Zool. Surv. India
Fauna of Uttar Pradesh, State Fauna Series, 22 (Part-2) : 395-426, 2015
INSECTA : COLEOPTERA : CARABIDAE
R. K. KUSHWAHA and V. D. HEGDE*
* Zoological Survey of India, M-Block, New Alipore, Kolkata-700 053
E-Mail : ravi_kumar3566@yahoo.com; hegde67@yahoo.co.in
INTRODUCTION
Family Carabidae is one of the largest family
under order Coleoptera. Beetles of the Family
Carabidae are also known as Ground Beetles. They
are elongate, oblongate, oval, flat, cylindrical and
sometimes rounded. The size varies from 1.5 mm
to 45-50 mm. These beetles are carnivorous in both
larval and adult stages and very few are carnivore
in larval stage and herbivore in adult stage. Most
of them are nocturnal, hiding under any objects
during day time and very few are diurnal. They live
under stone and bricks,under bark, under logs, at
light, vicinity of water and sand bank of water
bodies etc.
These beetles can be recognized by hind coxae
fused to metasternum and not dilated; first
abdominal sternite completely divided; antennae
filiform, 11 segmented and generally segment 1-4
glabrous, rest pubescent; 5-5-5 tarsal arrangement.
Family Carabidae comprises 34,275 species in
the world (Lorenz, 2005), however nearly 2000
species are represented in India (Saha, 1995). These
beetles are cosmopolitan in distribution. Carabid
beetles were already reported from following
districts of Uttar Pradesh state, Allahabad, Bareilly,
Bijnor, Chandauli, Chitrakoot, Faizabad, Fatehpur,
Gorakhpur, Lakhimpur Kheri, Lucknow,
Saharanpur, Sitapur, Sultanpur and Lalitpur.
STUDY AREA
Uttar Pradesh is the fourth largest state of India
with 75 districts. It is surrounded by Nepal,
Uttarakhand, Himachal Pradesh, Haryana and Delhi
in the north; by Rajasthan and Madhya Pradesh in
west; by Chhattisgarh in south; by Jharkhand and
Bihar in the East. Most of the land are dominated
by plain land. The state was surveyed by different
survey parties of Zoological Survey of India. During
study the specimens from Allahabad, Agra,
Auraiya, Azamgarh, Bahraich, Ballia, Barabanki,
Bareilly, Basti, Chandauli, Chitrakoot, Deoria,
Etawah, Fatehpur, Ghazipur, Gonda, Hardoi,
Jalaun, Jhansi, Lakhimpur Kheri, Lalitpur,
Lucknow, Mathura, Mau, Meerut, Mirzapur,
Pilibhit, Rae Bareilly, Saharanpur, Shahjahanpur,
Siddharth nagar, Sitapur, Sultanpur and Varanasi
districts were collected and identified.
MATERIAL AND METHODS
During the study period, different districts were
surveyed and specimens collected by different
teams of Zoological Survey of India. The specimens
collected and deposited in Coleoptera Section from
earlier surveys were also taken into consideration.
Specimens were collected by different method like
sweeping, dusting, hand picking and at light.
Preserved specimens were relaxed, set-pinned and
labelled. Specimens were sorted out in different
categories and identification was carried out with
the help of available literatures i.e. Andrewes (1919,
1920, 1922, 1929, 1935) and others. Taxa, first time
reported from State are marked with one asterisk
(*) and first time reported from India are marked
with two asterisk (**). Specimens were studied
under Lietz Stereo Binocular. The photographs of
specimens are taken with the help of Leica M 205A
stereo-zoom microscope, Nokia N73 and C5 phone
camera.
396
Fauna of Uttar Pradesh, State Fauna Series, 22
SYSTEMATIC LIST OF TAXA
Subfamily 2. HARPALINAE Bonelli, 1810
Subfamily 1. PTEROSTICHINAE Bonelli, 1810
Tribe 3. Harpalini Bonelli, 1810
Tribe 1. Abacetini Chaudoir, 1872
Subtribe. Stenolophina Kirby, 1837
Genus 1. Abacetus Dejean, 1828
Genus 6. Acupalpus Latreille, 1829.
1. Abacetus submetallicus (Nietner)
9. Acupalpus horni Andrewes
2. Abacetus vitreus Andrewes *
Genus 7. Pachytrachelus Chaudoir, 1852
3. Abacetus artus Andrewes *
Pachytrachelus Sp.
4. Abacetus guttula Chaudoir *
Genus 2. Cosmodiscus Sloane, 1907
5. Cosmodiscus picturatus Andrewes
Tribe 2. Pterostichini Bonelli, 1810
Subtribe 1. Molopina Bonelli, 1810
Genus 3. Euryaptus Bates, 1892
6. Euryaptus assamensis Bates
Subtribe 2. Pterostichina s.str.
Genus 4. Pterostichus Bonelli, 1810
7. Pterostichus harmandi Tchitcherin
Genus 5. Trigonotoma Dejean, 1828
8. Trigonotoma indica Brulle
Genus 8. Stenolophus Latreille, 1829
10. Stenolophus bajaurae Andrewes
Subtribe. Amblystomina Fauvel, 1889
Genus 9. Amblystomus Erichson, 1837 *
11. Amblystomus indicus (Nietner)*
Genus 10. Anoplogenius Chaudoir, 1852
12. Anoplogenius patinalis Bates
Genus 11. Barysomus Dejean, 1829
13. Barysomus semivittatus (Fabricius)
Subtribe Harpalina Bonelli, 1810
Genus 12. Dioryche Macleay, 1825
14. Dioryche chinnada Andrewes
KUSHWAHA and HEGDE : Insecta : Coleoptera : Carabidae
Genus 13. Hypolithus Dejean, 1829
15. Hypolithus acutangulus Bates
Genus 14. Kareya Andrewes, 1919
16. Kareya sublaevis (Bates)
17. Kareya sublaevis var. major (Bates)
Genus 15. Lamprophonus Bates, 1889
18. Lamprophonus lucens Bates
Genus 16. Oxycentrus Chaudoir, 1854 *
Oxycentrus sp.
Genus 17. Platymetopus Dejean, 1829
19. Platymetopus pictus Andrewes
397
32. Tetragonoderus taeniatus (Wiedeman) *
33. Tetragonoderus rhombophorus SchmidtGoebel**
Genus 28. Mnuphorus Chaudoir, 1873
34. Mnuphorus discophorus Chaudoir
Subfamily 4. ANTHIINAE Bonelli, 1813
Tribe 7. Anthiini Bonelli, 1813
Genus 29. Anthia Weber, 1801
35. Anthia sexguttata (Fabricius)
Tribe 8. Helluonini Hope, 1838
Genus 30. Macrocheilus Hope, 1838
20. Platymetopus rugosus (Nietner)
36. Macrocheilus chaudoiri Andrewes*
21. Platymetopus flavilabris (Fabricius)*
37. Macrocheilus niger Andrewes
Subtribe Anisodactylina Lacordaire, 1854
Genus 18. Gnathaphanus Macleay, 1825
22. Gnathaphanus rusticus Andrewes
Gnathaphanus Sp.
38. Macrocheilus trimaculatus Olivier
Genus 31. Omphra Reiche, 1842
Omphra Sp.
Subfamily 5. APOTOMINAE Leconte, 1853
Subfamily 3 . LEBIINAE Bonelli, 1810
Tribe 9. Apotomini Leconte, 1853
Tribe 4. Lebiini Bonelli, 1810
Genus 32. Apotomus Illiger, 1807
Genus 19. Anchista Nietner, 1856
23. Anchista brunnea (Wiedeman)
Genus 20. Bothynoptera Schaum, 1863
24. Bothynoptera dorsigera Schaum
Genus 21. Catascopus Kirby, 1825
25. Catascopus facialis (Wiedeman)
Genus 22. Dolichoctis Schmidt-Goebel, 1846 *
39. Apotomus hirsutulus Bates
Subfamily 6. TRECHINAE Bonelli, 1810
Tribe 10. Bembidiini Stephens, 1827
Genus 33. Asaphidion Des Gozis, 1886 *
40. Asaphidion championi Andrewes*
41. Asaphidion indicum (Chaudoir)*
Genus 34. Bembidion Latreille, 1802
26. Dolichoctis striata Schmidt-Goebel *
42. Bembidion loricatum Andrewes
Genus 23. Miscelus Klug, 1834
43. Bembidion luniferum Andrewes
27. Miscelus javanus var. unicolor Putzeys
Genus 24. Lachnoderma Macleay, 1873 *
28. Lachnoderma biguttatum Bates *
Genus 25. Cymindoidea Laporte, 1833 *
29. Cymindoidea distigma Chaudoir *
Tribe 5. Odacanthini Laporte, 1834
Genus 26. Odacantha Paykull, 1798
30. Odacantha bimaculata (Redtenbacher)
Tribe 6. Cyclosomini Laporte, 1834
Genus 27. Tetragonoderus Dejean, 1829 *
31. Tetragonoderus arcuatus Dejean *
44. Bembidion niloticum (Dejean)*
45. Bembidion notatum (Andrewes)*
46. Bembidion sobrinum Boheman
47. Bembidion xanthacrum Chaudoir
48. Bembidion xanthochiton Andrewes *
Genus 35. Tachys Stephens, 1828
49. Tachys peryphinus Bates**
50. Tachys babaulti Andrewes *
51. Tachys blandus Andrewes*
52. Tachys dulcis Andrewes
53. Tachys fumigatus Motchulsky
398
Fauna of Uttar Pradesh, State Fauna Series, 22
54. Tachys klugi (Nietner)
76. Chlaenius agilis Chaudoir
55. Tachys latus Peyron
77. Chlaenius bimaculatus lynx Chaudoir
56. Tachys latus var. virgatus Andrewes*
78. Chlaenius bioculatus Chaudoir
57. Tachys opalescens Andrewes**
79. Chlaenius chlorodius Dejean
58. Tachys ornatus Apetz
80. Chlaenius duvauceli (Dejean)
59. Tachys poecilopterus Bates
81. Chlaenius laetiusculus Chaudoir
60. Tachys politus Motchulsky
82. Chlaenius luculentus Andrewes
61. Tachys politus var. aspilotus Andrewes
83. Chlaenius pretiosus Chaudoir
62. Tachys ceylanicus (Nietner)*
84. Chlaenius scapularis Chaudoir*
63. Tachys latissimus var. gracilis (Motchulsky)*
85. Chlaenius trinototus Chaudoir*
Subfamily 7. BROSCINAE Hope, 1838
Tribe 11. Broscini Hope, 1838
Genus 36. Axonya Andrewes, 1923
64. Axonya championi Andrewes
Genus 37. Broscus Panzer, 1813
65. Broscus punctatus (Dejean)
Subfamily 8. BRACHININAE Bonelli, 1810
Tribe 12. Brachinini Bonelli, 1810
Subtribe Mastacina Erwin, 1970
Genus 38. Mastax Fischer von Waldheim, 1825 *
Mastax sp.
Subtribe Brachinina Bonelli, 1810
Genus 39. Brachinus Weber, 1801
66. Brachinus hexagrammus Chaudoir
67. Brachinus modestusSchmidt-Goebel *
68. Brachinus stevensi Andrewes*
69. Brachinus suturellus Chaudoir
Subtribe Pheropsophina Jeannel, 1948
Genus 40. Pheropsophus Solier, 1833
70. Pheropsophus assimilis Chaudoir **
86. Chlaenius variipes Chaudoir
87. Chlaenius xanthospilus (Wiedeman)
Genus 43. Harpaglossus Motchulsky, 1858
88. Harpaglossus opacus Chaudoir
Genus 44. Rhopalopalpus Laferte, 1851
89. Rhopalopalpus janthinus (Redtenbacher)
Tribe14. Oodini Laferte-Senectere, 1851
Genus 45. Oodes Dejean, 1826
90. Oodes subolivaceus Laferte
Genus 46. Systolocranius Chaudoir, 1857
91. Systolocranius linea Wiedeman
Tribe 15. Licinini Bonelli, 1810
Genus 47. Badister Schellenberg, 1806 *
Badister sp.
Genus 48. Diplocheila Brulle,
92. Diplocheila polita (Fabricius)
93. Diplocheila cordicollis Laferte *
94. Diplocheila (s. str.) daldorfi (Crotch)*
Subfamily 10. CARABINAE Latreille, 1802
71. Pheropsophus catoirei (Dejean)
Tribe 16. Carabini Latreille, 1802
72. Pheropsophus consularis Schmidt-Goebel*
Genus 49. Carabus Linnaeus, 1758
73. Pheropsophus hilaris (Fabricius)
74. Pheropsophus lineifrons Chaudoir
95. Carabus barysomus Bates
Subfamily 11. CICINDELINAE Latreille, 1802
Subfamily 9. LICININAE Bonelli, 1810
Tribe 17. Cicindelini Latreille, 1802
Tribe 13. Chlaeniini Brulle, 1834
Subtribe Cicindelina Latreille, 1802
Genus 41. Callistomimus Chaudoir, 1872
Genus 50. Cicindela Linnaeus, 1758
75. Callistomimus d’abreui Andrewes
Genus 42. Chlaenius Dejean, 1826
96. Cicindela (Calochroa) octonotata Wiedemann
97. Cicindela (Calochroa) flavomaculata Hope
KUSHWAHA and HEGDE : Insecta : Coleoptera : Carabidae
399
98. Cicindela (Chaetodera) albina Wiedemann
124. Scarites laticeps Andrewes *
99. Cicindela (Chaetodera) vigintiguttata Herbst
125. Scarites picicornis Dejean
100. Cicindela (Lophyra) catena Fabricius
126. Scarites planus Bonelli
101. Cicindela (Lophyra) cancellata intemperata
Acciavatti & Pearson
127. Scarites punctum Wiedeman
102. Cicindela (Spilodia) vittigera Dejean
129. Scarites selene Schmidt-Goebel
103. Cicindela (Spilodia) multiguttata Dejean
130. Scarites semirugosus Chaudoir *
104. Cicindela (Lophyridia) angulata Fabricius
131. Scarites striaticeps Chaudoir*
128. Scarites puncticollis Chaudoir
132. Scarites subnitens Chaudoir
105. Cicindela (Lophyridia) plumigera
macrograptina Acciavatti & Pearson
133. Scarites sulcatus Olivier*
106. Cicindela (Lophyridia) chloris Hope
134. Scarites semicircularis Mcleay **
107. Cicindela (Eugrapha) bigemina Klug
108. Cicindela (Eugrapha) brevis Horn
109. Cicindela (Eugrapha) minuta Olivier
110. Cicindela (Eugrapha) cognata Wiedemann
111. Cicindela (Eugrapha) erudita Wiedemann
112. Cicindela (Eugrapha) agnata Fleutiaux
Tribe 20. Clivinini Rafinasque, 1815
Genus 55. Clivina Latreille, 1802
135. Clivina attenuata Herbst (Scarites)
136. Clivina gamma Andrewes *
137. Clivina lobata Bonelli
138. Clivina striata Putzeys *
139. Clivina tranquebarica Bonelli
113. Cicindela (Eugrapha) venosa Kollar
Tribe 21. Dyschiriini Kolbe, 1880
114. Cicindela (Eugrapha) grammophora Chaudoir
Genus 56. Dyschirius Panzer, 1813
115. Cicindela (Eriodera) albopunctata Chaudoir*
140. Dyschirius hingstoni Andrewes
116. Cicindela (Myriochile) melancholica Fabricius
141. Dyschirius mitodes Andrewes
117. Cicindela (Myriochile) undulata Dejean
142. Dyschirius perfectus Andrewes *
118. Cicindela (Myriochile) atelesta Chaudoir
143. Dyschirius variabilisAndrewes
Tribe 18. Megacephalini Laporte, 1834
144. Dyschirius verticalis Putzeys
Subtribe Megacephalina Laporte, 1834
145. Dyschirius mahratta Andrewes *
Genus 51. Megacephala Latreille, 1802
146. Dyschirius mahratta var. paucipunctus
Andrewes*
119. Megacephala euphratica var. armeniaca
Castelnau
Subfamily 12. SCARITINAE Bonelli, 1810
Tribe 19. Scaritini Bonelli, 1810
Genus 52. Haplogaster Chaudoir, 1879
120. Haplogaster ovata Chaudoir
Genus 53. Oxylobus Chaudoir, 1855
121. Oxylobus porcatus Fabricius
Genus 54. Scarites Fabricius, 1775
122. Scarites dyschromus Chaudoir
123. Scarites indus Olivier
147. Dyschirius bengalensis Andrewes*
Subfamily 13. MELAENINAE Allaud, 1934
Tribe 22. Cymbionotini Andrewes, 1933
Genus 57. Cymbionotum Baudi, 1864
148. Cymbionotum fascigerum (Chaudoir)
Cymbionotum Sp.
Tribe 23. Melaenini Dejean, 1831
Genera 58. Melaenus Dejean, 1831 *
149. Melaenus piger (Fabricius) *
Subfamily 14. DRYPTINAE Bonelli, 1810
Tribe 24. Dryptini Bonelli, 1810
400
Fauna of Uttar Pradesh, State Fauna Series, 22
Genus 59. Drypta Latreille, 1796
Subfamily 19. OMOPHRONINAE Bonelli, 1810
150. Drypta argillacea Andrewes
Tribe 31. Omophronini Bonelli, 1810
151. Drypta lineola Macleay
Genus 69. Omophron Latreille, 1802
Tribe 25. Galeritini Leconte, 1853
Genus 60. Galerita Fabricius, 1801
152. Galerita indica Chaudoir
Subfamily 15. CTENODACTYLINAE
Castelnau, 1834
Tribe 26. Hexagonini Horn, 1881
Genus 61. Hexagonia Kirby, 1825 *
153. Hexagonia uninotata Andrewes *
Subfamily 16. SIAGONINAE Bonelli, 1813
Tribe 27. Siagonini Bonelli, 1813
Genus 62. Luperca Castelnau, 1840
154. Luperca laevigata Fabricius
Genus 63. Siagona Latreille, 1804
155. Siagona atrata Dejean*
156. Siagona baconi Chaudoir *
157. Siagona fabricii Andrewes
158. Siagona flesus (Fabricius)
159. Siagona pumila Andrewes *
160. Siagona dilutipes var. aligera Andrewes*
Subfamily 17. PLATYNINAE Bonelli, 1810
Tribe 28. Platyninni Bonelli, 1810
Genus 64. Megalonychus Chaudoir, 1843
161. Megalonychus birmanicus Bates
Genus 65. Onycholabis Bates, 1873 *
162. Onycholabis acutangulus Andrewes*
Genus 66. Orthotrichus Peyron, 1856
163. Orthotrichus indicus Bates
Subfamily 18. NEBRIINAE Laporte, 1834
Tribe 29. Nebriini Laporte, 1834
Genus 60. Nebria Latreille, 1802
164. Nebria xanthacra Chaudoir
165. Nebria xanthacra var. nigricollis Andrewes
*
Tribe 30. Notiophilini Motchulsky, 1850
Genus 68. Notiophilus Dumeril, 1806
166. Notiophilus orientalis Chaudoir
167. Omophron maculosus Chaudoir
Subfamily 20. PAUSSINAE Latreille, 1807
Tribe 32. Paussini Latreille, 1807
Genus 70. Merismoderus Westwood, 1847
168. Merismoderus bensoni Westwood
Genus 71. Platyrhopalus Westwood, 1838
169. Platyrhopalus denticornis Donovan
170. Platyrhopalus angustus Westwood
171. Platyrhopalus intermedius Benson
Genus 72. Paussus Linnaeus, 1775
172. Paussus hearseyanus Westwood
173. Paussus soleatus Wasmann*
174. Paussus ploiophorus Benson
Indications: * (First report from Uttar Pradesh)
** (First report from India)
SYSTEMATIC ACCOUNTS
Subfamily 1. PTEROSTICHINAE Bonelli, 1810
Tribe 1. Abacetini Chaudoir, 1872
Genus 1. Abacetus Dejean, 1828
1828. Abacetus Dejean, Spec. Gen. Col. 3: 195.
1. Abacetus submetallicus (Nietner)
1858. Distrigus submetallicus Nietner, Ann. Mag nat. Hist.,
(3) 2: 177.
1869. Abacetus chalceolus Chaudoir, Bull. Soc. Imp. Nat.
Mosc., 1: 384.
Material examined:1 ex., Allahabad, Fapamore,
04.ii.2009, Coll. V. D. Hegde & party (ZSI Reg.
no. 20030/H4A); 5 exs., Sitapur, Sinhanipur Jheel,
23.ii.2011, Coll. V. D. Hegde & party, Ex. Wet
Field (ZSI Reg. No. 20031-20035/H4A).
Distribution: INDIA: Throughout India
including Uttar Pradesh; Elsewhere: Cambodia,
Laos, Myanmar, Nepal, Sri Lanka and Vietnam.
2. Abacetus vitreus Andrewes*
1942. Abacetus vitreus Andrewes, Proc. Roy. Ent. Soc. Lond.,
11 (2): 32.
KUSHWAHA and HEGDE : Insecta : Coleoptera : Carabidae
Material examined: 1 ex., Sitapur, Sinhanipur
Jheel, 23.ii.2011, Coll. V. D. Hegde & party, Ex.
Wet Field (ZSI Reg. No. 20039/H4A).
401
Subtribe 2. Pterostichina s. str.
Genus 4. Pterostichus Bonelli, 1810
1810. Pterostichus Bonelli, Mem. Acad. Sci. Turin. 18: 68.
Distribution: INDIA: Manipur and Uttar
Pradesh.
7. Pterostichus harmandi Tchitcherin
3. Abacetus artus Andrewes*
1900. Pterostichus harmandi Tchitcherin, Horae Soc. Ent.
Ross., 34: 467.
1942. Abacetus artus Andrewes, Proc. Roy. Ent. Soc. Lond.,
11(2): 32.
Distribution: INDIA: Meghalaya, Uttar Pradesh
and West Bengal.
Material examined: 4 exs., Sitapur, Sinhanipur
Jheel, 23.ii.2011, Coll. V. D. Hegde & party, Ex.
Wet Field (ZSI Reg. No. 20040-20043/H4A).
Genus 5. Trigonotoma Dejean, 1828
Distribution: INDIA (Nevinson Coll., British
Museum) and Uttar Pradesh.
4. Abacetus guttula Chaudoir*
1869. Abacetus guttula Chaudoir, Bull. Soc. Imp. Nat. Mosc.,
1: 374.
Material examined: 1 ex., Chitrakoot, Pahari,
26.xi.2010, Coll. O. P. Srivastava and party, Ex.
Crevices in Wheat Field Soil (ZSI Reg. No. 20044/
H4A).
1828. Trigonotoma Dejean, Spec. Gen. Col., 3:182.
8. Trigonotoma indica Brulle
1834. Trigonotoma indica Brulle, Hist. Nat des Ins., 4: 333.
Distribution: INDIA: Assam, Delhi, Madhya
Pradesh, Maharashtra, Tamil Nadu, Uttarakhand,
Uttar Pradesh and West Bengal; Elsewhere:
Sri Lanka.
Subfamily 2. HARPALINAE Bonelli, 1810
Tribe 3. Harpalini Bonelli, 1810
Subtribe Stenolophina Kirby, 1837
Distribution: INDIA : Northern India including
Madhya Pradesh and Uttar Pradesh; Elsewhere: Sri
Lanka, Myanmar, and Iran.
1829. Acupalpus Latreille, in Cuviers’ Regne Animal
(Ed. 2) 4: 391.
Genus 2. Cosmodiscus Sloane, 1907
9. Acupalpus horni Andrewes
1907. Cosmodiscus Sloane, Proc. Linn. Soc. N. S. Wales,
32: 371.
5. Cosmodiscus picturatus Andrewes
1920. Cosmodiscus picturatus Andrewes, Ann. Mag nat.
Hist. (9)5: 447.
Genus 6. Acupalpus Latreille, 1829.
1923. Acupalpus horni Andrewes, Spol. Zeyl., 12: 237.
Distribution: INDIA: Karnataka, Tamil Nadu,
Uttarakh and and Uttar Pradesh; Elsewhere:
Myanmar and Sri Lanka.
Genus 7. Pachytrachelus Chaudoir, 1852
Distribution: INDIA : Andhra Pradesh, Odisha,
Maharashtra and Uttar Pradesh.
1852. Pachytrachelus Chaudoir, Bull. Soc. Imp. Nat. Mosc.,
1: 85.
Tribe 2. Pterostichini Bonelli, 1810
Pachytrachelus Sp.
Subtribe 1. Molopina Bonelli, 1810
Material examined: 1 ex. Jalaun, Orai, Machhar
Chauraha, 17.x.2011, Coll. B. Lal & Party, Ex. At
light.
Genus 3. Euryaptus Bates, 1892
1892. Euryaptus Bates, Ann. Mus. Civ. Sto. Nat. Gen., 32:
355.
Genus 8. Stenolophus Latreille, 1829
6. Euryaptus assamensis Bates
1829. Stenolophus Latreille, in Cuvier’s Regne Anim.
(Ed. ii) 4: 391.
1892. Euryaptus assamensis Bates, Ann. Mus. Civ. Sto. Nat.
Gen., 32: 356 (note).
10. Stenolophus bajaurae Andrewes
Distribution: INDIA: Assam, Uttarakhand and
Uttar Pradesh.
1924. Stenolophus bajaurae Andrewes, Mission Prov.
Centr. l’Inde Reg. Occi. l’Himalaya 1914, par Guy
Babault: 95.
402
Fauna of Uttar Pradesh, State Fauna Series, 22
Distribution: INDIA : Bihar, Himachal Pradesh,
Jammu-Kashmir, Uttarakhand and Uttar Pradesh.
Subtribe Amblystomina Fauvel, 1889
Genus 9. Amblystomus Erichson, 1837*
1837. Amblystomus Erichson, Kaf. Mark. Brand., 1: 59.
11. Amblystomus indicus (Nietner)*
1858. Megaristerus indicus Nietner, Ann. Mag. Nat. Hist.
(3) 2: 428.
1930. Amblystomus indicus, Andrewes, Cat. Ind. Insects, Part
18, Carabidae: 19.
Material examined: 1 ex., Jalaun, Chili near
Orai, 15.x.2011, Coll. B. Lal & party, Ex. at light;
1 ex., Jhansi, Pathrora near Barua sagar, 11.x.2011,
Alt. 165m., Coll. B. Lal & party, Ex. under leaf
litter.
Distribution: INDIA: Jharkhand and Uttar
Pradesh; Elsewhere: Australia, Myanmar, Sri Lanka
and Vietnam.
Genus 10. Anoplogenius Chaudoir, 1852
1852. Anoplogenius Chaudoir, Bull. Soc. Imp. Nat. Mosc.,
1: 88.
12. Anoplogenius patinalis Bates
1892. Anoplogenius patinalis Bates, Ann. Mus. Civ. Sto. Nat.
Gen., 32: 346.
Distribution: INDIA: Bihar, Uttarakhand and
Uttar Pradesh; Elsewhere: Myanmar.
Genus 11. Barysomus Dejean, 1829
1829. Barysomus Dejean, Spec. Gen. Col., 4: 56.
13. Barysomus semivittatus (Fabricius)
1798. Carabus semivittatus Fabricius, Suppl. Ent. Syst., 59.
1930. Barysomus semivittatus, Andrewes, Cat. Ind. Ins., Part
18. Carabidae: 38.
Distribution: INDIA: Bihar, Madhya Pradesh,
Odisha, Tamil Nadu and West Bengal; Elsewhere:
China, Sri Lanka and Vietnam.
Subtribe Harpalina Bonelli, 1810
Genus 12. Dioryche Macleay, 1825
1825. Dioryche Macleay, Ann. Jav., 21.
14. Dioryche chinnada Andrewes
1921. Dioryche chinnada Andrewes, Rec. Ind. Mus., 22: 342.
Distribution: INDIA: Bihar, Madhya Pradesh,
Maharashtra, Tamil Nadu, Uttar Pradesh and West
Bengal; Elsewhere: Sri Lanka.
Genus 13. Hypolithus Dejean, 1829
1829. Hypolithus Dejean, Spec. Gen. Col., 4: 166.
15. Hypolithus acutangulus Bates
1891. Hypolithus acutangulus Bates, Compt. Rend Soc. Ent.
Belg., 331.
Distribution: INDIA: Bihar, Delhi, Gujarat,
Jharkhand, Madhya Pradesh, Maharashtra, Tamil
Nadu, Uttarakhand and Uttar Pradesh; Elsewhere:
Bangladesh, Indonesia and Sri Lanka.
Genus 14. Kareya Andrewes, 1919
1919. Kareya Andrewes, Ann. Mag nat. Hist., (9) 3: 473.
16. Kareya sublaevis (Bates)
1891. Platymetopus sublaevis Bates, Compt. Rend soc. Ent.
Belg., 330.
1919. Kareya sublaevis Andrewes, Ann. Mag nat. Hist., (9)
3: 473.
Distribution: INDIA : Andhra Pradesh,
Arunachal Pradesh, Bihar, Jharkhand, Karnataka,
Madhya Pradesh, Maharashtra, Uttarakhand and
Uttar Pradesh; Elsewhere: Cambodia, China, Laos,
Myanmar, Thailand and Vietnam.
17. Kareya sublaevis var. major (Bates)
1891. Platymetopus sublaevis var. major Bates, Compt. Rend
Soc. Ent. Belg., 330.
1919. Kareya sublaevis var. major Andrewes, Ann. Mag nat.
Hist., (9) 3: 473.
Distribution: INDIA: Jharkhand, Madhya
Pradesh and Uttar Pradesh.
Genus 15. Lamprophonus Bates, 1889
1889. Lamprophonus Bates, Ann. Mus. Civ. Sto. Nat. Gen.,
27: 101.
18. Lamprophonus lucens Bates
1889. Lamprophonus lucens Bates, Ann. Mus. Civ. Sto. Nat.
Gen., 27: 102.
Distribution: INDIA: Andhra Pradesh, Bihar,
Jharkhand, Madhya Pradesh, Maharashtra, Odisha,
Tamil Nadu, Uttar Pradesh and West Bengal;
Elsewhere: Cambodia, Indonesia, Laos, Myanmar,
Sri Lanka and Vietnam.
KUSHWAHA and HEGDE : Insecta : Coleoptera : Carabidae
Genus 16. Oxycentrus Chaudoir, 1854*
403
1854. Oxycentrus Chaudoir, Bull. Soc. Imp. Nat. Mosc., 1:
345.
Distribution: INDIA: Bihar, Gujarat, Karnataka,
Madhya Pradesh, Maharashtra, Odisha,
Uttarakhand, Uttar Pradesh; Elsewhere: Sri Lanka.
Oxycentrus sp.
Gnathaphanus Sp.
Distribution: 5 exs., Hardoi, Hardoi, I. H.
Bunglow, 10.x.2006, Coll. P. Mukhopadhyay &
Party, Ex. At light.
Material examined: 1 ex., Allahabad, Meja, near
FRH, 21.viii.2007, Coll. V. D. Hegde & Party, Ex.
Under leaf litter; 1 ex., Mau, Social Forestry,
Vandevi, 21.ix.2010, Coll. V. D. Hegde & party,
Ex. under bricks.
Genus 17. Platymetopus Dejean, 1829
1829. Platymetopus Dejean, Spec. Gen. Col., 4: 68.
19. Platymetopus pictus Andrewes
1923. Platymetopus pictus Andrewes, Spol. Zeyl., 233.
Distribution: INDIA: Karnataka, Tamil Nadu
and Uttar Pradesh; Elsewhere: Saudi Arabia.
20. Platymetopus rugosus (Nietner)
Subfamily 3. LEBIINAE Bonelli, 1810
Tribe 4. Lebiini Bonelli, 1810
Genus 19. Anchista Nietner, 1856
1856. Anchista Nietner, Journ. Asia. Soc. Beng., 25: 523.
23. Anchista brunnea (Wiedeman)
1857. Ophonus rugosus Nietner, Journ. Asia. Soc. Beng.,
26: 150.
1823. Lebia brunnea Wiedeman, Zool. Mag., ii, 1: 59.
1927. Platymetopus rugosus, Andrewes, Ann. Mag nat. Hist.,
19: 106.
Distribution: INDIA: Bihar, Maharashtra and
Uttar Pradesh; Elsewhere: Bangladesh and Sri
Lanka.
Distribution: INDIA: Andhra Pradesh,
Jharkhand, Karnataka, Nilgiri Hills, Odisha, Tamil
Nadu, Uttarakhand, Uttar Pradesh and West Bengal;
Elsewhere: Sri Lanka and Victoria Point.
21. Platymetopus flavilabris (Fabricius)*
1798. Carabus flavilabris Fabricius, Suppl. Ent. Syst., 59.
1921. Platymetopus flavilabris, Andrewes, Rec. Ind. Mus.,
22: 344.
Material examined: 1 ex., Hardoi, Hardoi I. H.
Bunglow, 10.x.2006, Coll. P. Mukhopadhyay &
Party, Ex. At light.
Distribution: INDIA: including Uttar Pradesh;
Elsewhere: Sr Lanka.
Subtribe Anisodactylina Lacordaire, 1854
Genus 18. Gnathaphanus Macleay, 1825
1825. Gnathaphanus Macleay, Ann. Jav., 20.
22. Gnathaphanus rusticus Andrewes
1920. Gnathaphanus rusticus Andrewes, Ann. Soc. Ent.
Belg., 60: 107, 110.
Material examined: 1 ex., Ballia, Ballia,
11.xi.2009, Coll. V. D. Hegde & Party; 3 exs.,
Ballia, Surhatal WLS, 12.ii.2009, Coll. V. D. Hegde
& Party.
Genus 20. Bothynoptera Schaum, 1863
1863. Bothynoptera Schaum, Journ. Ent., 2: 75.
24. Bothynoptera dorsigera Schaum
1863. Bothynoptera dorsigera Schaum, Journ. Ent., 2: 76,
t.4, f.3.
Distribution: INDIA: Meghalaya, Uttar Pradesh
and West Bengal; Elsewhere: Vietnam.
Genus 21. Catascopus Kirby, 1825
1825. Catascopus Kirby, Trans. Linn. Soc. Lond., 14: 94.
25. Catascopus facialis (Wiedeman)
1819. Carabus facialis Wiedemen, Zool. Mag. i, 3: 165.
1930. Catascopus facialis, Andrewes, Cat. Ind. Insects Part
18, Carabidae: 76.
Distribution: INDIA: Assam, Throughout
Northern India including Uttar Pradesh; Elsewhere:
Myanmar.
Genus 22. Dolichoctis Schmidt-Goebel, 1846*
1846. Dolichoctis Schmidt-Goebel, Faun. Col. Birm., 62.
26. Dolichoctis striata Schmidt-Goebel*
1846. Dolichoctis striata Schmidt-Goebel, Faun. Col. Birm.,
62.
404
Fauna of Uttar Pradesh, State Fauna Series, 22
Material examined: 1 ex. Chitrakoot, 10.x.2001,
Coll. M. Prasad and party.
Distribution: INDIA: Andaman Island, Kerala
and Uttar Pradesh; Elsewhere: Australia, Indonesia,
Japan, Laos, Malaysia, Myanmar, Philippines, Sri
Lanka, Thailand and Vietnam.
Genus 23. Miscelus Klug, 1834
1834. Miscelus Klug, Jahrb. Ins., 82.
27. Miscelus javanus var. unicolor Putzeys
1845. Miscelus javanus var. unicolor Putzeys, Mem Soc.
Liege., 2: 375.
Distribution: INDIA: Bihar, Karnataka,
Kashmir, Madhya Pradesh and Uttar Pradesh;
Elsewhere: Australia,Cambodia, China, Laos,
Malaysia, Myanmar, Sri Lanka, Thailand and
Vietnam.
Genus 24. Lachnoderma Macleay, 1873*
1873. Lachnoderma Macleay, Trans. Ent. Soc. N. S. Wales,
2: 321
28. Lachnoderma biguttatum Bates*
1892. Lachnoderma biguttatum Bates, Ann. Mus. Civ. Sto.
Nat. Gen., 32: 424.
Material examined: 1 ex., Lakhimpur Kheri,
Dudhwa National Park, Salukapur, 02.iii.2013,
Coll. V. D. Hegde & Party.
Distribution: INDIA: Assam, Sikkim,
Uttarakhand and Uttar Pradesh; Elsewhere:
Myanmar.
Genus 25. Cymindoidea Laporte, 1832*
1832. Cymindoidea Laporte, Ann. Soc. Ent. Fr., 1: 390.
29. Cymindoidea distigma Chaudoir*
1875. Cymindoidea distigma Chaudoir, Bull. Soc. Imp. Nat.
Mosc., 2: 15.
Material examined: 1 ex. Jalaun, Orai,
Vohadpura, 16.x.2011, Alt. 82m, Coll. Coll. B. Lal
& party.
Distribution: INDIA: Uttarakhand, Uttar
Pradesh and West Bengal.
Tribe 5. Odacanthini Laporte, 1834
Genus 26. Odacantha Paykull, 1798
1798. Odacantha Paykull, Faun. Suec., 1: 169.
30. Odacantha bimaculata (Redtenbacher)
1844. Casnonia bimaculata Redtenbacher, Hugel’s
kaschmir, 4, 2: 498. t.23. f.2.
1930. Odacantha bimaculata Andrewes, Cat. Ind. Insects,
Part 18, Carabidae: 229.
Distribution: INDIA: Throughout Northern
India, extending Southwards to Karnataka and
Malabar; Elsewhere: Borneo, Cambodia, Indonesia,
Myanmar, Philippine Island and Sri Lanka.
Tribe 6. Cyclosomini Laporte, 1834
Genus 27. Tetragonoderus Dejean, 1829*
1829. Tetragonoderus Dejean, Spec. Gen. Col., 4: 485.
31. Tetragonoderus arcuatus Dejean*
1829. Tetragonoderus arcuatus Dejean, Spec. Gen. Col., 4:
495.
Material examined: INDIA: Uttar Pradesh,
4exs., Lucknow, Khurram Nagar, 16.ii.2011, Coll.
V.D. Hegde, Ex. Under leaf litter; 1 ex., Fatehpur,
Garhiwa, 16.iii.2012, GPS N 25° 54´ 55.3´´ E 80°
48´ 51.7´´, Coll. A. Sar & party; 2 exs., Barabanki,
Banki, Mohammadpur, 20.iii.2012, N26°56´ 45.1´´
E81°10´25.0´´, Coll. A. Sar & Party; 1 ex.,
Allahabad, Fapamore, 04.ii.2009, Coll. V. D. Hegde
& Party, Ex. Bank of Ganga River; 1 ex., Mau, Mau,
13.ii.2009, Coll. V. D. Hegde; 1 ex., Ballia, Surhatal
WLS, 12.ii.2009, Coll. V. D. Hegde & Party; 1 ex.,
Mathura, Kumaran Nagar, 16.xi.2010, Coll. O. P.
Srivastava & Party; 1 ex., Rae Bareilly, Bacthrawan,
09.vi.1989, Coll. A. K. Sanyal & Party; 1 ex.,
Azamgarh, Azamgarh, 10.ii.2009, Coll. V.D. Hegde
& Party; 1 ex., Siddharth Nagar, 13.iii.2010, Coll.
V. D. Hegde & Party, Ex. Agricultural Field; 1
ex.,Lucknow, Kuckrail Forest Area, 17.ii.2011,
Coll. V. D. Hegde& Party, Ex. Under Leaf Litter;
1 ex., Lucknow, Khurram Nagar, 16.ii.2011, Coll.
V. D. Hegde & party, Ex. under leaf litter (Reg. no.
20055/H4A); 1 ex., Lakhimpur Kheri, Dudhwa
National Park, Sathiyana, 01.iii.2013, Coll. V. D.
Hegde & Party.
Distribution: INDIA : Assam, Bihar, Himachal
Pradesh, Jharkhand, Madhya Pradesh, Maharashtra,
Meghalaya, Pondicherry, Uttarakhand, Uttar
Pradesh and West Bengal; Elsewhere: Bangladesh,
Egypt, Iraq, Laos, Myanmar and Sudan.
KUSHWAHA and HEGDE : Insecta : Coleoptera : Carabidae
32. Tetragonoderus taeniatus (Wiedeman)*
1823. Bembidium taeniatus Wiedeman, Zool. Mag. (2) 1: 62.
1850. Tetragonoderus taeniatus Chaudoir, Bull. Soc. Imp.
Nat. Mosc., 1: 455.
Material examined: 1 ex., Chandauli, 16.ii.2009,
Coll. V.D. Hegde & Party, ZSI Reg. No. 20056/
H4A; 1 ex., Lakhimpur Kheri, Dudhwa National
Park, Sathiyana, 27.ii.2011, Coll. V.D. Hegde &
Party.
Distribution: INDIA : Bihar, Himachal Pradesh
Uttarakhand, Uttar Pradesh and West Bengal;
Elsewhere: Myanmar.
33. Tetragonoderus rhombophorus
Schmidt-Goebel**
1846. Tetragonoderus rhombophorus Schmidt-Goebel,
Faun. Col. Birm., 93.
Material examined: 1 ex., Laknimpur Kheri,
Salukapur, Dudhwa National Park, GPS N28° 25´
5´´ E80° 43´9´´, 02.iii.2013, Coll. V. D. Hegde &
party (Reg. no. 20054/H4A).
Distribution: INDIA : Uttar Pradesh; Elsewhere:
Bangladesh, Laos and Myanmar.
Genus 28. Mnuphorus Chaudoir, 1873
1873. Mnuphorus Chaudoir, Berl. Ent. Zeit., 17: 55.
34. Mnuphorus discophorus Chaudoir
1876. Mnuphorus discophorus Chaudoir, Bull. Soc. Imp. Nat.
Mosc., 2: 69.
Distribution: INDIA (Bihar, Himachal Pradesh,
Uttarakhand and Uttar Pradesh).
Subfamily 4. ANTHIINAE Bonelli, 1813
Tribe 7. Anthiini Bonelli, 1813
Genus 29. Anthia Weber, 1801
1801. Anthia Weber, Obs. Ent.: 17.
35. Anthia sexguttata (Fabricius)
1775. Carabus sexguttata Fabricius, Syst. Ent., 236.
405
Distribution: INDIA : Throughout Indian Plains
including Uttar Pradesh.
Tribe 8. Helluonini Hope, 1838
Genus 30. Macrocheilus Hope, 1838
1838. Macrocheilus Hope, Col. Man., 2: 166.
36. Macrocheilus chaudoiri Andrewes*
1919. Macrocheilus chaudoiri Andrewes, Trans. Ent. Soc.,
Lond. 67: 130.
Material examined: 1 ex., Gonda, East of
Ambedkar Chauraha, Civil Lines, 12.viii.2007,
Coll. V. D. Hegde & Party, Ex. Under dry leaves
(Reg. no. 20064/H4A).
Distribution: INDIA: Bihar, Maharashtra, and
Uttar Pradesh; Elsewhere: Bangladesh, Myanmar,
Indonesia, Philippines, Laos, Cambodia, and China.
37. Macrocheilus niger Andrewes
1920. Macrocheilus niger Andrewes, Ann. Mag. Nat. Hist.
(9)6: 497, 499.
Distribution: INDIA: Andhra Pradesh, Bihar,
Jharkhand, Karnataka, Madhya Pradesh,
Maharashtra, Nilgiri Hills, Tamil Nadu andUttar
Pradesh) and Sri Lanka.
38. Macrocheilus trimaculatus Olivier
1790. Carabus trimaculatus Olivier, Enc. Meth., 347. t. 179.
f. 11.
1924. Macrocheilus trimaculatus, Andrewes, Ann. Mag nat.
Hist., (9) 13: 469.
Material examined: 1ex., Etah, Irrigation
Bunglow, 08.iii.2008, Coll. P. Mukhopadhyay &
party, Ex. at light.
Distribution: INDIA : including Uttar Pradesh;
Elsewhere: China, Laos, Myanmar, Malaysia,
Philippines, Sri Lanka and Vietnam.
Genus 31. Omphra Reiche, 1842
1842. Omphra Reiche, Ann. Soc. Ent. Fr., 330.
1930. Anthia sexguttata, Andrewes, Cat. Ind. Ins. Part 18
Carabidae: 31.
Omphra Sp.
Material examined: INDIA: 1ex., Sultanpur,
Panchopiran, 23.iii.2012, GPS N26°16´19.7´´ E82°
05´13.0´´, Coll. A. Sar and Party, Ex. Under dry
leaves.
Material examined: 1 ex., Auraiya, Shankarpur,
07.ix.2009, Coll. V. D. Hegde & Party; 1ex.,
Lalitpur, Jailwara, 14.x.2011, Alt. 296m, Coll. B.
Lal & Party, Ex. Under stone.
406
Fauna of Uttar Pradesh, State Fauna Series, 22
Subfamily 5. APOTOMINAE Leconte, 1853
42. Bembidion loricatum Andrewes
Tribe 9. Apotomini Leconte, 1853
1922. Bembidion loricatum Andrewes, Ent. Month. Mag.:
239.
Genus 32. Apotomus Illiger, 1807
1807. Apotomus Illiger, Mag. Ins., 6: 348.
2013. Bembidion loricatum, Kushwaha & Hegde, in
Souvenir: Water and Biodiversity: 158.
1892. Apotomus hirsutulus Bates, Ann. Mus. Civ. Sto. Nat.
Gen., 32: 287.
Material examined: 1 ex., Barabanki, Banki,
Mohammadpur, 20.iii.2012, N26°56´45.1´´ E
81°10´25´´, Coll. A. Sar and Party (Reg. no. 19581/
H4A).
Distribution: INDIA: Throughout India
including Uttar Pradesh; Elsewhere: Myanmar.
Distribution: INDIA: Bihar, Uttarakhand and
Uttar Pradesh; Elsewhere: Pakistan.
Subfamily 6. TRECHINAE Bonelli, 1810
43. Bembidion luniferum Andrewes
Tribe 10. Bembidiini Stephens, 1827
1924. Bembidion luniferum Andrewes, Ent. Month. Mag.:
192.
39. Apotomus hirsutulus Bates
Genus 33. Asaphidion Des Gozis, 1886*
1886. Asaphidion Des Gozis, Reche. espe. typi. dequelq.
ancien. genr., recti. syno., notes diver.: 6.
1831. Tachypus Dejean, Spec. Gen. Col., 5: 35.
Distribution: INDIA: Uttarakhand and Uttar
Pradesh.
44. Bembidion niloticum (Dejean)*
1831. Bembidium niloticum Dejean, Spec. Gen. Col., 5: 73.
40. Asaphidion championi Andrewes*
1924. Asaphidion championi Andrewes, in Mission Prov.
Centr. l‘Inde Reg. Occi. l’Himalaya, 1914, par Guy
Babault, Carab : 84. t.2.f.2.
Material examined: 1ex., Saharanpur,
Beharigarh, 17.vi.1971, Coll. J. C. Tripathi & Party.
Distribution: INDIA: Assam, Himachal Pradesh,
Uttarakhand, Uttar Pradesh and West Bengal.
41. Asaphidion indicum (Chaudoir)*
1850. Tachypus indicum Chaudoir, Bull. Soc. Imp. Nat.
Mosc., 2: 189 (Note).
1930. Asaphidion indicum, Andrewes, Cat. Ind. Insects, Part
18, Carabidae: 35.
Material examined: 1ex., Barabanki, Banki,
Mohammadpur, 20.iii.2012, N26°56´45.1´´ E81°
10´ 25.0´´, Coll. A. Sar & Party; 1 ex., Lakhimpur
Kheri, Dudhwa National Park, Sathiyana,
27.ii.2011, Coll. V. D. Hegde & party, Ex. Under
leaf litter.
Distribution: INDIA : Bihar, Haryana, Himachal
Pradesh, Uttarakhand, Uttar Pradesh and West
Bengal.
Genus 34. Bembidion Latreille, 1802
1802. Bembidion Latreille, Hist. Nat. Crust. et. Ins., 3: 82.
2013. Bembidion niloticum, Kushwaha & Hegde, in
Souvenir: Water and Biodiversity: 159.
Material examined: 1 ex., Barabanki, Sultanpur,
21.iii.2012, A. Sar and Party; 1 ex., Hardoi, Neer,
Bhais ka nala, 20.iii.2011, Coll. V. D. Hegde &
Party, Ex. Wet land; 1 ex., Siddharth Nagar,
13.iii.2010, Coll. V. D. Hegde & Party, Ex.
Agricultural Field.
Distribution:INDIA: Haryana, Nagaland,
Uttarakhand, Uttar Pradesh and West Bengal;
Elsewhere: Cambodia, China, Egypt, Iraq, Iran,
Japan, Laos, Pakistan, Saudi Arabia, Turkey and
Vietnam.
45. Bembidion notatum (Andrewes)*
1922. Bembidium notatum Andrewes, Ent. Month. Mag.:
175.
2013. Bembidion loricatum, Kushwaha & Hegde, in
Souvenir: Water and Biodiversity: 159.
Material examined: 1 ex., Hardoi, Neer, Bhais
ka Nala, 20.ii.2011, Coll. V. D. Hegde & party,
Ex. in soil crevices; 2 exs., Siddharth nagar,
Siddharth nagar, 13.iii.2010, Coll. V. D. Hegde &
Party, Ex. Agricultural Field; 1 ex., Shahjahanpur,
Khannaut River bank, 01.iii.2011, Coll. V. D.
Hegde & Party, Ex. On river bed; 2 exs., Hardoi,
Neer, Bhais ka Nala, 20.ii.2011, Coll. V. D. Hegde
KUSHWAHA and HEGDE : Insecta : Coleoptera : Carabidae
407
& party, Ex. Wet land; 2 exs., Hardoi, Neer, Bhais
Ka Nala, 20.ii.2011, Coll. V. D. Hegde & Party,
Ex. in soil crevices (Reg. No. 19607-19608/H4A);
3 exs., Hardoi, Neer, Bhais ka Nala, 20.ii.2011,
Coll. V. D. Hegde & party, Ex. Wet land (Reg. no.
19632-19634/H4A); 1 ex., Sitapur, Tendua Jheel,
24.ii.2011, Coll. V. D. Hegde & party (Reg. no.
19635/H4A).
Sitapur, Tendua Jheel, 24.ii.2011, Coll. V.D. Hegde
& party, Ex. wet land (Reg. no. 19618/H4A).
Distribution: INDIA: Himachal Pradesh,
Meghalaya, Uttarakhand, Uttar Pradesh and West
Bengal.
1850. Bembidion xanthacrum Chaudoir, Bull. Soc. Imp. Nat.
Mosc., 2: 175 (note).
46. Bembidion sobrinum Boheman
1848. Bembidium sobrinum Boheman, Ins. Caffr., 1: 232.
2013. Bembidion sobrinum, Kushwaha & Hegde, in
Souvenir: Water and Biodiversity: 159.
Material examined: 1 ex., Mathura, Kishori
Kunj, 17.xi.2010, Coll. O. P. Srivastava & Party; 5
exs., Mathura, Kishori Kunj, 17.xi.2010, Coll. O.
P. Srivastava & party, (Reg. No. 19613-19617/
H4A); 2 exs., Lucknow, Kuckrail Forest Area,
17.ii.2011, Coll. V. D. Hegde & Party, Ex. At pond
edge; 3exs., Lucknow, Kuckrail Forest Area,
17.ii.2011, Coll. V. D. Hegde & Party, Ex. At pond
edge (Reg. No. 19586, 19588, 19620/H4A); 1 ex.,
Fatehpur, Garhiwa, 16.iii.2012, GPS N25°54´
55.3°´´ E80°48´51.7´´, Coll. A. Sar & Party; 1 ex.,
Barabanki, Sultanpur, 21.iii.2012, Coll. A. Sar &
Party; 3 exs., Shahjahanpur, Khannaut River bank,
01.iii.2011, Coll. V. D. Hegde & Party, Ex. On river
bed; 2 exs., Shahjahanpur, Khannaut River bank,
01.iii.2011, Coll. V. D. Hegde & Party, Ex. On river
bed (Reg. no. 19621, 19605/H4A); 1 ex., Siddharth
Nagar, Siddharth Nagar, 13.iii.2010, Coll. V. D.
Hegde & Party, Ex. Agricultural Field (Reg. No.
19587/H4A); 1 ex., Ballia, 13.ii.2009, Coll. V. D.
Hegde & party (Reg. no. 19589/H4A); 17 exs.,
Hardoi, Neer, Bhais Ka Nala, 20.ii.2011, Coll. V.
D. Hegde & Party; 5 exs., Hardoi, Neer, Bhais Ka
Nala, 20.ii.2011, Coll. V. D. Hegde & Party, Ex. In
soil Crevices (Reg. No. 19601-19604, 19610/H4A);
20 exs., Hardoi, Neer, Bhais Ka Nala, 20.ii.2011,
Coll. V. D. Hegde & Party, Ex. wet land (1959319600, 19609,19611, 19619,19622-19630/H4A); 1
ex., Etah, Awagarh, 22.iii.1990, Coll. B. K.
Biswas& party (Reg. no. 19612/H4A); 1 ex.,
Distribution: INDIA : Plains of India including
Uttar Pradesh; Elsewhere: Cambodia, East African
Countries, Indonesia, Sri Lanka, Myanmar, South
Africa and Thiland.
47. Bembidion xanthacrum Chaudoir
2013. Bembidion xanthacrum, Kushwaha & Hegde, in
Souvenir: Water and Biodiversity: 159.
Material examined: 1ex., Chandauli, 17.iv.2007,
Coll. S.K. Chatterjee & party, Ex. Bank of Ganga
River (Reg. No. 19606/H4A); 1 ex., Shahjahanpur,
Khannaut river Bank, 01.iii.2011, Coll. V. D. Hegde
& Party, Ex. On river bed; 1 ex., Allahabad,
Fapamore, 04.ii.2009, Coll. V. D. Hegde & Party;
1 ex., Chandauli, Chandauli, 16.ii.2009, Coll. V.D.
Hegde & Party (Reg. no. 19582/H4A); 1 ex.,
Allahabad, Karela Bag, 05.ii.2009, Coll. V. D.
Hegde & Party, Ex. Yamuna River Bed (Reg. No.
19583/H4A); 1 ex., Sultanpur, Sultanpur,
15.x.2006, Coll. P. Mukhopadhyay & Party, Ex.
Bank of Gomati River (Reg. No. 19584/H4A); 1
ex., Ballia, Ballia, 13.ii.2009, Coll. V. D. Hegde &
Party, Ex. Bank of Ganga River (Reg. no. 19585/
H4A);1 ex., Shahjahanpur, Khannaut river Bank,
01.iii.2011, Coll. V. D. Hegde & Party, Ex. in soil
crevices (Reg. no. 19631/H4A); 1 ex., Lakhimpur
Kheri, Salukapur, Dudhwa National Park, GPS
N28°25´5´´ E80°43´9´´, 02.iii.2013, Coll. V. D.
Hegde & party (Reg. no. 20052/H4A).
Distribution: INDIA : Andhra Pradesh, Assam,
Bihar, Himachal Pradesh, Kerala, Madhya Pradesh,
Maharashtra, Uttarakhand, Uttar Pradesh, Tamil
Nadu and West Bengal; Elsewhere: Cambodia,
Laos, Myanmar, Pakistan, Thailand and Vietnam.
48. Bembidion xanthochiton Andrewes*
1922. Bembidium xanthochiton Andrewes, Ent. Month.
Mag.: 176.
2013. Bembidion xanthochiton, Kushwaha & Hegde, in
Souvenir: Water and Biodiversity: 159-160.
Material examined: 1 ex., Hardoi, Neer, Bhais
Ka Nala, 20.ii.2011, Coll. V. D. Hegde & party,
408
Fauna of Uttar Pradesh, State Fauna Series, 22
Ex. wet land (Reg. no. 19637/H4A); 1 ex.,
Lucknow, Kuckrail Forest Area, 17.ii.2011, Ex. at
pond edge (Reg. no. 19636/H4A).
Distribution: INDIA: Himachal Pradesh,
Uttarakhand, Uttar Pradesh and West Bengal.
Genus 35. Tachys Stephens, 1828
1828. Tachys Stephens, Ill. Brit. Ent. 2: 2, 4.
49. Tachys peryphinus Bates**
1886. Tachys peryphinus Bates, Ann. Mag. Nat. Hist., (5)17:
153.
Material examined: INDIA: Uttar Pradesh, 2
exs., Lalitpur, Jailwara, 14.x.2011, Alt. 296 m.,
Coll. B. Lal & Party, Ex. Near stream; 1 ex.,
Shahjahanpur, Khannaut river bank, 01.iii.2011,
Coll. V. D. Hegde & Party, Ex. In soil crevices.
Distribution: INDIA : Uttar Pradesh; Elsewhere:
Sri Lanka.
50. Tachys babaulti Andrewes*
1924. Tachys babaulti Andrewes, in Mission Prov. Centr.
l‘Inde Reg. Occi. l’Himalaya, 1914, par Guy Babault,
Carab., : 87, t.2, f.1.
2013. Tachys babaulti, Kushwaha & Hegde, in Souvenir:
Water and Biodiversity: 160.
Material examined: 1 ex., Lalitpur, Jailwara,
14.x.2011, Alt. 296m., Coll. B. Lal & party, Ex.
near stream (Reg. No. 19592/H4A).
Distribution: INDIA : Kerala, Madhya Pradesh,
Maharashtra, Tamil Nadu, Uttarakhand and Uttar
Pradesh; Elsewhere: Pakistan.
51. Tachys blandus Andrewes*
1924. Tachys blandus Andrewes, Mission Prov. Centr. l’Inde
Reg. Occi. l’Himalaya, 1914, par Guy Babault,
Carab.: 85, t.1, f.8.
2013. Tachys blandus, Kushwaha & Hegde, in Souvenir:
Water and Biodiversity: 160.
Material examined: 2 exs., Mathura, Kishori
Kunj, 17.xi.2010, Coll. O. P. Srivastava & party,
(Reg. No. 19676, 19677/H4A); 1 ex., Lucknow,
Kuckrail Forest Area, 17.ii.2011, Coll. V. D. Hegde
& party, Ex. at pond edge (Reg. no. 19675/H4A);
2 exs., Jalaun, Orai, Ajnari, Alt. 89 m., 17.x.2011,
Coll. B. Lal & party, Ex. leaf litter (reg. no. 19673,
19674/H4A).
Distribution: INDIA: Andhra Pradesh,
Jharkhand, Karnataka, Madhya Pradesh,
Maharashtra, Punjab, Uttarakhand and Uttar
Pradesh; Elsewhere: Pakistan.
52. Tachys dulcis Andrewes
1925. Tachys dulcis Andrewes, Ann. Mus. Civ. Sto. Nat. Gen.
51(3, 10): 395.
Distribution: INDIA : Uttar Pradesh.
53. Tachys fumigatus Motchulsky
1851. Tachys fumigatus Motchulsky, Bull. Mosc., 2: 509.
2013. Tachys fumigatus, Kushwaha & Hegde, in Souvenir:
Water and Biodiversity: 160.
Material examined: 6 exs., Lucknow, Kuckrail
Forest Area, 17.ii.2011, Coll. V. D. Hegde & party,
Ex. at pond edge; 2 exs., Hardoi, Neer, Bhais Ka
nala, 20.ii.2011, Coll. V. D. Hegde & Party, Ex. In
soil crevices; 2 exs., Shahjahanpur, Khannaut River
bank, 01.iii.2011, Coll. V. D. Hegde & Party, Ex.
On river bed; 1 ex., Hardoi, Neer, Bhais Ka nala,
20.ii.2011, Coll. V. D. Hegde & Party, Ex. Wet
land; 1 ex., Mathura, Kishori Kunj, 17.xi.2010, Coll.
O. P. Srivastava & Party; 1 ex., Barabanki, Banki,
Mohammadpur, GPS N26°56´45.1´´ E81°10´
25.0´´, 20.iii.2012, Coll. A. Sar & Party (Reg. No.
19591/H4A); 06 exs., Sitapur, Tendua Jheel,
24.ii.2011, Coll. V. D. Hegde & Party, Ex. Wet
land (Reg. no. 19651-19656/ H4A); 1 ex., Mathura,
Kishori Kunj, 17.xi.2010, Coll. O. P. Srivastava &
Party (Reg. no. 19657/H4A); 2 exs., Barabanki,
Sultanpur, 21.iii.2012, Coll. A. Sar & party (Reg.
no. 19658, 19659/H4A).
Distribution: INDIA : including Uttar Pradesh;
Elsewhere: Southern Asia from Japan to New
Guinea and from Philippines to East Africa and
Madagascar.
54. Tachys klugi (Nietner)
1858. Bembidium klugi Nietner, Ann. Mag. Nat. Hist. (3) 2:
423.
1886. Tachys klugi Bates, Ann. Mag. Nat. Hist. (5) 17: 155.
2013. Tachys klugi, Kushwaha & Hegde, in Souvenir: Water
and Biodiversity: 160.
Material examined: 4 ex., Mathura, Kishori
Kunj, 17.xi.2010, Coll. O. P. Srivastava & party,
(Reg. No. 19664-19667/H4A); 1 ex., Jalaun, Orai,
KUSHWAHA and HEGDE : Insecta : Coleoptera : Carabidae
Ajnari, Alt. 89 m., 17.x.2011, Coll. B. Lal & party
(Reg. no. 19663/H4A).
Distribution: INDIA: Common throughout
India; Elsewhere: South East Asia.
55. Tachys latus Peyron
1858. Tachys latus Peyron, Ann. Soc. Ent. Fr.: 364, t. 9, f. 3.
Distribution: INDIA: Bihar, Kashmir,
Uttarakhand, Uttar Pradesh and West Bengal;
Elsewhere: Bangladesh, Indonesia, Iraq, Myanmar,
Sri Lanka, Turkey and Vietnam.
56. Tachys latus var. virgatus Andrewes*
1925. Tachys latus var. virgatus Andrewes, Ann. Mus. Civ.
Gen., 51: 440.
2013. Tachys latus var. virgatus, Kushwaha & Hegde, in
Souvenir: Water and Biodiversity: 161.
Material examined: 1 ex., Sitapur, Tendua Jheel,
24.ii.2011, Coll. V. D. Hegde & party, Ex. Wet
Land (Reg. no. 19668/H4A).
Distribution: INDIA: Uttar Pradesh and West
Bengal.
57. Tachys opalescens Andrewes**
1925. Tachys opalescens Andrewes, Ann. Mus. Civ. Gen.,
51: 342 and 352.
2013. Tachys opalescens, Kushwaha & Hegde, in Souvenir:
Water and Biodiversity: 161.
Material examined: 1 ex., Auraiya, Fatehpur
near Muradganj, 20.x.2011, Coll. B. Lal & party,
Ex. Leaf litter (Reg. No. 19672/H4A); 1 ex.,
Lakhimpur Kheri, Dudhwa National Park,
Sathiyana, 27.ii.2011, Coll. V. D. Hegde & party,
Ex. Under leaf litter; 1 ex., Hardoi, Neer, Bhais Ka
nala, 20.ii.2011, Coll. V. D. Hegde & Party, Ex. In
soil crevices.
Distribution: INDIA: Uttar Pradesh; Elsewhere:
Sri Lanka.
58. Tachys ornatus (Apetz)
1854. Bembidium ornatus Apetz, Col. Brehm.: 12.
1935. Tachys ornatus, Andrewes, Faun. Brit. India,
Coleoptera, Carabidae 2: 16, f. 4a.
2013. Tachys ornatus, Kushwaha & Hegde, in Souvenir:
Water and Biodiversity: 161.
Material examined: 1 ex., Shahjahanpur,
409
Khannaut River Bank, 01.iii.2011, Coll. V. D.
Hegde & party, Ex. in soil crevices (Reg. No.
19662/H4A); 7 exs., Shahjahanpur, Khannaut River
bank, 01.iii.2011, Coll. V. D. Hegde & Party, Ex.
On river bed; 3 exs., Lucknow, Kuckrail Forest
Area, 17.ii.2011, Coll. V. D. Hegde & party, Ex. at
pond edge; 1 ex., Mathura, Kishori Kunj,
17.xi.2010, Coll. O. P. Srivastava & Party; 1 ex.,
Auraiya, Fatehpur nr. Muradganj, 20.x.2011, Coll.
B. Lal & party, Ex. leaf litter (Reg. no. 19660/H4A);
1 ex., Lucknow, Kuckrail Forest Area, 17.ii.2011,
Coll. V. D. Hegde & party, Ex. at pond edge (Reg.
no. 19661/H4A).
Distribution: INDIA: Jharkhand, Karnataka,
Madhya Pradesh, Maharashtra, Odisha, Tamil
Nadu, Uttarakhand and Uttar Pradesh; Elsewhere:
Egypt, Ethiopia, Namibia, Philippines, Saudi
Arabia, Sudan and Italy (?).
59. Tachys poecilopterus Bates
1873. Tachys poecilopterus Bates, Trans. Ent. Soc. Lond.,
21: 331.
2013. Tachys poecilopterus, Kushwaha & Hegde, in
Souvenir: Water and Biodiversity: 161.
Material examined: 3exs., Barabanki, Sultanpur,
21.iii.2012, Coll. A. Sar and party, (Reg. No. 1966919671/H4A).
Distribution: INDIA: Assam, Sikkim,
Uttarakhand, Uttar Pradesh and West Bengal;
Elsewhere: Cambodia, China, Indonesia, Laos,
Malaysia, Myanmar, Singapore, Thailand and
Vietnam.
60. Tachys politus Motchulsky
1851. Tachys politus Motchulsky, Bull. Soc. Imp. Nat. Mosc.,
2: 509.
2013. Tachys politus, Kushwaha & Hegde, in Souvenir:
Water and Biodiversity: 161.
Material examined: 7 exs., Auraiya, Fatehpur
near Muradganj, 20.x.2011, Coll. B. Lal & party,
Ex. leaf litter (Reg. No. 19638-19641, 19643,
19644, 19647/H4A); 3 exs., Mathura, Kishori Kunj,
Coll. O. P. Srivastava & party (Reg. no. 19642,
19645, 19646/ H4A); 1 ex., Jhansi, Ambabai,
12.x.2011, Coll. B. Lal & party, Ex. Dusting (Reg.
no. 19648/H4A); 1 ex., Shahjahanpur, Sugarcane
410
Fauna of Uttar Pradesh, State Fauna Series, 22
Research Institute rest house campus, 09.x.2006,
Coll. P. Mukhopadhyay & party (Reg. no. 19649/
H4A).
Distribution: INDIA: including Uttar Pradesh;
Elsewhere: throughout South-East Asia except
Japan.
61. Tachys politus var. aspilotus Andrewes
1925. Tachys politus var. aspilotus Andrewes, Ann. Mus. Civ.
Gen., 51: 449.
2013. Tachys politus var. aspilotus, Kushwaha & Hegde, in
Souvenir: Water and Biodiversity: 160.
Material examined: 1 ex., Auraiya, Fatehpur
near Muradganj, 20.x.2011, Coll. B. Lal & party,
Ex. leaf litter (Reg. No.19650/H4A); 1 ex., Lalitpur,
Jailwara, Alt. 296m, 14.x.2011, Coll. B. Lal &
Party, Ex. near stream (Reg. no. 19590/H4A).
Subfamily 7. BROSCINAE Hope, 1838
Tribe 11. Broscini Hope, 1838
Genus 36. Axonya Andrewes,1923
1923. Axonya Andrewes, Ann. Mag. Nat. Hist., (9) 12: 679.
64. Axonya championi Andrewes
1923. Axonya championi Andrewes, Ann. Mag. Nat. Hist.,
(9) 12: 680.
2011. Boscus punctatus, Kushwaha & Pal, Advancements in
Invertebrate Taxonomy and Biodiversity:139-146.
Distribution: INDIA: Arunachal Pradesh,
Assam, Himachal Pradesh, Mizoram, Sikkim,
Uttarakhand, Uttar Pradesh and West Bengal.
Genus 37. Broscus Panzer, 1813
1813. Broscus Panzer, Index Ent.: 62.
1828. Cephalotes Dejean, Spec. Gen. Col., 3: 426.
Distribution: INDIA: Throughout India
including Uttar Pradesh; Elsewhere: throughout
South-East Asia except Japan.
62. Tachys ceylanicus (Nietner)*
1858. Bembidium ceylanicus Nietner, Ann. Mag. Nat. Hist.,
(3)2: 423.
1935. Tachys ceylanicus, Andrewes, Faun. Brit. Ind.
Carabidae 2, Harpalinae, 1: 257.
Material examined: 1 ex., Lakhimpur Kheri,
Salukapur, Dudhwa National Park, GPS N28° 25´
5´´E80°43´9´´, 02.iii.2013, Coll. V. D. Hegde &
party (reg. no. 20053/H4A).
Distribution: INDIA: Karnataka, Uttarakhand
and Uttar Pradesh; Elsewhere: Sri Lanka.
63. Tachys latissimus var. gracilis
(Motchulsky)*
1862. Elaphropus gracilis Motchulsky, Etud. Ent., 11: 36.
1928. Tachys latissimus var. gracilis, Andrewes, Trans. Ent.
Soc. Lond. : 4, 20.
Material examined: 1 ex., Lakhimpur Kheri,
Dudhwa National Park, Sathiyana, 01.iii.2013, Coll.
V. D. Hegde & Party.
Distribution: INDIA: Bihar, Maharashtra,
Meghalaya, Nicobar Island, Uttarakhand, Uttar
Pradesh and West Bengal; Elsewhere: Indonesia,
Myanmar and Thailand.
65. Broscus punctatus (Dejean)
1828. Cephalotes punctatus Dejean, Spec. Gen. Col., 3: 431.
1930. Broscus punctatus, Andrewes, Cat. Ind. Ins., Part 18.
Carabidae: 55.
Distribution: INDIA: north of the Ganges,
including the whole Himalayan tract and
Meghalaya; Elsewhere: China, Egypt, Iraq, Saudi
Arabia and Turkey.
Subfamily 8. BRACHININAE Bonelli, 1810
Tribe 12. Brachinini Bonelli, 1810
Subtribe Mastacina Erwin, 1970
Genus 38. Mastax Fischer von Waldheim, 1825*
1825. Mastax Fischer von Waldheim, Ent. Russ., 3: 111.
Mastax sp.
Material examined:1ex., Saharanpur, Rajaji
sanctuary, Saharanpur road, 11.vi.1964, Coll. T. D.
Soota & party, Ex. under stone; 1 ex., Chitrakoot,
Pahari, 26.xi.2010, Coll. O. P. Srivastava & party,
Ex. Crevices in wheat field soil; 1 ex., Rae Bareilly,
Lohanipur, 06.iii.2010, Coll. V. D. Hegde & party,
Ex. bank of Sai River.
Subtribe Brachinina Bonelli, 1810
Genus 39. Brachinus Weber, 1801
1801. Brachinus Weber, Obs. Ent.: 22.
KUSHWAHA and HEGDE : Insecta : Coleoptera : Carabidae
66. Brachinus hexagrammus Chaudoir
1876. Brachinus hexagrammus Chaudoir, Ann. Soc. Ent.
Belg.: 55.
Material examined: 8 exs., Jalaun, Chili near
Orai, 15.x.2011, Coll. B. Lal & party, Ex. at light
(Reg. no. 20070-20077/H4A).
Distribution: INDIA : Throughout INDIA
including Uttar Pradesh.
67. Brachinus modestus Schmidt-Goebel*
1846. Brachinus modestus Schmidt-Goebel, Faun. Col.
Birm.: 73.
Material examined: 1 ex., Jhansi, Pathrora near
Barua Sagar, 11.x.2011, Alt. 165m, Coll. B. Lal &
party, Ex. under leaf litter (Reg. no. 20082/H4A).
Distribution: INDIA: Uttar Pradesh and West
Bengal; Elsewhere: Myanmar and Vietnam.
68. Brachinus stevensi Andrewes*
1924. Brachinus stevensi Andrewes, Mission Prov. Centr.
l‘Inde Reg. Occi. l‘Himalaya 1914, par Guy Babault:
107, t. 4. f. 2.
Material examined: 4 exs., Sitapur, Sinhanipur
Jheel, 23.ii.2011, Coll. V. D. Hegde & party, Ex.
Crevices in soil (Reg. no. 20078-20081/H4A).
Distribution: INDIA: Arunachal Pradesh,
Assam, Himachal Pradesh, Uttarakhand, Uttar
Pradesh and West Bengal.
69. Brachinus suturellus Chaudoir
1876. Brachinus suturellus Chaudoir, Ann. Soc. Ent. Belg.:
69.
Distribution: INDIA: Himachal Pradesh,
Jharkhand, Karnataka, Madhya Pradesh,
Uttarakhand and Uttar Pradesh; Elsewhere:
Myanmar and Vietnam(?).
Subtribe Pheropsophina Jeannel, 1948
Genus 40. Pheropsophus Solier, 1833
1833. Pheropsophus Solier, Ann. Soc. Ent. Fr., 2(1): 461.
70. Pheropsophus assimilis Chaudoir**
1876. Pheropsophus assimilis Chaudoir, Ann. Soc. Ent. Belg.:
38.
Material examined: 2 exs., Hardoi, Sandi Bird
Sanctuary, 21.ii.2011, Coll. V. D. Hegde & party,
411
Ex. Soil crevices of ploughed field (Reg. no. 20083,
20084/H4A).
Distribution: INDIA : Uttar Pradesh; Elsewhere:
Thailand and China.
71. Pheropsophus catoirei (Dejean)
1825. Brachinus catoirei Dejean, Spec. Gen. Col., 1: 301.
1924. Pheropsophus catoirei, Andrewes, Mission Prov.
Centr. l‘Inde Reg. Occi. l‘Himalaya 1914, par Guy
Babault: 55.
Material examined: 13 exs., Hardoi, Sandi Bird
Sanctuary, 21.ii.2011, Coll. V. D. Hegde & party,
Ex. Crevices of Soil; 1 ex., Hardoi, Sandi Bird
Sanctuary, 21.ii.2011, Coll. V. D. Hegde & party,
Ex. Crevices of Soil (Reg. no. 20088/H4A); 4 exs.,
Hardoi, Sandi Bird Sanctuary, 21.ii.2011, Coll. V.
D. Hegde & party, Ex. crevices of ploughed field;
1 ex., Ballia F.R.H., 15.ix.2010, Coll. V. D. Hegde
& party; 1 ex., Mirzapur, Chunar, 08.ii.2009, Coll.
V. D. Hegde & party, Ex. Ganga river bed; 1 ex.,
Chandauli, Bhansi, 27.ix.2010, Coll. V. D. Hegde
& party, Ex. under bricks; 1 ex., Bijnor, Indira park
near Director Wildlife & forest office, 22.vii.2008,
Coll. V. D. Hegde & party; 1 ex., Sitapur, Tendua
Jheel, 24.ii.2011, Coll. V. D. Hegde & party, Ex.
soil crevices of ploughed field; 1 ex., Ghaziabad,
Moorta, 10.ii.2009, Coll. R. S. Mridha & party.
Distribution: INDIA: Throughout India,
including Westward to Kashmir and the Khyber
Pass, Meghalaya, Andaman Island and Uttar
Pradesh; Elsewhere: Myanmar, Pakistan and
Sri Lanka.
72. Pheropsophus consularis Schmidt-Goebel*
1846. Pheropsophus consularis Schmidt-Goebel, Faun. Col.
Birm., : 75.
Material examined: 2 exs., Pilibhit, Mala Forest,
27.ix.1974, Alt. 250m, Coll. P. K. Maiti and party
(Reg. no. 20085, 20086/H4A).
Distribution: INDIA: Arunachal Pradesh,
Assam, Bihar, Himachal Pradesh, Jharkhand,
Odisha, Uttarakhand, Uttar Pradesh and West
Bengal; Elsewhere: Bangladesh and Myanmar.
73. Pheropsophus hilaris (Fabricius)
1798. Carabus hilaris Fabricius, Suppl. Ent. Syst., 56.
412
Fauna of Uttar Pradesh, State Fauna Series, 22
1921. Pheropsophus hilaris, Andrewes, Trans. Ent. Soc.
Lond., 69: 159.
Uttar Pradesh and West Bengal; Elsewhere:
Bangladesh.
Material examined: 3exs.,Chandauli, Bansi,
27.ix.2010, Coll. V. D. Hegde & party, Ex. Under
bricks; 1 ex., Chandauli, Bansi, 27.ix.2010, Coll.
V. D. Hegde & party, Ex. Under bricks (Reg. no.
20087/H4A); 2 exs., Lakhimpur Kheri, Dudhwa
National park, Sathiyana, GPS N28° 32´ 30´´ E80°
32´ 30´´, 01.iii.2013, Coll. V. D. Hegde & party.
78. Chlaenius bioculatus Chaudoir
Distribution: INDIA: Throughout India
including N. Andaman and Uttar Pradesh;
Elsewhere: Myanmar and Pakistan.
74. Pheropsophus lineifrons Chaudoir
1850. Pheropsophus lineifrons Chaudoir, Bull. Soc. Imp. Nat.
Mosc., 1: 80.
Distribution: INDIA: Assam, Bihar, Himachal
Pradesh, Jharkhand, Maharashtra, Uttarakhand,
Uttar Pradesh and West Bengal.
Subfamily 9. LICININAE Bonelli, 1810
Tribe 13. Chlaeniini Brulle, 1834
Genus 41. Callistomimus Chaudoir, 1872
1872. Callistomimus Chaudoir, Bull. Soc. Imp. Nat. Mosc.,
1: 382.
75. Callistomimus d’abreui Andrewes
1921. Callistomimus d’abreui Andrewes, Proc. Zool. Soc.
Lond., 1: 237 and 242, t.1, f.9.
Distribution: INDIA: Bihar, Madhya Pradesh
and Uttar Pradesh.
Genus 42. Chlaenius Dejean, 1826
1826. Chlaenius Dejean, Spec. Gen. Col., 2: 297.
76. Chlaenius agilis Chaudoir
1856. Chlaenius agilis Chaudoir, Bull. Soc. Imp. Nat. Mosc.,
2: 246.
Distribution: INDIA: Arunachal Pradesh,
Assam, Himachal Pradesh, Nim Nadi,Uttarakhand,
Uttar Pradesh and West Bengal; Elsewhere:
Cambodia, Laos and Vietnam.
77. Chlaenius bimaculatus lynx Chaudoir
1856. Chlaenius bimaculatus var. lynx Chaudoir, Bull. Soc.
Imp. Nat. Mosc., 2:199.
Distribution: INDIA: Bihar, Manipur, Odisha,
1856. Chlaenius bioculatus Chaudoir, Bull. Soc. Imp. Nat.
Mosc., 2:198.
Distribution: INDIA: Throughout India
including Andaman Island; Elsewhere: Borneo,
Cambodia, China, Indonesia, Japan, Laos,
Myanmar, Sri Lanka and Vietnam.
79. Chlaenius chlorodius Dejean
1826. Chlaenius chlorodius Dejean, Spec. Gen. Col., 2: 365.
Distribution: INDIA: Assam, Bihar, Madhya
Pradesh, Maharashtra, Uttarakhand, Uttar Pradesh
and West Bengal; Elsewhere: Bangladesh.
80. Chlaenius duvauceli (Dejean)
1831. Epomis duvauceli Dejean, Spec. Gen. Col., 5: 668.
1924. Chlaenius duvauceli, Andrewes, Mission Prov. Centr.
l‚ Inde Reg. Occi. l‚Himalaya 1914, par Guy Babault:
25.
Distribution: INDIA: Assam, Bengal, Bihar,
Gujarat, Haryana, Maharashtra, Tamil Nadu and
Uttar Pradesh.
81. Chlaenius laetiusculus Chaudoir
1856. Chlaenius laetiusculus Chaudoir, Bull. Soc. Imp. Nat.
Mosc., 2: 248.
Distribution: INDIA: Throughout Himalayan
Tract including Assam and Uttar Pradesh;
Elsewhere: Pakistan and Sri Lanka.
82. Chlaenius luculentus Andrewes
1920. Chlaenius luculentus Andrewes, Ent. Month. Mag.:
235, 236.
Distribution: INDIA (Bihar, Maharashtra,
Odisha, Uttar Pradesh and West Bengal).
83. Chlaenius pretiosus Chaudoir
1856. Chlaenius pretiosus Chaudoir, Bull. Soc. Imp. Nat.
Mosc., 2: 288.
Material examined: 1 ex. Agra, Kakuabaad,
29.viii.2009, Coll. V. D. Hegde & party, Ex. In soil
crevices.
Distribution: INDIA: Delhi, Gujarat, Rajasthan,
Uttarakhand and Uttar Pradesh; Elsewhere:
Pakistan and Sri Lanka.
KUSHWAHA and HEGDE : Insecta : Coleoptera : Carabidae
84. Chlaenius scapularis Chaudoir*
1876. Chlaenius scapularis Chaudoir, Ann. Soc. Ent. Belg.,
8: 98.
Material examined: 3 ex., Chitrakoot, Pahari,
26.xi.2010, Coll. O. P. Srivastava & party, Ex.
Crevices in Wheat field soil; 2 exs., Hardoi, Pihani,
30.vi.1989, Coll. K. K. Ray & party.
Distribution: INDIA: Bihar, Chhattisgarh, Delhi,
Himachal Pradesh, Karnataka, Madhya Pradesh,
Tamil Nadu, Uttarakhand, Uttar Pradesh and West
Bengal; Elsewhere: Sri Lanka.
85. Chlaenius trinototus Chaudoir*
1876. Chlaenius trinototus Chaudoir, Ann. Soc. Ent. Belg.,
8: 279.
Material examined: 1 ex., Lakhimpur Kheri,
Dudhwa FRH Compound, 23.ix.1974, Coll. P. K.
Maiti and Party, Ex. at light
Distribution: INDIA: Bihar, Uttarakhand and
Uttar Pradesh.
86. Chlaenius variipes Chaudoir
1856. Chlaenius variipes Chaudoir, Bull. Soc. Imp. Nat.
Mosc., 2: 268.
413
1930. Harpaglossus opacus, Andrewes, Cat. Ind. Insects,
Part 18, Carabidae: 175.
Distribution: INDIA: Gujarat, Rajasthan, Tamil
Nadu, Uttar Pradesh and West Bengal; Elsewhere:
Sri Lanka.
Genus 44. Rhopalopalpus Laferte, 1851
1851. Rhopalopalpus Laferte, Ann. Soc. Ent. Fr., 9(2): 212.
89. Rhopalopalpus janthinus (Redtenbacher)
1844. Chlaenius janthinus Redtenbacher, Hugel’s kaschmir
iv, 2: 500.
1930. Rhopalopalpus janthinus, Andrewes, Cat. Ind. Insects
Part 18, Carabidae: 290.
Distribution: INDIA: Himachal Pradesh,
Kashmir, Punjab, Uttarakhand and Uttar Pradesh;
Elsewhere: Pakistan.
Tribe 14. Oodini Laferte-Senectere, 1851
Genus 45. Oodes Dejean, 1826
1826. Oodes Dejean, Spec. Gen. Col., 2: 374.
90. Oodes subolivaceus Laferte
1851. Oodes subolivaceus Laferte, Ann. Soc. Ent. Fr.: 271
(note).
Distribution: INDIA: Bihar, Delhi, Gujarat,
Jharkhand, Madhya Pradesh and Uttar Pradesh;
Elsewhere: Pakistan.
Distribution: INDIA: Bihar, Jharkhand,
Karnataka, Maharashtra, Uttarakhand, Uttar
Pradesh and West Bengal; Elsewhere: Nepal and
Sri Lanka.
87. Chlaenius xanthospilus (Wiedeman)
Genus 46. Systolocranius Chaudoir, 1857
1821. Carabus xanthospilus Wiedeman, Germ. Mag. Ent.,
4: 115.
1857. Systolocranius Chaudoir, Bull. Soc. Imp. Nat. Mosc.
2: 23.
1921. Chlaenius xanthospilus Andrewes, Trans. Ent. Soc.
Lond.: 167.
91. Systolocranius linea (Wiedeman)
1821. Oodes linea Wiedeman, Germ. Mag., 4: 113.
Material examined: 1ex., Azamgarh, Kusum
Lodge, 21.iii.2009, Coll. A. Bal and Party, Ex. at
light.
Distribution: INDIA: Throughout India
including Uttar Pradesh; Elsewhere: Cambodia,
Laos, Sri Lanka, Thailand and Vietnam.
Genus 43. Harpaglossus Motchulsky, 1858
1858. Harpaloglossus Motchulsky, Etud. Ent.: 173.
88. Harpaglossus opacus Chaudoir
1857. Ceroglossus opacus Chaudoir, Bull. Soc. Imp. Nat.
Mosc., 2: 8.
1921. Systolocranius linea, Andrewes, Trans. Ent. Soc.
Lond.: 167.
Distribution: INDIA: Bihar, Madhya Pradesh,
Maharashtra, Uttar Pradesh and West Bengal.
Tribe 15. Licinini Bonelli, 1810
Genus 47. Badister Schellenberg, 1806*
1806. Badister Schellenberg, Helv. Ent., 2: 90 and 91.
Badister sp.
Material examined: 1 ex., Auraiya, near Bus
station, 18.x.2011, Coll. B. Lal & party, Ex. At light;
414
Fauna of Uttar Pradesh, State Fauna Series, 22
1 ex., Jalaun, Chili near Orai, 15.x.2011, Coll. B.
Lal & party, Ex. At light; 1 ex., Hardoi, Hardoi I.
H. Bunglow, 10.x.2006, Coll. P. Mukhopadhyay
& party, Ex. At light; 2 exs., Lucknow, Alambag,
12.vii.1989, Coll. K. K. Ray & party; 10 exs.,
Sultanpur, Panchopiran, 23.iii.2012, Coll. A. Sar
& party, Ex. Under bricks.
Genus 48. Diplocheila Brulle,
1834. Diplocheila Brulle, Hist. Nat des ins., 4: 407.
92. Diplocheila polita (Fabricius)
1792. Carabus polita Fabricius, Ent. Syst., 1(1): 146.
1922. Diplocheila polita, Andrewes, Ann. Mag nat. Hist. (9)
9: 283.
Material examined: 1 ex., Hardoi, Neer, Bhais
ka nala, 20.ii.2011, Coll. V. D. Hegde & party, Ex.
In soil crevices (Reg. No. 20090/H4A).
Distribution: INDIA: Throughout India
including Uttar Pradesh; Elsewhere: Myanmar and
Sri Lanka.
93. Diplocheila cordicollis Laferte*
Subfamily 10. CARABINAE Latreille, 1802
Tribe 16. Carabini Latreille, 1802
Genus 49. Carabus Linnaeus, 1758
1758. Carabus Linnaeus, Syst. Nat.Ed. 10: 413.
95. Carabus barysomus Bates
1889. Carabus barysomus Bates, Proc. Zool. Soc. Lond.:
210.
Distribution:INDIA : Jammu-Kashmir and Uttar
Pradesh; Elsewhere: Pakistan.
Subfamily 11. CICINDELINAE Latreille, 1802
Tribe 17. Cicindelini Latreille, 1802
Subtribe Cicindelina Latreille, 1802
Genus 50. Cicindela Linnaeus, 1758
1758. Cicindela Linnaeus, Syst. Nat., 1: 407.
96. Cicindela (Calochroa) octonotata
Wiedemann
1819. Cicindela octonotata Wiedemann, Zool. Mag., i, 3:
168.
1989. Cicindela (Calochroa) octonotata, Acciavatti and
Pearson, Ann. Carn. Mus., 58: 116.
1851. Diplocheila cordicollis Laferte, Ann. Soc. Ent. Fr.: 279.
1959. Diplocheila (Isorembus) cordicollis, Ball, Mem. Amer.
Ent. Soc., (16): 54.
Material examined: 1 ex., Jhansi, Babina,
21.xi.2010, Coll. O. P. Srivastava & party (Reg.
no. 20049/H4A).
Distribution: INDIA: Bihar, Delhi, Maharashtra,
Uttar Pradesh and West Bengal; Elsewhere:
Bangladesh.
94. Diplocheila (s. str.) daldorfi (Crotch)*
1871. Rembus daldorfi Crotch, List of all Coleoptera
Described A.D. 1758-1821, Referred to other Modern
Genera: 9.
1959. Diplocheila (s.str.) daldorfi, Ball, Mem. Amer. Ent.
Soc., (16): 25.
Material examined: 1 ex., Varanasi,
Dashashwamedh Ghat, 28.ix.2010, Coll. V. D.
Hegde & party, Ex. Under light (Reg. No. 20089/
H4A).
Distribution: INDIA: Assam, Bihar, Tamil
Nadu, Uttar Pradesh and West Bengal; Elsewhere:
Myanmar, Singapore and Vietnam.
Distribution: INDIA: Arunachal Pradesh,
Assam, Bihar, Himachal Pradesh, Manipur,
Meghalaya, Nagaland, Sikkim, Uttar Pradesh and
West Bengal; Elsewhere: Bangladesh, Myanmar
and Nepal.
97. Cicindela (Calochroa) flavomaculata Hope
1831. Cicindela flavomaculata Hope, Zool. misc., 1: 21.
1989. Cicindela (Calochroa) flavomaculata, Acciavatti and
Pearson, Ann. Carn. Mus., 58: 116.
Material examined: 2 exs., Varanasi, Babatpur,
24.x.2006, Coll. A. Bal & party; 2 exs., Allahabad,
Ramnagar, 22.x.2006, Coll. A. Bal & party; 4 exs.
Shahjahanpur, Jalalabad, 15.x.2006, Coll. A. Bal
& party; 10 exs., Ballia, Surhatal W.L.S., 12.ii.2009,
Coll. V. D. Hegde & party; 4 exs., Hardoi, I. H.
Dak Bunglow, 10.x.2006, Coll. P. Mukhopadhyay
& party, Ex. at light.
Distribution: INDIA: Assam, Haryana,
Jharkhand, Karnataka, New Delhi, Madhya
Pradesh, Punjab, Uttarakhand, Uttar Pradesh and
West Bengal; Elsewhere: South East Asia and
Philippines.
KUSHWAHA and HEGDE : Insecta : Coleoptera : Carabidae
415
98. Cicindela (Chaetodera) albina Wiedemann
103. Cicindela (Spilodia) multiguttata Dejean
1819. Cicindela albina Wiedemann, Zool. Mag., i, 3: 169.
1825. Cicindela (Spilodia) multiguttata Dejean, Spec. Gen.
Col., 1: 109.
1989. Cicindela (Chaetodera) albina, Acciavatti and
Pearson, Ann. Carn. Mus. 58: 140.
Distribution: INDIA: Bihar, Haryana, Odisha,
Punjab, Uttar Pradesh and West Bengal; Elsewhere:
Bangladesh and Pakistan.
99. Cicindela (Chaetodera) vigintiguttata Herbst
1806. Cicindela (Chaetodera) vigintiguttata Herbst, Nat.
Syst. Ins. Kaf., 10: 174, pl. 171, fig. 9.
1989. Cicindela (Chaetodera) vigintiguttata, Acciavatti and
Pearson, Ann. Carn. Mus., 58: 141.
Material examined: 3 exs., Sultanpur, 15.x.2006,
Coll. P. Mukhopadhyay & party, Ex. Gomati River
Bank.
Distribution: INDIA: Assam, Bihar, Haryana,
Odisha, Punjab, Sikkim, Uttar Pradesh and West
Bengal; Elsewhere: Bangladesh, Bhutan and
Pakistan.
100. Cicindela (Lophyra) catena Fabricius
1775. Cicindela (Lophyra) catena Fabricius, Syst. Ent.: 226.
1989. Cicindela (Lophyra) catena, Acciavatti and Pearson,
Ann. Carn. Mus., 58: 143-144.
Distribution: INDIA: Andhra Pradesh,
Jharkhand, Karnataka, Kerala, Madhya Pradesh,
Puducherry, Sikkim, Tamil Nadu, Uttarakhand,
Uttar Pradesh and West Bengal; Elsewhere:
Sri Lanka.
101. Cicindela (Lophyra) cancellata
intemperata Acciavatti & Pearson
1882. Cicindela cancellata viridula Mandl, Kol. Rund., 56:
65.
1989. Cicindela (Lophyra) cancellata intemperata Acciavatti
& Pearson, Ann. Carn. Mus., 58: 147, 148.
Distribution: INDIA : Assam, Gujarat, Haryana,
Sikkim, Uttar Pradesh and West Bengal; Elsewhere:
Nepal, Pakistan, Sri Lanka and Thailand.
102. Cicindela (Spilodia) vittigera Dejean
1825. Cicindela (Spilodia) vittigera Dejean, Spec. Gen. Col.,
1: 107.
1989. Cicindela (Spilodia) vittigera, Acciavatti & Pearson,
Ann. Carn. Mus., 58: 154.
Distribution: INDIA: Bihar, Haryana, Manipur,
Punjab, Uttar Pradesh and West Bengal; Elsewhere:
Bangladesh, Nepal and Pakistan.
1989. Cicindela (Spilodia) multiguttata, Acciavatti &
Pearson, Ann. Carn. Mus., 58: 155.
Distribution: INDIA: Arunachal Pradesh,
Assam, Haryana, Punjab, Uttar Pradesh and West
Bengal; Elsewhere: Bangladesh, Bhutan and Nepal.
104. Cicindela (Lophyridia) angulata Fabricius
1798. Cicindela (Lophyridia) angulata Fabricius, Suppl. Ent.
Syst.: 62.
1989. Cicindela (Lophyridia) angulata, Acciavatti &
Pearson, Ann. Carn. Mus., 58: 157.
Material examined: 1 ex.♂, 2 exs.♀, Gorakhpur,
Raj Ghat, Rapti River Bed, 17.iii.2010, Coll. V. D.
Hegde & party, Ex. by hand net; 1 ex., Sultanpur,
Sultanpur, 15.x.2006, Coll. P. Mukhopadhyay &
party, Ex. Gomati River Bank.
Distribution: INDIA : Andhra Pradesh, Assam,
Bihar, Goa, Gujarat, Haryana, Karnataka, Kerala,
Manipur, Puducherry, Punjab, Sikkim, Tamil Nadu,
Uttar Pradesh and West Bengal; Elsewhere:
Bangladesh, China, Myanmar, Pakistan, Nepal and
Vietnam.
105. Cicindela (Lophyridia) plumigera
macrograptina Acciavatti & Pearson
1989. Cicindela (Lophyridia) plumigera macrograptina
Acciavatti & Pearson, Ann. Carn. Mus., 58: 162.
Distribution: INDIA: Assam, Haryana, Uttar
Pradesh and West Bengal; Elsewhere: Bangladesh
and Nepal.
106. Cicindela (Lophyridia) chloris Hope
1831. Cicindela (Lophyridia) chloris Hope, Zool. misc., 1:
21.
1989. Cicindela (Lophyridia) chloris, Acciavatti & Pearson,
Ann. Carn. Mus., 58: 164.
Distribution: INDIA: Arunachal Pradesh,
Assam, Bihar, Haryana, Himachal Pradesh, JammuKashmir, Punjab, Sikkim, Uttar Pradesh and West
Bengal; Elsewhere: Bhutan, Nepal and Pakistan.
107. Cicindela (Eugrapha) bigemina Klug
1834. Cicindela bigemina Klug, Jahrb. Ins.:30.
1989. Cicindela (Eugrapha) bigemina, Acciavatti & Pearson,
Ann. Carn. Mus., 58: 287.
416
Fauna of Uttar Pradesh, State Fauna Series, 22
Distribution: INDIA: Bihar, Jammu-Kashmir,
Karnataka, Madhya Pradesh, Punjab, Uttar Pradesh
and West Bengal; Elsewhere: Nepal and Pakistan.
108. Cicindela (Eugrapha) brevis Horn
Distribution: INDIA: Andhra Pradesh, Bihar,
Haryana, Karnataka, Kerala, Odisha, Punjab, Tamil
Nadu, Uttar Pradesh and West Bengal; Elsewhere:
Myanmar and Pakistan.
1905. Cicindela bigemina brevis Horn, Deut. Ent. Zeit.: 34.
113. Cicindela (Eugrapha) venosa Kollar
1989. Cicindela (Eugrapha) brevis, Acciavatti & Pearson,
Ann. Carn. Mus., 58: 288.
Material examined: 1 ex., Saharanpur,
Beharigarh, 20.vii.1971, Coll. J.C. Tripathi and
Party.
Distribution: INDIA : Bihar, Haryana, Himachal
Pradesh, Punjab and Uttar Pradesh; Elsewhere:
Pakistan.
109. Cicindela (Eugrapha) minuta Olivier
1790. Cicindela minuta Olivier, Coleopteres, 2: 31, pl. 2,
fig. 13a, b.
1989. Cicindela (Eugrapha) minuta, Acciavatti & Pearson,
Ann. Carn. Mus., 58: 291.
Distribution: INDIA : Andhra Pradesh, Gujarat,
Karnataka, Kerala, Puducherry, Tamil Nadu, Uttar
Pradesh and West Bengal; Elsewhere: Bangladesh,
Indonesia, Kampuchea, Nepal and Vietnam.
110. Cicindela (Eugrapha) cognata Wiedemann
1823. Cicindela cognata Wiedemann, Zool. Mag. (1) 2: 66.
1989. Cicindela (Eugrapha) cognata, Acciavatti & Pearson,
Ann. Carn. Mus., 58: 292.
Distribution: INDIA: Andhra Pradesh, Bihar,
Haryana, Odisha, Punjab, Tamil Nadu, Uttar
Pradesh and West Bengal; Elsewhere: Bangladesh
and Nepal.
111. Cicindela (Eugrapha) erudita Wiedemann
1823. Cicindela erudita Wiedemann, Zool Mag.(1) 2: 68.
1989. Cicindela (Eugrapha) erudita, Acciavatti & Pearson,
Ann. Carn. Mus., 58: 294.
Distribution: INDIA: Assam, Bihar, Haryana,
Himachal Pradesh, Jammu-Kashmir, Jharkhand,
Madhya Pradesh, Punjab, Sikkim, Uttar Pradesh
and West Bengal; Elsewhere: Nepal.
112. Cicindela (Eugrapha) agnata Fleutiaux
1890. Cicindela (Eugrapha) agnata Fleutiaux, Ann. Soc. Ent.
Belg., 34: 168.
1989. Cicindela (Eugrapha) agnata, Acciavatti & Pearson,
Ann. Carn. Mus., 58: 295.
1836. Cicindela venosa Kollar,Wien. Mus. Ann., 1: 331.
1989. Cicindela (Eugrapha) venosa, Acciavatti & Pearson,
Ann. Carn. Mus., 58: 298.
Distribution: INDIA: Assam, Gujarat, Haryana,
New Delhi, Punjab, Uttar Pradesh and West Bengal;
Elsewhere: Bangladesh, Nepal and Pakistan.
114. Cicindela (Eugrapha) grammophora
Chaudoir
1852. Cicindela grammophora Chaudoir, Bull. Soc. Imp. Nat.
Mosc., (1) 25: 7.
1989. Cicindela (Eugrapha) grammophora, Acciavatti &
Pearson, Ann. Carn. Mus., 58: 300.
Distribution: INDIA: Bihar, Haryana, New
Delhi, Odisha, Punjab, Rajasthan, Uttar Pradesh and
West Bengal; Elsewhere: Bangladesh, Nepal and
Pakistan.
115. Cicindela (Eriodera) albopunctata
Chaudoir*
1852. Cicindela albopunctata Chaudoir, Bull. Soc. Imp. Nat.
Mosc., (1) 25: 10.
1989. Cicindela (Eriodera) albopunctata, Acciavatti &
Pearson, Ann. Carn. Mus., 58: 304.
Material examined: 2 ex., Shahjahanpur,
Khannaut river bank, 01.iii.2011, Coll. V. D. Hegde
& party, Ex. on river bed.
Distribution: INDIA: Assam, Himachal Pradesh,
Punjab, Sikkim, Uttarakhand, Uttar Pradesh and
West Bengal; Elsewhere: Nepal and Bhutan.
116. Cicindela (Myriochile) melancholica
Fabricius
1798. Cicindela melancholica Fabricius, Ent. Syst. Suppl.:
63.
1989. Cicindela (Myriochile) melancholica, Acciavatti &
Pearson, Ann. Carn. Mus., 58: 308.
Material examined: 1 ex., Basti, Basti,
19.vi.2002, Coll. M. Prasad and Party.
Distribution: INDIA : Bihar, Haryana, Madhya
KUSHWAHA and HEGDE : Insecta : Coleoptera : Carabidae
Pradesh, Maharashtra, New Delhi, Punjab,
Rajasthan, Uttar Pradesh and West Bengal;
Elsewhere: Pakistan and other Palaearctic Regions.
117. Cicindela (Myriochile) undulata Dejean
1826. Cicindela undulata Dejean, Spec. Gen. Col., l: 96.
1989. Cicindela (Myriochile) undulata, Acciavatti &
Pearson, Ann. Carn. Mus., 58: 309.
Distribution: INDIA : Andhra Pradesh, Haryana,
Jharkhand, Karnataka, Kerala, Madhya Pradesh,
Odisha, Puducherry, Punjab, Tamil Nadu, Uttar
Pradesh and West Bengal; Elsewhere: Bangladesh,
China and Sri Lanka.
118. Cicindela (Myriochile) atelesta Chaudoir
417
121. Oxylobus porcatus Fabricius
1798. Scarites porcatus Fabricius, Suppl. Ent. Syst.: 43.
Distribution: INDIA: Andhra Pradesh,
Jharkhand, Karnataka, Kerala, Madhya Pradesh,
Maharashtra, Odisha, Punjab, Tamil Nadu, Uttar
Pradesh and West Bengal; Elsewhere: Pakistan and
Sri Lanka.
Genus 54. Scarites Fabricius, 1775
1775. Scarites Fabricius, Syst. Ent.: 249.
122. Scarites dyschromus Chaudoir
1855. Scarites dyschromus Chaudoir, Bull. Soc. Imp. Nat.
Mosc., 1: 78.
1854. Cicindela atelesta Chaudoir, Bull. Soc. Imp. Nat.
Mosc., (1) 27: 115.
Material examined: 1 ex., Sultanpur, Payagpur,
20.ix.2007, Coll. A. Bal & party.
1989. Cicindela (Myriochile) atelesta, Acciavatti & Pearson,
Ann. Carn. Mus., 58: 311.
Distribution: INDIA: North India including
Uttar Pradesh.
Distribution: INDIA: Bihar, Sikkim, Uttar
Pradesh and West Bengal; Elsewhere: Nepal.
123. Scarites indus Olivier
Tribe 18. Megacephalini Laporte, 1834
Subtribe Megacephalina Laporte, 1834
Genus 51. Megacephala Latreille, 1802
1802. Megacephala Latreille, Hist. Nat. Ins., 3: 79.
119. Megacephala euphratica var.armeniaca
Castelnau
1834. Megacephala euphratica var.armeniaca Castelnau,
Rev Ent. Silb., 2: 28.
Distribution: INDIA : Uttar Pradesh; Elsewhere:
Afganistan, Armenia and Iran.
1795. Scarites Indus Olivier, Ent.: 36 and 9, t. 1, f. 2.
Material examined: 1 ex., Etawah, Saraiya
Village, 22.x.2011, Coll. B. Lal & party.
Distribution: INDIA: Throughout India
including Uttar Pradesh; Elsewhere: Sri Lanka.
124. Scarites laticeps Andrewes*
1929. Scarites laticeps Andrewes, Faun. Brit. India,
Coleoptera, Carabidae, 1: 283.
Material examined: 1 ex., Mau, Social Forestry,
21.ix.2010, Coll. V. D. Hegde & party.
Subfamily 12. SCARITINAE Bonelli, 1810
Distribution: INDIA: Madhya Pradesh, Tamil
Nadu and Uttar Pradesh.
Tribe 19. Scaritini Bonelli, 1810
125. Scarites picicornis Dejean
Genus 52. Haplogaster Chaudoir, 1879
1879. Haplogaster Chaudoir, Ann. Soc. Ent. Belg., 22: 149.
120. Haplogaster ovata Chaudoir
1879. Haplogaster ovata Chaudoir, Ann. Soc. Ent. Belg., 22:
150.
Distribution: INDIA : Meghalaya, Uttar Pradesh
and West Bengal; Elsewhere: Nepal and
Bangladesh.
Genus 53. Oxylobus Chaudoir, 1855
1855. Oxylobus Chaudoir, Bull. Soc. Imp. Nat. Mosc., 1: 5.
1831. Scarites picicornis Dejean, Spec. Gen. Col., 5: 493.
Distribution: INDIA: Bihar, Uttar Pradesh and
West Bengal; Elsewhere: Angola, Bangladesh,
Ethiopea, Mozambique, River Niger, Senegal, Sri
Lanka, South Africa, Sudan and Tanzania.
126. Scarites planus Bonelli
1813. Scarites planus Bonelli, Mem. Acad. Sci. Turin., 20:
470.
Material examined: 2 exs., Chitrakoot, Pahari,
26.xi.2010, Coll. O. P. Srivastava & party, Ex.
418
Fauna of Uttar Pradesh, State Fauna Series, 22
Crevices in wheat Field soil; 4 exs., Jalaun, Orai,
Ajnari, 17.x.2011, Alt. 89m., Coll. B. Lal & party,
Ex. Leaf Litter; 1ex., Hardoi, Sandi Bird Sanctuary,
21.ii.2011, Coll. V. D. Hegde & party, Ex. Under
cow dung.
Distribution: INDIA : Uttar Pradesh; Elsewhere:
Afghanistan, Egypt, Ethiopia, Iraq, Mozambique,
Saudi Arabia, Sudan and Syria.
127. Scarites punctum Wiedeman
1823. Scarites punctum Wiedeman, Zool. Mag., 2: 1,38.
Material examined: 1 ex., Sultanpur, Sultanpur,
15.x.2006, Coll. P. Mukhopadhyay and Party, Ex.
Bank of Gomati River.
Distribution: INDIA: Assam, Bihar, Himachal
Pradesh, Punjab, Uttarakhand, Uttar Pradesh and
West Bengal; Elsewhere: Nepal
128. Scarites puncticollis Chaudoir
1855. Taeniolobus puncticollis Chaudoir, Bull. Soc. Imp. Nat.
Mosc., 1: 47.
1929. Scarites puncticollis, Andrewes, Faun. Brit. India,
Coleoptera, Carabidae, 1: 234, 287.
Material examined: 1 ex., Hardoi, Sandi Bird
Sanctuary, 21.ii.2011, Coll. V. D. Hegde & party,
Ex. Cow dung; 1 ex., Hardoi, Neer, Bhais ka Nala,
20.ii.2011, Coll. V. D. Hegde & party, Ex. Wet
Land; 1 ex., Faizabad, Guptar Ghat, 09.iii.2010,
Coll. V. D. Hegde & party, Ex. Hand picking; 4
exs., Barabanki, Mohammadpur, 20.iii.2012, N 26°
56´45´´ E 81°10´25´´, Coll. A. Sar & Party; 1 ex.,
Lakhimpur Kheri, Salukapur, Dudhwa National
Park, GPS N28° 25´5´´ E80° 43´ 9´´, 02.iii.2013,
Coll. V. D. Hegde & party (Reg. no. 20051/H4A);
2 exs., Lakhimpur Kheri, Sathiyana, Dudhwa
National Park, GPS N28° 32´6´´ E80° 32´30´´,
01.iii.2013, Coll. V. D. Hegde & party.
Uttar Pradesh, Tamil Nadu and West Bengal;
Elsewhere: Bangladesh, Myanmar and Sri Lanka.
130. Scarites semirugosus Chaudoir*
1855. Scarites semirugosus Chaudoir, Bull. Soc. Imp. Nat.
Mosc., 1: 90.
Material examined: 1 ex., Hardoi, Sandi Bird
Sanctuary, 21.ii.2011, Coll. V. D. Hegde & party,
Ex. Soil Crevices of Ploughed field.
Distribution: INDIA: Karnataka, Maharashtra,
Meghalaya, Tamil Nadu, Uttar Pradesh and West
Bengal; Elsewhere: Myanmar, Malaysia,
Philippines and Thailand.
131. Scarites striaticeps Chaudoir*
1880. Scarites striaticeps Chaudoir, Ann. Soc. Ent. Belg., 23:
52.
Material examined: 1 ex., Lakhimpur Kheri,
Dudhwa FRH, 24.ix.1974, Coll. P. K. Maiti and
party, Ex. At light on 7.30 PM.
Distribution: INDIA: Assam, Jharkhand and
Uttar Pradesh; Elsewhere: Bangladesh and
Myanmar.
132. Scarites subnitens Chaudoir
1855. Scarites subnitens Chaudoir, Bull. Soc. Imp. Nat.
Mosc., 1: 87.
Material examined: 1 ex., Saharanpur, Rajaji
Sanctuary, Saharanpur Road, 11.vi.1964, Coll. T.
D. Soota, Ex. Under Stone; 1 ex., Mirzapur, Chunar,
8.ii.2009, Coll. V. D. Hegde & party, Ex. Ganga
River Bed.
Distribution: INDIA : Bihar, Uttarakhand, Uttar
Pradesh and West Bengal; Elsewhere: Myanmar.
133. Scarites sulcatus Olivier*
1795. Scarites sulcatus Olivier, Ent. iii, 36: 7. t. 1. f. 11.
Distribution: INDIA : Bihar, Gujarat, Karnataka,
Kerala, Madhya Pradesh, Maharashtra, Odisha,
Pondicherry, Tamil Nadu, Uttar Pradesh and West
Bengal; Elsewhere: Bangladesh, Nepal and Sri
Lanka.
129. Scarites selene Schmidt-Goebel
1846. Scarites selene Schmidt-Goebel, Faun. Col. Birm.: 94.
Distribution: INDIA: Assam, Bihar, Meghalaya,
Material examined: 1 ex., Lakhimpur Kheri,
Dudhwa National Park, 27.ii.2011, Coll. V. D.
Hegde & party, Ex. Under log; 1 ex., Shahjahanpur,
Shahjahanpur, 27.x.2007, Coll. P. Mukhopadhyay
& party, Ex. Horse dung.
Distribution: INDIA: Arunachal Pradesh,
Assam, Himachal Pradesh, Uttarakhand, Uttar
Pradesh and West Bengal; Elsewhere: Cambodia,
KUSHWAHA and HEGDE : Insecta : Coleoptera : Carabidae
419
China, Indonesia, Korea, Laos, Myanmar, Nepal,
Pakistan and Vietnam.
Distribution: INDIA: including Gujarat and
Uttar Pradesh; Elsewhere: Myanmar, and Thailand.
134. Scarites semicircularis Mcleay**
138. Clivina striata Putzeys*
1825. Scarites semicircularis Mcleay, Ann. Jav.: 24.
1846. Clivina striata Putzeys, Mem. Liege., 2: 592.
Material examined: 1 ex., Lakhimpur Kheri,
Forest along Saheli river bank, 23.ix.1974, Alt.
250m, Coll. P. K. Maiti & party.
Material examined: 1 ex., Ghazipur, Social
Forest Nursery, 24.ix.2010, Coll. V. D. Hegde &
party, Ex. Under leaf litter; 2 exs., Allahabad, Meja,
Near FRH, 21.viii.2007, Coll. V. D. Hegde & party,
Ex. Under Leaf Litter.
Distribution: INDIA : Uttar Pradesh; Elsewhere:
Cambodia, Indonesia, Laos, Malaysia, Myanmar,
Singapore, Thailand and Vietnam.
Tribe 20. Clivinini Rafinasque, 1815
Genus 55. Clivina Latreille, 1802
1802. Clivina Latreille, Hist. nat. Crust. et Ins., 3: 96.
135. Clivina attenuata Herbst
1806. Scarites attenuata Herbst, Nat. Syst. Ins. Kaf., 10: 264,
t. 176, f.7.
1929. Clivina attenuata Andrewes, Faun. Brit. India,
Coleoptera, Carabidae, 1: 353, 355, f 51.
Material examined: 12 exs., Mau, Sheetla
Mandir Campus, 22.ix.2010, Coll. V. D. Hegde &
party, Ex. Under bricks (Reg. No. 19550-19561/
H4A); 2 exs., Mau Social Forestry, Vandevi,
21.ix.2010, Coll. V. D. Hegde & party, Ex. Under
bricks.
Distribution: INDIA : Throughout Northern and
Central India including Uttar Pradesh; Elsewhere:
Cambodia, Indonesia, Iran, Laos, Myanmar and
Vietnam.
136. Clivina gamma Andrewes*
1929. Clivina gamma Andrewes, Faun. Brit. Ind.,
Coleoptera, Carabidae, 1: 368-369.
Material examined: 3 exs., Siddharth nagar,
13.iii.2010, Coll. V. D. Hegde & party, Ex.
Agricultural Field.
Distribution: INDIA : including Uttar Pradesh.
137. Clivina lobata Bonelli
Distribution: INDIA: Bihar, Jharkhand,
Southern India and Uttar Pradesh; Elsewhere: Sri
Lanka.
139. Clivina tranquebarica Bonelli
1813. Clivina tranquebarica Bonelli, Mem. Acad. Sci. Turin.,
20: 484.
Material examined: 1 ex., Hardoi, Neer, Bhains
ka nala, 20.ii.2011, Coll. V. D. Hegde & party, Ex.
Wet Land; 2 exs., Sitapur, Tendua Jheel, 24.ii.2011,
Coll. V. D. Hegde & party, Ex. Wet Land; 5exs.,
Lalitpur, Govind Sagar, 13.x.2011, Alt. 306m., Coll.
Coll. B. Lal & party, Ex. Under stone near water
stream; 3 exs., Lalitpur, Jailwara, 14.x.2011, Alt.
296m., Coll. B. Lal & party, Ex. Near water stream;
1 ex., Mirzapur, Chunar, 8.ii.2009, Coll. V. D.
Hegde & party, Ex. Ganga River Bed; 4 exs.,
Deoria, by the side of Jugantar Chetana High
School, 30.vi.1989, Coll. H. C. Ghosh & party; 1
ex., Siddharth Nagar, Siddharth Nagar, 13.iii.2010,
Coll. V. D. Hegde & party, Ex. Agricultural Field;
1 ex., Ballia, Ballia, 13.ii.2009, Coll. V. D. Hegde
& party, Bank of Ganga River.
Distribution: INDIA: including Uttar Pradesh;
Elsewhere: Cambodia, Laos, Malaysia, Myanmar,
Sri Lanka and Vietnam.
Tribe 21. Dyschiriini Kolbe, 1880
Genus 56. Dyschirius Panzer, 1813
1813. Dyschirius Panzer, Index Ent.: 67.
1813. Clivina lobata Bonelli, Mem. Acad. Sci. Turin., 20:
481.
140. Dyschirius hingstoni Andrewes
1892. Clivina laeviceps Bates, Ann. Mus. Civ. Sto. Nat. Gen.,
32: 277.
1929. Dyschirius hingstoni Andrewes, Faun. Brit. India,
Coleoptera, Carabidae, 1: 392, 400.
Material examined: 1 ex., Allahabad, Ramnagar,
22.x.1986, Coll. A. Bal & party.
Material examined: 1 ex., Auraiya, Jamui,
19.x.2011, Coll. B. Lal & party, Ex. Dhan Paddy.
420
Fauna of Uttar Pradesh, State Fauna Series, 22
Distribution: INDIA : Bihar, Uttar Pradesh and
West Bengal.
Distribution: INDIA: Maharashtra and Uttar
Pradesh; Elsewhere: Sri Lanka.
141. Dyschirius mitodes Andrewes
147. Dyschirius bengalensis Andrewes*
1929. Dyschirius mitodes Andrewes, Faun. Brit. India,
Coleoptera, Carabidae, 1: 394, 409.
1929. Dyschirius bengalensis Andrewes, Faun. Brit. India,
Coleoptera, Carabidae, 1: 397-398.
Distribution: INDIA (Uttar Pradesh).
142. Dyschirius perfectus Andrewes*
1929. Dyschirius perfectus Andrewes, Faun. Brit. India,
Coleoptera, Carabidae, 1: 392, 399.
Material examined: 1 ex., Hardoi, Neer, Bhais
Ka Nala, 20.ii.2011, Coll. V. D. Hegde & party,
Ex. Wet Land; 1 ex., Jhansi, Ambabai, Alt. 189 m.,
12.x.2011, Coll. B. Lal & party (Reg. no. 20046/
H4A).
Distribution: INDIA : Maharashtra, Uttarakhand
and Uttar Pradesh.
143. Dyschirius variabilis Andrewes
1929. Dyschirius variabilis Andrewes, Faun. Brit. India,
Coleoptera, Carabidae, 1: 393, 405.
Distribution: INDIA: Uttarakhand, Uttar
Pradesh and West Bengal; Elsewhere: Bangladesg.
144. Dyschirius verticalis Putzeys
1878. Dyschirius verticalis Putzeys, Compt. Rend soc. Ent.
Belg.: 172.
Distribution: INDIA : Uttar Pradesh.
145. Dyschirius mahratta Andrewes*
1929. Dyschirius mahratta Andrewes, Faun. Brit. India,
Coleoptera, Carabidae, 1: 396.
Material examined: 1 ex., Lakhimpur Kheri,
Salukapur, Dudhwa National Park, GPS N28° 25´
5´´ E80°43´9´´, 02.iii.2013, Coll. V. D. Hegde &
party (Reg. no. 20050/H4A).
Distribution: INDIA: Maharashtra and Uttar
Pradesh; Elsewhere: Sri Lanka.
146. Dyschirius mahratta var. paucipunctus
Andrewes*
1929. Dyschirius mahratta var. paucipunctus Andrewes,
Faun. Brit. India, Coleoptera, Carabidae, 1: 397.
Material examined: 1 ex., Hardoi, Neer, Bhais
Ka Nala, 20.ii.2011, Coll. V. D. Hegde & party,
Ex. wet land (reg. no. 20047/H4A).
Material examined: 1 ex., Hardoi, Neer, Bhais
Ka Nala, 20.ii.2011, Coll. V. D. Hegde & party,
Ex. wet land (reg. no. 20048/H4A).
Distribution: INDIA: Uttar Pradesh and West
Bengal.
Subfamily 13. MELAENINAE Allaud, 1934
Tribe 22. Cymbionotini Andrewes, 1933
Genus 57. Cymbionotum Baudi, 1864
1864. Cymbionotum Baudi, Berl. Ent. Zeit.: 211.
148. Cymbionotum fascigerum (Chaudoir)
1852. Coscinia fasciger Chaudoir, Bull. Soc. Imp. Nat. Mosc.,
1: 92.
1935. Cymbionotum fascigerum, Andrewes, Faun. Brit.
India, Coleoptera, Carabidae, 2: 25.
Distribution: INDIA: Uttar Pradesh.
Cymbionotum Sp.
Material examined: 1 ex. Lucknow, Kakor,
23.vi.1989, Coll. K. K. Ray & party.
Tribe 23. Melaenini Dejean, 1831
Genera 58. Melaenus Dejean, 1831*
1831. Melaenus Dejean, Spec. Gen. Col., 5: 481.
149. Melaenus piger (Fabricius)*
1801. Brachinus piger Fabricius, Syst. Eleuth., 1: 219.
1847. Melaenus piger Schaum, Stett. Ent. Zeit.: 49.
Material examined: 1 ex.,Ghazipur, Social
Forest Nursery, 24.ix.2010, Coll. V. D. Hegde &
party, Ex. Under leaf litter (ZSI Reg. No. 20045/
H4A) ; 1 ex., Jalaun, Orai, near Machhar Chauraha,
15.x.2011, Coll. B. Lal & party, Ex. at light; 1 ex.,
Etawah, Bakewar, 22.x.2011, Coll. B. Lal & party.
Distribution: INDIA: Andhra Pradesh,
Chhattisgarh, Goa, Karnataka, Madhya Pradesh,
Maharashtra, North India, Odisha, Puducherry,
Tamil Nadu, Uttar Pradesh and West Bengal;
Elsewhere: Sri Lanka.
KUSHWAHA and HEGDE : Insecta : Coleoptera : Carabidae
Subfamily 14. DRYPTINAE Bonelli, 1810
Tribe 24. Dryptini Bonelli, 1810
Genus 59. Drypta Latreille, 1796
1796. Drypta Latreille, Prec. Cara. Gen. Insectes: 75.
150. Drypta argillacea Andrewes
1924. Drypta argillacea Andrewes, MissionProv. Centr.
l’Inde Reg. Occi. l’Himalaya 1914, par guy Babault:
106, t.3, f.5.
Distribution: INDIA: Maharashtra, Odisha,
Uttarakhand and Uttar Pradesh; Elsewhere:
Bangladesh.
151. Drypta lineola Macleay
1825. Drypta lineola Macleay, Ann. Jav.: 27.
Material examined: INDIA: Uttar Pradesh, 1 ex.,
Auraiya, near Dibiyapur , 19.x.2011, Coll. B. Lal
& Party; 1 ex., Hardoi, Sandi Bird Sanctuary,
21.ii.2011, Coll. V. D. Hegde & Party, Ex. Under
Cow dung; 1 ex., Lakhimpur Kheri, Dudhwa Tiger
Reserve, 26.ii.2011, Coll. V.D. Hegde & party, Ex.
Dusting.
Distribution: INDIA: Himachal Pradesh,
Maharashtra, and Uttar Pradesh; Elsewhere:
throughout South East Asia.
421
Khannaut River Bank, 01.iii.2011, Coll. V. D.
Hegde & party, Ex. Dusting; 2 exs., Shahjahanpur,
Khannaut River Bank, 01.iii.2011, Coll. V. D.
Hegde & party, Ex. In soil crevices; 1 ex., Sitapur,
Tendua Jheel, 24.ii.2011, Coll. V. D. Hegde &
party, Ex. in leaf node of sugarcane; 1 ex.,
Lakhimpur Kheri, Dudhwa National Park,
Belrayan, GPS N28°23´14´´ E80°54´40.6´´,
28.ii.2013, Coll. V. D. Hegde & party; 1 ex.,
Lakhimpur Kheri, Dudhwa National Park,
Sathiyana, GPS N28°32´6´´ E80°32´30´´,
01.iii.2013, Coll. V. D. Hegde & party.
Distribution: INDIA: Bihar, Uttarakhand and
Uttar Pradesh.
Subfamily 16. SIAGONINAE Bonelli, 1813
Tribe 27. Siagonini Bonelli, 1813
Genus 62. Luperca Castelnau, 1840
1840. Luperca Castelnau, Hist. Nat. Ins. Col., 1: 63.
154. Luperca laevigata (Fabricius)
1781. Carabus laevigata Fabricius, Spec. Ins., 1: 304.
1929. Luperca laevigata, Andrewes, Faun. Brit.
India,Coleoptera, Carabidae, 1: 206, f.35.
Tribe 25. Galeritini Leconte, 1853
Distribution: INDIA: Bihar, Karnataka, Madhya
Pradesh, Maharashtra, Odisha, Tamil Nadu and
Uttar Pradesh.
Genus 60. Galerita Fabricius, 1801
Genus 63. Siagona Latreille, 1804
1801. Galerita Fabricius, Syst. Eleuth., 1: 214.
152. Galerita indica Chaudoir
1861. Galerita indica Chaudoir, Bull. Soc. Imp. Nat. Mosc.,
1: 557.
Distribution: North India.
Subfamily 15. CTENODACTYLINAE
Castelnau, 1834
Tribe 26. Hexagonini Horn, 1881
Genus 61. Hexagonia Kirby, 1825*
1825. Hexagonia Kirby, Trans. Linn. Soc. Lond. 14: 563.
153. Hexagonia uninotata Andrewes*
1935. Hexagonia uninotata Andrewes, Fauna Brit. India,
Carabidae, 2, Harapalini 1: 14, Pl. 2, f. 3.
Material examined: 2exs., Shahjahanpur,
1804. Siagona Latreille, Nouv. Dict. Hist. Nat. 24. Tab.
Meth.: 141.
155. Siagona atrata Dejean*
1825. Siagona atrata Dejean, Spec. Gen. Col., 1: 360.
Material examined: 1 exs., Hardoi, Sandi Bird
Sanctuary, 21.ii.2011, Coll. V. D. Hegde & party,
Ex. crevices in soil (Z.S.I. Reg. No. 19548/H4A);
1 exs., Hardoi, Sandi Bird Sanctuary, 21.ii.2011,
Coll. V. D. Hegde & party, Ex. Soil crevices of
Ploughed field; 1 ex., Lakhimpur Kheri, Dudhwa
National park, Chandan Chauki, GPS N28°32´1´´
E80°46´13´´, 27.ii.2013, Coll. V. D. Hegde & party,
Ex. under bark.
Distribution: INDIA : Assam, Bihar, Jharkhand,
Odisha, Uttar Pradesh and West Bengal; Elsewhere:
Bangladesh.
422
Fauna of Uttar Pradesh, State Fauna Series, 22
156. Siagona baconi Chaudoir*
1876. Siagona baconi Chaudoir, Bull. Mosc., 1: 89.
Material examined: 1 ex., Ballia, FRH,
15.ix.2010, Coll. V. D. Hegde & party, (Reg. no.
19549/H4A); 1 ex., Hardoi, Sandi Bird Sanctuary,
21.ii.2011, Coll. V. D. Hegde & party, Ex. Crevices
in soil; 1 ex., Hardoi, Sandi Bird Sanctuary,
21.ii.2011, Coll. V. D. Hegde & party, Ex. under
cow dung; 1 ex., Jalaun, Orai, Machhar Chauraha,
17.x.2011, Coll. B. Lal & party, Ex. at light; 1 ex.,
Lakhimpur Kheri, Dudhwa National park, Chandan
Chauki, GPS N28°32´1´´ E80°46´13´´, 27.ii.2013,
Coll. V. D. Hegde & party, Ex. under bark.
Distribution: INDIA: Bihar, Jharkhand, Madhya
Pradesh, Odisha, Uttar Pradesh and West Bengal;
Elsewhere: Bangladesh and Myanmar.
157. Siagona fabricii Andrewes
1921. Siagona fabricii Andrewes, Trans. Ent. Soc. Lond.:
158.
Material examined: 1 ex., Auraiya, near Bus
station, 18.x.2011, Coll. B. Lal & party, Ex. at light;
4 exs., Jalaun, Orai, Machhar Chauraha, 17.x.2011,
Coll. B. Lal & party, Ex. at light; 17 exs., Jalaun,
Orai, near Machhar Chauraha, 17.x.2011, Coll. B.
Lal & party, Ex. at light (Reg. No. 19562-19578/
H4A); 2 exs., Jalaun, Orai, Vohadpura, 16.x.2011,
Alt. 82m., Coll. B. Lal & party; 2exs., Jalaun, Orai,
Ajnari, 17.x.2011, Alt. 89m., Coll. B. Lal & party,
Ex. Under leaf litter; 12 exs., Jalaun, Chili near Orai,
15.x.2011, Coll. B. Lal & party, Ex. at light; 1 ex.,
Allahabad, Meja, near FRH, 21.viii.2007, Coll. V.
D. Hegde & party, Ex. Under leaf litter; 20 exs.,
Lalitpur, near Railway Station, 14.x.2011, Coll. B.
Lal & party, Ex. at light; 10 exs., Chitrakoot, Pahari,
26.xi.2010, Coll. O. P. Srivastava & party, Ex.
Crevices in wheat field soil; 01 ex.,Ghazipur, Social
Forest Nursery, 24.ix.2010, Coll. V. D. Hegde &
party, Ex. Under leaf litter; 4 exs., Ballia, Jamui,
17.ix.2010, Coll. V. D. Hegde & party; 1 ex., Ballia,
Bharatpura, 16.ix.2010, Coll. V. D. Hegde & party,
Ex. Under stone; 9 exs., Ballia, FRH, 15.ix.2010,
Coll. V. D. Hegde & party; 1 ex., Lakhimpur Kheri,
Dudhwa National Park, Gauri Phanta, GPS N28°
38´53´´ E80°34´12´´, 26.ii.2013, Coll. V. D. Hegde
& party.
Distribution: INDIA: throughout India including
Uttar Pradesh; Elsewhere: Sri Lanka.
158. Siagona flesus (Fabricius)
1801. Galerita flesus Fabricius, Syst. Eleuth., 1: 216.
1929. Siagona flesus, Andrewes, Faun. Brit. India,
Coleoptera, Carabidae 1: 180, 182, 196. f. 32 (13), f.
34.
Material examined: 1ex., Allahabad, Meja, near
FRH, 21.viii.2007, Coll. V. D. Hegde & party; 1
ex., Ballia, FRH, 15.ix.2010, Coll. V. D. Hegde &
party; 1 ex., Hardoi, Sandi Bird Sanctuary,
21.ii.2011, Coll. V. D. Hegde & party, Ex. Crevices
in soil; 1 ex., Jalaun, Chili near Orai, 15.x.2011,
Coll. B. Lal & party, Ex. at light; 1 ex., Jalaun, Orai,
Vohadpura, 16.x.2011, Alt. 82m., Coll. B. Lal &
party; 1 ex., Lalitpur, near Railway Station,
14.x.2011, Coll. B. Lal & party, Ex. at light; 1 ex.,
Jalaun, Orai, near Machhar Chauraha, 15.x.2011,
Coll. B. Lal & party, Ex. at light, (Reg. no. 19580/
H4A).
Distribution: INDIA: throughout India including
Uttar Pradesh; Elsewhere: Cambodia, Laos,
Myanmar, Malaysia, Senegal, Tanzania, Thailand
and Vietnam.
159. Siagona pumila Andrewes*
1921. Siagona pumilus Andrewes, Trans. Ent. Soc. Lond.:
164.
1929. Siagona pumila, Andrewes, Fauna Brit. India,
Coleoptera Carabidae, 1: 180, 199, f. 32(14), Pl. VI
Fig. 2.
Material examined: 2 exs., Lalitpur, near
Railway Station, 14.x.2011, Coll. B. Lal & party,
Ex. at light; 1 ex., Deoria, Hanuman Temple,
26.vi.1989, Coll. H.C. Ghosh & party, Ex. Near
pond; 1 ex., Ballia, Jamui, 17.ix.2010, Coll. V. D.
Hegde & party, (Reg. no. 19547/H4A); 1 ex.,
Hardoi, Sandi Bird Sanctuary, 21.ii.2011, Coll. V.
D. Hegde & party, Ex. Under Cow dung; 2 exs.,
Hardoi, Sandi Bird Sanctuary, 21.ii.2011, Coll. V.
D. Hegde & party, Ex. Crevices in Soil; 1 ex.,
Auraiya, Bamuripur, 6.ix.2009, Coll. V. D. Hegde
& party, Ex. Under dead leaves (Reg. no. 19579/
H4A); 1 ex., Lakhimpur Kheri, Dudhwa National
Park, Gauri Phanta, GPS N28°38´53´´ E80° 34´
12´´, 26.ii.2013, Coll. V. D. Hegde & party.
KUSHWAHA and HEGDE : Insecta : Coleoptera : Carabidae
423
Distribution: INDIA: Bihar, Karnataka,
Maharashtra, Tamil Nadu, Uttarakhand, Uttar
Pradesh and West Bengal; Elsewhere: Bangladesh.
Under dry Sugarcane leaves; 1 ex., Saharanpur,
Ghogregi vill., 31.x.2009, Coll. O. P. Srivastava &
party.
160. Siagona dilutipes var. aligera Andrewes*
Distribution: INDIA: throughout northern and
central India including Manipur, Nilgiri Hills and
Uttar Pradesh; Elsewhere: China, Laos, Myanmar
and Vietnam.
1929. Siagona dilutipes var. aligera Andrewes, Fauna Brit.
India, Coleoptera Carabidae, 1: 203.
Material examined: 1 ex., Jalaun, Chili near
Orai, 15.x.2011, Coll. B. Lal & party, Ex. at light.
Distribution: INDIA: Maharashtra and Uttar
Pradesh.
Subfamily 17. PLATYNINAE Bonelli, 1810
Subfamily 18. NEBRIINAE Laporte, 1834
Tribe 29. Nebriini Laporte, 1834
Genus 60. Nebria Latreille, 1802
1802. Nebria Latreille, Hist. Nat. Crust. et Ins.: 89.
Tribe 28. Platynini Bonelli, 1810
164. Nebria xanthacra Chaudoir
Genus 64. Megalonychus Chaudoir, 1843
1850. Nebria xanthacra Chaudoir, Bull. Soc. Imp. Nat.
Mosc., 1: 423.
1843. Megalonychus Chaudoir, Bull. Soc. Imp. Nat. Mosc.,
2: 418.
161. Megalonychus birmanicus Bates
1892. Megalonychus birmanicus Bates, Ann. Mus. Civ. Sto.
Nat. Gen. 32: 369.
Distribution: INDIA: Arunachal Pradesh,
Assam, Bihar, Meghalaya, Nagaland, Uttarakhand,
Uttar Pradesh and West Bengal; Elsewhere: Laos
and Myanmar.
Genus 65. Onycholabis Bates, 1873*
1873. Onycholabis Bates, Trans. Ent. Soc. Lond.: 329.
162. Onycholabis acutangulus Andrewes*
1923. Onycholabis acutangulus Andrewes, Ann. Mag. Nat.
Hist., (9)12: 682.
Material examined: 1 ex., Saharanpur, Rajaji
Sanctuary, Saharanpur Road, 27.xi.1964, Coll. T.
D. Soota and party, Ex. under stone at bank of
stream.
Distribution: INDIA: Arunachal Pradesh,
Uttarakhand and Uttar Pradesh.
Genus 66. Orthotrichus Peyron, 1856
1856. Orthotrichus Peyron, Ann. Soc. Ent. Fr., 4 (3): 717.
163. Orthotrichus indicus Bates
1891. Orthotrichus indicus Bates, Compt. Rend soc. Ent.
Belg.: 334.
Material examined: 01 ex., Meerut, Meerut,
07.xi.2007, Coll. P. Mukhopadhyay and Party, Ex.
Material examined: 7exs., Saharanpur,
Beharigarh, 17.vi.1971, Coll. J. C. Tripathi and
party.
Distribution: INDIA: Himachal Pradesh,
Kashmir, Uttarakhand and Uttar Pradesh.
165. Nebria xanthacra var. nigricollis
Andrewes*
1925. Nebria xanthacra var. nigricollis Andrewes, Ann.
Mag. nat. Hist., (9)15: 312.
Material examined: 8 exs., Saharanpur,
Beharigarh, 17.vi.1971, Coll. J. C. Tripathi & party;
2 exs., Saharanpur, Beharigarh, 17.vi.1971, Coll.
J. C. Tripathi & party, (Reg. no. 5364-5365/H4A).
Distribution: INDIA: Himachal Pradesh,
Uttarakhand and Uttar Pradesh.
Tribe 30. Notiophilini Motchulsky, 1850
Genus 68. Notiophilus Dumeril, 1806
1806. Notiophilus Dumeril, Zool. Analyt.: 194.
166. Notiophilus orientalis Chaudoir
1850. Notiophilus orientalis Chaudoir, Bull. Soc. Imp. Nat.
Mosc., 1: 428.
Distribution: INDIA: Himachal Pradesh,
Kashmir, Uttarakhand and Uttar Pradesh;
Elsewhere: Pakistan.
Subfamily 19. OMOPHRONINAE Bonelli, 1810
Tribe 31. Omophronini Bonelli, 1810
424
Fauna of Uttar Pradesh, State Fauna Series, 22
Genus 69. Omophron Latreille, 1802
1802. Omophron Latreille, Hist. Nat. Crust. et Ins., 3: 89.
167. Omophron maculosus Chaudoir
1850. Omophron maculosus Chaudoir, Bull. Soc. Imp. Nat.
Mosc., 1: 424.
Distribution: INDIA : Bihar and Uttar Pradesh;
Elsewhere: Bangladesh.
Subfamily 20. PAUSSINAE Latreille, 1807
Tribe 32. Paussini Latreille, 1807
Genus 70. Merismoderus Westwood, 1847
1847. Ceratoderus (Merismoderus) Westwood, Trans. Ent.
Soc. Lond., 5: 23.
1854. Merismoderus Lacordaire, Gen Col., 2: 11.
168. Merismoderus bensoni Westwood
1847. Merismoderus bensoni Westwood, Trans. Ent. Soc.
Lond., 5: 23.
Distribution: INDIA : Uttar Pradesh.
Genus 71. Platyrhopalus Westwood, 1838
1838. Platyrhopalus Westwood, Trans. Linn. Soc. Lond., 16:
654.
169. Platyrhopalus denticornis Donovan
1800. Paussus denticornis Donovan, Ins Ind., 1: 8, pl. 5,
fig. 1.
Material examined: 2 exs., Ballia, Bharatpur,
16.ix.2010, Coll. V. D. Hegde & party, Ex. Under
bark.
Distribution: INDIA: Assam, Bihar, Delhi,
Madhya Pradesh, Maharashtra, Punjab, Tamil
Nadu, Uttar Pradesh and West Bengal.
170. Platyrhopalus angustus Westwood
1839. Platyrhopalus angustus Westwood, Trans. Ent. Soc.
Lond., 2: 92, pl 10, fig. 6.
Distribution: INDIA: Andhra Pradesh, Assam,
Bihar, Kashmir, Uttarakhand and Uttar Pradesh.
171. Platyrhopalus intermedius Benson
1846. Platyrhopalus intermedius Benson, Calcutta Journ.
Nat. Hist., 6: 468.
Distribution: INDIA: Uttar Pradesh.
Genus 72. Paussus Linnaeus, 1775
1775. Paussus Linnaeus, Bigae Insect: 7.
172. Paussus hearseyanus Westwood
1842. Paussus hearseyanus westwood, Proc. Linn. Soc.
Lond.: 133.
Distribution: INDIA : Gujarat, Madhya Pradesh,
Maharashtra and Uttar Pradesh.
173. Paussus soleatus Wasmann*
1894. Paussus soleatus Wasmann, Krit. Verz. Myrm. Termit
Aith.: 216.
Material examined: 1 ex., Auraiya, Fatehpur
near Muradganj, 20.x.2011, Coll. B. Lal & party,
Ex. Leaf litter.
Distribution: INDIA: Maharashtra and Uttar
Pradesh.
174. Paussus ploiophorus Benson
1846. Paussus ploiophorus Benson, Calcutta Journ. Nat.
Hist., 6: 463.
Distribution: INDIA: Uttar Pradesh.
SUMMARY
The present paper deals with 174 species of 72
genera under 32 tribes and 20 subfamilies of Family
Carabidae. During the study 551examples has been
identified under 95 species and 36 genera. A total
of 48 species and 12 genera has been recorded for
the first time from Uttar Pradesh, out of which
during study 22 species had been reported for the
first time from Uttar Pradesh by Kushwaha &
Hegde (2010, 2012 and 2013). A total 05 species
has been recorded for the first time from India out
of which during the study 02 had been reported for
the first time from India by Kushwaha & Hegde
(2013).
ACKNOWLEDGEMENTS
We are thankful to the Director, Zoological
Survey of India, Kolkata for the facilities. We are
indebted to Dr. Kailash Chandra, Addl. Director
and Head, Entomology Division ‘A’ for his valuable
suggestions in preparing the manuscript. We thank
PCCF and Wild life Warden of Uttar Pradesh state
for permission to collect the specimens and all the
forest officials for providing help during our survey.
We thank all the staffs of Coleoptera section for
their constant encouragement.
KUSHWAHA and HEGDE : Insecta : Coleoptera : Carabidae
425
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