Notes on Early Land Plants Today. 73. Genera of Lejeuneaceae established in the period 1884–1893: dates of validation and implications

Matt Von Konrat

Dates of validation of generic names initially published as names in subgenera or sections of Lejeunea (mainly by Spruce) in the period 1884–1893 are established. Consequences for the dates of validations of specific names are analysed. Names not validly published (both generic and specific names) in this period are also identified. A few species names must be changed from those in common use today and two new combinations, Rectolejeunea versifolia and Lejeunea hyalina, become necessary.

Phytotaxa 220 (2): 143–198 www.mapress.com/phytotaxa/ Copyright © 2015 Magnolia Press ISSN 1179-3155 (print edition) Article PHYTOTAXA ISSN 1179-3163 (online edition) http://dx.doi.org/10.11646/phytotaxa.220.2.4 Notes on Early Land Plants Today. 73. Genera of Lejeuneaceae established in the period 1884–1893: dates of validation and implications LARS SÖDERSTRÖM1, FRED R. BARRIE2,3, ANDERS HAGBORG2, BARBARA J. CRANDALL-STOTLER4, S. ROBBERT GRADSTEIN5, RAYMOND E. STOTLER†4 & MATT VON KONRAT2 1 Department of Biology, Norwegian University of Science and Technology, N-7491 Trondheim, Norway; lars.soderstrom@ntnu.no Department of Science and Education, The Field Museum, 1400 South Lake Shore Drive, Chicago, Illinois 60605–2496, U.S.A.; fbarrie@fieldmuseum.org, hagborg@pobox.com, mvonkonrat@fieldmuseum.org 3 Missouri Botanical Garden, P. O. Box 299, St. Louis, Missouri 63166, U.S.A. 4 Department of Plant Biology, Southern Illinois University, Carbondale, Illinois 62901-6509, U.S.A.; crandall@siu.edu 5 Muséum National d’Histoire Naturelle, Department Systématique et Evolution, C.P. 39, 57 Rue Cuvier, 75231 Paris 05, France; gradstein@mnhn.fr 2 Abstract Dates of validation of generic names initially published as names in subgenera or sections of Lejeunea (mainly by Spruce) in the period 1884–1893 are established. Consequences for the dates of validations of specific names are analysed. Names not validly published (both generic and specific names) in this period are also identified. A few species names must be changed from those in common use today and two new combinations, Rectolejeunea versifolia and Lejeunea hyalina, become necessary. Introduction Taxonomic recognition of well-defined segregates within the broadly defined genus Lejeunea Libert (1820: 373) began in large part with the monumental treatment of the liverworts of the Amazon and Andes by Spruce (1884). In that work he divided the genus into 38 subgenera distributed between two groups, the Holostipae and the Schizostipae, applying to them names formed by the addition of a descriptive prefix hyphenated to the generic name Lejeunea. In an epithet, e.g. in the name of a subgenus, the inclusion of the hyphen is now treated as an error to be corrected by deletion of the hyphen (ICN [Melbourne Code] Art. 60.9, McNeill et al. 2012). One additional subgenus, Lejeunea subg. Anomalolejeunea, was named by Spruce in Pearson (1887). In addition, Stephani (1890a: 19) described a further “Gruppe” that he called “Cauda-Lejeunea” and treated at a rank commensurate with Spruce’s subgenera. However, he failed explicitly to connect “Cauda-Lejeunea” with a generic name (cf. ICN Art. 35.2) so this is not a validly published name of a generic subdivision. All 39 subgenera named by Spruce, as well as Stephani’s “Caudalejeunea” and Spruce’s “§ 3. Strepsi-Lejeunea”, one of three groups prefixed by ‘§’ and subordinate to Lejeunea subg. HarpaLejeunea (Spruce 1884: 168), but listed as a subgenus on p. 585 and its two species presented in subgeneric form on p. 179 (e.g. “Strepsi-Lejeunea inflexa”) of Spruce (1884) were recognized as genera by subsequent authors and have since been typified. However, who should be credited with the first publication of each of these as generic names and when these names, as well as those of certain species, were validly published, has been equivocal for more than a century. Schiffner (1893b) treated 31 of Spruce’s explicit Lejeunea subgenera, his Lejeunea ‘§ Strepsi-Lejeunea’ and Caudalejeunea that he attributed to Stephani as distinct genera and has generally been regarded as the author of each (see Grolle 1983). He considered Lejeunea subg. Microlejeunea (‘Micro-Lejeunea’) to be a subgenus of Eulejeunea. He treated the subgenera that Spruce referred to as Bryo-Lejeunea, Omphalo-Lejeunea, Ptycho-Lejeunea, ThysanoLejeunea and Homalo-Lejeunea under their original (pre-Sprucean) names—i.e., Bryopteris, Omphalanthus, Ptychanthus, Thysananthus and Marchesinia, respectively. These five Sprucean subgenera along with L. subg. Dendrolejeunea (‘Dendro-Lejeunea’) and L. subg. Potamolejeunea (‘Potamo-Lejeunea’) were later recognized at the generic rank by Lacouture (1908). Bonner et al. (1961) argued that Stephani should be considered the author for 32 Accepted by Christine Cargill: 30 May 2015; published: 20 Jul. 2015 Licensed under a Creative Commons Attribution License http://creativecommons.org/licenses/by/3.0 143 of these names, based on either the description of a single new species, thus validating the generic and specific names with a single description (ICN Art. 38.5), or by a clear reference to the previously and effectively published description of the subgenus by Spruce (currently ICN Art. 38.14). However, for either of these scenarios to apply, Stephani must have unambiguously treated the taxon as a genus, rather than a subgenus. Unfortunately, the application of the terms “genus” and “subgenus” by Stephani was very inconsistent. For example, in his “Westindische Hepaticae” (Stephani 1888c: 287) he referred to Cololejeunea as a “Subgenus” and in the same paragraph as a genus (“Gattung”). In that same work (l.c: 289), following his binomial treatment “Pycnolejeunea Schwaneckei St. n. sp.” he referred to the figure on Tab. XIII as “Lejeunea Schwaneckei”. After reviewing comparable inconsistencies in the publications by Stephani between 1888 and 1894, Zijlstra (1982) concluded that Stephani generally considered the Sprucean segregates as subgenera, despite sometimes referring to them as genera. Many of Stephani’s genus names predate Schiffner (1893b), to whom authorship of these names has often been ascribed. Schuster (1980a: 711) suggested solving the problem by arbitrarily conserving these names as genera with Spruce as author, a solution recommended earlier by Verdoorn (1934b), and formally proposed to the ICBN by Scott & Zijlstra (1986). This, of course, is not a permissible solution from a nomenclatural point of view, as affirmed by rejection of the proposal at the XIV International Botanical Congress in Berlin in 1987. Only names validly published at the rank involved may be conserved, i.e., a name published as a subgenus cannot be conserved as a genus. Moreover, Spruce (1884: 73) made it clear that he never intended elevating his Lejeunea segregates to generic rank and maintained them as subgenera in all his later publications. Earlier discussions of Spruce’s subgenera have been centered on the possibility that some generic names have been created through a combined generic and specific description (ICN Art. 38.5) (e.g. Gradstein et al. 1982). However, there is also a possibility that 1) there might be a clear (but sometimes “cryptic”) reference to Spruce (1884) to justify a combination or 2) there is enough “description” of a generic name (not to be confused with the description of specific names). We think very little is needed to add to the issue of combined generic and specific description. The view that there might be a clear (but sometimes “cryptic”) reference to Spruce to create a new combination has been controversial, primarily because of the issue of rank at which the name is being used. Bonner et al. (1961) accepted it for the Stephani (1890a, b, c) papers and Grolle (1983) indicated that there may be enough reference for two taxa in Stephani (1889b) for such a treatment. These views were rejected by Zijlstra (1982) since she concluded that the names were being used by Stephani at the subgeneric rank. This seemingly irresolvable conflict of views has led us to propose that some of Stephani’s works should be suppressed (ICN Art. 34) at the rank of genus. The possibility that there is enough “description” of a generic name in any of the 1884–1893 publications has not been discussed in any earlier publication. There is some evidence that this might be the case in one publication (Stephani 1888c), but the interpretation of the rank of the taxon being described is controversial. This is one of the publications we propose be suppressed at the rank of genus. According to ICN Art. 41.4, a provision first added, as Art. 33.2, to the St Louis Code (Greuter et al. 2000), if conditions for valid publication of the name of a new genus are fulfilled, the name can be treated as a new combination or name at a new rank when this was the author’s presumed intent, even without direct reference to a basionym. An example is Taxilejeunea (Spruce) Stephani (1889b: 262). Stephani published the name Taxilejeunea convexa Stephani (1889b: 282) for a novel species, providing a detailed description, but failed to supply any reference to Lejeunea subg. Taxilejeunea Spruce (1884: 212), the presumed basionym for the genus name. The name satisfies the provisions of ICN Art. 38.5 for a combined generic-specific description because the genus at that time was monotypic and no other name had previously been published based on the same type. However, as it clearly was Stephani’s presumed intent to base the generic name on one of Spruce’s subgeneric names, ICN Art. 41.4 is applicable, as, if the name is not so treated, it would otherwise be the name of a new genus (J. McNeill, pers. comm.). As a consequence, the name Taxilejeunea is typified by the Sprucean type (Lejeunea pterigonia (Lehm. et Lindenb. in Lehmann 1834: 44) Montagne (1840: 337) [≡ Taxilejeunea pterigonia (Lehm. et Lindenb.) Jack & Stephani (1892: 13)]), not by the type of the name of the species described by Stephani. Prior to what is now ICN Art. 41.4 being added to the Code, Taxilejeunea (Stephani 1889b: 362) was arguably not a combination but a new name at the rank of genus, validly published by a combined generic-specific description, the type of which was Taxilejeunea convexa Stephani (1889b: 362). As such, the name had priority over a combination based on Spruce’s name and type, Taxilejeunea (Spruce) Schiffner (1893b: 125). On this basis, Gradstein et al. (1982) proposed conservation of Taxilejeunea (Spruce) Schiffn. and three other names, Acrolejeunea (Spruce) Schiffn., Lopholejeunea (Spruce) Schiffn. and Trachylejeunea (Spruce) Schiffn., over the putatively heterotypic Stephani homonyms. Conservation of all four was approved by the Tokyo Congress and Schiffner’s combinations were added to App. III in the Berlin Code (Greuter et al. 1988). The addition of ICN Art. 41.4, however, obliges the acceptance of 144 • Phytotaxa 220 (2) © 2015 Magnolia Press SÖDERSTRÖM ET AL. Stephani as the combining author, the names dating from his publications. As a consequence, the entries in Appendix III of ICN for the four generic names that were conserved in 1987 (Acrolejeunea, Lopholejeunea, Taxilejeunea, Trachylejeunea) are now changed and ascribed to “(Spruce) Steph.”, not “(Spruce) Schiffn.” (cf. also ICN Art. 14.4, Note 2). These names are typified by Sprucean types following ICN Art. 41.4, not by the types of the names of the species described by Stephani (Wiersema et al. 2015). Example 9 of ICN Art. 41 cites Brachiolejeunea as a name validly published by Stephani (1889a), based on a combined generic-specific description. This particular example is flawed, however, as Stephani (1889a) included a list of species names under Brachiolejeunea that he said belonged to the genus, several of which would be new, validly published combinations if the name of the genus had been validly published here. Because Brachiolejeunea does not represent a case where the conditions of ICN Art. 38.5 are met, this example will be deleted from the next edition of the ICN (W. Greuter & J. McNeill, pers. comm.). There are several, similar cases of other generic names commonly regarded as validly published by a single combined generic-specific description (cf. Grolle 1983) that are actually validly published in later publications. The purpose of this paper is to identify where, when and by whom the Lejeunea subgenera of Spruce (1884) were established as genera and to identify which specific names (and when) have been validly described/combined in those genera. Some Stephani species were described before the generic names in which they were placed were validly published. Over the years, the names of such species have been treated as validly published, as validly published binomials in Lejeunea, or as not validly published, creating confusion that today still causes well-known taxa to be referred to by incorrect citations. This paper deals with the nomenclature of all the names that were originally described as subgenera (or sections) of Lejeunea between 1884 and October 1893. Consequences for the dates of valid publication of binomials at species rank are also analysed. Publications containing Lejeuneaceae taxa 1884–October 1893 We have not found any publication earlier than 1888 in which effectively (validly or not according to the Code) published names in “Xxxlejeunea” (= collective term for Lejeunea subgenera being treated at genus level) are proposed, either intentionally or accidentally. Papers published in the period 1888–1893 by Stephani are often difficult to interpret in this respect due to variable typography, sometimes in the same paper, and seemingly random use of the terms genus (“Gattung”) and subgenus (“Untergattung”) (see also Bonner et al. 1961 and Zijlstra 1982). In Stephani (1890b) he even confused the Latin and German terms for the same thing (“Die Anzahl der neuen Species ist gering; noch weniger zahlreich sind die Arten”). Other authors are generally more clear and consistent. Bescherelle consistently treated the Sprucean subgenera as subgenera in all his publications except his paper from 1892 (which probably was written by Stephani). Spruce also clearly treated his subgenera as such, as did Schiffner until his 1893 treatment. As it is not possible to find clear “internal evidence” that Stephani was treating his taxa as Lejeunea species the only possible interpretation is to take what he writes literally. Thus, his “Xxxlejeunea” must often be taken as generic names. In addition, it is not possible to treat his combinations as Lejeunea combinations when he is not explicitly connecting the genus name to the species (cf. ICN Art. 35.2). This has led us to the conclusion that, to avoid further confusion, the most confusing papers are best proposed for addition to Appendix VI (opera utique oppressa; cf. ICN Art. 34). Details, with evidence for effective publication of names and whether they are fulfilling the requirements for valid publication or not are given below. We give the ICN Article whenever needed except for ICN Art. 35.1 (generic name not validly published in the publication or earlier) which concerns the majority of the cases for species names not validly published in those publications. The names in the indices are direct references to the effectively published descriptions in earlier or current issues, so they can be taken as new combinations or validating names under Lejeunea, when such is missing. We have tried to identify such cases here. In several publications only one new species is described by Stephani and, in addition, one or more taxa are mentioned as belonging to the genus, as in the case of Brachiolejeunea (Stephani 1889a: 167), given as Example 9 of ICN Art. 41. Brachiolejeunea plagiochiloides is the only new species described, but Stephani further indicates that nine species of Phragmicoma Dumortier (1822: 112) and five species of “Brachio–L.” also belong to the genus. Thus, the example is in conflict with the ICN Art. 38.5 and will be removed from the Code (see above). We have also tried to establish when each paper was issued. Bonner et al. (1961) gave dates for most publications, but they seem to have used the dates on the printed issue, not the effective publication dates. TL-2 (Stafleu & Cowan GENERA OF LEJEUNEACEAE ESTABLISHED IN 1884–1893 Phytotaxa 220 (2) © 2015 Magnolia Press • 145 1976–2009) has more accurate information on the effective publication dates and we are following them whenever possible. For each publication we first summarize the nomenclatural impact before we list all new effectively published names, whether validly published or not. In the list of papers containing names in Lejeuneaceae from the period concerned, we enumerate all generic and binomials at specific rank that we consider validly published, as well as those that we consider to be not validly published names. We present all generic names in bold but only validly published generic names in italic. Stephani (1888a; Feb.) is the first of two papers on African hepatics. It uses the “Sprucean” subgenera without hyphens in a seemingly generic form. There is no reference, direct or indirect to Spruce (1884), so no new combinations are made and there is nothing that can be taken as a description of a genus. The index indicates that new binomial species names are intended under “Xxxlejeunea” by the typography. Moreover, Lejeunea entries for these names are lacking. Zijlstra (1982) and Bonner et al. (1961) considered that three generic names were established in this and the following publication. They did not regard Eulejeunea as established here since Spruce’s subgeneric name was not validly published (ICN Art. 21.3 and 22.1; type of genus included) as Spruce (1884) used the name subg. Eu-Lejeunea for what should have been the autonym. However, we consider the genus established on its own merits here. It is not a change in rank; the name is first proposed here, validly published by a combined generic-specific description. One additional generic name is also established by combined generic-specific descriptions (ICN Art. 38.5) and thus are also the two binomial specific names validly published here. Names validly published:—Eulejeunea Stephani (1888a: 60), E. hepaticola Stephani (1888a: 60), Microlejeunea (Spruce 1884: 286) Stephani (1888a: 61), M. africana Stephani (1888a: 61). Names not validly published:—None. Stephani (1888b; Mar–Apr) is the second paper on African hepatics. Typography and index are the same as in the previous paper except that hyphens are used in the “Xxx-Lejeunea” names in this publication. No reference to Spruce (1884) or any possible description of any genus exists. Two generic names (Archilejeunea and Homalolejeunea) are established here by combined generic-specific descriptions, with which Bonner et al. (1961) and Zijlstra (1982) also agreed. In addition three binomials at specific rank in earlier validly published generic names are validly published here. Three published generic names and six binomials at specific rank were not validly published here. Names validly published:—Archilejeunea Stephani (Spruce 1884: 88) Stephani (1888b: 113), A. erronea Stephani (1888b: 113), Eulejeunea flava (Swartz 1788: 144) Stephani (1888b: 109), E. rodriguezii Stephani (1888b: 108), Microlejeunea cochlearifolia Stephani (1889b: 113), Homalolejeunea (Spruce 1884: 132) Stephani (1888b: 112), H. henriquesii Stephani (1888b: 112). Names not validly published:—Acrolejeunea (Spruce 1884: 115) Stephani (1888b: 107, 112), A. occulta Stephani (1888b: 112), A. renauldii Stephani (1888b: 107), Cheilolejeunea (Spruce 1884: 251) Stephani (1888b: 109), C. newtonii Stephani (1888b: 109), C. principensis Stephani (1888b: 109), Mastigolejeunea (Spruce 1884: 100) Stephani (1888b: 110, 111), M. buettneri Stephani (1888b: 110), M. crispula Stephani (1888b: 111). Stephani (1888c; Nov–Dec) is a paper on Caribbean hepatics where a large number of species are described or combined under generic names that are effectively but not yet validly published. On page 289 Stephani gives a brief but clear description of Cololejeunea (“Soweit ich die Arten von Cololejeunea kenne, haben sie alle die …”) and it may seem that he establishes the genus, but on page 287 he talks about Cololejeunea both as a subgenus and “Gattung”. There is also a paragraph that might be interpreted as a description of Ceratolejeunea on page 283 (“wesentliche Merkmale von Ceratolejeunea, dahin gehören…”) which could be sufficient to establish the generic name. Those two genera were not accepted as validly published here by Bonner et al. (1961) or Zijlstra (1982) because there is ambiguity regarding the rank at which the names Cololejeunea and Ceratolejeunea are being described. Neither Ceratolejeunea, nor Cololejeunea have been accepted as described here by any earlier author, so adding the publication to Appendix VI (ICN Art. 34), suppressing any new generic name from this publication, will have a stabilizing effect on the nomenclature. Binomials in specific rank under earlier validly published generic names can be regarded as validly published here. The index indicates that they should be treated as new species under the suggested “Xxxlejeunea” although most genera were not yet established. Lejeunea schwaneckei is validly published here as Stephani uses this name in the legend to the illustration, although he placed it under the name Pycnolejeunea in the description itself, a generic name not validly published at this time. Forty-three binomials at specific rank published under 20 generic names are effectively but not validly published new names or combinations since their generic names had not been validly published. Names validly published:—Eulejeunea glaucescens (Gottsche in Gottsche et al. 1845b: 378) Stephani (1888c: 286), E. muscicola (Spruce 1884: 281) Stephani (1888c: 300), E. urbanii Stephani (1888c: 301), Lejeunea schwaneckei Stephani (1888c: 290), Microlejeunea ovifolia Gottsche ex Stephani (1888c: 287). Names not validly published:—Bryolejeunea (Spruce 1884: 111) Stephani (1888c: 285), B. diffusa (Swartz 1788: 144) Stephani (1888c: 300), B. filicina (Swartz 1788: 145) Stephani (1888c: 285), Ceratolejeunea (Spruce 1884: 198) Stephani (1888c: 283), C. breutelii (Gottsche in Gottsche et al. 1845b: 324) Stephani (1888c: 283), C. ceratantha (Nees et Mont. in Montagne 1840: 335) Stephani 146 • Phytotaxa 220 (2) © 2015 Magnolia Press SÖDERSTRÖM ET AL. (1888c: 283), C. cornuta (Lindenberg 1829: 23) Stephani (1888c: 285), C. cubensis (Montagne 1842: 481) Stephani (1888c: 300), C. kegelii (Lindenberg & Gottsche 1851: 635) Stephani (1888c: 300), C. spinosa (Gottsche in Gottsche et al. 1845b: 402) Stephani (1888c: 291), C. variabilis (Lindenb. in Gottsche et al. 1845b: 399) Stephani (1888c: 293), Cheilolejeunea (Spruce 1884: 251) Stephani (1888b: 109), C. duriuscula (Nees in Gottsche et al. 1845b: 364) Stephani (1888c: 285, 300), C. lineata (Lehm. et Lindenb. in Lehmann 1832: 53) Stephani (1888c: 287), Cololejeunea (Spruce 1884: 251) Stephani (1888c: 289), C. marginata (Lehm. et Lindenb. in Lehmann 1833: 11) Stephani (1888c: 287), C. sicifolia Stephani (1888c: 290), C. sintenisii Stephani (1888c: 291), C. stylosa Stephani (1888c: 289), Diplasiolejeunea (Spruce 1884: 301) Stephani (1888c: 288), D. pellucida (Sprengel 1827: 325) Stephani (1888c: 288), Drepanolejeunea (Spruce 1884: 186) Stephani (1888c: 286), D. hamatifolia (Hooker 1813: tab. 51) Stephani (1888c: 286), D. inchoata (Lehm. et Lindenb. in Lehmann 1833: 19) Stephani (1888c: 287), D. tenuis (Nees 1830: 57) Stephani (1888c: 292), Euosmolejeunea (Spruce 1884: 241) Stephani (1888c: 292), E. trifaria (Reinwardt et al. 1824: 226) Stephani (1888c: 292), Harpalejeunea (Spruce 1884: 164) Stephani (1888c: 288), H. patentissima (Hampe & Gottsche 1853: 355) Stephani (1888c: 288), H. stricta (Lindenb. et Gottsche in Gottsche et al. 1847: 756) Stephani (1888c: 291), Hygrolejeunea (Spruce 1884: 230) Stephani (1888c: 284), H. cerina (Lehm. et Lindenb. in Lehmann 1833: 16) Stephani (1888c: 284), Leptolejeunea (Spruce 1884: 193) Stephani (1888c: 285), L. elliptica (Lehm. et Lindenb. in Lehmann 1833: 13) Stephani (1888c: 285), Macrolejeunea (Spruce 1884: 224) Stephani (1888c: 292), M. subsimplex (Mont. et Nees in Montagne 1843: 264) Stephani (1888c: 292, 301), Neurolejeunea (Spruce 1884: 84) Stephani (1888c: 283), N. portoricensis (Hampe & Gottsche 1853: 352) Stephani (1888c: 288), Odontolejeunea (Spruce 1884: 142) Stephani (1888c: 281, 284), O. accedens (Gottsche in Gottsche et al. 1845b: 339) Stephani (1888c: 281), O. berteroana Stephani (1888c: 282), O. convexistipa (Lehm. et Lindenb. in Lehmann 1834: 43) Stephani (1888c: 284), O. lunulata (Weber 1815: 33) Stephani (1888c: 287), Omphalolejeunea (Spruce 1884: 86) Stephani (1888c: 282), O. filiformis (Swartz 1788: 144) Stephani (1888c: 285), Platylejeunea (Spruce 1884: 124) Stephani (1888c: 282), P. barbiflora (Lindenberg & Gottsche 1851: 630) Stephani (1888c: 282), P. conferta (C.F.W.Meissn. ex Sprengel 1827: 325) Stephani (1888c: 284), P. granulata (Nees 1833: 352) Stephani (1888c: 285), P. transversalis (Swartz 1788: 144) Stephani (1888c: 292, 301), P. vincentina (Gottsche in Gottsche et al. 1845b: 313) Stephani (1888c: 293), Prionolejeunea (Spruce 1884: 152) Stephani (1888c: 285), P. denticulata (Weber 1815: 30) Stephani (1888c: 285), Pycnolejeunea (Spruce 1884: 246) Stephani (1888c: 289), P. macroloba (Nees et Mont. in Montagne 1843: 260) Stephani (1888c: 290), P. schwaneckei Stephani (1888c: 289), Stictolejeunea (Spruce 1884: 81) Stephani (1888c: 287), S. squamata (Willd. in Weber 1815: 33) Stephani (1888c: 291), Strepsilejeunea (Spruce 1884: 168) Stephani (1888c: 287), S. involuta (Gottsche in Gottsche et al. 1845b: 350) Stephani (1888c: 287), Taxilejeunea (Spruce 1884: 212) Stephani (1888c: 281), T. antillana Stephani (1888c: 281), T. eggersiana Stephani (1888c: 285). Stephani (1889a; Mar) is the second in a series of three papers about Australian hepatics (the first part does not include any Lejeuneaceae). Brachiolejeunea has been considered established here by a combined generic-specific description following ICN Art. 41 Ex. 9, but since more than one Brachiolejeunea species is accepted, the name Brachiolejeunea is not validly published here (see above). Lopholejeunea seems to be validly published here through the specific description of Lopholejeunea norfolkensis treated as a combined generic-specific one, but as Stephani writes “dieses grossen Subgenus” it is apparent that he treated it at subgeneric rank. The index has new taxa under “Xxxlejeunea” and sometimes also under Lejeunea, but identifies taxa as new only under “Xxxlejeunea”. Bonner et al. (1961) also regarded Drepanolejeunea as validly published here, but since more than one species are described the conditions for a combined generic-specific description are not fulfilled. One genus, Euosmolejeunea, is validly published by a combined generic-specific description and three additional binomials under earlier validly published generic names are validly named or combined here. Twenty-six binomials published under six generic names are not validly named or combined since their genera had not been validly published. Names validly published:—Eulejeunea armitii Stephani (1889a: 169), E. denticalyx Stephani (1889a: 169), E. drummondii (Taylor 1846: 400) Stephani (1889a: 170), E. subelobata (Carrington & Pearson 1888: 1039) Stephani (1889a: 170), Euosmolejeunea (Spruce 1884: 241) Stephani (1889a: 170), E. sayeri Stephani (1889a: 170). Names not validly published:—Acrolejeunea (Spruce 1884: 115) Stephani (1889a: 164, 165), A. hartmannii Stephani (1889a: 164), A. novae-guineae Stephani (1889a: 165), A. occulta Stephani (1889a: 166), A. pappeana (Nees in Gottsche et al. 1845a: 296) Stephani (1889a: 166), A. peradeniensis (Mitten 1860b: 111) Stephani (1889a: 166), A. polycarpa (Nees 1833: 350) Stephani (1889a: 166), A. pulopenangensis (Gottsche in Gottsche et al. 1845a: 299) Stephani (1889a: 166), A. terminalis (Spruce 1884: 116) Stephani (1889a: 166), A. wildii Stephani (1889a: 165), Brachiolejeunea (Spruce 1884: 129) Stephani (1889a: 167), B. birmensis Stephani (1889a: 167), B. caledonica Stephani (1889a: 167), B. papilionacea Stephani (1889a: 167), B. plagiochiloides Steph. et Spruce in Stephani (1889a: 167), B. securifolia Stephani (1889a: 168), B. succisa Stephani (1889a: 168), Cololejeunea (Spruce 1884: 251) Stephani (1889a: 168), C. bistyla Stephani (1889a: 168), C. trichomanis (Gottsche 1882: 362) Stephani (1889a: 168), Drepanolejeunea (Spruce 1884: 186) Stephani (1889a: 168), D. grossidens Stephani (1889a: 168), D. ternatensis (Gottsche in Gottsche et al. 1845b: 346) Stephani (1889a: 169), Hygrolejeunea (Spruce 1884: 230) Stephani (1889a: 171), H. chalmersii Stephani (1889a: 171), H. norfolkensis Stephani (1889a: 171), Hygrolejeunea rostrata Stephani (1889a: 172), H. sacculifera Stephani (1889a: 172), H. sayeri Stephani (1889a: 173), Leptolejeunea (Spruce 1884: 193) Stephani (1889a: 173), L. australis Stephani (1889a: 173), L. denticulata GENERA OF LEJEUNEACEAE ESTABLISHED IN 1884–1893 Phytotaxa 220 (2) © 2015 Magnolia Press • 147 Stephani (1889a: 174), L. rosulans Stephani (1889a: 174), Lopholejeunea (Spruce 1884: 119) Stephani (1889a: 175), L. norfolkensis Stephani (1889a: 175). Stephani (1889b; Apr) is the third and last paper in the series on Australian hepatics. It follows the same style as the previous paper. There is an indirect reference to Spruce’s publication (p. 259), but the connection to any generic name is weak and at best ambiguous, which Grolle (1983) also points out. Bonner et al. (1961) recognized more genera as valid (for the same reason as the Brachiolejeunea example above), but Zijlstra (1982) agreed with our interpretation. New species are only treated under “Xxxlejeunea” in the index. Two generic names, Taxilejeunea and Trachylejeunea, are established by a combined generic-specific descriptions in addition to one binomial at specific rank under an earlier validly published generic name. In addition one name in Lejeunea is validly published here. Twenty-six binomials in specific rank under five generic names are not validly published. Names validly published:—Lejeunea plagiochiloides Stephani (1889b: 257), Microlejeunea erectifolia (Spruce 1887: 36) Stephani (1889b: 259), Taxilejeunea (Spruce 1884: 212) Stephani (1889b: 262), T. convexa Stephani (1889b: 262), Trachylejeunea (Spruce 1884: 180) Stephani (1889b: 262), T. elegantissima Stephani (1889b: 262). Names not validly published:—Mastigolejeunea (Spruce 1884: 100) Stephani (1889b: 257), M. anguiformis Stephani (1889b: 257), M. buettneri Stephani (1889b: 257), M. crispula Stephani (1889b: 257), M. innovans (Spruce 1884: 103) Stephani (1889b: 258), M. longidens Stephani (1889b: 258), M. nigra Stephani (1889b: 258), M. phaea Stephani (1889b: 257), M. plicatiflora (Spruce 1884: 104) Stephani (1889b: 258), Ptycholejeunea (Spruce 1884: 97) Stephani (1889b: 258), P. birmensis Stephani (1889b: 258), P. perrottetii Stephani (1889b: 259), P. recondita Stephani (1889b: 259), P. stephensoniana (Mitten 1852: 155) Stephani (1889b: 258), P. theobromae (Spruce 1884: 99) Stephani (1889b: 259), Pycnolejeunea (Spruce 1884: 246) Stephani (1889b: 259), P. bidentula Stephani (1889b: 259), P. ceylanica (Gottsche in Gottsche et al. 1845a: 259) Stephani (1889b: 259), P. curvatiloba Stephani (1889b: 260), P. longidens Stephani (1889b: 260), Strepsilejeunea (Spruce 1884: 168) Stephani (1889b: 260), S. austrina Stephani (1889b: 260), S. luchmannii Stephani (1889b: 261), Thysanolejeunea (Spruce 1884: 105) Stephani (1889b: 262), T. abietina Stephani (1889b: 263), T. amazonica (Spruce 1884: 106) Stephani (1889b: 263), T. cucullata Stephani (1889b: 263), T. dissoptera (Spruce 1884: 108) Stephani (1889b: 263), T. pterobryoides (Spruce 1884: 109) Stephani (1889b: 263), T. reversa Stephani (1889b: 263), T. vittata (Mitten 1873: 411) Stephani (1889b: 263). Stephani (1890a, b, c) is a series of papers about the genus Lejeunea in Lindenberg’s herbarium. These are very messy papers, even by the Stephani standard. In the introduction Stephani only talks about Spruce’s subgenera. In these papers he uses the terms “Genus”, “Subgenus”, “Gattung” and “Abtheilung” seemingly interchangeably. Bonner et al. (1961) considered 22 subgeneric names to be validly combined to form new generic names, but Zijlstra (1982) and Grolle (1983) concluded that no generic names were validly published here. In this series of papers Stephani also uses the same specific epithet for taxa in several (sub)genera, suggesting that he interpreted them as something in between genera and subgenera as we think about them today. As his intention is so unclear, and no consensus has been reached in spite of more than 50 years of debate, this series of publications will be proposed for suppression at the rank of genus (Appendix VI of the ICN). New names or combinations under Lejeunea are validly published here, but only when definitely associated with the generic name (cf. ICN Art. 35.2). Binomials at specific rank under “Xxxlejeunea” names validly published earlier can be regarded as validly published in this series of publications if they are definitely associated with the genus. Stephani (1890a; Jan fide TL-2) treated all species under their pre-Sprucean names, including Lejeunea among others, but said that they belong to various “Xxxlejeunea” placing the hyphenated and abbreviated Sprucean names after them; e.g., “Lejeunea squamata (Willd.) Nees zu Sticto-Lej. Spr.” (p. 17). There is considerable disagreement as to whether the Sprucean names are being used at generic or subgeneric rank, even among the authors of this paper. As discussed above, it is because of this uncertainty, we are proposing that this publication be suppressed at the rank of genus. A new subgenus, Lejeunea subg. Caudalejeunea Stephani (1890a: 18) is generally accepted as validly published here, but there is no definite connection to the genus Lejeunea, which renders it not validly published (cf. ICN Art. 35.2). Three binomials are validly published here since the generic name was validly published earlier. Another 13 binomials are used in the text but not validly published since the generic name had not been validly published (some of these are also not validly published for additional reasons). Names validly published:—Archilejeunea bongardii Stephani (1890a: 20), A. clypeata (Schweinitz 1821: 12) Stephani (1890a: 21), A. mariana (Gottsche in Gottsche et al. 1845b: 337) Stephani (1890a: 17). Names not validly published:—Acrolejeunea (Spruce 1884: 115) Stephani (1890a: 7), A. borgenii Stephani (1890a: 10), A. cumingiana (Montagne 1845a: 7) Stephani (1890a: 20), A. fertilis (Reinwardt et al. 1824: 211) Stephani (1890a: 10), A. polycarpa (Nees 1833: 350) Stephani (1890a: 22), Brachiolejeunea (Spruce 1884: 129) Stephani (1890a: 7), B. corticalis (Lehm. et Lindenb. in Lehmann 1832: 50) Stephani (1890a: 7), B. laxifolia (Taylor 1847: 341) Stephani (1890a: 7), B. tristis Stephani (1890a: 8), Bryolejeunea (Spruce 1884: 111) Stephani (1890a: 2), Caudalejeunea Stephani (1890a: 18), C. leiboldii Stephani (1890a: 19), Ceratolejeunea (Spruce 1884: 198) Stephani (1890a: 16, 18), Cololejeunea (Spruce 1884: 251) Stephani (1890a: 18), Dicranolejeunea (Spruce 1884: 138) Stephani (1890a: 16), D. mexicana (Lindenb. in Gottsche et al. 1845a: 325) Stephani (1890a: 13), D. phyllorhiza (Nees 1833: 348) Stephani (1890a: 7), Homalolejeunea brachiata (Swartz 1788: 144) Stephani (1890a: 14) ICN Art. 41.1; basionym not valid, H. guilleminiana (Montagne 1841: 128) Stephani (1890a: 14) ICN 41.1 (basionym not valid), Hygrolejeunea (Spruce 148 • Phytotaxa 220 (2) © 2015 Magnolia Press SÖDERSTRÖM ET AL. 1884: 230) Stephani (1890a: 23), Lopholejeunea (Spruce 1884: 119) Stephani (1890a: 16), L. nigricans (Lindenb. in Gottsche et al. 1845a: 316) Stephani (1890a: 16), L. zollingeri Stephani (1890a: 14), Mastigolejeunea (Spruce 1884: 100) Stephani (1890a: 8), M. ligulata (Lehm. et Lindenb. in Lehmann 1834: 39) Stephani (1890a: 8), M. versicolor (Lehm. et Lindenb. in Gottsche et al. 1845a: 297) Stephani (1890a: 8), Odontolejeunea (Spruce 1884: 142) Stephani (1890a: 19), O. convexistipa (Lehm. et Lindenb. in Lehmann 1834: 43) Stephani (1890a: 20), O. peruviana (Lehm. et Lindenb. in Lehmann 1833: 18) Stephani (1890a: 19), O. sagittistipula Stephani (1890a: 19), Peltolejeunea (Spruce 1884: 85) Stephani (1890a: 23), Platylejeunea (Spruce 1884: 124) Stephani (1890a: 13), P. setosa Stephani (1890a: 14), Ptycholejeunea (Spruce 1884: 97) Stephani (1890a: 5), P. perrottetii Stephani (1890a: 5), Pycnolejeunea (Spruce 1884: 246) Stephani (1890a: 10, 11), P. macroloba (Nees et Mont. in Montagne 1843: 260) Stephani (1890a: 10), P. meyeniana (Gottsche et al. 1843: 472) Stephani (1890a: 11), Stictolejeunea (Spruce 1884: 81) Stephani (1890a: 17), Thysanolejeunea (Spruce 1884: 105) Stephani (1890a: 4). Stephani (1890b; Feb fide TL-2) is the continuation of his review of Lindenberg’s herbarium. Here he changed the style of writing the names and only used “Xxxlejeunea” as binomials at specific rank, except in discussions of a few taxa, and he seems to treat them differently than in the first part (as if he changed his mind between publication of the two parts). He did not refer to any generic name without binomials and thus, no generic name can be considered validly published here. However, binomials at specific rank are validly published if the generic name was published validly earlier. Fifty-five binomials under five generic names are validly published here as the generic names were validly published earlier. As many as 139 binomials at specific rank under 18 generic names (including three names in Lejeunea) are effectively but not validly published here. Names validly published:—Eulejeunea angusta (Lehm. et Lindenb. in Lehmann 1832: 52) Stephani (1890b: 86), E. caespitosa (Lindenb. in Gottsche et al. 1845b: 384) Stephani (1890b: 86), E. capensis (Gottsche in Gottsche et al. 1845b: 374) Stephani (1890b: 83), E. clavata (Lindenb. in Gottsche et al. 1845b: 379) Stephani (1890b: 85), E. concava (Lindenb. et Gottsche in Gottsche et al. 1847: 759) Stephani (1890b: 78), E. corallina Stephani (1890b: 83), E. diversistipa (Lindenb. in Gottsche et al. 1847: 765) Stephani (1890b: 85), E. eckloniana (Lindenb. in Gottsche et al. 1845b: 381) Stephani (1890b: 86), E. glaucophylla (Gottsche 1857: 345) Stephani (1890b: 87), E. gottscheana (Lindenb. in Gottsche et al. 1845b: 382) Stephani (1890b: 86), E. haitica (Nees et Mont. in Montagne 1843: 263) Stephani (1890b: 89), E. huctumalcensis (Lindenb. et Gottsche in Gottsche et al. 1847: 762) Stephani (1890b: 81), E. laeta (Lehm. et Lindenb. in Lehmann 1834: 45) Stephani (1890b: 85), E. laetevirens (Nees et Mont. in Montagne 1842: 469) Stephani (1890b: 87), E. laxa (Nees 1830: 43) Stephani (1890b: 85), E. lepida (Lindenb. et Gottsche in Gottsche et al. 1847: 766) Stephani (1890b: 87), E. lucens (Taylor 1846: 399) Stephani (1890b: 84), E. magnoliae (Lindenb. et Gottsche in Gottsche et al. 1847: 763) Stephani (1890b: 81), E. nudipes (Hooker & Taylor 1844c: 568) Stephani (1890b: 82), E. olivacea Stephani (1890b: 85), E. pulvinata (Nees & Montagne 1836: 61) Stephani (1890b: 86), E. radicans (Lindenb. et Gottsche in Gottsche et al. 1847: 766) Stephani (1890b: 87), E. seriata (Lindenb. et Gottsche in Gottsche et al. 1847: 762) Stephani (1890b: 81), E. serpyllifolia (Libert 1820: 374) Stephani (1890b: 83), E. spiniloba (Lindenb. et Gottsche in Gottsche et al. 1847: 770) Stephani (1890b: 98), E. tabularis (Sprengel 1827: 325) Stephani (1890b: 83), E. vogelii (Taylor 1846: 397) Stephani (1890b: 98), E. wallichiana (Lehmann 1831: 5) Stephani (1890b: 78), E. wightii (Lindenb. in Gottsche et al. 1845b: 379) Stephani (1890b: 85), E. zacuapana Stephani (1890b: 87), Euosmolejeunea beyrichii (Lindenb. in Gottsche et al. 1845b: 371) Stephani (1890b: 82), E. contigua (Nees 1833: 360) Stephani (1890b: 78), E. duriuscula (Nees in Gottsche et al. 1845b: 364) Stephani (1890b: 80), E. implexicaulis (Hooker & Taylor 1844a: 397) Stephani (1890b: 84), E. lutea (Mont. ex Gottsche et al. 1845b: 383) Stephani (1890b: 86), E. opaca (Gottsche in Gottsche et al. 1845b: 352) Stephani (1890b: 79), E. orientalis (Gottsche in Gottsche et al. 1845b: 371) Stephani (1890b: 82), E. parvistipula (Lindenb. et Gottsche in Gottsche et al. 1847: 761) Stephani (1890b: 80), E. tenella (Taylor 1846: 398) Stephani (1890b: 87), Lejeunea papilliloba Stephani (1890b: 73), L. poeppigiana (Nees in Gottsche et al. 1845b: 396) Stephani (1890b: 92), L. pyriformis Stephani (1890b: 93), nom. illeg. (ICN Art. 53.1; non Lindenb. et Gottsche in Gottsche et al. 1847: 767), Microlejeunea albicans (Nees in Gottsche et al. 1845b: 386) Stephani (1890b: 88), M. albovirens (Hooker & Taylor 1844a: 397) Stephani (1890b: 88), M. bullata (Taylor 1846: 398) Stephani (1890b: 90), M. microstipula Stephani (1890b: 89), M. primordialis (Hooker & Taylor 1845: 92) Stephani (1890b: 84), M. punctiformis (Taylor 1846: 398) Stephani (1890b: 90), M. stricta Stephani (1890b: 90), M. ulicina (Taylor 1844: 115) Stephani (1890b: 88), Taxilejeunea caripensis (Lindenb. et Gottsche in Gottsche et al. 1847: 758) Stephani (1890b: 75), T. chamissonis (Lindenb. in Gottsche et al. 1845b: 378) Stephani (1890b: 85), T. conformis (Nees et Mont. in Montagne 1843: 260) Stephani (1890b: 75), T. cordistipula (Lindenb. et Gottsche in Gottsche et al. 1847: 758) Stephani (1890b: 75), T. flaccida (Lindenb. et Gottsche in Gottsche et al. 1847: 758) Stephani (1890b: 75), T. neumanniana (Nees in von Flotow et al. 1842: 44) Stephani (1890b: 75), T. tenera (Swartz 1788: 143) Stephani (1890b: 98), Trachylejeunea ambigua (Lindenb. et Gottsche in Gottsche et al. 1847: 764) Stephani (1890b: 84), T. raddiana (Lindenb. in Gottsche et al. 1845b: 342) Stephani (1890b: 70). Names not validly published:—Archilejeunea conferta (C.F.W.Meissn. ex Sprengel 1827: 325) Stephani (1890b: 73) ICN Art. 41.1; basionym not valid, Ceratolejeunea (Spruce 1884: 198) Stephani (1890b: 91–95), C. amazonica Stephani (1890b: 91), C. belangeriana (Gottsche in Gottsche et al. 1845b: 398) Stephani (1890b: 93), C. brasiliensis (Gottsche in Gottsche et al. 1845b: 398) Stephani (1890b: 93), C. ceratantha (Nees et Mont. in Montagne 1840: 335) Stephani (1890b: 92), C. coarina (Gottsche in Gottsche et al. 1845b: 395) Stephani (1890b: 91), C. cornuta (Lindenberg 1829: 23) Stephani (1890b: 94), C. filaria (Lehmann 1844: 28) GENERA OF LEJEUNEACEAE ESTABLISHED IN 1884–1893 Phytotaxa 220 (2) © 2015 Magnolia Press • 149 Stephani (1890b: 96), C. guianensis (Nees et Mont. in Montagne 1840: 335) Stephani (1890b: 92), C. haeckeriana (Gottsche et Lindenb. in Gottsche et al. 1845b: 401) Stephani (1890b: 95), C. involvens (Nees et Mont. in Montagne 1840: 336) Stephani (1890b: 95), C. longicornis (Gottsche in Gottsche et al. 1845b: 401) Stephani (1890b: 95), C. martiana (Gottsche in Gottsche et al. 1845b: 398) Stephani (1890b: 93), C. miradorensis Stephani (1890b: 93), C. pyriformis (Stephani (1890b: 93), C. radicosa (Nees ex Montagne 1842: 475) Stephani (1890b: 95), C. rigidula (Nees ex Montagne 1840: 336) Stephani (1890b: 82), C. singapurensis (Lindenb. in Gottsche et al. 1845b: 397) Stephani (1890b: 93), C. splitgerberiana (Montagne 1841: 112) Stephani (1890b: 95), C. variabilis (Lindenb. in Gottsche et al. 1845b: 399) Stephani (1888c: 293), Cheilolejeunea (Spruce 1884: 251) Stephani (1890b: 80–86), C. bethanica (Gottsche in Gottsche et al. 1845b: 381) Stephani (1890b: 86), C. clausa (Nees et Mont. in Montagne 1840: 337) Stephani (1890b: 80), C. duriuscula (Nees in Gottsche et al. 1845b: 364) Stephani (1890b: 81), C. gaudichaudii (Lindenb. in Gottsche et al. 1845b: 372) Stephani (1890b: 82), C. intertexta (Lindenb. in Gottsche et al. 1845b: 379) Stephani (1890b: 85), C. lineata (Lehm. et Lindenb. in Lehmann 1832: 53) Stephani (1890b: 82), C. lurida (Lindenb. in Gottsche et al. 1845b: 379) Stephani (1890b: 85), C. melastomatis (Lindenb. et Gottsche in Gottsche et al. 1847: 763) Stephani (1890b: 82), C. myriantha (Nees et Mont. in Gottsche et al. 1845b: 398) Stephani (1890b: 80), C. oxyloba (Lindenb. et Gottsche in Gottsche et al. 1847: 770) Stephani (1890b: 81, 98), C. sandvicensis Stephani (1890b: 88), C. trinitensis (Lindenb. in Gottsche et al. 1845b: 381) Stephani (1890b: 86), C. virescens (Lehm. et Lindenb. in Lehmann 1838: 21) Stephani (1890b: 82), C. xanthophylla (Lindenb. in Gottsche et al. 1845b: 370) Stephani (1890b: 81), Cololejeunea (Spruce 1884: 251) Stephani (1890b: 71–96), C. calcarea (Libert 1820: 373) Stephani (1890b: 71), C. cardiocarpa (Montagne 1842: 476) Stephani (1890b: 91), C. cuneata (Lehm. et Lindenb. in Lehmann 1832: 56) Stephani (1890b: 91), C. epiphyta (Gottsche in Gottsche et al. 1845b: 391) Stephani (1890b: 90), C. floccosa (Lehm. et Lindenb. in Lehmann 1833: 26) Stephani (1890c: 135), C. haskarliana (Lehmann 1844: 26) Stephani (1890b: 72), C. kegelii Stephani (1890b: 91), C. lhotzskiana (Hampe in Gottsche et al. 1845b: 389) Stephani (1890b: 89), C. marginata (Lehm. et Lindenb. in Lehmann 1833: 11) Stephani (1890b: 91), C. microscopica (Taylor 1836: 59) Stephani (1890b: 72), C. minutissima (Sm. in Smith & Sowerby 1806: tab. 1633) Stephani (1890b: 88), C. montagnei Stephani (1890b: 96), C. myriocarpa (Nees et Mont. in Montagne 1842: 473) Stephani (1890b: 89), C. papilliloba (Stephani 1890b: 73), C. pyriformis (Lindenb. et Gottsche in Gottsche et al. 1847: 767) Stephani (1890b: 89), Colurolejeunea (Spruce 1884: 303) Stephani (1890b: 97–98), C. acroloba Stephani (1890b: 97), C. calyptrifolia (Hooker 1813: tab. 43) Stephani (1890b: 97), C. ceratophora (Nees in Gottsche et al. 1845b: 405) Stephani (1890b: 97), C. junghuhniana Stephani (1890b: 97), C. tortifolia (Nees et Mont. in Montagne 1843: 265) Stephani (1890b: 98), Crossotolejeunea (Spruce 1884: 161) Stephani (1890b: 70), C. boryana (Montagne 1838: 47) Stephani (1890b: 70), C. crenata (Nees et Mont. in Montagne 1838: 48) Stephani (1890b: 70), Diplasiolejeunea (Spruce 1884: 301) Stephani (1890b: 90–91), D. kraussiana (Lindenb. in Gottsche et al. 1845b: 393) Stephani (1890b: 91), D. pellucida (C.F.W.Meissn. ex Sprengel 1827: 325) Stephani (1890b: 91), D. unidentata (Lehm. et Lindenb. in Lehmann 1834: 48) Stephani (1890b: 90), Drepanolejeunea (Spruce 1884: 186) Stephani (1890b: 70–90), D. bidens Stephani (1890b: 71), D. capulata (Taylor 1846: 394) Stephani (1890b: 71), D. hamatifolia (Hooker 1813: tab. 51) Stephani (1890b: 71), D. hampeana Stephani (1890b: 70), D. inchoata (C.F.W.Meissn. in Lehmann 1833: 19) Stephani (1890b: 70), D. latitans (Hooker & Taylor 1844a: 399) Stephani (1890b: 72), D. lyratifolia (Hook.f. et Taylor in Taylor 1846: 393) Stephani (1890b: 71), D. muricata (Gottsche in Gottsche et al. 1845b: 348) Stephani (1890b: 74), D. palmifolia (Nees 1833: 366) Stephani (1890b: 74), D. plicatiloba (Hooker & Taylor 1845: 92) Stephani (1890b: 81), D. tenuis (Nees 1830: 57) Stephani (1890b: 90), D. ternatensis (Gottsche in Gottsche et al. 1845b: 346) Stephani (1890b: 73), D. teysmannii (Gottsche 1882: 360) Stephani (1890b: 72), D. tridactyla (Gottsche in Gottsche et al. 1845b: 347) Stephani (1890b: 73), Eulejeunea remotifolia Stephani (1890b: 86) ICN Art. 38.1(a); no description, Harpalejeunea (Spruce 1884: 164) Stephani (1890b: 70–85), H. cinchonae (Nees in Gottsche et al. 1845b: 342) Stephani (1890b: 70), H. ovata (Hooker 1813: tab. 42) Stephani (1890b: 84), H. oxyphylla (Nees et Mont. in Montagne 1843: 264) Stephani (1890b: 76), H. parasitica (Hooker & Taylor 1844b: 477) Stephani (1890b: 85), H. stricta (Lindenb. et Gottsche in Gottsche et al. 1847: 756) Stephani (1890b: 70), Hygrolejeunea (Spruce 1884: 230) Stephani (1890b: 74–98), H. alata (Gottsche in Gottsche et al. 1845b: 406) Stephani (1890b: 98), H. cerina (Lehm. et Lindenb. in Lehmann 1833: 16) Stephani (1890b: 90), H. cordifissa (Taylor 1846: 395) Stephani (1890b: 80), H. delessertii (Nees et Mont. in Montagne 1843: 260) Stephani (1890b: 74), H. densistipula (Lehm. et Lindenb. in Lehmann 1838: 20) Stephani (1890b: 81), H. deplanata (Nees in Gottsche et al. 1845b: 368) Stephani (1890b: 81), H. discreta (Lindenb. in Gottsche et al. 1845b: 361) Stephani (1890b: 78), H. eluta (Nees 1833: 362) Stephani (1890b: 78), H. heterocheila (Taylor 1846: 394) Stephani (1890b: 95), H. leucophylla (Lindenb. in Gottsche et al. 1845b: 369) Stephani (1890b: 81), H. microloba (Taylor 1846: 399) Stephani (1890b: 85), H. orba (Gottsche in Gottsche et al. 1845b: 352) Stephani (1890b: 95), H. pallida (Lindenb. et Gottsche in Gottsche et al. 1847: 762) Stephani (1890b: 81), H. phyllobola (Nees et Mont. in Montagne 1842: 471) Stephani (1890b: 81), H. pterota (Hooker & Taylor 1845: 91) Stephani (1890b: 81), H. sordida (Nees 1830: 41) Stephani (1890b: 80), Lejeunea fallax-vellongicornis Stephani (1890b: 96) ICN Art. 38.1(a); no description, Leptolejeunea (Spruce 1884: 193) Stephani (1890b: 270), L. corynophora (Gottsche et al. 1843: 474) Stephani (1890b: 97), L. denticulata Stephani (1890b: 70), L. elliptica (Lehm. et Lindenb. in Lehmann 1833: 13) Stephani (1890b: 96), L. neesii (Mont. in Nees & Montagne 1836: 62) Stephani (1890b: 74), L. polyrhiza (Nees in Gottsche et al. 1845b: 403) Stephani (1890b: 96), L. stenophylla (Lindenb. et Gottsche in Gottsche et al. 1847: 769) Stephani (1890b: 96), L. trematodes (Nees in Nees & Montagne 1836: 62) Stephani (1890b: 74), L. vitrea (Nees 150 • Phytotaxa 220 (2) © 2015 Magnolia Press SÖDERSTRÖM ET AL. 1830: 56) Stephani (1890b: 96), Macrolejeunea (Spruce 1884: 100) Stephani (1890b: 90), M. subsimplex (Mont. et Nees in Montagne 1843: 264), Odontolejeunea (Spruce 1884: 142) Stephani (1890b: 69), O. accedens (Gottsche in Gottsche et al. 1845b: 339) Stephani (1890b: 69), O. lunulata (Weber 1815: 33) Stephani (1890b: 98), O. peruviana (Lehm. et Lindenb. in Lehmann 1833: 18) Stephani (1890b: 69), Prionolejeunea (Spruce 1884: 152) Stephani (1890b: 69–70), P. aemula (Gottsche in Gottsche et al. 1845b: 338) Stephani (1890b: 69), P. angulistipa Stephani (1890b: 69), P. decora (Taylor 1846: 393) Stephani (1890b: 69), P. denticulata (Weber 1815: 30) Stephani (1890b: 68), P. guadalupensis (Lindenb. in Gottsche et al. 1845b: 340) Stephani (1890b: 70), P. meissneri (Gottsche in Gottsche et al. 1845b: 340) Stephani (1890b: 69), P. microdonta (Gottsche in Gottsche et al. 1845b: 337) Stephani (1890b: 69), P. serrulata (Montagne 1842: 479) Stephani (1890b: 69), Pycnolejeunea (Spruce 1884: 246) Stephani (1890b: 76–86), P. adnata (Kunze ex Lehmann 1834: 46) Stephani (1890b: 81), P. callosa (Gottsche in Gottsche et al. 1845b: 359) Stephani (1890b: 77), P. ceylanica (Gottsche in Gottsche et al. 1845a: 259) Stephani (1890b: 77), P. densiuscula Stephani (1890b: 86) ICN Art. 38.1(a); no description, P. discoidea (Lehm. et Lindenb. in Lehmann 1834: 47) Stephani (1890b: 86), P. hyalina Stephani (1890b: 77), P. imbricata (Nees 1830: 42) Stephani (1890b: 77), P. incisa (Gottsche in Gottsche et al. 1845b: 360) Stephani (1890b: 77), P. macroloba (Nees et Mont. in Montagne 1843: 260) Stephani (1890b: 77), P. meyeniana (Gottsche et al. 1843: 472) Stephani (1890b: 77), P. trapezia (Nees 1830: 41) Stephani (1890b: 76), Strepsilejeunea (Spruce 1884: 168) Stephani (1890b: 74–85), S. acuminata (Lehm. et Lindenb. in Lehmann 1834: 49) Stephani (1890b: 75), S. acutangula (Nees 1833: 357) Stephani (1890b: 74), S. comitans (Hook.f. et Taylor in Taylor 1846: 400) Stephani (1890b: 79), S. inflexa (Hampe in Lehmann 1838: 22) Stephani (1890b: 74), S. involuta (Gottsche in Gottsche et al. 1845b: 350) Stephani (1888c: 287), S. krakakammae (Lindenb. in Gottsche et al. 1845b: 353) Stephani (1890b: 74), S. kunthiana (Lindenb. in Gottsche et al. 1845b: 353) Stephani (1890b: 74), S. lanceolata (Gottsche in Gottsche et al. 1845b: 350) Stephani (1890b: 75), S. lobulata (Lindenb. in Gottsche et al. 1845b: 350) Stephani (1890b: 74), S. mimosa (Hooker & Taylor 1844a: 398) Stephani (1890b: 85), S. neelgherriana (Gottsche in Gottsche et al. 1845b: 354) Stephani (1890b: 75), S. obtruncata (Montagne 1845b: 354) Stephani (1890b: 84), S. orba (Gottsche in Gottsche et al. 1845b: 350) Stephani (1890b: 74), S. owaihiensis (Gottsche in Gottsche et al. 1845b: 351) Stephani (1890b: 74), S. tenuistipula Lindenb. et Gottsche in Gottsche et al. 1845b: 351) Stephani (1890b: 74). Stephani (1890c; Mar fide TL-2) is the last part in the series of papers of Lindenberg’s herbarium. It is a list of all species of the “Gattung Lejeunea” in the herbarium. All names have references to their original publication page. However, how the list should be regarded is controversial; some authors, e.g., Bonner et al. (1961), have concluded that this is a list of generic names, while others, e.g., Zijlstra (1982) and Grolle (1983), consider this a list of subgeneric names, based on Stephani’s statement found on p. 98 in the second paper of the series (Stephani 1890b). Because of the controversial interpretations of rank in this three-part series of publications, we will propose that these Stephani publications (1890a, b, c) be suppressed at the rank of genus (ICN Art. 34). If accepted, no generic names will be considered validly published in this series and only binomials at specific rank under generic names validly published prior to this series of publications can be considered validly published here. Thirty-five binomials at specific rank in five generic names are validly combined here as the generic names were validly published before and 234 binomials at specific rank in 30 generic names are effectively but not validly published here. Names validly published:—Archilejeunea auberiana (Montagne 1842: 483) Stephani (1890c: 134), A. chrysophylla (Lehmann 1835: 423) Stephani (1890c: 134), A. cognata (Nees 1833: 353) Stephani (1890c: 134), A. conferta (C.F.W.Meissn. ex Sprengel 1827: 325) Stephani (1890c: 134), A. cruegeri (Lindenb. in Gottsche et al. 1845b: 319) Stephani (1890c: 134), A. guahamensis (Lindenb. in Gottsche et al. 1845b: 346) Stephani (1890c: 134), A. huanucensis (Gottsche. in Gottsche et al. 1845b: 346) Stephani (1890c: 134), A. olivacea (Hooker & Taylor 1844c: 568) Stephani (1890c: 134), A. parviflora (Nees 1833: 353) Stephani (1890c: 134), A. rotalis (Hooker & Taylor 1845: 89) Stephani (1890c: 134), A. rotundistipula (Lindenb. ex Lehmann 1829: 360) Stephani (1890c: 134), A. scutellata (Taylor 1846: 383) Stephani (1890c: 134), A. unciloba (Lindenb. in Gottsche et al. 1845b: 331) Stephani (1890c: 134), A. viridissima (Lindenberg 1843: 320) Stephani (1890c: 134), A. xanthocarpa (Lehm. et Lindenb. in Lehmann 1833: 8) Stephani (1890c: 134), Euosmolejeunea lindenbergii (Gottsche in Gottsche et al. 1845b: 336) Stephani (1890c: 137), E. montagnei (Montagne 1843: 261) Stephani (1890c: 137), Homalolejeunea acutiloba (Hooker & Taylor 1845: 90) Stephani (1890c: 138), H. brachiata (Swartz 1788: 144) Stephani (1890c: 138), H. fuscescens (Hampe in Lehmann 1838: 16) Stephani (1890c: 138), H. languida (Nees & Montagne 1836: 59) Stephani (1890c: 138), H. mackaii (Hooker 1813: tab. 53) Stephani (1890c: 138), Microlejeunea gracillima (Mitten 1860b: 115) Stephani (1890c: 139), Taxilejeunea affinis (Lindenb. et Gottsche in Gottsche et al. 1847: 748) Stephani (1890c: 141), T. debilis (Lehm. et Lindenb. in Lehmann 1832: 51) Stephani (1890c: 141), T. gracilipes (Taylor 1846: 385) Stephani (1890c: 141), T. isocalycina (Nees 1833: 356) Stephani (1890c: 141), T. leioscyphus (Gottsche 1864: 148) Stephani (1890c: 141), T. lumbricoides (Nees 1830: 40) Stephani (1890c: 141), T. lusoria (Lindenb. et Gottsche in Gottsche et al. 1847: 747) Stephani (1890c: 141), T. martinicensis (Gottsche 1864: 149) Stephani (1890c: 141), T. neumanniana (Nees in von Flotow et al. 1842: 44) Stephani (1890b: 75), T. pterigonia (Lehm. et Lindenb. in Lehmann 1834: 44) Stephani (1890c: 142), T. renistipula (Lindenb. in Gottsche et al. 1845b: 308) Stephani (1890c: 142), T. sulphurea (Lehm. et Lindenb. in Lehmann 1833: 14) Stephani (1890c: 142), T. surinamensis (Lindenberg & Gottsche 1851: 628) Stephani (1890c: 142). Names not validly published:—Acrolejeunea (Spruce 1884: 115) Stephani (1890c: 133), A. amplectens (Stephani 1886: 86) Stephani GENERA OF LEJEUNEACEAE ESTABLISHED IN 1884–1893 Phytotaxa 220 (2) © 2015 Magnolia Press • 151 (1890c: 133), A. aulacophora (Montagne 1843: 259) Stephani (1890c: 133), A. borgenii Stephani (1890c: 133), A. cucullata Stephani (1890c: 133), A. cumingiana (Montagne 1845a: 7) Stephani (1890c: 133), A. fertilis (Reinwardt et al. 1824: 211) Stephani (1890c: 133), A. haskarliana (Gottsche in Gottsche et al. 1845a: 299) Stephani (1890c: 133), A. juliformis (Nees 1833: 351) Stephani (1890c: 133), A. molleri (Stephani 1886: 86) Stephani (1890c: 133), A. pappeana (Nees in Gottsche et al. 1845a: 296) Stephani (1890c: 133), A. polycarpa (Nees 1833: 350) Stephani (1890c: 133), A. pulopenangensis (Gottsche in Gottsche et al. 1845a: 299) Stephani (1890c: 133), A. securifolia (Nees in Endlicher 1833: 5) Stephani (1890c: 133), A. torulosa (Lehm. et Lindenb. in Lehmann 1834: 41) Stephani (1890c: 133), A. tumida (Nees 1838: 213) Stephani (1890c: 133), Brachiolejeunea (Spruce 1884: 129) Stephani (1890c: 134), B. bicolor (Nees 1833: 349) Stephani (1890c: 134), B. corticalis (Lehm. et Lindenb. in Lehmann 1832: 50) Stephani (1890c: 134), B. laxifolia (Taylor 1847: 341) Stephani (1890c: 134), B. leiboldiana (Gottsche et Lindenb. in Gottsche et al. 1845a: 296) Stephani (1890c: 134), B. liebmaniana (Lindenb. et Gottsche in Gottsche et al. 1847: 744) Stephani (1890c: 134), B. tristis Stephani (1890c: 134), B. ustulata (Taylor 1846: 388) Stephani (1890c: 134), Bryolejeunea (Spruce 1884: 111) Stephani (1890c: 134), B. diffusa (Swartz 1788: 144) Stephani (1890c: 134), B. filicina (Swartz 1788: 145) Stephani (1890c: 134), B. flaccida (Lindenb. et Hampe in Hampe 1851: 640) Stephani (1890c: 134), B. gaudichaudii (Gottsche 1857: 340) Stephani (1890c: 134), B. liebmaniana (Lindenb. et Gottsche in Gottsche et al. 1847: 738) Stephani (1890c: 134), B. tenuicaulis (Gottsche et al. 1845a: 285) Stephani (1890c: 134); Stephani (1892e: 179), B. trinitensis (Lehm. et Lindenb. in Lehmann 1833: 12) Stephani (1890c: 134), Caudalejeunea (Stephani 1890c: 134), C. crescentiae (Lindenb. et Gottsche in Gottsche et al. 1847: 752) Stephani (1890c: 134), C. lehmanniana (Gottsche in Gottsche et al. 1845b: 325) Stephani (1890c: 134), C. recurvistipula (Gottsche in Gottsche et al. 1845b: 326) Stephani (1890c: 134), Ceratolejeunea (Spruce 1884: 198) Stephani (1890c: 134), C. amazonica Stephani (1890c: 134), C. belangeriana (Gottsche in Gottsche et al. 1845b: 398) Stephani (1890c: 134), C. brasiliensis (Gottsche in Gottsche et al. 1845b: 398) Stephani (1890c: 134), C. breutelii (Gottsche in Gottsche et al. 1845b: 324) Stephani (1890c: 134), C. catenulata (Nees in Gottsche et al. 1845b: 323) Stephani (1890c: 134), C. ceratantha (Nees et Mont. in Montagne 1840: 335) Stephani (1890c: 134),C. coarina (Gottsche in Gottsche et al. 1845b: 395) Stephani (1890c: 134), C. cornuta (Lindenberg 1829: 23) Stephani (1890c: 134), C. cubensis (Montagne 1842: 481) Stephani (1890c: 134), C. devexa (Lindenb. et Gottsche in Gottsche et al. 1847: 750) Stephani (1890c: 134), C. filaria (Taylor in Lehmann 1844: 28) Stephani (1890c: 134), C. guianensis (Nees et Mont. in Montagne 1840: 335) Stephani (1890c: 134), C. haeckeriana (Gottsche et Lindenb. in Gottsche et al. 1845b: 401) Stephani (1890c: 135), C. involvens (Nees et Mont. in Montagne 1840: 336) Stephani (1890c: 135), C. longicornis (Gottsche in Gottsche et al. 1845b: 401) Stephani (1890c: 135), C. martiana (Gottsche in Gottsche et al. 1845b: 398) Stephani (1890c: 135), C. miradorensis Stephani (1890b: 93); Stephani (1890c: 135), C. oculata (Gottsche in Gottsche et al. 1845b: 357) Stephani (1890c: 135), C. poeppigiana (Nees in Gottsche et al. 1845b: 396) Stephani (1890c: 135), C. pyriformis (Stephani (1890b: 93) Stephani (1890c: 135), C. radicosa (Nees ex Montagne 1842: 475) Stephani 1890c: 135), C. rigidula (Nees ex Montagne 1840: 336) Stephani (1890c: 135), C. singapurensis (Lindenb. in Gottsche et al. 1845b: 397) Stephani (1890c: 135), C. spinosa (Gottsche in Gottsche et al. 1845b: 402) Stephani (1890c: 135), C. variabilis (Lindenb. in Gottsche et al. 1845b: 399) Stephani (1890c: 135), Cheilolejeunea (Spruce 1884: 251) Stephani (1890c: 135), C. bethanica (Gottsche in Gottsche et al. 1845b: 381) Stephani (1890c: 135), C. clausa (Nees et Mont. in Montagne 1840: 337) Stephani (1890c: 135), C. duriuscula (Nees in Gottsche et al. 1845b: 364) Stephani (1890c: 135), C. gaudichaudii (Lindenb. in Gottsche et al. 1845b: 372) Stephani (1890c: 135), C. intertexta (Lindenb. in Gottsche et al. 1845b: 379) Stephani (1890c: 135),C. lineata (Lehm. et Lindenb. in Lehmann 1832: 53) Stephani (1890c: 135), C. lurida (Lindenb. in Gottsche et al. 1845b: 379) Stephani (1890c: 135), C. melastomatis (Lindenb. et Gottsche in Gottsche et al. 1847: 763) Stephani (1890c: 135), C. myriantha (Nees et Mont. in Gottsche et al. 1845b: 398) Stephani (1890c: 135), C. oxyloba (Lindenb. et Gottsche in Gottsche et al. 1847: 770) Stephani (1890c: 135), C. sandvicensis Stephani (1890c: 135), C. virescens (Lehm. et Lindenb. in Lehmann 1838: 21) Stephani (1890c: 135), C. xanthophylla (Lindenb. in Gottsche et al. 1845b: 370) Stephani (1890c: 135), Cololejeunea (Spruce 1884: 251) Stephani (1890c: 135), C. calcarea (Libert 1820: 373) Stephani (1890c: 135), C. cardiocarpa (Montagne 1842: 476) Stephani (1890c: 135), C. cuneata (Lehm. et Lindenb. in Lehmann 1832: 56) Stephani (1890c: 135), C. epiphyta (Gottsche in Gottsche et al. 1845b: 391) Stephani (1890c: 135, C. floccosa (Lehm. et Lindenb. in Lehmann 1832: 26) Stephani (1890c: 135), C. haskarliana (Lehmann 1844: 26) Stephani (1890c: 135), C. kegelii Stephani (1890b: 91); Stephani (1890c: 135), C. lhotzskiana (Hampe in Gottsche et al. 1845b: 389) Stephani (1890c: 135), C. marginata (Lehm. et Lindenb. in Lehmann 1833: 11) Stephani (1890c: 135), C. microscopica (Taylor 1836: 59) Stephani (1890c: 135), C. minutissima (Sm. in Smith & Sowerby 1806: tab. 1633) Stephani (1890c: 135), C. montagnei Stephani (1890c: 135), C. myriocarpa (Nees et Mont. in Montagne 1842: 473) Stephani (1890c: 135), C. papilliloba (Stephani 1890b: 73) Stephani (1890c: 135), C. pyriformis (Lindenb. et Gottsche in Gottsche et al. 1847: 767) Stephani (1890c: 135), Colurolejeunea (Spruce 1884: 303) Stephani (1890c: 135), C. acroloba Stephani (1890c: 135), C. calyptrifolia (Hooker 1813: tab. 43) Stephani (1890c: 135), C. ceratophora (Nees in Gottsche et al. 1845b: 405) Stephani (1890c: 135), C. junghuhniana Stephani (1890c: 135), C. tortifolia (Nees et Mont. in Montagne 1843: 265) Stephani (1890c: 135), Crossotolejeunea (Spruce 1884: 161) Stephani (1890c: 136), C. boryana (Montagne 1838: 47) Stephani (1890c: 135), C. crenata (Nees et Mont. in Montagne 1838: 48) Stephani (1890c: 135), Dicranolejeunea (Spruce 1884: 138) Stephani (1890c: 136), D. aberrans (Lindenb. et Gottsche in Gottsche et al.1847: 751) Stephani (1890c: 136), D. axillaris (Nees & Montagne 1836: 59) Stephani (1890c: 136), D. dubiosa (Lindenb. et 152 • Phytotaxa 220 (2) © 2015 Magnolia Press SÖDERSTRÖM ET AL. Gottsche in Gottsche et al. 1847: 752) Stephani (1890c: 136), D. incongrua (Lindenb. et Gottsche in Gottsche et al. 1847: 750) Stephani (1890c: 136), D. mexicana (Lindenb. in Gottsche et al. 1845a: 325) Stephani (1890c: 136), D. phyllorhiza (Nees 1833: 348) Stephani (1890c: 136), D. trigona (Nees & Montagne 1836: 61) Stephani (1890c: 136), Diplasiolejeunea (Spruce 1884: 301) Stephani (1890c: 136), D. kraussiana (Lindenb. in Gottsche et al. 1845b: 393) Stephani (1890c: 136), D. pellucida (C.F.W.Meissn. ex Sprengel 1827: 325) Stephani (1890c: 136), D. unidentata (Lehm. et Lindenb. in Lehmann 1834: 48) Stephani (1890c: 136), Drepanolejeunea (Spruce 1884: 186) Stephani (1890c: 136), D. bidens Stephani (1890c: 136), D. capulata (Taylor 1846: 394) Stephani (1890c: 136), D. hamatifolia (Hooker 1813: tab. 51) Stephani (1890c: 136), D. hampeana Stephani (1890c: 136), D. inchoata (C.F.W.Meissn. in Lehmann 1833: 19) Stephani (1890c: 136), D. latitans (Hooker & Taylor 1844a: 399) Stephani (1890c: 136), D. lyratifolia (Hook.f. et Taylor in Taylor 1846: 393) Stephani (1890c: 136), D. muricata (Gottsche in Gottsche et al. 1845b: 348) Stephani (1890c: 136), D. palmifolia (Nees 1833: 366) Stephani (1890c: 136), D. plicatiloba (Hooker & Taylor 1845: 92) Stephani (1890c: 136), D. tenuis (Nees 1830: 57) Stephani (1890c: 136), D. ternatensis (Gottsche in Gottsche et al. 1845b: 346) Stephani (1890c: 136), D. teysmannii (Gottsche 1882: 360) Stephani (1890c: 136), D. tridactyla (Gottsche in Gottsche et al. 1845b: 347) Stephani (1890c: 136), Eulejeunea remotifolia Stephani (1890c: 137) ICN Art. 38.1(a); no description, Harpalejeunea (Spruce 1884: 164) Stephani (1890c: 138), H. cinchonae (Nees in Gottsche et al. 1845b: 342) Stephani (1890c: 138), H. ovata (Hooker 1813: tab. 42) Stephani (1890c: 138), H. oxyphylla (Nees et Mont. in Montagne 1843: 264) Stephani (1890c: 138), H. parasitica (Hooker & Taylor 1844b: 477) Stephani (1890c: 138), H. stricta (Lindenb. et Gottsche in Gottsche et al. 1847: 756) Stephani (1890c: 138), Hygrolejeunea (Spruce 1884: 230) Stephani (1890c: 138), H. alata (Gottsche in Gottsche et al. 1845b: 406) Stephani (1890c: 138), H. cerina (Lehm. et Lindenb. in Lehmann 1833: 16) Stephani (1890c: 138), H. cordifissa (Taylor 1846: 395) Stephani (1890c: 138), H. delessertii (Nees et Mont. in Montagne 1843: 260) Stephani (1890c: 138), H. densistipula (Lehm. et Lindenb. in Lehmann 1838: 20) Stephani (1890c: 138), H. deplanata (Nees in Gottsche et al. 1845b: 368) Stephani (1890c: 138), H. discreta (Lindenb. in Gottsche et al. 1845b: 361) Stephani (1890c: 138), H. eluta (Nees 1833: 362) Stephani (1890c: 138), H. heterocheila (Taylor 1846: 394) Stephani (1890c: 138), H. leucophylla (Lindenb. in Gottsche et al. 1845b: 369) Stephani (1890c: 138), H. microloba (Taylor 1846: 399) Stephani (1890c: 138), H. orba (Gottsche in Gottsche et al. 1845b: 352) Stephani (1890c: 138), H. pallida (Lindenb. et Gottsche in Gottsche et al. 1847: 762) Stephani (1890c: 138), H. phyllobola (Nees et Mont. in Montagne 1842: 471) Stephani (1890c: 138), H. pterota (Hooker & Taylor 1845: 91) Stephani (1890c: 138), H. reflexistipula (Lehm. et Lindenb. in Lehmann 1833: 10) Stephani (1890c: 138), H. sordida (Nees 1830: 41) Stephani (1890c: 138), H. umbilicata (Nees 1830: 42) Stephani (1890c: 138), Leptolejeunea (Spruce 1884: 193) Stephani (1890c: 138), L. corynophora (Gottsche et al. 1843: 474) Stephani (1890c: 138), L. denticulata Stephani (1890c: 138), L. elliptica (Lehm. et Lindenb. in Lehmann 1833: 13) Stephani (1890c: 138), L. neesii (Mont. in Nees & Montagne 1836: 62) Stephani (1890c: 138), L. polyrhiza (Nees in Gottsche et al. 1845b: 403) Stephani (1890c: 138), L. stenophylla (Lindenb. et Gottsche in Gottsche et al. 1847: 769) Stephani (1890c: 138), L. trematodes (Nees in Nees & Montagne 1836: 62) Stephani (1890c: 138), L. vitrea (Nees 1830: 56) Stephani (1890c: 138), Lopholejeunea (Spruce 1884: 193) Stephani (1890c: 138), L. applanata (Reinwardt et al. 1824: 210) Stephani (1890c: 139), L. eulopha (Taylor 1846: 391) Stephani (1890c: 139), L. javanica (Nees in Gottsche et al. 1845b: 320) Stephani (1890c: 139), L. muelleriana (Gottsche 1863: 184) Stephani (1890c: 139), L. nigricans (Lindenb. in Gottsche et al. 1845a: 316) Stephani (1890c: 139), L. plicatiscypha (Taylor 1846: 386) Stephani (1890c: 139), L. sagrana (Montagne 1842: 464) Stephani (1890c: 139), L. sphaerophora (Lehm. et Lindenb. in Lehmann 1833) Stephani (1890c: 139), L. zollingeri Stephani (1890a: 14); Stephani (1890c: 139), Macrolejeunea (Spruce 1884: 224) Stephani (1890c: 139), M. subsimplex (Mont. et Nees in Montagne 1843: 264) Stephani (1890c: 139), Mastigolejeunea (Spruce 1884: 100) Stephani (1890c: 139), M. arcuata (Nees 1830: 38) Stephani (1890c: 139), M. auriculata (Wilson 1841: no. 170) Stephani (1890c: 139), M. frauenfeldii (Reichardt 1866: 958) Stephani (1890c: 139), M. humilis (Gottsche in Gottsche et al. 1845a: 239) Stephani (1890c: 139), M. ligulata (Lehm. et Lindenb. in Lehmann 1834: 39) Stephani (1890a: 8); Stephani (1890c: 139), M. repleta (Taylor 1846: 392) Stephani (1890c: 139), M. taitica (Gottsche 1864: 147) Stephani (1890c: 139), Odontolejeunea (Spruce 1884: 142) Stephani (1890c: 139), O. accedens (Gottsche in Gottsche et al. 1845b: 339) Stephani (1890c: 139), O. convexistipa (Lehm. et Lindenb. in Lehmann 1834: 43) Stephani (1890c: 139), O. lunulata (Weber 1815: 33) Stephani (1890c: 139), O. peruviana (Lehm. et Lindenb. in Lehmann 1833: 18) Stephani (1890c: 139), O. sagittistipula Stephani (1890c: 140), O. sieberiana (Gottsche in Gottsche et al. 1845b: 328) Stephani (1890c: 140), Omphalolejeunea (Spruce 1884: 86) Stephani (1890c: 140), O. filiformis (Swartz 1788: 144) Stephani (1890c: 140), Peltolejeunea (Spruce 1884: 85) Stephani (1890c: 140), P. ovalis (Lindenb. et Gottsche in Gottsche et al. 1847: 754) Stephani (1890c: 140), Platylejeunea (Spruce 1884: 124) Stephani (1890c: 140), P. baccifera (Taylor 1846: 387) Stephani (1890c: 140), P. barbiflora (Lindenberg & Gottsche 1851: 630) Stephani (1890c: 140), P. granulata (Nees 1833: 352) Stephani (1890c: 140), P. hobsoniana (Lindenb. in Gottsche et al. 1845b: 313) Stephani (1890c: 140), P. setosa Stephani (1890c: 140), P. subrotunda (Hook. in Kunth 1822: 41) Stephani (1890c: 140), P. transversalis (Swartz 1788: 144) Stephani (1890c: 140), P. vincentina (Gottsche in Gottsche et al. 1845b: 313) Stephani (1890c: 140), Prionolejeunea (Spruce 1884: 152) Stephani (1890c: 140), P. aemula (Gottsche in Gottsche et al. 1845b: 338) Stephani (1890c: 140), P. angulistipa Stephani (1890c: 140), P. decora (Taylor 1846: 393) Stephani (1890c: 140), P. denticulata (Weber 1815: 30) Stephani (1890c: 140), P. guadalupensis (Lindenb. in Gottsche et al. 1845b: 340) Stephani (1890c: 140), P. meissneri (Gottsche in Gottsche et al. 1845b: 340) Stephani (1890c: 140), P. microdonta (Gottsche in GENERA OF LEJEUNEACEAE ESTABLISHED IN 1884–1893 Phytotaxa 220 (2) © 2015 Magnolia Press • 153 Gottsche et al. 1845b: 337) Stephani (1890c: 140), P. serrulata (Montagne 1842: 479) Stephani (1890c: 140), Ptycholejeunea (Spruce 1884: 97) Stephani (1890c: 140), P. javanica (Nees in Gottsche et al. 1845a: 291) Stephani (1890c: 140), P. perrottetii Stephani (1890c: 140), P. pycnoclada (Taylor 1846: 385) Stephani (1890c: 140), P. retusa (Reinwardt et al. 1824: 214) Stephani (1890c: 140), P. semirepanda (Nees 1830: 39) Stephani (1890c: 140), P. striata (Lehm. et Lindenb. in Lehmann 1832: 16) Stephani (1890c: 140), P. sulcata (Nees 1830: 36) Stephani (1890c: 140), P. wightii (Gottsche in Gottsche et al. 1845a: 291) Stephani (1890c: 140), Pycnolejeunea (Spruce 1884: 246) Stephani (1890c: 140), P. adnata (Kunze ex Lehmann 1834: 46) Stephani (1890c: 140), P. callosa (Gottsche in Gottsche et al. 1845b: 359) Stephani (1890c: 140), P. ceylanica (Gottsche in Gottsche et al. 1845a: 259) Stephani (1890c: 140), P. densiuscula Stephani (1890c: 141) ICN Art. 38.1(a); no description, P. discoidea (Lehm. et Lindenb. in Lehmann 1834: 47) Stephani (1890c: 141), P. hyalina Stephani (1890c: 141), P. imbricata (Nees 1830: 42) Stephani (1890c: 141), P. incisa (Gottsche in Gottsche et al. 1845b: 360) Stephani (1890c: 141), P. macroloba (Nees et Mont. in Montagne 1843: 260) Stephani (1890c: 141), P. meyeniana (Gottsche et al. 1843: 472) Stephani (1890c: 141), P. trapezia (Nees 1830: 41) Stephani (1890c: 141), Stictolejeunea (Spruce 1884: 81) Stephani (1890c: 141), S. kunzeana (Gottsche et al. 1845b: 322) Stephani (1890c: 141), S. squamata (Willd. in Weber 1815: 33) Stephani (1890c: 141), Strepsilejeunea (Spruce 1884: 168) Stephani (1890c: 141), S. acuminata (Lehm. et Lindenb. in Lehmann 1834: 49) Stephani (1890c: 141), S. acutangula (Nees 1833: 357) Stephani (1890c: 141), S. comitans (Hook.f. et Taylor in Taylor 1846: 400) Stephani (1890c: 141), S. inflexa (Hampe in Lehmann 1838: 22) Stephani (1890c: 141), S. involuta (Gottsche in Gottsche et al. 1845b: 350) Stephani (1890c: 141), S. krakakammae (Lindenb. in Gottsche et al. 1845b: 353) Stephani (1890c: 141), S. kunthiana (Lindenb. in Gottsche et al. 1845b: 353) Stephani (1890c: 141), S. lanceolata (Gottsche in Gottsche et al. 1845b: 350) Stephani (1890c: 141), S. lobulata (Lindenb. in Gottsche et al. 1845b: 350) Stephani (1890c: 141), S. mimosa (Hooker & Taylor 1844a: 398) Stephani (1890c: 141), S. neelgherriana (Gottsche in Gottsche et al. 1845b: 354) Stephani (1890c: 141), S. obtruncata (Montagne 1845b: 354) Stephani (1890c: 141), S. orba (Gottsche in Gottsche et al. 1845b: 350) Stephani (1890c: 141), S. owaihiensis (Gottsche in Gottsche et al. 1845b: 351) Stephani (1890c: 141), S. tenuistipula Lindenb. et Gottsche in Gottsche et al. 1845b: 351) Stephani (1890c: 141), Taxilejeunea guadalupensis Stephani (1890c: 141) ICN Art. 38.1(a); no description, Thysanolejeunea (Spruce 1884: 105) Stephani (1890c: 142), T. anguiformis (Hooker & Taylor 1844c: 567) Stephani (1890c: 142), T. comosa (Lindenb. in Lehmann 1844: 25) Stephani (1890c: 142), T. convoluta (Lindenb. in Gottsche et al. 1845a: 288) Stephani (1890c: 142), T. dissoptera (Spruce 1884: 108) Stephani (1890c: 142), T. fruticosa (Lindenb. et Gottsche in Gottsche et al. 1847: 737) Stephani (1890c: 142), T. lehmanniana (Nees in Gottsche et al. 1845a: 302) Stephani (1890c: 142), T. plana (Sande Lacoste 1854: 419) Stephani (1890c: 142), T. spathulistipa (Reinwardt et al. 1824: 212) Stephani (1890c: 142). Schiffner (1890; May) is at first glance treating the taxa as genera, but when he described Phragmo-Lejeunia he clearly stated that it is a new subgenus, but he did not say of what genus. Thus, the name is not validly published (ICN Art. 35.2) as it is not definitely associated with a generic name. As he used the same typography here as for the other Lejeuneaceae species it is clear that he has all taxa at subgeneric level. There is a reference to Spruce under Colurolejeunea which might be taken as a combination, but he stated that he considers it a subgenus (see p. 35 “Hujus subgeneris …”). Thus, no generic name is validly published here. It is worth noting that Schiffner used the same specific epithet (“amboinensis”) in two of his new names under different subgenera. In several cases, the names have been treated as validly published Lejeunea names (e.g. by Gradstein et al. 2002) and some of them even as forming the Lejeunea basionym to taxa still accepted today. Although Schiffner (1898) and others are pointing back to those names as Lejeunea names, no new name can be regarded established under Lejeunea since the names are not definitely associated with the generic name in this publication (ICN Art. 35.2). There is no index. Three binomials are validly published since the generic names were validly published earlier. Thirty-three binomials in 11 generic names are effectively but not validly published. Most of the new names that are not validly published here are validly published in Sydow (1894) by the direct reference to Schiffner’s publication as most of the generic names were validly published by then. Names validly published:—Eulejeunea crenulata Schiffner (1890: 33), Euosmolejeunea trifaria (Reinwardt et al. 1824: 226) Schiffner (1890: 31), Microlejeunea parallela Schiffner (1890: 33). Names not validly published:—Acrolejeunea (Spruce 1884: 88) Schiffner (1890: 26), A. densifolia Schiffner (1890: 26), A. rostrata Schiffner (1890: 26), Ceratolejeunea (Spruce 1884: 207) Schiffner (1890: 30), C. auriculata Schiffner (1890: 30), Cololejeunea (Spruce 1884: 251) Schiffner (1890: 33, 34), C. angustibracteata Schiffner (1890: 34), C. pseudostipulata Schiffner (1890: 33), Colurolejeunea (Spruce 1884: 303) Schiffner (1890: 34), C. apiculata Schiffner (1890: 36), C. calyptrifolia (Hooker 1813: tab. 43) Schiffner (1890: 35), C. ceratophora (Nees in Gottsche et al. 1845b: 405) Schiffner (1890: 35), C. clavigera Schiffner (1890: 35), C. corynephora (Nees, Lindenb. et Gottsche in Gottsche et al. 1843: 474) Schiffner (1890: 35), C. minor Schiffner (1890: 36), C. naumannii Schiffner (1890: 36), C. sagittistipula (Spruce 1884: 304) Schiffner (1890: 35), C. superba (Montagne 1848: 115) Schiffner (1890: 36), C. zollingeri Schiffner (1890: 35), Dendrolejeunea (Spruce 1884: 110) Schiffner (1890: 25), D. vittata (Mitten 1873: 411) Schiffner (1890: 25), Harpalejeunea (Spruce 1884: 164) Schiffner (1890: 29), H. decurvicuspis (Bescherelle & Massalongo 1886: 639) Schiffner (1890: 29), H. massalongoana Schiffner (1890: 29), H. savatieriana (Bescherelle & Massalongo 1886: 638) Schiffner (1890: 29), H. subfenestrata (Massalongo 1885: 249) Schiffner (1890: 28), Hygrolejeunea (Spruce 1884: 124) Schiffner (1890: 35), H. amboinensis Schiffner (1890: 31), H. latistipula Schiffner (1890: 30), Lopholejeunea (Spruce 1884: 119) 154 • Phytotaxa 220 (2) © 2015 Magnolia Press SÖDERSTRÖM ET AL. Schiffner (1890: 28), L. eulopha (Taylor 1846: 391) Schiffner (1890: 28), L. fimbriata (Gottsche 1880: 64) Schiffner (1890: 28), L. sagrana (Montagne 1842: 464) Schiffner (1890: 27), Mastigolejeunea (Spruce 1884: 100) Schiffner (1890: 22), M. amboinensis Schiffner (1890: 22), M. atypos Schiffner (1890: 22), M. convoluta (Lindenb. in Gottsche et al. 1845a: 288) Schiffner (1890: 21), M. minuta Schiffner (1890: 23), Phragmolejeunea Schiffner (1890: 24), P. polymorpha (Sande Lacoste 1854: 420) Schiffner (1890: 25), Pycnolejeunea (Spruce 1884: 246) Schiffner (1890: 32), P. connivens Schiffner (1890: 32), P. schwaneckei Schiffner (1890: 32), P. ventricosa Schiffner (1890: 32). Stephani (1890d; Nov) is an enumeration of liverworts from western Indian Ocean Islands. There is no reference to Spruce in the paper and nothing that can be taken as a description of a new generic name. However, three new generic names (Acrolejeunea, Cheilolejeunea and Lopholejeunea) are established here by combined generic-specific descriptions. We agree with Zijlstra (1982) and Grolle (1983) who consider Lopholejeunea as established here. Two binomials at specific rank are validly published or combined under generic names that were validly published earlier. Three names are effectively but not validly published. In the index the new names are referenced under both Lejeunea and the appropriate “Xxxlejeunea” entries (see below). Names validly published:—Acrolejeunea (Spruce 1884: 115) Stephani (1890d: 286), A. parviloba Stephani (1890d: 286), Cheilolejeunea (Spruce 1884: 251) Stephani (1890d: 284), C. kurzii Stephani (1890d: 284), Eulejeunea ecarinata Stephani (1890d: 283), Lopholejeunea (Spruce 1884: 119) Stephani (1890d: 285), L. multilacera Stephani (1890d: 285), Taxilejeunea conformis Nees et Mont. in Montagne (1843: 260) Stephani (1890d: 283). Names not validly published:—Ceratolejeunea (Spruce 1884: 198) Stephani (1890d: 284, 285, 286), C. mascarena Stephani (1890d: 284), C. mauritiana Stephani (1890d: 285), C. renauldii Stephani (1890d: 286). Prantl (1890, Nov. earliest) is the register to Hedwigia vol. 29. Generic names are given as “Xxxlejeunea” names, but all binomials at specific rank are under Lejeunea. They are thus valid combinations in Lejeunea when pointing back to validly published names, or validating a name not validly published through a direct reference to an effectively published description. Thirteen binomials are validly published here, but as many as six of them are illegitimate homonyms, again showing that Stephani did not clearly distinguish between genera and subgenera. Names validly published:—Lejeunea acroloba Prantl (1890: xiv), L. amazonica Prantl (1890: xiv) nom. illeg. (ICN Art. 53.1; non Spruce 1884: 106), L. bidens Prantl (1890: xiv), L. amplectens (Stephani 1886: 86) Prantl (1890: xiv), L. hampeana Prantl (1890: xvi) nom. illeg. (ICN Art. 53.1; non Lindenb. in Gottsche et al. 1845b: 360), L. junghuhniana Prantl (1890: xvi), L. kegelii Prantl (1890: xvi) nom. illeg. (ICN Art. 53.1; non Lindenberg & Gottsche 1851: 635), L. microstipula Prantl (1890: xvi), , L. miradorensis Prantl (1890: xvi), L. montagnei Prantl (1890: xvi) nom. illeg. (ICN Art. 53.1; non Montagne 1843: 261), L. sandvicensis Prantl (1890: xvii), L. stricta Prantl (1890: xvii) nom. illeg. (ICN Art. 53.1; non Lindenb. et Gottsche in Gottsche et al. 1875: 756), L. taitica (Gottsche 1864: 147) Prantl (1890: xvii). and L. zacuapana (Stephani 1890b: 87) Prantl (1890: xviii). Names not validly published:—None in Lejeuneaceae. Coulter et al. (1890, Nov. earliest) is the register to Botanical Gazette vol. 15. New names are referenced under both Lejeunea and the appropriate “Xxxlejeunea”, in both cases indicating new names by using bold. They should then be regarded as valid combinations under Lejeunea since there are direct references to the basionyms. For not validly published “Xxxlejeunea” names, i.e. the three Ceratolejeunea, the supposed basionym is not validly published, but as the index makes direct references to the effectively published descriptions, it must be regarded as validating the names under Lejeunea (cf. ICN Art. 38.1(a)). The author(s) of the index is unclear although it probably was the editors (J.M. Coulter, C.R. Barnes and/or J.C. Arthur). As they at least should have been responsible for the index, we ascribe the names to them. Seven binomials in Lejeunea are validly published by a direct reference to an effectively published description or combined here while three names in Ceratolejeunea are not validly published. Names validly published:—Lejeunea ecarinata (Stephani 1890d: 283) Coulter et al. (1890: 349), L. kurzii (Stephani 1890d: 284) Coulter et al. (1890: 349), L. mascarena Coulter et al. (1890: 349), L. mauritiana Coulter et al. (1890: 349), L. multilacera (Stephani 1890d: 285) Coulter et al. (1890: 349), L. parviloba (Stephani 1890d: 286) Coulter et al. (1890: 349) nom. illeg. [ICN Art. 53.1; non Ångström (1877: 87)] and L. renauldii Coulter et al. (1890: 349). Names not validly published:—Ceratolejeunea (Coulter et al. 1890: 348) mascarena Coulter et al. (1890: 348), C. mauritiana Coulter et al. (1890: 348) and C. renauldii Coulter et al. (1890: 348). Stephani (1891a; Sep–Oct) is the first paper in a second series on African Hepatics. For the most part it is a list of specimens gathered by various collectors. The new species are fully described and for taxa previously published in other genera the author of the basionym is cited and thus the combinations are validly published. There is no reference to Spruce and nothing that can be taken as a description of a new generic name. Zijlstra (1982) considered five genera (Ceratolejeunea, Cololejeunea, Colurolejeunea, Eulejeunea and Drepanolejeunea) established in this and the following paper by a combined generic-specific descriptions, but we consider one of them, Eulejeunea, established earlier. We also cannot accept that Ceratolejeunea is validly published here by a combined genericspecific description as Stephani mentioned two binomials at specific rank under that generic name on page 202. We also accept Mastigolejeunea as validly published by a combined generic-specific description here as we reject the supposed generic-specific description in Stephani (1889b: 257) as not fulfilling the requirements of ICN Art. 38.5 (more than one taxon accepted). Seven GENERA OF LEJEUNEACEAE ESTABLISHED IN 1884–1893 Phytotaxa 220 (2) © 2015 Magnolia Press • 155 binomials at specific rank under six generic names are validly described or combined as the genera were validly published earlier. Five binomials at specific rank under four generic names are effectively but not validly published. The index has names both as Lejeunea and “Xxxlejeunea”. The new Lejeunea names are validly combined in the index (see below). Names validly published:—Cololejeunea (Spruce 1884: 251) Stephani (1891a: 208), C. elegans Stephani (1891a: 208), Colurolejeunea (Spruce 1884: 303) Stephani (1891a: 208), C. obtusa (Stephani 1891a: 208), Drepanolejeunea (Spruce 1884: 186) Stephani (1891a: 209), D. cristata Stephani (1891a: 209), Eulejeunea saccatiloba Stephani (1891a: 207), Homalolejeunea excavata (Mitten 1860a: 58) Stephani (1891a: 202), Lopholejeunea sagrana (Montagne 1842: 464) Stephani (1891a: 202), Mastigolejeunea (Spruce 1884: 100) Stephani (1891a: 206), M. nigra Stephani (1891a: 206), Taxilejeunea dusenii Stephani (1891a: 209), T. epiphyta Stephani (1891a: 206). Names not validly published:—Ceratolejeunea (Spruce 1884: 198) Stephani (1891a: 202, 207), C. diversicornua Stephani (1891a: 207), C. mascarena Stephani (1891a: 202), Eulejeunea cuculliloba Stephani (1891a: 202; ICN Art. 38.1(a); no description; probably an error for E. saccatiloba), Hygrolejeunea (Spruce 1884: 230) Stephani (1891a: 208), H. pulcherrima Stephani (1891a: 202), also ICN Art. 38.1(a); no description, Ptycholejeunea (Spruce 1884: 97) Stephani (1891a: 203), P. striata (Lehm. et Lindenb. in Lehmann 1833: 16) Stephani (1891a: 203); also ICN Art. 38.1(a); no description. Renauld & Cardot (1891) clearly used the subgeneric concept. Stephani (1891b; Nov–Dec) is the second paper in the second series on African hepatics. The typography is the same as in the first part. One generic name (Leptolejeunea) is validly published here by a combined generic-specific description and one additional binomial at specific rank is validly published since the genus was established earlier. Names validly published:—Cololejeunea crenatiflora Stephani (1891b: 270), Leptolejeunea (Spruce 1884: 193) Stephani (1891b: 270), L. quintasii Stephani (1891b: 270). Names not validly published:—None. Prantl (1891, Nov. earliest) is the register to Hedwigia vol. 30. Names are listed under both Lejeunea and “Xxxlejeunea”. For not validly published “Xxxlejeunea” names the supposed basionym is not validly published, but as the index makes direct references to the effectively published descriptions, it must be regarded as validating the names under Lejeunea. Eight binomials at specific rank are validly published, but two of them are later homonyms. Two generic names and five binomials at specific rank are effectively but not validly published. Names validly published:—Lejeunea cristata (Steph.) Prantl (1891: ix), L. dusenii (Steph.) Prantl (1891: ix), L. elegans Prantl (1891: ix) nom. illeg. (ICN Art. 53.1; non Gottsche in Gottsche et al. 1845b), L. epiphyta (Steph.) Prantl (1891: ix) nom. illeg. (ICN Art. 53.1; non Gottsche in Gottsche et al. 1845b), L. nigra (Steph.) Prantl (1891: ix), L. obtusa Prantl (1891: ix), L. quintasii (Steph.) Prantl (1891: ix), L. saccatiloba (Steph.) Prantl (1891: ix). Names not validly published:—Ceratolejeunea (Spruce 1884: 198) Prantl (1891: vi), C. diversicornua Prantl (1891: vi), C. mascarena Prantl (1891: vi), Eulejeunea cuculliloba Prantl (1891: viii), Hygrolejeunea Prantl (1891: ix), H. pulcherrima Prantl (1891: ix), Ptycholejeunea Prantl (1891: xiii), P. striata (Lehm. et Lindenb. in Lehmann 1833: 16) Prantl (1891: xiii). Stephani (1891c; Jan. 1892) is the treatment of hepatics in Renauld & Cardot’s paper on new or little known bryophytes. All species are described using their “Xxxlejeunea” names only and there is no reference to Spruce or any description of a new generic name. No new generic names and only one binomial at specific rank is validly published. Three binomials at specific rank are effectively but not validly published. Names validly published:—Acrolejeunea borgenii Stephani (1892a: 201). Names not validly published:—Ceratolejeunea (Spruce 1884: 198) Stephani (1891c: 203), C. mascarena Stephani (1892a: 203), C. mauritiana Stephani (1892a: 204), C. renauldii Stephani (1892a: 204). Jack & Stephani (1892; Jan–Apr) is primarily a list of species collected by Gustav Wallis in South America and the Philippines. All new species are described, most jointly by Jack and Stephani, but in a few cases by Stephani only. Although they state that all names are in alphabetic order, and place all “Xxxlejeunea” where they should be if treated as Lejeunea, they do not refer at all to Spruce. Further on, within the section of “Lejeunea”, the “Xxxlejeunea” names are alphabetic. The paper starts with a summary of the taxa collected in Philippines and South America, respectively, and all new taxa are then further described. There is no reference to Spruce and no trace of any description of a generic name. Two generic names, Ceratolejeunea and Thysanolejeunea, are each validly published by a combined generic-specific description which is in agreement with Zijlstra (1982), but which Bonner et al. (1961) considered validly published earlier. In addition there are five binomials at specific rank validly published in genera validly published earlier. Seven generic names and 15 binomials at specific rank are effectively but not validly published. New names and new combinations in Lejeunea appear in the index (see below). Names validly published:—Ceratolejeunea (Spruce 1884: 198) Jack & Stephani (1892: 16), C. grandiloba Jack & Stephani (1892: 16), Drepanolejeunea capulata (Taylor 1846: 394) Jack & Stephani (1892: 13), D. dactylophora (Gottsche et al. 1843: 473) Jack & Stephani (1892: 12), D. lancifolia (Gottsche 1864: 155) Jack & Stephani (1892: 13), D. tenuis (Nees 1830: 57) Jack & Stephani (1892: 12), Taxilejeunea apiculata (Gottsche 1864: 149) Jack & Stephani (1892: 13), Thysanolejeunea (Spruce 1884: 105) Jack & Stephani (1892: 20), T. gottschei Jack & Stephani (1892: 20). 156 • Phytotaxa 220 (2) © 2015 Magnolia Press SÖDERSTRÖM ET AL. Names not validly published:—Crossotolejeunea (Spruce 1884: 161) Jack & Stephani (1892: 16), C. inflexiloba Jack & Stephani (1892: 16), C. intricata Jack & Stephani (1892: 17), Dicranolejeunea (Spruce 1884: 138) Jack & Stephani (1892: 13), D. axillaris (Nees & Montagne 1836: 59) Jack & Stephani (1892: 13), D. dubiosa (Lindenberg & Gottsche 1846: 752) Jack & Stephani (1892: 13), Drepanolejeunea granatensis Jack & Stephani (1892: 13; ICN Art. 38.1(a); no description, but probably an error for Harpalejeunea granatensis), Harpalejeunea (Spruce 1884: 164) Jack & Stephani (1892: 13, 17, 18), H. cinchonae (Nees in Gottsche et al. 1845b: 342) Jack & Stephani (1892: 13), H. granatensis Jack & Stephani (1892: 17), H. tuberculata Jack & Stephani (1892: 18), Hygrolejeunea (Spruce 1884: 230) Jack & Stephani (1892: 13), H. reflexistipula (Lehm. et Lindenb. in Lehmann 1833: 10) Jack & Stephani (1892: 13), Peltolejeunea (Spruce 1884: 85) Jack & Stephani (1892: 13, 18, 19), P. jackii Steph. in Jack & Stephani (1892: 18), P. ovalis (Lindenb. et Gottsche in Gottsche et al. 1847: 754) Jack & Stephani (1892: 13), P. wallisii Jack & Stephani (1892: 19), Prionolejeunea (Spruce 1884: 152) Jack & Stephani (1892: 13), P. fabroniifolia (Spruce 1884: 154) Jack & Stephani (1892: 13), Strepsilejeunea (Spruce 1884: 168) Jack & Stephani (1892: 13, 19), S. inflexa (Hampe in Lehmann 1838: 22) Jack & Stephani (1892: 13), S. laevicalyx Jack & Stephani (1892: 19). Bescherelle (1892) is a list of species collected in Vietnam. There are no descriptions although several taxa are marked as “sp. n.”. Two binomials at specific rank under generic names that were validly published earlier are validly published here. Three generic names and five binomials at specific rank are effectively but not validly published. There is no index. Names validly published:—Cololejeunea trichomanis (Gottsche 1882: 362) Bescherelle (1892: 14), Leptolejeunea elliptica (Lehm. et Lindenb. in Lehmann 1833: 13) Bescherelle (1892: 14). Names not validly published:—Caudalejeunea Bescherelle (1892: 13), C. stephanii Bescherelle (1892: 13) also not validly published by ICN Art. 38.1(a); no description, Cololejeunea tonkinensis Steph. ex Bescherelle (1892: 14) ICN Art. 38.1(a); no description, Eulejeunea infestans Steph. ex Bescherelle (1892: 13) ICN Art. 38.1(a); no description, Leptolejeunea spicata Steph. ex Bescherelle (1892: 14) ICN Art. 38.1(a); no description, Ptycholejeunea (Spruce 1884: 97) Bescherelle (1892: 14), P. piriformis (Gottsche ex Stephani 1889b: 259) Bescherelle (1892: 14) ICN Art. 41.1; basionym not valid, Pycnolejeunea (Spruce 1884: 246) Bescherelle (1892: 14), P. meyeniana (Nees, Lindenb. et Gottsche in Gottsche et al. 1843: 472) Bescherelle (1892: 14). Stephani (1892a; May–Jun) is the third paper on African hepatics and follows the others in style. No new generic name is validly published here, but two binomials at specific rank are validly published. Two generic names with one binomial at specific rank each are effectively but not validly published. The index has Lejeunea names only. Names validly published:—Eulejeunea isomorpha (Gottsche 1882: 355) Stephani (1892a: 120), E. wilmsii Stephani (1892a: 124). Names not validly published:—Anomalolejeunea (Pearson 1887: 5) Stephani (1892a: 120), A. pluriplicata (Pearson 1887: 5) Stephani (1892a: 120), Ptycholejeunea (Spruce 1884: 97) Stephani (1892a: 120), P. striata (Lehm. et Lindenb. in Lehmann 1832: 16) Stephani (1892a: 120). Stephani (1892b; Jun) is a treatment of North American Lejeuneaceae. Stephani consistently used hyphens in the “Xxx-Lejeunea” names. He did not refer to Spruce and did not indicate the ranks except when writing about a couple of Lejeunea names that he could not place in any “suborder” [sic!]. No generic name is validly published here, but ten binomials at specific rank under four generic names are validly published. One generic name and two binomials under specific rank are effectively but not validly published. The index only gives Lejeunea entries and these can be treated as combinations where a Lejeunea combination does not exist (see below). Names validly published:—Cololejeunea calcarea (Libert 1820: 373) Stephani (1892b: 171), C. jooriana (Austin 1875: 20) Stephani (1892b: 171), C. minutissima (Sm. in Smith & Sowerby 1806: 1633) Stephani (1892b: 171), C. parvula (Wilson et Hook. in Wilson 1841: no. 172) Stephani (1892b: 172), Eulejeunea austinii (Lindberg 1875: 489) Stephani (1892b: 171), E. caroliniana (Austin 1876: 36) Stephani (1892b: 171), E. underwoodii Stephani (1892b: 172), Mastigolejeunea auriculata (Wilson. et Hook. in Wilson 1841: no. 170) Stephani (1892b: 171), Microlejeunea cardotii Stephani (1892b: 172), M. lucens (Taylor 1846: 399) Stephani (1892b: 171). Names not validly published:—Cololejeunea wrightii Stephani (1892b: 172); ICN Art. 38.1(a); no description, Neurolejeunea (Spruce 1884: 84) Stephani (1892b: 171), N. catenulata (Nees in Gottsche et al. 1845b: 323) Stephani (1892b: 171). Stephani (1892c; Jul–Aug) is a continuation of the African hepatic series in the same style as the preceding parts. No new generic name is validly published here, but seven binomials at specific rank are validly published. Two generic names, four binomials at specific rank and a variety are effectively but not validly published. Names validly published:—Acrolejeunea confertissima Stephani (1892c: 165), Ceratolejeunea cornutissima Stephani (1892c: 166), Cololejeunea cuneifolia Stephani (1892c: 166), C. filicaulis Stephani (1892c: 168), Colurolejeunea dusenii Stephani (1892c: 168), Leptolejeunea truncatiloba Stephani (1892c: 170), Mastigolejeunea turgida Stephani (1892c: 170). Names not validly published:—Hygrolejeunea (Spruce 1884: 230) Stephani (1892c: 169), H. lyratiflora Stephani (1892c: 169), H. papilionacea Stephani (1892c: 169), Odontolejeunea sieberiana var. africana Stephani (1892c: 171), Prionolejeunea (Spruce 1884: 152) Stephani (1892c: 171), P. denticulata (Weber 1815: 30) Stephani (1892c: 171), P. kindbergii Stephani (1892c: 171). Pearson (1892; Aug–Sep) is an enumeration of species from Madagascar. It is mainly a list of species recorded, but the new species are described properly under their “Xxxlejeunea” names. He refers to Spruce’s subgenera, but nowhere is he indicating that he follows GENERA OF LEJEUNEACEAE ESTABLISHED IN 1884–1893 Phytotaxa 220 (2) © 2015 Magnolia Press • 157 that rank. In a quote from Spruce, Pearson mentions a new subg. Callistolejeunea, which is not valid as it is not definitely connected with a genus (ICN Art. 35.2). References to previously validly published names makes combinations valid where the genus is already established. Nineteen binomials at specific rank under nine generic names are validly published. Six generic names and eight binomials at specific rank are effectively but not validly published. Names validly published:—Acrolejeunea abnormis (Gottsche 1882: 352) Pearson (1892: 4), A. fulva (Gottsche 1882: 550) Pearson (1892: 4), A. inflexa (Gottsche 1882: 351) Pearson (1892: 4), Archilejeunea nobilis (Gottsche 1882: 353) Pearson (1892: 4), Ceratolejeunea variabilis (Lindenb. in Gottsche et al. 1845b: 399) Pearson (1892: 8), Cololejeunea marginata (Lehm. et Lindenb. in Lehmann 1833: 11) Pearson (1892: 9), C. punctata (Gottsche 1882: 361) Pearson (1892: 9), Drepanolejeunea physifolia (Gottsche 1882: 357) Pearson (1892: 8), D. securifolia (Gottsche 1882: 360) Pearson (1892: 8), Eulejeunea obtusata (Gottsche 1882: 354) Pearson (1892: 8), E. vulgariformis (Gottsche 1882: 355) Pearson (1892: 9), Euosmolejeunea brachytoma (Gottsche 1882: 355) Pearson (1892: 8), Lopholejeunea fimbriata (Gottsche in Müller 1880: 64) Pearson (1892: 5), L. harpaphylla (Spruce 1884: 123) Pearson (1892: 7), L. lepidoscypha Kier et Pearson in Pearson (1892: 5), L. rusbyi (Spruce 1890: 122) Pearson (1892: 6), Microlejeunea byssoides (Gottsche 1882: 35) Pearson (1892: 9), M. oblongistipula (Gottsche 1882: 337) Pearson (1892: 9), M. papulosa (Gottsche 1882: 358) Pearson (1892: 9). Names not validly published:—Harpalejeunea (Spruce 1884: 164) Pearson (1892: 7), H. solitaria (Gottsche 1882: 356) Pearson (1892: 7), Omphalolejeunea (Spruce 1884: 86) Pearson (1892: 4), O. filiformis (Swartz 1788: 144) Pearson (1892: 4), Prionolejeunea (Spruce 1884: 152) Pearson (1892: 7), P. rutenbergiana (Gottsche 1882: 358) Pearson (1892: 7), P. grata (Gottsche 1882: 359) Pearson (1892: 7), Ptycholejeunea (Spruce 1884: 97) Pearson (1892: 4), P. striata (Lehm. et Lindenb. in Lehmann 1832: 16) Pearson (1892: 4), Pycnolejeunea (Spruce 1884: 246) Pearson (1892: 8), P. silvestris (Gottsche 1882: 354) Pearson (1892: 8), Strepsilejeunea (Spruce 1884: 168) Pearson (1892: 7), S. brevifissa (Gottsche 1882: 356) Pearson (1892: 7), S. cardiophylla (Gottsche 1882: 356) Pearson (1892: 7). Stephani (1892d; Sep–Oct) is the last part of series on the African hepatics and follows the others in style. No new generic name is validly published, but three binomials at specific rank are validly published. Names validly published:—Ceratolejeunea mascarena (Coulter et al. 1890: 349) Stephani (1892e: 204), C. mauritiana (Coulter et al. 1890: 349) Stephani (1892e: 205), C. renauldii (Coulter et al. 1890: 349) Stephani (1892e: 205). Names not validly published:—None. Prantl (1892, Nov. earliest) is the register to Hedwigia vol. 31. It mentions no name under “Xxxlejeunea”, but gives the subgeneric placement under the Lejeunea entries. Further, the Lejeunea entries are ordered alphabetically under their subgeneric placement, not after the binomials at specific rank. Nineteen binomials at specific rank are validly published here by the direct reference to the effective description, but four of them are illegitimate as later homonyms. Names validly published:—Lejeunea apiculata (Gottsche 1864: 149) Prantl (1892: xvii) nom. illeg. (ICN Art. 53.1; non Sande Lacoste 1854: 421), L. cuneifolia (Stephani 1892c: 166) Prantl (1892: xvi), L. dusenii (Stephani 1892c: 168) Prantl (1892: xvi) nom. illeg. (ICN Art. 53.1; non Prantl 1891: ix), L. filicaulis (Stephani 1892c: 168) Prantl (1892: xvi), L. gottschei (Jack & Stephani 1892: 20) Prantl (1892: xvii), L. granatensis Prantl (1892: xvi), L. grandiloba (Jack & Stephani 1892: 16) Prantl (1892: xvi), L. inflexiloba Prantl (1892: xvi), L. intricata Prantl (1892: xvi), L. jackii Prantl (1892: xvii), L. kindbergii Prantl (1892: xvii), L. laevicalyx Prantl (1892: xvii) nom. illeg. (ICN Art. 53.1; non Gottsche 1863: 221), L. lyratiflora Prantl (1892: xvi), L. papilionacea Prantl (1892: xvii), L. truncatiloba (Stephani 1892c: 170) Prantl (1892: xvii), L. tuberculata Prantl (1892: xvi), L. turgida (Stephani 1892c: 170) Prantl (1892: xvii) nom. illeg. (ICN Art. 53.1; non Mitten 1860b: 110), L. wallisii Prantl (1892: xvii), L. wilmsii (Stephani 1892a: 124) Prantl (1892: xvi). Names not validly published:—None. Coulter et al. (1892, Nov. earliest) is the register to Botanical Gazette vol. 17. It gives only entries under Lejeunea. It is not clear who compiled the index (although the editors J.M. Coulter, C.R. Barnes and/or J.C. Arthur were at least responsible for the index), so we prefer to refer to them. Two binomials at specific rank are validly published here. Names validly published:—Lejeunea cardotii (Stephani 1892b: 72) Coulter et al. (1892: 431), L. underwoodii (Stephani 1892b: 172) Coulter et al. (1892: 431). Names not validly published:—None. Bescherelle (1893a) clearly treated all Lejeunea as subgenera. Bescherelle (1893b) is a short list of hepatics from Brazzaville. It uses the hyphened “Xxx-Lejeunea”. One taxon is validly published here since the generic name is validly published earlier. Names validly published:—Hygrolejeunea pulcherrima (Stephani 1886: 87) Bescherelle (1893b: 60). Names not validly published:—None. Stephani (1892e; Jul. 1893) is a list of specimens collected in Costa Rica. The “Xxxlejeunea” are written in the generic form, without reference to Lejeunea except for a couple of species ascribed to Spruce (1884), where they were combined under their Lejeunea names. Zijlstra (1982) argued that as the list is generally alphabetic and that their placement where Lejeunea should occur indicates 158 • Phytotaxa 220 (2) © 2015 Magnolia Press SÖDERSTRÖM ET AL. that Stephani regarded them as subgenera. However, as the “Xxxlejeunea” are clearly written like genera and there is no reference to any subgenus, we regard them as genera, as Stephani by this time generally seems to have done. No new generic name is established here. There are two new binomials at specific rank in two generic names validly published here. Six generic names and eight binomials at specific rank are effectively but not validly published. Names validly published:—Ceratolejeunea brasiliensis (Gottsche in Gottsche et al. 1845b: 398) Stephani (1892e: 179), Drepanolejeunea inchoata (C.F.W.Meissn. in Lehmann 1838: 19) Stephani (1892e: 179). Names not validly published:—Bryolejeunea (Spruce 1884: 111) Stephani (1892e: 179), B. diffusa (Swartz 1788: 144) Spruce ex Stephani (1892e: 179), B. flaccida (Lindenb. et Hampe in Hampe 1851: 640) Stephani (1892e: 179), B. tenuicaulis (Taylor ex Gottsche et al. 1845a: 285) Stephani (1892e: 179), Dicranolejeunea (Spruce 1884: 128) Stephani (1892e: 179), D. axillaris (Nees & Montagne 1836: 59) Stephani (1892e: 179), Diplasiolejeunea (Spruce 1884: 301) Stephani (1892e: 179), D. pellucida (C.F.W.Meissn. ex Sprengel 1827: 325) Stephani (1892e: 179), Hygrolejeunea (Spruce 1884: 230) Stephani (1892e: 179), H. phyllobola (Nees et Mont. in Montagne 1842: 47) Stephani (1892e: 179), Mastigolejeunea crispula (Stephani 1888b: 111) Stephani (1892e: 179) ICN Art. 41.1; basionym not valid, Odontolejeunea (Spruce 1884: 142) Stephani (1892e: 180), O. sieberiana (Gottsche in Gottsche et al. 1845b: 328) Stephani (1892e: 180), Omphalolejeunea (Spruce 1884: 86) Stephani (1892e: 180), O. filiformis (Swartz 1788: 144) Stephani (1892e: 180). Schiffner (1893b; Sep) is the first author who clearly and intentionally used the Spruce’s subgenera as genera. Here 19 out of the 23 remaining subgeneric names are validly published at generic level together with a large number of binomials at specific rank. Names validly published:—Acrolejeunea fertilis (Reinwardt et al. 1824: 211) Schiffner (1893b: 128), A. hasskarliana (Gottsche in Gottsche et al. 1845a: 299) Schiffner (1893b: 128), A. polycarpa (Nees 1833: 350) Schiffner (1893b: 128), A. pycnoclada (Taylor 1846) Schiffner (1893b: 128), A. torulosa (Lehm. et Lindenb. in Lehmann 1834: 41) Schiffner (1893b: 128), Anomalolejeunea (Spruce in Pearson 1887: 5) Schiffner (1893b: 127), A. pluriplicata (Pearson 1887: 2) Schiffner (1893b: 127), Anoplolejeunea (Spruce 1884: 129) Schiffner (1893b: 131), A. herpestica (Spruce 1884: 129) Schiffner (1893b: 131), Archilejeunea leprieurii (Montagne 1835: 213) Schiffner (1893b: 130), Brachiolejeunea (Spruce 1884: 129) Schiffner (1893b: 128), B. bicolor (Nees 1833: 349) Schiffner (1893b: 128), B. laxifolia (Taylor 1847: 341) Schiffner (1893b: 128), B. spruceana (Massalongo 1885: 246) Schiffner (1893b: 128), Caudalejeunea Steph. ex Schiffner (1893b: 128), C. africana (Stephani 1886: 93) Schiffner (1893b: 129), C. crescentiae (Lindenb. et Gottsche in Gottsche et al. 1847: 752) Schiffner (1893b: 129), C. haenkeana (Schiffner 1886: 210) Schiffner (1893b: 129), C. hanningtonii (Mitten 1886: 324) Schiffner (1893b: 129), C. harpaphylla (Spruce 1884: 123) Schiffner (1893b: 129), C. recurvistipula (Gottsche in Gottsche et al. 1845b: 326) Schiffner (1893b: 129), Ceratolejeunea breutelii (Gottsche in Gottsche et al. 1845b: 324) Schiffner (1893b: 125), C. coarina (Gottsche in Gottsche et al. 1845b: 395) Schiffner (1893b: 125), C. cubensis (Montagne 1842: 481) Schiffner (1893b: 125), C. desciscens (Sande Lacoste 1856: 107) Schiffner (1893b: 126), Cheilolejeunea heteroclada (Spruce 1884: 256) Schiffner (1893b: 124), C. lineata (Lehm. et Lindenb. in Lehmann 1832: 53) Schiffner (1893b: 124), C. newtonii Steph. ex Schiffner (1893b: 124), C. novoguinensis (Schiffner 1893a: 258) Schiffner (1893b: 124), C. papulosa Schiffner (1893b: 124) Schiffner (1893b: 124), Cololejeunea microscopica (Taylor 1836: 59) Schiffner (1893b: 122), C. rossettiana (Massalongo 1889: 487) Schiffner (1893b: 122), C. venusta (Sande Lacoste 1854) Schiffner (1893b: 122), Colurolejeunea calyptrifolia (Hooker 1813: tab. 43) Schiffner (1893b: 121), C. naumannii Schiffner (1893b: 121), C. ornata (Goebel 1890: 26) Schiffner (1893b: 121), Crossotolejeunea (Spruce 1884: 161) Schiffner (1893b: 127), C. boryana (Montagne 1838: 47) Schiffner (1893b: 127), Dicranolejeunea (Spruce 1884: 138) Schiffner (1893b: 128), D. axillaris (Nees & Montagne 1836: 59) Schiffner (1893b: 128), D. phyllorhiza (Nees 1833: 348) Schiffner (1893b: 128), Diplasiolejeunea (Spruce 1884: 301) Schiffner (1893b: 121), D. pellucida (Sprengel 1827: 325) Schiffner (1893b: 121), Drepanolejeunea hamatifolia (Hooker 1813: tab. 51) Schiffner (1893b: 126), D. muricata (Gottsche in Gottsche et al. 1845b: 348) Schiffner (1893b: 126), D. palmifolia (Nees 1833: 366) Schiffner (1893b: 126), D. ternatensis (Gottsche in Gottsche et al. 1845b: 346) Schiffner (1893b: 126), Eulejeunea diversiloba (Spruce 1876b: 235) Schiffner (1893b: 124), E. ulicina (Taylor 1844: 115) Schiffner (1893b: 124), Harpalejeunea (Spruce 1884: 164) Schiffner (1893b: 127), H. ancistrodes (Spruce 1884: 169) Schiffner (1893b: 127), H. cinchonae (Nees in Gottsche et al. 1845b: 342) Schiffner (1893b: 127), H. holostipa (Spruce 1884: 171) Schiffner (1893b: 127), H. ovata (Hooker 1813: tab. 42) Schiffner (1893b: 127), H. tenuicuspis (Spruce 1884: 170) Schiffner (1893b: 127), Hygrolejeunea (Spruce 1884: 230) Schiffner (1893b: 124), H. cerina (Lehm. et Lindenb. in Lehmann 1833: 16) Schiffner (1893b: 125), H. reflexistipula (Lehm. et Lindenb. in Lehmann 1833: 10) Schiffner (1893b: 125), H. sordida (Nees 1830: 41) Schiffner (1893b: 125), H. spongia (Spruce 1876a: 132) Schiffner (1893b: 125), H. umbilicata (Nees 1830: 42) Schiffner (1893b: 125), Leptolejeunea corynophora (Gottsche et al. 1843: 474) Schiffner (1893b: 126), L. vitrea (Nees 1830: 56) Schiffner (1893b: 126), Lopholejeunea applanata (Reinwardt et al. 1824: 210) Schiffner (1893b: 129), L. eulopha (Taylor 1846: 391) Schiffner (1893b: 129), L. javanica (Nees in Gottsche et al. 1845b: 320) Schiffner (1893b: 129), Macrolejeunea (Spruce 1884: 224) Schiffner (1893b: 125), M. subsimplex (Mont. et Nees in Montagne 1843: 264) Schiffner (1893b: 125), Mastigolejeunea buettneri (Stephani 1888b: 110) Schiffner (1893b: 129), M. humilis (Gottsche in Gottsche et al. 1845a: 239) Schiffner (1893b: 129), M. ligulata (Lehm. et Lindenb. in Lehmann 1834: 39) Schiffner (1893b: 129), M. phaea Schiffner (1893b: 129), Neurolejeunea (Spruce 1884: 84) Schiffner (1893b: 131), N. catenulata (Nees in Gottsche et al. 1845b: 323) Schiffner GENERA OF LEJEUNEACEAE ESTABLISHED IN 1884–1893 Phytotaxa 220 (2) © 2015 Magnolia Press • 159 (1893b: 131), N. portoricensis (Hampe & Gottsche 1853: 352) Schiffner (1893b: 131), N. seminervis (Spruce 1884: 84) Schiffner (1893b: 131), Odontolejeunea (Spruce 1884: 142) Schiffner (1893b: 127), O. lunulata (Weber 1815: 33) Schiffner (1893b: 128), O. peruviana (Lehm. et Lindenb. in Lehmann 1833: 18) Schiffner (1893b: 128), O. sieberiana (Gottsche in Gottsche et al. 1845b: 328) Schiffner (1893b: 127), Otigoniolejeunea (Spruce 1884: 226) Schiffner (1893b: 125), O. schizotis (Spruce 1884: 227) Schiffner (1893b: 125), O. xiphotis (Spruce 1884: 227) Schiffner (1893b: 125), Peltolejeunea (Spruce 1884: 85) Schiffner (1893b: 131), P. jackii (Prantl 1892: xvii) Schiffner (1893b: 131), P. osculatiana (De Notaris 1855: 233) Schiffner (1893b: 131), P. ovalis (Lindenb. et Gottsche in Gottsche et al. 1847: 754) Schiffner (1893b: 131), P. pulcherrima (Stephani 1886: 87) Schiffner (1893b: 131), P. wallisii (Prantl 1892: xvii) Schiffner (1893b: 131), Platylejeunea (Spruce 1884: 124) Schiffner (1893b: 130), P. hobsoniana (Lindenb. in Gottsche et al. 1845b: 313) Schiffner (1893b: 130), P. vincentina (Gottsche in Gottsche et al. 1845b: 313) Schiffner (1893b: 130), Prionolejeunea (Spruce 1884: 152) Schiffner (1893b: 127), P. denticulata (Weber 1815: 30) Schiffner (1893b: 127), P. deplanata (Mitten 1886: 325) Schiffner (1893b: 127), P. fabroniifolia (Spruce 1884: 154) Schiffner (1893b: 127), P. grata (Gottsche 1882) Schiffner (1893b: 127), P. kindbergii (Coulter et al. 1892: xvii) Schiffner (1893b: 127), Pycnolejeunea (Spruce 1884: 246) Schiffner (1893b: 124), P. ceylanica (Gottsche in Gottsche et al. 1845a: 259) Schiffner (1893b: 124), P. imbricata (Nees 1830: 42), Schiffner (1893b: 124), P. macroloba (Nees et Mont. in Montagne 1843: 260) Schiffner (1893b: 124), P. trapezia (Nees 1830: 41) Schiffner (1893b: 124), Stictolejeunea (Spruce 1884: 81) Schiffner (1893b: 131), S. kunzeana (Gottsche et al. 1845b: 322) Schiffner (1893b: 131), S. squamata (Willd. in Weber 1815: 33) Schiffner (1893b: 131), Strepsilejeunea (Spruce 1884: 168) Schiffner (1893b: 127), S. kunthiana (Lindenb. in Gottsche et al. 1845b: 353) Schiffner (1893b: 127), S. mimosa (Hooker & Taylor 1844a: 398) Schiffner (1893b: 127), S. orba (Gottsche in Gottsche et al. 1845b: 350) Schiffner (1893b: 127), Trachylejeunea acanthina (Spruce 1884: 182) Schiffner (1893b: 126), T. asperiflora (Spruce 1884: 183) Schiffner (1893b: 126). Names not validly published:—Caudalejeunea stephanii Schiffner (1893b: 129) ICN Art. 38.1(a); no description, Drepanolejeunea blumei Steph. ex Schiffner (1893b: 126) ICN Art. 38.1(a); no description, Trachylejeunea prionocalyx Schiffner (1893b: 126) ICN Art. 38.1(a); no description. Generic names based on Lejeunea subgenera published between 1884 and 1893 Here we summarize the valid publication of each of the generic names based on Lejeunea subgenera published between 1884 and 1893. We give full nomenclature citation, the type information for all genera and comment on non-valid publications of the names. We mention under which genus the type species is placed if the genus is not accepted in the world checklist of hornworts and liverworts (Söderström et al. in press). For many of the names listed below the first published lectotypification statement was by A.W. Evans in a series of papers and floristic treatments published between 1903 and 1920 (Evans 1903b, 1904, 1906, 1907a, b, 1908a, 1911, 1912a, b, 1914, 1918, 1920). Evans adhered to the American Code of Botanical Nomenclature (Arthur et al. 1907), however, and consistently designated types according to the First Species Rule. Under the current Code, this method is ruled as mechanical and such typifications may be superseded under ICN Art. 10.5(b) and ICN Art. 10 *Example 7. We consider Evans’ typifications to have been superseded by the next effectively published typification, even when both type designations are the same. In many cases, the typifying author is Vanden Berghen (1945 [1948]). Acrolejeunea (Spruce) Steph., Bot. Gaz. 15: 286, 1890 (Stephani 1890d) nom. conserv. Basionym:—Lejeunea subg. Acrolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 115, 1884, ‘Acro-Lejeunea’ (Spruce 1884) ≡ Acrolejeunea (Spruce) Schiffn., Hepat. [Engl.Prantl]: 128, 1893 (Schiffner 1893b). Lectotype (Gradstein 1974: 332):—Jungermannia torulosa Lehm. et Lindenb. in Lehmann (1834: 41) ≡ Lejeunea torulosa (Lehm. et Lindenb.) Spruce (1884: 117) ≡ Acrolejeunea torulosa (Lehm. et Lindenb.) Schiffner (1893b: 128). Note:—The genus was established by the specific description of Acrolejeunea parviloba treated as a combined genericspecific one. Bonner et al. (1961) considered the genus established by Stephani (1890a: 7) because of direct reference of the name to the Sprucean subgenus (Art. 38.14), but in both the introductory and concluding paragraphs of this 3-part treatment Stephani (1890a, b, c) explicitly states that he considers the taxa to be subgenera, as already discussed by Gradstein et al. (1982), Zijlstra (1982) and Grolle (1983). Because of the controversial status of rank being used in this series of publications, we will propose that this series of publications be suppressed at the rank of genus. The generic name was used in five publications prior to its valid publication (Stephani 1888b: 107, 1889a: 164, 165, 1890a: 7, 1890c: 133, Schiffner 1890: 26). Anomalolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 127, 1893 (Schiffner 1893b). Basionym:—Lejeunea subg. Anomalolejeunea Spruce in Pearson, Skr. Vidensk.-Selsk. Christiana, Math.-Naturvidensk. Kl. 1887(9): 5, 1887 (Pearson 1887). Type (monotypic subgenus):—Lejeunea pluriplicata Pearson (1887: 5) ≡ Anomalolejeunea pluriplicata (Pearson) Schiffner (1893b: 127) ≡ Cheilolejeunea pluriplicata (Pearson) Schuster (1980b: 430). Note:—Stephani (1892b: 120) used but did not validly publish Anomalolejeunea. The type of the genus is now included in Cheilolejeunea (Spruce) Steph. 160 • Phytotaxa 220 (2) © 2015 Magnolia Press SÖDERSTRÖM ET AL. Anoplolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 131, 1893 (Schiffner 1893b). Basionym:—Lejeunea subg. Anoplolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 129, 1884, ‘Anoplo-Lejeunea’ (Spruce 1884). Type (monotypic subgenus):— Lejeunea herpestica Spruce (1884: 129) ≡ Anoplolejeunea herpestica (Spruce) Schiffner (1893b: 131) [= Anoplolejeunea conferta (C.F.W.Meissn. ex Sprengel 1827: 325) Evans (1908b: 175), cf. Evans (1908b)]. Archilejeunea (Spruce) Steph., Hedwigia 27: 113, 1888 (Stephani 1888b). Basionym:—Lejeunea subg. Archilejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 88, 1884, ‘Archi-Lejeunea’ (Spruce 1884). Lectotype (Vanden Berghen 1945 [1948]: 41):—Lejeunea porelloides Spruce (1884: 90) ≡ Archilejeunea porelloides (Spruce) Schiffner (1894: 181) ≡ Archilejeunea ludoviciana (De Not. in Lehmann 1857: 11) P.Geissler et Gradst. in Gradstein (1994: 2002) subsp. porelloides (Spruce) Gradstein (1994: 58). Note:— Stephani established the genus by the description of Archilejeunea erronea treated as a combined generic-specific description. It was regarded as a synonym of Leucolejeunea rotundistipula (Lindenb. ex Lehmann 1829: 360) Evans (1907c: 229) by Grolle (1983), so he indicated that it would have to be conserved against Stephani’s name. By the provisions of ICN Art. 41.4 this conservation is no longer necessary. Brachiolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 128, 1893 (Schiffner 1893). Basionym:—Lejeunea subg. Brachiolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 129, 1884, ‘Brachio-Lejeunea’ (Spruce 1884). Lectotype (Vanden Berghen 1945 [1948]: 43):—Phragmicoma laxifolia Taylor (1847: 341) ≡ Lejeunea laxifolia (Taylor 1847: 341) Spruce (1884: 130) ≡ Brachiolejeunea laxifolia (Taylor) Schiffner (1893b: 128). Note:—As also suggested in the current ICN Art. 41 Ex. 9, Bonner et al. (1961) considered Brachiolejeunea validly published by Stephani (1889a: 167) with a combined generic-specific description (ICN Art. 38.5). However, Stephani (1889a) enumerated several species in Brachiolejeunea in this treatment, so the requirements of ICN Art. 38.5 for a combined generic-specific description are not met. Conservation of Brachiolejeunea (Spruce 1884: 129) Schiffner (1893b: 128) against Brachiolejeunea (Spruce 1884: 129) Steph. et Spruce in Stephani (1889a: 167), as advocated by van Slageren (1985), is therefore unnecessary. The generic name was used three times (Stephani 1889a: 167, 1890a: 7, 1890c: 137) prior to its valid publication. Bryolejeunea (Spruce) Lacout., Rev. Bryol. Lichénol. 35: 106, 1908 (Lacouture 1908) nom. illeg. (ICN Art. 52.1; earlier name included, [Bryopteris (Nees 1838: 211) Lindenb. in Gottsche et al. 1845a: 284]). Basionym:—Lejeunea subg. Bryolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 111, 1884, ‘Bryo-Lejeunea’ (Spruce 1884), nom. illeg. ICN Art. 52.1; earlier name included, Frullania subg. Bryopteris (Nees 1838: 211)]. Lectotype (Grolle 1983: 7):––Jungermannia filicina Swartz (1788: 145) ≡ Lejeunea filicina (Sw.) Nees et Mont. in Montagne (1839: 66) ≡ Bryopteris filicina (Sw.) Nees in Gottsche et al. (1845a: 284). Note:—At both generic and subgeneric ranks the name Bryolejeunea is illegitimate as it included names at that rank that should be used. The generic name was used in four publications prior to its valid publication (Stephani 1888c: 285, 1890a: 2, 1890c: 134, 1892e: 179). The type of the genus is now included in Bryopteris (Nees) Lindenb. Caudalejeunea Schiffn., Hepat. (Engl.-Prantl): 129, 1893 (Schiffner 1893b). Lectotype (Vanden Berghen 1945 [1948]: 38):—Lejeunea lehmanniana Gottsche in Gottsche et al. (1845b: 325) ≡ Caudalejeunea lehmanniana (Gottsche) Evans (1907b: 554). Note:— This is not a Sprucean subgenus, but the name was introduced, but nowhere validly published, in the period 1884–1893 at both subgeneric (Stephani 1890a: 18; ICN Art. 35.2; not explicitly connected with a generic name) and generic level (Stephani 1890c: 134, Bescherelle 1892: 13). Ceratolejeunea (Spruce) J.B.Jack et Steph., Hedwigia 31: 16. 1892 (Jack & Stephani 1892). Basionym:—Lejeunea subg. Ceratolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 198, 1884, ‘Cerato-Lejeunea’ (Spruce 1884). Lectotype (Grolle 1983: 10):—Lejeunea cubensis Montagne (1842: 481) ≡ Ceratolejeunea cubensis (Mont.) Schiffner (1893b: 125). Note:—Bonner et al. (1961: 354) regarded the brief reference to Cerato-Lejeunea in Stephani (1890a: 16) as valid publication of the genus name. It can also be argued that the brief comparison with Neurolejeunea in Stephani (1888c: 283) would be enough of a description of the genus. However, because of controversy regarding the rank at which these names are treated in this publication, both these publications will be proposed for suppression at the rank of genus. As Grolle (1983) also concluded, Ceratolejeunea Jack & Stephani (1892: 16) is the earliest validly published name by the description of Ceratolejeunea grandiloba being treated as a combined generic-specific description. The generic name had been used in ten publications (Stephani 1888c: 283, 1890a: 16, 18, 1890b: 91–95, 1890c: 134, 1890d: 284–286, 1891a: 202, 207; 1891c: 203, Schiffner 1890: 30, Coulter et al. 1890: 348, Prantl 1891: vi) prior to its valid publication. Fulford (1945: 369) lectotypified the genus with Lejeunea cornuta, but this was rejected by Grolle (1983) as it was not definitely included in the subgenus by Spruce (1884). In his monograph of neotropical Ceratolejeunea, Dauphin (2003) followed Grolle (1983) in accepting Ceratolejeunea J.B.Jack et Steph. as a validly published generic name that, prior to the introduction of what is now ICN Art. 41.4, would be a heterotypic synonym with Ceratolejeunea (Spruce) Schiffn., and consequently designated Ceratolejeunea grandiloba J.B.Jack et Steph. as the generitype. Dauphin (2003) recognized two subgenera, Ceratolejeunea subg. Ceratolejeunea and Ceratolejeunea subg. Caduciloba Schuster (1978: 430; nine species). Following ICN Art. 41.1, Jack & Stephani’s name must now be regarded as a combination of Spruce’s subgenus and thus the choice of Ceratolejeunea grandiloba as type must be rejected. The name Ceratolejeunea subg. Ceratolejeunea must then replace the name Ceratolejeunea subg. Caduciloba and the name Ceratolejeunea subg. Ceratophora Schuster (1956: 294) must replace the name Ceratolejeunea subg. Ceratolejeunea sensu Dauphin. GENERA OF LEJEUNEACEAE ESTABLISHED IN 1884–1893 Phytotaxa 220 (2) © 2015 Magnolia Press • 161 Cheilolejeunea (Spruce) Steph., Bot. Gaz. 15: 284, 1890 (Stephani 1890d), nom. conserv. Basionym:—Lejeunea subg. Cheilolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 251, 1884, ‘Cheilo-Lejeunea’ (Spruce 1884). Lectotype (Grolle 1983: 10):—Lejeunea decidua Spruce (1884: 254) ≡ Cheilolejeunea decidua (Spruce) Evans (1905a: 188) [= Cheilolejeunea adnata (Kunze ex Lehmann 1834: 46) Grolle (1977: 529), cf. Gradstein (1989)]. Note:—Bonner et al. (1961) considered Cheilolejeunea validly published as a genus by Stephani (1890b: 88). However, on page 88 of this publication Stephani referred to “Cheilo-Lej. sandvicensis St. n. sp.”, as “Section Cheilo-Lejeunea” and not genus. As shown by Grolle (1979b: 342), Cheilolejeunea can be considered a validly published generic name with the description of Cheilolejeunea kurzii being treated as a combined generic-specific one (ICN Art. 38.5). Grolle (1979b) also determined that the original material of Lejeunea aneogyna in the Spruce herbarium [MANCH] is a mixture of three taxa. The plant that most closely matches the protologue of Lejeunea aneogyna Spruce (1884: 254), the species chosen as the lectotype of Cheilolejeunea by Evans (1906: 2), actually belongs to Trachylejeunea aneogyna (Spruce) Grolle (1979b: 338). The other species in the mixture are Cheilolejeunea decidua and Cheilolejeunea duriuscula (Nees in Gottsche et al. 1845b: 364) Schiffner (1897: 592). Grolle (1983) proposed Cheilolejeunea decidua as the new lectotype for the genus (ICN Art. 9.19, 9A.4). Grolle (1983) suggested that the generic name Cheilolejeunea (Spruce) Schiffner (1893b: 124) should be conserved against the older Euosmolejeunea Steph. (1889a), but a proposal was never formalized. The proposal of Ye et al. (2011) to conserve Cheilolejeunea (Spruce) Schiffn. against Omphalanthus Lindenb. et Nees in Gottsche et al. (1845: 303) has been accepted and added to Appendix III of the ICN (McNeill et al. 2012). However, Cheilolejeunea has not yet been conserved against the earlier name Euosmolejeunea, which is now included in it (Grolle 1979b). The generic name Cheilolejeunea was used but not validly published in four publications (Stephani 1888b: 109, 1888c: 285, 1890b: 80–86, 1890c: 135) prior to its validation. Cololejeunea (Spruce) Steph., Hedwigia 30: 208, 1891 (Stephani 1891a). Basionym:—Lejeunea subg. Cololejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 251, 1884, ‘Colo-Lejeunea’ (Spruce 1884). Lectotype (Vanden Berghen 1945 [1945]: 50):—Lejeunea calcarea Libert (1820: 373) ≡ Cololejeunea calcarea (Libert) Stephani (1892b: 171). Note:—Stephani (1888c: 285) gave a short diagnosis that may be taken as a description. Bonner et al. (1961) considered Cololejeunea validly published in Stephani (1890a: 18). However, as discussed earlier, whether the taxon is being treated as a genus or a subgenus is ambiguous. As indicated by Grolle (1983), the genus was validly established by the description of Cololejeunea elegans being treated as a combined generic-specific one (ICN Art. 38.5). Spruce recognized two sections in his subgenus, Lejeunea sect. Physocolea Spruce (1884: 292) and Lejeunea sect. Leptocolea Spruce (1884: 294). Evans (1911) applied the name Cololejeunea only to section Physocolea, for which he chose Lejeunea calcarea, the first species described by Spruce, as the type. Section Leptocolea was elevated to generic rank by Evans (1911: 261). Cololejeunea was used but not validly published in six publications (Stephani 1888c: 289, 1889a: 168, 1890a: 18, 1890b: 71–96, 1890c: 135, Schiffner 1890: 33–34) prior to its validation. Colurolejeunea (Spruce) Steph., Hedwigia 30: 208, 1891 (Stephani 1891a). Basionym:—Lejeunea subg. Colurolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 303, 1884, ‘Coluro-Lejeunea’ (Spruce 1884). Lectotype (Bonner 1963: 879):—Jungermannia calyptrifolia Hooker (1813: tab. 43) ≡ Lejeunea calyptrifolia (Hook.) Sprengel (1827: 234) ≡ Colurolejeunea calyptrifolia (Hook.) Stephani (1891a: 208) ≡ Colura calyptrifolia (Hook.) Dumortier (1835: 12). Note:—The genus was established by the description of Colurolejeunea obtusa being treated as a combined generic-specific description. The generic name was used but not validly published in three publications (Schiffner 1890: 34, Stephani 1890b: 97–98, 1890c: 135) prior to its valid publication. Stephani’s name, although technically not illegitimate (ICN Art. 52.3), is an obligate synonym of the earlier Colura (Dumortier 1831) Dumortier (1835), since the two genera share the same type. Crossotolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 127, 1893 (Schiffner 1893b). Basionym:—Lejeunea subg. Crossotolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 161, 1884, ‘Crossoto-Lejeunea’ (Spruce 1884). Lectotype (Vanden Berghen 1945 [1948]: 30):—Lejeunea boryana Montagne (1838: 47) ≡ Crossotolejeunea boryana (Mont.) Schiffner (1893b: 127). Note:—Bonner et al. (1961) considered the genus validly published in Stephani (1890b: 70), but this publication will be proposed for suppression (see earlier discussion). The generic name was used but not validly published in three publications (Stephani 1890b: 70, 1890c: 136, Jack & Stephani 1892: 16) prior to its valid publication. The type of the genus is now included in Lejeunea. Dendrolejeunea (Spruce) Lacout., Rev. Bryol. 35: 104 (Lacouture 1908). Basionym:—Lejeunea subg. Dendrolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 110, 1884, ‘Dendro-Lejeunea’ (Spruce 1884). Type (monotypic subgenus):—Bryopteris fruticosa Lindenb. et Gottsche in Gottsche et al. (1847: 737) ≡ Lejeunea fruticosa (Lindenb. et Gottsche) Spruce (1884: 110) ≡ Dendrolejeunea fruticosa (Lindenb. et Gottsche) Lacouture (1908: 105) ≡ Thysananthus fruticosus (Lindenb. et Gottsche) Schiffner (1893b: 130). Note:—The generic name was used but not validly published by Schiffner (1890: 25) prior to its valid publication. The type of the genus is now included in Thysananthus Lindenb. in Lehmann (1844: 24) Dicranolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 128, 1893 (Schiffner 1893b). Basionym:—Lejeunea subg. Dicranolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 138, 1884, ‘Dicrano-Lejeunea’ (Spruce 1884). Lectotype (Vanden Berghen 1945 [1948]: 45):—Lejeunea axillaris Nees & Montagne (1836: 59) ≡ Dicranolejeunea axillaris (Nees et Mont.) Schiffner (1893b: 128). Note:—Bonner et al. (1961) considered the genus validly published in Stephani (1890a: 7), but this publication will be proposed for suppression at the rank of genus (see earlier discussion). The generic name was used in four publications (Stephani 1890a: 16, 1890c: 136, 1892e: 179, Jack & Stephani 1892: 13) prior to its valid publication. 162 • Phytotaxa 220 (2) © 2015 Magnolia Press SÖDERSTRÖM ET AL. Diplasiolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 121, 1893 (Schiffner 1893b). Basionym:—Lejeunea subg. Diplasiolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 301, 1884, ‘Diplasio-Lejeunea’ (Spruce 1884). Lectotype (Vanden Berghen 1945 [1948]: 48):—Jungermannia pellucida C.F.W.Meissn. ex Sprengel (1827: 325) ≡ Lejeunea pellucida (C.F.W.Meissn. ex Spreng.) Gottsche et al. (1845b: 393) ≡ Diplasiolejeunea pellucida (C.F.W.Meissn. ex Spreng.) Schiffner (1893b: 121). Note:—Bonner et al. (1961) considered the generic name validly published in Stephani (1890b: 81; probably error for p. 91), but this publication will be proposed for suppression at the rank of genus (see earlier discussion). Diplasiolejeunea was used in its generic form in four publications (Stephani 1888c: 288, 1890b: 90–91, 1890c: 136, 1892e: 179) prior to its valid publication. Drepanolejeunea (Spruce) Steph., Hedwigia 30: 209, 1891 (Stephani 1891a). Basionym:—Lejeunea subg. Drepanolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 186, 1884 ‘Drepano-Lejeunea’ (Spruce 1884). Lectotype (Vanden Berghen 1945 [1948]: 31):—Jungermannia hamatifolia Hooker (1813: tab. 51) ≡ Lejeunea hamatifolia (Hook.) Sprengel (1827: 234) ≡ Drepanolejeunea hamatifolia (Hook.) Schiffner (1893b: 126). Note:—The genus was established by the description of Drepanolejeunea cristata being treated as a combined generic-specific description. Bonner et al. (1961) considered the genus validly published by the description of Drepanolejeunea grossidens in Stephani (1889a: 168). However, the “genus” was not monotypic in that publication and therefore ICN Art. 38.5 cannot be applied. The generic name was used in four publications (Stephani 1888c: 286, 1889a: 168, 1890b: 70–90, 1890c: 136) prior to its valid publication. Eulejeunea Steph., Hedwigia 27: 60, 1888 (Stephani 1888a). Holotype:—Eulejeunea hepaticola Stephani (1888a: 60) ≡ Lejeunea hepaticola (Steph.) Stephani (1915: 714). Note:—Neither Bonner et al. (1961) nor Zijlstra (1982) regarded Eulejeunea as validly published. However, we agree with Grolle (1983: 16) and consider the name to be validly established by Stephani (1888a) with the description of Eulejeunea hepaticola being treated as a combined generic-specific description. Since Lejeunea subg. Eu-Lejeunea Spruce (1884: 260) is not a validly published name for Spruce’s subgenus as it included the type of the genus Lejeunea (cf. ICN Art. 21.3), Stephani’s generic name cannot be considered a combination of Spruce’s name (cf. ICN Art. 41.4). The type of Stephani’s genus Eulejeunea is now included in Lejeunea Lib. Euosmolejeunea (Spruce) Steph., Hedwigia 28: 170, 1889 (Stephani 1889a). Basionym:—Lejeunea subg. Euosmolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 241, 1884 ‘Euosmo-Lejeunea’ (Spruce 1884). Lectotype (Vanden Berghen 1945 [1948]: 27):— Jungermannia trifaria Reinwardt et al. (1824: 226) ≡ Lejeunea trifaria (Reinw., Blume et Nees) Gottsche et al. (1845b: 341) ≡ Euosmolejeunea trifaria (Reinw., Blume et Nees) Stephani (1892b: 171) ≡ Cheilolejeunea trifaria (Reinw., Blume et Nees) Mizutani (1964: 132). Note:—Stephani established the genus by the generic-specific description of Euosmolejeunea sayeri. Stephani (1888c: 292) used the generic name prior to its valid publication. The type of the generic name is now included in Cheilolejeunea (Spruce) Steph. Since the name Euosmolejeunea has priority, a proposal must be made to conserve the generic name Cheilolejeunea against the older Euosmolejeunea to necessary name changes that would otherwise be necessary (see also under Cheilolejeunea). Harpalejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 126, 1893 (Schiffner 1893b). Basionym:—Lejeunea subg. Harpalejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh15: 164, 1884 (Spruce 1884) ‘Harpa-Lejeunea’. Lectotype (Grolle 1989: 90):—Lejeunea ancistrodes Spruce (1884: 169) ≡ Harpalejeunea ancistrodes (Spruce) Schiffner (1893b: 126). Note:—Bonner et al. (1961) considered the generic name validly published in Stephani (1890b: 70), but this publication will be proposed for suppression at the rank of genus (see earlier discussion). The generic name was used in six publications (Stephani 1888c: 288, 1890b: 70–85, 1890c: 138, Schiffner 1890: 29, Jack & Stephani 1892: 13–18, Pearson 1892: 7) prior to its valid publication. Evans (1903b) recognized Harpalejeunea ovata (Hook.) Schiffner (1893b: 127), based on Jungermannia serpyllifolia var. β ovata Hooker (1813: tab. 42), as the type of Harpalejeunea. However, the type of Schiffner’s taxon is technically Jungermannia ovata Dickson (1793: 11), which is a completely different taxon. As discussed in detail by Grolle (1989), the typification by Evans is contrary to the protologue of Harpalejeunea, requiring the designation of a new lectotype. Homalolejeunea (Spruce) Steph., Hedwigia 27: 112, 1888 (Stephani 1888b). Basionym:—Lejeunea subg. Homalolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 132, 1884, ‘Homalo-Lejeunea’ (Spruce 1884). Lectotype (Grolle 1983: 20):—Jungermannia mackaii Hooker (1813: tab. 53) ≡ Lejeunea mackaii (Hook.) Sprengel (1827: 233) ≡ Marchesinia mackaii (Hook.) Gray (1821: 689). Note:—Stephani established the genus by the description of Homalolejeunea henriquesii being treated as a combined genericspecific description. Stephani’s name, although technically not illegitimate (ICN Art. 52.3), is an obligate synonym of the earlier Marchesinia Gray (1821: 689), since the two genera share the same type. Hygrolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 124, 1893 (Schiffner 1893b). Basionym:—Lejeunea subg. Hygrolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 230, 1884, ‘Hygro-Lejeunea’ (Spruce 1884). Lectotype (Vanden Berghen 1945 [1948]: 27):—Jungermannia cerina Lehm. et Lindenb. in Lehmann (1833: 16) ≡ Lejeunea cerina (Lehm. et Lindenb.) Lehm. et Lindenb. in Gottsche et al. (1845b: 391) ≡ Hygrolejeunea cerina (Lehm. et Lindenb.) Schiffner (1893b: 125). Note:—Bonner et al. (1961) considered the generic name validly published in Stephani (1890a: 11), but this publication will be proposed for suppression at the rank of genus (see earlier discussion). The name was used in 11 publications (Stephani 1888c: 284, 1889a: 171, 1890a: 23, 1890b: 74–98, 1890c: 138, 1891a: 208, 1892c: 169, 1892e: 179, Schiffner 1890: 35, Prantl 1891: ix, Jack & Stephani 1892: 13) prior to its valid publication. The type of the generic name is now included in Lejeunea Lib. GENERA OF LEJEUNEACEAE ESTABLISHED IN 1884–1893 Phytotaxa 220 (2) © 2015 Magnolia Press • 163 Leptolejeunea (Spruce) Steph., Hedwigia 30: 270, 1891 (Stephani 1891b). Basionym:—Lejeunea subg. Leptolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 193, 1884, ‘Lepto-Lejeunea’ (Spruce 1884). Lectotype (Vanden Berghen 1945 [1948]: 31):— Jungermannia vitrea Nees (1830: 56) ≡ Lejeunea vitrea (Nees) Nees, Lindenb. et Gottsche in Meyen (1843: 475) ≡ Leptolejeunea vitrea (Nees) Schiffner (1893b: 126). Note:—The genus was established by the description of Leptolejeunea quintasii being treated as a combined generic-specific description. The generic name was used in four publications (Stephani 1888c: 285, 1889a: 173, 1890b: 96–97, 1890c: 138) prior to its valid publication. Lopholejeunea (Spruce) Steph., Bot. Gaz. 15: 285, 1890 (Stephani 1890d), nom. conserv. Basionym:—Lejeunea subg. Lopholejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 119, 1884, ‘Lopho-Lejeunea’ (Spruce 1884) ≡ Lopholejeunea (Spruce) Schiffn., Hepat. [Engl.-Prantl]: 129, 1893 (Schiffner 1893b). Lectotype (Vanden Berghen 1945 [1948]: 39):—Phragmicoma sagrana Montagne (1842: 464) ≡ Lejeunea sagrana (Mont.) Gottsche et al. (1845b: 314) ≡ Lopholejeunea sagrana (Mont.) Stephani (1891a: 202) [= Lopholejeunea subfusca (Nees) Schiffner (1897: 593), cf. Gradstein (1989)]. Note:—Stephani established the genus by the description of Lopholejeunea multilacera being treated as a combined generic-specific description. Stephani (1889a: 175) published the new species Lopholejeunea norfolkensis, but since he referred to the presumed generic name as subgenus, ICN Art. 38.5 cannot be applied and neither the generic nor the specific names are validly published (ICN Art. 35.1). The generic name was used in four publications (Stephani 1889a: 175, 1890a: 16, 1890c: 285, Schiffner 1890: 28) prior to its valid publication. Macrolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 125, 1893 (Schiffner 1893b). Basionym:—Lejeunea subg. Macrolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 224, 1884, ‘Macro-Lejeunea’ (Spruce 1884). Lectotype (Vanden Berghen 1945 [1948]: 29):—Lejeunea pallescens Mitten (1851: 360) ≡ Macrolejeunea pallescens (Mitt.) Schiffner (1893b: 125). Note:—The generic name was used in three publications (Stephani 1888c: 292, 1890b: 90, 1890c: 139) prior to its valid publication. The type of the generic name is now included in Lejeunea Lib. Mastigolejeunea (Spruce) Steph., Hedwigia 30: 206, 1891 (Stephani 1891a). Basionym:—Lejeunea subg. Mastigolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 100, 1884, ‘Mastigo-Lejeunea’ (Spruce 1884). Lectotype (Vanden Berghen 1945 [1948]: 41):—Jungermannia auriculata Wilson et Hook. in Wilson (1841: no. 170) ≡ Lejeunea auriculata (Wilson et Hook.) Sullivant (1856: 699) ≡ Mastigolejeunea auriculata (Wilson et Hook.) Stephani (1892b: 171). Note:—Stephani (1889b) established the genus by the generic-specific description of Mastigolejeunea nigra. Bonner et al. (1961) considered the generic name established by a generic-specific description of Mastigolejeunea phaea in Stephani (1889b: 257), but as Stephani also included other species under Mastigolejeunea that had been validly published earlier in other genera, ICN Art. 38.5 cannot be applied. Grolle (1983: 25) questioned whether the reference to Spruce’s subgenus in Stephani (1889b: 259) might be interpreted as a cryptic citation of a previously published description. However, Stephani refers to the taxon at both generic and subgeneric ranks. Since it not unequivocal that Mastigolejeunea is being treated as a genus in this publication, ICN Art. 38.14 does not apply. The generic name was used in five publications (Stephani 1888b: 110, 1889b: 257, 1890a: 8, 1890c: 139, Schiffner 1890: 22) prior to its valid publication. Microlejeunea (Spruce) Steph., Hedwigia 27: 61, 1888 (Stephani 1888a). Basionym:—Lejeunea subg. Microlejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 286, 1884, ‘Micro-Lejeunea’ (Spruce 1884). Lectotype (Vanden Berghen 1945 [1948]: 22):— Jungermannia ulicina Taylor (1844: 115) ≡ Lejeunea ulicina (Taylor) Gottsche et al. (1845b: 387) ≡ Microlejeunea ulicina (Taylor) Stephani (1892b: 171). Note:—Stephani established the genus by a description of Microlejeunea africana being treated as a combined generic-specific description. Neurolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 131, 1893 (Schiffner 1893b). Basionym:—Lejeunea subg. Neurolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 84, 1884, ‘Neuro-Lejeunea’ (Spruce 1884). Lectotype (Vanden Berghen 1945 [1948]: 47):— Lejeunea seminervis Spruce (1884: 84) ≡ Neurolejeunea seminervis (Spruce) Schiffner (1893b: 131). Note:—The generic name was used in two publications (Stephani 1888c: 283, 1892b: 171) prior to its valid publication. Odontolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 127, 1893 (Schiffner 1893b). Basionym:—Lejeunea subg. Odontolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 142, 1884, ‘Odonto-Lejeunea’ (Spruce 1884). Lectotype (Vanden Berghen 1945 [1848]: 36):—Jungermannia lunulata Weber (1815: 33) ≡ Lejeunea lunulata (F.Weber) Nees in Gottsche et al. (1845b: 326) ≡ Odontolejeunea lunulata (F.Weber) Schiffner (1893b: 128). Note:—The generic name was used in five publications (Stephani 1888c: 283, 1890a: 19, 1890b: 69,1890c: 139, 1892e, 180) prior to its valid publication. Omphalolejeunea (Spruce) Lacout., Rev. Bryol. 35: 105, 1908 (Lacouture 1908). Basionym:—Lejeunea subg. Omphalolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 86, 1884, ‘Omphalo-Lejeunea’ (Spruce 1884). Type (monotypic subgenus):—Jungermannia filiformis Swartz (1788: 144) ≡ Lejeunea filiformis (Sw.) Nees ex Montagne (1839: 64) ≡ Omphalolejeunea filiformis (Sw.) Lacouture (1908: 105) nom. superfl. Note:—The generic name was used in four publications (Stephani 1888c: 282, 1890c: 140, 1892e: 180, Pearson 1892: 4) prior to its valid publication. Lacouture’s name, although technically not illegitimate (ICN Art. 52.3), is an obligate synonym of the earlier Omphalanthus Lindenb. et Nees in Gottsche et al. (1845a: 303), since the two genera share the same type. Otigoniolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 125, 1893 (Schiffner 1893b). Basionym:—Lejeunea subg. Otigoniolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 226, 1884, ‘Otigonio-Lejeunea’ (Spruce 1884). Lectotype (Vanden Berghen 1945 [1948]: 30):—Lejeunea xiphotis Spruce (1884: 277) ≡ Otigoniolejeunea xiphotis (Spruce) Schiffner (1893b: 125) [= Otigoniolejeunea 164 • Phytotaxa 220 (2) © 2015 Magnolia Press SÖDERSTRÖM ET AL. huctumalcensis (Lindenb. et Gottsche in Gottsche et al. 1847: 762) Wei et al. (2014: 237), cf. Reiner-Drehwald & Ilkiu-Borges (2007)]. Peltolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 131, 1893 (Schiffner 1893b). Basionym:—Lejeunea subg. Peltolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 85, 1884, ‘Pelto-Lejeunea’ (Spruce 1884). Lectotype (Vanden Berghen 1945 [1948]: 28):— Lejeunea ovalis Lindenb. et Gottsche in Gottsche et al. (1847: 254) ≡ Peltolejeunea ovalis (Lindenb. et Gottsche) Schiffner (1893b: 131) ≡ Omphalanthus ovalis (Lindenb. et Gottsche) Gradst. in Gradstein et al. (1977: 411). Note:—Bonner et al. (1961) considered the generic name validly published in Stephani (1890a: 23), but this publication will be proposed for suppression at the rank of genus (see earlier discussion). The generic name was used in three publications (Stephani 1890a: 23, 1890c: 140, Jack & Stephani 1892: 13–19) prior to its valid publication. The type of the generic name is now included in Omphalanthus Lindenb. et Nees. Platylejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 130, 1893 (Schiffner 1893b). Basionym:—Lejeunea subg. Platylejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 124, 1884, ‘Platy-Lejeunea’ (Spruce 1884). Lectotype (Grolle 1983: 32):––Lejeunea taeniopsis Spruce (1884: 126) ≡ Symbiezidium taeniopsis (Spruce) Fulford (1942: 302) [= Symbiezidium transversale (Swartz 1788: 144) Trevisan (1877: 403) var. hookerianum (Gottsche et al. 1845b: 311) Gradstein & van Beek (1985), cf. Gradstein (1994)]. Note:—Bonner et al. (1961) considered the generic name validly published in Stephani (1890a: 8), but this publication will be proposed for suppression at the rank of genus (see earlier discussion). The generic name was used in three publications (Stephani 1888c: 282, 1890a: 13, 1890c: 140) prior to its valid publication. The type of the generic name is now included in Symbiezidium Trevisan (1877: 402). Potamolejeunea (Spruce) Lacout., Rev. Bryol. 35: 110, 1908 (Lacouture 1908). Basionym:—Lejeunea subg. Potamolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 249, 1884, ‘Potamo-Lejeunea’ (Spruce 1884). Lectotype (Vanden Berghen 1945 [1948]: 26):—Lejeunea polystachya Spruce (1884: 250) ≡ Potamolejeunea polystachya (Spruce) Stephani (1914: 638) ≡ Cheilolejeunea polystachya (Spruce) Gradstein & Ilkiu-Borges (2009: 62). Note:—The type of the generic name is now included in Cheilolejeunea (Spruce) Steph. Prionolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 127, 1893 (Schiffner 1893b). Basionym:—Lejeunea subg. Prionolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 152, 1884, ‘Priono-Lejeunea’ (Spruce 1884). Lectotype (Vanden Berghen 1945 [1948]: 35):—Lejeunea microdonta Gottsche in Gottsche et al. (1845b: 337) ≡ Prionolejeunea microdonta (Gottsche) Stephani (1897a: 842) [= Prionolejeunea denticulata (Weber 1815: 30) Schiffner (1893b: 127), cf. Ilkiu-Borges (2006)]. Note:—Bonner et al. (1961) considered the generic name validly published in Stephani (1890a: 8), but this publication will be proposed for suppression at the rank of genus (see earlier discussion). The generic name was used in six publications (Stephani 1888c: 285, 1890b: 69–70, 1890c: 140, 1892c: 171, Jack & Stephani 1892: 13, Pearson 1892: 7) prior to its valid publication. Ptycholejeunea (Spruce) Steph., in Engler, Pflanzenwelt Ostafrikas C: 65, 1895 (Stephani 1895b). Basionym:—Lejeunea subg. Ptycholejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 97, 1884, ‘Ptycho-Lejeunea’ (Spruce 1884). Lectotype (Grolle 1983: 33):—Jungermannia striata Lehm. et Lindenb. in Lehmann (1832: 16) ≡ Lejeunea striata (Lehm. et Lindenb.) Stephani (1890c: 140) ≡ Ptycholejeunea striata (Lehm. et Lindenb.) Stephani (1891a: 203) ≡ Ptychanthus striatus (Lehm. et Lindenb.) Nees (1838: 212). Note:—Ptycholejeunea was not established as a new genus by Stephani (1889b: 258) based on the generic-specific description of Ptycholejeunea stephensoniana, as proposed by Bonner et al. (1961) because several additional, validly published species names were referred to Ptycholejeunea in that publication. In this same publication, although he refers to Ptycholejeunea as “Gattung” on p. 258, he refers to it as subgenus on p. 259. As discussed under Mastigolejeunea, because it is not clear that the taxon is being treated as a genus, ICN Art. 38.14 does not apply. Schiffner (1893b) did not treat this taxon, so the earliest publication in which the genus name was validly published is Stephani (1895b: 65). Ptycholejeunea is an obligate synonym of the earlier Ptychanthus Nees (1838: 211), since the two genera share the same type. Pycnolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 124, 1893 (Schiffner 1893b). Basionym:—Lejeunea subg. Pycnolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 246, 1884, ‘Pycno-Lejeunea’ (Spruce 1884). Lectotype (Grolle 1983: 33):—Jungermannia contigua Nees (1833: 360) ≡ Lejeunea contigua (Nees) Montagne (1840: 335) ≡ Pycnolejeunea contigua (Nees) Grolle (1979a: 179). Note:—Bonner et al. (1961) considered the generic name validly published by Stephani (1890a: 10), but this publication will be proposed for suppression at the rank of genus (see earlier discussion). The generic name was used in eight publications (Stephani 1888c: 289, 1889b: 259, 1890a: 10–11, 1890b: 76–86, 1890c: 140, Schiffner 1890: 32, Bescherelle 1892: 14, Pearson 1892: 8) prior to its valid publication. Stictolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 131, 1893 (Schiffner 1893b). Basionym:—Lejeunea subg. Stictolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 81, 1884, ‘Sticto-Lejeunea’ (Spruce 1884). Lectotype (Vanden Berghen 1945 [1948]: 47):— Jungermannia squamata Willd. in Weber (1815: 33) ≡ Lejeunea squamata (Willd.) Nees ex Montagne (1835: 210) ≡ Stictolejeunea squamata (Willd.) Schiffner (1893b: 131). Note:—Bonner et al. (1961) considered the generic name validly published by Stephani (1890a: 17), but this publication will be proposed for suppression at the rank of genus (see earlier discussion). The generic name was used in three publications (Stephani 1888c: 291, 1890a: 17, 1890c: 141) prior to its valid publication. Strepsilejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 127, 1893 (Schiffner 1893b). Basionym:—Lejeunea sect. Strepsilejeunea Spruce, GENERA OF LEJEUNEACEAE ESTABLISHED IN 1884–1893 Phytotaxa 220 (2) © 2015 Magnolia Press • 165 Trans. & Proc. Bot. Soc. Edinburgh 15: 168, 1884, ‘Strepsi-Lejeunea’ (Spruce 1884). Lectotype (Grolle 1983: 39):—Lejeunea inflexa Hampe in Lehmann (1832: 22) ≡ Strepsilejeunea inflexa (Hampe) Stephani (1897b: 24) ≡ Cheilolejeunea inflexa (Hampe) Stephani (1897b: 24). Note:—Bonner et al. (1961) considered the generic name validly published by Stephani (1890b: 74), but this publication will be proposed for suppression at the rank of genus (see earlier discussion). The generic name was used in six publications (Stephani 1888c: 287, 1889b: 260, 1890b: 74–85, 1890c: 141, Jack & Stephani 1892: 13–19, Pearson 1892: 7) prior to its valid publication. The type of the generic name is now included in Cheilolejeunea (Spruce) Steph. Taxilejeunea (Spruce) Steph. Hedwigia 28: 262, 1889 (Stephani 1889b), nom. conserv. Basionym:—Lejeunea subg. Taxilejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 212, 1884, ‘Taxi-Lejeunea’ (Spruce 1884) ≡ Taxilejeunea (Spruce) Schiffn., Hepat. [Engl.Prantl]: 125, 1893 (Schiffner 1893b). Lectotype (Jones 1967: 292):—Jungermannia pterigonia Lehm. et Lindenb. in Lehmann (1834: 44) ≡ Lejeunea pterigonia (Lehm. et Lindenb.) Montagne (1840: 337) ≡ Taxilejeunea pterigonia (Lehm. et Lindenb.) Jack & Stephani (1892: 13). Note:—Stephani (1889b) established the genus by the description of Taxilejeunea convexa being treated as a combined generic-specific description. The generic name was used in Stephani (1888c: 281) prior to its valid publication. The type of the generic name is now included in Lejeunea Lib. Thysanolejeunea (Spruce) Steph., Hedwigia 31: 20, 1892 (Jack & Stephani 1892). Basionym:—Lejeunea subg. Thysanolejeunea Spruce, Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 105, 1884, ‘Thysano-Lejeunea’ (Spruce 1884). Lectotype (Grolle 1983: 41):— Thysananthus comosus Lindenb. in Lehmann (1844: 25) ≡ Lejeunea comosa (Lindenb.) Mitten (1860b: 109). Note:—Stephani established the genus by the description of Thysanolejeunea gottschei being treated as a combined generic-specific description. The generic name was used in three publications (Stephani 1889b: 262, 1890a: 4, 1890c: 142) prior to its valid publication. Stephani’s generic name, although technically not illegitimate (ICN Art. 52.3), is an obligate synonym of the earlier Thysananthus Lindenb. in Lehmann (1844: 24) since the two genera share the same type. Trachylejeunea (Spruce) Steph., Hedwigia 28: 262, 1889 (Stephani 1889b), nom. conserv. Basionym:—Lejeunea subg. Trachylejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 180, 1884, ‘Trachy-Lejeunea’ (Spruce 1884). Lectotype (Vanden Berghen 1945 [1948]: 29):—Lejeunea acanthina Spruce (1884: 182) ≡ Trachylejeunea acanthina (Spruce) Schiffner (1893b: 126) ≡ Cheilolejeunea acanthina (Spruce) Gradstein & Ilkiu-Borges (2009: 62). Note:—Stephani established the genus by the description of Trachylejeunea elegantissima being treated as a combined generic-specific description. The type of the generic name is now included in Cheilolejeunea (Spruce) Steph. Binomials at specific rank in Lejeuneaceae published between 1884 and 1893 and their current classification References in bold are the accepted reference where a binomial at specific rank is validly published, and the references in normal font and put into square brackets are those in which the binomial is used, but considered not validly published. The ICN Article is noted for names not validly published for reasons other than that the genus was not validly published (ICN Art. 35.1). Some names may also fail to fulfil other articles in addition to ICN Art. 35.1. We also give the accepted name of the taxon used in the world checklist of hornworts and liverworts (Söderström et al., in press) if other than listed here. Acrolejeunea abnormis (Gottsche 1882: 352) Pearson (1892: 4). Acrolejeunea amplectens (Stephani 1886: 86) [Stephani (1890c: 133)]. Note:—The name seems never to have been validly published. Current name (Gradstein 1975):—Schiffneriolejeunea polycarpa (Nees 1833: 350) Gradstein (1974: 335). Acrolejeunea aulacophora (Montagne 1843: 259) [Stephani (1890c: 133)] Stephani (1895c: 317). Acrolejeunea borgenii [Stephani (1890a: 10); Stephani (1890c: 133)] Stephani (1891c: 201). Note:—The name is validly published by a reference to Stephani (1890a: 10). Current name (Gradstein 1975):—Schiffneriolejeunea pappeana (Nees in Gottsche et al. 1845a: 296) Gradstein (1974: 335). Acrolejeunea confertissima Stephani (1892c: 165). Current name:—Acrolejeunea emergens (Mitten 1879: 397) Stephani (1895b: 65) var. confertissima (Stephani 1892c: 165) Gradstein (1975: 76). Acrolejeunea cucullata [Stephani (1890c: 133)]. Note:—The name seems never to have been validly published. Current name (Gradstein 1975):—Acrolejeunea pycnoclada (Taylor 1846: 385) Schiffner (1893b: 128) subsp. pycnoclada. Acrolejeunea cumingiana (Montagne 1845a: 7) [Stephani (1890a: 20), ICN Art. 41.1; basionym not valid; Stephani (1890c: 133)] Schiffner (1898: 283) Current name:—Schiffneriolejeunea cumingiana (Montagne 1845a: 7) Gradstein (1974: 333). Acrolejeunea densifolia [Schiffner (1890: 26); Sydow (1894: 346) ICN Art. 36.1(c); publ. in syn.]. Note:—The name seems never to have been validly published. Current name (Sydow 1894):—Schiffneriolejeunea pulopenangensis (Gottsche in Gottsche et al. 1845a) Gradstein (1974: 335). Acrolejeunea fertilis (Reinwardt et al. 1824: 211) [Stephani (1890a: 10), ICN Art. 41.1; basionym not valid; Stephani (1890c: 133)] Schiffner (1893b: 128). 166 • Phytotaxa 220 (2) © 2015 Magnolia Press SÖDERSTRÖM ET AL. Acrolejeunea fulva (Gottsche 1882: 550) Pearson (1892: 4). Current name (Geissler & Gradstein 1994):—Schiffneriolejeunea pappeana (Nees in Gottsche et al. 1845a: 296) Gradstein (1974: 335). Acrolejeunea hartmannii [Stephani (1889a: 164)]. Note:—The name seems never to have been validly published. Current name:— Acrolejeunea securifolia (Nees in Endlicher 1833: 5) Stephani (1895d: 59) subsp. hartmannii (Stephani 1912b: 44) Gradstein (1975). Acrolejeunea haskarliana (Gottsche in Gottsche et al. 1845a: 299) [Stephani (1890c: 133) ‘Hasskarliana’] Schiffner (1893b: 128) ‘Hasskarliana’. Current name:—Schiffneriolejeunea tumida var. haskarliana (Gottsche in Gottsche et al. 1845a: 299) Gradstein & Terken (1981: 77). Acrolejeunea inflexa (Gottsche 1882) Pearson (1892: 4). Acrolejeunea juliformis (Nees 1833: 351) [Stephani (1890c: 133)] Schiffner (1894: 183). Current name:—Archilejeunea juliformis (Nees 1833: 351) Gradstein (1975: 127). Acrolejeunea molleri (Stephani 1886: 86) [Stephani (1890c: 133)] Paris (1906: 41). Current name (Gradstein 1974):—Schiffneriolejeunea polycarpa (Nees 1833: 350) Gradstein (1974: 335). Acrolejeunea novae-guineae [Stephani (1889a: 165)] Stephani (1907: 295). Current name (Gradstein & Terken 1981):—Schiffneriolejeunea tumida (Nees 1838: 213) Gradstein (1974: 335). Acrolejeunea occulta [Stephani (1888b: 112); Stephani (1889a: 166)]. Note:—The name seems never to have been validly published. Current name:—Schiffneriolejeunea occulta (Stephani 1912b: 25) Gradstein (1974: 333). Acrolejeunea pappeana (Nees in Gottsche et al. 1845a: 296) [Stephani (1889a: 166); Stephani (1890c: 133)] Schiffner (1894: 182). Current name:—Schiffneriolejeunea pappeana (Nees in Gottsche et al. 1845a: 296) Gradstein (1974: 335). Acrolejeunea parviloba Stephani (1890d: 286). Current name:—Schiffneriolejeunea parviloba (Stephani 1890d: 286) Gradstein (1974: 335). Acrolejeunea peradeniensis (Mitten 1860b: 111) [Stephani (1889a: 166)] Schiffner (1898: 286). Current name (Gradstein & Terken 1981):—Schiffneriolejeunea tumida (Nees 1838: 213) Gradstein (1974: 335) var. tumida. Acrolejeunea polycarpa (Nees 1833: 350) [Stephani (1889a: 166); Stephani (1890a: 22); Stephani (1890c: 133)] Schiffner (1893b: 128). Current name:—Schiffneriolejeunea polycarpa (Nees 1833: 350) Gradstein (1974: 335). Acrolejeunea pulopenangensis (Gottsche in Gottsche et al. 1845a: 299) [Stephani (1889a: 166); Stephani (1890c: 133)] Schiffner (1894: 184). Current name:—Schiffneriolejeunea pulopenangensis (Gottsche in Gottsche et al. 1845a) Gradstein (1974: 335). Acrolejeunea pycnoclada (Taylor 1846: 385) Schiffner (1893b: 128). Acrolejeunea renauldii [Stephani (1888b: 107)]. Note:—The name seems never to have been validly published. Current name (Gradstein 1975):—Acrolejeunea aulacophora (Montagne 1843: 259) Stephani (1895c: 317). Acrolejeunea rostrata [Schiffner (1890: 26)] Schiffn. ex Sydow (1894: 246). Current name (Gradstein 1975):—Acrolejeunea pycnoclada (Taylor 1846: 385) Schiffner (1893b: 128) subsp. pycnoclada. Acrolejeunea securifolia (Nees in Endlicher 1833: 5) [Stephani (1890c: 133)] Stephani (1895b: 59). Acrolejeunea terminalis (Spruce 1884: 116) [Stephani (1889a: 166)] Schiffner (1894: 178). Current name (Verdoorn 1934a):—Acrolejeunea pycnoclada (Taylor 1846: 385) Schiffner (1893b: 128) subsp. pycnoclada. Acrolejeunea torulosa (Lehm. et Lindenb. in Lehmann 1834: 41) [Stephani (1890c: 133)] Schiffner (1893b: 128). Acrolejeunea tumida (Nees 1838: 213) [Stephani (1890c: 133)] Schiffner (1894: 185). Current name:—Schiffneriolejeunea tumida (Nees 1838: 213) Gradstein (1974: 335). Acrolejeunea wildii [Stephani (1889a: 165)] Steph. et Watts in Watts (1901: 215). Current name (Gradstein 1975):—Acrolejeunea securifolia (Nees in Endlicher 1833: 5) Stephani (1895d: 59) subsp. securifolia. Anomalolejeunea pluriplicata (Pearson 1887: 2) [Stephani (1892a: 120)] Schiffner (1893b: 127). Current name:—Cheilolejeunea pluriplicata (Pearson 1887: 2) Schuster (1980b: 430). Anoplolejeunea herpestica (Spruce 1884: 129) Schiffner (1893b: 131). Current name (Evans 1908a):—Anoplolejeunea conferta (C.F.W.Meissn. ex Sprengel 1827: 325) Evans (1908a: 175). Archilejeunea auberiana (Montagne 1842: 483) Stephani (1890c: 134). Archilejeunea bongardii Stephani (1890a: 20). Archilejeunea chrysophylla (Lehmann 1835: 423) Stephani (1890c: 134). Current name:—Acanthocoleus chrysophyllus (Lehmann 1835: 423) Kruijt (1988: 72). Archilejeunea clypeata (Schweinitz 1821: 12) Stephani (1890a: 21). Current name:—Cheilolejeunea clypeata (Schweinitz 1821: 423) Ye & Zhu (2010: 280). Archilejeunea cognata (Nees 1833: 353) Stephani (1890c: 134). Current name (Gradstein & Buskes 1985):—Anoplolejeunea conferta (C.F.W.Meissn. ex Sprengel 1827: 325) Evans (1908a: 175). Archilejeunea conferta (C.F.W.Meissn. ex Sprengel 1827: 325) [Stephani (1890b: 73) ICN Art. 41.1; basionym not valid] Stephani (1890c: 134). Current name:—Anoplolejeunea conferta (C.F.W.Meissn. ex Sprengel 1827: 325) Evans (1908a: 175). GENERA OF LEJEUNEACEAE ESTABLISHED IN 1884–1893 Phytotaxa 220 (2) © 2015 Magnolia Press • 167 Archilejeunea cruegeri (Lindenb. in Gottsche et al. 1845b: 319) Stephani (1890c: 134). Current name (Gradstein & Buskes 1985):— Archilejeunea parviflora (Nees 1833: 353) Stephani (1890c: 134). Archilejeunea erronea Stephani (1888b: 113). Current name (Gradstein & Geissler 1997):—Cheilolejeunea clypeata (Schweinitz 1821: 423) Ye & Zhu (2010: 280). Archilejeunea guahamensis (Lindenb. in Gottsche et al. 1845b: 346) Stephani (1890c: 134). Current name (Verdoorn 1934b):— Mastigolejeunea humilis (Gottsche in Gottsche et al. 1845a: 239) Schiffner (1893b: 129). Archilejeunea huanucensis (Gottsche. in Gottsche et al. 1845b: 346) Stephani (1890c: 134). Current name:—Omphalanthus huanucensis (Gottsche in Gottsche et al. 1845b: 346) Gradst. in Gradstein & Buskes (1985: 109). Archilejeunea leprieurii (Montagne 1835: 213) Schiffner (1893b: 130). Current name (Gradstein & Buskes 1985):—Archilejeunea parviflora (Nees 1833: 353) Stephani (1890c: 134). Archilejeunea mariana (Gottsche in Gottsche et al. 1845b: 337) Stephani (1890a: 17). Current name:—Cheilolejeunea mariana (Gottsche in Gottsche et al. 1845b: 337) Thiers & Gradstein (1989: 75). Archilejeunea nobilis (Gottsche 1882: 353) Pearson (1892: 4). Current name:—Marchesinia nobilis (Gottsche 1882: 353) Shi et al. (2015: 249). Archilejeunea olivacea (Hooker & Taylor 1844c: 568) Stephani (1890c: 134). Archilejeunea parviflora (Nees 1833: 353) Stephani (1890c: 134). Archilejeunea rotalis (Hooker & Taylor 1845: 89) Stephani (1890c: 134). Current name:—Cheilolejeunea rotalis (Hooker & Taylor 1845: 89) Wigginton (2012: 270). Archilejeunea rotundistipula (Lehmann 1829: 360) [Stephani (1890a: 20), ICN Art. 41.1; basionym not valid] Stephani (1890c: 134). Current name:—Cheilolejeunea rotundistipula (Lindenb. ex Lehmann 1829: 360) Malombe (2009: 322). Archilejeunea scutellata (Taylor 1846: 383) Stephani (1890c: 134). Current name (Gradstein & Buskes 1985):—Archilejeunea olivacea (Hooker & Taylor 1844c: 568) Stephani (1890c: 134). Archilejeunea unciloba (Lindenb. in Gottsche et al. 1845b: 331) Stephani (1890c: 134). Current name:—Cheilolejeunea unciloba (Lindenb. in Gottsche et al. 1845b: 331) Malombe (2009: 325). Archilejeunea viridissima (Lindenberg 1843: 320) Stephani (1890c: 134). Current name (Gradstein & Buskes 1985):—Archilejeunea parviflora (Nees 1833: 353) Stephani (1890c: 134). Archilejeunea xanthocarpa (Lehm. et Lindenb. in Lehmann 1833: 8) Stephani (1890c: 134). Current name:—Cheilolejeunea xanthocarpa (Lehm. et Lindenb. in Lehmann 1833: 8) Malombe (2009: 326). Brachiolejeunea bicolor (Nees 1833: 349) [Stephani (1890c: 134)] Schiffner (1893b: 128). Current name (van Slageren 1985):— Frullanoides densifolia Raddi (1822: 14) subsp. densifolia. Brachiolejeunea birmensis [Stephani 1889a: 167] Stephani (1895d: 63). Current name (Mizutani 1989):—Acrolejeunea infuscata (Mitten 1860b: 111) Jian Wang bis et Gradst. in Wang et al. (2014: 38). Brachiolejeunea caledonica [Stephani (1889a: 167)]. Note:—The name seems never to have been validly published and presumably refers to some other genus since Brachiolejeunea is a New World genus (van Slageren 1985). Brachiolejeunea corticalis (Lehm. et Lindenb. in Lehmann 1832: 50) [Stephani (1890a: 7); Stephani (1890c: 134)] Schiffner (1894: 180). Current name:—Frullanoides corticalis (Lehm. et Lindenb. in Lehmann 1832: 50) van Slageren (1985: 84). Brachiolejeunea laxifolia (Taylor 1847: 341) [Stephani (1890a: 7); Stephani (1890c: 134)] Schiffner (1893b: 128). Brachiolejeunea leiboldiana (Gottsche et Lindenb. in Gottsche et al. 1845a: 296) [Stephani (1890c: 134)] Schiffner (1894: 182). Brachiolejeunea liebmaniana (Lindenb. et Gottsche in Gottsche et al. 1847: 744) [Stephani (1890c: 134) ‘Liebmanniana’] Schiffner (1894: 183). Current name:—Frullanoides liebmaniana (Lindenb. et Gottsche in Gottsche et al. 1847: 744) van Slageren (1985: 102). Brachiolejeunea papilionacea [Stephani (1889a: 167)] Stephani (1895d: 64). Current name (van Slageren 1985):—Phaeolejeunea latistipula (Schiffn. ex Sydow 1894: 246) Mizutani (1968: 131). Brachiolejeunea plagiochiloides [Steph. et Spruce in Stephani (1889a: 167)] (Stephani 1889b: 257) Stephani (1912b: 141). Current name (van Slageren 1985):—Spruceanthus semirepandus (Nees 1830: 39) Verdoorn (1934b: 153). Brachiolejeunea securifolia [Stephani (1889a: 168)] Stephani (1912b: 128). Current name:—Blepharolejeunea securifolia (Stephani 1912b: 128) Schuster (1980b: 424). Brachiolejeunea spruceana (Massalongo 1885: 246) Schiffner (1893b: 128). Brachiolejeunea succisa [Stephani (1889a: 168)] Stephani (1895d: 65). Current name (van Slageren 1985):—Frullanoides densifolia Raddi (1822: 14) subsp. densifolia. Brachiolejeunea tristis [Stephani (1890a: 8); Stephani (1890c: 134)] Stephani (1912b: 112). Current name:—Frullanoides tristis (Stephani 1912b: 112) van Slageren (1985: 110). Brachiolejeunea ustulata (Taylor 1846: 388) [Stephani (1890c: 134)]. Note:—The name seems never to have been validly published. Current name (Gradstein 1975):—Acrolejeunea fertilis (Reinwardt et al. 1824: 211) Schiffner (1893b: 128). 168 • Phytotaxa 220 (2) © 2015 Magnolia Press SÖDERSTRÖM ET AL. Bryolejeunea diffusa (Swartz 1788: 144) [Stephani (1888c: 300); Stephani (1890c: 134); Stephani (1892e: 179)]. Note:—The name seems never to have been validly published. Current name:—Bryopteris diffusa (Swartz 1788: 144) Nees in Gottsche et al. (1845a: 286). Bryolejeunea filicina (Swartz 1788: 145) [Stephani (1888c: 285); Stephani (1890c: 134)]. Note:—The name seems never to have been validly published. Current name:—Bryopteris filicina (Swartz 1788: 145) Nees in Gottsche et al. (1845a: 284). Bryolejeunea flaccida (Lindenb. et Hampe in Hampe 1851: 640) [Stephani (1890c: 134); Stephani (1892e: 179)]. Note:—The name seems never to have been validly published. Current name (Gradstein 1994):—Bryopteris filicina (Swartz 1788: 145) Nees in Gottsche et al. (1845a: 284). Bryolejeunea fruticulosa (Taylor 1846: 382) [Stephani (1892e: 179)] Lacouture (1908: 107). Current name (Gradstein 1994):—Bryopteris filicina (Swartz 1788: 145) Nees in Gottsche et al. (1845a: 284). Bryolejeunea gaudichaudii (Gottsche 1857: 340) [Stephani (1890c: 134)]. Note:—The name seems never to have been validly published. Current name:—Bryopteris gaudichaudii Gottsche (1857: 340). Bryolejeunea liebmaniana (Lindenb. et Gottsche in Gottsche et al. 1847: 738) [Stephani (1890c: 134) ‘Liebmanniana’]. Note:—The name seems never to have been validly published. Current name (Gradstein 1994):—Bryopteris filicina (Swartz 1788: 145) Nees in Gottsche et al. (1845a: 284). Bryolejeunea tenuicaulis (Gottsche et al. 1845a: 285) [Stephani (1890c: 134); Stephani (1892e: 179)]. Note:—The name seems never to have been validly published. Current name (Gradstein 1994):—Bryopteris filicina (Swartz 1788: 145) Nees in Gottsche et al. (1845a: 284). Bryolejeunea trinitensis (Lehm. et Lindenb. in Lehmann 1833: 12) [Stephani (1890c: 134)]. Note:—The name seems never to have been validly published. Current name (Gradstein 1994):—Bryopteris filicina (Swartz 1788: 145) Nees in Gottsche et al. (1845a: 284). Caudalejeunea africana (Stephani 1886: 93) Schiffner (1893b: 129). Caudalejeunea crescentiae (Lindenb. et Gottsche in Gottsche et al. 1847: 752) [Stephani (1890c: 134)] Schiffner (1893b: 129). Current name (Stephani 1912b):—Caudalejeunea lehmanniana (Gottsche in Gottsche et al. 1845b: 325) Evans (1907b: 554). Caudalejeunea haenkeana (Schiffner 1886: 210) Schiffner (1893b: 129). Current name (Stephani 1912b):—Caudalejeunea lehmanniana (Gottsche in Gottsche et al. 1845b: 325) Evans (1907b: 554). Caudalejeunea hanningtonii (Mitten 1886: 324) Schiffner (1893b: 129). Caudalejeunea harpaphylla (Spruce 1884: 123) Schiffner (1893b: 129). Current name (Stephani 1912b).:—Caudalejeunea lehmanniana (Gottsche in Gottsche et al. 1845b: 325) Evans (1907b: 554). Caudalejeunea lehmanniana (Gottsche in Gottsche et al. 1845b: 325) [Stephani (1890c: 134)] Evans (1907b: 554). Caudalejeunea leiboldii [Stephani (1890a: 19)] Stephani (1895a: 233). Current name (Stephani 1912b):—Caudalejeunea lehmanniana (Gottsche in Gottsche et al. 1845b: 325) Evans (1907b: 554). Caudalejeunea recurvistipula (Gottsche in Gottsche et al. 1845b: 326) [Stephani (1890c: 134)] Schiffner (1893b: 129). Current name (Verdoorn 1934b):—Caudalejeunea reniloba (Gottsche in Gottsche et al. 1845b: 326) Stephani (1912b: 16). Caudalejeunea stephanii [Bescherelle 1892: 13; Schiffner (1893b: 129), ICN Art. 38.1(a); no description] Stephani (1912b: 17). Current name (Mizutani 1988):—Caudalejeunea reniloba (Gottsche in Gottsche et al. 1845b: 326) Stephani (1912b: 16). Ceratolejeunea amazonica [Stephani (1890b: 91); Stephani (1890c: 134)] Stephani (1913: 395). Current name (Dauphin 2003):— Ceratolejeunea cubensis (Montagne 1842: 481) Schiffner (1893b: 125). Ceratolejeunea auriculata [Schiffner (1890: 30)] Schiffn. ex Sydow (1894: 246). Current name (Mizutani 1977):—Lepidolejeunea bidentula (Steph. in Jack & Stephani 1894: 107) Schuster (1980b: 425). Ceratolejeunea belangeriana (Gottsche in Gottsche et al. 1845b: 398) [Stephani (1890b: 93); Stephani (1890c: 134)] Stephani (1913: 396). Ceratolejeunea brasiliensis (Gottsche in Gottsche et al. 1845b: 398) [Stephani (1890b: 93); Stephani (1890c: 134)] Stephani (1892e: 179). Current name (Dauphin 2003):—Ceratolejeunea fallax (Lehm. et Lindenb. in Lehmann 1833: 17) Bonner (1953: 189). Ceratolejeunea breutelii (Gottsche in Gottsche et al. 1845b: 324) [Stephani (1888c: 283, 300); Stephani (1890c: 134)] Schiffner (1893b: 125). Current name:—Neurolejeunea breutelii (Gottsche in Gottsche et al. 1845b: 324) Evans (1907a: 13). Ceratolejeunea catenulata (Nees in Gottsche et al. 1845b: 323) [Stephani (1890c: 134)]. Note:—The name seems never to have been validly published. Current name:—Neurolejeunea catenulata (Nees in Gottsche et al. 1845b: 323) Schiffner (1893b: 131). Ceratolejeunea ceratantha (Nees et Mont. in Montagne 1840: 335) [Stephani (1888c: 283, 300); Stephani (1890b: 92); Stephani (1890c: 134)] Schiffner (1897: 582). Ceratolejeunea coarina (Gottsche in Gottsche et al. 1845b: 395) [Stephani (1890b: 91); Stephani (1890c: 134)] Schiffner (1893b: 125). Ceratolejeunea cornuta (Lindenberg 1829: 23) [Stephani (1888c: 285); Stephani (1890b: 94); Stephani (1890c: 134)] Stephani (1895b: 65). Ceratolejeunea cornutissima Stephani (1892c: 166). Current name (Wigginton & Grolle 1996):—Ceratolejeunea coarina (Gottsche in Gottsche et al. 1845b: 395) Schiffner (1893b: 125). Ceratolejeunea cubensis (Montagne 1842: 481) [Stephani (1888c: 300); Stephani (1890c: 134)] Schiffner (1893b: 125). GENERA OF LEJEUNEACEAE ESTABLISHED IN 1884–1893 Phytotaxa 220 (2) © 2015 Magnolia Press • 169 Ceratolejeunea desciscens (Sande Lacoste 1856: 107) Schiffner (1893b: 126). Ceratolejeunea devexa (Lindenb. et Gottsche in Gottsche et al. 1847: 750) [Stephani (1890c: 134)] Stephani (1913: 408). Current name (Schuster 1980a):—Neurolejeunea breutelii (Gottsche in Gottsche et al. 1845b: 324) Evans (1907a: 13). Ceratolejeunea diversicornua (Prantl 1891: ix) [Stephani (1891a: 207); Prantl (1891: vi)] Schiffn. ex Sydow (1894: 239). Current name (Gradstein 2013a):—Ceratolejeunea coarina (Gottsche in Gottsche et al. 1845b: 395) Schiffner (1893b: 125). Ceratolejeunea filaria (Taylor in Lehmann 1844: 28) [Stephani (1890b: 96); Stephani (1890c: 134)] Stephani (1913: 412). Ceratolejeunea grandiloba Jack & Stephani (1892: 16). Ceratolejeunea guianensis (Nees et Mont. in Montagne 1840: 335) [Stephani (1890b: 92); Stephani (1890c: 134)] Stephani (1913: 416). Ceratolejeunea haeckeriana (Gottsche et Lindenb. in Gottsche et al. 1845b: 401) [Stephani (1890b: 95); Stephani (1890c: 135)] Stephani (1913: 417). Current name (Reiner-Drehwald 1999):—Ceratolejeunea cornuta (Lindenberg 1829: 23) Stephani (1895b: 65). Ceratolejeunea involvens (Nees et Mont. in Montagne 1840: 336) [Stephani (1890b: 95); Stephani (1890c: 135)] Stephani (1913: 336). Current name (Dauphin 2003):—Ceratolejeunea cornuta (Lindenberg 1829: 23) Stephani (1895b: 65). Ceratolejeunea kegelii (Lindenberg & Gottsche 1851: 635) [Stephani (1888c: 300)] Stephani (1913: 420). Current name (Dauphin 2003):—Ceratolejeunea coarina (Gottsche in Gottsche et al. 1845b: 395) Schiffner (1893b: 125). Ceratolejeunea longicornis (Gottsche in Gottsche et al. 1845b: 401) [Stephani (1890b: 95); Stephani (1890c: 135)] Stephani (1913: 421). Current name (Reiner-Drehwald 1999):—Ceratolejeunea filaria (Lehmann 1844: 28) Stephani (1913: 412). Ceratolejeunea martiana (Gottsche in Gottsche et al. 1845b: 398) [Stephani (1890b: 93); Stephani (1890c: 135)] Stephani (1913: 422). Current name (Reiner-Drehwald 1999):—Ceratolejeunea cornuta (Lindenberg 1829: 23) Stephani (1895b: 65). Ceratolejeunea mascarena [Stephani (1890d: 284), Coulter et al. (1890: 348), Stephani (1891a: 202), Steph. ex Prantl (1891: vi), Stephani (1891c: 203)] Stephani (1892d: 204). Current name (Grolle 1978):—Ceratolejeunea belangeriana (Gottsche in Gottsche et al. 1845b: 398) Stephani (1913: 396). Ceratolejeunea mauritiana [Stephani (1890d: 285), Coulter et al. (1890: 348), Stephani (1891c: 204)] Stephani (1892d: 205). Current name (Grolle 1995):—Ceratolejeunea belangeriana (Gottsche in Gottsche et al. 1845b: 398) Stephani (1913: 396). Ceratolejeunea miradorensis [Stephani (1890b: 93); Stephani (1890c: 135)] (Prantl 1890: xvi) Stephani (1913: 426). Current name (Dauphin 2003):—Ceratolejeunea cornuta (Lindenberg 1829: 23) Stephani (1895b: 65). Ceratolejeunea oculata (Gottsche in Gottsche et al. 1845b: 357) [Stephani (1890c: 135)] Stephani (1897a: 842). Ceratolejeunea poeppigiana (Nees in Gottsche et al. 1845b: 396) [Stephani (1890c: 135)] Stephani (1913: 433). Current name (Dauphin 2003):—Ceratolejeunea ceratantha (Nees et Mont. in Montagne 1840: 335) Schiffner (1897: 582). Ceratolejeunea pyriformis (Stephani 1890b: 93) [Stephani (1890c: 135)] Stephani (1913: 435). Current name (Dauphin 2003):— Ceratolejeunea cubensis (Montagne 1842: 481) Schiffner (1893b: 125). Ceratolejeunea radicosa (Nees ex Montagne 1842: 475) [Stephani (1890b: 95); Stephani (1890c: 135)] Stephani (1913: 435). Current name:—Leptolejeunea radicosa (Nees ex Montagne 1842: 475) Grolle (1979a: 178). Ceratolejeunea renauldii [Stephani (1890d: 286), Coulter et al. (1890: 348), Stephani (1891c: 204)] Stephani (1892d: 205). Current name (Grolle 1977):—Lejeunea alata Gottsche in Gottsche et al. (1845b: 406). Ceratolejeunea rigidula (Nees ex Montagne 1840: 336) [Stephani (1890b: 82); Stephani (1890c: 135)] Stephani (1913: 437). Current name:—Cheilolejeunea rigidula (Nees ex Montagne 1840: 336) Schuster (1971: 102). Ceratolejeunea singapurensis (Lindenb. in Gottsche et al. 1845b: 397) [Stephani (1890b: 93); Stephani (1890c: 135)] Schiffner (1898: 273). Ceratolejeunea spinosa (Gottsche in Gottsche et al. 1845b: 402) [Stephani (1888c: 291); Stephani (1890c: 135)] Stephani (1895a: 238). Ceratolejeunea splitgerberiana (Montagne 1841: 112) [Stephani (1890b: 95)] Stephani (1913: 443). Current name (Reiner-Drehwald 1999):—Ceratolejeunea cornuta (Lindenberg 1829: 23) Stephani (1895b: 65). Ceratolejeunea variabilis (Lindenb. in Gottsche et al. 1845b: 399) [Stephani (1888c: 293); Stephani (1890b: 94); Stephani (1890c: 135)] Pearson (1892: 8). Current name (Dauphin 2003):—Ceratolejeunea cornuta (Lindenberg 1829: 23) Stephani (1895b: 65). Cheilolejeunea bethanica (Gottsche in Gottsche et al. 1845b: 381) [Stephani (1890b: 86); Stephani (1890c: 135)] Stephani 1914: 652. Current name:—Lejeunea bethanica Gottsche in Gottsche et al. (1845b: 381). Cheilolejeunea clausa (Nees et Mont. in Montagne 1840: 337) [Stephani (1890b: 80); Stephani (1890c: 135)] Schuster (1980a: 863). Cheilolejeunea duriuscula (Nees in Gottsche et al. 1845b: 364) [Stephani (1888c: 285, 300); Stephani (1890b: 81); Stephani (1890c: 135)] Schiffner (1897: 592). Current name (Gradstein 1989):—Cheilolejeunea rigidula (Nees ex Montagne 1840: 336) Schuster (1971: 102). Cheilolejeunea gaudichaudii (Lindenb. in Gottsche et al. 1845b: 372) [Stephani (1890b: 82); Stephani (1890c: 135)]. Note:—The name seems never to have been validly published. Current name (Schultze-Motel & Menzel 1987):—Lejeunea flava (Swartz 1788: 144) Nees (1838: 277). Cheilolejeunea heteroclada (Spruce 1884: 256) Schiffner (1893b: 124). 170 • Phytotaxa 220 (2) © 2015 Magnolia Press SÖDERSTRÖM ET AL. Cheilolejeunea intertexta (Lindenb. in Gottsche et al. 1845b: 379) [Stephani (1890b: 85); Stephani (1890c: 135)] Stephani (1897c: 79). Cheilolejeunea kurzii Stephani (1890d: 284). Current name (Kitagawa 1981):—Cheilolejeunea intertexta (Lindenb. in Gottsche et al. 1845b: 379) Stephani (1897c: 79). Cheilolejeunea lineata (Lehm. et Lindenb. in Lehmann 1832: 53) [Stephani (1888c: 287); Stephani (1890b: 82); Stephani (1890c: 135)] Schiffner (1893b: 124). Cheilolejeunea lurida (Lindenb. in Gottsche et al. 1845b: 379) [Stephani (1890b: 85); Stephani (1890c: 135)] Stephani (1914: 658). Cheilolejeunea melastomatis (Lindenb. et Gottsche in Gottsche et al. 1847: 763) [Stephani (1890b: 82); Stephani (1890c: 135)]. Note:— The name seems never to have been validly published. Current name (Reiner-Drehwald & Grolle 2012):—Lejeunea flava (Swartz 1788: 144) Nees (1838: 277). Cheilolejeunea myriantha (Nees et Mont. in Gottsche et al. 1845b: 398) [Stephani (1890b: 80); Stephani (1890c: 135)] Schiffner (1897: 590). Current name (Gradstein & Hekking 1989):—Cheilolejeunea discoidea (Lehm. et Lindenb. in Lehmann 1834: 47) R.M.Schust. et Kachroo in Kachroo & Schuster (1961: 509). Cheilolejeunea newtonii [Stephani (1888b: 109) ‘Newtoni’] Schiffner (1893b: 124) ‘Newtoni’. Current name (Malombe 2007):— Cheilolejeunea intertexta (Lindenb. in Gottsche et al. 1845b: 379) Stephani (1897c: 79). Cheilolejeunea novoguineensis (Schiffner 1893a: 258) Schiffner (1893b: 124). Current name:—Lejeunea novoguineensis Schiffner (1893a: 258). Cheilolejeunea oxyloba (Lindenb. et Gottsche in Gottsche et al. 1847: 770) [Stephani (1890b: 81, 98); Stephani (1890c: 135)] Stephani (1914: 659). Current name (Grolle 1981):—Lejeunea spiniloba Lindenb. et Gottsche in Gottsche et al. (1847: 770). Cheilolejeunea papulosa Schiffner (1893b: 124). Cheilolejeunea principensis [Stephani (1888b: 109)] Schiffner (1893b: 124). Current name (Grolle 1979b):—Cheilolejeunea rigidula (Nees ex Montagne 1840: 336) Schuster (1971: 102). Cheilolejeunea sandvicensis [Stephani (1890b: 88); Stephani (1890c: 135)] (Prantl 1890: xvii) Stephani (1897a: 842). Cheilolejeunea trinitensis (Lindenb. in Gottsche et al. 1845b: 381) [Stephani (1890b: 86) ‘trinidensis’]. Note:—The name seems never to have been validly published. Current name:—Lejeunea trinitensis Lindenb. in Gottsche et al. (1845b: 381). Cheilolejeunea virescens (Lehm. et Lindenb. in Lehmann 1838: 21) [Stephani (1890b: 82); Stephani (1890c: 135)] Stephani (1914: 662). Cheilolejeunea xanthophylla (Lindenb. in Gottsche et al. 1845b: 370) [Stephani (1890b: 81); Stephani (1890c: 135)] Stephani (1914: 663). Cololejeunea angustibracteata [Schiffner (1890: 34)] Schiffn. ex Sydow (1894: 246). Cololejeunea bistyla [Stephani (1889a: 168)]. Note:—The name seems never to have been validly published. Current name:—Cololejeunea hamata Stephani (1895a: 248). Cololejeunea calcarea (Libert 1820: 373) [Stephani (1890b: 71); Stephani (1890c: 135)] Stephani (1892b: 171). Cololejeunea cardiocarpa (Montagne 1842: 476) [Stephani (1890b: 91); Stephani (1890c: 135)] Evans (1902: 172). Cololejeunea crenatiflora Stephani (1891b: 270). Current name (Tixier 1989):—Cololejeunea obliqua (Nees et Mont. in Montagne 1843: 264) Schiffner (1897: 586). Cololejeunea cuneata (Lehm. et Lindenb. in Lehmann 1832: 56) [Stephani (1890b: 91); Stephani (1890c: 135)] Herzog (1947: 320). Cololejeunea cuneifolia Stephani (1892c: 166). Cololejeunea elegans Stephani (1891a: 208). Cololejeunea epiphyta (Gottsche in Gottsche et al. 1845b: 391) [Stephani (1890b: 90); Stephani (1890c: 135)]. Note:—The name seems never to have been validly published. Current name (Evans 1911):—Myriocoleopsis minutissima (Sm. in Smith & Sowerby 1806: tab. 1633) R.L.Zhu, Y.Yu et Pócs in Yu et al. (2014: 293) subsp. myriocarpa (Nees et Mont. in Montagne 1842: 473) R.L.Zhu, Y.Yu et Pócs in Yu et al. (2014: 294). Cololejeunea filicaulis Stephani (1892c: 168). Current name (Tixier 1989):—Cololejeunea hildebrandii (Austin 1876: 35) Stephani (1897a: 842) ‘Hillebrandii’. Cololejeunea floccosa (Lehm. et Lindenb. in Lehmann 1832: 26) [Stephani (1890c: 135)] Schiffner (1898: 243). Cololejeunea haskarliana (Lehm. et Lindenb. in Lehmann 1844: 26) [Stephani (1890b: 72) ‘Hasskarliana’; Stephani (1890c: 135) ‘Hasskarliana’] Schiffner (1898: 244) ‘Hasskarliana’. Cololejeunea jooriana (Austin 1875: 20) Stephani (1892b: 171). Current name (Evans 1923):—Cololejeunea cardiocarpa (Montagne 1842: 476) Evans (1902: 172). Cololejeunea kegelii [Stephani (1890b: 91); Stephani (1890c: 135)] Stephani (1895a: 249). Cololejeunea lhotzskiana (Hampe in Gottsche et al. 1845b: 389) [Stephani (1890b: 89); Stephani (1890c: 135)] Schuster (1963a: 241). Current name (Thiers 1988):—Myriocoleopsis minutissima (Sm. in Smith & Sowerby 1806: tab. 1633) R.L.Zhu, Y.Yu et Pócs in Yu et al. (2014: 293). Cololejeunea marginata (Lehm. et Lindenb. in Lehmann 1833: 11) [Stephani (1888c: 287); Stephani (1890b: 91); Stephani (1890c: 135)] Pearson (1892: 9). Cololejeunea microscopica (Taylor 1836: 59) [Stephani (1890b: 72); Stephani (1890c: 135)] Schiffner (1893b: 122). GENERA OF LEJEUNEACEAE ESTABLISHED IN 1884–1893 Phytotaxa 220 (2) © 2015 Magnolia Press • 171 Cololejeunea minutissima (Sm. in Smith & Sowerby 1806: tab. 1633) [Stephani (1890b: 88); Stephani (1890c: 135)] Stephani (1892b: 171). Current name:—Myriocoleopsis minutissima (Sm. in Smith & Sowerby 1806: tab. 1633) R.L.Zhu, Y.Yu et Pócs in Yu et al. (2014: 293). Cololejeunea montagnei [Stephani (1890b: 96); Stephani (1890c: 135)]. Note:—The name seems never to have been validly published. Current name (Schuster 1980b):—Cololejeunea camillii (Lehmann 1857: 15) Evans (1912b: 59) ‘Camilli’. Cololejeunea myriocarpa (Nees et Mont. in Montagne 1842: 473) [Stephani (1890b: 89); Stephani (1890c: 135)] Massalongo (1906: 9). Current name:—Myriocoleopsis minutissima (Sm. in Smith & Sowerby 1806: tab. 1633) R.L.Zhu, Y.Yu et Pócs in Yu et al. (2014: 293) subsp. myriocarpa (Nees et Mont. in Montagne 1842: 473) R.L.Zhu, Y.Yu et Pócs in Yu et al. (2014: 294). Cololejeunea papilliloba (Stephani 1890b: 73) [Stephani (1890c: 135)] Stephani (1895a: 250). Cololejeunea parvula (Wilson et Hook. in Wilson 1841: no. 172) Stephani (1892b: 172). Current name (Evans 1902):—Myriocoleopsis minutissima (Sm. in Smith & Sowerby 1806: tab. 1633) R.L.Zhu, Y.Yu et Pócs in Yu et al. (2014: 293) subsp. minutissima. Cololejeunea pseudostipulata [Schiffner (1890: 33)] Schiffn. ex Sydow (1894: 246). Cololejeunea punctata (Gottsche 1882: 361) Pearson (1892: 9). Cololejeunea pyriformis (Lindenb. et Gottsche in Gottsche et al. 1847: 767) [Stephani (1890b: 89); Stephani (1890c: 135)]. Note:—The name seems never to have been validly published. Current name:—Microlejeunea pyriformis (Lindenb. et Gottsche in Gottsche et al. 1847: 767) Stephani (1915: 824). Cololejeunea rossettiana (Massalongo 1889: 487) Schiffner (1893b: 122). Cololejeunea sicifolia [Stephani (1888c: 290) ‘sicaefolia’] (Gottsche ex Evans 1911: 277) Pócs & Bernecker (2009: 8) ‘sicaefolia’. Cololejeunea sintenisii [Stephani (1888c: 291)] (Stephani 1916: 861) Pócs in Dauphin et al. (2008: 235). Cololejeunea stylosa [Stephani (1888c: 289)] Steph. in Evans (1900: 454). Cololejeunea trichomanis (Gottsche 1882: 362) [Stephani (1889a: 168)] Bescherelle (1892: 14). Cololejeunea venusta (Sande Lacoste 1854) Schiffner (1893b: 122). Current name (Tixier 1985):—Cololejeunea haskarliana (Lehm. et Lindenb. in Lehmann 1844: 26) Schiffner (1898: 244) ‘Hasskarliana’. Cololejeunea wrightii [Stephani (1892b: 172); ICN Art. 38.1(a); no description]. Note:—The name seems never to have been validly published. Current name (Evans 1911):—Myriocoleopsis minutissima (Sm. in Smith & Sowerby 1806: tab. 1633) R.L.Zhu, Y.Yu et Pócs in Yu et al. (2014: 293) subsp. myriocarpa (Nees et Mont. in Montagne 1842: 473) R.L.Zhu, Y.Yu et Pócs in Yu et al. (2014: 294). Colurolejeunea acroloba [Stephani (1890b: 97); Stephani (1890c: 135)] (Prantl 1890: xiv) Schiffner (1900: 198). Current name:—Colura acroloba (Prantl 1890: xiv) Jovet-Ast (1953: 297). Colurolejeunea apiculata [Schiffner (1890: 36)] Schiffner (1900: 198). Current name:—Colura apiculata (Schiffner 1900: 198) Stephani (1916: 936). Colurolejeunea calyptrifolia (Hooker 1813: tab. 43) [Schiffner (1890: 35); Stephani (1890b: 97); Stephani (1890c: 135)] Schiffner (1893b: 121). Current name:—Colura calyptrifolia (Hooker 1813: tab. 43) Dumortier (1835: 12). Colurolejeunea ceratophora (Nees in Gottsche et al. 1845b: 405) [Schiffner (1890: 35); Stephani (1890b: 97); Stephani (1890c: 135)]. Note:—The name seems never to have been validly published. Current name (Jovet-Ast 1953):—Colura tenuicornis (Evans 1900: 455) Stephani (1916: 942). Colurolejeunea clavigera (Spruce 1884: 304) [Schiffner (1890: 36)]. Note:—The name seems never to have been validly published. Current name (Grolle & Zhu 2002):—Macrocolura sagittistipula (Spruce 1884: 3041) Schuster (1994: 233). Colurolejeunea corynophora (Nees, Lindenb. et Gottsche in Gottsche et al. 1843: 474) [Schiffner (1890: 35) ‘corynephora’]. Note:—The name seems never to have been validly published. Current name:—Colura corynophora (Gottsche et al. 1843: 474) Trevisan (1877: 402) ‘corynephora’. Colurolejeunea dusenii Stephani (1892c: 198). Current name:—Colura dusenii Stephani (1892c: 168). Colurolejeunea junghuhniana [Stephani (1890b: 97); Stephani (1890c: 135)] (Prantl 1890: xvi) Stephani (1896a: 74). Current name:— Colura junghuhniana (Prantl 1890: xvi) Stephani (1916: 938). Colurolejeunea minor [Schiffner (1890: 37)] Schiffn. ex Sydow (1894: 246). Current name (Solari 1983):—Colura naumannii (Schiffner 1893b: 121) Stephani (1916: 935). Colurolejeunea naumannii [Schiffner (1890: 36)] Schiffner (1893b: 121). Current name:—Colura naumannii (Schiffner 1893b: 121) Stephani (1916: 935). Colurolejeunea obtusa Stephani (1891a: 208). Current name (Jovet-Ast 1953):—Colura digitalis (Mitten 1886: 325) Stephani (1916: 931). Colurolejeunea ornata (Goebel 1890: 26) Schiffner (1893b: 121). Current name:—Colura ornata Goebel (1890: 26). Colurolejeunea sagittistipula (Spruce 1884: 304) [Schiffner (1890: 35)]. Note:—The name seems never to have been validly published. Current name:—Macrocolura sagittistipula (Spruce 1884: 3041) Schuster (1994: 233). Colurolejeunea superba (Montagne 1848: 115) [Schiffner (1890: 36)], Stephani (1896b:309) Current name:—Colura superba (Montagne 1848: 115) Stephani (1916: 941). 172 • Phytotaxa 220 (2) © 2015 Magnolia Press SÖDERSTRÖM ET AL. Colurolejeunea tortifolia (Nees et Mont. in Montagne 1843: 265) [Schiffner (1890: 35); Stephani (1890b: 98); Stephani (1890c: 135)]. Note:—The name seems never to have been validly published. Current name:—Colura tortifolia (Nees et Mont. in Montagne 1843: 265) Trevisan (1877: 402). Colurolejeunea zollingeri [Schiffner (1890: 35)]. Note:—The name seems never to have been validly published and we do not know where to refer it. Crossotolejeunea boryana (Montagne 1838: 47) [Stephani (1890b: 70); Stephani (1890c: 135)] Schiffner (1893b: 127). Current name:— Lejeunea boryana Montagne (1838: 47). Crossotolejeunea crenata (Nees et Mont. in Montagne 1838: 48) [Stephani (1890b: 70); Stephani (1890c: 135)] Lacouture (1908: 109). Current name:—Xylolejeunea crenata (Nees et Mont. in Montagne 1838: 48) He & Grolle (2001: 36). Crossotolejeunea inflexiloba (Prantl 1892: xvi) Stephani (1913: 239). Current name:—Lejeunea inflexiloba Prantl (1892: xvi). Crossotolejeunea intricata (Prantl 1892: xvi) [Jack & Stephani (1892: 17)] Stephani (1896a: 75). Current name:—Lejeunea intricata Prantl (1892: xvi). Dendrolejeunea vittata (Mitten 1873: 411) [Schiffner (1890: 25); Schiffn. ex Sydow (1894: 246)]. Note:—The name seems never to have been validly published. Current name (Schiffner 1898):—Thysananthus fruticosus (Lindenb. et Gottsche in Gottsche et al. 1847: 737) Schiffner (1893b: 130). Dicranolejeunea aberrans (Lindenb. et Gottsche in Gottsche et al.1847: 751) [Stephani (1890c: 136)] Stephani (1912b: 158). Current name:—Acanthocoleus aberrans (Lindenb. et Gottsche in Gottsche et al. 1847: 751) Kruijt (1988: 62). Dicranolejeunea axillaris (Nees & Montagne 1836: 59) [Stephani (1890c: 136); Jack & Stephani (1892: 13); Stephani (1892e: 179)] Schiffner (1893b: 128). Dicranolejeunea dubiosa (Lindenb. et Gottsche in Gottsche et al. 1847: 752) [Stephani (1890c: 136); Jack & Stephani (1892: 13)] Stephani (1912b: 157). Current name (Kruijt 1985):—Dicranolejeunea axillaris (Nees & Montagne 1836: 59) Schiffner (1893b: 128). Dicranolejeunea incongrua (Lindenb. et Gottsche in Gottsche et al. 1847: 750) [Stephani (1890c: 136)] Stephani (1912b: 160). Current name:—Blepharolejeunea incongrua (Lindenb. et Gottsche in Gottsche et al. 1847: 750) van Slageren et Kruijt (1985: 133). Dicranolejeunea mexicana (Lindenb. in Gottsche et al. 1845a: 325) [Stephani (1890a: 13); Stephani (1890c: 136); Fulford (1956: 229) ‘maxicana’ ICN Art. 38.1(a); no description]. Note:—The name seems never to have been validly published. Current name (Stephani 1912b):—Dicranolejeunea axillaris (Nees & Montagne 1836: 59) Schiffner (1893b: 128). Dicranolejeunea phyllorhiza (Nees 1833: 348) [Stephani (1890a: 7); Stephani (1890c: 136)] Schiffner (1893b: 128). Current name:— Brachiolejeunea phyllorhiza (Nees 1833: 348) Kruijt & Gradstein (1986: 299). Dicranolejeunea trigona (Nees & Montagne 1836: 61) [Stephani (1890c: 136)]. Note:—The name seems never to have been validly published. Current name:—Acanthocoleus trigonus (Nees & Montagne 1836: 61) Gradstein (1992a: 101). Diplasiolejeunea kraussiana (Lindenb. in Gottsche et al. 1845b: 393) [Stephani (1890b: 91); Stephani (1890c: 136)] Stephani (1916: 919). Diplasiolejeunea pellucida (C.F.W.Meissn. ex Sprengel 1827: 325) [Stephani (1888c: 288); Stephani (1890b: 91); Stephani (1890c: 136); Stephani (1892e: 179)] Schiffner (1893b: 121). Diplasiolejeunea unidentata (Lehm. et Lindenb. in Lehmann 1834: 48) [Stephani (1890b: 90); Stephani (1890c: 136)] Schiffner (1897: 583). Drepanolejeunea bidens [Stephani (1890b: 71); Stephani (1890c: 136)] (Prantl 1890: xiv) Evans (1903a: 29). Drepanolejeunea blumei [Steph. ex Schiffner (1893a: 231) ICN Art. 38.1(a); no description; Steph. ex Schiffner (1893b: 126) ICN Art. 38.1(a); no description] Stephani (1896a: 81). Drepanolejeunea capulata (Taylor 1846: 394) [Stephani (1890b: 71); Stephani (1890c: 136)] Jack & Stephani (1892: 13). Drepanolejeunea cristata Stephani (1891a: 209). Current name (Jones 1958):—Drepanolejeunea cultrella (Mitten 1863: 168) Stephani (1913: 324). Drepanolejeunea dactylophora (Nees, Lindenb. et Gottsche in Gottsche et al. 1843: 473) Jack & Stephani (1892: 12). Drepanolejeunea granatensis [Jack & Stephani (1892: 13) ICN Art. 38.1(a); no description] (Prantl 1892: xvi) Bischler (1964: 150). Drepanolejeunea grossidens [Stephani (1889a: 168)] Stephani (1913: 358). Drepanolejeunea hamatifolia (Hooker 1813: tab. 51) [Stephani (1888c: 286); Stephani (1890b: 71); Stephani (1890c: 136)] Schiffner (1893b: 126) ‘hamattfolia’. Drepanolejeunea hampeana [Stephani (1890b: 70); Stephani (1890c: 136)] Stephani (1913: 345). Drepanolejeunea inchoata (C.F.W.Meissn. in Lehmann 1833: 19) [Stephani (1888c: 287); Stephani (1890b: 70); Stephani (1890c: 136)] Stephani (1892e: 179). Drepanolejeunea lancifolia (Gottsche 1864: 155) Jack & Stephani (1892: 13). Drepanolejeunea latitans (Hooker & Taylor 1844a: 399) [Stephani (1890b: 72); Stephani (1890c: 136)] Stephani (1913: 358). Current name:—Microlejeunea latitans (Hooker & Taylor 1844a: 399) Heinrichs, Schäf.-Verw., Pócs et S.Dong in Dong et al. (2013: 52). Drepanolejeunea lyratifolia (Hook.f. et Taylor in Taylor 1846: 393) [Stephani (1890b: 71); Stephani (1890c: 136)]. Note:—The name seems never to have been validly published. Current name (Grolle 1979a):—Diplasiolejeunea plicatiloba (Hooker & Taylor 1845: 92) Grolle (1979a: 175). GENERA OF LEJEUNEACEAE ESTABLISHED IN 1884–1893 Phytotaxa 220 (2) © 2015 Magnolia Press • 173 Drepanolejeunea muricata (Gottsche in Gottsche et al. 1845b: 348) [Stephani (1890b: 74); Stephani (1890c: 136)] Schiffner (1893b: 126). Current name:—Dactylophorella muricata (Gottsche in Gottsche et al. 1845b: 348) Schuster (1980b: 427). Drepanolejeunea palmifolia (Nees 1833: 366) [Stephani (1890b: 74); Stephani (1890c: 136)] Schiffner (1893b: 126). Drepanolejeunea plicatiloba (Hooker & Taylor 1845: 92) [Stephani (1890b: 81); Stephani (1890c: 136)] Stephani (1913: 359). Current name:—Diplasiolejeunea plicatiloba (Hooker & Taylor 1845: 92) Grolle (1979a: 175). Drepanolejeunea securifolia (Gottsche 1882: 360) Pearson (1892: 8). Current name (Grolle 1995):—Drepanolejeunea physifolia (Gottsche 1882: 357) Pearson (1892: 8) ‘physaefolia’. Drepanolejeunea tenuis (Nees 1830: 57) [Stephani (1888c: 292); Stephani (1890b: 90); Stephani (1890c: 136)] Jack & Stephani (1892: 12). Current name (Mizutani 1963):—Drepanolejeunea angustifolia (Mitten 1860b: 116) Grolle (1965: 206). Drepanolejeunea ternatensis (Gottsche in Gottsche et al. 1845b: 346) [Stephani (1889a: 169); Stephani (1890b: 73); Stephani (1890c: 136)] Schiffner (1893b: 126). Drepanolejeunea teysmannii (Gottsche 1882: 360) [Stephani (1890b: 72); Stephani (1890c: 136)] Stephani (1896a: 84). Drepanolejeunea tridactyla (Gottsche in Gottsche et al. 1845b: 347) [Stephani (1890b: 73); Stephani (1890c: 136)] Stephani (1913: 354). Eulejeunea angusta (Lehm. et Lindenb. in Lehmann 1832: 52) Stephani (1890b: 86). Current name:—Lejeunea angusta (Lehm. et Lindenb. in Lehmann 1832: 52) Montagne (1842: 469). Eulejeunea armitii Stephani (1889a: 169). Current name:—Lejeunea armitii Stephani (1889a: 169) Stephani (1915: 768). Eulejeunea austinii (Lindberg 1875: 489) Stephani (1892b: 171). Current name (Schuster 1980a):—Cheilolejeunea rigidula (Nees ex Montagne 1840: 336) Schuster (1971: 102). Eulejeunea caroliniana (Austin 1876: 36) Stephani (1892b: 171). Current name:—Lejeunea caroliniana Austin (1876: 36). Eulejeunea caespitosa (Lindenb. in Gottsche et al. 1845b: 384) Stephani (1890b: 86). Current name (Reiner-Drehwald & SchäferVerwimp 2008):—Lejeunea capensis Gottsche in Gottsche et al. (1845b 374). Eulejeunea capensis (Gottsche in Gottsche et al. 1845b: 374) Stephani (1890b: 83). Current name:—Lejeunea capensis Gottsche in Gottsche et al. (1845b 374). Eulejeunea clavata (Lindenb. in Gottsche et al. 1845b: 379) Stephani (1890b: 85). Current name:—Lejeunea clavata Lindenb. in Gottsche et al. (1845b: 379). Eulejeunea concava (Lindenb. et Gottsche in Gottsche et al. 1847: 759) Stephani (1890b: 78). Current name:—Lejeunea concava Lindenb. et Gottsche in Gottsche et al. (1847: 759). Eulejeunea corallina Stephani (1890b: 83). Current name (Mizutani 1972):—Cheilolejeunea intertexta (Lindenb. in Gottsche et al. 1845b: 379) Stephani (1897c: 79). Eulejeunea crenulata Schiffner (1890: 33). Current name (Hürlimann 1993):—Lejeunea patersonii (Stephani 1896a: 92) Stephani (1915: 784). Eulejeunea cuculliloba [Stephani (1891a: 202) ICN Art. 38.1(a); no description; Prantl (1891: viii) ICN Art. 38.1(a); no description]. Note:—This is probably an error for Eulejeunea saccatiloba. Current name (Pócs 2010):—Lejeunea anisophylla Montagne (1843: 263). Eulejeunea denticalyx Stephani (1889a: 169). Current name:—Lejeunea denticalyx (Stephani 1889a: 169) Stephani (1915: 794). Eulejeunea diversiloba (Spruce 1876b: 235) Schiffner (1893b: 124). Current name:—Microlejeunea diversiloba (Spruce 1876b: 235) Müller (1915: 654). Eulejeunea diversistipa (Lindenb. in Gottsche et al. 1847: 765) Stephani (1890b: 85). Current name (Grolle & Reiner-Drehwald 1999):— Lejeunea ambigua Lindenb. et Gottsche in Gottsche et al. (1847: 764). Eulejeunea drummondii (Taylor 1846: 400) Stephani (1889a: 170). Current name:—Lejeunea drummondii Taylor (1846: 400). Eulejeunea ecarinata Stephani (1890d: 283). Current name:—Lejeunea ecarinata (Stephani 1890d: 283) Coulter et al. (1890: 349). Eulejeunea eckloniana (Lindenb. in Gottsche et al. 1845b: 381) Stephani (1890b: 86) ‘ecklonii’. Current name:—Lejeunea eckloniana Lindenb. in Gottsche et al. (1845b: 381). Eulejeunea flava (Swartz 1788: 144) Stephani (1888b: 109). Current name:—Lejeunea flava (Swartz 1788: 144) Nees (1838: 277). Eulejeunea glaucescens (Gottsche in Gottsche et al. 1845b: 378) Stephani (1888c: 286). Current name:—Lejeunea glaucescens (Gottsche in Gottsche et al. 1845b: 378) Stephani (1888c: 286).. Eulejeunea glaucophylla (Gottsche 1857: 345) Stephani (1890b: 87). Current name (Evans 1908b):—Lejeunea laetevirens Nees et Mont. in Montagne (1842: 469) ‘laete-virens’. Eulejeunea gottscheana (Lindenb. in Gottsche et al. 1845b: 382) Stephani (1890b: 86). Current name:—Lejeunea gottscheana Lindenb. in Gottsche et al. (1845b: 382). Eulejeunea haitica (Nees et Mont. in Montagne 1843: 263) Stephani (1890b: 89) ‘haitensis’. Current name:—Lejeunea haitica Nees et Mont. in Montagne (1843: 263). Eulejeunea hepaticola Stephani (1888a: 60). Current name:—Lejeunea hepaticola (Stephani 1888a: 60) Stephani (1915: 714). 174 • Phytotaxa 220 (2) © 2015 Magnolia Press SÖDERSTRÖM ET AL. Eulejeunea huctumalcensis (Lindenb. et Gottsche in Gottsche et al. 1847: 762) Stephani (1890b: 81). Current name:—Otigoniolejeunea huctumalcensis (Lindenb. et Gottsche in Gottsche et al. 1847: 762) Wei et al. (2014: 237). Eulejeunea infestans [Steph. ex Bescherelle (1892: 13) ICN Art. 38.1(a); no description] Stephani (1896a: 90). Current name (Lee 2013):—Lejeunea papilionacea Prantl (1892: xvii). Eulejeunea isomorpha (Gottsche 1882: 355) Stephani (1892a: 120). Current name (Jones 1968):—Lejeunea flava (Swartz 1788: 144) Nees (1838: 277) subsp. tabularis (Sprengel 1827: 325) Arnell (1963: 199). Eulejeunea laeta (Lehm. et Lindenb. in Lehmann 1834: 45) Stephani (1890b: 85). Current name:—Lejeunea laeta (Lehm. et Lindenb. in Lehmann 1834: 45) Lehm. et Lindenb. in Gottsche et al. (1845b: 380). Eulejeunea laetevirens (Nees et Mont. in Montagne 1842: 469) Stephani (1890b: 87). Current name:—Lejeunea laetevirens Nees et Mont. in Montagne (1842: 469) ‘laete-virens’. Eulejeunea laxa (Nees 1830: 43) Stephani (1890b: 85). Current name:—Lejeunea laxa (Nees 1830: 43) Lindenb. in Gottsche et al. (1845b: 378). Eulejeunea lepida (Lindenb. et Gottsche in Gottsche et al. 1847: 766) Stephani (1890b: 87). Current name (Reiner-Drehwald 2010a):— Lejeunea laetevirens Nees et Mont. in Montagne (1842: 469) ‘laete-virens’. Eulejeunea lucens (Taylor 1846: 399) Stephani (1890b: 84). Current name (Evans 1908b):—Lejeunea laetevirens Nees et Mont. in Montagne (1842: 469) ‘laete-virens’. Eulejeunea magnoliae (Lindenb. et Gottsche in Gottsche et al. 1847: 763) Stephani (1890b: 81). Current name (Reiner-Drehwald 2009):—Lejeunea adpressa Nees in von Flotow et al. (1842: 45). Eulejeunea muscicola (Spruce 1884: 281) Stephani (1888c: 300). Current name:—Lejeunea muscicola Spruce (1884: 281). Eulejeunea nudipes (Hooker & Taylor 1844c: 568) Stephani (1890b: 82). Current name:—Nephelolejeunea nudipes (Hooker & Taylor 1844c: 568) L.Söderstr. et A.Hagborg in Pócs et al. (2015: 66). Eulejeunea obtusata (Gottsche 1882: 354) Pearson (1892: 8). Lejeunea obtusata (Gottsche 1882: 354). Eulejeunea olivacea Stephani (1890b: 85). Current name:—Lejeunea subolivacea Mizutani (1965: 121). Eulejeunea pulvinata (Nees & Montagne 1836: 61) Stephani (1890b: 86). Current name:—Lejeunea pulvinata Nees & Montagne (1836: 61). Eulejeunea radicans (Lindenb. et Gottsche in Gottsche et al. 1847: 766) Stephani (1890b: 87). Current name:—Lejeunea radicans Lindenb. et Gottsche in Gottsche et al. (1847: 766). Eulejeunea remotifolia Stephani (1890b: 86) ICN Art. 38.1(a); no description; Stephani (1890c: 137) ICN Art. 38.1(a); no description. Note:—The name seems never to have been validly published. Current name:—Lejeunea remotifolia Stephani (1915: 756). Eulejeunea rodriguezii Stephani (1888b: 108). Current name (Pócs 2011):—Lejeunea anisophylla Montagne (1843: 263). Eulejeunea saccatiloba Stephani (1891a: 207). Current name (Pócs 2010):—Lejeunea anisophylla Montagne (1843: 263). Eulejeunea seriata (Lindenb. et Gottsche in Gottsche et al. 1847: 762) Stephani (1890b: 81). Current name:—Lejeunea seriata Lindenb. et Gottsche in Gottsche et al. (1847: 762). Eulejeunea serpyllifolia (Libert 1820: 374) Stephani (1890b: 83). Current name (Bonner & Miller 1960):—Lejeunea cavifolia (Ehrhart 1789: 45) Lindberg (1871: 43). Eulejeunea spiniloba (Lindenb. et Gottsche in Gottsche et al. 1847: 770) Stephani (1890b: 98). Current name:—Lejeunea spiniloba Lindenb. et Gottsche in Gottsche et al. (1847: 770). Eulejeunea subelobata (Carrington & Pearson 1888: 1039) Stephani (1889a: 170). Current name:—Lejeunea subelobata Carrington & Pearson (1888: 1039). Eulejeunea tabularis (Sprengel 1827: 325) Stephani (1890b: 83). Current name:—Lejeunea flava (Swartz 1788: 144) Nees (1838: 277) subsp. tabularis (Sprengel 1827: 325) Arnell (1963: 199). Eulejeunea ulicina (Taylor 1844: 115) Schiffner (1893b: 124). Current name:—Microlejeunea ulicina (Taylor 1844: 115) Stephani (1892b: 171). Eulejeunea underwoodii Stephani (1892b: 172). Current name (Schuster 1980a):—Cheilolejeunea rigidula (Nees ex Montagne 1840: 336) Schuster (1971: 102). Eulejeunea urbanii Stephani (1888c: 301) ‘Urbani’. Current name:—Lejeunea urbanii (Stephani 1888c: 301) Stephani (1915: 766) ‘Urbani’. Eulejeunea vogelii (Taylor 1846: 397) Stephani (1890b: 98). Current name (Pócs 2010):—Lejeunea anisophylla Montagne (1843: 263). Eulejeunea vulgariformis (Gottsche 1882: 355) Pearson (1892: 9). Current name:—Lejeunea vulgariformis Gottsche (1882: 355). Eulejeunea wallichiana (Lehmann 1831: 5) Stephani (1890b: 78). Current name:—Microlejeunea wallichiana (Lehmann 1831: 5) R.L.Zhu et Y.M.Wei in Wei & Zhu (2013: 308). Eulejeunea wightii (Lindenb. in Gottsche et al. 1845b: 379) Stephani (1890b: 85). Current name:—Lejeunea wightii Lindenb. in Gottsche et al. (1845b: 379). Eulejeunea wilmsii Stephani (1892a: 124). Current name (Jones 1974):—Lejeunea eckloniana Lindenb. in Gottsche et al. (1845b: 381). GENERA OF LEJEUNEACEAE ESTABLISHED IN 1884–1893 Phytotaxa 220 (2) © 2015 Magnolia Press • 175 Eulejeunea zacuapana Stephani (1890b: 87). Current name:—Lejeunea zacuapana (Stephani 1890b: 87) Prantl (1890: xviii). Euosmolejeunea beyrichii (Lindenb. in Gottsche et al. 1845b: 371) Stephani (1890b: 82). Current name:—Cheilolejeunea beyrichii (Lindenb. in Gottsche et al. 1845b: 371) Reiner-Drehwald (2006: 474). Euosmolejeunea brachytoma (Gottsche 1882: 355) Pearson (1892: 8). Current name (Grolle 1978):—Cheilolejeunea trifaria (Reinwardt et al. 1824: 226) Mizutani (1964: 132). Euosmolejeunea contigua (Nees 1833: 360) Stephani (1890b: 78). Current name:—Pycnolejeunea contigua (Nees 1833: 360) Grolle (1979a: 179). Euosmolejeunea duriuscula (Nees in Gottsche et al. 1845b: 364) Stephani (1890b: 80). Current name (Gradstein 1989):—Cheilolejeunea rigidula (Nees ex Montagne 1840: 336) Schuster (1971: 102). Euosmolejeunea implexicaulis (Hooker & Taylor 1844a: 397) Stephani (1890b: 84). Current name:—Cheilolejeunea implexicaulis (Hooker & Taylor 1844a: 397) Schuster (1963a: 245). Euosmolejeunea lindenbergii (Gottsche in Gottsche et al. 1845b: 336) Stephani (1890c: 137). Current name:—Cheilolejeunea lindenbergii (Gottsche in Gottsche et al. 1845b: 336) Mizutani (1970: 226). Euosmolejeunea lutea (Mont. ex Gottsche et al. 1845b: 383) Stephani (1890b: 86). Current name (Evans 1908b):—Cheilolejeunea clausa (Nees et Mont. in Montagne 1840: 337) Schuster (1980a: 863). Euosmolejeunea montagnei (Montagne 1843: 261) Stephani (1890c: 137). Current name:—Cheilolejeunea montagnei (Montagne 1843: 261) Schuster (1963a: 112). Euosmolejeunea opaca (Gottsche in Gottsche et al. 1845b: 352) Stephani (1890b: 79). Current name (Evans 1908b):—Cheilolejeunea clausa (Nees et Mont. in Montagne 1840: 337) Schuster (1980a: 863). Euosmolejeunea orientalis (Gottsche in Gottsche et al. 1845b: 371) Stephani (1890b: 82). Current name:—Cheilolejeunea orientalis (Gottsche in Gottsche et al. 1845b: 371) Mizutani (1972: 399). Euosmolejeunea parvistipula (Lindenb. et Gottsche in Gottsche et al. 1847: 761) Stephani (1890b: 80). Note:—The name has never been used in Cheilolejeunea and we do not know where to refer it. Euosmolejeunea sayeri Stephani (1889a: 170). Current name (Grolle 1979b):—Cheilolejeunea trifaria (Reinwardt et al. 1824: 226) Mizutani (1964: 132). Euosmolejeunea tenella (Taylor 1846: 398) Stephani (1890b: 87). Current name:—Lejeunea tenella Taylor (1846: 398). Euosmolejeunea trifaria (Reinwardt et al. 1824: 226) [Stephani (1888c: 292)] Schiffner (1890: 31). Current name:—Cheilolejeunea trifaria (Reinwardt et al. 1824: 226) Mizutani (1964: 132). Harpalejeunea ancistrodes (Spruce 1884: 169) Schiffner (1893b: 127) ‘ancystrodes’. Harpalejeunea cinchonae (Nees in Gottsche et al. 1845b: 342) [Stephani (1890b: 70); Stephani (1890c: 138); Jack & Stephani (1892: 13)] Schiffner (1893b: 127). Harpalejeunea decurvicuspis (Bescherelle & Massalongo 1886: 639) [Schiffner (1890: 29)] Schiffn. ex Sydow (1894: 246). Harpalejeunea granatensis [Jack & Stephani (1892: 17)] (Prantl 1892: xvi) Stephani (1913: 251). Current name:—Drepanolejeunea granatensis (Prantl 1892: xvi) Bischler (1964: 150). Harpalejeunea holostipa (Spruce 1884: 171) Schiffner (1893b: 127). Current name:—Cheilolejeunea holostipa (Spruce 1884: 171) Grolle et R.L.Zhu in Grolle et al. (2001: 1071). Harpalejeunea massalongoana [Schiffner (1890: 29)] Schiffn. ex Sydow (1894: 246). Current name:—Lejeunea massalongoana (Schiffn. ex Sydow 1894: 246) Solari (1983: 542). Harpalejeunea ovata (Hooker 1813: tab. 42) [Stephani (1890b: 84); Stephani (1890c: 138)] Schiffner (1893b: 127). Current name (Grolle 1989):—Harpalejeunea molleri (Stephani 1887: 3) Grolle (1989: 89). Harpalejeunea oxyphylla (Nees et Mont. in Montagne 1843: 264) [Stephani (1890b: 76); Stephani (1890c: 138)] Stephani (1913: 255). Harpalejeunea parasitica (Hooker & Taylor 1844b: 477) [Stephani (1890b: 85); Stephani (1890c: 138)] Stephani (1913: 268). Harpalejeunea patentissima (Hampe & Gottsche 1853: 355) [Stephani (1888c: 288)] Steph. in Schiffner (1897: 591). Current name:— Ceratolejeunea patentissima (Hampe & Gottsche 1853: 355) Evans (1905b: 286). Harpalejeunea savatieriana (Bescherelle & Massalongo 1886: 638) [Schiffner (1890: 29)] Schiffn. ex Sydow (1894: 246). Current name (Solari 1983):—Cheilolejeunea obtruncata (Montagne 1845b: 354) Solari (1983: 539). Harpalejeunea solitaria (Gottsche 1882: 356) [Pearson (1892: 7)] Stephani (1913: 246). Harpalejeunea stricta (Lindenb. et Gottsche in Gottsche et al. 1847: 756) [Stephani (1888c: 291); Stephani (1890b: 70); Stephani (1890c: 138)] Stephani (1913: 259). Harpalejeunea subfenestrata (Massalongo 1885: 249) [Schiffner (1890: 28)] Schiffn. ex Sydow (1894: 246). Current name (Solari 1983):—Harpalejeunea parasitica (Hooker & Taylor 1844b: 477) Stephani (1913: 268). Harpalejeunea tenuicuspis (Spruce 1884: 170) Schiffner (1893b: 127). Harpalejeunea tuberculata [Jack & Stephani (1892: 18)] (Prantl 1892: xvi) Stephani (1913: 261). Current name (Grolle & ReinerDrehwald 1999):—Harpalejeunea cinchonae (Nees in Gottsche et al. 1845b: 342) Schiffner (1893b: 127). 176 • Phytotaxa 220 (2) © 2015 Magnolia Press SÖDERSTRÖM ET AL. Homalolejeunea acutiloba (Hooker & Taylor 1845: 90) Stephani (1890c: 138). Current name (Geissler & Gradstein 1994):—Marchesinia brachiata (Swartz 1788: 144) Schiffner (1893b: 128). Homalolejeunea brachiata (Swartz 1788: 144) [Stephani (1890a: 14), ICN Art. 41.1; basionym not valid] Stephani (1890c: 138). Current name:—Marchesinia brachiata (Swartz 1788: 144) Schiffner (1893b: 128). Homalolejeunea excavata (Mitten 1860a: 58) Stephani (1891a: 202). Current name:—Marchesinia excavata (Mitten 1860a: 58) Schiffner (1893b: 128). Homalolejeunea fuscescens (Hampe in Lehmann 1838: 16) Stephani (1890c: 138). Current name (Gradstein & Buskes 1985):— Archilejeunea juliformis (Nees 1833: 351) Gradstein (1975: 127). Homalolejeunea guilleminiana (Nees et Mont. in Montagne 1841: 128) [Stephani (1890a: 14), ICN Art. 41.1; basionym not valid]. Note:—The name seems never to have been validly published. Current name (Geissler & Gradstein 1994):—Marchesinia brachiata (Swartz 1788: 144) Schiffner (1893b: 128). Homalolejeunea henriquesii Stephani (1888b: 112). Current name (Stephani 1896a):—Marchesinia excavata (Mitten 1860a: 58) Schiffner (1893b: 128). Homalolejeunea languida (Nees & Montagne 1836: 59) Stephani (1890c: 138). Current name:—Marchesinia languida (Nees & Montagne 1836: 59) Stephani (1912b: 149). Homalolejeunea mackaii (Hooker 1813: tab. 53) Stephani (1890c: 138) ‘mackayi’. Current name:—Marchesinia mackaii (Hooker 1813: tab. 53) Gray (1821: 689). Hygrolejeunea alata (Gottsche in Gottsche et al. 1845b: 406) [Stephani (1890b: 98); Stephani (1890c: 138)] Stephani (1914: 521). Current name:—Lejeunea alata Gottsche in Gottsche et al. (1845b: 406). Hygrolejeunea amboinensis [Schiffner (1890: 31)] Schiffn. ex Sydow (1894: 246). Current name (Mizutani 1970):—Lejeunea sordida (Nees 1830: 41) Nees (1838: 278). Hygrolejeunea cerina (Lehm. et Lindenb. in Lehmann 1833: 16) [Stephani (1888c: 284); Stephani (1890b: 90); Stephani (1890c: 138)] Schiffner (1893b: 125). Current name:—Lejeunea cerina (Lehm. et Lindenb. in Lehmann 1833: 16) Lehm. et Lindenb. in Gottsche et al. (1845b: 391). Hygrolejeunea chalmersii [Stephani (1889a: 171)] Stephani (1914: 558). Current name (Grolle 1979a):—Lejeunea microloba Taylor (1846: 399). Hygrolejeunea cordifissa (Taylor 1846: 395) [Stephani (1890b: 80); Stephani (1890c: 138)] Stephani (1914: 544). Current name:— Lepidolejeunea cordifissa (Taylor 1846: 395) Reiner-Drehwald (2006: 478). Hygrolejeunea delessertii (Nees et Mont. in Montagne 1843: 260) [Stephani (1890b: 74); Stephani (1890c: 138)]. Note:—The name seems never to have been validly published. Current name:—Lepidolejeunea delessertii (Nees et Mont. in Montagne 1843: 260) Grolle (1984: 505). Hygrolejeunea densistipula (Lehm. et Lindenb. in Lehmann 1838: 20) [Stephani (1890b: 81); Stephani (1890c: 138)]. Note:—The name seems never to have been validly published. Current name:—Pycnolejeunea densistipula (Lehm. et Lindenb. in Lehmann 1838: 20) Stephani (1914: 602). Hygrolejeunea deplanata (Nees in Gottsche et al. 1845b: 368) [Stephani (1890b: 81); Stephani (1890c: 138)] Stephani (1914: 534). Current name:—Lejeunea deplanata Nees in Gottsche et al. (1845b: 368). Hygrolejeunea discreta (Lindenb. in Gottsche et al. 1845b: 361) [Stephani (1890b: 78); Stephani (1890c: 138)] Schiffner (1898: 266). Current name:—Lejeunea discreta Lindenb. in Gottsche et al. (1845b: 361). Hygrolejeunea eluta (Nees 1833: 362) [Stephani (1890b: 78); Stephani (1890c: 138)] Massalongo (1911: 11). Current name:— Lepidolejeunea eluta (Nees 1833: 362) Schuster (1963b: 139). Hygrolejeunea heterocheila (Taylor 1846: 394) [Stephani (1890b: 95); Stephani (1890c: 138)]. Note:—The name seems never to have been validly published. Current name:—Lejeunea heterocheila Taylor (1846: 394). Hygrolejeunea latistipula [Schiffner (1890: 30)] Schiffn. ex Sydow (1894: 246). Current name:—Phaeolejeunea latistipula (Schiffn. ex Sydow 1894: 246) Mizutani (1968: 131). Hygrolejeunea leucophylla (Lindenb. in Gottsche et al. 1845b: 369) [Stephani (1890b: 81); Stephani (1890c: 138)] Stephani (1914: 537). Current name (Reiner-Drehwald 2010b):—Lejeunea deplanata Nees in Gottsche et al. (1845b: 368) var. deplanata. Hygrolejeunea lyratiflora [Stephani (1892c: 169)] (Prantl 1892: xvi) Stephani (1914: 524). Current name:—Lejeunea lyratiflora Prantl (1892: xvi). Hygrolejeunea microloba (Taylor 1846: 399) [Stephani (1890b: 85); Stephani (1890c: 138)] Stephani (1907: 294). Current name:— Lejeunea microloba Taylor (1846: 399). Hygrolejeunea norfolkensis [Stephani (1889a: 171)] Stephani (1914: 573) ‘norfolkiensis’. Current name (Renner 2013):—Lejeunea gracilipes (Taylor 1846: 385) Spruce (1884: 213). Hygrolejeunea orba (Gottsche in Gottsche et al. 1845b: 352) [Stephani (1890b: 95); Stephani (1890c: 138)] Stephani (1914: 547). Current name (Reiner-Drehwald 2006):—Lepidolejeunea involuta (Gottsche in Gottsche et al. 1845b: 350) Grolle (1984: 504). GENERA OF LEJEUNEACEAE ESTABLISHED IN 1884–1893 Phytotaxa 220 (2) © 2015 Magnolia Press • 177 Hygrolejeunea pallida (Lindenb. et Gottsche in Gottsche et al. 1847: 762) [Stephani (1890b: 81); Stephani (1890c: 138)] Stephani (1914: 538). Current name:—Lejeunea pallida Lindenb. et Gottsche in Gottsche et al. (1847: 762). Hygrolejeunea papilionacea [Stephani (1892c: 169)] (Prantl 1892: xvii) Stephani (1914: 525). Current name:—Lejeunea papilionacea Prantl (1892: xvii). Hygrolejeunea phyllobola (Nees et Mont. in Montagne 1842: 471) [Stephani (1890b: 81); Stephani (1890c: 138); Stephani (1892e: 179)] Massalongo (1897: 33). Current name:—Lejeunea phyllobola Nees et Mont. in Montagne (1842: 471). Hygrolejeunea pterota (Hooker & Taylor 1845: 91) [Stephani (1890b: 81); Stephani (1890c: 138)] Stephani (1914: 527). Current name (Grolle 1988):—Cheilolejeunea ascensionis (Hooker & Taylor 1845: 91) Grolle (1988: 45). Hygrolejeunea pulcherrima (Stephani 1886: 87) [Stephani (1891a: 202)] Bescherelle (1893b: 60). Current name (Vanden Berghen 1961):—Lejeunea acuta Mitten (1863: 167). Hygrolejeunea reflexistipula (Lehm. et Lindenb. in Lehmann 1833: 10) [Stephani (1890c: 138); Jack & Stephani (1892: 13)] Schiffner (1893b: 125). Current name:—Lejeunea reflexistipula (Lehm. et Lindenb. in Lehmann 1833: 10) Lehm. et Lindenb. in Gottsche et al. (1845b: 335). Hygrolejeunea rostrata [Stephani (1889a: 172)] Stephani (1896a: 104). Current name (Renner 2013):—Lejeunea lumbricoides (Nees 1830: 40) Nees in Gottsche et al. (1845b: 342). Hygrolejeunea sacculifera [Stephani (1889a: 172)] Stephani (1914: 574). Current name (Grolle 1981):—Lejeunea sordida (Nees 1830: 41) Nees (1838: 278). Hygrolejeunea sayeri [Stephani (1889a: 173)] Stephani (1914: 575). Current name (Piippo 1986):—Lepidolejeunea bidentula (Steph. in Jack & Stephani 1894: 107) Schuster (1980b: 425). Hygrolejeunea sordida (Nees 1830: 41) [Stephani (1890b: 80); Stephani (1890c: 138)] Schiffner (1893b: 125). Current name:—Lejeunea sordida (Nees 1830: 41) Nees (1838: 278). Hygrolejeunea spongia (Spruce 1876a: 132) Schiffner (1893b: 125). Current name (Reiner-Drehwald 2006):—Lepidolejeunea sullivantii (Gottsche 1863: 196) Reiner-Drehwald (2006: 479). Hygrolejeunea umbilicata (Nees 1830: 42) [Stephani (1890c: 138)] Schiffner (1893b: 125). Current name:—Lejeunea umbilicata (Nees 1830: 42) Nees in Gottsche et al. (1843: 472). Lejeunea acroloba Prantl (1890: xiv). Current name:—Colura acroloba (Prantl 1890: xiv) Jovet-Ast (1953: 297). Lejeunea amazonica Prantl (1890: xiv), nom. illeg. (ICN Art. 53.1; non Spruce 1884: 106). Current name (Dauphin 2003):—Ceratolejeunea cubensis (Montagne 1842: 481) Schiffner (1893b: 125). Lejeunea apiculata (Gottsche 1864: 149) Prantl (1892: xvii) nom. illeg. (ICN Art. 53.1; non Sande Lacoste 1854: 421). Current name:— Taxilejeunea apiculata (Gottsche 1864: 149) Jack & Stephani (1892: 13). Lejeunea bidens Prantl 1890: xiv. Current name:—Drepanolejeunea bidens (Prantl 1890: xiv) Evans (1903a: 29). Lejeunea cardotii (Stephani 1892b: 72) Coulter et al. (1892: 431) ‘cardoti’. Current name (Reiner-Drehwald 1994):—Microlejeunea globosa (Spruce 1889: cxciii) Stephani (1915: 821). Lejeunea cristata (Stephani 1891a: 210) Prantl (1891: ix). Current name (Jones 1958):—Drepanolejeunea cultrella (Mitten 1863: 168) Stephani (1913: 324). Lejeunea cuneifolia (Stephani 1892c: 166) Prantl (1892: xvi). Current name:—Cololejeunea cuneifolia Stephani (1892c: 166). Lejeunea dusenii (Stephani 1891a: 209) Prantl (1891: ix). Current name (Jones 1967):—Lejeunea ramosissima Stephani (1886: 88). Lejeunea dusenii (Stephani 1892c: 168) Prantl (1892: xvi), nom. illeg. (ICN Art. 53.1; non Prantl 1891: ix). Current name:—Colura dusenii Stephani (1892c: 168). Lejeunea ecarinata (Stephani 1890d: 283) Coulter et al. (1890: 349). Lejeunea elegans (Stephani 1891a: 208) Prantl (1891: ix), nom. illeg. (ICN Art. 53.1; non Gottsche in Gottsche et al. 1845b: 364). Current name:—Cololejeunea elegans Stephani (1891a: 208). Lejeunea epiphyta (Stephani 1891a: 206) Prantl (1891: ix), nom. illeg. (ICN Art. 53.1; non Gottsche in Gottsche et al. 1845b: 391). Current name (Jones 1967):—Lejeunea conformis Nees et Mont. in Montagne (1843: 260). Lejeunea fallax-vellongicornis [Stephani (1890b: 96) ICN Art. 38.1(a); no description]. Note:—The name seems never to have been validly published and Stephani (1890b) also placed it in synonymy. Current name (Stephani 1890b):—Ceratolejeunea cornuta (Lindenberg 1829: 23) Stephani (1895b: 65). Lejeunea filicaulis (Stephani 1892c: 168) Prantl (1892: xvi). Current name (Tixier 1985):—Cololejeunea hildebrandii (Austin 1876: 35) Stephani (1897a: 842) ‘Hillebrandii’. Lejeunea gottschei (Jack & Stephani 1892: 20) Prantl (1892: xvii). Current name:—Thysananthus gottschei (Jack & Stephani 1892: 20) Stephani (1912a: 787). Lejeunea granatensis Prantl (1892: xvi). Current name:—Drepanolejeunea granatensis (Prantl 1892: xvi) Bischler (1964: 150). Lejeunea grandiloba (Jack & Stephani 1892: 16) Prantl (1892: xvi). Current name:—Ceratolejeunea grandiloba Jack & Stephani (1892: 16). 178 • Phytotaxa 220 (2) © 2015 Magnolia Press SÖDERSTRÖM ET AL. Lejeunea hampeana Prantl (1890: xvi), nom. illeg. (ICN Art. 53.1; non Lindenb. in Gottsche et al. 1845b: 360). Current name:— Drepanolejeunea hampeana Stephani (1913: 345). Lejeunea inflexiloba Prantl (1892: xvi). Lejeunea intricata Prantl (1892: xvi). Lejeunea jackii Prantl (1892: xvii). Current name:—Omphalanthus jackii (Prantl 1892: xvii) Gradst. in Gradstein et al. (1977: 410). Lejeunea junghuhniana Prantl (1890: xvi). Current name:—Colura junghuhniana (Prantl 1890: xvi) Stephani (1916: 938). Lejeunea kegelii Prantl (1890: xvi), nom. illeg. (ICN Art. 53.1; non Lindenberg & Gottsche 1851: 635). Current name:—Cololejeunea kegelii Stephani (1895a: 249). Lejeunea kindbergii Prantl (1892: xvii). Current name (Grolle 1978):—Prionolejeunea grata (Gottsche 1882: 359) Schiffner (1893b: 127). Lejeunea kurzii (Stephani 1890d: 284) Coulter et al. (1890: 349). Current name (Kitagawa 1981):—Cheilolejeunea intertexta (Lindenb. in Gottsche et al. 1845b: 379) Stephani (1897c: 79). Lejeunea laevicalyx Prantl (1892: xvii), nom. illeg. (ICN Art. 53.1; non Gottsche 1863: 221). Current name:—Cheilolejeunea laevicalyx (J.B.Jack et Steph. in Stephani 1913: 283) Grolle (1988: 403). Lejeunea lyratiflora Prantl (1892: xvi). Lejeunea mascarena Coulter et al. (1890: 349). Current name (Grolle 1978):—Ceratolejeunea belangeriana (Gottsche in Gottsche et al. 1845b: 398) Stephani (1913: 396). Lejeunea mauritiana Coulter et al. (1890: 349). Current name (Grolle 1995):—Ceratolejeunea belangeriana (Gottsche in Gottsche et al. 1845b: 398) Stephani (1913: 396). Lejeunea microstipula (Stephani 1890b: 89) Prantl (1890: xvi). Current name (Pócs et al. 2011):—Lejeunea exilis (Reinwardt et al. 1824: 227) Grolle (1979b: 353). Lejeunea miradorensis Prantl (1890: xvi). Current name (Dauphin 2003):—Ceratolejeunea cornuta (Lindenberg 1829: 23) Stephani (1895b: 65). Lejeunea montagnei Prantl (1890: xvi), nom. illeg. (ICN Art. 53.1; non Montagne 1843: 261). Current name (Schuster 1980b):— Cololejeunea camillii (Lehmann 1857: 15) Evans (1912b: 59) ‘Camilli’. Lejeunea multilacera (Stephani 1890d: 285) Coulter et al. (1890: 349). Current name:—Lopholejeunea multilacera Stephani (1890d: 285). Lejeunea nigra (Stephani 1891a: 206) Prantl (1891: ix). Current name:—Mastigolejeunea nigra Stephani (1891a: 206). Lejeunea obtusa (Stephani 1891a: 208) Prantl (1891: ix). Current name (Jovet-Ast 1953):—Colura digitalis (Mitten 1886: 325) Stephani (1916: 931). Lejeunea papilionacea Prantl (1892: xvii). Lejeunea papilliloba Stephani (1890b: 73). Current name:—Cololejeunea papilliloba Stephani (1890b: 73) Stephani (1895a: 250). Lejeunea parviloba (Stephani 1890d: 286) Coulter et al. (1890: 349), nom. illeg. [ICN Art. 53.1; non Ångström (1877: 87)]. Current name:—Schiffneriolejeunea parviloba (Stephani 1890d: 286) Gradstein (1974: 335). Lejeunea plagiochiloides Stephani (1889b: 257). Current name (van Slageren 1985):—Spruceanthus semirepandus (Nees 1830: 39) Verdoorn (1934b: 153). Lejeunea poeppigiana (Nees in Gottsche et al. 1845b: 396) Stephani (1890b: 92). Current name (Dauphin 2003):—Ceratolejeunea ceratantha (Nees et Mont. in Montagne 1840: 335) Schiffner (1897: 582). Lejeunea pyriformis Stephani (1890b: 93), nom. illeg. [ICN Art. 53.1; non Lindenb. et Gottsche in Gottsche et al. (1847: 767)]. Current name (Dauphin 2003):—Ceratolejeunea cubensis (Montagne 1842: 481) Schiffner (1893b: 125). Lejeunea quintasii (Stephani 1889b: 270) Prantl (1891: ix). Current name (Tixier 1995):—Leptolejeunea epiphylla (Mitten 1860b: 118) Stephani (1913: 380). Lejeunea renauldii Coulter et al. (1890: 349). Current name (Grolle 1977):—Lejeunea alata Gottsche in Gottsche et al. (1845b: 406). Lejeunea saccatiloba (Stephani 1891a: 207) Prantl (1891: ix). Current name (Pócs 2010):—Lejeunea anisophylla Montagne (1843: 263). Lejeunea sandvicensis Prantl (1890: xvii). Current name:—Cheilolejeunea sandvicensis (Prantl 1890: xvii) Stephani (1897a: 842). Lejeunea schwaneckei Stephani (1888c: 290). Current name:—Pycnolejeunea schwaneckei (Stephani 1888c: 290) Stephani (1914: 606). Lejeunea stricta Prantl (1890: xvii) nom. illeg. (ICN Art. 53.1; non Lindenb. et Gottsche in Gottsche et al. 1875: 756). Current name: Microlejeunea diversiloba (Spruce 1876a: 235) Müller (1915: 654). Lejeunea taitica (Gottsche 1864: 147) Prantl (1890: xvii). Current name (Mizutani 1986):—Mastigolejeunea virens (Ångström 1873: 131) Stephani (1912a: 776). Lejeunea truncatiloba (Stephani 1892c: 170) Prantl (1892: xvii). Current name (Grolle 1976a):—Leptolejeunea maculata (Mitten 1860b: 118) Schiffner (1898: 275). GENERA OF LEJEUNEACEAE ESTABLISHED IN 1884–1893 Phytotaxa 220 (2) © 2015 Magnolia Press • 179 Lejeunea tuberculata Prantl (1892: xvi). Current name (Grolle & Reiner-Drehwald 1999):—Harpalejeunea cinchonae (Nees in Gottsche et al. 1845b: 342) Schiffner (1893b: 127). Lejeunea turgida (Stephani 1892c: 170) Prantl (1892: xvii) nom. illeg. (ICN Art. 53.1; non Mitten 1860b: 110). Current name:— Mastigolejeunea turgida Stephani (1892c: 170). Lejeunea underwoodii (Stephani 1892b: 172) Coulter et al. (1892: 431). Current name (Schuster 1980a: 171):—Cheilolejeunea rigidula (Nees ex Montagne 1840: 336) Schuster (1971: 102). Lejeunea wallisii Prantl (1892: xvii). Current name:—Omphalanthus filiformis (Swartz 1788: 144) Nees in Gottsche et al. (1845a: 304) var. wallisii (Prantl 1892: xvii) Gradstein (2013b: 46). Lejeunea wilmsii (Stephani 1892a: 124) Prantl (1892: xvi). Current name (Jones 1974):—Lejeunea eckloniana Lindenb. in Gottsche et al. (1845b: 381). Lejeunea zacuapana (Stephani 1890b: 87) Prantl (1890: xviii). Leptolejeunea australis [Stephani (1889a: 173)] Stephani (1913: 389). Leptolejeunea corynophora (Gottsche et al. 1843: 474) [Stephani (1890b: 97) ‘corynephora’; Stephani (1890c: 138) ‘corynephora’] Schiffner (1893b: 126) ‘corynephora’. Current name:—Colura corynophora (Gottsche et al. 1843: 474) Trevisan (1877: 402) ‘corynephora’. Leptolejeunea denticulata [Stephani (1889a: 174); Stephani (1890b: 70); Stephani (1890c: 138)] Stephani (1913: 389). Leptolejeunea elliptica (Lehm. et Lindenb. in Lehmann 1833: 13) [Stephani (1888c: 285); Stephani (1890b: 96); Stephani (1890c: 138)] Bescherelle (1892: 14). Leptolejeunea neesii (Mont. in Nees & Montagne 1836: 62) [Stephani (1890b: 74); Stephani (1890c: 138)] Stephani (1913: 372). Current name:—Siphonolejeunea neesii (Mont. in Nees & Montagne 1836: 62) Bischler (1969: 338). Leptolejeunea polyrhiza (Nees in Gottsche et al. 1845b: 403) [Stephani (1890b: 96); Stephani (1890c: 138)] Schiffner (1898: 275). Current name:—Drepanolejeunea polyrhiza (Nees in Gottsche et al. 1845b: 403) Grolle & Zhu (2000: 392). Leptolejeunea quintasii Stephani (1891b: 270). Current name (Tixier 1995):—Leptolejeunea epiphylla (Mitten 1860b: 118) Stephani (1913: 380). Leptolejeunea rosulans [Stephani (1889a: 174)] Stephani (1913: 390). Leptolejeunea spicata [Bescherelle (1892: 14)] Stephani (1896a: 108). Current name:—Drepanolejeunea spicata (Stephani 1896a: 108) Grolle & Zhu (2000: 384). Leptolejeunea stenophylla (Lindenb. et Gottsche in Gottsche et al. 1847: 769) [Stephani (1890b: 96); Stephani (1890c: 138)] Schiffner (1893b: 117). Current name (Bischler 1967):—Drepanolejeunea orthophylla (Nees et Mont. in Montagne 1843: 265) Bischler (1967: 102). Leptolejeunea trematodes (Nees in Nees & Montagne 1836: 62) [Stephani (1890b: 74); Stephani (1890c: 138)] Stephani (1913: 376). Current name:—Drepanolejeunea trematodes (Nees in Nees & Montagne 1836: 62) Bischler (1967: 125). Leptolejeunea truncatiloba Stephani (1892c: 170). Current name (Grolle 1976a):—Leptolejeunea maculata (Mitten 1860b: 118) Schiffner (1898: 275). Leptolejeunea vitrea (Nees 1830: 56) [Stephani (1890b: 96); Stephani (1890c: 138)] Schiffner (1893b: 126). Lopholejeunea applanata (Reinwardt et al. 1824: 210) [Stephani (1890c: 139) ‘adplanata’] Schiffner (1893b: 129). Lopholejeunea eulopha (Taylor 1846: 391) [Schiffner (1890: 28); Stephani (1890c: 139)] Schiffner (1893b: 129). Lopholejeunea fimbriata (Gottsche 1880: 64) [Schiffner (1890: 28)] Pearson (1892: 5). Current name (Verdoorn 1934b):—Lopholejeunea eulopha (Taylor 1846: 391) Schiffner (1893b: 129). Lopholejeunea harpaphylla (Spruce 1884: 123) Pearson (1892: 7). Current name (Stephani 1912b):—Caudalejeunea lehmanniana (Gottsche in Gottsche et al. 1845b: 325) Evans (1907b: 554). Lopholejeunea javanica (Nees in Gottsche et al. 1845b: 320) [Stephani (1890c: 139)] Schiffner (1893b: 129). Current name (Gradstein et al. 2002):—Lopholejeunea nigricans (Lindenb. in Gottsche et al. 1845b: 316) Schiffner (1898: 293). Lopholejeunea lepidoscypha Kiaer et Pearson in Pearson (1892: 5). Lopholejeunea muelleriana (Gottsche 1863: 184) [Stephani (1890c: 139)] Schiffner (1897: 599). Current name (Zhu & Gradstein 2005):—Lopholejeunea nigricans (Lindenb. in Gottsche et al. 1845b: 316) Schiffner (1898: 293). Lopholejeunea multilacera Stephani (1890d: 285). Lopholejeunea nigricans (Lindenb. in Gottsche et al. 1845a: 316) [Stephani (1890a: 16); Stephani (1890c: 139)] Schiffner (1898: 293). Lopholejeunea norfolkensis [Stephani (1889a: 175)] Stephani (1912b: 96) ‘Norfolkiensis’. Current name (Thiers & Gradstein 1989):— Lopholejeunea eulopha (Taylor 1846: 391) Schiffner (1893b: 129). Lopholejeunea plicatiscypha (Taylor 1846: 386) [Stephani (1890c: 139)] Stephani (1912b: 96). Lopholejeunea rusbyi (Spruce 1890: 122) Pearson (1892: 6). Current name (Gradstein 1994):—Lopholejeunea nigricans (Lindenb. in Gottsche et al. 1845b: 316) Schiffner (1898: 293). Lopholejeunea sagrana (Montagne 1842: 464) [Schiffner (1890: 27) ‘sagreana’; Stephani (1890c: 139) ‘sagraeana’] Stephani (1891a: 202) ‘sagraeana’. Current name (Verdoorn 1934a):—Lopholejeunea subfusca (Nees 1830: 36) Schiffner (1897: 593). 180 • Phytotaxa 220 (2) © 2015 Magnolia Press SÖDERSTRÖM ET AL. Lopholejeunea sphaerophora (Lehm. et Lindenb. in Lehmann 1833) [Stephani (1890c: 139)] Stephani (1912b: 68). Lopholejeunea zollingeri [Stephani (1890a: 14); Stephani (1890c: 139)] Schiffner (1898: 296). Macrolejeunea pallescens (Mitten 1851: 360) Schiffner (1893b: 125). Current name:—Lejeunea pallescens Mitten (1851: 360). Macrolejeunea subsimplex (Mont. et Nees in Montagne 1843: 264) [Stephani (1888c; 292, 301); Stephani (1890b: 90); Stephani (1890c: 139)] Schiffner (1893b: 125). Current name (Grolle 1987):—Lejeunea cerina (Lehm. et Lindenb. in Lehmann 1833: 16) Lehm. et Lindenb. in Gottsche et al. (1845b: 391). Mastigolejeunea amboinensis [Schiffner (1890: 22); Schiffn. ex Sydow (1894: 245), ICN Art. 36.1(a); not accepted by author]. Note:— The name seems never to have been validly published. Current name (Stephani 1912a):—Thysananthus spathulistipus (Reinwardt et al. 1824: 212) Lindenb. in Gottsche et al. (1845a: 287). Mastigolejeunea anguiformis (Hooker & Taylor 1844c: 567) [Stephani (1889b: 257)] Thiers & Gradstein (1989: 75). Current name:— Thysananthus anguiformis (Hooker & Taylor 1844c: 567) Taylor ex Gottsche et al. (1845a: 289). Mastigolejeunea arcuata (Nees 1830: 38) [Stephani (1890c: 139)] Schiffner (1894: 183). Current name:—Acrolejeunea arcuata (Nees 1830: 38) Grolle et Gradst. in Gradstein (1974: 332). Mastigolejeunea atypos [Schiffner (1890: 22)] Schiffn. ex Sydow (1894: 246). Current name (Grolle & Piippo 1984):—Mastigolejeunea repleta (Taylor 1846: 392) Evans (1902: 131). Mastigolejeunea auriculata (Wilson 1841: no. 170) [Stephani (1890c: 139)] Stephani (1892b: 171). Mastigolejeunea buettneri [Stephani (1888b: 110) ‘Büttneri’; Stephani (1889b: 257)] Schiffner (1893b: 129) ‘Büttneri’. Current name (Sukkharak & Gradstein 2014):—Mastigolejeunea humilis (Gottsche in Gottsche et al. 1845a: 239) Schiffner (1893b: 129). Mastigolejeunea convoluta (Lindenb. in Gottsche et al. 1845a: 288) [Schiffner (1890: 21)] Schiffn. ex Sydow (1894: 245). Current name:—Thysananthus convolutus Lindenb. in Gottsche et al. (1845a: 288). Mastigolejeunea crispula [Stephani (1888b: 111); Stephani (1889b: 257); Stephani (1892e: 180) ICN Art. 41.1; basionym not valid] Stephani (1912a: 759). Current name (Vanden Berghen 1949, Gradstein & Inoue 1980):—Mastigolejeunea auriculata (Wilson 1841: no. 170) Stephani (1892b: 171). Mastigolejeunea frauenfeldii (Reichardt 1866: 958) [Stephani (1890c: 139)] Verdoorn (1934a: 230). Mastigolejeunea humilis (Gottsche in Gottsche et al. 1845a: 239) [Stephani (1890c: 139)] Schiffner (1893b: 129). Mastigolejeunea innovans (Spruce 1884: 103) [Stephani (1889b: 258)] Stephani (1912a: 765). Mastigolejeunea ligulata (Lehm. et Lindenb. in Lehmann 1834: 39) [Stephani (1890a: 8); Stephani (1890c: 139)] Schiffner (1893b: 129). Mastigolejeunea longidens [Stephani (1889b: 258) also ICN Art. 38.1(a); no description]. Note:—The name seems never to have been validly published and we do not know where to refer it. Mastigolejeunea minuta [Schiffner (1890: 23)] Schiffn. ex Sydow (1894: 246). Current name (Grolle & Piippo 1984):—Mastigolejeunea humilis (Gottsche in Gottsche et al. 1845a: 239) Schiffner (1893b: 129). Mastigolejeunea nigra [Stephani (1889b: 258)] Stephani (1891a: 206). Mastigolejeunea novo-hibernica [Schiffner (1890: 23); Schiffn. ex Sydow (1894: 246) ICN Art. 36.1(a); not accepted by author] Stephani (1912b: 770). Current name (Grolle & Piippo 1984):—Mastigolejeunea ligulata (Lehm. et Lindenb. in Lehmann 1834: 39) Schiffner (1893b: 129). Mastigolejeunea phaea [Gottsche ex Stephani (1889b: 257)] Steph. ex Schiffner (1893b: 129). Current name (Thiers & Gradstein 1989):—Spruceanthus thozetianus (Gottsche et F.Muell. in Gottsche 1880: 63) Thiers & Gradstein (1989: 62). Mastigolejeunea plicatiflora (Spruce 1884: 104) [Stephani (1889b: 258)] Stephani (1912a: 766). Mastigolejeunea repleta (Taylor 1846: 392) [Stephani (1890c: 139)] Evans (1902: 131). Mastigolejeunea taitica (Gottsche 1864: 147) [Stephani (1890c: 139)] Jack & Stephani (1894: 104)). Current name (Mizutani 1986):— Mastigolejeunea virens (Ångström 1873: 131) Stephani (1912a: 776). Mastigolejeunea turgida Stephani (1892c: 170). Mastigolejeunea versicolor (Lehm. et Lindenb. in Gottsche et al. 1845a: 297) [Stephani (1890a: 8)]. Note:—The name seems never to have been validly published. Current name (Schuster 1980a):—Mastigolejeunea auriculata (Wilson 1841: no. 170) Stephani (1892b: 171). Microlejeunea africana Stephani (1888a: 61). Microlejeunea albicans (Nees in Gottsche et al. 1845b: 386) Stephani (1890b: 88). Current name (Miller et al. 1962):—Metalejeunea cucullata (Reinwardt et al. 1824: 227) Grolle (1995: 100). Microlejeunea albovirens (Hooker & Taylor 1844a: 397) Stephani (1890b: 88) ‘albo-virens’. Current name:—Cheilolejeunea albovirens (Hooker & Taylor 1844a: 397) Hodgson (1962: 127). Microlejeunea bullata (Taylor 1846: 398) Stephani (1890b: 90). Microlejeunea byssoides (Gottsche 1882: 358) Pearson (1892: 9). Microlejeunea cardotii Stephani (1892d: 172) ‘Cardoti’. Current name (Reiner-Drehwald 1994):—Microlejeunea globosa (Spruce 1889: cxciii) Stephani (1915: 821). GENERA OF LEJEUNEACEAE ESTABLISHED IN 1884–1893 Phytotaxa 220 (2) © 2015 Magnolia Press • 181 Microlejeunea cochlearifolia Stephani (1888b: 113). Microlejeunea erectifolia (Spruce 1887: 36) Stephani (1889b: 259). Current name (Grolle 1995):—Metalejeunea cucullata (Reinwardt et al. 1824: 227) Grolle (1995: 100). Microlejeunea gracillima (Mitten 1860b: 115) Stephani (1890c: 139). Current name (Hattori 1966):—Microlejeunea punctiformis (Taylor 1846: 398) Stephani (1915: 832). Microlejeunea lucens (Taylor 1846: 399) Stephani (1892b: 171). Current name (Evans 1908b):—Lejeunea laetevirens Nees et Mont. in Montagne (1842: 469) ‘laete-virens’. Microlejeunea microstipula Stephani (1890b: 89). Current name (Pócs et al. 2011):—Lejeunea exilis (Reinwardt et al. 1824: 227) Grolle (1979b: 353). Microlejeunea oblongistipula (Gottsche 1882) Pearson (1892: 9). Microlejeunea ovifolia Stephani (1888c: 287). Current name (Bischler et al. 1961):—Microlejeunea bullata (Taylor 1846: 398) Stephani (1890b: 90). Microlejeunea papulosa (Gottsche 1882: 358) Pearson (1892: 9). Microlejeunea parallela Schiffner (1890: 33). Current name (Miller et al. 1962):—Metalejeunea cucullata (Reinwardt et al. 1824: 227) Grolle (1995: 100). Microlejeunea primordialis (Hooker & Taylor 1845: 92) Stephani (1890b: 84). Current name:—Lejeunea primordialis (Hooker & Taylor 1845: 92) Gottsche et al. (1845b: 375). Microlejeunea punctiformis (Taylor 1846: 398) Stephani (1890b: 90). Microlejeunea stricta Stephani (1890b: 90). Current name:—Microlejeunea diversiloba (Spruce 1876b: 235) Müller (1915: 654). Microlejeunea ulicina (Taylor 1844: 115) Stephani (1890b: 88). Neurolejeunea catenulata (Nees in Gottsche et al. 1845b: 323) [Stephani (1892b: 171)] Schiffner (1893b: 131). Neurolejeunea portoricensis (Hampe & Gottsche 1853: 352) [Stephani (1888c: 288)] Schiffner (1893b: 131). Current name:— Otigoniolejeunea portoricensis (Hampe & Gottsche 1853: 352) Wei et al. (2014: 237). Neurolejeunea seminervis (Spruce 1884: 84) Schiffner (1893b: 131). Odontolejeunea accedens (Gottsche in Gottsche et al. 1845b: 339) [Stephani (1888c: 281); Stephani (1890b: 69); Stephani (1890c: 139)]. Note:—The name seems never to have been validly published. Current name:—Cyclolejeunea accedens (Gottsche in Gottsche et al. 1845b: 339) Evans (1904: 201). Odontolejeunea berteroana [Stephani (1888c: 282)]. Note:—The name seems never to have been validly published. Current name (see below):—Rectolejeunea versifolia (Schiffner 1897: 597) L.Söderstr. et A.Hagborg, comb. nov. Odontolejeunea convexistipa (Lehm. et Lindenb. in Lehmann 1834: 43) [Stephani (1888c: 284); Stephani (1890a: 20); Stephani (1890c: 139)].Schiffner (1897: 583). Current name:—Cyclolejeunea convexistipa (Lehm. et Lindenb. in Lehmann 1834: 43) Evans (1904: 198). Odontolejeunea lunulata (Weber 1815: 33) [Stephani (1888c: 287); Stephani (1890b: 98); Stephani (1890c: 139)] Schiffner (1893b: 128). Odontolejeunea peruviana (Lehm. et Lindenb. in Lehmann 1833: 18) [Stephani (1890a: 19); Stephani (1890b: 69); Stephani (1890c: 139)] Schiffner (1893b: 128). Current name:—Cyclolejeunea peruviana (Lehm. et Lindenb. in Lehmann 1833: 18) Evans (1904: 196). Odontolejeunea sagittistipula [Stephani (1890a: 19); Stephani (1890c: 140)]. Note:—The name seems never to have been validly published. Current name (Teeuwen 1989):—Odontolejeunea rhomalea (Spruce 1884: 147) Stephani (1912b: 176). Odontolejeunea sieberiana (Gottsche in Gottsche et al. 1845b: 328) [Stephani (1890c: 140); Stephani (1892e: 180)] Schiffner (1893b: 127). Current name (Vanden Berghen 1952):—Odontolejeunea lunulata (Weber 1815: 33) Schiffner (1893b: 128). Omphalolejeunea filiformis (Swartz 1788: 144) [Stephani (1888c: 285); Stephani (1890c: 140); Pearson (1892: 4); Stephani (1892e: 180)] Lacouture (1908: 105). Current name:—Omphalanthus filiformis (Swartz 1788: 144) Nees in Gottsche et al. (1845a: 304). Otigoniolejeunea schizotis (Spruce 1884: 227) Schiffner (1893b: 125). Current name (Reiner-Drehwald & Ilkiu-Borges 2007):— Otigoniolejeunea huctumalcensis (Lindenb. et Gottsche in Gottsche et al. 1847: 762) Wei et al. (2014: 237). Otigoniolejeunea xiphotis (Spruce 1884: 227) Schiffner (1893b: 125). Current name (Reiner-Drehwald & Ilkiu-Borges 2007):— Otigoniolejeunea huctumalcensis (Lindenb. et Gottsche in Gottsche et al. 1847: 762) Wei et al. (2014: 237). Peltolejeunea jackii [Steph. in Jack & Stephani (1892: 18)] (Prantl 1892: xvii) Schiffner (1893b: 131). Current name:—Omphalanthus jackii (Prantl 1892: xvii) Gradst. in Gradstein et al. (1977: 410). Peltolejeunea osculatiana (De Notaris 1855: 233) Schiffner (1893b: 131). Current name:—Lejeunea osculatiana De Notaris (1855: 233). Peltolejeunea ovalis (Lindenb. et Gottsche in Gottsche et al. 1847: 754) [Stephani (1890c: 140); Jack & Stephani (1892: 13)] Schiffner (1893b: 131). Current name:—Omphalanthus ovalis (Lindenb. et Gottsche in Gottsche et al. 1847: 754) Gradst. in Gradstein et al. (1977: 411). Peltolejeunea pulcherrima (Stephani 1886: 87) Schiffner (1893b: 131). Current name (Vanden Berghen 1963):—Lejeunea acuta Mitten 182 • Phytotaxa 220 (2) © 2015 Magnolia Press SÖDERSTRÖM ET AL. (1863: 167). Peltolejeunea wallisii [Jack & Stephani (1892: 19)] (Prantl 1892: xvii) Schiffner (1893b: 131). Current name:—Omphalanthus filiformis (Swartz 1788: 144) Nees in Gottsche et al. (1845a: 304) var. wallisii (Prantl 1892: xvii) Gradstein (2013b: 46). Phragmolejeunea polymorpha (Sande Lacoste 1854: 420) Schiffner [(1890: 25)]. Note:—The name seems never to have been validly published. Current name:—Spruceanthus polymorphus (Sande Lacoste 1854: 420) Verdoorn (1934b: 155). Platylejeunea baccifera (Taylor 1846: 387) [Stephani (1890c: 140)] Steph. ex Evans (1900: 417). Current name (Gradstein & van Beek 1985):—Symbiezidium transversale (Swartz 1788: 144) Trevisan (1877: 403) var. transversale. Platylejeunea barbiflora (Lindenberg & Gottsche 1851: 630) [Stephani (1888c: 282); Stephani (1890c: 140)] Schiffner (1897: 595). Current name:—Symbiezidium barbiflorum (Lindenberg & Gottsche 1851: 630) Evans (1907b: 540). Platylejeunea conferta (C.F.W.Meissn. ex Sprengel 1827: 325) [Stephani (1888c: 284)]. Note:—The name seems never to have been validly published. Current name:—Anoplolejeunea conferta (C.F.W.Meissn. ex Sprengel 1827: 325) Evans (1908a: 175). Platylejeunea granulata (Nees 1833: 352) [Stephani (1888c: 285); Stephani (1890c: 140)]. Schiffner (1897: 583). Current name (Gradstein & van Beek 1985):—Symbiezidium transversale (Swartz 1788: 144) Trevisan (1877: 403) var. transversale. Platylejeunea hobsoniana (Lindenb. in Gottsche et al. 1845b: 313) [Stephani (1890c: 140)] Schiffner (1893b: 131). Current name (Gradstein & van Beek 1985):—Symbiezidium transversale (Swartz 1788: 144) Trevisan (1877: 403) var. transversale. Platylejeunea setosa [Stephani (1890a: 14); Stephani (1890c: 140)]. Note:—The name seems never to have been validly published. Current name (Gradstein & van Beek 1985):—Symbiezidium barbiflorum (Lindenberg & Gottsche 1851: 630) Evans (1907b: 540). Platylejeunea subrotunda (Hook. in Kunth 1822: 41) [Stephani (1890c: 140)]. Schiffner (1897: 596). Current name (Uribe & Gradstein 1998):—Symbiezidium transversale (Swartz 1788: 144) Trevisan (1877: 403). Platylejeunea transversalis (Swartz 1788: 144) [Stephani (1888c: 292, 301); Stephani (1890c: 140)] Schiffner (1893b: 131). Current name:—Symbiezidium transversale (Swartz 1788: 144) Trevisan (1877: 403). Platylejeunea vincentina (Gottsche in Gottsche et al. 1845b: 313) [Stephani (1888c: 293); Stephani (1890c: 140)] Schiffner (1893b: 131). Current name (Gradstein & van Beek 1985):—Symbiezidium transversale (Swartz 1788: 144) Trevisan (1877: 403) var. hookerianum (Gottsche et al. 1845b: 311) Gradstein & van Beek (1985: 237). Prionolejeunea aemula (Gottsche in Gottsche et al. 1845b: 338) [Stephani (1890b: 69); Stephani (1890c: 140)] Evans (1904: 219). Prionolejeunea angulistipa [Stephani (1890b: 69); Stephani (1890c: 140)] Stephani (1896a: 118). Current name (Ilkiu-Borges 2006):— Prionolejeunea denticulata (Weber 1815: 30) Schiffner (1893b: 127). Prionolejeunea decora (Taylor 1846: 393) [Stephani (1890b: 69); Stephani (1890c: 140)] Stephani (1913: 207). Prionolejeunea denticulata (Weber 1815: 30) [Stephani (1888c: 285); Stephani (1890b: 68); Stephani (1890c: 140); Stephani (1892c: 171)] Schiffner (1893b: 127). Prionolejeunea deplanata Schiffner (1893b: 127). Current name (Grolle 1978):—Prionolejeunea grata (Gottsche 1882: 359) Schiffner (1893b: 127). Prionolejeunea fabroniifolia (Spruce 1884: 154) [Jack & Stephani (1892: 13) ‘fabroniaefolia’] Schiffner (1893b: 127) ‘fabroniaefolia’. Current name (Schultze-Motel & Menzel 1987):—Prionolejeunea schlimiana (Gottsche 1864: 154) Stephani (1913: 214). Prionolejeunea grata (Gottsche 1882) [Pearson (1892: 7)] Schiffner (1893b: 127). Prionolejeunea guadalupensis (Lindenb. in Gottsche et al. 1845b: 340) [Stephani (1890b: 70); Stephani (1890c: 140)] Stephani (1913: 211). Prionolejeunea kindbergii [Stephani 1892c: 171] (Prantl. 1892: xvii) Schiffner (1893b: 127). Current name (Grolle 1978):—Prionolejeunea grata (Gottsche 1882: 359) Schiffner (1893b: 127). Prionolejeunea meissneri (Gottsche in Gottsche et al. 1845b: 340) [Stephani (1890b: 69); Stephani (1890c: 140)] Stephani (1913: 212). Prionolejeunea microdonta (Gottsche in Gottsche et al. 1845b: 337) [Stephani (1890b: 69); Stephani (1890c: 140)] Stephani (1897a: 842). Current name (Ilkiu-Borges 2006):—Prionolejeunea denticulata (Weber 1815: 30) Schiffner (1893b: 127). Prionolejeunea rutenbergiana (Gottsche 1882: 358) [Pearson (1892: 7) ‘rutenbergii’] Stephani (1913: 201). Current name (Grolle 1978):—Prionolejeunea grata (Gottsche 1882: 359) Schiffner (1893b: 127). Prionolejeunea serrulata (Montagne 1842: 479) [Stephani (1890b: 69); Stephani (1890c: 140)] Stephani (1913: 224). Current name (Ilkiu-Borges 2006):—Prionolejeunea denticulata (Weber 1815: 30) Schiffner (1893b: 127). Ptycholejeunea birmensis [Stephani (1889b: 258)] Stephani (1896a: 120). Current name (Mizutani 1961):—Ptychanthus striatus (Lehm. et Lindenb. in Lehmann 1832: 16) Nees (1838: 212). Ptycholejeunea javanica (Nees in Gottsche et al. 1845a: 291) [Stephani (1890c: 140)]. Note:—The name seems never to have been validly published. Current name (Verdoorn 1933):—Ptychanthus striatus (Lehm. et Lindenb. in Lehmann 1832: 16) Nees (1838: 212). Ptycholejeunea perrottetii [Stephani (1889b: 259) ICN Art. 38.1(a); no description; Stephani (1890a: 5); Stephani (1890c: 140)] Stephani (1896a: 121). Current name (Mizutani 1961):—Ptychanthus striatus (Lehm. et Lindenb. in Lehmann 1832: 16) Nees (1838: 212). Ptycholejeunea pycnoclada (Taylor 1846: 385) [Stephani (1890c: 140)]. Note:—The name seems never to have been validly published. Current name:—Acrolejeunea pycnoclada (Taylor 1846: 385) Schiffner (1893b: 128). GENERA OF LEJEUNEACEAE ESTABLISHED IN 1884–1893 Phytotaxa 220 (2) © 2015 Magnolia Press • 183 Ptycholejeunea pyriformis [Stephani (1889b: 259) ICN Art. 38.1(a); no description; Bescherelle (1892: 14) ‘piriformis’ ICN Art. 38.1(a); no description] Stephani (1896a: 122). Current name (Awasthi & Srivastava 1987):—Ptychanthus striatus (Lehm. et Lindenb. in Lehmann 1832: 16) Nees (1838: 212). Ptycholejeunea recondita [Stephani (1889b: 259) ICN Art. 38.1(a); no description] Stephani (1896a: 122). Current name:—Mastigolejeunea recondita (Stephani 1896a: 122) Mizutani (1969: 134). Ptycholejeunea retusa (Reinwardt et al. 1824: 214) [Stephani (1890c: 140)]. Note:—The name seems never to have been validly published. Current name:—Thysananthus retusus (Reinwardt et al. 1824: 214) Thiers & Gradstein (1989: 67). Ptycholejeunea semirepanda (Nees 1830: 39) [Stephani (1890c: 140)]. Note:—The name seems never to have been validly published. Current name:—Spruceanthus semirepandus (Nees 1830: 39) Verdoorn (1934b: 153). Ptycholejeunea stephensoniana (Mitten 1852: 155) [Stephani (1889b: 258)]. Note:—The name seems never to have been validly published. Current name:—Ptychanthus stephensonianus (Mitten 1852: 155) Stephani (1912a: 754). Ptycholejeunea striata (Lehm. et Lindenb. in Lehmann 1832: 16) [Stephani (1890c: 140); Stephani (1891a: 203); Steph. ex Prantl (1891: xiii); Stephani (1892a: 120); Pearson (1892: 4)] Stephani (1895b: 65). Current name:—Ptychanthus striatus (Lehm. et Lindenb. in Lehmann 1832: 16) Nees (1838: 212). Ptycholejeunea sulcata (Nees 1830: 36) [Stephani (1890c: 140)]. Note:—The name seems never to have been validly published. Current name:—Spruceanthus sulcatus (Nees 1830: 36) Gradstein (1985: 26). Ptycholejeunea theobromae (Spruce 1884: 99) [Stephani (1889b: 259)]. Note:—The name seems never to have been validly published. Current name:—Spruceanthus theobromae (Spruce 1884: 99) Gradstein (1985: 26). Ptycholejeunea wightii (Gottsche in Gottsche et al. 1845a: 291) [Stephani (1890c: 140)]. Note:—The name seems never to have been validly published. Current name (Mizutani 1961):—Ptychanthus striatus (Lehm. et Lindenb. in Lehmann 1832: 16) Nees (1838: 212). Pycnolejeunea adnata (Kunze ex Lehmann 1834: 46) [Stephani (1890b: 81); Stephani (1890c: 140)] Stephani (1914: 612). Current name:—Cheilolejeunea adnata (Kunze ex Lehmann 1834: 46) Grolle (1977: 529). Pycnolejeunea bidentula [Stephani (1889b: 259)] Steph. in Jack & Stephani (1894: 107). Current name:—Lepidolejeunea bidentula (Steph. in Jack & Stephani 1894: 107) Schuster (1980b: 425). Pycnolejeunea callosa (Gottsche in Gottsche et al. 1845b: 359) [Stephani (1890b: 77); Stephani (1890c: 140)] Stephani (1914: 609). Current name (He 1999):—Pycnolejeunea macroloba (Nees et Mont. in Montagne 1843: 260) Schiffner (1893b: 124). Pycnolejeunea ceylanica (Gottsche in Gottsche et al. 1845a: 259) [Stephani (1889b: 259); Stephani (1890b: 77); Stephani (1890c: 140)] Schiffner (1893b: 124). Current name:—Cheilolejeunea ceylanica (Gottsche in Gottsche et al. 1845a: 259) R.M.Schust. et Kachroo in Kachroo & Schuster (1961: 509). Pycnolejeunea connivens [Schiffner (1890: 32)] Schiffn. ex Sydow (1894: 246). Pycnolejeunea curvatiloba [Stephani (1889b: 260)] Stephani (1914: 635). Current name (Thiers 1992):—Cheilolejeunea trapezia (Nees 1830: 41) Kachroo & Schuster (1961: 509). Pycnolejeunea densiuscula [Stephani (1890b: 86) ICN Art. 38.1(a); no description; Stephani (1890c: 141) ICN Art. 38.1(a); no description] Stephani (1914: 613). Current name (Grolle 1979a):—Pycnolejeunea contigua (Nees 1833: 360) Grolle (1979a: 179). Pycnolejeunea discoidea (Lehm. et Lindenb. in Lehmann 1834: 47) [Stephani (1890b: 86); Stephani (1890c: 141)] Stephani (1914: 612). Current name:—Cheilolejeunea discoidea (Lehm. et Lindenb. in Lehmann 1834: 47) R.M.Schust. et Kachroo in Kachroo & Schuster (1961: 509). Pycnolejeunea hyalina [Stephani (1890b: 77); Stephani (1890c: 141)] Stephani (1914: 614). Current name (see below):—Lejeunea hyalina (Stephani 1914: 614) L.Söderstr. et A.Hagborg, comb. nov. Pycnolejeunea imbricata (Nees 1830: 42) [Stephani (1890b: 77); Stephani (1890c: 141)] Schiffner (1893b: 124). Current name (Zhu & Grolle 2004):—Cheilolejeunea trapezia (Nees 1830: 41) Kachroo & Schuster (1961: 509). Pycnolejeunea incisa (Gottsche in Gottsche et al. 1845b: 360) [Stephani (1890b: 77); Stephani (1890c: 141)] Schiffner (1898: 259). Current name:—Cheilolejeunea incisa (Gottsche in Gottsche et al. 1845b: 360) R.M.Schust. et Kachroo in Kachroo & Schuster (1961: 509). Pycnolejeunea longidens [Stephani (1889b: 260)] Stephani (1914: 634). Current name:—Cheilolejeunea longidens (Stephani 1914: 634) R.M.Schust. et Kachroo in Kachroo & Schuster (1961: 509). Pycnolejeunea macroloba (Nees et Mont. in Montagne 1843: 260) [Stephani (1888c: 290); Stephani (1890a: 10); Stephani (1890b: 77); Stephani (1890c: 141)] Schiffner (1893b: 124). Pycnolejeunea meyeniana (Gottsche et al. 1843: 472) [Stephani (1890a: 11) also ICN Art. 41.1; basionym not valid; Stephani (1890b: 77); Stephani (1890c: 141); Bescherelle (1892: 14)] Stephani (1914: 628). Current name:—Cheilolejeunea meyeniana (Gottsche et al. 1843: 472) R.M.Schust. et Kachroo in Kachroo & Schuster (1961: 509). Pycnolejeunea schwaneckei [Stephani (1888c: 289); Schiffner (1890: 32)] Schiffn. ex Sydow (1894: 246). Pycnolejeunea silvestris (Gottsche 1882: 354) [Pearson (1892: 8)] Stephani (1914: 601). Current name (Jones 1976):—Cheilolejeunea surrepens (Mitten 1879: 399) Jones (1976: 49). 184 • Phytotaxa 220 (2) © 2015 Magnolia Press SÖDERSTRÖM ET AL. Pycnolejeunea trapezia (Nees 1830: 41) [Stephani (1890b: 76); Stephani (1890c: 141)] Schiffner (1893b: 124). Current name:— Cheilolejeunea trapezia (Nees 1830: 41) Kachroo & Schuster (1961: 509). Pycnolejeunea ventricosa [Schiffner (1890: 32)] Schiffn. ex Sydow (1894: 246). Current name:—Cheilolejeunea ventricosa (Schiffn.) He (1999: 60). Stictolejeunea kunzeana (Gottsche et al. 1845b: 322) [Stephani (1890c: 141)] Schiffner (1893b: 131). Current name (Gradstein 1985):— Stictolejeunea squamata (Willd. in Weber 1815: 33) Schiffner (1893b: 131). Stictolejeunea squamata (Willd. in Weber 1815: 33) [Stephani (1888c: 291); Stephani (1890c: 141)] Schiffner (1893b: 131). Strepsilejeunea acuminata (Lehm. et Lindenb. in Lehmann 1834: 49) [Stephani (1890b: 75); Stephani (1890c: 141)] Stephani (1913: 293). Current name:—Lejeunea acuminata (Lehm. et Lindenb. in Lehmann 1834: 49) Lehmann (1838: 22). Strepsilejeunea acutangula (Nees 1833: 357) [Stephani (1890b: 74); Stephani (1890c: 141)] Stephani (1913: 278). Current name:— Cheilolejeunea acutangula (Nees 1833: 357) Grolle (1979a: 173). Strepsilejeunea austrina [Stephani (1889b: 260)] Spruce ex Stephani (1913: 29). Current name (Grolle 1982):—Cheilolejeunea mimosa (Hooker & Taylor 1844a: 398) Schuster (1963a: 245). Strepsilejeunea brevifissa (Gottsche 1882: 356) [Pearson (1892: 7)] Stephani (1895b: 65). Current name (Jones 1988):—Cheilolejeunea krakakammae (Lindenb. in Gottsche et al. 1845b: 353) Schuster (1963a: 112). Strepsilejeunea cardiophylla (Gottsche 1882: 356) [Pearson (1892: 7)] Stephani (1913: 276). Current name (Grolle 1991):—Lejeunea conformis Nees et Mont. in Montagne (1843: 260). Strepsilejeunea comitans (Hook.f. et Taylor in Taylor 1846: 400) [Stephani (1890b: 79); Stephani (1890c: 141)] Stephani (1913: 291). Current name (Zhu 2006):—Cheilolejeunea krakakammae (Lindenb. in Gottsche et al. 1845b: 353) Schuster (1963a: 112). Strepsilejeunea inflexa (Hampe in Lehmann 1838: 22) [Stephani (1890b: 74); Stephani (1890c: 141); Jack & Stephani (1892: 13)] Stephani (1897b: 24). Current name:—Cheilolejeunea inflexa (Hampe in Lehmann 1838: 22) Grolle (1979a: 174). Strepsilejeunea involuta (Gottsche in Gottsche et al. 1845b: 350) [Stephani (1888c: 287); Stephani (1890b: 74); Stephani (1890c: 141)] Stephani (1896a: 76). Current name:—Lepidolejeunea involuta (Gottsche in Gottsche et al. 1845b: 350) Grolle (1984: 504). Strepsilejeunea krakakammae (Lindenb. in Gottsche et al. 1845b: 353) [Stephani (1890b: 74); Stephani (1890c: 141)] Stephani (1913: 276). Current name:—Cheilolejeunea krakakammae (Lindenb. in Gottsche et al. 1845b: 353) Schuster (1963a: 112). Strepsilejeunea kunthiana (Lindenb. in Gottsche et al. 1845b: 353) [Stephani (1890b: 74); Stephani (1890c: 141)] Schiffner (1893b: 127). Current name (Grolle 1979a):—Cheilolejeunea acutangula (Nees 1833: 357) Grolle (1979a: 173). Strepsilejeunea laevicalyx [Jack & Stephani (1892: 19)] J.B.Jack et Steph. in Stephani (1913: 283). Current name:—Cheilolejeunea laevicalyx (J.B.Jack et Steph. in Stephani 1913: 283) Grolle (1988: 403). Strepsilejeunea lanceolata (Gottsche in Gottsche et al. 1845b: 350) [Stephani (1890b: 75); Stephani (1890c: 141)] Stephani (1913: 283). Strepsilejeunea lobulata (Lindenb. in Gottsche et al. 1845b: 350) [Stephani (1890b: 74); Stephani (1890c: 141)] Stephani (1913: 284). Current name (Grolle 1979a):—Cheilolejeunea acutangula (Nees 1833: 357) Grolle (1979a: 173). Strepsilejeunea luchmannii [Stephani (1889b: 261)] Stephani (1913: 292). Current name (Grolle 1982):—Lejeunea drummondii Taylor (1846: 400). Strepsilejeunea mimosa (Hooker & Taylor 1844a: 398) [Stephani (1890b: 85); Stephani (1890c: 141)] Schiffner (1893b: 127). Current name:—Cheilolejeunea mimosa (Hooker & Taylor 1844a: 398) Schuster (1963a: 245). Strepsilejeunea neelgherriana (Gottsche in Gottsche et al. 1845b: 354) [Stephani (1890b: 75) ‘nilgerriana’; Stephani (1890c: 141) ‘nilgerriana’] Schiffner (1899: 206) ‘Nilgherriana’. Current name:—Lejeunea neelgherriana Gottsche in Gottsche et al. (1845b: 354). Strepsilejeunea obtruncata (Montagne 1845b: 354) [Stephani (1890b: 84); Stephani (1890c: 141)] Stephani (1913: 295). Current name:— Cheilolejeunea obtruncata (Montagne 1845b: 354) Solari (1983: 539). Strepsilejeunea orba (Gottsche in Gottsche et al. 1845b: 350) [Stephani (1890b: 74); Stephani (1890c: 141)] Schiffner (1893b: 127). Current name (Reiner-Drehwald 2006):—Lepidolejeunea involuta (Gottsche in Gottsche et al. 1845b: 350) Grolle (1984: 504). Strepsilejeunea owaihiensis (Gottsche in Gottsche et al. 1845b: 351) [Stephani (1890b: 74); Stephani (1890c: 141)] Stephani (1897a: 842) ‘oweihiensis’. Current name (Piippo 1986):—Lepidolejeunea involuta (Gottsche in Gottsche et al. 1845b: 350) Grolle (1984: 504). Strepsilejeunea tenuistipula Lindenb. et Gottsche in Gottsche et al. (1845b: 351) [Stephani (1890b: 74); Stephani (1890c: 141)]. Note:— The name seems never to have been validly published. Current name (He & Grolle 2001):—Xylolejeunea crenata (Nees et Mont. in Montagne 1838: 48) He & Grolle (2001: 36). Taxilejeunea affinis (Lindenb. et Gottsche in Gottsche et al. 1847: 748) Stephani (1890c: 141). Current name (Stephani 1914):—Lejeunea caracensis Lindenb. in Gottsche et al. (1845b: 355). Taxilejeunea antillana [Stephani (1888c: 281)] Stephani (1914: 482). Taxilejeunea apiculata (Gottsche 1864: 149) Jack & Stephani (1892: 13). GENERA OF LEJEUNEACEAE ESTABLISHED IN 1884–1893 Phytotaxa 220 (2) © 2015 Magnolia Press • 185 Taxilejeunea caripensis (Lindenb. et Gottsche in Gottsche et al. 1847: 758) Stephani (1890b: 75). Current name:—Lejeunea caripensis Lindenb. et Gottsche in Gottsche et al. (1847: 758). Taxilejeunea chamissonis (Lindenb. in Gottsche et al. 1845b: 378) Stephani (1890b: 85). Current name:—Lejeunea chamissonis Lindenb. in Gottsche et al. (1845b: 378). Taxilejeunea conformis (Nees et Mont. in Montagne 1843: 260) Stephani (1890b: 75). Current name:—Lejeunea conformis Nees et Mont. in Montagne (1843: 260). Taxilejeunea convexa Stephani (1889b: 262). Current name (Grolle 1979a):—Lejeunea microloba Taylor (1846: 399). Taxilejeunea cordistipula (Lindenb. et Gottsche in Gottsche et al. 1847: 758) Stephani (1890b: 75). Current name:—Lejeunea cordistipula Lindenb. et Gottsche in Gottsche et al. (1847: 758). Taxilejeunea debilis (Lehm. et Lindenb. in Lehmann 1832: 51) Stephani (1890c: 141). Current name:—Lejeunea debilis (Lehm. et Lindenb. in Lehmann 1832: 51) Nees & Montagne (1836: 60). Taxilejeunea eggersiana [Stephani (1888c: 285)] Schiffner (1897: 579). Taxilejeunea flaccida (Lindenb. et Gottsche in Gottsche et al. 1847: 758) Stephani (1890b: 75). Current name:—Lejeunea flaccida Lindenb. et Gottsche in Gottsche et al. (1847: 758). Taxilejeunea gracilipes (Taylor 1846: 385) Stephani (1890c: 141). Current name:—Lejeunea gracilipes (Taylor 1846: 385) Spruce (1884: 213). Taxilejeunea guadalupensis [Stephani (1890c: 141) ICN Art. 38.1(a); no description]. Note:—The name seems never to have been validly published. Current name (Stephani 1914):—Lejeunea lusoria (Lindenb. et Gottsche in Gottsche et al. 1847: 747) Stephani (1890c: 141). Taxilejeunea isocalycina (Nees 1833: 356) Stephani (1890c: 141). Current name (Costa 2009):—Taxilejeunea serpillifolioides (Raddi 1822: 17) Costa (2009: 230). Taxilejeunea leioscyphus (Gottsche 1864: 148) Stephani (1890c: 141) ‘leioscypha’. Current name (Gradstein & Hekking 1979):— Lejeunea pallescens Mitten (1851: 360). Taxilejeunea lumbricoides (Nees 1830: 40) Stephani (1890c: 141). Current name:—Lejeunea lumbricoides (Nees 1830: 40) Nees in Gottsche et al. (1845b: 342). Taxilejeunea lusoria (Lindenb. et Gottsche in Gottsche et al. 1847: 747) Stephani (1890c: 141). Current name:—Lejeunea lusoria (Lindenb. et Gottsche in Gottsche et al. 1847: 747) Stephani (1890c: 141). Taxilejeunea martinicensis (Gottsche 1864: 149) Stephani (1890c: 141). Current name (Evans 1921):—Lejeunea debilis (Lehm. et Lindenb. in Lehmann 1832: 51) Nees & Montagne (1836: 60). Taxilejeunea neumanniana (Nees in von Flotow et al. 1842: 44) Stephani (1890b: 75). Current name:—Lejeunea neumanniana Nees in von Flotow et al. (1842: 44). Taxilejeunea pterigonia (Lehm. et Lindenb. in Lehmann 1834: 44) Stephani (1890c: 142) ‘pterogonia’. Current name:—Lejeunea pterigonia (Lehm. et Lindenb. in Lehmann 1834: 44) Montagne (1840: 337). Taxilejeunea renistipula (Lindenb. in Gottsche et al. 1845b: 308) Stephani (1890c: 142). Taxilejeunea sulphurea (Lehm. et Lindenb. in Lehmann 1833: 14) [Stephani (1888c: 301, 292)] Stephani (1890c: 142). Current name:— Lejeunea sulphurea (Lehm. et Lindenb. in Lehmann 1833: 14) Spruce (1884: 217). Taxilejeunea surinamensis (Lindenberg & Gottsche 1851: 628) Stephani (1890c: 142). Taxilejeunea tenera (Swartz 1788: 143) Stephani (1890b: 98). Current name:—Lejeunea tenera (Swartz 1788: 143) Gottsche et al. (1845b: 406). Thysanolejeunea abietina [Stephani (1889b: 263)]. Note:—The name seems never to have been validly published. Current name (Verdoorn 1934b):—Thysananthus fruticosus (Lindenb. et Gottsche in Gottsche et al. 1847: 737) Schiffner (1893b: 130). Thysanolejeunea amazonica (Spruce 1884: 106) [Stephani (1889b: 263)]. Note:—The name seems never to have been validly published. Current name:—Thysananthus amazonicus Spruce (1884: 106). Thysanolejeunea anguiformis (Hooker & Taylor 1844c: 567) [Stephani (1890c: 142)]. Note:—The name seems never to have been validly published. Current name:—Thysananthus anguiformis (Hooker & Taylor 1844c: 567) Hook.f. et Taylor ex Gottsche et al. (1845a: 289). Thysanolejeunea comosa (Lindenb. in Lehmann 1844: 25) [Stephani (1890c: 142)]. Note:—The name seems never to have been validly published. Current name:—Thysananthus comosus Lindenb. in Lehmann (1844: 25). Thysanolejeunea convoluta (Lindenb. in Gottsche et al. 1845a: 288) [Stephani (1890c: 142)]. Note:—The name seems never to have been validly published. Current name:—Thysananthus convolutus Lindenb. in Gottsche et al. (1845a: 288). Thysanolejeunea cucullata [Stephani (1889b: 263)]. Note:—The name seems never to have been validly published and it is unclear to us where to refer it. Thysanolejeunea dissoptera (Spruce 1884: 108) [Stephani (1889b: 263); Stephani (1890c: 142)]. Note:—The name seems never to have been validly published. Current name (Gradstein & Hekking 1989):—Thysananthus comosus Lindenb. in Lehmann (1844: 25). 186 • Phytotaxa 220 (2) © 2015 Magnolia Press SÖDERSTRÖM ET AL. Thysanolejeunea fruticosa (Lindenb. et Gottsche in Gottsche et al. 1847: 737) [Stephani (1890c: 142)]. Note:—The name seems never to have been validly published. Current name:—Thysananthus fruticosus (Lindenb. et Gottsche in Gottsche et al. 1847: 737) Schiffner (1893b: 130). Thysanolejeunea gottschei Jack & Stephani (1892: 20). Current name:—Thysananthus gottschei (Jack & Stephani 1892: 20) Stephani (1912a: 787). Thysanolejeunea lehmanniana (Nees in Gottsche et al. 1845a: 302) [Stephani (1890c: 142)]. Note:—The name seems never to have been validly published. Current name (Geissler & Gradstein 1994):—Spruceanthus semirepandus (Nees 1830: 39) Verdoorn (1934b: 153). Thysanolejeunea plana (Sande Lacoste 1854: 419) [Stephani (1890c: 142)]. Stephani (1896b: 313). Current name (Thiers & Gradstein 1989):—Thysananthus retusus (Reinwardt et al. 1824: 214) Thiers & Gradstein (1989: 67). Thysanolejeunea pterobryoides (Spruce 1884: 109) [Stephani (1889b: 263)]. Stephani (1912a: 786). Current name:—Fulfordianthus pterobryoides (Spruce 1884: 109) Gradstein (1992a: 44). Thysanolejeunea reversa [Stephani (1889b: 263)] Stephani (1896a: 139). Current name (Sukkharak 2015):—Thysananthus gottschei (Jack & Stephani 1892: 20) Stephani (1912a: 787). Thysanolejeunea spathulistipa (Reinwardt et al. 1824: 212) [Stephani (1890c: 142)] Jack & Stephani (1894: 104). Current name:— Thysananthus spathulistipus (Reinwardt et al. 1824: 212) Lindenb. in Gottsche et al. (1845a: 287). Thysanolejeunea vittata (Mitten 1873: 411) [Stephani (1889b: 263)]. Note:—The name seems never to have been validly published. Current name (Schiffner 1898):—Thysananthus fruticosus (Lindenb. et Gottsche in Gottsche et al. 1847: 737) Schiffner (1893b: 130). Trachylejeunea acanthina (Spruce 1884: 182) Schiffner (1893b: 126). Current name:—Cheilolejeunea acanthina (Spruce 1884: 182) Gradstein & Ilkiu-Borges (2009: 62). Trachylejeunea ambigua (Lindenb. et Gottsche in Gottsche et al. 1847: 764) Stephani (1890b: 84). Current name:—Lejeunea ambigua Lindenb. et Gottsche in Gottsche et al. (1847: 764). Trachylejeunea asperiflora (Spruce 1884: 183) Schiffner (1893b: 126). Current name:—Cheilolejeunea asperiflora (Spruce 1884: 183) Gradstein & Ilkiu-Borges (2009: 62). Trachylejeunea elegantissima Stephani (1889b: 262). Current name:—Siphonolejeunea elegantissima (Stephani 1889b: 262) Grolle (1976b: 405). Trachylejeunea prionocalyx [Schiffner (1893b: 126) ICN Art. 38.1(a); no description] Schiffner (1897: 592). Current name (ReinerDrehwald & Goda 2000):—Lejeunea sporadica Bescherelle & Spruce (1889: clxxx). Trachylejeunea raddiana (Lindenb. in Gottsche et al. 1845b: 342) Stephani (1890b: 70). Current name:—Lejeunea raddiana Lindenb. in Gottsche et al. (1845b: 342). Nomenclatural consequences of establishing of Sprucean subgenera One consequence of the conclusions above is that the correct citations for many names, both generic and specific, need to be changed from citations commonly accepted in recent publications. In a couple of cases, where the name usually regarded as the basionym of the accepted name proves to be not validly published, the correct name turns out to be a name usually included in synonymy. A few taxa turn out to lack validly published names in the genus in which they usually are placed today. Below we only comment on the taxa where the correct name changes or where a new name or combination is needed. The basionym for Acrolejeunea securifolia subsp. hartmannii is Ptychocoleus hartmannii Stephani (1912b: 44). If recognized at species rank the correct name for the taxon would be Acrolejeunea micholitzii Stephani (1895a: 58). As Cololejeunea bistyla Stephani (1889a: 168) is not validly published, the correct name is Cololejeunea hamata Stephani (1895a: 248). Grolle (1975) considered Colurolejeunea apiculata Schiffner (1900: 198) not to be a validly published name, but we regard it as validly published as the generic name was validly published earlier. As it is an earlier name than Colurolejeunea leratii Stephani (1908: 30), Colura apiculata (Schiffner 1900: 198) Stephani (1916: 936) needs to replace Colura leratii (Stephani 1908: 30) Stephani (1916: 939). The name Lejeunea hyalina has been considered validly published in Stephani (1890a: 77) by e.g. Gradstein & Hekking (1979) and Uribe & Gradstein (1998). However, in that publication it is referred to the genus Pycnolejeunea, which at that time was not yet validly published. The earliest validly published name for the taxon is Pycnolejeunea hyalina (Stephani 1914: 614) and we cannot find it validly published in the genus Lejeunea. Thus we validate it here. GENERA OF LEJEUNEACEAE ESTABLISHED IN 1884–1893 Phytotaxa 220 (2) © 2015 Magnolia Press • 187 Lejeunea hyalina (Steph.) L.Söderstr. et A.Hagborg, comb. nov. Basionym:—Pycnolejeunea hyalina Steph., Sp. Hepat. [Stephani] 5: 614, 1914 (Stephani 1914). Type:—COLOMBIA. Bogotá, Lindig (G-00128273, lectotype here designated [if not holotype]). As Microlejeunea cochlearifolia Stephani (1888b: 113) is validly published, it must replace Microlejeunea kamerunensis Stephani (1915: 812), if the synonymy is correct (see Jones 1969 “perhaps synonymous?”). Odontolejeunea berteroana Stephani (1888c: 282) is not validly published and therefore is not available as a basionym. Rectolejeunea berteroana Evans (1906: 12), published with a description, is not a combination based on Stephani’s name, but a new species. However, because Evans cites Cheilolejeunea versifolia Schiffner (1897: 597) in synonymy, Evans name is illegitimate (ICN Art. 52.1) and a new combination in Rectolejeunea based on Schiffner’s name is required. Rectolejeunea versifolia (Schiffn.) L.Söderstr. et A.Hagborg comb. nov. Basionym:—Cheilolejeunea versifolia Schiffn., Bot. Jahrb. Syst. 23 (5): 597, 1897 (Schiffner 1897). Type:—CUBA. Wright 1202 (G), 131 (YU), s.n. (MANCH–12950), syntypes. = Rectolejeunea berteroana A.Evans, Bull. Torrey Bot. 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Taxon 31 (4): 718–722. http://dx.doi.org/10.2307/1219693 198 • Phytotaxa 220 (2) © 2015 Magnolia Press SÖDERSTRÖM ET AL.

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