ISSN 0869-5938, Stratigraphy and Geological Correlation, 2015, Vol. 23, No. 2, pp. 155–191. © Pleiades Publishing, Ltd., 2015.
Original Russian Text © V.V. Arkadiev, E.Yu. Baraboshkin, M.I. Bagaeva, T.N. Bogdanova, A.Yu. Guzhikov, A.G. Manikin, V.K. Piskunov, E.S. Platonov, Yu.N. Savel’eva,
A.A. Feodorova, O.V. Shurekova, 2015, published in Stratigrafiya. Geologicheskaya Korrelyatsiya, 2015, Vol. 23, No. 2, pp. 43–80.
New Data on Berriasian Biostratigraphy, Magnetostratigraphy,
and Sedimentology in the Belogorsk Area (Central Crimea)
V. V. Arkadieva, E. Yu. Baraboshkinb, M. I. Bagaevac, T. N. Bogdanovad, A. Yu. Guzhikovc,
A. G. Manikinc, V. K. Piskunovb, E. S. Platonova, Yu. N. Savel’evae,
A. A. Feodorovae, and O. V. Shurekovae
a
St. Petersburg State University, Universitetskaya nab. 7–9, St. Petersburg, 199034 Russia
e-mail: arkadievvv@mail.ru
b Moscow State University, Moscow, 119991 Russia
c Saratov State University, ul. Astrakhanskaya 83, Saratov, 410012 Russia
d All-Russian Institute of Geology, Srednii pr. 74, St. Petersburg, 199106 Russia
e Research and Production Enterprise Geologorazvedka, ul. Knipovicha 11/2, St. Petersburg, 192019 Russia
Received July 9, 2013; in final form, December 17, 2013
Abstract—The most complete composite Berriasian bio- and magnetostratigraphic section of central Crimea
is characterized for the first time with a description of the contact between the carbonate Bedenekyr and terrigenous Bechku formations. The section contains all the standard ammonite zones: jacobi, occitanica, and
boissieri. The Malbosiceras chaperi Beds are attributed to the occitanica Zone. The Berriasian section is
characterized by six foraminiferal assemblages, ostracods (Costacythere khiamii–Hechticythere belbekensis
and Costaythere drushchitzi–Reticythere marfenini beds), and dinocysts (Phobercysta neocomica Beds).
The magnetostratigraphic section contains analogs of Chrons M17 and M16 reliably correlated with ammonite zones. On the basis of paleomagnetic data, the Berriasian section of central Crimea is correlated with
coeval sections of the Mediterranean Region. The sedimenological analysis confirms accumulation of Berriasian sediments mostly in shallow shelf environments of the carbonate platform.
Keywords: Berriasian, central Crimea, ammonites, bivalves, ostracods, foraminifers, palynomophs, biostratigraphy, magnetostratigraphy, sedimentology, paleomagnetism, magnetic polarity, magnetic chrons
DOI: 10.1134/S0869593815020033
INTRODUCTION
The Berriasian sections in the central part of the
Crimean Peninsula were investigated by many
researchers (Drushchits and Yanin, 1959; Kvantaliani
and Lysenko, 1979; Bogdanova et al., 1981; Bogdanova
and Kvantaliani, 1983; etc.); their paleomagnetic
study was first conducted by V.N. Eremin in the 1980s
(Molostovskii et al., 1989). The evolution of views on
their subdivision is considered in detail in the recently
published collective monograph (Arkadiev et al., 2012).
We conducted complex bio- and magnetostratigraphic
investigations of Berriasian sections in central Crimea
in 2002 and 2011–2012. In 2002, a team of geologists
from Moscow and Saratov State Universities examined sections near the settlements of Balki and
Pasechnoe. Unfortunately, fragmentary magnetostratigraphic records were obtained only for the
Pasechnoe section. No paleomagnetic measurements
appropriate for interpretation of magnetic polarity
were obtained for the Balki section because of insufficient sensitivity of the laboratory equipment used at
that time (Yampolskaya, 2005).
The joint efforts of specialists from St. Petersburg,
Saratov, and Moscow State Universities and Research
and Production Enterprise Geologorazvedka resulted in
the complex investigation of sections in the outskirts of
the settlements of Balki, Mezhgor’e, and Novoklenovo
in 2011 and sections in the Enisarai Ravine and on the
northern slope of the Karabi-Yaila Plateau (southsouthwest of the Balki settlement) in 2012 (Fig. 1). The
field works were dedicated to the thorough description
of sections (including the contact between limestones of
the Bedenekyr Formation and sandy–clayey Bechku
Formation first discovered at two localities in the outskirts of the Balki settlement), sampling of organic
remains (ammonites, bivalves, corals) and rocks for the
study of micro- (foraminifers, ostracods, dinocysts,
calpionellids, spores, and pollen) and ichnofossils,
paleomagnetic measurements, and sedimentological
analysis. The subsequent analysis of sampled material
made it possible, first, to correlate isolated outcrops
between each other and compile the most complete
Berriasian section for central Crimea and, second, to
obtain its micropaleontological and magentostratigraphic characteristics. The lithological description of
155
156
ARKADIEV et al.
Zelenogorskoe
Burul’ch
a R.
Novoklenovo
Balki
2944
2940
2420
2943
2952
Pasechnoe
Yakovlevka
2948
2947
2951
Mezhgor’e
0
2949
2950
1
2
3
4 km
SEA OF AZOV
Crimean
Peninsula
Kerch
Evpatoriya
Feodosiya
Belogorsk
Sudak
Sevastopol
Alushta
Yalta
BLACK
SEA
Fig. 1. Location of Berriasian outcrops in the Sary-Su River basin. Numerals correspond to section numbers.
the section was accomplished by E.Yu. Baraboshkin
and V.K. Piskunov. The paleomagnetic data were
obtained by M.I. Bagaeva, A.Yu. Guzhikov, and
A.G. Manikin. Organic remains were identified by the
following specialists: ammonites by V.V. Arkadiev and
T.N. Bogdanova; bivalves by T.N. Bogdanova; brachiopods and echinoderms by S.V. Lobacheva; corals by
I.Yu. Bugrova; crinoids by V.G. Klikushin; ostracods by
Yu.N. Savel’eva; dinocysts, spores, and pollen by
O.V. Shurekova; foraminifers by A.A. Feodorova;
ichnofossils by E.Yu. Baraboshkin. E.S. Platonov made
an attempt to find calpionellids in thin sections (200 in
total), but failed. The ammonite specimen, foraminifers, and ostracods illustrated in this work (collection
no. 13244) are stored at the Central Research Geological Museum (St. Petersburg); the collection of palynomorphs (no. 13220) is stored at the same place.
STRUCTURE OF THE SUCCESSION
In the Sary-Su River basin (Mezhgor’e–Balki–
Novoklenovo settlement area) (Fig. 1), the Berriasian
Stage includes (from the base upward) limestones and
clayey limestones of the upper Bedenekyr Formation,
siltstones and sandstones of the Bechku Formation, and
sponge horizon, clays, siltstones, and biohermal limestones of the Kuchki Formation (Arkadiev, 2007).
According to (Bogdanova et al., 1981), limestones of
the Bedenekyr Formation cropping out south of the
Balki Settlement on the Karabi-Yaila Plateau contain
ammonites Pseudosubplanites ponticus (Ret.) and Berriasella jacobi (Maz.) of the jacobi zone. Unfortunately,
more exact localization of these ammonite forms
remains unknown. It is conceivable that E.Yu. Baraboshkin investigated the same section or its analogs on
the northwestern slope of the Karabi-Yaila Plateau in
the vicinity of the military camp in 1996–1997. In this
area, white and pinkish upper Tithonian bioclastic
limestones with Anchispirocyclina lusitanica (Egger) are
overlain by white limestones at least 50–60 m thick with
ammonites Berriasella sp. and Protetragonites tauricus
(Kulj.-Vor.), brachiopods Loriolithyris sp., bivalves Gervilella sp. and Pinna sp., and gastropods.
The Bechku Formation in central Crimea is characterized by ammonites Dalmasiceras tauricum (Bogd.
et Ark.), Malbosiceras chaperi (Pict.), M. malbosi
(Pict.), M. pictetiforme Tav., Neocosmoceras euthymi
(Pict.), N. minutus Ark. et Bogd., Hegaratia bidi-
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NEW DATA ON BERRIASIAN BIOSTRATIGRAPHY, MAGNETOSTRATIGRAPHY
chotoma Bogd. et. Kvant., Fauriella simplicicostata
(Maz,), F. boissieri (Pict.), and others.1 On the basis of
this ammonite assemblage, the Bechku Formation was
correlated with the jacobi (Malbosiceras chaperi
Beds), occitanica, and boissieri zones. No guide
ammonite species were found in the Kuchki Formation developed in the Sary-Su River basin. The sponge
horizon contains abundant brachiopod remains
belonging to Symhythiris arguinensis (Moiss.), which
is characteristic of the synonymous beds. Ammonites
in the horizon are represented only by Hegaratia sp.
and Spiticeras sp. In 2002, E.Yu. Baraboshkin found in
its basal layer Riasanites crassicostatum (Kyant. et Lys.)
together with Loriolithyris valdensis (Lor.) and Symphythiris arguinensis (Miss.). This find allows at least
the base of the sponge horizon to be attributed to the
crassicostatum Subzone. The siltstone member overlying the sponge horizon near the settlement of Mezhgor’e yielded rare poorly preserved ammonites: Haploceras ex gr. cristifer (Opp.), Protetragonites tauricus
(Kulj.-Vor.), Spiticeras sp., and Subalpinites sp. (identifications by T.N. Bogdanova). These species of the
genera Haploceras, Protetragonites, and Spiticeras
occur through the entire Berriasian section of Crimea.
In France, representatives of the genus Subalpinites
are known from all the Berriasian zones (Le Hégarat,
1973). In Crimea, the Sualpinites taxa are described by
V.V. Arkadiev (Arkadiev et al., 2012) from the occitanica Zone in the outskirts of the Balki settlement. Nevertheless, the stratigraphic position of beds and their
ammonites are consistent also with their attribution to
the boissieri Zone. The biohermal limestones occurring in the upper part of the section are barren of
ammonite remains. Therefore, in the previously proposed stratigraphic scale (Arkadiev et al., 2012), they
are conditionally attributed to the Berriasian. At the
same time, finds of brachiopods Symphythiris kojnautensis (Moiss.),Weberithyris moissevi (Weber), Zeillerina baksanensis Smirn., bivalves Megadiceras koinautense Pchel., and others in this sequence have been
known for a long time (Yanin and Smirnova, 1981). The
correlation of these limestones with the Bel’bek River
section, where similar facies and faunal assemblages
occur below the undoubtedly lower Valanginan layers,
allows this sequence to be attributed to the upper Berriasian Megadiceras koinautense Beds (Yanin and Baraboshkin, 2000).
The upper surface of limestones is eroded and
affected by karst processes; the rocks are crossed by
deep (over 6 m) cracks filled with quartz sandstone.
The change in the sedimentation patterns in southwestern Crimea is typical of the Berriasian–Valanginian transition.
The total thickness of the Berriasian section in the
Sary-Su River basin is approximately 600 m (taking into
consideration gaps in observations). Unfortunately, the
1 E.Yu.
Baraboshkin believes that species euthymi and minutus
should be attributed to the genus Euthymiceras, and species bidichotoma and nerodenkoi, to the genus Balkites.
STRATIGRAPHY AND GEOLOGICAL CORRELATION
157
section does not represent a continuous succession and
is investigated in isolated outcrops correlated on the
basis of faunal finds. In the examined sections, the layers dip at an angle of 10°–12° in the NNE direction.
Below, we describe the lithological composition of
fragments constituting the composite section, some of
which are well known (outcrops 2420, 2940, 2943,
2944, 2947), while others were visited for the first time
(outcrops 2948–2952). All these outcrops are located in
the outskirts of the Balki, Mezhgor’e, and Novoklenovo
settlements (Fig. 2).
Section 2950 (44°58′36.40′′ N,
34°28′45.90′′ E; Fig. 3)
M e m b e r 1 (Samples 2950/1–5). Yellowish gray
bedded bioclastic wacke- to, less commonly, packstones with abundant thalassinoid burrows. Limestones contain ooids, bioclasts of brachiopod and
bivalve shells, skeletal detritus, and single foraminiferal tests of the Everticyclammina virguliana–Retrocyclammina recta–Bramkampella arabica Assemblage. The thickness is 4.5 m.
Section 2951 (44°58′41.40′′ N,
34′28°44.50′′ E; Fig. 3)
M e m b e r 2 (Samples 2951/1–7). Wacke- and
packstones similar to rocks constituting Member 1.
The talus yielded ammonite Malbosiceras ex gr. malbosi (Pictet). The thin sections demonstrate abundant
sections of foraminifers belonging to the Everticyclammina virguliana–Retrocyclammina recta–Bramkampella arabica Assemblage. The thickness is 12 m.
Section 2947 (Enisarai; 44°58′54.80′′ N,
34°28′18.00′′ E; Fig. 3)
The lower part of the section is similar to Member 2
(Samples 2947/1–9). These sediments are overlain by
the following units:
M e m b e r 3 (Samples 2951/9–40). Gray bedded
packstones and, less commonly, wackestones (layers
10–50 cm thick) with abundant Thalassinoides burrows, skeletal detritus up to 2 mm across, and rare larger
bioclasts. The boundaries between layers are wavy. The
layers contain abundant ferrugenous ooids and substantially rarer marcasite concretions. The taluses from the
lower and upper parts of the member yielded bivalves
Prohinnites renevieri (Coq.) and Tortarctica weberae
Mordv., respectively. The thin sections exhibit abundant
sections of foraminifers of the Everticyclammina virguliana–Retrocyclammina recta–Bramkampella arabica
Assemblage. The thickness is 63 m.
Section 2949 (44°58′56.10′′ N,
34°28′59.10′′ E; Fig. 3)
This section comprises five members (from the base
to the top):
M e m b e r 4 (Samples 2949/1–3). Thick-bedded
(30–40 cm) packstones with ferruginous ooids, bioVol. 23
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2015
2943
Beds with
Zeillerina
baksanensis
Beds with
Malbosiceras chaperi
2948
2952
occitanica
2940
2949
2947
2950
jacobi
2951
Polarity
Riasanites crassicostatum,
Riasanites sp., Hegaratia balkensis,
H. taurica, H. bidichotoma,
H. nerodenkoi,
Fauriella simplicicostata
Kuchki
Phoberocysta neocomica
?
25 15
22
24 7.6
17
23 8.4
15
22 5.5
4
Bechku
Triplasia emslandensis acuta
Lenticulina
andromede
3
Quadratina
Fauriella boissieri,
Neocosmoceras euthymi,
N. minutus, Hegaratia balkensis,
H. bidichotoma, H. nerodenkoi,
H. taurica, Haploceras carachtheis,
H. cristifer, Euphylloceras serum,
Leiophylloceras calypso,
Bochianites neocomiensis,
B. laevis, Spiticeras multiforme,
S. subspitiense, S. obliquelobatum,
S. cf. tenuicostatum
Protetragonites tauricus,
Haploceras ex gr elimatum,
Lytoceras liebigi, Spiticeras sp.
21 18
9
20 8
Dalmasiceras sp.
tunassica
19 4
18 5
17 0.3
16 1.65
15 3.6
14 1.5
13 2
?
15
5.6
2
11–12 1
10 2.8
9 11.5
7–8
45
7
6 6
Dalmasiceras tauricum,
Dalmasiceras sp.,
Malbosiceras malbosi,
M. pictetiforme,
Pomeliceras breveti,
Subalpinites amplus,
S. insolitus,
Euphylloceras serum,
Leiophylloceras calypso,
Protetragonites tauricus
5 6.25
4 2.7
65
3 63
2 12
1 4.5
grandis
Haploceras ex gr. cristifer,
Protetragonites tauricus,
Spiticeras sp., Subalpinites sp.
26 30
Bedenekyr
Dalmasiceras
tauricum
2944
Costacythere drushchitzi–Reticythere marfenini
Conorboides
hofkeri
Neocosmoceras
euthymi
Textularia crimica–Belorussiella taurica
B e r r i a s i a n
Riasanites
crassicostatum
Evirticyclammina virguliana,
Lenticulina
Rectocyclammina recta,
muensteri
Bramkampella arabica
Costacythere khiamii–
Hechticythere belbekensis
Beds with
Symphythiris
arguinensis
Zeillerina baksanensis
Symphythiris arguinensis
?
27 15
2420
Zeillerina baksanensis
Hegaratia sp., Spiticeras sp.
28 30
boissieri
(Yampol’skaya et al., 2006)
5
–
29 40
Lithology
Gaps in sampling, m
Ostracod
beds
Dinocyst
beds
Albat Formation, sequence
Member
Thickness, m
Foraminiferal
beds/
assemblages
Subzone,
beds with fauna
(Arkadiev et al.,
2012)
ARKADIEV et al.
Stage
Zone
158
Malbosiceras ex gr. malbosi
10
50–60
>50
Malbosiceras chaperi,
M. malbosi,
Ptychophylloceras semisulcatum,
Protetragonites tauricus
Pseudosubplanites ponticus,
Berriasella jacobi
Fig. 2. Composite bio- and magnetostratigraphic Berriasian section of central Crimea. For legend, see Fig. 4.
STRATIGRAPHY AND GEOLOGICAL CORRELATION
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2015
m
5
0
m
5
0
occitanica
occitanica
Zone
Lithology
?
?
?
15
14
13
12B
12А
12
11
10
9
8
7
6
5
4
3
2
1
10
9
8
7
6
5
4
3
2
1
270 0
10
Lithology
Texture
Packstone
m
5
0
occitanica
12
11
10
9
Paleomagnetic
samples
Zone
Member
90
Polarity
D°
1
2
3
4
5
6
7
Paleomagnetic
samples
270 0
180 270 –90 –45
2015
Rudstone
I°
45
90
270
–45 45
180
–90
0
90
D°
90
0
I°
159
Fig. 3. Lithological–sedimentological and paleomagnetic data on outcrops 2947–2952. For legend, see Fig. 4.
Texture
Grainstone
0
No. 2
Clay/wackestone
Outcrop 2948
Lithology
270
Outcrop 2951
7
4A
5
1
1
Vol. 23
Polarity
I°
270
–45 45
180 –90
0
90
2
1
Member
D°
90
Zone
Wackestone
Outcrop
2950
Paleomagnetic
samples
Texture
?
?
Texture
40
35
m
5
occitanica
jacobi
Floatstone
Polarity
30
25
18 20
19
15
10
5
1
Outcrop 2947
Clay/wackestone
Packstone
Polarity
Paleomagnetic
samples
Zone
Member
NEW DATA ON BERRIASIAN BIOSTRATIGRAPHY, MAGNETOSTRATIGRAPHY
5.6
6
45
4.8
6
8.3
7
5
5
7.5
Lithology
Packstone
18
17
16
15
10–12(?)
14
13
9
8
7
6
5
4
3
Member
Wackestone
0
STRATIGRAPHY AND GEOLOGICAL CORRELATION
Outcrop 2949 Outcrop 2952
160
ARKADIEV et al.
clast detritus up to 2 mm across, and rare larger bioclasts. The rocks contain intact shells of bivalves Tortarctica weberae Mordv. The thin section contains rare
sections of foraminifers belonging to the Everticyclammina virguliana–Retrocyclammina recta–Bramkampella arabica Assemblage The thickness is 2.7 m.
M e m b e r 5. Gray clays containing carbon detritus, ferruginous ooids, abundant foraminifers of the
Everticyclammina virguliana–Retrocyclammina recta–
Bramkampella arabica Assemblage, and ostracods
Cytherella lubimovae Neale, Cytherelloidea mandelstami Neale, Costacythere khiamii Tes. et Rach.),
C. foveata Tes. et Rach, Hechticythere belbekensis Tes.
et Rach., Quasigermanites bicarinatus moravicus Pok.,
and others. They are accompanied by single spores
and pollen grains, dinocysts of the Muderongia Assemblage, and prasinophytes. The thickness is 0.25 m.
Further, there is an unexposed interval 6 m wide.
M e m b e r 6 (Samples 2949/4–5). Light gray bedded marlstones (layers of approximately 20 cm thick)
with foraminifers of the Everticyclammina virguliana–
Retrocyclammina
recta–Bramkampella
arabica
Assemblage with calcareous forms characterized by
dwarfish sizes. The ostracod assemblage consists of
Cytherella krimensis Neale, C. lubimovae Neale, C. fragilis Neale, and others. The microfossils include also
single spores, pollen, and dinocysts of the Muderonia
assemblage. The thickness is 1.2 m.
There is an unexposed interval 4.8 m wide.
M e m b e r 7 (Samples 2949/6–7). Light gray
clayey limestones with Thalassoinoides burrows and single ferruginous ooids. The thin sections exhibit single
sections of foraminifers from the Everticyclammina virguliana–Retrocyclammina recta–Bramkampella arabica Assemblage. The thickness is 1 m.
There is an unexposed interval 3 m wide.
M e m b e r 8 (Samples 2949/8–10) is poorly
exposed, being represented by isolated outcrops of
gray wacke- and packstones 0.3–0.4 m thick with ferruginate ichnofossils (Thalassinoides?), skeletal detritus, unidentifiable bivalve shells, and ferruginous
oolites. The two upper layers enclose lenses of shelly
floatstones. The thin section demonstrates single foraminifers belonging to the Everticyclammina virguliana–Retrocyclammina recta–Bramkampella arabica
Assemblage. The thickness is 3 m.
Section 2948 (44°59′00.06′′ N,
34°29′10.50′′ E; Fig. 3)
The section comprises seven members (from bottom to top):
Member
9 (Samples 2948/6–7) is poorly
exposed, being represented by two outcrops of gray
wacke- and packstones (0.2–0.3 m thick) with rare
Thalassinoides burrows. Thin sections yield foraminifers of the impoverished Everticyclammina virguliana–
Retrocyclammina
recta–Bramkampella
arabica
Assemblage and ostracods represented by Costacythere
andreevi Tes., C. khaimii Tes. et Rach., Quasigermanites
bicarinatus moravicus Pok., and others. They are
accompanied by dinocysts from the Mudergonia Complex, Systematophora areolata Klement, Kleithriasphaeridium eoinodes (Eisenback), Prolixosphaeridium
parvispinum (Deflandre), Achomosphaera sp., and
Cometodinium habibii Montail. The thickness is 11.5 m.
There is an unexposed interval 2.7 m wide.
M e m b e r 10 (Samples 2948/2–5) is composed of
gray clays (layers 20–50 cm thick) with intercalations
of dark clays (up to 2–3 cm). Some levels yielded ferruginate unidentifiable casts of gastropod and bivalve
shells. The middle part of the member encloses a single lens-shaped intercalation of light gray grainstones
with gastropod shells, impoverished Everticyclammina
virguliana–Retrocyclammina
recta–Bramkampella
arabica foraminiferal assemblage dominated by simple
lituolids and poorly preserved Lenticulina tests, and
ostracods Costacythere khaimii Tes. et Rach. and others. The thickness is 2.8 m.
M e m b e r 11. Yellowish gray slightly consolidated
sandstone with foraminifers of the Everticyclammina
virguliana–Retrocyclammina
recta–Bramkampella
arabica Assemblage and ostracods Costacythere khiamii Tes. et Rach., Hechticythere belbekensis Tes. et
Rach., Schuleridea ex gr. juddi Neale, and others. The
thickness is 0.2 m.
M e m b e r 12 (Sample 2948/1) is composed of
horizontally bedded mixed carbonate–terrigenous
sandstones with an incised channel filled with trough
cross-bedded sediments containing bivalve shells,
which are oriented parallel to bedding surfaces and
saturate some laminae. The sandstones contain rare
crustacean Ophiomorpha sp. burrows. Bivalves are represented by Prohinnites renevieri (Coq.) and Entolium
germanicum (Woll.). The member yielded also a fragment of ammonite Fauriella (?) sp. and foraminifers of
the impoverished Everticyclammina virguliana–Retrocyclammina recta–Bramkampella arabica Assemblage
age dominated by simple lituolids and poorly preserved Lenticulina tests, ostracods Costacythere khiamii Tes. et Rach., Hechticythere belbekensis Tes. et
Rach., Schuleridea ex gr. juddi Neale, and others, and
dinocysts Spiniferites ex gr. ramosus (Ehren.). The
thickness is 0.8 m.
Section 2952 (44°59′09.24′′ N,
34°28′13.36′′ E; Fig. 3)
Similar to Section 2948, this section 2952 encloses
the contact between the calcareous and terrigenous
sequences. On the basis of this feature, its lower part is
correlated with the former section.
M e m b e r 9 (Samples 2952/1–3) is poorly
exposed, being represented by isolated outcrops of
gray wacke- and packstones 0.2–1.0 m thick with
thalassinoid burrows, ferruginous ooids, bioclast
detritus up to 2 mm across, less common larger biocalsts, and intact bivalve shells. The sediments are frequently ferruginate along bedding surfaces. The thickness is 11.5 m.
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NEW DATA ON BERRIASIAN BIOSTRATIGRAPHY, MAGNETOSTRATIGRAPHY
There is an unexposed interval 4 m wide with a single intercalation of packstone, which represents presumably an analog of grainstone in Member 10 (Sample 2952/4) of Section 2948. The thin sections exhibit
single sections of foraminifers Melathrokerion spirialis
Gorb.
M e m b e r 13 (Samples 2952/5–6) is composed of
light greenish gray calcareous clays with sand admixture, carbon detritus, intact unidentifiable casts of
bivalve shells, and their fragments. The rocks contain
the foraminifers of the Lenticulina muensteri Assemblage. Many Lenticulina shells are poorly preserved.
Simple lituolids demonstrate gigantism. The ostracod
assemblage includes Costacythere khiamii Tes. et
Rach., C. foveata Tes. et Rach., Hechticythere belbekensis Tes. et Rach., and others. Microphytoplankton is
represented by unidentifiable proximate dinocysts and
prasinophytes. The thickness is 2 m.
M e m b e r 14 (Sample 2952/7) is represented by
light gray bioturbated floatstones with intact unidentifiable bivalve casts covered by vague encrustations and
shell detritus. The thickness is 0.7 m.
There is an unexposed interval 0.75 m wide.
M e m b e r 15 (Samples 2952/8–10) consists of
greenish gray calcareous clays with abundant carbon
detritus, shelly detritus, and ferruginate structures
(bioturbation?). Accumulations of bivalve shells form
coquina. Bivalves are represented by Gervillella anceps
(Desh. et Leym.), Neithea simpliex Mordv., and Integricardium deshayesianum (Lor.). Up the section, clays
become sandy and contain abundant solitary corals
Montlivaltia crimea Kusm. The upper 0.5 m of the
member is poorly exposed and contains carbon detritus, and single unidentifiable remains of ammonites,
belemnites, bivalves Gervillella anceps (Desh. et
Leym.) and Neithea simpliex Mordv., and foraminifers
of the Lenticulina muensteri Assemblage with single
planktonic forms. The representatives of this assemblage are dwarfish and abnormal in shape. Many Lenticulina and Hoeglundina tests are poorly preserved.
The ostracod assemblage includes Costacythere khiamii
Tes. et Rach., C. foveata Tes. et Rach., C. drushchitzi
(Neale), and other species. There are also spores of
Schizaleales ferns Lygodium sp. and bisaccate coniferous and Classopollis spp. pollen. The thickness is 3.6 m.
M e m b e r 16 (Sample 2952/11) is poorly
exposed, being represented by sandy floatstones with
unidentifiable bivalve shells. The thickness is 0.4 m.
There is an unexposed interval 1.65 m wide.
M e m b e r 17 (Sample 2952/12) is composed of
slightly calcareous clays with carbon detritus, shelly
detritus, and ferruginate structures (sediment feeding
burrows?). The foraminiferal Lenticulina muensteri
Assemblage includes dwarfish forms. Ostracods are
represented by Cytherella krimensis Neale, C. flexuosa
Neale, Pontocypris felix Neale, and other species. The
sediments contain also bisaccate coniferous pollen.
The thickness is 0.3 m.
STRATIGRAPHY AND GEOLOGICAL CORRELATION
161
There is an unexposed interval approximately 5.6 m
wide.
M e m b e r 18 (Samples 2952/12A–15) consists of
weathered pack- and wackestones with vague (due to
poor exposure) clay intercalations (5–15 cm thick).
The pack- and wackestones demonstrate thalassinoid
burrows, skeletal detritus, and single ferruginous
ooids. They contain foraminifers of the Everticyclammina virguliana–Retrocyclammina recta–Bramkampella arabica Assemblage dominated by Melathrokerion spirialis Gorb. The ostracod assemblage includes
Cytherella lubimovae Neale, Cytherelloidea mandelstami Neale, Costacythere khiamii Tes. et. Rach.,
C. foveata Tes. et Rach., Hechticythere belbekensis Tes.
et Rach., Reticythere marfenini Tes. et Rach.,
Schuleridea ex gr. juddi Neale, and other forms. The
middle part of the member is unexposed (interval 1.2 m
wide). The thickness is 5 m.
The X-ray phase analysis of clay samples from Sections 2948, 2949, and 2952 revealed that all of them are
characterized by a practically identical composition:
quartz, calcite, and minerals of the kaolinite group,
mica, chlorite, rarely albite, anorthite, and gibbsite. A
single sample yielded dolomite. The samples were analyzed with a Rigaku MiniFlex II diffractometer.
Section 2944 (Novoklenovo; 44°59′46.80′′ N,
34°30′16.40′′ E; Fig. 4)
M e m b e r 19 (Samples 2944/1–3) is largely composed of yellow to brown unconsolidated clayey siltstones and brownish gray clays. The lower part of analogs of this member near the Balki settlement contains
an intercalation of brown calcareous sandstones (0.4 m
thick) with compact marlstone concretions, which
yielded ammonites Malbosiceras chaperi (Pict.),
M. malbosi (Pict.), and others; bivalves Entolium germanicum (Woll.), Aetostreon subsinuatum Leym., and
Integricardium deshayesianum (Lor.); and brachiopods
Sellithyris cf. uniplicata Smirn. The thickness is 4 m.
M e m b e r 20 (Samples 2944/4–9, Novoklenovo;
Samples 2940/1–7, Balki) consists of brown clays and
siltstones with marlstone concretions. Near the
Novoklenovo settlement, the member yielded ammonites Dalmasiceras tauricum Bogd. et Ark. and its analogs near the Balki settlement, Dalmasiceras tauricum
Bogd. et Ark., Malbosiceras malbosi (Pict.), M. pictetiforme Tav., Pomeliceras breveti (Pom.), Subalpinites
amplus Ark., S. insolitus Ark. and others; bivalves Pycnodonte weberae Yanin; and brachiopods Loriolithyris
cf. valdensis (Lor.) and Sellithyris ex gr. gratianopolitensis (Pict.). The member contains also diverse
microfossils: foraminifers of the Quadratina tunassica
Assemblage; ostracods Cytherelloidea flexuosa Neale,
Pontocyprella nova Neale, Cypridea funduklensis Tes.
et. Rach., Acrocythere alexandrae Neale et Kolp., and
other forms; single spores and Classopollis spp. pollen;
dinocyst of the Phoberocysta neocomica Assemblage;
prasinophytes; and acritarchs. The thickness is 8 m.
Vol. 23
No. 2
2015
Polarity
Paleomagnetic
samples
Lithology
Member
Zone
162
Lithology
D°
270
20
28
boissieri
15
9m
30 m
25
45
90
Bivalves
Calcareous clay
Brachiopods
Siltstone
Ammonites
Sandstone
Gastropods
Calcareous
sandstone
Quartz
conglomerate
Limestone
10
26
90 180 270 –90 –45 0
Clay
5
1
65
25
Corals
Belemnites
Bioclasts
Clayey limestone
Plant detritus
Reefal limestone
Skeletal detritus
Sponge limestone
Large plant remains
60
22 m
24
boissieri
17 m
23
55
50
45
Coquina
40
Silty marlstone
35
Marlstone concretions
Marcasite
concretions
Ferruginous ooids
Marlstone
15 m
30
25
22
4m
10
9m
5
20
No. 2
m
5
1
Lithology
15
occitanica Zone
Member
occitanica
Vol. 23
Outcrop 2940
21
Polarity
Paleomagnetic
samples
20
Outcrop 2944
Stratigraphic
unconformities or
erosional surfaces
9
20
19
1
0
2015
Fig. 4. Lithological–sedimentological and paleomagnetic data on outcrops 2940, 2943, and 2944.
Ichnofossils
Thalassinoides
Unidentifiable
ichnofossils
Structures
Trough cross bedding
Geomagnetic polarity
Normal polarity
of geomagnetic field
Reverse polarity
of geomagnetic field
Abnormal polarity
of geomagnetic field
Data on polarity omitted
from consideration
No data on polarity
ARKADIEV et al.
STRATIGRAPHY AND GEOLOGICAL CORRELATION
Outcrop 2943
?m
27
0
I°
Fossils
NEW DATA ON BERRIASIAN BIOSTRATIGRAPHY, MAGNETOSTRATIGRAPHY
Section 2940 (Balki; 44°59′21.94′′ N,
34°28′07.11′′ E; Fig. 4)
M e m b e r 21 (Samples 2940/8–23) comprises
dark gray, dark brown, and brown-gray (dominant)
clays with rare marlstone concretions and greenish
gray calcareous siltstones. In its middle part, the member encloses several intercalations (0.15–0.20 cm
thick) of calcareous siltstones representing oyster
accumulations with Pycnodonte weberae Yanin. The
member contains diverse macro- and microfossils:
ammonites Dalmasiceras sp. in its lower part; ammonites Protetragonites tauricus (Kulj.-Vor.), Haploceras
ex gr. elimatum (Opp.), Lytoceras liebigi (Opp.), and
Spiticeras sp. in the upper part; bivalves Entolium germanicum (Woll.), Spondylus complanatus (d’Orb.),
and others; brachiopods Loriolithyris valdenses (Lor.),
Sellithyris uniplicata Smirn., Belbekella airgulensis
Moiss, and other species; foraminifers of the Triplasia
emslandensis acuta Assemblage (some layers contain
giant agglutinated forms and planktonic species);
ostracods Pontocypris cuneata Neale, Acrocythere
alexandrae Neale et Kolp., Costacythere khiamii Tes.
et Rach., Hechticythere belbekensis Tes. et Rach., and
Reticythere marfenini Tes. et Rach.; single spores
including Schizaleales ferns Cicatricosisporites sp. and
pollen of Classopolllis spp.; dinocysts of the Phoberocysta neocomica Assemblage. The thickness is 18 m.
M e m b e r 22 (Samples 2940/24–33) is composed
of dark greenish gray and brown (dominant) viscous
clays and dark gray and brownish siltstones. The member contains different organic remains scattered
through its entire sections: abundant small ferruginate
casts of ammonites Neocosmoceras euthymi (Pict.),
N. minutus Ark. et Bogd., Hegaratia bidichotoma (Bogd.
et Kvant.), H. nerodenkoi (Bogd. et Kvant.), H. balkensis
(Bogd. et Kvant.), Spiticeras multiforme Djan., S. subspitiense (Uhl.), S. obliquelobatum (Uhl.), Bochinites
neocomiensis (d’Orb.), B. laevis Liu, and others (Fauriella boissieri (Pictet) from the collection by V.V. Drushchits originates likely from the same member); bivalves
Pycnodonte weberae Yanin, Aetostreon subsinuatum
(Leym.), and others; brachiopods Loriolithyris valdensis
(Lor.), Symphythiris arguinensis (Moiss.), Terebratuliopsis quadrata quadrata Smirn., and other forms; rare
foraminifers from the impoverished Triplasia emslandensis acuta assemblage; ostracods Cytherelloidea flexuosa Neale, Bythoceratina ex gr. variabilis Donze,
Eucytherura aff. trinodosa Pok., and others; spores of
Schizaleales ferns Cicatricosisporites sp., pollen of Classopolllis spp., and prasinophytes. The thickness is 5.5 m.
M e m b e r 23 (Samples 2940/34–44) is represented by dark greenish gray and brown clays alternating with dark gray siltstones. The member contains the
impoverished Triplasia emslandensis acuta Assemblage
with dominant Spirillina kubleri Mjatl. and ostracods
Cytherella krimensis Neale, C. fragilis Neale, and others. The thickness is 8.4 m.
M e m b e r 24 (Samples 2940/45–58) consists of
dark gray and brown clays alternating with brown calcareous siltstones. The rocks contain ammonites
STRATIGRAPHY AND GEOLOGICAL CORRELATION
163
Riasanites crassicostatum (Kvant. et Lys.), Riasanites sp.,
Hegaratia taurica (Bogd. et Kvant.), H. bidichotoma
(Bogd. et Kvant.), and others and brachiopods Loriolithyris valdensis (Lor.), Symphythiris arguinensis
(Moiss.), Terebratuliopsis quadrata quadrata Smirn.,
and other forms. In addition, ammonite Faurella simplicostata (Maz.) was found by B.T. Yanin in this member. The member under consideration contains also
the impoverished foraminiferal Triplasia emslandensis
acuta Assemblage dominated by agglutinated forms
and ostracods represented by Cytherella lubimovae
Neale, C. fragilis Neale, Costacythere andreevi Tes., and
others. The thickness is 7.6 m.
M e m b e r 25 (sponge horizon, Samples 2940/59–
65) in the outskirts of the Balki settlement is characterized by the following structure (from the base upward):
(1) Greenish gray unconsolidated clays (approximately 5 m) with abundant brachiopod Symphythiris
arguinensis (Moiss.) remains, foraminifers of the Triplasia emslandensis acuta Assemblage dominated by
Lenticulina macra Coup., Spirillina kubleri Mjatl.
accompanied by single planktonic forms, spores of
Schizaleales ferns Cicatricosisporites sp. and pollen of
Classopolllis spp., dinocysts Systematophora areolata
Klement and Phallocysta elongata (Beju), prasinophytes, and acritarchs.
(2) Light gray compact clotted limestones with
abundant sponge skeletons, small oysters Aetostreon
subsinuatum (Leym.), spines of echinoderms Diplocidaris (?) bicarinata Web., gastropods, and other
organic remains such as brachiopods Loriolithyris
valdensis (Lor.) and Symphythiris arguinensis (Moiss.),
single ammonites Riasanites crassicostatum (Kvant. et.
Lys.), and belemnite rostra. The limestones constitute
isolated bioherms 1.0–1.5 m across submerged into
the matrix of greenish clays. The thickness is at least
10–12 m.
(3) Alternating greenish gray clays and compact
gray calcareous siltstones (4–6 m) crossed by abundant vertical Ophiomorpha and Thalassinoides burrows. The sediments contain single weathered pyrite
concretions. The impoverished Triplasia emslandensis
acuta foraminiferal assemblage is dominated by agglutinated forms accompanied by poorly preserved calcareous tests.
(4) Greenish gray unconsolidated clays (5 m) with
ammonites Hegaratia sp. and Spiticeras sp. The Lenticulina andromede foraminiferal assemblage includes
single specimens of planktonic forms.
Ostracods scattered through the entire sponge
member are represented by the following species:
Cytherella krimensis Neale, C. lubimovae Neale,
Cytherelloidea flexuosa Neale, Neocythere pyrena Tes.
et Rach. The integral thickness of the member is 28 m.
Vol. 23
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2015
164
ARKADIEV et al.
Section 2943 (Mezhgor’e; 44°58′49.95′′ N,
34°24′27.60′′ E; Fig. 4)
Samples 2943/1–5 were taken in the upper part of
Member 25. The overlying sediments of Member 26
remained unsampled.
M e m b e r 26 (Samples 2943/6–13 were taken in
the uppermost part of the member; samples for paleomagnetic measurements were taken from this member
in 2002 in section 2420 near the Pasechnoe settlement
located 600–700 m northeast of Section 2943) is composed of greenish gray unconsolidated clays and yellowish gray fine-grained sandy siltstones. Higher in
the section, siltstones become progressively more calcareous to grade into marlstones. The member contains diverse organic remains: poorly preserved
ammonites Haploceras ex gr. cristifer (Opp.), Protetragonites tauricus (Kulj.-Vor.), Spiticeras sp., and Subalpinites sp.; abundant bivalves Gervillella cf. terekensis
(Renng.), Entolium germanicum (Woll.), Chlamys
goldfussi (Desh.), Neithea neocomiensis (d’Orb.),
N. simplex Mordv., Plagiostoma dubisiensis (Pict. et
Camp.), Ceratostreon minos (Coq.), Aetostreon subsinuatum Leym., and others; brachiopods Loriolithyris
valdensis (Lor.), Terebratuliopsis quadrata quadrata
Smirn., Weberithyris moisseevi (Web.), and others;
echinoderms Acrocidaris minor Ag., Rhabdocidaris aff.
burganensis Web., and Diplocidaris (?) bicarinata Web.;
crinoids Apiocrinus cf. valangiensis Lor. Sediments
from the lower part of the member contain foraminifers of the Lenticulina andromede Assemblage; in its
upper part, the latter is replaced by the Conorboides
hofkeri Assemblage. Ostracods are represented by the
following species: Cytherelloidea mandelstami Neale,
Bairdia menneri Tes. et. Rach., B. kuznetsovae Tes.
et. Rach., Cypridea funduklensis Tes. et Rach.,
Eucytherura paula Lueb., Neocythere dispar Donze,
Costacythere drushchitzi (Neale), and others. There
are also spores of Schizaleales ferns and other plants,
Classopollis spp. pollen, dinocysts of the Phoberocysta neocomica Assemblage, prasinophytes, and
acritarchs. The thickness is 30 m.
M e m b e r 27 (Samples 2943/14–15) consists of
light gray and yellowish gray massive and slightly consolidated marlstones. Its upper part is characterized by
diverse benthic macrofossils; corals; brachiopods Terebratuliopsis quadrata quadrata Smirn., Weberithyris
moisseevi (Web.), Zeillerina baksanensis Smirn., and
others; bivalves Chlamys goldfussi (Desh.), Neithea
atava (Roem.), N. neocomiensis (d’Orb.), Ceratostreon
minos (Coq.), and others; echinoderms Rhabdocidaris
aff. burganensis Web. and Pygopyrina incisa (Ag.); and
crinoids A. neocomiensis (d’Or.).
In 2002, E.Yu. Baraboshkin found in this member
brachiopods Loriolithyris valdensis (Lor.), Cyclothyris
rectimarginata Smirn., Septaliphoria gerassimovi
Moiss., Symphythiris koinautensis (Moiss.), Advenina
villersensis (Lor.), and other species.
The sediments contain also the impoverished foraminiferal association with Textularia crimica (Gorb.)
and dominated by simple lituolids. The thickness is 15 m.
M e m b e r 28 (Samples 2943/16–20) is represented by light brown-gray compact clotted biohermal
limestones with frequent accumulations of brachiopods Zeillerina baksanensis Smirn. and others in the
lower part of the member, where they form coquinas,
and abundant rudists in its upper part. Bioherms 1.5–
2.0 m high are formed by corals, rudists, and algae and
surrounded by organogenic–detrital and detrital limestones. The thickness is 25–30 m.
The Zeillerina baksanensis Beds are barren of guide
ammonite species; therefore, many researchers considered them Valanginian (Drushchits and Yanin,
1959; Gorbachik et al., 1975). Subsequently, limestones were attributed to the Megadiceras koinautense
Beds, which were dated back to the late Berriasian
(Yanin and Baraboshkin, 2000).
The biohermal limestones of the Kuchki Formation near the Mezhgor’e settlement are overlain with
the karst- and erosion-affected surface by quartz conglomerates (5–40 m), which may likely be considered
as analogous to the Albat Member of the Bel’bek River
basin, where it is, in turn, overlain by lower Valanginian strata (Yanin and Baraboshkin, 2000). Near the
Mezhgor’e settlement, the surface of limestones is
similarly uneven and bears indications of activity of
borers. The conglomerates are overlain by a sandstone
and clay sequence, which is overlain near the Balki settlement by oncolitic limestones with gastropods and
rudists. In the opinion of T.N. Bogdanova in
(Bogdanova et al., 1981), the conglomerates are
replaced west of the Mezhgor’e settlement by white
gastropod limestones, which are traceable up to the
Petrovo settlement.
BIOSTRATIGRAPHY
Ammonites. Of extreme interest is the finding of
ammonite Malbosiceras ex gr. malbosi (Pictet) (Plate I)
in the carbonate Bedenekyr Formation that underlies
the terrigenous Bechku Formation in the outskirts of
the Balki settlement (Section 2951). This ammonite
was found in talus stratigraphically below the Malbosiceras chaperi Beds. The last unit defined by
V.V. Arkadiev and T.N. Bogdanova in the Sary-Su River
basin was correlated with the upper part of the jacobi
Zone above the grandis Zone (Arkadiev et al., 2002,
2006). Previously, these sediments defined as the Malbosiceras (?) sp. Beds were attributed to the occitanica
Zone (Bogdanova et al., 1981). Their correlation with
any of these zones was ambiguous. V.V. Arkadiev identified from these beds M. malbosi (Pictet) together
with M. chaperi. In Berriasian sections of Western
Europe, the species M. malbosi is characteristic of the
boissieri Zone (paramimounum Subzone), while
M. chaperi occurs in the upper part of the jacobi Zone
(Le Hégarat, 1973; Tavera, 1985). In Crimea, M. malbosi is, in addition, documented in the occitanica
STRATIGRAPHY AND GEOLOGICAL CORRELATION
Vol. 23
No. 2
2015
NEW DATA ON BERRIASIAN BIOSTRATIGRAPHY, MAGNETOSTRATIGRAPHY
165
Plate I
(a)
(b)
1 cm
Plate I. Malbosiceras ex gr. malbosi (Pictet), specimen 1/13244.
(a) Ventral view (×0.7); (b) lateral view (×0.7); Balki settlement, Member 2, Berriasian, occitaniica Zone.
Zone (Arkadiev et al., 2012). Therefore, it is more reasonable to assume that it occurs at the lower level up to
the jacobi Zone, not that M. chaperi continues occurring up to the occitanica Zone. In such a situation, the
find of Malbosiceras ex gr. malbosi indirectly correlates
the carbonate part of the examined section with the
occitanica Zone. Magnetostratigraphic data, which
register reverse polarity in outcrops 2950 and 2951
(Fig. 3), are consistent with correlation between the
STRATIGRAPHY AND GEOLOGICAL CORRELATION
Malbosiceras chaperi Beds and occitanica Zone. In
the paleomagnetic scale, Chron M17 corresponds to
the upper part of the jacobi Zone and largest part of the
occitanica Zone (Ogg and Hinnov, 2012).
Bivalves. The identified bivalve remains include
both guide and marker species. In (Arkadiev et al.,
2012), B.T. Yanin defined in the Berrisian Stage of
Crimea three stratigraphic assemblages: lower–middle Berriasian, middle Berriasian, and upper BerriaVol. 23
No. 2
2015
166
ARKADIEV et al.
sian. Neithea simplex Mord. is a guide form for the
lower assemblage. This species is characteristic of Berriasian strata of Kopetdag, Mangyshlak, and the
northern Caucasus, in addition to the Crimean
Mountains. Prohinnites renevieri (Coq.) occurs from
the lower strata of the Berriasian Stage and characterizes its entire section in Crimea. Entolium germanicum
(Woll.) is also characterized by a wide stratigraphic
range occurring beyond the limits of the Berriasian
Stage. At the same time, in Crimea this species
appearing in the jacobi Zone forms peculiar coquinas
in clayey sediments of the central Crimean Mountains; i.e., it represents a marker species for this part of
the region under consideration. The species is less
characteristic of the middle and upper parts of the
Berriasian Stage, where it occurs as single specimens.
Gervillella anceps (Desh. in Leym.) and Integricardium deshayesianum (Lor.) are characteristic species
of the upper middle Berriasian Dalmasiceras tauricum
Subzone and largest part of the upper Berriasian. In
sections of the Bel’bek River basin in southwestern
Crimea, the first species (marker) forms coquinas. The
second species is less frequent and never forms
coquinas, being mostly distributed in the middle part
of the Berriasian Stage in this area of the Crimean
Mountains. Tortarctica weberi Mord. is also abundant
in the same layers of the Berriasian section.
Foraminifers. These microfossils from Berriasian
sediments of central Crimea (Plate II) were investigated
in residues and thin sections using variably oriented sections. For samples from some levels, shells extracted
from rocks were used for preparing polished sections. In
total, foraminifers from the composite Berriasian sec-
Plate II. Foraminifers from Berriasian sediments of central Crimea. Magnification: ×20 for figs. 1–25, ×35 for figs. 26–49, ×60
for fig. 50.
(1) Haplophragmium subaequale (Mjatl.), specimen 27/1324: (1a) lateral view, (1b) peripheral view, (1c) polished thin section,
Balki settlement, Member 18; (2, 3) Charentia evoluta Gorb: (2) specimen 31/1324: (2a) lateral view, (2b) peripheral view, Enisarai Ravine, Member 9; (3) specimen 32/1324, thin section, transverse section, Enisarai Ravine, Member 1; (4, 5) Stomatostoecha
compressa Gorb.: (4) specimen 35/1324: (4a) lateral view, (4b) peripheral view, Enisarai Ravine, Member 9; (5) specimen
36/1324, thin section, transverse section, Enisarai Ravine, Member 1; (6–8) Stomatostoecha enisalensis Gorb.: (6) specimen
38/1324, thin section, transverse section, Balki settlement, Member 18; (7) specimen 39/1324, oblique section close to the longitudinal one, Enisarai Ravine, Member 1; (8) specimen 37/1324: (8a, 8b) lateral view, (8c) peripheral view, Enisarai Ravine,
Member 11; (9, 10) Melathrokerion spirialis Gorb.: (9) specimen 33/1324: (9a, 9b) lateral view, (9c) peripheral view, Balki settlement, Member 18; (10) specimen 34/1324, thin section, transverse section, Enisarai Ravine, Member 1; (11) Rectocylammina
chouberti Hott., specimen 48/1324: (11a) lateral view, (11b) apertural view, (11c) polished section, Enisarai Ravine, Member 5;
(12) Rectocylammina recta Gorb., specimen 52/1324: (12a) lateral view, (12b) apertural view, (12c) polished section, Enisarai
Ravine, Member 10; (13) Rectocylammina arrabidensis Remalho, specimen 46/1324: (13a) lateral view, (13b) apertural view,
(13c) polished section, Enisarai Ravine, Member 11; (14, 16) Everticyclammina virguliana (Koechl.): (14) specimen 44/1324:
(14a) lateral view, (14b) apertural view, (14c) polished section, Enisarai Ravine, Member 5; (16) specimen 42/1324, lateral view,
Enisarai Ravine, Member 6; (15) Everticyclammina elongata Gorb., specimen 45/1324, polished section, Enisarai Ravine, Member 4; (17) Alveosepta jaccardi (Schrodt), specimen 62/1324: (17a, 17b) lateral view, (17c) peripheral view, Balki settlement,
Member 18; (18) Amijiella amiji (Henson), specimen 54/1324, thin section, longitudinal section, Enisarai Ravine, Member 4;
(19) Bramkampella arabica Radm., specimen 55/1324, thin section, longitudinal section, Enisarai Ravine, Member 11;
(20, 21) Textularia crimica (Gorb.), specimen 56/1324: (20a) lateral view, (20b) apertural view, Mezhgor’e settlement, Member 26;
(21) specimen 57/1324, thin section, longitudinal section, ibid; (22–24) Belorussiella taurica Gorb.: (22) specimen 58/1324, lateral view, Mezhgor’e Settlement, Member 26; (23) specimen 59/1324, thin section, ibid; (24) specimen 60/1324, thin section,
ibid; (25) Triplasia emslandensis acuta Brat. et Brand, specimen 30/1324: (25a, 25b) lateral view, (25c) apertural view, Balki settlement, Member 21; (26) Lenticulina ongkodes Esp. et Sigal, specimen 66/13244, Balki settlement, Member 21; (27) Lenticuina
aquilonica Mjatl., specimen 67/13244: (27a) lateral view, (27b) peripheral view, Enisarai Ravine, Member 15; (28) Lenticulina aff.
uspenskajae K. Kuzn., specimen 68/13244: (28a) lateral view, (28b) peripheral view, Enisarai Ravine, Member 15; (29) Lenticulina muensteri (Roemer), specimen 63/13244: (29a) lateral view, (29b) peripheral view, Mezhgor’e settlement, Member 26;
(30) Lenticulina andromede Esp. et. Sigal, specimen 64/13244: (30a) lateral view, (30b) peripheral view, Mezhgor’e settlement,
Member 26; (31) Lenticulina colligoni Esp. et Sigal, specimen 65/13244: (31a) lateral view, (31b) peripheral view, Enisarai Ravine,
Member 15; (32) Lenticulina bifurcata Bart. et Brand, specimen 69/13244: (32a) lateral view, (32b) peripheral view, Enisarai
Ravine, Member 15; (33) Lenticulina sp. (L. sp. 1 Gorb.), specimen 80/13244: (33a) lateral view, (33b) peripheral view, Balki settlement, Member 21; (34) Lenticulina macra Gorb., specimen 70/13244: (34a) lateral view, (34b) peripheral view, Enisarai Ravine,
Member 13; (35) Lenticulina fracta Esp. et Sigal, specimen 73/13244: (35a) lateral view, (35b) peripheral view, Enisarai Ravine,
Member 13; (36, 37) Lenticulina ambanjabensis (Esp. et Sigal): (36) specimen 77/13244: (36a) lateral view, (36b) peripheral view,
Mezhgor’e settlement, Member 26; (37) specimen 78/13244, thin section close to the orthogonal one, Enisarai Ravine, below
Member 1; (38) Lenticulina eichenbergi Bart. et Brand, specimen 82/13244: (38a) lateral view, (38b) peripheral view, Mezhgor’e settlement, Member 26; (39) Lenticulina neocomina Rom., specimen 74/13244: (39a) lateral view, (39b) peripheral view, Mezhgor’e
settlement, Member 26; (40) Astacolus mutilatus Esp. et Sigal, specimen 84/13244, Balki settlement, Member 18; (41) Astacolus proprius K. Kuzn., specimen 85/13244, Balki settlement, Member 18; (42) Astacolus folium (Wisn.), specimen 86/13244, Balki settlement, Member 18; (43) Saracenaria latruncula (Chalilov), specimen 87/13244, Balki settlement, Member 21; (44) Saracenaria
inflata Pathy, specimen 90/13244: (44a) lateral view, (44b) peripheral view, Mezhgor’e settlement, Member 26; (45) Saracenaria aculata Esp. et Sigal, specimen 89/13244: (45a) lateral view, (45b) peripheral view, Enisarai Ravine, Member 15; (46) Saracenaria compacta (Esp. et Sigal), specimen 91/13244: (46a) lateral view, (46b) peripheral view, Enisarai Ravine, Member 15; (47) Saracenaria
tsarmandrosoensis Esp. et Sigal, specimen 93/13244, Mezhgor’e settlement, Member 26; (48) Saracenaria provoslavlevi Furs. et Pol.,
specimen 94/13244: (48a) lateral view, (48b) peripheral view, Enisarai Ravine, Member 13; (49) Pseudosaracenaria truncata Pathy,
specimen 92/13244, Balki settlement, Member 21; (50) Conorboides hofkeri (Bart. et Brand), specimen 111/13244: (50a) dorsal view,
(50b) ventral view, (50c) peripheral view, Mezhgor’e settlement, Member 26.
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NEW DATA ON BERRIASIAN BIOSTRATIGRAPHY, MAGNETOSTRATIGRAPHY
167
Plate II
5
4a 4b
2a
9a
1a
1b
3
2b
9b
6
8a
8b
9c
7
8c
10
1c
13b
11b
14b
12b
16
15
11a
11c
12a
12c
19
13a
18
13c
22
17a
17b
23
14a
20a
24
14c
21
20b
17c
25c
26
30a
29a
25a
25b
27a 27b
28a 28b
30b
32a 32b
29b
31a
31b
36a
34a
33a
34b
35a
50a
38b
37
35b
33b
38a
36b
39a
50b
50c
39b
44a 44b
40
41
42
43
STRATIGRAPHY AND GEOLOGICAL CORRELATION
45a
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No. 2
2015
47
48a
48b
49
168
ARKADIEV et al.
tion are represented by over 200 species of 63 genera
(Fig. 5). As a whole, the identified foraminiferal assemblage is characteristic of the Textularia crimica–
Belorussiella taurica Beds developed through the entire
Crimean Region (Feodorova, 2004). The following six
successive foraminiferal assemblages may be defined in
the section under consideration on the basis of changes
in the taxonomic composition and quantitative parameters (from the base upward) (Fig. 5):
1. The Everticyclammina virguliana–Retrocyclammina recta–Bramkampella arabica Assemblage
(Members 1–12 and 18). This assemblage is characterized by the prevalence of lituolids (including complex) over nodosariid. The assemblage numbers 70 species of 45 genera in total. The characteristic species are
abundant and diverse representatives of the genera
Everticyclammina and Retrocyclammina, including
Retrocyclammina ex gr. chouberti Hott, R. recta Gorb.,
Everticyclammina virguliana (Koechl.), and E. elongata Gorb., as well as Haplophragmium subaequale
(Mjatluk), Melathrokerion spirialis Gorb., Charentia
evoluta (Gorb.), Pseudocyclammina lituus (Yok.), Stomatostoecha rotunda Gorb., S. compressa Gorb.,
S. enisalensis Gorb., Bramkapella arabica Radm.,
Amijella amiji (Henson), Belorussiella taurica Gorb.,
Alveosepta jaccardi (Schrodt), Astacolus mulitatus Esp.
et Sigal, A. inspissatus (Loeblich et Tappan), A. favoritus Gorb., Discorbis miser Gorb., Trocholina alpina
(Leup.), T. elongata (Leup.), T. molesta Gorb.,
T. infragranulata Noth, and others. Thin sections
from the lower part of the section (Member 2) yielded
single specimens of Protopeneroplis ultragranulatus
(Gorb.) and Pseudosiphoninella antiqua (Gorb.). The
assemblage is named after species E. virguliana,
R. recta, and B. arabica.
The Rectocyclammina Beds transitional from the
upper Tithonian to lower Berriasian, which are established on the Ai-Petri Plateau, and the Bramkapella
Beds from the overlying limestone sequence (Gorbachik and Mokhamad, 1999) are characterized by a
similar foraminiferal assemblage. The Bramkapella
Beds are taken by the last authors to be the early Berriasian in age.
It should be noted that the Everticyclammina virguliana–Retrocyclammina recta–Bramkampella arabica Assemblage is registered also in Member 18,
which is located higher in the section. The assemblage
from this member is characterized by the presence of
many (several hundred) specimens of Melathrokerion
spirialis Gorb. and species Flabellammina lidiae Gerke
et Pol. and Triplasia elegans (Mjatl.), which are known
from terminal Jurassic strata of the Boreal and Arctic
provinces, against the background of dominant complex Lituolidae. These data combined with petromagnetic measurements may presumably indicate that
Member 18 is repeated in the section, although this
assumption requires additional investigations. We
leave Member 18 in the general stratigraphic succession, although with the question mark.
2. The Lenticulina muensteri Assemblage (Members 13–17, 19) is notably dominated by Nodosariidae
with representatives of the genera Lenticulina being
particularly abundant and diverse and genera Saracenaria and Pseudonodosaria being subdominant. The
assemblage is named after the characteristic species,
which is present in all the examined samples. In total,
the assemblage includes approximately 65 species of
22 genera with Ramulna aculeata Wright, Lenticulina
nimbifera Esp. et Sigal, L. fracta Esp. et Sigal, Pseudonodosaria diversa (Hoff.), Saracenaria compacta Esp.
et Sigal, and Hoeglundina ex gr. caracolla (Roemer)
being dominant. The following species are also characteristic: Dorothia ex. oxycona (Reuss), D. kummi
Zedler (minima), Nodosaria raristriata Chapman,
Tristix acutangulus (Reuss), Lenticulina muensteri
(Roemer), L. colligoni Esp. et Sigal, Astacolus proprius
Kun., A. incurvatus (Reuss), Marginulina striatocostata
Reuss, M. micra Tairov, Marginulinopsis sigali Bart.,
Bett. et Bolli, Saracenaria provoslavlevi Furs. et Pol.,
S. provoslavlevi Furs. et Pol. var. minima, S. aculata
Esp. et Sigal, Dentalina gracilis d’Orb., D. guttinfera
d’Orb., Citharinella pectinatimornata Esp. et Sigal,
Trocholina micra Dulub., and others. The characteristic feature of this assemblage is alternative development of normal, dwarfish, and giant forms.
3. The Quadratina tunassica Assemblage
(Member 20) is the least representative one, consisting only of approximately 30 species belonging to
23 genera. The assemblage is characterized by gigantism among simple lituolids. It is named after the
index species of the Quadratina tunassica–Siphonella
antiqua Zone (Drushchits and Gorbachik, 1979),
which corresponds approximately to the upper part of
the grandis Zone and lower part of the occitanica
Zone. It is defined on the basis of the appearance of
Quadratina tunassica Schokhina and Lenticulina protodecimae Dieni et Massari and the presence of single
transit species such as Textularia crimica (Gorb.),
T. densa Hoff., Citharinella pectinatimornata Esp. et
Sigal, Citharina flexuosa (Bruck.), Tristix acutangulus
(Reuss), Lenticulina colligoni Esp. et Sigal, L. muensteri (Roemer), Astacolus incurvatus (Reuss), Nodosaria paupercula Reuss, Saracenaria latruncula (Chalilov), Planularia crepidiularis Roemer, Lagena sztejnae Dieni et Massari, and Spirillina kubleri Mjatl.
4. The Triplasia emslandensis acuta Assemblage
(Members 21–24 and lower part of Member 25) is distributed discretely through the section. In total, the
assemblage includes over 100 species of 47 genera,
among which Lenticulina representatives are dominant and species of the genera Saracenaria and Verneuilina are subdominant. It is named after the index
species of the synonymous Triplasia emslandensis
acuta Subzone (Kuznetsova and Gorbachik, 1985),
approximately correlated with the upper part of the
occitanica Zone and lower part of the boissieri Zone.
The assemblage is defined on the basis of the
appearance of Recurvoides ex gr. paucus Dubr., Haplophragmoides subchapmani K. Kuzn., Triplasia emslan-
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NEW DATA ON BERRIASIAN BIOSTRATIGRAPHY, MAGNETOSTRATIGRAPHY
densis acuta Brat. et Brand., Pseudolamarckina reussi
(Ant.), Lenticulina nuda (Reuss), L. nodosa (Reuss),
and Saracenaria inflata Pathy and the presence of
abundant specimens of transit species: Lenticulina
macra Gorb., L. neocomina Rom., Vaginulina kochii
Roemer, Saracenaria latruncula (Chalilov), Spirillina
kubleri Mjatl.
5. The Lenticulina andromede Assemblage (upper
part of Member 25 and lower part of Member 26) is
represented by over 70 species of 40 genera with the
distinctly dominant share of Lenticulina representatives. The assemblage is marked by the appearance of
abundant Lenticulina andromede Esp. et Sigal, Tristix
valanginica Schokhina, Lenticulina guttata guttata
(Dam), L. ex gr. ouachensis Sigal, L. praegaultina Bart.,
Falsopalmula costata Gorb., and Istriloculina rectoangularia Mats. et Temirb. They are accompanied by
species inherited from underlying sediments: abundant Ramulina aculeata Wright, Ammobaculites inconstans gracilis Bart. et Brand, Textularia crimica
(Gorb.), Belorussiella taurica Gorb., Lenticulina nuda
(Reuss), L. macra Gorb., L. neocomina Rom., Discorbis
praelongus Gorb., Spirillina kubleri Mjatl., and others.
6. The Conorboides hofkeri Assemblage (upper part
of Member 26) is named after one of the index species
of the Conorbina heteromorpha–Conorboides hofkeri Zone (Drushchits and Gorbachik, 1979), correlated with the upper part of the occitanica Zone and
boissieri Zone.
The assemblage includes over 50 species of 30 genera being barren of distinct dominant forms. It is
marked by the appearance of Dorothia kummi Zedler,
Dentalina marginulinoides Reuss, Miliospirella caucasica Ant., Discorbis agalarovae Ant., Epistomina
tenuicostata Bart. et Brand, E. ornata (Roemer), and
Conorboides hoffkeri (Bart. et Brand), accompanied by
abundant transit species Belorussiella taurica Gorb.,
Pseudolamarckina reussi (Ant.), Saracenaria inflata
Pathy, Hoeglundina ex gr. caracolla (Roemer), and
Spirillina kubleri Mjatl. and common Bulbabaculites
inconstans (Bart. et Brand), Nautiloculina oolithica
Mochler, Textularia crimica (Gorb.), Lenticulina
macra Gorb., L. muensteri (Roemer), Astacolus
ambanjabensis (Esp. et Sigal), Trocholina giganta
Gorb. et Manz., and others.
In central Crimea, Berriasian foraminifers were
investigated by T.N. Gorbachik (Kuznetsova and Gorbachik, 1985). The comparison of our data with the
results of this author is difficult since she mentions
only 31 species for the entire Berriasian section of central Crimea, including taxa identified with the open
nomenclature.
The foraminiferal species documented in the
examined sections are known from the Tithonian–
Valanginian sections of Crimea, the Caucasus, the
Caspian region, Syria, Germany, France, Italy, and
Madagascar. Within the Tethyan region, the Everticyclammina
virguliana–Retrocyclammina
recta–
Bramkampella arabica Assemblage of central Crimea
is most similar to Berriasian foraminiferal assemblages
STRATIGRAPHY AND GEOLOGICAL CORRELATION
169
of Syria, the northern Caspian region, southeastern
France, and Italy. The assemblages from the middle
and upper parts of Berriasian sections in central
Crimea are best comparable with their counterparts
from the “Cenozone D” of Madagascar (Espitalie and
Sigal, 1963).
Thus, the foraminiferal assemblages allow successful subdivision and correlation of sections within particular regions, while their potential in interregional
correlations without ammonites is very low.
Ostracods. The discovery of previously unknown
parts of the Berriasian section in central Crimea in 2012
substantially widened the range and characteristics of
the previously defined Costacythere khiamii–Hechticythere belbekensis Beds (Arkadiev et al., 2012). The
identified ostracod remains belong to 16 families. Their
assemblage includes 85 species of 33 genera in total.
The core of their assemblages is represented by smoothwalled forms characterized by a wide facies and stratigraphic distribution (Cytherella) and abundant representatives of the tropical (subtropical) genus Cytherelloidea. Of interest is the presence of brackish- and freshwater genus Cypridea, which is characterized by rare
specimens. Ornamented forms are mostly represented
by genera of the families Protocytheridae (Protocythere,
Reticyhere, Hechticythere, Costacythere) and Cytheruridae (Eucytherura) (Plate III).
The taxonomic and quantitative analyses of ostracod
assemblages allow two biostratigraphic units of the beds
rank to be defined in the examined Berriasian section
(Fig. 6). The lower part of the section conditionally correlated with the ammonite occitanica Zone contains
45 ostracod species belonging to 24 genera. It is united
into the Costacythere khiamii–Hechticythere belbekensis Beds on the basis of the co-occurrence of characteristic species and high abundance of Costacythere khiamii specimens.
The ostracod assemblage from the upper part of the
section corresponding to the part of the ammonite
boissieri Zone numbers 71 species of 28 genera. It is
characterized by the dominant role of the genera
Cytherella, Cytherelloidea, Paracypris, Costacythere,
and Reticythere. Many species are inherited from
underlying sediments (36 species of 20 genera in common), while others appear at this level for the first time
(35 species of 22 genera). This part of the section is
attributed to the Costacythere drushchitzi–Reticythere marfenini Beds on the basis of high abundance
and joint occurrence of these characteristic species.
The ostracod species registered in the examined section are mostly known from Lower Cretaceous (Berriasian–Hauterivian) deposits of Crimea (Neale, 1966;
Tesakova and Rachenskaya, 1996a, 1996b), the North
Caucasus (Kolpenskaya, 2000), Central Asia (Andreev,
1986), England (Neale, 1962, 1967, 1978; Slipper,
2009), France (Atlas…, 1985; Donze, 1964, 1965; etc.),
Germany (Triebel, 1938; Gründel, 1964; etc.), and
Poland (Kubiatowicz, 1983). Acrocythere diversa
Donze and Bythoceratina variabilis Donze are first
described from Berriasian strata of France (Donze,
Vol. 23
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STRATIGRAPHY AND GEOLOGICAL CORRELATION
Stage
Zone
Vol. 23
No. 2
2015
Berriasian
jacobi
boissieri
Subzone,
beds
with fauna
(Arkadiev
et al.,
2012)
Beds with
Zeillerina
baksanensis
?
Beds with
Symphythiris
arguinensis
Riasanites
crassicostatum
Neocosmoceras
euthymi
Beds with Malbosiceras chaperi
?
grandis
Dalmasiceras
tauricum
occitanica
Textularia crimica–Belorussiella taurica
Lenticulina Conorandromede boi des
hofkeri
Foraminiferal beds/
assemblages
?
Triplasia emslandensis
acuta
Kuchki
Bechku
28 30
27 15
26 30
25 15
24 7.6
23 8.4
22 5.5
21 18
20 8
17 0.3
19 10
18 5
16 1.65
15 3.6
14 1.5
13 2
11–12 1
10 2.8
9 11.5
7
7–8
6 6
5
4 2.7
3 63
2 12
50
1 4.5
Formation, sequence
Member
Thickness, m
Lithology
1 2 3 4 5 6 7 8 9 1011121314151617181920212223242526272829303132333435363738394041424344 45464748 49 50 51 525354 5556 57585960616263646566676869707172 73 74757677 787980 81 82 83 84
4
6
8
1314
23242526
2122
20
1516171819
121314
21
24
293031 33
394041
394041
424344 4546
4748 49 50
51 52
53545556
57585960616263646566676869707172 73 74757677 7879 80 81 82 83 84
ARKADIEV et al.
272829303132333435363738
Gaps in sampling, m
Stomatostoecha enisalensis
Trocholina elongata
Trocholina alpina
Bramkampella arabica
Ammobaculites inconstans gracilis
Melathrokerion spirialisilis
Bulbabaculites inconstans
Rectocyclammina recta
Pseudocyclammina sp. (cf. P. lituus)
Moechlerina basilinensis
Belorussiella taurica
Charentia evoluta
Everticyclammina virguliana
Rectocyclammina ex gr. chouberti
Nautiloculina oolithica
Protopeneroplis ultragranulatus
Pseudosiphoninella antiqua
Feurltillia frequens
Planularia madagascariensis
Trocholina molesta
Amijiella amiji
Trocholina giganta
Astacolus ambanjabensis
Pseudocyclammina spharoidalis
Trocholina burlini
Astacolus hamililis
Astacolus favoritus
Astacolus inspissatus
Triplasia minima sp. nov.
Stomatostoecha rotunda
Haplophragmium subaequale
Everticyclammina elongata
Astacotys mutilatus
Discorbis infracretaceus
Astacolus calliopsis
Dentalina nana
Lenticulina macra
Lenticulina neocomina
Lenticulina ex gr. subalata
Lenticulina sp. 2 (Gorb., 1978)
Stomatostoecha compressa
Discorbis miser
Astacolus ex gr. proprius
Lenticulina aff. andromede minima
Discorbis crimicus
Lenticulina aff. akmetchetica
Trocholina infragranulata
Triplasia elegans
Flabellammina lidiae
Alveosepta jaccardi
Lenticulina sp. 1 (Gorb., 1978)
Lenticulina muensteri
Marginulina striatocostata
Saracenaria provoslavlevi
Dorothia ex gr. oxycona
Ramulina aculeata
Vaginulina sp. 2385 E et S? 1963
Marginulinopsis sigali
Nodosaria raristriata
Saracenaria aculata
Lenticulina aff. ambanjabensis
Lenticulina aff. uspenskajae (minima)
Lenticulina fracta
Saracenaria provoslavlevi minima
Astacolus proprius
Pseudonodosaria diversa
Dentalina communis
Pseudonodosaria mutabilis
Saracenaria tsaramandrosoensis
Frondicularia complexa
Saracenaria compacta
Lenticulina colligoni–protodecimae
Vaginulina kochii
Planularia crepidularis
Tristix acutangulus
Citharina flexuosa
Planktonic forms
Saracenaria latruncula
Spirillina kubleri
Dorothia kummi (minima)
Tristix acutangulus micrus
Trocholina micra
Lenticulina ex gr. nimbifera
Discorbis praelongus
170
22
17
4
15
9
15
5.6
2
45
65
8 9 1011
10
50–
60 1 2 3 4 5 6 7
Fig. 5. Stratigraphic distribution of main foraminiferal species in Berriasian sections of central Crimea. For legend, see Fig. 4.
6.25
Lenticulina
muensteri
Quadratina
tunassica
Evirticyclammina virguliana,
Rectocyclammina recta,
Bramkampella arabica
Bedenekyr
STRATIGRAPHY AND GEOLOGICAL CORRELATION
Stage
Zone
Vol. 23
No. 2
2015
Berriasian
jacobi
occitanica
boissieri
Subzone,
beds
with fauna
(Arkadiev
et al.,
2012)
Beds with
Zeillerina
baksanensis
?
Beds with
Symphythiris
arguinensis
Riasanites
crassicostatum
Neocosmoceras
euthymi
Dalmasiceras
tauricum
Beds with Malbosiceras chaperi
?
grandis
Textularia crimica–Belorussiella taurica
Foraminiferal beds/
assemblages
?
Lenticulina Conorandromede boides
hofkeri
Kuchki
Triplasia emslandensis
acuta
Bechku
28 30
27 15
26 30
25 15
24 7.6
23 8.4
22 5.5
21 18
19 10
11.5
20 8
9
18 5
17 0.3
16 1.65
15 3.6
14 1.5
13 2
1
10 2.8
11–12
2.7
7
6 6
7–8
5
4
3 63
50
4.5
2 12
1
6.25
Lenticulina
muensteri
Quadratina
tunassica
Evirticyclammina virguliana,
Rectocyclammina recta,
Bramkampella arabica
Bedenekyr
Formation, sequence
Member
Thickness, m
Lithology
22
17
4
15
9
15
5.6
2
45
65
10
50–
60
123
119120121122
106107108109110111112113114115116117118
98 99 100 101102103104105
90 91 92 93 94 95 96 97
128
125126127
124
130131132
129
133
138 139140141142
135 136 137
134
Abundance of species:
1–10 specimens in sample
10–50 specimens in sample
over 100 specimens in sample
Occurrence of species:
permanent
discrete
85 86 87 88 89 90 91 92 93 94 95 96 97 98 99 100101102103104105106107108109110111112113114115116117118119120121122123124125126127128129130131132133134135136 137138139140141142
85 86 87 88 89
Fig. 5. (Contd.)
Gaps in sampling, m
Dentalina gracilis
Dentalina guttinfera
Marginulina micra
Citharinella pectinatimornata
Hoglundina ex gr. caracolla
Globulina fusica
Lenticulina colligoni
Astacolus incurvatus
Globulina prisca
Astacolus laudatus
Frondicularia caspidiata
Tristix crassa
Pseudosaracenaria truncata
Quadratina tunassica
Lenticulina protodecimae
Lagena sztejnae
Astacolus sp. 1
Nodosaria sceptium
Textularia densa
Nodosaria paupercula
Textularia crimica
Verneuilina angularis
Citharina rodocostata
Rhizammina individa
Lenticulina singulare
Astacolus planiusculus
Saracenaria valanginiana
Planularia aff. plana
Discorbis agalarovae
Textularia notha
Marssonella pseudocostata
Saracenaria inflata
Pseudolamarkina reussi
Lamarckina (?) asteriaformis
Recurvoides ex gr. paucus
Haplophragmoides subchapmani
Lenticulina nodosa
Triplasia emslandensis acuta
Lenticulina sp. 3 f
Lenticulina nuda
Tristix insignus
Lenticulina guttata guttata
Istriloculina rectoangularia
Lenticulina andromede
Lenticulina ex gr. ouachensis
(cf. L. o. inchoata)
Tristix valanginica
Lenticulina praegaultina
Falsopalmula costata
Istriloculina fabaria
Conorboides hofkeri
Dentalina pseudodebilis
Tristix acutangulus
Miliospirella caucasica
Epistomina ornata
Dentalina marginuloides
Discorbis agalarovae
Dorothia kummi
Epistomina tenuicostata
NEW DATA ON BERRIASIAN BIOSTRATIGRAPHY, MAGNETOSTRATIGRAPHY
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172
ARKADIEV et al.
1964). Metacytheropteron sp. A Pok., Quasigermanites
bicarinatus moravicus Pok., Eucytherura trinodosa Pok.,
and two close species Eucytherura ex gr. trinodosa Pok.
and Eucytherura aff. soror Pok. are identified in Tithonian sections of the Czech Republic (Pokorny, 1973).
Of these species, E. trinodosa, E. ex gr. trinodosa, and
E. aff. soror were found in the upper Tithonian–lower
Berriasian section of eastern Crimea (Arkadiev et al.,
2012). The subspecies Quasigermanites bicarinatus
moravicus Pok. was previously recorded in the upper
part of the Berriasian section of eastern Crimea, while
the close species Quasigermanites aff. bicarinatus Pok.
was documented in the middle part of the Berriasian
section in southwestern Crimea. Neocythere dispar
Donze was first described from the basal part of the
Valanginian Stage in the stratotype region of the Berriasian Stage (Donze, 1965) and subsequently registered in Berriasian layers of Mangyshlak (Andreev and
Oertli, 1970). We found this species in the upper and
middle–upper parts of the Berriasian section in the
central and western parts of the Crimean Peninsula,
respectively. Many ostracod species were first
described from the Berriasian section of central
Crimea: Cytherella krimensis Neale, C. lubimovae
Neale, Cytherelloidea flexuosa Neale, C. mandelstami
Neale, Bairdia menneri Tes. et Rach., B. kuznetsovae
Tes. et Rach., Cypridea funduklensis Tes. et Rach.,
Pontocyprella nova Neale, Pontocypris cuneata Neale,
Neocythere pyrena Tes. et Rach., Costacythere khiamii
Tes. et Rach., C. drushchitzi (Neale), C. andreevi Tes.
et Rach., C. foveata Tes. et Rach., Hechticythere belbekensis Tes. et Rach., Reticythere marfenini (Tes. et
Rach.), Eocytheropteron sp. A Neale (Neale, 1966;
Tesakova and Rachenskaya, 1996a, 1996b).
The defined ostracod assemblages of central
Crimea exhibit the most similarity to their assemblage
from the Berriasian stratotype (13 genera and 2 species
in common) (Grekoff and Magne, 1966; Neale,
1967). The similarity at the species level is noted to the
Berriasian assemblage from the section cropping out
along the Urukh River in the North Caucasus (10 genera and 7 species in common) (Kolpenskaya, 2000).
Nevertheless, a reliable correlation between ostracod
beds defined in central Crimea and coeval stratigraphic units in the Urukh section is impossible. The
assemblage of the Costacythere khiamii– Hechticythere belbekensis Beds is comparable with the similar Berriasian assemblage from the Berriasian tauricum Subzone defined in southwestern Crimea
(Bel’bek River basin) (Arkadiev et al., 2012).
It should be noted that the ostracod assemblage
from the upper part of Section 2952 (Sample 49-9-1)
is similar in its taxonomic composition and abundance
of Cytherella lubimovae, Costacythere khiamii, and
C. foveata to the assemblage from Section 2949 (Sample 39-2-1).
Palynomorphs. In total, 28 samples were subjected
to the palynological analysis.
The samples were treated in accordance with the
traditional technique used in palynological investigations, which is based on the hydrofluoric method and
modified technology (Raevskaya and Shurekova,
2011). No palynomorphs are observed in 12 samples.
Other samples contain variable quantities of spores,
pollen, and microphytoplankton represented by wellpreserved cysts of dinoflagellates, prasinophytes, and
acritarchs (Plate IV).
The proportions of palynomorphs in different parts
of the section are variable. In the lower part of the
ammonite Dalmasiceras tauricum Subzone, the
palynomorph assemblage is represented by Classopollis
pollen (47%), spores and bisaccate coniferous pollen
(1%), and marine microphytoplankton (52%). In the
remaining part of the section, Classopollis pollen constitutes up to 90% and spores + bisaccate coniferous
pollen are 1–5%. The share of microphytoplankton
varies from 15% in the Malbosiceras chaperi Beds to
Plate III. Ostracods from Berriasian sediments of central Crimea.
(1) Cytherella krimensis Neale, specimen 2/13244, left valve, lateral view, Balki settlement, occitanica Zone; (2) Cytherella lubimovae Neale, specimen 176/13220, left valve, lateral view, Balki settlement, boissieri Zone, Symphythris arguinensis Beds;
(3) Paracypris felix (Neale), specimen 3/13244, left lateral view, Balki settlement, occitanica Zone; (4) Cytherelloidea flexuosa
Neale, specimen 4/13244, left valve, lateral view, Balki settlement, occitanica Zone; (5) Cytherelloidea mandelstami Neale, specimen 5/13244, left valve, lateral view, Balki settlement, occitanica Zone; (6) Eucytherura ex gr. trinodosa Pokorny, specimen 6/13244,
left valve, lateral view, Balki settlement, occitanica Zone; (7) Eucytherura sp. 1, specimen 7/13244, right valve, lateral view, Balki
settlement, occitanica Zone; (8) Eucytherura sp., specimen 8/13244, left valve, lateral view, Balki settlement, occitanica Zone;
(9) Paranotacythere sp., specimen 9/13244, left valve, lateral view, Balki settlement, occitanica Zone; (10) Eocytheroteron sp., specimen 10/13244, left valve, lateral view, Balki settlement, occitanica Zone; (11) ?Furbergiella sp., specimen 11/13244, left valve, lateral view, Balki settlement, occitanica Zone; (12) Neocythere pyrena Tes. et Rach., specimen 212/13220, left valve, lateral view,
Balki settlement, bossieri Zone, euthymi Subzone; (13) Costacythere drushchitzi (Neale), specimen 230/13220, right valve, lateral
view, male, Mezhgor’e settlement, bossieri Zone; (14) Costacythere drushchitzi (Neale), specimen 12/13244, right lateral view,
Balki settlement, occitanica Zone; (15) Reticythere marfenini Tes. et Rach., specimen 13/13244, left lateral view, male, Balki
settlement, bossieri Zone, euthymi Subzone; (16–18) Costacythere khiami Tes. et Rach.: (16) specimen 14/13244;
(17) specimen 15/13244, left lateral views; (18) specimen 16/13244, right valve, lateral view, females, Balki settlement, occitanica
Zone; (19, 20) Costacythere foveata Tes. et Rach.: (19) specimen 17/13244, left valve, lateral view, female; (20) specimen 18/13244,
left valve, lateral view, male, Balki settlement, occitanica Zone; (21) Costacythere andreevi Tes. et Rach., specimen 19.13244, left
valve, lateral view, Balki settlement, occitanica Zone; (22) Hechticythere belbekensis Tes. et Rach., specimen 20.13244, right valve,
lateral view, Balki settlement, occitanica Zone, tauricum Subzone; (23) Cythereis sp. B, specimen 21/13244, right valve, lateral
view, Balki settlement, occitanica Zone; (24) Schuleridea ex gr. juddi Neale, specimen 22/13244, right lateral view, Balki settlement, occitanica Zone.
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NEW DATA ON BERRIASIAN BIOSTRATIGRAPHY, MAGNETOSTRATIGRAPHY
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Plate III
1
2
100 µm
100 µm
100 µm
5
7
30 µm
14
16
15
100 µm
100 µm
12
50 µm
13
17
50 µm
11
50 µm
100 µm
8
100 µm
10
9
100 µm
100 µm
100 µm
6
50 µm
100 µm
4
3
100 µm
100 µm
19
18
20
100 µm
100 µm
21
100 µm
100 µm
22
STRATIGRAPHY AND GEOLOGICAL CORRELATION
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100 µm
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100 µm
2015
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ARKADIEV et al.
5% in the upper part of the ammonite tauricum Subzone and boissieri Zone.
Through the entire section (Fig. 7), spores are represented by smooth grains of Leiotriletes spp., Cyathidites sp., and Schizaleales ferns with the costate (Cicatricososporites sp.), grumous (Verrucosisporites sp.),
and foveate (Klukisporites variegatus Coup.) exine.
The sediments of the Dalmasiceras tauricum Subzone
and boissieri Zone are marked by the appearance of
Pteropsida spores Eboracia torosa (Sach. et Iljina),
Concavissimisporites punctatus (Delcourt et Sprumont), Kraeuselisporites sp., and Microlepidites crassirimosus Timosch. and spores of Gleicheniaceae and
Lycopodiaceae ferns (Lycopodiumsporites sp., Densiosporites velatus Weyland et Krieger).
The pollen spectrum includes grains of Classopollis spp., Piceapollenites spp., Pinuspollenites spp.,
Callialasporites dampieri (Balme), and Quadraeculina anellaeformis Mal.
The Phoberocysta neocomica dinocyst assemblage
defined in the interval of the Berriasian section most
saturated with marine phytoplankton remains (lower
part of the ammonite Dalmasiceras tauricum Subzone) is represented by the following groups:
(1) chorate and proximoshorate cysts of dinoflagellates: Hystrichosphaerina? orbifera (Klement), Systematophora areolata Klement, Kleithriasphaeridium eoinodes (Eisenack), Tanyosphaeridium isocalamum
(Deflandre et Cookson), Dichadogonyaulax? pannea
(Norris), Dichadogonyaulax culmula (Norris), Achomosphaera sp., Cleistosphaeridium varispinosum (Sarjent), Ctenidodinium sp., Epiplosphaera reticulospinosa
Klement, Bourkidinium sp., Sentusidinium spp.;
(2) cavate cysts of dinoflagellates: Scriniodinium
campanula Gocht, Gonyaulacysta sp., Phoberocysta
neocomica (Gocht);
(3) proximate cysts of dinoflagellates: Pseudoceratium cf. pelliferum Gocht, Apteodinium sp., Rhynchodiniopsis martonensis Bailey et al., Rh. cladophora
(Deflandre), Cribroperidinium sp., Nannoceratopsis
deflandrei Evitt subsp. deflandrei, Durotrigia sp.;
(4) prasinophytes of the genus Pterospermella;
(5) acritarchs Micrhystridium sp.
In the upper part of the ammonite Dalmasiceras
tauricum Subzone and in the boissieri Zone, microphytoplankton is represented by cysts of the dinoflagellate
species Systematophora areolata Klement, Epiplosphaera spp., Kleithriasphaeridium eoinodes (Eisenack), Phoberocysta neocomica (Gocht), Rhynchodiniopsis cladophora (Deflandre), Durotrigia sp., and
Oligosphaeridium patulum Riding et Thomas; prasinophytes belonging to the genus Pterospermella; and
diverse acritarchs. Despite the scarcity of the taxonomic composition of microphytoplankton remains
in this interval of the section, it contains species characteristic of the above-mentioned Phoberocysta neocomica dinocyst assemblage from the lower part of the
Dalmasiceras tauricum Subzone. The impoverishment of the assemblage is most likely explained by
changes in depositional environments.
The Phoberocysta neocomica Beds established in
central Crimea are also recognizable in southwestern
and eastern parts of the peninsula (Arkadiev et al.,
2012). The dinocyst assemblage from this unit is correlated with the assemblage from the Berriasian
Dichadogonyaulax bensoni dinocyst zone of France
(Monteil, 1992), upper Riazanian–lower Valanginian
Pheoberocysta neocomica Zone of northwestern
Europe (Fisher and Riley, 1980), and synonymous
dinocyst zone of eastern Canada (Williams, 1975).
The microphytoplankton assemblage documented
in the Malbosiceras chaperi Beds and underlying
strata (Sections 2949, 2948) includes the following
groups: proximate cysts of dinoflagelaltes species Batiacasphaera sp., Cribroperidinium sp., Muderongia simplex Alberti, and Muderongia Complex; chorate and
proximate cysts Kleithriasphaeridium eoinodes (Eisenack), Cometodinium habibi Montail, Systematophora
areolata Klement, Prolixosphaeridium parvispinum
(Deflandre), Prolixosphaeridium spp., Achomosphaera sp., Spiniferites ex gr. ramosus (Ehrenberg);
prasinophytes Pterospermella.
As a whole, the dinocyst assemblage in Berrasian
sediments of central Crimea is characterized by an
impoverished composition. Nevertheless, it is evident
that by its taxonomic composition this assemblage is
close to that from the Phoberocysta neocomica Beds.
DEPOSITIONAL ENVIRONMENTS
The depositional environments of Berriasian sediments were relatively diverse, although as a whole they
Plate IV. Dinocysts from Berriasian sediments of central Crimea.
All the specimens originate from Berriasian section 2940 near the Balki settlement in central Crimea.
(Figs. 1–5, 8, 10–14, 17–23) occitanica Zone, tauricum Subzone, Member 20, Sample 149/13220; (figs. 6, 7, 9) occitanica
Zone, tauricum Subzone, Member 21, Sample 147/13220; (figs. 15, 16) bossieri Zone, Symphythiris arguinensis Beds, Member 25,
Sample 148/13220.
(1a, 1b) Phoberocysta neocomica (Gocht); (2) Achomosphaera sp.; (3) Tanyosphaeridium isocalamum (Defl. et Cook.); (4, 5) Hystrichsphaerina? orbifera (Klement); (6) Oligosphaeridium patulum Riding et Thomas; (7) Epiplosphaera gochti (Fens.); (8) Cleistosphaeridium varispinosum (Sarjent); (9) Epiplosphaera ?areolata (Klement); (10) Kleithriasphaeridium eoinodes (Eisen);
(11) Circulodinium distinctum (Defl. et Cook.); (12) Circulodinium brevispinosum (Pocock); (13) Dichadogonyaulax culmula (Norris); (14) Scrinodinium campanula Gocht; (15, 16) Systematophora sp.; (17) Apteodinium sp.; (18) Ctenidodinium sp.;
(19) Dichadogonyaulax? pannea (Norris); (20) Nannoceratopsis deflandrei Evitt subsp. deflandrei; (21) Tanyosphaeridium sp.;
(22) Micrhystridium sp.; (23) Pterospermella sp.
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NEW DATA ON BERRIASIAN BIOSTRATIGRAPHY, MAGNETOSTRATIGRAPHY
175
Plate IV
1a
1b
2
3
5
6
4
8
9
10
7
13
11
12
14
15
16
17
18
23
19
20
STRATIGRAPHY AND GEOLOGICAL CORRELATION
21
Vol. 23
22
No. 2
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50 µm
STRATIGRAPHY AND GEOLOGICAL CORRELATION
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No. 2
e
r
r
i
a
s
i
a
boissieri
Costacythere drushchitzi–
Reticythere marfenini
Bechku
Subzone,
beds
with fauna
(Arkadiev
et al.,
2012)
Beds with
Zeillerina
baksanensis
Beds with
Symphythiris
arguinensis
Bedenekyr
Riasanites
crassicostatum
Neocosmoceras
euthymi
Dalmasiceras
tauricum
Beds with Malbosiceras chaperi
grandis
Costacythere khiamii–
Hechticythere belbekensis
Ostracod beds
Kuchki
28 30
27 15
26 30
25 15
24 7.6
23 8.4
22 5.5
21 18
19 10
18 5
20 8
17 0.3
16 1.65
15 3.6
14 1.5
13 2
11–12 1
10 2.8
9 11.5
7
6 6
7–8
5
4 2.7
3 63
2 12
50
1 4.5
6.25
Stage
Zone
n
Formation, sequence
Member
Thickness, m
22
17
4
15
9
15
5.6
2
45
Gaps in sampling, m
12
7
Cytherelloidea mandelstami
2
1
4
2?
4j
3?
Paracypris sp.
Patellacythere sp.
Costacythere drushchitzi
2
34
1 4
2 9
2 2 3 13 7 39 1 7
2
13 3 2 2 10 2
2 3 7
5
5 6
10 15
9
3
6
35 1 142
7
2 8
14 10
3 39
7
39 15 6 13 14
1
2
30
3 40 7
25 2 3
6
16 2 5
3 1 4
2
6
1 1 1 13
1
12
1 12 2 3 10
10
9 2
78
37 1
7 10
121
10
1
8 2
11
1 62 4
2
16 15 1 2 1
1
22 10 1 2
1 1
1 1
15
2 2
5
3
10 15
7
7
93
103 77
26 8
6 986 262
5
5
1
1
1
1
1
71
1
1 1
5 1 123
16 2
3
72
15
57
3 2
5
2
4
4
1
6
14
1
30
149 27 1
6
39
32
13
1
2
163
82
340 15? 1 33
2
70
1009 4?
2
Costacythere andreevi
Bythoceratina sp.
Costacythere foveata
Costacythere khiamii
Hechticythere belbekensis
?Fuhrbergiella sp.
Paranotacythere sp.
Cythereis sp. B
Schuleridea ex gr. juddi
Quasihermanites bicarinatus moravicus
Pontocyprella sp.
“Cypridea” sp.
Costacythere sp.
Reticythere marfenini
50 7 7
36 23
50 242
12
52 330
14 13
55 5
220
83 1565
267
3 1
1
260
76
230 11
93
24
3
226 46 4 40 57 1 1
2 23
1
30
24
353
Cytherella lubimovae
Cytherella krimensis
Cytherella fragilis
Cytherella sp.
2
6
11
1 1
131
Protocythere sp. 1
Bairdia sp.
Schuleridea aff. juddi
2
1
8
1
3
Eucytherura aff. trinodosa
2
1
2
Eocytheropteron sp.
7
4
1
39 1
21
3
40 1
1
1
8
40
21
13
8 1
15 1
16
12
4
1
81
13
13
1
1
Cytherelloidea flexuosa
Neocythere sp.
Pontocypris felix
5
Cythereis aff. senckenbergi
Paracypris aff. caerulea
Acrocythere alexandrae
Schuleridea? sp. indet.
2
1 2
4 2
1
8 1
1
3 1 3
2
14 1
1 3
Eucytherura sp. 2
Paracypris aff. parallela
Acrocythere aff. hauteriviana
Pontocyprella nova
15 1 6
6 1 2
1
1 2
3
5
Cypridea funduklensis
Eucytherura aff. soror
Cytherelloidea sp.
9
3 1
1
3
2
1
36 1
42
17 7 1
9
1
1
1 2
14 1
2 10
1 5 11 15 7 1
4 1 1 2
Ostracods
8
2 5 12
1
Eucytherura sp. 1
Metacytheropteron sp. A
Bythoceratina ex gr. variabilis
2 4
3
1
Neocythere pyrena
Pontocyprella elongata
Pontocypris cuneata
Pontocyprella aff. superba
Macrocypris sp.
Pontocyprella aff. nova
Pontocyprella sp. (aff. P. harrisiana)
Paracypris sp. (P. ex gr. elegans)
Neocythere dispar
Pterygocythere? sp.
Eucytherura aff. ardescae
Bairdia menneri
1
1
1
1
1
4
6
3 3
10 2
6 3 2 21 7
3 1 1 3 15 10 1 3 11
1 31 3 22 4 1
3 1
ARKADIEV et al.
65
10
50–
60
Fig. 6. Stratigraphic distribution of main ostracod species in Berriasian sections of central Crimea. For legend, see Fig. 4.
Lithology
Acrocythere aff. alexandrae
Bairdia kuznetsovae
Eocytheropteron sp. A.
Acrocythere diversa
Eucytherura ardescae
Eucytherura paula
Paranotacythere diglypta
Vocontiana sp.
Eucytherura trinodosa
Bythocypris sp. (B. arcuta)
176
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jacobi
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Zone
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Berriasian
occitanica
jacobi
boissieri
Subzone,
beds
with fauna
Beds with
Zeillerina
baksanensis
Beds with
Symphythiris
arguinensis
Riasanites
crassicostatum
Neocosmoceras
euthymi
Dalmasiceras
tauricum
Beds with Malbosiceras chaperi
grandis
Phoberocysta neocomica
Bedenekyr
Kuchki
28 30
27 15
26 30
25 15
24 7.6
23 8.4
22 5.5
21 18
20 8
17 0.3
19 4
18 5
16 1.65
15 3.6
14 1.5
13 2
1
11–12
10 2.8
9 11.5
6 6
6.25
7–8 7
5
4 2.7
3 63
2 12
1 4.5
50
Dinocysts beds
Bechku
Formation, sequenc
Member
Thickness, m
22
17
4
15
9
15
1
2
45
65
10
50–
60
Gaps in sampling, m
31-12-1
31-8-1
31-7-1
31-5-1
Sample number
Cicatricosisporites sp.
Klukisporites variegatus
Cyathidites spp.
Verrucosispontes sp.
Leiotriletes spp.
Microlepidites crassirimosus
Concavissimisporites punctatus
Gleicheniidites sp.
Contignisporites sp.
Eboracia torosa
Densoisporites velatus
Disaccites gen. gen. spp.
Classopollis spp.
Pollen
Callialasporites dampieri
Quadraeculina anellaeformis
Spores
Lycopodiumsporites sp.
31-3-1
31-2-1
31-1-1
27-1-3
27-1-2
27-1-1
25-11-2
25-11-1
25-9-2
25-9-1
25-7-1
25-3-1
25-2-1
25-1-1
26-1-1
29-1-3
29-1-2
29-1-1
41-9-1
41-7-2
41-7-1
41-5-2
41-5-1
41-3-2
41-3-1
38-4
38-238-3
38-1-2
38-1-1
39-3-1
39-2-1
Kraeuselisporites sp.
Proximate cyst gen. et sp. indet.
Muderongia Complex
<1%
1–10%
>50%
Chorate cyst gen. et sp. indet.
Systematophora areolata
Kleithriasphaeridium eoinodes
Prolixosphaeridium parvispinum
Batiacasphaera sp.
Cribroperidinium sp.
Achomosphaera sp.
Cometodinium habibii
Spiniferites ex gr. ramosus
Phoberocysta neocomica
Rhynchodiniopsis cladophora
Durotrigia sp.
Sentusidinium spp.
Circulodinium distinctum
Ctenidodinium sp.
Systematophora sp.
Gonyaulacysta sp.
Hystrichosphaerina? orbifera
Nannoceratopsis deflandrei subsp. deflandrei
Psendoceratium cf. pelliferum
Apteodinium sp.
Scriniodinium campanula
Tanyosphaeridium isocalamum
Rhynchodiniopsis martonensis
Epiplosphaera reticulospinosa
Dichadogonyaulax? pannea
Dichadogonyaulax culmula
Circulodinium brevispinosum
Cleistosphaeridium varispinosum
Epiplosphaera ? areolata
Oligosphaeridium patulum
Epiplosphaera gochtii
Phallocysta elongata
Tubotuberella sp.
Muderongia sp.
Batioladinium jaegeri
Leptodinium sp.
Dichadogonyaulax sellwoodii
Dinocysts
Proportions of palynomorphs in palynological spectra:
Fig. 7. Stratigraphic distribution of palynomorphs in Berriasian sections of central Crimea. For legend, see Fig. 4.
Lithology
Pterospermella spp.
Prasinophytes
Micrhystridium spp.
Acritarchs
NEW DATA ON BERRIASIAN BIOSTRATIGRAPHY, MAGNETOSTRATIGRAPHY
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178
ARKADIEV et al.
were always associated with development of the carbonate platform.
The lower part of the section (Members 1–18)
demonstrates a relatively uniform composition of calcareous sediments in both the lateral and the vertical
direction. The presence of clay and marlstone intercalations and the absence of features indicating extremely
shallow areas, landslides, and accumulation of detrital
sediments imply the formation of this part of the section
in settings of an almost flat homoclinal ramp.
The examined limestone varieties belong to two
ramp microfacies (RMF; Flügel, 2010):
(1) RMF 3 (Members 1–4, 18) is represented by
wacke- and packstones with skeletal detritus bioturbated by crustaceans (Thalassinoides, Plate V, fig. 1).
The presence of micrite, skeletal detritus, and ooids
indicates calm to moderate hydrodynamics. This
microfacies is characteristic of the middle and/or
outer ramp.
(2) RMF 9 (Members 8, 9, 14, 16) comprises pack-,
wacke-, and floatstones with bioclasts and intraclasts
of the ramp, rare thalassinoid burrows, and lenses of
shelly floatstones (SMF 8 after Flügel, 2010). This
microfacies characterizes the middle and/or outer
ramp and calm to moderate hydrodynamics. Skeletal
detritus in both microfacies represents typical material
redeposited from inner parts of the ramp.
The rare sandstone intercalations (Members 11,
12, Plate V, figs. 5, 6) with well-developed trough to
horizontal bedding locally disturbed by crustacean
Ophiomorpha burrows (Plate V, fig. 5) were probably
deposited in tidal channels (Baraboshkin, 2011). The
effect of high-energy currents is evident from the presence of oyster accumulations with separated valves
(Plate V, fig. 3).
Clays and marlstones (Members 5, 6, 10, 13, 15, 17,
partly 18) were deposited in inner areas of the basin
and/or in depressions; in either case, they indicate
deeper sedimentation settings as compared with depositional environments of limestones. Corals of the genus
Montlivaltia occur frequently in pelitic rocks (Wright
and Burgess, 2005); therefore, their co-occurrence with
belemnites and ammonites in Member 15 implies their
autocthonous nature. At the same time, it is conceivable
that accumulation of some clay members as well as the
entire terrigenous part of the section was stimulated by
climate humidization, which explains the presence of
carbonaceous detritus, sand admixture, and coquinas.
Thus, the lower calcareous part of the section
(Members 1–4) was accumulated in environments of
the middle and/or outer ramp, while its overlying part
dominated by clays and marlstones was deposited in
deeper settings of inner areas of the basin or under climate humidization. Taking into consideration the
insignificant lateral variability of sediments in outer
parts of the ramp (Tucker and Wright, 1990; Flügel,
2010) and low probability of the replacement by carbonate facies, the vertical replacement of limestone by
clays resulted from transgressive processes or climate
humidizatiuon. The basin deepening is consistent with
the transgressive part of the megacycle in other regions
corresponding to the jacobi and occitanica phases
(Ogg and Hinnov, 2012).
The overlying part of the section (Members 19–24)
is largely represented by the terrigenous succession. Its
basal layers are composed of bioturbated glauconite–
quartz carbonate sandstones grading up the section
into clays with abundant small pyritized shells of
ammonites and other faunal remains. This trend most
likely reflects the basin deepening stage.
The appearance of the sponge horizon (Member 25)
reflects the change in the basin dynamics. The absence
of features indicating the dynamic influence of water
and the presence of a significant mud component in
sediments indicate that the sponge horizon was
formed at depths exceeding 50 m. The growth of Spongia bioherms proceeds usually under the influence of
high-energy bottom currents, which transport food
particles and nutrients. Such bioherms grow near the
slope bends; it is conceivable that, this area corresponded to the middle–lower ramp transition or was
slightly deeper.
The overlying section (Members 26–28) demonstrates a well-expressed shoaling trend, which is also
consistent with the transgressive–regressive megacycle
of the boissieri phase (Ogg and Hinnov, 2012). The
regressive features are already evident in the upper part
of Member 25 (Plate V, fig. 7), which was formed in
relatively shallow environments of the warm basin.
However, the substantial share of silty–sandy terrigenous admixture prevented accumulation of pure carbonate sediments in this zone. At the same time, the
presence of Ophiomorpha burrows implies an unconsolidated mobile substrate. The higher carbonate content in Members 26–27 and the presence of abundant
remains of diverse normal-marine organisms, including oysters, brachiopods, and echinoderms, indicate
open basin (ramp?) settings with the dominant role of
tidal currents. The section is crowned by Member 28,
represented by coral–rudist–algae limestones
Plate V. Fragments of the section structure and some ichnofossils in Berriasian sediment.
(1) Thalassinoides suevicus (Rieth) burrow, Member 3; (2) Gyrolithes sp. burrow, presumably Members 10–12; (3) accumulation of
oyster Pycnodonte weberae Yanin shells, Member 12; (4) limestones entirely bioturbated by Thalassinoides burrows, Member 2;
(5) horizontally bedded sandstones with Ophiomorpha sp. (arrow), Member 12; (6) trough cross bedding in sandstones, some layers
are enriched with bioclasts, Member 12; (7) outcrops of Member 25 near the Pasechnoe settlement; (8) outcrops of Member 28 (biohermal limestones) in the quarry of the Pasechnoe settlement, the top of the quarry wall corresponds to the presumable Berriasian–
Valanginian boundary. (Figs. 1–6) Balki settlement area, photo by V.K. Piskunov, 2012; (figs. 7, 8) photo by E.Yu. Baraboshkin,
2002.
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NEW DATA ON BERRIASIAN BIOSTRATIGRAPHY, MAGNETOSTRATIGRAPHY
179
Plate V
1
2
3
4
5
6
7
8
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180
ARKADIEV et al.
(Plate V, fig. 8), which were deposited on the coastal
shoal raised intermittently above the sea level.
The Berriassian sedimentation stage terminates
with desiccation of the carbonate platform, which was
accompanied by karst processes and reorganization of
the sedimentary system: in the Valanginian, terrigenous sedimentation became prevalent.
Most identified ostracod taxa are characteristic of
basins with normal salinity. Single representatives of
their genera able to resist salinity variations or dwell in
brackish-water to fresh-water environments do not
effect on in their general distribution (Morkhoven,
1963; Prakticheskoe…, 1999 ). The results of the paleoecological analysis of ostracod assemblages are also
consistent with presumable basin deepening (Members 5–18) and shoaling (Member 26) and confirm the
conclusion on deposition of the sponge horizon
(Member 25) at depths exceeding 50 m.
The palynological data (Fig. 7), including abundant Classopollis spp. pollen (up to 90% in spectra)
produced by Cheirolepidaceae plants and dominant
role of chorate dinocysts with long processes, imply
warm (tropical) environments.
MAGNETOSTRATIGRAPHY
Petromagnetic and Magnetic–
Mineralogical Investigations
The petromagnetic and magnetic–mineralogical
investigations included the study of magnetic susceptibility (K) and its anisotropy (AMS), measurement of
remanent magnetization (Jn), experimental magnetic
saturation with subsequent determinations of remanent
magnetization of saturation (Jrs) and remanent coercive
force (Hcr), and differential thermomagnetic analysis
(DTMA). K was measured on MFK1-FB equipment
(kappabridge), Jn was measured on a JR-6 spin magnetometer, and a TAF-2 thermoanalyzer of fractions
(“magnetic weights”) was used for DTMA. The AMS
analysis was conducted using the Anisoft 4.2 program.
The composite section is well differentiated with
respect to its petromagnetic properties (Fig. 8).
The lower (carbonate) part of the section (Members 1–9) is slightly magnetized (K = 0.2–8 × 10–5 SI
units, Jn = 0.002–0.2 × 10–3 A/m).
The overlying terrigenous sediments (Members 10–
26) are characterized by higher K values increasing up
the section to 25–33 × 10–5 SI units in Members 20–22
and then gradually decreasing to 11–15 × 10–5 SI units
in Member 26. The similar trend in the distribution of
Jn values is complicated by the presence of several intervals with abnormally high values, which results in the
increase in the Konigsberger ratio (Q factor) to several
units, while its background values never exceed fractions of unity. The upper carbonate members (27 and
28) are slightly magnetic (K = 1–3 × 10–5 SI units, Jn =
0.01–0.1 × 10 3 A/m).
The K/Jrs parameter proportional to the average
size of ferromagnetic particles is lowest in clays of the
Bechku Formation (Members 19–24) and highest in
limestones of Members 3 and 4 and terrigenous–carbonate varieties of Members 26 and 27.
Hcr value is highly variable (20–325 × 103 A/m),
being highest in some carbonate varieties of Members 1–4, 9, 27, and 28. Many samples are magnetically soft (saturation is achieved in fields of ~100 ×
103 A/m) (Fig. 9a), which is determined by the presence of magnetite or close minerals. The magnetically hard samples do not achieve saturation in fields
up to 600 × 103 A/m (Fig. 9b), which is explained by
the presence of hematite or strongly dehydrated
Fe hydroxides. On saturation curves obtained for
limestones from the lower part of the section (Members 1–3), both phases (magnetically soft and hard)
are well recognizable (Fig. 9c).
The presence of magnetite is also evident on
DTMA curves. Unfortunately, the peak near 550–
578°C corresponding to the Curie point of Fe3O4 or
close minerals is masked on all of them by thermomagnetic effects related likely to pyrite. The presence
of the latter is evident from the magnetization increment above 400°C owing to transformation of FeS2
into Fe3O4 (Figs. 9d, 9e). The magnetically hard phase
(hematite, martite, or dehydrated Fe hydroxides) is
reflected in the poorly expressed peak on DTMA
curves in the area of 650°C (Fig. 9e).
The ASM patterns are different in carbonate and
terrigenous varieties. Limestones exhibit chaotic magnetic patterns (Fig. 9f), while in clays and sandstones
projections of short axes of magnetic ellipsoids are regularly grouped in the center of the stereographic projection and projections of long axes strive to the equator and are marked by slight but distinct anisotropy
along the sublatitudinally oriented line that corresponds with the strike of beds (Fig. 9g). In order to
exclude suspicions that chaotic AMS patterns (Fig. 9f)
are determined by the instrumental error in measurements of slightly magnetic limestones (mostly К < 3 ×
10–5 SI units), we have analyzed two selections of samples with the measurement errors of <5% and >5%,
respectively. The testing revealed that the magnetic
patterns in both selections are the same.
The anisotropy of magnetic susceptibility in limestones is most likely determined by the irregular distribution of ferromagnetic minerals due to their concentration in bioturbations, which are characteristic of
carbonate rocks in the examined section. The enrichment of ichnofossils with ferromagnetic minerals is
explained by the concentration of biogenic magnetite
in many crustacean organisms (Biskirk and O’Brian,
1989); in addition, some burrows become populated
by magnetite-producing bacteria (Stolz et al., 1986).
Bioturbation of sediments provides no obstacle for
paleomagnetic investigations since ferromagnetic particles remain oriented in accordance with the field in
semiliquid sediments acquiring the post-depositional
detrital remanent magnetization Jn.
The anisotropy of long axes of magnetic ellipsoids
in the terrigenous part of the section indicates com-
STRATIGRAPHY AND GEOLOGICAL CORRELATION
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2015
Member
Thickness, m
Gaps in
sampling, m
Polarity
K, 10–5 SI units
0
10 20 30
Jn, 10–3 A/m
K/Jrs, 10–5 m/A
0 1 2 3 4 5 6 7
–0.05 0
2420
2943
28 30
27 15
26 30
3
?
25 15
22
24 7.6
2940
17
23 8.4
15
19.96
21.84
40.97
22 5.5
4
21 18
2952
11–12
2948
3.6
1.5
2
1
10 2.8
9 11.5
45
4 2.7
2949
6
5
65
21.2
0.50
0 50 150 250 350
Jrs, 10–3 A/m
Q
0
1
2
3
0
1000
2000
2947
6
6.25
3 63
2 12
10
50–
60
181
>50
1 4.5
2950
2949
7–8 7
2947
2015
17 0.3
1.65
16
15
14
13
20.3
15
5.6
2
2952
2944
19 4
18 5
2950 2951
No. 2
9
20 8
0.25
Hcr, 103 A/m
NEW DATA ON BERRIASIAN BIOSTRATIGRAPHY, MAGNETOSTRATIGRAPHY
Vol. 23
Fig. 8. Composite magnetostratigraphic Berriasian section of central Crimea: paleomagnetic and petromagnetic characteristics.
For legend, see Fig. 4.
STRATIGRAPHY AND GEOLOGICAL CORRELATION
Outcrops
182
ARKADIEV et al.
pression of rocks in the submeridional direction,
which is consistent with widely shared views on geodynamics of the Crimean Peninsula at the neotectonic
stage (Nikishin et al., 1997). In contrast to hard limestones, clays and clayey sands changed their magnetic
structure in response to collisional compressions. At
the same time, judging from the vague anisotropy of
long axes, deformations which they experienced could
not have resulted in significant distortion of Jn vectors
able to affect polarity determination.
An atypical AMS distribution is documented in
outcrop 2952 represented by alternating carbonate
and terrigenous rocks. As in the remaining section,
limestones in this outcrop are characterized by chaotic
magnetic patterns. At the same time, projections of
short axes in clays (Members 13, 15, 17) are displaced
from the center of the stereographic projection and are
arranged along a large circle in the SE–NW direction,
while long axes are distributed transversely in the SW–
NE direction (Fig. 9h). Such magnetic patterns indicate that these rocks were subjected to extremely
intense deformations due to local compression along
the SE–NW axis (Lanza and Meloni, 2006), which
was probably accompanied by the formation of the
upthrown (thrust) structure. This conclusion is indirectly confirmed by development of cleavage in limestones (Member 18) and poor quality of the paleomagnetic record in this outcrop. In such a situation, the
data on this interval of the section, including documented succession of layers, should be taken with precaution. It is conceivable that limestones with intense
cleavage represent an exotic block (klippe) of older
strata, which is consistent with the paleontological
data: the foraminiferal assemblage from Member 18
differs from that in underlying terrigenous rocks, being
identical to the assemblage in older limestones.
Similar AMS patterns are observed in clays, siltstones (upper part of Member 25 and Member 26), and
marlstones (Member 27) cropping out in the Mezhgor’e and Pasechnoe areas (outcrops 2943, 2420): projections of short axes of magnetic ellipsoids demonstrate as in outcrop 2952 a tendency for the shift along
the large circle, although they are less remote from the
center of stereographic projection (Fig. 9i). This indicates that unconsolidated plastic sediments are
deformed by compression in the SE–NW direction (as
in outcrop 2952, but to a lower degree).
Thus, the AMS investigations make it possible to
obtain nontrivial information on intensity of tectonic
movements, which affected rocks in the region under
consideration, and, in fact, to specify the structure of
the Berriasian section in central Crimea. At the same
time, outcrops 2952, 2943, and 2420 (Members 13–
18, 26–28) appeared to be unfavorable objects for
paleomagnetic investigations. As a whole, variations in
petromagnetic parameters through the composite section promote individualization of its particular intervals and recognition of sedimentation cyclicity
(Fig. 8), which is of undoubted interest for substantiation of stratigraphic units and paleogeographic reconstructions.
Paleomagnetic Investigations
The paleomagnetic investigations were aimed at
obtaining magnetic polarity characteristics of the section. Each of 181 oriented samples taken from different stratigraphic levels (Figs. 3, 4) was cut into three
or four cubes with edges of 20 mm. The laboratory
investigation of the samples included Jn measurements on the JR-6 spin magnetometer after a series of
successive magnetic demagnetization sessions by an
alternating field mostly up to 45–60 mT with a step of
5 mT (H demagnetization) on LDA-3 AF equipment
and temperature ranging from 100°C to 500–550°C
with a step of 50°C (T° demagnetization) in an
Aparin furnace. The samples were subjected to the
impact of high fields and temperatures until their
magnetization became comparable with the instrumental measurement accuracy. For the control over
the possible laboratory magnetization of samples, two
cubes from the same sample with mutually opposite
orientation along two Jn constituents were placed into
the furnace. For the control over the quality of results,
some samples were measured on the cryogenic magnetometer (2G Enterprises) in the Paleomagnetic
Laboratory (Institute of Petroleum Geology and
Geophysics, Siberian Branch, Russian Academy of
Sciences, Novosibirsk). The data were processed
using the Remasoft 3.0 program. The natural remanent magnetization (Jn) retained after the impact of
strong fields and high temperatures was accepted as
the stable component of magnetization (SCM)
(Figs. 10a, 10b).
Carbonate rocks of the basal part of the section
appeared to be most favorable for paleomagnetic
investigations. In samples from these rocks, the stable
components of magnetization projected onto the
upper hemisphere are usually defined with the appropriate accuracy (maximum deviation angles up to 15°)
after both demagnetization procedures (Fig. 10a). The
results of thermal demagnetization of samples from
Members 1–3 yield better paleomagnetic statistics as
compared with that for backup samples, where Jn was
Fig. 9. Results of the magnetic–mineralogical analysis. (a–c) Curves of magnetic saturations; (d, e) DTMA curves (integral
curves and first derivatives according to thermomagnetic analysis curves); (f–j) distributions of directions for axes of anisotropy
of magnetic susceptibility (in the stratigraphic coordinate system) for limestones (f) and terrigenous rocks (g) from outcrops 2952 (h)
and 2943 (i). DTMA was conducted simultaneously for several lithologically and magnetically similar samples. (1, 2) Long (K1)
and short (K3) axes of ellipsoid of anisotropy of magnetic susceptibility, respectively.
STRATIGRAPHY AND GEOLOGICAL CORRELATION
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2015
Jrs, 10–3 A/m
80
(b)
400
500 600
H, 103 A/m
2949-4
(c)
0 100 200 300 400 500 600
H, 103 A/m
–3
Jrs, 10 A/m
2950-3
30
0
–2
N = 220
270
90
2
1
0
–2
–3
–6
–5
(e)
3
combined samples
2947-3, 10
J/
Ts
eco
nd
2
he
ati
ng
ses
sio
n
10
8
6
4
1
2
100 200 300 400 500 600 700
–1
–2
0
–1
–3
dJ/
dT
–2
sec
ond
hea
ting
sess
ion
180
–3
N
N
(g)
N
(h)
(i)
N = 273
270
–1
–4
A/m (first heating session)
3
0
–5
ssion
ating se
first he
dJ/dT
(f)
se
ssi
on
–4
J, 10
4
N
he
at
in
g
–7
0 100 200 300 400 500 600
H, 103 A/m
(first heating session)
1
2
se
co
nd
–7
2947-10
dJ/dT, 10–7 A/m
300 200 100
dT
–3
20
10
100 200 300 400 500 600 700
dJ
/
session
200 100
0.5
1.0
–1
(first heating session)
40
dJ/dT, 10–7 A/m
2943-11
1.5
dJ/dT, 10–6 A/m
300
(second heating session)
200
1.0
dJ/dT, 10–7 A/m J, 10–7 A/m (second heating session)
0 100
2.0
1.5
se
co
fir
nd
st h
he
ea
ati
tin
ng
gs
ess
se
ssi
ion
on
heating
session
2943-5
J/
T
combined samples
2940-21, 22, 25
J/
T
J/T firs
t
40
2.5
st heating
dJ/dT fir
J, 10–6 A/m (first heating session)
2952-1
100
2.0
(d)
(second heating session)
Jrs, 10–3 A/m
80
(a)
183
J, 10–5 A/m (second heating session)
NEW DATA ON BERRIASIAN BIOSTRATIGRAPHY, MAGNETOSTRATIGRAPHY
N = 28
90 270
180
STRATIGRAPHY AND GEOLOGICAL CORRELATION
N = 59
90 270
180
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ARKADIEV et al.
destroyed by the alternating field (table). The relatively high precision parameter and steeper (than after
H demagnetization) paleomagnetic inclination are
determined by the fact that magnetization related to
hard ferromagnetic minerals is destroyed by temperature more effectively. This is evident from the correlation between the precision parameter and Hcr values in
results of demagnetization by the alternating field: in
the magnetically softest samples, precision parameters
are the highest and paleomagnetic directions correspond to the vector derived from thermal demagnetization sessions, the precision parameter decreases
with addition of magnetically harder samples to the
selection, and the average paleomagnetic inclination
becomes gentler (table).
In terrigenous varieties, the reliability of paleomagnetic measurements is worse than in limestone,
although the K and Jn values in them are substantially
higher. No stable component was defined in approximately one-third of terrigenous samples. The remaining samples may be divided into two groups. In samples of the first group, Jn projections either were initially located on the northern sector of the lower
hemisphere or left it after the session of demagnetization by weak temperature and alternating field. Such
behavior of paleomagnetic vectors during demagnetization was interpreted as the presence of the Jn component corresponding to the reversed polarity (R). In
some sample, SCM is anomalous: for example, southerly declination combined with positive inclinations in
Sample 2940-30 (Fig. 10c). In samples of the second
group, projections of paleomagnetic vectors remained
in the northern sector of the lower hemisphere up to
the last session of demagnetization (Fig. 10b). These
directions are interpreted as corresponding to normal
polarity of the magnetic field (N).
We believe that the significant scatter of R vectors
(Figs. 10e, 10f) is determined by the different degree
of SCM “contamination” with the stabilized secondary component related to products of oxidation
of magnetic grains and by impossibility to separate
the components during demagnetization. This effect
leaves the paleomagnetic statistics on normally magnetized samples practically unchanged (Fig. 10g,
table) since SCMs corresponding to normal polarity
are close to the direction of rock magnetization
reversal by the recent field, although statistically
being different (table).
It is of importance that the results of the sample
magnetization reversal by the alternating field and
temperature demonstrate principal similarity
(Figs. 10c, 10d). This increases substantially the reliability of paleomagnetic measurements as compared
with the results based only on one of the demagnetization procedures. Nevertheless, the anomalous
reversed polarity is shown as the half-shaded paleomagnetic column in Figs. 3 and 4. The rejected and
unreliable polarity determinations are scattered more
or less regularly through the section; therefore, the
absence of information hardly affects its paleomagnetic structure.
Small gaps in polarity determinations and single
intervals with its opposite sign were ignored during
compilation of the composite paleomagnetic column
(Figs. 2, 11). Owing to the biostratigraphic control,
identification of the paleomagnetic column with successive magnetic chrons was successful (at least for
Members 1–23) despite large unexposed intervals and
doubts in the reliability of results obtained for some
intervals of outcrops 2952, 2943, and 2420. The long
R magnetic zone corresponding to the largest part of
the occitanica Zone (and, probably, uppermost jacobi
Zone) and N magnetic zone in the uppermost part of
the occitanica Zone represent undounbted analogs of
Chron M17 (M17r and M17n, respectively). The Dalmasiceras tauricum Subzone is an analog of the Dalmasiceras dalmasi Subzone of the Mediterranean
standard and consequently Chron M17n cannot correspond to large gaps in sampling between Members 3
and 4 or 8 and 9 (Figs. 2, 11), which are located considerably below the first finds of Dalmasiceras representatives. The overlying R magnetic zone corresponds to Chron M16r established previously in the
Feodosiya area (Arkadiev et al., 2010) (Fig. 11). The
examined section includes also analogs of Chrons
M16n and M15r (it is conceivable that M15n and
M14r are included as well), although their position
cannot be determined because of large gaps in sampling (Fig. 11).
The obtained data make it possible to carry out
magnetostratigraphic correlation of the Berriasian
section of central Crimea and the Berriasian stratotype
Fig. 10. Results of the component analysis.
(a, b) (From left to right) stereographic images of changes in Jn vectors during demagnetization sessions, Zijderveld diagrams,
demagnetization plots; (c, d) comparison of results of demagnetization by the alternating field (h) and temperature (t); (e–g) stereographic projections of SCM corresponding to R polarity in Members 1–3 (limestones) according to results of T° and
H demagnetization (e), R polarity in Members 5–25 (terrigenous rocks) according to results of T° and H demagnetization (f),
R polarity in Members 1–3 (limestones) according to results of H demagnetization of magnetically softest samples (Hcr = 19.6–
26.8 × 103 A/m) and T° demagnetization, and N polarity in Members 5–25 (terrigenous rocks) according to results of T° and
H demagnetization (g). All the Zijderveld diagrams and Jn stereographic projections are presented in the stratigraphic coordinate
system. (1, 2) Jn projections in stereograms on the lower (1) and upper (2) hemispheres; (3, 4) projections of average SCM directions for all R and N populations of vectors, respectively; (5) projections of directions of rock magnetization reversal by the recent
geomagnetic field (“crosses” of magnetization reversal).
Paleomagnetic statistics for the stereogram (c) and interpretation of statistical parameters are presented in the table.
STRATIGRAPHY AND GEOLOGICAL CORRELATION
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NEW DATA ON BERRIASIAN BIOSTRATIGRAPHY, MAGNETOSTRATIGRAPHY
(a)
N
J/Jmax
1.0
0.8
E 0.6
E 0.4
0.2 Jmax = 35.1e–06 A/m
W
W
0
180
N
Sample 2949-8
S Base
N Top
N
N Top
E
E
W
W
180
N
Sample 2944-5
Sample 2940-30
h
(d)
J/Jmax
1.0
0.8
Jmax = 41.0e–03 A/m
0.6
0.4
0.2
h
t
180
180
N
N
Sample 2943-8
h
t
t
Sample 2940-60
E
E
jr-6
J/Jmax
1.0
0.8
0.6
0.4
Jmax = 1.18e–03 A/m
0.2
0
jr-6
SQUID
Sample 2944-4
SQUID
100 200 300 400 500 600
Division value =
175.e–06 A/m
°C
S Base
180
(e)
t
Sample 2944-7
S Base
N Top
W
W
h
0 10 20 30 40 50 60 70
mT
Division value =
7.89e–03 A/m
t
Sample 2947-10
10 20 30 40 50 60
mT
Division value =
42.9e–06 A/m
S Base
Sample 2940-21
t
Sample 2952-13
J/Jmax
1.0
0.8
0.6
0.4 J = 256.e–06 A/m
0.2 max
0
(b)
h
h
E
E
N
Sample 2950-5
100200300400500
°C
Division value =
5.13e–06 A/m
W
W
180
N
(c)
Sample 2947-10 N Top
185
N
(f)
N
N
(g)
1
n = 40
D = 169.1°
I = –28.5°
k = 1.6
α95 = 29.9°
n = 47
D = 176.3°
I = –41.4°
k = 6.3
α95 = 9.0°
2
270
90
3
4
5
180
180
STRATIGRAPHY AND GEOLOGICAL CORRELATION
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ARKADIEV et al.
in France (Galburn, 1985) (Fig. 11) and combined
with the available data on eastern Crimea (Arkadiev
et al., 2010; Bagaeva et al., 2011; Guzhikov et al.,
2012) allow the statement that analogs of all the Berriasian magnetic chrons are present in Crimea.
The ancient nature of the stable component of
magnetization (SCM) is substantiated by the following arguments.
(1) The direction of geomagnetic polarity is independent of both the lithological composition of rocks
and variations in their petromagnetic properties
(Figs. 2, 8). This does not prove the hypothesis of the
old magnetization nature, but is consistent with the
latter since the geomagnetic reversal is a global phenomenon; therefore the probability of interrelations
between magnetic polarity and lithological–magnetic properties determined by local and regional
factors is negligible.
(2) The data on magnetic patterns of limestones
(Fig. 9f) imply the confinement of the magnetically
soft fraction to bioturbations, indicating the biogenic
nature of magnetite particles, which could not have
been formed later than the diagenetic stage.
The entire section exhibits features characteristic
of detrital magnetization and, in contrast, atypical of
chemical magnetization: low Q factor values (fractions of unity) except for narrow intervals (to 1–5)
sporadically scattered through the section (Fig. 8) and
low paleomagnetic interstratal precision parameters
(up to 30) (table).
The substantiation of the hypothesis of sediment
magnetization prior to diagenesis cessation is identical
to the proof that Jn reflects the direction of the geomagnetic field in the Berriasian.
(3) The most reliable paleomagnetic result
obtained for limestones (table) is statistically identical
to the average paleomagnetic direction available for
the Late Jurassic in western Crimea (Pecherskii and
Safonov, 1993) (it should be noted that, according to
D.M. Pecherskii, “western Crimea” includes the
Ai-Petri Yala, Karabi-Yaila, Demerdzhi, Chatyr Dag,
and Cape Fiolent areas).
(4) The paleomagnetic reversal (inversion) test is
positive (table), which represents a very solid argument in favor of the primary nature of magnetization.
(5) The paleomagnetic zonality of the composite
section is well consistent with traditional views on the
regime of the Berriasian geomagnetic field (Ogg and
Hinnov, 2012) (Fig. 11).
The index of paleomagnetic confidence for the
obtained data is formally equal to 6 (of 7 possible)
according to the classification in (Van der Voo, 1997)
and 7 (of possible 8) according to A.N. Khramov in
(Dopolneniya…, 2000). Thus, magnetic polarity determinations for the largest part of the Berriasian section
(Members 1–23) deserve credence despite the generally low paleomagnetic quality of examined rocks.
CONCLUSIONS
(1) The contact between the carbonate Bedenekyr
and terrigenous Bechku formations is described for the
first time for central Crimea with the age of the upper
part of the former unit attributed to the Berriasian
occitanica Zone being specified.
(2) On the basis of the combined paleontological
and magnetostratigraphic data, the Malbosiceras chaperi Beds are attributed to the occitanica Zone.
(3) Six successive foraminiferal assemblages are
defined through the Berriasian section (from the base
upward): (1) Everticyclammina virguliana–Retrocyclammina recta–Bramkampella arabica; (2) Lenticulina muensteri; (3) Quadratina tunassica; (4) Triplasia
emslandensis acuta; (5) Lenticulina andromede;
(6) Conorboides hofkeri.
(4) The ostracod assemblages provide basis for
defining the following biostratigraphic units in the
Beriassian section (from the base upward): Costacythere khiamii–Hechticythere belbekensis Beds
correlated partly with the ammonite occitanica Zone
and the Costacythere drushchitzi–Reticythere
marfenini Beds correlated partly with the ammonite
boissieri Zone.
(5) The dinocysts assemblage allows the Phoberocysta neocomica Beds correlated partly with the ammonite occitanica and boissieri zones to be defined.
(6) A series of isolated outcrops (Novoklenovo,
Balki, and Mezhgor’e settlements) are united into a
composite section, which represents now the most
complete Berriasian succession for central Crimea.
Owing to magnetostratigraphic data obtained for this
composite section, the presence of all the Berriasian
magnetic chrons in central and eastern Crimea is substantiated for the first time.
(7) On the basis of paleomagnetic data, the Berriasian section of central Crimea is correlated with its
stratotype in the Mediterranean region with the position of standard Berriasian zones being specified.
(8) The established anisotropy in magnetic susceptibility is used for reconstructing directions of deformations of terrigenous (clayey) rocks in response to
tectonic movements.
ACKNOWLEDGMENTS
We are grateful to S.V. Lobacheva (posthumously)
for identifying brachiopods, V.A. Perminov (Center of
Learning Youth (TSENTUM) Intellekt, Feodosiya,
Russia) for his help in organizing field works,
A.Yu. Kurazhkovskii (Borok Geophysical Observatory) for help in conducting magnetic–mineralogical
investigations, and N.V. Platonova (assistant to the
director of the Resource Center X-ray Diffraction
Research Methods of St. Petersburg State University)
for accomplishing the X-ray phase analysis of samples.
STRATIGRAPHY AND GEOLOGICAL CORRELATION
Vol. 23
No. 2
2015
Statistical parameters of the distribution of the stable component of magnetization (SCM) directions
No. 2
Iav
(°)
k
α95
(°)
R
33
177.9
–37.1
5.0
12.5
Hcr = 19.6–26.8 × 103 A/m
R
5
177.5
–45.7
30.7
14.0
Hcr = 19.6–27.7 × 103 A/m
R
7
182.6
–39.5
11.3
18.7
Hcr = 19.6–30.9 × 103 A/m
R
9
182.9
–38.2
10.6
16.6
R
14
172.0
–47.9
16.1
10.2
Stable component of magnetization (SCM) calculated
using H demagnetization data on magnetically softest
R
samples (Hcr =19.6–26.8 × 103 A/m) and T° demagnetization
19
173.5
–47.4
19.0
7.9
Magnetically
soft
T° demagnetization
2015
Members 5–28 (terrigenous rocks)
Polarity
Dav
(°)
H demagnetization
All samples Hcr = 45.4 – 325.4 × 103 A/m
Members 1–3 (limestones)
Vol. 23
n
N
When an angle exceeds the
Results of the reversal test error value (±), the vectors
(McFadden
demonstrate significant difand McElhinny, 1990) ference; in the opposite situation, they are statistically
coincident (Debiche and
Watson, 1995)
A (°) Ac (°)
Cl
Angle with Angle with
RF (°) SCM + 180 (°)
5.6
91
SCM + 180°
1.7
46.8
353.5
47.9
Magnetization reversal by the recent field (RF)
2
4.3
58.7
Western Crimea (J3) (Pecherskii and Safonov, 1993)
11
352
47
14.7
4.0
8.8
B
11.8 ± 4.4
4.5 ± 6.7
(A) angle between vectors 12.5 ± 6.7
(Ac) critical angle
(Cl) classification
10
102.8
4.5
12.6 ± 6.1
13.7 ± 3.2
1.1 ± 6.8
NEW DATA ON BERRIASIAN BIOSTRATIGRAPHY, MAGNETOSTRATIGRAPHY
STRATIGRAPHY AND GEOLOGICAL CORRELATION
(n) number of samples in the selection
(Dav, Iav) average paleomagnetic declination
and inclination, respectively
(k) precision parameter
(α95) radius of the vector confidence circle
The amplitude of the secular geomagmetic variation (10%, according to V.G. Bakhmutov, 2006; (1200) summer variation is characterized by the amplitude of 8%).
187
Tirnovella occitanica
188
m
450
400
350
300
250
200
150
100
50
0
Fauriella boissieri
Member
Polarity
ARKADIEV et al.
?
?
Stage
Zone
Polarity
Berriasian
Tirnovella occitanica
T. subalpina
Stage
Fauriella boissieri
B. privasensis D. dalmas M. paramimounum
Eastern Crimea
Sultanovka
Berriasian–
Valanginian
(Bagaeva et al., 2011)
(Arkadiev et al.,
2010)
Zavodskaya Balka
50
0
Berriasian
F. boissieri
P. grandis
Vol. 23
B. picteti
Zone
Subzone
B. callisto
Polarity
Tithonian
Stage
Berriasian
Berriasella jacobi
Subthurmannia occitanica
Zone
Subthurmannia boissieri
Polarity
M14
M15
M16
M17
M18
M19
No. 2
Ma
140
141
142
143
144
145
2015
Magnetochronological
scale
28
27
26
25
24
23
22
21
20
19
18
–
10
9
8–4
3
2
1
STRATIGRAPHY AND GEOLOGICAL CORRELATION
Dinocysts beds
(Ogg and Hinnov, 2012)
Ostracod beds
Phoberocysta neocomica
Berriasian stratotype
(France)
Costacythere drushchitzi–Reticythere marfenini
(Galbrun, 1985)
Foraminiferal beds/
assemblages
?
Costacythere khiamii–Hechticythere belbekensis
Conorboides
hofkeri
Lenticulina
andromede
?
Lenticulina Quadratina
muensteri tunassica Triplasia emslandensis acuta
Central Crimea, Russia
Beds with
Zeillerina
baksanensis
Beds with
Symphythiris
arguinensis
Riasanites
crassicostatum
Neocosmoceras
euthymi
Dalmasiceras
tauricum
Beds with
Malbosiceras
chaperi
grandis
Textularia crimica–Belorussiella taurica
Evirticyclammina virguliana, Rectocyclammina recta,
Bramkampella arabica
Composite section (this work)
Stage
Zone
Subzone,
beds with fauna
(Arkadiev et al.,
2012)
Berriasian
B. jacobi
Magnetic chrons
NEW DATA ON BERRIASIAN BIOSTRATIGRAPHY, MAGNETOSTRATIGRAPHY
189
Fig. 11. Correlation of magnetostratigraphic data available for the Berriasian section of central and eastern Crimea with the Berriasian stratotype in southeastern France and standard magnetostratigraphic scale.
In the paleomagnetic column of the composite section of central Crimea, gaps 5 m wide and less in determinations of polarity are
not shown. For legend, see Fig. 4.
This work was supported by the Russian Foundation for Basic Research (project nos. 11-05-00405-a,
13-05-00745-a, 14-05-31152_a_mol) and Ministry of
Education and Science of the Russian Federation in
the base part (project 1582) and the scientific field
(project 1757).
Reviewers M.A. Rogov, A.Yu. Kazanskii,
and V.A. Zakharov
REFERENCES
Andreev, Yu.N. and Ertli, Kh.Yu., Some Cretaceous ostracods of Central Asia and related species of Europe, Vopr.
Mikropaleontol., 1970, no. 13, pp. 95–121.
Andreev, Yu.N., Cretaceous ostracods of Central Asia
(composition, distribution, evolution, geological significance), Doctoral (Geol.-Mineral.) Dissertation, Moscow:
MGU, 1986.
Arkadiev, V.V., Bagaeva, M.I., Guzhikov, A.Yu., et al., Bioand magnetostratigraphic characterization of the Upper
Berriasian section “Zavodskaya balka” (Eastern Crimea,
Feodosiya), Vestn. Sankt-Peterburg. Univ., Ser. 7. Geol.
Geogr., 2010, Iss. 2, pp. 3–16.
Arkadiev, V.V., Stage subdivision of Berriasian deposits of
Crimean Mountains, Vestn. Sankt-Peterburg. Univ., Ser. 7.
Geol. Geogr., 2007, Iss. 2, pp. 27–43.
Arkad’ev, V.V., Bogdanova, T.N., Lobacheva, S.V., et al., The
Berriasian of Crimean Mountains: problems of zonal subdivision and correlation, in Mat. Tret’ego Vseross. soveshch.
“Melovaya sistema Rossii i blizhnego zarubezh’ya: problemy
stratigrafii i paleogeografii”, Saratov, 26–30 sentyabrya 2006
(Proc. 3rd All-Russ. Conf. “The Cretaceous System of Russia and Adjacent Countries: Problems of Stratigraphy and
Paleogeography”, Saratov, September 26–30, 2006), Musatov, V.A., Ed., Saratov: Izd-vo SO EAGO, 2006, pp. 18–20.
Arkadiev, V.V., Bogdanova, T.N., Lobacheva, S.V., et al.,
Berriasian Stage of the Crimean Mountains: Zonal subdivisions and correlation, Stratigr. Geol. Correl., 2008, vol. 16,
no. 4, pp. 400–422.
Arkadiev, V.V., Bogdanova, T.N., Guzhikov, A.Yu., et al.,
Berrias Gornogo Kryma (The Berriasian of the Crimean
Mountains), St. Petersburg: Izd-vo “LEMA”, 2012 [in Russian].
Atlas des Ostracodes de France, Oertli, H.J., Ed., Bull. Centre Rech. Explor.-Prod. Elf.-Aquit. Mem., 1985, no. 9.
Bagaeva, M.I., Arkadiev, V.V., Baraboshkin, E.Yu., et al.,
New data on bio- and magnetostratigraphy of Berriasian—
Valanginian boundary deposit of Eastern Crimea, in Paleontologiya, stratigrafiya i paleogeografiya mezozoya i kainozoya
boreal’nykh raionov. Materialy nauchn. sessii (18–22 aprelya
2011 g.). T. I. Mezozoi (Proc. Sci. Sess. “Paleontology,
Stratigraphy, and Paleogeography of the Mesozoic and CenSTRATIGRAPHY AND GEOLOGICAL CORRELATION
ozoic of the Boreal Regions” (April 18–22, 2011). Vol. I:
Mesozoic), Novosibirsk: INGG SO RAN, 2011, pp. 23–26.
Baraboshkin, E.Yu., Prakticheskaya sedimentologiya. Terrigennye rezervuary. Posobie po rabote s kernom (Practical
Sedimentology. Clastic Reservoirs. The manual on the core
analysis), Tver: GERS, 2011 [in Russian].
Bogdanova, T.N., Lobacheva, S.V., Prozorovskii, V.A., and
Favorskaya, T.A., On the Berriasian Stage subdivisions in
the Crimean Mountains, Vestn. Leningrad. Univ., Ser. Geol.,
Geogr., 1981, no. 6, pp. 5–14.
Bogdanova, T.N. and Kvantaliani, I.V., New Berriasian
ammonites of the Crimea, Byull. Mosk. O–va Ispyt. Prir.,
Otd. Geol., 1983, vol. 58, no. 3, pp. 70–83.
Buskirk, R. E. and O’Brien Jr., W. P., Magnetic remanence
and response to magnetic fields in crustacea, in Magnetite
Biomineralization and Magnetoreception in Organisms, Kirschvink, J.L., Jones, D.S., MacFadden B.M., Eds., New
York, London: Plenum Press., 1985, pp. 365–383.
Debiche, M.G. and Watson, G.S., Confidence limits and
bias correction for estimating angles between directions
with applications to paleomagnetism, J. Geophys. Res.,
1995, vol. 100, no. B12, pp. 24405–24430.
Donze, P., Ostracodes berriasiens des subalpins septentrionaux (Bauges et Chartreuse), Trav. Lab. Géol. Fac. Sc.
Lyon. NS, 1964, no. 11, pp. 103–158.
Donze, P., Especes nouvelles d’Ostracodes des couches de
base du Valanginien de Berrias, Trav. Lab. Géol. Fac. Sc.
Lyon. NS, 1965, no. 12, pp. 87–107.
Dopolneniya k stratigraficheskomu kodeksu Rossii (Supplements to the Stratigraphic Code of Russia), St. Petersburg:
VSEGEI, 2000 [in Russian].
Drushchits, V.V. and Yanin, B.T., Lower Cretaceous deposits of the Central Crimea, Vestn. MGU, Ser. Biol., Pochvoved., Geol., Geogr., 1959, no. 1, pp. 115–120.
Drushchits, V.V. and Gorbachik, T.N., Zonal subdivision
of the Lower Cretaceous of the South of the USSR based
on ammonites and foraminifers, Izv. Akad. Nauk SSSR,
Ser. Geol., 1979, no. 12, pp. 95–105.
Espitalie, J. and Sigal, J., Contribution a l’etude des Foraminiferas du Jurassique superieur et du Neocomien du
Bassin de Majunga (Madagascar), Ann. Geol. de Madagascar, 1963, no. 32.
Feodorova, A.A., Reference sections of Jurassic–Cretaceous
boundary sediments in the Crimea as a basis for detailed stratification and correlation of productive sequences in the Caspian shelf, in Stratigrafiya neftegazonosnykh basseinov Rossii
(Stratigraphy of the Oil-and-Gas Basins in Russia), Vinogradov, N.V., Ed., St. Petersburg: Nedra, 2004, pp. 61–80.
Fisher, M.J. and Riley, L.A., The stratigraphic distribution
of dinoflagellate cysts at the boreal Jurassic–Cretaceous
boundary, Proc. Fourth Int. Palynol. Conf., Lucknow, 1980,
no. 2, pp. 313–329.
Vol. 23
No. 2
2015
190
ARKADIEV et al.
Flügel, E., Microfacies of Carbonate Rocks: Analysis, Interpretation, and Application. Second Edition, Berlin, Heidelberg: Springer, 2010.
Galbrun, B., Magnetostratigraphy of the Berriasian stratotype section (Berrias, France), Earth Planet. Sci. Lett.,
1985, vol. 74, pp. 130–136.
Gorbachik, T.N. and Mokhamad, G.K., Lituolida Foraminifera from the Tithonian and Berriasian of the Crimea;
structure, importance for stratigraphy and paleobiogeography, in Problemy stratigrafii i paleontologii mezozoya (Problems of Stratigraphy and Paleontology of the Mesozoic),
Kozlov, G.E. and Prozorovskii, V.A., Eds., St. Petersburg:
VNIGRI, 1999, pp. 165–186.
Gradstein, F., Ogg, J.G., Schmitz, M.D., and Ogg, G.M.,
The Geologic Time Scale 2012, Amsterdam: Elsevier, 2012.
Grekoff, N. and Magne, J., Les ostracodes du stratotype du
Berriasien, Rev. de Micropaleontol., 1966, vol. 9, no. 3,
pp. 177–185.
Gründel, J., Neue ostracoden aus der deutschen Unterkreide II, Monatsb. Deutschen Akad. Wiss. Berlin, 1964, vol. 6,
no. 11, pp. 849–858.
Guzhikov, A.Yu., Arkad’ev, V.V., Baraboshkin, E.Yu., et al.,
New sedimentological, bio-, and magnetostratigraphic data
on the Jurassic-Cretaceous boundary interval of Eastern
Crimea (Feodosiya), Stratigr. Geol. Correl., 2012, vol. 20,
no. 3, pp. 261–294.
Kolpenskaya, N.N., Ostracods. The Berriasian of Northern
Caucasus (Urukh section), in Biokhronologiya i korrelyatsiya
fanerozoya neftegazonosnykh basseinov Rossii. Vyp. 2 (Biochronology and Correlation of the Phanerozoic of the Oiland-Gas Basins of Russia), Kirichkov, A.I., Ed., St. Petersburg: VNIGRI, 2000, pp. 42–52, 115–129.
Kubiatowicz, W., Upper Jurassic and Neocomian ostracodes from Central Poland, Acta Geol. Pol., 1983, vol. 33,
nos. 1–4, pp. 1–72.
Kuznetsova, K.I. and Gorbachik, T.N., Stratigrafiya i foraminifery verkhnei yury i nizhnego mela Kryma (Upper Jurassic and Lower Cretaceous Stratigraphy and Foraminifers of
the Crimea), Moscow: Nauka, 1985 [in Russian].
Kvantaliani, I.V. and Lysenko, N.I., On the problem of
zonal subdivision of the Berriasian of the Crimea, Soobshch.
Akad. Nauk Gruz. SSR, 1979, vol. 94, no. 3, pp. 629–632.
Lanza, R. and Meloni, A., The Earth’s Magnetism: An Introduction for Geologist, Berlin-Heidelberg-New York: Springer,
2006.
Le Hégarat, G., Le Berriasien du Sud-East de la France,
Doc. Lab. Geol. Fac. Sci., 1973, vol. 43/1.
McFadden, P.L. and McElhinny, M.W., Classification of
the reversal test in palaeomagnetism, Geophys. J. Int., 1990,
vol. 103, pp. 725–729.
Molostovskii, E.A., Eremin, V.N., Guzhikov, A.Yu., et al.,
Paleomagnetic stratigraphy of Mesozoic and Cenozoic
deposits in southern and southeastern parts of the European
USSR, in Otchet o NIR po teme “Provesti detalizatsiyu i
utochnenie magnitostratigraficheskoi shkaly mezozoya i kainozoya SSSR” (Report on R&D Work: Detalization and
Clarification of Magnetostratigraphic Scale of Mesozoic
and Cenozoic of the USSR), Saratov: NII Geol. Saratov
State Univ., 1989 [in Russian].
Monteil, E., Kystes de dinoflagelles index (Tithonique–
Valanginien) du Sud-Est de la France. Proposition d’une
nouvelle zonation palynologique, Revue de Paleobiol.,
1992, vol. 11, no. 1, pp. 299–306.
Morkhoven, F.P.C.M., Generic descriptions, in Post-Paleozoic Ostracoda. Vol. II, Amsterdam–London–New York:
Elsevier, 1963.
Neale, J.W., Ostracoda from the Speeton Clay (Lower Cretaceous) of Yorkshire, Micropaleontol., 1962, vol. 8, no. 4,
pp. 425–484.
Neale, J.W., Ostracodes from the type Berriasian (Cretaceous) of Berrias (Ardeche, France) and their significance,
Univ. Kansas. Depart. Geol. Spec. Publ., 1967, no. 2, pp. 539–
569.
Neale, J.W., The Cretaceous, in A Stratigraphical Index of
British Ostracoda, Bate, R.N. and Robinson, E., Eds., Geol.
J. Spec. Iss., 1978, no. 8, pp. 325–384.
Nikishin, A.M., Bolotov, S.N., Baraboshkin, E.Yu., et al.,
Geological history of the Scythian-Black Sea Region, in
Ocherki geologii Kryma. Vyp. 1 (Essays on Geology of the
Crimean Peninsula), Moscow: MGU, 1997, pp. 207–227.
Neale, J.W., Ostracodes from the Lower Valanginian of the
Central Crimea, Paleontol. Zh., 1966, no. 1, pp. 87–100.
Pecherskii, D.M. and Safonov, V.A., Palinspastic reconstructions of the position of Crimean Mountains in Middle
Jurassic–Early Cretaceous based on the paleomagnetic
data, Geotektonika, 1993, no. 1, pp. 96–105.
Pokorny, V., The ostracoda of the Klentnice Formation
(Tithonian?), Czechoslovakia, Rozp. Ustred. Ust. Geol., 1973,
vol. 40, pp. 1–107.
Prakticheskoe rukovodstvo po mikrofaune. T. 7. Ostrakody
mezozoya (Practical Manual on Microfauna. Vol. 7. Mesozoic Ostracoda), Nikolaev, I.A. and Neustruev, I.Yu., Eds.,
St. Petersburg: VSEGEI, 1999 [in Russian].
Raevskaya, E.G. and Shurekova, O.V., Modern technology
and equipment for palynological analysis of carbonate-terrigenous deposits, in Problemy sovremennoi palinologii.
Materialy XIII Rossiiskoi palinologicheskoi konferentsii
(Proc. XIII Russ. Palynol. Conf. “Problems of Modern
Palynology”), Syktyvkar: Inst. Geol. Komi NTs UrO RAN,
2011, pp. 103–107.
Slipper, I.J., Marine Lower Cretaceous, in Ostracods in British Stratigraphy, Whittaker, J.E. and Hart, M.B., Eds., Publ.
Micropaleont. Soc. Geol. Soc. London, 2009, pp. 309–344.
Stolz, J.F., Chang, S.B.R., and Kirschvink, J.L., Bacterial
magnetite as trace fossil and paleooxygen indicator, Origins of
Life and Evolution of the Biosphere, 1986, vol. 16, nos. 3–4.
Tavera, J.M., Los ammonites del Tithonico Superior-Berriasense de la zona Subbetica (Cordilleras Beticas). Tesis
Doctoral, Granada: Univ. Granada, 1985.
Tesakova, E.M. and Rachenskaya, L.P., New Ostracodes
(Crustacea, Ostracoda) of the Genus Costacythere Gründel
from Berriasian sediments of the Crimea, Paleontol. Zh.,
1996, vol. 30, no. 4, pp. 416–428.
Tesakova, E.M. and Rachenskaya, L.P., New Ostracodes
(Crustacea, Ostracoda) of the genera Bairdia M’Coy, Neocythere Mertens, Macrodentina Martin, Hechticythere Gründel, and Cypridea Bosquet from the Berriasian of Central
Crimea, Paleontol. J., 1996, vol. 30, no. 5, pp. 542—551.
STRATIGRAPHY AND GEOLOGICAL CORRELATION
Vol. 23
No. 2
2015
NEW DATA ON BERRIASIAN BIOSTRATIGRAPHY, MAGNETOSTRATIGRAPHY
Triebel, E., Ostracoden untersuchungen, Protocythere und
Exophtalmocythere, zwei neue Ostracoden gattungen aus der
Deutschen Kreide, Senckenbergiana Lethaea, 1938, vol. 20,
no. 1/2, pp. 179–199.
Tucker, M.E. and Wright, V.P., Carbonate Sedimentology,
Oxford: Blackwell Science, 1990.
Van der Voo, R., Palaeomagnetism of the Atlantic, Tethys, and
Iapetus oceans, Cambridge: Cambridge Univ. Press, 1993.
Williams, G.L., Dinoflagellate and spore stratigraphy of the
Mesozoic–Cenozoic, Offshore Eastern Canada, Geol. Surv.
Canada, 1975, Pap. 74–30, pp. 107–161.
Wright, V.P. and Burgess, P.M., The carbonate factory continuum, facies mosaics and microfacies: an appraisal of
some of the key concepts underpinning carbonate sedimentology, Facies, 2005, vol. 51, pp. 17–23.
Yampolskaya, O.B., Paleomagnetism and petromagnetism
of Lower Cretaceous deposits of Crimean Mountains:
STRATIGRAPHY AND GEOLOGICAL CORRELATION
191
stratigraphic and paleogeographic aspects, Cand. Sci.
(Geol.-Mineral.) Dissertation , Saratov: SGU, 2005.
Yampolskaya, O.B., Baraboshkin, E.Yu., Guzhikov, A.Yu.,
et al., Lower Cretaceous paleomagnetic section in the
southwest Crimea, Moscow Univ. Geol. Bull., 2006, vol. 61,
no. 1, pp. 1–16.
Yanin, B.T. and Smirnova, T.N., Stratigraphic distribution
of bivalves and brachiopods in Berriasian and Valanginian
deposits of the Crimea, Byull. Mosk. O-va Ispyt. Prir., Otd.
Geol., 1981, vol. 56, no. 1, pp. 82–94.
Yanin, B.T. and Baraboshkin, E.Yu., The Berriasian section
in the Bel’bek River Basin, Southwestern Crimea, Stratigr.
Geol. Correl., 2000, vol. 8, no. 2, pp. 165–176.
Translated by I. Basov
Vol. 23
No. 2
2015