CYKKYJIEHTbI / SUCCULENTS (TOM \ vol III) 2000 XI! 1-2 NOMENCLATURAL ADJUSTMENTS IN CACTEAE (CACTOIDEAE-CACTACEAE), II. A.B. Doweld, National Institute of Carpology (Gaemerian Institution), Moscow, Russia This note is a continuation of the series of papers (Doweld, 1999), dealing with the nomenclatural changes in various taxa of the tribe Cacteae, in order to bring current nomenclature in accord with a new taxonomic treatment. ASTROPHYTUM The infrageneric system ofthe genus necessitates the validation of series, proposed, but never validated by Backeberg (1942). The fonner proposals of the subdividing the genus into two subgenera or even sections seems too excessive to emphasize the distinctness of two group of closely related species within the genus. , Astrophytum ser. Astrophytum =T Astrophytum ser. Lutei-faucinati Backeb. 1942, Cactaceae (Berlin), 1941, 2: 55, nom. inval. =T Astrophytum sect. Austrastrophytum M. Megata, 1944, Mem. CoIl. Agric. Kyoto Imp. Univ. 56 : 36, nom. inval. T y pus: A, myriostigma Lem. 1. Astrophytum ser. Anastrophytum Doweld, ser. nov. =T Subgen. Neoastrophytum Backeb. 1950, Caet. Succ. lourn. (Los Angeles), 22, 5 : 154 [quoad Lat. descr.]. =T Sect. Septentriastrophytum M. Megata, 1944, Mem. CoIl. Agric. Kyoto Imp. Univ. 56 : 42 [quoad Lat. descr.]. =T Ser. Rubro-faucinati Backeb. 1942, Cactaceae (Berlin), 1941,2: 55, nom. nud. T y pus: A. asterias (Zucc.) Lem. 2. An additional necessary transfer to subspecific rank should be made: Astrophytum capricorne ssp. senile (A. V. Fric) Doweld, stat. nov. =T Astrophytum senile A. V. Fric, 1925, Zivot v Prirod. 29, 10: 35; 29,12: 44 (nom, valid.!). - Lectotypus (designated here): photo in A. V. Fric, 1925,29,10 : 35. BISNAGA Bisnaga latispina (Haw.) Doweld, comb. nov. =T Cactus latispinus Haw. 1824, Philos. Mag. Journ. 63, 309: 41. - Neotypus (Lindsay ex Unger, 1992, Gross. Kugelkakt. : 302): G. Lindsay 2583 (DS). Bisnaga latispina (Haw.) Doweld ssp. spiralis (Karwinsky ex Pfeiff.) Doweld, comb. nov. =T Echinocactus spiralis Karwinsky ex Pfeiff. 1837, Enum. Cact. : 60-61, 180. - Lectotypus (designated here): tab. 16(4) in Pfeiff. 1839, Nov. Act. Acad. Caes. Leop. Carol. 19: 124. PARRYCACTUS Parrycactus alamosanus (Britton & Rose) Doweld, comb. nov. =T Echinocactus alamosanus Britt. et Rose, 1913, Contr. US Natl. Herb. 16: 239.,- Holotypus: Rose, Standley & Russel 12850 (US). Parrycactus alamosanus ssp. reppenhagenii (G. Unger) Doweld, comb. et stat. nov. =T Ferocactus reppenhagenii G. Unger, 1974, Kakteen Sukk. 25, 3: 50. - Holotypus: W. Reppenhagen & E. Priessnitz s.n. in 25 luI. 1973 (ZSS). TORREYCACTUS Torreycactus conothelos ssp. conothelos Torreycactus conothelos ssp. garciae (Glass & Mendoza-Garcia) Doweld, comb. et stat. nov. =T Thelocactus garciae Glass & Mendoza-Garcia, 1997 (' 1998'), Guia Ident. Cact. Mex. 1 : sine pag. [TH/GA]. - Holotypus: Garcia Luna 383 (CANTE). = Thelocactus flavus Mosco & ZanoveIlo, 1999, Cactus & Co. 3, I : 20, 22. Holotypus: Kuenzler 362 (HG-PAD). Torreycactus conothelos ssp. aurantiacus (Glass & Foster) Doweld, comb. et stat. nov. =T Thelocactus conothelos var. aurantiacus Glass & Foster, 1972, Cact. Suec. 1. (Los Angeles), 44, 2: 48. - Holotypus: Glass & Foster 3183 (ZSS). 36 CYKKYJlEHTbl / SUCCULENTS (TOM \ vol III) 2000 N2 1-2 Torreycactus conothelos ssp. argenteus (Glass & Foster) Doweld, comb. et stat. nov. =T The/ocactus conothe/os var. argenteus Glass & Foster, 1972, Cact. Succ. J. (Los Angeles), 44, 2: 47-48. - Ho)otypus: G/ass & Foster 3176 (ZSS). GYMNOCACTUS Gymnocactus vierecki; (Werd.) Backeb. ssp. viereckii Gymnocactus viereckii (Werd.) Backeb. ssp. major (Glass & Foster) Doweld, stat. nov. =T Gymnocactus viereckii var. major Glass & Foster, 1978, Cact. Succ. J. (Los Angeles), 50, 6 : 285. - Ho)otypus: G/ass & Foster 4110 (POM). KADENICARPUS Kadenicarpus pseudomacrochele ssp. pseudomacrochele; Kaden/carpus pseudomacrochele ssp. lausseri (Diers & Frank) Doweld, comb. et stat. nov. =T Turbinicarpus pseudomacroche/e var.lausseri Diers & Frank, 1991, Kakteen Sukk. 42, 5 : 108. - Ho)otypus: [unseen] (KOELN). With the recognition ofa ､ｩｳｴｮｾ＠ NEOBESSEYA genus Neobesseya in present system of the Cacteae, new combinations are needed: Neobesseya robbinsorum (W. H. Earle) Doweld, comb. nov. =T Cochiseia robbinsorum W. H. Earle, 1976, Saguaroland Bull. 30 : 64-66. - Ho)otypus: Robbins S.n. (ARIZ). Neobesseya missouriensis (Sweet) Britt. & Rose ssp. simi/is (Engelm.) Doweld, comb. et stat. nov. =T Mammillaria similis Engelm. 1845, Boston J. Nat. Hist. 5 : 246. - Ho)otypus: F. J Lindheimer S.n. in 1845 (MO). DOLICHOTHELE With the resurrection of the genus Dolichothele, a new formal nomenclatural combination is needed: Dolichothele carretli (Rebut ex K. Schum.) Doweld, comb. nov. =T Mammillaria carretii Rebut ex K. Schum. 1898, Gesamtbeschr. Kakt. : 542. - Neotypus (designated here): "Mexico, Nuevo Leon, near Icamole", 3 II 1907, W. E. Safford 1250 (US) (as 'Mammillaria saffordii'). SOLISIA With the amplification of the genus (present issue), a new combination is needed: Solisla solisioides (Backeb.) Doweld, comb. nov. =T Mammillaria solisioides Backeb. 1952, Cactus (Paris), 31, Supp!. : 3. - Lectotypus (Mottram, 1980, Mamm. Index: 86): fig. 3174 in Backeb. 1961, Cactaceae, 5 : 3434. BARTSCHELLA The revision made of the genus (present issue) requires new combinations should be made: Bartschella albicans (Britt. & Rose) Doweld, comb. nov. =T Neomammillaria albicans Britt. & Rose, 1923, Cactaceae,4: 138. - Holotypus: Rose 16842 (US). Bartschella albicans (Britt. & Rose) Doweld ssp. !rai/eana (Britt. & Rose) Doweld, comb. et stat. nov. =T Neomammillariafraileana Britt. & Rose, 1923, Cactaceae, 4: 157. - Ho)otypus: Rose 16508 (US). Bartschella armil/ata (K. Brandegee) Doweld, comb. nov. =T Mammillaria arm illata K. Brandegee, 1900, Zoe, 5 : 7. - Holotypus: T. S. Brandegee s. n. in 1893 (UC 108185). Bartschella boolii (G. Lindsay) Doweld, comb. nov. =T Mammillaria boolii G. Lindsay, 1953, Cact. Succ. J. (Los Angeles), 25, 2 : 48. - Ho)otypus: Lindsay 2220 (DS). Bartschella cerralboa (Britt. & Rose) Doweld, comb. nov. =T Neomammillaria cerralboa Britt. & Rose, 1923, Cactaceae,4 : 116. - Ho)otypus: 1. M Johnston 4038 (US). Bartschella deherdtiana (Farwig) Doweld, comb. nov. =T Mammillaria deherdtiana Farwig, 1969, Cact. Succ. J. (Los Angeles), 41, 1 : 28. - Ho)otypus: Schwarz S.n. in 1959 (POM). Bartschella deherdtiana (Farwig) Doweld ssp. dodsonii (Bravo) Doweld, comb. et stat. nov. =T Mammillaria dodson;; Bravo, 1970, Cact. Suc. Mex. 15, I : 3-5. - Holotypus: J. W. Dodson 401 (MEXU 118775). Bartschella glass;; (R. A. Foster) Doweld, comb. nov. =T Mammillaria glassii R. A. Foster, 1968, Cact. Succ. J. (Los Angeles), 40, 4: 132. - Holotypus: Glass & Foster 631 (RSA). Bartschella glass;; (R. A. Foster) Doweld ssp. ascension is (Reppenhagen) Doweld, comb. et stat. nov. =T 37 CYKKYJJEHTbl1 SUCCULENTS (TOM \ vol III) 2000 N!! 1-2 Mammillaria ascensionis Reppenhagen, 1979, Kakteen Sukk. 30, 3 : 61-62. - Holotypus: Reppenhagen 1062 (ZSS). Bartschella glassii (R. A. Foster) Doweld ssp. nominis-dulcis (Lau) Doweld, comb. et stat. nov. =T Mammillaria glassii var. nominis-dulcis Lau, 1985, Caet. Suee. J. (Los Angeles), 57,5 : 198. - Holotypus: Lau 1186a (MEXU). Bartschella glass;; (R. A. Foster) Doweld ssp. siberiensis (Lau) Doweld, comb. et stat. nov. =T Mammillaria glassii var. siberiensis Lau, 1985, Caet. Suee. J. (Los Angeles), 57, 5 : 198. - Holotypus: Lau 1322 (MEXU). Bartschella guelzowiana (Werdenn.) Doweld, comb. nov. =T Mammillaria guelzowiana Werdenn. 1928, Zeitsehr. Sukkulentenk.3 : 356. - Lectotypus (Mottram, 1980, Mamm. Index: 37 as 'typus'): fig. [unnumbered] in Werdenn.l.e. Bartschella heidiae (Krainz) Doweld, comb. nov. =T Mammillaria heidiae Krainz, 1975, Kakteen Sukk. 26, 10 : 217. - Holotypus: KrMenbilhl280174 (ZSS). Bartschella hernandezii (Glass & Foster) Doweld, comb. nov. =T Mammillaria hernandezii Glass & Foster, 1983, Caet. Suee. J. (Los Angeles), 55, 1 : 22. - Holotypus: Otero & Hernandez FO 023 (POM). Bartschella hutchisoniana (Gates) Doweld, comb. nov. =T Neomammillaria hutchisoniana Gates, 1934, Caet. Suee. 1. (Los Angeles), 6, 1 : 4. - Holotypus: Gates 121 (DS). Bartschella hutchisoniana (Gates) Doweld ssp. louisiae (Lindsay) Doweld, comb. et stat. nov. =T Mammillaria louisiae Lindsay, 1960, Caet. Suee. J. (Los Angeles), 32, 6 : 169. - Holotypus: Lindsay 2022 (SD). Bartschella insularis (Gates) Doweld, comb. nov. =T Mammillaria insularis Gates, 1938, Caet. Suee. J. (Los Angeles), 10: 25. _. Holotypus: Gates 523 (DS). Bartschella multidigitata (G. Lindsay) Doweld, comb. nov. =T Mammillaria multidigitata G. Lindsay, 1947, Caet. Suee. 1. (Los Angeles), 19, 10 : 152. - Holotypus: G. Lindsay & Bool499 (DS 314192). Bartschella napina (J. A. PUJ-pus) Doweld, comb. nov. =T Mammillaria napina J. A. Purpus, 1912, Monatssehr. Kakteenk. 22 : 161. - Lectotypus (Mottram, 1980, Mamm. Index: 59 as 'typus '): fig. [unnumbered] in Monatssehr. Kakteenk.23: 123. 1913. Bartschella palmeri (Coulter) Doweld, comb. nov. =T Cactus palmeri Coulter, 1894, Contr. US Natl. Herb. 3, 2 : 108. =T Mammillaria palmeri (Coult.) Bod. 1933 non Jacobi, 1856 (=T M neopalmeri Craig, 1945). Holotypus: Palmer 921 (US 1821091). Bartschella tetrancistra (Engelm.) Doweld, comb. nov. =T Mammillaria tetrancistra Engelm. 1852, Amer. J. Sci. Ser.2, 14: 337. - Neotypus (L. Bens. 1969, Cacti Ariz. ed. 3: 163): L. Benson 15716 (POM 288535). Bartschella yaquensis (Graig) Doweld, comb. nov. =T Mammillaria yaquensis Graig, 1945, Mammillaria Handb. : 320. - Holotypus: Hilton & Craig in 1937 (DS). Bartschella zephyranthoides (Seheidw.) Doweld, comb. nov. =T Mammillaria zephyranthoides Seheidw. 1841, AUg. Gartenzeit. 9 : 41. - Neotypus (required): Validation of infrageneric divisions: Bartschella ser.Glassella Doweld, ser. nov. - A ser. Bartschello aculeis radialibus vulgo pluribus (55-75) et aculeis centralibus pluribus (8-10) differt. Typus: Bartschella glass;; (R. A. Foster) Doweld (In honorem Ch. Glass nominatur). Bartschella ser. Napinae (Keizer) Doweld, comb. nov. =T Mammillaria ser. Napinae Keizer, 1991, Sueculenta, 70,6: 125. Typus: Bartschella napina (J. A. Purpus) Doweld. Bartschella ser. Probartschella Doweld, ser. nov. - A ser. Bartschello aculeis radialibus vulgo minoribus (lOIS), raro ad 20, aculeis centralibus pluribus (3-4), foveolis testae profundibus, ornamentatione cuticularis minusfavosibus differt. Typus: Bartschella arm illata (K. Brandegee) Doweld. Bartschella ser. Phellosperma (Britt. & Rose) Doweld, stat. nov. =T Phellosperma Britt. & Rose, 1923, Cactaeeae, 4 : 60. Typus: Bartschella tetrancistra (Engelm.) Doweld. Bartschella ser. Zephyranthoides (E. Kuhn & B. Hofmann) Doweld, comb. nov. =T Mammillaria ser. Zephyranthoides E. Kuhn & B. Hofmann, 1979, Infonn.-Brief ZAG Mammill. 5,3 : 41--42. Typus: Bartschella zephyranthoides (Scheidw.) Doweld. COCHEMIEA After new delimitation and fundamental revision of Mammillaria sect. Cochemiea (present issue), the genus Cochemiea is amplified a<; follows: Coehemiea barbata (Engelm.) Doweld, comb. nov. =T Mammillaria barbata Engelm. 1848, in Wislizenus, Mem. Tour North. Mex. : 105. - Holotypus: Wislizenus S.n. in 1846 (MO). 38 CYKKYJIEHTbI/ SUCCULENTS (TOM \ vol III) 2000 N!! 1-2 Cochemiea capensis (Gates) Doweld, comb. nov. =T Neomammillaria capensis Gates, 1933, Cact. Succ. J. (Los Angeles),4, 11 : 372. - Holotypus: Gates 68 (DS). Cochemiea diolca (K. Brandegee) Doweld, comb. nov. =T Mammillaria dioica K. Brandegee, 1897, Erythea, 5 : 115.- Lectotypus (L. Bens. 1969, Cacti Ariz. ed. 3: 159): Wright S.n. in 1852 (MO). Cochemiea dioiea (K. Brandegee) Doweld ssp. angelensis (Craig) Doweld, comb. et stat. nov. =T Mammillaria angelensis Craig, 1945, Mammillaria Handb. : 165. - Holotypus: G. Lindsay [s.n.?J in 1936 (DS). Coehemiea dioica (K. Brandegee) Doweld ssp. estebanensis (Lindsay) Doweld, comb. et stat. nov. =T Mammillaria estebanensis Lindsay, 1967, Cact. Succ. J. (Los Angeles), 39, 1 : 31. - Holotypus: Lindsay 3002 (DS). Cochemiea graham;; (Engelm.) Doweld, comb. nov. =T Mammillaria grahamii Engelm. 1856, Proc. Amer. Acad. Sci. 3 : 262. - Lectotypus (L. Bens. 1969, Cacti Ariz. ed. 3: 159): Wright S.n. in 1852 (MO). Cochemiea phitauiana (Baxter) Doweld, comb. nov. =T Neomammillaria phitauiana Baxter, 1931, Cact. Succ. J. (Los Angeles), 2, II : 472. - Holotypus: E. M Baxter S.n. in 5 IV 1930 (US 1491115). Coehemiea saboae (C. Glass) Doweld, comb. nov. =T Mammillaria saboae C. Glass, 1966, Cact. Suc. Mex. 11 : 55. - Holotypus: K. Sabo S.n. (MEXU). Cochemiea saboae (c. Glass) Doweld ssp. gold;; (c. Glass & Foster) Doweld, comb. nov. =T Mammillaria goldii C. Glass & Foster, 1968, Cact. Succ. J. (Los Angeles), 40, 4 : 151. - Holotypus: Glass & Gold 102711 (POM). Cochemiea saboae (C. Glass) Doweld ssp. haudeana (Lau & Wagner) Doweld, comb. et stat. nov. =T Mammillaria haudeana Lau & Wagner, 1978, Kakteen Sukk. 29, 11 : 253. - Holotypus: Lau 777 (HElD 46636). Cochemiea swinglei (Britt. & Rose) Doweld, comb. nov. =T Neomammillaria swinglei Britt. & Rose, 1923, Cactaceae,4 : 158. - Holotypus; Rose & al. 12568 (US). Cochemiea theresae (Cutak) Doweld, comb. nov. =T Mammillaria theresae Cutak, 1967, Cact. Succ. 1. (Los Angeles), 39, 6 : 239. - Holotypus: E. Gay & B. Gay 2411 (MO). Cochemiea wrightii (Engelm. & Bigelow) Doweld, comb. nov. =T Mammillaria wrightii Engelm. & Bigelow, 1856, Proc." Amer. Acad. Sci. 3 : 262. - Lectotypus (Coulter, 1894, Contr. US Natl. Herb. 3 : 101 as 'typus '): Wright s' n. in VIII 1851 (MO 2016409). Cochemiea wrlghtii (Engelm. & Bigelow) ssp. wilcoxi; (Tourney ex K. Schum.) Doweld, comb. et stat. nov. =T Mammillaria wilcoxii Tourney ex K. Schum. 1898, Gesamtbeschr. Kakt. : 545. - Neotypus (L. Bens. 1977, Cact. Succ. J. (Los Angeles), 49, I : 35): D. A. Zimmermann & A. D. Zimmermann 2788 (POM 322896). Validation of infrageneric groupings: Cochemiea sect. Protocochemiea Doweld, sect. nov. - Plantae plerumque cylindricae, caespitosae, aculeis radialibus 4-6-multiplibus, minoribus (/2-18) vel secondarie pluribus (40-60), aculeis centralibus ad normam 4-6, vel minoribus 1-2 (ad absentibus). - Typus: Cochemiea capensis (Gates) Doweld. Cochemiea sect. Mamillopsis (Morren) Doweld, comb. nov. =T Mammillaria sect. Mamillopsis Morren, 1874, Belg. Hortic. 24: 33. Typus: Cochemiea:senilis (Loddiges ex Salm-Dyck) Orcutt. Cochemiea ser. Barbatae Doweld, ser. nov. - Plantae solitariae vel caespitosae, aculeis centralibus plerumque (1-)2(-4), radialibus in duos series dispositis 50-60 vel minoribus (12-14), 4-6-multiplibus. - Typus: Cochemiea barbata (Engelm.) Doweld. Cochemiea ser. Cochemieoidei Doweld, ser. nov. - Plantae cylindricae, aculeis centralibus absentis vel raro solitariis, radialibus plerumque 12-18, 4-6-multiplibus. - Typus: Cochemiea capensis (Gates) Doweld. Cochemiea ser. Saboae Doweld, ser. nov. - Plantae caespitosae, aculeis centralibus nullis (raro I), radialibus pluribus (4-46). - Typus: Cochemiea saboae (C. Glass) Doweld. Cochemiea ser. Seniles Doweld, ser. nov. - Plantae cylindricae vel globulares, plerumque caespitosae, aculeis centralibus 4-6, radialibus pluribus 40-60, 4-6-multiplibus. - Typus: Cochemiea senilis (Loddiges ex SalmDyck) Orcutt. ESCOBARIOPSIS After re-c1assification of the genus Mammillaria s. I. (present issue), the species, formerly included in the series Herrerae, Lasiacanthae, Sphacelatae, Proliferae, and Bombycinae of Mammillaria s.I., are transferred into a newly established genus Escobariopsis. Available infrageneric groupings (series) are also transferred to maintain the nomenclatural stability; three sections of the genus are recognized and validated. Escobariopsis albicoma (Bod.) Doweld, comb. nov. =T Mammillaria albicoma Bod. 1929, Monatsschr. Deutsch. Kakt. Ges. I : 241. - Lectotypus (designated here): fig. [unnumbered] in Bod. I.c. Epitypus (Reppenhagen, 1991, Gatt. Mamm. 1 : 256 as 'neotypus'): Reppenhagen 1794 (KL). 39 CYKKYJIEHTbI/ SUCCULENTS (TOM \ vol III) 2000 N!! 1-2 Escobariopsis albifiora (Werdenn.) Doweld, comb. et stat. nov. =T Mammillariah,errerae var. albiflora Werdenn. 1931, Notizbl. Bot. Gart. Mus. Berlin, 11, 104: 277. - Neotypus Ｈｲ･ｱｵｩｾ､ＩＮ＠ Escobariopsi'i aureitanata (Backeb.) Doweld, comb. nov. =T Mammillaria aureilanata Backeb. 1938, Beitr. Sukkulentenk. -Pflege, 1938 : 13. - Lectotypus (designated here): fig. [unnumbered] in Quehl, 1914, Monatssehr. Kakteenk. 24 : 158 (as 'Mammillaria cephalophora' Quehl non Salm-Dyck). Escobariopsis bombycina (Que hi) Doweld, comb. nov. =T Mammillaria bombycina Quehl, 1910, Monatsschr. Kakteenk. 20 : 149. - Lectotypus (Mottram, 1980, Mamm. Index: 10 as 'typus '): fig. [unnumbered] in Quehl, I.e. : 151. Escobariopsis carmenae (Castaneda) Doweld, comb. nov. =T Mammillaria carmenae Castaneda, 1954, Anales Inst. BioI. Univ. Mex. 24, 2: 233. - Lectotypus (designated here): fig. in Castaneda, I.c. Epitypus (Ahuatzin, 1977, Cact. Suc. Mex. 22, 4: 86 as 'lectotypus'): Lau JJ92 (MEXU). Escobariopsisjittkaui (Glass & Foster) Doweld, comb. nov. =T Mammillariafittkaui Glass & Foster, 1971, Caet. Succ. J. (Los Angeles), 43, 3 : 116. - Holotypus: Fittkau [Abbey Gard 69-1169J (POM). Escobariopsisjittkaui (Glass & Foster) Doweld ssp. limonem'i'i (Reppenhagen) Doweld, comb. et stat. nov. =T Mammillaria limonensis Reppenhagen, 1985, Kakteen Sukk. 36, 3 : 44-46. - Holotypus: Reppenhagen 1620 (K). Escobariopsisjittkaui (Glass & Foster) Doweld ssp. mathi/dae (Krahenbuhl & Krainz) Doweld, comb. et stat. nov. =T Mammillaria mathi/dae KrahenbUhl & Krainz, 1973, Kakteen Sukk. 24, 12 : 265-266. - Holotypus: Wagner S.n. (ZSS). Escobariopsis gracilis (Pfeiff.) Doweld, comb. nov. =T Mammillaria gracilis Pfeiff. 1838, Allg. Gartenzeit. 6 : 275. - Neotypus (required). Escobariopsis herrerae (Werdenn.) Doweld, comb. nov. =T Mammillaria herrerae Werderm. 1931, Notizbl. Bot. Gart. Mus. Berlin, 11 : 276. - Holotypus: Schmoll S.n. (B). Escobariopsis humboldtii (Ehrenb.) Doweld, comb. nov: =T Mammillaria humboldt;i Ehrenb. 1840, Linnaea, 14 : ＳＷＸｾ＠ - Neotypus (required). Escobariopsisjaliscana (Britt. & Rose) Doweld, comb. nov. =T Neomammillaria bombycina Britt. & Rose, 1923, Caetaceae,4 : 160. - Holotypus: Rose & Painter 858 (US 399574). Escobariopsis jaliscqlla (Britt. & Rose) Doweld ssp. zacatecasensis (Shurly) Doweld, comb. 'et stat. nov. =T Mammillaria zacatecas ens is Shurly, 1960, Caet. Suce. J. Grt. Brit. 22, 3 : 51. - Holotypus: Schwarz s,n. (K 6620). Escobariopsis kraehenbuehlii (Krainz) Doweld, comb. nov. =T Pseudomammillaria kraehenbuehlii Krainz, 1970, Kakteen Sukk. 1970, 12 : sine pag. (Gesellschaftsnachr.) [4]. - Holotypus: Krahenbuhl 153.68 (,153:69,) (ZSS). Escobariopsis lasiacantha (Engelm.) Doweld, comb. nov. =T Mammillaria lasiacantha Engelm. 1856, Proc. Amer. Aead. Sci. 3 : 261. -- Holotypus: Wright S.n. (MO). Escobariopsis lasiacantha (Engelm.) Doweld ssp:egregia (Baekeb. ex Rogozinski & Appenzeller) D(lWeld, comb. et stat. nov. =T Mammillaria egregia Baekeb.ex Rogozinski & Appenzeller, 1989, Mitteilungsbl. Arbeitskr. Mammill. 13,3: 108. - Holotypus: Rogozinski 14/86 (ZSS). Escobariopsis lasiacantha (Engelm.) Doweld ssp. magal/anii (Craig) Doweld, comb. et stat. nov. =T Mammillaria magallanii Craig, 1945, Mammillaria Handb. : 225. - Lectotypus (Mottram, 1980, Mamm. Index: 50 as 'typus 1: fig. 205 in Craig, I.e. Escobariopsis lasiacantha (Engelm.) Doweld ssp. wohlschlageri (Reppenhagen) Doweld, comb. et stat. nov. =T Mammillaria wohlschlageri Reppenhagen, 1987, Gatt. Mammillaria: 49-50. - Holotypus: Reppenhagen 2132 (ZSS). , Escobariopsis laui (D. R. Hunt) Doweld, comb. nov. =T Mammillaria laui D. R. Hunt, 1979, Caet. Succ. J. Grt. Brit. 41,4 : 100. - Holotypus: Lau 1171 (K). Escobar/opsis laui (D. R. Hunt) Doweld ssp. dasyacantha (D. R. Hunt) Doweld, comb. et stat. nov. =T Mammillaria laui f. dasyacantha D. R. Hunt, 1979, Caet. Suce. J. Grt. Brit. 41,4: 100. -Holotypus: Lau 1219 (K). Escobariopsis laui (D. R. Hunt) Doweld ssp. subducta (D. R. Hunt) Doweld, comb. et stat. nov. =T Mammillaria laui f. subducta D. R. Hunt, 1979, Cact. Suce. J. Grt. Brit. 41,4 : 100. - Holotypus: Lau 1222 (K). Escobariopsis lenta (K. Brandegee) Doweld, comb. nov. =T Mammillaria lenta K. Brandegee, 1904, Zoe, 5 : 194. - Holotypus: C. A. Purpus II (UC 125367). Escobariopsis mazatlanensis (K. Schum. ex Gurke) Doweld, comb. nov. =T Mammillaria mazatlanensis K. Schum. ex Gurke, 1905, Monatsschr. Kakteenk. 15, 10: 154. - Lectotypus (Mottram, 1980, Mamm. Index: 53 as 'typus 1: fig. [unnumbered] in Gurke, I.e. : 155. . 40 CYKKYJlEHTb1 I SUCCULENTS (TOM \ vol III) 2000N!! 1-2 Escobariopsis mazatlanensis (K. Schum. ex Gurke) Doweld ssp. patonii (Bravo) Doweld, comb. et stat. nov. =T Neomammillaria patonii Bravo, 1931, Anales lnst. BioI. Mex. 2 : 129. - Lectotypus (Mottram, 1980, Mamm. Index: 65 'as typus '): fig. 2 in Bravo, I.e. Escobariopsis mercadensis (Patoni) Doweld, comb. nov. =T Mammillaria mercadensis Patoni, 1910, BoL Alianza Cient. Univ. Durango, I : 54. - Neotypus (Reppenhagen, 1991, Gatt. Mamm. 1 : 184): Reppenhagen 1772 (KL). Escobariopsis moelleriana (Bod.) Doweld, comb. nov. =T Mammillaria moe/leriana Bod. 1927, Zeitsehr. Sukkulentenk.l : 213. -Lectotypus (Mottram, 1980, Mamm. Index: 56 as 'typus'): fig. [unnumbered] in Bod. l.e.:2I4. Escobariopsis oteroi (Glass & Foster) Doweld, comb. nov. =T Mammillaria oteroi Glass & Foster, 1975, Caet. Suee. 1. (Los Angeles), 47, 2 : 94. - Holotypus: Otero S.n. (POM). Escobariopsis pennispinosa (Krainz) Doweld, comb. nov. =T Mammillaria pennispinosa Krainz, 1948, Sukkulentenk. 2 : 20. - Lectotypus (Mottram, 1980, Mamm. Index: 66 as 'typus '): fig. [unnumbered] in Krainz, I.e. Escobariopsis pennispinosa (Krainz) Doweld ssp. nazasensis (Glass & Foster) Doweld, comb. et stat. nov. =T Mammillaria pennispinosa var. nazasensis Glass & Foster, 1975, Caet. Suee. J. (Los Angeles), 47, 2 : 96. Holotypus: Glass & Foster 802 (ZSS). Escobariopsis perezdelarosae (Bravo & Seheinvar) Doweld, comb. nov. =T Neomammillaria perezdelarosae Bravo & Seheinvar, 1985, Caet. Sue. Mex. 30, 4 : 76. - Holotypus: Bravo & Scheinvar 3938 (MEXU). Escobariopsis picta (Meinshausen) Doweld, comb. nov. =T Mammillaria picta Meinshausen, 1858, Woehensehr. Gart. Pfl. 1 : 27. - Neotypus (required). Escobariopsis pilispina (J. A. Purpus) Doweld, comb. nov. =T Mammillaria pilispina 1. A. Purpus, 1912, Monatssehr. Kakteenk. 22 : 150. - Neotypus: (required). Escobariopsis plumo!J'a (F. A. C. Weber) Doweld, comb. nov. =T Mammillaria plumosa F. A. C. Weber, 1898, in Bois, Diet. Hort. : 804. - Neotypus (Reppenhagen, 1991, Gatt. Mamm. 1 : 290): Reppenhagen 1217 (KL). Escobariopsis prolifera (Miller) Doweld, comb. nov. =T Cactus proliferus Miller, 1768, Gard. Diet. ed. 8 : sine pag. [Cactus sp. N 6]. - Neotypus (required ?). Escobariopsis prolifera (Miller) Doweld ssp. texana (Engelm.) Doweld, comb. et stat. nov. =T Mammillaria pusilla var. texana Engelm. 1859, Caet. Mex. Bound. : 5. - Lectotypus (designated here): pI. 5 in Engelm. I.e. Escobar/opsis prolifera (Miller) Doweld ssp. haitiemis (K. Schum.) Doweld, comb. et stat. nov. =T Mammillaria pusil/a var. haitiensis K. Schum. 1904, BlUh. Kakt. I : sub pI. 46. - Lectotypus (designated here): pI. 46 in K. Sehum.l.c. Escobariopsis prolifera (Miller) Doweid ssp. arachnoidea (D. R. Hunt) Doweld, comb. et stat. nov. =T Mammillaria prolijera var. arachnoidea D. R. Hunt, 1978, Caet. Succ. J. Grt. Brit. 40, 1 : 11. - Holotypus: D. R. Hunt 8598 (K). Escobariopsis prolifera (Miller) Doweld ssp. zublerae (Reppenhagen) Doweld, comb. et stat. nov. =T Mammillaria zublerae ("zubleri ") Reppenhagen, 1987, Gatt. Mammillaria: 48-49. - Holotypus: Zubler s. n. (ZSS). Escobariopsis rayonesensis (Reppenhagen) Doweld, comb. nov. =T Mammillaria rayonesensis Reppenhagen, - Holotypus: Reppenhagen 1290 (KL). 199 I, Gatt. Mammillaria, 1 : ＲＶｾＱＮ＠ Escobariopsis rettigiana (Bod.) Doweld, comb. nov. =T Mammillaria retligiana Bod. 1930, Monatssehr. Deutsch. Kakt. Ges. 2 : 98. - Lectotypus (Mottram, 1980, Mamm. Index: 76 as 'typus '): fig. [unnumbered] in Bod. I.c. Escobariopsis sanchez-mejoradae (R. Gonzalez G.) Doweld, comb. nov. =T Mammillaria sanchez-mejoradae R. Gonzalez G. 1992, Cact. Sue. Mcx. 37, 3 : 55-59. - Holotypus: R. Gonzruez G. 4300 (MEXU). Escobariopsis schiedeana (Ehrenb.) Doweld, comb. nov. =T Mammillaria schiedeana Ehrenb. 1838, AUg. Gartenzeit. 6 : 249. - Neotypus (required). Escobariopsis sinistrohamata (Bod.) Dowdd, comb. nov. =T Mammillaria sinistrohamata Bod. 1932, Monatsschr. Deutsch. Kakt. Ges. 4 : 162. - Lectotypus (Mottram, 1980, Mamm. Index: 85 as 'typus '): fig. [unnumbered] in Bod. I.c. Epitypus (Reppenhagen, 1991, Gatt. Mamm. 1 : 190 as 'neotypus '): Reppenhagen 1254 (KL). Escobariopsis sphacelata (Martius) Doweld, comb. nov. =T Mammillaria vetula Martius, 1832, Nov. Act. Acad. Caes. Leop. Carol. 16, I : 339. - Lectotypus (Hunt, 1974, J. Mamm. Soc. 14: 35 as 'typus): tab. 25, fig. I in Martius, I.c. Escobariopsis tonalensis (D. R. Hunt) Doweld, comb. nov. =T Mammillaria tonalensis D. R. Hunt, 1979, Caet. Suce. 1. Grt. Brit. 41,4 : 103. - Holotypus: Lau 1114 (K). Escobariopsis vetula (Martius) Doweld, comb. nov. =T Mammillaria vetula Martius, 1832, Nov. Act. Acad. Caes. 41 CYKKYJlEHTbI / SUCCULENTS (TOM \ vol III) 2000 N2 1-2 Leop. Carol. 16, I : 338. - Lect()typus (Mottram, 1980, Mamm. Index: 97 as 'typus '): tab. 24 in Martius, I.c. Escobar/opsis viereck;; (Bod.) Doweld, comb. nov. =T Mammillaria viereckii Bod. 1927, Zeitschr. Sukkulentenk. 3 : 73. - Lectotypus (Mottram, 1980, Mamm. Index: 97 as 'typus '): fig. [unnumbered] in Bod. I.c. : 74. Escobariopsis viperina (J. A. Purpus) Doweld, comb. nov. =T Mammillaria viperina J. A. Purpus, 1912, Monatsschr. Deutsch. Kakt. Ges. 22: 148. - Lectotypus (Mottram, 1980, Mamm. Index: 97 as 'typus '): fig. [unnumbered] in J. A. Purpus, I.c. 23 : 21. 1913. Epitypus (Reppenhagen, 1991, Gatt. Mamm. 1 : 329 as 'neotypus'): Reppenhagen 118 (KL). Escobariopsis weingartiana (Bod.) Doweld, comb. nov. =T Mammillaria weingartiana Bod. 1932, Monatsschr. Deutsch. Kakt. Ges. 4: 219. -Lectotypus (Mottram, 1980, Mamm. Index: 99 as 'typus'): fig. [unnumbered] in BOd.l.c. Validation oflnfrageneric taxa: Escobariopsb; sect. Acentracantha (F. Buxb.) Doweld, comb. et stat. nov. =T Chi/ita subgen. Acentracantha F. Buxb. 1954, Sukkulentenk. 5 : 25. Typus: Escobariopsis lasiacantha (Engelm.) Doweld. Escobariopsis ser. Sphacelatae (D. R. Hunt) Doweld, comb. nov. =T Mammillaria ser. Sphacelatae D. R. Hunt, 1977, Cact. Succ. J. Grt. Brit. 39, 3 : 73. Typus: Escobariopsis sphacelata (Martius) Doweld. Escobariopl'is ser. Lasiacanthae (D. R. Hunt) Doweld, comb. nov. =T Mammillaria ser. Lasiacanthae D. R. Hunt, 1971, Cact. Succ. J. Grt. Brit. 33, 3: 63. Typus: Escobariopsis lasiacantha (Engelm.) Doweld. Escobariopsis ser. Herrerae (LUthy) Doweld, comb. nov. =T Mammillaria ser. Hem:!rae LUthy, 1995, Tax. Untersuch. Gatt. Mammill. : 141. Typus: Escobariopsis herrerae (Werderm.) Doweld. Escobariopsis sect. Polycentracantha Doweld, sect. nov. - Plantae globulosae, hemilactescentes, aculeis radialibus 20-50 (5 mulptiplibus), acu/eis centralibus (plerumque 4 vel duplicibus 8). - Typus: Escobariopsis pennispinosa (Krainz) Doweld. Escobariopsis ser. Pennispinosae r;>oweld, seT. nov.- A ser. Sphacelato, Lasiacantho Herreroque aculeis centralibus pluribus (4-8) differt, sed nolae supra indicatae affinitatem proximam series nostri cum Escobariopsido monstrant. - Typus: Escobariopsis pennispinosa (Krainz) Doweld. KRAINZIA The delimitation of the species of polyphyletic genus Mammillaria s.l. necessitates the transference of the species, formerly recognized as distinct series Longiflorae, Stylothelae, Leptoc/adodae, and Decipientes, into a single distinct generic entity. The most earlier generic name for that group of species is Krainzia Backeb. (1938), and although only two species were usually recognized in the genus since Backeberg's times, and the most of names are available for later validly published genus Ebnerella F. Buxb. (195 I), I prefer to amplify the genus Krainzia by including all species of series cited, because of confused limits of Ebnerella and later proposal of Buxbaum (1954) to regard it a junior synonym of his polyphyletic Chilita Orcutt emend. F. Buxb. Four sections of Krainzia are validated, pending further revision of the series Stylothelae, which is seems to be heterogeneous, but provisionally accepted here as circumscribed by LUthy (1995). Krainzia bocasana (Poselger) Poweld, comb. nov. =T Mammillaria bocasana Poselger, 1853, Allg. Gartenzeit. 21 : 94. - Neotypus (Reppenhagen, 1991, Gatt. Mamm. I : 204): Reppenhagen 1232 (KL). Krainzia bocasana (Poselger) Doweld ssp. eschauzieri (Coulter) Doweld, comb. et stat. nov. =T Cactus eschauzieri Coulter, 1894, Contr. US Natl. Herb. 3, 2 : 104. - Neotypus Cw. A. Maurice & B. F. Maurice, 1995, J. Mamm. Soc. 35, I : 17): W A. Maurice et B. F Maurice 2186 (herb. CANTE). Krainzia crinita (DC.) Doweld, comb. nov. =T Mammillaria crinita DC. 1828, Mem. Mus. Hist. Nat. Paris, 17 : 112. - Lectotypus (Mottram, 1980, Mamm. Index: 21 as 'typus'): tab. 3 in DC. 1834, Mem. Cact.: 7. Epitypus (Reppenhagen, 1991, Gatt, Mamm. 1 : 240 as 'neotypus'): Reppenhagen 1704 (KL). Krainzia crinita (DC.) Doweld ssp. leucantha (Bod.) Doweld, comb. et stat. nov. =T Mammillaria leucantha Bod. 1933, Kakteenk. 1933, 1 : 233. - Lectotypus (Mottram, 1980, Mamm. Index: 47 as 'typus'): fig. [unnumbered] in Bod. I.e. Krainzia crinita (DC.) Doweld ssp. wildii (A. Dietrich) Doweld, comb. et stat. nov. =T Mammillaria wildii A. Dietrich, 1836, Allg. Gartenzeit. 4 : 137. - Neotypus (required). Krainzia decipiens (Scheidw.) Doweld, comb. nov. =T Mammillaria decipiens Scheidw. 1839, Bull. Acad. Sci. Brux. 5 : 496. - Neotypus (required). Krainzia decipiens (Scheidw.) Doweld ssp. albescens (Tiegel) Doweld, comb. et stat. nov. =T Mammillaria a/bescens Tiegel, 1933, Moller's Deutsch. Gartn.-Zeit. 48, 22 : 260. - Neotypus (required). 42 CYKKYJlEHTbJ / SUCCULENTS (TOM \ vol III) 2000 N2 1-2 Krainzia decipiens (Scheidw.) Doweld ssp. camptotricha (Dams) Doweld, comb. et stat. nov. =T Mammillaria camptotricha Dams, 1905, Gartenwelt, 10 : 14. - Neotypus (Mottram, 1980, Mamm. Index: 12 as' lectotypus '): J N. Rose & J H. Painter 11536 (US). Krainzia densilpina (Coulter) Doweld, comb. nov. =T Cactus densispinus Coulter, 1894, Contr. US Nat!. Herb. 3, 2 : 96. - Holotypus: Eschanzier S.n. (F). Krainzia eiongata (DC.) Doweld, comb. nov. =T Mammillaria elongata DC. 1828, Mem. Mus. Hist. Nat. Paris, 17: 109. - Neotypus (Reppenhagen, 1991, Gatt. Mamm. 1 : 338): Reppenhagen 1174 (KL). Krainzia erythrosperma (Bod.) Doweld, comb. nov. =T Mammillaria erythrosperma Bod. 1918, Monatsschr. Kakteenk. 28 : 101. - Lectotypus (designated here): fig. [unnumbered, right] in Bod. I.c. : 103. Epitypus (Reppenhagen, 1991, Gatt. Mamm. I : 202 as 'neotypus '): Reppenhagen 1239 (KL). Krainzia longiJlora (Britt. & Rose) Backeb. ssp. stamp/eri (Reppenhagen) Doweld, comb. et stat. nov. =T Mammillaria stampleri Reppenhagen, 1979, Kakteen Sukk. 30, 8 : 187. - Holotypus: Reppenhagen /358 (ZSS). Krainzia microhelia (Werderm.) Doweld, comb. nov. =T Mammillaria microhelia Werderm. 1930, Monatsschr. Deutsch. Kakt. Ges. 2 : 236. - Holotypus: Singer s.n. (B). Krainzia pottsii (Scheer ex Salm-Dyck) Doweld, comb. nov. =T Mammillaria pottsii Scheer ex Salm-Dyck, 1850, Cact. Hort. Dyck. : 104. - Lectotypus (L. Bens. 1982, Cacti US Can. : 967): ex coll. Dyck. Jan. 1857 (MO). Krainzia schwarzii (Shurly) Doweld, comb. nov. =T Mammillaria schwarzii Shuriy, 1949, Cact. Succ. J. Grt. Brit. 11, 1 : 17.- Holotypus: F Schwarz s.n. (K). Krainzia tepexicensis (Meyrcin) Doweld, comb. nov. =T Mammillaria tepexicensis Meyran, 1991, Cact. Suc. Mex. 36, 3 : 62-64. - Holotypus: F Otero 177 (MEXU). Validation of Supraspecific taxa: Krainzia sect. Stylothelae (Pfeiff.) Doweld, stat. et comb. nov. =T Mammillaria § Stylothelae Pfeiff. 1837, Enum. Cact. : 36. Typus: Krainzia crinita (DC.) Doweld. Krainzia sect. Leptociadodae (Lem.) Doweld, stat. et comb. nov. =T Mammillaria § Leptocladodae Lem. 1839, Cact. Gen. Nov. Sp. Nov. : 100. Typus: Krainzia elongata (DC.) Doweld. Krainzia sect. Decipientes (D. R. Hunt) Doweld, stat. et comb. nov. =T Mammillaria ser. Decipientes D. R. Hunt, 1979, Cact. Succ. J. Grt. Brit. 41, 4: 95. Typus: Krainzia decipiens (Scheidw.) Doweld. References 1. 2. 3. 4. 5. 6. B a c k e b erg C. Cactaceae Lindley. Systematic synopsis II BIatt. Kakteenforsch: 1938. Jahrg. 5. Pt. 6. (unpaged). Systematische Ubersicht (Neubearbeitung) mit BeschreibungschlUssel B a c k e b erg C. Cactaceae ｌｩｮ､ｬ･ｹｾ＠ II Cactaceae (Berlin). 1942. Jahrg. 1941. H. 2. S. 1-72. B u x b a urn F. Die Phylogenie der nordamerikanischen Echinocacteen. Trib. Euechinocactineae F. Buxb. II Osterr. Bot. Zeitschr. 1951. Bd. 98, H. 1-2. S. 44-104. B u x b a urn F. Die Gattungen der "Mammillaria-Stufe" III: Chilita (Orcutt 1926) emend. F. Buxbaum (Syn.: Ebnerella F. Buxb.) II Sukkulentenkunde. 1954. Bd 5. S. 3-33. Dowel d A. B. (,[( 0 Y 3 n b tJ. A. E.) Nomenclatural adjustments in Cacteae (Cactoideae-Cactaceae) II CYI<KYneHTbl. 1999. Ng 1. C. 27-31. LU thy J. M. Taxonomische Untersuchung der Gattung Mammillaria (Cactaceae). [Germany], 1995.230 S. 43