109
Ribera et al.: Checklist of Mediterranean seaweeds I. Fucophyceae
Botanica Marina
Vol. 35, pp. 109- 130, 1992
Offprint
Check-list of Mediterranean Seaweeds
I. Fucophyceae (Warming, 1884)
M. A. Ribera, A. Gomez Garreta, T. Gallardo*, M. Cormaci*", G. Furnari** and G. Giaccone**
Laboratorio de Bothnica, Facultad de Farmacia Universidad de Barcelona 08028 Bar-celom, Spain
* Departamerlto de Biologia Vegetal, I Facultad de Biologia Universidad Conzplulense 08040 Madrid, Spain
** Istituto e Orto Botanico dell'Univer-sita, Via A. Longo 19 95125 Catania, Italy
(Accepted 18 January 1992)
Abstract
A check-list of.Mediterranean brown algae, based on literature records, is given. The distribution of each
taxon in the area (which is divided into 16 regions) is reported. The number of taxa accepted for the
Mediterranean Sea under current taxonomy is 265. This list has benefited from the suggestions on taxonomy,
nomenclature and regional distribution made by algological advisers from each region.
Introduction
This check-list of Mediterranean seaweeds is intended
to be a catalogue of benthic algal taxa of the Mediterranean Sea, including only the Rhodophyceae, Fucophyceae (Phaeophyceae) and Chlorophyceae. The
project emerged at the XXXth Congress and Plenary
Assembly of International Commission for the Scientific Exploration of the Mediterranean Sea in Palma
de Mallorca (Spain) in 1986, and has been coordinated by a group of Spanish algologists.
The geographical area covered by this check-list is the
Mediterranean Sea, including both the Sea of Marmara and the Black Sea. The area is bounded on the
west by the Straits of Gibraltar. The Mediterranean
coasts have been divided in a similar way to that used
for the Med-Checklist of Vascular Plants (Greuter et
al., 1984). However, some modifications have been
made to account for seaweed biogeography and historical treatment of seaweed floras. For example, the
Adriatic Sea is considered as a single region because
of its biogeographical unity, while Malta and Sicily
are treated together, as they traditionally have been
treated when dealing with seaweed floras. As a result,
Botanica Marina / Vol. 35 / 1992 / Fasc. 2
Copyright 0 1992 Walter de Gruyter . Berlin . New York
the area has been split into 16 regions (Fig. 1) that
are labelled with two-letter symbols in the following
manner:
(Sp) Spain, (BI) Balearic Islands, (Fr) France, (CS)
Corsica and Sardinia, (WI) Western Italy, (Si) Sicily
and adjacent islands, (Ad) Adriatic Sea (including
Albania), (Gr) Greece, (BS) Black and Azov seas,
(TU) Turkey (Sea of Marmara and Mediterranean
coast), (LS) Levant States (Syria, Lebanon and Israel), (Eg) Egypt, (Li) Libya, (Tn) Tunisia, (Ag) Algeria, (MO) Morocco.
This first part comprises the Fucophyceae (Phaeophyceae). The number of specific and infraspecific
taxa accepted for the Mediterranean Sea under current taxonomy is 265. This work benefited from the
suggestions on taxonomy, nomenclature and regional
distribution of an advisory board of algologists. The
advisers were: V. Aysel (Tu), A. Bavaru (BS), A.
Bologa (BS), C. F. Boudouresque (Fr, CS, Tn), F.
Cinelli (WI), S. Cirik (Tu), A. Cossu (CS), A. Diapoulis (Gr), V. Gazale (CS), S. Haritonidis (Gr), I.
Munda (Ad) and G. Pardi (WI).
110
Ribera ef al.: Checklist of Mediterranean seaweeds I. Fucophyceae
Fig. 1. Geographical regions considered: (Sp) Spain, (BI) Balearic Islands, (Fr) France, (CS) Corsica and Sardinia, (WI) Western
Italy, (Si) Sicily and adjacent islands, (Ad) Adriatic Sea (including Albania). (Gr) Grecce, (BS) Black and Azov seas, (TU) Turkey
(Sea of Marmara and Mediterranean coast), (LS) Levant States (Syria, Lebanon and Israel), (Eg) Egypt, (Li) Libya, (Tn) Tunisia,
(Ag) AIgeria, (MO) Morocco.
Knowledge of Mediterranean seaweeds had increased
in the last few years to the point where it is useful, if
not essential, to assemble this information in one
publication. Our primary objective was to take the
available pool of information and produce a single
list and geographic index for the Mediterranean Sea.
However, this list is not a definitive catalogue, but
rather it is permanently open for additions and corrections.
Another objective was to standardize the nomenclature as far as possible, reconciling the numerous taxonomic and nomenclatural divergences between the
different local floras. In our opinion, recent taxonomic research has demonstrated clearly the need for
resolving problems on a global rather than a local
basis. This check-list has particular biogeographical
interest because the distribution of each species is
detailed for the different zones of the Mediterranean
Sea.
This list is based upon literature records. To minimize
the citation of synonyms and misapplied names, only
references published since 1950 have generally been
used. For Egypt, however, we have included works
by Nasr (1940) and Nasr and Aleem (1949) because
contemporary studies for this area are rare. When
available, regional floras have been used as the basis
for the compilation. References used for the areas are
shown in Tdble I.
Taxa are named with their authors, but dates of valid
publication are not given because of uncertainty in
many cases. When authors' names have been abbreviated, TL2 (Stafleu and Cowan, 1976- 1988) was
followed.
Both in the checklist and in the index names of accepted taxa are in italics whereas names of synonyms,
misapplied names and doubtful or unaccepted taxa
are in roman type.
The orders are arranged according to Wynne and
Kraft (1981) while families, genera and species are
arranged alphabetically.
When a specific or infraspecific taxon has definitely
been recorded for a geographical area, a nurnbcr
corresponding to the most important bibliographic
reference is cited in the list. Because of space limitations, only one reference is cited for each area, but
additional references are available from the authors
on request. Abbreviations of journals follow those of
B. P. H. (Lawrence er al., 1968).
When a specific or infraspecific taxon has been recorded under a synonym or misapplied name, the
reference nurnbcr in the list is in italics and the synonym or misapplied name, is given in roman below
the current namc.
Superscript numbers refer to notes.
A list of raxa excludenda as well as a list of raxa
inquirenda is also given.
Botanica Marina / VoI. 35 / 1992 / Fasc. 2
$
List of Taxa and Their Distribution
g.
Taxa
5.
ECTOCARPALES Setchell et Gardner
F
L
E!
-<
0
r
.
.
L
\O
\O
N
n
Geographic regions
Sp
B1
Fr
CS
Ectocarpaceae C. Agardh
Acinetospora Bornet
A. crinita (Carmich. ex Harv.) Sauv. . . . . . . . . . . . . . . . . . . . . . . . . . .
8
= A. vidovichii (Menegh.) Sauv.
= Haplospora vidovichii (Menegh.) Bornet
Ectocarpus Lyngb.
E. arabicus Fig. et De Not. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . E. caspicus Henckel . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . E. fmciculatus Harv.
8
var. fasciculatus . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
var. abbreviatus (Kiitz.) Sauv. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . = E. abbreviatus Kiitz.
var. pycnocarpus (Rosenvinge) Cardinal . . . . . . . . . . . . . . . . . . . . . . E. parvus (Saunders) Hollenberg . . . . . . . . . . . . . . . . . . . . . . . . . . . . . E. rhodochortonoides Boerg. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . E. siliculosus (Dillwyn) Lyngb.
= E. confe~oidesLe Jolis
var. siliculosus . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
8
var. adriaticus (ErcegoviC) Cormaci et Furnari . . . . . . . . . . . . . . . . . . var. arctus (Kiitz.) Gallardo4' . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 105
= E. arctus Kiitz.
var. crassus (Kjellm.) Gallardo4' . . . . . . . . . . . . . . . . . . . . . . . . . . . . var. crouanii (Thur.) Gallardo4' . . . . . . . . . . . . . . . . . . . . . . . . . . . . = E. crouanii Thur.
var. dasycarpus (Kuck.) Gallardo4' . . . . . . . . . . . . . . . . . . . . . . . . . . = E. dasycarpus Kuck.
var. hiernalis (Crouan frat. ex Kjellm.) Gallardo4' . . . . . . . . . . . . . . . . = E. hiemalis (Crouan frat. ex Kjellrn.) Kjellm.
var. penicillatus C. Agardh . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 105
= E. penicillatus (C. Agardh) KjelIm.
var. pygmaeus (Areschoug) Gallardo4' . . . . . . . . . . . . . . . . . . . . . . . . 17
var. subulatus (Kiitz.) Gallardo4' . . . . . . . . . . . . . . . . . . . . . . . . . . . var. venetus (Kiitz.) gal lard^^^ . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Feldrnanrlia Hamel
F. battersiides (Ercegovic) Cormaci et Furnari . . . . . . . . . . . . . . . . . . . . F. caespitula ( J . Agardh) Knoep.-Pkg.
8
var. caespitula3 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
var. Iebelii (Areschoug ex Crouan frat.) Knoep.-P&. . . . . . . . . . . . . . 8
= F. lebelii (Areschoug ex Crouan frat.) Hamel
8
F. globifera (Kiitz.) Hamel . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
F. irregularis (Kiitz.) Hamel . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
8
E padinae (Buffham) Hamel . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 105
8
F. paradoxa (Mont.) Hamel . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
90
22
-
-
-
-
90
22
-
21
-
-
-
-
90
-
90
-
-
22
-
-
21
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
34
-
-
-
-
90
-
22
22
21
-
22
22
-
21
21
114
-
-
90
90
-
90
-
Taxa
Geographic regions
Sp
B1
Fr
CS
E paradoxoides (ErcegoviC) Corrnaci et Furnari . . . . . . . . . . . . . . . . . . E simplex (Crouan frat.) Hamel . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
9
Herponema J. Agardh
H. grangerum Kuck. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
H. solitariunz (Sauv.) Hamel . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
8
H. valiantei (Born.) Hamel . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
8
= Ectocarpus valiantei Born.
H. velutinum (Grev.) J. Agardh . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
= Ectocarpus simpliciusculus C. Agardh
Hincksia J. E. Gray
H. dalmatica (ErcegoviC) Cormaci et Furnari4* . . . . . . . . . . . . . . . . . . . = Giffordia dalmatica (ErcegoviC) Cormaci et Furnari
H. fuscara (Zanardini) Silva . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
= Giffordia fuscata (Zanardini) Kuck.
H. geniculata (ErcegoviC) Cormaci et F ~ r n a r i. ~. ~
................. = Giffordia geniculata (ErcegoviC) Cormaci et Furnari
H. gratzulosa (Sm.) Silva . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8
= Giffordia granulosa (Sm.) Hamel
H. hauckii (ErcegoviC) Cormaci et Furnari" . . . . . . . . . . . . . . . . . . . . .
= Giffordia hauckii (ErcegoviC) Cormaci et Furnari
H. hincksiae (Harv.) Silva . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
= Giffordia hincksiae (Harv.) Hamel
H.rnitchelliae(Harv.)Silva . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
8
= Giffordia mitchelliae (Harv.) Hamel
H.ovata(Kjellm.)Silva . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
8
= Giffordia ovata (Kjellm.) Kylin
= G. intermedia (Rosenvinge) Lund
H. sandriana (Zanardini) Silva . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
8
= Giffordia sandriana (Zanardini) Hamel
H. secunda (Kiitz.) Silva . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 105
= Giffordia secunda (Kiitz.) Batters
Kuckuckia Hamel
K. kylinii Cardinal . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . K. spinosa (Kiitz.) Kuck. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8
Kuetzingiella Kornmann
K. battersii (Born.) Kornmann
= Feldrnannia battersii (Born.) Hamel
var. battersii . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
var. rnediterranea (Sauv.) G6mez et Kibera4' . . . . . . . . . . . . . . . . . . . 8
= Ectocarpus battersii var. rnediterraneus Sauv.
p
W1
Si
Ad
Gr
-
52
-
55
-
-
-
5
-
-
116
21
-
-
-
-
22
21
-
55
43
-
-
-
-
46
55
42
-
-
-
34
-
-
52
-
-
22
-
-
55
52
-
-
-
-
-
-
52
-
-
22
-
-
55
52
-
-
-
34
-
-
52
-
-
-
-
-
55
71
-
90
22
21
50
55
52
5
7
116
34
-
55
-
-
90
22
21
50
55
52
-
-
-
34
-
55
100
-
7
22
22
-
-
-
55
-
-
21
-
-
22
114
-
-
52
-
-
-
-
-
BS
-
-
5
-
Tu
-
Eg
Tn
Ag
MO
-
-
-
-
-
-
85
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
LS
-
84
-
84
S4
68
-
P
-
-
-
68
84
-
-
-
"51
2
-
15
85
-
-
-
-
-
-
-
102
15
85
-
-
-
-
85
-
-
-
-
-
-
-
85
-
-
-
-
P
-
Li
85'
-
-
$
5'
.
g
.
W
\O
\O
N
?I
N
Phaeostroma Kuck.
P. berfholdii Kuck. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
Pilayella Bory
P. littoralis (L.) Kjellm. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 27
Spongonema Kiitz.
S. tornenfosunz(Huds.) Kiitz. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
Streblonema Pringsheim
S. effusunz Kylin . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
= Entonema effusum (Kylin) Kylin
S. infestans (Gran) Batters . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . S. oligosporum Stromfelt . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
= Entonema oligosporum (Stromfelt) Kylin
S. parasiticum (Sauv.) De Toni . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
= Entonema parasiticurn (Sauv.) Hamel
S. sphaericum (DerbCs et Solier) Thur. . . . . . . . . . . . . . . . . . . . . . . . . .
8
S. stilophorae (Crouan frat.) De Toni . . . . . . . . . . . . . . . . . . . . . . . . . . 105
S. tenuissimurn Hauck . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
S. fhurefiiSauv. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Streblonemopsis Valiante
S. irritans Valiante . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
8
p
p
p
Ralfsiaceae Farlow
Acrospongium Schiffner
A. ralfsioides Schiffner . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 98
Mesospora Weber
M. tnacrocarpa (J. Feldm.) Hartog . . . . . . . . . . . . . . . . . . . . . . . . . . .
8
= M. mediterranea J. Feldm.
Microspongium Reinke
M. gelatinosum Reinke . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Nemoderma Schousboe ex Bornet
N. tingitanum Schousboe ex Bornet . . . . . . . . . . . . . . . . . . . . . . . . . . .
8
Pseudolithoderma Svedelius
P. adriaticum (Hauck) Verlaque . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
8
= Lithoderma adriaticum Hauck
P. exfensum (Crouan frat.) Lund . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
Ralfsia Berkeley
R. disciformis Crouan frat. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . R. expansa ( J . Agardh) J . Agardh . . . . . . . . . . . . . . . . . . . . . . . . . . . . 98
R.verrucosa(Areschoug)J.Agardh . . . . . . . . . . . . . . . . . . . . . . . . . . .
8
Stragularia Stromfelt
S. clavafa (Ham.) Hamel . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 28
= Ralfsia bornetii Kuck.
= R. clavata (Ham.) Crouan frat.
-
-
-
-
-
-
52
-
-
-
-
-
-
-
55
-
-
-
84
68
-
-
-
85
-
-
-
-
-
-
-
-
-
-
-
15
85
-
-
-
68
-
-
15
85
-
66
-
-
-
-
-
12
84
68
-
15
-
-
68
-
90
22
21
-
-
-
21
-
-
22
21
50
55
-
-
-
22
21
18
55
52
5
-
-
-
-
-
90
22
21
46
-
-
2
1
-
5
-
5
6
-
55
55
5
-
52
-
6
-
5
5
-
-
-
-
-
85
-
Taxa
Geographic regions
Sp
B1
Fr
CS
W1
Si
Ad
Gr
BS
Tu
46
-
31"
-'l
38
-
-
52
-
12
6
-
52
-
66
-
LS
Eg
Li
Tn
Ag
MO
85
-
CHORDARIALES Setchell et Gardner
Chordariaceae Grev.
Cladosiphon Kiitz.
C. chordariaeformis Crouan frat. . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
= Castagnea chordariaeformis (Crouan frat.) Thur.
C. contortus (Thur.) Kylin . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . = Castagnea contorta Thur.
C. cylindricus (Sauv.) Kylin . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8
= Castagnea cylindrica Sauv.
C. irregularis (Sauv.) Kylin . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
8
= Castagnea irregularis Sauv.
C. mediterraneus Kiitz. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8
= Castagnea mediterranea (Kiitz.) Hauck
= C. fistulosa (Zanardini) DerbCs et Solier
C. zosterae (J. Agardh) Kylin . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . = Castagnea zosterae (J. Agardh) Thur.
Eudesme J. Agardh
E. virescens (Carmich. ex Berk.) J. Agardh . . . . . . . . . . . . . . . . . . . . . . = Castagnea virescens (Carmich. ex Berk.) Thur.
Gorztrania Sauv.
G. lubrica Sauv. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Liebmannia J. Agardh
L. leveillei J. Agardh . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8
Mesogloia C. Agardh
M . lanosa Crouan frat. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . M . vermiculata (Sm.) S. F. Gray . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8
Sauvageaugloia Hamel ex Kylin
S. griffithsiana (Grev. ex Harv.) Hamel ex Kylin . . . . . . . . . . . . . . . . . . 8
Sphaerotrichia Kylin
S. divaricata (C. Agardh) Kylin . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Strepsithalia Bornet ex Sauv.
S. liagorae Sauv. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8
S. liebmanniae Miranda . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Corynophlaeaceae Oltmanns
Corynophlaea Kiitz.
C. Jaccida (C. Agardh) Kiitz. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 17
= Myriactis adriatica (J. Agardh) De Toni
C. hamelii J. Feldm. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . C. umbellata (C. Agardh) Kiitz. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8
Cylindrocarpus Crouan frat.
C. microscopicus Crouan frat. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8
-
-
21
-
22
22
34
-
22
-
-
-
46
P
115
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Botanica Marina / Vol. 35 / 1992 / Fasc. 2
I
1
Taxa
M . liechtensternii Hauck . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
= Phaeosphaerium liechtensternii (Hauck) De Toni
M . orbiculare J. Agardh . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
= M. magnusii (Sauv.) Loiseaux
= M. seriatum (Reinke) Kylin
= Ascocyclus orbicularis (J. Agardh) Magnus
M . strangulans Grev. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
Protectocarpus Kuck.
P. speciosus (Boerg.) Kornmann . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
Zosterocarpus Bornet
Z . oedogonium (Menegh.) Born. . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
Geographic regions
Sp
B1
Fr
CS
W1
Si
Ad
Gr
BS
Tu
LS
Eg
9
-
22
21
-
-
52
5
-
-
-
-
8
90
22
21
46
55
52
5
1214
-
68
2
8
90
22
21
46
55
52
5
12
-
68
-
-
-
-
-
-
-
-
-
38
-
-
-
8
90
13
21
46
-
52
-
-
-
-
-
-
22
21
46
55
52
5
-
6
-
-
-
-
-
-
-
-
22
-
46
55
52
90
22
-
21
l5
55
52
-
-
-
-
Spermatochnaceae Kjellm.
Nemacystus DerbCs et Solier
N.jlexuosus (C. Agardh) Kylin . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8
= N. rarnulosus Derbts et Solier
N. hispanicus (Sauv.) Kylin . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Spermatochnus Kutz.
S . paradoxus (Roth) Kutz. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Stilophora J. Agardh
S. rhizodes (Turner) J. Agardh" . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8
S. tuberculosa (Hornem.) Reinke . . . . . . . . . . . . . . . . . . . . . . . . . . . . . -
-
-
-
-
-
-
-
5
65
59
39
-
5
12
59
-
38
-
68
-
73
-
SPOROCHNALES Sauv
Sporochnaceae Grev.
Carpomitra Kiitz.
C . costata (Stackh.) Batters
var. costata . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
var. mediterranea J. Feldm. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
= Sporochnus dichotomus Zanardini
Nereia Zanardini
N.filiformis ( J . Agardh) Zanardini . . . . . . . . . . . . . . . . . . . . . . . . . . .
Sporochnus C. Agardh
S. gaertneri (Gmelin) C. Agardh . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
S. pedunculatus (Huds.) C. Agardh . . . . . . . . . . . . . . . . . . . . . . . . . . .
Stilopsis Kuck.
S. lejolisii (Thur.) Kuck. e x Hamel . . . . . . . . . . . . . . . . . . . . . . . . . . .
8
22
21
52
52
-
-
-
-
-
50
55
55
-
90
-
-
8
90
22
21
46
55
52
5
38
-
39
3
-
-
-
-
-
8
90
22
21
46
46
55
55
52
5
-
-
-
-
-
-
-
-
-
55
-
-
-
-
-
-
Desmarestiaceae (Thuret) Kjellm.
Desmarestia Lamour.
D . aculeata (L.) Lamour. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 27
D . adriatica ErcegoviC . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
-
-
-
-
-
-
-
-
52
-
-
-
-
-
-
-
p
DESMARESTIALES Setchell et Gardner
-
Li
Tn
Ag
MO
D . dresnayi Lamour. ex Leman . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
D . ligulata Lamour. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
D . viridis 0. F. Miill." . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
-
-
-
Arthrocladiaceae Chauvin
Arthrocladia Duby
A. villosa (Huds.) Duby" . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8
90
Dictyosiphonaceae Kiitz.
Dictyosiphon Grev.
D . chordaria Areschoug . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
-
-
Giraudiaceae Hamel ex J. Feldm.
Giraudia Derbts et Solier
G . sphacelarioides Derbts et Solier . . . . . . . . . . . . . . . . . . . . . . . . . . . .
8
90
DICTYOSIPHONALES Setchell et Gardner
Myriotrichiaceae Kjellm.
Myriotrichia Harvey
M. udriatica Ha'uck . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . M. clavaefornzis Harv. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . = M. repens Hauck
= Dichosporangium repens (Hauck) Hauck
Prutasperococcus Sauv.
P. rnyriotrichiiformis Sauv. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . = Myriotrichia protasperococcus Kuck.
Punctariaceae (Thuret) Kjellm.
Asperococcus Lamour.
A. bullosus Lamour.
= A. turneri (Sm.) Hook.
f. bullosus . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
f. profundus J. Feldm. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
A. compressus Griff. e x Hook. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
= Haloglossum compressum (Griff. ex Hook.) Hamel
A. fistulosus (Huds.) Hook. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
= A. echinatus (Mertens) Grev.
A. scaber Kuck. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
Punctaria Grev.
P. latifolia Grev. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
P. plantaginea (Roth) Grev. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
P. tenuissinia (C. Agardh) Grev. . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
= Desmotrichum undulatum (J. Agardh) Reinke
8
8
8
90
-
-
90
-
8
-
8
-
-
-
-
-
Striariaceae Kjellm.
Stictyosiphon Kiitz.
S. adriuticus Kiitz. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . S. suriferus (Reinke) Rosenvinge . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8
S. tortilis (Rupr.) Reinke9 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . -
-
7
p
p
112
-
Taxa
Geographic regions
Sp
B1
Fr
CS
Striaria Grev.
S. attenuata (Grev.) Grev.
f.attenuata . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . f. crinita (J. Agardh) Hauck. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . f. ramosissima (Kiitz.) Hauck. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . -
90
-
W1
Si
11621
-
46
55
-
46
-
p
-
-
SCYTOSIPHONALES Feldm.
Scytosiphonaceae Farlow
Colpomenia (Endlicher) Derb6s et Solier
C. peregrina (Sauv.) Hamel . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8
C. sinuosa (Mertens ex Roth) Derbbs et Solier . . . . . . . . . . . . . . . . . . . 8
Hydroclathrus Bory
H. clathratus (Bory ex C. Agardh) Howe . . . . . . . . . . . . . . . . . . . . . . . 8
Petalonia DerbCs et Solier
P. fascia (0. F. Miill.) Kuntze . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8
P. zosterifolia (Reinke) Kuntze . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Rosenvingea Boerg.
R. intricata (J. Agardh) Boerg. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Scy~osiphonC. Agardh
S. dotyi Wynne9 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . S. simplicissimus (Clemente) Cremades2*
8
= S. lomentaria (Lyngb.) Link.
TILOPTERIDALES Sauv.
Tilopteridaceae Sauv.
Climacosorzts Sauv.
C. mediterraneus Sauv. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . -
-
CUTLERIALES Kjellm.
Cutleriaceae Hauck
Cutleria Grev.
C. adspersa (Roth) De Not.23 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
C. chilosa (Falkenb.) Silva" . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
= C. monoica Ollivier
C.mullifda(Sm.)Grev .25 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
Zanardinia Nardo ex Crouan frat.
Z. prototypus (Nardo) Nardo . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
8
8
9026
902'
22
21
22
21
46
46
55
8
90
22
21
46
55
8
90
22
21
24
55
8
90
30
21
46
-
SPHACELARIALES Oltmanns
Choristocarpaceae Kjellm.
Choristocarpus Zanardini
C. tenellus (Kiitz.) Zanardini . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
Ad
Gr
BS
Tu
LS
Eg
Li
Tn
Ag
MO
Ribera et al.: Checklist of Mediterranean seaweeds I. Fucophyceae
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V ) ,
Botanica Marina / Vol. 35 / 1992 / Fasc. 2
mr-
m
m
119
Taxa
Geographic regions
Sp
B1
Fr
CS
W1
Si
Ad
Gr
-
-
BS
D. repens (J. Agardh) J . Agardh . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
8
= D. fasciola var. repens (J. Agardh) J. Fcldm.
D. spiralis (Mont.) Hamel . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
8
= D. ligulatus (Kutz.) J. Feldm.
Lobophora J. Agardh
L. variegata (Lamour.) Womersley . . . . . . . . . . . . . . . . . . . . . . . . . . . . 104
= Pocockiella variegata (Lamour.) Papenfuss
Padina Adanson
P. boergesenii Allendcr et Kraft . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . = P. gymnospora (Kiitz.) Vickers
P. pavonica (L.) Lamour. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
8
P. tenuis Bory . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
Spatoglossurn Kutz.
S. schroederi (C. Agardh) Kutz. . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
S. solieri (Chauv. ex Mont.) Kiitz. . . . . . . . . . . . . . . . . . . . . . . . . . . . .
8
S. variabile Fig. et De Not. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
Stypopodiunl (Kiitz.) J . Agardh
S. schitperi (Buchinger ex Kiitz.) Verlaque et Boudour. . . . . . . . . . . . . . = S. tubruquense Nizamuddin et Godeh
Taonia J. Agardh
7: atomaria (Woodw.) J . Agardh
f. atomaria . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
8
f. ciliata (Kutz.) Nizamuddin . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
= var. ciliata Kiitz.
Zonaria Drapanald
Z. tournefortii (Lamour.) Mont. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 27
-
LAMINARIALES Kylin
Alariaceae Setchell et Gardner
Undaria Suringar emend. Okamura
U. pinnatiJda (Harv.) S ~ r i n g a r ' ~. . . . . . . . . . . . . . . . . . . . . . . . . . . . .
14
-
Chordaceae Dumortier
Chorda Stackh.
C . Jlum (L.) Stackh." . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
14
-
14
-
13
21
86
55
55
52
-
-
-
46
55
-
5
-
Laminariaceae Bory
Laminaria Lamour.
L . japonica Areschoug3' . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
L . ochroleuca Pylaie . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
L.rodrigueziiBorn . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
Saccorhiza Pylaie
S. polyschides (Lightf.) Batters . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
= S. bulbosa (Huds.) Pylaie
-
27
-
8
90
-
-
-
-
Tu
LS
Eg
Li
Tn
Ag
MO
g
0
2.
51.
2
.
.
.
Phyllariaceae Tilden
Phyllariopsis Henry et South
P. brvvipes (C. Agardh) Henry et South . . . . . . . . . . . . . . . . . . . . . . .
= Phyllaria renifonnis (Lamour.) Rost.
P. purpurascens (C. Agardh) Henry et South . . . . . . . . . . . . . . . . . . . .
= Phyllaria purpurascens (C. Agardh) Rost.
FUCALES Kylin
?
h,
-r
E
0
h,
Cystoseiraceae Kiitz.
Cystoseira C. Agardh
C . abies-marina (S. G. Gmelin) C. Agardh" . . . . . . . . . . . . . . . . . . . . .
C . algeriensis J . Feldm. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
C . amentacea Bory
= C. stricta Sauv. v. amentacea (Bory) Giaccone
var. amentacea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
var. spicata (Ercegovii:) G i a c ~ o n e. ~. .~. . . . . . . . . . . . . . . . . . . . . . .
= C. spicata Ercegovii:
= C. spicata subsp. elegans Ercegovii:
= C. stricta Sauv. v. spicata (ErcegoviC) Giaccone
var. stricta Montagne . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
= C. stricta Sauv. v. stricta
C . barbata C. Agardh
var. barbata . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
f. aurantia (Kiitz.) Giaccone . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
= f. repens Zinova et Kalugina
f. j7accida (Kiitz.) Woronich. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
f. insularum ErcegoviC . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
var. tophuloidea (ErcegoviC) Giaccone . . . . . . . . . . . . . . . . . . . . . . . .
= subsp. tophuloidea Ercegovii:
C . barbatula Kiitz.16 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
= C. graeca Schiffn. ex Gcrloff et Nizamuddin
C . brachycarpa J. Agardh emend. Giaccone
var. brachycarpa . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
var. balearica (Sauv.) Giaccone . . . . . . . . . . . . . . . . . . . . . . . . . . . .
= C. balearica Sauv. v. balearica
= C. caespitosa
var. claudiae (Giaccone) Giaccone . . . . . . . . . . . . . . . . . . . . . . . . . .
= C. balearica Sauv. v. claudiae Giaccone
C . compressa (Esper) Gerloff et Nizamuddin
= C. fimbriata (Desf.) Bory
f.compressa . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
f. plana (ErcegoviC) Cormaci et al.'? . . . . . . . . . . . . . . . . . . . . . . . .
=
subsp. plana ErcegoviC
f. rosetta (ErcegoviC) Cormaci et
=
subsp. rosetta Ercegovii:
aLS6 .
.......................
Taxa
C. corniculata (Wulfen) Z a n a r d i ~38~ i.~. ~. . . . . . . . . . . . . . . . . . . . . . . .
C. crinita (Desf.) Bory
f. crinita . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
f. bosphorica (Sauv.) Zinova et Kalugina . . . . . . . . . . . . . . . . . . . . . .
C. crinitopliylla ErcegoviC . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
C. dubia Valiante . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
= C. fucoides ErcegoviC
C. elegans Sauv. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
C. foeniculacea (L.) Grev. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
C. humilis Kiitz.
vachumilis . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
= C. compressa var. pustulata Ercegovii:
var. myriophylloides (Saw.) Price et John39 . . . . . . . . . . . . . . . . . . . .
= C. myriophylloides Sauv.
C. hyblaea Giaccone . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
C . jabukae ErcegoviC
f. jabukae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
f. tenuissima (Ercegovik) Cormaci et al. . . . . . . . . . . . . . . . . . . . . . . .
= subsp. tenuissima ErcegoviC
C. mauritanica S a ~ v . ~.' . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
C. mediterranea Sauv.
var. mediterranea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
var. valiautei Sauv. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
C . nodicattlis (With.) Roberts . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
C. pelagosae ErcegoviC . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
C . sauvageauana Hamel . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
= C. selaginoides Valiante
C . schiffneri Hamel
= C. discors (L.) C. Agardh f. discors
= C. discors (L.) C. Agardh f. dubia ErcegoviC
= C. ercegovicii Giaccone f. ercegovicii
= C. ercegovicii Giaccone f. schiffneri (Hamel) Giaccone
f.schiffneri . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
f. latiramosa (Ercegovik) G i a c c ~ n e.~. ~. . . . . . . . . . . . . . . . . . . . . . .
= C. ercegovicii f. latiramosa (ErcegoviC) Giaccone
f. tenuiramosa (ErcegoviC) Giaccone4* . . . . . . . . . . . . . . . . . . . . . . . .
= C. ercegovicii f. tenuiramosa (ErcegoviC) Giaccone
C . sedoides (Desf.) C. Agardh . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
C. spinosa Sauv.
var. spinosa4' . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
var. compressa (ErcegoviC) Cormaci et ~ 1 . 4.~. . . . . . . . . . . . . . . . . . .
= C. adriatica var. compressa (ErcegoviC) Giaccone
= C. platyramosa ErcegoviC
var. tenuior (ErcegoviC) Cormaci et al." . . . . . . . . . . . . . . . . . . . . . .
= C. adriatica f. tenuior (ErcegoviC) Giaccone
= C. gerlofii Nizamuddin
= C. rechingeri Schiffn. ex Gerloff et Nizamuddin
Geographic regions
Sp
B1
Fr
CS
8
-
8
27
56
-
-
51
8
27
8
8
8
8
Ribera et al.: Checklist of Mediterranean seaweeds I. Fucophyc
Notes
;
; ;jZl
Botanica Marina / Vol. 35 / 1992 / Fasc. 2
;l
1. Giaccone (1985: 697) cites both Acinetospora crinita and A.
vidovichii, which are synonyms.
2. Giaccone (1978: 80, 89) cites both Acinetospora crinita and
Haplospora vidovichii, which are synonyms.
3. We follow Knoepffler-Pi-guy (1970) in considering the records of Feldmannia paradoxa (Mont.) Hamel sensu Hamel
to be referred to Feldmannia caespitula (J. Agardh) Knoep.P6g.
4. According to Ben Maiz et al. (1987), the presence of this
species in Tunisia needs to be confirmed.
5. Perret-Boudouresque and Seridi (1989: 57) include in their
citation var. mediterranea.
6. According to Athanasiadis (1987), the presence of this
species in Greece needs to be confirmed.
7. Athanasiadis (1987: 108) does not consider this species to
belong to the Greek flora.
8. Although there is no post-1950 record from that area,
Hauck's record has been taken into account because the
Adriatic Sea is the type locality.
9. This species is believed to have been accidentally introduced
from Japan into the Thau lagoon (France), where it thrives
under favourable environmental conditions (Riouall, 1985:
86).
10. Athanasiadis (1987: 114) excludes this species from the
Aegean Sea.
11. Athanasiadis (1987: 112) excludes this species from both
Greece and the Adriatic Sea.
12. Giaccone (1978: 93, 94) records both this species and E.
kuckuckiana Schiffner, but the latter is a synonym of the
former.
13. Elachista neglecta Kuck. is possibly a synonym of E. intermedia Crouan frat., according to Feldmann (1937: 272).
14. Kalugina-Gutnik (1975: 227) records separately both Myrionema seriatum (Reinke) Kylin and Ascocyclus orbicularis
(J. Agardh) Magn.
15. Hamel (1937: 185) pointed out that Mediterranean specimens belong to var. adriatica J. Agardh.
16. Perret-Boudouresque and Seridi (1989: 52) include in their
citation var. mediterranea.
17. Verlaque (1981) suggests that D. adriatica Ercegovii: may
be a deep-water form of D. viridis.
18. This species includes f. australis (Kiitz.) Hauck (Ad 92, BS
116) and f. tenuissima Woronich (BS 116).
19. Funk (1955: 41) records both this species and Dichosporangium repens (Hauck) Hauck, but the latter is a synonym
of the former.
20. Giaccone (1978: 85, 86) records both this species and Myriotrichia protasperococcus Berthold, but the latter is a
synonym of the former.
21. Athanasiadis (1987: 118) does not recognize this species
either in Greece or in Turkey considering it a cold-water
alga that occurs in the Mediterranean Sea only in Sicily.
22. Regarding this combination, see Cremades and PCrez-Cirera (1990: 492).
23. This species includes the sporophytic stage, Aglaozonia
melanoidea Schousboe ex Bornet.
24. This specific name is based on the sporophytic stage, Aglaozonia chilosa Falkenberg. The species includes the gametophytic stage, Cutleria monoica Ollivier.
25. The species includes the sporophytic stage, Aglaozonia parvula (Grev.) Zanardini.
26. Sporophyte not recorded.
27. Gametophyte not recorded.
28. Prud'homme van Reine (1982: 232) suggests that S. hystrix
is an ecad of S. cirrosa. Its geographical distribution is:
Sp 8, B1 94, Fr 22, CS 34, W1 45, S1 55, Ad 52, Gr 47, Tu 26,
LS 68, Tn 15.
29. Prud'homme van Reine (1982: 103) considers the three
Mediterranean records doubtful.
124
Ribera et al.: Checklist of Mediterranean seaweeds I. Fucophyceae
According to Hornig and Schnetter (1988: 284) the Mediterranean Dictyola would consist of only D. dichoroma,
including two varieties, namely, var. dicholoma and var.
intricata. The latter would include D. linearis (C. Agardh)
Grev. and D. pusilla Lamour. The records of the taxon: D.
linearis f. divaricala Kiitz., which was never described but
is reported in some papers (see references: 47, 50, 52, 55),
are referred to D. linearis.
According to Boudouresque and Perret-Boudouresque
(1987: 64), this record requires substantiation, since it may
possibly involve misinterpretation or labelling error.
This species was accidentally introduced into the Mediterranean Sea from Japan (Boudouresque, Gerbal and
Knoepffler-PBguy, 1985: 364).
The Thau lagoon apparently provides a boreal environment
in the Mediterranean Sea for this species (Riouall, 1985:
86).
Amico et al. (1985: 899) suggest that the presence of this
species in the Mediterranean Sea needs to be confirmed.
Giivcn and t)ztig (1971: 124) and Nizamuddin er al. (1979:
469) record both this species and C . hoppii C. Agardh, but
the latter is a synonym of the former.
For the synonymization of Cystoseira barbatula and C.
graeca, see Cormaci et al. (1992).
This species includes C. corniculata f. imperfecta ErcegoviC
and C. corniculta f. clydonophila Cormaci et al. (Cormaci
et al., 1992).
C. cornicula1a subsp. Iaxior Ercegovik and C . corniculata
subsp. divergens Ercegovii: are synonyms of C. corniculala
(Wulfen) Zanardini (Cormaci et al., 1992).
The presence of this variety in the Mcditerranean Sea needs
to be confirmed. Athanasiadis (1987: 135) states that the
Greek record might be an error.
This species is included in C . gibraltarica (Sauv.) Dangeard
by Roberts (1978: 402).
This species includes C. adriatica Sauv. (Cormaci et al.,
1992).
This variety includes C. adriatica Sauv. var. intermedia
(ErcegoviC) Giaccone (Cormaci et al., 1992).
This variety includes C. adriatica Sauv. f. reducta (Ercegovik) Giaccone (Cormaci et al., 1992).
Although Hamza (1987: 64) records this species [as C.
ericoides (L.) C. Agardh] from Tunisia, we believe that his
material may be referable to C. mediterranea. Formerly, C.
ericoides had been recorded from Tunisia (Pottier, 1929),
but Ben Maiz et al. (1987: 291) consider this record to
apply to either C. medilerranea or C . stricta.
This species includes var. axillaris (J. Agardh) Mangenot.
Funk (1955: 53) cited this species from Naples on the basis
of a dead specimen carried by a ship.
This species deserves careful taxonomic study. Its complex
taxonomy has not allowed us to consider its varieties here.
At least var. vulgare is recorded everywhere in the Mediterranean Sea except LS (68), where only var. diversijolium
Bory and var. megalophyllum (Mont.) Grun. have been
recorded.
Regarding this combination see Gallardo, T. 1992. Nomenclatural notes on some Mediterranean algae, I: Phaeophyceae. Taxon, 41: in press.
According to Verlaque el Boudouresque (1991), Mediterranean records of Slypopodium zonale (Lamour.) Papenfuss
should be referred to S. schimperi.
Nomenclatural changes
On the basis of a comparative study (unpublished),
Cystoseira brachycarpa J . Agardh and C , balearica
Sauv. are distinguishable only at varietal level. The
following new combinations are proposed:
Cystoseira brachycarpa J . Agardh emend. Giaccone
var. balearica (Sauv.) Giaccone comb. et stat. nov.
[Basionym: C. balearica Sauv. (1912: 396)l.
Cystoseira brachycarpa J. Agardh emend. Giaccone
var. claudiae (Giaccone) Giaccone comb. nov.
[Basionym: C. balearica Sauv. var. claudiae Giaccone (in Amico et al., 1985: 906)l.
Taxa excludenda
Chilionema reptans (Crouan) Sauv.: (Gr 47). Athanasiadis (1987: 108) excludes this taxon from the
Greek flora (as Ectocarpus reptans P. et H .
Crouan).
Cystoseira baccata (Gmel.) Silva: (Gr 62). Restricted
to the Atlantic Ocean (Oliveras PIS and G6mez
Garreta, 1989: 92).
Cystoseira Junkii Schiffn, ex Gerloff et Nizamuddin:
(W1 49). Giaccone (1985: 436) considers this species to be referable to C. mediterranea var, valiantei
Sauv.
Cystoseira planiramea Schiffn. ex Gerloff et Nizamuddin: (Ad 48, Gr 48). This species may refer to
C . humilis Kutz. according to Giaccone (1985:
436). Athanasiadis (1987) considers C . planiramea
to lie with the range of C . compressa (Esper)
Gerloff et Nizamuddin.
Cystoseira pycnoclada Schiffn. ex Gerloff et Nizamuddin: (Ad 49) = C. schzj'Jneri Hamel f. schff
neri according to Giaccone (1985: 435).
Cystoseira sauvageauana var. polyoedenzatis (Sauv.)
Hamel: (Sp 27, Fr 99, W1 46, Si 55, Ag 85). According to Motta (1989: 325) this taxon does not
differ from the type variety.
Cystoseira sicula Schiffn. ex Gerloff et Nizamuddin:
(Si 79). Giaccone (1985: 436) states that it is a
juvenile form of C . sauvageauana Hamel.
Dictyota major Taylor: (Si 55). Hornig et Schnetter
(1988: 282) point out that the record of D. major
from Sicily (Giaccone et al. 1985: 713) should be
referred to D. dichotoma (Huds.) Lamour.
Sphacelariaplumosa Lyngbye: [W1 46, as Chaetopteris
plumosa (Lyngb.) Kiitz.]. Athanasiadis (1987: 123)
excludes this species from the Greek flora since it
is a boreal species. Prud'homme van Reine (1982:
116) states that its Mediterranean records must be
erroneous.
Stypopodium zonale (Lamour.) Papenfuss. According
to Verlaque et Boudouresque (1991), Mediterranean records of this species should be referred to
S. schimperi.
Botanica Marina / Vol. 35 / 1992 / Fasc. 2
125
Ribera et al.: Checklist of Mediterranean seaweeds I. Fucophyceae
Taxa inquirenda
Table I. References used for the different regions.
Adriogloia adriatica Ercegovik (Ad 52).
Cystoseira epiphytica Schiffn. e x Gerloff et Nizamuddin (Ad 49). Giaccone (1985: 435) believes that
this species is a juvenile form of C . humilis Kiitz.
Cystoseira gibraltarica (Sauv.) Dangeard may be a
synonym of C . mauritanica Sauv. (Sp 99, MO 76)
according to Roberts (1978: 402).
Cystoseira nzontagnei J . Agardh (B1 90, CS 21, Li 82,
Tn 15, Ag 85).
Cystoseira platyclada Sauv. (Sp 27, Si 55, Ag 85,
MO 76).
Dalmatogloia bryozoi Ercegovik (Ad 52).
Desmotrichum punctarioides A. Zin. et Dim.-Kon.
This species probably is a stage of some Punctaria
species.
Dictyopteris tripolitana Nizamuddin (B1 79, W1 79,
Si 79, Ad 79, Gr 79, Li 79, Tn 79, Ag 79).
Dilophus mediterraneus Schiffn. var. crassus Schiffn.
(Ad 52).
Ectocarpus congestus Crouan frat. (Ad 100).
Ectocarpus humilis Kiitz. (W1 46).
Ectocarpus siliculosus var. divergens Schiffn. (Ad 52).
Ectocarpus siliculosus var. elongatus Schiffn. (Ad 52).
Ectocarpus siliculosus var. megacarpus Schiffn. (Ad
52).
Microsporangiunz kuckuckianum Schiffn. (Ad 52).
Myriactula microscopica (Ercegovik) Ercegovik
(Ad 52).
Padinopsis adriatica Ercegovik (Ad 52).
Spermatocht~usparadoxus var. adriaticus ErcegoviC
(Ad 52).
Sp:
BI:
Fr:
CS:
WI:
Si:
Ad:
Gr:
BS:
Tu:
LS:
Eg:
Li:
Tn:
8
7
13
34
10
4
31
5
11
6
39
1
78
15
Ag: 85
MO: 76
9
35
14
21
18
33
41
16
12
26
68
2
79
20
17
56
22
114
23
55
42
37
38
58
77
3
80
61
27 28
79 90
29 30
51
94
79
97 99 111 112 113
24
79
43
47
65
59
89
73
81
79
40
45 46
25
56 98
9 9 1 0 4 1 0 5 106
50
54
79
52 71 72 79
62 109 110
66 101 116
84 108
92 100 103
74 75
82 102
95
79 88
86 96 99
88
Acknowledgements
We sould like to thank Dr P. C. Silva for his nomenclatural observations and suggestions; Dr M. Alvarez
and Mr R. Rycroft for improving the English of this
manuscript; Dr F. Conde for his bibliographical support; Mr J. Rull, Mr J. Marquez and Mr J. Sim6n
for the computer support; Dr J. Va1li.s for translating
Russian works; Dr C. Bened6 and Dr C. Blanch6 for
their aid in resolving taxonimic problems.
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Alphabetical List of Species
Acinetospora crinita
Acinetospora vidovichii
Acrospongium ralfsioides
Adriogloia adriatica
Aglaozonia chilosa
Aglaozonia melanoidea
Aglaozonia parvula
Arthrocladia villosa f. australis
Arthrocladia villosa f. tenuissima
Arthrocladia villosa f. villosa
Ascocyclus orbicularis
Asperococcus brrllosus f. bullosus
Asperococc~lsb~rllosusf. profundus
Asperococcus compressus
Asperococcus echinatus
Asperococcus fistulosus
Asperococcus scaber
Aspetococcus turneri
Carpomitra costata var. costata
Carpomitra costata var. mediterranea
Castagnea chordariaeformis
Castagnea contorta
Castagnea cylindrica
Castagnea fistulosa
Castagnea irregularis
Castagnea mediterranea
Castagnea virescens
Castagnea zosterae
Chaetopteris plumosa
Chilionema hispanicum
Chilionema reptans
Chorda /ilum
Choristocarpus tenellus
Cladosiphon chordariaeformis
Cladosiphon contortus
Cladosiphon cylindricus
Cladosiphon irregularis
Cladosiphon mediterraneus
Cladosiphon zosterae
Cladostephus hirsutus
Cladostephus spongiosus f. spongiosus
Cladostephus spongiosus f. verticillatus
Cladostephus verticillatus
Climacosorus mediterraneus
Colpomenia peregrina
Colpomenia sinuosa
Compsonema gracilis
Compsonema minutum
Corynophlaea jlaccida
Corynophlaea hamelii
Corynophlaea umbellata
Cutleria adspersa
Cutleria chilosa
Cutleria monoica
Cutleria multifida
Cylindrocarpus berkeleyi
Cylindrocarpus microscopicus
Cystoseira abies-marina
Cystoseira adriatica var. adriatica
Cystoseira adriatica var. compressa
Cystoseira adriatica var. intermedia
Cystoseira adriatica f. reducta
Cystoseira adriatica f. tenuior
Cystoseira algeriensis
Cystoseira arnentacea var. amentacea
Cystoseira amerltacea var. spicata
Cystoseira amentacea var. stricta
Cystoseira baccata
Cystoseira balearica var. balearica
Cystoseira balearica var. claudiae
Cystoseira barbata f. auranlia
Cystoseira barbata var. barbata
Cystoseira barbata f. jlaccida
Cystoseira barbata f. insularum
Cystoseira barbata f. repens
Cystoseira barbata subsp. tophuloidea
Cystoseira barbata var. tophuloidea
Cystoseira barbatula
Cystoseira brachycarpa var. balearica
Cystoseira brachycarpa var. brachycarpa
Cystoseira hrachycarpa var. claudiae
Botanica Marina / Vol. 35 / 1992 / Fasc. 2
Ribera et al.: Checklist of Mediterranean seaweeds I. Fucophyceae
Cystoseira caespitosa
Cystoseira cornpressa f. compressa
Cystoseira compressa f. plana
Cystoseira compressa subsp. plana
Cystoseira compressa f. rosetta
Cystoseira compressa subsp. rosetta
Cystoseira compressa var. pustulata
Cvstoseira corniculata
Cystoseira corniculata f. clydonophila
Cvstoseira corniculata f. im~erfecta
Cystoseira corniculata subsp. divergens
Cvstoseira corniculata subs^. laxior
Cystoseira crinita f. bosphorica
Cystoseira crinita f. crinita
Cystoseira crinitophylla
Cystoseira discors f. discors
Cystoseira discors f. dubia
Cystoseira dubia
Cystoseira elegans
Cystoseira epiphytica
! Cystoseira ercegovicii f. ercegovicii
Cystoseira ercegovicii f. latiramosa
Cystoseira ercegovicii f. schiffneri
Cystoseira ercegovicii f. tenuiramosa
Cystoseira ericoides
Cystoseira fimbriata
Cystoseira foeniculacea
Cystoseira fucoides
Cystoseira funkii
Cystoseira gerlofii
Cystoseira gibraltarica
Cystoseira graeca
Cystoseira hoppii
Cystoseira humilis var. humilis
Cystoseira humilis var. myriophylloides
Cystoseira hyblaea
Cystoseira jabukae f. jabukae
Cystoseira jabukae f. tenuissima
Cystoseira jabukae subsp. tenuissima
Cystoseira mauritanica
Cystoseira mediterranea var.
mediterranea
Cystoseira mediterranea var. valiantei
Cystoseira montagnei
Cystoseira myriophylloides
Cystoseira nodicaulis
Cystoseira pelagosae
Cystoseira planiramea
Cystoseira platyclada
Cystoseira platyramosa
Cystoseira pycnoclada
Cystoseira rechingeri
Cystoseira sauvageauana
Cystoseira sauvageauana var.
polyoedematis
Cystoseira schiffneri f. latiramosa
Cystoseira schiffneri f. schiffneri
Cystoseira schiffneri f. tenuiramosa
Cystoseira sedoides
Cystoseira selaginoides
Cystoseira sicula
Cystoseira spicata
Cystoseira spicata subsp. elegans
Cystoseira spinosa var. compressa
Cystoseira spinosa var. spinosa
Cystoseira spinosa var. squarrosa
Cystoseira spinosa var. tenuior
Cystoseira squarrosa
Cystoseira stricta var. amentacea
Cystoseira stricta var. spicata
Cystoseira stricta var. stricta
Cystoseira susanensis
Cystoseira tamariscifolia
Botanica Marina / Vol. 35 / 1992 / Fasc. 2
Cystoseira irsneoides
Cystoseira zosteroides
Dalrnatogloia bryozoi
Desmarestia aculeata
Desmarestia adriatica
Desrnarestia dresnayi
Desmarestia ligulata
Desmarestia viridis
Desmotrichum punctarioides
Desmotrichum undulatum
Dichosporangium repens
Dictyopteris membranacea
Dictyopteris polypodioides
Dictyopteris tripolitana
Dictyosiphon chordaria
Dictyota dichotoma var. dichotoma
Dictyota dichotoma var. implexa
Dictyota dichotoma var. intricata
Dictyota divaricata
Dictyota linearis
Dictyota major
Dictyota pusilla
Dilophus fasciola
Dilophus fasciola var. repens
Dilophus ligulatus
Dilophus mediterraneus
Dilophus mediterraneus var. crassus
Dilophus repens
Dilophus spiralis
Discosporangium mesarthrocarpum
Ectocarpus abbreviatus
Ectocarpus arabicus
Ectocarpus arctus
Ectocar~usbattersii var. mediterraneus
Ectocarpus caspicus
Ectocar~usconfervoides
Ectocarpus congestus
Ectocar~uscrouanii
Ectocarpus dasycarpus
Ectocar~usfasciculatus var. abbreviatus
~ctocar>us>ascictrlatus var. fascilulatus
Ectocarpus fasciculatus var. pycnocarpus
Ectocarpus hiemalis
Ectocarpus humilis
Ectocarpus paradoxus
Ectocarpus parvus
Ectocarpus penicillatus
Ectocarpus reptans
Ectocarpus rhodochortonoides
Ectocarpus siliculosus var. adriaticus
Ectocarpus siliculosus var. arctus
Ectocarpus siliculosus var. crassus
Ectocarpus siliculosus var. crouanii
Ectocarpus siliculosus var. dasycarpus
Ectocarpus siliculosus var. divergens
Ectocarpus siliculosus var. elongatus
Ectocarpus siliculosus var. hiemalis
Ectocarpus siliculosus var. megacarpus
Ectocarpus siliculosus var. penicillatus
Ectocarpus siliculosus var. pygmaeus
Ectocarpus siliculosus var. siliculosus
Ectocarpus siliculosus var. subulatus
Ectocarpus siliculosus var. venetus
Ectocarpus simpliciusculus
Ectocarpus valiantei
Elachista flaccida
Elachista fucicola
Elachista intermedia var. clavaeformis
Elachista intermedia var. intermedia
Elachista jabukae
Elachista kuckuckiana
Elachista negkcta
Elachista scutulata
129
Elachista stellaris
Entonema effusum
Entonema oligosporum
Entonema parasiticum
Eudesme virescens
Feldmannia battersii
Feldmannia battersiides
Felhannia caespitula var. caespitula
Feldmannia caespitula var. lebelii
Feldmannia globifera
Feldmannia irregularis
Feldmannia lebelii
Feldmannia padinae
Feldmannia paradoxa
Feldmannia paradoxoides
Felhannia simplex
Fuczrs spiralis
Fucus vesiculosus
Fucus vesiculosus var. axillaris
Fzicus virsoides
Giffordia dalmatica
Giffordia fuscata
Giffordia geniculata
Giffordia granulosa
Giffordia hauckii
Giffordia hincksiae
Giffordia intermedia
Giffordia mitchelliae
Giffordia ovata
Giffordia sandriana
Giffordia secunda
Giraudia sphacelarioides
Gontrania lubrica
Haloglossum compressum
Halopteris filicina
Halopteris scoparia
Halothrix lumbricalis
Haplospora vidovichii
Hecatonema liagorae
Hecatonema maculans
Herponema graniferum
Heroonema solitarium
Herponema valiantei
Herponema velutinum
Hincksia dalmatica
Hincksia fuscata
Hincksia geniculata
Hincksia granulosa
Hincksia hauckii
Hincksia hincksiae
Hincksia mitchelliae
Hincksia ovata
Hincksia sandriana
Hincksia secunda
Hydroclathrus clathratus
Kuckuckia kylinii
Kuckuckia spinosa
Kuetzingiella battersii var. battersii
Kuetzingiella batt~rsiivar. rnediterranea
Laminaria japonica
Laminaria ochroleuca
Laminaria rodriguezii
Leathesia difformis
Leathesia mucosa var. condensata
Leathesia mucosa var. mucosa
Leptonema neapolitanum
Leptonematella fasciculata
Leptonematella neapolitana
Liebmannia leveillei
Lithoderma adriaticum
Lobophora variegata
Mesogloia lanosa
Mesogloia vermiculata
130
Mesospora macrocarpa
Mesospora mediterranea
~ i c r o s ~ o n ~ gelatinosum
ium
Microsporangium kuckuckianum
~ ~ r i a c iadriatica
is
Myriactrda arabica
Myriactula elongata
Myriactula gracilis
Myriactula microscopica
Myriactula rigida
Myriactula rivrrlariue
Myriactula stellulata
Myrionema balticum
Myrionema conchicola
Myrionema liechtensternii
Myrionema magnusii
Mvrionema orbiculrre
Myrionema seriatum
Myrionema strangulans
Myriotrichia adriatica
Myriotricl~iaclavaeformis
Myriotrichia protasperococcus
Myriotrichia repens
Nemacystus Jlexuosus
Nemacystus kispanicus
Nemacystus ramulosus
Nemoderma tingitanum
Nereia Jilijbrmis
Padina boergesenii
Padina gymnospora
Padina pavonica
Padina tenuis
Padinopsis adriatica
Petalonia fbscia
Petalonia zosterijolia
Petrospongium berkeleyi
Phaeosphaerium liechtensternii
Phaeostroma bertholdii
Phyllaria purpurascens
Phyllaria reniformis
Phyllariopsis brevipes Phyllariopsis purpurascens
Pilayella littoralis
Pocockiella variegata
Ribera et al.: Checklist of Mediterranean seaweeds I. Fucophyceae
Protasperococcus myriotrichiijbrmis
Protectocarpus speciosus
Pseudoli~hoderrnaadriaticum
Pseudolithoderrna extensum
Punctaria lalijolia
Punctaria plantaginea
Punctaria lenuissima
Ralfsia bornetii
Ralfsia clavata
RaljXa discijorrnis
Ra&ia expansa
Ra!i.sia verrucosa
Rosen vingea intricata
Saccorhiza bulbosa
Saccorhiza polyschides
Sargassum acinarium
Sargassum jlavijolium
Sargassum hornschuchii
Sargassum linifolium
Sargassum mulicunr
Sargassum salicifolium
Sargassum trichocarpum
Sargassum vulgare var. diversifolium
Sargassum vulgare var. megalophyllum
Sargassum vulgare v. vulgare
Sauvageaugloia grifjithsiana
Scytosiphon dotyi
Scytosiphon lomentaria
Scytosiphon simplicissimus
Spatoglossum schroederi
Sparoglossum solieri
Spatoglossum variabile
Spermatochnus paradoxus
Spermatochnus paradoxus var.
adriaticus
Sphacelaria brachygonia
Sphacelaria britannica
Sphacelaria cirrosa
Sphacelaria cirrosa var. hystrix
Sphacelaria furcigera
Sphacelaria fusca
Sphacelaria hystrix
Sphacelaria nana
Sphacelaria plumosa
Sphacelaria plumula
Sphacelaria rigidula
Sphacelaria saxatilis
Sphacelaria tribuloides
Sphacella subtilissima
Sphaerotrickia divaricata
Spongonema tomenlosum
Sporochnus dichotomus
Sporochnus gaerrneri
Sporochnus pedunculatus
Stictyosiphon adriaticus
Stictyosipl~onsorijerus
Stictyosiphon tortilis
Stilophora rhizodes
Stilophora rhizodes var. adriatica
Stiloplrora tuberculosa
Stilopsis lejolisii
Stragularia clavata
Streblonema eflusum
Streblonema infestans
Streblonema oligosporum
Streblonema oarasiticum
Streblonema sphaericum
Streblonema stilophorae
S~reblonematenrrissimurn
Streblonema rhrtretii
Streblonemopsis irritans
Strepsithalia liagorae
Strepsilhalia liebmanniae
Striaria allenuala f. ullertuata
Striaria attenuala f. crinila
Striaria atlenuala f. rarnosissima
Stypocaulon scoparium
Stypopodium schimperi
Stypopodium tubruquense
Stypopodium zonale
Symphoricoccus stellaris
Taonia atomaria f. atomaria
Taonia atomaria f. ciliala
Taonia atomaria var. ciliata
Undaria pinnatgida
Zanardinia prototypus
Zonaria tournefortii
Zosterocarpus oedogonium
Botanica Marina / Vol. 35 / 1992 / Fasc. 2
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