THE LAURACEOUS COLLECTIONS OF
CARL FRIEDRICH PHILIPP VON MARTIUS
IN THE FLORA BRASILIENSIS
peDro luíS roDriGueS De moraeS1 anD anDerSon falcaDe1
Abstract. This study refers to specimens of Lauraceae collected in Brazil by Carl Friedrich Philipp von Martius between 1817 and
1820. He collected 57 currently accepted species of Lauraceae, which belong to nine genera: Aniba, Cassytha, Cinnamomum, Cryptocarya, Dicypellium, Licaria, Nectandra, Ocotea, and Persea, for which Martius is known to have made about 99 different collections.
The vast majority of them are nomenclatural types and, taking into account the synonyms involved, encompassing 26 holotypes, 23 lectotypes, and 30 syntypes. Besides Martius himself, early descriptions and citations of these materials were made by Nees von Esenbeck,
Meissner, and Mez, the irst monographers of the family. Martius’s collections were the basis for the Flora Brasiliensis, where the taxonomic treatment of Lauraceae was made by the Swiss botanist Carl Daniel Friedrich Meissner in 1866, being of particular interest for
indicating all the collections examined and available at that time. Here, we provide an annotated and updated list of 60 entries with information on the status of the names and their types, currently accepted names, collections found in different herbaria, accompanied by an
Index to Scientiic names and an Index to Collections. Although there are no cases of specimens of Lauraceae collected by Martius from
his “Martii Herbarium Florae Brasiliensis,” they are also listed here since they have been attributed to him in literature. Lectotypes for
two names, Cryptocarya speciosa, and Nectandra nitidula var. maior, are designated, and a new synonymy is proposed.
Resumo. O presente estudo refere-se aos espécimes de Lauraceae que Carl Friedrich Philipp von Martius coletou no Brasil entre 1817
e 1820. Ele coletou 57 espécies atualmente aceitas de Lauraceae, pertencentes a nove gêneros: Aniba, Cassytha, Cinnamomum, Cryptocarya, Dicypellium, Licaria, Nectandra, Ocotea, e Persea, para os quais Martius é sabido de ter feito cerca de 99 coletas diferentes. A
grande maioria das mesmas são tipos nomenclaturais e, levando-se em consideração os sinônimos envolvidos, englobando 26 holótipos,
23 lectótipos e 30 síntipos. Além do próprio Martius, as primeiras descrições e citações desses materiais foram feitas por Nees von Esenbeck, Meissner, e Mez, que foram os primeiros monograistas da família. As coleções de Martius serviram de base para a Flora Brasiliensis, onde o tratamento taxonômico das Lauraceae icou a cargo do botânico Suíço Carl Daniel Friedrich Meissner em 1866, sendo de
particular interesse por indicar todas as coleções examinadas e disponíveis à epoca. Aqui, apresentamos uma lista anotada e atualizada
de 60 entradas com informações sobre o estado dos nomes e de seus tipos, nomes atualmente aceitos, coleções dos diferentes herbários,
acompanhadas de um índice de Nomes Cientíicos e de um índice de Coleções. Embora não existam casos de espécimes de Lauraceae
oriundos do “Martii Herbarium Florae Brasiliensis,” que tenham sido coletados por Martius, esses também são aqui listados por terem
sido atribuídos a ele na literatura. Lectótipos de dois nomes, Cryptocarya speciosa, e Nectandra nitidula var. maior, são designados,
assim como uma sinonímia nova é proposta.
Keywords: Brazil, Herbarium Florae Brasiliensis, historical collections, nomenclature, taxonomy
Carl Friedrich Philipp von Martius was born on the 17th
of April 1794 in Erlangen, Germany, and died in Munich
on the 13th of December 1868. He is one of the most
famous naturalists of the 19th century, having his name in
close association with Brazil and its lora. In 1817, King
Maximilian Joseph I of Bavaria, took the occasion of the
marriage of the Archduchess Caroline Josepha Leopoldine
von Habsburg-Lothringen with the Crown Prince, Dom
Pedro de Alcântara, the future Emperor of Brazil, to send the
zoologist Johann Baptist Spix and the botanist Carl Martius
to join the Austrian scientiic commission that was organized
to accompany the entourage of the Archduchess on her
departure to Brazil. These [and many other] scientists came
to explore the Brazilian Kingdom, making collections of
animals, plants, and minerals, investigating the productions
of the countries visited, the culture, the inhabitants, etc.
(Papavero, 1971). Martius arrived in Rio de Janeiro on the
frigate “Austria” on the 15th of July 1817, accompanied by
the Austrian entomologist and botanist Johann Christian
Mikán, and his wife, along with the Austrian painter
Thomas Ender, and Spix (Martius and Rodrigues, 1956:
433; Papavero, 1971: 61). After some weeks in Rio, Spix
and Martius decided to initiate their voyage through Brazil.
Spix and Martius collected in Brazil between 1817 and
1820. Detailed information about their itineraries and the
places visited can be found in Spix and Martius (1823, 1828,
1831a, 1831b, 1968), Urban (1906), Papavero (1971), and
in Tiefenbacher (1983). The journey itself began on the 8th
of December 1817, with São Paulo as the irst destination,
reaching it by the end of the year (31st of December). In
1818, they left São Paulo for southern Bahia, gathering large
collections especially in Minas Gerais and Bahia, where they
arrived in November 1818. For a few months they collected
in the northeastern states of Bahia, Pernambuco, Piauí and
Maranhão, after which embarking for Belém in July 1819.
The collecting followed by the Ilha de Marajó and other
Our thanks to Henk van der Werff (MO), for communicating the specimens on loan to him and for sending their images, and to Katharine E.S. Donahue
(History & Special Collections, Louise M. Darling Biomedical Library, UCLA) for the help with the requested manuscript of Martius. Pedro L.R. de
Moraes is supported by a grant (PQ2) from CNPq, and is grateful for the sponsorship received by the CAPES/DAAD Agreement for the years 2003, 2005,
and 2008; a grant of FUNDUNESP (Proc. Nr. 00073/11-DFP), for his visit to MEL in 2011; stipends from the Belgian Focal Point to the Global Taxonomy
Initiative (GTI), which supported the visit to the Botanic Garden Meise in 2004 and 2012; and a sponsorship from CNPq (Proc. Nr. 450515/2013-3) for a
visit to LE.
1
Universidade Estadual Paulista “Júlio de Mesquita Filho”, Instituto de Biociências, Departamento de Botânica, Av. 24 A 1515, Bela Vista, Caixa Postal
199, 13506-900 Rio Claro, SP, Brazil. Email: pmoraes@rc.unesp.br. Author for correspondence.
Harvard Papers in Botany, Vol. 20, No. 2, 2015, pp. 167–197.
© President and Fellows of Harvard College, 2015
ISSN: 1938-2944, DOI: 10.3100/hpib.v20iss2.2015.n7, Published online: 31 December 2015
168
HARVARD PAPERS IN BOTANy
localities in the vicinity of Belém, Pará, then beginning
the inal stage of the expedition up the Rio Amazonas to
Manaus, where Spix and Martius followed separate paths:
the irst followed by the Rio Negro and its tributaries and
Martius followed the Rio Solimões and Rio Jupará. Both
returned to Belém in April 1820 and in June of that year,
embarked for Europe, arriving in Lisbon on the 23rd of
August, and inally in Munich in December 1820 (Spix and
Martius, 1823, 1828, 1831a; Urban, 1906; Shepherd, 2002).
The literature about the life and writings of Carl Martius
is quite extensive, given the amount and importance of his
work. Among the primary sources, we can highlight the
texts by Lasègue (1845), de Candolle (1856a, b, 1857),
Carus (1869), Eichler (1869a, c, d), Meissner (1869, 1870),
Seemann (1869), Schramm (1869a, b), Spring (1870,
1871), Rau (1871), Wunschmann (1884), and Ross (1917).
More recent literature and contextualized approaches
about Martius’s works can be found in Dutra (1942),
Hoehne (1942), Rambo (1952), Sommer (1953), Biblioteca
Nacional (1968), Staleu and Cowan (1981), Wuschek
(1989), Mägdefrau (1990), Förther (1994), and Lisboa
(1995, 1997).
The collections of Martius during the travel to Brazil
may be estimated at nearly 8,000 species, of which 800
were collected in port sites of Europe in Istria, Malta,
Gibraltar, Madeira, and Portugal, at the beginning and the
end of the travel (Martius, 1850; Wallich, 1851; Förther,
1994) (c. 20,000 to 25,000 specimens), thus becoming the
foundation for the production of the Flora Brasiliensis.
Martius originally edited this monumental work from 1840
to 1868, with the printing of 46 fascicles, where more than
about 9,000 species were treated, illustrated in about 1,400
igures and 1,071 lithographs. The edition of the Flora was
continued posthumously by August Wilhelm Eichler (1839–
1887) and Ignatz Urban (1848–1931), with completion of
the work in 1906. The Flora Brasiliensis in its inal form
consists of 15 volumes (in 40 parts) in folio, originally
published in 130 individual fascicles (in 130 dated parts),
the text consisting of 20,733 “pages” (the page numbers
actually refer to columns, two on each page), and 3,811
lithographs and nature-prints, illustrating 6,246 species
in full size. Given the magnitude of the undertaking, the
work had the collaboration of 65 researchers including the
most illustrious European botanists of the time, who have
described a total of 22,767 species (Urban, 1908; Martius
and Rodrigues, 1956: 434; Baldus in Spix and Martius,
1968: 8; Shepherd, 2002). The majority of those species are
angiosperms, of which 19,629 are native to Brazil and 3,138
are from adjacent regions or introduced in the country.
The treatment of Lauraceae in the Flora Brasiliensis was
made by the Swiss botanist Carl Daniel Friedrich Meissner
(1800–1874), which was published in Volume 5, Part 2, in
1866, with the description of 327 species. Among them,
there are 64 taxa with the indication of collections made by
or attributed to Martius.
During the expedition to Brazil, Martius made quick
notes in situ for 3,320 species of plants, in his “Adversaria
Botanica” (Martius, 1850: 723; Wallich, 1851: 68; Förther,
1994). Later, in his writings, he refers to this unpublished
manuscript again. A handwritten copy made by A. Progel,
VOL. 20, NO. 2
entitled “Observationes botanicae, Plantae in itinere
Brasiliensi annis 1817–1820 a Car. Frid. Phil. Martio
descriptae, vol. 1–6,” bound in six volumes in quarto, is
housed at the Library of the Botanische Staatssammlung
München (Martius, 1817–1820). A xerographic copy of
these volumes is also deposited in the Library of the Royal
Botanic Gardens, Kew. Of key importance is the fact that
many of the specimens collected by Martius have labels
that refer to those comments, providing an additional
source of information on the species (see Förther, 1994).
For Lauraceae, all these cases have been registered here and
referred to as “Martius Obs.,” followed by their numbers.
Additionally, as pointed out by Förther (1994: 12),
Martius received many collections sent from Brazil, in
between the end of 1830 and the end of 1840, made by
Ackermann (gathered in 1830–1836), P. Claussen (gathered
in 1834–1843), B. Luschnath (gathered in 1831–1837),
A. L. P. da Silva Manso (gathered ca. 1830–1832), and L.
Riedel (gathered ca. 1821–1836), which were distributed by
him to major European herbaria (e.g., B, BM, G, L, LE, M,
W, etc.). Among them, there were a series of new species
that Martius entitled as “Herbarium Florae Brasiliensis,”
being published in the journal Flora between 1837 and
1841 (Martius, 1837–1841). These exsiccatae distributed
to herbaria bear printed labels with “Martii Herbar.
Florae Brasil. Nº,” followed by a handwritten number,
corresponding to what has been published in Flora. Although
for Lauraceae there are no cases where the species were
collected by Martius, often in the literature these collections
were assigned to him. For this reason, they have been also
listed in this work and, when known, their collectors are
indicated. As pointed out by Hind (2011), the specimens
at herbarium BR usually provide evidence identifying the
collectors, since they bear both the labels of the original
collector and of the “Herb. Fl. Bras.”
Martius’s own collections from Brazil are held in the
Botanische Staatssammlung München (M), with duplicates
at B, BM, BR, G-DC, K, L, LE, P, and W (Urban, 1906;
Förther, 1994). Martius’s private herbarium (Herbarium
Martii), estimated to comprise about 300,000 specimens,
containing 60,000 species, nearly half of which are Brazilian,
was purchased by the Belgian Government for a sum of
£1,200, in 1870, to form the basis of a national collection,
and is now at the Botanic Garden Meise (BR) (see Eichler,
1869b; Bommer, 1870; Seemann, 1870; Staleu and Cowan,
1981; Förther, 1994). In Förther (1994), a detailed account
on the development and content of Martius’s private
herbarium and his numerous, fruitless attempts to sell
his collections to the Bavarian Government is presented.
According to Bommer (1870), these efforts of selling the
private herbarium to the Bavarian Kings, Maximilian II
and his successor Ludwig I, were unsuccessful because
Bavaria already had the Brazilian herbarium formed by
Martius and Spix. However, Martius’s strained relationship
with King Maximilian II might help to partially explain
the king's refusal to buy the collections. The king had
refused to increase Martius’s starting salary since his initial
appointment in 1836 to succeed Schrank as the irst curator
of the Munich Botanical Garden (see Förther, 1994).
2015
DE MORAES AND FALCADE, LAURACEAE COLLECTED By MARTIUS
The aim of the present study was to review all Lauraceae
collections made by Martius in Brazil, updating and
commenting their taxonomic status, detailing and clarifying
the localities and dates of the collections, as well as
indicating all herbaria known to possess their duplicates.
Meissner’s treatment of the family in Martius’s Flora
Brasiliensis (Meissner, 1866) was taken as a key reference
for citing the material collected by Martius along with the
specimens examined and available from other collectors.
This study is based on herbarium material from the
herbaria A, AWH, B, BC, BM, BR, C, CGE, COL, E, F, FI,
G, G-DC, GH, GOET, GZU, HAL, HBG, HRCB, JE, K,
KIEL, KR, L, LE, LINN, LISU, LZ, M, MA, MEL, MO,
Ny, OUPR, OXF, P, S, SBT, STR, TO, TUB, U, US, VT,
W, and WRSL (herbarium acronyms according to Thiers,
[continuously updated]). “Original material,” indicated
within the text, relies on Art. 9.2 & 9.3 ICN (McNeill et al.,
2012). All species names and their taxonomic treatment were
checked in the classical taxonomic works on Lauraceae, such
as those by Nees von Esenbeck (1833, 1836, 1848), Meissner
(1864), and Mez (1889), as well as in the more recent published or available revisions: Aiouea Aubl. and Aniba Aubl.
(Kubitzki and Renner, 1982), Cassytha L. (Weber, 1981),
Cinnamomum Schaeff. (Lorea-Hernández, 1996), Cryptocarya R. Br. (Kostermans, 1937, 1938a; Moraes, 2005, 2007),
Dicypellium Nees & Mart., Phyllostemonodaphne Kosterm.,
Systemonodaphne Mez and Urbanodendron Mez (Rohwer,
1988), Endlicheria Nees (Chanderbali, 2004), Licaria Aubl.
(Kostermans, 1936; Kurz, 2000), Nectandra Rol. ex Rottb.
(Rohwer, 1993), Ocotea Aubl. (Rohwer, 1986), and Persea
Mill. (Kopp, 1966).
169
As results, we show below that Martius collected 57
currently accepted species of Lauraceae, 55 of which
are native to Brazil (not introduced or subspontaneous).
Taking into account the synonyms involved, among the 99
different collections gathered by Martius, 26 are holotypes,
23 are lectotypes, and 30 are syntypes of names of the
corresponding taxa, i.e., the vast majority are nomenclatural
types. These numbers alone would be enough to demonstrate
the importance of the Lauraceae collections that Martius
gathered in Brazil over the three years of his expedition with
Spix. However, when comparing the expedition of Spix and
Martius with what is known about those of other naturalists
who visited Brazil in the early 19th century (see Mez, 1892,
for a list of collectors and their numbered collections of
Lauraceae), it is evident that just Friedrich Sellow (1789–
1831) and Ludwig Riedel (1791–1861), who remained in
Brazil for much longer, would have surpassed Martius in
number of species collected for the family. Sellow collected
in Brazil, from 1814 to 1831, a total of 61 currently accepted
species of Lauraceae (Moraes, 2008). Riedel, in turn, came to
Brazil in 1821 and remained in the country until his death in
1861. His collection period was from 1821 to 1836, in which
he collected 92 species of Lauraceae (Moraes, 2012a, b).
For convenience, the current accepted names of species
are listed in alphabetic order. Accepted names appear in
boldface italics, synonyms in italics. Homotypic synonyms
are indicated by “≡,” while the heterotypic are under the
entries “Synonym” or “Synonyms.” Whenever available,
barcode numbers are quoted with the herbarium acronyms.
An Index to Scientiic Names and an Index to Collections
are provided at the end.
an annotateD liSt of lauraceae collecteD by martiuS or attributeD to him
Synonyms: Aiouea brasiliensis Meisn. var. attenuata Meisn.
1. Aiouea laevis (Mart.) Kosterm., Recueil Trav. Bot. Néerl.
in DC., Prodr. 15(1): 83. 1864; ibid. in Martius, Fl.
35: 84. 1938 ≡ Aiouea brasiliensis Meisn. in DC., Prodr.
Bras. 5(2): 170. 1866 ≡ Aiouea riedelii Mez, Jahrb.
15(1): 83. 1864, nom. illegit.; ibid. in Martius, Fl. Bras.
Königl. Bot. Gart. Berlin 5: 37. 1889, nom. illegit.
5(2): 170. 1866; Mez, Jahrb. Königl. Bot. Gart. Berlin 5:
superl. TyPE: BRAZIL. Loco haud indicato, s.d.,
43. 1889; Moraes, Harvard Pap. Bot. 17(1): 189. 2012; ibid.
L. Riedel s.n. (Holotype: LE; Isotypes, B† [F neg.
in Pl. Ecol. Evol. 146(3): 361. 2013.
3788], G 00020674, GOET [2 sheets], K 000601965,
Basionym: Cryptocarya laevis Nees ex Mart., Flora 21(2,
L 0035527, LE [2 sheets], Ny 00354799). Specimen:
Beibl.): 64. 1838; Meissner in DC., Prodr. 15(1):
BRAZIL. Bahia, Ilhéus, “ad ripas luvi Itahype” [Rio
83. 1864; ibid. in Martius, Fl. Bras. 5(2): 170. 1866;
Itaípe], March 1822, L. Riedel 688 (LE n.v.).
Renner, Fl. Neotrop. Monogr. 31: 109. 1982. TyPE:
For a complete list of synonyms, see Kubitzki and
BRAZIL. Rio de Janeiro, “Crescit in sylvis Caa-poera
Renner (1982).
dictis ad Sebastianop.,” 12 June 1817, Martii Herbar.
None of the located specimens of Martii Herbar. Florae
Florae Brasil. Nº 237 (Holotype: BR 0000008686169
Brasil. Nº 237 bears information on who the collector is,
[annotated by Nees as “Cryptocarya pyriformis levis
even in BR. Nonetheless, it has been wrongly attributed to
N. ab. E.”]; Isotypes, AWH 28362, AWH 28363, B† [F
Martius by Kubitzki and Renner (1982). In fact, once Martius
neg. 3782], BM 000894059, BR 0000008685834, E
(1838) had indicated the gathering as from June, in Rio de
00259412, FI 005252, G, G 00021045, G 00021046,
Janeiro, this excluded him as the collector, since he has
G 00021047, G-DC 00200460, GH 00041126, HAL
arrived in Rio de Janeiro in July 1817. From information on
0101918, JE 00004805 [and probably JE 00000523,
the sites and dates of the collections reported by Martius in
ex Herbar. Bernhardi, Anonymous s.n.], K 000512712,
Herbarium Florae Brasiliensis, and based on the respective
K 000512713, K 000512714, KR, L 0035524, L
specimens located in BR, the most likely collector of the
0035525, LE 00000096, LE 00000135, LZ†, M
present species would be Bernhard Luschnath. As pointed
0111049, M 0111050, M 0111051, M 0111052, MO
out by Moraes (2012: 189), in Martius’s Flora Brasiliensis,
247250, Ny 00354800, P 00128383, P 00128384, W†,
Meissner (1866: 170) mistakenly attributed to Martius the
collection of Riedel 688 from Rio Itaípe.
WRSL).
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HARVARD PAPERS IN BOTANy
2. Aniba canelilla (Kunth) Mez, Jahrb. Königl. Bot. Gart.
Berlin 5: 53. 1889; Kostermans, Recueil Trav. Bot. Néerl.
35: 898. 1938; Kubitzki, Fl. Neotrop. Monogr. 31: 79. 1982;
Rohwer, Mitt. Inst. Allg. Bot. Hamburg 20: 13, 111. 1986;
Moraes, Harvard Pap. Bot. 17(1): 190. 2012.
Basionym: Cryptocarya canelilla Kunth in Humboldt,
Bonpland and Kunth, Nov. Gen. Sp. (quarto ed.)
7: 192, t. 645. 1825. Lectotype (designated by
Kostermans, 1938b: 899): VENEZUELA. “crescit
ad orientem Esmeraldæ, in monte de la Canelilla,
juxta ripam Cauræ et erevati; item prope catarractam
Maypurensium,” s.d., immat. fr., F. W. H. A. von
Humboldt & A. J. A. Bonpland 1018 (B-W 07784
00 0 & B-W 07784 01 0 [“Laurus canelilla Willd.,”
herbarium name, “Orinoco”]; Isolectotypes, P
00307242 [F neg. 35005], P 00506929).
Synonyms: Cryptocarya pretiosa Mart. in Buchner, Repert.
Pharm. 31: 356. 1829, syn. nov. ≡ Mespilodaphne
pretiosa Nees & Mart., Linnaea 8: 45. 1833, nom.
illegit. superl.; Martius, Syst. Mat. Med. Bras.: 111.
1843; ibid. in Fl. Bras. 5(2): 317. 1868 ≡ Mespilodaphne
pretiosa Nees & Mart. var. angustifolia Nees, Syst.
Laur.: 237. 1836, nom. invalid.; Meissner in DC.,
Prodr. 15(1): 103. 1864; ibid. in Martius, Fl. Bras. 5(2):
198–199. 1866; Moraes, Komarovia 6(1): 16. 2008.
LECTOTyPE (designated here): BRAZIL. Pará, “in
sylvis ad oppidum Pará rarius, frequentius in Provinciae
Paraensis interioribus,” 1819, immat. fr., fr., C. F. P.
von Martius s.n. (M 0121069 [“Cryptocarya pretiosa
Mart.,” “Páo v. Casca preciosa Lusitan.”]). Original
material: BRAZIL. Pará, s.d., immat. fr., C. F. P. von
Martius s.n. (M 01121070 [fragment], M 0121071,
M 0121072, MEL 2318499); idem, “ad luvium Rio
Negro,” s.d., immat. fr., C. F. P. von Martius s.n. (G,
MEL 2318499); idem, Amazonas, “inveni prope
praedium Capitany in prov. Alto Amaz.,” 1819, immat.
fr., C. F. P. von Martius s.n. (BR 0000008685865
[“Laurus pretiosa,” herbarium name]).
For a complete list of synonyms, see Kubitzki and
Renner (1982).
Vernacular names: “Casca preciosa” (Martius, 1829),
“Pereiorá Tupin.,” “Páo v. Casca pretiosa” (Martius, 1843,
1868).
Hitherto, the name Cryptocarya pretiosa Mart. has
been interpreted as a herbarium name mentioned in the
synonymy of Mespilodaphne pretiosa Nees & Mart. var.
angustifolia Nees as “Cryptocarya pretiosa Mart. in Herb.
Brasil. Monac.” (Nees von Esenbeck, 1836), therefore being
overlooked by Nees as already published by Martius (1829).
Moraes (2008) pointed out that, as indicated by Rohwer
(1986: 13, 111, 229), the name Mespilodaphne pretiosa
Nees & Mart. is superluous, thus illegitimate, since
Laurus quixos Lam., Encycl. 3: 455. 1792 was cited as a
synonym {≡ Ocotea quixos (Lam.) Kosterm., Recueil
Trav. Bot. Néerl. 35: 900. 1938. TyPE: [ECUADOR] “ex
Provincia de los Canelos in Peru,” “Provincia Riobaruba,”
12 April 1748, sterile, Joseph de Jussieu s.n. (Holotype:
P-JU 00307277 [Cat. No. 4245]); Isotype, P-LA 00381527,
sterile.; see Rohwer, 1986: 229, 249; 1993: 297–298])}.
VOL. 20, NO. 2
The earlier name Laurus quixos was not included in a
subordinated taxon under Mespilodaphne pretiosa, so the
latter name should be authomatically typiied by the type
of the irst one (Art. 7.5). The name Ocotea pretiosa Mez,
Jahrb. Königl. Bot. Gart. Berlin 5: 250. 1889, based on
Mespilodaphne pretiosa Nees & Mart. with exclusion of its
variety angustifolia Nees (with Laurus quixos in synonymy)
is therefore legitimate. The former name in sense of Mez
is Ocotea odorifera (Vell.) Rohwer, Mitt. Inst. Allg.
Bot. Hamburg 20: 111. 1986. Another name, Aydendron
suaveolens Nees, Linnaea 8: 37. 1833, which was included
by Mez into the concept of Ocotea pretiosa does not make
the new name illegitimate because the combination Ocotea
suaveolens (Meisn.) Benth. ex Hieron., Bol. Acad. Nac.
Ci. 4: 437. 1881 (≡ Oreodaphne suaveolens Meisn. in DC.,
Prodr. 15(1): 136. 1864) was already published to that date.
The combination “Ocotea pretiosa Benth. et Hook. f.”
recorded in Index Kewensis has not been validly published
by Bentham in Bentham and Hooker (1880: 158), being
substituted by the phrase “species typica generis” without
any nomenclatural status under the ICN (Art. 35.2).
Martius (1843a), in his Systema materiae medicae
vegetabilis brasiliensis, cited the existence of several
lithograph plates that were already inished from his
book that was never published, “Plantae Medicae et
Oeconomicae Brasiliae” (Mart. Pl. med. et oec. Br. ined.),
with the plate 33 being indicated for Mespilodaphne
pretiosa Nees & Mart. Meissner (1866: 301–302) also
mentioned this unpublished book as “Tabulas LXVI. LXX.
XCV. XCVI. CI. CII. receptae sunt e Martii opere inedito:
Plantae medicinales et oeconomicae Brasilienses” and
used those Martius’s plates in his monograph, which were
acknowledged as “Tabula nostra […] ex Martii Pl. med. et
oec. Bras. ined. recepta” (for Aydendron cujumary [Tabula
LXVI], Mespilodaphne opifera [LXX], Nectandra puchurymajor [XCV], Nectandra cymbarum [XCVI], Nectandra
puchury-minor [CI], and Dicypellium caryophyllaceum
[CII]). However, for “Mespilodaphne pretiosa Meissn.,”
Meissner only quoted “Tabula nostra LXXIV,” therefore
evidencing that another plate was prepared for the species
in question. Actually, plate 74 in Flora Brasiliensis refers
to Ocotea odorifera (Vell.) Rohwer, which most likely was
based on Riedel’s collection from “Mandiocca,” since it is
the only lowering material among the collections cited by
Meissner (see Moraes, 2012b).
3. Aniba citrifolia (Nees) Mez, Jahrb. Königl. Bot. Gart.
Berlin 5: 74. 1889; Kostermans, Recueil Trav. Bot. Néerl.
35: 890. 1938; Kubitzki, Fl. Neotrop. Monogr. 31: 23. 1982;
Moraes, Harvard Pap. Bot. 17(1): 191. 2012.
Basionym: Aydendron citrifolium Nees, Syst. Laur.: 257.
1836; Meissner in DC., Prodr. 15(1): 93. 1864; ibid. in
Martius, Fl. Bras. 5(2): 183. 1866. TyPE: BRAZIL.
Amazonas, “Rio Negro” (Alto Amazonas), “In sylvis
Japurensibus, ad amnem Juì,” February 1820, C. F.
P. von Martius s.n. (Holotype: M 0147109 [F neg.
6583]).
Additional synonyms in Kubitzki and Renner (1982) and
Moraes (2012).
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DE MORAES AND FALCADE, LAURACEAE COLLECTED By MARTIUS
4. Aniba desertorum (Nees) Mez, Jahrb. Königl. Bot. Gart.
Berlin 5: 76. 1889; Kostermans, Recueil Trav. Bot. Néerl.
35: 892. 1938; Kubitzki, Fl. Neotrop. Monogr. 31: 38. 1982;
Rohwer, Mitt. Inst. Allg. Bot. Hamburg 20: 247. 1986.
Basionym: Aydendron desertorum Nees, Syst. Laur.: 260.
1836; Meissner in DC., Prodr. 15(1): 92. 1864; ibid.
in Martius, Fl. Bras. 5(2): 182, t. 65. 1866 ≡ ‘Ocotea
desertorum Mart.,’ nom. nud. ≡ ‘Ocotea obtusifolia
Mart.,’ nom. nud. Lectotype (designated by Kubitzki
and Renner 1982: 38): BRAZIL. Minas Gerais,
“in sylvis Capões ad luv. Rio Verde pequeno et in
Chapada do Paranan,” July 1818, l., fr., C. F. P. von
Martius s.n. (M 0147106; Isolectotypes, M 0147104
[ex Herb. Zuccarinii], M 0147105, M 0147108).
Syntype: BRAZIL. Minas Gerais, Rio São Francisco,
“in sylvis capões versus f. S. Francisci,” August 1818,
fr., C. F. P. von Martius s.n. (M 0147107).
For a complete list of synonyms, see Kubitzki and Renner
(1982).
5. Aniba permollis (Nees) Mez, Jahrb. Königl. Bot. Gart.
Berlin 5: 55. 1889; Kostermans, Recueil Trav. Bot. Néerl.
35: 896. 1938; Kubitzki, Fl. Neotrop. Monogr. 31: 73. 1982.
Basionym: Aydendron permolle Nees, Syst. Laur.: 246.
1836; Meissner in DC., Prodr. 15(1): 88. 1864; ibid.
in Martius, Fl. Bras. 5(2): 176. 1866 ≡ ‘Cryptocarya
aü-üva Mart. ex Nees,’ Syst. Laur.: 246. 1836, nom.
invalid. Lectotype (designated by Kostermans,
1938b: 896): BRAZIL. Amazonas, Tefé, “in sylvis
ad Ega,” December 1819, C. F. P. von Martius Obs.
[Observationes botanicae] 2908: “Laurus Aüiuva
/ Arbor magna frondosa. Rami teretes tomento
ferruginei. …” (M 0147098; Isolectotypes, B† [F
neg. 3806], M 0147099, M 0147100, Ny 00354890
[fragment], Ny 00354891 [fragment]). Syntype:
BRAZIL. Amazonas, Tefé, “Ega,” November 1831,
l., E. F. Poeppig 2769 (B†, BM, F 0061359F ex
W, G [2 sheets], G-DC, HAL 0103802, KIEL, LE
00006806, LE, LZ†, M, OXF 00000388, P 00745599,
P 00745600, W, WRSL).
6. Aniba puchury-minor (Mart.) Mez, Jahrb. Königl. Bot.
Gart. Berlin 5: 70. 1889; Kostermans, Recueil Trav. Bot.
Néerl. 35: 903. 1938; Kubitzki, Fl. Neotrop. Monogr. 31:
57. 1982; Moraes, Pl. Ecol. Evol. 146(3): 362. 2013 ≡
Nectandra puchury-minor (Mart.) Nees, Syst. Laur.: 336.
1836; Martius, Syst. Mat. Med. Bras.: 109–110. 1843; ibid.
in Fl. Bras. 5(2): 317. 1868; Meissner in DC., Prodr. 15(1):
166. 1864; ibid. in Martius, Fl. Bras. 5(2): 277, t. 101. 1866.
Basionym: Ocotea puchury-minor Mart. in Buchner,
Repert. Pharm. 35: 172. 1830. Lectotype (designated
by Mez, 1889: 70): BRAZIL. Amazonas, “in sylvis
Japurensibus,” December 1819, fr., C. F. P. von
Martius s.n. (M 0113213; Isolectotype, B†). Syntypes:
BRAZIL. Amazonas, “in sylvis secus l. Solimões,”
November 1819, fr., C. F. P. Martius s.n. (GZU
000254329 [fragment], M 0112564); BRAZIL.
Amazonas, loco haud indicato, 1819, fr., C. F. P. von
Martius s.n. (M 0112563).
171
Synonymy in Kubitzki and Renner (1982) and Moraes
(2013).
Vernacular names: “Fabas Pechurim v. Puchury”
(Martius, 1868).
7. Cassytha iliformis L., Sp. Pl. 1: 35. 1753. Lectotype
(designated by Imkhanitskaya, 1974: 208): INDIA. P.
Osbeck s.n. (Herb. Linn No. 519.1, LINN). Original
material: P. Osbeck s.n. (S [05-4065]).
Synonyms: Cassytha brasiliensis Mart. ex Nees, Syst.
Laur.: 648. 1836; Mez, Jahrb. Königl. Bot. Gart.
Berlin 5: 490. 1889 ≡ Cassytha americana Nees
var. brasiliensis (Nees) Meisn. in DC., Prodr. 15(1):
256. 1864; ibid. in Martius, Fl. Bras. 5(2): 296. 1866.
TyPE: BRAZIL. Minas Gerais, “in Campis editis
Minarum novarum,” July–August 1818, C. F. P. von
Martius s.n. (Holotype: M).
Cassytha americana Nees var. puberula (Nees) Meisn.
in DC., Prodr. 15(1): 256. 1864; ibid. in Martius, Fl.
Bras. 5(2): 296. 1866. TyPE: BRAZIL. Bahia, C. F.
P. von Martius s.n. (Syntype: M); BRAZIL. Bahia, “in
collibus siccis pr. Ilheos. J. Domingo” [Fazenda João
Domingo], June 1821, L. Riedel 374 (Syntypes: LE,
Ny 00354928).
Synonyms and further information in Stapf (1912),
Kostermans (1950), and Weber (1981).
8. Cinnamomum erythropus (Nees & Mart.) Kosterm.,
Reinwardtia 6: 21. 1961; Lorea-Hernández (1996: 81);
Moraes, Harvard Pap. Bot. 17(1): 193. 2012 ≡ Phoebe
erythropus (Nees & Mart.) Mez, Jahrb. Königl. Bot. Gart.
Berlin 5: 189. 1889.
Basionym: Persea erythropus Nees & Mart. in Nees,
Linnaea 8: 49. 1833. Lectotype (designated by Moraes,
2008: 48): BRAZIL. Minas Gerais, Ouro Preto, “in
ferruginosis ad Antonio Pereira,” April 1818, l., C. F.
P. von Martius Obs. 883 (GZU 000249338 [fragment
on the top, annotated by Nees]; Isolectotypes, M
0147194, M 0147196 [“Persea erythropus M. Reise I.
p. 401. (nomen)”]). Syntypes: BRAZIL. Minas Gerais,
loco haud indicato, s.d., fr., C. F. P. von Martius s.n.
(M 0147195); BRAZIL. Loco haud indicato, l., F.
Sellow 1360 (B†, F [619716; fragment ex B†, F neg.
58035, Sellow s.n.], GZU 000265498 [fragment on
the bottom, annotated by Nees, Anonymous s.n.]).
Synonyms: Persea erythropus Nees & Mart. var. angustata
Meisn. in DC., Prodr. 15(1): 55. 1864 ≡ Persea
erythropus Nees & Mart. var. mixtinervia Meisn.
in Martius, Fl. Bras. 5(2): 162. 1866, nom. illegit.
superl. TyPE: BRAZIL. Minas Gerais, “Serra
da Piedade et S. Itacolomi,” October–November
1834, l., L. Riedel 2777 (Syntypes: C, G [2 sheets],
K 000602109, K 000602110, LE 00000480, LE
00000481, LE 00000482, Ny 00355827, OXF
00000375 [Anonymous s.n.], P 01978620, US
00099112); BRAZIL. Minas Gerais, Ouro Preto, “in
ferruginosis ad Ouro Preto,” “Serra de Ouro Preto,”
June 1839, l., fr., [Ackermann s.n.] P. Claussen 108
in Martii Herbar. Florae Brasil. Nº 1238 (Syntypes:
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HARVARD PAPERS IN BOTANy
BR 0000008762092, BR 0000008764317, G-DC
00200453, LE, Ny 00355828, P 00128549).
For a complete list of synonyms, see Lorea-Hernández
(1996) and Moraes (2008).
9. Cinnamomum verum J. Presl in Berchtold & J. Presl,
Přir. Rostlin 2: 36–37, t. 7. 1825 [1823?]; Moraes, Harvard
Pap. Bot. 17(1): 194. 2012; ibid. in Pl. Ecol. Evol. 146(3):
364. 2013. Specimen: BRAZIL. Rio de Janeiro, “in
horto (Chacara) secundum viam ad S. Clemente, prope
Sebastianopolin, cultum,” July 1833, l., November–
December 1833, fr., B. Luschnath s.n. in Martii Herbar.
Florae Brasil. Nº 477 (BR 0000013332587, M 0147231).
Basionym: Laurus cinnamomum L., Sp. Pl. 1: 369. 1753.
Lectotype (designated by Wuu-Kuang, 2011:
261): Herb. Clifford: 154, Laurus 6, sheet B (BM
000558701).
Synonyms: Cinnamomum zeylanicum Breyn., Ephem. Nat.
Cur. Dec. I, Ann. 4: 139. 1676, nom. invalid. (see
Howard, 1981).
Cinnamomum zeylanicum Blume; Bijdr. Fl. Ned. Ind.
11: 568. 1825; Nees in Nees and Th. Nees, Cinnam.
Disp.: 48. 1823; Nees in Wallich, Pl. Asiat. Rar. 2: 74.
1831; Nees, Syst. Laur.: 45. 1836; Meissner in DC.,
Prodr. 15(1): 13. 1864; ibid. in Martius, Fl. Bras. 5(2):
147. 1866. Specimen: BRAZIL. Rio de Janeiro, “...
haec est ex horto botanico prope Rio de J.o Provinciae
ejusdem nominis”, 1817, l., C. F. P. von Martius s.n.
(BR, M 0147232).
Cinnamomum zeylanicum Blume var. microphyllum
Nees, Syst. Laur.: 47. 1836; Meissner in DC., Prodr.
15(1): 14. 1864; ibid. in Martius, Fl. Bras. 5(2): 147.
1866. Specimen: BRAZIL. Pará, “in cultis ad Pará”,
l., C. F. P. von Martius s.n. (L 0036030 [“Laurus
cassioides Mart.,” herbarium name, “Pará”; ex
Herbarium Reinwardtianum], M 0147236) [‘Laurus
cassioides Mart. ex Miq.’, Ann. Mus. Bot. LugdunoBatavi 1: 256. 1864, nom. invalid.].
For a complete list of synonyms, see Kostermans (1964,
1986).
The specimen cultivated in Pará that Martius collected
was cited by Nees von Esenbeck (1836: 48), Miquel (1864),
and Meissner (1866: 147).
10. Cryptocarya mandioccana Meisn. in DC., Prodr. 15(1):
75. 1864; ibid. in Martius, Fl. Bras. 5(2): 165. 1866; Mez,
Jahrb. Königl. Bot. Gart. Berlin 5: 10. 1889; Kostermans,
Recueil Trav. Bot. Néerl. 34: 572. 1937; ibid. in Recueil
Trav. Bot. Néerl. 35: 125–126. 1938; Moraes, Taxon
54(3): 792. 2005; ibid. in Abc Taxa 3: 70. 2007; ibid. in
Komarovia 6(1): 8. 2008; ibid. in Harvard Pap. Bot. 17(1):
194. 2012. TyPE: BRAZIL. Rio de Janeiro, Magé, “in
sylvis Mandiocensis,” October 1823, l., L. Riedel s.n.
(Holotype: LE; Isotypes, B† [F neg. 3844], GOET 007344,
GOET 007345, K 000601800, L 0036185, L 0036186, LE,
Ny 00355045, OXF 00057026, OXF 00057027 [righthand specimen], U, US 00051077). Specimen: BRAZIL.
Minas Gerais, Visconde do Rio Branco, “habitat in sylvis
aboriginibus ad Praesid. Joannis Bapt. et alibi. Provinciae
VOL. 20, NO. 2
Minar.,” April 1818, fr., C. F. P. von Martius Obs. 1095 b:
“Noz moschada do Brazil. Litsaeae sp. ? Arbor alta trunco
recto, cortice 2–3 lin. crasso castâneo epidermide laevigata
cinereo-subrufesc. laevia absque rimis, ligno albido, ibra
dura compacta, speculis transversal. minutis, gravi. …” (M
0152460 [syntype of Cryptocarya moschata Nees & Mart.
in Nees, Linnaea 8: 37. 1833; Martius in Spix and Martius,
Reise Bras. 2: 543. 1828; ibid. in Syst. Mat. Med. Bras.:
110. 1843; ibid. in Fl. Bras. 5(2): 319. 1868; Meissner in
DC., Prodr. 15(1): 74. 1864; ibid. in Martius, Fl. Bras. 5(2):
164, t. 56. 1866]).
Further information in Moraes (2005, 2007).
Vernacular name: “Noz mocada do Brazil” (Martius,
1843, 1868).
Originally, Cryptocarya moschata was based on two
collections, a fruiting one by Martius and a lowering one by
Sellow. Unfortunately, these collections represent different
taxa, which had not been acknowledged either by Meissner
or by Mez. The lectotypiication of C. moschata in the sense
of the Martius collection was done by Kostermans (1937).
He proposed the specimen of Martius as (lecto)type, without
analyzing it, based on the description of Nees von Esenbeck
and solely on the available specimen of Sellow in hand.
Later (Kostermans, 1938a), after studying the specimen
of Martius, he was “obliged to alter the synonymy” of C.
moschata as “it proved to have pilose leaves,” justifying his
earlier decision in favor of the Martius collection because
“…as the description of Nees refers especially to this
species and not to that of Sellow...” However, contrary to
Kostermans’s statement, Nees’s paper deals mainly with
the Sellow collections at Berlin (Revisio Laurinarum a b.
Sellowio in Brasilia collectarum et iam in Herbario Regio
Berolinensi asservatarum), from which the protologue
allowed a clear decision (see below) on which material the
species was primarily described [see also Rohwer, 1993b in
his lectotypiication of Nectandra leucantha Nees & Mart.,
Linnaea 8: 48. 1833]. In Moraes (2005), after studying
both syntypes, he has concluded that the Martius collection
was in conlict with the protologue, while the Sellow
one was not and, therefore he selected a specimen of the
Sellow collection as lectotype. According to the original
diagnosis, which is not much elucidative, C. moschata has
“foliis ovali-oblongis brevi-cuspidatis utrinque reticulatis
glabris concoloribus, pedunculis axillaribus petiolo paulo
longioribus pauciloris, fructu obovato umbonato costato.”
The Sellow collection has leaves ovate-oblong, shortcuspidate, on both surfaces reticulate, glabrous, concolor,
whereas the Martius collection has leaves ovate-oblong,
acute to pointleted, on both surfaces reticulate, but pubescent
on the lower surface, discolour (brownish and shining
above, somewhat olive-green and dull below). Still, the
specimen of Sellow has short peduncles, slightly longer than
petioles, whereas the specimen of Martius has peduncles
considerably longer than petioles, and fruits umbonicobovate, ribbed. Other authors might reject Moraes’s
supersession of the existing lectotype, with the allegation
that “the specimen chosen by Kostermans as lectotype is
not in ‘serious’ conlict with regards to the protologue,”
since the main evidence relies on the leaf pubescence. In
fact, as pointed out by Moraes, according to the protologue
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DE MORAES AND FALCADE, LAURACEAE COLLECTED By MARTIUS
C. moschata is a glabrous species, which was clearly
based on the Sellow collection. This indument character,
or its absence, plays a major part in the delimitation of
Cryptocarya moschata in the sense of Sellow’s syntype
and C. mandioccana, the species represented by Martius’s
collection. If the lectotypiication of C. moschata in the
sense of the Martius collection would not be rejected for the
reasons explained, it will be necessary to coin a new name
for the distinctive entity represented by Sellow’s collection.
Unfortunately, in Moraes’s lectotypiication he selected the
specimen preserved at LE as the lectotype, reasoning that it
was the only material, from the available isosyntypes he had
accessed, that still had a lower. Only later he found out that
CGE houses a duplicate of Sellow’s syntype still bearing
three lowers, which would be the best choice.
In Martius (1843a: 110), the plate 105 was indicated
for “Cryptocarya moschata Mart.” (see entry 2; present
work). However, in Meissner (1866: 164), he has only
quoted “Tabula nostra LVI,” thus evidencing that another
plate was prepared for this species. Although Kostermans
(1937, 1953, 1964) had also cited plate 105 under C.
moschata Nees & Mart., this manuscript has not been found
within the reliquiae of Martius either at the the Bayerische
Staatsbibliothek (Abteilung fuer Handschriften und Seltene
Drucke) or at the library of the Botanische Staatssammlung
München. Wuschek (1989), in his treatment of Martius’s
Systema materiae medicae, did not mention the unpublished
manuscript, either.
The illustration of Cryptocarya moschata in Flora
Brasiliensis (Meissner, 1866; plate 56) was delineated
from different lowering and fruiting material, based on the
specimens analyzed and cited by Meissner. In this sense,
only the lowering habit and the lower pieces can be ascribed
to C. moschata Nees & Mart., since they were drawn from
either Sellow’s or Widgren’s collections. On the other hand,
the fruiting elements of the plate could be either ascribed to
C. mandioccana (from Martius s.n.), C. riedeliana P.L.R.
Moraes (from Riedel 485), or to C. guianensis Meisn. (from
Martin s.n.). However, judging from the outlined shape of
leaves and fruits, it would be proved that they were based on
Martius’s collection.
Another manuscript of Martius (1843b), which is kept at
the University of California at Los Angeles Library (Louise
M. Darling Biomedical Library), is a bound volume that
has a simple bookplate stating: “E Bibliotheca C. Ph. F.
Martii.” Additionally it is stamped: “Urban Library 1933.”
The holographic titlepage reads: “Plantae medicinales
Florae Brasiliensis secundum Ordines naturales dispositae,”
“1843. 10. Junii.” The second leaf of the work has a
handwritten note: “Conferenda haec lora medica Brasil.
cum Schöpii lora med. Amer. bor., cum Lunan Jamaic.,”
likely refering to a comparison among the Flora Medica of
Brazil (probably from Martius’s Systema materiae medicae
vegetabilis brasiliensis), Johann David Schöpf’s Materia
medica americana (1787), and John Lunan’s Hortus
Jamaicensis (1814). Leaf 3–5 present a holographic list of
plants arranged by family and genus and species. Leaf 6
begins as a numbered Leaf 1, after that each opening (left
page and right page facing) is numbered on the right leaf.
There are 80 openings and the left side of the opening is
173
blank or contains handwritten notes made by Martius relating
to some of the printed text, which has been cut from some
other book and pasted on the right hand side. Regarding
the species of Lauraceae, all the 11 species treated in Syst.
mat. med. bras. compose the printed matter, with a different
order of entries of the species from the former. Additionally,
there are handwritten notes about Silvia navalium Allemão
[≡ Mezilaurus navalium (Allemão) Taub. ex Mez], and
“Nectandra rodgiaei Schomb.” [≡ Nectandra rodioei R.H.
Schomb. ≡ Chlorocardium rodiei (R.H. Schomb.) Rohwer,
H.G. Richt. & van der Werff]. The clipped printed text of
Cryptocarya moschata Mart. is pasted on the numbered leaf
16.
11. Dicypellium caryophyllaceum (Mart.) Nees & Mart. in
Nees, Pl. Laur. Expos.: 14. 1833; ibid. in Syst. Laur.: 344.
1836; Martius, Syst. Mat. Med. Bras.: 111. 1843; ibid. in
Fl. Bras. 5(2): 316. 1868; Meissner in DC., Prodr. 15(1):
170. 1864; ibid. in Martius, Fl. Bras. 5(2): 281, t. 102. 1866;
Mez, Jahrb. Königl. Bot. Gart. Berlin 5: 473. 1889; Rohwer,
Bot. Jahrb. Syst. 110: 169. 1988; Moraes, Harvard Pap. Bot.
17(1): 195. 2012 ≡ Ocotea caryophyllacea (Mart.) Kostel.,
Allg. Med.-Pharm. Fl. 2: 482. 1833.
Basionym: Persea caryophyllacea Mart. in Buchner,
Repert. Pharm. 31: 355. 1829; ibid. in Spix and
Martius, Reise Bras. 3: 915. 1831. TyPE: BRAZIL.
Amazonas, Maués, “in sylvis ad Maué hace culta in
hortis Parae.,” April 1820, C. F. P. von Martius s.n.
(Holotype: M 0147268; Isotypes, B 10 0185010, B
10 0185011, G, GZU 000254389). Other original
material: BRAZIL. C. F. P. von Martius Obs. 2617:
“Persea caryophyllat. Mart./Dicipellium caryophyllat.
Nees/Laurus sanguinea Sw. ? … Colitur in horto regio
Pará, ex interioribus silvis advecta” (M 0147264, M
0147265, M 0147266 [as Martius s.n.]; BRAZIL.
Amazonas, “ad Coari et Ega,” November 1819, C. F.
P. von Martius s.n. (M 0147267 [ex Herb. Zuccarinii]).
Vernacular names: “Cravo do Maranhao” (Martius,
1829), “Páo Cravo Bras.,” “Imyra-Quiynha v. Kiynja i. e.
lignum Capsici Tupin.” (Martius, 1843).
12. Licaria brasiliensis (Nees) Kosterm., Recueil Trav.
Bot. Néerl. 34: 601. 1937 ≡ Aydendron brasiliense (Nees)
Meisn. in DC., Prodr. 15(1): 87. 1864; ibid. in Martius, Fl.
Bras. 5(2): 180, t. 64. 1866 ≡ Misanteca brasiliensis (Nees)
Lundell, Wrightia 4(3): 99. 1969.
Basionym: Acrodiclidium brasiliense Nees, Syst. Laur.:
267. 1836; Mez, Jahrb. Königl. Bot. Gart. Berlin 5:
92. 1889; Kostermans, Recueil Trav. Bot. Néerl. 33:
746. 1936; Kurz, Mitt. Inst. Allg. Bot. Hamburg 28/29:
157. 2000. TyPE: BRAZIL. Pará, “in sylvis secus
luvium Amazonum,” September–October 1819,
l., C. F. P. von Martius s.n. (Holotype: M 0147427;
Isotypes, L 0308064 [“Laurus riparia M.,” herbarium
name, “Amaz. l.”], M 147428, M 0147429, MEL
2324397 [“Laurus riparia,” herbarium name, “Para”],
MEL 2324398 [“Laurus riparia Mart.”, herbarium
name, “Para”], S [R-7276]).
Further synonyms in Kurz (2000).
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HARVARD PAPERS IN BOTANy
13. Licaria crassifolia (Poir.) P.L.R. Moraes, Komarovia
6(1): 55. 2008. Specimen: BRAZIL. Amazonas, “in sylvis
ad Coari,” November 1819, sterile, C. F. P. von Martius s.n.
(GZU 000256425 [annotated by Nees; leaf fragment], M
0147426) [Syntype of Nectandra nitidula Nees & Mart. var.
maior Nees, Syst. Laur.: 313. 1836].
Basionym: Laurus crassifolia Poir. in Lam. Encycl., Suppl.
3: 323. 1813. Lectotype (designated by Moraes, 2008:
55): FRENCH GUIANA. “Caienne”, 1791, fr., L.
C. M. Richard s.n. (P 00128465 [ex Herb. Poiret];
Isolectotypes, P 00128463 [ex Herb. Richard], P
00128464 [ex Herb. E. Drake], P-JU 00662838 [Cat.
No. 1226]).
Synonyms: Aydendron cannella Meisn. in DC., Prodr.
15(1): 90. 1864; ibid. in Martius, Fl. Bras. 5(2): 180.
1866 ≡ Acrodiclidium cannella (Meisn.) Mez, Jahrb.
Königl. Bot. Gart. Berlin 5: 90. 1889; Kostermans,
Recueil Trav. Bot. Néerl. 33: 728. 1936 ≡ Licaria
cannella (Meisn.) Kosterm., Recueil Trav. Bot. Néerl.
34: 583. 1937; Kurz, Mitt. Inst. Allg. Bot. Hamburg
28/29: 121. 2000 ≡ Misanteca cannella (Meisn.)
Lundell, Wrightia 4(3): 99. 1969. TyPE: FRENCH
GUIANA. Acarouany, August 1858, l., P. A. Sagot
1190 (Holotype: G-DC; Isotypes, B† [F neg. 3837],
BM 000993917, K 000602022, Ny 00354870, P
00128461, P 00128462, U 0002746, W n.v.).
Aydendron cayennense Meisn. in DC., Prodr. 15(1):
95. 1864; ibid. in Martius, Fl. Bras. 5(2): 186. 1866
≡ Acrodiclidium cayennense (Meisn.) Mez, Jahrb.
Königl. Bot. Gart. Berlin 5: 91. 1889; Kostermans,
Recueil Trav. Bot. Néerl. 33: 730. 1936 ≡ Licaria
cayennensis (Meisn.) Kosterm., Recueil Trav. Bot.
Néerl. 34: 583. 1937 ≡ Misanteca cayennensis
(Meisn.) Lundell, Wrightia 4(3): 99. 1969. TyPE:
FRENCH GUIANA. Caiena, s.d., fr., J.-B. Patris 41
(Holotype: G-DC [F neg. 7288]; Isotypes, B†, G, L
0036456, Ny 00354869, P).
14. Licaria polyphylla (Nees) Kosterm., Recueil Trav. Bot.
Néerl. 34: 584. 1937; Kurz, Mitt. Inst. Allg. Bot. Hamburg
28/29: 167. 2000 ≡ Misanteca polyphylla (Nees) Lundell,
Wrightia 4(3): 101. 1969.
Basionym: Nectandra polyphylla Nees, Syst. Laur.: 332.
1836; Meissner in DC., Prodr. 15(1): 160. 1864;
ibid. in Martius, Fl. Bras. 5(2): 270. 1866; Mez,
Jahrb. Königl. Bot. Gart. Berlin 5: 97. 1889. TyPE:
BRAZIL. Amazonas, Coari, “in sylvis ad Coari,”
November 1819, sterile, C. F. P. von Martius s.n.
(Holotype: M 0147422; Isotype, GZU 000254280).
Synonyms: Acrodiclidium guianense Nees var. caudatum
Meisn. in DC., Prodr. 15(1): 85. 1864; ibid. in Martius,
Fl. Bras. 5(2): 173. 1866; Mez, Jahrb. Königl. Bot.
Gart. Berlin 5: 98. 1889; Kostermans, Recueil Trav.
Bot. Néerl. 33: 734. 1936 ≡ Acrodiclidium meissneri
Mez, Jahrb. Königl. Bot. Gart. Berlin 5: 98. 1889.
Lectotype (designated by Kostermans, 1936: 738):
VENEZUELA. Amazonas, “prope San Carlos, ad Rio
Negro,” August 1853, R. Spruce 3058 (K 000602019;
VOL. 20, NO. 2
Isolectotypes, B† [F neg. 3841], BM 000947197,
BR 0000008812810, C 10013587, E 00259391,
F [0061307F], G 00368869, G 00368870, G-DC,
GH 00041577, GOET 004500, K 000602020, LE
00000141, Ny 00354749, Ny 00354750, P 00757118,
P 00757119, P 00757120, S [S-R-7278], U, W n.v.).
Further synonyms in Kurz (2000).
15. Licaria puchury-major (Mart.) Kosterm., Recueil
Trav. Bot. Néerl. 34: 583. 1937; Kurz, Mitt. Inst. Allg. Bot.
Hamburg 28/29: 126. 2000; Moraes, Harvard Pap. Bot.
17(1): 198. 2012 ≡ Nectandra puchury-major (Mart.) Nees
& Mart. in Nees, Syst. Laur.: 328. 1836; Martius, Syst. Mat.
Med. Bras.: 109–110. 1843; ibid. in Fl. Bras. 5(2): 317.
1868; Meissner in DC., Prodr. 15(1): 156. 1864; ibid. in
Martius, Fl. Bras. 5(2): 265, t. 95. 1866 ≡ Acrodiclidium
puchury-major (Mart.) Mez, Jahrb. Königl. Bot. Gart.
Berlin 5: 88. 1889; Kostermans, Recueil Trav. Bot. Néerl.
33: 727. 1936 ≡ Misanteca puchury-major (Mart.) Lundell,
Wrightia 4: 101. 1969.
Basionym: Ocotea puchury-major Mart. in Buchner,
Repert. Pharm. 35: 171. 1830. TyPE: BRAZIL.
Amazonas, “in sylvis ad Tabatinga,” December 1819,
fr., C. F. P. von Martius s.n. (Holotype: M 0147421
[F neg. 19259]; Isotypes, B†, BR 0000008812841,
G 00368872, GZU 000254328, L 0036458 [“Laurus
Pichurim vera,” herbarium name, “Rio Negro”; ex
Herbarium Reinwardtianum]). Probable isotypes: M
0147419, M 0147420.
For a complete list of synonyms, see Kurz (2000).
Vernacular names: “Fabas Pechurim v. Puchury”
(Martius, 1868).
16. Nectandra acuminata (Nees & Mart.) J.F. Macbr., Publ.
Field Mus. Nat. Hist., Bot. Ser., 11(1): 19. 1931; Rohwer,
Mitt. Inst. Allg. Bot. Hamburg 20: 49. 1986; ibid. in Fl.
Neotrop. Monogr. 60: 270. 1993.
Basionym: Persea acuminata Nees & Mart. in Nees, Syst.
Laur.: 170. 1836; Mez, Jahrb. Königl. Bot. Gart.
Berlin 5: 409. 1889 [as synonym of Nectandra
acutifolia (Ruiz & Pav.) Mez]. TyPE: BRAZIL.
Amazonas, Rio Japurá, “in sylvis ad l. Japurá,”
December 1819, monstr. fr., C. F. P. von Martius s.n.
(Holotype: M 0147319) [also syntype of Nectandra
berchemiifolia Meisn. in DC., Prodr. 15(1): 154. 1864,
as “berchemiaefolia”; ibid. in Martius, Fl. Bras. 5(2):
262. 1866; Rohwer, Mitt. Inst. Allg. Bot. Hamburg
20: 50. 1986; ibid. in Fl. Neotrop. Monogr. 60: 267.
1993].
17. Nectandra amazonum Nees, Syst. Laur.: 282. 1836;
Meissner in DC., Prodr. 15(1): 150. 1864; ibid. in Martius,
Fl. Bras. 5(2): 256, t. 92. 1866; Mez, Jahrb. Königl. Bot.
Gart. Berlin 5: 420. 1889; Rohwer, Mitt. Inst. Allg. Bot.
Hamburg 20: 49. 1986; ibid. in Fl. Neotrop. Monogr. 60: 236.
1993; Moraes, Pl. Ecol. Evol. 146(3): 369. 2013. Lectotype
(designated by Rohwer, 1986: 49, as “Holotyp M”):
BRAZIL. Pará, “in luminis Amazonum omni ripa a Pará,
2015
DE MORAES AND FALCADE, LAURACEAE COLLECTED By MARTIUS
ad Tabatinga,” September–October 1819, fr., C. F. P. von
Martius s.n. (M 0147314; Isolectotype, GZU 000254251).
Syntypes: BRAZIL. Pará, l., C. F. P. von Martius s.n. (B
10 0185108, G, GZU 000254252, L 0037115 [“Laurus
amazonum M / secus totum l. Amazon.,” herbarium name],
M 0147315, M 0147316, M 0147317, M 0147318, MEL
2324402 [“Laurus amazonum,” herbarium name, “Para”],
U 0002889 [fragment]).
Synonymy in Rohwer (1993).
18. Nectandra canescens Nees & Mart. in Nees, Linnaea 8:
47. 1836; ibid. in Syst. Laur.: 280. 1836; ibid. in Linnaea 21:
501. 1848; Meissner in DC., Prodr. 15(1): 147. 1864; ibid.
in Martius, Fl. Bras. 5(2): 251. 1866; Mez, Jahrb. Königl.
Bot. Gart. Berlin 5: 408. 1889; Rohwer, Mitt. Inst. Allg.
Bot. Hamburg 20: 51. 1986; ibid. in Fl. Neotrop. Monogr.
60: 274. 1993; Moraes, Komarovia 6(1): 23. 2008; ibid.
in Harvard Pap. Bot. 17(1): 199. 2012; ibid. in Pl. Ecol.
Evol. 146(3): 369. 2013. Lectotype (designated by Rohwer,
1993: 274): BRAZIL. Pará, “in sylvis secus l. Amazonum,
Provinciae Paraënsis et RN,” 20 October 1819, l., immat.
fr., C. F. P. von Martius Martius Obs. 2750: “Pomatium
canescens … arbor 15–20 ped. … In silvis ad marginem
l. Amazonum” (M 0112453 [annotated by Martius as
“Persea canescens,” and by Nees as “Pomatium canescens,”
herbarium names]; Isolectotypes, GZU 000254254
[annotated by Nees as “Pomatium canescens M. / in sylvis
secus lumen Amazonum Provinciae Paraensis. Sept. et
Oct.; in sylv. Ad luv. Madeiram Prov. Flumini Nigri Aprili /
a Martius”], M 0112458, M 0112459). Specimen: BRAZIL.
Amazonas, “in sylvis ad luv. Madeira,” April 1820, sterile,
C. F. P. von Martius s.n. (B 10 0185095 [F neg. 3750]).
Synonymy in Rohwer (1993) and Moraes (2012a).
The other syntype of Nectandra canescens Nees & Mart.
is the gathering by “Sellow B 433 c 410”, which is the type
of Nectandra rigida (Kunth) Nees var. canescens Mez,
Jahrb. Königl. Bot. Gart. Berlin 5: 407. 1889. According to
Rohwer (1993: 288), although N. rigida var. canescens is
based on the Sellow syntype of N. canescens Nees & Mart.,
it has to be treated as a new variety and not as a combination
based on N. canescens, because Mez accepted Nectandra
canescens in the sense of its Martius syntype in the same
publication.
19. Nectandra cuspidata Nees & Mart. in Nees, Syst.
Laur.: 330. 1836; Meissner in DC., Prodr. 15(1): 159. 1864;
ibid. in Martius, Fl. Bras. 5(2): 268, t. 97. 1866; Mez,
Jahrb. Königl. Bot. Gart. Berlin 5: 449. 1889; Rohwer, Fl.
Neotrop. Monogr. 60: 100. 1993 ≡ Nectandra membranacea
(Sw.) Griseb. ssp. cuspidata (Nees & Mart.) Rohwer, Mitt.
Inst. Allg. Bot. Hamburg 20: 72. 1986; Moraes, Pl. Ecol.
Evol. 146(3): 369. 2013. Lectotype (designated by Rohwer,
1993: 100): BRAZIL. Amazonas, Tefé, “in ripa luminis
Amazonum, praesertim ad villam Ega,” January 1820, l.,
fr., C. F. P. von Martius s.n. (M 0147303; Isolectotypes,
B 10 0086138, GZU 000254262, L 0037121, L 0037122
[“Laurus cuspidata M. / R. Amazon.,” herbarium name], L
1802480 [“Laurus egensis Mart.,” herbarium name, “Prov.
175
Rio Negro”], M 0147304). Syntypes: BRAZIL. Amazonas,
Rio Japurá, “in sylvis Japurensibus,” January 1820, l., fr.,
C. F. P. von Martius s.n. (GZU 000254262, M 0147302, M
0147306); BRAZIL. Amazonas/Pará, “in sylvis secus luv.
Maué et Amazonum,” September 1819, immat. fr., C. F. P.
von Martius s.n. (GZU 000254261, M 0147305).
Synonymy in Rohwer (1993).
“Martius Obs. 2721 Ocotea”, which has not been
annotated in any specimen in Munich, most likely refers to
this species.
20. Nectandra hihua (Ruiz & Pav.) Rohwer, Fl. Neotrop.
Monogr. 60: 196. 1993 ≡ Nectandra leucantha Nees var.
peruviana Meisn. in DC., Prodr. 15(1): 151. 1864.
Basionym: Laurus hihua Ruiz & Pav., Fl. Peruv. 4(2): t.
366. 1804[–1830] (Ruiz and Pavón, 1956: 33, t. 366).
TyPE: ECUADOR. Guayas, “in Huayaquilensibus
nemoribus,” 1799, J. Tafalla s.n. (Holotype: MA
811751; Isotypes, B 10 0185060, F [0040309F;
fragment], FI-W 005165, G 00369022, MA 811752).
Synonyms: Nectandra lucida Nees, Syst. Laur.: 334. 1836,
non ibid. p. 295, non 1833, hom. post. ≡ Nectandra
schomburgkii Meisn. in DC., Prodr. 15(1): 151.
1864; ibid. in Martius, Fl. Bras. 5(2): 258. 1866.
Lectotype (designated by Mez, 1889: 429): BRAZIL.
Amazonas, “ad lumen Japurá,” December 1819,
fr., C. F. P. von Martius s.n. (M 0121041 [“Ocotea
lucida Mart.,” herbarium name]; Isolectotype, GZU
000249367). Syntype: BRAZIL. Amazonas, “in
sylvis Japurensibus,” February 1820, l., C. F. P. von
Martius s.n. (M 0147294).
Complete synonymy in Rohwer (1993).
21. Nectandra japurensis Nees & Mart. in Nees, Syst.
Laur.: 335. 1836; Meissner in DC., Prodr. 15(1): 163. 1864;
ibid. in Martius, Fl. Bras. 5(2): 274, t. 99. 1866; Mez, Jahrb.
Königl. Bot. Gart. Berlin 5: 440. 1889; Rohwer, Mitt. Inst.
Allg. Bot. Hamburg 20: 65. 1986; ibid. in, Fl. Neotrop.
Monogr. 60: 125. 1993. Lectotype (designated by Rohwer,
1993: 125): BRAZIL. Amazonas, “in sylvis Japurensibus
ad Maripi, S. Joh. Princ. cet. Provinciae Fluminis nigri
Brasiliae,” December–January 1819/1820, l., C. F. P. von
Martius s.n. (M 0147292; Isolectotypes, B 10 0185058
[Mus. Bot. Berol. Film Nr. 1232/13; Ny neg. 8343], B 10
0185059, GZU 000254272, HBG [fragment], L 0037128
[“Laurus japurensis M. / R. Negro,” herbarium name; ex
Herbarium Reinwardtianum], M 0147293).
Synonymy in Rohwer (1993).
22. Nectandra leucantha Nees & Mart. in Nees, Linnaea
8: 48. 1833; ibid. in Syst. Laur.: 310. 1836; Meissner in
DC., Prodr. 15(1): 150. 1864; ibid. in Martius, Fl. Bras.
5(2): 257. 1866; Mez, Jahrb. Königl. Bot. Gart. Berlin 5:
431. 1889; Rohwer, Mitt. Inst. Allg. Bot. Hamburg 20: 52.
1986; ibid. in Fl. Neotrop. Monogr. 60: 201. 1993; Moraes,
Komarovia 6(1): 19. 2008 ≡ ‘Persea leucantha Mart.’,
nom. nud. Lectotype (designated by Rohwer, 1993: 201):
BRAZIL. Loco haud indicato, s.d., l., F. Sellow s.n. (B 10
176
HARVARD PAPERS IN BOTANy
0185048 [B neg. 6873]). Syntypes: BRAZIL. Minas Gerais,
“In camposus Tabuleiro ad Salgado,” August 1818, l., C. F.
P. von Martius Obs. 1704: “arbor 15–20 ped. … In campis
locis Tabuleiro consitis” (GZU 000254273 [annotated by
Nees; Anonymous s.n.], M 0111058) [= Nectandra hihua
(Ruiz & Pav.) Rohwer, Fl. Neotrop. Monogr. 60: 196.
1993]). Specimen: BRAZIL. Bahia, “m. ad. V. do Rio de
Contas interionis,” October–November 1818, l., C. F. P.
von Martius Obs. 1704 (GZU 000254273 [Anonymous s.n.],
M 0111057).
Synonymy in Rohwer (1993).
23. Nectandra membranacea (Sw.) Griseb., Fl. Brit. W. I.
282. 1860; Meissner in DC., Prodr. 15(1): 161. 1864; Mez,
Jahrb. Königl. Bot. Gart. Berlin 5: 450. 1889; Rohwer, Mitt.
Inst. Allg. Bot. Hamburg 20: 70. 1986; ibid. in Fl. Neotrop.
Monogr. 60: 94. 1993 ≡ Persea membranacea (Sw.) Spreng.,
Syst. Veg. (ed. 16) 2: 270. 1825 ≡ Phoebe membranacea
(Sw.) Nees, Syst. Laur.: 118. 1836 ≡ Ocotea membranacea
(Sw.) R.A. Howard, J. Arnold Arbor. 62: 59. 1981.
Basionym: Laurus membranacea Sw., Prodr. 65. 1788.
TyPE: JAMAICA. Loco haud indicato, s.d., l., O.
P. Swartz s.n. (Holotype: S [R-3166; Ny neg. 8559];
Isotypes, BM 000758874, S [09-16300]). Specimen:
“Ex India occidentalis”, s.d., l., O. P. Swartz s.n.
(SBT 11582).
Synonym: Nectandra cuspidata Nees & Mart. var.
macrocarpa Nees, Syst. Laur.: 331. 1836. TyPE:
BRAZIL. Rio de Janeiro, “in sylvis caeduis,” October
1817, fr., C. F. P. von Martius s.n. (Holotype: M
0147307).
Complete synonymy in Rohwer (1993).
24. Nectandra nitidula Nees & Mart. in Nees, Linnaea 8:
48. 1833; ibid. in Syst. Laur.: 313. 1836; Meissner in DC.,
Prodr. 15(1): 160. 1864; ibid. in Martius, Fl. Bras. 5(2):
271, t. 98. 1866; Mez, Jahrb. Königl. Bot. Gart. Berlin 5:
436. 1889; Rohwer, Mitt. Inst. Allg. Bot. Hamburg 20:
57. 1986; ibid. in Fl. Neotrop. Monogr. 60: 180. 1993;
Moraes, Komarovia 6(1): 20. 2008; ibid. in Harvard Pap.
Bot. 17(1): 200. 2012; ibid. in Pl. Ecol. Evol. 146(3): 369.
2013 ≡ Nectandra nitidula Nees & Mart. var. minor Nees,
Syst. Laur.: 313. 1836, nom. illegit. superl., = var. nitidula.
Lectotype (designated by Rohwer, 1993: 180): BRAZIL.
São Paulo, “in campis prope Taubaté,” December 1817, fr.,
C. F. P. von Martius Obs. 546: “arbor 20 ped. cal. viridis”
(M 0111064). Other syntypes: BRAZIL. São Paulo, 1829,
l., F. Sellow 28 (B 10 0185228 [B neg. 1581/3, Ny neg.
8319]); BRAZIL. Loco haud indicato, s.d., l., F. Sellow
s.n. (B 10 0185227 [B neg. 5197]). Probable syntypes:
BRAZIL. Loco haud indicato, s.d., l., F. Sellow s.n. (B 10
0185226 [ex Herb. Baschant]); idem, s.d., l., F. Sellow s.n.
(BR 0000008808479, E 00259309); idem, s.d., l., F. Sellow
s.n. (CGE); idem, s.d., l., Anonymous s.n. (GZU 000259200
[annotated by Nees]).
Synonyms: Nectandra nitidula Nees & Mart. var. maior
Nees, Syst. Laur.: 313. 1836; Rohwer, Mitt. Inst.
Allg. Bot. Hamburg 20: 57. 1986; ibid. in Fl. Neotrop.
VOL. 20, NO. 2
Monogr. 60: 181. 1993; Moraes, Komarovia 6(1): 20.
2008; ibid. in Harvard Pap. Bot. 17(1): 200. 2012; ibid.
in Pl. Ecol. Evol. 146(3): 369. 2013. LECTOTyPE
(designated here): BRAZIL. Minas Gerais, Serro, “in
montosis Serro frio Provinciae Minarum generalium,”
May 1818, l., C. F. P. von Martius s.n. (M 0111060
[annotated by Nees]; Isolectotype, GZU 000249360
[annotated by Nees with “Nectandra nitidula, nondum
lorens / Petalanthera paniculigera N. ab Es. / Minas
Geraes”]). Syntypes: BRAZIL. Minas Gerais,
Diamantina, “in campis, Tabuleiro, terrae Adamantin.
Prope Tejuco,” June 1818, fr., C. F. P. von Martius
s.n. (GZU 000256425 [annotated by Nees; fragment;
Anonymous s.n.], M 0111063 [annotated by Nees]);
BRAZIL. Bahia, “in sylvis Catingas locis editioribus
deserto Min. Nov.,” October 1818, l., C. F. P. von
Martius s.n. (GZU 000249359 [annotated by Nees],
M 0111061 [F neg. 6590; annotated by Martius as
“Persea paniculigera,” herbarium name]); BRAZIL.
Paraná, l., F. Sellow 4495 (B 10 0088601 [B neg.
1581/5, Ny neg. 8321; mixtum compositum, only the
lower fragment]; BRAZIL. Loco haud indicato, s.d.,
l., F. Sellow 1368* (B†, L 0037147); BRAZIL. Santa
Catarina, “Insula S. Catharinae,” s.d., l., J. Tweedie
s.n. (E 00259433, GZU 000249358 [ex Herb. Arnott.,
annotated by Nees as “Nectandra nitidula Var. a;”
Anonymous s.n.]) [= Nectandra megapotamica
(Spreng.) Mez, Bull. Herb. Boissier, Sér. 2, 2: 824.
1902]. Specimens: BRAZIL. Minas Gerais, August
1839, P. Claussen in Martii Herbar. Florae Brasil.
Nº 1239 [as 1293 in Mez, 1892] (AWH 28295, BM
000947260, G, G-DC, JE 00004387 [Anonymous
1239], K [not located], L 0037144, L 0037145, L
0037146, LE 00006796, LE, M 0111059, M 0111062,
MO 247468, Ny 00355481, OXF 00083028, P
00711051) = Claussen 106 [BR 0000008808240],
Claussen 107 [BR 0000008807946], Claussen s.n.
[BR 0000008808271; attributed to Riedel]).
Nectandra sarcocalyx Nees, Syst. Laur.: 338. 1836;
Meissner in DC., Prodr. 15(1): 167. 1864; ibid. in
Martius, Fl. Bras. 5(2): 278. 1866; Mez, Jahrb. Königl.
Bot. Gart. Berlin 5: 436. 1889. TyPE: BRAZIL.
Bahia, “in alpestribus campis ad Va. Do Rio das
Contas et Sincorâ,” October 1818, fr. monstr., C. F. P.
von Martius s.n. (Holotype: M 0111065 [annotated by
Martius as “[Laurus] Persea sarcocalyx,” herbarium
name]; Isotype, GZU 000254287 [annotated by Nees;
fragment]).
Further synonymy in Rohwer (1993), and Moraes
(2012a).
Rohwer (1993: 181) designated the collection
Martii Herbar. Florae Brasil. Nº 1239 as the lectotype
of Nectandra nitidula Nees & Mart. var. maior Nees.
However, this collection refers to a gathering undertaken
by Peter Claussen in Cachoeira do Campo, Minas Gerais,
in August 1839, which therefore is not an original material
for the lectotypiication of the variety named by Nees von
Esenbeck.
2015
DE MORAES AND FALCADE, LAURACEAE COLLECTED By MARTIUS
25. Nectandra oppositifolia Nees & Mart. in Nees, Linnaea
8: 47. 1833; ibid. in Syst. Laur.: 279. 1836; Meissner in DC.,
Prodr. 15(1): 146. 1864; ibid. in Martius, Fl. Bras. 5(2): 250.
1866; Mez, Jahrb. Königl. Bot. Gart. Berlin 5: 406. 1889;
Rohwer, Fl. Neotrop. Monogr. 60: 286. 1993; Moraes,
Komarovia 6(1): 22. 2008; ibid. in Harvard Pap. Bot.
17(1): 201. 2012; ibid. in Pl. Ecol. Evol. 146(3): 369. 2013
≡ Nectandra oppositifolia Nees & Mart. var. angustifolia
Nees & Mart. in Nees, Linnaea 8: 47. 1833, nom. illegit.
superl., = var. oppositifolia ≡ Nectandra mollis (Kunth)
Nees ssp. oppositifolia (Nees & Mart.) Rohwer, Mitt. Inst.
Allg. Bot. Hamburg 20: 61. 1986. Lectotype (designated
by Rohwer, 1993: 286): BRAZIL. São Paulo, l., F. Sellow
5722 (B 10 0185222 [Ny neg. 8327; annotated by Mez
as “Nectandra oppositifolia Nees fa. intermedia,” nom.
nud.]; Isolectotypes, B 10 0185221 [ex Herb. Kunth], BM
000895363, G-DC 00130422, K 000512726, VT 001549).
Other syntypes: BRAZIL. “Brasilia orientalis”, l., F. Sellow
LVI (BR 0000008808226); BRAZIL. Santa Catarina, l., F.
Sellow 4428 (B 10 0185220). Probable syntypes: BRAZIL.
Loco haud indicato, l., F. Sellow s.n. (GZU 000249356,
HAL 0010385 [both with labels by Schlechtendal annotated
as “Pomatium oppositifolium”]); idem, l., F. Sellow s.n.
(GZU 000249357 [annotated by Nees as “Pomatium
oppositifolium”]). Specimens: BRAZIL. Loco haud
indicato, sterile., F. Sellow s.n. (B 10 0185172); idem, l., F.
Sellow s.n. (B 10 0185175); BRAZIL. São Paulo, Iperó, “in
sylvis primaevis ad ypanema,” January 1818, sterile, C. F.
P. von Martius s.n. (M 0121029); BRAZIL. Minas Gerais,
“in sylv. abor. ad l. Doce,” 1818, l., C. F. P. von Martius
s.n. (M 0121027).
Synonym: Nectandra schottii Meisn. in DC., Prodr. 15(1):
147. 1864; ibid. in Martius, Fl. Bras. 5(2): 252. 1866;
Mez, Jahrb. Königl. Bot. Gart. Berlin 5: 407. 1889 ≡
‘Nectandra acutissima Nees ex Meisn.’ in DC., Prodr.
15(1): 147. 1864, nom. invalid. Lectotype (designated
by Rohwer, 1993: 286): BRAZIL. Rio de Janeiro,
“prope Cantagallo,” s.d, H. W. Schott 5607 (M).
Syntype: BRAZIL. Rio de Janeiro, “in sylvis caeduis
ad Sebastianopolin,” December 1817, fr., C. F. P. von
Martius s.n. (GZU 000249322, M 0121030).
Further synonyms and detailed information in Rohwer
(1993) and Moraes (2008).
26. Nectandra psammophila Nees & Mart. in Nees, Syst.
Laur.: 303. 1836; Meissner in DC., Prodr. 15(1): 161. 1864;
ibid. in Martius, Fl. Bras. 5(2): 272. 1866; Mez, Jahrb.
Königl. Bot. Gart. Berlin 5: 434. 1889; Rohwer, Mitt. Inst.
Allg. Bot. Hamburg 20: 68. 1986; ibid. in Fl. Neotrop.
Monogr. 60: 160. 1993; Moraes, Harvard Pap. Bot. 17(1):
201. 2012; ibid. in Scripta Bot. Belg. 49: 120. 2013.
Lectotype (designated by Rohwer, 1993: 160): BRAZIL.
Espírito Santo, Vila Velha, “Barra de Jucu,” March 1816
[sic; December 1815], l., M. A. P., Prinz zu Wied s.n. (BR
0000008808066; Isolectotypes, B 10 0185169 [F neg. 3763],
BR 0000008808363, GOET 000753, GZU 000254281,
MEL 2324403 [“Laurus amoena Mart.,” herbarium
name], MEL 2324386). Probable isolectotypes: BRAZIL.
177
[Bahia], with label “Ad l. Ilheos,” s.d., l., Anonymous
s.n. [“Communic. Martius 1827”] (K 000642237; “Laurus
amoena Mart.,” herbarium name); idem, [Bahia], with
label “In silvis prope Rio das Contas,” s.d., l., Anonymous
s.n. [“Communic. Martius 1827”] (K 000642238; “Laurus
saporosa Mart.,” herbarium name). Syntype: BRAZIL. Rio
de Janeiro/São Paulo, “in sylvis subalpinis alt. 1500 pedum
supra Serra do Mar”, December 1817, immat. fr., C. F. P.
von Martius Obs. 411: “Arbor 30 ped. Flores in racemis
6–8. In alpinis alt. 1500’” (GZU 000260031, M 0121034).
Specimen: BRAZIL. São Paulo, Iperó, “in sylv. primaevis
ad ypanema,” January 1817, sterile, C. F. P. von Martius
s.n. (M 0121033 [annotated by Rohwer as “probabiliter
Nectandra aff. psammophila Nees vel N. megapotamica
(Spr.) Mez”]).
Further synonymy in Rohwer (1993).
Specimens at K 000642237 and K 000642238 have
Herbarium Martii labels, typically printed with “Communic
……182…,” and handwritten with “Martius” and “7” (i.e.
“Communicavit Martius 1827”). Moraes et al. (2013),
and in subsequent papers, have reported part of the sets
of Brazilian plants that Martius distributed in 1827 to
several European herbaria and correspondents, particularly
concerning the sets received by Luigi Colla and published
in Herbarium Pedemontanum. In those papers, it has been
demonstrated that most of the specimens communicated by
Martius in 1827 came from collections of Prince Maximilian
zu Wied, from evidence of the localities (toponyms)
indicated on labels, being the same as those visited by Wied
in Brazil, and from matching of the specimens distributed
by Martius with those that are clearly attributed to Wied.
In the present case, the specimen at K 000642237 closely
matches Wied collections at BR and MEL, while the one at
K 000642238 closely matches the collections at B, GOET,
and GZU. Collections from Ilhéus and Rio de Contas can
be attributed either to Martius or to Wied, since both visited
these localities.
27. Nectandra puberula (Schott) Nees, Syst. Laur.: 332.
1836; Rohwer, Mitt. Inst. Allg. Bot. Hamburg 20: 58.
1986; ibid. in Fl. Neotrop. Monogr. 60: 220. 1993; Moraes,
Harvard Pap. Bot. 17(1): 202. 2012.
Basionym: Persea puberula Schott in Sprengel, Syst. Veg.
(ed. 16) 4(2): 405. 1827. TyPE: BRAZIL. Rio de
Janeiro, loco haud indicato, s.d., H. W. Schott s.n.
(Holotype: B 10 0185167 [B neg. 5200, ex Herb.
Sprengel nr. 40]; Isotypes, GZU 000254414, and
probably P 00711066, P 00711067).
Synonym: Nectandra amara Meisn. var. chartacea Meisn. in
DC., Prodr. 15(1): 158. 1864; ibid. in Martius, Fl. Bras.
5(2): 267. 1866. Lectotype (designated by Rohwer,
1993: 221): BRAZIL. Rio de Janeiro, “São Cristóvão,”
s.d., H. W. Schott 5609 (G 00369012; Isolectotypes,
F [0061480F], F [0061481F; fragment], LE, US
00048566, US 00997590). Syntypes: BRAZIL. Rio
de Janeiro, “in sylvaticis collibusque umbrosis circa
Rio de Janeiro,” May–July 1832, L. Riedel 480 (B 10
0185163 & GOET [2 sheets; Riedel s.n.; annotated
178
HARVARD PAPERS IN BOTANy
as “Nectandra amara β. subevenia Meissn.”], C, G
00368979, K 000576215, K 000576216 [as Riedel
s.n.], L 0037068, LE, MEL 2324391, MICH 1104567,
MICH 1104568, Ny 00355407, Ny 00355408, Ny
00355409, P 00711053, P 00711054, US 00288970,
W n.v.); BRAZIL. Rio de Janeiro, “Campos ad St.
João et Cabo Frio,” October 1833, l., B. Luschnath
s.n. in Martii Herbar. Florae Brasil. No 1307 (BR
0000008810496); BRAZIL. Mato Grosso, Cuiabá,
1836, l., A. L. P. da Silva Manso s.n. in Martii Herbar.
Florae Brasil. No 1307 (BR 0000008810878, G-DC,
LE, M 0121045, MICH 11104567, Ny 00355406, P
00711056); BRAZIL. Loco haud indicato, s.d., l., P.
Claussen 47 (B 10 0185166, KIEL); BRAZIL. Loco
haud indicato, s.d., l., P. Claussen 78 (B 10 0185165,
KIEL); BRAZIL. Minas Gerais, loco haud indicato,
s.d., l., P. Claussen 2078 (F [0061484F; fragment],
G 00368980, G 00368981, G 00369008, MO 247433,
P 00711064, P 00711065); BRAZIL. Rio de Janeiro,
“Près le Jardin bot. de Rio de Janeiro,” March 1839, l.,
J. B. A. Guillemin 667 (F [0061482F], F [0061483F],
G 00368978, G-DC, P 00711057, P 00711058, P
00711059).
For a complete list of synonyms, see Rohwer (1993).
Specimen at BR 0000008810526 bears a label annotated
anonymously as “Nectandra canescens Nees / Brasilia
Martius,” being identiied by Kostermans in 1935 as
“Nectandra puberula Nees”.
28. Nectandra purpurea (Ruiz & Pav.) Mez, Jahrb. Königl.
Bot. Gart. Berlin 5: 443. 1889; Rohwer, Mitt. Inst. Allg.
Bot. Hamburg 20: 77. 1986; ibid. in Fl. Neotrop. Monogr.
60: 46. 1993; Moraes, Harvard Pap. Bot. 17(1): 202. 2012.
Basionym: Laurus purpurea Ruiz & Pav., Fl. Peruv. 4(2):
t. 351. 1804[–1830] (Ruiz and Pavón, 1956: 17,
t. 351). Lectotype (designated by Bernardi, 1967:
60, as “specimine holotypico herbarii berolinensis
addicto”): PERU. Huánuco, “in Andium nemoribus
ad Cuchero et Chinchao,” 1778, H. Ruíz L. & J.
A. Pavón s.n. (B 10 0185160 [B neg. 1232/25];
Isolectotypes, BM 000947299, F [0040316F; #13/25
ex MA 811806], G 00369268 [F neg. 27609], MA
811806, OXF 00000255); Icon: Laurographia t. 7 (G
[F neg. 34254]).
Synonyms: Ocotea latifolia Kunth in Humboldt, Bonpland
and Kunth, Nov. Gen. Sp. (quarto ed.) 2: 165. 1817
≡ Nectandra latifolia (Kunth) Mez, Jahrb. Königl.
Bot. Gart. Berlin 5: 454. 1889. TyPE: COLOMBIA.
Cundinamarca, “in alta planitie Bogotensi,” s.d.,
F. W. H. A. von Humboldt & A. J. A. Bonpland s.n.
(Holotype: P 00128755).
Nectandra polita Nees & Mart. in Nees, Syst. Laur.:
325. 1836; Meissner in DC., Prodr. 15(1): 163. 1864;
ibid. in Martius, Fl. Bras. 5(2): 275. 1866. TyPE:
BRAZIL. Bahia, Cachoeira, “in sylvis Capões ad
Villa da Caxoeira,” November 1818, C. F. P. von
Martius s.n. (Holotype: M 0121031 [F neg. 19268];
Isotype, GZU 000249355). Specimens: BRAZIL.
VOL. 20, NO. 2
Bahia, Ilhéus, “in collib. pr. Ilheos,” May 1821, l.,
L. Riedel 307 (GOET, K 000642337, L 1802307 [as
Riedel s.n.], LE [2 sheets], Ny 00887019); idem, April
1837, l., B. Luschnath s.n. in Martii Herbar. Florae
Brasil. No.420 (B 10 0185159, BM 000947255, BR
0000008810489, BR 0000008810519, G, G-DC, GZU
000249354 [fragment], K 000642336, L 0037070,
LE 00006797, LE, M 0121032, MO 247434, Ny
00355493, P 00711025); idem, 1836, B. Luschnath
s.n. (BR 0000008808042).
Further synonyms in Rohwer (1993), and further
information on the priority of Ruiz and Pavón’s name over
Kunth’s name in Moraes (2012, 2013).
As pointed out by Moraes (2012a), Rohwer (1993)
indicated that Martii Herbar. Florae Brasil. No.420 would
probably be the same collection of Martius s.n., collected in
Cachoeira. However, Martius (1837–[1841]; 1841: 7) clearly
indicated “420. Oreodaphne dispersa Nees ab Esenb. Laur.
427 var. grandifolia ? ipso deiniente. … In silvis ad Ilheos.
Aprili. Dryas,” which does therefore represent a different
collection gathered by Luschnath.
29. Nectandra reticulata (Ruiz & Pav.) Mez, Jahrb. Königl.
Bot. Gart. Berlin 5: 404. 1889; Rohwer, Fl. Neotrop.
Monogr. 201: 276. 1993; Moraes, Komarovia 6(1): 24.
2008; ibid. in Harvard Pap. Bot. 17(1): 203. 2012; ibid. in
Pl. Ecol. Evol. 146(3): 370. 2013.
Basionym: Laurus reticulata Ruiz & Pav., Fl. Peruv. 4(2):
t. 348. 1804[–1830] (Ruiz and Pavón, 1956: 14, t.
348). Lectotype (designated by Rohwer, 1993: 276, as
“MA”; second-step lectotype designated by Moraes,
2008: 24): PERU. Cuchero, May[–September ?] 1780,
l., H. Ruíz L. & J. A. Pavón s.n. (MA 811776 [with
handwritten labels “Laurus reticulata Laurographia
Pavon et Sp. Pl. Fl. Per. / de Cuchero” by Pavón,
and “Enneandria Monogyn. Casyta ? Laurus F P. c.
L. N. 160 Ex Vitoc [Tarma]. Año 94” by Tafalla];
Isolectotypes, F [0040293F, ex MA, ex Vitoc,
fragments of leaf and lower], MA 811778 [numbered
“13/19”], MA 811780 [F neg. 29398; numbered
“13/19”]). Syntypes: PERU. Cuchero and Chinchao,
s.d., l., H. Ruíz L. & J. A. Pavón s.n. (A 00041960
[Pavón 504 ex G], B 10 0185150, BC 872724, BM
000947252, BR 0000008808370 [ex Herb. Reg.
Berolinense ex Herb. Lamberti], F [0040319F; F
neg. 58079], F [0040320F; F neg. 58078, with lower
and fruit], FI-W n.v., G 00020756 (Pavón 504, 2
sheets), G 00020755 & G 00020758 (Pavón s.n.; 7
sheets), G (4 sheets), GZU 000261284 [fragment],
HAL 0010378 [ex Herb. Reg. Berolinense ex Herb.
Lamberti], MA 811777 & MA 811779 [ex Herbarium
Peruvianum Ruiz et Pavon, numbered “13/20”], MA
811781 [Pavón’s label annotated as “Laurus reticulata
/ Laurographia Pavon et Sp. Pl. Fl. Per.”], P 00571439
[ex Herb. Pavon], P 00571440 [ex Herb. Pavon No.
36, annotated by Mez], P 00571441 [ex Herb. Reg.
Berolinense ex Herb. Lamberti]; PERU. Cuchero,
July, l., J. Dombey s.n. (B 10 0185149, G-DC, GZU
2015
DE MORAES AND FALCADE, LAURACEAE COLLECTED By MARTIUS
179
000254296 [fragment], MA 811775 [ex Herb. Ruiz &
Pavón, with Dombey’s label as “Laurus pubescens.
F. p. 1780/in Sylvis Cochero,” and “Enneandria
monogynia… /habitat in Sylvis Cochero, loret julio;”
see Alvarez Lopez, 1956: 75, and Ruiz and Pavón,
1989], P 00745320, P 00745321, P-JU 00662847;
also syntypes of Nectandra villosa Nees & Mart.
var. venosa Nees, Syst. Laur.: 291. 1836). Icon:
Laurographia t. 3 (G [F neg. 34250]).
Synonyms: Ocotea mollis Kunth in Humboldt, Bonpland
and Kunth, Nov. Gen. Sp. (quarto ed.) 2: 164. 1818
≡ Persea mollis (Kunth) Spreng., Syst. Veg. (ed. 16)
2: 270. 1825 ≡ Nectandra mollis (Kunth) Nees, Pl.
Laur. Expos. 14. 1833; ibid. in Syst. Laur.: 287. 1836;
Martius, Syst. Mat. Med. Bras.: 112. 1843; Meissner
in DC., Prodr. 15(1): 148. 1864; ibid. in Martius, Fl.
Bras. 5(2): 253, t. 91. 1866; Rohwer, Mitt. Inst. Allg.
Bot. Hamburg 20: 60. 1986. Lectotype (designated
by Rohwer, 1993: 276): COLOMBIA. Tolima,
“prope El Azufral in Andibus Quindiuensibus, alt.
1050 hexap.,” s.d., l., F. W. H. A. von Humboldt &
A. J. A. Bonpland s.n. (P 00128743; Isolectotypes,
B 10 0185155 [Mus. Bot. Berol. Film Nr. 5202],
B-W 07796 -01 0 [Mus. Bot. Berol. Film Nr. 657/33;
“Laurus villosa Willd.,” herbarium name] & B-W
07796 -00 0, F [0061634F; F neg. 63073], GZU
000254324 [fragment], Ny 00355629, Ny 00355630,
P 00128746, U 0002912, US 00028321). Specimens:
BRAZIL. Rio de Janeiro, April 1836, Martii Herbar.
Florae Brasil. No.169 [= Ackermann V no. 121] (AWH
28300, BR 0000008808080, BR 0000008808387,
G-DC 00147172, K 000576187, L 1802317, LE, M
0147273, M 0147274, M 0147275, M 0147276, MO
257354, OXF 00000288, STR).
Complete synonymy in Rohwer (1993) and further
information in Moraes (2013).
Vernacular name: “Canella preta” (Martius, 1843).
00000301, P 00756760) [Lectotype of Ocotea
petalanthera (Meisn.) Mez, Jahrb. Königl. Bot. Gart.
Berlin 5: 358. 1889].
Further information in Rohwer (1986) and Moraes
(2013).
32. Ocotea ceanothifolia (Nees) Mez, Jahrb. Königl. Bot.
Gart. Berlin 5: 365. 1889; Rohwer, Mitt. Inst. Allg. Bot.
Hamburg 20: 214. 1986 ≡ ‘Persea ceanothifolia Mart. ex
Nees’, Syst. Laur.: 236. 1836, nom. invalid.
Basionym: Mespilodaphne ceanothifolia Nees, Syst. Laur.:
236. 1836; Meissner in DC., Prodr. 15(1): 106. 1864;
ibid. in Martius, Fl. Bras. 5(2): 199. 1866. TyPE:
BRAZIL. Amazonas, “in sylvis Japurensibus ad
Porto dos Miranhas,” February 1820, l., C. F. P. von
Martius Obs. 3174: “Laurus ceanothifolia m. … Arbor
sive frutex ramis densis …” (Syntypes: B 10 0185262
[F neg. 3640], G 00369362, L [2 sheets; “Laurus
ceanothifolia M.,” herbarium name, “Japura”], M
0147148, M 0147150, Ny 00355332 [fragment]);
BRAZIL. Pará, “in sylvis ad Flumen Amazonum,”
September 1819, l., C. F. P. von Martius Obs. 3174
(Syntypes: M 0147149, Ny 00355332 [fragment]).
30. Ocotea bracteolata (Nees) Mez, Jahrb. Königl. Bot.
Gart. Berlin 5: 354. 1889; Rohwer, Mitt. Inst. Allg. Bot.
Hamburg 20: 216. 1986; Moraes, Pl. Ecol. Evol. 146(3):
372. 2013 ≡ Oreodaphne bracteolata Meisn., DC., Prodr.
15(1): 124. 1864; ibid. in Martius, Fl. Bras. 5(2): 220. 1866.
Basionym: Petalanthera bracteolata Nees, Syst. Laur.: 347.
1836. Lectotype (designated by Meissner [irst-step],
and by Mez [second-step]): BRAZIL. Amazonas, “in
sylvis aeternis Coari,” November–December 1819, l.,
C. F. P. von Martius s.n. (M 0147154; Isolectotypes,
B 10 0185271 [F neg. 3633], GZU 000254422, HBG
[fragment], M 0147155, Ny 00355873 [fragment]).
Syntypes: BRAZIL. Amazonas, Rio Japurá, “in sylvis
Japurensibus,” January–February 1820, l., C. F. P.
von Martius s.n. (GZU 000265483, M 0147153);
PERU. Loreto, “in Peruvia orient. prov. Maynas pr.
yurimaguas,” 1839, E. F. Poeppig 2426 (AWH 28268,
B† [F neg. 3702], F [0040353F], G 00369324, G
00369325, G-DC, GOET 004541, GZU 000254361,
KIEL [2 sheets], Ny 00355874 [fragment], OXF
33. Ocotea cernua (Nees) Mez, Mitt. Bot. Vereins Kreis
Freiburg 47/48: 422. 1888, s. lat.; ibid. in Jahrb. Königl.
Bot. Gart. Berlin 5: 377. 1889; Rohwer, Mitt. Inst. Allg.
Bot. Hamburg 20: 207. 1986; Moraes, Harvard Pap. Bot.
17(2): 247. 2012; ibid. in Pl. Ecol. Evol. 146(3): 371. 2013.
Basionym: Oreodaphne cernua Nees, Syst. Laur.: 424.
1836. Lectotype (designated by Mez, 1888: 422):
MARTINIQUE. Loco haud indicato, s.d., F. W. Sieber
Flora mixta. No 385 (KR; Isolectotypes, G 00369366,
G-DC 00130426 [only the lowering sheet], LE
00006768, LE [2 sheets], MEL, Ny 00074413, P
01959499). Syntypes: MARTINIQUE. Loco haud
indicato, s.d., F. Kohaut in F. W. Sieber Flora martin.
No 106 (AWH 28401, B 10 0185257, B 10 0185260,
B 10 0185261, BR 0000008813053, G 00369384, GH
00042066, GZU 000254342 [fragment], JE 00004386,
KIEL, L 0056294, L 0056295, LE [2 sheets], M
0147144, M 0147145, MO 277515); MARTINIQUE.
Loco haud indicato, s.d., J. W. Hornemann s.n. (C,
GZU 000260297 [fragment]).
31. Ocotea camphoromoea Rohwer, Mitt. Inst. Allg. Bot.
Hamburg 20: 179. 1986 ≡ ‘Persea subtriplinervia Mart.
ex Nees’, Syst. Laur.: 466. 1836, nom. invalid. ≡ Ocotea
subtriplinervia (Nees) Mez, Jahrb. Königl. Bot. Gart. Berlin
5: 333. 1889, nom. illegit.
Basionym: Camphoromoea subtriplinervia Nees, Syst.
Laur.: 466. 1836; Meissner in DC., Prodr. 15(1):
144. 1864; ibid. in Martius, Fl. Bras. 5(2): 246, t. 87.
1866. TyPE: BRAZIL. Pará, “in sylvis secus Flumen
Amazonum,” September 1819, l., C. F. P. von Martius
s.n. (Holotype: M 0147134; Isotypes, B 10 0185426,
M 0147135, M 0147136, M 0147137, M 0147138, M
0147139).
180
HARVARD PAPERS IN BOTANy
Synonyms: ? Laurus nitida Ruiz & Pav., Fl. Peruv. 4(2):
t. 353. 1804[–1830] (Ruiz and Pavón, 1956: 19, t.
353) ≡ Mespilodaphne nitida (Ruiz & Pav.) Meisn. in
DC., Prodr. 15(1): 106. 1864. Lectotype (designated
by Moraes, 2012b: 247): PERU. “in Chacahuassi,”
1787, H. Ruíz L. & J. A. Pavón s.n. (503 in G-Del)
(MA 811788; Isolectotypes, B 10 0247502 [B neg.
5266B], BM 000947285, F [0061414F; fragment],
F [0061419F; fragment], G [2 sheets], G [ex Herb.
Moricand, F neg. 27619], HAL 0103800, MA 811787,
Ny 00355164 [ex G ex Herb. Moricand], P 00756828
[Pavón 36]). Syntype: PERU. “Cuchero,” J. Dombey
s.n. (P 00756826, P 00756827). Icon: Laurographia t.
13 (G [F neg. 34260]).
? Laurus leptobotra Ruiz & Pav., Fl. Peruv. 4(2):
t. 368. 1804[–1830] (Ruiz and Pavón, 1956: 35, t.
368) ≡ Oreodaphne leptobotra (Ruiz & Pav.) Nees,
Linnaea 21: 520. 1848; Meissner in DC., Prodr. 15(1):
138. 1864; ibid. in Martius, Fl. Bras. 5(2): 239. 1866 ≡
Ocotea leptobotra (Ruiz & Pav.) Mez, Jahrb. Königl.
Bot. Gart. Berlin 5: 376. 1889. Lectotype (designated
by Moraes, 2012b: 247): PERU. “In Andium
nemoribus ad Macora praedium,” s.d., H. Ruíz L. &
J. A. Pavón s.n. (MA 811791; Isolectotypes, B 10
0247503 [B neg. 5266c], F [0061415F; fragment],
F [0061416F; fragment], G, GZU 000254142 [ex
B, fragment], MA 811790, Ny 00355162 [ex G,
fragment]). Icon: Laurographia t. 21 (G [F neg.
34268]).
? Leptodaphne subalpina Nees, Syst. Laur.: 361. 1836
≡ Oreodaphne subalpina (Nees) Meisn. in DC., Prodr.
15(1): 137. 1864; ibid. in Martius, Fl. Bras. 5(2): 238.
1866 ≡ Ocotea subalpina (Nees) Mez, Jahrb. Königl.
Bot. Gart. Berlin 5: 375. 1889. TyPE: BRAZIL.
Bahia, Rio de Contas, “Serra Grande prope Villam do
Rio de Contas,” October 1818, C. F. P. von Martius
s.n. (Holotype: M 0147140; Isotype, GZU 000254159
[fragment]).
? Leptodaphne tenuilora Nees, Syst. Laur.: 359.
1836 ≡ ‘Persea tenuilora Mart. ex Nees’, Syst. Laur.:
359. 1836, nom. invalid. ≡ Camphoromoea tenuilora
(Nees) Meisn. in DC., Prodr. 15(1): 144. 1864; ibid. in
Martius, Fl. Bras. 5(2): 247. 1866 ≡ Ocotea tenuilora
(Nees) Mez, Jahrb. Königl. Bot. Gart. Berlin 5: 383.
1889; Rohwer, Mitt. Inst. Allg. Bot. Hamburg 20:
208. 1986. TyPE: BRAZIL. Rio de Janeiro, “in sylva
Brasiliae densissima prope cataractas Tijucenses ad
Sebastianopolin,” November 1817, l., C. F. P. von
Martius Obs. 261: “Persea tenuilora Calyx clavato
infundibuliformis limbo 6ido …” (Holotype: M
0111179; Isotypes, B 10 0185423, GZU 000254160
[fragment], M 0111180).
For a complete list of probable synonyms, see Rohwer
(1986).
Rohwer (1986) has circumscribed Ocotea cernua (Nees)
Mez in a very broad sense. Once conirmed that the taxa
described and collected by Ruiz & Pavón in Peru are the
same as the one described by Nees from collections of
VOL. 20, NO. 2
Sieber “Flora Martinicensis & Mixta”, Ocotea leptobotra
(Ruiz & Pav.) Mez must be the correct name for having
priority (see Moraes, 2012b, 2013).
34. Ocotea citrosmoides (Nees) Mez, Jahrb. Königl. Bot.
Gart. Berlin 5: 349. 1889; Rohwer, Mitt. Inst. Allg. Bot.
Hamburg 20: 174. 1986.
Basionym: Oreodaphne citrosmoides Nees, Syst. Laur.:
384. 1836; Meissner in DC., Prodr. 15(1): 122. 1864;
ibid. in Martius, Fl. Bras. 5(2): 217. 1866. TyPE:
BRAZIL. Minas Gerais, Sabará, “in sylvis caeduis
prope Sabará,” May 1818, l. ♂, C. F. P. von Martius
Obs. 1223: “Arb. parva ramio paucis patul. Flores, qui
in siccis examinandi, pall. ochroleuco-viridialbi, anth.
tandem nigrescentibus. Prope Sabará” (Holotype: M
0111194; Isotypes, GZU 000254343, M 0111193).
35. Ocotea complicata (Meisn.) Mez, Jahrb. Königl. Bot.
Gart. Berlin 5: 247. 1889; Rohwer, Mitt. Inst. Allg. Bot.
Hamburg 20: 109. 1986; Moraes, Harvard Pap. Bot. 17(2):
248. 2012.
Basionym: Mespilodaphne complicata Meisn. in DC., Prodr.
15(1): 103. 1864; ibid. in Martius, Fl. Bras. 5(2): 199.
1866. Lectotype (designated by Assis and Mello Silva,
2010: 378): BRAZIL. Bahia, Ilhéus, “in sylv. arenosis
maritimis pr. Ilheos,” May 1822, L. Riedel 766 (G
00369363; Isolectotypes, B 10 0185255 [F neg. 3644],
F [0061435F, fragment], G [as Riedel 776], GOET,
K 000575058 [as Riedel s.n.], L 0036971 [as Riedel
s.n.], LE 00000570, Ny). Syntype: BRAZIL. Bahia,
“in sylvis Catingas ad Sincorâ locis altis,” November
1818, fr., C. F. P. von Martius s.n. (M 0111195 [F
neg. 19271; “Ocotea irmula Mart.,” herbarium name;
“Aydendron irmulum Nees,” herbarium name], M
0111196).
As pointed out by Moraes (2012b), Assis and MelloSilva (2010) reduced Ocotea complicata (Meisn.) Mez
to synonymy under O. fasciculata (Nees) Mez, which
is here reiteratively not accepted. The reason is based on
differences between both taxa in habit, lower, fruit and
vegetative characters, as well as in their habitats.
36. Ocotea cujumary Mart. in Buchner, Repert. Pharm. 35:
178. 1830; ibid. in Spix and Martius, Reise Bras. 3: 1128.
1831; Mez, Jahrb. Königl. Bot. Gart. Berlin 5: 286.1889;
Rohwer, Mitt. Inst. Allg. Bot. Hamburg 20: 161. 1986 ≡
Aydendron cujumary (Mart.) Nees, Syst. Laur.: 247. 1836;
Martius, Syst. Mat. Med. Bras.: 110. 1843; ibid. in Fl. Bras.
5(2): 318. 1868; Meissner in DC., Prodr. 15(1): 94. 1864;
ibid. in Martius, Fl. Bras. 5(2): 185, t. 66. 1866. TyPE:
BRAZIL. Amazonas, “in sylvis Japurensibus et ad Ega,
tum etiam in carapestribus sylvisque caeduis prope Barra
do Rio negro,” December 1819, fr., C. F. P. von Martius
s.n. (Holotype: M 0121076 [F neg. 6584]; Isotypes, B 10
0185244, M 0121077, Ny 00355591 [fragment]).
Complete synonymy in Rohwer (1986).
Vernacular name: “Cujumary” (Martius, 1843, 1868).
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DE MORAES AND FALCADE, LAURACEAE COLLECTED By MARTIUS
37. Ocotea cymbarum Kunth in Humboldt, Bonpland and
Kunth, Nov. Gen. Sp. (quarto ed.) 2: 166. 1817 ≡ Nectandra
cymbarum (Kunth) Nees, Syst. Laur.: 305. 1836; Martius,
Syst. Mat. Med. Bras.: 111. 1843; Meissner in DC., Prodr.
15(1): 156. 1864; ibid. in Martius, Fl. Bras. 5(2): 265, t.
96. 1866; Mez, Jahrb. Königl. Bot. Gart. Berlin 5: 464.
1889; Rohwer, Mitt. Inst. Allg. Bot. Hamburg 20: 106.
1986; ibid. in Fl. Neotrop. Monogr. 60: 298. 1993; Moraes,
Pl. Ecol. Evol. 146(3): 371. 2013 ≡ Licaria cymbarum
(Kunth) Pittier, Bol. Soc. Venez. Ci. Nat. 7: 135. 1941 ≡
Misanteca cymbarum (Kunth) Lundell, Wrightia 4(3):
100. 1969. Lectotype (designated by Moraes, 2013:
371): VENEZUELA. “in sylvis Orinocensibus prope San
Fernando de Atabapo,” May, fr., F. W. H. A. von Humboldt
& A. J. A. Bonpland 904 (B-W 07787 -01 0 [Mus. Bot.
Berol. Film Nr. 657/28], B-W 07787 -00 0 [annotated by
Willdenow as “Laurus coruscans” ex Bonpland, herbarium
name]; Isolectotype, P 00128760).
Synonym: Ocotea amara Mart. in Buchner, Repert. 35:
180. 1830. TyPE: BRAZIL. Amazonas, “in sylvis
aboriginibus Provinciae Fluminis nigri,” 1820, fr.,
C. F. P. von Martius s.n. (Holotype: M 0147301;
Isotypes, B 10 0185242 [sterile], L 0036963 [“Laurus
longifolia M. / Solimões”, herbarium name]).
Synonymy in Rohwer (1986: 106–107).
Vernacular name: “Páo Sassafras” (Martius, 1843).
38. Ocotea delexa Rohwer, Mitt. Inst. Allg. Bot. Hamburg
20: 214. 1986 ≡ Ocotea declinata (Meisn.) Mez, Jahrb.
Königl. Bot. Gart. Berlin 5: 352. 1889, nom. illegit., hom.
post.
Basionym: Oreodaphne declinata Meisn. in DC., Prodr.
15(1): 114. 1864; ibid. in Martius, Fl. Bras. 5(2): 208.
1866. TyPE: BRAZIL. Bahia, Ilhéus, September
1839, l., B. Luschnath s.n. in Martii Herbar. Florae
Brasil. No 1311 (Syntypes: BR [3 sheets]); BRAZIL.
Bahia, loco haud indicato, “partie merid. de la prov. de
Bahia,” s.d. [1840], l., J. S. Blanchet 3192 (Syntypes:
F [0061684F], G [4 sheets], G-DC 00201332 [F neg.
7280; Blanchet 3192A], Ny 00355707 [fragment],
OXF 00000309, P 00757003, W n.v.); idem, July
1839, l., B. Luschnath 32 (Syntypes: B 10 0185239,
B 10 0185240 [F neg. 3653], B 10 0185241, LE, M
0147123, M 0147124, M 0147125, M 0147126, M
0147127, Ny 00355708, Ny 00355709, P 00757004,
US 00047713).
39. Ocotea guianensis Aubl., Hist. Pl. Guiane 2: 781, t. 310.
1775 [spelled “Guyannensis” in the plate]; Rohwer, Mitt.
Inst. Allg. Bot. Hamburg 20: 157. 1986; Moraes, Pl. Ecol.
Evol. 146(3): 371. 2013 ≡ Laurus ocotea Rich., Actes Soc.
Hist. Nat. Paris 1: 108. 1792, nom. illegit. ≡ Oreodaphne
guianensis (Aubl.) Nees, Linnaea 21: 268, 516. 1848 ≡
Oreodaphne guianensis (Aubl.) Nees var. argentea Meisn.
in DC., Prodr. 15(1): 112. 1864, nom. illegit. superl., =
var. guianensis; ibid. in Martius, Fl. Bras. 5(2): 204. 1866.
Lectotype (designated by Rohwer, 1986: 157, as “Holotyp”):
FRENCH GUIANA. Loco haud indicato, April, l., J. B. F.
181
Aublet s.n. (BM 000947302). Syntype ?: PATRIA HAUD
INDICATA. Loco haud indicato, s.d. [June ?], fr., J. B. F.
Aublet s.n. (P 00756908). Specimen: BRAZIL. Pará, loco
haud indicato, May 1820, C. F. P. von Martius s.n. (M
0147117).
For a complete list of synonyms, see Rohwer (1986).
40. Ocotea hypoglauca (Nees & Mart.) Mez, Jahrb. Königl.
Bot. Gart. Berlin 5: 285. 1889; Rohwer, Mitt. Inst. Allg.
Bot. Hamburg 20: 191. 1986; Moraes, Komarovia 6(1): 26.
2008 ≡ Persea scrobiculata Meisn. in DC., Prodr. 15(1):
50. 1864, nom. illegit.; ibid. in Martius, Fl. Bras. 5(2): 161.
1866.
Basionym: Persea hypoglauca Nees & Mart. in Nees,
Linnaea 8: 49. 1833. Lectotype (designated by
Rohwer, 1986: 191): BRAZIL. Minas Gerais, “in
summo Monte Itacolomi 4500’ alt.,” fr., C. F. P. von
Martius Obs. 903: “arbor 10–15 ped. … In summo
Monte” (M 0111040 [annotated by Martius as “Persea
hypoglauca”, by Meissner as “Persea scrobiculata
nob. prodr.,” and by Mez as “Ocotea hypoglauca”];
Isolectotype, GZU 000254394). Syntypes: BRAZIL.
Paraná, l., F. Sellow 4803 (XXI) (B 10 0185196
[annotated by Nees], B 10 0185197, G 00020787
[as Sellow s.n.] [= Persea venosa Nees & Mart.]);
BRAZIL. Loco haud indicato, l., F. Sellow s.n. (B 10
0185313 [B neg. 1233/31; F neg. 3668]); idem, l., F.
Sellow s.n. (B 10 0185314 [B neg. 1233/32]).
41. Ocotea indecora (Schott) Mez, Jahrb. Königl. Bot. Gart.
Berlin 5: 249. 1889; Rohwer, Mitt. Inst. Allg. Bot. Hamburg
20: 110. 1986; Moraes, Komarovia 6(1): 14. 2008; ibid. in
Harvard Pap. Bot. 17(2): 251. 2012 ≡ Oreodaphne indecora
(Schott ) Nees, Syst. Laur.: 407. 1836 ≡ Mespilodaphne
indecora (Schott) Meisn. in DC., Prodr. 15(1): 102.
1864; ibid. in Martius, Fl. Bras. 5(2): 196, t. 72. 1866 ≡
Ocotea pretiosa Mez var. indecora (Schott) Vattimo-Gil,
Rodriguésia 37: 87. 1966.
Basionym: Persea indecora Schott in Spreng., Syst. Veg.
(ed. 16) 4(2): 405. 1827. Lectotype (designated by
Moraes, 2008: 14): BRAZIL. Loco haud indicato, s.d.,
l., H. W. Schott s.n. (B 10 0000017 [ex Herb. Sprengel
no. 43, B neg. 5254 B]). Syntype: BRAZIL. Loco
haud indicato, s.d., l., H. W. Schott 5610 (M 0121079
[F neg. 19273], M 0121080, Ny, US 00099223) [also
syntype of Mespilodaphne indecora (Schott) Meisn.
var. stricta Meisn. in DC., Prodr. 15(1): 102. 1864].
Probable syntype: BRAZIL. Loco haud indicato, s.d.,
l., H. W. Schott 5600 (M 0121078, Ny).
Synonym: ‘Aiouea tenuilora Nees ex Meisn.’ in DC.,
Prodr. 15(1): 510. 1864, nom. invalid.; Mez, Jahrb.
Königl. Bot. Gart. Berlin 5: 49. 1889. Specimen:
BRAZIL. Rio de Janeiro, “in sylvis caeduis ad
Sebastianopolin.,” September 1817, C. F. P. von
Martius s.n. (M 0147112).
Other synonyms in Assis and Mello-Silva (2010),
Moraes (2012b).
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HARVARD PAPERS IN BOTANy
42. Ocotea lancifolia (Schott) Mez, Jahrb. Königl. Bot. Gart.
Berlin 5: 289. 1889, s. lat.; Rohwer, Mitt. Inst. Allg. Bot.
Hamburg 20: 188. 1986; Moraes, Komarovia 6(1): 34. 2008;
ibid. in Harvard Pap. Bot. 17(2): 254. 2012 ≡ Oreodaphne
lancifolia (Schott) Nees, Syst. Laur.: 410. 1836; Meissner
in DC., Prodr. 15(1): 116. 1864; ibid. in Martius, Fl. Bras.
5(2): 210. 1866. Specimens: BRAZIL. Minas Gerais, Serra
do Caraça, May 1818, l., C. F. P. von Martius Obs. 1112
(M 0121035 [“Ocotea subacris Mart.,” herbarium name]);
idem, loco haud indicato, s.d., l., Anonymous s.n. (MEL
2324399; “L[aurus]. dura M[art].,” herbarium name).
Basionym: Persea lancifolia Schott in Spreng., Syst. Veg.
(ed. 16) 4(2): 405. 1827. TyPE: BRAZIL. Loco
haud indicato, l. ♂, H. W. Schott s.n. (Holotype: B
10 0000020 [B neg. 1234/1, F neg. 3675, Ny neg.
8379, ex Herb. Sprengel nr. 42, annotated as “Persea
lancifolia Schott”]).
Synonyms: Strychnodaphne lanceolata Nees, Linnaea 8:
39. 1833; Meissner in DC., Prodr. 15(1): 143. 1864;
ibid. in Martius, Fl. Bras. 5(2): 245. 1866; Moraes,
Komarovia 6(1): 53. 2008 ≡ Ocotea lanceolata
(Nees) Nees, Syst. Laur.: 474. 1836, non Nees in
Wallich (1831); Mez, Jahrb. Königl. Bot. Gart. Berlin
5: 334. 1889. TyPE: BRAZIL. Loco haud indicato,
s.d., immat fr., F. Sellow s.n. (Holotype: B† [possibly
F neg. 3675, ex Herb. Sprengel]; Isotypes, GZU
000249333 [annotated by Nees, Anonymous s.n.], VT
001552).
Oreodaphne martiana Nees, Linnaea 8: 41. 1833,
e.p., mixtum compositum; Meissner in DC., Prodr.
15(1): 134. 1864; ibid. in Martius, Fl. Bras. 5(2): 235.
1866. Lectotype (designated by Rohwer, 1986: 189,
as “Holotip M”): BRAZIL. Minas Gerais, “in alpinis
Serra do Caraça,” April 1818, l. ♀, C. F. P. von Martius
Obs. 1112: “Frutex 4–6 ped. ramis patentibus glab.
Serra de Carass.” (M 0111038 [“Ocotea daphnoides
Mart.,” herbarium name]; Isolectotypes, GZU
000259998 [fragment], M 0121036, Ny 00355742
[fragment]). Syntype: BRAZIL. Minas Gerais, loco
haud indicato, s.d. [April], fr., C. F. P. von Martius
s.n. (B 10 0185347, GZU 000254355 [fragment]) [=
Ocotea odorifera (Vell.) Rohwer, Mitt. Inst. Allg.
Bot. Hamburg 20: 111. 1986].
Oreodaphne nitidula Nees & Mart. in Nees, Linnaea
8: 41. 1833 ≡ Oreodaphne nitidula Nees & Mart. var.
lanceolata Meisn. in DC., Prodr. 15(1): 136. 1864,
nom. invalid. Lectotype (designated by Moraes, 2008:
53): BRAZIL. Minas Gerais, “editis Serro Frio, ad
Tapanhoacanga,” May 1818, l. ♂, C. F. P. von Martius
s.n. (M 0111039; Isolectotype: GZU 000254358).
Oreodaphne thymelaeoides Nees & Mart. in Nees,
Linnaea 8: 42. 1833. Lectotype (designated by
Moraes, 2008: 43): BRAZIL. Minas Gerais, Serro,
“in Serro Frio,” May 1818, l., C. F. P. von Martius
s.n. (M 0111037 [annotated by Martius as “Ocotea
daphnoides”, by Nees as “Oreodaphne angustifolia
thÿmelaeoides N. ab. Es. / nomen mutandum …,
ob Oreod. angustifol. Schott. in Spr. Cur. post.,”
and by Meissner as “Oreodaphne Martiana Nees β
VOL. 20, NO. 2
thymelaeoides Meissner ipse”]; Isolectotype, GZU
000254370 [annotated by Nees as “Oreodaphne
angustifolia thymelaeoides / Serro frio, Prov. Min. /
Majo”]).
Further synonyms in Rohwer (1986).
Rohwer (1986) has circumscribed Ocotea lancifolia in
a broad sense and in such terms that it is still needed to
be established whether several of the indicated synonyms
really belong to this taxon (i.e., Ocotea lancifolia).
In the protologue of Oreodaphne martiana, Nees has
described a species with lowers and fruits. The specimen at
B 10 0185347 (annotated and cited by Mez 1889: 250–251,
under Ocotea pretiosa Mez) bears a handwritten label by
Nees with “Oreodaphne martiana N. et M. Var. α / Min.
Ger. Apr. / a Mart.” However, it is a fruiting specimen
which belongs to Ocotea odorifera (Vell.) Rohwer, with a
duplicate in GZU. Therefore, Oreodaphne martiana Nees
must be taken only from the lowering material of Martius.
Another specimen cited by Mez (1889: 250–251, under
Ocotea pretiosa Mez), Martius “n. 79”, has not been located
so far.
43. Ocotea laxa (Nees) Mez, Jahrb. Königl. Bot. Gart.
Berlin 5: 381. 1889; Rohwer, Mitt. Inst. Allg. Bot. Hamburg
20: 211. 1986; Moraes, Komarovia 6(1): 8. 2008; ibid. in
Harvard Pap. Bot. 17(2): 264. 2012 ≡ ‘Persea laxa Mart. ex
Nees’, Syst. Laur.: 468. 1836, nom. invalid.
Basionym: Camphoromoea laxa Nees, Syst. Laur.: 468.
1836; Meissner in DC., Prodr. 15(1): 144. 1864; ibid. in
Martius, Fl. Bras. 5(2): 247, t. 88, 89. 1866. Lectotype
(designated by Moraes, 2008: 8): BRAZIL. Minas
Gerais, “in sylvis Capoeira prope Faz. de S. Barbara
et haud procul a Villa de Campanha,” February 1818,
l., C. F. P. von Martius Obs. 678: “arbor cortice dilute
cinereo. Altitudo 15–20 ped. Rami laxi dependentes.
In Capoeira” (M 0111035 [“Persea laxa Mart.,”
herbarium name]; Isolectotypes, B 10 0000022, GZU
000253885). Syntypes: BRAZIL. Minas Gerais e São
Paulo, “in sylvis caeduis prope luv. Sapucahy, prope
Camapuão”, l., C. F. P. von Martius s.n. (M 0111036
[F neg. 19275]); BRAZIL. Loco haud indicato, s.d.,
sterile, F. Sellow s.n. (B 10 0000024); idem, s.d.,
sterile, F. Sellow s.n. (B 10 0000025); idem, s.d.,
sterile, F. Sellow s.n. (HAL 0010445).
44. Ocotea longifolia Kunth in Humboldt, Bonpland and
Kunth, Nov. Gen. Sp. (quarto ed.) 2: 164. 1817; Rohwer,
Mitt. Inst. Allg. Bot. Hamburg 20: 158. 1986; Moraes,
Harvard Pap. Bot. 17(2): 254. 2012 ≡ Persea longifolia
(Kunth) Spreng., Syst. Veg. 2: 270. 1825 ≡ Oreodaphne
longifolia (Kunth) Nees, Syst. Laur.: 391. 1836. TyPE:
COLOMBIA. Tolima, “in saxosis inter Mariquitam
Novogranatensium et Santa Ana,” June 1801, fr., F. W. H.
A. von Humboldt & A. J. A. Bonpland 1729 (Holotype: P
00128764; Isotypes, B-W 07789-01 0 [B neg. 657/26] &
B-W 07789-00 0, P 00128765, P 00128766, P 00128767).
Synonyms: ? Laurus aurantiodora Ruiz & Pav., Fl. Peruv.
4(2): t. 349. 1804[–1830] (Ruiz and Pavón, 1956: 15,
t. 349) ≡ Aydendron suaveolens Nees var. β Nees,
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DE MORAES AND FALCADE, LAURACEAE COLLECTED By MARTIUS
Linnaea 21: 499. 1848, nom. invalid. ≡ Mespilodaphne
aurantiodora (Ruiz & Pav.) Meisn. in DC., Prodr.
15(1): 101. 1864 ≡ Ocotea aurantiodora (Ruiz &
Pav.) Mez, Jahrb. Königl. Bot. Gart. Berlin 5: 295.
1889. Lectotype (designated by Moraes, 2012b: 255):
PERU. “in Peruviae Andium nemoribus ad Chinchao
et Cuchero,” 1794, H. Ruíz L. & J. A. Pavón s.n. (MA
811782; Isolectotypes, B 10 0247398 [B neg. 5215], F
[0061413F; fragment], FI 005224, G, MA [2 sheets]).
Icon: Laurographia t. 15 (G [F neg. 34262]).
Ocotea opifera Mart. in Buchner, Repert. 35: 179.
1830; Mez, Jahrb. Königl. Bot. Gart. Berlin 5: 291.
1889; Rohwer, Mitt. Inst. Allg. Bot. Hamburg 20: 158.
1986; Moraes, Harvard Pap. Bot. 17(2): 255. 2012 ≡
Oreodaphne opifera (Mart.) Nees, Syst. Laur.: 390.
1836; Martius, Syst. Mat. Med. Bras.: 110. 1843 ≡
Mespilodaphne opifera (Mart.) Meisn. in DC., Prodr.
15(1): 510. 1864; ibid. in Martius, Fl. Bras. 5(2):
194, t. 70. 1866; Martius, Fl. Bras. 5(2): 318. 1868 ≡
‘Laurus opifera Mart. ex Meisn.’ in Martius, Fl. Bras.
5(2): 195. 1866, nom. invalid. Lectotype (designated
by Rohwer, 1986: 158, as “Holotyp M”): BRAZIL.
Amazonas, “in sylvis secus luv. Amazonum et Rio
Negro,” October 1819, fr., C. F. P. von Martius Obs.
2796: “Arbor 10–15 pedalis loribus in paniculis
magnis terminalibus pallide lavo-ochroleucis,
fructibus ellipticis in cupulis hemisphaericis.
Rami angulati” (M 0121057). Syntypes: BRAZIL.
Amazonas, “in sylvis ad Barra do Rio Negro,”
October 1819, fr., C. F. P. von Martius Obs. 2796
(GZU 000254327, M 0121055); idem, Amazonas, “in
sylvis Japurensibus” [Rio Japurá], December 1819,
l., C. F. P. von Martius Obs. 2796 (GZU 000254326,
M 0121056, M 0174126); idem, Amazonas, “in sylvis
Japurensibus” [Rio Japurá], January 1820, l., C. F. P.
von Martius s.n. (M 0121058); idem, Pará, “in sylvis
secus l. Amazonum,” May 1820, C. F. P. von Martius
s.n. (B 10 0185382 [fr.], M 0121059 [l.], M 0121061
[l.]); idem, loco haud indicato, s.d., l., Anonymous
s.n. (B 10 0185383 [annotated by Nees]).
Further synonyms in Rohwer (1986).
Vernacular name: “Canella de cheiro” (Martius, 1843,
1868).
Mez (1889), Rohwer (1986), and van der Werff
(2002b) have indicated Ocotea longifolia Kunth and O.
aurantiodora (Ruiz & Pav.) Mez as probable synonyms.
However, since there is no recent revision of the collections
of these taxa (of specimens from Brazil, Colombia, Peru,
and elsewhere), we keep here the traditional use of the irst
name for the Brazilian specimens, indicating the second
name as a doubtful synonym. Once the synonymy is
conirmed, Ocotea aurantiodora must be the correct name
to be adopted for being the oldest one (Moraes, 2012b).
45. Ocotea minarum (Nees & Mart.) Mez, Jahrb. Königl.
Bot. Gart. Berlin 5: 305. 1889; Rohwer, Mitt. Inst. Allg.
Bot. Hamburg 20: 148. 1986; Moraes, Komarovia 6(1): 13.
2008; ibid. in Harvard Pap. Bot. 17(2): 256. 2012.
183
Basionym: Gymnobalanus minarum Nees & Mart. in Nees,
Linnaea 8: 38. 1833; ibid. in Syst. Laur.: 480. 1836;
Meissner in DC., Prodr. 15(1): 140. 1864; ibid. in
Martius, Fl. Bras. 5(2): 242, t. 84. 1866. Lectotype
(designated by Rohwer, 1986: 148 [irst-step];
second-step lectotype designated by Moraes, 2008:
12): BRAZIL. São Paulo, l. ☿, F. Sellow 5636 (B
10 0000037 [B neg. 5276 B]; Isolectotypes, B 10
0000036, B 10 0000038, VT). Probable isolectotypes,
F [0061409F; as Sellow s.n.], HAL 0010433 [as
Sellow s.n.]. Other syntypes: BRAZIL. Minas Gerais,
“in sylvis ad Mariana,” April 1818, l. ♀, C. F. P.
von Martius s.n. (GZU 000253871, M 0111045);
BRAZIL. Minas Gerais, Serro, “in sylvis capões
Serro Frio,” May 1818, l. ♀, C. F. P. von Martius
s.n. (GZU 000253871, M 0111044); BRAZIL. São
Paulo, “campis ad ypanema et ytu” [Real Fábrica
de Ferro de São João de ypanema, currently Floresta
Nacional de Ipanema, Iperó; Itu], January 1818, fr., C.
F. P. von Martius s.n. (GZU 000253871 [fragment], M
0111041 [“Ocotea tubigera Mart.,” herbarium name]).
Possible synonyms in Rohwer (1986).
46. Ocotea neesiana (Miq.) Kosterm., Meded. Bot. Mus.
Herb. Utrecht 25: 16. 1936; Rohwer, Mitt. Inst. Allg. Bot.
Hamburg 20: 183. 1986; Moraes, Komarovia 6(1): 32. 2008;
ibid. in Harvard Pap. Bot. 17(2): 257. 2012; ibid. in Pl. Ecol.
Evol. 146(3): 372. 2013 ≡ Oreodaphne fallax Miq., Stirp.
Surinam. Select.: 203. 1851, nom. illegit. superl. ≡ Ocotea
fallax Miq. ex Mez, Mitt. Bot. Vereins Kreis Freiburg 47/48:
422. 1888, nom. illegit. superl.
Basionym: Nectandra neesiana Miq., Linnaea 18: 745.
1844. TyPE: SURINAME. “distr. Para,” February–
April 1844, l. ♂, F. W. R. Hostmann & A. Kappler
1433 (Holotype: U 0008441; Isotypes, AWH 28263,
B† [ex Herb. Döll, ide Mez 1888: 422; not located
in KR], G 00021042, G 00021043, GOET 000744,
JE 00004381, KIEL, LE, P 00571434, P 00571435,
P 01964216).
Synonyms: Oreodaphne japurensis Meisn. in DC., Prodr.
15(1): 124. 1864; ibid. in Martius, Fl. Bras. 5(2): 219.
1866; Mez, Jahrb. Königl. Bot. Gart. Berlin 5: 309.
1889 ≡ Oreodaphne dispersa Nees & Mart. in Nees,
Linnaea 8: 43. 1833, e.p. TyPE: BRAZIL. Amazonas,
“Rio Negro, in sylvis Japurensibus,” l. ♀, C. F. P. von
Martius s.n. (Holotype: M 0111069; Isotypes, GZU
000254347, L [“Laurus paucilora M.,” herbarium
name, “R. Negro;” ex Herbarium Reinwardtianum],
M 0111070, MEL 2318483 [“Laurus paucilora,”
herbarium name, “Para”], Ny 00355728 [fragment])
[also syntype of Oreodaphne dispersa Nees & Mart.].
Further synonyms in Rohwer (1986) and Moraes (2008,
2012b).
The specimen at L seems slightly different from the
others at GZU, M, MEL, and Ny. It was identiied by
Kostermans in February 1935 as “Ocotea tenuilora (Nees)
Mez”, but deinitely does not belong to it (see entry number
33, this work).
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47. Ocotea nutans (Nees) Mez, Jahrb. Königl. Bot. Gart.
Berlin 5: 362. 1889; Rohwer, Mitt. Inst. Allg. Bot. Hamburg
20: 196. 1986; Moraes, Komarovia 6(1): 33. 2008; ibid.
in Harvard Pap. Bot. 17(2): 259. 2012 ≡ Mespilodaphne
nutans (Nees) Meisn. in DC., Prodr. 15(1): 98. 1864; ibid.
in Martius, Fl. Bras. 5(2): 189. 1866.
Basionym: Oreodaphne nutans Nees, Syst. Laur.: 421.
1836. TyPE: BRAZIL. São Paulo, “In campis
Provinciae S. Pauli,” January 1818, l. ♀, C. F. P. von
Martius s.n. (Holotype: M 0111192; Fig. 1; Isotypes,
L [“Laurus paulina M.,” herbarium name, “S. Paulo;”
ex Herbarium Reinwardtianum; Fig. 2], MEL [“L.
paulina M.,” herbarium name, “St. Pauli”]).
Synonyms: Oreodaphne nutans Nees var. sylvestris Nees,
Syst. Laur.: 421. 1836 ≡ Mespilodaphne glauca (Nees)
Meisn. var. virescens Meisn. in DC., Prodr. 15(1): 98.
1864, nom. illegit.; ibid. in Martius, Fl. Bras. 5(2):
188. 1866. TyPE: BRAZIL. Minas Gerais, Serro, “in
sylvis Capões ad Tapanhoacanga, Serro Frio,” May
1818, l. ♀, C. F. P. von Martius s.n. (Holotype: M
0111191).
Mespilodaphne glauca (Nees) Meisn. var. virescens
Meisn. in DC., Prodr. 15(1): 98. 1864, excl. syn.
Oreodaphne nutans Nees var. sylvestris Nees ≡
Ocotea glauca (Nees) Mez var. virescens (Meisn.)
Mez, Jahrb. Königl. Bot. Gart. Berlin 5: 362. 1889.
TyPE: BRAZIL. Bahia, “circa Igreja Velha,” s.d.,
l., J. S. Blanchet 3250 (Syntypes: B 10 0185330,
BM 000993989, F [0061438F; F neg. 58155], G
00369419, G 00369420, G-DC 00200476, M 0147118
[F neg. 19272]); Bahia, Jacobina, s.d., l., J. S.
Blanchet 3250B (Syntypes: BR 0000008823410, BR
0000008823021, BR 0000008823366, F [0061439F],
FI-W 005242, G 00369421, G [2 sheets], G-DC
00200475, K 000442015, KIEL, M 014 7118 [F neg.
19272], MO 247521, Ny 00355343, P 00756893, P
00756894, P 00756895, P 00756896); idem, Igreja
Velha, s.d., J. S. Blanchet 301 (Syntypes: G, P).
Oreodaphne kunthiana Meisn. in DC., Prodr. 15(1):
119. 1864; ibid. in Martius, Fl. Bras. 5(2): 213. 1866.
TyPE: BRAZIL. Loco haud indicato, s.d., l. ♂, F.
Sellow s.n. (Holotype: B 10 0185388 [ex Herb. Kunth,
B neg. 5286 B]; Isotypes, G 00021062, P 01958844).
Ocotea subdecumbens M.A. Lisboa & J. Badini,
Rev. Esc. Minas 31(4): 32. 1974; Moraes, Harvard
Pap. Bot. 17(2): 259. 2012. TyPE: BRAZIL. Minas
Gerais, Mariana, entre Bento Rodrigues e Sta. Rita
Durão, February 1971, l. ♂, M. A. Lisboa & J. Badini
s.n. (Holotype: OUPR 26600 [right-hand specimen;
ex Herbário da Escola de Minas de Ouro Preto No.
4030]; Isotype, OUPR 18212); idem, Minas Gerais,
Mariana, entre Bento Rodrigues e Sta. Rita Durão,
February 1971, l. ♀, M. A. Lisboa & J. Badini s.n.
(Paratype: OUPR 26600 [left-hand specimen; ex
Herbário da Escola Federal de Farmácia e Bioquímica
de Ouro Preto No. 27003], HRCB).
VOL. 20, NO. 2
48. Ocotea phillyreoides (Nees) Mez, Jahrb. Königl. Bot.
Gart. Berlin 5: 315. 1889; Rohwer, Mitt. Inst. Allg. Bot.
Hamburg 20: 168. 1986; Moraes, Abc Taxa 3: 126. 2007;
ibid. in Komarovia 6(1): 36. 2008; ibid. in Harvard Pap. Bot.
17(2): 260. 2012 ≡ ‘Cryptocarya monticola Mart. ex Nees’,
Syst. Laur.: 400. 1836, nom. invalid. ≡ Mespilodaphne
phillyreoides (Nees) Meisn. in DC., Prodr. 15(1): 100. 1864;
ibid. in Martius, Fl. Bras. 5(2): 193. 1866 ≡ Mespilodaphne
phillyreoides Nees (Meisn.) var. oblonga Meisn. in DC.,
Prodr. 15(1): 100. 1864 [homotypic by typiication;
lectotype designated by Moraes, 2008: 36]; ibid. in Martius,
Fl. Bras. 5(2): 193. 1866.
Basionym: Oreodaphne phillyreoides Nees, Syst. Laur.:
400. 1836. Lectotype (designated by Moraes, 2008:
36): BRAZIL. São Paulo, “Rio das Pedras,” l. ♂, F.
Sellow B 1362 c 393 (B 10 0185369; Isolectotypes, B
10 0185367 [B neg. 5295], K 000575057). Probable
isolectotypes: BRAZIL. Loco haud indicato, l. ♂, F.
Sellow s.n. (G-DC 00201230, HAL 0103830); idem,
loco haud indicato, l., F. Sellow s.n. (B 10 0185370,
ex Herb. Baschant). Syntype: BRAZIL. Minas
Gerais, “in montosis, alt. 2800–3600’,” “Cryptocarya
monticola Martius” [herbarium name], l. ♀, immat.
fr., C. F. P. von Martius s.n. (M 0111178 [annotated by
Nees as “Oreodaphne philÿraeoides” and by Meissner
as “Mespilodaphne phillyreoides β oblonga”).
Further synonyms and information in Rohwer (1986)
and Moraes (2008).
Ocotea phillyreoides (Nees) Mez belongs to the Ocotea
tristis (Nees & Mart.) Mez Complex sensu Rohwer (1986),
whose synonymy must be conirmed.
49. Ocotea pulchella (Nees & Mart.) Mez, Jahrb. Königl.
Bot. Gart. Berlin 5: 317. 1889 ≡ Mespilodaphne pulchella
(Nees & Mart.) Meisn. in DC., Prodr. 15(1): 99. 1864; ibid.
in Martius, Fl. Bras. 5(2): 191, t. 68. 1866; Rohwer, Mitt.
Inst. Allg. Bot. Hamburg 20: 166. 1986; Moraes, Komarovia
6(1): 38. 2008; ibid. in Harvard Pap. Bot. 17(2): 261. 2012.
Basionym: Oreodaphne pulchella Nees & Mart. in Nees,
Linnaea 8: 40. 1833; ibid. in Syst. Laur.: 397. 1836;
Martius, Flora 20(2, Beibl.): 102. 1837. Lectotype
(designated by Moraes, 2008: 39): BRAZIL. São
Paulo, “inter ypanema et ytu, in Capões,” January–
February 1818, l. ♂, C. F. P. von Martius Obs. 595: “...
arbor 6–10 ped. …” (M 0121047 [annotated by Nees]).
Syntypes: BRAZIL. São Paulo, “in campestribus prope
Caja pintada versus Civ. S. Pauli,” December 1817,
l. ♀, C. F. P. von Martius Obs. 512: “...arbor 10–16
ped. in Campis” (M 0121046, M 0121048 [annotated
by Martius as “in virgultetis prope Civ. S. Pauli”]);
BRAZIL. São Paulo, fr., F. Sellow 144 (B 10 0185456);
BRAZIL. Minas Gerais, Serra da Moeda, fr., F. Sellow
B 1378 c 413 (B 10 0185472, B 10 0185471) [syntype
of Mespilodaphne pulchella (Nees & Mart.) Meisn.
var. ferruginea Meisn. in DC., Prodr. 15(1): 100.
1864]; BRAZIL. Rio Grande do Sul, l. ♀, immat. fr.,
F. Sellow d 2324 (B 10 0185458); BRAZIL. São Paulo,
l., F. Sellow 459 (B 10 0185455). Probable syntype:
2015
DE MORAES AND FALCADE, LAURACEAE COLLECTED By MARTIUS
fiGure 1. Oreodaphne nutans Nees: Holotype (M 0111192).
185
186
HARVARD PAPERS IN BOTANy
fiGure 2. Oreodaphne nutans Nees: Isotype (L, ex Herbarium Reinwardtianum).
VOL. 20, NO. 2
2015
DE MORAES AND FALCADE, LAURACEAE COLLECTED By MARTIUS
BRAZIL. Loco haud indicato, fr., F. Sellow s.n. (TUB
019041). Specimens: BRAZIL. Goiás, “prope Rio
Claro,” June, l. ♀, fr., Martii Herbar. Florae Brasil.
No.117 [= A. L. P. da Silva Manso 278] (B 10 0185473,
BR 000000880656, E 00259510, G 00021044, G-DC
00200973, K 000512688, LE 00006778, M 0121049,
Ny 00355381 [Ny 00355365], P 00571418) [syntype
of Mespilodaphne pulchella (Nees & Mart.) Meisn.
var. elliptica Meisn. in DC., Prodr. 15(1): 100. 1864].
Further synonyms and additional information in Rohwer
(1986) and Moraes (2008, 2012b).
50. Ocotea spixiana (Nees) Mez, Jahrb. Königl. Bot. Gart.
Berlin 5: 260. 1889; Rohwer, Mitt. Inst. Allg. Bot. Hamburg
20: 193. 1986 ≡ ‘Ocotea rufotomentosa Mart.’, nomen.
Basionym: Oreodaphne spixiana Nees, Syst. Laur.: 382.
1836; Meissner in DC., Prodr. 15(1): 117. 1864;
ibid. in Martius, Fl. Bras. 5(2): 212. 1866. TyPE:
BRAZIL. Minas Gerais, Serro, “in sylvis Capões,
locis montosis, Serro Frio,” May 1818, l., C. F. P. von
Martius s.n. (Holotype: M 0111182 [F neg. 6587];
Isotypes, GZU 000254368, M 0111181 [annotated by
Nees as “Aperiphracta martiana,” herbarium name],
M 0111183, M 0111184, Ny 00355782 [fragment]).
51. Ocotea tabacifolia (Meisn.) Rohwer, Mitt. Inst. Allg.
Bot. Hamburg 20: 173. 1986; Moraes, Harvard Pap. Bot.
17(2): 263. 2012.
Basionym: Persea tabacifolia Meisn. in DC., Prodr. 15(1):
51. 1864. TyPE: BRAZIL. Minas Gerais, “monte
Itambé,” November 1820, immat. fr., J. B. E. Pohl
3604 (Holotype: W†; Lectotype, designated by
Moraes, 2008: 47: Ny 00355862).
Synonyms: Oreodaphne umbrosa Nees, Syst. Laur.: 388.
1836, non (Kunth) Nees, Syst. Laur.: 456. 1836;
Meissner in DC., Prodr. 15(1): 122. 1864; ibid. in
Martius, Fl. Bras. 5(2): 216. 1866 ≡ Ocotea umbrosa
(Nees) Mart. ex Mez, Jahrb. Königl. Bot. Gart.
Berlin 5: 350. 1889, non Kunth, 1818 ≡ Oreodaphne
minarum Steud., Nomencl. Bot. (ed. 2) 2: 226. 1841.
TyPE: BRAZIL. Minas Gerais, “inter Taparoca et
Tapanhoacanga, locis altis sylvaticis,” 13 May 1818,
l. ♂, C. F. P. von Martius Obs. 1188: “arbor alt.
10–15 ped. … In Capoeira” (Holotype: M 0111067;
Isotypes, GZU 000254371 [fragment], M 0111068,
MEL 2324416).
Further synonym in Rohwer (1986).
52. Ocotea teleiandra (Meisn.) Mez, Jahrb. Königl. Bot.
Gart. Berlin 5: 382. 1889 ≡ Oreodaphne teleiandra Meisn.
in DC., Prodr. 15(1): 138. 1864; ibid. in Martius, Fl. Bras.
5(2): 239. 1866; Rohwer, Mitt. Inst. Allg. Bot. Hamburg
20: 211. 1986; Moraes, Komarovia 6(1): 55. 2008; ibid. in
Harvard Pap. Bot. 17(2): 263. 2012.
Basionym: Teleiandra glauca Nees & Mart. in Nees,
Linnaea 8: 46. 1833. TyPE: BRAZIL. São Paulo, l.
♂, F. Sellow 399 (Holotype: B 10 0000045; Isotypes,
B 10 0000044, B 10 0000046 [ex Herb. Kunth, B neg.
5322], K 000442012, VT 001553). Probable isotypes:
187
BRAZIL. Loco haud indicato, l. ♂, F. Sellow s.n. (BR
0000008767165, F [0061819F; fragment], G, GZU
000249332); idem, l., F. Sellow s.n. (HAL 0103829,
wrongly mounted with the label of “Oreodaphne
glauca Nees ab Es.!” written by Schlechtendal).
Synonyms: Camphoromoea venulosa Nees, Syst. Laur.: 469.
1836 ≡ Oreodaphne venulosa (Nees) Meisn. in DC.,
Prodr. 15(1): 126. 1864; ibid. in Martius, Fl. Bras.
5(2): 222. 1866 ≡ Ocotea venulosa (Nees) Baitello,
Fl. Fanerog. Estado São Paulo 3: 208. 2003. TyPE:
BRAZIL. São Paulo, “in sylvis Montis formosi”
[Morro Formoso, prope Bananal], December 1817, C.
F. P. von Martius s.n. (Holotype: M 0147133 [F neg.
19278]; Isotype, GZU 000253888).
As pointed out by Moraes (2012b: 264), Rohwer (1986:
211) synonymized Ocotea teleiandra (Meisn.) Mez with
Ocotea laxa (Nees) Mez. However, according to Baitello and
Marcovino (2003: 195, 197, 205), although both are similar in
vegetative characters, O. teleiandra may be distinguished by
tepals thick and papilose in the inner surface, leaves glabrous
abaxially, absence of bullate and barbellated domatia, and
larger fruits with cupule of smooth margin, whereas O. laxa
has leaves pubescent abaxially, with conspicuous domatia,
and fruits with hexalobed cupules.
53. Ocotea tristis (Nees & Mart.) Mez, Jahrb. Königl.
Bot. Gart. Berlin 5: 316. 1889; Rohwer, Mitt. Inst. Allg.
Bot. Hamburg 20: 167. 1986; Moraes, Komarovia 6(1):
43. 2008; ibid. in Harvard Pap. Bot. 17(2): 264. 2012 ≡
Mespilodaphne tristis Meisn. in DC., Prodr. 15(1): 100.
1864; ibid. in Martius, Fl. Bras. 5(2): 193, t. 69. 1866 ≡
Oreodaphne tristis Nees & Mart. var. oblongifolia Nees &
Mart. in Nees, Linnaea 8: 40. 1833, nom. illegit. superl., =
var. tristis.
Basionym: Oreodaphne tristis Nees & Mart. in Nees,
Linnaea 8: 40. 1833. Lectotype (designated by
Moraes, 2008: 45): BRAZIL. Minas Gerais, Serro,
“Serro Frio,” “Ocotea tristis s. Aeneae (Virg. VI.)
493.) In campis tristibus, ubi animae illorum, quos
amor peredit…,” May 1818, l., C. F. P. von Martius
s.n. (M 0111177).
Further synonyms and detailed information in Rohwer
(1986) and Moraes (2008, 2012b).
54. Ocotea variabilis (Nees) Mez, Jahrb. Königl. Bot. Gart.
Berlin 5: 288. 1889; Rohwer, Mitt. Inst. Allg. Bot. Hamburg
20: 189. 1986 ≡ ‘Ocotea variabilis Mart. ex Nees’, Syst.
Laur.: 429. 1836, nom. invalid.
Basionym: Oreodaphne variabilis Nees, Syst. Laur.: 429.
1836; Meissner in DC., Prodr. 15(1): 116. 1864;
ibid. in Martius, Fl. Bras. 5(2): 211. 1866. TyPE:
BRAZIL. Minas Gerais, Serro, “in campestribus
altis Serro Frio,” May 1818, C. F. P. von Martius s.n.
(Holotype: M 0121081 [F neg. 6588]; Isotypes, B 10
0185417 [F neg. 3737], GZU 000254373, L [“Laurus
adamantina M.,” herbarium name, “Diam. District,”
ex Herbarium Reinwardtianum], M 0121082, MEL
2324404 [“Laurus adamantina Mart.,” herbarium
name, “Serro Frio”).
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VOL. 20, NO. 2
Although Rohwer (1986) had proposed Ocotea variabilis
as a synonym of Ocotea lancifolia (Schott) Mez, their fruits
are different, mainly in relation to the cups, being cupolashaped, with simple margin, without remnant tepals in the
former, and dish-shaped, with double margin, and with
remnant tepals in the latter, making it possible to distinguish
and take them as belonging to different taxa.
TyPE: BRAZIL. Minas Gerais, “inter frutices in
planitie Chapada do Paranán,” September 1818, fr.,
C. F. P. von Martius Obs. 1770: “Frutex 4–6 pedalis,
foliis subcomplicatis subtus glauco rore adspersis,
calycibus roseo-igneis. In camp. Carrasco” (Holotype:
M 0111175 [F neg. 6589]; Isotypes, GZU 000254298,
M 0111176).
55. Ocotea velutina (Nees) Rohwer, Mitt. Inst. Allg. Bot.
Hamburg 20: 172. 1986; Moraes, Komarovia 6(1): 47. 2008.
Basionym: Oreodaphne velutina Nees, Syst. Laur.: 386.
1836; Meissner in DC., Prodr. 15(1): 132. 1864;
ibid. in Martius, Fl. Bras. 5(2): 231, t. 79. 1866;
Mez, Jahrb. Königl. Bot. Gart. Berlin 5: 348. 1889.
Lectotype (designated by Rohwer, 1986: 172):
BRAZIL. Minas Gerais, “in Sumo Brasiliae Monte
Itambé, Serro Frio,” May 1818, l. ☿–♂, C. F. P.
von Martius s.n. (M 0121084 [annotated by Martius
with “Ocotea velutina” and by Nees as “Aperiphracta
velutina”]; Isolectotypes, GZU 000254374 [ex Herb.
Nees, annotated by Nees as “Aperiphracta velutina,”
“hermaphr.-fem.,” “In summo Bras. Monte Itambé
Serro Frio... Maio”], GZU 000254375 [fragments]).
Syntypes: BRAZIL. Minas Gerais, “in montosis
sylvaticis prope Taparaoca präd. Serro Frio,” May
1818, l. ☿–♀, C. F. P. von Martius Obs. 1183:
“arbor 15–20 ped. ram. paucis distantibus, …” (GZU
000260730 [fragment], M 0121085); BRAZIL.
Minas Gerais, loco haud indicato, s.d., l., C. F. P.
von Martius s.n. (M 0121086); BRAZIL. São Paulo,
Iperó, “ypanema,” fr., F. Sellow B 1384 c 419 (B
10 0185416 [Ny neg. 8383]); BRAZIL. São Paulo,
“a S. Paulo ad meridiem,” l., F. Sellow 5867 (B 10
0185415 [Ny neg. 8384], K 000602367); BRAZIL.
São Paulo, F. Sellow 5928 (B†); BRAZIL. Loco haud
indicato, fr., F. Sellow s.n. (B 10 0185414); idem, fr.,
F. Sellow s.n. (CGE); idem, l., fr., F. Sellow s.n. (E
00259499); idem, fr., F. Sellow s.n. (HAL 0010486);
idem, fr., F. Sellow s.n. (LE 00006785).
Further synonyms in Rohwer (1986).
58. Persea americana Mill., Gard. Dict. (ed. 8) s.n. 1768;
Kopp, Mem. New york Bot. Gard. 14: 15. 1966; Moraes,
Harvard Pap. Bot. 17(1): 204. 2012; ibid. in Pl. Ecol. Evol.
146(3): 372. 2013.
Basionym: Laurus persea L., Sp. Pl. 1: 370. 1753. Lectotype
(designated by van der Werff, 2002a: 580): “Prunifera
arbor, fructu maximo pyriformi viridi, pericarpio
esculento butyraceo nucleum unicum maximum
nullo ossiculo tectum, cingente. The Avocado or
Alegator pear-tree,” in Sloane (1725: 132, t. 222, ig.
2). “Typotype” (original material): JAMAICA. Herb.
Sloane Vol. VII: 77 (BM-SL 000594115).
Synonym: Persea gratissima Gaertn., Fruct. Sem. Pl. 3:
222, t. 221. 1807, nom. illegit. superl. for Persea
americana Mill.; Nees, Syst. Laur.: 128. 1836;
Martius, Syst. Mat. Med. Bras.: 26, 112. 1843; ibid.
in Fl. Bras. 5(2): 320. 1868; Meissner in DC., Prodr.
15(1): 52. 1864; ibid. in Martius, Fl. Bras. 5(2): 158, t.
105, ig. 1. 1866; Mez, Jahrb. Königl. Bot. Gart. Berlin
5: 145. 1889. Specimen: BRAZIL. Pará, “circa Pará,”
“in sylvis archipelagi Insularum Paraënsium,” August
1819, C. F. P. von Martius s.n. (GZU 000254384, M).
Further list of synonyms in van der Werff (2002a).
As pointed out by van der Werff (2002a), Kopp (1966:
17) designated the description in Clusius (1601: 2) as
lectotype of Laurus persea. However, since descriptions
are not accepted as lectotypes, according to Art. 8.1 of ICN
(McNeill et al., 2012), the illustration in Sloane (1725)
has been designated as lectotype of Laurus persea L. and,
therefore of Persea americana Mill.
56. Ocotea virgultosa (Nees) Mez, Jahrb. Königl. Bot. Gart.
Berlin 5: 252. 1889; Rohwer, Mitt. Inst. Allg. Bot. Hamburg
20: 226. 1986; ibid. in Fl. Neotrop. Monogr. 60: 301. 1993;
Assis and Mello-Silva, Novon 20(4): 379. 2010.
Basionym: Nectandra virgultosa Nees, Syst. Laur.: 304.
1836; Meissner in DC., Prodr. 15(1): 167. 1864; ibid.
in Martius, Fl. Bras. 5(2): 278. 1866. TyPE: BRAZIL.
Minas Gerais, “in sylvulis udis depressis Capões
dictis,” s.d., C. F. P. von Martius s.n. (Holotype: M
0121088 [F neg. 19279]; Isotype, GZU 000249348
[fragment]).
57. Ocotea xanthocalyx (Nees & Mart.) Mez, Jahrb.
Königl. Bot. Gart. Berlin 5: 288. 1889; Rohwer, Mitt. Inst.
Allg. Bot. Hamburg 20: 161. 1986.
Basionym: Nectandra xanthocalyx Nees & Mart. in Nees,
Syst. Laur.: 312. 1836; Meissner in DC., Prodr. 15(1):
167. 1864; ibid. in Martius, Fl. Bras. 5(2): 277. 1866.
59. Persea rufotomentosa Nees & Mart. in Nees, Syst.
Laur.: 153. 1836; Meissner in DC., Prodr. 15(1): 45. 1864;
ibid. in Martius, Fl. Bras. 5(2): 155. 1866; Mez, Jahrb.
Königl. Bot. Gart. Berlin 5: 141. 1889; Kopp, Mem. New
york Bot. Gard. 14: 67. 1966; Moraes, Harvard Pap. Bot.
17(1): 205. 2012 ≡ Mutisiopersea rufotomentosa (Nees &
Mart.) Kosterm., Rheedea 3(2): 135. 1993. TyPE: BRAZIL.
Minas Gerais, Serro, “in sylvis Capões et in Tabuleiro locis
editis Montanis Serro Frio,” August 1818, l., C. F. P. von
Martius s.n. (Holotype: M 0147211 [F neg. 6582]; Isotypes,
GZU 000254417 [fragment], M 0147212, M 0147213).
60. Persea venosa Nees & Mart. in Nees, Linnaea 8: 50.
1833; ibid. in Syst. Laur.: 144. 1836; Meissner in DC.,
Prodr. 15(1): 48. 1864; ibid. in Martius, Fl. Bras. 5(2): 153,
t. 48, ig. 1. 1866; Mez, Jahrb. Königl. Bot. Gart. Berlin 5:
172. 1889; Kopp, Mem. New york Bot. Gard. 14: 30. 1966;
Moraes, Komarovia 6(1): 52. 2008; ibid. in Harvard Pap.
Bot. 17(1): 206. 2012. TyPE: BRAZIL. São Paulo, Iperó,
“ypanema,” s.d., l., F. Sellow B 1387 c 422 (Holotype:
2015
DE MORAES AND FALCADE, LAURACEAE COLLECTED By MARTIUS
B†; Lectotype, designated by Moraes, 2008: 52: G-DC
00135213; Isolectotypes, GZU 000254420, HAL 0103845,
K 000602076, K 000512722, Ny 00355864 [fragment ex
G-DC]).
Synonym: Persea venosa Nees & Mart. var. β tersa Nees &
Mart. in Nees, Linnaea 8: 50. 1833; Meissner in DC.,
Prodr. 15(1): 49. 1864; ibid. in Martius, Fl. Bras. 5(2):
153, t. 48, ig. 2. 1866; Kopp, Mem. New york Bot.
Gard. 14: 30. 1966; Moraes, Komarovia 6(1): 52. 2008;
ibid. in Harvard Pap. Bot. 17(1): 206. 2012. TyPE:
189
BRAZIL. Loco haud indicato, s.d., l., F. Sellow XXIV
(Holotype: B†; Lectotype designated by Moraes, 2008:
52: GZU 000273810 [annotated by Nees with “β”]).
Probable isolectotype: LISU [24000] ex Herbarium
Brasiliense Dr. Alexandre Rodrigues Ferreira nr. 693.
Specimen: BRAZIL. “in campestribus altis ad sylv.
margines / Provinciae S. Pauli et Min. Ger.,” s.d., fr., C.
F. P. von Martius s.n. (M 0147206 [“Persea hypoglauca
Mart.,” herbarium name]).
Further information in Moraes (2008).
remarKS
1. Martii Herbar. Florae Brasil. Nº 478: BRAZIL. Rio
3. Specimen at M 0111066 has been wrongly attributed to
de Janeiro, “in horto (Chacara) secundum viam ad S.
Martius. In fact, it is a Sellow collection belonging to
Clemente, prope Sebastianopolin, cultum,” March–
Ocotea acutifolia (Nees) Mez (see Moraes, 2008).
April 1833, fr., L. Riedel s.n. & B. Luschnath s.n. (BR
4. The name Laurus paniculigera Mart. ex Colla, Herb.
0000013332549, LE [4 sheets], Ny 459793), belongs to
Pedem. 5: 62. 1836, according to Moraes et al. (2014),
Cinnamomum camphora (L.) J. Presl in Berchtold and
remains unresolved since no material has been located
J. Presl, Přir. Rostlin 2: 36, 47–56, t. 8. 1825 [1823?].
at TO. However, as indicated by the former authors, in
2. Meissner (1866: 283) has reported a collection of Martius
Herbarium Sonder at MEL, there are two specimens that
from Rio de Janeiro for Goeppertia hirstuta (Schott)
agree with the protologue and belong to Metrodorea nigra
Nees, Syst. Laur.: 366. 1836; Meissner in DC., Prodr.
A. St.-Hil., Fl. Bras. Merid. (quarto ed.) 1: 81, t. 16. 1825.
15(1): 172. 1864 [= Endlicheria paniculata (Spreng.)
5. Specimen at MEL 2324410 has two handwritten labels,
J.F. Macbr., Publ. Field Mus. Nat. Hist., Bot. Ser. 13
one annotated with “Strychnodaphne Nees?”, the other
(2/3): 850. 1938]. However, it seems to be a mistake since
with “Laurus Martius”. Although its identiication is not
no other author has mentioned this material, neither it has
solved yet, it resembles the pattern of specimen at M
been located in Munich, nor in any other herbarium.
0111065, thus of Nectandra sarcocalyx Nees.
literature citeD
alvarez lopez, e. 1956. Comentario sobre “Laurus”, de Ruiz y
Pavón, con notas de Dombey acerca de algunas de sus especies.
Anales Inst. Bot. Cavanilles 13: 71–78.
aSSiS, l. c. S. anD r. mello-Silva. 2010. Taxonomic and
nomenclatural changes in the Ocotea indecora group
(Lauraceae). Novon 20: 377–380.
baitello, J. b. anD J. r. marcovino. 2003. Ocotea Aubl. Pages
179–208 in m. G. l. wanDerley, G. J. ShepherD, a. m.
Giulietti, t. S. melhem, eDS. Flora Fanerogâmica do Estado de
São Paulo, vol. 3. FAPESP/RiMa, São Paulo.
bentham, G. 1880. Ordo CXLIII. Laurineae. Pages 146–165
in G. bentham anD J. D. hooKer. Genera plantarum ad
exemplaria imprimis in herbariis Kewensibus servata deinita.
Voluminis Tertii Pars I.... L. Reeve & Co., Williams & Norgate,
Londini.
bernarDi, l. 1967. Emendationes laureae imprimis de Nectandra.
Candollea 22: 49–67.
biblioteca nacional [DiviSão De publicaçõeS e DivulGação;
Seção De expoSiçõeS]. 1968. Exposição comemorativa do
centenário da morte de Karl Friedrich Philipp von Martius 1868
– 1968. Homenagem ao sesquicentenário do Museu Nacional.
Iguassú, Rio de Janeiro.
bommer, J.-e. 1870. Notice sur le Jardin Botanique de Bruxelles.
Bull. Soc. Roy. Bot. Belgique 9: 418–444.
canDolle, a. l. p. p. De. 1856a. Notice sur la vie et les ouvrages de
M. de Martius… Imprimerie de Ramboz et Schuchardt, Genève.
––––––. 1856b. Sketch of the life and writings of M. de Martius,
Secretary to the Bavarian Academy of Science; by Alphonse de
Candolle. Hooker’s J. Bot. Kew Gard. Misc. 8: 362–369.
––––––. 1857. Sketch of the life and writings of M. de Martius,
Secretary to the Bavarian Academy of Science; by Alphonse de
Candolle. Hooker’s J. Bot. Kew Gard. Misc. 9: 6–10, 41–47.
caruS, c. G. 1869. Erinnerung an Carl Friedrich Philipp von
Martius. Nov. Actorum Acad. Caes. Leop.-Carol. German. Nat.
Cur. 35: 103–111.
chanDerbali, a. S. 2004. Endlicheria (Lauraceae). Fl. Neotrop.
Monogr. 91:1–141.
cluSiuS, c. 1601. Caroli ClvsI Atrebatis, Impp. Cæss. Avgg.
Maximiliani. II. Rudolphi. II. Aulæ quondam familiaris, Rariorvm
Plantarvm Historia. Quæ accesserint, proxima pagina docebit.
Ex oficina Plantiniana, apud Ioannem Moretum, Antverpiæ.
Dutra, J. S. 1942. Martius. Emiel, Rio de Janeiro.
eichler, a. w. 1869a. C. F. Ph. von Martius. Eine biographische
Skizze. Druck der Fr. Neubauer’schen Buchdruckerei (Krug’s
Wittwe), Regensburg.
––––––. 1869b. Das Herbarium Martii. Kgl. Hofbuchdruckerei von
Dr. C. Wolf & Sohn., München.
––––––. 1869c. Karl Friedrich Philipp von Martius. Gard. Chron.
1869: 164–165, 191–192, 221–222, 248–249.
––––––. 1869d. Karl Friedrich Philipp v. Martius. Allg. Zeitung
1869, Beilage Nr. 19, 21, 22: 277–279, 309–310, 325–326.
förther, h. 1994. Die Geschichte des Martius-Herbariums:
seine Brasilienkollektion und Empfehlungen zur Typuswahl.
Sendtnera 2: 5–24.
Garcia, r. 1922. Historia das Explorações Scientiicas. Pages
856–910 in inStituto hiStorico e GeoGraphico braSileiro,
Diccionario Historico, Geographico e Ethnographico do Brasil
(Commemorativo do Primeiro Centenario da Independencia).
Introducção geral. Primeiro volume. Imprensa Nacional, Rio de
Janeiro.
hinD, D. J. n. 2011. An annotated preliminary checklist of the
Compositae of Bolivia. Version 2. Kew Gardens, London. Available
at www.kew.org/science/tropamerica/boliviacompositae; www.
kew.org/science/tropamerica/boliviacompositae/checklist.pdf
190
HARVARD PAPERS IN BOTANy
hoehne, f. c. 1942. Notas bio-bibliográicas de naturalistas
botânicos que pretendemos homenagear com a denominação de
caminhos e picadas no Jardim Botânico e na Estação Biológica
do Alto da Serra. Pages 19–246 in f. c. hoehne, m. Kuhlmann
anD o. hanDro. O Jardim Botânico de São Paulo. Secretaria da
Agricultura, Indústria e Comércio, Departamento de Botânica
do Estado, São Paulo.
hooKer, w. J. 1841. I. Martius on the Botany of Brazil. J. Bot.
(Hooker) 4: 1–37.
howarD, r. a. 1981. Nomenclatural notes on the Lauraceae of the
Lesser Antilles. J. Arnold Arbor. 62: 45–61.
imKhanitSKaya, n. n. 1974. Lavrovye Kuby (Lauraceae Cubae).
Novosti Sist. Vyssh. Rast. 11: 192–209. (in Russian).
Kopp, l. e. 1966. A Taxonomic revision of the genus Persea in
the Western hemisphere. (Perseae–Lauraceae). Mem. New york
Bot. Gard. 14: 1–120.
KoStermanS, a. J. G. h. 1936. Revision of the Lauraceae I. Recueil
Trav. Bot. Néerl. 33: 719–757.
––––––. 1937. Revision of the Lauraceae II. The genera Endlicheria,
Cryptocarya (American species) and Licaria. Recueil Trav. Bot.
Néerl. 34: 500–609.
––––––. 1938a. Revision of the Lauraceae III. The genera Aiouea,
Systemonodaphne, Urbanodendron, Mezilaurus; additions and
corrections to Licaria and Cryptocarya. Recueil Trav. Bot.
Néerl. 35: 56–129.
––––––. 1938b. Revision of the Lauraceae V. A monograph of the
genera: Anaueria, Beilschmiedia (American species) and Aniba.
Recueil Trav. Bot. Néerl. 35: 834–931.
––––––. 1950. 81e Famille.–Lauracées. Pages 1–86 in h. humbert,
eD. Flore de Madagascar et des Comores. Typographie FirminDidot et Cie, Paris.
––––––. 1953. Notas sobre as Lauraceae-Lauroideae sul americanas.
Bol. Técn. Inst. Agron. N. 28: 49–75.
––––––. 1964. Bibliographia Lauracearum. Departemen Urusan
Research Nasional; P. T. Djulie “Archipel,” Bogor.
––––––. 1986. A monograph of the genus Cinnamomum Schaeff.
(Lauraceae), Part I. Ginkgoana 6: 1–171. Academia Scientiic
Book Inc., Tokyo.
KubitzKi, K. anD S. renner. 1982. Lauraceae I (Aniba and Aiouea).
Fl. Neotrop. Monogr. 31: 1–124.
Kurz, h. 2000. Revision der Gattung Licaria (Lauraceae). Mitt.
Inst. Allg. Bot. Hamburg 28–29: 89–221.
laSèGue, a. 1845. Musée Botanique de M. Benjamin Delessert.
Notices sur les collections de plantes et la bibliothèque qui
le composent; contenant em outre des documents sur les
principaux herbiers d’Europe Et l’exposé des Voyages entrepris
dans l’intérét de la Botanique. Librairie de Fortin, Masson et
Cie, Paris.
liSboa, K. m. 1995. Viagem pelo Brasil de Spix e Martius:
Quadros da natureza e esboço de uma civilização. Rev. Bras.
Hist. 15: 73–91.
––––––. 1997. A Nova Atlântida de Spix e Martius: natureza e
civilização na Viagem pelo Brasil (1817-1820). Editora Hucitec/
FAPESP, São Paulo.
lorea-hernánDez, f. G. 1996. A systematic revision of the
Neotropical species of Cinnamomum Schaeffer (Lauraceae).
Unpublished PhD. thesis, University of Missouri, Saint Louis.
lunan, J. 1814. Hortus Jamaicensis, or a botanical description,
(according to the Linnean system) and an account of the virtues,
&c. of its indigenous plants hitherto known, as also of the
most useful exotics. Compiled from the best authorities, and
alphabetically arranged. In Two Volumes. Printed at the Ofice
of the St. Jago de la Vega Gazette, Jamaica.
mäGDefrau, K. 1990. Martius, Carl Ritter von. Neue Deutsche
Biogr. 16: 310–312.
VOL. 20, NO. 2
martiuS, c. f. p. von. 1817–1820. Observationes botanicae,
Plantae in Itinere Brasiliensi annis 1817–1820 a Car. Frid.
Phil. Martio descriptae. 6 Vols. (Manuscript; copy handwritten
by Progel in M, copy in K).
––––––. 1829. Ueber einige brasilianische Arzneimittel. Buchner,
Repert. Pharm. 31: 337–383.
––––––. 1830. Dr. von Martius, über einige von ihm in der
brasilianischen Provinz von Rio Negro beobachtete ArzneiPlanzen. Buchner, Repert. Pharm. 35: 169–199.
––––––. 1837–[1841]. Herbarium Florae Brasiliensis. Plantae
brasilienses exsiccatae, quas denominatas, partim diagnosi
aut obsevationibus instructas Botanophilis offert Dr. C. Fr. Ph.
de Martius. Monachii. 1837: Flora 20(2, Beibl.): 1–128; 1838:
Flora 21(2, Beibl.): 49–96; 1839: Flora 22(1, Beibl.): 1–64;
1841: Flora 24(2, Beibl.): 1–112.
––––––. 1843a. Systema materiae medicae vegetabilis brasiliensis.
Apud F. Fleischer, Lipsiae; apud F. Beck in comm., Vindobonae.
––––––. 1843b. Plantae medicinales Florae Brasiliensis secundum
Ordines naturales dispositae. 10 Junii 1843. (Manuscript in
University of California at Los Angeles Library–Louise M.
Darling Biomedical Library).
––––––. 1850. Ueber das königliche Herbarium zu München. Gel.
Anz. 31: 717–747.
––––––. 1868. De Lauracearum qualitatibus et usu. Pages 315–
320 in c. f. p. von martiuS, eD. Flora Brasiliensis, Vol. 5(2).
Typographia Regia C. Wolf et il. in Ofic. Lithograph. S.
Minsinger tum B. Keller, Munich; apud Frid. Fleischer in
Comm., Leipzig.
––––––. anD J. h. roDriGueS. 1956. Como se deve escrever a
Historia do Brasil. Rev. Hist. América 42: 433–458.
mcneill, J., f. r. barrie, w. r. bucK, v. Demoulin, w. Greuter,
D. l. hawKSworth, p. S. herenDeen, S. Knapp, K. marholD,
J. praDo, w. f. pruD’homme van reine, G. f. Smith, J. h.
wierSema, anD n. J. turlanD. 2012. International Code of
Nomenclature for algae, fungi, and plants (Melbourne Code)
adopted by the Eighteenth International Botanical Congress
Melbourne, Australia, July 2011. Koeltz Scientiic Books,
Königstein. [Regnum Vegetabile 154].
meiSSner, c. f. 1864. Lauraceae. Pages 1–260 in a. l. p. p.
De canDolle, eD. Prodromus Systematis Naturalis Regni
Vegetabilis, Vol. 15(1). Sumptibus Victoris Masson et ilii, Paris.
––––––. 1866. Lauraceae et Hernandiaceae. Pages 137–302, 309–
310, tab. 45–107 in c. f. p. von martiuS, eD. Flora Brasiliensis,
Vol. 5(2). Typographia Regia C. Wolf et il. in Ofic. Lithograph.
S. Minsinger tum B. Keller, Munich; apud Frid. Fleischer in
Comm., Leipzig.
––––––. 1869. Denkschrift auf Carl Friedr. Phil. von Martius.
Akademische Buchdruckerei von F. Straub, in Commission bei
G. Franz, München.
––––––. 1870. Notice sur Charles-Frédéric-Philippe de Martius,
traduit de l’allemand par le docteur Postel. Bull. Soc. Linn.
Normandie, Ser. 2, 5: 308–341.
mez, c. 1888. Die amerikanischen Lauraceen des Döll’schen
Herbars. Mitt. Bot. Vereins Kreis Freiburg 47/48: 420–422.
––––––. 1889. Lauraceae americanae: monographice descripsit.
Jahrb. Königl. Bot. Gart. Berlin 5: 1–556.
––––––. 1892. Spicilegium laureanum. Arbeiten Königl. Bot. Gart.
Breslau 1: 71–166.
miquel, f. a. w. 1864. Cinnamomi generis revisio. Ann. Mus.
Bot. Lugduno-Batavi 1: 254–270.
moraeS, p. l. r. De. 2005. Lectotypiication of names of Brazilian
species of Cryptocarya (Lauraceae). Taxon 54: 789–795.
––––––. 2007. Taxonomy of Cryptocarya species of Brazil. Abc
Taxa 3: 1–191.
2015
DE MORAES AND FALCADE, LAURACEAE COLLECTED By MARTIUS
––––––. 2008. The lauraceous collections of Friedrich Sellow.
Komarovia 6: 1–67.
––––––. 2012a. The Lauraceae collected in Brazil by Ludwig
Riedel–I. Harvard Pap. Bot. 17: 185–216.
––––––. 2012b. The Lauraceae collected in Brazil by Ludwig
Riedel–II. Ocotea. Harvard Pap. Bot. 17: 245–273.
––––––. 2013. The collections of Lauraceae in the Herbarium of
Henri Van Heurck (AWH). Pl. Ecol. Evol. 146: 360–383.
––––––, S. De SmeDt, h.-J. eSSer, c. GalaGher, anD l. GuGlielmone.
2013. On some Brazilian plants distributed by Martius in 1827
and published by Colla in 1833. Harvard Pap. Bot. 18: 23–36.
––––––, S. De SmeDt, anD l. GuGlielmone. 2014. On some Brazilian
plants distributed by Martius in 1827 and published by Colla in
Herbarium Pedemontanum–V. Harvard Pap. Bot. 19: 143–155.
neeS von eSenbecK, c. G. D. 1833. Revisio laurinarum ab Sellowio
in Brasilia collectarum et iam in Herbario Regio Berolinensi
asservatarum. Linnaea 8: 36–51.
––––––. 1836. Systema Laurinarum. Sumtibus Veitii et sociorum,
Berolini.
––––––. 1848. Laurinae Vent. / Laurineae Kunth in Beiträge zu
einer Flora der Aequinoctial-Gegenden der neuen Welt, von Joh.
Friedr. Klotzsch. Linnaea 21: 487–526.
papavero, n. 1971. Essays on the history of Neotropical
Dipterology, with special reference to collectors (1750–1905),
Vol. 1. Museu de Zoologia, Universidade de São Paulo, São
Paulo.
prantl, c. 1872. Geschichte der Ludwig-Maximilians-Universität
in Ingolstadt, Landshut, München, zur Festfeier ihres
vierhundertjährigen Bestehens im Auftrage des Akademischen
Senates. Zweiter Band. Christian Kaiser, München.
rambo, b. 1952. Martius. Inst. Hans Staden, São Paulo.
rau, c. 1871. Memoir of C. F. P. von Martius. Annu. Rep. Board
Regents Smithson. Inst. [for the year 1869] (1871): 169–178.
rohwer, J. G. 1986. Prodromus einer Monographie der Gattung
Ocotea Aubl. (Lauraceae) sensu lato. Mitt. Inst. Allg. Bot.
Hamburg 20: 1–278.
––––––. 1988. The genera Dicypellium, Phyllostemonodaphne,
Systemonodaphne and Urbanodendron (Lauraceae). Bot. Jahrb.
Syst. 110: 157–171.
––––––. 1993. Lauraceae: Nectandra. Fl. Neotrop. Monogr. 60:
1–333.
roSS, h. 1917. Zum Andenken der Forschungsreise von Spix und
Martius in Brasilien 1817–1820. Ber. Deutsch. Bot. Ges. 35:
119–128.
ruiz, h. anD J. pavón. 1956. Flora Peruviana, et Chilensis, sive
descriptiones et icones plantarum Peruvianarum, et Chilensium,
secundum Systema Linnaeanum digestae, cum characteribus
plurium generum evulgatorum reformatis. Tomus IV (Pars II).
Anales Inst. Bot. Cavanilles 13: 5–70.
–––––– anD ––––––. 1989. Laurographia Florae Peruviana et
Chilensis. Introducción, transcripción y notas by a. González
bueno anD m. c. navarro aranDa. Ayuntamento de la Villa de
Belorado, Burgos.
Schoepf, J. D. 1787. D. Io. Davidis Schoepf… Materia medica
americana potissimvum Regni Vegetabilis. Svmtibvs Io. Iac.
Palmii, Erlangae.
Schramm, h. 1869a. C. F. Ph. v. Martius. Sein Lebens-und
Characterbild insbesondere seine Reiseerlebnisse in Brasilien.
Vol. 1. Ludwig Denicke, Leipzig.
––––––. 1869b. C. F. Ph. v. Martius. Sein Lebens-und Characterbild
insbesondere seine Reiseerlebnisse in Brasilien. Vol. 2. Ludwig
Denicke, Leipzig.
Seemann, b. 1869. Carl Friedrich Phillipp von Martius. J. Bot. 7:
17–21.
191
––––––. 1870. Botanical news. J. Bot. 8: 136, 165.
ShepherD, G. J. 2002. A brief history of the Flora Brasiliensis.
Available at http://lorabrasiliensis.cria.org.br/info?history
(accessed in May 2015).
Sloane, h. 1725. A voyage to the Islands Madera, Barbadoes,
Nieves, St Christophers, and Jamaica; with the Natural History
of the herbs and trees, four-footed beasts, ishes, birds, insects,
reptiles, &c. of the last of those Islands. To which is preix’d,
and Introduction, wherein is an Account of the inhabitants, air,
waters, diseases, trade, &c. of that Place; with some Relations
concerning the Neighbouring Continent, and Islands of America.
Illustrated with the igures of the Things described, which have
not been heretofore engraved. In large Copper-Plates as big
as the Life. By Sir Hans Sloane, Bart. In Two Volumes. Vol. II.
Printed for the Author, London.
Sommer, f. 1953. A vida do Botânico Martius: Pai das Palmeiras.
Melhoramentos, São Paulo.
Spix, J. b. von anD c. f. p. von martiuS. 1823. Reise in Brasilien
auf Befehl Sr. Majestät Maximilian Joseph I. Königs von
Baiern in den Jahren 1817 bis 1820… Erster Theil. Mit einer
geographischen Charte und fünfzehn Abbildungen. Gedruckt bei
M. Lindauer, München.
–––––– anD ––––––. 1828. Reise in Brasilien auf Befehl Sr. Majestät
Maximilian Joseph I. Königs von Baiern in den Jahren 1817 bis
1820… Zweiter Theil, bearbeitet und herausgegeben von Dr.
C. F. P. von Martius. Mit vier Charten und vier und zwanzig
Abbildungen. Gedruckt bei I. J. Lentner, München.
–––––– anD ––––––. 1831a. Reise in Brasilien auf Befehl Sr.
Majestät Maximilian Joseph I. Königs von Baiern in den
Jahren 1817 bis 1820… Dritter und letzter Theil, bearbeitet und
herausgegeben von Dr. C. F. P. von Martius. Mit einer Karte
vom Amazonenstrome. Bei dem Verfasser, München; in Comm.
bei Friedr. Fleischer, Leipzig.
–––––– anD ––––––. 1831b. Atlas zur Reise in Brasilien. Gedruckt
bei M. Lindauer, München.
–––––– anD ––––––. 1968. Viagem pelo Brasil 1817–1820: Excertos
e ilustrações. Tradução de l. f. lahmeier, revisado por b. f. r.
Galvão e b. De maGalhãeS – segundo versão de e. winKler.
Introdução de h. balDuS. Edições Melhoramentos, São Paulo.
SprinG, a. 1870. Prologue a la mémoire de Charles-Fréd.-Phil. von
Martius, 1794-1868. Ann. Hort. Belge Étrangère 20: V–XXIV.
––––––. 1871. Notice sur Charles-Fréd.-Phil. von Martius. Annuaire
Acad. Roy. Sci. Belgique 37: 257–294.
Stafleu, f. a. anD cowan, r. S. 1981. Taxonomic literature. 2nd
ed. Vol. 3: Lh–O. Regnum Vegetabile 105. Dr. W. Junk b.v.,
Bohn, Scheltema & Holkema, Utrecht, The Hague.
Stapf, o. 1912. Order CXVI. Laurineæ. Pages 493–502 in w. t.
thiSelton-Dyer, eD. Flora Capensis, Vol. 5(1). Lovell Reeve &
Co., Ltd., London.
thierS, b. [continuously updated]. Index Herbariorum: A global
directory of public herbaria and associated staff. New york
Botanical Garden’s Virtual Herbarium. Available at http://
sweetgum.nybg.org/ih/ (accessed in May 2015).
tiefenbacher, l. 1983. Die Brasilienexpedition von J. B. v. Spix
und C. F. Ph. v. Martius in den Jahren 1817 bis 1820. Spixiana,
Supplement 9: 35–42.
urban, i. 1906. Vitae itineraque collectorum botanicorum, Notae
collaboratorum biographicae, Flora Brasiliensis ratio edendi
chronologica, Systema, Index Familiarum. Pages 1–268 in c.
f. p. von martiuS, a. w. eichler, anD i. urban, eDS. Flora
Brasiliensis, Vol. 1 (1). Apud R. Oldenbourg in comm., München
et Leipzig.
––––––. 1908. Martii Flora Brasiliensis. Verh. Bot. Vereins Prov.
Brandenburg 49: 1–6.
192
HARVARD PAPERS IN BOTANy
werff, h. 2002a. A synopsis of Persea (Lauraceae) in
Central America. Novon 12: 575–586.
––––––. 2002b. A synopsis of Ocotea (Lauraceae) in Central
America and Southern Mexico. Ann. Missouri Bot. Gard. 89:
429–451.
vanzolini, p. e. 1996. A contribuição zoológica dos primeiros
naturalistas viajantes no Brasil. Revista USP 30: 190–238.
voGel, a. 1869. Carl Friedrich von Martius. Nekrolog. Neues
Repert. Pharm. 18: 121–128.
wallich, n. 1851. Das Königliche Herbarium zu München
geschildert. (Sketch of the Royal Herbarium at Munich.) By
Dr. C. F. Ph. von Martius. Translated from the German in the
Gelehrten Anzeigen, Bd. XXXI. No. 89–93 [separately printed
at Munich 1850]; by N. Wallich,… Hooker’s J. Bot. Kew Gard.
Misc. 3: 65–74, 102–111.
van Der
VOL. 20, NO. 2
weber, J. z. 1981. A taxonomic revision of Cassytha (Lauraceae)
in Australia. J. Adelaide Bot. Gard. 3: 187–262.
wittrocK, v. b. 1903. Catalogus illustratus iconothecæ botanicæ
horti Bergiani Stockholmiensis anno 1903; notulis biographicis
adjectis. Acta Horti Berg. 3: 136.
wunSchmann, e. 1884. Karl Friedrich Philipp von Martius. Allg.
Deutsche Biogr. 20: 517–527.
wuScheK, e. 1989. Die Beiträge des Botanikers Carl Friedrich
Philipp von Martius (1794-1868) zur Pharmakognosie und zur
Erforschung tropischer Nutzplanzen, mit einer Bearbeitung
seines Systema Materiae Medicae Vegetabilis Brasiliensis von
1843. Dissertation der Fakultät für Chemie und Pharmazie der
Ludwig-Maximilians-Universität München.
wuu-KuanG, S. 2011. Taxonomic revision of Cinnamomum
(Lauraceae) in Borneo. Blumea 56: 241–264.
inDex to Scientific nameS
Current accepted names in boldface italics. Synonyms in italics. Unpublished (herbarium) names in regular typeset.
Numbers in parentheses indicate entries of the respective species in the text.
Acrodiclidium brasiliense Nees (12)
Acrodiclidium cannella (Meisn.) Mez (13)
Acrodiclidium cayennense (Meisn.) Mez (13)
Acrodiclidium guianense Nees var. caudatum Meisn. (14)
Acrodiclidium meissneri Mez (14)
Acrodiclidium puchury-major (Mart.) Mez (15)
Aiouea laevis (Mart.) Kosterm. (1)
Aiouea brasiliensis Meisn. (1)
Aiouea brasiliensis Meisn. var. attenuata Meisn. (1)
Aiouea riedelii Mez (1)
Aiouea tenuiflora Nees ex Meisn. (41)
Aniba canelilla (Kunth) Mez (2)
Aniba citrifolia (Nees) Mez (3)
Aniba desertorum (Nees) Mez (4)
Aniba permollis (Nees) Mez (5)
Aniba puchury-minor (Mart.) Mez (6)
Aperiphracta martiana Nees (50)
Aperiphracta velutina Nees (55)
Aydendron brasiliense (Nees) Meisn. (12)
Aydendron cannella Meisn. (13)
Aydendron cayennense Meisn. (13)
Aydendron citrifolium Nees (3)
Aydendron cujumary (Mart.) Nees (2, 36)
Aydendron desertorum Nees (4)
Aydendron irmulum Nees (35)
Aydendron permolle Nees (5)
Aydendron suaveolens Nees (2)
Aydendron suaveolens Nees var. β Nees (44)
Camphoromoea laxa Nees (43)
Camphoromoea subtriplinervia Nees (31)
Camphoromoea tenuilora (Nees) Meisn. (33)
Camphoromoea venulosa Nees (52)
Cassytha americana Nees var. brasiliensis (Nees) Meisn.
(7)
Cassytha americana Nees var. puberula (Nees) Meisn. (7)
Cassytha brasiliensis Mart. ex Nees (7)
Cassytha filiformis L. (7)
Cinnamomum camphora (L.) J. Presl (Remarks)
Cinnamomum erythropus (Nees & Mart.) Kosterm. (8)
Cinnamomum verum J. Presl (9)
Cinnamomum zeylanicum Blume (9)
Cinnamomum zeylanicum Blume var. microphyllum
Nees (9)
Cinnamomum zeylanicum Breyn. (9)
Chlorocardium rodiei (R.H. Schomb.) Rohwer, H.G.
Richt. & van der Werff (10)
Cryptocarya aü-üva Mart. ex Nees (5)
Cryptocarya canelilla Kunth (2)
Cryptocarya guianensis Meisn. (10)
Cryptocarya laevis Nees ex Mart. (1)
Cryptocarya mandioccana Meisn. (10)
Cryptocarya monticola Mart. (48)
Cryptocarya monticola Mart. ex Nees (48)
Cryptocarya moschata Mart. (10)
Cryptocarya moschata Nees & Mart. (10)
Cryptocarya pretiosa Mart. (2)
Cryptocarya pyriformis levis Nees (1)
Cryptocarya riedeliana P.L.R. Moraes (10)
Dicipellium caryophyllatum Nees (11)
Dicypellium caryophyllaceum (Mart.) Nees & Mart.
(2, 11)
Endlicheria paniculata (Spreng.) J.F. Macbr. (Remarks)
Goeppertia hirstuta (Schott) Nees (Remarks)
Gymnobalanus minarum Nees & Mart. (45)
Laurus adamantina Mart. (54)
Laurus Aüiuva Mart. (5)
Laurus amazonum Mart. (17)
Laurus amoena Mart. (26)
Laurus aurantiodora Ruiz & Pav. (44)
Laurus canelilla Willd. (2)
Laurus cassioides Mart. (9)
Laurus cassioides Mart. ex Miq. (9)
Laurus ceanothifolia Mart. (32)
Laurus cinnamomum L. (9)
Laurus coruscans Bonpl. ex Willd. (37)
Laurus crassifolia Poir. (13)
Laurus cuspidata Mart. (19)
Laurus dura Mart. (42)
Laurus egensis Mart. (19)
Laurus hihua Ruiz & Pav. (20)
2015
DE MORAES AND FALCADE, LAURACEAE COLLECTED By MARTIUS
Laurus japurensis Mart. (21)
Laurus leptobotra Ruiz & Pav. (33)
Laurus longifolia Mart. (37)
Laurus membranacea Sw. (23)
Laurus nitida Ruiz & Pav. (33)
Laurus ocotea Rich. (39)
Laurus opifera Mart. ex Meisn. (44)
Laurus paniculigera Mart. ex Colla (Remarks)
Laurus pauciflora Mart. (46)
Laurus paulina Mart. (47)
Laurus persea L. (58)
Laurus pichurim Mart. (15)
Laurus pretiosa Mart. (2)
Laurus pubescens Dombey (29)
Laurus purpurea Ruiz & Pav. (28)
Laurus quixos Lam. (2)
Laurus reticulata Ruiz & Pav. (29)
Laurus riparia Mart. (12)
Laurus sanguinea (Rol.) Sw. (11)
Laurus saporosa Mart. (26)
[Laurus] sarcocalyx Mart. (24)
Laurus villosa Willd. (29)
Leptodaphne subalpina Nees (33)
Leptodaphne tenuilora Nees (33)
Licaria brasiliensis (Nees) Kosterm. (12)
Licaria cannella (Meisn.) Kosterm. (13)
Licaria cayennensis (Meisn.) Kosterm. (13)
Licaria crassifolia (Poir.) P.L.R. Moraes (13)
Licaria cymbarum (Kunth) Pittier (37)
Licaria polyphylla (Nees) Kosterm. (14)
Licaria puchury-major (Mart.) Kosterm. (15)
Mespilodaphne aurantiodora (Ruiz & Pav.) Meisn. (44)
Mespilodaphne ceanothifolia Nees (32)
Mespilodaphne complicata Meisn. (35)
Mespilodaphne glauca (Nees) Meisn. var. virescens Meisn.
(47)
Mespilodaphne indecora (Schott) Meisn. (41)
Mespilodaphne indecora (Schott) Meisn. var. stricta
Meisn. (41)
Mespilodaphne nitida (Ruiz & Pav.) Meisn. (33)
Mespilodaphne nutans (Nees) Meisn. (47)
Mespilodaphne opifera (Mart.) Meisn. (2, 44)
Mespilodaphne phillyreoides (Nees) Meisn. (48)
Mespilodaphne phillyreoides Nees (Meisn.) var. oblonga
Meisn. (48)
Mespilodaphne pretiosa Nees & Mart. (2)
Mespilodaphne pretiosa Nees & Mart. var. angustifolia
Nees (2)
Mespilodaphne pulchella (Nees & Mart.) Meisn. (49)
Mespilodaphne pulchella (Nees & Mart.) Meisn. var.
elliptica Meisn. (49)
Mespilodaphne pulchella (Nees & Mart.) Meisn. var.
ferruginea Meisn. (49)
Mespilodaphne tristis Meisn. (53)
Metrodorea nigra A. St.-Hil. (Remarks)
Mezilaurus navalium (Allemão) Taub. ex Mez (10)
Misanteca brasiliensis (Nees) Lundell (12)
Misanteca cannella (Meisn.) Lundell (13)
193
Misanteca cayennensis (Meisn.) Lundell (13)
Misanteca cymbarum (Kunth) Lundell (37)
Misanteca polyphylla (Nees) Lundell (14)
Misanteca puchury-major (Mart.) Lundell (15)
Mutisiopersea rufotomentosa (Nees & Mart.) Kosterm.
(59)
Nectandra acuminata (Nees & Mart.) J.F. Macbr. (16)
Nectandra acutifolia (Ruiz & Pav.) Mez (16)
Nectandra acutissima Nees ex Meisn. (25)
Nectandra amara Meisn. var. chartacea Meisn. (27)
Nectandra amara Meisn. var. subevenia Meisn. (27)
Nectandra amazonum Nees (17)
Nectandra berchemiifolia Meisn. (16)
Nectandra berchemiaefolia Meisn. (16)
Nectandra canescens Nees & Mart. (18, 27)
Nectandra cuspidata Nees & Mart. (19)
Nectandra cuspidata Nees & Mart. var. macrocarpa Nees
(23)
Nectandra cymbarum (Kunth) Nees (2, 37)
Nectandra hihua (Ruiz & Pav.) Rohwer (20, 22)
Nectandra japurensis Nees & Mart. (21)
Nectandra latifolia (Kunth) Mez (28)
Nectandra leucantha Nees & Mart. (10, 22)
Nectandra leucantha Nees var. peruviana Meisn. (20)
Nectandra lucida Nees (20)
Nectandra megapotamica (Spreng.) Mez (24, 26)
Nectandra membranacea (Sw.) Griseb. (23)
Nectandra membranacea (Sw.) Griseb. ssp. cuspidata
(Nees & Mart.) Rohwer (19)
Nectandra mollis (Kunth) Nees (29)
Nectandra mollis (Kunth) Nees ssp. oppositifolia (Nees &
Mart.) Rohwer (25)
Nectandra neesiana Miq. (46)
Nectandra nitidula Nees & Mart. (24)
Nectandra nitidula Nees & Mart. var. maior Nees (13, 24)
Nectandra nitidula Nees & Mart. var. minor Nees (24)
Nectandra nitidula Nees & Mart. var. nitidula (24)
Nectandra oppositifolia Nees & Mart. (25)
Nectandra oppositifolia Nees & Mart. fo. intermedia Mez
(25)
Nectandra oppositifolia Nees & Mart. var. angustifolia
Nees & Mart. (25)
Nectandra oppositifolia Nees & Mart. var. oppositifolia
(25)
Nectandra polita Nees & Mart. (28)
Nectandra polyphylla Nees (14)
Nectandra psammophila Nees & Mart. (26)
Nectandra puberula (Schott) Nees (27)
Nectandra puchury-major (Mart.) Nees & Mart. (2, 15)
Nectandra puchury-minor (Mart.) Nees (2, 6)
Nectandra purpurea (Ruiz & Pav.) Mez (28)
Nectandra reticulata (Ruiz & Pav.) Mez (29)
Nectandra rigida (Kunth) Nees var. canescens Mez (18)
Nectandra rodgiaei Schomb. (10)
Nectandra rodioei R.H. Schomb. (10)
Nectandra sarcocalyx Nees (24, Remarks)
Nectandra schomburgkii Meisn. (20)
Nectandra schottii Meisn. (25)
194
HARVARD PAPERS IN BOTANy
Nectandra villosa Nees & Mart. var. venosa Nees (29)
Nectandra virgultosa Nees (56)
Nectandra xanthocalyx Nees & Mart. (57)
Ocotea acutifolia (Nees) Mez (Remarks)
Ocotea amara Mart. (37)
Ocotea aurantiodora (Ruiz & Pav.) Mez (44)
Ocotea bracteolata (Nees) Mez (30)
Ocotea camphoromoea Rohwer (31)
Ocotea caryophyllacea (Mart.) Kostel. (11)
Ocotea ceanothifolia (Nees) Mez (32)
Ocotea cernua (Nees) Mez (33)
Ocotea citrosmoides (Nees) Mez (34)
Ocotea complicata (Meisn.) Mez (35)
Ocotea cujumary Mart. (36)
Ocotea cymbarum Kunth (37)
Ocotea daphnoides Mart. (42)
Ocotea declinata (Meisn.) Mez (38)
Ocotea delexa Rohwer (38)
Ocotea desertorum Mart. (4)
Ocotea fallax Miq. ex Mez (46)
Ocotea fasciculata (Nees) Mez (35)
Ocotea firmula Mart. (35)
Ocotea glauca (Nees) Mez var. virescens (Meisn.)
Mez (47)
Ocotea guianensis Aubl. (39)
Ocotea hypoglauca (Nees & Mart.) Mez (40)
Ocotea indecora (Schott) Mez (41)
Ocotea lanceolata (Nees) Nees (42)
Ocotea lancifolia (Schott) Mez (42, 54)
Ocotea latifolia Kunth (28)
Ocotea laxa (Nees) Mez (43, 52)
Ocotea leptobotra (Ruiz & Pav.) Mez (33)
Ocotea longifolia Kunth (44)
Ocotea lucida Mart. (20)
Ocotea membranacea (Sw.) R.A. Howard (23)
Ocotea minarum (Nees & Mart.) Mez (45)
Ocotea mollis Kunth (29)
Ocotea neesiana (Miq.) Kosterm. (46)
Ocotea nutans (Nees) Mez (47)
Ocotea obtusifolia Mart. (4)
Ocotea odorifera (Vell.) Rohwer (2, 42)
Ocotea opifera Mart. (44)
Ocotea petalanthera (Meisn.) Mez (30)
Ocotea phillyreoides (Nees) Mez (48)
Ocotea pretiosa (Nees & Mart.) Benth. (2)
Ocotea pretiosa Mez (2, 42)
Ocotea pretiosa Mez var. indecora (Schott)
Vattimo-Gil (41)
Ocotea puchury-major Mart. (15)
Ocotea puchury-minor Mart. (6)
Ocotea pulchella (Nees & Mart.) Mez (49)
Ocotea quixos (Lam.) Kosterm. (2)
Ocotea rufotomentosa Mart. (50)
Ocotea spixiana (Nees) Mez (50)
Ocotea suaveolens (Meisn.) Benth. ex Hieron. (2)
Ocotea subacris Mart. (42)
Ocotea subalpina (Nees) Mez (33)
Ocotea subdecumbens M.A. Lisboa & J. Badini (47)
VOL. 20, NO. 2
Ocotea subtriplinervia (Nees) Mez (31)
Ocotea tabacifolia (Meisn.) Rohwer (51)
Ocotea teleiandra (Meisn.) Mez (52)
Ocotea tenuilora (Nees) Mez (33, 46)
Ocotea tristis (Nees & Mart.) Mez (48, 53)
Ocotea tubigera Mart. (45)
Ocotea umbrosa Kunth (51)
Ocotea umbrosa (Nees) Mart. ex Mez (51)
Ocotea variabilis Mart. ex Nees (54)
Ocotea variabilis (Nees) Mez (54)
Ocotea velutina Mart. (55)
Ocotea velutina (Nees) Rohwer (55)
Ocotea venulosa (Nees) Baitello (52)
Ocotea virgultosa (Nees) Mez (56)
Ocotea xanthocalyx (Nees & Mart.) Mez (57)
Oreodaphne bracteolata Meisn. (30)
Oreodaphne cernua Nees (33)
Oreodaphne citrosmoides Nees (34)
Oreodaphne declinata Meisn. (38)
Oreodaphne dispersa Nees & Mart. (46)
Oreodaphne fallax Miq. (46)
Oreodaphne glauca Nees (52)
Oreodaphne guianensis (Aubl.) Nees (39)
Oreodaphne guianensis (Aubl.) Nees var. argentea
Meisn. (39)
Oreodaphne guianensis (Aubl.) Nees var. guianensis (39)
Oreodaphne indecora (Schott ) Nees (41)
Oreodaphne japurensis Meisn. (46)
Oreodaphne kunthiana Meisn. (47)
Oreodaphne lancifolia (Schott) Nees (42)
Oreodaphne leptobotra (Ruiz & Pav.) Nees (33)
Oreodaphne longifolia (Kunth) Nees (44)
Oreodaphne martiana Nees (42)
Oreodaphne martiana Nees var. thymelaeoides Meisn. (42)
Oreodaphne minarum Steud. (51)
Oreodaphne nitidula Nees & Mart. (42)
Oreodaphne nitidula Nees & Mart. var. lanceolata
Meisn. (42)
Oreodaphne nutans Nees (47)
Oreodaphne nutans Nees var. sylvestris Nees (47)
Oreodaphne opifera (Mart.) Nees (44)
Oreodaphne phillyreoides Nees (48)
Oreodaphne pulchella Nees & Mart. (49)
Oreodaphne spixiana Nees (50)
Oreodaphne suaveolens Meisn. (2)
Oreodaphne subalpina (Nees) Meisn. (33)
Oreodaphne teleiandra Meisn. (52)
Oreodaphne thymelaeoides Nees & Mart. (42)
Oreodaphne tristis Nees & Mart. (53)
Oreodaphne tristis Nees & Mart. var. oblongifolia
Nees & Mart. (53)
Oreodaphne tristis Nees & Mart. var. tristis (53)
Oreodaphne umbrosa Nees (51)
Oreodaphne umbrosa (Kunth) Nees (51)
Oreodaphne variabilis Nees (54)
Oreodaphne velutina Nees (55)
Oreodaphne venulosa (Nees) Meisn. (52)
Persea acuminata Nees & Mart. (16)
2015
DE MORAES AND FALCADE, LAURACEAE COLLECTED By MARTIUS
Persea americana Mill. (58)
Persea canescens Mart. (18)
Persea caryophyllacea Mart. (11)
Persea caryophyllata Mart. (11)
Persea ceanothifolia Mart. ex Nees (32)
Persea erythropus Mart. (8)
Persea erythropus Nees & Mart. (8)
Persea erythropus Nees & Mart. var. angustata Meisn. (8)
Persea erythropus Nees & Mart. var. mixtinervia
Meisn. (8)
Persea gratissima Gaertn. (58)
Persea hypoglauca Mart. (60)
Persea hypoglauca Nees & Mart. (40)
Persea indecora Schott (41)
Persea lancifolia Schott (42)
Persea laxa Mart. (43)
Persea laxa Mart. ex Nees (43)
Persea leucantha Mart. (22)
Persea longifolia (Kunth) Spreng. (44)
Persea membranacea (Sw.) Spreng. (23)
Persea mollis (Kunth) Spreng. (29)
195
Persea paniculigera Mart. (24)
Persea puberula Schott (27)
Persea rufotomentosa Nees & Mart. (59)
Persea sarcocalyx Mart. (24)
Persea scrobiculata Meisn. (40)
Persea subtriplinervia Mart. ex Nees (31)
Persea tabacifolia Meisn. (51)
Persea tenuiflora Mart. (33)
Persea tenuilora Mart. ex Nees (33)
Persea venosa Nees & Mart. (40, 60)
Persea venosa Nees & Mart. var. β tersa
Nees & Mart. (60)
Petalanthera bracteolata Nees (30)
Petalanthera paniculigera Nees (24)
Phoebe erythropus (Nees & Mart.) Mez (8)
Phoebe membranacea (Sw.) Nees (23)
Pomatium canescens Mart. (18)
Pomatium oppositifolium Nees (25)
Silvia navalium Allemão (10)
Strychnodaphne lanceolata Nees (42)
Teleiandra glauca Nees & Mart. (52)
inDex to collectionS
Numbers in bold in parentheses refer to the species number in treatment.
Ackermann V no. 121 = Martii Herbar. Florae Brasil. Nº
169 (29)
Ackermann s.n. = Martii Herbar. Florae Brasil. Nº 1238 (8)
Anonymous s.n. [JE 00000523] (1)
Anonymous s.n. [GZU 000265498] (8)
Anonymous s.n. [GZU 000249358] (24)
Anonymous s.n. [GZU 000259200] (24)
Anonymous s.n. [K 000642237] (26)
Anonymous s.n. [K 000642238] (26)
Anonymous s.n. [MEL 2324399] (42)
Anonymous s.n. [B 10 0185383] (44)
Aublet s.n. [BM 000947302] (39)
Aublet s.n. [P 00756908] (39)
Blanchet 301 (47)
Blanchet 3192 (38)
Blanchet 3192A (38)
Blanchet 3250 (47)
Blanchet 3250B (47)
Claussen 47 (27)
Claussen 78 (27)
Claussen 106 = Martii Herbar. Florae Brasil. Nº 1239 (24)
Claussen 107 = Martii Herbar. Florae Brasil. Nº 1239 (24)
Claussen 108 (8)
Claussen 2078 (27)
Claussen s.n. = Martii Herbar. Florae Brasil. Nº 1239 (24)
Dombey s.n. [B 10 0185149, G-DC, GZU 000254296, MA
811775, P 00745320, P 00745321, P-JU 00662847] (29)
Dombey s.n. [P 00756826, P 00756827] (33)
Guillemin 667 (27)
Herb. Clifford: 154, Laurus 6, sheet B [BM 000558701] (9)
Herb. Sloane Vol. VII: 77 [BM-SL 000594115] (58)
Hornemann s.n. [C, GZU 000260297] (33)
Hostmann & Kappler 1433 (46)
Humboldt & Bonpland 904 (37)
Humboldt & Bonpland 1018 (2)
Humboldt & Bonpland 1729 (44)
Humboldt & Bonpland s.n. [P 00128755] (28)
Humboldt & Bonpland s.n. [B 10 0185155, B-W 07796 -01
0 & B-W 07796 -00 0, F (0061634F), GZU 000254324,
Ny 00355629, Ny 00355630, P 00128743, P 00128746,
U 0002912, US 00028321] (29)
Kohaut s.n. = Sieber Flora martin. No. 106 (33)
Lisboa & Badini s.n. [OUPR (26600), OUPR (18212)] (47)
Lisboa & Badini s.n. [HRCB, OUPR (26600)] (47)
Luschnath 32 (38)
Luschnath s.n. = Martii Herbar. Florae Brasil. Nº 420 (28)
Luschnath s.n. = Martii Herbar. Florae Brasil. Nº 477 (9)
Luschnath s.n. = Martii Herbar. Florae Brasil. Nº 478
(Remarks # 1)
Luschnath s.n. = Martii Herbar. Florae Brasil. Nº 1307 (27)
Luschnath s.n. = Martii Herbar. Florae Brasil. Nº 1311 (38)
Luschnath s.n. [BR 0000008808042] (28)
Martii Herbar. Florae Brasil. Nº 117 (49)
Martii Herbar. Florae Brasil. Nº 169 (29)
Martii Herbar. Florae Brasil. Nº 237 (1)
Martii Herbar. Florae Brasil. Nº 420 (28)
Martii Herbar. Florae Brasil. Nº 477 (9)
Martii Herbar. Florae Brasil. Nº 478 (Remarks # 1)
Martii Herbar. Florae Brasil. Nº 1238 (8)
Martii Herbar. Florae Brasil. Nº 1239 (24)
Martii Herbar. Florae Brasil. Nº 1307 (27)
Martii Herbar. Florae Brasil. Nº 1311 (38)
Martius Obs. 261 (33)
Martius Obs. 411 (26)
Martius Obs. 512 (49)
Martius Obs. 546 (24)
Martius Obs. 595 (49)
Martius Obs. 678 (43)
196
HARVARD PAPERS IN BOTANy
Martius Obs. 883 (8)
Martius Obs. 903 (40)
Martius Obs. 1095 b (10)
Martius Obs. 1112 (42)
Martius Obs. 1183 (55)
Martius Obs. 1188 (51)
Martius Obs. 1223 (34)
Martius Obs. 1704 (22)
Martius Obs. 1770 (57)
Martius Obs. 2617 (11)
Martius Obs. 2750 (18)
Martius Obs. 2796 (44)
Martius Obs. 2908 (5)
Martius Obs. 3174 (32)
Martius “n. 79” (42)
Martius s.n. [M 0121069, M 01121070, M 0121071, M
0121072, MEL 2318499] (2)
Martius s.n. [G, MEL 2318499] (2)
Martius s.n. [BR 0000008685865] (2)
Martius s.n. [M 0147109] (3)
Martius s.n. [M 0147104, M 0147105, M 0147106, M
0147107, M 0147108] (4)
Martius s.n. [M 0147107] (4)
Martius s.n. [B†, M 0113213] (6)
Martius s.n. [GZU 000254329, M 0112564] (6)
Martius s.n. [M 0112563] (6)
Martius s.n. [M] (7)
Martius s.n. [M] (7)
Martius s.n. [M 0147195] (8)
Martius s.n. [BR, M 0147232] (9)
Martius s.n. [L 0036030, M 0147236] (9)
Martius s.n. [B 10 0185010, B 10 0185011, G, GZU
000254389, M 0147268] (11)
Martius s.n. [M 0147266] (11)
Martius s.n. [M 0147267] (11)
Martius s.n. [L 0308064, M 0147427, M 147428, M
0147429, MEL 2324398, S (R-7276)] (12)
Martius s.n. [GZU 000256425, M 0147426] (13)
Martius s.n. [GZU 000254280, M 0147422] (14).
Martius s.n. [B†, BR 0000008812841, G 00368872, GZU
000254328, L 0036458, M 0147419, M 0147420, M
0147421] (15)
Martius s.n. [M 0147319] (16)
Martius s.n. [GZU 000254251, M 0147314] (17)
Martius s.n. [B 10 0185108, G, GZU 000254252, L 0037115,
M 0147315, M 0147316, M 0147317, M 0147318, MEL
2324402, U 0002889] (17)
Martius s.n. [B 10 0185095] (18)
Martius s.n. [B 10 0086138, GZU 000254262, L 0037121,
L0037122, L 1802480, M 0147303, M 0147304] (19)
Martius s.n. [GZU 000254262, M 0147302, M 0147306]
(19)
Martius s.n. [GZU 000254261, M 0147305] (19)
Martius s.n. [GZU 000249367, M 0121041] (20)
Martius s.n. [M 0147294] (20)
Martius s.n. [B 10 0185058, B 10 0185059, GZU 000254272,
HBG, L 0037128, M 0147292, M 0147293] (21)
Martius s.n. [M 0147307] (23)
VOL. 20, NO. 2
Martius s.n. [GZU 000249360, M 0111060] (24)
Martius s.n. [GZU 000256425, M 0111063] (24)
Martius s.n. [GZU 000249359, M 0111061] (24)
Martius s.n. [GZU 000254287, M 0111065] (24)
Martius s.n. [M 0121027] (25)
Martius s.n. [M 0121029] (25)
Martius s.n. [GZU 000249322, M 0121030] (25)
Martius s.n. [M 0121033] (26)
Martius s.n. [BR 0000008810526] (27)
Martius s.n. [GZU 000249355, M 0121031] (28)
Martius s.n. [B 10 0185271, GZU 000254422, HBG, M
0147154, M 0147155, Ny 00355873] (30)
Martius s.n. [GZU 000265483, M 0147153] (30)
Martius s.n. [B 10 0185426, M 0147134, M 0147135, M
0147136, M 0147137, M 0147138, M 0147139] (31).
Martius s.n. [GZU 000254159, M 0147140] (33)
Martius s.n. [M 0111195, M 0111196] (35)
Martius s.n. [B 10 0185244, M 0121076, M 0121077, Ny
00355591] (36)
Martius s.n. [B 10 0185242, L 0036963, M 0147301] (37)
Martius s.n. [M 0147117] (39)
Martius s.n. [M 0147112] (41)
Martius s.n. [B 10 0185347, GZU 000254355] (42)
Martius s.n. [GZU 000254358, M 0111039] (42)
Martius s.n. [GZU 000254370, M 0111037] (42)
Martius s.n. [M 0111036] (43)
Martius s.n. [M 0121058] (44)
Martius s.n. [B 10 0185382, M 0121059, M 0121061] (44)
Martius s.n. [GZU 000253871, M 0111041] (45)
Martius s.n. [GZU 000253871, M 0111044] (45)
Martius s.n. [GZU 000253871, M 0111045] (45)
Martius s.n. [GZU 000254347, L, M 0111069, M 0111070,
MEL 2318483, Ny 00355728] (46)
Martius s.n. [L, M 0111192, MEL] (47)
Martius s.n. [M 0111191] (47)
Martius s.n. [M 0111178] (48)
Martius s.n. [GZU 000254368, M 0111181, M 0111182, M
0111183, M 0111184, Ny 00355782] (50)
Martius s.n. [GZU 000253888, M 0147133] (52)
Martius s.n. [M 0111177] (53)
Martius s.n. [B 10 0185417, GZU 000254373, L, M
0121081, M 0121082, MEL 2324404] (54)
Martius s.n. [GZU 000254374, GZU 000254375, M
0121084] (55)
Martius s.n. [M 0121086] (55)
Martius s.n. [GZU 000249348, M 0121088] (56)
Martius s.n. [GZU 000254384, M] (58)
Martius s.n. [GZU 000254417, M 0147211, M 0147212, M
0147213] (59)
Martius s.n. [M 0147206] (60)
Osbeck s.n. [LINN 519.1] (7)
Osbeck s.n. [S (05-4065)] (7)
Patris 41 (13)
Poeppig 2426 (30)
Poeppig 2769 (5)
Pohl 3604 (51)
Richard s.n. [P 00128463, P 00128464, P 00128465, P-JU
00662838] (13)
2015
DE MORAES AND FALCADE, LAURACEAE COLLECTED By MARTIUS
Riedel 307 (28)
Riedel 374 (7)
Riedel 480 (27)
Riedel 688 (1)
Riedel 766 (35)
Riedel 776 (35)
Riedel 2777 (8)
Riedel s.n. [B†, G 00020674, GOET [2 sheets], K 000601965,
L 0035527, LE [3 sheets], Ny 00354799] (1)
Riedel s.n. [B†, GOET 007344, GOET 007345, K
000601800, L 0036185, L 0036186, LE [2 sheets], Ny
00355045, OXF 00057026, OXF 00057027, U, US
00051077] (10)
Riedel & Luschnath s.n. = Martii Herbar. Florae Brasil. Nº
478 (Remarks # 1)
Ruíz & Pavón s.n. [B 10 0185160, BM 000947299, F
(0040316F), G 00369268, MA 811806, OXF 00000255]
(28)
Ruíz & Pavón s.n. [F (0040293F), MA 811776, MA 811778,
MA 811780] (29)
Ruíz & Pavón s.n. [A 00041960, B 10 0185150, BC 872724,
BM 000947252, BR 0000008808370, F (0040319F),
F (0040320F), FI-W, G 00020755, G 00020756, G
00020758, G (14 sheets), GZU 000261284, HAL 0010378,
MA 811777, MA 811779, MA 811781, P 00571439, P
00571440, P 00571441] (29)
Ruíz & Pavón s.n. [B 10 0247502, BM 000947285, F
(0061414F), F (0061419F), G [2 sheets], G, HAL
0103800, MA 811787, MA 811788, Ny 00355164, P
00756828] (33)
Ruíz & Pavón s.n. [B 10 0247503, F (0061415F), F
(0061416F), G, GZU 000254142, MA 811790, MA
811791, Ny 00355162] (33)
Ruíz & Pavón s.n. [B 10 0247398, F (0061413F), G, MA [2
sheets], MA 811782] (44)
Sagot 1190 (13)
Schott 5600 (41)
Schott 5607 (25)
Schott 5609 (27)
Schott 5610 (41)
Schott s.n. [B 10 0185167, GZU 000254414, P 00711066,
P 00711067] (27)
Schott s.n. [B 10 0000017] (41)
Schott s.n. [B 10 0000020] (42)
Sellow XXIV (60)
Sellow LVI (25)
Sellow 28 (24)
Sellow 144 (49)
Sellow 399 (52)
Sellow 459 (49)
Sellow 1360 (8)
Sellow 1368* (24)
Sellow 4428 (25)
197
Sellow 4495 (24)
Sellow 4803 (XXI) (40)
Sellow 5636 (45)
Sellow 5722 (25)
Sellow 5867 (55)
Sellow 5928 (55)
Sellow B 1362 c 393 (48)
Sellow B 1378 c 413 (49)
Sellow B 1384 c 419 (55)
Sellow B 1387 c 422 (60)
Sellow d 2324 (49)
Sellow s.n. [B 10 0185048] (22)
Sellow s.n. [B 10 0185227] (24)
Sellow s.n. [B 10 0185226, BR 0000008808479, CGE, E
00259309] (24)
Sellow s.n. [GZU 000249356, HAL 0010385] (25)
Sellow s.n. [GZU 000249357] (25)
Sellow s.n. [B 10 0185172] (25)
Sellow s.n. [B 10 0185175] (25)
Sellow s.n. [B 10 0185313] (40)
Sellow s.n. [B 10 0185314] (40)
Sellow s.n. [B†, GZU 000249333, VT 001552] (42)
Sellow s.n. [B 10 0000024] (43)
Sellow s.n. [B 10 0000025] (43)
Sellow s.n. [HAL 0010445] (43)
Sellow s.n. [B 10 0185388, G 00021062, P 01958844] (47)
Sellow s.n. [G-DC 00201230, HAL 0103830] (48)
Sellow s.n. [B 10 0185370] (48)
Sellow s.n. [TUB 019041] (49)
Sellow s.n. [BR 0000008767165, F (0061819F), G, GZU
000249332] (52)
Sellow s.n. [HAL 0103829] (52)
Sellow s.n. [B 10 0185414] (55)
Sellow s.n. [CGE] (55)
Sellow s.n. [E 00259499] (55)
Sellow s.n. [HAL 0010486] (55)
Sellow s.n. [LE 00006785] (55)
Sellow s.n. [M 0111066] (Remarks # 3)
Sieber Flora martin. Nº 106 (33)
Sieber Flora mixta. Nº 385 (33)
Silva Manso 278 = Martii Herbar. Florae Brasil. Nº 117 (49)
Silva Manso s.n. = Martii Herbar. Florae Brasil. Nº 1307
(27)
Spruce 3058 (14)
Swartz s.n. [BM 000758874, S (R-3166), S (09-16300)]
(23)
Swartz s.n. [SBT 11582] (23)
Tafalla s.n. [B 10 0185060, F [0040309F], FI-W 005165, G
00369022, MA 811751, MA 811752] (20)
Tweedie s.n. [E 00259433] (24)
Wied s.n. [B 10 0185169, BR 0000008808066, BR
0000008808363, GOET 000753, GZU 000254281, MEL
2324403, MEL 2324386] (26)