Bull. Natl. Mus. Nat. Sci., Ser. B, 38(3), pp. 93–119, August 22, 2012
Rare and Threatened Pteridophytes of Asia 1. An Enumeration of
Narrowly Distributed Taxa
Atsushi Ebihara1,*, Christopher Roy Fraser-Jenkins2, Barbara S. Parris3,
Xian-Chun Zhang4, Yue-Hong Yang5, Wen-Liang Chiou6, Ho-Ming Chang7,
Stuart Lindsay8, David Middleton8, Masahiro Kato1, Titien Ngatinem Praptosuwiryo9,
Victor B. Amoroso10, Julie F. Barcelona11, Rajapaksha Haddokara Gedara Ranil12,
Chan-Ho Park13, Noriaki Murakami14 and Akihiko Hoya1
1
Department of Botany, National Museum of Nature and Science,
4–1–1 Amakubo, Tsukuba, Ibaraki 305–0005, Japan
2
Student Guest House, Thamel, P.O. Box no 5555, Kathmandu, Nepal
3
Fern Research Foundation, 21 James Kemp Place, Kerikeri, Bay of Islands 0230, New Zealand
4
Institute of Botany, Chinese Academy of Sciences, Beijing 100093, China
5
Shanghai Chenshan Plant Science Research Center, Chinese Academy of Sciences,
Shanghai Botanical Garden, Shanghai 201602, China
6
Taiwan Forestry Research Institute, Taipei 10066, Taiwan
7
Endemic Species Research Institute, Nantou 55244, Taiwan
8
Royal Botanic Garden Edinburgh, 20A Inverleith Row, Edinburgh, EH3 5LR, Scotland, U.K.
9
Center for Plant Conservation, Bogor Botanical Gardens,
LIPI, Jl. Ir. H. Juanda No. Bogor, West Java, Indonesia
10
Central Mindanao University, Bukidnon, Philippines
11
School of Biological Sciences, University of Canterbury, Christchurch 8140, New Zealand
12
Department of Crop Science, Faculty of Agriculture, University of Peradeniya,
Peradeniya, 20400, Sri Lanka
13
National Institute of Biological Resources, Korea
14
Makino Herbarium, Tokyo Metropolitan University, Japan
* E-mail: ebihara@kahaku.go.jp
(Received 17 May 2012; accepted 26 June 2012)
Abstract A list of narrowly distributed Asian pteridophyte taxa towards an assessment of globally threatened species is provided. Of the total 886 taxa (835 species and 51 infraspecific taxa)
enumerated, 577 occur in Southeast Asia, 215 in East Asia and 101 in South Asia.
Key words : endangered species, ferns, lycophytes, monilophytes, pteridophytes.
The pteridophyte (monilophyte and lycophyte)
flora of Asia is remarkably rich as demonstrated
by the fact that an estimated 4500 species, more
than one third of the worldʼs diversity, occurs in
Southeast Asia (Moran, 2008) alone. Half a century has passed since the first publication of the
Flora Malesiana series II (pteridophytes). These
publications, however, do not yet even cover half
of the species that occur in the region and it
seems a considerable amount of time will be
required for their completion (Roos, 1996). On
the other hand, the speed of deforestation in Asia
is remarkably high (Sodhi and Brook, 2006), and
pteridophytes, which are major constituents of
the forest understorey, are undoubtedly under
threat. Even though there are only a limited number of Asian pteridophyte species that have been
assessed and included in the IUCN Red List,
many Asian countries/areas have already published or compiled local red lists of vascular
Atsushi Ebihara et al.
94
plants including pteridophytes (e.g. Lee, 2009 for
South Korea; Japanese Ministry of Environment,
2007 for Japan; Wang, 2010 for Taiwan; Fernando et al., 2008 for the Philippines; Lindsay
and Middleton, 2010- onwards for Thailand
[assessed global threats following the IUCN criteria (IUCN Standards and Petitions Subcommittee, 2011)]; Chandra et al., 2008 for India).
Threatened species appearing in regional red lists
are not necessarily globally endangered; there are
many species which are locally endangered (at
least in a part of the distributed countries/area)
but are assessed as Least Concern over their
entire distribution. In the present paper, we
extracted ʻnarrowly distributedʼ taxa (i.e. endemic
to one country/area or distributed only in a few
localities even when their ranges straddle parts of
two or more countries/areas) from previously
published regional red lists. We have also
attempted to include possible ʻrare and threatenedʼ taxa in the countries/areas whose red lists
have not yet been compiled based on our current
knowledge from field and herbarium studies.
Although this provisional list covers neither all
of the countries/areas nor all the taxa we present
it as a starting point for compilation of a red list
of Asian pteridophytes using the IUCN criteria
(IUCN Standards and Petitions Subcommittee,
2011) through international collaboration.
Table 1. A provisional list of narrowly distributed monilophytes and lycophytes in Asia that require assessments
for a global red list. The system by Christenhusz et al. (2011) is adopted for family-level classification, but
generic concepts are as supplied by contributing authors due to current lack of consensus on generic delimitation in some cases and the lack of published combinations in some genera. Taxon concepts of Indian species
mostly follow Fraser-Jenkins (2008). Distributions are indicated by islands or island groups rather than by
countries. Published categories of threatened species in regional red lists are indicated in ʻNoteʼ for those distributed only in one country/area if available (Japanese Ministry of Environment, 2007 for Japan; Wang, 2010
for Taiwan; Fernando et al., 2008 for the Philippines; Parris et al., 2010 for Peninsuolar Malaysia; Johns et al.,
2006 for New Guinea [Prov. Papua of Indonesia]) and also for those globally assessed (Lindsay and Middleton, 2010- onwards for Thailand, Laos and Cambodia).
Name
Lycopodiaceae
Huperzia bucahwangensis Ching
Huperzia crispata (Ching ex H. S. Kung)
Ching
Huperzia cunninghamioides (Hayata) Holub
Huperzia elmeri (Herter) Holub
Huperzia lucidula (Michx.) Trev.
Huperzia medogensis Ching et Y. X. Lin
Huperzia muscicola Ching et W. M. Chu
Huperzia nilagirica (Spring) R. D. Dixit
Huperzia quasipolytrichoides (Hayata)
Ching var. rectifolia (J. F. Cheng) H. S.
Kung et L. B. Zhang et H. S. Kung
Huperzia rubicaulis S. K. Wu et X. Cheng
Huperzia taiwanensis (C. M. Kuo) C. M.
Kuo
Huperzia tetrastichua (Kunze ex Alderw.)
Holub
Huperzia vernicosa (Hook. et Grev.) Trevis
Huperzia verticillata (L. f.) Trevisan var.
giganteum Herter (=Huperzia gigantea
(Herter) Holub)
Lycopodium halconense Copel.
Phlegmariurus austrosinicus (Ching) L. B.
Zhang
Distribution
China (Yunnan)
China (Chongqing, Guizhou, Hubei,
Hunan, Jiangxi, Sichuan, Yunnan)
Japan (Amami Isl. [extinct]), Taiwan
Philippines (Benguet, Tayabas, Mindoro,
Palawan, Cotabato, Davao)
China (Jilin)
China (Xizang)
China (Yunnan)
S. India (Kerala, Tamil Nadu)
China (Jiangxi, Hunan)
China (Yunnan)
Taiwan
Note
CR in Taiwan
DD in Taiwan
Maluku Islands (Seram) to Polynesia
S. India (Kerala, Tamil Nadu), ?Sri Lanka
Philippines (Davao, Cotabato, Misamis
Occidental)
Philippines (Mindoro)
China (Chongqing, Guangdong, Guangxi,
Guizhou, Hunan, Jiangxi, Sichuan,
Hongkong, Yunnan)
EN in the Philippines
Narrowly Distributed Asian Pteridophytes
Table 1.
Name
Lycopodiaceae (continued)
Phlegmariurus guangdongensis Ching
Phlegmariurus ovatifolius (Ching) L. B.
Zhang
Phlegmariurus yunnanensis Ching
Selaginellaceae
Selaginella aitchisonii Hieron.
Selaginella apoensis Hieron.
Selaginella atimonanensis B. C. Tan et
Jermy
Selaginella calcicola Hieron.
Selaginella canaliculata (L.) Spring
95
(Continued)
Distribution
Note
China (Guangdong, Guangxi, Hainan)
China (Yunnan)
China (Yunnan)
Tien Shan, Sinkiang, Afghanistan,
Pakistan, India (Jammu , Kashmir)
Philippines (Mindanao: Mt. Apo);
Maluku Islands (Seram)
Philippines (Quezon)
EN in the Philippines
Selaginella cataractarum Alston
Maluku Islands (Seram), Borneo
Maluku Islands (Seram, Ambon, Miangas, Morotai, Halmahera, Ternate)
S. India (Tamil Nadu)
Selaginella chaii Jermy
Selaginella curtisii Ridl.
Borneo (Sarawak)
Peninsular Malaysia
Selaginella devolii H. M. Chang, P. F. Lu et
W. L. Chiou
Selaginella doederleinii Hieron. var. opaca
Seriz.
Selaginella gaudichaudiana Spring
Selaginella hainanensis X. C. Zhang et
Noot.
Selaginella ingens Alston
Selaginella latifolia (Hook. et Grev.) Spring
Selaginella lineariformis Jermy
Selaginella miniatospora (Dalzell) Baker
Selaginella ornithopodioides (L.) Spring
Selaginella pilosula Alderw.
Selaginella pricei B. C. Tan et Jermy
Selaginella ridleyii Baker
Taiwan
CR in Peninsular
Malaysia
EN in Taiwan
Japan (Izu Islands)
NT in Japan
Borneo (Sabah)
Sri Lanka
Borneo (Sarawak)
S. India (Maharashtra, Goa, Karnataka)
Sri Lanka
Maluku Islands (Seram, Ternate)
Philippines (Western Samar)
Peninsular Malaysia
Selaginella rivalis Ridl.
Peninsular Malaysia
Selaginella robinsonii Alderw.
Selaginella selangorensis Bedd. ex Ridl. var.
selangorensis
Selaginella selangorensis Bedd. ex Ridl. var.
ciliata Alston
Selaginella wattii Baker
Isoetaceae
Isoetes hopei J. R. Croft
Possibly extinct in
known localities
Maluku Islands (Seram, Morotai)
China (Hainan)
Maluku Islands (Ambon, Seram)
Peninsular Malaysia
Peninsular Malaysia
N. E. India (Manipur), ?Myanmar.
New Guinea (Prov. Papua)
Isoetes hypsophila Hand.-Mazz.
Isoetes neoguineensis Baker
Isoetes orientalis H. Liu et Q. F. Wang
Isoetes philippinensis Merr. et L. M. Perry
China (Jiangxi, Sichuan, Yunnan)
New Guinea (Papua New Guinea)
China (Zhejiang)
Philippines (Lanao del Norte)
Isoetes pseudojaponica M. Takamiya, Mitsu.
Watan. et K. Ono
Isoetes sahyadriensis Mahabale
Isoetes sinensis Palmer var. sinensis
Japan
C. India (Maharashtra, Madhya Pradesh)
China (Anhui, Guangxi, Hunan, Jiangxi,
Zhejiang), Japan
EN in the Philippines
EN in Peninsular
Malaysia
CR in Peninsular
Malaysia
VU in Peninsular
Malaysia
VU in Peninsular
Malaysia
EN in New Guinea
(Prov. Papua)
CR in the Philippines,
possibly extinct in
known locality
VU in Japan
Atsushi Ebihara et al.
96
Table 1.
(Continued)
Name
Isoetaceae (continued)
Isoetes sinensis Palmer var. coreana (Y. H.
Chung et H. K. Choi) M. Takamiya,
Mitsu. Watan. et K. Ono
Isoetes stevensii J. R. Croft
Isoetes taiwanensis DeVol
Isoetes yunguiensis Q. F. Wang et W. C. Taylor
Ophioglossaceae
Botrychium microphyllum (Sahashi) K.
Iwats.
Mankyua chejuensis B. Y. Sun, M. H. Kim
et C. H. Kim
Ophioglossum kawamurae Tagawa
Ophioglossum namegatae M. Nishida et Kurita
Ophioglossum parvum M. Nishida et Kurita
Marattiaceae
Angiopteris caudatiformis Hieron.
Angiopteris hokouensis Ching
Angiopteris ruttenii Alderw.
Angiopteris sparsisora Ching
Archangiopteris bipinnata Ching
Archangiopteris hokouensis Ching
Archangiopteris itoi W. C. Shieh
Archangiopteris somai Hayata
Marattia boninensis Nakai
Osmundaceae
Osmunda bromeliifolia (C. Presl) Copel.
Osmunda hybrida Tsutsumi et al.
Osmunda mildei C. Chr.
Todea papuana Hennipman
Hymenophyllaceae
Abrodictyum boninense Tagawa et K. Iwats.
Abrodictyum clathratum (Tagawa) Ebihara
et K. Iwats.
Abrodictyum kalimantanense (K. Iwats. et
Kato) Ebihara et K. Iwats.
Cephalomanes crassum (Copel.) Price
Crepidomanes megistostomum (Copel.)
Copel.
Crepidomanes pervenulosum (Alderew.)
Copel.
Hymenophyllum archboldii (Copel.) C. V.
Morton
Hymenophyllum batuense Rosenst.
Hymenophyllum bontocense Copel.
Hymenophyllum campanulatum Christ
Hymenophyllum copelandii C. V. Morton
Hymenophyllum edanoi (Copel.) C. V.
Morton
Hymenophyllum hieronymii (Brause) C. Chr.
Hymenophyllum levingei C. B. Clarke
Polyphlebium werneri (Rosenst.) Ebihara et
K. Iwats.
Distribution
Note
Korea, Japan
New Guinea (Papua New Guinea)
Taiwan
China (Guizhou, Yunnan)
CR in Taiwan
Japan (Hokkaido)
CR in Japan
S. Korea (Cheju-do Isl.)
CR in S. Korea
Japan
Japan
Japan
EN in Japan
VU in Japan
VU in Japan
China (Yunnan)
China (Guangxi, Yunnan)
Maluku Islands (Seram)
China (Yunnan)
China (Yunnan)
China (Yunnan)
Taiwan
Taiwan
Japan (Bonin Islands)
Borneo, Philippines (Luzon, Mindoro,
Panay, Mindanao)
Laos, Myanmar
China (Guangdong, Jiangxi, Hongkong)
New Guinea (Papua New Guinea)
Japan (Bonin Islands)
Taiwan, Philippines (Luzon)
CR in Taiwan, a hybrid
taxon
EN in Taiwan
VU in Japan
Hybrid between
O. japonica and
O. vachellii
EN in Japan
Borneo (E. Kalimantan)
Philippines (Samar, Leyte), Sulawesi
Thailand (Phitsanulok, Phangnga)
Borneo (E. Kalimantan), Maluku Islands
(Ambon)
New Guinea (Prov. Papua)
Sumatra
Philippines (Luzon), Thailand (Nakhon Si
Thammarat)
Philippines (Panay, Negros)
New Guinea
Philippines (Mindoro, Luzon)
New Guinea (Papua New Guinea)
N. Indian Himalaya (Sikkim), Bhutan,
Tibet, China
New Guinea (Papua New Guinea)
DD in Thailand
Known only from the
type
Narrowly Distributed Asian Pteridophytes
Table 1.
(Continued)
Name
Hymenophyllaceae (continued)
Trichomanes wallii Thwaites ex Trimen
Gleicheniaceae
Dicranopteris clemensiae Holttum
Dicranopteris seramensis M. Kato
Diplopterygium angustilobum (Holttum)
Parris, comb. nov.
Diplopterygium clemensiae (Copel.) Parris,
comb. nov.
Diplopterygium deflexum (Holttum) Parris,
comb. nov.
Gleichenia peltophora Copel. var. schizolepis
C. Chr. ex Holttum
Gleichenia peltophora Copel. var. seramensis
M. Kato
Sticherus reflexipinnulus (C. Chr.) Copel.
Dipteridaceae
Dipteris lobbiana (Hook.) Moore
Matoniaceae
Phanerosorus major Diels
Marsileaceae
Marsilea scalaripes D. M. Johnson
Salviniaceae
Azolla japonica (Franch. et Sav.) Franch. et
Sav. ex Nakai
Plagiogyriaceae
Plagiogyria egenolfioides (Baker) Copel.
var. egenolfioides
Cyatheaceae
Cyathea acanthophora Holttum
Cyathea acuminata Copel.
Cyathea albosetacea (Scott. ex Bedd.)
Copel.
Cyathea alleniae Holttum
Cyathea angustipinna Holttum
Cyathea apoensis Copel.
Cyathea ascendens Domin
Cyathea atropurpurea Copel.
Cyathea binayana M. Kato
Cyathea binuangensis Alderw.
Cyathea bisquamata M. Kato
Cyathea brachyphylla Holttum
Cyathea callosa Christ
Cyathea carrii Holttum
Cyathea catillifera Holttum
Cyathea caudata (J. Sm. ex Hook.) Copel.
Cyathea christii Copel. (including Cyathea
negrosiana Christ)
97
Distribution
Note
Sri Lanka
Borneo (Sabah)
Maluku Islands (Seram)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
Borneo (Sarawak)
Maluku Islands (Seram)
New Guinea (Papua New Guinea)
Philippines (Negros Oriental), Borneo,
Peninsular Malaysia
Maluku Islands (Seram), New Guinea
(Prov. Papua)
Peninsular Malaysia
Japan
VU in Japan
Borneo (Sarawak)
Borneo (Sabah)
Philippines (Northern Samar, Capiz)
Nicobar Islands
Peninsular Malaysia
Borneo (Sarawak)
Philippines (Aurora, Laguna, Quezon,
Negros, Mindoro, Davao, Misamis
Occidental)
New Guinea (Papua New Guinea)
Philippines (Oriental Mindoro, Agusan
del Norte, Davao)
Maluku Islands (Seram)
Philippines (Northern Quezon, Tawitawi)
Maluku Islands (Seram)
Borneo (Sabah)
Philippines (Kalinga, Benguet, Bataan,
Laguna, Quezon, Camarines Sur, Catanduanes, Palawan, Camiguin)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
Philippines (Luzon, Mindoro, Palawan,
Negros, Mindanao)
Philippines (Negros Oriental, Davao
Oriental, Davao del Sur, North Cotabato,
Bukidnon)
EN in the Philippines
EN in the Philippines
EN in the Philippines
EN in the Philippines
EN in the Philippines
EN in the Philippines
EN in the Philippines
Atsushi Ebihara et al.
98
Table 1.
Name
Cyatheaceae (continued)
Cyathea cinerea Copel.
(Continued)
Distribution
Philippines (Agusan, Misamis Occidental)
Note
EN in the Philippines,
but known only from
the type
Cyathea coactilis Holttum
Cyathea coriacea M. Kato
Cyathea croftii Holttum
Cyathea cucullifera Holttum
Cyathea curanii Copel.
New Guinea (Papua New Guinea)
Maluku Islands (Seram)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
Philippines (Quezon)
Cyathea discophora Holttum
Cyathea edanoi Copel.
Cyathea eriophora Holttum
Cyathea excavata Holttum
Cyathea ferruginea Christ
Cyathea fuliginosa (Christ) Copel.
Borneo (Sabah)
Philippines (Cagayan, Quezon)
EN in the Philippines
New Guinea (Papua New Guinea)
Peninsular Malaysia
Philippines (Palawan, Negros, ?Luzon)
EN in the Philippines
Philippines (Luzon, Biliran, Leyte,
EN in the Philippines
Negros, Mindanao)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
Philippines (Mountain Province, Benguet, EN in the Philippines
Nueva Vizcaya, Mindoro)
Borneo (Sabah)
Philippines (Luzon, Panay, Negros,
EN in the Philippines
Mindanao)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
Philippines (Luzon, Catanduanes, Mindoro, EN in the Philippines
Samar, Biliran, Panay, Negros, Mindanao)
Philippines (Sibuyan) +
VU in the Philippines
Borneo (Sabah)
Maluku Islands (Seram)
New Guinea (Papua New Guinea)
Philippines (Benguet, ?Bukidnon)
EN in the Philippines
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
Philippines (Catanduanes)
CR in the Philippines
New Guinea (Papua New Guinea)
Maluku Islands (Seram)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
Philippines (Sibuyan)
VU in the Philippines
Japan (Bonin Islands)
VU in Japan
Maluku Islands (Seram)
New Guinea (Papua New Guinea)
Philippines (Ifugao, Mountain Province,
VU in the Philippines
Benguet, Aurora, Nueva Viscaya, Laguna,
Quezon, Camarines Sur, Sorsogon,
Bukidnon, Camiguin, Davao Oriental,
Misamis Occidental)
Maluku Islands (Seram)
New Guinea (Papua New Guinea)
Maluku Islands (Seram)
New Guinea (Papua New Guinea)
Philippines (Ilocos Norte, Isabela, Aurora, VU in the Philippines
Northern Quezon, Tawi-Tawi, Bukidnon)
New Guinea (Papua New Guinea)
Philippines (Zamboanga)
VU in the Philippines
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
Cyathea fusca Baker
Cyathea glaberrima Holttum
Cyathea gregaria (Brause) Domin.
Cyathea halconensis Christ
Cyathea havilandii Baker
Cyathea heterochlamydea Copel.
Cyathea hunsteiniana Brause
Cyathea inaequalis Holttum
Cyathea integra J.Sm. ex Hook.
Cyathea latipinnula Copel.
Cyathea longipes Copel.
Cyathea mapahuensis M. Kato
Cyathea marginata (Brause) Domin.
Cyathea masapilidensis Copel.
Cyathea media W. H. Wagner et Grether
Cyathea mesosora Holttum
Cyathea microchlamys Holttum
Cyathea moseleyi Baker
Cyathea murkelensis M. Kato
Cyathea nigropaleata Holttum
Cyathea nothofagorum Holttum
Cyathea obliqua Copel.
Cyathea ogurae (Hayata) Domin
Cyathea ohaensis M. Kato
Cyathea parvipinna Holttum
Cyathea philippinensis Baker
Cyathea pukuana M. Kato
Cyathea pycnoneura Holttum
Cyathea ramispinoides M. Kato
Cyathea recurvata (Brause) Domin
Cyathea robinsonii Copel.
Cyathea rubella Holttum
Cyathea rufopannosa Christ
Cyathea scandens (Brause) Domin
Cyathea schlechteri (Brause) Domin
EN in the Philippines,
known only from the type
Narrowly Distributed Asian Pteridophytes
Table 1.
Name
Cyatheaceae (continued)
Cyathea setulosa Copel.
Cyathea sibuyanensis Copel.
Cyathea stipitipinnula Holttum
Cyathea subtripinnata Holttum
Cyathea tenuicaulis Domin
Cyathea ternatea Alderw.
Cyathea tripinnatifida Roxb.
Cyathea vandeusenii Holttum
Cyathea womersleyi Holttum
Cyathea zamboangana Copel.
Dicksoniaceae
Dicksonia archboldii Copel.
Lindsaeaceae
Lindsaea eberhardtii (Christ) K. U. Kramer
Lindsaea hewittii Copel.
Lindsaea jamesonioides Baker
Lindsaea malabarica (Bedd.) Baker
Lindsaea obscura Brause
Lindsaea papuana Copel.
Lindsaea repanda Kunze
Lindsaea sarawakensis K. U. Kramer
Lindsaea securifolia Presl ex Goldm. var.
kusukusensis (Hayata) Shieh
Lindsaea tenera Dryand.
Lindsaea venusta Kaulf. ex Kuhn
Odontosoria minutula (Sa. Kurata) Ebihara,
comb. nov.
Odontosoria quadripinnata Lehtonen
Odontosoria veitchii (Baker) Parris
Odontosoria yaeyamensis (S. J. Lin, M.
Kato et K. Iwats.) Ebihara, comb. nov.
Tapeinidium buniifolium K. U. Kramer
Dennstaedtiaceae
Dennstaedtia melanostipes Ching
Histiopteris incisa (Thunb.) J. Sm. var.
seramensis M. Kato
Microlepia caudigera T. Moore
Microlepia matthewii Christ
Microlepia membranacea B. S. Wang
Microlepia modesta Ching
Microlepia rheophila K. Iwats. et M. Kato
Paesia taiwanensis W. C. Shieh
Pteridaceae
Adiantum chienii Ching
Adiantum davidii Franch. var. longispinum
Ching
Adiantum fengianum Ching
Adiantum formosanum Tagawa
Adiantum fragiliforme C. Chr.
99
(Continued)
Distribution
Philippines (Kalinga, Isabela, Quezon)
Philippines (Sibuyan)
Borneo (Sabah)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
Maluku Islands (Seram, Ternate)
Maluku Islands (Seram, Ambon)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
Philippines (Zamboanga, Agusan)
New Guinea (Prov. Papua)
China (Hainan)
Borneo (Sarawak)
Borneo (Sabah)
C. and S. India (Madhya Pradesh, ?Andhra
Pradesh, Karnataka, Tamil Nadu, Kerala)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
Japan (Bonin Islands)
Borneo (Sarawak)
Taiwan
Andaman Islands, Nicobar Islands
Sri Lanka, S. India
Japan (Amami Isl.)
Note
VU in the Philippines
VU in the Philippines,
known only from the type
VU in the Philippines
VU in New Guinea
(Prov. Papua)
VU in Japan
CR in Taiwan
Taxonomically somewhat uncertain as to
whether it is really
specifically distinct
from L. bouillodii
CR in Japan
New Guinea (Papua New Guinea)
Borneo (Sabah)
Japan (Iriomote Isl.)
New Guinea (Papua New Guinea)
China (Yunnan)
Maluku Islands (Seram)
Bhutan, N.E. India (Arunachal Pradesh,
?Meghalaya)
China (Guangdong, Guangxi)
China (Guangdong, Hunan)
China (Anhui, Jiangxi, Zhejiang)
Borneo (E. Kalimantan)
Taiwan
China (Guangdong, Hunan)
China (Chongqing, Guizhou, Sichuan,
Yunnan)
China (Yunnan)
Taiwan
Thailand (Trat), Cambodia
CR in Taiwan
VU in Taiwan
DD in Thailand and
Cambodia
Atsushi Ebihara et al.
100
Table 1.
Name
Pteridaceae (continued)
Adiantum gomphophyllum Baker
Adiantum juxtapositum Ching
Adiantum lianxianense Ching et Y. X. Lin
Adiantum meishanianum F. S. Hsu ex Y. C.
Liu et W. L. Chiou
Adiantum membranifolium S. Linds. et
Suksathan
Adiantum mindanaoense Copel.
Adiantum ogasawarense Tagawa
Adiantum reniforme L. var. sinense Y. X. Lin
Adiantum roborowskii Maxim. var. taiwanianum (Tagawa) W. C. Shieh
Adiantum siamense Tagawa et K. Iwats.
Adiantum thongthamii Suksathan
Aleuritopteris duthiei (Baker) Ching
Aleuritopteris ebenipes X. C. Zhang
Aleuritopteris squamosa (C. Hope et C. H.
Wright) Ching
Aleuritopteris thwaitesii (Mett. ex Kuhn)
Saiki
Antrophyum lancifolium Rosenst. ex M.
Kato
Antrophyum nanum Fée
Antrophyum vittarioides Baker
Austrogramme boerlageana (Alderw.)
Hennipman
Cerosora chrysosorum (Baker) Domin
Cheilanthes delicatula Tagawa et K. Iwats.
Cheilanthes fragilis Hook.
Cheilanthes krameri Franch. et Sav.
Cheilosoria patula (Baker) P. S. Wang
Cheilanthes pseudoargentea (S. K. Wu) K.
Iwats.
Cheilanthes siamensis (S. K. Wu) K.Iwats.
Coniogramme gracilis Ogata
Coniogramme macrophylla (Blume) Hieron.
Haplopteris hainanensis (C. Chr. ex Ching)
E. H. Crane
Haplopteris mediosora (Hayata) X. C.
Zhang
Haplopteris winitii (Tagawa et K. Iwats.) S.
Linds.
Pteris angustipinna Tagawa
Pteris austrosinica (Ching) Ching
Pteris cadieri Christ var. hainanensis
(Ching) S. H. Wu
Pteris calocarpa (Copel.) M. G. Price.
Pteris clemensiae Copel.
Pteris endoneura M. G. Price
(Continued)
Distribution
Thailand (Phangnga, Satun)
China (Fujian, Guangdong, Hunan,
Jiangxi, Zhejiang)
China (Guangdong, Guizhou, Hunan)
Taiwan
Thailand (Chiang Mai [Doi Chiang Dao,
Doi Ang Khang], Chiang Rai [Mae Sai])
Philippines (San Ramon, Zamboanga)
Japan (Bonin Islands)
China (Chongqing, Hubei, Sichuan)
Taiwan
Note
VU in Thailand
CR in Taiwan, a hybrid
taxon
VU in Thailand
VU in the Philippines
EN in Japan
VU in Taiwan
Thailand (Nakhon Si Thammarat, Krabi)
Thailand (Trat [Koh Chang])
N.W. India (Uttarakhand), Nepal, Bhutan,
Tibet
China (Guizhou)
China (Hainan, Yunnan)
EN in Thailand
VU in Thailand
Sri Lanka, S. India (Tamil Nadu)
Genus uncertain
Maluku Islands (Seram)
Philippines (Luzon, Palawan, Mindanao),
Indonesia (Java, Seram)
Borneo, Vietnam
Maluku Islands (Seram, Ambon)
Borneo (Sarawak)
Thailand (Chiang Mai)
Thailand (Chiang Mai, Tak), Myanmar
Japan
China (Chongqing, Guizhou, Hubei,
Hunan, Sichuan)
Thailand (Chiang Mai)
DD in Thailand
EN in Japan
DD in Thailand
Thailand (Nan, Lampang)
Japan (Amami Isl.)
China (Hainan, Yunnan)
China (Guangdong, Hainan, Yunnan)
DD in Thailand
CR in Japan
Taiwan
VU in Taiwan
Thailand (Chiang Mai, Chiang Rai,
Lampang)
Taiwan
China (Guangdong, Guangxi, Hunan,
Jiangxi)
China (Guangdong, Guangxi, Hainan)
DD in Thailand
Philippines (Zamboanga)
Borneo (Sabah)
Philippines (Quezon, Western Samar,
Camarines Norte)
Pteris ensiformis Burmann var. rheophila M. Borneo (E. Kalimantan)
Kato, Darnaedi et K. Iwats.
Pteris geminata Wall. ex J. Agardh
S. India (Tamil Nadu)
CR in Taiwan
CR in the Philippines
EN in the Philippines
Narrowly Distributed Asian Pteridophytes
Table 1.
(Continued)
Name
Pteridaceae (continued)
Pteris griffithii Hook.
Pteris guangdongensis Ching
Pteris hui Ching
Pteris giasii Fraser-Jenk.
Pteris kawabatae Sa. Kurata
Pteris lepidopoda M. Kato et K.U. Kramer
Pteris mertensioides Willd. subsp. polylepis
M. Kato
Pteris montis-wilhelminae Alston
Pteris nakasimae Tagawa
Pteris pachysora (Copel.) M. G. Price
Pteris reptans T. G. Walker
Pteris tarandus M. Kato et K. U. Kramer
Pteris viridissima Ching
Pteris wallichiana Agardh var. obtusa S. H. Wu
Pteris wulaiensis C. M. Kuo
Pteris xichouensis W. M. Chu et Z. R. He
Pteris yamatensis (Tagawa) Tagawa
Pteris zippellii (Miquel) M. Kato et K. U.
Kramer
Sinopteris grevilleoides (Christ) C. Chr. et
Ching
Syngramma minima Holttum
Syngramma valleculata (Baker) C. Chr.
Taenitis brooksii Copel.
Taenitis dimorpha Holttum
Taenitis hosei (Baker) Holttum
Taenitis intermedia M. Kato
Taenitis mediosora M. Kato
Taenitis trilobata Holttum
Taenitis vittarioides Holttum subsp. moluccana M. Kato
Cystopteridaceae
Cystoathyrium chinense Ching
Aspleniaceae
Asplenium adnatum Copel.
Asplenium cardiophyllum (Hance) Baker
Asplenium daghestanicum Christ subsp.
hunzanum (Reichst. et Fraser-Jenk.)
Fraser-Jenk.
Asplenium daghestanicum Christ subsp.
iskardense (Reichst.) Fraser-Jenk.
Asplenium dichotomum Hook.
Asplenium disjunctum Sledge
Asplenium exiguum Bedd. subsp. exiguum
Asplenium gardneri Baker
Asplenium humbertii Tardieu
Asplenium kinabaluense Holttum
Asplenium klossii C. Chr.
Asplenium micantifrons (Tuyama) Tuyama
ex H. Ohba
101
Distribution
N. E. India (Arunachal Pradesh),
Myanmar; ?China
China (Guangdong, Guangxi)
China (Guangxi)
S. E. Bangladesh (Chittagong Hills)
Japan (Yakushima Isl.)
Maluku Islands (Seram)
Maluku Islands (Seram)
New Guinea (Prov. Papua)
Japan (Kyushu)
Philippines (Zamboanga)
Sri Lanka, S. India (Kerala)
Maluku Islands (Seram)
China (Guizhou, Yunnan)
China (Jiangxi, Sichuan, Yunnan)
Taiwan
China (Yunnan)
Japan
Maluku Islands (Seram), New Guinea
Note
So far known to S.E.
Bangladesh, but almost
certainly occurring
nearby in W. Myanmar
CR in Japan
VU in New Guinea
(Prov. Papua)
EN in Japan
CR in the Philippines
CR in Taiwan
CR in Japan
China (Sichuan, Yunnan)
Peninsular Malaysia
Borneo (Sarawak)
Borneo (Sarawak)
Peninsular Malaysia
Borneo (Sarawak)
Maluku Islands (Ambon)
Maluku Islands (Seram)
Borneo (Sarawak)
Maluku Islands (Ambon, Seram)
China (Sichuan)
Extinct
China (Guangdong, Hongkong, Yunnan)
China (Hainan), Japan (Bonin Islands,
Ryukyu)
N. Pakistan, ?N. W. India (Uttarakhand,
Nanda Devi)
N. Pakistan
Borneo (Sabah, Sarawak)
Sri Lanka
S. India (Kerala, Tamil Nadu)
Sri Lanka
Thailand (Chiang Mai), N. Vietnam
Borneo (Sabah)
Borneo (Sabah)
Japan (Bonin Islands)
Not found since the
type collection in the
first half of the 19th
Century
VU in Japan
Atsushi Ebihara et al.
102
Table 1.
Name
Aspleniaceae (continued)
Asplenium oligophlebium Baker var. iezimaense
(Tagawa) Tagawa
Asplenium pachychlamys C. Chr.
Asplenium rebeccae Fraser-Jenk. et Wangdi
Asplenium rivulare Fraser-Jenk.
Asplenium siamense Tagawa et K. Iwats.
Asplenium speluncae Christ
Asplenium subaquatile Cesati
Asplenium (Hymenasplenium) unilaterale
Lam. var. saxicola M. Kato et K. Iwats.
Ceterachopsis qiujiangensis Ching et S. H.
Fu ex Ching et S. H. Wu
Hymenasplenium inthanonense N. Murak. et
J. Yokoy.
Thelypteridaceae
Chingia bewaniensis Holttum
Chingia paucipaleata Holttum
Chingia pricei Holttum
Chingia supraspinigera (Rosenst.) Holttum
Chingia urens Holttum
Christella boninensis (Kodama ex Koidz.)
Holttum
Christella minima Holttum
Christella peekelii (Alderw.) Holttum
Christella subdentata Holttum
Coryphopteris andersonii Holttum
Coryphopteris andreae Holttum
Coryphopteris athyrioides Holttum
Coryphopteris borealis Holttum
Coryphopteris coriacea (Brause) Holttum
Coryphopteris engleriana (Brause) Holttum
Coryphopteris gymnopoda (Baker) Holttum
var. bintangensis Holttum
Coryphopteris gymnopoda (Baker) Holttum
var. humilis Holttum
Coryphopteris inopinata Holttum
Coryphopteris lauterbachii (Brause)
Holttum
Coryphopteris ledermannii (Hieron.)
Holttum
Coryphopteris meiobasis Holttum
Coryphopteris pectiniformis (C. Chr.)
Holttum var. pectiniformis
Coryphopteris pectiniformis (C. Chr.)
Holttum var. hirsuta Holttum
Coryphopteris plumosa (C. Chr.) Holttum
Coryphopteris seramensis M. Kato
Coryphopteris squamipes (Copel.) Holttum
Coryphopteris stereophylla (Alderw.)
Holttum
Coryphopteris tahanensis Holttum
Coryphopteris unidentata (Bedd.) Holttum
Coryphopteris viscosa (Baker) Holttum var.
poiensis Holttum
Cyclogramma chunii (Ching) Tagawa
(Continued)
Distribution
Japan (Ryukyu)
Borneo (Sabah)
Bhutan
S. India (Kerala, Tamil Nadu)
Thailand (Loei)
China (Guangdong, Guangxi, Guizhou,
Hunan, Jiangxi)
Borneo (Sarawak, ?Kalimantan)
Maluku Islands (Seram)
Note
EN in Japan
DD in Thailand
China (Yunnan)
Thailand (Chiang Mai)
New Guinea (Papua New Guinea)
Philippines (Quezon)
Philippines (Benguet, Misamis Occidental)
New Guinea (Papua New Guinea)
Philippines (Laguna, Bukidnon)
Japan (Bonin Islands)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
Philippines (Cavite, Lubang Island,
Mindoro Occidental)
Borneo (Sarawak)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
Philippines (Ifugao, Benguet)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
Peninsular Malaysia
VU in Thailand
VU in the Philippines
VU in the Philippines
EN in the Philippines
EN in Japan
VU in the Philippines
CR in the Philippines
Peninsular Malaysia
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
Peninsular Malaysia
Peninsular Malaysia
Borneo (Sarawak)
Maluku Islands (Seram)
Philippines (Zambales, Benguet, Davao
del Sur, South Cotabato, Bukidnon)
New Guinea (Prov. Papua)
Peninsular Malaysia
Peninsular Malaysia
Borneo (Sarawak)
China (Guangdong, Guangxi)
VU in the Philippines
VU in New Guinea
(Prov. Papua)
Narrowly Distributed Asian Pteridophytes
Table 1.
Name
Thelypteridaceae (continued)
Cyclosorus distinctus (Copel.) Copel.
(=Amphineuron distinctum (Copel.)
Holttum, nom. illeg.)
Cyclosorus thailandicus S. Linds.
Dictyocline mingchegensis Ching
Mesophlebion falcatilobum Holttum
Mesophlebion oligodictyon (Baker) Holttum
Parathelypteris subimmersa (Ching) Ching
Plesioneuron altum (Brause) Holttum
Plesioneuron bipinnatum (Copel.) Holttum
Plesioneuron costulisorum (Copel.) Holttum
Plesioneuron croftii Holttum
Plesioneuron cystodioides Holttum
Plesioneuron dryas Holttum
Plesioneuron falcatipinnulum (Copel.)
Holttum
Plesioneuron fuchsii Holttum
Plesioneuron fulgens (Brause) Holttum
Plesioneuron kundipense Holttum
Plesioneuron medusella Holttum
Plesioneuron murkelense M. Kato
Plesioneuron notabile (Brause) Holttum
Plesioneuron platylobum Holttum
Plesioneuron pullei Holttum
Plesioneuron sandsii Holttum
Plesioneuron saxicola M. Kato
Plesioneuron stenura Holttum
Plesioneuron translucens Holttum var.
translucens
Plesioneuron translucens Holttum var.
seramense M. Kato
Plesioneuron varievestitum (C. Chr.)
Holttum
Plesioneuron wantotense (Copel.) Holttum
Plesioneuron wariense (Copel.) Holttum
Plesioneuron woodlarkense (Copel.)
Holttum
Pneumatopteris angusticaudata Holttum
Pneumatopteris boridensis Holttum
Pneumatopteris brooksii (Copel.) Holttum
Pneumatopteris caudata (Holttum) Holttum
Pneumatopteris eburnea Holttum
Pneumatopteris finisterrae (Brause) Holttum
Pneumatopteris jermyii Holttum
Pneumatopteris latisquamata Holttum
Pneumatopteris micropaleata Holttum
Pneumatopteris mingendensis (Gilli)
Holttum
Pneumatopteris nephrolepioides (C. Chr.)
Holttum
Pneumatopteris papuana Holttum
Pneumatopteris patentipinna Holttum
Pneumatopteris pergamacea Holttum
Pneumatopteris psilophylla Holttum
103
(Continued)
Distribution
New Guinea (Papua New Guinea)
Thailand (Phetchaburi)
China (Fujian, Guangdong, Jiangxi,
Zhejiang)
Borneo (Sarawak)
Borneo (Sarawak, Brunei, Kalimantan)
China (Hainan)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
Borneo (Sabah)
Maluku Islands (Seram), New Guinea
(Papua New Guinea)
New Guinea (Papua New Guinea)
Borneo (Sarawak)
Maluku Islands (Seram)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
Maluku Islands (Seram), New Guinea
New Guinea (Papua New Guinea)
Maluku Islands (Seram)
New Guinea (Papua New Guinea)
Maluku Islands (Ambon)
Maluku Islands (Seram)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
Borneo (Sarawak)
New Guinea (Papua New Guinea)
Borneo (Sarawak)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
Borneo (Sabah)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
Borneo (Sarawak)
Note
The genus Amphineuron
Holttum is a later
homonym of Amphineurion (A. DC.) Pichon,
Apocynaceae (Brummitt,
2007)
DD in Thailand
Atsushi Ebihara et al.
104
Table 1.
Name
Thelypteridaceae (continued)
Pneumatopteris regis (Copel.) Holttum
Pneumatopteris walkeri Holttum
Pronephrium acanthocarpum (Copel.)
Holttum
Pronephrium aquatiloides (Copel.) Holttum
Pronephrium brauseanum Holttum
Pronephrium giluwense Holttum
Pronephrium hewittii (Copel.) Holttum
Pronephrium hosei (Baker) Holttum
Pronephrium longipetiolatum (K. Iwats.)
Holttum
Pronephrium micropinnatum Holttum
Pronephrium millarae Holttum
Pronephrium peltatum (Alderw.) Holttum
var. persitiferum Holttum
Pronephrium thysanoides Holttum
Pseudophegopteris cyclocarpa Holttum
Pseudophegopteris kinabaluensis Holttum
Pseudophegopteris yunkweiensis (Ching)
Ching
Sphaerostephanos alpinus Holttum
Sphaerostephanos alticola Holttum
Sphaerostephanos aquatilis (Copel.) Holttum
Sphaerostephanos austerus (Brause) Holttum
Sphaerostephanos carrii Holttum
Sphaerostephanos dimidiolobatus Holttum
Sphaerostephanos efogensis Holttum
Sphaerostephanos ekutiensis Holttum
Sphaerostephanos flavoviridis Holttum
Sphaerostephanos gymnorachis Holttum
Sphaerostephanos hispidifolius (Alderw.)
Holttum
Sphaerostephanos isomorphus Holttum
Sphaerostephanos lithophyllus (Copel.)
Holttum
Sphaerostephanos lobangensis (C. Chr.)
Holttum
Sphaerostephanos mengenianus Holttum
Sphaerostephanos mjobergii Holttum
Sphaerostephanos muluensis Holttum
Sphaerostephanos nakaikei Holttum
Sphaerostephanos novaebritanniae Holttum
Sphaerostephanos obtusifolius (Rosenst.)
Holttum
Sphaerostephanos omatianus Holttum
Sphaerostephanos plurivenosus Holttum
Sphaerostephanos potamios Holttum
Sphaerostephanos pterosporus (Alderw.)
Holttum var. altilobus Holttum
Sphaerostephanos pullenii Holttum
Sphaerostephanos reconditus Holttum
Sphaerostephanos semicordatus Holttum
Sphaerostephanos squamatellus Holttum
Sphaerostephanos sudesticus Holttum
Sphaerostephanos telefominicus Holttum
Sphaerostephanos trimetralis Holttum
Sphaerostephanos uaniensis Holttum
(Continued)
Distribution
Note
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
Borneo (Sarawak)
Borneo (Sarawak)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
Borneo (Sarawak)
Borneo (Sarawak), Philippines (Zamboanga, Basilan)
Taiwan
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
Borneo (Sabah, Sarawak)
Borneo (Sarawak)
New Guinea (Papua New Guinea)
Borneo (Sabah)
China (Guangxi, Guizhou, Hunan,
Yunnan)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
Borneo (Sabah)
Borneo (Sarawak, Kalimantan)
New Guinea (Papua New Guinea)
Borneo (Sabah)
Borneo (Sabah)
New Guinea (Papua New Guinea)
Borneo (Sarawak)
Borneo (Sarawak)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
Peninsular Malaysia
New Guinea (Papua New Guinea)
Borneo (Sarawak)
New Guinea (Papua New Guinea)
Borneo (Sabah)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
CR in Taiwan
Narrowly Distributed Asian Pteridophytes
Table 1.
Name
Thelypteridaceae (continued)
Sphaerostephanos uniauriculatus (Copel.)
Holttum
Sphaerostephanos wauensis Holttum
Sphaerostephanos woitapensis Holttum
Stegnogramma cyrtomioides (C. Chr.) Ching
Stegnogramma gymnocarpa (Copel.) K.
Iwats. subsp. amabilis (Tagawa) K. Iwats.
Thelypteris didymochlaenoides (C .B.
Clarke) Holttum
(Continued)
Distribution
Note
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
China (Chongqing, Guizhou, Hubei,
Hunan, Sichuan)
Japan
N. E. India (Meghalaya)
Thelypteris kingii C. F. Reed
N. India (Sikkim)
Thelypteris kurzii (Holttum) Fraser-Jenk.
Nicobar Islands
Thelypteris namburensis (Bedd.) C. F. Reed
N. E. India (Assam, Arunachal Pradesh)
Thelypteris parasitica (L.) Fosberg subsp.
manickirudorum Fraser-Jenk. et Benniamin
Thelypteris sledgei Fraser-Jenk.
Thelypteris srilankensis Panigrahi
Thelypteris thwaitesii (Hook.) C. F. Reed
S. India (Tamil Nadu)
Sri Lanka
Sri Lanka, ?Nicobar Islands
Sri Lanka
Thelypteris zeylanica Ching
Sri Lanka, Arunachal Pradesh
Cheilanthopsis indusiosa (Christ) Ching
Woodsia okamotoi Tagawa
Rhachidosoraceae
Rhachidosorus pulcher (Tagawa) Ching
Rhachidosorus truncatus Ching
Blechnaceae
Blechnum acanthopodum T. C. Chambers et
P. A. Farrant
Blechnum puniceum T. C. Chambers, P. J.
Edwards et R. J. Johns
105
China (Sichuan, Yunnan)
Taiwan
NT in Japan
As known so far
endemic to N.E. India,
but very probably also
occuring in Myanmar
and perhaps also China
As known at present
endemic to Sikkim, but
likely to occur in Tibet
and China
Very rare or perhaps
extinct
As known so far
endemic to N. E. India,
but likely to occur in
Myanmar and/or China
Apparently endemic to
S. India
Reports from S. India
(Kerala, very rare),
refer to the very similar
sterile hybrid,
T. × insularis.
Reported from Assam
by Chandra et al.
(2008) in error for an
unidentified species
near to T. triphylla
Although this may be a
good species, it is
conceivable that these
plants might only be
depauperate forms of
T. calcarata and
T. caudipinna respectively, and require
further investigative
study
CR in Taiwan
Yunnan, Taiwan
China (Guangxi, Guizhou, Yunnan)
New Guinea (Papua New Guinea)
New Guinea (Prov. Papua)
EN in New Guinea
(Prov. Papua)
Atsushi Ebihara et al.
106
Table 1.
Name
Athyriaceae
Allantodia dushanensis Ching ex W. M. Chu
et Z. R. He
Allantodia heterocarpa (Ching) Ching
(Continued)
Distribution
China (Guangxi, Guizhou, Hunan)
Athyrium hainanense Ching
Athyrium kenzo-satakei Sa. Kurata
Athyrium khasimontanum Fraser-Jenk.
China (Chongqing, Guizhou, Hunan,
Sichuan)
China (Guangdong, Hunan)
China (Hainan)
China (Chongqing, Guizhou, Hubei,
Hunan, Sichuan, Yunnan)
Borneo (Sabah)
Taiwan
Borneo (Sabah)
China (Guangxi, Guizhou, Hunan,
Jiangxi, Zhejiang)
China (Hainan)
Japan (Kyushu)
N. E. India (Meghalaya)
Athyrium kumaonicum Holttum ex Punetha
N. W. India (Uttarakhand)
Athyrium masamunei Seriz.
Athyrium minimum Ching
Athyrium neglectum Seriz.
Athyrium palustre Seriz.
Athyrium yakusimense Tagawa
Cornopteris seramensis M. Kato
Deparia biserialis (Baker) M. Kato
Deparia bonincola (Nakai) M. Kato
Deparia hainanensis (Ching) R. Sano
Deparia lancea (Thunb.) Fraser-Jenk. var.
rheophila (M. Kato, Darnaedi et K.
Iwats.) M. Kato, comb. nov.
Deparia macdonellii (Bedd.) M. Kato
Japan (Yakushima Isl.)
Taiwan
Japan
Japan
Japan (Yakushima Isl.)
Maluku Islands (Seram)
Borneo (Sabah)
Japan (Bonin Islands)
China (Hainan)
Borneo (E. Kalimantan)
Allantodia metcalfii (Ching) Ching
Allantodia wangii Ching
Athyriopsis erecta Z. R. Wang
Athyrium amoenum C. Chr.
Athyrium auriculatum Seriz.
Athyrium clemensiae Copel.
Athyrium elongatum Ching
VU in Taiwan
CR in Japan
As known so far
endemic to N.E. India
Range imperfectly
known. As known so
far endemic to N.W.
India
CR in Japan
EN in Taiwan
CR in Japan
EN in Japan
CR in Japan
NT in Japan
Pakistan, N.W. India (Jammu , Kashmir,
Himachal Pradesh)
Japan (Kyushu)
EN in Japan
Japan
EN in Japan
Sri Lanka
Very rare or extinct
Maluku Islands (Seram)
Borneo
Peninsular Malaysia
Borneo (Sabah)
S. India (Tamil Nadu, Agastyamalai Hills)
Deparia minamitanii Seriz.
Deparia otomasui (Sa. Kurata) Seriz.
Deparia zeylanica (Hook.) M. Kato
Diplazium aculeatum Alderw.
Diplazium aequibasale (Baker) C. Chr.
Diplazium angustipinna (Holttum) Holttum
Diplazium atrosquamosum C. Chr.
Diplazium austrosylvaticum Fraser-Jenk. et
Benniamin
Diplazium beddomei C. Chr.
Sri Lanka, S. India (?Kerala, Tamil Nadu)
Diplazium burmanicum Ching ex W. M. Chu Arunachal Pradesh, Myanmar
et Z. R. He
Diplazium ceramicum (Miq.) C. Chr.
Maluku Islands (Seram)
Diplazium cognatum (Hieron.) Sledge
Sri Lanka S. India (Kerala, Tamil Nadu)
Diplazium costulisorum (Copel.) C. Chr.
Philippines (Davao, Mt. Apo)
Diplazium cultratum C. Presl
Philippines (Biliran, Laguna, Quezon,
Camarines Norte)
Philippines (Nueva Viscaya, Laguna)
Maluku Islands (Seram)
Diplazium egenolfioides M. G. Price.
Diplazium esculentoides M. Kato
Note
Closely related to or a
synonym of
D. leptophyllum
VU in the Philippines,
but known only from
two type collections
VU in the Philippines
EN in the Philippines
Narrowly Distributed Asian Pteridophytes
Table 1.
Name
Athyriaceae (continued)
Diplazium flavoviride Alston
Diplazium furculicolum Alderw.
Diplazium insigne Holttum
Diplazium kunstleri Holttum
Diplazium longicarpum Kodama
Diplazium moluccanum Rosenst.
Diplazium paradoxum Fée
Diplazium pariens Copel.
Diplazium propinquum (Copel.) Alderw.
Diplazium subtripinnatum Nakai
Diplazium travancoricum Bedd.
Diplazium tricholepis C. Chr.
Diplazium velutinum Holttum
Diplazium wahauense M. Kato, Darnaedi et
K. Iwats.
Hypodematiaceae
Hypodematium crenatum (Forssk.) Kuhn
subsp. fauriei (Kodama) K. Iwats.
Hypodematium crenatum (Forssk.) Kuhn
subsp. mehrae Fraser-Jenk.
Dryopteridaceae
Arachniodes cantilenae Sa. Kurata
Arachniodes haniffii (Holttum) Ching.
Arachniodes hiugana Sa. Kurata
Arachniodes nigrospinosa (Ching) Ching
Ataxipteris dianguiensis W. M. Chu et H. G.
Zhou
Bolbitis christensenii (Ching) Ching
Bolbitis hainanensis Ching et Chu H. Wang
Bolbitis hekouensis Ching
Bolbitis interlineata (Copel.) Ching
Bolbitis malaccensis (C. Chr.) Ching
Bolbitis medogensis (Ching et S. K. Wu)
S. Y. Dong
Bolbitis nitens Holttum
Bolbitis presliana (Fée) Ching
Bolbitis semicordata (Baker) Ching
Bolbitis tibetica Ching et S. K. Wu
Bolbitis tonkinensis (C. Chr. ex Ching) K.
Iwats.
Ctenitis calcicola M. Kato
Ctenitis clathrata M. Kato
Ctenitis coriacea M. Kato
Ctenitis dumrongii Tagawa et K. Iwats.
Ctenitis ferruginea (Bedd.) Ching
Ctenitis iriomotensis (H. Itô) Nakaike
Ctenitis mannii (C. Hope) Ching
Ctenitis microlepigera (Nakai) Ching
Ctenitis minutiloba Holttum
Ctenitis obtusiloba (Baker) Ching
107
(Continued)
Distribution
Maluku Islands (Seram), New Guinea
(Papua New Guinea)
Maluku Islands (Seram)
Peninsular Malaysia
Peninsular Malaysia
Japan (Bonin Islands)
Maluku Islands (Seram, Tanimbar)
Sri Lanka
Philippines (Bukidnon, Negros, Davao
Oriental, North Cotabato [Mt. Apo])
Philippines (Agusan)
Japan (Bonin Islands)
Sri Lanka, S. India (Kerala, Tamil Nadu)
Borneo (Sabah)
Peninsular Malaysia
Borneo (E. Kalimantan)
Japan
Note
EN in Japan
VU in the Philippines
CR in Japan
VU in Japan
N.W. India (Uttarakhand)
Japan (Kyushu)
Peninsular Malaysia
Japan (Kyushu)
China (Guangdong, Guangxi, Guizhou,
Hunan, Hongkong), Taiwan
China (Guangxi, Hainan, Yunnan)
EN in Japan
CR in Japan
China (Guangxi, Guizhou)
China (Hainan, Yunnan)
China (Hainan, Yunnan)
Borneo (Sarawak)
Peninsular Malaysia
Arunachal Pradesh, Tibet
Peninsular Malaysia
S. India (Karnataka, Kerala)
S. India (Kerala, Tamil Nadu)
Nepal, Arunachal Pradesh, Tibet
Thailand (Chiang Rai), N. Vietnam,
Cambodia
Maluku Islands (Seram)
Maluku Islands (Seram)
Maluku Islands (Seram)
Thailand (Chiang Mai, Chiang Rai)
S. India (Tamil Nadu)
Japan (Iriomote Isl.)
N. E. India (Assam State, ?Meghalaya),
?China
Japan (Bonin Islands)
Borneo (Sabah)
Sri Lanka
DD in Thailand
VU in Japan
As known so far
perhaps endemic to
N. E. India (if C. fulgens
is not synonymous), but
likely to occur in
Myanmar and/or China
CR in Japan
Atsushi Ebihara et al.
108
Table 1.
Name
Dryopteridaceae (continued)
Ctenitis paleolata Copel.
Ctenitis paucisora (Copel.) Copel.
Ctenitis pseudorhodolepis Ching et Chu H.
Wang
Ctenitis scabrosa (Kuntze) Ching
Ctenitis tabacifera (Alderw.) Ching
Ctenitis muluensis Holttum
Cyrtogonellum inaequalis Ching
Cyrtomidictyum basipinnatum (Baker)
Ching
Cyrtomidictyum conjunctum Ching
Cyrtomium chingianum P. S. Wang
Cyrtomium grossum Christ
Cyrtomium sinningense Ching et K. H.
Shing
Cyrtomium taiwanianum Tagawa
Cyrtomium urophyllum Ching
Cyrtomium wulingense S. F. Wu
Dryopteris anadroma Mitsuta, nom. nud.
Dryopteris angustifrons (T. Moore) Kuntze
Dryopteris anthracinisquama Miyam.
Dryopteris austroindica Fraser-Jenk.
Dryopteris deparioides (T. Moore) Kuntze
subsp. deparioides
Dryopteris deparioides (T. Moore) Kuntze
subsp. concinna C. Chr.
Dryopteris enneaphylla (Baker) C. Chr. var.
pseudosieboldii (Hayata) Tagawa et K.
Iwats.
Dryopteris gymnosora (Makino) C. Chr. var.
angustata H. Itô
Dryopteris hadanoi Sa. Kurata
Dryopteris insularis Kodama
Dryopteris khullarii Fraser-Jenk.
Dryopteris meghalaica Fraser-Jenk. et
Gibby
Dryopteris odontoloma (Bedd.) C. Chr.
Dryopteris permagna M. G. Price
Dryopteris rheophila Mitsuta ex Darnaedi,
M. Kato et K. Iwats.
Dryopteris shibipedis Sa. Kurata
(Continued)
Distribution
Philippines (Benguet)
N. E. India (Arunachal Pradesh), Borneo
China (Guangxi, Guizhou, Sichuan)
CR in the Philippines;
known only from the
type
S. India (Kerala, Tamil Nadu)
Maluku Islands (Seram, Ambon)
Borneo (Sarawak)
China (Chongqing, Guangxi, Guizhou)
China (Guangdong, Guangxi, Hunan,
Hongkong)
China (Jiangxi, Zhejiang)
China (Guizhou)
China (Guizhou, Yunnan)
China (Hunan)
Taiwan
China (Chongqing, Guangxi, Guizhou,
Hunan, Sichuan, Yunnan)
China (Hunan, Sichuan)
Japan (Yakushima Isl.)
China (Yunnan), Myanmar
Japan (Kyushu)
S. India (Tamil Nadu)
Sri Lanka
VU in Taiwan
VU in Japan
CR in Japan
Sri Lanka, S. India (Tamil Nadu)
Taiwan
NT in Taiwan
Japan (Yakushima Isl.)
CR in Japan
Japan
Japan (Bonin Islands)
N. W. India (Uttarakhand)
N. E. India (Manipur, Meghalaya),
Myanmar
S. India (Kerala, Tamil Nadu)
Philippines (Kalinga, Benguet, Mountain
Province, South Cotabato)
Thailand (Loei)
NT in Japan
VU in Japan
Japan (Kyushu)
Dryopteris sledgei Fraser-Jenk.
Dryopteris tsugiwoi Sa. Kurata
Dryopteris tsutsuiana Sa. Kurata
Dryopteris vidyae Fraser-Jenk.
Sri Lanka, S. India (Tamil Nadu)
Japan
Japan (Kyushu)
N. India (Darjeeling), N. E. India
(Arunachal Pradesh, Meghalaya)
Dryopteris whangshangensis Ching
China (Anhui, Fujian, Hubei, Jiangxi,
Zhejiang)
Japan (Yakushima Isl.)
S. India (Kerala, Tamil Nadu)
Dryopteris yakusilvicola Sa. Kurata
Elaphoglossum beddomei Sledge
Note
EN in the Philippines
DD in Thailand
EW in Japan (listed as
EX)
CR in Japan
CR in Japan
As known so far
endemic to N.E. India,
but is also likely to
occur in Tibet, S.W.
China or Myanmar
Narrowly Distributed Asian Pteridophytes
Table 1.
(Continued)
Name
Dryopteridaceae (continued)
Elaphoglossum brevifolium Holttum
Elaphoglossum dumrongii Tagawa et K.
Iwats.
Elaphoglossum mcclurei Ching
Elaphoglossum murkelense M. Kato
Elaphoglossum nilgiricum Krajina ex Sledge
Elaphoglossum robinsonii Holttum
Elaphoglossum stigmatolepis (Fée) T. Moore
Elaphoglossum tosaense (Yatabe) Makino
Lastreopsis subrecedens Ching
Lomagramma brassii Holttum
Peranema cyatheoides D. Don var. luzonicum (Copel.) Ching et S. H. Wu
Phanerophlebiopsis polyodon (Ching)
Fraser-Jenk., comb. nov.
Polystichum adungense Ching et FraserJenk. ex H.S.Kung et Li Bing Zhang
Polystichum alcicorne (Baker) Diels
Polystichum altum Ching ex Li Bing Zhang
et H. S. Kung
Polystichum annapurnicola Fraser-Jenk.
Polystichum anomalum (Hook. et Arn.) J.
Sm.
Polystichum centronepalense Fraser-Jenk.
Polystichum cuneatum Nakaike
Polystichum fibrillosopaleaceum (Kodama)
Tagawa var. marginale Seriz.
Polystichum gracilipes C. Chr.
Polystichum inaense (Tagawa) Tagawa
Polystichum kwangtungense Ching
Polystichum manickamianum Beniamin,
Fraser-Jenk. et Irud.
Polystichum obae Tagawa
Polystichum palniense Fraser-Jenk.
Polystichum papuanum C. Chr.
Polystichum subinerme (Kunze) Fraer-Jenk.
Polystichum tortuosum Nakaike
Teratophyllum hainanense S. Y. Dong et X.
C. Zhang
Lomariopsidaceae
Cyclopeltis mirabilis Copel.
Lomariopsis chinensis Ching
Tectariaceae
Aenigmopteris elegans Holttum
Arthropteris caudata Rosenst.
Ctenitopsis acrocarpa Ching
Ctenitopsis matthewi (Ching) Ching
Heterogonium ceramense Holttum
Heterogonium hennipmanii Tagawa et K.
Iwats.
Heterogonium wenzelii (Copel.) Holttum
109
Distribution
Peninsular Malaysia
Thailand (Loei)
China (Guangdong, Hainan)
Maluku Islands (Seram)
S. India (Kerala, Tamil Nadu)
Peninsular Malaysia
S. India (Tamil Nadu)
Japan
China (Hainan)
New Guinea (Papua New Guinea)
Philippines (Benguet: Mt. Pulag)
N. E. India (Arunachal Pradesh, Manipur,
Meghalaya)
Arunachal Pradesh, N. Myanmar, China
Note
DD in Thailand
VU in Japan
CR in the Philippines.
As known so far
endemic to N.E. India,
though to be expected
in Myanmar and/or
S.W. China
China (Chongqing, Guizhou, Sichuan)
China (Chongqing, Sichuan, Yunnan)
C. Nepal
Sri Lanka, S. India (Keral, Tamil Nadu)
C. Nepal
New Guinea (Papua New Guinea)
Japan (Honshu)
As known so far
endemic to Nepal,
pending further study
CR in Japan
Japan (Honshu)
EN in Japan
Japan (Honshu)
NT in Japan
China (Guangdong)
S. India (Tamil Nadu, Agastyamalai Hills)
Japan (Amami Isl.)
S. India (Tamil Nadu: Palni Hills)
New Guinea (Papua New Guinea)
S. India (Kerala, Tamil Nadu)
New Guinea (Papua New Guinea)
China (Hainan)
CR in Japan
Borneo (Sarawak)
China (Yunnan)
Borneo (Sabah)
New Guinea (Papua New Guinea)
China (Yunnan)
China (Guangdong)
Maluku Islands (Seram)
Thailand (Tak)
Philippines (Leyte)
CR in Thailand
EN in the Philippines,
known only from the
type
Atsushi Ebihara et al.
110
Table 1.
Name
Tectariaceae (continued)
Psomiocarpa apiifolia (J. Sm.) C. Presl
Pteridrys lofouensis (Christ) C. Chr. et
Ching
Pteridrys olivacea (Rosenst.) Copel.
Pteridrys zeylanica Ching
Tectaria andersonii Holttum
Tectaria cherasica Holttum
Tectaria craspedocarpa Copel.
Tectaria crenata Cav. var. petiolata M. Kato
Tectaria croftii Holttum
Tectaria curtisii Holttum var. curtisii
Tectaria curtisii Holttum var. hendersonii
Holttum
Tectaria danfuensis Holttum
Tectaria exauriculata Holttum
Tectaria guachana Rusea
Tectaria hekouensis Ching et Chu H. Wang
Tectaria holttumii C. Chr.
Tectaria jacobsii Holttum
Tectaria laotica C. Chr. et Tardieu
Tectaria lobbii (Hook.) Copel.
Tectaria luchunensis S. K. Wu
Tectaria manilensis (C. Presl) Holttum var.
chupangensis (Ridl.) Holttum
Tectaria media Ching
Tectaria melanocauloides M. Kato
Tectaria microchlamys Holttum
Tectaria minuta Copel.
Tectaria nesiotica Holttum
Tectaria nitens Copel.
Tectaria pseudosiifolia sp. nov., in prep.
Tectaria rheophytica Holttum
Tectaria seramensis M. Kato
Tectaria simaoensis Ching et Chu H. Wang
Tectaria squamipes Holttum
Tectaria stalactica M. G. Price
Tectaria subcordata Holttum
(Continued)
Distribution
Philippines (Bataan, Isabela, Laguna,
Quezon, Masbate, Samar, Leyte, Bohol,
Panay, Zamboanga)
China (Guizhou, Yunnan)
Note
A monotypic Philippine endemic genus
Maluku Islands (Seram), New Guinea
Sri Lanka, S. India (Kerala, Andhra
Pradesh)
Borneo (Sarawak)
Peninsular Malaysia
New Guinea (Papua New Guinea)
Maluku Islands (Seram)
New Guinea (Papua New Guinea)
Peninsular Malaysia
Peninsular Malaysia
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
Peninsular Malaysia
China (Yunnan)
Borneo (Sabah)
Borneo (Brunei)
Thailand (Loei), Laos (Salavan)
Borneo (Sarawak), Philippines (Bohol)
China (Yunnan)
Peninsular Malaysia
China (Hainan)
Maluku Islands (Seram, Ambon)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
Borneo (Sabah)
Bhutan, Arunachal Pradesh
New Guinea (Papua New Guinea)
Maluku Islands (Seram)
China (Yunnan)
New Guinea (Papua New Guinea)
Philippines (Cagayan, Ilocos Norte,
Isabela, Polillo)
New Guinea (Papua New Guinea)
DD in Thailand and
Laos
As known so far
endemic to N.E. India,
but likely to occur in
China or Myanmar.
Differs from T. siifolia
in the fronds not being
at all dimorphic;
narrow scales at
stipe-base concolorous
dark-brown or blackish, without scarious
margins; pinnae never
with a proliferous
bulbil; costae and veins
beneath with more
prominent and dense
hairs
VU in the Philippines
Narrowly Distributed Asian Pteridophytes
Table 1.
Name
Tectariaceae (continued)
Tectaria subdigitata (Baker) Copel.
Tectaria translucens Holttum
Tectaria trimenii (Bedd.) C. Chr.
Tectaridium macleanii Copel.
Oleandraceae
Oleandra hainanensis Ching
Oleandra intermedia Ching
Davalliaceae
Davallia dimorpha Holttum
Davallia petelotii Tardieu et C. Chr.
Davallia seramensis M. Kato
Davallia sessilifolioides M. Kato
Davallodes seramense M. Kato
Humata dissecta Alston
Polypodiaceae
Acrosorus pectinatus sp. nov. in prep.
[Simpson & Marsh s. n. (K)]
Acrosorus sclerophyllus (Alderw.) Parris,
comb. nov.
Acrosorus subtriangularis (Alderw.) Parris,
comb. nov.
Acrosorus tenuis sp. nov. in prep. [S.
Dransfield et al. 947 (K)]
Acrosorus vallatus sp. nov. in prep. [Kato et
al. C-12130 (TI)]
Aglaomorpha cornucopia (Copel.) Roos
Arthromeris cyrtomioides S. G. Lu et C. D.
Xu
Belvisia platyrhynchos (Kunze) Copel.
(=Belvisia glauca (Copel.) Copel.)
Calymmodon acrosoroides sp. nov. in prep.
[Ledermann 8855c (B)]
Calymmodon acutangularis sp. nov. in prep.
[Schultze 3348a (B)]
Calymmodon binaiyensis sp. nov. in prep.
[Kato et al. C-3863 (TI)]
Calymmodon borneensis sp. nov. in prep.
[Parris 6719 (K)]
Calymmodon coriaceus sp. nov. in prep.
[Vink 17467 (BM)]
Calymmodon debilis sp. nov. in prep. [Kato
et al. C-7750 p. p. (TI)]
Calymmodon decipiens sp. nov. in prep.
[Parris 8484 (K)]
Calymmodon fragilis Copel.
Calymmodon gibbsiae sp. nov. in prep.
[Gibbs 6119 (K)]
Calymmodon holttumii sp. nov. in prep.
[Parris 11590 (K)]
111
(Continued)
Distribution
Borneo (Sarawak)
Peninsular Malaysia
Sri Lanka, S. India (Tamil Nadu)
Philippines (Cagayan, Isabela, Quezon,
Camarines Sur, Samar, Panay)
Note
EN in the Philippines
China (Hainan)
China (Guangdong, Guangxi, Guizhou,
Hongkong, Yunnan)
Peninsular Malaysia
Thailand (Phitsanulok), Laos
Sulawesi, Maluku Islands (Seram), New
Guinea
Maluku Islands (Seram)
Maluku Islands (Seram)
Maluku Islands (Ambon), Waigeu
DD in Thailand and
Laos
Borneo (Brunei: Mt Galagas)
New Guinea (Prov. Papua)
Maluku Islands (Ambon)
Borneo (Brunei, Sabah)
Maluku Islands (Seram: Murkele Ridge)
Philippines (Kalinga, Abra, Mountain
Province, Benguet, Bukidnon, Davao,
Misamis Occidental, Cotabato, Zamboanga)
China (Yunnan), Arunachal Pradesh
Philippines (Luzon, Mindoro, Agusan del
Norte, Bukidnon, Davao), New Guinea
(Prov. Papua: Yapen Island)
New Guinea (Papua New Guinea:
Kameelrucken)
New Guinea (Prov. Papua: Tami-Mundung)
Maluku Islands (Seram: Gunung Binaiya)
Borneo (Sabah, Sarawak)
New Guinea (Papua New Guinea: Mt.
Ambua)
Maluku Islands (Seram: Murkele Ridge)
Borneo (Sabah: Mt. Kinabalu)
New Guinea (Prov. Papua)
New Guinea (Prov. Papua: Angi Lakes)
Borneo (Sabah: Mt. Kinabalu)
VU in the Philippines
Atsushi Ebihara et al.
112
Table 1.
Name
Polypodiaceae (continued)
Calymmodon hyalinus Copel.
Calymmodon hygroscopicus Copel.
Calymmodon ichthyorhachioides Alston
Calymmodon innominatus sp. nov. in prep.
[Parris 8646 (K)]
Calymmodon kanikehensis sp. nov. in prep.
[Kato et al. C-2999 (TI)]
Calymmodon kinabaluensis sp. nov. in prep.
[Parris 8716]
Calymmodon ledermannii sp. nov. in prep.
[Ledermann 8457 (B)]
Calymmodon linearis sp. nov. in prep.
[Ledermann 10118 (B)]
Calymmodon mniodes Copel.
Calymmodon morobensis sp. nov. in prep.
[Parris 9573 (K)]
Calymmodon murkelensis sp. nov. in prep.
[Kato et al. C-12467 (TI)]
Calymmodon ohaensis sp. nov. in prep.
[Kato et al. C-12050 (TI)]
Calymmodon pallidivirens sp. nov. in prep.
[Sundue 2218 (KNP)]
Calymmodon papuanus sp. nov. in prep.
[Carr 15230 (B)]
Calymmodon persimilis Tagawa
Calymmodon pseudoclavifer sp. nov. in
prep. [Ratcliffe 230 (K)]
Calymmodon pseudordinatus sp. nov. in
prep. [Parris 6138 (K)]
Calymmodon ramifer Copel.
Calymmodon redactus sp. nov. in prep.
[Clemens 39161 p. p. (K)]
Calymmodon schultzei sp. nov. in prep.
[Schultze 33 32 (B)]
Calymmodon seramensis sp. nov. in prep.
[Kato et al. C-5330 (TI)]
Calymmodon subalpinus sp. nov. in prep.
[Croft 112 (K)]
Calymmodon tehoruensis sp. nov. in prep.
[Kato et al. C-13804 (TI)]
Colysis acuminata (Baker) Holttum var.
fluminalis (Ridley) Hennipmann
Colysis fluviatilis (Laut.) Ching
Colysis longipes Ching
Craspedosorus sinensis Ching et W. M. Chu
Ctenopterella cornigera (Baker) Parris
Ctenopterella fluvialis sp. nov. in prep.
[Johns et al. 7392 (K)]
Ctenopterella pediculata (Baker) Parris
Ctenopteris thwaitesii (Bedd.) Sledge
Dasygrammitis kinabaluensis sp. nov. in
prep. [Parris 11584 (K)]
Dasygrammitis seramensis sp. nov. in prep.
[Kato et al. C-14387 (TI)]
Goniophlebium someyae (Yatabe) Ebihara,
comb. nov.
(Continued)
Distribution
Note
Borneo (Sarawak)
New Guinea (Papua New Guinea)
New Guinea (Prov. Papua)
Borneo (Sabah: Mt. Kinabalu)
Maluku Islands (Seram: Kanikeh)
Borneo (Sabah: Mt. Kinabalu)
New Guinea (Papua New Guinea: East
Sepik, Eastern Highlands, Morobe)
New Guinea (Papua New Guinea:
Lordberg)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea:
Morobe, Central, New Ireland)
Maluku Islands (Seram: Murkele Ridge)
Maluku Islands (Seram: Murkele Ridge)
Borneo (Sabah: Mt. Kinabalu or Gunung
Alab)
New Guinea (Papua New Guinea:
Western Highlands, Central)
New Guinea (Papua New Guinea)
New Guinea (Prov. Papua: Cyclops Mts.)
Philippines (Luzon: Mt. Banahaw)
New Guinea (Prov. Papua)
Borneo (Sabah: Mt. Kinabalu)
New Guinea (Prov. Papua: Tami-Mundung)
Maluku Islands (Seram: Sawai)
New Guinea (Papua New Guinea: Mt.
Piora)
Maluku Islands (Seram: Murkele Ridge)
Malay Peninsula, Borneo
Borneo
China (Guangxi, Hainan)
China (Yunnan)
Sri Lanka
Borneo (Brunei: Temburong River)
Known only from a
single locality
Borneo (Sarawak)
Sri Lanka
Borneo (Sabah: Mt. Kinabalu)
Maluku Islands (Seram: Murkele Ridge)
Japan
EN in Japan
Narrowly Distributed Asian Pteridophytes
Table 1.
Name
Polypodiaceae (continued)
Goniophlebium terrestre Copel.
Grammitis demissa Parris
Grammitis kairatuensis M. Kato et Parris
Lecanopteris lomarioides (Kunze) Copel.
Lecanopteris luzonensis Hennip.
Lecanopteris mirabilis (C. Chr.) Copel.
Lepisorus affinis Ching
Lepisorus boninensis (Christ) Ching
Lepisorus cespitosus Y. X. Lin
Lepisorus confluens W. M. Chu
Lepisorus elegans Ching et W. M. Chu ex
W. M. Chu
Lepisorus likiangensis Ching et S. K. Wu
Lepisorus longus Ching
Lepisorus luchunensis Y. X. Lin
Lepisorus medogensis Ching et Y. X. Lin
Lepisorus oosphaerus (C. Chr.) Ching
Lepisorus sordidus (C. Chr.) Ching
Lepisorus subconfluens Ching
Lepisorus tricholepis K. H. Shing ex Y. X. Lin
Lepisorus waltonii (Ching) S. L. Yu
Lepisorus xiphiopteris (Baker) W. M. Chu
ex Y. X. Lin
Leptochilus metallicus (Bedd.) C. Chr.
Leptochilus wallii (Baker) C. Chr.
Loxogramme antrophyoides (Baker) C. Chr.
var. rheophila M. Kato, Darnaedi et K.
Iwats.
Loxogramme boninensis Nakai
Micropolypodium pulogense (Copel.) A. R.
Sm.
Microsorum krayanense M. Kato, Darnaedi
et K. Iwats.
Microsorum membranaceum (D. Don)
Ching var. carinatum W. M. Chu et Z. R.
He
Microsorum paucijugum (Alderw.) K. Iwats.
et M. Kato
Microsorum reticulatum Ching ex L. Shi
Microsorum siamense Boonkerd
Microsorum submarginale M. Kato,
Darnaedi et K. Iwats.
Microsorum thailandicum Boonkerd et
Noot.
Neocheiropteris palmatopedata (Baker)
Christ
Neocheiropteris triglossa (Baker) Ching
Oreogrammitis alstonii sp. nov. in prep.
[Alston 16878 (BM)]
Oreogrammitis antipodalis (Copel.) Parris
113
(Continued)
Distribution
Philippines (Quirino, ?Isabela, Laguna)
New Guinea (Papua New Guinea)
Maluku Islands (Seram)
Philippines (Nueva Ecija, Nueva Vizcaya), Sulawesi
Philippines (Zambales, Bataan, Rizal,
Quezon, Laguna, Lanao)
Maluku Islands (Seram, Ambon), New
Guinea
China (Hainan)
Japan (Bonin Islands)
China (Xizang)
China (Yunnan)
China (Yunnan)
China (Sichuan, Yunnan)
China (Hainan)
China (Yunnan)
China (Xizang)
Thailand (Chiang Mai)
China (Sichuan, Yunnan), Myanmar,
India (Arunachal Pradesh, Nagaland,
Manipur)
China (Yunnan), India (Arunachal
Pradesh, ?Sikkim), Myanmar, ?Nepal,
?Bhutan, ?Thailand
China (Xizang)
China (Xizang)
China (Xizang, Yunnan)
Sri Lanka
Sri Lanka
Borneo (E. Kalimantan)
Japan (Bonin Islands)
Philippines (Luzon)
Borneo (E. Kalimantan)
Note
VU in the Philippines
EN in the Philippines
EN in the Philippines
NT in Japan
VU in Thailand
Possibly extinct
VU in Japan
Known only from three
collections from Mt.
Pulog
China (Yunnan)
Borneo
China (Yunnan)
Thailand (Yala)
Borneo (E. Kalimantan)
DD in Thailand
Thailand (Chumphon)
EN in Thailand
China (Guizhou, Sichuan, Yunnan)
China (Yunnan)
Maluku Islands (Halmahera: Jailolo Peak)
New Guinea (Prov. Papua)
Atsushi Ebihara et al.
114
Table 1.
Name
Polypodiaceae (continued)
Oreogrammitis atrostipes sp. nov. in prep.
[Klimas 176 (COLO)]
Oreogrammitis attenuata (Kunze) Parris
Oreogrammitis austroindica (Parris) Parris
Oreogrammitis carrii sp. nov. in prep. [Carr
15219 (K)]
Oreogrammitis celebica sp. nov. in prep.
[Hennipman 5388 p. p. (K)]
Oreogrammitis clandestina sp. nov. in prep.
[Ranker 2175 (COLO)]
Oreogrammitis clavata (Parris) Parris
Oreogrammitis crinifera (Parris) Parris
Oreogrammitis crispatula (Holttum) Parris
Oreogrammitis dictymioides (Copel.) Parris
Oreogrammitis distincta sp. nov. in prep.
[Johns 6742 (K)]
Oreogrammitis excelsa (Parris) Parris
Oreogrammitis fenicis (Copel.) Parris
Oreogrammitis gardettei sp. nov. in prep.
[Gardette 1061 (K)]
Oreogrammitis glossophylla (Parris) Parris
Oreogrammitis graniticola sp. nov. in prep.
[Parris 8763 (K)]
Oreogrammitis habbemensis (Copel.) Parris
Oreogrammitis hainanensis sp. nov. in prep.
[Smith 1548 (K)]
Oreogrammitis hispida (Copel.) Parris
Oreogrammitis imberbis (Parris) Parris
Oreogrammitis incognita sp. nov. in prep.
[Ranker 2185 (COLO)]
Oreogrammitis inconstans (Alderw.) Parris
Oreogrammitis jeraiensis (Parris) Parris
(Continued)
Distribution
New Guinea (Papua New Guinea: Crater
Mountain)
Sri Lanka, S. India (Tamil Nadu),
S. India (Tamil Nadu)
New Guinea (Papua New Guinea: Owen
Stanley Range)
Sulawesi: Mt. Rorokatimbu
Note
Known from a single
19th century collection,
probably extinct
Borneo (Sabah: Mt. Kinabalu)
New Guinea (Prov. Papua)
New Guinea (Prov. Papua)
Peninsular Malaysia
New Guinea (Prov. Papua)
Borneo (Brunei: Gunung Retak)
EX in Peninsular
Malaysia
New Guinea (Prov. Papua)
Philippines, Taiwan
Sumatra (Gunung Tujuh)
New Guinea (Papua New Guinea)
Borneo (Sabah: Mt. Kinabalu)
New Guinea (Papua New Guinea)
China (Hainan: Five Finger Mountain)
New Guinea (Prov. Papua)
New Guinea (Papua New Guinea)
Borneo (Sabah: Mt. Kinabalu)
New Guinea (Prov. Papua)
Peninsular Malaysia
Oreogrammitis katoi sp. nov. in prep. [Kato
et al. 13619]
Oreogrammitis kunstleri Parris
Maluku Islands (Seram: Kairatu)
Oreogrammitis malayensis Parris
Peninsular Malaysia
Oreogrammitis manuselensis sp. nov. in
prep. [Kato et al. C-11761bis (TI)]
Oreogrammitis medialis (Baker) Parris
Oreogrammitis meijer-dreesii (Copel.) Parris
Maluku Islands (Seram: Murkele Ridge)
Oreogrammitis mesocarpa (Alderw.) Parris
Oreogrammitis mollis sp. nov. in prep.
[Parris 8482 (K)]
Oreogrammitis montana (Parris) Parris
Oreogrammitis murrayana (C. Chr.) Parris
Oreogrammitis murudensis sp. nov. in prep.
[SAR 88632 p. p. (KEP)]
Oreogrammitis muscicola sp. nov. in prep
Parris 8525 (K).
Oreogrammitis nana (Fée) Parris
Oreogrammitis natunae sp. nov. in prep.
[Everett s. n. (E)]
New Guinea (Prov. Papua)
Borneo (Sabah: Mt. Kinabalu)
Peninsular Malaysia
Sri Lanka
New Guinea (Prov. Papua)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
Borneo (Sarawak: Gunung Murud)
Borneo (Sabah: Mt. Kinabalu)
Java
Borneo (Kalimantan: Natuna Islands)
EN in Peninsular
Malaysia
EX in Peninsular
Malaysia
EN in Peninsular
Malaysia
VU in New Guinea
(Prov. Papua)
Narrowly Distributed Asian Pteridophytes
Table 1.
(Continued)
Name
Polypodiaceae (continued)
Oreogrammitis nigropaleata (Copel.) Parris
Oreogrammitis nipponica (Tagawa et K.
Iwats.) Parris
Oreogrammitis nubicola sp. nov. in prep.
[Parris 8531 (K)]
Oreogrammitis oncobasis (M. G. Price)
Parris
Oreogrammitis pilifera (Ravi et J. Joseph)
Parris
Oreogrammitis piruensis (M. Kato et Parris)
Parris
Oreogrammitis polita Parris
Oreogrammitis pseudospiralis (Alderw.)
Parris
Oreogrammitis ramicola sp. nov. in prep.
[Parris 8501 (K)]
Oreogrammitis reptans (Parris) Parris
Oreogrammitis rupestris (Parris) Parris
Oreogrammitis salticola (Parris) Parris
Oreogrammitis similis sp. nov. in prep.
[Hennipman 5388 p. p. (K)]
Oreogrammitis simplicivenia sp. nov. in
prep. [Majuakim 1 (KEP)]
Oreogrammitis sledgei (Parris) Parris
Oreogrammitis subreticulata (Copel.) Parris
Oreogrammitis taeniophylla (Parris) Parris
Oreogrammitis tenuis (Parris) Parris
Oreogrammitis tidorensis sp. nov. in prep.
[Alston 16730 (BM)]
Oreogrammitis trogophylla (Copel.) Parris
Oreogrammitis tuyamae (H. Ohba) Parris
Oreogrammitis ultramaficola sp. nov. in
prep. [Parris 8500 (K)]
Oreogrammitis velutina (Parris) Parris
Oreogrammitis wallii (Beddome) Parris
Oreogrammitis xiphopteroides sp. nov. in
prep. [Croft & Hope 51 (K)]
Oreogrammitis zeylanica (Fée) Parris
Phymatopteris cruciformis (Ching) Pic.
Serm.
Phymatopteris digitata (Ching) Pic. Serm.
Platycerium grande (Fée) Kunze.
Podosorus angustatus Holttum
Polypodiodes falcipinnula S. K. Wu et J.
Murata
Polypodiodes niponica (Mett.) Ching var.
glandulosa P. S. Wang
Polypodiodes paramoena Ching et Y. X. Lin
Polypodiodes pseudolachnopus S. G. Lu
115
Distribution
New Guinea (Papua New Guinea)
Japan
Note
CR in Japan
Borneo (Sabah: Mt. Kinabalu)
Philippines
S. India (Kerala, Tamil Nadu)
Maluku Islands (Seram)
New Guinea (Papua New Guinea)
New Guinea (Prov. Papua)
Borneo (Sabah: Mt. Kinabalu)
New Guinea (Prov. Papua)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
Sulawesi: Mt. Rorokatimbu
Borneo (Sabah: Gunung Silam)
Sri Lanka
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
Maluku Islands (Tidore: Gunung Kiematuba)
New Guinea (Papua New Guinea)
Japan (Bonin Islands)
Borneo (Sabah: Mt. Kinabalu)
CR in Japan
New Guinea (Prov. Papua)
Sri Lanka
New Guinea (Papua New Guinea: Star
Mountains)
Sri Lanka
China (Guangdong)
China (Guangdong, Guizhou, Zhejiang,
Hunan)
Philippines (?Masbate: Ticao Isl., Lanao
del Sur, Davao, Zamboanga)
Philippines (Isabela: San Mariano,
Barangay Disulap)
China (Yunnan)
China (Guizhou)
China (Xizang)
China (Sichuan, Xizang, Yunnan)
CR in the Philippines
CR in the Philippines,
a monotypic genus
known only from the
type
Atsushi Ebihara et al.
116
Table 1.
Name
Polypodiaceae (continued)
Polypodiodes simonsiana Fraser-Jenk., in
prep.
(Continued)
Distribution
Note
N. E. India (Arunachal Pradesh, Manipur,
Meghalaya), Myanmar
Similar to P. niponica,
but with a curved
rachis and mid and
lower pinnae backward-deflexed and very
acutely pointed
CR in Japan
Polypodium amamianum Tagawa
Prosaptia archboldii Copel.
Prosaptia binaiyensis sp. nov. in prep.
[Parris 11166 (K)]
Prosaptia borneensis sp. nov. in prep.
[Parris 6683 (K)]
Prosaptia ceylanica Parris
Prosaptia crenata sp. nov. in prep. [LAE
61379 (K)]
Prosaptia fungosa sp. nov. in prep. [Brass
13207 (MICH)]
Prosaptia fusca (Copel.) Parris
Japan (Amami Islands)
New Guinea (Prov. Papua)
Maluku Islands (Seram: Gunung Binaiya)
Prosaptia giluwensis sp. nov. in prep. [Parris
9369 (K)]
Prosaptia idenbergensis sp. nov. in prep.
[Brass 13653 (UC)]
Prosaptia khasyana (Hook.) C. Chr. et
Tardieu
New Guinea (Papua New Guinea: Mt.
Giluwe)
New Guinea (Prov. Papua: Idenberg
River)
N. E. India (Meghalaya), Myanmar
Prosaptia koitakiensis sp. nov. in prep. [Carr
12650 (K)]
Prosaptia micropora (Copel.) Parris
Prosaptia morobensis sp. nov. in prep.
[Woods 1640 (E)]
Prosaptia mulierum sp. nov. in prep. [Gay
775 (K)]
Prosaptia negrosensis (Copel.) M. G. Price
Prosaptia novae-hiberniae sp. nov. in prep.
[LAE 68305 (K)]
Prosaptia pinnata sp. nov. in prep. [Kluge
7275 (UC)]
Prosaptia repens sp. nov. in prep. [Parris
8112 (K)]
Prosaptia rivularis sp. nov. in prep. [NGF
45058 (K)]
Prosaptia seramensis sp. nov. in prep. [Kato
et al. C-12532]
Prosaptia shawii (Copel.) Parris
Prosaptia subglabra (M. Kato et Parris)
Parris
Prosaptia subulatipinna (Alderw.) Parris
Prosaptia sulawesica sp. nov. [Hennipman
5303 (L)]
Prosaptia wobbensis sp. nov. in prep.
[Schlechter 16390 (K)]
Pyrrosia boothii (Hook.) Ching
New Guinea (Papua New Guinea: Owen
Stanley Range)
Philippines
New Guinea (Papua New Guinea: Mt.
Kaindi)
New Guinea (Papua New Guinea:
Western Highlands, Eastern Highlands)
Philippines
New Guinea (Papua New Guinea: New
Ireland)
Sulawesi: Mt. Roroka Timbu
Pyrrosia ceylanica (Giesenh.) Sledge
Borneo (Sabah, Sarawak)
Sri Lanka
New Guinea (Papua New Guinea: Mt.
Albert Edward)
New Guinea (Prov. Papua: Idenberg
River)
New Guinea (Prov. Papua)
New Guinea (Papua New Guinea: West
Sepik, Western Highlands, Enga)
New Guinea (Papua New Guinea:
Garaina)
Maluku Islands (Seram: Murkele Ridge,
Gunung Binaiya, Sawai)
New Guinea (Papua New Guinea)
Maluku Islands (Seram, Bacan)
New Guinea (Prov. Papua)
Sulawesi: Mt. Rorokatimbu
New Guinea (Papua New Guinea:
Wobbe)
Sikkim, Bhutan, Arunachal Pradesh,
Myanmar
Sri Lanka, S. India (Karnataka, Kerala,
Tamil Nadu)
EN in New Guinea
(Prov. Papua)
Records of this species
from elsewhere,
including S. China and
S. E. Asia refer to
P. barathrophylla
(Baker) M .G. Price
Narrowly Distributed Asian Pteridophytes
Table 1.
(Continued)
Name
Polypodiaceae (continued)
Pyrrosia splendens (C. Presl) Ching
Radiogrammitis alepidota (M. G. Price)
Parris
Radiogrammitis beddomeana (Alderw.)
Parris
Radiogrammitis celata sp. nov. in prep.
[Ranker 2181 p. p. (COLO)]
Radiogrammitis coredrosora (Alderw.)
Parris
Radiogrammitis exilis sp. nov. in prep.
[Sundue 2223 (KNP)]
Radiogrammitis kanikehensis sp. nov. in
prep. [Kato et al. C-2998 p. p. (TI)]
Radiogrammitis kinabaluensis (Copel.)
Parris
Radiogrammitis peninsularis (Copel.) Parris
117
Distribution
Note
Philippines (Luzon, Samar, Leyte, Bohol,
Mindanao)
Vietnam, Philippines (Luzon, Negros)
Sri Lanka
Borneo (Sabah: Mt. Kinabalu)
New Guinea (Prov. Papua)
Borneo (Sabah: Gunung Alab)
Maluku Islands (Seram: Kanikeh)
Borneo (Sabah)
Peninsular Malaysia
Radiogrammitis plana (Alderw.) Parris
Radiogrammitis pseudoxiphopteris (Parris)
Parris
Radiogrammitis reducta (Alderw.) Parris
Radiogrammitis silvicola (Parris) Parris
Radiogrammitis tehoruensis (M. Kato et
Parris) Parris
Radiogrammitis tuberculata (Parris) Parris
Selliguea yakuinsularis (Masam.) H. Ohashi
et K. Ohashi
Themelium abebaion (Alderw.) Parris
Themelium allocotum (Alderw.) Parris
Sumatra
Peninsular Malaysia
Themelium crenulatum (Parris) Parris
Themelium ctenoideum (Brause) Parris
Themelium elatum sp. nov. in prep. [van
Royen & Sleumer 8180a (K)]
Themelium flagelliforme (Brause) Parris
Themelium gibbsiae sp. nov. in prep. [Gibbs
6119 (K)] [Gibbs 5664 (K)]
Themelium loherianum (Christ) Parris
Themelium pseudallocotum sp. nov. ined.
[Kato et al. C-14165 (TI)]
Themelium pullei (Alston) Parris
Tomophyllum bipinnatum (Copel.) Parris
Tomophyllum capillatum (Brause) Parris
Tomophyllum congregatifolium (Alderw.)
Parris
Tomophyllum cumingii sp. nov. in prep.
[Cuming 205 (E)]
Tomophyllum daymanense sp. nov. in prep.
[Brass 22634 (L)]
Tomophyllum dissitum sp. nov. in prep. [de
Wilde & de Wilde-Duyfjes 14254 (L)]
Tomophyllum epaleatum (Parris) Parris
Tomophyllum havilandii (Baker) Parris
Tomophyllum hymenophylloides (Parris)
Parris
Tomophyllum perplexum (Parris) Parris
Tomophyllum plicatum sp. nov. in prep.
[Lack & Grimes 1752 (K)]
New Guinea (Papua New Guinea)
New Guinea (Papua New Guinea)
New Guinea (Prov. Papua: Nettoti Range)
New Guinea (Prov. Papua)
New Guinea (Papua New Guinea)
Maluku Islands (Seram)
New Guinea (Papua New Guinea)
Japan
New Guinea (Prov. Papua)
New Guinea (Prov. Papua)
New Guinea (Papua New Guinea)
New Guinea (Prov. Papua: Angi Lakes)
Philippines
Maluku Islands (Seram: Muselleinan
Pass)
New Guinea (Prov. Papua)
New Guinea (Prov. Papua)
New Guinea (Papua New Guinea)
New Guinea (Prov. Papua)
Philippines (Luzon)
New Guinea (Papua New Guinea)
Sumatra: Gunung Leuser
Sri Lanka, Java
Borneo (Sabah)
New Guinea (Prov. Papua)
Sri Lanka, S. India (Tamil Nadu)
Sulawesi: Gunung Tambusisi
EN in Peninsular
Malaysia
CR in Peninsular
Malaysia
EN in Japan
VU in New Guinea
(Prov. Papua)
Atsushi Ebihara et al.
118
Table 1.
Name
Polypodiaceae (continued)
Tomophyllum sakaguchianum (Koidz.)
Parris
Tomophyllum sesquipinnatum (Copel.)
Parris
Tomophyllum terpsichoroides sp. nov. in
prep. [Croft 1336 p. p. (K)]
Tomophyllum themelioides sp. nov. in prep.
[Parris 11189 (K)]
Tomophyllum warburgii sp. nov. in prep.
[Warburg 4720 (B)]
Xiphopterella alternidens (Cesati) Parris
Xiphopterella coriacea sp. nov. in prep.
[Sundue 2231 (KNP)]
Xiphopterella corneri sp. nov. in prep.
[RSNB 1869A (K)]
Xiphopterella gracilis Parris
Xiphopterella lenbrassii sp. nov. in prep.
[Brass 25701 (K)]
Xiphopterella major sp. nov. in prep.
[Ranker 2194 (COLO)]
Xiphopterella murudensis (Copel.) Parris,
comb. nov.
Xiphopteris nudicarpa (P. M. Zamora et Co)
Parris
(Continued)
Distribution
Japan
Note
EN in Japan
New Guinea (Prov. Papua)
New Guinea (Papua New Guinea: Mt.
Saruwaged)
Maluku Islands (Seram: Gunung Binaiya)
Java: Gunung Wilis
Borneo (Sarawak, Kalimantan)
Borneo (Sabah: Mt Kinabalu or Gunung
Alab)
Borneo (Sabah: Mt Kinabalu)
Peninsular Malaysia
New Guinea (Papua New Guinea:
Normanby Island)
Borneo (Sabah: Mt. Kinabalu)
Borneo (Sarawak)
Borneo (Sabah), Philippines (Palawan)
Acknowledgments
This study was supported by the Environment
Research and Technology Development Fund
(S-9) of the Ministry of the Environment, Japan.
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Appendix 1.
species
New combinations in threatened
Acrosorus sclerophyllus (Alderw.) Parris, comb.
nov., based on Polypodium sclerophyllum
Alderw., Nova Guinea 14: 45 (1924).
Acrosorus subtriangularis (Alderw.) Parris,
comb. nov., based on Polypodium subtriangularis Alderw., Bull. Dép. Agric. Indes Néerl.
18: 20 (1908).
Deparia lancea (Thunb.) Fraser-Jenk. var.
rheophila (M. Kato, Darnaedi et K. Iwats.)
M. Kato, comb. nov., based on Diplazium
subsinuatum (Wall. ex Hook. et Grev.) Tagawa
var. rheophilum M. Kato, Darnaedi et K. Iwats.,
J. Fac. Sci. Univ. Tokyo III 15(1): 100 (1991).
Diplopterygium angustilobum (Holttum) Parris,
View publication stats
119
comb. nov., based on Gleichenia angustiloba
Holttum, Reinwardtia 4: 266 (1957).
Diplopterygium clemensiae (Copel.) Parris,
comb. nov., based on Hicriopteris clemensiae
Copel., Univ. California Publ. Botany 18: 217
(1942).
Diplopterygium deflexum (Holttum) Parris,
comb. nov., based on Gleichenia deflexa Holttum, Reinwardtia 4: addendum p. 280 (1957).
Goniophlebium someyae (Yatabe) Ebihara,
comb. nov., based on Polypodium someyae
Yatabe, Bot. Mag. (Tokyo) 5: 245 (1891).
Odontosoria minutula (Sa. Kurata) Ebihara,
comb. nov., based on Sphenomeris minutula
Sa. Kurata, J. Geobot. 13: 101 (1965).
Odontosoria yaeyamensis (S. J. Lin, M. Kato et
K. Iwats.) Ebihara, comb. nov., based on
Sphenomeris yaeyamensis S. J. Lin, M. Kato
et K. Iwats., J. Plant Res. 109: 207, f. 3 (1996).
Phanerophlebiopsis polyodon (Ching) FraserJenk., comb. nov., based on Polystichum
polyodon Ching, Bull. Chinese Bot. Soc. 2: 93
(1936).
Xiphopterella murudensis (Copel.) Parris,
comb. nov., based on Polypodium murudense
Copel., Philipp. J. Sci. C, Bot. 12(1): 61
(1917).