Ernstbrunn Limestone and Klentnice beds
(KimmeridgianBerriasian; Waschberg-dánice Unit;
NE Austria and SE Czech Republic):
state of the art and bibliography
SIMON SCHNEIDER, MATHIAS HARZHAUSER, ANDREAS KROH, ALEXANDER LUKENEDER
& MARTIN ZUSCHIN
This paper summarises the knowledge of the Ernstbrunn Limestone and Klentnice beds and provides a comprehensive
scientific bibliography on these strata. At outcrop both lithostratigraphic units occur as so-called “tectonic klippen” inserted in the autochthonous sedimentary succession of the Waschberg-Ždánice Unit. The latter is a distal, transitional
Alpine-Carpathian tectonic nappe that extends between the Danube and Thaya rivers in Lower Austria and southern
Moravia. Both strata have also been identified from several drillings and belong to the autochthonous Mesozoic succession deposited on the southern slope of the Bohemian Massif. Ammonite biostratigraphy and micropalaeontology reveal
a Kimmeridgian to early Late Tithonian age for the Klentnice beds and a Middle Tithonian to Berriasian (?Hauterivian)
age for the Ernstbrunn Limestone. The Ernstbrunn-Pavlov Carbonate Platform gradually developed from the Klentnice
beds and persisted during the Jurassic–Cretaceous transition. The rock record provides evidence for lagoonal and patch
reef facies and fringing ooid-oncoid bars, all attributed to the Ernstbrunn Limestone. The gradual transition to more distal, siliciclastically-influenced settings is formed by the upper portion of the Klentnice beds that developed as lateral
equivalents of the carbonates. In places, both strata are highly fossiliferous. The lagoonal limestones preserve a
megadiverse, mollusc-dominated assemblage of more than 500 species of invertebrates and calcareous algae. Most
abundant taxa include Heterodiceras and Epidiceras bivalves (basal rudists), nerineid gastropods, decapods,
ammonites, corals, and dasycladaceans. As a result of diagenetic aragonite loss, the fauna of the Klentnice beds appears
impoverished and is dominated by echinoderms, calcareous sponges, and brachiopods. A remarkably large portion of
the complex depositional and natural environment of the Ernstbrunn Limestone and Klentnice beds is preserved both at
outcrop and in subsurface and still awaits systematic scientific effort in various fields. • Key words: tectonics, stratigraphy, Tithonian–Berriasian, Upper Jurassic–Lower Cretaceous, northern Tethys margin, carbonate platform.
SCHNEIDER, S., HARZHAUSER, M., KROH, A., LUKENEDER, A. & ZUSCHIN, M. 2013. Ernstbrunn Limestone and
Klentnice beds (Kimmeridgian–Berriasian; Waschberg-Ždánice Unit; NE Austria and SE Czech Republic): state of the
art and bibliography. Bulletin of Geosciences 88(1), 105–130 (9 figures, 1 table). Czech Geological Survey, Prague.
ISSN 1214-1119. Manuscript received April 17, 2012; accepted in revised form July 12, 2012; published online November 23, 2012; issued December 6, 2012.
Simon Schneider (corresponding author), Mathias Harzhauser, Andreas Kroh & Alexander Lukeneder, Natural History
Museum Vienna, Department of Geology & Palaeontology, Burgring 7, 1010 Vienna, Austria; simon.schneider@nhm-wien.ac.at • Martin Zuschin, University of Vienna, Department of Palaeontology, Althanstrasse 14,
1090 Vienna, Austria
This article intends to summarise the current state of knowledge on the facies, biota, and age of the Ernstbrunn Limestone and Klentnice beds both at outcrop and from drill
cores, and to provide a bibliography for these strata, similar
to the compilation on the coeval Štramberk Limestone (Silesian Unit, NE Czech Republic) by Vašíček & Skupien
(2004, 2005). All Upper Jurassic klippen of the
Waschberg-Ždánice Unit, stretching from Niederfellabrunn to Kleinschweinbarth in Lower Austria, and further
DOI 10.3140/bull.geosci.1360
on to the Pavlovské vrchy (Pavlov Hills) of the Mikulov
area in Czech Republic are considered, and data on their
tectonic formation are compiled. The summary is split into
four parts, covering (1) the geological overview and tectonic evolution, (2) the stratigraphy of the klippen, (3) the sedimentology and microfacies of the Ernstbrunn Limestone
and Klentnice beds, and (4) the fossil flora and fauna that
have been preserved in these rocks. Furthermore, we outline the geological and palaeontological significance of the
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Bulletin of Geosciences Vol. 88, 1, 2013
A
B1
B2
Figure 1. Epidiceratidae: Character fossils of the Ernstbrunn Limestone. • A – internal mould of Epidiceras sp.; Dörfles; NHMW 2012/0108/0002.
• B – small cluster of five specimens of Heterodiceras lucii (Defrance, 1819) from a karst fissure at Dörfles V quarry; NHMW 2012/0111/0001.
• B1 – small double-valved specimen (upper left), adult left valve (upper right), and two fragmentary specimens (left- and right-central) growing on a single right valve (below). • B2 – same cluster as in B1; side view. Scale bar = 10 mm.
Ernstbrunn Limestone and Klentnice beds, and highlight
potential future directions of research.
Most people familiar with the Upper Jurassic
Ernstbrunn Limestone likely will think of epidiceratid bivalves when this unit is mentioned – and with a bivalve it
all began. When Karl Haidinger (1782) reported on an
epidiceratid bivalve unearthed from the carbonates of the
Semmelberg near Ernstbrunn, he considered this find a rarity. In fact, he had discovered the character fossil of the
stratum, i.e. a large internal mould of a basal rudist
(Fig. 1A). However, it took almost 50 more years, until
geological investigations on the Ernstbrunn Limestone
started. It was Ami Boué, who “painted” a first picture of
these carbonates in his “Geognostisches Gemälde von
Deutschland” (Boué 1829) and established the term
“Calcaire d’Ernstbrunn” (= Ernstbrunn Limestone) in his
follow-up article on the Mesozoic stratigraphy of the “Austrian Alps” (Boué 1830). Since then, many famous Austrian and Czech geologists have contributed to the discussion about the nature and origin of the enigmatic Upper
Jurassic hill chain that occurs to the north of Vienna, between the Danube and Thaya rivers (see bibliography below). Finally, these rocks were recognised as unrooted tectonic klippen that have been sheared off the Mesozoic
basement below the Vienna Basin, and dragged up in the
course of the thrusting of the Waschberg-Ždánice Nappe
onto the Alpine-Carpathian Foredeep (= Molasse Unit;
Figs 2, 3) (Poul et al. 2011, Wessely 2006).
Besides the vivid discussion on structural geology, first
essays on the lithology and fauna of several outcrops of
Ernstbrunn Limestone were compiled by Ferstl (1845),
106
Prinzinger (1851), Hingenau (1852), Foetterle (1853), and
Hobza (1876). In the late 19th century, Abel (1899a, b) provided a comprehensive description of the sediments and
listed the fossil fauna of the Ernstbrunn Limestone. Scattered over the 20th century, several classical, macroscopic
facies descriptions of the stratum were published (e.g.,
Jüttner 1922, 1942; Schön 1927; Tollmann 1963), until the
microfacies of the Ernstbrunn Limestone was studied in
detail in Moravia (Eliáš 1992; Eliáš & Eliášová 1984,
1986; Řehánek 1987a, b) and in Austria (Hofmann 1990a,
Moshammer & Schlagintweit 1999). The latter authors
provide modern lithologic characterisations of the Ernstbrunn Limestone at Dörfles and a few other small outcrops
(Fig. 4). Information on the microfacies of several major
outcrops (e.g., Falkenstein, Staatz), however, is still wanting, and a comprehensive facies model for the Ernstbrunn-Pavlov Carbonate Platform has not been proposed.
After the first description by Haidinger (1782), the
abundant, large fossils preserved in the Ernstbrunn Limestone soon had attracted both collectors and scientists (e.g.,
Hoernes 1874; Makowsky 1874; Peters 1855, 1867; Suess
1858). In the 20th century, the fossil fauna of the Ernstbrunn
Limestone was treated in several unpublished Ph.D. theses
(Bachmayer 1940, Dürrmayer 1931, Matzka 1934, Moeller
1911). However, only a few peculiar species or genera were
formally described, and several moderately specious taxa
were monographed (see palaeontology section below). A
milestone for further investigation was set, when the
ammonite fauna and biostratigraphy of the Ernstbrunn
Limestone were revised (Zeiss & Bachmayer 1989, Zeiss
2001). Recently, a re-evaluation of the extraordinarily rich
Simon Schneider et al. Ernstbrunn Limestone and Klentnice beds state of the art and bibliography
Figure 2. Main tectonic units of the study region, including the Waschberg-Ždánice-Unit (WZU). Dashed line (A–B) indicates position of cross-section
in Fig. 3.
A
B
Figure 3. Schematic cross-section of the Molasse, Waschberg-Ždánice, and Flysch units and adjacent Vienna Basin. Modified from Zimmer & Wessely
(1996).
decapod fauna of the Ernstbrunn Limestone has been
started, which already produced a wealth of results
(Feldmann & Schweitzer 2009; Robins et al. 2012, 2013;
Schweitzer & Feldmann 2008a, b, 2009a–d, 2010a–d).
Major portions of the fauna, however, including several of
the most specious groups (e.g., Scleractinia, Bivalvia,
Gastropoda, Brachiopoda, Echinoidea), have never been
studied in detail. As a result, knowledge on the palaeobiodiversity and palaeoecology of the Ernstbrunn-Pavlov
Carbonate Platform remains piecemeal, despite of more
than 200 years of research.
Any comprehensive essay on the Ernstbrunn Limestone will necessarily also expand on the closely associated
Klentnice beds. Generally, the Klentnice beds comprise a
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Bulletin of Geosciences Vol. 88, 1, 2013
Table 1. Locality names and GPS coordinates of current and former outcrops of Ernstbrunn Limestone (E) and Klentnice beds (K), inferred from
labels of fossils in the collections of NHMW. Numbers in the first column
correspond to Fig. 4. Localities without number and coordinates could not
be matched with a particular outcrop.
NR Stratum Locality
Coordinates
1E
Dörfles I
48°32´43.81˝ N 16°21´00.52˝ E
2E
Dörfles II
48°32´45.94˝ N 16°20´58.67˝ E
3E
Dörfles III
48°32´53.77˝ N 16°20´50.86˝ E
4E
Dörfles IV
48°32´57.21˝ N 16°20´53.47˝ E
5E
Dörfles V
48°33´03.53˝ N 16°20´47.59˝ E
6E
Ernstbrunn Kalkwerk II,
Ernstbrunn II
48°32´52.34˝ N 16°21´21.08˝ E
7 E, K
Ernstbrunn (Semmelberg)
48°31´58.36˝ N 16°20´54.76˝ E
–E
Kottingneusiedl
–
–
–E
Grafensulz
–
–
8E
Michelstetten (Galgenberg)
48°35´44.05˝ N 16°26´38.05˝ E
9E
Falkenstein (Steinbruch)
48°43´38.86˝ N 16°35´01.29˝ E
–E
Falkenstein (not specified)
–
10 E
Klafterbrunn I
48°33´36.43˝ N 16°21´07.68˝ E
11 E
Klafterbrunn II
48°33´33.74˝ N 16°21´09.40˝ E
12 E
Klafterbrunn III
48°33´29.36˝ N 16°21´13.42˝ E
13 E
Klafterbrunn IV
48°33´25.36˝ N 16°21´14.61˝ E
14 E, K
Klement I
48°34´06.31˝ N 16°21´02.43˝ E
Mikulov (Marienmühle)
48°48´39.34˝ N 16°39´31.21˝ E
15 E
–E
16 E, K
–
Asparn an der Zaya
–
Leiser Berg bei Niederleis
48°33´33.25˝ N 16°22´25.77˝ E
–
17 E
Staatz
48°40´34.71˝ N 16°29´17.14˝ E
18 K–E
Südmährerkreuz
48°45´43.75˝ N 16°36´48.35˝ E
19 K–E
Steinbruch bei Schletz
48°34´49.78˝ N 16°27´10.81˝ E
20 K
Niederfellabrunn
48°28´04.64˝ N 16°18´14.19˝ E
21 K
Buschberg NE Klement
48°34´37.42˝ N 16°23´44.84˝ E
22 K
Stützenhofen
48°44´30.60˝ N 16°36´30.90˝ E
23 K
Niederfellabrunn (Hundsberg) 48°28´37.40˝ N 16°18´28.70˝ E
24 K
Mikulov (Gemeindeberg)
48°48´36.73˝ N 16°38´15.59˝ E
25 K
Mikulov (Heiliger Berg)
48°48´27.11˝ N 16°38´55.39˝ E
26 K
Děvín (Maydenberg)
48°52´09.94˝ N 16°38´07.94˝ E
–E
Perná N Nikolsburg
[Makowsky 1874]
–
–
27 K
Niederfellabrunn
48°27´45.11˝ N 16°18´12.13˝ E
28 E
Mikulov (Quarry Holy Mt.)
48°48´31.06˝ N 16°39´16.44˝ E
29 E, K
Mikulov (Turold Quarry)
48°49´00.51˝ N 16°38´23.91˝ E
30 K
Klentnice (Table Mountain)
48°50´56.38˝ N 16°38´11.98˝ E
mixture of fossiliferous siliciclastics and carbonates (see
sedimentology section below) and grade vertically and
laterally into the pure carbonates of the Ernstbrunn Limestone (Glaessner 1931, Jüttner 1922; cf. Poul et al.
2011). The term “Klentnitzer Mergel” (= Klentnitz Marls)
was introduced by Abel (1899a), who was among the first
108
scientists to provide a detailed description of this stratum,
including a list and several drawings of the fauna (Abel
1897). The name refers to the informal type locality,
Klentnice (Klentnitz), located to the north of Mikulov
(Czech Republic).
Since the 1950s, the sediments of the Molasse and
Waschberg units in Austria and Czech Republic have been
penetrated by numerous wells reaching down to the crystalline basement of the Bohemian Massif. In several drill
cores, the autochthonous Mesozoic succession on the crystalline slope contains both Ernstbrunn Limestone and
Klentnice beds, which finally proved the tectonic scenario
outlined above (e.g., Adámek 1979, 1986; Eliáš & Wessely
1990; Malzer et al. 1993; Poul et al. 2011; Wessely 2006).
Recently, new data retrieved from subsurface rocks and
surface geology have been integrated, producing a more
complete view of the klippen strata (Adámek 2005, Poul et
al. 2011, Wessely 2006). Since the 1970s, an additional set
of lithostratigraphic units and terms has been developed for
the autochthonous Mesozoic strata in subsurface (see Eliáš
& Wessely 1990 for an overview).
Abbreviations used in the text. – GBA = Geologische Bundesanstalt, Vienna; NHMW = Natural History Museum
Vienna; WZU = Waschberg-Ždánice Unit.
State of the art
Geographic overview, geology and tectonics
At surface, the Ernstbrunn Limestone and Klentnice beds
occur within a SW–NE trending hill chain that starts at the
Waschberg north of Stockerau (Lower Austria) and ends at
the Děvín (Maydenberg) west of Pavlov (Pollau) in southern Moravia (Czech Republic), thus extending between the
Danube and Thaya rivers over approximately 60 km
(Fig. 2), and probably even a few kilometres further to the
north (Pícha & Hanzlíková 1965). Together with the elongated strip of surrounding sediments these hills form a distinct tectonic unit, named Waschberg Zone according to
the concepts developed by Tercier (1936) and Grill (1953)
[note that at that time the strict restriction of the term
“Zone” to the biostratigraphic nomenclature was not established]. Czech geologists have designed an alternative
term, i.e. Ždánice “Zone” (also Ždánice-Subsilesian
“Zone”), for the same unit (Roth 1962). Since there is no
official agreement on a common name, a combination of
both names is used herein. With regard to structural geology, the Waschberg-Ždánice Unit (WZU) represents the
most distal Alpine-Carpathian nappe in the boundary region between these two orogens. Moreover, the WZU is the
only structural element that forms a direct connection of
the Alps and Carpathians at surface (Prey 1960, 1965; Roth
Simon Schneider et al. Ernstbrunn Limestone and Klentnice beds state of the art and bibliography
1967; Tollmann 1971; Fig. 2). Towards the north, the
WZU is bordered by the Alpine-Carpathian Foredeep,
which has been partly overthrust by the WZU during the
Late Styrian tectonic phase at the Karpatian/Badenian
(= Early/Middle Miocene) boundary (Pokorný 1973, Tollmann 1966; Figs 2, 3). In its easternmost part, the Pouzdřany thrust sheet is inserted between the Alpine-Carpathian Foredeep and the WZU. In turn, the WZU is
overthrust by the Rhenodanubian Flysch, as can be seen
in the south-western part of the WZU. Because of its sandwiched tectonic position between the Molasse and Flysch
zones (Fig. 3), the WZU has been thought to represent an
eastern prolongation of the Helvetic Unit by several scholars (Tercier 1936, Tollmann 1963, Trauth 1948). In its
Figure 4. Detailed geologic map of the study region. Numbers of outcrops correspond to Table 1. Modified from Grill
(1961), Poul et al. (2011) and Schnabel (2002).
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Bulletin of Geosciences Vol. 88, 1, 2013
southern part, the Austrian WZU is tectonically partitioned
into two subunits, i.e. the outer Roselsdorf Unit and the inner Waschberg Unit sensu stricto. Further partitioning into
several blocks occurred along reactivated faults of Variscan strike (Poul et al. 2011, Wessely 2006).
The regular sedimentary succession of the WZU consists of Miocene strata. Several of the hills, which are
composed of Oligocene, Eocene, Palaeocene, Upper Cretaceous, and Upper Jurassic rocks (Fig. 4), rear up as more
or less steep blocks, and thus are termed “cliffs” in common speech. The Upper Jurassic strata form major parts of
the Leis Hills complex (Leiser Berge), including
Semmelberg, Dörfles (Fig. 5A, B), Klement, and
Buschberg localities, and the hills of Staatz (Wessely &
Hofmann 2007; Fig. 5D), Falkenstein (Fig. 5C),
Stützenhofen (Fig. 5F), and Kleinschweinbarth in Lower
Austria. In the Czech Republic, in and around Mikulov
(Nikolsburg) in southern Moravia, they comprise (Fig. 4)
the Pavlov Hills (Pavlovské vrchy, Pollauer Berge),
i.e. the Saint Hill (Svatý kopeček, Heiligenberg), Turold,
Table Mountain (Stolová hora, Tafelberg), and Děvín
(Maydenberg; Fig. 5E). Stratigraphically, both the Klentnice beds and the Ernstbrunn Limestone have not yet been
formalised and type localities and sections have not been
officially designated or described. Consequently, both
units are not termed formations herein. Moreover, we favour the term “Klentnice beds” versus “Klentnice Marls”,
since the stratum comprises a variety of different facies
(see sedimentology section below). Recently, Poul et al.
(2011) proposed a separate informal unit termed “nodular
limestones” for a transitional zone between the Klentnice
beds and Ernstbrunn Limestone in the Pavlov Hills. The
term “nodular limestone” is commonly attributed to condensed limestone facies deposited around the calcite compensation depth (Jenkyns 1974). This does not account
for the facies found in the Pavlov Hills. Moreover, any approach to separate different units along a gradual transition will remain arbitrary. Herein, we thus assign all
well-bedded portions of Upper Jurassic rocks in the WZU
to the Klentnice beds.
During the last 40 years, a series of lithostratigraphic
units and terms have been established in order to characterise the subsurface, autochthonous Mesozoic strata on
the southern slope of the Bohemian Massif (e.g., Eliáš
1974, Ladwein 1976), which have been summarized and
harmonized by Eliáš & Wessely (1990). According to the
resulting scheme, the Upper Jurassic portion of the succession starts with the Vranovice limestones and
dolomites, which have no equivalent at surface. These
carbonates are overlain by the Mikulov Marls and the
Kurdějov Arenite. The latter two units are considered
equivalents of the Klentnice beds. Due to the lack of information on the typical succession of the Klentnice
beds, however, it remains enigmatic whether these strata
110
are effectively facially equivalent. The Jurassic succession is topped by the Ernstbrunn Limestone, which is
capped by erosion, and overlain by limestones of Late
Cretaceous age (Adámek 1979, 1986; Eliáš & Wessely
1990; Pokorný 1958; Poul et al. 2011).
The initial, most controversial and still enduring discussion concerns the origin of the Jurassic rocks of the
WZU, and several contrasting theories were proposed on
that subject. The “island theory” – most popular during
the early years of research – identified the Jurassic
klippen as erosional relics of a formerly continuous cover
of Jurassic rocks on the crystalline basement, which acted
as islands in the Cenozoic Paratethys Sea (e.g., Stejskal
1932, Suess 1929, Uhlig 1904). In this context, the
Ernstbrunn Limestone has also been thought to represent
a prolongation of the Upper Jurassic strata of the
Ortenburg-Passau region (Lower Bavaria, SE Germany;
Suess 1885), which, however, are true erosional relics of
different facies and age resting directly on the crystalline
slope (Gröschke 1985). Other scholars regarded the
klippen as large olistolithes that were derived from a now
eroded Jurassic shelf in superior position (e.g., Stejskal
1928, 1930, 1931, 1935b; Uhlig 1903). When a first set of
boreholes penetrated several unrooted Carpathian klippen
in the Silesian flysch unit in the early 20th century, scientists began to discuss a similar tectonic model for the
Ernstbrunn Limestone (Friedl 1927; Jüttner 1930, 1931,
1932, 1933b, 1934; Petrascheck 1914, 1921; Schnabel
1929, 1933a, b). Starting from 1948, numerous wells also
penetrated the sedimentary successions of the AlpineCarpathian Foredeep and WZU both in Austria and Czech
Republic, mainly in order to explore the hydrocarbon resources of the region (Eliáš & Wessely 1990, Tollmann
1985). The new data confirmed the presence of Upper Jurassic strata identical to the Ernstbrunn Limestone and
Klentnice beds in the autochthonous Mesozoic cover of
the slope of the Bohemian Massif (Brix & Götzinger
1964; Brix et al. 1977; Eliáš 1977, 1981; Eliáš & Wessely
1990; Grill 1958; Kapounek et al. 1967; Kröll 1980;
Malzer et al. 1993; Wessely 1993, 1997, 2006; Zimmer &
Wessely 1996). It is therefore confirmed that the Upper
Jurassic blocks originate from below the WZU and thus
truly represent un-rooted tectonic klippen, but not
olistolithes (Kapounek et al. 1967, Poul et al. 2011,
Wessely 2006). Based on detailed geologic mapping,
borehole data, and seismic analysis that were in part already presented by Stráník et al. (1979), Poul et al. (2011)
recently developed a comprehensive model of the tectonic evolution of the Pavlov Hills expanding on details of
the partitioning of the Jurassic rocks due to thrusting and
strike-slip movements. The resulting scheme shows that
the Pavlov klippen extend much further in subsurface
than those in the southern part of the WZU, with only relatively small horst structures cropping out.
Simon Schneider et al. Ernstbrunn Limestone and Klentnice beds state of the art and bibliography
A
B
C
D
E
F
Figure 5. Major outcrops of Ernstbrunn Limestone (A–E) and Klentnice beds (F). • A – Ernstbrunn Kalkwerk II quarry. • B – Dörfles V quarry.
• C – Falkenstein quarry. • D – Staatz Klippe. • E – Děvín north of Mikulov. • F – roadcut at Stützenhofen.
Stratigraphy
Biostratigraphy of the Upper Jurassic klippen strata is still
not settled and alternative stratigraphic approaches, e.g.,
sequence stratigraphy, isotope geochemistry, or palaeomagnetics, have never been tested for the Jurassic rocks of
the WZU. The first author who clearly stated a Jurassic age
for the Klentnice beds was Suess (1852). A few years later,
Zittel (1868), based on the macrofauna of the Klentnice
beds, regarded the Jurassic klippen as older than Tithonian.
Typically, Jurassic biostratigraphy is based on ammonites.
They are, however, relatively rare in most outcrops of the
Ernstbrunn Limestone and Klentnice beds and therefore
ammonite biostratigraphy on the klippen strata started relatively late.
Most ammonites from the Klentnice beds have been
unearthed from wine cellars at Niederfellabrunn. These
relatively poorly preserved specimens and a few other
finds have been extensively discussed, and determinations of earlier publications were usually emended by
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Bulletin of Geosciences Vol. 88, 1, 2013
subsequent authors (Abel 1897, Arkell 1956, Bachmayer
1957, Krafft 1897, Kutek & Zeiss 1988, Spath 1933,
Vašíček 1971, Vetters 1905, Zeiss 1977). According to the
latest update by Kutek & Zeiss (1988) and Zeiss (2001), the
seven ammonite taxa known from the Klentnice beds
largely suggest an early Late Tithonian age (Micracanthoceras microcanthum Zone, Simplisphinctes Subzone; Ogg
et al. 2008).
Besides ammonites, different microfossils were locally
employed for biostratigraphy. From a single locality in the
Pavlov Hills a rich fauna of Foraminifera and Ostracoda
from the lower part of the Klentnice beds was studied
(Hanzlíková 1962, 1965; Pokorný 1973), which produced
conflicting results. While a latest Oxfordian to Late Kimmeridgian age was deduced from the Foraminifera
(Hanzlíková 1965), a Late Tithonian to Berriasian age was
inferred from the Ostracoda (Pokorný 1973). A re-evaluation, however, confirmed a Kimmeridgian age for both
groups (Gramann & Luppold 1991, Schudack & Schudack
1997).
Relatively sound biostratigraphic data are also available for the autochthonous Mesozoic in subsurface.
Ammonites from several wells in southern Moravia described by Vašíček (1971, 1980) indicate a ?Callovian to
Oxfordian age for the Vranovice limestones and dolomites,
while Brix et al. (1977) report an Oxfordian to latest
Kimmeridgian age for the same stratum in Austria. The
overlying Mikulov Marls, corresponding to the lower portion of the Klentnice beds have been dated as Kimmeridgian to early Middle Tithonian (Holzknecht & Hamršmíd
1988; Vašíček 1980) or Early Tithonian (Brix et al. 1977).
The Kurdějov Arenite, representing the upper portion of
the Klentnice beds at subsurface, is interpreted to comprise
upper Middle to lower Upper Tithonian deposits (cf. Brix
et al. 1977, Vašíček 1980). Papp & Turnovsky (1964)
studied the microfauna from drill cores from the Austrian
part of the WZU and concluded on a Late Jurassic
(?Tithonian) to Berriasian (?Hauterivian) age for the
equivalents of the Klentnice beds. Brix et al. (1977) regard
the microfauna derived from these cores as stratigraphically largely indistinct, but propose an exclusively
Late Jurassic age for the Klentnice beds.
In contrast to the specimens from the Klentnice beds,
the ammonites from the Ernstbrunn Limestone are usually
well preserved. Up to the 1950s, only few specimens were
available for study (Abel 1899a, b; Arkell 1956; Bachmayer 1957; Moeller 1911; Spath 1933). The most comprehensive assessment of the ammonites by Zeiss is mainly
based on the extensive collections of Friedrich Bachmayer
(Zeiss 1999, 2001; Zeiss & Bachmayer 1989). In conclusion, Zeiss (2001) proposes middle Middle to early Late
Tithonian age (Richterella richteri Zone to Micracanthoceras microcanthum Zone, Simplisphinctes Subzone) for
the Ernstbrunn Limestone. It should be mentioned that the
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majority of the ammonite specimens were collected at
Ernstbrunn-Dörfles Werk II quarry, while most of the other
fossils, usually representing benthic taxa were found at
other quarries. This suggests significant variation in facies
between the different klippen and outcrops – and probably
also differences in age.
An alternative biostratigraphic approach based on calpionellids is restricted to several exposures of Ernstbrunn
Limestone in the Mikulov region, and revealed a much
younger, Tithonian to Hauterivian age (Eliáš & Eliášova
1985, Houša & Řehánek 1987, Houša et al. 1963). Likewise, a study of microflora and microfauna from several
small outcrops of Ernstbrunn Limestone (Kleinschweinbarth, surroundings of Ernstbrunn, Galgenberg, surroundings of Falkenstein and Schletz) points to a Tithonian to
Early/Middle Berriasian age (Moshammer & Schlagintweit 1999).
In conclusion, the Klentnice beds are thought to comprise the Kimmeridgian to early Late Tithonian, while the
Ernstbrunn Limestone may have been deposited during the
Middle Tithonian to Berriasian (?Hauterivian). According
to this scenario, the Ernstbrunn-Pavlov Carbonate Platform
persisted during the Jurassic–Cretaceous transition and is
thus entirely coeval to the Štramberk Limestone of northern Moravia.
Sedimentology and facies
The sedimentology and microfacies of the Pavlov Hills
have been explored in detail (Eliáš 1962; Eliáš & Eliášová
1984, 1986; Fabian & Jüttner 1937; Jüttner 1933a; Matějka
1926, 1929; Matějka & Stráník 1961; Řehánek 1987a, b),
but published data from the Ernstbrunn Limestone and
Klentnice beds in Austria are sparse. Comprehensive general sedimentological characterisation of several outcrops in
Austria has been provided by Glaessner (1937). The texture and faulting of the Ernstbrunn Limestone are accurately described by Schön (1927) and Havíř & Stráník
(2003). At several outcrops, for example at Staatz or the
southern flank of the Děvín, large, severely brecciated rock
portions provide evidence of the tectonic movements outlined above (Petrascheck 1921, Zapletal 1930).
In the 1980s, modern microfacies analysis of the
Ernstbrunn Limestone of the Pavlov Hills started, and numerous carbonate thin sections were screened by Eliáš &
Eliášová (1984, 1986) and Řehánek (1987a, b). In Austria,
the microfacies of the five classical, now abandoned quarries at Dörfles was analysed in a diploma thesis by
Hofmann (1990a). He provided schematic sections and
identified different types of lagoonal microfacies, i.e.
wackestones, packstones, grainstones, algal bindstones,
and “diceratid” facies. However, only a brief overview of
his results is published in Hofmann (1992b, 1993a, 2001).
Simon Schneider et al. Ernstbrunn Limestone and Klentnice beds state of the art and bibliography
Several small outcrops of Ernstbrunn Limestone in Austria
have been more or less randomly sampled by Moshammer
& Schlagintweit (2003), who recorded different types of
inner and outer platform carbonates. The microfacies of
most of the important outcrops of Ernstbrunn Limestone
(e.g., Falkenstein, Michelstetten, Klement, Staatz, Stützenhofen), however, was never studied. Consequently, an integrative facies model for the Ernstbrunn-Pavlov Carbonate
Platform is still lacking, as is a detailed comparison and
correlation of autochthonous, subsurface rocks with those
sediments available at outcrop.
According to the publications listed above, the Klentnice beds consist of claystones and marls, which dominate
the lower portion of the stratum, while intercalations of
limestone (ooid-, intraclast-, and bioclast-wacke- to packstones; Fig. 6B, D, H) become gradually more abundant towards the top. These facies are interpreted as base-of-slope
to pelagic mud facies by Eliáš & Eliášová (1984). However, because the shallow-water carbonates of the Ernstbrunn Limestone gradually developed from the Klentnice
beds, the latter may rather represent shelf sediments deposited at moderate depth. This is underlined by the composition of the rich macrofauna (see below).
Similar to the Klentnice beds, the individual outcrops
of Ernstbrunn Limestone are composed of several different
types of facies (Fig. 6A, C, G). Fossiliferous or oncoidal
wackestones and packstones, algal bindstones, and
epidiceratid- and coral-framestones may represent different microhabitats of lagoonal settings, i.e. epidiceratid and
coral patch reefs, and the carbonate sands and mud in between. Ooid-oncoid grainstones and packstones of variable
grain size and degree of sorting may have been deposited in
the form of fringing ooid-oncoid bars along the margins of
the carbonate platform. Altogether, these sediments, preserved in numerous isolated klippen, provide a fairly complete view of the complex facies architecture of the Ernstbrunn-Pavlov Carbonate Platform.
In addition to the Jurassic rocks, the discordantly overlying Cretaceous strata and their relation to the Ernstbrunn
Limestone were studied (Glaessner 1930). Moreover, several fissure fillings inserted in the carbonates at Staatz and
Dörfles were analysed, and found to contain mixed assemblages of Tithonian, Late Cretaceous (Upper Campanian to
Santonian), Palaeogene and Neogene micro- and nanofossils (Bachmayer 1964a, Hofmann et al. 1999).
Closely related to the sedimentology of the rocks, the
present-day surface morphology and carstification of the
Jurassic klippen, including several small caves, have been
subjects of an unpublished Ph.D. thesis (Riedl 1958a) and
several publications (Riedl 1957, 1958b, 1960). The geography and underlying geologic structures of the Pavlov
Hills were detailed by Mikula (1927).
During the Middle Ages the Ernstbrunn Limestone has
been used as a building stone, e.g., in the castles of Staatz
and Falkenstein and the church of Michelstetten (Rohatsch
& Thinschmidt 1997). Since more than 100 years, the limestone is intensively mined at Ernstbrunn-Dörfles, and is
used as a natural resource for various construction materials and technical products (Bullinger 1997, Ernstbrunner
Kalktechnik 2011). For the first time, Moshammer &
Lobitzer (1997) have published data on the chemical composition of the Ernstbrunn Limestone along with its colorimetric properties, and thus provided important information
on the technical applicability of the rocks.
Palaeontology
The first fossil described from the Upper Jurassic rocks of
the Waschberg Unit is a “Gienmuschel” [old German trivial name for Chama; Adelung 1793–1801], i.e. a fossil bivalve of the genus Epidiceras (Haidinger 1782; Fig. 1A),
which is a character fossil of the Ernstbrunn Limestone.
Usually, these bivalves occur together with Heterodiceras
(Fig. 1B) as major components of the epidiceratid-nerineid
biofacies, and are preserved as internal moulds of doublevalved specimens (Schneider 2012). According to Bachmayer (1957), the fossil fauna of the Ernstbrunn Limestone comprises more than 500 species, which is rather an
underestimation according to our opinion (Fig. 7). Although several major taxonomic groups have not yet been
studied in detail, more than 300 taxa have been identified
and published.
As already outlined by Bachmayer (1940), the total
abundance of fossils and composition of the fauna vary
strongly among the different klippen and outcrops, which
can easily be demonstrated by the following examples.
(1) Corals are less diverse but much more abundant at
Klafterbrunn 3 quarry than at any other locality. (2) In contrast, epidiceratids and nerineids typically occur in abundance in all quarries at Dörfles (Fig. 1A, B) as well as at
Falkenstein and Mikulov Marienmühle (Mariánský mlýn)
quarry, but have not been recorded from Klafterbrunn,
Staatz, Turold, or Děvín. (3) Pharyngeal teeth of Lepidotes
are very common at Falkenstein (Fig. 8F), but rare at most
other localities. (4) Decapods are particularly abundant at
Dörfles I and Klement I quarries (Bachmayer 1940)
(Fig. 8B). (5) A major portion of the ammonites described
by Zeiss (2001) have been collected from Dörfles Werk II
quarry (see list below; Fig. 8D); a single specimen (described as Ernstbrunnia) comes from Galgenberg near
Michelstetten (Lower Austria).
In particular, the Dörfles quarries are famous for their
numerous large bivalves (Fig. 8C) and gastropods (Fig. 8I)
preserved as internal moulds. Careful investigation of
the limestone from Dörfles, however, predominantly reveals molluscs preserved with shell. Due to complete lithification of the carbonate sediment, the shells cannot be
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extracted and are thus usually not collected. Taking all
these observations into account, the different klippen and
outcrops of Ernstbrunn Limestone may represent different
habitats and facies, which may have experienced different
diagenetic histories (Schneider et al. 2011). Moreover, the
individual klippen may deviate in age. Up to now, these
differences have neither been systematically assessed nor
discussed in the literature.
Flora and fauna described
from the Ernstbrunn Limestone
and Klentnice beds
In the following overview only those publications which
employ scientific names at genus or species level are included.
Trace fossils. – Moshammer & Schlagintweit (1999) recorded a single microcoprolite taxon from thin sections of Ernstbrunn Limestone. Several types of borings in shells or coral
skeletons can be identified, which have been caused by Lithophaga, gastrochaenid bivalves, clionid sponges, and yet
unidentified microborers (personal observation SS).
Nanofossils. – Stradner (1961) recorded “Coccolithus cf.
pelagicus (Wallisch)” from unspecified “Tithonian strata”
from a well. However, since Coccolithus pelagicus (Wallich, 1877) Schiller, 1930 is an entirely Cenozoic taxon
(Bukry 1973), this is surely an artefact. Nine species of calcareous nanofossils were recorded by Holzknecht & Hamršmíd (1988) from the Mikulov Marls derived from a borehole near Mikulov.
Microflora. – To date, no macrofloral remains have been
discovered from the Ernstbrunn Limestone and Klentnice
beds. However, major portions of the Ernstbrunn Limestone are characterised by abundant calcareous algae.
Bachmayer (1944) described two new species of Dasycladales. Another two species of calcareous algae, again one
of them new to science, are detailed by Kamptner (1951).
Subsequently, Grill (1963) and Eliáš & Eliášová (1986)
listed calcareous algae from the Ernstbrunn Limestone.
Thirty-three different taxa of algae, some of them of enigmatic affinities, have been identified from the Dörfles
quarries by Hofmann (1990a, b, 1991a, b, 1992a, b,
1993b). Most of these taxa also belong to the Dasycladales
and one of them is regarded as a new species by Hofmann
(1994; emended by Schlagintweit 2011). Moshammer &
Schlagintweit (1999) also report five dasyclad species that
were collected from several smaller outcrops of Ernstbrunn
Limestone.
Foraminifera. – Similar to the algal flora, the microfauna
of the Ernstbrunn Limestone can only be assessed by studying thin sections of the rocks. Grill (1963) was the first
who studied Foraminifera from the Ernstbrunn Limestone
and he reported several taxa in open nomenclature from
Dörfles. Řehánek (1987a) listed six foraminifer taxa from
the Pavlov Hills, Hofmann (1990a) identified seven taxa
from Dörfles, and Moshammer & Schlagintweit (1999) recorded five taxa from a few other localities. Noth (1951)
mentions a foraminifer sample from the Upper Jurassic of
Niederleis, but does not provide additional information. A
list of Foraminifera from the Klentnice beds in Austria has
been provided by Grill (1953), while the Foraminifera of
an outcrop in Moravia have been monographed by Hanzlíkova (1965). Moreover, Papp & Turnovsky (1964) and
Holzknecht & Hamršmíd (1988) provided data on microfauna from drill cores taken from autochthonous Mesozoic
strata (Mikulov Marls).
Tintinnida. – Eliáš & Eliášová (1985) and Řehánek
(1987a) list more than 30 species of calpionellids from the
Ernstbrunn Limestone from the Pavlov Hills.
Porifera. – The “chaetetid” fauna of the Ernstbrunn Limestone comprises four species of these coralline sponges
(Bachmayer & Flügel 1961b; Fig. 6F). However, during the
following decades, many additional specimens have been
collected by Bachmayer and are stored in the collections of
NHMW. At several outcrops of the Klentnice beds a rich
fauna of silicified calcareous sponges occurs (personal observation AK, SS; Fig. 9B, I, J), which is unstudied.
Hydrozoa. – Four species of Hydrozoa from the Ernstbrunn Limestone, including a new species, were described
by Bachmayer & Flügel (1961a). Similar to the “chaetetids”, there is rich additional, unstudied material in the collections of NHMW.
Figure 6. Typical (micro-) facies of Ernstbrunn Limestone and Klentnice beds. • A – ooid-onkoid grainstone; Ernstbrunn Limestone; southern end of
Děvín; NHMW 2012/0106/0003. • B – bioclast-ooid packstone; upper part of Klentnice beds; eastern flank of Děvín; NHMW 2012/0106/0001.
• C – onkoid-grainstone; Ernstbrunn Limestone; Falkenstein quarry; NHMW 2011/0309/0001. • D – macro-ooid packstone; upper part of Klentnice beds;
eastern flank of Děvín; NHMW 2012/0106/0004. • E – calcareous sponge (Barroisia sp.); longitudinal section; Dörfles; NHMW 2012/0108/0006.
• F – coralline sponge (Chaetetopsis krimholzi Javorskij, 1947; det. F. Bachmayer); vertical section, with boring lithophagid bivalve in situ; Ernstbrunn
Limestone; Dörfles I; NHMW 2012/0110/0002. • G – bioclast wackestone; Ernstbrunn Limestone; Dörfles I; NHMW 2012/0110/0001. • H – marlstone
with ooids, peloids and sponge spicules; upper part of Klentnice beds; western flank of Děvín; NHMW 2012/0106/0002. Scale bars = 2 mm.
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Figure 7. Bar chart showing
estimated numbers of species
for higher taxonomic groups occurring in the Ernstbrunn Limestone and Klentnice beds. Estimates are based on literature
and collection surveys.
Scleractinia. – Moeller (1911) described 29 taxa of corals
from the Ernstbrunn Limestone, but did not provide figures. A single coral from the Ernstbrunn Limestone of Staatz
was reported by Kühn (1939). Only a small portion of the
rich fauna of Scleractinia from the Ernstbrunn Limestone
(Fig. 8H) has been studied by Eliášová (1986, 1990), since
she solely considered material from localities in the Czech
Republic. Although the major outcrops of this unit do not
preserve proper coral reefs, corals must have occurred in
small patch reefs (Eliášová 1986) or in non-biohermal assemblages, because they are present in considerable abundance and diversity. The now restored outcrop at Klafterbrunn likely exposed a proper coral reef (as interpreted by
Bachmayer 1940) or at least its parautochthonous debris,
since samples from this locality are predominantly composed of several taxa of well-preserved branching Scleractinia.
Bivalvia. – Epidiceratid bivalves were the first fossils to
be described and figured from the Ernstbrunn Limestone
(Haidinger 1782) and represent the dominant macrofossils
of the epidiceratid-nerineid facies at Dörfles (Fig. 1A, B).
They were mentioned by numerous authors working on the
stratum, and figured by several of them (e.g., Hofmann
1992b, Plöchinger & Karanitsch 2002, Wessely 2006). An
overview of the bivalve fauna was given in the unpublished
theses of Dürrmayer (1931), who identified 24 taxa, and
Bachmayer (1940), who extended this list to 35. Moreover,
Bachmayer (1948a) speculated about the mode of life of
the Epidiceratidae from Ernstbrunn. A survey of the material stored at the NHMW revealed approximately 70 species of Bivalvia (Fig. 8C, G).
Due to the loss of aragonite, the bivalve fauna of the
Klentnice beds is usually restricted to oysters and
pectinids. In places, however, relatively well-preserved,
mainly infaunal bivalves occur, which may reach a diversity of some 20 species (personal observation SS). A few
bivalves from the Klentnice beds at Niederfellabrunn have
been reported by Abel (1897), who also described a new
propeamussiid species. Furthermore, he recorded a species
of “Aucella”, which is a junior synonym of Buchia, a genus
that is regarded a typical constituent of boreal Upper Jurassic faunas (e.g., Kelly 1990). Together with several new
Figure 8. Typical fossils from the Ernstbrunn Limestone. • A – Metriomphalus transitorius (Zittel, 1873); rubber cast of external mould; Dörfles;
NHMW 2012/0108/0005. • B – Gastrodorus neuhausense von Meyer, 1864; specimen figured by Feldmann & Schweitzer (2009, fig. 3.8); Dörfles;
NHMW 1990z0041/4244. • C – Fimbria sp., internal mould. Dörfles; NHMW 2012/0108/0004. • D – Kutekiceras aff. steinbergense Zeiss, 2001;
microconch; rubber cast of external mould; Dörfles; NHMW 2012/0108/0001. • E – Eunerinea hoheneggeri (Peters, 1855); original of Wieczorek (1997,
pl. 1, fig. 8); Dörfles?; NHMW 1997z/0166/0005. • F – Lepidotes maximus Wagner, 1863; pharyngeal tooth; Falkenstein; NHMW 2012/0109/0001.
• G – Chlamys cf. textoria (von Schlotheim, 1820); Dörfles; NHMW 2012/0108/0007. • H – indeterminable colonial coral; internal mould; Dörfles I
quarry; NHMW 2012/0110/0003. • I – nerineid gastropod; internal mould; Dörfles; NHMW 2012/0108/0003. Scale bars: A, C–I = 10 mm; B = 1 mm.
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discoveries, these specimens were redescribed and refigured by Vetters (1905). Vašíček (1971) documented
nuculoid and gryphaeid bivalves from drill cores penetrating the autochthonous Klentnice beds below the AlpineCarpathian Foredeep in the Czech Republic.
Gastropoda. – The largest fossils preserved in the Ernstbrunn
Limestone are the more than 250 mm high Leviathania
gigantea (Makowsky, 1874), which were first described
from the Pavlov Hills (Makowsky 1874), figured by several authors (e.g., Bachmayer 1964b, 1969; Plöchinger &
Karanitsch 2002; Wessely 2006), and recently revised
by Harzhauser & Schneider (accepted). Most abundant,
however, are nerineid gastropods (Fig. 8E, I), several taxa
of which were recorded from the Ernstbrunn Limestone
of the Mikulov area by Peters (1855). Recently, the nerineids from the Ernstbrunn-Dörfles quarries have been
revised by Wieczorek (1998), who recognised 16 species. The total number of 45 gastropod species described
by Dürrmayer (1931) was raised to 52 by Bachmayer
(1940). The gastropod fauna of the Ernstbrunn Limestone (Fig. 8A) still awaits a comprehensive description,
which would likely more or less double the number of
species.
Because aragonitic shells have usually been dissolved,
no gastropods have been recorded from the Klentnice beds
so far, but a few unspecific internal moulds of turreted
shells from Südmährerkreuz (see Fig. 4 for location) are
stored in the collections of the NHMW.
Cephalopoda. – Altogether, four species of nautilids
(Hercoglossa spp.) have been reported from the Ernstbrunn Limestone (Bachmayer 1940, 1957; Dürrmayer
1931). An exceptionally well preserved specimen was figured by Bachmayer (1964b, 1969). The ammonite
fauna of the Ernstbrunn Limestone was revised by Zeiss
(2001), who also provided a review of earlier work dealing
with these fossils. These are the unpublished theses of
Moeller (1911), Dürrmayer (1931), and Bachmayer
(1940), as well as the fauna listed by Bachmayer (1957)
and Zeiss & Bachmayer (1989). Altogether, Zeiss
(2001) reported 37 species of ammonites in 14 genera
from Dörfles Werk II quarry (Fig. 8D), and a single specimen of Ernstbrunnia from the Galgenberg. Additionally, he recognised 13 more or less dubious taxa from
the literature, where specimens have been lost. The most
celebrated ammonite from the Ernstbrunn Limestone is
Ernstbrunnia fasciculata Zeiss, 2001, since it was figured on a stamp issued on the occasion of the 100th anniversary of the Naturhistorisches Museum Wien in 1976
(Bachmayer 1976, Thenius & Vavra 1996) and formed
the template of a logo of the Naturhistorisches Museum
Wien in the 1980s. The specimen was refigured by Hofmann (1995) and Wessely (2006).
Ammonites from the Klentnice beds, namely from the
surroundings of Niederfellabrunn, have been described by
Abel (1897), Krafft (1897, dealing with a single specimen),
Vetters (1905), Spath (1933), and Arkell (1956). Additionally, Vašíček (1971, 1980) detailed several species
from boreholes in southern Moravia. The latest revision of
the fauna was produced by Zeiss (1977) and recollected by
Kutek & Zeiss (1988). Additionally, several types of
aptychi from outcrops and drill cores were detailed (Bachmayer 1963; Vašíček 1971, 1980; Vetters 1905).
Furthermore, Abel (1897) lists several belemnite species from the Klentnice beds of Niederfellabrunn, but does
not provide figures. From the same outcrops, Vetters
(1905) reported on five species of belemnites and described and illustrated three of them as new.
Brachiopoda. – A few brachiopods from the Ernstbrunn
Limestone are listed by Prinzinger (1851). Suess (1858)
mentions three species from the Ernstbrunn Limestone
(one from Ernstbrunn and two from Mikulov) and four
from the Klentnice beds. Dürrmayer (1931) lists a single
species from the Ernstbrunn Limestone. Large terebratulids are relatively common at Dörfles V quarry, but have
never been studied. Brachiopods from the Klentnice beds
(Fig. 9A, H) are also mentioned by Hoernes (1874). Vetters
(1905) described and illustrated three species of terebratulids, while Jüttner (1922) provided figures of a single rhynchonellid taxon.
Ostracoda. – Ostracods have so far only been recorded
from the Klentnice beds of a single outcrop in the Mikulov
region. In two papers Pokorný (1959, 1971) provided preliminary results on the ostracod fauna from this locality and
formally described a new genus and a new species. Subsequently, Pokorný (1973) monographed the Ostracoda and
recorded a total of 56 species.
Figure 9. Typical fossils from the Klentnice beds. • A – terebratulid brachiopod; Klentnice (Table Mountain); NHMW 1931/0001/0041. • B–J – all
specimens (presumably) from Ernstbrunn (Semmelberg). • B – Corynella sp.; NHMW 2012/0107/0007. • C – Acropeltis aequituberculatus L. Agassiz in
L. Agassiz & Desor, 1846, aboral view; NHMW 2012/0107/0002. • D – millericrinid pluricolumnal, articular surface; NHMW 2012/0107/0004.
• E – Hemicidaris (Sphaerotiaris) sp. [= Hemicidaris crenularis auctt.]; lateral view of spine; NHMW 2012/0107/0005. • F – millericrinid pluricolumnal,
lateral view; with serpulid (lower left); NHMW 2012/0107/0006. • G – Plegiocidaris crucifera (L. Agassiz, 1840); oral view; NHMW 2012/0107/0001.
• H – rhynchonelliform brachiopod; NHMW 2012/0107/0009. • I – calcareous sponge; NHMW 2012/0107/0003. • J – calcareous sponge; polished section; NHMW 2012/0107/0008. Scale bars = 10 mm.
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Isopoda. – Bachmayer (1949) described three species of
Isopoda from the Ernstbrunn Limestone, including a new
genus and two new species. Two additional species were
newly established by Bachmayer (1955b) and the total
number of isopods was updated to seven taxa.
Decapoda. – The first decapod species from the Ernstbrunn Limestone has been listed by Moericke (1897). Later, Glaessner (1931) and Bachmayer (1945, 1955c) provided more or less comprehensive lists of crustaceans.
Moreover, Bachmayer (1948b) reported on teratologic decapod specimens. Bachmayer (1958b, 1959) described a
single new decapod. Wehner (1988) mentioned several
taxa from Ernstbrunn in her Ph.D. thesis on the “Prosopidae” (Brachyura). A few years ago, the several thousands of carapaces collected by Friedrich Bachmayer
during four decades were “rediscovered” in the collections of the NHMW Vienna and a wealth of species was
treated in numerous publications since then (Feldmann
& Schweitzer 2009; Robins et al. 2007, 2010, 2012,
2013; Schweitzer & Feldmann 2008a, b, 2009a–d,
2010a–d). Nevertheless, numerous additional forms are
hitherto unpublished.
Echinodermata. – The first short list of echinoderms
from the Ernstbrunn Limestone was provided by Prinzinger (1851). Much later, Bachmayer (1958a) reported
two crinoid species, one of them described as new. Remains of Saccocoma were mentioned from Tithonian
strata from a borehole by Řehánek (1987a). Besides, the
Ernstbrunn Limestone yields a moderately diverse fauna
of echinoids, including exceptionally large “regular”
and irregular taxa. From the Klentnice beds, Rolle
(1855) describes 9 species of “regular” echinoids. Moreover, crinoids and echinoids from the Klentnice beds are
listed by Hoernes (1874) and Jüttner (1922), who also figures several species. Within this unit large hemicidarid
spines (Fig. 9E) are most common, as are stem fragments
of millericrinid crinoids (Fig. 9D, F). Apart from Plegiocidaris (Fig. 9G) other echinoderm remains (Fig. 9C) are
generally rare.
Pisces. – Dürrmayer (1931) and Bachmayer (1940) list a
single species of fish, i.e. Lepidotes maximus Wagner,
1863. The typical circular teeth of these large fishes occur
in high abundance at Falkenstein (Fig. 8F) and were obviously also quite common in particular outcrops of the Mikulov region, judging from old collections at Vienna
(NHMW, GBA). Occasionally, the teeth are preserved
more or less in situ, while the bony parts of the skeleton
disintegrated entirely during diagenesis. Several small
pharyngeal teeth of unknown affinity and a few poorly
preserved shark teeth are present in the collections of
NHMW.
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Significance of the biota
Anybody having a close look at the “classical” Ernstbrunn
Limestone, i.e. the lagoonal facies of Dörfles, Falkenstein,
or Marienmühle at outcrop will easily recognize the outstanding palaeontological significance of this stratum,
simply because of its unusual richness in fossils (Fig. 7).
Looking at detail, the assemblage yields not just an enormous number of partly remarkably large specimens but
also an extraordinarily high diversity. Estimated to comprise more than 500 species (Bachmayer 1957), the fossil
assemblage of the Ernstbrunn Limestone ranges among the
most speciose Mesozoic environments ever recorded and
certainly deserves to be termed an Upper Jurassic marine
biodiversity hotspot (sensu Roberts et al. 2002; Fig. 7). However, up to the 21st century, this has largely remained “hidden diversity”, since despite of more than 40 years of intense, enthusiastic, and accurate collecting by Bachmayer
none of the most specious groups of fossils was detailed.
First and most important from a stratigraphical point of
view, the ammonites of the Ernstbrunn Limestone were
monographed (Zeiss 2001). Recently, the decapod faunas
from Ernstbrunn-Dörfles (represented by more than 10.000
specimens in the collections of the NHMW) and from the
coeval Štramberk Limestone (Silesian Unit, northern Moravia, Czech Republic) have been found to be of extraordinary significance for phylogenetic studies. The Tithonian
reefs and reef-associated strata at the northern Tethys margin may comprise an ancient radiation centre of the Decapoda and accordingly acted as a cradle for several major
clades of modern crabs (Feldmann & Schweitzer 2009;
Schweitzer & Feldmann 2008a, b, 2009a–d, 2010a–d).
Likewise, the Jurassic-Cretaceous lagoons on the Tethys
shelf may have played a key role in the development of the
Requienioidea, i.e. the left-valve-attached clade of rudist
bivalves (Hippuritida). Basal representatives of this group,
now placed in the family Epidiceratidae (Bieler et al.
2010), flourished in the lagoons of the Ernstbrunn-Pavlov
Carbonate Platform, forming small, paucispecific patch
reefs. Together with the highly abundant Nerineidae (Gastropoda), which also came to bloom during the Tithonian–Berriasian, and with a rich flora of dasyclad algae, the
Epidiceratidae dominate the biofacies of these lagoons.
Although the fossil assemblage preserved in the
Klentnice beds is much less spectacular due to the nearcomplete diagenetic loss of aragonite, the stratum is wellknown for its more or less strongly silicified, originally
calcitic skeletons, i.e. mainly echinoderms, brachiopods,
and calcareous sponges. At places ammonites or bivalves
add to a more complex view of the fauna, providing valuable insight into bottom life of the outer shelf area dominated by silciclastic sediments.
Information on inner and outer platform carbonates
(= Ernstbrunn Limestone) and on their intergrading with
Simon Schneider et al. Ernstbrunn Limestone and Klentnice beds state of the art and bibliography
siliciclastic sediments of the outer shelf (= Klentnice beds,
Mikulov Marls, Kurdějov Arenite) is available from several localities at surface and numerous cores. As a whole,
they offer an exceptionally complete view on the facies architecture on and around a Tithonian–Berriasian marginal
carbonate platform in the Tethys Ocean.
Strata of similar age and composition are fairly restricted in occurrence but geographically widespread,
which increases the scientific importance of the Jurassic–Cretaceous klippen of the WZU. A coeval highly diverse assemblage has been reported from the Štramberk
Limestone, cropping out in several klippen associated with
the Silesian and Subsilesian nappes of the Carpathians in
the Štramberk (NE Czech Republic) and Andrychow
(S Poland) regions (e.g., Boehm 1883; Zittel 1868, 1873).
However, the Štramberk Limestone predominantly preserves near-shore strata, and the more distal facies,
well-known from the Ernstbrunn Limestone and Klentnice
beds, is lacking. Lagoonal fossil assemblages of similar
age, diversity, and composition have been described from
Mont Salève (Hauté-Savoie, SE France; e.g., Joukowsky &
Favre 1913), Valfin (W Switzerland, Loriol & Bourgeat
1886–1888), southern Ukraine (Alth 1881, 1882) and the
Crimea peninsula (e.g., Pčelinzev 1924, 1963). Slightly
older, Lower Tithonian assemblages are reported from the
Palermo region (Sicily, Italy; Gemmellaro 1868–1876) and
the southern Franconian Alb (Kelheim and Ingolstadt regions; e.g., Boehm 1882). Most of these outcrops were less
intensely sampled and/or are significantly smaller than the
Ernstbrunn, Falkenstein and Pavlov klippen, making the
Ernstbrunn Limestone one of the most important strata for
exploration of these facies and their typical biota.
Future directions
Due to intensive drilling in the course of hydrocarbon exploration, knowledge of the standard geologic succession
of the Upper Jurassic rocks at the southern slope of the Bohemian Massif and the tectonic evolution of the WZU klippen seem largely settled. With regard to sedimentology, a
next crucial step would be a comprehensive study of the
(micro-)facies of all major and minor outcrops of the Ernstbrunn Limestone, in order to identify the total range of
existing lithofacies and to develop a facies model. Additionally, outcrops and historical wine cellars exposing the
Klentnice beds need to be systematically accessed and/or
excavated in order to figure out stratification and sedimentology.
With regard to palaeontology, a systematic account of
the Ernstbrunn fossil fauna and flora would be necessary, to
establish a solid base for comprehensive studies of palaeoecology, palaeoclimatology and palaeobiogeography. Extensive collections of fossils are stored in the palaeontological
repositories of the Natural History Museum Vienna and the
Moravian Museum at Brno, and to a minor extent at the
Geologische Bundesanstalt Wien, the University of Vienna,
and the Krahuletz Museum at Eggenburg. Preliminary data
from Bivalvia, which are usually reliable indicators of
(palaeo-)ecology suggest close faunistic and ecologic relationships to the coeval Štramberk Limestone, but also indicate significant differences in community structure, which
need to be evaluated in detail. Likewise, all other groups of
fossil organisms may add important information about the
Ernstbrunn ecosystem.
Finally, sedimentological and palaeontological data
should be integrated to enable a preferably complete reconstruction of the (1) size and extent, (2) facies architecture,
and (3) palaeoenvironment of the former Ernstbrunn-Pavlov Carbonate Platform. The resulting model may be compared to those from coeval carbonate platforms, in order to
identify their typical as well as unique features. Finally,
data may also be checked against those from earlier or later
ecosystems of similar composition to distinguish directional from non-directional developments.
Acknowledgements
We owe sincere thanks to the following people and organisations:
Thomas Hofmann (Geologische Bundesanstalt, Vienna) and
Radek Vodrážka (Czech Geological Survey, Prague) made numerous rare publications accessible. Fred Rögl (Natural History
Museum Vienna), Fritz F. Steininger (Eggenburg) and Godfried
Wessely (Vienna) provided information on regional geology.
Ulla Schudack and Michael Schudack (both Freie Universität
Berlin) provided information on micropalaeontology. Oleg
Mandic (Vienna) provided the photograph for Figure 5F. Alice
Schumacher (Natural History Museum Vienna) photographed the
fossils depicted in Figures 1, 8 and 9. Irene Zorn (Geologische
Bundesanstalt, Vienna) and Karl Rauscher (University of Vienna) enabled access to the collections under their care. Petr
Skupien (Vysoká škola báňská, Technická univerzita, Ostrava,
Czech Republic) organised and guided a highly informative field
trip to the Štramberk region attended by SS Careful reviews by
István Főzy (Hungarian Natural History Museum, Budapest)
and Matúš Hyžný (Department of Geology and Palaeontology,
Comenius University, Bratislava) helped to significantly improve
the manuscript. The work of SS was financially supported by the
Deutsche Forschungsgemeinschaft (DFG research fellowship
SCHN 1264/1-1).
References
As far as we are aware, this bibliography covers all aspects of
geoscientific research particularly concerned with the Ernstbrunn
Limestone and Klentnice beds at surface and subsurface. Furthermore, compilations of regional geology (e.g., Andrusov 1959;
Cicha et al. 1964; Fuchs 1976; Hauer 1875, 1878; Kober 1926;
121
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