Land of wood and water: A checklist of liverworts and hornworts of Jamaica

Lars Söderström; Anders Hagborg; Jiří Váňa; Matt von Konrat

The first ever liverwort and hornwort checklist is provided for Jamaica. We report 332 taxa in 314 species, 14 of which are invalid, illegitimate or in orphaned genera but we do not know where to place them. An additional 16 of the reported taxa are rejected as erroneously reported from Jamaica. The list is based on over 270 literature references, including monographs, regional studies, and molecular investigations. Except for a few early works in the 18th and early 19th century, and a scattering of other studies, the liverwort and hornwort flora of Jamaica has received negligible attention, despite its apparent diversity. The knowledge of the liverwort and hornwort flora, both in Jamaica and regionally in the Caribbean, remains so poor it is too early to reliably comment about range-restricted or endemic species. We predict that a number of new records remain to be reported, based on regionally widespread species that have been recorded elsewhere. On the other hand, it is envisaged that a number of the taxa presented here maybe synonyms of other taxa as they are taxonomically insufficiently known. As a consequence, Anastrophyllum harrisanum Steph. is reduced to a new synonym of A. tubulosum (Nees) Grolle.

Land of wood and water: A checklist of liverworts and hornworts of Jamaica Author(s): Lars Söderström, Anders Hagborg, Jiří Váňa and Matt von Konrat Source: The Bryologist, Vol. 114, No. 1 (Spring 2011), pp. 67-91 Published by: American Bryological and Lichenological Society Stable URL: http://www.jstor.org/stable/41289758 Accessed: 10-08-2016 20:31 UTC REFERENCES Linked references are available on JSTOR for this article: http://www.jstor.org/stable/41289758?seq=1&cid=pdf-reference#references_tab_contents You may need to log in to JSTOR to access the linked references. Your use of the JSTOR archive indicates your acceptance of the Terms & Conditions of Use, available at http://about.jstor.org/terms JSTOR is a not-for-profit service that helps scholars, researchers, and students discover, use, and build upon a wide range of content in a trusted digital archive. We use information technology and tools to increase productivity and facilitate new forms of scholarship. For more information about JSTOR, please contact support@jstor.org. American Bryological and Lichenological Society is collaborating with JSTOR to digitize, preserve and extend access to The Bryologist This content downloaded from 107.0.125.4 on Wed, 10 Aug 2016 20:31:07 UTC All use subject to http://about.jstor.org/terms Land of wood and water: A checklist of liverworts and hornworts of Jamaica Lars Soderstrom1,4, Anders Hagborg2, Jiri Vana3, and Matt von Konrat2 1 Department of Biology, Norwegian University of Science and Technology, N-7491 Trondheim, Norway; 2 Department of Botany, The Field Museum, 1400 South Lake Shore Drive, Chicago, IL 60605-2496, USA; 3 Department of Botany, Charles University, Benatska 2, CZ-128 01 Praha 2, Czech Republic Abstract. The first ever liverwort and hornwort checklist is provided for Jamaica. We report 332 taxa in 314 species, 14 of which are invalid, illegitimate or in orphaned genera but we do not know where to place them. An additional 16 of the reported taxa are rejected as erroneously reported from Jamaica. The list is based on over 270 literature references, including monographs, regional studies, and molecular investigations. Except for a few early works in the 18th and early 19th century, and a scattering of other studies, the liverwort and hornwort flora of Jamaica has received negligible attention, despite its apparent diversity. The knowledge of the liverwort and hornwort flora, both in Jamaica and regionally in the Caribbean, remains so poor it is too early to reliably comment about range-restricted or endemic species. We predict that a number of new records remain to be reported, based on regionally widespread species that have been recorded elsewhere. On the other hand, it is envisaged that a number of the taxa presented here maybe synonyms of other taxa as they are taxonomically insufficiently known. As a consequence, Anastrophyllum harrisanum Steph. is reduced to a new synonym of A. tubulosum (Nees) Grolle. Keywords. Jamaica, liverworts, hornworts, checklist. ♦ The Caribbean diverse forests global f t Islands, and prior supporting to the excep arriv ecosystems, ranging monta island was from completely to cactus scrublands, are one of thi 1995). Despite recent c biodiversityawareness, hotspots (Mittermeier forest clear Jamaica, originally "Xaymaca", an Arawa leading to species shift (aboriginal and 1 water," Caribbean. Indian) a is word meaning "land significant threat (C in the world, the regions third largest island in it Jamaica has flora the a land mass ofpo 1 remains very vascular flora. We checklist of liverworts 4 Corresponding author e-mail: lars.soderstrom@bio.ntnu.no an effort to further p DOI: 10.1639/0007-2745-114.1.67 bryological research in The Bryologist 114(1), pp. 67-91 0007-2745/1 1 /$2.65/0 Copyright ©2011 by The American Bryological and Lichenological Society, Inc. This content downloaded from 107.0.125.4 on Wed, 10 Aug 2016 20:31:07 UTC All use subject to http://about.jstor.org/terms 68 The Bryologist 114(l)' 2011 one recent collecting trip increased the number of checklists is summarized by Soderstrom et al. (2007, for Dominican Republic by over 120%, 2008), including outlining the utility of checklistsrecords as recording 258 taxa (Schafer-Verwimp & Pocs 2009). powerful and important tools, and their applicability to taxonomy, systematics, and conservation. We and Methods predict that a vast number of new records are yetMaterials to be reported for Jamaica, given that at least 50 other Sources include over 270 publications found liverwort and hornwort species occur on the through the work with Early Land Plants Today surrounding islands, or regionally, in Central and(ELPT) database of liverwort taxonomy and and with some consultation with South America. In this checklist we report 332 taxadistribution, in 314 species (out of which the occurrence of 3 is taxonomic experts. Some publications that only list names and mention Jamaica as types questioned) and 92 genera (out of which 3 genera published are either not longer recognized or the species here (Bonner 1962-1977; Crosby & Engel 2006; Geissler & 1985-1990) are not included in the checklist. included belongs to other genera but we are unableBischler to place them at present). Fourteen of those taxa are However, we cite all types and make an attempt to between holotypes, lectotypes, syntypes and invalid, illegitimate or in orphaned genera but we separate do paratypes. If only the word type is used we are not not know where to place them and 16 of the reported sure if it is a holotype or paratype but in the latter taxa are rejected as erroneously reported from Jamaica. The first apparent report of liverworts from case all paratypes should come from Jamaica. For invalid taxa that technically do not have any types, Jamaica, albeit only generic records, was by Browne (1756). Swartz (1788, 1806) provided the first we put the word type in quotation marks to indicate where the original material comes from. The significant work, with over 35 species, including the checklist distinguishes between records that are based description of many new species. It was not until the on specimens (reference in bold) and second-hand late 19th century and early 20th century that further reports, e.g., citation of the island (reference in influential works appeared (Boswell 1887; Pearson normal type) and we give the name used in the 1931). It is alarming, considering the liverwort publication. The four taxa in parenthesis are reported diversity, that with the exception of Schuster (1971), from Jamaica but the reports are doubtful although no one to our knowledge has investigated specifically the occurrence on the island is not unlikely. Taxa are Jamaican liverworts and hornworts since 1931! Even arranged in alphabetic order. Invalid and illegitimate these works that represent the most influential have all dealt with less than 40 taxa for the island. The names that we are not able to place in any synonymy are included, but no attempts have been made to seemingly most significant collections of the island, validate these names. Significantly, each accepted remain those of Swartz in 1783, Evans in 1903 and Schuster in 1967. Prior to the date of the current taxon is qualified using a four level ranking system checklist, Jamaican treatments have been confinedthat to indicates our level of knowledge about a taxon. The coding convention is outlined in detail by von references to old reports, broader monographic and taxonomic works, or distributional studies from Konrat et al. (2010). Briefly, ? = Problem with the taxon name; * = Serious doubts about the value of neighboring regions. Interestingly, Jamaica is the type locality for approximately 160 described taxa. the taxon; ** = Knowledge problem; *** = Accepted, a good taxon as currently understood. Schuster (1996) was the latest work to describe a new taxon from Jamaica, although that name remains CHECKLIST invalid. Reiner-Drehwald (2009) provided the most recent report for Jamaica, on Lejeunea adpressa Nees, Anthocerotophyta a widely distributed species of tropical America, but ** Anthoceros cavernosus Steph.: type of A. confuses , she only cites an earlier record from 1968. It remains Stephani 1898-1923 as A. confusus. very evident that the Caribbean, in general, remains** Anthoceros harrisanus (Steph.) Parihar: type of Aspirunder collected and much is yet to be discovered omitus harrisanus , Stephani 1898-1923 as Aspiromitus floristically, as evident by Schafer-Verwimp who in harrisanus. This content downloaded from 107.0.125.4 on Wed, 10 Aug 2016 20:31:07 UTC All use subject to http://about.jstor.org/terms Soderstrom et al.: Liverworts and hornworts of Jamaica 69 *** Anthoceros punctatus L.: type of A. monkeyanus, Stephani *** Bazzania aurescens Spruce: Pagan 1942 as B. quadricre- 1898-1923 as A. monkeyanus , Schafer-Verwimp & Pocs natay Fulford 1946, 1959 also as B. quadricrenata , Arnell 2009. 1959 as B. quadricrenata, Jovet-Ast 1960 as B. quadricrenata. *** Dendroceros crispus (Sw.) Nees: type of Anthoceros ** Bazzania cubensis (Gottsche ex Steph.) Pagan: Fulford crispus , Swartz 1788, 1806 both as A. crispus , Gottsche et 1946, 1963, Jovet-Ast 1960. al. 1844-1847, Pagan 1942, Miller et al. 1983 also as A. *** Bazzania cuneistipula (Gottsche & Lindenb.) Trevis.: type crispus. *** Leiosporoceros dussii (Steph.) Hassel: Stephani 1898- of Mastigobryum cuneistipulum , Boswell 1887 as M. cuneistipulumy Stephani 1898-1923 as M. tenerum and 1923 as Aspiromitus rigidus. M. cuneistipulumy Pearson 1931, Pagan 1942, Fulford 1946, 1963, Arnell 1956. ** Megaceros jamaicensis Steph.: type, Stephani 1898-1923. *** Bazzania denticulata (Lindenb. 8c Gottsche) Trevis.: Marchantiophyta Fulford 1946, 1963, Jovet-Ast 1960, Fulford & Sharp 1990. *** Acanthocoleus aberrans (Lindenb. & Gottsche) Kruijt: ** Bazzania eggersiana (Steph.) Pagan: Fulford 1963. Kruijt 1985 as Dicranolejeunea aberransy 1988, Fulford & *** Bazzania gracilis (Hampe 8c Gottsche) Steph.: LindenSharp 1990 as Dicranolejeunea aberrans. berg 8c Gottsche 1851 as Mastigobryum cuneifoliumy *** - var. aberrans: Gradstein 1994, Schafer-Verwimp 1999, Fulford 1946, 1963, Jovet-Ast 1949, Arnell 1956, Schafer- Zhu & Grolle 2001. Verwimp 8c Pocs 2009. *** - var. laevis Gradst.: Gradstein 1994. *** Bazzania hookeri (Lindenb.) Trevis.: Fulford 1946, 1959, *** Acrolejeunea emergens (Mitt.) Steph.: Gradstein 1994. 1963, 1987 also as B. latidens and B. breuteliana. *** Adelanthus pittieri (Steph.) Grolle: Fulford 8c Sharp 1990, *** Bazzania jamaicensis (Lehm. & Lindenb.) Trevis.: type of Schuster 2002 as Calyptrocolea pittieri. Herpetium jamaicense, Lehmann 1838 as H. jamaicensey *** Anastrophyllum piligerum (Nees) Steph.: Vana 1984, Gottsche et al. 1844-1847 as Mastigobryum jamaicensey 1991, Vana 8c Piippo 1989, Bapna 8c Kachroo 2000. Lindenberg 8c Gottsche 1851 as M. jamaicense , Boswell *** Anastrophyllum tubulosum (Nees) Grolle: type of A. 1887 as M. jamaicense, Stephani 1898-1923 as M. harrisanum , Stephani 1898-1923 as A. harrisanum , Vana jamaicensey Fulford 1946, 1959, 1963, Arnell 1956, Jovet1984. Note: Anastrophyllum harrisanum Steph. syn. nov. Ast 1960. ** Aneura bipinnata (Sw.) Nees: type of Jungermannia *** Bazzania longa (Nees) Trevis.: Pagan 1942, Fulford et al. bipinnatay Swartz 1788, 1806 both as /. bipinnata , Lunan 1970. Note: Bonner (1963) reports var. papillata (Steph.) 1814 as /. bipinnatay Weber 1815 as /. bipinnata, Fulford but this is a misreading of Fulford (1946, 1963). Gottsche et al. 1844-1847, Stephani 1898-1923. ** Anomoclada portoricensis (Hampe et Gottsche) Vana: Schuster 1972 as Odontoschisma portoricensis. * - var. longa: Fulford 1946, 1987. *** Bazzania longistipula (Lindenb.) Trevis.: type of Masti- *** Anoplolejeunea conferta (C. F. W.Meissn. ex Spreng.) A. Evans: type of Lejeunea proteoides , Lehmann 1838 as L. gobryum elegantulum and M. hanseniiy Gottsche 1864 as M. elegantulumy Stephani 1886 as M. elegantulum and M hanseniiy Pagan 1942, Fulford 1946, 1959, 1963, Arnell proteoideSy Evans 1908b, Pagan 1939, Thiers 1986, 1956. Schuster 1992c, Schafer-Verwimp 8c Pocs 2009. ** Bazzania pycnophylla (Taylor) Trevis.: type of Mastigo? Aphanolejeunea lancifera R. M. Schust.: type, Schuster bryum pycnophyllumy Taylor 1846 as M. pycnophyllum, 1980a, Schuster 1992c. Note: The genus Aphanolejeunea Gottsche et al. 1844-1847 as M. pycnophyllumy Linden- is now completely synonymized with Cololejeunea (e.g. berg 8c Gottsche 1851 as M. pycnophyllumy Stephani Pocs & Bernecker 2009) but they think this taxon might 1898-1923 as M. pycnophyllumy Fulford 1946, 1963, be similar or identical with Aphanolejeunea sintenisii Jovet-Ast 1960. (Steph.) Steph. and did not transfer it. *** Archilejeunea fuscescens (Hampe ex Lehm.) Fulford: *** Bazzania roraimensis (Steph.) Fulford: syntype of B. platystipuky Fulford 1941, 1946 both as B. platystipula , Lehmann 1838 as Lejeunea fuscescens. 1963, Fulford et al. 1970. *** Archilejeunea parviflora (Nees) Steph.: Pagan 1939 as A. *** Bazzania stolonifera (Sw.) Trevis.: type of Jungermannia viridissimay Jovet-Ast 1960 as A. viridissima, Gradstein & Buskes 1985, Thiers 1985, Gradstein 1994. *** Asterella elegans (Spreng.) Trevis.: Schuster 1992b, Bischler-Causse et al. 2005, Schafer-Verwimp 8c Pocs 2009. stoloniferay Swartz 1788, 1806 both as /. stoloniferay Lunan 1814 as /. stolonifera , Gottsche et al. 1844-1847 as Mastigobryum stoloniferumy Lindenberg 8c Gottsche 1851 as M. stoloniferumy Boswell 1887 as M. stoloniferum and M. vincentianumy Stephani 1898-1923 as M. stoloni- *** Bazzania affinis (Lindenb. 8c Gottsche) Trevis.: Forrest et al. 2006. ferum , Pagan 1939, 1942 as B. vincentinay Fulford 1946, 1959, 1963, 1987, Miller et al. 1983 also as Herpetium ** Bazzania armatistipula (Steph.) Fulford: type of Mastigobryum armatistipulumy Stephani 1898-1923 as M. armatistipulumy Fulford 1946, 1963. stoloniferumy Fulford 8c Sharp 1990, Piippo et al. 2002. ** Bazzania sublonga Fulford: holotype, Fulford 1959, 1963. This content downloaded from 107.0.125.4 on Wed, 10 Aug 2016 20:31:07 UTC All use subject to http://about.jstor.org/terms JO The Bryologist 114(1): 2011 *** Ceratolejeunea brevinervis (Spruce) A. Evans: Evans 1905, ** Bazzania wrightii (Gottsche ex Steph.) Steph.: Fulford 1946, 1963, 1987, Jovet-Ast 1960. Pagan 1939, 1942, Fulford 1945, 1987, Jovet-Ast 1960, Dauphin 2003. *** Brachiolejeunea laxifolia (Taylor) Schiffn.: Fulford 8c *** Ceratolejeunea coarina (Gottsche) Steph.: Fulford 1987 as Sharp 1990, Gradstein 1994, Schafer-Verwimp 8c Pocs C. plumula. 2009. *** Brachiolejeunea leiboldiana (Gottsche & Lindenb.) *** Ceratolejeunea cornuta (Lindenb.) Steph.: lectotype of Schiffn.: type of B. sulcata , Stephani 1898-1923 as B. Jungermannia cornuta, Boswell 1887 as Lejeunea cornuta, sulcata , Fulford 8c Sharp 1990, Gradstein 1994. Fulford 1945 also as C. maritima, 1987 also as C. variabilis, *** Brachiolejeunea phyllorhiza (Nees) Kruijt 8c Gradst.: Schuster 1980b, 1992c as C. schwaneckei , Lucking 1995 as C. maritima, Dauphin 2003, Zhu et al. 2005. Kruijt & Gradstein 1986, Fulford 8c Sharp 1990 as Dicranolejeunea phyllorhiza , Gradstein 1994, Schafer- *** Ceratolejeunea cubensis (Mont.) Schiffn.: Gottsche et al. 1844-1847 as Lejeunea cubensis, Stephani 1898-1923 as Verwimp 8c Pocs 2009. C. guadelupensis, Pagan 1942 as C. guadelupensis, Fulford *** Bryopteris diffusa (Sw.) Nees: syntype of Jungermannia 1945, Schuster 1980b, Fulford 8c Sharp 1990, Dauphin diffusa , Swartz 1788, 1806 both as /. diffusa , Lunan 1814 as /. diffusa, Weber 1815 as /. diffusa, Gottsche et al. 1844-1847, Miller et al. 1983, Yano 1987, Fulford 8c Sharp 1990, Reiner-Drehwald 1993, 1999, Gradstein 1994, Schafer-Verwimp 8c Pocs 2009. *** Bryopteris filicina (Sw.) Nees: type of Jungermannia filicina and syntype of B. filicina a hookeriana, Swartz 2003, Schafer-Verwimp 8c Pocs 2009. *** Ceratolejeunea fallax (Lehm. 8c Lindenb.) Bonner: Fulford 1987. *** Ceratolejeunea patentissima (Hampe 8c Gottsche) A. Evans: Pagan 1942, Fulford et al. 1970, Schuster 1992c. *** Ceratolejeunea spinosa (Gottsche) Steph.: Fulford 1945, 1987, Dauphin 2003. 1788, Swartz 1806 both as /. filicina , Lunan 1814 as /. ** Cheilolejeunea aciculifera R. M. Schust.: holotype, Schuster 1980a, 1992c. filicina , Weber 1815 as /. filicina , Nees 1830 as /. filicina , Gottsche et al. 1844-1847 as B. filicina p arguta and B. ** Cheilolejeunea acutangula (Nees) Grolle: Fulford 1987. filicina a hookeriana , Evans 1908a, Pagan 1939, Fulford ** Cheilolejeunea adnata (Kunze ex Lehm.) Grolle: Schuster 1987 as B. trinitensis , Fulford & Sharp 1990 also as B. fruticulosa and B. trinitensis , Reiner-Drehwald 1993, 1999, Gradstein 1994, Lucking 1995. 1992c. *** Cheilolejeunea clausa (Nees 8c Mont.) R. M. Schust.: Schuster 1980b. *** Calypogeia laxa Gottsche & Lindenb.: Fulford 1968, * Cheilolejeunea curnowii Steph.: type, Stephani 1895, 1987, Schuster 2000, Schafer-Verwimp 8c Pocs 2009. Stephani 1898-1923. Note: Neither E. Reiner-Drehwald *** Calypogeia peruviana Nees & Mont.: Evans 1907a as C. (pers. comm.), T. Pocs (pers. comm.) nor R.L. Zhu portoricensis , Stephani 1898-1923 as C. biapiculata , (pers. comm.) knows what it is and it is not mentioned Pagan 1939 as C. portoricensis , 1942 as C. biapiculata and in a manuscript of Cuban hepatics by Grolle given to C. portoricensis , Bischler 1962b also as C. heterophylla var. heterophylla, Fulford 1968, Schuster 1969, 1972, 2000 as C. heterophylla, Schafer-Verwimp & Pocs 2009. Pocs or in any notes he gave Reiner-Drehwald. *** Cheilolejeunea holostipa (Spruce) Grolle 8c R. L. Zhu: Schuster 1971, 1992c both as Cyrtolejeunea holostipa, *** Caudalejeunea lehmanniana (Gottsche) A. Evans: Thiers 1986 as Cyrtolejeunea holostipa, Grolle et al. 2002, Schuster 1980b, Gradstein 1994. Schafer-Verwimp 8c Pocs 2009. *** Cephalozia crassifolia (Lindenb. 8c Gottsche) Fulford: ** Cheilolejeunea jamaicensis Steph.: type, Stephani 1895, Fulford 1968, 1987 as C. caribbeania and C. subforficata, Fulford et al. 1970 as C. caribbeania , Schuster 2002 as C. 1898-1923, Pagan 1939, Thiers 1986. *** Cheilolejeunea lineata (Lehm. 8c Lindenb.) Schiffn.: subforficata, Schafer-Verwimp 8c Pocs 2009. Fulford 1987 as Cystolejeunea lineata. *** Cephaloziella antillana (Besch. 8c Spruce) Fulford: "type" ** Cheilolejeunea mammifera R. M. Schust.: holotype, of Evansia jamaicensis , Douin 1920 as E. jamaicensis, Schuster 1980a. Fulford 1976, Schuster 2002, Schafer-Verwimp & Pocs *** Cheilolejeunea oncophylla (Angstr.) Grolle 8c M. E. Reiner: 2009. Grolle & Reiner-Drehwald 1997, Reiner-Drehwald 1998, *** Cephaloziella granatensis (J.B.Jack ex Steph.) Fulford: Schafer-Verwimp 8c Pocs 2009, Schafer-Verwimp 8c "type" of Prionolobus evansii, Douin 1920 as P. evansii , Fulford 1976, Fulford 8c Sharp 1990, Nieuwkoop 8c Arts Reiner-Drehwald 2009. ? Cheilolejeunea papillifera R. M. Schust. nom. inval.: 1995, Grolle 8c Long 2000, Schuster 2002, Schafer- "syntype", Schuster 1992c. Note: Schuster (1992c) Verwimp & Pocs 2009. provided no Latin description describing the taxon. We ** Cephaloziella inaequalis R. M. Schust.: holotype, Schuster 1972, Fulford 1976. * Cephaloziella secundifolia Pearson: holotype, Pearson do not know where to refer it. *** ( Cheilolejeunea trifaria (Reinw., Blume 8c Nees) Mizut.: Schuster 1992c?). Note: Schuster (1992c) seems to be the 1931, Fulford 1976. Note: Fulford (1976) could not study only one reporting the taxon from Jamaica and he uses C. it and did not know what it is. cf. trifaria. Thus, we do not know of any confirmed report. This content downloaded from 107.0.125.4 on Wed, 10 Aug 2016 20:31:07 UTC All use subject to http://about.jstor.org/terms Soderstrom et al.: Liverworts and hornworts of Jamaica Jl *** Chiloscyphus breutelii (Gottsche) J. J. Engel & R. M. Schust.: Fulford 1976, 1987 both as Lophocolea trapezoideay Schafer-Verwimp 8c Pocs 2009. ** Cololejeunea papulosa (K. I. Goebel) Mizut.: "holotype", Eggers et al. 2004 as Aphanolejeunea angustissima. ? Cololejeunea papulosa R. M. Schust. nom. illeg .: type, *** Chiloscyphus coadunatus (Sw.) J. J. Engel 8c R. M. Schust.: Schusterl980a. Note: This is a later homonym of C. type of Jungermannia coadunata , Swartz 1806 as /. papulosa Tixier from Indochina. We do not know the coadunata , Weber 1815 as /. coadunata , Gottsche et al. value of Schuster's taxon. 1844-1847 as Lophocolea coadunata , Stephani 18981923 as L. coadunata , Pearson 1931 as L. coadunata , Pagan 1942 as L. coadunata , Fulford 1976 as I. coadunata , Engel 1978 as L. coadunata , Miller et al. 1983 as L. coadunata , Damsholt 2002. *** Chiloscyphus connatus (Sw.) J. J. Engel & R. M. Schust.: type of Jungermannia connata , Swartz 1788, 1806 both as /. connata , Lunan 1814 as /. connata , Weber 1815 as /. connata , Stephani 1898-1923 as Lophocolea connata , Pagan 1939, 1942 both as L. connata , Fulford 1976, 1987 both as I. connata , Fulford & Sharp 1990 as I. connata. *** Chiloscyphus guadeloupensis (Steph.) J. J. Engel & R. M. Schust.: Fulford 1976 as Lophocolea guadeloupensis. *** Chiloscyphus liebmannianus (Gottsche) J. J. Engel & R. M. Schust.: Fulford 1976, 1987 both as Lophocolea liebmannianay Fulford & Sharp 1990 as L. liebmanniana. *** Chiloscyphus martianus (Nees) J. J. Engel 8c R. M. Schust.: Fulford 1976 as Lophocolea martiana , Schuster 1980b as L. martiana subsp. martiana. *** Chiloscyphus muricatus (Lehm.) J. J. Engel 8c R. M. ** Cololejeunea parallelifolia R. M. Schust.: holotype, Schuster 1980a. *** Cololejeunea sicaefolia (Gottsche) Pocs et Bern.-Liick.: Jovet-Ast 1948 as Aphanolejeunea sicaefolia , Schuster 1992c as Aphanolejeunea sicaefolia. *** Cololejeunea sicaefolia subsp. jamaicensis (R. M. Schust.) Bern.-Liick. 8c Pocs: holotype of Aphanolejeunea jamaicensis, Schuster 1980a, 1992c both as A. jamaicensis. *** Cyclolejeunea accedens (Gottsche) A. Evans: lectotype of Phragmicoma affixa , Taylor 1847, 1850 both as P. affixa , Stephani 1898-1923 as C. affixa and Dicranolejeunea affixa , Fulford 1987, Kruijt 1988, Grolle 1991. *** Cyclolejeunea chitonia (Taylor ex Gottsche, Lindenb. 8c Nees) A. Evans: Pagan 1939. *** Cyclolejeunea convexistipa (Lehm. 8c Lindenb.) A. Evans: Boswell 1887 as Lejeunea mougeotii , Stephani 1890 as L. convexistipa , Grolle 1984, Schuster 1992c, Lucking 1995, Schafer-Verwimp 8c Pocs 2009. *** Cyclolejeunea luteola (Spruce) Grolle: Grolle 1984. Schust.: Fulford 1976 as Lophocolea muricata , Schuster *** Dicranolejeunea axillaris (Nees 8c Mont.) Schiffn.: Kruijt 1988, Gradstein 1994. 1992c, Schafer-Verwimp & Pocs 2009. *** Diplasiolejeunea brunnea Steph.: Reyes 1983, Tixier 1985 as *** Cololejeunea appressa (A. Evans) Benedix: holotype of Leptocolea appressa , Evans 1912b as L. appressa , Stephani D. cubensis, Lucking 1995, Schafer-Verwimp 8c Pocs 2009. *** Diplasiolejeunea cavifolia (Steph.) Steph.: Evans 1912a, 1898-1923 as L. appressa , Miller et al. 1983 as L. appressa , 1912b both as D. brachyclada , Pagan 1939 as D. Mizutani 1984, Fulford 8c Sharp 1990 as I. appressa , Zhu brachyclada , Jovet-Ast 1957, 1960 both as D. brachyclada , 1995, Bapna 8c Kachroo 2000, Zhu & So 2001, Eggers et al. 2001, Asthana 8c Srivastava 2003. Grolle 1966 as D. brachyclada , Bizot 8c Pocs 1974, Miller *** Cololejeunea camillii (Lehm.) A. Evans: Evans 1912c, Fulford 8c Sharp 1990, Eggers et al. 2004 as Aphanolejeunea camillii , Schafer-Verwimp & Pocs 2009, Schafer- Verwimp 8c Reiner-Drehwald 2009. *** Cololejeunea cardiocarpa (Mont.) A. Evans: Schuster 1980b, Miller et al. 1983, Schafer-Verwimp 8c Pocs 2009. *** Cololejeunea gracilis (Ast) Pocs: holotype of Aphanolejeunea gracilis var. linearifolia , Schuster 1980a as A. gracilis var. linearifolia , 1992c as A. gracilis , Eggers et al. 1998 as A. gracilis. et al. 1983, Reyes 1983, Fulford 8c Sharp 1990 as D. brachyclada , Schafer-Verwimp 8c Pocs 2009. *** Diplasiolejeunea inermis Tixier: holotype, Tixier 1985, Lucking 1995, Schafer-Verwimp 2004. *** Diplasiolejeunea johnsonii A. Evans: Reyes 1983, Gradstein et al. 1995, Schafer-Verwimp 2004, Schafer- Verwimp 8c Pocs 2009. ** - var. johnsonii: holotype, Evans 1912b, Fulford 8c Sharp 1990. *** Diplasiolejeunea pellucida (C. F. W.Meissn. ex Spreng.) Schiffn.: Grolle 1966, Reyes 1983, Fulford 1987, Fulford ** Cololejeunea microscopica (Taylor) Schiffn. var. africana (Pocs) Pocs et Bern.-Liick.: Schuster 1992c as Aphanolejeunea verrucosa. 8c Sharp 1990, Lucking 1995, Bapna 8c Kachroo 2000, Schafer-Verwimp 8c Pocs 2009. *** - var. malleiformis A. Evans: Evans 1912a, Pagan 1939, *** Cololejeunea minutissima (Sm.) Schiffn.: Miller et al. 1983. Fulford 1987. *** - subsp. myriocarpa (Nees 8c Mont.) R. M. Schust.: *** - var# pellucida: Evans 1912a, Pagan 1939, Fulford 1987. Evans 1911 as C. myriocarpa , Pagan 1939 as C. *** Diplasiolejeunea rudolphiana Steph.: Evans 1912a, Pagan myriocarpa. , Schuster 1980b, Fulford 8c Sharp 1990, 1939, Schuster 1980b, Reyes 1983, Fulford 1987, Zhu 8c Schafer-Verwimp 8c Pocs 2009. So 2001, Schafer-Verwimp 8c Pocs 2009. ** Cololejeunea obliqua (Nees & Mont.) Schiffn.: Evans 1911 as Leptocolea scabriflora , Pagan 1939 as L. scabri- *** Diplasiolejeunea unidentata (Lehm. 8c Lindenb.) Schiffn.: Evans 1912a, Pagan 1939, 1942, Jovet-Ast 1960, Fulford flora , Jovet-Ast 1948 as L. scabriflora , Fulford 8c Sharp 1987, Fulford et al. 1970, Fulford 8c Sharp 1990, Schafer- 1990 as L. scabriflora , Lucking 1995. Verwimp 8c Pocs 2009. This content downloaded from 107.0.125.4 on Wed, 10 Aug 2016 20:31:07 UTC All use subject to http://about.jstor.org/terms 72 The Bryologist 114(l). 2011 swartzii, Clark & Svihla 1945, 1948a, Miller et al. 1983, *** Diplophyllum obtusatum (R. M. Schust.) R. M. Schust.: Gradstein & Vana 1987, Engel 8c Merrill 1998, Schafer- Fulford 1987, Fulford 8c Sharp 1990, Uribe 8c Gradstein Verwimp 8c Pocs 2009. *** Drepanolejeunea anoplantha (Spruce) Steph.: Bischler 1964, Fulford et al. 1970, Schuster 1992c, Schafer- 2003, Schafer-Verwimp 8c Pocs 2009. *** Frullania brasiliensis Raddi: Lehmann 1838 as F. tenuis, Gottsche et al. 1844-1847, Boswell 1887, Pearson 1931, Clark 8c Svihla 1944b, Stotler 1970. Verwimp 8c Pocs 2009. ? - var. pseudocrassiretis R. M. Schust. nom. inval. : "holotype", Schuster 1996. Note: Schuster (1996) did ** Frullania brevipinna Steph.: type, Stephani 1898-1923. *** Frullania caulisequa (Nees) Nees: Clark & Svihla 1948b not indicate any herbarium when describing the variety. as F. gymnotis, Clark 1958 as F. gymnotis, Jovet-Ast 1960 We do not know the value of the taxon. *** Drepanolejeunea bidens (Steph.) A. Evans: Fulford & Sharp 1990. as F. gymnotis, Schuster 1992a as F. gymnotis. ** Frullania ecuadorensis Steph.: Stotler 1970. *** Frullania ericoides (Nees) Mont.: Hampe 1852, Miller et al. 1983. *** Drepanolejeunea biocellata A. Evans: Bischler 1964, Winkler 1968, Fulford & Sharp 1990. ** Frullania flexicaulis Spruce: Stotler 1970, Fulford 8c *** Drepanolejeunea crucianella (Taylor) A.Evans: Jovet-Ast 1948, Bischler 1964. Sharp 1990. *** Frullania gibbosa Nees: syntype, Gottsche et al. 18441847, Boswell 1887, Frye & Clark 1947, Clark 8c Svihla *** Drepanolejeunea fragilis Bischl.: Fulford et al. 1970. ** Drepanolejeuneahamulata Gottsche ex Steph.: Schuster 1948a, Clark 1956, Arnell 1959, Fulford 8c Sharp 1990, 1992c. Yuzawa 1991, Schuster 1992a, Schafer-Verwimp 8c Pocs 2009. *** ( Drepanolejeunea inchoata (C. F. W.Meissn. ex Lehm. & Lindenb.) Schiffn.: Winkler 1968, Miller et al. 1983, *** Frullania intumescens (Lehm. 8c Lindenb.) Lehm. 8c Fulford 1987. Note: Bischler (1964) questions the Lindenb.: type of /. intumescens, Lehmann 1834 as /. occurrence on Jamaica and we have not found any report intumescens, Gottsche et al. 1844-1847, Clark & Svihla confirming it. 1952, Stotler 1970, Fulford 8c Sharp 1990, Schuster ** - var. inchoata : Bischler 1964?). *** Drepanolejeunea lichenicola (Spruce) Steph.: Bischler 1992a. *** Frullania kunzei (Lehm. 8c Lindenb.) Lehm. 8c Lindenb.: 1964, Winkler 1968, Fulford 8c Sharp 1990, Lucking Stotler 1970 as F. neesii, Fulford 1987 as F. neesii, Fulford 1995, Schafer-Verwimp 8c Pocs 2009. 8c Sharp 1990 as F. neesii, Schafer-Verwimp 8c Pocs 2009. *** Drepanolejeunea mosenii (Steph.) Bischl.: Bischler 1968, ** Frullania macrocephala (Lehm. 8c Lindenb.) Lehm. 8c Lindenb.: Stotler 1970, Schafer-Verwimp 8c Pocs 2009. Lucking 1995, Schafer-Verwimp 8c Pocs 2009. *** Drepanolejeunea orthophylla (Nees 8c Mont.) Bischl.: Bischler 1968, Fulford 8c Sharp 1990. ** Frullania moritziana Lindenb. 8c Gottsche: Stotler 1970. *** Frullania riojaneirensis (Raddi) Angstr.: type of Junger- mannia obscura, Swartz 1806 as /. obscura, Weber 1815 *** Drepanolejeunea trigonophylla Steph.: Bischler 1964, as /. obscura, Nees 1830 as /. obscura, Yuzawa 1991. Winkler 1968, Fulford 1987. *** Dumortiera hirsuta (Sw.) Nees: Swartz 1788, 1806 both ** Frullania subtilissima (Nees ex Mont.) Lindenb.: Clark 8c Svihla 1948a as F. atrata var. subtilissima. as Marchantia hirsuta, Lunan 1814 as M. hirsuta. , Weber 1815 as M. hirsuta, Nees 1830, Gottsche et al. 1844- *** Frullanoides bahamensis (A. Evans) Van Slageren: 1847, Hitchcock 1898, Easa 2003, Bischler-Causse et al. syntype of Brachiolejeunea longispica, Stephani 1898- 2005, Schafer-Verwimp 8c Pocs 2009. 1923 as B. longispica. ** - subsp. hirsuta: lectotype of Marchantia hirsuta, Evans *** Frullanoides corticalis (Lehm. 8c Lindenb.) Van Slageren: 1919, Haynes et al. 1923, Grolle 1976, Miller et al. 1983, Schuster 1992b, Perold 1999. type of Jungermannia corticalis, Lehmann 1832 as J. corticalis, Gottsche et al. 1844-1847 as Phragmicoma ** - subsp. nepalensis (Taylor) R. M. Schust.: Evans 1919 corticalis, Boswell 1887 as Lejeunea corticalis, Stephani as D. nepalensis , Haynes et al. 1923 as D. nepalensis , 1890 as P. corticalis, Frye 8c Clark 1947 as Brachiolejeu- Schuster 1992b. Note: The subspecies is often treated as a nea corticalis, Schuster 1980b as B. corticalis, Fulford 8c synonym of the very variable D. hirsuta but recent studies Sharp 1990, Gradstein 1994, Reiner-Drehwald 1999, (unpubl.) indicate that more than one taxon is involved. Schafer-Verwimp 8c Pocs 2009. *** Frullania arecae (Spreng.) Gottsche: Clark & Svihla *** Frullanoides liebmaniana (Lindenb. 8c Gottsche) Van 1947, 1948a, Fulford 8c Sharp 1990, Schafer-Verwimp 8c Slageren: Evans 1908b as Brachiolejeunea insularis, Pocs 2009. Stephani 1898-1923 as B. insularis, Pagan 1939 as B. *** - var. arecae. Boswell 1887 as F. hians , Yuzawa 1991. insularis, Fulford 8c Sharp 1990, Gradstein 1994, Bapna *** - var. spiniloba (Steph.) Yuzawa: Yuzawa 1991. 8c Kachroo 2000. *** Frullanoides tristis (Steph.) Van Slageren: Fulford 8c *** Frullania atrata (Sw.) Nees: holotype of Jungermannia atrata, "type" of F. atrata a swartzii , Swartz 1788, 1806 Sharp 1990, Gradstein 1994. both as /. atrata, Lunan 1814 as /. atrata, Weber 1815 as ? Harpalejeunea reflexa R. M. Schust. nom. inval. : "type", /. atrata, Gottsche et al. 1844-1847 as F. atrata a Schuster 1992c, Grolle 8c Reiner-Drehwald 1999. Note: This content downloaded from 107.0.125.4 on Wed, 10 Aug 2016 20:31:07 UTC All use subject to http://about.jstor.org/terms Soderstrom et al.: Liverworts and hornworts of Jamaica 73 Schuster (1992c) failed to provide a Latin diagnosis when piligera var. jamaicensis, Guerke 1978, Crandall-Stotler describing the species and Grolle & Reiner-Drehwald & Guerke 1980, Fulford 8c Sharp 1990, Schuster 1992a, (1999) could not study it. Eggers et al. 2004. *** Harpalejeunea reflexula A. Evans: holotype, Evans 1908c, Stephani 1898-1923, Thiers 1986, Grolle & ReinerDrehwald 1999. ? Jungermannia aberrans (3 erosula Gottsche, Lindenb. & Nees: type, Gottsche et al. 1844-1847. Note: The species {Andrewsianthus aberrans (Nees & Mont.) Grolle) occurs *** Harpalejeunea stricta (Lindenb. & Gottsche) Steph.: Grolle & Reiner-Drehwald 1999, Schafer-Verwimp & Pocs 2009. in Madagascar, Reunion and Mauritius. It is not likely that this taxon is the same. ? Jungermannia exigua Austin nom. illeg .: type, Austin *** Harpalejeunea tridens (Besch. 8c Spruce) Steph.: Schuster1876. Note: Austin indicates that it may be a Lophocolea 1992c as H. heterodonta. *** Herbertus juniperoideus (Sw.) Grolle: Swartz 1788, 1806 both as Jungermannia juniperina, Lunan 1814 as /. but the description is too vague to tell what it is. Further, it is a later homonym of /. exigua Taylor. *** Kurzia capillaris (Sw.) Grolle: Swartz 1788, 1806 both as juniperoidea , Weber 1815 as /. juniperoidea , Gottsche et Jungermannia capillaris, Lunan 1814 as /. capillaris, al. 1844-1847 as Sendtnera juniperina , Boswell 1887 as Weber 1815 as /. capillaris, Lindenberg 8c Gottsche 1846 Sendtnera adunca , Stephani 1898-1923 as Schisma as Lepidozia capillaris, Boswell 1887 as L. capillaris, juniperinum , Pagan 1939 as H. juniperina , Pagan 1942 as Stephani 1898-1923 as L. capillaris, Pagan 1939 as L. H. juniperina, Arnell 1956 as H. juniperina , Fulford 1963, capillaris, Fulford 8c Sharp 1990, Schafer-Verwimp & Van Reenen 1982, Miller et al. 1983 also as H. Pocs 2009. juniperinus and Sendtnera juniperina, Fulford 8c Sharp 1990. *** - var> capillaris: type of Jungermannia capillaris, Fulford 1966 as Microlepidozia capillaris , 1987 as K. capillaris. *** - subsp. juniperoideus : type of Jungermannia juniperoidea , "type" of Sendtnera juniperina a swartziana , Schafer-Verwimp & Pocs 2009. *** - subsp. pensilis (Taylor) K. Feldberg & Heinrichs: Fulford 1987 as H. pensilis , Fulford & Sharp 1990 as H. *** - Vflr> verrucosa (Steph.) Pocs: Stephani 1898-1923 as Lepidozia verrucosa, Arnell 1956 as I. verrucosa, Fulford 1966 as Microlepidozia verrucosa, Bizot 8c Pocs 1974 as K. verrucosa, Fulford 1987 as K. verrucosa. *** Leiolejeunea grandiflora A. Evans: type, Evans 1908c, Stephani 1898-1923, Thiers 1986, Gradstein et al. 2001. pensilis . ** Heteroscyphus combinatus (Nees) Schiffn.: Fulford 1976, Fulford & Sharp 1990. ** Heteroscyphus elliottii (Steph.) Pagan: Fulford 1987, Schafer-Verwimp & Reiner-Drehwald 2009. ** Heteroscyphus miradorensis (Steph.) Schiffn.: Fulford 8c *** Leiomitra tomentosa (Sw.) Lindb.: type of Jungermannia tomentosa and "type" of /. swarzii, Swartz 1788, 1806 both as J. tomentosa, Raeuschel 1797 as /. swarzii, Lunan 1814 as /. tomentosa, Weber 1815 as /. tomentosa, Boswell 1887 as Trichocolea tomentella p tomentosa, Stephani 1898-1923 as T. tomentosa, Pearson 1931 as T. Sharp 1990. *** Isotachis multiceps (Lindenb. 8c Gottsche) Gottsche: Hatcher 1960, Fulford 1963, Fulford & Sharp 1990. *** Isotachis serrulata (Sw.) Gottsche: type of I. urbanii , tomentosa, Pagan 1939 as T. tomentosa, Hatcher 1958 as Trichocolea tomentosa, Fulford 1963, 1987 both as T. tomentosa, Engel 1978 as T. tomentosa, Fulford & Sharp Jungermannia serrulata var.a swartziana and /. serrulata , 1990 as T. tomentosa, Schafer-Verwimp & Pocs 2009 as Swartz 1788, 1806 both as /. serrulata, Lunan 1814 as J. T. tomentosa. serrulata, Weber 1815 as /. serrulata, Gottsche et al. 1844-1847 as /. serrulata var.a swartziana , Boswell 1887, Stephani 1898-1923 as I. urbanii, Hatcher 1960, Fulford 1963, 1987, Rico & Pocs 2004. *** Jubula pennsylvanica (Steph.) A. Evans: Ahonen2006. Note: Patsch et al. (2010) studied the /. pennsylvanica / hutchinsiae complex and concluded that it was only one species with 5 subspecies (out of which one didn't seem *** Lejeunea adpressa Nees: Schafer-Verwimp & Pocs 2009, Reiner-Drehwald 2009. ** Lejeunea androgyna R. M. Schust.: holotype, Schuster 1980a, 1992c, Reiner-Drehwald 1999. *** Lejeunea asperrima Spruce: Fulford 8c Sharp 1990 as Echinocolea asperrima, Ilkiu-Borges 2005. ? Lejeunea capillaris subsp. antillana R. M. Schust.: holotype, Schuster 1980a, Reiner-Drehwald 1999. Note: to have a name). They did not draw the full conclusion Lejeunea capillaris belongs to Microlejeunea but the out of their study and make the necessary nomenclatural variety was never transferred. We do not know the value changes. We prefer to keep /. pennsylvanica as a species with two subspecies, separated from /. huchinsiae with 2- of the taxon. *** Lejeunea cerina (Lehm. 8c Lindenb.) Gottsche, Lindenb. 3 subspecies. The complex is only represented on Jamaica & Nees: Fulford 1987 as Hygrolejeunea cerina, Schafer- by subsp. bogotensis and all reports of /. pennsylvanica are Verwimp 8c Pocs 2009. sensu lato. ** Lejeunea cyanomontana R. M. Schust.: holotype, Schus- *** - subsp. bogotensis (Steph.) W. R. Guerke: type of Frullania jamaicensis and /. piligera var. jamaicensis, Stephani 1898-1923 as F. jamaicensis, Pearson 1931 as /. ter 1980a, 1992c, Reiner-Drehwald 1999. *** Lejeunea flava (Sw.) Nees: Swartz 1788, 1806 both as Jungermannia flava, Lunan 1814 as /. flava, Weber 1815 This content downloaded from 107.0.125.4 on Wed, 10 Aug 2016 20:31:07 UTC All use subject to http://about.jstor.org/terms 74 The Bryologist 114(1): 2011 as J. flava, Gottsche et al. 1844-1847, Boswell 1887, Frye *** Lepidozia jamaicensis Steph.: type, Stephani 1898-1923, Fulford 1966. & Clark 1947, Miller et al. 1983 also as /. flava, Fulford 1987, Schuster 1992c, Lucking 1995, Reiner-Drehwald *** Lepidozia patens Lindenb.: Fulford 1966, 1987, Fulford et 1999, Bapna 8c Kachroo 2000, Zhu & So 2001, SchaferVerwimp &Pocs 2009. al. 1970, Schafer-Verwimp 8c Pocs 2009. *** Leptolejeunea elliptica (Lehm. 8c Lindenb.) Schiffn.: Bischler 1969, Schuster 1980b, Miller et al. 1983, ** - subsp. flava : lectotype of /. flava, Grolle 1976, Schuster 1980b, Fulford & Sharp 1990, Reiner-Drehwald Schuster 1984, Lucking 1995, Schafer-Verwimp 8c Pocs 2000. 2009. ** Lejeunea glaucescens Gottsche: Fulford 1987. *** Leptolejeunea exocellata (Spruce) A. Evans: Bischler 1969, Fulford 8c Sharp 1990, Schafer-Verwimp 8c Pocs *** Lejeunea laetevirens Nees 8c Mont.: Boswell 1887 as I. lucens, Bischler et al. 1962, Schuster 1962, 1980b, 2009. * Leptolejeunea jamaicensis R. M. Schust.: holotype, Fulford 1987, Lucking 1995, Schafer-Verwimp 8c Pocs Schuster 1967. Note: This taxon does not seem to have 2009. been mentioned anywhere since it was described. *** Lejeunea maxonii (A. Evans) Xiao L. He: holotype of Rectolejeunea maxonii, Evans 1918, 1912b both as R. *** Leptolejeunea radicosa (Nees 8c Mont, ex Mont.) Grolle: holotype of L. obovata, Bischler 1969 as L. obovata, maxoniiy Pagan 1939 as R. maxonii , Frye 8c Clark 1947 as R. maxonii, Jovet-Ast 1960 as R. maxonii , Schuster 1980b Dauphin et al. 1998, Eggers et al. 2004, Schafer-Verwimp as R. maxonii, He 1997, Reiner-Drehwald 1999, Yano 8c Reiner-Drehwald 2009. 2004. *** Leptoscyphus cuneifolius (Hook.) Mitt.: Schuster 1980b, Fulford & Sharp 1990. * Lejeunea oerstediana Lindenb. 8c Hampe ex Hampe: type, Hampe 1852, Lehmann 1857, Stephani 1898-1923 as *** Leptoscyphus ovatus (Spruce) Grolle: Grolle 1963, Fulford 1976, 1987. Euosmolejeunea oerstediana, Reiner-Drehwald 1999 as E. oerstediana. Note: A poorly known taxon often placed in *** Leucolejeunea xanthocarpa (Lehm. 8c Lindenb.) A. Evans: Evans 1908b, Winkler 1968, Schuster 1980b, Miller et al. Euosmolejeunea ( = Cheilolejeunea). We can not tell where 1983, Fulford 1987, Schafer-Verwimp 8c Pocs 2009. it belongs until the type is re-studied. ** Lejeunea phyllobola Nees 8c Mont.: Boswell 1887. ? Lejeunea ruthii R. M. Schust. subsp. jamaicensis R. M. *** Lopholejeunea eulopha (Taylor) Schiffn.: Pagan 1939 as L. howei, Jovet-Ast 1960 as I. howei. Schust. nom. inval:. Schuster 1992c. Note: Lejeunea *** Lopholejeunea nigricans (Lindenb.) Schiffn.: Evans 1907b ruthii (A. Evans) R. M. Schust. is treated as Microlejeunea as I. muelleriana, Pagan 1939 as I. muelleriana, Jovet-Ast ruthii A. Evans here but the subspecies is neither 1948 as L. muelleriana, Schuster 1980b as L. muelleriana, transferred, nor valid as Schuster (1992c) failed to give a Gradstein 1994, Lucking 1995. Latin diagnosis when describing it. *** Lejeunea subspathulata Spruce: holotype of Trachylejeu- *** Lopholejeunea subfusca (Nees) Schiffn.: Schuster 1980b, Gradstein 1994, Zhu 8c Gradstein 2005, Schafer- nea dilatata, Evans 1908c as T. dilatata, Stephani 1898- Verwimp 8c Pocs 2009. 1923 as T. dilatata, Jovet-Ast 1948 as T. dilatata, Thiers * Lophozia jamaicensis (Nees) Steph.: type of Jungerman- 1986 as Echinocolea dilatata, Fulford 1987 as T. dilatata, nia jamaicensis, Gottsche et al. 1844-1847 as J. Lucking 1995 as E. dilatata , Ilkiu-Borges 2005. jamaicensis , Stephani 1898-1923. Note: Stephani transferred Jungermannia jamaicensis Nees to Lophozia with a ** Lejeunea trinitensis Lindenb.: Evans 1940 as Stylolejeunea pililoba, Schuster 1962 as Lejeunea pililoba , Fulford 1987 ? stating that it is missing in Nees's herbarium and may as S. trinitensis. be an Anastrophyllum but the description does not fit Anastrophyllum. *** Lepidolejeunea involuta (Gottsche) Grolle: type of Ceratolejeunea mollis, Pearson 1931 as C. mollis, Piippo *** Lunularia cruciata (L.) Dumort. ex Lindb.: Haynes et al. 1923, Bischler-Causse et al. 2005. 1986, Schuster 1992c as I. punctata, Dauphin 2003 as C. mollis , Schafer-Verwimp 8c Pocs 2009. *** Marchantia breviloba A. Evans: holotype, Evans 1917, Haynes et al. 1923, Bischler-Causse et al. 2005. *** Lepidozia armata Steph.: Fulford 1966, Fulford 8c Sharp 1990. *** Lepidozia cupressina (Sw.) Lindenb.: lectotype of Jun- *** Marchantia chenopoda L.: type of M. swartzii and M. dillenii, Lehmann 1832 as M. swartzii , Gottsche et al. and /. cupressina a swartziana, Swartz 1788, 1806 both as 1844-1847, Hampe 1852, Lindberg 1883 as M. dillenii, Hitchcock 1898, Evans 1917, Miller et al. 1983, Bischler- /. cupressina, Lunan 1814 as /. cupressina, Weber 1815 as Causse et al. 2005. germannia cupressina, "type" of J. cupressina a* major, J. cupressina, Lehmann 1832 as J. cupressina, J. cupressina *** Marchantia inflexa Nees et Mont.: Stephani 1898-1923 as M disjuncta, Evans 1910a as M. disjuncta, Schuster a swartziana and /. cupressina a* major, Gottsche et al. 1844-1847, Lindenberg 8c Gottsche 1846, Boswell 1887, 1992b as M. papillata subsp. inflexa, Bischler-Causse et Fulford 1966, Engel 1978, Fulford 8c Sharp 1990, al. 2005. Damsholt 2002. *** Marchantia paleacea Bertol.: Evans 1917, Haynes et al. ** - subsp. cupressina: type of J. cupressina, Grolle 1976. 1923, Schafer-Verwimp 8c Pocs 2009. This content downloaded from 107.0.125.4 on Wed, 10 Aug 2016 20:31:07 UTC All use subject to http://about.jstor.org/terms Soderstrom et al.: Liverworts and homworts of Jamaica 75 * - var. jamaicense Reimers: holotype, Reimers 1933, *** - subsp. paleacea : Evans 1917 as M. domingensis , Schuster 1992b, Bischler-Causse et al. 2005. Jovet-Ast 1960. Note: Reimers (1933) split Micropterygium trachyphyllum into several varieties but they do *** Marchantia polymorpha L.: Evans 1917, Schuster 1992b, not seem to have been recognized by any later author, Bischler-Causse et al. 2005, Schafer-Verwimp & Pocs 2009. except that Jovet-Ast (1960) repeated his reports in her *** - subsp. ruderalis Bischl. 8c Boissel.-Dub.: Damsholt overview of Antillean hepatic distribution. We are not 2002 as M latifolia. sure about the value of the varieties. *** Marchesinia brachiata (Sw.) Schiffn.: type of Junger- mannia brachiata and M. quadridens , Swartz 1788, 1806 *** Mnioloma cellulosum (Spreng.) R. M. Schust.: Bischler both as /. brachiata, Lunan 1814 as /. brachiata , Weber 1962a as Calypogeia cellulosa, Fulford 1987 as C. cellulosa, 1815 as /. brachiata, Gottsche et al. 1844-1847 as Fulford 8c Sharp 1990. Lejeunea brachiata , Boswell 1887 as I. brachiata and I. *** Mnioloma crenulatum (Bischl.) R. M. Schust.: Fulford 1968 as Calypogeia crenulata. guilleminiana, Stephani 1898-1923 as M. quadridens , Evans 1908a, Fulford 1987, Fulford & Sharp 1990, *** Mnioloma cyclostipum (Spruce) R.M. Schust.: Schuster Reiner-Drehwald 1993, 1999, Gradstein 1994. ? Mastigobryum novae-hollandiae a6 abnorme Lindenb. 8c 1972 as Calypogeia cyclostipa, 1995 " cyclostipa" , 2000. *** Mnioloma parallelogramum (Spruce) R. M. Schust.: Fulford 1968 as Calypogeia parallelogramma. Gottsche: syntype, Lindenberg 8c Gottsche 1851. Note: Mastigobryum novae-hollandiae Lindenb. 8c Gottsche ex *** Monoclea gottschei Lindb.: Stephani 1898-1923, Pagan Nees is a synonym of Bazzania adnexa (Lehm. 8c Lindenb.) Trevis. from Australia. This unranked infra- 1939. *** - subsp. elongata Gradst. 8c Mues: Bischler-Causse et taxon must belong to some other species. *** Mastigolejeunea auriculata (Wilson) Schiffn.: Evans al. 2005, Schafer-Verwimp 8c Pocs 2009. *** Mytilopsis albifrons Spruce: Fulford 1966, Dauphin et al. 1908a, Pagan 1939, Schuster 1980b, Gradstein 1994. 1998, Vital 8c Visnadi 2000, Gradstein et al. 2001. *** Metzgeria albinea Spruce: Kuwahara 1986, Costa 2008. *** Metzgeria aurantiaca Steph.: Costa 2008. ** Neesioscyphus bicuspidatus (Steph.) Grolle: Fulford 1987. *** Neurolejeunea breutelii (Gottsche) A. Evans: Evans *** Metzgeria ciliata Raddi: type of M. oligotricha, Evans 1907b, Pagan 1939, 1942, Schuster 1980b, Fulford 1987, 1910b as M. oligotricha , Pagan 1939 as M. oligotricha , Fulford 8c Sharp 1990, Gradstein 1994, Lucking 1995, Kuwahara 1983, 1984, 1986 all as M. decipiens , Costa 2008. Schafer-Verwimp 8c Pocs 2009. *** Neurolejeunea catenulata (Nees) Schiffn.: Evans 1907b, *** Metzgeria dichotoma (Sw.) Nees: type of Jungermannia dichotoma , Swartz 1788, 1806 both as /. dichotoma , Pagan 1939, Jovet-Ast 1960, Fulford 1987, Gradstein 1994. Gottsche et al. 1844-1847, Boswell 1887, Stephani ** Nowellia evansii Grolle: holotype, Grolle 1968, Fulford 1898-1923, Evans 1910b, Pagan 1939, Miller et al. 1983, 1968 as N. caribbeana, Schuster 1984, 2002, SchaferKuwahara 1986, Costa 2008. Verwimp 8c Pocs 2009. *** Metzgeria furcata (L.) Corda: Weber 1815 as Junger- *** Odontolejeunea decemdentata (Spruce) Steph.: holotype mannia fur cat a, Costa 2008. of O. longispica, Stephani 1898-1923 as O. longispica, *** Metzgeria herminieri Schiffn.: Costa 2008. Evans 1908c as O. longispica, Thiers 1985 as O. *** Metzgeria leptoneura Spruce: Engel 1976, Miller et al. longispica, Teeuwen 1989, Gradstein 1994, Lucking 1983, Kuwahara 1982, 1984 both as M. papulosa , 1986 1995, Schafer-Verwimp 8c Pocs 2009. also as M papulosa, M. sandei and M. australis, So 2002, *** Odontolejeunea lunulata (F. Weber) Schiffn.: Evans 1904 Costa 2008, Schafer-Verwimp 8c Pocs 2009. as O. sieberiana, Pagan 1939 as O. sieberiana, Fulford 1987 as O. sieberiana, Teeuwen 1989, Schuster 1992c as ** - var. leptoneura: syntype of M. furcata f.a extensa , Nees 1838 as M. furcata f.a extensa , Engel 1978, Schuster 1992a. O. sieberiana, Gradstein 1994, Lucking 1995. *** Odontoschisma denudatum (Mart.) Dumort.: Fulford *** Metzgeria procera Mitt.: Stephani 1898-1923 as M. hamata, Pearson 1931 as M. hamata, Kuwahara 1984 1968, 1987, Schafer-Verwimp 8c Pocs2009. ** - var. denudatum: syntype of O. subrotundifolium, also as M. hamata, 1985 as M. hamata, 1986, Costa 2008, Stephani 1898-1923 as O. subrotundifolium. Pagan 1942 as Schafer-Verwimp 8c Pocs 2009. O. subrotundifolium, Schuster 1974, Fulford 8c Sharp 1990. *** Metzgeria rufula Spruce: Kuwahara 1985, 1986 both as*** Odontoschisma longiflorum (Taylor) Steph.: type of M quadrifaria , Costa 2008. Sphagnoecetis longiflora, Taylor 1846 as S. longiflora, *** Microlejeunea acutifolia Steph.: Schuster 1992c as Lejeunea acutifolia. Gottsche et al. 1844-1847 as S. longiflora , Stephani 1898-1923, Fulford 1968, 1987. *** Microlejeunea bullata (Taylor) Steph.: Schuster 1962 as*** Odontoschisma prostratum (Sw.) Trevis.: type of JungerLejeunea ulicina subsp. bullata, Lucking 1995 as Lejeunea mannia prostrata, Swartz 1788, 1806 both as /. prostrata, bullata. Lunan 1814 as /. prostrata, Weber 1815 as /. prostrata , *** Micropterygium trachyphyllum Reimers: Fulford 1966, 1987. Gottsche et al. 1844-1847 as Sphagnoecetis prostrata, Stephani 1898-1923, Frye & Clark 1945, Fulford 1968, This content downloaded from 107.0.125.4 on Wed, 10 Aug 2016 20:31:07 UTC All use subject to http://about.jstor.org/terms j6 The Bryologist 114(1)' 2011 Schuster 1972, 1974, Fulford et al. 1970, Fulford 8c Sharp** Plagiochila bancroftii Steph.: type, Gottsche et al. 1844- 1847, Stephani 1898-1923. 1990, Grolle 8c Long 2000. *** Omphalanthus filiformis (Sw.) Nees: syntype of O. *** Plagiochila bifaria (Sw.) Lindenb.: lectotype of Junger- filiformis a communis and type of Jungermannia filiformis , mannia bifaria, Swartz 1788, 1806 both as /. bifaria, Swartz 1788, 1806 both as J. filiformis , Lunan 1814 as /. Lunan 1814 as /. bifaria, Weber 1815 as /. bifaria, filiformis , Weber 1815 as /. filiformis, Gottsche et al. 1844- Lindenberg 1843, Gottsche et al. 1844-1847, Boswell 1847 as O. filiformis a communis , Evans 1907b, Pearson 1887, Stephani 1898-1923, Dugas 1929, Fulford 8c 1931, Pagan 1939, 1942, Fulford 1987, Fulford 8c Sharp Sharp 1990, Heinrichs et al. 1998a, Schafer-Verwimp 8c 1990, Reiner-Drehwald 1993, 1999, Lucking 1995, Yano 2004, Schafer-Verwimp & Pocs 2009. Pocs 2009. *** Plagiochila breuteliana Lindenb.: Fulford 1987, Heinrichs 2002, Schafer-Verwimp 8c Pocs 2009. ? Omphalanthus sulphureus y laevis Gottsche, Lindenb. 8c Nees: type, Gottsche et al. 1844-1847. Note: The species ** Plagiochila caribbeania R. M. Schust.: holotype, Schuster 1980b. is a synonym of Taxilejeunea sulphurea (Lehm. 8c Lindenb.) Schiffn. but we do not know where the * Plagiochila conferta Steph.: type, Stephani 1898-1923. unranked infrataxon belongs. T. sulphurea is not Note: The species does not seem to have been dealt with otherwise reported from Jamaica but occurs on other Caribbean Islands. since its description in 1918. *** Plagiochila cristata (Sw.) Lindenb.: lectotype of Jungermannia cristata , Swartz 1788, 1806 both as /. cristata, *** Pallavicinia lyellii (Hook.) Carruth.: Hitchcock 1898, Stephani 1898-1923, Miller et al. 1983, Bapna 8c Lunan 1814 as /. cristata, Nees 1830 as J. cristata, Weber Kachroo 2000. 1815 as /. cristata, Lindenberg 1839, Gottsche et al. 1844-1847, Dugas 1929, Pagan 1942, Fulford 8c Sharp 1990, Heinrichs et al. 1998a, Heinrichs 2002, Schafer- ? Paracromastigum bifidum subsp. densum (Pearson) R. M. Schust. nom. inval.: holotype of Cephalozia densa, Pearson 1931 as C. densay Schuster 2000. Note: Paracromastigum bifidum (Steph.) R. M. Schust. is a Verwimp 8c Pocs 2009. *** Plagiochila deflexa Mont. 8c Gottsche: Heinrichs et al. 2002a, Schafer-Verwimp 8c Pocs 2009. synonym to P. pachyrhizum (Nees) Fulford (Gradstein & Costa 2003) but the identity of the subspecies is *** Plagiochila dichotoma (P. Beauv.) Nees 8c Mont.: Lunan unknown. This is the only report of a Paracromastigum 1814 as Jungermannia dichotoma , Pagan 1939, 1942, from Jamaica that is known to us. Bapna 8c Kachroo 2000. *** Plagiochasma jamaicense (Haynes) A. Evans: lectotype ** Plagiochila divaricata Lindenb.: Fulford 1987. (and paratype) of Aytonia jamaicensisy Haynes 1907 as A.*** Plagiochila dominicensis Taylor: Fulford 1987, Heinrichs jamaicensis, Evans 1915, Haynes et al. 1923, Bischler 2002, Schafer-Verwimp 8c Pocs 2009. 1979b, Bischler-Causse et al. 2005. *** Plagiochila gymnocalycina (Lehm. 8c Lindenb.) Mont.: *** Plagiochasma rupestre (J. R. Forst. 8c G. Forst.) Steph.: holotype of P. lambertina, type of P. rutilans (3 laxa, Haynes et al. 1923, Bischler 1979a, Miller et al. 1983, Lindenberg 1840 as P. rutilans p laxa , Gottsche et al. Schuster 1992b, Schafer-Verwimp 8c Pocs 2009. 1844-1847 as P. rutilans P laxa , Gottsche 1857 as P. *** - var. rupestre. type of Aytonia evansii, Haynes 1907 as lambertina, Stephani 1898-1923 as P. lambertina, Schiffner 8c Arnell 1964 as P. rutilans var. laxa, Heinrichs Aytonia evansii , Stephani 1898-1923 as P. evansii , Evans 1915, Bischler 1979b, Bischler-Causse et al. 2005. et al. 2001. ** Plagiochila jamaicensis Lindenb. 8c Hampe: type, Hampe ** Plagiochila abrupta Lehm. 8c Lindenb.: Boswell 1887, Fulford 1987. 1852, Stephani 1898-1923, Dugas 1929, Fulford et al. 1970. *** Plagiochila adianthoides (Sw.) Lindenb.: lectotype of Jungermannia adianthoides , Swartz 1788, 1806 as /. *** Plagiochila macrostachya Lindenb.: lectotype, Lindenberg adianthoides , Lunan 1814 as /. adianthoides , Weber 1815 1840, Gottsche et al. 1844-1847, Stephani 1898-1923, as /. adianthoides , Lindenberg 1840, Gottsche et al. Pearson 1931, Pagan 1942, Schafer-Verwimp 1989, 1844-1847, Boswell 1887, Stephani 1898-1923 as P. Heinrichs 2002, Schafer-Verwimp 8c Pocs 2009. procera, Pagan 1942 as P. procera , Fulford 1987, Fulford*** Plagiochila montagnei Nees: Stephani 1898-1923 as P. 8c Sharp 1990 also as P. procera, Heinrichs et al. 1998b, Heinrichs 2002. florida , Schuster 1960, 1980b both as P. hypnoides, Fulford 8c Sharp 1990 as P. florida, Heinrichs 8c *** Plagiochila aerea Taylor: lectotype of /. adiantoides var.y Gradstein 2000. distans and "type" of /. adiantoides var.c, Swartz 1806 as*** Plagiochila patula (Sw.) Lindenb.: lectotype of Junger/. adiantoides var.c, Boswell 1887 as P. bur sat a, Fulford mannia patula , Swartz 1806 as /. patula , Weber 1815 as 1987 as P. bursatay Fulford 8c Sharp 1990, Heinrichs et /. patula, Nees 1831 as /. patula, Lindenberg 1839, al. 1998b, Grolle & Heinrichs 1999, Heinrichs 2002. Gottsche et al. 1844-1847, Boswell 1887, Stephani 1898-1923, Schiffner & Arnell 1964, Arnell 1956, *** Plagiochila alternans Lindenb. 8c Gottsche: Stephani 1898-1923 as P. columbica, Pearson 1931 as P. columbica, Miiller et al. 1999. Heinrichs et al. 1998a, Heinrichs et al. 2002b, SchaferVerwimp 8c Pocs 2009. This content downloaded from 107.0.125.4 on Wed, 10 Aug 2016 20:31:07 UTC All use subject to http://about.jstor.org/terms Soderstrom et al.: Liverworts and horn worts of Jamaica 77 *** Plagiochila rutilans Lindenb.: Pearson 1931, Fulford *** Prionolejeunea denticulata (F. Weber) Schiffh.: type of 1987. Jungermannia denticulata, Gottsche et al. 1844-1847 as Lejeunea denticulata, Boswell 1887 as L. denticulata , *** - var. rutilans: holotype of P. harrisana , Stephani 18981923 as P. harrisana , Jovet-Ast 1960 as P. harrisana, Evans 1904 as Cyclolejeunea angulistipa, Pearson 1931, Heinrichs et al. 1998b as P. harrisana , Heinrichs et al. Pagan 1939 as C. angulistipa, Jovet-Ast 1948 as C. 2001. angulistipa, Fulford 1987 as C. angulistipa and P. *** Plagiochila simplex (Sw.) Lindenb.: "type" of P. simplex var.a minor , syntype of P. distinctifolia and P. distincti- aequitexta, Schuster 1992c as C. angulistipa, Reiner- Drehwald 1999, Ilkiu-Borges 2006. folia a gracilior , lectotype of Jungermannia simplex and /.*** Prionolejeunea galliotii Steph.: Ilkiu-Borges 2006. adiantoides var. (3 angustior, Swartz 1788, 1806 both as /. *** Prionolejeunea guadalupensis (Lindenb.) Steph.: Ilkiu- Borges 2006. simplex, Lunan 1814 as /. simplex, Weber 1815 as J. simplex, Lindenberg 1839 as P. distinctifolia and P. *** Pycnolejeunea contigua (Nees) Grolle: Boswell 1887 as distinctifolia a gracilior, 1840 as P. simplex var. a minor, Lejeunea contigua. Gottsche et al. 1844-1847 as P. distinctifolia, Boswell *** Pycnolejeunea monophthalma (R. M. Schust.) Xiao L. He: 1887 as P. distinctifolia, Stephani 1898-1923 as P. holotype of Trachylejeunea monophthalma, Schuster distinctifolia, Stephani 1898-1923, Dugas 1929 , Pearson 1971 as T. monophthalma, Thiers 1986 as T. mono- 1931 as P. distinctifolia, Fulford & Sharp 1990 as P. distinctifolia var. distinctifolia and P. distinctifolia a gracilior, Heinrichs et al. 1998b, Yano 2004, SchaferVerwimp & Pocs 2009 also as P. distinctifolia. ** Plagiochila solitaria Gottsche ex Steph.: type, Stephani 1898-1923, Arnell 1956, Jovet-Ast 1960. *** Plagiochila stricta Lindenb.: lectotype, type of P. phthalma, He 1999. *** Pycnolejeunea schwaneckei (Steph.) Steph.: Evans 1906, Stephani 1898-1923, Pagan 1939, Thiers 1986, He 1999. *** Radula antilleana Castle: Castle 1960a, 1969, Fulford 1987, Yamada 1988. *** Radula fendleri Gottsche ex Steph.: Castle 1963, 1969 both as R. saccatiloba, Fulford et al. 1970 as R. saccatiloba, cinchoniensis, Lindenberg 1839, Gottsche et al. 1844- Fulford 1987 as R. saccatiloba, Yamada 1987, 1988 both 1847, Stephani 1898-1923, Pearson 1931 as P. cincho- as R. saccatiloba, Schafer-Verwimp 8c Pocs 2009. niensis, Pagan 1942, Arnell 1959, Heinrichs et al. 1998b, *** Radula flaccida Lindenb. 8c Gottsche: Castle 1939, Rycroft et al. 2002, Sim-Sim et al. 2004. *** Plagiochila subbidentata Taylor: lectotype, Taylor 1847, 1850, Stephani 1898-1923, Fulford et al. 1970, Heinrichs 2002, Schafer-Verwimp 8c Pocs 2009. *** Plagiochila superba (Nees ex Spreng.) Mont. 8c Nees: Boswell 1887, Pearson 1931. *** - var. superba: Heinrichs 2002. ** Plagiochila tamariscina Steph.: Fulford 1987, Schafer- Verwimp 8c Pocs 2009. ** Plagiochila tenuis Lindenb.: Boswell 1887, Fulford 1987. *** Plagiochila vincentina Lindenb.: Heinrichs 2002. ** Plagiochilion heteromallum (Lehm. 8c Lindenb.) Hassel: Boswell 1887 as Plagiochila heteromalla. *** Porella crispata (Hook.) Trevis.: type of Madotheca Schuster 1980b. *** Radula grevilleana Taylor: type, Taylor 1847, 1850, Stephani 1898-1923, Castle 1939, 1969, Jovet-Ast 1960. *** Radula husnotii Castle: Castle 1960a, 1969, Fulford 1987, Yamada 1988. *** Radula inflexa Gottsche ex Steph.: Stephani 1898-1923, Pagan 1942, Castle 1963, 1969, Yamada 1980, Fulford 1987, Fulford & Sharp 1990. *** Radula jamaicensis Pearson: type, Pearson 1931. *** Radula javanica Gottsche: Castle 1966 as R. longifolia, 1968 as R. macrostachya, 1969 as R. longifolia, Fulford 1987 as R. longifolia, Yamada 1982 as R. macrostachya , 1988 as R. amazonica and R. macrostachya. *** Radula mexicana Lindenb. 8c Gottsche: Castle 1964, commutata, Gottsche et al. 1844-1847 as M. commutata , 1969, Yamada 1988, 1993, Fulford 8c Sharp 1990, Stephani 1898-1923 as M. commutata, Swails 1970, Schafer-Verwimp 8c Pocs 2009. Fulford 8c Sharp 1990, So 2005. *** Porella leiboldii (Lehm. & Lindenb.) Trevis.: Fulford & Sharp 1990. *** Radula microlobula Castle: holotype, Castle 1960a, 1969, Yamada 1988. *** Radula pallens (Sw.) Nees 8c Mont.: type of Jungerman- *** Porella swartziana (F.Weber) Trevis.: lectotype of nia pallens, Swartz 1788, 1806 both as J. pallens, Lunan Jungermannia swartziana, Weber 1815 as /. swartziana, 1814 as /. pallens, Weber 1815 as /. pallens , Gottsche et Gottsche et al. 1844-1847 as Madotheca swartziana , al. 1844-1847, Boswell 1887, Castle 1960b, 1968, 1969, Barbour 1902, Clark & Svihla 1944a, Frye & Clark 1946, Swails 1970, Schuster 1980b, Fulford 8c Sharp 1990, Yano 2004, So 2005. Fulford 1987, Fulford 8c Sharp 1990, Dauphin 2005. ? - var. p minor Gottsche, Lindenb. et Nees nom. illeg.: syntype, Gottsche et al. 1844-1847. *** Prionolejeunea aemula (Gottsche) A. Evans: Fulford 1987, *** Radula portoricensis Steph.: Castle 1937, 1969, Pagan Ilkiu-Borges 2006. 1939, 1942, Jovet-Ast 1960, Yamada 1981, 1988, Schafer*** Prionolejeunea ampliretis Herzog: Ilkiu-Borges 2006. *** Prionolejeunea decora (Taylor) Steph.: Pagan 1942, Thiers 1986, Grolle 1991, Ilkiu-Borges 2006. Verwimp 8c Pocs 2009. *** Radula quadrata Gottsche: "type" of R. propagulifera, Castle 1965 as R. mollis and R. propagulifera , Schuster This content downloaded from 107.0.125.4 on Wed, 10 Aug 2016 20:31:07 UTC All use subject to http://about.jstor.org/terms 78 The Bryologist 114(1)' 2011 1980b as R. mollis , Miller et al. 1983 as R. mollis , Reiner- as /. gracillima , Paton 1999 as /. gracillima, Damsholt Drehwald 1994. 2002 as /. gracillima . *** Radula tectiloba Steph.: Castle 1964, 1969, Yamada 1981,*** Solenostoma hyalinum (Lyell) Mitt.: Vana 1974 as 1988, 2000, Fulford 1987, Fulford 8c Sharp 1990, Schafer- Jungermannia hyalina , Vana 1975b as /. hyalina, Verwimp 8c Pocs 2009. Gradstein 8c Vana 1987 as /. hyalina, Van Zanten 8c *** Radula underwoodii Castle: holotype, Castle 1960a, 1969, Yamada 1988. Gradstein 1988 as /. hyalina, Fulford 8c Sharp 1990 as /. hyalina, Paton 1999 as /. hyalina, Schuster 2002. *** Radula voluta Taylor ex Gottsche, Lindenb. 8c Nees: *** Steereella lilliana (Steph.) Kuwah.: holotype of Metzgeria lilliana, Stephani 1898-1923 as M lilliana, Kuwahara 1986, Grolle & So 2002 as M. lilliana, Costa 2008 as M. Castle 1965 as R. ramulina , Fulford 8c Sharp 1990 as R. ramulina , Schafer-Verwimp 8c Pocs 2009. lilliana . *** Reboulia hemisphaerica (L.) Raddi: Bischler-Causse et al. 2005, Schafer-Verwimp 8c Pocs 2009. *** Steereella linearis (Sw.) Kuwah.: lectotype of Jungermannia linearis and syntype of Metzgeria furcata p2 ** Rectolejeunea emarginuliflora (Gottsche ex Schiffn.) A.Evans: Thiers 1986. minor, Swartz 1788 as /. linearis, Lunan 1814 as /. linearis , Nees 1830 as Jungermannia furcata y line- *** Riccardia fucoidea (Sw.) Schiffn.: type of Jungermannia fucoidea , Swartz 1788, 1806 both as /. fucoidea , Lunan aris, 1838 as M. furcata 02 minor, Stephani 1898-1923 as 1814 as /. fucoidea , Weber 1815 as J. fucoides , Nees 1830 M linearis, Evans 1910b as M. linearis, Engel 1978 as M. as /. fucoides, Gottsche et al. 1844-1847 as Metzgeria linearis, Kuwahara 1979, Kuwahara 1986, Costa 2008 as fucoides , Boswell 1887 as Pseudoneura fucoides, Stephani M. linearis, Schafer-Verwimp 8c Pocs 2009 as M. linearis. 1898-1923 as Aneura fucoides , Pearson 1931 as A. *** Stictolejeunea squamata (Willd. ex F. Weber) Schiffn.: fucoides, Pagan 1939, Arnell 1959, Engel 1978 as M. Boswell 1887 as Lejeunea squamata, Stephani 1890 as L. fucoidea , Meenks 1987, Forrest et al. 2006, Schafer- squamata, Evans 1907b, Pagan 1939, 1942, Jovet-Ast Verwimp 8c Pocs 2009. 1960, Fulford 1987, Gradstein 1985, 1994, Lucking 1995, Bapna 8c Kachroo 2000. *** Riccardia palmata (Hedw.) Carruth.: type of Junger- *** Symbiezidium barbiflorum (Lindenb. 8c Gottsche) A. mannia polyphylla, Swartz 1788, 1806 both as /. palmata, Evans: Fulford 1942, Gradstein & van Beek 1985, Lunan 1814 as /. polyphylla . Lucking 1995. ** Riccardia virgata (Gottsche ex Steph.) Pagan: Ste- phanil898-1923 as Aneura virgata , Pagan 1939, 1942.*** Symbiezidium transversale (Sw.) Trevis.: Swartz 1788, 1806 both as Jungermannia transversalis, Lunan 1814 as /. *** (Riccia fluitans L.: Haynes et al. 1923). Note: Bischler- transversalis, Weber 1815 as /. transversalis, Evans 1908a, Causse et al. (2005) doubts the correctness of the report. *** Scapania portoricensis Hampe 8c Gottsche: holotype of S. Fulford 1942, 1987 both also as S. subrotundum, Schuster 1992c, Lucking 1995. grandis , Boswell 1887 as S. grandis , Evans 1912c, Pagan *** - var. hookerianum (Gottsche) Gradst. 8c Beek: Evans 1939, Fulford et al. 1970, Fulford 8c Sharp 1990, Schuster 1908a as S. vincentinum, Pearson 1931 as Platylejeunea 2002, Schafer-Verwimp 8c Pocs 2009. vincentina, Gradstein & van Beek 1985, Gradstein 1994. ** - var. portoricensis: Hong 1988 as S. portoricensis var. portoricensis f. integrifolia and f. grossealata. *** - var. transversale. lectotype of Jungermannia transver- *** Schiffneriolejeunea polycarpa (Nees) Gradst.: Evans 1908b salis, Pagan 1939, Gradstein 8c van Beek 1985, Gradstein 1994, Reiner-Drehwald 1999. as Ptychocoleus polycarpus , Pagan 1939, 1942 both as P. ** Symphyogyna digitisquama Steph.: type, Stephani 1898poly carpus, Fulford 8c Sharp 1990, Gradstein 1974, 1994. 1923, Evans 1927, Pagan 1942. *** Solenostoma amoenum (Lindenb. 8c Gottsche) R. M. Schust. ex Vana, Hentschel 8c Heinrichs: Vana 1974 as ** Symphyogyna fuscovirens A. Evans: holotype, Evans 1927. ** Symphyogyna sinuata (Sw.) Mont. 8c Nees: type of Jungermannia amoena , Fulford 8c Sharp 1990 as /. amoena , Schuster 2002. Jungermannia sinuata, Swartz 1788, 1806 both as /. *** Solenostoma amplexifolium (Hampe ex Lehm.) Vana 8c sinuata, Lunan 1814 as /. sinuata, Weber 1815 as /. Schaf.-Verw.: type of Aplozia boweri, Pearson 1931 as A. sinuata, Gottsche et al. 1844-1847, Stephani 1898-1923, boweri, Pagan 1942 as Plagiochila amplexifolia, Vana Evans 1927. 1974 as Jungermannia linguifolia , Fulford 1987 as *** Syzygiella perfoliata (Sw.) Spruce: type of Jungermannia Lethocolea amplexifolia , Schafer-Verwimp 8c Pocs 2009. *** Solenostoma callithrix (Lindenb. 8c Gottsche) Steph.: perfoliata, Swartz 1788, 1806 both as /. perfoliata, Lunan 1814 as /. perfoliata, Weber 1815 as /. perfoliata, Pearson 1931 as Eucalyx callithrix, Vana 1974 as Gottsche et al. 1844-1847 as /. perfoliata, Stephani Jungermannia callithrix, Fulford 1987 as J. callithrix, 1898-1923, Pagan 1939, 1942, Inoue 1966, Fulford et al. Fulford 8c Sharp 1990 as /. callithrix, Schumacker 8c Vana 1970, Fulford 1987, Fulford 8c Sharp 1990. 1999 as /. callithrix, Schafer-Verwimp 8c Pocs 2009 as /. *** Syzygiella rubricaulis (Nees) Steph.: "type" of Jamesoniella callithrix. decurrens , Grolle 1971 as /. rubricaulis, Fulford 1987 as /. *** Solenostoma gracillimum (Sm.) R. M. Schust.: Vana 1975a as Jungermannia gracillima, Fulford 8c Sharp 1990 rubricaulis, Fulford 8c Sharp 1990 as /. rubricaulis, Schafer- Verwimp 8c Pocs 2009 as /. rubricaulis. This content downloaded from 107.0.125.4 on Wed, 10 Aug 2016 20:31:07 UTC All use subject to http://about.jstor.org/terms Soderstrom et al.: Liverworts and hornworts of Jamaica 79 *** Taxilejeunea debilis (Lehm. & Lindenb.) Steph.: "type" of Omphalanthus sulphureus var. integerrima , Stephani 1890 as O. sulphureus var. integerrima , Evans 1921, Fulford 1987, Lucking 1995, Schafer-Verwimp & Pocs *** Trichocolea flaccida (Spruce) J. B. Jack & Steph.: Fulford 1963, 1987. *** Trichocolea paraphyllina (Spruce) Steph.: Hatcher 1958, Fulford 1963, 1987, Fulford & Sharp 1990. 2009. Note: In a recent molecular study (Wilson et al. *** Tylimanthus laxus (Lehm. & Lindenb.) Spruce: type of T. 2007), the type of Taxilejeunea was nested in Lejeunea jamaicensis , Boswell 1887 as Plagiochila approximata , and it was therefore treated as a synonym by Crandall- Stephani 1898-1923 as T. jamaicensis , Fulford 1987 also Stotler et al. (2009). However, in the study T. pterigonia (the generic type) was only identified with a "cf.". In addition, many of the names lacks combinations in Lejeunea and a substantial part of them will even need new names if moved to Lejeunea. Thus, we prefer to keep Taxilejeunea for now. ** Taxilejeunea densiflora A. Evans: holotype, Evans 1921, Thiers 1986 as Lejeunea densiflora. as T. approximates, Burghardt & Gradstein 2008. *** Xylolejeunea aquarius (Spruce) Xiao L. He & Grolle: Schuster 1980b as Trachylejeunea aquarius. *** Xylolejeunea crenata (Nees & Mont.) Xiao L. He & Grolle: He & Grolle 2001. ** Zoopsidella antillana (Steph.) H.Rob.: Schuster 1972. ** - subsp. jamaicensis R. M. Schust.: holotype, Schuster 1999. ** Taxilejeunea graminicolor (Spruce) Steph.: Pearson 1931. ** Taxilejeunea jamaicensis A. Evans: holotype, Evans 1921. ? Taxilejeunea jamaicensis Steph. nom. illeg .: "type", Stephani 1898-1923. Note: The two Taxilejeunea jamaicensis are heterotypic and we are not sure if they represent the same taxon or not. * Taxilejeunea langiana Pearson: type, Pearson 1931. It does not seem that the taxon has been re-studied since its description in 1931. *** Taxilejeunea obtusangula (Spruce) A. Evans: Schuster 1980b, Schafer-Verwimp & Pocs 2009. *** Taxilejeunea pterigonia (Lehm. & Lindenb.) Schiffn.: Evans 1921, Pearson 1931, Thiers 1986 as Lejeunea pterigonia. Taxa rejected from Jamaica *** Acromastigum inaequilaterum (Lehm. & Lindenb.) A. Evans. The report of this Asiatic taxon by Bonner (1963, as Bazzania elegantula) is a clear error. *** Bazzania trilobata (L.) Gray. The report of this boreal taxon by Weber (1815, as Jungermannia trilobata ) is rejected here. *** Dendromastigophora flagellifera (Hook.) R. M. Schust. The report of this Australian taxon by Gottsche et al. (1844-1847, as Sendtnera flagellifera) is rejected here. *** Frullania hypoleuca Nees: The report of this Asiatic taxon by Miller et al. (1983) is rejected here. *** Frullania serrata Gottsche: Reported from Jamaica by ** Taxilejeunea tenera (Sw.) Schiffn.: type of Jungermannia tenera , Swartz 1788, 1806 both as /. tenera , Lunan 1814 as /. tenera , Weber 1815 as J. tenera , Gottsche et al. 1844-1847 as Lejeunea tenera , Stephani 1898-1923, Fulford & Sharp 1990, Reiner- Drehwald 1999, 2000b as Neopotamolejeunea tenera. *** Telaranea nematodes (Gottsche ex Austin) M. Howe: Schuster 1969, Fulford 1987. Note: The taxonomy of the two Telaranea taxa reported from Jamaica has been Clark & Svihla (1948a) but tropical America are doubtful fide Hattori (1977) and we here reject all reports. ** Haplomitrium mnioides (Lindb.) R. M. Schust. The report of this Asiatic taxon by Hitchcock (1898, as Calobryum mnioides) is rejected here. *** Herbertus aduncus (Dicks.) Gray. Boswell (1887, as Sendtnera adunca) reports it from Jamaica but he includes S. juniperina in his concept. *** Mannia androgyna (L.) A. Evans. The report by Weber confused in the past and the two species was generally (1815, as Marchantia androgyna) is rejected by Bischler- considered synonyms until Engel & Smith Merrill Causse et al. (2005). (2004) lectotypified T. sejuncta and showed that they were different species. Thus, which species the reports here really belongs to, and if two species occur on the island, is unclear until the specimens have been reexamined. *** Marchantia linearis Lehm. et Lindenb. The report of this rare Asiatic taxon by Lehmann (1832) is rejected here. *** Monoclea forsteri Hook. The reports of this Australian taxon by Miller et al. (1983) and Campbell (1987) are due to inclusion of M. gottschei in their concept. *** Telaranea sejuncta (Angstr.) S.W.Arnell: Schuster 1972 (s.lat.). ? Trachylejeunea jamaicensis Pearson: type, Pearson 1931. Note: Trachylejeunea is now considered a synonym to Cheilolejeunea (Gradstein & Ilkiu-Borges 2009) but this species has not been transferred there. It does not seem that it has been re-studied since its description in 1931. *** Trichocolea brevifissa Steph.: Pearson 1931 as T. cubensis , Hatcher 1958, Fulford 1963, 1987. *** Trichocolea elliottii Steph.: Pearson 1931, Hatcherl958, Fulford 1963, 1987. *** Plagiochila arbuscula (Brid. ex Lehm. & Lindenb.) Lindenb. The report of this Asiatic taxon by Lindenberg (1839) is rejected here. *** Podomitrium phyllanthus (Hook.) Mitt. The report of this Australian taxon by Gottsche et al. (1844-1847, as Blyttia phyllanthus) is rejected here. *** Radula lindenbergiana Gottsche ex C.Hartm. Reported by Boswell (1887, as R. complanata var. a* propagulifera) but we reject it. *** Riccardia chamedryfolia (With.) Grolle. Syntype of Jungermannia multifida var.y pinnatifida and as such This content downloaded from 107.0.125.4 on Wed, 10 Aug 2016 20:31:07 UTC All use subject to http://about.jstor.org/terms 80 The Bryologist 114(1): 2011 reported by Gottsche et al. (1844-1847, as Aneura Bryopteris filicina a hookeriana Gottsche, Lindenb. 8c Nees = B. filicina pinnatifida f.p denticulata ) and Engel (1978, as R. pinnatifida). However, it is lectotypified with its B. filicina p arguta Gottsche, Lindenb. 8c Nees = B. filicina European syntype and the syntype from Jamaica belongs B. fruticulosa Taylor = B. filicina to some other species. B. trinitensis (Lehm. 8c Lindenb.) Lehm. 8c Lindenb. = B. - Riccia reticulata Sw. The type of this taxon described by filicina Swartz (1788) and repeatedly reported is a fern protallium Calobryum mnioides (Lindb.) Steph. = Haplomitrium mnioides (Gradstein 8c Costa 2003). Calypogeia biapiculata (Spruce) Steph. = C. peruviana *** Spruceanthus semirepandus (Nees) Verd. The type of C. cellulosa (Spreng.) Steph. = Mnioloma cellulosum Phragmicoma lehmanniana is from Jamaica but that isC. crenulata Bischl. = Mnioloma crenulatum probably mislabelled fide Geissler 8c Gradstein (1994).C. cyclostipa (Spruce) Steph. = Mnioloma cyclostipum The taxon occurs otherwise in SE Asia. C. heterophylla (Steph.) = C. peruviana C. parallelogramma (Spruce) Steph. = Mnioloma parallelogramum Synonyms Acrostolia fucoidea (Sw.) Dumort. = Riccardia fucoidea Anastrophyllum harrisanum Steph. = A. tubulosum, syn. nov. Aneura fucoidea (Sw.) Besch. = Riccardia fucoidea A. pinnatifida P denticulata Nees = Riccardia chamedryfolia A. virgata Gottsche ex Steph. = Riccardia virgata C. portoricensis (Steph.) A. Evans = C. peruviana Calyptrocolea pittieri (Steph.) R. M. Schust. = Adelanthus pittieri Candollea simplex (Sw.) Raddi = Plagiochila simplex Cephalozia caribbeania Fulford = C. crassifolia C. densa Pearson = Paracromastigum bifidum subsp. densum Anthoceros confusus Steph. = A. cavernosus C. subforficata Herzog = C. crassifolia A. crispus Sw. = Dendroceros crispus Ceratolejeunea guadelupensis Steph. = C. cubensis A. monkeyanus Steph. = A. punctatus A. rigidus Lehm. ex Gottsche = Leiosporoceros dussii C. maritima (Spruce) Steph. = C. cornuta C. mollis Pearson = Lepidolejeunea involuta Aphanolejeunea angustissima Steph. = Cololejeunea papillosa C. plumula (Spruce) Steph. = C. coarina A. camillii (Lehm.) R.M.Schust. = Cololejeunea camillii C. schwaneckei Steph. = C. cornuta A. gracilis Ast = Cololejeunea gracilis C. variabilis (Lindenb.) Schiffn. = C. cornuta A. gracilis var. linearifolia R. M. Schust. = Cololejeunea gracilisCololejeunea myriocarpa (Nees 8c Mont.) C.Massal. = C. A. jamaicensis R. M. Schust. = Cololejeunea sicaefolia subsp. jamaicensis minutissima subsp. myriocarpa Cyclolejeunea affixa (Taylor) Steph. = C. accedens Aphanolejeunea sicaefolia (Gottsche) A. Evans = Cololejeunea sicaefolia C. angulistipa (Steph.) A. Evans = Prionolejeunea denticulata Cyrtolejeunea holostipa (Spruce) A. Evans = Cheilolejeunea Aphanolejeunea verrucosa Ast = Cololejeunea microscopica var. africana holostipa Cystolejeunea lineata (Lehm. 8c Lindenb.) A. Evans = Aplozia boweri Pearson = Solenostoma amplexifolium Archilejeunea viridissima (Lindenb.) A. Evans = A. parviflora Aspiromitus harrisanus Steph. = Anthoceros harrisanus Cheilolejeunea lineata Dicranolejeunea aberrans (Lindenb. 8c Gottsche) Steph. = Acanthocoleus aberrans A. rigidus (Lehm. ex Gottsche) Steph. = Leiosporoceros dussii D. affixa (Taylor) Steph. = Cyclolejeunea accedens Aytonia evansii Haynes = Plagiochasma rupestre var. rupestre D. phyllorhiza (Nees) Schiffn. = Brachiolejeunea phyllorhiza A. jamaicensis Haynes = Plagiochasma jamaicense Diplasiolejeunea brachyclada A. Evans = D. cavifolia Basichiton tomentosa (Hoffm.) Trevis. = Leiomitra tomentosa D. cubensis Tixier = D. brunnea Bazzania breuteliana (Lindenb. 8c Gottsche) Trevis. = B. Dumortiera nepalensis (Taylor) Nees = D. hirsuta subsp. hookeri nepalensis B. elegantula (De Not.) Trevis. = Acromastigum inaequilaterum B. latidens (Gottsche ex Steph.) Fulford = B. hookeri Echinocolea asperrima (Spruce) R. M. Schust. = Lejeunea asperrima B. platystipula Fulford = B. roraimensis E. dilatata (A.Evans) R.M.Schust. = Lejeunea subspathulata B. quadricrenata (Gottsche ex Steph.) Pagan = B. aurescens Echinomitrion furcatum var. lineare (Sw.) Huebener = B. tenera (Gottsche 8c Lindenb.) Trevis. = B. cuneistipula B. vincentina (Lehm. 8c Lindenb.) Trevis. = B. stolonifera Bellincinia commutata (Gottsche) Kuntze = Porella crispata Steereella linearis Eucalyx callithrix (Lindenb. 8c Gottsche) Pearson = Solenostoma callithrix Blyttia phyllanthus (Hook.) Nees = Podomitrium phyllanthus Eulejeunea flava (Sw.) Schiffn. = Lejeunea flava Brachiolejeunea corticalis (Lehm. 8c Lindenb.) Schiffti. = Euosmolejeunea oerstediana (Lindenb. 8c Hampe ex Hampe) Frullanoides corticalis B. insularis A. Evans = Frullanoides liebmaniana Steph. = Lejeunea oerstediana B. longispica Steph. = Frullanoides bahamensis Evansia jamaicensis Douin = Cephaloziella antillana Frullania atrata (Sw.) Dumort. = F. atrata B. sulcata A. Evans ex Steph. = B. leiboldiana F .atrata var. subtilissima Nees ex Mont. = F. subtilissima This content downloaded from 107.0.125.4 on Wed, 10 Aug 2016 20:31:07 UTC All use subject to http://about.jstor.org/terms Soderstrom et al.: Liverworts and hornworts of Jamaica 8l F. atrata a swartzii Nees = F. atrata /. filiformis Sw. = Omphalanthus filiformis F. cubensis Steph. = F. riojaneirensis J. flava Sw. = Lejeunea flava F. gymnotis Nees & Mont. = F. caulisequa J. fucoidea Sw. = Riccardia fucoidea F. hians (Lehm. & Lindenb.) Lehm. & Lindenb. = F. arecaev ar. L. = Metzgeria furcata J. furcata J. furcata y linearis (Sw.) Nees = Steereella linearis arecae F. jamaicensis Steph. = Jubula pennsylvanicaJ. subsp. bogotensis gracillima Sm. = Solenostoma gracillimum F. neesii Lindenb. = F. kunzei J. hyalina Lyell = Solenostoma hyalinum F. obscura (Sw.) Dumort. = F. riojaneirensis J. intumescens Lehm. & Lindenb. = Frullania intumescens F. tenuis Hampe ex Lehm. = F. brasiliensis J. jamaicensis Nees = Lophozia jamaicensis Gamoscyphus perfoliatus (Sw.) Trevis. = Syzygiella perfoliataJ. juniperina (Sw.) Sw. = Herbertus juniperoideus Harpalejeunea heterodonta A. Evans = H. tridens J. juniperoidea Sw. = Herbertus juniperoideus Herbertus juniperinus (Sw.) Spruce = H. juniperoideus J. linearis Sw. = Steereella linearis H. pensilis (Taylor) Spruce = H. juniperoideus subsp. pensilis J. linguifolia Gottsche = Solenostoma amplexifolium Herpetium jamaicense Lehm. & Lindenb. = Bazzania jamaicensis J. multifida var. y pinnatifida Weber = Riccardia chamedryfolia /. obscura Sw. = Frullania riojaneirensis H. stoloniferum (Sw.) Nees ex Mont. = Bazzania stolonifera J. pallens Sw. = Radula pallens Hygrolejeunea cerina (Lehm. & Lindenb.) Steph. = Lejeunea J. palmata Hedw. = Riccardia palmata cerina J. palmata var. P polyphylla (Sw.) Steud. = Riccardia palmata Isotachis swartziana (Gottsche, Lindenb. & Nees) Steph. = I. serrulata J. patula Sw. = Plagiochila patula J. perfoliata Sw. = Syzygiella perfoliata I. urbanii Steph. = I. serrulata J. polyphylla Sw. = Riccardia palmata Jamesoniella decurrens Steph. ex Bonner = Syzygiella rubricaulis J. prostrata Sw. = Odontoschisma prostratum J. rubricaulis (Nees) Grolle = Syzygiella rubricaulis J. serrulata Sw. = Isotachis serrulata Jubula bogotensis Steph. = /. pennsylvanica subsp. bogotensis J. serrulata var. a swartziana Gottsche, Lindenb. & Nees = }. hutchinsiae subsp. bogotensis (Steph.) Verd. = /. pennsylvanica subsp. bogotensis J. piligera var. jamaicensis Pearson = /. pennsylvanica subsp. bogotensis Isotachis serrulata J. simplex Sw. = Plagiochila simplex /. sinuata Sw. = Symphyogyna sinuata J. stolonifera Sw. = Bazzania stolonifera Jungermannia adianthoides Sw. = Plagiochila adianthoides J. swartziana F.Weber = Porella swartziana /. adiantoides var. c Sw. = Plagiochila aerea J. swarzii Raeusch. = Leiomitra tomentosa J. adiantoides var. P angustior Sw. ex Steud. = Plagiochila simplex /. tenera Sw. = Taxilejeunea tenera J. tomentella var. p tomentosa (Sw.) Lindenb. = Leiomitra J. adiantoides var. y distans Sw. ex Steud. = Plagiochila aerea tomentosa J. amoena Lindenb. & Gottsche = Solenostoma amoenum J. tomentosa Sw. = Leiomitra tomentosa J. atrata Sw. = Frullania atrata J. transversalis Sw. = Symbiezidium transversale /. bifaria Sw. = Plagiochila bifaria J. trilobata L. = Bazzania trilobata J. bipinnata Sw. = Aneura bipinnata J. brachiata Sw. = Marchesinia brachiata Lejeunea acutifolia (Steph.) R. M. Schust. = Microlejeunea J. callithrix Lindenb. & Gottsche = Solenostoma callithrix Kurzia verrucosa (Steph.) Grolle = K. capillaris var. verrucosa acutifolia J. capillaris Sw. = Kurzia capillaris Lejeunea bullata Taylor = Microlejeunea bullata J. coadunata Sw. = Chiloscyphus coadunatus L. brachiata (Sw.) Nees = Marchesinia brachiata J. connata Sw. = Chiloscyphus connatus L. contigua (Nees) Mont. = Pycnolejeunea contigua J. cornuta Lindenb. = Ceratolejeunea cornuta J. corticalis Lehm. & Lindenb. = Frullanoides corticalis L. convexistipa (Lehm. & Lindenb.) Gottsche, Lindenb. & Nees J. cristata Sw. = Plagiochila cristata L. cornuta (Lindenb.) Nees = Ceratolejeunea cornuta J. cupressina Sw. = Lepidozia cupressina L. corticalis (Lehm. & Lindenb.) Bosw. = Frullanoides corticalis J. cupressina a swartziana Lehm. & Lindenb. = Lepidozia L. cubensis Mont. = Ceratolejeunea cubensis cupressina = Cyclolejeunea convexistipa Lejeunea densiflora (A.Evans) B.M.Thiers, nom. inval. = J. cupressina a* major Lehm. & Lindenb. = Lepidozia cupressina J. denticulata F. Weber = Prionolejeunea denticulata Taxilejeunea densiflora L. denticulata (F. Weber) Nees = Prionolejeunea denticulata J. dichotoma (P. Beauv.) Steud. = Plagiochila dichotoma L. filicina (Sw.) Nees & Mont. = Bryopteris filicina J. dichotoma Sw. = Metzgeria dichotoma L. fuscescens Hampe ex Lehm. = Archilejeunea fuscescens J. diffusa Sw. = Bryopteris diffusa L. guilleminiana Nees = Marchesinia brachiata J. distinctifolia (Lindenb.) Hook. f. & Taylor = Plagiochila simplex I. lehmanniana Steph. = Spruceanthus semirepandus L. lucens Taylor = L. laetevirens J. filicina Sw. = Bryopteris filicina L. mougeotii Lindenb. & Gottsche = Cyclolejeunea convexistipa This content downloaded from 107.0.125.4 on Wed, 10 Aug 2016 20:31:07 UTC All use subject to http://about.jstor.org/terms 82 The Bryologist 114(l) : 2011 Lejeunea pililoba Spruce = Lejeunea trinitensis L. proteoides Lehm. 8c Lindenb. = Anoplolejeunea conferta Lejeunea pterigonia Lehm. et Lindenb. = Taxilejeunea M. jamaicense (Lehm. 8c Lindenb.) Gottsche, Lindenb. 8c Nees = Bazzania jamaicensis M. pycnophyllum Taylor = Bazzania pycnophylla M. stoloniferum (Sw.) Lindenb. 8c Gottsche = Bazzania pterigonia L. squamata (Willd. ex F. Weber) Nees ex Mont. = Stictolejeunea squamata L. squamata y neesii Gottsche, Lindenb. & Nees = Stictolejeunea stolonifera M. tenerum Gottsche 8c Lindenb. = Bazzania cuneistipula M. vincentianum (Lehm. 8c Lindenb.) Gottsche, Lindenb. 8c Nees = Bazzania stolonifera squamata L. tenera (Sw.) Gottsche, Lindenb. & Nees = Taxilejeunea Mastigophora capillaris (Sw.) Trevis. = Kurzia capillaris M. cupressina (Sw.) Trevis. = Lepidozia cupressina tenera L. transversalis (Sw.) Nees = Symbiezidium transversale M. juniperina (Sw.) Nees = Herbertus juniperoideus Metzgeria australis Steph. = M. leptoneura L. ulicina subsp. bullata (Taylor) R.M.Schust. = Microlejeunea ulicina M. decipiens (C.Massal.) Schiffh. = M. furcata M. fucoidea (Sw.) Mont. 8c Nees = Riccardia fucoidea Lepidolejeunea punctata (Herzog) R. M. Schust. = L. involuta Lepidozia capillaris (Sw.) Trevis. = Kurzia capillaris I. verrucosa Steph. = Kurzia capillaris var. verrucosa Leptocolea appressa A. Evans = Cololejeunea appressa M. furcata a extensa (Nees) Gottsche, Lindenb. 8c Nees = M leptoneura M furcata a extensa Nees = M. leptoneura var. leptoneura M. furcata 02 minor Nees = Steereella linearis L. scabriflora (Gottsche ex Steph.) A. Evans = Cololejeunea M. hamata (Mitt.) Lindb. = M. procera obliqua Leptolejeunea obovata Bischl. = L. radicosa M. lilliana Steph. = Steereella lilliana Lethocolea amplexifolia (Hampe ex Lehm.) Grolle = M. linearis (Sw.) Austin = Steereella linearis Solenostoma amplexifolium Lophocolea coadunata (Sw.) Nees 8c Mont. = Chiloscyphus coadunatus M. oligotricha A. Evans = M. ciliata M. papulosa Steph. = M leptoneura M. quadrifaria Steph. = M. rufula L. connata (Sw.) Mont. & Nees = Chiloscyphus connatus M. sandei Schiffn. = M. leptoneura L. guadeloupensis Steph. = Chiloscyphus guadeloupensis Microlepidozia capillaris (Sw.) Fulford = Kurzia capillaris L. liebmanniana Gottsche = Chiloscyphus liebmannianus M. verrucosa (Steph.) Fulford = Kurzia capillaris var. verrucosa Lophocolea martiana Nees = Chiloscyphus martianus Neopotamolejeunea tenera (Sw.) M.E.Reiner = Taxilejeunea L. muricata (Lehm.) Nees = Chiloscyphus muricatus tenera L. trapezoidea Mont. = Chiloscyphus breutelii Nowellia caribbeana Fulford = N. evansii Lopholejeunea howei A. Evans = L. eulopha Odontolejeunea longispica A. Evans = 0. decemdentata L. muelleriana (Gottsche) Schiffn. = L. nigricans O. sieberiana (Gottsche) Schiffh. = O. lunulata Madotheca commutata Gottsche = Porella crispata Odontoschisma portoricensis (Hampe et Gottsche) Steph. = M. swartziana (F. Weber) Lehm. & Lindenb. = Porella swartziana Anomoclada portoricensis O. subrotundifolium Steph. = O. denudatum var. denudatum Marchantia androgyna L. = Mannia androgyna M. dillenii Lindb. = M. chenopoda Omphalanthus filiformis a communis Gottsche, Lindenb. 8c Nees = O. filiformis M. disjuncta Sull. = M. inflexa O. sulphureus var. integerrima Steph. = Taxilejeunea debilis M. domingensis Lehm. 8c Lindenb. = M. paleacea subsp. Phragmicoma affvxa Taylor = Cyclolejeunea accedens P. corticalis (Lehm. 8c Lindenb.) Lehm. 8c Lindenb. = paleacea M. hirsuta Sw. = Dumortiera hirsuta M. latifolia Gray = M. polymorpha subsp. ruderalis Frullanoides corticalis P. filiformis (Sw.) Nees = Omphalanthus filiformis M. papillata subsp. inflexa (Nees 8c Mont.) R. M. Schust. = M.P. lehmanniana Nees = Spruceanthus semirepandus P. transversalis (Sw.) Nees = Symbiezidium transversale inflexa M. swartzii Lehm. 8c Lindenb. = M. chenopoda Marchesinia corticalis (Lehm. 8c Lindenb.) Kuntze = Frullanoides corticalis M. lehmanniana (Nees) Kuntze = Spruceanthus semirepandus M. quadridens Steph. = M. brachiata Plagiochasma evansii (Haynes) Steph. = Plagiochasma rupestre var. rupestre Plagiochila amplexifolia Hampe ex Lehm. = Solenostoma amplexifolium P. approximata Lindenb. = Tylimanthus laxus Mastigobryum armatistipulum Steph. = Bazzania armatistipula P. bursata (Lam.) Lindenb. = P. aerea M. cuneifolium Gottsche ex Duss = Bazzania gracilis Plagiochila cinchoniensis Pearson = P. stricta M. cuneistipulum Gottsche 8c Lindenb. = Bazzania cuneistipula P. columbica (A. Evans) Gottsche = P. alternans M. elegantulum (Gottsche) Gottsche ex Steph. = Bazzania P. distinctifolia Lindenb. = P. simplex P. distinctifolia a gracilior Lindenb. = P. simplex longistipula Mastigobryum elegantulum Gottsche = Bazzania longistipula P. florida Spruce = P. montagnei M. hansenii Steph. = Bazzania longistipula P. harrisana Steph. = P. rutilans var. rutilans This content downloaded from 107.0.125.4 on Wed, 10 Aug 2016 20:31:07 UTC All use subject to http://about.jstor.org/terms Soderstrom et at.: Liverworts and hornworts of Jamaica 83 P. heteromalla (Lehm. & Lindenb.) Lehm. & Lindenb. = Plagiochilion heteromallum P. hypnoides Willd. ex Lindenb. = P. montagnei P. lambertina Gottsche = P. gymnocalycina P. procera Lindenb. = P. adianthoides P. rutilans P laxa Lindenb. = P. gymnocalycina Thysananthus lehmannianus (Nees) Steph. = Spruceanthus semirepandus Thysanolejeunea lehmanniana Steph. = Spruceanthus semirepandus Trachylejeunea aquarius (Spruce) A. Evans = Xylolejeunea aquarius P. simplex var. a minor Lindenb. = P. simplex T. dilatata A. Evans = Lejeunea subspathulata Platylejeunea transversalis (Sw.) Schiffn. = Symbiezidium T. monophthalma R. M. Schust. = Pycnolejeunea transversale monophthalma P. vincentina (Gottsche) Schiffn. = Symbiezidium transversale var. hookerianum Trichocolea tomentella P tomentosa (Sw.) Gottsche, Lindenb. & Nees = Leiomitra tomentosa Pleuroschisma prostratum (Sw.) Mitt. = Odontoschisma T. tomentosa (Sw.) Gottsche = Leiomitra tomentosa Tylimanthus approximatus (Lindenb.) Besch. = T. laxus prostratum Prionolejeunea aequitexta A. Evans = P. denticulata T. jamaicensis Steph. = T. laxus Prionolobus evansii Douin = Cephaloziella granatensis Viviania sinuata (Sw.) Raddi = Symphyogyna sinuata Pseudoneura fucoidea (Sw.) Gottsche = Riccardia fucoidea Ptychocoleus corticalis (Lehm. & Lindenb.) Trevis. = Frullanoides corticalis P. polycarpus (Nees) Trevis. = Schiffneriolejeunea polycarpa Radula amazonica Spruce = R. javanica R. complanata var. a* propagulifera Nees = R. lindenbergiana R. longifolia Steph. = R. javanica R. macrostachya Lindenb. & Gottsche = R. javanica R. mollis Lindenb. & Gottsche = R. quadrata R. propagulifera Castle = R. quadrata Acknowledgments We would like to thank John Engel for access to his personal library, Jorn Hentschel, Tamas Pocs, Elena Reiner-Drehwald for their taxonomic advice and contributions, Alfons-SchaferVerwimp for calling our attention to several missing papers and some synonyms, and to Ana Seneca for help with literature search. We dedicate this paper to Dr. Ray Stotler and Dr. R. ramulina Taylor = R. voluta Barbara Crandall- Stotler for their dedication to bryology, and R. saccatiloba Steph. = R. fendleri we hope that this paper encourages research of liverworts and Rectolejeunea maxonii A. Evans = Lejeunea maxonii hornworts in the islands of the Caribbean, just as their own Riccardia bipinnata (Sw.) Trevis. = Aneura bipinnata research has encouraged research across generations. This work R. pinnatifida (Weber) Trevis. = R. chamedryfolia was supported, in part, by GBIF (Seed Money: 2007-41), and is Sarcomitrium pinnatifidum (Weber) Mitt. = Riccardia part of the Early Land Plants Today project. chamedryfolia Scapania grandis Bosw. = S. portoricensis Literature Cited S. portoricensis f. grossealata Herzog = S. portoricensis var. Ahonen, I. 2006. The taxonomic position of the genus portoricensis Nipponolejeunea Hatt. Journal of the Hattori Botanical S. portoricensis f. integrifolia Miill.Frib. = S. portoricensis var. portoricensis Schisma juniperinum (Sw.) Dumort. = Herbertus juniperoideus Sendtnera adunca (Dicks.) Gottsche & Rabenh. = Herbertus aduncus S. flagellifera (Hook.) Nees = Dendromastigophora flagellifera S. juniperina (Sw.) Nees = Herbertus juniperoideus S. juniperina a swartziana Gottsche = Herbertus juniperoideus Sphagnoecetis longiflora Taylor = Odontoschisma longiflorum S. prostrata (Sw.) Nees = Odontoschisma prostratum Stephanina pallens (Sw.) Kuntze = Radula pallens Stylolejeunea pililoba (Spruce) A. Evans = Lejeunea trinitensis S. trinitensis (Lindenb.) Fulford = Lejeunea trinitensis Symbiezidium brachiatum (Sw.) Trevis. = Marchesinia Laboratory 99: 319-342. Arnell, S. 1956. Hepaticae collected in Cuba and Hispaniola by E. L. Ekman. The Bryologist 59: 271-276. Martinique and in French Guiana in 1958 Tidskrift 53: 499-506. Asthana, G. & S. C. Srivastava. 2003. Indian Cololejeunea: A taxonomic study. Bryophytorum Bibliotheca 60: 1-155. Austin, C. F. 1876. Notes on Hepaticology. Botanical Bulletin, Hanover 1(7): 31-32. Bapna, K. R. & P. Kachroo. 2000. Hepaticology of India II. Himanshu Publications, Udaipur. Barbour, W. C. 1902. Porella L. The Bryologist 5: 32-36. Berke, P. 8c T. Beatley. 1995. Sustaining Jamaica's Forests: The brachiata S. subrotundum (Hook.) Trevis. = S. transversale Protected Area and Resource Conservation Project. S. vincentinum (Gottsche) Trevis. = S. transversale var. Environmental Management 19: 527-546. hookerianum Bischler, H. 1962a. The genus Calypogeja Raddi in Central and Taeniolejeunea appressa (A. Evans) Zwickel = Cololejeunea appressa South America I. Introduction and subgenera Mnioloma and Caracoma. Candollea 18: 19-51. This content downloaded from 107.0.125.4 on Wed, 10 Aug 2016 20:31:07 UTC All use subject to http://about.jstor.org/terms 84 The Bryologist H4(l)' 2011 South America II. Subgenus Calypogejay Acroradula. subgroups Section 1,8. 2 Acutilobulae. and Revue Br 3. Candollea 18: 53-93. Lichenologique 33: 185-210. Acroradula. et Section 9. Densifoliae. Revue Bryo Centrale et Meridionale. Revue Bryologique Lichenologique 33: 15-179. Lichenologique 33: 328-398. Section 10. Ampliatae. Revue Bry Centrale et Meridionale. II. RevueAcroradula. Bryologique et Lichenologique 34: 1-81. Lichenologique 35 "1967": 95-134. Acroradula. Section 11. Complanatae. Revue B Amerique. Nova Hedwigia 17: 265-350. Lichenologique 35 "1967": 1-94. d'Asie et d'Oceanie. Journal of the Hattori Botanical taxonomic revision of the genus. Revue B Laboratory 45: 25-79. Lichenologique 36 "1968": 5-44. Chai, S.-L., E. Tanner 8c K. McLaren Chai. americains. Revue Bryologique et Lichenologique 45: forest clearance and fragmentation pre- a 255-333. Park establishment: The case of a Jamaic rainforest. Biological Conservation 142: 2 Lejeuneaceae III. A historical account of Lejeunea cucullata Clark, L. 1956. Frullania gibbosa. The Bryo (Reinwardt, Blume & Nees) Nees and its varieties. Nova Hedwigia 3 "1961": 445-462. The Bryologist 61: 67. Bischler-Causse, H., S. R. Gradstein, S. Jovet-Ast, D. G. Long & N. Salazar Allen. 2005. Marchantiidae. Flora Neotropica Bryologist 47: 62-66. Monographs 97: 1-267. Bizot, M. & T. Pocs. 1974. East African Bryophytes I. Egri 47: 108-112. tanarkepzo foiskola tudomanyos kozlemenyei. Acta Academiae Paedagogicae Agriensis 12: 383-449. 187-189. Bonner, C. E. B. 1962-1977. Index Hepaticarum. Pars II-VIIa. J. Cramer, Weinheim. 82-85. Boswell, H. 1887. Jamaica mosses and hepaticae. Journal of Botany 25: 45-50. Middle America. The Browne, P. 1756. The civil and natural history of Jamaica in three parts. London503. Bry 51: 239-242. Burghardt, M. 8c S. R. Gradstein. 2008. A revision of Tylimanthus (Acrobolbaceae,55:Marchantiophyta) in tropical 186-189. America, Africa and Macaronesia. Fieldiana, Botany, n.s. Costa, D. P. 2008. Metzgeriaceae. Flora Neotropica 102: 1-169. 47: 199-210. Crandall-Stotler, B. J. 8c W. R. Guerke. 1980. Developmental Campbell, E. O. 1987. Monoclea (Hepaticae); distribution and anatomy of Jubula Dum. (Hepaticae). The Bryologist 83: number of species. The Bryologist 90: 371-373. 179-201. Castle, H. 1937. A revision of the genus Radula. Introduction and part I. Subgenus Cladoradula. Annales Bryologiciclassification 9 of the Marchantiophyta. Edinbu "1936": 13-56. Botany 66: 155-198. Crosby, M. R. 8c J. J. Engel. 2006. Index of H Acroradula. Annales Bryologici 12: 21-47. Botanical Laboratory, Nichinan. Damsholt, K. 2002. Illustrated flora of Nordic Subgenus Acroradula . Section 3. Dichotomae. Journal the hornworts. Nordicof Bryological Society, Lun Hattori Botanical Laboratory 21Dauphin, "1959": 1-52. G. 2003. Ceratolejeunea. Flora Neot Monographs 90: 1-86. unrecognized species of the leafy hepatics. Revue Bryologique et Lichenologique 29: 10-25. Anthocerotae. Tropical Bryology 26: 1411998. Additions to Acroradula. Section 6. Saccatae. Revue Bryologique etthe hepatic flora of Costa Lichenologique 32: 1-48. Lindbergia 23: 74-80. This content downloaded from 107.0.125.4 on Wed, 10 Aug 2016 20:31:07 UTC All use subject to http://about.jstor.org/terms Soderstrom et al.: Liverworts and hornworts of Jamaica 85 Douin, C. 1920. La famille des Cephaloziellacees. Memoires de la Societe Botanique de France 29: 1-90. Leptocolea and Aphanolejeunea. Bulletin of Botanical Club 38: 251-286. Dugas, M. 1929. Contribution a l'etude du genre " Plagiochila" Dum. Annales des Sciences Naturelles, Botaniqe, ser. 10 11: 1-199. Bulletin of the Torrey Botanical Club 39 Easa, P. S. 2003. Biodiversity documentation for Kerala. Part 4 Torrey Botanical Club 39: 603-611. Bryophytes. Kerala Forest Research Handbook 17: 1-47. Eggers, J., J.-P. Frahm 8c R. A. Pursell. 1998. New bryophyte taxon records for tropical coounties II. Tropical Bryology Bryologist 15: 54-63. 14: 81-84. species. Bulletin of the Torrey Botanical C records for tropical countries 5. Tropical Bryology 25: 19-23. Transactions of the Connecticut Academ Sciences 21: 253-261. bryophyte records for the tropical countries IV. Tropical Bryology 20: 97-100. Journal of Botany 5: 131-150. Engel, J. J. 1976. Metzgeria hamata Lindb., an illegitimate name of Hepaticae. Lindbergia 3: 219-220. Torrey Botanical Club 46: 167-182. Brunswick Peninsula (Strait of Magellan) Hepaticae and Botanical Club 48: 1 Bulletin of the Torrey Anthocerotae. Fieldiana, Botany 41: 1-319. Symphyogyna. Transactions of the Connect Arts and Sciencesof 28: 301-353. Krunodiplophyllum Grolle and a revision the Australasian species of Diplophyllum (Dum.) Dum. (Scapaniaceae, Jungermanniales). Journal of the Hattori Botanical Bryologist 43: 1-4. Laboratory 84: 241-283. Forrest, L. L., E. C. Davis, D. G. Long, B. J. Cr Evans, A. W. 1904. Hepaticae of Puerto Rico. Clark 8c M. L.IV. Hollingsworth. 2006. Unrave Odontolejeunea, Cyclolejeunea and Prionolejeunea. Bulletin evolutionary history of the liverworts (Mar of the Torrey Botanical Club 31: 183-226. multiple taxa, genomes and analyses. The B 303-334. Bulletin of the Torrey Botanical Club 273-290. Frye, T. C. &32: L. Clark. 1945. Hepaticae of North America part III. University of Washington Publications in Biology 6: 337-564. Rectolejeunea, Cystolejeunea , and Pycnolejeunea. Bulletin of the Torrey Botanical Club 33: 1-25. University Bryologist 10: 24-30. of Washingto 565-733. Neurolejeunea , Omphalanthus , and Lopholejeunea. Bulletin University of Washingt 735-1022. of the Torrey Botanical Club 34: 1-34. Fulford, M. 1941. Studies on American Hepaticae. II. A new Marchesinia , Mastigolejeunea , Caudalejeuttea, species of Bazzania and from the West Indies. The Bryologist 145-146. Bryopteris. Bulletin of the Torrey44:Botanical Club 34: 533-568. the genus Symbiezidium. Lloydia 5: 293PtychocoleuSy Archilejeunea , Leucolejeunea and Anoplolejeunea. Bulletin of the Torrey Botanical Club 35: Ceratolejeunea. Brittonia 5: 368-403. 155-179. America. Annales Cryptogamici et Phyt 1-175. Torrey Botanical Club 35: 371-389. Bryologist 13: 33-36. supplement to 'The genus Bazzania in C America". Bulletin of the Torrey Botan Botany. Oxford 24: 271-303. 308-341. This content downloaded from 107.0.125.4 on Wed, 10 Aug 2016 20:31:07 UTC All use subject to http://about.jstor.org/terms 86 The Bryologist 114(l) : 2011 I. Memoirs of the New York Botanical Garden 11: 1-172. Central French Guiana Part 4. Liverworts and Hornworts. Memoirs of the New York Botanical Garden 76 II. Memoirs of the New York Botanical Garden 11: Additions to the hepatic flora of Costa R 173-276. "1994": 73-86. III. Memoirs of the New York Botanical Garden 11: Symbiezidium Trevis. Nova Hedwigia, Beihe 277-392. IV. Memoirs of the New York Botanical Garden 11: Liverworts in the Tropics. Memoirs o Botanical Garden 45: 388-425. 395-535. Grolle, R. 1963. Monographic der Lebermoosgattung Mitt.. Nova Acta Leopoldina. Abhandlungen Hepaticae Elliottianae by Richard Leptoscyphus Spruce in 1895. Memoirs der Kaiserlich Leopoldinisch-Carolinisch Deutschen of the New York Botanical Garden 45: 264-278. Akademie der Naturforscher 25 "1962": 1-143. hundred and twenty-seven years after C. M. Gottsche. Repertorium 73: 78-89. Memoirs of the New York Botanical Garden 63: 1-86. the Hattori Botanical Laboratory 31: 20-49. forest in Puerto Rico, 11. The leafy hepaticae of Pico del Oeste. Journal of the Arnold Arboretum 51: 56-69. Repertorium 82: 1-100. Geissler, P. 8c H. Bischler. 1985-1990. Index Hepaticarum vol. 8/9-12. J. Cramer, Berlin. benachbarter Gebiete. Feddes Repertoriu Phragmicoma lehmanniana Nees and other species Cyclolejeunea. Wissenschaftliche Zeitschrift described in the genus Phramicoma (Studies in Schiller-Universitat Jena/Thuringen. Mate Lejeuneaceae subfam. Pthycanthoideae 23). Journal of the naturwissenschaftliche Reihe 33: 759-764. Hattori Botanical Laboratory 75: 201-209. Gottsche, C. M. 1857. Pugillus Novarum Hepaticarum e the Hattori Botanical Laboratory 69: 185-1 recensione Herbarii Musei Parisiensis, congestus. Annales des Sciences Naturelles, Botaniqe, ser. 4 8: 318-348. Plagiochila aerea Taylor 1846 (Hepaticae), fir Lycopodium pinnatum by Lamarck 1792. N Prodromus Florae Novo-Granatensis). Annales des Sciences 68: 511-526. Naturelles, Botaniqe, ser. 5 1: 95-198. Hepaticae and Anthocerotae of Europe and 1847. Synopsis Hepaticarum, Hamburg. Journal of Bryology 22: 103-140. Gradstein, S. R. 1974. Studies on Lejeuneaceae subfam. Ptychanthoideae. I. Nomenclature and taxonomy of 8c Reiner comb. nov. (Lej (Angstr.) Grolle Ptychocoleus , Acrolejeunea and Schiffneriolejeunea. Journal the neotropics. Journal of Bryology 19: 78 of the Hattori Botanical Laboratory 38: 327-336. (Lejeuneaceae) including t Schiffn. Nova Hedwigia, Beiheft 80:new 195-220. species from the pa Bryology 21: 31-45. Brachiolejeuneaceae. Flora Neotropica Monographs 62: 1-217. Metzgeria species (Hepaticae): Jungermannia 1753, /. linearis Sw. 1788 and /. pubescens Sc Archilejeunea (Spruce) Schiffn. Nova Hedwigia, Beiheft 23: 80: 119-121. Cryptogamie, Bryologie 89-112. Evans (Lejeuneaceae, Hepaticae). Taxon 50 Brazil. Memoirs fo the New York Botanical Garden 87: 1-316. 1067-1074. Guerke, W. R. 1978. A monograph of the genus Jubula Dumortier. Bryophytorum Bibliotheca 17: 1-118. Hampe, E. 1852. Hepaticae Oerstedianae. Linnaea 24 "1851": bryophytes of tropical America. Memoirs of the New York Botanical Garden 86: 1-577. 300-304, 640-641. This content downloaded from 107.0.125.4 on Wed, 10 Aug 2016 20:31:07 UTC All use subject to http://about.jstor.org/terms Soderstrom et al.: Liverworts and hornworts of Jamaica 87 ribosomal DNA. Plant Systematics and Evol Hatcher, R. E. 1958. The genus Trichocolea in North, Central and South America - (Hepaticae). Lloydia 20 "1957": 139-185. 221-230. Hitchcock, A. S. 1898. List of the cryptogams collected in the Bahamas, Jamaica and Grand Cayman. Annual Report of the Missouri Botanical Garden 9: 111-120. Hedwigia 2: 573-608. Hattori, S. 1977. Dr. H. Hurlichmann's of Newportoricensis Hampe et Hong, W.collections C. 1988. Studies on Scapania Caledonian Frullaniaceae. Journal of the Hattori Botanical Gott. of Latin America. Lindbergia 14: 69-72. Laboratory 43: 409-438. Ilkiu-Borges, A. L. 2005. A taxonomic revision of Echinocolea Haynes, C. C. 1907. Two new species of AytoniaHepaticae). from Jamaica. (Lejeuneaceae, Nova Hedwigia 80: 45-71. Bulletin of the Torrey Botanical Club 34: 57-60. Prionolejeunea (Lejeuneaceae, Jungerman Marchantiales. North American Flora 14: 1-66. Cuvillier Verlag, Gottingen. He, X.-L. 1997. Type studies on Pycnolejeunea (Lejeuneaceae, Inoue, H. 1966. A monograph of the hepat Hepaticae), IV. Annales Botanci Fennici 34: 65-74. Spruce. Journal of the Hattori Botanical 171-213. Pycnolejeunea (Lejeuneaceae, Hepaticae). Acta Botanica Jovet-Ast, S. 1948. Hepatiques des Antilles fran9aisee recoltees Fennica 163: 1-77. para P. et V. Allorge en 1936 I. Revue Bryologique et Lichenologique 16 "1947": 17-46. Lejeuneaceae (Hepaticae) from the Neotropics, Madagascar and the Seychelles. Annales Botanci Fennici 38: 25-44. P. et V. Allorge en 1936 II. Revue Bryolog Heinrichs, J. 2002. A taxonomic revision of Plagiochila sect. Lichenologique 17 "1948": 24-34. Hylacoetes , sect. Adiantoidea and sect. Fuscoluteae in the Neotropics with a preliminary subdivision of neotropical geographique. Compte Rendu Sommaire Plagiochilaceae into nine lineages. Bryophytorum Societe de Biogeographie 36 "1959": 12 Bibliotheca 58: 1-184. Kruijt, R. C. 1985. A preliminary revision o Dicranolejeunea (Spruce) Schiffn.. Nova Crispatae and sect. Hypnoides (Hepaticae) in the 80: 155-163. Neotropics. I. Plagiochila disticha , P. montagnei and P. raddiana. Nova Hedwigia 70: 161-184. Acanthocoleus. Bryophytorum Bibliotheca neotropical species of Plagiochila (Du subfam. Ptychanhoideae X. On Brachiole described by Olof Swartz. (Nees) Kruijt 8c Gradstein comb, nov (H Botanical Laboratory 85: 1-32 (Hepaticae) Hattori Hedwigia 43: 299-309. Kuwahara, Y. 1979. Notes on the genus St Plagiochila killarniensis Pearson and P. bifaria (Sw.) Bryologist 82: 613-615. Lindenb. (Hepaticae). Journal of Bryology 20: 495-497. Metzgeria collected by Dr. Antoine M H. Anton. 2001. Plagiochila rutilans Andean (Hepaticae): a poorly and 1973. Proceedings, Koninklijke Nederla known species from tropical America. The Bryologist 104: van Wetenschappen. Series C, biological an 350-361. sciences 85(3): 357-380. Proschold. 2002a. moritziana , sequence and P. The systematic position of with a wide geographical No trichostoma , and P. distribution. deflexa ba variation lipophilic ribosomal metabolites. DNA, The Br genus Metzgeria Raddi (Hepaticae). Hikob Molecular, broad The nuclear secondary 189-203. a 603-616. of Metzgeriaceae Japan, East Himalay morphological, and from phytochemical e species Bryologist Asia, of and Plagiochila Tropical Oceania.bifaria Nova Hedw concept ( 107: 273-285. 28-40. 2002b. Plagiochila virginica A. EvansBryophytorum rather than P. dubia Bibliotheca 28: 1-254. Lindenb. 8c Gottsche occurs in Macaronesia; in Lehmann, placement J. G. C. 1832. Novarum et minu sect. Contigua Carl is supported by ITS sequences of nuclear stirpium pugillus IV. Hamburg. This content downloaded from 107.0.125.4 on Wed, 10 Aug 2016 20:31:07 UTC All use subject to http://about.jstor.org/terms 88 The Bryologist 114(1): 2011 besonderer Beziehung auf Schlesien u VI addita enumeratione plantarum mit omnium in his pugillis descriptarum. Hamburg. Ortlichkeiten des Riesengebirge, vol. 3. Br Nieuwkoop, J. 8c T. Arts. 1995. Additions to t of Madeira. Lindbergia 20: 35-39. VII addita enumeratione plantarum omnium in his pugillis descriptarum. Hamburg. Pagan, F. M. 1939. A preliminary list of the Rico including Vieques and Mona Island. 42: 1-12, 37-50, 71-82. X addita enumeratione plantarum omnium in his pugillis descriptarum. Hamburg. 45: Lindberg, S. O. 1883. Kritisk granskningBryologist af mossorna uti76-110. Dillenii Historia muscorum. J. C. Frenckell 8c Paton, Son, J. Helsingfors. A. 1999. The liverwort flora of the B Books, Colchester. Lindenberg, J. B. C. 1839. Species hepaticarum. Fasc. 1. Henry & Cohen, Bonn. Patsch, R., J. Hentschel, R. Linares-Palomino, Heinrichs. 2010. Diversification and taxono Bonn. Liverwort Jubula Dumort. (Jungermannio inferred from nuclear and chloroplast DN Bonn. Systematic Botany 35: 6-12. Pearson, W. H. 1931. Notes on a collection of Annales 4: 95-112. Iungermanniae Trichomanoideae Jamaica. Lepidozia. HenryBryologici 8c Cohen, Bonn. Perold, S. M. 1999. Flora of Southern Africa. H 1: Marchantiopsida. Fascicle 1: Marchantii Henry 8c Lucking, Cohen, A. 1995. Botanical Institute, Pretoria. Bonn. Piippo, S. 1986. A monograph of Lepidolejeu Diversitat und Mikrohabita Luteolejeunea Hepaticae). Ac in einem(Lejeuneaceae: tropischen Reg epiphyller Moose Rica Fennica 132: 1-69. besonderer Beriicksichtigung unter d Ulm, 211. Pocs. 2002. Hepatic and hornwort flora o Lunan, J. 1814. Hortus Jamaicensis. St. Jago de la Vega Gazette.. Annales Botanci Fennici 39: 101-128. Meenks, J. L. D. 1987. Studies on Colombian cryptogams XXVIII. A guide to the tropical Andean species of Riccardia Pocs, T. 8c A. Bernecker. 2009. Knowledge of Aphanolejeunea (Hepaticae). Journal of the Hattori Botanical Laboratory (Jungermanniopsida) after 25 years. Polish Botanical 62: 161-182. Journal 54: 1-11. Miller, H. A., H. O. Whittier 8c B. A. Whittier. 1983. Raeuschel, E. A. 1797. Nomenclator botanicus omnes plantas ab illustr. Carolo a Linne descriptas aliisque botanicis temporis Prodromus Florae Hepaticarum Polynesiae. Bryophytorum Bibliotheca 25: 1-423. Mittermeier, R. A., P. R. Gil, M. Hoffmann, J. Pilgrim, T. recentioris detectas enumerans. I.G. Feind; Lipsiae, 414 pp. Reimers, H. 1933. Revision der Lebermoosgattung Brooks, C. G. Mittermeier, J. Lamoreux 8c G. A. B. da Micropterygium. Hedwigia 73: 133-204. Fonseca. 2004. Hotspots Revisited: Earth's Biologically Reiner-Drehwald, M. E. 1993. Las Lejeuneaceae (Hepaticae) de Richest and Most Endangered Ecoregions. Mexico City Misiones, Argentina I. Las especies holostipas. Tropical (Mexico): CEMEX. Bryology 7: 13-45. Mizutani, M. 1984. Notes on the Lejeuneaceae. 8. Japanese species of the subgenus Taeniolejeunea of the genus Noreste de Argentina. Tropical Bryology 9 Cololejeunea. Journal of the Hattori Botanical Laboratory 57: 153-170. V. Cheiblejeunea y Lepidolejeunea. Tropical Bryology Miiller, J., J. Heinrich 8c S. R. Gradstein. 1999. A revision of Plagiochila sect. Plagiochila in the Neotropics. The of Latin America. Bryophytorum Bibliothec Bryologist 102: 729-746. Nees von Esenbeck, C. G. 1830. Enumeratio plantarum Argentina VI. Lejeunea y Taxilejeunea. Tr 19: 81-131. cryptogamicarum Javae et insularum adiacentium, quas a Blumio et Reinwardtio collectas describi edique curavit gen. nov. (Lejeuneaceae, Hepaticae). Nova Christ. Godfr. Nees Ab Esenbeck professor vratislaviensis. Fasciculus prior, hepaticas complectens, ab editore 447-464. illustratis. Grass, Barth 8c Co., Breslau, 86 pp. distributed species of tropical America. Cryp cryptogamicarum Javae. Linnaea 6:Bryologie 602-623. 30: 329-336. This content downloaded from 107.0.125.4 on Wed, 10 Aug 2016 20:31:07 UTC All use subject to http://about.jstor.org/terms Soderstrom et al.: Liverworts and hornworts of Jamaica 89 Reyes, D. M. 1983. El genero Diplasiolejeunea en Cuba. Acta botanica Academiae Scientiarum Hungarica 28 "1982": America, vol. III. Columbia University P 145-180. Rico, R. & T. Pocs. 2004. Briofitos de las tierras altas de la Phytologia 45: 415-437. Guayana Venezuelana: hepaticas del Roraima-tepui I. America, vol. IV. Columbia University P Cryptogamie, Bryologie 25: 249-269. Rycroft, D. S., W. J. Cole, J. Heinrichs, H. Groth, C. Renker & T. Proschold. 2002. Phytochemical, morphological, and Pages 463-626. in: Schuster, R. M. (ed.), molecular evidence for the occurrence of the neotropical Bryology, vol. 1. Hattori Botanical Labora liverwort Plagiochila stricta in the Canary Islands, new to Macaronesia. The Bryologist 105: 363-372. America. V. Columbia University Press, Schafer-Verwimp, A. 1989. New or interesting records of Brazilian Bryophytes, II. Journal of the Hattori Botanical America. VI. Columbia University Press, Laboratory 67: 313-321. Lejeuneaceae (part 2). Journal of the Ha Dominica, West Indies. Haussknechtia, Beiheft 72: 9: 317-332. Laboratory 163-359. Marchantiopsida) in the Tropical Andes, Drepanolejeunea with description (Spr) Schiffn. Nova Hed of two new species. Cryptogamie, Bryologie 25: 3-17. subfam. Zoopsidoideae (3): Zoopsidella. 101-150. the Dominican Republic, West Indies. Acta Botanica Hungarica 51: 367-425. Beiheft 118: 1-524. bryophyte synonyms flora in the (Lejeuneaceae). Schiffner, V. Expedition nach of & West Indies, 119: 1-606. genera Beiheft Diplasiolejeunea and and new Lejeunea Cryptogamie, Bryologie 30: 357-375. Sim-Sim, M., M. G. Esquivel, S. Fontinha & S. Carvalho. 2004. S. der W. Arnell. Plagiochila 1964. stricta Lindenb. Ergebnisse new to Madeira: der morphological Botanisch kaiserlichen Akademie der Wissenschaften and molecular evidence. Nova Hedwigia 79: 497-505. Sudbrasilien Hepaticae Guadeloupe, 1901. So, II. Thallophyta Bryophyta. M. L. 2002. Metzgeria (Hepaticae) und in Australasia and the (Lebermoose). Osterreichische Akademie Pacific. New Zealand Journal of Botany 40: 603-627. der Wissenschaften. Mathematisch-Naturwissenschaftliche Klasse. Denkschriften 111: 1-156. Schumacker, R. & J. Vana. 1999. Two new liverworts for America. New Zealand Journal of Botan Soderstrom, L., A. Hagborg, M. von Konrat Europe in Macaronesia: Odontoschisma prostratum (Sw.) Trevis. on the Azores and Jungermannia callithrix Lindenb. Early Land Plants Today: Liverwort check Fieldiana 47: 105-130. & Gottsche on the Azores and Madeira. Tropical Bryology 17: 115-127. and Anthocerotae in Europe and Macarone Schuster, R. M. 1960. A monograph of the Nearctic 427. Cryptogamie, Bryologie 28: 299-350. Plagiochilaceae. Part. III. Sectio Contiguae to Conclusion. Stephani, F. 1886. Hepaticae species novae ve American Midland Naturalist 63: 1-130. V. Hedwigia 25: 5-9. subgenera Microlejeunea and Chaetolejeunea. Journal of the Lindenberg. Hedwigia 29: 1-23. Hattori Botanical Laboratory 25: 1-80. 232-253. Drepanolejeunea, and Leptolejeunea. Journal of the Elisha Mitchell Scientific Society 83: 192-229. Stotler, R. E. 1970. The genus Frullania s Latin America. Nova Hedwigia 18 "19 America, vol. 2. Columbia University Swails,Press, L. F. New 1970.York. The genus Porella in L Hedwigia 19: 201-291. Lejeuneaceae from Dominica and Jamaica. Bulletin ofGenera the Swartz, O. 1788. Nova et Species Torrey Botanical Club 97 "1970": 336-352. Prodromus descriptionum vegetabilium incognitorum quae sub itinere in Indiam "1971": 121-265. annis 1783-1787. Uppsala. This content downloaded from 107.0.125.4 on Wed, 10 Aug 2016 20:31:07 UTC All use subject to http://about.jstor.org/terms 90 The Bryologist 114(1): 2011 and Lophoziaceae (Hepaticae). Annales Bo 263-290. sive descriptiones plantarum in Prodromo recensitarum, vol 3. J. Palm, Erlingen. Vital, D. M. & S. R. Visnadi. 2000. New records and notes on Taylor, T. 1846. New Hepaticae. London of Botany 5: BrazilianJournal bryophytes. Journal of the Hattori Botanical 258-284, 365-417. Laboratory 88: 191-197. von Konrat, M., L. Soderstrom & A. Hagborg. 2010. The Early cryptogamic plants from Jamaica. Annals and Magazine of A community-driven Land Plants Today project (ELPT): Natural History 20: 379-381. effort and a new partnership with Phytotaxa. Phytotaxa XX: XX-XX. cryptogamic plants from Jamaica.Weber, Transactions of the F. 1815. Historia muscorum hepaticarum prodromus , Botanical Society of Edinburgh 3: 23-25. Kiel. Teeuwen, M. 1989. A revision of the genus Odontolejeunea Wilson, R., S. R. Gradstein, H. Schneider 8c J. Heinrichs. 2007. (Spruce) Schiffn. (Lejeuneaceae: Hepaticae). Nova Unravelling the phylogeny of Lejeuneaceae Hedwigia 32: 607-622. (Jungermanniopsida): evidence for four lineages. Molecular Thiers, B. M. 1985. Branching in Lejeuneaceae III. Nova Phylogenetics and Evolution 43: 270-282. Hedwigia, Beiheft: 80: 31-61. Winkler, S. 1968. Die epiphyllen Moose der Nebelwalder von El Salvador, C. A. Revue Bryologique et Lichenologique 35 Lejeuneoideae. Nova Hedwigia 42: 237-275. "1967": 303-369. Tixier, P. 1985. Contribution a la Connaissance des Yamada, K. 1980. Notes on the type specimens of Radula taxa Cololejeunoideae. Bryophytorum from Bibliotheca 27: 1-439. Latin America (1). Journal of the Hattori Botanical Uribe, J. & S. R. Gradstein. 2003. Type studies on Frullania Laboratory 48: 243-257. subgenus Meteoriopsis (Hepaticae). I. The lectotypification of the genus Frullania , F. subgen. Thyopsiella and F. subgen. Latin America (2). Journal of the Hattori Bot Meteoriopsis , and some species transferred from subgen. Laboratory 49: 385-398. Meteoriopsis to subgen. Thyopsiella. Cryptogamie, Bryologie 24: 193-207. Latin America (3). Journal of the Hattori Bot Van Reenen, G. B. A. 1982. Studies on Colombian cryptogams Laboratory 52: 449-463. XII. High Andean species of Herbertus S. F. Gray (Hepaticae). Lindbergia 8: 110-120. Latin America. 4. Journal of the Hattori Botan Van Zanten, B. O. & S. R. Gradstein. 1988. Experimental Laboratory 62: 289-298. dispersal geography of neotropical liverworts. Nova Hedwigia, Beiheft 90: 41-94. Hattori Botanical Laboratory 65: 379-390. Vana, J. 1974. Studien uber die Jungermannioideae (Hepaticae). 4. Jungermannia subg. Plectocolea und subg. Latin America 6. Journal of the Hattori Botani Solenostoma: Allgemeines, slid- und mittelamerikanische Laboratory 73: 125-137. Arten. Folia Geobotanica et Phytotaxonomica 9: 179-208. (Radulaceae; Hepaticae) made by Marko L (Hepaticae). 6. Jungermannia subg. Solenostoma : Journal of the Hattori Botanical Laborator Europaische und nordamerikanische Arten. Folia 257-265. Geobotanica et Phytotaxonomica 9 "1974": 369-423. Yano, O. 1987. Additions to the Brazilian Hepaticae. The Bryologist 90: 374-375. (Hepaticae). 7. Jungermannia subg. Plectocolea : Europaische und nordamerikanische Arten. Folia Geobotanica et Phytotaxonomica 10: 67-99. estados do Brasil. Acta Amazonica 34: 559-576. Yuzawa, Y. 1991. A monograph of subgen. Chonanthelia of gen. Frullania of the world. Journal of the Hattori preliminary information. Pages 99-106. In: J.(Hepaticae) Vana (ed.), Laboratory 70: 181-291. Proceedings of the third meeting of the Botanical bryologists from Zhu, R.-L. 1995. Notes on some species of the genus Central and East Europe. Univerzita Karlova, Praha. Cololejeunea (Lejeuneaceae, Hepaticae) in China. Journal of the Hattoriof Botanical Laboratory 78: 83-109. special reference to the BRYOTROP transect Mt. Kinabalu. XV. Lophoziaceae (Hepaticae). Willdenowia 20: 199-219. (Spruce) Schiffn. (Lejeuneaceae, Hepaticae) Monographs in Systematic Botany from th Peninsula, Papua New Guinea. XXXI. Cephaloziaceae Botanical Garden 74: 1-98. subfam. Allobielloideae, Cephaloziellaceae, Antheliaceae This content downloaded from 107.0.125.4 on Wed, 10 Aug 2016 20:31:07 UTC All use subject to http://about.jstor.org/terms Soderstrom et al.: Liverworts and hornworts of Jamaica 91 distribution of Acanthocoleus yoshinganus and Lejeunea Ceratolejeunea (Lejeuneaceae, Hepaticae) in papilionacea (Hepaticae, Lejeuneaceae). Annales Botanci Cryptogamie, Bryologie 26: 91-96. Fennici 38: 329-336. ms received February 5, 2010; accepted June 8 Nova Hedwigia, Beiheft 121: 1-418. This content downloaded from 107.0.125.4 on Wed, 10 Aug 2016 20:31:07 UTC All use subject to http://about.jstor.org/terms

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