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Zool. Surv. India Fauna of Uttar Pradesh, State Fauna Series, 22 (Part-2) : 395-426, 2015 INSECTA : COLEOPTERA : CARABIDAE R. K. KUSHWAHA and V. D. HEGDE* * Zoological Survey of India, M-Block, New Alipore, Kolkata-700 053 E-Mail : ravi_kumar3566@yahoo.com; hegde67@yahoo.co.in INTRODUCTION Family Carabidae is one of the largest family under order Coleoptera. Beetles of the Family Carabidae are also known as Ground Beetles. They are elongate, oblongate, oval, flat, cylindrical and sometimes rounded. The size varies from 1.5 mm to 45-50 mm. These beetles are carnivorous in both larval and adult stages and very few are carnivore in larval stage and herbivore in adult stage. Most of them are nocturnal, hiding under any objects during day time and very few are diurnal. They live under stone and bricks,under bark, under logs, at light, vicinity of water and sand bank of water bodies etc. These beetles can be recognized by hind coxae fused to metasternum and not dilated; first abdominal sternite completely divided; antennae filiform, 11 segmented and generally segment 1-4 glabrous, rest pubescent; 5-5-5 tarsal arrangement. Family Carabidae comprises 34,275 species in the world (Lorenz, 2005), however nearly 2000 species are represented in India (Saha, 1995). These beetles are cosmopolitan in distribution. Carabid beetles were already reported from following districts of Uttar Pradesh state, Allahabad, Bareilly, Bijnor, Chandauli, Chitrakoot, Faizabad, Fatehpur, Gorakhpur, Lakhimpur Kheri, Lucknow, Saharanpur, Sitapur, Sultanpur and Lalitpur. STUDY AREA Uttar Pradesh is the fourth largest state of India with 75 districts. It is surrounded by Nepal, Uttarakhand, Himachal Pradesh, Haryana and Delhi in the north; by Rajasthan and Madhya Pradesh in west; by Chhattisgarh in south; by Jharkhand and Bihar in the East. Most of the land are dominated by plain land. The state was surveyed by different survey parties of Zoological Survey of India. During study the specimens from Allahabad, Agra, Auraiya, Azamgarh, Bahraich, Ballia, Barabanki, Bareilly, Basti, Chandauli, Chitrakoot, Deoria, Etawah, Fatehpur, Ghazipur, Gonda, Hardoi, Jalaun, Jhansi, Lakhimpur Kheri, Lalitpur, Lucknow, Mathura, Mau, Meerut, Mirzapur, Pilibhit, Rae Bareilly, Saharanpur, Shahjahanpur, Siddharth nagar, Sitapur, Sultanpur and Varanasi districts were collected and identified. MATERIAL AND METHODS During the study period, different districts were surveyed and specimens collected by different teams of Zoological Survey of India. The specimens collected and deposited in Coleoptera Section from earlier surveys were also taken into consideration. Specimens were collected by different method like sweeping, dusting, hand picking and at light. Preserved specimens were relaxed, set-pinned and labelled. Specimens were sorted out in different categories and identification was carried out with the help of available literatures i.e. Andrewes (1919, 1920, 1922, 1929, 1935) and others. Taxa, first time reported from State are marked with one asterisk (*) and first time reported from India are marked with two asterisk (**). Specimens were studied under Lietz Stereo Binocular. The photographs of specimens are taken with the help of Leica M 205A stereo-zoom microscope, Nokia N73 and C5 phone camera. 396 Fauna of Uttar Pradesh, State Fauna Series, 22 SYSTEMATIC LIST OF TAXA Subfamily 2. HARPALINAE Bonelli, 1810 Subfamily 1. PTEROSTICHINAE Bonelli, 1810 Tribe 3. Harpalini Bonelli, 1810 Tribe 1. Abacetini Chaudoir, 1872 Subtribe. Stenolophina Kirby, 1837 Genus 1. Abacetus Dejean, 1828 Genus 6. Acupalpus Latreille, 1829. 1. Abacetus submetallicus (Nietner) 9. Acupalpus horni Andrewes 2. Abacetus vitreus Andrewes * Genus 7. Pachytrachelus Chaudoir, 1852 3. Abacetus artus Andrewes * Pachytrachelus Sp. 4. Abacetus guttula Chaudoir * Genus 2. Cosmodiscus Sloane, 1907 5. Cosmodiscus picturatus Andrewes Tribe 2. Pterostichini Bonelli, 1810 Subtribe 1. Molopina Bonelli, 1810 Genus 3. Euryaptus Bates, 1892 6. Euryaptus assamensis Bates Subtribe 2. Pterostichina s.str. Genus 4. Pterostichus Bonelli, 1810 7. Pterostichus harmandi Tchitcherin Genus 5. Trigonotoma Dejean, 1828 8. Trigonotoma indica Brulle Genus 8. Stenolophus Latreille, 1829 10. Stenolophus bajaurae Andrewes Subtribe. Amblystomina Fauvel, 1889 Genus 9. Amblystomus Erichson, 1837 * 11. Amblystomus indicus (Nietner)* Genus 10. Anoplogenius Chaudoir, 1852 12. Anoplogenius patinalis Bates Genus 11. Barysomus Dejean, 1829 13. Barysomus semivittatus (Fabricius) Subtribe Harpalina Bonelli, 1810 Genus 12. Dioryche Macleay, 1825 14. Dioryche chinnada Andrewes KUSHWAHA and HEGDE : Insecta : Coleoptera : Carabidae Genus 13. Hypolithus Dejean, 1829 15. Hypolithus acutangulus Bates Genus 14. Kareya Andrewes, 1919 16. Kareya sublaevis (Bates) 17. Kareya sublaevis var. major (Bates) Genus 15. Lamprophonus Bates, 1889 18. Lamprophonus lucens Bates Genus 16. Oxycentrus Chaudoir, 1854 * Oxycentrus sp. Genus 17. Platymetopus Dejean, 1829 19. Platymetopus pictus Andrewes 397 32. Tetragonoderus taeniatus (Wiedeman) * 33. Tetragonoderus rhombophorus SchmidtGoebel** Genus 28. Mnuphorus Chaudoir, 1873 34. Mnuphorus discophorus Chaudoir Subfamily 4. ANTHIINAE Bonelli, 1813 Tribe 7. Anthiini Bonelli, 1813 Genus 29. Anthia Weber, 1801 35. Anthia sexguttata (Fabricius) Tribe 8. Helluonini Hope, 1838 Genus 30. Macrocheilus Hope, 1838 20. Platymetopus rugosus (Nietner) 36. Macrocheilus chaudoiri Andrewes* 21. Platymetopus flavilabris (Fabricius)* 37. Macrocheilus niger Andrewes Subtribe Anisodactylina Lacordaire, 1854 Genus 18. Gnathaphanus Macleay, 1825 22. Gnathaphanus rusticus Andrewes Gnathaphanus Sp. 38. Macrocheilus trimaculatus Olivier Genus 31. Omphra Reiche, 1842 Omphra Sp. Subfamily 5. APOTOMINAE Leconte, 1853 Subfamily 3 . LEBIINAE Bonelli, 1810 Tribe 9. Apotomini Leconte, 1853 Tribe 4. Lebiini Bonelli, 1810 Genus 32. Apotomus Illiger, 1807 Genus 19. Anchista Nietner, 1856 23. Anchista brunnea (Wiedeman) Genus 20. Bothynoptera Schaum, 1863 24. Bothynoptera dorsigera Schaum Genus 21. Catascopus Kirby, 1825 25. Catascopus facialis (Wiedeman) Genus 22. Dolichoctis Schmidt-Goebel, 1846 * 39. Apotomus hirsutulus Bates Subfamily 6. TRECHINAE Bonelli, 1810 Tribe 10. Bembidiini Stephens, 1827 Genus 33. Asaphidion Des Gozis, 1886 * 40. Asaphidion championi Andrewes* 41. Asaphidion indicum (Chaudoir)* Genus 34. Bembidion Latreille, 1802 26. Dolichoctis striata Schmidt-Goebel * 42. Bembidion loricatum Andrewes Genus 23. Miscelus Klug, 1834 43. Bembidion luniferum Andrewes 27. Miscelus javanus var. unicolor Putzeys Genus 24. Lachnoderma Macleay, 1873 * 28. Lachnoderma biguttatum Bates * Genus 25. Cymindoidea Laporte, 1833 * 29. Cymindoidea distigma Chaudoir * Tribe 5. Odacanthini Laporte, 1834 Genus 26. Odacantha Paykull, 1798 30. Odacantha bimaculata (Redtenbacher) Tribe 6. Cyclosomini Laporte, 1834 Genus 27. Tetragonoderus Dejean, 1829 * 31. Tetragonoderus arcuatus Dejean * 44. Bembidion niloticum (Dejean)* 45. Bembidion notatum (Andrewes)* 46. Bembidion sobrinum Boheman 47. Bembidion xanthacrum Chaudoir 48. Bembidion xanthochiton Andrewes * Genus 35. Tachys Stephens, 1828 49. Tachys peryphinus Bates** 50. Tachys babaulti Andrewes * 51. Tachys blandus Andrewes* 52. Tachys dulcis Andrewes 53. Tachys fumigatus Motchulsky 398 Fauna of Uttar Pradesh, State Fauna Series, 22 54. Tachys klugi (Nietner) 76. Chlaenius agilis Chaudoir 55. Tachys latus Peyron 77. Chlaenius bimaculatus lynx Chaudoir 56. Tachys latus var. virgatus Andrewes* 78. Chlaenius bioculatus Chaudoir 57. Tachys opalescens Andrewes** 79. Chlaenius chlorodius Dejean 58. Tachys ornatus Apetz 80. Chlaenius duvauceli (Dejean) 59. Tachys poecilopterus Bates 81. Chlaenius laetiusculus Chaudoir 60. Tachys politus Motchulsky 82. Chlaenius luculentus Andrewes 61. Tachys politus var. aspilotus Andrewes 83. Chlaenius pretiosus Chaudoir 62. Tachys ceylanicus (Nietner)* 84. Chlaenius scapularis Chaudoir* 63. Tachys latissimus var. gracilis (Motchulsky)* 85. Chlaenius trinototus Chaudoir* Subfamily 7. BROSCINAE Hope, 1838 Tribe 11. Broscini Hope, 1838 Genus 36. Axonya Andrewes, 1923 64. Axonya championi Andrewes Genus 37. Broscus Panzer, 1813 65. Broscus punctatus (Dejean) Subfamily 8. BRACHININAE Bonelli, 1810 Tribe 12. Brachinini Bonelli, 1810 Subtribe Mastacina Erwin, 1970 Genus 38. Mastax Fischer von Waldheim, 1825 * Mastax sp. Subtribe Brachinina Bonelli, 1810 Genus 39. Brachinus Weber, 1801 66. Brachinus hexagrammus Chaudoir 67. Brachinus modestusSchmidt-Goebel * 68. Brachinus stevensi Andrewes* 69. Brachinus suturellus Chaudoir Subtribe Pheropsophina Jeannel, 1948 Genus 40. Pheropsophus Solier, 1833 70. Pheropsophus assimilis Chaudoir ** 86. Chlaenius variipes Chaudoir 87. Chlaenius xanthospilus (Wiedeman) Genus 43. Harpaglossus Motchulsky, 1858 88. Harpaglossus opacus Chaudoir Genus 44. Rhopalopalpus Laferte, 1851 89. Rhopalopalpus janthinus (Redtenbacher) Tribe14. Oodini Laferte-Senectere, 1851 Genus 45. Oodes Dejean, 1826 90. Oodes subolivaceus Laferte Genus 46. Systolocranius Chaudoir, 1857 91. Systolocranius linea Wiedeman Tribe 15. Licinini Bonelli, 1810 Genus 47. Badister Schellenberg, 1806 * Badister sp. Genus 48. Diplocheila Brulle, 92. Diplocheila polita (Fabricius) 93. Diplocheila cordicollis Laferte * 94. Diplocheila (s. str.) daldorfi (Crotch)* Subfamily 10. CARABINAE Latreille, 1802 71. Pheropsophus catoirei (Dejean) Tribe 16. Carabini Latreille, 1802 72. Pheropsophus consularis Schmidt-Goebel* Genus 49. Carabus Linnaeus, 1758 73. Pheropsophus hilaris (Fabricius) 74. Pheropsophus lineifrons Chaudoir 95. Carabus barysomus Bates Subfamily 11. CICINDELINAE Latreille, 1802 Subfamily 9. LICININAE Bonelli, 1810 Tribe 17. Cicindelini Latreille, 1802 Tribe 13. Chlaeniini Brulle, 1834 Subtribe Cicindelina Latreille, 1802 Genus 41. Callistomimus Chaudoir, 1872 Genus 50. Cicindela Linnaeus, 1758 75. Callistomimus d’abreui Andrewes Genus 42. Chlaenius Dejean, 1826 96. Cicindela (Calochroa) octonotata Wiedemann 97. Cicindela (Calochroa) flavomaculata Hope KUSHWAHA and HEGDE : Insecta : Coleoptera : Carabidae 399 98. Cicindela (Chaetodera) albina Wiedemann 124. Scarites laticeps Andrewes * 99. Cicindela (Chaetodera) vigintiguttata Herbst 125. Scarites picicornis Dejean 100. Cicindela (Lophyra) catena Fabricius 126. Scarites planus Bonelli 101. Cicindela (Lophyra) cancellata intemperata Acciavatti & Pearson 127. Scarites punctum Wiedeman 102. Cicindela (Spilodia) vittigera Dejean 129. Scarites selene Schmidt-Goebel 103. Cicindela (Spilodia) multiguttata Dejean 130. Scarites semirugosus Chaudoir * 104. Cicindela (Lophyridia) angulata Fabricius 131. Scarites striaticeps Chaudoir* 128. Scarites puncticollis Chaudoir 132. Scarites subnitens Chaudoir 105. Cicindela (Lophyridia) plumigera macrograptina Acciavatti & Pearson 133. Scarites sulcatus Olivier* 106. Cicindela (Lophyridia) chloris Hope 134. Scarites semicircularis Mcleay ** 107. Cicindela (Eugrapha) bigemina Klug 108. Cicindela (Eugrapha) brevis Horn 109. Cicindela (Eugrapha) minuta Olivier 110. Cicindela (Eugrapha) cognata Wiedemann 111. Cicindela (Eugrapha) erudita Wiedemann 112. Cicindela (Eugrapha) agnata Fleutiaux Tribe 20. Clivinini Rafinasque, 1815 Genus 55. Clivina Latreille, 1802 135. Clivina attenuata Herbst (Scarites) 136. Clivina gamma Andrewes * 137. Clivina lobata Bonelli 138. Clivina striata Putzeys * 139. Clivina tranquebarica Bonelli 113. Cicindela (Eugrapha) venosa Kollar Tribe 21. Dyschiriini Kolbe, 1880 114. Cicindela (Eugrapha) grammophora Chaudoir Genus 56. Dyschirius Panzer, 1813 115. Cicindela (Eriodera) albopunctata Chaudoir* 140. Dyschirius hingstoni Andrewes 116. Cicindela (Myriochile) melancholica Fabricius 141. Dyschirius mitodes Andrewes 117. Cicindela (Myriochile) undulata Dejean 142. Dyschirius perfectus Andrewes * 118. Cicindela (Myriochile) atelesta Chaudoir 143. Dyschirius variabilisAndrewes Tribe 18. Megacephalini Laporte, 1834 144. Dyschirius verticalis Putzeys Subtribe Megacephalina Laporte, 1834 145. Dyschirius mahratta Andrewes * Genus 51. Megacephala Latreille, 1802 146. Dyschirius mahratta var. paucipunctus Andrewes* 119. Megacephala euphratica var. armeniaca Castelnau Subfamily 12. SCARITINAE Bonelli, 1810 Tribe 19. Scaritini Bonelli, 1810 Genus 52. Haplogaster Chaudoir, 1879 120. Haplogaster ovata Chaudoir Genus 53. Oxylobus Chaudoir, 1855 121. Oxylobus porcatus Fabricius Genus 54. Scarites Fabricius, 1775 122. Scarites dyschromus Chaudoir 123. Scarites indus Olivier 147. Dyschirius bengalensis Andrewes* Subfamily 13. MELAENINAE Allaud, 1934 Tribe 22. Cymbionotini Andrewes, 1933 Genus 57. Cymbionotum Baudi, 1864 148. Cymbionotum fascigerum (Chaudoir) Cymbionotum Sp. Tribe 23. Melaenini Dejean, 1831 Genera 58. Melaenus Dejean, 1831 * 149. Melaenus piger (Fabricius) * Subfamily 14. DRYPTINAE Bonelli, 1810 Tribe 24. Dryptini Bonelli, 1810 400 Fauna of Uttar Pradesh, State Fauna Series, 22 Genus 59. Drypta Latreille, 1796 Subfamily 19. OMOPHRONINAE Bonelli, 1810 150. Drypta argillacea Andrewes Tribe 31. Omophronini Bonelli, 1810 151. Drypta lineola Macleay Genus 69. Omophron Latreille, 1802 Tribe 25. Galeritini Leconte, 1853 Genus 60. Galerita Fabricius, 1801 152. Galerita indica Chaudoir Subfamily 15. CTENODACTYLINAE Castelnau, 1834 Tribe 26. Hexagonini Horn, 1881 Genus 61. Hexagonia Kirby, 1825 * 153. Hexagonia uninotata Andrewes * Subfamily 16. SIAGONINAE Bonelli, 1813 Tribe 27. Siagonini Bonelli, 1813 Genus 62. Luperca Castelnau, 1840 154. Luperca laevigata Fabricius Genus 63. Siagona Latreille, 1804 155. Siagona atrata Dejean* 156. Siagona baconi Chaudoir * 157. Siagona fabricii Andrewes 158. Siagona flesus (Fabricius) 159. Siagona pumila Andrewes * 160. Siagona dilutipes var. aligera Andrewes* Subfamily 17. PLATYNINAE Bonelli, 1810 Tribe 28. Platyninni Bonelli, 1810 Genus 64. Megalonychus Chaudoir, 1843 161. Megalonychus birmanicus Bates Genus 65. Onycholabis Bates, 1873 * 162. Onycholabis acutangulus Andrewes* Genus 66. Orthotrichus Peyron, 1856 163. Orthotrichus indicus Bates Subfamily 18. NEBRIINAE Laporte, 1834 Tribe 29. Nebriini Laporte, 1834 Genus 60. Nebria Latreille, 1802 164. Nebria xanthacra Chaudoir 165. Nebria xanthacra var. nigricollis Andrewes * Tribe 30. Notiophilini Motchulsky, 1850 Genus 68. Notiophilus Dumeril, 1806 166. Notiophilus orientalis Chaudoir 167. Omophron maculosus Chaudoir Subfamily 20. PAUSSINAE Latreille, 1807 Tribe 32. Paussini Latreille, 1807 Genus 70. Merismoderus Westwood, 1847 168. Merismoderus bensoni Westwood Genus 71. Platyrhopalus Westwood, 1838 169. Platyrhopalus denticornis Donovan 170. Platyrhopalus angustus Westwood 171. Platyrhopalus intermedius Benson Genus 72. Paussus Linnaeus, 1775 172. Paussus hearseyanus Westwood 173. Paussus soleatus Wasmann* 174. Paussus ploiophorus Benson Indications: * (First report from Uttar Pradesh) ** (First report from India) SYSTEMATIC ACCOUNTS Subfamily 1. PTEROSTICHINAE Bonelli, 1810 Tribe 1. Abacetini Chaudoir, 1872 Genus 1. Abacetus Dejean, 1828 1828. Abacetus Dejean, Spec. Gen. Col. 3: 195. 1. Abacetus submetallicus (Nietner) 1858. Distrigus submetallicus Nietner, Ann. Mag nat. Hist., (3) 2: 177. 1869. Abacetus chalceolus Chaudoir, Bull. Soc. Imp. Nat. Mosc., 1: 384. Material examined:1 ex., Allahabad, Fapamore, 04.ii.2009, Coll. V. D. Hegde & party (ZSI Reg. no. 20030/H4A); 5 exs., Sitapur, Sinhanipur Jheel, 23.ii.2011, Coll. V. D. Hegde & party, Ex. Wet Field (ZSI Reg. No. 20031-20035/H4A). Distribution: INDIA: Throughout India including Uttar Pradesh; Elsewhere: Cambodia, Laos, Myanmar, Nepal, Sri Lanka and Vietnam. 2. Abacetus vitreus Andrewes* 1942. Abacetus vitreus Andrewes, Proc. Roy. Ent. Soc. Lond., 11 (2): 32. KUSHWAHA and HEGDE : Insecta : Coleoptera : Carabidae Material examined: 1 ex., Sitapur, Sinhanipur Jheel, 23.ii.2011, Coll. V. D. Hegde & party, Ex. Wet Field (ZSI Reg. No. 20039/H4A). 401 Subtribe 2. Pterostichina s. str. Genus 4. Pterostichus Bonelli, 1810 1810. Pterostichus Bonelli, Mem. Acad. Sci. Turin. 18: 68. Distribution: INDIA: Manipur and Uttar Pradesh. 7. Pterostichus harmandi Tchitcherin 3. Abacetus artus Andrewes* 1900. Pterostichus harmandi Tchitcherin, Horae Soc. Ent. Ross., 34: 467. 1942. Abacetus artus Andrewes, Proc. Roy. Ent. Soc. Lond., 11(2): 32. Distribution: INDIA: Meghalaya, Uttar Pradesh and West Bengal. Material examined: 4 exs., Sitapur, Sinhanipur Jheel, 23.ii.2011, Coll. V. D. Hegde & party, Ex. Wet Field (ZSI Reg. No. 20040-20043/H4A). Genus 5. Trigonotoma Dejean, 1828 Distribution: INDIA (Nevinson Coll., British Museum) and Uttar Pradesh. 4. Abacetus guttula Chaudoir* 1869. Abacetus guttula Chaudoir, Bull. Soc. Imp. Nat. Mosc., 1: 374. Material examined: 1 ex., Chitrakoot, Pahari, 26.xi.2010, Coll. O. P. Srivastava and party, Ex. Crevices in Wheat Field Soil (ZSI Reg. No. 20044/ H4A). 1828. Trigonotoma Dejean, Spec. Gen. Col., 3:182. 8. Trigonotoma indica Brulle 1834. Trigonotoma indica Brulle, Hist. Nat des Ins., 4: 333. Distribution: INDIA: Assam, Delhi, Madhya Pradesh, Maharashtra, Tamil Nadu, Uttarakhand, Uttar Pradesh and West Bengal; Elsewhere: Sri Lanka. Subfamily 2. HARPALINAE Bonelli, 1810 Tribe 3. Harpalini Bonelli, 1810 Subtribe Stenolophina Kirby, 1837 Distribution: INDIA : Northern India including Madhya Pradesh and Uttar Pradesh; Elsewhere: Sri Lanka, Myanmar, and Iran. 1829. Acupalpus Latreille, in Cuviers’ Regne Animal (Ed. 2) 4: 391. Genus 2. Cosmodiscus Sloane, 1907 9. Acupalpus horni Andrewes 1907. Cosmodiscus Sloane, Proc. Linn. Soc. N. S. Wales, 32: 371. 5. Cosmodiscus picturatus Andrewes 1920. Cosmodiscus picturatus Andrewes, Ann. Mag nat. Hist. (9)5: 447. Genus 6. Acupalpus Latreille, 1829. 1923. Acupalpus horni Andrewes, Spol. Zeyl., 12: 237. Distribution: INDIA: Karnataka, Tamil Nadu, Uttarakh and and Uttar Pradesh; Elsewhere: Myanmar and Sri Lanka. Genus 7. Pachytrachelus Chaudoir, 1852 Distribution: INDIA : Andhra Pradesh, Odisha, Maharashtra and Uttar Pradesh. 1852. Pachytrachelus Chaudoir, Bull. Soc. Imp. Nat. Mosc., 1: 85. Tribe 2. Pterostichini Bonelli, 1810 Pachytrachelus Sp. Subtribe 1. Molopina Bonelli, 1810 Material examined: 1 ex. Jalaun, Orai, Machhar Chauraha, 17.x.2011, Coll. B. Lal & Party, Ex. At light. Genus 3. Euryaptus Bates, 1892 1892. Euryaptus Bates, Ann. Mus. Civ. Sto. Nat. Gen., 32: 355. Genus 8. Stenolophus Latreille, 1829 6. Euryaptus assamensis Bates 1829. Stenolophus Latreille, in Cuvier’s Regne Anim. (Ed. ii) 4: 391. 1892. Euryaptus assamensis Bates, Ann. Mus. Civ. Sto. Nat. Gen., 32: 356 (note). 10. Stenolophus bajaurae Andrewes Distribution: INDIA: Assam, Uttarakhand and Uttar Pradesh. 1924. Stenolophus bajaurae Andrewes, Mission Prov. Centr. l’Inde Reg. Occi. l’Himalaya 1914, par Guy Babault: 95. 402 Fauna of Uttar Pradesh, State Fauna Series, 22 Distribution: INDIA : Bihar, Himachal Pradesh, Jammu-Kashmir, Uttarakhand and Uttar Pradesh. Subtribe Amblystomina Fauvel, 1889 Genus 9. Amblystomus Erichson, 1837* 1837. Amblystomus Erichson, Kaf. Mark. Brand., 1: 59. 11. Amblystomus indicus (Nietner)* 1858. Megaristerus indicus Nietner, Ann. Mag. Nat. Hist. (3) 2: 428. 1930. Amblystomus indicus, Andrewes, Cat. Ind. Insects, Part 18, Carabidae: 19. Material examined: 1 ex., Jalaun, Chili near Orai, 15.x.2011, Coll. B. Lal & party, Ex. at light; 1 ex., Jhansi, Pathrora near Barua sagar, 11.x.2011, Alt. 165m., Coll. B. Lal & party, Ex. under leaf litter. Distribution: INDIA: Jharkhand and Uttar Pradesh; Elsewhere: Australia, Myanmar, Sri Lanka and Vietnam. Genus 10. Anoplogenius Chaudoir, 1852 1852. Anoplogenius Chaudoir, Bull. Soc. Imp. Nat. Mosc., 1: 88. 12. Anoplogenius patinalis Bates 1892. Anoplogenius patinalis Bates, Ann. Mus. Civ. Sto. Nat. Gen., 32: 346. Distribution: INDIA: Bihar, Uttarakhand and Uttar Pradesh; Elsewhere: Myanmar. Genus 11. Barysomus Dejean, 1829 1829. Barysomus Dejean, Spec. Gen. Col., 4: 56. 13. Barysomus semivittatus (Fabricius) 1798. Carabus semivittatus Fabricius, Suppl. Ent. Syst., 59. 1930. Barysomus semivittatus, Andrewes, Cat. Ind. Ins., Part 18. Carabidae: 38. Distribution: INDIA: Bihar, Madhya Pradesh, Odisha, Tamil Nadu and West Bengal; Elsewhere: China, Sri Lanka and Vietnam. Subtribe Harpalina Bonelli, 1810 Genus 12. Dioryche Macleay, 1825 1825. Dioryche Macleay, Ann. Jav., 21. 14. Dioryche chinnada Andrewes 1921. Dioryche chinnada Andrewes, Rec. Ind. Mus., 22: 342. Distribution: INDIA: Bihar, Madhya Pradesh, Maharashtra, Tamil Nadu, Uttar Pradesh and West Bengal; Elsewhere: Sri Lanka. Genus 13. Hypolithus Dejean, 1829 1829. Hypolithus Dejean, Spec. Gen. Col., 4: 166. 15. Hypolithus acutangulus Bates 1891. Hypolithus acutangulus Bates, Compt. Rend Soc. Ent. Belg., 331. Distribution: INDIA: Bihar, Delhi, Gujarat, Jharkhand, Madhya Pradesh, Maharashtra, Tamil Nadu, Uttarakhand and Uttar Pradesh; Elsewhere: Bangladesh, Indonesia and Sri Lanka. Genus 14. Kareya Andrewes, 1919 1919. Kareya Andrewes, Ann. Mag nat. Hist., (9) 3: 473. 16. Kareya sublaevis (Bates) 1891. Platymetopus sublaevis Bates, Compt. Rend soc. Ent. Belg., 330. 1919. Kareya sublaevis Andrewes, Ann. Mag nat. Hist., (9) 3: 473. Distribution: INDIA : Andhra Pradesh, Arunachal Pradesh, Bihar, Jharkhand, Karnataka, Madhya Pradesh, Maharashtra, Uttarakhand and Uttar Pradesh; Elsewhere: Cambodia, China, Laos, Myanmar, Thailand and Vietnam. 17. Kareya sublaevis var. major (Bates) 1891. Platymetopus sublaevis var. major Bates, Compt. Rend Soc. Ent. Belg., 330. 1919. Kareya sublaevis var. major Andrewes, Ann. Mag nat. Hist., (9) 3: 473. Distribution: INDIA: Jharkhand, Madhya Pradesh and Uttar Pradesh. Genus 15. Lamprophonus Bates, 1889 1889. Lamprophonus Bates, Ann. Mus. Civ. Sto. Nat. Gen., 27: 101. 18. Lamprophonus lucens Bates 1889. Lamprophonus lucens Bates, Ann. Mus. Civ. Sto. Nat. Gen., 27: 102. Distribution: INDIA: Andhra Pradesh, Bihar, Jharkhand, Madhya Pradesh, Maharashtra, Odisha, Tamil Nadu, Uttar Pradesh and West Bengal; Elsewhere: Cambodia, Indonesia, Laos, Myanmar, Sri Lanka and Vietnam. KUSHWAHA and HEGDE : Insecta : Coleoptera : Carabidae Genus 16. Oxycentrus Chaudoir, 1854* 403 1854. Oxycentrus Chaudoir, Bull. Soc. Imp. Nat. Mosc., 1: 345. Distribution: INDIA: Bihar, Gujarat, Karnataka, Madhya Pradesh, Maharashtra, Odisha, Uttarakhand, Uttar Pradesh; Elsewhere: Sri Lanka. Oxycentrus sp. Gnathaphanus Sp. Distribution: 5 exs., Hardoi, Hardoi, I. H. Bunglow, 10.x.2006, Coll. P. Mukhopadhyay & Party, Ex. At light. Material examined: 1 ex., Allahabad, Meja, near FRH, 21.viii.2007, Coll. V. D. Hegde & Party, Ex. Under leaf litter; 1 ex., Mau, Social Forestry, Vandevi, 21.ix.2010, Coll. V. D. Hegde & party, Ex. under bricks. Genus 17. Platymetopus Dejean, 1829 1829. Platymetopus Dejean, Spec. Gen. Col., 4: 68. 19. Platymetopus pictus Andrewes 1923. Platymetopus pictus Andrewes, Spol. Zeyl., 233. Distribution: INDIA: Karnataka, Tamil Nadu and Uttar Pradesh; Elsewhere: Saudi Arabia. 20. Platymetopus rugosus (Nietner) Subfamily 3. LEBIINAE Bonelli, 1810 Tribe 4. Lebiini Bonelli, 1810 Genus 19. Anchista Nietner, 1856 1856. Anchista Nietner, Journ. Asia. Soc. Beng., 25: 523. 23. Anchista brunnea (Wiedeman) 1857. Ophonus rugosus Nietner, Journ. Asia. Soc. Beng., 26: 150. 1823. Lebia brunnea Wiedeman, Zool. Mag., ii, 1: 59. 1927. Platymetopus rugosus, Andrewes, Ann. Mag nat. Hist., 19: 106. Distribution: INDIA: Bihar, Maharashtra and Uttar Pradesh; Elsewhere: Bangladesh and Sri Lanka. Distribution: INDIA: Andhra Pradesh, Jharkhand, Karnataka, Nilgiri Hills, Odisha, Tamil Nadu, Uttarakhand, Uttar Pradesh and West Bengal; Elsewhere: Sri Lanka and Victoria Point. 21. Platymetopus flavilabris (Fabricius)* 1798. Carabus flavilabris Fabricius, Suppl. Ent. Syst., 59. 1921. Platymetopus flavilabris, Andrewes, Rec. Ind. Mus., 22: 344. Material examined: 1 ex., Hardoi, Hardoi I. H. Bunglow, 10.x.2006, Coll. P. Mukhopadhyay & Party, Ex. At light. Distribution: INDIA: including Uttar Pradesh; Elsewhere: Sr Lanka. Subtribe Anisodactylina Lacordaire, 1854 Genus 18. Gnathaphanus Macleay, 1825 1825. Gnathaphanus Macleay, Ann. Jav., 20. 22. Gnathaphanus rusticus Andrewes 1920. Gnathaphanus rusticus Andrewes, Ann. Soc. Ent. Belg., 60: 107, 110. Material examined: 1 ex., Ballia, Ballia, 11.xi.2009, Coll. V. D. Hegde & Party; 3 exs., Ballia, Surhatal WLS, 12.ii.2009, Coll. V. D. Hegde & Party. Genus 20. Bothynoptera Schaum, 1863 1863. Bothynoptera Schaum, Journ. Ent., 2: 75. 24. Bothynoptera dorsigera Schaum 1863. Bothynoptera dorsigera Schaum, Journ. Ent., 2: 76, t.4, f.3. Distribution: INDIA: Meghalaya, Uttar Pradesh and West Bengal; Elsewhere: Vietnam. Genus 21. Catascopus Kirby, 1825 1825. Catascopus Kirby, Trans. Linn. Soc. Lond., 14: 94. 25. Catascopus facialis (Wiedeman) 1819. Carabus facialis Wiedemen, Zool. Mag. i, 3: 165. 1930. Catascopus facialis, Andrewes, Cat. Ind. Insects Part 18, Carabidae: 76. Distribution: INDIA: Assam, Throughout Northern India including Uttar Pradesh; Elsewhere: Myanmar. Genus 22. Dolichoctis Schmidt-Goebel, 1846* 1846. Dolichoctis Schmidt-Goebel, Faun. Col. Birm., 62. 26. Dolichoctis striata Schmidt-Goebel* 1846. Dolichoctis striata Schmidt-Goebel, Faun. Col. Birm., 62. 404 Fauna of Uttar Pradesh, State Fauna Series, 22 Material examined: 1 ex. Chitrakoot, 10.x.2001, Coll. M. Prasad and party. Distribution: INDIA: Andaman Island, Kerala and Uttar Pradesh; Elsewhere: Australia, Indonesia, Japan, Laos, Malaysia, Myanmar, Philippines, Sri Lanka, Thailand and Vietnam. Genus 23. Miscelus Klug, 1834 1834. Miscelus Klug, Jahrb. Ins., 82. 27. Miscelus javanus var. unicolor Putzeys 1845. Miscelus javanus var. unicolor Putzeys, Mem Soc. Liege., 2: 375. Distribution: INDIA: Bihar, Karnataka, Kashmir, Madhya Pradesh and Uttar Pradesh; Elsewhere: Australia,Cambodia, China, Laos, Malaysia, Myanmar, Sri Lanka, Thailand and Vietnam. Genus 24. Lachnoderma Macleay, 1873* 1873. Lachnoderma Macleay, Trans. Ent. Soc. N. S. Wales, 2: 321 28. Lachnoderma biguttatum Bates* 1892. Lachnoderma biguttatum Bates, Ann. Mus. Civ. Sto. Nat. Gen., 32: 424. Material examined: 1 ex., Lakhimpur Kheri, Dudhwa National Park, Salukapur, 02.iii.2013, Coll. V. D. Hegde & Party. Distribution: INDIA: Assam, Sikkim, Uttarakhand and Uttar Pradesh; Elsewhere: Myanmar. Genus 25. Cymindoidea Laporte, 1832* 1832. Cymindoidea Laporte, Ann. Soc. Ent. Fr., 1: 390. 29. Cymindoidea distigma Chaudoir* 1875. Cymindoidea distigma Chaudoir, Bull. Soc. Imp. Nat. Mosc., 2: 15. Material examined: 1 ex. Jalaun, Orai, Vohadpura, 16.x.2011, Alt. 82m, Coll. Coll. B. Lal & party. Distribution: INDIA: Uttarakhand, Uttar Pradesh and West Bengal. Tribe 5. Odacanthini Laporte, 1834 Genus 26. Odacantha Paykull, 1798 1798. Odacantha Paykull, Faun. Suec., 1: 169. 30. Odacantha bimaculata (Redtenbacher) 1844. Casnonia bimaculata Redtenbacher, Hugel’s kaschmir, 4, 2: 498. t.23. f.2. 1930. Odacantha bimaculata Andrewes, Cat. Ind. Insects, Part 18, Carabidae: 229. Distribution: INDIA: Throughout Northern India, extending Southwards to Karnataka and Malabar; Elsewhere: Borneo, Cambodia, Indonesia, Myanmar, Philippine Island and Sri Lanka. Tribe 6. Cyclosomini Laporte, 1834 Genus 27. Tetragonoderus Dejean, 1829* 1829. Tetragonoderus Dejean, Spec. Gen. Col., 4: 485. 31. Tetragonoderus arcuatus Dejean* 1829. Tetragonoderus arcuatus Dejean, Spec. Gen. Col., 4: 495. Material examined: INDIA: Uttar Pradesh, 4exs., Lucknow, Khurram Nagar, 16.ii.2011, Coll. V.D. Hegde, Ex. Under leaf litter; 1 ex., Fatehpur, Garhiwa, 16.iii.2012, GPS N 25° 54´ 55.3´´ E 80° 48´ 51.7´´, Coll. A. Sar & party; 2 exs., Barabanki, Banki, Mohammadpur, 20.iii.2012, N26°56´ 45.1´´ E81°10´25.0´´, Coll. A. Sar & Party; 1 ex., Allahabad, Fapamore, 04.ii.2009, Coll. V. D. Hegde & Party, Ex. Bank of Ganga River; 1 ex., Mau, Mau, 13.ii.2009, Coll. V. D. Hegde; 1 ex., Ballia, Surhatal WLS, 12.ii.2009, Coll. V. D. Hegde & Party; 1 ex., Mathura, Kumaran Nagar, 16.xi.2010, Coll. O. P. Srivastava & Party; 1 ex., Rae Bareilly, Bacthrawan, 09.vi.1989, Coll. A. K. Sanyal & Party; 1 ex., Azamgarh, Azamgarh, 10.ii.2009, Coll. V.D. Hegde & Party; 1 ex., Siddharth Nagar, 13.iii.2010, Coll. V. D. Hegde & Party, Ex. Agricultural Field; 1 ex.,Lucknow, Kuckrail Forest Area, 17.ii.2011, Coll. V. D. Hegde& Party, Ex. Under Leaf Litter; 1 ex., Lucknow, Khurram Nagar, 16.ii.2011, Coll. V. D. Hegde & party, Ex. under leaf litter (Reg. no. 20055/H4A); 1 ex., Lakhimpur Kheri, Dudhwa National Park, Sathiyana, 01.iii.2013, Coll. V. D. Hegde & Party. Distribution: INDIA : Assam, Bihar, Himachal Pradesh, Jharkhand, Madhya Pradesh, Maharashtra, Meghalaya, Pondicherry, Uttarakhand, Uttar Pradesh and West Bengal; Elsewhere: Bangladesh, Egypt, Iraq, Laos, Myanmar and Sudan. KUSHWAHA and HEGDE : Insecta : Coleoptera : Carabidae 32. Tetragonoderus taeniatus (Wiedeman)* 1823. Bembidium taeniatus Wiedeman, Zool. Mag. (2) 1: 62. 1850. Tetragonoderus taeniatus Chaudoir, Bull. Soc. Imp. Nat. Mosc., 1: 455. Material examined: 1 ex., Chandauli, 16.ii.2009, Coll. V.D. Hegde & Party, ZSI Reg. No. 20056/ H4A; 1 ex., Lakhimpur Kheri, Dudhwa National Park, Sathiyana, 27.ii.2011, Coll. V.D. Hegde & Party. Distribution: INDIA : Bihar, Himachal Pradesh Uttarakhand, Uttar Pradesh and West Bengal; Elsewhere: Myanmar. 33. Tetragonoderus rhombophorus Schmidt-Goebel** 1846. Tetragonoderus rhombophorus Schmidt-Goebel, Faun. Col. Birm., 93. Material examined: 1 ex., Laknimpur Kheri, Salukapur, Dudhwa National Park, GPS N28° 25´ 5´´ E80° 43´9´´, 02.iii.2013, Coll. V. D. Hegde & party (Reg. no. 20054/H4A). Distribution: INDIA : Uttar Pradesh; Elsewhere: Bangladesh, Laos and Myanmar. Genus 28. Mnuphorus Chaudoir, 1873 1873. Mnuphorus Chaudoir, Berl. Ent. Zeit., 17: 55. 34. Mnuphorus discophorus Chaudoir 1876. Mnuphorus discophorus Chaudoir, Bull. Soc. Imp. Nat. Mosc., 2: 69. Distribution: INDIA (Bihar, Himachal Pradesh, Uttarakhand and Uttar Pradesh). Subfamily 4. ANTHIINAE Bonelli, 1813 Tribe 7. Anthiini Bonelli, 1813 Genus 29. Anthia Weber, 1801 1801. Anthia Weber, Obs. Ent.: 17. 35. Anthia sexguttata (Fabricius) 1775. Carabus sexguttata Fabricius, Syst. Ent., 236. 405 Distribution: INDIA : Throughout Indian Plains including Uttar Pradesh. Tribe 8. Helluonini Hope, 1838 Genus 30. Macrocheilus Hope, 1838 1838. Macrocheilus Hope, Col. Man., 2: 166. 36. Macrocheilus chaudoiri Andrewes* 1919. Macrocheilus chaudoiri Andrewes, Trans. Ent. Soc., Lond. 67: 130. Material examined: 1 ex., Gonda, East of Ambedkar Chauraha, Civil Lines, 12.viii.2007, Coll. V. D. Hegde & Party, Ex. Under dry leaves (Reg. no. 20064/H4A). Distribution: INDIA: Bihar, Maharashtra, and Uttar Pradesh; Elsewhere: Bangladesh, Myanmar, Indonesia, Philippines, Laos, Cambodia, and China. 37. Macrocheilus niger Andrewes 1920. Macrocheilus niger Andrewes, Ann. Mag. Nat. Hist. (9)6: 497, 499. Distribution: INDIA: Andhra Pradesh, Bihar, Jharkhand, Karnataka, Madhya Pradesh, Maharashtra, Nilgiri Hills, Tamil Nadu andUttar Pradesh) and Sri Lanka. 38. Macrocheilus trimaculatus Olivier 1790. Carabus trimaculatus Olivier, Enc. Meth., 347. t. 179. f. 11. 1924. Macrocheilus trimaculatus, Andrewes, Ann. Mag nat. Hist., (9) 13: 469. Material examined: 1ex., Etah, Irrigation Bunglow, 08.iii.2008, Coll. P. Mukhopadhyay & party, Ex. at light. Distribution: INDIA : including Uttar Pradesh; Elsewhere: China, Laos, Myanmar, Malaysia, Philippines, Sri Lanka and Vietnam. Genus 31. Omphra Reiche, 1842 1842. Omphra Reiche, Ann. Soc. Ent. Fr., 330. 1930. Anthia sexguttata, Andrewes, Cat. Ind. Ins. Part 18 Carabidae: 31. Omphra Sp. Material examined: INDIA: 1ex., Sultanpur, Panchopiran, 23.iii.2012, GPS N26°16´19.7´´ E82° 05´13.0´´, Coll. A. Sar and Party, Ex. Under dry leaves. Material examined: 1 ex., Auraiya, Shankarpur, 07.ix.2009, Coll. V. D. Hegde & Party; 1ex., Lalitpur, Jailwara, 14.x.2011, Alt. 296m, Coll. B. Lal & Party, Ex. Under stone. 406 Fauna of Uttar Pradesh, State Fauna Series, 22 Subfamily 5. APOTOMINAE Leconte, 1853 42. Bembidion loricatum Andrewes Tribe 9. Apotomini Leconte, 1853 1922. Bembidion loricatum Andrewes, Ent. Month. Mag.: 239. Genus 32. Apotomus Illiger, 1807 1807. Apotomus Illiger, Mag. Ins., 6: 348. 2013. Bembidion loricatum, Kushwaha & Hegde, in Souvenir: Water and Biodiversity: 158. 1892. Apotomus hirsutulus Bates, Ann. Mus. Civ. Sto. Nat. Gen., 32: 287. Material examined: 1 ex., Barabanki, Banki, Mohammadpur, 20.iii.2012, N26°56´45.1´´ E 81°10´25´´, Coll. A. Sar and Party (Reg. no. 19581/ H4A). Distribution: INDIA: Throughout India including Uttar Pradesh; Elsewhere: Myanmar. Distribution: INDIA: Bihar, Uttarakhand and Uttar Pradesh; Elsewhere: Pakistan. Subfamily 6. TRECHINAE Bonelli, 1810 43. Bembidion luniferum Andrewes Tribe 10. Bembidiini Stephens, 1827 1924. Bembidion luniferum Andrewes, Ent. Month. Mag.: 192. 39. Apotomus hirsutulus Bates Genus 33. Asaphidion Des Gozis, 1886* 1886. Asaphidion Des Gozis, Reche. espe. typi. dequelq. ancien. genr., recti. syno., notes diver.: 6. 1831. Tachypus Dejean, Spec. Gen. Col., 5: 35. Distribution: INDIA: Uttarakhand and Uttar Pradesh. 44. Bembidion niloticum (Dejean)* 1831. Bembidium niloticum Dejean, Spec. Gen. Col., 5: 73. 40. Asaphidion championi Andrewes* 1924. Asaphidion championi Andrewes, in Mission Prov. Centr. l‘Inde Reg. Occi. l’Himalaya, 1914, par Guy Babault, Carab : 84. t.2.f.2. Material examined: 1ex., Saharanpur, Beharigarh, 17.vi.1971, Coll. J. C. Tripathi & Party. Distribution: INDIA: Assam, Himachal Pradesh, Uttarakhand, Uttar Pradesh and West Bengal. 41. Asaphidion indicum (Chaudoir)* 1850. Tachypus indicum Chaudoir, Bull. Soc. Imp. Nat. Mosc., 2: 189 (Note). 1930. Asaphidion indicum, Andrewes, Cat. Ind. Insects, Part 18, Carabidae: 35. Material examined: 1ex., Barabanki, Banki, Mohammadpur, 20.iii.2012, N26°56´45.1´´ E81° 10´ 25.0´´, Coll. A. Sar & Party; 1 ex., Lakhimpur Kheri, Dudhwa National Park, Sathiyana, 27.ii.2011, Coll. V. D. Hegde & party, Ex. Under leaf litter. Distribution: INDIA : Bihar, Haryana, Himachal Pradesh, Uttarakhand, Uttar Pradesh and West Bengal. Genus 34. Bembidion Latreille, 1802 1802. Bembidion Latreille, Hist. Nat. Crust. et. Ins., 3: 82. 2013. Bembidion niloticum, Kushwaha & Hegde, in Souvenir: Water and Biodiversity: 159. Material examined: 1 ex., Barabanki, Sultanpur, 21.iii.2012, A. Sar and Party; 1 ex., Hardoi, Neer, Bhais ka nala, 20.iii.2011, Coll. V. D. Hegde & Party, Ex. Wet land; 1 ex., Siddharth Nagar, 13.iii.2010, Coll. V. D. Hegde & Party, Ex. Agricultural Field. Distribution:INDIA: Haryana, Nagaland, Uttarakhand, Uttar Pradesh and West Bengal; Elsewhere: Cambodia, China, Egypt, Iraq, Iran, Japan, Laos, Pakistan, Saudi Arabia, Turkey and Vietnam. 45. Bembidion notatum (Andrewes)* 1922. Bembidium notatum Andrewes, Ent. Month. Mag.: 175. 2013. Bembidion loricatum, Kushwaha & Hegde, in Souvenir: Water and Biodiversity: 159. Material examined: 1 ex., Hardoi, Neer, Bhais ka Nala, 20.ii.2011, Coll. V. D. Hegde & party, Ex. in soil crevices; 2 exs., Siddharth nagar, Siddharth nagar, 13.iii.2010, Coll. V. D. Hegde & Party, Ex. Agricultural Field; 1 ex., Shahjahanpur, Khannaut River bank, 01.iii.2011, Coll. V. D. Hegde & Party, Ex. On river bed; 2 exs., Hardoi, Neer, Bhais ka Nala, 20.ii.2011, Coll. V. D. Hegde KUSHWAHA and HEGDE : Insecta : Coleoptera : Carabidae 407 & party, Ex. Wet land; 2 exs., Hardoi, Neer, Bhais Ka Nala, 20.ii.2011, Coll. V. D. Hegde & Party, Ex. in soil crevices (Reg. No. 19607-19608/H4A); 3 exs., Hardoi, Neer, Bhais ka Nala, 20.ii.2011, Coll. V. D. Hegde & party, Ex. Wet land (Reg. no. 19632-19634/H4A); 1 ex., Sitapur, Tendua Jheel, 24.ii.2011, Coll. V. D. Hegde & party (Reg. no. 19635/H4A). Sitapur, Tendua Jheel, 24.ii.2011, Coll. V.D. Hegde & party, Ex. wet land (Reg. no. 19618/H4A). Distribution: INDIA: Himachal Pradesh, Meghalaya, Uttarakhand, Uttar Pradesh and West Bengal. 1850. Bembidion xanthacrum Chaudoir, Bull. Soc. Imp. Nat. Mosc., 2: 175 (note). 46. Bembidion sobrinum Boheman 1848. Bembidium sobrinum Boheman, Ins. Caffr., 1: 232. 2013. Bembidion sobrinum, Kushwaha & Hegde, in Souvenir: Water and Biodiversity: 159. Material examined: 1 ex., Mathura, Kishori Kunj, 17.xi.2010, Coll. O. P. Srivastava & Party; 5 exs., Mathura, Kishori Kunj, 17.xi.2010, Coll. O. P. Srivastava & party, (Reg. No. 19613-19617/ H4A); 2 exs., Lucknow, Kuckrail Forest Area, 17.ii.2011, Coll. V. D. Hegde & Party, Ex. At pond edge; 3exs., Lucknow, Kuckrail Forest Area, 17.ii.2011, Coll. V. D. Hegde & Party, Ex. At pond edge (Reg. No. 19586, 19588, 19620/H4A); 1 ex., Fatehpur, Garhiwa, 16.iii.2012, GPS N25°54´ 55.3°´´ E80°48´51.7´´, Coll. A. Sar & Party; 1 ex., Barabanki, Sultanpur, 21.iii.2012, Coll. A. Sar & Party; 3 exs., Shahjahanpur, Khannaut River bank, 01.iii.2011, Coll. V. D. Hegde & Party, Ex. On river bed; 2 exs., Shahjahanpur, Khannaut River bank, 01.iii.2011, Coll. V. D. Hegde & Party, Ex. On river bed (Reg. no. 19621, 19605/H4A); 1 ex., Siddharth Nagar, Siddharth Nagar, 13.iii.2010, Coll. V. D. Hegde & Party, Ex. Agricultural Field (Reg. No. 19587/H4A); 1 ex., Ballia, 13.ii.2009, Coll. V. D. Hegde & party (Reg. no. 19589/H4A); 17 exs., Hardoi, Neer, Bhais Ka Nala, 20.ii.2011, Coll. V. D. Hegde & Party; 5 exs., Hardoi, Neer, Bhais Ka Nala, 20.ii.2011, Coll. V. D. Hegde & Party, Ex. In soil Crevices (Reg. No. 19601-19604, 19610/H4A); 20 exs., Hardoi, Neer, Bhais Ka Nala, 20.ii.2011, Coll. V. D. Hegde & Party, Ex. wet land (1959319600, 19609,19611, 19619,19622-19630/H4A); 1 ex., Etah, Awagarh, 22.iii.1990, Coll. B. K. Biswas& party (Reg. no. 19612/H4A); 1 ex., Distribution: INDIA : Plains of India including Uttar Pradesh; Elsewhere: Cambodia, East African Countries, Indonesia, Sri Lanka, Myanmar, South Africa and Thiland. 47. Bembidion xanthacrum Chaudoir 2013. Bembidion xanthacrum, Kushwaha & Hegde, in Souvenir: Water and Biodiversity: 159. Material examined: 1ex., Chandauli, 17.iv.2007, Coll. S.K. Chatterjee & party, Ex. Bank of Ganga River (Reg. No. 19606/H4A); 1 ex., Shahjahanpur, Khannaut river Bank, 01.iii.2011, Coll. V. D. Hegde & Party, Ex. On river bed; 1 ex., Allahabad, Fapamore, 04.ii.2009, Coll. V. D. Hegde & Party; 1 ex., Chandauli, Chandauli, 16.ii.2009, Coll. V.D. Hegde & Party (Reg. no. 19582/H4A); 1 ex., Allahabad, Karela Bag, 05.ii.2009, Coll. V. D. Hegde & Party, Ex. Yamuna River Bed (Reg. No. 19583/H4A); 1 ex., Sultanpur, Sultanpur, 15.x.2006, Coll. P. Mukhopadhyay & Party, Ex. Bank of Gomati River (Reg. No. 19584/H4A); 1 ex., Ballia, Ballia, 13.ii.2009, Coll. V. D. Hegde & Party, Ex. Bank of Ganga River (Reg. no. 19585/ H4A);1 ex., Shahjahanpur, Khannaut river Bank, 01.iii.2011, Coll. V. D. Hegde & Party, Ex. in soil crevices (Reg. no. 19631/H4A); 1 ex., Lakhimpur Kheri, Salukapur, Dudhwa National Park, GPS N28°25´5´´ E80°43´9´´, 02.iii.2013, Coll. V. D. Hegde & party (Reg. no. 20052/H4A). Distribution: INDIA : Andhra Pradesh, Assam, Bihar, Himachal Pradesh, Kerala, Madhya Pradesh, Maharashtra, Uttarakhand, Uttar Pradesh, Tamil Nadu and West Bengal; Elsewhere: Cambodia, Laos, Myanmar, Pakistan, Thailand and Vietnam. 48. Bembidion xanthochiton Andrewes* 1922. Bembidium xanthochiton Andrewes, Ent. Month. Mag.: 176. 2013. Bembidion xanthochiton, Kushwaha & Hegde, in Souvenir: Water and Biodiversity: 159-160. Material examined: 1 ex., Hardoi, Neer, Bhais Ka Nala, 20.ii.2011, Coll. V. D. Hegde & party, 408 Fauna of Uttar Pradesh, State Fauna Series, 22 Ex. wet land (Reg. no. 19637/H4A); 1 ex., Lucknow, Kuckrail Forest Area, 17.ii.2011, Ex. at pond edge (Reg. no. 19636/H4A). Distribution: INDIA: Himachal Pradesh, Uttarakhand, Uttar Pradesh and West Bengal. Genus 35. Tachys Stephens, 1828 1828. Tachys Stephens, Ill. Brit. Ent. 2: 2, 4. 49. Tachys peryphinus Bates** 1886. Tachys peryphinus Bates, Ann. Mag. Nat. Hist., (5)17: 153. Material examined: INDIA: Uttar Pradesh, 2 exs., Lalitpur, Jailwara, 14.x.2011, Alt. 296 m., Coll. B. Lal & Party, Ex. Near stream; 1 ex., Shahjahanpur, Khannaut river bank, 01.iii.2011, Coll. V. D. Hegde & Party, Ex. In soil crevices. Distribution: INDIA : Uttar Pradesh; Elsewhere: Sri Lanka. 50. Tachys babaulti Andrewes* 1924. Tachys babaulti Andrewes, in Mission Prov. Centr. l‘Inde Reg. Occi. l’Himalaya, 1914, par Guy Babault, Carab., : 87, t.2, f.1. 2013. Tachys babaulti, Kushwaha & Hegde, in Souvenir: Water and Biodiversity: 160. Material examined: 1 ex., Lalitpur, Jailwara, 14.x.2011, Alt. 296m., Coll. B. Lal & party, Ex. near stream (Reg. No. 19592/H4A). Distribution: INDIA : Kerala, Madhya Pradesh, Maharashtra, Tamil Nadu, Uttarakhand and Uttar Pradesh; Elsewhere: Pakistan. 51. Tachys blandus Andrewes* 1924. Tachys blandus Andrewes, Mission Prov. Centr. l’Inde Reg. Occi. l’Himalaya, 1914, par Guy Babault, Carab.: 85, t.1, f.8. 2013. Tachys blandus, Kushwaha & Hegde, in Souvenir: Water and Biodiversity: 160. Material examined: 2 exs., Mathura, Kishori Kunj, 17.xi.2010, Coll. O. P. Srivastava & party, (Reg. No. 19676, 19677/H4A); 1 ex., Lucknow, Kuckrail Forest Area, 17.ii.2011, Coll. V. D. Hegde & party, Ex. at pond edge (Reg. no. 19675/H4A); 2 exs., Jalaun, Orai, Ajnari, Alt. 89 m., 17.x.2011, Coll. B. Lal & party, Ex. leaf litter (reg. no. 19673, 19674/H4A). Distribution: INDIA: Andhra Pradesh, Jharkhand, Karnataka, Madhya Pradesh, Maharashtra, Punjab, Uttarakhand and Uttar Pradesh; Elsewhere: Pakistan. 52. Tachys dulcis Andrewes 1925. Tachys dulcis Andrewes, Ann. Mus. Civ. Sto. Nat. Gen. 51(3, 10): 395. Distribution: INDIA : Uttar Pradesh. 53. Tachys fumigatus Motchulsky 1851. Tachys fumigatus Motchulsky, Bull. Mosc., 2: 509. 2013. Tachys fumigatus, Kushwaha & Hegde, in Souvenir: Water and Biodiversity: 160. Material examined: 6 exs., Lucknow, Kuckrail Forest Area, 17.ii.2011, Coll. V. D. Hegde & party, Ex. at pond edge; 2 exs., Hardoi, Neer, Bhais Ka nala, 20.ii.2011, Coll. V. D. Hegde & Party, Ex. In soil crevices; 2 exs., Shahjahanpur, Khannaut River bank, 01.iii.2011, Coll. V. D. Hegde & Party, Ex. On river bed; 1 ex., Hardoi, Neer, Bhais Ka nala, 20.ii.2011, Coll. V. D. Hegde & Party, Ex. Wet land; 1 ex., Mathura, Kishori Kunj, 17.xi.2010, Coll. O. P. Srivastava & Party; 1 ex., Barabanki, Banki, Mohammadpur, GPS N26°56´45.1´´ E81°10´ 25.0´´, 20.iii.2012, Coll. A. Sar & Party (Reg. No. 19591/H4A); 06 exs., Sitapur, Tendua Jheel, 24.ii.2011, Coll. V. D. Hegde & Party, Ex. Wet land (Reg. no. 19651-19656/ H4A); 1 ex., Mathura, Kishori Kunj, 17.xi.2010, Coll. O. P. Srivastava & Party (Reg. no. 19657/H4A); 2 exs., Barabanki, Sultanpur, 21.iii.2012, Coll. A. Sar & party (Reg. no. 19658, 19659/H4A). Distribution: INDIA : including Uttar Pradesh; Elsewhere: Southern Asia from Japan to New Guinea and from Philippines to East Africa and Madagascar. 54. Tachys klugi (Nietner) 1858. Bembidium klugi Nietner, Ann. Mag. Nat. Hist. (3) 2: 423. 1886. Tachys klugi Bates, Ann. Mag. Nat. Hist. (5) 17: 155. 2013. Tachys klugi, Kushwaha & Hegde, in Souvenir: Water and Biodiversity: 160. Material examined: 4 ex., Mathura, Kishori Kunj, 17.xi.2010, Coll. O. P. Srivastava & party, (Reg. No. 19664-19667/H4A); 1 ex., Jalaun, Orai, KUSHWAHA and HEGDE : Insecta : Coleoptera : Carabidae Ajnari, Alt. 89 m., 17.x.2011, Coll. B. Lal & party (Reg. no. 19663/H4A). Distribution: INDIA: Common throughout India; Elsewhere: South East Asia. 55. Tachys latus Peyron 1858. Tachys latus Peyron, Ann. Soc. Ent. Fr.: 364, t. 9, f. 3. Distribution: INDIA: Bihar, Kashmir, Uttarakhand, Uttar Pradesh and West Bengal; Elsewhere: Bangladesh, Indonesia, Iraq, Myanmar, Sri Lanka, Turkey and Vietnam. 56. Tachys latus var. virgatus Andrewes* 1925. Tachys latus var. virgatus Andrewes, Ann. Mus. Civ. Gen., 51: 440. 2013. Tachys latus var. virgatus, Kushwaha & Hegde, in Souvenir: Water and Biodiversity: 161. Material examined: 1 ex., Sitapur, Tendua Jheel, 24.ii.2011, Coll. V. D. Hegde & party, Ex. Wet Land (Reg. no. 19668/H4A). Distribution: INDIA: Uttar Pradesh and West Bengal. 57. Tachys opalescens Andrewes** 1925. Tachys opalescens Andrewes, Ann. Mus. Civ. Gen., 51: 342 and 352. 2013. Tachys opalescens, Kushwaha & Hegde, in Souvenir: Water and Biodiversity: 161. Material examined: 1 ex., Auraiya, Fatehpur near Muradganj, 20.x.2011, Coll. B. Lal & party, Ex. Leaf litter (Reg. No. 19672/H4A); 1 ex., Lakhimpur Kheri, Dudhwa National Park, Sathiyana, 27.ii.2011, Coll. V. D. Hegde & party, Ex. Under leaf litter; 1 ex., Hardoi, Neer, Bhais Ka nala, 20.ii.2011, Coll. V. D. Hegde & Party, Ex. In soil crevices. Distribution: INDIA: Uttar Pradesh; Elsewhere: Sri Lanka. 58. Tachys ornatus (Apetz) 1854. Bembidium ornatus Apetz, Col. Brehm.: 12. 1935. Tachys ornatus, Andrewes, Faun. Brit. India, Coleoptera, Carabidae 2: 16, f. 4a. 2013. Tachys ornatus, Kushwaha & Hegde, in Souvenir: Water and Biodiversity: 161. Material examined: 1 ex., Shahjahanpur, 409 Khannaut River Bank, 01.iii.2011, Coll. V. D. Hegde & party, Ex. in soil crevices (Reg. No. 19662/H4A); 7 exs., Shahjahanpur, Khannaut River bank, 01.iii.2011, Coll. V. D. Hegde & Party, Ex. On river bed; 3 exs., Lucknow, Kuckrail Forest Area, 17.ii.2011, Coll. V. D. Hegde & party, Ex. at pond edge; 1 ex., Mathura, Kishori Kunj, 17.xi.2010, Coll. O. P. Srivastava & Party; 1 ex., Auraiya, Fatehpur nr. Muradganj, 20.x.2011, Coll. B. Lal & party, Ex. leaf litter (Reg. no. 19660/H4A); 1 ex., Lucknow, Kuckrail Forest Area, 17.ii.2011, Coll. V. D. Hegde & party, Ex. at pond edge (Reg. no. 19661/H4A). Distribution: INDIA: Jharkhand, Karnataka, Madhya Pradesh, Maharashtra, Odisha, Tamil Nadu, Uttarakhand and Uttar Pradesh; Elsewhere: Egypt, Ethiopia, Namibia, Philippines, Saudi Arabia, Sudan and Italy (?). 59. Tachys poecilopterus Bates 1873. Tachys poecilopterus Bates, Trans. Ent. Soc. Lond., 21: 331. 2013. Tachys poecilopterus, Kushwaha & Hegde, in Souvenir: Water and Biodiversity: 161. Material examined: 3exs., Barabanki, Sultanpur, 21.iii.2012, Coll. A. Sar and party, (Reg. No. 1966919671/H4A). Distribution: INDIA: Assam, Sikkim, Uttarakhand, Uttar Pradesh and West Bengal; Elsewhere: Cambodia, China, Indonesia, Laos, Malaysia, Myanmar, Singapore, Thailand and Vietnam. 60. Tachys politus Motchulsky 1851. Tachys politus Motchulsky, Bull. Soc. Imp. Nat. Mosc., 2: 509. 2013. Tachys politus, Kushwaha & Hegde, in Souvenir: Water and Biodiversity: 161. Material examined: 7 exs., Auraiya, Fatehpur near Muradganj, 20.x.2011, Coll. B. Lal & party, Ex. leaf litter (Reg. No. 19638-19641, 19643, 19644, 19647/H4A); 3 exs., Mathura, Kishori Kunj, Coll. O. P. Srivastava & party (Reg. no. 19642, 19645, 19646/ H4A); 1 ex., Jhansi, Ambabai, 12.x.2011, Coll. B. Lal & party, Ex. Dusting (Reg. no. 19648/H4A); 1 ex., Shahjahanpur, Sugarcane 410 Fauna of Uttar Pradesh, State Fauna Series, 22 Research Institute rest house campus, 09.x.2006, Coll. P. Mukhopadhyay & party (Reg. no. 19649/ H4A). Distribution: INDIA: including Uttar Pradesh; Elsewhere: throughout South-East Asia except Japan. 61. Tachys politus var. aspilotus Andrewes 1925. Tachys politus var. aspilotus Andrewes, Ann. Mus. Civ. Gen., 51: 449. 2013. Tachys politus var. aspilotus, Kushwaha & Hegde, in Souvenir: Water and Biodiversity: 160. Material examined: 1 ex., Auraiya, Fatehpur near Muradganj, 20.x.2011, Coll. B. Lal & party, Ex. leaf litter (Reg. No.19650/H4A); 1 ex., Lalitpur, Jailwara, Alt. 296m, 14.x.2011, Coll. B. Lal & Party, Ex. near stream (Reg. no. 19590/H4A). Subfamily 7. BROSCINAE Hope, 1838 Tribe 11. Broscini Hope, 1838 Genus 36. Axonya Andrewes,1923 1923. Axonya Andrewes, Ann. Mag. Nat. Hist., (9) 12: 679. 64. Axonya championi Andrewes 1923. Axonya championi Andrewes, Ann. Mag. Nat. Hist., (9) 12: 680. 2011. Boscus punctatus, Kushwaha & Pal, Advancements in Invertebrate Taxonomy and Biodiversity:139-146. Distribution: INDIA: Arunachal Pradesh, Assam, Himachal Pradesh, Mizoram, Sikkim, Uttarakhand, Uttar Pradesh and West Bengal. Genus 37. Broscus Panzer, 1813 1813. Broscus Panzer, Index Ent.: 62. 1828. Cephalotes Dejean, Spec. Gen. Col., 3: 426. Distribution: INDIA: Throughout India including Uttar Pradesh; Elsewhere: throughout South-East Asia except Japan. 62. Tachys ceylanicus (Nietner)* 1858. Bembidium ceylanicus Nietner, Ann. Mag. Nat. Hist., (3)2: 423. 1935. Tachys ceylanicus, Andrewes, Faun. Brit. Ind. Carabidae 2, Harpalinae, 1: 257. Material examined: 1 ex., Lakhimpur Kheri, Salukapur, Dudhwa National Park, GPS N28° 25´ 5´´E80°43´9´´, 02.iii.2013, Coll. V. D. Hegde & party (reg. no. 20053/H4A). Distribution: INDIA: Karnataka, Uttarakhand and Uttar Pradesh; Elsewhere: Sri Lanka. 63. Tachys latissimus var. gracilis (Motchulsky)* 1862. Elaphropus gracilis Motchulsky, Etud. Ent., 11: 36. 1928. Tachys latissimus var. gracilis, Andrewes, Trans. Ent. Soc. Lond. : 4, 20. Material examined: 1 ex., Lakhimpur Kheri, Dudhwa National Park, Sathiyana, 01.iii.2013, Coll. V. D. Hegde & Party. Distribution: INDIA: Bihar, Maharashtra, Meghalaya, Nicobar Island, Uttarakhand, Uttar Pradesh and West Bengal; Elsewhere: Indonesia, Myanmar and Thailand. 65. Broscus punctatus (Dejean) 1828. Cephalotes punctatus Dejean, Spec. Gen. Col., 3: 431. 1930. Broscus punctatus, Andrewes, Cat. Ind. Ins., Part 18. Carabidae: 55. Distribution: INDIA: north of the Ganges, including the whole Himalayan tract and Meghalaya; Elsewhere: China, Egypt, Iraq, Saudi Arabia and Turkey. Subfamily 8. BRACHININAE Bonelli, 1810 Tribe 12. Brachinini Bonelli, 1810 Subtribe Mastacina Erwin, 1970 Genus 38. Mastax Fischer von Waldheim, 1825* 1825. Mastax Fischer von Waldheim, Ent. Russ., 3: 111. Mastax sp. Material examined:1ex., Saharanpur, Rajaji sanctuary, Saharanpur road, 11.vi.1964, Coll. T. D. Soota & party, Ex. under stone; 1 ex., Chitrakoot, Pahari, 26.xi.2010, Coll. O. P. Srivastava & party, Ex. Crevices in wheat field soil; 1 ex., Rae Bareilly, Lohanipur, 06.iii.2010, Coll. V. D. Hegde & party, Ex. bank of Sai River. Subtribe Brachinina Bonelli, 1810 Genus 39. Brachinus Weber, 1801 1801. Brachinus Weber, Obs. Ent.: 22. KUSHWAHA and HEGDE : Insecta : Coleoptera : Carabidae 66. Brachinus hexagrammus Chaudoir 1876. Brachinus hexagrammus Chaudoir, Ann. Soc. Ent. Belg.: 55. Material examined: 8 exs., Jalaun, Chili near Orai, 15.x.2011, Coll. B. Lal & party, Ex. at light (Reg. no. 20070-20077/H4A). Distribution: INDIA : Throughout INDIA including Uttar Pradesh. 67. Brachinus modestus Schmidt-Goebel* 1846. Brachinus modestus Schmidt-Goebel, Faun. Col. Birm.: 73. Material examined: 1 ex., Jhansi, Pathrora near Barua Sagar, 11.x.2011, Alt. 165m, Coll. B. Lal & party, Ex. under leaf litter (Reg. no. 20082/H4A). Distribution: INDIA: Uttar Pradesh and West Bengal; Elsewhere: Myanmar and Vietnam. 68. Brachinus stevensi Andrewes* 1924. Brachinus stevensi Andrewes, Mission Prov. Centr. l‘Inde Reg. Occi. l‘Himalaya 1914, par Guy Babault: 107, t. 4. f. 2. Material examined: 4 exs., Sitapur, Sinhanipur Jheel, 23.ii.2011, Coll. V. D. Hegde & party, Ex. Crevices in soil (Reg. no. 20078-20081/H4A). Distribution: INDIA: Arunachal Pradesh, Assam, Himachal Pradesh, Uttarakhand, Uttar Pradesh and West Bengal. 69. Brachinus suturellus Chaudoir 1876. Brachinus suturellus Chaudoir, Ann. Soc. Ent. Belg.: 69. Distribution: INDIA: Himachal Pradesh, Jharkhand, Karnataka, Madhya Pradesh, Uttarakhand and Uttar Pradesh; Elsewhere: Myanmar and Vietnam(?). Subtribe Pheropsophina Jeannel, 1948 Genus 40. Pheropsophus Solier, 1833 1833. Pheropsophus Solier, Ann. Soc. Ent. Fr., 2(1): 461. 70. Pheropsophus assimilis Chaudoir** 1876. Pheropsophus assimilis Chaudoir, Ann. Soc. Ent. Belg.: 38. Material examined: 2 exs., Hardoi, Sandi Bird Sanctuary, 21.ii.2011, Coll. V. D. Hegde & party, 411 Ex. Soil crevices of ploughed field (Reg. no. 20083, 20084/H4A). Distribution: INDIA : Uttar Pradesh; Elsewhere: Thailand and China. 71. Pheropsophus catoirei (Dejean) 1825. Brachinus catoirei Dejean, Spec. Gen. Col., 1: 301. 1924. Pheropsophus catoirei, Andrewes, Mission Prov. Centr. l‘Inde Reg. Occi. l‘Himalaya 1914, par Guy Babault: 55. Material examined: 13 exs., Hardoi, Sandi Bird Sanctuary, 21.ii.2011, Coll. V. D. Hegde & party, Ex. Crevices of Soil; 1 ex., Hardoi, Sandi Bird Sanctuary, 21.ii.2011, Coll. V. D. Hegde & party, Ex. Crevices of Soil (Reg. no. 20088/H4A); 4 exs., Hardoi, Sandi Bird Sanctuary, 21.ii.2011, Coll. V. D. Hegde & party, Ex. crevices of ploughed field; 1 ex., Ballia F.R.H., 15.ix.2010, Coll. V. D. Hegde & party; 1 ex., Mirzapur, Chunar, 08.ii.2009, Coll. V. D. Hegde & party, Ex. Ganga river bed; 1 ex., Chandauli, Bhansi, 27.ix.2010, Coll. V. D. Hegde & party, Ex. under bricks; 1 ex., Bijnor, Indira park near Director Wildlife & forest office, 22.vii.2008, Coll. V. D. Hegde & party; 1 ex., Sitapur, Tendua Jheel, 24.ii.2011, Coll. V. D. Hegde & party, Ex. soil crevices of ploughed field; 1 ex., Ghaziabad, Moorta, 10.ii.2009, Coll. R. S. Mridha & party. Distribution: INDIA: Throughout India, including Westward to Kashmir and the Khyber Pass, Meghalaya, Andaman Island and Uttar Pradesh; Elsewhere: Myanmar, Pakistan and Sri Lanka. 72. Pheropsophus consularis Schmidt-Goebel* 1846. Pheropsophus consularis Schmidt-Goebel, Faun. Col. Birm., : 75. Material examined: 2 exs., Pilibhit, Mala Forest, 27.ix.1974, Alt. 250m, Coll. P. K. Maiti and party (Reg. no. 20085, 20086/H4A). Distribution: INDIA: Arunachal Pradesh, Assam, Bihar, Himachal Pradesh, Jharkhand, Odisha, Uttarakhand, Uttar Pradesh and West Bengal; Elsewhere: Bangladesh and Myanmar. 73. Pheropsophus hilaris (Fabricius) 1798. Carabus hilaris Fabricius, Suppl. Ent. Syst., 56. 412 Fauna of Uttar Pradesh, State Fauna Series, 22 1921. Pheropsophus hilaris, Andrewes, Trans. Ent. Soc. Lond., 69: 159. Uttar Pradesh and West Bengal; Elsewhere: Bangladesh. Material examined: 3exs.,Chandauli, Bansi, 27.ix.2010, Coll. V. D. Hegde & party, Ex. Under bricks; 1 ex., Chandauli, Bansi, 27.ix.2010, Coll. V. D. Hegde & party, Ex. Under bricks (Reg. no. 20087/H4A); 2 exs., Lakhimpur Kheri, Dudhwa National park, Sathiyana, GPS N28° 32´ 30´´ E80° 32´ 30´´, 01.iii.2013, Coll. V. D. Hegde & party. 78. Chlaenius bioculatus Chaudoir Distribution: INDIA: Throughout India including N. Andaman and Uttar Pradesh; Elsewhere: Myanmar and Pakistan. 74. Pheropsophus lineifrons Chaudoir 1850. Pheropsophus lineifrons Chaudoir, Bull. Soc. Imp. Nat. Mosc., 1: 80. Distribution: INDIA: Assam, Bihar, Himachal Pradesh, Jharkhand, Maharashtra, Uttarakhand, Uttar Pradesh and West Bengal. Subfamily 9. LICININAE Bonelli, 1810 Tribe 13. Chlaeniini Brulle, 1834 Genus 41. Callistomimus Chaudoir, 1872 1872. Callistomimus Chaudoir, Bull. Soc. Imp. Nat. Mosc., 1: 382. 75. Callistomimus d’abreui Andrewes 1921. Callistomimus d’abreui Andrewes, Proc. Zool. Soc. Lond., 1: 237 and 242, t.1, f.9. Distribution: INDIA: Bihar, Madhya Pradesh and Uttar Pradesh. Genus 42. Chlaenius Dejean, 1826 1826. Chlaenius Dejean, Spec. Gen. Col., 2: 297. 76. Chlaenius agilis Chaudoir 1856. Chlaenius agilis Chaudoir, Bull. Soc. Imp. Nat. Mosc., 2: 246. Distribution: INDIA: Arunachal Pradesh, Assam, Himachal Pradesh, Nim Nadi,Uttarakhand, Uttar Pradesh and West Bengal; Elsewhere: Cambodia, Laos and Vietnam. 77. Chlaenius bimaculatus lynx Chaudoir 1856. Chlaenius bimaculatus var. lynx Chaudoir, Bull. Soc. Imp. Nat. Mosc., 2:199. Distribution: INDIA: Bihar, Manipur, Odisha, 1856. Chlaenius bioculatus Chaudoir, Bull. Soc. Imp. Nat. Mosc., 2:198. Distribution: INDIA: Throughout India including Andaman Island; Elsewhere: Borneo, Cambodia, China, Indonesia, Japan, Laos, Myanmar, Sri Lanka and Vietnam. 79. Chlaenius chlorodius Dejean 1826. Chlaenius chlorodius Dejean, Spec. Gen. Col., 2: 365. Distribution: INDIA: Assam, Bihar, Madhya Pradesh, Maharashtra, Uttarakhand, Uttar Pradesh and West Bengal; Elsewhere: Bangladesh. 80. Chlaenius duvauceli (Dejean) 1831. Epomis duvauceli Dejean, Spec. Gen. Col., 5: 668. 1924. Chlaenius duvauceli, Andrewes, Mission Prov. Centr. l‚ Inde Reg. Occi. l‚Himalaya 1914, par Guy Babault: 25. Distribution: INDIA: Assam, Bengal, Bihar, Gujarat, Haryana, Maharashtra, Tamil Nadu and Uttar Pradesh. 81. Chlaenius laetiusculus Chaudoir 1856. Chlaenius laetiusculus Chaudoir, Bull. Soc. Imp. Nat. Mosc., 2: 248. Distribution: INDIA: Throughout Himalayan Tract including Assam and Uttar Pradesh; Elsewhere: Pakistan and Sri Lanka. 82. Chlaenius luculentus Andrewes 1920. Chlaenius luculentus Andrewes, Ent. Month. Mag.: 235, 236. Distribution: INDIA (Bihar, Maharashtra, Odisha, Uttar Pradesh and West Bengal). 83. Chlaenius pretiosus Chaudoir 1856. Chlaenius pretiosus Chaudoir, Bull. Soc. Imp. Nat. Mosc., 2: 288. Material examined: 1 ex. Agra, Kakuabaad, 29.viii.2009, Coll. V. D. Hegde & party, Ex. In soil crevices. Distribution: INDIA: Delhi, Gujarat, Rajasthan, Uttarakhand and Uttar Pradesh; Elsewhere: Pakistan and Sri Lanka. KUSHWAHA and HEGDE : Insecta : Coleoptera : Carabidae 84. Chlaenius scapularis Chaudoir* 1876. Chlaenius scapularis Chaudoir, Ann. Soc. Ent. Belg., 8: 98. Material examined: 3 ex., Chitrakoot, Pahari, 26.xi.2010, Coll. O. P. Srivastava & party, Ex. Crevices in Wheat field soil; 2 exs., Hardoi, Pihani, 30.vi.1989, Coll. K. K. Ray & party. Distribution: INDIA: Bihar, Chhattisgarh, Delhi, Himachal Pradesh, Karnataka, Madhya Pradesh, Tamil Nadu, Uttarakhand, Uttar Pradesh and West Bengal; Elsewhere: Sri Lanka. 85. Chlaenius trinototus Chaudoir* 1876. Chlaenius trinototus Chaudoir, Ann. Soc. Ent. Belg., 8: 279. Material examined: 1 ex., Lakhimpur Kheri, Dudhwa FRH Compound, 23.ix.1974, Coll. P. K. Maiti and Party, Ex. at light Distribution: INDIA: Bihar, Uttarakhand and Uttar Pradesh. 86. Chlaenius variipes Chaudoir 1856. Chlaenius variipes Chaudoir, Bull. Soc. Imp. Nat. Mosc., 2: 268. 413 1930. Harpaglossus opacus, Andrewes, Cat. Ind. Insects, Part 18, Carabidae: 175. Distribution: INDIA: Gujarat, Rajasthan, Tamil Nadu, Uttar Pradesh and West Bengal; Elsewhere: Sri Lanka. Genus 44. Rhopalopalpus Laferte, 1851 1851. Rhopalopalpus Laferte, Ann. Soc. Ent. Fr., 9(2): 212. 89. Rhopalopalpus janthinus (Redtenbacher) 1844. Chlaenius janthinus Redtenbacher, Hugel’s kaschmir iv, 2: 500. 1930. Rhopalopalpus janthinus, Andrewes, Cat. Ind. Insects Part 18, Carabidae: 290. Distribution: INDIA: Himachal Pradesh, Kashmir, Punjab, Uttarakhand and Uttar Pradesh; Elsewhere: Pakistan. Tribe 14. Oodini Laferte-Senectere, 1851 Genus 45. Oodes Dejean, 1826 1826. Oodes Dejean, Spec. Gen. Col., 2: 374. 90. Oodes subolivaceus Laferte 1851. Oodes subolivaceus Laferte, Ann. Soc. Ent. Fr.: 271 (note). Distribution: INDIA: Bihar, Delhi, Gujarat, Jharkhand, Madhya Pradesh and Uttar Pradesh; Elsewhere: Pakistan. Distribution: INDIA: Bihar, Jharkhand, Karnataka, Maharashtra, Uttarakhand, Uttar Pradesh and West Bengal; Elsewhere: Nepal and Sri Lanka. 87. Chlaenius xanthospilus (Wiedeman) Genus 46. Systolocranius Chaudoir, 1857 1821. Carabus xanthospilus Wiedeman, Germ. Mag. Ent., 4: 115. 1857. Systolocranius Chaudoir, Bull. Soc. Imp. Nat. Mosc. 2: 23. 1921. Chlaenius xanthospilus Andrewes, Trans. Ent. Soc. Lond.: 167. 91. Systolocranius linea (Wiedeman) 1821. Oodes linea Wiedeman, Germ. Mag., 4: 113. Material examined: 1ex., Azamgarh, Kusum Lodge, 21.iii.2009, Coll. A. Bal and Party, Ex. at light. Distribution: INDIA: Throughout India including Uttar Pradesh; Elsewhere: Cambodia, Laos, Sri Lanka, Thailand and Vietnam. Genus 43. Harpaglossus Motchulsky, 1858 1858. Harpaloglossus Motchulsky, Etud. Ent.: 173. 88. Harpaglossus opacus Chaudoir 1857. Ceroglossus opacus Chaudoir, Bull. Soc. Imp. Nat. Mosc., 2: 8. 1921. Systolocranius linea, Andrewes, Trans. Ent. Soc. Lond.: 167. Distribution: INDIA: Bihar, Madhya Pradesh, Maharashtra, Uttar Pradesh and West Bengal. Tribe 15. Licinini Bonelli, 1810 Genus 47. Badister Schellenberg, 1806* 1806. Badister Schellenberg, Helv. Ent., 2: 90 and 91. Badister sp. Material examined: 1 ex., Auraiya, near Bus station, 18.x.2011, Coll. B. Lal & party, Ex. At light; 414 Fauna of Uttar Pradesh, State Fauna Series, 22 1 ex., Jalaun, Chili near Orai, 15.x.2011, Coll. B. Lal & party, Ex. At light; 1 ex., Hardoi, Hardoi I. H. Bunglow, 10.x.2006, Coll. P. Mukhopadhyay & party, Ex. At light; 2 exs., Lucknow, Alambag, 12.vii.1989, Coll. K. K. Ray & party; 10 exs., Sultanpur, Panchopiran, 23.iii.2012, Coll. A. Sar & party, Ex. Under bricks. Genus 48. Diplocheila Brulle, 1834. Diplocheila Brulle, Hist. Nat des ins., 4: 407. 92. Diplocheila polita (Fabricius) 1792. Carabus polita Fabricius, Ent. Syst., 1(1): 146. 1922. Diplocheila polita, Andrewes, Ann. Mag nat. Hist. (9) 9: 283. Material examined: 1 ex., Hardoi, Neer, Bhais ka nala, 20.ii.2011, Coll. V. D. Hegde & party, Ex. In soil crevices (Reg. No. 20090/H4A). Distribution: INDIA: Throughout India including Uttar Pradesh; Elsewhere: Myanmar and Sri Lanka. 93. Diplocheila cordicollis Laferte* Subfamily 10. CARABINAE Latreille, 1802 Tribe 16. Carabini Latreille, 1802 Genus 49. Carabus Linnaeus, 1758 1758. Carabus Linnaeus, Syst. Nat.Ed. 10: 413. 95. Carabus barysomus Bates 1889. Carabus barysomus Bates, Proc. Zool. Soc. Lond.: 210. Distribution:INDIA : Jammu-Kashmir and Uttar Pradesh; Elsewhere: Pakistan. Subfamily 11. CICINDELINAE Latreille, 1802 Tribe 17. Cicindelini Latreille, 1802 Subtribe Cicindelina Latreille, 1802 Genus 50. Cicindela Linnaeus, 1758 1758. Cicindela Linnaeus, Syst. Nat., 1: 407. 96. Cicindela (Calochroa) octonotata Wiedemann 1819. Cicindela octonotata Wiedemann, Zool. Mag., i, 3: 168. 1989. Cicindela (Calochroa) octonotata, Acciavatti and Pearson, Ann. Carn. Mus., 58: 116. 1851. Diplocheila cordicollis Laferte, Ann. Soc. Ent. Fr.: 279. 1959. Diplocheila (Isorembus) cordicollis, Ball, Mem. Amer. Ent. Soc., (16): 54. Material examined: 1 ex., Jhansi, Babina, 21.xi.2010, Coll. O. P. Srivastava & party (Reg. no. 20049/H4A). Distribution: INDIA: Bihar, Delhi, Maharashtra, Uttar Pradesh and West Bengal; Elsewhere: Bangladesh. 94. Diplocheila (s. str.) daldorfi (Crotch)* 1871. Rembus daldorfi Crotch, List of all Coleoptera Described A.D. 1758-1821, Referred to other Modern Genera: 9. 1959. Diplocheila (s.str.) daldorfi, Ball, Mem. Amer. Ent. Soc., (16): 25. Material examined: 1 ex., Varanasi, Dashashwamedh Ghat, 28.ix.2010, Coll. V. D. Hegde & party, Ex. Under light (Reg. No. 20089/ H4A). Distribution: INDIA: Assam, Bihar, Tamil Nadu, Uttar Pradesh and West Bengal; Elsewhere: Myanmar, Singapore and Vietnam. Distribution: INDIA: Arunachal Pradesh, Assam, Bihar, Himachal Pradesh, Manipur, Meghalaya, Nagaland, Sikkim, Uttar Pradesh and West Bengal; Elsewhere: Bangladesh, Myanmar and Nepal. 97. Cicindela (Calochroa) flavomaculata Hope 1831. Cicindela flavomaculata Hope, Zool. misc., 1: 21. 1989. Cicindela (Calochroa) flavomaculata, Acciavatti and Pearson, Ann. Carn. Mus., 58: 116. Material examined: 2 exs., Varanasi, Babatpur, 24.x.2006, Coll. A. Bal & party; 2 exs., Allahabad, Ramnagar, 22.x.2006, Coll. A. Bal & party; 4 exs. Shahjahanpur, Jalalabad, 15.x.2006, Coll. A. Bal & party; 10 exs., Ballia, Surhatal W.L.S., 12.ii.2009, Coll. V. D. Hegde & party; 4 exs., Hardoi, I. H. Dak Bunglow, 10.x.2006, Coll. P. Mukhopadhyay & party, Ex. at light. Distribution: INDIA: Assam, Haryana, Jharkhand, Karnataka, New Delhi, Madhya Pradesh, Punjab, Uttarakhand, Uttar Pradesh and West Bengal; Elsewhere: South East Asia and Philippines. KUSHWAHA and HEGDE : Insecta : Coleoptera : Carabidae 415 98. Cicindela (Chaetodera) albina Wiedemann 103. Cicindela (Spilodia) multiguttata Dejean 1819. Cicindela albina Wiedemann, Zool. Mag., i, 3: 169. 1825. Cicindela (Spilodia) multiguttata Dejean, Spec. Gen. Col., 1: 109. 1989. Cicindela (Chaetodera) albina, Acciavatti and Pearson, Ann. Carn. Mus. 58: 140. Distribution: INDIA: Bihar, Haryana, Odisha, Punjab, Uttar Pradesh and West Bengal; Elsewhere: Bangladesh and Pakistan. 99. Cicindela (Chaetodera) vigintiguttata Herbst 1806. Cicindela (Chaetodera) vigintiguttata Herbst, Nat. Syst. Ins. Kaf., 10: 174, pl. 171, fig. 9. 1989. Cicindela (Chaetodera) vigintiguttata, Acciavatti and Pearson, Ann. Carn. Mus., 58: 141. Material examined: 3 exs., Sultanpur, 15.x.2006, Coll. P. Mukhopadhyay & party, Ex. Gomati River Bank. Distribution: INDIA: Assam, Bihar, Haryana, Odisha, Punjab, Sikkim, Uttar Pradesh and West Bengal; Elsewhere: Bangladesh, Bhutan and Pakistan. 100. Cicindela (Lophyra) catena Fabricius 1775. Cicindela (Lophyra) catena Fabricius, Syst. Ent.: 226. 1989. Cicindela (Lophyra) catena, Acciavatti and Pearson, Ann. Carn. Mus., 58: 143-144. Distribution: INDIA: Andhra Pradesh, Jharkhand, Karnataka, Kerala, Madhya Pradesh, Puducherry, Sikkim, Tamil Nadu, Uttarakhand, Uttar Pradesh and West Bengal; Elsewhere: Sri Lanka. 101. Cicindela (Lophyra) cancellata intemperata Acciavatti & Pearson 1882. Cicindela cancellata viridula Mandl, Kol. Rund., 56: 65. 1989. Cicindela (Lophyra) cancellata intemperata Acciavatti & Pearson, Ann. Carn. Mus., 58: 147, 148. Distribution: INDIA : Assam, Gujarat, Haryana, Sikkim, Uttar Pradesh and West Bengal; Elsewhere: Nepal, Pakistan, Sri Lanka and Thailand. 102. Cicindela (Spilodia) vittigera Dejean 1825. Cicindela (Spilodia) vittigera Dejean, Spec. Gen. Col., 1: 107. 1989. Cicindela (Spilodia) vittigera, Acciavatti & Pearson, Ann. Carn. Mus., 58: 154. Distribution: INDIA: Bihar, Haryana, Manipur, Punjab, Uttar Pradesh and West Bengal; Elsewhere: Bangladesh, Nepal and Pakistan. 1989. Cicindela (Spilodia) multiguttata, Acciavatti & Pearson, Ann. Carn. Mus., 58: 155. Distribution: INDIA: Arunachal Pradesh, Assam, Haryana, Punjab, Uttar Pradesh and West Bengal; Elsewhere: Bangladesh, Bhutan and Nepal. 104. Cicindela (Lophyridia) angulata Fabricius 1798. Cicindela (Lophyridia) angulata Fabricius, Suppl. Ent. Syst.: 62. 1989. Cicindela (Lophyridia) angulata, Acciavatti & Pearson, Ann. Carn. Mus., 58: 157. Material examined: 1 ex.♂, 2 exs.♀, Gorakhpur, Raj Ghat, Rapti River Bed, 17.iii.2010, Coll. V. D. Hegde & party, Ex. by hand net; 1 ex., Sultanpur, Sultanpur, 15.x.2006, Coll. P. Mukhopadhyay & party, Ex. Gomati River Bank. Distribution: INDIA : Andhra Pradesh, Assam, Bihar, Goa, Gujarat, Haryana, Karnataka, Kerala, Manipur, Puducherry, Punjab, Sikkim, Tamil Nadu, Uttar Pradesh and West Bengal; Elsewhere: Bangladesh, China, Myanmar, Pakistan, Nepal and Vietnam. 105. Cicindela (Lophyridia) plumigera macrograptina Acciavatti & Pearson 1989. Cicindela (Lophyridia) plumigera macrograptina Acciavatti & Pearson, Ann. Carn. Mus., 58: 162. Distribution: INDIA: Assam, Haryana, Uttar Pradesh and West Bengal; Elsewhere: Bangladesh and Nepal. 106. Cicindela (Lophyridia) chloris Hope 1831. Cicindela (Lophyridia) chloris Hope, Zool. misc., 1: 21. 1989. Cicindela (Lophyridia) chloris, Acciavatti & Pearson, Ann. Carn. Mus., 58: 164. Distribution: INDIA: Arunachal Pradesh, Assam, Bihar, Haryana, Himachal Pradesh, JammuKashmir, Punjab, Sikkim, Uttar Pradesh and West Bengal; Elsewhere: Bhutan, Nepal and Pakistan. 107. Cicindela (Eugrapha) bigemina Klug 1834. Cicindela bigemina Klug, Jahrb. Ins.:30. 1989. Cicindela (Eugrapha) bigemina, Acciavatti & Pearson, Ann. Carn. Mus., 58: 287. 416 Fauna of Uttar Pradesh, State Fauna Series, 22 Distribution: INDIA: Bihar, Jammu-Kashmir, Karnataka, Madhya Pradesh, Punjab, Uttar Pradesh and West Bengal; Elsewhere: Nepal and Pakistan. 108. Cicindela (Eugrapha) brevis Horn Distribution: INDIA: Andhra Pradesh, Bihar, Haryana, Karnataka, Kerala, Odisha, Punjab, Tamil Nadu, Uttar Pradesh and West Bengal; Elsewhere: Myanmar and Pakistan. 1905. Cicindela bigemina brevis Horn, Deut. Ent. Zeit.: 34. 113. Cicindela (Eugrapha) venosa Kollar 1989. Cicindela (Eugrapha) brevis, Acciavatti & Pearson, Ann. Carn. Mus., 58: 288. Material examined: 1 ex., Saharanpur, Beharigarh, 20.vii.1971, Coll. J.C. Tripathi and Party. Distribution: INDIA : Bihar, Haryana, Himachal Pradesh, Punjab and Uttar Pradesh; Elsewhere: Pakistan. 109. Cicindela (Eugrapha) minuta Olivier 1790. Cicindela minuta Olivier, Coleopteres, 2: 31, pl. 2, fig. 13a, b. 1989. Cicindela (Eugrapha) minuta, Acciavatti & Pearson, Ann. Carn. Mus., 58: 291. Distribution: INDIA : Andhra Pradesh, Gujarat, Karnataka, Kerala, Puducherry, Tamil Nadu, Uttar Pradesh and West Bengal; Elsewhere: Bangladesh, Indonesia, Kampuchea, Nepal and Vietnam. 110. Cicindela (Eugrapha) cognata Wiedemann 1823. Cicindela cognata Wiedemann, Zool. Mag. (1) 2: 66. 1989. Cicindela (Eugrapha) cognata, Acciavatti & Pearson, Ann. Carn. Mus., 58: 292. Distribution: INDIA: Andhra Pradesh, Bihar, Haryana, Odisha, Punjab, Tamil Nadu, Uttar Pradesh and West Bengal; Elsewhere: Bangladesh and Nepal. 111. Cicindela (Eugrapha) erudita Wiedemann 1823. Cicindela erudita Wiedemann, Zool Mag.(1) 2: 68. 1989. Cicindela (Eugrapha) erudita, Acciavatti & Pearson, Ann. Carn. Mus., 58: 294. Distribution: INDIA: Assam, Bihar, Haryana, Himachal Pradesh, Jammu-Kashmir, Jharkhand, Madhya Pradesh, Punjab, Sikkim, Uttar Pradesh and West Bengal; Elsewhere: Nepal. 112. Cicindela (Eugrapha) agnata Fleutiaux 1890. Cicindela (Eugrapha) agnata Fleutiaux, Ann. Soc. Ent. Belg., 34: 168. 1989. Cicindela (Eugrapha) agnata, Acciavatti & Pearson, Ann. Carn. Mus., 58: 295. 1836. Cicindela venosa Kollar,Wien. Mus. Ann., 1: 331. 1989. Cicindela (Eugrapha) venosa, Acciavatti & Pearson, Ann. Carn. Mus., 58: 298. Distribution: INDIA: Assam, Gujarat, Haryana, New Delhi, Punjab, Uttar Pradesh and West Bengal; Elsewhere: Bangladesh, Nepal and Pakistan. 114. Cicindela (Eugrapha) grammophora Chaudoir 1852. Cicindela grammophora Chaudoir, Bull. Soc. Imp. Nat. Mosc., (1) 25: 7. 1989. Cicindela (Eugrapha) grammophora, Acciavatti & Pearson, Ann. Carn. Mus., 58: 300. Distribution: INDIA: Bihar, Haryana, New Delhi, Odisha, Punjab, Rajasthan, Uttar Pradesh and West Bengal; Elsewhere: Bangladesh, Nepal and Pakistan. 115. Cicindela (Eriodera) albopunctata Chaudoir* 1852. Cicindela albopunctata Chaudoir, Bull. Soc. Imp. Nat. Mosc., (1) 25: 10. 1989. Cicindela (Eriodera) albopunctata, Acciavatti & Pearson, Ann. Carn. Mus., 58: 304. Material examined: 2 ex., Shahjahanpur, Khannaut river bank, 01.iii.2011, Coll. V. D. Hegde & party, Ex. on river bed. Distribution: INDIA: Assam, Himachal Pradesh, Punjab, Sikkim, Uttarakhand, Uttar Pradesh and West Bengal; Elsewhere: Nepal and Bhutan. 116. Cicindela (Myriochile) melancholica Fabricius 1798. Cicindela melancholica Fabricius, Ent. Syst. Suppl.: 63. 1989. Cicindela (Myriochile) melancholica, Acciavatti & Pearson, Ann. Carn. Mus., 58: 308. Material examined: 1 ex., Basti, Basti, 19.vi.2002, Coll. M. Prasad and Party. Distribution: INDIA : Bihar, Haryana, Madhya KUSHWAHA and HEGDE : Insecta : Coleoptera : Carabidae Pradesh, Maharashtra, New Delhi, Punjab, Rajasthan, Uttar Pradesh and West Bengal; Elsewhere: Pakistan and other Palaearctic Regions. 117. Cicindela (Myriochile) undulata Dejean 1826. Cicindela undulata Dejean, Spec. Gen. Col., l: 96. 1989. Cicindela (Myriochile) undulata, Acciavatti & Pearson, Ann. Carn. Mus., 58: 309. Distribution: INDIA : Andhra Pradesh, Haryana, Jharkhand, Karnataka, Kerala, Madhya Pradesh, Odisha, Puducherry, Punjab, Tamil Nadu, Uttar Pradesh and West Bengal; Elsewhere: Bangladesh, China and Sri Lanka. 118. Cicindela (Myriochile) atelesta Chaudoir 417 121. Oxylobus porcatus Fabricius 1798. Scarites porcatus Fabricius, Suppl. Ent. Syst.: 43. Distribution: INDIA: Andhra Pradesh, Jharkhand, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Odisha, Punjab, Tamil Nadu, Uttar Pradesh and West Bengal; Elsewhere: Pakistan and Sri Lanka. Genus 54. Scarites Fabricius, 1775 1775. Scarites Fabricius, Syst. Ent.: 249. 122. Scarites dyschromus Chaudoir 1855. Scarites dyschromus Chaudoir, Bull. Soc. Imp. Nat. Mosc., 1: 78. 1854. Cicindela atelesta Chaudoir, Bull. Soc. Imp. Nat. Mosc., (1) 27: 115. Material examined: 1 ex., Sultanpur, Payagpur, 20.ix.2007, Coll. A. Bal & party. 1989. Cicindela (Myriochile) atelesta, Acciavatti & Pearson, Ann. Carn. Mus., 58: 311. Distribution: INDIA: North India including Uttar Pradesh. Distribution: INDIA: Bihar, Sikkim, Uttar Pradesh and West Bengal; Elsewhere: Nepal. 123. Scarites indus Olivier Tribe 18. Megacephalini Laporte, 1834 Subtribe Megacephalina Laporte, 1834 Genus 51. Megacephala Latreille, 1802 1802. Megacephala Latreille, Hist. Nat. Ins., 3: 79. 119. Megacephala euphratica var.armeniaca Castelnau 1834. Megacephala euphratica var.armeniaca Castelnau, Rev Ent. Silb., 2: 28. Distribution: INDIA : Uttar Pradesh; Elsewhere: Afganistan, Armenia and Iran. 1795. Scarites Indus Olivier, Ent.: 36 and 9, t. 1, f. 2. Material examined: 1 ex., Etawah, Saraiya Village, 22.x.2011, Coll. B. Lal & party. Distribution: INDIA: Throughout India including Uttar Pradesh; Elsewhere: Sri Lanka. 124. Scarites laticeps Andrewes* 1929. Scarites laticeps Andrewes, Faun. Brit. India, Coleoptera, Carabidae, 1: 283. Material examined: 1 ex., Mau, Social Forestry, 21.ix.2010, Coll. V. D. Hegde & party. Subfamily 12. SCARITINAE Bonelli, 1810 Distribution: INDIA: Madhya Pradesh, Tamil Nadu and Uttar Pradesh. Tribe 19. Scaritini Bonelli, 1810 125. Scarites picicornis Dejean Genus 52. Haplogaster Chaudoir, 1879 1879. Haplogaster Chaudoir, Ann. Soc. Ent. Belg., 22: 149. 120. Haplogaster ovata Chaudoir 1879. Haplogaster ovata Chaudoir, Ann. Soc. Ent. Belg., 22: 150. Distribution: INDIA : Meghalaya, Uttar Pradesh and West Bengal; Elsewhere: Nepal and Bangladesh. Genus 53. Oxylobus Chaudoir, 1855 1855. Oxylobus Chaudoir, Bull. Soc. Imp. Nat. Mosc., 1: 5. 1831. Scarites picicornis Dejean, Spec. Gen. Col., 5: 493. Distribution: INDIA: Bihar, Uttar Pradesh and West Bengal; Elsewhere: Angola, Bangladesh, Ethiopea, Mozambique, River Niger, Senegal, Sri Lanka, South Africa, Sudan and Tanzania. 126. Scarites planus Bonelli 1813. Scarites planus Bonelli, Mem. Acad. Sci. Turin., 20: 470. Material examined: 2 exs., Chitrakoot, Pahari, 26.xi.2010, Coll. O. P. Srivastava & party, Ex. 418 Fauna of Uttar Pradesh, State Fauna Series, 22 Crevices in wheat Field soil; 4 exs., Jalaun, Orai, Ajnari, 17.x.2011, Alt. 89m., Coll. B. Lal & party, Ex. Leaf Litter; 1ex., Hardoi, Sandi Bird Sanctuary, 21.ii.2011, Coll. V. D. Hegde & party, Ex. Under cow dung. Distribution: INDIA : Uttar Pradesh; Elsewhere: Afghanistan, Egypt, Ethiopia, Iraq, Mozambique, Saudi Arabia, Sudan and Syria. 127. Scarites punctum Wiedeman 1823. Scarites punctum Wiedeman, Zool. Mag., 2: 1,38. Material examined: 1 ex., Sultanpur, Sultanpur, 15.x.2006, Coll. P. Mukhopadhyay and Party, Ex. Bank of Gomati River. Distribution: INDIA: Assam, Bihar, Himachal Pradesh, Punjab, Uttarakhand, Uttar Pradesh and West Bengal; Elsewhere: Nepal 128. Scarites puncticollis Chaudoir 1855. Taeniolobus puncticollis Chaudoir, Bull. Soc. Imp. Nat. Mosc., 1: 47. 1929. Scarites puncticollis, Andrewes, Faun. Brit. India, Coleoptera, Carabidae, 1: 234, 287. Material examined: 1 ex., Hardoi, Sandi Bird Sanctuary, 21.ii.2011, Coll. V. D. Hegde & party, Ex. Cow dung; 1 ex., Hardoi, Neer, Bhais ka Nala, 20.ii.2011, Coll. V. D. Hegde & party, Ex. Wet Land; 1 ex., Faizabad, Guptar Ghat, 09.iii.2010, Coll. V. D. Hegde & party, Ex. Hand picking; 4 exs., Barabanki, Mohammadpur, 20.iii.2012, N 26° 56´45´´ E 81°10´25´´, Coll. A. Sar & Party; 1 ex., Lakhimpur Kheri, Salukapur, Dudhwa National Park, GPS N28° 25´5´´ E80° 43´ 9´´, 02.iii.2013, Coll. V. D. Hegde & party (Reg. no. 20051/H4A); 2 exs., Lakhimpur Kheri, Sathiyana, Dudhwa National Park, GPS N28° 32´6´´ E80° 32´30´´, 01.iii.2013, Coll. V. D. Hegde & party. Uttar Pradesh, Tamil Nadu and West Bengal; Elsewhere: Bangladesh, Myanmar and Sri Lanka. 130. Scarites semirugosus Chaudoir* 1855. Scarites semirugosus Chaudoir, Bull. Soc. Imp. Nat. Mosc., 1: 90. Material examined: 1 ex., Hardoi, Sandi Bird Sanctuary, 21.ii.2011, Coll. V. D. Hegde & party, Ex. Soil Crevices of Ploughed field. Distribution: INDIA: Karnataka, Maharashtra, Meghalaya, Tamil Nadu, Uttar Pradesh and West Bengal; Elsewhere: Myanmar, Malaysia, Philippines and Thailand. 131. Scarites striaticeps Chaudoir* 1880. Scarites striaticeps Chaudoir, Ann. Soc. Ent. Belg., 23: 52. Material examined: 1 ex., Lakhimpur Kheri, Dudhwa FRH, 24.ix.1974, Coll. P. K. Maiti and party, Ex. At light on 7.30 PM. Distribution: INDIA: Assam, Jharkhand and Uttar Pradesh; Elsewhere: Bangladesh and Myanmar. 132. Scarites subnitens Chaudoir 1855. Scarites subnitens Chaudoir, Bull. Soc. Imp. Nat. Mosc., 1: 87. Material examined: 1 ex., Saharanpur, Rajaji Sanctuary, Saharanpur Road, 11.vi.1964, Coll. T. D. Soota, Ex. Under Stone; 1 ex., Mirzapur, Chunar, 8.ii.2009, Coll. V. D. Hegde & party, Ex. Ganga River Bed. Distribution: INDIA : Bihar, Uttarakhand, Uttar Pradesh and West Bengal; Elsewhere: Myanmar. 133. Scarites sulcatus Olivier* 1795. Scarites sulcatus Olivier, Ent. iii, 36: 7. t. 1. f. 11. Distribution: INDIA : Bihar, Gujarat, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Odisha, Pondicherry, Tamil Nadu, Uttar Pradesh and West Bengal; Elsewhere: Bangladesh, Nepal and Sri Lanka. 129. Scarites selene Schmidt-Goebel 1846. Scarites selene Schmidt-Goebel, Faun. Col. Birm.: 94. Distribution: INDIA: Assam, Bihar, Meghalaya, Material examined: 1 ex., Lakhimpur Kheri, Dudhwa National Park, 27.ii.2011, Coll. V. D. Hegde & party, Ex. Under log; 1 ex., Shahjahanpur, Shahjahanpur, 27.x.2007, Coll. P. Mukhopadhyay & party, Ex. Horse dung. Distribution: INDIA: Arunachal Pradesh, Assam, Himachal Pradesh, Uttarakhand, Uttar Pradesh and West Bengal; Elsewhere: Cambodia, KUSHWAHA and HEGDE : Insecta : Coleoptera : Carabidae 419 China, Indonesia, Korea, Laos, Myanmar, Nepal, Pakistan and Vietnam. Distribution: INDIA: including Gujarat and Uttar Pradesh; Elsewhere: Myanmar, and Thailand. 134. Scarites semicircularis Mcleay** 138. Clivina striata Putzeys* 1825. Scarites semicircularis Mcleay, Ann. Jav.: 24. 1846. Clivina striata Putzeys, Mem. Liege., 2: 592. Material examined: 1 ex., Lakhimpur Kheri, Forest along Saheli river bank, 23.ix.1974, Alt. 250m, Coll. P. K. Maiti & party. Material examined: 1 ex., Ghazipur, Social Forest Nursery, 24.ix.2010, Coll. V. D. Hegde & party, Ex. Under leaf litter; 2 exs., Allahabad, Meja, Near FRH, 21.viii.2007, Coll. V. D. Hegde & party, Ex. Under Leaf Litter. Distribution: INDIA : Uttar Pradesh; Elsewhere: Cambodia, Indonesia, Laos, Malaysia, Myanmar, Singapore, Thailand and Vietnam. Tribe 20. Clivinini Rafinasque, 1815 Genus 55. Clivina Latreille, 1802 1802. Clivina Latreille, Hist. nat. Crust. et Ins., 3: 96. 135. Clivina attenuata Herbst 1806. Scarites attenuata Herbst, Nat. Syst. Ins. Kaf., 10: 264, t. 176, f.7. 1929. Clivina attenuata Andrewes, Faun. Brit. India, Coleoptera, Carabidae, 1: 353, 355, f 51. Material examined: 12 exs., Mau, Sheetla Mandir Campus, 22.ix.2010, Coll. V. D. Hegde & party, Ex. Under bricks (Reg. No. 19550-19561/ H4A); 2 exs., Mau Social Forestry, Vandevi, 21.ix.2010, Coll. V. D. Hegde & party, Ex. Under bricks. Distribution: INDIA : Throughout Northern and Central India including Uttar Pradesh; Elsewhere: Cambodia, Indonesia, Iran, Laos, Myanmar and Vietnam. 136. Clivina gamma Andrewes* 1929. Clivina gamma Andrewes, Faun. Brit. Ind., Coleoptera, Carabidae, 1: 368-369. Material examined: 3 exs., Siddharth nagar, 13.iii.2010, Coll. V. D. Hegde & party, Ex. Agricultural Field. Distribution: INDIA : including Uttar Pradesh. 137. Clivina lobata Bonelli Distribution: INDIA: Bihar, Jharkhand, Southern India and Uttar Pradesh; Elsewhere: Sri Lanka. 139. Clivina tranquebarica Bonelli 1813. Clivina tranquebarica Bonelli, Mem. Acad. Sci. Turin., 20: 484. Material examined: 1 ex., Hardoi, Neer, Bhains ka nala, 20.ii.2011, Coll. V. D. Hegde & party, Ex. Wet Land; 2 exs., Sitapur, Tendua Jheel, 24.ii.2011, Coll. V. D. Hegde & party, Ex. Wet Land; 5exs., Lalitpur, Govind Sagar, 13.x.2011, Alt. 306m., Coll. Coll. B. Lal & party, Ex. Under stone near water stream; 3 exs., Lalitpur, Jailwara, 14.x.2011, Alt. 296m., Coll. B. Lal & party, Ex. Near water stream; 1 ex., Mirzapur, Chunar, 8.ii.2009, Coll. V. D. Hegde & party, Ex. Ganga River Bed; 4 exs., Deoria, by the side of Jugantar Chetana High School, 30.vi.1989, Coll. H. C. Ghosh & party; 1 ex., Siddharth Nagar, Siddharth Nagar, 13.iii.2010, Coll. V. D. Hegde & party, Ex. Agricultural Field; 1 ex., Ballia, Ballia, 13.ii.2009, Coll. V. D. Hegde & party, Bank of Ganga River. Distribution: INDIA: including Uttar Pradesh; Elsewhere: Cambodia, Laos, Malaysia, Myanmar, Sri Lanka and Vietnam. Tribe 21. Dyschiriini Kolbe, 1880 Genus 56. Dyschirius Panzer, 1813 1813. Dyschirius Panzer, Index Ent.: 67. 1813. Clivina lobata Bonelli, Mem. Acad. Sci. Turin., 20: 481. 140. Dyschirius hingstoni Andrewes 1892. Clivina laeviceps Bates, Ann. Mus. Civ. Sto. Nat. Gen., 32: 277. 1929. Dyschirius hingstoni Andrewes, Faun. Brit. India, Coleoptera, Carabidae, 1: 392, 400. Material examined: 1 ex., Allahabad, Ramnagar, 22.x.1986, Coll. A. Bal & party. Material examined: 1 ex., Auraiya, Jamui, 19.x.2011, Coll. B. Lal & party, Ex. Dhan Paddy. 420 Fauna of Uttar Pradesh, State Fauna Series, 22 Distribution: INDIA : Bihar, Uttar Pradesh and West Bengal. Distribution: INDIA: Maharashtra and Uttar Pradesh; Elsewhere: Sri Lanka. 141. Dyschirius mitodes Andrewes 147. Dyschirius bengalensis Andrewes* 1929. Dyschirius mitodes Andrewes, Faun. Brit. India, Coleoptera, Carabidae, 1: 394, 409. 1929. Dyschirius bengalensis Andrewes, Faun. Brit. India, Coleoptera, Carabidae, 1: 397-398. Distribution: INDIA (Uttar Pradesh). 142. Dyschirius perfectus Andrewes* 1929. Dyschirius perfectus Andrewes, Faun. Brit. India, Coleoptera, Carabidae, 1: 392, 399. Material examined: 1 ex., Hardoi, Neer, Bhais Ka Nala, 20.ii.2011, Coll. V. D. Hegde & party, Ex. Wet Land; 1 ex., Jhansi, Ambabai, Alt. 189 m., 12.x.2011, Coll. B. Lal & party (Reg. no. 20046/ H4A). Distribution: INDIA : Maharashtra, Uttarakhand and Uttar Pradesh. 143. Dyschirius variabilis Andrewes 1929. Dyschirius variabilis Andrewes, Faun. Brit. India, Coleoptera, Carabidae, 1: 393, 405. Distribution: INDIA: Uttarakhand, Uttar Pradesh and West Bengal; Elsewhere: Bangladesg. 144. Dyschirius verticalis Putzeys 1878. Dyschirius verticalis Putzeys, Compt. Rend soc. Ent. Belg.: 172. Distribution: INDIA : Uttar Pradesh. 145. Dyschirius mahratta Andrewes* 1929. Dyschirius mahratta Andrewes, Faun. Brit. India, Coleoptera, Carabidae, 1: 396. Material examined: 1 ex., Lakhimpur Kheri, Salukapur, Dudhwa National Park, GPS N28° 25´ 5´´ E80°43´9´´, 02.iii.2013, Coll. V. D. Hegde & party (Reg. no. 20050/H4A). Distribution: INDIA: Maharashtra and Uttar Pradesh; Elsewhere: Sri Lanka. 146. Dyschirius mahratta var. paucipunctus Andrewes* 1929. Dyschirius mahratta var. paucipunctus Andrewes, Faun. Brit. India, Coleoptera, Carabidae, 1: 397. Material examined: 1 ex., Hardoi, Neer, Bhais Ka Nala, 20.ii.2011, Coll. V. D. Hegde & party, Ex. wet land (reg. no. 20047/H4A). Material examined: 1 ex., Hardoi, Neer, Bhais Ka Nala, 20.ii.2011, Coll. V. D. Hegde & party, Ex. wet land (reg. no. 20048/H4A). Distribution: INDIA: Uttar Pradesh and West Bengal. Subfamily 13. MELAENINAE Allaud, 1934 Tribe 22. Cymbionotini Andrewes, 1933 Genus 57. Cymbionotum Baudi, 1864 1864. Cymbionotum Baudi, Berl. Ent. Zeit.: 211. 148. Cymbionotum fascigerum (Chaudoir) 1852. Coscinia fasciger Chaudoir, Bull. Soc. Imp. Nat. Mosc., 1: 92. 1935. Cymbionotum fascigerum, Andrewes, Faun. Brit. India, Coleoptera, Carabidae, 2: 25. Distribution: INDIA: Uttar Pradesh. Cymbionotum Sp. Material examined: 1 ex. Lucknow, Kakor, 23.vi.1989, Coll. K. K. Ray & party. Tribe 23. Melaenini Dejean, 1831 Genera 58. Melaenus Dejean, 1831* 1831. Melaenus Dejean, Spec. Gen. Col., 5: 481. 149. Melaenus piger (Fabricius)* 1801. Brachinus piger Fabricius, Syst. Eleuth., 1: 219. 1847. Melaenus piger Schaum, Stett. Ent. Zeit.: 49. Material examined: 1 ex.,Ghazipur, Social Forest Nursery, 24.ix.2010, Coll. V. D. Hegde & party, Ex. Under leaf litter (ZSI Reg. No. 20045/ H4A) ; 1 ex., Jalaun, Orai, near Machhar Chauraha, 15.x.2011, Coll. B. Lal & party, Ex. at light; 1 ex., Etawah, Bakewar, 22.x.2011, Coll. B. Lal & party. Distribution: INDIA: Andhra Pradesh, Chhattisgarh, Goa, Karnataka, Madhya Pradesh, Maharashtra, North India, Odisha, Puducherry, Tamil Nadu, Uttar Pradesh and West Bengal; Elsewhere: Sri Lanka. KUSHWAHA and HEGDE : Insecta : Coleoptera : Carabidae Subfamily 14. DRYPTINAE Bonelli, 1810 Tribe 24. Dryptini Bonelli, 1810 Genus 59. Drypta Latreille, 1796 1796. Drypta Latreille, Prec. Cara. Gen. Insectes: 75. 150. Drypta argillacea Andrewes 1924. Drypta argillacea Andrewes, MissionProv. Centr. l’Inde Reg. Occi. l’Himalaya 1914, par guy Babault: 106, t.3, f.5. Distribution: INDIA: Maharashtra, Odisha, Uttarakhand and Uttar Pradesh; Elsewhere: Bangladesh. 151. Drypta lineola Macleay 1825. Drypta lineola Macleay, Ann. Jav.: 27. Material examined: INDIA: Uttar Pradesh, 1 ex., Auraiya, near Dibiyapur , 19.x.2011, Coll. B. Lal & Party; 1 ex., Hardoi, Sandi Bird Sanctuary, 21.ii.2011, Coll. V. D. Hegde & Party, Ex. Under Cow dung; 1 ex., Lakhimpur Kheri, Dudhwa Tiger Reserve, 26.ii.2011, Coll. V.D. Hegde & party, Ex. Dusting. Distribution: INDIA: Himachal Pradesh, Maharashtra, and Uttar Pradesh; Elsewhere: throughout South East Asia. 421 Khannaut River Bank, 01.iii.2011, Coll. V. D. Hegde & party, Ex. Dusting; 2 exs., Shahjahanpur, Khannaut River Bank, 01.iii.2011, Coll. V. D. Hegde & party, Ex. In soil crevices; 1 ex., Sitapur, Tendua Jheel, 24.ii.2011, Coll. V. D. Hegde & party, Ex. in leaf node of sugarcane; 1 ex., Lakhimpur Kheri, Dudhwa National Park, Belrayan, GPS N28°23´14´´ E80°54´40.6´´, 28.ii.2013, Coll. V. D. Hegde & party; 1 ex., Lakhimpur Kheri, Dudhwa National Park, Sathiyana, GPS N28°32´6´´ E80°32´30´´, 01.iii.2013, Coll. V. D. Hegde & party. Distribution: INDIA: Bihar, Uttarakhand and Uttar Pradesh. Subfamily 16. SIAGONINAE Bonelli, 1813 Tribe 27. Siagonini Bonelli, 1813 Genus 62. Luperca Castelnau, 1840 1840. Luperca Castelnau, Hist. Nat. Ins. Col., 1: 63. 154. Luperca laevigata (Fabricius) 1781. Carabus laevigata Fabricius, Spec. Ins., 1: 304. 1929. Luperca laevigata, Andrewes, Faun. Brit. India,Coleoptera, Carabidae, 1: 206, f.35. Tribe 25. Galeritini Leconte, 1853 Distribution: INDIA: Bihar, Karnataka, Madhya Pradesh, Maharashtra, Odisha, Tamil Nadu and Uttar Pradesh. Genus 60. Galerita Fabricius, 1801 Genus 63. Siagona Latreille, 1804 1801. Galerita Fabricius, Syst. Eleuth., 1: 214. 152. Galerita indica Chaudoir 1861. Galerita indica Chaudoir, Bull. Soc. Imp. Nat. Mosc., 1: 557. Distribution: North India. Subfamily 15. CTENODACTYLINAE Castelnau, 1834 Tribe 26. Hexagonini Horn, 1881 Genus 61. Hexagonia Kirby, 1825* 1825. Hexagonia Kirby, Trans. Linn. Soc. Lond. 14: 563. 153. Hexagonia uninotata Andrewes* 1935. Hexagonia uninotata Andrewes, Fauna Brit. India, Carabidae, 2, Harapalini 1: 14, Pl. 2, f. 3. Material examined: 2exs., Shahjahanpur, 1804. Siagona Latreille, Nouv. Dict. Hist. Nat. 24. Tab. Meth.: 141. 155. Siagona atrata Dejean* 1825. Siagona atrata Dejean, Spec. Gen. Col., 1: 360. Material examined: 1 exs., Hardoi, Sandi Bird Sanctuary, 21.ii.2011, Coll. V. D. Hegde & party, Ex. crevices in soil (Z.S.I. Reg. No. 19548/H4A); 1 exs., Hardoi, Sandi Bird Sanctuary, 21.ii.2011, Coll. V. D. Hegde & party, Ex. Soil crevices of Ploughed field; 1 ex., Lakhimpur Kheri, Dudhwa National park, Chandan Chauki, GPS N28°32´1´´ E80°46´13´´, 27.ii.2013, Coll. V. D. Hegde & party, Ex. under bark. Distribution: INDIA : Assam, Bihar, Jharkhand, Odisha, Uttar Pradesh and West Bengal; Elsewhere: Bangladesh. 422 Fauna of Uttar Pradesh, State Fauna Series, 22 156. Siagona baconi Chaudoir* 1876. Siagona baconi Chaudoir, Bull. Mosc., 1: 89. Material examined: 1 ex., Ballia, FRH, 15.ix.2010, Coll. V. D. Hegde & party, (Reg. no. 19549/H4A); 1 ex., Hardoi, Sandi Bird Sanctuary, 21.ii.2011, Coll. V. D. Hegde & party, Ex. Crevices in soil; 1 ex., Hardoi, Sandi Bird Sanctuary, 21.ii.2011, Coll. V. D. Hegde & party, Ex. under cow dung; 1 ex., Jalaun, Orai, Machhar Chauraha, 17.x.2011, Coll. B. Lal & party, Ex. at light; 1 ex., Lakhimpur Kheri, Dudhwa National park, Chandan Chauki, GPS N28°32´1´´ E80°46´13´´, 27.ii.2013, Coll. V. D. Hegde & party, Ex. under bark. Distribution: INDIA: Bihar, Jharkhand, Madhya Pradesh, Odisha, Uttar Pradesh and West Bengal; Elsewhere: Bangladesh and Myanmar. 157. Siagona fabricii Andrewes 1921. Siagona fabricii Andrewes, Trans. Ent. Soc. Lond.: 158. Material examined: 1 ex., Auraiya, near Bus station, 18.x.2011, Coll. B. Lal & party, Ex. at light; 4 exs., Jalaun, Orai, Machhar Chauraha, 17.x.2011, Coll. B. Lal & party, Ex. at light; 17 exs., Jalaun, Orai, near Machhar Chauraha, 17.x.2011, Coll. B. Lal & party, Ex. at light (Reg. No. 19562-19578/ H4A); 2 exs., Jalaun, Orai, Vohadpura, 16.x.2011, Alt. 82m., Coll. B. Lal & party; 2exs., Jalaun, Orai, Ajnari, 17.x.2011, Alt. 89m., Coll. B. Lal & party, Ex. Under leaf litter; 12 exs., Jalaun, Chili near Orai, 15.x.2011, Coll. B. Lal & party, Ex. at light; 1 ex., Allahabad, Meja, near FRH, 21.viii.2007, Coll. V. D. Hegde & party, Ex. Under leaf litter; 20 exs., Lalitpur, near Railway Station, 14.x.2011, Coll. B. Lal & party, Ex. at light; 10 exs., Chitrakoot, Pahari, 26.xi.2010, Coll. O. P. Srivastava & party, Ex. Crevices in wheat field soil; 01 ex.,Ghazipur, Social Forest Nursery, 24.ix.2010, Coll. V. D. Hegde & party, Ex. Under leaf litter; 4 exs., Ballia, Jamui, 17.ix.2010, Coll. V. D. Hegde & party; 1 ex., Ballia, Bharatpura, 16.ix.2010, Coll. V. D. Hegde & party, Ex. Under stone; 9 exs., Ballia, FRH, 15.ix.2010, Coll. V. D. Hegde & party; 1 ex., Lakhimpur Kheri, Dudhwa National Park, Gauri Phanta, GPS N28° 38´53´´ E80°34´12´´, 26.ii.2013, Coll. V. D. Hegde & party. Distribution: INDIA: throughout India including Uttar Pradesh; Elsewhere: Sri Lanka. 158. Siagona flesus (Fabricius) 1801. Galerita flesus Fabricius, Syst. Eleuth., 1: 216. 1929. Siagona flesus, Andrewes, Faun. Brit. India, Coleoptera, Carabidae 1: 180, 182, 196. f. 32 (13), f. 34. Material examined: 1ex., Allahabad, Meja, near FRH, 21.viii.2007, Coll. V. D. Hegde & party; 1 ex., Ballia, FRH, 15.ix.2010, Coll. V. D. Hegde & party; 1 ex., Hardoi, Sandi Bird Sanctuary, 21.ii.2011, Coll. V. D. Hegde & party, Ex. Crevices in soil; 1 ex., Jalaun, Chili near Orai, 15.x.2011, Coll. B. Lal & party, Ex. at light; 1 ex., Jalaun, Orai, Vohadpura, 16.x.2011, Alt. 82m., Coll. B. Lal & party; 1 ex., Lalitpur, near Railway Station, 14.x.2011, Coll. B. Lal & party, Ex. at light; 1 ex., Jalaun, Orai, near Machhar Chauraha, 15.x.2011, Coll. B. Lal & party, Ex. at light, (Reg. no. 19580/ H4A). Distribution: INDIA: throughout India including Uttar Pradesh; Elsewhere: Cambodia, Laos, Myanmar, Malaysia, Senegal, Tanzania, Thailand and Vietnam. 159. Siagona pumila Andrewes* 1921. Siagona pumilus Andrewes, Trans. Ent. Soc. Lond.: 164. 1929. Siagona pumila, Andrewes, Fauna Brit. India, Coleoptera Carabidae, 1: 180, 199, f. 32(14), Pl. VI Fig. 2. Material examined: 2 exs., Lalitpur, near Railway Station, 14.x.2011, Coll. B. Lal & party, Ex. at light; 1 ex., Deoria, Hanuman Temple, 26.vi.1989, Coll. H.C. Ghosh & party, Ex. Near pond; 1 ex., Ballia, Jamui, 17.ix.2010, Coll. V. D. Hegde & party, (Reg. no. 19547/H4A); 1 ex., Hardoi, Sandi Bird Sanctuary, 21.ii.2011, Coll. V. D. Hegde & party, Ex. Under Cow dung; 2 exs., Hardoi, Sandi Bird Sanctuary, 21.ii.2011, Coll. V. D. Hegde & party, Ex. Crevices in Soil; 1 ex., Auraiya, Bamuripur, 6.ix.2009, Coll. V. D. Hegde & party, Ex. Under dead leaves (Reg. no. 19579/ H4A); 1 ex., Lakhimpur Kheri, Dudhwa National Park, Gauri Phanta, GPS N28°38´53´´ E80° 34´ 12´´, 26.ii.2013, Coll. V. D. Hegde & party. KUSHWAHA and HEGDE : Insecta : Coleoptera : Carabidae 423 Distribution: INDIA: Bihar, Karnataka, Maharashtra, Tamil Nadu, Uttarakhand, Uttar Pradesh and West Bengal; Elsewhere: Bangladesh. Under dry Sugarcane leaves; 1 ex., Saharanpur, Ghogregi vill., 31.x.2009, Coll. O. P. Srivastava & party. 160. Siagona dilutipes var. aligera Andrewes* Distribution: INDIA: throughout northern and central India including Manipur, Nilgiri Hills and Uttar Pradesh; Elsewhere: China, Laos, Myanmar and Vietnam. 1929. Siagona dilutipes var. aligera Andrewes, Fauna Brit. India, Coleoptera Carabidae, 1: 203. Material examined: 1 ex., Jalaun, Chili near Orai, 15.x.2011, Coll. B. Lal & party, Ex. at light. Distribution: INDIA: Maharashtra and Uttar Pradesh. Subfamily 17. PLATYNINAE Bonelli, 1810 Subfamily 18. NEBRIINAE Laporte, 1834 Tribe 29. Nebriini Laporte, 1834 Genus 60. Nebria Latreille, 1802 1802. Nebria Latreille, Hist. Nat. Crust. et Ins.: 89. Tribe 28. Platynini Bonelli, 1810 164. Nebria xanthacra Chaudoir Genus 64. Megalonychus Chaudoir, 1843 1850. Nebria xanthacra Chaudoir, Bull. Soc. Imp. Nat. Mosc., 1: 423. 1843. Megalonychus Chaudoir, Bull. Soc. Imp. Nat. Mosc., 2: 418. 161. Megalonychus birmanicus Bates 1892. Megalonychus birmanicus Bates, Ann. Mus. Civ. Sto. Nat. Gen. 32: 369. Distribution: INDIA: Arunachal Pradesh, Assam, Bihar, Meghalaya, Nagaland, Uttarakhand, Uttar Pradesh and West Bengal; Elsewhere: Laos and Myanmar. Genus 65. Onycholabis Bates, 1873* 1873. Onycholabis Bates, Trans. Ent. Soc. Lond.: 329. 162. Onycholabis acutangulus Andrewes* 1923. Onycholabis acutangulus Andrewes, Ann. Mag. Nat. Hist., (9)12: 682. Material examined: 1 ex., Saharanpur, Rajaji Sanctuary, Saharanpur Road, 27.xi.1964, Coll. T. D. Soota and party, Ex. under stone at bank of stream. Distribution: INDIA: Arunachal Pradesh, Uttarakhand and Uttar Pradesh. Genus 66. Orthotrichus Peyron, 1856 1856. Orthotrichus Peyron, Ann. Soc. Ent. Fr., 4 (3): 717. 163. Orthotrichus indicus Bates 1891. Orthotrichus indicus Bates, Compt. Rend soc. Ent. Belg.: 334. Material examined: 01 ex., Meerut, Meerut, 07.xi.2007, Coll. P. Mukhopadhyay and Party, Ex. Material examined: 7exs., Saharanpur, Beharigarh, 17.vi.1971, Coll. J. C. Tripathi and party. Distribution: INDIA: Himachal Pradesh, Kashmir, Uttarakhand and Uttar Pradesh. 165. Nebria xanthacra var. nigricollis Andrewes* 1925. Nebria xanthacra var. nigricollis Andrewes, Ann. Mag. nat. Hist., (9)15: 312. Material examined: 8 exs., Saharanpur, Beharigarh, 17.vi.1971, Coll. J. C. Tripathi & party; 2 exs., Saharanpur, Beharigarh, 17.vi.1971, Coll. J. C. Tripathi & party, (Reg. no. 5364-5365/H4A). Distribution: INDIA: Himachal Pradesh, Uttarakhand and Uttar Pradesh. Tribe 30. Notiophilini Motchulsky, 1850 Genus 68. Notiophilus Dumeril, 1806 1806. Notiophilus Dumeril, Zool. Analyt.: 194. 166. Notiophilus orientalis Chaudoir 1850. Notiophilus orientalis Chaudoir, Bull. Soc. Imp. Nat. Mosc., 1: 428. Distribution: INDIA: Himachal Pradesh, Kashmir, Uttarakhand and Uttar Pradesh; Elsewhere: Pakistan. Subfamily 19. OMOPHRONINAE Bonelli, 1810 Tribe 31. Omophronini Bonelli, 1810 424 Fauna of Uttar Pradesh, State Fauna Series, 22 Genus 69. Omophron Latreille, 1802 1802. Omophron Latreille, Hist. Nat. Crust. et Ins., 3: 89. 167. Omophron maculosus Chaudoir 1850. Omophron maculosus Chaudoir, Bull. Soc. Imp. Nat. Mosc., 1: 424. Distribution: INDIA : Bihar and Uttar Pradesh; Elsewhere: Bangladesh. Subfamily 20. PAUSSINAE Latreille, 1807 Tribe 32. Paussini Latreille, 1807 Genus 70. Merismoderus Westwood, 1847 1847. Ceratoderus (Merismoderus) Westwood, Trans. Ent. Soc. Lond., 5: 23. 1854. Merismoderus Lacordaire, Gen Col., 2: 11. 168. Merismoderus bensoni Westwood 1847. Merismoderus bensoni Westwood, Trans. Ent. Soc. Lond., 5: 23. Distribution: INDIA : Uttar Pradesh. Genus 71. Platyrhopalus Westwood, 1838 1838. Platyrhopalus Westwood, Trans. Linn. Soc. Lond., 16: 654. 169. Platyrhopalus denticornis Donovan 1800. Paussus denticornis Donovan, Ins Ind., 1: 8, pl. 5, fig. 1. Material examined: 2 exs., Ballia, Bharatpur, 16.ix.2010, Coll. V. D. Hegde & party, Ex. Under bark. Distribution: INDIA: Assam, Bihar, Delhi, Madhya Pradesh, Maharashtra, Punjab, Tamil Nadu, Uttar Pradesh and West Bengal. 170. Platyrhopalus angustus Westwood 1839. Platyrhopalus angustus Westwood, Trans. Ent. Soc. Lond., 2: 92, pl 10, fig. 6. Distribution: INDIA: Andhra Pradesh, Assam, Bihar, Kashmir, Uttarakhand and Uttar Pradesh. 171. Platyrhopalus intermedius Benson 1846. Platyrhopalus intermedius Benson, Calcutta Journ. Nat. Hist., 6: 468. Distribution: INDIA: Uttar Pradesh. Genus 72. Paussus Linnaeus, 1775 1775. Paussus Linnaeus, Bigae Insect: 7. 172. Paussus hearseyanus Westwood 1842. Paussus hearseyanus westwood, Proc. Linn. Soc. Lond.: 133. Distribution: INDIA : Gujarat, Madhya Pradesh, Maharashtra and Uttar Pradesh. 173. Paussus soleatus Wasmann* 1894. Paussus soleatus Wasmann, Krit. Verz. Myrm. Termit Aith.: 216. Material examined: 1 ex., Auraiya, Fatehpur near Muradganj, 20.x.2011, Coll. B. Lal & party, Ex. Leaf litter. Distribution: INDIA: Maharashtra and Uttar Pradesh. 174. Paussus ploiophorus Benson 1846. Paussus ploiophorus Benson, Calcutta Journ. Nat. Hist., 6: 463. Distribution: INDIA: Uttar Pradesh. SUMMARY The present paper deals with 174 species of 72 genera under 32 tribes and 20 subfamilies of Family Carabidae. During the study 551examples has been identified under 95 species and 36 genera. A total of 48 species and 12 genera has been recorded for the first time from Uttar Pradesh, out of which during study 22 species had been reported for the first time from Uttar Pradesh by Kushwaha & Hegde (2010, 2012 and 2013). A total 05 species has been recorded for the first time from India out of which during the study 02 had been reported for the first time from India by Kushwaha & Hegde (2013). ACKNOWLEDGEMENTS We are thankful to the Director, Zoological Survey of India, Kolkata for the facilities. We are indebted to Dr. Kailash Chandra, Addl. Director and Head, Entomology Division ‘A’ for his valuable suggestions in preparing the manuscript. We thank PCCF and Wild life Warden of Uttar Pradesh state for permission to collect the specimens and all the forest officials for providing help during our survey. We thank all the staffs of Coleoptera section for their constant encouragement. KUSHWAHA and HEGDE : Insecta : Coleoptera : Carabidae 425 REFERENCES Andrewes H. E. 1919. 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