Hepatics and Anthocerotes of the Mediterranean, an annotated checklist

Montserrat Brugues

The Collective Article “New Mediterranean Biodiversity Records” of the Mediterranean Marine Science journal offers the means to publish biodiversity records in the Mediterranean Sea. The current article has adopted a country-based classification and the countries are listed according to their geographic position, from west to east. New biodiversity data are reported for 7 different countries, although one species reported from Malta is new for the entire Mediterranean basin, and is presumably also present in Israel and Lebanon (see below, under Malta). Italy: the rare native fish Gobius kolombatovici is first reported from the Ionian Sea, whilst the alien jellyfish Rhopilema nomadica and the alien fish Oplegnathus fasciatus are first reported from the entire country. The presence of O. fasciatus from Trieste is concomitantly the first for the entire Adriatic Sea. Finally, the alien bivalve Arcuatula senhousia is reported for the first time from Campania (Tyrrhenian Sea). Tunisia: a bloom of the alien crab Portunus segnis is first reported from the Gulf of Gabes,, where it was considered as casual. Malta: the alien flatworm Maritigrella fuscopunctata is recorded in the Mediterranean Sea for the first time, on the basis of 25 specimens. At the same time, web searches include possible unpublished records from Israel and Lebanon. The alien crab P. segnis, already mentioned above, is first formally reported from Malta based on specimens collected in 1972. Concomitantly, the presence of Callinectes sapidus in Maltese waters is excluded since based on misidentifications. Greece: the Atlantic northern brown shrimp Penaeus atzecus, previously known from the Ionian Sea from sporadic records only, is now well established in Greek and international Ionian waters. The alien sea urchin Diadema setosum is reported for the second time from Greece, and its first record from the country is backdated to 2010 in Rhodes Island. The alien lionfish Pterois miles is first reported from Greece and concomitantly from the entire Aegean Sea. Turkey: the alien rhodophyte Antithamnion hubbsii is first reported from Turkey and the entire eastern Mediterranean. New distribution data are also provided for the native fishes Alectis alexandrina and Heptranchias perlo. In particular, the former record consists of a juvenile Collective Article Α Mediterranean Marine Science Indexed in WoS (Web of Science, ISI Thomson) and SCOPUS The journal is available on line at http://www.medit-mar-sc.net DOI: http://dx.doi.org/10.12681/mms.1477 Medit. Mar. Sci., 16/3, 2015, 682-702 683 measuring 21.38 mm total length, whilst the latter by a mature male. Cyprus: the rare native cephalopod Macrotritopus defilippi, and the alien crab Atergatis roseus, sea slug Plocamopherus ocellatus and fish Cheilodipterus novemstriatus are first recorded from the entire country. Lebanon: the alien crabs Actaea savignii and Matuta victor, as well as the alien fish Synanceia verrucosa, are first recorded from the entire country. In addition, the first Mediterranean record of A. savignii is backdated to 2006, whilst the high number of M. victor specimens observed in Lebanon suggest its establishment in the Basin. The Atlantic fishes Paranthias furcifer and Seriola fasciata, and the circumtropical Rachycentron canadum, are also first reported from the country. The P. furcifer record backdates its presence in the Mediterranean to 2007, whilst S. fasciata records backdate its presence in the eastern Mediterranean to 2005. Finally, two of these latter species have been recently ascribed to alien species, but all three species mayfit the cryptogenic category, if not a new one, better.

Received 08.04.2016; accepted 21.07.2016; printed 30.03.2017 Proceedings of the 3rd International Congress “Biodiversity, Mediterranean, Society”, September 4th-6th 2015, NotoVendicari (Italy) I review here examples of recent progress in the taxonomy and biogeography of Mediterranean taxa. Morphological approaches have still much to offer, as shown by a study of the Sicilian species of the wingless weevil genus Pseudomeira Stierlin, 1881 (Coleoptera Curculionidae). A systematic analysis of molecular markers, however, is revealing a huge number of previously unsuspected cryptic species, as in the scarab genus Pachypus Dejean, 1821 (Coleoptera Pachypodidae). Other molecular studies have revealed very deep phylogeographic structure in the Corsican brook salamander; the presence of six or more species hitherto lumped under Rumina decollata (Linnaeus, 1758) (Pulmonata Subulinidae), in a snail genus in which biparental and uniparental reproduction coexist; the conservation of the same mal...

The Collective Article “New Mediterranean Biodiversity Records” of the Mediterranean Marine Science journal offers the means to publish biodiversity records in the Mediterranean Sea. The current article has adopted a country-based classification and the countries are listed according to their geographic position, from west to east. New biodiversity data are reported for 7 different countries, although one species reported from Malta is new for the entire Mediterranean basin, and is presumably also present in Israel and Lebanon (see below, under Malta). Italy: the rare native fish Gobius kolombatovici is first reported from the Ionian Sea, whilst the alien jellyfish Rhopilema nomadica and the alien fish Oplegnathus fasciatus are first reported from the entire country. The presence of O. fasciatus from Trieste is concomitantly the first for the entire Adriatic Sea. Finally, the alien bivalve Arcuatula senhousia is reported for the first time from Campania (Tyrrhenian Sea). Tunisia: a bloom of the alien crab Portunus segnis is first reported from the Gulf of Gabes, where it was considered as casual. Malta: the alien flatworm Maritigrella fuscopunctata is recorded in the Mediterranean Sea for the first time, on the basis of 25 specimens. At the same time, web searches include possible unpublished records from Israel and Lebanon. The alien crab P. segnis, already mentioned above, is first formally reported from Malta based on specimens collected in 1972. Concomitantly, the presence of Callinectes sapidus in Maltese waters is excluded since based on misidentifications. Greece: the Atlantic northern brown shrimp Penaeus atzecus, previously known from the Ionian Sea from sporadic records only, is now well established in Greek and international Ionian waters. The alien sea urchin Diadema setosum is reported for the second time from Greece, and its first record from the country is backdated to 2010 in Rhodes Island. The alien lionfish Pterois miles is first reported from Greece and concomitantly from the entire Aegean Sea. Turkey: the alien rhodophyte Antithamnion hubbsii is first reported from Turkey and the entire eastern Mediterranean. New distribution data are also provided for the native fishes Alectis alexandrina and Heptranchias perlo. In particular, the former record consists of a juvenile measuring 21.38 mm total length, whilst the latter by a mature male. Cyprus: the rare native cephalopod Macrotritopus defilippi, and the alien crab Atergatis roseus, sea slug Plocamopherus ocellatus and fish Cheilodipterus novemstriatus are first recorded from the entire country. Lebanon: the alien crabs Actaea savignii and Matuta victor, as well as the alien fish Synanceia verrucosa, are first recorded from the entire country. In addition, the first Mediterranean record of A. savignii is backdated to 2006, whilst the high number of M. victor specimens observed in Lebanon suggest its establishment in the Basin. The Atlantic fishes Paranthias furcifer and Seriola fasciata, and the circumtropical Rachycentron canadum, are also first reported from the country. The P. furcifer record backdates its presence in the Mediterranean to 2007, whilst S. fasciata records backdate its presence in the eastern Mediterranean to 2005. Finally, two of these latter species have been recently ascribed to alien species, but all three species may fit the cryptogenic category, if not a new one, better.

Cryptogamie, Bryologie, 2007, 28 (4): 351-437 © 2007 Adac. Tous droits réservés Hepatics and Anthocerotes of the Mediterranean, an annotated checklist R.M. ROSa*, V. MAZIMPAKAb, U. ABOU-SALAMAc, M. ALEFFId, T.L. BLOCKEELe, M. BRUGUÉSf, M.J. CANOa, R.M. CROSf, M.G. DIAg, G.M. DIRKSEh, W. EL SAADAWIc, A. ERDA∆i, A. GANEVAj, J.M. GONZÁLEZ-MANCEBOk, I. HERRNSTADT l, K. KHALILm, H. KÜRSCHNERn, E. LANFRANCOo, A. LOSADA-LIMAk, M.S. REFAIc, S. RODRÍGUEZ-NUÑEZk, M. SABOVLJEVI± p, C. SÉRGIOq, H. SHABBARAc, M. SIM-SIMq, L. SÖDERSTRÖMr In Memoriam to Creu Casas and Carmela Cortini Pedrotti aUniversidad de Murcia, Facultad de Biología, Departamento de Biología Vegetal, Campus de Espinardo, 30100-Murcia, Spain. rmros@um.es; mcano@um.es bUniversidad Autónoma de Madrid, Facultad de Ciencias, Departamento de Biología, Ctra. de Colmenar Viejo Km 15, 28049 Madrid, Spain. vicente.mazimpaka@uam.es cBotany Department, Faculty of Science, Ain Shams University, Abbasia, Cairo, Egypt. wagelsaadawi@yahoo.com, hshabbara@yahoo.com dDipartimento di Scienze Ambientali, Sezione di Botanica ed Ecologia, Laboratorio di Briologia, Università degli Studi di Camerino, Via Pontoni, 5 - 62032 Camerino (MC), Italy. michele.aleffi@unicam.it e9 Ashfurlong Close, Dore, Sheffield S17 3NN, United Kingdom. tblockeel@aol.com fDepartament de Biologia Animal, Vegetal i Ecologia, Unitat de Botànica, Universitat Autònoma de Barcelona, 08193 Bellaterra, Barcelona, Spain. montserrat.brugues@uab.es, rosamaria.cros@uab.es gDipartimento di Scienze Botaniche, Università degli Studi di Palermo, Via Archirafi, 38, 90133 Palermo, Italy. mgdia@unipa.it hInstitute iAdnan for Forestry and Nature Research (IBN), P.O. Box 23, NL-6700, AA Wageningen, Netherland. gerard.dirkse@wur.nl Menderes University, Faculty of Arts & Sciences, Biology Department, 09010, Aydin,Turkey. aerdag@adu.edu.tr jInstitute of Botany, Bulgarian Academy of Science, 23 Acad. G. Bonchev Str., 1113 Sofia, Bulgaria. animoss@bio.bas.bg kUniversidad de La Laguna, Departamento de Biología Vegetal (Botánica), Calle Astrofísico Francisco Sánchez s/n, 38071-La Laguna, Tenerife, Canary Islands, Spain. alosada@ull.es, jglezm@ull.es sofia.rodrigueznunez@gobiernodecanarias.org * Correspondence and reprints: rmros@um.es R.M. Ros, V. Mazimpaka et al. 352 lThe mEcology Hebrew University of Jerusalem, Israel. ilanahs@vms.huji.ac.il and Forestry Department, Faculty of Agriculture, University of Aleppo, Aleppo, Syria. nInstitut für Biologie der Freien Universität Berlin, Systematische Botanik und Pflanzengeographie, Altensteinstraße 6, D-14195 Berlin, Germany. kuersch@zedat.fu-berlin.de oBiology Department, University of Malta. edwinlanfranco@gmail.com pInstitute of Botany and Botanical Garden, Faculty of Biology, University of Belgrade, Takovska 43, 11000 Belgrade, Serbia. sabmar@hotmail.com qMuseu e Jardim Botânico, Rua da Escola Politécnica, 58, 102-1250 Lisboa, Portugal. msimsim@fc.ul.pt, csergio@fc.ul.pt rDepartment of Biology, Norwegian University of Science and Technology, N-7491 Trondheim, Norway. lars.soderstrom@bio.ntnu.no (Received 25 May 2007, accepted 19 September 2007) Abstract – The names of all hepatics and anthocerotes published to the end of may 2007 in the Mediterranean countries and Bulgaria are compiled in an annotated checklist list. The list comprises accepted names and synonyms and provides explanatory annotations for ambiguous and disputed names. A total of 403 accepted species and 12 infraspecific taxa are reported from the area. Liverworts / distribution / nomenclature / checklist / Mediterranean INTRODUCTION In the second half of the twentieth century, several authors published checklists summarizing the current state of knowledge of the distribution of bryophytes in Europe and Macaronesia (Podp{ra, 1954; Corley et al., 1981; Düll, 1983, 1984, 1985, 1992; Grolle, 1983; Corley & Crundwell, 1991; Grolle & Long, 2000; Dierßen, 2001; Söderström et al., 2002; Hill et al., 2006). Within the Mediterranean region many lists have been produced at national level (Casas, 1991, 1998; Herrnstadt et al., 1991; Aleffi & Schumacker, 1995; Düll, 1995; ElSaadawi et al., 1999; Sabovljevi† & Stevanovi†, 1999; Sabovljevi†, 2000, 2003; Cortini Pedrotti, 2001a; Ganeva & Natcheva, 2003; Sérgio & Carvalho, 2003; Kürschner & Erda∑, 2005; Natcheva & Ganeva, 2005; Aleffi, 2005; Cekova, 2005; Colacino & Sabovljevi†, 2006; Dragicevi† & Velji†, 2006; Sérgio et al., 2007), and several new floras have been published or are currently in the process of publication (Casas et al., 2001, 2004, 2006; Cortini Pedrotti, 2001b, 2006; Heyn & Herrnstadt, 2004; Guerra & Cros, 2006). There have also been lists covering different sectors of the Mediterranean region, as well some of the larger islands (Bischler & Jovet-Ast, 1973; Eggers, 1982; Dia et al., 1987; Frey & Kürschner, 1991; Dirkse et al., 1993; Düll et al., 1999; Ros et al., 1999; Losada-Lima et al., 2001, Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 353 2004; Sabovljevi† & Natcheva, 2006; Sotiaux et al., 2007). However no attempt has been made to produce taxonomically critical lists of bryophytes for the whole Mediterranean region. With respect to liverworts, only two works can be regarded as covering the whole area, and these are mainly focused on ecological and biogeographical aspects (Bischler-Causse, 1995; Bischler, 2004). Because of the intensive research activity that is currently taking place in the Mediterranean area, and the need for updated red lists to inform conservation studies and policies, it is important that accurate and comprehensive lists of published bryophyte names should be available for the whole Mediterranean region. The OPTIMA Bryophyte Commission, during a meeting held at Palermo in September 2001, proposed that the gap should be filled by the publication of a checklist of Mediterranean bryophytes in two stages: initially for liverworts and subsequently for mosses. After an interval of several years the first stage has been completed and we present here a checklist of Mediterranean liverworts in the form of a catalogue of recognized names and synonyms. The catalogue includes all the names that have been published for any one or more of the countries on the periphery of the Mediterranean Sea. The project During the OPTIMA meeting in 2001, the Bryophyte Commission considered two important matters: 1) the definition of the area to be included, and 2) the authorship of the list and the system to be adopted in its preparation. The point at issue in the first question was whether the list should include the whole of each country bordering the Mediterranean littoral, or only the areas within these countries subject to a Mediterranean climate. It was clear that there would be serious difficulties in correlating the boundaries of study areas and their climatic regimes, and we concluded that the only practical option was to define the area using the national boundaries of the countries bordering the northern, eastern and southern shores of the Mediterranean. We included Bulgaria since part of that country is subject to a Mediterranean climate (see Fig. 1 for a map of the study area). We decided to treat the larger islands as distinct units because of their biogeographical interest. However the Azores archipelago was excluded because of its non-Mediterranean climate. With regard to the second question, it was felt appropriate to designate authors to be responsible for each country and island group (Table 1). These authors would compile a list of accepted names and synonyms according to criteria proposed by the Commission and send them to the Secretary who, assisted by the remaining members of the Commission, would consolidate them into a single paper. From the time of these initial decisions, the project has proceeded through 5 stages: 1) the Commission prepared the criteria for the list of accepted names and synonyms and distributed them to the designated authors; 2) the authors compiled their lists and sent them to the Secretary of the Commission (R. M. Ros), who in collaboration with a second member of the commission (V. Mazimpaka), consolidated the country and islands lists into a master list of accepted names and a separate list of synonyms; 3) the resulting lists were sent back to the authors for data checking, correction and update; 4) having rechecked and re-integrated the author amendments and additions in the master list, the Commission edited the final version of the paper, which (5) was returned to the authors for text checking. This resulted in the version that was submitted for publication. 354 R.M. Ros, V. Mazimpaka et al. Fig. 1. Map of the study area indicating the abbreviation used the countries and islands for which a separate list is included in the checklist (cf. also Table 1). The lengthy duration of the project is explained by its complexity and the fact that the members of the OPTIMA Bryophyte Commission had other concurrent activities and responsibilities. However the experience gained during the project will serve as a sound basis for the checklist of mosses still to be produced. Taxonomy and Nomenclature Criteria for the list of accepted names The list includes published names and records to the end of May 2007. The checklist by Grolle & Long (2000) was the basic reference work for accepted names in Europe and Macaronesia. We have made some exceptions to this list in the light of more recently published taxonomic papers, and we have added the names of taxa from African and Asian countries that were outside the geographical scope of Grolle & Long’s list. We have also included some names that are not cited in Grolle & Long’s list in view of their frequent use in Mediterranean areas and the lack of consensus in recent literature on their taxonomic value. For example, Grolle & Long (2000) cited very few infra-specific names. We took the view that the varieties and subspecies of some polymorphic taxa that have been recognized in relatively recent studies merited inclusion (e.g., those of Bazzania trilobata (L.) Gray, Blepharostoma trichophyllum (L.) Dumort., Jubula hutchinsiae (Hook.) Dumort., Marchantia polymorpha L. and several species of Riccia). In cases where infraspecific taxa have not been distinguished in the literature for a Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 355 Table 1. Countries and island groups treated in the checklist, with the names of the responsible authors. Country abbreviations are based on the two-letters code defined by ISO 3166-1 for country names. For the islands that are not independent countries a two-letter code as used in Söderström et al. (2002) has been used except for Madeira for which MD has been used. For the purpose of the checklist, Andorra has not been considered. Sinai has been included in Egypt. Israel includes some adjacent areas (Gaza, the West Bank, the former Cis-Jordan, and parts of both the Golan Heights and Mt. Hermon). The list of Malta includes the Maltese islands: Malta, Gozo and Comino. The Spanish North-African territories have been included in Morocco; Kosovo in Serbia; Gibraltar in Spain. Turkey includes the European and Asian parts, although no segregation between them has been done. Countries and Islands Abbreviations Authors Albania AL M. Sabovljevi† Algeria DZ R.M. Ros & M.J. Cano Baleares BL M. Brugués & R.M. Cros Bosnia-Herzegovina BA M. Sabovljevi† Bulgaria BG A. Ganeva Canary Islands CN A. Losada-Lima, G.M. Dirkse, S. Rodríguez-Nuñez & J.M. González-Mancebo Corsica CO L. Söderström Crete CT T.L. Blockeel Croatia HR M. Sabovljevi† Cyprus CY T. L. Blockeel Egypt EG W. El Saadawi, H. Shabbara, U. Abou-Salama & M.S. Refai France FR L. Söderström Greece GR T.L. Blockeel I. Herrnstadt Israel IL Italy IT M. Aleffi Jordan JO H. Kürschner Lebanon LB H. Kürschner & A. Erda∑ Libya LY R.M. Ros & M.J. Cano Macedonia MK M. Sabovljevi† Madeira MD M. Sim-Sim Malta MT E. Lanfranco Montenegro ME M. Sabovljevi† Morocco MA R.M. Ros & M.J. Cano Portugal PT C. Sérgio Sardinia SA M. Aleffi Serbia RS M. Sabovljevi† Sicilia SC M.G. Dia Slovenia SI M. Sabovljevi† Spain ES M. Brugués & R.M. Cros Syria SY W. El Saadawi, H. Shabbara, U. Abou-Salama, M.S. Refai & K. Khalil Tunisia TN R.M. Ros & M.J. Cano Turkey TR H. Kürschner & A. Erda∑ 356 R.M. Ros, V. Mazimpaka et al. given territory, then the relevant occurrences have been indicated in the checklist at species level. For the most part, however, we have been restrictive in our inclusion of infraspecific taxa. Thus, if the majority of recent authors have not recognized a taxon, then we have also ignored it here. We have not recognized the category of “form”. We have compared the contents of our list as they relate to European territories with the data and subsequent amendments published by Söderström et al. (2002; 2007) (see http://www.bio.ntnu.no/ECCB/ for a complete checklist). Taxa erroneously reported from the territories covered by Söderström et al. (2007) and noted as such by them are not repeated here. Taxa whose presence in a given area is doubtful have been excluded at the discretion of the author responsible for that area. We have ignored names previously rejected in published papers, but have included an explanatory annotation for those rejected here. The names of the accepted taxa and their occurrence in individual territories are presented in tabular format. Taxa are arranged alphabetically within taxonomic unit, and territories are listed in alphabetic sequence of their respective abbreviations. The reported presence of a taxon in a territory is marked by one of the following symbols indicating the nature of the record. The absence of a symbol means that the taxon has not been reported in the territory or has been rejected in previous publications. Annotations concerning taxonomic criteria and floristic data are indicated by a number following the name of the taxon, or in the relevant cell of the grid. o n (o) (n) + Report based on collections published before 1962. Report based on at least one collection published during or after 1962. Doubtful report based on collections published before 1962. Doubtful report based on at least one collection published during or after 1962. Report without precise collection data or locality. Criteria for the list of synonyms Our objective was to compile all the names published for Mediterranean liverworts, and we have therefore included all available synonyms. The authors of the national and island lists were asked to include all the names published for their designated territories. All these names were checked using standard works on liverworts and other important recent publications, including general and regional floras (Müller, 1954-1957; Schuster, 1966, 1969, 1980, 1988, 1992a, 1992b; Paton, 1999; Schumacker & Vá≈a, 2000, 2005; Damsholt, 2002), generic monographs (Grolle, 1972; Jovet-Ast, 1986; Bischler-Causse, 1993; Sim-Sim, 1999; Engel & Merrill, 2004; Long, 2006, etc.) and earlier lists (e.g., Grolle, 1983, Grolle & Long, 2000; Söderström et al., 2002). Our intention was to include all the names used in the literature, even if they had been misapplied or had not been validly published. However it became clear that the authorities given for names cited in floristic publications were sometimes erroneous, and we have therefore ignored such errors. The names and authorities of taxa were first checked against Index Hepaticarum (Bonner, 1962a, 1962b, 1963a, 1963b, 1965, 1966; Bischler, 1977; Geissler & Bischler, 1985, 1987, 1989, 1990) and Index of Hepatics (Crosby & Engel, 2006), and the nomenclature was then further verified using the publications cited above. Author abbreviations follow Brummitt & Powell (1992), and annotations are included for problematic names. Invalid combinations have been included, especially basionyms and their subsequent combinations, with the aim of facilitating their interpretation. Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 357 For ease of consultation, all the synonyms and disputed names used in the Mediterranean countries have been arranged alphabetically in a single consolidated list, with a cross-reference where appropriate to the annotation in which the taxonomic problem is discussed. This list also includes names rejected for the whole of our area. Summarizing the results of three centuries work in a critical list is a complex and difficult task, especially considering that advances in taxonomy and nomenclature have run in parallel with floristic exploration. While we have attempted to check all names against the available literature, we do not assume that the result is perfect. We would welcome any further observations regarding omissions, errors and inconsistencies, since they will serve to improve future editions of the checklist. 358 R.M. Ros et al. Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 359 JO TR List of accepted names AL BA BG BL CN CO CT CY DZ EG ES FR GR HR IL IT LB LY MA MD ME MK MT PT RS SA SC SI SY TN j ● j ● j ● ● j ● ● ● ● Anthocerotae Anthoceros L. agrestis Paton [2] caucasicus Steph. punctatus L. [2] ● ● ● ● ● ● + ● ● j ● ● ● ● ● Notothylas Sull. ex A. Gray orbicularis (Schwein.) A. Gray Phaeoceros Prosk. carolinianus (Michx.) Prosk. laevis (L.) Prosk. [4] [5] Phymatoceros Stotler, W. T. Doyle & Crand.-Stotl. [6] bulbiculosus (Brot.) Stotler, W. T. Doyle & Crand.-Stotl. [5] + ● ● [1] j ● ● ● ● j ● ● + j + ● ● + ● ● ● ● [3] ● + ● ● ● ● j ● ● ● ● ● ● ● ● ● ● j ● j ● j ● ● + ● ● ● ● ● ● (●) ● ● ● (●) ● ● ● ● ● ● ● Hepaticae Acanthocoleus R. M. Schust. aberrans (Lindenb. & Gottsche) Kruijt ● ● Acrobolbus Nees wilsonii Nees j) (j ● Adelanthus Mitt. decipiens (Hook.) Mitt. ● ● Anastrepta (Lindb.) Schiffn. orcadensis (Hook.) Schiffn. ● ● Anastrophyllum (Spruce) Steph. assimile (Mitt.) Steph. hellerianum (Nees ex Lindenb.) R. M. Schust. michauxii (F. Weber) H. Buch minutum (Schreb.) R. M. Schust. saxicola (Schrad.) R. M. Schust. Aneura Dumort. maxima (Schiffn.) Steph. pinguis (L.) Dumort. Anthelia (Dumort.) Dumort. julacea (L.) Dumort. [7] juratzkana (Limpr.) Trevis. [7] Aphanolejeunea A. Evans azorica (V. Allorge & Ast) Bernecker & Pócs ● ● ● ● ● ● ● ● ● ● ● ● ● j j ● ● j ● j ● ● ● ● j ● ● ● ● ● ● ● ● ● ● ● ● ● j ● ● ● + j ● ● ● ● ● ● ● j ● j j ● ● R.M. Ros et al. 360 AL BA BG BL madeirensis (Schiffn.) Grolle microscopica (Taylor) A. Evans sintenisii Steph. CN CO CT CY DZ EG ES ● ● IT JO TR LB LY MA MD ME ● MK MT PT RS SA SC SI ● j SY TN ● ● ● Asterella P. Beauv. [8] africana (Mont.) A. Evans gracilis (F. Weber) Underw. [9] lindenbergiana (Corda ex Nees) Arnell saccata (Wahlenb.) A. Evans [10] ● ● ● ● j ● ● ● ● j ● j ● ● ● ● ● ● + ● ● ● ● ● ● ● j ● ● ● j j ● j) (j ● ● ● ● ● ● j j ● (●) ● ● ● ● ● ● ● j j) (j ● ● ● j j ● ● ● j ● ● j ● j) (j ● j ● [1] ● ● j ● ● j j + j [1] j ● ● ● ● j j j ● ● ● j ● ● j ● j j j ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● j j ● ● ● ● j ● ● j j ● ● ● ● ● ● ● j ● ● j j ● + (●) ● + j ● ● j j ● ● ● ● j ● j) (j ● ● ● j j ● ● ● ● + j j ● ● ● ● Blasia L. pusilla L. Blepharostoma (Dumort.) Dumort. trichophyllum (L.) Dumort. subsp. brevirete (Bryhn & Kaal.) R. M. Schust. subsp. trichophyllum IL ● Arnellia Lindb. fennica (Gottsche) Lindb. Bazzania Gray flaccida (Dumort.) Grolle tricrenata (Wahlenb.) Lindb. trilobata (L.) Gray var. depauperata (Müll. Frib.) Grolle var. trilobata HR ● Apometzgeria Kuwah. pubescens (Schrank) Kuwah. Barbilophozia Loeske atlantica (Kaal.) Müll. Frib. attenuata (Mart.) Loeske barbata (Schmidel ex Schreb.) Loeske binsteadii (Kaal.) Loeske floerkei (F. Weber & D. Mohr) Loeske hatcheri (A. Evans) Loeske kunzeana (Huebener) Müll. Frib. lycopodioides (Wallr.) Loeske quadriloba (Lindb.) Loeske rubescens (R. M. Schust. & Damsh.) Kartt. & L. Söderstr. GR 361 ● Apomarsupella R. M. Schust. revoluta (Nees) R. M. Schust. Athalamia Falconer hyalina (Sommerf.) S. Hatt. [9] spathysii (Lindenb.) S. Hatt. FR Hepatics and Anthocerotes of the Mediterranean, an annotated checklist [1] j j j ● ● j ● j j ● ● j ● j j (●) j j ● ● ● [1] + ● ● ● ● ● ● ● ● ● ● ● R.M. Ros et al. 362 AL Calypogeia Raddi arguta Nees & Mont. azorica Bischl. azurea Stotler & Crotz [11] [12] fissa (L.) Raddi integristipula Steph. muelleriana (Schiffn.) Müll. Frib. [11] neesiana (C. Massal. & Carestia) Müll. Frib. sphagnicola (Arnell & J. Perss.) Warnst. & Loeske suecica (Arnell & J. Perss.) Müll. Frib. Cephalozia (Dumort.) Dumort. affinis Lindb. ex Steph. ambigua C. Massal. bicuspidata (L.) Dumort. catenulata (Huebener) Lindb. connivens (Dicks.) Lindb. crassifolia (Lindenb. & Gottsche) Fulford lacinulata J. B. Jack ex Spruce leucantha Spruce loitlesbergeri Schiffn. lunulifolia (Dumort.) Dumort. macrostachya Kaal. pleniceps (Austin) Lindb. Cephaloziella (Spruce) Schiffn. baumgartneri Schiffn. calyculata (Durieu & Mont.) Müll. Frib. [14] dentata (Raddi) Steph. divaricata (Sm.) Schiffn. elachista (J. B. Jack ex Gottsche & Rabenh.) Schiffn. elegans (Heeg) Schiffn. [15] granatensis (J. B. Jack ex Steph.) Fulford grimsulana (J. B. Jack ex Gottsche & Rabenh.) Lacout. hampeana (Nees) Schiffn. [14] integerrima (Lindb.) Warnst. massalongi (Spruce) Müll. Frib. phyllacantha (C. Massal. & Carestia) Müll. Frib. rubella (Nees) Warnst. [15] spinigera (Lindb.) Warnst. stellulifera (Taylor ex Spruce) Schiffn. turneri (Hook.) Müll. Frib. varians (Gottsche) Steph. BA BG BL CN CO ● ● CT CY DZ ● EG ES FR ● ● GR HR IL j IT Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 363 JO TN TR ● ● j) (j ● ● ● LB LY MA MD ME j ● MK ● MT PT RS ● SA SC SI ● ● j SY ● j j ● ● j ● ● ● ● ● (●) ● ● ● ● ● ● ● ● ● ● ● ● ● [1] ● ● ● ● ● ● ● ● ● ● ● ● j ● j j) (j (●) j) (j ● ● ● j ● ● j j ● ● j) (j ● j ● ● ● ● j ● (●) (●) ● ● ● ● j ● j ● ● j ● ● j ● ● ● ● ● ● ● j ● ● ● j ● j ● j j ● ● ● ● [1] ● ● ● ● ● ● ● ● ● (●) ● ● ● ● ● ● ● ● ● j ● ● ● ● j) (j j ● ● ● j j ● ● ● ● ● ● j ● j ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● + ● ● ● ● ● ● j ● j ● ● ● ● j) (j ● ● ● ● ● ● ● ● ● j j j ● [13] ● ● ● ● ● + j j ● j ● ● ● + ● ● ● ● ● ● ● ● j ● j ● ● ● ● j [1] j ● ● ● ● ● ● ● ● ● ● ● ● j ● ● ● ● ● ● ● j ● ● ● ● ● j ● ● j ● j j ● + ● j + ● ● ● ● ● ● ● ● ● ● ● ● ● j (●) ● ● ● j ● ● + ● ● ● ● ● ● ● + ● j ● ● ● ● ● ● ● ● ● ● j ● ● ● (●) ● ● ● ● ● ● ● ● j ● ● ● j ● ● ● ● ● ● ● ● j ● j j ● ● + j ● ● ● ● ● ● ● + R.M. Ros et al. 364 Chiloscyphus Corda pallescens (Ehrh. ex Hoffm.) Dumort. polyanthos (L.) Corda AL BA BG BL CN CO [16] j ● ● j j ● ● CT CY ● j ● HR j ● ● ● j ● j ● ● j ● ● ● j ● j ● j j ● ● ● ● ● Corsinia Raddi coriandrina (Spreng.) Lindb. ● ● ● ● ● j ● ● ● ● ● j ● ● ● ● ● j ● ● ● ● ● ● ● ● ● ● ● ● j ● ● ● j ● MK j MT PT RS SA SC SI ● ● ● ● j ● ● ● ● j SY TN ● ● ● ● ● ● ● [1] ● j ● j ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● j ● ● ● ● j + ● ● ● ● ● ● ● (●) ● ● ● ● ● ● ● ● ● j ● ● ● ● ● ● ● ● ● ● ● ● j ● ● ● j ● ● ● [1] ● ● j j j ● ● j ● ● (●) ● ● j ● ● ● Douinia (C. E. O. Jensen) H. Buch ovata (Dicks.) H. Buch Drepanolejeunea (Spruce) Schiffn. hamatifolia (Hook.) Schiffn. ● Dumortiera Nees hirsuta (Sw.) Nees ● Exormotheca Mitt. pustulosa Mitt. welwitschii Steph. MA MD ME ● Cyathodium Kunze foetidissimum Schiffn. Eremonotus Lindb. & Kaal. ex Pearson myriocarpus (Carrington) Pearson LY ● Cryptothallus Malmb. mirabilis Malmb. Diplophyllum (Dumort.) Dumort. albicans (L.) Dumort. obtusatum (R. M. Schust.) R. M. Schust. obtusifolium (Hook.) Dumort. taxifolium (Wahlenb.) Dumort. LB ● Colura (Dumort.) Dumort. calyptrifolia (Hook.) Dumort. Conocephalum Hill conicum (L.) Dumort. salebrosum Szweyk., Buczkowska & Odrzykoski [17] ● TR GR Cladopodiella H. Buch fluitans (Nees) H. Buch francisci (Hook.) Jörg. Cololejeunea (Spruce) Schiffn. calcarea (Lib.) Schiffn. minutissima (Sm.) Schiffn. rossettiana (C. Massal.) Schiffn. schaeferi Grolle JO FR EG IT 365 ES DZ IL Hepatics and Anthocerotes of the Mediterranean, an annotated checklist ● ● ● ● ● ● ● ● ● ● ● ● [1] ● ● ● j ● j ● ● j j j ● j ● j) (j ● ● ● ● j ● ● ● ● ● j j ● ● ● ● R.M. Ros et al. 366 AL Fossombronia Raddi [18] [19] angulosa (Dicks.) Raddi caespitiformis De Not. ex Rabenh. [20] [21] subsp. caespitiformis subsp. multispira (Schiffn.) J. R. Bray & D. C. Cargill crispa Nees [22] echinata Macvicar [20] foveolata Lindb. incurva Lindb. leucoxantha Lehm. mittenii Tind. pusilla (L.) Nees [21] [23] wondraczekii (Corda) Lindb. [20] [23] Frullania Raddi azorica Sim-Sim et al. [24] dilatata (L.) Dumort. [25] ericoides (Nees) Mont. fragilifolia (Taylor) Gottsche et al. inflata Gottsche jackii Gottsche [26] microphylla (Gottsche) Pearson oakesiana Austin parvistipula Steph. polysticta Lindenb. riparia Hampe ex Lehm. sergiae Sim-Sim et al. tamarisci (L.) Dumort. [25] teneriffae (F. Weber) Nees BA ● ● BG BL CN CO CT ● ● ● ● ● ● ● ● ● ● ● ● ● (●) ● ● ● j ● CY DZ j ● Gymnocolea (Dumort.) Dumort. inflata (Huds.) Dumort. Gymnomitrion Corda apiculatum (Schiffn.) Müll. Frib. concinnatum (Lightf.) Corda corallioides Nees crenulatum Gottsche ex Carrington obtusum Lindb. FR GR HR ● ● ● j IL IT 367 JO TN TR ● ● ● ● j ● ● ● j ● ● ● ● ● ● ● ● ● ● LY [1] ● ● LB ● MA MD ME ● ● ● ● ● ● [1] j j ● ● j ● j j MK MT ● PT RS ● j SA SC ● ● ● ● ● ● ● ● j ● ● ● SI SY ● ● (●) + ● ● ● ● + ● ● ● ● ● ● ● ● ● ● j ● j ● ● ● ● j ● ● ● j ● ● + (●) j ● ● ● ● ● ● ● ● j ● ● ● ● j j ● ● ● ● ● j ● ● ● ● ● ● ● ● ● ● ● ● j ● + ● ● j ● ● ● ● ● ● ● ● ● ● ● ● ● j ● j j j ● ● j ● j ● ● ● ● ● ● ● ● ● ● j ● ● ● ● ● j ● ● ● ● j ● j ● ● ● [24] ● ● j ● ● ● ● ● ● ● ● j ● ● ● ● j) (j ● ● ● ● j ● ● j) (j ● ● ● ● j ● ● ● ● j ● + ● ● ● ● ● ● ● ● ● ● ● j ● j ● ● ● ● j ● ● ● ● ● ● ● ● ● ● ● ● ● ● j) (j ● ● + j ● ● j ● ● ● ● j ● + j j Haplomitrium Nees hookeri (Sm.) Nees Harpalejeunea (Spruce) Schiffn. molleri (Steph.) Grolle ES j Geocalyx Nees graveolens (Schrad.) Nees Gongylanthus Nees ericetorum (Raddi) Nees EG Hepatics and Anthocerotes of the Mediterranean, an annotated checklist ● ● ● ● ● ● ● ● ● ● ● ● ● j) (j ● ● ● ● R.M. Ros et al. 368 AL Harpanthus Nees flotovianus (Nees) Nees scutatus (F. Weber & D. Mohr) Spruce BA BG j + BL CN CO CT CY DZ EG ES HR IL IT 369 JO TR LB LY MA MD ME MT PT RS SA SC SI SY TN j j) (j j j ● j ● ● j ● Jamesoniella (Spruce) F. Lees autumnalis (DC.) Steph. undulifolia (Nees) Müll. Frib. MK ● j ● ● Hygrobiella Spruce laxifolia (Hook.) Spruce ● ● j ● ● j) (j ● ● ● Jubula Dumort. hutchinsiae (Hook.) Dumort. [27] subsp. hutchinsiae subsp. javanica (Steph.) Verd. ●[27] ● ● ● [27] ● ● [27] ● ● ● ● ● ● ● j ● ● ● ● ● ● ● j ● ● ● ● ● ● j ● ● ● ● ● ● ● ● j ● ● ● ● ● ● j ● ● j ● j ● ● j ● ● ● ● ● j ● j ● j j ● ● ● j j ● ● ● j ● j ● j ● ● j ● ● ● ● ● ● ● ● ● j ● [1] ● ● ● ● j (●) ● ● ● j ● ● ● j j) (j j ● ● ● j ● ● [1] ● j j ● ● [1] j ● ● ● ● ● ● ● j ● ● ● ● j) (j ● ● ● j ● ● j ● j ● j ● j) (j ● ● ● ● j) (j j j ● j ● ● ● ● j Kurzia G. Martens pauciflora (Dicks.) Grolle sylvatica (A. Evans) Grolle trichoclados (Müll. Frib.) Grolle Leiocolea (Müll. Frib.) H. Buch badensis (Gottsche) Jörg. bantriensis (Hook.) Jörg. [31] collaris (Nees) Schljakov [31] gillmanii (Austin) A. Evans GR ● ● Heteroscyphus Schiffn. denticulatus (Mitt.) Schiffn. Jungermannia L. [28] [29] atrovirens Dumort. borealis Damsh. & Vá≈a caespiticia Lindenb. callithrix Lindenb. & Gottsche caucasica Vá≈a confertissima Nees exsertifolia Steph. subsp. cordifolia (Dumort.) Vá≈a gracillima Sm. handelii (Schiffn.) Amakawa hyalina Lyell [30] leiantha Grolle lignicola (Schiffn.) Grolle obovata Nees paroica (Schiffn.) Grolle polaris Lindb. pumila With. sphaerocarpa Hook. subelliptica (Lindb. ex Kaal.) Levier subulata A. Evans subtilissima (Schiffn.) Grolle FR Hepatics and Anthocerotes of the Mediterranean, an annotated checklist j j ● ● ● ● ● [1] ● ● ● ● ● ● j ● ● ● ● ● ● ● ● ● ● ● ● ● j) (j ● ● ● ● j ● j ● ● ● ● ● (●) ● j ● ● + ● ● ● ● R.M. Ros et al. 370 AL heterocolpos (Thed. ex Hartm.) H. Buch turbinata (Raddi) H. Buch Lejeunea Lib. canariensis (Steph.) Steph. cavifolia (Ehrh.) Lindb. [33] eckloniana Lindenb. flava (Sw.) Nees hibernica Bischl. et al. ex Grolle lamacerina (Steph.) Schiffn. mandonii (Steph.) Müll. Frib. patens Lindb. [33] BA BG BL j ● j ● ● CN CO CT CY DZ ● ● ● ● ● ● ● HR ES FR GR ● ● ● j ● ● ● ● j ● ● ● ● ● j ● EG IL IT Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 371 JO TN TR LB LY MA MD ME ● ● ● ● j [32] ● ● ● ● ● ● Lunularia Adans. cruciata (L.) Lindb. PT j j RS SA SC j ● SI SY j ● ● ● ● ● ● ● ● j ● ● ● ● ● ● j ● ● ● ● [34] ● ● ● ● ● ● j ● ● ● ● (●) j ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● j ● ● ● ● ● Leptoscyphus Mitt. cuneifolius (Hook.) Mitt. Lophozia (Dumort.) Dumort. [36] ascendens (Warnst.) R. M. Schust. bicrenata (Schmidel ex Hoffm.) Dumort. capitata (Hook.) Macoun decolorans (Limpr.) Steph. elongata Steph. excisa (Dicks.) Dumort. grandiretis (Lindb. ex Kaal.) Schiffn. incisa (Schrad.) Dumort. laxa (Lindb.) Grolle longidens (Lindb.) Macoun longiflora (Nees) Schiffn. obtusa (Lindb.) A. Evans opacifolia Culm. ex Meyl. perssonii H. Buch & S. W. Arnell polaris (R. M. Schust.) R. M. Schust. & Damsh. silvicola H. Buch [37] sudetica (Nees ex Huebener) Grolle ventricosa (Dicks.) Dumort. [38] wenzelii (Nees) Steph. MT ● ● Lepidozia (Dumort.) Dumort. [35] cupressina (Sw.) Lindenb. reptans (L.) Dumort. Lophocolea (Dumort.) Dumort. bidentata (L.) Dumort. fragrans (Moris & De Not.) Gottsche et al. heterophylla (Schrad.) Dumort. minor Nees MK ● j j ● ● ● ● ● ● ● ● j ● j ● j j ● j ● ● ● ● ● ● ● ● ● ● ● ● ● j ● ● ● ● ● ● j ● ● j ● j ● j ● ● ● ● ● ● ● ● ● j ● ● ● j ● ● ● j ● j) (j ● j ● [1] ● [1] j ● ● [1] ● ● ● ● ● ● ● j ● ● j ● ● j ● j ● ● ● ● j ● ● ● ● ● ● ● ● j j j + j j ● ● ● ● ● ● j ● j ● j ● (●) ● ● j ● j j) (j ● j ● j ● j j ● ● j j ● ● ● ● ● ● ● j) (j ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● j ● j) (j j ● j + j j ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● j ● ● ● ● ● ● ● j j + ● ● ● j ● j ● ● j j j ● ● j) (j ● j ● j ● ● j ● ● ● ● ● ● ● ● ● ● ● ● ● ● j ● ● ● ● R.M. Ros et al. 372 AL Mannia Opiz androgyna (L.) A. Evans californica (Gottsche ex Underw.) L. C. Wheeler fragrans (Balbis) Frye & L. Clark [10] pilosa (Hornem.) Frye & L. Clark triandra (Scop.) Grolle Marchantia L. debilis Goegel paleacea Bertol. polymorpha L. subsp. montivagans Bischl. & Boisselier subsp. polymorpha subsp. ruderalis Bischl. & Boisselier BA BG ● Moerckia Gottsche blyttii (Moerch ex Hornem.) Brockm. hibernica (Hook.) Gottsche CO CT CY DZ TN TR ● ● EG ES FR GR HR IL IT JO LB ● ● ● ● ● ● ● ● j j) (j j LY MA MD ME MK MT PT RS SA SC ● ● SI SY j ● ● ● ● ● ● j j ● ● ● ● ●l ● ● ● ● ● ● j ● ● ● ● j ● ● ● ● ● ● ● ● ● j ● ● ● ● j ● ● + ● ● ● ● ● ● ● j ● ● ● ● ● ● ● ● ● j ● ● j ● ● [1] ● ● ● ● ● + (●) + ● + ● + ● ● ● j ● ● ● ● ● ● ● ● ● j + j ● ● + ● ● ● ● ● ● ● ● ● ● ● ● ● j ● ● ● ● ● ● ● j ● j ● ● ● ● ● ● ● ● ● ● ● ● ● ● j ● ● ● ● ● (●) j) (j ● j ● j (●) ● j ● ● ● ● ● ● ● j ● j ● ● j ● j ● ● ● ● ● ● j ● ● ● ● ● ● ● ● ● ● ● j ● ● ● j ● ● ● ● ● ● ● ● ● j ● ● ● ● ● ● ● ● ● j) (j j j ● + ● ● ● ● (●) ● j ● + j j ● ● ● ● ● j ● ● ● ● ● ● j ● j ● ● j ● ● ●l ● Marsupella Dumort. adusta (Nees emend. Limpr.) Spruce alpina (Gottsche ex Husn.) Bernet boeckii (Austin) Kaal. brevissima (Dumort.) Grolle commutata (Limpr.) Bernet condensata (Ångstr. ex C. Hartm.) Kaal. emarginata (Ehrh.) Dumort. funckii (F. Weber & D. Mohr) Dumort. profunda Lindb. sparsifolia (Lindb.) Dumort. sphacelata (Gieseke ex Lindenb.) Dumort. sprucei (Limpr.) Bernet Microlejeunea Steph. ulicina (Taylor) A. Evans CN 373 [39] Marchesinia mackaii (Hook.) Gray Metzgeria Raddi conjugata Lindb. furcata (L.) Dumort. leptoneura Spruce simplex Lorb. ex Müll. Frib. temperata Kuwah. violacea (Ach.) Dumort. [40] BL Hepatics and Anthocerotes of the Mediterranean, an annotated checklist ● ● ● ● ● ● ● ● ● ● j j ● (●) ● ● ● j) (j ● ● ● j ● ● ● ● ● + ● j j ● R.M. Ros et al. 374 AL Mylia Gray anomala (Hook.) Gray taylorii (Hook.) Gray BA BG j ● CN CO CT CY DZ EG ● ● j) (j j IL ● IT JO TR LB LY MA MD ME MK MT PT RS SA SC SY TN j ● j SI ● j ● ● ● ● j) (j ● ● ● ● ● j ● ● ● ● ● ● ● ● [1] ● j ● j j ● j ● ● ● j j ● ● ● ● ● j ● ● ● ● j ● ● ● ● ● ● ● j ● j j ● j ● j ● + ● j ● ● ● ● ● ● ● ● ● j ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● j ● ● ● ● ● ● ● ● ● ● ●[42] ● ● ● ● ● j ● j ● ● ● j ● [1] ● ● ● j ● j j ● ● ● ● + ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● + ● (●) ● ● ● ● j Petalophyllum Nees & Gottsche ex Lehm. ralfsii (Wilson) Nees & Gottsche j ● ● ● Peltolepis Lindb. quadrata (Saut.) Müll. Frib. Plagiochila (Dumort.) Dumort. [44] asplenioides (L. emend. Taylor) Dumort. [45] HR ● Pallavicinia Gray lyellii (Hook.) Carruth. Plagiochasma Lehm. & Lindenb. appendiculatum Lehm. & Lindenb. microcephalum (Steph.) Steph. var. tunesicum Bischl. rupestre (J. R. Forst. & G. Forst.) Steph. GR ● ● Odontoschisma (Dumort.) Dumort. [12] denudatum (Mart.) Dumort. elongatum (Lindb.) A. Evans macounii (Austin) Underw. prostratum (Sw.) Trevis. sphagni (Dicks.) Dumort. Pellia Raddi endiviifolia (Dicks.) Dumort. epiphylla (L.) Corda [43] neesiana (Gottsche) Limpr. FR 375 ● ● Nowellia Mitt. curvifolia (Dicks.) Mitt. Pedinophyllum (Lindb.) Lindb. interruptum (Nees) Kaal. ES ● Nardia Gray breidleri (Limpr.) Lindb. compressa (Hook.) Gray geoscyphus (De Not.) Lindb. [41] insecta Lindb. [41] scalaris Gray Oxymitra Bisch. ex Lindenb. incrassata (Brot.) Sérgio & Sim-Sim BL Hepatics and Anthocerotes of the Mediterranean, an annotated checklist j ● ● ● ● ● j ● ● ● ● ● ● ● ● ● ● ● ● ● ● j ● ● ● ● j ● ● j) (j j ● ● j ● j ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● (●) ● ● ● ● j) (j ● ● ● ● j ● ● ● (●) j ● ● + j) (j ● ● ● j ● ● (●) (●) ● ● ● ● ● R.M. Ros et al. 376 AL bifaria (Sw.) Lindenb. britannica Paton exigua (Taylor) Taylor heterophylla Lindenb. ex Lehm. [46] maderensis Gottsche ex Steph. [47] porelloides (Torrey ex Nees) Lindenb. [45] punctata (Taylor) Taylor retrorsa Gottsche spinulosa (Dicks.) Dumort. stricta Lindenb. virginica A. Evans BA BG BL CN CO CT CY DZ ● EG GR HR IL IT Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 377 JO TR LB LY MA MD ME ES FR ● ● ● ● ● ● ● MK MT PT RS SA SC SI SY TN [1] ● ● ● ● + ● j j ● ● ● ● + j ● ● ● ● ● ● ● ● ● ● [1] ● ● ● ● j ● ● ● ● ● ● ● ● ● j ● ● ● Pleurocladula Grolle albescens (Hook.) Grolle ● ● ● ● ● ● j ● ● ● ● ● ● ● ● ● ● ● ● j) (j Porella L. arboris-vitae (With.) Grolle ● ● xbaueri (Schiffn.) C. E. O. Jensen canariensis (F. Weber) Underw. cordaeana (Huebener) Moore inaequalis (Gottsche ex Steph.) Perss. obtusata (Taylor) Trevis. pinnata L. platyphylla (L.) Pfeiff. ● ● ● (●) [48] j ● ● ● j ● Ptilidium Nees ciliare (L.) Hampe pulcherrimum (Weber) Vain. Radula Dumort. [52] aquilegia (Hook. f. & Taylor) Gottsche et al. carringtonii J. B. Jack complanata (L.) Dumort. holtii Spruce jonesii Bouman et al. lindenbergiana Gottsche ex C. Hartm. nudicaulis Steph. visianica C. Massal. wichurae Steph. Reboulia Raddi hemisphaerica (L.) Raddi [51] j ● ● ● ● ● [49] ● ● ● ● j ● ● ● j ● [50] ● ● ● ● ● ● ● ● ● ● ● j ● ● ● ● j ● ● j ● ● ● ● ● j ● ● [16] ● ● ● + ● ● ● ● ● ● ● [48] ● ● ● j ● ● j ● ● ● ● ● ● ● ● j ● ● ● ● [50] ● ● ● ● ● ● [1] ● ● ● ● ● ● ● ● ● ● j ● j ● ● ● ● j ● ● Prasanthus Lindb. suecicus (Gottsche) Lindb. Preissia Corda quadrata (Scop.) Nees [51] [1] ● ● j ● ● ● j j ● ● ● ● ● ● j ● j ● j j ● ● ● j ● ● ● ● ● ● ● ● ● ● + j ● ● ● ● ● [1] ● ● ● j ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● j ● ● ● ● j ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● j ● ● j ● ● ● ● ● ● ● ● ● ● ● j ● ● ● ● ● ● ● j j ● ● j R.M. Ros et al. 378 AL Riccardia Gray chamedryfolia (With.) Grolle incurvata Lindb. latifrons (Lindb.) Lindb. multifida (L.) Gray palmata (Hedw.) Carruth. Riccia L. [53] [54] [55] atlantica Sérgio & Perold atromarginata Levier beyrichiana Hampe ex Lehm. bicarinata Lindb. bifurca Hoffm. [56] breidleri Jur. ex Steph. canaliculata Hoffm. cavernosa Hoffm. ciliata Hoffm. [57] ciliifera Link ex Lindenb. congoana Steph. crinita Taylor [57] crozalsii Levier crustata Trab. crystallina L. emend. Raddi duplex Lorb. fluitans L. frostii Austin glauca L. var. ciliaris Warnst. var. glauca gougetiana Durieu & Mont. var. armatissima Levier ex Müll. Frib. var. gougetiana huebeneriana Lindenb. lamellosa Raddi ligula Steph. macrocarpa Levier melitensis C. Massal. michelii Raddi nigrella DC. papillosa Moris perennis Steph. rhenana Lorb. ex Müll. Frib. sommieri Levier sorocarpa Bisch. var. heegii Schiffn. var. sorocarpa subbifurca Warnst. ex Croz. teneriffae S. W. Arnell trabutiana Steph. warnstorfii Limpr. ex Warnst. var. subinermis Warnst. var. warnstorfii BA j BG BL ● CN CO CT ● ● ● CY DZ EG j j j j j j j ● ● GR HR ● j IL IT ● ● ● [1] ● ● j ● j ● ● ● ● ● ● j ● ● ● ● j ● ● ● j ● ● ● ● ● ● j ● ● ● j + j ● ● ● ● j ● ● ● ● ● ● ● ● j ● ● FR ● j) (j ● ES ● ● Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 379 JO TN TR j) (j ● LB LY MA MD ME j ● ● MK MT j) (j PT ● ● ● ● j SA SC SI ● ● j ● ● SY ● ● ● ● RS ● j ● (●) j ● j ● ● j ● ● j ● ● ● ● ● j ● j ● ● ● j ● ● ● ● ● + j ● ● ● ● ● j ● ● ● ● ● ● ● ● ● ● ● ● ● j ● [1] ● ● j) (j ● ● ● ● ● ● ● j ● ● ● j ● ● ● ● ● ● ● ● j ● ● + ● + ● ● ● ● ● + ● j) (j j ● ● + ● ● ● j ● ● ● ● + ● + ● ● ● ● j ● ● ● ● ● ● ● ● j ● ● ● j + + ● ● ● + ● ● ● j ● ● ● ● j ● ● ● ● ● ● ● ● ● ● j ● ● ● ● ● j ● ● ● ● + j ● ● ● ● ● j ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● + j ● ● ● ● ● ● + ● ● [1] ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● + ● ● ● ● ● ● ● ● ● ● ● j) (j ● ● ● j ● [1] ● ● ● ● ● ● j j) (j ● + ● ● + j (●) ● ● ● j ● ● ● ● j ● j ● ● j ● ● + ● ● ● ● ● ● j) (j + ● ● ● + [58] ● j ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● j) (j ● ● j [1] ● j ● ● ● + ● ● ● ● ● ● ● j ● ● ● ● ● j ● ● ● ● + j ● ● ● j ● + ● ● ● j ● + ● ● ● ● ● ● ● ● ● + ● ● ● ● ● ● + ● ● ● ● j ● ● ● j ● j ● ● j ● ● ● ● + ● ● ● ● ● ● ● ● ● ● ● + ● ● ● ● ● ● ● ● ● ● ● j ● ● ● ● j ● j ● + ● ● ● ● j ● ● ● ● ● ● [1] ● j ● ● ● ● j ● j ● ● ● ● j j ● ● + ● ● ● ● j ● j ● ● ● ● ● ● ● ● ● ● ● ● ● j ● ● ● ● ● ● ● ● + ● ● j + ● ● ● ● ● ● ● ● ● ● ● ● ● j + ● ● ● ● + j ● ● ● ● + + ● ● ● ● ● j j j) (j ● ● ● ● ● [1] ● j ● j j ● ● ● j) (j ● 380 R.M. Ros et al. AL Ricciocarpos Corda natans (L.) Corda BA BG j ● Riella Mont. affinis M. Howe & Underw. bialata Trab. cossoniana Trab. cyrenaica Maire helicophylla (Bory & Mont.) Mont. notarisii (Mont.) Mont. numidica Trab. [60] parisii Gottsche sersuensis Trab. [60] BL CN CO CT CY DZ EG j ● ES FR GR ● ● j IL IT ● ● [59] ● ● ● 381 JO TN TR ● ● LB LY MA MD ME j ● MK MT PT RS SA SC SI (●) j SY j ● ● [1] ● + ● ● ● ● ● ● j j j ● ● ● ● ● j) (j ● ● ● ● ● ● j j ● j ● j j j ● j ● ● + ● ● j [1] j j Saccogyna Dumort. viticulosa (L.) Dumort. ● ● Sauteria Nees alpina (Nees) Nees Scapania (Dumort.) Dumort. aequiloba (Schwägr.) Dumort. apiculata Spruce aspera Bernet & M. Bernet brevicaulis Taylor calcicola (Arnell & J. Perss.) Ingham carinthiaca J. B. Jack ex Lindb. compacta (A. Roth) Dumort. [63] crassiretis Bryhn curta (Mart.) Dumort. cuspiduligera (Nees) Müll. Frib. gracilis Lindb. gymnostomophila Kaal. helvetica Gottsche irrigua (Nees) Nees lingulata H. Buch mucronata H. Buch nemorea (L.) Grolle obscura (Arnell & C. E. O. Jensen) Schiffn. paludicola Loeske & Müll. Frib. paludosa (Müll. Frib.) Müll. Frib. praetervisa Meyl. scandica (Arnell & H. Buch) Macvicar scapanioides (C. Massal.) Grolle subalpina (Nees ex Lindenb.) Dumort. uliginosa (Sw. ex Lindenb.) Dumort. umbrosa (Schrad.) Dumort. undulata (L.) Dumort. verrucosa Heeg HR Hepatics and Anthocerotes of the Mediterranean, an annotated checklist ● j ● ● ● ● j ● ● ● ● ● ● ● ● ● j ● j j ● ● ● j) (j ● j [1] j ● ● ● j ● ● [1] j) (j ● ● j) (j ● j ● +[61] ● j ● ● ● ● [62] ● ● ● j ● ● ● ● ● ● j ● ● ● ● ● j j j j ● ● ● ● ● j [1] ● j ● ● j j ● ● ● ● ● j) (j ● ● ● j) (j j ● ● ● ● j ● j ● ● ● (●) ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● j ● ● j) (j j ● ● ● ● j ● j j j ● j) (j ● ● j ● j ● ● ● ● ● ● j ● ● j j ● ● j ● ● j ● ● j ● ● ● + ● ● ● j ● ● ● ● ● ● ● (●) ● ● ● ● ● ● ● ● ● ● j ● ● ● ● ● j ● ● ● j ● j) (j ● (●) ● ● ● ● ● ● ● j ● ● ● ● j ● j ● j ● j ● ● j ● ● j j ● ● ● ● j j ● ● ● R.M. Ros et al. 382 BL CN CO CT CY DZ ● ● ● ● ● ● ● ● ● ● Sphaerocarpos Boehm. michelii Bellardi stipitatus Bisch. ex Lindenb. ● ● ● ● texanus Austin ● ● ● ● ● ● ● ● ● ● ● + ● ● ● Southbya Spruce nigrella (De Not.) Henriq. tophacea (Spruce) Spruce Targionia L. hypophylla L. lorbeeriana Müll. Frib. AL BA ● j BG ● ● j j TN TR ES FR GR HR IL IT JO LB LY ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● j ● ● EG ● ● ● MA MD ME j ● ● j ● ● MK MT ● j ● PT RS SA SC SI SY ● ● ● j ● ● ● j) (j ● j ● ● ● ● ● ● ● j) (j [64] ● + ● ● ● j j ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● j j) (j ● [1] ● ● ● ● ● ● ● j ● ● ● ● ● ● ● j ● ● ● j ● ● j ● j ● ● Tetralophozia (R. M. Schust.) Schljakov filiformis (Steph.) Urmi setiformis (Ehrh.) Schljakov ● ● (●) Trichocolea Dumort. tomentella (Ehrh.) Dumort. Tylimanthus Mitt. madeirensis Grolle & Perss. 383 ● Telaranea Spruce ex Schiffn. azorica (H. Buch & Perss.) Pócs ex Schumacker & Vá≈a [65] europaea J. J. Engel & G. L. S. Merril [66] [67] Tritomaria Schiffn. ex Loeske exsecta (Schmidel ex Schrad.) Loeske exsectiformis (Breidl.) Loeske polita (Nees) Jörg. quinquedentata (Huds.) H. Buch scitula (Taylor) Jörg. Hepatics and Anthocerotes of the Mediterranean, an annotated checklist ● j j ● ● ● ● ● ● ● ● ● ● ● ● j j) (j ● ● j ● j ● j ● ● ● ● j ● ● ● ● + ● ● ● ● [68] j j ● j ● ● ● j) (j j ● ● ● ● j j j 384 R.M. Ros et al. List of synonyms and names used in the studied area Symbols used in the list: l Name used in the Mediterranean literature, but not found in reference works and there are uncertainties about the correct synonymy and authorship. ☞ The author/s of the name is/are doubtful. ? Name used in the Mediterranean literature, but not found in reference works and the authorship is therefore uncertain, although there is no doubt about the taxon referred (the name was probably created in a floristic publication). # Name used in the Mediterranean literature and found in independent reference works, but the synonymy could not be confirmed. $ Synonym used in the Mediterranean literature, but not formally published (the name was undoubtedly created in a floristic publication) therefore no authors are indicated. Anthocerotae Anthoceros beltrani Casares-Gil [4] Anthoceros bulbiculosus Brot. = Phymatoceros bulbiculosus (Brot.) Stotler, W. T. Doyle & Crand.-Stotl. Anthoceros caespiticius De Not. = Anthoceros punctatus L. Anthoceros carolinianus Michx. = Phaeoceros carolinianus (Michx.) Prosk. Anthoceros constans Lindb. = Anthoceros punctatus L. Anthoceros crispulus (Mont.) Douin = Anthoceros punctatus L. Anthoceros crispulus auct. non (Mont.) Douin = Anthoceros agrestis Paton Anthoceros dichotomus Raddi = Phymatoceros bulbiculosus (Brot.) Stotler, W. T. Doyle & Crand.-Stotl. Anthoceros dichotomus f. stricta Casares-Gil = Phymatoceros bulbiculosus (Brot.) Stotler, W. T. Doyle & Crand.-Stotl. Anthoceros dichotomus var. gussonei Zodda = Phaeoceros laevis (L.) Prosk. [5] Anthoceros husnotii Steph. = Anthoceros punctatus L. Anthoceros laevis L. = Phaeoceros laevis (L.) Prosk. Anthoceros mandonii Steph. = Anthoceros caucasicus Steph. Anthoceros multifidus L. = Anthoceros punctatus L. [2] Anthoceros multifidus auct. = Anthoceros agrestis Paton [2] Anthoceros multilobulus Lindb. = Anthoceros punctatus L. Anthoceros polymorphus Raddi = Anthoceros punctatus L. Anthoceros punctatus var. cavernosus auct. non (Nees) Gottsche et al. = Anthoceros agrestis Paton Anthoceros punctatus var. crispatus Mont. = Anthoceros punctatus L. Anthoceros punctatus var. crispatus auct. non Mont. = Anthoceros agrestis Paton Anthoceros punctatus var. crispulus Mont. = Anthoceros punctatus L. Anthoceros punctatus var. crispulus auct. non Mont. = Anthoceros agrestis Paton Anthoceros punctatus var. husnotii (Steph.) Zodda = Anthoceros punctatus L. Anthoceros punctatus f. multifidus (L.) Nees = Anthoceros punctatus L. [2] Anthoceros punctatus f. multifidus auct. non (L.) Nees = Anthoceros agrestis Paton [2] Anthoceros punctatus var. multifidus (L.) Schmidel ex Swartz = Anthoceros punctatus L. [2] Anthoceros punctatus var. multifidus auct. non (L.) Schmidel ex Swartz = Anthoceros agrestis Paton [2] Anthoceros validus Steph. = Phaeoceros carolinianus (Michx.) Prosk. Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 385 # Anthoceros torreyi Austin = Anthoceros punctatus L. Aspiromitus madeirensis Steph. = Anthoceros caucasicus Steph. Carpobolus orbicularis (Schwein.) Schwein. = Notothylas orbicularis (Schwein.) A. Gray Carpolipum fertile Lehm. = Notothylas orbicularis (Schwein.) A. Gray Carpolipum valvatum (Sull.) Nees = Notothylas orbicularis (Schwein.) A. Gray Chamaeceros fertilis (Lehm.) Milde = Notothylas orbicularis (Schwein.) A. Gray Notothylas fertilis (Milde) Lehm. = Notothylas orbicularis (Schwein.) A. Gray Notothylas valvata Sull. = Notothylas orbicularis (Schwein.) A. Gray Phaeoceros bulbiculosus (Brot.) Prosk. = Phymatoceros bulbiculosus (Brot.) Stotler, W.T. Doyle & Crand.-Stotl. Phaeoceros dichotomus (Raddi) Prosk. = Phymatoceros bulbiculosus (Brot.) Stotler, W.T. Doyle & Crand.-Stotl. Phaeoceros laevis subsp. carolinianus (Michx.) Prosk. = Phaeoceros carolinianus (Michx.) Prosk. Hepaticae Achiton quadratum (Scop.) Corda = Preissia quadrata (Scop.) Müll. Frib. [51] Acolea adusta (Nees emend. Limpr.) C. Massal. hom. illeg. non (Gray) Trevis. = Marsupella adusta (Nees emend. Limpr.) Spruce Acolea alpina (Gottsche ex Husn.) C. Massal. = Marsupella alpina (Gottsche ex Husn.) Bernet Acolea brevissima Dumort. = Marsupella brevissima (Dumort.) Grolle Acolea concinnata (Lightf.) Dumort. = Gymnomitrion concinnatum (Lightf.) Corda Acolea conferta (Limpr.) C. Massal. & Carestia = Marsupella brevissima (Dumort.) Grolle Acolea corallioides (Nees) Dumort. = Gymnomitrion corallioides Nees Acolea revoluta (Nees) Steph. = Apomarsupella revoluta (Nees) R. M. Schust. Acolea varians (Lindb.) Steph. = Marsupella brevissima (Dumort.) Grolle Aitonia italica (Sassi) Lindb. = Plagiochasma rupestre (J. R. Forst. & G. Forst.) Steph. Alicularia breidleri Limpr. = Nardia breidleri (Limpr.) Lindb. Alicularia compressa (Hook.) Nees = Nardia compressa (Hook.) Gray Alicularia geoscypha De Not. = Nardia geoscyphus (De Not.) Lindb. Alicularia geoscypha f. insecta (Lindb.) Müll. Frib. = Nardia insecta Lindb. Alicularia minor (Nees) Limpr. = Nardia geoscyphus (De Not.) Lindb. Alicularia pachyphylla De Not. = Nardia compressa (Hook.) Gray Alicularia rotaeana De Not. = Nardia scalaris Gray Alicularia scalaris (Gray) Corda = Nardia scalaris Gray Alicularia scalaris var. minor (Nees) Nees = Nardia geoscyphus (De Not.) Lindb. Anastrophyllum minutum var. grande (Gottsche ex Lindb.) R. M. Schust. = Anastrophyllum minutum (Schreb.) R. M. Schust. Anastrophyllum minutum var. weberi (Mart.) Kartt. = Anastrophyllum minutum (Schreb.) R. M. Schust. Anastrophyllum myriocarpum (Carrington) R. M. Schust. ex Vá≈a = Eremonotus myriocarpus (Carrington) Pearson Anastrophyllum nardioides (Lindb.) Kaal. = Anastrophyllum assimile (Mitt.) Steph. Anastrophyllum reichardtii (Gottsche in Jur.) Steph. = Anastrophyllum assimile (Mitt.) Steph. Aneura incurvata (Lindb.) Steph. = Riccardia incurvata Lindb. Aneura latifrons Lindb. = Riccardia latifrons (Lindb.) Lindb. Aneura multifida (L.) Dumort. = Riccardia multifida (L.) Gray 386 R.M. Ros et al. Aneura multifida var. major (Dicks. ex Hook.) Nees = Riccardia chamedryfolia (With.) Grolle ☞ Aneura multifida var. rivularis Rabenh. = Riccardia chamedryfolia (With.) Grolle Aneura palmata (Hedw.) Dumort. = Riccardia palmata (Hedw.) Carruth. Aneura pinguis var. angustior (Hook.) Dumort. = Aneura pinguis (L.) Dumort. Aneura pinnatifida (Sw.) Dumort. hom. illeg. = Riccardia chamedryfolia (With.) Grolle Aneura sinuata (Dicks. ex Hook.) Dumort. = Riccardia chamedryfolia (With.) Grolle Anomomarsupella cephalozielloides R. M. Schust. = Eremonotus myriocarpus (Carrington) Pearson Anthelia andina Herzog = Anthelia juratzkana (Limpr.) Trevis. Anthelia julacea subsp. juratzkana (Limpr.) Meyl. = Anthelia juratzkana (Limpr.) Trevis. Anthelia nivalis (Sw. ex F. Weber & D. Mohr) Lindb. [7] Anthelia phyllacantha C. Massal. & Carestia = Cephaloziella phyllacantha (C. Massal. & Carestia) Müll. Frib. Anthelia setiformis (Ehrh.) Dumort. = Tetralophozia setiformis (Ehrh.) Schljakov Anthelia turneri (Hook.) Dumort. = Cephaloziella turneri (Hook.) Müll. Frib. Antoiria vulgaris Raddi nom. illeg. = Porella platyphylla (L.) Pfeiff. Antrocephalus italicus Sassi = Plagiochasma rupestre (J. R. Forst. & G. Forst.) Steph. Aphanolejeunea teotonii V. Allorge & Ast = Aphanolejeunea sintenisii Steph. Aplozia amplexicaulis (Dumort.) Dumort. = Jungermannia sphaerocarpa Hook. Aplozia anomala (Hook.) Warnst. = Mylia anomala (Hook.) Gray Aplozia atrovirens (Dumort.) Dumort. = Jungermannia atrovirens Dumort. Aplozia autumnalis (DC.) Heeg = Jamesoniella autumnalis (DC.) Steph. Aplozia confertissima (Nees) Dumort. = Jungermannia confertissima Nees Aplozia cordifolia Dumort. = Jungermannia exsertifolia subsp. cordifolia (Dumort.) Vá≈a Aplozia crenulata (Mitt.) Lindb. = Jungermannia gracillima Sm. Aplozia crenulata var. inundata (Schiffn.) Macvicar = Jungermannia gracillima Sm. Aplozia crenulata var. gracillima (Sm.) Heeg = Jungermannia gracillima Sm. Aplozia gracillima (Sm.) Dumort. = Jungermannia gracillima Sm. Aplozia hyalina (Lyell) Dumort. = Jungermannia hyalina Lyell Aplozia lanceolata (L.) Dumort. = Jungermannia atrovirens Dumort. Aplozia lanceolata auct. non (L.) Dumort. = Jungermannia leiantha Grolle Aplozia lurida (Dumort.) Dumort. = Jungermannia sphaerocarpa Hook. Aplozia lurida sensu Breidl. = Jungermannia confertissima Nees Aplozia nana (Nees) Breidl. = Jungermannia sphaerocarpa Hook. Aplozia oblongifolia (Müll. Frib.) Jörg. = Jungermannia pumila With. Aplozia obovata (Nees) Loeske = Jungermannia obovata Nees Aplozia polaris (Lindb.) Bryhn & Kaal. = Jungermannia polaris Lindb. Aplozia pumila (With.) Dumort. = Jungermannia pumila With. Aplozia riparia (Taylor) Dumort. = Jungermannia atrovirens Dumort. Aplozia riparia var. potamophila Müll. Arg. ex Bernet = Jungermannia atrovirens Dumort. Aplozia riparia var. rivularis Bernet = Jungermannia atrovirens Dumort. Aplozia scalariformis (Nees) Dumort. = Jungermannia confertissima Nees Aplozia schiffneri Loitl. = Jungermannia polaris Lindb. Aplozia sphaerocarpa (Hook.) Dumort. = Jungermannia sphaerocarpa Hook. Aplozia sphaerocarpa var. nana (Nees) Macvicar = Jungermannia sphaerocarpa Hook. Aplozia sphaerocarpoidea (De Not.) Dumort. = Jungermannia atrovirens Dumort. Aplozia tristis (Nees) Dumort. = Jungermannia atrovirens Dumort. Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 387 Asterella bonjeanii (De Not.) Trevis. = Asterella lindenbergiana (Corda ex Nees) Arnell Asterella elegans (Spreng.) Trevis. REJECTED FROM THE AREA [8] Asterella fragrans (Schleich.) Trevis. = Asterella saccata (Wahlenb.) A. Evans [10] Asterella hemisphaerica (L.) P. Beauv. = Reboulia hemisphaerica (L.) Raddi Asterella ludwigii (Schwägr.) Underw. ex A. Evans nom. inval. = Mannia triandra (Scop.) Grolle Asterella ludwigii auct. non (Schwägr.) Underw. ex A. Evans nom. inval. = Asterella gracilis (F. Weber) Underw. Asterella nana (Lindenb.) C. Massal. = Athalamia hyalina (Sommerf.) S. Hatt. [9] Asterella pilosa (Wahlenb.) Trevis. = Asterella gracilis (F. Weber) Underw. Aytonia rupestris J. R. Forst. & G. Forst. = Plagiochasma rupestre (J. R. Forst. & G. Forst.) Steph. Barbilophozia baueriana (Schiffn.) Loeske = Barbilophozia hatcheri (A. Evans) Loeske Barbilophozia gracilis (Schleich. ex Steph.) Müll. Frib. = Barbilophozia attenuata (Mart.) Loeske Barbilophozia hatcheri var. grandiretis H. Buch ex Lammes = Barbilophozia rubescens (R. M. Schust. & Damsh.) Kartt. & L. Söderstr. Barbilophozia lycopodioides var. hatcheri (Evans) Schljakov = Barbilophozia hatcheri (A. Evans) Loeske ? Barbilophozia lycopodioides var. parvifolia Schiffn. = Barbilophozia hatcheri (A. Evans) Loeske Barbilophozia obtusa (Lindb.) H. Buch = Lophozia obtusa (Lindb.) A. Evans Bazzania deflexa (Mart.) Trevis. = Bazzania tricrenata (Wahlenb.) Lindb. Bazzania denudata auct. non (Torrey ex Gottsche et al.) Trevis. = Bazzania flaccida (Dumort.) Grolle Bazzania implexa (Nees) Müll. Frib. = Bazzania flaccida (Dumort.) Grolle Bazzania triangularis (Schleich.) Lindb. nom. illeg. = Bazzania tricrenata (Wahlenb.) Lindb. Bazzania triangularis var. implexa (Nees) Zahlbr. = Bazzania flaccida (Dumort.) Grolle $ Bazzania tricrenata var. implexa = Bazzania flaccida (Dumort.) Grolle Bellincinia laevigata (Schrad.) Kuntze = Porella arboris-vitae (With.) Grolle Bellincinia montana Raddi = Porella arboris-vitae (With.) Grolle Bellincinia platyphylla (L.) Kuntze = Porella platyphylla (L.) Pfeiff. Bellincinia porella (Dicks.) Kuntze = Porella pinnata L. Bellincinia thuja (Hook.) Kuntze = Porella arboris-vitae (With.) Grolle Blasia pusilla var. gemmifera Nees = Blasia pusilla L. Blepharostoma arachnoideum var. brevirete (Bryhn & Kaal.) Frye & L. Clark = Blepharostoma trichophyllum subsp. brevirete (Bryhn & Kaal.) R. M. Schust. Blepharostoma trichophyllum var. brevirete Bryhn & Kaal. = Blepharostoma trichophyllum subsp. brevirete (Bryhn & Kaal.) R. M. Schust. Blepharozia ciliaris (L.) Dumort. = Ptilidium ciliare (L.) Hampe ? Blepharozia ciliaris var. wallrothiana Nees = Ptilidium pulcherrimum (Weber) Vain. Blepharozia pulcherrima (Weber) Lindb. = Ptilidium pulcherrimum (Weber) Vain. Blyttia lyellii (Hook.) Endl. ex Gottsche et al. = Pallavicinia lyellii (Hook.) Carruth. Blyttia lyellii var. hibernica (Hook.) Nees = Moerckia hibernica (Hook.) Gottsche Blyttia moerckii Nees = Moerckia blyttii (Moerch ex Hornem.) Brockm. Brissocarpus riccioides Bisch. = Corsinia coriandrina (Spreng.) Lindb. Calycularia blyttii (Moerch ex Hornem.) Steph. = Moerckia blyttii (Moerch ex Hornem.) Brockm. Calycularia hibernica (Hook.) Steph. = Moerckia hibernica (Hook.) Gottsche 388 R.M. Ros et al. Calypogeia adscendens (Nees) Warnst. = Calypogeia muelleriana (Schiffn.) Müll. Frib. Calypogeia ericetorum Raddi = Gongylanthus ericetorum (Raddi) Nees Calypogeia flagellifera Raddi = Gongylanthus ericetorum (Raddi) Nees Calypogeia meylanii H. Buch = Calypogeia integristipula Steph. Calypogeia muelleriana f. sphagnicola (Arnell & J. Perss.) Schljakov = Calypogeia sphagnicola (Arnell & J. Perss.) Warnst. & Loeske Calypogeia neesiana var. laxa Meyl. ex Müll. Frib. = Calypogeia muelleriana (Schiffn.) Müll. Frib. Calypogeia neesiana var. meylanii (H. Buch) R. M. Schust. = Calypogeia integristipula Steph. Calypogeia paludosa Warnst. = Calypogeia sphagnicola (Arnell & J. Perss.) Warnst. & Loeske Calypogeia tenuis (Austin) A. Evans. = Calypogeia sphagnicola (Arnell & J. Perss.) Warnst. & Loeske Calypogeia trichomanis (L.) Corda nom. rejic. = Calypogeia fissa (L.) Raddi Calypogeia trichomanis auct. non (L.) Corda = Calypogeia azurea Stotler & Crotz [11] # Calypogeia trichomanis var. communis Nees = Calypogeia fissa (L.) Raddi Calypogeia trichomanis var. erecta Müll. Frib. = Calypogeia muelleriana (Schiffn.) Müll. Frib. Calypogeia trichomanis var. fissa (L.) De Not. = Calypogeia fissa (L.) Raddi # Calypogeia trichomanis f. papillosa Warnst. = Calypogeia azurea Stotler & Crotz Calypogeia trichomanis var. sphagnicola (Arnell & J. Perss.) Meyl. = Calypogeia sphagnicola (Arnell & J. Perss.) Warnst. & Loeske Calypogeia trichomanis var. sprengelii (Mart.) C. Massal. = Calypogeia fissa (L.) Raddi Calypogeia trichomanis var. suecica (Arnell & J. Perss.) Casares-Gil = Calypogeia suecica (Arnell & J. Perss.) Müll. Frib. Candollea carinata Raddi = Scapania compacta (A. Roth) Dumort. Cephalozia albescens (Hook.) Dumort. = Pleurocladula albescens (Hook.) Grolle # Cephalozia asperifolia f. belsensis Douin = Cephaloziella divaricata (Sm.) Schiffn. Cephalozia baueri (Mart.) C. Massal. = Nowellia curvifolia (Dicks.) Mitt. Cephalozia bicuspidata subsp. ambigua (C. Massal.) R. M. Schust. = Cephalozia ambigua C. Massal. Cephalozia bicuspidata subsp. lammersiana (Huebener) R. M. Schust. = Cephalozia bicuspidata (L.) Dumort. Cephalozia bicuspidata var. alpicola (C. Massal.) C. Massal. & Carestia = Cephalozia pleniceps (Austin) Lindb. Cephalozia bicuspidata var. lammersiana (Huebener) Breidl. = Cephalozia bicuspidata (L.) Dumort. Cephalozia bicuspidata var. setulosa Spruce = Cephalozia bicuspidata (L.) Dumort. Cephalozia bifida Lindb. = Cephaloziella rubella (Nees) Warnst. Cephalozia borealis Lindb. = Cladopodiella francisci (Hook.) Jörg. Cephalozia bryhnii Kaal. = Cephaloziella rubella (Nees) Warnst. Cephalozia byssacea (A. Roth) Dumort. nom. rejic. = Cephaloziella hampeana (Nees) Schiffn. Cephalozia byssacea auct. non (A. Roth) Dumort. = Cephaloziella divaricata (Sm.) Schiffn. Cephalozia columbae Camus = Cephaloziella massalongi (Spruce) Müll. Frib. Cephalozia compacta Warnst. = Cephalozia connivens (Dicks.) Lindb. Cephalozia curvifolia (Dicks.) Dumort. = Nowellia curvifolia (Dicks.) Mitt. Cephalozia dentata (Raddi) Lindb. = Cephaloziella dentata (Raddi) Steph. Cephalozia divaricata (Sm.) Dumort. = Cephaloziella divaricata (Sm.) Schiffn. Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 389 Cephalozia divaricata Heeg nom. inval. non (Sm.) Dumort. = Cephaloziella rubella (Nees) Warnst. Cephalozia divaricata var. scabra M. Howe = Cephaloziella divaricata (Sm.) Schiffn. Cephalozia divaricata var. stellulifera Taylor ex Spruce = Cephaloziella stellulifera (Taylor ex Spruce) Schiffn. Cephalozia divaricata var. starkei auct. non (Funck ex Nees) Spruce nom. illeg. = Cephaloziella divaricata (Sm.) Schiffn. Cephalozia elegans Heeg = Cephaloziella elegans (Heeg) Schiffn. Cephalozia fluitans (Nees) Spruce = Cladopodiella fluitans (Nees) H. Buch Cephalozia francisci (Hook.) Dumort. = Cladopodiella francisci (Hook.) Jörg. Cephalozia granatensis J.B. Jack ex Steph. = Cephaloziella granatensis (J.B. Jack ex Steph.) Fulford Cephalozia helleri (Nees ex Lindenb.) Lindb. nom. illeg. = Anastrophyllum hellerianum (Nees ex Lindenb.) R. M. Schust. Cephalozia hibernica Spruce ex Pearson = Cephalozia crassifolia (Lindenb. & Gottsche) Fulford Cephalozia hypogyna Steph. = Cephaloziella stellulifera (Taylor ex Spruce) Schiffn. Cephalozia integerrima Lindb. = Cephaloziella integerrima (Lindb.) Warnst. Cephalozia lammersiana (Huebener) Carrington = Cephalozia bicuspidata (L.) Dumort. Cephalozia laxifolia (Hook.) Lindb. = Hygrobiella laxifolia (Hook.) Spruce ☞ Cephalozia limprichtii (Warnst.) Steph. = Cephaloziella stellulifera (Taylor ex Spruce) Schiffn. Cephalozia lucens (A. Evans) Steph. = Metahygrobiella lucens (A. Evans) H. A. Miller REJECTED FROM THE AREA # Cephalozia lunulaefolia var. gasilieni Corb. = Cephalozia affinis Lindb. ex Steph. Cephalozia massalongi Spruce = Cephaloziella massalongi (Spruce) Müll. Frib. Cephalozia media Lindb. = Cephalozia lunulifolia (Dumort.) Dumort. # Cephalozia media var. gasilieni Corb. = Cephalozia affinis Lindb. ex Steph. Cephalozia multiflora (Huds.) Lindb. = Cephalozia connivens (Dicks.) Lindb. Cephalozia multiflora Spruce nom. illeg. = Cephalozia lunulifolia (Dumort.) Dumort. Cephalozia myriocarpa (Carrington) Lindb. = Eremonotus myriocarpus (Carrington) Pearson Cephalozia nematodes sensu auct. eur. non Gottsche ex Austin = Teleranea europea J.J. Engel & G.L.S. Merrill [66] Cephalozia notarisiana C. Massal. = Hygrobiella laxifolia (Hook.) Spruce ☞ Cephalozia nudis Lindb. = Cephalozia lunulifolia (Dumort.) Dumort. Cephalozia obtusiloba Lindb. = Cladopodiella fluitans (Nees) H. Buch Cephalozia pallida (Spruce) Pearson = Cephalozia lunulifolia (Dumort.) Dumort. Cephalozia papillosa Douin = Cephaloziella divaricata (Sm.) Schiffn. Cephalozia patula Steph. = Cephaloziella stellulifera (Taylor ex Spruce) Schiffn. Cephalozia phyllacantha (C. Massal. & Carestia) C. Massal. & Carestia = Cephaloziella phyllacantha (C. Massal. & Carestia) Müll. Frib. Cephalozia pleniceps var. macrantha (Kaal. & Nicholson) Müll. Frib. = Cephalozia pleniceps (Austin) Lindb. Cephalozia raddiana (C. Massal.) C. Massal. = Cephaloziella rubella (Nees) Warnst. Cephalozia reclusa (Taylor) Dumort. = Cephalozia catenulata (Huebener) Lindb. Cephalozia serriflora Lindb. = Cephalozia catenulata (Huebener) Lindb. Cephalozia sphagni (Dicks.) Spruce = Odontoschisma sphagni (Dicks.) Dumort. Cephalozia starkei auct. non (Funck ex Nees) Dumort. ex Cogn. nom. illeg. = Cephaloziella divaricata (Sm.) Schiffn. 390 R.M. Ros et al. Cephalozia symbolica (Gottsche) Breidl. = Cephalozia lunulifolia (Dumort.) Dumort. Cephalozia symbolica var. sphagnorum C. Massal. = Cephalozia pleniceps (Austin) Lindb. Cephalozia turneri (Hook.) Lindb. = Cephaloziella turneri (Hook.) Müll. Frib. Cephaloziella algeriensis (Douin) Trab. = Cephaloziella baumgartneri Schiffn. Cephaloziella alpina Douin = Cephaloziella varians (Gottsche) Steph. Cephaloziella arctica Bryhn & Douin = Cephaloziella varians (Gottsche) Steph. Cephaloziella baumgartneri var. algeriensis Douin = Cephaloziella baumgartneri Schiffn. Cephaloziella bifidoides Douin = Cephaloziella rubella (Nees) Warnst. Cephaloziella byssacea (A. Roth) Warnst. nom. rejic. = Cephaloziella hampeana (Nees) Schiffn. Cephaloziella byssacea auct. non (A. Roth) Warnst. = Cephaloziella divaricata (Sm.) Schiffn. ☞ Cephaloziella byssacea var. asperifolia (Taylor) R. M. Schust. = Cephaloziella divaricata (Sm.) Schiffn. # Cephaloziella carnutensis Douin = Cephaloziella divaricata (Sm.) Schiffn. Cephaloziella columbae (Camus) Müll. Frib. = Cephaloziella massalongi (Spruce) Müll. Frib. Cephaloziella compacta (Jörg.) Müll. Frib. = Cephaloziella massalongi (Spruce) Müll. Frib. Cephaloziella curnowii Douin = Cephaloziella hampeana (Nees) Schiffn. Cephaloziella divaricata var. asperifolia (Taylor) Damsh. = Cephaloziella divaricata (Sm.) Schiffn. Cephaloziella divaricata var. polystratosa (R. M. Schust. & Damsh.) Potemkin = Cephaloziella divaricata (Sm.) Schiffn. Cephaloziella divaricata var. rupestris (C. E. O. Jensen) Damsh. nom. inval. = Cephaloziella divaricata (Sm.) Schiffn. Cephaloziella divaricata var. scabra (M. Howe) S.W. Arnell = Cephaloziella divaricata (Sm.) Schiffn. Cephaloziella erosa Limpr. ex Warnst. = Cephaloziella hampeana (Nees) Schiffn. ? Cephaloziella francisci (Hook.) Dumort. = Cladopodiella francisci (Hook.) Jörg. Cephaloziella gallica (Douin) Douin = Cephaloziella hampeana (Nees) Schiffn. [14] Cephaloziella gracillima (Douin) Douin = Cephaloziella stellulifera (Taylor ex Spruce) Schiffn. Cephaloziella hampeana var. subtilis (Velen.) Macvicar = Cephaloziella hampeana (Nees) Schiffn. Cephaloziella latzeliana Schiffn. = Cephaloziella divaricata (Sm.) Schiffn. ☞ Cephaloziella limprichtii Warnst. = Cephaloziella stellulifera (Taylor ex Spruce) Schiffn. Cephaloziella limprichtii var. stellulifera (Taylor ex Carrington & Pearson) Müll. Frib. = Cephaloziella divaricata (Sm.) Schiffn. Cephaloziella massalongi var. algarvica Douin = Cephaloziella massalongi (Spruce) Müll. Frib. Cephaloziella massalongi var. compacta (Jörg.) Müll. Frib. = Cephaloziella massalongi (Spruce) Müll. Frib. Cephaloziella myriantha (Lindb.) Schiffn. = Cephaloziella rubella (Nees) Warnst. [15] Cephaloziella nigrimonasteriensis Douin = Cephaloziella divaricata (Sm.) Schiffn. Cephaloziella papillosa (Douin) Schiffn. = Cephaloziella divaricata (Sm.) Schiffn. Cephaloziella patula (Steph.) Schiffn. = Cephaloziella stellulifera (Taylor ex Spruce) Schiffn. Cephaloziella piriflora Douin = Cephaloziella integerrima (Lindb.) Warnst. Cephaloziella pulchella (C. E. O. Jensen) Douin = Cephaloziella rubella (Nees) Warnst. Cephaloziella raddiana (C. Massal.) Schiffn. = Cephaloziella rubella (Nees) Warnst. Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 391 Cephaloziella rubella var. elegans (Heeg) R. M. Schust. nom. illeg. = Cephaloziella elegans (Heeg) Schiffn. Cephaloziella rubella var. pulchella (C. E. O. Jensen) R. M. Schust. = Cephaloziella rubella (Nees) Warnst. Cephaloziella rubella var. sullivantii (Austin) Müll. Frib. = Cephaloziella rubella (Nees) Warnst. Cephaloziella starkei auct. non (Funck ex Nees) Schiffn. nom. illeg. = Cephaloziella divaricata (Sm.) Schiffn. # Cephaloziella starkei var. examphigastriata Douin ex Müll. Frib. = Cephaloziella divaricata (Sm.) Schiffn. ☞ Cephaloziella starkei var. papillosa (Douin) Casares-Gil = Cephaloziella divaricata (Sm.) Schiffn. Cephaloziella starkei var. scabra (M. Howe) Frye & L. Clark = Cephaloziella divaricata (Sm.) Schiffn. # Cephaloziella starkei var. serriflora Douin = Cephaloziella divaricata (Sm.) Schiffn. ☞ Cephaloziella stellulifera var. gracillima Douin = Cephaloziella stellulifera (Taylor ex Spruce) Schiffn. # Cephaloziella stellulifera var. hericensis Douin = Cephaloziella stellulifera (Taylor ex Spruce) Schiffn. Cephaloziella stellulifera var. gracillima Douin = Cephaloziella stellulifera (Taylor ex Spruce) Schiffn. Cephaloziella stellulifera var. limprichtii (Warnst.) Macvicar = Cephaloziella stellulifera (Taylor ex Spruce) Schiffn. Cephaloziella striatula (C. E. O. Jensen) Douin = Cephaloziella spinigera (Lindb.) Warnst. Cephaloziella subdentata Warnst. = Cephaloziella spinigera (Lindb.) Warnst. Cephaloziella vaucherioides Cout. = Cephaloziella baumgartneri Schiffn. Cephaloziella verrucosa (C.E.O. Jensen) Bryhn & Kaal. hom. illeg. = Cephaloziella varians (Gottsche) Steph. Cesia adusta (Nees emend. Limpr.) Lindb. = Marsupella adusta (Nees emend. Limpr.) Spruce Cesia concinnata (Lightf.) Lindb. = Gymnomitrion concinnatum (Lightf.) Corda l Cesia conferta (Limpr.) Pearson = Marsupella brevissima (Dumort.) Grolle Cesia corallioides (Nees) Carruth. = Gymnomitrion corallioides Nees Cesia revoluta (Nees) Lindb. = Apomarsupella revoluta (Nees) R. M. Schust. Cesia varians (Lindb.) Lindb. = Marsupella brevissima (Dumort.) Grolle Cesius concinnatus (Lightf.) Gray nom. rejic. = Gymnomitrion concinnatum (Lightf.) Corda Chaetopsis trichophylla (L.) Mitt. = Blepharostoma trichophyllum (L.) Dumort. Chandonanthus setiformis (Ehrh.) Lindb. = Tetralophozia setiformis (Ehrh.) Schljakov Cheilolejeunea boaventurae Steph. = Lejeunea eckloniana Lindenb. Chiloscyphus canariensis Bryhn = Lophocolea fragrans (Moris & De Not.) Gottsche et al. Chiloscyphus denticulatus Mitt. = Heteroscyphus denticulatus (Mitt.) Schiffn. Chiloscyphus fragilis (A. Roth) Schiffn. = Chiloscyphus pallescens (Ehrh. ex Hoffm.) Dumort. Chiloscyphus lophocoleoides Nees = Chiloscyphus pallescens (Ehrh. ex Hoffm.) Dumort. Chiloscyphus minor (Nees) J. J. Engel & R. M. Schust. = Lophocolea minor Nees Chiloscyphus pallescens var. fontana Müll. Frib. = Chiloscyphus pallescens (Ehrh. ex Hoffm.) Dumort. Chiloscyphus pallescens var. fragilis (A. Roth) Müll. Frib. = Chiloscyphus pallescens (Ehrh. ex Hoffm.) Dumort. 392 R.M. Ros et al. Chiloscyphus polyanthos var. erectus Boulay = Chiloscyphus pallescens (Ehrh. ex Hoffm.) Dumort. # Chiloscyphus polyanthos var. gracilescens Boulay = Chiloscyphus polyanthos (L.) Corda Chiloscyphus polyanthos var. pallescens (Ehrh. ex Hoffm.) C. Hartm. = Chiloscyphus pallescens (Ehrh. ex Hoffm.) Dumort. Chiloscyphus polyanthos var. rivularis (Schrad.) Nees = Chiloscyphus polyanthos (L.) Corda Chiloscyphus rivularis (Schrad.) Hazsl. = Chiloscyphus polyanthos (L.) Corda Chomiocarpon commutatus (Lindenb.) Lindb. = Preissia quadrata (Scop.) Müll. Frib. Chomiocarpon quadratus (Scop.) Lindb. = Preissia quadrata (Scop.) Nees [51] Cincinnulus argutus (Nees & Mont.) Dumort. = Calypogeia arguta Nees & Mont. Cincinnulus suecicus (Arnell & J. Perss.) Müll. Frib. = Calypogeia suecica (Arnell & J. Perss.) Müll. Frib. Cincinnulus trichomanis (L.) Dumort. = Calypogeia fissa (L.) Raddi Cincinulus trichomanis auct. non (L.) Dumort. = Calypogeia azurea Stotler & Crotz [11] Cincinnulus trichomanis var. fissus (L.) Boulay = Calypogeia fissa (L.) Raddi Cladopus francisci (Hook.) Meyl. = Cladopodiella francisci (Hook.) Jörg. Clevea crassa Trab. nom. inval. = Athalamia hyalina (Sommerf.) S. Hatt. Clevea hyalina (Sommerf.) Lindb. = Athalamia hyalina (Sommerf.) S. Hatt. Clevea rousselliana (Mont.) Leitg. ex Steph. = Athalamia spathysii (Lindenb.) S. Hatt. Clevea spathysii (Lindenb.) Müll. Frib. = Athalamia spathysii (Lindenb.) S. Hatt. Clevea suecica (Lindb.) Lindb. = Athalamia hyalina (Sommerf.) S. Hatt. Clevea trabutiana Steph. = Athalamia hyalina (Sommerf.) S. Hatt. Codonia pusilla (L.) Dumort. nom. inval. = Fossombronia pusilla (L.) Nees Codonia ralfsii (Wilson) Dumort. = Petalophyllum ralfsii (Wilson) Nees & Gottsche Codonia wondraczekii (Corda) Dumort. = Fossombronia wondraczekii (Corda) Lindb. Coleochila anomala (Hook.) Dumort. = Mylia anomala (Hook.) Gray Coleochila stillicidiorum (Raddi) Dumort. = Southbya tophacea (Spruce) Spruce Cololejeunea azorica V. Allorge & Ast = Aphanolejeunea azorica (V. Allorge & Ast) Bernecker & Pócs Cololejeunea inconspicua (Raddi) Schiffn. = Cololejeunea minutissima (Sm.) Schiffn. Cololejeunea madeirensis Schiffn. = Aphanolejeunea madeirensis (Schiffn.) Grolle Cololejeunea microscopica (Taylor) Schiffn. = Aphanolejeunea microscopica (Taylor) A. Evans Cololejeunea teotonii (V. Allorge & Ast) Grolle = Aphanolejeunea sintenisii Steph. ☞ Conocephalum hemisphaericum Dumort. = Preissia quadrata (Scop.) Nees Conocephalum trioicum F. H. Wigg. nom. superfl. = Conocephalum conicum (L.) Dumort. Conocephalus officinalis Trevis. = Conocephalum conicum (L.) Dumort. Conocephalus vulgaris Bisch. = Conocephalum conicum (L.) Dumort. Corbierella algeriensis Douin & Trab. = Exormotheca welwitschii Steph. ? Corbierella welwitschii (Steph.) Douin & Trab. ex Paris = Exormotheca welwitschii Steph. Cordaea blyttii (Moerch ex Hornem.) Corda = Moerckia blyttii (Moerch ex Hornem.) Brockm. Cordaea flotoviana Nees = Moerckia hibernica (Hook.) Gottsche Corsinia coriandrina var. latifrons Trab. nom. inval. = Corsinia coriandrina (Spreng.) Lindb. Corsinia marchantioides Raddi = Corsinia coriandrina (Spreng.) Lindb. Corsinia reticulata (J.F. Gmel.) Dumort. nom. illeg. = Corsinia coriandrina (Spreng.) Lindb. Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 393 Crossocalyx hellerianus (Nees ex Lindenb.) Meyl. = Anastrophyllum hellerianum (Nees ex Lindenb.) R. M. Schust. Crossogyna autumnalis (DC.) Schljakov = Jamesoniella autumnalis (DC.) Steph. Cyathophora commutata (Lindenb.) Trevis. = Preissia quadrata (Scop.) Müll. Frib. ☞ Dichiton calyculatum (Durieu & Mont.) Trevis. = Cephaloziella calyculata (Durieu & Mont.) Müll. Frib. Dichiton gallicum Douin = Cephaloziella hampeana (Nees) Schiffn. [14] Dichiton integerrimum (Lindb.) H. Buch = Cephaloziella integerrima (Lindb.) Warnst. Dichiton perpusillum Mont. nom. illeg. = Cephaloziella calyculata (Durieu & Mont.) Müll. Frib. Dichominum cruciatum (L.) Trevis. = Lunularia cruciata (L.) Lindb. Dicranolejeunea johnsoniana (Mitt.) Vanden Berghen ex S. W. Arnell = Acanthocoleus aberrans (Lindenb. & Gottsche) Kruijt Dilaena blyttii (Moerch ex Hornem.) Dumort. = Moerckia blyttii (Moerch ex Hornem.) Brockm. Dilaena hibernica (Hook.) Dumort. = Moerckia hibernica (Hook.) Gottsche Dilaena lyellii (Hook.) Dumort. = Pallavicinia lyellii (Hook.) Carruth. Diplolaena blyttii (Moerch ex Hornem.) Nees = Moerckia blyttii (Moerch ex Hornem.) Brockm. Diplolaena lyellii (Hook.) Dumort. = Pallavicinia lyellii (Hook.) Carruth. Diplolaena lyellii var. hibernica (Hook.) Nees = Moerckia hibernica (Hook.) Gottsche ☞ Diplolaena lyellii var. lamellata (Nees) Macvicar = Petalophyllum ralfsii (Wilson) Nees & Gottsche Diplolaena lyellii f. lamellata Nees = Petalophyllum ralfsii (Wilson) Nees & Gottsche Diplomitrium blyttii (Moerch ex Hornem.) Corda = Moerckia blyttii (Moerch ex Hornem.) Brockm. Diplomitrium hibernicum (Hook.) Corda = Moerckia hibernica (Hook.) Gottsche Diplophylleia albicans (L.) Trevis. = Diplophyllum albicans (L.) Dumort. Diplophylleia minuta (Schreb.) Trevis. = Anastrophyllum minutum (Schreb.) R. M. Schust. Diplophylleia obtusifolia (Hook.) Trevis. = Diplophyllum obtusifolium (Hook.) Dumort. Diplophylleia taxifolia (Wahlenb.) Trevis. = Diplophyllum taxifolium (Wahlenb.) Dumort. Diplophyllum apiculatum var. obtusatum R. M. Schust. = Diplophyllum obtusatum (R. M. Schust.) R. M. Schust. Diplophyllum dicksoni (Hook.) Dumort. = Douinia ovata (Dicks.) H. Buch Diplophyllum gymnostomophilum (Kaal.) Kaal. = Scapania gymnostomophila Kaal. Diplophyllum ovatum (Dicks.) Lindb. ex Steph. = Douinia ovata (Dicks.) H. Buch Diplophyllum scapanioides (C. Massal.) Steph. = Scapania scapanioides (C. Massal.) Grolle Dumortiera irrigua (Wilson ex Hook.) Nees = Dumortiera hirsuta (Sw.) Nees Dumortiera spathysii (Lindenb.) Nees = Athalamia spathysii (Lindenb.) S. Hatt. Duriaea helicophylla Bory & Mont. = Riella helicophylla (Bory & Mont.) Mont. Duriaea notarisii (Mont.) Bory & Mont. nom. illeg. = Riella notarisii (Mont.) Mont. Duvalia pilosa (Hornem.) Lindb. = Mannia pilosa (Hornem.) Frye & L. Clark Duvalia rupestris Nees = Mannia triandra (Scop.) Grolle Eucalyx crenulatus (Mitt.) Loeske = Jungermannia gracillima Sm. Eucalyx gracillimus (Sm.) Loeske = Jungermannia gracillima Sm. ☞ Eucalyx hyalinus (Lyell) Breidl. = Jungermannia hyalina Lyell Eucalyx muellerianus (Schiffn.) Müll. Frib. = Jungermannia hyalina Lyell Eucalyx obovatus (Nees) Breidl. = Jungermannia obovata Nees 394 R.M. Ros et al. Eucalyx subellipticus (Lindb. ex Kaal.) Breidl. = Jungermannia subelliptica (Lindb. ex Kaal.) Levier Eucephalozia bicuspidata (L.) Schiffn. = Cephalozia bicuspidata (L.) Dumort. Eulejeunea canariensis Steph. = Lejeunea canariensis (Steph.) Steph. Eulejeunea cavifolia (Ehrh.) Lindb. = Lejeunea cavifolia (Ehrh.) Lindb. Eulejeunea flava (Sw.) Steph. = Lejeunea flava (Sw.) Nees Eulejeunea lamacerina Steph. = Lejeunea lamacerina (Steph.) Schiffn. Eulejeunea pitardii Steph. = Lejeunea eckloniana Lindenb. Eulejeunea serpyllifolia (Dicks.) Schiffn. nom. illeg. = Lejeunea patens Lindb. [33] ? Eulejeunea serpyllifolia var. cavifolia (Ehrh.) Lindb. = Lejeunea cavifolia (Ehrh.) Lindb. ? Eulejeunea serpyllifolia var. patens (Lindb.) A. L. M. Guim. = Lejeunea patens Lindb. Eulejeunea ulicina (Taylor) Schiffn. = Microlejeunea ulicina (Taylor) A. Evans Evansia dentata (Raddi) Douin = Cephaloziella dentata (Raddi) Steph. Exormotheca africana Steph. = Exormotheca pustulosa Mitt. Exormotheca bullosa sensu Müll. Frib. = Exormotheca welwitschii Steph. Fegatella conica (L.) Corda = Conocephalum conicum (L.) Dumort. Fegatella officinalis Raddi nom. superfl. = Conocephalum conicum (L.) Dumort. Fimbriaria africana Mont. = Asterella africana (Mont.) A. Evans Fimbriaria blumeana Nees = Asterella blumeana (Nees) Kachroo REJECTED FROM THE AREA Fimbriaria bonjeanii De Not. = Asterella lindenbergiana (Corda ex Nees) Arnell Fimbriaria elegans auct. eur. non Spreng. = Asterella africana (Mont.) A. Evans Fimbriaria fragrans (Balbis) Nees = Mannia fragrans (Balbis) Frye & L. Clark [10] Fimbriaria fragrans auct. = Asterella saccata (Wahlenb.) A. Evans. [10] Fimbriaria lindenbergiana Corda ex Nees = Asterella lindenbergiana (Corda ex Nees) Arnell Fimbriaria ludwigii auct. non (Schwägr.) Limpr. = Asterella gracilis (F. Weber) Underw. Fimbriaria ludwigii (Schwägr.) Limpr. ex Müll. Frib. nom. inval. = Mannia triandra (Scop.) Grolle Fimbriaria nana Lindenb. = Athalamia hyalina (Sommerf.) S. Hatt. [9] Fimbriaria pilosa (Wahlenb.) Nees = Asterella gracilis (F. Weber) Underw. Fimbriaria raddii Corda ex Nees = Asterella wallichiana (Lehm. & Lindenb.) Pandé et al. REJECTED FROM THE AREA Fimbriaria saccata (Wahlenb.) Nees = Asterella saccata (Wahlenb.) A. Evans Fimbriaria stahlii Steph. = Asterella pringlei Underw. REJECTED FROM THE AREA Fimbriaria tenella auct. non L. = Asterella gracilis (F. Weber) Underw. Fossombronia angulosa var. caespitiformis Raddi = Fossombronia caespitiformis De Not. ex Rabenh. Fossombronia caespitiformis var. husnotii (Corb.) Corb. = Fossombronia caespitiformis subsp. multispira (Schiffn.) J.R. Bray and D.C. Cargill in Stotler et al. Fossombronia caespitiformis var. intermedia (Corb. ex Trab.) Trab. ex Bonner nom. inval. = Fossombronia echinata Macvicar [20] Fossombronia caespitiformis var. subcristata Schiffn. = Fossombronia caespitiformis De Not. ex Rabenh. subsp. caespitiformis Fossombronia corbulaeformis Trab. = Petalophyllum ralfsii (Wilson) Nees & Gottsche # Fossombronia crispula (Brot.) R. M. Schust. [21] Fossombronia cristata Lindb. = Fossombronia wondraczekii (Corda) Lindb. Fossombronia cristata var. wondraczekii (Corda) Lindb. = Fossombronia wondraczekii (Corda) Lindb. Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 395 Fossombronia crozalsii Corb. = Fossombronia mittenii Tind. Fossombronia dumortieri Huebener & Genth ex Lindb. nom. inval. = Fossombronia foveolata Lindb. Fossombronia husnotii Corb. = Fossombronia caespitiformis subsp. multispira (Schiffn.) J.R. Bray and D.C. Cargill in Stotler et al. Fossombronia intermedia Corb. ex Trab. nom. inval. = Fossombronia echinata Macvicar [20] Fossombronia intestinalis Taylor REJECTED FROM THE AREA [19] Fossombronia loitlesbergeri Schiffn. = Fossombronia pusilla (L.) Nees [23] Fossombronia mauritanica Trab. nom. inval. = Fossombronia caespitiformis subsp. multispira (Schiffn.) J. R. Bray and D. C. Cargill in Stotler et al. Fossombronia maritima (Paton) Paton = Fossombronia pusilla (L.) Nees Fossombronia pusilla var. corsa Lindb. ex Bonner nom. nud. = Fossombronia pusilla (L.) Nees Fossombronia pusilla var. decipiens Corb. = Fossombronia pusilla (L.) Nees Fossombronia pusilla var. maritima Paton = Fossombronia pusilla (L.) Nees l Fossombronia pusilla var. major Jur. nom. nud. = Fossombronia pusilla (L.) Nees Fossombronia verrucosa Lindb. = Fossombronia caespitiformis subsp. multispira (Schiffn.) J. R. Bray and D. C. Cargill in Stotler et al. Frullania aeolotis Mont. & Nees = Frullania riparia Hampe ex Lehm. Frullania bryhnii Müll. Frib. = Frullania polysticta Lindenb. Frullania calcarifera Steph. = Frullania tamarisci (L.) Dumort. Frullania caucasica Steph. = Frullania parvistipula Steph. Frullania cesatiana De Not. = Frullania riparia Hampe ex Lehm. Frullania cesatiana var. muscicola (Steph.) Bisang et al. nom. nud. REJECTED FROM THE AREA [24] Frullania cesatiana var. muscicola auct. eur. non (Steph.) Bisang et al. nom. nud. = Frullania azorica Sim-Sim et al. [24] Frullania cleistostoma Schiffn. & Wollny = Frullania inflata Gottsche Frullania davurica auct. eur. non Hampe = Frullania jackii Gottsche [26] Frullania davurica subsp. jackii (Gottsche) S. Hatt. = Frullania jackii Gottsche Frullania dilatata var. anomala Corb. = Frullania dilatata (L.) Dumort. Frullania dilatata var. briziana C. Massal. = Frullania dilatata (L.) Dumort. Frullania dilatata var. elongata Losada Lima & Vanden Berghen [25] Frullania dilatata var. microphylla (Wallroth) Nees = Frullania dilatata (L.) Dumort. Frullania eboracencis subsp. caucasica (Steph.) R. M. Schust. = Frullania parvistipula Steph. Frullania eboracencis subsp. parvistipula (Steph.) R. M. Schust. = Frullania parvistipula Steph. Frullania fragilifolia var. parvistipula (Steph.) Schiffn. = Frullania parvistipula Steph. Frullania funchalensis Gottsche nom. nud. = Frullania dilatata (L.) Dumort. Frullania germana (Taylor) Taylor = Frullania teneriffae (F. Weber) Nees Frullania hispanica Nees = Frullania tamarisci (L.) Dumort. Frullania hispanica Hampe non Nees = Frullania polysticta Lindenb. Frullania hutchinsiae (Hook.) Nees = Jubula hutchinsiae (Hook.) Dumort. Frullania illyrica Grolle = Frullania inflata Gottsche ? Frullania inflata subsp. illyrica (Grolle) R. M. Schust. = Frullania inflata Gottsche Frullania inflata var. illyrica (Grolle) R. M. Schust. = Frullania inflata Gottsche Frullania major Raddi nom. superfl. = Frullania tamarisci (L.) Dumort. 396 R.M. Ros et al. Frullania maritima Steph. p. p. = Frullania tamarisci (L.) Dumort. Frullania microphylla var. deciduifolia Grolle = Frullania microphylla (Gottsche) Pearson Frullania muscicola Steph. REJECTED FROM THE AREA [24] Frullania muscicola sensu Sérgio non Steph. = Frullania azorica Sim-Sim et al. [24] Frullania nervosa Mont. = Frullania tamarisci (L.) Dumort. Frullania nervosa var. microcellularis Müll. Frib. nom. nud. = Frullania tamarisci (L.) Dumort. Frullania nervosa var. polysticta Nees nom. herb. = Frullania polysticta Lindenb. Frullania obscurifolia Mitt. REJECTED FROM THE AREA Frullania saxicola Austin = Frullania inflata Gottsche Frullania squarrosa Dumort. = Frullania ericoides (Nees) Mont. Frullania tamarisci var. atrovirens Carrington = Frullania tamarisci (L.) Dumort. Frullania tamarisci var. calcarifera Steph. = Frullania tamarisci (L.) Dumort. Frullania tamarisci var. hispanica (Nees) Nees = Frullania tamarisci (L.) Dumort. Frullania tamarisci var. mediterranea (De Not.) Nees = Frullania tamarisci (L.) Dumort. Frullania tamarisci var. mediterranea f. robustior Schiffn. = Frullania tamarisci (L.) Dumort. Frullania tamarisci var. heterophylla Corb. = Frullania tamarisci (L.) Dumort. Frullania tamarisci var. heterophylla f. brunnea Schiffn. = Frullania tamarisci (L.) Dumort. Frullania tamarisci var. microphylla Gottsche = Frullania microphylla (Gottsche) Pearson Frullania tamarisci var. robusta Lindb. = Frullania tamarisci (L.) Dumort. Frullania tamarisci var. sardoa (De Not.) De Not. = Frullania tamarisci (L.) Dumort. Frullania tamarisci var. schiffneri Nicholl = Frullania tamarisci (L.) Dumort. Frullania tenera Lindb. ex Steph. non Spruce = Frullania parvistipula Steph. Frullania trabutiana Steph. = Frullania dilatata (L.) Dumort. Frullania welwitschii Steph. = Frullania ericoides (Nees) Mont. Frullania willkommii Steph. = Frullania tamarisci (L.) Dumort. Gongylanthus flagelliferus (Raddi) Nees = Gongylanthus ericetorum (Raddi) Nees Grimaldia androgyna (L.) Lindb. = Mannia androgyna (L.) A. Evans Grimaldia angustifolia (Neck.) Lindb. ex Schiffn. = Mannia androgyna (L.) A. Evans Grimaldia atlantica Trab. nom. inval. [39] Grimaldia barbifrons Bisch. = Mannia fragrans (Balbis) Frye & L. Clark Grimaldia carnica C. Massal. = Mannia pilosa (Hornem.) Frye & L. Clark Grimaldia dichotoma Raddi nom. illeg. = Mannia androgyna (L.) A. Evans Grimaldia fragrans (Balbis) Corda = Mannia fragrans (Balbis) Frye & L. Clark [10] Grimaldia hemisphaerica (L.) Lindenb. = Reboulia hemisphaerica (L.) Raddi Grimaldia pilosa (Hornem.) Lindb. = Mannia pilosa (Hornem.) Frye & L. Clark Grimaldia punicea Wallr. = Athalamia hyalina (Sommerf.) S. Hatt. Grimaldia rupestris (Nees) Lindenb. = Mannia triandra (Scop.) Grolle Grimaldia triandra (Scop.) Lindb. = Mannia triandra (Scop.) Grolle Grimaldia ventricosa Wallr. = Reboulia hemisphaerica (L.) Raddi Guentheria graveolens Trevis. = Corsinia coriandrina (Spreng.) Lindb. Gymnocolea acutiloba (Schiffn.) Müll. Frib. = Gymnocolea inflata (Huds.) Dumort. Gymnocolea huebeneriana (Nees) Dumort. = Hygrobiella laxifolia (Hook.) Spruce Gymnocolea inflata subsp. acutiloba (Schiffn.) R. M. Schust. & Damsh. ex L. Söderstr. & Vá≈a = Gymnocolea inflata (Huds.) Dumort. Gymnocolea inflata var. acutiloba (Schiffn.) S. W. Arnell = Gymnocolea inflata (Huds.) Dumort. Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 397 Gymnocolea inflata var. heterostipa (Carrington & Spruce) Müll. Frib. = Gymnocolea inflata (Huds.) Dumort. Gymnocolea laxifolia (Hook.) Dumort. = Hygrobiella laxifolia (Hook.) Spruce Gymnomitrion adustum Nees emend. Limpr. = Marsupella adusta (Nees emend. Limpr.) Spruce Gymnomitrion alpinum (Gottsche ex Husn.) Schiffn. = Marsupella alpina (Gottsche ex Husn.) Bernet Gymnomitrion cochleare (Lindb.) Müll. Frib. = Marsupella brevissima (Dumort.) Grolle Gymnomitrion commutatum (Limpr.) Schiffn. = Marsupella commutata (Limpr.) Bernet Gymnomitrion concinnatum var. intermedium Limpr. = Gymnomitrion concinnatum (Lightf.) Corda Gymnomitrion condensatum Ångstr. ex C. Hartm. = Marsupella condensata (Ångstr. ex C. Hartm.) Kaal. Gymnomitrion confertum (Limpr.) Limpr. = Marsupella brevissima (Dumort.) Grolle Gymnomitrion crassifolium Carrington = Marsupella brevissima (Dumort.) Grolle Gymnomitrion revolutum (Nees) Philib. = Apomarsupella revoluta (Nees) R. M. Schust. Gymnomitrion varians (Lindb.) Schiffn. = Marsupella brevissima (Dumort.) Grolle Haplozia amplexicaulis (Dumort.) Warnst. = Jungermannia sphaerocarpa Hook. Haplozia atrovirens (Dumort.) Müll. Frib. = Jungermannia atrovirens Dumort. Haplozia autumnalis (DC.) Warnst. = Jamesoniella autumnalis (DC.) Steph. Haplozia breidleri Müll. Frib. = Jungermannia confertissima Nees Haplozia cordifolia (Dumort.) Müll. Frib. = Jungermannia exsertifolia subsp. cordifolia (Dumort.) Vá≈a Haplozia crenulata (Mitt.) Müll. Frib. = Jungermannia gracillima Sm. Haplozia crenulata var. gracillima (Sm.) C. E. O. Jensen = Jungermannia gracillima Sm. Haplozia crenulata f. gracillima (Sm.) Müll. Frib. = Jungermannia gracillima Sm. Haplozia hyalina (Lyell) Warnst. = Jungermannia hyalina Lyell Haplozia levieri (Steph.) Müll. Frib. = Jungermannia confertissima Nees Haplozia lurida (Dumort.) Müll. Frib. = Jungermannia sphaerocarpa Hook. Haplozia muelleriana (Schiffn.) Casares-Gil = Jungermannia hyalina Lyell Haplozia nigrella (De Not.) Dumort. = Southbya nigrella (De Not.) Henriq. Haplozia oblongifolia Müll. Frib. = Jungermannia pumila With. Haplozia obovata (Nees) Casares-Gil = Jungermannia obovata Nees Haplozia polaris (Lindb.) Müll. Frib. = Jungermannia polaris Lindb. Haplozia pumila (With.) Müll. Frib. = Jungermannia pumila With. Haplozia riparia (Taylor) Müll. Frib. = Jungermannia atrovirens Dumort. Haplozia riparia var. rivularis (Bernet) Müll. Frib. = Jungermannia atrovirens Dumort. Haplozia rivularis (Schiffn.) Casares-Gil = Jungermannia pumila With. Haplozia schiffneri (Loitl.) Müll. Frib. = Jungermannia polaris Lindb. Haplozia schraderi (Mart.) Dumort. = Jamesoniella autumnalis (DC.) Steph. Haplozia sphaerocarpa (Hook.) Müll. Frib. = Jungermannia sphaerocarpa Hook. Haplozia sphaerocarpa var. amplexicaulis (Dumort.) Müll. Frib. = Jungermannia sphaerocarpa Hook. Haplozia sphaerocarpa var. nana (Nees ex Flot.) Müll. Frib. = Jungermannia sphaerocarpa Hook. Haplozia subelliptica (Lindb. ex Kaal.) Casares-Gil = Jungermannia subelliptica (Lindb. ex Kaal.) Levier Haplozia tristis (Nees) Müll. Frib.= Jungermannia atrovirens Dumort. 398 R.M. Ros et al. Harpalejeunea ovata (Hook.) Schiffn. non (Dicks.) Schiffn. = Harpalejeunea molleri (Steph.) Grolle Hepatica conica (L.) Lindb. = Conocephalum conicum (L.) Dumort. Herpetium deflexum (Mart.) Nees = Bazzania tricrenata (Wahlenb.) Lindb. Herpetium reptans (L.) Nees = Lepidozia reptans (L.) Dumort. Herpetium trilobatum (L.) Nees = Bazzania trilobata (L.) Gray Homalolejeunea mackaii (Hook.) Gray ex Casares-Gil = Marchesinia mackaii (Hook.) Gray Hygrobiella myriocarpa (Carrington) Spruce = Eremonotus myriocarpus (Carrington) Pearson Hygrophila irrigua (Wilson ex Hook.) Taylor nom. superfl. = Dumortiera hirsuta (Sw.) Nees Hypenantron africanum (Mont.) Kuntze = Asterella africana (Mont.) A. Evans ? Hypenantron lindenbergianum (Corda ex Nees) Kuntze = Asterella lindenbergiana (Corda ex Nees) Arnell Hypnenatron nanum (Lindenb.) Trevis. = Athalamia hyalina (Sommerf.) S. Hatt. Isopaches bicrenatus (Schmidel ex Hoffm.) H. Buch = Lophozia bicrenata (Schmidel ex Hoffm.) Dumort. Isopaches decolorans (Limpr.) H. Buch = Lophozia decolorans (Limpr.) Steph. Isopaches hellerianus (Nees ex Lindenb.) H. Buch = Anastrophyllum hellerianum (Nees ex Lindenb.) R. M. Schust. Jamesoniella autumnalis f. subapicalis (Nees) Müll. Frib. = Jamesoniella autumnalis (DC.) Steph. Jamesoniella subapicalis (Nees) Schiffn. = Jamesoniella autumnalis (DC.) Steph. Jubula hutchinsiae subsp. hutchinsiae var. integrifolia (Steph.) S. Hatt. [27] Jubula hutchinsiae var. sullivanti Spruce [27] Jubula hutchinsiae var. warburgii auct. eur. non Schiffn. = Jubula hutchinsiae (Hook.) Dumort. subsp. hutchinsiae Jubula javanica Steph. = Jubula hutchinsiae subsp. javanica (Steph.) Verd. Jubula pennsylvanica (Steph.) A. Evans [27] Jungermannia acuta Lindenb. nom. illeg. = Leiocolea collaris (Nees) Schljakov Jungermannia acuta var. aeruginosa Lindenb. = Leiocolea turbinata (Raddi) H. Buch Jungermannia acutiloba Kaal. non Hook. f. & Taylor = Gymnocolea inflata (Huds.) Dumort. Jungermannia aequiloba Schwägr. = Scapania aequiloba (Schwägr.) Dumort. Jungermannia affinis Wilson ex Hook. nom. inval. = Leiocolea turbinata (Raddi) H. Buch Jungermannia albescens Hook. = Pleurocladula albescens (Hook.) Grolle Jungermannia albicans L. = Diplophyllum albicans (L.) Dumort. # Jungermannia albicans var. rubescens Grognot = Diplophyllum albicans (L.) Dumort. Jungermannia albicans var. taxifolia (Wahlenb.) Wallr. = Diplophyllum taxifolium (Wahlenb.) Dumort. Jungermannia algeriensis Gottsche & Rabenh. = Leiocolea turbinata (Raddi) H. Buch Jungermannia alicularia De Not. = Southbya tophacea (Spruce) Spruce Jungermannia alpestris Schleich. ex F. Weber = Leiocolea collaris (Nees) Schljakov Jungermannia alpestris auct. non Schleich. ex F. Weber = Lophozia sudetica (Nees ex Huebener) Grolle Jungermannia amplexicaulis Dumort. = Jungermannia sphaerocarpa Hook. Jungermannia angulosa Dicks. = Fossombronia angulosa (Dicks.) Raddi Jungermannia anomala Hook. = Mylia anomala (Hook.) Gray Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 399 Jungermannia aquilegia Hook. f. & Taylor = Radula aquilegia (Hook. f. & Taylor) Gottsche et al. Jungermannia arboris-vitae With. = Porella arboris-vitae (With.) Grolle Jungermannia arenaria Nees = Lophozia excisa (Dicks.) Dumort. Jungermannia asplenioides L. emend. Taylor = Plagiochila asplenioides (L. emend. Taylor) Dumort. Jungermannia assimilis Mitt. = Anastrophyllum assimile (Mitt.) Steph. Jungermannia attenuata (Mart.) Lindenb. = Barbilophozia attenuata (Mart.) Loeske Jungermannia attenuata var. gracilis Schleich. ex Lindenb. = Barbilophozia attenuata (Mart.) Loeske # Jungermannia attenuata var. gradata Desm. = Barbilophozia attenuata (Mart.) Loeske Jungermannia autumnalis DC. = Jamesoniella autumnalis (DC.) Steph. Jungermannia badensis Gottsche = Leiocolea badensis (Gottsche) Schiffn. Jungermannia bantriensis Hook. = Leiocolea bantriensis (Hook.) Jörg. Jungermannia bantriensis var. acuta Lindb. = Leiocolea collaris (Nees) Schljakov Jungermannia bantriensis var. muelleri (Nees ex Lindenb.) Lindb. = Leiocolea collaris (Nees) Schljakov Jungermannia barbata Schmidel ex Schreb. = Barbilophozia barbata (Schmidel ex Schreb.) Loeske Jungermannia barbata var. attenuata (Mart.) Nees = Barbilophozia attenuata (Mart.) Loeske Jungermannia barbata var. floerkei (F. Weber & D. Mohr) Dumort. = Barbilophozia floerkei (F. Weber & D. Mohr) Loeske Jungermannia barbata var. lycopodioides (Wallr.) Nees = Barbilophozia lycopodioides (Wallr.) Loeske Jungermannia barbata var. quinquedentata (Huds.) Nees = Tritomaria quinquedentata (Huds.) H. Buch Jungermannia barbata var. schreberi Nees nom. illeg. = Barbilophozia barbata (Schmidel ex Schreb.) Loeske Jungermannia bartlingii Hampe ex Nees = Scapania cuspiduligera (Nees) Müll. Frib. Jungermannia baueri Mart. = Nowellia curvifolia (Dicks.) Mitt. Jungermannia baueriana (Schiffn.) Arnell = Barbilophozia hatcheri (A. Evans) Loeske Jungermannia bicalyculata Raddi = Cephalozia bicuspidata (L.) Dumort. Jungermannia bicornis F. Weber & D. Mohr = Anastrophyllum minutum (Schreb.) R. M. Schust. Jungermannia bicrenata Schmidel ex Hoffm. = Lophozia bicrenata (Schmidel ex Hoffm.) Dumort. Jungermannia bicuspidata L. = Cephalozia bicuspidata (L.) Dumort. Jungermannia bidentata L. = Lophocolea bidentata (L.) Dumort. Jungermannia bifaria Sw. = Plagiochila bifaria (Sw.) Lindenb. Jungermannia blasia DC. = Blasia pusilla L. Jungermannia blyttii Moerch ex Hornem. = Moerckia blyttii (Moerch ex Hornem.) Brockm. Jungermannia breidleri (Müll. Frib.) Gams = Jungermannia confertissima Nees Jungermannia byssacea A. Roth nom. rejic. = Cephaloziella hampeana (Nees) Schiffn. Jungermannia byssacea auct. non A. Roth = Cephaloziella divaricata (Sm.) Schiffn. Jungermannia calycina Taylor = Pellia endiviifolia (Dicks.) Dumort. Jungermannia calyculata Durieu & Mont. = Cephaloziella calyculata (Durieu & Mont.) Müll. Frib. Jungermannia calypogeia Raddi nom. illeg. = Calypogeia fissa (L.) Raddi Jungermannia catenulata Huebener = Cephalozia catenulata (Huebener) Lindb. 400 R.M. Ros et al. Jungermannia cavifolia Ehrh. = Lejeunea cavifolia (Ehrh.) Lindb. Jungermannia chamedryfolia With. = Riccardia chamedryfolia (With.) Grolle Jungermannia ciliaris L. = Ptilidium ciliare (L.) Hampe Jungermannia collaris Nees = Leiocolea collaris (Nees) Schljakov Jungermannia compacta A. Roth = Scapania compacta (A. Roth) Dumort. Jungermannia compacta Mazziari nom. nud. non Roth [28] Jungermannia commutata Huebener = Lophozia bicrenata (Schmidel ex Hoffm.) Dumort. Jungermannia complanata L. = Radula complanata (L.) Dumort. Jungermannia compressa Hook. = Nardia compressa (Hook.) Gray Jungermannia concinnata Lightf. = Gymnomitrion concinnatum (Lightf.) Corda Jungermannia confervoides Raddi = Cephaloziella divaricata (Sm.) Schiffn. Jungermannia connivens Dicks. = Cephalozia connivens (Dicks.) Lindb. Jungermannia corcyraea Nees = Leiocolea turbinata (Raddi) H. Buch Jungermannia cordaeana Huebener = Porella cordaeana (Huebener) Moore # Jungermannia cordifolia Brot. non Hook. [63] Jungermannia cordifolia Hook. non Brot. nec Ehrh. ex F. Weber nom. illeg. = Jungermannia exsertifolia subsp. cordifolia (Dumort.) Vá≈a Jungermannia crassifolia Lindenb. & Gottsche = Cephalozia crassifolia (Lindenb. & Gottsche) Fulford Jungermannia crenulata Sm. nom. illeg. non Schmidel = Jungermannia gracillima Sm. Jungermannia crenulata var. gracillima (Sm.) Hook. = Jungermannia gracillima Sm. # Jungermannia crispula Brot. [21] Jungermannia crocata De Not. = Lophocolea minor Nees Jungermannia cuneifolia Hook. = Leptoscyphus cuneifolius (Hook.) Mitt. Jungermannia cupressina Sw. = Lepidozia cupressina (Sw.) Lindenb. Jungermannia curta Mart. = Scapania curta (Mart.) Dumort. Jungermannia curvifolia Dicks. = Nowellia curvifolia (Dicks.) Mitt. Jungermannia cuspiduligera Nees = Scapania cuspiduligera (Nees) Müll. Frib. Jungermannia cylindracea Dumort. = Lophozia excisa (Dicks.) Dumort. Jungermannia decipiens Hook. = Adelanthus decipiens (Hook.) Mitt. Jungermannia decolorans Limpr. = Lophozia decolorans (Limpr.) Steph. Jungermannia deflexa var. flaccida Schleich. ex Lindenb. nom. illeg. = Bazzania flaccida (Dumort.) Grolle Jungermannia dentata Raddi = Cephaloziella dentata (Raddi) Steph. Jungermannia denudata (Mart.) Nees = Odontoschisma denudatum (Mart.) Dumort. Jungermannia dilatata L. = Frullania dilatata (L.) Dumort. Jungermannia distans Schwein. = Porella pinnata L. Jungermannia divaricata Sm. = Cephaloziella divaricata (Sm.) Schiffn. Jungermannia emarginata Ehrh. = Marsupella emarginata (Ehrh.) Dumort. Jungermannia endiviifolia Dicks. = Pellia endiviifolia (Dicks.) Dumort. Jungermannia epiphylla L. = Pellia epiphylla (L.) Corda Jungermannia epiphylla var. furcigera Hook. = Pellia endiviifolia (Dicks.) Dumort. Jungermannia epiphylla var. longifolia Lam. & DC. = Pellia endiviifolia (Dicks.) Dumort. Jungermannia ericoides Nees = Frullania ericoides (Nees) Mont. Jungermannia excisa Dicks. = Lophozia excisa (Dicks.) Dumort. Jungermannia exigua Taylor = Plagiochila exigua (Taylor) Taylor Jungermannia exsecta Schmidel ex Schrad. = Tritomaria exsecta (Schmidel ex Schrad.) Loeske Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 401 Jungermannia exsectiformis Breidl. = Tritomaria exsectiformis (Breidl.) Loeske Jungermannia falcata Raddi nom. illeg. = Diplophyllum albicans (L.) Dumort. Jungermannia fennica Gottsche = Arnellia fennica (Gottsche) Lindb. Jungermannia fissa (L.) Scop. = Calypogeia fissa (L.) Raddi Jungermannia fissidentoidea Huebener = Diplophyllum albicans (L.) Dumort. Jungermannia flaccida Huebener = Jungermannia obovata Nees Jungermannia flava Sw. = Lejeunea flava (Sw.) Nees Jungermannia floerkei F. Weber & D. Mohr = Barbilophozia floerkei (F. Weber & D. Mohr) Loeske Jungermannia flotoviana Nees = Harpanthus flotovianus (Nees) Nees Jungermannia fluitans Nees = Cladopodiella fluitans (Nees) H. Buch Jungermannia foscardiana Mazziari nom. nud. [28] Jungermannia fragilifolia Taylor = Frullania fragilifolia (Taylor) Gottsche et al. Jungermannia fragilis A. Roth = Chiloscyphus pallescens (Ehrh. ex Hoffm.) Dumort. Jungermannia fragrans Moris & De Not. = Lophocolea fragrans (Moris & De Not.) Gottsche et al. Jungermannia francisci Hook. = Cladopodiella francisci (Hook.) Jörg. Jungermannia fruticulosa auct. eur. non (O. F. Müll.) Sm. = Metzgeria violacea (Ach.) Dumort. [40] Jungermannia fuciformis Bory [43] Jungermannia funckii F. Weber & D. Mohr = Marsupella funckii (F. Weber & D. Mohr) Dumort. Jungermannia furcata L. = Metzgeria furcata (L.) Dumort. Jungermannia germana Taylor = Frullania teneriffae (F. Weber) Nees Jungermannia gillmanii Austin = Lophozia gillmanii (Austin) R. M. Schust. Jungermannia gracilis (Schleich. ex Lindenb.) Lindb. = Barbilophozia attenuata (Mart.) Loeske Jungermannia grandiretis Lindb. ex Kaal. = Lophozia grandiretis (Lindb. ex Kaal.) Schiffn. Jungermannia graveolens Schrad. = Geocalyx graveolens (Schrad.) Nees Jungermannia grimsulana J. B. Jack ex Gottsche & Rabenh. = Cephaloziella grimsulana (J.B. Jack ex Gottsche & Rabenh.) Lacout. Jungermannia guilfortiana Mazziari nom. nud. [28] Jungermannia guttulata Lindb. & Arnell = Lophozia longiflora (Nees) Schiffn. Jungermannia hamatifolia Hook. = Drepanolejeunea hamatifolia (Hook.) Schiffn. Jungermannia hampeana Nees = Cephaloziella hampeana (Nees) Schiffn. Jungermannia hatcheri A. Evans = Barbilophozia hatcheri (A. Evans) Loeske Jungermannia helleriana Nees ex Lindenb. = Anastrophyllum hellerianum (Nees ex Lindenb.) R. M. Schust. Jungermannia heterocolpos Thed. ex Hartm. = Leiocolea heterocolpos (Thed. ex Hartm.) H. Buch Jungermannia heterophylla Schrad. = Lophocolea heterophylla (Schrad.) Dumort. Jungermannia hibernica Hook. = Moerckia hibernica (Hook.) Gottsche Jungermannia hoffmannii Wallr. = Ptilidium ciliare (L.) Hampe Jungermannia hornschuchiana Nees = Leiocolea bantriensis (Hook.) Jörg. Jungermannia hutchinsiae Hook. = Jubula hutchinsiae (Hook.) Dumort. subsp. hutchinsiae Jungermannia incisa Schrad. = Lophozia incisa (Schrad.) Dumort. Jungermannia inconspicua Raddi = Cololejeunea minutissima (Sm.) Schiffn. Jungermannia inflata Huds. = Gymnocolea inflata (Huds.) Dumort. 402 R.M. Ros et al. Jungermannia inflata var. fluitans (Nees) Gottsche et al. = Cladopodiella fluitans (Nees) H. Buch Jungermannia integrifolia Lehm. & Lindenb. = Chiloscyphus integrifolius (Lehm. & Lindenb.) Lehm. & Lindenb. REJECTED FROM THE AREA Jungermannia jenseniana Grolle REJECTED FROM THE AREA [29] Jungermannia intermedia Lindenb. nom. illeg. = Lophozia excisa (Dicks.) Dumort. Jungermannia interrupta Nees = Pedinophyllum interruptum (Nees) Kaal. Jungermannia irrigua Nees = Scapania irrigua (Nees) Nees Jungermannia julacea L. = Anthelia julacea (L.) Dumort. Jungermannia julacea var. clavuligera Nees = Anthelia juratzkana (Limpr.) Trevis. Jungermannia juratzkana Limpr. = Anthelia juratzkana (Limpr.) Trevis. Jungermannia karl-muelleri Grolle = Jungermannia pumila With. Jungermannia kaurinii Limpr. = Leiocolea gillmanii (Austin) R. M. Schust. Jungermannia kunzeana Huebener = Barbilophozia kunzeana (Huebener) Müll. Frib. Jungermannia lacera Mazziari nom. nud. [28] Jungermannia laevigata Schrad. = Porella arboris-vitae (With.) Grolle Jungermannia lammersiana Huebener = Cephalozia bicuspidata (L.) Dumort. Jungermannia lanceolata L. nom. rejic. = Jungermannia atrovirens Dumort. Jungermannia lanceolata auct. non L. = Jungermannia leiantha Grolle Jungermannia laurentiana De Not. = Leiocolea collaris (Nees) Schljakov Jungermannia laxifolia Hook. = Hygrobiella laxifolia (Hook.) Spruce Jungermannia leersii A. Roth = Ptilidium ciliare (L.) Hampe Jungermannia laevigata Schrad. = Porella arboris-vitae (With.) Grolle Jungermannia levieri (Steph.) H. Buch, A. Evans & Verd. = Jungermannia confertissima Nees Jungermannia limosa Mazziari nom. nud. [28] Jungermannia limprichtii (Lindb. ex C. Massal. & Carestia) Warnst. = Lophozia excisa (Dicks.) Dumort. Jungermannia longidens Lindb. = Lophozia longidens (Lindb.) Macoun Jungermannia longiflora Nees = Lophozia longiflora (Nees) Schiffn. Jungermannia lunulifolia Dumort. = Cephalozia lunulifolia (Dumort.) Dumort. Jungermannia lurida Dumort. nom. illeg. = Jungermannia sphaerocarpa Hook. Jungermannia lycopodioides Wallr. = Barbilophozia lycopodioides (Wallr.) Loeske Jungermannia lycopodioides var. floerkei (F. Weber & D. Mohr) Lindb. = Barbilophozia floerkei (F. Weber & D. Mohr) Loeske Jungermannia lyellii Hook. = Pallavicinia lyellii (Hook.) Carruth. Jungermannia lyonii Taylor = Tritomaria quinquedentata (Huds.) H. Buch Jungermannia mackaii Hook. = Marchesinia mackaii (Hook.) Gray Jungermannia macounii Austin = Cephalozia macounii (Austin) Austin Jungermannia menzelii Corda = Cephalozia bicuspidata (L.) Dumort. Jungermannia michauxii F. Weber = Anastrophyllum michauxii (F. Weber) H. Buch Jungermannia minuta Schreb. = Anastrophyllum minutum (Schreb.) R. M. Schust. Jungermannia minuta var. procera Nees = Anastrophyllum michauxii (F. Weber) H. Buch Jungermannia minuta var. robusta C. Massal. & Carestia nom. inval. = Anastrophyllum assimile (Mitt.) Steph. Jungermannia minutissima Sm. = Cololejeunea minutissima (Sm.) Schiffn. Jungermannia muelleri Nees ex Lindenb. = Leiocolea collaris (Nees) Schljakov Jungermannia multifida L. = Riccardia multifida (L.) Gray Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 403 Jungermannia multifida var. sinuata Dicks. ex Hook. = Riccardia chamedryfolia (With.) Grolle Jungermannia myriocarpa Carrington = Eremonotus myriocarpus (Carrington) Pearson Jungermannia nana Nees ex Flot. = Jungermannia sphaerocarpa Hook. Jungermannia nardioides Lindb. = Anastrophyllum assimile (Mitt.) Steph. Jungermannia naumannii Nees ex Mart. = Barbilophozia floerkei (F. Weber & D. Mohr) Loeske Jungermannia nemorea L. = Scapania nemorea (L.) Grolle Jungermannia nemorosa L. nom. illeg. = Scapania nemorea (L.) Grolle Jungermannia nemorosa var. recurvifolia Hook. = Scapania gracilis Lindb. Jungermannia nevicensis Carrington = Marsupella boeckii (Austin) Lindb. Jungermannia nigrella De Not. = Southbya nigrella (De Not.) Henriq. Jungermannia nigricans Lam. nom. illeg. = Frullania tamarisci (L.) Dumort. Jungermannia obtusa Lindb. = Lophozia obtusa (Lindb.) A. Evans Jungermannia obtusifolia Hook. = Diplophyllum obtusifolium (Hook.) Dumort. # Jungermannia oppositifolia Lacroix = Gongylanthus ericetorum (Raddi) Nees Jungermannia orcadensis Hook. = Anastrepta orcadensis (Hook.) Schiffn. Jungermannia pallescens Ehrh. ex Hoffm. = Chiloscyphus pallescens (Ehrh. ex Hoffm.) Dumort. Jungermannia pallescens var. rivularis Schrad. = Chiloscyphus polyanthos (L.) Corda Jungermannia palmata Hedw. = Riccardia palmata (Hedw.) Carruth. Jungermannia pauciflora Dicks. = Kurzia pauciflora (Dicks.) Grolle Jungermannia pinguis L. = Aneura pinguis (L.) Dumort. Jungermannia platyphylla L. = Porella platyphylla (L.) Pfeiff. Jungermannia platyphylla var. thuja Hook. = Porella arboris-vitae (With.) Grolle Jungermannia platyphylla var. canariensis F. Weber = Porella canariensis (F. Weber) Underw. Jungermannia platyphylloidea Schwein. = Porella platyphylla (L.) Pfeiff. Jungermannia pleniceps Austin = Cephalozia pleniceps (Austin) Lindb. Jungermannia plicata Hartm. = Barbilophozia kunzeana (Huebener) Müll. Frib. Jungermannia polita Nees = Tritomaria polita (Nees) Schiffn. Jungermannia polyanthos L. = Chiloscyphus polyanthos (L.) Corda Jungermannia porella Dicks. nom. illeg. = Porella pinnata L. Jungermannia porelloides Torrey ex Nees = Plagiochila porelloides (Torrey ex Nees) Lindenb. Jungermannia porphyroleuca Nees. nom. illeg. = Lophozia longiflora (Nees) Schiffn. Jungermannia porphyroleuca var. guttulata (Lindb. & Arnell) Warnst. = Lophozia longiflora (Nees) Schiffn. Jungermannia prostrata Sw. = Odontoschisma prostratum (Sw.) Trevis. Jungermannia pubescens Schrank = Apometzgeria pubescens (Schrank) Kuwah. Jungermannia pulcherrima Weber = Ptilidium pulcherrimum (Weber) Vain. Jungermannia pulvinata Raddi = Marsupella emarginata (Ehrh.) Dumort. Jungermannia pumila subsp. polaris (Lindb.) R. M. Schust. = Jungermannia polaris Lindb. Jungermannia pumila var. rivularis (Schiffn.) Frye & L. Clark = Jungermannia pumila With. Jungermannia punctata Taylor = Plagiochila punctata (Taylor) Taylor Jungermannia pusilla L. = Fossombronia pusilla (L.) Nees Jungermannia quinquedentata Huds. = Tritomaria quinquedentata (Huds.) H. Buch 404 R.M. Ros et al. Jungermannia quinquedentata var. attenuata (Mart.) Steud. = Barbilophozia attenuata (Mart.) Loeske Jungermannia raddiana C. Massal. = Cephaloziella rubella (Nees) Warnst. Jungermannia ralfsii Wilson = Petalophyllum ralfsii (Wilson) Nees & Gottsche Jungermannia reclusa Taylor = Cephalozia catenulata (Huebener) Lindb. Jungermannia reichardtii Gottsche ex Jur. = Anastrophyllum assimile (Mitt.) Steph. Jungermannia reptans L. = Lepidozia reptans (L.) Dumort. Jungermannia resupinata Hook. nom. inval. = Scapania compacta (A. Roth) Dumort. Jungermannia resupinata L. = Scapania undulata (L.) Dumort. Jungermannia rigida Wallr. nom. illeg. = Aneura pinguis (L.) Dumort. Jungermannia rigidula Huebener nom. illeg. = Cephalozia bicuspidata (L.) Dumort. Jungermannia riparia Taylor = Jungermannia atrovirens Dumort. $ Jungermannia riparia var. rotundifolia = Jungermannia atrovirens Dumort. Jungermannia rosacea Corda = Scapania curta (Mart.) Dumort. Jungermannia rostellata Huebener = Jungermannia pumila With. Jungermannia rubella Nees = Cephaloziella rubella (Nees) Warnst. Jungermannia saxicola Schrad. = Anastrophyllum saxicola (Schrad.) R. M. Schust. Jungermannia scalariformis Nees = Jungermannia confertissima Nees Jungermannia scalaris Schmidel ex Schreb. nom. dub. [12] Jungermannia scalaris Hook. homon. illeg. = Nardia scalaris Gray Jungermannia scalaris var. denudata Mart. = Odontoschisma denudatum (Mart.) Dumort. Jungermannia scalaris var. minor Nees = Nardia geoscyphus (De Not.) Lindb. Jungermannia scapanioides C. Massal. = Scapania scapanioides (C. Massal.) Grolle Jungermannia schiffneri (Loitl.) A. Evans = Jungermannia polaris Lindb. Jungermannia schraderi Mart. = Jamesoniella autumnalis (DC.) Steph. Jungermannia schraderi auct. non Mart. = Jamesoniella undulifolia (Nees) Müll. Frib. Jungermannia schreberi S. Hatt. nom. inval. = Barbilophozia barbata (Schmidel) Loeske Jungermannia scitula Taylor = Tritomaria scitula (Taylor) Jörg. Jungermannia scutata F. Weber & D. Mohr = Harpanthus scutatus (F. Weber & D. Mohr) Spruce Jungermannia serpyllifolia var. ovata auct. non Hook. = Harpalejeunea molleri (Steph.) Grolle Jungermannia setiformis Ehrh. = Tetralophozia setiformis (Ehrh.) Schljakov Jungermannia sinuata Dicks. nom. illeg. = Riccardia chamedryfolia (With.) Grolle Jungermannia socia Nees = Lophozia excisa (Dicks.) Dumort. Jungermannia sphacelata Gieseke ex Lindenb. = Marsupella sphacelata (Gieseke ex Lindenb.) Dumort. Jungermannia sphaerocarpa var. amplexicaulis (Dumort.) Frye & L. Clark = Jungermannia sphaerocarpa Hook. Jungermannia sphaerocarpoidea De Not. = Jungermannia atrovirens Dumort. Jungermannia sphagni Dicks. = Odontoschisma sphagni (Dicks.) Dumort. Jungermannia spinulosa Dicks. = Plagiochila spinulosa (Dicks.) Dumort. Jungermannia sprengelii Mart. = Calypogeia fissa (L.) Raddi Jungermannia starkei auct. non Funk ex Nees nom. illeg. = Cephaloziella divaricata (Sm.) Schiffn. Jungermannia stellulifera Taylor ex Carrington & Pearson = Cephaloziella divaricata (Sm.) Schiffn. Jungermannia stillicidiorum (Raddi) De Not. = Southbya tophacea (Spruce) Spruce Jungermannia stipulacea Hook. = Harpanthus scutatus (F. Weber & D. Mohr) Spruce Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 405 Jungermannia subalpina Nees ex Lindenb. = Scapania subalpina (Nees ex Lindenb.) Dumort Jungermannia subapicalis Nees = Jamesoniella autumnalis (DC.) Steph. Jungermannia sudetica Nees ex Huebener = Lophozia sudetica (Nees ex Huebener) Grolle Jungermannia sullivantii Austin = Cephaloziella rubella (Nees) Warnst. Jungermannia tamarisci L. = Frullania tamarisci (L.) Dumort. Jungermannia tamarisci var. mediterranea De Not. = Frullania tamarisci (L.) Dumort. Jungermannia tamarisci var. sardoa De Not. = Frullania tamarisci (L.) Dumort. Jungermannia tamariscifolia L. nom. illeg. = Frullania tamarisci (L.) Dumort. Jungermannia taxifolia Wahlenb. = Diplophyllum taxifolium (Wahlenb.) Dumort. Jungermannia taylorii Hook. = Mylia taylorii (Hook.) Gray Jungermannia taylorii var. anomala (Hook.) Nees = Mylia anomala (Hook.) Gray Jungermannia teneriffae F. Weber = Frullania teneriffae (F. Weber) Nees Jungermannia tenerrima Raddi nom. illeg. = Jungermannia gracillima Sm. Jungermannia tersa Nees = Jungermannia sphaerocarpa Hook. Jungermannia thuja Dicks. nom. illeg. = Porella arboris-vitae (With.) Grolle Jungermannia thuja auct. non Dicks. nom. illeg. = Porella obtusata (Taylor) Trevis. Jungermannia tomentella Ehrh. = Trichocolea tomentella (Ehrh.) Dumort. Jungermannia tophacea Spruce = Southbya tophacea (Spruce) Spruce Jungermannia trichomanis (L.) Scop. = Calypogeia fissa (L.) Raddi Jungermannia trichomanis auct. non (L.) Scop. = Calypogeia azurea Stotler & Crotz [11] Jungermannia trichophylla L. = Blepharostoma trichophyllum (L.) Dumort. Jungermannia tricrenata Wahlenb. = Bazzania tricrenata (Wahlenb.) Lindb. Jungermannia trilobata L. = Bazzania trilobata (L.) Gray Jungermannia tristis Nees = Jungermannia atrovirens Dumort. Jungermannia turbinata Raddi = Leiocolea turbinata (Raddi) H. Buch Jungermannia turneri Hook. = Cephaloziella turneri (Hook.) Müll. Frib. Jungermannia ulicina Taylor = Microlejeunea ulicina (Taylor) A. Evans Jungermannia uliginosa Sw. ex Lindenb. = Scapania uliginosa (Sw. ex Lindenb.) Dumort. Jungermannia umbrosa Schrad. = Scapania umbrosa (Schrad.) Dumort. Jungermannia undulata L. = Scapania undulata (L.) Dumort. Jungermannia varians Gottsche = Cephaloziella varians (Gottsche) Steph. Jungermannia ventricosa Dicks. = Lophozia ventricosa (Dicks.) Dumort. Jungermannia ventricosa var. guttulata (Lindb. & Arnell) C. Massal. = Lophozia longiflora (Nees) Schiffn. Jungermannia ventricosa var. longidens (Lindb.) C. Massal. & Carestia = Lophozia longidens (Lindb.) Macoun Jungermannia ventricosa var. porphyroleuca Hartm. = Lophozia longiflora (Nees) Schiffn. Jungermannia viticulosa L. = Saccogyna viticulosa (L.) Dumort. Jungermannia wenzelii Nees = Lophozia wenzelii (Nees) Steph. Jungermannia wilsonii (Nees) Hook. f. & Taylor = Acrobolbus wilsonii Nees Jungermannia wilsoniana Nees = Leiocolea turbinata (Raddi) H. Buch Jungermannia wondraczekii Corda = Fossombronia wondraczekii (Corda) Lindb. Jungermannia zeyheri Huebener = Jungermannia pumila With. Kantius(Kantia) argutus(arguta) (Nees & Mont.) Lindb. = Calypogeia arguta Nees & Mont. Kantius(Kantia) calypogeus(calypogea) (Raddi) Lindb. = Calypogeia fissa (L.) Raddi Kantius(Kantia) fissus(fissa) (L.) Lindb. = Calypogeia fissa (L.) Raddi 406 R.M. Ros et al. Kantius(Kantia) muellerianus(muelleriana) Schiffn. = Calypogeia muelleriana (Schiffn.) Müll. Frib. Kantius (Kantia) neesianus (neesiana) (C. Massal. & Carestia) Arnell & C. E. O. Jensen = Calypogeia neesiana (C. Massal. & Carestia) Müll. Frib. Kantius(Kantia) sphagnicolus(sphagnicola) Arnell & J. Perss. = Calypogeia sphagnicola (Arnell & J. Perss.) Warnst. & Loeske Kantius(Kantia) sprengelii (Mart.) Trevis. = Calypogeia fissa (L.) Raddi Kantius(Kantia) submersus(submersa) Arnell = Calypogeia sphagnicola (Arnell & J. Perss.) Warnst. & Loeske Kantius(Kantia) suecicus(suecica) Arnell & J. Perss. = Calypogeia suecica (Arnell. & J. Perss.) Müll. Frib. Kantius(Kantia) trichomanis (L.) Gray = Calypogeia fissa (L.) Raddi Kantius(Kantia) trichomanis auct. non (L.) Gray = Calypogeia azurea Stotler & Crotz [11] Kantius(Kantia) trichomanis var. fissus(fissa) (L.) Lindb. = Calypogeia fissa (L.) Raddi Kantius(Kantia) trichomanis var. neesianus(neesiana) C. Massal. & Carestia = Calypogeia neesiana (C. Massal. & Carestia) Müll. Frib. Leiocolea alpestris (Schleich. ex F. Weber) Isov. = Leiocolea collaris (Nees) Schljakov Leiocolea kaurinii (Limpr.) Jörg. = Leiocolea gillmanii (Austin) A. Evans Leiocolea muelleri (Nees ex Lindenb.) Jörg. = Leiocolea collaris (Nees) Schljakov Leiocolea obtusa (Lindb.) H. Buch = Lophozia obtusa (Lindb.) A. Evans Leiocolea turbinata f. aquatica Jelenc nom. nud. [32] Leioscyphus anomalus (Hook.) Steph. = Mylia anomala (Hook.) Gray Leioscyphus cuneifolius (Hook.) Steph. = Leptoscyphus cuneifolius (Hook.) Mitt. Lejeunea aberrans Lindenb. & Gottsche = Acanthocoleus aberrans (Lindenb. & Gottsche) Kruijt Lejeunea azorica Steph. = Lejeunea lamacerina (Steph.) Schiffn. Lejeunea calcarea Lib. = Cololejeunea calcarea (Lib.) Schiffn. Lejeunea calcarea var. rossettiana (C. Massal.) Mansion & Clerbois = Cololejeunea rossettiana (C. Massal.) Schiffn. Lejeunea cavifolia var. loitlesbergeri Schiffn. nom. inval. = Lejeunea cavifolia (Ehrh.) Lindb. Lejeunea diversiloba Spruce = Lejeunea hibernica Bischl. et al. ex Grolle Lejeunea echinata (Hook.) Taylor ex Gottsche et al. nom. inval. = Cololejeunea calcarea (Lib.) Schiffn. Lejeunea flava subsp. moorei (Lindb.) R. M. Schust. [34] Lejeunea hamatifolia (Hook.) Spreng. = Drepanolejeunea hamatifolia (Hook.) Schiffn. Lejeunea hamatifolia Dumort. nom. inval. = Drepanolejeunea hamatifolia (Hook.) Schiffn. Lejeunea holtii Spruce = Lejeunea eckloniana Lindenb. Lejeunea inconspicua (Raddi) De Not. ex Rabenh. = Cololejeunea minutissima (Sm.) Schiffn. Lejeunea lamacerina var. azorica (Steph.) Greig-Smith = Lejeunea lamacerina (Steph.) Schiffn. Lejeunea mackaii (Hook.) Spreng. = Marchesinia mackaii (Hook.) Gray Lejeunea macvicarii Pearson = Lejeunea mandonii (Steph.) Müll. Frib. Lejeunea microscopica (Taylor) Gottsche et al. = Aphanolejeunea microscopica (Taylor) A. Evans Lejeunea minutissima (Sm.) Spreng. = Cololejeunea minutissima (Sm.) Schiffn. Lejeunea molleri Steph. = Harpalejeunea molleri (Steph.) Grolle Lejeunea moorei Lindb. [34] Lejeunea ovata (Dicks.) Taylor ex Gottsche = Douinia ovata (Dicks.) H. Buch Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 407 Lejeunea planiuscula (Lindb.) H. Buch = Lejeunea lamacerina (Steph.) Schiffn. Lejeunea rossettiana C. Massal. = Cololejeunea rossettiana (C. Massal.) Schiffn. Lejeunea serpyllifolia (Dicks.) Lib. nom. illeg. = Lejeunea patens Lindb. [33] Lejeunea serpyllifolia Scop. = Lejeunea cavifolia (Ehrh.) Lindb. [33] Lejeunea serpyllifolia var. cavifolia (Ehrh.) Lindb. nom. inval. = Lejeunea cavifolia (Ehrh.) Lindb. ☞ Lejeunea serpyllifolia var. loitlesbergeri Schiffn. = Lejeunea cavifolia (Ehrh.) Lindb. Lejeunea serpyllifolia var. planiuscula C. Massal. nom. illeg. = Lejeunea lamacerina (Steph.) Schiffn. Lejeunea serpyllifolia f. planiuscula Lindb. nom. illeg. = Lejeunea lamacerina (Steph.) Schiffn. Lejeunea taylorii Spruce = Cololejeunea minutissima (Sm.) Schiffn. Lejeunea ulicina (Taylor) Gottsche et al. = Microlejeunea ulicina (Taylor) Steph. Lepidozia azorica H. Buch & Perss. = Telaranea azorica (H. Buch & Perss.) Pócs ex Schumacker & Vá≈a [65] Lepidozia pearsonii Spruce REJECTED FROM THE AREA [35] Lepidozia pinnata (Hook.) Dumort. = Lepidozia cupressina (Sw.) Lindenb. Lepidozia reptans var. tenera (Huebener) Nees = Lepidozia reptans (L.) Dumort. Lepidozia setacea (Weber) Mitt. = Kurzia pauciflora (Dicks.) Grolle Lepidozia sylvatica A. Evans = Kurzia sylvatica (A. Evans) Grolle Lepidozia trichoclados Müll. Frib. = Kurzia trichoclados (Müll. Frib.) Grolle Leptoscyphus anomalus (Hook.) Lindb. = Mylia anomala (Hook.) Gray Leptoscyphus interruptus (Nees) Lindb. = Pedinophyllum interruptum (Nees) Kaal. Leptoscyphus taylorii (Hook.) Mitt. = Mylia taylorii (Hook.) Gray Liochlaena lanceolata Nees = Jungermannia leiantha Grolle Liochlaena subulata (A. Evans) Schljakov = Jungermannia subulata A. Evans Lophocolea alata Mitt. ex Larter = Lophocolea bidentata (L.) Dumort. ☞ Lophocolea bidentata var. alata (Mitt. ex Larter) Nees = Lophocolea bidentata (L.) Dumort. Lophocolea bidentata var. cuspidata (Nees) Nees = Lophocolea bidentata (L.) Dumort. Lophocolea bidentata var. fluitans F. A. Camus ex Husn. = Lophocolea bidentata (L.) Dumort. Lophocolea bidentata var. latifolia (Nees) Rabenh. nom. inval. = Lophocolea bidentata (L.) Dumort. Lophocolea bidentata var. rivularis (Raddi) Warnst. = Lophocolea bidentata (L.) Dumort. Lophocolea crocata (De Not.) Nees = Lophocolea minor Nees Lophocolea cuspidata (Nees) Limpr. = Lophocolea bidentata (L.) Dumort. l Lophocolea heterophylla f. gemmifera Andres = Lophocolea heterophylla (Schrad.) Dumort. Lophocolea heterophylla var. minor (Nees) Douin = Lophocolea minor Nees Lophocolea heterophylla f. propagulifera Feld nom. inval. = Lophocolea heterophylla (Schrad.) Dumort. Lophocolea hirticalyx Corb. & Steph. = Lophocolea fragrans (Moris & De Not.) Gottsche et al. Lophocolea hookeriana Nees = Lophocolea bidentata (L.) Dumort. Lophocolea incisa Lindb. = Lophocolea heterophylla (Schrad.) Dumort. Lophocolea lateralis Dumort. = Lophocolea bidentata (L.) Dumort. Lophocolea latifolia Nees = Lophocolea bidentata (L.) Dumort. Lophocolea preauxiana Mont. = Saccogyna viticulosa (L.) Dumort. 408 R.M. Ros et al. ☞ Lophocolea spicata Gottsche et al. = Lophocolea fragrans (Moris & De Not.) Gottsche et al. Lophocolea vogesiaca Huebener ex Nees = Harpanthus flotovianus (Nees) Nees Lophozia acutiloba Schiffn. = Gymnocolea inflata (Huds.) Dumort. Lophozia alpestris (Schleich. ex F. Weber) A. Evans = Leiocolea collaris (Nees) Schljakov Lophozia alpestris auct. non (Schleich. ex F. Weber) A. Evans = Lophozia sudetica (Nees ex Huebener) Grolle Lophozia attenuata (Mart.) Dumort. = Barbilophozia attenuata (Mart.) Loeske Lophozia atlantica (Kaal.) Schiffn. = Barbilophozia atlantica (Kaal.) Müll. Frib. Lophozia badensis (Gottsche) Schiffn. = Leiocolea badensis (Gottsche) Jörg. Lophozia bantriensis (Hook.) Steph. = Leiocolea bantriensis (Hook.) Jörg. Lophozia barbata (Schmidel ex Schreb.) Dumort. = Barbilophozia barbata (Schmidel ex Schreb.) Loeske Lophozia baueriana (Schiffn.) Schiffn. = Barbilophozia hatcheri (A. Evans) Loeske Lophozia canariensis Bryhn = Marsupella emarginata (Ehrh.) Dumort. Lophozia capitata subsp. laxa (Lindb.) Bisang = Lophozia laxa (Lindb.) Grolle Lophozia collaris (Nees) Dumort. = Leiocolea collaris (Nees) Schljakov Lophozia confertifolia Schiffn. = Lophozia ventricosa (Dicks.) Dumort. [38] Lophozia ehrhartiana (F. Weber) Inoue & Steere = Lophozia ventricosa (Dicks.) Dumort. Lophozia elongata Steph. REJECTED FROM THE AREA Lophozia excisa var. cylindracea (Dumort.) Müll. Frib. = Lophozia excisa (Dicks.) Dumort. Lophozia excisa var. jurensis (Meyl. ex Müll. Frib.) Müll. Frib. = Lophozia excisa (Dicks.) Dumort. Lophozia exsecta (Schmidel ex Schrad.) Dumort. = Tritomaria exsecta (Schmidel ex Schrad.) Loeske Lophozia exsectiformis (Breidl.) Jaap = Tritomaria exsectiformis (Breidl.) Loeske Lophozia floerkei (F. Weber & D. Mohr) Schiffn. = Barbilophozia floerkei (F. Weber & D. Mohr) Loeske Lophozia gillmanii (Austin) R. M. Schust. = Leiocolea gillmanii (Austin) A. Evans Lophozia gracillima H. Buch = Lophozia ascendens (Warnst.) R. M. Schust. Lophozia gracilis Schleich. ex Steph. nom. illeg. = Barbilophozia attenuata (Mart.) Loeske Lophozia guttulata (Lindb. & Arnell) A. Evans = Lophozia longiflora (Nees) Schiffn. Lophozia hatcheri (A. Evans) Steph. = Barbilophozia hatcheri (A. Evans) Loeske Lophozia heterocolpos (Thed. ex Hartm.) M. Howe = Leiocolea heterocolpos (Thed. ex Hartm.) H. Buch Lophozia hornschuchiana (Nees) Macoun = Leiocolea bantriensis (Hook.) Jörg. Lophozia incisa subsp. opacifolia (Culm. ex Meyl.) R. M. Schust. & Damsh. = Lophozia opacifolia Culm. ex Meyl. Lophozia inflata (Huds.) M. Howe = Gymnocolea inflata (Huds.) Dumort. Lophozia jurensis Meyl. ex Müll. Frib. = Lophozia excisa (Dicks.) Dumort. Lophozia kaurinii (Limpr.) Steph. = Leiocolea gillmanii (Austin) A. Evans Lophozia kiaeri Jörg. p. p. = Lophozia excisa (Dicks.) Dumort. Lophozia kunzeana (Huebener) A. Evans = Barbilophozia kunzeana (Huebener) Müll. Frib. Lophozia longiflora sensu auct. = Lophozia ventricosa (Dicks.) Dumort. Lophozia lycopodioides (Wallr.) Cogn. = Barbilophozia lycopodioides (Wallr.) Loeske Lophozia lycopodioides var. parvifolia Schiffn. = Barbilophozia hatcheri (A. Evans) Loeske Lophozia lyoni (Taylor) Steph. = Tritomaria quinquedentata (Huds.) H. Buch Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 409 Lophozia minuta (Schreb.) Schiffn. = Anastrophyllum minutum (Schreb.) R. M. Schust. Lophozia muelleri (Nees ex Lindenb.) Dumort. = Leiocolea collaris (Nees) Schljakov Lophozia porphyroleuca (Nees) Schiffn. nom. illeg. = Lophozia longiflora (Nees) Schiffn. Lophozia quadriloba (Lindb.) A. Evans = Barbilophozia quadriloba (Lindb.) Loeske Lophozia propagulifera (Gottsche) Steph. = Lophozia latifolia R. M. Schust. [36] Lophozia quinquedentata (Huds.) Cogn. = Tritomaria quinquedentata (Huds.) H. Buch Lophozia rufescens Schljakov = Lophozia sudetica (Nees ex Huebener) Grolle Lophozia schreberi Boulay nom. illeg. = Barbilophozia barbata (Schmidel ex Schreb.) Loeske Lophozia scutata (F. Weber & D. Mohr) Dumort. = Harpanthus scutatus (F. Weber & D. Mohr) Spruce Lophozia turbinata (Raddi) Steph. = Leiocolea turbinata (Raddi) H. Buch Lophozia turbinata var. wilsoniana (Nees) C. Massal. = Leiocolea turbinata (Raddi) H. Buch Lophozia ventricosa var. longiflora (Nees) Macoun = Lophozia longiflora (Nees) Schiffn. Lophozia ventricosa var. longiflora auct. non (Nees) Macoun = Lophozia ventricosa (Dicks.) Dumort. Lophozia ventricosa var. porphyroleuca (Nees) Müll. Frib. = Lophozia longiflora (Nees) Schiffn. Lophozia ventricosa var. silvicola (H. Buch) E. W. Jones ex R. M. Schust. = Lophozia silvicola H. Buch Lophozia ventricosa var. uliginosa Breidl. ex Schiffn. = Lophozia ventricosa (Dicks.) Dumort. Lunularia alpina Nees = Sauteria alpina (Nees) Nees Lunularia dillenii Le Jol. = Lunularia cruciata (L.) Lindb. Lunularia michelii Le Jol. = Lunularia cruciata (L.) Lindb. Lunularia vulgaris Raddi nom. illeg. = Lunularia cruciata (L.) Lindb. Madotheca baueri Schiffn. = Porella xbaueri (Schiffn.) C. E. O. Jensen Madotheca canariensis (F. Weber) Nees = Porella canariensis (F. Weber) Underw. # Madotheca canariensis var. subsquarrosa Schiffn. = Porella canariensis (F. Weber) Underw. Madotheca cordaeana (Huebener) Dumort. = Porella cordaeana (Huebener) Moore Madotheca cordaeana var. distans (Müll. Frib.) Müll. Frib. = Porella cordaeana (Huebener) Moore Madotheca dentata (Hartm.) C. Massal. = Porella cordaeana (Huebener) Moore Madotheca inaequalis Gottsche ex Steph. = Porella inaequalis (Gottsche ex Steph.) Perss. Madotheca jackii Schiffn. = Porella platyphylla (L.) Pfeiff. Madotheca kabylica Trab. nom. inval. = Porella cordaeana (Huebener) Moore Madotheca laevigata (Schrad.) Dumort. = Porella arboris-vitae (With.) Grolle Madotheca laevigata var. obscura Nees = Porella arboris-vitae (With.) Grolle Madotheca laevigata var. killarniensis (Pearson) Macvicar = Porella arboris-vitae (With.) Grolle Madotheca laevigata var. subintegra (Kaal.) Schiffn. = Porella arboris-vitae (With.) Grolle ☞ Madotheca laevigata var. thuja (Nees) Gottsche et al. = Porella arboris-vitae (With.) Grolle Madotheca levieri (J. B. Jack & Steph.) Copp. = Porella cordaeana (Huebener) Moore Madotheca obscura (Nees) Boulay = Porella arboris-vitae (With.) Grolle Madotheca obscura (Nees) Warnst. nom. inval. = Porella arboris-vitae (With.) Grolle Madotheca obtusata Taylor = Porella obtusata (Taylor) Trevis. 410 R.M. Ros et al. Madotheca platyphylla (L.) Dumort. = Porella platyphylla (L.) Pfeiff. Madotheca platyphylla var. subsquarrosa Schiffn. = Porella platyphylla (L.) Pfeiff. Madotheca platyphylloidea (Schwein.) Nees = Porella platyphylla (L.) Pfeiff. Madotheca porella Nees nom. illeg. = Porella pinnata L. Madotheca porelloides De Not. ex Lindb. nom. inval. = Porella platyphylla (L.) Pfeiff. Madotheca rivularis Nees = Porella cordaeana (Huebener) Moore Madotheca simplicior J. E. Zetterst. = Porella cordaeana (Huebener) Moore Madotheca thuja (Hook.) Dumort. nom. illeg. = Porella arboris-vitae (With.) Grolle Madotheca thuja auct. non (Hook.) Dumort. = Porella obtusata (Taylor) Trevis. Madotheca torva De Not. ex Bertol. nom. inval. = Porella obtusata (Taylor) Trevis. Mannia atlantica Jelenc nom. inval. [39] Mannia dichotoma Udar & V. Chandra nom. inval. = Mannia androgyna (L.) A. Evans Mannia dichotoma S. W. Arnell = Mannia androgyna (L.) A. Evans Mannia rupestris (Nees) Frye & L. Clark = Mannia triandra (Scop.) Grolle Marchantia alpestris (Nees) Burgeff = Marchantia polymorpha subsp. montivagans Bischl. & Boisselier Marchantia androgyna L. = Mannia androgyna (L.) A. Evans Marchantia angustifolia Neck. nom. illeg. = Mannia androgyna (L.) A. Evans Marchantia aquatica (Nees) Burgeff = Marchantia polymorpha L. subsp. polymorpha Marchantia circumscissa Biv. = Mannia androgyna (L.) A. Evans Marchantia commutata Lindenb. = Preissia quadrata (Scop.) Müll. Frib. Marchantia conica L. = Conocephalum conicum (L.) Dumort. Marchantia cruciata L. = Lunularia cruciata (L.) Lindb. Marchantia cruciata var. hyalina Sommerf. = Athalamia hyalina (Sommerf.) S. Hatt. $ Marchantia dillenii = Lunularia cruciata (L.) Lindb. Marchantia fragrans Balbis = Mannia fragrans (Balbis) Frye & L. Clark [10] Marchantia glauca Biv. = Plagiochasma rupestre (J. R. Forst. & G. Forst.) Steph. Marchantia gracilis F. Weber = Asterella gracilis (F. Weber) Underw. Marchantia hemisphaerica L. = Reboulia hemisphaerica (L.) Raddi Marchantia hirsuta Sw. = Dumortiera hirsuta (Sw.) Nees Marchantia hyalina Sommerf. = Athalamia hyalina (Sommerf.) S. Hatt. Marchantia irrigua Wilson ex Hook. = Dumortiera hirsuta (Sw.) Nees Marchantia pilosa Hornem. = Mannia pilosa (Hornem.) Frye & L. Clark Marchantia polycephala Schleich. nom. inval. = Asterella gracilis (F. Weber) Underw. Marchantia polymorpha auct. = Marchantia polymorpha subsp. ruderalis Bischl. & Boisselier Marchantia polymorpha var. alpestris (Nees) Nees = Marchantia polymorpha subsp. montivagans Bischl. & Boisselier Marchantia polymorpha var. aquatica (Nees) Gottsche et al. = Marchantia polymorpha L. subsp. polymorpha Marchantia polymorpha f. domestica Nees nom. illeg. = Marchantia polymorpha subsp. ruderalis Bischl. & Boisselier Marchantia polymorpha var. mamillata I. Hag. ex Schiffn. = Marchantia polymorpha subsp. montivagans Bischl. & Boisselier Marchantia quadrata Scop. = Preissia quadrata (Scop.) Müll. Frib. [51] Marchantia spathysii Lindenb. = Athalamia spathysii (Lindenb.) S. Hatt. Marchantia stellata Scop. = Marchantia polymorpha subsp. ruderalis Bischl. & Boisselier Marchantia triandra Scop. = Mannia triandra (Scop.) Grolle Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 411 Marchantia umbellata Scop. = Marchantia polymorpha subsp. ruderalis Bischl. & Boisselier Marchantia vittata Raddi = Marchantia polymorpha L. subsp. polymorpha Marchesinia johnsoniana (Mitt.) Herz. = Acanthocoleus aberrans (Lindenb. & Gottsche) Kruijt ☞ Marsilia epiphylla (L.) Lindb. nom. illeg. = Pellia epiphylla (L.) Corda Marsupella aemula (Limpr.) Lindb. ex Kaal. = Marsupella condensata (Ångstr. ex C. Hartm.) Kaal. Marsupella aquatica (Lindenb.) Schiffn. = Marsupella emarginata (Ehrh.) Dumort. Marsupella badensis Schiffn. = Marsupella funckii (F. Weber & D. Mohr) Dumort. Marsupella capillaris (Limpr.) H. Bern. = Marsupella boeckii (Austin) Lindb. Marsupella conferta (Limpr.) Spruce = Marsupella brevissima (Dumort.) Grolle Marsupella emarginata subsp. aquatica (Lindenb.) Meyl. = Marsupella emarginata (Ehrh.) Dumort. Marsupella emarginata var. aquatica (Lindenb.) Dumort. = Marsupella emarginata (Ehrh.) Dumort. Marsupella emarginata var. minor C. Massal. ex Schiffn. nom. illeg. = Marsupella emarginata (Ehrh.) Dumort. Marsupella erythrorhiza Schiffn. = Marsupella sphacelata (Gieseke ex Lindenb.) Dumort. ☞ Marsupella filiformis (Lindb.) Lindb. ex Kaal. = Marsupella boeckii (Austin) Lindb. Marsupella funckii var. major (Nees) Boulay = Marsupella funckii (F. Weber & D. Mohr) Dumort. ☞ Marsupella gracilis (C. Massal. & Carestia) Pearson = Marsupella sprucei (Limpr.) Bernet Marsupella hungarica Boros & Vajda nom. inval. = Marsupella funckii (F. Weber & D. Mohr) Dumort. Marsupella joergensenii Schiffn. = Marsupella sphacelata (Gieseke ex Lindenb.) Dumort. Marsupella latifolia Lindb. nom. inval. = Marsupella boeckii (Austin) Lindb. ☞ Marsupella neglecta (Limpr.) Lindb. = Marsupella sprucei (Limpr.) Bernet Marsupella nevicensis (Carrington) Pearson = Marsupella boeckii (Austin) Lindb. Marsupella pearsonii Schiffn. ex Macvicar = Marsupella emarginata (Ehrh.) Dumort. Marsupella pygmaea (Limpr.) Steph. = Marsupella funckii (F. Weber & D. Mohr) Dumort. Marsupella revoluta (Nees) Dumort. = Apomarsupella revoluta (Nees) R. M. Schust. Marsupella robusta (De Not.) A. Evans = Marsupella emarginata (Ehrh.) Dumort. Marsupella styriaca (Limpr.) Kaal. = Marsupella sparsifolia (Lindb.) Dumort. Marsupella sullivantii (De Not.) A. Evans = Marsupella sphacelata (Gieseke ex Lindenb.) Dumort. Marsupella ustulata (Huebener) Spruce ex Pearson hom. illeg. = Marsupella emarginata (Ehrh.) Dumort. Marsupella ustulata Spruce non (Huebener) Spruce ex Pearson hom. illeg. = Marsupella sprucei (Limpr.) Bernet Marsupella ustulata var. neglecta (Limpr.) Müll. Frib. = Marsupella sprucei (Limpr.) Bernet Marsupella varians (Lindb.) Müll. Frib. = Marsupella brevissima (Dumort.) Grolle Martinellia aequiloba (Schwägr.) Lindb. nom. illeg. = Scapania aequiloba (Schwägr.) Dumort. ☞ Martinellia apiculata (Spruce) Trevis. = Scapania apiculata Spruce ☞ Martinellia aspera (Bernet & M. Bernet) Broth. = Scapania aspera Bernet & M. Bernet Martinellia calcicola Arnell & J. Perss. = Scapania calcicola (Arnell & J. Perss.) Ingham Martinellia convexa (Scop.) Lindb. = Scapania umbrosa (Schrad.) Dumort. Martinellia crassiretis (Bryhn) Arnell = Scapania crassiretis Bryhn Martinellia curta (Mart.) Lindb. = Scapania curta (Mart.) Dumort. 412 R.M. Ros et al. Martinellia cuspiduligera (Nees) Arnell = Scapania cuspiduligera (Nees) Müll. Frib. Martinellia gracilis (Lindb.) Lindb. = Scapania gracilis Lindb. ☞ Martinellia helvetica (Gottsche) Trevis. = Scapania helvetica Gottsche Martinellia mucronata (H. Buch) Arnell & C. E. O. Jensen = Scapania mucronata H. Buch Martinellia nemorosa Lindb. nom. illeg. = Scapania nemorea (L.) Grolle Martinellia obliqua Arnell = Scapania uliginosa (Sw. ex Lindenb.) Dumort. Martinellia obscura Arnell & C. E. O. Jensen = Scapania obscura (Arnell & C. E. O. Jensen) Schiffn. Martinellia paludicola (Loeske & Müll. Frib.) C. E. O. Jensen = Scapania paludicola Loeske & Müll. Frib. ☞ Martinellia paludosa (Müll. Frib.) Arnell & C. E. O. Jensen = Scapania paludosa (Müll. Frib.) Müll. Frib. Martinellia resupinata (L.) Gray ex Lindb. = Scapania undulata (L.) Dumort. ☞ Martinellia resupinata (Hook.) Lindb. nom. inval. = Scapania compacta (A. Roth) Dumort. Martinellia rosacea (Corda) Lindb. = Scapania curta (Mart.) Dumort. Martinellia scandica Arnell & H. Buch = Scapania scandica (Arnell & H. Buch) Macvicar ☞ Martinellia subalpina (Nees ex Lindenb.) Trevis. = Scapania subalpina (Nees ex Lindenb.) Dumort. ☞ Martinellia uliginosa (Sw. ex Lindenb.) Trevis. = Scapania uliginosa (Sw. ex Lindenb.) Dumort. ☞ Martinellia umbrosa (Schrad.) Gray = Scapania umbrosa (Schrad.) Dumort. Martinellia undulata (L.) Gray = Scapania undulata (L.) Dumort. Massula capitata (Hook.) Schljakov = Lophozia capitata (Hook.) Macoun Massula grandiretis (Lindb. ex Kaal.) Schljakov = Lophozia grandiretis (Lindb. ex Kaal.) Schiffn. Massula incisa (Schrad.) Schljakov nom. illeg.= Lophozia incisa (Schrad.) Dumort. Massula opacifolia (Culm. ex Meyl.) Schljakov nom. illeg. = Lophozia opacifolia Culm. ex Meyl. Mastigobryum deflexum (Mart.) Nees = Bazzania tricrenata (Wahlenb.) Lindb. Mastigobryum triangulare Schleich. ex Steph. nom. illeg. = Bazzania tricrenata (Wahlenb.) Lindb. Mastigobryum trilobatum (L.) Nees = Bazzania trilobata (L.) Gray Mesophylla crenulata (Mitt.) Corb. = Jungermannia gracillima Sm. Mesophylla crenulata var. gracillima (Sm.) Boulay nom. illeg. = Jungermannia gracillima Sm. Mesophylla hyalina (Lyell) Corb. = Jungermannia hyalina Lyell Mesophylla hyalina var. gracillima (Schiffn.) Boulay = Jungermannia hyalina Lyell Mesophylla nigrella (De Not.) Boulay = Southbya nigrella (De Not.) Henriq. Mesophylla obovata (Nees) Corb. = Jungermannia obovata Nees Mesophylla rotaeana (De Not) Dumort. = Nardia scalaris Gray Mesophylla scalaris (Gray) Dumort. = Nardia scalaris Gray Mesophylla stillicidiorum (Raddi) Boulay = Southbya tophacea (Spruce) Spruce Metzgeria conjugata subsp. simplex (Lorb. ex Müll. Frib.) R. M. Schust. = Metzgeria simplex Lorb. ex Müll. Frib. # Metzgeria conjugata f. subpinnata Jörg. = Metzgeria conjugata Lindb. Metzgeria densiseta Steph. = Metzgeria furcata (L.) Dumort. Metzgeria fruticulosa auct. eur. non (O. F. Müll.) Evans = Metzgeria violacea (Ach.) Dumort. [40] Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 413 Metzgeria furcata var. fruticulosa auct. eur. non (O. F. Müll.) Lindb. = Metzgeria violacea (Ach.) Dumort. [40] ☞ Metzgeria furcata var. ulvula (Nees) Gottsche et al. = Metzgeria furcata (L.) Dumort. Metzgeria glabra Raddi nom. illeg. = Metzgeria furcata (L.) Dumort. Metzgeria glabra f. ulvula (Nees) Zodda = Metzgeria furcata (L.) Dumort. Metzgeria hamata Lindb. nom. illeg. = Metzgeria leptoneura Spruce Metzgeria nitida Mitt. = Metzgeria leptoneura Spruce Metzgeria pubescens (Schrank) Raddi = Apometzgeria pubescens (Schrank) Kuwah. Microlejeunea diversiloba auct. non (Spruce) Müll. Frib. = Lejeunea hibernica Bischl. et al. ex Grolle Microlejeunea mandonii Steph. = Lejeunea mandonii (Steph.) Müll. Frib. Microlepidozia setacea auct. non (G. Web.) Jørg. = Kurzia pauciflora (Dicks.) Grolle Microlepidozia sylvatica (A. Evans) Jørg. = Kurzia sylvatica (A. Evans) Grolle Microlepidozia trichoclados (Müll. Frib.) Jørg. = Kurzia trichoclados (Müll. Frib.) Grolle Mnium fissum L. = Calypogeia fissa (L.) Raddi Mnium trichomanis L. = Calypogeia fissa (L.) Raddi Mnium trichomanis auct. non L. = Calypogeia azurea Stotler & Crotz [11] Moerckia flotoviana (Nees) Schiffn. = Moerckia hibernica (Hook.) Gottsche Mylia cuneifolia (Hook.) Gray = Leptoscyphus cuneifolius (Hook.) Mitt. Nardia alpina (Gottsche ex Husn.) Trevis. nom. inval. = Marsupella alpina (Gottsche ex Husn.) Bernet Nardia boeckii (Austin) Lindb. = Marsupella boeckii (Austin) Lindb. Nardia brevissima (Dumort.) Lindb. = Marsupella brevissima (Dumort.) Grolle Nardia cochlearis Lindb. = Marsupella brevissima (Dumort.) Grolle Nardia commutata (Limpr.) C. Massal. & Carestia = Marsupella commutata (Limpr.) Bernet Nardia compressa var. rigida Lindb. = Nardia compressa (Hook.) Gray Nardia crassula Lorb. nom. inval. = Nardia insecta Lindb. Nardia crenulata (Mitt.) Lindb. = Jungermannia gracillima Sm. Nardia crenulata var. cristulata (Dumort.) Schiffn. = Jungermannia gracillima Sm. Nardia crenulata var. gracillima (Sm.) Lindb. = Jungermannia gracillima Sm. Nardia crenulata var. inundata Schiffn. = Jungermannia gracillima Sm. ☞ Nardia densifolia (Nees) Trevis. = Marsupella emarginata (Ehrh.) Dumort. Nardia emarginata (Ehrh.) Gray = Marsupella emarginata (Ehrh.) Dumort. Nardia emarginata var. aquatica (Lindenb.) Carrington = Marsupella emarginata (Ehrh.) Dumort. Nardia filiformis Lindb. = Marsupella boeckii (Austin) Lindb. ☞ Nardia funckii (F. Weber & D. Mohr) Carrington ex Lindb. = Marsupella funckii (F. Weber & D. Mohr) Dumort. Nardia geoscyphus var. insecta (Lindb.) Schiffn. = Nardia insecta Lindb. Nardia gracilis C. Massal. & Carestia = Marsupella sprucei (Limpr.) Bernet Nardia gracillima (Sm.) Lindb. = Jungermannia gracillima Sm. Nardia haematosticta Nees ex Lindb. nom. inval. = Nardia geoscyphus (De Not.) Lindb. Nardia hyalina (Lyell) Carrington = Jungermannia hyalina Lyell Nardia hyalina var. colorata Samp. nom. inval. [30] Nardia latifolia Lindb. nom. inval. = Marsupella boeckii (Austin) Lindb. Nardia levieri Steph. = Jungermannia confertissima Nees Nardia lignicola Schiffn. = Jungermannia lignicola (Schiffn.) Grolle 414 R.M. Ros et al. Nardia muelleriana Schiffn. = Jungermannia hyalina Lyell Nardia obovata (Nees) Lindb. = Jungermannia obovata Nees Nardia obovata var. minor Carrington = Jungermannia subelliptica (Lindb. ex Kaal.) Levier Nardia pachyphylla (De Not.) Trevis. = Nardia compressa (Hook.) Gray Nardia revoluta (Nees) Lindb. = Apomarsupella revoluta (Nees) R. M. Schust. ☞ Nardia robusta (De Not.) Lindb. ex Carrington = Marsupella emarginata (Ehrh.) Dumort. ☞ Nardia silvrettae (Gottsche ex Dumort.) Trevis. [41] Nardia sparsifolia (Lindb.) Lindb. = Marsupella sparsifolia (Lindb.) Dumort. Nardia sparsifolia var. minor (Schleich.) Lindb. nom. inval. = Marsupella brevissima (Dumort.) Grolle Nardia sphacelata (Gieseke ex Lindenb.) Carrington = Marsupella sphacelata (Gieseke ex Lindenb.) Dumort. Nardia sprucei (Limpr.) C. Massal. & Carestia = Marsupella sprucei (Limpr.) Bernet Nardia subelliptica Lindb. ex Kaal. = Jungermannia subelliptica (Lindb. ex Kaal.) Levier Nardia subtilissima Schiffn. = Jungermannia subtilissima (Schiffn.) Grolle Nardia varians Lindb. = Marsupella brevissima (Dumort.) Grolle Neesiella carnica (C. Massal.) Schiffn. = Mannia pilosa (Hornem.) Frye & L. Clark Neesiella pilosa (Hornem.) Schiffn. = Mannia pilosa (Hornem.) Frye & L. Clark Neesiella rupestris (Nees) Schiffn. = Mannia triandra (Scop.) Grolle Obtusifolium obtusum (Lindb.) S. W. Arnell = Lophozia obtusa (Lindb.) A. Evans Odontoschisma cavifolium Steph. = Odontoschisma denudatum (Mart.) Dumort. Odontoschisma denudatum subsp. elongatum (Lindb.) Potemkin = Odontoschisma elongatum (Lindb.) A. Evans Odontoschisma denudatum var. elongatum (Lindb.) Lindb. = Odontoschisma elongatum (Lindb.) A. Evans Odontoschisma denudatum f. elongatum Lindb. = Odontoschisma elongatum (Lindb.) A. Evans Odontoschisma remotifolium Warnst. = Odontoschisma sphagni (Dicks.) Dumort. Odontoschisma sphagni var. denudatum (Mart.) C. Massal. & Carestia = Odontoschisma denudatum (Mart.) Dumort. Odontoschisma sphagni var. macrior Bernet = Odontoschisma denudatum (Mart.) Dumort. Odontoschisma tessellatum (Berggr.) C. E. O. Jensen = Odontoschisma macounii (Austin) Underw. Orthocaulis atlanticus (Kaal.) H. Buch = Barbilophozia atlantica (Kaal.) Müll. Frib. Orthocaulis attenuatus (Mart.) A. Evans = Barbilophozia attenuata (Mart.) Loeske Orthocaulis floerkei (F. Weber & D. Mohr) H. Buch = Barbilophozia floerkei (F. Weber & D. Mohr) Loeske Orthocaulis gracilis (Schleich. ex DC.) H. Buch = Barbilophozia attenuata (Mart.) Loeske Orthocaulis kunzeanus (Huebener) H. Buch = Barbilophozia kunzeana (Huebener) Müll. Frib. Otiona crinita (Michx.) Corda = Reboulia hemisphaerica (L.) Raddi Otiona italica (Sassi) Dumort. = Plagiochasma rupestre (J. R. Forst. & G. Forst.) Steph. Otiona rupestris (J. R. Forst. & G. Forst.) Dumort. = Plagiochasma rupestre (J. R. Forst. & G. Forst.) Steph. Oxymitra paleacea Bisch. ex Lindenb. = Oxymitra incrassata (Brot.) Sérgio & Sim-Sim Oxymitra pyramidata (Raddi) Huebener non (Willd.) Huebener = Oxymitra incrassata (Brot.) Sérgio & Sim-Sim Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 415 Oxymitra pyramidata var. linkii (Lindenb.) Lindenb. = Oxymitra incrassata (Brot.) Sérgio & Sim-Sim Oxymitra pyramidata var. paleacea (Bisch. ex Lindenb.) Gottsche et al. = Oxymitra incrassata (Brot.) Sérgio & Sim-Sim Pallavicinia blyttii (Moerch ex Hornem.) Lindb. = Moerckia blyttii (Moerch ex Hornem.) Brockm. Pallavicinia flotoviana (flotowii error pro flotoviana) (Nees) Lindb. = Moerckia hibernica (Hook.) Gottsche Pallavicinia flotoviana var. hibernica (Hook.) Lindb. nom. illeg. = Moerckia hibernica (Hook.) Gottsche Pallavicinia hibernica (Hook.) Gray = Moerckia hibernica (Hook.) Gottsche Pedinophyllum interruptum var. pyrenaicum (Spruce) Kaal. = Pedinophyllum interruptum (Nees) Kaal. Pedinophyllum pyrenaicum (Spruce) Lindb. = Pedinophyllum interruptum (Nees) Kaal. Pedinophyllum pyrenaicum var. interruptum (Nees) Schiffn. nom. incorrect. = Pedinophyllum interruptum (Nees) Kaal. Pellia borealis Lorb. = Pellia epiphylla (L.) Corda Pellia calycina (Taylor) Nees = Pellia endiviifolia (Dicks.) Dumort. Pellia endiviifolia f. furcigera (Hook.) Schiffn. = Pellia endiviifolia (Dicks.) Dumort. Pellia epiphylla subsp. borealis (Lorb.) Messe = Pellia epiphylla (L.) Corda Pellia epiphylla f. neesiana Gottsche = Pellia neesiana (Gottsche) Limpr. Pellia fabbroniana auct. non Raddi = Pellia endiviifolia (Dicks.) Dumort. $ Pellia fabbroniana var. furcigera = Pellia endiviifolia (Dicks.) Dumort. Pellia fabbroniana f. furcigera (Hook.) C. Massal. = Pellia endiviifolia (Dicks.) Dumort. Pellia fabbroniana f. undulata Jelenc nom. inval. [42] Pellia fuciformis (Bory) Nees [43] Peltolepis grandis (Lindb.) Lindb. = Peltolepis quadrata (Saut.) Müll. Frib. Peltolepis sibirica Lindb. = Peltolepis quadrata (Saut.) Müll. Frib. Petalophyllum lamellatum (Nees) Lindb. = Petalophyllum ralfsii (Wilson) Nees & Gottsche Phragmicoma mackaii (Hook.) Dumort. = Marchesinia mackaii (Hook.) Gray Physocolea rossettiana (C. Massal.) Steph. = Cololejeunea rossettiana (C. Massal.) Schiffn. Plagiochasma algericum Steph. = Plagiochasma rupestre (J. R. Forst. & G. Forst.) Steph. Plagiochasma aitonia Lindenb. & Nees = Plagiochasma rupestre (J. R. Forst. & G. Forst.) Steph. Plagiochasma beccarianum Steph. REJECTED FROM THE AREA Plagiochasma italicum (Sassi) De Not. = Plagiochasma rupestre (J. R. Forst. & G. Forst.) Steph. Plagiochasma roussellianum Mont. = Athalamia spathysii (Lindenb.) S. Hatt. Plagiochasma sauteria Gottsche ex Jelenc nom. inval. = Athalamia spathysii (Lindenb.) S. Hatt. Plagiochila aequiloba (Schwägr.) Mont. & Nees = Scapania aequiloba (Schwägr.) Dumort. Plagiochila africana Steph. REJECTED FROM THE AREA [44] Plagiochila asplenioides auct. = Plagiochila porelloides (Torrey ex Nees) Lindenb. Plagiochila asplenioides var. humilis (Nees) Lindenb. = Plagiochila porelloides (Torrey ex Nees) Lindenb. Plagiochila asplenioides var. major Nees = Plagiochila asplenioides (L. emend. Taylor) Dumort. Plagiochila asplenioides var. minor Lindenb. = Plagiochila porelloides (Torrey ex Nees) Lindenb. 416 R.M. Ros et al. Plagiochila asplenioides subsp. porelloides (Torrey ex Nees) Kaal. = Plagiochila porelloides (Torrey ex Nees) Lindenb. Plagiochila asplenioides var. riparia Breidl. = Plagiochila porelloides (Torrey ex Nees) Lindenb. Plagiochila atlantica F. Rose = Plagiochila heterophylla Lindenb. ex Lehm. [46] Plagiochila compacta (A. Roth) Mont. & Nees = Scapania compacta (A. Roth) Dumort. Plagiochila corniculata auct. non (Dumort.) Dumort. = Plagiochila exigua (Taylor) Taylor Plagiochila curta (Mart.) Mont. & Nees = Scapania curta (Mart.) Dumort. Plagiochila diacantha Taylor ex Steph. = Plagiochila bifaria (Sw.) Lindenb. Plagiochila dubia Lindenb. & Gottsche = Plagiochila patula (Sw.) Lindenb. REJECTED FROM THE AREA Plagiochila interrupta (Nees) Mont. & Nees = Pedinophyllum interruptum (Nees) Kaal. Plagiochila javanica (Sw.) Nees & Mont. REJECTED FROM THE AREA Plagiochila killarniensis Pearson = Plagiochila bifaria (Sw.) Lindenb. Plagiochila major (Nees) S. W. Arnell nom. illeg. = Plagiochila asplenioides (L. emend. Taylor) Dumort. Plagiochila punctata var. owenii (Steph.) Macvicar = Plagiochila punctata (Taylor) Taylor Plagiochila pyrenaica Spruce = Pedinophyllum interruptum (Nees) Kaal. Plagiochila sharpii H. L. Blomq. = Plagiochila retrorsa Gottsche Plagiochila spinulosa var. inermis Carrington = Plagiochila bifaria (Sw.) Lindenb. Plagiochila spinulosa var. killarniensis (Pearson) Macvicar = Plagiochila bifaria (Sw.) Lindenb. Plagiochila spinulosa var. macaronesiana Allorge & V. Allorge = Plagiochila bifaria (Sw.) Lindenb. Plagiochila spinulosa var. punctata (Taylor) Carrington = Plagiochila punctata (Taylor) Taylor Plagiochila spinulosa var. tridenticulata auct. non (Michx.) Husn. nom. illeg. = Plagiochila exigua (Taylor) Taylor Plagiochila stableri Pearson = Plagiochila exigua (Taylor) Taylor Plagiochila tridenticulata auct. non (Michx.) Dumort. = Plagiochila exigua (Taylor) Taylor Plagiochila undulata (L.) Mont. & Nees = Scapania undulata (L.) Dumort. Plectocolea crenulata (Mitt.) A. Evans = Jungermannia gracillima Sm. Plectocolea crenulata var. gracillima (Sm.) Frye & L. Clark = Jungermannia gracillima Sm. Plectocolea hyalina (Lyell) Mitt. = Jungermannia hyalina Lyell Plectocolea obovata (Nees) Lindb. = Jungermannia obovata Nees Plectocolea subelliptica (Lindb. ex Kaal.) A. Evans = Jungermannia subelliptica (Lindb. ex Kaal.) Levier Pleuroclada albescens (Hook.) Spruce nom. illeg. = Pleurocladula albescens (Hook.) Grolle Pleuroschisma flaccidum Dumort. = Bazzania flaccida (Dumort.) Grolle Pleuroschisma implexum (Nees) Meyl. = Bazzania flaccida (Dumort.) Grolle Pleuroschisma sphagni (Dicks.) Dumort. = Odontoschisma sphagni (Dicks.) Dumort. $ Pleuroschisma triangulare var. implexa = Bazzania flaccida (Dumort.) Grolle Pleuroschisma tricrenatum (Wahlenb.) Dumort. = Bazzania tricrenata (Wahlenb.) Lindb. Pleuroschisma tricrenatum var. implexum (Nees) Müll. Frib. = Bazzania flaccida (Dumort.) Grolle Pleuroschisma trilobatum (L.) Dumort. = Bazzania trilobata (L.) Gray Porella arboris-vitae var. killarniensis (Pearson) M. F. V. Corley = Porella arboris-vitae (With.) Grolle Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 417 Porella arboris-vitae var. obscura (Nees) M.F.V. Corley = Porella arboris-vitae (With.) Grolle Porella cordaeana var. distans (Müll. Frib.) Jelenc = Porella cordaeana (Huebener) Moore Porella cordaeana var. simplicior (J.E. Zetterst.) Arnell = Porella cordaeana (Huebener) Moore Porella dentata (Hartm.) Lindb. = Porella cordaeana (Huebener) Moore Porella kabylica Jelenc nom. inval. = Porella cordaeana (Huebener) Moore Porella laevigata (Schrad.) Pfeiff. = Porella arboris-vitae (With.) Grolle Porella laevigata var. obscura (Nees) Arnell = Porella arboris-vitae (With.) Grolle ☞ Porella laevigata var. thuja (Nees) Arnell = Porella arboris-vitae (With.) Grolle Porella levieri J. B. Jack & Steph. = Porella cordaeana (Huebener) Moore Porella notarisii Trevis. nom. inval. = Porella platyphylla (L.) Pfeiff. Porella platyphylla var. subsquarrosula (Schiffn.) Arnell = Porella platyphylla (L.) Pfeiff. Porella platyphylloidea (Schwein.) Lindb. = Porella platyphylla (L.) Pfeiff. ? Porella porelloides De Not. = Porella platyphylla (L.) Pfeiff. Porella rivularis (Nees) Pfeiff. = Porella cordaeana (Huebener) Moore Porella thuja (Dicks.) Lindb. nom. illeg. = Porella arboris-vitae (With.) Grolle Porella thuja auct. non (Dicks.) Lindb. = Porella obtusata (Taylor) Trevis. Preissia commutata (Lindenb.) Nees = Preissia quadrata (Scop.) Müll. Frib. Preissia cucullata Nees & Mont. = Lunularia cruciata (L.) Lindb. ☞ Preissia hemisphaerica Cogn. = Preissia quadrata (Scop.) Müll. Frib. Preissia italica Corda = Preissia quadrata (Scop.) Müll. Frib. Prionolobus turneri (Hook.) Schiffn. = Cephaloziella turneri (Hook.) Müll. Frib. ☞ Ptilidium ciliare var. pulcherrimum (Weber) Warnst. = Ptilidium pulcherrimum (Weber) Vain. Ptilidium trichophyllum (L.) Mitt. = Blepharostoma trichophyllum (L.) Dumort. Radula bornmuelleri Schiffn. = Radula lindenbergiana Gottsche ex C. Hartm. Radula commutata Gottsche ex J. B. Jack = Radula lindenbergiana Gottsche ex C. Hartm. Radula complanata subsp. lindbergiana (Gottsche ex C. Hartm.) R. M. Schust. = Radula lindenbergiana Gottsche ex C. Hartm. Radula complanata f. propagulifera Nees = Radula lindenbergiana Gottsche ex C. Hartm. Radula complanata var. propagulifera (Nees) Gottsche et al. = Radula lindenbergiana Gottsche ex C. Hartm. Radula germana J. B. Jack = Radula lindenbergiana Gottsche ex C. Hartm. Radula limbata Schiffn. = Radula wichurae Steph. Radula lindbergiana Gottsche ex Jack nom. inval. = Radula lindenbergiana Gottsche ex C. Hartm. Radula lindbergii Gottsche ex Lindb. nom. inval. = Radula lindenbergiana Gottsche ex C. Hartm. Radula notarisii Steph. = Radula complanata (L.) Dumort. Radula ovata J. B. Jack ex Steph. = Radula lindenbergiana Gottsche ex C. Hartm. Radula physoloba Mont. REJECTED FROM THE AREA [52] Reboulia charrieri Douin & R.C.V. Douin = Reboulia hemisphaerica (L.) Raddi ☞ Reboulia hemisphaerica var. macrocephala C. Massal. = Reboulia hemisphaerica (L.) Raddi Reboulia hemisphaerica var. microspora Schiffn. = Reboulia hemisphaerica (L.) Raddi Reboulia hemisphaerica var. pangiensis (Kashyap) Kashyap = Reboulia hemisphaerica (L.) Raddi 418 R.M. Ros et al. Reboulia hemisphaerica subsp. paradoxa R. M. Schust. nom. inval. = Reboulia hemisphaerica (L.) Raddi Reboulia occidentalis Douin & R. C. V. Douin = Reboulia hemisphaerica (L.) Raddi Reboulia quadrata (Scop.) Bertol. = Preissia quadrata (Scop.) Nees [51] Rhakiocarpon conspersum Corda = Reboulia hemisphaerica (L.) Raddi Riccardia latifrons var. sinuata (Dicks. ex Hook.) Lindb. = Riccardia chamedryfolia (With.) Grolle Riccardia major Lindb. = Riccardia chamedryfolia (With.) Grolle Riccardia pinguis (L.) Gray = Aneura pinguis (L.) Dumort. Riccardia pinguis f. angustior (Hook.) P. Allorge = Aneura pinguis (L.) Dumort. l Riccardia pinguis f. submersa Loeske ex Jaap nom. inval. = Aneura pinguis (L.) Dumort. Riccardia sinuata (Dicks. ex Hook.) Trevis. = Riccardia chamedryfolia (With.) Grolle Riccia aculeata Trab. nom. nud. = Riccia crozalsii Levier [55] Riccia aegyptiaca S. W. Arnell = Riccia congoana Steph. Riccia affinis Milde = Riccia ciliata Hoffm. Riccia aggregata Underw. = Riccia nigrella DC. Riccia americana (M. Howe) M. Howe = Riccia lamellosa Raddi Riccia armatissima (Levier ex Müll. Frib.) Trab. = Riccia gougetiana var. armatissima Levier ex Müll. Frib. Riccia arvensis Austin = Riccia bifurca Hoffm. Riccia atromarginata var. glabra Levier ex Müll. Frib. = Riccia trabutiana Steph. Riccia austinii Steph. = Riccia lamellosa Raddi Riccia battandieri Trab. nom. nud. [55] Riccia baumgartneri Schiffn. = Riccia subbifurca Warnst. ex Croz. Riccia bavarica Warnst. = Riccia warnstorfii Limpr. ex Warnst. Riccia beckeriana Steph. = Riccia frostii Austin Riccia bifurca f. subinermis (Heeg) Müll. Frib. = Riccia bifurca var. subinermis Heeg Riccia bischoffii Huebener = Riccia ciliifera Link ex Lindenb. Riccia bischoffii f. armata Trab. nom. nud. = Riccia gougetiana var. armatissima Levier ex Müll. Frib. [55] Riccia bischoffii var. ciliifera (Link. ex Lindenb.) Müll. Frib. = Riccia ciliifera Link ex Lindenb. Riccia bischoffii var. fimbriata Gottsche et al. = Riccia ciliifera Link ex Lindenb. Riccia bischoffii f. maxima Levier = Riccia gougetiana Durieu & Mont. Riccia bischoffii var. subtumida Milde = Riccia ciliata Hoffm. Riccia bullosa Link ex Lindenb. REJECTED FROM THE AREA Riccia bullosa sensu Müll. Frib. = Exormotheca welwitschii Steph. Riccia cana Durieu ex Trab. nom. illeg. = Riccia crinita Taylor Riccia canescens Steph. = Riccia crinita Taylor Riccia capensis Steph. = Riccia limbata Bisch. ex Krauss REJECTED FROM THE AREA Riccia capillata Schmidel nom. illeg. = Ricciocarpos natans (L.) Corda Riccia centrifuga Arnell = Riccia fluitans L. Riccia chinensis Herz. [54] Riccia chudoana Steph. [54] Riccia ciliata Raddi nom. illeg. = Riccia michelii Raddi Riccia ciliata var. intumescens Bisch. = Riccia ciliata Hoffm. [57] Riccia ciliata var. epilosa Warnst. = Riccia ciliata Hoffm. Riccia ciliifera f. pedemontana (Steph.) Müll. Frib. = Riccia ciliifera Link ex Lindenb. Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 419 Riccia ciliata var. violacea Kny = Riccia ciliata Hoffm. Riccia commutata J. B. Jack ex Levier = Riccia warnstorfii Limpr. ex Warnst. Riccia commutata var. acrotricha Levier ex Sommier = Riccia warnstorfii var. subinermis Warnst. Riccia concava Bisch. REJECTED FROM THE AREA Riccia convexa Steph. [54] Riccia coriandrina Spreng. = Corsinia coriandrina (Spreng.) Lindb. l Riccia crozalsii f. major C. Massal. nom. nud. = Riccia crozalsii Levier Riccia crystallina auct. = Riccia cavernosa Hoffm. Riccia crystallina var. angustior Lindenb. ex Nees = Riccia cavernosa Hoffm. Riccia crystallina subsp. austrigena Schiffn. = Riccia crystallina L. emend. Raddi Riccia crystallina var. major Lindb. = Riccia crystallina L. emend. Raddi Riccia crystallina var. vulgaris Lindenb. = Riccia crystallina L. emend. Raddi Riccia dufourii Nees = Riccia lamellosa Raddi Riccia echinulata Trab. nom. nud. = Riccia ciliifera Link ex Lindenb. [55] Riccia epicarpa Wallr. ex Gottsche et al. = Riccia sorocarpa Bisch. Riccia erinacea Schiffn. = Riccia gougetiana var. armatissima Levier ex Müll. Frib. Riccia eudichotoma Bisch. nom. illeg. = Riccia canaliculata Hoffm. Riccia eudichotoma var. fluitans (L.) Bisch. nom. incorrect. & nom. illeg. = Riccia fluitans L. Riccia fluitans var. canaliculata (Hoffm.) A. Roth = Riccia canaliculata Hoffm. Riccia fluitans var. purpurascens Klinggr. = Riccia huebeneriana Lindenb. Riccia frostii var. crystallinoides Schiffn. [58] Riccia fruticulosa auct. eur. non O. F. Müll. = Metzgeria violacea (Ach.) Dumort. [40] Riccia glauca var. subinermis (Lindb.) Warnst. = Riccia glauca var. ciliaris Warnst. Riccia glaucescens Carrington = Riccia beyrichiana Hampe ex Lehm. Riccia gougetiana var. erinacea Schiffn. nom. inval. = Riccia gougetiana var. armatissima Levier ex Müll. Frib. Riccia henriquesii Levier = Riccia bicarinata Lindb. Riccia henriquesii var. mediterranea C. Massal. = Riccia bicarinata Lindb. Riccia heyeri (Huebener ex Genth) Müll. Frib. = Riccia canaliculata Hoffm. Riccia huebeneriana var. cavernosa (Hoffm.) Ast = Riccia cavernosa Hoffm. Riccia huebeneriana var. pseudo-frostii Schiffn. = Riccia huebeneriana Lindenb. Riccia hyi Levier ex Müll. Frib. nom. inval. = Riccia glauca var. ciliaris Warnst. Riccia incrassata Brot. = Oxymitra incrassata (Brot.) Sérgio & Sim-Sim Riccia insularis Levier ex Steph. = Riccia sorocarpa Bisch. Riccia intumescens (Bisch.) Heeg = Riccia ciliata Hoffm. [57] Riccia intumescens Underw. nom. inval. = Riccia crinita Taylor [57] Riccia klinggraeffii Gottsche = Riccia huebeneriana Lindenb. Riccia lamellosa var. americana M. Howe = Riccia lamellosa Raddi Riccia latzelii Schiffn. = Riccia ciliifera Link ex Lindenb. Riccia lescuriana Austin = Riccia beyrichiana Hampe ex Lehm. Riccia lescuriana var. glaucescens (Carrington) Müll. Frib. nom. incorrect. = Riccia beyrichiana Hampe ex Lehm. Riccia lesquereuxii Lindb. = Riccia beyrichiana Hampe ex Lehm. Riccia levieri Schiffn. = Riccia beyrichiana Hampe ex Lehm. Riccia levieri var. algarvica Schiffn. ex Nicholson = Riccia beyrichiana Hampe ex Lehm. Riccia lindenbergiana Saut. = Riccia sorocarpa Bisch. Riccia linkii Lindenb. = Oxymitra incrassata (Brot.) Sérgio & Sim-Sim 420 R.M. Ros et al. Riccia lusitanica Levier ex Steph. = Riccia bicarinata Lindb. Riccia major A. Roth = Riccia glauca L. Riccia mamillata Trab. ex Steph. [54] Riccia michelii var. ciliaris Levier nom. illeg. = Riccia michelii Raddi Riccia michelii var. subinermis Levier nom. illeg. = Riccia michelii Raddi Riccia microspora Steph. = Riccia frostii Austin Riccia minima L. p. p. = Riccia sorocarpa Bisch. Riccia minima L. p. p. = Riccia nigrella DC. Riccia minutissima Steph. = Riccia bifurca Hoffm. [56] Riccia montagnei Steph. = Riccia cavernosa Hoffm. Riccia munita Trab. nom. nud. = Riccia sorocarpa Bisch. [55] Riccia natans L . = Ricciocarpos natans (L.) Corda Riccia negrii Gola = Riccia crystallina L. emend. Raddi Riccia palaestina S. W. Arnell = Riccia frostii Austin Riccia palmata Lindenb. = Riccia michelii Raddi Riccia palmata var. minor Lindenb. = Riccia michelii Raddi Riccia panormitana Levier & Jack ex Lojacono-Pojero = Riccia sorocarpa Bisch. Riccia paradoxa De Not. = Riccia michelii Raddi Riccia pearsoni Steph. = Riccia nigrella DC. Riccia pedemontana Steph. = Riccia ciliifera Link ex Lindenb. Riccia plana Taylor = Riccia crystallina L. emend. Raddi Riccia polycarpa (Trab.) Jelenc [54] Riccia pseudo-frostii (Schiffn.) Müll. Frib. = Riccia huebeneriana Lindenb. Riccia pseudopapillosa Levier ex Steph. = Riccia papillosa Moris Riccia pusilla Warnst. = Riccia bifurca Hoffm. Riccia pyramidata Raddi non Willd.= Oxymitra incrassata (Brot.) Sérgio & Sim-Sim Riccia raddiana J. B. Jack & Levier ex Steph. = Riccia sorocarpa Bisch. Riccia reticulata J. F. Gmel. nom. illeg. = Corsinia coriandrina (Spreng.) Lindb. Riccia riatensis Trab. nom. nud. = Riccia lamellosa Raddi [55] Riccia ruppinensis Steph. = Riccia glauca L. var. glauca Riccia sanguinea Kashyap = Riccia frostii Austin Riccia sorocarpa var. raddiana (J. B. Jack & Levier ex Steph.) Croz. ex Casares-Gil = Riccia sorocarpa Bisch. Riccia sorocarpa f. raddiana (J. B. Jack & Levier ex Steph.) Chevassut & Quézel nom. inval. = Riccia sorocarpa Bisch. Riccia spinosissima Steph. = Riccia ciliata Hoffm. Riccia subciliata Trab. nom. inval. = Riccia glauca var. ciliaris Warnst. Riccia subcrispula Warnst. = Riccia bifurca Hoffm. Riccia subinermis Lindb. = Riccia glauca var. ciliaris Warnst. Riccia subtumida Milde = Riccia ciliata Hoffm. Riccia tellinii Gola = Riccia cavernosa Hoffm. Riccia terracianoi Gola = Riccia cavernosa Hoffm. Riccia trichocarpa M. Howe = Riccia crinita Taylor Riccia trichophylla Gasp. ex Müll. Frib. nom. inval. = Riccia papillosa Moris Riccia tumida Lindenb. = Riccia michelii Raddi Riccia velutina Wilson ex Hook. = Ricciocarpos natans (L.) Corda Riccia venosa A. Roth = Riccia glauca L. var. glauca Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 421 Riccia warnstorfii f. subinermis (Warnst.) Warnst. ex Kavina = Riccia warnstorfii var. subinermis Warnst. Riccia watsonii Austin = Riccia frostii Austin Riccia zachariae Lorb. ex Müll. Frib. = Riccia gougetiana Durieu & Mont. var. gougetiana Ricciella fluitans (L.) A. Braun = Riccia fluitans L. Ricciella fluitans var. canaliculata (Hoffm.) Hartm. = Riccia canaliculata Hoffm. Ricciella fluitans f. terrestris Jaap nom. inval. = Riccia fluitans L. Ricciella heyeri Huebener ex Genth = Riccia canaliculata Hoffm. Ricciella huebeneriana (Lindenb.) Dumort. = Riccia huebeneriana Lindenb. Ricciella huebeneriana var. cavernosa (Hoffm.) Casares-Gil = Riccia cavernosa Hoffm. Ricciella polycarpa Trab. = Riccia polycarpa (Trab.) Jelenc Ricciella rautanenii Steph. = Riccia cavernosa Hoffm. Riccinia perennis (Steph.) Trab. = Riccia perennis Steph. Ricciocarpos velutina (Wilson ex Hook.) Steph. = Ricciocarpos natans (L.) Corda Riella battandieri Trab. = Riella notarisii (Mont.) Mont. Riella clausonis Letourn. = Riella parisii Gottsche Riella cossoniana var. echinata Müll. Frib. nom. inval. = Riella helicophylla (Bory & Mont.) Mont. Riella gallica Balansa = Riella notarisii (Mont.) Mont. Riella reuteri Mont. = Riella notarisii (Mont.) Mont. Riella reuteri f. gallica (Balansa) Trab. = Riella notarisii (Mont.) Mont. Roemeria multifida (L.) Raddi nom. illeg. = Riccardia multifida (L.) Gray Roemeria palmata (Hedw.) Raddi nom. illeg. = Riccardia palmata (Hedw.) Carruth. Ruppinia italica (Sassi) Trevis. = Plagiochasma rupestre (J. R. Forst. & G. Forst.) Steph. Rupinia pyramidata (Raddi) Corda non (Willd.) Corda = Oxymitra incrassata (Brot.) Sérgio & Sim-Sim Rupinia rupestris (J. R. Forst. & G. Forst.) Sw. = Plagiochasma rupestre (J. R. Forst. & G. Forst.) Steph. Saccobasis polita (Nees) H. Buch = Tritomaria polita (Nees) Jörg. Saccogyna graveolens (Schrad.) Lindb. = Geocalyx graveolens (Schrad.) Nees Sarcocyphos aemulus Limpr. = Marsupella condensata (Ångstr. ex C. Hartm.) Kaal. Sarcocyphos alpinus Gottsche ex Husn. = Marsupella alpina (Gottsche ex Husn.) Bernet Sarcocyphos boeckii Austin = Marsupella boeckii (Austin) Lindb. Sarcocyphos capillaris Limpr. = Marsupella boeckii (Austin) Lindb. Sarcocyphos commutatus Limpr. = Marsupella commutata (Limpr.) Bernet Sarcocyphos confertus Limpr. = Marsupella brevissima (Dumort.) Grolle Sarcocyphos densifolius Nees = Marsupella emarginata (Ehrh.) Dumort. ☞ Sarcocyphos emarginatus (Ehrh.) C. Hartm. = Marsupella emarginata (Ehrh.) Dumort. Sarcocyphos ehrhardtii Corda nom. illeg. = Marsupella emarginata (Ehrh.) Dumort. Sarcocyphos ehrhardtii var. aquaticus (Nees) Gottsche et al. = Marsupella emarginata (Ehrh.) Dumort. Sarcocyphos ehrhardtii var. robustus De Not. = Marsupella emarginata (Ehrh.) Dumort. Sarcocyphos funckii (F. Weber & D. Mohr) Nees = Marsupella funckii (F. Weber & D. Mohr) Dumort. Sarcocyphos muelleri Nees = Marsupella funckii (F. Weber & D. Mohr) Dumort. Sarcocyphos neglectus Limpr. = Marsupella sprucei (Limpr.) Bernet Sarcocyphos pygmaeus Limpr. = Marsupella funckii (F. Weber & D. Mohr) Dumort. Sarcocyphos revolutus Nees = Apomarsupella revoluta (Nees) R. M. Schust. 422 R.M. Ros et al. Sarcocyphos sparsifolius Lindb. = Marsupella sparsifolia (Lindb.) Dumort. Sarcocyphos sphacelatus (Gieseke ex Lindenb.) Nees = Marsupella sphacelata (Gieseke ex Lindenb.) Dumort. Sarcocyphos sprucei Limpr. = Marsupella sprucei (Limpr.) Bernet Sarcocyphos styriacus Limpr. = Marsupella sparsifolia (Lindb.) Dumort. Sarcocyphos sullivantii De Not. = Marsupella sphacelata (Gieseke ex Lindenb.) Dumort. Sarcocyphos ustulatus auct. = Marsupella sprucei (Limpr.) Bernet Sauteria grandis Lindb. = Peltolepis quadrata (Saut.) Müll. Frib. Sauteria hyalina (Sommerf.) Lindb. = Athalamia hyalina (Sommerf.) S. Hatt. Sauteria quadrata Saut. = Peltolepis quadrata (Saut.) Müll. Frib. Sauteria seriata Lindb. nom. illeg. = Athalamia hyalina (Sommerf.) S. Hatt. Scapania aconiensis De Not. = Scapania nemorea (L.) Grolle Scapania bartlingii (Hampe ex Nees) Nees nom. illeg. = Scapania cuspiduligera (Nees) Müll. Frib. Scapania biroliana C. Massal. & Carestia = Scapania compacta (A. Roth) Dumort. Scapania buchii Müll. Frib. nom. illeg. = Scapania lingulata H. Buch Scapania carestiae De Not. = Scapania cuspiduligera (Nees) Müll. Frib. Scapania carinthiaca var. massalongi Müll. Frib. = Scapania carinthiaca J. B. Jack ex Lindb. Scapania casaresana Steph. = Scapania gracilis Lindb. Scapania compacta var. biroliana (C. Massal. & Carestia) C. Massal. = Scapania compacta (A. Roth) Dumort. Scapania convexa (Scop.) Pearson = Scapania umbrosa (Schrad.) Dumort. Scapania curta var. geniculata (C. Massal.) Müll. Frib. = Scapania helvetica Gottsche ☞ Scapania curta var. spinulosa (Nees) Gottsche et al. = Scapania carinthiaca J. B. Jack ex Lindb. Scapania dentata (Dumort.) Dumort. = Scapania undulata (L.) Dumort. Scapania franzoniana De Not. = Scapania subalpina (Nees ex Lindenb.) Dumort. Scapania geniculata C. Massal. = Scapania helvetica Gottsche ☞ Scapania gracilis var. laxifolia (Dumort.) Dumort. = Scapania gracilis Lindb. Scapania gracilis var. minor (Pearson) Macvicar = Scapania gracilis Lindb. Scapania intermedia (Husn.) Pearson = Scapania undulata (L.) Dumort. Scapania joergensenii Schiffn. ex Müll. Frib. = Scapania nemorea (L.) Grolle Scapania massalongii (Müll. Frib.) Müll. Frib. = Scapania carinthiaca J. B. Jack ex Lindb. Scapania microphylla Warnst. = Scapania lingulata H. Buch Scapania microscopica Culm. = Diplophyllum obtusifolium (Hook.) Dumort. Scapania mucronata var. praetervisa (Meyl.) H. Buch = Scapania praetervisa Meyl. Scapania nemorea subsp. crassiretis (Bryhn) Potemkin = Scapania crassiretis Bryhn Scapania nemorosa Dumort. nom. illeg. = Scapania nemorea (L.) Grolle Scapania nemorosa var. aconiensis (De Not.) C. Massal. = Scapania nemorea (L.) Grolle Scapania nemorosa var. fallaciosa Schiffn. = Scapania nemorea (L.) Grolle Scapania oakesii Austin = Scapania undulata (L.) Dumort. Scapania obliqua (Arnell) Schiffn. = Scapania uliginosa (Sw. ex Lindenb.) Dumort. Scapania recurva Steph. ex Müll. Frib. = Scapania gracilis Lindb. Scapania resupinata (L.) Dumort. = Scapania undulata (L.) Dumort. Scapania resupinata sensu A. L. M. Guim. = Scapania gracilis Lindb. Scapania rosacea (Corda) Nees nom. inval. = Scapania curta (Mart.) Dumort. Scapania rotundata Warnst. = Scapania paludicola Loeske & Müll. Frib. Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 423 Scapania subalpina var. undulifolia (Nees) Gottsche et al. = Scapania subalpina (Nees ex Lindenb.) Dumort. Scapania tyrolensis (Nees) Nees = Scapania aequiloba (Schwägr.) Dumort. Scapania undulata var. aequatiformis (De Not.) C. Massal. & Carestia = Scapania undulata (L.) Dumort. Scapania undulata var. dentata (Dumort.) Douin = Scapania undulata (L.) Dumort. Scapania undulata var. dentata (Dumort.) McArdle. nom. inval. = Scapania undulata (L.) Dumort. Scapania undulata var. integrifolia Lindb. = Scapania undulata (L.) Dumort. Scapania undulata var. paludosa Müll. Frib. = Scapania paludosa (Müll. Frib.) Müll. Frib. Scapania undulata var. purpurascens (Huebener) Cogn. = Scapania undulata (L.) Dumort. Scapania undulata var. purpurea Nees nom. inval. = Scapania undulata (L.) Dumort. Scapania vexata C. Massal. nom. illeg. = Scapania scapanioides (C. Massal.) Grolle Scapaniella carinthiaca (J. B. Jack ex Lindb.) H. Buch = Scapania carinthiaca J. B. Jack ex Lindb. Scapaniella massalongi (Müll. Frib.) H. Buch = Scapania carinthiaca J. B. Jack ex Lindb. Schistochilopsis grandiretis (Lindb. ex Kaal.) Konstantinova = Lophozia grandiretis (Lindb. ex Kaal.) Schiffn. Schistochilopsis incisa (Schrad.) Konstantinova = Lophozia incisa (Schrad.) Dumort. Schistochilopsis opacifolia (Culm. ex Meyl.) Konstantinova = Lophozia opacifolia Culm. ex Meyl. Solenostoma amplexicaule (Dumort.) Steph. = Jungermannia sphaerocarpa Hook. Solenostoma atrovirens (Dumort.) Müll. Frib. nom. illeg. = Jungermannia atrovirens Dumort. Solenostoma caespiticium (Lindenb.) Steph. = Jungermannia caespiticia Lindenb. Solenostoma confertissimum (Nees) Schljakov = Jungermannia confertissima Nees Solenostoma cordifolium (Dumort.) Steph. = Jungermannia exsertifolia subsp. cordifolia (Dumort.) Vá≈a Solenostoma crenulatum (Sm.) R. M. Schust. = Jungermannia gracillima Sm. Solenostoma crenulatum (Sm.) Mitt. nom. illeg. = Jungermannia gracillima Sm. Solenostoma handelii (Schiffn.) Müll. Frib. = Jungermannia handelii (Schiffn.) Amakawa Solenostoma hyalinum (Lyell) Mitt. = Jungermannia hyalina Lyell Solenostoma lanceolatum (L.) Steph. = Jungermannia atrovirens Dumort. Solenostoma lanceolatum auct. non (L.) Steph. = Jungermannia leiantha Grolle Solenostoma levieri (Steph.) Steph. = Jungermannia confertissima Nees Solenostoma oblongifolium (Müll. Frib.) Müll. Frib. = Jungermannia pumila With. Solenostoma obovatum (Nees) C. Massal. = Jungermannia obovata Nees Solenostoma ontariense R. M. Schust. = Jungermannia hyalina Lyell Solenostoma polare (Lindb.) R. M. Schust. = Jungermannia polaris Lindb. Solenostoma pumilum (With.) Müll. Frib. = Jungermannia pumila With. Solenostoma pusillum (C. E. O. Jensen) Steph. = Jungermannia jenseniana Grolle Solenostoma pusillum var. vinaceum R. M. Schust. = Jungermannia confertissima Nees Solenostoma scalare (Gray) Mitt. = Nardia scalaris Gray Solenostoma schiffneri (Loitl.) Müll. Frib. = Jungermannia polaris Lindb. Solenostoma sphaerocarpoideum (De Not.) Paton & E. F. Warb. = Jungermannia atrovirens Dumort. Solenostoma sphaerocarpum (Hook.) Steph. = Jungermannia sphaerocarpa Hook. Solenostoma subellipticum (Lindb. ex Kaal.) R.M. Schust. = Jungermannia subelliptica (Lindb. ex Kaal.) Levier 424 R.M. Ros et al. Solenostoma triste (Nees) Müll. Frib. = Jungermannia atrovirens Dumort. Southbya alicularia C. Massal. = Southbya tophacea (Spruce) Spruce Southbya crenulata (Mitt.) Bernet = Jungermannia gracillima Sm. Southbya fennica (Gottsche) C. Hartm. = Arnellia fennica (Gottsche) Lindb. Southbya hyalina (Lyell) Husn. = Jungermannia hyalina Lyell Southbya obovata (Nees) Lindb. = Jungermannia obovata Nees Southbya stillicidiorum (Raddi) Lindb. = Southbya tophacea (Spruce) Spruce Sphaerocarpos algeriensis Trab. nom. nud. = Sphaerocarpos texanus Austin [64] Sphaerocarpos californicus Austin = Sphaerocarpos texanus Austin Sphaerocarpos europaeus Lorb. = Sphaerocarpos texanus Austin Sphaerocarpos notarisii Mont. = Riella notarisii (Mont.) Mont. Sphaerocarpos sphaerocarpos (Dicks.) M. Howe nom. illeg. = Sphaerocarpos michelii Bellardi Sphaerocarpos terrestris Sm. nom. illeg. = Sphaerocarpos michelii Bellardi Sphagnoecetis communis Nees nom. illeg. = Odontoschisma sphagni (Dicks.) Dumort. Sphagnoecetis macounii Austin = Odontoschisma macounii (Austin) Underw. Sphenolobus ascendens Warnst. = Lophozia ascendens (Warnst.) R.M. Schust. Sphenolobus exsectiformis (Breidl.) Steph. = Tritomaria exsectiformis (Breidl.) Loeske Sphenolobus exsectiformis var. aequilobus (Culm.) Müll. Frib. = Tritomaria scitula (Taylor) Jörg. Sphenolobus exsectus (Schmidel ex Schrad.) Steph. = Tritomaria exsecta (Schmidel ex Schrad.) Loeske Sphenolobus hellerianus (Nees ex Lindenb.) Steph. = Anastrophyllum hellerianum (Nees ex Lindenb.) R. M. Schust. Sphenolobus michauxii (F. Weber) Steph. = Anastrophyllum michauxii (F. Weber) H. Buch Sphenolobus minutus (Schreb.) Berggr. = Anastrophyllum minutum (Schreb.) R. M. Schust. Sphenolobus politus (Nees) Steph. = Tritomaria polita (Nees) Jörg. Sphenolobus saxicolus (Schrad.) Steph. = Anastrophyllum saxicola (Schrad.) R. M. Schust. Sphenolobus scitulus (Taylor) Steph. = Tritomaria scitula (Taylor) Jörg. Staurophora pulchella Willd. nom. illeg. = Lunularia cruciata (L.) Lindb. Stephanina complanata (L.) Kuntze = Radula complanata (L.) Dumort. Strozzia conica (L.) Müll. Frib. = Conocephalum conicum (L.) Dumort. Strozzia hemisphaerica (L.) Müll. Frib. = Reboulia hemisphaerica (L.) Raddi Strozzius conicus (L.) Gray = Conocephalum conicum (L.) Dumort. Strozzius hemisphaericus (L.) Gray = Reboulia hemisphaerica (L.) Raddi Targionia bifurca Nees & Mont. = Targionia hypophylla L. Targionia convoluta Lindenb. & Gottsche = Targionia hypophylla L. Targionia germanica Corda nom. inval. = Targionia hypophylla L. Targionia hypophylla var. fimbriata (Müll. Frib.) Müll. Frib. = Targionia hypophylla L. Targionia hypophylla var. lorbeeriana (Müll. Frib.) Aleffi & Schumacker nom. inval. = Targionia lorbeeriana Müll. Frib. Targionia michelii Corda = Targionia hypophylla L. Targionia sphaerocarpos Dicks. = Sphaerocarpos michelii Bellardi Telaranea nematodes sensu auct. eur. non (Gottsche ex Austin) M. Howe = Telaranea europaea J.J. Engel & G.L.S. Merril [66] Telaranea sejuncta auct. eur. non (Ångstr.) S. W. Arnell = Telaranea europaea J.J. Engel & G.L.S. Merril [67] Telaranea setacea auct. non (Weber) Müll. Frib. = Kurzia pauciflora (Dicks.) Grolle Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 425 Telaranea sylvatica (A. Evans) Müll. Frib. = Kurzia sylvatica (A. Evans) Grolle Telaranea trichoclados (Müll. Frib.) Müll. Frib. = Kurzia trichoclados (Müll. Frib.) Grolle Temnoma setiforme (Ehrh.) M. Howe = Tetralophozia setiformis (Ehrh.) Schljakov Tesselina coriandrina (Spreng.) Dumort. = Corsinia coriandrina (Spreng.) Lindb. ☞ Tesselina paleacea auct. = Oxymitra incrassata (Brot.) Sérgio & Sim-Sim Tesselina pyramidata (Raddi) Dumort. non (Willd.) Dumort. = Oxymitra incrassata (Brot.) Sérgio & Sim-Sim Tesselina pyramidata var. paleacea Lindb. = Oxymitra incrassata (Brot.) Sérgio & Sim-Sim Tesselina pyramidata var. paleacea (Bisch. ex Lindenb.) Crozals nom. illeg. = Oxymitra incrassata (Brot.) Sérgio & Sim-Sim Tesselina pyramidata var. polycarpa (Bisch. ex Lindenb.) Lindb. = Oxymitra incrassata (Brot.) Sérgio & Sim-Sim Thedenia blyttii (Moerch ex Hornem.) Hartm. nom. inval. = Moerckia blyttii (Moerch ex Hornem.) Brockm. ☞ Trigonanthus curvifolius (Dicks.) Spruce = Nowellia curvifolia (Dicks.) Mitt. Trigonanthus francisci (Hook.) Hartm. = Cladopodiella francisci (Hook.) Jörg. Tritomaria polymorpha (R. M. Schust.) Grolle = Tritomaria polita (Nees) Jörg. Tritomaria scitula var. savoica Müll. Frib. = Tritomaria scitula (Taylor) Jörg. Annotations 1. Bischler (2004) reported several taxa from Mediterranean countries without details of locality. The following require verification as they are the only known reports for their respective countries. – For Algeria: Calypogeia muelleriana (Schiffn.) Müll. Frib., Cephalozia connivens (Dicks.) Lindb., C. lunulifolia (Dumort.) Dumort., Diplophyllum albicans (L.) Dumort., Dumortiera hirsuta (Sw.) Nees, Jungermannia obovata Nees, J. pumila With., Leiocolea collaris (Nees) Schljakov, Lophocolea heterophylla (Schrad.) Dumort., L. minor Nees, Marsupella emarginata (Ehrh.) Dumort., Riccardia latifrons (Lindb.) Lindb., Riccia ligula Steph. and Scapania gracilis Lindb. – For Lebanon: Anthoceros punctatus L., Athalamia hyalina (Sommerf.) S. Hatt., Barbilophozia barbata (Schmidel ex Schreb.) Loeske, Bazzania flaccida (Dumort.) Grolle, Blepharostoma trichophyllum (L.) Dumort., Cephaloziella calyculata (Durieu & Mont.) Müll. Frib., Cololejeunea minutissima (Sm.) Schiffn., Fossombronia angulosa (Dicks.) Raddi, F. caespitiformis subsp. multispira (Schiffn.) J. R. Bray & D. C. Cargill (as F. husnotii Corb.), Jungermannia leiantha Grolle, Lophocolea bidentata (L.) Dumort., Nardia compressa (Hook.) Gray, Pellia epiphylla (L.) Corda, Plagiochila porelloides (Torrey ex Nees) Lindenb., Porella arboris-vitae (With.) Grolle, P. platyphylla (L.) Pfeiff., Radula complanata (L.) Dumort., Riccia bifurca Hoffm., R. crystallina L. emend. Raddi, R. michelii Raddi, Scapania aequiloba (Schwägr.) Dumort. and S. aspera Bernet & M. Bernet. – For Morocco: Riccia huebeneriana Lindenb., Riella cossoniana Trab. and R. parisii Gottsche. – For Tunisia: Riccia warnstorfii Lindenb. and Sphaerocarpos texanus Austin. – For Turkey: Plagiochila bifaria (Sw.) Lindenb. – For Europe, see Söderström et al. (2007). 2. The nomenclatural problems of Anthoceros agrestis Paton and its relationship with A. punctatus L. have been discussed by Söderström et al. (2002). In the 426 R.M. Ros et al. Mediterranean area the two species have been regularly confused, both on taxonomic and nomenclatural basis, but most records probably belong to A. punctatus. Further studies of Mediterranean material are needed (including A. multifidus L.) to clarify the distribution of the two taxa. 3. Phaeoceros carolinianus (Michx.) Prosk. was reported from Crete by Jovet-Ast & Bischler (1968). This is a doubtful record that is rejected here. Jovet-Ast & Bischler include Crete in the distribution of “P. laevis subsp. carolinianus”, citing the paper by Düll (1966, p. 743). However in this paper Düll does not distinguish subsp. carolinianus and subsp. laevis. He lists only P. laevis. 4. Anthoceros beltrani Casares-Gil was described by Casares-Gil (1915) from Spain, and later synonymized by Proskauer (1958) with Phaeoceros laevis (L.) Prosk. Nevertheless this synonymy is questionable because of the paroicous condition of A. beltrani. Further studies are needed. 5. Anthoceros dichotomus var. gussonei Zodda was synonymized with Phaeoceros laevis (L.) Prosk. by Proskauer (1958), although it often has been erroneously interpreted in the Mediterranean area as a synonym of Phymatoceros bulbiculosus (Brot.) Stotler, W. T. Doyle & Crand.-Stotl. 6. For the taxonomy of the genus Phymathoceros, we have followed Stotler et al. (2005) and Crandall-Stotler et al. (2006). 7. According to Schuster (1988), Anthelia nivalis (F. Weber & D. Mohr) Lindb. is a nomen dubium and could correspond to Anthelia juratzkana (Limpr.) Trevis. p. p. and A. julacea (L.) Durmort. p. p. (type probably A. julacea). 8. The presence of the American Asterella elegans (Spreng.) Trevis. was rejected from Corsica by Söderström et al. (2002) and the report from Turkey (Bornmüller, 1931) must also be rejected. 9. There has been some confusion in the literature with the name Fimbriaria nana Lindenb., that has been considered by some authors to be a synonym of Asterella gracilis (F. Weber) Underw., but Grolle (1980) clearly established its identity with Athalamia hyalina (Sommerf.) S. Hatt. 10. Fimbriaria fragrans (Balbis) Nees 1825 (≡ Marchantia fragrans Balbis) is Mannia fragrans (Balbis) Frye & L. Clark while F. fragrans (Schleich.) Nees 1838 (≡ Marchantia fragrans Schleich.) is a synonym to Asterella saccata (Wahlenb.) A. Evans. Both variants of F. fragrans have been used in the study area. It is likely that most of the Mediterranean reports correspond to Mannia fragrans rather than Asterella saccata, but further revision is needed. 11. The problems in applying the name Calypogeia trichomanis (L.) Corda are discussed by Söderström et al. (2002), who note that this name has been variously interpreted in Europe as both C. azurea Stotler & Crotz (Grolle & Long, 2000) and C. muelleriana (Schiffn.) Müll. Frib. The same is true of the Mediterranean area. 12. Jungermannia scalaris Schmidel ex Schreb. is a nomen dubium and could correspond either to Calypogeia trichomanis (L.) Corda nom. rejic. (currently equated by most authors with Calypogeia azurea Stotler & Crotz), or to Odontoschisma sp. (Vá≈a, 1976). 13. Cephalozia macrostachya Kaal. was erroneously included in the checklist of Spain by Casas (1998). Therefore it should be excluded from Spain. Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 427 14. Cephaloziella gallica (Douin) Douin was considered by Müller (1954-1957) to be a synonym of Cephaloziella calyculata (Durieu & Mont.) Müll. Frib., but it has been synonymized with C. hampeana (Nees) Schiffn. by recent authors (Schumacker & Vá≈a, 2000, 2005; Söderström et al., 2002) and that is the criterion followed here. 15. Cephaloziella myriantha (Lindb.) Schiffn. was considered by Schuster (1980) to be a synonym of C. rubella var. elegans (Heeg) R. M. Schust. (= C. elegans (Heeg) Schiffn.), but Damsholt (2002) referred it to C. rubella (Nees) Warnst. var. rubella, which is the interpretation followed here. 16. Chiloscyphus pallescens (Ehrh. ex Hoffm.) Dumort. and Porella obtusata (Taylor) Trevis., have been reported from localities formerly included in the Albanian territory, but now belong to Montenegro. 17. Conocephalum salebrosum Szweyk., Buczkowska & Odrzykoski has been recently described by Szweykowski et al. (2005) as a separated species from C. conicum (L.) Dumort. The new species is in some areas of Europe more common than C. conicum and it is impossible to know which species is meant by the latter name prior to 2005. Conocephalum salebrosum will certainly turn out to be more widespread in the Mediterranean area. 18. Recent studies on the genus Fossombronia Raddi by Stotler & Crandall-Stotler (2006) propose a new treatment for several taxa, and this treatment has been followed here. 19. Fossombronia intestinalis Taylor was erroneously reported from Algeria by Söderström et al. (2002). It is an Australian taxon that is also known from South Africa and it should be rejected from the Mediterranean area. 20. According to Scott & Pike (1988), Fossombronia intermedia Corb. ex Trab. nom. inval. (= Fossombronia caespitiformis var. intermedia Corb. ex Trab. nom. inval.) could be ascribed either to F. caespitiformis De Not. ex Rabenh. or to F. wondraczekii (Corda) Lindb. However Stotler & Crandall-Stotler (2006) consider it to be a synonym of F. echinata Macvicar. At present it is not possible to know the identity of the Mediterranean specimens so named. 21. Jungermannia crispula Brot. was described from Portuguese material and was later transferred to Fossombronia as F. crispula (Brot.) R. M. Schust. by Schuster (1992a), but no other report of this species has been found in European works. Machado (1925) included J. crispula in Fossombronia pusilla (L.) Nees p. p. and F. caespitiformis p. p., but according to Sérgio & Carvalho (2003), no original material of J. crispula has been found. Therefore it should be considered a nomen dubium. 22. Fossombronia crispa is problematic and should be considered doubtful for Europe (cf. Söderström et al., 2007). 23. The identity of Fossombronia loitlesbergeri Schiffn. is controversial. Paton (1999) and Scott & Pike (1988) considered it to be related to Fossombronia wondraczekii (Corda) Lindb., but Stotler & Crandall-Stotler (2006), whom we follow here, treated it as a synonym of F. pusilla (L.) Nees. 24. Frullania muscicola Steph. is an Asian species that was reported from Macaronesia by Sérgio (1985). Later Bisang et al. (1989) considered that the Asian and Macaronesian specimens would be better considered as Frullania cesatiana var. muscicola Bisang et al., and Sim-Sim et al. (1995) assigned the Macaronesian specimens to the endemic F. azorica Sim-Sim et al. 428 R.M. Ros et al. The reports from the Canary islands of Frullania riparia Hampe ex Lehm. by Arnell (1961) also correspond to this taxon (Casas et al., 1989). 25. Several infraspecific taxa have been recognized in Frullania dilatata (L.) Dumort. and F. tamarisci (L.) Dumort. for Portugal and Madeira by Sim-Sim (1999). In addition Losada Lima & Vanden Berghen (1985) described Frullania dilatata var. elongata from the Canary Islands, though it was subsequently synonymized with F. ericoides (Nees) Mont. by Bisang et al. (1989). However these territories form only a relatively small part of the Mediterranean Basin, and the criteria used by Sim-Sim (1989) are difficult to apply to the whole region. We have therefore adopted a conservative treatment in this checklist. 26. Grolle & Long (2000) referred Frullania davurica Hampe to F. jackii Gottsche. The Mediterranean material identified as F. davurica probably corresponds to F. jackii, which is also the interpretation of Schumacker & Vá≈a (2005). Further confirmation is needed. 27. Jubula pennsylvanica (Steph.) A. Evans was reported doubtfully from Turkey by Schiffner (1909) sub Jubula hutchinsiae var. sullivanti Spruce. According to Grolle & Long (2000), it could be the same taxon as Jubula hutchinsiae subsp. javanica (Steph.) Verd. The relationship between the two taxa needs to be assessed. Two varieties have also been recognized in Macaronesia by Sim-Sim et al. (2002): Jubula hutchinsiae subsp. hutchinsiae var. hutchinsiae and Jubula hutchinsiae subsp. hutchinsiae var. integrifolia Lindb. However the correct author citation of the latter name is Jubula hutchinsiae subsp. hutchinsiae var. integrifolia (Steph.) S. Hatt., and it has previously been synonymized with subsp. javanica (see Söderström et al., 2007). 28. Jungermannia compacta Mazziari nom. nud. non Roth., J. foscardiana Mazziari nom. nud., J. guilfortiana Mazziari nom. nud., J. lacera Mazziari nom. nud., and J. limosa Mazziari nom. nud. were reported from Greece by Mazziari (1851) but no other reference to these names has been found in the literature. 29. Jungermannia jenseniana Grolle was reported from Spain by Lloret et al. (1997), but after revision of the specimen by M. Infante, it was assigned to Nardia scalaris Gray. Therefore the species is at the moment rejected from the area. 30. Nardia hyalina var. colorata Samp. is a nomen nudum and is based on a separate type (from Portugal) than Nardia hyalina var. colorata (Mart.) Dalla Torre & Sarnth. (type from Germany). Vá≈a (1975) treated the latter as a synonym of Jungermannia gracillima. Later Sérgio & Carvalho (2003) stated that “Nardia hyalina var. colorata Samp. = ?? Jungermannia hyalina Lyell”. Consequently the identity of the name remains uncertain and requires further studies. 31. Leiocolea collaris (Nees) Schljakov is sometimes treated as a synonym of L. bantriensis (Hook.) Jørg. Thus, the literature reports of the latter must be treated with caution. 32. Leiocolea turbinata f. aquatica Jelenc nom. nud. was reported by Jelenc (1955) from Morocco but no other record or reference to it has been found. 33. According to Schuster (1980), Jungermannia serpyllifolia Dicks. nom. illeg. corresponds to Lejeunea patens Lindb. but the name Lejeunea serpyllifolia sensu Lib. nom. illeg. refers to another taxon, which led Bonner & Miller (1960) to propose the new name Lejeunea libertiae Bonner & H. A. Miller. This was later synonymized with L. cavifolia (Ehrh.) Lindb. The literature reports of L. serpyllifolia are difficult to place but do mostly correspond to L. cavifolia. Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 429 34. Schuster (1980) confirmed the presence of L. flava subsp. moorei (Lindb.) R.M. Schust. in Ireland. In Macaronesia it has also been reported from Madeira. According to Grolle & Long (2000), the identity of the European and Macaronesian specimens referred to L. flava remains to be confirmed. In this paper they have been included in Lejeunea flava (Sw.) Nees. 35. Lepidozia pearsonii Spruce was reported by Reinoso (1985) from Spain and considered by Casas (1998) as a species whose occurrence in Spain needed confirmation, which led to Söderström et al. (2002) to exclude it from Spain. After that, the specimen has been revised by R. Schumacker as Lepidozia reptans (L.) Dumort. Therefore the species should be rejected from the Mediterranean area. 36. The report of Lophozia latifolia R. M. Schust. (sub L. propagulifera (Gottsche) Steph.) from Serbia given in Sabovljevi† & Natcheva (2006) is almost certainly erroneous. This is an arctic taxon that has been synonymized with Lophozia jurensis Müll. Frib. (whose type is from the Alps), but this synonymy is very questionable and “true” L. propagulifera has never been reported from central Europe (see Söderström et al., 2007). 37. Recent molecular studies (Roo et al., 2007) show that Lophozia silvicola H. Buch is a species well separated from L. ventricosa (Dicks.) Dumort. 38. Lophozia confertifolia Schiffn. is not conspecific with true L. wenzelii (Nees) Steph. as stated by Müller (1954-1957), and we have followed Schuster (1969) and Schumacker & Vá≈a (2005) who consider Lophozia confertifolia to be a synonym of Lophozia ventricosa (Dicks.) Dumort (see Söderström et al., 2007). 39. Grimaldia atlantica Trab. nom. inval. (Mannia atlantica Jelenc nom. inval.) was reported by Trabut (1941) from Algeria, but no other record or reference to it have been found. 40. According to Grolle & So (2003), Metzgeria violacea (Ach.) Dumort. is the correct name for M. fruticulosa auct. eur. 41. The identity of the plants named Nardia silvrettae (Gottsche ex Dumort.) Trevis. is doubtful since it has been synonymized with both N. geoscyphus (De Not.) Lindb. (Vá≈a, 1976) and N. insecta Lindb. (Schumacker & Vá≈a, 2005). According to Geissler & Bischler (1989), its basionym is Marsupella silvrettae Gott. ex Dumort., 1874 (≡ Jungermannia silvrettae Gottsche & Rabenh. nom. inval.). Schumacker & Vá≈a (2000, 2005) refer it to Nardia insecta Lindb., but cite it with a different authority – Nardia silvrettae Douin. 42. Pellia fabbroniana f. undulata Jelenc nom. inval. was reported by Jelenc (1955) from Algeria but no other record or reference to it has been found. 43. Jungermannia fuciformis Bory (= Pellia fuciformis (Bory) Nees) is an uncertain European name reported from Greece by Bory de Saint-Vicent (1832). Its identity is still to be assessed but Coppey (1908) stated that it could be a sterile form of Pellia epiphylla (L.) Corda. 44. Plagiochila africana Steph. was erroneously reported from Libya by Söderström et al. (2002). It is a west African tropical species that should be rejected from the Mediterranean area. 45. The nomenclatural history of Plagiochila asplenioides (L. emend. Taylor) Dumort. is complex and it is not always possible to know whether a particular report belongs to P. asplenioides s. str. or P. porelloides (Nees) Lindenb. Most earlier reports probably belong to the latter (cf. Söderström et al., 2002). 430 R.M. Ros et al. 46. Plagiochila atlantica Lindenb. ex Lehm. has been synonymized with P. heterophylla Lindenb. ex Lehm by Heinrichs (2002). 47. Plagiochila maderensis Gottsche ex Steph. has recently been reinstated for Madeira and the Canary Islands by Rycroft et al. (2004). 48. Fontinha (2005) excluded Porella arboris-vitae (With.) Grolle and Porella platyphylla (L.) Pfeiff. from Madeira. Bischler et al. (2006) stated that the first species is restricted to southern continental areas while the second has a “European and Asiatic (North American?)” distribution. Further studies are needed to establish conclusively their presence in the Canary Islands, in view of the ecological differences that characterize the individual islands. 49. Madotheca canariensis (F. Weber) Nees has been reported from Northern Africa by Trabut (1941) and others without mention of locality or country and Bischler (2004) placed it in Morocco. Trabut used this name for specimens intermediate between Madotheca laevigata (Schrad.) Dumort. and Madotheca thuja (Dicks.) Dumort. The two latter names are synonyms of Porella arboris-vitae (With.) Grolle, but M. thuja was earlier frequently used for P. obtusata (Taylor) Trevis. The identity of the North African plants is therefore questionable and Porella canariensis (F. Weber) Underw. should be rejected until it is positively verified. 50. Ros et al. (1999) erroneously reported Porella cordaeana (Huebener) Moore from Libya based on a supposed record of Rungby (1962). But this author reported this species only for Morocco in Northern Africa. In addition, Porella pinnata L. was reported erroneously by Ros et al. (1999) from Morocco and should be rejected from this country. 51. In several works Achiton quadratum (Scop.) Corda has been erroneously assigned to Reboulia hemisphaerica (L.) Raddi. According to Bonner (1962b) the basionym of this name is Marchantia quadrata Scop., which actually is the basionym of Preissia quadrata (Scop.) Nees. 52. Düll (1983) included the name Radula physoloba auct. as a synonym of R. aquilegia (Hook. f. & Taylor) Gottsche et al. Later Sérgio & Carvalho (2003) excluded this species from Portugal and considered it to be a synonym of Radula lindenbergiana Gottsche ex C. Hartm. In both cases the use of this name seems to be erroneous, as Radula physoloba Mont. is a species of New Zealand. Thus, it should be excluded from the Mediterranean lists. 53. For the taxonomy of the genus Riccia L., the monographic treatment by JovetAst (1986) has been followed here. 54. Several species that have been reported from Northern Africa were not included in the monograph of the genus Riccia L. in the Mediterranean area by Jovet-Ast (1986), but mentioned by Söderström et al. (2002), and their identity remains doubtful. – Riccia chinensis Herz. was proposed as new to Algeria by Hariot (1913) as Riccia chinensis Steph. ex Hariot from Tahount Arak (Mouydir), with the observation that the specific name given by Stephani was provisional because it seemed to be unsuitable for an African species. Moreover it lacked a Latin diagnosis and was therefore an invalid name. Herzog subsequently validated the name but using material from China as type. The identity of the Algerian material is therefore doubtful. Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 431 – Riccia chudoana Steph. was reported from Sahara sine loco by Jelenc (1955) but no other record or reference to it has been found. It is certainly a nom. nov. for R. chinensis Herz. as it is typified by the same specimen. – Riccia convexa Steph. was invalidly published by Hariot (1913) on the basis of material from Dayet El-Tofla (Algeria). Later Stephani (1912-1917) validated the species but selecting material collected in China, transmitted by Hariot, as the type. The identity of the Algerian material is therefore doubtful. – Riccia mamillata Trab. ex Steph. was reported from Algeria by Trabut (1941) but according to Casares-Gil (1919) it is similar to Riccia macrocarpa Levier. – Riccia polycarpa (Trab.) Jelenc was reported from Algeria by Jelenc (1955). No other record or reference to it has been found. 55. Invalidly published names in the genus Riccia that have been used for material from Northern Africa are: – Riccia aculeata Trab. nom. nud., reported by Trabut (1941) from Algeria. Grolle (1976) indicated that, according to Jovet-Ast in litt., it is probably identical with Riccia crozalsii Levier. – Riccia battandieri Trab. nom. nud., reported by Trabut (1941) from Algeria. Grolle (1976) indicated that, according to Jovet-Ast in litt., it is a nomen dubium. – Riccia bischoffii f. armata Trab. nom. nud., reported by Trabut (1941) from North Africa without indication of country. Jovet-Ast (1983) stated that this name corresponds, according to the description, with Riccia gougetiana var. erinacea Schiffn. nom. inval. (= var. armatissima Schiffn.). – Riccia echinulata Trab. nom. nud., reported by Trabut (1941) from Algeria. Grolle (1976) indicated that, according to Jovet-Ast in litt., it is a nomen dubium (it probably corresponds to R. ciliifera Link ex Lindenb.). – Riccia munita Trab. nom. nud., reported by Trabut (1941) from Algeria. Grolle (1976) indicated that, according to Jovet-Ast in litt., it is a nomen dubium (it probably corresponds to R. sorocarpa Bisch.). – Riccia riatensis Trab. nom. nud., reported by Trabut (1941) from Morocco. Grolle (1976) indicated that, according to Jovet-Ast in litt., it is a nomen dubium (it probably corresponds to R. lamellosa Raddi). 56. In the original description of Riccia minutissima Steph., the author cited two specimens, which Schuster (1992b) considered to belong to two different species (Riccia bifurca Hoffm. and R. glauca L.). Later Jovet-Ast (1986) stated that the correct synonymy of Riccia minutissima is with R. bifurca, and consequently the second specimen cited in the protologue was misidentified. 57. Riccia ciliata var. intumescens Bisch. was combined as Riccia intumescens by Heeg in 1894 but also Underwood in 1884 had used this name invalidly. According to Schuster (1992b), R. intumescens (Bisch.) Heeg is R. ciliata, whereas R. intumescens Underw. nom. inval. is R. ciliata subsp. trichocarpa (= R. crinita Taylor). Until the Mediterranean material identified as Riccia intumescens is re-examined, it is impossible to know the species to which the individual records refer. 58. Riccia frostii var. crystallinoides Schiffn. was reported from Syria by Schiffner (1913), but no other record of or reference to it has been found. 59. Riella affinis M. Howe & Underw. was erroneously included in Casas (1998). The only Spanish reports of this species are from the Canary Islands, and it must be excluded from mainland Spain. 432 R.M. Ros et al. 60. Despite the statements by Grolle & Long (2000) and Geissler & Bischler (1990), Riella numidica Trab. and Riella sersuensis Trab. were validly described, since the protologue of both species includes a Latin diagnosis (in agreement with the article 36.1 of the Missouri ICBN). 61. Scapania brevicaulis Taylor has been reported from Italy by Potemkin (2002), and this is the only report for the Mediterranean area. Because the author did not give any indication of the locality, the report should be considered doubtful until specimens can be checked. 62. Scapania calcicola (Arnell & J. Perss.) Ingham was erroneously reported from the Balearic Islands by Casas et al. (2004) without giving any locality, and should be rejected. 63. Jungermannia cordifolia Brot. non Hook. was described from Portugal. On the basis of the original description, Sim-Sim (1989) stated that this taxon corresponded to Scapania compacta (A. Roth) Dumort. and proposed this synonymy without studying the type. Since the type cannot be located a formal synonymyzation should be made following designation of a neotype. 64. Sphaerocarpos algeriensis Trab. nom. nud. was reported by Trabut (1941) from Algeria. Jovet-Ast (1956) indicated that the morphological characters given for S. algeriensis fall within the range of variation observed in S. texanus Austin in Morocco and that it is probably only a synonym of that species. 65. Schumacker & Vá≈a (2005) proposed a new combination for Lepidozia azorica in the genus Telaranea as T. azorica (H. Buch & Perss.) Pócs ex Schumacker & Vá≈a. 66. Telaranea nematodes (Gottsche ex Austin) M. Howe was considered to be the correct name for the species present in Europe and Macaronesia until Engel & Merrill (2004) established a new name for it, Telaranea europaea J. J. Engel & G. L. S. Merrill. 67. Telaranea sejuncta auct. non (Angst.) Arnell is a confusing taxon. It was assigned to T. nematodes (Gottsche ex Austin) M. Howe by Arnell (1963), and this synonymy has been widely used. According to Engel & Merrill (2004), “all the published statements of distribution, descriptions and illustrations of T. sejuncta (apart from the type) are suspect, since they may include some or all the taxa formerly included in T. nematodes, as well as T. diacantha, when, in fact, it is none of these”. In our current state of knowledge, it can be assumed that all the Mediterranean reports of T. nematodes and T. sejuncta correspond to T. europaea. 68. Trichocolea tomentella (Ehrh.) Dumort. was recorded from Malta by Gulia (1858-59) and Sommier & Caruana Gatto (1915) as “Jungermannia tomentella (Ehr.)”. These reports are almost certainly erroneous and are here rejected. Acknowledgements. The authors wish to express their gratitude to David Long and Jesús Muñoz who kindly provided information concerning various taxa, and to Susana Rams and Olaf Werner for their help with the tables and figure. Ana Séneca helped to coordinate the list with the European checklist. 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Hepatics and Anthocerotes of the Mediterranean, an annotated checklist

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