texas, St. Anna East Mexico.
ventricosum Hall, 1847. - Replaced by "Cardium ,. halli.
Deshayes. Combination to be revised. » Regarding
C. ventricosum: Cardiidae. Silurian. RECORDS. "1857
Deshayes, p. 49: Cardium ventricosum (Edmondia).
Hall . 1847. - Replaced by C. halli. Desh.
ventricosum Orbigny, 1843. - New name : "Cardium"
insujJlatum Deshayes. Combination to be revised. »
Regarding C. ventricosum: Cardiidae. Cretaceous .
RECORDS. "1857. Deshayes , p. 49 : Cardium
ventricosum. d ' Orb. non Brug. --1857 Deshayes, p. 59:
Cardium insufflatum Desh. New name for Cardium
venlricosum d'Orbigny, 1843. --1922-32 Sherborn, p.
6843: Cardium ventricosum d'Orbigny, 1844. "1999
Jacques Vidal (pers. com.) MNHN, Paris: Fossils from
the Paleontological Collection. Cardium ventricosum
d'Orbigny. Senonien. Algeria, 819825.
ventricosum Tolmachoff, 1927. - New name :
Conocardium ventriculosum Branson , 1942.
Conocardiidae. Paleozoic. RECORDS. "1927 ZR :
Conocardium ventricosum Tolmachoff, 1927 sp. n.
Palaeozoic Arctic. --1942 ZR: Conocardium
ventricosum Branson, 1942 p. 387 Tolmachoff 1926
non Hall 1860 replaced by Conocardium ventriculosum
nom. nov. Branson, 1942.
ventriculosum Branson, 1942. - Conocardium
ventricosum Branson, 1942. Conocardiidae. Fossil.
RECORDS. --1942 ZR: Conocardium ventriculosum nom.
nov. Branson, 1942 p. 387, for Conocardium
ventricosum Tolmachoff 1926 non Hall 1860.
venusta Talent, 1963. - Cypricardinia venusta Talent,
1963. Cardiacea. Devonian. RECORDS. --1963 ZR:
Cypricardinia venusta Talent, 1963 sp. nov. Kilgower
Member, Tabberabbera Formation, Victoria, Australia,
Devonian.
*venustum. - Homonymy to be noticed. Cardium
venustum from 3 sources: Deshayes, 1858. - Dunker,
1861. - Gabb, 1873 .
venustum Deshayes, 1858. - Synonym of Papyridea
capsoides (Bayan, 1873). » Regarding Cardium
venustum: Cardiidae. Eocene . RECORDS . --1886
Cossmann: Cardium venustum Deshayes = Papyridea
capsoides (8ayan, 1873) . --1922-32 Sherborn :
Cardium venustum [In index only. Not located in text].
-- J980 Ruhoff: Cardium venustum Deshayes, J858;
[1856-1865] vol. 1: 575, pl. 55 , figs . 22-24.
venustum Dunker, 1861. - Synonym of Trigoniocardia
medium (Linne, 1758). » Regarding Cardium
venustum: Cardiidae. Recent. RECORDS. --1861 Dunker:
Cardium venustllm. "1902 Dall : Cardium venustum
Dunker, 1861 = Cardium medium Linnaeus, 1758.
--1939 McLean: Cardium venustum Dunker, 1861 =
Trigoniocardia medium Linne, 1758. --1977 FischerPiette: Cardium venustum Dunker = Corculum medium
828
Tropical Marine Mollusc Programme (TMMP)
venustum - verioti
Linne. - CHARACTERS AND OCCURRENCE DISTRIBUTION. ··1939 McLean: C. medium: North
Carolina USA, West Indies, Brazil.
venustum Gabb, 1873. - Synonym of Laevicardium
serra tum (Linnaeus, 1758). OTHER JUDGEMENTS: New
name Laevicardium venustulum (Gabb, 1873). »
Regarding Cardium venustum: Cardiidae. Neogene.
RECORDS. "1873 Gabb: Cardium venustum Gabb, 1873.
"1902 Dall : Cardium venuslum of Gabb, 1873 (not of
Dunker, 1861) = Cardium serratum Linnaeus, 1758.
"1922 Pilsbry, p. 421-422, pI. 25, figs. 2, 7: Cardium
(Laevicardium) venustum Gabb, 1873. ··1925
Woodring , p. 145: Cardium venustum Gabb .
Dominician Republic. ··1939 McLean: Cardium
venustum in Gabb, 1873 = Laevicardium serratum
sen'atum (Linne, 1758). ··1989 Vokes, p.129:
Laevicardium venustum (Gabb, 1873) nec Dunker
1861. ··1989 Vokes. p.129: Cardium venus tum Gabb,
1873 preoccupied name. "1989 Vokes. p.129:
Laevicardium venustum (Gabb, 1873) preoccupied
name. ··1989 Vokes. p. 129: Cardium venustum Gabb,
1873 replaced by Laevicardium venustulum nom. nov.
venzoi Tavani, 1948. - New combination: Protocardia
vemoi Tavani, 1948. Cardiidae. Cretaceous. RECORDS.
··1948 ZR: Cardium (Protocardia) venzoi Tavani, 1948
sp. nov. Cretaceous Somaliland.
Vepricardiini Kafanov & Starobogatov, 1977 (tribus) .
- Cardiidae. RECORDS. ··1977 Kafanov et Starobogatov:
Vepricardiini. ··1977 ZR: Vepricardiini Kafanov &
Starobogatov Tribe nov. of Cardiidae, type genus
Vepricardium Iredale, 1929. Ref to Kafanov & Popov
p. 62 [Trans!. p. 312].
Vepricardium Iredale, 1929 (genus). - Cardiidae.
Paleocene and Recent. GENOTYPE. Vepricardium
pulchricostatum Iredale, 1929. Original designation.
Recent, Australia. RECORDS. ··1929 ZR: Vepricardium
gen. n. [type] pulchricostatum sp. n. Iredale, 1929a E.
Australia. ··1977 ZR : Vepricardium Ireda1e, 1929.
Designated type genus for new tribe Vepricardiini
Kafanov & Starobogatov. Ref. to Kafanov & Popov p.
62 [Transl. p. 312]. ··1980 Vokes: Vepricardium Ireda1e,
1929 in subfamily Cardiinae. ··1995 Pacaud & Renard:
Vepricardium Iredale, 1929 in subfamily Cardiinae.
··1998 Schneider: Vepricardium. No Paleogene cardiids
belong to this genus. ··1999 Higo et al.: Vepricardium
Iredale, 1929. Type species: Vepricardium
pulchricostatum Iredale, 1929 (0. d.). ··2000 Vidal, p.
447: Vepricardium Iredale, 1929. Type species:
Vepricardium pulchricostatum Ireda1e, 1929 =
Vepricardium multispinosum (Sowerby, 1841). ··200 I
Vidal, p. 407: Vepricardium. - CHARACTERS AND
OCCURRENCE - CHARACTERS of diagnostic value:
shell like Bucardium in outline (semicircular) but large.
Posterior ribs not digitate, dorsal margins with smooth
areas, Rib sculpture of close-set spines. Hinge plate
rounding into anterior and posterior margins (Moore
et al. 1969, emend. Hylleberg 1994). DESCRIPTIONS.
··1978 Maxwell: the genus Vepricardium has nearly
equilateral valves with the posterior end well rounded
to weakly truncate. ··2000 Vidal : ornaments of the ribs
always situated along the posterior flank or on the
posterior part of the top, consisting of rounded tubercles
or festooned periostracal or calcareous insertions on
the juveni les, becoming more or less elaborate
tubercles, spines and lamellae in the adults. Surface of
ribs and interstices almost always covered by numerous
pustules or tiny acute spines. ··200 I Vidal: tentacles
numerous. Eyes on tip of tentacles. ··2002 Schneider:
Vepricardium has a conical right anterior cardinal tooth
that projects horizontally (i.e. into the socket of the
hinge of the left valve) a greater distance than the height
or width of the tooth. The tooth is directly above the
right anterior cardinal socket and does not obscure the
socket face. The tip of the right anterior cardinal tooth
is sharp, and the right anterior cardinal tooth is not
connected to the right posterior cardinal. The 'lunule
flap' is strongly folded over the exterior shell surface
anterior of the beak, covering the entire lunule, and the
lunule flap has several nearly vertical ridges on its
surface. The dorsal surface of the nymph is distinctly
concave. The posterior end of the nymph has a sharp
dorsal extension (nymph flare). The radial ribs are
strongly expressed from the ventral margin all the way
to the umbo. It has strong, sharp spines with notches
on the posterodorsal portion of their spines. The
adductor muscle scars are not impressed into the shell.
Vepricardium has the right caecum located in the middle
of the stomach floor. The lengths of the labial palps
are more than half the length of the animal. REMARKS,
··2000 Vidal : Vepricardium is close to Trachycardium
s. s., particularly to the subgenus Dal/ocardia (Vidal
1999), close to Fulvia (Vidal 1994) and very close to
the group of the Neogene Cardium multicostatum
Brocchi, 1843 (Vidal 1999). DISTRIBUTION. EuropeAfrica-East Indies-Australia.
Vepricardium Iredale, 1929 (subgenus). - Cardiidae.
Paleocene and Recent. RECORDS . ··1995 Pacaud &
Renard: (Vepricardium) Iredale, 1929.
Vepricardium in Iredale & MacMichael 1962. - Printer's
error. RECORDS. ··1962 Iredale & MacMichael, p. 21:
Cardium Vepricardium Iredale. [Vepricardium is not a
species].
vera Kvaliashvili, 1978. - Digressodacna vera
K valiashvili, 1978. Lymnocardiidae. Pliocene.
RECORDS. ··1978 ZR: Digressodacna vera KvaJiashvili,
1978 sp. nov. Georgia SSR. Ref. to Kvaliashvili p. 79.
verbeeki Martin, 1879. - Combination to be revised:
"Cardium" verbeeki Martin, 1879. Cardiidae. Upper
Miocene. TYPE LOCALITY. Tjilanangsch., Java, Indonesia.
TYPE REPOSITORY. ?Geologischen Reichs-Museum,
Leiden, the Netherlands. RECORDS. ··1919 Martin:
Cardium verbeeki Mart.
verecunda Papaianopol, 1978. - Caladacna verecunda
Papaianopol, 1978, Lymnocardiidae. Upper Miocene.
RECORDS. ··1978 ZR: Caladacna verecunda sp. nov.
Romania. Ref. to Papaianopol, I. p. 138. fig ' d.
verecunda Papaianopol , 1981 . - Pontalmyra verecunda
Papaianopol , 1981. Lymnocardiidae. Upper Miocene.
RECORDS. ··1981 ZR : Pontalmyra (Crassadacna) verewnda sp. nov. Romania. Ref to Papaianopol, p. 41.
*verioti. - Homonymy to be noticed. Cardium verioti
from 2 sources: Buvignier, 1852. - Loriol & Cotteau,
1867.
verioti Buvignier, 1852. - Combination to be revised:
"Cardium" verioti Buvignier, 1852. Cardiidae. Fossil.
RECORDS. ··1980 Ruhoff: Cardium verioti Buvignier,
1852; atlas p. 16, pI. 16, figs. 1-5.
verioti Loriol & Cotteau, 1867. - Combination to be
Phuket Marine Biological Center Special Publication vol. 30 (2004)
829
verneuili - vicksburgellse
revised: "Cardium" verioti Loriol & Cotteau, 1867.
Cardiidae. Fossil. RECORDs. "1980 Ruhoff: Cardium
verioti Loriol & Cotteau, 1867; vol. 21 : 561, pI. 9, f. 8.
verneuili Deshayes, 1838. - Combination to be revised:
"Cardium" verneuili Deshayes, 1838. Cardiidae.
Miocene. RECORDS. "1886 Fischer: Cardium verneuili
Deshayes. "1922-32 Sherborn, p. 6860: Cardium
verneuili Deshayes, 1838. "1999 Jacques Vidal (pers.
com.) MNHN, Paris: Fossils from the Paleontological
Collection. Cardium verneuilli(i}. URSS, the Crimean,
B32855, -6.
verneuilianum d'Orbigny, 1845. - Combination to be
revised: "Cardium" verneuilianum d' Orbigny, 1845.
Cardiidae. Fossil. RECORDS. "1922-32 Sherborn, p.
6861: Cardium verneuilianum d'Orbigny, 1845 .
verrucosa meridiolla/is Strougo, 1976. - Acanthocardia
verrucosa meridionalis Strougo, 1976. Cardiidae.
Upper Eocene. RECORDS. "1976 ZR: Acanthocardia
(Agnocardia) verrucosa meridionalis Strougo, 1976
subsp. nov. Egypt, Maadi. Ref. to Strougo, p. 106.
*verrucosum. - Homonymy to be noticed. Cardium
verrucosum from 2 sources: Deshayes , 1829. Andrzejoswky, 1835.
verrucosum Andrzejoswky 1835. - Replaced by
"Cardium" podolicum Deshayes. Combination to be
revised. » Regarding "c." verrucosum: Cardiacea.
Fossil. RECORDS. "1857. Deshayes, p. 49: Cardium
verrucosum. Andrz. 1835. - Replaced by C. podolicum.
Desh.
verrucosum Deshayes, 1829. - Changed combination:
Freneixicardia verrucosum (Deshayes, 1829). OTHER
JUDGEMENTS: Cardium asperulum Brongniart. »
Regarding Cardium verrucosum: Cardiidae. Eocene of
France. Eocene of Egypt, and Rupelian (Lower
Oligocene) of Italy. RECORDS . "1829 Deshayes :
Cardium verrucosum. Lutetian (Middle Eocene) (fide
Schneider 2002). "1835 Lamarck: Cardium
verrucosum Desh . = C. asperulum Brongniart non
Lamarck. "1857. Deshayes, p. 49: Cardium
verrucosum Deshayes, 1830 sic. "1886 Cossmann :
Cardium verrucosum Deshayes. "1922-32 Sherbom,
p. 6865 : Cardium verrucosum Deshayes, 1829. "1977
Popov: Bucardium (Vepricardium) verrucosum.
Bucardium (Agnocardia) verrucosum (Deshayes ,
1829) ... 1995 Le Renard et al. .' Vepricardium (s. str.)
verrucosum [Cardium verrucosumJ Deshayes, 1829.
"1999 Jacques Vidal (pers. com.) MNHN, Paris:
Fossils from the Paleontological Collection .
Trachycardium verrucosum Deshayes. Lutetien. France
(Bassin de Paris). "2002 Schneider: Freneixicardia
verrucosa (Deshayes, 1829).
verruculosum Deshayes. - Combination to be revised :
"Cardium " verruculosum Deshayes. Cardiidae. Fossil.
RECORDS. "1839 Anton: Cardium verruculosum
Deshayes.
vertebratum Jonas, 1844. - Vasticardium vertebra tum
(Jonas, 1844). OTHER JUDGEMENTS: Laevicardiumjlavum
Linnaeus auctorum. - Not a synonym. » Regarding
Cardium vertebratum: Cardiidae. Recent. HABITAT. 810 fathoms (Melvill & Standen 1906). DISTRIBUTION.
Australia (Grasset 1884); Red Sea, Australia (Hylleberg
1994). RECORDS . "1844 Jonas, p. 33: Cardium
vertebra tum. "1858 Adams & Adams, p. 454: Cardium
(Pectunculus) vertebratum Jonas ... 1867 Mitchell:
Cardium vertebratum Jonas . "1870 March: c.
vertebratum Jonas junior synonym of Cardium jlavum
L. "1872 Tryon : Cardium vertebratum Jonas. "1884
Grasset: Cardium vertebra tum . R v. "1890 Paetel:
Cardium vertebratum Jon. "1906 Melvill & Standen,
p. 837: Cardium (Trachycardium) vertebra tum Jonas.
"1922-32 Sherbom, p. 6873: Cardium vertebratum
Jonas, 1844. "1928 Melvill: Cardium vertebratum
Jonas ... 1977 Fischer-Piette: Cardium vertebra tum
Jonas = Laevicardium jlavum Linne auctorum. .. 1992
Oliver: Trachycardium vertebratum. "1997 Vidal:
Vasticardium vertebratum (Jonas, 1844). "1999 Vidal:
Vasticardium vertebratum (Jonas, 1844).
verveceum Loriol, 1868 . - Changed combination:
Protocardia verveceum Loriol , 1868. Cardiidae. Fossil.
RECORDS. "1980 Ruhoff: Cardium verveceum Loriol,
1868; ser. 4: 28, pI. 2, fig. 6. "1958 ZR: Protocardia
vervecea from the Cretaceous Arzier, Swiss Jura, a redescription [Ref. Roessinger 1958].
veselillovici Brusina . - New combination: Lymnocardium veselinovici (Brus.). Lymnocardiidae. Fossil.
RECORDS. "1987 Taktakishvili : Limnocardium
veselinovici Brus.
veslellse Staadt in Cossmann & Pissarro, 1913. Parvicardium veslense Staadt in Cossmann & Pissarro,
1913. Cardiidae. Fossil. RECORDS. "1992 Renard:
Cardium (Plagiocardium) veslense Staadt in C. & P.,
1913: pI. LXIII fig. 69-30. "1995 Le Renard et al.:
Parvicardium veslense [Cardium veslense] Staadt in
Cossmann & Pissarro, 1913 ... 1995 Pacaud & Renard:
Parvicardium veslense.
vesta Rigaux & Sauvage, 1868. - Combination to be
revised: "Cardium " vesta Rigaux & Sauvage, 1868.
Cardiidae. FossiL RECORDS. "1980 Ruhoff: Cardium
vesta Rigaux & Sauvage, 1868, vol. 3: 74, pI. 6, figs.
15-16.
vetustum Papaianopol, 1984. - Dacicardium vetustum
Papaianopol, 1984. Lymnocardiidae. Upper Miocene.
RECORDS. "1984 ZR: Dacicardium vetustum sp. nov.
Romania. Ref. to Papaianopol p. 75 .
vexatum - Conocardium vexatum. Conocardiidae. Fossil.
RECORDS. "1889 ZR: Conocardium vexatum with fig.
[Ref. Barrois].
viatorium Olsson, 1964. - Laevicardium viatorium
Olsson, 1964. Cardiidae. Neogene. RECORDS. "1964
ZR: Laevicardium viatorium Olsson, 1964 sp. nov.
Esmeraldas formation , Neogene, Punta Gorda,
Ecuador.
vicaryi Palmer, 1973 . - Protocardia vicaryi Palmer,
1973. Cardiidae. Lower Cretaceous. RECORDS. "1973
ZR: Protocardia vicaryi Palmer, 1973 sp. nov. England,
Devon, Haldon Hills p. 176. Ref. to Palmer.
vicillum Fuchs, 1870. - Euxinicardium vicinum (Fuchs,
1870). Lymnocardiidae. Fossil. RECORDS. "1987
Anaya: Euxinicardium vicinum Fuchs. "1987
Taktakishvili: Limnocardium vicinum (Fuchs). "1991
Muller & Magyar: Lymnocardium vicinum (Fuchs
1870).
vicksburgellse Conrad, 1847. - Combination to be
revised: "Cardium" vicksburgense Conrad, 1847.
Cardiidae. Fossil. RECORDS ... 1857 Deshayes: Cardium
vicksburgensis. "1922-32 Sherbom, p. 6896: Cardium
vicksburgense Conrad, 1847. "1982 ZR: Cardium
vicksburgense Conrad 1848. Neotype designated. Ref.
830
Tropical Marine Mollusc Programme (TMMP)
victor - villeneuvianum
to Dockery, p. 74.
victor Angas, 1872. [Figs. Part 3 p. 904]. - Changed
combination: Cten ocardia victor (Angas, 1872).
Cardiidae. Recent. TYPE LOCA LITY. Mauritius. TYPE
REPOSITORY. Holotype, BMNH reg. no. 1872.5.18.3
(Wilson & Stevenson 1977). RECORDS. --1872 Angas,
p. 612, pI. 42, fig . 9: Cardium victor. --1872 ZR:
Cardium (Ctenocardia) victor sp. n. Angas, 1872.
Mauritius. --1880 Martens: Cardium victor Angas. The
lunule is conspicuously different in the Rand L valves.
The species stands rather isolated among cardiids, is,
however, somewhat related to C. multispinosum Sow.,
fimbriatum Wood and asiaticum Desh. --1890 Paetel:
Hemicardium(Fragum) victor Ang . "1904 Smith:
Cardium (Ctenocardia) victor . .. 1909 Melvill: Cardium
(Ctenocardia) victor Angas. --1932 Prashad: Cardium
(Ctenocardia) victor. "1937 Viader: Cardium victor,
Angas. --1946 Abrard: Cardium (Ctenocardium) victor
Angas. --1977 Fischer-Piette: Corculum (Ctenocardia)
victor Angas. --1977 Wilson & Stevenson: "Cardium"
victor Angas, 1872; must remain in doubt. --1977
Wilson & Stevenson: Cardium (Ctenocardia) victor
Angas, 1872. --1988 Drivas & Jay: Ctenocardia victor
(Angas, 1872). --1990 Abbott & Dance: Ctenocardia
victor. --1991 Matsukuma, Okutani & Habe :
Ctenocardia victor (Angas, 1872). --1992 Lamprell &
Whitehead : Ctenocardia (Ctenocardia) victor. "1999
Higo et al.: Ctenocardia victor (Angas, 1872). --2001
Vidal, p. 408: Ctenocardia victor. - CHARACTERS
AND OCCURRENCE - CHARACTERS of diagnostic
value: shell squarish, inflated, 41-52 strong ribs armed
with long pointed dorsally recurved spines which are
longer at the keel; interstices and intermediate ribs with
fine arcuate sculpture. DESCRIPTIONS. --1977 Wilson &
Stevenson: shell quadrate, umbo approximately central,
touching, prosogyrate. Angled umbonal keel marks
posterior slope, anterior and ventral margins rounded,
posterior margin nearly straight with pointed serration,
anterior and ventral margins with truncated serration.
Lunule ovate with 2 spinose ribs in each valve ;
escutcheon long, narrow, slightly wider in right valve,
single spinose rib in each valve. fibS 48 (41-52)
Anterior to umbonal keel, each 2n or 3 rd rib bears
long, pointed spines which curve dorsally. At anterior
end spines thicker, shorter and dorso-ventrally flattened
at their ends. Longest spines on ribs along the keel.
Ribs on posterior slope with short, fine, pointed spines.
Interstices and spineless median ribs with numerous
thin and fragile concentric lamellae, some of which
run up and coalesce on sides of spines. Hinge line
sinuous. Laterals approximately equidistant from umbo.
LV; anterior cardinal very strong, long pointed and
curved dorsally, separated by deep cleft from the weak,
low and pointed posterior cardinal. Laterals: One large
anterior and one + one, weak posterior teeth . RV;
anterior cardinal small and joined by saddle with large,
pointed, dorsally curved posterior cardinal. Anterior
laterals: one + one, ventral tooth large, truncated, curved
from shell cavity; dorsal tooth weak. One large posterior
dorsal tooth. --200 I Vidal : only the exhalant aperture
is tube-shaped. Inhalant siphon open towards the foot
aperture. No eyes. DIMENSIONS. --1988 Drivas & Jay:
ant.-post. 42.5 mm, umbo-ventr. 57 mm. REMARKS.
"1909: Melvill obtained a large and fine specimen,
peculiarly oblique in form, which might be
characterised as vaT. pulcherrima. HABITAT. --1904
Smith: 3 fathoms , off south coast of Ceylon. --1988
Drivas & Jay: Reunion, western Indian Ocean, in 6080 m depth. --1909 Melvill : 28-30 fathoms, coral-rockrubbl e, broken shell, to a depth of 78 fathoms. --1999
Higo et al.: 10-100 m, sand. DISTRIBUTION. "1994
Hylleberg: East Africa, Japan, Australia. --1999 Higo
et at.: Okinawa islands and southwards, Philippines,
southeast Asia, northern Australia.
victor pulcherrima Melvill, 1909 (variety). No evidence
of sub specific status. Identical with Ctenocardia victor
(Angas, 1872). » Regarding Cardium victor var.
pulcherrima: Cardiidae. Recent. TYPE LOCALITY.
Providence, western Indian Ocean, 50-78 fathoms
(Melvill 1909). REMARK S. Melvill(1909) obtained a
large and fine specimen, peculiarly oblique in form,
which might be characterised as var. pulcherrima.
RECORDS. "1909 Melvill , p. 135: Cardium (Ctenocardia) victor Angas vaT. pulcherrima.
victoriae Tate, 1887. - Combination to be revised:
"Cardium " victoriae Tate. Cardiidae. Eocene. RECORDS.
--1897 Harris: Cardium viCloriae Tate, 1887. --1903
Dennant & Clark : Cardium victoriae Tate.
vidali Cossmann & Peyrot , 191 I . - Changed
combination : Acanthocardia vidali (Cossmann &
Peyrot, 1911). Cardiidae. Neogene. RECORDS. --1911
ZR: Cardium vidali Cossmann & Peyrot, 1911 sp. n.
France. Neogene. --1977 Popov: Acanthocardia
(Acanthocardia) vidali (Cossmann & Peyrot, 1911).
vidali Poorten & Dekker, 2002. - Vepricardium vidali
Poorten & Dekker, 2002. Cardiidae. Recent. RECORDS.
--2002 Poorten & Dekker: Vepricardium vidali spec.
nov.
viennensis Papp, 1953 . - Monodacna viennensis Papp,
1953. Lymnocardiidae. Fossil. RECORDS. "1956 ZR:
Monodacna viennensis Papp, 1953 sp. nov.
Vietnamicardium Vu-Khuc, 1977 (genus). - Synonym
of Tulongocardium. » Regarding Vietnamicardium:
Cardiidae. Fossil. RE CORDS. --1980 Vokes :
Vietnamicardium Vu-khuc, 1977. Fossil. "1977 ZR:
Vietnamicardium gen. nov. ofCardiidae. Type species
Cardium vietnamicllm Vu-Huc 1965, Upper Triassic.
Ref. to Vu-Huc , D. p . 678 . --1995 Schneider :
Vietnamicardium Scott, 1978 Cretaceous. --1995
Schneider: Vietnamicardium synonymized with
Tulongocardillm since there are no autopomorphies
(characters unique to a taxon) to distinguish the two
genera. "1995 Schneider: Vietn amicardium type
species V vietnamicum Vu-Khuc in Vu-Khuc et ai. 1965
vietnamicum Vu-Huc, 1965. - Vietnamicardium
vietnamicum (Vu-Huc, 1965). Cardiidae. Upper
Triassic. RECORDS. --1976 Ma et al. p. 256: Cardium
vietnamicum Vukhuc, 1965. --1977 ZR : Cardium
vielnamicum Vu-Huc, 1965. Designated type species
for new genus Vietnamicardium. Ref to Vu-Huc, D. p.
678 . --1995 Schneider: Vietnamicardium type species
V vietnamicum Vu-Khuc in Vu-Khuc et al. 1965.
vigeni 8agdasaryan, 1979. - Sarmaticardium vigeni
Bagdasaryan, 1979. Cardiacea. Upper Miocene .
RECORDS. --1979 ZR : Sarmaticardium vigeni
Bagdasaryan, 1979. Sp. nov. Ukraine . Ref. to
Bagdasaryan, p. 41 .
villeneuvianum Matheron, 1843. - Combination to be
Phuket Marine Biological Center Special Publication vol. 30 (2004)
831
villicum - virgineum
revised: "Cardium" villeneuvianum Matheron, 1843.
Cardiidae. Cretaceous. RECORDS. 001922-32 Sherborn,
p. 6903: Cardium villeneuvianum Matheron, 1843.
001999 Jacques Vidal (pers. com.) MNHN, Paris:
Fossils from the Paleontological Collection. Cardium
villeneuvianum Matheron. Senonien. France, S09263,
-4, B17626.
villicum Papaianopol, 1982. - New combination:
Lymnocardium villicum Papaianopol, 1982.
Lymnocardiidae. Upper Miocene. TYPE LOCALITY.
Ponti en et Dacien. Basin Dacique, Romania. RECORDS.
001982 Papaianopol, p. 194, fig.: Limnocardium
(Euxinicardium) villicum sp. nov.
villmarense d'Archiac & Verneuil, 1842. - Changed
combination: Conocardium villmarense d'Orbigny,
1850. Conocardiidae. Fossil. RECORDS. 001922-32
Sherborn, p. 6904: Cardium villmarense d'Archiac &
Verneuil, 1842. 001922-32 Sherborn, p. 6904:
Conocardium (A & V) villmarense d'Orbigny, 1850.-
Cardium villmarense.
viminea (mode). - See under vimineum Martyn.
vimineum Martyn, 1784. Non binominal. - Not
available. RECORDS. 001784 Martyn, p. 50, pI. 47, fig.
3: Cardium viminea. 001858 Adams & Adams, p. 455:
Cardium (Trachycardium) vimineum Mart. 001869
Dunker, p. 112: Cardium virmineum sic (fide FischerPiette 1977). 001890 Paetel: Cardium vimineum Mrts.
001977 Fischer-Piette, p. 155: Cardium virmineum sic
= species insufficiently known. - CHARACTERS
AND OCCURRENCE. - DISTRIBUTION. China.
vincenti Cossmann, 1891. - Combination to be revised:
"Cardium" vincenti Cossmann, 1891. Cardiidae.
Oligocene. RECORDS. 001891 ZR: Cardium vincenti.
Oligocene of Etampes [Ref. Cossmann 1891].
vinculum Papaianopol, 1982. - New combination:
Lymnocardium vinculum Papaianopol, 1982.
Lymnocardiidae. Upper Miocene. TYPE LOCALITY.
Pontien et Dacien. Basin Dacique, Romania. RECORDS.
001982 Papaianopol, p. 183, fig.: Limnocardium
(Euxinicardium) vinculum sp. nov.
vindinense d'Orbigny, 1844. - Combination to be
revised: "Cardium" vindinense d'Orbigny, 1844.
Cardiidae. Fossil. RECORDS. 001871 Stoliczka, 30: C.
vindinense (Acanthocardium sic). 001922-32 Sherborn,
p. 6911: Cardium vindinense d'Orbigny, 1844.
vindobonense. - Homonymy to be noticed. Cardium
vindobonense from 3 sources: Hoernes in Partsch, 1848.
- Schwetz, 1913. - Possibly Laskarev (?year).
vindobonense Hoernes in Partsch, 1848. - Combination
to be revised: "Cardium" vindobonense Partsch.
Cardiidae. Miocene. RECORDS. 001922-32 Sherborn, p.
6911: Cardium vindobonense M. Hoernes (ex Partsch
MS), 1848. 001965 ZR: Cardium vindobonense figd.
Sarmatian, Yugoslavia [Ref. Spajic 1965]. 001999
Jacques Vidal (pers. com.) MNHN, Paris: Fossils from
the Paleontological Collection. Cardium vindobonense
Partsch. Austria; former Czechoslovakia. B31085,
B31396.
vindobonensis Laskarev. - Changed combination:
Cerastoderma vindobonensis (Laskarev). Cardiidae.
Miocene. RECORDS. 001998 Schultz: Cerastoderma
vindobonensis vindobonensis (Laskarev).
vindobonense Schwetz, 1913. - Combination to be
revised: "Cardium" vindobonense Schwetz, 1913.
Cardiidae. Tertiary. RECORDS. 001913 ZR: Cardium
vindobonense Schwetz, 1913 sp. n. Tertiary, Kertsch.
vindobonense breviformis Papp, 1954 - Combination
to be revised: "Cardium" vindobonense brevi/armis
Papp, 1954. Cardiidae. Sarmatian. REcORDs. 001949 ZR:
Cardium vindobonense brevi/armis 'no sp.' [nomen
nud.] [Ref. Papp 1949]. 001954 ZR: Cardium
vindobonense brevi/ormis Papp, 1954 subsp. nov.
Sarmatian, Vienna. 001965 ZR: Cardium vindobonense
brevi/armis figd. Sarmatian, Yugoslavia [Ref. Spajic
1965].
vindobonense jekeliusi Papp, 1954. - Combination to
be revised: "Cardium" vindobonense jekeliusi Papp,
1954. Cardiidae. Sarmatian. RECORDS. 001949 ZR:
Cardium vindobonensejekeliusi 'no sp.' [nomen nud.]
[Ref. Papp 1949]. 001954 ZR: Cardium vindobonense
jekeliusi Papp, 1954 subsp. nov. Sarmatian, Vienna.
001965 ZR: Cardium vindobonense jekeliusi figd.
Sarmatian, Yugoslavia [Ref. Spajic 1965].
violacea Martyn, 1784. Non binominal. - Not available.
Not a cardiid. RECORDS. 001784 Martyn: Cardium
violacea.
viquesne/i Pavlovic, 1929. - New combination:
Lymnocardium viquesneli Pavlovic, 1929.
Lymnocardiidae. Pontian. RECORDS. 001928 ZR:
Limnocardium viquesneli [Ref. Pavlovitch 1928].
001929 ZR: Limnocardium viquesneli Pavlovic, 1929
sp. n. Pontian Serbia.
virescens Gmelin, 1791. - Not a cardiid. REcORDs. 001742
Gualtieri, pI. 84, fig. A: Concha cordiformis. 001742
Gualtieri. ID by Gmelin 1791: Cardium virescens t.
84. f. A. 001791 Gmelin, p. 3256, No. 48.: Cardium
virescens. 001815 Wood: Cardium virescens L. 001815
Wood: Cardium virescens Linn. Gmel., inside silverish.
001817 Dillwyn: Cardium virescens Gme1in should be
ignored. 001828 Wood: Cardium virescens Gmelin.
001856 Wood & Hanley, No. 46: Modiola virescens
Gmel. 3256, from Gualt. 84, f. A. (= M. marmorata,
Forbes). Execrable copy. 001902 Sherborn, p. 1050:
Cardium virescens Gmelin, 1790 sic. 001977 FischerPiette: Cardium virescens Gmelin = species
insufficiently known. - CHARACTERS AND
OCCURRENCE - REMARKS. 001977 Fischer-Piette:
Gmelin referred to Gualtieri, but the illustrated species
has silvery interior surface. This character was also
mentioned by Wood. 001856. Wood & Hanley: since
silvery coatings do not occur in cardiids, the species
virescens must belong to another family. DISTRIBUTION.
Britain &c.
virgatum Papaianopol, 1982. - New combination:
Lymnocardium virgatum Papaianopo1, 1982.
Lymnocardiidae. Pliocene. TYPE LOCALITY. Pontien et
Dacien. Basin Dacique, Romania. RECORDS. 001982
Papaianopol, p. 197, fig.: Limnocardium (Euxinicardium) virgatum sp. nov.
virgineum Linne, 1758. - Fulvia aperta (Bruguiere,
1789). OTHER JUDGEMENTS: Fulvia bullata (Linnaeus,
1758). - Cerastoderma glaucum (Bruguiere, 1789). Dubious species. - Genus Mactra. » Regarding
Cardium virgineum: Recent. TYPE LOCALITY.
Mediterranean (Smith 1945). TYPE REPOSITORY.
Unknown. RECORDS. 001758 Linne, p. 682, No. 81:
Cardium virgineum. 001767 Linne, p. 1124, No. 93:
Cardium virgineum Linne. 001786 Schroter, No. XXI:
832
Tropical Marine Mollusc Programme (TMMP)
virgineum beta - virile
Cardium virgineum. Linn. XII. pag. 1124. Spec. 93.
"1791 Gmelin, p. 3253, No. 25: Cardium virgineum
Linne. ··1789 Bruguiere, dubious species, p. 235.
Cardium virgineum Linn. pag. 1124, num. 93. ··1815
Wood: Cardium virgineum L. is a doubtful species
completely confused by Gmelin. ··1815 Wood: C.
virgineum of Linne agrees with Cardium glaucum,
except that longitudinal ribs are wanting in virgineum.
··1817 Dillwyn: Cardium virgineum Linnaeus =
Cardium glaucum Poiret. ··1850 Jay: Cardium
virgineum Gmel. = Cardium ruga tum Gron. ··1855
Han ley: Cardium virgineum Linne. ··1869 Romer:
Cardium virgineum L. ed. X, N. 81, is a Mactra. ··1902
Sherborn, p. 1052: Cardium virgineum Linnaeus, 1758.
··1926 ZR: Cardium virgineum Lin. [Ref. Salisbury &
Woodward 1926]. ··1945 Smith: Cardium virgineum
Linne, 1758; Gmelin 1790, p. 3253 syn. = Fulvia
bullata Linne. ··1952 Dodge: Cardium virgineum
Linne. ··1977 Fischer-Piette: Cardium virgineum
Linne. ··1994 Vidal: ?Cardium virgineum Linne, 1758
= Fulvia (Fulvia) aperta (Bruguiere, 1789). CHARACTERS AND OCCURRENCE - REMARKS.
··1977 Fischer-Piette: Dodge (1952) stated that the
name virgineum cannot be retained owing to the
vagueness of its diagnosis, but "that the shell known
as Cardium apertum (Fulvia aperta) is probably the
species which Linnaeus described, and that the latter
is not a synonym of Solen bullatus Linne".
virgineum beta Gmelin, 1791 (variety). [Figs. Part I p.
135]. - Synonym of Fulvia aperta (Bruguiere, 1789).
OTHER JUDGEMENTS: Fulvia bullata (Linnaeus, 1758).Cardium ruga tum Gronovius, 1781 [not available].»
Regarding Cardium virgineum var. B: Cardiidae.
Recent. TYPE LOCALITY. Indian and American Oceans.
RECORDS. ··1781 Gronovius ZOop. t. 18. f. 5. ID by
Gmelin 1791: Cardium virgineum var. beta. ··1782
Chemnitz. ID by Gmelin 1791: Cardium virgineum var.
beta t. 18. f. 181. "1791 Gmelin No. 25: Cardium
virgineum var. B. ··1817 Dillwyn: Cardium virgineum
var. beta Gmelin = Cardium ruga tum Gronovius.
··1839 Anton: Cardium virgineum var. b. Gmelin =
Cardium apertum Chemnitz. ··1844 Reeve: Cardium
virgineum var B. Gmelin, 1791 = Cardium ruga tum
Gronovius, 1781. ··1874 Lischke: Cardium virgineum
var. beta. Gmelin = Cardium ruga tum Meuschen, 1781.
··1945 Smith: Cardium virgineum var. beta Chemnitz,
1782 syn. = Fulvia bullata Linne. ··1977 Wilson &
Stevenson: Cardium virgineum var. B. Gmelin, 1791 =
Fulvia aperta (Bruguiere, 1789). ··1979 Richardson
et al.: Cardium virgineum var. B Gmelin, 1791 = Fulvia
aperta (Bruguiere, 1789).
virginian a Conrad, 1864. - Protocardia virginiana
Conrad, 1864. Cardiidae. Recent. RECORDS. ··1980
Ruhoff: Protocardia virginiana Conrad, 1864; p. 211.
virginianum Conrad, 1839. - Changed combination:
Planicardium virginianum (Conrad, 1839). Neogene.
Cardiidae. Type species of Planicardium Olsson
(1967). Classifications according to Schneider (2202):
Cardium (Cerastoderma) virginianum, section
Cerastoderma (Dall 1900, Mansfield 1932).
Cerastoderma virginianum (Gardner 1944).
Planicardium virginianum (Campbell et al. 1975.
Stanley 1986). Chesacardium virginianum (Ward
1992). RECORDS. ··1863 Conrad: Cardium
(Cerastoderma) virginianum Conrad, 1839. (fide
Schneider 2002). ··1922-32 Sherborn, p. 6930:
Cardium virginianum Conrad, 1838. "2002 Schneider:
Planicardium virginianum (Conrad, 1839).
virgo Reeve, 1845. [Figs. Part I p. 144]. - New name
and changed combination: Ctenocardia perornata
(Jredale, 1929). OTHER JUDGEMENTS: Cardium
Jornicatum Sowerby, 1840. » Regarding Cardium
virgo: TYPE LOCALITY. ··1999 Higo et ai.: none
designated. TYPE MATERIAL. ··1977 Wilson & Stevenson:
not located. Cardiidae. Recent. RECORDS. ··1845 Reeve:
Cardium virgo. ··1858 Adams & Adams, p. 459:
Hemicardia (Ctenocardia) virgo Reeve. ··1884
Grasset: Hemicardium virgo. Rv. ··1890 Paetel:
Hemicardium virgo Rv. ··1909 Lynge: Cardium virgo
Reeve, 1845 = Cardium Jornicatum Sowerby, 1840.
··1922-32 Sherborn, p. 6931: Cardium virgo Reeve,
1845. ··1977 Fischer-Piette, No. 142: Cardium virgo
Reeve = Cardium Jornicatum Sowerby. ··1977 Wilson
& Stevenson, p. 26: Cardium virgo Reeve, 1845; has
been renamed Ctenocardia perornata (lredale, 1929).
··1992 Lamprell & Whitehead: Ctenocardia
(Ctenocardia) virgo. ··1999 Higo et at.: Ctenocardia
virgo (Reeve, 1845). ··2003 Hylleberg & Kilburn, p.
177: Ctenocardium virgo (Reeve) = Ctenocardia
perornatum (Iredale, 1929). - CHARACTERS AND
OCCURRENCE - DESCRIPTIONS. ··1845 Reeve: shell
somewhat squarely heart-shaped, posteriorly concavely
angulated, radiately ribbed, ribs about 38 in number,
narrow, compressed, very closely scaled, middle ribs
with the scales more erect, interstices between the ribs
transversely elevately striated, pure snowy white
without, blood-red within. ··Of authors: shell rounded
squarish, thick, anterior end rounded, posterior slope
angulated, concave; 38 ribs very closely scaled, middle
ribs with the scales more erect, interstices transversely
striped. Colour white, internally and externally, but
yellow or rose colours may occur centrally and
posteriorly internally. Shell length to 35 mm. REMARKS.
··1845 Reeve: Care should be taken not to mistake this
very delicate and beautiful shell for a white variety of
the Cardium hystrix, to which it is most nearly allied.
··1977: Wilson & Stevenson: C. virgo Reeve and C.
Jornicata Sowerby are conspecific. However, the type
illustration and descriptions given by Reeve (1845) are
close to perornata Iredale (= C. imbricatum Sowerby),
and certainly do not match the species C. Jornicata
which is an easily identifiable species that should not
be confused with any other cockle. HABITAT. ··1999
Higo et al.: 10-80 m, mud and sand. DISTRIBUTION.
··1845 Reeve: unknown. ··1884 Grasset: China. ··1992
Lamprell & Whitehead: Northern to North Western
Australia. ··1994 Hylleberg: Indo-Pacific, Australia.
··1999 Higo et at.: Taiwan and southwards, Philippines,
South China Sea, Beibu gulf, Hainan, Thailand,
Australia.
virgulata Martyn, 1784. Non binominal. - Not available.
Not a cardiid. RECORDS. ··1784 Martyn: Cardium
virgulata.
virile Dall, 1898. - Changed combination:
Trachycardium virile (Dall, 1898). Cardiidae.
Oligocene. RECORDS. ··1898 Dall: Cardium
(Trachycardium) virile Dall, 1898 n. sp. ··1977 Vokes:
"Trachycardium" virile (Dall)
Phuket Marine Biological Center Special Publication vol. 30 (2004)
833
virmineum - v/amingi
virmineum (error). - Printer's mistake for vimineum
Martyn in Fischer-Piette 1977.
visterniceniensis Gillet, 1938. - New combination :
Lymnocardium visterniceniensis Gillet, 1938.
Lymnocardiidae. Miocene . RECORDS . --1939 ZR:
Limnocardium visterniceniensis Gillet, 1938 sp. n.
Sarmatian [Miocene] Roumania.
visternicense chisinau Simionescu & Barbu, 1940
(forma). - Upgraded. New combination: Lymnocardium visternicense chisinau Simionescu & Barbu,
1940. Cardiidae. Sarmatian. RECORDS . --1946 ZR:
Cardium visternicense n. f. chisinau Simionescu &
Barbu, 1940 Sarmatian fossil, Rumania.
viteLlinum Reeve, 1844. [Figs. Part I p. 130. Part 3 p.
930]. - Synonym of Laevicardium crassum (Gmelin,
1791). OTHER JUDGEMENTS: Cardium laevigatum
Pennant. - Cardium oblongllm Chemnitz [not
avai lable]. - Cardium norvegicltm Spengler, 1799. Laevicardium brasilianum (Lamarck). » Regarding
Cardium vitellinum : Cardiidae. Recent. TYPE
REPOSITORY. 3 syntypes. 1978126. BMNH London. TYPE
LOCALITY . Unknown (Reeve 1844). RECORDS. --1844
Reeve, p. 168: Cardium vitellinum . --1844 Reeve,
Conch. Icon. sp. 37: Cardium vitellinum. -- J 856 Wood
& Hanley: C. vitellinum Reeve I. Car. f. 37 = Cardium
laevigatum Penn. --1858 Adams & Adams, p. 457:
Laevicardium vitellinum, Reeve. --1867 Hidalgo:
Cardium vitellinum, Reeve Guv., 1844, fig . 37) =
Cardium oblongum Chemnitz. --1867 Weinkauff:
Cardium vitellinum Reeve = Cardium norvegicum
Spengler. --1868 ZR: Cardium vile/linum = Cardium
norvegicum (Spengl.) = Cardium laevigatum of
Pennant not Linne [Ref. Romer 1868]. --1869 Romer:
Cardium vitellinum Reeve fig. 37 = Cardium
norvegicum Spengler. --1888 Kobe\t: Cardium
vitellinum Reeve = Cardium norvegicum Spengler.
--1889 Martens: Cardium vitellinum Reeve (p. 13 : Kap
Verden, Bonavista, 86 m, coarse sand). --1890 Paetel:
Laevicardium vitellinum R v. --1912 Dautzenberg &
Fischer: Cardium vitellinum Reeve J 844 = Cardium
(Laevicardium) norvegicum Spengler 1799. --1917
Hidalgo: Cardium vitellinum Reeve = Cardium
norvegicum Spengler. --1922-32 Sherborn, p. 6954:
Cardium vitellinum Reeve, 1844. --1953 Olsson &
Harbison: Laevicardium vitellinum (Reeve, 1844).
--1958 Abbott: Laevicardium vitellinum Reeve (in
Olsson and Harbison 1953). --1969 Work :
Laevicardium vitellinum (Reeve, 1843) in Olsson &
Harbison 1953: 107 = Laevicardium laevigatum
(Linnaeus , 1758). --1977 Popov : Laevicardium
vitellinum (Reeve, ) 845). --) 977 Fischer-Piette:
Cardium vitellinum Reeve = Laevicardium brasilianum
Lamarck. --1977 Fischer-Piette: Laevicardium
vitellinum Rve in Hall & Standen 1907 = Laevicardium
crassum Gmelin. --1977 Fischer-Piette, p.12: Cardium
vitellinum Rve in Martens 1889 = Laevicardium
crassum Gmelin. - CHARACTERS AND OCCURRENCE - DESCRIPTIONS. --1844 Reeve, sp. 37: shell
obliquely ovate, rather thin, radiately striated; yellowish
white, copiously freckled with numerous small pinkishbrown spots, covered with a yellow epidermis; margins
crenulated within (Descriptions in Latin as well as in
English). REMARKS. Cardium vitellinum is less
ventricose than the related C. oviputamen and farther
distinguished by its different tinge and style of
colouring. DISTRIBUTION. --1844 Reeve: unknown. --Of
authors: South East USA, Brazil.
vitrea Eichwald, 1829. [Figs. Part 1 p. 122]. - Changed
combination: Adacna vitrea (Eichwald, 1829) .
Lymnocardiidae. Recent. TYPE LOCALITY. Caspian Sea.
RECORDS. --1829 Eichwald : Glycimeris vitrea. --1838
Eichwald: Adacna vitrea m. --1838 Eichwald:
Glycymeris vitrea Zool. spec. --1841 Eichwald:
Glycimeris vitrea Eichwald, 1829. --) 841 Eichwald:
Adacna vitrea. --1849 Middendorff: Pholadomya vitrea
Eichw. (fide Andrussoff 1903). --1858 Adams &
Adams, p. 460: Adacna vitrea, Eichw. --1866 Issei:
Adacna vitrea, Eichwald, 1842. --1876 ZR: Adacna
vitrea (Eichw.) [Ref. Grimm 1876]. --1876 ZR:
Monodacna vitrea Eichw.) [Ref. Vest 1876]. --1877
Grimm : Adacna vitrea Eichwald . --1890 Paetel :
Adacna vitrea Eichw. -- J 890 Woodward: Adacna
vitrea, Eichwald - Caspian, Aral. --1903 Andrussoff:
Glycimeris vitrea Eichwald, 1829. --1903 Andrussoff:
Eichwald, 1838 moved Glycimeris vitrea to genus
Adacna. --1922-32 Sherborn, p. 6956: Adacna vitrea
Eichwald, 1838. - Glycimeris vitrea Eichwald, 1829.
--1968 Logvinenko & Starobogatov: Hypanis vitrea
vitrea (Eichwald). --1974 Starobogatov: Hypanis vitrea
(Eichwald). --1978 ZR: Hypanis vitrea. Spawning.
Larval numbers in plankton, relationships, Caspian Sea.
Ref. to Galperina. - CHARACTERS AND OCCURRENCE - DISTRIBUTION. --1969 Nordsieck: Black Sea.
vitrea bergi Starobogatov, 1974. - Hypanis vitrea bergi
Starobogatov. Lymnocardiidae. Recent. TYPE LOCALITY.
Caspian Sea. RECORDS. --1974 Starobogatov, p. 245 ,
fig. 212: Hypanis vitrea bergi Starobogatov subsp. nov.
--1978 ZR: Hypanis vitrea bergi Starobogatov, 1978
subsp. nov. Aral Sea. Ref. to Starobogatov p. 245.
vitrea ellxinica Nevesskaya, 1963 (variety). - Adacna
vitrea euxinica Nevesskaya, 1963. Lymnocardiidae.
Quaternary. RECORDS. --1963 ZR: Adacna vitrea
euxinica Nevesskaya, J963 subsp. nov. Quaternary,
Black Sea basin. --1969 Nordsieck: Adacna vitrea
(Eichwald, 1829) var. euxinica Nevessk.
vitrea glabra Ostroumoff. - Hypanis vitrea glabra
(Ostroumoft). Lymnocardiidae. Recent. TYPE LOCALITY.
Caspian Sea. RECORDS. -- J 968 Logvinenko &
Starobogatov: Hypanis vitrea glabra (Ostroumoft).
--1974 Starobogatov: Hypanis vitrea glabra
(Ostroumoft).
vitrea minima auct. (variety). - Changed combination:
Hypanis minima minima (Ostroumoff, 1907). OTHER
JUDGEMENTS: Hypanis vitrea glabra (Ostroumoft). »
Regarding Adacna vitrea var. minima: Lymnocardiidae.
Recent. TYPE LOCALITY. Caspian Sea. RECORDS. --1968
Logvinenko & Starobogatov : Adacna vitrea var.
minima auct. non Ostroumoff, 1907 = Hypanis vitrea
glabra (Ostr.). --1974 Starobogatov: Adacna vitrea var.
minima auct. = Hypanis minima minima (Ostroumoff,
1907).
vittatllm (variety). - See tuberculatum var. villatllm
Brusina, 1865.
vlamingi Wilson & Stevenson, 1977. [Figs. Part 3 p.
871]. - Synonym of Acrosterigma impolitum (Sowerby,
1833). » Regarding Acrosterigma vlamingi:
Cardiidae. Recent. TYPE LOCALITY. South Passage, Shark
Bay, Western Australia. TYPE REPOSITORY . Holotype,
834
Tropical Marine Mollusc Programme (TMMP)
vlastoides - vulgaris
Western Australian Museum, 1029-66. TYPE MATERJAL.
Holotype in WAM 1029-66, a single left valve from
Shark Bay, Western Australia. Eight paratypes, all from
the central West coast of Western Australia, from
Abrolhos Islands to Shark Bay (Vidal 1999).
DIMENSIONS. Length 24 .2, height 27.4, width
(calculated) 16.6 mm (Vidal 1999). DISTRIBUTION. West
Australia (Hylleberg 1994). REcORDs. ··1977 Wilson
& Stevenson, p. 92-95, pI. 6, figs. 21-22, text fig. 16:
Acrosterigma vlamingi sp. nov. ··1978 ZR:
Acrosterigma vlamingi sp. nov. Western Australia. Ref.
to Wilson et al. p. 92, fig'd. ··1992 Lamprell &
Whitehead: Acrosterigma vlamingi. ··1999 Vidal:
Wilson & Stevenson 1977, p. 92, pI. 6, figs 21-22 :
Acrosterigma vlamingi = Acrosterigma impolitum
(Sowerby, 1833).
vlastoides Reed, 1915. -Shanina vias to ides Reed, 1915.
Cardiacea. Ordovician. ··1915 ZR: Shanina vias to ides
Reed, 1915 gen. & sp. nov. Ordovician Shan States.
vodopici Brusina 1902. - Changed combination:
Prosodacna vodopici (Brusina 1902). Lymnocardiidae.
Pannonian. REMARKS. ··1991 Muller & Magyar, p. 368:
Prosodacna vodopici (Brusina 1902).
voesendorfense Papp, 1951 . - New combination: Lymnocardium voesendorfense (Papp, 1951). Lymnocardiidae. Pannonian. REMARKS. The relationship with
Monodacna voesendorfensis Papp, 1953 remains to be
determined. RECORDS. ··1948 ZR: Limnocardium
voesendorfensis Papp, 1948 'no sp.' [nomen nud.].
··1951 ZR: Limnocardium voesendorfense Papp, 1951
sp. nov. Pannonian Vienna Basin.
voesendorfensis Papp, 1953. - Monodacna voesendorfensis Papp, 1953. Lymnocardiidae. Fossi I.
RECORDS. ··1956 ZR : Monodacna voesendorfensis
Papp, 1953 sp. nov.
vogdti Andrussow, 1902. - Combination to be revised:
"Cardium" vogdti Andrussow, 1902. Cardiidae.
Neogene. REMARKS. The relationship with "Cardium
vogdti" Andrusov, 1961 and 1963 remains to be
determined. RECORDS. ··1905 Dollfus, commenting
Andrussow 1902: Cardium vogdti n. sp. tend s towards
Cerastoderma. ··1961 ZR: Cardium vogdti Andrusov,
1961 "sp. nov."[in a table] Miocene Caspian region,
Russia. ··1963 ZR: Cardium vogdli Andrusov, 1963
sp. nov. Neogene, Russia.
vogeli. - Protocardium vogeli. Cardiidae . Upper
Cretaceous. RECORDS . ··1966 Rasmussen:
Protocardium vogeli (Faxe Lime, Denmark).
volarovid Androussov, 1923. - Apscheronia volarovici
Andrusov, 1963. Lymnocardiidae. Pliocene. RECORDS.
··1930 ZR: Apscheronia volarovici Androussov, 1923
sp. n. ··1963 ZR: Apscheronia volarovici Andrusov,
1963 sp. nov. Apsheron layers, Pliocene, Russia.
volaticum Papaianopol, 1982. - New combination:
Lymnocardium volaticum. Lymnocardiidae. Upper
Miocene. TYPE LOCALITY . Pontien et Dacien. Basin
Dacique, Romania. RECORDS. ··1982 Papaianopol, p.
1·93, fig.: Limnocardium (Euxinicardium) volaticum sp.
nov.
volgensis (subspecies). - See under catillus volgensis
Svitoc, 1967.
volhynicum Grishkevich, 1967. - Combination to be
revised: "Cardium" volhynicum Grishkevich, 1967.
Cardiidae. Upper Sarmatian. RECORDS. ··1967 ZR:
Cardium volhynicum Grishkevich, 1967 sp. nov. Upper
Sarmatian, Volhyno-Podolia, USSR.
voltz;; Leymerie , 1846. - Changed combination:
Pleuriocardia voltzi (Leymerie, 1846). Cardiidae .
Cretaceous. RECORDS. ··1922-32 Sherborn, p. 6975:
Cardium voltzii de Leymerie, 1841 [n.n.]. ··1922-32
Sherborn, p. 6975: Cardium voltzii de Leymerie, 1846.
··1978 Scott: Pleuriocardia (Pleuriocardia) voltzi
(Leymerie). ··1978 Scott: Cardium voltzi Leymerie.
··1999 Jacques Vidal (pers. com .)MNHN, Paris: Fossils
from the Paleontological Collection. Cardium voltzi
Leymerie. Aptien. France (Yonne), S07314, -15,
S06041 , R50326.
voskoboinikovi (spelling) . - See under voskobojnikovi.
voskobojnikovi Andrusov, 1964. - Changed combination: Pontalmyra voskobojnikovi (Andrusov, 1964).
Lymnocardiidae. Tertiary. RECORDS. ··1964 ZR:
Didacna voskobojnikovi Andrusov, 1964 sp . nov.
Tertiary, USSR. ··1987 Taktakishvili: Didacna
voskoboinikovi Andrus. - Pontalmyra voskoboinikovi
(Andrus.)
voskuili Healy & Lamprell, 1992. - Synonym of Fulvia
papyracea (Bruguiere, 1789). » Regarding Fulvia
voskuili: Cardiidae. Recent. DISTRIBUTION. North East
Australia. RECORDS. ··1992 Healy & Lamprell: Fulvia
voskuili. ··1994 Vidal: Fulvia voskuili Healey &
Lamprell, 1992 = Fulvia (Fulvia) papyracea
(Bruguiere, 1789).
vostreysi (subspecies). - See under laqueatum vostreysi
Ward, 1992.
vroomi Aartsen, Menkhorst, & Gittenberger, 1984. Parvicardium vroomi Aartsen, Menkhorst, &
Gittenberger, 1984. Cardiidae. Recent. REMARKS.
Parvicardium vroomi is related to Cardium transversale
Deshayes, 1854. RECORDS. ··1984 Aartsen, Menkhorst,
& Gittenberger: Parvicardium vroomi. ··1984 ZR:
Parvicardium vroomi nom. nov. for Cardium
transversale Deshayes, 1854 nec Roemer 1849. Ref.
to van Aartsen et al. ··1998 Macedo et al.:
Parvicardium vroomi van Aartsen, Menkhorst &
Gittenberger, 1984. ··2000 Aartsen & Goud :
Parvicardium vroomi van Aartsen , Menkhorst &
Gittenberger, 1984. Replacement name for Cardium
transversale Deshayes 1855. - CHARACTERS AND
OCCURRENCE - DISTRIBUTlON. Mediterranean,
Canary Islands (Aartsen, Menkhorst, & Gittenberger,
1984).
vulgare (mode). - See under vulgaris.
vulgare Chemnitz, 1782. Non binominal. - Not available.
Cardiidae. RECORDS. ··1782 Chemnitz, Tab. 19 Fig. 194:
Cardium vulgare, edule Lin.
vulgaris Costa, 1778. - Synonym of Cerastoderma edule
(Linne, 1758). » Regarding Cardium vulgaris:
Cardiidae. Recent. DISTRIBUTION . Unknown. RECORDS.
··1678 Lister, pI. 5, fig. 34: Cardium vulgaris = edule
(fide Spengler 1799). ··1778 da Costa: Cardium
vulgaris. ··1788 Schrater: Cardium vulgare. [Reference
to Chemnitz (1782) fig. 198). ··1803 Montagu :
Cardium vulgare Da Costa p. 180 = Cardium edule
Lin. ··1817 Dillwyn: Cardium vulgare da Costa =
Cardium edule Linnaeus. ··1856 Wood & Hanley: C.
vulgare Da Cos. = Cardium edule Lin. ··1859 Sowerby:
C. vulgare = Cardium edule, Linn. ··1867 Weinkauff:
Cardium vulgare Pennant, da Costa = Cardium edule
835
Phuket Marine Biological Center Special Publication vol. 30 (2004)
vulgaris - wamillatum
Linne. ··1886 Locard: Cardium vulgare, da Costa, 1787
Cardium edule, Linne, 1767. ··1888 Kobelt: Cardium
vulgare de Costa sic = Cardium edule Linne. ··1890
Paetel: Cardium vulgare D. Cost. = edule L. "1902
Sherborn, p. 1063: Cardium vulgare da Costa, 1778.
"1969 Nordsieck: Cerastoderma vulgaris (da Costa,
1778). ··1977 Fischer-Piette: Cardium vulgare da Costa
= Cardium edule Linne.
vulgaris Sinz. - Didacnomya vulgaris. Lymnocardiidae.
Fossil. RECORDS. ··1964 Davidaschvili & Kitovani:
Didacnomya vulgaris Sinz. ··1967 Ebersin:
Didacnomya vulgaris (Sinz.). "1977 Popov:
Didacnomya vulgaris.
vulgaris. - Lunulacardium vulgaris. Probably not in
Cardiidae. ?Lunulacardiidae. Fossil. RECORDS. ··192232 Sherborn, p. 648: Lunulacardium vulgaris [In index
only. Not located in text].
vulgaris. - Monodacna vulgaris. Lymnocardiidae.
Pliocene . RECORDS. ··1929 ZR : Monodacna
(Didacnomya) vulgaris Pliocene [Ref. Krestovnikov
1929] .
vulgaris Lister, 1678. Non binominal. - See under
vulgaris Costa, 1778.
vulgaris planior Krestovnikov, 1929 (variety). Upgraded. Monodacna vulgaris planior Krestovnikov,
1929. Lymnocardiidae. Pliocene. RECORDS. ··1929 ZR:
Monodacna (Didacnomya) vulgaris var. planior
Krestovnikov, 1929 nov. Pliocene S. Russia [Ref.
Krestovnikov 1929].
vulgaroides Chelidze, 1975. - Didacnomya vulgaroides
Chelidze, 1975 . Lymnocardiidae . Lower Pliocene.
RECORDS. ··1975 ZR : Didacnomya vulgaroides
Chelidze, 1975 sp. nov. Georgia SSR, Khobs region.
Ref. to Chelidze, p. 195.
vulgatum. - Synonym of Cerastoderma edule (Linne,
1758). Cardiidae . Fossil. RECORDS. "1844 Philippi:
Cardium vulgatum = Cardium edule L. [fossil] .
Vulgodacna Iredale, 1937 (genus). - Tridacnidae .
Recent. RECORDS. "1937 ZR: Vulgodacna Iredale, 1937
gen. n. Tridacnidae, type Tridacna maxima var.fossor
Hedley (Vulgodacna fossor). ··1999 Higo et al.:
Vulgodacna Iredale, 1937 = Flodacna (subgenus)
Iredale, 1937. Tridacnidae.
vulva Jousseaume, 1898. [Figs. Part 3 p. 852]. Synonym of Clinocardium buelowi (RoUe, 1896). »
Regarding Cardium vulva: Cardiidae. Recent. TYPE
LOCALITY. Japan. TYPE REPOSITORY. Holotype in the Paris
Museum. REMARKS . Clinocardium californiense
buelowi in Kira (1962) = Clinocardium californiense
(Deshayes, 1839) (Kuroda et al. 1971). DISTRIBUTION.
North Japan, Alaska. RECORDS. ··1898 Jousseaume, p.
81: Cardium vulva . ··1898 ZR: Cardium vulva
Jousseaume, 1898 n. sp. Japan. ··1952 Kuroda & Habe:
vulva Jousseaume = Clinocardium buelowi (Rolle sic).
··1971 Kuroda, Habe & Oyama: Cardium vulva
Jousseaume, 1898 = Clinocardium californiense
buellowi sic (Rolle). "1977 Fischer-Piette: Cardium
vulva Jousseaume = Corculum californiense Deshayes.
··1977 Habe : Cardium vulva Jousseaume, 1898 =
Clinocardium (Keenocardium) buellowi sic (Rolle,
1896). "1978 ZR: Cardium vulva Jousseaume, 1898.
New synonym of C. (Keenocardium) buellowi Rolle,
1896. Ref. to Habe, T. p. 194. ··1980 Keen: Cardium
vulva = Clinocardium buelowi (Rolle, 1896). ··1999
=
Higo et al.: Cardium vulva Jousseaume, 1898 =
Clinocardium (Keenocardium) buellowi (RoUe, 1896).
vutskitsi. - Prosodacna vutskitsi. Lymnocardiidae.
Pannonian. RECORDS. ··1966 ZR: Prosodacna vutskitsi,
Pannonian, Mecsek Mts. Yugoslavia, pI. [Ref. Bartha
1966].
vosendorfense (invalid spelling) . - See under voesendorfense .
W
wade; Stephenson, 1941. - Changed combination:
Pachycardium wadei Stephenson, 1941. Cardiidae.
Cretaceous . RECORDS. ··1946 ZR: Cardium
(Pachycardium) wadei Stephenson, 1941 Navarro
formations, Cretaceous, Texas, N. edge of Corsicana,
Nacatoch sand. "1947 ZR: Cardium (Pachycardium)
wadei Stephenson, 1941 sp. nov. Upper Cretaceous,
Navarro Group Texas.
wagnerianum (subspecies). - See under laevigatum
wagnerianum Olsson and Harbison 1953.
wai/andi Woodring, 1925. - Acrosterigma wai/andi
(Woodring, 1925). Cardiidae. Miocene. TYPE LOCALITY.
Bowden, Jamaica (Vidal 1999). RECORDS. ··1999 Vidal:
Acrosterigma wailandi (Woodring, 1925)
waiparana Freneix & Grant-Mackie , 1978. Schedocardia? waiparana Freneix & Grant-Mackie.
Cardiidae. Upper Cretaceous. RECORDS. ··1978 Freneix
& Mackie, p. 514, Figs. 25, 26: Schedocardia (?)
waiparana n. sp. ··1978 ZR: Schedocardia? waiparana
Freneix, 1978 sp. nov. New Zealand. Ref. to Freneix et
al. p. 54. ··1992 Schneider: Schedocardia? waiparana
Freneix & Grant-Mackie.
waitakiense Suter, 1907. - Changed combination:
Hedecardium waitakiense (Suter, 1907). Cardiidae.
Chattian (Middle Oligocene) . Marwick (1944)
designated C. waitakiense type species of subgenus
Hedecardium of genus Vepricardium . Upgraded to
generic level by authors. TYPE LOCALITY. South
Canterbury, New Zealand. RECORDS. ··1907 Suter:
Cardium (Trachycardium) waitakiense Suter, 1907.
"1907 ZR: Cardium (Trachycardium) waitakiense
Suter, 1907 n. sp. Oamaru series, New Zealand. ··1926
McClelland: Cardium wailakiense Suter, 1907 n. sp.
··1944 ZR: Cardium waitakiense Suter Oligocene,
designated genotype of Hedecardium by Marwick,
1944a. ··1969 Moore et al.: Vepricardium (Hedecardium) waitakiense (Suter, 1907). "1977 Popov:
Hedecardium waitakiense (Suter, 1907). Cardium
waitakiense Suter, 1907. Type of Hedecardium
Marwick, 1944 . "1978 Maxwell: Hedecardium
(Hedecardium) waitakiense (Suter, 1907). ··1998
Schneider: Hedecardium waitakiense (Suter, 1907).
Type species of Hedecardium. Cardiidae. Oligocene.
··2002 Schneider: Hedecardium (Hedecardium)
waitakiense (Suter, 1907).
waltonianum DaB, 1900. - Combination to be revised:
"Cardium" waltonianum Da1l, 1900. Cardiidae. Fossil.
RECORDS. ··1930 Stewart: C. waltonianum Dall. ··2002
Schneider: citing Dall (1900): Cardium (Cerasloderma)
waltonianum sp. nov. section Cerasloderma .
wamillatllm Furon, 1948. - Combination to be revised:
"Cardium" wamillatum Furon, 1948 . Cardiidae.
Eocene.
RE CORDS . "1950 ZR : Cardium
836
Tropical Marine Mollusc Programme (TMMP)
wapeti - whitleyi
(Trachycardium) wamillatum Furon, 1948 sp. nov.
Eocene, Togo.
wapeti Cox, 1961. - Protocardia wapeti Cox, 1961.
Cardiidae. Cretaceous. RE COR DS . ··1961 ZR:
Proto cardia wapeli Cox, 1961 sp. nov. Nanutarra
fonnation Western Australia, Cretaceous.
warrenana Meek & Hayden, 1860. - Tancredia
warrenana Meek & Hayden, 1860. Cardiacea. Jurassic.
RECORDS. ··1860 Meek & Hayden: Tan credia
warrenana Meek & Hayden, 1860.
wassoevitschi Ebersin, 1967 . - Didacnomya wassoevitschi Ebersin, 1967. Lymnocardiidae. Pliocene.
RECORDS. ··1967 Ebersin: Didacnomya wassoevitschi
sp. n. ··1967 ZR: Didacnomya wassoevilschi Eberzin,
1967 sp. nov. figs . Pliocene, USSR.
wateleti Oeshayes, 1858 . - Changed combination:
Nemocardium wateleli (Oeshayes, 1858). Cardiidae.
Eocene. RECORDS. ··1930 Stewart: C. wallonianum
Oall . ··1886 Cossmann : Prolocardillm wateleti
(Oeshayes). ··1899 Sacco, p. 57: Nemocardium wateleti
(Oeshayes) Eocene (also spelled walheleli by Sacco).
··1977 Popov: Nemocardium (Nemocardium) wate/eli
(Oeshayes, 1858). ··1980 Ruhoff: Cardium wateleli
Oeshayes, 1858; [1856-1865] vol. I: 570, pI. 54, figs .
16-18 ... 1995 Le Renard et al. .' Nemocardium (s. str.)
wateleti [Cardium wateleti] Oeshayes, 1858 ... 1995
Pacaud & Renard : Nemoeardium (Nemocardium)
wateleti . .. 1999 Jacques Vidal (pers. com.) MNHN,
Paris: Fossils from the Paleontological Collection .
Nemocardium wateleti Oeshayes. Ypn!sien. France
(Bassin de Paris), B52859 to -61.
watsoni Richards, 1947. - Combination to be revised:
"Cardium" watsoni Richards, 1947. ?Conocardiidae.
Permian. RECORDS. ··1947 ZR: Cardium watsoni
Richards, 1947 sp. nov. Coniacean Pennian Andes.
waylandi Woodring, 1925. - New combination:
Trachycardium waylandi (Woodring, 1925). Cardiidae.
Miocene. TYPE LOCALITY. Jamaica. RECORDS. ··1925
Woodring, p. 139, PI. 19, Figs. 3-4: Cardium
(Traehycardium) waylandi Woodring, 1925.
waynense (subspecies). - See under diversum waynense
Mansfield, 1940.
waynense Mansfield, 1940. Nemocardium waynense
Mansfield, 1940. Cardiidae. Middle Oligocene.
RECORDS. ··1954 Keen: Nemocardium (Nemoeardium)
waynense Mansfield, 1940.
weaveri Anderson & Martin, 1914. - Changed
combination: Nemocardium weaveri (Anderson &
Martin, 1914). Cardiidae. Lower Oligocene and
Cenozoic. RECORDS. ··1914 ZR: Cardium weaveri
Anderson & Martin, 1914 sp. n. Cenozoic California.
.. 1954 Keen: Nemoeardium (Nemocardium) weaveri
(Anderson & Martin, 1914). ··1977 Popov :
Nemocardium (Habecardium) weaveri (Anderson &
Martin, 1914).
weberi Beet, 1953 . - Corcu/um weberi Beet, 1953.
Cardiidae. Mio-Pliocene . RECORDS. ··1956 ZR :
Coreulum (Ctenocardia) weberi Beet, 1953 sp. nov.
Mio-Pliocene Waisioe, Buton Malay Arch.
weisenense Papp. - Synonym of "Cardium"
(Replidacna) aurelianum Jekelius. Cardiacea. Fossil.
RECORDS. ··1977 ZR : Cerastoderma (lnaequieostata)
weisenense (Papp) syn. nov. Cardium (Rep/idacna)
aurelianum Jekelius. Ref to Paramo nova p. 70 [Transl.
p.320].
welleri Stephenson, 1941. - Changed combination:
Granocardium welleri (Stephenson, 1941). Cardiidae.
Fossil. RECORDS. ··1947 ZR: Cardium (Ethmocardium)
welleri Stephenson, 1941 sp. nov. Upper Cretaceous,
Navarro Group Texas. ··1954 Keen: Cardium
(Ethmocardium) welleri Stephenson, 1941. ··1954
Keen : Granoeardium (Ethmocardium) welleri
(Stephenson, 1941).
wenonah Weller, 1907. - Changed combination: Pleuriocardia wenonah (Weller). Cardiidae. Cretaceous.
RECORDS. ··1954 Keen: Cardium wenonah Weller, 1907
(Criocardium wenonah) . "1978 Scott: Cardium
wenonah Weller. ··1978 Scott: Pleuriocardia (Dochmocardia) wenonah (Weller).
whitei Oall , 1900. - Changed combination:
Granocardium whitei Oall, 1900 (Meek & Hayden,
1857). OTHER JUDGEMENTS: Granoeardium speeiosum
Adams and Reeve. - Not a synonym of Cardium
speciosum Adams and Reeve. » Regarding Cardium
whitei: Cardiidae. Cretaceous. RECORDS. "1900 Oall:
Cardium whitei Oa11 not a synonym of Cardium
speciosum Adams and Reeve. ··1902 Oall: Cardium
whitei Dall type of subgenus Ethmocardium White,
1880. ··1954 Keen: Cardium whiteiOall, 1900. ··1954
Keen: Granocardium (Ethmocardium) whitei (Oall,
1900). ··1969 Moore et al.: C. whitei = Granocardium
(Ethmocardium) speciosum (Meek & Hayden, 1857).
··1970 Speden: Cardium speciosum Adams & Reeve
= Cardium (Ethmocardium) whitei Dall, 1900. ··1993
Jaillard et a/.: Granoeardium (Ethmocardium) whitei.
··1998 Schneider: Granocardium (Ethmocardium)
whitei (Oall, 1900) USNM 315529 (holotype).
whitei (aff.) Oall, 1900. - Synonym of ?Ethmocardium
ursaniense Landes (1940). » Regarding whitei aff.:
Cardiidae. Cretaceous. RECORDS. ··1970 Speden:
Granoeardium (Ethmocardium) aff. whitei (Oall).
Ethmoeardium ursaniense Landes (1940) may be
conspecific.
whitfieldi Weller.
- Changed
combination: Pleuriocardia whitfieldi (Weller).
Cardiidae. Cretaceous. RECORDS. ··1963 ZR: Cardium
whitfieldi, first record in Delaware, Upper Cretaceous
[Ref. Richards & Shapiro 1963]. ··1978 Scott: Cardium
whitfieldi Weller. ··1978 Scott: Pleuriocardia
(Pleuriocardia) whitfieldi (Weller).
Cardiidae
whitleyi Iredale, 1929. - Synonym of Fragum seruposum
(Oeshayes, 1855). OTHER JUDGEMENTS: Fragumfragum
(Linnaeus). » Regarding Fragum whitleyi: Cardiidae.
Recent. Type locality. Australia. TYPE REPOSITORY .
Australian Museum, Sydney. Syntypes C057817.
RECORDS. ··1929 Iredale, p. 264, pI. 30, fig. 14: Fragum
whitleyi sp. nov. ··1929 ZR: Fragum whitleyi lreda1e,
1929b sp. n. N. Queensland. ··1977 Fischer-Piette, No.
149: Cardium whitleyi Iredale = Coreulum fragum
Linne. ··1977 Wilson & Stevenson: Fragum whitleyi
Iredale, 1929. ··1992 Lamprell & Whitehead: Fragum
whitleyi Iredale, 1929 = Fragum (Fragum) scruposum
Oeshayes, 1855. - CHARACTERS AND OCCURRENCE - OISTRlBUTION. ··1994 Hylleberg: Red Sea,
Indo-Pacific. [Figs. Part 3 p. 908].
Tridacnidae
whitleyilredale, 1937. [Figs. Part 3 p. 908]. - Synonym
837
Phukel Marine Biological Center Special Publication vol. 30 (2004)
wiesenense - xiangyunensis
of Tridacna derasa (Roding, 1798). » Regarding
Tridacna whitleyi : Tridacnidae. Recent. RECORDS.
"1937 ZR: Persikima whitleyi Iredale, 1937 sp. n.
Tridacnidae, type, Middleton Reef, S. Pacific. 001963
ZR: Tridacna whitleyi found to house commensal
shrimps of the genus Archistus in the Swain Reefs,
Australia, [Ref. McMichael 1963]. 001999 Higo et al.:
Persikima whitleyi Iredale, 1937 = Tridacna (Flodacna)
derasa (Roding, 1798).
wiesenense Papp, 1954. - Combination to be revised:
"Cardium" wiesenense Papp, 1954. Cardiidae.
Sarmatian. RECORDS. 001954 ZR: Cardium wiesenense
Papp, 1954 sp. nov. Sarmatian, Vienna.
wilckensi Zinsmeister, 1984. - Lahillia wilckensi
Zinsmeister, 1984. Cardiidae. Upper Eocene. REcORDs.
001984 ZR: Lahillia wilckensi Zinsmeister, 1984 sp.
nov. Antarctica. Ref. to Zinsmeister. 001995 Schneider:
Lahillia wilckensi Zinsmeister, 1984. USNM 365500
(holotype).
wilcoxi DaB 1900. - Trigoniocardia wilcoxi DaU, 1900.
Cardiidae. Tertiary. RECORDS. 001900 Dall : Trigoniocardia wilcoxi. 001953 Olsson & Harbison: Trigoniocardia (Trigoniocardia) wilcoxi (DaU, 1900). 001958
Abbott: Trigoniocardia willcoxi Dall 1900. 001977
Vokes p. 177: Trigoniocardia wilcoxi Dall, 1900.
williami Fischer-Piette, 1977. - Synonym of
Microcardium panamense (Dall, 1908). » Regarding
Microcardium williami: Cardiidae. Recent. RECORDS.
001977 Fischer-Piette, p. 4: Microcardium williami new
name for Microcardium panamense (Dall, 1908).
001977 ZR: Microcardium williami Fischer-Piette, 1977
nom. nov. for Microcardium panamensis (Dall, 1908)
nec Cardium panamense Sowerby, 1833. Ref to
Fischer-Piette, E. p. 4. - CHARACTERS AND
OCCURRENCE - REMARKS. 001980: Keen claimed that
Microcardium williami is an unnecessary new name
for a supposed homonym: Pr%cardia panamensis
Dall, 1908, versus Cardium panamense Sowerby, 1833,
species that now fall not only in different genera but in
different subfamilies. HABITAT. 001971 Keen & Mclean:
333 m. DISTRIBUTION. Panama (Keen & McLean 1971).
wilmarense d' Archiac. Conocardium wilmarense
d' Archiac . Conocardiidae. Fossil. Original classification in Cardiidae. RECORDS . "1859 Chenu:
Conocardium wilmarense, d' Archiac.
wilsoni (subspecies). - See also under elongatum wilsoni
Voskuil & Onverwagt, 1991.
wilsoniVoskuil & Onverwagt, 1991. [Figs. Part 3 p. 872].
- Changed combination: Vasticardium wilsoni (Voskui I
& Onverwagt 1991). Cardiidae. Recent. TYPE LOCALITY.
Broome, western Australia. TYPE REPOSITORY. Holotype
NNM (No. 56583), Leiden, Holland; paratype No.4
(WAM No . 732-66) from Rosemary lsI. Western
Australia; paratype No. 12 (NNMNo. 56584) from Fiji;
paratype No. 13 (NNM No. 56585) from the Moluccas.
REMARKS. See also Vas/icardium elongatum wilsoni.
REcORDS. 001991 Voskuil & Onverwagt: Trachycardium
wilsoni. 001992 Lamprell & Whitehead: Acrosterigma
wilsoni. 001991 Voskuil & Onverwagt: Trachycardium
wilsoni. 001993 Vidal : Trachycardium wilsoni Voskuil
& Onverwagt, 1991 = Acrosterigma elongatum
(Bruguiere, 1789). 001993 Vidal: Acrosterigma wilsoni
(Voskuil & Onverwagt, 1991); downgraded to
subspecies A. elongatum wilsoni. "1999 Vidal:
Trachycardium wilsoni Voskuil & Onverwagt, 1991 =
Va sticardium elongatum (Bruguiere 1789). CHARACTERS AND OCCURRENCE - DISTRIBUTION. 001994 Hylleberg: North West Australia.
winkleri lukas Stevanovic, 1982. - New combination:
Lymnocardium winkleri lukas (Stevanovic, 1982).
Lymnocardiidae. Upper Miocene. RECORDS. 001982 ZR:
Limnocardium winkleri lukas Stevanovic, 1982 subsp.
nov. Yugoslavia. Ref. to Stevanovic, p. 128.
witchelli Lycett, 1863. - Combination to be revised:
"Cardium" witchelli Lycett, 1863. Cardiidae. Fossil.
RECORDS. 001980 Ruhoff: Cardium witchelli Lycett,
1863; p. 55, pI. 40, fig. 36.
wolfartiChenu. [Figs. Part I p. 152]. - Tridacna wolfarti
Chenu. Tridacnidae. Fossil. RECORDS. "1845 Chenu pI.
8 f. 5: Tridacna wolfarti Chenu fossil Lower Hessen,
Germany.
woodringi Loel & Corey, 1932. - New combination:
Trachycardium woodringi (Loel & Corey, 1932).
Cardiidae. Miocene. RECORDS. 001933 ZR: Cardium
(Trachy cardium) woodringi Loel & Corey, 1932
Miocene California [Ref. Loel & Corey 1932]. 001944
Keen & Bentson : Cardium (Trachycardium) woodringi
Loel & Corey, 1932 (n. sp.).
woodsi Marwick, 1944. - Ethmocardium woodsi
(Marwick, 1944) . Cardiidae. Upper Cretaceous.
RECORDS. 001944 ZR: Ethmocardium woodsi Marwick,
1944a sp. nov. Selinqu Rapids, Piripanan, Upper
Cretaceous, = Cardium sp. 2 Woods N.Z. Geol. Surv.
Pal. Bull. p. 38 pI. 18 f. 4, 5. "1954 Keen :
Ethmocardium woodsi Marwick, 1944. 001954 Keen:
Granocardium (Ethmocardium) woodsi (Marwick,
1944). 001978 Freneix & Mackie : Granocardium
(Ethmocardium) woodsi (Marwick). "1978 Freneix &
Mackie: Ethmocardium woodsi (Marwick 1944, p. 259,
pI. 36. fig. 21).
woolnoughi Bell, 1917. - Combination to be revised :
"Cardium " woolnoughi Bell, 1917. Cardiidae .
Pliocene. RECORDS. 001917 ZR: Cardium woolnoughi
Bell, 1917 n. sp. Pliocene England.
wop/neri Ludbrook, 1966. - Protocardia wop/neri
Ludbrook, 1966. Cardiidae. Albian. RECORDS. 001966
ZR: Protocardia wop/neri Ludbrook, 1966 sp. nov.
Albian, S. Australia.
x
xanthocheilum Carpenter, 1857. - Synonym of
Trachycardium quadragenarium (Conrad, 1837). »
Regarding Cardium xanthocheilum: Cardiidae. Recent.
RECORDS. 001857 Carpenter, (Gould ms. ) p. 232:
Cardium xanthocheilum. 001864 Carpenter: Cardium
xanthocheilum Gld. = Cardium quadragenarium Conr.
001872 Carpenter: Cardium xanthocheilum. 00190 I
Dall: Cardium xanthocheilum (Gould MS.) Carpenter,
1856 = Cardium quadragenarium Conrad, 1837 .
001977 Fischer-Piette: Cardium xanthocheilum
. Carpenter = Laevicardium quadragenarium Conrad.
002000 Coan et al.: C. xanthocheilum Carpenter, 1857
= Trachycardium (Dal/ocardia) quadragenarium
(Conrad, 1837).
xiangyunensis Chen, 1976. - Combination to be revised:
"Cardium" xiangyunensis Chen, 1976. Cardiidae.
Upper Triassic. TYPE LOCALI TY . Yunnan, China .
838
TropicaL Marine Mollusc Programme (TMMP)
yagatagensis - zavalui
RECORDS. ··1976 Ma et af. p. 256, pI. 23, figs. 5-10, 17:
Cardium xiangyunensis Chen, 1976 sp. nov.
Y
yagatagensis (error). - Printer's mistake for yakatagensis
in ZR for 1972 with ref to Clark, 1932.
Yagudinella Kafanov, 1975 (genus). - Cardiidae. Fossil.
RECORDs. ··1975 Kafanov: Yagudinella. ··1977 Vokes:
Yagudinella Kafanov, 1975. ··1980 Vokes: Yagudinella
Kafanov, 1975. ··1975 ZR: Yagudinella Kafanov, 1975
gen. nov. ofCardiidae. Type species Cardium (Serripes)
notabile Saverby (sic = Sowerby) 1915. Ref. to
Kafanov, p. 147. ··1980 ZR: Yagudinella Kafanov
1975. Key to species & subspecies. Ref. to Kafanov.
··1998 Schneider (l998b): Yagudinella. ··1999 Higo
et at.: Yagudinella Kafanov, 1975 = Serripes Gould,
1841.
Yagudinella Kafanov, 1975 (subgenus). - Cardiidae.
Fossil. RECORDS. ··2000 Coan et af.: Subgenus Yagudinella Kafanov, 1975.
yakatagensis Clark, 1932. - Changed combination:
Clinocardium yakatagensis (Clark, 1932). Cardiidae.
Upper Oligocene. RECORDS. ··1954 Keen: Cardium
(Cerastoderma) yakatagensis Clark, 1932. ··1972 ZR:
Clinocardium yagatagensis sic (Clark, 1932).
Combination nov. Transferred from Cardium
(Ceratoderma sic) [Ref. Addicott 1972]. ··1974 ZR:
Ciliatocardium yakatagense (Clark, 1932). Comb. nov.
Ref. to Kafanov, p. 1469.
yamasakii Makiyama, 1934. - Changed combination:
Clinocardium yamasakii (Makiyama, 1934). Cardiidae.
Oligocene. ?Miocene. REcORDs. ··1934 Makiyama:
Cardium yamasakii. ··1935 ZR: Cardium
(Cerastoderma) yamasakii Makiyama, 1934 sp. n.
Oligocene Japan. ··1973 Keen: Clinocardium yamasakii (Makiyama, 1934). ··1974 ZR: Ciliatocardium
yamasakii. Makiyama 1934. Comb. nov. Ref. to
Kafanov, p. 1469.
yatsushiroensis Hayami, 1965. - Nemocardium
yatsushiroensis Hayami, 1965. Cardiidae. Cretaceous.
RECORDS. ··1965 ZR: Nemocardium yatsushiroense
Hayami, 1965 sp. nov. Albian, Yatsushiro formation,
Kumamoto Pref. Japan. ··1973 Keen: Nemocardium
(Nemocardium) yatsushiroense Hayami, 1965 Lower
Cretaceous. ··1986 Scott: Nemocardium yatsushiroensis Hayami, 1965.
Yavorskielia Khalfin, 1950 (genus). - [Ambonychiacea,
not mentioned in the ZR]. RECORDS. ··1969 ZR:
Yavorskiella Khalfin, 1950 in subfamily Aenigmoconchiidae Betekhtina & Sukov, 1969 [Ref. Betekhtina
& Sukov. 1969]
ykatagense Clark, 1932. - Changed combination:
Clinocardium ykatagense (Clark, 1932). Cardiidae.
Fossil. RECORDS. ··1936 ZR: Clinocardium ykatagense
(Clark, 1932) [Ref. Keen 1936].
yokoyamai Cossmann, 1924. - Combination to be
revised: "Cardium" yokoyamai Cossmann, 1924.
Cardiidae. Pliocene. RECORDS. ··1924 ZR: Cardium
yokoyamai Cossmann, 1924 Pliocene. India.
yokoyamai Otuka, 1935. - Serripes yokoyamai Otuka,
1935. Cardiidae. Middle Miocene to Lower Pliocene.
RECORDS. ··1935 ZR: Serripes yokoyamai Otuka, 1935
sp. n. Miocene Japan. ··1973 Keen: Serripes yokoyamai
Otuka, 1935. [Middle Miocene to Lower Pliocene].
··1975 ZR: Serripes yokoyamai Otuka, (CHECK) 1935
Referred to Yagudinella. Ref. to Kafanov, p. 147. ··1975
ZR: Yagudinella yokoyamai (Otuka, 1935) comb. nov.
transferred from Serripes. Ref. to Kafanov, p. 147.
··1977 Popov: Serripes yokoyamai Otuka, 1935. ··1978
Kafanov: Yagudinella yokoyamai (Otuka 1935).
yokoyama; Takeda, 1953. - Nemocardium yokoyamai
Takeda, 1953. Cardiidae. Oligocene. RECORDS. ··1953
ZR: Nemocardium yokoyamai Takeda, 1953 sp. nov.
Oligocene, Japan. ··1954 Keen: Nemocardium
(Arctopratufum) yokoyamai Takeda, 1953. ··1960 ZR:
Nemocardium yokoyamai Takeda; lectotype designated
[Ref. Oyama et at. 1960]. ··1973 Keen: Nemocardium
(Nemocardium) yokoyamai Takeda, 1953. ··1977 Habe:
Arctopratufum yokoyamai Takeda, 1953.
yoshidense Kanno, 1958. - Laevicardium yoshidense
Kanno, 1958. Cardiidae. Upper Oligocene. REcORDs.
··1960 ZR: Laevicardium yoshidense Kanno, 1958 sp.
nov. p. 176, pI. 2, figs. 5-7; Nenokami, Hikokubo,
Yoshida-machi, Japan, Upper Oligocene. ··1973 Keen:
Laevicardium yoshidense Kanno, 1958.
yukatagensis Clark, 1932. - Combination to be revised:
"Cardium" yukatagensis Clark, 1932. Cardiidae.
Oligocene. RECORDS. ··1932 ZR: Cardium (Cerastoderma) yukatagensis Clark, 1932 sp. n. Oligocene S.
Alaska.
yunnanensis Chen, 1976. - Myophoricardium
yunnanensis Chen, 1976. Cardiidae. Fossil. RECORDS.
··1976 Ma et al. p. 260: Myophoricardium yunnanensis
Chen sp. nov.
Z
zaccagnai Regny, 1922. - Combination to be revised:
"Cardium" zaccagnai Regny, 1922. Cardiidae.
Cretaceous. REcoRDs. ··1922 ZR: Cardium zaccagnai
Vinassa de Regny, 1922 sp. n. Cretaceous Tripoli.
··1924 ZR: Cardium zaccagnai Vinassa de Regny, 1924
sp. n. Cretaceous Tripoli.
Zagrabicardium Basch, 1990 (genus). - Lymnocardiidae. Fossil. RECORDS. GENOTYPE. Basch (1990 p.
26) designated Cardium riegeli Homes, 1870 type
species of subgenus (Zagrabicardium) of genus
Limnocardium.
zagrabiense Brusina, 1874. - New combination:
Lymnocardium zagrabiense (Brusina, 1874). Lymnocardiidae. Fossil. RECORDS. ··1977 Popov: Limnocardium zagrabiense. ··1990 Basch: Limnocardium
(Hungarocardium) zagrabiense (Brusina, 1874).
Zamphiridacna Motas in Andreescu, 1974 (genus). Lymnocardiidae. Tertiary. RECORDS. ··1974 ZR:
Zamphiridacna Motas. Referred to new subfamily
Prosodacninae. Ref. to Andreescu, p. 128. ··1980
Vokes: Zamphiridacna Motas in Andreescu, 1974.
Subfamily uncertain. ··1980 ZR: Zamphiridacna Motas
1974. Key to species. Ref. to Andreescu.
zavalui Liteanu & Schoverth, 1964. - Horiodacna
zavafui Liteanu & Schoverth, 1964. Lymnocardiidae.
Fossil. REMARKS. Representatives of genus Horiodacna
Sabba from Dack in eastern Bulgaria. RECORDS. ··1964
ZR: Horiodacna zavafui Liteanu & Schoverth, 1964
sp. nov. Zavalu, Rumania, Dacian. ··1987 Lutskanov:
Horiodacna zavafui Liteanu & Schoverth, 1960. ··1987
Phukel Marine Biological Cenler Special Publication vol. 30 (2004)
839
zecki - Zllzica
Lutskanov: Horiodacna zavalui.
zecki Oppenheim, 1919. - Combination to be revised:
"Cardium" zecki Oppenheim, 1919. Cardi idae. Eocene.
RECORDS. ··1919 ZR: Cardium zecki Oppenheim, 1919
sp. n. Eocene, Togo.
zetes Lorio1, 1889. - Combination to be revised :
"Cardium " zetes Lorio1, 1889. Cardiidae. Fossil.
RECORDS. ··1889 ZR: Cardium zetes De Lorio1, 1889
n. sp. with fig .
zhidkovae Kafanov, 1994. - Ciliatocardium zhidkovae
Kafanov, 1994. Cardiidae. Fossil. RECORDS. ··1994
Kafanov in Arkhipova et al.: Ciliatocardium zhidkovae.
zhuzhunae Popov, 1982. - Cerastoderma zhuzhunae
Popov, 1982. Cardiidae. Oligocene. RECORDS. ··1982
ZR : Cerastoderma zhuzhunae Popov, 1982 sp. nov.
Georgia SSR. Ref. to Popov et al. p. 42.
zlatarskii Andrussow. - Pseudocatillus zlatarskii
(Andrussow). Lymnocardiidae. Fossil. REcORDs. ··1967
Ebersin : Pseudocatillus zlatarskii (Andrus.).
zlatarskii Wassoievic & Eberzin, 1930. - Monodacna
zlatarskii Wassoievic & Eberzin, 1930. Lymnocardiidae. Pliocene. RECORDS. ··1931 ZR: Monodacna
zlatarskii Wassoievic & Eberzin, 1930 sp. n. URSS
Pliocene.
zonariaeforme Schauroth, 1865. - Unicardium
zonariaeforme Schauroth, 1865. Cardiacea. Fossil.
REcORDs. ··1980 Ruhoff: Unicardium zonariaeforme
Schauroth, 1865; p. 130, pI. 3, fig. 11.
zonata (mode and variety). - See under zonatum Brown,
1844 and papillosum zonala Monterosato.
*zonatum. - Homonymy to be noticed. Cardium
zonatum from 2 sources: Brown, 1827. - Leach, 1852.
zonatllm Brown, 1827. - Synonym of Cerastoderma
edule (Linne, 1758). » Regarding Cardium zonatum:
Cardiidae. Recent. TYPE LOCALITY. Devonshire coast.
REMARKS. Brown (1844): May be distinguished from
the young of C. edule by being invariably much thinner,
and more inflated; smoother, and its colour of a
gamboge-yellow, and zoned. Its habitat is at the mouth
of rivers, where the water is brackish; and it never
reaches an inch in diameter. REcORDs. ··1827 Brow'a
Cardium zonatum Brown, First ed . ··1844 Brown, 2 n
ed. p. 88, pI. 35, fig. 8: Cardium zona/um. ··1852 Leach:
Cardium zona/um Brown. ··1859 Sowerby: C. zona/um
= Cardium edule, Linn. ··1867 Weinkauff: Cardium
zona/um Brown = Cardium edule Linne. ··1869 Romer:
Cardium zonatum Brown = Cardium edule L. ··1890
Paetel : Cardium zonatum Brown. = edule L. "1917
Hidalgo: Cardium zona/um Brown = Cardium edule
Linne. ··1922-32 Sherborn, p. 7049: Cardium zonatum
Brown, 1827. ··1977 Fischer-Piette: Cardium zona tum
Brown = Cardium edule Linne.
zonatum Leach, 1852. - Synonym of Parvicardium ovale
(Sowerby). » Regarding Cardiumzonatum: Cardiidae.
Recent. RECORDS. ··1852 Leach, p. 319: Cardium
zonatum Leach, 1852. ··1867 Weinkauff: Cardium
zona tum Leach = Cardium fasciatum Montagu. ··1888
Kobel!: Cardium zonatum Leach (1852) = Cardium
fasciatum Montagu.
zonatllm beta Leach, 1852 (variety). Infrasubspecific.
» Regarding Cardium zonatum var. beta: No evidence
of subspecific status. Parvicardium ovale (Sowerby).
Cardiidae. Recent. RECORDS. ··1852 Leach: Cardium
zonatum var. beta L.
zorritensis Spieker, 1922 . - New combination:
Trachycardium zorritensis (Spieker, (922). Cardiidae.
Tertiary. RECORDS. "1922 ZR : Cardium (Trachycardium) zorritensis Spieker, 1922 Tertiary, Peru .
zlljovici Brusina, 1897 . - New combination:
Lymnocardium zujovici (Brusina, (897) non Fuchs.
Lymnocardiidae. Ponti an. RECORDS. ··1953 ZR:
Limnocardium zujovici Brus . non Fuchs Ponti an,
Serbia. ··1979 ZR: Limnocardium zujovici Brusina,
1897. Neotype. Ref. to Stevanovic, p. 336.
Zllzica (variety).- See planicostata var. zuzica Chelidze,
1957.
840
Tropical Marine Mollusc Programme (TMMP)
These sentences are extracted from pages 388-389 in
G. Schenk & A.M. Keen J 940. (Ref in Part 1 p. 310)
We need ilJustrations of unfigured or little-known species.
Proper identification of specimens obviously must Qrecede all
atfempts to delimit the geographical distrioution of species.
Species must bear correct names if check lists are to be meaningful.
Taxonomic revisions of more species and genera should be
undertaken.
We need! in short, nomenclatural stability to prevent the continual
shutt ing of species from one generic allocation to another.
Nomenclatural sadists tend to keep the names of genera in perpetual
chaos.
Phuket Marine Biological Center Special Publication vol. 30 (2004)
841
O.A.L M0rch: an important cuartor of the Shell Collections ofthe University of Copenhagen
[Universitetsmuseets Conchyliesamlinger] and the Royal Natural History Museum
[Kg/. naturhistoriske Museum] .
In 1844 M0rch became amanuensis of Dr. Beck who was in charge of the shell collection of King
Christian 8, who reigned from 1839 to 1848 [many specimens shown on the present Plates have C VIII
written on the inside]. Thanks to the knowledge achieved by Morch regarding the old cardiids dating
back to Chemnitz (1782), Spengler (1799), and the King's Collection, scientists of present times are
able to trace these shells in the collection of the Zoological Museum the University of Copenhagen
(ZMUC).
Morch had great knowledge and he was critical with a good sense for shapes and variations. He was
also good at making order in collections. E.g., he wrote "Sp." in ink inside the shells when he had traced
the material Spengler worked with. Morch's labels are also an invaluable source of information. Ifit had
not been for Morch we would have had a hard time allocating much of the old material at ZMUC to the
right source of origin. M0rch, or Morch as his name also is spelled, lived from 1828 to 1875.
The picture of Otto Andreas Lowson M0rch is extracted from a biography written in Danish by Jonas
Collin: Konchyliologen O.A.L. M0rch. En biografisk Skizze (38 pp).
842
Tropical Marine Mollusc Programme (TMMP)
The Plates
I have taken photos of specimens deposited at museums of natural history in Austria (Vienna), Belgium
(Brussels), Denmark (Copenhagen) France (Paris), United Kingdom (London) and Thailand (Phuket
Marine Biological Center).
I addition to my own records the present Plates show species provided by courtesy of the following
people and institutions:
Australian Museum Sydney. Photographer of the
Museum by courtesy of Peter Middelfart.
Natural History Museum, London. BIW and colour
pictures. Photographer of the Museum by courtesy of Kathie Way & Amelia MacLellan.
Zoological Museum, Copenhagen. Photographer
of the Museum by courtesy of Tom Schi0tte.
Legend: abbreviated names ofphotographers
A. Ie G. = Amou Ie Goff for Museum National
d'Histoire Naturelle, Paris
G.D. = G. Dajoz for Museum d'Histoire Naturelle
Geneve. BIW pictures by courtesy of Godtfred
H0pner Petersen
G.H.P. = Godtfred H0pner Petersen, Zoological
Museum University of Copenhagen
H.T. = Harry Taylor for the Natural History Museum, London
J.H. = Jorgen Hylleberg, Marine Ecology, Aarhus
University
J.V = Jacques Vidal, Museum National d'Histoire
Naturelle, Paris
L.K. = Lisa Kirkendale for the Natural History
Museum, London
P.S.D. = P.S. Dance for the Linnean Society of
London
Cardium & Ringicardium
843
& Acanthocardia
Acanthocardia
844
845
Acanthocardia
Acanthocardia
846
847
Acanthocardia
Acanthocardia
848
849
Cerastoderma
Cerastoderma
850
851
Cerastoderma
Cerastoderma & ?Parvicardium &
852
Clinocardium & Ciliatocardium
853
Ciliatocardium & Serripes
Serripes & Trachycardium
854
855
Trachycardium
Dinocardium & Trigoniocardia
856
Trachycardium & Trigoniocardia
857
Dallocardia & Trachycardium
858
859
Vas ticardium
Vas ticardium
860
861
Vas ticardium
Vas ticardium
862
863
Vas ticardium
Vas ticardium
864
865
Vas ticardium
Vasticardium& Acrosterigma
866
867
Vas ticardium
Vas ticardium
868
869
Vas ticardium
Vas ticardium
870
871
Vasticardium & Acrosterigma
Vasticardium & Acrosterigma
872
873
Acrosterigma
Acrosterigma
874
875
Acrosterigma
Vasticardium & Acrosterigma
876
877
Acrosterigma
Acrosterigma
878
879
Acrosterigma
Acrosterigma & Fulvia
880
881
11
... _hi.
, ...
Acrosterigma & Vepricardium
Vepricardium
882
883
Vepricardium
Vepricardium
884
885
Vepricardium
Vepricardium
886
887
Europicardium & Fulvia
Fulvia & Microfragum
888
889
Fulvia
Fulvia & Parvicardium
890
Fulvia
Laevicardium & Acrosterigma
892
& Clinocardium
893
Laevicardium
Laevicardium
894
895
Laevicardium
Laevicardium
896
897
Laevicardium
Laevicardium
898
899
Lyrocardium
Nemocardium & Frigidocardium
900
& Pratulum
Microcardium, & Frigidocardium
901
& Plagiocardium
Plagiocardium & Ctenocardia
902
903
Ctenocardia
Ctenocardia & Americardia
904
セ@
セBM
M-f-Rvz
s..... "'7 j-lJ.p./r
r).,;..,,/ Mヲ・ッャjセ@
セ@
,s'- I{ 411. t..-..r
l'Jw-.rOY..J
,/,t, //,
Americardia & Fragum
905
Fragum & ? Americardia
906
907
Fragum
Fragum & Hemicardium
908
909
Hemicardium & Fragum
Hemicardium & Lunulicardia
910
911
Lunulicardia
Lunulicardia
912
913
Corculum
Corculum
914
915
Corculum & Parvicardium
Parvicardium & Papyridea
916
& Fulvia
Papyridea & Parvicardium
917
& Afrocardium
Afrocardium & Hemidonax
918
919
Cardium fasciatum Montagu. [= C. ovale Sowerby]
a & b. A specimen of the short, strongly ventricose form. F<eroes. 20-30 fm
c & d. A specimen of the somewhat elongated, much tumid form.
West Iceland (0nundarfjardur), ca. 12 fm.
e. A specimen with tubercles not only on the anterior and posterior area,
but also on the middle area. West Iceland (0nundarfjardur), ca. 12 fm.
f. That form which was confused by March with C. edule L. F<eroes.
g. The same half from in front.
h. The same half from behind.
i. A specimen with an oblique cordiform contour. F<eroes, 135 m.
k. A specimen from behind showing an intercostal sculpture in the form of
a fine picking. West Iceland (0nundarfjardur), ca. 12 fm.
Parvicardium
Drawlllgs from
A.S. Jensen (1912):
The Danish Ingolf
Expedition.
Note the characteristic oblique
scales on anterior ribs (g).
K. 'V. OckelmalUl, pers. comm.
"Cardium"
920
921
"Cardium"
Cardium & Serripes
922
& Trachycardium
923
Trachycardium
Trachycardium & Acrosterigma
924
925
Vasticardium
Trachycardium & Clinocardium
926
& Vaslicardium
927
Acrosterigma
Fulvia & Microcardium
928
929
Fulvia & Laevicardium
Laevicardium
930
931
Trigoniocardia & Fragum
Lunulicardia
932
933
Parvicardium & Afrocardium
Papyridea
934
935
Phuket Marine Biological Center Special Publication vol. 30 (2004)
INDEX
Specimens identified by Spengler and Lamarck are marked as [Spengler 's 10] and [Lamarck's 10] respectively.
TYPE is short for any kind of type illustrated on the Plates. Bold script refers to photographs of type material.
A
abrolhense Vidal, 1999. TYPE. - 872
aculeatum A. Linnaeus, 1758. [Spengler's 10]. - 844
aculeatum cf . Linnaeus, 1758. - 844
aculeatum Linnaeus, 1758. - 844
aculeatum Linnaeus, 1758. [Lamarck 's 10]. - 844
aeuleatum Linnaeus, 1758. TYPE. - 843
aeuleatum var. B. Spengler, 1799. TYPE. - 844
aculeatum var. C. Spengler, 1799. TYPE. - 844
adamsii Adams & Reeve, 1850. TYPE. - 902
aeolicum Born, 1778. [Lamarck 's 10]. - 899
aeolicum Born, 1778. TYPE. - 899
aequale Deshayes, 1854. TYPE. - 913
aequiliratum Poutiers, 1981. TYPE. - 90 I
albohamatum Hylleberg & Vidal, 2000. TYPE. - 881
andreae Dujardin, 1837. - 920
angulatum Lamarck, 1819. - 859
angulatllm Lamarck, 1819. [Lamarck's 10]. - 859
angulatum Lamarck, 1819. TYPE. - 859
aperta Bruguiere, 1789. TYPE. - 889
arcticum Sowerby, 1834. TYPE. - 852
arcuatum Montagu, 1803 TYPE. - 921
arenicolum Reeve, 1845. TYPE. - 872, 924
asiaticum Bruguiere, 1789. - 882
assimile Reeve, 1844. TYPE. - 859, 925
assimilis Reeve, 1844. - 859
auri Fischer-Piette, 1977. TYPE. - 915
auricula ForsskaJ. TYPE. - 932
australe Sowerby, 1834. - 880
australis Sowerby, 1834. - 929
aviculare Lamarck, 1805. - 921
B
ballieni Vidal, 1994. TYPE. - 887
beauforti Prashad, 1932. TYPE. - 873
belticum Reeve, 1845 . TYPE. - 851
biradiatum Bruguiere, 1789. ? TYPE. - 879
boholensis Vidal, 1994. TYPE. - 887
boreale Reeve, 1845. TYPE. - 922
brasilianum Lamarck, 1819. TYPE. - 929
buelowi Rolle, 1896. - 852
bul/alllm Lamarck, 1819. - 917
burdigalinum Lamarck, J 8 19. TYPE. - 920
burnupi Sowerby, 1897. TYPE. - 883
C
californiense Deshayes, 1839. - 926
californiense Deshayes, 1841. - 852
caparti Nickles, 1955. - 887
caparti Nickles, 1955. TYPE. - 887
cardissa Linnaeus, 1758. (Lamarck's 10]. - 913
cardissa Linnaeus, 1758. [Spengler's 10]. - 914
cardissa Linnaeus, 1758. - TYPE. 913
cardissa Linnaeus, 1858. - 921
cardissa var. B. Spengler 1799. - TYPE. 914
cardissa var. C. Spengler 1799. - TYPE. 914
eardissa var. D. Spengler 1799. - TYPE. 914
cardissa var. E. Spengler 1799. - TYPE. 914
cardissa var. F. Spengler 1799. - TYPE. 914
cardissa var. H. [Spengler's 10]. - 914
carditaeforme Reeve, 1845. TYPE. - 917,933
centumliratum Melvill & Standen, 1906. - TYPE. 900
ciliare Linnaeus, 1758. - 844
ciliare Linnaeus, 1758. (Lamarck's 10]. - 844
cilialum Fabricius, 1780. - 852, 853 , 890
compunctum Kira, 1954. TYPE. - 859
consors Sowerby, 1833. - 857,922, 923
consors Sowerby, 1833. TYPE. - 922
coronatum Spengler 1799. - 883
coronatum Spengler, 1799. TYPE. - 884
costatum Linnaeus, 1758. (Spengler's 10]. - 922
costatum Linnaeus, 1758. TYPE. - 843
couvrili Fischer-Piette, 1977. TYPE. - 874
crassum Gmelin, 1791. - 895, 896, 930
crenelloides Melvill . 1909. TYPE. - 917
crenulatum Lamarck, 1819. TYPE. 850
cygnorum Deshayes, 1855. - 874, 892
cygnorum Deshayes, 1855. TYPE. - 927
D
dampierense Wilson & Stevenson 1977. TYPE. - 874
delicatulum Smith, 1915. TYPE. - 854
deshayesii Payraudeau, 1826. TYPE. - 845
dianthinum Melvill & Standen, 1899. TYPE. - 875
diluvianum Lamarck, 1819. TYPE. - 920
discus Vidal, 1999. TYPE. - 875
dOllaceum Spengler 1799. TYPE. - 918
donaceum Spengler, 1799 - 896
dOllaceum var. B Spengler 1799. TYPE. 918
dulce Deshayes, 1863. TYPE. - 888
dupuchense Reeve, 1845. TYPE. - 925
E
ebaranum Yokoyama, 1927. - 917
echinatum Linnaeus, 1758. [Lamarck's 10]. - 846
echinatum Linnaeus, 1758. [Spengler's 10]. - 845
echinatum Linnaeus, 1758. TYPE.- 848
echinatum var. [al Lamarck. TYPE. - 845
echinatum var. [hI Lamarck. TYPE. - 845
edule Linnaeus, 1758. - 849
edule Linnaeus, 1758. (Lamarck's 10]. - 851
edule Linnaeus, 1758. [Spengler's 10]. - 850
936
Tropical Marine Mollusc Programme (TMMP)
edule Linnaeus, 1758. TYPE. - 850
edule var. Lamarck. TYPE. - 851
edule var. Linnaeus. - 850
eichwaldii Reeve, 1845. TYPE. - 851
elatum Sowerby, 1833. TYPE. - 892
elenensis Sowerby, 1840. TYPE. - 892
elongatum Bruguiere, 1789. - 860
elongatum Bruguiere, 1789. TYPE. - 861
elongatum cipangense Vidal, 1999. TYPE. - 873
elongatum coralense Vidal, 1993. TYPE. - 873
elongatum elongatum Bruguiere, 1789. - 860
elongatum ellode Vidal, 1993. TYPE. - 860
enode Sowerby, 1840. TYPE. - 860
erinaceum Lamarck. - 848
erinaceum Lamarck. TYPE. - 848
euglyptum Sowerby, 1914. TYPE. - 917
exasperatum Sowerby, 1838. TYPE. - 90 I
exiguum Gmelin, 1791. - 915
exochum Melvill, 1906. TYPE. - 857
F
fabricii Deshayes, 1854. TYPE. - 854
Jasciatum Montagu, 1803. - 919
festivum Deshayes, 1855. TYPE. - 888
fidele Vidal, 1992. TYPE. - 861
fimbriatum Lamarck, 1819. TYPE. - 884
flavum Born 1780. - 898
flavum Born 1780. TYPE. - 897
flavum Born, 1780. [Lamarck's 10]. - 863
flavum dupuchense Reeve, 1845 . - 925
flavum Linnaeus, 1758. [Spengler's 10]. - 862
flavllm Linnaeus, 1758. TYPE. - 861
flavum subrugosum Sowerby, 1838. TYPE. - 861
fornicatum Sowerby, 1840. - 902
fornicatum Sowerby, 1840. TYPE. - 902
foveolatum Sowerby, 1840. TYPE. - 874
Joveolatum Sowerby, 1841. - 927
fragiformis Vidal, 1994. TYPE. - 889
fragile ForskaJ in Niebuhr, 1775. TYPE. - 888
fragile Reeve, 1845. TYPE. - 888
fragilis congoensis Cosel, 1995. TYPE. - 887, 888
fragum Linnaeus 1758. - 906
fragum Linnaeus, 1758. [Spengler's 10]. - 905
fragum Linnaeus, 1758. TYPE. - 905
fraseri Garrard, 1963. TYPE. - 90 I
fucatllm Spengler, 1799. TYPE. - 875
G
gaillardi Fischer-Piette, 1977. TYPE. - 892
glaucum Bruguiere, 1789. - 851
glaucum mareotica Pallary, 1912. TYPE. - 851 .
graniferum Broderip & Sowerby, 1829. - 857
gratiosum Deshayes, 1855. TYPE. - 862
groenlandicllm Bruguiere, 1789. - 922
groenlandicum Bruguiere, 1789. [Spengler's IDJ. - 853
groenlandicum Bruguiere, 1789. TYPE. - 853
guanacastensis Hertlein & Strong, 1947. - 904
guichardi Bernardi, 1857. - 908
guichardi Bernardi, 1857. TYPE. - 908
H
hemicardium Linnaeus, 1758. [Spengler's IOJ. - 909
hemicardillm Linnaeus, 1758. TYPE. - 909
hians Spengler 1799. TYPE. - 889
hibernicum Sowerby, 1815. - 921
hiulcum Reeve, 1845. TYPE. - 934
hobbsae Vidal, 1999. TYPE. - 875
hlldsoniellse Deshayes, 1855. TYPE. - 890
hungerfordi Sowerby, 190 I. TYPE. - 890
hystrix Reeve, 1844. TYPE. - 902,903
hystrix Solander. - 903
imbricatum Born, 1780. [Spengler's 10]. - 906
imbricatu", Born, 1780. TYPE. - 905
imbricatum Sowerby 1840. TYPE. - 903
impolitum Sowerby, 1833. - 871,873,874
impoUtum Sowerby, 1833. TYPE. - 874
incarnatum Reeve, 1844. TYPE. - 885
indicllm Lamarck, 1819. TYPE. - 843
indioceanum Vidal, 1993. TYPE. - 864
intercoslatum Dujardin. - 921
interrogatorium Fischer-Piette, 1977. TYPE. - 892
islandicum Bruguiere, 1789. [Spengler's 10]. - 853
isocardia Linnaeus, 1758. [Lamarck's IOJ. - 854
isocardia Linnaeus, 1758. [Spengler's ID]. - 854
isthmicum IsseI, 1869. TYPE. - 852
J
junoniae Lamarck. [Lamarck's 10]. - 913
K
kalamantanum Adams & Reeve, 1850. TYPE. - 899
kerslakae Healey & LampreU, 1992. TYPE. - 864
L
lacullosum Reeve, 1845. TYPE. - 864, 876, 926
laevigatum Linnaeus, 1758. [Lamarck's 10]. - 893
laevigatum Linnaeus, 1758. [Spengler's 10]. - 893
laevigatum Linnaeus, 1758. TYPE. - 890
laevigatum Pennant, 1777 - 896
laevigatum var. Spengler, 1799. TYPE. - 893
lamarckii Reeve, 1845. [Lamarck's ID]. - 851
lamarckii Reeve, 1845. TYPE. - 851
lamellatum Spengler, 1799. TYPE. - 882
laperousii Deshayes, 1841. - 853
latum Born, 1778. [Spengler's ID]. - 902
latum Born, 1778. TYPE. - 917
lalum Bruguiere, 1789. [Lamarck's 10] . - 901
leucostomum Born, 1780. - 869,870
leucostomum Born, 1780. [Spengler's ID]. - 855
leucostomum Born, 1780. TYPE. - 855
leucostomum var. B Spengler, 1799. - 866
Phuket Marin e Biological Center Special Publication vol. 30 (2004)
levisulcatum Smith, 1903 . TYPE. - 916
lima Reeve, 1845 . TYPE. - 923
lima Spengler in Schroter, 1786. TYPE. - 882
Iineonotata Vidal, 1994. TYPE. - 890
lobulatum Deshayes, 1854. TYPE. - 892
wmboke Vidal, 2003 . TYPE. - 862
luteomarginatum Voskuil & Onverw., 1991. - 868
luteomarginatum insulare Vidal, 1997. TYPE. - 868
luteomarginatum marerubrumVosk. & 0., 1991. - 869
M
maculatum Sowerby, 1840. TYPE. - 876
maculosum howense Vidal, 1999. TYPE. - 877
maculosum Sowerby, 1834. - 876
maculosum Wood, 1815. - 872, 924
maculosum Wood, 1815. TYPE. - 876
magnijicum Deshayes. - 904
magnum Born, 1780 - 856
magnum Born, 1780. TYPE. - 856
magnum var. b. Spengler, 1799. TYPE. - 859
manthelaniense Dollfus & Dautz., 1888. TYPE. - 920
mareoticum Pallary, 1926. TYPE. - 851
marerubrum Voskuil & Onverwagt, 1991 . TYPE.- 869
marielae Wilson & Stevenson, 1977. TYPE. - 877
marmoreum Lamarck. - 869
marmoreum var. [2] Lamarck, 1819. TYPE. - 870
mauritianum Deshayes, 1855. TYPE. - 877
medium Linnaeus, 1758. [Lamarck 's IO]. - 905
medium Linnaeus, 1758. [Spengler's 10]. - 905
mendanense Sowerby, 1896. TYPE. - 867
mindanense Reeve, 1844. TYPE. - 857,858,923
minimum Philippi, 1836 - 890, 915
mirabile Deshayes, 1854. TYPE. - 885
multispinosum Sowerby, 1838. TYPE. - 886
mundum Reeve, 1845. TYPE. - 906,931
muricalum Linnaeus, 1758. [Lamarck 's 10]. - 858
muricatum Linnaeus, 1758. TYPE. - 858
muricatum var. [2]. Lamarck. - 858
muticum Reeve, 1844. TYPE. - 928
N
natalense Krauss, 1848. - 916
nebulosum Reeve, 1845 TYPE. - 877,927
nemo Fischer-Piette. 1977. TYPE. - 930
nivale Reeve, 1845. TYPE. - 909, 931
nodulosum Lamarck. - 847
nodulosum Spengler 1799. TYPE. - 906
norvegicum Spengler, 1799. [Lamarck's 10]. - 896
norvegicum Spengler, 1799. TYPE. - 895
norvegicum var. Spengler, 1799. TYPE. - 895
notabile Sowerby, 1915. TYPE. - 854
nuttallii Conrad, 1837. - 852
o
obliquum Lamarck, 1805. TYPE.- 920
obliquum Spengler, 1799. TYPE. - 856
937
oblongum Bruguiere, 1789. - 897,898
oblongum Gmelin, 1791. [Lamarck's IO]. - 896
obovale Broderip & Sowerby, 1833. TYPE. - 856, 931
obsoletum Spengler, 1799. TYPE. - 898
okinawense Kira, 1959. - 867
olivifer Iredale, 1936. TYPE. - 864
omane/lse Melvill & Standen, 1906. TYPE. - 915
orbita Broderip & Sowerby, 1833. TYPE. - 866
ovale Sowerby, 1840. - 919
oviputamen Reeve, 1844. - 894
oviputamen Reeve, 1844. TYPE. - 894
ovuloides Reeve, 1845. TYPE. - 856, 931
oxygonum Sowerby, - 866
oxygonum Sowerby, 1834. TYPE. - 878
P
pallidum Reeve, 1845. TYPE. - 928
panamense Sowerby, 1833. - 857
panamense Sowerby, 1833. TYPE. - 924
papillosum Poli, 179 I. - 933
papillosum Poli, 1791 . - 915
papuanum Vidal, 1996. TYPE. - 869
papyraceum Bruguiere 1789. - 890,893
papyraceum Bruguiere 1789. [Spengler's IO]. - 889
papyraceum Bruguiere 1789. TYPE. - 889
pectinatum Lamarck, 1819. TYPE. - 850
pectinatum Linnaeus, 1758. [Spengler's 10]. - 899
pectinatum Linnaeus, 1758. TYPE. - 899
pectiniforme Born, 1780. - 863
pectiniforme Born, 1780. TYPE. - 863
pellnanti Reeve. TYPE. - 930
peregrinum Jousseawne, 1888. TYPE. - 864
perornatum Iredale, 1929. TYPE. - 903
philippinense Hedley, 1899 (Deshayes MS). - 866
pianicostatll11l Sowerby, 1833. TYPE. - 904
polii Payraudeau, 1826. TYPE. - 933
politum Spengler, 1799. TYPE. - 896
porulosum Solander in Br., 1766. [Lamarck 's 10]. - 920
pristipleura Dall, 190 I. - 876
pristis Valenciennes in B.St Vincent, 1827. TYPE. - 895
probatu11l Iredale, 1927. TYPE. - 900
procerum Sowerby in Brod. & S., 1833 . TYPE. - 924
productllm Deshayes, 1854. TYPE. - 915
pro/undu11l Vidal, 1999. TYPE. - 878
pseud%ssile Reeve, 1844. TYPE. - 926
pseudo lima Lamarck, 1819. - 902
pulchellum Reeve, 1844. TYPE. - 928
pulchricosUltum Iredale, 1929. TYPE. - 885
pulchrum Reeve, 1845. TYPE. - 929
Q
quadrarium Reeve, 1845. TYPE. - 933
R
radialum Reeve, 1845. TYPE. - 929
radula Deshayes, 1832. - 921
938
rastrum Reeve, 1845. TYPE. - 923
replicatum Spengler 1799. TYPE. - 914
retusumLinnaeus,1758.-910
retusum Linnaeus, 1758. [Spengler's ID]. - 911
retusum Linnaeus, 1758. TYPE. 911, 932
retusum var. Spengler 1799. [Spengler's ID]. 911
rhomboides Lamarck, 1819. TYPE. - 921
richardi (Audouin, 1826). TYPE. - 918
ringens Bruguiere, 1789. [Spengler's ID]. - 843
robustum Lightfoot in Solander, 1786 - 856
rosemariensis Wilson & Stevenson 1977. TYPE. - 878
roseolum Melvill, 1909. TYPE. - 907
roseum Lamarck, 1819. TYPE. - 915
. rubescells Smith, 1911. TYPE. - 917
rubicundum Reeve, 1844. - 858, 923
rubicundum Reeve, 1844. TYPE. - 865,926
rubrohamatum Voskuil & OnvelWagt, 1988. - 881
rudelltis Fischer-Piette, 1977. TYPE. - 884
rugosum Lamarck, 1819. [Lamarck's ID] . - 863
rugosum var. I and 2. [Lamarck's ID). - 870
rusticum Linnaeus, 1758. - 849
rusticum Linnaeus, 1758. [Lamarck's IDJ. - 847, 849
rusticum Linnaeus, 1758. [Spengler's ID]. - 849
rusticum Linnaeus, 1758. TYPE. - 848
S
scalata Vidal, 1999. TYPE. - 890
scruposum Deshayes, 1855. TYPE. - 907
selelle Vidal, 1999. TYPE. - 877
semisulcata Gray, 1825. TYPE. - 916
senticosum Sowerby, 1833. - 923
serratum Linnaeus, 1758- 896
serra/urn Linnaeus, 1758. - 893
serratum Linnaeus, 1758. [Lamarck's ID]. - 894
serra tum Linnaeus, 1758. [Spengler's ID]. - 879
serratum Linnaeus, 1758. TYPE. - 894
setosum Redfield, 1848. - 902
seurati Vidal, 1999. TYPE. - 880
sewelli Prashad, 1932. TYPE. - 865
shepstonense Tomlin, 1931. TYPE. - 918
simplex Spengler, 1799. - 880
simplex Spengler, 1799. TYPE. - 879
sinense Sowerby, 1838. - 886
skeeti Hedley, 1906. TYPE. - 918
soleniforme Bruguiere, 1789. - 917
sorenselli Powell. TYPE. - 865
soyeri Fischer-Piette, 1977. TYPE. - 880
speciosum Adams & Reeve, 1848. TYPE. - 904
spinosus Deshayes. - 921
squamosum Gmelin 1791. - 922
stellatum Reeve, 1845. TYPE. - 916
Tropical Marine Mollusc Programme (TMMP)
striatlllum thetidis Hedley, 1902. TYPE. - 900
striatum Spengler, 1799. TYPE. - 880
sllbangulatum Scacchi, 1833 . TYPE. - 916
sllbassimile Vidal, 2003. TYPE. - 868
subelongatum Sowerby, 1840. TYPE. - 867
subretusum Sowerby, 1834. TYPE. - 912
suediense Reeve, 1845. TYPE. - 915
sueziensis Issei, 1869. - 852
sulcatinllm Lamarck, 1819. TYPE. - 921
sulcalum Lamarck, 1819. [Lamarck's ID]. - 898
sulcatum Lamarck, 1819. TYPE. - 898
suluanum Vidal , 1999. TYPE. - 880
symbolica Iredale, 1929. TYPE. - 903
T
tegulatum Dautzenberg, 1900. TYPE. - 907
tenuicosta Lamarck, 1819. - 929
tenuicostatum Lamarck, 1819. [Lamarck's ID) - 891
telluicostatum Lamarck, 1819. TYPE. - 891
tenuicostatum var. Lamarck. [Lamarck's ID]. - 891
tenuilamellosum Poutiers, 1981. TYPE. - 901
tltomassini Vidal, 1998. TYPE. - 868
torresi Smith, 1885. TYPE. - 900
torresi. Smith, 1885. - 900
trallslatum Prashad, 1932. TYPE. - 870
transversale Deshayes, 1854. - 917
tuberculatum Linnaeus, 1758. - 847,849
tuberculatum Linn., 1758. [Lamarck's ID]. - 846, 847
tuberculatum Linnaeus, 1758. [Spengler's ID]. - 846
tuberclliatum Linnaeus, 1758. TYPE. - 846
tumidllln Deshayes, 1854. TYPE. - 871
tumoriferum Lamarck, 1819. - 910
U
unedo Linnaeus, 1758. - 905
unedo Linnaeus, 1758. [Spengler's ID]. - 907
unedo Linnaeus, 1758. TYPE. - 907
unicolor Sowerby, 1840. TYPE. - 881
ullimaculatum Broderip & Sowerby, 1833. TYPE. - 914
uniornatum Vidal, 1999. TYPE. - 881
V
variegatum Sowerby, 1840. TYPE. - 870
ventricosum Bruguiere , 1789. - 856
victor Angas, 1872. TYPE. - 904
vitellinum Reeve, 1844. TYPE. - 930
vlamingi Wilson & Stevenson, 1977. TYPE. - 871
vlllva Jousseaume, 1898. TYPE. - 852
W
whitleyi Iredale, 1929. TYPE. - 908
wilsoni Voskuil & OnvelWagt, 1991. TYPE . - 872
939
Phuket Marine Biological Center Special Publication vol, 30 (2 004)
RECTIFICATIONS
Corrections regarding Part 1:
p. 44 fig, No.9: correct
p.146 ex Plate 1: correct
correct
p.351 legend:
C cardissa
Tridacna giga
Vigmette
into C isocardia
into Tridacna gigas
into Vignette
June 2004
Printed by Funny Publishing
549/1 Soi Senanikom, Pholyothin Rd.
Chatuchak, Bangkok 10900 Thailand
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Vasticardium e/ongatum (Bruguiere, 1789)
The British malaeologist Hugh Cuming (1791-1865) collected and sold shells, many of which
ended up in cabinets, for example of the Danish King Christian 8 (C VIII written inside the
shells). The specimens are now deposited in Copenhagen at the ZMUC. The shells are from the
Phillipines. Reeve (1844 fig. 46) illustrated the species [Sec Part I p. 132J. I made the pictures
on a flatbed scanner (Agfa StudioScan IIsi) and converted the colours to gray scale.