Neodiversity 3: 3-12
www.neodiversity.org
10th Mar 2008
NEW COMBINATIONS IN THE GENUS
CATTLEYA
LINDL. (ORCHIDACEAE)
C. VAN DEN BERG1,2
1Departamento
de Ciências Biológicas, Universidade Estadual de FeiraSantana,
de
BR-116 Km3, Feira
de Santana, 44031-460, Brazil. e-mail: vcassio@gmx.de
ABSTRACT
New combinations and names are proposed in Cattleya to accommodate esspeci
previously
assigned to Laelia and Sophronitis. These are needed in order to maintain
monophyly
the
of the genus in light of recent phylogenetic hypotheses. Comments on nomenclatural
problems are also provided where necessary.
Since van den Berg et al. (2000) Cattleya, as currently circumscribed, is
published sequence data of ITS nuclear paraphyletic to Sophronitis s.l. Therefore,
DNA, it became clear that the Brazilian even though Sophronitis s.l. (sensu van den
species of Laelia Lindl. did not belong withBerg et al. 2000) was not contradicted, its
the Mexican group of species that include maintenance would require the creation of
the type of the genus; they were related to new
a
genera for the various subgroups of
completely different group. The initial Cattleya. A more practical (and simpler)
solution proposed was to place them in solution is to lump all Sophronitis species
Sophronitis Lindl. (van den Berg & Chase with Cattleya and deal with these
2000, 2001). Other authors, however, groupings at the infrageneric level. In this
preferred to split these species into several way, future infrageneric changes would not
segregate genera (Chiron & Castro 2002).
require further binomial changes, providing
Recent DNA studies with nine plastid also nomenclatural stability for artificial
regions (van den Berg, in prep.) plus thehybrids of species in this alliance.
ITS dataset (van den Berg et al. 2000) Several species of the former Brazilian
indicate that even though Sophronitis s.l. is Laelia
and Sophronitis already had
always recovered using different molecular previous valid combinations available in
datasets, the relationships among this Cattleya and they shall be used. Recent or
group, Cattleya Lindl., and Brassavola critical synonyms are presented here with
R.Br. are different from those produced the new combinations and new names. A
with the ITS dataset alone. Brassavola is review of the infrageneric taxonomy in
placed outside the Cattleya-Sophronitis Cattleya and combinations for natural
clade and Sophronitis s.l. is embedded hybrids will be published in a separate
among species traditionally recognized as paper.
Cattleya.
Cattleya Lindl., Coll. Bot. t. 33. 1821.
_________________________________
Sophronitis Lindl., Bot. Reg. 14: t.
2CNPq Research Fellow PQ1D
1147. 1828.
Maelenia Dumort., Mem. Acad. Sc.
3
VAN DEN BERG - NEW COMBINATIONS IN CATTLEYA
Brux. 9: 13, t. 10. 1834.
Eunannos Porto & Brade, Arq. Inst.
Biol. Veg. 12: 206. 1933.
Hoffmannseggella H.G.Jones, Acta Bot.
Acad. Sci. Hung. 14: 69. 1968.
Dungsia Chiron & V.P.Castro, Richardiana
2: 11. 2002.
Hadrolaelia (Schltr.) Chiron & V.P.Castro,
Richardiana 2: 11. 2002.
Microlaelia (Schltr.) Chiron & V.P.Castro,
Richardiana 2: 11. 2002.
Cattleyella Van den Berg & M.W.Chase,
Boletim CAOB 52: 100. 2004.
Schluckebieria Braem, Richardiana 4: 49.
2004.
Chironiella Braem, Richardiana 6: 108.
2006.
Brasilaelia Campacci & Gutfreund, Col.
Orquid. Brasileiras 4(pré-anexo): 99. 2006.
Suppl. 3: 12. 1996.
Sophronitis alaorii (Brieger & Bicalho)
Van den Berg & M.W.Chase f. dietliana
(O.Gruss) Van den Berg & M.W.Chase,
Lindleyana 16: 110 (2001).
Cattleya alvaroana (F.E.L.Miranda) Van
den Berg,comb. nov.
Basionym: Laelia alvaroana F.E.L.Miranda,
Bradea 8: 124. 1999.
Sophronitis alvaroana (F.E.L.Miranda)
Van den Berg & M.W.Chase, Lindleyana
15: 115. 2000. Hoffmannseggella alvaroana
(F.E.L.Miranda) V.P.Castro & Chiron,
Richardiana 2: 21. 2002.
Cattleya angereri (Pabst) Van den Berg,
comb. nov.
Basionym:
Laelia
angereri
Pabst,
Orchid. Dig. 39: 153. 1975.
Cattleya acuensis (Fowlie) Van den Berg,
Sophronitis angereri (Pabst) Van den
comb. nov.
Berg & M.W.Chase, Lindleyana 15: 115.
Basionym: Sophronitis acuensis Fowlie, 2000. Hoffmannseggella angereri (Pabst)
Orchid Dig. 39:150. 1975.
V.P.Castro & Chiron, Richardiana 2: 21.
Hadrolaelia acuensis (Fowlie) Chiron & 2002.
V.P.Castro, Richardiana 2: 16. 2002.
Cattleya bicalhoi Van den Berg,
nom. nov.
Cattleya alagoensis (V.P.Castro & Chiron)
Basionym: Laelia dayanaRchb.f., Gard.
Van den Berg,comb. nov.
Chron. n. s., 6: 772. 1876.
Basionym: Sophronitis alagoensis V.P.Castro
Sophronitis dayana (Rchb.f.) Van den
& Chiron, Richardiana 3: 101. 2003.
Berg & M.W.Chase, Lindleyana 15: 116.
2000. Hadrolaelia dayana (Rchb.f.) Chiron
Cattleya alaorii (Brieger & Bicalho) Van & V.P.Castro, Richardiana 2: 20. 2002.
den Berg,comb. nov.
Cattleya × dayana Rolfe was previously
Basionym: Laelia alaorii Brieger & used for a natural hybrid, and therefore a
Bicalho, Bradea 2: 108. 1976.
new name is necessary when Laelia dayana
Sophronitis alaorii (Brieger & Bicalho) is transferred to Cattleya. The name honors
Van den Berg & M.W.Chase, Lindleyana H.D.Bicalho, who published a detailed
15: 115. 2000. Hadrolaelia alaorii (Brieger account about the differences of this
& Bicalho) Chiron & V.P.Castro, Richardiana species in relation to C. pumila Hook. and
2: 20. 2002.
C. praestans (Rchb.f.) Van den Berg.
Cattleya alaorii (Brieger & Bicalho) Van
den Berg f. dietliana (O.Gruss) Van den Cattleya blumenscheinii (Pabst) Van den
Berg, comb. nov.
Berg, comb. nov.
Basionym: Laelia alaorii Brieger &
Basionym: Laelia blumenscheinii Pabst,
Bicalho f. dietliana O.Gruss, Die Orchidee Bradea 1: 487.1975.
4
VAN DEN BERG - NEW COMBINATIONS IN CATTLEYA
Sophronitis blumenscheinii (Pabst) Van Cattleya caulescens (Lindl.) Van den Berg,
den Berg & M.W.Chase, Lindleyana 15: comb. nov.
116. 2000. Hoffmannseggella blumenscheinii
Basionym: Laelia caulescens Lindl.,
(Pabst) V.P.Castro & Chiron, Richardiana 2: Edward’s Bot. Reg. 27: sub t. 1. 1841.
21. 2002.
Hoffmannseggella caulescens (Lindl.)
H.G.Jones, Caldasia 10: 493. 1970.
Cattleya bradei (Pabst) Van den Berg, Sophronitis caulescens (Lindl.) Van den
comb. nov.
Berg & M.W.Chase, Lindleyana 15: 116.
Basionym: Laelia bradei Pabst, Bradea 2000.
1: 332. 1973.
Sophronitis bradei (Pabst) Van den Berg Cattleya cernua (Lindl.) Van den Berg,
& M.W.Chase, Lindleyana 15: 116. 2000. comb. nov.
Hoffmannseggella bradei (Pabst) V.P.Castro
Basionym: Sophronitis cernua Lindl.,
& Chiron, Richardiana 2: 21. 2002.
Bot. Reg. 14: sub t. 1147. 1828.
" Cattleya cernua" (Lindl.) Beer, Prakt
Orch. 209. 1854. nom. inval.
Cattleya brevicaulis (H.G.Jones) Van den
Sophronitis pterocarpa Lindl. & Paxton,
Berg, comb. nov.
Paxton’s Fl. Gard. 3: 11. 1853.
Basionym: Hoffmannseggella brevicaulis
The name " Cattleya cernua" (Lindl.)
H.G.Jones, Rhodora 74: 284. 1972.
Beer is listed by the International Plant
Sophronitis brevicaulis (H.G.Jones) Van Names Index (IPNI), but a careful
den Berg & M.W.Chase, Lindleyana 15: examination of the original publication
116. 2000. Dungsia brevicaulis (H.G.Jones) (Beer 1854) reveals it to be an invalid
Chiron & V.P.Castro, Richardiana 2: 19. name. The combination is simply a cross2002.
reference
to
the
treatment
under
Sophronitis, implying the name was not
Cattleya brevipedunculata (Cogn.) Van accepted by the author, which renders the
den Berg,comb. nov.
combination invalid under the International
Basionym: Sophronitis wittigiana Barb.Rodr. Code of Botanical Nomenclature (ICBN,
var. brevipedunculata Cogn. in Mart., Fl. McNeill et al. 2006), article 34.1, provision
bras. 3(5): 318. 1901.
a): “A name is not validly published when it
Sophronitis brevipedunculata (Cogn.) is not accepted by the author in the original
Fowlie, Orchid. Digest 36: 192. 1972. publication”. Zappi (1994) proposed to the
Hadrolaelia
brevipedunculata
(Cogn.) IAPT committee the conservation of
Chiron & V.P.Castro, Richardiana 2: 21. Cattleya violacea (Kunth) Rolfe against " C.
2002.
violacea" Beer, for the same reasons as in
" C. cernua" Beer. The committee rejected
this proposal on the grounds that " C.
Cattleya briegeri (Blumensch. ex Pabst) violacea" Beer was surely invalid and
Van den Berg,comb. nov.
conservation was unnecessary (Brummitt
Basionym: Laelia briegeri Blumensch. 1996). Sophronitis pterocarpa is a name
ex Pabst, Bradea 1: 367. 1973.
that has been applied to an allegedly
Sophronitis briegeri (Blumensch. ex Pabst) distinct taxon from Paraguay and western
Van den Berg & M.W.Chase,
Lindleyana 15: Brazil. However, examination of the type
116.
2000.
Hoffmannseggella
briegeri revealed it to be conspecific with C. cernua.
(Blumensch. ex Pabst) V.P.Castro & Chiron, The type specimen was also collected in
Richardiana 2: 22. 2002.
Rio de Janeiro, where only C. cernua
5
VAN DEN BERG - NEW COMBINATIONS IN CATTLEYA
occurs. If the taxon in Paraguay represents athese authors explicitly did not accept in
new taxonomic entity, it will require a other papers), if not all the combinations
different name.
proposed in that paper, should be
considered invalid.
Cattleya cinnabarina (Bateman ex Lindl.)
Van den Berg,comb. nov.
Cattleya conceicionensis (V.P.Castro &
Basionym: Laelia cinnabarina Batem. ex Campacci) Van den Berg,comb. nov.
Lindl., Sert. Orch. t. 28. 1839.
Basionym: Hoffmannseggella conceicionensis
" Cattleya
cinnabarina"
(Batem. ex V.P.Castro & Campacci, Richardiana 2:
Lindl.) Beer, Prakt. Orch. 209. 1854. nom. 113.
2002.
Sophronitis
conceicionensis
inval.
Hoffmannseggella
cinnabarina (V.P.Castro & Campacci) Van den Berg &
(Bateman ex Lindl.) H.G.Jones, Acta Bot. M.W.Chase, Kew Bull. 59: 566. 2005.
Acad. Sci. Hung. 14: 69. 1968. Sophronitis
cinnabarina (Bateman ex Lindl.) Van den Cattleya crispata (Thunb.) Van den Berg,
Berg & M.W.Chase, Lindleyana 15: 116. comb. nov.
2000.
Basionym: Cymbidium crispatum Thunb.,
The combination proposed by Beer Pl. Bras. 2: 18. 1818.
(1854) is invalid by the same reason Hoffmannseggella
crispata
(Thunb.)
explained under the comments on C. cernua. H.G.Jones, Hawaii Orchid J. 3: 15. 1974.
Therefore, a valid combination is proposed Sophronitis crispata (Thunb.) Van den Berg
here.
& M.W.Chase, Lindleyana 15: 116. 2000.
Laelia flava Lindl., Bot. Reg. 25: misc.
Cattleya coccinea Lindl. f. rossiteriana
88. 1839. " Cattleya flava" (Lindl.) Beer,
(Barb.Rodr.) Van den Berg,comb. nov.
Prakt. Stud. Orchid: 210. 1854. nom. inval.
Basionym:
Sophronitis
rossiteriana Hoffmannseggella flava (Lindl.) H.G.Jones,
Barb.Rodr., Gen. Sp. Orchid. 1: 77. 1877.
Caldasia 10: 494. 1970.
Cattleya colnagoi (Chiron & V.P.Castro) Cattleya dichroma Van den Berg,nom.
Van den Berg,comb. nov.
nov.
Basionym: Hoffmannseggella colnagoi
Basionym: Sophronitis bicolor F.E.L.
Chiron & V.P.Castro, Richardiana 5: 8. 2005. Miranda, Die Orchidee 42: 227. 1991.
" Sophronitis
colnagoi"
(Chiron
& Hadrolaelia
bicolor
(F.E.L.Miranda)
V.P.Castro) Campacci, Bol. CAOB 58: 51. Chiron & V.P.Castro, Richardiana 2: 20.
2005. nom. inval.
2002.
When Baptista et al. (2005) proposed 22 Because of the pre-existing Cattleya
new combinations (including " Sophronitis bicolor Lindl., a new name is necessary
colnagoi" ), they clearly stated in thewhen transferring Sophronitis bicolor to
introduction: “despite our belief that the Cattleya. We used a Greek version of the
name by us used is the correct one, wesame epithet for this purpose.
decided to make new combinations for
some of these species” (my translation). Cattleya duveenii (Fowlie) Van den Berg,
This implies that the authors did not acceptcomb. nov.
the names as they were proposing, which Basionym: Laelia duveenii Fowlie,
renders them invalid under Article 34.1 of Orchid Digest 52: 180. 1988.
the ICBN. For this reason, at least the Sophronitis duveenii (Fowlie) Van den
combinations
in
Sophronitis
and Berg & M.W.Chase, Lindleyana 15: 116.
Prosthechea Knowles & Westc. (which 2000. Hoffmannseggella duveenii (Fowlie)
6
VAN DEN BERG - NEW COMBINATIONS IN CATTLEYA
V.P.Castro & Chiron, Richardiana 2: 22. Cattleya fournieri (Cogn.) Van den Berg,
2002.
comb. nov.
Basionym: Laelia longipes Rchb.f. var.
Cattleya endsfeldzii (Pabst) Van den Berg, fournieri Cogn., Chron. Orchid. 1: 27. 1897.
comb. nov.
Laelia fournieri (Cogn.) F.E.L.Miranda,
Basionym: Laelia endsfeldzii Pabst, Bradea 6: 156. 1993. Sophronitis fournieri
Bradea 2: 51. 1975.
(Cogn.) Van den Berg & M.W.Chase,
Sophronitis endsfeldzii (Pabst) Van den Lindleyana 15: 117. 2000. Hoffmannseggella
Berg & M.W.Chase, Lindleyana 15: 116. fournieri (Cogn.) V.P.Castro & Chiron,
2000. Hoffmannseggella endsfeldzii (Pabst) Richardiana 2: 22. 2002.
V.P.Castro & Chiron, Richardiana 2: 22.
2002.
Cattleya ghillanyi (Pabst) Van den Berg,
comb. nov.
Cattleya esalqueana (Blumensch. ex Pabst)
Basionym:
Laelia
ghillanyi
Pabst,
Van den Berg,comb. nov.
Bradea 1: 332. 1973.
Basionym: Laelia esalqueana Blumensch.ex
Hoffmannseggella
ghyllanyi
(Pabst)
Pabst, Bradea 1: 367. 1973.
H.G.Jones, Hawaii Orchid J. 3: 16. 1974.
Sophronitis esalqueana (Blumensch. ex Sophronitis ghyllanyi (Pabst) Van den Berg
Pabst) Van den Berg & M.W.Chase, & M.W.Chase, Lindleyana 15: 117. 2000.
Lindleyana 15: 116. 2000. Hoffmannseggella
esalqueana (Blumensch. ex Pabst) V.P.Castro& Cattleya gloedeniana (Hoehne) Van den
Chiron, Richardiana 2: 22. 2002.
Berg, comb. nov.
Basionym: Laelia gloedeniana Hoehne,
Bol. Agric. (São Paulo) 34: 624. 1934.
Cattleya fidelensis (Pabst) Van den Berg,
Sophronitis gloedeniana (Hoehne) Van
comb. nov.
den Berg & M.W.Chase, Lindleyana 15:
Basionym:
Laelia
fidelensis
Pabst, 117. 2000. Hoffmannseggella gloedeniana
Orquídea (Rio de Janeiro) 29: 11, t.6. 1967. (Hoehne) V.P.Castro & Chiron, Richardiana
Sophronitis fidelensis (Pabst) Van den 2: 22. 2002.
Berg & M.W.Chase, Lindleyana 15: 117.
2000. Hadrolaelia fidelensis (Pabst) Chiron Cattleya gracilis (Pabst) Van den Berg,
& V.P.Castro, Richardiana 2: 17. 2002. comb. nov.
Chironiella
fidelensis
(Pabst)
Braem,
Basionym: Laelia gracilis Pabst, Bradea
Richardiana 6: 109. 2006. Brasilaelia 2: 314. 1979.
fidelensis
(Pabst)
Gutfreund,
Colet.
Sophronitis gracilis (Pabst) Van den
Orquídeas Brasil. 4(pré-anexo): 99. 2006.
Berg & M.W.Chase, Lindleyana 15: 117.
2000. Hoffmannseggella gracilis (Pabst)
V.P.Castro & Chiron, Richardiana 2: 21.
Cattleya flavasulina (F.E.L.Miranda & 2002.
K.G.Lacerda) Van den Berg,comb. nov.
Basionym: Hoffmannseggella flavasulina Cattleya harpophylla (Rchb.f.) Van den
F.E.L.Miranda & K.G.Lacerda, Orchids, Berg, comb. nov.
Mag. Amer. Orchid Soc. 72: 850. 2003.
Basionym: Laelia harpophylla Rchb.f.,
Sophronitis flavasulina (F.E.L.Miranda Gard. Chron. (1873): 542. 1873.
& K.G.Lacerda) Van den Berg &
Hoffmannseggella harpophylla (Rchb.f.)
M.W.Chase, Kew Bull. 56: 566. 2005.
H.G.Jones, Caldasia 10: 494. 1970.
Sophronitis harpophylla (Rchb.f.) Van den
7
VAN DEN BERG - NEW COMBINATIONS IN CATTLEYA
Berg & M.W.Chase, Lindleyana 15: 117. V.P.Castro & Chiron, Richardiana 2: 23.
2000. Dungsia harpophylla (Rchb.f) Chiron 2002.
& V.P.Castro, Richardiana 2: 11. 2002.
Laelia cardimii Pabst & A.F.Mello,
Bradea 2: 226. 1978. " Sophronitis cardimii"
Cattleya hatae (V.P.Castro & K.G.Lacerda) (Pabst & A.F.Mello) Campacci, Bol. CAOB
Van den Berg,comb. nov.
58: 51. 2005. nom. inval.
Basionym:
Hoffmannseggella
hatae
Although the name Laelia cardimii
V.P.Castro & K.G.Lacerda, Icon. Orchid. Pabst & A.F.Mello has been commonly
Bras. 2: t. 155. 2006.
used for a distinctive species under
cultivation (Miranda 1998), the type and
Cattleya hegeriana (Campacci) Van den description of this name is conspecific with
Berg, comb. nov.
Laelia itambana and therefore does not
Basionym: Hoffmannseggella hegeriana belong to the current concept being applied
Campacci, Colet. Orquídeas Brasil. 5: 158. to this species, which remains undescribed
2007.
to date. Apparently unaware of this fact,
Campacci in Baptista et al. (2005)
Cattleya hispidula (Pabst & A.F.Mello) published a new combination for it, which
Van den Berg,comb. nov.
was also invalid for the same reasons
Basionym: Laelia hispidula Pabst & explained under the comments of C.
A.F.Mello, Bradea 2: 227. 1978.
colnagoi.
Sophronitis hispidula (Pabst & A.F.Mello)
Van den Berg & M.W.Chase, Lindleyana Cattleya jongheana (Rchb.f.) Van den
15: 117. 2000. Hoffmannseggella hispidula Berg, comb .nov.
(Pabst & A.F.Mello) V.P.Castro & Chiron,
Basionym: Laelia jongheana Rchb.f.,
Richardiana 2: 22. 2002.
Gard. Chron. (1872): 425. 1872.
Sophronitis jongheana (Rchb.f.) Van den
Cattleya hoehneiVan den Berg,nom. nov.
Berg & M.W.Chase, Lindleyana 15: 117.
Basionym: Laelia mixta Hoehne, Arch. 2000. Hadrolaelia jongheana (Rchb.f.)
Bot. São Paulo 1: 20. 1938.
Chiron & V.P.Castro, Richardiana 2: 20.
Sophronitis mixta (Hoehne) Van den 2002.
Berg & M.W.Chase, Lindleyana 15: 118.
2000. Hoffmannseggella mixta (Hoehne) Cattleya
kautskyiana (V.P.Castro
&
V.P.Castro & Chiron, Richardiana 2: 23. Chiron) Van den Berg,comb. nov.
2002.
Basionym: Hoffmannseggella kautskyana
This nomen novum is necessary because V.P.Castro & Chiron, Richardiana 3: 65.
Cattleya ×mixta L.C.Men. was already taken
for 2003.
a natural hybrid. The name honors F.C. " Sophronitis kautskyana" (V.P.Castro &
Hoehne, who originally described this Chiron) Baptista, Bol. CAOB 58: 51. 2005.
species.
nom. inval.
Cattleya itambana (Pabst) Van den Berg,
comb. nov.
Basionym:
Laelia
itambana
Pabst,
Bradea 1: 333. 1973.
Sophronitis itambana (Pabst) Van den
Berg & M.W.Chase, Lindleyana 15: 117.
2000. Hoffmannseggella itambana (Pabst)
8
Cattleya kettieana (Pabst) Van den Berg,
comb. nov.
Basionym:
Laelia
kettieana
Pabst,
Bradea 2: 1. 1976.
Sophronitis kettieana (Pabst) Van den
Berg & M.W.Chase, Lindleyana 15: 117.
2000. Hoffmannseggella kettieana (Pabst)
VAN DEN BERG - NEW COMBINATIONS IN CATTLEYA
V.P.Castro & Chiron, Richardiana 2: 23. Warm.) Chiron & V.P.Castro, Richardiana
2002.
2: 11. 2002.
Cattleya liliputana (Pabst) Van den Berg, Cattleya lundii f. alba (L.C.Men.) Van den
comb. nov.
Berg, comb. nov.
Basionym: Laelia
liliputana
Pabst, Basionym: Laelia lundii var. alba L.C.Men.
,
Bradea 1: 334. 1973.
Bol. CAOB 33: 68. 1998.
Hoffmannseggella
liliputana
(Pabst)
H.G.Jones, Hawaii Orchid J. 3: 16. 1974. Cattleya macrobulbosa (Pabst) Van den
Sophronitis liliputana (Pabst) Van den Berg Berg, comb. nov.
& M.W.Chase, Lindleyana 15: 117. 2000.
Basionym: Laelia macrobulbosa Pabst,
Bradea 1: 335. 1973.
Cattleya longipes (Rchb.f.) Van den Berg,
Hoffmannsegella macrobulbosa (Pabst)
comb. nov.
H.G.Jones, Hawaii Orchid J. 3: 16. 1974.
Basionym: Laelia longipes Rchb.f., Sophronitis macrobulbosa (Pabst) Van den
Xenia Orchid. 2: 59. 1874.
Berg & M.W.Chase, Lindleyana 15: 117.
Sophronitis longipes (Rchb.f.) Van den 2000.
Berg & M.W.Chase, Lindleyana 15: 117.
2000. Hoffmannseggella longipes (Rchb.f.)
V.P.Castro & Chiron, Richardiana 2: 23. Cattleya mantiqueirae (Fowlie) Van den
Berg, comb. nov.
2002.
Basionym: Sophronitis coccinea subsp.
mantiqueirae Fowlie, Orchid. Digest 32:
Cattleya luetzelburgii Van den Berg,nom.
273. 1968.
nov.
Sophronitis mantiqueirae (Fowlie) Fowlie,
Basionym: Laelia bahiensis Schltr.,
Repert. Spec. Nov. Regni Veg. 17: 272. Orchid. Dig. 36: 183. 1972. Hadrolaelia
mantiqueirae (Fowlie) Chiron & V.P.Castro,
1921.
Hoffmannseggella
bahiensis (Schltr.) Richardiana 2: 21. 2002.
H.G.Jones, Novosti Sist. Vyssh. Rast. 12:
141. 1975. Sophronitis bahiensis (Schltr.) Cattleya marcaliana (Campacci & Chiron)
Van den Berg & M.W.Chase, Lindleyana Van den Berg,comb. nov.
Basionym: Dungsia marcaliana Campacci
15: 116. 2000.
A nomen novum is necessary because the & Chiron, Richardiana 2: 77. 2002.
Sophronitis marcaliana (Campacci &
name Cattleya × bahiensis Rolfe is already
taken for a natural hybrid. We honor Chiron) Van den Berg & M.W.Chase, Kew
Luetzelburg, who was the first to collect Bull. 56: 566. 2005.
this species in Bahia.
Cattleya milleri (Blumensch. ex Pabst)
Cattleya lundii (Rchb.f. & Warm.) Van den Van den Berg,comb. nov.
Basionym: Laelia milleri Blumensch. ex
Berg, comb. nov.
Basionym: Bletia lundii Rchb.f. & Pabst, Bradea 1: 367. 1973.
Sophronitis milleri (Blumensch. ex
Warm., Otia Bot. Hamburg. 2: 92. 1881.
Laelia lundii (Rchb.f. & Warm.) Rchb.f., Pabst) Van den Berg & M.W.Chase,
Otia
Bot.
Hamburg.
2:
92.
1881. Lindleyana 15: 118. 2000. Hoffmannseggella
Sophronitis lundii (Rchb.f. & Warm.) Van milleri (Blumensch. ex Pabst) V.P.Castro &
den Berg & M.W.Chase, Lindleyana 15: Chiron, Richardiana 2: 23. 2002.
117. 2000. Microlaelia lundii (Rchb.f. &
9
VAN DEN BERG - NEW COMBINATIONS IN CATTLEYA
Cattleya
mirandae (K.G.Lacerda
&
V.P.Castro) Van den Berg,comb. nov.
Basionym: Hoffmannseggella mirandae
K.G.Lacerda & V.P.Castro, Richardiana
5(1): 16. 2005.
" Sophronitis
neomirandae"
Baptista,
Bol. CAOB 58: 51. 2005. nom. inval.
K.G.Lacerda) Van den Berg & M.W.Chase,
Kew Bull. 59: 566. 2005.
Cattleya pfisteri (Pabst & Senghas) Van
den Berg,comb. nov.
Basionym: Laelia pfisteri Pabst &
Senghas, Orchidee 26: 254. 1975.
Sophronitis pfisteri (Pabst & Senghas)
Van den Berg & M.W.Chase, Lindleyana
15: 118. 2000. Hoffmannseggella pfisteri
(Pabst & Senghas) V.P.Castro & Chiron,
Richardiana 2: 23. 2002.
Cattleya munchowiana (F.E.L.Miranda)
Van den Berg,comb. nov.
Basionym:
Laelia
munchowiana
F.E.L.Miranda, Bradea 8: 126. 1999.
Sophronitis munchowiana (F.E.L.Miranda)
Van den Berg & M.W.Chase, Lindleyana Cattleya praestans (Rchb.f.) Van den
15: 118. 2000. Hoffmannseggella munchowiana Berg, comb. nov.
Basionym: Laelia praestans Rchb.f.,
(F.E.L.Miranda) V.P.Castro & Chiron,
Allg. Gartenzeitung 25: 336. 1857.
Richardiana 2: 23. 2002.
Sophronitis praestans (Rchb.f.) Van den
Berg & M.W.Chase, Lindleyana 15: 118.
Cattleya neokautskyi Van den Berg,nom.
2000. Hadrolaelia praestans (Rchb.f.)
nov.
Basionym:
Laelia
kautskyi
Pabst, Chiron & V.P.Castro, Richardiana 2: 20.
2002.
Orchid. Rev. 78: 321. 1970.
Hoffmannseggella
kautskyi
(Pabst)
H.G.Jones, Rhodora 74: 285. 1972. Cattleya presidentensis (Campacci) Van
Sophronitis kautskyi (Pabst) Van den Berg den Berg,comb. nov.
Basionym: Hoffmannseggella presidentensis
& M.W.Chase, Lindleyana 15: 117. 2000.
Dungsia
kautskyi (Pabst) Chiron & Campacci, Bol. CAOB 57: 28. 2005.
" Sophronitis presidentensis" (Campacci)
V.P.Castro, Richardiana 2: 19. 2002.
Cattleya × kaustkyi Pabst was described Campacci, Bol. CAOB 58: 51. 2005. nom.
for a natural hybrid, and therefore a newinval.
name is necessary for this species.
Cattleya purpurata (Lindl. & Paxton) Van
Cattleya
pabstii (F.E.L.Miranda
& den Berg,comb. nov.
Basionym: Laelia purpurata Lindl. &
K.G.Lacerda) Van den Berg,comb. nov.
Basionym: Hoffmannseggella
pabstii Paxton, Paxton’s Fl. Gard. 3: 111: 1852.
" Cattleya purpurata" (Lindl. & Paxton)
F.E.L.Miranda & K.G.Lacerda, Orchids,
Beer, Prakt. Orch. p.213. 1854. nom. inval.
Mag. Amer. Orchid. Soc. 72: 851. 2003.
Sophronitis pabstii (F.E.L.Miranda & Sophronitis purpurata (Lindl. & Paxton)
K.G.Lacerda) Van den Berg & M.W.Chase, Van den Berg & M.W.Chase, Lindleyana
15: 118. 2000. Hadrolaelia purpurata
Kew Bull. 59: 566. 2005.
(Lindl. & Paxton) Chiron & V.P.Castro,
Cattleya pendula (R.C.Mota, P.L.Viana & Richardiana 2: 19. 2002. Chironiella
purpurata (Lindl. & Paxton) Braem,
K.G.Lacerda) Van den Berg,comb. nov.
Basionym: Hoffmannseggella pendula Richardiana 6: 109. 2006. Brasilaelia
R.C.Mota, P.L.Viana & K.G.Lacerda, purpurata (Lindl. & Paxton) Campacci,
Richardiana 4: 1. 2003.
Colet. Orquídeas Brasil. 4(pré-anexo): 100.
Sophronitis pendula (R.C.Mota, P.L.Viana& 2006.
10
VAN DEN BERG - NEW COMBINATIONS IN CATTLEYA
The new combination in Cattleya for this (Withner) V.P.Castro & Chiron, Richardiana
species was listed in the International Plant 2: 23. 2002.
Name Index (IPNI) as proposed by Beer
(1854) but it is invalid for the reasonsCattleya sincorana (Schltr.) Van den Berg,
explained in the comments under C. cernua. comb. nov.
Beer's combination was used by Chadwick
Basionym: Laelia sincorana Schltr.,
& Chadwick (2006), but because they did Orchis 11: 72. 1917.
not include the original basionym in their Sophronitis sincorana (Schltr.) Van den
listing, it could not also constitute a validBerg & M.W.Chase, Lindleyana 15: 118.
publication of this combination.
2000. Hadrolaelia
sincorana
(Schltr.)
Chiron & V.P.Castro, Richardiana 2: 20.
Cattleya pygmaea (Pabst) Van den Berg, 2002.
comb. nov.
Basionym: Sophronitis coccinea subsp. Cattleya tereticaulis (Hoehne) Van den
pygmaea Pabst, Bradea 2(12): 70. 1976.
Berg, comb. nov.
Sophronitis pygmaea (Pabst) Withner,
Basionym: Laelia tereticaulis Hoehne,
Cattleyas & Relatives 3: 77. 1993. Arch. Bot. São Paulo, n.s., 2: 163. 1952.
Hadrolaelia pygmaea (Pabst) Chiron & (“teretecaulis”).
V.P.Castro, Richardiana 2: 21. 2002.
Hoffmannseggella tereticaulis (Hoehne)
H.G.Jones, Florida Orchidist 17: 156. 1974.
Cattleya reginae (Pabst) Van den Berg, Sophronitis tereticaulis (Hoehne) Van den
comb. nov.
Berg & M.W.Chase, Lindleyana 15: 118.
Basionym: Laelia reginae Pabst, Bradea 2000.
2: 24. 1975.
Sophronitis reginae (Pabst) Van den Berg Cattleya verboonenii (F.E.L.Miranda) Van
& M.W.Chase, Lindleyana 15: 118. 2000. den Berg,comb. nov.
Basionym: Laelia verboonenii F.E.L.Miranda,
Hoffmannseggella
reginae
(Pabst)
V.P.Castro & Chiron, Richardiana 2: 23. Bradea 6: 157. 1993.
Sophronitis verboonenii (F.E.L.Miranda)
2002.
Van den Berg & M.W.Chase, Lindleyana
Cattleya rupestris (Lindl.) Van den Berg, 15: 118. 2000. Hoffmannseggella verboonenii
(F.E.L.Miranda) V.P.Castro & Chiron,
comb. nov.
Basionym: Laelia rupestris Lindl., Bot. Richardiana 2: 23. 2002.
Reg. 28: sub. t. 62. 1842.
Sophronitis mirandae Van den Berg & Cattleya virens (Lindl.) Van den Berg,
M.W.Chase, Lindleyana 15: 118. 2000. comb. nov.
Basionym: Laelia virens Lindl., Edward´s
(“mirandai”). Hoffmannseggella rupestris
(Lindl.) V.P.Castro & Chiron, Richardiana Bot. Reg. 30: misc. 1. 1844.
Sophronitis virens (Lindl.) Van den Berg
2: 23. 2002.
& M.W.Chase, Lindleyana 15: 118. 2000.
Cattleya sanguiloba (Withner) Van den Hadrolaelia virens (Lindl.) Chiron &
V.P.Castro, Richardiana 2: 17. 2002.
Berg, comb. nov.
virens
(Lindl.)
Braem,
Basionym: Laelia sanguiloba Withner, Chironiella
Richardiana 6: 110. 2006. Brasilaelia virens
Cattleyas and Relatives 2: 122. 1990.
Sophronitis sanguiloba (Withner) Van (Lindl.) Campacci, Colet. Orquídeas Brasil.
den Berg & M.W.Chase, Lindleyana 15: 4(pré-anexo): 100. 2006.
118. 2000. Hoffmannseggella sanguiloba
11
VAN DEN BERG - NEW COMBINATIONS IN CATTLEYA
Cattleya viridiflora (Verola & Semir) Van Boletim CAOB 58: 49-51.
Beer, J.G. 1854. Praktische Studien an der
den Berg,comb. nov.
Familie der Orchideen. A. Hartinger, Vienna.
Basionym: Hoffmannseggella viridiflora
Brummitt, R.K. 1996. Report of the committee
Verola & Semir, Novon 17(1): 126. 2007.
for Spermatophyta: 44. Taxon 45: 671-681.
Chadwick, A.A. & Chadwick, A.E. 2006. The
Cattleya wittigiana (Barb.Rodr.) Van den
classic Cattleyas. Timber Press, Portland.
Berg, comb. nov.
Chiron, G.R. & Castro, V.P. 2002. Révision des
Basionym: Sophronitis wittigiana Barb.Rodr.,
espèces brésiliènnes du genre Laelia Lindley.
Richardiana 2: 4-28.
Rev. Hort. (1878): 24. 1878.
Barrie, F.R.; Burdet, H.M.;
Hadrolaelia
wittigiana
(Barb.Rodr.) McNeill, J.;
Chiron & V.P.Castro, Richardiana 2: 21. Demoulin, V.; Hawksworth, D.L.; Marhold,
K.; Nicolson, D.H.; Prado, J.; Silva, P.C.;
2002.
Skog, J.E.; Wiersema, J.H.; Turland., N.J.
2006.
International
Code
of Botanical
Cattleya wittigiana (Barb.Rodr.) Van den
Nomenclature (Vienna Code) adopted by the
Berg f. candida (Roeth & O.Gruss) van den Seventeenth International Botanical Congress
Berg, comb. nov.
Vienna, Austria, July 2005. Gantner, Ruggell.
Basionym: Sophronitis wittigiana (Barb.Rodr.) Miranda, F.E.L. 1998. Present status of Brazilian
Fowlie f. candida Roeth & O.Gruss, Die Laelia species. Pages 57-61 In: C.E.B. Pereira.
Proceedings of the 15th World Orchid
Orchidee 56(1): 54. 2005.
Conference, Rio de Janeiro 1996. Naturalia
Publications, Turriers.
Cattleya xanthina (Lindl.) Van den Berg, van den Berg, C.; Higgins, W.E.; Dressler, R.L.;
comb. nov.
Whitten, W.M.; Soto Arenas, M.A.; Culham,
Basionym: Laelia xanthina Lindl., Bot. A.; Chase, M.W. 2000. A phylogenetic analysis
Mag. 85: t. 5144. 1859.
of Laeliinae (Orchidaceae) based on sequence
Sophronitis xanthina (Lindl.) Van den data from internal transcribed spacers (ITS) of
Berg & M.W.Chase, Lindleyana 15: 119. nuclear ribosomal DNA. Lindleyana 15: 962000. Hadrolaelia xanthina (Lindl.) Chiron 114.
& V.P.Castro, Richardiana 2: 20. 2002. van den Berg, C. & Chase, M.W. 2000.
Nomenclatural notes on Laeliinae - I.
(Lindl.)
Braem,
Chironiella
xanthina
Lindleyana 15: 115-119.
Richardiana 6: 109. 2006. Brasilaelia van den Berg, C. & Chase, M.W. 2001.
xanthina
(Lindl.)
Campacci,
Colet.
Nomenclatural notes on Laeliinae - II.
Orquídeas Brasil. 4(pré-anexo): 100. 2006.
Additional
combinations
and
notes.
Lindleyana 16(2): 109-112.
Zappi, D.C. 1994. (1105). Proposal to conserve
LITERATURE CITED
Cattleya violacea (Kunth) Rolfe. Taxon 43:
479.
Baptista, D.H.; Campacci, M.A.; Oliveira, S.M.
2005. Novas combinações de nomenclatura.
ISSN 1809-5348
12
Cássio Van Den Berg