Pyramidellidae (Mollusca, Gastropoda, Heterobranchia) collected during the Dutch C A N C A P and MAURITANIA expeditions in the south-eastern part of the North Atlantic Ocean (part 1) C A N C A P - p r o j e c t . Contributions, no. 119 J.J. van Aartsen, E. Gittenberger & J. G o u d Aartsen, J.J. v a n , E . Gittenberger & J. G o u d . Pyramidellidae (Mollusca, Gastropoda, Heterobranchia) collected d u r i n g the D u t c h C A N C A P and M A U R I T A N I A expeditions i n the south-eastern part of the N o r t h Atlantic Ocean (part 1). Z o o l . V e r h . L e i d e n 321, 15.vi.1998:1-57, figs 1-68.— ISSN 0024-1652/ISBN 90-73239-66-4. J.J. v a n Aartsen, A d m i r a a l Helfrichlaan 33, N L 6952 G B Dieren, The Netherlands. E. Gittenberger, Department of Evertebrata, N a t i o n a l M u s e u m of N a t u r a l History, P . O . Box 9517, N L 2300 R A Leiden, The Netherlands (e-mail: gittenberger@naturalis.nnm.nl). J. G o u d , Department of Evertebrata, N a t i o n a l M u s e u m of N a t u r a l History, P . O . Box 9517, N L 2300 R A Leiden, The Netherlands (e-mail: goud@naturalis.nnm.nl). K e y w o r d s : Pyramidellidae; new species; N o r t h Atlantic Ocean. The species of the Pyramidellidae collected d u r i n g several expeditions i n the south-eastern part of the N o r t h Atlantic Ocean are listed, w i t h locality data, depth ranges, a n d notes o n nomenclature, systematics a n d distribution. The samples classified w i t h the genera Pyramidella, Tiberia, Adelactaeon, Odetta, Folinella, Ondina, Odostomia, Puposyrnola a n d Eulimella (partly) are dealt w i t h i n this paper. In total 64 species are reported f r o m the research area, 32 of w h i c h are described as n e w to science; one n o m e n n o v u m is introduced. Lectotypes of Aclis tricarinata Watson, 1897, Monoptygma puncturata Smith, 1872, Odetta sulcata de Folin, 1870, a n d Odostomia sulcifera Smith, 1872, are designated a n d figured. In an appendix, the complete locality data of the M a u r i t a n i a I & II expeditions are published for the first time. Contents Introduction 4 Materials and methods 4 Systematic p a r t 5 G e n u s Pyramidella L a m a r c k , 1799 5 G e n u s Tiberia M o n t e r o s a t o , 1875 7 G e n u s Pseudoscilla Boettger, 1901 G e n u s Liamorpha P i l s b r y , 1898 9 9 G e n u s Adelactaeon C o s s m a n n , 1895 11 G e n u s Odetta d e F o l i n , 1870 13 G e n u s Folinella D a l i & Bartsch, 1904 G e n u s Ondina de F o l i n , 1870 16 18 G e n u s Odostomia F l e m i n g , 1813 19 G e n u s Puposyrnola C o s s m a n n , 1921 40 G e n u s Eulimella Forbes & M a c A n d r e w , 1846 42 Acknowledgements 49 References 49 Appendix 53 4 Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) Introduction The seven C A N C A P expeditions to the C A N a r i a n a n d the C A P e - V e r d i a n archipelagos, d u r i n g the p e r i o d 1976-1986, a i m i n g at a better k n o w l e d g e of the m a r i n e b i o d i v e r s i t y i n that area, were o r g a n i z e d b y staff members of the N a t i o n a a l N a t u u r h i s t o risch M u s e u m (= N N M ) , L e i d e n . Research material w a s collected i n v a r i o u s w a y s , f r o m the coastal zone d o w n to 4000 m d e p t h . F o r a d d i t i o n a l i n f o r m a t i o n a n d detailed locality data concerning the C A N C A P expeditions, see the extensive report b y v a n der L a n d (1987). A d d i t i o n a l l y , the N N M participated i n the D u t c h M a u r i t a n i a I & II expeditions to the N W A f r i c a n coast. The locality data of the M a u r i t a n i a I & II expeditions h a d not yet been p u b l i s h e d completely a n d are p r o v i d e d n o w i n an a p p e n d i x . The g r o u p of genera dealt w i t h i n this p a p e r is not considered a m o n o p h y l e t i c entity. The genera are c o m b i n e d here for pragmatic reasons o n l y . The species are arranged a c c o r d i n g to s i m i l a r i t y a n d p a r t l y alphabetically. Material and methods The C A N C A P material has been deposited i n the N N M collections. The material collected d u r i n g the M a u r i t a n i a - I e x p e d i t i o n is i n the Z o o l o g i c a l M u s e u m i n A m s t e r d a m (= Z M A ) , whereas that of the M a u r i t a n i a - I I e x p e d i t i o n is i n the N N M . O n l y the samples not collected d u r i n g the expeditions m e n t i o n e d here, are cited u n d e r the head i n g ' M a t e r i a l ' w i t h a reference to the collection w h e r e that material is kept. In the f o l l o w i n g species descriptions m u c h emphasis is p l a c e d o n the type of e m b r y o n i c w h o r l s or protoconch, w h i c h is k n o w n to be i n d i c a t i v e of the k i n d of e m b r y o n i c d e v e l o p m e n t of the snails (Thorson, 1946, 1950; M a l e i k o v s k y , 1976). The course of the g r o w t h l i n e s , w h i c h are m o s t l y orthocline or prosocline a n d rarely opisthocline, is also i n d i c a t e d . O n e of the characteristics of the P y r a m i d e l l i d a e is a m o r e or less p r o m i n e n t tooth o n the c o l u m e l l a . W e d i s t i n g u i s h e d between a tooth (or teeth) p r o p e r l y speaking, a f o l d , e.g. a slight t h i c k e n i n g o n the c o l u m e l l a , a n d n o t h i c k e n i n g at a l l . Some species have "teeth" or "riblets" at the i n s i d e of the outer l i p , w h i c h are f o r m e d at a g r o w t h - s t o p ; these cannot be seen if they are too far i n s i d e the shell. The d i m e n s i o n s of the shells are g i v e n o n the basis of f u l l - g r o w n specimens; because it m a y be uncertain whether a shell s h o u l d be considered f u l l - g r o w n , the m i n i m u m sizes i n d i c a t e d are based o n a m o r e or less subjective judgement. In the P y r a m i d e l l i d a e there are over 300 n o m i n a l genera. W e agree w i t h several w o r k e r s s u c h as Schänder (1994), W i s e (1996), a n d others, that most of these taxa, w h i c h are based o n shell characters o n l y , are i l l - d e f i n e d a n d p r o b a b l y d o not represent p h y l o g e n e t i c a l l y v a l i d g r o u p s . W e have therefore u s e d genera a n d subgenera i n a fairly b r o a d sense, m a i n l y to denote m o r p h o l o g i c a l l y w e l l - d e f i n e d g r o u p s of species. The t e r m i n o l o g y u s e d for the protoconch, or e m b r y o n i c w h o r l s , f o l l o w s v a n A a r t sen (1981,1987) a n d Schänder (1994). - T y p e A : angle of the p r o t o c o n c h w i t h the shell axis r a n g i n g f r o m 90° to c. 110°. A - I is p l a n i s p i r a l or p l a n o r b i d , w i t h a l l p r o t o c o n c h w h o r l s m a i n l y i n one plane (fig. 68). A - I I is h e l i c o i d , w i t h a clearly p r o t r u d i n g nucleus (fig. 57). In some cases the first Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) 5 teleoconch w h o r l overlaps the protoconch, b u t e v e n i n those cases the t o p w h o r l c a n still be seen. - T y p e B: angle of the p r o t o c o n c h w i t h the axis of the shell r a n g i n g f r o m 120° to c. 160°. H e r e the t o p w h o r l u s u a l l y cannot be seen because it is h i d d e n w i t h i n the first teleoconch w h o r l (fig. 66); the rest of the e m b r y o n i c w h o r l s is visible, h o w e v e r . - T y p e C : angle almost 180°. H e r e the t o p w h o r l s have almost d i s a p p e a r e d into the first teleoconch w h o r l (fig. 61). The types B a n d C are u s u a l l y referred to as 'intorted'. It s h o u l d be e m p h a s i z e d that there is v e r y little intraspecific v a r i a t i o n i n the structure of the protoconch, w h i c h is often diagnostic. The f o l l o w i n g abbreviations are used for collections w i t h p y r a m i d e l l i d s used i n the present paper: A D , J.J. v a n Aartsen, Dieren; B M N H , The N a t u r a l H i s t o r y M u s e u m , L o n d o n ; H U J , H e b r e w U n i v e r s i t y , Jerusalem; M N H N , Muséum N a t i o n a l d'Histoire N a t u relle, Paris; N M W , N a t i o n a l M u s e u m of Wales, Cardiff; R M N H , N a t i o n a l M u s e u m of N a t u r a l H i s t o r y , L e i d e n ; U S N M , U . S . N a t i o n a l M u s e u m of N a t u r a l H i s t o r y , W a s h i n g ton; Z M A , Z o o l o g i c a l M u s e u m , A m s t e r d a m . The abbreviation M W H is u s e d for the f o r m e r M a l a c o l o g i c a l W o r k i n g - g r o u p , the H a g u e . T h i s g r o u p w a s active i n the sixties, w h e n the first author w a s a m e m b e r . M a n y n e w species i n this paper are n a m e d after f e l l o w - m e m b e r s of that time, i n remembrance of the m a n y pleasant h o u r s spent together. Systematic part G e n u s Pyramidella L a m a r c k , 1799 T y p e species (by m o n o t y p y ) : Trochus dolabratus Linné, 1758. The generic name Pyramidella L a m a r c k , 1799, has been-validated u n d e r the P l e n a r y P o w e r s of the I C Z N , O p i n i o n 386 (1955). T h e n a m e Obeliscus A n o n y m u s , 1797 [= Obeliscus G r a y , 1847] has also been u s e d for this genus, whereas the n a m e Pyramidella w a s u s e d for the genus n o w k n o w n as Otopleura Fischer, 1885. Pyramidella (Pyramidella) dolabrata (Linné, 1758) Trochus dolabratus Linné, 1758: 760, spec. 522. Pyramidella terebelloides; K i r a , 1962: 203, p i . 69 fig. 22. Pyramidella terebella; Habe, 1962: 86, p l . 41 fig. 40. Pyramidella terebellum; Cernohorsky, 1972:199, p i . 56 fig. 12,12a; L i n d n e r , 1982: 212, p i . 51 fig. 3. Pyramidella dolabrata; K a y , 1979: 412, 413, fig. 133A; Abbott & Dance, 1982: 277, figure. M a t e r i a l . — Cape Verde Islands: 6.058/2, 6.064/2, 7.076/2, 7.077/1. N o m e n c l a t u r e . — W e agree w i t h C e r n o h o r s k y (1972), K a y (1979), a n d others, that P. dolabrata (Linné, 1758) is a s o m e w h a t variable species a n d that Helix terebellum Müller, 1774, Obeliscus terebelloides A . A d a m s , 1854, a n d several a d d i t i o n a l n o m i n a l taxa described as separate species b y L a s e r o n (1959) s h o u l d be considered s y n o n y m s . D i s t r i b u t i o n . — I n the East A t l a n t i c P. dolabrata has been recorded f r o m L u a n d a , 6 Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) West A f r i c a , b y S o w e r b y i n Reeve (1865: spec. 13) a n d D a u t z e n b e r g (1912: 55), f r o m St. H e l e n a b y S m i t h (1890: 275), f r o m S. Thome b y T o m l i n et a l . (1914: 256), f r o m the G u l f of Guinée b y T a l a v e r a (1981:166), a n d f r o m the C a p e V e r d e Islands b y D a u t z e n berg (1912: 55), Saunders (1977: 7) a n d v o n C o s e l (1982: 21). T h i s i m p l i e s that the type locality of this species, v i z . " i n A f r i c a " (Linné, 1767: 1231, n o . 601) m a y be right after all, a n d not erroneous as suggested b y D o d g e (1958: 204). D e p t h r a n g e . — 29-44 m . Pyramidella (Longchaeus) schanderi n o m . n o v . (fig-1) Obeliscus suturalis v o n M a l t z a n , 1885: 26. Schänder, 1994: 49, 66, fig. 7d. N o t Pyramidella suturalis Lea, 1846: 258, p i . 36 fig. 63. M a t e r i a l . — Cape Verde Islands: 6.014/1, 6.025/1, 6.027/1, 6.054/1, 6.056/2, 6.059/>25, 6.064/14, 6.082/3, 6.083/2, 6.084/4, 6.107/14, 6.128/2, 6.130/7, 6.132/2, 6.141/4, 6.156/5, 6.159/1, 6.160/6, 6.164/3, 6.171/1, 7.031/1, 7.042/3, 7.043/9, 7.064/1, 7.065/17, 7.067/1, 7.068/4, 7.075/>25, 7.079/>25, 7.080/>25, 7.088/1, 7.089/6, 7.090/1, 7.093/1, 7.097/3, 7.106/1, 7.115/1, 7.128/4, 7.160/4, 7.161/2. West A f r i c a (Mauritania): 3.133/1. D i s t r i b u t i o n . — T h i s species w h i c h w a s o r i g i n a l l y described f r o m Gorée, West A f r i c a , turns out to be one of the most c o m m o n p y r a m i d e l l i d species of the C a p e V e r d e Islands. In o u r material it is represented b y n u m e r o u s specimens. N o m e n c l a t u r e . — A t first w e w e r e n o t sure about the differences between this species a n d Pharcidella (?)inopinata (Schänder, 1994). Schander (1994: 48, 66, f i g . 7c) describes a n d figures a s p e c i m e n of a rather different f o r m , b u t this c o u l d have been a n aberrant specimen. Schänder h i m s e l f (in litt., 1997) considers o u r shells to b e l o n g to Pyramidella suturalis ( v o n M a l t z a n , 1885). W e agree w i t h this o p i n i o n . U n f o r t u n a t e l y , the w e l l - k n o w n name Pyramidella suturalis ( v o n M a l t z a n , 1885) is p r e o c c u p i e d b y Pyramidella suturalis L e a , 1846. Therefore, w e here p r o p o s e Pyramidella schanderi n o m . n o v . for the f o r m e r t a x o n (6.2 x 2.4 m m ; f i g . 1). W e d o not classify this species w i t h Pharcidella D a l l , 1889, as this (sub)genus is characterized b y p r o m i n e n t axial ribs, w h i c h P. schanderi n o m . n o v . does not possess. A tentative placement into the subgenus Longchaeus Mörch,1875, seems m o r e a p p r o priate i n v i e w of the clear p e r i p h e r a l sulcus. D i f f e r e n t i a t i o n . — Pyramidella schanderi n o m . n o v . is v e r y s i m i l a r to the West A t l a n tic P. (Longchaeus) Candida Mörch, 1875, b u t shells of the latter species are somewhat larger a n d less slender, w h i l e the sutures are even m o r e h o r i z o n t a l (see W a r m k e & A b b o t t , 1962: p i . 28, f i g . d). M o r e o v e r , the g r o w t h l i n e s i n o u r shells are orthocline to slightly opisthocline, whereas they are prosocline i n P. Candida. T h e same character distinguishes the West A t l a n t i c P. crenulata H o l m e s , 1859 (see W i s e , 1996: 475, f i g . 23). T h e figure p u b l i s h e d b y A b b o t t (1974: 291, 290, f i g . 3461) does not represent P. Candida. D e p t h r a n g e . — 18-95 m . E t y m o l o g y . — T h i s species is r e n a m e d after D r . C . Schänder, f r i e n d of the first author a n d w e l l - k n o w n for h i s contributions to the k n o w l e d g e of the W e s t A f r i c a n Pyramidellidae. 7 Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) G e n u s Tiberia Monterosato, 1875 T y p e species (by subsequent designatio n [ I C Z N A r t . 69(a)(i)(l)]): Pyramidella minuscula Monterosato,1880. Pyramidella minuscula Monterosato, 1874, is a n o m e n n u d u m . T h i s species w a s form a l l y described b y Monterosato o n l y i n 1880, as Pyramidella (Tiberia) minuscula. A s discussed b y v a n A a r t s e n & C o r g a n (1996: 180), the genus g r o u p names Tiberia Jeffreys, 1884, a n d Tiberiola C o s s m a n n , 1900 (= Tiberia Jeffreys, 1884, not Monterosato, 1875) are s y n o n y m s . T h e genus TiberiellaC o e n , 1933, w i t h the type species Tiberia pretiosa C o e n , 1933, is not closely related because the h o l o t y p e of T. pretiosa ( H U J 7301) is a f o r m of the w e l l - k n o w n Chrysallida obtusa ( B r o w n , 1827) ( v a n A a r t s e n , u n p u b l i s h e d results). Tiberiella is a j u n i o r s y n o n y m of Parthenina B u c q u o y , D a u t z e n b e r g & D o l l fus,1883, because b o t h taxa have the same type species. Tiberia minusculoides spec. n o v . (fig. 2) M a t e r i a l . — H o l o t y p e ( N N M 57270): 7.049, Cape Verde Islands, S W of M a i o , Ponta Inglez/ Ponta Prêta; 15°06'N 23°24'W; depth 273 m , yellow-grey sandy clay; v a n V e e n grab; 25.viii.1986. Paratypes ( N N M 57271-57278). Canary Islands: 2.065/1, 2.066/1, 2.074/1, 4.060/6, 4.082/1. Cape Verde Islands: 6.124/1, 7.038/1, 7.061/5 fragments. D e s c r i p t i o n . — Shell s o l i d ; the spire f o r m i n g a slender cone w i t h straight sides a n d the p e r i p h e r y o n l y w e a k l y angled. C o l o u r y e l l o w i s h - w h i t e w i t h a y e l l o w - b r o w n s p i ral b a n d a l o n g the p e r i p h e r y ; o n the a d a p i c a l part of the l o w e r w h o r l s , b o r d e r i n g the suture, another b a n d of the same colour. E m b r y o n i c w h o r l s p l a n o r b i d , type B to C . Teleoconch f o r m e d b y five to six strongly flattened w h o r l s . Suture s i m p l e a n d straight. G r o w t h l i n e s c h a n g i n g f r o m s o m e w h a t prosocline o n the t o p w h o r l s to somew h a t opisthocline a n d flexuous o n the l o w e r w h o r l s . O n l y m i c r o s c o p i c s p i r a l striae can be detected. O u t e r l i p gently c u r v e d . There are n o teeth o n the i n s i d e of the outer l i p . U m b i l i c u s r o u n d a n d deep. There is one tooth-like f o l d at the a d a p i c a l side of the c o l u m e l l a ; m o r e basally there are t w o m u c h w e a k e r folds. D i m e n s i o n s : H . 5.0-6.2 m m , W . 2.5-2.75 m m ; h o l o t y p e : 5.2 x 2.7 m m . D i f f e r e n t a t i o n . — T h i s species is rather s i m i l a r to the w e l l - k n o w n Tiberia minuscula (Monterosato, 1880) a n d has most p r o b a b l y been confused w i t h that species i n the past. Shells of T. minuscula, h o w e v e r , are smaller, v i z . about 4 m m h i g h w i t h the same n u m b e r of w h o r l s , u s u a l l y m o r e c y l i n d r i c a l a n d not p r o v i d e d w i t h a y e l l o w - b r o w n s p i r a l b a n d b e l o w the u p p e r suture of the l o w e r w h o r l s . D e p t h r a n g e . — 273-886 m . E t y m o l o g y . — T h e epithet minusculoides refers to the s i m i l a r i t y w i t h minuscula. Tiberia apicifusca spec. n o v . (figs 3, 52) M a t e r i a l . — H o l o t y p e ( N N M 57279): 6.012, Cape Verde Islands, S of Säo-Tiago; 14°53'N depth c. 500 m , grey b r o w n m u d ; v a n V e e n grab; 5.vi.l981. 23°30'W; Paratypes ( N N M 57280-57291). Cape Verde Islands: 6.010/2, 6.011/3, 6.012/7, 6.013/1, 6.017/5, 8 Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) 6.027/1, 7.003/3, 7.006/1, 7.014/1, 7.049/1, 7.100/2, 7.129/2 fragments. D e s c r i p t i o n . — Shell rather t h i n ; the spire f o r m s a slender cone w i t h rather straight sides. The regular, conical, general shape m a y be d i s t u r b e d s o m e w h a t w h e n the l o w e r part of the w h o r l s is relatively b r o a d . C o l o u r cream to w h i t e , w i t h o u t s p i r a l c o l o u r b a n d s b u t w i t h a y e l l o w i s h - b r o w n top. E m b r y o n i c w h o r l s intorted, of type C , a n d c o l o u r e d light b r o w n . Teleoconch f o r m e d b y five to six w h o r l s ; the i n i t i a l ones are nearly flat, whereas the f o l l o w i n g ones are increasingly m o r e convex. Suture s i m ple a n d clearly m a r k e d . G r o w t h l i n e s m a r k e d l y opisthocline; shell w i t h o u t a d d i t i o n a l sculpture. O u t e r l i p gently c u r v e d , w i t h o u t teeth. U m b i l i c u s w i d e , o p e n a n d deep. O n l y a single tooth-like f o l d is clearly visible; t w o a d d i t i o n a l , v e r y w e a k folds, b e l o w the a d a p i c a l one, m a y also be discernible. D i m e n s i o n s : H . 4.5-5.0 m m , W . 2.3-2.5 m m ; h o l o t y p e 4.5 x 2.3 m m . D i f f e r e n t i a t i o n . — Tiberia apicifusca spec. n o v . can be r e c o g n i z e d easily because of its light b r o w n protoconch, thus d i f f e r i n g f r o m b o t h T. minuscula a n d T. minusculoides spec. n o v . The p r o t o c o n c h is also m o r e strongly intorted than i n these t w o species. N o n e of the 28 specimens that c o u l d be s t u d i e d h a d a colour b a n d . D e p t h r a n g e . — 273-970 m . E t y m o l o g y . — T h i s species is n a m e d apicifusca because of the b r o w n c o l o u r of the apex. Tiberia micalii (Penas & R o l a n , 1997) (fig. 4) Sayella micalii Penas & Rolan, 1997: 36, figs 1-5. M a t e r i a l . — Cape Verde Islands: 7.065/1, 7.067/1 (fig. 4). D e s c r i p t i o n . — O n l y t w o fragmentary specimens were f o u n d , b u t these are quite characteristic. The complete shell p r o b a b l y h a d a n elongated o b l o n g shape, w i t h a relatively h i g h last w h o r l . C o l o u r cream, w i t h a n a r r o w y e l l o w - b r o w n s p i r a l b a n d just above the p e r i p h e r y . E m b r y o n i c w h o r l s p l a n o r b i d , of type B, diameter about 450 jum. O n l y a f e w w e a k l y convex teleoconch w h o r l s are k n o w n . Suture i m p r e s s e d . G r o w t h lines orthocline to s l i g h t l y prosocline. There is n o particular m i c r o s c u l p t u r e . O u t e r l i p rather straight. U m b i l i c u s n a r r o w b u t deep. O n the c o l u m e l l a t w o teeth can be seen, b u t b o t h fragments are b r o k e n at the l o w e r c o l u m e l l a r part. N o m e n c l a t u r e . — A l t h o u g h Penas & R o l a n (1997: 37) describe t w o c o l u m e l l a r teeth (their figure even s h o w s three), they place their species i n Sayella D a l l , 1885. H o w e v e r , i n species of that genus, hitherto k n o w n o n l y f r o m eastern N o r t h A m e r i c a , there is o n l y one m o d e r a t e l y p r o m i n e n t f o l d o n the c o l u m e l l a . Because of the three c o l u m e l l a r teeth, the s m o o t h , b a n d e d w h o r l s , a n d the f o r m of the u m b i l i c u s , w e place this species i n Tiberia. It differs f r o m T. minuscula a n d related forms b y its p r o m i n e n t teeth a n d the m u c h m o r e slender a n d o v a l f o r m . D e p t h r a n g e . — 39-42 m . Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) 9 G e n u s Pseudoscilla Boettger, 1901 T y p e species (by m o n o t y p y ) : Oscilla (Pseudoscilla) miocaenica Boettger, 1901. Oscilla (Pseudoscilla) miocaenica Boettger, 1901, is f i g u r e d b y Z i l c h (1935: 227, f i g . 2). T h i s species is s u r p r i s i n g l y s i m i l a r to Odostomia (Parthenia) exarata Carpenter, 1857, as f i g u r e d b y D a l i & Bartsch (1909: p l . 19 f i g . 2). Because O. exarata is the type species (by o r i g i n a l designation) of Miraldiella C o s s m a n n , 1921, the latter n o m i n a l taxon is a j u n ior s y n o n y m of Pseudoscilla. Pseudoscilla babylonia (C.B. A d a m s , 1845) (figs 5, 53) Chemnitzia babylonia C . B. A d a m s , 1845: 6. Aclis tricarinata Watson, 1897: 255, p i . 20 fig. 23. Cingulina babylonia; W a r m k e & Abbott, 1961:148; de Jong & Coomans, 1988:120, p i . 19 fig. 637. M a t e r i a l . — Canary Islands: 4.001/1, 4.020/1, 4.027/1, 4.036/1, 4.073/1, 4.K04/1; G r a n C a n a d a , AD4888/3; Lanzarote, AD24813/1 a n d AD27559/2; L a Graciosa, AD27510/1; Fuertaventura, AD27665/3. Cape V e r d e Islands: 6.004/1, 6.005/2, 6.006/2, 6.010/3, 6.015/1, 6.024/2, 6.040/7, 6.103/1, 6.105/2, 6.132/2, 6.134/3, 6.149/3, 7.007/2, 7.008/2, 7.028/2, 7.030/2, 7.031/1, 7.050/1, 7.064/1, 7.079/6, 7.080/25, 7.100/3, 7.101/1, 7.102/1, 7.119/1, 7.121/2. D i f f e r e n t i a t i o n . — Pseudoscilla babylonia is one of the relatively f e w species of P y r a m i d e l l i d a e i n w h i c h the e m b r y o n i c w h o r l s are not s m o o t h b u t decorated ( w i t h s p i r a l ribs). T h e sculpture of the p r o t o c o n c h can o n l y be seen i n w e l l preserved specimens, h o w e v e r (fig. 53). N o m e n c l a t u r e . — A c c o r d i n g to Faber (1988: 81), w h o p l a c e d this species i n Liamorpha P i l s b r y , 1898, Pyramidelloides judithae U s t i c k e , 1959, is a s y n o n y m . See hereafter w i t h Liamorpha elegans (de F o l i n , 1870). A s t u d y of the three syntypes of Aclis tricarinata W a t s o n , 1897 ( N M W 1955.158. 00655), as w e l l as t w e n t y conspecific other specimens i n the same collection ( N M W 1955.158), a l l o r i g i n a t i n g f r o m M a d e i r a , p r o v e d c o n c l u s i v e l y that Aclis tricarinata W a t son, 1897, is a j u n i o r s y n o n y m of Pseudoscilla babylonia (C. B. A d a m s , 1845). O n e of the syntypes m e n t i o n e d has been selected as lectotype (2.2 x 1.0 m m ; fig.5). D i s t r i b u t i o n . — T h i s w e l l - k n o w n C a r i b b e a n species turns out to be w i d e l y d i s t r i b u t e d t h r o u g h o u t the C a n a r y Islands a n d the C a p e V e r d e Islands. D e p t h r a n g e . — 23-354 m . G e n u s Liamorpha P i l s b r y , 1898 T y p e species (by m o n o t y p y ) : Lia decorata de F o l i n , 1873. O r i g i n a l l y described as Lia (de F o l i n , 1873), this n a m e t u r n e d out to be p r e o c c u p i e d b y b o t h Lia Eschscholtz, 1829, a n d Lia Mörch, 1852. F o r that reason P i l s b r y (1898: 323) i n t r o d u c e d the replacement n a m e Liamorpha. Liamorpha elegans (de F o l i n , 1870) Mathilda elegans de Folin, 1870: 212, p i . 26 fig. 11 [not p i . 28 fig. 15]. 10 Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) Figs 1-5. Pyramidella, Tiberia a n d Pseudoscilla species. 1, Pyramidella schanderi n o m . nov., 6.2 m m , sta. 7.079, Cape V e r d e Islands, W of Ilhéu de Sal Rei, 16°10'N 23°00'W, depth 60 m , 28-viii-1986; 2, Tiberia minusculoides spec, nov., holotype ( N N M 57270), 5.2 m m , sta. 7.049, C a p e V e r d e Islands, S W of M a i o , Ponta Inglez/Ponta Prêta, 15°06'N 23°14'W, depth 273 m , 25-viii-1986; 3, Tiberia apicifusca spec, nov., holotype ( N N M 57280), 4.5 m m , sta. 6.012, Cape V e r d e Islands, S of Säo Tiago, 14°53'N 23°30'W, depth about 500 m , 5-vi-1981; 4, Tiberia micalii (Penas & Rolân, 1997), 2.2 m m , sta. 7.067, Cape V e r d e Islands, W of Boa Vista, S W of Ilhéu Calheta d o Velho, 16°11'N 22°59'W, depth 42 m , 27-viii-1986; 5, Pseudoscilla babylonia (C.B. A d a m s ) (lectotype of Aclis tricarinata Watson), 2.2 m m , M a d e i r a , coli N M W 1955.158.00655. 11 Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) Miraida elegans; Hoenselaar & Moolenbeek 1990: 65, figs 1-4; R o l a n & Fernandes, 1993: 6, p l . 11 figs 13; N o f r o n i & Schander, 1994: 9, fig. e. M a t e r i a l . — West A f r i c a (Mauritania): 3.109/10, 3.128/1, 3.172/>25, 3.178/5, 3.179/4, 3.189/2; M.011/1, M.030/2, M.032/2, M.033/2, M.043/8, M.044/1, M.045/2, M.106/6, M.113/3, M.115/4, M.118/7,M.121/3, M.122/1. D i s t r i b u t i o n . — A v e r y characteristic a n d relatively c o m m o n W e s t A f r i c a n species, o r i g i n a l l y described f r o m Senegal. H o e n s e l a a r & M o o l e n b e e k (1990: 65, 66) reported the f i n d i n g of a s p e c i m e n f r o m the M e d i t e r r a n e a n , Balearic i s l a n d Formentera (Spain). N o m e n c l a t u r e . — The generic placement i n Liamorpha P i l s b r y , 1898, is based o n the h e l i c o i d p r o t o c o n c h of b o t h L. elegans a n d L. decorata, the type species. V a n A a r t sen (1984:137) already m e n t i o n e d the p o s s i b i l i t y that Liamorpha P i l s b r y , 1898, is s y n o n y m o u s w i t h Miralda A . A d a m s , 1863, as suggested b y D a l i & Bartsch (1906: 356). The type species of Miralda, Parthenia diadema A. A d a m s , 1860, is problematic, h o w e v er. The (possible) type s p e c i m e n [ B M N H 1878.1.28.348, ex H . A d a m s coll.] has a n intorted top a n d teeth at the i n s i d e of the outer l i p . A s this is quite different f r o m the d e s c r i p t i o n b y D a l i & Bartsch (1906: 356, p l . 17 f i g . 2) of a s p e c i m e n that they also considered to b e l o n g to Miralda diadema, w e prefer to use Liamorpha P i l s b r y , 1898. D e p t h r a n g e . — 14-114 m . G e n u s Adelactaeon C o s s m a n n , 1895 T y p e species (by o r i g i n a l designation): Tornatella papyraceus Basterot, 1825. The genus Adelactaeon, w i t h the type species A. papyraceus (Basterot, 1825) f r o m the M i o c e n e of France, has been considered s y n o n y m o u s w i t h Myonia A . A d a m s , 1860, not D a n a , 1847, b y C o s s m a n n (1895: 54) himself, as w e l l as b y several later authors. T h i s v i e w is accepted here as w e l l . B o t h A . A d a m s (1860: 406) a n d C o s s m a n n (1895: 55) classify their genus w i t h the A c t e o n i d a e . G l i b e r t (1952: 387, 388, p i . 14 f i g . 8) discusses a n d figures the type species, w h i c h he considers i d e n t i c a l w i t h Tornatella elatav o n K o e n e n , 1882 (3.0 x l . 6 m m ; f i g . 6), k n o w n f r o m the north-west E u r o p e a n M i o c e n e . W e consider Phasianema W o o d , 1842, a n d Leucotina A . A d a m s , 1860, different genera. Leucotina has recently been s h o w n b y H o r i et a l . (1995) to b e l o n g to the A m a t h i n i d a e . W h e t h e r this is also true for Adelactaeon, w e d o not k n o w . The genus Phasianema is based o n Phasianema sulcata W o o d , 1842, its type species b y m o n o t y p y . T h i s species is s i m i l a r to Adelactaeon i n general shape, b u t it differs b y the n a r r o w s p i r a l ribs, separated b y m u c h broader interstices, w h i c h are not crossed b y t h i c k e n e d lamellar g r o w t h l i n e s as i n Adelactaeon. Phasianema sulcata is f i g u r e d b y G l i b e r t (1958: p i . 2 f i g . 12). Nerita costata B r o c c h i , 1814, not G m e l i n , 1791, is i n d i c a t e d as type species of Phasianema b y Thiele (1929: 233) a n d W e n z (1940: 849), b u t this is a n error. Brocchi's species, the correct n a m e of w h i c h is Fossarus clathratus P h i l i p p i , 1844 (see M i e n i s , 1973: 87), is the type species of Clathrella Récluz, 1864 (by m o n o t y p y : see v a n A a r t s e n et al., 1984: 50, p l . 123 f i g . 241). Recently it has been suggested b y C a m p b e l l et al. (1995:168) that Carinorbis C o n r a d , 1862, is a n senior s y n o n y m of Clathrella Récluz, 1864. 12 Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) Adelactaeon puncturata (Smith, 1872) (fig- 7) Monoptygma (Myonia) puncturata Smith, 1872: 734, p l . 75 fig. 16. N o m e n c l a t u r e . — T h e type lot [ B M N H 1870.1.14.11] consists of five shells w i t h a n intorted top of type C , a p r o m i n e n t f o l d o n the c o l u m e l l a , a n d a s p i r a l sculpture consisting of seven to eight spirals, w h i c h are broader than the interstices. I n these interstices, the lamellar g r o w t h l i n e s are thickened a n d v e r y conspicuous, w h e n c e the name. W e have selected the largest s p e c i m e n (4.4 x 2.2 m m ) as lectotype (fig. 7). D i s t r i b u t i o n . — O r i g i n a l l y described f r o m W h y d a h , West A f r i c a , this species has never been f o u n d again as far as w e k n o w . I n o u r material w e f o u n d a n u m b e r of fragments or t o p w h o r l s w h i c h m i g h t b e l o n g to it. A d d i t i o n a l l y w e d i s c o v e r e d t w o other Adelactaeon species, w h i c h clearly differ f r o m A. puncturata a n d are described below. Adelactaeon elongata spec. n o v . (fig. 8) M a t e r i a l . — H o l o t y p e ( N N M 57294): M.002, M a u r i t a n i a , Baie de Cansado; 20°53'N 17°02'W; depth 8 m , anoxic fine m u d ; v a n V e e n grab; 5.vi.l988. Paratypes. M a u r i t a n i a : M . 0 0 2 / l ( N N M 57263); III/2 ( Z M A 3.97.005). D e s c r i p t i o n . — Shell slender conical to s l i g h t l y o v a l . C o l o u r whitish-transparent. E m b r y o n i c w h o r l s intorted, of type C . Teleoconch of four to five m o d e r a t e l y convex w h o r l s , w h i c h r a p i d l y increase i n size. T h e height of the w h o r l s is 60% of their w i d t h . Suture i n d e n t e d . G r o w t h l i n e s thickened, r i b - l i k e , a n d clearly prosocline. Sculpture f o r m e d b y s p i r a l ribs a n d interstices w h i c h are about equal i n w i d t h . A b o u t eight interstices, crossed b y the thickened g r o w t h l i n e s , are present o n each w h o r l . O n the base of the last w h o r l , six to eight a d d i t i o n a l spirals are seen. O u t e r l i p gently c u r v e d a n d sharp, c o l u m e l l a nearly straight. U m b i l i c u s n a r r o w b u t conspicuous. T o o t h w e l l - d e v e l o p e d , v e r y h i g h o n the c o l u m e l l a . D i m e n s i o n s : H . 7.6 m m , W . 3.1 m m ; h o l o t y p e : 7.6 x 3.1 m m . D i f f e r e n t i a t i o n . — A l t h o u g h e v i d e n t l y congeneric w i t h A. puncturata (Smith, 1872), this n e w species clearly differs b y the m o r e slender shells, w i t h spirals a n d interstices w h i c h are about e q u a l l y b r o a d . I n A. puncturata the interstices are n a r r o w e r than the spirals. D e p t h r a n g e . — 0-8 m . E t y m o l o g y . — The epithet elongata refers to the shape of the shell. Adelactaeon lilyae spec. n o v . (fig. 9) M a t e r i a l . — H o l o t y p e ( Z M A 3.97.006): B3, M a u r i t a n i a , Banc d ' A r g u i n ; 19°37.4'N 16°52.1'W; depth 21-34 m ; 17.V.1988. Paratypes ( N N M 57295-57296). M a u r i t a n i a : M.065/1, 3.109/3. Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) 13 D e s c r i p t i o n . — Shell o v a l to egg-shaped, w i t h p u p o i d a l t o p . C o l o u r w h i t i s h . E m b r y o n i c w h o r l s intorted, of type C . Teleoconch of three to f o u r w h o r l s w h i c h are m o d e r a t e l y convex, the height m e a s u r i n g s l i g h t l y m o r e t h a n half the w i d t h . Suture i n d e n t e d . G r o w t h l i n e s prosocline, t h i c k e n e d to t h i n lamellae. Sculpture consisting of s p i r a l ribs a n d equally b r o a d interstices; o n the i n i t i a l teleoconch w h o r l s there are about eight ribs a n d o n the last w h o r l there are eight a d d i t i o n a l ones. O u t e r l i p gently c u r v e d ; the c o l u m e l l a nearly straight. U m b i l i c u s s m a l l b u t conspicuous. T o o t h p r o m i nent b u t situated v e r y d e e p l y inside the shell, so that it cannot easily be detected. D i m e n s i o n s : H . 3.5-5.0 m m , W . 1.9-2.5 m m ; h o l o t y p e : 3.5 x 1.9 m m . D i f f e r e n t i a t i o n . — Shells of this species differ f r o m those of Adelactaeon elongata spec. n o v . b y their m o r e egg-shaped f o r m , the s o m e w h a t l o w e r w h o r l s as c o m p a r e d to their w i d t h , a n d the m o r e p u p o i d a l apical part, w i t h smaller e m b r y o n i c w h o r l s . The h o l o t y p e is a relatively slender specimen; some of the paratypes are comparativel y broader. D e p t h r a n g e . — 14-25 m . E t y m o l o g y . — T h i s species is n a m e d after M r s . L i l y de Klein-Steenbergen, w h o a l w a y s s u p p o r t s the first author b y collecting shells a n d shellgrit w h e r e v e r she goes. G e n u s Odetta de F o l i n , 1870 T y p e species (by subsequent d e s i g n a t i o n [ I C Z N A r t . 69 (a)(i)(l); see v a n A a r t s e n , 1984]): Odetta sulcata de F o l i n , 1870 . T h i s generic n a m e has been discussed b y v a n A a r t s e n (1984: 135). T h e status of the type species c o u l d be ascertained. See also the notes o n O. sulcata. Odetta dekleini spec. n o v . (fig. I D M a t e r i a l . — H o l o t y p e ( N N M 57297): 7.007, Cape Verde Islands, S W of Säo Tiago; 14°54'N 23°38'W; depth 420 m , m u d d y sand; v a n V e e n grab; 20.viii.1986. Paratypes ( N N M 57298-57313). Cape V e r d e Islands: 6.005/2, 6.006/2, 6.009/1, 6.015/4, 6.040/1, 7.004/1, 7.007/6, 7.050/3, 7.066/1, 7.067/1, 7.119/2, 7.121/2, 7.128/5, 7.129/6, 7.161/1. D e s c r i p t i o n . — Shell v e r y s m a l l , egg-shaped w i t h a s o m e w h a t p u p o i d a l t o p . C o l o u r w h i t i s h transparent. E m b r y o n i c w h o r l s intorted, of type B. Teleoconch consisting of three to three a n d a half o n l y s l i g h t l y convex w h o r l s . Suture not w e l l m a r k e d , m o s t l y c o i n c i d i n g w i t h one of the s p i r a l grooves. G r o w t h l i n e s thickened, prosocline, especially p r o n o u n c e d w i t h i n the s p i r a l grooves. W i t h three to f o u r s p i r a l grooves, w h i c h measure about half the w i d t h of the s p i r a l b a n d s . O n the last w h o r l there are about five a d d i t i o n a l grooves b e l o w the p e r i p h e r y . I n a l l these grooves the t h i c k e n e d g r o w t h l i n e s c a n easily be seen. O u t e r l i p t h i n a n d s m o o t h i n s i d e . There is no u m b i l i c u s . O n l y a n i n c o n s p i c u o u s tooth is discernible, situated d e e p l y w i t h i n the aperture. D i m e n s i o n s : H . 1.1-1.3 m m , W . 0.6 m m ; h o l o t y p e 1.1 x 0.6 m m . D i f f e r e n t i a t i o n . — T h i s Odetta species differs f r o m b o t h O. sulcata a n d O. marci spec. n o v . b y the fact that the grooves between the s p i r a l ribs are not s m o o t h a n d narr o w b u t relatively b r o a d , w i t h the t h i c k e n e d g r o w t h l i n e s p r o m i n e n t l y present. 14 Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) D e p t h r a n g e . — 42-420 m . E t y m o l o g y . — T h i s species is n a m e d after D r W . J. de K l e i n , a f r i e n d a n d former colleague of the first author, w h o a l w a y s s h o w s a k e e n interest i n n a t u r a l history. Odetta marcispec. n o v . (fig. 12) M a t e r i a l . — H o l o t y p e ( N N M 57314): M.078, M a u r i t a n i a , off Banc d ' A r g u i n ; 20°00'N 17°21'W; depth 41 m , dark-grey m u d d y fine sand; v a n V e e n grab; 14.vi.1988. Paratypes ( N N M 57315). M a u r i t a n i a : M.064/2. D e s c r i p t i o n . — Shell f o r m i n g a slender cone, slightly n a r r o w e d a n d s o m e w h a t p u p o i d a l at the t o p . C o l o u r w h i t i s h . E m b r y o n i c w h o r l s intorted, of type B . Teleoc o n c h consisting of five a n d a half o n l y s l i g h t l y convex w h o r l s . Suture clearly m a r k e d , channelled. G r o w t h l i n e s d i f f i c u l t to detect, most p r o b a b l y about orthocline. W i t h one or t w o incised s p i r a l grooves at the u p p e r part of the i n i t i a l teleoconch w h o r l s ; the l o w e r w h o r l s w i t h four to five s u c h grooves, w h i c h cover the total height of the w h o r l . Shell base w i t h a d d i t i o n a l l y eight to ten grooves, w h i c h are increasingly m o r e n a r r o w l y spaced t o w a r d s the u m b i l i c u s . O u t e r l i p t h i n , s m o o t h inside. There is a w e l l - m a r k e d u m b i l i c a l chink. C o l u m e l l a r tooth conspicuous. D i m e n s i o n s : H . 3.8 m m , W . 1.5 m m ; h o l o t y p e 3.8 x 1.5 m m . D i f f e r e n t i a t i o n . — Odetta marci spec. n o v . is k n o w n w i t h o n l y one f u l l g r o w n a n d t w o i m m a t u r e specimens. Still this species is so m u c h different f r o m the w e l l - k n o w n O. sulcata that w e d o not hesitate to describe it as n e w to science. I n O. marci spec, n o v . the first groove appears just b e l o w the u p p e r suture a n d the l o w e r w h o r l s carry four to five grooves, whereas O. sulcata has the first s p i r a l groove at or b e l o w the m i d d l e of the w h o r l a n d the f i n a l w h o r l s carry o n l y t w o to three grooves. T h e e m b r y o n i c w h o r l s are also quite different, i n d i c a t i n g a different m o d e of d e v e l o p ment. D e p t h r a n g e . — 24-41 m . E t y m o l o g y . — T h i s species is n a m e d after M a r c S. S. L a v a l e y e , participant i n the M a u r i t a n i a expeditions a n d keen collector of P y r a m i d e l l i d a e . Odetta sulcata (de F o l i n , 1870) (figs 10,13, 54) Ondina sulcata de Folin, 1870: 214. Odetta sulcata; de Folin, 1873: p i . 29 fig. 1. Pyrgulina infrasulcata Dautzenberg, 1912: 68, p i . 3 figs 17, 18; Schänder, 1994: 45, p l . 6 f i g . e [citing Dautzenberg, 1913]. M a t e r i a l . — M a u r i t a n i a : M.031/2, M.032/1, M.043/1, M.064/1, M.065/2; V/14 (figured specimen: 2.1 x 0.8 mm), VII/26; B l / 4 , B8/1. D e p t h r a n g e . — 0-70 m . N o m e n c l a t u r e . — A s s h o w n b y v a n A a r t s e n (1984: 135), Ondina sulcata de F o l i n , 1870, s h o u l d be considered the type species of Odetta de F o l i n , 1870. T h i s species w a s Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) 15 Figs 6-13. Adelactaeon a n d Odetta species. 6, Tornatella elata v o n Koenen, 3.0 m m , Miste, Winterswijk, the Netherlands, coll. v a n Aartsen; 7, Adelactaeon puncturata (Smith), lectotype, 4.4 m m , W h y d a h , West A f r i c a , coll. B M N H 1870.1.14.11; 8, Adelactaeon elongata spec, nov., holotype ( N N M 57294), 7.6 m m , sta. M A U . 0 0 2 , M a u r i t a n i a , Baie de Cansado, 20°53'N 17°02'W, 5-vi-1988; 9, Adelactaeon lilyae spec, nov., holotype ( Z M A 3.97.006), 3.5 m m , sta. B3, M a u r i t a n i a , Banc d ' A r g u i n , 19°37,4'N 16°52.1'W, 21-34 m , 17-V-1988; 10, Odetta sulcata de Folin, lectotype (coll. de F o l i n - M N H N ) , 1.7 m m , Ile Cagnabac, Senegal; 11, Odetta dekleini spec, nov., holotype ( N N M 57297), 1.1 m m , sta. 7.007, Cape V e r d e Islands, S W of Säo Tiago, 14°54'N 23°38'W, depth 420 m , 20-viii-1986; 12, Odetta marci spec, nov., holotype ( N N M 57314), 3.8 m m , Sta. M A U . 0 7 8 , M a u r i t a n i a , off Banc d ' A r g u i n , 20°00'N 17°21'W, depth 41 m , 14-vi-1988; 13, Odetta sulcata (de Folin), 2.1 m m , Sta. R V , M a u r i t a n i a , Banc d ' A r g u i n , 19°47'30"N 17°32'30"W, 23-V-1988. 16 Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) o r i g i n a l l y described f r o m Cagnabac a n d C a p V e r g a , West A f r i c a . The t w o type specimens i n M N H N c o r r e s p o n d perfectly w e l l w i t h the s p e c i m e n f i g u r e d as Pyrgulina infrasulcatab y Schänder (1994: p l . 6 f i g . e) a n d w i t h o u r specimens. T h e lectotype of O. sulcata is designated a n d f i g u r e d here (1.7 x 0.6 m m ; f i g . 10). T h e type s p e c i m e n of Dautzenberg's P. infrasulcata, also i n M N H N , is f u l l y destroyed. Still, o n the basis of the o r i g i n a l data, w e are c o n v i n c e d that P. infrasulcata is a j u n i o r s y n o n y m of O. sulcata. Several of o u r specimens are larger t h a n the type specimens a n d some s h o w a less p r o n o u n c e d s p i r a l sculpture, b u t this clearly concerns intraspecific v a r i a t i o n o n l y . G e n u s Folinella D a l i & Bartsch, 1904 T y p e species: Amoura anguliferens de F o l i n , 1873. The generic n a m e Folinella w a s i n t r o d u c e d to replace Amoura de F o l i n , 1873, n o t G r a y , 1847, a n d , consequently, b o t h n o m i n a l taxa have the same type species, v i z . Amoura anguliferens de F o l i n , 1873. Ividella D a l i & Bartsch, 1907 [a n o m e n n o v u m for Funicularia Monterosato, 1884, not Forbes, 1845] is a j u n i o r s y n o n y m of Folinella. Rissoa excavata P h i l i p p i , 1836, is the type species of Funicularia M o n t e r o s a t o a n d thus also of Ividella. See v a n A a r t s e n (1984). Folinella holthuisi spec. n o v . (fig. 14) M a t e r i a l . — H o l o t y p e ( N N M 57316): M.087, M a u r i t a n i a , off Banc d ' A r g u i n ; 19°32'N 16°54'W; depth 65 m, sticky grey m u d w i t h shells, ophiuroids, small bivalves; v a n Veen grab; 15.vi.1988. Paratypes. M a u r i t a n i a : M.031/3 ( N N M 57317); B l / 6 ( Z M A 3.97007). D e s c r i p t i o n . — Shell slender o v o i d , strongly sculptured. C o l o u r w h i t i s h . E m b r y o n i c w h o r l s intorted, of type B. Teleoconch consisting of three to three a n d a half w h o r l s . Suture deeply channelled. G r o w t h l i n e s slightly prosocline. O n the i n i t i a l teleoconch w h o r l s , the u p p e r three quarters of the w h o r l s h o w axial ribs o n l y . These ribs are crossed b y a s p i r a l r i b at the l o w e r e n d , f o l l o w e d further d o w n b y another p r o m i n e n t s p i r a l rib, b o r d e r i n g the s u t u r a i channel. O n the l o w e r w h o r l s , the axial ribs split u p into beads w h i c h extend laterally, f o r m i n g s p i r a l ribs that w a y . There are three p l u s one s p i r a l ribs, crossing e q u a l l y strong axials w h i c h e n d at about the p e r i p h e r y ; o n the last w h o r l , o n the shell base, there are about f o u r a d d i t i o n a l , s m o o t h , s p i r a l ribs. O u t e r l i p t h i n a n d s m o o t h i n s i d e . U m b i l i c u s l a c k i n g . There is a m o d e r a t e l y strong tooth o n the c o l u m e l l a . D i m e n s i o n s : H . 2.0-2.2 m m , W . 1.0-1.2 m m ; h o l o t y p e 2.2 x 1.1 m m . D i f f e r e n t i a t i o n . — T h i s species is most s i m i l a r to Folinella ghisottii v a n A a r t s e n , 1984, b u t that species has a different s p i r a l sculpture. O n the o l d e r w h o r l s there are t w o w i d e l y spaced s p i r a l ribs, as i n Folinella excavata ( P h i l i p p i , 1836). O n the l o w e r w h o r l s , a t h i r d s p i r a l develops i n between these t w o , so that o n the p e n u l t i m a t e w h o r l there are three nearly e q u a l l y p r o m i n e n t s p i r a l ribs. W e figure a s p e c i m e n f r o m Isola d i G i g l i o (Italy) for c o m p a r i s o n (fig. 15). See also v a n A a r t s e n (1977: p i . 60 f i g . 4) and Pefias et al. (1996: p l . 29 f i g . 62) Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) 17 D e p t h r a n g e . — 53-70 m . E t y m o l o g y . — T h i s species is n a m e d after Prof. D r L . B . H o l t h u i s , i n a c k n o w l e d g e ment of his v a l u a b l e advices i n n o m e n c l a t u r a l questions . Folinellamoolenbeekispec. n o v . (figs 16, 55) M a t e r i a l . — H o l o t y p e ( Z M A 3.97.008): II, M a u r i t a n i a , Banc d ' A r g u i n ; 19°46'N 17°30'W; littoral; 23.V.1988. Paratypes ( Z M A 3.97.009-013). M a u r i t a n i a : 1/1, II/2, V / 2 , VII/4, IX/10. D e s c r i p t i o n . — Shell compact, s o m e w h a t egg-shaped w i t h rather straight sides, w i t h a p u p o i d t o p . C o l o u r w h i t e . E m b r y o n i c w h o r l s intorted, of type B to C . Teleoconch consisting of about four w h o r l s that l o o k concave because of the sculpture. Suture incised, not channelled. G r o w t h l i n e s a n d axial ribs prosocline. W i t h 15 to 20 prosocline, axial ribs w h i c h continue to the base of the shell. There are t w o w i d e l y spaced, v e r y p r o m i n e n t s p i r a l ribs, i.e. one o n each half of the w h o r l . O n the last w h o r l there are three a d d i t i o n a l spirals b e l o w the p e r i p h e r y . O u t e r l i p as thick as the b o r d e r i n g shell w a l l , f o r m i n g a c o n t i n u o u s peristome w i t h the inner l i p . U m b i l i c u s l a c k i n g . T o o t h weak, situated d e e p l y inside the aperture. D i m e n s i o n s : H . 2.0-2.5 m m , W . 1.0-1.1 m m ; h o l o t y p e 2.0 x 1.0 m m . D i f f e r e n t i a t i o n . — I n general shape a n d especially i n sculpture this species is v e r y s i m i l a r to the w e l l - k n o w n Folinella excavata ( P h i l i p p i , 1836), b u t its a p i c a l w h o r l s are m u c h m o r e compact a n d m o r e p u p o i d as c o m p a r e d to the slender, r e g u l a r l y conical t o p w h o r l s of F. excavata (compare figs 55 a n d 56). T h e axial ribs are u s u a l l y m o r e p r o socline too, b u t this is variable i nF.excavata. D e p t h r a n g e . — littoral. E t y m o l o g y . — T h i s species is n a m e d after R . G . M . M o o l e n b e e k , f r i e n d a n d c o l league of the authors a n d curator of the department of M o l l u s c a at Z M A . Folinella excavata ( P h i l i p p i , 1836) (figs 17, 56) Rissoa excavata P h i l i p p i , 1836:154, p i . 10 fig. 6. Chrysallida excavata; v a n Aartsen, 1977: 50, p i . 60 fig. 3; Penas et al., 1996:18, p l . 29 figs 60, 61. Chrysallida (Ividella) excavata; Savona & Ciccone, 1981: 41-49, p i . 49. M a t e r i a l . — O n l y one specimen, dredged at 645 m depth i n the Selvagens archipelago (St. 3.070). D i s t r i b u t i o n . — T h i s w e l l - k n o w n species is w i d e s p r e a d a l o n g the A t l a n t i c a n d M e d i t e r r a n e a n coasts of E u r o p e . It w a s reported p r e v i o u s l y f r o m the A t l a n t i c coast of M o r o c c o , f r o m Tanger to Es-Saouira ( M o g a d o r ) b y P a l l a r y (1920: 60) a n d B e l l o n - H u m b e r t (1974: 42). V a r i a t i o n . — T h e variety harveyi T h o m p s o n , 1840, as discussed a n d f i g u r e d b y Savona & C i c c o n e (1981: 49) is a f o r m w i t h m u c h m o r e n u m e r o u s axial ribs w h i c h is f o u n d every n o w a n d then together w i t h the u s u a l f o r m . W e d i d not f i n d this f o r m i n o u r material. 18 Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) G e n u s Ondina d e F o l i n , 1870 T y p e species (by subsequent d e s i g n a t i o n [ I C Z N A r t . 69 (a)(i)(l); see v a n A a r t s e n , 1984]): Ondina semiornata de F o l i n , 1872 [= Ondina warreni ( T h o m p s o n , 1845)]. The genus Ondina de F o l i n , 1870 a n d its type species have been discussed b y v a n A a r t s e n (1984: 136). T h e E u r o p e a n species have been m o n o g r a p h e d b y the same author (1987). Ondina mosti spec. n o v . (fig. 18) M a t e r i a l . — H o l o t y p e ( N N M 57318): 7.129, Cape Verde Islands, S of Säo N i c o l a u , S. Jorge Bay; 16°33'N 24°16'W; depth 405 m , m u d d y bottom; rectangular dredge; 2.ix.l986. Paratypes ( N N M 57319-57323). Cape Verde Islands: 7.049/1, 7.050/7, 7.120/1, 7.121/1, 7.129/2. D e s c r i p t i o n . — Shell elongated o v o i d w i t h a flat top. C o l o u r h y a l i n e , transparent. E m b r y o n i c w h o r l s intorted, of type C . Teleoconch consisting of about three to three a n d a half m o d e r a t e l y convex w h o r l s . Suture w e l l - m a r k e d b u t not deep. G r o w t h l i n e s nearly orthocline a n d s i n u o u s near the u p p e r suture. W i t h f o u r to five i n c i s e d spirals o n the u p p e r w h o r l s ; last w h o r l w i t h about ten a d d i t i o n a l i n c i s e d lines b e l o w the p e r i p h e r y . O u t e r l i p t h i n a n d flexuous, s m o o t h inside. U m b i l i c u s f o r m i n g a slight chink. There is a slight f o l d o n the c o l u m e l l a , as i n a l l Ondina species. D i m e n s i o n s : H . 2.0-2.4 m m , W . 1.0-1.2 m m ; h o l o t y p e 2.4 x 1.1 m m . D i f f e r e n t i a t i o n . — T h i s species belongs to the Ondina s.s. species g r o u p , w i t h a.o. O. obliqua ( A l d e r , 1844), O. warreni ( T h o m p s o n , 1845) a n d O. divisa (J. A d a m s , 1797) [= O. insculpta ( M o n t a g u , 1808)], k n o w n f r o m A t l a n t i c - M e d i t e r r a n e a n E u r o p e . These species are f i g u r e d b y v a n A a r t s e n (1987: figs 46, 49, 48) a n d W a r é n (1991: figs 34C, D , E). T h e y a l l differ f r o m O. mosti spec. n o v . b y a p r o p o r t i o n a l l y smaller aperture a n d last w h o r l , a n d a m o r e slender general shape, w i t h telescope-like u p p e r w h o r l s . T h e incised spirals r e m i n d of those i n O. divisa, b u t shells of that species are also m o r e slender conical (see Warén, 1991: f i g . 39D). D e p t h r a n g e . — 200-400 m . E t y m o l o g y . — T h i s species is n a m e d after B. v a n der M o s t , former m e m b e r of the MWH. Ondina cf. obliqua ( A l d e r , 1844) Odostomia obliqua A l d e r , 1844: 327, p i . 7 fig. 12. Ondina obliqua; v a n Aartsen, 1987: 17, 19, p i . 33 fig. 46; Warén, 1991: 104, 105, fig. 34D; Penas et al., 1996: 53, 56, fig. 143. M a t e r i a l . — Canary Islands: 4.116/1, 420 m . N o t e . — Ondina obliqua can be r e c o g n i z e d b y its tilted p r o t o c o n c h w h i c h is intorte d of type B, instead of type C as i n a l l other Ondina species. It is a w e l l - k n o w n a l l be it rare species of the A t l a n t i c - M e d i t e r a n e a n coasts of E u r o p e , n o t reported before f r o m o u r s t u d y area. Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) 19 W e f o u n d a single s p e c i m e n w h i c h has a n u m b e r of its characteristics. That shell is m u c h m o r e slender t h a n u s u a l i n O. obliqua, h o w e v e r , a n d its w h o r l s d o not increase i n size so r a p i d l y . W e therefore d o u b t its identification. Ondina warreni ( T h o m p s o n , 1845) Rissoa warreni Thompson, 1845: 315, p i . 19 fig. 4. Ondina warreni; v a n Aartsen, 1987:17,19, p i . 33 fig. 49; Warén, 1991:104,105, fig. 34E; Penas, 1996: 53, 57, 61, figs 146,147. M a t e r i a l . — M a d e i r a Archipelago: 4.K27/7, beached. N o t e . — O n l y m e n t i o n e d f r o m o u r s t u d y area b y N o r d s i e c k & Talavera (1979:186) f r o m Porto Santo, M a d e i r a archipelago. T h e shell f i g u r e d b y W a r é n (1991: f i g . 34E) is exceptionally slender a n d not representative of this w e l l - k n o w n species of A t l a n t i c E u r o p e a n coasts. Ondina dilucida (Monterosato, 1884) (fig. 19) Auriculina dilucida Monterosato, 1884: 97 . Ondina s p e c ; v a n Aartsen et al., 1984: 53, pl.125 fig.257. Ondina diaphana dilucida; v a n Aartsen, 1987:18, pl.33 fig. 54. M a t e r i a l . — Canary Islands: 2.011/1 (fig. 19), 2.085/2. N o t e . — T h e s m o o t h , transparent Ondina species, for w h i c h the subgenus Glabrondina Sacco, 1892, w a s i n t r o d u c e d , are v e r y d i f f i c u l t to d i s t i n g u i s h . There are three shells i n o u r material that w e i d e n t i f y w i t h O. dilucida w i t h some doubt. O n e of these specimens is f i g u r e d (2.4 x 1.2 m m ; f i g . 19). T h e species h a d not been reported f r o m our s t u d y area before. D e p t h range.—125-500 m . G e n u s Odostomia F l e m i n g , 1813 T y p e species (designated b y G r a y , 1847): Turbo plicatus M o n t a g u , 1803. The type species of this "classic" p y r a m i d e l l i d genus is also the type species of the n o m i n a l genus Turritodostomia Sacco, 1892. Therefore, Turritodostomia is a j u n i o r s y n o n y m of Odostomia, as already i n d i c a t e d b y B u c q u o y et a l . (1898: 767) a n d D a l i & Bartsch (1909:131). The E u r o p e a n species have been m o n o g r a p h e d b y v a n A a r t s e n (1987) a n d are also p a r t l y discussed b y Penas et al. (1996). Odostomia (Odostomia) acuta Jeffreys, 1848 (figs 20, 57) Odostomia acuta Jeffreys, 1848: 338; Warén, 1980: 37, p l . 6 figs 23, 24; v a n Aartsen et al., 1984: 51, p l . 125 fig. 246; 1987: 8,11, p l . 29 fig. 15; Pefias et al., 1996: 39, 41, figs 108,109. 20 Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) Figs 14-19. Folinella a n d Ondina species. 14, Folinella holthuisi spec, nov., holotype ( N N M 57316), 2.2 m m , sta. M A U . 0 8 7 , M a u r i t a n i a , off Banc d ' A r g u i n , 19°32'N 16°54'W, depth 65 m , 15-vi-1988; 15, Folinella ghisottii v a n Aartsen, 1984, 2.0 m m , Italy, Isola d i G i g l i o , coll. v a n Aartsen; 16, Folinella moolenbeeki spec, nov., holotype ( Z M A 3.97.008), 2.0 m m , Sta. R II, M a u r i t a n i a , Banc d ' A r g u i n , 19°46'N 17°30'W, 23-V-1988 ; 17, Folinella excavata (Philippi), 3.0 m m , Porto Polio, Corse, France, littoral, 13-vii-1978, coll. v a n Aartsen; 18, Ondina mosti spec, nov., holotype ( N N M 57318), 2.4 m m , sta. 7.129, C a p e V e r d e Islands, S of Säo N i c o l a u , S. Jorge Bay, 16°33'N 24°16'W, depth 405 m , 2-ix-1986; 19, Ondina dilucida (Monterosato), 2.4 m m , sta. 2.011, C a n a r y Islands, S of Fuerteventura, Punta de Jandia, 28°02'N 14°29'W, depth 125 m , 24-viii-1977. Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) 21 M a t e r i a l . — M a u r i t a n i a : M.002/4, M.014/1, M.031/6, M.033/1, M.044/1, M.045/2, M.065/2, M.097/7, M.099/2, M . l l l / 2 , M.113/2, M.121/1, 3.109/4, 3.133/1, 3.172/1; 1/3, I I / l , V / l , S t . VII/6; B7/6, B8/7. Canary Islands: 2.012/1, 2.021/1, 2.034/2, 2.043/2, 2.080/1, 4.041/1, 4.088/4, 4.138/2, 4.159/1. M a d e i ra Archipelago: 1.017/5, 4K27/5; Bay of Funchal, -40 m/>25 [in Z M A ] . Cape Verde Islands: 6.014/5, 6.084/1, 6.101/3, 6.103/6, 6.105/10, 7.085/1, 7.086/4, 7.088/2, 7.093/25, 7.094/3, 7.095/3, 7.102/6, 7.106/4. N o m e n c l a t u r e . — W e figure a s p e c i m e n f r o m the R i a de A r o s a (Spain) for c o m p a r i s o n (figs 20, 57). D i s t r i b u t i o n . — T h i s species w a s reported already f r o m the C a n a r y Islands b y Jeffreys (1884: 347), f r o m M a d e i r a b y W a t s o n (1897: 296) a n d f r o m the A z o r e s b y N o r d sieck & Talavera (1979:187). D e p t h r a n g e . — 0-200 m . Odostomia (Odostomia) duureni spec. n o v . (fig. 21) M a t e r i a l . — H o l o t y p e ( N N M 57324): 5.053, Azores, S of Säo M i g u e l ; 37°42'N 25°27'W; depth 50 m , coarse, m a i n l y volcanic sand; v a n Veen grab; 31.V.1981. Paratypes ( N N M 57325-57334): Azores: 5.037/2, 5.053/1, 5.077/6, 5.078/2, 5.079/2, 5.092/2, 5.102/1, 5.130/5, 5.131/2, 5.138/1. D e s c r i p t i o n . — Shell f o r m i n g a slender cone w i t h flat sides. C o l o u r r o s e - b r o w n to y e l l o w i s h b r o w n . E m b r y o n i c w h o r l s p l a n o r b i d , of type B. Teleoconch consisting of three to four flat w h o r l s , a n g l e d at the p e r i p h e r y . Suture clear, b u t not d e e p l y incised. G r o w t h l i n e s clearly prosocline; w i t h o u t a d d i t i o n a l sculpture. O u t e r l i p almost straight adapically, a b r u p t l y c u r v e d at the p e r i p h e r y a n d gently r o u n d e d basally. T h e c o l u m e l l a r part of the inner l i p is straight. U m b i l i c u s l a c k i n g . T o o t h clearly d e v e l o p e d at the a d a p i c a l side of the c o l u m e l l a . D i m e n s i o n s : H . 1.8-2.5 m m , W . 0.9-1.4 m m ; h o l o t y p e 1.8 x 0.9 m m . D i f f e r e n t i a t i o n . — W i t h its prosocline g r o w t h l i n e s , the flat a n d p e r i p h e r a l l y a n g l e d w h o r l s , a n d a straight c o l u m e l l a r b o r d e r of the aperture, the present species is s i m i l a r to Odostomia unidentata ( M o n t a g u , 1803). H o w e v e r , the latter species has a h e l i c o i d p r o t o c o n c h of type A - I I , i n d i c a t i v e of a different e m b r y o n i c d e v e l o p m e n t . D e p t h r a n g e . — 50-125 m . E t y m o l o g y . — T h i s species is n a m e d after L . v a n D u u r e n , former m e m b e r of the MWH. Odostomia (Odostomia) omphaloessa W a t s o n , 1897 (figs 22, 58) Odostomia omphaloessa Watson, 1897: 261, p i . 20 fig. 30. M a t e r i a l . — Canary Islands: 2.022/9, 2.033/5, 2.044/1, 2.073/3, 2.075/1, 4.013/1, 4.014/1, 4.048/2, 4.067/5, 4.073/5, 4.080/1, 4.088/20, 4.089/1, 4.090/10, 4.091/1, 4.092/5. Selvagens Archipelago: 3.065/2, 3.080/1, 3.081/1, 3.087/4, 3.099/2, 4.103/1. M a d e i r a : N o r t h coast littoral/17[AD28150] a n d Bay of Funchal, -40 m . />25 i n Z M A . 22 Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) N o t e s . — O r i g i n a l l y described f r o m M a d e i r a , w h e r e it has still been f o u n d recentl y i n shell grit f r o m the n o r t h coast littoral, this is one of the v e r y f e w c o l o u r e d Odostomia species. Whereas W a t s o n (1897: 261) indicates the c o l o u r as " w i t h a slight r u d d y tinge, so that the general appearance is a little h o r n y " , several of o u r specimens are rosa c o l o u r e d , others w h i t i s h . A s noted already b y W a t s o n , this species is v e r y s i m i l a r to O. acuta Jeffreys, 1848, b u t the shells are m u c h smaller a n d differ also i n the structure of the e m b r y o n i c w h o r l s , w h i c h are intorted of type B, instead of h e l i c o i d of type A II. D e p t h r a n g e . — 0-215 m . Odostomia (Odostomia) romburghi spec. n o v . (fig. 23) M a t e r i a l . — H o l o t y p e ( N N M 57335): 6.107, Cape Verde Islands, S W of Santa L u z i a ; 16°44'N 24°46'W; depth 50 m , sand; v a n Veen grab; 16.vi.1982. Paratypes ( N N M 57336-57401). Cape Verde Islands: 6.001/3, 6.003/1, 6.004/7, 6.005/8, 6.006/3, 6.007/7, 6.008/10, 6.009/6, 6.010/2, 6.014/7, 6.015/12, 6.024/>25, 6.040/6, 6.054/25, 6.056/5, 6.057/8, 6.066/1, 6.077/10, 6.082/7, 6.083/8, 6.085/3, 6.101/5, 6.103/>25, 6.105/>25, 6.107/18, 6.128/4, 6.130/13, 6.132/5, 6.134/8, 6.138/5, 6.143/6, 6.147/3, 6.149/14, 6.156/1, 6.160/3, 6.162/2, 7.028/17, 7.030/17, 7.031/6, 7.032/1, 7.037/7, 7.042/5, 7.043/12, 7.049/4, 7.050/15, 7.068/2, 7.080/5, 7.085/2, 7.088/1, 7.089/1, 7.091/5, 7.093/6, 7.094/2, 7.100/14, 7.101/5, 7.102/7, 7.109/8, 7.110/2, 7.115/13, 7.116/>25, 7.119/24, 7.120/5, 7.121/12, 7.122/3, 7.128/>25, 7.129/8, 7.141/15, 7.142/4, 7.143/4, 7.160/3. D e s c r i p t i o n . — Shell f o r m i n g a slender cone w i t h straight sides. C o l o u r bluei s h - w h i t e . E m b r y o n i c w h o r l s h e l i c o i d , of type A II. Teleoconch consisting of six to seven w h o r l s , w h i c h are almost flat. T h e p e r i p h e r y of the last w h o r l is p r o m i n e n t l y carinated. Suture clearly m a r k e d a n d accentuated b y the carina of the w h o r l above it. G r o w t h l i n e s prosocline; w i t h o n l y a (very) faint a d d i t i o n a l s p i r a l striation. O u t e r l i p straight a d a p i c a l l y , strongly angled at the p e r i p h e r a l carina a n d then gently c u r v e d to the c o l u m e l l a , w h i c h is rather straight. U s u a l l y there is a n a r r o w u m b i l i c a l chink. T o o t h clearly m a r k e d , situated rather h i g h o n the c o l u m e l l a . . D i m e n s i o n s : H . 3.2-4.0 m m , W . 1.4-1.6 m m ; h o l o t y p e 3.2 x 1.4 m m . D i f f e r e n t i a t i o n . — T h i s species is v e r y s i m i l a r to O. unidentata ( M o n t a g u , 1803), but differs i n the m u c h m o r e slender shape of the shell a n d the quite different d i m e n sions. A t the stage of six w h o r l s , a n average O. unidentata measures 5.0 m m i n height, whereas O. romburghi spec. n o v . reaches o n l y 3.0 m m . F o r the protoconchs the sizes are m o r e than 350 jum versus c. 200 jum. See also the notes o n O. unidentata v a r . elata Jeffreys, 1867, w i t h O. verhoeveni spec. n o v . D e p t h r a n g e . — 18-400 m . E t y m o l o g y . — T h i s species is n a m e d after D . v a n R o m b u r g h , former m e m b e r of the M W H . Odostomia (Odostomia) suboblonga Jeffreys, 1884 Odostomia suboblonga Jeffreys, 1884: 345, p i . 26 fig. 3; Warén, 1980: 39, p l . 6 fig. 29; v a n Aartsen, 1987: 12, p i . 30 fig. 21. Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) 23 M a t e r i a l . — Canary Islands: 4.049/4, 313 m . D i f f e r e n t i a t i o n . — T h i s is a v e r y rare species w h i c h is most s i m i l a r to O. conoidea (Brocchi, 1814). B o t h species share the general shape of the shell, w i t h a s m o o t h a n d s h i n y surface a n d orthocline g r o w t h l i n e s . H o w e v e r , O. suboblonga has a n intorted protoconch, of type B, a n d i n s i d e the outer l i p there are n o teeth. D i s t r i b u t i o n . — Odostomia suboblonga has been m e n t i o n e d f r o m off the C a p e V e r d e Islands b y Jeffreys (1884: 345), a n d f r o m Porto Santo a n d the A z o r e s b y N o r d s i e c k & Talavera (1979:188). Odostomia (Odostomia) unidentata ( M o n t a g u , 1803) (fig. 24) Turbo unidentata M o n t a g u , 1803: 324. Odostomia unidentata; v a n Aartsen, 1987: 8,11, p i . 29 fig. 17; Pefias et al., 1996: 54, 47, figs 114,115. M a t e r i a l . — M a u r i t a n i a : 3.172/2, 3.178/7; M.030/7, M.031/9, M.043/1, M.046/2, M.077/3, M.078/2, M.079/3, M.084/2, M.097/6, M . l l l / 1 , M.113/6, M.115/4, M.121/2, M.135/4; 1/5, II/6, III/l, V / 2 5 ; B7/6, B8/6. Canary Islands: 2.003/3, 2.034/2, 2.048/1, 2.103/1, 2.120/2, 4.024/2, 4.038/7, 4.041/3, 4.044/3, 4.048/4, 4.068/1, 4.088/4, 4.090/7, 4.092/3, 4.112/1, 4.137/1, 4.138/5, 4.139/6, 4.159/1; Tenerife, -98 m/12 i n Z M A . Selvagens Archipelago: 3.063/4. M a d e i r a Archipelago: 1.017/2; Bay of F u n chal, -40 m/2 i n Z M A . Cape Verde Islands: 6.004/1. D i f f e r e n t i a t i o n . — See the notes o n O. unidentata v a r . elata Jeffreys, 1867, w i t h the next species. D i s t r i b u t i o n . — A w e l l - k n o w n species, p r e v i o u s l y reported f r o m the "western coasts of A f r i c a " b y Jeffreys (1884: 346), f r o m the Canaries a n d M a d e i r a b y Jeffreys (1884: 346) a n d W a t s o n (1885: 481), a n d f r o m the A z o r e s b y D a u t z e n b e r g (1889: 59). D e p t h r a n g e . — 0-200 m . Odostomia (Odostomia) verhoeveni spec. n o v . (figs 25, 59) M a t e r i a l . — H o l o t y p e ( N N M 57402): St. M.097, M a u r i t a n i a , off Banc d ' A r g u i n ; 19°24'N 16°48'W; depth 26 m m u d d y sand w i t h some shellgravel, ophiuroids; v a n V e e n grab; 16.vi.1988. Paratypes ( N N M 57403-57405). M a u r i t a n i a : M.078/1, M.097/5, M.099/1. D e s c r i p t i o n . — Shell f o r m i n g a slender, s o m e w h a t p a g o d a - l i k e cone. C o l o u r w h i t e a n d s h i n y . E m b r y o n i c w h o r l s h e l i c o i d , of type A II. Teleoconch consisting of f o u r a n d a half to five flat w h o r l s . Suture i m p r e s s e d , nearly h o r i z o n t a l . G r o w t h l i n e s slightly opisthocline a n d s i n u o u s . There is h a r d l y a n y sculpture; a faint s p i r a l striation c a n be detected u n d e r the microscope. O u t e r l i p t h i n , s m o o t h inside. U m b i l i c u s obsolete, o n l y a slight c h i n k is sometimes present. T o o t h w e l l - d e v e l o p e d , situated v e r y h i g h o n the c o l u m e l l a . D i m e n s i o n s : H . 1.6-1.8 m m , W . 0.6-0.7 m m ; h o l o t y p e 1.7 x 0.7 m m . D i f f e r e n t i a t i o n . — T h i s species looks l i k e a m i n i a t u r e of a v e r y slender O. unidentata ( M o n t a g u , 1803). It is n o t o n l y m u c h smaller than that species, b u t it also s h o w s opisthocline g r o w t h l i n e s instead of the p r o m i n e n t prosocline ones of O. unidentata. 24 Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) Figs 20-25. Odostomia species. 20, Odostomia acuta Jeffreys, 4.5 m m , R i a de A r o s a , Spain, coll. v a n A a r t sen; 21, Odostomia duureni spec, nov., holotype ( N N M 57324), 1.8 m m , sta.5.053, Azores, S of Säo M i g u e l , 37°42'N 25°27'W, depth 50 m , 31-V-1981; 22, Odostomia omphaloessa Watson, 1.5 m m , sta. 4.088, Canary Islands, E of Lanzarote, 29°08'N 13°26'W, depth 51 m , 22-V-1980; 23, Odostomia romburghi spec, nov., holotype ( N N M 57335), 3.2 m m , sta. 6.107, C a p e V e r d e Islands, S W of Santa L u z i a , 16°44'N 24°46'W, depth 50 m , 16-vi-1982; 24, Odostomia unidentata (Montagu), 4.8 m m , L o c h Creran (Scotland), off R u b l a M o r , 56°31.2'N 05°23.5'W, 23-viii-1993, coll. v a n Aartsen; 25, Odostomia verhoeveni spec, nov., holotype ( N N M 57402), 1.7 m m , sta. M A U . 0 9 7 , M a u r i t a n i a , off Banc d ' A r g u i n , 19°24'N 16°48'W, depth 26 m , 16-vi-1988. Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) 25 O u r material also contains t w o specimens of a s o m e w h a t larger f o r m (2.2 x 1.0 m m ) w i t h clearly prosocline g r o w t h l i n e s . These t w o shells m a y b e l o n g to O. unidentata var. elata Jeffreys, 1867, w h i c h is o r i g i n a l l y described as "Spire m o r e elongated, a n d base n a r r o w e r " (Jeffreys, 1867: 134). U n f o r t u n a t e l y , a c c o r d i n g to Warén (1980: 38), there are n o type specimens k n o w n of this taxon. M o r e o v e r , Jeffreys' n a m e is preocc u p i e d b y O. elata A . A d a m s , 1860. F o r the m o m e n t b e i n g these t w o specimens cannot be i d e n t i f i e d . D e p t h r a n g e . — 26-42 m . E t y m o l o g y . — T h i s species is n a m e d after M . V e r h o e v e n , former m e m b e r of the MWH. Odostomia (Odostomia) brandhorsti spec. n o v . (fig. 26) M a t e r i a l . — H o l o t y p e ( N N M 57406): 7.119, Cape Verde Islands, S of Razo; 16°36'N 24°36'W; depth 140-160 m , coarse calcareous sand, gravel a n d stones w i t h Foraminifera; v a n V e e n grab; l.ix.1986. Paratypes ( N N M 57407-57429). Cape Verde Islands: 6.004/3, 6.005/4, 6.006/1, 6.008/2, 6.015/7, 6.024/1, 6.040/2, 6.052/1, 6.061/2, 6.068/18, 6.105/7, 6.134/5, 7.004/1, 7.005/1, 7.007/1, 7.042/2, 7.050/2, 7.080/8, 7.106/2, 7.119/11, 7.121/7, 7.122/1, 7.128/5. D e s c r i p t i o n . — Shell f o r m i n g a short cone w i t h straight sides a n d a flat top. C o l o u r white-transparent. E m b r y o n i c w h o r l s intorted, of type C . Teleoconch consisting of about four, slightly convex w h o r l s . Suture clearly m a r k e d . G r o w t h l i n e s strongly p r o socline. O n the i n i t i a l w h o r l s a f e w s p i r a l l y i n c i s e d lines are sometimes discernible; apart f r o m that the surface is s m o o t h . O u t e r l i p t h i n . O n l y a slight u m b i l i c a l c h i n k is sometimes present. There is a s m a l l b u t distinct c o l u m e l l a r tooth. D i m e n s i o n s : H . 1.4-1.5 m m , W . 0.6-0.7 m m ; h o l o t y p e : 1.4 x 0.7 m m . D i f f e r e n t i a t i o n . — T h i s rather c o m m o n species looks s o m e w h a t l i k e some forms of O. turrita H a n l e y , 1844, b u t it has a n intorted p r o t o c o n c h instead of the h e l i c o i d p r o toconch of that species. O. striolata Forbes & H a n l e y , 1850, is also s o m e w h a t similar, but m u c h larger a n d less slender. D e p t h r a n g e . — 60-400 m . E t y m o l o g y . — T h i s species is n a m e d after A . L . Brandhorst, former m e m b e r of the MWH. Odostomia (Odostomia) dijkhuizeni spec. n o v . (fig. 27) M a t e r i a l . — H o l o t y p e ( N N M 57430): M.065, M a u r i t a n i a , off Banc d ' A r g u i n ; 20°00'N 17°11'W; depth 25 m , m u d d y sand w i t h Cardium a n d Lanice; v a n V e e n grab; 13.vi. 1988. Paratypes ( N N M 57431- 57445; Z M A 3.97.027-3.97.032). M a u r i t a n i a : 3.133/1, M.011/2, M.013/1, M.030/3, M.031/3, M.033/1, M.043/3, M.044/1, M.045/2, M.046/1, M.065/3, M.078/1, M.079/1, M.084/1, M.085/3; 1/12, II/4, III/9, V/>25, VII/24; Bl/>25, B8/4. D e s c r i p t i o n . — Shell f o r m i n g a slender cone w i t h straight sides. C o l o u r w h i t e , s h i n y . E m b r y o n i c w h o r l s intorted, of type C . Teleoconch of f o u r to five almost flat w h o r l s . Suture clearly incised b u t not at a l l channelled. G r o w t h l i n e s orthocline to 26 Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) v e r y slight prosocline. T h e surface is s m o o t h a n d s h i n y . O u t e r l i p t h i n , straight over most of its length. O n l y a slight u m b i l i c a l c h i n k c a n sometimes be detected. T o o t h conspicuous, situated at the u p p e r part of the c o l u m e l l a . D i m e n s i o n s : H . 2.3-2.5 m m , W . 1.0-1.2 m m ; h o l o t y p e : 2.3 x 1.0 m m . D i f f e r e n t i a t i o n . — T h i s species is s i m i l a r to O. turriculata Monterosato, 1869, b u t it has nearly orthocline g r o w t h l i n e s , whereas the last w h o r l is relatively m u c h larger. There is a certain resemblance to O. paardekooperi spec. n o v . too, b u t shells of that species are m o r e slender a n d have a relatively shorter last w h o r l , whereas the g r o w t h lines are clearly prosocline. D e p t h r a n g e . — 0-75 m . E t y m o l o g y . — T h i s species is n a m e d after T. D i j k h u i z e n , former m e m b e r of the MWH. Odostomia (Odostomia) paardekooperi spec. n o v . (fig. 28) M a t e r i a l . — H o l o t y p e ( N N M 57446): 6.101, Cape Verde Islands, S W of Santa L u z i a ; 16°45'N 24°46'W; depth 20 m , coarse sand; v a n Veen grab; 16.vi.1982. Paratypes ( N N M 57447-57489). Cape Verde Islands: 6.004/3, 6.005/1, 6.007/3, 6.009/2, 6.015/3, 6.019/1, 6.024/2, 6.056/3, 6.061/10, 6.066/7, 6.068/2, 6.077/5, 6.082/7, 6.083/5, 6.101/20, 6.103/22, 6.105/13, 6.107/4, 6.130/7, 6.134/4, 6.147/2, 6.156/5, 6.158/4, 6.159/13, 6.160/6, 6.164/5, 6.171/3, 7.032/1, 7.049/2, 7.050/6, 7.093/6, 7.100/1, 7.106/3, 7.115/1, 7.116/3, 7.119/24, 7.120/4, 7.121/5, 7.128/7, 7.141/22, 7.142/9, 7.143/5. D e s c r i p t i o n . — Shell f o r m i n g a slender cone w i t h straight sides. C o l o u r w h i t i s h . E m b r y o n i c w h o r l s intorted, of type C . Teleoconch of about six flat w h o r l s , s o m e w h a t a n g l e d at the p e r i p h e r y of the last w h o r l . Suture clearly i m p r e s s e d . G r o w t h l i n e s m a r k e d l y prosocline. N o sculpture, except for the g r o w t h l i n e s . O u t e r l i p straight adapically, strongly c u r v e d at the p e r i p h e r a l angle a n d evenly r o u n d e d abapically. U m b i l i c u s s m a l l b u t distinct. T o o t h v e r y p r o m i n e n t . D i m e n s i o n s : H . 2.7-3.1 m m , W . 1.1-1.3 m m ; h o l o t y p e : 2.8 x 1.1 m m . D i f f e r e n t i a t i o n . — T h i s fairly c o m m o n species is most s i m i l a r to O. turriculata Monterosato, 1869, as described a n d f i g u r e d b y v a n A a r t s e n (1987: 5, 28, f i g . 3). Shells of that species, h o w e v e r , have flexuous a n d opisthocline g r o w t h l i n e s a n d s i m i l a r b u t larger e m b r y o n i c w h o r l s . D e p t h r a n g e . — 20-400 m . E t y m o l o g y . — T h i s species is n a m e d after L . S. Paardekooper, former m e m b e r of the M W H . Odostomia (Odostomia) schrami spec. n o v . (figs 29, 60) M a t e r i a l . — H o l o t y p e ( N N M 57490): 3.172, off M a u r i t a n i a ; 20°21'N 17°17'W; depth 34 m , sand w i t h shell gravel; triangular dredge; l.xi.1978. Paratypes ( N N M 57491-57502; Z M A 3.97.033). M a u r i t a n i a : 3.110/2, 3.133/1, 3.172/37, 3.178/2, M.097/1, M.106/5, M.109/2, M . l l l / 4 , M.113/6, M.115/7, M.121/5, M.122/2; B4/4. Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) 27 D e s c r i p t i o n . — Shell slender o v a l , w i t h the last w h o r l o c c u p y i n g nearly t w o thirds of the total length. C o l o u r w h i t e to transparent, glossy. E m b r y o n i c w h o r l s intorted, of type B. Teleoconch consisting of four a n d a half to five a n d a half w h o r l s , w h i c h are o n l y slightly convex. Suture clearly incised a n d m a r k e d l y i n c l i n e d . G r o w t h l i n e s orthocline, b e c o m i n g flexuous or sinuous o n the l o w e r w h o r l s . T h e surface is s m o o t h a n d glossy. O u t e r l i p sharp, s m o o t h inside. M o u t h n a r r o w l y o v a l , o c c u p y i n g just over one t h i r d of the total height. U m b i l i c u s l a c k i n g . T o o t h conspicuous, p l a c e d h i g h o n the c o l u m e l l a . D i m e n s i o n s : H . 2.9-3.0 m m , W . 0.8-1.1 m m ; h o l o t y p e : 2.9 x 0.9 m m . D i f f e r e n t i a t i o n . — O. schrami spec. n o v . is s i m i l a r to O. kromi v a n A a r t s e n et al., 1984. C o m p a r e d w i t h that species, it is larger, w i t h a relatively higher b o d y - w h o r l a n d a larger aperture. T h e t o p w h o r l s are s i m i l a r (in b o t h species type B), b u t the g r o w t h l i n e s are m o r e orthocline a n d become m o r e or less flexuous o n the l o w e r w h o r l s , i n contrast to the straight, s o m e w h a t prosocline g r o w t h l i n e s i n O. kromi (see also v a n A a r t s e n et a l , 1984: 52, p l . 124 f i g . 249). D e p t h r a n g e . — 15-75 m . E t y m o l o g y . — This species is n a m e d after A . A . W . Schram, former m e m b e r of the MWH. Odostomia (Odostomia) striolata Forbes & H a n l e y , 1850 Odostomia striolata Forbes & Hanley, 1850: 267, p i . 95 fig. 5; v a n Aartsen et al., 1984: 51, pl.124 fig. 247; 1987: 9,13, p i . 30 fig. 25; Penas et al., 1996: 49, 52, figs 127,128. M a t e r i a l . — Canary Islands: 2.003/3, 2.010/1, 2.032/2, 2.033/13, 2.043/3, 2.044/1, 2.075/2, 2.147/3; 4.001/3, 4.013/3 4.014/19, 4.016/8, 4.020/>25, 4.022/>25, 4.024/11, 4.027/19, 4.029/10, 4.034/1, 4.038/7, 4.088/4, 4.090/2, 4.110/7, 4.112/21; Fuertaventura/>25, Tenerife, -100m/>25, E l Hierro/2, Isla de L a Palma/>25 [in Z M A ] . Selvagens Archipelago: 3.062/4, 3.063/10, 3.099/9, 4.103/1. M a d e i r a , Bay of Funchal, -6-20 m/>25 [in Z M A ] . Azores: A Z O 020/1, A Z O 021/4, A Z O 022/2; 5.006/7, 5.008/6, 5.017/8, 5.036/2, 5.050/8, 5.051/9, 5.054/6, 5.077/1, 5.078/16, 5.079/>25, 5.080/4, 5.083/6, 5.084/8, 5.092/7, 5.093/8, 5.094/4, 5.102/2, 5.112/16, 5.113/7, 5,116/2, 5.119/4, 5.121/16, 5.126/4, 5.128/6, 5.129/1, 5.132/>25, 5.135/17, 5.138/15, 5.139/6, 5.140/5, 5.146/3, 5.150/6, 5.159/5, 5.162/1, 5.176/5; Säo M i g u e l , Ponta Delgada, -8 m/5, Sâo M i g u e l , Praia do Populo, beach/10; Säo M i g u e l , Ponta Delgada, reef edge/3, Pico, lagoon/5, Graciosa, beach/6, Terceira, beach/11 [in Z M A ] . Cape Verde Islands: 6.093/3, 7.028/12, 7.030/3, 7.031/2, 7.037/2, 7.038/6, 7.100/2, 7.110/1, 7.115/2, 7.119/8, 7.120/1, 7.121/3, 7.129/3. D i s t r i b u t i o n . — O. striolata has o n l y been reported once f r o m Tenerife, C a n a r y Islands ( N o r d s i e c k & Talavera, 1979:189). F r o m the present s t u d y the species appears to be m u c h m o r e w i d e l y d i s t r i b u t e d . D e p t h r a n g e . — 0-400 m . Odostomia (Odostomia) turrita H a n l e y , 1844 Odostomia turrita Hanley, 1844: 18; v a n Aartsen et al., 1984: 51, p l . 124 fig. 248; 1987: 8,12, p l . 29 fig. 18; Penas et al., 1996: 47, 54, figs 116,117. M a t e r i a l . — Mauritania: M.011/2, M.013/2, M.032/2, M.033/1, M.084/2, M.106/13, M.118/12, M.121/4; 3.172/>25, 3.174/2, 3.178/17, 3.179/9. Canary Islands: 2.033/2, 2.043/1, 2.064/3, 2.075/4, 28 Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) 4.013/15, 4.014/21, 4.016/>25, 4.020/16, 4.022/>25, 4.024/>25, 4.027/19, 4.028/2, 4.029/11, 4.034/1, 4.036/>25, 4.038/25, 4.041/8, 4.048/5, 4.067/7, 4.073/2, 4.088/19, 4.090/4, 4.091/1, 4.110/2, 4.112/5; Fuertaventura/8, Tenerife/>25 [in Z M A ] . M a d e i r a , B a y of Funchal, -2 to - 4 0 m./>25 [in Z M A ] . Azores: 5.006/>25, 5.008/2, 5.009/2, 5.017/3, 5.026/6, 5.033/12, 5.036/7, 5.037/8, 5.039/4, 5.040/>25, 5.047/12, 5.050/19, 5.051/10, 5.064/6, 5.077/>25, 5.078/>25, 5.079/>25, 5.080/7, 5.083/5, 5.084/11, 5.091/>25, 5.093/24, 5.094/5, 5.096/19, 5.102/6, 5.111/4, 5.112/>25, 5.113/12, 5.116/5, 5.117/6, 5.118/3, 5.119/20, 5.121/13, 5.122/2, 5.126/1, 5.128/6, 5.129/5 5.131/5, 5.132/6, 5.133/2, 5.135/4, 5.136/4, 5.138/7, 5.139/16, 5.140/17, 5.144/>25, 5.148/5, 5.150/6, 5.158/>25, 5.159/8, 5.162/14, 5.164/12, 5.190/6, 5K01/7; A Z O 021/3, A Z O 022/>25; Säo M i g u e l , Ponta Delgada -8 m/1, Säo M i g u e l , Ponta Delgada, reef edge/25; Säo M i g u e l , Säo Roque/10, Säo M i g u e l , Praia de Popula, beach/15, Pico, lagoon/7, Graciosa, beach/5 [in Z M A ] . D i s t r i b u t i o n . — R e p o r t e d a l r e a d y f r o m the C a n a r y Islands b y Jeffreys (1884: 346), W a t s o n (1885: 481) a n d N o r d s i e c k & T a l a v e r a (1979: 188), f r o m M a d e i r a b y W a t s o n (1897: 298) a n d f r o m M o r o c c o (Casablanca a n d Rabat) b y P a l l a r y (1920: 60) a n d B e l l o n - H u m b e r t (1974: 45). F r o m o u r w o r k this species appears to be w i d e l y d i s t r i b u t e d t h r o u g h o u t o u r s t u d y area except f o r the C a p e V e r d e Islands. D e p t h r a n g e . — 0-250 m . Odostomia (Odostomia) angusta Jeffreys, 1867 Odostomia pallida var. angusta Jeffreys, 1867:125. Odostomia angusta; Warén, 1980: 37, p l . 6 fig. 18 [not fig. 22 (Warén, i n litt.)]; v a n Aartsen, 1987: 9, 13, p i . 30 fig. 23; Penas et al., 1996: 40, 49, figs 123,124. M a t e r i a l . — Mauritania: M.106/>25, M.109/5, M . l l l / 7 , M.115/2, M.118/>25, M.121/4, 3.172/24, 3.174/>25, 3.178/>25, 3.179/21, 3.181/17; B6/>25. Canary Islands: 4.022/2, 4.088/1, 4.090/1. Cape Verde Islands: 6.105/1, 6.149/1, 7.115/1, 7.142/1. D i s t r i b u t i o n . — T h i s species h a d n o t b e e n m e n t i o n e d before f r o m W e s t A f r i c a o r the n e i g h b o u r i n g i s l a n d s D e p t h r a n g e . — 13-80 m . Odostomia (Odostomia) boermani spec. n o v . (figs 30, 61) M a t e r i a l . — H o l o t y p e ( N N M 57503): M.033, off M a u r i t a n i a ; 18°47'N 16°34'W; depth 114 m , m u d d y sand a n d fine shell gravel; v a n V e e n grab; 9.vi.l988. Paratypes ( N N M 57504-57508; Z M A 3.97.034). Mauritania: 3.133/1, M.030/2, M.032/5, M.033/2, M.046/1; B5/1). D e s c r i p t i o n . — Shell strong, forming a slender, somewhat pagoda-like cone. C o l o u r m i l k y - w h i t e a n d shiny. E m b r y o n i c w h o r l s intorted, type C . Teleoconch cons i s t i n g of a b o u t f i v e w h o r l s w h i c h are o n l y s l i g h t l y c u r v e d i n their u p p e r t w o - t h i r d s a n d m a r k e d l y m o r e c o n v e x i n the l o w e r t h i r d . S u t u r e w e l l - m a r k e d . O n l y g r o w t h l i n e s c a n be detected, w h i c h are about o r t h o c l i n e . O u t e r l i p t h i n , s m o o t h i n s i d e . U m b i l i c u s a s m a l l c h i n k . T h e r e is a m o d e r a t e l y p r o m i n e n t t o o t h o n the c o l u m e l l a . D i m e n s i o n s : H . 2.5-2.7 m m , W . 1.0-1.3 m m ; h o l o t y p e : 2.7 x 1.3 m m . D i f f e r e n t i a t i o n . — O. boermani spec. n o v . is s i m i l a r to O. lukisii Jeffreys, 1859, as far Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) 29 as its s h i n y , m i l k y - w h i t e w h o r l s a n d the flat, intorted top w h o r l s are concerned. H o w ever, the shape of the w h o r l s is quite different a n d the shells are clearly larger than those of O. lukisii. D e p t h r a n g e . — 25-114 m . E t y m o l o g y . — T h i s species is n a m e d after D . J . B o e r m a n , former m e m b e r of the MWH. Odostomia (Odostomia) carrozzai v a n A a r t s e n , 1987 Odostomia carrozzai v a n Aartsen, 1987: 10, 13, p i . 31 fig. 30 [nomen n o v u m for Odostomia albella auct., n o n Lovén, 1846]. M a t e r i a l . — Canary Islands: 4.073/1, 4.088/4, 4.091/1, 4.092/1. Selvagens Archipelago: 3.070 (fragments). D i s t r i b u t i o n . — T h i s species is cited f r o m M a d e i r a b y Jeffreys (1884: 348) o n the authority of W a t s o n , w h o cited this species too (Watson, 1897: 296) as v e r y abundant at M a d e i r a . W e f o u n d o n l y a f e w specimens, none of w h i c h f r o m M a d e i r a . D e p t h r a n g e . — 48-92 m . Odostomia (Odostomia) eulimoides H a n l e y , 1844 Odostomia eulimoides H a n l e y , 1844: 18; v a n Aartsen et al., 1984: 53, p l . 125 fig. 252; 1987: 10, 14, p l . 31 figs 31, 32; Penas et al., 1996: 44, 51, figs 132,133. M a t e r i a l . — M a u r i t a n i a : M.078/5, M.084/1; B7/1, B8/1. Azores: 5.051/12; Säo M i g u e l , Ponta Delgada,-8 m/1, Säo M i g u e l , Säo Roque/1, Graciosa, beach/2, Terceira, beach/>25 [in Z M A ] . D i s t r i b u t i o n . — Odostomia eulimoides has been reported f r o m Tanger B a y ( M o r o c co) b y Jeffreys (1884: 348 'Odostomia pallida') as its most southern locality. I n o u r mater i a l there are six specimens f r o m the West A f r i c a n coast a n d m a n y f r o m the A z o r e s , w h i c h extends the area of this species to the Southwest. D e p t h r a n g e . — 0-41 m . Odostomia (Odostomia)kuiperispec. n o v . (fig. 31) M a t e r i a l . — H o l o t y p e ( Z M A 3.97.014): 6, Azores, N E of Terceiro, Säo Sebastiao, O s Salgueiros, 29.iii.1989 (leg. T. Piersma). Paratypes ( N N M 57509-57511, 58051; Z M A 3.97.015-016). Azores: 5.130/2, 5.138/5, 5.176/4, A Z O . 2 2 / 1; 6, N E of Terceira, Säo Sebastiäo/21, St. 9 Pico, lagoon, E . of Lajes d o Pico/4. D e s c r i p t i o n . — Shell slender egg-shaped, rather s m a l l . C o l o u r w h i t e . E m b r y o n i c w h o r l s intorted, of type B. Teleoconch of three a n d a half to four, moderately convex w h o r l s ; the last w h o r l measures m o r e than half the total height of the shell. Suture clearly d e f i n e d . G r o w t h l i n e s s l i g h t l y prosocline, w i t h a tendency to become orthocline o n the l o w e r w h o r l s ; n o further sculpture. O u t e r l i p t h i n , s m o o t h inside. O n l y a 30 Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) Figs 26-31. Odostomia species. 26, Odostomia brandhorsti spec, nov., 1.4 m m , holotype ( N N M 57406), sta. 7.119, Cape V e r d e Islands, S of Razo, 16°36'N 24°36'W, depth 140-160 m , l-ix-1986; 27, Odostomia dijkhuizeni spec, nov., holotype ( N N M 57430), 2.3 m m , sta. M A U . 0 6 5 , Mauritania, off Banc d ' A r g u i n , 20°00'N 17°11'W, depth 25 m , 13-vi-1988; 28, Odostomia paardekooperi spec, nov., holotype ( N N M 57446), 2.8 m m , sta. 6.101, Cape V e r d e Islands, S W of Santa L u z i a , 16°45'N 24°46'W, depth 20 m , 16-vi-1982; 29, Odostomia schrami spec, nov., holotype ( N N M 57490), 1.9 m m , sta. 3.172, off Mauritania, 20°21'N 17°17'W, depth 34 m , l-xi-1978; 30, Odostomia boermani spec, nov., holotype ( N N M 57503), 2.7 m m , sta. M A U . 0 3 3 , off Mauritania, 18°47'N 16°34'W, depth 114 m , 9-vi-1988; 31, Odostomia kuiperi spec, nov., holotype ( Z M A 3.97.014), 1.4 m m , sta. 6, Azores, Pico, N E of Terceira, Säo Sebastio, 29.iii.1989. Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) 31 slight u m b i l i c a l chink. T o o t h s m a l l b u t c o n s p i c u o u s . D i m e n s i o n s : H . 1.4-1.8 m m , W . 0.7-0.9; h o l o t y p e : 1.4 x 0.8 m m . D i f f e r e n t i a t i o n . — T h i s species l o o k s like a slender m i n i a t u r e of O. angusta Jeffreys, 1867, b u t w i t h the same n u m b e r of w h o r l s it measures about half the size of that species. D e p t h r a n g e . — 0-142 m . E t y m o l o g y . — T h i s species is n a m e d after J.G.J. K u i p e r , f o r m e r m e m b e r of the MWH. Odostomia (Odostomia) lukisii Jeffreys, 1859 Odostomia lukisii Jeffreys, 1859: 112, p i . 3 figs 19a, b; Warén, 1980: 38, p l . 6 figs 25, 26; v a n Aartsen et al., 1984: 53, p l . 124 fig. 253; 1987: 8,12, p l . 30 fig. 19; Penas et al., 1996: 46, 53, figs 138,139. M a t e r i a l . — Mauritania: 3.172/2; B l / 1 , B2/2. Canary Islands: 2.032/1, 2.033/4, 2.043/1, 4.013/2, 4.014/2, 4.038/1, 4.073/4, 3K08/1, 4K06/1; G r a n Canaria/1, Tenerife/42, E l Hierro/>25, Isla de L a Palma/10, Fuertaventura/21, Gomera/2 [in Z M A ] . M a d e i r a Archipelago: 4K27/1; Bay of Funchal/18 [in Z M A ] . A z o r e s : 5.006/1, 5.010/1, 5.036/2, 5.037/3, 5.039/3, 5.040/2, 5.082/1, 5.102/1, 5.158/1, A Z O 005/2, A Z O 020/1, A Z O 033/4, 5K01/1; Säo M i g u e l , Ponta Delgada, -8 m/4, Pico, lagoon/13, Terceira, beach/9 [in Z M A ] . D i s t r i b u t i o n . — T h i s species has not been m e n t i o n e d f r o m o u r s t u d y area before. D e p t h r a n g e . — 0-150 m . Odostomia (Odostomia) scalaris M a c g i l l i v r a y , 1843 Odostomia scalaris 1843 M a c g i l l i v r a y , 1843: 154; v a n Aartsen et al., 1984: 53, p l . 125 fig. 254; 1987: 9, 12, p l . 30 fig. 22; Penas et al., 1996: 51, 52, fig. 136 (not 137). M a t e r i a l . — Mauritania: M.033/1, M.078/1; 3.172/>25. D i s t r i b u t i o n . — T h i s w i d e l y d i s t r i b u t e d species has been reported before f r o m "off the W e s t of Africa*' b y Jeffreys (1884: 348), f r o m M a d e i r a b y W a t s o n (1897: 298), f r o m the Selvagens archipelago b y N o r d s i e c k & T a l a v e r a (1979: 187) a n d f r o m the A z o r e s b y D a u t z e n b e r g (1889: 59). N o m e n c l a t u r e . — M a n y authors use the n a m e O. rissoides H a n l e y , 1844, for this species, b u t w e f o l l o w v a n A a r t s e n (1987a: 149, 150) i n the use of the senior n a m e O. scalaris. D e p t h r a n g e . — 34-114 m . Odostomia (Odostomia) verduini v a n A a r t s e n , 1987 Odostomia verduini v a n Aartsen, 1987: 5, 9,13, p i . 30 fig. 24; Penas et al., 1996: 49, 54, figs 125,126. M a t e r i a l . — Canary Islands: 4.069/2, 4.092/1; Mauritania: M.033/1, M.064/2, M.118/3; 3.172/3, 3.174/2, 3.178/5, 3.181/11; Cape Verde Islands: 6.059/1, 6.064/1, 7.068/1. D i f f e r e n t i a t i o n . — Shells of this species h a v e flat w h o r l s w i t h a n a d a p i c a l s p i r a l 32 Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) groove, a p r o m i n e n t c o l u m e l l a r tooth, a n d prosocline g r o w t h l i n e s . A m o n g o u r material w e f o u n d t w o specimens [M.065 a n d 3.172] w h i c h corres p o n d i n nearly a l l details w i t h O. verduini, except for their h e l i c o i d p r o t o c o n c h of type A II instead of the intorted type B of O. verduini. A s there are o n l y t w o specimens available for s t u d y , w e refrain f r o m d e s c r i b i n g these specimens as representing a species n e w to science. D e p t h r a n g e . — 12-100 m . Odostomia (Odostomia) meijeri spec. n o v . (fig. 32) M a t e r i a l . — H o l o t y p e ( N N M 57512): M.043, off M a u r i t a n i a ; 19°04'N 16°25'W; depth 15 m , greyish sand w i t h some shell gravel; v a n Veen grab; l l . v i . 1 9 8 8 . Paratypes ( N N M 57513-57519; Z M A 3.97.035-3.97.042): M a u r i t a n i a : 3.109/9, 3.128/1, 3.172/7, M.013/3, M . 0 4 3 / l l , M.044/6, M . l l l / 2 ; 1/11,11/16, III/8, St. V / 1 , VII/9, IX/4, B4/3, B7/3. A f e w specimens are not considered paratypes: M a u r i t a n i a : M.011/1, M.030/2, M.113/1; 3. 172/1. These shells are larger, v i z . u p to t w o m m h i g h at the same w i d t h , a n d thus m o r e slender too. The g r o w t h l i n e s become flexuous near the aperture o n the last w h o r l a n d the tooth seems to be s o m e w h a t m o r e p r o m i n e n t . Still w e consider it possible that these specimens b e l o n g to the same species a n d give a p h o t o g r a p h (fig. 33) for future recognition. D e s c r i p t i o n . — Shell compact, short o v o i d - c o n i c a l , w i t h an obtuse top. C o l o u r w h i t e to hyalin-transparent. E m b r y o n i c w h o r l s intorted, of type C . Teleoconch consisti n g of three to three a n d a half slightly convex w h o r l s , w h i c h are somewhat s h o u l dered. Suture conspicuous but not v e r y deep. G r o w t h l i n e s straight a n d nearly orthocline. W i t h a f e w s p i r a l l y incised lines a r o u n d the periphery, not a l w a y s easily discernible. O u t e r l i p thin, evenly r o u n d e d a n d s m o o t h inside. O n l y a slight u m b i l i c a l chink. T o o t h v e r y s m a l l , situated deep w i t h i n the aperture a n d h a r d l y visible therefore. D i m e n s i o n s : H . 1.5-1.6 m m , W . 0.8-1.0 m m ; h o l o t y p e : 1.6 x 0.9 m m . D i f f e r e n t i a t i o n . — Because of the s p i r a l l y i n c i s e d lines a r o u n d the p e r i p h e r y , this s m a l l species is s i m i l a r to O. nivosa ( M o n t a g u , 1803) as f i g u r e d b y v a n A a r t s e n et al. (1984: 53, p l . 123 f i g . 255). H o w e v e r , that species has m u c h m o r e slender shells, w i t h a sculpture consisting of a f e w s p i r a l ribs. A c c o r d i n g to the o r i g i n a l description, Ondina hemisculpta de F o l i n , 1887, does not c o r r e s p o n d w i t h O. meijeri spec, n o v . , but the figure is s o m e w h a t s i m i l a r . A c c o r d i n g to K i s c h (1959:108) the type s p e c i m e n is lost. D e p t h r a n g e . — 0-34 m . E t y m o l o g y . — T h i s species is n a m e d after A . W . J . Meijer, former m e m b e r of the MWH. Odostomia (Odostomia) truncatula Jeffreys, 1850 (fig. 34) Odostomia truncatula Jeffreys, 1850: 109; Warén, 1980: 39, p l . 6 fig. 21; Fretter et al., 1986: 595-597, figs 409- 411. 33 Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) M a t e r i a l . — Cape Verde Islands: 7.102/1, 7.119/1, 7.121/1 (fig. 34). D i s t r i b u t i o n . — Three specimens of a n Odostomia species f r o m the C a p e V e r d e Islands are so s i m i l a r to O. truncatula (type locality: P l y m o u t h ) that w e i d e n t i f y t h e m w i t h that rare species, a l t h o u g h it is not k n o w n f r o m the C a n a r y Islands, M a d e i r a or the A z o r e s . There are also n o p u b l i s h e d records f r o m M o r o c c o or the m o r e southern West A f r i c a n coasts. D e p t h r a n g e . — 140-200 m . Odostomia (Odostomia) vanurki spec. n o v . (fig. 35) M a t e r i a l . — H o l o t y p e ( N N M 57520): St. 7.065, Cape Verde Islands, W of Boa Vista, S W of Ilhéu Calheta do Velho; 16°10'N 22°58'W; depth 39 m , very fine, y e l l o w calcareous sand; v a n V e e n grab; 27.viii.1986. Paratypes ( N N M 57521-57523): Cape Verde Islands: 6.059/2, 7.042/1, 7.065/5. D e s c r i p t i o n . — Shell rather slender, nearly c y l i n d r i c a l w i t h a n obtuse t o p . C o l o u r w h i t i s h to transparent. E m b r y o n i c w h o r l s intorted. Teleoconch consisting of three a n d half, evenly r o u n d e d w h o r l s . Suture w e l l - m a r k e d . G r o w t h l i n e s s l i g h t l y prosocline, not v e r y p r o n o u n c e d ; apart f r o m these, the shell surface is s m o o t h a n d s h i n y . O u t e r l i p t h i n . U m b i l i c u s l a c k i n g . There is a w e a k tooth o n the c o l u m e l l a , situated deeply inside the aperture. D i m e n s i o n s : H . 1.7 m m , W . 0.6 m m ; h o l o t y p e : 1.7 x 0.6 m m . D i f f e r e n t i a t i o n . — Odostomia vanurki spec. n o v . differs f r o m O. nivosa ( M o n t a g u , 1803) a n d O. truncatula Jeffreys, 1850, as f i g u r e d b y v a n A a r t s e n (1987: p i . 28 figs 8 a n d 9, respectively) b y the m o r e slender general shape of the shell a n d its smooth, s h i n y surface w i t h o u t a s p i r a l sculpture. Odostomia vanurki is also v e r y s i m i l a r to LiostomiawareniSchänder, 1994, b u t it c a n be d i s t i n g u i s h e d f r o m that species b y the w e a k tooth o n the c o l u m e l l a , the sutures w h i c h are less steep, the relatively larger aperture a n d the m o r e convex w h o r l s . D e p t h range — 39-76 m . E t y m o l o g y . — T h i s species is n a m e d after R . M . v a n U r k , former m e m b e r of the MWH. Odostomia(Megastomia)conoidea (Brocchi, 1814) (fig. 36) Turbo conoideus Brocchi, 1814: 660, pl.16 fig. 2; Pinna & Spezia, 1978:162, p i . 53 fig. 4 [type]. Odostomia conoidea; v a n Aartsen, 1987: 7,10, p i . 29 fig. 12; Penas et al., 1996: 42, 45, figs 106,107. M a t e r i a l . — M a u r i t a n i a : M.011/9, M.013/4, M.014/2, M.030/12, M.031/>25, M.032/18, M.033/14, M.034/8, M.035/3, M.046/6, M.064/5, M.065/15, M.077/>25, M.078/12, M.079/17, M.084/7, M.087/1, M.097/7, M.099/14, M.106/1, M . l l l / 2 , M.113/9, M.118/2, M.121/5, M.135/>25; 3.109/13, 3.113/25, 3.123/3, 3.128/4, 3.133/>25, 3.159/1, 3.172/6, 3.192/3, 3.194/>25. Canary Islands: 2.010/1, 2.075/2, 4.044/2. Cape Verde Islands: 6.010/2, 6.057/4, 6.059/10, 6.061/4, 6.066/2, 6.103/1, 6.107/1, 6.128/2, 6.130/12, 6.156/1, 6.160/2, 6.164/2, 7.042/7, 7.065/4, 7.066/5,7.068/3, 7.079/>25, 7.080/25, 7097/1, 7.143/1, 7.160/3, 7.161/3. 34 Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) N o m e n c l a t u r e . — T h i s rather c o m m o n species s h o w s a b r o a d intraspecific v a r i a tion. W e therefore consider O. boteroi Schänder, 1994, O. corimbensis Schander, 1994, O. gilsoni D a u t z e n b e r g , 1912, a n d O. sulcifera S m i t h , 1872, conspecific. W e have selecte d the s p e c i m e n m a r k e d " F i g ' d spec." i n sample B M N H 1870.1.1229 as lectotype of O. sulcifera, described f r o m W h y d a h (West A f r i c a ) (4.6 x 1.9 m m ; f i g . 36). D i s t r i b u t i o n . — T h i s w i d e l y d i s t r i b u t e d species has been reported f r o m M a d e i r a b y W a t s o n (1897: 297), f r o m M o r o c c o (Casablanca, M o g a d o r a n d Rabat) b y P a l l a r y (1920: 59) a n d B e l l o n - H u m b e r t (1974: 44), a n d f r o m the C a n a r y Islands b y N o r d s i e c k & Talavera (1979:186). D e p t h r a n g e . — 15-150 m . Odostomia (Megastomia) conoidea (Brocchi, 1814) var. (fig. 37) M a t e r i a l . — M a u r i t a n i a : M.029/1, M.033/7, M.064/3, M.097/11 (fig. 37), 3.159/1 and Banc d ' A r g u i n , St. B8/>25 i n Z M A . N o t e s . — The present specimens are m u c h m o r e slender than any w e have seen f r o m A t l a n t i c - M e d i t e r r a n e a n o r i g i n . The length is 2.7-3.1 m m a n d the w i d t h is n o m o r e than 1.1-1.4 m m . Still they s h o w the same p r o t o c o n c h as Odostomia (Megastomia) conoidea, the same orthocline g r o w t h l i n e s a n d the same v e r y s m o o t h a n d s h i n y surface. A w e a k carinal s p i r a l rib is also present. K n o w i n g the great v a r i a b i l i t y of the P y r a m i d e l l i d a e , w e refrain f r o m d e s c r i b i n g these specimens as a n e w species b u t p o i n t out their different shape (2.7 x 1.1 m m ; f i g . 37). Odostomia (Megastomia) conspicua A l d e r , 1850 Odostomia conspicua A l d e r , 1850: 359; v a n Aartsen, 1987: 7, 8, p i . 28 figs 10,11. M a t e r i a l . — Canary Islands: 4.013/2, 4.014/4, 4.038/5, 4.041/1, 4.088/7, 4.099/1, 4.137/1, 4.138/4, 4D02/3; Tenerife, -98 m/15 [in Z M A ] . M a d e i r a Archipelago: 3.002/1; Bay of Funchal, -35 m/8 [in ZMA]. N o t e s . — Odostomia conspicua, one of the largest E u r o p e a n Odostomia species, has already been m e n t i o n e d f r o m M a d e i r a b y W a t s o n (1897: 297) a n d N o r d s i e c k & T a l a v era (1979:186). It is a rather rare species. The shells are r o s a - b r o w n i n colour. D e p t h r a n g e . — 0-120 m . Odostomia (Megastomia) desmiti spec. n o v . (figs 38, 62) M a t e r i a l . — H o l o t y p e ( N N M 57524): M.065, M a u r i t a n i a , off Banc d ' A r g u i n ; 20°00'N 17°11'W; depth 25 m, m u d d y sand w i t h C a r d i u m and Lanice; v a n V e e n grab; 13.vi.1988. Paratypes ( N N M 57525-57532; Z M A 3.97.043). M a u r i t a n i a : M.030/4, M.035/6, M.064/6, M.097/1, M.113/3; 4.112/1, 4.159/1; B8/5. M.065/5, D e s c r i p t i o n . — Shell f o r m i n g a slender cone w i t h straight sides. C o l o u r w h i t i s h i n Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) 35 Figs 32-38. Odostomia species. 32, Odostomia meijeri spec, nov., holotype ( N N M 57512), 1.6 m m , sta. M A U . 0 4 3 , off Mauritania, 19°04'N 16°25'W, depth 15 m , l l - v i - 1 9 8 8 ; 33, Odostomia meijeri spec. nov. var., 1.8 m m , sta. 3.172, Mauritania, 20°21'N 17°17'W, depth 34 m , l-xi-1978. ; 34, Odostomia truncatula Jeffreys, 2.3 m m , sta. 7.121, Cape V e r d e Islands, S of Razo, 16°36'N 24°37'W, depth 200-230 m , l-ix-1986; 35, Odostomia vanurki spec, nov., holotype ( N N M 57520), 1.7 m m , sta. 7.065, Cape V e r d e Islands, W of Boa Vista, S W of Ilhéu Calheta do Velho, 16°10'N 22°58'W, depth 39 m , 27-viii-1986; 36, Odostomia conoidea (Brocchi) (lectotype of O. sulcifera Smith, B M N H 1870.1.1229), 4.6 m m , West Africa, W h y d a h ; 37, Odostomia conoidea var., 2.7 m m , sta. M A U . 0 9 7 , Mauritania, off Banc d ' A r g u i n , 19°24'N 16°48'W, depth 26 m , 16-vi-1988; 38, Odostomia desmiti spec, nov., holotype ( N N M 57524), 3.7 m m , sta. M A U . 0 6 5 , Mauritania, off Banc d ' A r g u i n , 20°00'N 17°11'W, depth 25 m , 13-vi-1988. 36 Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) d e a d material. E m b r y o n i c w h o r l s intorted, of type B. Teleoconch of about five w h o r l s , w h i c h are s l i g h t l y convex. Suture w e l l - m a r k e d , b u t not v e r y d e e p l y i n c i s e d . G r o w t h lines orthocline to s l i g h t l y prosocline. A p a r t f r o m the g r o w t h l i n e s , the sculpture consists of a single s p i r a l line a r o u n d the p e r i p h e r y . O u t e r l i p e v e n l y r o u n d e d , w i t h a n u m b e r of s p i r a l riblets inside. U m b i l i c u s m o d e r a t e l y p r o m i n e n t . T o o t h clearly marked. D i m e n s i o n s : H . 2.7-3.7 m m , W . 1.5-1.8 m m ; h o l o t y p e : 3.7 x 1.7 m m . D i f f e r e n t i a t i o n . — T h i s species differs f r o m the w e l l - k n o w n O. conoidea (Brocchi, 1814) a n d related forms b y its type B (instead of type A ) protoconch, i n d i c a t i v e of a non-pelagic e m b r y o n i c d e v e l o p m e n t . D e p t h r a n g e . — 24-200 m . E t y m o l o g y . — T h i s species is n a m e d after H . G . de Smit, former m e m b e r of the MWH. Odostomia (Megastomia) zijpi spec. n o v . (fig. 39) M a t e r i a l . — H o l o t y p e ( N N M 57533): St. M.065, M a u r i t a n i a , off Banc d ' A r g u i n ; 20°00'N 17°11'W; depth 25 m , m u d d y sand w i t h Cardium and Lanice; v a n Veen grab; 13.vi.1988. Paratypes ( N N M 57534): Mauritania: M.065/3. D e s c r i p t i o n . — Shell f o r m i n g a v e r y b r o a d cone w i t h straight sides. C o l o u r c h a l k y - w h i t e . E m b r y o n i c w h o r l s intorted, of type B. Teleoconch consisting of about three to three a n d a quarter, rather flat w h o r l s . Suture i m p r e s s e d b u t not deep. G r o w t h l i n e s s l i g h t l y prosocline. N o sculpture except for the g r o w t h l i n e s . O u t e r l i p sharp, i n s i d e w i t h about seven s p i r a l riblets. W i t h o u t a n u m b i l i c u s or w i t h o n l y a v e r y s m a l l u m b i l i c a l c h i n k . T o o t h rather p r o m i n e n t . D i m e n s i o n s : H . 1.8-2.1 m m , W . 1.2-1.4 m m ; h o l o t y p e : 2.1 x 1.4 m m . D i f f e r e n t i a t i o n . — A remarkable species. T h e shell is m u c h broader t h a n i n O. (M.) desmiti spec, n o v . , w i t h w h i c h it c a n be c o m p a r e d . W e f o u n d o n l y f o u r specimens, so it is possible that f u l l - g r o w n shells reach larger d i m e n s i o n s . Its s t u m p y shape differentiates this species clearly f r o m O. (M.) conoidea (Brocchi, 1814), O. (M.) desmiti a n d s i m i l a r species . D e p t h . — 25 m . E t y m o l o g y . — T h i s species is n a m e d after J . C . M . v a n Z i j p , former m e m b e r of the MWH. Odostomia (Liostomia) clavula ( L o v e n , 1846) Turbonilla clavula L o v e n , 1846:18. Odostomia clavula; v a n Aartsen, 1987: 6, p i . 28 figs 6, 7; Penas et al., 1996: 42, 43, figs 96, 97. Liostomia clavula; Warén, 1991:106,107, fig. 35C, D , G . M a t e r i a l . — Mauritania: M.030/1, M.031/7, M.032/1, M.034/1, M.077/3, M.078/4, M.084/1, M.085/4, M.087/1, M.097/3, M.099/4, M.135/1, 3.113/1, 3.133/2, 3.172/1, 3.194/>25; B l / 4 , B2/1, B5/4, B7/6, St. B8/4. Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) 37 N o m e n c l a t u r e . — F o r the time b e i n g w e d o not f o l l o w Warén (1991:106) i n the use of Liostomia Monterosato, 1884, as a f u l l genus b u t prefer to use it w i t h the majority of the authors as a subgenus of Odostomia F l e m i n g , 1813. D i s t r i b u t i o n . — O. clavula has not been reported before f r o m o u r research area. D e p t h r a n g e . — 19-100 m . Odostomia (Pyramistomia) fehrae spec. n o v . (fig. 40) M a t e r i a l . — H o l o t y p e ( N N M 57535): St. 5.140, Azores, W of Pico; 38°34'N 28°33'W; depth 88 m ; v a n Veen grab; 7.vi.l981. Paratypes ( N N M 57536-57543). Azores: 5.051/6, 5.058/2, 5.093/1, 5.111/1, 5.122/1, 5.131/2, 5.135/2, 5.140/10. D e s c r i p t i o n . — Shell strong, f o r m i n g a slender cone w i t h straight sides, interrupte d b y b r o a d channels. C o l o u r w h i t i s h . E m b r y o n i c w h o r l s p l a n o r b i d , of type B. Teleoconch of four s h o u l d e r e d w h o r l s , f o r m i n g a right angle near the u p p e r suture, continu i n g w i t h a slightly concave part t o w a r d s the l o w e r p e r i p h e r y , w h i c h is accentuated b y a p r o m i n e n t s p i r a l rib, a n d reaches the l o w e r suture nearly vertically. Suture o n l y slightly i m p r e s s e d , b u t b r o a d l y channelled because of the sculpture. G r o w t h l i n e s clearly prosocline. Except for the g r o w t h l i n e s , the sculpture consists of the strong, s p i r a l rib at the abapical side of the w h o r l s , a n d a less p r o m i n e n t one d e l i m i t i n g the shell base. The f o r m of the outer l i p reflects the shape of the w h o r l s . U m b i l i c u s lacking. Tooth well-marked. D i m e n s i o n s : H . 1.7-1.9 m m , W . 0.7-0.9 m m ; h o l o t y p e : 1.7 x 0.9 m m . D i f f e r e n t i a t i o n . — T h i s species is u n l i k e any other recent E u r o p e a n p y r a m i d e l l i d . It has some analogy w i t h the fossil O. deubeli Boettger, 1901, as f i g u r e d b y C o s s m a n n (1921: p i . 6 figs 34, 35) a n d Z i l c h (1934: 238, pl.12 fig.14). H o w e v e r , that species has a definite s p i r a l rib at the a d a p i c a l side of the w h o r l s , w h i c h splits into t w o ribs o n the l o w e r w h o r l s ; i n o u r species there is o n l y a c o n s p i c u o u s l y angular part b e l o w the suture b u t no u p p e r s p i r a l rib. Still, the general m o r p h o l o g y of the t w o species s u g gests a close relationship a n d because O. deubeli is the type species of Pyramistomia C o s s m a n n , 1921, w e classify o u r species w i t h that taxon, w h i c h is g i v e n a subgeneric status here. D e p t h r a n g e . — 92-400 m . E t y m o l o g y . — T h i s species is n a m e d after M r s . M . C . Fehr-de W a l , former m e m b e r of the M W H . P e n d i n g further research o n the generic s u b d i v i s i o n of the genus Odostomia, the f o l l o w i n g species are not classified i n a subgenus. Odostomia bernardi spec. n o v . (figs 41, 63) M a t e r i a l . — H o l o t y p e ( N N M 57544): 5.091, Azores, E of Faial; 38°51'N 28°36'W; depth 33 m , fine v o l canic sand w i t h shell gravel; v a n Veen grab; 2.vi.l981. Paratypes ( N N M 57545-57577; Z M A 3.97.018-025). Azores: A Z O 20/12, A Z O 24a/l, 5.006/1, 5.008/3, 38 Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) 5.026/1, 5.039/1, 5.050/7, 5.051/22, 5.053/1, 5.054/21, 5.055/1, 5.064/1, 5.077/2, 5.084/15, 5.091/>25, 5.092/>25, 5.094/1, 5.096/13, 5.100/4, 5.112/16, 5.113/11, 5.121/1, 5.122/3, 5.126/1, 5.128/7, 5.130/6, 5.132/13, 5.135/10, 5.138/4, 5.139/1, 5.140/4, 5.160/1, 5.190/1; Säo M i g u e l , C a l o u r a littoral/3, Terceira, A n g r a de Heroismo harbour/>25, Säo M i g u e l , Säo R o g u e / l , Faial, Porto P i n near H o r t a , beach/3, Säo M i g u e l , Praia d o Populo/14, Säo M i g u e l , Praia d o Populo/>25, Terceira/3, Säo M i g u e l , Ponta Delgada (-8 m)/4 [in Z M A ] . D e s c r i p t i o n . — Shell egg-shaped. C o l o u r w h i t i s h i n d e a d material. Embryonic w h o r l s of type C . Teleoconch consisting of t w o a n d a half to three s l i g h t l y convex w h o r l s , the last one relatively v e r y large. Suture w e l l - m a r k e d , moderately incised. G r o w t h l i n e s orthocline to v e r y s l i g h t l y prosocline. Except f o r the w e l l - m a r k e d g r o w t h l i n e s there is n o clear sculpture. O u t e r l i p rather straight adapically, basally m o r e strongly c u r v e d . U m b i l i c u s l a c k i n g . There is a clear tooth, situated deeply inside the aperture. D i m e n s i o n s : H . 1.4-1.6 m m , W . 0.7-0.8 m m ; holotype: 1.4 x 0.7 m m . D i f f e r e n t i a t i o n . — This relatively c o m m o n species is s i m i l a r to the E u r o p e a n Odostomia glabrata Forbes & H a n l e y , 1850, as f i g u r e d b y v a n A a r t s e n (1987: 30, f i g . 26). T h e latter species, h o w e v e r , has m o r e slender shells, w h i c h are not egg-shaped a n d p r o v i d e d w i t h clearly prosocline g r o w t h l i n e s . T h e e m b r y o n i c w h o r l s of O. glabrata are also of type C , b u t m u c h larger. D e p t h r a n g e . — 0-300 m . E t y m o l o g y . — This species is n a m e d after J.J. B e r n a r d , former m e m b e r of the MWH. Odostomia dalsumi spec. n o v . (figs 42, 64) M a t e r i a l . — H o l o t y p e ( N N M 57578): 7.028, Cape Verde Islands, SE of C i m a ; 14°57'N 24°39'W; depth 225 m , y e l l o w sand w i t h shell gravel; v a n V e e n grab; 23.viii.1986. Paratypes ( N N M 57579): Cape Verde Islands: 7.028/6. D e s c r i p t i o n . — Shell short, egg-shaped. C o l o u r w h i t e . E m b r y o n i c w h o r l s intorted, of type C . Teleoconch of little m o r e than t w o s h o u l d e r e d w h o r l s . Suture clearly m a r k e d , moderately incised. G r o w t h l i n e s prosocline a n d v e r y clearly m a r k e d . O n l y the p r o m i n e n t g r o w t h l i n e s f o r m a k i n d of sculpture. O u t e r l i p t h i n . U m b i l i c u s s m a l l but conspicuous. T o o t h weak, situated deeply inside the aperture. D i m e n s i o n s : H . 0.9-1.0 m m , W . 0.7 m m ; h o l o t y p e : 0.9 x 0.7 m m . D i f f e r e n t i a t i o n . — F o r differences of this v e r y s m a l l species w i t h O. mamoi M i f s u d , 1993, a n d O. prinsi spec, nov., see the notes o n the latter species. D e p t h . — 225 m . E t y m o l o g y . — T h i s species is n a m e d after J. v a n D a l s u m , former m e m b e r of the MWH. Odostomia mamoi M i f s u d , 1993 (fig. 43) Odostomia mamoi M i f s u d , 1993:16-18, figs; N o f r o n i & Tringali, 1995: 36, 37, p i . 7 figs 29-33. 39 Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) M a t e r i a l — Canary Islands: 2.022/2 (fig. 43), 2.085/1. D e s c r i p t i o n . — Shell b r o a d l y egg-shaped. C o l o u r w h i t i s h transparent. E m b r y o n i c w h o r l s intorted, of type C . Teleoconch of t w o to t w o a n d a half w h o r l s . Suture clearly m a r k e d , moderately i n c i s e d . G r o w t h l i n e s prosocline a n d w e l l - m a r k e d ; there is n o other sculpture. O u t e r l i p t h i n , r e g u l a r l y c u r v e d . U m b i l i c u s rather s m a l l b u t w e l l m a r k e d . N o tooth o n the c o l u m e l l a . D i m e n s i o n s : H . 1.5 m m , W . 1.0 m m ; f i g u r e d s p e c i m e n (fig. 43): 1.5 x 1.0 m m . D i s t r i b u t i o n . — O r i g i n a l l y described f r o m the M e d i t e r r a n e a n , this species has recently been m e n t i o n e d f r o m Tenerife, C a n a r y Islands, b y N o f r o n i & T r i n g a l i (1995). O u r shells also originate f r o m the C a n a r y Islands. T h i s species is n o t k n o w n f r o m the C a p e V e r d e Islands, b u t t w o m o r e or less s i m i l a r taxa, v i z . O. dalsumi spec. n o v . a n d O. prinsi spec. n o v . are reported f r o m that region. D e p t h r a n g e . — 83-500 m . Odostomia prinsi spec. n o v . (figs 44, 65) M a t e r i a l . — H o l o t y p e ( N N M 57580): 6.093, Cape Verde Islands, S W of Razo; 16°36'N 24°37'W; depth 400-430 m , sand and shell gravel; v a n V e e n grab; 15.vi. 1982. Paratypes ( N N M 57581-57582): Cape Verde Islands: 6.093/4, 7.128/3. D e s c r i p t i o n . — Shell r i s s o i d , rather s m a l l . C o l o u r w h i t e . E m b r y o n i c w h o r l s intorted, of type C . Teleoconch consisting of t w o to t w o a n d a half w h o r l s . Suture clearly d e f i n e d , b u t not i n c i s e d . G r o w t h l i n e s v e r y p r o n o u n c e d , r i b - l i k e , a n d clearly prosocline; n o a d d i t i o n a l sculpture present. O u t e r l i p t h i n , r e g u l a r l y c u r v e d . U m b i l i c u s s m a l l , b u t clearly present. T o o t h s m a l l b u t w e l l - d e v e l o p e d , situated deeply i n s i d e the aperture. D i m e n s i o n s : H . 1.2-1.3 m m , W . 0.8 m m ; h o l o t y p e : 1.2 x 0.8 m m . D i f f e r e n t i a t i o n . — W i t h O. mamoi a n d O. dalsumi spec, n o v . , O. prinsi spec. n o v . belongs to a g r o u p of three species w i t h s m a l l shells w i t h rather p r o m i n e n t , prosocline g r o w t h l i n e s . I n O. mamoi a n d O. prinsi the w h o r l s are r e g u l a r l y r o u n d e d , w h e r e as they are clearly s h o u l d e r e d i n O. dalsumi. I n O. dalsumi the smallest of the three, the tooth is less clearly d e v e l o p e d t h a n i n O. prinsi. I n O. mamoi, the largest species, there is n o tooth o n the c o l u m e l l a . D e p t h r a n g e . — 400 m . E t y m o l o g y . — T h i s species is n a m e d after B. P r i n s , former m e m b e r of the M W H . Odostomia nitens Jeffreys, 1870 Odostomia nitens Jeffreys, 1870: 79; v a n Aartsen, 1987: 6, p i . 28 fig. 4. M a t e r i a l . — Canary Islands: 2.011/1, 2.085/1, 4.058/3, 4.060/11, 4.087/1. Azores: 5.012/>25, 5.051/>25, 5.122/2. Cape Verde Islands: 6.093/1, 7.039/3, 7.052/1. N o m e n c l a t u r e . — T h i s w e l l - k n o w n species is the type species (by m o n o t y p y ) of Doliella Monterosato, 1880. 40 Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) D i s t r i b u t i o n . — Odostomia nitens is k n o w n f r o m rather deep water a n d has been reported f r o m the A z o r e s b y Jeffreys (1884: 349), W a t s o n (1885: 480) a n d D a u t z e n b e r g (1889: 59). D e p t h r a n g e . — 125-700 m . Odostomia marthinae ( N o f r o n i & Schänder, 1994) (fig. 45) Chrysallida marthinae N o f r o n i & Schänder, 1994: 2, 8, 9, figs l a , b, 2f. M a t e r i a l . — M a u r i t a n i a : M.013/5, M.030/19, M.031/>25, M.032/4, M.033/4, M.043/16, M.044/18, M.045/1, M.046/2, M.077/>25, M.078/>25, M.079/>25, M.084/19, M.085/6, M.097/>25, M.099/4, M . l l l / > 2 5 , M.113/>25, M.114/1, M.115/9, M.121/21, M.135/8; 3.109/4, 3.113/7, 3.123/1, 3.128/1, 3.133/5, 3.172/>25, 3.194/>25; B I /5, B2/15, B3/2, B5/5, B7/>25, B8/>25. N o m e n c l a t u r e . — T h i s recently described species is v e r y characteristic b u t d i f f i c u l t to classify w i t h one of the m a n y m o r p h o l o g i c a l l y d e f i n e d (sub)genera i n the f a m i l y . W e d o not consider it a Chrysallida species, a n d tentatively classify it w i t h Odostomia. The f i g u r e d s p e c i m e n (fig. 45) measures 2.5 x 1.2 m m . D i s t r i b u t i o n . — T h i s is one of the m o r e c o m m o n West A f r i c a n p y r a m i d e l l i d s . W h i l e s t u d y i n g the M o l l u s c a of the b a y of A l g e c i r a s , the first author d i s c o v e r e d a perfectly w e l l preserved shell of this species, b u t as this w a s the o n l y s p e c i m e n f o u n d , it w a s not m e n t i o n e d i n the p u b l i s h e d report b y v a n A a r t s e n et al. (1984). D e p t h r a n g e . — 14-100 m . G e n u s Puposyrnola C o s s m a n n , 1921 T y p e species (by o r i g i n a l designation): Auricula acicula L a m a r c k , 1804 . The type species of this genus is the eocene fossil Auricula acicula L a m a r c k , 1804. This species is f i g u r e d b y C o s s m a n n (1921: 229, p i . 5 figs 60-64), w h o based the 'subgenus' Puposyrnola o n the p u p o i d a l outline of the shell, i n contrast to Syrnola A. A d a m s , 1860. U s u a l l y , b o t h Syrnola a n d Puposyrnola species are s u p p o s e d to have a w e l l - d e v e l o p e d tooth o n the c o l u m e l l a . T h i s tooth is m i s s i n g i n the t w o species that are described b e l o w . T h e i r classification is based o n the general shape of the shells, w h i c h is clearly p u p o i d . Syrnola minuta H . A d a m s , 1869, has p r e v i o u s l y been p l a c e d i n Puposyrnola b y v a n A a r t s e n (1994: 93) a n d Penas et al. (1996: 38). Puposyrnola minuta ( H . A d a m s , 1869) Syrnola minuta H . A d a m s , 1869: 274, p l . 19 fig. 10. Puposyrnola minuta; v a n Aartsen, 1994: 93, p i . 107 fig. 10; Penas et al., 1996: 38, 41, figs 89. 90, 93. M a t e r i a l . — Canary Islands: 4.060/1. M a u r i t a n i a : M.035/2, 2.036/4, 3.194/1. N o t e s . — T h i s w e l l - k n o w n A t l a n t i c - M e d i t e r r a n e a n species, once reported f r o m o u r research area, f r o m O r o t a v a , C a n a r y Islands ( H . A d a m s , 1869) is present i n o u r Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) 41 Figs 39-45. Odostomia species. 39, Odostomia zijpi spec, nov., holotype ( N N M 57533), 2.1 m m , sta. M A U . 0 6 5 , M a u r i t a n i a , off Banc d A r g u i n , 20°00'N 17°11'W, depth 25 m , 13-vi-1988; 40, Odostomia fehrae spec, nov., holotype ( N N M 57535), 1.7 m m , sta 5.140, Azores, W of Pico, 38°34'N 28°33'W, depth 88 m , 7-vi-1981; 41, Odostomia bernardi spec, nov., holotype ( N N M 57544), 1.4 m m , sta. 5.091, Azores, E of Faial, 38°51'N 28°36'W, depth 33 m , 2-vi-1981; 42, Odostomia dalsumi spec, nov., holotype ( N N M 57578), 0.9 m m , sta. 7.028, Cape V e r d e Islands, SE of C i m a , 14°57'N 24°39'W, depth 225 m , 23-viii-1986; 43, Odostomia mamoi M i f s u d , 1.5 m m , sta. 2.022, Canary Islands, S of Fuerteventura, Punta de Jandia, 28°03'N 14°30'W, depth 83-97 m , 25-viii-1977; 44, Odostomia prinsi spec, nov., holotype ( N N M 57580), 1.2 m m , sta. 6.093, C a p e V e r d e Islands, S W of Razo, 16°36'N 24°37'W, depth 400-430 m , 15-vi-1982; 45, Odostomia (?) marthinae (Nofroni & Schänder), 2.5 m m , sta. M A U . 1 1 4 , M a u ritania, off Banc d ' A r g u i n , 20°29"N 17°14'W, depth 36 m , 18-vi-1988. 42 Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) material too. T h e shells v a r y i n slenderness, b u t are a l w a y s characteristic. T h e g o l d e n - y e l l o w b a n d m e n t i o n e d i n the o r i g i n a l d e s c r i p t i o n is rarely present i n e m p t y shells. O u r largest s p e c i m e n measures 4.0 m m i n height. D e p t h r a n g e . — 78-580 m . Puposyrnola kaasi spec. n o v . (figs 46, 66) M a t e r i a l . — H o l o t y p e ( N N M 57583): 7.100, Cape V e r d e Islands, W of Sal, off Palmeira; 16°46'N 23°01'W; depth 354 m , volcanic a n d calcareous m u d , sand a n d gravel; v a n V e e n grab; 30.viii.1986. Paratype ( N N M 57584): Cape Verde Islands: 6.079/1. Description.— Shell subcylindrical w i t h a p u p o i d a l top. C o l o u r m i l k y white. E m b r y o n i c w h o r l s p l a n o r b i d , i n t o r t e d , of type B. Teleoconch of six to seven s l i g h t l y concave w h o r l s . Suture clearly i n c i s e d , b u t not c h a n n e l l e d . G r o w t h l i n e s prosocline; w i t h a n a d d i t i o n a l , m i c r o s c o p i c s p i r a l striation. O u t e r l i p straight. U m b i l i c u s l a c k i n g . N e i t h e r a c o l u m e l l a r tooth n o r a f o l d are present. D i m e n s i o n s : H . 4.4 m m , W . 1.4 m m ; h o l o t y p e : 4.4 x 1.4 m m . D i f f e r e n t i a t i o n . — C o m p a r e d w i t h the o n l y other E u r o p e a n Puposyrnola species, v i z . P. minuta ( H . A d a m s , 1869), shells of this species are larger, m o r e c y l i n d r i c a l , a n d p r o v i d e d w i t h a less clearly p u p o i d a l t o p . D e p t h r a n g e . — 270-354 m . E t y m o l o g y . — T h i s species is n a m e d after P. K a a s , former m e m b e r of the M W H . G e n u s Eulimella Forbes & M a c A n d r e w , 1846 T y p e species (by o r i g i n a l d e s i g n a t i o n [see v a n A a r t s e n , 1988: 173]): Eulima macandrei Forbes, 1844 [= E. Scillae (Scacchi, 1835)]. T h i s genus comprises species w i t h slender, s m o o t h shells w i t h m a n y w h o r l s , a n d w i t h o u t a c o l u m e l l a r tooth. W e f o l l o w the majority of the authors i n c o n s i d e r i n g the fossil Eulimella scillae (Scacchi, 1835) a n d Eulimella macandrei (Forbes, 1844) one a n d the same species. T h e subgenus n a m e Ptycheulimella Sacco, 1892, is sometimes u s e d for species w i t h shells w i t h a w e a k f o l d o n the c o l u m e l l a a n d frequently a b r o w n i s h s p i r a l c o l o u r b a n d . A s l o n g as the i d e n t i f i c a t i o n of the type species of the subgenus, v i z . Tornatella pyramidata Deshayes, 1835, is most uncertain, w e prefer not to s u b d i v i d e Eulimella s.l. Eulimella unifasciata (Forbes, 1844) Eulima unifasciata Forbes, 1844:188. Syrnola unifasciata; C a r r o z z a , 1977:178, p i . 2 fig. 1; Terreni, 1981: 46, pl.7 fig.10. Eulimella unifasciata; v a n Aartsen, 1994: 99, p i . 109 fig. 19. M a t e r i a l . — C a n a r y Islands: 2.114/3, 2.155/1, 4.080/1,4.082/1,4.115/2. N o m e n c l a t u r e . — Several authors p l a c e d this species i n the genus Syrnola A . A d a m s , 1860. W e d o not agree w i t h this classification, because Syrnola is s u p p o s e d to Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) 43 be characterized b y the presence of a p r o m i n e n t tooth, w h i c h is l a c k i n g i n this species. Distribution.—Eulimella unifasciata is k n o w n f r o m rather deep water a n d has prev i o u s l y been reported f r o m "off the Sahara" a n d f r o m the A z o r e s b y Jeffreys (1884: 351) a n d D a u t z e n b e r g (1889: 60), a n d f r o m M a u r i t a n i a b y A l t i m i r a (1978:182). D e p t h r a n g e . — 200-700 m . Eulimella verduini spec. n o v . (fig. 47) Material — H o l o t y p e ( N N M 57585): 2.010, Canary Islands, S of Fuerteventura, Punta de Jandia; 28°03"N 14°29'W; depth 100-300 m , sand and shell gravel; v a n Veen grab; 24.viii.1977. Paratypes ( N N M 57586-56590). Canary Islands: 2.010/1, 2.034/1, 4.044/2, 4.048/3, 4.067/1. Three specimens f r o m Station 4.080 are tentatively classified w i t h this species. D e s c r i p t i o n . — Shell f o r m i n g a v e r y slender, elongated cone, w i t h a rather obtuse top. C o l o u r w h i t i s h , w i t h a w e l l - d e l i m i t e d , n a r r o w , s u p r a p e r i p h e r a l , b r o w n b a n d . Between this b a n d a n d the abapical suture there is a l w a y s a n a r r o w w h i t i s h zone. E m b r y o n i c w h o r l s p l a n o r b i d , of type A I; rather large, v i z . 400 jum i n w i d t h . Teleoconch f o r m e d b y six to seven w h o r l s w h i c h are s l i g h t l y concave. Suture w e l l - m a r k e d b e l o w the p e r i p h e r y of the p r e c e d i n g w h o r l . G r o w t h l i n e s orthocline a n d rather p r o m inent, s i m u l a t i n g axial ribs. Except for the clearly m a r k e d g r o w t h l i n e s , there is o n l y a vague s u p r a p e r i p h e r a l angle, just above the abapical suture of each w h o r l . O u t e r l i p straight a d a p i c a l l y a n d gently c u r v e d b e l o w . U m b i l i c u s l a c k i n g . O n the c o l u m e l l a there is n o tooth, but u s u a l l y a n indistinct f o l d . D i m e n s i o n s : H . 3.1-4.0 m m , W . 0.9-1.0 m m ; holotype: 3.1 x 1.0 m m . D i f f e r e n t i a t i o n . — In this species there is a n a r r o w , b r o w n i s h , s p i r a l b a n d o n the l o w e r part of each w h o r l , just as i n E. unifasciata (Forbes, 1844), E. neoattenuata G a g l i n i , 1992, a n d E. vanhareni spec, nov., but it differs f r o m a l l of these b y the clearly concave w h o r l s a n d the m u c h more accentuated g r o w t h l i n e s . D e p t h r a n g e . — 47-300 m . E t y m o l o g y . — This species is n a m e d after the late A . V e r d u i n , w e l l - k n o w n m a l a cologist a n d g o o d f r i e n d of the authors. Eulimella robusta spec. n o v . (figs 48, 67) M a t e r i a l . — H o l o t y p e ( N N M 57591): M.034, off M a u r i t a n i a ; 18°46'N 16°40'W; depth 167 m , m u d d y fine sand, some shell gravel; v a n Veen grab; 9.vi.l988. Paratypes ( N N M 57592-57593): M a u r i t a n i a : M . 0 3 4 / l , M.035/1. D e s c r i p t i o n . — Shell strong, conical w i t h straight sides a n d a rather obtuse top. C o l o u r w h i t i s h , no colour-bands c o u l d be seen. E m b r y o n i c w h o r l s p l a n o r b i d , of type A I, w i t h a w i d t h of about 450 jum. Teleoconch of five to six flat or s l i g h t l y concave w h o r l s w i t h a s o m e w h a t angled p e r i p h e r y . Suture w e l l - m a r k e d . G r o w t h l i n e s somew h a t prosocline; a d d i t i o n a l l y there is o n l y a fine microscopic s p i r a l striation. O u t e r l i p straight a d a p i c a l l y a n d gently c u r v e d t o w a r d s the c o l u m e l l a , w h i c h is nearly 44 Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) straight. U m b i l i c u s l a c k i n g . O n the c o l u m e l l a there is n o tooth, b u t o n l y a n indistinct fold. D i m e n s i o n s : H . 4-5 m m , W . 1.7-2.0 m m ; h o l o t y p e : 4.7 x 1.8 m m . D i f f e r e n t i a t i o n . — T h i s species is most s i m i l a r to the recently described Eulimella telum Schänder, 1994, b u t the shells are larger, have a larger p r o t o c o n c h a n d also a different shape (see Schänder, 1994: 62, f i g . 3e). D e p t h r a n g e . — 167-200 m . E t y m o l o g y . — T h e epithet robusta refers to the strong appearance of the shells. Eulimella sinuata spec. n o v . (fig. 49) M a t e r i a l . — H o l o t y p e ( N N M 57594): 7.039, Cape Verde Islands, SE of C i m a ; 14°56'N 24°38'W; depth 590-610 m , volcanic a n d calcareous sand; v a n Veen grab; 24.viii.1986. Paratypes ( N N M 57595-57598). Cape Verde Islands: 6.095/1, 7.015/2, 7.029/1, 7.039/3. D e s c r i p t i o n . — Shell rather stout, f o r m i n g a v e r y slender, elongated cone. C o l o u r w h i t i s h , sometimes w i t h a v e r y faint y e l l o w i s h - b r o w n , s p i r a l c o l o u r b a n d a r o u n d the p e r i p h e r y . E m b r y o n i c w h o r l s s o m e w h a t h e l i c o i d , of type A . Teleoconch of six to seven nearly flat w h o r l s w h i c h are s l i g h t l y a n g l e d at the p e r i p h e r y . Suture v e r y clearl y m a r k e d . G r o w t h l i n e s prosocline; w i t h o u t further sculpture. O u t e r l i p straight a d a p i c a l l y , gently c u r v e d b e l o w . U m b i l i c u s l a c k i n g . There is n o c o l u m e l l a r tooth b u t o n l y a n indistinct f o l d . D i m e n s i o n s : H . 3.3-4.5 m m , W . 1.1-1.4 m m ; h o l o t y p e : 3.3 x 1.1 m m . D i f f e r e n t i a t i o n . — F r o m the other Eulimella species w i t h a g o l d e n b r o w n s p i r a l b a n d , this species differs m a i n l y b y its prosocline g r o w t h l i n e s ; these are opisthocline i n E. unifasciata a n d E. vanhareni spec. n o v . T h e w h o r l s are neither flat as i n E. unifasciata n o r concave as i n E. verduini spec. n o v . D e p t h r a n g e . — 225-930 m . E t y m o l o g y . — T h e epithet sinuata refers to the s l i g h t l y sinuate outline of the whorls. Eulimella vanhareni spec. n o v . (figs 50, 68) M a t e r i a l . — H o l o t y p e ( N N M 57599): 4.116, Canary Islands, S of Palma; 28°26'N 17°51'W; depth 420 m , gravel, sand a n d shells; v a n Veen grab; 28.V.1980. Paratypes ( N N M 57600-57607). Canary Islands: 2.014/2, 2.075/1, 2.084/1, 2.103/1, 2.120/1, 4.048/5, 4.116/5, 4.156/2. Selvagens Archipelago: 3.099/1. D e s c r i p t i o n . — Shell f o r m i n g a v e r y m u c h elongated cone w i t h straight sides. C o l o u r w h i t i s h , w i t h a g o l d e n - y e l l o w s p i r a l b a n d just above the p e r i p h e r y . E m b r y o n ic w h o r l s p l a n o r b i d , of type A I to B. Teleoconch consisting of seven to eight w h o r l s w h i c h are flat to v e r y s l i g t h l y convex. Suture v a g u e l y incised. G r o w t h l i n e s opisthocline i n the l o w e r w h o r l s ; w i t h o u t a d d i t i o n a l sculpture. O u t e r l i p gently c u r v e d , w i t h several tooth-like riblets i n s i d e . U m b i l i c u s l a c k i n g . There is n o c o l u m e l l a r tooth, b u t a clear f o l d is u s u a l l y present. Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) 45 D i m e n s i o n s : H . 4 - 6 m m , W . 1.1 - 1 . 5 m m ; h o l o t y p e : 4.6 x 1.3 m m . D i f f e r e n t i a t i o n . — Because of its c o l o u r e d s p i r a l b a n d , this species is clearly s i m i lar to the w e l l - k n o w n E. unifasciata (Forbes, 1844). That species has the same type of p r o t o c o n c h as w e l l as opisthocline g r o w t h l i n e s ( v a n A a r t s e n , 1994: 99, 109, fig.19). H o w e v e r the c o l o u r e d b a n d is broader a n d not clearly d e l i m i t e d i n E. unifasciata a n d there are n o riblets o n the i n s i d e of the outer l i p . Eulimella vanhareni spec. n o v . is s o m e w h a t s i m i l a r to "Syrnola" wenzi N o r d s i e c k , 1972 [= Odostomia crassa Jeffreys, 1884, not Odostomia crassa ( T h o m p s o n , 1845)], b u t that species (only fragments are k n o w n ) has orthocline to s l i g t h l y prosocline g r o w t h lines a n d a w e l l - d e v e l o p e d tooth o n the c o l u m e l l a (van A a r t s e n , 1994: 94). Eulimella species are s u p p o s e d to have shells w i t h o u t teeth o n the i n s i d e of the outer l i p . Nevertheless, this species is classified here. T h e present lack of k n o w l e d g e about the m o r e t h a n 300 n o m i n a l genera of the P y r a m i d e l l i d a e leaves u s n o m o r e acceptable alternative t h a n c o n s i d e r i n g this a n aberrant Eulimella species. D e p t h r a n g e . — 200-585 m . E t y m o l o g y . — T h i s species is n a m e d after H . v a n H a r e n , former m e m b e r of the MWH. Eulimella tydemani spec. n o v . (fig. 51) M a t e r i a l . — H o l o t y p e ( N N M 57608): 7.068, Cape Verde Islands, W of Boa Vista, S W of Ilhéu Calheta do V e l h o ; 16°11'N 22°59'W; depth 40 m , fine yellow-grey sand, foramin.; v a n V e e n grab; 27.viii.1986. Paratypes ( N N M 57609-57610). Cape Verde Islands: 7.042/1, 7.079/10, 7.080/2. D e s c r i p t i o n . — Shell s u b c y l i n d r i c a l , w i t h a rather obtuse t o p . A n a r r o w light b r o w n s p i r a l b a n d r u n s a r o u n d the p e r i p h e r y a n d just above the l o w e r suture at the older w h o r l s . E m b r y o n i c w h o r l s p l a n o r b i d , of type A I, rather large, v i z . about 500 jum i n diameter a n d , characteristically, c o l o u r e d light b r o w n . Teleoconch of six to seven relatively h i g h flat w h o r l s . Suture incised, not channeled as i n the other Eulimella species here described. G r o w t h l i n e s v e r y s l i g h t l y prosocline to orthocline. O n l y an indistinct m i c r o s c o p i c s p i r a l striation is discernible. Palatal part of the outer l i p nearly straight, then a b r u p t l y c u r v e d u p w a r d s to reach the c o l u m e l l a . U m b i l i c u s lacki n g . O n the c o l u m e l l a there is n o tooth b u t o n l y a n indistinct f o l d . D i m e n s i o n s : H . 3.5 - 4.0 m m , W . 1.0 -1.15 m m ; h o l o t y p e : 3.7 x 1.1 m m . D i f f e r e n t i a t i o n . — A l t h o u g h the present species is s o m e w h a t s i m i l a r to Puposyrnola minuta ( H . A d a m s , 1869) i n general shape (see: v a n A a r t s e n , 1994:107, f i g . 10; Pefias et al., 1996: 41, f i g . 89) that species has a clearly different f o r m a n d a totally different apertural structure. M o r e o v e r , the t o p w h o r l of P. minuta is not c o l o u r e d l i g h t b r o w n , as it is i n E. tydemani spec. n o v . Because of its n o n - p u p o i d a l outline w e d o n o t place Eulimella tydemani i n Puposyrnola. D e p t h r a n g e . — 40-76 m . E t y m o l o g y . — T h e epithet tydemani refers to the research vessel T y d e m a n , u s e d d u r i n g the C A N C A P II-VII expeditions. 46 Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) Figs 46-51. Puposyrnola and Eulimella species. 46, Puposyrnola kaasi spec, nov., holotype ( N N M 57583), 4.4 m m , sta 7.100, Cape Verde Islands, W of Sal, off Palmeira, 16°46'N 23°01'W, depth 354 m , 30-viii-1986; 47, Eulimella verduini spec, nov., holotype ( N N M 57585), 3.1 m m , sta. 2.010, Canary Islands, S of Fuerteventura, Punta de Jandia, 28°03'N 14°29'W, depth 100-300 m , 24-viii-1977; 48, Eulimella robusta spec, nov., holotype ( N N M 57591), 4.7 m m , sta. M A U . 0 3 4 , off M a u r i t a n i a , 18°46'N 16°40'W, depth 167 m , 9-vi-1988; 49, Eulimella sinuata spec, nov., holotype ( N N M 57594), 3.3 m m , sta. 7.039, Cape Verde Islands, SE of C i m a , 14°56'N 24°38'W, depth 590-610 m , 24-viii-1986; 50, Eulimella vanhareni spec, nov., holotype ( N N M 57599), 4.6 m m , sta. 4.116, Canary Islands, S of Palma, 28°26'N 17°51'W, depth 420 m , 28-V-1980; 51, Eulimella tydemani spec, nov., holotype ( N N M 57608), 3.7 m m , sta. 7.068, Cape Verde Islands, W of Boa Vista, S W of Ilhéu Calheta do V e l h o , 16°11'N 22°59'W, depth 40 m , 27-viii-1986. Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) 47 Figs 52-60. Protoconchs. 52, Tiberia apicifusca spec, nov., holotype ( N N M 57279), sta. 6.012; 53, Pseudoscilla babylonia (C.B. A d a m s ) (lectotype of Aclis tricarinata Watson), coll N M W 1955.158.00655; 54, Odetta sulcata (de Folin), sta. R V , M a u r i t a n i a ( Z M A ) ; 55, Folinella moolenbeeki spec, nov., holotype ( Z M A 3.97.008); 56, Folinella excavata (Philippi), Porto Polio, Corse, France; 57, Odostomia acuta Jeffreys, Ria de A r o s a , Spain; 58, Odostomia omphaloessa Watson, sta. 4.088; 59, Odostomia verhoeveni spec, nov., holotype ( N N M 57402), sta. M A U . 0 9 7 ; 60, Odostomia schrami spec, nov., holotype ( N N M 57490), sta. 3.172. A l l scale-bars indicate 0.1 m m . 48 Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) Figs 61-68. Protoconchs. 61, Odostomia boermani spec, nov., holotype ( N N M 57503), sta. M A U . 0 3 3 ; 62, Odostomia desmiti spec, nov., holotype ( N N M 57524), sta.MAU.065; 63, Odostomia bernardi spec, nov., holotype ( N N M 57544), sta. 5.091; 64, Odostomia dalsumi spec, nov., holotype ( N N M 57578), sta. 7.028; 65, Odostomia prinsi spec, nov., holotype ( N N M 57580), sta. 6.093; 66, Puposyrnola kaasi spec, nov., h o l otype ( N N M 57583), sta 7.100; 67, Eulimella robusta spec, nov., holotype ( N N M 57591), sta. M A U . 0 3 4 ; 68, Eulimella vanhareni spec, nov., holotype ( N N M 57599), sta. 4.116. A l l scale-bars indicate 0.1 m m . Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) 49 Acknowledgements W e thank all colleagues w h o s u p p o r t e d o u r research i n any w a y , i n particular D r . P h . Bouchet ( M u s e u m N a t i o n a l d ' H i s t o i r e N a t u r e l l e , Paris), D r . M . G . H a r a s e w y c h ( N a t i o n a l M u s e u m of N a t u r a l H i s t o r y , W a s h i n g t o n ) , M r . H . K . M i e n i s ( H e b r e w U n i versity, Jerusalem), M r s . A . J. T r e w ( N a t i o n a l M u s e u m of Wales, C a r d i f f ) , a n d M r s . K . W a y (British M u s e u m ( N a t u r a l H i s t o r y ) , L o n d o n ) . References Aartsen, J.J. van, 1977. European Pyramidellidae: I. Chrysallida.— Conchiglie 13: 49-64. Aartsen, J.J. van, 1981. European Pyramidellidae: II. Turbonilla.— Boll, malac. 17: 61-88. Aartsen, J.J. van, 1984. The p y r a m i d e l l i d genera described by the marquis L . de F o l i n . — Boll, malac. 20:131-138. Aartsen, J.J. van, 1987. European Pyramidellidae:III. Odostomia and Ondina. — Boll, malac. 23:1-34. Aartsen, J.J. van, 1987a. Nomenclatural notes, 4. Melania scalaris P h i l i p p i , 1836, and Odostomia scalaris M a c g i l l i v r a y , 1843.— Basteria 51:149,150. Aartsen, J.J. van, 1988. Nomenclatural notes, 6. The generic name Eulimella (Gastropoda, Opisthobranchia, Pyramidellidae), authorship and type species.— Basteria 52:171-174. Aartsen, J.J. van, 1994. European Pyramidellidae: IV. The genera Eulimella, Anisocycla, Syrnola, Cingulina, Oscilla and Careliopsis.— Boll, malac. 30: 85-109. Aartsen, J.J. van, 1995. Anisocycla Monterosato, 1880 or Ebala Leach i n Gray, 1847: that is the quest i o n . — Boll, malac. 31: 65-68. Aartsen, J.J. v a n & J.X. Corgan, 1996. Thiele's pyramidellacean gastropod n a m e s . — Basteria 60: 177-182. Aartsen, J.J. van, H . P . M G . Menkhorst & E. Gittenberger, 1984. The marine M o l l u s c a of the bay of Algeciras, Spain, w i t h general notes o n Mitrella, Marginellidae and T u r r i d a e . — Basteria, S u p p l . 2: 1-135. Abbott, R.T., 1974. A m e r i c a n Seashells (Second edition):l-663.— N e w York, L o n d o n . Abbott, R.T. & S.P. Dance, 1982. C o m p e n d i u m of Seashells: i-x, 1-411.— N e w Y o r k . A d a m s , A . , 1860. 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Z i l c h , A . , 1934. Z u r Fauna des Mittel-Miocäns v o n Kostej ( B a n a t ) . — Senckenbergiana 16:193-302. Z i l c h , A . , 1935. Nachtrag z u r Typus-Bestimmung der Fauna des Mittel-Miocäns v o n Kostej (Banat).— Senkenbergiana 17: 226-228. Received: 18.iii.1997 Accepted: 20.iii.1997 Edited: J . C d e n H a r t o g 53 Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) Appendix The locality data of the M a u r i t a n i a I & II e x p e d i t i o n s Tyro Mauritania-I Expedition 1988 d i v i n g , snorkelling; 6.vi.l988. (material in Z M A ) M.004: Passe d u Lévrier, E of C a p Blanc; 20°47'N B l : Banc d ' A r g u i n ; 19°33.8'N 17°54.5'W; depth grab; 6.vi.l988. 53-64 m ; 13.V.1988. M.005: Passe d u Lévrier, E of C a p Blanc; 20°47'N B2: Banc d ' A r g u i n ; 19°32.7'N 16°59.9'W; depth 17°02'W; dept 15 m , sandy m u d ; V a n V e e n grab; 17°02'W; depth 15 m , sandy m u d ; V a n V e e n 99-152 m ; 15.V.1988. 6.vi.l988. B3: Banc d ' A r g u i n ; 19°37.4'N 16°52.1*W; depth M.006: Passe d u Lévrier, E of C a p Blanc; 20°49'N 21-34 m ; 17.V.1988. 17°01'W; dept 15 m , fine sandy m u d w i t h tube- B4: Banc d ' A r g u i n ; 19°40'N 16°56'W; depth 37-62 w o r m s , hermit-crabs, bivalves; 1.2 m Agassiz m ; 19.V.1988. trawl; 7.vi.l988. B5: Banc d ' A r g u i n ; 19°56.7'N 17°28.2'W; depth M.007: E coast of C a p Blanc; 20°47'N 17°03'W; lit- 85-154 m ; 22.V.1988. toral zone; hand-collecting, snorkelling; 7.vi.l988. B6: Banc d ' A r g u i n ; 20°15'N 17°08'W; depth 18-20 M.008: E coast of C a p Blanc; 20°47'N 17°03'W; m ; 24.V.1988. sublittoral zone, depth 0-2 m ; scuba d i v i n g , snor- B7: Banc d ' A r g u i n ; 20°00'N 17°26'W; depth 50-62 kelling, snorkelling; 7.vi.l988. m ; 24.V.1988. M.009: Passe d u Lévrier, E of C a p Blanc; 20°48'N B8: Banc d ' A r g u i n ; 20°05.4'N 17°21.6'W; depth 33 17°02'W; d e p t h 17 m , sandy m u d a n d shell grav- i n ; 28.V.1988. el; 1.2 m Agassiz trawl; 7.vi.l988. I: Banc d'Arguin, Baie d'Awatil; 19°46'N M.010: Passe d u Lévrier, E of C a p Blanc; 20°49'N littoral; siz trawl; 7.vi.l988. 17°29'W; littoral; 23.V.1988. II: Banc d'Arguin; 19°46'N 17°01'W; depth 17 m , m u d d y sand; 1.2 m A g a s 17°30'W; 23.V.1988. M.011: O f f coast; 18°50'N 16°18'W; depth 15 m , III: Banc d ' A r g u i n ; 19°46'30"N 17°31'W; littoral; h a r d bottom w i t h b r o w n i s h y e l l o w sand; V a n 23.V.1988. V e e n grab; 8.vi.l988. IV: Banc d ' A r g u i n ; 19°47'N 17°32'W; littoral; M.012: Off coast; 18°50'N 16°19'W; depth 18 m , 23.V.1988. brownish IX: Banc d ' A r g u i n ; 19°50"30"N 17°34'30"W; litto- chaetes; V a n V e e n grab; 8.vi.l988. yellow sand, hermit crabs, poly- ral; 23.V.1988. M.013: O f f coast; 18°50'N 16°20'W; depth 20 m , V : Banc d ' A r g u i n ; 19°47'30"N 17°32'30"W; litto- coarse y e l l o w sand w i t h shell gravel; V a n V e e n ral; 23.V.1988. grab; 8.vi.l988. V I : Banc d ' A r g u i n ; 19°48'N 17°33'W; littoral; 23.V.1988. M.014: Off coast; 18°50'N 16°21'W; depth 21 m , b r o w n i s h yellow, somewhat m u d d y sand; V a n VII: Banc d ' A r g u i n ; 19°49'N 17°33'30"W; littoral; V e e n grab; 8.vi.l988. 23.V.1988. M.015: Off coast; 18°50'N 16°21'W; d e p t h 21 m , VIII: Banc d ' A r g u i n ; 19°49'50"N 17°34'W; littoral; sandy bottom w i t h calcareous algae o n shells, 23.V.1988. sponges; 1.2 m A g a s s i z t r a w l ; 8.vi.l988. M.016: O f f Mauritania;18°51'N 16°21'W; depth 21 m, sandy bottom; 1.2 m Agassiz trawl; 8.vi.l988. Tyro Mauritania-II Expedition 1988 M.017: O f f coast; 18°50'N 16°19'W; depth 18 m , (material in N N M ) sandy bottom w i t h folious bryozoans, sponges; M.001: Baie de Cansado; 20°53'N 17°02'W; depth M.018: O f f coast; 18°50'N 16°22'W; d e p t h 26 m , 1.2 m A g a s s i z t r a w l ; 8.vi.l988. 8 m ; fishing lines; 5.vi.1988. sandy bottom w i t h shells, hermit-crabs, b r y o - M.002: Baie de Cansado; 20°53'N 17°02'W; depth zoans, sponges; 1.2 m A g a s s i z trawl; 8.vi.l988. 8 m , anoxic fine m u d ; V a n V e e n grab; 5.vi.l988. M.019: O f f coast; 18°50'N 16°23'W; depth 30 m , M.003: Baie de Cansado; 20°54'N 17°02'W; d e p t h sandy bottom w i t h sponges; 1.2 m A g a s s i z trawl; 0-6 m , o n shipwreck (partly above water); scuba 8.vi.l988. 54 Aartsen et al. P y r a m i d el l i d ae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) M.020: O f f coast; 18°50'N 16°24'W; depth 37 m , M.040: O f f coast; 18°51'N 16°53*W; depth 500 m , sandy bottom; 1.2 m A g a s s i z trawl; 8.vi. 1988. fossil coral debris, tubeworms; 3.5 m A g a s s i z M.021: O f f coast; 18°50*N 16°25'W; depth 37 m , trawl; 9.vi.l988. m u d d y sand w i t h Foraminifers, crabs, opitho- M.041: O f f coast; 18°51'N 16°56'W; depth 800-840 branchs, hermit-crabs; 1.2 m A g a s s i z trawl; m , m u d d y bottom; 3.5 m Agassiz trawl; 10.vi.1988. 8.vi.l988. M.042: Off coast; 18°03*N 16°58'W; depth 820-990 M.022: O f f coast; 18°50'N 16°28'W; depth 60-66 m , m u d d y bottom; 3.5 m Agassiz trawl; 10.vi.1988. m , m u d d y sand, crabs, echinoids, opithobranchs, M.043: O f f coast; 19°04'N 16°25'W; depth 15 m , hermit-crabs, sponges, flatfish (soleids); 1.2 m greyish sand w i t h some shell gravel; V a n V e e n Agassiz trawl; 8.vi.l988. grab; l l . v i . 1 9 8 8 . M.023: Off coast; 18°50'N 16°30'W; depth 71 m , M.044: O f f coast; 19°04'N 16°25'W; depth 18 m , small catch; 1.2 m A g a s s i z trawl; 8.vi.l988. m u d d y fine grey sand, some shell gravel; V a n M.024: O f f coast; 18°50*N 16°18'W; depth 14 m ; 3 V e e n grab; l l . v i . 1 9 8 8 . baited shrimp-traps, overnight; 8/9.vi.l988. M.045: O f f coast; 19°04'N 16°26'W; depth 22 m , M.025: O f f coast; 18°50'N 16°18'W; depth 14 m ; deep-sea camera plus video-camera (ship at anchor); 8.vi.l988. m u d d y fine grey sand, shell gravel; V a n V e e n grab; l l . v i . 1 9 8 8 . M.046: O f f coast; 19°04'N 16°27'W; depth 25 m , M.026: O f f coast; 18°50'N 16°18'W; depth 14 m ; m u d d y fine sand a n d shell gravel; V a n V e e n line fishing; 8.vi.l988. grab; l l . v i . 1 9 8 8 . M.027: Off coast; 18°50'N 16°18'W; depth 16 m , M.047: O f f coast; 19°04'N 16°27'W; depth 25 m , hard bottom w i t h sand, h y d r o i d s and soft corals; hermit-crabs w i t h epizoans, octopods, 1.2 m A g a s s i z trawl; 9.vi.l988. pods; 2.4 m Agassiz trawl; l l . v i . 1 9 8 8 . gastro- M.028: O f f coast; 18°50'N 16°20'W; depth 20 m , M.048: O f f coast; 19°04'N 16°25'W; depth 24 m , sandy bottom, soft corals, bryozoans, crabs; 1.2 flatfish, hermit-crabs, sponges; 2.4 m A g a s s i z m A g a s s i z trawl; 9.vi.l988. trawl; l l . v i . 1 9 8 8 . M.029: O f f coast; 18°49'N 16°22'W; depth 23 m , M.049: O f f coast; 19°05'N 16°25'W; depth 12-18 greyish-yellow sand w i t h few animals; V a n V e e n m , sand and sandstone, red algae, gorgonians, grab; 9.vi.l988. etc; rectangular dredge; l l . v i . 1 9 8 8 . M.030: O f f coast; 18°49'N 16°24'W; depth 36 m , M.050: O f f coast; 19°05'N 16°25'W; depth 20 m , m u d d y grey sand, tubeworms macro-foramini- shells, gorgonians, hermit-crabs, some red algae; fers; V a n V e e n grab; 9.vi.l988. 2.4 m A g a s s i z trawl; l l . v i . 1 9 8 8 . M.031: O f f coast; 18°48'N 16°28'W; depth 70 m , M.051: O f f coast; 19°04'N 16°25'W; depth 15 m , m u d d y dark-grey sand w i t h tubeworms forami- sandy bottom a n d sandstone ridge; scuba d i v - nifer; V a n V e e n grab; 9.vi.l988. i n g , underwater photography; 11 / 12.vi.1988. M.032: O f f coast; 18°47'N 16°34'W; depth 113 m , M.052: O f f coast; 19°04*N 16°25'W; depth 18 m ; greyish-yellow sand; V a n V e e n grab; 9.vi.l988. rectangular dredge; l l . v i . 1 9 8 8 . M.033: O f f coast; 18°47'N 16°34'W; depth 114 m , M.053: O f f coast; 19°05'N 16°28'W; depth 30 m , m u d d y sand a n d fine shell gravel; V a n V e e n many grab; 9.vi.l988. ll.vi.1988. M.034: O f f coast; 18°46*N 16°40'W; depth 167 m , M.054: O f f coast; 19°05'N 16°29'W; depth 30 m , small flatfish; 2.4 m Agassiz trawl; m u d d y fine sand, some shell gravel; V a n V e e n small flatfish (soleids), gastropods; 2.4 m A g a s - grab; 9.VÏ.1988. siz t r a w l ; l l . v i . 1 9 8 8 . M.026: O f f coast; 18°50'N 16°18'W; depth 14 m ; M.055: O f f coast; 19°05'N 16°25*W; depth 15 m ; line fishing; 8.vi.l988. line fishing; l l . v i . 1 9 8 8 . M.036: O f f coast; 18°45'N 16°42'W; depth 203 m ; M.056: O f f coast; 19°05'N 16°25'W; depth 15 m ; 5 5 shrimp-traps, overnight; 9/10.vi.l988. s h r i m p traps, overnight; 11 /12.vi.l988. M.037: O f f coast; 18°45'N 16°42'W; bottom at 200 M.057: Off coast; 19°05'N 16°25'W; depth 15 m ; m ; line fishing; 9.vi.l988. deep-see camera, underwater-video; l l . v i . 1 9 8 8 . M.038: O f f coast; 18°46'N 16°45*W; depth 260 m , M.058: O f f coast; 19°05'N 16°25'W; at surface; m u d d y bottom; 3.5 m A g a s s i z trawl; 9.vi. 1988. neuston net (30 minutes); 12.vi. 1988. M.039: O f f coast; 18°48'N 16°43'W; depth 260-280 M.059: O f f coast; 19°05'N 16°39'W; depth 110 m , m , m u d d y bottom, tubeworms; 3.5 m A g a s s i z small catch, hermit-crabs, flatfish; 2.4 m A g a s s i z trawl; 9.vi.l988. trawl; 12.vi.1988. 55 Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) M.060: Off coast; 19°06'N 16°46'W; depth 280-350 lar dredge; 14.vi.1988. m , rather steep continental slope, m a n y irregular M.077: off Banc d ' A r g u i n ; sea-urchins; 2.4 m Agassiz trawl; 12.vi.1988. depth 35 m , dark-grey m u d d y sand; V a n V e e n 20°00'N 17°17'W; M.061: Off coast; 19°09'N 16°52'W; depth 400-750 grab; 14.vi.1988. m , rather steep canyon slope, fishes, pennatul- M.078: off Banc d ' A r g u i n ; ids, small gastropods; 2.4 depth 41 m , dark-grey m u d d y fine sand; V a n m Agassiz trawl; 20°00'N 17°21'W; 12. vi.1988. V e e n grab; 14.vi.1988. M.062: Off coast; 19°09'N 16°52'W; depth 800- M.079: off Banc 1200 m , steep slope, only some pelagic fauna; depth 53 m , dark-grey m u d d y fine sand, tube- failure (net d i d not touch bottom), 2.4 m A g a s - worms, siz-trawl; 12.vi.1988. 14.vi.1988. M.063: off Banc d ' A r g u i n ; 20°00'N 17°09'W; depth 20 m , h a r d bottom w i t h some m u d d y d'Arguin; ophiurids, Tellina; M.080: off Banc d ' A r g u i n ; 20°01'N V a n V e e n grab; 20°02'N 3.5 m Agassiz trawl; 14.vi.1988. M.064: off Banc d ' A r g u i n ; M.081: off Banc 17°11'W; depth 24 m , m u d d y sand w i t h C a r d i u m and Lanice; V a n V e e n grab; 13.vi.1988. M.065: off Banc d'Arguin; 20°00'N 17°26'W; depth 60-70 m , gastropods, hermit-crabs, flatfish; sand, gorgonians; V a n V e e n grab; 13.vi.1988. 20°00'N 17°23'W; d'Arguin; 19°59'N 17°27'W; depth 120 m , failure: gear lost; 3.5 m Agassiz trawl; 14.vi.1988. 17°11'W; M.082: off Banc d ' A r g u i n ; 19°59'N 17°30'W; depth 25 m , m u d d y sand w i t h C a r d i u m and depth 100 m ; 2.4 m Agassiz trawl; 14.vi.1988. Lanice; V a n V e e n grab; 13.vi.1988. M.083: off Banc d ' A r g u i n ; M.066: off Banc d'Arguin; 20°00'N 17°11'W; depth 25 m , m u d d y sand w i t h C a r d i u m and trawl; 14.vi.1988. Lanice; 3.5 m Agassiz trawl; 13.vi.1988. M.084: off Banc d ' A r g u i n ; M.067: off Banc d'Arguin; 20°00'N 17°12'W; grab; 14.vi.1988. m Agassiz trawl; 13.vi.1988. M.085: off Banc off Banc d'Arguin; 20°01'N 17°15'W; ophiurids, 13. vi.1988. 15.vi.1988. 20°00'N 17°17'W; 19°35'N 16°50'W; d'Arguin; 19°35'N 16°51'W; depth 35 m , light grey sandy m u d w i t h shells, depth 30 m , m u d d y sand; 3.5 m Agassiz trawl; M.069: off Banc d ' A r g u i n ; 17°42'W; depth 19 m , light-grey sandy m u d ; V a n V e e n depth 28 m , m u d d y sand w i t h C a r d i u m , etc.; 3.5 M.068: 20°00'N depth 720-760 m , failure: gear lost; 2.4 m Agassiz small M.086: off Banc bivalves; V a n V e e n d'Arguin; 19°32'N grab; 16°52'W; depth 32 m , m u d d y sand w i t h small bivalves; 3.5 depth 52 m , sticky grey m u d w i t h shell gravel, m Agassiz trawl; 13.vi.1988. ophiuroids, M.070: off Banc d ' A r g u i n ; depth 38-41 20°00'N 17°18'W; m , m u d d y sand; 3.5 m Agassiz trawl; 13.vi.1988. small bivalves; V a n V e e n grab; 15.vi.1988. M.087: off Banc d ' A r g u i n ; 19°32'N 16°54'W; depth 65 m , sticky grey m u d w i t h shells, ophiur- M.071: off Banc d ' A r g u i n ; depth 43 m , m u d d y 20°00'N 17°21'W; sand w i t h tubeworms, oids, small bivalves; V a n V e e n grab; 15.vi.1988. M.088: off Banc d ' A r g u i n ; 19°33'N 16°55'W; crabs, flatfish; 3.5 m Agassiz trawl; 13.vi.1988. depth 64 m , sticky grey m u d w i t h shells, ophiur- M.072: off Banc d ' A r g u i n ; oids, echinoids, bivalves, tubeworms; 1.2 20°00'N 17°24'W; depth 48-52 m , m u d d y sand; 3.5 m Agassiz Agassiz trawl; 15.vi.1988. trawl; 13.vi.1988. M.089: off Banc d ' A r g u i n ; M.073: off Banc d'Arguin; 20°03'N 17°09'W; 19°34'N w o r m s ; 1.2 m Agassiz trawl; 15.vi.1988. siz trawl; 13.vi.1988. M.090: off Banc d ' A r g u i n ; d'Arguin; 20°02'N 17°09'W; 19°35'N depth 22 m , hermit-crabs w i t h bryozoans; 2.4 m fishes; 1.2 m Agassiz trawl; 15.vi.1988. M.091: off Banc off Banc d'Arguin; 20°01'N 17°09'W; d'Arguin; depth 22 m , L u i d i a , hermit-crabs w i t h bryo- trawl; 15.vi.1988. M.092: off Banc d ' A r g u i n ; crabs, small catch; rectangular dredge; 14.vi.1988. M.076: off Banc d ' A r g u i n ; 19°35'N 16°50'W; depth 30 m , very small catch; 1.2 m Agassiz zoans, some 16°51'W; depth 38 m , ophiuroids, echinoids, bivalves, Agassiz trawl; 13.vi.1988. M.075: 16°55'W; depth 62 m , ophiuroids, Ebalia, bivalves, tube- depth 19 m , hard calcareous bottom; 1.2 m A g a s M.074: off Banc m 19°35'N 16°50'W; depth 32 m , very small catch; 1.2 m Agassiz 20°00'N 17°10'W; depth 24 m , tubeworms, hermit-crabs; rectangu- trawl; 15.vi.1988. M.093: off Banc d'Arguin; 19°35'N 16°57'W; 56 Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) depth 85-90 m , echinoids, small bivalves, gastro- M.110: off Banc d ' A r g u i n ; pods, depth 22 m , shell gravel; 2.4 m Agassiz trawl; tubeworms; 1.2 m Agassiz trawl; 15. vi.1988. 20°30'N 17°05'W; 18.vi.1988. M.094: off Banc d ' A r g u i n ; 19°32'N 17°01'W; depth 110 m ; 1.2 m Agassiz trawl; 15.vi.1988. M.095: off Banc d ' A r g u i n ; 19°31'N 17°01'W; M.lll: off Banc d ' A r g u i n ; 20°31'N 17°09'W; depth 29 m , coarse sand, somewhat m u d d y , a n d shell gravel; V a n V e e n grab; 18.vi.1988. depth 200-110 m , small catch, only hermit-crabs; M.112: of Banc d ' A r g u i n ; 20°31'N 17°10'W; depth rectangular dredge; 15.vi. 1988. 29 m , shell gravel, hermit-crabs, echinoids; 2.4 m M.096: off Banc d ' A r g u i n ; 19°33'N 17°02'W; Agassiz trawl; 18.vi.1988. depth 200-110 m ; 1.2 m Agassiz trawl; 16.vi.1988. M.113: off Banc d ' A r g u i n ; M.097: off Banc d ' A r g u i n ; depth 36 m , coarse sand a n d shell gravel; V a n 19°24'N 16°48'W; depth 26 m m u d d y sand w i t h some shell gravel, V e e n grab; 18.vi.1988. ophiuroids; V a n V e e n grab; 16.vi.1988. M.114: off Banc d ' A r g u i n ; M.098: off Banc d ' A r g u i n ; 19°25'N 16°48'W; depth 26 m , m u d d y sand, gorgonians, hermitcrabs, tubeworms; 2.4 m Agassiz trawl; 16. vi.1988. M.099: off Banc d ' A r g u i n ; 20°30'N 20°29'N 17°13'W; 17°14'W; depth 36 m , shell gravel; 2.4 m Agassiz trawl; 18.vi.1988. M.115: off Banc d ' A r g u i n ; 20°28'N 17°17'W; depth 38 m , m u d d y sand w i t h fine shell gravel 19°25'N 16°49'W; a n d tubeworms; V a n V e e n grab; 18.vi.1988. depth 42 m , dark-grey sandy m u d w i t h shell M.116: off Banc gravel tubeworms; V a n depth 38 m , shell gravel, tubeworms, hermit- 19°26'N M.117: off Banc d ' A r g u i n ; (Pinna), ophiuroids, Veen grab; 16.vi.1988. M.100: off Banc d ' A r g u i n ; d'Arguin; trawl; 18.vi.1988. 1988. M.118: off Banc d ' A r g u i n ; 19°43'N 16°59'W; depth 61-78 m , sticky m u d w i t h some sand a n d 20°24'N 17°19'W; depth 35-40 m , sandy bottom; 3.5 m Agassiz fers, gastropods, etc.; 2.4 m Agassiz trawl; 16.vi. off Banc d ' A r g u i n ; 17°18'W; crabs; 2.4 m Agassiz trawl; 18.vi.1988. 16°50'W; depth 45 m , m u d d y sand w i t h macro-foramini- M.101: 20°26'N 28°25'N 17°04'W; depth 17 m , y e l l o w sand a n d shell gravel; V a n V e e n grab; 19.vi.1988. shell gravel; 2.4 m Agassiz trawl; 16.vi.1988. M.119: off Banc M.102: depth 17 m , practically no catch; 2.4 m Agassiz off Banc d ' A r g u i n ; 19°45'N 17°05'W; d'Arguin; depth 24 m , m u d d y bottom w i t h dense p o p u l a - trawl; 19.vi.1988. tion; 2.4 m Agassiz trawl; 16.vi.1988. M.120: off Banc d ' A r g u i n ; M.103: off Banc d ' A r g u i n ; 19°46'N 20°25'N 20°24'N 17°06'W; 17°08'W; 17°06'W; depth 22 m , shell gravel w i t h m a n y V e n u s shells, depth 400-100 m , steep slope of canyon, tube- few small fish, echinoids; 3.5 m Agassiz trawl; w o r m s , scleractinians, ophiuroids; rectangular 19.vi.1988. dredge; 16.vi.1988. M.121: off Banc d ' A r g u i n ; M.104: off Banc d ' A r g u i n ; 19°43'N 17°30'W; 20°27'N depth 1500 m , m a n y fish etc.; 3.5 m Agassiz what m u d d y ; V a n V e e n grab; 19.vi.1988. trawl; 17.vi.1988. M.122: off Banc d ' A r g u i n ; M.105: off Banc d ' A r g u i n ; 19°43'N 17°44'W; 17°14'W; depth 31 m , fine sand w i t h shell gravel, some20°27'N 17°15'W; depth 32 m , shell gravel; 3.5 m Agassiz trawl; depth 1600-1900 m , m a n y holothurians a n d o p h - 19.vi.1988. iuroids; 3.5 m Agassiz trawl; 17.vi.1988. M.123: off Banc d ' A r g u i n ; 20°26'N 17°20'W; sur- M.106: off Banc d ' A r g u i n ; 20°31'N 17°02'W; face (bottom at 36 m); neuston net (during the depth 15 m , shell gravel; V a n V e e n grab (4x); day); 19.vi.1988. 18.vi.1988. M.124: off Banc d ' A r g u i n ; M.107: off Banc d ' A r g u i n ; 20°31'N 17°03'W; depth 18 m , very small catch (Pyura, crab); 2.4 m 19.vi.1988. Agassiz trawl; 18.vi.1988. M.125: off Banc d ' A r g u i n ; M.108: off Banc d ' A r g u i n ; 20°31'N 17°04'W; depth 20 m , small catch, shell gravel; 2.4 m calcareous 19.vi.1988. 20°31'N 17°05'W; 17°21'W; 20°25'N 17°26'W; depth 47 m , sandy bottom w i t h some shell- a n d Agassiz trawl; 18.vi.1988. M.109: off Banc d ' A r g u i n ; 20°26'N depth 37 m , m u d d y sand; 3.5 m Agassiz trawl; gravel; 3.5 m M.126: off Banc d ' A r g u i n ; Agassiz 20°24'N trawl; 17°30'W; depth 22 m , m u d d y sand w i t h shell gravel; V a n depth 55-60 m , fine sediment, flatfish, hermit- Veen grab; 18.vi.1988. crabs, etc.; 3.5 m Agassiz trawl; 19.vi.1988. Aartsen et al. Pyramidellidae of the N o r t h Atlantic Ocean. Z o o l . V e r h . L e i d e n 321 (1998) M.127: off Banc d'Arguin; 20°24'N 17°34'W; 57 M.136: off C a p Blanc; 20°43'N 17°21'W; depth 50 depth 62-75 m , fine sediment, many fishes, crabs, m ; deep-sea camera (lOx); 21.vi.1988. gastropods, etc; 3.5 m Agassiz trawl; 19.vi.1988. M.137: off C a p Blanc; 20°43'N 17°23'W; depth 50 M.128: off Banc d ' A r g u i n ; 20°25'N 17°35'W; sur- m , flatfish, hermit-crabs, octopus, etc.; 2.4 m face (bottom at 75 m); neuston net (during the Agassiz trawl; 21.vi.1988. night); 19.vi.1988. M.138: off C a p Blanc; 20°44'N 17°25*W; depth 54- M.129: depth off Banc d'Arguin; 20°25'N 17°40'W; 140 m , failure; V a n V e e n grab (4x); 20.vi.1988. M.130: off Banc 62 m ; 3.5 m Agassiz trawl; 21.vi.1988. M.139: off C a p Blanc; 20°42'N 17°30'W; depth 6371 m , Xenophora, flatfish, hermit-crabs, etc.; 3.5 d'Arguin; 20°25'N 17°40'W; m A g a s s i z trawl; 21.vi.1988. depth 95-100 m , m u d d y sand; 3.5 m Agassiz M.140: off C a p Blanc; 20°40'N 17°31'W; depth 73 trawl; 20.vi. 1988. m , m u d d y sand w i t h some shell gravel; H a m o n M.131: off Banc d'Arguin; 20°29'N 17°41'W; grab; 21.vi.1988. depth 225-235 m ; 3.5 m Agassiz trawl; 20.vi.1988. M.141: off C a p Blanc; 20°40'N 17°31'W; depth 72 M.132: off C a p Blanc; 20°34'N 17°45'W; depth m ; deep-sea camera (lOx); 21.vi.1988. 305-325 m ; 3.5 m Agassiz trawl; 20.vi.1988. M.142: off C a p Blanc; 20°39'N 17°36'W; depth 75- M.133: off C a p Blanc; 20°39'N 17°48'W; depth 100 m , Xenophora, flatfish, octopus, etc.; 3.5 m 400-450 m ; 3.5 m Agassiz trawl; 20.vi.1988. Agassiz trawl; 21.vi.1988. M.134: off C a p Blanc; 20°44*N 17°48'W; depth M.143: off C a p Blanc; 20°40'N 17°31'W; depth 530-700 m , m a i n l y fish, shrimp, small asteroids, 150-175 m , net torn, only small catch, mostly cri- tubeworms; 3.5 m Agassiz trawl; 20.vi.1988. noids; 3.5 m Agassiz trawl; 21.vi.1988. M.135: off C a p Blanc; 20°42'N 17°21'W; depth 50 M.144: off C a p Blanc; 20°43'N 17°39'W; depth m , m u d d y , dark-grey fine sand, little shell grav- 200-210 m ; 2.4 m A g a s s i z trawl; 21.vi.1988. el; v a n V e e n grab; 21.vi.1988.