NAT. CROAT. VOL. 20 No 2 237¿354 ZAGREB December 31, 2011
original scientific paper / izvorni znanstveni rad
CATALOGUE AND ATLAS
OF CAVE-DWELLING TERRESTRIAL ISOPODS
(CRUSTACEA: ONISCIDEA) FROM CROATIA
JANA BEDEK1, STEFANO TAITI2 & SANJA GOTTSTEIN3
1
2
Croatian Biospeleological Society, Demetrova 1, HR-10 000 Zagreb
(jana.bedek@hbsd.hr)
Istituto per lo Studio degli Ecosistemi, CNR, Via Madonna del Piano 10,
I-50019 Sesto Fiorentino (stefano.taiti@ise.cnr.it)
3
Department of Zoology, Faculty of Science, Rooseveltov trg 6, HR-10 000
Zagreb (sgottst@biol.pmf.hr)
Bedek, J., Taiti, S. & Gottstein, S.: Catalogue and atlas of cave-dwelling terrestrial isopods
(Crustacea: Oniscidea) from Croatia. Nat. Croat., Vol. 20, No. 2., 237–354, 2011, Zagreb.
This paper presents a checklist with historical overview and new records on the cave-dwelling
terrestrial isopods (troglophiles and troglobionts) from Croatia, based on detailed analyses of available literature and unpublished data from collections. For each species the following data are
quoted: the name of the taxon; synonyms; general distribution; type locality; ecological status; list
of locality records; species names used in the cited literature; the distribution map using 10 × 10 km
UTM grid squares. Annexes with the locality synonyms used in the examined literature and collection data are enclosed. Altogether this paper presents data from 557 caves, 6 artificial underground
structures and 15 other epigean and subterranean localities. A checklist of cave terrestrial isopod
taxa in Croatia is composed of 35 species and five subspecies distributed in five families, including
22 Croatian endemic and 20 troglobiotic taxa. The family Trichoniscidae and the subfamily Trichoniscinae are the most representative with 26 and 20 species, respectively. The most represented genus is Alpioniscus with 10 species. The South Croatian Littoral macroregion has the highest number
of species and endemics among Croatian macroregions. The most widely distributed species in
Croatia are the troglophiles Mesoniscus graniger and Androniscus roseus found in three macroregions.
Only four troglobiotic species (Alpioniscus balthasari, A. strasseri, Titanethes albus and T. dahli) are distributed in two macroregions, while the rest are distributed only in single macroregions. All the 40
Croatian cave taxa are distributed in the Dinaric karst area, and only three troglophilic species are
found also outside. No Croatian endemic taxa are found outside the Dinaric karst area.
Key words: checklist, endemic taxa, Dinaric karst, biodiversity, distribution
Bedek, J., Taiti, S. & Gottstein, S.: Katalog i atlas {piljskih kopnenih jednakono`nih rakova
(Crustacea: Oniscidea) Hrvatske. Nat. Croat., Vol. 20, No. 2., 237–354, 2011, Zagreb.
U ovom radu je prikazan popis vrsta s povijesnim pregledom i novim nalazima {piljskih jednakono`nih kopnenih rakova, koji se temelji na detaljnoj analizi dostupne literature o {piljskim jednakono`nim rakovima te neobjavljenim podacima iz zbirki. Popisani su svi poznati {piljski jednakono`ni
rakovi (troglofili i troglobionti). Za svaku vrstu se navode sljede}i podaci: ime svojte; sinonimi; op}a
Croatian Natural History Museum, Demetrova 1, Zagreb, Croatia
238
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
rasprostranjenost; tipski lokalitet; ekolo{ki status; popis nalaza; ime vrste upotrijebljeno u citiranoj
literaturi; karta rasprostranjenosti na UTM mre`i 10 × 10 km. Prilo`eni su dodaci s popisom sinonima
objekata upotrijebljenih u citiranoj literaturi te podaci iz zbirki. Sveukupno rad obuhva}a podatke
iz 557 speleolo{ka objekta, {est podzemnih gra|evina i 15 ostalih nadzemnih i podzemnih
lokaliteta. Popis {piljskih svojti utvr|enih do danas za podru~je Hrvatske obuhva}a 35 vrsta i pet
podvrsta unutar pet porodica koje uklju~uju 22 endema Hrvatske i 20 troglobionata. S obzirom na
broj vrsta najzastupljenija je porodica Trichoniscidae (26 vrsta) i potporodica Trichoniscinae (20 vrsta).
Rod Alpioniscus sa 10 vrsta je naju~estaliji. Makroregija ju`no hrvatsko primorje ima mnogo vi{e
vrsta i endema od ostalih regija. Naj{ire rasprostranjene vrste su troglofilne vrste Mesoniscus graniger,
Androniscus roseus i troglobiontna vrsta Androniscus stygius, rasprostranjene u tri makroregije. Samo
~etiri troglobiontne vrste (Alpioniscus balthasari, A. strasseri, Titanethes albus i T. dahli) su rasprostranjene u dvije makroregije, ostale su utvr|ene samo u jednoj. Od 40 {piljskih svojti sve su rasprostranjene u Dinarskom kr{u a samo tri izvan Dinarskog kr{a. Izvan Dinarskog kr{a nema Hrvatskih endemskih svojti.
Klju~ne rije~i: popis vrsta, endemi, dinarski kr{, bioraznolikost, rasprostranjenost
CONTENTS
INTRODUCTION . . . . . .
HISTORICAL REVIEW . . . .
MATERIALS AND METHODS .
RESULTS . . . . . . . . . .
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Mesoniscus graniger (Frivaldsky, 1865) . . . . . .
Androniscus dentiger Verhoeff, 1908. . . . . . . .
Androniscus roseus buccariensis Verhoeff, 1930 . . .
Androniscus roseus histrianorum (Arcangeli, 1932) . .
Androniscus roseus ocellatus (Kesselyak, 1930) . . .
Androniscus roseus roseus (C. Koch, 1838) . . . . .
Androniscus stygius microcavernicolus Kesselyak, 1931
Androniscus stygius stygius (Nemec, 1897) . . . . .
Androniscus wolfi Strouhal, 1939 . . . . . . . . .
Aegonethes antilocapra Frankenberger, 1938 . . . .
Aegonethes cervinus (Verhoeff, 1931) . . . . . . .
Alpioniscus balthasari (Frankenberger, 1937) . . . .
Alpioniscus bosniensis (Frankenberger, 1939) . . . .
Alpioniscus christiani Poto~nik, 1983 . . . . . . .
Alpioniscus haasi (Verhoeff, 1931) . . . . . . . .
Alpioniscus heroldi (Verhoeff, 1931) . . . . . . . .
Alpioniscus herzegowinensis (Verhoeff, 1931) . . . .
Alpioniscus kratochvili (Frankenberger, 1938) . . . .
Alpioniscus magnus (Frankenberger, 1938) . . . . .
Alpioniscus strasseri (Verhoeff, 1927) . . . . . . .
Alpioniscus trogirensis Buturovi}, 1955 . . . . . .
Alpioniscus verhoeffi (Strouhal, 1938) . . . . . . .
Titanethes albus (C. Koch, 1841) . . . . . . . . .
Titanethes dahli Verhoeff, 1926 . . . . . . . . . .
Cyphonethes herzegowinensis (Verhoeff, 1900) . . . .
Trichoniscus matulici matulici Verhoeff, 1901 . . . .
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Trichoniscus matulici metkovicensis Buturovi}, 1955 . . . .
Strouhaloniscellus biokovoensis Bedek et Taiti, 2009 . . . .
Troglocyphoniscus absoloni Strouhal, 1939 . . . . . . . .
Cyphopleon kratochvili (Frankenberger, 1939) . . . . . . .
Cyphoniscellus herzegowinensis (Verhoeff, 1900) . . . . . .
Buddelundiella cataractae Verhoeff, 1930 . . . . . . . . .
Thaumatoniscellus speluncae Karaman, Bedek et Horvatovi},
Chaetophiloscia cellaria (Dollfus, 1884) . . . . . . . . . .
Oroniscus dalmaticus Strouhal, 1937 . . . . . . . . . .
Oroniscus meledensis Strouhal, 1937. . . . . . . . . . .
Oroniscus stentai (Arcangeli, 1926) . . . . . . . . . . .
Typhlarmadillidium kratochvili (Frankenberger, 1938) . . . .
Typhlarmadillidium trebinjanum (Verhoeff, 1900). . . . . .
Echinarmadillidium fruxgalii (Verhoeff, 1900) . . . . . . .
Cyphodillidium absoloni (Strouhal, 1934) . . . . . . . . .
DISCUSSION . . . . . . . . . . . . . . . . .
ACKNOWLEDGEMENTS . . . . . . . . . . . .
REFERENCES . . . . . . . . . . . . . . . . .
SUMMARY . . . . . . . . . . . . . . . . . .
APPENDIX 1. List of synonyms for locality records .
APPENDIX 2. Collection records . . . . . . . . .
APPENDIX 3. Alphabetical order of localities . . .
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INTRODUCTION
In Croatia 130 species and 30 subspecies of terrestrial isopods have been recorded to date, 59 species and 17 subspecies being endemic to Croatia (ARCANGELI, 1939a,
POTO^NIK, 1989, SCHMALFUSS, 2003, FARKAS & KR^MAR, 2004, FARKAS & VILISICS,
2008, BEDEK & TAITI, 2009, KARAMAN et al., 2009). This paper presents an overview
of the present knowledge on cave-dwelling terrestrial isopods, and is based on data
available in the literature and unpublished data from collections.
HISTORICAL REVIEW
The first known data about terrestrial isopods from Croatian caves were given
by the school teacher Eugen Adolf Jurinac from Vara`din. He performed the first
systematic biospeleological research in the period from 15th to 31st August 1883 in
the Kordun region. Among other cave animals he collected Titanethes albus (C.
Koch, 1841) in three caves in the Ogulin region (JURINAC, 1886, 1888). In the beginning of the 20th century the director of the Croatian Natural History Museum August Langhoffer led major biospeleological surveys in Kordun, Gorski Kotar, Lika
and Zagreb (LANGHOFFER, 1912, 1915a, 1915b) and the entomologist Victor Stiller
from Zagreb also explored Kordun and Gorski Kotar (STILLER, 1911a, 1911b, 1913,
1914). The Dalmatian cave fauna was first investigated by Umberto Girometta, a
school teacher and speleologist from Split, and his students from 1912 to 1914
(GIROMETTA, 1913, 1914). A mammalogist from Vienna, Otto Wettstein, explored
240
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
caves on the island of Bra~ in 1912 in an expedition organized by the biologist Prof.
Dr. Franz Werner (WETTSTEIN, 1914). They all collected different species of the genera
Alpioniscus, Androniscus and Titanethes, but mostly misidentified them as Titanethes
albus or Trichoniscus roseus. All known data from that period were reviewed by the
Hungarian isopodologist Ernö Csiki in his famous paper »Magyarország szárazföldi
Isopodái. (Isopoda terrestria Hungariae)« (CSIKI, 1926).
From the beginning of the 20th century many foreign biologists (mostly entomologists) collected numerous specimens from caves, including terrestrial isopods. The
collected material resulted in a lot of new species being described, mostly in the
thirties of the last century. The first scientific analyses date to 1926, when the Italian
isopodologist Alceste Arcangeli described the troglophilic species Oroniscus stentai
(as Porcellio stentai) from Bra~; this had been collected in 1903 by Josef (Giuseppe)
Müller, coleopterologist from Trieste, born in Zadar (ARCANGELI, 1926). The German isopodologist Karl Verhoeff analyzed material from Istria collected by the
diplopodologist Karl Strasser (VERHOEFF, 1929a, 1929b). The Hungarian isopodologist Adorján Kesselyák described the troglophilic subspecies Androniscus stygius
microcavernicolus (as Androniscus roseus microcavernicolus), which he had collected on
Mt. U~ka in 1928 and 1929 (KESSELYÁK, 1931). Verhoeff described two new troglobiotic species from the island of Kor~ula collected by German zoologist Fritz Haas,
Alpioniscus haasi (as Illyrionethes haasi) and Aegonethes cervinus (as Illyrionethes cervinus) (VERHOEFF, 1931a, 1931b). A new troglophilic subspecies from Istria, Androniscus roseus histrianorum (as Trichoniscus roseus histrianorum), collected by Italian biologist Pietro Parenzan, was described by Arcangeli (ARCANGELI, 1932). Czech biologist Karel Absolon, one of the most famous explorers of Dinaric caves, collected
fauna on the island of Mljet in 1917. Out of that material the Austrian isopodologist
Hans Strouhal in 1934 described the troglophilic species Echinarmadillidium absoloni,
for which Verhoeff in 1939 described a new genus, Cyphodillidium (STROUHAL, 1934,
VERHOEFF, 1939). The Slovak isopodologist Zdenìk Frankenberger made a twoweek-long fieldtrip in Dalmatia in June 1937 and described the new troglobiotic
species Alpioniscus balthasari (as Illyrionethes balthasari) (FRANKENBERGER, 1937). In
the same year Strouhal described two new troglophilic species from the genus
Oroniscus collected by Absolon in 1914 from central Dalmatia (O. dalmaticus) and in
1917 from the island of Mljet (O. meledensis) (STROUHAL, 1937). The German biospeleologist Beno Wolf listed all cave isopods known up to 1935 in his famous
»Animalium Cavernarum Catalogus« (WOLF, 1938). The Czech arachnologist Josef Kratochvíl collected material from Dalmatia on his fieldtrips through the former Yugoslavia. His material was analyzed by Frankenberger who described five new species: Typhlarmadillidium kratochvili (as Cylisticus kratochvili) from the island of Kor~ula, Alpioniscus magnus (as Illyrionethes magnus) from the island of Bra~, Alpioniscus
kratochvili (as Illyrionethes kratochvili) from the island of Hvar, and Cyphopleon kratochvili (as Cyphoniscellus kratochvili) from Mt. Svilaja. He also erected the new genus
Aegonethes fot the species Illyrionethes cervinus from the islands of Kor~ula and Vis.
The new species A. antilocapra is also described by FRANKENBERGER (1938a, 1938b,
1939). Strouhal analyzed material from the Natural History Museum in Vienna,
collected by Czech coleopterologist Joseph Klimesch in 1913 and Werner in 1927,
and described the new troglobiotic species Alpioniscus verhoeffi (as Illyrionethes
verhoeffi) from the Neretva River collected by Austrian biologist Joseph Erber in
1861 (STROUHAL, 1938). He also analyzed material from the collection »Biospeleo-
Nat. Croat. Vol. 20(2), 2011
241
logica balcanica« gathered by Absolon and published the results in a series of papers named »Landasseln aus Balkanhöhlen, in der Sammlung Biospeleolgica balcanica«
and »Landasseln aus Balkanhöhlen, gesammelt von Prof. Dr. K. Absolon«. Many researchers besides Absolon contributed in collecting terrestrial isopods. Strouhal published
many new biogeographical data for a lot of species and described many new genera
and species from the Balkan region. From Croatia he described the new genus and
species Troglocyphoniscus absoloni from the island of Mljet (STROUHAL, 1939a, 1939b,
1939c, 1939d, 1939e). In 1940 Frankenberger and Strouhal summarized all the data,
which ended a very important period for the knowledge of cave terrestrial isopods
in Croatia.
In the fifties and the sixties of the last century the isopodologist Adem Buturovi}
from Sarajevo continued the study of terrestrial isopods on material collected by the
arachnologist Franjo Nikoli} from Dubrovnik from 1951 to 1953, by the Serbian biologist Stanko Karaman from 1924 to 1953, and material collected by himself in 1954.
He described the new troglophilic subspecies Trichoniscus matulici metkovicensis from
the surroundings of Metkovi} and the troglobiotic species Alpioniscus trogirensis from
the surroundings of Trogir, and added many new biogeographical data for other
known species (BUTUROVI], 1955b, 1955c, 1963). In 1966 the biologist Mladen Karaman, son of Stanko Karaman, published a complete review of terrestrial isopods
from Yugoslavia (KARAMAN, 1966). After that there was one long period of silence,
which was ended by the Slovenian isopodologist Franc Poto~nik with the description of the troglobiotic species Alpioniscus christiani, collected on the island of Krk by
the Austrian collembologist Erhard Christian (POTO^NIK, 1983). Poto~nik carried
out a survey of the fauna of terrestrial isopods of Yugoslavia (POTO^NIK, 1989).
Since 1970 the biospeleologist and technician from the Croatian Natural History
Museum Branko Jal`i} has collected material of terrestrial isopods from Croatia
(JAL@I] & BO@I^EVI], 1975, KEROVEC et al., 1999, JAL@I], 2001). With many colleagues
he established the Croatian Biospeleological Society in 1996. During biospeleological research since 1989 Roman Ozimec has collected terrestrial isopods (OZIMEC,
2003, OZIMEC, 2009), and assembled a great amount of references. Since the 1990s,
terrestrial isopods have been collected also by Ton}i Ra|a (VUJ^I]-KARLO & RA\A,
2004). Through the work of the Croatian Biospeleological Society many people have
become involved in the collection of cave animals, including terrestrial isopods:
Marijana Frani~evi}, Tomica Rubini}, Hrvoje Cvitanovi}, Gordan Poli}, Stipe Bu{eli}, Helena Biland`ija, Marko Luki}, Martina Pavlek, Predrag Rade, Ana Komeri~ki, Kazimir Miculini}, Robert Bakovi}, Vedran Jal`i}, Fanica Kljakovi} Ga{pi},
Alen Kirin, Petra Kova~ Konrad, and some young biology students such as Petra
Bregovi}, Nikolina Ragu`, Tamara ^ukovi}, An|ela ]uku{i}, Lana \ud, Dajana
Hmura, Marta Malenica, Petra Kutle{a and many others. Most of these samples
have been examined by J. Bedek since 2000 (BEDEK et al., 2000, GOTTSTEIN MATO^EC
et al., 2002, OZIMEC, 2004, 2005, OZIMEC & BEDEK, 2005, BEDEK 2005a, 2005b, 2007a,
2007b, JAL@I] et al., 2007, BAK[I] et al., 2008, BEDEK et al., 2008, OZIMEC et al., 2008, LUKI] et.al., 2009), which resulted in the descriptions of two new species in 2009: Strouhaloniscellus biokovoensis from Mt. Biokovo (BEDEK & TAITI, 2009), and Thaumatoniscellus speluncae from Mt. U~ka (KARAMAN et al., 2009) The present paper deals with
part of the results from field work carried out in the last 15 years by the Croatian
Biospeleological Society and the Croatian Natural History Museum; the remaining
results with descriptions of several new taxa will be published in future papers.
242
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
MATERIALS AND METHODS
This paper is based on a review of data of all available literature and on unpublished data. Cave-dwelling taxa of terrestrial isopods (troglophiles and troglobionts)
are listed in this catalogue with all known locality records, both from subterranean
and epigean environment.
Unpublished data come from projects of the Croatian Biospeleological Society and
the Croatian Natural History Museum on the cave fauna of Croatia. Only taxa identified to the level of species and subspecies are listed, while unidentified specimens
and new species are not included. A lot of material was collected in cooperation with
other organizations, especially the speleological societies: SO PDS Velebit, SO HPD
@eljezni~ar, DDSKF, SK Samobor, SK Ursus spelaeus, SU Estavela, SU Spelunka, SO
PD Dubovac, SD Karlovac and the Biological Students Society BIUS. Part of the
Crustacea Collection from the Croatian Natural History Museum was also examined,
collected in 1883 by Adolf Jurinac, in 1902 by August Langhoffer, and by Dragutin
Hirc (unknown date).
Isopods in caves were collected using tweezers in all habitats and preserved in
75% ethanol with glycerol. Micropreparations in Hoyer’s medium of the most important body parts were used for identifications. The collected material has been
deposited in the collections of the Croatian Biospeleological Society, the Croatian
Natural History Museum, the Museo di Storia Naturale dell’Università di Firenze,
Sezione di Zoologia »La Specola« in Italy, the Department of Biology and Ecology,
Faculty of Sciences, University of Novi Sad, Serbia and the Ton}i Ra|a private collection. The collection of the Croatian Biospeleological Society has been deposited
in the Croatian Natural History Museum.
For each species the following data are quoted:
1. Ordinal number. All taxa are marked with Arabic numerals. Taxa that are
only doubtfully present in Croatia are not marked with a number.
2. Taxa names. Taxonomic arrangement and nomenclature follows SCHMALFUSS
(2003).
3. Synonyms.
4. Distribution.
5. Type locality. The valid name is given with original names of type localities
in square brackets.
6. Ecological status. If there are no literature data, ecological status is given according to SKET (2008).
7. List of localities.
• The valid names of the localities in Croatian language according to topographic and speleological nomenclature are included. Frequent terms for
types of feature are: {pilja, spilja, spila, pe}, pe}ina = cave; jama, jametina,
rupa = pit; kaverna = cavern; ledenica = ice cave; snije`nica = snow cave;
izvor {pilja = spring cave; ponor = sinkhole; pukotina = crevice; sustav =
cave system; rudnik = mine.
• Localities are listed by level of precision of given locality (the most precise localities are at the beginning), then by alphabetical order of UTM grid squares,
then by alphabetical order of the name of the locality. Misidentifications,
doubtful data and misinterpreted data are placed at the end of the table.
Nat. Croat. Vol. 20(2), 2011
243
• The UTM grid with squares of a 10 × 10 km is used. Data are taken from
the UTM 1:500000 map of the world. If it is not established in which UTM
square the feature is located, when possible the UTM reference for a 100 ×
100 km field is given.
• The macroregion reference according to ROGLI] (1974) is given.
• Source of the record is given.
8. Species names used in cited literature, whether they are misidentifications or
misprints, are listed below the table with listed localities.
9. The Croatian distribution of each species is mapped using the 10 × 10 km
UTM grid squares on the map of Croatian territory with macroregions according to ROGLI] (1974) (Fig. 1). The other less precise localities are not
mapped, except if they are distant from other localities of that taxa. The localities for which the 10 × 10 km UTM square is precise are marked with a
circle or triangle and localities for which the 10 × 10 km UTM square is not
precise are marked with a transparent square.
10. Locality names used in the references with their present real name are listed
by alphabetical order in Appendix 1. In Appendix 2 collection records with
collector(s) and dates are given. All localities with their present real name are
listed in alphabetical order in Appendix 3.
Fig. 1. Croatian macroregions defined according to ROGLI] (1974) with the distribution of karst areas according to GOTTSTEIN MATO^EC et al. (2002). The Dinaric karst is
divided with a thick line. E (Eastern Croatia); C (Central Croatia); M (Mountainous
Croatia); NL (Northern Croatian Littoral); SL (Southern Croatian Littoral).
244
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
RESULTS
MESONISCIDAE
1. Mesoniscus graniger (Frivaldsky, 1865) (Tab. 1, Fig. 2)
Synonyms:
Titanethes graniger Frivaldsky, 1865; M. alpicola meridionalis Chappuis, 1944;
M. alpicola vulgaris Chappuis, 1944; M. histrianorum Arcangeli, 1939; M.
prenjanus Arcangeli, 1939; M. triangulifer (Verhoeff, 1930); Nematoniscus illyricus Verhoeff, 1933; N. prenjanus (Verhoeff, 1901); N. triangulifer Verhoeff,
1930; Schiödtia graniger (Frivaldsky, 1865); Titanethes graniger Heller, 1858;
Trichoniscus prenjanus Verhoeff, 1901
Distribution:
Slovenia, Serbia, Bosnia and Herzegovina, Slovakia, Romania (SCHMALFUSS,
2003); Croatia (ARCANGELI, 1939a)
Type locality:
Baradla barlang (cave system), Aggtelek, Hungary [Aggteleki barlang (FRIVALDSKY, 1865)]
Ecological status: In caves, but also under big rocks in deep soil (GIURGINCA, 2009); troglophilic
Tab. 1. List of recorded localities for Mesoniscus graniger
Locality
Jastrebina (cave), Frketi} Selo, Novigrad na Dobri, Karlovac
UTM
WL33
Region
C
Jama bijele sige (cave), Ponikve, Gornji Stenjevac, Medvednica,
Zagreb
WL67
C
Petnaestica (cave), Ponikve, Gornji Stenjevac, Medvednica,
Zagreb
WL67
C
Veternica (cave), Gornji Stenjevac, Medvednica, Zagreb
WL67
C
Francuski rudnik IV (mine), Sljeme, Medvednica, Zagreb
WL87
C
Medvednica (cave), Horvatove stube, Medvednica, Zagreb
WL87
C
Rudnik Zrinski (mine), Sljeme, Medvednica, Zagreb
WL87
C
Velika pe} (cave), Ka{ina, Medvednica, Zagreb
WL88
C
Rudnik Kra{evski zviri (mine), Lipovje, Prigorec, Ivan{~ica, Ivanec
Gradusa (cave), Sjeverovac, Petrinja, Banovina
WM81
XL12
C
C
Bijeli bunar (spring cave), Vo}in
XL95
E
Luda bu`a (endogean), Papuk, Velika, Papuk
YL04
E
Medvje|i brlog (cave), Papuk, Velika, Papuk
YL04
E
Uviraljka (endogean), Papuk, Velika, Papuk
Jama iznad Jezerca (cave), Jezerac, Jankovac, Velika, Papuk
Jama na Bilu (cave), Bilo, Velika, Papuk
YL04
YL14
YL14
E
E
E
Jan~ikin guz (cave), Jankovac, Velika, Papuk
YL14
E
Jelenova {pilja (cave), Jankovac, Velika, Papuk
Sedrena 1 kod slapa Skakavac (cave), Jankovac, Velika, Papuk
YL14
YL14
E
E
Source
CBSS collection 1
CBSS collection 2
BEDEK, 2007b
CBSS collection 3
BEDEK, 2007b
CBSS collection 4
BEDEK, 2007b
CBSS collection 5
BEDEK, 2007b
CBSS collection 6
BEDEK, 2007b
CBSS collection 7
BEDEK, 2007b
CBSS collection 8
BEDEK, 2007b
CBSS collection 9
CBSS collection 10
CBSS collection 11
BEDEK, 2007b
CBSS collection 12
BEDEK, 2007b
CBSS collection 13
BEDEK, 2007b
CBSS collection 14
CBSS collection 15
CBSS collection 16
CBSS collection 17
BEDEK, 2007b
CBSS collection 18
CBSS collection 19
245
Nat. Croat. Vol. 20(2), 2011
U~ka (endogean)
VL
NL
MISINTERPRETED LOCALITIES
Jajce, Bosnia and Herzegovina
Ra~i{ka pe~ina (cave, Slovenian cave cadastre no. 942), Ra~ice,
Ra~ice, Starod, Slovenia
ARCANGELI, 1939a*
GRUNER &
TABACARU, 1963**
GIURGINCA, 2009**
GIURGINCA, 2009
BEDEK et al., 2006
GIURGINCA, 2009
*Monte Maggiore is most probably U~ka Mt. in Istria, as indicated by the species name (M. histrianorum,
junior synonym of M. graniger)
**In this paper the locality is placed in Italy, Julian Alps instead in Istria, Croatia.
Fig. 2. Croatian distribution of Mesoniscus graniger.
TRICHONISCIDAE: TRICHONISCINAE
2. Androniscus dentiger Verhoeff, 1908 (Tab. 2, Fig. 3)
Synonyms:
Androniscus africanus Arcangeli, 1939; A. alpinus Verhoeff, 1908; A. carli Strouhal, 1929; A. weberi Verhoeff, 1908; Itea dentigera (Verhoeff, 1908); Stenasellus
hazeltoni (Collinge, 1946); Trichoniscus dentiger (Verhoeff, 1908), Androniscus
dentiger croaticus Strouhal, 1939
246
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
Distribution:
Europe and North Africa, often synanthropically, also known from greenhouses in North America (SCHMALFUSS, 2003)
Type locality:
Santa Margherita, Liguria; Massa; Carrara and Forno, Tuscany, Italy […westlich der Apenninen …. bei S. Margherita… bei Massa und Carrara, oberhalb
Massa ... bei Forno in den apuanischen Bergen (VERHOEFF, 1908)]
Ecological status: Endogean to troglophilic species (VANDEL, 1960b)
Note:
According to VANDEL (1960a) and examined specimens from its type locality, the subspecies A. d. croaticus Strouhal, 1939 is a junior synonym of A. d.
dentiger.
Fig. 3. Croatian distribution of Androniscus dentiger.
Tab. 2. List of recorded localities for Androniscus dentiger
Locality
UTM
Cimerjanka (cave), Cimerplac, Lepoglava, Ravna gora
WM72 C
Region Source
CBSS collection 20
Duplja (cave), Male Pe}ine, Bednjica, Klenovnik, Ravna gora
WM72 C
CBSS collection 21
Ledenica (cave), Male Pe}ine, Bednjica, Lepoglava, Ravna
gora
WM72 C
CBSS collection 22
Rudnik u Velikoj Sotinskoj (mine), Velika Sotinska, Lepoglava,
Ravna gora
WM72 C
CBSS collection 23
[pilja kod Dobre vode (cave), potok Kamenica, Lepoglava,
Ravna gora
WM72 C
CBSS collection 24
247
Nat. Croat. Vol. 20(2), 2011
CBSS collection 25
LANGHOFFER, 1915a1
Velika {pilja (cave), Veliki Goranec, Klenovnik, Ravna gora
WM72 C
STROUHAL, 1939d
ARCANGELI, 1940
KARAMAN, 1966
BEDEK et al., 2006
Vihra (cave), potok Kamenica, Lepoglava, Ravna gora
WM72 C
CBSS collection 26
Vila (cave), Pintari}i, Kameni~ko Podgorje, Klenovnik, Ravna gora
WM72 C
CBSS collection 27
Han`ekova {pilja (cave), Belec grad, Ivan{~ica, Zlatar
WM81 C
CBSS collection 28
Jama ispod Belec grada (cave), Belec grad, Ivan{~ica, Zlatar
WM81 C
CBSS collection 29
CBSS collection 30
LANGHOFFER, 1915a1
Cerjanska {pilja (cave), Klenovnik, Ravna gora
WM82 C
STROUHAL, 1939d
ARCANGELI, 1940
KARAMAN, 1966
Cinkalica 1 (cave), Ku~a gora, @akli, Veliki Goranec, Klenovnik WM82 C
CBSS collection 31
Cinkalica 3 (cave), Lepoglava, Ravna gora
WM82 C
CBSS collection 32
Goranska Klepa (cave), Lepoglava, Ravna gora
WM82 C
CBSS collection 33
Kukelj sjever (cave), Kukelj, Galinci, Lepoglava, Ravna gora
WM82 C
CBSS collection 34
[pilja pod Butinom (cave), Mala Sotinska, Klenovnik,
Lepoglava, Ravna gora
WM82 C
CBSS collection 35
CBSS collection 36
LANGHOFFER, 1915a1
Vindija (cave), Zlogonje, Klenovnik, Ravna gora
WM82 C
STROUHAL, 1939d
ARCANGELI, 1940
KARAMAN, 1966
CBSS collection 37
LANGHOFFER, 1915a1
Zdenec pri Ciglaru (cave), Ciglar, Klenovnik, Ravna gora
WM82 C
STROUHAL, 1939d
STROUHAL, 1939e
ARCANGELI, 1940
KARAMAN, 1966
Plitvi~ka {pilja (cave), Plitvica, Pe{~enica Vini~ka, Vinica
WM83 C
CBSS collection 38
[koljka~a (cave), Plitvica, Pe{~enica Vini~ka, Vinica
WM83 C
CBSS collection 39
Cingulica pod @idovinom (cave), @idovina, Mikul~i}i Gornji,
Ivan{~ica, Zlatar
WM91 C
CBSS collection 40
From SE Italy to the Cetina region
STROUHAL, 1940a
Istria
FRANKENBERGER &
STROUHAL, 1940
STROUHAL, 1939d
NL
KARAMAN, 1966
FRANKENBERGER &
STROUHAL, 1940
Croatia
VANDEL, 1960a
SCHMÖLZER, 1965a
POTO^NIK, 1989
1
Trichoniscus roseus
248
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
3. Androniscus roseus buccariensis Verhoeff, 1930 (Tab. 3, Fig. 4)
Distribution:
Type locality:
Slovenia, Croatia (POTO^NIK, 1989)
Uvala kod Bakra (bay), Bakar, Hrvatsko primorje, Croatia [Bucht von Buccari (VERHOEFF, 1930)]
Ecological status: Troglophilic (STROUHAL, 1940a; BUTUROVI], 1957)
Tab. 3. List of recorded localities for Androniscus roseus buccariensis
Locality
UTM
Region Source
Vrana (lake), Cres
VK56
NL
[pilja iznad ceste (cave), brodogradili{te V. Lenac, Kostrena, Rijeka VL51
NL
Bunker blizu uvale Perilo (bunker), Perilo, Kostrena, Rijeka
VL61
NL
Rudnik iznad uvale Martin{~ica (mine), Martin{~ica, Kostrena,
Rijeka
VL61
NL
[pilja u uvali Sve`anj (cave), uvala Sve`anj, Paveki, Bakar
VL61
NL
Urinjska {pilja (cave), Urinj, Kostrena, Rijeka
VL61
NL
VERHOEFF, 1938
KARAMAN, 1966
CBSS collection 41
CBSS collection 42
LUKI] et al., 2011
CBSS collection 43
LUKI] et al., 2011
CBSS collection 44
LUKI] et al., 20111
CBSS collection 45
LUKI] et al., 20111
VERHOEFF, 1930
STROUHAL, 1938
Uvala kod Bakra (bay), Bakar
VL61
NL
FRANKENBERGER &
STROUHAL, 1940
KARAMAN, 1966
Cres
NL
FRANKENBERGER &
STROUHAL, 1940
VERHOEFF, 1940
From SE Italy to the Cetina region
STROUHAL, 1940a
VERHOEFF, 1931c
SCHMÖLZER, 1965a
Croatia
POTO^NIK, 1989
KARAMAN &
^EMERLI], 2002
1
Androniscus sp.
4. Androniscus roseus histrianorum (Arcangeli, 1932) (Tab. 4, Fig. 4)
Synonyms:
Trichoniscus roseus histrianorum Arcangeli, 1932; Androniscus histrianorum (Arcangeli, 1932)
Distribution:
Croatia (POTO^NIK, 1989)
Type locality:
[pilja na Gradini kod Premanture (cave), Premantura, Pula, Istria, Croatia
[caverna presso Promontore, dintorini di Pola, Istria (ARCANGELI, 1932)]
Ecological status: Troglophilic (STROUHAL, 1940a; BUTUROVI], 1957)
249
Nat. Croat. Vol. 20(2), 2011
Tab. 4. List of recorded localities for Androniscus roseus histrianorum
Locality
UTM
Region Source
CBSS collection 46
ARCANGELI, 1932
ARCANGELI, 1938
[pilja na Gradini kod Premanture (cave), Premantura, Pula
VK16
NL
WOLF, 1938
FRANKENBERGER &
STROUHAL, 1940
GOTTSTEIN et al., 2001
BEDEK et al., 2006
From SE Italy to the Cetina region
STROUHAL, 1940a
Croatia
POTO^NIK, 1989
5. Androniscus roseus ocellatus (Kesselyak, 1930) (Tab. 5, Fig. 4)
Synonyms:
Distribution:
Type locality:
Ecological status:
Androniscus stygius ocellatus Kesselyak, 1930
Croatia (KESSELYAK, 1930)
Velebit?, Croatia [in Kroatien, vermutlich im Velebitgebirge (KESSELYAK, 1930)]
unknown
Tab. 5. List of recorded localities for Androniscus roseus ocellatus
Locality
UTM
Velebit?
Region Source
NL?
KESSELYÁK, 1930
ARCANGELI, 1938
6. Androniscus roseus roseus (C. Koch, 1838) (Tab. 6, Fig. 4)
Synonyms:
Itea rosea C. Koch, 1838; Trichoniscus roseus (C. Koch, 1838); T. roseus var.
nanus Carl, 1908; Androniscus carynthiacus Verhoeff, 1908; Trichoniscus (Androniscus) carynthiacus (Verhoeff, 1908); Philougria rosea (C. Koch, 1838)
Distribution:
From eastern France to Romania, north to southern Germany (not in Bulgaria and Greece) (SCHMALFUSS, 2003)
Type locality:
Germany [Deutschlands (C. KOCH, 1838)]
Ecological status: Troglophilic (STROUHAL, 1940a)
Tab. 6. List of recorded localities for Androniscus roseus roseus
Locality
UTM
Region Source
Porto Madonna (cave), Savudrija
UL83
NL
Dru{ka pe} (cave), Mo{}eni~ka draga, Potoki, Lovran
VL41
NL
VERHOEFF, 1929b
WOLF, 1938
CBSS collection 47
Oporovina (cave), Oraji, Tuli{evica, U~ka, Lovran
VL41
NL
CBSS collection 48
Vele vode (sinkhole), Crni Lug
VL73
M
CBSS collection 49
250
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
LANGHOFFER, 1912
Ote{ka pe}ina (cave), Peru{i}, Gospi}
WK34
M
LANGHOFFER, 1915b
CSIKI, 1926
WOLF, 1938
Su{ik (sinkhole), Dre`ni~ko polje, Dre`nica, Velika Kapela
WL00
M
CBSS collection 50
Hajdu~ka pe}ina (cave), Bistrac, Gornje Dubrave, Ogulin
WL21
M
LANGHOFFER, 1915a
Izvor {pilja kod Tro{marije (cave), kanjon Dobre, Tro{marija,
Ogulin
WL21
M
CBSS collection 51
Su{ica (sinkhole), Rebi}a Glavica, Generalski Stol
WL21
M
CBSS collection 52
Tounj~ica (cave), Kanjon Tounj~ice, Tounj
WL21
M
CBSS collection 53
Bibi~ka {pilja (cave), kanjon Ribnjaka, Munjasi, Tro{marija
WL22
M
CBSS collection 54
Danetska pe}ina (cave), kanjon Dobre, Dani, Bosiljevo
WL22
M
CBSS collection 55
Dragina pe}ina na Dobri (cave), kanjon Dobre, Grabrk,
Tro{marija, Karlovac
WL22
M
CBSS collection 56
\ukina velika {pilja (cave), kanjon Ribnjaka, Tro{marija,
Karlovac
WL22
M
CBSS collection 57
[pilja kod Podumolskog mlina (cave), kanjon Dobre,
Podumol, Bosiljevo
WL22
M
CBSS collection 58
[pilja kod slapa kod Kasuni}a (cave), kanjon Dobre, Kasuni}i,
Generalski Stol
WL22
M
CBSS collection 59
[pilja u kanjonu Dobre ispod Podjak{i}a (cave), kanjon Dobre,
WL22
Podjak{i}i, Generalski Stol
M
CBSS collection 60
Cepinka (cave), Boi}i, So{ice, @umberak
WL26
C
CBSS collection 61
Jama u Vrloj strani (cave), Gudalji Radatovi}i, @umberak
WL26
C
CBSS collection 62
Pu{ina (cave), Jezernice, So{ice, @umberak
WL26
C
CBSS collection 63
Ledeni~ka {pilja (cave), Rebrovi}i, Tounj
WL31
M
LANGHOFFER, 1915a
Lipa na Protulipi (cave), Stare{ini}i, Protulipa, Bosiljevo
WL32
C
CBSS collection 64
Jastrebina (cave), Frketi} Selo, Novigrad na Dobri, Karlovac
WL33
C
CBSS collection 65
Jama pod Gradom (cave), Dugi vrh, Cerovica, Kra{i},
@umberak
WL35
C
CBSS collection 66
LANGHOFFER, 1912
WL35
C
CSIKI, 1926
Stankova {pilja (cave), kanjon Kupe, Kamanje, Ozalj, Karlovac WL35
C
CBSS collection 67
Vrlovka (cave), kanjon Kupe, Kamanje, Ozalj, Karlovac
WL35
C
CBSS collection 68
[pilja pod Vrajinim stijenama (cave), Toma{evci, So{ice,
@umberak
WL36
C
CBSS collection 69
Vranja~ka {pilja (cave), Slapnica, Medven Draga, Kra{i},
@umberak
WL36
C
CBSS collection 70
@idovske ku}e (cave), Cerovica, So{ice, @umberak
WL37
C
CBSS collection 71
Dvoga}a (cave), Ponikve, Glavica, Zagreb, Medvednica
WL67
C
CBSS collection 72
Horvatove stube (soil), Sljeme, Zagreb, Medvednica
WL78
C
CBSS collection 73
Ozaljska {pilja (cave), Ozalj, Karlovac
WOLF, 1938
Croatia
PAOLETTI, 1978
251
Nat. Croat. Vol. 20(2), 2011
Fig. 4. Croatian distribution of Androniscus roseus.
7. Androniscus stygius microcavernicolus Kesselyak, 1931
(Tab. 7, Fig. 5)
Synonyms:
Androniscus roseus microcavernicolus Kesselyak, 1931; A. microcavernicolus
Kesselyak, 1931
Distribution:
Croatia (POTO^NIK, 1989) [in this paper the author considers this subspecies
to be present also in Slovenia, but he does not confirm this in a subsequent
review (POTO^NIK, 1992). No further records for Slovenia are known].
Type locality:
[pilja na U~ki (cave), Lovran, U~ka, Istria, Croatia [namenlosen Höhle an
dem Monte Maggiore (Istrien) (KESSELYAK, 1931)]
Ecological status: This subspecies has been considered to be troglophilic (STROUHAL, 1940a;
BUTUROVI], 1957), but since all specimens were collected only deep in
caves and are depigmented and blind, we prefer to consider this taxon as
troglobiotic.
Tab. 7. List of recorded localities for Androniscus stygius microcavernicolus
Locality
UTM
Region Source
Jama kod crvenog gloga (cave), Mala U~ka, U~ka
VL31
NL
CBSS collection 74
Jama na Balagu{tini (cave), Dobre~, Poklon, U~ka
VL31
NL
CBSS collection 75
252
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
Mala pe} (cave), Molinarska draga, U~ka
VL31
NL
CBSS collection 76
Ov~ja pe} (cave), Molinarska draga, U~ka
VL31
NL
CBSS collection 77
Petr~eva jama (cave), Petrc, Podmaj, Mala U~ka, U~ka
VL31
NL
CBSS collection 78
Rupa I (cave), Podmaj, Mala U~ka, U~ka
VL31
NL
CBSS collection 79
Rupa II (cave), Podmaj, Mala U~ka, U~ka
VL31
NL
CBSS collection 80
Topolska pe} (cave), Topol, Poklon, U~ka
VL31
NL
CBSS collection 81
Jama ispod Tomini}evog brega (cave), Tomini}ev breg, Brest
pod U~kom, Lupoglav, ]i}arija
VL32
NL
CBSS collection 82
Klanj~eva pe} (cave), Brest pod U~kom, ]i}arija
VL32
NL
CBSS collection 83
Pe}ina na Crekvici 1 (cave), Vojni poligon iznad Bresta, Brest
pod U~kom, ]i}arija
VL32
NL
CBSS collection 84
Pe}ina poli Vrti}a (cave), Brest pod U~kom, ]i}arija
VL32
NL
CBSS collection 85
Jama pod Velikim Knezgradom (cave), Veliki Knezgrad,
Tuli{evica, Lovran, U~ka
VL41
NL
CBSS collection 86
Jama na Mahencu 2 (cave), Mahenac, Veprinac, Opatija
VL42
NL
CBSS collection 87
Jama u Velim Sapcima 3 (cave), Mahenac, Veprinac, Opatija
VL42
NL
CBSS collection 88
Pe}ina kod sela Puhari (cave), Puhari, Veprinac, Opatija
VL42
NL
CBSS collection 89
KESSELYÁK, 1931
ARCANGELI, 1938
WOLF, 19381
FRANKENBERGER &
STROUHAL, 1940
[pilja na U~ki (cave), U~ka
VL
NL
SCHMÖLZER, 1965a
KARAMAN, 1966
GOTTSTEIN et al., 2001
SCHMIDT & LEISTIKOW,
2004
BEDEK et al., 2006
JAL@I] et al., 2010
From SE Italy to the Cetina region
STROUHAL, 1940a
Croatia
POTO^NIK, 1989
8. Androniscus stygius stygius (Nemec, 1897) (Tab. 8, Fig. 5)
Synonyms:
Distribution:
Type locality:
Trichoniscus stygius Nemec, 1897; Androniscus cavernarum Verhoeff, 1908
SE-Austria, NE-Italy, Slovenia (SCHMALFUSS, 2003); Croatia (POTO^NIK, 1989)
Grotta dell'Orso (cave, Friuli Venezia Giulia cadastre no. 33, historical cadastre no. 7/VG), Trieste, Italy [Gabroviza-Grotte (NEMEC, 1897)]
Ecological status: This subspecies has been considered to be troglophilic (STROUHAL, 1940a;
BUTUROVI], 1957), but since all specimens were collected deep in caves
and are depigmented and blind, we prefer to consider this taxon as troglobiotic.
253
Nat. Croat. Vol. 20(2), 2011
Tab. 8. List of recorded localities for Androniscus stygius stygius
Locality
UTM
Region Source
Vidovi}a {pilja (cave), Dre`ni~ko polje, Dre`nica, Velika
Kapela, Ogulin
WL00
M
CBSS collection 90
\ulin ponor – Medvedica (cave system), Ogulin
WL11
M
CBSS collection 91
Jama 3 (cave), Malik, Ku{trovka, Tro{marija
WL12
M
CBSS collection 92
Jama na Bertiji 1 (cave), Bertija, Malik, Ku{trovka, Tro{marija
WL12
M
CBSS collection 93
Jama na Bertiji 2 (cave), Bertija, Malik, Ku{trovka, Tro{marija
WL12
M
CBSS collection 94
Ku{trovka (cave), Ku{trovka, Tro{marija
WL12
M
CBSS collection 95
Ambarac (sinkhole), O{tarije, Ogulin
WL20
M
CBSS collection 96
Mandelaja (cave), O{tarije, Ogulin
WL20
M
CBSS collection 97
[pilja u Mekoti (cave), Ko{are, Tounj
WL20
M
CBSS collection 98
Guge~ka pe}ina (cave), Gornje Dubrave, Tounj
WL21
M
CBSS collection 99
Hajdu~ka pe}ina (cave), Bistrac, Gornje Dubrave, Ogulin
WL21
M
CBSS collection 100
Mika{inovi}a {pilja (cave), Bistrica, Mika{inovi}i, Gornje
Dubrave, Tounj
WL21
M
CBSS collection 101
Pe}ina u Guge~kom usjeku (cave), Gornje Dubrave, Tounj
WL21
M
Per~evi}a {pilja (cave), Kanjon Tounj~ice, Tounj
WL21
M
CBSS collection 103
Su{ica (sinkhole), Rebi}a Glavica, Generalski Stol
WL21
M
CBSS collection 104
[utova~ka pe}ina (cave), kanjon Dobre, Tro{marija
WL21
M
CBSS collection 105
Tamnica (cave), Potok Tounjski, Tounj
WL21
M
CBSS collection 106
Tounj~ica (cave), Kanjon Tounj~ice, Tounj
WL21
M
CBSS collection 107
CNHM collection 1
CBSS collection 102
Vreli} (cave), Donje Dubrave, Generalski Stol
WL21
M
CBSS collection 108
Bibi~ka {pilja (cave), kanjon Ribnjaka, Munjasi, Tro{marija
WL22
M
CBSS collection 109
Jama II kod Koli}a (cave), Koli}i, Generalski Stol
WL22
M
CBSS collection 110
Jama kod Kasuni}a (cave), Kasuni}i, Generalski Stol
WL22
M
CBSS collection 111
Jama pod Gradanom (cave), Koli}i, Generalski Stol
WL22
M
CBSS collection 112
Jama u kanjonu Dobre ispod Grabrka (cave), kanjon Dobre,
Podumol, Bosiljevo
WL22
M
CBSS collection 113
Ledenica u [peharima (cave), [pehari, Bosiljevo
WL22
M
CBSS collection 114
Pave}a {pilja (cave), kanjon Dobre, Soline, Bosiljevo
WL22
M
CBSS collection 115
[pilja u kanjonu Dobre ispod Podjak{i}a (cave), kanjon Dobre,
Podjak{i}i, Generalski Stol
WL22
M
CBSS collection 116
Vodena jama (cave), Koli}i, Generalski Stol
WL22
M
CNHM collection 2
@u~ki (cave), ^a`inka, Generalski Stol
WL22
M
CBSS collection 117
Ledeni~ka {pilja (cave), Rebrovi}i, Tounj
WL31
M
CBSS collection 118
Lipa na Protulipi (cave), Stare{ini}i, Protulipa, Bosiljevo
WL32
C
CBSS collection 119
Vrlovka (cave), kanjon Kupe, Kamanje, Ozalj, Karlovac
WL35
C
CBSS collection 120
Croatia
ARGANO et al., 1982
254
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
Fig. 5. Croatian distribution of Androniscus stygius.
9. Androniscus wolfi Strouhal, 1939 (Tab. 9, Fig. 6)
Synonyms:
Distribution:
Type locality:
Androniscus dentiger wolfi Strouhal, 1939
Croatia (POTO^NIK, 1989)
Vla{ka pe} (cave), Kozica, Senj, Hrvatsko primorje, Croatia [Vla{ka Pe}ina
bei Novi; BB Nr. 551 (STROUHAL, 1939d)]
Ecological status: Troglophilic (STROUHAL, 1940a; BUTUROVI], 1957)
Note:
After examination of specimens from the type locality, A. wolfi is a valid
species belonging to the subgenus Roseoniscus and not Dentigeroniscus, as
already stated by VANDEL (1960a).
Tab. 9. List of recorded localities for Androniscus wolfi
Locality
Vla{ka pe} (cave), Kozica, Senj
Mokra~a (cave), Paklenica, Starigrad, Velebit
UTM
Region Source
CNHM collection 3
STROUHAL, 1939d
FRANKENBERGER &
STROUHAL, 1940
A
RCANGELI, 1940
VK99 NL
KARAMAN, 1966
GOTTSTEIN et al., 2001
VUJ^I]-KARLO & RA\A, 2004
BEDEK et al., 2006
WK41 SL
CBSS collection 121
255
Nat. Croat. Vol. 20(2), 2011
Southern Velebit, Velebit
National Park Paklenica, Velebit
From SE Italy to the Cetina region
BEDEK et al., 2000
VUJ^I]-KARLO & RA\A, 20041
STROUHAL, 1940a
SCHMÖLZER, 1965a
POTO^NIK, 1989
Croatia
1
Androniscus cf. wolfi
Fig. 6. Distribution of Androniscus wolfi.
10. Aegonethes antilocapra Frankenberger, 1938 (Tab. 10, Fig. 7)
Distribution:
Type locality:
Croatia: the islands of Kor~ula and Vis (SCHMALFUSS, 2003)
Tabaina {pilja (cave), Vela Luka, the island of Kor~ula; [pilja pod Humom
(cave), Komi`a, the island of Vis; Caretova {pilja (cave), Vis, the island of
Vis and [pilja na Bardarovici (cave), Bardarovica, Korita, Vis, the island of
Vis, Dalmatia, Croatia [Tabaina spilja, environs de la ville de Vela Luka, île
de Kor}ula. – Spilja pod Humom, environs de la ville de Komi`e, île de Vis
(Lissa). – Ceretova spilja kod Visa, environs de la ville de Vis, île de Vis. –
Bardarovica, environs du village Korita, îlle de Vis. (FRANKENBERGER, 1938a)]
Ecological status: Troglobiotic (STROUHAL, 1940a; BUTUROVI], 1957)
Tab. 10. List of recorded localities for Aegonethes antilocapra
Locality
UTM
[pilja uz cestu kod Komi`e (cave), Komi`a, Vis
WH86 SL
Region Source
CBSS collection 122
256
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
FRANKENBERGER, 1938a
Caretova {pilja (cave), Vis, Vis
WH96 SL
Kraljicina {pilja (cave), Oklju~na, Komi`a, Vis
WH96 SL
CBSS collection 123
Rupa na Ribinovoj bandi (cave), Ribinova banda, Kostirna,
Vis, Vis
WH96 SL
CBSS collection 124
KARAMAN, 1966
GIROMETTA, 19141
[pilja na Bardarovici (cave), Bardarovica, Korita, Vis, Vis
WH96 SL
FRANKENBERGER, 1938a
[pilja od Vore (cave), Kostirna, Vis, Vis
WH96 SL
CNHM collection 4
[pilja pod Humom (cave), Komi`a, Vis
WH96 SL
Tanki rot (cave), Podhumlje, Komi`a, Vis
WH96 SL
CBSS collection 125
Veliki rudnik (mine, cave), Kostirna, Vis, Vis
WH96 SL
CBSS collection 126
KARAMAN, 1966
FRANKENBERGER, 1938a
KARAMAN, 1966
CBSS collection 127
FRANKENBERGER, 1938a
Tabaina {pilja (cave), Vela Luka, Kor~ula
XH45
SL
FRANKENBERGER &
STROUHAL, 1940
KARAMAN, 1966
BEDEK et al., 2006
OZIMEC, 2010
Vis
SL
Fig. 7. Distribution of Aegonethes antilocapra.
FRANKENBERGER &
STROUHAL, 1940
257
Nat. Croat. Vol. 20(2), 2011
STROUHAL, 1939a
SCHMÖLZER, 1965a
TABACARU, 1996
Kor~ula and Vis
SL
GOTTSTEIN et al., 2001
GOTTSTEIN MATO^EC et
al., 2002
SCHMALFUSS, 2003
From the Neretva to the S Montenegro region
SL
POTO^NIK, 1989
Croatia
1
STROUHAL, 1940a
Titanethes albus
11. Aegonethes cervinus (Verhoeff, 1931) (Tab. 11, Fig. 8)
Synonyms:
Distribution:
Illyrionethes cervinus Verhoeff, 1931
Italy: Gargano, Croatia: Dubrovnik and the islands of Kor~ula and Mljet
(SCHMALFUSS, 2003)
Type locality:
Pi{urka (cave), Kor~ula, the island of Kor~ula, Dalmatia, Croatia [Paganetti-Höhle auf der dalmatinischen Insel Curzola (VERHOEFF, 1931a, b)]
Ecological status: Troglobiotic (STROUHAL, 1940a; BUTUROVI], 1957)
Tab. 11. List of recorded localities for Aegonethes cervinus
Locality
UTM
Garden of Biological Institute Lapad (endogean?), Lapad,
Dubrovnik
BN52
Karle (cave), Gru`, Dubrovnik
BN62
\urovi}a {pilja (cave), \urovi}i, Cavtat, Konavle
BN71
[ipun (cave), Cavtat, Dubrovnik
BN71
Hrastovska {pilja (cave), Hrastovsko polje, Lastovo, Lastovo
XH53
Jama pod Velji vrh (cave), Velji vrh, Lastovo, Lastovo
XH53
Medvi|a ropa (cave), Skrivena Luka, Lastovo
XH53
Ra~a (cave), Rada{ do, Skrivena Luka, Lastovo
XH53
[pilja na Pozalicu (cave), Pr`ina, Skrivena Luka, Lastovo
XH53
[pilja pod Ka{tel (cave), Ka{tel, Lastovo, Lastovo
XH53
Region Source
BUTUROVI], 1955b
SL
KARAMAN, 1966
STROUHAL, 1939a
STROUHAL, 1939e
SL
ARCANGELI, 1947
KARAMAN, 1966
CBSS collection 128
STROUHAL, 1939a
STROUHAL, 1939e
SL
ARCANGELI, 1947
KARAMAN, 1966
STROUHAL, 1939a
STROUHAL, 1939e
SL
ARCANGELI, 1947
KARAMAN, 1966
CBSS collection 129
SL
OZIMEC et al., 2008
CBSS collection 130
SL
OZIMEC et al., 2008
CBSS collection 131
SL
OZIMEC et al., 2008
CBSS collection 132
SL
OZIMEC et al., 2008
CBSS collection 133
SL
OZIMEC et al., 2008
CBSS collection 134
SL
OZIMEC et al., 2008
258
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
Jama na Odvoru (cave), Strmena strana, Ra~i{}e, Kor~ula
Samograd (cave), Ra~i{}e, Kor~ula
Jakasova {pilja (cave), Glava, Rasohatica, @rnovo, Kor~ula
Jama pod Kola~ (cave), Vu~ja, Postrane, @rnovo, Kor~ula
XH66
XH66
XH75
XH75
SL
SL
SL
SL
Pi{urka (cave), Kor~ula, Kor~ula
XH75 SL
Jama na Brekalcima (cave), Pola~e, Mljet
XH93 SL
Jama u Velikom Desajdubu (cave), Veliki Desajdub, Pola~a,
Mljet
XH93 SL
Male ponte (cave), Grabova glava, Ropa, Mljet
XH93 SL
[pilja kod Nerezinog dola (cave), Nerezin do, Ropa, Mljet
XH93 SL
[pilja kod Solina (cave), Soline, Mljet
Jama za Svetim Spasom (cave), Babino Polje, Mljet
Mala {pilja (cave), Blata, Mljet
XH93 SL
YH03 SL
YH03 SL
Movrica (cave), Dra~evac, Babino Polje, Mljet
YH03 SL
Osta{evica (cave), Dra~evac, Babino Polje, Mljet
YH03 SL
Velika {pilja (cave), Blata, Mljet
YH03 SL
Gali~njak (cave), Okuklje, Mljet
Jama za Svetim Ivanom (cave), Babino Polje, Mljet
Me|ugra|en (cave), Veliki grad, Babino Polje, Mljet
YH13 SL
YH13 SL
YH13 SL
CBSS collection 135
CBSS collection 136
CBSS collection 137
CBSS collection 138
CBSS collection 139
VERHOEFF, 1931a
VERHOEFF, 1931b
FRANKENBERGER, 1938a
STROUHAL, 1938
STROUHAL, 1939a
FRANKENBERGER &
STROUHAL, 1940
ARCANGELI, 1947
KARAMAN, 1966
ARGANO et al., 1982
SCHMIDT & LEISTIKOW,
2004
BEDEK et al., 2006
OZIMEC, 2010
CNHM collection 5
CBSS collection 140
CNHM collection 6
CBSS collection 141
CBSS collection 142
CBSS collection 143
FRANKENBERGER, 1938a
STROUHAL, 1939a
ARCANGELI, 1947
KARAMAN, 1966
CBSS collection 144
CBSS collection 145
CBSS collection 146
CBSS collection 147
STROUHAL, 1939a
STROUHAL, 1939c
STROUHAL, 1939e
ARCANGELI, 1947
CBSS collection 148
FRANKENBERGER, 1938a
STROUHAL, 1939a
STROUHAL, 1939e
ARCANGELI, 1947
KARAMAN, 1966
CBSS collection 149
FRANKENBERGER, 1938a
STROUHAL, 1939a
ARCANGELI, 1947
KARAMAN, 1966
CBSS collection 150
CBSS collection 151
CBSS collection 152
259
Nat. Croat. Vol. 20(2), 2011
Mljet, surroundings of Dubrovnik
SL
Kor~ula and Mljet
SL
Kor~ula and Mljet, surroundings of Dubrovnik
Mljet
From The Neretva to the S Montenegro region
Croatia
SL
SL
SL
FRANKENBERGER &
STROUHAL, 1940
SCHMÖLZER, 1965a
TABACARU, 1996
GOTTSTEIN et al., 2001*
GOTTSTEIN MATO^EC et
al., 2002*
SCHMALFUSS, 2003
OZIMEC & BEDEK, 2005
STROUHAL, 1940a
POTO^NIK, 1989
* Vis is also given, but there are no literature data for the island of Vis, so this must be a mistake.
Fig. 8. Croatian distribution of Aegonethes cervinus.
12. Alpioniscus balthasari (Frankenberger, 1937) (Tab. 12, Fig. 9)
Synonyms:
Distribution:
Type locality:
Illyrionethes balthasari Frankenberger, 1937
Southern Croatia, Bosnia, Herzegovina (SCHMALFUSS, 2003)
Kotlu{a (cave), Kotlu{a, Civljane, Vrlika, Dalmatia, Croatia [eine Höhle in
der Umgebung der Stadt Knin; es ist dies jene Höhle, die in der Nähe des
Dorfes Kotlu{a am westlichen Fusße des Dinara-Gebirges; »Höhle mit
Wasser« (FRANKENBERGER, 1937)]
Ecological status: Troglobiotic (STROUHAL, 1940a; BUTUROVI], 1957)
260
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
Tab. 12. List of recorded localities for Alpioniscus balthasari
Locality
^ude (cave), Zrmanja, Obrovac
Rastova~a (cave), Zaton, Rastovac, Obrovac
UTM
WJ59
WJ59
Region Source
SL
CBSS collection 153
SL
CBSS collection 154
CBSS collection 155
SL
BEDEK, 2007a
CBSS collection 156
SL
BEDEK, 2007a
SL
CBSS collection 157
CBSS collection 158
SL
BEDEK, 2007a
SL
CBSS collection 159
CBSS collection 160
SL
BEDEK, 2007a
CBSS collection 161
SL
BEDEK, 2007a
CBSS collection 162
SL
BEDEK, 2007a
CBSS collection 163
SL
BEDEK, 2007a
CBSS collection 164
SL
BEDEK, 2007a
SL
CBSS collection 165
CBSS collection 166
SL
BEDEK, 2007a
GIROMETTA, 19131
LANGHOFFER, 1915a1
SL
STROUHAL, 1939b
Miljacka I (cave), Krka, Oklaj, Knin
WJ67
Miljacka II (cave), Krka, Oklaj, Knin
WJ67
[pilja kod Mlina na Miljacki (cave), Krka, Oklaj, Knin
WJ67
Golubnja~a (cave), Bukovica, Ka{tel @egarski, Obrovac
WJ68
Velika pe}ina (cave), Gornji ^abri}i, Crnopac, Velebit, Gra~ac
WJ69
Martina jama (cave), Krka, Skradin
WJ75
Drenova~a (cave), Lali}a stanovi, Lali}i, Krka, Kistanje
WJ76
Tavnica (cave), Lali}a ograde, Lali}i, Krka, Kistanje
WJ76
Tri{i}a jama (cave), Lali}a ograde, Lali}i, Krka, Kistanje
WJ76
Pr~ina (cave), Kistanje
WJ77
Kusa nad Manastirskim lukama (cave), Krupa, Obrovac
WJ79
Stra`benica (cave), Vrpolje, [ibenik
WJ83
Zve~ara (cave), Perkovi}, Sitno Donje, [ibenik
WJ83
Stara jametina (cave), Pun~ka draga, G. Krni}i, Konjevrate,
[ibenik
WJ84
SL
[karin Samograd (cave), [kari}i, Konjevrate, [ibenik
WJ84
SL
Topla pe} (cave), Klju~, ^ikola, Skradin
WJ85
SL
Jama na polju (cave), Vrsine, Trogir
WJ92
SL
Koli{tina (cave), Prgomet, Trogir
WJ92
SL
Golubinjka jama (cave), Koprno, [ibenik
WJ94
SL
Smokvica (cave), Koprno, [ibenik
WJ94
SL
CBSS collection 167
MZUF collection 1
CBSS collection 168
BEDEK, 2007a
CBSS collection 169
GIROMETTA, 19141
LANGHOFFER, 1915a1
STROUHAL, 1939b
GIROMETTA, 19131
LANGHOFFER, 1915a1
STROUHAL, 1939b
GIROMETTA, 19141
LANGHOFFER, 1915a1
STROUHAL, 1939b
GIROMETTA, 19141
LANGHOFFER, 1915a1
STROUHAL, 1939a
STROUHAL, 1939b
KARAMAN, 1966
BEDEK, 2007a
261
Nat. Croat. Vol. 20(2), 2011
Kad jaganjci utihnu (cave), Lug Paripov, Ljuboti}, Tribalj
Kru{~ica
Ivina jama (cave), Velika Paklenica, Velebit, Starigrad
WK21
SL
CBSS collection 170
WK30
SL
Manita pe} (cave), Velika Paklenica, Velebit, Starigrad
WK30
SL
Pisanica (cave), Jurline, Paklenica, Velebit, Starigrad
Bu{ljeta (cave), Pisak, Bojinac, Velebit, Starigrad
Jama kod Jatare 2 (cave), Jatara, Velebit, Starigrad
Jatara (cave), Jatara, Velebit, Starigrad
Vaganka (cave), Veliki Vaganac, Velebit, Starigrad
WK30
WK31
WK31
WK31
WK31
SL
SL
SL
SL
SL
Gornja Cerova~ka {pilja (cave), Cerovac, Crnopac, Velebit,
Gra~ac
WK70
M
XH69
XH69
SL
SL
CNHM collection 7
CBSS collection 171
MZUF collection 2
BO@I^EVI], 19652
CBSS collection 172
CBSS collection 173
CBSS collection 174
CBSS collection 175
CBSS collection 176
CNHM collection 8
CBSS collection 177
BEDEK et al., 2008
CBSS collection 178
CBSS collection 179
Benova jama (cave), Zavodi, Ravna vla{ka, Biokovo, Tu~epi
Bubnjava~a (cave), Veprinac, Biokovo, Makarska
Jama na putu za Staru {kolu (cave), Sv. Jure, Biokovo,
Makarska
Krjava 1 (cave), Veliko Brdo, Biokovo, Makarska
Krjava 2 (cave), Veliko Brdo, Biokovo, Makarska
Kuna (cave), ^edor, Biokovo, Makarska
Ledenica kod Stare {kole (cave), Sv. Jure, Biokovo, Makarska
Podrum (cave), Gornja Podgora, Biokovo, Podgora
Bo{ak (cave), Lozovci, @upa, Biokovo, Zagvozd
XH69
SL
CNHM collection 9
XH69
XH69
XH69
XH69
XH69
XH79
SL
SL
SL
SL
SL
SL
Gradska spila (cave), @upa, Biokovo, Zagvozd
XH79
SL
CBSS collection 180
CBSS collection 181
CBSS collection 182
CNHM collection 10
CBSS collection 183
CBSS collection 184
CNHM collection 11
CBSS collection 185
XH79
SL
CBSS collection 186
XH79
SL
CBSS collection 187
Jama iznad Sarana~a (cave), Sarana~, Gornje Igrane,
Podgora
Jama kod Tu~epske vilenja~e (cave), Ravna vla{ka, Biokovo,
Tu~epi
Jama na Docima (cave), Sarana~, Gornje Igrane, Biokovo,
Podgora
Jama na grebenu (cave), Prisika, Biokovo, Podgora
Jama za Supinom (cave), Vrata Biokova, Biokovo, Podgora
XH79
SL
CBSS collection 188
XH79
XH79
SL
SL
Jujnovi}a {pilja (cave), Jujnovi}i, Kozica, Biokovo, Podgora
XH79
SL
Mala jama (cave), @upa, Biokovo, Zagvozd
XH79
SL
Matija{eva {pilja (cave), @upa, Biokovo, Zagvozd
XH79
SL
Spila 1 (cave), Jujnovi}i, Kozica, Biokovo, Podgora
XH79
SL
Spila 2 (cave), Jujnovi}i, Kozica, Biokovo, Podgora
XH79
SL
Stonjska pe} donja (cave), @upa, Biokovo, Zagvozd
Stonjska pe} gornja (cave), @upa, Biokovo, Zagvozd
Tu~epska vilenja~a (cave), Ravna vla{ka, Biokovo, Tu~epi
XH79
XH79
XH79
SL
SL
SL
Pe}ina u Carevoj Poski (cave), Jurilji, Vrgorac
XH88
SL
Mala {pilja (cave), Antunovi}i, Kozica, Biokovo, Podgora
Martina jama (cave), Stilja, Vrgorac
XH89
XH89
SL
SL
CBSS collection 189
CBSS collection 190
CNHM collection 12
CBSS collection 191
CBSS collection 192
CNHM collection 13
CBSS collection 193
CNHM collection 14
BUTUROVI], 1955c
KARAMAN, 1966
CNHM collection 15
CBSS collection 194
CBSS collection 195
CBSS collection 196
CBSS collection 197
BUTUROVI], 1955c
KARAMAN, 1966
CNHM collection 16
CBSS collection 198
262
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
Bunarina (cave), Podanak, Uble, Le}evica, Split
XJ03
SL
Jama na Kuku (cave), Rado{i}, Le}evica, Split
XJ03
SL
Jama pod Kragljevcem (cave), Ramljane, Mose}, Sinj
XJ04
SL
Kosilova jama (cave), Brezovac, Dinara, Knin
XJ08
SL
Petrinjka (cave), Vu~evica, Ka{tel Luk{i}, Kozjak, Split
XJ12
SL
Jazinka (cave), Ljuta, Mu}, Mose}, Sinj
XJ13
SL
Kotlu{a (cave), Kotlu{a, Civljane, Cetina, Vrlika
XJ16
SL
Kranjica (cave), Crvene grede, Jare, Vrlika
XJ16
SL
Medi}a pe}ina (cave), Civljane, Cetina, Vrlika
XJ16
SL
Rudeli}a {pilja (cave), Civljane, Cetina, Vrlika
XJ16
SL
Gornja Tuti}eva {pilja (cave), Tuti}i, Civljane, Cetina, Vrlika
XJ17
SL
Gospodska {pilja (cave), ^etnik, Civljane, Cetina, Vrlika
XJ17
SL
Pe}ina iznad Kule (cave), Pekasova draga, Glava{, Civljane,
Cetina, Vrlika
XJ17
SL
Pe}ina na izvoru Cetine (cave), Civljane, Cetina, Vrlika
XJ17
SL
Velika ^ulumova {pilja (cave), Krivodol, Kijevo, Vrlika
XJ17
SL
Bali}eva {pilja (cave), Bali}i, Dugopolje, Mosor, Split
XJ22
SL
T. Ra|a collection 1
GIROMETTA, 19131
LANGHOFFER, 1915a1
STROUHAL, 1939b
GIROMETTA, 19131
LANGHOFFER, 1915a1
STROUHAL, 1939b
GIROMETTA, 19141
STROUHAL, 1939a
KARAMAN, 1966
CNHM collection 17
CBSS collection 199
GIROMETTA, 19131
LANGHOFFER, 1915a1
STROUHAL, 1939b
GIROMETTA, 19141
LANGHOFFER, 1915a1
STROUHAL, 1939a
STROUHAL, 1939b
KARAMAN, 1966
CNHM collection 18
CBSS collection 200
FRANKENBERGER, 1937
FRANKENBERGER, 1939
STROUHAL, 1939a
KARAMAN, 1966
BEDEK et al., 2006
JAL@I] et al., 2010
CBSS collection 201
STROUHAL, 1939a
KARAMAN, 1966
CNHM collection 19
CBSS collection 202
STROUHAL, 1939a
KARAMAN, 1966
CNHM collection 20
CBSS collection 203
CNHM collection 21
CBSS collection 204
STROUHAL, 1939a
KARAMAN, 1966
CBSS collection 205
FRANKENBERGER, 1939
KARAMAN, 1966
CNHM collection 22
CBSS collection 206
GIROMETTA, 19131
GIROMETTA, 19141
LANGHOFFER, 1915a1
STROUHAL, 1939a
STROUHAL, 1939b
KARAMAN, 1966
263
Nat. Croat. Vol. 20(2), 2011
GIROMETTA, 19131
LANGHOFFER, 1915a1
STROUHAL, 1939a
STROUHAL, 1939b
KARAMAN, 1966
STROUHAL, 1939a
KARAMAN, 1966
GIROMETTA, 19141
LANGHOFFER, 1915a1
STROUHAL, 1939a
STROUHAL, 1939b
KARAMAN, 1966
GIROMETTA, 19141
LANGHOFFER, 1915a1
STROUHAL, 1939a
KARAMAN, 1966
GIROMETTA, 19141
LANGHOFFER, 1915a1
STROUHAL, 1939b
GIROMETTA, 19141
LANGHOFFER, 1915a1
STROUHAL, 1939b
GIROMETTA, 19131
GIROMETTA, 19141
LANGHOFFER, 1915a1
STROUHAL, 1939a
STROUHAL, 1939b
KARAMAN, 1966
STROUHAL, 1939a
KARAMAN, 1966
GIROMETTA, 19141
LANGHOFFER, 1915a1
STROUHAL, 1939a
STROUHAL, 1939b
STROUHAL, 1939e
KARAMAN, 1966
CBSS collection 207
STROUHAL, 1939a
KARAMAN, 1966
CBSS collection 208
CBSS collection 209
ERHARD, 1997
STROUHAL, 1939a
KARAMAN, 1966
Jama na Ko{taku (cave), Ko{tak, Klis, Split
XJ22
SL
Kitlica jama (cave), Potravlje, Sinj
XJ25
SL
Trojama (cave), Srijane, Mosor, Split
XJ31
SL
Ledenica na podno`ju Jabukovca (cave), Jabukovac, Mosor,
Split
XJ32
SL
Makluta~a (cave), Dugopolje, Mosor, Split
XJ32
SL
Snije`nica pod Ljubljanom (cave), Ljubljan, Mosor, Split
XJ32
SL
Vranja~a (cave), Kotlenice, Dugopolje, Mosor, Split
XJ32
SL
Jama kod Rado{i}a (cave), Rado{i}, Sinj
XJ33
SL
Jama na Visokoj (cave), Visoka, Rado{i}, Sinj
XJ33
SL
Crvenku{a-Tamnica-Suhi Rumin (cave system), Rumin,
Biteli}, Sinj
XJ34
SL
Govedija jama (cave), Malin, Bajagi}, Sinj
Kosinac (spring), Han, Obrovac Sinjski, Sinj
Suvi Rumin (cave), Rumin, Biteli}, Sinj
XJ34
XJ34
XJ34
SL
SL
SL
Zimska pe}ina (cave), Obrovac Sinjski, Sinj
XJ34
SL
XJ35
SL
CNHM collection 23
XJ35
SL
CBSS collection 210
XJ35
SL
CBSS collection 211
Bila lipotica (cave), Kod`omanove staje, Vrdovo, Dinara,
Biteli}, Sinj
Jama u Konjiku{i gornjoj (cave), Malovani, Vrdovo, Dinara,
Biteli}, Sinj
Od`i}a ponor (sinkhole), Buljani, Gornji Biteli}, Sinj
264
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
Pastirski stan (cave), Vukovi}i, Vrdovo, Dinara, Biteli}, Sinj
Pid`in pit (cave), Biteli}, Sinj
Pometena golubinka (cave), Lovri}a staje, Vrdovo, Dinara,
Biteli}, Sinj
XJ35
XJ35
SL
SL
CBSS collection 212
CBSS collection 213
XJ35
SL
CBSS collection 214
Vodena pe}a (cave), Kod`oman, Bajagi}, Rumin, Sinj
XJ35
SL
@ablja jama (cave), Vrdovo, Dinara, Biteli}, Sinj
XJ35
SL
[pilja kod izvora Rude (cave), Ruda, Otok, Sinj
XJ43
SL
]uli}a jama (cave), Grljak, Bartulovi}i, Biokovo, Brela
XJ50
SL
Drinova II (cave), Drinova, Gornja Brela, Biokovo, Brela
XJ50
SL
Gornja jama na Grljaku (cave), Grljak, Bartulovi}i, Biokovo, Brela
Jama iznad Prozor~i}a (cave), Nevistine stine, Gornja Brela,
Biokovo, Brela
XJ50
SL
CNHM collection 24
CBSS collection 215
CBSS collection 216
CNHM collection 25
CBSS collection 217
CBSS collection 218
CNHM collection 26
CBSS collection 219
CBSS collection 220
XJ50
SL
CBSS collection 221
Jama u Dovnju (cave), Dovanj, Rogoznica, Brela
XJ50
SL
Jamina (cave), Nevistine stine, Gornja Brela, Biokovo, Brela
Plo{nica (cave), Nevistine stine, Gornja Brela, Biokovo, Brela
Pozjata (cave), Jagodnik, Bartulovi}i, Biokovo, Brela
Spasiteljica (cave), Drinova, Gornja Brela, Biokovo, Brela
Svetica (cave), Gornja Brela, Biokovo, Brela
Duboka trepetljika (cave), Kaoci, Muci}i, Biokovo, Zagvozd
Ga}ina jama (cave), Ga~e{e, Biokovo, Zagvozd
Jama iznad Lipog do~i}a (cave), Sv. Jure, Biokovo, Makarska
Jama na Brezdanu (cave), Brezdan, Topi}i, Biokovo, Ba{ka Voda
Jama na Ka~koj glavici 3 (cave), Ka~ka glavica, Kaoci, Muci}i,
Biokovo, Zagvozd
Jama na Mr`dolcu (cave), Mr`dolac, Dedi}i, Sridnja gora,
Zagvozd
Jama pod Sv. Jurom (cave), Sv. Jure, Biokovo, Makarska
Jama u ^ike{ima (cave), ^ike{i, Biokovo, Zagvozd
Jama u trepetljikama (cave), Kaoci, Muci}i, Biokovo, Zagvozd
Kukor (cave), Bast, Biokovo, Ba{ka Voda
Mala jama u stijenama (cave), Spiljice, Kaoci, Muci}i,
Biokovo, Zagvozd
Mu`ina {pilja (cave), Sridnja gora, Svagu{a, Biokovo, Zagvozd
Pasji manastir (cave), Topi}i, Biokovo, Ba{ka Voda
XJ50
XJ50
XJ50
XJ50
XJ50
XJ60
XJ60
XJ60
XJ60
SL
SL
SL
SL
SL
SL
SL
SL
SL
CBSS collection 222
MZUF collection 3
CBSS collection 223
CNHM collection 27
CBSS collection 224
CBSS collection 225
CBSS collection 226
CBSS collection 227
CNHM collection 28
CBSS collection 228
CBSS collection 229
XJ60
SL
CBSS collection 230
XJ60
SL
CBSS collection 231
XJ60
XJ60
XJ60
XJ60
SL
SL
SL
SL
CBSS collection 232
CBSS collection 233
CBSS collection 234
CBSS collection 235
XJ60
SL
CNHM collection 29
XJ60
XJ60
SL
SL
Pe}ina isto~no od sela Svagu{a (cave), Svagu{a, Biokovo,
Zagvozd
XJ60
SL
Pe}ina u Jurkovoj dolini (cave), Jurkova dolina, Biokovo,
Zagvozd
XJ60
SL
Pe}ina u Sridnjoj gori (cave), Sridnja gora, Biokovo, Zagvozd
XJ60
SL
Potkameni~ina jama (cave), Dedi}i, Biokovo, Zagvozd
Pru`ina (cave), Prodani, Biokovo, Zagvozd
Pukotina kod doca Stra`benica (cave), Kaoci, Muci}i,
Biokovo, Zagvozd
Raputinova {pilja (cave), Rastovac, Biokovo, Zagvozd
[pilja do Obilje`ene (cave), Kaoci, Muci}i, Biokovo, Zagvozd
XJ60
XJ60
SL
SL
CBSS collection 236
CBSS collection 237
BUTUROVI], 1955c
KARAMAN, 1966
BUTUROVI], 1955c
KARAMAN, 1966
BUTUROVI], 1955c
KARAMAN, 1966
CBSS collection 238
CBSS collection 239
XJ60
SL
CNHM collection 30
XJ60
XJ60
SL
SL
CBSS collection 240
CBSS collection 241
265
Nat. Croat. Vol. 20(2), 2011
[pilja na Mr`dolcu (cave), Mr`dolac, Dedi}i, Sridnja gora,
Zagvozd
[pilja u Bastu (cave), Bast, Biokovo, Ba{ka Voda
[pilja u Gaju (cave), Brnasi, Biokovo, Zagvozd
Velika {pilja (cave), Bukovci, Sridnja gora, Dedi}i, Zagvozd
Vodena trepetljika (cave), Kaoci, Muci}i, Biokovo, Zagvozd
Mravinju{a jama (cave), Lozovci, @upa, Biokovo, Zagvozd
XJ60
SL
CBSS collection 242
XJ60
XJ60
XJ60
XJ60
XJ70
SL
SL
SL
SL
SL
Bari~evi}a pe}ina (cave), Cetina
XJ
SL
Damljinhova jama (cave), Sinj
XJ
SL
Golubinka (cave), Cetina
XJ
SL
Jama kod Dugopolja (cave), Dugopolje, Split
XJ
SL
Jama na Bala~i (cave), Mosor, Split
XJ
SL
Kozjak pe}ina (cave), Cetina
XJ
SL
Pe}ina kod Sinja (cave), Sinj
XJ
SL
Pe}ina na isto~noj padini Mosor planine (cave), Mosor, Split
XJ
SL
Pe}ina na sjeveroisto~nim padinama Biokova (cave), Biokovo,
XJ
Zagvozd
SL
Pe}ina na Svilaji planini (cave), Svilaja
XJ
SL
Pe}ina sjeveroisto~no od Sinja (cave), Sinj
XJ
SL
Vu~kovi} jama (cave), Cetina
XJ
SL
Zakanova jama (cave), Sinj
XJ
SL
Zrilina pe}ina (cave), Cetina
XJ
SL
CBSS collection 243
CBSS collection 244
CBSS collection 245
CBSS collection 246
CBSS collection 247
STROUHAL, 1939a
KARAMAN, 1966
STROUHAL, 1938
KARAMAN, 1966
STROUHAL, 1939a
KARAMAN, 1966
STROUHAL, 1939a
STROUHAL, 1939a
STROUHAL, 1939e
KARAMAN, 1966
STROUHAL, 1939a
KARAMAN, 1966
STROUHAL, 1939a
KARAMAN, 1966
FRANKENBERGER, 1939
KARAMAN, 1966
FLASAROVÁ, 1967
BUTUROVI], 1955c
KARAMAN, 1966
FRANKENBERGER, 1939
KARAMAN, 1966
FRANKENBERGER, 1939
KARAMAN, 1966
STROUHAL, 1939a
KARAMAN, 1966
STROUHAL, 1938
KARAMAN, 1966
STROUHAL, 1939a
KARAMAN, 1966
STROUHAL, 1939a
STROUHAL, 1939a
KARAMAN, 1966
STROUHAL, 1939a
STROUHAL, 1939e
KARAMAN, 1966
STROUHAL, 1939a
KARAMAN, 1966
STROUHAL, 1939a
KARAMAN, 1966
FRANKENBERGER, 1939
FRANKENBERGER, 1939
FRANKENBERGER, 1939
FRANKENBERGER, 1939
Bobija jama (cave)
Crnomarkovi}a pe}ina (cave)
D`amije (cave)
Pe}ina Stojakovi} (cave)
Podbirnja~a {pilja (cave)
[pilja na Dinari 254 (cave), Dinara
[pilja na Dinari 256 (cave), Dinara
[pilja na Dinari 258 (cave), Dinara
[pilja na Dinari 260 (cave), Dinara
SL
SL
SL
SL
266
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
Vr{ine (cave)
Biokovo
SL
Dinara, W slopes, Vrlika
SL
Cetina spring, surroundings, Civljane, Vrlika
SL
Knin, surroundings
SL
Krka, surroundings
SL
Mosor
SL
Sinj, surroundings
SL
Svilaja
SL
Central Dalmatia
Dalmatia
From Southern Velebit to Biokovo
From the Cetina to the Neretva region
Southern Croatia
SL
SL
Croatia
1
2
SL
STROUHAL, 1939a
STROUHAL, 1939e
KARAMAN, 1966
OZIMEC, 2008
BEDEK, 2011
FRANKENBERGER &
STROUHAL, 1940
KARAMAN, 1966
FRANKENBERGER &
STROUHAL, 1940
STROUHAL, 1938
BEDEK, 2005a, JAL@I] et
al., 2007
FRANKENBERGER &
STROUHAL, 1940
FRANKENBERGER, 1939
FRANKENBERGER &
STROUHAL, 1940
SCHMÖLZER, 1965a
VANDEL, 1946
JAL@I] et al., 2007
STROUHAL, 1940a
SCHMALFUSS, 2003
POTO^NIK, 1989
TABACARU, 1996
Titanethes albus
Titanethes (albus?) dahli
Fig. 9. Croatian distribution of Alpioniscus balthasari.
267
Nat. Croat. Vol. 20(2), 2011
13. Alpioniscus bosniensis (Frankenberger, 1939) (Tab. 13, Fig. 10)
Synonyms:
Distribution:
Type locality:
Illyrionethes bosniensis Frankenberger, 1939
Southern Croatia, Bosnia and Herzegovina (SCHMALFUSS, 2003)
Pe}ina u Donjem Bri{niku (cave), Donji Bri{nik, Tomislavgrad, Bosnia and
Herzegovina [Höhle 12 km weit nach O. von Tomislavgrad (Sw.-Bosnien)
bei der Schule von D. Bri{nik; BB Nr. 272 (FRANKENBERGER, 1939)]
Ecological status: Troglobiotic (STROUHAL, 1940a; BUTUROVI], 1957)
Tab. 13. List of recorded localities for Alpioniscus bosniensis
Locality
UTM
Baba {pilja (cave), [o{i}i, Gornje Igrane, Biokovo, Podgora
XH79
Jama pod Osojem (cave), Vrata Biokova, Biokovo, Podgora
Jama za Supinom (cave), Vrata Biokova, Biokovo, Podgora
Tu~epska vilenja~a (cave), Ravna vla{ka, Biokovo, Tu~epi
XH79
XH79
XH79
Ze~ica (cave), Ze~ica, [o{i}i, Biokovo, Podgora
XH79
Velika {pilja (cave), Antunovi}i, Biokovo, Kozica
XH89
Pozjata (cave), Jagodnik, Bartulovi}i, Biokovo, Brela
XJ50
Region Source
CNHM collection 31
CBSS collection 248
SL
BUTUROVI], 1955c
KARAMAN, 1966
SL
CBSS collection 249
SL
CBSS collection 250
SL
CBSS collection 251
CBSS collection 252
SL
OZIMEC et al., 2009
CBSS collection 253
MZUF collection 4
SL
BUTUROVI], 1955c
KARAMAN, 1966
SL
BUTUROVI], 1955c
Fig. 10. Croatian distribution of Alpioniscus bosniensis.
268
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
Mravinju{a jama (cave), Lozovci, @upa, Biokovo, Zagvozd
XJ70
SL
Biokovo
SL
Biokovo and Vrgorac surroundings
From the Cetina to the Neretva region
Southern Croatia
Croatia
SL
SL
SL
CNHM collection 32
CBSS collection 254
MZUF collection 5
OZIMEC, 2008
BEDEK, 2011
SKET, 1986
STROUHAL, 1940a
SCHMALFUSS, 2003
POTO^NIK, 1989
14. Alpioniscus christiani Poto~nik, 1983 (Tab. 14, Fig. 11)
Distribution:
Type locality:
Croatia: the island of Krk (SCHMALFUSS, 2003)
Biserujka (cave), Rudine, ^i`i}i, Omi{alj, the island of Krk, Kvarner, Croatia
[Höhle »Vitezi~eva pe~ina« bei Rudine, Omi{alj, Insel Krk (POTO^NIK, 1983)]
Ecological status: Troglobiotic
Fig. 11. Distribution of Alpioniscus christiani.
Tab. 14. List of recorded localities for Alpioniscus christiani
Locality
UTM
Biserujka (cave), Rudine, ^i`i}i, Omi{alj, Krk
VL60
Region Source
CBSS collection 255
POTO^NIK, 1983
GOTTSTEIN et al., 2001
NL
GOTTSTEIN MATO^EC et
al., 2002
BEDEK et al., 2006
BO@I] et al., 2010
269
Nat. Croat. Vol. 20(2), 2011
Krk
NL
Croatia
TABACARU, 1996
SCHMALFUSS, 2003
POTO^NIK, 1989
TVRTKOVI] et al., 2004
15. Alpioniscus haasi (Verhoeff, 1931) (Tab. 15, Fig. 12)
Synonyms:
Distribution:
Type locality:
Illyrionethes haasi Verhoeff, 1931
Croatia: the island of Kor~ula (SCHMALFUSS, 2003)
Pi{urka (cave), Kor~ula, the island of Kor~ula, Dalmatia, Croatia [Paganetti-Höhle auf der dalmatinischen Insel Curzola (VERHOEFF, 1931a, b)]
Ecological status: Troglobiotic (STROUHAL, 1940a; BUTUROVI], 1957)
Tab. 15. List of recorded localities for Alpioniscus haasi
Locality
Jama na Kamenoj (cave), Kamena, ^ara, Kor~ula
Jama na Mehovnju (cave), Veliki hom, Postrane, @rnovo,
Kor~ula
Jama u Dubokom dolu (cave), Duboki dol, Pupnat, Kor~ula
[pilja pod Veli vrh (cave), Veli vrh, Pupnat, Kor~ula
UTM
XH55
Region Source
SL
CBSS collection 256
XH65
SL
CBSS collection 257
XH65
XH65
SL
SL
Samograd (cave), Ra~i{}e, Kor~ula
XH66
SL
Jakasova {pilja (cave), Glava, Rasohatica, @rnovo, Kor~ula
XH75
SL
Jama pod Bilu{inim vrtom (cave), Postrane, @rnovo, Kor~ula
Jama pod Kola~ (cave), Vu~ja, Postrane, @rnovo, Kor~ula
Jama u Mikulin dolac (cave), Mikulina glava, Postrane,
@rnovo, Kor~ula
Jama u Vrisju (cave), @rnovo, Kor~ula
XH75
XH75
SL
SL
CBSS collection 258
CBSS collection 259
CBSS collection 260
OZIMEC, 2004
CBSS collection 261
FRANKENBERGER, 1938a
KARAMAN, 1966
CBSS collection 262
CBSS collection 263
XH75
SL
CBSS collection 264
XH75
SL
Pi{urka (cave), Kor~ula, Kor~ula
XH75
SL
CBSS collection 265
CNHM collection 33
CBSS collection 266
VERHOEFF, 1931a
VERHOEFF, 1931b
FRANKENBERGER, 1938a
ARCANGELI, 1947
KARAMAN, 1966
OZIMEC, 2004
BEDEK et al., 2006
OZIMEC, 2010
STROUHAL, 1939a
FRANKENBERGER &
STROUHAL, 1940
SCHMÖLZER, 1965a
TABACARU, 1996
SCHMALFUSS, 2003
VANDEL, 1946
STROUHAL, 1940a
POTO^NIK, 1989
Kor~ula
SL
Dalmatia
From The Neretva to the S Montenegro region
Croatia
SL
270
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
Fig. 12. Distribution of Alpioniscus haasi.
16. Alpioniscus heroldi (Verhoeff, 1931) (Tab. 16, Fig. 13)
Synonyms:
Distribution:
Type locality:
Illyrionethes heroldi Verhoeff, 1931
Southern Croatia, Bosnia and Herzegovina (SCHMALFUSS, 2003)
Bilek (cave), south Herzegovina, Bosnia and Herzegovina [Bilek-Höhle der
Südherzegowina (VERHOEFF, 1931b)]
Ecological status: Troglobiotic (STROUHAL, 1940a; BUTUROVI], 1957)
Tab. 16. List of recorded localities for Alpioniscus heroldi
Locality
UTM
Region Source
Jama u Zabira|u (cave), Osojnik, Dubrovnik
BN53
SL
CBSS collection 267
Tunel u Pera~kom blatu (tunnel, cave), Pera~ko blato, Plo~e
XH97 SL
CBSS collection 268
[pilja na Zalokavlju (cave), Raba, Opuzen
YH06 SL
CNHM collection 34
Vi{ti}ina jama (cave), Dukati, Opuzen
YH06 SL
CNHM collection 35
Kornjatu{a (cave), Dobranje, Opuzen
YH16 SL
CNHM collection 36
Pavli~evica (cave), Dragovija – Vid, Metkovi}
YH17 SL
Vilenska pe} (cave), Ogra|enica, Gumanca, Majkovi, Slano
YH34 SL
CBSS collection 270
Kukova pe} (cave), Brse~ine, Trsteno, Dubrovnik
YH43 SL
CBSS collection 271
Zadubravica (cave), Zadubravica, Ri|ica, Dubrovnik
YH44 SL
CBSS collection 272
Dubrovnik region
SL
CNHM collection 37
CBSS collection 269
SKET, 1986
SKET, 2003
271
Nat. Croat. Vol. 20(2), 2011
Southern Croatia
SL
Croatia
SCHMALFUSS, 2003
POTO^NIK, 1989
MISINTERPRETED LOCALITY
Benetina pe}ina (cave), Gumanca Slano*, Bosnia and
Herzegovina
STROUHAL, 1939a
* The locality is probably Benetina pe}ina in Popovo polje, Herzegovina, near Slano.
Fig. 13. Croatian distribution of Alpioniscus heroldi.
Alpioniscus herzegowinensis (Verhoeff, 1931) (Tab. 17)
Synonyms:
Distribution:
Type locality:
Illyrionethes herzegowinensis Verhoeff, 1931
Bosnia and Herzegovina (SCHMALFUSS, 2003)
Pe}ine u [umi (caves), [uma, Trebinje, Bosnia and Herzegovina [Gebiet der
Schuma der südwestlichsten Herzegowina in verschiedenen Höhlen im
Dunkeln (VERHOEFF, 1931b)]
Ecological status: Troglobiotic (STROUHAL, 1940a; BUTUROVI], 1957)
Tab. 17. List of recorded localities for Alpioniscus herzegowinensis
Locality
UTM
Region Source
Lukina jama – Trojama (cave system), Hajdu~ki kukovi,
Velebit, Krasno*
VK95
NL
KEROVEC et al., 19991
*Most probably a misidentification. All examined material from the cave system Lukina jama – Trojama and
other caves on Northern Velebit Mt. belongs to other species, not yet described, related to A. strasseri.
1
Alpioniscus hercegowiniensis
272
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
17. Alpioniscus kratochvili (Frankenberger, 1938) (Tab. 18, Fig. 14)
Synonyms:
Distribution:
Type locality:
Illyrionethes kratochvili Frankenberger, 1938
Croatia: the island of Hvar (SCHMALFUSS, 2003)
Markova {pilja (cave), Pelegrin, Hvar; Gr~ka pe}ina (cave), Hvar and Grap~eva {pilja (cave), Gramin Dolac, Humac, Jelsa, the island of Hvar, Dalmatia,
Croatia [Markova spilja, près de Pellegrino, environs de la ville de Hvar. –
Gr~ka pe}ina, environs de la ville de Hvar. – Grab}ina pe}ina, environs de
la ville de Hvar. (FRANKENBERGER, 1938a)]
Ecological status: Troglobiotic (STROUHAL, 1940a; BUTUROVI], 1957)
Fig. 14. Distribution of Alpioniscus kratochvili.
Tab. 18. List of recorded localities for Alpioniscus kratochvili
Locality
UTM
Gr~ka pe}ina (cave), Hvar, Hvar
XH18
Markova {pilja (cave), Pelegrin, Hvar, Hvar
XH18
[pilja pod Kapelu (cave), Smokovnik, Hvar, Hvar
XH18
Grap~eva {pilja (cave), Gromin Dolac, Humac, Jelsa, Hvar
XH47
Jama na Teruncu (cave), Bogomolje, Hvar
XH67
Propod (cave), Su}uraj, Hvar
XH77
Region Source
FRANKENBERGER, 1938a
SL
KARAMAN, 1966
CBSS collection 273
FRANKENBERGER, 1938a
SL
KARAMAN, 1966
BEDEK et al., 2006
SL
UBIC & OZIMEC, 2005
FRANKENBERGER, 1938a
SL
KARAMAN, 1966
SL
CNHM collection 38
GIROMETTA, 19141
SL
LANGHOFFER, 1915a1
STROUHAL, 1939b
273
Nat. Croat. Vol. 20(2), 2011
Hvar
SL
Dalmatia
From the Cetina to the Neretva region
Croatia
SL
SL
1
STROUHAL, 1939a
FRANKENBERGER &
STROUHAL, 1940
SCHMÖLZER, 1965a
TABACARU, 1996
SCHMALFUSS, 2003
VANDEL, 1946
STROUHAL, 1940a
POTO^NIK, 1989
Titanethes albus
18. Alpioniscus magnus (Frankenberger, 1938) (Tab. 19, Fig. 15)
Synonyms:
Distribution:
Type locality:
Illyrionethes magnus Frankenberger, 1938
Croatia (SCHMALFUSS, 2003)
Bazgova~a jama (cave), Nere`i{}a; Kopa~ina (cave), Donji Hum, Supetar; Tanki ratac (cave), Supetar; ^injadra (cave), Nere`i{}a; Filipovi}eva jama (cave),
Nere`i{}a; Jama za Mahrincem (cave), Pu~i{}a and [pilja pod Vrhom (cave),
Pu~i{}a, the island of Bra~, Dalmatia, Croatia [Razgova}a spilja, environs du
village Nezeri{}e. – Kopacina près du village de Dolni Humac, environs de la
ville de Supetar. – Tanki ratac, près de la ville de Supetar. ^injadra spilja, environs du village de Nezeri{}e. – Filipovi}eva jama, environs de Nezeri{}e. –
Jama za Mahrinacom, environs du village de Pu~i{}e. – Spilja pod Vrhom, environs de Pu~i{}e. (FRANKENBERGER, 1938a)]
Ecological status: Troglobiotic (STROUHAL, 1940a; BUTUROVI], 1957)
Fig. 15. Distribution of Alpioniscus magnus.
274
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
Tab. 19. List of recorded localities for Alpioniscus magnus
Locality
UTM
[pilja izme|u Lokvica i Blisine glavice (cave), Milna, Bra~
XH19 SL
[pilja izme|u Milne i Sv. Josipa, Milna (cave), Bra~
Dobra jama (cave), Nere`i{}a, Bra~
Region Source
XH19 SL
XH29 SL
WETTSTEIN, 19141
STROUHAL, 1939b
WETTSTEIN, 19141
STROUHAL, 1939b
STROUHAL, 1939a
KARAMAN, 1966
FRANKENBERGER, 1938a
Filipovi}eva jama (cave), Nere`i{}a, Bra~
XH29 SL
STROUHAL, 1939a
KARAMAN, 1966
CNHM collection 39
CBSS collection 274
Bazgova~a jama (cave), Nere`i{}a, Bra~
XH39 SL
FRANKENBERGER, 1938a
STROUHAL, 1939a
KARAMAN, 1966
BEDEK et al., 2006
Jama kod Vidove gore (cave), Vidova gora, Bol, Bra~
Je{kalovica (cave), Selca, Bra~
XH39 SL
XH49 SL
WETTSTEIN, 19141
STROUHAL, 1939b
STROUHAL, 1939a
KARAMAN, 1966
STROUHAL, 1939a
Studentski ratac (cave), Selca, Bra~
XH49 SL
STROUHAL, 1939e
KARAMAN, 1966
@ejava (cave), Osridke, Selca, Bra~
XH49 SL
STROUHAL, 1939a
KARAMAN, 1966
FRANKENBERGER, 1938a
Kopa~ina (cave), Donji Hum, Supetar, Bra~
XJ20
SL
STROUHAL, 1939a
KARAMAN, 1966
FRANKENBERGER, 1938a
Tanki ratac (cave), Supetar, Bra~
XJ20
SL
STROUHAL, 1939a
KARAMAN, 1966
CBSS collection 275
WETTSTEIN, 19141
FRANKENBERGER, 1938a
^injadra (cave), Nere`i{}a, Bra~
XJ30
SL
STROUHAL, 1939a
STROUHAL, 1939b
KARAMAN, 1966
HLAVÁ^ & JAL@I], 2009
Minjera (mine, cave), [krip, Postira, Bra~
XJ30
SL
CBSS collection 276
CBSS collection 277
Jama za Mahrincem (cave), Pu~i{}a, Bra~
XJ40
SL
FRANKENBERGER, 1938a
STROUHAL, 1939a
KARAMAN, 1966
FRANKENBERGER, 1938a
[pilja pod Vrhom (cave), Pu~i{}a, Bra~
XJ40
SL
STROUHAL, 1939a
VANDEL, 1946
KARAMAN, 1966
275
Nat. Croat. Vol. 20(2), 2011
Bra~
SL
FRANKENBERGER &
STROUHAL, 1940
SCHMÖLZER, 1965a
TABACARU, 1996
From SE Italy to the Cetina region
STROUHAL, 1940a
POTO^NIK, 1989
Croatia
1
SCHMALFUSS, 2003
Titanethes albus
19. Alpioniscus strasseri (Verhoeff, 1927) (Tab. 20, Fig. 16)
Synonyms:
Distribution:
Type locality:
Illyrionethes strasseri Verhoeff, 1927; I. enriquesi Arcangeli, 1932
NE-Italy, Slovenia, Croatia including the island of Cres (SCHMALFUSS, 2003)
Grotta ad ovest di Basovizza (cave, Friuli Venezia Giulia cadastre no. 68, historical cadastre no. 53/VG), Longera, Trieste, Italy and Belinca jama (cave,
Slovenian cave cadastre no. 950), [tarje, Se`ana, Slovenia [Höhle am Mt.
Spaccato in Istrien, eine Stunde von Triest entfernt; Höhle Belinca bei Storje
(VERHOEFF, 1927)]
Ecological status: Troglobiotic (STROUHAL, 1940a; BUTUROVI], 1957)
Tab. 20. List of recorded localities for Alpioniscus strasseri
Locality
UTM
Region
Porto Madonna (cave), Savudrija
UL83
NL
Markova jama (cave), [tancija [pin, Tar, Pore~
UL91
NL
Marmo (cave), Brtonigla
UL92
NL
Source
VERHOEFF, 1929b
WOLF, 1938
CBSS collection 278
VERHOEFF, 1929a
WOLF, 1938
Manzin (cave), Vodnjan
VK07
NL
STROUHAL, 1939a
St. Lucia (cave), Vodnjan
VK07
NL
STROUHAL, 1939a
Jama kod Buri}i (cave), Buri}i, Kanfanar
VK09
NL
CBSS collection 279
OZIMEC et al., 2011b
ARCANGELI, 1932
BRIAN, 1938
WOLF, 1938
[pilja na Gradini kod Premanture (cave), Premantura, Pula
VK16
NL
STROUHAL, 1939a
FRANKENBERGER &
STROUHAL, 1940
KARAMAN, 1966
Ikinica (cave), Munti}
VK17
NL
CNHM collection 40
Fe{tinsko kraljevstvo (cave), Fe{tini, @minj
VK19
NL
CBSS collection 280
^ampari (cave), Petri}evi, Beli, Cres
VK49
NL
Ov~arica (cave), Belej, Cres
VK55
NL
CBSS collection 281
Piskovica (cave), Jakli}i, Gologorica, Cerovlje
VL21
NL
CBSS collection 282
Rabakova {pilja (cave), Rabaki, Ro~, Ro~ko polje, Buzet
VL22
NL
Denizza (cave), Kr{an, Plomin
VL30
NL
VERHOEFF, 1938
KARAMAN, 1966
CBSS collection 283
OZIMEC et al., 2011b
BRIAN, 1938
STROUHAL, 1939a
276
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
Jama iznad Vranjske drage (cave), Pikuli}i, Boljun, Vela
draga, U~ka
VL31
NL
Jama mrtvih golubova (cave), Mala U~ka, U~ka
VL31
NL
Jama na Balagu{tini (cave), Dobre~, Poklon, U~ka
VL31
NL
CBSS collection 284
CNHM collection 41
CBSS collection 285
CBSS collection 286
Mala pe} (cave), Molinarska draga, U~ka
VL31
NL
CBSS collection 287
Petr~eva jama (cave), Petrc, Podmaj, Mala U~ka, U~ka
VL31
NL
CBSS collection 288
Pupi}ina pe}ina (cave), Vranja, Boljun, Vela draga, U~ka
VL31
NL
CBSS collection 289
Rupa I (cave), Podmaj, Mala U~ka, U~ka
VL31
NL
CBSS collection 290
Rupa II (cave), Podmaj, Mala U~ka, U~ka
VL31
NL
CBSS collection 291
Topolska pe} (cave), Topol, Poklon, U~ka
VL31
NL
CBSS collection 292
Barinova jama (cave), Vojni poligon iznad Bresta, Brest pod
U~kom, ]i}arija
VL32
NL
CBSS collection 293
Boru{njak 1 (cave), Bon~i}a ku}a, Boru{njak, Brgudac,
]i}arija, Lupoglav
VL32
NL
CBSS collection 294
Boru{njak 3 (cave), Bon~i}a ku}a, Boru{njak, Brgudac,
]i}arija, Lupoglav
VL32
NL
CBSS collection 295
Crekvica (cave), Vojni poligon iznad Bresta, Brest pod
U~kom, ]i}arija
VL32
NL
CBSS collection 296
Jama na prevoji}u (cave), Popova voda, Brgudac, ]i}arija,
Lupoglav
VL32
NL
CBSS collection 297
Jama pod kri` (cave), Podbrus, Semi}i, ]i}arija, Lupoglav
VL32
NL
CBSS collection 298
Novogodi{nja jama (cave), Ba~venik, Dolenja Vas, Lupoglav,
]i}arija
VL32
NL
CBSS collection 299
O{tirova jama (cave), Vojni poligon iznad Bresta, Brest pod
U~kom, ]i}arija
VL32
NL
CBSS collection 300
Pe}ina kod planinarske ku}e Pavlovac (cave), Bon~i}a ku}a,
Boru{njak, Brgudac, ]i}arija, Lupoglav
VL32
NL
CBSS collection 301
Pe}ina na Crekvici 1 (cave), Vojni poligon iznad Bresta, Brest
pod U~kom, ]i}arija
VL32
NL
CBSS collection 302
Pe}ina pod Brlo`nikom (cave), Brlo`nik, Veprinac, ]i}arija,
Opatija
VL32
NL
CBSS collection 303
Pe}ina pod Sokoli}em (cave), Sokoli}, Semi}i, ]i}arija,
Lupoglav
VL32
NL
CBSS collection 304
Pe}ina pod Str`en (cave), Str`en, Brgudac, ]i}arija, Lupoglav VL32
NL
CBSS collection 305
Pe}ina u Topolovcu (cave), Topolovac, Brgudac, ]i}arija,
Lupoglav
NL
CBSS collection 306
VL32
SD 2 (cave), Mahenac, Veprinac, ]i}arija, Opatija
VL32
NL
CBSS collection 307
Vela pe} (cave), Brest pod U~kom, ]i}arija
VL32
NL
CBSS collection 308
Dru{ka pe} (cave), Mo{}eni~ka draga, Potoki, Lovran
VL41
NL
CBSS collection 309
Jama kod potoka Banine 3 (cave), Menderi, Veprinac, U~ka
VL41
NL
CBSS collection 310
Proljetna jama (cave), Veliki Knezgrad, Lovranska Draga,
U~ka, Lovran
VL41
NL
CBSS collection 311
Rudnik u Medveji (mine), Medveja, Lovran
VL41
NL
CBSS collection 312
Jama na Mahencu 2 (cave), Mahenac, Veprinac, Opatija
VL42
NL
CBSS collection 313
CNHM collection 42
MZUF collection 6
FRANKENBERGER, 1939
Pe}ina kod Pu`i (cave), Pu`i, Opatija
VL42
NL
FRANKENBERGER &
STROUHAL, 1940
KARAMAN, 1966
277
Nat. Croat. Vol. 20(2), 2011
Pe}ina kod sela Puhari (cave), Puhari, Veprinac, Opatija
[pilja iznad ceste (cave), brodogradili{te V. Lenac, Kostrena,
Rijeka
VL42
VL51
NL
NL
CBSS collection 314
CBSS collection 315
LUKI] et al., 20111
CBSS collection 316
Bunetova jama (cave), Hum~ine, Ro`mani}i, Bakar
VL61
NL
LUKI] et al., 2009
LUKI] et al., 2011
Karlova jama (cave), Sopalj, Kostrena, Rijeka
VL61
NL
Mali {tumbur (cave), Perilo, Kostrena, Rijeka
VL61
NL
CBSS collection 317
LUKI] et al., 2011
CBSS collection 318
LUKI] et al., 2011
CNHM collection 43
[pilja u uvali Predpe} (cave), uvala Predpe}, Paveki, Bakar
VL61
NL
CBSS collection 319
LUKI] et al., 2009
LUKI] et al., 2011
CBSS collection 320
[pilja u uvali Sve`anj (cave), uvala Sve`anj, Paveki, Bakar
VL61
NL
LUKI] et al., 2009
LUKI] et al., 2011
CBSS collection 321
Ton~akova jama (cave), Hum~ine, Ro`mani}i, Bakar
VL61
NL
Urinjska {pilja (cave), Urinj, Kostrena, Rijeka
VL61
NL
Vrtare male (cave), Dramalj, Crikvenica
VL70
NL
CBSS collection 323
Lokvarka (cave), Lokve, Delnice*
VL82
M
KARAMAN, 1966
LUKI] et al., 2011
CBSS collection 322
LUKI] et al., 2011
STROUHAL, 1939a
CASALE et al., 2004
[pilja na U~ki (cave), U~ka
VL
NL
Cres
NL
Istria
NL
From SE Italy to the Cetina region
KESSELYÁK, 1931
VERHOEFF, 1940
ARGANO et al., 1982
VERHOEFF, 1931c
ARGANO et al., 1982
STROUHAL, 1940a
SCHMÖLZER, 1965a
PAOLETTI, 1978
Croatia
POTO^NIK, 1989
TABACARU, 1996
SCHMALFUSS, 2003
MISIDENTIFICATION
Medvje|a {pilja (cave), Rt. Lokvica, Sv. Jakov, Lo{inj**
VK53
NL
JAL@I], 2007
*According to STROUHAL (1939a) only two female specimens were available for the identification of the
species. All examined material from the area of Gorski kotar belongs to other species, not yet described,
therefore STROUHAL's record might be a misidentification.
**Re-examination of this material shows that the specimens belong to another, not yet described species.
1
Alpioniscus sp.
278
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
Fig. 16. Croatian distribution of Alpioniscus strasseri.
20. Alpioniscus trogirensis Buturovi}, 1955 (Tab. 21, Fig. 17)
Distribution:
Type locality:
Croatia: Dalmatia, Trogira (probably Trogir) (SCHMALFUSS, 2003)
[pilja Sv. Filipa i Jakoba (cave), Marina and [pilja od Bilosoja (cave), St Anthony's Monastery, Trogir, Dalmatia, Croatia [Grotte près de la ville Trogir
(Dalmatie) (BUTUROVI], 1955a)*]
Ecological status: Troglobiotic (BUTUROVI], 1957)
Tab. 21. List of recorded localities for Alpioniscus trogirensis
Locality
[pilja Sv. Filipa i Jakoba (cave), Marina, Trogir
[pilja od Bilosoja (cave), St Anthony’s Monastery, Trogir
Trogir surroundings
Croatia
UTM Region Source
BUTUROVI], 1955a*
WJ82 SL
KARAMAN, 1966
BEDEK et al., 2006
CBSS collection 324
MZUF collection 7
XJ01 SL
BUTUROVI], 1955a*
KARAMAN, 1966
SL
SCHMALFUSS, 2003
POTO^NIK, 1989
TABACARU, 1996
* According to HAD@I (1933) the material collected in 1924 by Stanko Karaman, labeled as »pe}ina u neposrednoj blizini Trogira« (Grotte près de la ville Trogir, Cave near Trogir) is from one of two caves: [pilja Sv.
Filipa i Jakoba and [pilja od Bilosoja
279
Nat. Croat. Vol. 20(2), 2011
Fig. 17. Distribution of Alpioniscus trogirensis.
21. Alpioniscus verhoeffi (Strouhal, 1938) (Tab. 22, Fig. 18)
Synonyms:
Distribution:
Illyrionethes verhoeffi (Strouhal, 1938)
Croatia or Herzegovina (STROUHAL, 1938); Bosnia (SCHMALFUSS, 2003) is
probably a misinterpreted datum.
Type locality:
[pilja u dolini Neretve (cave), Dalmatia, Croatia or Bosnia and Herzegovina, [Narenta Grotten, Dalmatien (STROUHAL, 1938)]
Ecological status: Troglobiotic (STROUHAL, 1940a; BUTUROVI], 1957)
Tab. 22. List of recorded localities for Alpioniscus verhoeffi
Locality
UTM
Region Source
[pilja na Vr{inoj glavici (cave), Stra`benica, Otok, Sinj
XJ44
SL
CBSS collection 325
Jamica u docima (cave), Varda, Desne, Metkovi}
YH07
SL
CBSS collection 326
Kaverna pod vijaduktom kod Puljana (cave), Puljani, Jezero,
Plo~e
YH07
SL
CBSS collection 327
Maica golubinka (cave), Golovrh, Bebi}i, Metkovi}
YH07
SL
CBSS collection 328
Po`arova jama (cave), Desne, Opuzen
YH07
SL
CBSS collection 329
STROUHAL, 1938
[pilja u dolini Neretve (cave), Neretva*
YH
SL
FRANKENBERGER &
STROUHAL, 1940
BUTUROVI], 1955c
KARAMAN, 1966
280
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
Neretva region
SL
STROUHAL, 1939a
SCHMÖLZER, 1965a
Dalmatia
SL
VANDEL, 1946
From the Cetina to the Neretva region
SL
STROUHAL, 1940a
*It is possible that this locality is in Herzegovina
Fig. 18. Croatian distribution of Alpioniscus verhoeffi.
22. Titanethes albus (C. Koch, 1841) (Tab. 23, Fig. 19)
Synonyms:
Pherusa alba C. Koch, 1841; Titanethes absoloni Verhoeff, 1901; T. brevicornis
Joseph, 1882; T. fracticornis Joseph, 1882; T. nodifer Verhoeff, 1901
Distribution:
NE-Italy, Slovenia (SCHMALFUSS, 2003); Croatia (POTO^NIK, 1989)
Type locality:
Postojnska jama (cave system, Slovenian cave cadastre no. 747), Postojna, Slovenia [Adelsbergergrotte in Krain (KOCH, 1841)]
Ecological status: Troglobiotic (STROUHAL, 1940a; BUTUROVI], 1957)
Tab. 23. List of recorded localities for Titanethes albus
Locality
Crna pe}ina (cave), Korana, Plitvi~ka jezera, Korenica
Golubnja~a (cave), Korana, Plitvi~ka jezera, Korenica
Mra~na {pilja (cave), Korana, Plitvi~ka jezera, Korenica
UTM Region Source
WK47 M
REDEN[EK, 1958
CBSS collection 330
WK47 M
REDEN[EK, 1958
WK47 M
REDEN[EK, 1958
281
Nat. Croat. Vol. 20(2), 2011
LANGHOFFER, 1915a
[upljara (cave), Kalu|erovac, Plitvi~ka jezera, Korenica
WK47 M
STROUHAL, 1939b
Bara}eva {pilja (cave), Nova Kr{lja, Rakovica, Slunj*
WK58 M
STROUHAL, 1939b
REDEN[EK, 1958
CBSS collection 331
Donja Bara}eva {pilja (cave), Nova Kr{lja, Rakovica, Slunj
WK58 M
STROUHAL, 1939b
OZIMEC, 2003
HLAVÁ^ & JAL@I], 2010
CNHM collection 44
Dumen~i}a {pilja (cave), stara Kr{lja, Rakovica, Slunj
WK58 M
CBSS collection 332
LOHAJ & LAKOTA, 2010
LANGHOFFER, 1915a
Gornja Bara}eva {pilja (cave), Nova Kr{lja, Rakovica, Slunj
WK58 M
STROUHAL, 1939b
OZIMEC, 2003
Zobenica pe}ina (cave), Stara Kr{lja, Rakovica, Slunj
WK58 M
LANGHOFFER, 1915a
STROUHAL, 1939b
Zvirnjak pe}ina (cave), Rakovica, Slunj
WK58 M
STROUHAL, 1939b
[ari}a jama (cave), Risovac, Frka{i}, Plje{ivica, Korenica
WK65 M
MZUF collection 8
Kr~evina (cave), Kr~evina, Capa}u{a, Nebljusi, Donji Lapac,
WK74 M
Plje{ivica
BUTUROVI], 1963
[pilja pod Zimzelom (cave), Potok Musulinski, Ogulin**
WL01
M
HLAVÁ^ & JAL@I], 2010
Pe}inik (cave), Pu{kari}i, Ogulin
WL11
M
Jama 3 (cave), Malik, Ku{trovka, Tro{marija
WL12
M
CBSS collection 333
Jama na Bertiji 1 (cave), Bertija, Malik, Ku{trovka, Tro{marija WL12
M
CBSS collection 334
Jama na Bertiji 2 (cave), Bertija, Malik, Ku{trovka, Tro{marija WL12
M
CBSS collection 335
Ku{trovka (cave), Ku{trovka, Tro{marija
M
CBSS collection 336
WL12
CNHM collection 45
CASALE & JAL@I], 1999
CNHM collection 46
JURINAC, 1886
JURINAC, 1888
Ambarac (sinkhole), O{tarije, Ogulin
WL20
M
LANGHOFFER, 1912
LANGHOFFER, 1915b
CSIKI, 1926
STROUHAL, 1939b
Mandelaja (cave), O{tarije, Ogulin
WL20
M
CBSS collection 337
Munjave (sinkhole), Josipdol, Ogulin
WL20
M
STROUHAL, 1939b
[pilja u Mekoti (cave), Ko{are, Tounj
WL20
M
Ceri~ka pe}ina (cave), Tounj
WL21
M
Guge~ka pe}ina (cave), Gornje Dubrave, Tounj
WL21
M
STILLER, 19141
STROUHAL, 1939b
STROUHAL, 1939b
STILLER, 1911b1
STROUHAL, 1939b
CBSS collection 338
Mika{inovi}a {pilja (cave), Bistrica, Mika{inovi}i, Gornje
Dubrave, Tounj
WL21
M
Su{ica (sinkhole), Rebi}a Glavica, Generalski Stol
WL21
M
CBSS collection 339
[pilja u kamenolomu Tounj (cave), Kamenolom Tounj, Tounj WL21
M
CBSS collection 340
Tamnica (cave), Potok Tounjski, Tounj
WL21
M
CBSS collection 341
Vreli} (cave), Donje Dubrave, Generalski Stol
WL21
M
CBSS collection 342
Bibi~ka {pilja (cave), kanjon Ribnjaka, Munjasi, Tro{marija
WL22
M
CBSS collection 343
JAL@I] & BO@I^EVI], 1975
282
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
Danetska pe}ina (cave), kanjon Dobre, Dani, Bosiljevo
WL22
M
CBSS collection 344
WL22
M
CBSS collection 345
WL22
WL22
WL22
M
M
M
Jama u kanjonu Dobre ispod Grabrka (cave), kanjon Dobre,
WL22
Podumol, Bosiljevo
M
Dragina pe}ina na Dobri (cave), kanjon Dobre, Grabrk,
Tro{marija, Karlovac
Jama II kod Koli}a (cave), Koli}i, Generalski Stol
Jama na Gnojini (cave), Gnojina, Mate{e, Tro{marija
Jama na rubu (cave), Bartolovi}i, Gornje Dubrave, Tounj
WL22
M
CBSS collection 346
CBSS collection 347
CBSS collection 348
MZUF collection 9
CBSS collection 349
CBSS collection 350
WL22
M
CBSS collection 351
WL22
M
CBSS collection 352
WL22
WL22
WL24
WL24
M
M
C
C
CBSS collection 353
CBSS collection 354
CBSS collection 355
CBSS collection 356
WL24
C
CBSS collection 357
WL24
WL24
WL25
WL26
WL26
WL26
C
C
C
C
C
C
Drobovnik (cave), Kun~ani, Radatovi}i, @umberak
WL26
C
Jama kod Starog mlina (cave), Boi}i, So{ice, @umberak
WL26
C
Jama na Javorniku (cave), Kuljaji, Gudalji, Radatovi}i,
@umberak
WL26
C
Jama u Lukama (cave), Sekuli}i, Radatovi}i, @umberak
Jama u Vrloj strani (cave), Gudalji Radatovi}i, @umberak
Kotari (cave), Kuljaji, Radatovi}i, @umberak
WL26
WL26
WL26
C
C
C
Lalovi}eva jama (cave), Magovci, Radatovi}i, @umberak
WL26
C
Mili}ka (cave), Kuljaji, Radatovi}i, @umberak
Pavlovica (cave), Keseri, Radatovi}i, @umberak
Pe~enjevka (cave), Keseri, Radatovi}i, @umberak***
Pe~ine (spring cave), Malinci, Radatovi}i, @umberak
Plavinka (cave), Gaj, Sekuli}i, Radatovi}i, @umberak
Raki~ka (cave), Pilatovci, Radatovi}i, @umberak
[pilja kod Jura{eve livade (cave), Sekuli}i, Radatovi}i,
@umberak
WL26
WL26
WL26
WL26
WL26
WL26
C
C
C
C
C
C
CBSS collection 358
MZUF collection 10
CBSS collection 359
CBSS collection 360
CBSS collection 361
CBSS collection 362
CBSS collection 363
MZUF collection 11
CBSS collection 364
CNHM collection 47
CBSS collection 365
CBSS collection 366
CBSS collection 367
CBSS collection 368
CNHM collection 48
CBSS collection 369
CBSS collection 370
CBSS collection 371
CBSS collection 372
CBSS collection 373
CBSS collection 374
CBSS collection 375
WL26
C
CBSS collection 376
M
CNHM collection 49
JURINAC, 1886
JURINAC, 1888
LANGHOFFER, 1912
LANGHOFFER, 1915b
CSIKI, 1926
STROUHAL, 1939b
Ledenica u [peharima (cave), [pehari, Bosiljevo
Pe}ina pod Pe}anima (cave), kanjon Dobre, Pe}ani,
Generalski Stol
[pilja kod Podumolskog mlina (cave), kanjon Dobre,
Podumol, Bosiljevo
Trijama II (cave), [pehari, Bosiljevo
Vodena jama (cave), Koli}i, Generalski Stol
\ot (cave), Kuni} Ribni~ki, Netreti}, Lipnik, Karlovac
Jama bez imena (cave), Rosopoajnik, Netreti}, Lipnik, Karlovac
Jama na [krilama (cave), Kuni} Ribni~ki, Netreti}, Lipnik,
Karlovac
Vatrena jama (cave), Kuni} Ribni~ki, Netreti}, Lipnik, Karlovac
Zvone~ka II (cave), Gri~e, Netreti}, Lipnik, Karlovac
Pivnica (cave), @akanje, Kamanje
Brija~ica (cave), Gaj, Sekuli}i, Radatovi}i, @umberak
Cepinka (cave), Boi}i, So{ice, @umberak
Dakina jama (cave), Rajakovi}i, Radatovi}i, @umberak
Tr`i}ka {pilja (cave), [imi} brdo, Tr`i} Primi{ljanski, Tounj
WL30
283
Nat. Croat. Vol. 20(2), 2011
Ledeni~ka {pilja (cave), Rebrovi}i, Tounj
WL31
M
STROUHAL, 1939b
Lipa na Protulipi (cave), Stare{ini}i, Protulipa, Bosiljevo
WL32
C
CBSS collection 377
Izvor {pilja kod Trupkovi} mlina (cave), Novigrad na
Dobri, Karlovac
WL33
C
CBSS collection 378
Jastrebina (cave), Frketi} Selo, Novigrad na Dobri, Karlovac WL33
C
CBSS collection 379
Jama pod Gradom (cave), Dugi vrh, Cerovica, Kra{i},
@umberak
WL35
C
CBSS collection 380
Potok (cave), Furjani}i, Ozalj, @umberak
WL35
C
CBSS collection 381
Rogovac (cave), Lovi} Prekri{ki, Ozalj, @umberak
WL35
C
CBSS collection 382
[pilja pokraj Kozja~e (cave), kanjon Kupe, Kamanje, Ozalj,
Karlovac
WL35
C
CBSS collection 383
CNHM collection 50
CBSS collection 384
LANGHOFFER, 1912
Vrlovka (cave), kanjon Kupe, Kamanje, Ozalj, Karlovac
WL35
C
LANGHOFFER, 1915a
LANGHOFFER, 1915b
CSIKI, 1926
STROUHAL, 1939b2
BEDEK et al., 2009
Zverina (cave), Dugi vrh, Cerovica, Kra{i}, @umberak
WL35
C
CBSS collection 385
@lota (cave), Ferenci, Ozalj, @umberak
WL35
C
CBSS collection 386
Jamina (cave), Donji O{trc, Kra{i}, @umberak
WL36
C
CBSS collection 387
Provala (cave), Bu~ari, Kra{i}, @umberak
WL36
C
[pilja u Lukama (spring cave), Jezerine, Kra{i}, @umberak
WL36
C
CBSS collection 389
Vrulje (sinkhole), Jezerine, Kra{i}, @umberak
WL36
C
CBSS collection 390
Mla|enova pe}ina (cave), Bandino Selo, Slunj
WL40
C
Jopi}eva {pilja – Bent (cave system), Krnjak, Karlovac
WL41
C
Vra`i}a {pilja (cave), Barilovi}, Karlovac
WL42
C
Bribirska pe}ina (cave)
CBSS collection 388
MZUF collection 12
LANGHOFFER, 1915a
STROUHAL, 1939b
CBSS collection 391
LANGHOFFER, 1915a
STROUHAL, 1939b
STROUHAL, 1939b
Jezero (cave)
LANGHOFFER, 1915a
Kova~evka pe}ina (cave)
STROUHAL, 1939b
Velika pe}ina (cave)
STROUHAL, 1939b
Nature Park »@umberak-Samoborsko gorje«
C
BEDEK, 2005b
Tounj surroundings
M
SKET, 1986
Region between Josipdol and Plitvice
M
FRANKENBERGER &
STROUHAL, 1940
Northern Croatia
MUNI] et al., 2000
From SE Italy to the Cetina region
STROUHAL, 1940a
TABACARU, 1996
Croatia
POTO^NIK, 1989
MISIDENTIFICATION
Mala Ze~ica (cave), Labin{tica, Labin, Trogir****
WJ92
SL
GIROMETTA, 1913
Mar~ina jama (cave), Labin{tica, Labin, Trogir****
WJ92
SL
GIROMETTA, 1913
Velika Ze~ica (cave), Labin{tica, Labin, Trogir****
WJ92
SL
GIROMETTA, 1913
Jama kod Klja~ine (cave), Rado{i}, Trogir****
XJ02
SL
GIROMETTA, 1914
Jama pod Malim Kraljevcem (cave), Rado{i}, Trogir****
XJ02
SL
GIROMETTA, 1914
284
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
Jama pod ^ezinim humcem (cave), Gornji Mu}, Mose},
Sinj****
XJ13
SL
GIROMETTA, 1914
Orlova~a (cave), Bitunjska greda, Gornji Mu}, Mose}, Sinj**** XJ13
SL
GIROMETTA, 1914
Jama na Brlogu (cave), Stari gaj, Ramljane, Mose}, Sinj****
XJ14
SL
GIROMETTA, 1914
Golubinjka jama (cave), Jesenice, Polji~ka, Omi{****
XJ21
SL
Nesretnja jama (cave), Polji~ka, Omi{****
XJ
SL
Pe} na Glu{cu (cave), Sinj****
XJ
SL
Turska pe} (cave), Polji~ka, Omi{****
XJ
SL
Velika pe} (cave), Polji~ka, Omi{****
XJ
SL
GIROMETTA, 1913
LANGHOFFER, 1915a
GIROMETTA, 1913
LANGHOFFER, 1915a
GIROMETTA, 1914
GIROMETTA, 1913
LANGHOFFER, 1915a
GIROMETTA, 1913
LANGHOFFER, 1915a
*It is probably one of the following caves, Donja Bara}eva {pilja or Gornja Bara}eva {pilja
**The identification is based on one female and two juvenile specimens; considering the distribution of
Titanethes spp. this is probably a misidentification.
***This locality is on the Croatian-Slovenian border
****Misidentification (conclusion based according to the distribution of the genera Titanethes and Alpioniscus).
The material is not available for the identification of the species.
1
Titanethes
2
Titanethes dahli
Fig. 19. Croatian distribution of Titanethes albus.
285
Nat. Croat. Vol. 20(2), 2011
23. Titanethes dahli Verhoeff, 1926 (Tab. 24, Fig. 20)
Distribution:
Type locality:
NE-Italy, Slovenia, Croatia: Istria (SCHMALFUSS, 2003)
Diva{ka jama (cave, Slovenian cave cadastre no. 741), Diva~a, Slovenia [Divaca-Grotte, Krains (VERHOEFF, 1926)]
Ecological status: Troglobiotic (STROUHAL, 1940a; BUTUROVI], 1957)
Tab. 24. List of recorded localities for Titanethes dahli
Locality
UTM
Region
Source
CBSS collection 392
Velebita (cave system), Crikvena, Velebit, Krasno
VK95
NL
Jama kod Ra{pora (cave), Ra{por, Lupoglav, ]i}arija
VL23
NL
CBSS collection 393
Radota jama (cave), Rakitovec, Buzet, ]i}arija
VL23
NL
CBSS collection 394
Jama kod crvenog gloga (cave), Mala U~ka, U~ka
VL31
NL
CBSS collection 395
Jama kod dje~jeg oporavili{ta 2 (cave), Poklon, U~ka
VL31
NL
CBSS collection 396
Jama kod Male U~ke (cave), Mala U~ka, U~ka
VL31
NL
CBSS collection 397
Jama na Poklonu (cave), Poklon, U~ka
VL31
NL
CBSS collection 398
Jama za greben (cave), Stra`ica, Poklon, U~ka
VL31
NL
BAK[I] et al., 2008
CBSS collection 399
CBSS collection 400
K’U~ka (cave), Jazvina, Vela U~ka, U~ka, Opatija
VL31
NL
Pe}ina kod dje~jeg oporavili{ta (cave), Poklon, U~ka
VL31
NL
CBSS collection 401
Topolska pe} (cave), Topol, Poklon, U~ka
VL31
NL
CBSS collection 402
Babina jama (cave), Vojni poligon iznad Bresta, Brest pod
U~kom, ]i}arija
VL32
NL
CNHM collection 51
Barinova jama (cave), Vojni poligon iznad Bresta, Brest pod
U~kom, ]i}arija
VL32
NL
CBSS collection 403
Billova ponikva (cave), Vojni poligon iznad Bresta, Brest pod
VL32
U~kom, ]i}arija
NL
CBSS collection 404
Boru{njak 1 (cave), Bon~i}a ku}a, Boru{njak, Brgudac,
]i}arija, Lupoglav
VL32
NL
CBSS collection 405
Boru{njak 3 (cave), Bon~i}a ku}a, Boru{njak, Brgudac,
]i}arija, Lupoglav
VL32
NL
CBSS collection 406
Br{ljanovica (cave), Vojni poligon iznad Bresta, Brest pod
U~kom, ]i}arija
VL32
NL
CBSS collection 407
Jama ispod Tomini}evog brega (cave), Tomini}ev breg, Brest
pod U~kom, Lupoglav, ]i}arija
VL32
NL
CBSS collection 408
Jama na prevoji}u (cave), Popova voda, Brgudac, ]i}arija,
Lupoglav
VL32
NL
CBSS collection 409
Jama pod Boljunskim dolom (cave), Boljunski dol, Brgudac,
Lupoglav, ]i}arija
VL32
NL
CBSS collection 410
Jama pod kri` (cave), Podbrus, Semi}i, ]i}arija, Lupoglav
MZUF collection 13
VL32
NL
CBSS collection 411
Mali Boru{njak 6 (cave), Boru{njak, Brgudac, ]i}arija, Lupoglav VL32
NL
CBSS collection 412
Novogodi{nja jama (cave), Ba~venik, Dolenja Vas, Lupoglav,
]i}arija
VL32
NL
CBSS collection 413
O{tirova jama (cave), Vojni poligon iznad Bresta, Brest pod
U~kom, ]i}arija
VL32
NL
CBSS collection 414
Pe}ina pod Str`en (cave), Str`en, Brgudac, ]i}arija, Lupoglav
VL32
NL
CBSS collection 415
Pe}ina poli Vrti}a (cave), Brest pod U~kom, ]i}arija
VL32
NL
CBSS collection 416
SDB (cave), Golubine, Brgudac, Lupoglav, ]i}arija
VL32
NL
CNHM collection 52
CBSS collection 417
286
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
CBSS collection 418
SDB 2 (cave), Brgudac, Lupoglav, ]i}arija
VL32
NL
SKH (cave), Golubine, Brgudac, Lupoglav, ]i}arija
VL32
NL
Vela pe} (cave), Brest pod U~kom, ]i}arija
VL32
NL
CBSS collection 420
Grnja~a (cave), Grnja~, Lovranska Draga, U~ka, Lovran
VL41
NL
CBSS collection 421
Jama kod potoka Banine 3 (cave), Menderi, Veprinac, U~ka
VL41
NL
CBSS collection 422
Jama na Lovranskim lazi}ima (cave), Lovranski lazi}i,
Menderi, Veprinac, U~ka
VL41
NL
CBSS collection 423
Jama pod Velikim Knezgradom (cave), Veliki Knezgrad,
Tuli{evica, Lovran, U~ka
VL41
NL
CBSS collection 424
Kaverna u tunelu U~ka (cave), Veprinac, U~ka
VL41
NL
CBSS collection 425
Proljetna jama (cave), Veliki Knezgrad, Lovranska Draga,
U~ka, Lovran
VL41
NL
CBSS collection 426
Bla`eva jama (cave), Pu`i, Opatija
VL42
NL
Jama u Velim Sapcima 3 (cave), Mahenac, Veprinac, Opatija
VL42
NL
CBSS collection 427
Jama u Vetvi (cave), Vetven vrh, Poljane, Opatija, ]i}arija
VL42
NL
CBSS collection 428
Pe}ina Ivana Stani}a i Franja Stani} (cave), Pu`i, Opatija
VL42
NL
Pe}ina kod sela Puhari (cave), Puhari, Veprinac, Opatija
VL42
NL
CBSS collection 429
ZV – 1 (cave), Zvon~ev vrh, Veprinac, Opatija, ]i}arija
VL42
NL
CBSS collection 430
Casello ferrovario (cave), Permani, Opatija
VL43
NL
Pe}ina Marije Ru`i}, Permani (cave), Opatija
VL43
NL
MZUF collection 14
CBSS collection 419
FRANKENBERGER, 1939
KARAMAN, 1966
FRANKENBERGER, 1939
KARAMAN, 1966
WOLF, 19382
STROUHAL, 1939b
FRANKENBERGER, 1939
KARAMAN, 1966
CBSS collection 431
Vrelo (cave), Fu`ine
VL71
NL
Hircova jama (cave), Kali~ak, Crni Lug, Risnjak
VL73
M
CBSS collection 432
Kapaljka 1 (cave), Gerovo, Risnjak
VL73
NL
CBSS collection 433
Nacina jama (cave), Gerovo, Risnjak
VL73
NL
CBSS collection 434
Ru{evina (cave), Japetova {egina, Gerovo, Risnjak
VL73
M
CBSS collection 435
[temajzlinka (cave), Japetova {egina, Gerovo, Risnjak
VL73
M
CBSS collection 436
Trlica 1 (cave), Gerovo, Risnjak
VL73
M
CBSS collection 437
Vele vode (sinkhole), Crni Lug
VL73
M
CBSS collection 438
Bukovac (cave), Vrata, Lokve, Delnice
VL82
M
STROUHAL, 1939b
Ledena {pilja (cave), Golubinjak, Lokve, Delnice
VL82
M
OZIMEC et al., 2011a
STILLER, 1911a1
LANGHOFFER, 1915a2
STROUHAL, 1939b
STROUHAL, 1939a
Lokvarka (cave), Lokve, Delnice
VL82
M
Nove jame (cave), Lokve, Delnice
VL82
M
STROUHAL, 1939b
KARAMAN, 1966
CASALE et al., 2004
STROUHAL, 1939b
STILLER, 19131
Pustinja (cave), Delnice
VL82
M
LANGHOFFER, 1915a2
Jama iznad Hajdove hi`e (cave), Ku`elj, Delnice
VL83
M
CBSS collection 439
STROUHAL, 1939b
287
Nat. Croat. Vol. 20(2), 2011
Markov ponor (sinkhole), Lipovo Polje, Donji Kosinj, Oto~ac WK15 M
Sini~i}a {pilja (cave), Kalanj, Brinje
WK18 M
Crna~ka {pilja (cave), Crna~ko polje, Obajdini, Jezerane,
Velika Kapela
WK19 M
Rokina bezdana (cave), Crna~ko polje, Jezerane, Velika Kapela WK19 M
Veliki (cave), Kosinj, Oto~ac
WK25 M
Samograd (cave), Peru{i}, Gospi}
WK34 M
Sitvukova pe}ina (cave), Peru{i}, Gospi}
WK34 M
Lu{ka {pilja (cave), Debeli lug, Bjelsko, Ogulin
WL00
M
Su{ik (sinkhole), Dre`ni~ko polje, Dre`nica, Velika Kapela
WL00
M
Vidovi}a {pilja (cave), Dre`ni~ko polje, Dre`nica, Velika
Kapela, Ogulin
WL00
M
[pilja pod Ma~kovom dragom (cave), Ma~kova draga,
Jasenak, Ogulin
WL01
M
Li{trovica (cave), Brezno, Bjelsko, Velika Kapela, Ogulin
WL10
M
Zagorska pe} (cave), Desmerice, Zagorje, Ogulin
WL10
M
\ulin ponor – Medvedica (cave system), Ogulin
WL11
M
Ambarac (sinkhole), O{tarije, Ogulin
Mandelaja (cave), O{tarije, Ogulin
WL20
WL20
M
M
Gojak (spring cave), Gojak, Ogulin
WL21
M
Hajdu~ka pe}ina (cave), Bistrac, Gornje Dubrave, Ogulin
Per~evi}a {pilja (cave), Kanjon Tounj~ice, Tounj
Tounj~ica (cave), Kanjon Tounj~ice, Tounj
Tr`i}ka {pilja (cave), [imi} brdo, Tr`i} Primi{ljanski, Tounj
Pe}ina Bivio Beru}a (cave), Gro{ovi}i, Opatija
Likar ponor (sinkhole), Pazari{te
WL21
WL21
WL21
WL30
VL
WK
M
M
M
M
NL
M
CNHM collection 53
CBSS collection 440
MZUF collection 15
JAL@I], 2001
CNHM collection 54
CBSS collection 441
MZUF collection 16
CBSS collection 442
STROUHAL, 1939b
STROUHAL, 1939b
CNHM collection 55
LANGHOFFER, 19122
LANGHOFFER, 1915b2
CSIKI, 19262
STROUHAL, 1939b
CNHM collection 56
CBSS collection 443
LANGHOFFER, 19122
LANGHOFFER, 1915b2
CSIKI, 19262
STROUHAL, 1939b2
CBSS collection 444
CNHM collection 57
CBSS collection 445
CNHM collection 58
CBSS collection 446
MZUF collection 17
CBSS collection 447
CNHM collection 59
CBSS collection 448
JURINAC, 18862
JURINAC, 18882
LANGHOFFER, 19122
CSIKI, 19262
STROUHAL, 1939b2
OZIMEC, 20092
CBSS collection 449
MZUF collection 18
STROUHAL, 1939b
CNHM collection 60
CBSS collection 450
CNHM collection 61
CBSS collection 451
CBSS collection 452
CNHM collection 62
CBSS collection 453
CNHM collection 63
FRANKENBERGER, 1939
ERHARD, 1997
288
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
Langhofferova {pilja (cave), Ogulin
Zerngol {pilja (cave), Ogulin
[pilja kod Starigrada (cave)
WL
WL
M
M
Opatija surroundings
NL
Istria
NL
Gorski Kotar
Region from Lokve and Ogulin till Gospi} and Peru{i}
M
From SE Italy to the Cetina region
Croatia
STROUHAL, 1939b
STROUHAL, 1939b
STROUHAL, 1939b
FRANKENBERGER &
STROUHAL, 1940
HIRC, 19022
VERHOEFF, 1931c
SCHMALFUSS, 2003
SKET, 1986
FRANKENBERGER et
STROUHAL, 1940
STROUHAL, 1940a
TABACARU, 1996
POTO^NIK, 1989
MISIDENTIFICATION
Medina {pilja, Peru{i}, Gospi}*
WK34 M
LANGHOFFER, 19122
LANGHOFFER, 1915b2
CSIKI, 19262
STROUHAL, 1939b
*Examination of the CNHM collection (Crustacea, leg. A. Langhoffer, 14.08.1902.) shows that the female
specimen probably belongs to the genus Alpioniscus.
1
Titanethes
2
Titanethes albus
Fig. 20. Croatian distribution of Titanethes dahli.
289
Nat. Croat. Vol. 20(2), 2011
24. Cyphonethes herzegowinensis (Verhoeff, 1900) (Tab. 25, Fig. 21)
Synonyms:
Distribution:
Type locality:
Titanethes herzegowinensis Verhoeff, 1900
SW-Herzegovina (SCHMALFUSS, 2003); Croatia (POTO^NIK, 1989)
Ilijina pe}ina (cave); Absturz-Höhle (cave); Vu~ja pe}ina (cave) and Matuli}eva pe}ina (cave), Trebinje, Bosnia and Herzegovina [Bei Trebinje... Elias-Höhle, Absturz-, Wolfs- und Matulic-Höhle (VERHOEFF, 1900)]
Ecological status: Troglobiotic (STROUHAL, 1940a; BUTUROVI], 1957)
Fig. 21. Croatian distribution of Cyphonethes herzegowinensis.
Tab. 25. List of recorded localities for Cyphonethes herzegowinensis
Locality
UTM
Region Source
Aragonka (cave), Ljuba~, Dubrovnik
BN53
SL
Banova ljut (cave), Ljuba~, Dubrovnik
BN53
SL
CBSS collection 455
Debela ljut (cave), Ljuba~, Dubrovnik
BN53
SL
CBSS collection 456
Jama na vrh Vrguda (cave), Vrgud, Osojnik, Dubrovnik
BN53
SL
CBSS collection 457
Jama u Zabira|u (cave), Zabira|e, Osojnik, Dubrovnik
BN53
SL
CBSS collection 454
CBSS collection 458
CBSS collection 459
Mo~iljska {pilja (cave), Somino brdo, Osojnik, Dubrovnik
BN53
SL
STROUHAL, 1939b
Vilina {pilja – Ombla (cave system), Komolac, Dubrovnik
BN62
SL
CNHM collection 64
OZIMEC, 2005
290
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
Jezero (cave), Kuna Konavoska, Snije`nica, Cavtat
BN81
SL
FRANKENBERGER, 1939
STROUHAL, 1939b
[krabuljica (cave), Mali Orah, Kuna Konavoska, Snije`nica, Cavtat BN81
SL
FRANKENBERGER, 1939
Vilenska jama (cave), Ston, Pelje{ac
YH24
SL
STROUHAL, 1939b
Kukova pe} (cave), Brse~ine, Trsteno, Dubrovnik
YH43
SL
CBSS collection 460
Vitkova~a (cave), Mravinjac, Trsteno, Dubrovnik
YH43
SL
CBSS collection 461
Zadubravica (cave), Zadubravica, Ri|ica, Dubrovnik
YH44
SL
CBSS collection 462
Pelje{ac region
SL
FRANKENBERGER &
STROUHAL, 1940
Dubrovnik region
SL
KARAMAN, 1966
SKET, 2003
Wesern Pelje{ac
SL
SKET, 1986
Dalmatia
SL
KARAMAN, 1966
Southern Croatia
SL
TABACARU, 1996
From the Neretva to the S Montenegro region
SL
STROUHAL, 1940a
Croatia
POTO^NIK, 1989
25. Trichoniscus matulici matulici Verhoeff, 1901 (Tab. 26, Fig. 22)
Synonyms:
Trichoniscus omblae Verhoeff, 1901; T. remyi Verhoeff, 1933; T. sorrentinus
Verhoeff, 1908; T. stygivagus Verhoeff, 1901; T. turgidus Verhoeff, 1929
Distribution:
Southern Italy including Sicily, Croatia, Bosnia and Herzegovina, Serbia
and Montenegro (SCHMALFUSS, 2003); Slovenia (VILISICS & LAPANJE, 2005)
Type locality:
South Herzegovina (rock springs) [Südherzegowina an Felsenquellen (VERHOEFF, 1901)]
Ecological status: Troglophilic (STROUHAL, 1940a; BUTUROVI], 1957)
Tab. 26. List of recorded localities for Trichoniscus matulici matulici
Locality
UTM
Garden of Biological institute Lapad, Lapad, Dubrovnik
BN52 SL
Region
Source
BUTUROVI], 1955b
Rov na Lapadu, Lapad, Dubrovnik
BN52 SL
BUTUROVI], 1955b
Aragonka (cave), Ljuba~, Dubrovnik
BN53 SL
CBSS collection 463
Banova ljut (cave), Ljuba~, Dubrovnik
BN53 SL
CBSS collection 464
Debela ljut (cave), Ljuba~, Dubrovnik
BN53 SL
CBSS collection 465
Jama na vrh Prodoli (cave), Ljuba~, Dubrovnik
BN53 SL
CBSS collection 466
Jama na vrh Vrguda (cave), Vrgud, Osojnik, Dubrovnik
BN53 SL
CBSS collection 467
Jama u Zabira|u (cave), Osojnik, Dubrovnik
BN53 SL
CBSS collection 468
Jama uz stazu na Sv. Nikoli (cave), Ora{ac, Dubrovnik
BN53 SL
CBSS collection 469
Kaverna uz cestu iznad Groma~e (cave), Groma~a,
Dubrovnik
BN53 SL
CBSS collection 470
Mo~iljska {pilja (cave), Somino brdo, Osojnik, Dubrovnik
BN53 SL
CBSS collection 471
BUTUROVI], 1955b
KARAMAN, 1966
Rudina jama (cave), Groma~a, Dubrovnik
BN53 SL
CBSS collection 472
[pilja ispod Krsta (cave), Na Krst, Loznica, Zaton,
Dubrovnik
BN53 SL
CBSS collection 473
291
Nat. Croat. Vol. 20(2), 2011
[pilja na vrh Kr~evina (cave), Kr~evine, Groma~a, Ora{ac,
Dubrovnik
BN53 SL
CBSS collection 474
Karle (cave), Gru`, Dubrovnik
BN62 SL
STROUHAL, 1939e
VERHOEFF, 1901
Ombla (river bank), Dubrovnik
BN62 SL
Sumpora~a mala (cave), Komolac, Dubrovnik
BN62 SL
Vilina {pilja – Ombla (cave system), Komolac, Dubrovnik
BN62 SL
CSIKI, 1926
VANDEL, 1946
VANDEL, 1955
CBSS collection 475
CBSS collection 476
BUTUROVI], 1955b
KARAMAN, 1966
Kriva~a (cave), Osojnik, Dubrovnik
BN63 SL
[pilja u Gaju (cave), Osojnik, Dubrovnik
BN63 SL
CBSS collection 477
CBSS collection 478
\urovi}a {pilja (cave), \urovi}i, Cavtat, Konavle
BN71 SL
STROUHAL, 1939e
STROUHAL, 1939a
STROUHAL, 1939e
[ipun (cave), Cavtat, Dubrovnik
BN71 SL
BUTUROVI], 1955b
VANDEL, 1955
KARAMAN, 1966
Male ponte (cave), Grabova glava, Ropa, Mljet
XH93 SL
CBSS collection 479
[pilja kod Solina (cave), Soline, Mljet
XH93 SL
CBSS collection 480
Jara~ na ponti od Graca (cave), Pola~e, Mljet
XH94 SL
CBSS collection 481
Jama za Svetim Ilijom (cave), Babino Polje, Mljet
YH03 SL
CBSS collection 482
Jama za Svetim Spasom (cave), Babino Polje, Mljet
YH03 SL
CBSS collection 483
Movrica (cave), Dra~evac, Babino Polje, Mljet
YH03 SL
Osta{evica (cave), Dra~evac, Babino Polje, Mljet
YH03 SL
CBSS collection 485
Velika {pilja (cave), Blata, Mljet
YH03 SL
CBSS collection 486
Gali~njak (cave), Okuklje, Mljet
YH13 SL
Jama za Svetim Ivanom (cave), Babino Polje, Mljet
YH13 SL
CBSS collection 488
Kukova pe} (cave), Brse~ine, Trsteno, Dubrovnik
YH43 SL
CBSS collection 489
Pasja jama (cave), Brse~ine, Trsteno, Dubrovnik
YH43 SL
CBSS collection 490
Ze~ja jama (cave), Kli{evo, Dubrovnik
YH43 SL
CBSS collection 491
CBSS collection 484
STROUHAL, 1939e
CBSS collection 487
MICULINI], 2008
Zadubravica (cave), Zadubravica, Ri|ica, Dubrovnik
YH44 SL
CBSS collection 492
Lapad, Dubrovnik
BN52 SL
KARAMAN, 1966
Mljet
SL
FRANKENBERGER &
STROUHAL, 1940
VANDEL, 1946
KARAMAN, 1966
Southern Dalmatia
SL
From The Neretva to the S Montenegro region
SL
FRANKENBERGER &
STROUHAL, 1940
VANDEL, 1946
SKET, 2003
Croatia
STROUHAL, 1940a
POTO^NIK, 1989
SCHMALFUSS, 2003
292
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
26. Trichoniscus matulici metkovicensis Buturovi}, 1955 (Tab. 27, Fig. 22)
Distribution:
Type locality:
Croatia (POTO^NIK, 1989)
^o}ina jama (cave), Bebi}i, Metkovi}, Dalmatia, Croatia [^o~ina jama kod
Bebi}a sela (BUTUROVI], 1955b)]
Ecological status: Troglophilic
Tab. 27. List of recorded localities for Trichoniscus matulici metkovicensis
Locality
UTM
Region Source
Kuna (cave), ^edor, Biokovo, Makarska
XH69
SL
Mravinju{a jama (cave), Lozovci, @upa, Biokovo, Zagvozd
XJ70
SL
CBSS collection 493
CNHM collection 65
CBSS collection 494
BUTUROVI], 1955b
^o}ina jama (cave), Bebi}i, Metkovi}
YH07
SL
BUTUROVI], 1955c
KARAMAN, 1966
BEDEK et al., 2006
Biokovo
Croatia
1
SL
OZIMEC, 20081
BEDEK, 2011
POTO^NIK, 1989
Trichoniscus matulici
Fig. 22. Croatian distribution of Trichoniscus matulici.
293
Nat. Croat. Vol. 20(2), 2011
TRICHONISCIDAE: HAPLOPHTHALMINAE
27. Strouhaloniscellus biokovoensis Bedek et Taiti, 2009 (Tab. 28, Fig. 23)
Distribution:
Type locality:
Croatia: Mt. Biokovo (BEDEK & TAITI, 2009)
Jama iznad Sarana~a (cave), Sarana~, Gornje Igrane, Podgora, Biokovo, Dalmatia (BEDEK & TAITI, 2009)
Ecological status: Troglobiotic (BEDEK & TAITI, 2009)
Tab. 28. List of recorded localities for Strouhaloniscellus biokovoensis
Locality
UTM
Region
Jama iznad Sarana~a (cave), Sarana~, Gornje Igrane,
Podgora, Biokovo
XH79
SL
Velika {pilja (cave), Antunovi}i, Biokovo, Kozica
XH89
SL
Biokovo
SL
Source
CBSS collection 495
MZUF collection 19
BEDEK & TAITI, 2009
BEDEK, 2010
JAL@I] et al., 2010
CBSS collection 496
BEDEK, 2011
OZIMEC et al., 2011c
Fig. 23. Distribution of Strouhaloniscellus biokovoensis.
28. Troglocyphoniscus absoloni Strouhal, 1939 (Tab. 29, Fig. 24)
Distribution:
Type locality:
Croatia: the island of Mljet (SCHMALFUSS, 2003)
Movrica (cave), Dra~evac, Babino Polje, the island of Mljet, Dalmatia, Croatia [Movrica auf der Insel Mljet (Meleda); BB Nr. 661 (STROUHAL, 1939c)]
294
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
Ecological status: This species is known as troglobiotic (STROUHAL, 1940a; BUTUROVI], 1957),
but it has to be considered as troglophilic since it occurs at cave entrances
and eyes are present.
Fig. 24. Distribution of Troglocyphoniscus absoloni.
Tab. 29. List of recorded localities for Troglocyphoniscus absoloni
Locality
UTM
Region Source
Polu{pilja ispod ku}e u Bastu (cave), Bast, Biokovo, Ba{ka Voda
XJ60
SL
Jama na Mehovnju (cave), Veliki hom, Postrane, @rnovo, Kor~ula
XH65
SL
CBSS collection 498
Jama u Dubokom dolu (cave), Duboki dol, Pupnat, Kor~ula
XH65
SL
CBSS collection 499
Samograd (cave), Ra~i{}e, Kor~ula
XH66
SL
CBSS collection 500
Bubnjava~a (cave), Veprinac, Biokovo, Makarska
XH69
SL
CBSS collection 501
Jakasova {pilja (cave), Glava, Rasohatica, @rnovo, Kor~ula
XH75
SL
CBSS collection 502
[pilja kod Solina (cave), Soline, Mljet
XH93
SL
CBSS collection 497
CBSS collection 503
MZUF collection 20
STROUHAL, 1939a
STROUHAL, 1939c
STROUHAL, 1939e
STROUHAL, 1940b
Movrica (cave), Dra~evac, Babino Polje, Mljet
YH03
SL
VANDEL, 1946
ARCANGELI, 1947
KARAMAN, 1966
BEDEK et al., 2006
JAL@I] et al., 2010
295
Nat. Croat. Vol. 20(2), 2011
Gali~njak (cave), Okuklje, Mljet
YH13
SL
CBSS collection 504
SCHMÖLZER, 1965a
Mljet
SL
TABACARU, 1993
SCHMALFUSS, 2003
OZIMEC & BEDEK, 2005
From the Neretva to the S Montenegro region
SL
Croatia
STROUHAL, 1940a
POTO^NIK, 1989
29. Cyphopleon kratochvili (Frankenberger, 1939) (Tab. 30, Fig. 25)
Synonyms:
Distribution:
Type locality:
Cyphoniscellus kratochvili Frankenberger, 1939
Croatia: central Dalmatia (SCHMALFUSS, 2003)
[pilja na Svilaji planini (cave), Mt. Svilaja, Vrlika, Dalmatia, Croatia [einer
Höhle an der Svilaja Planina (Mitteldalmatien) (FRANKENBERGER, 1939)]
Ecological status: Troglophilic (STROUHAL, 1940a; BUTUROVI], 1957)
Fig. 25. Distribution of Cyphopleon kratochvili.
Tab. 30. List of recorded localities for Cyphopleon kratochvili
Locality
UTM
Region Source
Miljacka II (cave), Krka, Oklaj, Knin
WJ67
SL
[pilja kod Mlina na Miljacki (cave), Krka, Oklaj, Knin
WJ67
SL
CBSS collection 505
MZUF collection 21
CBSS collection 506
296
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Stara jametina (cave), Pun~ka draga, G. Krni}i, Konjevrate,
[ibenik
WJ84
SL
CBSS collection 507
Jama nasuprot Torka (cave), Torak, kanjon ^ikole, Klju~, Drni{
WJ85
SL
CBSS collection 508
Topla pe} (cave), Klju~, ^ikola, Skradin
WJ85
SL
CBSS collection 509
Sedrena {pilja iza mlina (cave), kanjon Krke, Burze, Kistanje,
WJ87
Knin
SL
CBSS collection 510
Golubnja~a (cave), Srebrni~ki potok, Hajdarovci, Srb, Plje{ivica
WK81 M
BUTUROVI], 1963
FRANKENBERGER, 1939
FRANKENBERGER, 1940
[pilja na Svilaji planini (cave), Svilaja, Vrlika
XJ
SL
KARAMAN, 1966
SCHMIDT & LEISTIKOW,
2004
BEDEK et al., 2006
Middle Dalmatia
SL
Central Dalmatia
SL
TABACARU, 1993
SCHMÖLZER, 1965a
SCHMALFUSS, 2003
From SE Italy to the Cetina region
STROUHAL, 1940a
Croatia
POTO^NIK, 1989
30. Cyphoniscellus herzegowinensis (Verhoeff, 1900) (Tab. 31, Fig. 26)
Synonyms:
Distribution:
Type locality:
Cyphoniscus herzegowinensis Verhoeff, 1900
Herzegovina (SCHMALFUSS, 2003); Croatia (POTO^NIK, 1989)
Vu~ja pe}ina (cave), Trebinje, Bosnia and Herzegovina [Wolfshöhle bei Trebinje (VERHOEFF, 1900)]
Ecological status: Troglophilic (STROUHAL, 1940a; BUTUROVI], 1957)
All known localities are subterranean, and the species is often found deep
in caves, so this species is most probably troglobiotic.
Tab. 31. List of recorded localities for Cyphoniscellus herzegowinensis
Locality
UTM
Region Source
Aragonka (cave), Ljuba~, Dubrovnik
BN53
SL
Jama na vrh Vrguda (cave), Vrgud, Osojnik, Dubrovnik
BN53
SL
CBSS collection 511
CBSS collection 512
CNHM collection 66
Mo~iljska {pilja (cave), Somino brdo, Osojnik, Dubrovnik
BN53
SL
CBSS collection 513
OZIMEC, 2005
STROUHAL, 1939c
STROUHAL, 1939e
[ipun (cave), Cavtat, Dubrovnik
BN71
SL
FRANKENBERGER &
STROUHAL, 1940
KARAMAN, 1966
FRANKENBERGER, 1939
Vilina pe}ina (cave), Gruda, Konavle, Cavtat
BN80
SL
FRANKENBERGER &
STROUHAL, 1940
Samograd (cave), Ra~i{}e, Kor~ula
XH66
SL
CBSS collection 514
KARAMAN, 1966
297
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Vilenska pe} (cave), Ogra|enica, Gumanca, Majkovi, Slano
YH34
SL
CBSS collection 515
Southern Dalmatia
SL
STROUHAL, 1940b
From the Neretva to the S Montenegro region
SL
STROUHAL, 1940a
POTO^NIK, 1989
Croatia
Fig. 26. Croatian distribution of Cyphoniscellus herzegowinensis.
TRICHONISCIDAE: BUDDELUNDIELLINAE
31. Buddelundiella cataractae Verhoeff, 1930 (Tab. 32, Fig. 27)
Distribution:
Type locality:
Europe (SCHMALFUSS, 2003)
Krka (river, between waterfalls), Skradin, Dalmatia, Croatia [zwischen den
Kerkawasserfällen Mitteldalmatiens oberhalb Skardona (VERHOEFF, 1930)]
Ecological status: Widespread species, already recorded in caves in Liguria (VANDEL, 1960b)
and in Sicily (CARUSO & BRISOLESE, 1974), common in caves in Croatia,
thus we consider it a troglophilic species.
Tab. 32. List of recorded localities for Buddelundiella cataractae
Locality
UTM
Region Source
Sumpora~a (cave), Komolac, Dubrovnik
BN62
SL
CBSS collection 516
\urovi}a {pilja (cave), \urovi}i, Cavtat, Konavle
BN71
SL
CBSS collection 517
298
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Urinjska {pilja (cave), Urinj, Kostrena, Rijeka
VL61
NL
Mandalina (cave), [ibenik
WJ74
SL
CBSS collection 518
LUKI] et al., 20111
CBSS collection 519
VERHOEFF, 1930
Krka (river, between waterfalls), Skradin
WJ75
SL
SCHMÖLZER, 1965a
FLASAROVÁ, 1966
KARAMAN, 1966
Dalmatia
SL
1
VERHOEFF, 1931c
PAOLETTI, 1978
Croatia
Buddelundiella cf. cataractae
Fig. 27. Croatian distribution of Buddelundiella cataractae.
TRICHONISCIDAE: THAUMATONISCELLINAE
32. Thaumatoniscellus speluncae Karaman, Bedek et Horvatovi}, 2009
(Tab. 33, Fig. 28)
Distribution:
Type locality:
Croatia: Mt. U~ka (KARAMAN et al., 2009)
Jama kod potoka Banine 3 (cave), Menderi, Veprinac, U~ka, Croatia [Pit 3
near Banina Stream (KARAMAN et al., 2009)]
Ecological status: Troglobiotic (KARAMAN et al., 2009)
299
Nat. Croat. Vol. 20(2), 2011
Fig. 28. Distribution of Thaumatoniscellus speluncae.
Tab. 33. List of recorded localities for Thaumatoniscellus speluncae
Locality
UTM
Region Source
Jama iznad Vranjske drage (cave), Pikuli}i, Boljun, Vela
draga, U~ka
VL31
NL
CBSS collection 520
MZUF collection 22
CBSS collection 521
Jama kod potoka Banine 3 (cave), Menderi, Veprinac, U~ka
VL41
NL
DBE collection 1
KARAMAN et al., 2009
JAL@I] et al., 2010
PHILOSCIIDAE
33. Chaetophiloscia cellaria (Dollfus, 1884) (Tab. 34, Fig. 29)
Synonyms:
Chaetophiloscia italica (Verhoeff, 1901); C. magnopunctata (Strouhal, 1929); C.
pallida Verhoeff, 1928; C. piligera Verhoeff, 1908; C. pseudocellaria (Arcangeli,
1934); C. solerii (Arcangeli, 1937); Neophiloscia magnopunctata Strouhal, 1929;
Philoscia cellaria Dollfus, 1884; P. italica Verhoeff, 1901; P. pseudocellaria Arcangeli, 1934; P. solerii Arcangeli, 1937
Distribution:
Northern Mediterranean region from Spain to Greece (SCHMALFUSS, 2003)
Type locality:
Beaune (basements), Côte-d’Or, France [caves de Beaune (Côte-d’Or) (DOLLFUS, 1884)]
Ecological status: Troglophilic (VANDEL, 1962)
300
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Tab. 34. List of recorded localities for Chaetophiloscia cellaria
Locality
UTM
Region Source
Rudnik u Medveji (mine), Medveja, Lovran
VL41
NL
CBSS collection 522
Opatija
VL42
NL
ARCANGELI, 1952
[pilja iznad ceste (cave), brodogradili{te V. Lenac,
Kostrena, Rijeka
VL51
NL
Mali {tumbur (cave), Perilo, Kostrena, Rijeka
VL61
NL
Urinjska {pilja (cave), Urinj, Kostrena, Rijeka
VL61
NL
Mandalina (cave), [ibenik
WJ74
SL
CBSS collection 523
LUKI] et al., 2011
CBSS collection 524
LUKI] et al., 2011
CBSS collection 525
LUKI] et al., 2011
CBSS collection 526
ARCANGELI, 1947
Istria
NL
ARCANGELI, 1950
VANDEL, 1962
Croatia
POTO^NIK, 1989
MISIDENTIFICATION
Gali~njak (cave), Okuklje, Mljet*
MICULINI], 20081
*Examination of the collection shows that the specimens belong to other not yet identified species
1
Chaetophiloscia cf. cellaria
Fig. 29. Croatian distribution of Chaetophiloscia cellaria.
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Nat. Croat. Vol. 20(2), 2011
ONISCIDAE
34. Oroniscus dalmaticus Strouhal, 1937 (Tab. 35, Fig. 30)
Synonyms:
Strouhalius dalmaticus (Strouhal, 1937); Strouhaloniscus dalmaticus (Strouhal,
1937)
Distribution:
Croatia: Dalmatia (SCHMALFUSS, 2003)
Type locality:
Vranja~a (cave), Kotlenice, Dugopolje, Split, Dalmatia, Croatia [Höhle Vranja~a« bei Dugopolje, am Fuße des Mosorgebirges; BB Nr. 334 (STROUHAL,
1937)]
Ecological status: Troglophilic (STROUHAL, 1940a)
Tab. 35. List of recorded localities for Oroniscus dalmaticus
Locality
UTM
Region Source
Jama u Bratu{u (cave), Bratu{, Makarska
XH69
SL
Kuna (cave), ^edor, Biokovo, Makarska
XH69
SL
CBSS collection 528
Gradska spila (cave), @upa, Biokovo, Zagvozd
XH79
SL
CBSS collection 529
Matija{eva {pilja (cave), @upa, Biokovo, Zagvozd
XH79
SL
CBSS collection 530
Poskokova rupa (cave), @upa, Biokovo, Zagvozd
XH79
SL
CBSS collection 531
Stonjska pe} donja (cave), @upa, Biokovo, Zagvozd,
Zagvozd
XH79
SL
CBSS collection 532
Tu~epska vilenja~a (cave), Ravna vla{ka, Biokovo,
Tu~epi
XH79
SL
CBSS collection 533
CBSS collection 527
MZUF collection 23
STROUHAL, 1937
ARCANGELI, 1939b
Vranja~a (cave), Kotlenice, Mosor, Dugopolje, Split
XJ32
SL
FRANKENBERGER &
STROUHAL, 1940
SCHMÖLZER, 1965a
KARAMAN, 1966
BEDEK et al., 2006
Drinova II (cave), Drinova, Gornja Brela, Biokovo, Brela
XJ50
SL
CBSS collection 534
Jama iznad Prozor~i}a (cave), Nevistine stine, Gornja
Brela, Biokovo, Brela
XJ50
SL
CBSS collection 535
Jamina (cave), Nevistine stine, Gornja Brela, Biokovo, Brela XJ50
SL
CBSS collection 536
Male spile (cave), Crvene stine, Topi}i, Biokovo, Ba{ka
Voda
XJ50
SL
CBSS collection 537
Kukor (cave), Bast, Biokovo, Ba{ka Voda
XJ60
SL
CBSS collection 538
Mala jama u stijenama (cave), Spiljice, Kaoci, Muci}i,
Biokovo, Zagvozd
XJ60
SL
CBSS collection 539
Ogra|ena {pilja pod Sv. Jurom (cave), Sv. Jure, Biokovo,
XJ60
Makarska
SL
CBSS collection 540
Pe}ina kod Zagvozda (cave), Zagvozd, Biokovo
SL
XJ60
BUTUROVI], 1955c
KARAMAN, 1966
302
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BUTUROVI], 1955c
Pe}ina u Sridnjoj gori (cave), Sridnja gora, Zagvozd,
Biokovo
XJ60
SL
Pru`ina (cave), Prodani, Biokovo, Zagvozd
XJ60
SL
CBSS collection 541
[pilja u Bastu (cave), Bast, Biokovo, Ba{ka Voda
XJ60
SL
CBSS collection 542
^o}ina jama (cave), Bebi}i, Metkovi}
YH07
SL
Biokovo
SL
Mosor
SL
KARAMAN, 1966
BUTUROVI], 1955c
KARAMAN, 1966
OZIMEC, 2008
BEDEK, 2011
FRANKENBERGER, 1938b
GOTTSTEIN et al., 2001
GOTTSTEIN MATO^EC et al.,
2002
Dalmatia
SCHMALFUSS, 2003
From SE Italy to the Cetina region
STROUHAL, 1940a
Croatia
POTO^NIK, 1989
Fig. 30. Distribution of Oroniscus dalmaticus.
35. Oroniscus meledensis Strouhal, 1937 (Tab. 36, Fig. 31)
Synonyms:
Distribution:
Strouhalius meledensis (Strouhal, 1937); Strouhaloniscus meledensis (Strouhal,
1937)
Croatia: the island of Mljet (SCHMALFUSS, 2003)
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Nat. Croat. Vol. 20(2), 2011
Type locality:
Osta{evica (cave), Dra~evac, Babino Polje, the island of Mljet, Dalmatia, Croatia [»[pilja Osta{evica« auf der Insel Meleda; BB Nr. 660 (STROUHAL, 1937)]
Ecological status: Troglophilic (STROUHAL, 1940a)
Tab. 36. List of recorded localities for Oroniscus meledensis
Locality
UTM
Region
Source
STROUHAL, 1937
STROUHAL, 1939e
ARCANGELI, 1939b
Osta{evica (cave), Dra~evac, Babino Polje, Mljet
YH03
SL
FRANKENBERGER &
STROUHAL, 1940
KARAMAN, 1966
GOTTSTEIN et al., 2001
GOTTSTEIN MATO^EC et
al., 2002
BEDEK et al., 2006
FRANKENBERGER, 1938b
Mljet
SL
From the Neretva to the S Montenegro region
SL
SCHMÖLZER, 1965a
SCHMALFUSS, 2003
Croatia
STROUHAL, 1940a
POTO^NIK, 1989
Fig. 31. Distribution of Oroniscus meledensis.
304
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36. Oroniscus stentai (Arcangeli, 1926) (Tab. 37, Fig. 32)
Synonyms:
Porcellio stentai Arcangeli, 1926; Strouhalius stentai (Arcangeli, 1926); Strouhaloniscus stentai (Arcangeli, 1926)
Distribution:
Croatia (SCHMALFUSS, 2003)
Type locality:
^injadra (cave), Nere`i{}a, the island of Bra~, Dalmatia, Croatia [Grotta
Cinjadra nell’Isola Brazza (ARCANGELI, 1926)]
Ecological status: Troglophilic (STROUHAL, 1940a)
Fig. 32. Distribution of Oroniscus stentai.
Tab. 37. List of recorded localities for Oroniscus stentai
Locality
UTM
Region
Source
FRANKENBERGER, 1938b1
ARCANGELI, 1939b
[pilja kod Murvice (cave), Murvica, Bol, Bra~
XH29 SL
FRANKENBERGER &
STROUHAL, 1940
KARAMAN, 1966
GOTTSTEIN et al., 2001
GOTTSTEIN MATO^EC et al.,
2002
305
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CBSS collection 543
ARCANGELI, 1926
ARCANGELI, 1939b
KARAMAN, 1966
^injadra (cave), Nere`i{}a, Bra~
XJ30
SL
GOTTSTEIN et al., 2001
GOTTSTEIN MATO^EC et al.,
2002
BEDEK et al., 2006
JAL@I] et al., 2010
Bra~
SCHMÖLZER, 1965a
SL
STROUHAL, 1940a
From SE Italy to the Cetina region
POTO^NIK, 1989
Croatia
1
SCHMALFUSS, 2003
Oroniscus dalmaticus
ARMADILLIDIIDAE: ELUMINAE
37. Typhlarmadillidium kratochvili (Frankenberger, 1938)
(Tab. 38, Fig. 33)
Synonyms:
Cylisticus kratochvili Frankenberger, 1938; Synarmadillidium kratochvili (Frankenberger, 1938); Troglarmadillidium kratochvili (Frankenberger, 1938)
Distribution:
Croatia: the island of Kor~ula (SCHMALFUSS, 2003)
Type locality:
Jakasova {pilja (cave), Glava, Rasohatica, @rnovo, the island of Kor~ula,
Dalmatia, Croatia [der Höhle bei Rasokatica im Gebirge Glava in der Umgebung des Dorfes Postrana auf der Insel Kor~ula (FRANKENBERGER, 1938b)]
Ecological status: Troglophilic (troglobiotic?) (STROUHAL, 1940a)
Tab. 38. List of recorded localities for Typhlarmadillidium kratochvili
Locality
UTM
Region Source
FRANKENBERGER, 1938b
ARCANGELI, 1939c
FRANKENBERGER, 1940
Jakasova {pilja (cave), Glava, Rasohatica, @rnovo, Kor~ula
XH75 SL
FRANKENBERGER &
STROUHAL, 1940
SCHMÖLZER, 1965b
KARAMAN, 1966
BEDEK et al., 2006
OZIMEC, 2010
From the Neretva to the S Montenegro region
SL
STROUHAL, 1940a
FERRARA & TAITI, 1996
GOTTSTEIN et al., 2001
Kor~ula
SL
GOTTSTEIN MATO^EC et
al., 2002
SCHMALFUSS, 2003
306
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Croatia
POTO^NIK, 1989
MISIDENTIFICATION
Samograd (cave), Ra~i{}e, Kor~ula*
XH66 SL
OZIMEC, 2004
* Re-examination of this material shows that no material belonging to this species was collected.
Fig. 33. Distribution of Typhlarmadillidium kratochvili.
38. Typhlarmadillidium trebinjanum (Verhoeff, 1900) (Tab. 39, Fig. 34)
Synonyms:
Armadillidium trebinjanum Verhoeff, 1900; Troglarmadillidium trebinjanum Verhoeff, 1900
Distribution:
Croatia, Herzegovina, Montenegro (SCHMALFUSS, 2003)
Type locality:
Trebinje (surroundings), Bosnia and Herzegovina [der Umgegend von Trebinje in der Herzegowina (VERHOEFF, 1900)]
Ecological status: Troglophilic (STROUHAL, 1940a)
Tab. 39. List of recorded localities for Typhlarmadillidium trebinjanum
Locality
UTM
Region
Source
[ipun (cave), Cavtat, Dubrovnik
BN71
SL
MZUF collection 24
Je{kalovica (cave), Selca, Bra~
XH49 SL
STROUHAL, 1939c
KARAMAN, 1966
307
Nat. Croat. Vol. 20(2), 2011
Bra~
SL
FRANKENBERGER &
STROUHAL, 1940
SCHMÖLZER, 1965b
FERRARA & TAITI, 1996
From SE Italy to the Cetina region
From the Neretva to the S Montenegro region
STROUHAL, 1940a
SL
STROUHAL, 1940a
POTO^NIK, 1989
Croatia
SCHMALFUSS, 2003
Fig. 34. Croatian distribution of Typhlarmadillidium trebinjanum.
ARMADILLIDIIDAE: ARMADILLIDIINAE
39. Echinarmadillidium fruxgalii (Verhoeff, 1900) (Tab. 40, Fig. 35)
Synonyms:
Distribution:
Type locality:
Armadillidium fruxgalii Verhoeff, 1900
Croatia: s-Dalmatia, Montenegro (SCHMALFUSS, 2003)
Ombla (river bank) and Lapad, Dubrovnik, Dalmatia, Croatia [Omblathale,
... der Halbinsel Lapad bei Gravosa (VERHOEFF, 1900)]
Ecological status: Troglophilic (STROUHAL, 1940a)
This species is often found in caves, so we agree with STROUHAL in considering it as a troglophilic species.
308
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
Tab. 40. List of recorded localities for Echinarmadillidium fruxgalii
Locality
UTM
Region Source
VERHOEFF, 1900
CSIKI, 1926
STROUHAL, 1934
Lapad, Dubrovnik
BN52
SL
FRANKENBERGER &
STROUHAL, 1940
KARAMAN, 1966
SCHMIDT & LEISTIKOW, 2004
VERHOEFF, 1900
CSIKI, 1926
STROUHAL, 1934
Ombla (river bank), Dubrovnik
BN62
SL
FRANKENBERGER, 1938b
FRANKENBERGER &
STROUHAL, 1940
KARAMAN, 1966
SCHMIDT & LEISTIKOW, 2004
FRANKENBERGER, 1938b
ARCANGELI, 1939c
Caretova {pilja (cave), Vis, Vis
WH96 SL
FRANKENBERGER &
STROUHAL, 1940
KARAMAN, 1966
Stra~in~ica (cave), Vela Luka, Kor~ula
XH45
SL
CBSS collection 544
CBSS collection 545
FRANKENBERGER, 1938b
Tabaina {pilja (cave), Vela Luka, Kor~ula
XH45
SL
ARCANGELI, 1939c
FRANKENBERGER &
STROUHAL, 1940
KARAMAN, 1966
CBSS collection 546
Hrastovska {pilja (cave), Hrastovsko polje, Lastovo,
Lastovo
XH53
SL
MZUF collection 25
OZIMEC et al., 2008
Jama Cvjetana Barbi}a (cave), Skrivena Luka, Lastovo
XH53
SL
Jama u uvali Kru~ica (cave), Zaklopatica, Lastovo, Lastovo XH53
SL
Jama za Smokvom (cave), Skrivena Luka, Lastovo
XH53
SL
Lastovo, Lastovo
XH53
SL
Medvi|a ropa (cave), Skrivena Luka, Lastovo
XH53
SL
CBSS collection 547
OZIMEC et al., 2008
CBSS collection 548
OZIMEC et al., 2008
CBSS collection 549
OZIMEC et al., 2008
CBSS collection 550
CBSS collection 551
OZIMEC et al., 2008
309
Nat. Croat. Vol. 20(2), 2011
[pilja na Pozalicu (cave), Pr`ina, Skrivena Luka, Lastovo
[pilja pod Ka{tel (cave), Ka{tel, Lastovo, Lastovo
XH53
XH53
SL
SL
Samograd (cave), Ra~i{}e, Kor~ula
XH66
SL
Jama u Bratu{u (cave), Bratu{, Makarska
XH69
SL
CBSS collection 552
OZIMEC et al., 2008
CBSS collection 553
OZIMEC et al., 2008
CBSS collection 554
OZIMEC, 2004
CBSS collection 555
FRANKENBERGER, 1938b
Marijan, Split
XJ11
SL
FRANKENBERGER &
STROUHAL, 1940
KARAMAN, 1966
Bast (endogean, forest), Biokovo, Ba{ka Voda
XJ60
SL
CBSS collection 556
[pilja u Bastu (cave), Bast, Biokovo, Ba{ka Voda
XJ60
SL
CBSS collection 557
VERHOEFF, 1939
Southern Dalmatia
SL
SCHMÖLZER, 1965b
SCHMALFUSS, 2003
Dalmatia
SL
VERHOEFF, 1931c
SL
STROUHAL, 1940a
From SE Italy to the Cetina region
From the Neretva to the S Montenegro region
Croatia
STROUHAL, 1940a
POTO^NIK, 1989
Fig. 35. Croatian distribution of Echinarmadillidium fruxgalii.
310
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
40. Cyphodillidium absoloni (Strouhal, 1934) (Tab. 41, Fig. 36)
Synonyms:
Distribution:
Type locality:
Echinarmadillidium absoloni Strouhal, 1934
Croatia: the island of Mljet (SCHMALFUSS, 2003)
Movrica (cave), Dra~evac, Babino Polje, the island of Mljet, Dalmatia, Croatia [Pe~ina Movrica auf der Insel Meleda (STROUHAL, 1934)]
Ecological status: Troglophilic
Tab. 41. List of recorded localities for Cyphodillidium absoloni
Locality
UTM
Region
Jama u Velikom Desajdubu (cave), Veliki Desajdub,
Pola~a, Mljet
XH93
SL
Male ponte (cave), Grabova glava, Ropa, Mljet
[pilja kod Solina (cave), Soline, Mljet
XH93
XH93
SL
SL
[pilja kod vrha Spile (cave), Vrije{}e, Pola~a, Mljet
XH93
SL
Bezdan (cave), Crna klada, Ropa, Mljet
Jama za Svetim Spasom (cave), Babino Polje, Mljet
YH03
YH03
SL
SL
Movrica (cave), Dra~evac, Babino Polje, Mljet
YH03
SL
Odisejeva {pilja (cave), Babino Polje, Mljet
YH03
SL
[pilja na brdu Bugara (cave), Bugara, Ropa, Mljet
YH03
SL
Velika {pilja (cave), Blata, Mljet
YH03
SL
Breg jama (cave), Babino Polje, Mljet
YH13
SL
Gali~njak (cave), Okuklje, Mljet
YH13
SL
Jama za Svetim Ivanom (cave), Babino Polje, Mljet
Polu{pilja kod Sobre (cave), Sobra, Mljet
YH13
YH13
SL
SL
Mljet
SL
Dalmatia
Croatia
SL
1
Cyphodillidium
Source
CNHM collection 67
CBSS collection 558
CBSS collection 559
CBSS collection 560
CNHM collection 68
CBSS collection 561
CBSS collection 562
CBSS collection 563
STROUHAL, 1934
FRANKENBERGER, 1938b
FRANKENBERGER &
STROUHAL, 1940
STROUHAL, 1939e
KARAMAN, 1966
SCHMIDT & LEISTIKOW, 2004
BEDEK et al., 2006
JAL@I] et al., 2010
CBSS collection 564
FRANKENBERGER, 1938b
ARCANGELI, 1939c
KARAMAN, 1966
CBSS collection 565
FRANKENBERGER, 1938b
ARCANGELI, 1939c
KARAMAN, 1966
CBSS collection 566
CBSS collection 567
MICULINI], 2008
CBSS collection 568
MZUF collection 26
FRANKENBERGER &
STROUHAL, 1940
SCHMÖLZER, 1965b
GOTTSTEIN et al., 20011
GOTTSTEIN MATO^EC et al.,
20021
SCHMALFUSS, 2003
OZIMEC & BEDEK, 2005
VERHOEFF, 1939
POTO^NIK, 1989
Nat. Croat. Vol. 20(2), 2011
311
Fig. 36. Distribution of Cyphodillidium absoloni.
DISCUSSION
The karst area in Croatia covers approximately 46% of the national territory
(GOTTSTEIN MATO^EC et al., 2002). More than 9000 caves have been registered in
Croatia (BAK[I] & NOVOSEL), but literature data concerning subterranean terrestrial
isopods include only 221 caves (~2,5 %) and 14 various localities (mines, endogean
and epigean environment). Altogether, with the new records from collections, this
paper presents data from 557 caves (~6,2 %), six artificial underground structures and
15 other epigean and subterranean localities (endogean, soil, etc.). Probably some localities from the collections are already present in the literature, but the old names
are in some cases impossible to be referred to names of localities known at present.
Altogether 765 different records (one locality with one species) were gathered.
Among them 275 records (~35,9 %) come from literature data, 601 (~78,6 %) from
collection data, and 112 (~14,6 %) from both literature and collection data.
Thirty-five species and five subspecies of the cave-dwelling Oniscidea have been
recorded. They are divided into five families and six subfamilies. The family Trichoniscidae and the subfamily Trichoniscinae are the most representative and include
13 (59 %) out of 22 Croatian endemic taxa, and 17 (85 %) out of 20 troglobiotic species
(Tab. 42). The most represented genus is Alpioniscus with 10 species, followed by
Androniscus with eight taxa (Fig. 37). Out of 26 cave species described in Croatia, 16
312
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
(~62 %) were described in the thirties of the last century, which was the result of intensive sampling done mostly in Dalmatia (Fig. 38).
Tab. 42. Number of genera, species, subspecies, endemic, troglobiotic and troglophilic taxa in
different families and subfamilies
Family/Subfamily
Genera
Spp.
(sspp.)
Endemic
spp. (sspp.)
Troglobiotic
spp. (sspp.)
Troglophilic
spp. (sspp.)
Mesoniscidae
1
1
0
0
1
Trichoniscidae
12
26 (5)
12 (5)
19 (1)
7 (4)
• Trichoniscinae
6
20 (5)
8 (5)
16 (1)
4 (4)
• Haplophthalminae
4
4
3
2
2
• Buddelundiellinae
1
1
0
0
1
• Thaumatoniscellinae
1
1
1
1
0
Philosciidae
1
1
0
0
1
Oniscidae
1
3
3
0
3
Armadillidiidae
3
4
2
2
2
• Eluminae
1
2
1
0
2
• Armadillidiinae
2
2
1
0
2
18
35 (5)
17 (5)
19 (1)
16 (4)
Total
Only 165 (~20,3 %) out of 814 Croatian 10 x 10 km UTM squares have one or more
cave species. However, plots with at least one cave species are more or less scattered all over the Dinaric region in Croatia. The UTM square VL41 belonging to the
Northern Croatian Littoral macroregion has the maximum number of species (six),
Fig. 37. Frequency distribution of genera of cave-dwelling terrestrial isopods in
Croatia. Numbers refer to taxa in each genus.
Nat. Croat. Vol. 20(2), 2011
313
Fig. 38. Comparison of the number of cave species and subspecies described in Croatia
from 1890 to present.
followed by plots BN 62, BN71, XH66, XH69 and YH03 belonging to the Southern
Croatian Littoral macroregion with five species (Fig. 39). The islands of Mljet and
Kor~ula, mountains of Biokovo and U~ka, surroundings of the town of Dubrovnik,
and the town of Ogulin are biodiversity hot spots of cave-dwelling terrestrial isopods
in Croatia. The present situation might be changed when intensive investigations
are carried out in still poorly explored karstic regions like Mt. Velebit.
The Southern Croatian Littoral and the Northern Croatian Littoral macroregions
have many more species than the other Croatian macroregions. Out of 40 cave-dwelling taxa in Croatia the Southern Croatian Littoral macroregion has 27 (67,5 %) and
the Northern Croatian Littoral macroregion has 15 (37,5 %) (Fig. 40).
The Southern Croatian Littoral macroregion has 16 (~72,7 %) out of 22 endemic
cave taxa in Croatia, and among them, two are widespread and 14 are endemic to
that region. Cavernicolous terrestrial isopods are recorded from nine Croatian islands
on a total of 79 islands and 523 islets. Eight species are endemic to a single island
and occur on five islands. All the 40 cave-dwelling taxa in Croatia are distributed
in the Dinaric karst area, and only three troglophilic species are also found outside
that area (Androniscus dentiger, A. roseus and Mesoniscus graniger). There are no
endemic taxa outside the Dinaric karst area (Fig. 41).
Distribution areas of the different species are unequal. Some troglophilic species
are widespread, such as Mesoniscus graniger in Slovenia, Croatia, Serbia, Bosnia and
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Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
Fig. 39. 10 × 10 km UTM grid with numbers of species and subspecies of cave-dwelling
terrestrial isopods in each square.
Herzegovina, Slovakia, Romania; Androniscus roseus from eastern France to Romania; Chaetophiloscia cellaria in the northern Mediterranean region; Trichoniscus matulici in southern Italy including Sicily, Slovenia, Croatia, Bosnia and Herzegovina
and Montenegro. Some species are probably spread synanthropically such as
Buddelundiella cataractae in Europe; Androniscus dentiger in Europe and North Africa,
and in greenhouses in North America. The troglobiotic species Aegonethes cervinus
is distributed across the Adriatic Sea in Italy (Gargano) and Croatia (Dubrovnik,
the islands of Kor~ula and Mljet). In Croatia 23 species (out of 35) are distributed
only in single macroregion. The most widely distributed taxa in Croatia are the
troglophilic species Mesoniscus graniger and Androniscus roseus and the troglobiotic
species A. stygius, found in three macroregions. Only four troglobiotic species (Alpio-
Nat. Croat. Vol. 20(2), 2011
315
Fig. 40. Frequency distribution of species and subspecies present in Croatian macroregions. Numbers refer to taxa present in each macroregion – E (Eastern Croatia); C (Central Croatia); M (Mointainous Croatia); NL (northern Croatian Littoral); SL (Southern
Croatian Littoral)
Fig. 41. Distribution of Croatian endemic taxa.
316
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
Fig. 42. Comparison of terrestrial isopod species inhabiting various number of Croatian
macroregions with indicated ecological adaptation to the cave environment.
niscus balthasari, A. strasseri, Titanethes albus and T. dahli) are distributed in two macroregions, the rest are distributed only in a single macroregion (Fig. 42).
Recent data (published after 2005 or in this paper) have revealed new insights
into several species. Although the species Mesoniscus graniger is not listed in some
reviews (KARAMAN, 1966; POTO^NIK, 1989; SCHMALFUSS, 2003) there are old literature
data for Croatia (ARCANGELI, 1939a). This record is from Istria, but in recent investigations in that region the species was not found, either in caves or in endogean
habitats. Mesoniscus graniger was instead found on Mt. Medvednica, Mt. Ivan{~ica,
Mt. Papuk, the Banovina region and near Karlovac. Only the last locality is part of
the Dinaric region (BEDEK, 2007b) (Tab. 1; Fig. 2). Mesoniscus graniger is one of the
few cave isopods found in Croatia outside the Dinaric region.
The only known record in the literature for Alpioniscus heroldi is Benetina pe}ina
near Slano (STROUHAL, 1939a). This cave is probably in Popovo polje in Herzegovina, near Slano, so there is no confirmed literature record of A. heroldi in Croatia.
However, during recent investigations this species was found in several Croatian
caves (Tab. 16; Fig. 13).
Alpioniscus herzegowinensis was recorded from the cave system Lukina jama –
Trojama, Hajdu~ki kukovi, Mt. Velebit, Krasno (KEROVEC et al., 1999). All collected
material of the genus Alpioniscus from Lukina jama – Trojama and other deep pits
in northern Mt. Velebit belong to other species, close to the strasseri-group (unpublished data). The species A. herzegowinensis is endemic for the [uma region in SW
Herzegovina, where microclimate conditions in caves are totally different from
those of the deep pits of northern Mt. Velebit where air temperature in deep pits is
more than 10°C lower, so this is probably a misidentification (Tab. 17).
The only known literature data for the species Alpioniscus verhoeffi is [pilja u
dolini Neretve, Neretva, Dalmatia (STROUHAL, 1938) which can be in Dalmatia or
Herzegovina, and Bosnia is probably a misinterpreted datum (SCHMALFUSS, 2003).
This species has been collected in several caves near Neretva River and, very surprisingly, also in [pilja na Vr{inoj glavici, Stra`benica, Otok, Sinj (Tab. 22; Fig. 18).
Nat. Croat. Vol. 20(2), 2011
317
The new records resulted in a better understanding of the distribution pattern of
several species listed in Croatia. For the species Alpioniscus balthasari, the limits of
its distribution were extended by 100 km (Tab. 12, Fig. 9). Androniscus wolfi was
found in the south part of Mt. Velebit (93 km from the only other known locality)
(Tab. 9, Fig. 6), Titanethes dahli on Mt. Velebit (Tab. 24, Fig. 20), Trichoniscus matulici
metkovicensis on Mt. Biokovo (Tab. 27, Fig. 22), Buddelundiella cataractae in the Dubrovnik and Rijeka regions (Tab. 32, Fig. 27), Echinarmadillidium fruxgalii and Aegonethes
cervinus on the island of Lastovo (Tab. 40, Fig. 35; Tab. 11, Fig. 8), Cyphoniscellus
herzegowinensis on the island of Kor~ula (Tab. 31, Fig. 26) and Troglocyphoniscus
absoloni on Mt. Biokovo and the island of Kor~ula (Tab. 29, Fig. 24).
Some species cited in the literature have not been confirmed by recent, often repeated explorations in those caves such as, Oroniscus meledensis from the island of
Mljet, and Typhlarmadillidium kratochvili from the island of Kor~ula.
In this paper only part of the CBSS collection is listed. It contains also many currently unidentified and several new species which will be treated in future papers.
Moreover, members of the Croatian Biospeleological Society are still collecting material through numerous projects. Many oniscidean genera need to be revised, and
many species to be redescribed. The state of understanding about cave terrestrial
isopods in Croatia will probably be modified as a result of a necessary extensive
taxonomic work.
ACKNOWLEDGEMENTS
Many acknowledgements to project leaders (in brackets) and financiers. Partially
supported by: Ministry of Environmental Protection and Physical Planning (Branko
Jal`i}), @umberak – Samoborsko gorje Nature Park (Roman Ozimec), Biokovo Nature Park (Roman Ozimec), the town of Dubrovnik (Marijana Frani~evi}, today
Cukrov), Rufford Small Grants for Nature Conservation (Jana Bedek), World Bank
(Nikola Tvrtkovi}), Netherlands Government (Nikola Tvrtkovi}), Medvednica Nature Park (Roman Ozimec), Dubrovnik – Neretva County (Roman Ozimec), Dubrovnik Airport (Tihomir Kova~evi}, Roman Ozimec), the town of Crikvenica (Branko Jal`i}), Krka National Park (Branko Jal`i}, Kazimir Miculini}), Papuk Nature
Park (Jana Bedek), U~ka Nature Park (Jana Bedek), Mljet National Park (Helena
Biland`ija, Ana Komeri~ki), Ministry of Culture (Helena Biland`ija), Kostrena Municipality Tourist Office (Marko Luki}, Ana Komeri~ki), State Institute for Nature
Protection (Branko Jal`i}, Roman Ozimec, Marko Luki}), Vara`din County (Roman
Ozimec), Sisak – Moslavina County (Roman Ozimec), Croatian Electricity Company
(Jana Bedek), European Union (Krunoslav Mla~ak, Ljiljana Dravec), Croatian Natural History Museum (Vedran Jal`i}), Velebit Nature Park (Jana Bedek).
We thank Marko Luki} for assistance in all stages of this work and for reviewing
the manuscript, Roman Ozimec who collected most of the literature and helped
with useful information, Branko Jal`i} who helped with identifying the majority of
localities from old literature and with other useful information. Dr Helmut Schmalfuss, Dr. Ferenc Vilisics, Dr Ivo Karaman, Darija ]aleta, Ivan Glava{ are particularly
to be thanked for their help in collecting the missing literature. We are grateful to
Slavko Polak, Dr. Fabio Stoch and Dr. Fulvio Gasparo for helping with names of
318
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
caves and localities, Dr Ivo Karaman for reviewing the manuscript, Hrvoje Cvitanovi} for correcting the names of caves, Ana Komeri~ki and Graham McMaster for
improving the English, and to the anonymous referees for valuable comments and
suggestions.
We would like to thank all collectors of the material (by alphabetical order): Robert Bakovi}, Marko Ba{i}, Helena Biland`ija, Igor Bor{i}, Petra Bregovi}, Ivan Budinski, Stipe Bu{eli}, Marijana Cukrov (Frani~evi}), Hrvoje Cvitanovi}, Tamara ^ukovi}, An|ela ]uku{i}, Ester Domina, Tvrtko Dra`ina, Marina Dr~eli}, Aljo{a
Dupli}, Lana \ud, Dajana Hmura, Ivica Jaklinovi}, Vatroslav Jakobovi}, Branko
Jal`i}, Vedran Jal`i}, Lucija Jankovi}-Rapan, Luka Katu{i}, Fanica Kljakovi} Ga{pi},
Marina Kipson, Alen Kirin, Ana Kolari}, Tommy Kolman, Ana Komeri~ki, Petra
Kova~ Konrad, Ivana Kova~i}, Marko Luki}, Marta Malenica, Marin Manger, Neven
Mato~ec, Kazimir Miculini}, Barbara Mikac, Sa{a Minihofer, Gorana Mi{}eni},
Kre{imir Moto~i}, Dr Jos Notenboom, Mislav Novokmet, Darijan Ozimec, Margita
Ozimec, Roman Ozimec, Goran Palijan, Martina Pavlek, Slavko Polak, Gordan
Poli}, Predrag Rade, Ton}i Ra|a, Maja Ra|enovi}, Nikolina Ragu`, Ilija Ra{i}, Tomica Rubini}, Dr Rajko Slapnik, S. [ore, Matija Vojnovi}, Milan Vojinovi}, Davorin
Vranekovi}, Snje`ana Vuj~i}-Karlo, Roko @aja and Dijana @upan.
And last but not least we thank all speleologists and colleagues from SO PDS
Velebit, SO HPD @eljezni~ar, DDSKF, SK Samobor, SK Ursus spelaeus, SU Estavela,
SU Spelunka, SO PD Dubovac, SD Karlovac and the Biological Students Society
BIUS who organised and co-financed the different speleological camps.
Received May 25, 2010
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S U M M A RY
Catalogue and atlas of cave-dwelling terrestrial isopods
(Crustacea: Oniscidea) from Croatia
J. Bedek, S. Taiti & S. Gottstein
In Croatia 130 species and 30 subspecies of terrestrial isopods have been recorded,
59 species and 17 subspecies of which are endemic to Croatia. Altogether, this paper
presents data from 557 caves, 6 artificial underground structures and 15 other epigean and subterranean localities. Among the cave taxa, 35 species and five subspecies have been recorded. The family Trichoniscidae and the subfamily Trichoniscinae
are the most abundant (13 (59%) out of 22 Croatian endemic taxa and 17 (85 %) out
of 20 troglobiotic taxa). The genus most represented is Alpioniscus with 10 species.
Only 165 out of 814 Croatian 10 × 10 km UTM squares (~20,3 %) have one or more
cave species. The UTM square VL41 presents the maximum with six species, belonging to the Northern Croatian Littoral macroregion. The UTM squares BN62, BN71,
XH66, XH69 and YH03 have five species, belonging to the Southern Croatian Littoral
macroregion. The Southern Croatian Littoral macroregion has 27 (67,5 %) out of 40
cave taxa in Croatia, and the Northern Croatian Littoral macroregion has 15 (37,5 %).
The Southern Croatian Littoral mesoregion has 16 (~72,7 %) out of 22 endemic cave
taxa in Croatia and among them, two are widespread and 14 are endemic to that region. Five islands have their own endemic species (eight species). The most widely
distributed species in Croatia are the troglophiles Mesoniscus graniger, Androniscus
roseus, and the troglobiont A. stygius found in three macroregions. Only four troglobiotic species (Alpioniscus balthasari, A. strasseri, Titanethes albus and T. dahli) are distributed in two macroregions, the remaining species are distributed only in single
macroregions. All the 40 cave-dwelling taxa in Croatia are distributed in the Dinaric
karst area, and only three troglophilic species are found also outside this area. No endemic taxa occur outside the Dinaric karst area. In this paper only part of the CBSS
collection is listed. The majority of genera need to be revised and many species
redescribed. The knowledge about cave terrestrial isopods in Croatia will probably
be modified as a result of an extensive taxonomic work which still needs to be done.
326
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
APPENDIX 1.
Tab. 43. List of synonyms for locality records of cave–dwelling terrestrial isopods in Croatia
with references associated with the recorded locality name.
Recorded locality name
Baba pe}ina (BUTUROVI], 1955c, KARAMAN, 1966)
Bali~eva pe}ina (STROUHAL, 1939a)
Bali}eva pe}ina (GIROMETTA, 1913, GIROMETTA, 1914, LANGHOFFER,
1915a, STROUHAL, 1939b, KARAMAN, 1966)
Bara~eva II (STROUHAL, 1939b)
Bara~eva Pe}ina (STROUHAL, 1939b)
Bara~eva Pe}ina I (STROUHAL, 1939b)
Bara~eva pe}ina II (LANGHOFFER, 1915a)
Bara}eva {pilja Mala (OZIMEC, 2003)
Bara}eva {pilja Velika (OZIMEC, 2003)
Bardarovica (FRANKENBERGER, 1938a, KARAMAN, 1966)
Bari}evi}a pe}ina (KARAMAN, 1966)
Barilovi} [pilja (STROUHAL, 1939b)
Bazdova~a (STROUHAL, 1939a, BEDEK et al., 2006)
Bazgova jama (BEDEK et al., 2006)
BB Nr. 6 (misprint, should be BB Nr. 661) (JAL@I] et al., 2010)
BB Nr. 16 (STROUHAL, 1939b)
BB Nr. 72 (STROUHAL, 1939a, STROUHAL, 1939c, STROUHAL, 1939e)
BB Nr. 91 (STROUHAL, 1939a, STROUHAL, 1939e)
BB Nr. 93 (STROUHAL, 1939a, STROUHAL, 1939e)
BB Nr. 99 (STROUHAL, 1939a, STROUHAL, 1939e)
BB Nr. 100 (STROUHAL, 1939a)
BB Nr. 101 (STROUHAL, 1939a, STROUHAL, 1939c)
BB Nr. 102 (STROUHAL, 1939a, BEDEK et al., 2006)
BB Nr. 105 (STROUHAL, 1939a)
BB Nr. 108 (STROUHAL, 1939a)
BB Nr. 109 (STROUHAL, 1939a)
BB Nr. 116 (STROUHAL, 1939a, STROUHAL, 1939e)
BB Nr. 146 (STROUHAL, 1939b)
BB Nr. 158 (STROUHAL, 1939a)
BB Nr. 324 (STROUHAL, 1939a)
BB Nr. 325 (STROUHAL, 1939a, STROUHAL, 1939e)
BB Nr. 326 (STROUHAL, 1939a, STROUHAL, 1939e)
BB Nr. 327 (STROUHAL, 1939a, STROUHAL, 1939e)
BB Nr. 329 (STROUHAL, 1939a)
BB Nr. 331 (STROUHAL, 1939a, STROUHAL, 1939b)
BB Nr. 333 (STROUHAL, 1939a)
BB Nr. 334 (STROUHAL, 1937, STROUHAL, 1939a, BEDEK et al., 2006)
BB Nr. 335 (STROUHAL, 1939a)
BB Nr. 345 (STROUHAL, 1939a)
BB Nr. 351 (STROUHAL, 1939b)
BB Nr. 410 (STROUHAL, 1939b)
BB Nr. 489 (STROUHAL, 1939d, ARCANGELI, 1940)
Present locality name
Baba {pilja
Bali}eva {pilja
Bali}eva {pilja
Gornja Bara}eva {pilja
Bara}eva {pilja
Donja Bara}eva {pilja
Gornja Bara}eva {pilja
Donja Bara}eva {pilja
Gornja Bara}eva {pilja
[pilja na Bardarovici
Bari~evi}a pe}ina
Vra`i}a {pilja
Bazgova~a jama
Bazgova~a jama
Movrica
Vilenska jama
[ipun
[ipun
\urovi}a {pilja
Studentski ratac
@ejava
Je{kalovica
Bazgova~a jama
Ledenica na podno`ju
Jabukovca
^injadra
Dobra jama
[ipun
Jezero
Osta{evica
Jama kod Rado{i}a
Jama na Visokoj
Jama na Bala~i
Vr{ine
Bali}eva {pilja
Smokvica
Trojama
Vranja~a
Jama na Ko{taku
Jama pod Kragljevcem
Nove jame
Veliki
Cerjanska {pilja
327
Nat. Croat. Vol. 20(2), 2011
BB Nr. 492 (STROUHAL, 1939b)
BB Nr. 494 (STROUHAL, 1939d, ARCANGELI, 1940)
BB Nr. 502 (STROUHAL, 1939d, ARCANGELI, 1940, BEDEK et al., 2006)
BB Nr. 503 (STROUHAL, 1939b)
BB Nr. 508 (STROUHAL, 1939b)
BB Nr. 511 (STROUHAL, 1939b)
BB Nr. 512 (STROUHAL, 1939b)
BB Nr. 520 (STROUHAL, 1939b)
BB Nr. 523 (STROUHAL, 1939b)
BB Nr. 526 (STROUHAL, 1939e, ARCANGELI, 1940)
BB Nr. 527 (STROUHAL, 1939b)
BB Nr. 529 (STROUHAL, 1939d)
BB Nr. 537 (STROUHAL, 1939b)
BB Nr. 538 (STROUHAL, 1939b)
BB Nr. 545 (STROUHAL, 1939b)
BB Nr. 547 (STROUHAL, 1939b)
BB Nr. 550 (STROUHAL, 1939b)
BB Nr. 551 (STROUHAL, 1939d, ARCANGELI, 1940, BEDEK et al., 2006)
BB Nr. 553 (STROUHAL, 1939b)
BB Nr. 556 (STROUHAL, 1939b)
BB Nr. 557 (STROUHAL, 1939b)
BB Nr. 562 (STROUHAL, 1939b)
BB Nr. 563 (STROUHAL, 1939b)
BB Nr. 566 (STROUHAL, 1939b)
BB Nr. 567 (STROUHAL, 1939b)
BB Nr. 569 (STROUHAL, 1939b)
BB Nr. 570 (STROUHAL, 1939b)
BB Nr. 573 (STROUHAL, 1939b)
BB Nr. 608 (STROUHAL, 1939a)
BB Nr. 642 (STROUHAL, 1939a)
BB Nr. 643 (STROUHAL, 1939a)
BB Nr. 644 (STROUHAL, 1939a)
BB Nr. 645 (STROUHAL, 1939a)
BB Nr. 646 (STROUHAL, 1939a)
BB Nr. 652 (STROUHAL, 1939a, STROUHAL, 1939c, STROUHAL, 1939e)
BB Nr. 653 (STROUHAL, 1939a, STROUHAL, 1939e)
BB Nr. 660 (STROUHAL, 1937, STROUHAL, 1939e, BEDEK et al., 2006)
BB Nr. 661 (STROUHAL, 1939a, STROUHAL, 1939c, STROUHAL, 1939e,
BEDEK et al., 2006)
BB Nr. 845 (STROUHAL, 1939a, STROUHAL, 1939c, STROUHAL, 1939e)
BB Nr. 849 (STROUHAL, 1939b)
BB Nr. 972 (STROUHAL, 1939a, STROUHAL, 1939c, STROUHAL, 1939e)
BB Nr. 1000 (STROUHAL, 1939a)
BB Nr. 1011 (STROUHAL, 1939a)
BB Nr. 1014 (STROUHAL, 1939a)
BB Nr. 1015 (STROUHAL, 1939a)
BB Nr. 1016 (STROUHAL, 1939a)
BB Nr. 1018 (STROUHAL, 1939a)
BB Nr. 1019 (STROUHAL, 1939a)
Bukovac
Vindija
Velika {pilja
Ceri~ka pe}ina
Munjave
Langhofferova {pilja
Ledeni~ka {pilja
Langhofferova {pilja
Zerngol {pilja
Zdenec pri Ciglaru
Pustinja
Zdenec pri Ciglaru
Vra`i}a {pilja
Vra`i}a {pilja
Ledena {pilja
Mla|enova pe}ina
Pustinja
Vla{ka pe}
Bribirska pe}ina
[upljara
Gornja Bara}eva {pilja
[upljara
Zvirnjak pe}ina
Donja Bara}eva {pilja
Ledena {pilja
Kova~evka pe}ina
Lokvarka
Velika pe}ina
Bali}eva {pilja
Bali}eva {pilja
Pe}ina kod Sinja
Zimska pe}ina
Kitlica jama
Crvenku{a-Tamnica-Suhi
Rumin
[ipun
Karle
Osta{evica
Movrica
[ipun
Mo~iljska {pilja
[ipun
Gospodska {pilja
Medi}a pe}ina
Rudeli}a {pilja
Rudeli}a {pilja
Bari~evi}a pe}ina
Crnomarkovi}a pe}ina
Zrilina pe}ina
328
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
BB Nr. 1023 (STROUHAL, 1939a)
BB Nr. 1024 (STROUHAL, 1939a)
BB Nr. 1032 (STROUHAL, 1939a, STROUHAL, 1939e)
BB Nr. 1034 (STROUHAL, 1939a)
BB Nr. 1038 (STROUHAL, 1939a)
BB Nr. 1042 (STROUHAL, 1939a)
BB Nr. 1043 (STROUHAL, 1939a)
Be`mek stan (BEDEK et al., 2006)
Bijele sige jama (BEDEK, 2007b)
Biospeologica balcanica loc. 557 (OZIMEC, 2003)
Biospeologica balcanica Lok. 549 (OZIMEC, 2003)
Biospeologica balcanica Lok. 554 (OZIMEC, 2003)
Biospeologica balcanica Lok. 566 (OZIMEC, 2003)
Bukova [pilja (STROUHAL, 1939b)
Caretova spilja (FRANKENBERGER, 1938b, ARCANGELI, 1939c)
cave at Mt. Ucka (GOTTSTEIN et al., 2001)
cave near village Murvica (GOTTSTEIN MATO^EC et al., 2002)
caverna presso il convento nelle vicinanze del villagio Murvica
(ARCANGELI, 1939b)
caverna presso Promontore (ARCANGELI, 1932, 1938, BRIAN, 1938, BEDEK
et al., 2006)
caverna presso Rasokatica nella montagna Glava (ARCANGELI, 1939c)
caverna sul monte Bugara (ARCANGELI, 1939c)
caves from Racice beside Castelnuovo (today Podgrad) (GIURGINCA, 2009)
Ceretova spilja (FRANKENBERGER, 1938a, KARAMAN, 1966)
Cicrusova jama (GIROMETTA, 1914)
Cinjadra (ARCANGELI, 1926, ARCANGELI, 1939b, BEDEK et al., 2006, JAL@I]
et al., 2010)
^injandra (KARAMAN, 1966, GOTTSTEIN MATO^EC et al., 2002)
^o~ina jama (BUTUROVI], 1955b, BUTUROVI], 1955c, KARAMAN, 1966)
Daljinhova jama (KARAMAN, 1966)
Devizza (BRIAN, 1938, STROUHAL, 1939a)
Dorfe Murvica (FRANKENBERGER & STROUHAL, 1940)
Durovi} jama (STROUHAL, 1939a, STROUHAL, 1939e, ARCANGELI, 1947)
D`amija (STROUHAL, 1939e)
\urovi} jama (KARAMAN, 1966)
Eishöhle bei Lokve (STILLER, 1911a)
environs du village de Nere`i{}e (BEDEK et al., 2006)
Golubinjka (GIROMETTA, 1914)
Golubinjka pod Lubljanom (LANGHOFFER, 1915a)
Golubinjka pod Ljubljanom (STROUHAL, 1939b)
Golubinjka pod malim Kraljevcom (GIROMETTA, 1914)
Golubnja~a kod @egara (BEDEK, 2007a)
Gorane Spiljas (ARCANGELI, 1940)
Gorane {pilja (STROUHAL, 1939d, KARAMAN, 1966, BEDEK et al., 2006)
Gospodska pe}ina (STROUHAL, 1939a, KARAMAN, 1966)
Grab~ina {pilja (KARAMAN, 1966)
Grab}ina spilja (FRANKENBERGER, 1938a)
Gr}ka pe}ina (FRANKENBERGER, 1938a, KARAMAN, 1966)
Grotta No. 1637 VG (BEDEK et al., 2006)
Kozjak pe}ina
Podbirnja~a {pilja
D`amije
Bobija jama
Pe}ina Stojakovi}
Vu~kovi} jama
Golubinka
Bazgova~a jama
Jama bijele sige
Gornja Bara}eva {pilja
Bara}eva {pilja
Bara}eva {pilja
Donja Bara}eva {pilja
Bukovac
Caretova {pilja
[pilja na U~ki
[pilja kod Murvice
[pilja kod Murvice
[pilja na Gradini kod
Premanture
Jakasova {pilja
[pilja na brdu Bugara
Ra~i{ka pe~ina
Caretova {pilja
Snije`nica pod Ljubljanom
^injadra
^injadra
^o}ina jama
Damljinhova jama
Denizza
[pilja kod Murvice
\urovi}a {pilja
D`amije
\urovi}a {pilja
Ledena {pilja
Bazgova~a jama
Snije`nica pod Ljubljanom
Snije`nica pod Ljubljanom
Snije`nica pod Ljubljanom
Jama pod Malim Kraljevcem
Golubnja~a
Velika {pilja
Velika {pilja
Gospodska {pilja
Grap~eva {pilja
Grap~eva {pilja
Gr~ka pe}ina
Biserujka
329
Nat. Croat. Vol. 20(2), 2011
Grotte bei Kotlu{a (FRANKENBERGER, 1939)
Grotte beim Dorfe O{tarija (JURINAC, 1888)
Grotte beim Dorfe Trschitsch (JURINAC, 1888)
grotte de l’île de Meleda (VANDEL, 1946)
Grotte de l’île Veglia (BEDEK et al., 2006)
Grotte près de la ville Trogir (BUTUROVI], 1955a, BEDEK et al., 2006)
Grotte près de la ville Trogir (BUTUROVI], 1955a)
Höhle am fusse des Dinara Gebirges (FRANKENBERGER, 1939)
Höhle am Monte Maggiore (WOLF, 1938)
Höhle am Mt. Maggiore (SCHMÖLZER, 1965a)
Höhle am Mte. Maggiore (FRANKENBERGER & STROUHAL, 1940)
Höhle am Ostabhang der Mosor planina (FRANKENBERGER, 1939
FLASAROVÁ, 1967)
Höhle an dem Monte Maggiore (KESSELYÁK, 1931, BEDEK et al., 2006,
JAL@I] et al., 2010)
Höhle an der Svilaja Planina (FRANKENBERGER, 1939)
Höhle an der Quelle des Cetina-Flusses (FRANKENBERGER, 1939)
Höhle auf dem Bugara-Berge (FRANKENBERGER, 1938b)
Höhle auf der Insel Veglia (Krk) (BEDEK et al., 2006)
Höhle auf der Svilaja planina (FRANKENBERGER, 1939)
Höhle auf der Svilaja Planina (FRANKENBERGER, 1940)
Höhle bei Bol (STROUHAL, 1939b)
Höhle bei Curzola (BEDEK et al., 2006)
Höhle bei dem Kloster in der Nähe des Dorfes Murvica
(FRANKENBERGER, 1938b)
Höhle bei der Bahnstation Josipdol (STILLER, 1914)
Höhle bei der Stadt Curzola (BEDEK et al., 2006)
Höhle bei Josipdol (STROUHAL, 1939b)
Höhle bei Petricevi (KARAMAN, 1966)
Höhle bei Pola (FRANKENBERGER & STROUHAL, 1940)
Höhle bei Promontore (STROUHAL, 1939a)
Höhle bei Pu`i (FRANKENBERGER, 1939)
Höhle bei Rasokatica (FRANKENBERGER, 1938b, FRANKENBERGER, 1940,
SCHMÖLZER, 1965b, BEDEK et al., 2006)
Höhle bei Sagorje (JURINAC, 1888)
Höhle bei Sinj (STROUHAL, 1939a)
Höhle bei Starigrad (STROUHAL, 1939b)
Höhle bei Suhi Rumin (ERHARD, 1997)
Höhle des Mosor-Gebirges (FRANKENBERGER & STROUHAL, 1940)
Höhle, die in der Nähe des Dorfes Kotlu{a (FRANKENBERGER, 1937)
Höhle im weichen Grund (STILLER, 1914, STROUHAL, 1939b)
Höhle in der Umgebung der Stadt Knin (FRANKENBERGER, 1937)
Höhle mit Wasser (FRANKENBERGER, 1937, BEDEK et al., 2006, JAL@I] et
al., 2010)
Höhle Namenlos (STILLER, 1914)
Höhle NO von Sinj (FRANKENBERGER, 1939)
Höhle NW. – Kroatiens (FRANKENBERGER & STROUHAL, 1940)
Höhle von Bivio Beru} (FRANKENBERGER, 1939)
Kotlu{a
Ambarac
Tr`i}ka {pilja
Movrica
Biserujka
[pilja Sv. Filipa i Jakoba
[pilja od Bilosoja
Kotlu{a
[pilja na U~ki
[pilja na U~ki
[pilja na U~ki
Pe}ina na isto~noj padini
Mosor planine
[pilja na U~ki
[pilja na Svilaji planini
Pe}ina na izvoru Cetine
[pilja na brdu Bugara
Biserujka
Pe}ina na Svilaji planini
[pilja na Svilaji planini
Jama kod Vidove gore
Pi{urka
[pilja kod Murvice
[pilja u Mekoti
Pi{urka
[pilja u Mekoti
^ampari
[pilja na Gradini kod
Premanture
[pilja na Gradini kod
Premanture
Pe}ina kod Pu`i
Jakasova {pilja
Zagorska pe}
Pe}ina kod Sinja
[pilja kod Starigrada
Suvi Rumin
Vranja~a
Kotlu{a
[pilja u Mekoti
Kotlu{a
Kotlu{a
[pilja u Mekoti
Pe}ina sjeveroisto~no od Sinja
Vla{ka pe}
Pe}ina Bivio Beru}a
330
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
Höhle von Marija Ru`i} (FRANKENBERGER, 1939)
Höhle von Ozalj (WOLF, 1938)
Höhlen auf der Dinara planina, hauptsächlich ihren westlichen
Abhängen; Lok.Nr. 254 (FRANKENBERGER, 1939)
Höhlen auf der Dinara planina, hauptsächlich ihren westlichen
Abhängen; Lok.Nr. 256 (FRANKENBERGER, 1939)
Höhlen auf der Dinara planina, hauptsächlich ihren westlichen
Abhängen; Lok.Nr. 258 (FRANKENBERGER, 1939)
Höhlen auf der Dinara planina, hauptsächlich ihren westlichen
Abhängen; Lok.Nr. 260 (FRANKENBERGER, 1939)
Höhlen bei Bol … auf das Plateau nach Neresi zu führen … (WETTSTEIN,
1914)
Höhlen bei Milna … auf dem südlichen Abhange des zwischen den
Namen »Lokvice« und »Blisina glavica« (WETTSTEIN, 1914)
Höhlen bei Milna … die links vom Wege nach S. Guiseppe liegt
(WETTSTEIN, 1914)
Höhlen von Ivan Stani} und von Franjo Stani} (FRANKENBERGER, 1939)
Höhleneingang … in Nordistrien bei Ra}i~e (BEDEK et al., 2006)
Jaice (GIURGINCA, 2009)
Jama na Bala}i (KARAMAN, 1966)
Jama pod Ko{takom (STROUHAL, 1939a, KARAMAN, 1966)
Jama pod malim Kraljevcom (GIROMETTA, 1914)
Jama za Mahrinacom (FRANKENBERGER, 1938a, STROUHAL, 1939a)
Jama za Mahrincom (KARAMAN, 1966)
Je{kalovi}a jama (KARAMAN, 1966)
Jezerina Jama (STROUHAL, 1939b)
Josipdol Ponor (STROUHAL, 1939b)
Kapacina (KARAMAN, 1966)
Karla Gru` (STROUHAL, 1939a, STROUHAL, 1939e, ARCANGELI, 1947,
KARAMAN, 1966)
Kerkawasserfällen (VERHOEFF, 1930)
kleine Höhle bei Tonnj (STILLER, 1911b)
kleine Höhle bei Tounj (STROUHAL, 1939b)
Koli{tini (GIROMETTA, 1913)
Kopacina (FRANKENBERGER, 1938a, STROUHAL, 1939a)
Kötlenicehöhle (BEDEK et al., 2006)
Kotlu{ka pe}ina (BEDEK et al., 2006, JAL@I] et al., 2010)
Kraljeva pe}ina (GIROMETTA, 1914, STROUHAL, 1939a, KARAMAN, 1966)
Krasulja (BEDEK et al., 2006)
Ledena Pe}ina (STROUHAL, 1939b)
Ledenica (LANGHOFFER, 1915a, STROUHAL, 1939b)
Ledenica (GIROMETTA, 1914, LANGHOFFER, 1915a)
Ledenica am Mosor (STROUHAL, 1939a)
Ledenica na Mosoru (KARAMAN, 1966)
Lok. Nr. 250 (FRANKENBERGER, 1939)
Lok. Nr. 253 (FRANKENBERGER, 1939)
Lok. Nr. 263 (FRANKENBERGER, 1939)
Pe}ina Marije Ru`i}
Ozaljska {pilja
[pilja na Dinari 254
[pilja na Dinari 256
[pilja na Dinari 258
[pilja na Dinari 260
Jama kod Vidove gore
[pilja izme|u Lokvica i
Blisine glavice
[pilja izme|u Milne i Sv.
Josipa
Pe}ina Ivana Stani}a i Franja
Stani}
Ra~i{ka pe~ina
Jajce
Jama na Bala~i
Jama na Ko{taku
Jama pod Malim Kraljevcem
Jama za Mahrincem
Jama za Mahrincem
Je{kalovica
Jezero
Munjave
Kopa~ina
Karle
Krka
Guge~ka pe}ina
Guge~ka pe}ina
Koli{tina
Kopa~ina
Vranja~a
Kotlu{a
Bali}eva {pilja
Biserujka
Ledeni~ka {pilja
Ledena {pilja
Ledenica na podno`ju
Jabukovca
Ledenica na podno`ju
Jabukovca
Ledenica na podno`ju
Jabukovca
Pe}ina na izvoru Cetine
Pe}ina na Svilaji planini
Pe}ina na isto~noj padini
Mosor planine
331
Nat. Croat. Vol. 20(2), 2011
Lok. Nr. 265 (FRANKENBERGER, 1939)
Lokve {pilja (STROUHAL, 1939a, STROUHAL, 1939b, KARAMAN, 1966)
Lukina jama (KEROVEC et al., 1999)
Ma~kova {pilja (BEDEK et al., 2006)
Mali ora (FRANKENBERGER, 1939)
Markova spilja (FRANKENBERGER, 1938a, BEDEK et al., 2006)
Medina pe}ina (LANGHOFFER, 1912, LANGHOFFER, 1915b, CSIKI, 1926,
STROUHAL, 1939b)
Medvednica gornja {pilja (BEDEK, 2007b)
Medvjedica (STROUHAL, 1939b)
Mo~ilje cave (OZIMEC, 2005)
Mo~ilje pe}ina (STROUHAL, 1939b)
Monte Maggiore (ARCANGELI, 1939a; GRUNER & TABACARU, 1963;
GIURGINCA, 2009)
Morica pe}ina (KARAMAN, 1966)
Mosorgebirge, in einer Höhle (SCHMÖLZER, 1965a)
Movricahöhle (BEDEK et al., 2006, JAL@I] et al., 2010)
Mra~na pe}ina (REDEN[EK, 1958)
Mra~na pe}ina (STROUHAL, 1939a, KARAMAN, 1966)
Mre{nica ponor (STROUHAL, 1939b)
Murcica (GOTTSTEIN et al., 2001)
Narenta-Grotten (KARAMAN, 1966)
Narenta Grotten (STROUHAL, 1938, FRANKENBERGER & STROUHAL, 1940,
BUTUROVI], 1955c)
Omi{aljska spilja Bisernica (BEDEK et al., 2006)
Ostasevica (GOTTSTEIN et al., 2001)
Ota{evica (KARAMAN, 1966)
Ote{ (LANGHOFFER, 1912)
Ozaljska pe}ina (CSIKI, 1926)
Paganetijeva pe}ina (KARAMAN, 1966)
Paganetijeva {pilja (KARAMAN, 1966, BEDEK et al., 2006)
Paganetti (VERHOEFF, 1931a, VERHOEFF, 1931b, FRANKENBERGER, 1938a,
STROUHAL, 1938, STROUHAL, 1939a, FRANKENBERGER & STROUHAL, 1940,
ARCANGELI, 1947, ARGANO et al., 1982, SCHMIDT & LEISTIKOW, 2004,
BEDEK et al., 2006)
Paganettijeva pe}ina (FRANKENBERGER, 1938a, BEDEK et al., 2006)
Pe}ina iznad Opatije (KARAMAN, 1966)
Pe}ina iznad Opatije (KARAMAN, 1966)
pe}ina kod Murcice (KARAMAN, 1966)
Pe}ina kod Nebljusa (BUTUROVI], 1963)
Pe}ina kod Ozlja (LANGHOFFER, 1912)
pe}ina kod Petri}evi (KARAMAN, 1966)
pe}ina kod Pule (KARAMAN, 1966)
pe}ina kod Rosokatice na brdu Glava (KARAMAN, 1966)
pe}ina kod s. Kotlu{a (KARAMAN, 1966)
pe}ina kod samostana u samom Trogiru (HAD@I, 1933)
pe}ina kod Tr`i}a (LANGHOFFER, 1912)
pe}ina kod Zagorja (LANGHOFFER, 1912)
Pe}ina sjeveroisto~no od Sinja
Lokvarka
Lukina jama – Trojama
Velika {pilja
[krabuljica
Markova {pilja
Medina {pilja
Medvednica
\ulin ponor – Medvedica
Mo~iljska {pilja
Mo~iljska {pilja
U~ka
Movrica
Vranja~a
Movrica
Mra~na {pilja
Crvenku{a-Tamnica-Suhi
Rumin
Ambarac
[pilja kod Murvice
[pilja u dolini Neretve
[pilja u dolini Neretve
Biserujka
Osta{evica
Osta{evica
Ote{ka pe}ina
Ozaljska {pilja
Pi{urka
Pi{urka
Pi{urka
Pi{urka
Bla`eva jama
Pe}ina Ivana Stani}a i Franja
Stani}
[pilja kod Murvice
Kr~evina
Ozaljska {pilja
^ampari
[pilja na Gradini kod
Premanture
Jakasova {pilja
Kotlu{a
[pilja od Bilosoja
Tr`i}ka {pilja
Zagorska pe}
332
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
pe}ina na sjeveroisto~noj padini Biokova (KARAMAN, 1966)
pe}ina na Sridnjoj Gori (KARAMAN, 1966)
pe}ina na Svilaji planini (KARAMAN, 1966, BEDEK et al., 2006)
pe}ina na U~ki (KARAMAN, 1966)
pe}ina: Sv. Filipa i Jakoba (HAD@I, 1933)
Piccola caverna sul Monte Maggiore (ARCANGELI, 1938)
Pisuska (STROUHAL, 1938)
pit near Pula (GOTTSTEIN et al., 2001)
Pit 3 near Banina Stream (KARAMAN et al., 2009, JAL@I] et al., 2010)
podzemni hod (JURINAC, 1886)
Ponor Mre`nice (JURINAC, 1886)
Porta Madonna (WOLF, 1938)
Promontore (WOLF, 1938, FRANKENBERGER & STROUHAL, 1940,
KARAMAN, 1966)
Pusi oberhalb Abbazia (FRANKENBERGER & STROUHAL, 1940)
Rasokatica im Gebirge Glava (FRANKENBERGER & STROUHAL, 1940)
Razgova~a {pilja (KARAMAN, 1966)
Razgova}a spilja (FRANKENBERGER, 1938a)
Razgova}a {pilja (BEDEK et al., 2006)
Repnjak (LANGHOFFER, 1915a, STROUHAL, 1939d, ARCANGELI, 1940,
KARAMAN, 1966)
Ro~ka {pilja (OZIMEC et al., 2011b)
Rudeli}a pe}ina (KARAMAN, 1966)
Rudeli}a Pe}ina N. I (STROUHAL, 1939a)
Rudeli}a Pe}ina N. II (STROUHAL, 1939a)
Sinjandra (GOTTSTEIN et al., 2001)
Sipun Aeskulapgrotte (VANDEL, 1955)
Sitvuki barlang (LANGHOFFER, 1915b)
Sitvuki pe}ina (CSIKI, 1926, STROUHAL, 1939b)
Skocmen Spiljas (ARCANGELI, 1940)
Sko~men (LANDHOFFER, 1915a; KARAMAN, 1966)
Smrde}a pe}ina (REDEN[EK, 1958)
Spela (KARAMAN, 1966)
Spila (BUTUROVI], 1955c)
Spila Gjurgjevica (OZIMEC, 2004)
Spilja Bezimena (STILLER, 1914)
Spilja kod Rasokatice (FRANKENBERGER, 1938a)
Spilja na Smokovniku (UBIC & OZIMEC, 2005)
Spilja pod Humom (FRANKENBERGER, 1938a)
Spilja pod Vrhom (FRANKENBERGER, 1938a, STROUHAL, 1939a)
Spilja pri Nerznom Dolu (FRANKENBERGER, 1938a, STROUHAL, 1939a,
ARCANGELI, 1947)
Spilja pri Smokovniku (UBIC & OZIMEC, 2005)
Spisur}a jama (BEDEK et al., 2006)
Spisurska spilja (BEDEK et al., 2006)
Sve~ara (STROUHAL, 1939b)
Svilanjska jama (BEDEK et al., 2006)
[pilja Bezimena bei Ko{are (STROUHAL, 1939b)
Pe}ina na sjeveroisto~nim
padinama Biokova
Pe}ina u Sridnjoj gori
[pilja na Svilaji planini
[pilja na U~ki
[pilja Sv. Filipa i Jakoba
[pilja na U~ki
Pi{urka
[pilja na Gradini kod
Premanture
Jama kod potoka Banine 3
Ambarac
Ambarac
Porto Madonna
[pilja na Gradini kod
Premanture
Pe}ina kod Pu`i
Jakasova {pilja
Bazgova~a jama
Bazgova~a jama
Bazgova~a jama
Cerjanska {pilja
Rabakova {pilja
Rudeli}a {pilja
Rudeli}a {pilja
Rudeli}a {pilja
^injadra
[ipun
Sitvukova pe}ina
Sitvukova pe}ina
Zdenec pri Ciglaru
Zdenec pri Ciglaru
Crna pe}ina
Spila 1
Spila 1
Samograd
[pilja u Mekoti
Jakasova {pilja
[pilja pod Kapelu
[pilja pod Humom
[pilja pod Vrhom
[pilja kod Nerezinog dola
[pilja pod Kapelu
Pi{urka
Pi{urka
Zve~ara
Biserujka
[pilja u Mekoti
333
Nat. Croat. Vol. 20(2), 2011
[pilja kod O{tarije ogulinske (JURINAC, 1886)
[pilja kod Premanture (BEDEK et al., 2006)
[pilja kod Ra~ica (BEDEK et al., 2006)
[pilja kod Rasohatice (KARAMAN, 1966, BEDEK et al., 2006)
[pilja kod Trogira (KARAMAN, 1966, BEDEK et al., 2006)
[pilja kod Trogira (KARAMAN, 1966)
[pilja kod V. Rasoke (BEDEK et al., 2006)
{pilja Mo~ilje (OZIMEC, 2005)
[pilja na izvoru Mre`nice (JURINAC, 1886)
{pilja pri Nereznom dolu (KARAMAN, 1966)
Tabaina spilja (FRANKENBERGER, 1938a, FRANKENBERGER, 1938b,
ARCANGELI, 1939c, FRANKENBERGER & STROUHAL, 1940, KARAMAN,
1966, BEDEK et al., 2006)
tamnik (JURINAC, 1886)
Trimbu{ova pe}ina (REDEN[EK, 1958)
Tr`i} pe}ina (CSIKI, E., 1926, STROUHAL, 1939b)
Tr`i}i barlang (LANGHOFFER, 1912)
unnamed small cave at Mt. Maggiore (SCHMIDT & LEISTIKOW, 2004)
Velika pe} (BUTUROVI], 1955c, KARAMAN, 1966)
Velika pe}ina (BEDEK et al., 2006, JAL@I] et al., 2010)
Velika pe}ina (BEDEK et al., 2006)
Velika pe}ina kod Velikih Goranaca (LANGHOFFER, 1915a)
Velika Spilja (FRANKENBERGER, 1938a, FRANKENBERGER, 1938b,
STROUHAL, 1939a, ARCANGELI, 1947)
Velika spilja (ARCANGELI, 1939c)
Vile Jezerkinje (REDEN[EK, 1958)
Vilina pe}ina (BUTUROVI], 1955b, KARAMAN, 1966)
Vindia Spiljas (ARCANGELI, 1940)
Vindia {pilja (STROUHAL, 1939d, KARAMAN, 1966)
Viteziceva pecina (GOTTSTEIN et al., 2001)
Vitezi~eva pe~ina (POTO^NIK, 1983, BEDEK et al., 2006)
Vitezi~ova jama (BEDEK et al., 2006)
Vitezi}a {pilja (BEDEK et al., 2006)
Vitezi}eva pe}ina (GOTTSTEIN MATO^EC et al., 2002)
Vlaska pecina (ARCANGELI, 1940, GOTTSTEIN et al., 2001)
Vlaska Pe~ina bei Zengg zwischen Kozica und Smokvica (BEDEK et al.,
2006)
Vla{ka pe}ina (STROUHAL, 1939d, KARAMAN, 1966, VUJ^I]-KARLO &
RA\A, 2004, BEDEK et al., 2006)
Vla{ka {pilja (BEDEK et al., 2006)
Vodena jama (JURINAC, 1886, LANGHOFFER, 1912, LANGHOFFER, 1915b,
CSIKI, 1926, STROUHAL, 1939b)
Vra`i}-pe}ina (LANGHOFFER, 1915a)
Vrlovkahöhle (STROUHAL, 1939b)
Zagorje pe}ina (CSIKI, 1926, STROUHAL, 1939b)
Zagorska pe} (VUJ^I]-KARLO & RA\A, 2004)
Zala (JAL@I] & BO@I^EVI], 1975)
Zejava (STROUHAL, 1939a, KARAMAN, 1966)
Zrinski rudnici (BEDEK, 2007b)
Ambarac
[pilja na Gradini kod
Premanture
Ra~i{ka pe~ina
Jakasova {pilja
[pilja Sv. Filipa i Jakoba
[pilja od Bilosoja
Jakasova {pilja
Mo~iljska {pilja
Zagorska pe}
[pilja kod Nerezinog dola
Tabaina {pilja
Ambarac
Crna pe}ina
Tr`i}ka {pilja
Tr`i}ka {pilja
[pilja na U~ki
Velika {pilja
Kotlu{a
Velika {pilja
Velika {pilja
Velika {pilja
Velika {pilja
Crna pe}ina
Vilina {pilja – Ombla
Vindija
Vindija
Biserujka
Biserujka
Biserujka
Biserujka
Biserujka
Vla{ka pe}
Vla{ka pe}
Vla{ka pe}
Vla{ka pe}
Ambarac
Vra`i}a {pilja
Vrlovka
Zagorska pe}
Vla{ka pe}
Mika{inovi}a {pilja
@ejava
Rudnik Zrinski
334
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
APPENDIX 2.
Tab. 44. Collection records
Croatian Biospeleological Society (CBSS) collection
CBSS collection 1
M. Luki}, 05.12.2008.
CBSS collection 2
J. Bedek, 24.01.2004.
CBSS collection 3
H. Biland`ija, 04.12.2004.
R. Ozimec, 04.12.1994.; 29.10.2000.; J. Bedek, I. Bor{i}, R. Ozimec,
14.07.2001.; I. Bor{i}, 07.10.2001.; M. Luki}, 23.02.2003.; R. Ozimec,
26.02.2003.; M. Luki}, 21.04.2003.; 15.06.2003.; 07.11.2003.; J. Bedek, M.
CBSS collection 4
Luki}, 12.11.2003.; M. Luki}, R. Ozimec, 28.12.2003.; M. Pavlek,
13.05.2004.; J. Bedek, 04.12.2004.; M. Luki}, 22.12.2004.; J. Bedek,
16.01.2005.; H. Biland`ija, 26.02.2005.; R. Ozimec, 04.11.2006.
CBSS collection 5
J. Bedek, 16.04.2005.
CBSS collection 6
J. Bedek, 22.05.2005.
CBSS collection 7
H. Biland`ija, 14.06.2004.
CBSS collection 8
H. Biland`ija, 20.03.2005.
CBSS collection 9
R. Ozimec, 11.05.2008.
CBSS collection 10
R. Ozimec, P. Rade, 27.02.2009.
CBSS collection 11
J. Bedek, 18.07.2004.
CBSS collection 12
H. Cvitanovi}, 20.04.2006.
CBSS collection 13
P. Rade, 20.04.2006.
CBSS collection 14
M. Pavlek, 23.04.2006.
CBSS collection 15
M. Luki}, 27.10.2008.
CBSS collection 16
M. Luki}, 25.10.2008.
CBSS collection 17
M. Pavlek, 21.04.2006.; R. Ozimec, H. Cvitanovi}, 25.10.2008.
CBSS collection 18
M. Luki}, 23.09.2010.
CBSS collection 19
A. Komeri~ki, 22.09.2010.
CBSS collection 20
L. Katu{i}, 13.05.2006.
R. Ozimec, M. Pavlek, 27.05.2007.; J. Bedek, 0.11.2007.; F. Kljakovi}
CBSS collection 21
Ga{pi}, 13.05.2008.
CBSS collection 22
R. Ozimec, 27.05.2007.
CBSS collection 23
F. Kljakovi} Ga{pi}, R. Ozimec, P. Rade, 28.05.2007.
R. Ozimec, 16.05.2006., 13.10.2006.; H. Cvitanovi}, R. Ozimec, 16.10.2006.;
CBSS collection 24
R. Ozimec, 04.11.2007.
H. Biland`ija, L. Katu{i}, R. Ozimec, M. Pavlek, 14.05.2006.; P. Bregovi},
CBSS collection 25
L. Katu{i}, A. Kolari}, P. Rade, 15.10.2006.
CBSS collection 26
H. Cvitanovi}, 03.05.2006.; H. Cvitanovi}, R. Ozimec, 14.10.2006.
CBSS collection 27
R. Ozimec, 16.11.2007.
R. Ozimec, 12.05.2008.; H. Cvitanovi}, E. Domina, M. Luki}, R. Ozimec,
CBSS collection 28
17.10.2008.; A. Komeri~ki, 20.05.2009.
CBSS collection 29
P. Rade, 14.10.2008.
R. Ozimec, 29.08.1993.; J. Bedek, 10.02.2002.; H. Biland`ija, R. Ozimec, M.
CBSS collection 30
Pavlek, 15.05.2006.; R. Ozimec, M. Pavlek, 27.05.2007.; H. Cvitanovi},
02.11.2007.; E. Domina, 16.10.2008.
CBSS collection 31
R. Ozimec, P Rade, 15.10.2006.
CBSS collection 32
H. Biland`ija, 14.05.2006.
CBSS collection 33
H. Cvitanovi}, P. Rade, 14.10.2006.
Nat. Croat. Vol. 20(2), 2011
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
34
35
36
37
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
CBSS collection 56
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
57
58
59
60
61
62
63
CBSS collection 64
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
65
66
67
68
69
70
71
72
73
74
75
76
77
R. Ozimec, 03.11.2007.
P. Bregovi}, R. Ozimec, 04.11.2007.
R. Ozimec, M. Pavlek, 17.05.2006.
H. Cvitanovi}, R. Ozimec, 26.05.2007.; E. Domina, P. Rade, 18.10.2008.
R. Ozimec, 25.05.2007.
H. Cvitanovi}, R. Ozimec, 25.05.2007.
H. Cvitanovi}, 17.10.2008.; R. Ozimec, P. Rade, 18.05.2009.
P. Bregovi}, 17.10.2009.
M. Luki}, 27.04.2009.
J. Bedek, P. Bregovi}, 29.04.2009.
H. Cvitanovi}, 28.04.2009.
A. Komeri~ki, 29.04.2009.
J. Bedek, 05.03.2011.
J. Bedek, 24.04.2009.
M. Luki}, 24.06.2009.
J. Bedek, 12.07.2005.
R. Ozimec, 08.09.2009.
J. Bedek, 06.12.2008.; J. Bedek, A. Kirin, 10.05.2009.
H. Biland`ija, 15.06.2008.
A. Kirin, 05.12.2009.
M. Pavlek, 22.06.2008.
J. Bedek, H. Cvitanovi}, 20.06.2008.; J. Bedek, 12.05.2009.
H. Cvitanovi}, R. Ozimec, 17.02.1999.; R. Ozimec, 27.02.2000.; J. Bedek,
16.02.2003.; H. Cvitanovi}, 18.06.2008.; H. Cvitanovi}, 08.12.2008.; M.
Luki}, 14.05.2009.
H. Cvitanovi}, K. Miculini}, 22.06.2008.
A. Kirin, 14.05.2009.
J. Bedek, 20.06.2008.; K. Miculini}, 09.05.2009.
J. Bedek, 09.05.2009.; A. Kirin, M. Luki}, 13.05.2009.
R. Ozimec, 19.07.1998.
T. Rubini}, 29.11.2003.
J. Bedek, 30.11.2002.
T. Dra`ina, M. Pavlek, 24.08.2008.; R. Ozimec, 26.08.2008.; M. Luki},
15.05.2009.; M. Pavlek, 24.05.2009.
H. Biland`ija, 14.06.2008.; M. Luki}, 05.12.2008.
R. Ozimec, T. Rubini}, 03.11.2002.
R. Ozimec, 08.02.2009.
R. Ozimec, 26.02.2009.; K. Miculini}, 06.06.2009.; J. Bedek, 17.09.2009.
I. Ra{i}, 07.12.2002.
T. Rubini}, 29.09.2002.
R. Ozimec, 21.12.2003.
M. Pavlek, 04.12.2004.
R. Ozimec, 13.04.2008.
J. Bedek, M. Pavlek, 07.11.2006.
D. Hmura, 16.09.2007.
M. Pavlek, 25.07.2006.
M. Pavlek, 25.07.2006.
335
336
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
CBSS collection 95
CBSS collection
CBSS collection
CBSS collection
CBSS collection
96
97
98
99
CBSS collection 100
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
101
102
103
104
105
106
107
108
109
110
111
112
113
114
CBSS collection 115
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
CBSS collection
116
117
118
119
120
121
D. Hmura, 17.09.2007.
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K. Miculini}, 14.05.2009.
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H. Cvitanovi}, 10.2000.
H. Biland`ija, 15.06.2008., 23.08.2008.; A. Kirin, 11.05.2009.; A. Komeri~ki,
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H. Cvitanovi}, 18.05.1997.; R. Ozimec, 11.11.1998.
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R. Ozimec, 07.06.2003.
R. Ozimec, I. Ra{i}, 09.06.2003.
T. Rubini}, 29.11.2003.
R. Ozimec, 26.04.2003.
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R. Ozimec, T. Rubini}, 20.09.2002.; M. Luki}, 22.09.2002.
K. Moto~i}, 09.06.2003.
T. Rubini}, 24.02.2002.; R. Ozimec, T. Rubini}, 20.09.2002.; T. Rubini},
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J. Bedek, 22.09.2002.
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K. Miculini}, 24.06.2008.; A. Komeri~ki, 15.05.2009.
M. Luki}, K. Miculini}, 14.06.2008.
R. Ozimec, T. Rubini}, 03.11.2002.
T. Rubini}, 07.09.2003.
T. Rubini}, 18.01.1998.; R. Ozimec, T. Rubini}, 15.09.2002.
H. Cvitanovi}, 17.09.2009.
J. Bedek,19.03.2009.
J. Bedek, 03.11.2002.
T. Rubini}, 02.02.2002.
T. Rubini}, 15.11.1997.; R. Ozimec, 19.04.1998.; J. Bedek, R. Ozimec, T.
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R. Ozimec, 02.02.1997.; J. Bedek, R. Ozimec, T. Rubini}, 02.11.2002.; J.
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H. Cvitanovi}, 27.04.1996.; R. Ozimec, 29.01.1998.; H. Cvitanovi},
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M. Luki}, 07.11.2006.
R. Ozimec, 14.09.2007.
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J. Bedek, P. Bregovi}, M. Pavlek, 23.04.2009.
J. Bedek, M. Luki}, 08.11.2006.
A. Kolari}, 26.07.2006.
J. Bedek, M. Luki}, 18.09.2007.
J. Bedek, 05.11.2006.
H. Biland`ija, P. Bregovi}, 25.06.2009.
M. Luki}, 25.06.2009.
J. Bedek, A. Kirin, 03.06.2010.
M. Luki}, 25.04.2009.
K. Miculini}, 01.06.2010.
J. Bedek, P. Bregovi}, 26.06.2009.
J. Bedek, M. Luki}, K. Miculini}, 04.06.2010.
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H. Biland`ija, D. Hmura, M. Pavlek, 18.09.2007.
H. Biland`ija, , 27.06.2009.
H. Cvitanovi}, M. Luki}, M. Pavlek, 06.11.2006.
H. Biland`ija, M. Luki}, 26.06.2009.
J. Bedek, H. Biland`ija, P. Bregovi}, 27.06.2009.
J. Bedek, 26.06.2009.
P. Bregovi}, I. Jaklinovi}, A. Kolari}, M. Pavlek, N. Ragu`, 24.07.2006.; A.
Kirin, A. Komeri~ki, 04.06.2010.
J. Bedek, P. Bregovi}, 24.06.2009.
H. Biland`ija, B. Jal`i}, M. Pavlek, 04.11.2006.; H. Biland`ija, 17.09.2007.
J. Bedek, 04.11.2006.
H. Biland`ija, P. Bregovi}, 28.06.2009.
H. Biland`ija, H. Cvitanovi}, M. Pavlek, 04.10.2006.
J. Bedek, M. Luki}, 28.06.2009.
M. Pavlek, 26.04.2009.
J. Bedek, 26.04.2009.; A. Kirin, A. Komeri~ki, 04.06.2010.
P. Bregovi}, A. Komeri~ki, 21.04.2009.
P. Bregovi}, 24.04.2009.
R. Ozimec, 03.07.1994., 09.04.1994., 28.09.1996.; J. Bedek, R. Ozimec,
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M. Luki}, 02.05.2004.
R. Ozimec, 01.06.1997.
S. Minihofer, 24.07.2004.
R. Ozimec, 03.08.2003.
R. Ozimec, 02.08.2003.
R. Ozimec, 31.08.1996.
J. Bedek, 12.07.2005.
G. Poli}, 18.05.1997.
J. Bedek, 09.12.2000.
M. Luki}, 31.01.2008.
J. Bedek, 07.09.2009.
R. Ozimec, 08.09.2009.; A. Komeri~ki, 09.09.2009
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J. Bedek, 08.09.2009.
A. Kirin, 17.05.2009.; M. Luki}, 11.09.2009.
M. Luki}, 09.09.2009.
R. Ozimec, 17.06.2006.
H. Cvitanovi}, 29.11.2003.; J. Bedek, M. Pavlek, 01.02.2004.
R. Ozimec, 28.01.1995.
J. Bedek, 01.02.2008.; H. Biland`ija, B. Jal`i}, 18.06.2008.
R. Ozimec, 17.06.2008.
H. Cvitanovi}, 14.04.1997., 21.06.1997.; J. Bedek, 31.01.2004.
M. Luki}, 26.03.2008.
H. Cvitanovi}, 26.03.2008., 31.10.2008.
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500
501
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J. Bedek, D. @upan, 01.11.2008.
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R. Ozimec, 25.06.2009.
M. Luki}, 03.11.2008.
J. Bedek, 26.03.2008.
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J. Bedek, R. Ozimec, 31.10.2008.
J. Bedek, 31.10.2008.; J. Bedek, M. Luki}, 04.11.2008.
R. Ozimec, 02.11.2008.
J. Bedek, 30.03.2008., 02.11.2008.
J. Bedek, H. Cvitanovi}, M. Luki}, 28.03.2008.
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J. Bedek, K. Miculini}, D. @upan, 01.11.2008.
H. Cvitanovi}, 27.03.2008.
A. Komeri~ki, 06.10.2009.
H. Cvitanovi}, 21.06.2009.
K. Miculini}, 29.03.2008.
R. Ozimec, 29.03.2008.; M. Luki}, R. Ozimec, 01.11.2008.; R. Ozimec,
22.06.2009.
J. Bedek, H. Cvitanovi}, M. Luki}, 31.03.2008.
J. Bedek, 01.11.2008.
M. Pavlek, 09.04.2007.
M. Pavlek, 11.04.2007.
P. Bregovi}, M. Luki}, 03.05.2008.
M. Luki}, 03.05.2008.
J. Bedek, 03.05.2008.
J. Bedek, H. Biland`ija, M. Pavlek, 08.04.2007.; M. Luki}, 28.04.2008.
H. Biland`ija, 28.04.2008.
J. Bedek, I. Bor{i}, T. Dra`ina, 19.06.2001.; M. Pavlek, 07.04.2007.; J.
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J. Bedek, 10.04.2007., 14.04.2007.; H. Biland`ija, 30.05.2007.; J. Bedek, H.
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H. Biland`ija, 30.04.2008.
R. Ozimec, 07.-08.10.2009.
R. Ozimec, 07.10.2009.
J. Bedek, H. Cvitanovi}, 27.03.2008.
R. Ozimec, 31.03.2008.
R. Ozimec, 27.08.1998.
J. Bedek, 21.03.2004.
R. Ozimec, 25.05.2004.; J. Bedek, 02.04.2006.; M. Luki}, 22.06.2007.; J.
Bedek, M. Luki}, 19.08.2007.
R. Ozimec, 13.11.2010.
R. Ozimec, 20.08.2005.
J. Bedek, H. Biland`ija, 24.04.2004.
R. Ozimec, 27.04.2004.
R. Ozimec, M. Pavlek, 26.04.2004.; H. Biland`ija, M. Pavlek, 30.09.2007.
H. Cvitanovi}, M. Luki}, 03.04.2006.; J. Bedek, H. Cvitanovi}, 27.10.2006.
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521
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523
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529
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532
533
534
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538
539
540
541
S. Gottstein, R. Ozimec, 03.06.2000.
M. Pavlek, 11.04.2007.; J. Bedek, H. Biland`ija, 26.04.2008.
J. Bedek, 10.04.2007.; H. Biland`ija, 30.05.2007.; J. Bedek, H. Biland`ija, P.
Bregovi}, 02.05.2008.
J. Bedek, R. Ozimec, 06.01.2005.; J. Bedek, A. Kirin, A. Komeri~ki, P.
Kova~ Konrad, 04.05.2010.; A. Kirin, 08.10.2010.
A. Kirin, 04.05.2010.
A. Kirin, 10.10.2010.
A. Kirin, 09.10.2010.
H. Biland`ija, R. Ozimec, M. Pavlek, 07.01.2005.; J. Bedek, M. Luki},
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M. Luki}, 03.05.2010.
J. Bedek, 26.03.2008.
M. Luki}, 02.11.2008.
B. Jal`i}, S. Gottstein Mato~ec, R. Ozimec, 21.06.2001.; J. Bedek, K.
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M. Pavlek, 26.04.2004.
R. Ozimec, 04.11.2008.; A. Komeri~ki, K. Miculini}, 23.06.2009.
H. Cvitanovi}, 07.10.2004.
R. Ozimec, 28.06.2003., 31.10.2003., 27.05.2004.; J. Bedek, M. Luki},
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M. Luki}, 22.01.2011.
J. Bedek, 06.10.2005.
A. Kirin, K. Miculini}, 02.06.2010.
M. Pavlek, 04.11.2006.; J. Bedek, 17.11.2007., 05.04.2008.
J. Bedek, 24.4.2009.
P. Bregovi}, M. Pavlek, 17.10.2009.
D. Hmura, 18.10.2009.
H. Cvitanovi}, A. Komeri~ki, M. Luki}, 29.04.2009.
J. Bedek, A. Kirin, P. Kova~ Konrad, M. Luki}, 08.05.2010.
R. Ozimec, 27.08.2005.; J. Bedek, M. Pavlek, 27.10.2006.
R. Ozimec, 28.04.2002.
H. Cvitanovi}, M. Pavlek, 19.11.2005.; R. Ozimec, 04.04.2006.
R. Ozimec, 19.11.2005.
R. Ozimec, 25.05.2004.
J. Bedek, 27.10.2003.
M. Frani~evi}, 23.05.2004.
R. Ozimec, 08.11.2009.
H. Biland`ija, I. Jaklinovi}, M. Luki}, 28.10.2003.; H. Biland`ija, R.
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26.10.2006.
R. Ozimec, 28.10.2003.; J. Bedek, 26.10.2006.
H. Biland`ija, I. Jaklinovi}, M. Luki}, R. Ozimec, 29.10.2003.; R. Ozimec,
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R. Ozimec, 16.05.2010.
R. Ozimec, 20.06.2003.
R. Ozimec, 21.06.2003.
J. Bedek, 22.03.2003.
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R. Ozimec, 22.08.1997., 26.08.1998.; R. Ozimec, G. Nonveiller, 07.08.2000.;
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27.07.2004., 08.09.2005., 06.04.2006., 28.07.2006., 04.11.2009.
CBSS collection 543
J. Bedek, 22.10.2007.
CBSS collection 544
H. Cvitanovi}, 26.04.2004.
CBSS collection 545
M. Pavlek, 01.10.2007.
D. Hmura, I. Jaklinovi}, R. Ozimec, 02.10.2007.; L. \ud, 03.03.2008.; L.
CBSS collection 546
Jankovi}-Rapan, M. Manger, 04.03.2008.
CBSS collection 547
M. Luki}, 01.10.2007.
CBSS collection 548
P. Bregovi}, 05.03.2008.
CBSS collection 549
T. ^ukovi}, 05.03.2008.
CBSS collection 550
M. Luki}, 27.09.2007.
CBSS collection 551
R. Ozimec, 03.03.2008.
P. Bregovi}, 29.09.2007.; L. \ud, D. Hmura, 03.03.2008.; L. \ud, R.
CBSS collection 552
Ozimec, 04.03.2008.
D. Hmura, M. Luki}, 27.09.2007.; M. Pavlek, 03.10.2007.; L. \ud, A.
CBSS collection 553
]uku{i}, 04.03.2008.
CBSS collection 554
R. Ozimec, M. Pavlek, 26.04.2004.
CBSS collection 555
J. Bedek, 27.10.2006.
CBSS collection 556
R. Ozimec, 26.08.2005.
R. Ozimec, G. Nonveiller, 07.08.2000.; R. Ozimec, 13.07.2003., 08.09.2005.,
CBSS collection 557
04.11.2009.
CBSS collection 558
B. Jal`i}, 09.04.2007.
J. Bedek, H. Biland`ija, M. Pavlek, 09.04.2007.; J. Bedek, M. Pavlek,
CBSS collection 559
27.04.2008.
H. Cvitanovi}, M. Pavlek, 11.04.2007.; J. Bedek, M. Luki}, M. Pavlek,
CBSS collection 560
26.04.2008.
CBSS collection 561
M. Pavlek, 09.04.2007.
CBSS collection 562
J. Bedek, P. Bregovi}, 01.05.2008.
CBSS collection 563
J. Bedek, I. Bor{i}, T. Dra`ina, 20.06.2001.
CBSS collection 564
P. Rade, 14.04.2007.
CBSS collection 565
M. Pavlek, 07.04.2007.; H. Biland`ija, 12.04.2007.
CBSS collection 566
J. Bedek, 30.04.2008.
J. Bedek, 14.04.2007.; J. Bedek, H. Biland`ija, P. Bregovi}, M. Pavlek,
CBSS collection 567
02.05.2008.
CBSS collection 568
H. Biland`ija, 30.04.2008.
Croatian Natural History Museum (CNHM) collection
CNHM collection 1
B. Jal`i}, 17.06.2008.
CNHM collection 2
B. Jal`i}, 15.06.2008.
CNHM collection 3
J. Bedek, R. Ozimec, 28.06.2004.
CNHM collection 4
B. Jal`i}, 14.06.1996.
CNHM collection 5
B. Jal`i}, 29.04.2008.
CNHM collection 6
B. Jal`i}, 09.04.2007.
CNHM collection 7
B. Jal`i}, 16.09.2006.
CNHM collection 8
B. Jal`i}, 06.06.2006.
CNHM collection 9
B. Jal`i}, 27.06.2002.
CNHM collection 10 B. Jal`i}, 28.06.2002.
CNHM collection 11 B. Jal`i}, 25.05.2004.
CBSS collection 542
348
CNHM
CNHM
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B. Jal`i}, 24.06.2002.
B. Jal`i}, 25.05.2004.
B. Jal`i}, 26.03.2003.
B. Jal`i}, 26.03.2003.
B. Jal`i}, 26.06.2002.
B. Jal`i}, 15.09.2005.
M. Pavlek, 13.08.2005.; H. Biland`ija, 14.08.2009.
H. Biland`ija, F. Kljakovi} Ga{pi}, 15.08.2005.
B. Jal`i}, 18.08.2004.
H. Biland`ija, F. Kljakovi} Ga{pi}, M. Pavlek, 14.08.2005.
H. Biland`ija, 01.08.2009.
B. Jal`i}, 06.03.2001.
H. Biland`ija, B. Jal`i}, F. Kljakovi} Ga{pi}, M. Pavlek, 17.08.2005.
M. Pavlek, 17.08.2005.
B. Jal`i}, 29.06.2002.
B. Jal`i}, 22.05.2004.
V. Jal`i}, B. Jal`i}, 15.10.2002.
B. Jal`i}, 20.06.2003.
B. Jal`i}, 19.06.2003.
B. Jal`i}, 27.04.2002.
B. Jal`i}, 21.03.2004.
S. Vuj~i} – Karlo, 12.02.1998.
B. Jal`i}, 17.07.1997.
B. Jal`i}, 16.07.1997.; S. Vuj~i} – Karlo, 08.02.1998.
B. Jal`i}, 19.07.1997.
B. Jal`i}, 21.08.1997.
B. Jal`i}, 16.10.2007.
B. Jal`i}, 04.08.2004.
B. Jal`i}, 13.08.2001.
B. Jal`i}, 07.11.2006.
B. Jal`i}, 24.06.2009.
B. Jal`i}, 09.07.2007.
B. Jal`i}, 21.07.2007.
J. Bedek, R. Ozimec, 02.07.2004.
A. Jurinac, 20.08.1883.
B. Jal`i}, 07.06.2003.
B. Jal`i}, 01.12.2002.
A. Jurinac, 20.08.1883.
A. Langhoffer, [nap, 16.07.1902.
B. Jal`i}, 05.11.2006.
B. Jal`i}, 26.06.2009.
B. Jal`i}, 26.05.2001.
J. Bedek, R. Ozimec, 29.06.2004.
A. Langoffer, 14.08.1902.
D. Hirc; J. Notenboom, 03.07.2004.
J. Bedek, 30.06.2004., 10.07.2004.
Nat. Croat. Vol. 20(2), 2011
CNHM collection 58 H. Cvitanovi}, S. Gottstein Mato~ec, R. Ozimec, 03.07.2004.
CNHM collection 59 A. Jurinac, 25.08.1883.; J. Bedek, B. Jal`i}, R. Ozimec, 01.07.2004.
CNHM collection 60 A. Jurinac, 20.08.1883.
CNHM collection 61 B. Jal`i}, 12.09.1999.
CNHM collection 62 J. Bedek, H. Cvitanovi}, 27.08.2004.
CNHM collection 63 A. Jurinac, 20.08.1883.
CNHM collection 64 B. Jal`i}, 10.02.1998.
CNHM collection 65 B. Jal`i}, 21.03.2004.
CNHM collection 66 B. Jal`i}, 09.02.1988.
CNHM collection 67 B. Jal`i}, 09.04.2007.
CNHM collection 68 B. Jal`i}, 27.04.2008.
Museo di Storia Naturale dell’Università di Firenze (MZUF) collection
MZUF collection 1
B. Jal`i}, 13.02.1998.
MZUF collection 2
M. Pavlek, 07.06.2006.
MZUF collection 3
M. Pavlek, 23.05.2004.
MZUF collection 4
R. Ozimec, 23.03.2004.
MZUF collection 5
J. Bedek, 21.03.2004.
MZUF collection 6
A. Komeri~ki, 24.04.2009.
MZUF collection 7
M. Luki}, 01.05.2010.
MZUF collection 8
J. Bedek, H. Biland`ija, B. Jal`i}, 22.06.2005.
MZUF collection 9
A. Kirin, 12.05.2009.
MZUF collection 10
H. Cvitanovi}, R. Ozimec, P. Rade, 19.03.2009.; J. Bedek, 06.06.2009.
MZUF collection 11
H. Biland`ija, F. Kljakovi} Ga{pi}, B. Jal`i}, 13.04.2008.
MZUF collection 12
H. Biland`ija, 11.12.2004.
MZUF collection 13
J. Bedek, M. Luki}, 23.04.2009.
MZUF collection 14
J. Bedek, 27.06.2009.
MZUF collection 15
J. Bedek, 15.08.2009.; H. Biland`ija, 05.09.2010.
MZUF collection 16
M. Luki}, 31.01.2008.
MZUF collection 17
R. Bakovi}, 20.06.2009.
MZUF collection 18
J. Bedek, 05.09.2009.
MZUF collection 19
R. Ozimec, 25.03.2003.; M. Luki}, 19.08.2007.
MZUF collection 20
M. Luki}, 26.04.2008.
MZUF collection 21
A. Komeri~ki, 04.05.2010.
MZUF collection 22
K. Miculini}, 02.06.2010.
MZUF collection 23
R. Ozimec, 20.10.2005.
MZUF collection 24
M. Luki}, 12.02.2005.
MZUF collection 25
H. Cvitanovi}, 04.03.2008.
MZUF collection 26
H. Biland`ija, B. Jal`i}, 26.05.2007.
Department of Biology and Ecology, Faculty of Sciences, University of Novi Sad collection
DBE collection 1
J. Bedek, 17.11.2007., 05.04.2008.
Ton}i Ra|a collection
T. Ra|a collection 1
T. Ra|a, S. [ore, 25.08.2001.
349
350
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
APPENDIX 3.
Alphabetical order of localities
Absturz-Höhle 289
Ambarac 253, 281, 287, 329, 331,
332, 333
Aragonka 289, 290, 296
Baba {pilja 267, 326
Babina jama 285
Bali}eva {pilja 262, 326, 327, 330
Banova ljut 289, 290
Bara}eva {pilja 281, 326, 328
Baradla barlang 244
Bari~evi}a pe}ina 265, 326, 327
Barinova jama 276, 285
Bast 309
Bazgova~a jama 273, 274, 326,
328, 332
Beaune 299
Belinca jama 275
Benetina pe}ina 271, 316
Benova jama 261
Bezdan 310
Bibi~ka {pilja 250, 253, 281
Bijeli bunar 244
Bila lipotica 263
Bilek 270
Billova ponikva 285
Biserujka 268, 328, 329, 330, 331,
332, 333
Bla`eva jama 286, 331
Bobija jama 265, 328
Boru{njak 1 276, 285
Boru{njak 3 276, 285
Bo{ak 261
Breg jama 310
Bribirska pe}ina 283, 327
Brija~ica 282
Br{ljanovica 285
Bubnjava~a 261, 294
Bukovac 286, 327, 328
Bunarina 262
Bunetova jama 277
Bunker blizu uvale Perilo 248
Bu{ljeta 261
Carrara 246
Caretova {pilja 255, 256, 308, 328
Casello ferrovario 286
Cepinka 250, 282
Ceri~ka pe}ina 281, 327
Cerjanska {pilja 247, 326, 332
Cimerjanka 246
Cingulica pod @idovinom 247
Cinkalica 1 247
Cinkalica 3 247
Crekvica 276
Crna pe}ina 280, 332, 333
Crna~ka {pilja 287
Crnomarkovi}a pe}ina 265, 327
Crvenku{a – Tamnica – Suhi Rumin 263, 327, 331
^ampari 275, 329, 331
^injadra 273, 274, 304, 305, 326,
328, 332
^o}ina jama 292, 302, 328
^ude 260
]uli}a jama 264
Dakina jama 282
Damljinhova jama 265, 328
Danetska pe}ina 250, 282
Debela ljut 289, 290
Denizza 275, 328
Diva{ka jama 285
Dobra jama 274, 326
Donja Bara}eva {pilja 281, 284,
326, 327, 328
Dragina pe}ina na Dobri 250, 282
Drenova~a 260
Drinova II 264, 301
Drobovnik 282
Dru{ka pe} 249, 276
Duboka trepetljika 264
Dumen~i}a {pilja 281
Duplja 246
Dvoga}a 250
D`amije 265, 328
\ot 282
\ukina velika {pilja 250
\ulin ponor – Medvedica 253,
287, 331
\urovi}a {pilja 257, 291, 297, 326,
328
Fe{tinsko kraljevstvo 275
Filipovi}eva jama 273, 274
Forno 246
Francuski rudnik IV 244
Ga}ina jama 264
Gali~njak 258, 291, 295, 300, 310
Garden of Biological institute Lapad 257, 290
Germany 249
Gojak 287
Golubinka 265, 328
Golubinjka jama 260, 284
Golubnja~a 260, 280, 296, 328
Goranska Klepa 247
Gornja Bara}eva {pilja 281, 284,
326, 327, 328
Gornja Cerova~ka {pilja 261
Gornja jama na Grljaku 264
Gornja Tuti}eva {pilja 261
Gospodska {pilja 262, 327, 328
Govedija jama 263
Gradska spila 261, 301
Gradusa 244
Grap~eva {pilja 272, 328
Gr~ka pe}ina 272, 328
Grnja~a 286
Grotta ad ovest di Basovizza 275
Grotta dell’Orso 252
Guge~ka pe}ina 253, 281, 330
Hajdu~ka pe}ina 250, 253, 287
Han`ekova {pilja 247
Hircova jama 286
Horvatove stube 250
Hrastovska {pilja 257, 308
Ikinica 275
Ilijina pe}ina 289
Ivina jama 261
Izvor {pilja kod Tro{marije 250
Izvor {pilja kod Trupkovi} mlina
283
Jajce 245, 330
Jakasova {pilja 258, 269, 294, 305,
328, 329, 331, 332, 333
Jama II kod Koli}a 253, 282
Jama 3 253, 281
Jama bez imena 282
Jama bijele sige 244, 328
Jama Cvjetana Barbi}a 308
Jama ispod Belec grada 247
Jama ispod Tomini}evog brega
252, 285
Jama iznad Hajdove hi`e 286
Jama iznad Jezerca 244
Jama iznad Lipog do~i}a 264
Nat. Croat. Vol. 20(2), 2011
Jama iznad Prozor~i}a 264, 301
Jama iznad Sarana~a 261, 293
Jama iznad Vranjske drage 276,
299
Jama kod Buri}i 275
Jama kod crvenog gloga 251, 285
Jama kod dje~jeg oporavili{ta 2
285
Jama kod Dugopolja 265
Jama kod Jatare 2 261
Jama kod Kasuni}a 253
Jama kod Klja~ine 283
Jama kod Male U~ke 285
Jama kod potoka Banine 3 276,
286, 298, 299, 332
Jama kod Rado{i}a 263, 326
Jama kod Ra{pora 285
Jama kod Starog mlina 282
Jama kod Tu~epske vilenja~e 261
Jama kod Vidove gore 274, 329,
330
Jama mrtvih golubova 276
Jama na Bala~i 265, 326, 330
Jama na Balagu{tini 251, 276
Jama na Bertiji 1 253, 281
Jama na Bertiji 2 253, 281
Jama na Bilu 244
Jama na Brekalcima 258
Jama na Brezdanu 264
Jama na Brlogu 284
Jama na Docima 261
Jama na Gnojini 282
Jama na grebenu 261
Jama na Javorniku 282
Jama na Ka~koj glavici 3 264
Jama na Kamenoj 269
Jama na Ko{taku 263, 326, 330
Jama na Kuku 262
Jama na Lovranskim lazi}ima
286
Jama na Mahencu 2 252, 276
Jama na Mehovnju 269, 294
Jama na Mr`dolcu 264
Jama na Odvoru 258
Jama na Poklonu 285
Jama na polju 260
Jama na prevoji}u 276, 285
Jama na putu za Staru {kolu 261
Jama na rubu 282
Jama na [krilama 282
Jama na Teruncu 272
Jama na Visokoj 263, 326
351
Jama na vrh Prodoli 290
Jama na vrh Vrguda 289, 290,
296
Jama nasuprot Torka 296
Jama pod Bilu{inim vrtom 269
Jama pod Boljunskim dolom 285
Jama pod ^ezinim humcem 284
Jama pod Gradanom 253
Jama pod Gradom 250, 283
Jama pod Kola~ 258, 269
Jama pod Kragljevcem 262, 326
Jama pod kri` 276, 285
Jama pod Malim Kraljevcem 283,
328, 330
Jama pod Osojem 267
Jama pod Sv. Jurom 264
Jama pod Velikim Knezgradom
252, 286
Jama pod Velji vrh 257
Jama u Bratu{u 301, 309
Jama u ^ike{ima 264
Jama u Dovnju 264
Jama u Dubokom dolu 269, 294
Jama u kanjonu Dobre ispod
Grabrka 253, 282
Jama u Konjiku{i gornjoj 263
Jama u Lukama 282
Jama u Mikulin dolac 269
Jama u trepetljikama 264
Jama u uvali Kru~ica 308
Jama u Velikom Desajdubu 258,
310
Jama u Velim Sapcima 3 252, 286
Jama u Vetvi 286
Jama u Vrisju 269
Jama u Vrloj strani 250, 282
Jama u Zabira|u 270, 289, 290
Jama uz stazu na Sv. Nikoli 290
Jama za greben 285
Jama za Mahrincem 273, 274, 330
Jama za Smokvom 308
Jama za Supinom 261, 267
Jama za Svetim Ilijom 291
Jama za Svetim Ivanom 258, 291,
310
Jama za Svetim Spasom 258, 291,
310
Jamica u docima 279
Jamina 264, 283, 301
Jan~ikin guz 244
Jara~ na ponti od Graca 291
Jastrebina 244, 250, 283
Jatara 261
Jazinka 262
Jelenova {pilja 244
Je{kalovica 274, 306, 326, 330
Jezero 283, 290, 326, 330
Jopi}eva {pilja – Bent 283
Jujnovi}a {pilja 261
K’U~ka 285
Kad jaganjci utihnu 261
Kapaljka 1 286
Karle 257, 291, 327, 330
Karlova jama 277
Kaverna pod vijaduktom kod
Puljana 279
Kaverna u tunelu U~ka 286
Kaverna uz cestu iznad Groma~e 290
Kitlica jama 263, 327
Klanj~eva pe} 252
Koli{tina 260, 330
Kopa~ina 273, 274, 330
Kornjatu{a 270
Kosilova jama 262
Kosinac 263
Kotari 282
Kotlu{a 259, 262, 329, 330, 331,
333
Kova~evka pe}ina 283, 327
Kozjak pe}ina 265, 328
Kraljicina {pilja 256
Kranjica 262
Kr~evina 281, 331
Kriva~a 291
Krjava 1 261
Krjava 2 261
Krka 297, 298, 330
Kukelj sjever 247
Kukor 264, 301
Kukova pe} 270, 290, 291
Kuna 261, 292, 301
Kusa nad Manastirskim lukama
260
Ku{trovka 253, 281
Lalovi}eva jama 282
Langhofferova {pilja 288, 327
Lapad 307, 308
Lastovo 308
Ledena {pilja 286, 327, 328, 330
Ledenica 246
Ledenica kod Stare {kole 261
Ledenica na podno`ju Jabukovca 263, 326, 330
Ledenica u [peharima 253, 282
352
Bedek, J. et al.: Catalogue and Altas od Cave – Dwelling Terrestrial Isopods from Croatia
Ledeni~ka {pilja 250, 253, 283,
327, 330
Likar ponor 287
Lipa na Protulipi 250, 253, 283
Li{trovica 287
Lokvarka 277, 286, 327, 331
Luda bu`a 244
Lukina jama – Trojama sustav
271, 316, 331
Lu{ka {pilja 287
Maica golubinka 279
Makluta~a 263
Mala jama 261
Mala jama u stijenama 264, 301
Mala pe} 252, 276
Mala {pilja 258, 261
Mala Ze~ica 283
Male ponte 258, 291, 310
Male spile 301
Mali Boru{njak 6 285
Mali {tumbur 277, 300
Mandalina 298, 300
Mandelaja 253, 281, 287
Manita pe} 261
Manzin 275
Mar~ina jama 283
Marijan 309
Markov ponor 287
Markova jama 275
Markova {pilja 272, 331
Marmo 275
Martina jama 260, 261
Massa 246
Matija{eva {pilja 261, 301
Matuli}eva pe}ina 289
Medi}a pe}ina 262, 327
Medina {pilja 288, 331
Medvednica 244, 331
Medvi|a ropa 257, 308
Medvje|a {pilja 277
Medvje|i brlog 244
Me|ugra|en 258
Mika{inovi}a {pilja 253, 281, 333
Mili}ka 282
Miljacka I 260
Miljacka II 260, 295
Minjera 274
Mla|enova pe}ina 283, 327
Mo~iljska {pilja 289, 290, 296, 327,
331, 333
Mokra~a 254
Movrica 258, 291, 293, 294, 310,
326, 327, 329, 331
Mra~na {pilja 280, 331
Mravinju{a jama 265, 268, 292
Munjave 281, 327, 330
Mu`ina {pilja 264
Nacina jama 286
Nesretnja jama 284
Nove jame 286, 326
Novogodi{nja jama 276, 285
Odisejeva {pilja 310
Od`i}a ponor 263
Ogra|ena {pilja pod Sv. Jurom
301
Ombla 291, 307, 308
Opatija 300
Oporovina 249
Orlova~a 284
Osta{evica 258, 291, 303, 326,
327, 331
O{tirova jama 276, 285
Ote{ka pe}ina 250, 331
Ov~arica 275
Ov~ja pe} 252
Ozaljska {pilja 250, 330, 331
Pasja jama 291
Pasji manastir 264
Pastirski stan 264
Pave}a {pilja 253
Pavli~evica 270
Pavlovica 282
Pe~enjevka 282
Pe~ine 282
Pe} na Glu{cu 284
Pe}ina Bivio Beru}a 287, 329
Pe}ina isto~no od sela Svagu{a
264
Pe}ina Ivana Stani}a i Franja Stani} 286, 330, 331
Pe}ina iznad Kule 262
Pe}ina kod dje~jeg oporavili{ta
285
Pe}ina kod planinarske ku}e Pavlovac 276
Pe}ina kod Pu`i 276, 329, 332
Pe}ina kod sela Puhari 252, 277,
286
Pe}ina kod Sinja 265, 327, 329
Pe}ina kod Zagvozda 301
Pe}ina Marije Ru`i} 286, 330
Pe}ina na Crekvici 1 252, 276
Pe}ina na isto~noj padini Mosor planine 265, 329, 330
Pe}ina na izvoru Cetine 262, 329,
330
Pe}ina na sjeveroisto~nim padinama Biokova 265, 332
Pe}ina na Svilaji planini 265, 329,
330
Pe}ina pod Brlo`nikom 276
Pe}ina pod Pe}anima 282
Pe}ina pod Sokoli}em 276
Pe}ina pod Str`en 276, 285
Pe}ina poli Vrti}a 252, 285
Pe}ina sjeveroisto~no od Sinja
265, 329, 331
Pe}ina Stojakovi} 265, 328
Pe}ina u Carevoj Poski 261
Pe}ina u Donjem Bri{niku 267
Pe}ina u Guge~kom usjeku 253
Pe}ina u Jurkovoj dolini 264
Pe}ina u Sridnjoj gori 264, 302,
332
Pe}ina u Topolovcu 276
Pe}ine u {umi 271
Pe}inik 281
Per~evi}a {pilja 253, 287
Petnaestica 244
Petr~eva jama 252, 276
Petrinjka 262
Pid`in pit 264
Pisanica 261
Piskovica 275
Pi{urka 257, 258, 269, 329, 331,
332
Pivnica 282
Plavinka 282
Plitvi~ka {pilja 247
Plo{nica 264
Podbirnja~a {pilja 265, 328
Podrum 261
Polu{pilja ispod ku}e u Bastu 294
Polu{pilja kod Sobre 310
Pometena golubinka 264
Porto Madonna 249, 275, 332
Poskokova rupa 301
Postojnska jama 280
Potkameni~ina jama 264
Potok 283
Pozjata 264, 267
Po`arova jama 279
Pr~ina 260
Proljetna jama 276, 286
Propod 272
Provala 283
353
Nat. Croat. Vol. 20(2), 2011
Pru`ina 264, 302
Pukotina kod doca Stra`benica
264
Pupi}ina pe}ina 276
Pustinja 286, 327
Pu{ina 250
Rabakova {pilja 275, 332
Ra~a 257
Ra~i{ka pe~ina 245, 328, 330, 333
Radota jama 285
Raki~ka 282
Raputinova {pilja 264
Rastova~a 260
Rogovac 283
Rokina bezdana 287
Rov na Lapadu 290
Rudeli}a {pilja 262, 327, 332
Rudina jama 290
Rudnik iznad uvale Martin{~ica 248
Rudnik Kra{evski zviri 244
Rudnik u Medveji 276, 300
Rudnik u Velikoj Sotinskoj 246
Rudnik Zrinski 244, 333
Rupa I 252, 276
Rupa II 252, 276
Rupa na Ribinovoj bandi 256
Ru{evina 286
Samograd 258, 269, 287, 294, 296,
306, 309, 332
Santa Margherita 246
SD 2 276
SDB 285
SDB 2 286
Sedrena 1 kod slapa Skakavac
244
Sedrena {pilja iza mlina 296
Sini~i}a {pilja 287
Sitvukova pe}ina 287, 332
SKH 286
Smokvica 260, 326
Snije`nica pod Ljubljanom 263,
328
South Herzegovina 290
Spasiteljica 264
Spila 1 261, 332
Spila 2 261
St. Lucia 275
Stankova {pilja 250
Stara jametina 260, 296
Stonjska pe} donja 261, 301
Stonjska pe} gornja 261
Stra~in~ica 308
Stra`benica 260
Studentski ratac 274, 326
Sumpora~a 298
Sumpora~a mala 291
Su{ica 250, 253, 281
Su{ik 250, 287
Suvi Rumin 263, 329
Svetica 264
[ari}a jama 281
[ipun 257, 291, 296, 306, 326, 327,
332
[karin Samograd 260
[koljka~a 247
[krabuljica 290, 331
[pilja do Obilje`ene 264
[pilja ispod Krsta 290
[pilja izme|u Lokvica i Blisine
glavice 274, 330
[pilja izme|u Milne i Sv. Josipa
274, 330
[pilja iznad ceste 248, 277, 300
[pilja kod Dobre vode 246
[pilja kod izvora Rude 264
[pilja kod Jura{eve livade 282
[pilja kod Mlina na Miljacki 260,
295
[pilja kod Murvice 304, 328, 329,
331
[pilja kod Nerezinog dola 258,
332, 333
[pilja kod Podumolskog mlina
250, 282
[pilja kod slapa kod Kasuni}a
250
[pilja kod Solina 258, 291, 294,
310
[pilja kod Starigrada 288, 329
[pilja kod vrha Spile 310
[pilja na Bardarovici 255, 256,
326
[pilja na brdu Bugara 310, 328,
329
[pilja na Dinari 254, 265, 330
[pilja na Dinari 256, 265, 330
[pilja na Dinari 258, 265, 330
[pilja na Dinari 260, 265, 330
[pilja na Gradini kod Premanture 248, 249, 275, 328, 329,
331, 332, 333
[pilja na Mr`dolcu 265
[pilja na Pozalicu 257, 309
[pilja na Svilaji planini 295, 296,
329, 332
[pilja na U~ki 251, 252, 277, 328,
329, 332, 333
[pilja na vrh Kr~evina 291
[pilja na Vr{inoj glavici 279, 316
[pilja na Zalokavlju 270
[pilja od Bilosoja 278, 329, 331,
333
[pilja od Vore 256
[pilja pod Butinom 247
[pilja pod Humom 255, 256, 332
[pilja pod Kapelu 272, 332
[pilja pod Ka{tel 257, 309
[pilja pod Ma~kovom dragom
287
[pilja pod Veli vrh 269
[pilja pod Vrajinim stijenama
250
[pilja pod Vrhom 273, 274, 332
[pilja pod Zimzelom 281
[pilja pokraj Kozja~e 283
[pilja Sv. Filipa i Jakoba 278, 329,
332, 333
[pilja u Bastu 265, 302, 309
[pilja u dolini Neretve 279, 316,
331
[pilja u Gaju 265, 291
[pilja u kamenolomu Tounj 281
[pilja u kanjonu Dobre ispod
Podjak{i}a 250, 253
[pilja u Lukama 283
[pilja u Mekoti 253, 281, 329, 332
[pilja u uvali Predpe} 277
[pilja u uvali Sve`anj 248, 277
[pilja uz cestu kod Komi`e 255
[temajzlinka 286
[upljara 281, 327
[utova~ka pe}ina 253
Tabaina {pilja 255, 256, 308, 333
Tamnica 253, 281
Tanki ratac 273, 274
Tanki rot 256
Tavnica 260
Ton~akova jama 277
Topla pe} 260, 296
Topolska pe} 252, 276, 285
Tounj~ica 250, 253, 287
Trebinje 306
Trijama II 282
Tri{i}a jama 260
Trlica 1 286
Trojama 263, 326
354
Bedek, J. et al.: Catalogue and atlas of cave–dwelling terrestrial isopods from Croatia
Tr`i}ka {pilja 282, 287, 329, 331,
333
Tu~epska vilenja~a 261, 267, 301
Tunel u Pera~kom blatu 270
Turska pe} 284
U~ka 245, 331
Urinjska {pilja 248, 277, 298, 300
Uvala kod Bakra 248
Uviraljka 244
Vaganka 261
Vatrena jama 282
Vela pe} 276, 286
Vele vode 249, 286
Velebit 249
Velebita 285
Velika ^ulumova {pilja 262
Velika pe} 244, 284
Velika pe}ina 260, 283, 327
Velika {pilja 247, 258, 265, 267,
291, 293, 310, 327, 328, 331,
333
Velika Ze~ica 283
Veliki 287, 326
Veliki rudnik 256
Veternica 244
Vidovi}a {pilja 253, 287
Vihra 247
Vila 247
Vilenska jama 290, 326
Vilenska pe} 270, 297
Vilina pe}ina 296
Vilina {pilja – Ombla 289, 291,
333
Vindija 247, 327, 333
Vi{ti}ina jama 270
Vitkova~a 290
Vla{ka pe} 254, 327, 329, 333
Vodena jama 253, 282
Vodena pe}a 264
Vodena trepetljika 265
Vrana 248
Vranja~a 263, 301, 326, 329, 330,
331
Vranja~ka {pilja 250
Vra`i}a {pilja 283, 326, 327, 333
Vreli} 253, 281
Vrelo 286
Vrlovka 250, 253, 283, 333
Vr{ine 266, 326
Vrtare male 277
Vrulje 283
Vu~ja pe}ina 289, 296
Vu~kovi} jama 265, 328
Zadubravica 270, 290, 291
Zagorska pe} 287, 329, 331, 333
Zakanova jama 265
Zdenec pri Ciglaru 247, 327, 332
Ze~ica 267
Ze~ja jama 291
Zerngol {pilja 288, 327
Zimska pe}ina 263, 327
Zobenica pe}ina 281
Zrilina pe}ina 265, 327
ZV – 1 286
Zve~ara 260, 332
Zverina 283
Zvirnjak pe}ina 281, 327
Zvone~ka II 282
@ablja jama 264
@ejava 274, 326, 333
@idovske ku}e 250
@lota 283
@u~ki 253