109 Ribera et al.: Checklist of Mediterranean seaweeds I. Fucophyceae Botanica Marina Vol. 35, pp. 109- 130, 1992 Offprint Check-list of Mediterranean Seaweeds I. Fucophyceae (Warming, 1884) M. A. Ribera, A. Gomez Garreta, T. Gallardo*, M. Cormaci*", G. Furnari** and G. Giaccone** Laboratorio de Bothnica, Facultad de Farmacia Universidad de Barcelona 08028 Bar-celom, Spain * Departamerlto de Biologia Vegetal, I Facultad de Biologia Universidad Conzplulense 08040 Madrid, Spain ** Istituto e Orto Botanico dell'Univer-sita, Via A. Longo 19 95125 Catania, Italy (Accepted 18 January 1992) Abstract A check-list of.Mediterranean brown algae, based on literature records, is given. The distribution of each taxon in the area (which is divided into 16 regions) is reported. The number of taxa accepted for the Mediterranean Sea under current taxonomy is 265. This list has benefited from the suggestions on taxonomy, nomenclature and regional distribution made by algological advisers from each region. Introduction This check-list of Mediterranean seaweeds is intended to be a catalogue of benthic algal taxa of the Mediterranean Sea, including only the Rhodophyceae, Fucophyceae (Phaeophyceae) and Chlorophyceae. The project emerged at the XXXth Congress and Plenary Assembly of International Commission for the Scientific Exploration of the Mediterranean Sea in Palma de Mallorca (Spain) in 1986, and has been coordinated by a group of Spanish algologists. The geographical area covered by this check-list is the Mediterranean Sea, including both the Sea of Marmara and the Black Sea. The area is bounded on the west by the Straits of Gibraltar. The Mediterranean coasts have been divided in a similar way to that used for the Med-Checklist of Vascular Plants (Greuter et al., 1984). However, some modifications have been made to account for seaweed biogeography and historical treatment of seaweed floras. For example, the Adriatic Sea is considered as a single region because of its biogeographical unity, while Malta and Sicily are treated together, as they traditionally have been treated when dealing with seaweed floras. As a result, Botanica Marina / Vol. 35 / 1992 / Fasc. 2 Copyright 0 1992 Walter de Gruyter . Berlin . New York the area has been split into 16 regions (Fig. 1) that are labelled with two-letter symbols in the following manner: (Sp) Spain, (BI) Balearic Islands, (Fr) France, (CS) Corsica and Sardinia, (WI) Western Italy, (Si) Sicily and adjacent islands, (Ad) Adriatic Sea (including Albania), (Gr) Greece, (BS) Black and Azov seas, (TU) Turkey (Sea of Marmara and Mediterranean coast), (LS) Levant States (Syria, Lebanon and Israel), (Eg) Egypt, (Li) Libya, (Tn) Tunisia, (Ag) Algeria, (MO) Morocco. This first part comprises the Fucophyceae (Phaeophyceae). The number of specific and infraspecific taxa accepted for the Mediterranean Sea under current taxonomy is 265. This work benefited from the suggestions on taxonomy, nomenclature and regional distribution of an advisory board of algologists. The advisers were: V. Aysel (Tu), A. Bavaru (BS), A. Bologa (BS), C. F. Boudouresque (Fr, CS, Tn), F. Cinelli (WI), S. Cirik (Tu), A. Cossu (CS), A. Diapoulis (Gr), V. Gazale (CS), S. Haritonidis (Gr), I. Munda (Ad) and G. Pardi (WI). 110 Ribera ef al.: Checklist of Mediterranean seaweeds I. Fucophyceae Fig. 1. Geographical regions considered: (Sp) Spain, (BI) Balearic Islands, (Fr) France, (CS) Corsica and Sardinia, (WI) Western Italy, (Si) Sicily and adjacent islands, (Ad) Adriatic Sea (including Albania). (Gr) Grecce, (BS) Black and Azov seas, (TU) Turkey (Sea of Marmara and Mediterranean coast), (LS) Levant States (Syria, Lebanon and Israel), (Eg) Egypt, (Li) Libya, (Tn) Tunisia, (Ag) AIgeria, (MO) Morocco. Knowledge of Mediterranean seaweeds had increased in the last few years to the point where it is useful, if not essential, to assemble this information in one publication. Our primary objective was to take the available pool of information and produce a single list and geographic index for the Mediterranean Sea. However, this list is not a definitive catalogue, but rather it is permanently open for additions and corrections. Another objective was to standardize the nomenclature as far as possible, reconciling the numerous taxonomic and nomenclatural divergences between the different local floras. In our opinion, recent taxonomic research has demonstrated clearly the need for resolving problems on a global rather than a local basis. This check-list has particular biogeographical interest because the distribution of each species is detailed for the different zones of the Mediterranean Sea. This list is based upon literature records. To minimize the citation of synonyms and misapplied names, only references published since 1950 have generally been used. For Egypt, however, we have included works by Nasr (1940) and Nasr and Aleem (1949) because contemporary studies for this area are rare. When available, regional floras have been used as the basis for the compilation. References used for the areas are shown in Tdble I. Taxa are named with their authors, but dates of valid publication are not given because of uncertainty in many cases. When authors' names have been abbreviated, TL2 (Stafleu and Cowan, 1976- 1988) was followed. Both in the checklist and in the index names of accepted taxa are in italics whereas names of synonyms, misapplied names and doubtful or unaccepted taxa are in roman type. The orders are arranged according to Wynne and Kraft (1981) while families, genera and species are arranged alphabetically. When a specific or infraspecific taxon has definitely been recorded for a geographical area, a nurnbcr corresponding to the most important bibliographic reference is cited in the list. Because of space limitations, only one reference is cited for each area, but additional references are available from the authors on request. Abbreviations of journals follow those of B. P. H. (Lawrence er al., 1968). When a specific or infraspecific taxon has been recorded under a synonym or misapplied name, the reference nurnbcr in the list is in italics and the synonym or misapplied name, is given in roman below the current namc. Superscript numbers refer to notes. A list of raxa excludenda as well as a list of raxa inquirenda is also given. Botanica Marina / VoI. 35 / 1992 / Fasc. 2 $ List of Taxa and Their Distribution g. Taxa 5. ECTOCARPALES Setchell et Gardner F L E! -< 0 r . . L \O \O N n Geographic regions Sp B1 Fr CS Ectocarpaceae C. Agardh Acinetospora Bornet A. crinita (Carmich. ex Harv.) Sauv. . . . . . . . . . . . . . . . . . . . . . . . . . . 8 = A. vidovichii (Menegh.) Sauv. = Haplospora vidovichii (Menegh.) Bornet Ectocarpus Lyngb. E. arabicus Fig. et De Not. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . E. caspicus Henckel . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . E. fmciculatus Harv. 8 var. fasciculatus . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . var. abbreviatus (Kiitz.) Sauv. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . = E. abbreviatus Kiitz. var. pycnocarpus (Rosenvinge) Cardinal . . . . . . . . . . . . . . . . . . . . . . E. parvus (Saunders) Hollenberg . . . . . . . . . . . . . . . . . . . . . . . . . . . . . E. rhodochortonoides Boerg. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . E. siliculosus (Dillwyn) Lyngb. = E. confe~oidesLe Jolis var. siliculosus . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 var. adriaticus (ErcegoviC) Cormaci et Furnari . . . . . . . . . . . . . . . . . . var. arctus (Kiitz.) Gallardo4' . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 105 = E. arctus Kiitz. var. crassus (Kjellm.) Gallardo4' . . . . . . . . . . . . . . . . . . . . . . . . . . . . var. crouanii (Thur.) Gallardo4' . . . . . . . . . . . . . . . . . . . . . . . . . . . . = E. crouanii Thur. var. dasycarpus (Kuck.) Gallardo4' . . . . . . . . . . . . . . . . . . . . . . . . . . = E. dasycarpus Kuck. var. hiernalis (Crouan frat. ex Kjellm.) Gallardo4' . . . . . . . . . . . . . . . . = E. hiemalis (Crouan frat. ex Kjellrn.) Kjellm. var. penicillatus C. Agardh . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 105 = E. penicillatus (C. Agardh) KjelIm. var. pygmaeus (Areschoug) Gallardo4' . . . . . . . . . . . . . . . . . . . . . . . . 17 var. subulatus (Kiitz.) Gallardo4' . . . . . . . . . . . . . . . . . . . . . . . . . . . var. venetus (Kiitz.) gal lard^^^ . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Feldrnanrlia Hamel F. battersiides (Ercegovic) Cormaci et Furnari . . . . . . . . . . . . . . . . . . . . F. caespitula ( J . Agardh) Knoep.-Pkg. 8 var. caespitula3 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . var. Iebelii (Areschoug ex Crouan frat.) Knoep.-P&. . . . . . . . . . . . . . 8 = F. lebelii (Areschoug ex Crouan frat.) Hamel 8 F. globifera (Kiitz.) Hamel . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . F. irregularis (Kiitz.) Hamel . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 E padinae (Buffham) Hamel . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 105 8 F. paradoxa (Mont.) Hamel . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 90 22 - - - - 90 22 - 21 - - - - 90 - 90 - - 22 - - 21 - - - - - - - - - - - - - - - - 34 - - - - 90 - 22 22 21 - 22 22 - 21 21 114 - - 90 90 - 90 - Taxa Geographic regions Sp B1 Fr CS E paradoxoides (ErcegoviC) Corrnaci et Furnari . . . . . . . . . . . . . . . . . . E simplex (Crouan frat.) Hamel . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 9 Herponema J. Agardh H. grangerum Kuck. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . H. solitariunz (Sauv.) Hamel . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 H. valiantei (Born.) Hamel . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 = Ectocarpus valiantei Born. H. velutinum (Grev.) J. Agardh . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . = Ectocarpus simpliciusculus C. Agardh Hincksia J. E. Gray H. dalmatica (ErcegoviC) Cormaci et Furnari4* . . . . . . . . . . . . . . . . . . . = Giffordia dalmatica (ErcegoviC) Cormaci et Furnari H. fuscara (Zanardini) Silva . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . = Giffordia fuscata (Zanardini) Kuck. H. geniculata (ErcegoviC) Cormaci et F ~ r n a r i. ~. ~ ................. = Giffordia geniculata (ErcegoviC) Cormaci et Furnari H. gratzulosa (Sm.) Silva . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 = Giffordia granulosa (Sm.) Hamel H. hauckii (ErcegoviC) Cormaci et Furnari" . . . . . . . . . . . . . . . . . . . . . = Giffordia hauckii (ErcegoviC) Cormaci et Furnari H. hincksiae (Harv.) Silva . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . = Giffordia hincksiae (Harv.) Hamel H.rnitchelliae(Harv.)Silva . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 = Giffordia mitchelliae (Harv.) Hamel H.ovata(Kjellm.)Silva . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 = Giffordia ovata (Kjellm.) Kylin = G. intermedia (Rosenvinge) Lund H. sandriana (Zanardini) Silva . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 = Giffordia sandriana (Zanardini) Hamel H. secunda (Kiitz.) Silva . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 105 = Giffordia secunda (Kiitz.) Batters Kuckuckia Hamel K. kylinii Cardinal . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . K. spinosa (Kiitz.) Kuck. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 Kuetzingiella Kornmann K. battersii (Born.) Kornmann = Feldrnannia battersii (Born.) Hamel var. battersii . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . var. rnediterranea (Sauv.) G6mez et Kibera4' . . . . . . . . . . . . . . . . . . . 8 = Ectocarpus battersii var. rnediterraneus Sauv. p W1 Si Ad Gr - 52 - 55 - - - 5 - - 116 21 - - - - 22 21 - 55 43 - - - - 46 55 42 - - - 34 - - 52 - - 22 - - 55 52 - - - - - - 52 - - 22 - - 55 52 - - - 34 - - 52 - - - - - 55 71 - 90 22 21 50 55 52 5 7 116 34 - 55 - - 90 22 21 50 55 52 - - - 34 - 55 100 - 7 22 22 - - - 55 - - 21 - - 22 114 - - 52 - - - - - BS - - 5 - Tu - Eg Tn Ag MO - - - - - - 85 - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - LS - 84 - 84 S4 68 - P - - - 68 84 - - - "51 2 - 15 85 - - - - - - - 102 15 85 - - - - 85 - - - - - - - 85 - - - - P - Li 85' - - $ 5' . g . W \O \O N ?I N Phaeostroma Kuck. P. berfholdii Kuck. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Pilayella Bory P. littoralis (L.) Kjellm. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 27 Spongonema Kiitz. S. tornenfosunz(Huds.) Kiitz. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Streblonema Pringsheim S. effusunz Kylin . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . = Entonema effusum (Kylin) Kylin S. infestans (Gran) Batters . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . S. oligosporum Stromfelt . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . = Entonema oligosporum (Stromfelt) Kylin S. parasiticum (Sauv.) De Toni . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . = Entonema parasiticurn (Sauv.) Hamel S. sphaericum (DerbCs et Solier) Thur. . . . . . . . . . . . . . . . . . . . . . . . . . 8 S. stilophorae (Crouan frat.) De Toni . . . . . . . . . . . . . . . . . . . . . . . . . . 105 S. tenuissimurn Hauck . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . S. fhurefiiSauv. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Streblonemopsis Valiante S. irritans Valiante . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 p p p Ralfsiaceae Farlow Acrospongium Schiffner A. ralfsioides Schiffner . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 98 Mesospora Weber M. tnacrocarpa (J. Feldm.) Hartog . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 = M. mediterranea J. Feldm. Microspongium Reinke M. gelatinosum Reinke . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Nemoderma Schousboe ex Bornet N. tingitanum Schousboe ex Bornet . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 Pseudolithoderma Svedelius P. adriaticum (Hauck) Verlaque . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 = Lithoderma adriaticum Hauck P. exfensum (Crouan frat.) Lund . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Ralfsia Berkeley R. disciformis Crouan frat. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . R. expansa ( J . Agardh) J . Agardh . . . . . . . . . . . . . . . . . . . . . . . . . . . . 98 R.verrucosa(Areschoug)J.Agardh . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 Stragularia Stromfelt S. clavafa (Ham.) Hamel . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 28 = Ralfsia bornetii Kuck. = R. clavata (Ham.) Crouan frat. - - - - - - 52 - - - - - - - 55 - - - 84 68 - - - 85 - - - - - - - - - - - 15 85 - - - 68 - - 15 85 - 66 - - - - - 12 84 68 - 15 - - 68 - 90 22 21 - - - 21 - - 22 21 50 55 - - - 22 21 18 55 52 5 - - - - - 90 22 21 46 - - 2 1 - 5 - 5 6 - 55 55 5 - 52 - 6 - 5 5 - - - - - 85 - Taxa Geographic regions Sp B1 Fr CS W1 Si Ad Gr BS Tu 46 - 31" -'l 38 - - 52 - 12 6 - 52 - 66 - LS Eg Li Tn Ag MO 85 - CHORDARIALES Setchell et Gardner Chordariaceae Grev. Cladosiphon Kiitz. C. chordariaeformis Crouan frat. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . = Castagnea chordariaeformis (Crouan frat.) Thur. C. contortus (Thur.) Kylin . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . = Castagnea contorta Thur. C. cylindricus (Sauv.) Kylin . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 = Castagnea cylindrica Sauv. C. irregularis (Sauv.) Kylin . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 = Castagnea irregularis Sauv. C. mediterraneus Kiitz. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 = Castagnea mediterranea (Kiitz.) Hauck = C. fistulosa (Zanardini) DerbCs et Solier C. zosterae (J. Agardh) Kylin . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . = Castagnea zosterae (J. Agardh) Thur. Eudesme J. Agardh E. virescens (Carmich. ex Berk.) J. Agardh . . . . . . . . . . . . . . . . . . . . . . = Castagnea virescens (Carmich. ex Berk.) Thur. Gorztrania Sauv. G. lubrica Sauv. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Liebmannia J. Agardh L. leveillei J. Agardh . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 Mesogloia C. Agardh M . lanosa Crouan frat. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . M . vermiculata (Sm.) S. F. Gray . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 Sauvageaugloia Hamel ex Kylin S. griffithsiana (Grev. ex Harv.) Hamel ex Kylin . . . . . . . . . . . . . . . . . . 8 Sphaerotrichia Kylin S. divaricata (C. Agardh) Kylin . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Strepsithalia Bornet ex Sauv. S. liagorae Sauv. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 S. liebmanniae Miranda . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Corynophlaeaceae Oltmanns Corynophlaea Kiitz. C. Jaccida (C. Agardh) Kiitz. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 17 = Myriactis adriatica (J. Agardh) De Toni C. hamelii J. Feldm. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . C. umbellata (C. Agardh) Kiitz. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 Cylindrocarpus Crouan frat. C. microscopicus Crouan frat. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 - - 21 - 22 22 34 - 22 - - - 46 P 115 Ribera et al.: Checklist of Mediterranean seaweeds I. Fucophyceae I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I l 1 l 1 I l 1 I I I I I I I I I I I I I I I l I I I I I I l 1 l I I l I I I I I I I 1 I l I 1 I I - 1 1 1 - 1 Botanica Marina / Vol. 35 / 1992 / Fasc. 2 I 1 Taxa M . liechtensternii Hauck . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . = Phaeosphaerium liechtensternii (Hauck) De Toni M . orbiculare J. Agardh . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . = M. magnusii (Sauv.) Loiseaux = M. seriatum (Reinke) Kylin = Ascocyclus orbicularis (J. Agardh) Magnus M . strangulans Grev. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Protectocarpus Kuck. P. speciosus (Boerg.) Kornmann . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Zosterocarpus Bornet Z . oedogonium (Menegh.) Born. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Geographic regions Sp B1 Fr CS W1 Si Ad Gr BS Tu LS Eg 9 - 22 21 - - 52 5 - - - - 8 90 22 21 46 55 52 5 1214 - 68 2 8 90 22 21 46 55 52 5 12 - 68 - - - - - - - - - 38 - - - 8 90 13 21 46 - 52 - - - - - - 22 21 46 55 52 5 - 6 - - - - - - - - 22 - 46 55 52 90 22 - 21 l5 55 52 - - - - Spermatochnaceae Kjellm. Nemacystus DerbCs et Solier N.jlexuosus (C. Agardh) Kylin . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 = N. rarnulosus Derbts et Solier N. hispanicus (Sauv.) Kylin . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Spermatochnus Kutz. S . paradoxus (Roth) Kutz. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Stilophora J. Agardh S. rhizodes (Turner) J. Agardh" . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 S. tuberculosa (Hornem.) Reinke . . . . . . . . . . . . . . . . . . . . . . . . . . . . . - - - - - - - - 5 65 59 39 - 5 12 59 - 38 - 68 - 73 - SPOROCHNALES Sauv Sporochnaceae Grev. Carpomitra Kiitz. C . costata (Stackh.) Batters var. costata . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . var. mediterranea J. Feldm. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . = Sporochnus dichotomus Zanardini Nereia Zanardini N.filiformis ( J . Agardh) Zanardini . . . . . . . . . . . . . . . . . . . . . . . . . . . Sporochnus C. Agardh S. gaertneri (Gmelin) C. Agardh . . . . . . . . . . . . . . . . . . . . . . . . . . . . . S. pedunculatus (Huds.) C. Agardh . . . . . . . . . . . . . . . . . . . . . . . . . . . Stilopsis Kuck. S. lejolisii (Thur.) Kuck. e x Hamel . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 22 21 52 52 - - - - - 50 55 55 - 90 - - 8 90 22 21 46 55 52 5 38 - 39 3 - - - - - 8 90 22 21 46 46 55 55 52 5 - - - - - - - - - 55 - - - - - - Desmarestiaceae (Thuret) Kjellm. Desmarestia Lamour. D . aculeata (L.) Lamour. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 27 D . adriatica ErcegoviC . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . - - - - - - - - 52 - - - - - - - p DESMARESTIALES Setchell et Gardner - Li Tn Ag MO D . dresnayi Lamour. ex Leman . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . D . ligulata Lamour. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . D . viridis 0. F. Miill." . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . - - - Arthrocladiaceae Chauvin Arthrocladia Duby A. villosa (Huds.) Duby" . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 90 Dictyosiphonaceae Kiitz. Dictyosiphon Grev. D . chordaria Areschoug . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . - - Giraudiaceae Hamel ex J. Feldm. Giraudia Derbts et Solier G . sphacelarioides Derbts et Solier . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 90 DICTYOSIPHONALES Setchell et Gardner Myriotrichiaceae Kjellm. Myriotrichia Harvey M. udriatica Ha'uck . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . M. clavaefornzis Harv. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . = M. repens Hauck = Dichosporangium repens (Hauck) Hauck Prutasperococcus Sauv. P. rnyriotrichiiformis Sauv. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . = Myriotrichia protasperococcus Kuck. Punctariaceae (Thuret) Kjellm. Asperococcus Lamour. A. bullosus Lamour. = A. turneri (Sm.) Hook. f. bullosus . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . f. profundus J. Feldm. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . A. compressus Griff. e x Hook. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . = Haloglossum compressum (Griff. ex Hook.) Hamel A. fistulosus (Huds.) Hook. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . = A. echinatus (Mertens) Grev. A. scaber Kuck. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Punctaria Grev. P. latifolia Grev. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . P. plantaginea (Roth) Grev. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . P. tenuissinia (C. Agardh) Grev. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . = Desmotrichum undulatum (J. Agardh) Reinke 8 8 8 90 - - 90 - 8 - 8 - - - - - Striariaceae Kjellm. Stictyosiphon Kiitz. S. adriuticus Kiitz. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . S. suriferus (Reinke) Rosenvinge . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 S. tortilis (Rupr.) Reinke9 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . - - 7 p p 112 - Taxa Geographic regions Sp B1 Fr CS Striaria Grev. S. attenuata (Grev.) Grev. f.attenuata . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . f. crinita (J. Agardh) Hauck. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . f. ramosissima (Kiitz.) Hauck. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . - 90 - W1 Si 11621 - 46 55 - 46 - p - - SCYTOSIPHONALES Feldm. Scytosiphonaceae Farlow Colpomenia (Endlicher) Derb6s et Solier C. peregrina (Sauv.) Hamel . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 C. sinuosa (Mertens ex Roth) Derbbs et Solier . . . . . . . . . . . . . . . . . . . 8 Hydroclathrus Bory H. clathratus (Bory ex C. Agardh) Howe . . . . . . . . . . . . . . . . . . . . . . . 8 Petalonia DerbCs et Solier P. fascia (0. F. Miill.) Kuntze . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 P. zosterifolia (Reinke) Kuntze . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Rosenvingea Boerg. R. intricata (J. Agardh) Boerg. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Scy~osiphonC. Agardh S. dotyi Wynne9 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . S. simplicissimus (Clemente) Cremades2* 8 = S. lomentaria (Lyngb.) Link. TILOPTERIDALES Sauv. Tilopteridaceae Sauv. Climacosorzts Sauv. C. mediterraneus Sauv. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . - - CUTLERIALES Kjellm. Cutleriaceae Hauck Cutleria Grev. C. adspersa (Roth) De Not.23 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . C. chilosa (Falkenb.) Silva" . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . = C. monoica Ollivier C.mullifda(Sm.)Grev .25 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Zanardinia Nardo ex Crouan frat. Z. prototypus (Nardo) Nardo . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 8 9026 902' 22 21 22 21 46 46 55 8 90 22 21 46 55 8 90 22 21 24 55 8 90 30 21 46 - SPHACELARIALES Oltmanns Choristocarpaceae Kjellm. Choristocarpus Zanardini C. tenellus (Kiitz.) Zanardini . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Ad Gr BS Tu LS Eg Li Tn Ag MO Ribera et al.: Checklist of Mediterranean seaweeds I. Fucophyceae IV) V ) , Botanica Marina / Vol. 35 / 1992 / Fasc. 2 mr- m m 119 Taxa Geographic regions Sp B1 Fr CS W1 Si Ad Gr - - BS D. repens (J. Agardh) J . Agardh . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 = D. fasciola var. repens (J. Agardh) J. Fcldm. D. spiralis (Mont.) Hamel . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 = D. ligulatus (Kutz.) J. Feldm. Lobophora J. Agardh L. variegata (Lamour.) Womersley . . . . . . . . . . . . . . . . . . . . . . . . . . . . 104 = Pocockiella variegata (Lamour.) Papenfuss Padina Adanson P. boergesenii Allendcr et Kraft . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . = P. gymnospora (Kiitz.) Vickers P. pavonica (L.) Lamour. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 P. tenuis Bory . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Spatoglossurn Kutz. S. schroederi (C. Agardh) Kutz. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . S. solieri (Chauv. ex Mont.) Kiitz. . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 S. variabile Fig. et De Not. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Stypopodiunl (Kiitz.) J . Agardh S. schitperi (Buchinger ex Kiitz.) Verlaque et Boudour. . . . . . . . . . . . . . = S. tubruquense Nizamuddin et Godeh Taonia J. Agardh 7: atomaria (Woodw.) J . Agardh f. atomaria . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 f. ciliata (Kutz.) Nizamuddin . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . = var. ciliata Kiitz. Zonaria Drapanald Z. tournefortii (Lamour.) Mont. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 27 - LAMINARIALES Kylin Alariaceae Setchell et Gardner Undaria Suringar emend. Okamura U. pinnatiJda (Harv.) S ~ r i n g a r ' ~. . . . . . . . . . . . . . . . . . . . . . . . . . . . . 14 - Chordaceae Dumortier Chorda Stackh. C . Jlum (L.) Stackh." . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 14 - 14 - 13 21 86 55 55 52 - - - 46 55 - 5 - Laminariaceae Bory Laminaria Lamour. L . japonica Areschoug3' . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . L . ochroleuca Pylaie . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . L.rodrigueziiBorn . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Saccorhiza Pylaie S. polyschides (Lightf.) Batters . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . = S. bulbosa (Huds.) Pylaie - 27 - 8 90 - - - - Tu LS Eg Li Tn Ag MO g 0 2. 51. 2 . . . Phyllariaceae Tilden Phyllariopsis Henry et South P. brvvipes (C. Agardh) Henry et South . . . . . . . . . . . . . . . . . . . . . . . = Phyllaria renifonnis (Lamour.) Rost. P. purpurascens (C. Agardh) Henry et South . . . . . . . . . . . . . . . . . . . . = Phyllaria purpurascens (C. Agardh) Rost. FUCALES Kylin ? h, -r E 0 h, Cystoseiraceae Kiitz. Cystoseira C. Agardh C . abies-marina (S. G. Gmelin) C. Agardh" . . . . . . . . . . . . . . . . . . . . . C . algeriensis J . Feldm. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . C . amentacea Bory = C. stricta Sauv. v. amentacea (Bory) Giaccone var. amentacea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . var. spicata (Ercegovii:) G i a c ~ o n e. ~. .~. . . . . . . . . . . . . . . . . . . . . . . = C. spicata Ercegovii: = C. spicata subsp. elegans Ercegovii: = C. stricta Sauv. v. spicata (ErcegoviC) Giaccone var. stricta Montagne . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . = C. stricta Sauv. v. stricta C . barbata C. Agardh var. barbata . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . f. aurantia (Kiitz.) Giaccone . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . = f. repens Zinova et Kalugina f. j7accida (Kiitz.) Woronich. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . f. insularum ErcegoviC . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . var. tophuloidea (ErcegoviC) Giaccone . . . . . . . . . . . . . . . . . . . . . . . . = subsp. tophuloidea Ercegovii: C . barbatula Kiitz.16 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . = C. graeca Schiffn. ex Gcrloff et Nizamuddin C . brachycarpa J. Agardh emend. Giaccone var. brachycarpa . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . var. balearica (Sauv.) Giaccone . . . . . . . . . . . . . . . . . . . . . . . . . . . . = C. balearica Sauv. v. balearica = C. caespitosa var. claudiae (Giaccone) Giaccone . . . . . . . . . . . . . . . . . . . . . . . . . . = C. balearica Sauv. v. claudiae Giaccone C . compressa (Esper) Gerloff et Nizamuddin = C. fimbriata (Desf.) Bory f.compressa . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . f. plana (ErcegoviC) Cormaci et al.'? . . . . . . . . . . . . . . . . . . . . . . . . = subsp. plana ErcegoviC f. rosetta (ErcegoviC) Cormaci et = subsp. rosetta Ercegovii: aLS6 . ....................... Taxa C. corniculata (Wulfen) Z a n a r d i ~38~ i.~. ~. . . . . . . . . . . . . . . . . . . . . . . . C. crinita (Desf.) Bory f. crinita . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . f. bosphorica (Sauv.) Zinova et Kalugina . . . . . . . . . . . . . . . . . . . . . . C. crinitopliylla ErcegoviC . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . C. dubia Valiante . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . = C. fucoides ErcegoviC C. elegans Sauv. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . C. foeniculacea (L.) Grev. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . C. humilis Kiitz. vachumilis . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . = C. compressa var. pustulata Ercegovii: var. myriophylloides (Saw.) Price et John39 . . . . . . . . . . . . . . . . . . . . = C. myriophylloides Sauv. C. hyblaea Giaccone . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . C . jabukae ErcegoviC f. jabukae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . f. tenuissima (Ercegovik) Cormaci et al. . . . . . . . . . . . . . . . . . . . . . . . = subsp. tenuissima ErcegoviC C. mauritanica S a ~ v . ~.' . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . C. mediterranea Sauv. var. mediterranea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . var. valiautei Sauv. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . C . nodicattlis (With.) Roberts . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . C. pelagosae ErcegoviC . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . C . sauvageauana Hamel . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . = C. selaginoides Valiante C . schiffneri Hamel = C. discors (L.) C. Agardh f. discors = C. discors (L.) C. Agardh f. dubia ErcegoviC = C. ercegovicii Giaccone f. ercegovicii = C. ercegovicii Giaccone f. schiffneri (Hamel) Giaccone f.schiffneri . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . f. latiramosa (Ercegovik) G i a c c ~ n e.~. ~. . . . . . . . . . . . . . . . . . . . . . . = C. ercegovicii f. latiramosa (ErcegoviC) Giaccone f. tenuiramosa (ErcegoviC) Giaccone4* . . . . . . . . . . . . . . . . . . . . . . . . = C. ercegovicii f. tenuiramosa (ErcegoviC) Giaccone C . sedoides (Desf.) C. Agardh . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . C. spinosa Sauv. var. spinosa4' . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . var. compressa (ErcegoviC) Cormaci et ~ 1 . 4.~. . . . . . . . . . . . . . . . . . . = C. adriatica var. compressa (ErcegoviC) Giaccone = C. platyramosa ErcegoviC var. tenuior (ErcegoviC) Cormaci et al." . . . . . . . . . . . . . . . . . . . . . . = C. adriatica f. tenuior (ErcegoviC) Giaccone = C. gerlofii Nizamuddin = C. rechingeri Schiffn. ex Gerloff et Nizamuddin Geographic regions Sp B1 Fr CS 8 - 8 27 56 - - 51 8 27 8 8 8 8 Ribera et al.: Checklist of Mediterranean seaweeds I. Fucophyc Notes ; ; ;jZl Botanica Marina / Vol. 35 / 1992 / Fasc. 2 ;l 1. Giaccone (1985: 697) cites both Acinetospora crinita and A. vidovichii, which are synonyms. 2. Giaccone (1978: 80, 89) cites both Acinetospora crinita and Haplospora vidovichii, which are synonyms. 3. We follow Knoepffler-Pi-guy (1970) in considering the records of Feldmannia paradoxa (Mont.) Hamel sensu Hamel to be referred to Feldmannia caespitula (J. Agardh) Knoep.P6g. 4. According to Ben Maiz et al. (1987), the presence of this species in Tunisia needs to be confirmed. 5. Perret-Boudouresque and Seridi (1989: 57) include in their citation var. mediterranea. 6. According to Athanasiadis (1987), the presence of this species in Greece needs to be confirmed. 7. Athanasiadis (1987: 108) does not consider this species to belong to the Greek flora. 8. Although there is no post-1950 record from that area, Hauck's record has been taken into account because the Adriatic Sea is the type locality. 9. This species is believed to have been accidentally introduced from Japan into the Thau lagoon (France), where it thrives under favourable environmental conditions (Riouall, 1985: 86). 10. Athanasiadis (1987: 114) excludes this species from the Aegean Sea. 11. Athanasiadis (1987: 112) excludes this species from both Greece and the Adriatic Sea. 12. Giaccone (1978: 93, 94) records both this species and E. kuckuckiana Schiffner, but the latter is a synonym of the former. 13. Elachista neglecta Kuck. is possibly a synonym of E. intermedia Crouan frat., according to Feldmann (1937: 272). 14. Kalugina-Gutnik (1975: 227) records separately both Myrionema seriatum (Reinke) Kylin and Ascocyclus orbicularis (J. Agardh) Magn. 15. Hamel (1937: 185) pointed out that Mediterranean specimens belong to var. adriatica J. Agardh. 16. Perret-Boudouresque and Seridi (1989: 52) include in their citation var. mediterranea. 17. Verlaque (1981) suggests that D. adriatica Ercegovii: may be a deep-water form of D. viridis. 18. This species includes f. australis (Kiitz.) Hauck (Ad 92, BS 116) and f. tenuissima Woronich (BS 116). 19. Funk (1955: 41) records both this species and Dichosporangium repens (Hauck) Hauck, but the latter is a synonym of the former. 20. Giaccone (1978: 85, 86) records both this species and Myriotrichia protasperococcus Berthold, but the latter is a synonym of the former. 21. Athanasiadis (1987: 118) does not recognize this species either in Greece or in Turkey considering it a cold-water alga that occurs in the Mediterranean Sea only in Sicily. 22. Regarding this combination, see Cremades and PCrez-Cirera (1990: 492). 23. This species includes the sporophytic stage, Aglaozonia melanoidea Schousboe ex Bornet. 24. This specific name is based on the sporophytic stage, Aglaozonia chilosa Falkenberg. The species includes the gametophytic stage, Cutleria monoica Ollivier. 25. The species includes the sporophytic stage, Aglaozonia parvula (Grev.) Zanardini. 26. Sporophyte not recorded. 27. Gametophyte not recorded. 28. Prud'homme van Reine (1982: 232) suggests that S. hystrix is an ecad of S. cirrosa. Its geographical distribution is: Sp 8, B1 94, Fr 22, CS 34, W1 45, S1 55, Ad 52, Gr 47, Tu 26, LS 68, Tn 15. 29. Prud'homme van Reine (1982: 103) considers the three Mediterranean records doubtful. 124 Ribera et al.: Checklist of Mediterranean seaweeds I. Fucophyceae According to Hornig and Schnetter (1988: 284) the Mediterranean Dictyola would consist of only D. dichoroma, including two varieties, namely, var. dicholoma and var. intricata. The latter would include D. linearis (C. Agardh) Grev. and D. pusilla Lamour. The records of the taxon: D. linearis f. divaricala Kiitz., which was never described but is reported in some papers (see references: 47, 50, 52, 55), are referred to D. linearis. According to Boudouresque and Perret-Boudouresque (1987: 64), this record requires substantiation, since it may possibly involve misinterpretation or labelling error. This species was accidentally introduced into the Mediterranean Sea from Japan (Boudouresque, Gerbal and Knoepffler-PBguy, 1985: 364). The Thau lagoon apparently provides a boreal environment in the Mediterranean Sea for this species (Riouall, 1985: 86). Amico et al. (1985: 899) suggest that the presence of this species in the Mediterranean Sea needs to be confirmed. Giivcn and t)ztig (1971: 124) and Nizamuddin er al. (1979: 469) record both this species and C . hoppii C. Agardh, but the latter is a synonym of the former. For the synonymization of Cystoseira barbatula and C. graeca, see Cormaci et al. (1992). This species includes C. corniculata f. imperfecta ErcegoviC and C. corniculta f. clydonophila Cormaci et al. (Cormaci et al., 1992). C. cornicula1a subsp. Iaxior Ercegovik and C . corniculata subsp. divergens Ercegovii: are synonyms of C. corniculala (Wulfen) Zanardini (Cormaci et al., 1992). The presence of this variety in the Mcditerranean Sea needs to be confirmed. Athanasiadis (1987: 135) states that the Greek record might be an error. This species is included in C . gibraltarica (Sauv.) Dangeard by Roberts (1978: 402). This species includes C. adriatica Sauv. (Cormaci et al., 1992). This variety includes C. adriatica Sauv. var. intermedia (ErcegoviC) Giaccone (Cormaci et al., 1992). This variety includes C. adriatica Sauv. f. reducta (Ercegovik) Giaccone (Cormaci et al., 1992). Although Hamza (1987: 64) records this species [as C. ericoides (L.) C. Agardh] from Tunisia, we believe that his material may be referable to C. mediterranea. Formerly, C. ericoides had been recorded from Tunisia (Pottier, 1929), but Ben Maiz et al. (1987: 291) consider this record to apply to either C. medilerranea or C . stricta. This species includes var. axillaris (J. Agardh) Mangenot. Funk (1955: 53) cited this species from Naples on the basis of a dead specimen carried by a ship. This species deserves careful taxonomic study. Its complex taxonomy has not allowed us to consider its varieties here. At least var. vulgare is recorded everywhere in the Mediterranean Sea except LS (68), where only var. diversijolium Bory and var. megalophyllum (Mont.) Grun. have been recorded. Regarding this combination see Gallardo, T. 1992. Nomenclatural notes on some Mediterranean algae, I: Phaeophyceae. Taxon, 41: in press. According to Verlaque el Boudouresque (1991), Mediterranean records of Slypopodium zonale (Lamour.) Papenfuss should be referred to S. schimperi. Nomenclatural changes On the basis of a comparative study (unpublished), Cystoseira brachycarpa J . Agardh and C , balearica Sauv. are distinguishable only at varietal level. The following new combinations are proposed: Cystoseira brachycarpa J . Agardh emend. Giaccone var. balearica (Sauv.) Giaccone comb. et stat. nov. [Basionym: C. balearica Sauv. (1912: 396)l. Cystoseira brachycarpa J. Agardh emend. Giaccone var. claudiae (Giaccone) Giaccone comb. nov. [Basionym: C. balearica Sauv. var. claudiae Giaccone (in Amico et al., 1985: 906)l. Taxa excludenda Chilionema reptans (Crouan) Sauv.: (Gr 47). Athanasiadis (1987: 108) excludes this taxon from the Greek flora (as Ectocarpus reptans P. et H . Crouan). Cystoseira baccata (Gmel.) Silva: (Gr 62). Restricted to the Atlantic Ocean (Oliveras PIS and G6mez Garreta, 1989: 92). Cystoseira Junkii Schiffn, ex Gerloff et Nizamuddin: (W1 49). Giaccone (1985: 436) considers this species to be referable to C. mediterranea var, valiantei Sauv. Cystoseira planiramea Schiffn. ex Gerloff et Nizamuddin: (Ad 48, Gr 48). This species may refer to C . humilis Kutz. according to Giaccone (1985: 436). Athanasiadis (1987) considers C . planiramea to lie with the range of C . compressa (Esper) Gerloff et Nizamuddin. Cystoseira pycnoclada Schiffn. ex Gerloff et Nizamuddin: (Ad 49) = C. schzj'Jneri Hamel f. schff neri according to Giaccone (1985: 435). Cystoseira sauvageauana var. polyoedenzatis (Sauv.) Hamel: (Sp 27, Fr 99, W1 46, Si 55, Ag 85). According to Motta (1989: 325) this taxon does not differ from the type variety. Cystoseira sicula Schiffn. ex Gerloff et Nizamuddin: (Si 79). Giaccone (1985: 436) states that it is a juvenile form of C . sauvageauana Hamel. Dictyota major Taylor: (Si 55). Hornig et Schnetter (1988: 282) point out that the record of D. major from Sicily (Giaccone et al. 1985: 713) should be referred to D. dichotoma (Huds.) Lamour. Sphacelariaplumosa Lyngbye: [W1 46, as Chaetopteris plumosa (Lyngb.) Kiitz.]. Athanasiadis (1987: 123) excludes this species from the Greek flora since it is a boreal species. Prud'homme van Reine (1982: 116) states that its Mediterranean records must be erroneous. Stypopodium zonale (Lamour.) Papenfuss. According to Verlaque et Boudouresque (1991), Mediterranean records of this species should be referred to S. schimperi. Botanica Marina / Vol. 35 / 1992 / Fasc. 2 125 Ribera et al.: Checklist of Mediterranean seaweeds I. Fucophyceae Taxa inquirenda Table I. References used for the different regions. Adriogloia adriatica Ercegovik (Ad 52). Cystoseira epiphytica Schiffn. e x Gerloff et Nizamuddin (Ad 49). Giaccone (1985: 435) believes that this species is a juvenile form of C . humilis Kiitz. Cystoseira gibraltarica (Sauv.) Dangeard may be a synonym of C . mauritanica Sauv. (Sp 99, MO 76) according to Roberts (1978: 402). Cystoseira nzontagnei J . Agardh (B1 90, CS 21, Li 82, Tn 15, Ag 85). Cystoseira platyclada Sauv. (Sp 27, Si 55, Ag 85, MO 76). Dalmatogloia bryozoi Ercegovik (Ad 52). Desmotrichum punctarioides A. Zin. et Dim.-Kon. This species probably is a stage of some Punctaria species. Dictyopteris tripolitana Nizamuddin (B1 79, W1 79, Si 79, Ad 79, Gr 79, Li 79, Tn 79, Ag 79). Dilophus mediterraneus Schiffn. var. crassus Schiffn. (Ad 52). Ectocarpus congestus Crouan frat. (Ad 100). Ectocarpus humilis Kiitz. (W1 46). Ectocarpus siliculosus var. divergens Schiffn. (Ad 52). Ectocarpus siliculosus var. elongatus Schiffn. (Ad 52). Ectocarpus siliculosus var. megacarpus Schiffn. (Ad 52). Microsporangiunz kuckuckianum Schiffn. (Ad 52). Myriactula microscopica (Ercegovik) Ercegovik (Ad 52). Padinopsis adriatica Ercegovik (Ad 52). Spermatocht~usparadoxus var. adriaticus ErcegoviC (Ad 52). Sp: BI: Fr: CS: WI: Si: Ad: Gr: BS: Tu: LS: Eg: Li: Tn: 8 7 13 34 10 4 31 5 11 6 39 1 78 15 Ag: 85 MO: 76 9 35 14 21 18 33 41 16 12 26 68 2 79 20 17 56 22 114 23 55 42 37 38 58 77 3 80 61 27 28 79 90 29 30 51 94 79 97 99 111 112 113 24 79 43 47 65 59 89 73 81 79 40 45 46 25 56 98 9 9 1 0 4 1 0 5 106 50 54 79 52 71 72 79 62 109 110 66 101 116 84 108 92 100 103 74 75 82 102 95 79 88 86 96 99 88 Acknowledgements We sould like to thank Dr P. C. Silva for his nomenclatural observations and suggestions; Dr M. Alvarez and Mr R. Rycroft for improving the English of this manuscript; Dr F. Conde for his bibliographical support; Mr J. Rull, Mr J. Marquez and Mr J. 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Meiiez. 1979. A list of marine algae from Libya. Bot. Mar. 22: 465-476. 83. Oliveras Pla, M. A. and A. Gomez Garreta. 1989. Corologia del genero Cystoseira C. Agardh (Phaeophyceae, Fucales). Anales Jard. Bot. Madrid 46: 89-97. 84. Oztiirk, M. and H. Giiner. 1986. Taxonomy and distribution of Ectocarpales (Phaeophyta) members on Aegean and Mediterranean coasts of Turkey. DOGA TU Bio. D.C . 10: 459-472. 85. Perret-Boudouresque, M. and H. Seridi. 1989. Inventaire des algues marines benthiques d'Algtrie. GIS Posidonie publ., Marseille. pp. 1 - 11 7. 86. Pignatti, A. and L. Rizzi-Longo. 1971 -72. Raccolte di alghe bentoniche nelle acque dell'Arcipelago Toscano. Atti Ist. Veneto Sci., Lett. and Arti 130: 31 3 -327. 87. Pottier, J. 1929. Etude sur les possibilites d'utilisation des plantes marines tunisiennes pour la nourriture de bttail. Ann. Inst. octanogr. 6: 321 -362. 88. Prud'Homme van Reine, W. F. 1982. List oJ Localities. 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Alphabetical List of Species Acinetospora crinita Acinetospora vidovichii Acrospongium ralfsioides Adriogloia adriatica Aglaozonia chilosa Aglaozonia melanoidea Aglaozonia parvula Arthrocladia villosa f. australis Arthrocladia villosa f. tenuissima Arthrocladia villosa f. villosa Ascocyclus orbicularis Asperococcus brrllosus f. bullosus Asperococc~lsb~rllosusf. profundus Asperococcus compressus Asperococcus echinatus Asperococcus fistulosus Asperococcus scaber Aspetococcus turneri Carpomitra costata var. costata Carpomitra costata var. mediterranea Castagnea chordariaeformis Castagnea contorta Castagnea cylindrica Castagnea fistulosa Castagnea irregularis Castagnea mediterranea Castagnea virescens Castagnea zosterae Chaetopteris plumosa Chilionema hispanicum Chilionema reptans Chorda /ilum Choristocarpus tenellus Cladosiphon chordariaeformis Cladosiphon contortus Cladosiphon cylindricus Cladosiphon irregularis Cladosiphon mediterraneus Cladosiphon zosterae Cladostephus hirsutus Cladostephus spongiosus f. spongiosus Cladostephus spongiosus f. verticillatus Cladostephus verticillatus Climacosorus mediterraneus Colpomenia peregrina Colpomenia sinuosa Compsonema gracilis Compsonema minutum Corynophlaea jlaccida Corynophlaea hamelii Corynophlaea umbellata Cutleria adspersa Cutleria chilosa Cutleria monoica Cutleria multifida Cylindrocarpus berkeleyi Cylindrocarpus microscopicus Cystoseira abies-marina Cystoseira adriatica var. adriatica Cystoseira adriatica var. compressa Cystoseira adriatica var. intermedia Cystoseira adriatica f. reducta Cystoseira adriatica f. tenuior Cystoseira algeriensis Cystoseira arnentacea var. amentacea Cystoseira amerltacea var. spicata Cystoseira amentacea var. stricta Cystoseira baccata Cystoseira balearica var. balearica Cystoseira balearica var. claudiae Cystoseira barbata f. auranlia Cystoseira barbata var. barbata Cystoseira barbata f. jlaccida Cystoseira barbata f. insularum Cystoseira barbata f. repens Cystoseira barbata subsp. tophuloidea Cystoseira barbata var. tophuloidea Cystoseira barbatula Cystoseira brachycarpa var. balearica Cystoseira brachycarpa var. brachycarpa Cystoseira hrachycarpa var. claudiae Botanica Marina / Vol. 35 / 1992 / Fasc. 2 Ribera et al.: Checklist of Mediterranean seaweeds I. Fucophyceae Cystoseira caespitosa Cystoseira cornpressa f. compressa Cystoseira compressa f. plana Cystoseira compressa subsp. plana Cystoseira compressa f. rosetta Cystoseira compressa subsp. rosetta Cystoseira compressa var. pustulata Cvstoseira corniculata Cystoseira corniculata f. clydonophila Cvstoseira corniculata f. im~erfecta Cystoseira corniculata subsp. divergens Cvstoseira corniculata subs^. laxior Cystoseira crinita f. bosphorica Cystoseira crinita f. crinita Cystoseira crinitophylla Cystoseira discors f. discors Cystoseira discors f. dubia Cystoseira dubia Cystoseira elegans Cystoseira epiphytica ! Cystoseira ercegovicii f. ercegovicii Cystoseira ercegovicii f. latiramosa Cystoseira ercegovicii f. schiffneri Cystoseira ercegovicii f. tenuiramosa Cystoseira ericoides Cystoseira fimbriata Cystoseira foeniculacea Cystoseira fucoides Cystoseira funkii Cystoseira gerlofii Cystoseira gibraltarica Cystoseira graeca Cystoseira hoppii Cystoseira humilis var. humilis Cystoseira humilis var. myriophylloides Cystoseira hyblaea Cystoseira jabukae f. jabukae Cystoseira jabukae f. tenuissima Cystoseira jabukae subsp. tenuissima Cystoseira mauritanica Cystoseira mediterranea var. mediterranea Cystoseira mediterranea var. valiantei Cystoseira montagnei Cystoseira myriophylloides Cystoseira nodicaulis Cystoseira pelagosae Cystoseira planiramea Cystoseira platyclada Cystoseira platyramosa Cystoseira pycnoclada Cystoseira rechingeri Cystoseira sauvageauana Cystoseira sauvageauana var. polyoedematis Cystoseira schiffneri f. latiramosa Cystoseira schiffneri f. schiffneri Cystoseira schiffneri f. tenuiramosa Cystoseira sedoides Cystoseira selaginoides Cystoseira sicula Cystoseira spicata Cystoseira spicata subsp. elegans Cystoseira spinosa var. compressa Cystoseira spinosa var. spinosa Cystoseira spinosa var. squarrosa Cystoseira spinosa var. tenuior Cystoseira squarrosa Cystoseira stricta var. amentacea Cystoseira stricta var. spicata Cystoseira stricta var. stricta Cystoseira susanensis Cystoseira tamariscifolia Botanica Marina / Vol. 35 / 1992 / Fasc. 2 Cystoseira irsneoides Cystoseira zosteroides Dalrnatogloia bryozoi Desmarestia aculeata Desmarestia adriatica Desrnarestia dresnayi Desmarestia ligulata Desmarestia viridis Desmotrichum punctarioides Desmotrichum undulatum Dichosporangium repens Dictyopteris membranacea Dictyopteris polypodioides Dictyopteris tripolitana Dictyosiphon chordaria Dictyota dichotoma var. dichotoma Dictyota dichotoma var. implexa Dictyota dichotoma var. intricata Dictyota divaricata Dictyota linearis Dictyota major Dictyota pusilla Dilophus fasciola Dilophus fasciola var. repens Dilophus ligulatus Dilophus mediterraneus Dilophus mediterraneus var. crassus Dilophus repens Dilophus spiralis Discosporangium mesarthrocarpum Ectocarpus abbreviatus Ectocarpus arabicus Ectocarpus arctus Ectocar~usbattersii var. mediterraneus Ectocarpus caspicus Ectocar~usconfervoides Ectocarpus congestus Ectocar~uscrouanii Ectocarpus dasycarpus Ectocar~usfasciculatus var. abbreviatus ~ctocar>us>ascictrlatus var. fascilulatus Ectocarpus fasciculatus var. pycnocarpus Ectocarpus hiemalis Ectocarpus humilis Ectocarpus paradoxus Ectocarpus parvus Ectocarpus penicillatus Ectocarpus reptans Ectocarpus rhodochortonoides Ectocarpus siliculosus var. adriaticus Ectocarpus siliculosus var. arctus Ectocarpus siliculosus var. crassus Ectocarpus siliculosus var. crouanii Ectocarpus siliculosus var. dasycarpus Ectocarpus siliculosus var. divergens Ectocarpus siliculosus var. elongatus Ectocarpus siliculosus var. hiemalis Ectocarpus siliculosus var. megacarpus Ectocarpus siliculosus var. penicillatus Ectocarpus siliculosus var. pygmaeus Ectocarpus siliculosus var. siliculosus Ectocarpus siliculosus var. subulatus Ectocarpus siliculosus var. venetus Ectocarpus simpliciusculus Ectocarpus valiantei Elachista flaccida Elachista fucicola Elachista intermedia var. clavaeformis Elachista intermedia var. intermedia Elachista jabukae Elachista kuckuckiana Elachista negkcta Elachista scutulata 129 Elachista stellaris Entonema effusum Entonema oligosporum Entonema parasiticum Eudesme virescens Feldmannia battersii Feldmannia battersiides Felhannia caespitula var. caespitula Feldmannia caespitula var. lebelii Feldmannia globifera Feldmannia irregularis Feldmannia lebelii Feldmannia padinae Feldmannia paradoxa Feldmannia paradoxoides Felhannia simplex Fuczrs spiralis Fucus vesiculosus Fucus vesiculosus var. axillaris Fzicus virsoides Giffordia dalmatica Giffordia fuscata Giffordia geniculata Giffordia granulosa Giffordia hauckii Giffordia hincksiae Giffordia intermedia Giffordia mitchelliae Giffordia ovata Giffordia sandriana Giffordia secunda Giraudia sphacelarioides Gontrania lubrica Haloglossum compressum Halopteris filicina Halopteris scoparia Halothrix lumbricalis Haplospora vidovichii Hecatonema liagorae Hecatonema maculans Herponema graniferum Heroonema solitarium Herponema valiantei Herponema velutinum Hincksia dalmatica Hincksia fuscata Hincksia geniculata Hincksia granulosa Hincksia hauckii Hincksia hincksiae Hincksia mitchelliae Hincksia ovata Hincksia sandriana Hincksia secunda Hydroclathrus clathratus Kuckuckia kylinii Kuckuckia spinosa Kuetzingiella battersii var. battersii Kuetzingiella batt~rsiivar. rnediterranea Laminaria japonica Laminaria ochroleuca Laminaria rodriguezii Leathesia difformis Leathesia mucosa var. condensata Leathesia mucosa var. mucosa Leptonema neapolitanum Leptonematella fasciculata Leptonematella neapolitana Liebmannia leveillei Lithoderma adriaticum Lobophora variegata Mesogloia lanosa Mesogloia vermiculata 130 Mesospora macrocarpa Mesospora mediterranea ~ i c r o s ~ o n ~ gelatinosum ium Microsporangium kuckuckianum ~ ~ r i a c iadriatica is Myriactrda arabica Myriactula elongata Myriactula gracilis Myriactula microscopica Myriactula rigida Myriactula rivrrlariue Myriactula stellulata Myrionema balticum Myrionema conchicola Myrionema liechtensternii Myrionema magnusii Mvrionema orbiculrre Myrionema seriatum Myrionema strangulans Myriotrichia adriatica Myriotricl~iaclavaeformis Myriotrichia protasperococcus Myriotrichia repens Nemacystus Jlexuosus Nemacystus kispanicus Nemacystus ramulosus Nemoderma tingitanum Nereia Jilijbrmis Padina boergesenii Padina gymnospora Padina pavonica Padina tenuis Padinopsis adriatica Petalonia fbscia Petalonia zosterijolia Petrospongium berkeleyi Phaeosphaerium liechtensternii Phaeostroma bertholdii Phyllaria purpurascens Phyllaria reniformis Phyllariopsis brevipes Phyllariopsis purpurascens Pilayella littoralis Pocockiella variegata Ribera et al.: Checklist of Mediterranean seaweeds I. Fucophyceae Protasperococcus myriotrichiijbrmis Protectocarpus speciosus Pseudoli~hoderrnaadriaticum Pseudolithoderrna extensum Punctaria lalijolia Punctaria plantaginea Punctaria lenuissima Ralfsia bornetii Ralfsia clavata RaljXa discijorrnis Ra&ia expansa Ra!i.sia verrucosa Rosen vingea intricata Saccorhiza bulbosa Saccorhiza polyschides Sargassum acinarium Sargassum jlavijolium Sargassum hornschuchii Sargassum linifolium Sargassum mulicunr Sargassum salicifolium Sargassum trichocarpum Sargassum vulgare var. diversifolium Sargassum vulgare var. megalophyllum Sargassum vulgare v. vulgare Sauvageaugloia grifjithsiana Scytosiphon dotyi Scytosiphon lomentaria Scytosiphon simplicissimus Spatoglossum schroederi Sparoglossum solieri Spatoglossum variabile Spermatochnus paradoxus Spermatochnus paradoxus var. adriaticus Sphacelaria brachygonia Sphacelaria britannica Sphacelaria cirrosa Sphacelaria cirrosa var. hystrix Sphacelaria furcigera Sphacelaria fusca Sphacelaria hystrix Sphacelaria nana Sphacelaria plumosa Sphacelaria plumula Sphacelaria rigidula Sphacelaria saxatilis Sphacelaria tribuloides Sphacella subtilissima Sphaerotrickia divaricata Spongonema tomenlosum Sporochnus dichotomus Sporochnus gaerrneri Sporochnus pedunculatus Stictyosiphon adriaticus Stictyosipl~onsorijerus Stictyosiphon tortilis Stilophora rhizodes Stilophora rhizodes var. adriatica Stiloplrora tuberculosa Stilopsis lejolisii Stragularia clavata Streblonema eflusum Streblonema infestans Streblonema oligosporum Streblonema oarasiticum Streblonema sphaericum Streblonema stilophorae S~reblonematenrrissimurn Streblonema rhrtretii Streblonemopsis irritans Strepsithalia liagorae Strepsilhalia liebmanniae Striaria allenuala f. ullertuata Striaria attenuala f. crinila Striaria atlenuala f. rarnosissima Stypocaulon scoparium Stypopodium schimperi Stypopodium tubruquense Stypopodium zonale Symphoricoccus stellaris Taonia atomaria f. atomaria Taonia atomaria f. ciliala Taonia atomaria var. ciliata Undaria pinnatgida Zanardinia prototypus Zonaria tournefortii Zosterocarpus oedogonium Botanica Marina / Vol. 35 / 1992 / Fasc. 2 View publication stats