СПИСАНИЕ НА БЪЛГАРСКОТО ГЕОЛОГИЧЕСКО ДРУЖЕСТВО, год. 74, кн. 1–3, 2013, с. xx–xx REVIEW OF THE BULGARIAN GEOLOGICAL SOCIETY, vol. 74, part 1–3, 2013, p. xx–xx Paleogene hyaline benthic foraminifera (LAGENINA and ROTALIINA) from the Republic of Macedonia Boris Valchev1, Violeta Stojanova2, Sava Juranov3 “St. Ivan Rilski” University of Mining and Geology, 1700 Sofia; E-mail: b_valchev@mgu.bg “Goce Delčev” University, K-2000 Štip, Republic of Macedonia; E-mail: stojanovavioleta@yahoo.com 3 “St. Kliment Ohridski” University of Sofia, 1504 Sofia; E-mail: juranov@gea.uni-sofia.bg 1 2 Палеогенски хиалинни бентосни фораминифери (LAGENINA и ROTALIINA) от Република Македония Борис Вълчев1, Виолета Стоянова2, Сава Джуранов3 Минно-геоложки университет „Св. Иван Рилски“, София 1700 Университет „Гоце Делчев“, Щип К-2000, Република Македония 3 Софийски университет „Св. Климент Охридски“, София 1504 1 2 Резюме. Настоящата статия представя 49 вида, принадлежащи на 28 рода, 15 подсемейства, 20 семейства, 12 надсемейства от подразредите LAGENINA Delage and Hérouard, 1896 и ROTALIINA Delage and Hérouard, 1896. 36 вида се описват за първи път в Република Македония. Останалите 13 вида са описани и фигурирани в по-ранна публикация, посветена на разрез Чардаклия от Овчеполския басейн, поради което тук е посочено само стратиграфското им разпространение в останалите изучени разрези. Изследваният материал е получен от 122 проби от горната флишка задруга и задругата на жълтите пясъчници от 11 разреза в 6 палеогенски басейна, развити във Вардарската зона и Сръбско-Македонския масив. Ключови думи: Палеоген, хиалинни бентосни фораминифери, таксономия, Република Македония. Abstract. The present article aims to represent 49 species belonging to 28 genera, 15 subfamilies, 20 families, 12 superfamilies of the suborders LAGENINA Delage and Hérouard, 1896 and ROTALIINA Delage and Hérouard, 1896. 36 of the species are first described in the Republic of Macedonia. The other 13 ones were described and figured before from Chardaklija section of the Ovche Pole basin, and therefore here is shown only their stratigraphical distribution in the other studied sections. The studied foraminiferal specimens have been obtained from 122 samples coming from the upper flysh unit and the unit of yellow sandstones of 11 sections from 6 Paleogene basins developed in the Vardar Zone and Serbo-Macedonian Massif. Key words: Paleogene, hyaline benthic foraminifera, taxonomy, Republic of Macedonia. Introduction The Paleogene sediments in the Republic of Mace­ donia crop out in 7 sedimentary basins (Fig. 1): Skopje–Kumanovo, Ovche Pole, Tikvesh, Valandovo– Gevgelija, Deve Bair, Delchevo, and Strumica. They are located in two tectonic zones (Dumurdzhanov et al., 2005): Vardar Zone (the first 4 basins) and SerbоMacedonian Massif (the last 3 ones). Five lithostratigraphical units (basal unit, lower flysh unit, unit of yellow sandstones, upper flysh unit and carbonatesandy unit) have been recognized, as their lithology, thickness and spatial relationships were discussed in several works (Maksimovič et al., 1954; Stojanova, 2008; Stojanova et al., 2011; Stojanova, Petrov, 2012; Stojanova et al., 2012; Stojanova et al., 2013). The age of the studied sediments (Late Eocene–Early Oligocene) has been determined by means of macrofossils and microfossils (planctonic and benthic foraminifers). First data for Late Eocene (Priabonian) age, based on gastropods, bivalves, corals, and nummulitids from the Ovche Pole and Tikvesh basins, gave Maksimovič et al. (1954). Later on Mitrovič-Petrovič et al. (1990), confirmed the Priabonian age on the base of echinoids, and noted that the uppermost levels of the sediments of the Ovche Pole basin are of Early Oligocene age. Stojanova (Stojanova, 2008; Stojanova et al., 2011; Stojanova, Petrov, 2012; Stojanova et al., 2012) also confirmed the Upper Eocene-Oligocene range of the sediments, but pointed out that it is difficult to place the E/O boundary by means of benthic foraminifers, because of their broad stratigraphical 53 The present study is focused on the taxonomy of the hyaline foraminifers from the Paleogene basins of this part of the Balkan Peninsula. 143 samples from 11 of totally 12 outcrop sections (Figs. 2–9) from 6 basins (the Paleogene sediments of the Deve Bair basin are poorly exposed) have been investigated. The foraminiferal specimens were obtained from 122 of them. 135 samples come from the upper flysh unit and only in the Vojnik section the samples have been picked from the unit of yellow sandstones. Taxonomy Fig. 1. Sketch with the location of the Paleogene basins in Republic of Macedonia and the studied sections (modified after Dumurdzanov et al., 2005) 1, distribution of Paleogene sediments; 2, tectonic boundary (WMZ, Western Macedonian Zone; VZ, Vardar Zone; SMM, Serbian-Macedonian Massif); 3, basin boundary; 4, basins (SKB, Skopje-Kumanovo; OPB, Ovche Pole; TB, Tikvesh; VGB, Valandovo-Gevgelija; DBB, Deve Bair; DB, Delchevo; SB, Strumica); 5, section Фиг. 1. Скица с разпространението на палеогенските басейни в Република Македония и изучените разрези (по Dumurdzanov et al., 2005, с изменения) 1 – разпространение на палеогенските седиментни скали; 2 – тектонска граница (WMZ – Западномакедонска зона, VZ – Вардарска зона, SMM – Сръбско-Македонски масив); 3 – басейнова граница; 4 – басейни (SKB – СкопскоКумановски, OPB – Овчеполски, TB – Тиквешки, VGB – Валандово-Гевгелийски, DBB – Девебаирски, DB – Делчевски, SB – Струмички–; 5 – разрез The following pages represent 49 species belonging to 28 genera, 15 subfamilies, 20 families, 12 superfamilies of the suborders LAGENINA Delage and Hérouard, 1896 and ROTALIINA Delage and Hérouard, 1896. 36 of the species are first described in the Republic of Macedonia. The other 13 ones were described and figured from Chardaklija section of the Ovche Pole basin (Džuranov et al., 1999) that is why here is shown only their stratigraphical distribution in the other studied sections. One species is first recorded from Priabonian sedimentary rocks. Taxonomical determination at generic level is based on the classification of Loeblich and Tappan (1988). Suborder LAGENINA Delage and Herouard, 1896 Superfamily NODOSARIACEA Ehrehberg, 1838 Family NODOSARIIDAE Ehrehberg, 1838 Subfamily NODOSARIINAE Ehrenberg, 1838 Genus Nodosaria Lamarck, 1812 Nodosaria ewaldi Reuss, 1851 Plate I, Fig. 1 1851. N. (Nodosaria) ewaldi m.; Reuss, S. 58, Taf. 3, Fig. 2. 1969. Nodosaria ewaldi Reuss; Krayeva, Zernetskij, p. 43, pl. 15, fig. 6 (in Russian). 1971. Nodosaria ewaldi Reuss; Kaptarenko-Chernousova, p. 62, pl. 1, figs. 16, 17 (in Ukrainian). 2001. Nodosaria ewaldi Reuss; Valchev, p. 112, pl. 1, fig. 5. distribution. Despite this fact, she indicated Bolivina antegressa Subzone from the Planulina costata Zone. Juranov (in Stojanova et al., 2013), using planktonic foraminifers, determined Early Oligocene age for the Nemanjici section of the Ovche Pole basin and it was proposed that the upper levels of the upper flysh unit are of the same age. The first foraminiferal taxonomical investigations of the Paleogene sediments from the Republic of Macedonia was published at the end of the 20-th century when Džuranov et al. (1999) gave descriptions of 21 species including 2 agglutinated, 6 porcelaneous, and 13 hyaline ones from Chardaklija section of the Ovche Pole basin. Later on Stojanova et al. (2013) presented 5 taxa of planktonic foraminifers from 5 basins, and Valchev et al. (2013) introduced one porcelaneous and 7 agglutinated taxa from 4 basins. 54 Nomenclature. The holotype is the specimen figured by Reuss (1851, Taf. 3, Fig. 2). The species was first described from the Oligocene (Septarien clays) near Berlin (Germany). Material. 7 specimens, with different degree of preser­ vation. Description. The test is elongated, straight or slightly arcuate. The initial chamber is spherical and bears a short spine. The late chambers are cylindrical or tubular, separated by grooved horizontal sutures. The surface is smooth. Distribution. The species is known from the Paleocene of Bulgaria and the Netherlands, the Upper Eocene of the USA and Ukraine, the Oligocene of Germany and the Netherlands. Occurrence. Valandovo-Gevgelija basin (Rabrovo section – samples 6, 8; Dedeli section – samples 4, 6). 55 Фиг. 2. Разпространение на установените хиалинни таксони в изучените разрези (*данните за разрез Чардаклия са публикувани от Džuranov et al., 1999) Fig. 2. Distribution of the established hyaline taxa in the studied sections (*the data for Chardaklija section were published by Džuranov et al., 1999) Fig. 3. Stratigraphical distribution of hyaline taxa in Vojnik section 1, clayey-carbonate sediments; 2, clayey-carbonate-sandy sediments; 3, siltstones; 4, thin bedded sandstones; 5, conglomerates; 6, limestones; 7, sandy limestones; 8, tuffs; 9, sample containing hyaline specimens; 10, sample barren of hyaline specimens Фиг. 3. Стратиграфско разпространение на хиалинните таксони в разрез Войник 1 – глинесто-карбонатни седименти; 2 – глинесто-карбонатно-песъчливи седименти; 3 – алевролити; 4 – тънкопластови пясъчници; 5 – конгломерати; 6 – варовици; 7 – песъчливи варовици; 8 – туфи; 9 – проба, съдържаща хиалинни екземпляри; 10 – проба без хиалинни екземпляри Nodosaria sp. Plate I, Fig. 2 Material. 10 well preserved specimens. Description. The test is elongate, straight or slightly arcuate. The chambers are subspgerical, separated by broad horizontal depressed sutures. Remarks. Only test fragments have been found. Occurrence. Ovche Pole basin (Nemanjici section, samples 2, 3, 6, 8, 16–18). Family VAGINULINIDAE Reuss, 1860 Subfamily LENTICULININAE Chapman, Parr, and Collins, 1934 Genus Lenticulina Lamarck, 1804 56 Lenticulina cf. wilcoxensis (Cushman, Ponton, 1932) Plate I, Fig. 3 1932. Robulus wilcoxensis Cushman, Ponton, n. sp.; Cushman, Ponton, p. 52, pl. 7, fig. 3. 1951. Robulus wilcoxensis Cushman, Ponton; Cushman, p. 15, pl. 4, fig. 17. 1962. “Darbyella” sp. H; Ebensberger, S. 62, Taf. 6, Fig. 8, 9. 1962. Robulus wilcoxensis Cushman, Ponton; Hillebrandt, S. 56, Taf. 3, Fig. 24. 1965. Robulus wilcoxensis Cushman, Ponton; Pozaryska, p. 65, pl. 7, fig. 2, pl. 8, fig. 3. 1971. Lenticulina wilcoxensis Cushman, Ponton; Kaptarenko– Chernousova, p. 99, pl. 5, fig. 2 (in Ukrainian). 57 Фиг. 4. Стратиграфско разпространение на хиалинните тексони в разрез Неманци (легенда на фиг. 3) Fig. 4. Stratigraphical distribution of hyaline taxa in Nemanjici section (legend on Fig. 3) Fig. 5. Stratigraphical distribution of hyaline taxa in Ezevo Brdo and Kadrifakovo sections (legend on Fig. 3) Фиг. 5. Стратиграфско разпространение на хиалинните тексони в разрези Ежево бърдо и Кадрифаково (легенда на фиг. 3) 58 Fig. 6. Stratigraphical distribution of hyaline taxa in Madzarica section (legend on Fig. 3) Фиг. 6. Стратиграфско разпространение на хиалинните тексони в разрез Маджарица (легенда на фиг. 3) 2004. Lenticulina wilcoxensis Cushman, Ponton; Valchev, p. 54, pl. 1, fig. 12. Nomenclature. The holotype (Cushman Coll. No. 16186) is from the Eocene of Alabama. Material. More than 30 well preserved specimens. Description. The test is close-coiled. The last whorl is composed of 9–10 chambers separated by flush or slightly depressed sutures, curved backwards and branching at the peripheral area. The periphery is acute. The aperture is terminal, radiate. Distribution. The species is known from the Paleocene of the USA, Poland, the Alps, Bulgaria, the Upper Eocene of the USA. It was also established in the deep-sea cores from the Atlantic (Paleocene). Occurrence. Ovche Pole basin (Nemanjici section, samples 2, 3, 5, 9–18; Kadrifakovo section, samples 5, 6, 8, 10). Lenticulina yaguatensis (Bermudez, 1949) Material. About 30 specimens with different degree of preservation. Remarks. The species was described from Chardaklija section of the Ovche Pole basin (Džuranov et al., 1999, p. 62, pl. 1, Fig. 9). Occurrence. Ovche Pole basin (Nemanjici section, samples 5, 6, 10, 11, 16–18; Ezevo Brdo section, samples 2–4, 7, 8, 10–12; Kadrifakovo section, samples 1–3, 6, 8, 10). Lenticulina sp. Plate I, Fig. 4 Material. 9 badly preserved specimens. Description. The test is laterally depressed, closecoiled. The last whorl is composed of 6–7 chambers, separated with slightly depressed sutures curved backwards. The periphery is slightly keeled. The aperture is not distinct. Occurrence. Tikvesh basin (Krivolak section, samples 1, 2, 4), Delchevo basin (Crna Skala section, samples 7, 8). Genus Percultazonaria Loeblich and Tappan, 1986 Percultazonaria fragaria (Gümbel, 1868) Plate I, Fig. 5 1949. Marginulinopsis fragaria (Gumbel); Cuvillier, Szakall, p. 69, pl. 26, figs. 15, 17–19. 1970. Marginulina fragaria Gümbel; Shutskaya, pl. 11, fig. 4 (in Russian). 59 Fig. 7. Stratigraphical distribution of hyaline taxa in Hadzi Jusufli and Krivolak sections (legend on Fig. 3) Фиг. 7. Стратиграфско разпространение на хиалинните тексони в разрези Хаджи Юсуфли и Криволак (легенда на фиг. 3) 60 Fig. 8. Stratigraphical distribution of hyaline taxa in Rabrovo and Dedeli sections (legend on Fig. 3) Фиг. 8. Стратиграфско разпространение на хиалинните тексони в разрези Раброво и Дедели (легенда на фиг. 3) Fig. 9. Stratigraphical distribution of hyaline taxa in Crna Skala and Stuka sections (legend on Fig. 3) Фиг. 9. Стратиграфско разпространение на хиалинните тексони в разрези Црна скала и Щука (легенда на фиг. 3) 61 PLATE I 62 ТАБЛИЦА I PLATE I 1. Nodosaria ewaldi Reuss, 1851 Valandovo–Gevgelija basin, Dedeli section, upper flysh unit, sample 6; SEMx65 2. Nodosaria sp. Ovche Pole basin, Nemanjici section, upper flysh unit, sample 6; SEMx200 3. Lenticulina cf. wilcoxensis (Cushman and Ponton, 1932) Ovche Pole basin, Kadrifakovo section, upper flysh unit, sample 6; SEMx340 4. Lenticulina sp. Tikvesh basin, Krivolak section, upper flysh unit, sample 2; SEMx180 5. Percultazonaria fragaria (Gümbel, 1868) Ovche Pole basin, Nemanjici section, upper flysh unit, sample 10; SEMx110 6. Palmula budensis (Hantken, 1875) Ovche Pole basin, Nemanjici section, upper flysh unit, sample 5; SEMx180 7. Favulina hexagona (Williamson, 1848) Ovche Pole basin, Nemanjici section, upper flysh unit, sample 5; SEMx340 8. Globulina gibba d’Orbigny, 1826 Tikvesh basin, Krivolak section, upper flysh unit, sample 7; SEMx170 9. Guttulina irregularis (d’Orbigny, 1846) Tikvesh basin, Krivolak section, upper flysh unit, sample 5; SEMx200 10. Glandulina ovula d’Orbigny Ovche Pole basin, Nemanjici section, upper flysh unit, sample 5; SEMx400 11. Bolivina cf. antegressa Subbotina, 1953 Ovche Pole basin, Nemanjici section, upper flysh unit, sample 8; SEMx140 12, 13. Bolivina cf. cookei Cushman, 1922 Ovche Pole basin, Nemanjici section, upper flysh unit; 14, sample 13, SEMx130; 15, sample 8, SEMx250 14. Bolivina gracilis Cushman and Applin, 1926 Ovche Pole basin, Nemanjici section, upper flysh unit, sample 6; SEMx185 15. Bolivina nobilis Hantken, 1875 Ovche Pole basin, Nemanjici section, upper flysh unit, sample 12; SEMx325 16. Bolivina reticulata Hantken, 1875 Ovche Pole basin, Nemanjici section, upper flysh unit, sample 12; SEMx250 17. Bolivina scalprata Scwager, 1883 Ovche Pole basin, Nemanjici section, upper flysh unit, sample 10; SEMx250 Scale bar – 100 µm ТАБЛИЦА I 1. Nodosaria ewaldi Reuss, 1851 Валандово–Гевгелийски басейн, разрез Дедели, горна флишка задруга, пр. 6; SEMx65 2. Nodosaria sp. Овчеполски басейн, разрез Неманци, горна флишка задруга, пр. 6; SEMx200 3. Lenticulina cf. wilcoxensis (Cushman and Ponton, 1932) Овчеполски басейн, разрез Кадрифаково, горна флишка задруга, пр. 6; SEMx340 4. Lenticulina sp. Тиквешки басейн, разрез Криволак, горна флишка задруга, пр. 2; SEMx180 5. Percultazonaria fragaria (Gümbel, 1868) Овчеполски басейн, разрез Неманци, горна флишка задруга, пр. 10; SEMx110 6. Palmula budensis (Hantken, 1875) Овчеполски басейн, разрез Неманци, горна флишка задруга, пр. 5; SEMx180 7. Favulina hexagona (Williamson, 1848) Овчеполски басейн, разрез Неманци, горна флишка задруга, пр. 5; SEMx340 8. Globulina gibba d’Orbigny, 1826 Тиквешки басейн, разрез Криволак, горна флишка задруга, пр. 7; SEMx170 9. Guttulina irregularis (d’Orbigny, 1846) Тиквешки басейн, разрез Криволак, горна флишка задруга, пр. 5; SEMx200 10. Glandulina ovula d’Orbigny Овчеполски басейн, разрез Неманци, горна флишка задруга, пр. 5; SEMx400 11. Bolivina cf. antegressa Subbotina, 1953 Овчеполски басейн, разрез Неманци, горна флишка задруга, пр. 8; SEMx140 12, 13. Bolivina cf. cookei Cushman, 1922 Овчеполски басейн, разрез Неманци, горна флишка задруга, 14 – пр. 13, SEMx130; 15 – пр. 8, SEMx250 14. Bolivina gracilis Cushman and Applin, 1926 Овчеполски басейн, разрез Неманци, горна флишка задруга, пр. 6; SEMx185 15. Bolivina nobilis Hantken, 1875 Овчеполски басейн, разрез Неманци, горна флишка задруга, пр. 12; SEMx325 16. Bolivina reticulata Hantken, 1875 Овчеполски басейн, разрез Неманци, горна флишка задруга, пр. 12; SEMx250 17. Bolivina scalprata Scwager, 1883 Овчеполски басейн, разрез Неманци, горна флишка задруга, пр. 10; SEMx250 63 1975. Marginulina fragaria Gümbel; Braga et al., p. 93, t. 1, fig. 35. 1983. “Cristellaria fragaria Gümb.”; Geroch, Verdenius, pl. 3, fig. 3. 1984. Marginulina fragaria Gümbel; Olszewska, Szymakow­ ska, p. 129, tabl. 1, fig. 17. 1985. Marginulinopsis fragaria (Gümbel); Grünig, p. 262, pl. 5, figs. 24–28. 1991. Marginulinopsis fragaria (Gümbel); Barbin, Keller– Grünig, pl. 2, figs. 6–12. 2006. Percultazonaria fragaria (Gümbel); Cimerman et al., p. 24, pl. 5, figs 4, 5. Nomenclature. The species was first described from the Eocene of the Alpes. Here it is identified after the images of Cimerman et al. (2006, pl. 5, Figs. 4, 5). Material. 15 very well preserved specimens. Description. The test is heteromorphous, slighltly inflated, with planispiral initial portion and short rectilinear later one. The chambers are broad and low. The periphery is subacute. The sutures are elevated and ornamented by nodes. The aperture is terminal, radiate. Remarks. We refered our specimens to genus Per­ cultazonaria Loeblich and Tappan because of the presence of nodes covering the sutures. Distribution. The species is known from the Paleocene and Eocene of Spain, California, the Upper Eocene of SE United States, the Eocene to Miocene of the Carpathians, France, the Paleocene to Oligocene of Italy, the Eocene of Caucasus, Crimea and Middle Asia. Occurrence. Ovche Pole basin (Nemanjici section, samples 1, 2, 5, 6, 9, 10, 12–14, 16–18). Subfamily PALMULINAE Saidova, 1981 Genus Palmula Lea, 1833 Palmula budensis (Hantken, 1875) Plate I, Fig. 6 1949. Frondicularia budensis (Hantken); Cuvillier, Szakall, p. 84, pl. 30, fig. 14. 1969. Frondicularia budensis (Hantken); Krayeva, Zernetskij, p. 52, pl. 17, figs. 1, 2a–b (in Russian). 1975. Frondicularia budensis (Hantken); Braga et al., p. 104. 1984b. Plectofrondicularia budensis (Hantken); Olszewska, p. 57, Tabl. 1, fig. 12. 1984. Plectofrondicularia budensis (Hantken); Olszewska, Szy­makowska, p. 130, Tabl. 1, fig. 13. 1985. Frondicularia budensis (Hantken). Grünig, p. 261, pl. 4, fig. 21. 1992. Palmula budensis (Hantken); Darakchieva, Juranov, p. 12, pl. I, fig. 9. not fig. 10. 1993. Palmula budensis (Hantken); Sztrаkos (In: Mathelin, Sztrаkos), p. 76, pl. 9, fig. 10. 2003. Palmula budensis (Hantken); Horvath, p. 18, pl. 3, fig. 8, pl. 4, fig. 8. 2006. Palmula budensis (Hantken); Cimerman et al., p. 24, pl. 5, fig. 7. Nomenclature. The species was first described from the Eocene of Hungary (Clavulina Szaboi Schichten). 64 Here it is identified after the SEM images of Horvath (2003, pl. 3, Fig. 8, pl. 4, Fig. 8). Material. 10 specimens with different degree of preser­ vation. Description. Test is elongate, flattened, with planispirally coiled early portion. Later the arrangement of the chambers becomes uncoiled, rectilinear. The chambers are broad, low and arched, 12–15 in number. Sutures are flush. Aperture is terminal, radiate. Distribution. The species is known from the Eocene of Bulgaria, Italy, Spain, France, the Upper Eocene of Slovenia, the Eocene and Oligocene of Poland, the Paleogene of Ukraine. Occurrence. Ovche Pole basin (Nemanjici section, samples 4–6, 11–13, 16, 17). Family LAGENIDAE Reuss, 1862 Genus Lagena Walker and Jacobs, 1798 Lagena humifera Bandy, 1949 Material. 7 very well preserved specimens. Remarks. The species was described from Chardaklija section of the Ovche Pole basin (Džuranov et al., 1999, p. 62, pl. 1, Fig. 10). Occurrence. Ovche Pole basin (Madzarica section, samples 10–12). Lagena striata (d’Orbigny, 1839) Material. 20 very well preserved specimens. Remarks. The species was described from Chardaklija section of the Ovche Pole basin (Džuranov et al., 1999, p. 62, pl. 1, Fig. 11). Occurrence. Skopje–Kumanovo basin (Vojnik section, samples 3, 4, 6, 7), Ovche Pole basin (Ezovo Brdo section, samples 7, 8, 11, 12; Kadrifakovo section, sam­ ples 3, 4, 7, 8; Madzarica section, samples 1, 2, 5). Family POLYMORPHINIDAE d’Orbigny, 1839 Subfamily POLYMORPHININAE d’Orbigny, 1839 Genus Globulina d’Orbigny, 1839 Globulina gibba d’Orbigny, 1826 Plate I, Fig. 8 1846. Globulina Gibba d’Orbigny; d’Orbigny, p. 227, tab. 13, fig. 13, 14. 1882. Globulina gibba d’Orbigny; Terquem, p. 130, pl. 13, figs. 22–27. 1934. Globulina gibba d’Orbigny; Cushman, Dusenbery, p. 59, pl. 8, fig. 4. 1948. Globulina gibba d’Orbigny; Brotzen, p. 46, fig. 10 – 1, 2. 1949. Globulina gibba d’Orbigny; Cuvillier, Szakall, p. 87, pl. 31, fig. 2. 1950. Globulina gibba d’Orbigny; Vassilenko, p. 196, pl. 2, figs. 1a–c (in Russian). 1956. Globulina gibba d’Orbigny; Haque, p. 107, pl. 30, fig. 4. 1960. Globulina gibba d’Orbigny; Olsson, p. 25, pl. 3, fig. 25. 1961. Globulina gibba d’Orbigny; Kaasschieter, p. 183, pl. 8, fig. 6–7. 1962. Globulina gibba d’Orbigny; Kiesel, S. 48, Taf. 7, Fig. 10. 1965. Globulina gibba d’Orbigny; Pozaryska, p. 86, pl. 13, fig. 3. 1969. Globulina gibba d’Orbigny; Krayeva, Zernetskij, p. 67, pl. 23, fig. 4 (in Russian). 1970. Globulina gibba d’Orbigny; Le Calvez, p. 84, pl. 17, figs. 3–4. 1984a. Globulina gibba d’Orbigny; Olszewska, p. 21, tabl. 2, fig. 12. 1985. Globulina gibba d’Orbigny; Grünig, p. 263, pl. 6, figs. 7–9. 1985. Globulina gibba d’Orbigny; Papp, Schmidt, p. 79, pl. 71, figs. 9–12. 1992. Globulina gibba d’Orbigny; Darakchieva, Juranov, p. 16, pl. 3, fig. 4. 2005. Globulina gibba d’Orbigny; Valchev, p. 161, pl. 1, fig. 1. Nomenclature. The species was first described from the Eocene of France (Bordeaux vicinities). Here it is identified after the refigured d’Orbigny’s specimens (Papp, Schmidt, 1985, pl. 71, Figs. 9–12). Material. About 25 specimens, most of them well preserved. Description. The test is rounded; the later chambers embrace the earlier ones. The sutures are slightly depressed, curved. The surface is smooth. The aperture is terminal, broad-ovate, radiate. Distribution. The species is known from the Senonian of Germany, the Paleocene of Alabama, the Netherlands, Sweden, Ukraine, Caucasus, Australia, the upper Paleocene of England, the Paleocene and Eocene of Bulgaria, the Eocene of Belgium, the Lower and Middle Eocene of France, the Upper Eocene of Ukraine, Caucasus, England, the Oligocene of Germany, Hungary, the Miocene of the Vienna Basin, the Lower Miocene of Dominican Republic, the Middle Miocene of the central Paratethys. Widely distributed in recent sediments. Occurrence. Ovche Pole basin (Nemanjici section, samples 3, 4, 7–9, 13, 14). 2005. Guttulina irregularis (d’Orbigny, 1846); Valchev, p. 162, pl. 1, fig. 4. Nomenclature. A holotype was not designated. The species was first described from the Badenian of the Vienna Basin. Here it is identified after the refigured d’Orbigny’s specimens (Papp, Schmidt, 1985, pl. 71, Figs. 1–4). Material. 8 well preserved specimens. Description. The test is inflated, asymmetrical, rounded in outline. Three chambers are visible on both sides. The sutures are slightly depressed, curved. The surface is smooth. The aperture is terminal, narrowovate, and radiate. Distribution. It is known from the Paleocene of the Netherlands, the Paleocene and Eocene of Bulgaria, the Middle Eocene of France, the Upper Eocene of Ukraine, England, USA, the Oligocene of Germany, the Miocene of Austria. Occurrence. Tikvesh basin (Krivolak section, samples 2, 4, 5). Family ELLIPSOLAGENIDAE A. Silvestri, 1923 Subfamily OOLININAE Loeblich and Tappan, 1961 Genus Favulina Patterson and Richarson, 1987 Favulina hexagona (Williamson, 1848) Plate I, Fig. 7 Genus Guttulina d’Orbigny, 1839 Guttulina irregularis (d’Orbigny, 1846) Plate I, Fig. 9 1956. Lagena hexagona (Williamson); Hagn, S. 141, Taf. 10, Fig. 22. 1959. Lagena hexagona (Williamson); Stancheva, p. 326, pl. 1, fig. 6 (in Bulgarian). 1969. Lagena hexagona (Williamson); Krayeva, Zernetskij, p. 48, pl. 16, fig. 11 (in Russian). 1971. Lagena hexagona (Williamson); Kaptarenko-Chernou­ sova, p. 52, pl. 1, fig. 12 (in Ukrainian). 1988. Favulina hexagona (Williamson); Loeblich, Tappan, pl. 462, figs. 1, 2. 1992. Lagena hexagona (Williamson); Darakchieva, Juranov, p. 15, pl. 2, fig. 6. 1999. Favulina hexagona (Williamson); Darakchieva, p. 36. 2002. Favulina hexagona (Williamson); Valchev, p. 25, pl. 1, fig. 11 (in Bulgarian). 1846. Globulina irregularis d’Orbigny; d’Orbigny, p. 266, tab. 13, fig. 9–10. 1935. Guttulina irregularis (d`Orbigny); Cushman, p. 24, pl. 9, figs. 13–16. 1959. Guttulina irregularis (d`Orbigny); Mallory, p. 177, pl. 14, fig. 13. 1961. Guttulina irregularis (d`Orbigny); Kaasschieter, p. 181, pl. 8, figs. 2–3. 1969. Guttulina irregularis (d’Orbigny); Krayeva, Zernetskij, p. 66, pl. 24, figs. 3, 4 (in Russian). 1970. Guttulina irregularis (d’Orbigny); Le Calvez, p. 92, pl. 20, fig. 3. 1985. Guttulina communis (d’Orbigny); Papp, Schmidt, p. 79, pl. 71, figs. 1–4. 1992. Guttulina irregularis (d’Orbigny); Darakchieva, Juranov, p. 17, pl. 3, fig. 1. Nomenclature. The species was first described from recent sediments of Scotland. Here it is identified after the images of Loeblich, Tappan (1988, pl. 462, Figs. 1, 2). Material. 15 very well preserved specimens. Description. The test is unilocular, moderately inflated, slightly elongated. The surface is covered with regular hexagonal reticulations. The aperture is terminal, on a short neck. Distribution. The species is known from the Maast­ richtian and Paleocene of the Netherlands, the Paleo­ cene of New Jersey, Ukraine, Poland, Australia, East Kamchatka, the upper Paleocene of England, the Paleocene, Eocene and Oligocene of Bulgaria, the Eocene of Germany, USA, the Upper Eocene of Donbas 65 basin, Hungary, the Oligocene of the Netherlands, Hungary, the Miocene of the Vienna basin, Bavaria, Dominican Republic, the Middle Miocene of the Netherlands. Nowadays it lives at depth 520–4200 m. Occurrence. Ovche Pole basin (Nemanjici section, samples 1, 2, 5, 6; Madzarica section, samples 11–13). Family GLANDULINIDAE Reuss, 1860 Subfamily GLANDULININAE Reuss, 1860 Genus Glandulina d’Orbigny, 1839 Glandulina ovula d’Orbigny, 1846 Plate I, Fig. 10 1846. Glandulina Laevigata d’Orbigny; d’Orbigny, p. 29, tab. 1, fig. 4, 5. 1846. Glandulina Ovula d’Orbigny; d’Orbigny, p. 29, tab. 1, fig. 6, 5. 1930. Glandulina laevigata d`Orbigny; Cushman, Ozawa, p. 143, pl. 40, figs. 1a–b. 1956. Glandulina laevigata d`Orbigny; Haque, p. 103, pl. 11, figs. 13–14. 1961. Glandulina laevigata d`Orbigny; Kaasschieter, p. 187, pl. 8, fig. 17. 1962. Glandulina laevigata d`Orbigny; Kiesel, S. 51, taf. 8, fig. 3. 1965. Glandulina laevigata d`Orbigny; Pozaryska, p. 91, pl. 22, fig. 7. 1985. Glandulina ovula d’Orbigny; Papp, Schmidt, p. 21, pl. 2, figs. 1–9. 1992. Glandulina laevigata d’Orbigny; Darakchieva, Juranov, p. 18, pl. 3, fig. 3. 2005. Glandulina laevigata d’Orbigny, 1846; Valchev, p. 163, pl. 1, fig. 8 (with synonymy). Nomenclature. The species was first described from the Badenian of the Vienna Basin. The lectotype was figured by Papp, Schmidt (1985, pl. 2, Fig. 8). Material. 41 well preserved specimens. Description. The test is elongate-oval, round in cross section. Both ends are tapered. The surface is smooth. The aperture is terminal, radiate. Remarks. We accept the Papp, Schmidt’s (1985) opinion that G. laevigata d’Orbigny is subjective junior synonym of G. ovula d’Orbigny. Distribution. The species is known from the Upper Cretaceous of Poland, the Upper Paleocene of England, the Eocene of Bulgaria, the Eocene and Oligocene of Belgium, the Upper Eocene of Ukraine, Caucasus, England, Poland, the Oligocene of Turkmenia, the Miocene of Vienna Basin and Bulgaria. It was also established in deep sea holes in Southeast Atlantic (Upper Eocene). Nowadays it is cosmopolitan. Occurrence. Ovche Pole basin (Nemanjici section, samples 2–6, 13–18). Suborder ROTALIINA Delage and Herouard, 1896 Superfamily BOLIVINACEA Glaessner, 1937 Family BOLIVINIDAE Glaessner, 1937 Genus Bolivina d’Orbigny, 1839 66 Bolivina cf. antegressa Subbotina, 1953 Plate I, Fig. 11 1953. Bolivina antegressa n. sp.; Subbotina, p. 226, pl. 10, figs. 11–16 (in Russian). 1975. Bolivina antegressa Subbotina; Braga et al., p. 94. 1978. Bolivina antegressa Subbotina; Proto Decima, Bolli, p. 790, pl. 2, fig. 2. 1985. Bolivina antegressa Subbotina; Grünig, p. 264. 1988. Bolivina antegressa Subbotina; Parisi, Coccioni, pl. 2, figs. 3–4. Nomenclature. The holotype (VNIGRI Coll. No. 3257) is from the Upper Eocene of Crimea. Material. About 30 specimens, most of them well preserved. Description. The test is biserial, elongate, tapered at the initial part, then broadening slowly. The chambers are low. The surface is covered with irregular longitudinal costae. The periphery is subacute. The apertute is low, loop-shaped, basal. Remarks. Our specimens are more elongate than the holotype. Distribution. The species is known from the Paleocene– Lower Oligocene of Italy, the Eocene of France, Spain, the Atlantic, the Upper Eocene of Crimea. Occurrence. Ovche Pole basin (Nemanjici section, samples 3, 4, 7–9, 15–18), Tikvesh basin (Krivolak section, samples 1, 4, 6, 7). Bolivina cf. cookei Cushman, 1922 Plate I, Figs. 12, 13 1961. Bolivina cookei Cushman; Kaasschieter, p. 195, pl. 8, figs. 25–26. 1975. Bolivina cookei Cushman; Samuel, p. 133, pl. 74, fig. 3. 1977. Bolivina cookei Cushman; Pozaryska, p. 27, pl. 9, fig. 3. 1984. Bolivina cookei Cushman; Odrzywolska–Bienkowa, Po­ zaryska, p. 128, pl. 3, figs. 9–10. 1984a. Bolivina cookei Cushman; Olszewska, tabl. 8, fig. 9. Nomenclature. The species was first described from the Oligocene of the USA (Mississippi). Here it is identified after the Kaasschieter’s images (1961, pl. 8, Figs. 25–26). Material. 20 well preserved specimens. Description. The test is biserial, slightly tapered at the initial part. The sutures are oblique, curved and slightly depressed, the chambers are moderately high. Remarks. Our specimens are characterized by the lack of distinct longitudinal costae that are typical for the species. Distribution. The species is known from the Middle– Upper Eocene of Belgium, the Upper Eocene and Oligocene of the Paris basin, the Carpathians (Poland, Slovakia) and the USA. Occurrence. Ovche Pole basin (Nemanjici section, samples 4, 5, 12–14), Valandovo–Gevgelija basin (Rab­ rovo section, samples 6, 9, 14; Dedeli section, samples 5, 8, 12), Delchevo basin (Crna Skala Section, samples 9, 12, 18, 25). Bolivina gracilis Cushman and Applin, 1926 Plate I, Fig. 14 1935. Bolivina gracilis Cushman et Applin; Cushman, p. 37, pl. 14, figs. 8–10. 1975. Bolivina gracilis Cushman et Applin; Braga et al., p. 106. 1985. Bolivina gracilis Cushman et Applin; Grünig, p. 265, pl. 5, figs. 9–10. Nomenclature. The species was first described from the Eocene of Texas. Here it is identified after the images of Grünig (1985, pl. 5, Figs. 9–10). Material. 15 specimens with different degree of preser­ vation. Description. The test is slender, tapered at the distal end. The chambers are distinct, separated by oblique depressed sutures. Initial part of the test is covered with indistinct longitudinal costae, while the later one is smooth. Remarks. Grünig (1985) noted the presence of intermediate forms between B. gracilis Cushman and Applin and B. nobilis Hantken and she pointed out that it is ecologically controlled lineage. We are not able to confirm or reject this statement, because our observation is based on limited number of specimens. Distribution. The species is known from the Eocene of Italy, France, Spain, and the USA. Occurrence. Ovche Pole basin (Nemanjici section, samples 2, 3, 6), Tikvesh basin (Hadzi Jusufli section, samples 3, 5, 6; Krivolak section, samples 1, 2, 4, 6). Bolivina nobilis Hantken, 1875 Plate I, Fig. 15 1947. Bolivina nobilis Hantken; Subbotina, p. 96, pl. 9, fig. 3 (in Russian). 1957. Bolivina nobilis Hantken; Sacal, Debourle, p. 27, pl. 8, fig. 20. 1970. Bolivina nobilis Hantken; Le Calvez, p. 115. 1975. Bolivina nobilis Hantken; Braga et al., p. 106, t. 5, figs. 1–2. 1975. Bolivina nobilis Hantken; Samuel, p. 134, pl. 74, figs. 4–6. 1983. Bolivina nobilis Hantken; Krhovsky, p. 77, pl. 1, fig. 8. 1984. Bolivina nobilis Hantken; Odrzywolska-Bienkowa, Po­ zaryska, p. 129, pl. 3, fig. 7. 1984. Bolivina nobilis Hantken; Olszewska, Szymakowska, p. 130, tabl. 1, fig. 15. 1985. Bolivina nobilis Hantken; Grünig, p. 265, pl. 5, figs. 12–14. 1990. Bolivina nobilis Hantken; Barbieri, t. 3, fig. 3. 2006. Bolivina nobilis Hantken; Cimerman et al., p. 26, pl. 6, figs. 9, 10. Nomenclature. The species was first describen from Eocene of Hungary (Clavulina Szaboi Schichten). Here it is identified after the images of Grünig (1985, pl. 5, Figs. 12–14). Material. Мore than 20 specimens with different degree of preservation. Description. The test is slightly elongated with almost parallel sides. The periphery is slightly acute. The surface is covered with fine longitudinal costae that become indistinct at the later part of the test. Distribution. The species is known from the Eocene of Italy, France, Spain, the Upper Eocene of Libya, Caucasus and Middle Asia, the Eocene and Oligocene of the Carpathians, the Oligocene of the Paris basin, the Miocene of France. Occurrence. Ovche Pole basin (Nemanjici section, samples 2–6, 9, 11–18; Madzarica section, samples 14–16). Bolivina reticulata Hantken, 1875 Plate I, Fig. 16 1957. Bolivina reticulata Hantken; Sacal, Debourle, p. 27, pl. 8, fig. 13. 1962. Bolivina reticulata Hantken; Kiesel, S. 61, Taf. 9, Fig. 8. 1975. Bolivina reticulata Hantken; Samuel, p. 134, pl. 73, figs. 7, 8. 1980. Bolivina reticulata Hantken; Jutson, p. 378, pl. 1, fig. 16. 1984a. Bolivina reticulata Hantken; Olszewska, p. 23, tabl. 3, fig. 5. 1984b. Bolivina reticulata Hantken; Olszewska, p. 58, tabl. 1, fig. 7. 1984. Bolivina reticulata Hantken; Olszewska, Szymakowska, p. 131, tabl. 1, fig. 18. 1985. Bolivina reticulata Hantken; Olszewska, p. 227, pl. 1, fig. 13. Nomenclature. The species was first described from Eocene of Hungary (Clavulina Szaboi Schichten). Here it is identified after the images of Samuel (1975, pl. 73, Figs. 7, 8). Material. 20 specimens, most of them with broken periphery. Description. The test is tapered at the initial part. The periphery is narrowly rounded. The chambers are low, separated by oblique, straight, and slightly depressed sutures. The surface is covered with simple longitudinal reticulations. Distribution. The species is known from the Upper Eocene–Miocene of the Carpathians, the Oligocene of Germany, the Oligocene–Miocene of Spain, the Miocene of France. Occurrence. Ovche Pole basin (Nemanjici section, samples 4, 5, 7, 12–15, 17, 18; Madzarica section, samples 13–16). Bolivina scalprata Schwager, 1883 Plate I, Fig. 17 1960. Bolivina scalprata Schwager; Subbotina et al., p. 110, pl. 7, figs. 1, 2 (in Russian). 67 1984a. Bolivina scalprata Schwager; Olszewska, p. 23, tabl. 3, fig. 6. Nomenclature. The species was first described from the Eocene of Egypt. Here it is identified after the images of Subbotina et al. (1960, pl. 7, Figs. 1, 2). Material. Аbout 20 specimens with different degree of preservation. Description. The test is elongated, tapered at the initial part and gradually broadening towards the late part. The chambers are distinct, low, separated by oblique and straight sutures, which are ornamented with fine and sharp costae. The chamber surface is also covered with irregularly distributed costae. The periphery bears a narrow keel. Distribution. The species is known from the Eocene of Egypt, the Eocene–Oligocene of the Carpathians, the Miocene of Hungary. Occurrence. Ovche Pole basin (Nemanjici section, samples 3, 4, 6, 7, 10, 12–16). Superfamily BULIMINACEA Jones, 1875 Family BULIMINIDAE Jones, 1875 Genus Bulimina d’Orbigny, 1826 Bulimina sculptilis Cushman, 1923 Plate II, Fig. 1 1947. Bulimina sculptilis Cushman; Cushman, Parker, p. 103, pl. 24, fig. 12. 1947. Bulimina sculptilis Cushman; Subbotina, p. 93, pl. 8, figs. 27–29 (in Russian). 1948. Bulimina sculptilis Cushman; Chalilov, pl. 12, fig. 7 (in Russian). 1953. Bulimina sculptilis Cushman; Subbotina, p. 212, pl. 9, figs. 17–19 (in Russian). 1968. Bulimina sculptilis Cushman; Hofker, p. 5, pl. 1, figs. 2–3. Nomenclature. The species was first described from the Oligocene of Mississippi (USA). Here it is identified after the image of Cushman, Parker (1947, pl. 24, Fig. 12). Material. 18 well preserved specimens. Description. The test is moderately elongate, slightly tapered at the initial part. The sutures are horizontal, the surface is ornamented with longitudinal straight, sharp and thin costae beginning at the base of the last formed chambers. The aperture is broad, loop-shaped. Distribution. The species is known from the Eocene of the Alps, the Upper Eocene and Oligocene of Caucasus and Middle Asia, the Oligocene of Ecuador, the Eocene to Miocene of the USA, the Olicocene to Miocene of Venezuela. Occurrence. Tikvesh basin (Hadzi Jusufli section, samples 2, 7; Krivolak section, samples 3–5). Bulimina trigona Terquem, 1882 Material. 25 specimens, most of them well preserved. 68 Remarks. The species was described from Chardaklija section of the Ovche Pole basin (Džuranov et al., 1999, p. 63, pl. 1, Fig. 22). Occurrence. Ovche Pole basin (Nemanjici section, samples 2–3, 12–14, 16–18; Ezevo Brdo section, samples 2–4, 7–12; Madzarica section, samples 12–14), Strumica basin (Stuka section, samples 2–5). Superfamily FURSENKOINACEA Loeblich and Tappan, 1961 Family FURSENKOINIDAE Loeblich and Tappan, 1961 Genus Fursenkoina Loeblich and Tappan, 1961 Fursenkoina dibollensis (Cushman et Applin, 1926) Material. About 50 specimens with different degree of preservation. Remarks. The species was described from Chardaklija section of the Ovche Pole basin (Džuranov et al., 1999, p. 63, pl. 1, Figs. 12, 13). Occurrence. Skopje–Kumanovo basin (Vojnik section, samples 4–6), Ovche Pole basin (Ezevo Brdo section, samples 7, 8, 10, 12; Kadrifakovo section, samples 1–5, 7; Madzarica section, samples 7-10), Strumica basin (Stuka section, samples 4–5). Superfamily DELOSINACEA Parr, 1950 Family CAUCASINIDAE N. K. Bykova, 1959 Subfamily BAGGATELLINAE N. K. Bykova, 1959 Genus Caucasina Khalilov, 1951 Caucasina eocaenica Chalilov, 1958 Plate II, Fig. 2 1967. Caucasina eocaenica Chalilov; Chalilov, p. 164, pl. 36, fig. 2 (in Russian). 1996. Caucasina eocaenica Chalilov; Sztrakos, pl. 3, figs. 27–28. 2004. Caucasina eocaenica Chalilov; Bugrova, p. 98, pl. 39, fig. 13 (in Russian). Nomenclature. The species was first described from the Upper Eocene of Azerbaijan. Here it is identified after the image of Chalilov (1967, pl. 36, Fig. 2). Material. Аbout 20 well preserved specimens. Description. The test is elongate, subcylindrical, high trochospiral. The chambers are slightly inflated, those from the last whorl comprise 1/3 of the test length. The sutures are almost horizontal, depressed. The surface is smooth. The aperture is elongate, loop-shaped. Distribution. The species is known from the Paleocene and Eocene of France, the Eocene of Crimea, Caucasus, Turkmenia, Azerbaijan. Occurrence. Ovche Pole basin (Nemanjici section, samples 2, 3, 5–7, 13–16), Delchevo basin (Crna Skala section, samples 21, 23, 25, 26). Caucasina tenebricosa Pishvanova, 1960 Plate II, Fig. 3 1960. Caucasina tenebricosa Pischvanova, sp. n.; Subbotina et al., p. 90, pl. 6, fig. 2 (in Russian). 1960. Caucasina tenebricosa Pischvanova; Subbotina, p. 212, pl. 4, figs. 2–4 (in Russian). 1972. Caucasina tenebricosa Pischvanova; Pishvanova, p. 265, pl. 21, fig. 1 (in Russian). 1975. Caucasina tenebricosa Pischvanova; Samuel, p. 140, pl. 80, fig. 11. 1979. Caucasina cf. tenebricosa Pischvanova; Sztrakos, p. 87, pl. 27, fig. 12. 1984a. Caucasina tenebricosa Pischvanova; Olszewska, tabl. 9, fig. 17. 1984b. Caucasina tenebricosa Pischvanova; Olszewska, p. 63, tabl. 4, fig. 3. 1984. Caucasina tenebricosa Pischvanova; Olszewska, Szy­ makowska, p. 133, tabl. 2, fig. 4. Description. The test is uniserial, slender, composed of globular chambers connected with short and thin neck. The surface is smooth. The aperture is round, terminal, on a short neck. Distribution. It is known from the Upper Senonian of Germany, the Paleocene of Texas (Midway Formation), Mexico, Poland, Bulgaria, the Eocene of Bulgaria, Ukraine, Caucasus, Crimea, Turkmenia, California, Venezuela, the Middle Eocene of Dalmatian, the Oligocene of the Netherlands, Hungary, the Miocene of the Vienna Basin, Bavaria, Bulgaria. Occurrence. Ovche Pole basin (Nemanjici section, samples 3, 4, 10, 14–16; Kadrifakovo section, samples 5–6, 8–9). Nomenclature. The holotype (VNIGRI Coll. No. 5197) is from the Oligocene of the Precarpathian area (Dzvinyach Village). Material. 15 well preserved specimens. Description. The test is elongate, tapered at the initial part, gradually broadening towards the apertural end. The chambers are inflated, increasing gradually in size. The sutures are deep, especially at the initial part. The surface is smooth. The aperture is broad, elongate, loop-shaped. Distribution. The species is known from the Upper Eocene–Miocene of the Carpathians, the Oligocene– Miocene of the Precarpathian region. Occurrence. Ovche Pole basin (Nemanjici section, samples 2, 3, 7, 8, 13–16). Superfamily DISCORBACEA Ehrenberg, 1838 Family BAGGINIDAE Cushman, 1927 Subfamily BAGGININAE Cushman, 1927 Genus Baggina Cushman, 1926 Baggina subconica (Terquem, 1882) Plate II, Figs. 4, 5 Superfamily STILOSTOMELLACEA Finlay, 1947 Family STILOSTOMELLIDAE Finlay, 1947 Genus Siphonodosaria A. Silvestri, 1924 Siphonodosaria adolphina (d’Orbigny, 1846) Plate II, Fig. 6 1846. Dentalina Adolphina d’Orbigny; d’Orbigny, p. 51, tab. 2, fig. 18–20. 1926. Nodosaria cf. adolphina (d’Orbigny); Cushman, p. 597, pl. 18, fig. 2. 1951. Dentalina adolphina d’Orbigny; Spasov, p. 100, pl. 1, fig. 11 (in Bulgarian). 1953. Siphonodosaria adolphina (d’Orbigny); Subbotina, p. 180, pl. 6, figs. 1, 2 (in Russian). 1969. Nodosaria adolphina (d’Orbigny); Krayeva, Zernetskij, p. 41, pl. 14, fig. 6 (in Russian). 1985. Stilostomella adolphina (d’Orbigny); Papp, Schmidt, p. 31, pl. 14, figs. 8–11. 1992. Siphonodosaria adolphina (d’Orbigny); Darakchieva, Juranov, p. 30, pl. 5, fig. 4. 2007. Siphonodosaria adolphina (d’Orbigny); Valchev, p. 133, pl. 1, fig. 14. Nomenclature. The species was first described from the Badenian of the Vienna Basin. The lectotype (GBA Coll. No. 1981/03/58) was designated by Papp, Schmidt (1985, pl. 14, Fig. 9). Material. 13 well preserved specimens. 1882. Rotalina subconica Terq.; Terquem, p. 61, pl. 4, fig. 5. 1961. Cancris subconicus (Terquem); Kaasschieter, p. 213, pl. 12, figs. 6–8. 1970. Cancris subconicus (Terquem); Le Calvez, p. 145, pl. 43, fig. 6. 1975. Cancris subconicus (Terquem); Samuel, p. 141, pl. 82, fig. 4. 1977. Baggina subconica (Terquem); Pozaryska, p. 33, pl. 11, fig. 1. 1981. Cancris subconicus (Terquem); Hughes, p. 202, pl. 15.4, figs. 5–7. 1983. Cancris subconicus (Terquem); King, p. 29, pl. 3, figs. 16–17. 1983. Cancris aff. subconicus (Terquem); Krhovsky, p. 79, pl. 4, figs. 3–5. 1984. Baggina subconica (Terquem); Odrzywolska-Bienkowa, Pozaryska, p. 134, pl. 6, figs. 4, 8–9. 1984a. Baggina subconica (Terquem); Olszewska, p. 25, tabl. 4, fig. 1. Nomenclature. The holotype is the specimen figured by Terquem (1882, pl. 4, Fig. 5). The species was first described from the Eocene (Lutetian) of the Paris Basin. Material. 17 specimens with different degree of preservation. Description. The test is very low trochospirally coiled, with 6–7 chambers in the last whorl. The final chamber comprises one-half of it. The periphery is broadly rounded. The surface is smooth. The aperture is broad, umbilical, at the base of the last chamber. Distribution. The species is known from the Eocene of Belgium, France, the Eocene and Oligocene of the Carpathians and the North Sea. Occurrence. Ovche Pole basin (Nemanjici section, samples 5–7, 13–15), Delchevo basin (Crna Skala section, samples 4, 5). 69 PLATE II 70 ТАБЛИЦА II PLATE II 1. Bulimina sculptilis Cushman, 1923 Ovche Pole basin, Nemanjici section, upper flysh unit, sample 6; SEMx300 2. Caucasina eocenica Chalilov, 1958 Ovche Pole basin, Nemanjici section, upper flysh unit, sample 6; SEMx240 3. Caucasina tenebricosa Pishvanova, 1960 Ovche Pole basin, Nemanjici section, upper flysh unit, sample 14; SEMx300 4, 5. Baggina subconica (Terquem, 1882) 8, Delchevo basin, Crna Skala section, upper flysh unit, sample 5, SEMx150; 9, Ovche Pole basin, Nemanjici section, upper flysh unit, sample 7, SEMx180 6. Siphonodosaria adolphina (d’Orbigny, 1846) Ovche Pole basin, Nemanjici section, upper flysh unit, sample 16; SEMx110 7. Valvulineria jacksoensis Cushman, 1933 Delchevo basin, Crna Skala section, upper flysh unit, sample 24; SEMx270 8. Eponides sp. Ovche Pole basin, Nemanjici section, upper flysh unit, sample 10; SEMx200 9, 10. Cibicidoides sp. 9, Ovche Pole basin, Nemanjici section, upper flysh unit, sample 4, spiral view, SEMx200; 10, Tikvesh basin, Krivolak section, upper flysh unit, sample 3, umbilical view, SEMx140 11, 12. Cibicides lobatulus (Walker and Jakobs, 1798) Tikvesh basin, Hadzi Jusufli section, upper flysh unit, sample 7: 11, spiral view; SEMx180; 12, umbilical view; SEMx160 13–15. Cibicides ungerianus (d’Orbigny, 1846) 13, 14, Delchevo basin, Crna Skla section, upper flysh unit, sample 24: 13, spiral view, SEMx200; 14, umbilical view, SEMx250; 15, Ovche Pole basin, Nemanjici section, upper flysh unit, sample 3, umbilical view, SEMx160 16, 17. Cibicides sp. Tikvesh basin, Krivolak section, upper flysh unit, sample 6: 16, umbilical view, SEMx160; 17, spiral view, SEMx180 Scale bar – 100 µm ТАБЛИЦА II 1. Bulimina sculptilis Cushman, 1923 Овчеполски басейн, разрез Неманци, горна флишка задруга, пр. 6; SEMx300 2. Caucasina eocenica Chalilov, 1958 Овчеполски басейн, разрез Неманци, горна флишка задруга, пр. 6; SEMx240 3. Caucasina tenebricosa Pishvanova, 1960 Овчеполски басейн, разрез Неманци, горна флишка задруга, пр. 14; SEMx300 4, 5. Baggina subconica (Terquem, 1882) 8 – Делчевски басейн, разрез Црна скала, горна флишка задруга, пр. 5, SEMx150; 9 – Овчеполски басейн, разрез Неманици, горна флишка задруга, пр. 7, SEMx180 6. Siphonodosaria adolphina (d’Orbigny, 1846) Овчеполски басейн, разрез Неманци, горна флишка задруга, пр. 16; SEMx110 7. Valvulineria jacksoensis Cushman, 1933 Делчевски басейн, разрез Црна скала, горна флишка задруга, пр. 24; SEMx270 8. Eponides sp. Овчеполски басейн, разрез Неманци, горна флишка задруга, пр. 10; SEMx200 9, 10. Cibicidoides sp. 9 – Овчеполски басейн, разрез Неманци, горна флишка задруга, пр. 4, спирална страна, SEMx200; 10 – Тиквешки басейн, разрез Криволак, горна флишка задруга, пр. 3, умбиликална страна, SEMx140 11, 12. Cibicides lobatulus (Walker and Jakobs, 1798) Тиквешки басейн, разрез Хаджи Юсуфли, горна флишка задруга, пр. 7: 11 – спирална страна, SEMx180; 12 – умбиликална страна, SEMx160 13–15. Cibicides ungerianus (d’Orbigny, 1846) 13, 14 – Делчевски басейн, разрез Црна скала, горна флишка задруга, пр. 24: 13 – спирална страна, SEMx200; 14 – умбиликална страна, SEMx250; 15 – Овчеполски басейн, разрез Неманци, горна флишка задруга, пр. 3, умбиликална страна, SEMx160 16, 17. Cibicides sp. Тиквешки басейн, разрез Криволак, горна флишка задруга, пр. 6: 16 – умбиликална страна, SEMx160; 17 – спирална страна, SEMx180 71 Genus Valvulineria Cuchman, 1926 Valvulineria jacksonensis Cushman, 1933 Plate II, Fig. 7 1933. Valvulineria jacksonensis Cushman, sp. n.; Cushman, p. 18, pl. 2, fig. 9. 1935. Valvulineria jacksonensis Cushman; Cushman, p. 44, pl. 18, fig. 2. 1957. Valvulineria aff. jacksonensis Cushman; Sacal, Debourle, p. 35, pl. 12, fig. 2. 1971. Valvulineria cf. jacksonensis Cushman; Ferrer, p. 51, lam. 6, figs. 16–17. 1984a. Valvulineria aff. jacksonensis Cushman; Olszewska, p. 25, tabl. 4, fig. 4. Nomenclature. The holotype (U.S.N.M. Coll. No. 371559) is from the Upper Jackson Eocene of the Southeastern Coastal Plain Region of the United States (Ocala Limestone, Brooklyn, Ala). Material. 9 specimens with different degree of preser­ vation. Description. The test is low-trochospiral, biconvex. The last whorl is composed of 8 chambers, gradually increasing in size. The sutures on the umbilical side are straight, radial slightly depressed, while on the dorsal one they are curved and flush. The surface is smooth. The aperture is umbilical covered with narrow lobe of the last-formed chamber. Distribution. The species is known from the Upper Paleocene–Middle Eocene of Spain, the Eocene of the USA, the Upper Eocene of France, The Eocene and Oligocene of the Carpathians. Occurrence. Delchevo basin (Crna Skala section, sam­ ples 20, 24–26). Family EPONIDIDAE Hofker, 1951 Subfamily EPONIDINAE Hofker, 1951 Genus Eponides de Montfort, 1808 Eponides minima Cushman, 1933 Material. 10 specimens, some of them badly preserved. Remarks. The species was described from Chardaklija section of the Ovche Pole basin (Džuranov et al., 1999, p. 63). Occurrence. Ovche Pole basin (Madzarica section, samples 12–15), Valandovo–Gevgelija basin (Rabrovo section, samples 2, 6; Dedeli section, sample 4). Eponides sp. Plate II, Fig. 8 Material. Аbout 15 well preserved specimens. Description. The test is low-trochospiral, biconvex. The spiral side comprises two whorls, as the last one is composed of 6–7 indistinct chambers separated by slightly curved backwards, oblique, almost flush sutures. The umbilicus is narrow, flush. The periphery is acute, the surface is smooth. The aperture is archshaped, interiomarginal. 72 Occurrence. Ovche Pole basin (Nemanjici section, sam­ples 4–6, 10–12, 17, 18). Superfamily DISCORBINELLACEA Sigal, 1952 Family PARRELBIDIDAE Hofker, 1956 Genus Cibicidoides Thalman, 1939 Cibicidoides sp. Plate II, Figs. 9, 10 Material. About 29 specimens with different degree of preservation. Description. The test is low trochospiral, biconvex, lenticular. The umbilical side is more elevated. The spiral side reveals 2–2.5 whorls. The last whorl comprises 10–11 chambers gradually increasing in size. The periphery is angled. The spiral side is coarsely perforated. The aperture is low arch, interimarginal. Occurrence. Ovche Pole basin (Nemanjici section, samples 3, 4, 8, 9, 15–18), Tikvesh basin (Krivolak section, samples 1, 3). Superfamily PLANORBULINACEA Schwager, 1877 Family CIBICIDIDAE Cushman, 1927 Subfamily CIBICIDINAE Cushman, 1927 Genus Cibicides de Montfort, 1808 Cibicides carinatus (Terquem, 1882) Material. 40 specimens, most of them well preserved. Remarks. The species was described from Chardaklija section of the Ovche Pole basin (Džuranov et al., 1999, p. 64, pl. 1, Fig. 14). Occurrence. Ovche Pole basin (Ezevo Brdo section, samples 2–6, 9–11; Kadrifakovo section, samples 3–5, 7, 9; Madzarica section, samples 1, 2, 7, 8, 11, 12), Strumica basin (Stuka section, samples 2–5). Cibicides lobatulus (Walker and Jakobs, 1798) Plate II, Figs. 11, 12 1935. Cibicides lobatulus (Walker and Jacobs); Cushman, p. 52, pl. 22, figs. 4–6. 1954. Cibicides (Cibicides) lobatulus (Walker and Jacobs) emend. d’Orb.; Vassilenko, p. 138, pl. 22, figs. 1–5, pl. 23, fig. 1 (in Russian). 1961. Cibicides lobatulus (Walker and Jacobs); Kaasschieter, p. 221, pl. 14, fig. 5. 1973. Cibicides lobatulus (Walker and Jacobs); Ferrer et al., p. 56, fig. 17(4). 1975. Cibicides lobatulus (Walker and Jacobs); Samuel, p. 148, pl. 86, fig. 5. 1979. Cibicides lobatulus (Walker and Jacobs); Sztrakos, p. 86, pl. 26, fig. 7. 1981. Cibicides lobatulus (Walker and Jacobs); Hughes, p. 202, pl. 15.4, figs. 13, 15–16. 1984. Cibicides lobatulus (Walker and Jacobs); OdrzywolskaBienkowa, Pozaryska, p. 140, pl. 12, fig. 8, pl. 13, fig. 2. 1984b. Cibicides lobatulus (Walker and Jacobs); Olszewska, p. 62, tabl. 3, fig. 10. 1991. Cibicides lobatulus (Walker and Jacobs); Barbin, KellerGrünig, pl. 1, figs. 17–18. Nomenclature. The species was first described from recent sediments of England. Here it is identified after the SEM images of Barbin, Keller-Grünig (1991, pl. 1, Figs. 17–18). Material. About 20 specimens, most of them well preserved. Description. The test is planoconvex, ovate to round in outline, with convex umbilical side and slightly concave spiral one. The last whorl is composed of 7–8 gradually increasing in size chambers, separated by slightly depressed sutures. The last ones are curved backwards on the spiral side and almost straight on the umbilical one. The periphery is acute to keeled. The aperture is slit-like, at the base on the last-formed chamber. Remarks. The species differs from C. carinatus (Terquem) by the more convex umbilical side and the slightly concave spiral one. Kaasschieter (1961) considered intergradation between the two species. Distribution. It is known from the Paleocene and Eocene of Spain, France, Italy, Pakistan, Ukraine, the Eocene of Belgium, the USA, Caucasus and Middle Asia, the Eocene–Miocene of the Carpathians, the Oligocene of Germany, the Paris Basin, the North Sea, the Oligocene–Miocene of the Netherlands, Hungary, Florida, Sahalin Peninsula, widespread in recent sediments. Occurrence. Ovche Pole basin (Nemanjici section, samples 1, 4, 6, 8–12, 14–17; Madzarica section, samples 8–10), Tikvesh basin (Hadzi Jusufli section, samples 3–4, 6–7; Krivolak section, samples 2, 4, 7). Cibicides tallahatensis Bandy, 1949 Material. About 50 specimens with different degree of preservation. Remarks. The species was described from Chardaklija section of the Ovche Pole basin (Džuranov et al., 1999, p. 64, pl. 1, Fig. 15). Occurrence. Skopje-Kumanovo basin (Vojnik section, samples 5, 6, 8), Ovche Pole basin (Ezevo Brdo secton, sam 7, 8, 12, 13; Kadrifakovo section, samples 5–7, 10; Madzarica section, samples 8–10), Tikvesh basin (Hadzi Jusufli section, samples 1, 4, 5, 7; Krivolak section, samples 2, 4, 6). Cibicides ungerianus (d’Orbigny, 1846) Plate II, Figs. 13–15 1846. Rotalina Ungeriana d’Orbigny; d’Orbigny, p. 157, pl. 8, figs. 16–18. 1954. Cibicidoides ungerianus (d’Orbigny); Vassilenko, p. 172, pl. 29, figs. 6, 7, pl. 30, fig. 1 (in Russian). 1961. Cibicides ungerianus (d’Orbigny); Kaasschieter, p. 220, pl. 14, fig. 3. 1975. Planulina ungeriana (d’Orbigny); Samuel, p. 148, pl. 87, figs. 1–3. 1976. Cibicides ungerianus (d’Orbigny); Salaj et al., p. 158, pl. 16, figs. 3–4. 1978. Planulina (?) ungeriana (d’Orbigny); Kantorova, S. 208, t. 46, figs. 1–2. 1978. Heterolepa ungeriana (d’Orbigny); Proto Decima, Bolli, p. 794, pl. 6, figs. 18–19. 1978. Cibicides ungerianus (d’Orbigny); Odrzywolska-Bien­ kowa, et al., p. 275, pl. 12, fig. 7. 1979. Cibicidoides ungerianus (d’Orbigny); Sztrakos, p. 88, pl. 30, fig. 3. 1981. Cibicides ungerianus (d’Orbigny); Hughes, p. 198, pl. 15.2, figs. 1–3. 1983. Cibicidoides ungerianus (d’Orbigny); Tjalsma, Lohmann, p. 28, pl. 18, fig. 1, pl. 21, figs. 5–6. 1984. Cibicides ungerianus (d’Orbigny); Odrzywolska-Bien­ kowa, Pozaryska, p. 142, pl. 12, fig. 11. 1984a. Cibicides ungerianus (d’Orbigny); Olszewska, p. 28, tabl. 5, fig. 4. 1985. Heterolepa ungeriana (d’Orbigny); Grünig, p. 276, pl. 11, figs. 10–12. 1985. Cibicides ungerianus (d’Orbigny); Papp, Schmidt, p. 60, pl. 51, figs. 7–11. 2001. Cibicidoides ungerianus (d’Orbigny); Scherbacher et al., p. 626, pl. 2, figs. 5–6. Nomenclature. The species was first described from the Badenian of the Vienna Basin. The lectotype (GBA 1981/03/236) was figured by Papp, Schmidt (1985, pl. 51, Figs. 8, 9). Material. More than 20 specimens, most of them well preserved. Description. The test is planoconvex, low-trochospiral. The last worl comprises 10–13 chambers. The sutures are radial; on the umbilical side they are straight, while on the spiral one curved backwards. The periphery is acute or with very narrow keel. The aperture is slit-like, at the base on the last-formed chamber. Remarks. The species have been referred to different genera, but the planoconvex test and the presence of a narrow keel is characteristic feature for the genus Cibicides de Montfort. Distribution. The species is known from the Paleocene of Tunisia, the Paleocene and Eocene of Italy, Spain, France, Caucasus and Middle Asia, the Paleocene– Oligocene of Malij Balhan, the Paleocene-Miocene of Ukraine, Sahalin Peninsula, the Eocene of Belgium, the Eocene–Oligocene of the Netherlands, the Atlantic, the bay of Biscay, the Eocene-Miocene of France, the Carpathians, the Oligocene of Germany, the Former Republic of Jugoslavia, the North Sea, the OligoceneMiocene of Hungary, Spain, Austria. Occurrence. Ovche Pole basin (Nemanjici section, samples 1–3, 5–7), Delchevo basin (Crna Skala section, samples 10, 13, 14, 19, 23, 26). Cibicides cf. westi Howe, 1939 Material. 30 specimens with different degree of preser­ vation. Remarks. The species was described from Chardaklija section of the Ovche Pole basin (Džuranov et al., 1999, p. 65, pl. 1, Fig. 16). 73 Occurrence. Ovche Pole basin (Nemanjici section, samples 1–3, 8–12, 15–17; Ezevo Brdo section, sam­ ples 7, 8, 12; Madzarica section, samples 9–11), Delchevo basin (Crna Skala Section, samples 9, 10, 13, 15, 16, 19, 23), Strumica basin (Stuka section, sam­ples 4–5). Cibicides sp. Plate II, Figures 16, 17 Material. 10 specimens, with different degree of preser­ vation. Description. The test is low-trochospiral, with convex umbilical side and slightly concave spital one. The last worl comprises 6–7 chambers very slowly increasing in size. They are separated by straight, radial, slightly depressed sutures. The periphery is acute or bears narrow keel. The aperture is indistinct. Occurrence. Tikvesh basin (Hadzi Jusufli section, sample 4; Krivolak section, samples 5, 6). Superfamily NONIONACEA Schultze, 1854 Family NONIONIDAE Schultze, 1854 Subfamily NONIONINAE Schultze, 1854 Genus Nonion de Montfort, 1808 Nonion graniferum (Terquem, 1882) Material. 20 specimens, most of them well preserved. Remarks. The species was described from Chardaklija section of the Ovche Pole basin (Džuranov et al., 1999, p. 65, pl. 1, Fig. 19). Occurrence. Ovche Pole basin (Kadrifakovo section, samples 5–7, 10; Madzarica section, samples 11–13, 16). Genus Nonionella Cushman, 1926 Nonionella winniana Howe, 1939 Material. More than 30 well preserved specimens. Remarks. The species was described from Chardaklija section of the Ovche Pole basin (Džuranov et al., 1999, p. 65, pl. 1, Figs. 17, 18). Occurrence. Skopje-Kumanovo basin (Vojnik section, samples 3–4, 6), Ovche Pole basin (Ezevo Brdo section, samples 2, 3, 5–7, 10, 11; Kadrifakovo section, samples 3–4, 7–8). Subfamily PULLENIINAE Schwager, 1877 Genus Mellonis de Montfort, 1808 Mellonis affine (Reuss, 1851) Plate III, Fig. 1 1851. N. (Nonionina) affinis m.; Reuss, S. 72, Taf. 5, Fig. 32. 1939. Nonion affine (Reuss); Cushman, p. 9, pl. 2, fig. 13. 1961. Nonion affine (Reuss); Kaasschieter, p. 203, pl. 11, figs. 3–4. 1962. Nonion affine (Reuss); Kiesel, S. 65, Taf. 9, Fig. 14. 1970. Nonion affine (Reuss); Le Calvez, p. 190, pl. 27, fig. 2. 1975. Nonion affine (Reuss); Samuel, p. 150, pl. 79, fig. 2. 1977. Melonis affine (Reuss); Matl, Smigielska, p. 25, pl. 2–2, fig. 4. 74 1978. Melonis affine (Reuss); Odrzywolska-Bienkowa et al., p. 278, pl. 14, figs. 1–2. 1984. Melonis affine (Reuss); Odrzywolska-Bienkowa, Poza­ ryska, p. 145, pl. 14, figs. 7–8. Nomenclature. The holotype is the specimen figured by Reuss (1851, Taf. 5, Fig. 32). The species was first described from the Oligocene (Septarien clays) of Germany (Hermsdorf surroundings). Material. 30 samples, most of them are well preserved. Description. The test is planispiral, involute, oval in outline. The last whorl comprises 11–12 chambers gradually increasing in size. The umbilicus is narrow, slightly depressed. The sutures are flush. The periphery is broadly rounded. The wall is moderately perforated. Distribution. The species is known from the Eocene of Belgium, the Eocene–Oligocene of the Carpathians, Germany, France, the Eocene–Miocene of the Nether­ lands, the Oligocene of the USA. Occurrence. Ovche Pole basin (Nemanjici section, samples 1–2, 5, 7–11, 15), Valandovo-Gevgelija basin (Rabrovo section, samples 7, 13, 15; Dedeli section, samples 6–7). Genus Pullenia Parker and Jones, 1862 Pullenia quinqueloba (Reuss, 1851) Plate III, Fig. 2 1851. N. (Nonionina) quinqueloba m.; Reuss, S. 71, Taf. 5, Fig. 31. 1942. Pullenia quinqueloba (Reuss); Dam ten, Reinhold, p. 94, t. 7, fig. 7. 1957. Pullenia quinqueloba (Reuss); Sacal, Debourle, p. 52, pl. 22, fig. 14. 1959. Pullenia quinqueloba (Reuss); Mallory, p. 246, pl. 34, figs. 1a–b. 1976. Pullenia quinqueloba (Reuss); Salaj et al., p. 160, pl. 11, fig. 3. 2008a. Pullenia quinqueloba (Reuss); Valchev, p. 116, pl. 1, fig. 7 (with synonymy). Nomenclature. The holotype is the specimen figured by Reuss (1851, Taf. 5, Fig. 31). The species was first described from the Oligocene (Septarien clays) of Germany (Hermsdorf surroundings). Material. About 30 specimens, most of them well preserved. Description. The test is planispiral, involute, elliptical in outline. The last whorl is composed of 5 chambers gradually increasing in size. The periphery is narrowly rounded. The aperture is arch-shaped, at the base of the last formed chamber. Remarks. The species differs from P. jarvisi Cushman by the presence of 5 chambers in the last whorl. Distribution. It is known from the Senonian of Saratov and Volgograd Districts, Turkmenia, France, the Paleocene of Denmark, North Caucasus, Turkmenia, Tunisia, the Netherlands, the Upper Paleocene of England, the Eocene of Bulgaria, Ukraine, Donets basin, the Carpathians, Belgium, the Netherlands, England, Turkmenia, the Oligocene of Germany, Belgium, the Netherlands, Trinidad, Turkmenia, the Upper Miocene of Dominican Republic, the Miocene of Vienna basin, Bavaria, the Miocene and Pliocene of the Netherlands. It was also established during the deep-sea drilling in the Atlantic (Eocene–Oligocene, Upper Miocene, Pleistocene), Bay of Biscay (Upper Eocene–Oligocene), Norwegian Sea (Lower Eocene). Nowadays it lives in calm waters and great depths. Occurrence. Ovche Pole basin (Nemanjici section, samples 1–3, 9–12, 14–16), Valandovo–Gevgelija basin (Rabrovo section, samples 6, 7, 9, 14; Dedeli section, sample 2), Delchevo basin (Crna Skala section, samples 16, 22, 26). Superfamily CHILLOSTOMELLACEA Brady, 1881 Family CHILOSTOMELLIDAE Brady, 1881 Subfamily CHILOSTOMELLINAE Brady, 1881 Genus Chilostomelloides Cushman, 1926 Chilostomelloides balkhanensis (Dain et Chalilov, 1952) Plate III, Fig. 3 1952. Chilostomella balkhanensis Dain et Chalilov, sp. n.; Dain, p. 125, pl. 2, figs. 3–4 (in Russian). 1967. Chilostomella balkhanensis Dain et Chalilov; Chalilov, p. 110, pl. 22, fig. 2 (in Russian). Nomenclature. The holotype (VNIGRI Coll. No. 4282) is from the Upper Eocene–Lower Oligocene (Bolivina Zone) of Western Turkmenia (Malij Balkhan). Material. 12 badly preserved specimens. Description. The test is elongate, fusiform, with strongly embracing chambers. Only the final two of them are visible from the exterior. The sutures are oblique, slightly depressed. The aperture is circular, situated at the base of the last formed chamber. Remarks. The circular aperture, as well as the form of the test made us to refer this species to genus Chilostomelloides Cushman. Distribution. The species is known from the Upper Eocene of Azerbaijan, the Upper Eocene–Lower Oligocene of Turkmenia. Occurrence. Ovche Pole basin (Nemanjici section, samples 1–4). Family HETEROLEPIDAE Gonzales–Donoso, 1969 Genus Anomalinoides Brotzen, 1942 Anomalinoides acutus (Plummer, 1926) Plate III, Fig. 4 1948. Anomalinoides acuta (Plummer); Brotzen, p. 87, pl. 14, fig. 2. 1960. Anomalinoides acuta (Plummer); Olsson, p. 51, pl. 11, figs. 4–5. 1961. Anomalina acuta Plummer; Kaasschieter, p. 216, pl. 12, figs. 12-13; pl.13, fig. 4. 1983. Anomalinoides acuta (Plummer); Hanzlikova, p. 60, pl. 10, fig. 12. 2008b. Anomalinoides acutus (Plummer); Valchev, p. 123, pl. 1, figs. 1, 2 (with synonymy). Nomenclature. The species was first described from the Paleocene of Texas. Here it is identified after the SEM images of Valchev (2008b, pl. 1, Figs. 1, 2). Material. 7 well preserved specimens. Description. The test is low trochospiral, compact, slightly convex on both sides. The spiral side comprises 2.5 whorls. The last whorl is composed of 10–11 inflated chambers separated by slightly depressed, oblique, and curved backwards sutures. The wall is coarsely perforate. The aperture is low interiomarginal arch. Distribution. It is known from the Upper Cretaceous and the Paleocene of Texas, the Upper Cretaceous, Paleocene and Eocene of Bulgaria, the Paleocene of Trinidad, Arkansas, the Netherlands, Sweden, Tunisia, Caucasus, Middle Asia, the Lower Eocene of England, the Eocene of Ukraine, Byelorussia, Caucasus, Ural, North Turkmenia, Germany, Belgium. It was also found during the deep-sea drilling in Norwegian Sea (Lower Eocene) and the Atlantic (Eocene). Occurrence. Delchevo basin (Crna Skala Section, samples 21, 25). Anomalinoides danicus (Brotzen, 1940) Plate III, Fig. 5 1968. Anomalina danica (Brotzen); Pozaryska, Szczechura, p. 86, pl. 14, figs. 6–11. 1970. Anomalina ex gr. danica (Brotzen); Shutskaya, pl. 4, fig. 10, pl. 12, fig. 10, pl. 37, fig. 3 (in Russian). 1975. Anomalina cf. danica (Brotzen); Braga et al., p. 97, t. 2, figs. 12a–c. 1975. Gavelinella danica (Brotzen); Berggren, Aubert, p. 155, pl. 6, figs. 3, pl. 11, fig. 1, pl. 13, fig.1, pl. 14, fig. 4, pl. 17, fig. 7; pl. 19, fig. 4. 1976. Anomalina danica (Brotzen); Salaj et al., p. 162, pl. 9, figs. 4–5. 1983. Gavelinella danica (Brotzen); King, p. 35, pl. 5, figs. 16–17. 2008b. Anomalinoides danicus (Brotzen); Valchev, p. 124, pl. 1, figs. 3, 4 (with synonymy). Nomenclature. The species was first described from the Maastrichtian of the Netherlands. Here it is identified after the SEM images of Valchev (2008b, pl. 1, Figs. 3, 4). Material. 15 specimens with different degree of pre­ servation. Description. The test is low trochospiral, compact. The spiral side comprises 2–2.5 worls. The last whorl is composed of 8–9 chambers gradually increasing in size. The sutures are radial, slightly curved backwards, depressed. The wall is coarsely perforated. The periphery is broadly rounded. The aperture is arch-shaped, interiomarginal. Distribution. It is known from the Upper Cretaceous of Germany, the Maastrichtian of Tunisia, Ukraine, the 75 PLATE III 76 ТАБЛИЦА III PLATE III 1. Mellonis affine (Reuss, 1851) Valandovo–Gevgelija basin, Rabrovo section, upper flysh unit, sample 15, spiral view; SEMx230 2. Pullenia quinqueloba (Reuss, 1851) Delchevo basin, Crna Skala section, upper flysh unit, sample 16, spiral view; SEMx140 3. Chilostomelloides balkhanensis (Dain et Chalilov, 1952) Ovche Pole basin, Nemanjici section, upper flysh unit, sample 3; SEMx250 4. Anomalinoides acutus (Plummer, 1926) Delchevo basin, Crna Skala section, upper flysh unit, sample 24, umbilical view; SEMx180 5. Anomalinoides danicus (Brotzen, 1940) Ovche Pole basin, Nemanjici section, upper flysh unit, sample 13, umbilical view; SEMx280 6, 7. Anomalinoides welleri (Plummer, 1926) Tikvesh basin, Krivolak section, upper flysh unit, sample 3: 6, umbilical view, SEMx150; 7, spiral view, SEMx190 8–11. Heterolepa dutemplei (d’Orbigny, 1846) 8, 9, Delchevo basin, Crna Skala section, upper flysh unit, sample 5: 8, spiral view, SEMx120; 9, umbilical view, SEMx130; 10, Ovche Pole basin Nemanjici section, upper flysh unit, sample 7, umbilical view, SEMx120; 11, Valandovo–Gevgelija basin, Dedeli section, upper flysh unit, sample 11, spiral view, SEMx150 12, 13. Heterolepa perlucida (Nautall,1932) Valandovo–Gevgelija basin, Rabrovo section, upper flysh unit, sample 15: 12, umbilical view, SEMx150; 13, spiral view, SEMx180 14. Gyroidinoides soldanii (d’Orbigny, 1826) Ovche Pole basin, Madzarca section, upper flysh unit, sample 12, umbilical view, SEMx220 Scale bar – 100 µm ТАБЛИЦА III 1. Mellonis affine (Reuss, 1851) Валандово–Гевгелийски басейн, разрез Раброво, горна флишка задруга, пр. 15, спирална страна; SEMx230 2. Pullenia quinqueloba (Reuss, 1851) Делчевски басейн, разрез Црна скала, горна флишка задруга, пр. 16, спирална страна; SEMx140 3. Chilostomelloides balkhanensis (Dain et Chalilov, 1952) Овчеполски басейн, разрез Неманци, горна флишка задруга, пр. 3; SEMx250 4. Anomalinoides acutus (Plummer, 1926) Делчевски басейн, разрез Црна скала, горна флишка задруга, пр. 24, умбиликална страна; SEMx180 5. Anomalinoides danicus (Brotzen, 1940) Овчеполски басейн, разрез Неманци, горна флишка задруга, пр. 13, умбиликална страна; SEMx280 6, 7. Anomalinoides welleri (Plummer, 1926) Тиквешки басейн, разрез Криволак, горна флишка задруга, пр. 3: 6 – умбиликална страна, SEMx150; 7 – спирална страна, SEMx190 8–11. Heterolepa dutemplei (d’Orbigny, 1846) 8, 9 – Делчевски басейн, разрез Црна скала, горна флишка задруга, пр. 5: 8 – спирална страна, SEMx120; 9 – умбиликална страна, SEMx130; 10 – Овчеполски басейн, разрез Неманци, горна флишка задруга, пр. 7, умбиликална страна, SEMx120; 11 – Валандово–Гевгелийски басейн, разрез Дедели, горна флишка задруга, пр. 11, спирална страна, SEMx150 12, 13. Heterolepa perlucida (Nautall, 1932) Валандово–Гевгелийски басейн, разрез Раброво, горна флишка задруга, пр. 15: 12 – умбиликална страна, SEMx150; 13 – спирална страна, SEMx180 14. Gyroidinoides soldanii (d’Orbigny, 1826) Овчеполски басейн, разрез Маджарица, горна флишка задруга, пр. 12; умбиликална страна, SEMx220 Netherlands, the Paleocene of North Europe, Poland, Ukraine, North Caucasus, Crimea, Tunisia, Bulgaria, the Eocene of Spain, France, Italy, Caucasus, Middle Asia, the Carpathians, deep-sea holes in the Atlantic (Paleocene–Lower Eocene), and Norwegian Sea (Lower Eocene). Occurrence. Ovche Pole basin (Nemanjici section, samples 6–8, 13–15). 77 Anomalinoides welleri (Plummer, 1926) Plate III, Figs. 6, 7 1951. Anomalina welleri (Plummer); Cushman, p. 63, l. 18, fig. 12. 1954. Anomalina (Anomalina) welleri (Plummer); Vassilenko, p. 62, pl. 3, figs. 6, 7. 1975. Anomalinoides welleri (Plummer); Berggren, Aubert, p. 151, pl. 5, fig. 3, pl. 13, fig. 7, pl. 18, fig. 6, pl. 19, fig. 1. 1976. Anomalinoides welleri (Plummer); Aubert, Berggren, p. 430, pl. 9, fig. 5. 1992. Anomalinoides welleri (Plummer); Kaiho, p. 255, pl. 4, fig. 3. 2008b. Anomalinoides welleri (Plummer); Valchev, p. 124, pl. 1, fig. 7. Nomenclature. The species was first described from the Paleocene of Texas. Here it is identified after the SEM images of Kaiho (1992). Material. 12 specimens with different degree of pre­ servation. Description. Test is trochospiral, biconvex, moderately flattened. Spiral side is evolute and it reveals 2–2.5 whorls, as the chambers are distinct in the last one only. 10–12 triangular chambers with gradually increasing sizes are visible on the umbilical side. Sutures are radial, slightly depressed, curved backwards. Umbilicus is narrow, deep, usually filled with secondary deposits. Periphery is broadly rounded. Wall is finely perforated. Aperture is slit-like, extended from periphery to umbilicus. Distribution. The species is known from the Maastrichtian and Paleocene of the Tethys region. It was also established in the deep-sea holes in the North Atlantic (Paleocene). Occurrence. Ovche Pole basin (Nemanjici section, samples 1, 2, 16–18), Tikvesh basin (Krivolak section, samples 3, 5). Genus Heterolepa Franzenau, 1894 Heterolepa dutemplei (d’Orbigny, 1846) Plate III, Figs. 8–11 1846. Rotalina Dutemplei d’Orbigny; d’Orbigny, p. 157, pl. 8, figs. 19–21. 1942. Cibicides dutemplei (d’Orbigny); Dam ten, Reinhold, p. 99, t. 8, fig. 3. 1954. Cibicides (Gemellides) dutemplei (d’Orbigny); Vassi­ lenko, p. 195, pl. 35, figs. 1, 3 (in Russian). 1957. Cibicides dutemplei (d’Orbigny); Sacal, Debourle, p. 68, pl. 32, fig. 7. 1961. Cibicides dutemplei (d’Orbigny); Kaasschieter, p. 218, pl. 12, fig. 15. 1962. Cibicides cf. dutemplei (d’Orbigny); Kiesel, S. 73, taf. 11, fig. 1. 1970. Heterolepa dutemplei (d’Orbigny); Le Calvez, p. 202. 1975. Heterolepa dutemplei (d’Orbigny); Braga et al., p. 109, t. 6, fig. 1–3. 1978. Heterolepa dutemplei (d’Orbigny); Kantorova, p. 206, t. 45, figs. 1–6. 78 1979. Heterolepa dutemplei (d’Orbigny); Sztrakos, p. 88, pl. 31, fig. 1. 1980. Heterolepa dutemplei (d’Orbigny); Jutson, p. 381, pl. 2, fig. 14. 1985. Heterolepa dutemplei (d’Orbigny); Grünig, p. 275, pl. 11, figs. 4–6. 1985. Heterolepa dutemplei (d’Orbigny); Papp, Schmidt, p. 61, pl. 52, figs. 1–6. Nomenclature. The species was first described from the Badenian of the Vienna Basin. The lectotype (GBA Coll. No. 1981/03/240) was designated by Papp, Schmidt (1985, pl. 14, Fig. 2). Material. About 25 specimens, most of them well preserved. Description. The test is low trochospiral, biconvex, as the umbilical side is more elevated. The spiral one is compose of 2–2.5 worls. The last worl comprises 10– 12 chambers separated by radial, curved backwards, slightly depressed (on the umbilical side) or flush (on the spiral side) sutures. The wall is moderately perforated. The periphery is subangular. The aperture is slit-shaped, interiomarginal. Distribution. The species is known from the Paleocene of Ukraine, the Carpathians, the Eocene of Belgium, France, Spain, Italy, the Oligocene of Germany, Turkmenia, the Former republic of Yugoslavia, the Eocene–Miocene of the Carpathians, Sahalin Peninsula, the Oligocene–Miocene of Spain, Hungary, the Netherlands, Occurrence. Ovche Pole basin (Nemanjici section, samples 3, 4, 6–8, 11–13, 17–18), ValandovoGevgelija basin (Rabrovo section, samples 3, 7–8; Dedeli section, samples 1, 2, 6, 7, 10–12), Delchevo basin (Crna Skala Section, samples 4, 9, 16, 18). Heterolepa perlucida (Nuttall, 1932) Plate III, Figs. 12, 13 1954. Cibicides (Gemellides) perlucidus Nuttall; Vassilenko, p. 191, pl. 34, figs. 2, 4. 1957. Cibicides perlucidus Nuttall; Sacal, Debourle, p. 69, pl. 31, fig. 8. 1971. Heterolepa perlucida (Nautall); Ferrer, p. 61, lam. 7, figs. 13–14. 1978. Cibicides perlucidus Nuttall; Shutskaya et al., pl. 3, fig.7 (in Russian). 1979. Cibicidoides perlucidus (Nuttall); Schnitker, pl. 11, figs. 10–12. 1984. Heterolepa perlucida (Nautall); Odrzywolska-Bienkowa, Pozaryska, p. 144, pl.14, fig. 4. 1984a. Heterolepa perlucida (Nautall); Olszewska, p. 31, tabl. 7, fig. 8. 2008b. Heterolepa perlucida (Nuttal); Valchev, p. 125, pl. 1, figs. 11, 12 (with synonymy). Nomenclature. The species was first described from the Lower Oligocene of Mexico. Here it is identified after the SEM images of Valchev (2008b, pl. 1, Figs. 11, 12). Material. 5 specimens with different degree of preser­ vation. Description. The test is low trochospiral, biconvex. The spiral one is compose of 2–2.5 worls. The last worl comprises 9–10 chambers separated by radial, curved backwards, slightly depressed (on the umbilical side) or flush (on the spiral side) sutures. The wall is moderately perforated. The periphery is subangular. The aperture is slit-shaped, interiomarginal. Remarks. The species differs from H. grimsdalei (Nuttall) by its smaller pore’s diameter on the spiral side, and from H. dutemplei (d’Orbigny) by the less convex test. Distribution. It is known from the Paleocene and Eocene of Bulgaria, Spain, Turkmenia, France, North Caucasus, Crimea, the Carpathians, the Upper Eocene of Poland, the Oligocene of Mexico and the Carpa­ thians, the Oligocene and Miocene of France. Occurrence. Valandovo-Gevgelija basin (Rabrovo section, samples 8, 13, 15). Family GAVELINELLIDAE Hofker, 1956 Subfamily GYROIDINOIDINAE Saidova, 1981 Genus Gyroidinoides Brotzen, 1942 Gyroidinoides soldanii (d’Orbigny, 1826) Plate III, Fig. 14 1846. Rotalina Soldanii d’Orbigny; d’Orbigny, p. 155, pl. 8, figs. 10–12. 1947. Gyroidina soldanii d’Orbigny; Subbotina, p. 99, pl. 3, figs. 20–22 (in Russian). 1953. Gyroidina soldanii d’Orbigny; Myatlyuk, p. 61, pl. 5, figs. 3–5 (in Russian). 1957. Gyroidina soldanii d’Orbigny; Sacal, Debourle, p. 36, pl. 13, figs. 7–8. 1962. Gyroidina soldanii d’Orbigny; Kiesel, S. 69, taf. 10, fig. 6. 1975. Gyroidina soldanii d’Orbigny; Samuel, p. 151, pl. 81, figs. 3–4. 1975. Gyroidinoides soldanii (d’Orbigny); Braga et al., p. 97, t. 2, figs. 19a–c; p. 109, t. 6, figs. 10–11. 1977. Gyroidina soldanii d’Orbigny; Pozaryska, p. 43, pl. 4, figs. 4a-c, pl. 13, figs. 12–13. 1978. Gyroidina soldanii d’Orbigny; Odrzywolska–Bienkowa et al., p. 277, pl. 13, fig. 6. 1978. Gyroidinoides soldanii (d’Orbigny); Proto Decima, Bolli, p. 794, pl. 5, figs. 1–2. 1979. Gyroidinoides soldanii (d’Orbigny); Sztrakos, p. 87, pl. 29, fig. 6. 1980. Gyroidina soldanii d’Orbigny; Jutson, p. 381, pl. 2, fig. 12. 1984a. Gyroidina soldanii d’Orbigny; Olszewska, p. 30, tabl. 6, fig. 13. 1984. Gyroidina soldanii d’Orbigny; Olszewska, Szymakowska, p. 134, tabl. 2, fig. 13. 1985. Gyroidinoides soldanii (d’Orbigny); Grünig, p. 275, pl. 10, figs. 12–14. 1985. Gyroidina soldanii d’Orbigny; Papp, Schmidt, p. 60, pl. 50, figs. 4–9. 1987. Gyroidina soldanii d’Orbigny; Ponomareva, p. 64, pl. 2, fig. 2 (in Russian). 1988. Gyroidinoides soldanii (d’Orbigny); Parisi, Coccioni, p. 104, pl. 3, figs. 20–22; pl. 4, figs. 1–3. Nomenclature. The species was first described from the Eocene of France (Bordeaux vicinities). Here it is identified after the images of Papp, Schmidt (1985, pl. 50, Figs. 4–9). Material. About 30 specimens with different degree of preservation. Description. The test is trochospiral, with flat or slightly convex spiral side and highly convex umbilical one. The spiral side comprises 2–2.5 whorls. The last whorl is composed of 11–12 chambers gradually increasing in size. The last formed one is subconical in shape. The umbilicus is open, narrow. The sutures are radial, flush to slightly depressed on the umbilical side, and oblique, depressed on the spiral side. The periphery is narrowly rounded, the surface is smooth. The aperture is slit-like, interiomarginal. Distribution. The species is known from the Paleocene and Eocene of France, Italy, Spain, the Paleocene– Oligocene of Caucasus and Middle Asia, the Upper Eocene of the Carpathians, the Eocene–Oligocene of the Atlantic, the Eocene-Pleistocene of the Carpa­ thians, the Oligocene of Germany, the Oligocene–Neo­ gene of Hungary, Spain, the Miocene of the Vienna Basin, Paris Basin. Occurrence. Ovche Pole basin (Nemanjici section, samples 3, 4, 6–10, 16–18; Madzarica section, samples 11–13), Tikvesh basin (Hadzi Jusufli section, samples 2, 4, 5, 7; Krivolak section, samples 2, 3, 5, 6). Superfamily ROTALIACEA, Ehrenberg, 1839 Family ROTALIIDAE Ehrenberg, 1839 Subfamily PARARPTALIINAE Reiss, 1963 Genus Pararotalia Le Calvez, 1949 Pararotalia audouini (d’Orbigny, 1826) Material. 40 specimens with different degree of pre­ servation. Remarks. The species was described from Chardaklija section of the Ovche Pole basin (Džuranov et al., 1999, p. 66, pl. 1, Fig. 20). Occurrence. Skopje–Kumanovo basin (Vojnik section, samples 7, 8), Ovche Pole basin (Ezevo Brdo section, samples 4–14; Kadrifakovo section, samples 5–10), Strumica basin (Stuka section, samples 3–5). Pararotalia subinermis Bhatia, 1955 Material. 25 specimens, most of them well preserved. Remarks. The species was described from Chardaklija section of the Ovche Pole basin (Džuranov et al., 1999, p. 66, pl. 1, Fig. 21). Occurrence. Skopje–Kumanovo basin (Vojnik section, samples 4, 5, 7), Ovche Pole basin (Kadrifakovo section, samples 3–7, 10; Madzarica section, samples 5, 7–9). 79 References Aubert, J., W. Berggren. 1976. Paleocene benthic foraminiferal biostratigraphy and paleoecology of Tunisia. – Bull. Centre Rech. Pau-SNPA, 10, 2, 379–469. Barbieri, R. 1990. L’Eocene medio e superiore del Bacino di Tripolitania (Libia nord occidentale): biostratigrafia e paleoecologia. – Boll. Soc. Paleontol. Italiana, 29 (3), 253–271. Barbin, V., A. Keller-Grünig. 1991. Benthic foraminiferal assemblages from the Brendola section (Priabonian stage stratotype area, Northern Italy): Distribution, palaeoenvironment end palaeoecology. – Marine Micropaleontol., 17, 237–254. Berggren, W., J. Aubert. 1975. Paleocene benthonic foraminiferal biostratigraphy, paleobiogeography and paleoecology of Atlantic–Tethyan region: Midway-type fauna. – Palaeogeogr., Palaeoclimatol., Palaeoecol., 18, 73–192. Braga, G., R. De Biase, A. Grünig, F. Proto Decima. 1975. Foraminiferi Bentonici del Paleocene e dell’ Eocene della Sezione di Possagno. – Schweiz. Palaontol. Abh., 97, 85–111. Brotzen, F. 1948. The Swedish Paleocene and its foraminiferal fauna. – Sverig. Geol. Unders., 493, 42, 2, 1–140. Bugrova, E. M. 2004. Paleogene foraminifers from the Central Caucasus and Precaucasus. – In: Bogdanova, T. N., W. A. Gavrilova (Eds.). Atlas of the most important groups of fauna of Mesozoic-Cenozoic deposits of the Northern Caucasus and Praecaucasus. Saint Petersburg, VSEGEI Press, 82–126 (in Russian). Chalilov, D. 1948. Upper Cretaceous stratigraphy of the Malij Balkhan based on foraminifers. Baku, Acad. Sci. AzSSR, 93 p. (in Russian). Chalilov, D. 1967. Paleogene Microfauna and stratigraphy of Azerbaijan – part II. Baku, Acad. Sci. AzSSR, 286 p. (in Russian). Cimerman, F., B. Jelen, D. Skaberne. 2006. Late Eocene benthic foraminiferal fauna from clastic sequence of the Socka – Dobrna area and its chronostratigraphic importance (Slovenia). – Geologija, 49, 1, Ljubljana, 7–44. Cushman, J. 1926. The Foraminifera of the Velasco shale of the Tampico embayment. – Bull. Amer. Ass. Petrol. Geol., 10, 581–612. Cushman, J. 1933. New foraminifera from the Upper Jackson Eocene of the southeastern coastal plain region of the United States. – Contr. Cush. Lab. For. Res., 9, 1, 1–21. Cushman, J. 1935. Upper Eocene Foraminifera of the Southeastern United States. – Geol. Surv. Prof. Paper, 181, 1–60. Cushman, J. 1939. A monograph of the foraminiferal Family Nonionidae. – Geol. Surv. Prof. Paper, 191, 1–69. Cushman, J. A. 1951. Paleocene foraminifera of the gulf coastal region of the United States and adjacent areas. Geol. Surv. Prof. Paper, 232, 75 p. Cushman, J. A., A. N. Dusenbery. 1934. Eocene foraminifera of the Poway conglomerate of California. – Contr. Cush. Lab. For. Res., 10, 3, 51–64. Cushman, J., Y. Ozawa. 1930. A monograph of the foraminiferal family Polymorphinidae. Recent and fossil. – Proc. U. S. Nat. Mus., 77 (2829), 1–185. Cushman, J., F. Parker. 1947. Bulimina and related foraminiferal genera. Geol. Surv. Prof. Pap., 210-D, 176 p. Cushman, J. A., G. Ponton. 1932. An Eocene foraminiferal fauna of Wilcox age from Alabama. – Contr. Cush. Lab. For. Res., 8, 3, 51–72. Cuvillier, J., V. Szakall. 1949. Foraminiféres d’Aquitaine. Soc. Nat. des Petroles d’Aaquitaine, 113 p. Dain, L. 1952. Chilostomellids. – Treat. VNIGRI, New Ser., 63, 114–147 (in Russian). 80 Dam ten, A., Th. Reinhold. 1942. Die stratigraphische Glie­ derung des niederlandischen Oligo-Miozans nach Fora­ miniferen. – Meded. Geol. Stichting, Ser. C-V-No 2, 1–106. Darakchieva, S. 1999. Small Tertiary foraminifers from Bulgaria. Atlas. Sofia, Bulgaria, 92 p. Darakchieva, S., S. Juranov. 1992. Small foraminifers from the Middle and Upper Eocene in the Burgas district. Calcareous benthic foraminifers – Part І. – Geologica Balc., 22, 4, 3–42. Dumurdzanov, N., T. Serafimovski, B. C. Burchfiel. 2005. Cenozoic tectonics of Macedonia and its relation to the South Balkan extensional regime. – Geosphere, 1, 1, 1–22. Džuranov, S., V. Tuneva, N. Dumurdžanov. 1999. Micro­ foraminifera findings near the village of Čardaklija in the Ovče Pole Paleogene basin, Republic of Macedonia. – Geologica Macedonica, 13, 55–68. Ebensberger, H. 1962. Stratigraphische und mikropalaeontologische untersuchungen in der Aachener Oberkreide, besonders der Maastricht-Stufe. – Palaeontographica, A, 120, 1–120. Ferrer, J. 1971. El Paleoceno y Eoceno del Borde Sur-oriental de la Depresion del Ebro (Cataluna). Schweiz. Palaontol. Abh., 90, 70 p. Ferrer, J., Y. Le Calvez, H. Luterbacher, I. Premoli Silva. 1973. Contribution de l’étude des Foraminifères Ilerdiens de la région de Tremp (Catalogne). Mém. Mus. Nat. d’Hist. Natur., C, 29, 78 p. Geroch, S., J. G. Verdenius. 1983. Note to the plates of J. Grzybowski’s micropaleontological publications. – In: Verdenius, J. G., J. E. van Hinte, A. R. Fortuin (Eds.). Proceedings of the First Workshop on Arenaceous Fora­minifera. Trondheim, Norway, Sept. 7–9, 1981, 273–303. Grünig, A. 1985. Systematical description of Eocene benthic foraminifera of Possagno (Northern Italy), Sansoain (Northern Spain) and Biarritz (Aquitaine, France). – Mém. Sci. Géol., 37, 251–302. Hagn, H. 1956. Geologische und palaontologische Unter­ suchungen im Tertiar des Monte Brione und seiner Umgebung (Gardasee, Oberitalien). – Palaeontographica, 107, A, 67–210. Hanzlikova, E. 1983. Paleogene stratigraphy and foraminifera of the Outer Flysch Belt. – Misc. Micropalaeont., Hodonin, 43–70. Haque, A. 1956. The Foraminifera of the Ranikot and the Laki of the Nammal Gorge, Salt Range. Mem. Geol. Surv. Pakistan, 1, 300 p. Hillebrandt, A. von. 1962. Das Paleozan und seine Fora­ miniferenfauna im Becken von Reichenhall und Salzburg. – Bayer. Acad. Wiss., math-natw. Cl., Abh., (N. F.), 108, 1–192. Hofker, J. 1968. Tertiary Foraminifera of Coastal Ecuador– Lower Oligocene and Lower Miocene. Palaeontographica, Abt. A, 130, 59 p. Horvath, M. 2003. Data to revision and distribution of small foraminifera species described by HANTKEN (1868, 1875). Part II. Nodosariidae and Vaginulinidae. – Fragm. Palaeont. Hung., 21, 5–32. Hughes, M. 1981. Contribution on the Oligocene and Eocene microfaunas of the southern North Sea. – In: Microfossils from recent and fossil shelf seas. British Micropaleontol. Soc. Ser., 186–204. Jutson, D. 1980. Oligo-Miocene benthonic foraminifera from Barranco Blanco, Province of Almeria, SE Spain. – Rev. Esp. Micropaleontol., 12, 3, 365–381. Kaasschieter, J. 1961. Foraminifera of the Eocene of Belgium. Inst. Royal Sc. Nat. Belg., Mem. 47, 271 p. Kaiho, K. 1992. A low extinction rate of intermediate-water benthic foraminifera at the Cretaceous/Tertiary boundary. – Marine Micropaleontol., 18, 229–259. Kantorova, V. 1978. Vnutorna morfologia a stavba stien niektorych zastupkov podradu Rotaliina. – In: Paleontol. Konf.’ 77. Praha, Univ. Karlova, 197–215. Kaptarenko-Chernousova, O. K. 1971. Paleocene Nodosariids from Ukraine. Kijv, Naukovaya Dumka, 147 p. (in Ukrainian). Kiesel, Y. 1962. Die Oligozänen Foraminiferen der Tiefbohrung Dobbertin (Mecklenburg). – Freib. Forsch.-H., Bd. 1, 1–123. King, C. 1983. Cainozoic micropalaeontological biostratigraphy of the North Sea. Inst. Geol. Sci., Rep. 82/7, 40 p. Krhovsky, J. 1983. Foraminiferida of the Eocene–Oligocene boundary from the Pouzdrany Formation. – Knihovnicka Zemniho plynu a nafty, 4, 71–98. Krayeva, E. Y., B. F. Zernetskij. 1969. Paleogene Foraminifers of Ukraine. Paleont. Sb., 3, 197 p. (in Russian). Le Calvez, Y. 1970. Contribution à l’étude des Foraminifères Paléogènes du Bassin de Paris. Cah. Paléontol. Edit. Centre Nat. Rech. Sci., 324 p. Loeblich, A. Jr., H. Tappan. 1988. Foraminiferal genera and their classification. New York, Van Nostrand Reinhold C., 970 p. Maksimovič, B., B. Sikošek, O. Markovič, M. Veselinovič. 1954. Geološki sastav i tektonska struktura jednog dela Ovceg Polja i Tikveša sa paleontološkom dokumentacijom. – Trudovi na Geološki Zavod na NRM, 4, Skopje, 1–177 (in Macedonian). Mallory, V. 1959. Lower Tertiary Biostratigraphy of the California Coast Ranges. Tusla, Oklahoma, AAPG, 416 p. Mathelin, J.-C., K. Sztrаkos. 1993. L’Eocène de Biarritz (Pyrénées Atlantiques, SW France). Stratigraphie et paléoenvironnement. Monographie des foraminifères. – Cahiers de Micropaléontol., 8, 5–85. Matl, K., T. Smigielska. 1977. Palaeogene marine sediments between Glogow and Sieroszowice (Lower Silesia – Poland). – Rocz. Pol. Tow. Geol., 57, 1, 11–25. Mitrovič-Petrovič, J., T. Ljubotenski, M. Pavlovič. 1990. Paleogeni ehinidi Istočne Makedonje. – In: XII Kongres na geolozi na Jugoslavija, Ohrid, kn. 1, 369–377 (in Macedonian). Myatlyuk, E. 1953. Spirilinids, Rotaliids, Epistomionids, and Asterigerinids. Treat. VNIGRI, New Ser., 71, 273 p. (in Russian). Odrzywolska-Bienkowa, E., K. Pozaryska, E. Martini. 1978. Middle Oligocene microfossils from the Polish Lowlands: their stratigraphical and palegeographical significance. – Acta Palaeontol. Polonica, 23, 3, 249–291. Odrzywolska-Bienkowa, E., K. Pozaryska. 1984. Priabonian foraminifers of the Polish Lowlands. – Acta Palaeontol. Polonica, 29, 3–4, 107–156. Olszewska, B. 1984a. Interpretacja paleoekologiczna otwornic kredy I paleogenu Polskich Karpat zewnetrznych. – Biul. Inst. Geol., 346, 7–62. Olszewska, B. 1984b. Biostratygrafia serii menilitowo-krosnienskiej w okolicy Przemysla (Jednostka Skolska, Polskie Karpaty Zewnetrzne). – Biul. Inst. Geol., 340, 45–87. Olszewska, B. 1985. Otwornice warstw menilitowych Polskich Karpat Zewnetrznych. – Rocz. Pol. Tow. Geol., 55, 1/2, 201–250. Olszewska, B., F. Szymakowska. 1984. Olistostroma w Kolaczycach Kolo Jasla (Karpaty Srodkowe) i czas jejpowstania w swietle nowych badan mikropaleontologicznych. – Biul. Inst. Geol., 346, 117–145. Olsson, R. 1960. Foraminifera of Latest Cretaceous and earliest Tertiary age in the New Jersey coastal plain. – J. Paleontology, 34, 1, 1–58. D’Orbigny, A. 1846. Foraminifères Fossiles du Bassin Tertiaire de Vienne. Paris, Gide et Comp., 312 p. Papp, A., A. E. Schmidt. 1985. Die fossilen Foraminiferen des Tertiaren Becken von Vien. Revision der Monographie von Alcide d’Orbigny (1846). Abh. Geol. Bundesanstalt, 37, 310 p. Parisi, G., R. Coccioni. 1988. Deep-water benthic foraminifera at the Eocene–Oligocene boundary in the Massignano section (Ancona, Italy). – Int. Subcomm. Paleog. Strat., E/O Meeting, Spec. Publ., 2, 3, 97–109. Pishvanova, L. 1972. Upper Eocene and Miocene foraminifers from Western USSR. – In: Materials of paleontology and Stratigraphy of oil-bearing areas of western USSR. Treat. VNIGRI, 27, 206–283 (in Russian). Ponomareva, L. 1987. Calcareous benthic foraminifers from the Sheshor Horizon in the Ukrainian Carpathians. – Paleontol. Sb., 24, Lvov, 60–65 (in Russian). Pozaryska, K. 1965. Foraminifera and Biostratigraphy of the Danian and Montian in Poland. Palaeontol. Polonica, 14, 156 p. Pozaryska, K. 1977. Upper Eocene foraminifera of East Poland and their palaeogeographical meaning. – Acta Paleontol. Polonica, 22, 1, 3–54. Pozaryska, K., J. Szczechura. 1968. Foraminifera from the Paleocene of Poland, their ecological and biostratigraphical meaning. Palaeontol. Polonica, 20, 107 p. Proto Decima, F., H. Bolli. 1978. Southeast Atlantic DSDP leg 40 Paleogene benthic foraminifers. – Init. Repts. DSDP, 40, 783–809. Reuss, A. 1851. Über die fossilen Foraminiferen und Ento­ mostraceen der Septarienthone der umgegent von Berlin. – Zeitschr. Deutsch. Geol. Ges., 3, 49–92. Sacal, V., A. Debourle. 1957. Foraminifères d’Aquitaine – Peneroplidae a Victoriellidae. Mém. Soc. Géol. Fr., nov. sér., 36, 1, mém. 78, 88 p. Salaj, J., K. Pozaryska, J. Szczechura. 1976. Foraminiferida, zonation and subzonation of the Paleocene of Tunisia. – Acta paleontol. Polonica, 21, 2, 127–190. Samuel, O. 1975. Foraminifera of Upper Priabonian from Lubietova (Slovakia). – Zap. Karpaty, Ser. paleontol., 1, 111–176. Scherbacher, M., G. Schmiedl, Ch. Hemleben. 2001. Early Oligocene benthic Foraminifera from the Lower Inn Valley area: implications for the paleoenvironmental evolution of the Inneralpine Molasse. – In: Piller, W. E., M. W. Rasser (Eds.). Paleogene of the Eastern Alps. Osterr. Akad. Wiss., Schriftenr. Erdwiss. Komm., 14, 611–640. Schnitker, D. 1979. Cenozoic deep water benthic foraminifers, Bay of Biscay. – Init. Repts. DSDP, 48, 377–413. Shutskaya, E. 1970. Lower Paleogene stratigraphy, foraminifers and paleogeography of Crimea, Precaucasus and western part of Middle Asia. Treat. VNIGRI, 70, 256 p. (in Russian). Shutskaya, E., G. Kurgalimova, A. Moksyakova, G. Korovina, G. Chasina. 1978. Eocene stratigraphical analogies between Armenia and the Mediterranean. – Geol. Min. Deposits Asia, Africa and Central America, 3, 3–21 (in Russian). Spasov, Ch. 1951. Thortonian foraminifers from Staropatitsa Village area, Kula District. – Bull. Geol. Inst., 1, 103–124 (in Bulgarian). Stancheva, M. 1959. Microfaunal characteristics of the Torthonian from the Northwest Bulgaria. – Trav. ser la Géol. de Bulg., Ser. paleontol., 1, 229–320 (in Bulgarian). Stojanova, V. 2008. Evolution and stratigraphy of the Paleo­ gene in the Republic of Macedonia. Resume of Ph.D. Thesis. Štip, 38 p. (in Macedonian). 81 Stojanova, V., G. Petrov. 2012. Correlation of the lithostratigraphic profiles in the Paleogene basins in the Republic of Macedonia. – In: Proceedings of National conference with international participation “Geosciences 2012”. Sofia, BGS, 99–100. Stojanova, V., G. Petrov, V. Stefanova. 2011. Small foraminifers from the Paleogene basins in the Republic of Macedonia. – In: Proceedings of National conference with international participation “Geosciences 2011”. Sofia, BGS, 93–94. Stojanova, V., G. Petrov, V. Stefanova. 2012. Biostratigraphy of the Ovche Pole Paleogene basin, Republic of Macedonia. – In: Second Congr. Geol. Sci. Maced., Krushevo, Geol. Macedonica, Sp. Publ., 53–62 (in Macedonian). Stojanova, V., B. Valchev, S. Juranov. 2013. Paleogene planktonic foraminifera of the Republic of Macedonia. – C. R. Acad. bulg. Sci., 66, 5, 717–724. Sztrakos, K. 1979. La Stratigraphie, Paléoécologie, Paléogéo­ graphie et les Foraminifères de l’Oligocène du Nord-Est de la Hongrie. Cahiers de Micropaléontologie, 3, 95 p. Sztrakos, K. 1996. Le Thanétien supérior et l’Yprésien du Bassin de l’Adour (Aquitaine, France). Stratigraphie et paléoénvironnement. Etude des foraminifères. – Cahiers de Micropaléontogie, 10, 25–63. Subbotina, N. 1947. Danian and Paleogene foraminifers from Caucasus. – In: Microfauna of oil-bearing deposits of Caucasus, Emba and Middle Asia. VNIGRI, 39–160 (in Russian). Subbotina, N. 1953. Upper Eocene Lagenids and Buliminids from the South USSR. – Treat. VNIGRI, new ser. 69, Microfauna USSR, 6, 115–255 (in Russian). Subbotina, N. 1960. Oligocene and Miocene microfauna of Vorotyste River Vally (Precarpathians). – Treat. VNIGRI, 153, Microfauna USSR, 11, 157–263 (in Russian). Subbotina, N., L. Pishvanova, L. Ivanova. 1960. Oligocene and Miocene foraminiferal stratigraphy of Precarpathians. – Treat. VNIGRI, 153, Microfauna USSR, 11, 5–156 (in Russian). Terquem, M. 1882. Les foraminifères de l’Eocène des environs de Paris. – Mém. Soc. Géol. France, Ser. 3, 2, 1–193. Tjalsma R., G. Lohman. 1983. Paleocene–Eocene Bathyal and Abyssal Benthic Foraminifera from the Atlantic Ocean. Micropaleontology, spec. publ. 4, 89 p. Valchev, B. 2001. Genus Nodosaria Lamarck, 1812 from the Paleocene part of Bjala Formation (East Balkan Mountains). – Rev. Bulg. Geol. Soc., 62, 1–3, 111–115. Valchev, B. 2002. Unilocular foraminifers from the Paleocene of the coastal part of East Stara Planina. – Ann. de l’Univ. de Sofia, Fac. Géol. et Géogr., 95, 1–Géol., 17–30 (in Bulgarian, with an English abstract). Valchev, B. 2004. Genus Lenticulina Lamarck, 1804 from the Paleocene of the coastal part of East Stara Planina. – Ann. Univ. of Mining and Geol., 47, І–Geol. and Geophys., 51–56. Valchev, B. 2005. Paleocene Polymorphinids and Glandulinids (Order Foraminiferida) from the coastal part of East Stara Planina (East Bulgaria). – Ann. Univ. of Mining and Geol., 48, І–Geol. and Geophys., 161–165. Valchev, B. 2007. Midway-type benthic foraminifera from the Paleocene of the coastal part of East Stara Planina (Eastern Bulgaria). Family TEXTULARIIDAE Ehrenberg, 1838 to family STILOSTOMELLIDAE Finlay, 1947. – Ann. Univ. of Mining and Geol., 50, I–Geol. and Geophys., 129–137. Valchev, B. 2008a. Midway-type benthic foraminifera from the Paleocene of the coastal part of East Stara Planina (Eastern Bulgaria). Family BAGGINIDAE Cushman, 1927 to family ORIDORSALIDAE Loeblich and Tappan, 1964. – Ann. Univ. of Mining and Geol., 51, I–Geol. and Geophys., 119–126. Valchev, B. 2008b. Midway-type benthic foraminifera from the Paleocene of the coastal part of East Stara Planina (Eastern Bulgaria). Family HETEROLEPIDAE Gonzales-Donoso, 1969 to family ROTALIIDAE Ehrenberg, 1839. – Ann. Univ. of Mining and Geol., 51, І–Geol. and Geophys., 127–132. Valchev, B., V. Stojanova, S. Juranov. 2013. New findings of Paleogene agglutinated and porcelaneous foraminifera from the Republic of Macedonia. – C. R. Acad. bulg. Sci., 66, 7, 1033–1042. Vassilenko, V. 1950. Paleocene foraminifers from the central part of Dnepr–Donets lowering. – Treat. VNIGRI, 51, Microfauna USSR, 4, 177–225 (in Russian). Vassilenko, V. 1954. Anomalinids. Treat. VNIGRI, New Ser., 80, 282 p. (in Russian). (Постъпила на 29.01.2014 г., приета за печат на 20.03.2014 г.) Отговорен редактор Димитър Синьовски 82 View publication stats