Tropical Bryology 28:15-47, 2006 15 Checklist of Hawaiian Anthocerotes and Hepatics G. W. Staples & C. T. Imada Hawaii Biological Survey, Bishop Museum, 1525 Bernice St., Honolulu, Hawai‘i, 96817-2704 U.S.A. Abstract. A literature-based checklist of Hawaiian hepatics and anthocerotes is presented. Geographic coverage includes the eight main Hawaiian Islands; the northwestern Hawaiian Islands are excluded. The checklist is alphabetically ordered by scientific names; the family is noted for each genus. Synonyms and misapplied names are cross-referenced to the accepted names. A bibliography of supporting references is included. Introduction The Hawaii Biological Survey (HBS) was established as a program of Bishop Museum by the Hawaii State Legislature in 1992, specifically to create and maintain accurate inventories of all species present in the Hawaiian Islands, and to document their presence with voucher specimen collections and bibliographies of relevant scientific literature. In the ensuing fourteen years, comprehensive taxonomic inventories have been compiled for many groups of organisms, both native and alien, that comprise the Hawaiian biota. Vascular plants (Wagner et al. 1999; Palmer 2003) and marine algae (Abbott 1999; Abbott & Huisman 2004) have been the primary focus of botanical inventory efforts; recent research has produced checklists and bibliographies for non-marine algae (Sherwood 2004) and mosses (Staples et al. 2004). Still much needed are similar checklists for Hawaiian Cyanophyta (blue-green algae), fungi, and lichens. This checklist fills one of the gaps in our knowledge of Hawaiian terrestrial plants—the hepatics and anthocerotes. These ongoing inventories will help the Hawaii Biological Survey to fulfill its mission by systematically compiling literature-based inventories for each of these groups of organisms. The products also provide a foundation for undertaking revisionary studies and floristic research. Modern comprehensive floristic accounts that incorporate the latest revisionary studies for all Hawaiian bryophytes are greatly needed. Genesis of the current checklist This checklist of Hawaiian hepatics and anthocerotes began as a list of names and island distributions for Hawaiian taxa extracted from the Prodromus florae hepaticarum polynesiae (Miller et al. 1983), which covers much of the Pacific, although the title suggests only Polynesia. Initial data extraction and creation of an electronic checklist were undertaken by HBS staff members with funding from the USDA-PLANTS database project in 1998. We then began making TROPICAL BRYOLOGY 28 (2006) preliminary updates to the taxonomy and distribution information while simultaneously compiling a bibliography of literature that supports the names in current use. Further USDA funding (2004–2006) allowed much more extensive literature searching that enabled assembly of taxonomic information, revision of nomenclature, addition of new taxa and distributional records, and integration of all this new information into the developing checklist. In the final version of the checklist, taxonomic decisions were made by Staples, with review and critical input from four independent bryological reviewers. Coverage It must be emphasized that this is a literature-based taxonomic checklist, not a specimen-based inventory. It is compiled primarily from literature published during the second half of the twentieth century to the present (2006). The checklist attempts to account for all scientific names in use for Hawaiian hepatics and anthocerotes in the latter decades of the twentieth century; it is not a comprehensive nomenclator for all names that have ever been published for, or applied to, Hawaiian hepatics and anthocerotes. No nomenclatural novelties or taxonomic changes are initiated in this checklist. Geographically, coverage focuses on the eight main Hawaiian Islands. There are, as yet, no published reports for bryophytes from the uninhabited northwestern Hawaiian Islands, although a few collections were made there. In the last decades of the twentieth century there was a renaissance of bryological research worldwide, and as a result many revisions, floristic accounts, and several monographs have been published that impact Hawaiian hepatic and anthocerote nomenclature. Noteworthy among these are the recent works of M. L. So (So 2000, 2001a, 2002a, 2002b, 2003a, 2003b, 2004, 2005b), which are systematically overhauling the taxonomy of Hawaiian hepatics and providing a much better understanding of the species and their distribution. These regional and global studies have 16 STAPLES & IMADA: HAWAIIAN ANTHOCEROTES AND HEPATICS resulted in significant taxonomic changes, as well as reporting the discovery of several taxa new to the Islands, either rare natives or newly naturalized alien species. This checklist aims to summarize all this information in a concise format, as follows. Format Accepted scientific name The main body of the checklist is alphabetically organized by genus, species, and infraspecific taxon. Accepted names appear in bold italic type. Authors are included for the genera, species, and infraspecific epithets. Authors of hepatic and anthocerote genera have been standardized against the classification proposed in Bryophyte Biology (Shaw & Goffinet 2000). All author names are abbreviated according to Authors of Plant Names style (Brummitt & Powell 1992). References that explain the taxonomic concept or provide geographic distribution data are cited after the accepted taxon name. For nearly all taxa, Miller et al. (1983) is cited. Late in the compilation it was decided to cite selected earlier papers (Miller 1953, 1963, 1967; Miller & Scott 1960; Miller et al 1962) that precede the Prodromus florae hepaticarum polynesiae (Miller et al. 1983). This compensates for the absence of voucher specimen data in the Prodromus itself, which is an uncritical compilation of names and literature. The earlier papers frequently cite specimens that document taxonomic concepts and island distribution, and their incorporation here enables interested researchers to follow up on information by studying the specimens cited in them. The first mention of a genus includes the family assignment on the same line. Most genera are still placed in the same families recognized by Miller et al. (1983), but a few changes in family placement have been made since then. In such cases, we provide first the family assignment used by Miller et al. (1983), and mention alternative family placements afterward, with a reference to the source. Our principal source for genera assigned to each family were the chapters in Bryophyte Biology (Shaw & Goffinet 2000) dealing with hornwort classification (Renzaglia & Vaughn 2000) and hepatic classification (Crandall-Stotler & Stotler 2000). In a few cases we could not locate generic names in Bryophyte Biology and these names were searched in the bryological nomenclator included in the W3MOST database of bryophytes (http://mobot.mobot.org/W3T/Search/most.html) on the TROPICOS website. Although coverage for hepatics in the latter source proved to be spotty, we were able to verify family assignments for some of the problem genera there. Synonyms and misapplied names (in italics) follow the accepted name in paragraph style, alphabetically ordered. These are limited primarily to names in use for Hawaiian hepatics and anthocerotes in the latter part of the twentieth century, although a few names from other parts of the world that enter into the nomenclature for particular taxa have been included to enable tracking names through the literature. Basionyms are not distinguished from synonyms and are alphabetized accordingly. Status Following the taxonomy is a column indicating status of the taxon in the Hawaiian flora: end indicates an endemic, ind indicates indigenous, and nat signifies naturalized. A ? following any of these signifies uncertainty about the status of the taxon in the Hawaiian flora. While some research has been devoted to the mode of introduction of pteridophytes and angiosperms to Hawai‘i, resulting in a list of species likely to have been introduced by Polynesian voyagers that settled the archipelago and another (much larger) list of plants that arrived following European contact, little attention has been devoted to bryophytes in this regard. A few hepatics collected for the first time late in the twentieth century are almost certainly alien species that have naturalized or are in the process of doing so. The authors have used their best judgment in cases where the status is not clear and appended a ? to whatever status we assigned, to draw attention to the species and invite investigation from specialists. Island distribution Following the status category is a summary of geographic distribution on the eight main Hawaiian Islands. The island name abbreviations are explained at the beginning of the checklist. An island abbreviation appears in the row to the right of the taxon name to signify that the taxon is reported in the literature from that island. Absence of the island name abbreviation indicates the species is not (yet) known from that island. Given the inadequate state of bryological collecting in the Hawaiian Islands, many new island distribution records are sure to be found in the future. And rare taxa, known only from types or historical specimens, will be recollected and their distributions better understood. Nearly all the island distributions are abstracted from the literature cited in brackets following the accepted name. In some case, Miller’s earlier papers provide more detailed specimen citation than the Prodromus florae hepaticarum polynesiae compilation; in such cases, the earlier papers are cited as well. And in a few cases (e.g., Plagiochila) specimens determined by specialists subsequent to their published works added new island distribution data; in such cases the relevant voucher is cited in brackets following the accepted name. Synonym list Following the main body of the checklist, an alphabetical list of all synonyms and misapplied names is provided. This list cross-references every synonym or misapplied name to the main entry found under the scientific name accepted in this checklist. TROPICAL BRYOLOGY 28 (2006) STAPLES & IMADA: HAWAIIAN ANTHOCEROTES AND HEPATICS References The checklist concludes with a bibliography of the references cited in the checklist; more than 230 bryological titles were consulted in the course of the project, but only those providing taxonomic or distribution data pertinent to this checklist are included here. Anomalies and incomplete records Numerous hepatic and anthocerote names without any island distribution are included, based on (mostly) older literature that reports the species from the Hawaiian (or sometimes Sandwich) Islands without stating on which island the species was found. For example, the names Lejeunea owaihiensis Gottsche and L. hawaiiensis Gottsche ex Mitt. are both based on Menzies specimens collected in 1793 from the Hawaiian Islands, without locality. Both names are considered synonyms of Lepidolejeunea involuta (Gottsche) Grolle in this checklist, another name based on a type specimen reported for the Hawaiian Islands without island locality. Likewise, Marchesinia mittenii A.Evans is based on a Hillebrand collection from the Hawaiian Islands, without locality. Until more recent collections are studied, the island distribution of taxa such as these remains unknown. Particularly problematic in this regard are numerous taxa that Miller et al. (1983) reported from the Hawaiian Islands, without indication of a voucher specimen to support the record. These records have been removed to the end of the main checklist in a section called Dubious Records. A much smaller list of names can definitely be excluded from the Hawaiian flora; these are listed as Excluded Species. References supporting these exclusions are cited. Numerical summaries In compiling the following summaries we tabulated the families, genera, species, and infraspecific taxa from the checklists and took into account the status Hawaiian Anthocerote Flora in Summary Endemic Indigenous 0 1 Families 0 4 Genera 1 3 Species 1 3 Total named taxa* 17 accorded each species or infraspecific taxon. For simplicity, the taxa with questionable status were tabulated with those of known status in the same category. Thus nat + nat? = the number of naturalized species. Dubious taxa have been compiled in a separate column; excluded taxa have not been included in these summaries. Due to the confusion that would ensue from trying to accommodate all family classifications available for the Anthocerotes, we simply counted all hornworts in one family, Anthocerotaceae. Although the total number of taxa has declined slightly, the perceived level of endemism has declined more sharply. A number of species that were described as endemic to the Hawaiian Islands were latter found to be synonyms of widespread tropical species. This trend will likely continue as additional revisionary and floristic research further clarifies the status of Hawaiian hepatics. The way forward It is hoped that by bringing together the literature and providing an updated checklist of names in current use, one or more competent bryologists might be inspired to undertake preparation of a hepatic and anthocerote flora for the Hawaiian Islands. Researchers studying anthocerotes and hepatics are invited to request loans from, or visit in person, the Herbarium Pacificum (BISH). There are extensive Hawaiian (and Pacific) collections, including many types, in BISH that are seldom consulted by bryological researchers. One of the goals of publishing this checklist is to encourage bryophyte researchers to make fuller use of the BISH resources. That said, one caveat must be mentioned. H. A. Miller described a number of Hawaiian hepatic taxa in his papers on Hawaiian bryology (e.g., Miller 1953, 1963, 1967, inter alia) and for some of these taxa the holotypes are stated to be in BISH. In fact, no type material has ever been located in BISH for most of Miller’s taxa. Recently, Miller’s personal Alien – 2 2 2 subtotal 1 6 6 6 dubious – 0 2 2 Total 1 6 8 8 Alien 1 2 6 6 subtotal 29 65 180 182 dubious 3 6 30 31 Total 32 71 210 213 * species + infraspecific taxa Hawaiian Hepatic Flora in Summary Endemic 0 Families 0 Genera 102 Species 103 Total named taxa* TROPICAL BRYOLOGY 28 (2006) Indigenous 28 63 72 73 18 STAPLES & IMADA: HAWAIIAN ANTHOCEROTES AND HEPATICS herbarium was donated to Miami University, Oxford,Ohio (M. Vincent, pers. comm.), and the ongoing incorporation of this collection raises the possibility that these missing type specimens will be found and deposited in BISH. Researchers wishing to study type material of Miller’s Hawaiian hepatics are advised to contact both BISH and MU regarding the whereabouts of any type specimens, which remain missing as this checklist is being finalized. Until such time as a comprehensive Hawaiian hepatic flora is available, taxonomic coverage for Hawaiian hepatics will remain spotty. While revisionary studies including Hawaiian taxa have been published in recent decades for some genera—Herbertus, Jubula, Marchantia, Metzgeria, Plagiochasma, Plagiochila, Porella, Radula—and more are in progress for Aneura, Frullania, and Megaceros, among others, there are still many genera for which no modern taxonomic accounts include Hawaiian taxa. By far the majority of articles cited in support of this checklist make only passing mention of Hawaiian taxa. And that provides a direction in which future studies might proceed. Acknowledgments Funding for initial data compilation and for further data refinement was made available by the USDAPLANTS program; we thank Scott Petersen and Mark Skinner for their support and assistance. The Marks Botanical Endowment, Bishop Museum, supported finalizing the checklist for publication. Our Bishop Museum colleagues Katie Anderson and Karin Kami assisted with data entry and bibliographic compilation. Bishop Museum librarian Patrice Belcher obtained numerous critical references through interlibrary loan services. William Weber (COLO) graciously placed his files on Hawaiian hepatics and his own working checklist at our disposal. Reviews of the draft checklist by S. R. Gradstein (GOET), M. L. So (HKBU), S. L. Timme (KSP), and R. L. Zhu (HSNU) brought to light several relevant articles overlooked during the compilation as well as suggestions for its improvement. B. Thiers (NY) verified the single BISH specimen labelled Pleurozia gigantea, thereby documenting that the species genuinely occurs in Hawai‘i. The authors gratefully acknowledge all assistance rendered; any errors of fact or interpretation are the sole responsibility of the authors. References Abbott, I. A. 1999. Marine red algae of the Hawaiian Islands. Bishop MuseuPress, Honolulu. Abbott, I. A. & J. M. Huisman. 2004. Marinegreen and brown algae of the Hawaiian Islands. Bishop Museum PressHonolulu. Bharadwaj, D.C. 1975. 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Journal of the Hattori Botanical Laboratory 94: 159–177. So, M.-L., 2004. Studies on the type specimens of Odontoschisma (Hepaticae) in Asia and the Pacific. Journal of the Hattori Botanical Laboratory 95: 249–261. So, M.-L. 2005a. A synopsis of Radula (Radulaceae, Marchantiophyta) in New Zealand and Tasmania. Journal of the Hattori Botanical Laboratory 98: 149–174. So, M.-L. 2005b. Radula (Radulaceae, Marchantiophyta) in Hawaii. Journal of the Hattori Botanical Laboratory 98: 175–191. Staples, G. W., C. T. Imada, W. J. Hoe, & C. W. Smith. 2004. A revised checklist of Hawaiian mosses. Tropical Bryology 25: 35–69. Stephani, F. 1924. Species hepaticarum. 6: 16–562. Thiers, B. M. 1993. A monograph of Pleurozia (Hepaticae; Pleuroziaceae) Bryologist 96: 517– 554. Tixier, P. 1985. Contribution à la connaissance des Cololejeunoideae. Bryophytarum Bibliotheca 27: 439 pp. Váňa, J. 1975. Studien über die Jungermannioideae (Hepaticae). 9. Jungermannia Subg. Plectocolea and Subg. Solenostoma in Hawaii Ergänzungen und Synopsis der Gattung Jungermannia. Folia Geobotanica et Phytotaxonomica, Praha 10: 357– 382. Váňa, J., S. Piippo, & T. Koponen. 2005. Bryophyte flora of Hunan province, China. 6. Jungermanniaceae and Gymnomitriaceae (Hepaticae). Acta Botanica Fennica 178: 57–78. Wagner, W.L., Jr., D.R. Herbst, & S.H. Sohmer. 1999. Manual of the Flowering Plants of Hawai‘i. 2 vols. Bishop Museum Press and Univ. Hawaii Press. Honolulu. Yamada, K. 1979. A revision of Asian taxa of Radula, Hepaticae. Journal of the Hattori Botanical Laboratory 45: 201–322. Yamada, K. 1989. The bryophytes of Sabah (North Borneo) with special reference to the BRYOTROP transect of Mount Kinabalu. VIII. Radula (Radulaceae, Hepaticopsida). Willdenowia 19: 219–236. Yoshida, L. & C.W. Smith. 1976. Two recent thalloid liverworts introduced to Hawaii. Bulletin of the Pacific Tropical Botanical Garden 6(1): 18–20. Yuzawa, Y.1991. Monograph of subgenus Chonanthelia of gen. Frullania (Hepaticae) of the World. Journal of the Hattori Botanical Laboratory 70: 181–291. Zhu, R.-L. & S.R. Gradstein. 2005. Monograph of Lopholejeunea (Lejeuneaceae, Hepaticae) in Asia. Systematic Botany Monographs 74: 1–98. Zhu, R.-L. & M.-J. Lai. 2005. Cheilolejeunea decursiva (Sande Lac.) R.M.Schust. and Cheilolejeunea ventricosa (Schiffn.) X.L. He (Lejeuneaceae, Hepaticae), new to Australia. Journal of the Hattori Botanical Laboratory 98: 193–203. TROPICAL BRYOLOGY 28 (2006) STAPLES & IMADA: HAWAIIAN ANTHOCEROTES AND HEPATICS 21 Apendix 1. Anthocerotae of the Hawaiian Islands Island abbreviations: Ni–Ni‘ihau; K–Kaua‘i; O–O‘ahu; Mo–Moloka‘i; L–Läna‘i; M–Maui; Ka–Kahoolawe; H–Hawai‘i. FullName ANTHOCEROS L. (Anthocerotaceae) Anthoceros spongiosus Steph. [Miller et al. 1983] DENDROCEROS Nees (Anthocerotaceae, or Dendrocerotaceae [Renzaglia & Vaughn 2000*]) Dendroceros crispus (Sw.) Nees [Miller et al. 1983; Piippo 1993] Dendroceros brevicapsulus Steph. Dendroceros clintoni Austin Dendroceros hawaicus Steph. FOLIOCEROS Bharadwaj (Anthocerotaceae, or Foliocerotaceae [Renzaglia & Vaughn 2000*]) Folioceros fuciformis (Mont.) D.C. Bharadw. [Miller et al. 1983; Bharadwaj 1975] Anthoceros badlwinii Baldwin Anthoceros fuciformis Mont. Anthoceros kaluensis Steph. Aspiromitus fuciformis (Mont.) Steph. MEGACEROS Campb. (Anthocerotaceae, or Dendrocerotaceae [Renzaglia & Vaughn 2000*]) Megaceros flagellaris (Mitt.) Steph. [Hasegawa 1983; Piippo 1993] NOTOTHYLAS Sull. (Anthocerotaceae, or Notothyladaceae [Renzaglia & Vaughn 2000*]) Notothylas breutelii (Gottsche) Gottsche [Miller 1967; Miller et al. 1983; Schuster 1992b: 855] Anthoceros breutelii Gottsche PHAEOCEROS Prosk. (Anthocerotaceae, or Notothyladaceae [Renzaglia & Vaughn 2000*]) Phaeoceros carolinianus (Michx.) Prosk. [Miller et al. 1983; Hasegawa 1984; Hasegawa 1991; Campbell 1989; Hässel de Menéndez 1989] Anthoceros communis Steph. Anthoceros havaiensis Reichardt Anthoceros oahuensis Steph. Anthoceros pusillus Steph. Anthoceros validus Steph. Aspiromitus hawaicus (Reichardt) Steph. Phaeoceros laevis (L.) Prosk. subsp. carolinianus (Michx.) Prosk. STATUS NI K O MO L M KA H end nat? M K O Mo ind O ind K O nat? O ind K O M H M H H M H * Renzaglia and Vaughn (2000: 16) contrasted four recent classifications for the hornworts in which genera are placed in various families. See there for details. TROPICAL BRYOLOGY 28 (2006) STAPLES & IMADA: HAWAIIAN ANTHOCEROTES AND HEPATICS 22 Dubious Records Folioceros glandulosus (Lehm. & Lindenb.) D.C. Bharadw. was listed for the Hawaiian Islands (as Anthoceros glandulosus Lehm. & Lindenb.) without island locality, by Miller et al. 1983: 49. Until voucher material is examined the status of this name remains dubious for the Hawaiian hornwort flora. Megaceros vincentianus (Lehm. & Lindenb.) Campb. was reported from 4 islands (Kaua‘i, Moloka‘i, Maui, Hawai‘i) by Miller et al. (1983). According to S. R. Gradstein (pers. comm. 2005) it is unlikely this neotropical hornwort occurs in the Hawaiian Islands. The identity of plants so-called remains to be determined as this checklist is being completed. Apendix 2 Hepaticae of the Hawaiian Islands STATUS NI K O MO L M KA H FULLNAME ACROMASTIGUM A.Evans (Lepidoziaceae) Acromastigum integrifolium (Austin) A.Evans [Miller et ind K O Mo M al. 1983; Schuster 2000: 296 (in key) says A. integrifolium may range to New Caledonia] Bazzania ?integrifolia (Austin) A.Evans Herpocladium bidens Mitt., in part Herpocladium tenacifolium misapplied Mastigobryum integrifolium Austin Mastigobryum rudimentare Steph. ANASTREPTA (Lindb.) Schiffn. (Jungermanniaceae) Anastrepta orcadensis (Hook.) Schiffn. [Schuster 1981; Miller et al. 1983] Jungermannia orcadensis Hook. Lophozia orcadensis (Hook.) Boulay ANASTROPHYLLUM (Spruce) Steph. (Jungermanniaceae) Anastrophyllum esenbeckii (Mont.) Steph. [Miller et al. 1983] Jungermannia esenbeckii Mont. Anastrophyllum fissum Steph. [Miller et al. 1983] Anastrophyllum piligerum misapplied Jungermannia piligera misapplied ARCHILEJEUNEA (Spruce) Schiffn. (Lejeuneaceae) Archilejeunea planiuscula (Mitt.) Steph. [Miller 1967; Miller et al. 1983; Gradstein et al. 2002] Archilejeunea mariana misapplied Archilejeunea owahuensis Gottsche ex Steph. 1 Spruceanthus marianus misapplied ASTERELLA P.Beauv. (Aytoniaceae) Asterella innovans (Austin) H.A.Mill. [Miller et al. 1983] Fimbriaria innovans (Austin) H.A.Mill. Marchantia innovans Austin ind K end K end K ind end Mo M H M O K M 1 The status of A. owahuensis is unclear. The name was reduced to S. marianus by Miller et al. However, Gradstein et al. state that S. marianus is misapplied to Pacific plants, which are Archilejeunea planiuscula. The type material of A. owahuensis has to be studied to decide its true identity. TROPICAL BRYOLOGY 28 (2006) STAPLES & IMADA: HAWAIIAN ANTHOCEROTES AND HEPATICS Asterella parva (Steph.) H.A.Mill. [Miller et al. 1983] Fimbriaria parva Steph. BAZZANIA Gray (Lepidoziaceae) Bazzania baldwinii Austin ex A.Evans [Miller et al. 1983] Bazzania deflexa misapplied Mastigobryum baldwinii (Austin ex A.Evans) Steph. Bazzania brighamii (Austin) A.Evans [Miller et al. 1983] Bazzania ligulata (Sande Lac. ex Steph.) Steph. Mastigobryum brighami Austin Mastigobryum ligulatum Sande Lac. ex Steph. Mastigobryum modestum Steph. Mastigobryum obtusidens Steph. Bazzania cordistipula (Mont.) Trevis. [Miller et al. 1983] Bazzania falcata A.Evans Herpetium cordistipulum Mont. Mastigobryum cordistipulum (Mont.) Lindenb. Mastigobryum honoluluense Steph. Mastigobryum newellii Steph. Mastigobryum subsandei Steph. Bazzania didericiana (Gottsche ex Steph.) Steph. [Miller et al. 1983] Bazzania didricksenii (Steph.) W.E.Nicholson Mastigobryum didericiana Gottsche ex Steph. Mastigobryum didricksenii Steph. Pleuroschisma pearsonii (Steph.) Müll.Frib. Bazzania emarginata (Steph.) C.M.Cooke [Miller et al. 1983] Mastigobryum bispinosum Steph. Mastigobryum didericiana Gottsche ex Steph. var. emarginata Steph. Mastigobryum emarginatum (Steph.) Steph. Mastigobryum hawaiicum Steph. Mastigobryum kalihiense Beauverd Bazzania inaequabilis Steph. ex C.M.Cooke [Miller et al. 1983] Mastigobryum inaequabilis (Steph. ex C.M.Cooke) Steph. Mastigobryum pacificum Steph. Bazzania minuta (Austin) A.Evans [Miller et al. 1983] Bazzania vittata misapplied Mastigobryum minutum Austin Bazzania nuuanuensis C.M.Cooke [Miller et al. 1983] Mastigobryum nuuanuense (C.M.Cooke) Steph. Mastigobryum repandifolium Steph. Bazzania patens (Mont.) Trevis. [Miller et al. 1983] Bazzania beecheyana Steph. Bazzania erosistipula Steph. Herpetium patens Mont. Mastigobryum beecheyanum (Steph.) Steph. Mastigobryum grossidens Steph. Mastigobryum parvistipulum Austin Mastigobryum patens (Mont.) Gottsche, Lindenb., & Nees Mastigobryum tantalum Steph. Bazzania praerupta (Reinw., Blume & Nees) Trevis. [Miller et al. 1983; Mizutani & Chang 1986; Long & TROPICAL BRYOLOGY 28 (2006) end 23 Mo end K Mo M ind K O Mo M H ind K O Mo M H end K O Mo M H end K O Mo M H ind K O end K M H Mo end O H end K O M ind K O Mo M H 24 STAPLES & IMADA: HAWAIIAN ANTHOCEROTES AND HEPATICS Grolle 1990] Bazzania decurva (Nees) Trevis. Jungermannia praerupta Reinw., Blume & Nees Bazzania sandvicensis (Gottsche ex Steph.) Steph. [Miller et al. 1983] Mastigobryum rockii Steph. Mastigobryum sandwicense Gottsche ex Steph. BLEPHAROSTOMA (Dumort.) Dumort. (Blepharostomataceae or Pseudolepicoleaceae [Crandall-Stotler & Stotler 2000]) Blepharostoma trichophyllum (L.) Dumort. [Miller 1967; Miller et al. 1983] Jungermannia trichophylla L. CALYPOGEIA Raddi (Calypogeiaceae) Calypogeia aeruginosa Mitt. [Miller et al. 1983] Calypogeia aeruginosa var. waialealeensis H.A.Mill. & Kuwah. Calypogeia waialealeensis (H.A.Mill. & Kuwah.) H.A.Mill. Calypogeia arguta Nees & Mont. ex Nees [Miller et al. 1983] Calypogeia bifurca Austin Calypogeia pacifica Steph. Kantia arguta (Nees & Mont. ex Nees) Lindb. Kantia bifurca (Austin) A.Evans Calypogeia cuspidata (Steph.) Steph. [Miller et al. 1983] Calypogeia confertifolia Steph. Calypogeia hawaica Steph. Kantia cuspidata Steph. Calypogeia tosana (Steph.) Steph. [Miller et al. 1983] Calypogeia bidentula Austin Kantia bidentula A.Evans Kantia tosana Steph. Kantia trichomanis misapplied CEPHALOZIA (Dumort.) Dumort. (Cephaloziaceae) Cephalozia sandvicensis (Mont.) Spruce [Miller et al. 1983] Blepharostoma sandwicense (Mont.) Trevis. Cephalozia connivens misapplied Cephalozia multiflora misapplied Jungermannia crassifolia Lindenb. & Gottsche Jungermannia sandwicensis Mont. CEPHALOZIELLA (Spruce) Schiffn. (Cephaloziellaceae) Cephaloziella baldwinii (C.M.Cooke) H.A.Mill. [Cooke 1904; Miller et al. 1983] Cephalozia baldwinii C.M.Cooke Cephaloziella heteroica (C.M.Cooke) Douin [Miller et al. 1983] Cephalozia heteroica C.M.Cooke Cephaloziella kilohanensis (C.M.Cooke) Douin [Miller et al. 1983] Cephalozia kilohanensis C.M.Cooke end K O Mo M H ind K M ind K ind K O M H end K O Mo M H ind K O Mo M H ind K O Mo M H end K Mo M H end K Mo M end K M TROPICAL BRYOLOGY 28 (2006) STAPLES & IMADA: HAWAIIAN ANTHOCEROTES AND HEPATICS Cephaloziella lilae (C.M.Cooke) Douin [Miller et al. 1983] Cephalozia lilae C.M.Cooke CHEILOLEJEUNEA (Spruce) Schiffn. (Lejeuneaceae) Cheilolejeunea decursiva (Sande Lac.) R.M.Schust. [Miller et al. 1983; Zhu & Lai 2005: 197] Trachylejeunea oahuensis A.Evans Cheilolejeunea hawaica Steph. [Miller et al. 1983] Cheilolejeunea intertexta (Lindenb.) Steph. [Miller et al. 1983] Lejeunea intertexta Lindenb. Cheilolejeunea pyriflora Steph. [Miller 1967; Miller et al. 1983] Cheilolejeunea intertexta misapplied Cheilolejeunea stenoschiza (Ångstr.) A.Evans [Miller et al. 1983] Lejeunea stenoschiza Ångstr. Pycnolejeunea stenoschiza (Ångstr.) Steph. CHILOSCYPHUS Corda (Geocalycaceae) Chiloscyphus autoicus (Steph.) J.J.Engel & R.M.Schust. [Miller et al. 1983; Engel & Schuster 1984] Lophocolea autoica Steph. Chiloscyphus baldwinii (Steph.) J.J.Engel & R.M.Schust. [Miller et al. 1983; Engel & Schuster 1984] Lophocolea baldwinii Steph. Lophocolea breutelii misapplied Lophocolea columbica misapplied Lophocolea orbigiana misapplied Chiloscyphus bartlettii (H.A.Mill.) J.J.Engel & R.M.Schust. [Miller et al. 1983; Engel & Schuster 1984] Lophocolea bartlettii H.A.Mill. Chiloscyphus bicuspidatus (Steph.) J.J.Engel & R.M.Schust. [Miller et al. 1983; Engel & Schuster 1984] Lophocolea bicuspidata Steph. Chiloscyphus ciliolatus (Nees) J.J.Engel & R.M.Schust. [Miller et al. 1983; Engel & Schuster 1984; Piippo 1985a] Chiloscyphus beecheyanus (Taylor) J.J.Engel & R. Schust. Chiloscyphus gaudichaudii (Mont.) J.J.Engel & R. Schust. Jungermannia ciliolata Nees Lophocolea beecheyana Taylor Lophocolea bidentata misapplied Lophocolea ciliolata (Nees) Gottsche Lophocolea gaudichaudii Mont. Lophocolea newellana Steph. Lophocolea trispinosa Steph. Chiloscyphus greenwelliae (H.A.Mill.) H.A.Mill. [Miller et al. 1983] Lophocolea connata misapplied Lophocolea greenwelliae H.A.Mill. Chiloscyphus hawaiicus (Steph.) J.J.Engel & R.M.Schust. [Miller et al. 1983; Engel & Schuster TROPICAL BRYOLOGY 28 (2006) 25 end K O end? K O Mo M ind ind K O Mo K O Mo M M ind? end H H O K O Mo ind M O H end K O Mo M H end Mo M H M H end ind K O Mo M H end K M H end K Mo H 26 STAPLES & IMADA: HAWAIIAN ANTHOCEROTES AND HEPATICS 1984] Lophocolea hawaiica Steph. Chiloscyphus kilauensis Steph. [Miller et al. 1983] Lophocolea kilauensis (Steph.) H.A.Mill. Chiloscyphus laceratus Steph. [Miller 1967; Miller et al. 1983] Chiloscyphus decurrens misapplied Chiloscyphus mitteni misapplied Chiloscyphus lambertonii H.A.Mill. [Miller et al. 1983] Chiloscyphus muricatus (Lehm.) J.J.Engel & R.M.Schust. [Miller et al. 1983; Engel & Schuster 1984] Jungermannia muricata Lehm. Lophocolea muricata (Lehm.) Nees Chiloscyphus skottsbergianus H.A.Mill. [Miller et al. 1983] Chiloscyphus spinosus (Gottsche) J.J.Engel & R.M.Schust. [Miller et al. 1983; Engel & Schuster 1984] Lophocolea spinosa Gottsche end O H end Mo M end ind Mo M H H end H end K O M H ind K O Mo M H end K COLOLEJEUNEA (Spruce) Schiffn. (Lejeuneaceae) Cololejeunea ceatocarpa (Ångstr.) Steph. [Miller et al. 1983; Tixier 1985] Lejeunea ceatocarpa Ångstr. Leptocolea ceatocarpa (Ångstr.) Steph. Physocolea ceatocarpa (Ångstr.) Steph. Cololejeunea cookei A.Evans [Miller 1963; Miller et al. 1983] Physocolea cookei (A.Evans) Steph. Cololejeunea dzumacensis Tixier [Tixier 1985] Cololejeunea hillebrandii (Austin) Steph. [Miller et al. 1983; Tixier 1985] Lejeunea hillebrandii Austin Lejeunea longifolia Austin Physocolea hillebrandii (Austin) Steph. Cololejeunea hoeana Tixier [Tixier 1985] Cololejeunea lanciloba Steph. [Miller et al. 1983] Leptocolea lanciloba (Steph.) A.Evans Pedinolejeunea lanciloba (Steph.) P.C. Chen & P.C.Wu Cololejeunea longistyla A.Evans [Miller et al. 1983] Cololejeunea kauaiensis (Steph.) Bischl. Physocolea kauaiensis Steph. Physocolea longistyla (A.Evans) Steph. Cololejeunea obcordata (Austin) A.Evans [Miller et al. 1983; Tixier 1985] Lejeunea obcordata Austin Leptocolea mamilliloba Steph. Leptocolea obcordata (Austin) A.Evans Physocolea mamilliloba Steph. Cololejeunea ovalifolia A.Evans [Miller et al. 1983] Leptocolea ovalifolia (A.Evans) A.Evans ind ind O K O Mo end ind? O K O end K O end K O COLURA (Dumort.) Dumort. (Lejeuneaceae) Colura tenuicornis (A.Evans) Steph. [Miller et al. 1983] Colura calyptrifolia misapplied ind K O Mo end H O Mo M M M H TROPICAL BRYOLOGY 28 (2006) STAPLES & IMADA: HAWAIIAN ANTHOCEROTES AND HEPATICS 27 Colura longicornis misapplied Colura tenuis Steph. Colurolejeunea tenuicornis A.Evans Lejeunea calyptrifolia misapplied DIPLOPHYLLUM (Dumort.) Dumort. (Scapaniaceae) Diplophyllum albicans (L.) Dumort. [Miller et al. 1983] Diplophyllum exiguum Steph. [Miller et al. 1983] DREPANOLEJEUNEA (Spruce) Schiffn. (Lejeuneaceae) Drepanolejeunea anderssonii (Ångstr.) A.Evans [Miller et al. 1983] Harpalejeunea anderssonii (Ångstr.) Steph. Lejeunea anderssonii Ångstr. Drepanolejeunea pseudoneura (A.Evans) Grolle [Miller et al. 1983; Grolle 1988] Harpalejeunea falsinervis A.Evans ex Bonner Harpalejeunea pseudoneura A.Evans Drepanolejeunea ungulata (Steph.) Grolle [Miller et al. 1983; Grolle 1991] Drepanolejeunea uncinata misapplied Harpalejeunea ungulata Steph. Lejeunea ungulata Ångstr. DUMORTIERA Nees (Marchantiaceae) Dumortiera hirsuta (Sw.) Nees [Miller et al. 1983] Dumortiera trichocephala (Hook.) Nees Marchantia hirsuta Sw. Marchantia trichocephala Hook. FOSSOMBRONIA Raddi (Fossombroniaceae) Fossombronia pusilla (L.) Nees [Krayesky et al. 2005] FRULLANIA Raddi (Frullaniaceae, or Jubulaceae [Crandall-Stotler & Stotler 2000]) Frullania apiculata (Reinw., Blume & Nees) Nees [Miller 1953; Miller et al. 1983] Frullania crenatiloba Steph. Frullania oceanica Mitt. Jubula vittata Steph. Jungermannia apiculata Reinw., Blume & Nees Frullania ericoides (Nees) Mont. [Miller 1953; Miller et al. 1983; Hattori & Piippo 1986] Frullania elliptica Steph. Frullania sandvicensis Ångstr., in part ? Frullania squarrosa (Reinw., Blume & Nees) Dumort. Jungermannia squarrosa Reinw., Blume & Nees Frullania hawaiiensis H.A.Mill. [Miller 1953; Miller et al. 1983] Frullania hypoleuca Nees [Miller 1953; Miller et al. 1983; Hattori 1981] Frullania explicata Mont. Frullania gymnotis Nees & Mont. Frullania parvilobula Steph. Frullania papulirostra Steph. TROPICAL BRYOLOGY 28 (2006) ind end end end M H K O Mo O M H M H end K O Mo M H ind K O M H ind M ind K O Mo M H ind K O Mo M H end O Mo M ind K O Mo M H 28 STAPLES & IMADA: HAWAIIAN ANTHOCEROTES AND HEPATICS Frullania ptychantha misapplied Frullania meyeniana Lindenb. [Miller 1953; Miller et al. 1983] Frullania helleri Steph. Frullania neurota Taylor [Miller 1953; Miller et al. 1983; Yuzawa 1991] Frullania aongstroemii A.Evans Frullania arietina misapplied Frullania sandvicensis Ångstr. Frullania oahuensis Hampe [Miller 1953; Miller et al. 1983] Frullania cookei A.Evans ex Steph. FULFORDISTRIA H.A.Mill. (Schistochilaceae) Fulfordistria cookei H.A.Mill. [Miller et al. 1983] Schistochila cookei (H.A.Mill.) Schust. GEOCALYX Nees (Geocalycaceae) Geocalyx graveolens (Schrad.) Nees [Miller 1967; Miller et al. 1983] Jungermannia graveolens Schrad. HERBERTUS Gray (Herbertaceae) Herbertus gracilis (Mont.) Steph. [Miller 1963; Miller et al. 1983] Herpocladium bidens Mitt., in part Herpocladium gracile (Mont.) Steph. Mastigobryum biapiculatum Steph. Mastigophora gracilis Mont. Mastigophora parvifolia Steph. Schisma gracilis Mont. Sendtnera gracilis (Mont.) Gottsche, Lindenb. & Nees Sendtnera tristicha Austin Herbertus hawaiiensis H.A.Mill. [Miller & Scott 1960; Miller 1967; Miller et al. 1983; Hong et al. 1993] Herbertus sakuraii misapplied Herbertus helleri (Steph.) W.E.Nicholson [Miller et al. 1983; So 2003a] Herbertus longifissus misapplied Herbertus parisii (Steph.) H.A.Mill., in part Herbertus sanguineus Austin Herbertus setigerus (Steph.) H.A.Mill., in part Schisma helleri Steph. Schisma juniperina misapplied Schisma longifissa misapplied Schisma setigera Steph. Herbertus herpocladioides E.B.Scott & H.A.Mill. [Scott & Miller 1959; Miller 1963; Miller et al. 1983] Herbertus sanguineus (Mont.) Austin [Miller 1963; Miller et al. 1983] Schisma juniperinum (Sw.) Nees var. sanguineum Austin Schisma sanguineum (Austin) Steph. HYGROLEJEUNEA (Spruce) Steph. (Lejeuneaceae) Hygrolejeunea pacifica Steph. [Miller et al. 1983] ind K O Mo M H ind K O Mo L M H end K O Mo H end Mo M ind? ind H K O Mo ind M H M end O M H ind K Mo M H ind K O Mo M H end O TROPICAL BRYOLOGY 28 (2006) STAPLES & IMADA: HAWAIIAN ANTHOCEROTES AND HEPATICS JAMESONIELLA (Spruce) Carrington in F. Lees (Jungermanniaceae) Jamesoniella labrifolia H.A.Mill. [Miller 1967; Miller et al. 1983] Jamesoniella robusta (Austin) Spruce ex Steph. [Miller et al. 1983] Anastrophyllum contractum misapplied Anastrophyllum coriaceum (Austin) Steph. Anastrophyllum macrophyllum (Ångstr.) Steph. Anastrophyllum ? rigidum (Austin) Steph. Crossogyna robusta (Austin) Schljakov Jamesoniella coriacea (Austin) Spruce Jamesoniella macrophylla (Ångstr.) Steph. Jungermannia contracta misapplied Jungermannia coriacea Austin Jungermannia macrophylla Ångstr. Jungermannia rigida Austin Jungermannia robusta Austin JUBULA Dumort. (Jubulaceae) Jubula hutchinsiae (Hook.) Dumort. subsp. javanica (Steph.) Verd. [Miller 1953; Guerke 1978; Miller et al. 1983] Frullania piligera Austin Jubula piligera (Austin) A.Evans Jubula setacea Steph. JUNGERMANNIA L. (Jungermanniaceae) Jungermannia hawaiica (H.A.Mill.) Váňa [Váňa 1975] Nardia exserta A.Evans Solenostoma exsertum (A.Evans) Steph. Solenostoma hawaiicum H.A.Mill. Jungermannia micrantha (Mitt.) Steph. [Váňa 1975] Jungermannia baldwinii Steph. Jungermannia callithrix misapplied Jungermannia lurida A.Evans Jungermannia mauii Austin Jungermannia nana misapplied Jungermannia newellana Steph. Jungermannia sexplicata Steph. Mylia taylori misapplied Nardia callithrix misapplied Nardia mauii (Austin) A.Evans Nardia micrantha (Mitt.) Steph. Nardia truncata misapplied Plectocolea mauii (Austin) H.A.Mill. Plectocolea micrantha Mitt. Plectocolea submicrantha H.A.Mill. KURZIA G.Martens (Lepidoziaceae) Kurzia hawaica (C.M.Cooke) Grolle [Miller et al. 1983] Lepidozia hawaica C.M.Cooke Lepidozia minutifolia Steph. Microlepidozia hawaica (C.M.Cooke) H.A.Mill. Kurzia mauiensis (H.A.Mill.) H.A.Mill. [Miller et al. 1983; Piippo 1985b] Microlepidozia mauiensis H.A.Mill. TROPICAL BRYOLOGY 28 (2006) 29 end Mo M H ind K O Mo M H ind K O Mo L M H end K O Mo M H ind K O Mo M H end K O Mo M H ind M 30 STAPLES & IMADA: HAWAIIAN ANTHOCEROTES AND HEPATICS Kurzia tayloriana (H.A.Mill.) H.A.Mill. [Miller et al. 1983] Microlepidozia tayloriana H.A.Mill. LEJEUNEA Lib. (Lejeuneaceae) Lejeunea anisophylla Mont. [Miller et al. 1983] Lejeunea pacifica Mont. [Miller et al. 1983] Eulejeunea pacifica (Mont.) Steph. LEPIDOLEJEUNEA R.M.Schust. (Lejeuneaceae) Lepidolejeunea involuta (Gottsche) Grolle [Miller et al. 1983; Piippo 1986–without island locality] Harpalejeunea owaihiensis (Gottsche) Steph. Hygrolejeunea owaihiensis (Gottsche) Steph. Lejeunea hawaiiensis Gottsche ex Mitt. Lejeunea owaihiensis Gottsche Lopholejeunea owaihiensis (Lindenb. in Gottsche) Steph. Strepsilejeunea owaihiensis (Gottsche) Steph. LEPIDOZIA (Dumort.) Dumort. (Lepidoziaceae) Lepidozia australis (Lehm. & Lindenb.) Mitt. [Miller et al. 1983] Jungermannia australis Lehm. & Lindenb. Lepidozia remyana Steph. Lepidozia reptans misapplied Lepidozia reptans Evans var. australis (Lehm. & Lindenb.) Gottsche, Lindenb. & Nees Lepidozia triceps Taylor Mastigophora triceps (Taylor) Trevis. Lepidozia sandvicensis Lindenb. ex Gottsche, Lindenb. & Nees [Miller et al. 1983] Lepidozia filipendula Taylor Lepidozia pusilla Steph. Mastigophora filipendula (Taylor) Trevis. Mastigophora sandvicensis (Lindenb.) Trevis. Lepidozia sellingiana H.A.Mill. [Miller et al. 1983] LIOCHLAENA Nees (Jungermanniaceae) Liochlaena subulata (A.Evans) Schljakov [Grolle & Váňa 1969; Miller et al. 1983; Long & Grolle 1990; Schljakov 1981] Jamesoniella subulata (A.Evans) Steph. Jungermannia subulata A.Evans LOPHOLEJEUNEA (Spruce) Schiffn. (Lejeuneaceae) Lopholejeunea nigricans (Lindenb.) Schiffn. [Miller et al. 1983; Zhu & Gradstein 2005] Lejeunea gibbosa Ångstr. Lejeunea hawaica Steph. Lejeunea mannii Austin Lopholejeunea gibbosa (Ångstr.) Steph. Lopholejeunea hawaica Steph. Lopholejeunea mannii (Austin) Steph. Lopholejeunea owahuensis Gottsche ex Steph. Lopholejeunea subnuda (Mitt.) Steph. Mastigolejeunea honoluluana Steph. end K M ind? ind K O Mo K O Mo M end K M ind K H H ind Mo M end K ind K O Mo M H ind K O Mo M H TROPICAL BRYOLOGY 28 (2006) STAPLES & IMADA: HAWAIIAN ANTHOCEROTES AND HEPATICS Phragmicoma subnuda Mitt. Lopholejeunea proxima Steph. [Miller et al. 1983] end LOPHOZIA (Dumort.) Dumort. (Jungermanniaceae) Lophozia perssoniana H.A.Mill. [Miller et al. 1983] end LUNULARIA Adans. (Lunulariaceae) Lunularia cruciata (L.) Dumort. ex Lindenb. [Miller 1967; Yoshida & Smith 1976] Marchantia cruciata L. MARCHANTIA L. (Marchantiaceae) Marchantia crenata Austin [Bischler-Causse 1989] Fimbriaria rechingeri Steph. Marchantia chenopoda misapplied Marchantia cuspidisquama Steph. Marchantia disjuncta misapplied Marchantia furciloba Steph. Marchantia punaluana Steph. Marchantia rechingeri (Steph.) H.A.Mill. Marchantia sandvicensis Reichardt Marchantia marginata Steph. [Bischler-Causse 1989] Marchantia antiqua Steph. Marchantia disjuncta misapplied Marchantia eatonii Austin ex H.A.Mill. Marchantia nitida misapplied Marchantia paleacea misapplied MARCHESINIA Gray (Lejeuneaceae) Marchesinia mittenii A.Evans [Evans 1900; Miller et al. 1983–without island locality] METACALYPOGEIA (S.Hatt.) Inoue (Calypogeiaceae) Metacalypogeia alternifolia (Nees ex Gottsche, Lindenb. & Nees) Grolle [Miller et al. 1983, Long & Grolle 1990] Bazzania alternifolia (Nees) Trevis. Calypogeia alternifolia (Nees ex Gottsche, Lindenb. & Nees) Steph. Calypogeia trichomanis misapplied Kantia alternifolia (Nees ex Gottsche, Lindenb. & Nees) Spruce Kantia rotundistipula Steph. Mastigobryum alternifolium Nees ex Gottsche, Lindenb. & Nees Metacalypogeia montana (Horik.) Inoue var. verruculosa (S.Hatt.) H.A.Mill. Metacalypogeia remotifolia (Herzog) Inoue METAHYGROBIELLA R.M.Schust. (Cephaloziaceae) Metahygrobiella lucens (A.Evans) H.A.Mill. [Miller et al. 1983] Cephalozia lucens (A.Evans) Bonner Jungermannia lucens A.Evans Metahygrobiella maxima (Steph.) N.Kitag. [Miller et al. 1983] TROPICAL BRYOLOGY 28 (2006) nat 31 K Mo O M H M ? end K O M end K O Mo M K O Mo M H end ind end M end M H 32 STAPLES & IMADA: HAWAIIAN ANTHOCEROTES AND HEPATICS Cephalozia maxima Steph. METALEJEUNEA Grolle (Lejeuneaceae) Metalejeunea cucullata (Reinw., Blume, & Nees) Grolle [Miller et al. 1962; Miller et al. 1983; Grolle 1995] Jungermannia cucullata Reinw., Blume, & Nees Microlejeunea albicans (Nees) Steph. Microlejeunea cucullata (Reinw., Blume, & Nees) Steph. Microlejeunea erectifolia misapplied Microlejeunea sundaica W.E.Nicholson METZGERIA Raddi (Metzgeriaceae) Metzgeria furcata (L.) Dumort. [Kuwahara 1960; Miller et al. 1983; Long & Grolle 1990; So 2002b] Metzgeria decipiens (C.Massal.) Schiffn. Metzgeria dichotoma misapplied Metzgeria mauina Steph. Metzgeria molokaiensis Kuwah. Metzgeria quadriseriata A.Evans Metzgeria leptoneura Spruce [Kuwahara 1960; Miller et al. 1983; Piippo 1991; So 2002b] Metzgeria papulosa Steph. Metzgeria lindbergii Schiffn. [Kuwahara 1960; Miller et al. 1983; Piippo 1991; So 2002b] Metzgeria oceanica Kuwah. Metzgeria pectinata Steph. Metzgeria pubescens (Schrank) Raddi [So 2003b– without island loc.] MICROLEJEUNEA Steph. (Lejeuneaceae) Microlejeunea lunulatiloba Horik. [Miller et al. 1962; Miller et al. 1983; So 2005b] MNIOLOMA Herzog (Calypogeiaceae) Mnioloma fuscum (Lehm. & Lindenb.) R.M.Schust. [Miller et al. 1983; Kitagawa 1988; Schuster 1995: 848–9] Calypogeia baldwinii Austin Calypogeia fusca (Lehm. & Lindenb.) Steph. Jungermannia fusca Lehm. & Lindenb. Kantia baldwinii (Austin) A.Evans Metacalypogeia fusca (Lehm. & Lindenb.) N. Kitag. Omphalanthus baldwinii nom. nud. NOTOSCYPHUS Mitt. (Jungermanniaceae) Notoscyphus lutescens (Lehm. & Lindenb.) Mitt. [Stephani 1924; Miller et al. 1983; Váňa et al. 2005] Calypogeia minutistipula Steph. Gymnanthe bolanderi Austin Jungermannia bolanderi Austin Jungermannia lutescens Lehm. & Lindenb. Notoscyphus paroicus Schiffn. Saccogyna bolanderi (Austin) A.Evans ind K O Mo M H ind K O Mo M H K O Mo M H O Mo M H ind ind O ind? ind ind ind M K O M H ODONTOSCHISMA (Dumort.) Dumort. TROPICAL BRYOLOGY 28 (2006) STAPLES & IMADA: HAWAIIAN ANTHOCEROTES AND HEPATICS (Cephaloziaceae) Odontoschisma sandvicense (Ångstr.) A.Evans [Miller et al. 1983; So 2004] Gamoscyphus sandvicense (Ångstr.) Trevis. Odontoschisma gracile (Mitt.) Steph. Odontoschisma grandistipulum Steph. Sphagnoecetis gracilis Mitt. Sphagnoecetis sandvicense Ångstr. Odontoschisma subjulaceum Austin [Miller et al. 1983; So 2004] Jungermannia caudifera Taylor ex Austin PALLAVICINIA Gray (Pallaviciniaceae) Pallavicinia lyellii (Hook.) Carruth. [Miller et al. 1983; Grolle & Piippo 1986] Pallavicinia cylindrica (Austin) A.Evans Pallavicinia simplex Steph. Steetzia cylindrica Austin PLAGIOCHASMA Lehm. & Lindenb. (Aytoniaceae) Plagiochasma japonicum (Steph.) C.Massal. [Bischler 1979; Long & Grolle 1990] Plagiochasma cordatum misapplied Rupinia cordata misapplied PLAGIOCHILA (Dumort.) Dumort. (Plagiochilaceae) Plagiochila baldwinii Austin ex A.Evans [Inoue 1976; So 2001a] Plagiochila acutiuscula Austin ex A.Evans Plagiochila flava misapplied Plagiochila conduplicata Steph. [Inoue 1976; Miller 1963; Miller et al. 1983; So 2001a] Plagiochila convoluta Steph. [Inoue 1976; So 2001a] Plagiochila deflexa Mont. & Gottsche [Inoue 1976; Miller et al. 1983; So 2001a] Plagiochila adiantoides misapplied Plagiochila arbuscula misapplied Plagiochila askenasii Steph. Plagiochila belangeriana misapplied Plagiochila caespitosa Steph. Plagiochila nahikuna Steph. Plagiochila patula (Sw.) Nees & Mont. Plagiochila rockii Steph. Plagiochila fissidentoides Taylor [Inoue 1976: 439 excl. Molokai; So 2001a] Plagiochila simplex misapplied Plagiochila subtilis Dugas Plagiochila variegata misapplied Plagiochila gaudichaudii Mont. & Gottsche [Inoue 1976; So 2001a] Plagiochila haleakalensis Steph. Plagiochila manni Steph. Plagiochila tenuis misapplied Plagiochila gracillima Austin ex A.Evans [Inoue 1976; So 2001a] Plagiochila grossispina Steph. [Inoue 1976; So 2001a; Hoe 3530.2 from Maui, det. M.L. So 3/2004] TROPICAL BRYOLOGY 28 (2006) 33 end K O Mo M H end K O Mo M H ind Mo M H ind O end M end K O Mo H end end K O K O Mo M H H M H end O end K O Mo M H end Mo M H M H end O 34 STAPLES & IMADA: HAWAIIAN ANTHOCEROTES AND HEPATICS Plagiochila modesta Dugas Plagiochila hawaica Steph. [Inoue 1976; So 2001a] Plagiochila hoei Inoue [Inoue 1976; So 2001a] Plagiochila deltoidea misapplied Plagiochila mauiensis Steph. (as “marriensis,” an orthogr. error) [Inoue 1976; So 2001a; Cooke 129 from Kaua‘i det. M.L. So 3/2004] Plagiochila frondescens misapplied Plagiochila mauiensis Dugas Plagiochila maunakeana Steph. (as “mannakeana,” an orthogr. error) [Inoue 1976; So 2001a; Cooke 88 from Kaua‘i det. M.L. So 3/2004] Plagiochila biserialis misapplied Plagiochila remyana Steph. [Inoue 1976; Schuster 1980; So 2001a; Svihla 35-282 from Maui, det. M.L. So 3/2004] Plagiochila mollucensis Steph. Plagiochila owaihiensis misapplied Plagiochila yokogurensis Steph. subsp. remyana (Steph.) R.M.Schust. PLAGIOCHILION S.Hatt. (Plagiochilaceae) Plagiochilion combinatum (Mitt.) Inoue [Miller 1963; Inoue 1964; Miller et al. 1983; So 2001a] Plagiochila combinata Mitt. Plagiochila oahuna Steph. Plagiochila oppositifolia Austin PLEUROZIA Dumort. (Pleuroziaceae) Pleurozia conchifolia (Hook. & Arn.) Austin var. conchifolia [Miller et al. 1983; Thiers 1993] Jungermannia conchaefolia Hook. & Arn. Physiotium conchaefolia (Hook. & Arn.) Nees Pleurozia cochleariformis (Hook. & Arn.) Trevis. Pleurozia purpurea misapplied Pleurozia gigantea (F.Weber) Lindb. [Miller et al. 1983; Baldwin 18 from Maui det. B. Thiers 3/2006] Jungermannia gigantea F.Weber Physiotium gigantea (F.Weber) Lindb. Pleurozia subinflata (Austin) Austin [Miller et al. 1983; Thiers 1993] Physiotium evolutum Mitt. Physiotium subinflatum Austin Pleurozia evoluta (Mitt.) Steph. PORELLA L. (Porellaceae) Porella acutifolia (Lehm. & Lindenb.) Trevis. [Miller et al. 1983; Hattori 1978; So 2002a] Madotheca acutifolia Lehm. & Lindenb. Madotheca hawaiiensis (A.Evans) Steph. Porella hawaiica sphalm. Porella hawaiiensis A.Evans Porella hoeana S.Hatt. [Hattori 1976; Miller et al. 1983] Porella laevigata misapplied RADULA Dumort. (Radulaceae) Radula acutangula Steph. [Miller et al. 1983; So 2005b] end end M M H end K O Mo M H end K M H end K O M H end K O Mo M H ind K O Mo M H Mo ind ind M K O M ind M end M end K O Mo M H H TROPICAL BRYOLOGY 28 (2006) STAPLES & IMADA: HAWAIIAN ANTHOCEROTES AND HEPATICS Radula pauciramea Steph. Radula cavifolia Hampe ex Gottsche, Lindenb., & Nees [So 2005b] Radula aneurismalis misapplied Radula cordata Mitt. [Yamada 1979; Miller et al. 1983; So 2005b] Radula excisiloba Steph. Radula mannii Austin Radula xalapensis misapplied ? ? ? Radula curvilobula M.L.So [So 2005b] Radula gracilis Mitt. ex Steph. [Miller et al. 1983; So 2005b] Radula lejeuneaformis Steph. Radula hawaiica M.L.So [So 2005b] Radula javanica Gottsche [Miller et al. 1983; So 2005b] Radula cordiloba Taylor Radula pallens misapplied Radula perrottetii misapplied Radula mauiensis M.L.So [So 2005b] Radula reflexa Nees & Mont. [Miller et al. 1983; So 2005b] Radula perrotettii misapplied Radula sullivantii Austin [So 2005b] Radula perrotettii misapplied RICCARDIA Gray (Aneuraceae) Riccardia alba (Colenso) E.A.Br. [Hürlimann 1976; Miller et al. 1983; Brown & Braggins 1989–all without island loc.] Aneura alba Colenso Aneura samoana Steph. Riccardia samoana (Steph.) H.A.Mill. Riccardia attenuata (Steph.) H.A.Mill. [Miller et al. 1983] Aneura attenuata Steph. Riccardia baldwinii (Steph.) H.A.Mill. [Miller et al. 1983] Aneura baldwinii Steph. Aneura tamariscina misapplied Riccardia brunnea (Steph.) S.Hatt. [Miller et al. 1983] Aneura brunnea Steph. Riccardia densiramea (Steph.) S.Hatt. [Miller et al. 1983] Aneura densiramea Steph. Riccardia flaccida (Steph.) S.Hatt. [Miller et al. 1983] Aneura flaccida Steph. Aneura multifida misapplied Riccardia multifida misapplied Riccardia fleischeri (Steph.) H.A.Mill. [Miller et al. 1983] Aneura fleischeri Steph. Riccardia hamatifolia (Steph.) H.A.Mill. [Miller et al. 1983] Aneura hamatifolia Steph. Aneura hamatiflora sphalm. Riccardia hawaica (Steph.) H.A.Mill. [Miller et al. 1983] Aneura hawaica Steph. TROPICAL BRYOLOGY 28 (2006) ind 35 K M H end? K O Mo M H end ind K O Mo M M H O O Mo L M H end ind end ind K O M M ind? K M ind end O Mo M end H end Mo end K end K O Mo M H H end O end O end H K O Mo M H H 36 STAPLES & IMADA: HAWAIIAN ANTHOCEROTES AND HEPATICS Riccardia multifida (L.) Gray [Miller et al. 1983; Schuster 1992a: 620–621] Jungermannia multifida L. Riccardia multispica (Steph.) S.Hatt. [Miller et al. 1983] Aneura multispica Steph. Aneura palmata misapplied Riccardia palmata misapplied Riccardia newellana (Steph.) H.A.Mill. [Miller et al. 1983] Aneura newellana Steph. Riccardia nigra (Steph.) H.A.Mill. [Miller et al. 1983] Aneura nigra Steph. Riccardia obtusifrons (Steph.) H.A.Mill. [Miller et al. 1983] Aneura obtusifrons Steph. Aneura pinguis misapplied Aneura plana Steph. Riccardia pauciramea (Steph.) H.A.Mill. [Miller et al. 1983] Aneura pauciramea Steph. Riccardia pectinata (Austin) H.A.Mill. [Miller et al. 1983] Aneura pectinata Austin Riccardia punahuina (Steph.) H.A.Mill. [Miller et al. 1983] Aneura punahuina Steph. Riccardia rockii (Steph.) H.A.Mill. [Miller et al. 1983] Aneura rockii Steph. RICCIA L. (Ricciaceae) Riccia fluitans L. [Hall et al. 1995] Riccia hawaiiensis Hürl. [Hürlimann 1986] Riccia rechingeri Steph. [Yoshida & Smith 1976; Miller et al. 1983] Riccia sorocarpa Bischl. [Hoe 1978; Miller et al. 1983; Schuster 1992b] RICCIOCARPUS Corda (Ricciaceae) Ricciocarpus natans (L.) Corda [Miller 1967; Miller et al. 1983] SCAPANIA (Dumort.) Dumort. (Scapaniaceae) Scapania grossidens Steph. [Miller et al. 1983] Scapania dentata Dumort. var. oakesii misapplied Scapania oakesii Austin Scapania hawaica Müll.Frib. [Miller et al. 1983] Scapania nemorosa misapplied Scapania mauina Steph. [Miller et al. 1983] Scapania ‘manina’ sphalm. Scapania ornithopodioides (With.) Waddell [Miller et al. 1983; Long & Grolle 1990] Jungermannia ornithopodioides With. Scapania planifolia (Hook.) Dumort. Scapania paucidens Steph. [Miller et al. 1983] Scapania undulata misapplied nat? O end? K O H end H end H end M end end O K O Mo M H M H end H end M nat? end nat O H H nat M nat? O end end M K M end M ind M end Mo H TROPICAL BRYOLOGY 28 (2006) STAPLES & IMADA: HAWAIIAN ANTHOCEROTES AND HEPATICS SOUTHBYA Spruce (Arnelliaceae) Southbya grollei N.Kitag. [Váňa 1975] SPRUCEANTHUS Verd. (Lejeuneaceae) Spruceanthus polymorphus (Sande Lac.) Verd. [Mizutani 1961; Miller et al. 1983] Brachiolejeunea aliena (Ångstr.) Steph. Brachiolejeunea apiculata Steph. Brachiolejeunea polymorpha (Sande Lac.) Steph. Dicranolejeunea didericiana Steph. Lejeunea aliena Ångstr. Lejeunea elongata (Austin) Austin Mastigolejeunea apiculata Steph. Phragmicoma elongata Austin Phragmicoma polymorpha Sande Lac. Ptycholejeunea elongata (Austin) Steph. Thysananthus elongatus (Austin) A.Evans Thysananthus paucidens Steph. ind? 37 M ind K O end K O Mo M SYZYGIELLA Spruce (Plagiochilaceae, or Jungermanniaceae [Crandall-Stotler & Stotler 2000]) Syzygiella eatonii (Austin ex A.Evans) Inoue [Inoue 1962; Miller 1963; Miller et al. 1983] Plagiochila eatonii Austin ex A.Evans end K M TARGIONIA L. (Targioniaceae) Targionia hypophylla L. [ Miller 1967; Miller et al. 1983] ind K O M SYMPHYOGYNA Nees & Mont. (Pallaviciniaceae) Symphyogyna semiinvolucrata Austin [Miller et al. 1983; Grolle & Piippo 1986] Pallavicinia baldwinii Austin Steetzia baldwinii Austin Symphyogyna baldwinii (Austin) Steph. Symphyogyna flavescens Steph. Symphyogyna picta Steph. TEMNOMA Mitt. (Blepharostomataceae, or Pseudolepicoleaceae [Crandall-Stotler & Stotler 2000]) Temnoma setigerum (Lindenb.) R.M.Schust. var. hawaiicum Inoue [Inoue 1974; Miller et al. 1983] TRICHOCOLEA Dumort. (Trichocoleaceae) Trichocolea gracillima Austin [Miller et al. 1983] Trichocolea tomentosa misapplied TROCHOLEJEUNEA Schiffn. (Lejeuneaceae) Trocholejeunea sandvicensis (Gottsche) Mizut. [Miller et al. 1983; Mizutani 1989] Acrolejeunea sandvicensis (Gottsche) Steph. Brachiolejeunea sandvicensis (Gottsche) A.Evans Brachiolejeunea sexplicata Steph. Cheilolejeunea sandvicensis Steph. Lejeunea cancellata Nees & Mont. Lejeunea sandvicensis (Gottsche) A.Evans TROPICAL BRYOLOGY 28 (2006) end M end M ind O H H H 38 STAPLES & IMADA: HAWAIIAN ANTHOCEROTES AND HEPATICS Lejeunea sandvicensis (Steph.) Steph. Lejeunea subligulata A.Evans Lejeunea subsquarrosa (Austin) Austin Mastigolejeunea sandvicensis (Gottsche) Steph. Phragmicoma bicolor Mont. Phragmicoma polygona Mitt. Phragmicoma sandvicensis Gottsche Phragmicoma subsquarrosa Austin TYLIMANTHUS Mitt. (Acrobolbaceae) Tylimanthus integrifolius A.Evans [Miller 1967; Miller et al. 1983] WIESNERELLA Schiffn. (Marchantiaceae, or Wiesnerellaceae [Crandall-Stotler & Stotler 2000]) Wiesnerella denudata (Mitt.) Steph. [Miller et al. 1983] Dumortiera denudata Mitt. Dubious Records Thirtyone hepatic taxa (30 species and 1 variety) were reported for the Hawaiian Islands by Miller et al. (1983), most without citing an island locality or a voucher specimen. We list these taxa here alphabetically, citing the page where Miller et al. reported the Hawaiian Islands as part of the distribution and, where applicable, cite any additional relevant literature. Bazzania javanica (Sande Lac.) Schiffn. was listed without island locality by Miller et al. (1983: 60). Bazzania mauiensis C.M.Cooke in W.E.Nicholson was listed for Hawai‘i island and tentatively from Maui by Miller et al. (1983: 61). Bazzania stolonifera (Sw.) Trevis. was listed without island locality by Miller et al. (1983: 64). Ceratolejeunea oculata (Gottsche) Steph. was listed without island locality by Miller et al. (1983: 78). Chiloscyphus coadunatus (Sw.) J.J.Engel & R.M.Schust. was listed (as Lophocolea coadunata) without island locality by Miller et al. (1983: 199). According to Gradstein (pers. comm. 2005) the occurrence of this heretofore exclusively neotropical hepatic in the Pacific is doubtful. Chiloscyphus pusillus (Steph.) J.J.Engel & R.M.Schust. was listed (as Lophocolea pusilla) without island locality by Miller et al. (1983: 202). Cololejeunea hirta (Steph.) S.Hatt. was listed for the island of Maui by Miller et al. (1983: 94). However, Tixier (1985: 290–291) did not cite any Hawaiian specimens for this tropical Asian and South Pacific species in his monograph of the Cololejeuneoideae. Cololejeunea planissima (Mitt.) Abeyw. was listed for the island of Kaua‘i by Miller et al. (1983: 98). However, Tixier (1985: 94–95) did not cite any Hawaiian specimens for this tropical Asian species in his monograph of the Cololejeuneoideae. end Mo M ind K O Mo M H Cyathodium aureonitens (Griff.) Schiffn. was listed without island locality by Miller et al. (1983: 104). Denotarisia linguifolia (De Not.) Grolle was listed without island locality by Miller et al. (1983: 108). Dumortiera hirsuta (Sw.) Nees var. latior Gottsche, Lindenb., & Nees was listed without island locality by Miller et al. (1983: 114). Frullania glomerata (Lehm. & Lindenb.) Nees & Mont. was listed without island locality by Miller et al. (1983: 127). Frullania intermedia (Reinw., Blume & Nees) Dumort. was listed without island locality by Miller et al. (1983: 130). Frullania stellatitexta Steph. was listed without island locality by Miller et al. (1983: 139). According to Miller (1953) the type specimen is from Fiji. Heteroscyphus argutus (Reinw., Blume & Nees) Schiffn. was listed without island locality by Miller et al. (1983: 149) and Piippo (1985a: 138). Lejeunea fissistipula (Steph.) Steph. was listed without island locality by Miller et al. (1983: 176). Lejeunea gibbiloba (Steph.) H.A.Mill. was listed without island locality by Miller et al. (1983: 177). Lepidozia holorhiza (Nees) Nees was listed without island locality by Miller et al. (1983: 190). Mastigophora diclados (Brid. ex F.Weber) Nees was listed without island locality by Miller et al. (1983: 225). Metzgeria comata Steph. was listed without island locality by Miller et al. (1983: 229). According to Kuwahara (1960) and So (2003c) this species is Melanesian (New Caledonia); its occurrence in Hawai‘i is dubious. Metzgeria scobina Mitt. was listed without island locality by Miller et al. (1983: 232). According to So (2003c) this species is distributed in SE Asia and Australasia; its occurrence in Hawai‘i is dubious. TROPICAL BRYOLOGY 28 (2006) STAPLES & IMADA: HAWAIIAN ANTHOCEROTES AND HEPATICS Monosolenium tenerum Griff. was listed without island locality by Miller et al. (1983: 236). Radula boryana (F.Weber) Nees ex Mont. was listed without island locality by Miller et al. (1983: 288). According to Grolle (2001) and Yamada (1979) the Asian reports of R. boryana are erroneous. The species’ presence in the Hawaiian Islands is doubtful. Radula complanata (L.) Dumort. was listed without island locality by Miller et al. (1983: 289). Yamada (1979) pointed out that it is unlikely this temperate species occurs in Hawai‘i. Radula decurrens Mitt. was listed without island locality by Yamada (1979: 314) and Miller et al. (1983: 289). The species was not accounted for in the most recent revision of Hawaiian Radula (So 2005b). Radula retroflexa Taylor was listed without island locality by Miller et al. (1983: 294), Yamada (1989: 230) and So (2005a). However the species was not included in the recent revision of Hawaiian Radula (So 2005b). Riccardia bipinnatifida (Colenso) Hewson was listed without island locality by Miller et al. (1983: 298). The occurrence of this New Guinea/Australia species (Hewson 1970) in the Hawaiian Islands is dubious. Riccardia grandiflora (Steph.) H.A.Mill. was listed (as Aneura grandiflora Steph.) without island locality by Miller et al. (1983: 300). Riccardia lilliena (Steph.) H.A.Mill. was listed (as Aneura lilliena Steph.) without island locality by Miller et al. (1983: 301). Riccardia loriana (Steph.) H.A.Mill. was listed without island locality by Miller et al. (1983: 301). The occurrence of this New Guinea/Australia species (Hewson 1970) in the Hawaiian Islands is dubious. Symphyogyna subsimplex Mitt. was listed without island locality by Miller et al. (1983: 323). Excluded Records Eight hepatics listed by Miller et al. (1983) from the Hawaiian Islands have been excluded from the flora based on reasonably conclusive evidence that they do not occur there. Relevant literature to support the exclusions is cited. Frullania astrolabea Steph. was described from a specimen collected during the voyage of the Astrolabe that purportedly came from the Hawaiian TROPICAL BRYOLOGY 28 (2006) 39 Islands. However, Miller (1953) and Hattori (1981) excluded the species from the Hawaiian flora because 1) the Astrolabe did not visit the Hawaiian Islands; 2) the species is otherwise unknown in Hawaii. Hattori (1981) suggested that the type material may have actually originated from New Zealand and it appears to be conspecific with F. scandens. Plagiochila gracilis Lindenb. & Gottsche was listed for the Hawaiian Islands without island locality by Miller et al. (1983: 260). According to So (2001b) this Asian species does not occur in Hawai‘i. Plagiochila owaihiensis Nees & Lindenb. was described from “oceani australis insula Owaihi.” According to Inoue (1976) and So (2000) this species is probably endemic to Tahiti; the type specimen was mislabeled as coming from “Owaihi.” Inoue definitely rejected this species from the Hawaiian flora; So tentatively did so. Plagiochila spathulifolia Mitt. was listed for Maui by Miller et al. 1983. According to Inoue (1982) and So (2001b) this Asian species does not occur in the Hawaiian Islands. Plagiochila tingens Steph. was listed without island locality by Miller et al. (1983: 270). Stephani’s protologue states the type came from the Hawaiian Islands, but no specimen has been located. So (2000) provisionally considered P. tingens a synonym of P. owaihiensis (q.v.) until such time as type material can be located for comparison. Prionolejeunea microdonta (Gottsche) Steph. was listed without island locality by Miller et al. (1983: 278). The species is excluded from the Hawaiian flora because Fulford & Sharp (1990: 70) do not mention Hawai‘i for the distribution of this neotropical species and it is unlikely to occur here. Stictolejeunea squamata (Willd. ex F.Weber) Schiffn. var. willdenoviana Gottsche was listed without island locality by Miller et al. (1983: 321). The neotropical S. squamata is not known to occur in Hawai‘i (Gradstein 1994). Symbiezidium transversale (Sw.) Trevis. (syn. S. bacciferum (Taylor) Steph., S. cryptocarpum (Mitt.) Steph.) was reported for the Hawaiian Islands based on a Menzies collection made in 1793. Gradstein and Van Beek (1985: 235–236) believe the label data are erroneous and have excluded this otherwise neotropical species from the Hawaiian flora. 40 STAPLES & IMADA: HAWAIIAN ANTHOCEROTES AND HEPATICS CROSS-REFERENCED LIST OF SYNONYMS AND MISAPPLIED NAMES Acrolejeunea sandvicensis See Trocholejeunea sandvicensis Anastrophyllum contractum misapplied See Jamesoniella robusta Anastrophyllum coriaceum See Jamesoniella robusta Anastrophyllum macrophyllum See Jamesoniella robusta Anastrophyllum piligerum misapplied See Anastrophyllum fissum Anastrophyllum ?rigidum See Jamesoniella robusta Aneura alba See Riccardia alba Aneura attenuata See Riccardia attenuata Aneura baldwinii See Riccardia baldwinii Aneura brunnea See Riccardia brunnea Aneura densiramea See Riccardia densiramea Aneura flaccida See Riccardia flaccida Aneura fleischeri See Riccardia fleischeri Aneura hamatiflora sphalm. See Riccardia hamatifolia Aneura hamatifolia See Riccardia hamatifolia Aneura hawaica See Riccardia hawaica Aneura multifida misapplied See Riccardia flaccida Aneura multispica See Riccardia multispica Aneura newellana See Riccardia newellana Aneura nigra See Riccardia nigra Aneura obtusifrons See Riccardia obtusifrons Aneura palmata misapplied See Riccardia multispica Aneura pauciramea See Riccardia pauciramea Aneura pectinata See Riccardia pectinata Aneura pinguis misapplied See Riccardia obtusifrons Aneura plana See Riccardia obtusifrons Aneura punahuina See Riccardia punahuina Aneura rockii See Riccardia rockii Aneura samoana See Riccardia alba Aneura tamariscina misapplied See Riccardia baldwinii Anthoceros baldwinii See Folioceros fuciformis Anthoceros breutelii See Notothylas breutelii Anthoceros communis See Phaeoceros carolinianus Anthoceros fuciformis See Folioceros fuciformis Anthoceros havaiensis See Phaeoceros carolinianus Anthoceros kaluensis See Folioceros fuciformis Anthoceros oahuensis See Phaeoceros carolinianus Anthoceros pusillus See Phaeoceros carolinianus Anthoceros validus See Phaeoceros carolinianus Archilejeunea mariana misapplied See Archilejeunea planiuscula Archilejeunea owahuensis See Archilejeunea planiuscula Aspiromitus hawaicus See Phaeoceros carolinianus Bazzania alternifolia See Metacalypogeia alternifolia Bazzania beecheyana See Bazzania patens Bazzania decurva See Bazzania praerupta Bazzania deflexa misapplied See Bazzania baldwinii Bazzania didricksenii See Bazzania didericiana Bazzania erosistipula See Bazzania patens Bazzania falcata See Bazzania cordistipula Bazzania ?integrifolia See Acromastigum integrifolium Bazzania ligulata See Bazzania brighamii Bazzania vittata misapplied See Bazzania minuta Blepharostoma sandwicense See Cephalozia sandvicensis Brachiolejeunea aliena See Spruceanthus polymorphus Brachiolejeunea apiculata See Spruceanthus polymorphus Brachiolejeunea polymorpha See Spruceanthus polymorphus Brachiolejeunea sandvicensis See Trocholejeunea sandvicensis Brachiolejeunea sexplicata See Trocholejeunea sandvicensis TROPICAL BRYOLOGY 28 (2006) STAPLES & IMADA: HAWAIIAN ANTHOCEROTES AND HEPATICS Calypogeia aeruginosa var. waialealeensis See Calypogeia aeruginosa Calypogeia alternifolia See Metacalypogeia alternifolia Calypogeia baldwinii See Mnioloma fuscum Calypogeia bidentula See Calypogeia tosana Calypogeia bifurca See Calypogeia arguta Calypogeia confertifolia See Calypogeia cuspidata Calypogeia fusca See Mnioloma fuscum Calypogeia hawaica See Calypogeia cuspidata Calypogeia minutistipula See Notoscyphus paroicus Calypogeia pacifica See Calypogeia arguta Calypogeia trichomanis misapplied See Metacalypogeia alternifolia Calypogeia waialealeensis See Calypogeia aeruginosa Cephalozia baldwinii See Cephaloziella baldwinii Cephalozia connivens misapplied See Cephalozia sandvicensis Cephalozia heteroica See Cephaloziella heteroica Cephalozia kilohanensis See Cephaloziella kilohanensis Cephalozia lilae See Cephaloziella lilae Cephalozia lucens See Metahygrobiella lucens Cephalozia maxima See Metahygrobiella maxima Cephalozia multiflora misapplied See Cephalozia sandvicensis Cheilolejeunea intertexta misapplied See Cheilolejeunea pyriflora Cheilolejeunea sandvicensis See Trocholejeunea sandvicensis Chiloscyphus beecheyanus See Chiloscyphus ciliolatus Chiloscyphus decurrens misapplied See Chiloscyphus laceratus Chiloscyphus gaudichaudii See Chiloscyphus ciliolatus Chiloscyphus mitteni misapplied See Chiloscyphus laceratus Cololejeunea kauaiensis See Cololejeunea longistyla Colura calyptrifolia misapplied See Colura tenuicornis Colura longicornis misapplied See Colura tenuicornis Colura tenuis See Colura tenuicornis Colurolejeunea tenuicornis See Colura tenuicornis Crossogyna robusta See Jamesoniella robusta Dendroceros brevicapsulus See Dendroceros crispus Dendroceros clintoni See Dendroceros crispus Dendroceros hawaicus See Dendroceros crispus Dicranolejeunea didericiana See Spruceanthus polymorphus Drepanolejeunea uncinata misapplied See Drepanolejeunea ungulata Dumortiera denudata See Wiesnerella denudata Dumortiera trichocephala See Dumortiera hirsuta Eulejeunea pacifica See Lejeunea pacifica Fimbriaria innovans See Asterella innovans Fimbriaria parva See Asterella parva Fimbriaria rechingeri See Marchantia crenata Frullania aongstroemii See Frullania neurota Frullania arietina misapplied See Frullania neurota Frullania cookei See Frullania oahuensis Frullania crenatiloba See Frullania apiculata Frullania elliptica See Frullania ericoides Frullania explicata See Frullania hypoleuca Frullania gymnotis See Frullania hypoleuca Frullania helleri See Frullania meyeniana Frullania oceanica See Frullania apiculata Frullania papulirostra See Frullania hypoleuca Frullania parvilobula See Frullania hypoleuca Frullania piligera See Jubula hutchinsiae subsp. javanica Frullania ptychantha misapplied See Frullania hypoleuca Frullania sandvicensis in part ? See Frullania ericoides Frullania sandvicensis See Frullania neurota Frullania squarrosa See Frullania ericoides Gamoscyphus sandvicense See Odontoschisma sandvicense TROPICAL BRYOLOGY 28 (2006) 41 42 STAPLES & IMADA: HAWAIIAN ANTHOCEROTES AND HEPATICS Gymnanthe bolanderi See Notoscyphus lutescens Harpalejeunea anderssonii See Drepanolejeunea anderssonii Harpalejeunea falsinervis See Drepanolejeunea pseudoneura Harpalejeunea owaihiensis See Lepidolejeunea involuta Harpalejeunea pseudoneura See Drepanolejeunea pseudoneura Harpalejeunea ungulata See Drepanolejeunea ungulata Herbertus longifissus misapplied See Herbertus helleri Herbertus parisii, in part See Herbertus helleri Herbertus sakuraii misapplied See Herbertus hawaiiensis Herbertus sanguineus See Herbertus helleri Herbertus setigerus, in part See Herbertus helleri Herpetium cordistipulum See Bazzania cordistipula Herpetium patens See Bazzania patens Herpocladium bidens, in part See Acromastigum integrifolium, Herbertus gracilis Herpocladium gracile See Herbertus gracilis Herpocladium tenacifolium misapplied See Acromastigum integrifolium Hygrolejeunea owaihiensis See Lepidolejeunea involuta Jamesoniella coriacea See Jamesoniella robusta Jamesoniella macrophylla See Jamesoniella robusta Jamesoniella subulata See Liochlaena subulata Jubula piligera See Jubula hutchinsiae subsp. javanica Jubula setacea See Jubula hutchinsiae subsp. javanica Jubula vittata See Frullania apiculata Jungermannia apiculata See Frullania apiculata Jungermannia australis See Lepidozia australis Jungermannia baldwinii See Jungermannia micrantha Jungermannia bolanderi See Notoscyphus lutescens Jungermannia callithrix misapplied See Jungermannia micrantha Jungermannia caudifera See Odontoschisma subjulaceum Jungermannia ciliolata See Chiloscyphus ciliolatus Jungermannia conchaefolia See Pleurozia conchifolia var. conchifolia Jungermannia contracta misapplied See Jamesoniella robusta Jungermannia coriacea See Jamesoniella robusta Jungermannia crassifolia See Cephalozia sandvicensis Jungermannia cucullata See Metalejeunea cucullata Jungermannia esenbeckii See Anastrophyllum esenbeckii Jungermannia fusca See Mnioloma fuscum Jungermannia gigantea See Pleurozia gigantea Jungermannia graveolens See Geocalyx graveolens Jungermannia lucens See Metahygrobiella lucens Jungermannia lurida See Jungermannia micrantha Jungermannia lutescens See Notoscyphus lutescens Jungermannia macrophylla See Jamesoniella robusta Jungermannia mauii See Jungermannia micrantha Jungermannia multifida See Riccardia multifida Jungermannia muricata See Chiloscyphus muricatus Jungermannia nana misapplied See Jungermannia micrantha Jungermannia newellana See Jungermannia micrantha Jungermannia orcadensis See Anastrepta orcadensis Jungermannia ornithopodioides See Scapania ornithopodioides Jungermannia piligera misapplied See Anastrophyllum fissum Jungermannia praerupta See Bazzania praerupta Jungermannia rigida See Jamesoniella robusta Jungermannia robusta See Jamesoniella robusta Jungermannia sandwicensis See Cephalozia sandvicensis Jungermannia sexplicata See Jungermannia micrantha Jungermannia squarrosa See Frullania ericoides Jungermannia subulata See Liochlaena subulata Jungermannia trichophylla See Blepharostoma trichophyllum Kantia alternifolia See Metacalypogeia alternifolia TROPICAL BRYOLOGY 28 (2006) STAPLES & IMADA: HAWAIIAN ANTHOCEROTES AND HEPATICS Kantia arguta See Calypogeia arguta Kantia baldwinii See Mnioloma fuscum Kantia bidentula See Calypogeia tosana Kantia bifurca See Calypogeia arguta Kantia cuspidata See Calypogeia cuspidata Kantia rotundistipula See Metacalypogeia alternifolia Kantia tosana See Calypogeia tosana Kantia trichomanis misapplied See Calypogeia tosana Lejeunea aliena See Spruceanthus polymorphus Lejeunea anderssonii See Drepanolejeunea anderssonii Lejeunea calyptrifolia misapplied See Colura tenuicornis Lejeunea cancellata See Trocholejeunea sandvicensis Lejeunea ceatocarpa See Cololejeunea ceatocarpa Lejeunea elongata See Spruceanthus polymorphus Lejeunea gibbosa See Lopholejeunea nigricans Lejeunea hawaica See Lopholejeunea nigricans Lejeunea hawaiiensis See Lepidolejeunea involuta Lejeunea hillebrandii See Cololejeunea hillebrandii Lejeunea intertexta See Cheilolejeunea intertexta Lejeunea longifolia See Cololejeunea hillebrandii Lejeunea mannii See Lopholejeunea nigricans Lejeunea obcordata See Cololejeunea obcordata Lejeunea owaihiensis See Lepidolejeunea involuta Lejeunea sandvicensis See Trocholejeunea sandvicensis Lejeunea stenoschiza See Cheilolejeunea stenoschiza Lejeunea subligulata See Trocholejeunea sandvicensis Lejeunea subsquarrosa See Trocholejeunea sandvicensis Lejeunea ungulata See Drepanolejeunea ungulata Lepidozia filipendula See Lepidozia sandvicensis Lepidozia hawaica See Kurzia hawaica Lepidozia minutifolia See Kurzia hawaica Lepidozia pusilla See Lepidozia sandvicensis Lepidozia remyana See Lepidozia australis Lepidozia reptans misapplied See Lepidozia australis Lepidozia reptans var. australis See Lepidozia australis Lepidozia triceps See Lepidozia australis Leptocolea ceatocarpa See Cololejeunea ceatocarpa Leptocolea mamilliloba See Cololejeunea obcordata Leptocolea obcordata See Cololejeunea obcordata Leptocolea ovalifolia See Cololejeunea ovalifolia Leptocolea lanciloba See Cololejeunea lanciloba Lophocolea autoica See Chiloscyphus autoicus Lophocolea baldwinii See Chiloscyphus baldwinii Lophocolea bartlettii See Chiloscyphus bartlettii Lophocolea beecheyana See Chiloscyphus ciliolatus Lophocolea bicuspidata See Chiloscyphus bicuspidatus Lophocolea bidentata misapplied See Chiloscyphus ciliolatus Lophocolea breutelii misapplied See Chiloscyphus baldwinii Lophocolea ciliolata See Chiloscyphus ciliolatus Lophocolea columbica misapplied See Chiloscyphus baldwinii Lophocolea connata misapplied See Chiloscyphus greenwelliae Lophocolea gaudichaudii See Chiloscyphus ciliolatus Lophocolea greenwelliae See Chiloscyphus greenwelliae Lophocolea hawaiica See Chiloscyphus hawaiicus Lophocolea kilauensis See Chiloscyphus kilauensis Lophocolea muricata See Chiloscyphus muricatus Lophocolea newellana See Chiloscyphus ciliolatus Lophocolea orbigiana misapplied See Chiloscyphus baldwinii Lophocolea spinosa See Chiloscyphus spinosus Lophocolea trispinosa See Chiloscyphus ciliolatus TROPICAL BRYOLOGY 28 (2006) 43 44 STAPLES & IMADA: HAWAIIAN ANTHOCEROTES AND HEPATICS Lopholejeunea gibbosa See Lopholejeunea nigricans Lopholejeunea hawaica See Lopholejeunea nigricans Lopholejeunea mannii See Lopholejeunea nigricans Lopholejeunea owahuensis See Lopholejeunea nigricans Lopholejeunea owaihiensis See Lepidolejeunea involuta Lopholejeunea subnuda See Lopholejeunea nigricans Lophozia orcadensis See Anastrepta orcadensis Madotheca acutifolia See Porella acutifolia Madotheca hawaiiensis See Porella acutifolia Marchantia antiqua See Marchantia marginata Marchantia chenopoda misapplied See Marchantia crenata Marchantia cruciata See Lunularia cruciata Marchantia cuspidisquama See Marchantia crenata Marchantia disjuncta misapplied See Marchantia crenata, Marchantia marginata Marchantia eatonii See Marchantia marginata Marchantia furciloba See Marchantia crenata Marchantia hirsuta See Dumortiera hirsuta Marchantia innovans See Asterella innovans Marchantia nitida misapplied See Marchantia marginata Marchantia paleacea misapplied See Marchantia marginata Marchantia punaluana See Marchantia crenata Marchantia rechingeri See Marchantia crenata Marchantia sandvicensis See Marchantia crenata Marchantia trichocephala See Dumortiera hirsuta Mastigobryum alternifolium See Metacalypogeia alternifolia Mastigobryum baldwinii See Bazzania baldwinii Mastigobryum beecheyanum See Bazzania patens Mastigobryum biapiculatum See Herbertus gracilis Mastigobryum bispinosum See Bazzania emarginata Mastigobryum brighami See Bazzania brighamii Mastigobryum cordistipulum See Bazzania cordistipula Mastigobryum didericiana See Bazzania didericiana Mastigobryum didericiana var. emarginata See Bazzania emarginata Mastigobryum didricksenii See Bazzania didericiana Mastigobryum emarginatum See Bazzania emarginata Mastigobryum grossidens See Bazzania patens Mastigobryum hawaiicum See Bazzania emarginata Mastigobryum honoluluense See Bazzania cordistipula Mastigobryum inaequabilis See Bazzania inaequabilis Mastigobryum integrifolium See Acromastigum integrifolium Mastigobryum kalihiense See Bazzania emarginata Mastigobryum ligulatum See Bazzania brighamii Mastigobryum minutum See Bazzania minuta Mastigobryum modestum See Bazzania brighamii Mastigobryum newellii See Bazzania cordistipula Mastigobryum nuuanuense See Bazzania nuuanuensis Mastigobryum obtusidens See Bazzania brighamii Mastigobryum pacificum See Bazzania inaequabilis Mastigobryum parvistipulum See Bazzania patens Mastigobryum patens See Bazzania patens Mastigobryum repandifolium See Bazzania nuuanuensis Mastigobryum rockii See Bazzania sandvicensis Mastigobryum rudimentare See Acromastigum integrifolium Mastigobryum sandwicense See Bazzania sandvicensis Mastigobryum subsandei See Bazzania cordistipula Mastigobryum tantalum See Bazzania patens Mastigolejeunea apiculata See Spruceanthus polymorphus Mastigolejeunea honoluluana See Lopholejeunea nigricans Mastigolejeunea sandvicensis See Trocholejeunea sandvicensis Mastigophora filipendula See Lepidozia sandvicensis TROPICAL BRYOLOGY 28 (2006) STAPLES & IMADA: HAWAIIAN ANTHOCEROTES AND HEPATICS Mastigophora gracilis See Herbertus gracilis Mastigophora parvifolia See Herbertus gracilis Mastigophora sandvicensis See Lepidozia sandvicensis Mastigophora triceps See Lepidozia australis Metacalypogeia fusca See Mnioloma fuscum Metacalypogeia montana var. verruculosa See Metacalypogeia alternifolia Metacalypogeia remotifolia See Metacalypogeia alternifolia Metzgeria decipiens See Metzgeria furcata Metzgeria dichotoma misapplied See Metzgeria furcata Metzgeria mauina See Metzgeria furcata Metzgeria molokaiensis See Metzgeria furcata Metzgeria oceanica See Metzgeria lindbergii Metzgeria papulosa See Metzgeria leptoneura Metzgeria pectinata See Metzgeria lindbergii Metzgeria quadriseriata See Metzgeria furcata Microlejeunea albicans See Metalejeunea cucullata Microlejeunea cucullata See Metalejeunea cucullata Microlejeunea erectifolia misapplied See Metalejeunea cucullata Microlejeunea sundaica See Metalejeunea cucullata Microlepidozia hawaica See Kurzia hawaica Microlepidozia mauiensis See Kurzia mauiensis Microlepidozia tayloriana See Kurzia tayloriana Mylia taylori misapplied See Jungermannia micrantha Nardia callithrix misapplied See Jungermannia micrantha Nardia exserta See Jungermannia hawaiica Nardia mauii See Jungermannia micrantha Nardia micrantha See Jungermannia micrantha Nardia truncata misapplied See Jungermannia micrantha Notoscyphus paroicus See Notoscyphus lutescens Odontoschisma gracile See Odontoschisma sandvicense Odontoschisma grandistipulum See Odontoschisma sandvicense Omphalanthus baldwinii See Mnioloma fuscum Pallavicinia baldwinii See Symphyogyna semiinvolucrata Pallavicinia cylindrica See Pallavicinia lyellii Pallavicinia simplex See Pallavicinia lyellii Pedinolejeunea lanciloba See Cololejeunea lanciloba Phaeoceros laevis subsp. carolinianus See Phaeoceros carolinianus Phragmicoma bicolor See Trocholejeunea sandvicensis Phragmicoma elongata See Spruceanthus polymorphus Phragmicoma polygona See Trocholejeunea sandvicensis Phragmicoma polymorpha See Spruceanthus polymorphus Phragmicoma sandvicensis See Trocholejeunea sandvicensis Phragmicoma subnuda See Lopholejeunea nigricans Phragmicoma subsquarrosa See Trocholejeunea sandvicensis Physiotium conchaefolia See Pleurozia conchifolia var. conchifolia Physiotium evolutum See Pleurozia subinflata Physiotium gigantea See Pleurozia gigantea Physiotium subinflatum See Pleurozia subinflata Physocolea ceatocarpa See Cololejeunea ceatocarpa Physocolea cookei See Cololejeunea cookei Physocolea hillebrandii See Cololejeunea hillebrandii Physocolea kauaiensis See Cololejeunea longistyla Physocolea longistyla See Cololejeunea longistyla Physocolea mamilliloba See Cololejeunea obcordata Plagiochasma cordatum misapplied See Plagiochasma japonicum Plagiochila acutiuscula See Plagiochila baldwinii Plagiochila adiantoides misapplied See Plagiochila deflexa Plagiochila arbuscula misapplied See Plagiochila deflexa Plagiochila askenasii See Plagiochila deflexa Plagiochila belangeriana misapplied See Plagiochila deflexa TROPICAL BRYOLOGY 28 (2006) 45 46 STAPLES & IMADA: HAWAIIAN ANTHOCEROTES AND HEPATICS Plagiochila biserialis misapplied See Plagiochila maunakeana Plagiochila caespitosa See Plagiochila deflexa Plagiochila combinata See Plagiochilion combinatum Plagiochila deltoidea misapplied See Plagiochila hoei Inoue Plagiochila eatonii See Syzygiella eatonii Plagiochila flava misapplied See Plagiochila baldwinii Plagiochila frondescens misapplied See Plagiochila mauiensis Plagiochila haleakalensis See Plagiochila gaudichaudii Plagiochila mannakeana See Plagiochila maunakeana Plagiochila mannii See Plagiochila gaudichaudii Plagiochila marriensis See Plagiochila mauiensis Plagiochila modesta See Plagiochila grossispina Plagiochila mollucensis See Plagiochila remyana Plagiochila nahikuna See Plagiochila deflexa Plagiochila oahuna See Plagiochilion combinatum Plagiochila oppositifolia See Plagiochilion combinatum Plagiochila owaihiensis misapplied See Plagiochila remyana Plagiochila patula See Plagiochila deflexa Plagiochila rockii See Plagiochila deflexa Plagiochila simplex misapplied See Plagiochila fissidentoides Plagiochila subtilis See Plagiochila fissidentoides Plagiochila tenuis misapplied See Plagiochila gaudichaudii Plagiochila variegata misapplied See Plagiochila fissidentoides Plagiochila yokogurensis subsp. remyana See Plagiochila remyana Plectocolea mauii See Jungermannia micrantha Plectocolea micrantha See Jungermannia micrantha Plectocolea submicrantha See Jungermannia micrantha Pleuroschisma pearsonii See Bazzania didericiana Pleurozia cochleariformis See Pleurozia conchifolia var. conchifolia Pleurozia evoluta See Pleurozia subinflata Pleurozia purpurea misapplied See Pleurozia conchifolia var. conchifolia Porella hawaiica sphalm. See Porella acutifolia Porella hawaiiensis See Porella acutifolia Porella laevigata misapplied See Porella hoeana Ptycholejeunea elongata See Spruceanthus polymorphus Pycnolejeunea stenoschiza See Cheilolejeunea stenoschiza Radula aneurismalis misapplied See Radula cavifolia Radula cordiloba See Radula javanica Radula excisiloba See Radula cordata Radula lejeuneaformis See Radula gracilis Radula mannii See Radula cordata Radula pallens misapplied See Radula javanica Radula pauciramea See Radula acutangula Radula perrotettii misapplied See Radula javanica, Radula reflexa, Radula sullivantii Radula xalapensis misapplied? See Radula cordata Riccardia multifida misapplied See Riccardia flaccida Riccardia palmata misapplied See Riccardia multispica Riccardia samoana See Riccardia alba Rupinia cordata misapplied See Plagiochasma japonicum Saccogyna bolanderi See Notoscyphus lutescens Scapania ‘manina’ sphalm. See Scapania mauina Scapania dentata var. oakesii misapplied See Scapania grossidens Scapania nemorosa misapplied See Scapania hawaica Scapania oakesii See Scapania grossidens Scapania planifolia See Scapania ornithopodioides Scapania undulata misapplied See Scapania paucidens Schisma gracilis See Herbertus gracilis Schisma helleri See Herbertus helleri Schisma juniperina misapplied See Herbertus helleri Schisma juniperinum var. sanguineum See Herbertus sanguineus TROPICAL BRYOLOGY 28 (2006) STAPLES & IMADA: HAWAIIAN ANTHOCEROTES AND HEPATICS Schisma longifissa misapplied See Herbertus helleri Schisma sanguineum See Herbertus sanguineus Schisma setigera See Herbertus helleri Schistochila cookei See Fulfordistria cookei Sendtnera gracilis See Herbertus gracilis Sendtnera tristicha See Herbertus gracilis Solenostoma exsertum See Jungermannia hawaiica Solenostoma hawaiicum See Jungermannia hawaiica Sphagnoecetis gracilis See Odontoschisma sandvicense Sphagnoecetis sandvicense See Odontoschisma sandvicense Spruceanthus marianus misapplied See Archilejeunea planiuscula Steetzia baldwinii See Symphyogyna semiinvolucrata Steetzia cylindrica See Pallavicinia lyellii Strepsilejeunea owaihiensis See Lepidolejeunea involuta Symphyogyna baldwinii See Symphyogyna semiinvolucrata Symphyogyna flavescens See Symphyogyna semiinvolucrata Symphyogyna picta See Symphyogyna semiinvolucrata Thysananthus elongatus See Spruceanthus polymorphus Thysananthus paucidens See Spruceanthus polymorphus Trachylejeuenea oahuensis See Cheilolejeunea decursiva Trichocolea tomentosa misapplied See Trichocolea gracillima TROPICAL BRYOLOGY 28 (2006) 47