18 NOAA Technical Report NMFS 18 A Temporal and Spatial Study of Invertebrate Communities Associated With Hard-Bottom Habitats in the South Atlantic Bight E. L. Wenner, P. Hinde, D. M. Knott, and R. F. Van Dolah November 1984 U.S. DEPARTMENT OF COMMERCE National Oceanic and Atmospheric Administration National Marine Fisheries Service NOAA TECHNICAL REPORTS NMFS The major responsibilities of the National Marine Fisheries Service (NMFS) are to monitor and assess the abundance and geographic distribution of fishery resources, to understand and predict fluctuations in the quantity and distribution of these resources, and to establish levels for optimum use of the resources. 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Individual copies may be obtained from: U.S. Department of Commerce, National Technical Information Service, 5285 Port Royal Road, Springfield, VA 22161. NOAA Technical Report NMFS 18 A Temporal and Spatial Study of Invertebrate Communities Associated With Hard-Bottom Habitats in the South Atlantic Bight E. L. Wenner, P. Hinde, D. M. Knott, and R. F. Van Dolah November 1984 u.s. DEPARTMENT OF COMMERCE Malcolm Baldrige, Secretary National Oceanic and Atmospheric Administration John V. Byrne, Administrator National Marine Fisheries Service William G. Gordon, Assistant Administrator for Fisheries The National Marine Fisheries Service (NMFS) does not approve, recommend or endorse any propriety product or proprietary material mentioned in this publication. No reference shall be made to NMFS, or to this publication furnished by NMFS, in any advertising or sales promotion which would indicate or imply that NMFS approves, recommends or endorses any proprietary product or proprietary material mentioned herein, or which has as its purpose an intent to cause directly or indirectly the advertised product to be used or purchased because of this NMFS publication. CONTENTS Introduction Materials and methods Collection of samples Data analysis Results and discussion Hydrographic measurements and description of study sites Large macrofauna sampled by dredge and trawl Biomass of larger macroinvertebrates Smaller macroinvertebrates sampled by suction and grab Conclusions Acknowledgments Literature cited Appendices . . . . . . . . . . . . . I I I 2 3 3 3 8 8 13 15 15 17-104 Figures 1. Location of hard-bottom stations sampled during 1981 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2. Normal cluster dendrogram showing station groups formed using the Jaccard similarity coefficient and flexible sorting. . . . . . 3. Inverse classification hierarchies and nodal diagram showing constancy and fidelity of station-species group coincidence based on seasonal dredge collections. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4. Inverse classification hierarchies and nodal diagram showing constancy and fidelity of station-species group coincidence based on seasonal trawl collections. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5. Normal cluster dendrogram of station groups formed using the Bray-Curtis similarity coefficient, log transformation, and flexible sorting . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 6. Inverse classification hierarchies and nodal diagram showing constancy and fidelity of station-species group coincidence based on seasonal suction and grab collections. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 4 6 8 13 15 Tables 1. Number of species and percent contribution for each taxonomic group collected by dredge and trawl at each station and season. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2. Species groups resulting from numerical classification of data from samples collected by dredge during 1981 3. Species groups resulting from numerical classification of data from samples collected by trawl during 1981 4. Percent contribution to epifaunal biomass of taxonomic groups in dredge and trawl collections. . . . . . . . . . . . . . . . . . . . . . . . . . 5. Average density and percent of total number for 10 numerically dominant invertebrates collected by suction and grab. . . . . . . 6. The 10 most abundant invertebrate species collected by suction and grab samplers at each station . . . . . . . . . . . . . . . . . . . . . . . 7. Seasonal community structure values for pooled replicate samples of invertebrates at each station .... . . . . . . . . . . . . . . . . . . . 8. Species groups resulting from numerical classification of data from samples collected by suction and grab samplers during all seasons, 1981 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . iii 3 5 7 9 10 II 12 14 A Temporal and Spatial Study of Invertebrate Communities Associated With Hard-Bottom Habitats in the South Atlantic Bight l E. L. WENNER, P. HINDE, D. M. KNOTT, and R. F. VAN DOLAH2 ABSTRACT Species composition, biomass, density, and diversity of bentblc invertebrates from six bard-bottom areas were evaluated. Seasonal collections using a dredge, trawl, and suction and grab samplers yielded 432, 525, and 845 taxa, respectively. Based on collections wltb tbe diCCerent gear types, species composition of invertebrates was found to cbange batbymetrically. Inner· and mlddle-sbelf sites were more similar to eacb otber In terms of invertebrate species composition tban tbey were to outer-sbelf sites, regardless of season. Sites on tbe Inner and outer sbelf were grouped according to latitude; bowever, results suggest tbat deptb Is apparently a more important determinant of Invertebrate species composition tban eltber season or latitude. Sponges generally dominated dredge and trawl collections In terms of biomass. Generally, cnldarlans, bryozoans, and sponges dominated at sites In terms of number of taxa collected. Tbe most abundant smaUer macrofauna collected In suction and grab samples were polycbaetes, ampbl. pods, and mollusks. Densities of tbe numerically dominant species cbanged botb seasonally and batbymetrlcally, witb very few of tbese species restricted to a specific batbymetrlc zone. Tbe blgb diversity of invertebrates from bard·bottom sites Is attributed to tbe large number of rare species. No consistent seasonal cbanges in diversity or number of species were noted for individual stations or deptb zones. In addition, n' and Its components sbowed no definite patterns related to deptb or latitude. However, more species were collected at mlddle-sbelf sites tban at Inner- or outer·sbelf sites, wbicb may be related to more stable bottom temperature or greater babltat complexity In tbat area. INTRODUCTION Although other studies have provided species lists of invertebrates from hard-bottom areas [see Continental Shelf Associates (1979) for literature review], Wenner et al. (1983) provided the first detailed information on structure of hard-bottom invertebrate communities in the South Atlantic Bight. Results indicated that the invertebrate fauna of these areas was more diverse than the surrounding sand biotope, Species composition was apparently related more to station depth than to season, although species associations did vary between winter and summer. Because sampling during their study was limited to only two seasons, it was not possible to determine seasonal trends in community composition, structure, and biomass which might be occurring over an annual cycle. This paper presents results from a second-year study of invertebrates associated with hard-bottom habitats anq focuses on seasonal as well as bathymetric changes in community parameters. Comparisons with results from the initial study of hard-bottom sites by Wenner et al. (1983) provide additional information on annual variations in the epibenthic invertebrate community. 'Contribution No. 178 from lbe Marine Resources Research Institute, Charleslon, S.C. 'Marine Resources Research Institute, P.O. Box 12559. Charleston, SC 29412. MATERIALS AND METHODS Collection of Samples Two study sites were selected between lat. 31 °24' and 32°49'N from each of three bathymetric zones: Inner shelf (ISO I, 17-18 m; IS02, 17-22 m), middle shelf (MS02, 23-29 m; MS06, 32-36 m), and outer shelf (OSOI, 59-67 m; OS06, 46-69 m) (Fig. 1). Four of the sites (ISOl, IS02, MS02, OSOI) were previously assessed during the initial study (Wenner et al. 1983). All sites were sampled quarterly during 1981 as follows: Winter (25 February to 17 March), spring (26 April to 28 May), summer (25 July to 13 August), and fall (6 October to 16 November). Prior to sampling for invertebrates, hydrographic profiles of temperature, salinity, and dissolved oxygen were obtained at each site by Niskin bottles with reversing thermometers. Qualitative samples of large and encrusting macroinvertebrates were taken quarterly with a heavy duty Cerame- Vivas dredge (Duke University Marine Laboratory 1968) having a mouth opening 90 cm x 37 cm and maximum mesh opening 40 mm x 30 mm. Two replicate dredge tows were attempted at each station with a total of 43 actually completed. Tow length was standardized to approximately 0.1 km using 10ran-C positioning. Additional qualitative collections of large macroinvertebrates were obtained as incidental catch by a trawl (12 m headrope, 16 m footrope, 12.8 m vertical height) equipped with steel doors and rubber rollers. Six replicate tows (three day and three night) were attempted quarterly at all sites except station OS06, where high bottom relief prevented use of the trawl. Altogether, 119 trawl tows were completed. Trawl tows were standardized to a distance of about 1 km. . \ \ each sample was narcotized with 10% magnesium chloride and preserved in a 10% seawater-Fonnalin solution. At each outer-shelf station, where water depth precluded the use of a diver-operated suction device, five quantitative 0.1 m2 samples were attempted with a Smith-McIntyre grab. Only 33 samples were collected due to ineffective sampling of the grab on hard substrates. After retrieval, each sample was placed on a 1.0 mm sieve and washed to remove the finer sediment prior to preservation in 10% buffered seawater Fonnalin. Macroinvertebrates collected by suction and grab samplers were identified and counted, except for encrusting fauna which were not easily quantified by counting (e.g., hydroids and colonial corals). \ j tI North / Carolina Data Analysis Community diversity was estimated by the Shannon index (It) (Pielou 1975) in conjunction with species richness (SR) (Margalef 1958), and evenness (J) (Pielou 1975). These measures were computed on data from pooled replicates taken with suction or grab samplers at each station, since pooling the replicates provided a larger sample size and a more representative estimate of community diversity at a site. The degree of dominance at a station was quantified with the dominance index (01) (McNaughton 1967). For qualitative collections by dredge and trawl, diversity was evaluated by comparing the number of species (s) among stations. The Kruskal- Wallis one- way analysis by ranks (Siegel 1956) was used to detennine whether s differed significantly between stations or seasons. Cluster analysis was used to detennine patterns of similarity among stations (pooled replicated collections) and among species using qualitative binary data (i.e., species presence or absence) for dredge and trawl samples, and quantitative abundance data for suction and grab samples. Due to the large number of species in collections made by the various sampling devices, it was necessary to reduce data sets which contained> 200 species prior to cluster analysis in order to remain within the computational limits of available computer programs. Data sets were reduced by both elimination of species which were infrequently collected and elimination of taxa having undetennined or questionable identity. The elimination of these taxa from our data sets was justifiable because "rare" species usually do not have definable distribution patterns and can confuse interpretation of cluster analysis. Data sets subjected to cluster analysis were reduced to include only those species represented in 5 or more dredge collections, 9 or more trawl collections, and 12 or more suction and grab collections. Species and collections were classified using flexible sorting (Lance and Williams 1967) with a cluster intensity coefficient (fJ) of -0.25. The Jaccard similarity coefficient (Clifford and Stephenson 1975) was used with presence-absence data collected by dredge and trawl. Species abundance data collected by suction and grab were transfonned using log 10 (x + 1) and clustered using the Bray-Curtis measure of similarity (Bray and Curtis 1957). Since attribute values covered a wide range, a strong transfonnation was used to reduce the sensitivity of the Bray-Curtis measure to larger values (Clifford and Stephenson 1975). Subsequent to cluster analysis, species and station groups were chosen using a variable stopping rule (Boesch 1977). Then nodal analyses (Williams and Lambert 1961; Lambert and Williams 1962) were employed to describe pooled collections at a station in tenns of their characteristic species and to describe species groups resulting from inverse cluster analysis in tenns of their patterns of occurrence at stations (Boesch 1977). Coincidence of station and / M502. 0501. I i ! Florida Figure I.-Location of bard-bottom stations sampled during 1981. Invertebrates collected by dredge and trawl were sorted into the following major taxonomic groups for wet-weight estimates: Porifera, Hydrozoa, Scleractinia!Octocorallia, Mollusca, Decapoda, Echinodennata, Ascidiacea, and other miscellaneous invertebrates. Encrusting organisms which could not easily be removed from rocks were not weighed. Representative specimens of each species were preserved in 10% buffered seawater Formalin and transferred to 70% isopropanol prior to identification. Quantitative samples of smaller benthic invertebrates not adequately sampled by dredge and trawl were obtained at inner- and middle-shelf sites by scuba divers using an airlift suction sampling device similar to that described by Chess (1979). Divers obtained five replicate samples at each station from a suitable area chosen to avoid large patches of sand commonly found at the sites. A disc with five equally spaced radial marks was dropped to the bottom and a 3 m line, fastened to the center of the disc, was then used to place five open-ended quadrat boxes (0.1 m2 , 10 cm walls) equidistantly around the disc. Exact positioning of the quadrat boxes was accomplished by randomly selecting one of nine possible quadrat areas from a larger grid frame attached to the 3 m line. Fauna within the quadrat was sampled by scraping the area while simultaneously sucking with the airlift suction device to a depth of 10 cm or until rocky substrate was encountered. Suction was applied until all sediment within a quadrat was removed to the designated depth. All suction samples were collected in 1.0 mm mesh bags and brought to the surface for preservation. On deck, 2 species groups was expressed by graded constancy and fidelity values, which were used to determine those species that were most consistelltly collected and most restricted to pooled collections at a station. The mathematical definitions for constancy and fidelity are found \n Boesch (1977). respect to their bottom topography. Station OSOI was a low relief hardground with only occasional exposed rock of little relief and relatively sparce epifaunal density. In contrast, OS06 was a typical shelf edge reef, with irregular bottom topography characterized by ledges and extensive rock outcropping of moderate to high relief which were often heavily encrusted with invertebrates. RESULTS AND DISCUSSION Large Macrofauna Sampled by Dredge and Trawl Hydrographic Measurements and Description of Study Sites Seasonal collections of epifauna from hard-bottom sites sampled by dredge and trawl yielded 432 and 525 taxa, respectively. A list of the identified taxa, arranged phylogenetically for each station and sampling gear, is found in Appendices I and 2. The Bryozoa (91 taxa), Porifera (89 taxa), and Cnidaria (70 taxa), dominated dredge collections from all seasons in terms of numbers of taxa. Porifera (111 taxa) were also well represented in trawl collections, along with other taxa such as decapod crustaceans (86 taxa), Bryozoa (85 taxa), and Mollusca (85 taxa). Both the present and previous study (Wenner et al. 1983) of hard-bottom communities have shown that Porifera, Bryozoa, and Cnidaria are the most diverse taxonomic groups encountered. The only invertebrate groups whose percent contribution of species in dredge and trawl collections changed noticeably across the continental shelf were the Porifera, Cnidaria, and Bryozoa (Table 1). In terms of number of taxa, sponges were most important at sites on the inner shelf. Their richness diminished at middle- and outer-shelf sites where cnidarian and bryozoan taxa dominated. No seasonal trends in percent composition among taxa were noted. The number of invertebrate taxa (s) collected at each station by dredge and trawl did not change significantly between seasons (Kruskal-Wallis test, P > 0.05). These results were consistent with those reported by Wenner et al. (1983) who found no significant difference in s between winter and summer sampling periods. However, in the present study we did find a significant difference Hydrographic parameters at the hard-bottom sites were similar to those reported by Wenner et al. (1983). Bottom temperatures on the inner shelf were more extreme (I 1.7 °-27.3 0c) than those on the outer shelf (17.0 o -23.9°C). Temperatures at middle-shelf sites were similar to those encountered at outer-shelf sites (16.0 o -23.4°C). Bottom salinity and dissolved oxygen were generally high at the sites during all seasons, ranging from 34.3 to 36.6% 0 and 3.9-6.4 mVI, respectively. Television transect videotapes showed that the bottom at station ISO I was smooth, flat, and overlaid by a layer of sand. Sessile invertebrates at ISOI were numerous and uniformly covered the broad expanse of sand-covered hardpan. Station IS02 had a more irregular bottom than ISOI with numerous rock ledges and discontinuous outcroppings up to 1 m in height. These outcroppings were distributed throughout areas of sand and were covered with a dense assemblage of epifauna, particularly adjacent to the ledges. Middle-shelf sites were characterized as having moderate relief ledges, although there were also areas of low rocky relief interspersed with sand. At MSOl, several ledges up to I m in height were observed, whereas ledges with a vertical drop of nearly 2 m were observed at MS06. Epifaunal density was usually high on emergent rock of moderate to high relief at MS06. Outer-shelf stations, which were located in the vicinity of the shelf break, wer;e distinctly different from one another with Table i.-Number of species and percent contribution for each taxonomic group collected by dredge (D) and trawl (T) at each station and season. Number samples Porifera Cnidaria Mollusca Cirripedia Decapoda Echinodermata Bryozoa Tunicata Station Season D T No. % No. % No. % No. % No. % No. % No. % No. % Total number ISOI Winter Spring 2 2 2 2 I 2 2 2 1 2 2 2 2 2 2 2 2 2 2 2 6 6 6 6 5 6 6 6 6 6 6 6 6 6 6 6 6 6 6 6 45 25 33 21 49 49 40 51 29 30 IS 24 20 38 36 38 26 29 28 36 28.1 17.9 20.8 15.7 24.4 25.5 24.2 27.4 17.7 14.2 8.8 14.2 12.4 16.2 20.3 19.7 13.7 15.3 14.0 18.7 26 31 36 42 37 25 37 37 38 37 43 44 43 49 45 46 54 48 53 SO 16.2 22.1 22.6 31.3 18.4 13.0 22.4 19.8 23.2 17.5 25.1 26.0 26.7 20.8 25.4 23.8 28.4 25.4 26.5 25.9 16 12 17 10 20 23 IS 20 16 28 14 13 IS 34 14 16 17 14 16 23 10.0 8.6 10.7 7.5 9.9 11.9 9.1 10.8 9.8 13.3 8.2 7.7 9.3 14.5 7.9 8.3 8.9 7.4 8.0 11.9 4 5 4 5 4 5 4 4 4 5 4 3 3 3 4 3 4 6 5 6 2.5 3.6 2.5 3.7 2.0 2.6 2.4 2.1 2.4 2.4 2.3 1.8 1.9 1.3 2.3 1.6 2.1 3.2 2.5 3.1 IS IS 12 12 25 22 20 IS IS 29 22 IS 17 17 13 18 25 35 39 25 9.4 10.7 7.5 8.9 12.4 II.S 12.1 8.1 9.1 13.7 12.9 8.9 10.6 7.2 7.3 9.3 13.2 18.5 19.5 13.0 8 9 10 6 7 13 7 8 IS 10 7 9 6 16 13 8 9 11 13 9 5.0 6.4 6.2 4.5 3.5 6.8 4.2 4.3 9.1 4.7 4.1 5.3 3.7 6.8 7.3 4.1 4.7 5.8 6.5 4.7 29 27 29 25 43 37 28 35 32 SO SO 43 49 63 43 54 44 38 38 32 18.1 19.3 18.2 18.7 21.4 19.3 16.9 18.8 19.5 23.6 29.2 25.4 30.4 26.8 24.3 27.9 23.2 20.1 19.0 16.6 17 16 18 13 16 18 14 16 IS 22 16 18 8 IS 9 10 11 8 8 12 10.6 11.4 11.3 9.7 7.9 9.4 8.5 8.6 9.1 10.4 9.4 10.6 4.9 6.4 5.1 5.2 5.8 4.2 4.0 6.2 160 140 159 134 201 192 165 186 164 211 171 169 161 235 177 193 190 189 200 193 3.6 11.2 40 22 39 38.5 26.5 31.2 4 4 7 3.8 4.8 5.6 2.9 3 14 2 6 11 1.9 7.2 8.8 I 8 8 0.9 9.6 6.4 54 34 42 51.9 40.9 33.6 6 3 7.2 2.4 104 83 125 Summer IS02 MS02 Fall Winter Spring Summer Fall Winter Spring Summer MS06 OSOI OS06 Fall Winler Spring Summer Fall Winter Spring Summer Fall Winler Spring Summer Fall 2 I 2 3 0.8 in s between stations sampled (Kruskal-Wallis test, P < 0.01) by trawl. Although the fewest species were collected at IS01, this is not necessarily indicative of generally lower richness at inner shelf stations, nor do these results imply that s is influenced solely by depth or latitude, since little difference was noted between numbers of species collected at IS02, middle-shelf sites, and OS01. Increases (or decreases) in s are probably also influenced by localized factors at each site such as topographic complexity and habitat heterogeneity. The extensive hard-bottom area of ISOI had little vertical relief compared with IS02, which was characterized by numerous rock ledges supporting dense epifaunal populations. These rock ledges provide complex microhabitats which allow many epifaunal species to coexist (Schoener 1974; Menge and Sutherland 1976). In addition, many sessile species settle in greater abundance on structurally complex substrata because there is more surface area to which they can securely attach (Seed 1976; Russ 1980; Dean 1981). Normal cluster analysis indicated that pooled replicate dredge and trawl collections were classified into three station groups corresponding to inner-shelf sites (group 1), middle-shelf sites (group 2), and outer-shelf sites (group 3) (Fig. 2). Stations were not grouped in a manner which would indicate that species composition changed seasonally in a consistent pattern. Even within each site group, no grouping of stations by season was evident. This DREDGE: STATION suggests that the qualitative composition of the epifaunal community sampled by dredge and trawl was generally similar throughout the year. In addition, there was no indication that stations were grouped in accordance with their latitudinal location on the continental shelf. Comparison with results of normal cluster analysis from the initial study indicates that depth is apparently a more important determinant of faunal composition than season or latitude at hard-bottom sites off South Carolina and Georgia. These results suggest that the epifaunal community of hard-bottom areas are persistent throughout the year. Osman (1977) suggested that changes in the composition of epifaunal invertebrates occur slowly. Growth and reproduction rate of epifauna are reduced during the winter with the result that few new species are added, but several are lost. Inverse cluster analysis of the 199 most frequently occurring species from seasonal dredge collections formed 10 groups (Table 2). Nodal analysis indicated that the three major branches of the inverse cluster dendrogram represent species assemblages found primarily at inner-shelf sites, those found across the shelf, and those found primarily at middle- and outer-shelf sites (Fig. 3). Species which were most consistently collected at inner-shelf stations partially comprise groups A, C, and D. However, these assemblages were not restricted to sites on the inner shelf, as indicated by moderate fidelity values. Species in these groups which were collected only at inner-shelf sites included the sponges Homaxinella waltonsmithi and Tedania ignis; the echinoderms Lytechinus variegatus, Encope michelini, and Luidia alternata; and the mollusk Pisania tincta. Except for the sponges H. waltonsmithi and T. ignis, these species are found on a wide variety of bottom types and are not limited to hard substrates. Although the cluster hierarchy indicated that group Band C species were most similar in their occurrence at hard-bottom sites, the nodal diagram illustrates differences in the distribution of species in these two groups. Species in group B were most constant at station OSO 1; whereas species in group C were most constant at station IS02. Species in groups E and F were consistently collected at most sites sampled. Group E species were highly constant at all sites and included the most ubiquitous species collected. Many of the species in this group were dominant epifaunal organisms in terms of their frequency of occurrence at hard-bottom sites across the shelf. However, because of their ubiquity they were not faithful to any site group. Species in group F were also fairly ubiquitous but were more commonly encountered at inner- and middle-shelf sites. Species groups G-J were most consistently collected at middleand outer-shelf sites. Group G species displayed high constancy at stations MS02 and MS06; however, the only species in this group which was collected exclusively at middle-shelf hard-bottom stations was the bryozoan Schizoporella errata. Group H species also were common at station MS06 but were most consistently encountered at station OS06. Species groups I and J displayed highest constancy and fidelity at outer shelf sites; however, only species in group J were restricted to these locations, as indicated by their very high fidelity values. Species in groups I and J which were collected exclusively at outer-shelf sites included the cnidarians Balanophyllia floridana, Diodogorgia sp., Ellisella sp., and Stylatula sp.; the decapod crustaceans Galathea rostrata, Dardanus insignis, and Parthenope fraterculus; the bryozoan Membranipora arborescens; and the echinoderm Astroporpa annulata. Comparisons of species assemblages identified by cluster and nodal analyses between the present and the previous study (Wenner et al. 1983) have shown that several species are most frequently encountered in one depth zone. Those which typified GROUPS SIMILARITY Group Season Fa W; ISOI w; 5, 5, Fa 1502 1502 ISOI Sp -WiSp 5, w; 2 Sp Fa 5, Fa W; Fa 5' Sp :3 Station Fa Sp 5, O.L ,8-' 0jL. 6-' l°i4_'- O '-i2_ - ~_ ' -_ O/ 1502 1502 Sp ISOI ISDI -MS02 MSQ2 MS02 MSQ6 MSQ6 MSQ6 MS06 M502 0501 -0501 0501 0506 -= ~: J 0506 0501 0506 TRAWL: STATION GROUPS SIMILARITY Group Season W, Sp 5, Fa W; Sp 5, Fa 5, Fa 2 :3 Sp Sp Stotion W; Fa Wi 6501 5, Fa 0501 w, Sp 0.8 , 0.6 0.4 , 0.2 a, -0.2 , ISOI ISOI ISOI ISOI 1502 1502 1502 1502 MS06MS06 MS02 MSQ6 MS02 MS02 M$06 M$02 5, 1.0 , 0'301 OSOI Figure 2.-Normal cluster dendrogram showing station groups formed using the Jaccard similarity coemclent and flexible sorting. Data from replicate dredge and trawl collections were pooled for each station prior to cluster BRalysis. 4 Table 2.-Specles groups resulting from numerical c1asslftcatlon of data from samples collected by dredge during 1981. (Ar Cnldarla; Ech Echinodermata; Mo MoUusca; Po Porifera.) data; Cn = Group A Lissodendoryx sigmala (Po) Clavelina giganlea (Ch) Scypha barbadensis (Po) Clalhrina c('riacea (Po) Pilumnus dasypodus (Ar) Celleporina hassalli (Bry) Membranipora lenuis (Bry) Synalpheus minus (Ar) Dislaplia bermudensis (Ch) Ocnus pygmaeus (Ech) Homaxinella wallonsmilhi (Po) Lylechinus variegalus (Ech) Crepidula aculeala (Mo) Styela partita (Ch) Celleporaria mordax (Bry) Bimeria humilis (Cn) Symplegma viride (Ch) Trididemnum savignii (Ch) Homaxinella sp. A (Po) Diplosoma macdonaldi (Ch) EudislOma carolinense (Ch) Sertularia dislans (Cn) Group B Microciona prolifera (Po) Teichaxinella corrugala (Po) Pseudaxinella lunaecharta (Po) Arca zebra (Mo) Axinella bookhouli (Po) Pandaros acanlhifolium (Po) Diodora cayenensis (Mo) Murex jlorifer dileclus (Mo) Slenocionops furcala coelata (Ar) Leplogorgia sp. (Cn) Antipalharia (Cn) Membraniporella aragoi (Bry) Eucidaris Iribuloides (Ech) Pachycheles rugimanus (Ar) Group C Spheciospongia vesparium (Po) Lissodendoryx isodictyalis (Po) Leodia sexiesperjorala (Ech) Cliona caribbaea (Po) Cinachyra kuekenthali (Po) Aplysina fislularis (Po) Aplousina tuberosa (Bry) Amphimedon compressa (Po) Thalysias juniperina (Po) Ircinia strobilina (Po) Synalpheus townsendi (Ar) Hiatella arctica (Mo) Amalhia allernata (Bry) = = = Anlropora tin cIa (Bry) Turbicellepora dicholoma (Bry) Clytia fragilis (Cn) Balanus venustus (Ar) Cribrilaria radiata (Bry) Microporella umbracula (Bry) Aelea anguina (8ry) Amalhia distans (Bry) Celleporaria albirostris (8ry) Pa'rasmittina spathulata (Bry) Teleslo sanguinea (Cn) Cleidochasma porcellanum (Bry) Halecium lenellum (Cn) Sertularia plumulifera (Cn) Halecium dysymmetrum (Cn) Pseudomedaeus agassizii (Ar) Floridina anliqua (Bry) Encope michelini(Ech) Pisania tincla (Mo) Hemectyon pearsei (Po) Dromidia an/illensis (Ar) Oculina sp. (Cn) Halichondria bowerbanki (Po) Ciocalapata gibbsi (Po) Ircinia campana (Po) Ircinia felix (Po) Arbacia punclulata (Ech) Higginsia slrigilala (Po) Ircinia ramosa (Po) Geodia gibberosa (Po) Group D Group F Homaxinella rudis (Po) Cinachyra alloclada (Po) Spiraslrella coccinea (Po) Cinachyra rhizophyla (Po) Luidia alternala (Ech) Tedania ignis (Po) Phakellia lobata (Po) Haliclona oculata (Po) LeplOgorgia virgulata (Cn) Vennicularia knorrii (Mo) Hippaliosina rostrigera (8ry) Reptadeonella haslingsae (8ry) Conopea merrilli (Ar) Pilumnus sayi (Ar) Celleporaria magnifica (8ry) Styela plicala (Ch) Pteria colymbus (Mo) Molguia occidentalis (Ch) Modiolus american us (Mo) Arca imbricala (Mo) Lophogorgia hebes (Cn) Mithrax pleuracanlhus (Ar) Eudendrium carneum (Cn) Macrocoeloma camptocerum (Ar) Pseudochama rodians (Mo) Haliclona sp. C (Po) Nolella slipala (Bry) Buskia sp. (Bry) Xytopseus griseus (Po) Sigmadocia caerula (Po) Group E Didemnum candidum (Ch) Chama congregata (Mo) Monostaechas quadridens (Cn) Microporella ciliata (Bry) Obelia dichotoma (Cn) Schizoporella cornula (Bry) Tryposlega venuSla (Bry) Tilanideum frauenfeldii (Cn) Balanus trigon us (Ar) Halecium sp. (Cn) TeleslO fruticulosa (Cn) Ophiolhrix angulata (Ech) Dynamena cornicina (Cn) Crisia sp. (Bry) Sertularia marginata (Cn) Bugula rylandi (Bry) Aglaophenia latecarinata (Cn) Salacia desmoides (Cn) Clytia cylindrica (Cn) Aglaophenia trifida (Cn) Senularella conica (Cn) Sertularella sp. A (Cn) Dynamena quodridentata (Cn) Group G Synthecium tubitheca (Cn) Petraliella bisinuata (Bry) Bugula fulva (8ry) Schizoporella jloridana (8ry) Aeverrillia seligera (8ry) Thyroscyphus marginatus (Cn) Chaperia sp. (8ry) Diaperoecia jloridana (8ry) Parellisina curviroslris (8ry) Cupuladria doma (8ry) Cupuladria canariensis (8ry) Crepidacanrha poissoni teres (Bry) Megalobrachium sorialum (Ar) Smittina smitliella (8ry) Schizoporella errala (8ry) Leuconia aspera (Po) Lyonsia beana (Mo) Alcyonidium sp. (8ry) Podochela riisei (Ar) Gonodactylus bredini (Ar) 5 = Arthropoda; Bry = Bryozoa; Ch = Chor· Group H Parellisina latiroSlris (8ry) Nellia len ella (8ry) Nemerlesia simplex (Cn) Caberia boryi (8ry) Microporella marsupiata (8ry) Proboscina jloridana (8ry) Gymnangium sinuosum (Cn) Cribrilaria jloridana (Bry) Lichenopora radiata (Bry) Exochella longirostris (8ry) Clypeasler subdepressus (Ech) Floridina parvicella (Bry) Turrilopsis nutricula (Cn) Hincksella cylindrica (Cn) Group I Bougainvillia sp. (Cn) Serlularella areyi (Cn) Antenella secundaria (Cn) Eudendrium tenellum (Cn) Dynamena dalmasi (Cn) Balanus calidus (Ar) Aplousina gigantea (8ry) Filellum serratum (Cn) Stylopoma informata (8ry) Smitlipora levinseni (8ry) Campanularia hincksii (Cn) Thesea sp. (Cn) Hippoporina contracta (Bry) Scandia mUlabilis (Cn) Hebella venusta (Cn) Hebella scandens (Cn) Mithrax acuticornis (Ar) Galalhea roslrala (Ar) Amalhia convoluta (8ry) Alderina smitti (Bry) Clenoslomala (8ry) Scrupocellaria regularis (8ry) Aglaophenia elongala (Cn) Group J Dardanus insignis (Ar) Membranipora arhorescens (Bry) Halopteris clarkei (Cn) Cycloperiella rubra (8ry) Astroporpa annulala (Ech) Stylatula sp. (Cn) Balanophyllia jloridana (Cn) Ellisella sp. (Cn) Diodogorgia sp. (Cn) Parlhenope fraterculus (Ar) SIMILARITY -l.8 -1.4 -1.0 -0.6 ,!, " STATIONS -O.Z ISO I IS02 MS02 MS06 OSO, OS06 r---- A A B C c 0 o In c.. ::J 0 a:: (!) E In W U W c.. In G H H I I FIDELITY CONSTANCY . . 20.1 Very Hioh (]]J20,1 _ D<O.IV.., Low ~ LO.5 HiOh Low ~2 ,20.3 Moderot. Moderot. Figure 3.-Inverse classification hierarchies and nodal diagram showing constancy and fidelity of station-species group coincidence based on seasonal dredge collections. inner-shelf hard-bottom sites included the sponges Homaxinella waltonsmithi, Spheciospongia vesparium, Cliona caribbaea, and Halichondria bowerbanki; the echinoderms Encope michelini, Lytechinus variegatus, Ocnus pygmaeus, and Arbacia punctulata; the bryozoan Membranipora tenuis; and the decapod crustacean Synalpheus minus. Middle-shelf hard-bottom sites were characterized by the bryozoans Chaperia sp., Aeverrillia setigera, Schizoporella floridana, Petraliella bisinuata, and Parellisina curvirostris; and the hydroids Thyroscyphus marginatus and Synthecium tubitheca. Species which were consistently collected at outer-shelf sites during both years of sampling by dredge included the bryozoans Smittipora levinseni, Stylopoma in/ormata, Cribrilaria floridana, and Cycloperiella rubra; the hydroids Dynamensa dalmasi and Filellum serratum; the echinoderm Astroporpa annulata; and the decapod crustacean Mithrax acuticornis. Few species assemblages identified by cluster analysis of dredge samples were restricted to anyone depth zone or hardbottom site. This suggests that few species assemblages, other than those displaying high fidelity values in nodal diagrams, could be considered characteristic of a particular depth zone in the South Atlantic Bight. Inverse cluster analysis of 184 species from seasonal trawl collections produced seven groups. Species and site group coincidences were interpreted in terms of nodal constancy and fidelity (Table 3, Fig. 4). Group A contained species which were characteristic of inner-shelf stations and station MS06. Species in group B also displayed high constancy for inner-shelf stations; however, no species in either group were restricted to specific hard-bottom sites. Only the tunicate Ascidiacea A was collected exclusively at inner-shelf sites. Groups C, D, and E were composed of species which were fairly ubiquitous at hard-bottom sites across the shelf. Species in group C were highly constant at inner- and middle-shelf stations, whereas those in group D were consistently collected at every live-bottom site sampled. This group contained many of the most frequently occurring epifaunal species collected during the study. Group E contained species which displayed high constancy at stations IS02, MS02, and MS06. Species in group F were highly constant at station MS06, but were not restricted to this station. Those species in group G were most frequently collected at both middleshelf sites and at the outer-shelf station. As indicated by the inverse dendrogram, species in group H, which were highly constant at station OSO I, were least similar to other groups in terms of their distribution. Several species, such as the bryozoans Aplousina gigantea and Cycloperiella rubra, the octocoralEllisella sp., and the decapod crustacean Mesopenaeus tropicalis, were collected only at station OS01. No species assemblage identified from our study was restricted to specific hard-bottom sites. However, comparison of species assemblages from cluster analysis of winter and summer from the first-year study (Wenner et al. 1983) with seasonal data from the present study show that the following species formed a consistently recurring assemblage on the inner shelf: The bryozoans Scrupocellaria regularis and Hippoporina contracta; the echinoderm Ocnus pygmaeus; and the crustaceans Synalpheus longicarpus, S. minus, Pilumnus pannosus, Megalobrachium soriatum, Trachypenaeus constrictus, and Conopea galeata.Other species, such as the cnidarian Sertularella areyi; the echinoderm Astroporpa annulata; the sponges Xytopsues griseus and Sigmadocia caerula; and the decapod crestaceans Mesopenaeus tropicalis, Sicyonia brevirostris, and Solenocera atlantidis, were consistently collected during both years at outer-shelf sites. These findings suggest that there is no sharp bathymetric zonation of epifaunal species groups in hard-bottom habitats of the South 6 = Table 3.-Specles groups resulting Crom numerical classification of data from samples coUected by trawl during 1981. (Ar Artbropoda; Bry data; Cn Cnldarla; Ech Echinodermata; Mo MoUusca; Po Porifera.) = Group A Anthosigmella varians (Po) Asterias forbesii (Ech) Cinachyra alloclada (Po) Pandaros acanthijoLium (Po) Leuconia aspera (Po) Crassimarginatella Leucocypha (Bry) Trypostega venusta (Bry) Microporella umbracuLa (Bry) Homaxinella waLlOnsmithi (Po) Higginsia strigiLara (Po) Lytechinus variegatus (Ech) CiocaLapota gibbsi (Po) CribriLaria radiata (Bry) Amphimedon compressa (Po) Reptadeonella hastingsae (Bry) Microciona proLijera (Po) Myriastra fibrosa (Po) Group B Scrupocellaria reguLaris (Bry) Gymnangium sinuosum (Cn) Hemectyon pearsei (Po) Spirastrella coccinea (Po) Ascidiacea A (Ch) CLiona caribbaea (Po) CLathrina coriacea (Po) Trachypenaeus constrictus (Ar) NoLella stipata (Bry) Synalpheus Longicarpus (Ar) Synalpheus minus (Ar) PiLumnus dasypodus (Ar) Celleporaria mordax (Bry) StyeLa panita (Ch) PiLumnus pannosus (Ar) Conopea gaLeara (Ar) Ocnus pygmaeus (Ech) Podochela riisei (Ar) Macrocoeloma trispinosum (Ar) Botryllus planus (Ch) MegaLobrachium soriatum (Ar) Scypha barbadensis (Po) Sundanella sibogae (Bry) Hippoporina conlracra (Bry) SympLegma viride (Ch) BOlrylloides nigrum (Ch) Membranipora lenuis (Bry) = = = Tiranideum frauenfeLdii (Cn) Obelia dichoroma (Cn) TeleSIO frU/iculosa (Cn) Didemnum candidum (Ch) Scandia murabiLis (Cn) CLytia cyLindrica (Cn) Dromidia antiLLensis (Ar) Group C StyeLa pLicala (Ch) Milhrax pLeuracanthus (Ar) MolguLa occidentaLis (Ch) Pilumnus sayi (Ar) Leprogorgia virguLora (Cn) Spheciospongia vesparium (Po) Sertularella conica (Cn) Crisia sp. (Bry) Synalpheus lownsendi (Ar) DislapLia bermudensis (Ch) Diodora cayenensis (Mo) DipLosoma macdonaLdi (Ch) Lophogorgia hebes (Cn) Arbacia punctuLara (Ech) Antropora lin cIa (Bry) HaLiclona ocuLara (Po) SerluLaria dislans (Cn) Cinachyra kuekenthaLi (Po) Arca imbricala (Mo) Group E Ircinia campana (Po) Pisanlia tincra (Mo) Ircinia strobiLina (Po) Aplysina fistuLaris (Po) Lissodendoryx isodictyaLis (Po) Homaxinella sp. A (Po) Chama congregara (Mo) Ircinia ramosa (Po) Ircinia feLix (Po) Group F Group D Sleganoporella magniLabris (Bry) Neofibularia nolirangere (Po) Smittina smittiella (Bry) StrongyLophora sp. B (Po) Caberia boryi (Bry) Callyspongia fallax (Po) PiLumnus floridanus (Ar) Geodia gibberosa (Po) Alpheus formosus (Ar) Schizoporella errala (Bry) CLaveLina gigantea (Ch) CrepiduLa acuLeara (Mo) Salacia desmoides (Cn) Amalhia disrans (Bry) Eudendrium sp. (Cn) Parasmittina spathulara (Bry) Ophiothrix angulata (Ech) Thalysias juniperina (Po) Halecium dysymmelrum (Cn) HippaLiosina rostrigera (Bry) SertuLaria pLumuLijera (Cn) TeLeslO sanguinea (Cn) Balanus venuSlus (Ar) Dynamena cornicina (Cn) Dynamena quadridentata (Cn) Eudendrium cameum (Cn) Conopea merrilli (Ar) Aelea anguina (Bry) Turbicellepora dichotoma (Bry) Pleria coLymbus (M 0) Balanus trigon us (Ar) Schizoporella comura (Bry) Microporella ciLiala (Bry) HaLecium sp. (Cn) Sertularia marginata (Cn) Monosraechas quadridens (Cn) AgLaophenia Irifida (Cn) Group G Eudendrium tellellum (Cn) Bimeria humiLis (Cn) BougaillvilLia sp. (Cn) Hiatella arclica (Mo) Serlularella sp. A (Cn) CLytia fragiLis (Cn) Celleporaria magnifica (Bry) Ctenostomata (Bry) Amalhia altemala (Bry) Stenociollops furcata coe/ala (Ar) AgLaophenia eLolIgala (Cn) FiLellum serralum (Cn) Diaperoecia floridana (Bry) Pseudomedaeus agassizii (Ar) Dynamella daLmasi (Cn) PachycheLes rugimanus (Ar) Scyllarides lIodifer (Ar) 7 = Bryozoa; Ch = Chor- Parellisina curvirostris (Bry) Polyandrocarpa floridana (Ch) Thyroscyphus marginalus (Cn) Schizoporella floridana (Bry) Hebella scandens (Cn) Synrhecium tubitheca (Cn) Aglaophenia allmani (Cn) Metapenaeopsis goodei (Ar) Campanularia hincksii (Cn) ASlrophylOn municatum (Ech) Hebella venusta (Cn) PetraLieJla bisinuata (Bry) Halecium tenellum (Cn) Celleporaria aLbiroslris (Bry) PLumularia selacea (Cn) Hincksella cylindrica (Cn) Lissodendoryx sigmala (Po) AgLaophenia lalecarinata (Cn) BuguLa ryLandi (Bry) Chaperia sp. (Bry) Bugula fuLva (Bry) Aeverillia seligera (Bry) Celleporina hassalli (Bry) Muricea penduLa (Cn) BuguLa microoecia (Bry) Group H Ellisella sp. (Cn) ApLousina gigantea (Bry) Mesopenaeus lropicalis (Ar) Sertularella areyi (Cn) Stylalula sp. (Cn) Cladocarpus fLexiLis (Cn) Diodogorgia sp. (Cn) Halopteris clarkei (Cn) Cycloperiella rubra (Bry) StyLopoma informata (Bry) Antipatharian (Cn) Sicyonia brevirostris (Ar) Alcyonidium sp. (Bry) Haliclona sp. C (Po) Leprogorgia sp. (Cn) Turritopsis nU/ricula (Cn) Thesea sp. (Cn) Astroporpa annuLata (Ech) SoLenocera alLantidis (Ar) Nellia ten ella (Bry) Xylopsues {lriseus (Po) .Teichaxinella co,rugata (Po) Sigmadocia caeruLa (Po) Axillella bookhouti (Po) Halicholldria bowerbanki (Po) Balanus calidus (Ar) SIMILARITY 2 .....L_-O'.L,.----'_-_OL L--.L\6-l._-...J'iL ..·.....L-l.?--JL...J°I· STATIONS STATIONS 1:501 1:502 MS02 M506 0501 :1:501 :l:S02 MS02 MS06 OSOI ..----A. A ----B B c c o o E E (/) lL :::> o a:: (.!) (/) LU U F LU lL (/) L------G G L------------_H H I -1.6 I -1.2 I -0.8 I -0.4 SIMILARITY I ° I 0.4 CONSTANCY 3.0 2~ FIDELITY _LO.7VERY HIGH EJ]2.0,1 LOW . 2 4 VERY HIGH B .2.0.5 HIGH 0<0,1 9123 0< IVERY MODERATE VERY LOW ~ HIGH 21l0W LOW 2.2 MODERATE Figure 4.-Inverse classification hierarchies and nodal diagram showing constancy and ftdeUty of station-species group coincidence hased on seasonal trawl collections. Atlantic Bight. However, the least similarity in species composition apparently occurs between inner- and outer-shelf sites while middle-shelf sites represent a transition zone between the other strata. Biomass of Larger Macroinvertebrates Comparisons of total biomass between taxa showed that Porifera were dominant at all stations during each season, except at station MS06 in spring, when biomass of Ascidiacea was highest (Table 4). These results are similar to those reported by Wenner et al. (1983) who found that Porifera accounted for a large percentage of the total invertebrate biomass from both the dredge and trawl collections. In the present study, the seasonal contribution of Porifera to the total biomass of invertebrates collected by dredge and trawl was as follows: Winter (78%), spring (59%), summer (67%), and fall (76%). The only taxa other than sponges and tunicates which formed a significant component of sample biomass in dredge or trawl collections were anthozoans and mollusks. Analysis of variance of logarithmically transformed biomass values from replicate samples (OS06 excluded) indicated no significant difference in biomass between seasons for invertebrates collected by dredge or trawl (P > 0.05). Because many of the large macrofauna sampled by dredge and trawl, such as sponges, appear to persist throughout the year, major changes in biomass were not observed. Undoubtedly, our sampling methods were not refined enough to detect minor seasonal changes in biomass due to growth or mortality. Significant differences in biomass were noted between stations (P < 0.05) for dredge-collected invertebrates, with the greatest average biomass recorded at stations IS02 (x = 22.3 kg) and ISOI (x = 13.6 kg). The lowest biomass of invertebrates collected by dredge occurred at station MS06 (x = 4.9 kg). Low biomass at this station is probably related to the patchy occurrence of invertebrates. Television transects showed that epifaunal organisms were sparse over parts of this station, with large expanses of sand interspersed with patches of live bottom. In addition, some sampling bias may have resulted from limiting dredge tows at MS02 and MS06 to areas of low relief. Since epifaunal growth was most dense on moderate to high relief ledges and rocks, an underestimation of biomass may have resulted from avoiding these areas. Biomass values for trawl-collected invertebrates were greatest at an inner-shelf site, which is consistent with results from the initial study (Wenner et al. 1983). Among stations sampled by trawl (OS06 excluded), average biomass values were significantly 35.8 kg), MS02 (x 23.2 kg), greater (P < 0.001) at IS02 (x and ISOI (x = 10.6 kg). The lowest average biomass was recorded at station OSOl (x 3.2 kg). = = = SmaUer Macroinvertebrates Sampled by Suction and Grab A total of 845 taxa was collected during seasonal sampling with suction sampler and grab. A listing of all fauna collected, including their ranks by abundance and density at each station, is presented in Appendix 3. The phylum Annelida, which was represented by more species than any other group during the initial study (Wenner et al. 1983), was also the most diverse group in the present study, where it contained 285 species. Annelids also dominated collections in terms of numerical abundance, accounting for 72.4% of the total 60,739 invertebrate specimens taken by suction sampler and grab. Mollusks ranked second among major Table 4 -Percent contribution to epifaunal biomass of taxonomic groups In dredge (D) and trawl (T) collections. Percent of the total biomass for both gear types Is also shown. The mean (x), standard deviation (SD), and number of samples (II) for which biomass measurements were taken are shown. ISOI Seaso. MS06 MS02 IS02 OSOI OS06 D T Total D T Total D T Total D T Total D T Total D T 27 <I 38 6 <I 5 24 66 <I 4 I <I I 26 54 <1 14 3 <I 3 25 58 <1 9 18 <I 7 7 89 <1 7. <I <I 1 7 86 <1 3 2 <I 2 7 13 3 14 24 <:1 21 25 86 <I 2 <I <I I 9 85 <I 3 1 <I 2 9 45 2 3 36 I 6 7 85 <1 2 1 <1 <I II 84 <I 2 7<1 <1 II 30 2 37 15 2 14 <I 37 1 15 5 2 13 27 34 () 0 0 0 <1 <I 0 0 0 0 0 0 0 0 0 1.09 0.55 0.52 2 77.79 12.96 11.96 6 78.88 9.86 11.63 8 6.27 3.14 0.62 2 11.94 1.99 1.76 6 18.22 2.28 1.59 8 25 <1 5 35 <I 10 25 32 <I 3 10 7 37 31 I 4 13 9 7 35 <I 2 41 I 24 31 <I 65 4 6 <1 16 8 I 57 4 10 <1 17 10 I 0 5 65 17 12 1 0 7.98 3.99 4.75 2 48.10 8.02 5.71 6 56.08 7.01 5.48 8 3.12 0.27 1.56 2 23.38 3.99 3.89 6 26.50 3.55 3.31 8 0.16 0.08 0.04 2 Total Winter Porifera Hydroidea Anthozoa Mollusca Decapoda Echinodermata Ascidiacea Other Invertebrata Total bio'mass L (kg) x SD n 47.80 111.09 158.89 23.90 18.50 19.86 10.78 11.08 15.33 2 6 8 29.55 284.64 314.19 56.93 52.36 45.01 41.45 5 6 2.47 179.54 182.01 29.90 26.00 38.08 36.28 6 7 2 23 8 2 14 18 Not sampled Spring Porifera Hydroidea Anthozoa Mollusca Decapoda Echinodermata Ascidiacea Total biomass (kg) L x SD n 60 <I 9 7 <I 4 20 43 1 I I <1 1 52 51 1 5 4 <1 2 37 57 0 6 11 <1 6 20 65 <I 2 1 <I I 30 36.53 18.26 1.17 2 42.76 7.13 5.88 6 79.29 9.91 7.18 8 78.14 39.07 10.33 2 52 <I 5 6 <I 4 33 31 2 4 22 <I 4 36 36 1 5 18 <I 4 36 29 1 12 7 <1 6 44 11.45 5.73 4.86 2 37.74 6.30 3.10 6 49.19 6.15 3.22 8 6.22 3.11 3.35 2 20 38 8 <1 2 30 76 <1 2 1 <I 1 19 66 <I 9 2 <I 1 21 66 <1 4 6 <1 8 15 45 <I 5 1 <1 3 44 13.08 6.54 5.10 2 61.63 10.27 8.54 6 74.72 9.34 7.67 8 42.34 21.17 7.95 2 56.31 9.38 6.64 6 62 <1 4 4 <I 3 27 32 1 18 8 <I 2 38 70 <I 5 3 <1 4 17 75 <I 4 2 <I 4 15 II '0 191.34 269.48 31.90 33.68 15.76 14.27 6 8 14.29 108.13 122.42 18.02 15.30 714 5.50 12.24 11.69 2 6 8 " 0. e 1Ji " Z Summer Porifera Hydroidea Anthozoa Mollusca Decapoda Echinodermata Ascidiacea Total biomass (kg) L x SD n 73 <I 2 1 <I I 23 2 9 1 2 <1 3 82 74 2 10 <1 2 2 10 70 2 10 <1 2 2 14 84 <1 6 3 <I <1 6 77 <I 2 <1 2 13 6 77 <I 2 1 2 12 6 80 0 14 <1 <I 2 3 36 2 13 <1 6 29 13 46 2 13 <1 5 23 II 49 12 21 6 <I 11 I 5.12 2.56 3.30 2 88.74 14.79 19.73 6 93.87 11.73 17.66 8 2.90 1.45 1.17 2 42.71 7.12 10.41 6 45.61 5.70 9.20 8 4.62 2.31 2.91 2 14.90 2.48 1.73 6 19.52 2.44 1.83 8 1.54 54 <1 5 4 <I 5 32 68 3 1 6 <I 4 18 88 2 3 <I 1 2 4 86 2 3 I I 2 5 95 <1 <1 2 0 I 1 93 <1 1 1 <1 2 3 93 <1 40 <1 10 35 4 2 9 54 <I 9 I 20 15 0 50 I 9 12 14 11 3 36 3 25 18 5 11 2 98.64 12.33 8.38 8 16.53 8.26 4.04 2 12.42 6.21 1.59 2 25.47 4.24 4.23 6 37.89 4.74 3.74 8 0.99 0.50 0.37 2 72 1 2 1 <I I 23 288.89 295.09 48.15 36.89 33.45 35.15 6 8 '0 " 0. e 1Ji " Z 1.54 Fall Porifera Hydroidea Anthozoa Mollusca Decapoda Echinodermata Ascidiacea Total biomass (kg) L x SD n 1 157.30 173.83 26.22 21.73 34.04 29.98 6 8 9 27.18 13.59 14.17 2 I 1 <1 2 3 90.30 117.48 15.05 14.68 16.09 14.64 6 8 "2 0. e 1Ji 0 Z 650), Chone americana (n = 543), Malacoceros glutaeus (n = 535), and Exogone dispar (n = 482); and the amphipods Luconacia incerta (n = 907), Photis sp. (n = 631), and Erichthonius brasiliensis (n = 523). These species accounted for about 62% of all invertebrate specimens caught in suction and grab samples. Densities of the 10 numerically dominant species changed seasonally and bathymetrically. In winter, the amphipods Luconacia incerta, Photis sp., and Erichthonius brasiliensis were important at inner-shelf sites (Table 5). Their densities at these sites decreased in spring; and during the rest of the year <5 individuals/0.1 m 2 were collected. The density of F. implexa was high in fall, when approximately 182 individuals/O.I m2 were collected at IS01. Another species whose density was highest in fall on the inner shelf was the polychaete Syllis spongicola. However, taxonomic groups with a total of 251 species; however, mollusks accounted for only 4.3% of the total number of invertebrate specimens and were outranked by amphipods (100 species) which represented 13.3% of total invertebrate abundance. Other groups represented in suction and grab collections included decapod crustaceans (111 species), arthropods other than amphipods and decapods (49 species), echinoderms (39 species), nemerteans (3 species), and sipunculans (7 species). Many of the taxa collected by suction and grab represent sand-bottom species which are also common at the hard-bottom sites, since there is generally a sand layer covering the rock bottom. The 10 invertebrate species which numerically outranked others at the hard-bottom sites during all seasons were the polychaetes Filograna implexa (n = 30,346), Pista palmata (n 1,439), Syllis spongicola (n = 1,358), Spiophanes bombyx (n = Table 5.-Average density (x) expressed as number per 0.1 m', and percent of total number (%) for 10 numericaUy dominant invertebrates collected by suction and grab. Number of coUections (n) is indicated. ISOI (n 5) = Species x % IS02 (n 5) MS02 (n 5) = x % 3.2 0.2 6.6 22.6 4.6 24.8 4.4 0.4 4 \.0 16.8 <1 <1 \.6 5.6 \.2 6.2 1.1 <1 10.2 4.2 0.4 0.8 \.4 68.0 0.4 20.4 <1 <1 <1 15.8 <I 4.7 0.2 27.0 6.6 <1 6.3 \.5 = x % x ------------------------------------------------------Filograna implexa Pista palmata Syllis spongicola Luconacia incena Spiophanes bombyx POOtis sp. Chane americana Malacoceros glutaeus Erichthonius brasiliensis Exogone dispar MS06 (n 5) = 53.2 \.4 35.4 17.8 3.0 3 \.8 7.8 19.0 2.0 18.0 4.8 4.6 0.4 0.8 \.8 0.2 3.4 4.6 2.4 2.2 <1 <1 <1 <1 \.7 2.2 0.2 0.4 \.2 2.0 0.2 10.8 0.4 0.4 16.8 6.0 <1 <1 <1 <1 <1 3.7 <I <1 5.7 2.0 8.9 <1 5.9 2.9 <1 5.3 1.3 3.2 <1 3.0 1.084.4 66.2 \.8 <1 \.0 17.2 7.0 <1 2.4 <1 4.2 <1 4 \.2 2.5 11.6 <1 \.6 <1 8.4 <1 -----------------------------------------------------Filograna implexa Pista pa/mata Syllis spongicola Luconacia incena Spiophanes bombyx Photis sp. Chone americana MaLacoceros gLutaeus Erichthonius brasiliensis Exogone dispar 0.2 14.8 0.8 0.2 0.8 \.4 2.8 4.0 \.6 <1 10.4 <1 <1 <1 \.0 \.9 2.8 1.1 3.6 0.2 0.2 1.8 \.2 2.2 <1 <1 1.1 <1 = x % OS06 (n 5) = x % Winler ---- --- ---- --------- -- ----- ------ ----- ---- ------ ---- --0.6 2.2 16.0 45.0 9.4 2.0 5.2 6.2 0.2 7.2 0.2 0.2 3.4 22.2 2A 3.8 2.0 0.6 0.6 Summer 0.2 1.0 7.4 0.8 0.6 <1 <1 3.7 <1 <1 \.0 2.4 0.2 0.8 <1 \.2 <1 <1 <1 <1 4.2 I\.9 2.5 <I \.4 \.6 <1 \.9 20.33 20.4 17.66 3.66 0.33 17.7 3.7 <1 ----------------------------------------------------_.. ----------------------------------------------------.-- Spring Filograna implexa Pista palmata Syllis spongicola Luconacia incena Spiophanes bombyx Photis sp. Chane americana Malacoceros glutaeus Erichrhonius brasiliensis Exogone dispar % OSOI (n 3) 248.4 7.6 43.3 1.3 48.8 15.8 0.6 8.5 2.7 <1 34.6 2.0 4.4 \.0 4.4 0.6 2.8 <1 0.4 <1 <1 \.8 I\.6 1.3 2.0 1.0 <1 <1 \.4 14.2 <1 \.8 <1 \.8 <I \.1 -----------------------------------------------------1.6 0.8 6.6 0.4 \.6 0.2 24.6 8.8 0.4 2.0 <1 <I 2.8 <1 <1 <1 10.4 3.7 <1 <I 14.0 8.0 10.1 5.7 11.6 8.4 0.2 0.2 0.4 <1 <1 <1 1.0 0.2 2.0 <1 <1 1.5 3.0 4.6 2.2 3.4 0.2 0.2 <1 <1 ----- ---- --- ---- --- ------ --- ---- -- --- --- ---- ----- -------- Fall ----- --- ---- ---- ---- ---- ---- ---- ---- ---- ----- ---- ---- ---- Filograna imp/exa Pista palmata Syllis spongicola Luconacia incena Spiophanes bombyx Photis sp. Chane americana Malacoceros glutaeus Erichthonius brasiliensis Exogone dispar 182.4 0.4 55.0 3.6 41.4 <I 12.5 <1 \.6 1.8 5.4 \.0 14.2 <I <1 1.2 <1 3.2 2.4 0.2 1.6 0.4 0.8 0.4 2.2 <1 1.5 <1 <1 <1 7.0 0.4 2.8 6.4 <1 2.6 53\.6 55.4 0.4 44.6 4.6 3.0 1.8 4.4 <1 3.4 <1 36.8 3.8 3.2 <1 3.8 <I 10 4.155.0 \.2 20.2 0.8 \.6 2.4 \.8 0.4 1.6 93.0 <1 <I <1 <1 <1 <1 <1 <1 24.6 <1 4.1 10.0 13.4 18.0 0.2 3.0 5.0 6.7 0.2 <1 0.4 <1 0.6 <I tion, especially by fishes; and 3) differing microhabitats within and between sites. Predation could influence the communities in this study since it has been found to be a major factor affecting benthic populations in soft- and hard-bottom communities of the intertidal zone (Paine 1966; Peterson 1977). However, studies on the effects of predation on the continental shelf are limited and largely conjectural. Frankenberg and Leiper (1977) hypothesized that fishes remove large numbers of small crustaceans, such as cumaceans and amphipods, from benthic assemblages. Predation on benthos can also be expected to alter species composition, resulting in greater abundance of those species having higher intrinsic rates of population increase (Slobodkin 1961); and, furthermore, a selective preference of predators for soft forms such as crustaceans or polychaetes may shift species composition toward forms with exoskeletons which are more resistant to predation (e.g., mollusks) (Richards 1963). Examination of stomach contents of 10 species of fishes known to inhabit live-bottom areas showed that amphipods, polychaetes, and decapods were most heavily preyed upon, including the numerically dominant species Exogone dispar and Erichthonius brasiliensis (G. Sedberry). neither of these species was consistently abundant at both innershelf sites. Filograna implexa was the dominant species at middle-shelf sites (Table 6). The maximum density of F. implexa occurred in fall at MS06 when 4,155 individuals/O.l m 2 were collected (Table 5). The high densities of F. implexa which were observed at midshelf sites did not occur on the outer shelf (Table 6). Species which were important at these deeper sites were the polychaetes Pista palmata and Spiophanes bombyx, with the highest densities for both occurring in spring. Comparisons with results presented by Wenner et aI. (1983) confirm that F. implexa is the numerically dominant invertebrate at middle-shelf sites. However, its importance at outer-shelf sites was considerably diminished during the second year of sampling when numbers of Pista palmata exceeded those of other species collected. Filograna implexa was also important on the inner shelf, but densities did not approach those noted in previous collections from these sites. Other investigators (Frankenberg 1971; Frankenberg and Leiper 1977; Tenore 1978) have noted that density variation over space and time is typical of numerically dominant species in softbottom communities on the continental shelf off Georgia. Wenner et al. (1983) speculated that variable abundances of benthic invertebrates could be influenced by 1) differentially successful and sequential recruitment by larvae of various species; 2) preda- 'G. Sedberry. Assistant Marine Scientist, Marine Resources Research Institute, Box 12SS9, Charleston, SC 29412, pers. commun. January 1983. Table 6.-The 10 most abundot Invertebrate species coUected by suction od grab samplers at eacb station. (An Ecblnodermata; 81 = 8IpuncuUda.) Species Total number Percent of total Species ISOI Filograna implexa (An) Syllis spongicoia (An) Erichthonius brasiliensis (Ar) Axiothella mucosa (An) Exogone dispor (An) Luconacia incena (Ar) Amphorete americana (An) PhOtis "p. (Ar) Elasmopus "p. A (Ar) Gammaropsis "p. (Ar) IS8 143 140 130 lOS IS.6 6.8 4.2 4.1 3.1 2.7 2.4 2.4 2.2 1.8 Luconacia incena (Ar) Elasmopus sp. A (At) Erichthonius brasiliensis (At) Lembos smithi (Ar) CapreIIa equilibra (At) Podocerus "p. (Ar) Photis sp. (At) Leptochelia "p. (Ar) Lumbrineris inflata (An) Polycirrus carolinensis (An) 49.2 3.1 2.1 1.6 1.2 1.2 1.2 l.l 1.0 0.9 Filograna implexa (An) Luconacia incena (Ar) Syllis spongicoia (An) Chone americana (An) Spiophones bombyx (An) Syllis hyalina (An) Aspidosiphon gosnoldi (Si) Polydora colonia (An) AutoIytus "p. (An) Malacoceros glutaeus (An) 32.4 8.9 4.0 3.0 2.7 2.1 2.0 1.6 1.4 1.4 Filograna implexa (An) Plakosyllis quadrioculata (An) Rildardanus Iaminosa (Ar) Protomedeia "p. A (Ar) Amphorete americana (An) Syllis spongicoia (An) Sphaerosyllis sp. (An) Syllidae L (An) Chone americana (An) Syllis hyalina (An) MS02 Percent of total 3S3 22S 7.1 221 179 16S 160 4.4 3.6 3.3 3.2 3.2 2.6 2.2 1.7 IS8 127 109 84 4.S MS06 8,347 S23 349 267 203 202 19S 192 16S ISS OSOI Pista polmata (An) Spiophones bombyx (An) Erichthonius "p. A (Ar) Syllis spongicoia (An) Onuphis polliduia (An) Photis "p. (Ar) Owenia fusiformis (An) Photis Iongicaudata (Ar) Mesochaetopterus "p. (An) Amphorete acutifrons (An) Total number IS02 929 406 247 246 186 Filograna implexa (An) Syllis spongicoia (An) Malacoceros glutaeus (An) Chone americana (An) Ophiothrix angulata (Ech) Photis "p. (Ar) Acanthohoustorius millsi (Ar) Megalobrachium soriatum (Ar) Lumbrineris inflata (An) Exogone dispor (An) = AnneUda; Ar = Artbropoda; Ecb = 20,787 248 21S 180 174 173 120 107 96 94 78.8 0.9 0.8 0.7 0.7 0.7 O.S 0.4 0.4 0.4 OS06 1.373 380 170 128 114 90 84 67 S9 S7 11 268 112 104 71 S4 S3 47 47 40 40 11.9 S.O 4.6 3.2 2.4 2.4 2.1 2.1 1.8 1.8 Much of the variability in populations of infaunal organisms can be attributed to our sampling methods which included too few replicates to accurately determine whether changes in density were due only to seasonality. In addition, the complex' physiography and diverse substrates of hard-bottom habitats create a habitat patchiness that could result in either selective settling by larvae or differential survival of different benthic invertebrate species. Other physical factors which have been reported to affect stability (i.e., persistence) of benthic populations are fluctuations in water temperature (Boesch et a1. 1977) and disturbance of sediments by storms (McCall 1977). Boesch et a1. (1977) attributed the temporal stability of macrofaunal species on the outer shelf of the Middle Atlantic Bight to decreasing thermal variation and reduced effects of storms in deeper water. Although upwelling of colder water does occur on the continental shelf in the South Atlantic Bight, most of the benthic infaunal species collected by suction sampler and grab have wide latitudinal ranges and are eurybathic, extending from the intertidal zone to the continental slope and beyond. Thus, most should be fairly tolerant of the environmental conditions found at hard-bottom sites on the shelf. Values of Shannon diversity (II'), evenness (J), richness (SR), dominance (01), number of species (s), and number of individuals (n) showed no consistent patterns between depth zones and seasons (Table 7). During all seasons, It was> 5 bits/individual at both inner-shelf stations, except ISOI in fall when It was 4.58. This comparatively low It value reflected an uneven distribution of individuals among species at ISOI and, consequently, an elevated dominance index (01) value rather than low SR or s. Low evenness and high 01 were attributed to large numbers of the colonial polychaete Filograna implexa. More species were collected at mid-shelf stations than at any others sampled. Approximately 42 and 48% of all species collected at MS02 and MS06, respectively, were represented by one or two individuals and are consequently considered rare. Both s and SR were greatest at mid-shelf stations in winter; however, the community-structure values were not seasonally consistent between stations. For example, values of It were high (> 6) at MS06 during all seasons·except fall when the lowest It value for any station sampled occurred. At MS02, high It values were noted only in winter and summer. As noted for inner-shelf stations, low It values were affected by low evenness which reflected strong dominance by the polychaete F. implexa. Community-structure values at outer-shelf stations were also not consistent between stations or seasons. At station OS06, It and J showed little seasonal variation and indicated that species were fairly diverse and evenly distributed. The dominance index dominated by also indicated that collections were not num~ricaly one or two species. However, sand SR at OS06 did change with season, being much lower in fall. At station OSO I, seasonal changes in It reflected concomitant changes in evenness and dominance. Thus, although more species and greater SR occurred at OSOI in spring, It was lower than in any other season at this station due to numerical dominance by the polychaete Pista palmata. Most community-structure values at outer-shelf sites were comparable with those obtained on the inner shelf; however, the fewest species obtained on the shelf were recorded at OSO 1 in winter and OS06 in fall. In addition, fewest individuals were collected at outer-shelf stations. In general, results from the present study are consistent with those reported in the initial study of hard-bottom habitats (Wenner et a1. 1983). Sampling during both years has shown that rare species are a conspicuous component of the hard-bottom community and contribute to elevated species richness at all of the sites sampled. In addition, no cons.istent seasonal changes in diversity or s (Kruskal-Wallis test, P > 0.05) were noted for stations or depth zones. The low It values which occurred at hardbottom sites sampled in both years of study apparently reflect localized changes in the population density of a few dominant species, such as F. implexa, and are probably not indicative of any seasonality. Although II' and its components did not show any definite patterns with regard to depth or latitude, the number of species was found to be significantly different between sites (Kruskal-Wallis test, P < 0.01). Specifically, more species were collected at midshelf sites, which may be related to more stable bottom temperatures in this area of the shelf (Miller and Richards 1980). In terms of community structure, these sites were characterized by I) a large number of species, many of which were rare, and 2) high numerical dominance by Filograna implexa. In contrast, innershelf sites, which were subject to more extreme temperature variations, generally had fewer species and less dominance than sites on the middle shelf. Outer-shelf sites are also thermally variable because of intrusions of colder upwelled water, which predominantly occurs in fan (Atkinson et a1. 1978). Fewest species were recorded at outer-shelf sites, and many of those which occurred were rare. In addition, dominance was relatively low. These results suggest that fluctuating temperatures at innerand outer-shelf sites may exert some control on community structure at live-bottom sites. However, it is also possible that the lower richness at outer-shelf sites was due to collection of samples with a grab which does not sample hard substrates as effectively as the suction device. Although within-reef habitat heterogeneity is one important factor in determining richness of benthic fauna, thermal stability is probably also important. Normal cluster analysis of pooled replicate collections taken with suction and grab samplers grouped stations according to depth zones with inner-shelf stations forming group I, middleshelf stations forming group 2, and two outer-shelf sites forming Table 7.-Seasonal community structure values [num"".. or individuals, number or species, diversity (H'), evenness (J'), rlcbness (SR), and dominance Index (DI)) ror pooled repUcate samples or Invertebrates at eacb station. Station Season IS01 winter spring summer IS02 ran winter spring summer MS02 ran winter spring summer MS06 ran winter spring summer OSOI ran winter spring summer OS06 ran winter spring summer rail No. individuals No. species It .r SR D1 2.003 1.020 713 2.204 2.158 1.468 808 547 2,987 8,190 990 4,799 1.896 954 1.188 22,328 299 2,869 694 373 245 178 172 222 176 197 172 148 313 306 261 307 319 230 251 297 77 255 135 125 6.09 5.90 6.17 4.58 5.58 6.41 6.42 6.25 6.55 3.22 7.19 3.99 6.75 6.61 6.71 0.85 4.88 4.48 5.78 6.10 0.77 0.79 0.83 0.59 0.75 0.84 0.87 0.87 0.79 0.39 0.90 0.48 0.81 0.84 0.84 0.10 0.78 0.56 0.82 0.88 32.10 25.55 26.03 28.71 22.80 26.88 25.54 23.30 39.00 33.85 37.70 36.10 42.13 33.38 35.31 29.46 13.33 31.90 20.48 20.94 17.2 24.2 20.5 53.8 22.6 11.2 12.9 15.2 14.8 68.7 8.5 60.0 17.5 17.4 14.6 93.5 38.1 51.8 18.4 20.1 1.215 670 366 198 189 99 6.03 6.69 5.27 0.79 0.88 0.80 27.74 28.89 16.60 21.3 7.5 32.2 12 species were limited to specific bathymetric zones during both years of study (Wenner et al. 1983). No species was found to be highly constant at inner-shelf sites, whereas the polychaete Eunice jilamentose, the amphipod Megaluropus sp., and the mollusk Chione grus were most consistently collected at middle-shelf sites during both years. In addition, the amphipod Erichthonius sp. A, the polychaete Syllis sp. D, and the echinoderm Genocidaris maculata were characteristically found at sites sampled on the outer shelf in both studies. These results indicate that ~any of the smaller epifaunal and infaunal species comprising benthic assemblages at hard-bottom sites are relatively ubiquitous across the shelf, although differences in overall community composition were observed between depth zones. groups 3 and 4 (Fig. S). The two outer-shelf station groups constituted a larger agglomeration in the dendrogram which was very dissimilar in species composition to the inner- and middle-shelf groups. None of the stations ,were grouped together by season; however, inner- and outer-shelf sites were grouped according to latitude with ISOI and OSOI clustering separately from IS02 and OS06, respectively. Major agglomerations in the dendrogram indicated bathymetric separation of sites, suggesting that depth is a major factor influencing species distribution. During the previous study, quantitative collections of smaller epibenthic and infaunal organisms were separated according to bathymetric zones during winter but not during summer, suggesting that the quantitative composition of the hard-bottom community was much more homogeneous across the shelf during summer (Wenner et al. 1983). However, based on the present study, we have found no evidence to suggest consistent seasonal change in community composition. Similarly, seasonal differences in species assemblages were not observed by Boesch et al. (1977) in a study of benthic communities in the Middle Atlantic Bight. Most species groups delineated by cluster analysis were not limited in their distribution to one depth zone (Table 8, Fig. 6). Species in group A were consistently collected at inner- and middle-shelf sites, whereas those in group B were highly constant at all sites. Group C species were .also ubiquitous, but displayed high constancy for middle-shelf stations. The only species in this group which was collected solely at middle-shelf sites was the polychaete Ceratonereis sp. A. Species in group D, which were widely distributed across the shelf, displayed highest constancy at middle-shelf stations and OSOL Groups E, F, and G were most consistently collected at inner- and middle-shelf stations but were not restricted in their distribution to these sites. The tanaidacean Apseudes sp. A, which occurred in group G, was the only species which was restricted to inner-shelf sites. Species in group H were found at all stations but were most frequently encountered at middle-shelf stations and OS06. Examination of inverse cluster groups and nodal diagrams from the initial survey of hard-bottom habitats also revealed that few SUCTION Group 2 3 4 SlolOn Station Sa F. S. Wi Wi Sp Sa F. 1501 -Wi -"502 - Sp F. Sa F. Wi Sa Sp - SPSa F. Wi -SPSa F. ISOI ISOI ISOI IS02 IS02 IS02 IS02 AND 0 •• I GRAB: STATION CONCLUSIONS I) A total of 1,307 invertebrate taxa was collected with all sampling devices documenting the diverse nature of offshore hardbottom habitats. Most sites were inhabited by rare species which were represented by only one or two individuals. Numerical dominance was limited to the polychaetes Filograna implexa and Pista palmata which were prevalent at middle- and outer-shelf sites, respectively. 2) Diversity of hard-bottom sites was generally high during all seasons. No consistent seasonal changes in diversity were noted for stations or depth zones throughout the study area. Low If values were attributable to low evenness caused by high numbers of species such as Filograna implexa and Pista palmata. Although If and its components did not show any well defined changes with depth, season, or latitude, more species were collected at middleshelf sites. This may be related to less variable bottom temperatures in this area of the shelf, coupled with relatively high habitat complexity due to the frequent occurrence of large sponges and octocorals, which offer refuge to other benthic invertebrates. The number of identifiable invertebrate taxa collected at each station by dredge and trawl did not change significantly between seasons. Changes in the number of species between stations were attributable to varying degrees of topographic complexity and habitat heterogeneity. 3) Porifera, Bryozoa, and Cnidaria were the most diverse taxonomic groups collected by dredge and trawl during both years in which sampling was conducted. Sponges were most diverse at sites on the inner shelf, but their diversity diminished in dredge and trawl collections at middle- and outer-shelf sites. Taxa accounting for the greatest number of species in the latter areas were cnidarians and bryozoans. 4) Comparisons of total biomass between taxa showed that Porifera were dominant at most stations. No significant difference in biomass estimates was observed between seasons; however, bathymetric differences did occur, with biomass being greatest on the inner shelf for dredge and trawl collections. This may reflect the more uniform distribution of epifaunal invertebrates at innershelf sites; however, differences between sites may also be caused by sampling bias since efforts were made not to sample over high relief ledges on the middle shelf in order to minimize loss of gear. S) The hard-bottom sites sampled throughout the entire study area were distinguished from each other primarily by trends in faunal composition related to depth. No evidence of seasonal grouping was noted. Epifaunal species censused by dredge and trawl appeared to be more faithful to particular depth zones than did benthic organisms sampled by suction sampler and grab. However, most species assemblages were not restricted to a particular depth zone. Inner- and middle-shelf sites were more GROUPS SIMILARITY 0.7 ! 0.5 I 0.. I 0.1 ! -0.1 I -0.' I - MS02 lIS02 llS02 IIS06 11$06 llS06 MS06 -osOi 0501 OSOI OSOI -0sii6 0506 OS06 Figure 5.-Normal cluster dendrogram of station groups formed using the Bray-Curtis slmUarity coemclent, log transformation, and flexible sorting. Data from repUcated suction and grab collections were pooled for eacb station prior to cluster analysis. 13 Table 8.-Species groups resulting from numerical classlncatlon of data from samples collected by suction and grab samplers during all seasons, 1981. (Am = Amphlpoda; Br = Branchlopoda; Cu = Cumacea; D = Decapoda; E = Echinodermata; I = Isopoda; M = Mollusca; My = Mysldacea; P = Polychaeta; Po = Porifera; Py = Pycnogonlda; SI = Sipuncullda; T = Tanaidacea.) Group A Ophiothrix angulata (E) Polycirrus carolinensis (P) Lumbrineris inflata (P) Exogone dispar (P) Ophiactis savignyi (E) Luconacia incena (Am) Podocerus sp. (Am) Megalobrachium soriatum (D) Paracerce;s caudata (I) Ampithoe sp. A (Am) Elasmopus sp. A (Am) Carpias bermudensis (I) Lembos smithi (Am) Websterinereis tridentata (P) Podarke obscura (P) Synalpheus IOwnsendi (D) Xanlhidae A (D) Melita appendiculata (Am) Sipunculida A (Si) Pherusa inflata (P) Maera sp. A (Am) Pista quadrilobata (P) Arca imbricata (M) Ampelisca schellenbergi (Am) Chama congregata (M) Gouldia cerina (M) Pseudomedaeus agassizii (D) Lysianopsis alba (Am) Crepidula aculeata (M) Syllis gracilis (P) Group B Aspidosiphon gosnoldi (Si) Syllis hyalina (P) Chone americana (P) Eunice vittata (P) Leucothae spinicarpa (Am) Syllis spongicola (P) Pagurus carolinensis (D) Laevicardium pictum (M) Malacoceros glutaeus (P) Megalomma bioculatum (P) Chrysopelalidae A (P) Amphiodia pulchella (E) Eulalia sanguinea (P) Crassinella lunulata (M) Axiothella mucosa (P) Spiophanes bombyx (P) Owenia fusiformis (P) Pista palmata (P) Filograna implexa (P) Harmothoe sp. A (P) Caulleriella ki/lariensis (P) Ampharete americana (P) Prionospio cristata (P) Axiognathus squamatus (E) Loimia medusa (P) Spio pettiboneae (P) Polydora caeca (P) Prionospio cirrifera (P) MicrodeulOpus myersi (Am) Armandia maculata (P) Erichthonius brasiliensis (Am) Photis sp. (Am) Gammaropsis sp. (Am) Caprella equilibra (Am) Leptochelia sp. (T) Anoplodactylus petiolatus (Py) Diplodonta punctata (M) Mitrella lunata (M) Suturoglypta iontha (M) Pherusa ehlersi (P) Caprella penantis (Am) Varicorbula operculata (M) Syllis cornuta (P) Schistomeringos rudolphi (P) Brania clavata (P) Group D Leptochela papulata (D) Bowmaniella portoricensis (My) Ampelisca cristoides (Am) Paraprionospio pinnata (P) Mesochaetopterus sp. (P) Onuphis nebulosa (P) Batrachanotus fragosus (D) Ampharete aCUlifrons (P) Spionidae B (P) Unciola sp. A (Am) Onuphis pallidula (P) Ampelisca vadorum (Am) Aonides mayaguezensis (P) Automate evennanni (D) Xanthidae B (D) Glycera capitata (P) Nassarius albus (M) Pomaroceros american us (P) Group C Nassarina minor (M) Lysidice ninetta (P) Lumbrineris coccinea (P) Tellina sybaritica (M) Laonice cirrata (P) Nephtys squamosa (P) Pelecypoda B (M) Marginella lavalleeana (M) Eunice filamentosa (P) Melphidippidae A (Am) Diodora cayenensis (M) Musculus lateralis (M) CeralOnereis versipedata (P) Ceratonereis sp. A (P) lnachoides forceps (D) Latreutes parvulus (D) Trichobranchus glacialis (P) Marphysa sp. A (P) Phyllodoce fragilis (P) A Ipheus normanni (D) Trypanosyllis zebra (P) Sicyonia laevigata (D) Branchiosyllis exilis (P) Websterinereis sp. B (P) Chevalia sp. (Am) Chione grus (M) Polycirrus eximius (P) Hiatella arctica (M) Tellina americana (M) Diastylidae A (Cu) Semele nuculoides (M) AcanthohauslOrius millsi (Am) Megaluropus sp. (Am) Syllis regulata carolinae (P) Botula fusca (M) Pitha Iherminieri (D) Pagurus hendersoni (D) CeralOnereis mirabilis (P) Ophiostigma isacanthum (E) Majidae A (D) Arabella mutans (P) Anchialina typica (My) Pelia mUlica (D) Lithophaga bisulcata (M) Gastrochaena hians (M) Group G Phyllodoce castanea (P) Pseudovermilia occidentalis (P) Apseudes sp. A (T) Arbacia punctulata (E) Eunice antennata (P) Pilumnus sayi (D) Hyboscolex longiseta (P) Amphipoda E (Am) Paraphaxus spinosus (Am) Amphipoda B (Am) Cerithium atratum (M) Vermiliopsis annulata (P) Cumingia tellinoides (M) Pseudovermilia sp. A (P) Mithrax pleuracanthus (D) Tanyslylidae A (Py) Pagurisrestortugae (D) Group H Group E Erichsonella filiform is (I) Marginella hartleyanum (M) Lucifer faxoni (D) Pseudeurythoe ambigua (P) Mangelia rugirima (M) Corbula contracta (M) Aspidosiphon albus (Si) Chione latilirata (M) Phyllodoce longipes (P) Glycera sp. B (P) Marginella roscida (M) PhyliochaelOpterus socialis (P) Subodyte pellucida (P) Group F Hydroides crucigera (P) Hydroides sp. A (P) 14 Melitidae A (Am) Syllidae L (P) Syllis sp. D (P) Genocidaris maculata (E) Glycera tesselata (P) Odontosyllis enopla (P) Syllis alternata (P) Syllis ferrugina (P) Rildardanus laminosa (Am) Erichthonius sp. A (Am) Glycera papillosa (P) Protadorvillea kefersteini (P) Lembos unicornis (Am) Goniadides carolinae (P) Thyone pseudofusus (E) SIMILARITY -1.0 ! -0.8 J I ! - Q6 1 -O~4 ! ! ! -0.2 ! STATIONS I en , 0.2 STATIONS ISOI IS02 MS02 MS06 0501 0506 ISOI A A B B IS02 MS02 MS06 -==.,='-1 ll. :;) o a:: Cl L--------c en LLJ U LLJ ll. ,----------0 en E ,-----G '------H [ I t I I I I -1,0 -0.8 -0.6 -0..4 I I 1 -0,2 I 0.2 CONSTANCY SIMILARITY FIDELITY _1.0.7 VERY HIGH 01.0.1 LOW _1.4VERY HIGH 01. I LOW II1II1.0.5 HIGH D 1IIilIIiI1.3HIGH 3.0 1~ MODERATE <0.1 VERY LOW 2~ MODERATE D < I VERY LOW Figure 6.-Inverse classification hierarchies and nodal diagram showing constancy and fldellty of station-species group coincidence based on seasonal suction and grab collections. similar in species composition to each other than they were to outer-shelf sites. 6) The densities of some numerically dominant species collected by suction and grab changed seasonally and bathymetrically. Such density variations may result from: I) Differentially successful and sequential recruitment among larvae of various species; 2) predation, especially by marine fishes; and 3) habitat complexity. Seasonal influences on densities of individual species were not large enough to cause detectable seasonal changes in overall community structure and composition. water pollution. U.S. Environmental Protection Agency Office of Research and Development, Report No. EPA-600/3-77-033, 113 p. BOESCH, D. F., J. N. KRAEUTER, and D. K. SERAFY. 1977. Distribution and structure of communities of macrobenthos on the outer continental shelf of the middle Atlantic Bight: 1973-1976 investigations. Va. Inst. Mar. Sci., Spec. Rep. Appl. Mar. Sci. Ocean Eng. 173, 111 p. BRAY, J. R., and J. T. CURTIS. 1937. An ordination of the upland forest communities of southern Wisconsin. Ecol. Monogr. 27:323-349. CHESS, J. R. 1979. An airlift sampling device for in situ collecting of biota from rocky substrata. Mar. Technol. Soc. J. 12(3):20-23. CLIFFORD, H. T., and W. STEPHENSON. 1973. An introduction to numerical classification. Acad. Press, N.Y., 229 p. CONTINENTAL SHELF ASSOCIATES. 1979. South Atlantic hard bottom study. Prepared for Bureau of Land Management (Contract AA331-CT8-23), Washington, D.C., 336 p. DEAN, T. A. 1981. Structural aspects of sessile invertebrates as organizing forces in an estuarine fouling community. J. Exp. Mar. BioI. Ecol. 33:163-180. DUKE UNIVERSITY MARINE LABORATORY. 1968. Manual for use of R/V Eastward. Duke University Cooperative Research and Training Program in Biological Oceanography, Beaufort, N.C., 146 p. FRANKENBERG, D. 1971. The dynamics of benthic communities off Georgia, USA. Thalassia Jugosl. 7:49-33. FRANKENBERG, D., and A. S. LEIPER. 1977. Seasonal cycles in benthic communities of the Georgia continental shelf. In B. C. Coull (editor), Ecology of marine benthos, p. 383-397. Univ. South Carolina Press, Columbia. LAMBERT, J. M., and W. T. WILLIAMS. 1962. Multivariate methods in plant ecology. IV. Nodal analysis. J. Ecol. 30:773-802. LANCE, G. N., and W. T. WILLIAMS. 1967. A general theory of classificatory sorting strategies. I. Hierarchical systems. Compu!. J. 9:373-380. MARGALEF, D. R. 1938. Information theory in ecology. Gen. Syst. 3:36-7 I. McCALL, P. L. 1977. Community patterns and adaptive strategies of the infaunal benthos of Long Island Sound. J. Mar. Res. 33:221-266. ACKNOWLEDGMENTS Weare grateful to all persons associated with the joint South Carolina-Georgia project to study hard-bottom habitats which was funded by the Minerals Management Service (Contract AA551-CTI-18). Those persons who helped with collection of samples and identification of invertebrate taxa included: B. B. Boothe, D. R. Calder, A. L. Edwards, J. V. Miglarese, F. L. Nicholson, C. B. O'Rourke, E. C. Roland, and S. Shipman. Additional technical help at sea and in the laboratory was provided by W. T. Hodges, Jr., M. S. Maclin, G. H. Steele, A. C. Vitucci, Jr., H. L. Ansley, C. F. Cowman, C. D. Harris, R. Mahood, and S. P. Olsson. We also thank M. J. Clise and G. Gash for help with data processing and coordination; L. H. Hodges and N. M. Beaumont for secretarial support; K. Swanson for graphics; and C. K. Biernbaum and G. R. Sedberry for critical review of the manuscript. LITERATURE CITED ATKINSON, L. P., R. T. BARBER, and M. WADE. 1978. Hydrography, nutrients, chlorophyll, and organic carbon. South Atlantic Benchmark Program. Vol. 3 Draft Report, p. 13-72. Bureau of Land Manqement, Outer Continental Shelf Office, New Orleans, La. BOESCH, D. F. 1977. Application of numerical classification in ecological investigations of 15 McNAUGHTON, S. J. 1967. Relationships among functional properties of California grasslandB. Nature (Lond.) 216:168-169. MENGE, B. A., and J. P. SUTHERLAND. 1976. Species divenity gradients: synthesis of the roles of pmlalion, competition, and temporal heterogeneity. Am. Nat. 110:351-369. MIlLER, G. C., and W. J. RICHARDS. 1980. Reef fish habitat, faunal assemblages, and facton determining distributions in the South Atlantic Bight. Proc. Gulf Caribb. Fish. Insl. 32: 114-130. OSMAN,R. W. 1977. The establishment and development of a marine epifaunal community. Ecol. Monogr. 47:37-63. PAINE, R. T. 1966. Food web complexity and species divenity. Am. Nat. 100:65-75. PETERSON, C. H. 1977. Competitive organization of the soft·bollom macrobenthic communities of southern California lagoons. Mar. BioI. (Bed.) 43:343-359. PIELOU, E. C. 1975. Ecological diveni.lY. John Wiley & Sons, N.Y., 165 p. RICHARDS, S. W. 1963. The demersal fish population of Long Island Sound. Bull. Bingham Oceanogr. Collect. Yale Univ. 18(2), 101 p. RUSS, G. R. 1980. Effects of predation by fishes, competition, and structural complexity of the substratum on the establishment of a marine epifaunal community. Exp. Mar. BioI. Ecol. 42:55-69. J. SEED, R. 1976. Ecology. In B. L. Bayne (editor), Marine mussels: their ecology and physiology, p. 13-66. Cambro Univ. Press, Cambridge, Engl. SCHOENER, T. W. 1974. Resource partitioning in ecological communities. Science (Wash., D.C.) 185:27-39. SIEGEL, S. 1956. Nonparametric statistics for the behavioral sciences. McGraw-Hili, N.Y., 312 p. SLOBODKIN, L. B. 1961. Growth and regulation of animal populations. Holt, Rinehart and Winston, N.Y., 184 p. TENORE,K. R. 1978. Macroinfaunal benthos. South Atlantic OCS Benchmark Program, 1977 report. Vol. III, p. 257-288. Bureau of Land Management, New Orleans, La. WENNER, E. L., D. M. KNOTT, R. F. VAN DOLAH, and V. G. BURRELL, Jr. 1983. Invertebrate communities associated with hard bollom habitats in the South Atlantic Bight. Estuarine Coastal Shelf Sci. 17:143-158. WILLIAMS, W. T., and J. M. LAMBERT. 1961. Multivariate methods in plant ecology. III. Inverse association-analysis. J. Ecol. 49:717-729. 16 Appendix 1. Phylogenetic list of invertebrate taxa collected by dredge during 1981 cruises to the stations off South Carolina and Georgia. w = winter, sp = spring, s = summer and f = fall. No dredge samples were collected during winter samples at station OS06. IS0l IS02 MS02 f f f - w,sp,f - f f - sp,s,f f f w w,sp w,s w,sp,f f w,sp,s,f w,sp,s,f w,sp,s,f f sp,f sp,f sp w,f f - - - - - - sp,f sp,f sp sp sp w w f w,sp ,s,f MS06 OSOl OS06 Porifera Demospongea - Ceractinomorpha .... -..I Keratosa Aplysillidae Chelonaplysilla sp. Darwinellidae Darwinella sp. Spongiidae Aplysina fistularis Euspongia officenalis 1rcinia campana 1rcinia felix 1rcinia ramosa 1rcinia sp. B 1rcinia strobilina Oligoceras sp. Spongia tubulifera Haplosclerida Adociidae Sigmadocia caerula Strongylophora sp. B Desmacidonidae Xytopsues griseus Haliclonidae Amphimedon compressa Callyspongia fallax Haliclona areolata Haliclona oculata Haliclona sp. C undetermined Haliclonidae Nepheliospongiidae Xestospongia halichondrio1des Poecilosclerida Poecilosclerida H Amphilectidae Tenaciella obliqua Mic roc ion idae Microciona prolifera Microciona sp. D Microciona spinosa Pandaros acanthifolium - - w,s 1;11 - f s - - - - - f sp - w,s,f f f sp - w,s,f f f f f - s,f - w,sp,s,f w,sp w w w,sp,s,f w w - - - sp - - w - w f w,s,f s w w,f - sp f s - w,sp,s,f sp,f f w,sp - - - - - f f s,f Appendix l. (Continued) IS01 Thalyseurypon sp. Thalyseurypon foliacea Thalysias juniperina Thalysias sp. Mycalidae Neofibularia noli tangere Myxillidae undetermined Myxillidae Plocamiidae Holoplocamia sp. Holoplocamia sp. A Tedaniidae Lissodendoryx isodictyalis Lissodendoryx sigmata Tedania ignis 00 Halichondriida Halichondriidae Ciocalapata gibbsi Halichondria bowerbanki IS02 MS02 - MS06 OSOI OS06 - - - f w,sp,f sp - sp w,sp,s,f f - f - - sp - sp - - - - f sp w - f s w w,sp,f w,sp,f - - - w sp s,f w,sp,f f sp,f sp,f w - s w,sp,s,f w,sp,s,f f f - f w f f - Demospongea - Tetractinomorpha Axinellida Axinellidae Axinella bookhouti Homaxinella rudis Homaxinella sp. A Homaxinella waltonsmithi Phakellia folium Phakellia lobata Pseudaxinella lunaecharta Pseudaxinella sp. A Pseudaxinella sp. B Pseudaxinella wilsoni Teichaxinella corrugata Teichaxinella morchella Teichaxinella shoemakeri undetermined Axinellidae Demoxyidae Higginsia strigilata Euryponidae Tricheurypon viride Hemiasterellidae undetermined Hemiasterellidae w,sp,f w sp,f w,sp,s,f - w w sp - s - w,sp,f w,sp,f s,f w,sp,f w w,sp,f - w sp sp sp,f - - - f f sp w,s,f f w,f - - - f - - - - f f w,f f - - f sp w sp w,sp,s,f w w,sp,s,f f - sp sp,s - - f Appendix l. (Continued) 1S0l Raspaliidae Ectyoplasia ferox Hemectyon pearsei Raspaciona sp. A Hadromerida Clionidae Cliona caribbaea Spirastrellidae Anthosigmella varians Columnitis sp. Spheciospongia othella (?) Spheciospongia vesparium Spirastrella coccinea Timea mixta Tethyidae--Tethya crypta \0 Choristida - Astrophorina Ancorinidae Myriastra fibrosa Myriastra incrustata Myriastra kallitetilla Myriastra sp. A Geodiidae Erylus sp. Geodia gibberosa Stelleta grubii Stelleta sp. A Choristida - Spirophorina Craniellidae Cinachyra alloclada Cinachyra kuekenthali Cinachyra rhizophyta Choristida - Chondrosiina Chondrillidae Chondrilla nucula Chondrosiidae Chondrosia collectrix sp sp 1S02 MS02 MS06 OS06 w,sp,f sp f - w w,sp,f sp,f f w,s,f w,sp w,sp,f - - - f w,sp w,sp,s,f f - sp,f f - s sp w w sp sp,f w w w w - - - sp sp s w,f - - w w sp - - - - - - s w,sp,s,f w,f - f f - - - w f w,s,f w w w w,sp,f w,sp,f w,sp,f - f f f w f - sp w sp,f - - - sp - - - - - Calcarea Leucettidae Leucetta sp. A OSOl f f f Appendix l. (Continued) Leucetta f10ridana Leuconiidae Leuconia aspera MS02 MS06 OSOl OS06 - - - f w,sp w,f s f - - w,f sp - - f f w,sp f - f s,f w s,f f 1S01 1S02 - - w,s sp Calcarea - Ca1cinea C1athrinida C1athrinidae C1athria sp. C1athrina coriacea s,f w - Calcarea - Ca1caronea Scyettida Scyettidae Scypha barbadensis s,f Cnidaria Hydrozoa N 0 Anthomedusa/Athecata Bougatnvi11iidae Bimeria humi1is Bougainvi11ia sp. C1avidae Turritopsis fascicu1aris Turritopsis nutricu1a Eudendriidae Eudendrium carneum Eudendrium sp. Eudendrium tene11um Hydractiniidae C1avactinia sp. Hydractiniidae B Leptomedusae/Thecata Campanu1ariidae Campanu1aria hincksii C1ytia cy1indrica C1ytia fragilis Obelia dichotoma Ha1eciidae Ha1ecium diminutivum Ha1ecium dysymmetrum Ha1ecium sp. Ha1ecium tene11um f - s,f w,s,f s,f - sp - - s sp,s,f w,sp,s,f s - - - s w,sp s,f sp,s sp,f w,sp,s,f s sp,s f sp sp sp s,f sp,f - s,f w,f sp,s,f w,sp,s - - sp - sp w,sp,s,f w w,s w,sp,s w,sp,s,f w,sp,s,f w,sp,f w,sp,s,f w,sp,s,f w,sp,f w,f w,f w,sp,s,f w,sp,f w,sp,s,f w,sp,s w,sp,s sp,s,f sp,s,f sp sp,s,f s w,sp,s w,sp,s,f w,sp w,sp,s w,sp,s,f w,f sp,s w,sp,s w,sp sp,s,f sp,s w,sp,s,f w,sp,s,f sp,f sp,f sp,f s,f - W,sp,S Appendix l. (Continued) 1S01 N Lafoeidae Fi1e11um serra tum Hebe11a scandens Hebe11a venusta Lafoea fruticu10sa Scandia mutabi1is Laodiceidae Modeeria rotunda P1umu1ariidae Ag1aophenia a11mani Ag1aophenia e10ngata Ag1aophenia 1atecarinata Ag1aophenia trifida Antene11a secundaria Gymnangium sinuosum Ha10pteris c1arkei Monostaechas quadridens Nemertesia simplex P1umu1aria margaretta P1umu1aria setacea Schizotricha tene11a Sertu1ariidae Dynamena cornicina Dynamena da1masi Dynamena quadridentata Sa1acia desmoides Sertu1are11a areyi Sertu1are11a conica Sertu1are11a pinnigera Sertu1are11a sp. A Sertu1aria dis tans Sertu1aria marginata Sertu1aria p1umu1ifera Thyroscyphus marginatus Syntheciidae Hinckse11a cy1indrica Synthecium iubitheca - - 1S02 w w w MS02 MS06 OSOl OS06 s sp w,s,f - w,sp,s,f w,sp,f w,sp,f sp,f sp,s,f sp sp,f sp,f w,sp,s,f sp,s,f - - sp w,f w,sp,s,f f w,f - - - f w - - sp,f f sp,s,f w,sp,s,f sp sp sp,s,f w,sp,s,f - - - w,sp,f w,sp,f sp s s f w,sp,s,f f w w f w w,sp,s,f - - - - s - - sp w,sp,s, f w,sp,s,f w,sp,s,f - - w, sp f w,sp,s,f w,s w w,sp,s,f w,sp w,sp,s,f s sp,s,f w,sp,s,f - w,sp,s,f f w,f sp,s w,sp,s,f sp,s w,sp,s,f sp,s,f f sp,s,f w,sp,s,f w,sp,s,f w,sp w,sp,s,f s,f w,sp,s,f sp,f w,sp,s,f w,sp,s,f w,sp,s sp,s w,f - - w,sp,s,f f w,s,f w,sp,f w,sp,s,f w,sp,f s,f s - - w,s,f w,sp w,f - - - - w,sp,s,f - - - - w,sp,s,f f f - - w,sp s w,sp,s,f s w,sp,s,f w,s,f sp sp,s,f sp,s sp,s,f sp,f sp,s,f sp,s,f sp,s,f sp,s,f w,sp s w,s sp,s w,s w,sp,s,f w,sp,s w,s s,f sp,f sp,f sp,s,f sp,f sp sp,s,f f s w,sp,s w,f sp,f - sp,f sp,f - f sp,s sp,f Anthozoa - Octora11ia Te1estacea Te1estidae Te1esto fruticu10sa Te1esto sanguinea w,s,f w,s w,sp,s,f w,sp,s,f w,sp,s,f w,sp,s,f sp,f sp,s,f Appendix l. (Continued) Alcyonacea Nidaliidae Nidalia occidentalis Gorgonacea Anthothelidae Diodogorgia sp. Titanideum frauenfeldii Ellisellidae Ellisella sp. Gorgoniidae Leptogorgia virgulata Lophogorgia cardinalis Lophogorgia hebes Paramuriceidae Thesea sp. Plexauridae Muricea pendula N N Pennatulacea Renillidae Renilla reniformis Virgulariidae Stylatula sp. ISOl IS02 MS02 - - - MS06 OSOl OS06 - - s sp,s,f - w,f w,sp,s,f s,f sp,s,f - w,f sp,s,f w,sp,s,f - f sp,f w,sp,s,f f f w,sp w,sp,s,f sp - - f sp,f w,sp,s,f - f - - w,sp - - - - w - - - - w,s,f w,sp,f sp w,sp w,s w sp,f f - sp,f s - - w,·sp,s,f w,sp,s,f - - w,sp,s,f w,sp,s,f w,sp,s,f - - w,sp,f - - - sp,s,f sp,f Anthozoa - Zoantharia Actiniaria undetermined Actiniaria Scleractinia - Faviina Faviidae Solenastria hyades Oculinidae Oculina arbuscula Oculina sp. w,sp,f w,f w,sp Rhizangiidae Astrangia astreiformis Phyllangia americana - - - f f Scleractinia - Dendrophylliina Dendrophylliidae Balanophyllia floridana - - - - w,s,f f Appendix l. (Continued) Antipatharia undetermined Antipatharian IS0l IS02 MS02 f s w,f - w,sp,f w w,sp - w,sp w,s,f - sp w,sp,s w,sp,f MS06 OSOI OS06 Mollusca Gastropoda - Prosobranchia Archaeogastropoda Fissurellidae Diodora cayenensis Trochidae Calliostoma euglyptum Calliostoma roseolum IV w Mesogastropoda Crepulidae Crepidula aculeata Crepidula plana Cymatiidae Cyphoma mcgintyi Distorsio constricta mcgintyi Cypraeidae Cypraea spurca acicularis Eratoidae Erato maugeriae Mytridae Vexillum ~ern atum Ovulidae Simnia uniplicata Turritellidae Vermicularia knorrii Velutinidae Lamellaria ~ i c u a Vermetidae Spiroglyphus irregularis Neogastropoda Buccinidae Pisania tincta Columbellidae---Anachis avara Anachis lafresnayi Nassarina minor Suturoglypta iontha Conidae Conus floridanus - - - w w,sp - - - sp - - - sp - sp - sp,f - - - - - f - - - - s - - - w f w,sp,s,f - sp,s,f f - - - - s - - sp - w sp,f sp - sp - w sp w,sp,s,f - w s - - - - f f - s - - - - Appendix l. (Continued) Faseio1ariidae Faseio1aria 1i1ium hunteria Faseio1aria tu1ipa P1europ1oea gigantea Marginellidae Margine11a apieina Murieidae Ca1otrophon ostrearum Favartia ce11u1osa Murex f10rifer di1eetus Murex pomum Murex sp:-B Nassariidae Nassarius a1bus Turridae Kurtzie11a 1imonite11a IS01 IS02 MS02 MS06 s sp w,sp - - - - s - - - w w - - - - w w,sp w - - w,f - - - w - - - sp - - - OSOl OS06 f sp s Pe1eeypoda tv ~ Areoida Areidae Anadara notabilis Area imbricata Area zebra Areopsis adamsi Barbatia candida Mytiloida Mytilidae Lithophaga bisu1eata Modiolus amerieanus Musculus 1ateralis Pterioida Anomiidae Anomia squamula Pododesmus rudis Peetinidae Ch1amys benedieti Plieutulidae P1ieatula ~ib osa Pteriidae Pteria eo1ymbus Spondy1idae Spondy1us amerieanus w,sp,s,f w,s - - sp w,sp,s,f f - w,sp,s,f - - - w,sp w,sp,f - - sp,f w,sp,s,f - - f w,f w,s,f s,f - - f w,sp,f w,sp,s,f w,sp,f f f - - f sp sp - - - s w,f - f - - f - w,SP,S,f - w,sp,s,f w w,sp,s w,sp,f s,f f s s f s Appendix l. (Continued) ISOI Ostreina Ostreidae Lopha frons Ostrea permolis IV VI Veneroida Cardiidae Americardia media Laevicardium pictum Chamidae Chama congregata Chama macerophylla Chama sarda Pseudochama radians Tellinidae Tellina americana Veneridae Chione grus Chione latilirata Macrocallista maculata Myoida Corbulidae Corbula chittyana Gastrochaenidae Gastrochaena hians Hiatellidae Hiatella arctica Pholadomyoida Lyonsiidae Lyonsia beana Octopoda Octopodidae Octopus joubini Octopus sp. s IS02 - f W,sp,S w,s,f s - - W - sp sp f - sp,f w,s,f w,sp,f w,f f w,sp - - - - w,sp OS06 MS06 - w - OSOI MS02 sp,f sp - w s,f w - - w - - - - - - - - - w,f - sp,s,f sp sp,s,f w,s,f w,s,f W,sp w,sp sp,S sp - - - sp f - - - - - f f - w Arthropoda - Mandibulata Crustacea - Cirripedia Thoracia - Lepadomorpha Scalpellidae Scalpellum diceratum f Appendix l. (Continued) Thoracia - Ba1anomorpha Archaeoba1anidae Conopea ga1eata Conopea merri11i Ba1anidae Balanus ca1idus Balanus trigonus Balanus venustus 1S01 1S02 MS02 w w,f s w,sp,s,f w,sp,s,f sp,f w,sp,s,f w,sp,f - sp w,sp,s,f w,s,f w,sp,s - w,s w,sp - - sp - w,sp MS06 OSOl - w,sp,s s w,sp,f s w,sp,s,f sp,f SP,B w,sp,s,f w,sp sp sp,f sp f s OS06 Crustacea - Malacostraca Stomatopoda Gonodacty1idae Gonodacty1us bredini IV 0'1 Decapoda - Pleocyemata - Caridea A1pheidae A1pheus formosus ?yna1pheus fritzmue11eri ~na!pE.e_u~ lon&.!..carpus Syna1pheu~ minus Synalpheus townsendi Decapoda - Pleocyemata - Pa1inura Scy11aridae Scy11arus chacei Scy11arus depressus sp sp sp,s,f - s,f sp w, sp, s, f - .- - sp,s w,sp,s - w sp - sp,s - - - - - - - sp s w,s,f f f - f Decapoda - P1eocyemata - Anomura Diogenidae Dardanus fucosus Dardanus insignis Paguristes sericeus Paguristes tortugae Petrochirus diogenes Ga1atheidae Ga1athea rostrata Paguridae Pagurus caro1inensis Py10pagurus cora11inus Porcellanidae Mega10brachium soriatum Pachyche1es rugimanus Petro1isthes ga1athinus Porce11ana sigsbeiana sp - sp w,sp - - - - - sp - sp - - - - - - w w,sp,f sp s - sp - - w,sp,s - - w sp - w - s,f s - - sp,f - w,sp w,s - w w,s,f sp f Appendix t-) -..I l. (Continued) Decapoda - Pleocyemata - Brachyura Calappidae Calappa angusta Osachila semi levis Cancridae Cancer irroratus Dromiidae Dromidia antillensis Grapsidae Euchirograpsus americanus Majidae Macrocoeloma camptocerum Macrocoeloma trispinosum Metaporaphis calcarata Mithrax acuticornis Mithrax forceps Mithrax pleuracanthus Pelia mutica Podochela gracilipes Podochela riisei Stenocionops furcata coelata Stenorhynchus seticornis Parthenopidae Parthenope fraterculus Pinnotheridae Pinnotheres maculatus Pinnotheres ostreum Portunidae Portunus ordwayi Raninidae Hypochonca sabulosa Symethis variolosa Xanthidae Carpoporus papulosus Glyptoxanthus ~ Pilumnus dasypodus Pilumnus floridanus Pilumnus pannosus Pilumnus sayi Pseudomedaeus agassizii Xanthidae A ISOl IS02 - - - MS02 - MS06 - OSOl OS06 - sp,f - w - w,s,f s s w,s s,f w w,sp sp,s f - - - w w,sp f sp - - w w,s w,sp,f f f w,sp,f sp - - w,sp,s,f - - - - W,sp - - W,sp - s f s f - w,sp f - w,s ,f - w - w s - - - - w,s,f sp - w w - w - - - - - - - f - - f - w,sp - - - - s sp w,sp,s,f - - - w,sp,s - s s w,sp w,sp,f w,sp,s,f w,sp sp,s,f sp,s,f w,sp,s - - - w,sp s,f w,sp,f f s,f sp sp,s s s,f Appendix 1. (Continued) ISOl IS02 MS02 MS06 OSOl OS06 w,sp,f w,sp,s,f w,sp,s,f w,sp,s,f w,s s,f w,sp,f w,sp,s,f w,sp,s sp,f Bryozoa Stenolaemata Cyclostomata Crisiidae Crisia sp. Diaperoeciidae Diaperoecia floridana Lichenoporidae Lichenopora radiata Oncousoeciidae Proboscina floridana Plagioeciidae Plagioecia dispar - w - sp s w,sp,s,f - s,f - - - sp,s,f sp,f - - - - - w,sp w sp,f sp,s,f s,f w w,sp - - sp w w w,sp,s - w f w w,sp,s - w,s - sp w,sp,s - w,sp,s,f w w,s,f w,sp,s,f f s sp,s,f Gymnolaemata IV 00 Ctenostomata undetermined Ctenostomata Alcyonidiidae Alcyonidium sp. Buskiidae Buskia sp. Nollelidae Nolella stipata Valkeriidae Aeverrillia setigera Vesiculariidae Amathia alternata Amathia convoluta Amathia distans Victorellidae Sundanella sibogae Cheilostomata - Anasca Aeteidae Aetea anguina Alderinidae Alderina smitti Antropora tincta Copidozoum tenuirostre Crassimarginatella leucocypha Parellisina curvirostris Parellisina latirostris w,sp w - sp,s W,sp w,sp,f w,sp w,sp - - - f - w,sp sp,s,f w,sp,f - sp - s w w,s,f sp sp,s sp,s,f sp w,sp,s,f w,sp,s w,sp,s,f sp sp,s,f - - w,sp,s,f w,sp,s,f w,s W,sp,S sp,s,f sp,f - sp sp,f sp,f sp,f f w,sp,s,f s,f w.s sp,f sp,s,f - sp,f w - Appendix l. (Continued) MS02 MS06 OSOl OS06 w w,sp,s,f f w,sp,f f - sp - - sp,f w w sp w,sp,s,f - s - - - - sp s,f sp - - w,sp w,sp,f f sp f w,s - - - - - sp s - w,sp,s,f w,sp,s,f w - w sp sp w,sp,f w,sp,f w - IS01 N ID Bugulidae Bugu1a fu1va Bugu1a ~ Bugu1a johnstonae Bugu1a microoecia Bugu1a ry1andi Bugu1a sp. Bugu1a turrita Cau1ibugu1a dendrograpta Cau1ibugu1a pearsei Ce11ariidae Ce11aria nodosa Chaperiidae Chaperia sp. Cupu1adriidae Cupu1adria canariensis Cupu1adria doma Discopore11a umbe11ata Farciminariidae Nellia tene11a Hincksinidae Ap10usina gigantea Ap10usina tuberosa Membraniporidae Membranipora arborescens Membranipora tenuis Microporidae Micropora coriacea Onychoce1lidae F10ridina antiqua F10ridina parvice11a Smittipora 1evinseni Scrupoce11ariidae Caberia boryi Scrupoce11aria bertho11eti Scrupoce11aria regu1aris Steganopore11idae Steganopore11a magni1abris Chei1ostomata - Cribrimorpha Cribrilinidae Be11u1opora be11u1a Cribri1aria f10ridana Cribri1aria radiata Membranipore11a aragoi undetermined Cribri1inidae - - - w,sp,s,f IS02 - - f sp sp,f sp s,f - sp,s,f sp,s f Bp w,f w,sp,s,f f sp,s,f s - f sp,f - - - - f w w,s - - w - - - - - sp s - - - - - sp,s,f - w,sp,s,f s w,sp s sp,s sp sp w,sp,f w,sp,s,f sp w,sp,s,f sp sp,f sp,s,f w,sp,f - - f w,sp,s,f - - - sp,s sp,s w w,f sp,f sp w sp,s,f - - - s,f - sp,s,f f sp,s,f - w,sp - sp w,sp,s,f w sp s sp,s - w,sp,s,f w,sp,s,f w w,sp w,sp,f - Appendix 1. ....o ( Continued) Cheilostomata - Ascophora Adeonidae Reptadeonella hastingsae Celleporariidae Celleporaria albirostris Ce~lepora ia magnifica Celleporaria mordax Trematoecia turrita Celleporinidae Celleporina hassalli Turbicellepora dichotoma Cheiloporinidae Hippaliosina rostrigera Crepidacanthidae Crepidacantha sp. Crepidacantha poissoni teres Exochellidae Exochella longirostris Triporula stellata Hippoporinidae Hippopleurifera mucronata Hippoporidra calcarata Hippoporidra janthina Hippoporina contracta Hippoporina verrilli Hippothoidae Tryposostega venusta Microporellidae Microporella ciliata Microporella marsupiata Microporella umbracula Petraliidae Petraliella bisinuata Petraliella marginata Phylactellidae Phylactella aviculifera Schizoporellidae Arthropoma cecili Cleidochasma-porcellanum Cycloperiella rubra Escharina porosa Schizop re~rnuta Schizoporella errata Schizoporella fIOridana Stylopoma informata ISOl IS02 MS02 MS06 OSOl Oso6 w,sp,f w,sp,s,f w,sp,s,f w,sp,s,f w,sp,s,f sp w,f w,sp,s,f w,sp,s,f sp,s,f s w,sp,s,f w,sp,s,f w,sp,s,f w,s sp,s,f w,sp,s s,f w,s,f s f w,sp,s w,sp,s w,sp,s,f w,sp,f w,sp,s w,sp,s w,sp,s,f w,sp,s,f w,sp,s,f w,sp,f sp w,sp s sp sp,f w,sp,s sp sp sp,f sp,s sp,s,f sp,f s w w,sp f w,sp,s,f s w,sp,s,f w,sp,s,f sp,s,f w,sp,f w,sp,s,f sp,s,f w,sp,f w,sp,s,f w,sp,s,f w,sp,s,f sp,s,f w,sp,s w,sp,s,f sp,s,f w,sp,s,f sp,f w,sp,s,f w,sp,s,f w,sp,s,f w w,sp,s,f w,sp,s,f sp sp sp,s,f sp s w,sp,s w s w,sp,s,f sp w,f w,sp,s,f w,sp,f w,sp,s,f s,f w,f sp sp,s w sp,s,f f sp,f f f f f w,sp w,sp,s,f sp w,sp,f w,f w,sp,s,f w,sp,f sp,s,f w,sp,s,f s w,sp,s sp,s,f sp,s,f sp,s sp,s,f w,sp,s,f sp,s,f sp Appendix l. (Continued) IS02 MS02 - sp sp - sp - s w,sp,s,f sp w,sp,s,f w,f - - - - - sp,s,f w w,sp,f - - - - - - ISOl Sertellidae Rhyncozoon rostratum Rhyncozoon tuberculatum Smit tinidae Parasmittina nit ida Parasmittina spathulata Smittina smittiella Stomachetosellidae Cigclisula serrulata - - w - w,sp,s,f w MS06 - OSOl OS06 w w,sp,s,f s s sp,s,f - - - f sp,f Echinodermata Crinoidea Comasteridae Comactinia meridionalis lM Stelleroidea - Asteroidea Platyasterida Luidiidae Luidia alternata Luidia bernasconiae Luidia clathrata Paxillosida Astropectinidae Astropecten articulatus Astropecten comptus Astropecten duplicatus Astropecten nitidus Astropecten sp. Valvatida Ophidiasteridae Narcissia trigonaria Spinulosida Echinasteridae Echinaster brasiliensis Echinaster modestus Echinaster serpentarius Echinaster sp. - s - - - - - w sp s - - w,sp W,s - f sp,f w sp f sp,S - - - - - - - - - sp sp,f - sp wts,f s w w,sp,f s,f Appendix l. (Continued) Forcipulata Asteriidae Asterias forbesii Phyronophiurida - Euryalina Gorgonocephalidae Astrophyton muricatum Astroporpa annulata w IV Ophiurida Goniasteridae Tosia parva Gorgonocephalidae Astrocyclus caecilia Ophiac tidae Ophiactis savignyi Ophiochitonidae Ophionereis reticulata Ophioderma tidae Ophioderma brevispinum Ophiothricidae Ophiothrix angulata Ophiothrix suensonii ISOl IS02 w,sp,s,f sp - - MS02 MS06 OSOl OS06 w,sp,s,f sp,s,f - f w - - - - - - - - - - sp - - s,f - f - - - - - f - f - - f w,sp,s,f sp,s,f sp,s,f s,f w,sp,s,f - - - f s,f f - - - sp w,sp, f s w,sp,s w,sp,s,f f w, f s w,sp,s,f w,sp,f - sp s W,s sp - f sp,s s w,sp,f sp f sp,f - - - s Echinoidea Cidaroida Cidaridae Eucidaris tribuloides ------ Arbacoida Arbaciidae Arbacia punctulata Temnopleuroida Toxopneustidae Lytechinus variegatus Clypeasteroida Clypeasteridae Clypeaster prostratus Clypeaster subdepressus Mellitidae Encope michelini Leodia sexiesperforata Cassiduloida Echinolampadidae Echinolampas depressa - - Appendix l. (Continued) ISOI Spa tango ida undetermined Spatangonidae - IS02 - MS02 MS06 OSOI OS06 - - - f sp sp - s Holothuroidea Dendroehirotida Cumariidae Oenus pygmaeus Aspidoehirotida Synallaetidae Isostiehopus badinotus w,sp,s,f w,sp,f - - - w,f - - w f - sp,s,f s sp w,sp,s,f w,sp,s,f sp,s - w sp,s,f w,sp,s,f sp,f w sp,f sp sp,s sp,s sp,s,f sp s,f f w,sp,f f sp,s s,f s,f s s,f s· f Chordata Aseidiaeea Aseidiaeea A undetermined Aseidiaeea t..> t..> Enterogona Aseidiidae Aseidia eurvata Clavelinidae Clavelina gigantea Clavelina pieta Clavelina sp. Distaplia bermudensis Eudistoma earolinense Eudistoma hepatieum Eudistoma sp. Didemnidae Didemnum eandidum Didemnum sp. Didemnum soeeiosum Diplosoma maedonaldi Tridemnum savignii Tridemnum sp. Polyelinidae Aplidium eonstellatum Aplidium pellueidum Aplidium sp. A Aplidium sp. B Aplidium stellatum w,sp,s,f - w,sp,f - - - w,sp sp sp w f f f w,sp,s,f S sp,s w,sp,s,f s,f w,sp,s,f sp w,s - sp - s sp,f - - sp - f - f - f s,f f s,f - w w w - '- w Appendix l. (Continued) Pleurogona Molgulidae Molgula occidentalis Styelidae Botrylloides nigrum Botryllus planus--Polyandrocarpa floridana Polyandrocarpa gravei Polyandrocarpa tincta Styela partita Styela plicata Symplegma viride 1M ~ IS01 IS02 MS02 MS06 w,sp,s,f w,sp,s,f w,sp,f w,sp,s,f s sp w w,sp,f w,sp,s,f f - sp sp - - w,sp - sp,s,f w,sp,s,f f s f sp sp w,sp,s - - sp,s w,sp,f f OSOI OS06 f f - w,sp s Appendix 2. Phylogenetic list of invertebrate taxa collected by trawl during 1981 cruises to the stations off South Carolina and Georgia. w = winter, sp = spring, s = summer and f = fall. No trawling was done at station OS06 during any season. 1SOl 1S02 MS02 MS06 OSOl Porifera Demospongea - Ceractinomorpha w Ul Keratosa Keratosa D Aplysillidae Chelonaplysilla sp. Darwinellidae Darwinella sp. Dysideidae Dysidea variabilis (?) Polyfibrospongia echina Spongiidae Aplysina fistularis Aplysina lacunosa Euspongia officenalis Hyatella intestinallis 1rcinia campana 1rcinia felix 1rcinia ramosa 1rcinia strobilina Spongia tubulifera Haplosclerida Adociidae Sigmadocia caerula Strongylophora sp. B Desmacidonidae Xytopsues griseus Xytopsues sp. Haliclonidae Amphimedon compressa Callyspongia fallax Callyspongia vagInalis Haliclona areolata Haliclona oculata Haliclona palmata Haliclona sp. C Niphantes digitalis Niphantes erecta Nepheliospongiidae Xestospongia halichondrioides Poecilosclerida Poecilosclerida H Amphilec tidae Tenaciella obliqua f sp,s s,f w,f s w,sp,f sp,f s w w w f w s w f w,sp,s f sp s s w,sp sp w,sp,s,f s w,sp,s,f w,sp s w,sp,s,f w,sp,s,f s w,sp,s s sp,s,f w,sp,f sp w,f s w,sp,s,f w w sp w W,sp,S w s s,f w,sp w w s,f f Appendix 2. (Continued) 1501 w 0'1 Nic roc ionidae Microciona prolifera Microciona sp. Microciona spinosa Pandaros acanthi folium Thalyseurypon foliacea Thalysias juniperina Thalysias sp. Mycalidae Mycale sp. Mycale sp. A Neofibularia noli tangere Zygomycale parishii Plocamiidae Holoplocamia sp. Tedaniidae Acarnus innominatus lotrochota birotulata Lissodendoryx isodictyalis Lissodendoryx sigmata Lissodendoryx sp. B Tedania ~ i s Halichondriida Hali.chondriidae Ciocalapata gibbsi Halichondria bowerbanki Halichondria sp. 1502 M502 M506 0501 5, 5 - w, f s,f - - - 5 5 - W W W,sp,S 5 sp s,f W,sp,s,f w,sp,s 5 w,sp,s,f - - - - - - - w,sp,s,f f - - - sp - - - W s,f - - sp,s,f sp,f W sp W,sp sp,s,f - - - f 5 sp,s f 5 sp sp sp 5 sp w,sp,s,f sp,s,f sp w,sp,s,f w,sp,s,f w,f s,f sp,s,f sp,s 5 w,sp,s w,S, f sp w,sp,s,f w,sp,s - - - W 5 Demospongea - Tetractinomorpha Axinellida Axinellidae Axinella bookhouti Homaxinella rudis Homaxinella sp. A Homaxinella waltonsmithi Phakellia lobata Pseudaxinella lunaecharta Pseudaxinella sp. A Pseudaxinella sp. B Pseudaxinella wilsoni Teichaxinella corrugata Teichaxinella morchella undetermined Axinellidae Demoxyidae Higginsia strigilata sp,f - W w,sp,s,f w f w 5 w,sp,s w,s w,sp,s,f w,sp,s,f w,s,f s,f w sp - W w,s,f 5 s,f sp,f sp,s,f sp,s,f - - - - - sp,f 5 - - - s,f - sp,s,f s,f - w,sp,s,f f w w 5 5 sp,s,f 5 f w,sp,s,f 5 w,f Appendix 2. (Continued) 1S0l Raspaliidae Ectyoplasia ferox Ectyoplasia sp. A Hemectyon pearsei Raspaciona sp. A Raspaciona sp. B W -..I Hadromerida Clionidae Cliona caribbaea Cliona celata Cliona lampa Cliona sp. Coppatidae Spirasigma sp. (?) Placospongiidae Placospongia carinata Podospongiidae Podospongia sp. Spirastrellidae Anthosigmella varians Spheciospongia othella (?) Spheciospongia vesparium Spirastrella coccinea Timea mixta Suberit d~ Polymastia sp. undetermined Suberitidae Tethyidae Aaptos aaptos Tethya actinia Tethya crypta Choristida - Astrophorina Ancorinidae Myriastra fibrosa Myriastra incrustata Myriastra kallitetilla Myrias tra sp. A Geodiidae Geodia gibberosa Stelleta grubii Stelleta sp:-A W w,s W - w,sp,s 1S02 s W w,sp,s,f s f w,sp,s,f MS02 - w,sp,s,f MS06 - sp - w,sp,s s - - - - - - sp - - - - - - - s - w,s,f w,sp,s,f s,f sp W,sp - - - w,sp,f f w,sp,s,f w,s w - - - sp,f - w,sp,s,f w,sp,s,f w,sp f - - s s,f f - s - - - - w,sp,s,f - - - - sp sp sp sp - f w - - - OSOl - - - f w,s,f w,f s sp w sp - sp s - f w,sp,s - sp w,sp,s ,f - - w,sp s w,s,f Appendix 2. (Continued) Choristida - Spirophorina Craniellidae Cinachyra alloclada Cinachyra kukenthali Cinachyra rhizophyta ISOI IS02 MS02 MS06 w,sp,s w,s,f w,s w,sp,s,f w,s,f s w w,sp,s s,f w,sp,s,f Choristida - Chondrosiina Chondrillidae Chondrilla nucula Chondrosiidae Chondrosia collectrix - w w,sp,f w - w,sp sp Lithistida Scleritodermidae Scleritoderma paccardi (7) - - - - - - sp sp sp - w,sp OSOI s sp Calcarea IN 00 Leucettidae Leucetta Leucetta Leuconiidae Leuconia Leuconia sp. A floridana aspera crustacea w,sp,s w - sp,f sp - w,sp w W,sp sp Calcarea - Calcinea Clathrinida CIa thrinidae Clathria sp. A Clathrina coriacea Leucosolenia canariensis Leucosolenia cancellata - w,sp,s,f - s s sp,s,f - w,sp,s - Calcarea - Calcaronea Scyettida Scyettidae Scypha barbadensis Scypha sp. A w,sp,s,f sp sp,s,f - w sp Cnidaria Scyphozoa Semaeostomeae Cyaneidae Cyanea capillata w Appendix 2. (Continued) IS0l Ulmaridae Aurelia aurita Rhizostomeae Stomolophidae Stomolophus meleagris IS02 MS02 MS06 OSOI - - s,f f - w,s,f Cubozoa Carybdeidae Tamoya haplonema - f Hydrozoa IN \0 Anthomedusa/Athecata Bougainvilliidae Bimeria humilis Bougainvillia sp. Clavidae Turritopsis fascicularis Turritopsis nutricula Eudendriidae Eudendrium carneum Eudendrium sp. Eudendrium tenellum Hydractiniidae CIa vac tinia sp. Hydractiniidae A Tubulariidae Ectopleura sp. Leptomedusae/Thecata Campanulariidae Campanularia hincksii Clytia 'cylindrica Clytia edwardsi Clytia fragilis Clytia paulensis Obelia dichotoma Campulinidae Cuspidella humilis Haleciidae Halecium diminutivum Halecium dysymmetrum Halecium sp. Halecium tenellum sp,s,f w,sp,s,f s,f f s,f w,s,f s,f f w,sp,s,f w,sp,s - s sp s,f W,sp w,s,f w w,sp,f s w,sp,s,f w,sp,s,f w,sp,f sp,s,f w,sp,s,f f sp,s,f sp,s,f f w,sp,s,f w,sp,s,f s,f w,sp,s,f w,s,f sp w,s,f - - - - w - - - - W,sp w,sp,s,f w,sp,s,f w,sp,s,f sp w,sp,f w,sp,s,f w,sp,s,f w,sp,s,f w,sp,s,f s w,sp,s,f w,sp w,sp,s,f w,sp,s w,sp,s,f w,sp,s,f sp,s w,sp,s,f w,sp,s,f sp w,sp,f - w,s s,f s w,s,f w,sp,s,f w,sp,s,f - f w,s w,sp,s,f f - w,sp,s,f w,sp,s,f w w,sp,s,f w,sp,s,f - - - w,sp sp,s,f w,sp,s,f w,sp,s,f Appendix 2. (Continued) ISOI .j>. 0 Lafoeidae FiIeIIum serra tum HebeIIa scandens HebeIIa venusta Scandia mutabilis Laodiceidae Modeeria rotunda PIumuIariidae Aglaophenia aIImani Aglaophenia elongata Aglaophenia Iatecarinata Aglaophenia trifida Antenella secundaria Cladocarpus flexilis Gymnangium sinuosum Halopteris clarkei Halopteris diaphana Monostaechas quadridens Nemertesia simplex Plumularia margaretta Plumularia setacea Sertulariidae Diphasia tropica Dynamena cornicina Dynamena dalmasi Dynamena quadridentata Salacia desmoides SertuIareIIa areyi Sertularella conica Sertularella pinnigera SertuIareIIa sp. A Sertularia distans Sertularia marginata Sertularia plumulifera Thyroscyphus marginatus Thyroscyphus ramosus Syntheciidae Hincksella cylindrica Synthecium tubitheca - IS02 MS02 MS06 OSOI - w,sp,s s w,sp,s w,s,£ w,sp,s,f w,sp,s,f w,sp,s,f w,sp,f w,sp,s,f sp,s,f f w,sp,s,f s,f w,sp,s,f w,sp,s,f w,sp,s,f - - - - s s sp,f s,f w,sp,s,f s s sp,s,f w,sp,s,f w,sp,s s sp,s,f w,sp,s,f w,sp,s,f sp,s w,sp,s,f w,sp,s,f sp f w,sp,s,f w,sp,s,f - sp,s,f - - - - - s,f s,f s - w,sp,s,f - f - - w,sp,s,f w,sp,s,f - - - f - s,f - s - - w,sp,s,f sp,s w,sp,s,f w w,sp,s,f w,sp,s,f w,sp,s,f w,sp,s,f w,sp,s,f w,s w,sp,s,f w,sp,s,f sp w,sp,s,f s,f sp,s w,sp w,sp,s,f w,sp,s,f w,sp,s,f w w,sp,s,f - - - sp,s w,sp,s,f w,sp,s,f w,sp,s,f s,f - f w,sp,s,f w,sp,s,f w,sp,s s sp w,sp,s,f f w w,sp,s,f w w,sp,s,f w,sp,s,f w,sp,s,f w,sp,s,f w,sp,s,f f w,sp,s,f w,sp,s w,sp,s,f w,sp,s w,sp,s,f f - - s,f s - s,f w,sp,s,f w,sp,s,f w,sp,s,f w,sp,s,f w,sp,s,f w,sp,s,f - sp,s,f - - w,s,f w,s,f w,sp,s,f w,sp,f w,sp,s w,sp,s,f w,s,f w,s,f w,s,f w,sp,s,f w,s,£ w,sp,s,f w,sp,s,f w,sp - w,sp,s,f s w,sp,s,f w,sp,s,f w,s,f w,sp,s Anthozoa - Octocorallia Telestacea Telestidae Telesto fruticulosa Telesto riisei Telesto sanguinea - w,sp,s,f w,sp,s,f sp,s,f - sp,s w,sp,s,f w,sp,s w,sp,s,f Appendix 2. (Continued) ISOI Gorgonacea Anthothelidae Diodogorgia sp. Titan1QUm rrauenfeldii Ellisellidae Ellisella sp. Gorgoniidae Leptogorgia setacea Leptogorgia sp. A Leptogorgia virgulata Lophogorgia hebes Paramuriceidae Thesea sp. Plexauridae Muricea pendula Pennatulacea Virgulariidae Stylatula sp. "" IS02 MS02 MS06 OSOI - f w,sp,s,f w,s,f w,sp,s,f - - w,sp,s,f w,sp,s,f w,sp,s,f - - - - - - - - w,sp,s,f w,sp,s,f w,sp,s,f w,sp,s,f w,sp,s,f w,sp,s,f w,sp,s,f sp w,sp,s,f sp,s,f f s,f - - s sp,s,f w,sp,s,f sp,f - w,sp,s,f w,sp,f w - - f w,sp,s,f w,sp,s,f w,sp,s,f w,sp,s,f sp,s sp f w - w,sp,s,f Anthozoa - Zoantharia Actiniaria undetermined Actiniaria Scleractinia - Faviina Faviidae Solenastria hyades Oculinidae Oculina arbuscula Oculina sp. Rhizangiidae Phyllangia americana w,sp,s,f w sp,s - - w,sp Scleractinia - Dendrophylliina Dendrophylliidae Balanophyllia floridana - - - - s,f Antipatharia undetermined Antipatharian - f f - sp,s,f Arthropoda - Chelicerata Merostomata - Xiphosura sp,s Appendix 2. (Continued) IS0l Limulidae Limulus E2!YEhemus IS02 MS02 MS06 OSOI - w - - - - sp - - - - sp,s,f w,sp,s,f w,sp,s,f w,sp,s,f w,sp,f w,sp w,sp,s,f w,sp,s,f f w,sp,s,f sp W,sp,s, f w,sp,s,f sp w,sp,s,f w,sp,f s w,sp,s,f w,sp,s,f s w,sp,s,f w,s sp,s,f w,sp,s,f w,sp,f Arthropoda - Mandibulata Crustacea - Cirripedia Thoracia - Lepadomorpha Lepadidae Lepas pectinata Scalpellidae Scalpellum diceratum "'" tv Thoracia - Balanomorpha Archaeobalanidae Conopea galeata Conopea merrilli Balanidae Balanus calidus -----~al nus trigonus Balanus venustus Crustacea - Malacostraca Stomatopoda Gonodactylidae Gonodactylus bredini Decapoda - Dendrobranchiata Penaeidae Mesopenaeus tropical is Metapenaeopsis goodei Penaeus duorarum duorarum Trachypenaeus constrictus Sicyonidae Sicyonia brevirostris Solenoceridae Solenocera atlantidis Decapoda - Pleocyemata - Car idea Alpheidae Alpheidae A Alpheus formosus Alpheus normanni s - - - w,f - w,sp,f f w,sp,s,f w,s,f s,f w,sp,s,f w w,f w,sp - - s - - - w,s sp w,sp - - - w,s,f w,s,f w w,sp,s,f w,sp,s,f s w - w,sp,s,f s,f w,sp,s,f w,sp,f s sp s - Appendix 2. (Continued) IS01 A1pheus sp. A Automate evermanni Syna1pheus apioceros Syna1pheus fritzmue11eri Syna1pheus longicarpus Syna1pheus minus Syna1pheus sp. A Syna1pheus townsendi Hippo1ytidae Lysmata wurdemanni Tozeuma serratum Panda1idae Parapanda1us sp. P1esionika tenuipes Pa1aemonidae Anchistioides antiguensis Peric1imenes perryae (?) Pontonia margarita Processidae Processa tenuioes ~ w Decapoda - P1eocyemata - Pa1inura Scy11aridae Scy11arides nodifer Scy11arus chacei Scy11arus depressus Decapoda - P1eocyemata - Anomura A1buneidae A1bunea ~ibesi Axiidae Axiopsis hirsutimana Diogenidae Dardanus fucosus Dardanus ins ignis Paguristes sericeus Paguristes tortugae Ga1atheidae Ga1athea rostrata Paguridae Pagurus caro1inensis Pagurus defensus Py10pagurus cora11inus Py10pagurus ho1thuisi Porce11anidae Mega10brachium soria tum Pachyche1es rugimanus Petro1isthes ga1athinus - f w,sp,f sp,f - IS02 MS02 MS06 - - - - f - sp,s,f w,sp,s,f w,sp,s,f - w,sp,s,f w,sp,s,f - - OSOl s s sp,s sp s sp,f f sp,s w,sp,s w,sp,f w,sp,f - - f - sp - sp,s - - - - f sp - f sp - - - - - - s - - - w - w,sp,s,f - sp,s w,sp,s sp,s,f s s,f - - - - - f - - - - - - -- s,f - f w,sp,s - - - - sp,s s f sp,s,f - w w sp,s - - - w,sp sp,s - w,s,f - - w,s - - - - - - - - - sp,s sp w,sp - f sp w,sp,f sp w,sp,s,f Appendix .j:> .j:> 2. (Continued) Decapoda - Pleocyemata - Brachyura Cancridae Cancer irroratus Dorippidae Ethusa mascarone americana Dromiidae Dromidia antillensis Goneplacidae Glyptoplax smithi Grapsidae Euchirograpsus americanus Planes minutus Homolidae Homola barbata Homolidae A Leucosiidae Iliacantha intermedia Majidae Batrochonotus fragosus Collodes trispinosus Libinia dubia Libinia emarginata Macrocoeloma camptocerum Macrocoeloma trispinosum Metaporaphis calcaratta Mithrax acuticornis Mithrax forceps Mithrax pleuracanthus Pelia mutica Podochela gracilipes Podochela riisei Podochela sidneyi Stenocionops furcata coelata Stenorhynchus seticornis Palicidae Palicus faxoni Parthenopid~ Parthenope fraterculus Portunidae Ovalipes stephensoni Portunus gibbesii Portunus ordwayi Portunus spinicarpus Portunus ~in manus Xanthidae Carpoporus papulosus Glyptoxanthus erosus Lobopilumnus agassizii Pilumnus dasypodus MS02 ISOI IS02 MS06 w sp - - - - w w,sp,s,f sp,s,f w,sp,f w,sp,s,f sp,s,f - - - - - - - - f sp,s f - - - - - - - - w - - - - s s - s sp,s - - - s w w w,sp sp - w,sp,s,f w,sp w sp - OSOI - - w,sp sp,s,f w,sp - s w,sp,s,f sp - - s,f sp sp w,sp w,sp,s,f sp - - w,sp,s,f w w,sp - - sp,s sp w,sp w,sp,s,f - sp sp,s - - - w - - w,f w,sp,s,f sp,s,f - - - s - - - - sp,s w,s s s s s s sp - - sp,s sp - - - - - - sp - - - - - - w,sp,s,f w,sp,f w - - sp,s,f - f - - sp f sp,s s s w s Appendix 2. (Continued) Pi1umnus f10ridanus Pi1umnus pannosus Pi1umnus sayi Pseudomedaeus agassizii Xanthidae A IS01 IS02 MS02 MS06 OSOl s sp,s,f w,sp,s,f w,sp,s w,sp,s,f w,sp,s,f w,sp sp sp,s w,sp,s,f w,sp,s sp w,sp,f sp w,sp,s,f w,sp,f - sp,f w,sp,s,f sp - - Bryozoa Steno1aemata ~ VI Cyclostomata Crisiidae Crisia sp. Diaperoeciidae Diaperoecia f10ridana Lichenoporidae Lichenopora radiata Oncousoeciidae Proboscina f10ridana P1agioeciidae P1agioecia dispar w,sp,s,f w,sp,s,f w,sp,s,f w,sp - - w,sp,s w,sp,s,f w,sp,s,f - - - w,sp,f - - - sp - - - s,f s w,f w,sp,s sp,f w,sp,s,f f - w,sp - w,s sp sp sp w,s,f w,sp,s w w,sp w,sp,s,f Gymnolaemata Ctenostomata undetermined Ctenostomata Alcyonidiidae Alcyonidium sp. Buskiidae Buskia sp. Nollelidae Nolel1a stipata Valkeriidae Aeverrillia setigera Vesiculariidae Amathia alternata Amathia convoluta Amathia dis tans Victore11idae Sundane11a sibogae s,f w,sp,s,f sp,s,f Cheilostomata - Anasca Aeteidae Aetea anguina Aetea truncata w,sp,f w w,sp f s,f w,sp,s,f sp,s,f w s s,f w,s,f w,sp,s,f w,sp,s,f f sp w,sp,s,f sp,s,f sp,f sp sp,s,f w,sp,f w,sp,s,f sp w,sp,s,f w w,sp,s,f - - - - w,sp,s,f Appendix 2. (Continued) ISOI Alderinidae Alderina smitti Antropora tincta Copidozum tenuirostre Crassimarginatella leucocypha Parellisina curvirostris Parellisina latirostris Bugulidae Bugula fulva Bugula grayi Bugula marcusi Bugula microoecia Bugula rylandi Bugula sp. Bugula turrita Caulibugula dendrograpta Caulibugula pearsei - - sp,s,f w,sp,s,f - w,sp,s - s - - s,f sp f sp,f ~ 0- Farcimn~ Nellia tenella Hincksinidae Aplousina gigantea Aplousina tuberosa Membraniporid~ Membranipora savartii Membranipora tenuis Microporidae Micropora coriacea Onychocellidae Floridina antiqua Smittipora levinseni Scrupocellariidae Caberia boryi Scrupocellaria bertholleti Scrupocellaria regular is Steganoporellidae Steganoporella magnilabris w,sp - w,sp - sp w,sp,f - MS02 MS06 sp,f w,sp,s w,s,f sp,s w,sp,f w,sp w,sp,s,f - w w,sp,s,f f w,sp,s,f sp,f - w,sp,s w,sp,s,f sp,s,f - f - - - - s,f sp,s,f - w,sp,s,f sp sp w,sp w,sp,s,f f OSOI sp,s w sp,s,f w w w w sp,f Cel1ari ~ Cella ria nodosa Chaperiidae Chaperia sp. Cupuladriidae Cupuladria doma Discoporella umbellata Electrinidae Electra lacinosa Epistomiidae Synnotum aegyptiacum IS02 w,sp,s,f s sp,f - sp w,sp,s,f w w,sp w s,f - - - - - - w,sp,s - - f - - s,f s,f sp,s - - - w,sp,s,f sp,s,f - - - - w,sp - w,sp sp,f w - - - - sp f - - f w,s,f w f w,s,f - w - sp sp - - sp w,sp s w sp,s s s w,sp,s,f f sp w,sp,s,f Appendix 2. (Continued) ISOl Cheilostomata - Cribrimorpha Cribrilinidae Cribrilaria floridana Cribrilaria radiata Membraniporella aragoi ~ -..I Cheilostomata - Ascophora Adeonidae Reptadeonella hastingsae Celleporariidae Celleporaria albirostris Celleporaria magnifica Celleporaria mordax Celleporinidae Celleporina hassalli Turbicellepora dichotoma Cheiloporinidae Hippaliosina rostrigera Crepidacanthidae Crepidacantha Eoissoni teres Exochellidae Exochella longirostris Hippoporinidae Hippopleurifera mucronata Hippoporella uvulifera Hippoporidra janthina Hippoporina contracta Hippoporinidae A Hippothoidae Trypostega venusta Microporellidae Microporella ciliata Microporella marsupiata Microporella umbracula Petraliidae Petraliella bisinuata Petraliella marginata Phylactellidae Phylactella aviculifera Schizoporellidae Arthropoma cecili Cleidochasma porcellanum Cycloperiella rubra Schizoporella cornuta Schizoporella errata Schizoporella floridana Stylopoma informata IS02 MS02 MS06 w,sp,s,f w,sp,f - - - s s,f - OSOl s s,f - - s w,s w,sp,f w,sp,s,f s,f - w,sp,s,f w,sp,s,f - - w,s,f w,s sp,s,f w,s,f w,sp,s,f w,sp,s,f w,sp,s,f s,f w,sp,s,f w,sp,s,f w sp,s,f w,sp,s,f s w,sp,s,f w,sp,s,f w,sp,s,f sp,s w,sp,s,f w,sp,s,f w,sp,s w,sp,s w,sp,s,f w,sp,s sp,s - - - s,f s - - - sp s s,f w,s,f - s w,sp sp sp,f - w,sp,s,f - s sp,s,f s,f - sp,f - w,sp,s sp,s,f - w,sp,s,f f w,sp,s,f w w,sp,s w,sp,s,f w,sp,s,f w,sp,s,f sp,f w,sp,s,f sp,f w,sp,s,f w,sp,s,f w,sp,s,f w,sp,s,f s,f f - w,s,f - - - - - - - sp,f - - w,s,f sp - s w,sp,s,f - s,f - - w,sp,s,f - - - - sp - w,sp,s,f w,sp w,sp,s,f - - w,sp,s,f w,sp,f w,sp,s,f w f w,s,f sp,s,f w,sp,s,f sp w,f sp,s,f Appendix 2. (Continued) ISOI Sertellidae Rhyncozoon rostratum Rhyncozoon tubercula tum Smittinidae Parasmittina spathulata Smittina s m i t i e l ~ Stomachetosellidae Bracebridgia subsulcata IS02 - f MS02 f - - - s w,sp,f w, f w,sp,s,f - MS06 sp sp,f - w,sp,s,f w,sp,s,f f - - - w,sp,s,f w,sp,s,f w,sp,s,f OSOI sp,s,f Mollusr.a Gastropoda - Prosobranchia Archaeogastropoda Fissurellidae Diodora cayenensis Trochidae Calliostoma euglyptum Calliostoma pulchrum Calliostoma roseolum - w,sp,f s - - - sp - - - s "01,5 ''',sp - w,sp sp - - - - w - - - - s f sp ~ 0Cl Mesogastropoda Crepidu1idae Crepidula aculeata Crepidula forn:Lcata Crepi~ula plaaa Cymatiidae Cymatium labiosum Dis torsio constric ta macgi ntyi Cypraeidae Cypraea cervus Cypraea cinerea Cypraea spurca acicularis Eratoidae Erato maugeriae Trivia maltbiana Trivia nix - - Trivia pendiculus Ovulidae Simnia acicularis Simnia uniplicata Turritellidae Vermicularia knorrii Triphoridae Triphora nigrocincta Vermetidae Spiroglyphus irregular is - s - - - - - - f s - f - - w - w sp f - - sp s - s - s - s sp s,f - - s - sp - s,f - w,f - - - sp - - f Appendix 2. (Continued) IS01 Neogastropoda Bueeinidae Cantharus multangu1us Pisania tineta Co1umbel1id~ .". \0 Anaehis avara Anaehis 1afresnayi Mitre11a lunata Nassarina minor Suturog1ypta iontha Coral1iophi1idae-----Coral1iophila earibaea Faseio1ariidae Faseio1aria li1ium hunteria Faseio1aria tu1ipa Pleurop1oea gigantea Margine11idae Margine11a rose ida Murieidae Ca1otrophon ostrearum Murex f10rifer di1eetus Murex pomum Murex recurvirostris rubidus Thais haemastoma florida Urosa1pinx perrugata Nassariidae Nassarius a1bus Nassarius vibex ---- f sp - IS02 MS02 MS06 OSOl sp w,sp,s,f sp,s,f sp,f sp,f - sp,s s - - f s s - s sp - - f sp - sp - s - - sp - - - f - - sp - - - - - - s f - - - w - - - - - - w,sp - sp,s w w - s - s f f sp sp Gastropoda - Opisthobranehia Aplysiaeea Aplysiidae Aplysia wi11eoxi Ty10dinidae P1eurobranehaea hedgpeth Nudibranehia undetermined Nudibranehia Bornel1idae Bornella ea1earata Dendrodorididae Dendrodoris krebsii Dendrodoris warta Dorididae Anisodoris worki Hypse1odoris edentieu1ata Taringa aiviea s sp - - s - - - - w - - sp f sp Appendix 2. (Continued) 1501 Dotodae Doto doerga (?) - 1502 - M502 M506 0501 s Pelecypoda Arcoida Arcidae Anadara notabilis Arca imbricata Arca - -zebra -Barbatia candida Barbatia domingensis UI 0 Mytiloida Mytilidae Gregariella coralliophaga Lithophaga antillarum Lithophaga aristata Lithophaga bisulcata Modiolus americanus Musculus lateralis Pinnidae Pinna - - -carnea --Pterioida Anomiidae Anomia simplex Anomia squamula Pododesmus rudis Pectinidae Chlamys benedicti Lyropecten nodosus Plicutulidae Plicatula ~ib osa Pteriidae Pteria colymbus Ostreina Ostreidae Crassostrea virginica Lopha frons Ostrea equestris Ostrea - - -frons -Ostrea .Eermolis Veneroida Chamidae Chama congregata Chama macerophylla Pseudochama radians - - - w,s w w,sp,s,f - w,sp,s w - - - - - - - sp sp s - f f - sp,f sp s,f - sp sp,s w,f f sp sp sp sp sp w,s sp sp sp,s - - w - - s - - sp f sp sp - - - sp f w,sp,s,f w,f w,sp,s,f w,sp,s,f w,sp,s,f w,sp,s,f - - w w,sp,s,f - - s - - w s w w,sp,s,f - - - - w - w,sp w,s w w w,sp,s,f sp w s sp,s w w,sp sp s Appendix 2. (Continued) 1SOl Veneridae Chione grus Gouldia cerina Myoida Corbulidae Corbula chittyana Gastrochaenidae Gastrochaena hians Spengleria rostrata Hiatellidae Hiatella arctica Pholadomyoida Lyonsiidae Lyonsia beana 1S02 MS02 MS06 OSOl - - - - sp f - - - sp - sp - - sp sp sp - w,s s,f sp,s w,sp,s,f sp,s,f w,s,f - f Cephalopoda VI Sepioidea Sepiolidae Rossia sp. Teuthoidea Loliginidae Loligo pealei Loligo ~ undetermined Loliginidae Ommastrephidae 1llex illecebrosus Octopoda Octopodidae Octopus sp. w - w,sp,s,f s s w,sp,s,f sp sp sp w,sp,f w,sp - sp sp w sp,s w,sp,s,f w w,sp,s w,sp,s,f sp,s w,sp,s,f - - f - w - - S - Echinoderma ta Crinoidea Comasteridae Comactinia meridionalis Stelleroidea - Asteroidea Platyasterida Luidiidae Luidia alternata Luidia clathrata w,sp,s - sp,f sp - s Appendix 2. (Continued) 1S01 Paxillosida As tropec tinidae Astropecten articulatus Astropecten comptus Astropecten sp. Ul N Forcipulata Asteriidae Asterias forbesii MS02 MS06 OSOI - - - - w,s - - - - - - s - s,f 'lalvatida Ophidiasteridae Narcissia trigonaria Spinulosida Echinasteridae Echinaster brasiliensis Echinaster modestus Echinaster serpentarius Echinaster sp. 1S02 W - - - - - W W W sp,s,f sp,s,f w,sp sp - - - w w sp s - w,sp,s,f - - - sp,s,f sp,s w,sp sp,s W sp,s,f - sp w,sp,s,f sp,s,f w,sp,s,f w,sp,s,f Stelleroidea - Ophiuroidea Ophiuroidea A Ophiuroidea B Phyronophiurida - Euryalina Gorgonocephalidae Astrophyton muricatum Astroporpa annulata Ophiurida Amphiuridae Amphiodia pulchella Axiognathus squamatus Ophiostigma isacanthum Ophiactidae Ophiactis savignyi Ophiodermatidae Ophioderma brevispinum Ophio thricidae Ophiothrix angulata Oohiothrix suensonii - - - s sp,s s - - - - - sp sp,s - - w,sp sp,s w,sp,s w,sp,s,f w,sp,s w,sp,s,f - - - - w,sp,s w,s Appendix 2. (Continued) ISOI IS02 MS02 MS06 OSOI w,sp,s w,sp,f sp,s s,f s Echinoidea Cidaroida Cidaridae Eucidaris tribuloides Arbacoida Arbaciidae Arbacia punctulata Temnopleuro ida Temnopleuridae Genocidaris maculata Toxopneustidae Lytechinus var1egatus - - w,sp,s,f w,sp,s,f w,sp,s - - - s w,s,f w,sp,s,f - w,sp,s,f w,sp,s,f w,sp,s,f w w - Holothuroidea V. Dendrochirotida Cumariidae Ocnus pygmaeus IN Aspidochirotida Hole thuridae Holothuria dakarensis Holothuria princeps Holothuria surinamensis Synallac tidae Isostichopus badinotus - sp - - f sp - - w,f - s s w,sp,f w,f f f sp,s,f s,f s sp s w w,sp,s,f s w,sp,s,f w w w,sp,s,f w,sp,s,f w,sp,f sp - - Chordata Ascidiacea Ascidiacea A undetermined Ascidiacea Enterogona Ascidiidae Ascidia curvata Ascidia sp. A Clavelinidae Clavelina gigantea Clavelina picta--Clavelina sp. Clavelina sp. D Distaplia bermudensis Eudistoma carolinense - - w,sp,s.f sp,s,f w,sp,s,f sp w,sp,s w sp sp,f s w sp,s Appendix 2. (Continued) Euristoma olivaceum Euristoma sp. Didemnidae Didemnum candidum Didemnum sp. Didemnum speciosum Diplosoma macdonaldi Trididemnum savignii Trididemnum sp. Polyclinidae Aplidium bermudae Aplidium constellatum Aplidium exile Aplidium pellucidum Aplidium sp. A Aplidium sp. B Aplidium sp. D ~lid um stellatum U\ ~ Pleurogona Molgulidae Molgula occidentalis Pyuridae Pyura yittata Styelidae Botrylloides nigrum Botryllus planus Polyandrocarpa floridana Polyandrocarpa gravei Polyandrocarpa tincta Styela partita Styela plicata Symplegma vir ide ISOl IS02 MS02 w w,sp,s,f w,sp,s,f w,sp,s,f w,sp,s,f w,sp,s,f w,s,f w,sp,s,f sp w,sp,s,f - - sp,s w,sp,f sp sp,s w,sp,s,f sp sp,s,f w w w w,s - - - f w,sp,s,f - w,sp,s MS06 - sp,f s sp - w sp W,sp w - - - - w w w sp w w,sp,s,f w,sp,s,f w,sp,s,f w,sp,s,f w sp sp,f f s sp,f sp,s,f sp,s w,s,f w,sp,s.f w,sp,s,f f sp,f w,sp,s,f sp,s f w,sp,s,f w,sp,s,f sp,s,f - w,sp,s,f w,sp,s,f s,f w,s,f w,sp,s,f w s w,sp,s sp,s w,s,f w - - OSOl - sp sp w,sp,s,f sp,s w f sp w,sp,s,f - f Appendix 3. Ranked abundance of invertebrate species collected by suction and grab samplers at each station off South Carolina and Georgia during 1981. Average m2 , and ranked abundance for each season are indicated along with rank for all seasons combined. STATION OVERALL RANK 1.0 2.0 3.0 4.0 ~.o lJl lJl 6.0 7.0 8.0 9.0 10.0 11.0 12.0 13.0 14.0 15.0 16.0 17.0 18.0 19.0 20.0 21.5 21.5 23.0 24.0 25.0 26.5 26.5 28.0 29.0 30.5 30.5 32.0 33.0 34.0 35.5 35.5 38.0 38.0 38.0 40.0 41.0 42.0 43.5 43.5 45.0 46.0 47.5 47.5 50.0 50.0 50.0 52.5 52.5 :>4.5 54.5 SPECIES NAE~ FILOGRANA IMPLEXA SYLLIS'SPONGICOLA ERICHTHONIUS BRASILIENSIS AXIOTHELLA MUCOSA EXOGONE DISPAR LUCON~CIA INCERTA AMPHARETE AMERICANA PHOTIS SP. ELASMOPUS SP. A GAMMAROPSIS SP. ~APREL A EQUILIBRA ODOCERUS SP. LEUCOTHOE SPINICARPA RILDAROANUS LAMINOSA MICROOEUTOPUS MYERS I AMPELISCA SCHELLENBERG I OWENIA FUSIFORMIS CARPIAS BERMUDENSIS PRIONOSPIO CRISTATA NEMATODA ASPIDOSIPHON GOSNOLDI MEGALOBRACHIUM SORIATUM SIPUNCULIDA GOULDIA CERINA CHONE AMERICANA MALACOCEROS GLUTAEUS PARACERCEIS CAUDATA OPHIOTHRIX ANGULATA SYLLIS A~ILYH ERICHSONELLA FILIFORMlS PAGURUS CAROLINENSIS EUNICE VITUTA LUCIFER FAXONI AMPHIODIA PULCHELLA HYOROIDES CRUCIGERA MARGINELLA HARTLEYANUM CHAMA CONGREGATA LEPTOCHELIA SP. MELIT. APPENDICULATA BOTULA FUSCA HYDROIDES St->. A TURI3ELLARJA NEMERTINEA SPIOPHANES XYB~O LAEVICAROIUM PICTUM CHRYSOPETALIDAE A OPHIOSTIGMA ISACANTHUM PILUMNUS SP. LUMBRINERIS Il'\FLATA LYSIANOPSIS ALBA PRJONOSPIO CIRRIFERA LITHOPHAGA BISULCATA POLYCIRRUS CAROLINENSIS CERATONEREIS MIRABILIS PAGURIDEA wINT,EI1 ST ERR 3.2 6.6 41.0 3.6 16.8 22.6 28.0 24.8 1.4 9.0 16.8 13.6 1.4 6.2 14.2 4.2 8.4 0.6 12.0 4.8 4.4 1.4 7.b 1.4 4.4 0.4 1.8 1.4 1.8 0.2 0.4 1.4 · 5.0 1.6 1.8 1.2 3.0 1.0 1.6 1.4 2.8 3.4 4.6 0.2 2.0 1.4 U.2 1.8 • 4.0 0.2 1.6 0.6 · 2.9 5.6 n.3 2.0 6.2 11.5 9.5 11.3 0.9 3.6 8.2 4. 1 1.2 1.3 6.9 1.3 4.8 0.4 8.0 1.9 2.1 0.9 5.4 0.4 1.6 0.2 0.7 0.7 1.0 0.2 0.2 0.5 · 4.1 1.1 0.5 0.7 2.0 0.3 1.0 0.5 1.1 1.2 2.6 0.2 1.5 1.4 0.2 1.0 • 2.4 0.2 0.7 0.2 · (x) density, expressed as number per 0.10 1501 SPRING ST ERR MEAN · • 4.8 3.4 29.4 1t.6 4.6 0.2 0.8 20.0 11.2 2.8 1.0 7.4 5.8 3.2 1.1 17.6 1.9 2.4 0.2 0.4 11.2 8.0 1.6 0.0 6.2 3.0 1.2 3.2 7.6 0.7 2.2 2.8 2.6 1.0 4.4 0.2 1.4 1.8 0.2 2.0 2.4 1.8 3.0 1.2 0.6 0.2 0.6 0.6 2.0 0.8 1.8 5.0 0.8 1.0 1.2 0.4 3.0 0.2 0.7 1.2 0.2 0.8 1.2 1.0 1.3 1.0 0.4 0.2 0.6 0.2 0.7 0.6 1.0 4.3 0.5 0.4 · · • 0.2 0.4 1.4 0.8 0.8 1.6 0.6 0.6 · 1.2 0.2 0.2 0.4 · · • 0.2 0.4 0.7 0.6 0.5 0.9 0.4 0.2 · 1.0 0.2 0.2 0.2 SUMMER MEAN ST ERR 0.2 11t.8 It.O 11t.4 1.6 0.6 0.4 0.8 2.1t 0.1t 0.6 5.2 1.2 2.6 1.2 1.6 • 1.6 2.6 0.6 0.6 5.6 1.1t 2.8 1.4 0.2 0.8 · · 3.4 0.1t 2.6 6.6 0.2 1.0 2.0 1.0 0.8 • 0.2 0.8 0.8 1.2 0.2 1.0 1.2 1.4 0.2 1.2 2.6 0.2 0.4 0.2 0.4 • 0.2 14.3 2.6 4.8 0.6 0.4 0.2 0.5 0.8 0.2 • 0.2 5.2 0.2 · 0.5 0.6 1.1 • 0.7 0.9 0.6 0.2 0.1 0.4 1.5 0.7 0.2 0.4 1.5 0.4 2.6 4.1 0.2 0.4 0.8 0.5 0.4 • 0.2 0.5 0.4 0.5 0.2 0.5 0.6 0.5 0.2 0.6 1.4 0.2 0.2 0.2 0.4 · FALL ST ERR MEAN 182.4 55.0 1.0 1.6 H.2 3.6 • 1.6 2.2 0.4 • 0.6 3.0 0.4 3.0 0.4 121t.8 53.3 0.8 0.8 2.7 1.4 • 0.9 1.9 0.2 • 0.9 2.2 0.6 0.2 1.8 0.2 3.1 0.8 1.1 2.5 5.1 1.5 0.6 1.2 2.1 1.1 3.-3 1.5 0.2 2.8 0.8 • 0.1t 2.1 0.4 2.3 0.4 • 2.4 0.8 1.0 2.6 0.8 1.2 0.2 2.4 2.2 2.8 3.6 1.0 1.1 1.1 0.2 • 0.8 0.4 1.0 1.3 0.4 0.6 0.2 0.7 1.1 1.1 2.6 • 2.2 2.2 2.2 0.2 5.6 0.6 '3.4 1.0 3.8 4.2 5.8 3.6 1.2 1.8 5.4 3.6 3.6 3.2 0.6 5.2 2.4 • 3.2 2.4 1.8 0.4 • Appendix 3. (Continued) STATION OVERALL RANK VI 01 57.0 57.0 ";7.0 61.5 61.5 01.5 61.5 01 .5 61.5 65.0 66.0 67.5 07.5 fO.O 70.0 70.0 73.0 73.0 13.0 76.0 76.0 76.0 79.5 79.5 79.5 79.5 85.5 85.5 85.5 85.5 85.5 85.5 85.5 85.5 93.0 93.0 93.0 93.0 93.0 93.0 93.0 100.0 100.0 100.0 100.0 100.0 100.0 100.0 110.0 110.0 110.0 11 0.0 110.0 110.0 110.0 SPECIES AMP TTHOE SP. A PAGURUS HENDERSON I PITHO LHERMINIERI MEGALOMMA BIOCULATUM PELI A MUT I CA POTAMILLA 5P. PSEUOOMEDAEUS AGASSIZII SPIO SP. A SUTUROGLYPTA IONTHA ARMANDIA MACULATA EULALIA SANGUINEA PAGURIDAE SERPULIDAE CREPIDULA ACULEATA DIPLODONTA SEMIASPERA PODARt<E OBSCURA CRASSINELLA LUNULATA NOTOMA5TU5 SP. PAGURlJS SP. AMPELISCA VADORUM PAGURISTES SP. PAGURISTES TORTUGAE CERITHIUM ATRATUM PHYLLOCHAETUPTERUS 50CIALIS POTAMILLA RENIFORMIS S I PUNCULI OA A ACANTHOCHITONA PYGMAEA HYDROIDES UNCINATA LOIMIA MEDUSA MAJIDAE A MITHRAX PLEURACANTHU5 SICYONIA SP. VERMILIOPSIS ANNULATA WEBSTERINEREIS TRIDENTATA ARABELLA MUTANS ARCA IMBRICATA BRACHYURA GASTROCHAENA HIANS LEMBOS UNICORNIS RUOILEMBOIOES NAGLEI TANYSTYLUM CALICIROSTRE UIPLODONTA AT C~UP OPHIACTIS SAVIGNYI PARAPHOXUS SPINOSUS PILUMNUS SAYI POLYDORA WEBSTER I SYLLIS GkACILIS THELEPUS SET05US AMPHIPODA B ANCISTROSYLLI5 RIGIOA BOWMANIELLA PORTORICENSIS CAPITELL IDAE 0 CHIONE GI-lU5 CORBULA CONTRACTA HYBOSCOLEX ATESIG~OL WINTER MEAN ST ERR 0.6 1.0 1.6 0.8 0.2 0.2 · 0.4 0.4 1.2 0.6 0.2 0.2 ISOI SPRING SUMMER "'1EAN ST ERR MEAN 5T ERR 0.6 0.4 0.2 0.4 1.4 1.0 0.6 0.6 0.6 0.9 · 1.6 1.4 0.2 1.0 0.4 1.4 • 0.5 0.7 0.2 0.8 0.4 0.5 3.2 1.8 3.4 2.2 2.4 0.4 0.6 0.4 0.6 0.4 0.8 1.2 2.0 0.6 1.4 0.4 1.0 • 1.9 0.6 3.1 0.9 0.9 0.2 0.4 0.2 0.4 0.4 0.6 0.5 0.8 0.4 0.7 0.4 0.6 0.6 1.4 0.2 1.8 0.6 1.0 0.2 0.7 0.8 • 0.4 0.8 0.2 0.4 0.4 0.6 0.6 0.2 0.4 0.4 0.2 0.8 0.8 • • 0.6 0.6 0.2 0.4 1.6 0.8 • 0.2 0.4 0.9 0.6 0.2 · · • • 0.2 <l.2 · · · ·• • • 0.4 0.6 0.6 0.2 0.4 0.2 • 0.6 0.4 • 1.6 0.4 · 0.6 0.2 0.4 • 0.2 0.8 · · • 0.7 0.2 ·• 0.4 0.2 0.4 · 0.2 0.5 • • • 0.6 · 0.6 1.4 1.4 0.2 0.8 0.2 0.4 0.9 0.2 0.2 0.4 0.2 · 1.0 1.2 0.4 0.4 0.2 0.4 0.8 1.2 0.2 0.2 0.2 0.2 • • 0.8 0.2 0.6 0.2 0.2 0.6 0.6 0.2 0.6 0.2 • 0.6 0.6 0.2 0.6 1.0 0.2 0.2 0.4 0.5 0.6 • 0.6 1.2 ~.1 • • 0.4 · ·· • • ·• • 1.6 • ·· • · ·· · · 1.1 0.2 0.8 0.6 0.4 0.2 1.0 0.6 0.2 0.2 0.4 0.2 0.8 0.2 0.4 0.6 0.2 0.2 0.2 0.2 0.6 0.2 0.2 0.4 0.4 0.2 0.2 0.2 0.2 0.4 0.2 0.4 0.4 0.2 0.2 0.2 0.2 0.2 0.4 0.4 0.2 0.2 0.2 0.2 0.2 • • 0.2 • 0.8 0.4 • • 0.2 0.4 • • · • 0.4 0.2 • • 0.2 0.2 ·• • • 0.2 • 0.2 0.5 0.2 0.4 0.2 0.2 ·• • ·•• · 0.4 0.4 0.6 0.2 • 0.8 • O.~ • · • • · · FALL MEAN 5T ERR 1.2 1.4 1.6 0.6 1.4 3.2 1.0 • 0.4 • 0.8 0.6 0.7 0.5 0.4 1.4 2.5 0.3 0.2 · · • 0.5 • 1.2 1.4 2.0 0.4 0.8 1.2 0.6 0.2 0.8 0.6 0.6 1.0 1.6 0.2 0.4 0.8 1.0 0.2 0.4 0.6 1.2 0.4 1.6 1.4 0.8 0.2 0.6 0.4 1.2 0.5 0.4 0.2 · • o.e • • • 0.4 • • • 0.2 • • 0.4 0.6 0.2 1.0 0.2 1.0 1.0 0.4 0.6 1.4 1.0 0.6 0.4 0.4 0.5 1.2 0.6 0.4 0.4 0.7 0.6 0.2 0.5 0.2 0.4 o.e · • 0.2 0.4 0.2 0.2 0.8 0.6 0.4 0.4 0.6 0.5 0.6 0.4 0.4 Appendix 3. (Continued) STATlOr-. OVERAll RANK u. -.I 110.0 11 0.0 11 0.0 110.0 110.0 110.0 123.5 123.5 123.5 123.5 123.5 123.5 123.5 123.5 123.5 123.5 123.5 123.5 123.5 123.5 141.5 141.5 141.5 141.5 141.5 141.5 141.5 141.5 141.5 141.5 141.5 141.5 141.5 14.~ 141.5 141.5 141.5 141.5 141.5 141.5 141.5 141.5 166.5 166.5 166.5 166.5 166.5 166.5 166.5 166.5 166.5 166.5 166.5 166.5 166.5 wINTER MElIN S1 ERR SPECIES lEMBOS SMITHI MAERA wIlLIAMSI POL YDORA OHA STROMBIfORMIS BIlINEATUS SYLLIS CORNUH SYNALPHEUS TOwNSEND I AMPHIPODA E ANCHIALINA TYPICA CANCER IRRORATUS CAP ITELLIDAE CAULlERIELLA KILlARIENSIS CUMINGIA TELLINOIDES MAERA SP. A MANGELIA RUGIRIMA MICROPANOPE SP. PAGURUS HENDERSON I PHERUSA INFLATA SPIO PETTIBONEAE SYNALPHEUS SP. TURBONILLA HEMPHILLI ASPJDOSIPHON ALBUS AUTOl nus SP. BODOTR IIOAE A CAPITELLA CAPITATA CAR IDEA COLOMASTI X SP. INACHOIDES FORCEPS ISCHNOCHITONIDAE A lYSIANASSA SP. MARGINELLA AUREOCINCTA MARGINELLA ROSCIDA MEGAlUROPUS SP. MICROPHOLIS PACHYBACTERA MITRELLA LUNAT A MUREX fLORIFER DILECTUS MUSCULUS LATERALIS OOWS PYGMAElJS TELLINA AMERICANA THOR SP. TRICHOBRANCHUS GLACIAlIS VERMICUlARIA KNORRII XANTHIDAE AMPELISCA SP. C ARBACIA PUNCTUlATA BATRACHONOTUS FRAGOSUS CRATENA KAORUAE DODECACERIA CORALLII ERATO MAUGERIAE EUNICE ANTENNATA GLYCYMERIS PECTINATA HARMOTHOE SP. A HIATELLA ARCTICA HYPOCONCHA SABULOSA lATREUTES PARVUlUS LEPTOCHElA PAPULATA .. 1.2 0.2 0.6 1.2 0.8 0.2 0.6 0.6 0.2 0.2 0.7 0.4 0.2 0.4 1.2 0.2 1.2 0.2 0.2 0.4 0.2 0.4 0.2 1.0 0.6 0.2 0.6 0.2 0.2 0.2 0.2 0.4 0.2 0.5 0.2 0.4 0.2 0.4 · · o.t! · · ·• · · ·• · · · 0.2 0.6 0.8 0.8 · · • 0.4 0.2 0.6 • 0.4 0.8 ·· • ISOI SPHING ST ERR MEAN 0.2 ·· · ·· · · ··• · ··· · 0.6 0.6 0.6 0.2 0.2 ·· · 0.4 0.4 0.4 0.2 ·· · · ·· ··• ·· 0.2 0.2 0.2 1.0 0.2 0.8 1.2 0:6 0.2 0.8 0.2 0.2 0.8 0.2 1.0 0.6 1.0 0.4 • ··· ··· ·· · · · · · · · ··• ··• ·· · · · · · · 0.6 0.4 1.0 0.8 0.2 1.0 0.4 0.2 0.2 0.6 0.2 0.8 0.2 0.6 0.2 0.8 0.4 0.2 0.2 0.2 0.2 0.2 0.4 0.4 0.4 0.8 0.6 0.4 • 0.4 0.6 0.6 0.2 0.4 0.2 0.2 0.2 U.2 0.8 0.2 0.2 0.2 0.5 0.2 0.2 0.2 0.4 0.4 0.4 0.2 0.2 0.2 0.4 0.4 0.4 0.2 0.2 0.2 0.2 · • · · · SUMMER ME.AN 5T ERR · ·· · · · · ·· ··• · ·· · ··• ·· ·•• • ·• ··•• · ··• • MEAN · · ·• FAll ST ERR • · 1.2 1.2 • 0.8 0.2 0.2 0.2 0.4 1.0 0.4 0.4 0 ... 0.6 1.0 0.2 0 ... 1.0 0.2 0.6 0.2 0.8 0.4 0.2 0.4 0.8 0.2 0.6 0.8 0.2 0.5 0.2 1.0 0.2 0.2 0.2 1.0 0.2 0.2 0.2 0.2 0.2 0.2 0:2 0.2 0.4 0.2 0.4 0.2 0.2 1.0 0.4 0.2 0.8 1.0 0.2 0.5 0.6 0.4 0.2 0.2 • 0.2 ··• • · · ·• ·• · · · • ··· ·· ··· ·· · · ··· · ·· ·· 0.4 0.2 0.2 0.2 0.6 • 0.2 ·• 0.4 • ·• ··• · · • 0.4 0.8 · 1.0 0.4 0.8 0.2 0.2 • • ·• · 0.6 0.2 • • • 0.6 • 0.2 • 0.4 • 0.8 0.4 0.4 0.2 • • • • • · ·• · · 0.2 0.6 1.0 0.4 0.6 0.2 0.2 • • · · 0.4 0.4 0.2 0.6 0.0 0.4 0.2 0.2 0.2 0.2 Appendix 3. (Continued) STATION OvERALL RANK Ib6.5 166.5 Ib6.:' 166.S 166.5 Ib6.5 16b.5 166.5 16b.':) 166.5 166.5 Ib6.5 166.5 166.5 166.5 1~7.0 VI 00 197.0 197.0 197.0 197.0 197.0 197.0 197.0 197.0 197.0 1~7.0 197.0 197.0 197.0 197.0 197.0 197.0 197.0 197.0 197.0 197.0 197.0 197.0 197.0 197.0 197.0 197.0 197.0 197.0 197.0 197.0 197.0 197.0 244.5 244.5 244.5 244.5 244.5 244.5 244.5 SPECIE.S ~EDIOMASTUS CALIFORNIENSIS MICROP4NOPE SP. it SP. ~ICROPROT PUS OLIGOCHAETA OSTRACODA PHYLLODOCE SP. PINNIXA SP. PLEUROMERIS TRIDENTATA POLYCHERIA SP. POLYPLACOPHORA A PROCERAEA SP. PROCESS A SP. SABELLARIA VULGARIS 8EAUFORTENSIS TANYSTYLIDAE A XANTHIDAE A ALPHEUS NORMANNI AMERICARDIA MEDIA AMPHIPODA J ANOPLODACTYLUS PETIOLATUS CERATONEREIS SP. CHRYSOPETALIDAE B CORBULA BARRATTIANA DIASTYLIDAE A EULALIA MACROCEROS GONIADIDES CAROLINAE HYOROIOES SP. B LEMBOs UNIFASCIATUS REDUCTUS LITHOPHAGA ARISTATA LUCAPINELLA LIMATUL4 LYONSIA BEANA MAJIDAE MALOANIDAE MARGINELLA CARNEA MEGALOMMA SP. A MONOClJLODES SP. NEOPONTONIOES BEAUFORTENSIS PELECYPODA 8 PETALOPROCTUS SOCIALIS PISTA PALMATA PSEUOEURYTHOE AMBIGUA PSEUDOVERMILIA SP. A SABELLIDAE SERPULIDAE H SERPULIDAE K SICYONIA TYPICA SPIONIOAE 8 TIRON TROPAKIS VARICORBULA OPERCULATA ACANTHOHAUSTORIUS MILLSI AMPHIODIA TRYCHNA ANOPLODACTYLUS INSIGNIS APANTHURA MAGNIFICA AUSTRAEOLIS CATINA CADULUS QUAORIDENTATUS CHAETOPTERIOAE wINTER MEAN ST ERR 0.8 0.8 0.8 0.2 0.2 • 0.6 0.4 0.4 • 0.4 0.8 0.2 0.2 0.2 ·· · 0.2 0:4 0.4 0.2 0.2 ·· ·• ·· · · ··· · 0.2 0.2 0.4 • 0.2 0.2 0.2 0.2 0.2 0.4 0.2 0.4 0.2 • • 0.2 0.2 0.4 0.2 0.2 0.2 0.2 0.2 0.4 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.6 0.4 0.4 0.2 0.6 0.2 0.2 0.2 0.2 0.2 · 0.4 0.4 0.6 0.4 0.4 0.4 0.2 0.2 0.4 ·• 0.2 0.'+ 0.2 • 0.2 0.2 0.2 0.2 0.4 0.2 · ·•• · 0.4 0.2 0.2 0.2 0.4 0.2 0.2 0.2 0.2 0.2 0:4 0.2 0.2 0.2 0.4 SUMME.R MEAN Sf ERR 0.2 U.2 0.2 0.2 0.4 0.4 • 0.2 SPRINli ST ERR MEAN · · · ·· ·• ···• ·• ·• · 0.4 0.4 0.4 0.2 0.2 0.8 0.6 · ·· · ·• ·· ·· ·· · ISOl • 0.2 • · ·· ··· · · ···• · ··• · ·· ·· · · ··· ·· ·•• · ·• ·· ·• ··· ··· · 0.2 0.2 U.6 0:2 0.4 0.2 U.2 0.6 0.2 0.2 0.2 0.4 0.6 0.4 0.4 • • • · ·• ··• ·· ·· 0.4 ·• · · ·· ·· · 0.2 0.2 0.4 0.2 0.4 0.2 0.2 0.4 0.4 0.2 0.4 0.2 0.2 0.2 0:4 0.2 0.2 0.2 0:2 0.2 0.2 0.2 ·• ·• ··• · · ··· ··· · · • ·· ···• ·• • • • · ·· · ·• 0.4 0.2 0.2 0.2 FALL "lEAN ST ERR · 0.2 0.2 0.6 • • · 0.2 0.2 0.4 · ·• • • ·• 0.8 0.4 0.4 0.4 0.4 0.4 0.2 0.2 0.2 0.2 0.2 0.2 • O.b 0.2 0.4 0.4 • 0.4 0.2 0.4 0.4 0.4 0.4 0.6 0.2 0.2 0.2 0.2 0.4 0.2 0.2 ··• ·· ·• · · ·• • ·• · • • • 0.4 0.2 0.4 0.4 0.2 0.2 ·• 0.6 • 0.4 0.6 0.6 0.2 0.2 • • · · · 0.4 0.2 0.4 0.2 ·• 0.2 0.2 Appendix 3. (Continued) STAT ION OVERALL RANK 5.4 ~ VI \(;) 244.5 244.5 244.'5 ;:44.5 244.5 244.5 244.5 244.5 244.5 244.5 ;:44.5 244.5 244.5 244.5 244.5 244.5 244.5 244.5 244.5 244.5 244.5 244.5 244.5 244.5 ;:44.5 244.5 244.5 244.5 244.5 244.5 244.5 244.5 244.5 244.5 244.5 244.5 244.5 244.5 244.5 244.5 244.5 244.5 244.5 244.5 244.5 244.5 244.5 244.5 244.5 244.5 244.5 244.5 244.5 244.5 wINTER MEliN S1 ERR SPECIES CHIONE LATILIRA1A CRASSISPIRA ALBOMACULATA CREPIDULA PLANA CREPIDULA SP. OIODORA CAYlNENSIS OROMIDIA ANTILLENSIS EBALIA STI!Io\PSONI ENOEIS SPINOSA ERICHTHONIUS SP. A FAVARTIA ASOLU ~EC HEXAPANOP1U SP.· HOLOTHURO DEA HYALINA ALBOLINEATA INACHOIDES FORCEPS ~ LEMBOS SPINICARPUS INERMIS LUMBRINERIS COCCINEA LYONSIA SP. LYSIANASSIDAE A MARPHYSA SP. A MICROPHRYS ANTILLENSIS MITHRAX SP. NEREIDAE NEREIS SP. NICOMACHE TRISPINATA NOTOMASTUS HEMIPOOUS OLIVELLA MINUTA OPHIlJRO IDEA PERICLIMENES LONGICAUDATUS PERICLIMENES SP. PHERlISA EHLERSI POLYCHAETA POLYCIRRUS EXIMIUS PONTOGENIA SERICOMA PROCESSA 8ERMUDENSIS SABELLARIA VULGARIS SCOLOPLOS SP. B SEMELE NUCULOIDES SICYONIA BREVIROSTRIS SPIRORBIS SP. STENOTHOE GEORGIANA SYLLIDAE SYLLIDAE L SYLL IDAE N SYLLIS AL TERNATA SYLLIS FERRUGINA SYLLIS REGULATA CAROLINAE SYNCHELIDIUM AMERICANUM TERf-BELL 1O.4E TRITONIA BAYER I MISA TRYPANOSYLLIS SP. TRYPANOSYLLIS ZE8RA TURBO CASTANEA TURBONILLA INCISA WEBSTERINEREIS SP. A XANTHIDAE 8 ~ ·· · ·•• 0.4 U.4 0.4 0.2 0.4 • 0.2 • • 0.2 ··• ·• ·• • • 0.4 0.2 • • • • 0.4 0.2 0.2 • 0.2 • • 0.2 • ·• ·· IS01 SPRING ST ERR MEAN ·• ··• ·•• ·•• ·• ·· ·· ··• · ·•• ··• • 0.2 0.2 0.2 0.2 0.4 0.2 0.2 0.4 0.2 0.4 0.4 0.2 0.2 • 0.2 0.4 0.4 • 0.2 0.4 0.2 • ·• · ·· ·· ·• · · ·• 0.2 0 ... · • • • 0.2 · ·· ··• · • · ·• 0.2 0.4 0.4 0.4 0.4 0.4 0.4 0.2 0.4 0.2 0.4 · ·· ·•• ·· 0.2 0.2 • 0.4 · 0.2 0.2 0.4 • • • • • • • • 0.2 • • • • 0.2 • • • • · · · · ·• • • • • 0.2 • • • • • 0.2 • • · · · ·• ·•• ·•• ·• • · ··• • 0.2 0.2 0.2 0.2 0.2 • 0.2 • • · · 0.2 SUMMER MEAN ST ERR FALL ST ERR MEAN 0.4 0.2 0.2 0.2 0.2 0.4 0.4 0.2 0.2 • 0.2 0.2 • • • 0.2 • 0.2 • 0.2 0.2 0.4 0.4 0.2 0.2 0.2 • 0.2 • 0.2 ·• • • • 0.2 • • • · ·• ··• ·• ·• ·• ·• · ·· ··· · 0.4 0.2 · • ·• ··•• ··• • • ·• 0.2 0.2 0.2 0.2 0.2 0.4 0.4 0.2 • • • • 0.2 • • · • 0.4 0.2 • • • • 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 ··• · ·· ·• ·• • · 0.2 0.2 • • 0.2 0.2 • • • • • • • 0.4 0.4 0.2 • 0.4 0.4 0.2 0.4 • 0.4 0.4 0.4 0.4 0.4 0.2 • ·•• 0.2 • 0.4 • • • 0.2 0.2 • • • • 0.2 0.4 • 0.4 · • • • 0.2 • 0.4 • • • 0.2 0.2 • • • • 0.2 0.4 • 0.4 0.2 0.2 0.2 0.2 • 0.2 0.2 0.2 0.2 0.2 0.2 0.2 · 0.2 0.2 0.2 0.2 0.2 0.2 • • • 0.2 • • • • 0.2 • • Appendix 3. (Continued) STATION OVERALL RANK 351.0 351.0 351.0 3') 1.0 351.0 351.0 3':>1.0 351.0 3"1.0 3~1.0 351.0 3~1.0 0\ 0 3S1.u 351.0 351.0 3'51.0 351.0 351.0 3'51.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 3'51.0 3~ 1.0 31;;1.0 351.U 351.0 3'51.0 351.0 3" 1.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 3"1.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351. () 3~ 1. 0 351.0 ISOI WINTER SPECIES ST ERR N~EM ACROCIRRIDAE A ALPHEIDAE AMAEA RET IH.RA AMPELISCA CRISTOIDES AMPHARETE ACUTIFRONS AMPHILOCHlJS SP. AMPHIPODA G AMPHIPODA H ANACHIS AVARA A~ CHIS FLORIDANA ARCA SP. ARO ZEBkA ARCOPSIS ADMoiSI ASTERTAS FORBESII ASTHENOTHAEHUS HEMPHILLI AUTOMATE EVERMANNI AXIOGNATHUS SQUAMATUS BRANCHIOSYLLIS OCULATA BRANU CLAVATA CALOTROPHON ~URAE TSO CANTHARUS MULTANGULUS CER4DOCUS SP. B CERAPUS TU8ULARIS CERATONEREIS VERSIPEOATA CERODRILLI4 CERVINA CHRYSOPETALIOAE C CHRYSOPETALIDAE 0 CIROLANA PARVA CIRRIFORMIA FILIGERA CONUS JASPIDEUS STEARNS I COOPERELLA ATLANTICA COROPHIIDAE COROPHIUM SP. CREP IDULA SP. CRYOTURRIS ELATA DELECTOPECTEN SP. A DENDROCHIROTIOA DENORODORIS SP. A DIASTYLIDAE DIODORA LISTERI DONDICE UCCIDENTALIS DORIOPSILLA ALBOPUNCTATA EBALlA CARIOSA EUNICE WEBSTER I EUNICIDAE EUPHROSINE TRILOBA EURYDICE LITTORALIS EXOGONE SP. GASTROPTERON RUBRUM GLYCERA TESSELATA GONEPlAX SIGS8EI HETEROMYSIS FORMOSA HIPPOLYTIDAE JAEROPSIS CORALICOLA KURTZIELLA ATROSTYLA . 0.2 0.2 0:2 ~.O 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 ·• · · · ··· · · ·· • ·• ·• ·• ··• ·•• ·• · ·• ·• ··· • • • · · 0.2 0.2 0.2 0.2 0.2 0.2 ~:O 0.2 ··· · ·· · ·· · ·• ·· ··• ·· ·· U.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 • 0.2 0.2 NAE~ SPRING ST ERR • • 0.2 0.2 0.2 0.2 ··• • • ··• ·• ··• ·• ·•• ··• ··· ·• 0.2 0.2 ·· · ··· · ··· · ·•• · ··· ·· ·• ·• ··· ·• ·•• · ··• ·· ·•• • · 0.2 0.2 0.2 0.2 0.2 0.2 • • 0.2 ·• ··• ··•• • ·• ·• 0.2 0.2 0.2 0.2 0.2 0.2 0.2 • 0.2 0.2 SUMMEH MEAN ST ERR ··• ··· · ··•• • ··• ·· · ·· ···• • • ·• • • ··•• ·• 0.2 0.2 0.2 0.2 • 0.2 • • • 0.2 • • • • • 0.2 0.2 0.2 0.2 0.2 • • • • · · ··•• • • · ·• ·· ·· · ·• ··• ·•• ·•• · ··• ·· ···• ·• ·· • 0.2 0.2 0.2 0.2 0.2 • ·• FALL MEAN ST ERR ·• ·• 0.2 • • • • • 0.2 ·· ·• ·••• · ·• 0.2 0.2 • • 0.2 • • • • • • 0.2 • • • 0.2 • • • • • • 0.2 • • • 0.2 • 0.2 0.2 0.2 · • • 0.2 • 0.2 0.2 • 0.2 0.2 • 0.2 0.2 • • • 0.2 • • • • 0.2 • • • • • • 0.2 • • 0.2 · · • • • 0.2 • • 0.2 • • • • • 0.2 • • • Appendix 3. (Continued) STAT lOr-. OVERALL HANK 0'1 ..... 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 SPECIES KURTZIELLA LIMONITELLA LAEVICAROIUM SP. LAMELlARIA PERSPICUA lATREUTES FUCORUM • LEM80S SP. LjPTOCHELA SP. L LJE80RGIA SP. B LINGA AMIANTUS LUMBRINERIS SP. LYSIDTCE NINETTA MACROCOELOMA SP. MACROCOELOMA CAMPTOCERUM MARGINELLA LAVALLEEANA MARGINELLA SP. MARPHYSA SP. 8 MEGALOMMA SP. MESOCHAETOPT~RUS SP. MICROPHRYS SP. MICROPHRYS SP • • MITHRAX ACUTICORNIS MITHRAX SP. • MUNIDA SP. MUREX RECURVIROSTRIS RUHIDUS MYSIOACEA MYSIDOPSIS FURCA NASSAR IUS ALBUS NASSARIUS VIBEX NEMERTINEA C NEMERTINEA E NEOORILLIA CYOIA NEPHTYIDAE NEPHTYS SQUAt-lOSA NEREIS SUCCIr-.EA NUCULA PROXIMA NYMPHOPSIS DUODORSOSPINOSA OCTOPUS SP. OOONTOSYLLIS SP. ODONTOSYLLIS ENOPLA OLIVA SP. OLTVELLA BULLULA ONUPHIDAE ONUPHIS SP. OPHIOCNIDA SCABRIUSCULA OPHIOLEPIS ELEGANS OXYUROSTYLIS SMITHI I-'ENAEOIDEA PERICLIMENAEUS SP. A PHERUSA SP. PHTISICA MARINA PHYLLODOCE FRAGILIS PHYLLODOCE LOr-.GIPES PHYLLODOCIDAE PILUMNUS DASYPODUS PILUMNUS FLORIDANUS PILUMNUS PANNOSUS " INTER MUN ST ERR • ~.o ·• ·• ··• ·· ·· ·· ··•• ··• ·· · ·· ·· ··• ·• · ·• · ·• · ·• · 0.2 0.2 0.2 ·• ··• • • • ·• ·· ·• ··· · 0.2 O.~ 0.2 0.2 • • • ··• ·• ·•• ·• ··• · ··• ·· ·• ·· 0:2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 1501 SPRING ST ERR MEAN · ·• · ·· ·•• ·• · ·· · ·· ·• ·· · · ·· ··• · ·· • ·· ···• ·· ·· ·· ·•• ·· ·• 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 • 0.2 ·· ·· · · ·· · ·•• ··• ·· ·• ·· ··· ·• ·• 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 SUMMER ME-AN ST ERR 0.2 0.2 ·· · · ··· ·· ··· ·• ··• · ·•• O.~ O.~ O.~ 0.2 • • 0.2 • • ·· · ·•• 0.2 O.~ 0.2 0.2 0.2 0.2 • 0.2 · · 0.2 · 0.2 0.2 0.2 · · 0.2 0.2 0.2 0.2 ·• 0.2 · 0.2 0.2 • • 0.2 • · · · 0.2 • • • • ·· ··• ·• ·• ·•• • ·· ···• 0:2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 ··· • ···• ··• ··· ··• FALL ST ERR 0.2 0.2 0.2 ·· ·• ·· MEAN • • • 0.2 0.2 • 0.2 0.2 · ·• · · • 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 • • ·· • · 0.2 0.2 0.2 • 0.2 0.2 0.2 • • • Appendix 3. (Continued) STATION OVERALL RANK C'I tv 3':>1.0 351.0 351.0 351.0 3~ 1.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 351.0 ST ERR N~E'" PINNIXA CYLINDRICA ~IN OTHE~ S SP. PIONOSYLLIS SP. A PISANIA TINCTA PLICATULA GIBBOSA PODOCHELA SP. ~OLIN CES SP. PORCELLANIDAE PORTUNIDAE PROCERAEA FASCIATA PROCESSA HEMPHILLI PROCESSA VIC INA ~ROTULA TUBULARIA PSEUDOVERMILIA OCCIDENTALIS PTERIA COLYMBUS SABELLARIA <.;RACILLIS SABELL.RIA VULGARIS VULGARIS SABELLARIIDAE SCALI Mj(ER~ IOAE SCOLOPLOS SP. A SElLA ADAMSI SEMELE BELLASTRIATA SEMELE PURPURASCENS SERPULIDAE A SICYONIA L.AEVIGATA 5 I PUNCULI DA C SIRAIUS BICULOR SPELOEOPHORUS ELEVATUS STENOTHOIDAE STENOTHOIDAE A S't'LLIDAE H TELL INA AEQUISTRIATA TELL INA 5P. THARYX MARIONI THYONE PSEUOOFUSUS TOZEUMA SP. TRITONIA BAYERI TURBONILLA ABRUPTA TURBONILLA INTERRUPTA WILLI.MIA ISeE~K XANTHIOAE it SPRING WINTER SPECIES • · · 0." ·· · ·· ·· ·· · ··· ·· O.C' ·· · ·· • · ·• 0.2 0.2 0.2 0.2 0.2 0.2 0 • .2 0:2 0.2 0.2 ~.o 0.2 • • 0.2 0.02 0.2 0.2 • • ·• ·· · 0.2 I5Ul 0.2 0.2 • • · · ··•• ·• 0.2 0.2 0.2 0.2 0.2 0.2 • ·· ··• · 0.2 SUMMER MEAN ST ERR 0.2 0.2 ·· 0.2 ·· ·· ·· ·•• ·• ·• · ··• ·• · · ···• • · 0.2 0.2 0.2 ·• ·· ··· ·· ··• ·· ·• ·· ·· ·• ·• ··· ·· · 0.2 0.2 0:2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 ME.AN FALL ST ERR · ·• ·· ··• ·• · 0.2 0.02 0.2 0.2 • 0.2 • 0.2 · · 0.2 o.£! • 0.2 • 0.2 0:2 • • · ··· · 0.2 • • 0.2 · ··· ··· ·• ·· MEAN 5T ERR • • • • 0.2 • • • • 0.2 0.2 · · · ·· · ··• · ·• 0.2 0.2 0.2 0.2 ·• 0.2 • 0.2 • 0.2 • 0.2 · ·•• · · ·• · 0.2 O.~ 0.2 0.2 0.2 • • • • • 0.2 · · 0.2 0.2 • • · 0.2 0.2 Appendix 3. (Continued) STATION UVERALL RANK 1.0 2.0 3.0 4.0 ~.o 6.0 7.0 8.0 q.O 01 w 10.0 11 .0 12.0 13.0 14.5 14.5 16.0 17.0 18.0 19.0 20.0 21.0 22.0 24.0 24.0 24.0 26.0 27 .'J 27.5 29.0 30.5 30.5 32.0 33.0 34.5 34.$ 30.5 3h.~ 38.0 39.0 40.0 42.0 42.0 42.0 44.5 44.5 46.5 46.':; 48.0 50.5 ~0.5 50.$ 50.5 53.0 54.5 54.5 SPECIE.S LUCONACIA INCERTA ELASMOPUS .~S A ERICHTHONIUS BRASILIENSIS LEM80S SM IT HI CAPRELLA EOUILIBRA PODOCERUS SP. PHOTIS SP. LEPTOCHELIA SP. LUM8RINERIS INFLATA POLYCIRRUS CAROLINENSIS EXOGONE DIStJAR GAMMAROPSIS SP. SPIO SP. A AMPITHOE SP. ,\\ MEGALOBRACHIU~ SORIATUM PSEUDOMEDAEUS AGASSIZII CARPIAS BERMUOENSIS ARCA IMBRICATA SYLLlS GRAC IllS MICRODEUTOPUS MYERS I OXYUROSTYLIS SMITHI MAERA SP. A PILUMNUS SP. S I PUNCULI DA A SYlLlS HYALlNA OPHIACTIS SAVIGNYI ASPIDOSIPHON GOSNOLOI OPHIOTHRIX ANGULATA MALACOCEROS GLUTAEUS NEMATODA POOARKE OBSCURA CHAMA CONGREGATA SYNALPHEUS TOwNSEND I ASTERIAS FORBESII SYLLIS SPONGICOLA APSEUOES SP. A CREPIDULA ACULEATA AMPELISCA SCHELLENBERG I EULALIA SANGUINEA PARACERCEIS CAUDATA MELITA APPENOICULATA PISTA QUAORILOBATA XANTHIDAE A ARMANDIA MACULATA TURBONILLA INTERRUPTA AMPHARETE AMERICANA JASSA FALCA TA PRIONOSPIO CIRRIFERA AXIOGNATHUS SQUAMATUS GOULDIA CERINA LEMBOS UNICORNIS TURAONILU I"'CISA CHRYSOPETALID,\\E A AMPHIODIA PULCHELLA LYSIANOPSIS ALBA wINTER IS02 SUMMER SPRING FALL ME,\\N ST ERR MEAN ST ERR MEAN ST ERR MEAN ST ERR 68.0 14.0 27.0 15.2 29.8 26.4 20.4 17.2 8.0 5.6 6.6 0.2 4.4 3.2 7.2 4.2 0.6 2.6 6.4 10.0 0.2 4.0 4.2 4.8 3.0 6.2 1.4 4.4 0.2 6.2 0.4 3.2 1.4 19.5 4.5 11.3 7.8 9.3 7.8 5.0 7.3 3.6 1.7 2.6 0.2 4.4 1.1 2.1 1.7 0.4 1.0 1.6 1.9 0.2 1.9 1.3 1.5 0.5 1.5 0.7 1.4 0.2 4.7 0.2 1.2 1.2 0.2 4.7 0.4 5.0 0.4 5.2 0.4 2.2 0.4 2.6 1.2 1.2 1.9 1.1 0.6 0.6 1.0 0.8 5.4 3.2 4.0 5.2 4.0 1.2 0.8 1.2 4.6 0.4 0.6 0.6 3.9 2.9 1.0 2.3 1.6 0.7 0.8 1.2 4.3 0.2 0.4 7.2 7.1 2.9 0.9 1.3 4.9 4.6 3.4 4.2 3.9 10.0 10.8 3.4 0.9 0.9 5.3 1.3 0.7 0.4 1.0 1.3 1.3 1.8 1.0 0.7 2.0 1.3 0.2 1.0 1.7 0.7 0.4 1.3 0.7 1.9 1.1 1.0 0.7 0.2 3.3 1.4 1.2 0.2 12.2 1.4 2.8 3.4 3.8 1.0 1.8 2.0 13.8 16.8 6.8 3.2 4.0 10.8 7.8 8.6 8.4 6.0 16.0 11.0 6.2 2.0 1.6 9.2 4.2 1.8 0.4 1.6 4.4 3.0 2.8 2.2 2.4 5.8 3.8 0.4 2.4 4.8 1.6 0.6 7.8 1.2 3.4 1.6 1.4 2.0 0.4 4.0 2.8 1.2 0.8 0.4 · • · · · · · · 1.2 1.2 • 0.6 1.0 0.4 3.0 0.8 2.2 0.4 2.5 0.5 1.5 • 2.6 2.8 0.8 · · 1.7 2.8 0.4 • 8.6 ·• 0.8 0.4 3.0 1.8 1.2 • • • 5.6 • 0.6 • 0.4 1.0 0.6 0.6 ·· 2.2 4.4 7.0 2.4 5.8 3.2 1.3 2.5 2.0 2.0 3.0 2.4 1.8 1.1 1.1 0.6 • • • 2.4 0.8 1.8 0.2 3.4 3.2 4.0 • 3.6 1.2 2.6 1.8 1.6 0.8 1.2 1.4 3.2 ·• 1.0 • 0.4 • 3.0 1.0 • 0.8 · 1.6 4.~ 1.3 2.1 1.3 • • • 0.9 0.5 0.9 g.2 .9 1.4 1.9 • 2.4 0.5 1.0 0.8 0.7 0.4 0.7 0.7 1.5 • 1.0 • • 0.4 • 2.0 0.4 • 0.6 • 0.7 • 0.8 0.4 • 0.4 0.2 • 2.2 1.0 2.8 • 0.9 0.5 1.4 2.4 0.4 0.8 1.2 0.6 0.8 1.4 9.6 0.6 1.6 0.2 3.2 1.6 1.1 0.2 0.6 0.4 0.6 0.5 0.5 0.7 0.2 0.7 0.2 1.1 1.0 • 1.6 0.2 • 1.0 0.2 • • • 0.6 7.0 0.2 0.4 0.4 2.0 • • • 0.4 2.5 0.2 0.4 0.4 1.0 • 0.4 1.0 3.0 0.4 0.4 0.4 • 0.4 0.6 2.5 0.4 o.~ o. • • • 0.2 • • • • 0.4 • 0.2 0.4 • 0.4 1.2 0.8 • 0.4 0.4 0.6 • 0.2 • 0.2 Appendix 3. (Continued) STATION OvERALL KANK :'7.U 'j -, • 0 ')7.U bO.:' bO.S 6U.5 60.~ 6'+.0 64.0 04.0 b7.0 67.0 67.0 6~.5 09.5 0'1 .j::o. 72.5 72.5 12.5 72.5 7'3.5 78.5 78.5 78.5 78.5 78.5 78.5 78.5 ti5.0 85.0 d').O >i5.0 d5.0 90.5 ~0.5 90.5 90.5 90.5 90.5 98.5 98.5 98.5 98.5 98.5 98.5 98.5 98.5 98.5 98.5 106.0 lu6.0 106.0 106.0 lU6.0 113.5 113.5 ANOPLODACTYLUS PETIOLATUS MITRELLA LUNATA PHERLJSA EHLERSI 8RACHYURA ERICHTHONIUS SP. A LEUCOTHOE SPINICARPA SYNALPHEUS sr. MEGALOMMA BIOCULATUM PILUMNUS FLOkIOANUS SYLLIS SP. A AMPHIPODA E FILOGRANA IMPLEXA SERPULIDAE APANTHURA ACIF NGA~ PHYLLODOCE CASTANEA A,",PELISCA SP. C LYSJANASSA SP. PHERUSA INFLATA SYLLIS ALTERNATA CERATONEREIS MIRABILIS JAEROPSIS CORALICOLA LATREUTES PARVULUS MEGALUROPlIS SP. NEMERTINEIl PAGURUS SP. POLYDORA TETRABRANCHIA IlIE8STERINEREIS TRIOENTATA ALPHEUS IN A~RON CAPRELLA PENANTIS GONJAOIDES CAROLINAE PILUMNUS DASYPODUS THARYX MARIONI BHANCHIOSYLLIS EXILIS DIPLODONTA PUNCTATA GA~MARlJS SP. LOIMIA MEDUSA PAGURUS SI ~ENILORAC PRIONOSPIO CRISTATA AXIOTHELLA MUCOSA CERADOCUS SP. HETEROHYSIS A50M~OF HEXAPANOPEUS SP • MAERA CAROLINIANA PEllA MUTICA PISANIA TINCH PSElJOOVERMJlIA OCCIDENTALIS THOR SP. TURBELLARJA AUTOL nus SP. CRASSISPIRA ALBOMACULATA DASYbRANCHUS gP. SICYONIA LAEVIGATA SPIOPHANES BOM8YX BRANCHIOSYLLIS OCULATA INACHOIDE.S FORCEPS . SUMMER SPRING WINTER SPECIES IS02 MEAN ST ERR MEAN ST ERR MEAN 1.t! 3.0 1.4 2.13 0.2 0.2 0.2 0.1+ 2.6 0.7 1.7 0.5 1.8 0.2 0.2 0.2 0.2 1.1 1.8 0.6 1.0 0.4 1.2 0.9 0.4 0.8 0.4 1.0 O.ti 1.6 0.4 0.8 2.0 0.2 2.2 0.1:3 1.2 0.7 0.2 0.4 1.3 0.2 1.3 0.6 0.4 · 0.4 0.4 0.2 1.2 0.2 • 0.8 0.4 1.2 ·• · • 0.4 0.4 0.2 0.8 0.2 · ·· · 0.8 0.2 1.0 0.2 0.2 0.8 1.4 0.4 1.0 • 0.2 0.6 1.0 · · · · · · · • 0.2 0.4 0.5 · · · · ·• · O.b 1.4 0.9 2.2 0.4 0.7 0.4 0.4 0.6 0.4 0.2 0.2 1.8 1.8 0.4 0.4 0.2 0.2 0.4 O.b 0.6 0.2 1•~ 0.8 0.4 0.2 0.4 0.2 0.4 0.2 0.2 0.5 0.6 0.2 0.2 2.0 2.2 o.~ ·· • 1.6 1.0 ·· ·· · 0.9 0.6 • 1.6 1.6 0.2 0.6 0.2 2.2 1.0 0.6 1.2 0.8 0.2 1:0 0.9 0.2 0.4 0.2 1.6 0.4 0.6 0.6 0.8 0.2 0:2 0.6 2.0 0.8 0.6 0.6 0.2 0.2 0.4 1.8 0.6 0.4 0.4 0.2 • 1.8 · · · 1.3 ·· MEAN ST ERR · • 0.4 • 0.4 0.6 ·· · O.b 0.6 0.2 2.4 2.4 0.2 2.1t 0.2 1.4 0.7 0.2 0.8 1.2 0.2 0.6 0.6 0.4 2.2 1.0 0.4 0.2 1.0 0.5 0.2 · ··• • • 0.2 2.0 2.4 3.2 0.6 0.6 0.6 0.4 0.8 1.1t 1.2 0.2 1.0 0.6 1.2 0.8 0.2 0.2 0.8 0.7 1.2 0.2 0.4 0.6 0.6 0.8 0.2 0.1t 0.2 1.0 0.2 0.4 0.2 0.4 0.2 0.4 0.4 1.0 0.1t 0.2 0.8 • • • 0.1t 1.4 0.2 0.8 1.4 1.2 0.2 0.8 1.4 • 0.8 • 0.6 0.8 0.2 • 0.4 • 0.4 0.5 0.2 0.4 • 0.4 0.6 • 0.4 0.2 0.8 0.4 0.4 0.4 1.2 0.2 0.6 0.2 0.2 0.2 1.0 0.6 0.2 0.2 • 0.4 0.2 0.2 o.c 0.4 ·· • ·• · ·· ··· ·• · • 0.8 0.9 0.7 0.5 0.2 0.4 0.4 0.2 0.2 0.2 0.2 0.4 0.2 0.4 0.2 0.4 · · ST ERR • 0.5 0.2 2.0 1.5 1.6 0.4 0.8 1.4 1.2 1.0 0.4 0.6 0.4 0.2 0.2 · · FALL · · ·• · ·• ·•• · · • 0.4 · ·· · · ·•• • • 0.2 0.8 ·· ·•• • 0.2 0.4 0.8 1.2 0.2 • ·• 0.4 0.2 1.0 · ·•• • • 0.2 0.4 0.8 • • • 0.2 0.4 0.4 0.5 0.2 Appendix 3. (Continued) STATION OVEriALL RANK SPECIES WINTER MEliN 113.5 113."> 113.5 113.5 113.5 113.5 113.5 0Vl 113.5 127 .5 121.5 127.5 127 .5 127.5 127.5 127.5 127.5 127.5 127.5 127.5 127.5 127.5 127.5 J27.5 127.5 127.5 127 .5 143.0 143.0 143.0 143.0 143.0 143.0 143.0 143.0 143.0 143.0 143.0 143.0 143.0 158.0 158.0 158.0 l~tl.O 1')tl.0 1':18.0 1~8.0 1':>8.0 15R.O 1,8.0 1513.0 1';8.0 158.0 1511.0 158.0 158.0 MARGINELLA LIIVALLEEANA PHYLLODOCE FHAGILIS PISTA PAU1ATA POLYCH£RIA SP. POLYCIRRU5 EXIMIUS POLYDORA CAi:.CA SABELLMlI A VULGARIS SI NETHOFUA~B SYLLIS R£GULATA CAROLINAE AMPELISCA ~tURODAV ARABELLA MUTAr-.S CHAMA MACEROPHYLLA DIODORA SI NE~ YAC DODECACERIA CORALLII EUNICE FILAMENTOSA HYDROIDES SP. B LITHOPHAGA 81SULCATA MARPHY5A SP. A O~ENIA FUSIFORMIS PAGURI5TES TORTUGAE PAGURUS HENOEKSONI PILUMNU5 SUC~NAP PSEUOOVERMILIA SP. A 5"'10 PETTIBONEAE SPIONIDAE TELLINA AMERICANA TRYPANOSYLLIS ZEBRA AMPHARETE ACUTIFRONS HOLOTHUROIDEA LUCIFER FAXONI LUMBRINERIS COCCINEA MEDIOMASTUS CALIFORNIENSIS MELPHIDIPPIOAE A NYMPHONIOAE A PAGURISTES SP. RILDARDANUS ASONI~AL SUTUROGLYPTA IONTHA THY ONE PSEUOOFUSUS VERMICULARIA KNORRII XANTHIDAE ANOPLODACTYLUS INSIGNIS AHBACI~ PUNCTULATA CHIONE GRUS CHRYSOPETALIDAE 8 CUMINGIA TELLINOIOES ENGINA TURBINELLA EUNICE VITTATA GLYCERA PAPILLOSA GNATHIA SP. A MALDANIDAE PILUMNUS SAYI PROCtSSA SP. SABELLARIA FLORIDENSIS SABELLARIA VULGARIS SABELLAPIA VULGARIS VULGARIS SEBA SP. 1502 '5T SPfolING ~RE 0.4 0.6 0.2 0.4 1.4 0.8 0.4 0.4 1.2 0.6 0.2 0.4 1.0 0.4 1.0 0.6 O.CJ · · o.e · O.d · ·• 0.6 0.4 0.2 0.2 SUMMEfol MEAN ST ERR MEAN ST ERR 0.2 0.2 0.8 0.5 o.e ·· ·• · ·· ·· · 0.4 0.2 • 0.2 0.2 0.4 0.2 0.6 0.4 0.2 0.2 0.4 0.2 0.2 0.2 0.4 0.4 0.2 0.2 0.2 0.2 0.2 0.6 0.4 0.2 0.4 0.4 • 0.4 • 0.4 0.8 0.6 0.4 O.b 0.4 0.4 0.4 0.4 0.6 0.6 0.4 0.2 0.4 0.6 0.4 0.4 0.2 0.4 1.0 0.2 0.6 O.t! 0.2 0.2 0.6 0.4 0.2 0.4 0.2 0.2 0.6 (l.2 0.2 0.2 0.6 0.2 0.2 0.2 0.2 0.4 0.2 0.2 0.2 0.2 1.0 0.2 0.2 0.8 O.d 0.6 0.6 0.6 0.2 0.6 0.2 0.2 0.2 O.I::! 0.8 0.2 0.2 0.2 0.2 0.2 0.8 0.6 0.2 0.2 0.5 0.2 0.2 0.2 0.2 0.2 0.4 0.2 ·· · ·· · · · ·· ··· 0.4 0.6 O.b · ·· · ·· • · · ·· ··· · 0.2 0 ... 0.4 FALL • · ·· · · ·· 0.2 0.4 0.2 0.4 · 0.8 ·· O.C · · ·· · · 0:4 0.8 0.4 0.2 0.4 0.4 0:2 0.4 0.2 0.2 0.6 0.6 0.4 0.6 0.2 0.6 0.2 0.6 0.4 0.4 0.8 0.2 0.8 0.4 0.6 0.2 0.8 0.4 ·· 0.4 • 0.6 0:2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.6 0.4 0.4 0.6 0.4 0.2 0.8 0.2 0.6 · · · (J.b · · 0.2 0.8 0.4 0.2 0.2 · · ·· · · • · ·· · ·• · • • 0.2 0.8 0.8 0.4 0.4 0.2 o.~ · ·• · ·· · · · 0:2 · ·· ·· · · • 0.8 · · · o.~ 0.4 ··· · MEAN · · · ·• · · 0.6 0.2 ST ERR • 0.4 0.2 0.2 • 0.2 0.2 1.4 0.2 0.2 0.2 0.4 0.2 0.2 0.2 0.2 0.2 0.2 0:2 0.8 0.6 o.e 1.0 · 1.0 0.8 0.6 o.e 0.2 · 0.2 0.2 · 0.2 · • · • • ·• • 0.2 0.4 ·• 0.4 · · · · 0.2 0.4 0.2 0.4 0.2 0.2 · 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 · · 0.2 • • 0.2 Appendix 3. (Continued) STATION OvERALL RA/'IIK l':>d.O li-l3.5 ItJ3.S IdJ.5 1"3.5 183.5 183.5 Hn.~ ltD.':) lfU.~ 0\ 0\ 183.5 11'33 .5 Id3.5 183.5 IM3.5 IH3.5 1>j).':) l1i3.5 183.5 11'\3.5 1(j3.5 183.':> 183.5 11j3.5 1fl3.5 liD.S lii3.5 1113.5 1113.5 183.5 183.5 183.':) 183.5 183.5 lti3.5 220.5 220.5 220.5 220.5 220.5 220.5 220.5 £'20.5 220.5 220.5 220.5 220.5 220.':> t20.5 220.5 220.':) 220.5 220.5 220.5 220.5 5PFCH_5 XMHHIDAE [) ANCHIALINA TYPICA ANOPL00ACTYLUS LENTUS 8RANIA ~TAv LC CI\PRELL ~\1)lJ A CARIN08ATEA CARINATA CERAPUS TUauLARIS CERITHIOPSIS EMERSON I CEPITHIOPSIS ~RE NI CERTTHIUM ATRATUM COLOMAS TIX ':>P. COROPHI WAE ERATO EAI~EGUAM EUNICE ANTENNllfll FAVARTIA CELLULOSA GTHNOPSIS SP. GLYCERA TESSELATA HIPPOLVTIDAE HVROSCOLEX LONGISfTA LAEVICARDIUM PICTUM LEMBOS SP. LUM8kINE~IS LATkEILLI MICROPANOf'E SP. NEMERTINEA E OPHIOSTIGMA SP. A PAGURIDAE PAGURIDEA PHYLLOCHAETOPTERUS SOCIALIS PHYLLODOCE LONGIPES PILUMNUS SP. It POLYDORA IkET5bE~ RUOJLEMBOIDES I~LGAN SIPUNCULIDA C 5VLLlDAE TIMARETE SP. ACCALATHURA CHENULATA AMPHILOCHUS SP. AMPHINOMIDAE 8 ANCISTROSYLLIS RIGIDA 80WMANIELLA PORTORICENSIS CAPITELLA CAPITATA CARWEA CERATONEREIS SP. CHAETOPLEURA APICUlATA CHONE AMt.RTCANA CRASSINELLA LUNULATA DENDROChlROTIDA DIOOORA CAY£NENSIS It £BALIA SP. EUNICE SP. EURYDICE LITTORALIS HARIolOTHOE 5P. A HIPPOLVTE NICHOLSONI * HYPOCONCHA SABULOSA MELlTlDAE wINTER ;'olEAN 5T ERR O.C ·· · O.C 0.4 U.6 0.,+ 0.4 U.6 0.6 · ..· · ··· ·· ·· · 0.6 0.2 ·· · · ··· ·· · ··· ·• ·· ·• ·· · ··· ·· ··• ··•• ·· ·· 0.0 O.~ O.C ~.O O.C 0.2 0.2 O.~ 0.6 U.6 0.4 0.6 ·· ·· ·· · ·· ·· ·· · ··· ·· ·· ·· O.t:: 0.2 0.6 0.4 U." 0.4 0.2 0.2 0.2 0.2 U.2 0.2 15U2 N\IE~i SPRING Sf ERR ··· ·· ·· ·· · · ·· · ·· 0.4 0.4 0.6 0.4 0.6 U.2 0.6 0.2 0.6 0.4 0.2 iJ.2 · · · O.c · O.c ·· · · • ·· · · ·· · · ··· 0:4 0.2 0.4 0.4 0.2 0.4 · 0.2 0.2 0.2 U.2 0.2 0:2 U.2 0.2 0.4 0.4 0.4 0.2 0.4 0.2 0:2 0.6 0.2 0.4 · · 0.4 0.2 0.4 0.2 0.6 0.6 0.6 0.4 0.4 0.4 · · ·· ··· · 0:6 · 0.4 · · 0:2 0.6 0.4 o.~ 0.2 0:2 0.4 0.4 0.2 0.2 0.2 0.6 0.0 · U... · ·· 0.4 0.4 0.4 0.2 0:2 0.2 0.2 · · · 0.2 0.4 0.4 0.2 0.2 0.2 0.2 0.2 0.4 0.2 0.4 0:2 0.2 0.4 0:2 0.2 0.2 0.4 0.2 0.4 0.2 · 0.2 0.2 0.2 0.2 0.2 0.2 0.2 · · · · · · · · · ·· 0:2 FALL MEAN 5T ERR 0.4 0.4 0.2 0.2 ·· ·· 0.2 0.6 0.2 · · · ···• ·• SUMMER l"1EIIN 5T ERR O.t! O.t! 0.2 · · 0.2 0.2 · 0.2 0.2 0:2 0.0 0.2 0.4 0:2 0.2 o.~ ~.O · · · ·· • · · · · · 0.2 0.2 0.4 0.2 0.4 0.2 0.2 0.4 0.2 0.2 0.4 • 0.4 · · • · 0.2 0.2 0.4 0.4 · Appendix 3. (Continued) IS02 STA TION OVERALL ~ANK 220.5 ~20.5 01 -...I 220.5 220.':) 220.S 220.5 220.5 aO.5 220.5 220.5 220.5 220.5 220.5 220.5 220.5 220.5 220.5 220.5 220.5 220.5 295.0 295.0 295.0 295.0 295.0 295.0 2~5.0 295.0 295.0 295.0 295.0 295.0 295.0 295.0 295.0 295.0 295.0 295.0 295.0 295.0 295.0 295.6 295.0 295.0 295.0 295.0 29S.0 295.0 29S.0 295.0 29':>.0 2'15.0 295.0 2~5.0 295.0 SPECIES METOPORHAPHIS CALCARATA MICROPHRYS SP. NASSARIllS ALBUS NEMEIH INEA C NYMPHONIOAE OllGOCHAETA PARAPHOXUS SPINOSUS PERICLIMENES SP. PHYLLODOCE SP. PINNIXA SP. PORTUNUS SP. RUPELLARIA TYP ICA SABELLIDAf SElLA ADAMSI 51CYONJA SP. STENOCIONOPS FURCATA CDELATA • SYLLIS FERRUGINA TIRON TROPAt\IS TRICHOBRANCHUS GLACIALIS XANTHIDAE B ACANTHOHAUSTORIUS BOUSFIELDI AEOLI 0 I ACE A A AMPITHOE SP. ANACHIS lAFRESNAYI ARABELLA IRICOLOI-l AUSTRAEOlIS CAT INA BARBA TI A CAND IDA BATRACHONOTUS ~USOGARF CANTHARUS SUl GNATLU~ CAPITflLIDAE 0 CAULLERIELlA KIllARIENSIS CERATONEREIS VERSIPEOATA CHEVALIA SP. CHRYSOPETALIDAE CLYPEASTER SP. CRA5S0STREA VIRGINICA DONDICE OCCIDENTALIS DURVILLEA SOCIABILIS DROMIDIA ANTILLENSIS ECHINASTER SP. EPITONIUM HUMPHREYS I ERICH50NELLA FIL FO~MIS EUNICE TENUlS GLYCERIDAE GONODACTYLUS 8REDINI HEMIPOOUS ROSE US HES IONJDAE HETEROCRYPTA GRANULATA HIATELLA ARCTlCA HOLOTHURIA OAKARENSIS HYDROIOES SP. A HYPSICO~US PHAEOTAENIA ISCHNOCHITONIDAE A LAMELLARIA PERSPICUA lEPTOCHELA Sf'. SPRING WINTER MEAN 51 ERR ·• ·• · ·• ··• ·· ···• • ··• ·· ·• ·• · ··• ··· · ··• ·•• ··• ·• · ·· ·· ···• ·•• • ·· · ·• ·• · 0.2 0.4 0.2 0.4 0.2 0.2 0.2 0.4 0:2 0.2 0.2 0.2 0.2 0.2 • ··• • ···• ··• ·· ·· ·· ·• · 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 SUMMER MEAN ST ERR 0.4 0.4 • 0.2 · · ·•• 0.2 • 0.4 0.4 0.2 0.4 0.4 0.4 0.2 0.2 0.4 0.2 0.2 ·• 0.2 ··· ·· • ··• ·• · ·• ·· ·• · ··· ··• ·· ··• ·• 0.2 0.2 0.2 0.2 0.2 · · ·• • ··• ··• • ··• ·• · ··· ·• ··• ·· · ·· 0.2 ··· • • • • • 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 MEAN · · ·•• FAll 5T ERR 0.2 • 0.2 • • 0.2 0.2 0.2 0.4 0.4 · 0.4 0.2 0.2 0.2 ·• • • • 0.2 • 0.2 0.2 0.2 • • • · ··• ·• • ·•• ·•• 0.2 0.2 • • • 0.2 0.2 · ·· · ·• • • · 0.2 0.2 0.2 • · ·• ·•• • • 0.2 • • • • • 0.2 · · · ··• 0.2 0.2 • • • 0.2 0.2 0.2 0.2 0.2 • 0.2 0.2 0.2 0.2 0.2 5T ERR ·• 0.4 • • • MEAN • • • • · 0.2 ·• ·· • • • • • • 0.4 • 0.2 • • • • • • • 0.2 • 0.2 • 0.2 0.4 0.4 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 · ·•• • • • • • 0.2 • 0.2 • 0.2 0.2 • • • • • 0.2 · · ·• • ·• 0.2 • • • 0.2 • 0.2 0.2 • • • • • • 0.2 • • · 0.2 0.2 • • • 0.2 • • • 0.2 0.2 0.2 • • 0.2 0.2 • • Appendix 3. (Continued) 1502 STATION O\lE~AL RANK 295.0 295.0 295.(j 295.0 29~.0 29S.lJ 295.0 295.0 295.0 21.J5.0 295.(j 295.0 295.0 295.0 295.0 295.0 .:'95.0 29'j.0 295.0 295.0 29S.0 0\ 00 295.0 295.0 295.0 29!:l.0 295.0 295.{) 295.0 295.0 2<J5.0 295.0 295.0 295.0 295.0 295.0 295.0 295.(j 295.0 295.0 295.0 295.0 295.0 205.0 295.0 295.U 295.0 295.0 295.0 295.0 295.0 295.0 295.0 295.0 245.0 295.0 WINTEF< SPECIES LlNGA AMIANIU5 LUIDIA ALTEkf'..ATA LUM8RINU<IDAE LUMBRINEkIS IMPATIENS LYONSIA ':>P. LYSMAfA INTE.RI'EOIA LYSMATA SP. MACROCOELOMA TRISPINOSUM MA,J I DAE A ME5ANTHURA OECORATA MITHRAX FORCEPS MITHRAX PLEUkACANTHU5 MITRELLA RAVENELl MODIOLUS MODIOLUS SQUAM05US MONOCULOOES SP. MUREX FLORIFER DILECTuS MUSCULUS LATEkALIS NAINERIS SP. NEMAfONEREIS StJ. NEREIS ATNI~UCA NEREIS LAMELLOSA NEREIS SP. NEREfS SUCCINEA NOTOI:lRYON IO~Aw NUCULA CkENULATA O NTOSYL I~ ENOPLA UNUPHIDAE OPHIOCNIDA SCABRIUSCULA OPHIUROIDEA ORAINI IDAf: PALAEMONIOAE PALEANOTUS SP. 8 PANANTHURA fORMOSA PELECYPODA C I~ DfSCENS PERICLI"1EN ~ PERICLIMENES SP. it PETALOPROCTUS SOCIALI':> PETROLISTHES GALATHINUS PHYLLODOCE SP. E PINNOTHEf-lES USTREUM PINNOTHEKES 5P. PINNOTHERIOAE PITHO LHERMINIERI PLEUROPLOCA GIGANTEA POOOCHELA RIISEI PODOCHELA SP. POOODESMUS SP. POLYCHAETA G POLYCIRRUS SP. C POLYDORA COLONIA POLYDORA SOCIALIS POLYDORA SP. POLY NO lOAE POLYPLACOPHUHA C PORTUNIDAE MEAN Sf ERR 0.2 0.2 0.2 0.2 0.2 0.2 • ·· ·· · ·· ··· · ··• ·· ··· ·· ·· ·· ·• · ··· · ·· ··· · ·· · · ·• ·• · ·· ··• · ··• • ·· ··· ·· ··· ·· · ··· · ···• 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 5UMMEf-l GNI~PS • • 0.2 • • · MEAN · ··· · ·• ··• ··• ··• ·· · · ·•• ·· ·· · 0.2 Sf ER~ · ··· · ·· ·• · ·• ·• · • ·• • ·• · ·· · · ·· ··• ··· ·• 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0:2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0:2 0.2 0.2 0.2 0.2 • • • ··· ·• • · 0.2 0.2 0.2 0.2 0.2 0.2 FALL ME.AN 5T ERR · · ·· ·· · ··· · ··• ·· ·· ·· ·• ·• · · ··•• ·• ·•• ·•• · · ··•• · ··· · ··• ·· ·• 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 • • 0.2 0.2 • 0.2 0.2 • 0.2 0.2 0.2 0.2 • • • • 0.2 0.2 • 0.2 0.2 0.2 0.2 0.2 ·• ·• · ·• • • · · MEAN Sf ER~ • • • • • • ·• 0.2 • • • • 0.2 · • · ·••• ·• ··• 0.2 0.2 0.2 • • • 0.2 • 0.2 • • • • • 0.2 • • 0.2 • 0.2 0.2 • • ·· ·• • • 0.2 • • • • • 0.2 • • • • • 0.2 0.2 • • • 0.2 • • 0.2 • 0.2 • • • 0.2 • • 0.2 0.2 0.2 • • • • • • Appendix 3. (Continued) STATION OVE~ AL RANK 2':15.0 295.0 2~5.0 295.0 295.0 295.0 295.0 295.0 295.0 295.0 295.0 295.0 295.0 295.0 295.0 295.0 295.0 295.0 295.0 01 10 SPECIES \lINTER ~IAEM POTAMILLA RtNIFORMIS PRIONOSPIO SP. PROCERAEA SP. PYRGOSPIRA TAMPAENSIS SCHISTOMERINGOS RUDOLPHI SICYONIA BREVIROSTRIS SICYONIA BREVIROSTRIS * SICYONIIDAE SPELOEOPHORUS PONTIFEk SPIRORBIS SP. STENORHYNCHUS SETICQRNIS SYLLIDAE H TELLINA SYBARITICA TEREBELL IOAE THOR MANNINGI • TRIPHORA NIGROCINCTA TURBONILLA SP. ". VERMILIOPSIS ANNULATA VEXILLUM WANOOENSIS · ··• 0.2 ··• ·• SPkING ST ERR "4EAN ST ERR · ··• ·· ··· · · ·• ·· · ··· · ··· · ·• ·· ·· ·· ·· 0.2 ··· · 0.2 0:2 0.2 0.2 0.2 · ·• ··• 1502 0.2 0:2 · ·· ·· ·· 0.2 0.2 0.2 0.2 0.2 SUMf'lER MEAN ST ERR ·· ·· ·· 0.2 0.2 ·· ··· · ·• ·· 0.2 0.2 0.2 0.2 0.2 0.2 0:2 0.2 ·· · 0.2 0.2 0.2 0.2 FALL MEAN ST ERR 0.2 0.2 • · 0.2 0.2 0.2 • 0.2 0.2 0.2 • 0.2 0.2 . Appendix 3. (Continued) MS02 STATION UV'ERALL RANK ~ 0 1.ll 2.0 3.0 4.0 5.0 0.0 7.0 8.0 9.0 10.0 11.0 12.0 13.0 14.0 15.0 16.0 17.0 18.0 19.0 20.0 21.0 23.0 23.0 23.0 25.0 26.0 27 .0 21:l.0 29.0 30.0 31.0 32.0 33.0 34.0 35.0 36.0 37.5 37.5 39.5 39.5 4105 41.5 43.:i 43.5 45.0 40.0 47.0 48.0 49.0 50.0 51.5 51.5 53.0 55.0 55.0 MEAN FILOGRANA IMPLEXA SYLLIS SPONGICOLA MALACOCEHOS GLUTAEUS CHONE Ar-1EkICANA OPHIOTHRIX ANGULATA PHOTIS SP. ACANTHOHAUSTORIUS MILLSI MEGALOBRACHIUM SOklATUM LUMBRINEkIS INFLATA EXOGONE DISl-'AR OWENIA FUSIfUHMIS CHEVALIA st-'. LAEvrCARDIUM PICTUM LUCONAcIA INCERTA BRANCHIOSYLLIS EXILIS LEMMOS SMITHI MICRODEUTOPUS MYERSI AMPITHOE SI-'. A PAGURUS CARuLINENSIS SP. GAM~AROPSI THOR SP. AMPHIODIA PULCHELLA OPHIACTIS SAV'IGNYI PAGURUS SP. LEUCOTHOE SPINICAkPA XANTHJl)AE A MITRELLA LUNATA SUTUROGLYPfA IONTrlA NEMERTINEA SYLLIS HYALlr-.A PODARKE 085CLJRA POLYCIRRUS CAROLINENSlS PILUt-1NUS AUTOL nus SP. CAPRELLA PENANTIS CAPRELLA EQUILIBRA ASPIDOSIPHON GOSNOLOI WEBSTERINEREIS SP. B PSEUDOMEOAEUS AGASSIZII ~EBSTERINER IS TRIDENTATA HARMOTHOE SP. A PRJONOSPIO CRISTATA SPHAERODORIDIUM CLAPAREDII SYNALPHEUS IDNES ~OT EUNICE VITTATA ELASMOPUS SP. A POLYDORA CAECA EULALIA SAN&uINEA A~PHARETt. AMERICANA HIATELLA ARCTICA MEGALOMMA BIOCULATUM PROCERAEA SP. CHIONE GkUS AXIO&NATHUS SQUAMATUS SEMELE NUCULOIDES 5.-. ST ERR 1.8 26.4 31.8 4.2 5.6 11.2 18.0 5.2 21.4 1.1:l 17.1:l 9.4 4.4 12.0 6.0 4.4 11.0 4.2 2.8 3.6 9.0 3.2 33.2 19.6 t\ • 1 2.9 15.7 13.0 2.7 4.2 6.3 9.6 2.2 20.6 1.1 7.7 4.9 2.0 7.3 2.4 2.5 5.0 3.2 2.5 1.5 5.7 1.8 10.8 7.2 7.2 4.8 2.0 2.4 2.6 2.6 3.0 2.0 3.2 3.2 0.4 3.2 5.2 0.8 10.4 1.8 2.2 2.6 0.4 1.6 2.2 1.2 1./j 4.6 3.0 3.1 2.7 1.2 1.3 1.4 1.5 1.5 0.7 1.4 1.5 0.4 1.1 2.9 0.6 9.9 1.1 1.0 1.7 0.2 0.5 1.1 1.2 0.8 1.4 2.4 2.2 0.6 1.2 0.7 53.2 35.4 U.~l . . SUMMER SPRING WINTER SPECIE:.S . . FALL ST ERR ME:. AN ST ERR MEAN 1084.4 17.2 11.6 41.2 11.8 4.2 20.4 5.8 10.0 8.4 23.0 4.8 21.6 7.0 8.4 7.4 747.0 8.0 10.9 20.9 6.8 3.4 13.5 5.1 4.1 3.1 11.4 4.8 2.9 5.3 4.1 3.2 0.2 7.4 2.4 1.0 1.0 0.2 5.0 1.2 0.8 0.8 12.4 5.d 6.4 7.4 3.2 10.6 1.6 3.8 7.2 3.2 6.4 3.8 3.6 4.2 7.4 1.0 3.8 1.6 9.4 5.4 3.4 6.8 3.2 4.4 4.4 6.6 2.3 3.4 3.7 1.9 3.5 0.9 2.4 4.7 1.6 2.6 1.3 2.1 2.0 4.0 0.4 1.6 6.4 8.4 2.9 1.3 4.9 1.6 1.9 2.6 2.0 7.0 3.4 0.8 0.8 0.2 2.0 0.8 3.2 9.4 0.2 0.8 1.2 1.2 4.8 1.1 0.4 0.4 0.2 0.8 0.6 2.0 5.9 0.2 0.0 0.8 2.8 3.2 1.2 0.4 7.2 1.8 1.9 1.8 0.8 0.4 4.5 1.3 u.2 0.2 2.0 2.2 1.0 2.4 1.4 0.2 0.2 1.5 1.7 O.d 1. y 0.7 S31.6 44.6 36.8 3.4 1.4 4.'" 12.4 20.0 8.4 3.8 1.8 3.8 4.2 3.0 2.8 1.0 Y.6 1.6 7.2 0.8 3.6 1.8 1.6 5.4 2.4 6.6 1.4 1.0 1.0 2.0 2.6 1.2 0.4 1.0 0.5 0.9 1.7 0.6 0.2 0.3 2.8 1.6 0.8 1.7 1.0 0.4 1.0 0.8 0.6 0.2 0.4 1.2 0.6 1.6 0.1:l 0.5 0.4 0.2 0.2 0.7 0.6 0.9 NAE~ . • 3.6 2.2 3.8 0.6 5.8 0.6 6.6 5.8 5.8 3.0 1.8 2.6 . . 3.3 0.7 2.8 0.4 3.4 0.4 4.9 3.1 3.2 1.8 1 .1 2.1 · · • ·· · · · · · ·· · • 2.tl 2.6 3.8 1.8 6.4 4.0 1.0 • 1.6 2.6 1.0 3.2 0.6 4.4 . 2.2 4.2 2.8 8.6 1.0 5.4 0.2 0.6 2.2 2.4 3.0 1.4 ST ~RE 355.4 42.6 lY.3 1.6 0.7 3.1 10.1 10.8 3.3 1.6 0.6 3.3 2.0 1.5 1.2 0.6 4.0 1 .1 3.1 0.8 2.3 2.3 1.0 2.5 1.2 4.8 0.5 0.6 1.3 1.4 1.4 1 .1 2.6 1.9 1.0 . 0.6 1.0 0.6 1.5 0.2 1.a • 1.3 1.6 2.3 3.6 0.8 1.5 0.2 0.4 0.8 0.8 1.3 0.5 Appendix 3. (Continued) STATION UVERALL RANK 55.0 57.0 58.0 59.0 60.0 61.0 63.0 63.0 63.0 67.0 67.0 b7.0 67.0 67.0 70.5 70.5 72.0 73.0 75.0 75.0 75.0 -..I 77 .5 77.5 tlO.O 80.0 80.0 83.0 83.0 I'D. 0 85.5 85.5 13 7 • 0 1'l9.0 tl9.0 1'l9.0 '12.0 92.0 92.0 95.5 95.5 <.,)5.5 95.5 101.5 101.5 101.5 101.5 lOl.S 101.5 101.5 101.5 108.5 lu8.5 10tl.5 108.5 108.5 SPECIES SPIOPHANES BOMHYX TUR8ELLARIA PODOCERUS .P~ ERICHTHONIUS BRASILIENSIS CERATONEREIS VERSIPEDATA POLYCIRRUS ExiMIUS CERITHIUM ATHATUM MAJ IOAE A TRAVISIA PARI/A DlAS TYLI OAE A OPHIOSTIGMA SP. A PARACERCEIS CAUDATA PHERUSA EHLERSI TELL INA AMERICANA ELASMOPUS SP. l:l GOULDIA CERII'IA OPHIOSTIGMA ISACANTHUM NEMATODA BRACHYLJRA CHRYSOPETALIDAE A LUMBRINEkIS COCCINEA LEPTOCHELA PAPULATA SYNALPHElJS SP. GONIADIDES CAROLINAE LEPTOCHELIA SP. PAGURIDEA ODONTOSYLLIS ENOPLA PISTA PAU1ATA S I PUNCUL! DA A MEL! TI OAE A XANTHIDAE CARPIAS 8ERMUDENSIS INACHOIDES FORCEPS LATREUTES PARVULUS PRIONOSPIO CIRRIFERA AXIOTHELLA MUCOSA NOTOMASTUS SP. TRYPANOSYLLIS ZEBRA BODOTPIIDAE 8 CIRROPHORUS LYRIFORMIS MARPHYSA SP. A PHERUSA INFLATA CERATONEREIS SP. CERATor~ER IS SP. A CERATONEREIS SILBAR~ COLOMASTlX SP. LOIM 1A MEDUSA MELITA APPENOICULATA OPHIOSTIGMA SP. OPHIUROIDEA BOWMANIELLA PORTORICENSIS CIRRATUL!DAE MAERA SP. A MUSCULUS LATERAL IS NASSAR IUS ALBUS wINTER MS02 SPRING SUMMEH MEAN ST ERR MEAN ST ERR MEAN 3.0 2.4 2.2 2.0 1.8 1.8 1.8 2.0 0.9 1.5 1.4 1.0 0.9 1.b 2.4 0.4 2.0 1.b 2.4 2.0 3.b 1.3 0.4 1.4 0.5 1.1 1.4 2.4 0.6 1.2 1.2 0.2 0.6 2.2 5.8 2.l:j 3.4 1.6 0 ... 0.4 • 0.8 4.0 1.2 2.7 1.0 0.4 0.4 0.6 1.8 1.8 1.4 0.4 1.b 1.b O.b 0.6 1.2 1.4 1.4 1.6 4."+ 1.2 1.2 1.tI 1.6 0.£ 1.4 2.0 1.6 0.7 1.0 1.0 1.0 3.9 0.7 0.7 1 .1 1.0 0.2 0.7 1.5 0.5 I.!:! 1.6 0.2 1.2 0.2 0.9 0.7 0.2 O.b 0.2 0.6 1.2 0.6 0.8 0.4 • 0.6 0.8 0.4 0.4 0.2 2.6 O.b 1.4 0.9 1.0 0.2 2.1 0.6 2.2 0.4 3.4 3.0 1.4 1.6 0.8 3.0 0.6 0.8 0.6 1."+ 0.2 1.4 0.4 1.6 0:7 0.2 1.4 0.2 0.8 ·· • 2.2 ·• ·· • • 2.2 · · · · ·• ·· · • 1.5 · · FALL ST ERR MEAN ST ERR · 2.4 1.8 1.2 5.4 0.2 1.4 2.8 2.0 1.2 2.2 0.4 1.6 0.8 4.8 0.6 1.0 1.8 2.8 0.8 • 0.8 0.8 0.7 5.4 0.2 1.0 1.7 1.4 0.4 1.6 0.2 1.0 0.4 4.1 0.4 0.8 0.9 1.5 0.5 • 0.4 0.2 1.4 • 0.2 0.2 0.5 1.8 3.6 2.0 3.2 1.8 1.8 0.2 4.8 1.4 3.6 0.2 0.4 0.6 1.8 o.a O.b 1.4 0.2 0.6 0.7 0.7 0.2 0.4 0.6 O.b 1.0 0.2 4.4 2.0 0.2 1.4 0.4 0.4 2.2 2.4 2.6 2.1 1.1 0.2 0.5 0.2 0.2 1.6 1.7 2.3 1.6 1.2 1.0 1.0 0.6 0.4 1.8 1.8 1.4 0.4 0.8 0.8 2.0 1.6 0.4 0.6 0.4 0.8 O.b 1.4 0.9 0.2 0.4 0.8 0.5 • 4.b 0.2 · · · • 1.0 · 4.3 0.2 • 0.9 • 0.3 o.a · · · ·• · · · ·• • 0.8 2.2 0.2 1.2 1.8 0.8 2.4 1.0 0.8 • 1.4 1.3 1.6 1.3 0.9 0.2 3.2 1.4 0.6 0.2 0.4 0.4 1.3 • 0.4 1.4 O.~ o. 0.7 0.5 1.2 0.4 0.4 • 2.6 1.2 0.4 0.4 0.2 1.0 0.1 0.2 0.2 0.2 0.6 • 0.8 2.4 1.2 1.6 1.0 0.8 1.4 1.4 1.0 0.6 0.8 1.4 1.4 1.4 2.8 0.2 1.0 0.6 1.5 0.6 0.1 0.4 0.4 0.1 1.4 0.3 0.6 0.6 1.0 0.1 0.9 2.5 0.2 1.0 1.2 1.6 1.2 0.6 • 0.2 0.2 0.4 0.8 0.4 0.4 0 ... 0.2 0.4 0.4 0.2 • 0.2 • O.b • • · · · • · 1.2 1.4 0.6 • • 0.2 • 0.4 • ·· 2.6 0.2 • • 0.5 1.0 0.6 · • • • • • • • • • • 0.4 Appendix 3. (Continued) MS02 STATION OVEHALL RANK -...I N 108.5 115.':> 115.S 115.5 115.5 11':>.S 115.5 115.5 115.5 123.5 123.5 123.5 123.5 123.5 123.5 123.5 123.5 129.0 129.0 129.0 134.0 134.0 134.0 ) 34.0 134.U 134.0 134.0 140.5 140.5 14L1.5 140.5 140." 140.5 ISI.0 151.0 151.0 151.0 1')1.0 151.0 151.0 151.0 151.0 151.0 151.0 1"1.0 151.0 151.0 151.0 164.0 164.0 164.0 164.0 164.0 164.0 164.0 S1 ERR N~EM XANTHIDAE 8 ALPHEUS NORMAl';NI ARC A IMBRICATA CRASSTNELLA LUNULA1A MEGALUROPUS SP. PISTA QUADRIL08A1A TANYSTYLlDAl: A TELL INA SYBARITICA TRICH08RANCHUS GLACIALIS EUNICE FILAMENTOSA GASTROCHAENA HIANS MARGINELLA LAVALLEE ANA NASSAR INA RO~rI NICOMACHE TRISPINATA PAGURUS HENOERSONI PEllA MUTICA SICYONIA SP. AMPELTSCA VAOOKUM DIPLODONTA PUNCTATA sYLLlS GRACILIS ANCHIALINA TYPICA LAONICE CIRKATA LEUCOTHOIDES POTTS I PETALOPROCTUS SOCIALIS PROCESSA SP. SCOLOPLOS SP. A SYNCHELIDJUM AMERICANUM AMPHIPODA 8 ANOPLODACTYLUS PE110LATUS SP. A HYDROI E~ LITHOPHAGA 8ISULCATA MITHRAX PLEUHACANTHUS POMATOCEkOS SUNACIRE~A AMPELJSCA SCHELLENBERG I ARABELLA MUTANS CHONE SP. B COLOMASTIX .~S COMACTINIA MERIOIONALIS LyslANOPSIS ALBA MALOANJDAE MELPHIDIPPIDAE A NEPHTYS ASO~tAUQS PELECYPODA 8 PHYLLODOCIDAE SERPULIDAE SIPUNCULIOA SYLLTS ALTEkNATA SYLLIs REGULATA CAROLINAE AMPHARETE ACUTIFRONS CERITHIOPSIS GREENI LEMBOS UNICORNIS LYS I ANASS IOAE A LYSIDICE NINETTA PAGURUS HENDERSON I ~ PINNIXA SP. SPRING WINTER SPfC YES 0.8 2.0 LI.2 0.8 1.4 1.2 0.4 1.4 1.0 0.2 1.0 1.2 1.8 0.2 1.0 0.6 2.2 2.0 1.8 0.6 O.d 1.3 0.2 0.6 0.7 0.4 0.4 1.2 0.8 0.2 0.6 1.0 1.6 0.2 0.8 0.4 O.d 1.8 1.1 1.2 1.0 (J.O · ··• · · ··• · 1.8 1.2 1.2 0.8 0.2 1.4 0.6 1.0 1.0 0.7 0.6 0.2 1.0 0.2 ·· · ·· • • 0.2 • 0.2 0.0 0.4 0.6 0.4 0.4 0.8 0.4 0.8 0.2 0.2 1.4 0.2 0.2 0.2 0.2 0.8 0.2 0.4 0.2 0.2 0.9 0.2 ·· 0.4 0.4 1.8 0.4 ·· 0.2 0.2 1.8 0.4 ST ERR NAE~r 0.8 0.6 1.0 2.0 0.2 1.6 1.6 1.4 0.4 0.4 1.2 1.6 1.2 0.6 0.2 0.4 0.9 0.2 0.7 0.7 0.5 0.2 0.4 1.2 0.7 0.6 0.4 0.8 0.8 0.2 0.6 0.2 0.8 0.2 1.8 1.8 0.2 0.6 0.5 0.2 0.4 0.2 0.5 0.2 1.3 1.8 0:6 0.2 0.6 0.4 0.2 0.4 · · · · ·· 0.2 0.2 1.2 1.2 1.0 • 0.2 0.2 0.7 1.0 0.8 1.6 0.2 1.0 0.4 0.4 0.2 0.4 1.2 0.4 0.6 1.6 0.2 0.6 1.8 1.6 0.2 1.0 0.4 0.2 0.2 0.2 1.2 0.4 0.4 0.9 0.2 0.4 1.8 0.8 0.4 0.4 0.2 · · • 0.8 FALL SUMMER · · • 0.2 ME.AN · · ··• S1 ERR · · ··• 0.6 0.4 0.8 0.4 0.4 0.2 0.4 0.2 0.2 0.4 0.4 0.6 0.4 0.2 0.2 0.2 0.2 0.2 0.2 0.4 0.2 0.4 0.2 1.2 0.2 0.4 0.6 0.2 0.4 0.2 0.2 0.4 ·•• • 0.4 · · · ·•• MEAN ST ERR 1.8 1.0 1.0 0.5 ·• · · o.a 2.0 0.6 0.2 0.2 0.8 1.0 1.0 1.2 0.4 1.0 0.5 • 0.6 1.6 0.6 1.4 0.2 0.4 0.9 0.6 0.9 0.2 · ·• • 0.4 0.4 0.8 0.2 • • 0.2 0.2 0.6 0.2 · • 0.4 2.4 2.0 1.0 1.5 · · 0.2 0.6 1.4 0.6 0.2 0.8 0.4 • 0.2 0.6 0.5 0.6 0.2 0.5 0.2 • 0.6 0.2 0.2 0.6 0.8 0.4 0.2 0.2 0.8 0.6 0.2 0.2 0.4 0.5 0.4 0.2 0.2 0.6 0.8 0.2 0.8 0.4 • 0.6 0.2 0.5 0.4 0.2 0.2 0.4 2.0 0.4 0.6 0.8 0.8 0.6 0.6 0.4 0.2 0.6 0.4 0.2 0.2 2.0 0.4 0.6 0.8 0.6 0.4 0.6 0.4 0.2 0.6 0.2 0.2 1.0 • • 1.4 • 0.6 0.2 0.8 • • 1.0 0.2 0.6 • • • • • 0.4 • • 0.4 0.4 0.2 • 0.6 • • • • 0.4 • • 0.2 0.2 0.2 • 0.4 Appendix 3. (Continued) STATION OVERALL RANK SPECIES 'NINTER N~E / 164.0 164.0 164.0 164.0 177 .5 177.5 177.5 177 .5 177.5 ..., -.I 177 .5 177 .5 177.5 177 .5 177 .s 177 .5 177.5 177.5 177.5 177 .5 177.5 190.0 190.0 190.0 190.0 190.0 19U.0 190.0 190.0 1':10.0 203.0 203.0 203.0 203.0 203.0 203.0 203.0 203.0 203.0 203.0 203.0 203.0 203.0 203.0 203.0 203.0 203.0 229.0 229.0 229.0 <:'29.0 229.0 229.0 2~.0 229.0 229.0 SICYONIA LAEVIGATA SPIO PETIBO~AE TIRON TROPAKIS TURBONILLA INCISA ASCORHYNCHUS CASTELLI CAPRELLIDtlE A CAULLERIELLA KILLARIENSIS CERITHIOPSIS EMERSON I CREPIDULA ACULEATA OIOOORA CA1ENENSIS EULALIA MACROCEROS GALA THEA ROSTRATA GL YCERA CAP IT ATA HYB05COLEX LONGISETA LUMBRINERIS LATREILLI MARGINELLA AuREOCINCTA ONUPHIS NEBULOSA OSTRACODA PERICLIMENES SP. PHYLLOOOCE FRAGILIS CARIOEA CHAMA CONGREGATA LINGA AMIANTUS NEREIOAE C PAGURIOAt: PHYLLODOCE SP. SEMELE BELLASTRIATA SYLLlDAE I SYLLIDAE L 8RANCHIOSYLLlS OCULATA CHAMA MACER0PHYLLA COLUBRARIA LANCEOLATA GLYCERA TESSELATA HOLOTHUROIDEA HYDROIDES CkUCIGERA LUCIFER FAXONI MANGELIA RUGIRIMA PHYLLOCARIDA ~HYL OD CE CASTANEA PSEUDOVERMILIA SP. A PSEUDOVERMILIA OCCIDENTALIS SABELLARIA VULGARIS VULGARIS STENOTHOE SP. A SYLLIS SP. a SYLLIS SP. I VARICORBULA OPERCULATA APANTHURA MAGNIfICA ARMANDIA MACULATA CAPITELLIDAE A CERAPUS TLltlULARIS CHRYSOPETALIDAE B COLOMASTIX SP. A CREPIDULA PLANA CRUCIGERA ,.EtlSTERI CUMINGIA TELLINOIDES 101502 I.e 0.0 0.2 0.4 SPRING 5T ERR O.d SUMMER 5T ERR NAE~( ·• · · · ·· · 0.4 0.4 O.d 0.4 0.2 0.2 0.6 0.4 0.4 1.0 0.5 0.4 0.4 0.4 0.4 0.2 0.4 0.6 0.4 0.4 0.4 0.2 0.2 0.2 0.6 0.2 1.6 0.2 0.2 0.6 1.2 1.2 0.8 0.8 0.2 1.6 0.2 0.2 0.4 1.2 1.2 0.4 0.6 1.0 0.8 0.2 0.4 0.6 0.4 0.2 0.2 0.8 0:2 0.2 0.4 · 0:6 · · o.~ ·· · · ·• ·· · ·· ·· · · · · O.d · · • · · · ·· ·· · ·· ·• 0.4 0.2 1.4 U.4 1.4 0.4 1.0 1.0 · · • 0.6 · · · · ··· 1.0 0.6 0.6 0.4 0:2 0.2 0.8 0.8 0.4 0.2 0.4 O.b 0.2 0.2 0.4 · 0.2 0.2 0.6 0.4 0.4 0.2 0.8 0.4 0.6 0.2 O.d ·· · NAE~ ··• FALL ST ERR MEAN ST ERR ·· 0.6 0.8 1.6 1.0 0.2 0.6 0.8 0.4 0.2 0.4 0.2 1.6 0.2 0:4 0.2 1.6 0.2 1.0 0.8 0.4 0.4 0.4 0.2 0.2 0.2 · · ·· · ·• • 1.0 1.0 0.8 0.6 0.2 0.4 0.8 0.6 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0:4 0.4 0.2 0.4 1.2 0.2 0.4 0.2 1.2 0.2 0.6 1.2 0.2 0.2 0.2 0.4 0.2 0.4 · ·· · o.~ 0.2 · ·• · O.d 0.5 0.6 0.4 • 0.4 · 0.8 0.2 · 0.2 0.4 0.8 0.4 0.2 0.2 0:2 0.4 0.2 0.2 0.2 0.2 0.2 0.4 0.2 0.2 0.2 0.2 0.2 0.4 0.4 0.2 0.4 1.0 0.2 0.2 0.6 · O.d · · · O.d · · o.~ · • 0.2 0.8 0:2 0.6 1.0 0.4 1.0 0.2 ·· · ·· · 0.4 · • 0.2 ·• ·• ·· · · ·· ··· ·· · 0.2 • 0.2 ·· ·• 0.2 0.2 0.2 0.2 0.4 0.2 0.6 0.2 0.2 0.2 0.4 0.2 • 1.0 0.2 0.6 • 0.6 0.4 0.2 0.4 0.6 0.4 0.6 1.2 0.2 0.8 0.4 0.6 0.2 0.4 O.d ·• • 0.8 0.6 0.6 0.2 0.2 ·· · · · ·· ·· ·· · O.~ ·· · · ·· • · · · 0.4 0.4 0.4 0.4 0.2 0.2 0.2 0.2 0.6 0.6 0.2 0.5 0.2 0.2 0.2 0.1 0.4 0.6 0.2 0.4 o.~ 1.0 0.4 0.4 0.2 1.2 0.4 0.8 0.6 0.4 • 0.6 · 0.2 · 0.4 • 0.2 Appendix 3. (Continued) MSU2 :::'TATION KNA~. 0.92~ 229.0 22Y.0 C29.0 229.0 229.0 229.0 229.0 229.0 a9.0 229.0 229.0 C29.0 229.0 ~29.0 -..J ~ 229.0 229.0 229.0 229.0 229.0 229.0 229.0 229.0 229.0 229.0 229.0 ~63.5 ~63.5 i:'63.5 203.5 eA3.5 263.5 203.5 263.5 263.5 263.5 263.5 263.5 203.5 263.5 263.5 263.5 263.5 ~63.5 203.5 263.5 ~63.5 263.5 263.5 263.'S 263.5 263.5 263.5 ~63.5 263.5 SPRING wINTER SPECIES L A~EVl MEAN EURYDICE LITTORALIS HAPLOSCOlOPLOS FHAGILIS HESIONIDAE HIPPO!. 't' TE. SP. HIPPOLYTIDAE HYPOCON1HA SA8ULOSA JAEROPS 5 CuRAlICOLA LIMA PELLlJCIDA LYONSIA HYALlNA MAGlLONA SP. A MAJ LJAE MITHRAX SP. MODIOLUS AMERICANUS NEREIS SP. OPHELIA DENTICULATA PAll CUS SP. PAkAPRIONOSPIO piNNATA PERSICULA CANTENATA POLYLJORA SP. POLYDORA TETRA8RANCHIA SCHISTOMERINGOS RUDOLPH I SPIONIDAE STREljLOSOlo1A SP. SYLLIDAE C SYNALPHEUS fRITZMUELLERI VERM Il. lOPS IS ANNULATA AONIDES MAYAGUEZENSIS ASPIUOSIPHON AL8US AUTOMATE EVERMANNI BATRACHONOTUS FRAGOSUS CAPlTELLIDAE DIPLODONTA SEtoAIASfJERA EULALIA SP. EUN IC TOAE GLYCERA SP. C GLYCERA PAPILLOSA H[TEROCRYPTA GRANULATA HETEHOMYSIS FORMOSA KURTZIELLA ATRuSTYLA LEMAOS UNIFASCIATUS REDUCTUS MELf TIDAE. MICROPANOPE SP. MICROPHOLIS FACHYAACTEHA NEREIDAE NEHEIS RIISU OLI GOCHAET A PALAEMONIDAE PARAONIDAE PERICLIMENAEUS SCHMITTI PHERUSA SP. PISANlA TINCTA PLEUROMERIS THIDENTATA PRIONOSPIO SP. PROTODORVILLEA KEFEHSIE.INI SCOLOPLOS RUI3RA .. O.~ ··· ST MEAN ~RE 0.2 0.2 0.6 0.2 0.6 0.6 0.4 0:4 0.4 0.4 0.4 0.2 1.0 0.2 0.6 0.4 0.6 v.2 0.4 · 0.2 0:4 0.2 1.0 0.8 0.4 0.2 0.6 0.6 1: 0 0.5 0.2 0.4 0.2 0.2 O.~ 0:2 0.2 0.4 0.4 1.0 0.6 · ·· · ·· ·· ·· · ··· · 0.4 0.4 0.2 U.6 0.2 0.4 0.6 · · 0.6 0.8 0.5 o.e 0.2 0.6 0.4 0.2 0.2 0.2 0.6 0.2 0.2 0.2 0.2 0.6 0.2 · · 0.4 0.2 0.2 0.2 0.2 0.4 U.8 0.2 U.2 0.2 0.2 0.4 0.6 0.2 0.2 0.2 · · ·· ·• ··• · · 0.2 0.2 O.tj 0.6 ·• 0.4 0.6 0.6 0.4 0.4 0.4 0.2 0.2 0.2 0.2 0.6 0.2 0.4 0.4 0.2 0.2 ·· ·· · ·· · · ·• · 0.6 0.2 0.2 0.2 · ·· · · · • U.4 0.2 0.13 0.4 0.4 0:4 0.2 0.8 0.2 0.2 0.6 0.4 0.2 0.2 0.2 ·· · ·· · ·· · · · ·· · · ·· O.b 0.4 · 1.0 · · · · 1.0 0.6 0.8 • 0.8 0.2 · 1.0 0.4 o.a 1.0 0.6 0.2 • · · · ·• · • · 0.6 0.2 1.0 0.2 0.2 0.2 0.2 1.0 0.2 0.2 · 1.• u 0.4 1.0 0.2 0.2 0.2 • 0.6 0." 0.8 0.6 0.2 0.2 0.2 0.0 0.2 0." 0.2 • 0.2 ·· · · ··• 1: U ·· ·• 0.2 0.2 0.6 0.2 0.2 0.4 0.2 0.2 0.2 0.2 0.6 0." · · · ·• • • • 0.4 0.4 U.4 0.2 0:4 0.2 0.6 0.2 0.2 0.4 0.2 0.2 0.2 0.2 0.6 0.2 0.2 0.4 0.4 0.4 0.4 ·· · ·· ST ERR 0:4 0.4 · · · · ·· ·· · MEAN · · o.a · O.b 0.4 0.4 · · ·· ·· · · ·• · · ·• ·· · O.e · · 0.2 0.6 0.4 · · ST ERk 0.2 0.2 0.2 0.2 ().2 · MEAfi 0.6 0.2 0.2 0.2 0.2 ·· ·· · ·· · · ST ERR 0.2 • • FALL SUM I: ~ · · • · ·• ·• · ·· · · · · 0.4 0.6 0.8 0.2 • • 0.4 0.6 • o.a 0.2 · · 0.4 0.4 0.4 0.2 Appendix 3. (Continued) STATION OVERALL RANK -.J VI 263.5 263.5 c63.5 263.5 263.5 310.':> JI0.5 .310.5 310.5 .jl0.5 310.5 310.5 310.5 310.5 .310.5 310.':> JI0.5 310.5 310.5 310.5 310.5 310.5 310.':> 310.5 310.5 310.5 310.5 310.':5 31u.") 310.':> JI0.5 310.5 .HO.,,) 310.':> 310.':'> .HO.5 310.5 310.5 310.") 310.5 ::n0.5 310.5 310.5 310.5 310.5 310.5 310.5 310.5 310.':> 310.5 310.5 310.5 .310.5 310.5 JI0.5 SPECIES SIPUNCULIDA E SPIONIDAE I:J SYLLIS CORNuTA TRYPTON SP. wE8SrERINEREIS SP. A ALPHEUS SP. AMPELISCA CRISTOIDES AMPHIPODA E ANAIHXIS SP. AIIlOMIA S(,lUAMULA ARBACIA AT LUTCNU~ ASTERIAS FORBESII BRAChIOPOOA A BRANIA CLAVATA CHIONE SP. * CHIONE LATILIRATA CIRRIFORMIA FILIGERA CORBULA CONTRACTA CUMACEA CYCLASPIS VARIANS DENDRODORIS KREBSII DIASTYLIOAE EBALlA STIMPSONI ENGINA TUR8INELLA EUNICE SP. EUNICE ANTENNATA EUPHROSINE TRILOBA EXOGONE SP. FAVARTIA CELLULOSA GLYCERA SP. B HEMUS CRISTULIPES HIPPOMEDUN SP. A HYDROIDES UNCINATA LAEVICARDIUM MORTUNI LASAE IOAE LEUCOTHOE SP. A MAERA CAROLINIANA METOPORHAPHIS CALCARATA NEOPONTONIDES SINET~OFUAE8 NEPHTYS INCISA OD ~TOSYL IS SPa OGYRIDE5 SP. ONUPHIS PALLIDULA PAGURISnS TORTUGAE PEPICLIMENES AMERICANUS PILUMNUS SAYI PINNOTHERIDAE PISIONE REMUTA PITHO LHERMINIERI SABELLIDAE SEMELE PURPURASCENS SEROLI S MGR AYI SPEOCA~CINUS CAROLINENSIS STENORHYNCHuS SETICORNIS STROM8JFORMIS BILINEAfUS MS02 WINTER MUN ST ERR 0.4 0.2 0.2 0.2 ·· ·· · ··· · ·· · 0.6 0.2 0.2 0.6 0.6 0.6 U.2 • 0.2 • 0.2 * SPkING ·· ··• · ··• 0.4 0.2 0.2 0.6 • ·· 0.6 0.4 0.2 • • 0.2 0.2 0.4 0.2 0.6 0.2 0.2 0.2 0.6 0.2 0.2 0.2 0.6 0.6 ·• · ·· · ·• · ·· · ·· ··• ·• · 0.6 0.4 0.6 0.2 0.2 0.6 0.2 0.2 0.2 0.4 • • 0.2 • 0.4 ·· · ·· 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 MEAN • SUMMER ST ERR · · ·· · ·· · ·· ·· ··• · 0.4 0.2 0.4 0.6 0.2 0.2 0.4 0.4 0.4 0.4 ·• • • 0.2 0.6 0.2 0.2 0.2 0.6 0.2 0.2 • 0.6 0.4 • • • · ·· ·• ·· 0.4 0.2 0.6 0.6 0.4 0.2 0.2 0.4 0.2 0.2 0.2 0.2 0.4 0.2 • · ·· ··• · · • · ··· · 0.6 ·• ··• ·• · 0.6 0.4 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 • ··• ·· · 0.2 0.2 MEAN • • 0.2 0.2 0.2 ·•• • FALL ST ERR ·• ··· · · ·· · · ·•• 0.2 0.2 0.2 0.4 0.2 0.4 0.2 0.2 0.2 MEAN ST ERR 0.4 0.4 0.4 0.4 0.2 0.4 0.6 0.2 0.4 0.2 • • · ·• • • 0.6 0.2 0.2 • 0.4 0.2 0.2 0.2 0.2 0.2 0.2 • • • • • 0.4 • 0.4 0.2 0.2 0.2 0.2 0.2 • • • • 0.2 0.2 0.4 0.2 0.2 0.2 0.2 0.2 0.2 0.4 0.2 0.2 0.4 0.4 ·•• ·• ··•• ·•• ·• ··· ·• ·• ·· 0.2 0.2 0.2 0.2 0.1t 0.2 0.4 • • · 0.4 · · ·· ···• • • • • 0.2 0.2 • • • • • 0.2 • • • 0.4 0.4 ·• ·•• ·• · 0.2 • • 0.4 0.4 · ·• 0.6 • • • 0.6 0.6 • 0.6 0.4 0.4 0.4 • 0.2 0.6 · • 0.4 0.4 0.4 • • • 0.2 0.2 0.6 0.2 0.2 0.2 0.2 0.4 0.2 0.4 ·• • • • 0.4 0.4 0.4 • 0.2 0.6 • 0.4 0.4 0.2 • 0.2 0.2 0.6 0.2 0.2 Appendix 3. (Continued) STATION UVEHALL HANK 310.5 310.5 310.5 310.5 310.5 310.5 310.5 . n O.5 310.5 310.5 36~.5 369.5 "36'1.5 369.5 369.5 369.5 369.5 3,,9.5 369.5 3b';i.5 -...l 0'1 369.5 369.5 369.5 369.5 369.5 3h9.5 369.5 3b~.5 369.5 364.5 369.5 369.5 369.5 309.5 369.5 369.5 369.5 369.5 369.5 369.5 369.5 369.5 369.5 369.5 369.5 369.5 369.5 369.5 369.5 369.5 369.5 369.5 J6'1.5 369.5 369.5 MEAN SUBADYTE PELLUCIDA SYLlIDAE SYllI5 SP. tJ 5YLLIS 5P. G TANAIDACEA l::l TEREBELLIDAE THOR FLOklDANUS THOR MANNINGI TRIPrlORA NIGkOCINCTA TURAONILLA ATPUR~ETNI ACANTHOHAU5TORIU5 SHOtMAKERI AMPHIODIA TRYCHNA AMPHIPODA F ARCA ZERRA ARICJDEA SP. ASTROPECTEN SI-'. BARNEA TkUNCATA BODOTRIJDAE A CHIONE AERUP~ ATNI CIRRATULUS SP. OIOOORA LISTERI DORIOPSILLA AL80PUNCTATA DORIOP5ILLA ROwENA AMEHICANA ETHUS" MASC~ONE EULALIA VIRIDIS HARMOTHOE :'P. HEMJPOOUS RuSEUS HYDRO IDES 51-'. C LATRE.UTES :'P. LEPTOGNAIHIIOAE A LUMBRINEkIOPSIS PA~ DOXA LYONSI" SP. LYONSI" 5P. ... LYONSIA 8EANA LYSIDJCE SP. A MADRELLA .~S MEDIOMASTUS CALIFOHNItNSlS MITHRJlX FORCEPS MUN ILl A 5P. MURE X PO~IUM NATICA PUSILLA NOTOMASTUS HEMIPuOUS OCNUS PYGMAtUS ONUPHIDAE OPHIODERMA MUNIP5 VE~it PARAPHOXUS SPINOSUS F'ELECYPOOA G PERICLlMl::NAEUS SP. A PHIDIANA LYNCEUS PILUMNUS DASYPODUS PODOCHELA .P~ PODOCHELA SEPJLICA~G POLYCERA AURISULA PORTUNUS SP. PRION05PIO SP. C · · ·· ·· ST ERR · · ··· · 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.4 0.4 0.2 0.2 0.2 0.2 0.2 0.2 0.4 0.2 0.4 0.4 0.2 0.4 ·· ·· ·· ·• ·· ·· · · ·· ·· MEAN 5T ERR MEAN 0.6 0.2 0.6 0.2 0.2 0.6 0.2 0.2 0.6 0.2 0.6 · · • 0.6 0.6 ·•• · ·•• ··· ·· ·• · · ··• ··· • ·• ··· ··• • 0.4 0.4 0.4 · · · · ··• ·· · · ·••• 0.6 0.4 0.4 0.4 0.4 0.4 U.4 0.2 0.4 0.2 0.2 0.,+ • 0.4 • 0.4 • • 0.4 0.2 0.2 0.2 0.2 0.4 0.2 0.4 0.2 0.4 0.2 0.2 0.4 0.2 0.2 0.2 0.4 0.2 0.4 0.2 0.2 0.2 0.2 0.4 0.4 0.4 ~.o · ·· ··• · 0.,+ · · ·· ·· · · ·• ··• ·· 0.4 0.2 O.c? ·• ·· ··· · ···• ·· ·•• · · 0.2 0.2 0.4 0.4 0.4 0.2 0:2 0.2 0.2 0.2 · FALL SUMMER GNI~PS WINTER SPECIES MS02 • · ·· ··· ··• ·• ·•• ··· ·• ·• ···• · · ··• · ·• ··• • ·· ·· ·· 0.2 0.2 0.4 5T ERR • • • • 0.2 · ·• · ·•• ·••• 0.2 0.2 • • 0.4 • • • ··•• ·· · ·· ·• ·• ··• • ·• ·•• 0.2 0.1t 0.2 0.4 0.4 0.4 0.2 0.4 0.2 0.2 0.4 0.4 0.2 0.2 0.1t 0.2 • MEAN 5T ERR 0.4 • • • 0.4 • 0.4 0.2 0.4 0.2 • 0.1t • ·•• 0.2 • • • 0.2 · ·• · ·•• ·•• ·• ·•• ·• ·• • • • • • • • 0.2 • 0.2 • • • 0.2 • • 0.2 0.4 0.4 o.~ o. 0.4 0.4 0.4 · 0.2 • 0.2 0.2 0.2 0.2 0.2 0.4 • • • • 0.4 0.2 0.2 • • • • • 0.2 • • • 0.2 • • • • 0.2 • • • • 0.4 0.2 0.2 Appendix 3. (Continued) STATION OvERALL RANK 369.5 369.5 369.5 369.5 369.5 369.~ 369.5 369.5 369.5 36'i.5 -..I -..I 369.5 369.5 369.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 481:1.5 488.S 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 48.~ 488.5 48 .~ 488.5 488.S 488.5 488.5 488.5 4ti8.~ 488.5 4'38.5 SPECIES PSEUOElJRYTHOE AMljIGUA SCOLOPLOS ARMIGER SIRAIUS BICULOR SPEOCARCINUS CAROLINENSIS • STENOTHOE ::'P. 1:3 STENOTHOE GEOHGIANA SYNALPHEUS MINUS TELLINA AT ~RETLA THY ONE PSEUDOFUSUS THICHOPHOXUS FLORIDANUS TURAONILLA SP. 0 TURRIDAE F ZEBINA BROWNIANA ACTEON CANDEN5 ALABA INCERTA ALPHEIDAE ALPHEUS SP. 8 ALPHEUS FORMOSUS AMBLYOSYLLIS FORMOSA AMPELISCA VERRILLI AMPHILOCHUS SP. AMPHIPODA G AMPITHOIDAE A AMYGDALUM SAGITTATUM ANCISTROSYLLIS RIGIOA ANOfJLODACTYLUS INSIGNIS ARABELLA IRICOLOR ARCOPSIS ADAMSI ARENE TI-lICAHINATA ARICIOEA FRAGILIS ARICIDEA SUECICA AUSTRAEOLIS CAT INA BARLEfl A SP • • BODOTRI lOAE C BODOTRIIOAE 0 BOTULA FUSCA BRANCHIOSYLLIS SP. 8RANIA SP. CAECUM CUB IT ATUfoI CALLIOSTOMA SP. CALLIOSTOMA PULCHRUM • CAPITELLIDAE 1:3 CERADOCUS SP. CERATONEREIS IRRITAI::lILI5 CHAETOPTt::RIOAE CHAETOZONE SETOSA CHIONE CANCELLA1A CHRYSOPETALIDAE C CHRYSOPETALIDAE 0 CONUS DAlJCUS CRENELLA DIVARICATA CUMINGIA COARCTATA DASY8RANCHUS .~S DELECTOPECTt:N SP. DENDHODOHIS SP. A wINTER 1'1502 SPRIN(, SUMMER I'1UN ST ERR MEAN ST ERR ·· ·• · ·· ·• · ·· · ··· ·· ·· ··· ··• ·· · ·•• ·· ··• · · · ··• • · ·· ·· ·· ··· ··• · ··• ·• ··· · ·· ··• ··• 0.2 0.2 0.4 0.4 0.4 0.2 U.2 0.4 0.4 0.4 0.2 0.2 0.4 0.2 ·· ··· ·• ·• ··· ·· ··· · ·· · ·· ·• • · ··• ·• ··· · ··· · ··· ·· · ··• • ·· • ··• · 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0:2 0.2 ~:o 0.2 0.2 0.2 0.2 0.2 0.2 0.2 · ·· · • 0:4 0.4 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0:2 0.2 0.2 0.2 0:2 0.2 0.2 0.2 0.2 · ·· ·· · ·· 0.2 0.2 0.2 0.2 0.2 0.2 FALL MEAN ST ERR 0.4 0.4 0.2 0.4 0.2 0.2 0:4 0.4 · ·· ·· ·• ·· · ·· · ·· · ·· · · ·· ·· · · ·· · ·· · ·· · ·· 0.4 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 ·· 0.2 0.2 u.2 ·· ·• • u.~ · ·· ·· ST ERR ~IAEM ··• ·· ·• ·•• · ·· • 0.4 • 0.4 0.2 • • • U.2 • • ·• · 0.2 0.2 0.2 0.2 • 0.2 • • 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 · · · · · · · · · • • · 0.2 0.2 0.2 • 0.2 Appendix 3. (Continued) MS02 STATION OvERALL RANK 488."> 48.~ -..I oc '+88.5 488."> 488.5 488.5 488.5 488.5 488.5 488.5 488."> 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488."> '+88.5 488.5 488.';; 488.5 488.5 488.5 488.5 48d.5 488.5 '+88.5 488.5 4H8.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 4t:SB.5 488.5 488.5 488.5 488.5 48.~ 488.5 488.5 488.5 488.5 488.5 488.5 488.5 J'Il E~r DIODORA CAYENENSIS * DIOPATRA CUl-'i-<EA DONO ICE OCCluENTALIS OORTOPSILL4 LElA DORVILLEA SOCIABILIS DROMIDIA ANTtLLENSIS DROMIDIA ANTILLENSIS l> ECHJNASTF-R SPA EPJTONIUM FOLIACEICUSTUM ERATO EAIHEGUA~I ERICHSONELLA FILIFORMIS ErEONE LAC TEA GALATHEIDAE GASTROPODA GENOCIDARIS MACULATA GLOTTIDIA PyRAMIDATA GLYCERA SPA GL YOMER I S SPA GONEPLACIDAE GONEPLAX S l(iS8E I tJONTADIOAE GONODACTYLUS BREDINI C"RAtJSIOAE GREGARIElLA CORALLIOPHAGA GYPTIS BRfVIPALPA HAPLOSCOLOPLOS FOLI0SUS HERMODICE CARUNCULATA ~IYOR TDES SPA B HYOROIDES SPA D HYPERJIOEA JNACHOIDES FORCEPS * IRIDOPAGURUS DISPAR * ISCHNOCHITONIDAE A LEIOCAPITELLA AR~LG LEIOCHRIOES PALLIDICR LEM80S SPINICARPUS INEH~S LEPTOCHELA SPA LUCAPJNELLA LIMATULA LUMBRINEF<IDES ACUTA LYSIANASSA SPA MACROCOELOMA SPA * MACROCOELOMA EUTHECA ~ MAGELONA SP. MAGELONA SPA C MALLUVIUM BENTHOPHILUM * MARIONIA CUCULLATA MEGALOMMA SPA MEIOSQUILLA QUAORIDENS MELANELLA CONOIOEA MUREX FLORIfEH DILECTUS MURJCIDAE NAINERIS SPA NANOPLAX XANThiFOHMIS * NATANTIA NEMA10NEHEIS SPA O.~ ·· ·· ·· ·· ·· ··· ·· ·· · ·· · · 0.2 0.2 0.2 0.2 0.2 0.2 ST ERf.< 0.2 · · O.;c · ··· ·· ·· ·· ··· ·· ·· ·· · 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 ··• 0:2 ·•• ·· ·· ·· · ··• ·· ·• 0.2 0.2 O.~ SUMMER SPRING wINrE~ SPECIES • 0.2 ·· ·· ··· ·· · ·· -.· 0.2 0.2 0.2 E~ AN ··· ·· ·· ··· ·· ··· ·· ·• 0.2 0.2 ST ERR MEAN ST ERf.< ·· 0.2 · · · ·· · · · O.;c · ·· · · ·· ··· ··· · · · ·· ··· ·· ·· ·· · ··· 0.2 0.2 0.2 0.2 0.2 0:2 0.2 0.2 0.2 0.2 U.2 0.2 0:2 0.2 ·· · · ·· ·· ··• ··• ··· ·· · · 0.2 ·• ·• · ·· ·· ··• ··· ·· ·· ·· • 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 · ·· ·· 0.2 · 0.2 ··· • · 0.2 O.~ 0.2 0.2 0.2 0.2 MEAN FALL ST E.RR · 0.2 · 0.2 0.2 · 0.2 • 0.2 0.2 • 0.2 0:2 • 0.2 0.2 0.2 0.2 · 0.2 0.2 U.2 0.2 • · · · · • • 0.2 • ·• ·· · ··· ·· ·• ·• • 0.2 0.2 • 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 U.2 0.2 0.2 0.2 • ·• · ·• ·• · ·• · ·•• 0.2 ·• • · · 0.2 ·• 0.2 0.2 0.2 0.2 0.2 ·• • Appendix 3. (Continued) STATION OVERALL RANK SPECIES WINTER N~EM -J \Q 488.5 488.5 488.5 488.5 488.5 488.5 41:\8.S 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 '+88.5 488.5 4/18.5 488.5 488.5 488.5 488.5 NEMERTINEA E NEOPONTONIOlS BEAUFORTEN5IS NEREIDAE A NEREIS SUCCINEA NOTOMASTUS lATERICEUS ONUPHIS EREMITA OPHIOCNIDA SP. OPHIOCNIDA SCABRIUSCULA OPHIOPHRAG~US PULCHER ORBINIA RISERI ORBINIIDAE OSn~EA PERMOLI S OVALIPES SP. PACHYCHELES kUGIMANUS PAGURISTE.S SP. PAGURISTES TRIANGULATUS PALEANOTUS SP. 8 PALOLO SICILIENSIS PAPYRIDEA ~IMROFNELS PARAMETOPELLA CYPRIS PARAPIONOSYLLIS SP. A PECTINIDAE A PELECYPODA PELECYPODA C PELECYPOOA H PENAEIOAE PERICLIMENE':) IR IOESCENS PETROLISTHES GALATHINUS PHASCOLOPSIS GOULDI PHORONIS ARCHITECTA PHTISICA MARINA PHYLLODOCE SP. E PHYLLODOCE ARENAE PHYLLODOCE lONGIPE.S PILUMNUS FLORIDANUS PINNOTHERES SP. PI ONOS YLLI S SP. PIlAR FULMINATUS POLYCHAETA POLYCHAETA H POLYCIRRUS SP. t3 POLYCIRRUS EXIMIUS OUBIUS POLYNOIDAE PONTONIINAE POTAMILLA RENIFORMIS PROSCOLOPLOS SP. PSAMMOLYCE A~OHP OINETC PTE~IA COLYMtJUS PTE~OMERIS PE~PLAN PTILANTHURA TRICARINA PYRUNCULUS CAELATUS RILDAROANUS LAMINOSA RISSOINA SP. A RISSOINA CATEStlYANA SCOLOPLOS SP. 0.2 • • ··· · 0.2 0.2 • • 0.2 ··• ·· · · · · ··· ·· · ·· ·· ·· · ··· ·• ·• ·· • · SPRING 5T ERR MEAN 0.2 ·•• ·• ·•• ··• · • • ·• · ··• ·•• ·• ·· ·· · ·· ·• ··• ·· ·• ·• ·• ·· ·• · ··• ·· ·• · ··· ·• · ·• ·· ··• · ·· ·· ·• 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 MS02 0.2 0.2 o.~ 0.2 o.~ 0.2 0.2 0.2 • • 0.2 0.2 0.2 0.2 0.2 ·· 0.2 0.2 0.2 0.2 0.2 SUMMER 5T ERR • • 0.2 0.2 ·••• 0.2 0.2 ·· ··•• • · ·••• ·•• ·• ··•• • ·•• • ·•• ·• · ·· · ·· ··• ··• · ·· 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 MEAN 0.2 0.2 0.2 • • • • 0.2 FALL ST ERR ·••• • • • • • • 0.2 • MEAN • 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 • • • • • • ·••• ·• ·• ·• • • • • • • 0.2 • • • • • 0.2 0.2 0.2 0.2 0.2 • • 0.2 o.~ • • ··• • 0.2 0.2 · 0.2 0.2 0.2 0.2 · · 0.2 • • 0.2 0.2 0.2 • • • 0.2 0.2 0.2 · ·• 0.2 0.2 0.2 0.2 · · · · ··• ·· • • • • • • • • • • • • 0.2 0.2 0.2 0.2 0.2 ST ERR • • • 0.2 • 0.2 ··•• • • • • • • • • • • 0.2 · ·•• ·••• • 0.2 • • • • • · • • • • • • 0.2 0.2 • • • • 0.2 • • • • • 0.2 O.~ o. 0.2 0.2 • • • • • • • • • • 0.2 • • • • • • • 0.2 • • • • • • Appendix 3. (Continued) STATION tJVERALL qANK 4ijtl.5 488.5 ,+88.5 ,+d8.5 48tl.5 '+88.5 oc 0 4tl8.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.5 488.') 48tl.5 488.5 488.5 488.5 488.5 4H8.5 488.5 488.5 488.5 488.5 SICYONTA HREvlROSTRIS SICYONIA TYPICA SICYONIA TYPICA SIGALIONIDAE SIPUNCULUS NUDUS SIRAIUS BICOLOR .. SPELOEOPHORUS ?ONTIFEH SPhAEROSYLLI S SP. SPIONIDAE A SPJOPHANES SF. STENOCIONOPS FlJRCATA COELATA 5THENELAIS !:lOA STROMBIFORMIS AUHICINCTUS SYLLIOAE 0 SYLLIOAE J SYLLIS FERRUGINA TELLINA "P. TEREBELLIDAE A TEREBELLIDtaE. D TEREBELLIDAE G TEREBELLIDES STROEMI THECACEHA AHE&I~N.EP THELEPUS SE10SUS THUNOR SP. TRIvIA CANDIDULA TRIVIA PEDICULUS TURRONILLA SP. F VOLVULELLA PERSIMILIS ME.6N ST ERR MEAN ST ERR !"lEAN 5T ERR ·· ·· · ·• ·· · ··· ··· ·• ·· ·· ·· ·· ·· ·· ·· • ·· ·· · ·· ··· · ···• ·• ·· ·· · ·· ·• ·• ··• ··· ·· · ·· · ·· ·• ··· ··• ·• ·· ·• ·· ·• · . .. FALL SUMMER SPRING wINTER SPECIES ,..S02 ~.O ··· ·· ·· ·· ·· · ·· ·· 0.2 0:2 0.2 O.c O.~ 0.2 0.2 0.2 o.~ 0.2 0:2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 • • 0.2 0.2 ··• • ··• ··• ·· · ·· ·· 0.2 0:2 0.2 0.2 0.2 MEAN 0.2 0.2 0.2 • • • • • · ·· ·• • · • · ··• · ST ERR 0.2 0.2 0.2 • · 0.2 0.2 0.2 0.2 • 0.2 • 0.2 0.2 0.2 • 0.2 0.2 0.2 • 0.2 0.2 0.2 • ·• Appendix 3. (Continued) STATION OVERALL RANK 00 ..... 1.0 2.0 3.0 4.0 5.0 6.0 7.0 8.0 9.0 10.0 11.0 12.5 12.5 14.0 IS.0 16.0 17.0 18.5 18.5 20.0 21.0 22.0 23.0 24.0 25.0 26.0 27.0 28.5 28.5 30.S 30.5 32.0 33.0 34.5 34.5 36.0 31.0 38.5 38.5 40.0 41.5 41.5 43.0 44.U 45.0 47.5 47.5 47.5 47.5 :;2.0 :;2.0 52.0 52.0 52.0 55.0 SPECIES FILOGRANA AXEL~MI LUCONACIA INCERTA SYLLIS SPONGICOLA CHONE AMERICANA SPIOPHANES BOMBYX SYLLIS HYALINA ASPIDOSI~HON GOSNOLDI POLYDORA COLONIA AUTOLYTUS SPa MALACOCEROS GLUTAEUS LUMBRINERIS INFLATA EUNICE VITTATA IoIEBSTERINEREIS .~S 13 CHRYSOPETALIOAE A BRANIA CLAVATA ACANTHOHAUSTORIUS MILLSI OWENIA FUSIFORMIS NEMERTINEA PHYLLOCHAETOPTERUS SOCIALIS EXOGONE DISPAR NEMATODA THOR SPa OPHIACTIS SAvIGNYI OPHIOTHRIX ANGULATA PODOCERUS SPa SIPUNCULIDA LEUCOTHOE SPINICARPA ELASMOPUS SPa B EULALIA SANGUINEA AMPHIODIA PuLCHELLA HYDROTDES CRUCIGERA NOTOMASTUS SPa CHIONE GRUS SYLLIS SP. 0 SYLLIS REGULATA CAROLINAE SYLLInAE LAEVICAROIUM PICTUM BRANCHIOSYLLIS EXILIS PAGURUS CARULINENSIS PROTOMEDEIA SPa A MEGALOMMA 8IOCULATUM UNCIOLA SPa A OPHIOSTIGMA ISACANTHUM POLYCIRRUS CAROLINENSIS CRASSINELLA LUNULATA ELASMOPUS .~S A LEMBOS SMITHI LEMBOS SPINICARPUS INERMIS TELL INA AMERICANA AMPELISCA IGRE~NLHCS AXIOGNATHUS SQUAMATUS MEGALURO~US SP. PHOTIS SPa SERPULIDAE POLYCIRRUS EXIMIUS liINTEfl MS06 SPRING FALL SUMMER MEAN ST ERR MEAN ST ERR MEAN 5T ERR MEAN ST ERR 0.0 45.0 16.0 5.2 9.4 12./j 1.6 21.4 0.6 6.2 6.2 3.0 5.8 1.6 1.8 4.8 6.0 3.0 0.4 29.8 0.2 3.4 0.2 3.8 22.2 5.8 1.0 0.2 0.9 0.2 2.8 13.4 3.0 0.5 1.6 0.4 0.6 24.6 1.6 4.6 9.8 1.0 0.2 3.8 11.3 0.8 0.1 5.3 4155.0 0.8 20.2 2.4 1.6 11.'+ 5.6 2605.6 0.4 13.3 0.1 1.1 '+.5 2.e 2.0 0.4 4.0 1.2 11.0 0.4 7.6 2.8 2.4 0.2 0.6 1.4 1.2 0.4 1.9 0.6 6.5 0.2 4.1 1.7 1.5 0.2 0.4 0.6 0.6 0.6 0.2 0.2 1.2 0.4 0.2 0.2 0.2 1.0 4.4 1.5 0:4 0.4 1.4 0.2 0.2 0.7 • 7.2 5.2 0.8 6.8 2.U 1.8 3.2 0.6 0.6 1.8 0.6 5.2 3.0 1.0 3.0 5.4 1.6 0.2 3.2 0.4 2.6 1.4 0.2 · · 8.a 1.H 3.0 4.2 1.6 21.4 0.4 5.3 3.5 2.0 3.~ 0.8 1.3 2.1 3.1 1• 1 · 2.b 2.3 O.b 5.4 1.0 3.2 1.0 0.4 0.6 0.6 0.4 3.1 2.3 0.4 2.3 3.1 0.9 0.2 1.~ 0.4 1.9 o.~ 0.2 · · 2.0 1.4 0.4 1.2 1.0 1.0 0.4 1.2 1.2 1.4 1.8 4.b 2.0 1.4 1.6 0.5 1.0 1.1 2.7 1.5 1.4 0.5 ·• · ·· ·· 1.0 0.6 0.8 0.2 0.2 5.4 ·• · ··• • · 0.3 0.4 0.8 0.2 0.2 3.4 · · · 0.2 8.8 4.8 4.2 2.8 1.4 · 0.6 2.8 3.6 10.0 2.0 4.2 · · • 0.2 4.4 1.5 1.4 0.'1 1.2 • 0.4 1.1 0.9 5.5 o.a 1.3 · · 1.2 0.6 0.8 0.4 2.0 2.0 4.0 2.6 1.6 0.8 1.6 5.2 0.8 0.7 1.0 3.5 1.4 0.7 0.6 0.5 1.9 1.0 4.6 0.4 2.4 0.5 1.6 0.2 1.2 3:0 1.7 3.8 0.6 1.8 2.0 2.8 0.2 0.1 1.3 · · ·· · O.~ · · · ·· · • 0.2 1.4 1.8 2.8 0.2 0.1 1.3 2.1 2.4 0.4 0.6 0.6 0.4 0.2 1.2 1.0 1.8 0.4 0.6 1.3 · 0.2 0.2 0.4 0.2 0.2 0.2 · ·• 2.4 • 0.8 1.4 0.4 • 18.4 1.8 5.8 3.8 5.2 0.8 11.0 • 1.2 2.6 1.4 1.6 0.4 10.2 1.8 6.0 1.0 3.2 4.6 3.4 3.6 1.2 1.8 2.4 0.6 1.6 1.4 4.0 0.4 1.8 2.4 3.0 1.U · • 16.2 0.6 2.5 0.9 3.0 0.5 7.1 • 0.6 0.1 0.1 1.0 0.2 3.8 1.3 2.5 0.6 2.1 2.0 1.6 1.5 1.0 1.0 0.9 0.4 0.6 0.9 2.8 0.4 O.t! 1.7 2.0 o.e · 1.6 2.4 0.4 0.6 O./j 4./j 1.2 1.4 0.4 0.4 0.4 2.4 1.8 0.4 0.9 0.4 3.0 1.0 2:1 0.4 · ·• · · Appendix 3. (Continued) STATION llvERALL RANK ')H.O se.o 58.0 S8.0 58.0 62.5 b2.~ 62.5 62.~ 00 N 66.0 66.0 66.0 72.0 72.0 72.0 72.0 72.0 72.0 12.0 72.0 72.0 77 .5 77.5 H2.~ 82.5 82.5 82.5 82.5 82.5 82.5 82.5 89.0 89.0 89.0 89.0 89.0 q4.5 94.5 94.5 94.5 94.5 94.5 100.0 100.0 100.0 100.0 100.0 106.0 106.0 106.0 106.0 106.0 106.0 106.0 112.0 ST ERR I' AE~ BOw~ANI L PORTURICENSIS GENOCIDARIS MACULATA PISTIl PALMATA SALMACINA SP. A XANTHIDAE A ARCA IMBRICATA HIATELLA ARCTlCA OPHIOSTIGMA SP. A POLYOORA CAECA ANCHIALINA TYPICA SYLLlS ALTERNATA TURBELLARIA AMPELISCA VAQuRUM CHAMA CONGREGATA LOIMIA MEDUSA LYSIDICE NINElTA MELl TA APPEND I CUL ATA NASSAR INA MINOR PAGURUS SP. SCHISTO~ERINGOS RUDuLPHI SYNALPHEUS TOIINSENDI GONIADIDAE A PARAPIONOSYLLIS SP. AMPHARETE AMERICANA CHEVALI A SP. ERICHSONELLA FILIFORMIS GAMMAROPSIS SP. MUNIDA SP. PERICLIMENES AMERICANUS THOR MANNlt-.lGI VARICORBULA OPERCULATA A HARMOTHOE. .P~ LEPTOCHELA PAPULATA MITHRAX SP. PARAPRIONOSPIO PINNATA PROTODORVILLEA KEFERSTEINI AMPHIPOOA G ARABELLA MUTANS GASTROCHAENA HIANS HYDROIDES SP. A PAGURIDEA SIPUNCULIDA A BOTULA FUSCA MELITIDAE A ONUPHIS i\lEHULOSA ONUPHIS PALLIDULA SEMELE NUCULOIOES CAPHELLA EQUILIBkA CARPIAS BERMUOENSIS CAULLERIELLA KILLARIENSIS LITHOPHAGA 8ISULCATA NASSAR IUS ALBUS ODONTOSYLLIS ENOPLA PILUMNUS SP. ASCOkHYNCHUS CASTELLI O.t; 2.4 2.2 ·• SUMMER SPRING \lINTER SPECIES ~S06 o.~ 1.0 1.3 · · · FALL ,'1EAN ST ERR MEAN ST ERR MEAN ST ERR 1.8 0.4 1.1 0.2 1.2 1.0 0.8 O.d 0.,+ 0.6 0.6 0.2 0.4 3.2 1.4 2.0 1.0 0.2 0.2 0.8 4.2 1.S 0.6 • 0.4 0.8 1.0 0.4 3.4 0.2 0.,+ 0.4 2.0 0.2 0.2 0.4 0.4 0.4 1.2 At.2 3.2 0.6 0.4 0.4 1.0 0.2 2.7 0.2 0.2 0.2 1.5 0.4 2.4 1.0 1.0 0.2 1.6 0.4 1.4 0.4 1.5 0.8 0.5 0.2 1.6 0.2 0.9 0:4 0.6 1.4 1.6 0.4 0.6 0.9 0.6 O.~ 0.2 1.6 0.2 1.2 1.2 1.6 0.4 0.5 0.9 0.4 2.6 2.3 0.2 0.2 1.0 0.2 2.4 0.5 0.2 0.1 0.2 0.2 1.4 0.8 1.'+ 1.2 0.4 0.4 3.0 0.8 3.0 1.0 3.0 0.2 0.2 1.2 3.2 3.2 0.2 0.7 0.6 0.7 1.2 0.4 0.2 1.9 0.6 3.0 0.3 3.0 0.2 0.2 0.4 2.0 1.3 0:8 0.2 3.0 0.4 0.2 2.0 ·· ·· · ·· · ·· · 0.4 0.4 2.2 4.0 2.8 0.9 2.1 1.7 0.4 0.4 0.5 1.6 1.4 1.0 \l.2 1.2 1.At 1.2 0.6 1.2 0.6 1.0 0.2 0.4 1.0 0.4 0.8 0.2 1.2 0.2 0.2 0.6 0.2 0.4 1.8 1.6 0.2 1.2 1.6 1.6 1.6 0.6 1.4 0.9 0.4 0.4 0.2 1.6 0.4 0.2 0.4 1.2 0.4 0.6 0.6 0.4 0.2 1.2 2.8 0.6 0.6 1.6 0.6 O.B 1.0 O.~ 1.0 0:4 0.2 0.6 1.0 1.0 1.8 0.2 0.8 1.4 2.2 1.0 0.2 0.4 0.4 0.9 0.3 0.2 0.2 0.4 1.2 0.5 0.4 0.8 0.2 0.6 0.4 2.6 1.0 0.8 0.6 2.6 0.8 0.4 2.6 0.4 O.~ 0.8 0.8 1.8 0.4 1.1 0.4 0.4 0.2 0.4 0.4 0.2 1.0 1.0 0.6 0.2 1.0 0.8 0.2 · · O.t; · · ·· · ·• · ·· · 0.6 2.4 0.6 0.8 1.0 1.0 0.8 1.2 · • O.H 0.7 1.0 0.4- · · ·· · · ·· ·• • 0.6 1• 7 0.4 0.8 1.0 0.3 0.4 0.7 • · · · · ·• · ·· ··· · · · · · · · · ·· ··• ·· · • ·• · ··· · ·· ·• · · ·• ·· • ··· · ·• · · • · ·· ·· · • 1.2 0.6 0.4 0.2 2.0 0.2 • 2.0 1.6 • 0.2 • • 1.0 0.4 · 1.0 O.B 2.2 · • · • 1.2 0.4 0.4 0.2 0.5 0.2 · ·• · o.e · 1.0 1.1 0.2 0.8 0.4 0.6 1.9 Appendix 3. (Continued) STATION OVERALL RANK 112.0 112.0 112.0 112.0 122.5 122.5 122.5 122.5 122.5 122.5 122.5 122.5 122.5 122.5 122.5 122.5 122.5 122.':> 122.5 122.5 IJ~.O 00 ...., 135.0 135.0 135.0 135.0 135.0 135.0 135.0 135.0 146.0 146.0 146.0 146.0 146.0 146.0 146.0 146.0 146.0 1 /.6.0 146.0 146.0 146.0 160.5 160.5 160.5 160.5 160.5 160.5 160.5 160.5 160.~ 160.5 1bO.5 160.5 160.5 SPECIES OIASTYLIOAE A HEMUS CRISTULIPES PHERUSA INFLATA POMATOCEROS AMEHICANUS AMPHIPODA F A5PIOOSIPHON AL8US BRACHYLJRA CERATONEREIS SP. CHIONE AT~IL RINA GOUL9IA HOLO HURO Ot.A KURTZIELLA LI~ONTEA LU~BRINERIDES ACUTA MALDAN WAE MESOCHAEI0PTE~US SP. MUNIOA PUSILLA POLYDORA SP. SP lOPE TT I 80NtAE SYLLIS CORNUTA XAfIITHIDAE 8 AMP !THOE SP. A ARMANDIA MACULATA EUPHROSINE TRILOBA MAJIDAE A PAGURUS HENDERSON I SCOLOPLOS StJ. A SYLLIDAE L TEREI::lELL IOAE TEREBELLIDAE C AMPHARETE ACUTIFRONS CARIDEA CERATONER I~ MIRAtiIL1S ERICHT O~IUS BRASILIENSIS GLYCERA CAPITATA GLYCERA TESSELATA MITRELLA LUNATA NEPHTYS SQUAMOSA PERICLIMENES SP. PHYLLOCArlIOA PHYLLODOCIOAE PRIONOSPIO CIRRIFERA THYONE PSEUDOFUSUS ALPHEUS NORMAfIoNI AMPELJSCA CRISTOIOES ANOPLODACTYLU$ PETIOLATUS ARCA ZEBRA EUNICIDAE HETEROCRYPTA GRANULATA JAEROPSIS CORALICOLA LEMBOS UNICORNIS LEUCOTHOE SP. A LUMBRINERIOPSIS PARAOUXA LYONSIA SP. ~ MARGINELLA hARTLEY ANUM NEMATONEHE b SP. (i SPRING wINTER MON MS06 ST ERR MEAN ST ERR 0.6 0.2 0.8 1.0 1.8 0.8 0.4 1.0 0.4 0.6 0.9 0.8 0.2 0.4 0.'+ 1.6 0.2 1.6 0.2 0.2 0.4 0.8 2.u 0.6 0.2 0.2 0.4 0.2 0.4 0.8 1.5 0.4 0.2 0.2 0.2 0:4 0.2 0.2 0.2 1.8 0.8 1.2 1.3 0.6 0.4 1.0 0.6 0.6 0.6 · · ·· ··· · · · o.~ · · ·· ··•• · · 0.4 0.2 0.4 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.0 1.4 0.8 0.4 1.0 0.4 • 0.4 • 0.4 0.2 0.6 1.2 0.4 0.4 0.4 1.2 0.2 0.6 1.2 0.2 0.2 0.2 0.7 1.4 1.4 0.6 0.2 1.2 0.9 0.4 0.2 1.4 0.2 1.0 1:2 0.2 0.6 · · · ·· • • 1.2 · ·· · · ·· · 0.8 0.4 1.2 0.7 0.6 0.2 0.7 0.8 0.4 0.2 1.4 0.4 0.4 0.4 0.2 1.2 0.4 · ·· ·• ·· 0.4 1.2 0.4 0.2 1.0 0.2 1.0 0.4 0.2 0.6 0.2 0.8 0.4 0.2 0.2 0.2 0.2 0.4 0.6 0.2 0.2 0.6 0.4 0.4 0.6 0.2 · ·· ·· · ·· • · FALL SUMMER • • · ·· ·· · ·· · · 5T ERR MEAN 5T ERR 0.2 0.2 0.2 0.9 o.a 1.4 0.4 1.0 0.2 MEAN • 0.2 · · • 0.8 0.6 0.4 1.4 0.8 1.4 0.2 0.8 0.8 0.6 0.2 0.2 0.4 0.5 0.2 0.2 · ·· ·• ·· · ··· · ·· · · ·• · ·· 0.8 0.6 0.4 • 0.7 0.2 0.4 0.6 ·• • • • 0.6 0.5 · ·•• • · 0.2 0.2 0.2 0.2 0.4 0.2 0.6 0.2 0.2 0.2 0.4 ··• • • 0.7 ·• • • • 0.2 1.0 0.4 • 0.4 0.2 0.4 o.l:! 0.2 0.2 0.4 1.0 0.4 0.4 1.2 1.2 1.0 0.6 1.4 1.4 1.6 0.4 1.0 1.0 0.4 1.2 0.4 · 0.2 0.2 0.4 0.6 0.2 0.2 0.2 0.4 0.2 0.2 0.7 0.5 o.a 0.4 1.0 1.0 1.4 0.2 0.4 0.0 0.2 0.7 0.4 · 0.2 0.5 1.0 0.4 0.4 0.2 0.6 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.4 0.2 0.2 0.2 0.2 0.8 0.2 0.2 0.2 0.6 1.0 0.8 0.2 • 0.2 0.4 0.4 1.2 0.8 0.4 0.2 0.4 0.2 • • • 1.0 1.0 0.2 0.8 0.2 0.2 0.2 0.2 0.2 ·•• ·· 0.6 0.4 1.0 1.0 0.8 0.6 0.2 0.8 0.2 0.2 0.6 • • 0.2 • • • 1.4 0.2 0.6 • 0.4 · • • • • 0.6 • • Appendix 3. (Continued) STATION OVEKALL RANK 0Cl ~ 16U.':) 160.5 160.5 179.0 1H.0 179.0 179.0 179.0 179.0 179.0 179.0 119.0 179.0 179.0 179.0 179.0 179.0 179.0 119.0 179.0 179.0 179.0 1 19.0 1"79.0 200.5 200.5 2UO.5 200.5 <:'00.5 200.5 <:'00.5 200.5 200.5 200.5 200.5 200.5 200.5 200.5 200.5 200.!:J 200.5 200.5 200.':> 200.5 200.':> 200.5 233.5 233.5 233.5 233.5 233.5 233.5 233.5 233.5 233.5 OPH IURO I uE A S'l'LLlS FERRUGINA TANYSTYLI OAE A AONIOE5 MAYAGUEZEN5IS AXIOTHELLA MUCOSA CAPITfLLIDAE COLOMASTIX SP. DIODORA CAYENEN5lS GALA THEA ROSTRATA GONEPLACIDAE A GREGARIELLA AGHPOILA~C A LYSIAN S~IOAE. ""ARGINELLA LAVALLEEANA OSTRACODA IJELIA MUTICA PLEUROMERIS lR IDENTA TA POLYOORA TETRABRArKHIA PRIONOSPIO CRI5TATA PSEUDEJRYTHOE AM8lGUA PSEUDOMEDAEUS AGASSIZll SICYONIA LAEVIGATA SYLLIS GkACILIS TRYPANOSYLLIS ZEBRA '1EBSTERINEREIS TRIDENTATA ACANTHOHAUSTORIUS 80U5FIELDI ALPHEUS SP. ANCISTROSYLLIS RlblDA BATRACHONOTUS FRAGOSUS acDOTRllOAE 0 CHAETOZONE SETOSA CUMACEA DROMIDIA ANTILLENSIS LEPTOCHELIA SP. LYONSJA SP. !"1AGELONA SP. A MODIOLUS AMERICANUS MUSCULUS LATEHALIS NYMPHOPSIS OuODORSOSPINOSA PHYLLODOCE LONGIPES POOARI<.E OBSCURA PROCERAEA SP. SEROLIS MGRAl'I SICvONIA SP. SPIONIDAE SYNCHELIDIUM AMERICANUM TANAIDACEA 8 ACCALATHURA CRENULAJA ACTEON CANDEi'lS AMPHIPODA B AMYGCALUM SAGITTATUM ANAMIXIS SP. ASTROPECTEN SP. BARLEEIA sp. 0 ENCOPE SfJ. EUNICE FILAMENTOSA SPRING '1INTE~ SPECIES MON 5T ERR 1.2 0.4 0.2 0.4 0.4 0.2 0.4 0.2 0.6 0.2 ·· · ·· ·· ·· ·· · ··· · · · MS06 ·· · ·· ·· ·· ·· O.b 0.6 0.6 0.4 1.2 0.2 0.8 0.2 0.4 0.4 0.4 0.6 0.4 0.2 0.2 0.4 MEAN 5T ERR MEAN ·· ·• ··· · · ·· ·· · ··· · · · ·•• • · ··• · 0.2 0.8 0.2 0.6 0.4 0.2 0.6 0.4 0.4 1.0 • 0.6 0.2 0.2 0.5 · · ·· ·•• · · ·• · ·· · • · · ·• 1.0 0.2 0.4 0.2 0.8 0.2 0.2 0.2 0.2 0.2 0:2 0.2 0.2 0.2 0.6 0.6 0.6 0.4 0.4 0.6 0.2 0.2 • 0:4 0.4 0:2 0.4 0.4 0.2 0.6 0.4 0.2 0.4 0.2 0.2 0.2 0.4 0.2 0.2 0.2 0.4 0.8 0.4 0.4 0.2 0.2 0.4 0.2 0.2 0.2 ~.O ··· · · · ·· · ·· ·•• 0.2 0.2 0.2 0.2 1.0 0.8 0.2 0.5 0.6 0.2 • • • 0.2 0.6 • • • 0.4 ·· 0.2 0.6 0.2 FALL 5UM E~ · ·· ·• ·• · ·· ·· • ·· ·•• 0.6 0.4 0.2 0.6 • 0.4 0.2 0.2 0.4 0.2 0.2 0.4 0.2 0.2 0.2 ·· • 0.2 0.8 · · · 0.2 0.8 ·• • 0.2 0.8 • • 0.4 • 0.2 ST ERR • • ·· · ·• ·• 0.4 0.6 0.8 0.2 0.2 0.4 0.2 0.2 0.2 0.6 0.6 0.2 0.2 0.4 0.4 0.2 0.2 1.0 • • • 0.2 0.3 · · • • 0.8 0.2 0.4 0.4 0.4 0.2 0.4 0.4 0.6 0.2 0.4 0.6 0.2 0.4 0.6 0.2 0.2 0.2 0.2 0.8 0.2 0.6 0.4 0.2 O.it 0.4 · ·· · ""fAN 5T ERR 0.2 1.4 0.4 0.8 0.4 0.4 0.4 0.6 1.0 1.2 0.8 1.2 0.2 1.0 0.4 0.5 0.2 0.2 0.2 0.6 0.6 0.7 o.e 1.2 • • • 0.8 • 0.4 0.4 0.2 0.2 0.2 0.6 0.6 0.2 0.6 0.8 0.4 0.2 0.2 0.6 0.6 0.4 0.2 0.4 0.4 0.4 0.2 0.2 0.4 ·· • • ·• ·• • • • • 0.6 • • • 0.4 • 0.6 0.4 0.2 · 0.2 0.2 0.4 • • 0.4 • 0.8 • o.e ·· 0.2 1.0 0.6 0.4 • 0.4 ·• 0.4 0.4 0.2 • • 0.4 • • • • Appendix 3. (Continued) STATION 0VERALL RANK 00 VI 233.5 233.5 233.5 233.5 233.5 233.5 233.5 233.5 233.5 <:'33.5 <:'33.5 233.5 233.5 233.5 233.5 233.5 233.5 233.5 233.5 233.5 233.5 233.5 233.5 233.5 233.5 233.5 233.5 233.5 233.5 233.5 233.5 233.5 233.5 233.5 233.5 283.5 2!:13 .5 283.5 2l:l3.5 283.5 2~3.5 2l:lJ.5 283.5 2M3.5 283.5 c~3.5 21'\3.5 21i3.5 2·i3.5 2li3.5 283.5 283.5 283.5 21:\3.5 283.5 EURYDICE LITTORALIS GLYCERA SP. B GONIADA MACULATA HYDRO IDES ~P • A LATREUTES PARVULUS LlMA PELLUC lOA LITHOPHAGA ARISTATA LUCAPINELLA LIMATULA LYONSIA tlEANA MAERA CAROLINIANA MAJIOAE MELPHIDIPPIDAE A MICROPHRYS ~ISNELIT~A MITHRAX PLEURACANTHUS MUSCULUS SP. it NEMATONEREIS SP. A NE"'EkTINEA t NEREIDAE NOTOMASTUS LATEkICEUS OCNUS PYGMAEUS OLl GOCHAET A PALICUS SP. PELECYPODA A PELECYPODA tl PENAEOIDEA PISlONE KE,.,OTA POOOCHELA SP. POLT'DORA SP. 0 PsEUDOVERMILIA OCCIDENTAL IS SCOLOPLOS RUBRA slPUNCULIOA E SUdAOYTE PELLUCIDA SYLLlS SP. G SYLUS SP. I XANTHlOAE AMPHILOCHUS SP. AMP ITHO I DAE A APSEUDES SP. B ASTACILLA S~. A AUTOMATE IN A~REV BOOOTRIIDAE 8 CAP lTELL! OAE A CAP lTELLI flAt. 0 CERATONEREIS SP. A CHAETOPTER WilE CHAMA MACEROPHYLLA CHLAMYS 8ENEOICTI CIRRATULlOAE CORBULA CONTRACTA COROPHI WAE CRENELLA DECUSSATA OIASTYLlDAE DIPLODONTA SIM~OFELCUN ERATO MAUGEHIAE ETHUSA MASCAHONE AMEHICANA MElIN ST ERR M£AN ST ERR MEAN · · ·• 0.4 0.0 0.8 0.2 0.2 0.8 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.6 0.2 0.4 0.2 0.4 0.2 0.4 0.6 0.4 O.l:! 0.4 · ·· · • 0.2 0.8 0.2 0.8 0.2 0.6 0.2 0.8 0.2 0.6 0.2 0.6 0.4 0.0 0.4 0.0 0.2 0.4 0.2 0.6 0.4 0.4 0.2 0.4 ·· · ·· ·· · ·· ·· ·· ·• ·· ·· · · • • · ·· ·· ·· ·· ·· · · ··· · ·· ··· · · · ··· · ·· ·· 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.4 0.2 0.2 0.2 0.4 0.2 • 0.6 0.2 ··· · ·· ·· 0.4 0.2 0.8 0.8 0.0 0.0 0.2 0.8 0.4 0.2 0.2 0.6 0.2 0.2 0.4 0.2 0.2 0.4 0.4 U.2 0.4 0.2 0.4 0.2 0:2 0.4 0:2 0.2 0:2 0.2 ·· · ·· ·· ·• 0.2 0.4 0.2 0.4 0.6 0.2 0.6 0.2 · 0.2 · ·· ·· ·· ·· ·· 0.4 0.4 · FALL SUMMER SPRING wINTER SPECIES MS06 0.2 · ··· ·· · ·· ··· 0.4 0.4 · 0.6 · · · ·· ·· ·· ·· ··· · · ·· ·· · · ·· 0.2 0.2 ST ERR o.~ • • 0.2 0.4 0.2 0.2 • 0.2 0.2 0.2 0.4 0.4 0.4 0.2 0.2 0.2 · ·• ·· ·· ·· ·•• · · ·· · · · ··· ··· · ·· · 0.6 0.0 0.6 0.6 0.4 0.2 0.2 0:2 ·· · 0:6 0.6 0.2 0.6 0.4 0.6 0.2 0.4 0:6 0:4 0.4 0.4 · ·· · "1EAN ST ERR 0.2 0.2 · 0.2 0.2 0.2 0.2 • 0.2 0.8 0.4 0.6 · · · · 0.2 0.8 0.2 0.6 · 0.4 0.4 0.4 0.6 0.2 0.2 0.4 0.2 0:2 0.2 0.2 0:5 0.2 • • • D.!:! · · · · · · · · · · · ·• · 0.2 0.2 0.2 0.2 0.2 0.2 0.8 0.0 0 ... 0.6 0.4 0.2 0.6 0.6 0.6 0.4 0.2 0.0 0.4 0.2 0.6 0.2 • 0.4 0.4 • • 0.2 0.2 0.2 0.2 • 0.6 0.6 ·• Appendix 3. (Continued) ~TA ION UVERALL RANK 00 0- 283.5 283.5 283.5 2lj3.5 2"3.5 283.5 <:'1'3.5 283.5 283.5 2,,3.5 2a3.5 283.5 283.5 283.5 283.5 283.5 2H3.5 283.5 21-:13.5 283.5 283.5 283.5 2H3.5 <:,H3.5 283.5 283.5 283.5 <:'83.5 283.5 283.5 283.5 2tl3.5 283.5 2H3.5 283.5 2tU.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 SPECIES GEUKfNSIA DEMISSA GONODACTYLUS IN DE~B HI PPOL YT IDAE HIPPOMEDOIII SPa A HY80SCOLEX LONGISETA LAONICE CIRRATA LEM80S SPa LEPTOGNATHIIDAE A LEUCOTHOIOES POTTSI LYSIANOPSIS AL8A MANGELIA RUGIHIMA MARGINELLA ROSCIDA MUREX FLORIFER o ILECTUS NEREIS SPa NEREIS SUCCINEA NOTOMASTUS AMERICANuS OPHELIA DENTICULATA PARAONIOAE PELECYPODA C PER I CL EN ~P S IR IDESCENS PHYLLOOOCE SP. PILUMNUS SAYI PINNUTHEHIIJAE POLYCIRRUS tXIMIUS OUbIUS PONTOGENIA SERICOMA PORTUNUS 5f-'. PROCESSA SP. SEMELE PURPURASCENS SIPUNCULIOA B SPIO SPa SYLLIOAE C SYLLIOAE I SYLLIS SPa F TRACHYCARIS ~UTCIRTSER VERTICOROIA ORNATA WEf3STERINEREIS SPa A ACANTHOCHITOIIIA PYGMAEA AEGIOAE A ALPHEUS SPa B AMPHIPOOA E AR8ACIA PUNCTULATA ARICIOEA CERRUTI ASTHENOTHAEHUS HEMPHILLI it ATYS SANDERSON I 8000TRlIOAE A 8RACHYCYTHARA BARBARAE LYONS CAOULUS QUAORIDENTATUS CAPITELLA CAPITATA CERADOCUS SPa CERATONEREIS VERSIPEOATA CHAETOPTERUS VARIOPEDATUS CHIONE CANCELLATA CYCLOPECTEN NANUS DENTALI IOAE it OIPLODONTA PUNCTATA WINTEF; SPRING MEAN ST ERR 0.6 0.4 0.2 0.4 0.4 0.4 0.2 0.2 0:2 0.2 0:2 0.2 ·· · ·· ··· ·· ·· · ·· · 0.2 ·· · ·· ··· ··· · ·· · 0.2 0.2 0.2 O.l:' 0.2 0.2 0.2 0.4 0.2 0.4 0.4 0.4 0.4 · · ·• · 0.4 MS06 • • • 0.2 0.4 0.2 0.2 • 0.4 SUMMER MEAN ST ERR MEAN ST ERR · ·· ·· ·· · ·· · ·· ·• ·· ·• ·· · · ··• ·· ·· · ·· ·• · ·· ·· · · ··• · · ·· · ·· ·· ·· ·· ··· · ·· ··•• ·· ·· ··· ·· · ··· ·· ·· ··• • ·· ·· ·· · · ·• · ··· ·· ·· · 0.6 0.4 0.2 0.2 0:4 0.2 0.0 0.6 0.6 0.6 0.6 0.6 0.2 0.2 0.2 0.2 0.6 0.6 0.2 0.2 0.2 0.2 0.2 0.2 Ll.4 • 0.4 0.2 0.4 0.4 0.2 0.4 0.1+ 0.4 0.2 0.4 0.4 0.2 0.4 0.2 0.2 0.2 0.2 0.2 0.2 • 0.2 0.2 · 0.4 0.2 • 0.4 0.2 ·• · · • 0.4 0.2 • 0.2 0.2 FALL 0.2 0.4 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.6 0.2 0.2 0.2 0.6 0.2 • ·· · 0.4 · · 0:4 0.4 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 ·· ··• ·· ··· · • · ·· ·· ·· 0.2 0.2 0.2 0.2 0:2 0:2 0.4 0.2 0.4 0.2 0.2 · · ·· ·• ·· 0.2 MEAN · · ST ERH 0.2 0.6 • 0.2 0.6 0.2 0.6 0.2 0.6 0.4 0.2 0.2 0.6 0.2 0.6 0.2 0.2 • 0.4 • 0.2 • • 0.6 · · • 0.4 0.6 0.2 0.2 0.2 0.4 • 0.4 0.2 0.2 0.2 0.2 0.2 0.6 0.4 0.2 0.2 0.4 0.2 0.2 0.2 0.2 • 0.6 • 0.4 · · • · • • • 0.6 • 0.6 • 0.4 • 0.2 · • • • • 0.2 0:2 0.4 • • ·· 0.4 • Appendix 3. (Continued) STATION O~ERAL 00 -.I RANK 353.5 353.5 353.5 3':>3.5 353.5 353.5 353.5 3:,3.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 353.5 .$53.5 353.5 353.5 353.5 353.5 SPECIES DROM I I DIl,E EBALIA CARIOSA EBALIA STIMf'SOt'lI ENGINA TUR8INELLA EUCIDARI~ TkIBULOIDES EUNICE AI"TEN1'lA TA GASTROPOOA A GLYCERA ASOL IPA~ HAPLOSCOLOPLOS ROBUSTUS HpIONIDAE H PPOLYTIDAE it HYALINA AVENA HYOROIDES UNCINATA INACHOIDES FOkCEPS INACHOIDES FO~CEPS it ISCHNOCHITONIDAE A LUM8RINERIS SP. LUM8RINERIS COCCINEA LUMBRINERIS LATREILLI UONSIA HYALINA MAERA SP. A MAERA WILLIAMSI MICROPANOPE SCULPTIPES MYSIDACEA NANOPLAX XANTHIFORMIS NEREIS ACUMI"'ATA NICOMACHE TRISPINATA OPHIOCNIDA SCABRIUSCULA OPHIOPHRAGMUS SP. OPISTHOBRANCHIA 4 PARAPINNIXA BOUVIERI PARAPIONOSYLLIS LONGICIRRATA PELECYPODA PENAEIDAE PERICLIMENAEUS SP. PHERUSA EHLt:RSI PISTil, QUADRILOBATA POECILOCHAETUS S~. POTAMJLLA R£NIFORMIS PROCESSA HEMPHILLI it PSEUDOCHAMA RADIANS PSEUDOMEDAEUS AGASSIZII It PTEROMERIS PE~LAN PYRUNCULUS CAELATUS RILOARDANUS LAMINOSA SCOLELEPIS 51-'. S SEMELE BELLASTRIATA SPEOCARCINUS CAROLI~ENSr SPHAEROOORIDIUM CLAPAKEDII SPIO SP. A STHENELAIS BOA STROMBIFORMIS BILINEATUS SUTUROGLYPTA IONTHA SYLLIDAE D SYLLIDAE N ··· · ST ERR ··· • 0.2 0.2 0.4 0.2 0.2 0.4 0.2 0.4 0.4 0.2 0.2 0.2 0.2 0.4- 0.4 0.2 0.4 0.2 0.4 0.4 U.4 0.4 0.4 0.4 0.2 0.2 0.4 0.2 0.4 0.2 0.4 0.4 0.4 0.2 0.4 0.4 · · ·· ·· · · · · ·· ·· ··• ··• ··• · ·· · 0.4 0.2 SUMMER SPRING wINTEf< MEAN ~S06 · · ·• ·· · · · ·• • ·· ··• ·· ··· ·· ·· · 0.4 0.2 0.4 0.4 0.4 0.2 0.2 0.2 0.2 0.2 Mt:AN ST ERR 0.2 0.2 · ··•• ··• · · ··· · ··• • • ·· ·· ··• ·· ·• ·• 0.2 • • • · · ·· ·· ·• ··• · ··· ·· ··· ·· · ··• 0.2 0.4 0.4 0.2 0.2 0.2 0.4 0.2 0.4 0.4 0.2 0.2 0.2 0.2 0.2 0.2 • 0.4• • 0.2 ··· ··• · 0.2 • • 0.2 • · ·• ·•• ·· · · 0.4 0.2 0.2 FALL MEAN ST ERR ·· · ··• ·· · ·•• ·· ·• · ··• ·· ·• ·· ··• ·• ·· · ·• ·· ·• ··•• · ·•• ··• ·· · · ·• ·••• ·• 0.2 0.2 0.2 0.2 0.2 0.2 MEAN ST ERA 0.2 0.4 0.4 0.2 0.4 0.4 0.2 0.2 0.2 • 0.2 0.2 0.2 0.4 • 0.4 ·• • · ·• · ··• 0.2 • • 0.4 0.4 · ··• ·• 0.4 0.4- 0.4 0.4 0.4- 0.2 0.2 0.2 0.4 0.4 • 0.4 0.2 0.4 • 0.4 0.4 0.4 0.4 0.2 0.2 0.2 ·• 0.2 • • • 0.2 0.4 ·• 0.2 • • 0.2 • ·· 0.2 0.2 • • 0.2 • • 0.2 • 0.4 • • • 0.2 · · ·• 0.4 • • • • • • 0.2 • • 0,.2 0.2 • 0.2 • • • • 0.2 • • 0.2 0.2 • 0.2 • 0.4 • • • • 0.2 0.2 0.4 • 0.4 0.2 • 0.2 • 0.2 • 0.2 • • 0.2 • 0.2 Appendix 3. (Continued) M506 STATION ()VERAll RANK oc oc 3':>3.5 353.5 353.5 353.S 353.5 353.5 3S3.5 353.S 353.5 353.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 4'16.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 446.':> 4~6.5 496.5 496.5 496.5 496.5 496.':> 496.5 496.5 496.5 SPECIE.S WINTER ME/1N SYNAlPHEUS SP. SYNElMIS Alt:lI/';I TEREBElLlOAE A THARl'X MARIuNI TIkON TROPAI<IS TRAVISIA PARVA TRICHOPHOXUS FlORIDANUS TRIVIA MALTt:lIANA TlJRBONILLA ABRUPTA TURBONILLA INTERRUPTA ALPHEIOAE AMBLYOSYLLIS FORMOSA AMERICARDJA MEDIA AMPHILOCHIDAE AMPHIURA FI8ULATA Ar-1YGDALUM PAPYRIUM ANACHJS LAFRESNAYI ANOMIA SIMPLEX ANOPLODACTYLUS INSIGNIS APANTHURA ACIF ~GA ARCIDAE * ARCOPSIS ADAMS I ARENE TRICARIIliATA ASHC ILLA LAUFF I ASTHENOTHAERUS HEMPHIlLI AUSTRAEOllS CAT INA 8ARBATIA DOMINGENSIS BERGHIA Sil. I::lODOTRIIDAE BODOTRIIOAE C CADUlUS TETHASCHISTUS CAECUM COOPERI CALOCAklS HIRSU1IMANA CAPIfELLlDAE B CAP ITElLl OAE E CAPIfElLlDA£ G CERAPUS TUBUlARIS CERITHIOPSJS METAXAE TAENIOlATA CERITHIUM ATHATUM CHAETOPLEURA APICUlATA CHAMA SP. CHRYSOPETALIDAE CIROLANA PARVA ClkRATULUS SP. CIkRATULUS SP. A CIRRIFORMIA FILIGERA CLYPEASTER 5.... COLOMASTIX SP. I:l CREN£LLA OIvARICATA CREPIDUlA ACULEATA CRONIUS SP. CRON IUS RUAER CUMINGIA TEllINOIDES CYCLOPECTEN SUNA~ * OELECTOPECTE/'j SP. it · ·• ·• ·· ·• ··• ·· ·· ·• ·· o.c ·· ·· ··• ··• ·• ·· ··· ·• 0.2 SPRING 51 ERR · ·· • · ··• ··· ·· ·· · ··· ·· ·• ··· ·· · ·• · ··• ·• · 0.4 0.2 • 0.2 0.2 0.4 0.2 0.4 0.2 0.2 0.2 O.C 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 r-1EAN · · ··• ··• ··• ·• ··• ··•• ·• ··• ·• • 0.2 0.4 0.2 0.2 0.2 5T ERR ·• · 0.2 0.4 0.2 • • • 0.2 • • • · ·• ·• ·· ·• ·· ·· · ··• ·· • ··· ··• ·•• 0.2 0.2 0.2 0.2 • 0.2 0.2 0.2 • • • ··· • 0.2 • 0.2 • 0.2 0.2 0.2 0.2 • 0.2 0.2 0.2 0.2 · • • FALL ~EM US ·• ·• MEAN · 5T R~E · • 0.2 • 0.2 0.2 0.2 • 0.2 • 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 • 0.2 • • 0.2 · · ·•• ·• ·• ··• • • ··• ·•• • ·• ··•• ·· ·• ·•• 0.2 0.2 0.2 0.2 0.2 • • 0.2 ·• · ·••• ·•• 0.2 0.2 0.2 • • • • • • • • • • • · ··• ··• ·••• 0.2 • 0.2 0.2 0.2 • • • • MEAN 0.4 0.2 • • • Sf ERR 0.4 0.2 • • • • 0.4 • • • 0.2 • 0.2 • 0.2 • 0.2 0.2 • 0.2 0.2 • 0.2 0.2 • 0.2 • 0.2 • 0.2 0.2 • • ·• • • • • 0.2 • • • • 0.2 0.2 • 0.2 • • ·• 0.2 • 0.2 • • • • • 0.2 0.2 · ··• · • · • • • 0.2 • • • • 0.2 0.2 • 0.2 • • 0.2 • • 0.2 • • 0.2 • • • 0.2 • • • • Appendix 3. (Continued) ~TA ION OVERALL kANK ~.OY4 oc 'C 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 490.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 wINTEI1 SPECIES NAE~ DENTALIUM OCCIDENTALE DENTALIUM OCCIDENTALE * DIODOPA LISTEi-1I DIODORA LISTERI * DIOPATRA TRIDENTATA DONDICE OCCIOENTALIS ELASMOPUS SP. D EULALI A SP. EULALIA MACROCEROS EUNICE SP. EUNICE SP. t3 EURYDICE PIPERATA FLABELLIGERA SP. GASTROPODA E GLOTTIOIA PYRAMIDATA GLYCERA SP. C GLYCYMERIS SP. * GONIADIDI:.S CAROLINAE HARMOTHOE .P~ HESIONURA SP. A HYALINA ALBOLINEATA HYDROIDES SP. 0 HYPSICOMUS PHAEOTAENIA IRIDOPAGURUS DISPAR KURTZIELLA ATROSTYLA LASAEIDAE LEIOCHRIDES PALLIOIOR LEPTOCHELA SP. LILJEBORGIA SP. A LILJEBORG{A SP. 8 LIMEA BRONNIANA LINGA AMIANTUS LISSA BICARINATA LUCIFER FAXONI LUCINA NASSULA LYROPECTEN SP. * LYSIANASSIDAE. LYSMATA SP. MACROCALLISTA * MACULATA MACROCOELOMA SP. * MACROCOELOMA CAMPTOCEkUM MACROCOELOMA TRISPINOSU"I MARGINELLA SP. MARGINELLA SP. * MARGINELLA AUREOCINCTA MARGINELLA VIRGINIANA MARG INELLI DAE * MEGALOBRACHIUM SORIATUM MESORHOEA SEXSPINOSA MICRODEUTOPUS MYERS I MICROPANOPE SP. MICROPANOPE NUTTINGI MICROPHRYS SF. MICROPHRYS SP. * MODIOLUS MODIOLUS SQUAMOSUS · ·· · ·· ·· 0.2 0.2 0.2 0.2 SPkI!'lG MEAN ST I:.RR MEAN ST ERi< · ·· · · ·· · · ·• · · ··· · ··· · ··· ·· · ··· ··· · ·• · ··• ·· ·· ··· · ··· ·· · ·· ·· ·· ·•• · ··· ·· ·· ·· ··· ··• ·• · ··· · · · ·• · ···• ··• ··• ·· · ··· ··· · · 0.2 0.2 0.2 0.2 0:2 0.2 0:2 0.2 0.2 0.2 0.2 0.2 0.2 0:2 0.2 • 0.2 0.2 0.2 0.2 0.2 · ··· · ·· · ·· ··· · · ·· · O.e · · 0.2 0:2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 FALL SUMMER ST ERR 0.2 ·· MSU6 · ·· · 0.2 · 0.2 0:2 · · · ·•• · ··· ··• · ··· ·· ·•• ·•• ·· ··· ··· ··• ··· ·· • 0.2 0.2 0:2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 ··• · · ·• · · • ··• ·· · · ··• 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 ·• 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 · ·· · ··· · · 0.2 0.2 MEAN 0.2 · 0.2 ·• ··• ··• ·• 0.2 0.2 · · ·• ·• ··• • ·•• ST ERR 0.2 • 0.2 • ·• 0.2 • • • • • · 0.2 • 0.2 0.2 0.2 • 0.2 0.2 0.2 0.2 0.2 0.2 • 0.2 • 0.2 • • • • 0.2 · • 0.2 • 0.2 • · · • • Appendix 3. (Continued) STATION OVERALL RANK .. Y6.5 .. <.;6.5 496.5 4<';b.5 496.5 496.5 496.5 496.5 496.5 496.5 4'/6.5 496.5 496.5 4'/b.5 \Cl 0 496.5 496.5 496.5 4Y6.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 '~96.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 ,+96.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 496.5 SPRING wINTER SPECIES MUN IDA IkRASA "'USCULUS NIGER NAINERIS SP. NANNOSQUILLA SP. A NATTCA SP. A NEMERTINEA C NEOPONTONIDES dEAUFORTENSIS NEPHTYS INCISA NOTOHRYON WAHOI NOTOCTRRUS SPINIfEHUS NOTOHASTUS SUDO~I EH NOTOMASTuS LOBATU5 NYMPHONIDAE A NYMPHOPSIS SP. A OLIVELLA BULLULA ONUPHIS SP. ONUPHIS EREMIT A OPALI A PUM ILIO OPHELIIDAE OPHIOLEPIS ELEGANS OPHIONEREIS RETICULATA OPHIOSTlbM4 SP. OPHIURO IDEA A ORBINIIDAE ORBINIIDAE ti ORCHOMENE 'SP. A OSACHILA TUBEROSA OXYUHOSTYLI~ SMITHI PA6URTSTES TORTUGA£ PAGURUS HENDERSON I it PANANTHURA FORMOSA PAPYRTDEA SEMISULCATA PAREULEPIS SP. PARTHENOPE FRATERCULUS PARV I TURl:lO IOES INTERRUPTUS PERICLIMENES SP • PHIDIANA LYNCEUS PHORONIS ARCHITECTA PHTISICA MARI"'A PHYLLODOCE ARENAE PHYLLODOCE CASTANEA PHYLLODOCIDAE C PILUMNUS DASYPODU5 PINNIXA SP. PINNOTHERES MACULATUS • PISANIA TINCTA PITHO LHERMINIERI PLICATULA GIBBOSA PODOCHELA SE~IL CARG POL YCHAET A F POL YDORA SP. C POLYDORA SP. E POLYNOIDAE A POLYPLACOPHOHA A PORTUNIDAE 0 • "'tAN 5T ERR o.c O:c 0.2 ·· ·· · ·· ··· ·· · ·· 0:2 · ··· · ·· ··· ·· ·• O.C 0.2 O.~ 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0:2 0.2 0.2 o.~ ·· ··· ·· · · · · ··• ·· ··· ··· ·· ··· ·· · ·• ·· ··· ·· ··· ··· 0.2 0:2 0.2 0:2 0.2 0.2 0.2 0.2 • ·· MS06 0.2 0.2 SUMMER IoiEAN 5T ERR ·· ··· ··· ·· ·· ··· · ·· · ··· ·· ··· ·· · · ·• · ··· ··· ·· · 0:2 0.2 ·· ·· ·· ·· · ·· 0.2 0.2 0.2 0:2 0.2 0.2 · ·· · ··· ··· ·· ··• ··· · ··· ··· ·· · 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 MEAN · 0.2 · · ·· FALL ST ERR · · · ··· ·· O.C 0.2 0.2 0.2 0.2 • 0.2 0.2 0.2 0.2 ··· · · ·· · ··· · ··· o.c · 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 ·· · ·• · ·· · · ST ERR 0.2 0.2 ·· MEAN ·· · ·· · ·· · · 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 · · 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 · 0.2 0.2 • 0.2 0.2 0.2 0.2 0.2 0.2 • 0.2 0.2 0.2 0.2 ·• 0.2 0.2 0.2 · · 0.2 0.2 0.2 · · • · · · Appendix 3. (Continued) MS06 STATION L A~EVu RANK 5.b~4 496.5 4'16.5 496.5 4~6.5 446.5 446.5 496.5 496.5 496.5 496.5 496.5 4'-'b.5 496.5 496.5 496.5 4'i6.5 \0 ..... 496.5 4qb.5 4Y6.5 496.5 496.5 496.':J 496.5 496.5 496.5 496.5 4'i6.5 496.5 496.5 496.5 496.5 496.5 49b.':> 4~6.5 496.5 ... 96.5 446.5 496.5 496.5 496.5 49b.5 4Y6.:' 496.5 496.5 4'16.5 496.5 /-ION PuRTUNUS ORDWAYI PORTUNUS OROWAYI -l> PSAMMOLYCE CTENID0PHOHA PYLOPAGUkUS SP. -l> PYLOAGU~S HOLTHUISI RISSOINA SP. A RUDJLEM801 ~S NAGLEI SABELLARIJDAE SABELLIDAF -l> SALMON[US IN A~TRO SCALIBREGMA [I'<FLATUM SIPUNCULIOA C SPATANGOIOEA SPELOEOPHORUS NODOSUS SPIONIDAE a SPIROGLYPTUS SIRALUGE~RI SQUILLA hEPTACANTHA STENUCIONOPS FURCATA COELATA 5 TENOPLEUS T1::S SP. A STENORHYNCHUS SETICORNIS STENOTHOE SP. A STHENELAIS SP. SYLLIOAE K SYLLIDES SF'. SYLLIS SP. SYLLIS SP. tl SYLLIS SP. E SYNALPHEUS MINUS SYNAPTIDAE TELLINA SP. TELLINA SY8ARITICA THALENESSA SP. A TRACHYPENAEUS CONSTRICTUS TRICHOBRANCHUS GLACIALIS TRICHOPHOXUS EPISIOMUS TRICOLIA THALLASSICOLA TRIVIA CANDIOULA TRIVIA NIX TRYPTON CARNEUS TURBONILLA IL HP~EH UHLI AS LIMBA TUS UNCIOLA SP. UNCIOLA SPICATA VERMILlOPSIS ANNULATtI VE~IL UM GEMMA TUM XANTHTDAE. it XANTHTDAE C SPRING wINTER SPECIE') · ·· ·· ·· ·· ·· · • · · ·· ·· · ·· ·· ·· ST um 0.2 0:2 0:2 0.2 0.2 · ·· ·· ·• ·· ·· · ·· · ·· ·· · ·· ·· ·· 0.2 0.2 0.2 ~.u 0.2 0.2 ~.U o.~ 0.2 U.2 U.2 0.2 0.2 0.2 0.2 o.~ 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 ~.o 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 MEAN ·· ·· ·· ··• • ST ERR • · ·• ·· · ···• ··• · ··· ·· ·· ·· ···• ··• ··• ·· · 0.2 0.2 0.2 0.2 0.2 0.2 • 0.2 • ·• 0.2 • 0.2 U.2 0.2 0.2 0.2 0.2 0.2 0.2 ··• ·· • ··• ·• ·· ·• ·• ·· ·• · FALL SUMMER MEAN ·· ··• · ·•• ···• · ·• ·• ·•• ·• • ·• · · · · ··• ·· ST ERR ·•• ··•• • · ··· ·• ··• ··• ·• • ·• · · ·• ·• ··• 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 MEAN o.~ • • · ·• 0.2 0.2 • • • 0.2 • • 0.2 0.2 ·· ·••• • • • • 0.2 • • 0.2 • · • 0.2 ST ~RE 0.2 • • • 0.2 • • 0.2 • 0.2 • • 0.2 • 0.2 • • • • • • • • • 0.2 • · · ·• 0.2 0.2 • 0.2 0.2 • • • • • • • Appendix 3. (Continued) STATION L A~EVU 1 .0 2.0 3.0 4.0 5.0 6.0 7.0 0.0 ':1.0 10.0 11 .0 12.0 13. U 14.0 15.U 16.0 '4:l N ST ERR N~E" 11.0 IR.O IQ.O 20.0 21.':> 21.5 24.0 24.0 24.0 2,.,.0 ?7.S 27.5 2<).0 30.0 31. ':> 31.5 33.5 33.5 35.0 36.0 3t1.5 38.5 3e.5 3!i.5 42.0 42.0 42.0 44.0 45.5 4':'.5 <;0.0 .... 0.0 ~o. 50.0 '10.0 "0.0 50.0 ')7.5 ::;7.5 PISTA PALMATA SPIOPHANES bOMBYX ERICHTHONIUS SP. A SYLLIS SPONGICOLA ONUPHIS PALLICULA PHOTIS SP. OWENIA FUSIFORMIS PHOTIS LONGICAUDATA MESOCHA T P E~US SP. AMPHARETE ACUTIFkONS CRASSINELLA LUNULATA UNCIOLA SP. A OMJPH I S NERULOSA BOWMANIELLA PORIOklCENSIS AMPELISCA VADORUM AMPHIODIA PULCHELLA MUNIOA PUSILLA AMPHAPETE AMERICANA NE"'1flTODA AGLAO~HAMUS VERRILLI AXTOTHELLA MUCOSA NEMERTINEA APSfUDES SP. 1:3 PAPAPRIONOSPIO PINNATA PHOTOHAUSTORIUS wIGLEYI RILOAPDANUS LAMINOSA PHTISICA MAHINA SPIONIQAE B EURYDICE PIPERATA PRIONOSPIO CRISTATA HARMOTHOE SP. A MICROPANOPE SP. ~ AMYGDAl.UM SAGITTATUM EULALIA AENIU~NAS EUNTCE VITTATA TRICHOPHOXU~ FLOklOANUS ACAIIJTHOHAllSTORIUS MILLSI BATRACHONUTUS FRAGUSUS LAONICF. CIRkATA STHENEL/lIS [jOA ARMANOIA MACULATA EUCEHAMUS SUGNOLEA~P XANTHIDAE g GLYCERA CAPllATA 4UTOMATE IN A~REVf LOIMIA Mf..DUSA AMPfLISCA CHISTOIDE5 AMPEL TSCA VENETIENSIS GAMIAAROPSIS SP. bLOTTlnIA PYRAMIDATA PHYLLODOCE SEPIG~OL PSAM~OLYCE CTENIDOPHOkA THYONE PSEUDOFUSUS ASYCtiIS CAROLINAE. ERICHT ONIU~ BRASILIENSIS 20.3 17.7 4.0 20.3 16.2 2.1 2.3 3.7 2.U 2.3 3.7 2.0 1.0 1.0 1.0 1.0 0.3 2.0 0.3 0.3 2.0 0.3 0.3 0.3 2.7 2.7 1.3 0.7 0.3 0.3 2.7 2.7 1.7 1.7 0.3 0.3 · · · ··• · · · · · · ·· ··· ·· ·• ·· ·• ·· ·· · SUMMER SPRING WINTER SPECIES K~Ak OSOI · · · ··· · · · · ·• ·•• ··· ·· · ·· ST ERR MEAN ST ERR MEAN 248.4 48.8 20.0 7.6 20.0 15.8 7.4 9.6 10.2 1.4 7.8 7.0 4.8 4.8 4.8 4.6 6.2 1.8 5.0 0.8 4.0 2.2 3.4 3.2 US.8 18.9 15.5 7.6 10.4 12.0 5.5 4.1 5.5 0.9 4.3 1.5 1.8 2.1 2.0 1.3 6.2 1.8 4.8 0.4 1.6 1.0 1.2 1.0 14.0 11.6 9.8 7.4 4.7 8.0 0.6 5.0 0.4 10.0 0.8 0.6 2.7 0.6 0.4 2.6 1.0 2.9 1.2 1.4 0.6 0.9 1.2 0.6 0.5 0.4 MEAN • 0.8 · 1:0 0.3 1.0 0.3 4.4 0.8 0.4 2.6 2.0 3.2 3.4 2.6 1.6 1.6 1.4 1.2 1.4 0.6 2.0 1.5 1.4 0.7 ·• 1.2 0.6 0.2 0.4 1.2 1.4 1.0 1.4 0.4 1.6 0.8 1.4 1.0 0.4 0.2 0.2 1.2 0.9 0.6 0.6 0.2 1.6 0.8 0.9 2.0 1.7 · 1.1 ··• ·· · · 1.7 FALL · · ·• 11.2 8.0 0.6 • 6.0 3.0 0.6 6.2 1.2 1.0 1.2 2.4 1.4 1.2 · · 3.2 2.2 0.6 1.6 0.8 1.4 5.0 0.6 0.2 1.8 3.4 0.6 0.8 • 5.8 1.7 0.4 4.1 1.0 0.6 1.0 1.3 0.6 1.0 • • 2.2 1.8 0.6 1.0 0.6 0.9 3.1 0.4 0.2 1.3 1.9 0.6 0.6 • 2:2 0.8 0.4 3.2 0.6 1.0 1.0 ST ERR · 1.6 0.4 8.0 0.4 • 1.0 0.5 0.4 1.8 0.2 1.0 0.3 • 0.8 0.6 • 0.5 0.2 0.8 0.8 1.0 0.8 0.8 0.6 0.4 0.8 0.5 0.8 0.6 0.6 0.2 • 0.2 • • 1.0 1.8 • • 0.5 1.6 • 0.4 0.2 0.4 0.6 0.4 0.6 0.6 0.8 1.4 1.2 0.4 1.0 0.4 0.6 0.5 1.2 1.0 0.2 1.0 0.2 1.0 0.6 0.2 0.6 0.6 2.0 0.2 0.4 0.8 1.4 0.2 0.6 • · ·· · · • • • • 0.4 0.4 1.0 2.6 0.2 0.2 1.0 2.6 1.3 0.2 0.2 0.6 1.0 1.2 o.a 0.7 0.2 0.4 0.2 0.6 0.2 0.4 1.0 0.8 • • O.~ 0.6 1.2 0.4 • 0.4 1.0 0.4 0.4 0.4 · 0.2 0.4 0.8 • · • • • ·• ·• 0.4 • 0.4 0.7 • • • 0.2 • • • Appendix 3. (Continued) STATION OVERALL KNA~ SPECIE.S wlNTEH 57.5 57.5 57.5 57.5 57.5 57.5 66.5 6 .~ \Q w 66.S 66.5 66.5 h6.5 66.5 66.5 66.5 h6.5 77.0 77.0 77.0 77.0 77.0 71.0 71.0 77.0 77.0 77.0 77.0 d~.5 89.5 R9.5 119.5 119.5 ~9.5 B9.5 ~9.5 "\9.') tl9.S tl9.5 ~9.5 H9.5 89.5 lll3.5 103.5 103.5 103.5 103.5 103.5 11l).5 103.5 103.5 103.5 103.5 103.5 103.5 103.5 HOLOTHURuIDEA LEPTOCHELA fJAFULATA NEPHTYS ASCMAUQ~ NOTOMASTUS SP. PAGURIDEA XA"lTHIDAE A AXIOGNATHUS SQUAMATUS BRACHYURA OIOPATRA CUfJREA GLYCERA SP. C GON IAOA rvlACULA TA GONIAOIDAE A GONIADIDES CAROLINAE NOTOMASTUS AMERICANUS OPH I IJRO I DE A POMATOCEROS 5UNACIRf~A CHEVALIA SP. CHONE AMEPICAi'lA LU~BRINERIS COCCINEA PAGURUS Sf'. PARTHENOPE FRATERCULUS PRIONOSPIO SP. l:l PROCESS A SP. PROTOnORVILLEA KEFERSTEINI STO"1ATOPOOA A TEL I~A SYRARITICA THARYX MARIONI ASPIDOSIPHON GOSNOLDI ASTROPECTEN SP. CALOCAPIS Al'iAMITUS~IH CHIONE GRUS CLYTHROCERUS SUL ISUP~E GLYCERA SP. 1:3 LEMA05 SIN~UCINU MALDANIDAE NASSAR IUS ALBUS NOTOPYGOS ATI~ RC OLIGOCHAETA PAGURUS HENDERSON I PELECYPOOA TEREAELL lOAE F CAPITELLIOAE CAULLERIELLA KILLARIENSIS CHAETOPTERUS VARIOPEOATUS IDUNELLA SP. LUCIFER FAXO/l<I MARGINELUI kOSCIDA NEOPONTONIDES 8EAUFORTENSIS PHYLLOCHAETOPTERUS SOCIALIS POLYO ~A CAECA PSF.UDOMEDAEUS AGASS I ZI I SPI0PHANES SP. A SYLLl5 HYALINA THOR SP. TURBELLARIA ·· ·· ·· · ·· · ·· 0:3 ··· ·· ·· · · ··• · · ·· · 0.3 0.7 0.7 1.0 1.0 0.7 0.7 0.3 0.7 1 .7 0.3 0.7 1.7 0.3 0.3 0.7 0.7 ··· ·· · SUMMEk GNI~PS 51 ERR NAE~r OSOl r-AEAN 5T ERR 1.8 0.4 1.6 0.2 0.6 0.2 0.2 O.H 0.2 0.6 ·· ·· O.d O.b 0.2 0.5 0.7 0.4 1.0 1.0 0.2 0.8 0.6 0.6 0.'+ 0.2 1.4 1.0 1.4 1.4 1.4 0.4 0.8 0.8 1.2 1.2 0.4 0.2 1.0 1.0 1.0 0.6 1.2 0.6 0.2 1.0 0.4 0.4 0.2 1.2 0.5 0.4 0.9 0.4 0.4 0.6 0.4 0.6 0.7 0.2 0.2 0.8 0.5 1.0 0.4 1.0 0.6 0.2 0.5 0.4 0.8 0.2 0.2 0.5 0.2 0.2 0.2 1.4 1.6 0.6 1.0 1.6 0.2 · · ·· ··· ·· · ··· ·· ·· ··· ·· 0.3 0.3 (J.2 0.6 0:3 O.:J 1.7 1.2 0.'+ 0.4 0.4 0.2 1. u 1:0 0.6 0.4 0.3 1:7 · ··· · 0.3 0.7 ·• ··· · ··· · ··· · · ··· 1.7 · • 0.4 · · · O.b 0.4 1.0 · · · MEAN I:" 1.U 0.8 0.8 1.b 1.0 0.6 0.2 ·· ·· ·· FALL ST ERk MEAN l:U O.C 0.2 O.~ 0:6 0.4 0.6 0.2 0.2 0.8 0.2 0.2 0.,+ 0.6 0.,+ 0.6 1.4 0.4 0.,+ 0.9 0.2 0.4 0.4 0.6 0.2 0.4 0.2 0.2 0.4 0.6 1.8 1.0 0.2 0.2 ·· ··• · I.e · ·· · · ·• ·• · 0.6 1.4 0.2 0.2 0:2 0.2 ·· · 0.4 0.'+ · · • · • • ·• ·• · ST ERR · · • ·• · · • • · · O.b 0.6 0.4 0.4 0.4 0.6 0.4 0:2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.4 0.2 0.4 ·· 0.2 • 0.4 0:2 0.4 0.2 0.4 0.'+ 0.2 0.6 0.2 0.2 1.0 0.4 0.6 0.4 0.2 0.4 0.2 0.2 1.0 0.2 0.2 · 0.2 0.'+ · 0.4 0.6 · • • 0.2 0.2 · · · 0.2 0.2 0.6 0.4 0.8 0.6 0.2 o.a 0.'+ 0.6 0.4 0.2 • • • · · 0.6 0.2 Appendix 3. (Continued) OSOl STATION UvERALL KANK 11 H. ~ ll'i.5 11d.S 11R.S 118.5 Iltl.5 l1li.S l1R.S I11j.5 118.S 11 A. ':> 118.5 l1A.5 11A.5 118.5 llfi.5 14~.5 'J:J .;. 142.5 142.') 142.5 142.5 142.5 142.5 14~.5 142.5 142.5 142.5 142.5 142.5 142.5 142.5 142.5 142.5 142.5 142.5 142.5 142.5 142.5 \42.5 142.5 142.5 142.5 142.5 142.5 142.5 142.5 142.5 142.5 189.5 189.5 IH9.5 189.5 1119.5 189.5 189.5 SPfCIES \0 r-<E AI''l ACTEON CANOll\S AONIDfS 5ISNEZUGAY~ DELECTOPlCTEr-. SP. uLYCERA PAPILLOSA GLYCERA SP. A MACROCLYMfNE ZONALIS ODONTOSYLLIS E"lOPLA PALICUS SP. PELECYPOOA b POOOCHELA GRACILIPE5 POLYCIRRUS EXIMIUS SABELLARIA \/ULGARIS \/uLGARIS SEMELE AELLASTRIATA SPIO PETTIBONEAE SPIO SP. A STOMATOPOOA AMPHINO"1IDAE e AMPHIPODA C; ANOPLODACTYLUS PEfIOLATUS CAPRELLA F:QUILIBRA GLYCERIDAE uOULDIA ~NIREC LILJEBORGIA SP. 13 LIMA PELLUCIDA LUMBRINERIS INFLATA MEGALOBRACHIU~ SOR!ATUM M!TRELLA LUNATA NEPHTYS INClSA NUCULA PROXIMA ONUPHIS SP. OSTRACODA PAGURUS CAROLINENSIS PALAEMONIDAE PARAONIDAE tJHILINE SAGt-<A PHORONIS ARCHITECIA PHYLLODOCE ARENAE PHYLLODOCE .P~ PSEUDEURYTHOE Ai-HHGUA PTILANTHURA TRICAKINA SCHISTOMERINGOS RUDOLPH I SIPUNCULIDA SIPUNCULIDA E SPIONI[)lIE SPIONIDAE A SU8ADYTE PELLUCIOA TEREBELL IOAE. A TEREBELLIDAE Ii ACCALATHURA CRENULATA AMPHIO!)IA SP. AMPHIPODA E ARABELLA ,..lUT Ail< 5 ARGOPECTEN GIBBUS ARICIDEA FRAGILIS ASPJDOSIPHON ALBUS .. 0.3 0.7 ·· ··· 0.3 · 5T FRR l).3 ·· · ·· 0.7 0.3 1.U · · · 1.0 · ·· · ··· O.b · 1.0 1.0 0.7 0.7 ··· ·· ··· ·· · ··· ·· ·· ··· ·· ··· · ·· · ·· · ·· ·· 1.0 0.3 SUMMER SPRING II'ITER 0.7 0.7 · ···• ·· ··· · ··· ··· ·· · ·· · ·· ·· 0.3 0.3 0.3 0.3 0.3 0.7 0.3 FALL 5T ERR 0.2 0.2 0.6 0.4 0.4 0.8 0.6 0.2 0.2 MEAN 5T ERR ··· · ·· ·· 0.4 0.2 0.6 0.2 0.2 0.4 0.2 0.4 0.2 0.2 · 0.6 0.6 0.6 0.4 0.4 0.6 0.4 0.4 0.6 0.6 0.6 0.6 0.2 0.4 0.2 0.2 0.2 0.6 O.b 0.4 0.4 0.6 0.2 0.2 0.4 0.2 0.2 U.2 0.2 0.2 0.6 0.4 0.2 0.2 0.2 0.4 0.4 0.4 0.4 MEAN 0.6 0.2 0.6 0.4 0.4 0.8 0.8 0.6 O.~ · · · ··· · 0:2 0.4 0.6 0.6 0.6 ·• · 0.4 0.2 0.2 • 0.4 0.4 0.2 0.4 · · · ··· · · · 0:4 • 0.2 0.2 0.4 0.4 0.2 0.2 · 0.2 · 0.6 0.4 0.6 0.4 MEAN · · ST ~R£ · 0.6 0.6 0.2 0.2 0.2 O:~ 0.2 0:6 0.2 0.4 O.~ · · · 0.2 · · 0.2 0.2 ·· 0.6 · O.b ··· 0.6 • 0.2 0:4 0.2 0.2 • 0.4 0.2 0.2 0.2 0.4 0.2 0.2 · · · ·• ·· ··• ··· ·· ·• ·· 0.4 0:2 · • · ·• ·•• • · ·· · ·• ·· ·· 0.4 0:2 0.4 0.2 0.4 0.2 O.c · · · • 0.2 0.2 0.4 0.4 • 0.2 0.2 0.4 0.2 • 0.2 0.2 O.b 0.2 0.2 0.4 · · · 0.2 • 0.2 0.4 0.4 • 0.6 0.2 O.<l! · 0.2 · · 0.6 0.2 0.2 · 0.2 Appendix 3. (Continued) STA TION ~ OVERALL RANK U\ 189.5 189.5 189.5 189.5 189.5 1A9.5 1A9.5 189.5 189.5 189.5 189.5 189.5 189.5 189.5 189.5 189.5 189.5 189.5 189.5 189.5 189.5 189.5 189.5 189.5 189.5 189.5 189.5 189.5 189.5 lA9.5 189.5 189.5 189.5 1H9.5 189.5 189.5 189.5 189.5 189.5 189.5 189.5 189.5 189.5 1H9.5 lR9.5 189.5 189.5 1139.5 189.5 189.5 189.5 189.5 189.5 18Y.5 189.5 SPECIE:.S WINTER MUN CHRYSOPETALIDAE A CHRYSOPET ALIDAE B CORBULA CONTRACTA CRENELLA DIvARICATA DAPHNELLA MORRA EBALlA STIMPSONI EPITONIUM NOVANGLIAE ETHUSA MASCARONE AMERICANA GASTROCHAENA HIANS GLYCERA TESSELATA HIPPOMEDON SP. B IRIDOPAGURUS DISPAR LEMBOS SPINICARPUS INERMIS LUMBRTNERIOAE LUMBRINERIOES ACUTA LUMBRTNERTOPSIS PARADOXA LYONSJA HYALINA MAERA SP. A MAGfLONA SP. A MANGELIA AMIR ~UR MELTTA APPENDICULATA MICRODEUTOPUS MYERSI MICROPANOPE PUSILLA § MTCROSPIO PIGMENTATA NAINERIS SP. NEHERTINEA 0 OCNUS PYGMAEUS PAGURTSTES TRUNGULATU5 PARAPHOXUS SPINOSUS PECTINIDAE A PHILINE QUADRATA PHYLLOOOCE SP. F PHYLLODOCIDAE PILUMNUS SP. PISIONE REMOTA PLEUROMERIS TRIOENTATA PLICATULA GIBBOSA PODOCHELA SP. POECILOCHAETUS SP. PRIONOSPIO CIRRIFERA PROCESSA GUYANAE SCOLELEP1S TEXANA SCOLOPLOS SP. STCYONIA SP. SIGALIONIDAE SIPUNCULlDA 8 SOLENOCERA SP. STENOPLEUSTES SP. A STROMBIFORMIS BILINEATUS SYLLIDAE L SYNCHELIDIUM AMERICANUM TEREBELLIDAE C TEREBELLIDES STROEMI THARYX SP. TRTCHOPHOXUS EPISTOMUS ·· ·· · ·· ·• ···• · ·• ·• ···• ··· ·· · ···• ·· ··• ··• ··· ·· ·· 0.3 SUMMER GNI~PS ST ERR · ··• ·· ·· ··• ·· ·• • 0.3 0.7 0.7 0.3 0.3 0.7 0.7 0.3 0.3 0.7 0.7 0:3 0501 ··· ··• ··· · ··• ·· ·• · ··•• • • ··•• ··· · 0.3 MEAN ST ERR 0.4 0.4 0.4 0.4 0.4 0.4 0.2 0.4 0.4 0.4 0.2 0.4 0.2 0.4 0.2 0.2 0.4 0.4 0.4 0.4 0.4 0.2 0.2 0.4 0.2 0.2 0.4 0.4 0.4 0.4 0.4 0.4 0.4 0.4 0.4 0.2 0.4 0.2 0.4 0.4 0.2 0.2 0.2 0.4 0.4 0.4 0.2 0.4 0.4 0.4 0.2 0.4 0.2 0.4 0.4 0.2 0.2 0.2 0.4 0.4 0.2 0.2 · ·• ·· · ·• · ··• · • · · ·· • · • 0.2 ·· ·• 0.2 · ·• · · ··• ·· · ·• 0.4 0.2 0.4 0.4 0.2 0.2 0.2 0.2 0.2 0.2 0.2 MEAN 0.2 ··• · ·· FALL ST ERR · • • • 0.2 0.2 0.2 · ·• · 0.2 0.2 0.2 0.2 0.2 · 0.2 • 0.2 · ·· • 0.2 0.4 0.4 ·· 0.2 0.4 · · ·· ·· ·· 0.2 0.4 0.4 ·· · · ·· ·· ·· 0.2 0.2 0.4 0.2 0.2 0.2 0.2 0.2 0.2 • 0.2 {J.2 0.4 0.4 0.2 0.2 0.4 0.2 0.4 0.2 0.4 0.4 0.2 0.2 · · · · · ST ERR • 0.2 0.2 0.2 0.4 MEAN · · · · · · 0.2 · ·• ··• ·• · · • · 0.4 0.2 0:2 • • 0.2 • • · 0.4 · 0.2 0.2 · · 0.2 0.4 0.2 · · Appendix 3. (Continued) 0501 STATION OVERALL RANK \0 0- 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 SPECIES ALPHEUS SP. B AMPELISCA SP. C AMPELISCA VERRILLI AMPHINOMIOAE A A~PHIODIA TRYCHNA ANOMURA ARBACIA PUNCTULATA ARGOPECTEN IRRAOIANS ARICIOEA SUECICA ARMINA WATTLA AST~RIAS FORBESII ASTHENOTHAERU5 HEMPHILLI ASTROPECTEN COMPTUS BOOOTR I IOAE B BRANCHIOSYLLIS EXILIS CARIOEA CAVOLINIA LONGIROSTRI5 CHAETOPTERIDAE CHIONE LATILIRATA CIRSOTREMA OALLI COLLOOES SP. DENTALIUM OCCIDENTALE OIACRIA QUAORIDENTATA DIA5TYLIOAE A OIOOORA SI NE~ YAC ~ DIPLOCIRRUS CAPENSIS DONAX SP. DROMIIDAE EBALIA SP. EBALI ASP. * ECHINASTER SP. ECHINOIDEA ECHIURIDA EPITONIUM KREBSII EULALIA SP. EUNICE ANTENNAT A EUNICE SP. GALATHEA ROSTRATA GONEPL4X IEBS~I GONIADA TERES GONIAOIDAE B HIATELLA SP. A HIPPOLYTE SP. ~ HI PPOL YT IOAE HYOROIOE5 PROTULICOLA HYOROIDES UNCINATA HYPOCONCHA ARCUATA HYPOC ~CHA 5A8ULOSA IR DOPAGUR ~ OISPAR * LAEVICARDIUM PICTUM LEPTOCHELA BERMUDEN5IS LEPTOCHELIA SP. LEUCOTHOE SPINICARPA LYONSIA SP. LYSIOICE NINETTA MON ST ERR ··· ··· ··· · ··· ·· ··· ··· · ··· ·· ·· ·· ··• · · ·• ··· ··· ··· ·· · ·· · ··· ·· ·· ··· ·· ·· ·• ·· ·· ·· ·· · · ·· ··· ··· 0.3 0.3 0.3 0.3 0.3 0:3 0:3 0.3 0.3 0.3 0.3 0.3 0.3 0.3 0:3 0.3 0.3 0.3 0.3 0.3 !'lEAN ·· · 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 • 0.2 0.2 ··· · ·• ·· · ·· · ·• 5T ERR • · · · ·• ·· ·· ·· · 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 ·· 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 • 0.2 0.2 0.2 0.2 0.2 0.2 • • ·· · FALL SUMMER GNI~P5 wINTER · · ·• · • • • · MEAN 5T ERR ·· ·· · ·· ·· · ··· ·• ·• ·· ·· 0:2 0.2 0.2 0.2 ·· ·· 0.2 ·• ··• · ·•• ·• · ·• ·· ·•• ·• · ·• • ·· · O.~ ·· · · 0:2 0.2 0.2 0.2 0.2 0.2 MEAN 0.2 0.2 • • · 5T ERR 0.2 0.2 • • • • • • 0.2 · • 0.2 • 0.2 • 0.2 · ··• · 0.2 0.2 • • • 0.2 0.2 • · 0.2 • · 0.2 • 0.2 0.2 • • • · · · · 0.2 0.2 0.2 0.2 Appendix 3. (Continued) STATION OVERALL "lANK SPECIES \lINTER 29,.,.0 296.0 296.0 2<}/).0 29/:}.0 296.0 296.0 296.0 290.0 2"16.0 296.0 2'J6.0 ~ -.J 246.0 296.0 296.0 2"11->.0 296.0 290.0 296.0 296.0 296.0 296.0 29,.,.0 291'>.0 296.0 296.0 296.0 296.0 2'/6.0 296.0 296.0 296.0 29b.0 296.() 2<Jf:J.0 296.0 2l.J6.0 296.0 296.0 296.0 296.0 e96.0 296.() 2"16.0 2<}f,.0 296.1) 296.0 296.0 296.0 296.0 296.0 29h.0 29f:J.O 296.0 MAGFLONA SP. S MALACOCEROS GLUTAEUS MARGINELLA MU~IAYELTRAH MARGINF:LLIl :'P. MEDIOMASTUS CALIFORt--.IENSIS MEGALOMMA H10CULATUM MICROPANOPE SP. MICROPANOPE URINATOR * MUNIDA IRRASA MUNIDA SP. MUREX FLORIFtH DILECTUS MUSCULUS LATERAL IS NANNODIELLA ACIT NALE~ NANNOSQUILLA SP. A NASSAR INA MINOR NAsSARTUS VIBEX NE.PHTYlDAE NEREIDAE NEREIS ACUMINATA NEREJS SP. NOT ~ASTUs HE~IPODUS OD STO~IA LAEVIGATA ONUPHIDAE ONUPHIS SP. A OPHIOCNIDA SP. OPHIOTHRIX ANGULATA PAGURIDAE PAGURUS HENDERSON I * PAGURUS PIERCEI PAGURUS SP. * PALTCUS ALTERNATUS PALLIOLUM RETICULUM PAPANAITIS POLYNOIDES PARANA IT I5 SPECIOSA PARAPINNIX,A BOUVIERI PAROALISCIDAE A PAsIPHAEIOA£ PECTINIDAE PELECYPOOA F PEllA MUTT CA PERIn IMENE:. AMERICANUS PEqICLIMENES SP. PETROLISTHES GALATHINUS PHERUSJ\ EHLEr.iSI PHYLLODOCE MADEIRE.NSIS PHYLLODOCE SI NE~ANlIP PINNIXA SP. PINNOTHERIOAE PISTA QUAORILOBATA PLAKOSYLLIS QUADRIJCULATA POOARKE URSCURA POOOCHELA RIISEI POLINICES UBERINUS POLYCHAETA E PORTU~JUS SP. SPRING ST ERK NlIE~r O.6}<~ 0501 ·· ·· ··· ·· ·· · ·· ·· ·· ·· ··· ·· · ·· ··· ··· ··· ·· ··· ··· ·· ·· ·· ·· ··· ··· ·· ·· ·· 0.3 0.3 0.3 0.3 0.3 0.3 ·· · ··· ··· ··· ··• ·· ·· ·· ·· ··· ··• ·· ·· ·• ·• 0.3 0.3 SUMMER ST ERR NAE~r · · · · · 0:2 0.2 • 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 · · ·• • 0.2 0.2 0.2 ·· ·· ·• · · · ··• · 0.2 · · · · · · · · 0.2 0.2 0.2 ··• · · · ·• · · ·· · 0.2 0.2 0:2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0:2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 FALL MEAN ST ERR 0:2 0.2 ·.·· 0.2 0.2 ·· ·· ··• ·• · · · · · · ·· · 0.2 0.2 • ·· · ·· ·· · 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 ·· 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 · • · ·•• ·• MEAN ST ERI< 0.2 0.2 · ·· 0.2 · · · · • 0.2 · 0.2 0:2 0.2 · · · · · · · 0.2 0.2 0.2 0.2 · ·· ·• 0.2 • 0.2 · · 0.2 • · 0.2 · 0.2 • Appendix 3. (Continued) 5 TATION UVERALL RANK \Q 00 i96.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.0 296.U 296.0 296.0 2<,16.0 296.0 296.0 296.0 296.0 291'>.0 296.0 296.0 296.0 296.0 c96.0 296.0 £:'96.0 296.0 296.0 296.0 296.0 296.0 2~t).0 296.0 296.0 296.0 t?9f>.0 296.0 296.0 <:'96.0 £'96.0 296.0 5T ERR N~E'>t 5PINICARPUS PRIMNO LATREILLEI PRTOt-105PIO SP. PYLOPdGURUS HOLTHUI5I PYLOPAGUkUS SP. PYRUNCULUS CAELATU5 RANINIDAE SAAELL.IOAE SCALIRREGMA If\FLATU,.. SC()LELEPIS SR. [:3 5COLOPLOS SP. A SEMELE NUCULOIOES SERPULIOAE SICYONIA RREvIROSTRIS * SIGATICA SINE~LOkAC SIPUNCULIOA A 5 I PUNCULI DA C 50LfMYA VF.:LUM " 50LENOCERA ATLANTIUIS 5TENOCIONOPS FURCATA COELATA 5TENORHYNCHU5 SETICORNIS SP. A 5THEN LA~EL A SYLLIOAE SYLLI5 CORNUTA 5YNALPHEUS SP. TANAIDACEA 8 TELLINA SP. TELLINlI SR. • TEREBELLIDAE THOR FLORIDANU5 THOR MANNINGI TRACHYCAROIUM EGMONTIANUM • TRACHYCARI5 kE5TRICTUS TPAVISIA PAHVA IF< I CHOAR ANCHU5 GLACIALl5 TURRONILLA ABRUPT A VARICORHULA QFEI-<CULATA VOLVULELLA PAUPERCULA VOLVULELLA SIL MIS~EP wEBSTERINfREIS SP. XA/l1T '1 I OAE PORTUN ~ ·· ··· ··· ·· ·· ·· · ·· ·· ·· ·· ·• ··· ·· ·· ·· 0.3 0.3 0.3 ·· ··· ·· ·· ·· · ·· · ·· ·· ·· ·· ·· ···• ·· ··· 0.3 0.3 0.3 0.3 0.3 0.3 0.3 0.3 u.3 0.3 0.3 MEAN · · ·· ·· · ·· ··• · ·· · · ··· ·· · 5T ERR · · ··• · · ·· ··· ·· ·• · · ·· ··• · · 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 U:2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 • 0.2 FALL SUMMER SPRING wINTER SPECIES 0501 0.2 MEAN 5T ERR 0.2 0.2 ·· ·· ·· · ·•• · ·· 0.2 0.2 0.2 0.2 • 0.2 • ·· ··• ··· · ·• ·· ·• · 0.2 / ·• ··· · ··· ·• ·· · ·• ··• ·• ··• ·· ·• ··• 0.2 0.2 0.2 0.2 0.2 0.2 MEAN • · 0.2 • • 0.2 0.2 0.2 0.2 ST ERR • · 0.2 • 0.2 0.2 0.2 0.2 • • • • • ·• ·• ·• · ·• · ·· • 0.2 • • 0.2 • • • 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 • 0.2 0.2 0.2 · ·• · ·• ·• Appendix 3. (Continued) STATION OvERALL RANK 1.0 2.0 3.0 4.0 S.O 6.0 7.5 ~.5 IC IC 7.5 9.5 11.0 12.:' 12.5 14.0 15.0 16.5 16.5 19.5 19.5 19.':> 19.5 23.0 23.0 23.0 25.5 25.5 27.5 27.5 30.5 30.5 30.5 30.5 34.0 34.0 34.0 36.0 37.5 37.5 39.5 3~.5 44.0 44.0 44.0 44.0 44.0 44.0 44.0 50.0 50.0 so.o so.o ~o. 54.0 ':>4.0 54.0 SPECII:.S FILOGRANA I MPLEXA PLAKOSYLLIS QUAORIOCULATA RILDARDANUS LAMINOSA HESIONIDAE PROTOl-IEOE I A SP. A AMPHAPETE AMERICANA SPHAEROSYLLI S SP. SYLLIS S P O N ~ I C O L A CHONE AMERICANA SYLLIOAI:. L SYLLIS HYALIi'4A COROPHIIDAE GENOCIDARIS ~ACULT SERPULIDAE N PISTONE REMOTA CHRYSOPETALIDAE A NEMATODA GLYCERA TESSELATA MUNIDA Si-'. OLIGOCHAETA SPIOPHANES BOMBYX GLYCERA PAPILLOSA MEL IT IDAf A SYLL IS AL TEI·mAT A MUNIDA PUS ILLA PRIONOSPIO CRI5TATA PHOTIS SP. TANAIDACEA B CAULLERIELLA KILLARIENSIS EUNICE VITTATA NOTOMASTUS LATERICEUS SYLLI DAE GLYCERIDAE ODONTOSYLLIS ENOPLA SYLLIS Sf.'. 0 GLYCERA SP. B PROTODORVILLEA KEFEkSTEINI TURBELLAfo<IA EURYDICE PIPEkATA PISlA PALMATA Ek I CHTHOtH US SP. A GONIADIDES CAROLINAE HE5IONIDAE A LANICf SP. LEUCUTHUE APR CIN ~S SYLLIS FERRUG INA THOR SP. HAR~OTHOE SP. A PAGlJR IDEA PERICLIMENES AMERICANUS SPHAEROSYLLIS PIRIFERA XANTHIDAE A CERATONEREIS SIL8AR~ PAGUkISTES SP. SABELLIOAE 0506 WINTEF< SPRING MEAN 5T ERR ·· ·· ·•• ·· ··• ·•• ·• ··· · ··· ··• ·• ·• ·· ·• ·· ·· ·· ·· ·· ·· ··· ··· ·· ·· ··· ··· ·· ··•• • · ···• · ··•• ··• ··• ··· ·· ·• ··• ··· ·• ·· ·· ··• ·· SUMMER FALL Io1EAN ST ERF< MEAN 51 ERR MEAN 34.6 17 .2 15.8 13.4 5.0 5.0 9.0 4.4 2.8 3.6 3.8 6.6 1.8 2.6 2.2 1.8 0.4 2.2 2.4 1.6 4.4 1.8 2.0 2.0 2.2 2.8 0.6 0.8 2.0 2.2 30.7 16.9 8.8 13.4 3.1 4.0 7.8 2.9 1.5 3.3 2.6 6.6 1.6 2.3 2.2 1.3 0.4 1.7 2.1 1.6 2.9 1.0 2.0 1.1 1.3 2.3 0.4 0.6 1.8 1.5 1.0 5.2 4.0 0.8 0.2 4.4 1.0 2.3 1.2 0.6 0.2 3.5 18.0 6.6 1.0 0:4 2.8 2.8 1.6 1.8 2.5 2.8 1.4 0.8 1.2 2.4 0.2 2.0 2.2 0.2 0.6 2.0 0.6 1.4 0.2 1.0 0.4 1.0 1.9 0.2 1.3 1.2 0.2 0.6 1.8 0.4 0.9 0.2 1.0 0.4 0.2 0.2 1.6 0.6 0.7 0.4 · • ·· · · · ·· · · · 2.0 4.6 4.0 1.2 • 0.5 2.6 2.3 0.7 4.2 3.8 3.2 3.4 4.8 1.4 1.6 2.1:1 2.6 2.3 1.8 1.8 3.0 0.7 1.1 1.7 2.4 0.8 1.4 0.2 0.8 3.0 2.8 1.0 0.6 3.0 0.6 0.2 1.0 0.6 2.1 0.5 0.9 0.2 0.6 1.9 2.5 0.3 0.4 1.8 0.6 0.2 0.3 0.4 1.4 0.7 2.2 0.2 1.1 0.2 2.0 1.3 · · · ·• • • 0.6 0.2 0.8 1.0 1.2 0.4 0.6 0.4 1.6 0.2 1.2 · · ·• • • 0.6 0.2 0.6 0.4 0.6 0.4 0.4 0.2 0.2 0.2 0.8 · · ST ~RE 5.6 1.2 0.4 3.0 0.6 0.4 2.0 • 4.1 0.6 0.4 1.5 0.2 0.2 0.8 0.6 0.2 ·• ·•• 1.0 0.6 0.2 0.2 · ·• • • • 0.6 0.4 0.2 0.2 • 1.4 0.8 1.4 0.2 0.9 0.6 0.9 0.2 0.4 0.6 0.4 • 0.4 0.6 0.4 0.4 0.6 2.8 0.2 0.6 1.3 0.2 0.2 ·• • • • • 0.2 • • 1.6 • • • · ·• 0.2 1.2 1.4 0.2 1.0 0.4 0.4 • • • 0.2 1.0 O.b • 0.4 1.6 1.4 1.4 0.2 • 0.2 0.8 1.2 1.0 0.2 • Appendix 3. (Continued) STATION OVERALL fJANK .... 0 0 60.0 60.0 60.0 1>0.0 00.0 60.0 bO.O 60.U 60.0 "7.5 0-'.5 67.5 67.5 67.5 07.5 75.5 75.5 75.5 75.5 75.5 75.5 7S.5 7':>.5 7':>.5 75.5 ~6.0 ~6.0 86.0 lio.O 86.0 1;6.0 8b.0 86.0 86.0 86.0 116.0 104.5 104.5 104.5 104.5 104.5 104.5 104.5 104.5 104.5 104.5 104.5 104.5 104.5 104.5 104.5 lU4.5 104.5 104.5 1U4.5 ST ERR ~H,EM ·· ·· ·• ·• ·· ··· ·· ··· ··• · ··• ··· ·• ·• ·· ·• · ··· ·· ·· ··• • ·• ·· ·· · ·• ··· ··· ·• ···• ·•• ··• ··• ··• • ··•• ··• ··• · ··• ·· ·· ·· · ··• ··· · ST ERR NAE~! · · 1.4 1.0 0.6 0.8 1.6 1.2 1.4 0.4 0.6 0.6 1.1 1.0 1.2 1.0 1.2 1.4 1.0 0.4 0.2 0.6 0.2 0.2 1:2 0.4 1.2 1.4 0.8 0.2 0.2 0.6 0.2 0.2 0.8 0.8 1.2 0.4 1.2 0.4 1.2 0.4 0.8 0.2 · ·• · · 0.6 ·· 0.4 0.2 1.0 0.6 1.0 0.4 0.8 0.2 1.0 0.6 0.6 0.4 0.4 0.8 0.8 0.4 0.8 0.6 0.8 0.4 0.5 0.4 0.5 0.8 0.4 0.8 0.4 0:6 0.2 0.6 0.2 0.8 0.6 0.4 0.8 0.4 0.8 0.4 0.4 0.8 0.2 · ·• · · · FALL SUMMER SPt-<ING WINTEf< SPECIES AMPHIPODA G INGOLFIELLA SP. A LEPTOCHELlll SP. NEMEfJTINEA NOTOMtI.sTUS SP. PAGUtHDAE. SEMELE PUkPURASCENS SPATANGOIOEA XANTHIDAE. ARCOPsIS ADAMS[ CHASSINELLA LUNULATA OPHIOLEPIS ELEGANS PAGUfJUS SP. sERPULIDAE S I PUNCULI DA ASPIUOSIPHON GOSNOLDI AUTOLYTUS SP. GLYCERA CAPITATA LEUCUTHOE SP. A OPHIUROIDE.A PHYLLOCHAET0PTERUS SOClALlS PRIONOSPIO SP. PYLOPAGURUS HOLTHUISI SlPUNCULIDA 8 SU~ADYTE PELLUCIDA ALPHEUS NORMAf'<Nl AONIDES MAYAGUEZENSIS ARBACIA PUNCTULATA ELAsMOPUS SP. A LAONICE ClRRAlA LUCONAClA TNCERTA MlCROPANOPE SCULPTIPES PAGURISTES TuRTUGAE PHYLLOCARTDA TER ~EL IDES STROEMl )(ANTHIDAE. 8 ALPHEIDAE AMPELISCA VADORUM AMPHIP0DA K B~ANCHIOSYL IS EXILlS CAPIlELLlDAE A ECHINOIDEA GLANS DOMINGuENSIS ISCHNOCHITONlDAE A LEMBOs UNICORNIs LEPTOCHELA AT LUPA~ MAJTDAE. A MAPGINELLA HARTLEY ANUM NANOPLAX XANTHIFORHIS NEPHTYS SQUAMOSA ODONTOSYLLTS SP. OwENIA FUSIFORMIS PAGURUS HENDERSON I PHYLLODOCIDAE POLYDORA CAl::CA 0506 • ·• · · · ·· · ST E.Rt-< MEAN 1.6 0.2 0.4 0.4 0.6 1.4 0.2 0.2 0.4 0.4 0.2 0.6 0.2 0.4 1.6 0.2 0.2 0.4 0.9 0.2 0.2 0.4 1.0 0.2 0.2 1.0 0.4 0.4 0.2 0.2 0.8 0.2 1.0 NA.E~I · ·· · · ··· • · · ST ERR • 0.2 0.6 0.2 0.2 0.2 0.2 • 0.2 0.2 0.2 0.4 0.6 0.2 0.4 0.8 0.4 0.4 0.2 0.4 0.6 0.2 O.b 1.2 0.2 0.2 1.0 0.2 0.2 0.6 0.6 0.2 0.2 0.6 0.2 0.6 0.4 0.2 0.6 1.0 0.6 0.5 0.2 0.2 0.8 0.4 0.4 0.2 0.5 0.2 0.4 0.2 0.2 0.2 0.2 0.2 1.0 0.2 0.6 · ·• · · ·· · · · · · • ·· 0.8 0.4 • 0.6 0.2 0.2 0.2 · · · · 0.8 • • 0.8 · · · 0.4 0.8 0.6 0.2 0.4 0.0 0.6 0.2 0.6 0.2 0.2 0.4 · ·• • • 0.2 • 0.6 ·•• ·· ·•• ·•• · ·• · • ·• ·• 0.2 • • 0.4 • • • • • • • • 0.2 • 0.2 0.2 • 0.6 0.2 0.6 0.2 0.2 0.2 0.2 0.2 · • 0.4 • • ·• Appendix 3. (Continued) STATION OVERALL SPECIES KNA~ 104.5 104.5 104.5 104.~ 104.5 104.5 104.5 132.5 132.5 132.5 132.~ ... 0 132.5 132.5 132.5 132.5 132.5 132.5 132.5 132.5 132.5 132.5 132.5 132.5 132.5 132.5 132.5 1.32.5 132.5 132.5 132.5 132.5 132.5 132.5 132.5 132.5 132.5 132.') 173.5 173.5 173.5 173.5 173.5 113.5 173.5 173.5 173.5 173.5 173.5 173.5 173.5 173.5 173.5 173.':1 173.5 173.5 WINTER MEI\N POLYOORA SP. POMATOCEROS AMERICANUS PRTONOSPIO CIRRIFERA PRTONOSPIO SP. B SCHISTOMERINGOS RUDOLPH I SPIO PETTIBONEAE wEBST[RINEREIS SP. B AMPELISCA VENETIENSI5 AHPHIURA STIMPSONI ARMANDIA AT LUCA~ AUTO~ATE 80~MANIEL A EV R~AN I PORTUkICENSIS BRACHYURA CHEVALIA SP. ERICHTHONIUS 8RASILE~ EULALIA SANbUINEA GONlADlDAE A IRIOOPAGURUS OISPAR LEM80S SPINICARPUS INERMIS LOIMIA MEDUSA LUCIFER FAXONI lYSIOICE NINETTA MARGINELLA ROSCIDA OPH I OMUS IU" SP. OPHIOSTIG~A ISACANTHUM PALICUS SP. PERICLI~EN S SP. * POECILOCHAETUS SP. PROCE.SSA SP. PROTOMEDEIA SP. PYLOP~GURuS SP. ~ SYlLIS REGULATA EANILO~ C SYNAlPHEuS SP. TEREBEllIOAt. C THYONE PSEUDOFUSUS TRYPANOSYLlIS ZEBRA UNCIOlA SP. A ALPHEOPSIS SP. it AM PH I POOA ,j AMP ITHOE SP. A AMP ITHOJl)AE A ANACHIS LAFRESNAYI ANOPLOOACTYlUS PETIOlATUS ASTROPORPA AT LU~NA AXIOTHELlA MUCOSA BRANCHIOSYLLIS OCULATA CHLAMYS BENEDICTI CORBuLA CHITTYANA EUNICE ANTENNA fA EUPHROSINE. SP. GALA THEA ROSTHATA GASTROPODA K LILJEBORGIA SP. A lIlJERORGIA SP. ~ LUMBRINERIS LATREILLI ·· ·· ··· ··· ··· ·· ··· · ··• ··· ·· ··• ··• ·· ·• ·· ··· ··· ··· ·· ·· ·· · 0506 SUMMER SP ..<1NG ST ERR ·· ·· ··· ·. ··· ··· ·· ···,. ··• ·· ·· ·· ··· ··· ··· ··· ·• ·· ·· ·· ··· ·· FALL i"1EAN sr ERR MEAN ST ERR MEAN ST ERR 0.4 0.6 0.6 0.6 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.4 0.8 0.2 0.6 0.2 0.4 0.4 0.6 0.6 0.8 0.4 0.5 0.2 0.4 0.2 0.4 0.4 0.2 0.2 0.2 0.2 0.2 0.6 0.4 0.4 0.2 0.6 0.2 0.4 0.2 0.2 0.2 0.6 0.4 0.2 0.2 0.4 0.2 0.2 0.2 0.2 0.4 0.6 (j.6 0.4 0.4 0.6 0.6 0.2 0:2 0.2 0.4 0.2 0.2 0.4 0.2 • 0.2 • 0.2 · 0.2 0.2 0.2 0.4 0.2 U.2 0.2 0.,+ 0.6 0.2 0.4 0.,+ 0.4 0.2 0.2 0.4 0.2 0.2 0.6 0.6 0.2 0.2 0.6 0.6 O.b · · O.b 0.4 0.2 0.2 · · 0:6 0.2 0.2 0.2 0.4 0.4 O.t! ·· ·· · ·· ·· ·· ·· · ·· ·· 0.4 0.4 0.4 0.2 0.2 0.4 0.4 0.2 0.2 0.2 0.2 0.2 0:4 0.4 0.4 0:4 0.4 0.4 · ·· ··· · · · · · ·· ·· · · · · · 0.4 0.4 0.4 0.4 0.4 0.4 · · · · · · ·· O.~ · · · · · · · · 0.2 0.2 0:2 O.b 0.2 0.4 0.4 0.2 0.4 0:2 0.2 0.4 0.4 0.2 0.2 ·• 0:4 0.4 ·· · 0.2 · ·· o.c: · · 0:4 0.4 0.4 0.2 0.4 0.2 · • · • · o.c: 0.2 0.,+ 0.4 0.6 0.2 · · · · · · · · 0.4 0.4 0.4 0:2 0.2 U:2 Appendix 3. (Continued) 0506 STATION OVERALL RANK 1 (J.5 173.5 173.5 113.5 173.5 173.5 1 n.5 173.5 173.5 173.5 173.S 1 n.5 lU.5 173.5 173.5 173.5 173.5 173.5 173.~ .- 0 IV 1 73.~ 173.5 173.5 173.5 173.5 173.5 173.5 173.~ 173.5 173.5 173.5 173.5 173.5 173.5 173.5 t!.57.S c57.5 257.5 257.5 257.5 257.5 i!57.5 257.5 257.5 257.5 257.5 257.5 257.5 c'57.5 257.5 257.5 257.5 257.5 257.5 257.5 i!57.5 SPECIES MAJ lOAE MEL IT A APPENOICULATA MITHRAX ACUTICORNIS NASSAR INA RO~rIM NASSAR IUS .dLBUS NEREIS SLJCCINEA ONUPHIS PALLICULA ONUPHIS SP. OPHIACTIS S~VIGNYI OPHIOCNIDA SP. OPHIOTHRIX ANGULATA OSACHILA TUbEROSA PAGURUS CAROLINENSIS PALICUS FAXONI PARACERCEIS CAUDATA PARTHENOPE FHATERCULUS PELECYPODA lj PERICLIMENES IRWESCENS PILLJMNUS SP. PITAR SIMPSOrd POLYCIRRUS SUI~XE POLYPLACOPHURA A RIMULA FRENULATA SPIONIDAE STHENELAIS BOA STOMATOPODA SYlLIS CURNUTA TERf8ELL lOAE TETRATHYRUS SP. THARYX MAPIGNI TRICH08RANCHUS GLACIALIS VERMILIOPSIS ANNULATA wEBSTERINEREIS SP. WEBSTERINERUS TRIOENTATA AGLAOPHAMUS VERR ILLI ALBUNEIDAE ALPHEUS SP. 8 AMPELISCA SCHELLEN8ERGI AMPHINOMlDAE 8 AMPHIPODA AMPHIPODA 8 AMYGOALUM SAGITTATUM ANOPLODACTYLUS INSIGNIS ARC A IMBRICATA ASTERIAS FOR8E~I AUTOMATE 5P. AXIOGNATHUS SQUAMATUS CALLIOSTUMA PULCHRUM CALOCARIS HIHSUTIMANA CAPITfLLA CAPITATA CERAOOCUS SP. B CERATONEREIS SP. CERATONEREIS SP. 8 CEROORIlLIA PERRYAE VARIOPEUATUS CHAETOPTE.I-~US sur·H"1tR SPRING wINTER FALL MEAN ST ERH !'o1EAN ST ERR fit. AN ·· ·· ··· ···• ··· ··· ·· ··· ·· ·· ·· ··· ··· ·· ·· ·· ··· ··· ·· ··· · ··· ··· ·· ·· O.C 0.4 0.2 0.4 u.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.4 0.2 0.2 0.2 u.2 0.4 0.2 0.2 0.2 0.2 0.4 0.2 U.4 0.2 0.2 0:2 0.2 0.2 0.4 0.4 ·· · ··· ··· ·· ·· ·· ··· ··· · ··· ·• ·· ···• ·· ·· ·· ·• ··• ·· ·· · ·· · ··• • 0.4 • 0.4 ·· · ··• · ··• ··· · · ·· ··· · · ·· ·· ·· 0.2 0.4 · 0.2 0.2 0.2 0.2 0.2 0.2 0.4 · · o.c · · · ·· · 0.4 0.4 U.4 0:4 0.4 0.4 0:2 0.4 0.4 ·• · · · ·· · · · · ·· ·• ·· ·• ·• ·• 0.2 0.4 0.2 0.4 0.2 0:2 0.2 0.2 0.2 · ·• · · · ··· 0.2 0.2 0.2 U.2 0.2 0.2 0.2 0.2 0.2 ··• 0.2 • 0.2 • 0.2 • 0.2 0.2 0.2 0.2 · · · · 0.2 0.2 0.4 0.4 0.4 • 0.2 · · · · · 0.4 0.4 0.4 • 0.2 • • 0.2 0.4 0.2 • 0.4 0.2 ·• ·· · ·· · ·· · 0.2 0.2 0.2 0.2 O.C 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 · · · 0.4 0.2 0.2 0.4 0.2 0.2 0.4 0.2 0.4 0.4 0.2 0.4 0.4 U.2 · · · 0.2 o.c 0.2 ·· · ST ERR • 0.2 0:2 • ST ERR MEAN · ·· · 0.2 0.2 0.2 • • • 0.2 · · 0.2 • • • 0.2 0.2 • • • 0.2 Appendix 3. (Continued) STATION lJVEI-iALL RANK 0 w 257.5 ;:57.5 c.57.5 ':".17.5 C:57.5 t:,:>1.5 c':'7.S c'-,7.5 c.,::> 1.5 c.'::>7.5 25"7.5 c'::>7.5 c.'::>7.5 ;..,7.5 c5"7.5 2<;7.5 257.5 £'':>7.5 t!57.5 c57.5 c57.5 c57.5 c57.5 257.5 c.57.5 257.5 c.,::>7.5 257.5 257.5 c.,:>7.S iS7.S 257.5 c57.5 257.5 257.5 t!57.5 c57.5 i':J7.5 t!57.5 <:.57.5 c.':J7.5 c57.5 1.57.5 257.5 2S7.5 257.5 257.5 257.5 257.5 t!57.5 257.5 i!57.5 257.5 c57.5 257.5 SPECIES SETOSA CHAMA CONGREGATA CHIONE SP. CLYTHROCERUS PERPUSILLUS COLOMASTIX SP. CRASSINELLA MARTINICENSIS CRENELLA DECUSSATA CREPIDULA ACULEATA DAPDANUS INSIGI"IS DENDROCHIROTIDA DIPLOCIRRUS CAPENSIS EI-iTCHSONELLA SI~ROFTLIf EUl. ALI A SP. EULALIA SP. A EXOGONE DISPAH fAVARTIA CELLULOSA GAMMAROPSIS SP. GASTROCHAENA HIANS GONEPl.ACIDAE il GONIADIllAE GOULDIA CERI;-,A HETEROMYSIS FORMOSA HIPPOMEDON SP. A HOL01HUROIDE.A HYDROIDES SP. a HYPERI IOU HYPSICOMUS AI~EATOEAHP LAEVICARDIUM PICTUM LEPTOCHELA SP. LIMA SCAbRA LIMEA BRONNIA/liA LUl-'8k IN£R I OAE LYTECH!NUS VAkIEGATUS t-IACRURA MAERA SP. MAGELONA .~S 8 MEIOSQUILLA SNEDIR AU~ MEL IT TOAE MESOCHAETOPTEkUS SP. MICROJASSA SP. A MICROPANOfJE SP. MICROSPIO AT NE~GIP MITHkAX PLEURACANTHUS NATICA CANRt.NA NEOORILLIA CYDIA OPHIOPHRAG~US SP. PAGURUS PIEkCEI PAl.EANOTLJS SP. A PARTHENOPE SP. PELECYPODA A PERICLIME.NA£US ASCIOIAfWM PERICl.IMENAEUS ATLANrICUS PERICLIMENES SP. PH TIS I CA ~1 ARI NA PHYLLODOCE CASTANEA CHAETOZ ~.E wINTEF< SPRING MEAN ST ERR ·· ··· ·· ·· ··· ·· ··· ·· ··· ··· ·· · ··· ·· ·· ·· ·· ··· ·· · ·· ··· · ··· ·· ·· ·· ·· · ··• ··· · ··· · ··· ··· ···• ··· ·• ·· ·• ·· · ··· · ·· ·· · ··· ·· 0506 MEAN · · 0.2 0.2 0.2 0.2 0.2 0.2 0.2 SUMMER ST ERR · 0.2 0.2 0.2 0.2 0.2 0.2 0.2 · fALL MEAN ST ERR · 0.2 0:2 • · ·· · 0.2 0.2 0.2 0.2 0.2 0.2 0.2 o.~ 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 · ··· 0.2 ·· 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0:2 0.2 0.2 0.2 0.2 0.2 0.2 U:2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 ·· · 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 · ·· · ·· · ·· · ·· · · · ·· · ··• · ·· · ··· · · ··• 0.2 ·· 0.2 0.2 0.2 0.2 0.2 0.2 0.2 · · · o.~ · ·• · · · · ··· · · 0.2 0.2 0.2 0.2 0.2 0.2 ·· 0.2 · ST ERR 0.2 0.2 · • 0.2 · 0.2 • • · ·· ··· ··• ··· ·· MEAN · ·• · · · ··• ·• · ·· 0.2 0.2 0.2 0.2 0.2 0.2 0:2 • • 0.2 ··• · 0.2 · • • 0.2 • • · 0.2 Appendix 3. (Continued) STATION uvE~AL 0 J:> RANK 257.5 257.5 c57 .5 257.5 257.5 257.5 257.5 eST.5 257.5 257.5 257.5 257.5 c,:>7.5 257.':> 257.5 2':>7.5 2':>7.5 <:'57.5 257.5 257.5 cC,7.5 c57.5 257.5 <:'':>7.5 c.57.5 c.":>7.5 2":>7.5 257.5 <'.57.5 2':>7.5 257.5 257.5 <'.57.5 257.5 257.5 c.5"1.5 257.5 257.5 257.5 <:'57.':> SPECIES PHYLLODOCE MUCOSA PILlJMNUS SAn * PINNA CAl-mEA PINNIXfI SPa PINNOTHER[DAE POOARKE Of<SCuRA PODOCERUS SPa PODOCHELA GkACILIPES POLYCHAE1A (" POLYCIRRUS CAROLINEN5lS PRIONOSPIO CIRR08RANCHIATA PROCERAEA SPa PROCESSA PROFUNDA PROTULA TU8ULARIA PSAMMOLYCE C1ENIOOPHORA PSEUDEURYTHUE AMHIGUA PSEUDOMEDAEU5 AGASSIZII PSEUOOVEPMIL[A SPa PSEUDOVERMfLIA OCCIOEN1ALIS SABELLA A~L HT POKCIM SCALIBREGMA MUTALF~I SEMELE SPa SERPULA v[RMICULARIS GRANULOSA SICYONIA SPa SOLA5TER CARIBBAEU5 SPAIANGONIOAE SPIONIDAE [3 STHENELAIS SPa SYLLJOAE fJ 5YLLIDAE E SYLLIDAE 1 SYLLJS GRAC IllS SYNALPHEUS SUPRACIG~OL SYNALPHEUS MINUS SYNALPHEUS SPa * TANYSTYLIDAE A TRACHYCA~JS RESTRICTUS T~ITONIA 8AYE~I MISA TRIVIA CANOIDLJLA VITRINELLIOAt C ·· ·· ··• ··• ··• ·· ·· ·· ·· ·· ··· ··· ··· ···• · ··· SUMMER SPRING wINTER MON 0506 S1 ERR ·· ·· · ··· ·· ·· ··· ·· ·· ··· ··· ··· ·· · ··· ·· ··· · MEAN ST ERR 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0.2 0:2 MEAN · FALL 51 ERR · ·• · ··· · · · ·• ·· • 0.2 0.2 • 0.2 0.2 ·· 0.2 0.2 U.2 0.2 0.2 0.2 0:2 0.2 0:2 0.2 ·· 0:2 0.2 0.2 0:2 0.2 0.2 · ·· 0.2 0.2 · ·· ·· · ··· · ·· 0.2 · 0:2 0.2 · · · ·· ·· 0.2 0.2 0.2 0.2 · · ··· · · • 0.2 ST ERR 0.2 0.2 0.2 • 0.2 0:2 U.2 0.2 · · 0.2 v.2 0.2 0.2 0.2 0:2 0.2 · !'lEAN · · · · ·• • · 0.2 0.2 ·• · U.2 • • · 0.2 0.2 • • · 0.2 • 0:2 · 0.2