Moss Flora of Mauritius 1 The Moss Flora of Mauritius Jan-Peter Frahm1, Brian J. O'Shea 2 & Boon-Chuan Ho1 1 Nees Institut für Biodiversität der Pflanzen, Universität Bonn, Meckenheimer Allee 170, 2 53115 Bonn, Germany; 141 Fawnbrake Avenue, London SE24 0BG, United Kingdom. Abstract: The mosses reported from Mauritius were compiled from the literature and are listed with localities and references. Included are collections by the first author made in 2007 on the island. Barbula indica, Campylopus flavicoma, Racopilum ayresii and Ectropothecium chenagonii, Groutiella tomentosa, Schlotheimia ferruginosa and Trichostomum crispulum are reported as new to Mauritius. The list includes 238 species. A short survey of the bryological exploration of the island is given. Introduction Mauritius is (with Rodriguez and Réunion) part of the Mascarenes and situated east of Madagascar on 21° S. The climate is determined by the SE winds, resulting in a distinct dry period from May to October and a rainy season from October to April. The rocks are volcanic and originated about 20 million years ago (as in Rodriguez and Réunion). The island is relatively small, about 60 km from W to E and 80 km from N to S, and also relatively low with only a few mountains reaching 800 m altitude. Due to massive habitat destruction and deforestation, the natural forest is almost totally destroyed. Already Renauld (1897) stated "l'extension des cultures a forcément diminué la richesse de la vegetation spontanée". The lower altitudes are almost totally converted to sugar cane plantations. The largest semi-natural part of the island is the Black River National Park in the SW of the island, a high plateau with partial swampy forests, which is eroded by deep gorges. The SE flanks face the wind clouds and receive precipitation of up to 4000 mm or more. The NW parts are distinctly drier, particularly the higher mountains in the NW part of the Island (Le Pouce and Pieter Botha). The vegetation in the higher parts consists of a secondary growth of Sideroxylon bush which is partially forested with Pinus and Eucalyptus. History of the bryological exploration Mauritius apparently is much undercollected as compared with Réunion or Madagascar. This is shown by the fact that only a few records are included in Renauld (1897), who covered the Mascarenes and the Comores. [Either Renauld was following the report of Bescherelle 1879–80 ARCHIVE FOR BRYOLOGY 51 (2009) 2 Frahm, O´Shea, Ho adding on it specimens collected by Robillard & Rodriguez or Renauld has seen the same specimens reported by Bescherelle]. Even conspicuous species such as Leucoloma dichelymoides, which would be collected even by non-bryologists, were not known from Mauritius at that time, and were first recorded by Dixon (1938). Of the numerous species of Leucoloma, Campylopus or Fissidens listed by Renauld (1897), only few to very few originated from Mauritius. The uninhabited island was taken over by the Netherlands in 1598 but was abandoned in 1710. In 1715, France colonized the island, but lost it during the Napoleonic war in 1810 against Britain, until Mauritius became independent in 1968. In spite of the change in ownership of the island, the French continued the biological exploration, and French is still the common language on Mauritius. The bryological (and also general botanical or biological) exploration started at the end of the 18th century. Biographical data of the collectors can be found in Dorr (1997). The first collector of bryophytes on Mauritius was Philibert Commerson (1727–1773), a French physician and naturalist. He travelled with Bougainville around the world 1766–1769. At the end of the trip, he left the expedition to explore Madagascar, Réunion and Mauritius until 1773. Later, 15 species of mosses were attributed to him. Between 1793 and 1802, Chevalier Louis Marie Aubert-Aubert du Petit-Thouars (1758–1832) stayed on Mauritius. He was an officer in the French army who quit his career to study natural science, and planned an expedition together with his brother. He failed to meet him in the harbour at Brest due to the disturbances of the French revolution, and reached Mauritius alone in December 1793, where he started to collect plants, interrupted by trips to Madagascar and Réunion. He returned to France in 1802. From his large collection of plants, there were ten species of mosses new to science, of which two were later synonymized. Jean Baptiste Bory de Saint Vincent (1778–1846) was a French Baron and soldier. He joined an expedition to the "Terres Australes" (Australia) as a zoologist with the ships "Naturaliste" and "Géographe", but was left on the Mascarenes because of health reasons. He stayed a month in April 1801 on Mauritius and stayed on Réunion again in 1802 before his return to France. Next Louis Hyacinthe Boivin (1808–1852), a French botanist and plant collector, especially on the Indian Ocean Islands, took part as botanist on the Oise Expedition (1846–52) and visited Mauritius in 1847–49. In February 1853, Nils Johann Andersson (1821–1888) collected mosses on Mauritius during a voyage around the world. The species were identified and published by Ångström (1873). The only Mauritian bryophyte collector was Victor de Robillard. He was a conchologist and ichthyologist, but also collected cryptogams, during the years 1840–1880. His moss collections are cited by Renauld (1897) from the year 1876. Campylopus robillardei was named after him. Frère Rodriguez, a French priest, collected plants in various parts of the world where he was working, including on Réunion and Mauritius in 1891–92 (according to Dorr, 1997) or 1889 (according to Renauld, 1897). The mosses were named by Renauld, the hepatics by Stephani. Alfred Voeltzkow (1860–1947) was a German zoologist and traveller, especially in East Africa and Madagascar, who stayed in the Mascarenes 1890–93 and 1903–05. All nineteenth century records were summarized by Renauld (1897), who listed 127 species from Mauritius (amongst the more numerous records for Réunion, Madagascar and the other East African Islands). In the 20th century, the collecting activities were much less, although Mauritius was much easier to reach, especially during the recent decades, when the island became a tourist attraction. In 1935 and 1962, R.E. Vaughan collected mosses on Mauritius, which were identified and published by Dixon (1938), and was Honorary Curator at the Mauritius Herbarium at Reduit. ISSN 0945-3466 Moss Flora of Mauritius 3 Gillis Een visited Mauritius (and Réunion) in 1962, and his liverwort collections were published by Arnell (1965) and mosses by Bizot (1968). More identification of mosses were published later by Een in a series of papers (Een 1976, 1978, 1989, 1993, 1997) after retirement. He also sent specimens to C.C. Townsend, who also received specimens from Mark Coode, who worked on the “Flore des Mascareignes” at Kew. In 1966, Gunnar Erdtman, a Swedish palynologist, stayed in Mauritius to collect pollen and spores and also collected some bryophytes, which were 34 years later identified and published by Een (2000). In 1972, Marshall Crosby collected on Mauritius. His specimens are deposited in MO and were included in taxonomic revisions. In 1978 and 1985, Hiroshi Kanda from the Japanese Institute of Polar Research collected on Mauritius, apparently on the way to or from Antarctica. He collected about 150 specimens, which are listed in a printed database of Antarctic mosses (Kanda 1987); however, only two of them were identified. David H. Lorence collected between 1971 and 1976. Many bryophyte specimens are in MO. Finally, Tamas Pocs has also collected there in 1995–1996. In 2007, the first author spent his holidays on Mauritius. During this two-weeks trip in September, at the end of the dry season, he collected bryophytes in the southeast of the island, mainly within the Black River National Park, which belongs to the highest part of the island (up to 800 m elevation) and is the only larger relatively natural area. The identified collections are included here. The specimens are kept in the herbarium of the author in BONN. Almost all records from the 19th century (representing more than half of the total records) have no locality data. This makes it impossible today to locate rare species. However, the most frequently visited areas of Mauritius were the mountain of Le Pouce just above the capital and Curepipe south of Port St. Louis (an area which is completely settled today), whereas the mountains in the south of the island (Montagne de la Rivière Noir) were more difficult to access at that time. The data from the nineteenth century were taken from Renauld (1897) but nomenclaturally updated. In a number of occasions records for Mauritius were found to be incorrect, as early collections did not always distinguish between the three islands of the Mascarenes, and in particular a number of taxa recorded for Mauritius were in error, only occurring on Réunion; this has resulted in a number of deletions from the African checklist. Many records from the last half of the 20th century resulted from monographs and revisions, based on material collected earlier (mostly in the 19th century), showing the importance of herbarium specimens, even when they were initially misidentified. The publications from the "taxonomic period" between 1970 and 2000 added significantly to the publishing of previously unpublished records (number of records in brackets, data extracted from O'Shea 2006) Allen et al. 1986 (2) Argent 1973 (1) Bizot 1974 (1) Buck 1979 (1) Buck 1980 (1) Crosby et al. 1983 (2) De Sloover 1983 (3) De Sloover 1986 (8) Dirkse et al. 1991 (1) Eddy 1985 (1987) (5) Ellis 2003 (1) ARCHIVE FOR BRYOLOGY 51 (2009) 4 Frahm, O´Shea, Ho Enroth 1994 (1) Frahm 1985 (11) Goffinet 1996 (1) Hyvönen 1989 (3) Kruijer 1997, 2002 (2) LaFarge 2002 (1) Mohamed 1979 (1) Ochi 1972 (8) Orbán 1978 (1) Orbán 1981 (3) O'Shea et al. 1997 (3) Pursell 1987 (1) Reese 1991 (2) Salazar Allen 1993 (5) Snider 1975 (1) Sollman et al. 1996 (1) Tixier 1993 (1) Touw 1976 (1) Vitt 1981 (1) List of species The first and also last compilation of the mosses of the East African Islands dates back to Renauld in 1897, who published the famous "Prodrome de la flore bryologique de Madagascar, des Mascareignes et de Comores". The author lists 121 species of mosses, of which 46 species (one third!) were said to be endemic and 58 in common with Réunion. To get an overview of the moss flora of Mauritius, all species recorded by O'Shea (2006) for Mauritius were entered in a spreadsheet together with the references in chronological order. The list was completed with the records cited in the literature. The checklist by Crosby et al. (1983) could not be used because it does not differentiate between the islands of the Mascarenes. The extract of this database revealed 253 species which so far were recorded from Mauritius This is about a hundred species more than the number of hepatics known from Mauritius, of which 158 were counted in the checklist of the East African Island by Grolle (1995), updated by Wigginton (2009). (For comparison: Madagascar 372, Réunion 260, Rodriguez 27, Seychelles 108, Comores 143 liverwort species). The references from the literature were added to this list. In a few cases, no references were found for the records in the database, and in others cases, species were added. The regard of synonyms lowered the number of species again, so the provisional flora now comprises 238 species of mosses. Conspicuous is the high number of species which are known only from one record, showing the insufficient state of knowledge of the mosses of Mauritius. Also, records of ephemeral species are lacking, which means either that there are none, or the collections on the island has been done only during the dry season. * = taxon new to the island Acroporium megasporum (Duby) M.Fleisch. [Hypnum megasporum Duby, Sematophyllum megasporum (Duby) A. Jaeger; Macrohymenium megasporum (Duby) Kis; Rhaphidostegium megasporum (Duby) Besch.] ISSN 0945-3466 Moss Flora of Mauritius 5 Ad ligna, de Robillard hb. S; (Duby 1879: 12 as Hypnum megasporum, Bescherelle 1880: 307, Renauld 1897, Een 1997); Rodriguez (Renauld 1891: 212); on tree near Curepipe, leg. Vaughan 1933 (Dixon 1938); Macabé, upland forest 600 m, leg. Een (Bizot 1968); Macabé, upland forest 1800 ft., (Bizot 1968); Curepipe, leg. Vaughan 1962; Rivière du Poste near Grand Bassin (Een 1997). Aerobryopsis capensis (Müll.Hal.) M.Fleisch. [Aerobryum pseudocapense Müll.Hal.; Aerobryidium subpiligerum (Hampe) Cardot] Without locality, leg. de Robillard (Renauld 1891: 210); Rodriguez (Renauld 1871: 211); rain forest, Mt. Cocotte, leg. Vaughan 1935 (Dixon 1938); without locality, leg. Erdtman (Een 2000); Reserve naturelle de lande de Petrin 650 m (Bizot 1968); Le Pouce, forest halfway to the top; Mont Cocotte, plateau N of the mountain 2300 ft. (Een 1976); without localities, leg. Commerson, Perrollet, Andersson, Mad. Lecoultre (Bescherelle 1880: 268 as Aerobryum pseudocapense) Aerobryopsis cirrifolia (Schwägr.) Paris 'Insula Mauritii: Guil. Sieber' (Müller 1851: 138). Aerobryopsis longissima (Dozy & Molk.) M.Fleisch. [Aerobryopsis wallichii (Brid.) M.Fleisch.] Without locality, leg. Mme. Lecoultre 1871 (O'Shea et al. 1997). Anoectangium aestivum (Hedw.) Mitt. (? Anoectangium borbonense Besch.) Mt. Cocotte, leg. Vaughan 1935 (Dixon 1938); Le Pouce, sur les rochers minés pour faire passer la vieille route de Port Louis à St. Pierre, 550 m, leg. Een (Bizot 1968). Black River Nationalpark, Magdalena Falls, trail to Mt. Cocotte, leg. Frahm 2007. A. borbonense seems to be a local name for the widespread A. aestivum. Anomodon pseudotristis (Müll.Hal.) Kindb. [Haplohymenium pseudotriste (Müll.Hal.) var. robustum Bizot; Pseudoleskea tenuissima Besch.; Haplohymenium tenuissimum (Besch.) Broth.] Without locality, leg. Duisabo (Bescherelle 1880: 288, Renauld 1891: 210, 1897), de Robillard (Renauld 1891: 210, 1897); Le Pouce, forest halfway to the top (Bizot 1968). Archidium ohioense Schimp. ex Müll.Hal. [Pleuridium globiferum Brid.] Without locality, (Renauld 1897 as Pleuridium globiferum, Snider 1975); sur la terre nue mouillee, adhere aux frondes des Lichens crustaces, d’apres Bridel (Bescherelle 1879: 39 as Pleuridium globiferum) Atrichum androgynum (Müll.Hal.) A.Jaeger Jardin botanique de Curepipe, Onraedt 1973 (De Sloover 1986). Barbella capillicaulis var. angustifolia Dixon & Thér. On Litsoea near Curepipe, leg. Vaughan 1933 (Dixon 1938). Barbella capillicaulis var. capillicaulis (Renauld & Cardot) Cardot (Een 1993b). Barbula inclinans Schimp. ex Besch. Without locality, leg. Ayres (Bescherelle 1879: 343, Renauld 1897). *Barbula indica (Hook.) Spreng. Black River Nationalpark, Magdalena Falls, trail to Mt. Cocotte, leg. Frahm 2007. Bartramia gigantea Bory Without locality, (Fransén 2004). Brachymenium acuminatum Harv. in Hook. [B. borgenianum Hampe; Mielichhoferia densifolia Ångström; Brachymenium densifolium (Ångström) Broth., Bryum pungens Taylor ] Without locality, leg. Ångström as Brachymenium densifolium (Ochi 1972); leg. Robillard as Brachymenium borgenianum (Renauld 1891: 210, 1897; Ochi 1972); without locality, leg. Andersson 1853 (Bescherelle 1879: 377 as Mielichhoferia densifolia, Renauld 1897, not cited by Ochi 1972). J.Ängström s.n., isotype of B. densifolium, (S-PA; L); Robillard s.n., as Bryum borgenianum (H) (Ochi 1972: 4–5). Rosulabryum densifolium (Brid.) Ochyra is an accepted name and not a synonym, but Brachymenium densifolium (Ångström) Broth. is synonym of Brachymenium acuminatum. ARCHIVE FOR BRYOLOGY 51 (2009) 6 Frahm, O´Shea, Ho Brachymenium eurychelium Besch. Plateau au nord de Mt: Cocotte alt 700 m, leg. Een (Bizot 1968). Brachymenium exile (Dozy & Molk.) Bosch & Sande Lac. 2 km NW of le Petrin, slope in forest (Müller 2002). Brachymenium leptophyllum (Bruch & Schimp. ex Müll.Hal.) Bruch & Schimp. ex A.Jaeger [Brachymenium longicolle Thér.] Without locality, leg. Robillard as B. speirocladum (Ochi 1972). Brachymenium nepalense Hook. in Schwägr. Without locality, leg. Voeltzkow, type of Brachymenium microcarpum and B. submicrocarpum (Ochi 1972). Brachythecium borgenii (Hampe) A.Jaeger Without locality, Robillard 1876, 1881 (Een 1997). Brachythecium implicatum (Müll.Hal.) A.Jaeger [Brachythecium atrotheca (Duby) Besch.] Ad arborum cortices superficiem magnam saepe obtegens, de Robillard (Duby 1879: 8, Bescherelle 1880: 294, Renauld 1891: 201); Rodriguez 1889 (Renauld 1891: 212); Ravines de la Réunion, Vacoa, Darnty 1876 (Bescherelle 1880: 294, Renauld 1897). [Miss out in O’Shea 2006] Breutelia magdalenae De Sloover Yémen, alt. 500 m. base des arbres d'une savane, déc. 1973, Onraedt 73 ma 466 et 467 (Hb. Onraedt), (De Sloover 1975: 266–269, fig 230–250). Bryoxiphium norvegicum (Brid.) Mitt. Plaine Champagne Nature Reserve, on rock cliff by a stream, rather shaded place, October 1980, leg. Seto M-57 (OSA) (Kimura & Seto 2003). Bryum apiculatum Schwägr. [Bryum nitens Harv.; Bryum laetenitens Müll.Hal. in Besch.; Gemmabryum apiculatum (Schwägr.) J.R. Spence & H.P. Ramsay, Bryum nitens Harv. Bryum alteoperculatum Besch.; Bryum laetenitens Müll. Hal.] Without locality, Robillard, 1876, holotype of Bryum laetenitens (BM); Melville (H) (Ochi 1972: 64); above St Paul, alt. 2000 m. (Een 1976: 214). Without locality, leg. Aubert du Petit Thouars (Bescherelle 1880: 234, Renauld 1897 as Bryum alteoperculatum); without locality, leg. Robillard (type of Bryum laetenitens, Bescherelle 1880: 235, Renauld 1897, Ochi 1972). Bryum argenteum var. argenteum Hedw. Above St. Paul, alt. 2000 m (Een 1976: 214) Bryum aubertii (Schwägr.) Brid. [Rhodobryum aubertii (Schwägr.) Thér.] Without locality, leg. Aubert du Petit Thouars (Bescherelle 1880: 236, Renauld 1897, Ochi 1972). Bryum billardierei var. billardieri Schwägr. [Bryum truncorum (Brid.) Brid.; Rosulabryum billarderi (Schwägr.) J.R. Spence] Without locality, leg. Mairie; leg. Robillard as Bryum ischyospeiron (Renauld 1891: 210; Ochi 1972); Macabé, upland forest 1800 ft. (Een 1976); High Forest, on damp rocks det. Dixon as Bryum nanorrhodon; without locality leg. Vaughan (Een 1989); Black River National Park, Les Mares südlich Petrin; Magdalena Falls, trail to Mt. Cocotte, leg. Frahm 2007. Bryum coronatum Schwägr. [Bryum eurystomum Renauld & Cardot; Gemmabryum coronatum (Schwägr.) J.R. Spence & H.P. Ramsay] Rivière du Mat, leg. Rodriguez, type of Bryum eurystomum (Ochi 1972). Bryum erythrocaulon (Schwägr.) Brid. [Bryum nanorrhodon Müll.Hal. in Besch.] Without locality, leg. Commerson, Duisabo 1838; Montagne de la Rivière Noire, leg. Boivin (Bescherelle 1880: 237, Renauld 1897), Robillard (Renauld 1897); Robillard 1876, types of Bryum ischyrodon and Bryum nanorrhodon (Bescherelle 1880: 237, Mohamed 1979); leg. Commerson, type; Perrier Forest Reserve leg. Goode (Mohamed ISSN 0945-3466 Moss Flora of Mauritius 7 1979); on damp rocks, High Forests, leg. Vaughan (Dixon 1938); Macabé, upland forest 1800 ft. (Een 1989). This record has been published erroneously as Bryum billardierei by Een (1976). Bryum huillense Welw. & Duby [Rosulabryum huillense (Welw. & Duby) Ochyra] Without details, hb. Melville as Bryum nanorrhodon (Renauld 1891: 210; Ochi 1972). Bryum leptospeiron Müll.Hal. Without locality, leg. Robillard (type, Bescherelle 1880: 236, Renauld 1891: 210, 1897; Ochi 1972); bare rock in Ravenal swamp, near Midlands, leg. Vaughan 1935 (Dixon 1938). Callicostella brevipes (Broth.) Broth. Le Pouce 1800 ft. (Een 1997). Callicostella erosotruncata Cardot [Schizomitrium erosotruncatum (Cardot) Ochyra] Without locality, Robillard s.n., as Hookeria seychellensis (PC) (Renauld 1891: 210; [this must be an error] Demaret & Potier de la Varde 1952: 341 (Een 1993b). Callicostella fissidentella (Besch.) Kindb. Macabé, upland forest alt 550m; Le Pouce, forest halfway to the top, leg, Een (Bizot 1968). Callicostella lacerans (Müll.Hal.) A.Jaeger Malherbe, leg. Vaughan 1935 (Dixon 1938). Callicostella parvicellulata Demaret & P. de la Varde Macabé, upland forest 1800 ft.; Malherbe, pine woods near Curepipe, leg. Vaughan 1935 (Een 1997). Callicostella salaziae (Besch.) Kindb. Mont Cocotte, northern slope (Een 1997). Callicostella seychellensis (Besch.) Renauld [Hookeria seychellensis Besch.; Schizomitrium seychellense (Besch.) Ochyra] Without locality, de Robillard, as Hookeria seychellensis (Renauld 1891: 210). Calliergonella cuspidata (Hedw.) Loeske Curepipe, Botanical Garden, meadow (Müller 2002) Calymperes graeffeanum Müll.Hal. [Calymperes robillardei Besch.] Without locality, leg. Robillard (Renauld 1897). Calymperes hispidum Renauld & Cardot [Calymperes borgenii Kiaer; Calymperes crassilimbatum Renauld & Cardot] Le Pouce, sur les rochers minés pour faire passer la vieille route de Port Louis à St. Pierre, 550 m, leg. Een (Bizot 1968); without locality and collector (Reese 1991). Calymperes palisotii Schwägr. Le Pouce 1800 ft, det. Orbán, EGR (Een 1993: 48). Calymperes tenerum Müll.Hal. Ile aux Aigrettes, leg. Erdtman 1966 (Een 2000); St. Aubin, valley below the village leg. Frahm 2007. Calyptrochaeta asplenioides (Brid.) Crosby [Eriopus asplenioides (Brid.) Besch.] Bassin Anglais, Curepipe, near cave, alt. 500 m, leg. Vaughan 1935 (Dixon 1938). Campylopus arctocarpus ssp. madecassus (Besch.) J.-P.Frahm [Campylopus madecassus Besch.] Without locality, leg. Robillard, Voeltzkow as Campylopus boryanus; Central Plateau, Le Petrin, leg. Barclay 1968; Crown land «Perrier» leg. Barclay 1967; Curepipe, Park Hotel, leg. Henderson 1967 (Frahm 1985); Versant N du Mt. Cocotte (Bizot 1968); Black River National Park, Black River Visitors Center, Mackabee trail; Black River National Park, trail from road to Piton de la Petite Rivière Noire, leg. Frahm 2007. Campylopus arcuatus (Brid.) A.Jaeger [Campylopus capitiflorus Mont.] ARCHIVE FOR BRYOLOGY 51 (2009) 8 Frahm, O´Shea, Ho Without locality, leg. Rodriguez 1891; foot of tree near Mare aux Vacoas 600 m alt., leg. Vaughan (Dixon 1938); Les Mares, leg. Vaughan; Rivière du Poste, leg. Coode (Frahm 1985); Black River National Park, Plaine Champagne, Parakeet trail, leg. Frahm 2007. Campylopus aureonitens (Müll.Hal.) A.Jaeger [Campylopus lonchoclados Müll.Hal. in Geh.; Campylopus valentinii Besch.; Campylopus flavoviridis Schimp. in sched.; Campylopus aongstroemii Müll. Hal. ex Ångström] Without locality, leg. Rodriguez 1889 (Renauld 1891: 211, 1897); leg. Robillard 1876 as Campylopus lonchoclados (type), Campylopus valentinii and Campylopus flavo-viridis; leg. Bernier as Campylopus valentinii; leg. Darutz as Campylopus lonchoclados; leg. Rodriguez as Campylopus lonchoclados, leg. Pike; trail up Le Pouce leg. Crosby 1972; Corps du Garde; Midlands reservoir; Rivière du Poste, leg. Coode 1974; Petrin leg. Barclay (Frahm 1985); Black River National Park, Magdalena Falls, trail to Mt. Cocotte; between Case Noyale and Black River National Park, leg. Frahm 2007. Without locality, leg. Andersson as C. aongstroemii (Bescherelle 1879: 326); sur les rochers, ou il forme de larges plaques de plusieurs metres de longueur, leg. Robillard (Bescherelle 1879: 326 as C. lonchoclados). Campylopus cambouei Renauld & Cardot Perrier Nature Reserve at Lagoa de Vacoas (Een 1989); west flank of Mt. Laporte 1000 ft., leg. Barcley (Frahm 1985). *Campylopus flavicoma Müll.Hal. ex Broth. Black River National Park, Les Mares south of Petrin, Sideroxylon swamp, on soil, leg. Frahm 2007. This species was formerly included in the genus Bryohumbertia (and was therefore not included in the treatment of African Campylopus species (Frahm 1985), however, molecular studies (Stech 1999) revealed that this genus cannot be maintained but has to be included in Campylopus. The former species of Bryohumbertia belong to a distinct group with verticillate leaves and are represented in each tropical region with one species, Campylopus filifolius in the Neotropics, Campylopus flavicoma in Africa and Campylopus subcomosus in SE Asia. Campylopus flexuosus var. flexuosus (Hedw.) Brid. [Campylopus boivinianus Besch.; Campylopus brachymastyx Müll.Hal. in Besch.] Without locality, leg. de Robillard as Campylopus brachymastyx (Bescherelle 1879: 324, Renauld 1891: 210); Rodriguez (Renauld 1891: 210), without locality, leg. Boivin 1849 (type of Campylopus boivinianus, Bescherelle 1879: 320, Renauld 1897, Frahm 1985); leg. Voeltzkow 1904; leg. Duisabo; leg. Robillard 1876, leg. Rodriguez 1889 (Renauld 1897, Frahm 1985); Petrin to Grand Bassin Road 2100 ft., leg. Barclay; Le Petrin 2100 ft., leg. Barclay, 1969, 1974; leg. Vaughan 1962; near Rivière du Poste, leg. Coode 1974; Les Maries, leg. Vaughan (Frahm 1985); Black River National Park, Les Mares südlich Petrin, leg. Frahm 2007. Without locality, leg. Robillard (Bescherelle 1879). Campylopus flexuosus var. incacorralis (Herzog) J.-P.Frahm Le Petrin, upland heath vegetation, leg. Coode (Frahm 1985). Campylopus julaceus ssp. arbogastii (Renauld & Cardot) J.-P.Frahm Le Petrin; near Curepipe Point, leg. Barclay (Frahm 1985); Petrin Heath Nature Reserve (Een 1989); Black River National Park, Magdalena Falls, trail to Mt. Cocotte; Bel Ombre trail, leg. Frahm 2007. Campylopus pilifer ssp. pilifer Brid. Le Pouce, on top; without locality, Vaughan 1935 (Een 1989); near Curepipe Point, leg. Barclay (Frahm 1985). The record of Campylopus introflexus (Hedw.) Brid. var. mauritianus Dixon with leaves "haud reflexis" might belong here. Campylopus praetermissus J.-P.Frahm Malenga at foot of Deux Mamelles, leg. Coode (Frahm 1985); SE part of the island, on coastal plain, leg. Erdtman (Een 2000). ISSN 0945-3466 Moss Flora of Mauritius 9 Campylopus robillardei Besch. Without locality, leg. Robillard (type Bescherelle 1879: 322, Renauld 1891: 211, 1897, Frahm 1985); Rodriguez 1889 (Renauld 1891: 211, 1897, Frahm 1985); Chemin cheval near Grand Bassin, leg. Vaughan 1933 (Dixon 1938); Petrin, upland heath, leg. Barclay 1969, 1974; Piton Poule, leg. Barclay 1974; Petrin Heath Nature Reserve; ibid. leg. Vaughan 1962; Chemin Cheval near Grand Basin, leg. Vaughan (Een 1989, Frahm 1985); Black River National Park, Magdalena Falls, trail to Mt. Cocotte; Piton Savanne; between Case Noyale and Black River National Park, leg. Frahm 2007. Campylopus thwaitesii (Mitt.) A.Jaeger [Campylopus fuscolutescens Renauld & Cardot] Without locality, leg. Robillard (type of Campylopus fusco-lutescens) (Renauld 1897). Campylopus trachyblepharon ssp. comatus (Renauld & Cardot) J.-P.Frahm [Campylopus comatus Renauld & Cardot] Tourbière à Sphagnum sur le plateau du Mt. Cocotte alt. 700m (Bizot 1968); Petrin Heath Nature Reserve; in the uplands, leg. Vaughan 1936 (Een 1989); Mare aux Vacoas (Dixon 1938); Mare Longue Plateau, leg. Coode 1973 (Frahm 1985); Black River National Park, Les Mares S of Petrin, leg. Frahm 2007. Cardotiella appendiculata (Renauld & Cardot) Vitt Le Pouce, forest halfway to the top, leg. Een (Goffinet 1996). Catagonium nitens ssp. nitens (Brid.) Cardot [Acrocladium aubertii Brid.] Without localities, Bory, du Petit Thouars (Bescherelle 1880: 316, Renauld 1897). Ceratodon purpureus (Hedw.) Brid. [Ceratodon purpureus var. brevidens Besch.] Near Grand Bassin, on the path to Savanne, leg. Bouton 1864 (Dixon 1938). Without locality, leg. Ayres (Bescherelle 1879: 339 as C. purpureus var. brevidens) Chaetomitrium borbonicum Besch. On dense carpets in upland wood, leg. Vaughan 1935 (Dixon 1938). Cyclodictyon aubertii (P.Beauv.) Kuntze [Hookeria aubertii (P. Beauv.) Müll. Hal.] Without locality, leg. Aubert du Petit-Thouars (Bescherelle 1880: 282 as Hookeria aubertii); Grotte près di Bassin Anglais de Curepipe, alt. 550 m; Le Pouce, forêt à la base de la montagne (Bizot 1968); Malherbe, pine woods near Curepipe, leg. Vaughan 1935 (Een 1997). Cyclodictyon vallis-gratiae (Müll.Hal.) Kuntze Without locality, Robillard s.n. (Demaret & Potier de la Varde 1951: 42) Cyclodictyon vesiculosum (Brid.) Kuntze [Hookeria vesiculosa (Brid.) Müll. Hal.] Without locality, Belanger s.n. sterile (Bescherelle 1880: 283, Renauld 1898: 214) Ditrichum difficile (Duby) M.Fleisch. [Leptotrichum boryanum Müll.Hal.] Without locality, leg. Bory (Bescherelle 1879: 340, Renauld 1897). Ectropothecium intertextum Renauld & Cardot Macabé, upland forest (Bizot 1968); without locality, de Robillard (Renauld 1897); without locality, leg. Erdtman (Een 2000). Ectropothecium mauritianum Renauld & Cardot [Ectropothecium arcuatum Renauld & Cardot] Without locality, Rodriguez 1888 (Renauld 1897). Ectropothecium nishimurii O'Shea & Ochyra [Ctenidium mauritianum Broth.] Malherbe, colonizing base of pine stumps, leg. Vaughan 1935 (Dixon 1938 as Ctenidium mauritianum). Voeltzkow s.n., 1904 (Nishimura 1985: 75; O'Shea & Ochyra 2000: 121) Ectropothecium paillotii Renauld & Cardot Without locality, Mauritius (Renauld & Cardot 1915: 451) Ectropothecium regulare (Brid.) A.Jaeger Without locality, de Robillard, Rodriguez 1889 (Renauld 1891: 210–211, Renauld 1897); Grateloup; Grois bois, Boivin; Mme. Lecoutre; sur les bois morts, ravines de Réunion à Vacoa et près de Curepipe, Darnty (Bescherelle 1880: 319, Bizot 1968); Le Pouce, forest ARCHIVE FOR BRYOLOGY 51 (2009) 10 Frahm, O´Shea, Ho halfway to the top (Bizot 1968); Curepipe, cave near Bassin Anglais; Mare aux Vacoas; Macabé 1800 ft., Le Pouce (Een 1997). Ectropothecium regulare var. angustifolium Thér. Without locality, Rodriguez s.n. (type, Thériot 1923: 461) Ectropothecium valentinii Besch. Without locality, de Robillard (Renauld 1891: 210, 1897). Ectropothecium valentinii var. minus Renauld Without locality, de Robillard (type, Renauld 1889: 85, 1891: 210) Entodon dregeanus (Hornsch.) Müll.Hal. Phoenix, garden on old stone walls and base of trees, leg. Vaughan (Een 1997). Entodon motelayi (Besch.) Paris Without locality, de Robillard (type, Renauld 1897, Buck 1980, Bescherelle 1891). Entosthodon borbonicus Besch. Cascade à 1 kilométre à l'ouest de la Plaine Champagne (Bizot 1968); Le Pouce, on top (Een 1976 as Funaria borbonica). According to Brotherus (1925), Entosthodon borbonicus is autoicous, Entosthodon mauritianus synoicous. Entosthodon mauritianus Schimp. ex Besch. Sur la terre près du sommet de la montagne du Pouce, leg. Darnty 13 Sept 1874 (Bescherelle 1879: 375, Renauld 1897); Black River National Park, trail from road to Piton de la Petite Rivière Noire, leg. Frahm 2007. Fissidens crispulus Brid. [Fissidens sylvaticus Griff., Fissidens arbogastii Renauld & Cardot] Wet clay banks and rocks in stream, Curepipe, leg. Vaughan 1933 (Dixon 1938). Fissidens darntyi Schimp. Sur la terre humide à Plaisance, leg. Darnty (Bescherelle 1879: 334, Renauld 1897). Fissidens ovatus Brid. [Fissidens cymatophyllus Müll.Hal.; Fissidens glaucescens Hornsch.] Le Pouce, sur les rochers minés pour faire passer la vieille route de Port Louis à St. Pierre, 550 m, leg. Een (Bizot 1968); versant Nord du Mont Cocotte (Bizot 1968). Fissidens palmifolius (P.Beauv.) Broth. Without locality, leg. Robillard (Pursell 1987). Fissidens plumosus Hornsch. Cascade un kilométre à l'ouest de la Plaine Champagne (Bizot 1968). Fissidens serratus var. serratus Müll.Hal. [Fissidens papillosus Sande Lac., Fissidens eenii Bizot] Le Pouce, sur les rochers mines pour faire passer le vieux chemin entre Port Louis et St. Pierre, leg. Een 1962, holotype of Fissidens eenii (Bizot 1968, Dirkse et al. 1991); Le Pouce 1800 ft. (Een 1993, Bizot 1968 as Fissidens eenii); St. Aubin, valley below the village, leg. Frahm 2007. Fissidens subplanifrons Bizot & Onr. ex Brugg.-Nann. Cascade de Black River, a la base d'un tronc de Cyathea dans le lit du torrent, 600 m alt., leg. Onraedt (type, Bizot 1974). Floribundaria floribunda (Dozy & Molk.) M.Fleisch. [Papillaria fulvastra Besch., Papillaria robillardi Müll.Hal. in Besch.] Without locality leg. de Robillard, Rodriguez 1888 as Papillaria fulvastra (Renauld 1891: 210, 1897), de Robillard as Papillaria robillardi (Bescherelle 1880: 266, Renauld 1891: 210); on Holalium near Curepipe, leg. Vaughan 1933 (Dixon 1938); Grotte près du Bassin Anglais de Curepipe, alt. 550 m; Le Pouce, forest halfway to the top (Bizot 1968); Curepipe (Kanda 1987); Le Pouce; forest at the base (Een 1976); Le Pouce, forest halfway to the top, Een (1997). Floribundaria vaginans (Welw. & Duby) Broth. [Floribundaria patentissima (Müll.Hal.) M.Fleisch.) Le Pouce, forest halfway to the top (Bizot 1968). Funaria hygrometrica var. calvescens (Schwägr.) Kindb.. ISSN 0945-3466 Moss Flora of Mauritius 11 Montagne du Pouce, Boivin, sept.1849 (Bescherelle 1879: 376; Renauld 1909: 99). Funaria hygrometrica var. mauritiana (Besch.) Paris Montagne du Pouce, dans les Charbonnières, leg. Boivin Sept 1849 (Bescherelle 1879: 376, Renauld 1897); on wooden shingles, Curepipe, leg. Vaughan 1935 (Dixon 1938). Groutiella laxotorquata (Besch.) Wijk & Margad. [Macromitrium laxotorquatum Müll.Hal. in Besch.] Without locality, leg. de Robillard as Macromitrium laxotorquatum (Bescherelle 1879: 362, Renauld 1891: 210); Diego Suarez, leg. Chenagon 1890 (Renauld 1897); Le Pouce, sur les rochers minés pour faire passer la vieille route de Port Louis à St. Pierre, 550 m, leg. Een (Bizot 1968). Groutiella tomentosa (Hornsch.) Wijk. & Marg. Black River Visitor´s Center, Mackabee trail, leg. Frahm 2007. Helicodontium lanceolatum (Hampe & Müll.Hal.) A.Jaeger Phoenix, garden on old stone walls and base of trees, leg. Vaughan (Een 1997). Hildebrandtiella phleoides (Desv. ex Brid.) B.H.Allen & Magill [Hildebrandtiella pachyclada Besch.] Without locality, leg. de Robillard (type); Commerson (Bescherelle 1880: 264, Renauld 1897: 188; Allen & Magill 2007: 41–42). Holomitrium borbonicum Hampe in Besch. Without locality, Aubert du Petit Thouars (Bescherelle 1879: 328, Renauld 1897); base of trees near Petrin, leg. Vaughan 1962 (Een 1978). Holomitrium cylindraceum var. cylindraceum (P.Beauv.) Wijk & Margad. [Holomitrium vaginatum Brid.] Montagne de la Rivière noire, leg. Boivin, leg. Robillard (Bescherelle 1879: 327, Renauld 1897); Mont Cocotte, plateau N of the mountain, 2300 ft. (Een 1976); Le Pouce, near top (Een 1978). Holomitrium cylindraceum var. obtusifolium (Besch.) Wijk & Margad. [Schlotheimia commersoniana Besch.] Without locality, associé au Macromitrium aciculare, leg. Commerson (Bescherelle 1879: 362, Renauld 1897); Trou aux Cerfs near Curepipe leg. Vaughan 1962 (Een 1976). Holomitrium lepervanchei Thér. Reserve naturelle de la lande Petrin alt 650 m (Bizot 1968). Homaliodendron exiguum (Bosch & Sande Lac.) M.Fleisch. [Homalia valentinii Besch.] Without locality, leg. Robillard (Bescherelle 1880: 274 as Homalia valentinii); Grotte près du bassin anglais de Curepipe alt 550 m (Bizot 1968); Macabé, upland forest 1800 ft. (Een 1997); Black River National Park, Black River Visitors Center, Mackabee trail, leg. Frahm 2007 Homaliodendron flabellatum (Sm.) M.Fleisch. [Homaliodendron scalpellifolium (Mitt.) M.Fleisch.] Le Pouce, leg. Balfour s.n. (NY) (De Sloover 1982: 245). Homaliodendron piniforme (Brid.) Enroth [Pinnatella tamariscina (Hampe) Broth.] On Eugenia, bois pomme, Forest of Macabe, leg. Vaughan 1935 (Dixon 1938); Mont Cocotte, northern slope (Een 1978). Hymenostylium scaturiginosum (Besch.) Broth. Corps du Garde Mountain, anfractuosité de rochers, alt. 260 m, leg. Staub & Gueno 1964 (Bizot 1968, Een 1976). Hypnum gracilirameum Müll.Hal. in Renauld Without locality, leg. Robillard; Landmark (Renauld 1897). Hypopterygium tamarisci (Sw. ex Sw.) Brid. in Müll.Hal. [Hypopterygium laricinum (Hook.) Brid., Hypopterygium mauritianum Hampe in Besch., Hypopterygium mauritianum var. nanum ARCHIVE FOR BRYOLOGY 51 (2009) 12 Frahm, O´Shea, Ho Besch., Hypopterygium sphaerocarpum Renauld, Hypopterygium torulosum Schimp. in Besch., Hypopterygium pugiunculum Bosw.] Montagne de la Rivière noire, leg. Boivin 1847; Andersson; Mont Bambou, leg. Darnty, Rodriguez 1889 as Hypopterygium mauritianum (Bescherelle 1880: 327, Renauld 1891: 211, 1897), Rodriguez 1889 as Hypopterygium mauritianum (Renauld 1891: 211, 1897); without locality, leg. de Robillard as Hypopterygium mauritianum var. nanum (Bescherelle 1880: 327, Renauld 1897); Le Pouce, near top (Een 1978 as Hypopterygium laricinum); Black River Visitors Center, Macabee trail leg. Frahm 2007; without locality and collector (Kruijer 1997); without locality, leg. Balfour; Robillard; Onraedt 1971; Montagne de la Rivière Noire, leg. Crosby; Basin Anglais (Kruijer 2002); without locality, leg. Commerson (Bescherelle 1880: 326, Renauld 1897); Bassin Anglais, near cave, leg. Vaughan 1933 (Dixon 1938). Isopterygium argyroleucum Besch. Associé à Octoblepharum albidum, de Robillard (Bescherelle 1880: 314, Renauld 1891: 210, 1897). Isopterygium intortum (P.Beauv.) A.Jaeger Inter lichenes ad cortices dejectos, Robillard as Hypnum aoraton (Duby 1879: 13, Renauld 1891: 210); Bory; Grateloup; sur les paties élevées de la Montagne du Pouce, sur les branches des Sciophila, Boivin 1849; sur les bois morts près de Curepipe, Darnty 22, Mars 1877(Bescherelle 1880: 316, Renauld 1897). Jaegerina. For a key of the African species see Argent (1973). Jaegerina formosa Besch. Without locality, de Robillard (Bescherelle 1880: 263, Renauld 1897); Brotherus (1925); Allen et al. (1986); Perrier Nature reserve near Mare aux Vacoas (Een 1976); Petrin Heath Forest; Macabé, upland forest; Le Poue, near top; Mont Cocotte, northern slope; Le Pouce, near top; Piton Grand Bassin 2336 ft., leg. Vaughan 1962 (Een 1978); Black River National Park, Black River Gorge viewpoint, trail to cascade; trail from road to Piton de la Petite Rivière Noire; Black River Gorge viewpoint, trail to cascade, leg. Frahm 2007. Jaegerina retrosquarrosa B.H.Allen, Crosby & Magill Montagne Cocotte Nature Reserve, low cloud forest, 770 m alt. Type locality, only known from this record (Allen et al. 1986). Jaegerina robillardii Müll.Hal. (Jaegerina excurrens Dixon) Without locality, de Robillard (Bescherelle 1880: 263, Renauld 1897); Brotherus (1925); Allen et al. (1986); rain forest, Mt. Cocotte, leg. Vaughan 1933 (type, Dixon 1938). The species is not mentioned by Argent (1973). Jaegerina solitaria (Brid.) A.Jaeger Without locality, leg. Bory, leg. de Robillard associé à Hildebrantiella pachclada (Bescherelle 1880: 261, Renauld 1897); Rodriguez (Renauld 1891: 211); Brotherus (1925); Le Pouce, forest halfway to the top (Bizot 1968); Le Pouce, on top; Mont Cocotte, on top (Een 1976, Allen et al. 1986). Leiomitrium plicatum (P.Beauv.) Mitt. [Orthotrichum plicatum P. Beauv.] Without locality, leg. M. du Petit-Thouars (Bescherelle 1879: 355, Vitt 1981). Lepidopilidium subrevolutum (Renauld & Cardot) Cardot Macabé, upland forest alt. 550 m (Bizot 1968). Lepidopilum lastii Mitt. [Lepidopilum carrougeauanum Thér. & P. de la Varde] Le Pouce, forest near the top (Een 1997 as Lepidopilum carrougeauanum). Leptodon fuciformis (Brid.) Enroth [Thuidium fuciforme Brid., Pinnatella fuciforme (Brid.) Touw] Without details (Bridel 1812: 432, repeated by Renauld 1897: 224 and Touw 1976: 190 without comment). Leskea mauritiana Besch. ISSN 0945-3466 Moss Flora of Mauritius 13 Without locality, associé au Pterogoniella schimperi et au Leptohymenium fabronioides, leg. Darnty (Bescherelle 1880: 287, Renauld 1897). Leucobryum boryanum Besch. Without locality, leg. Erdtman (Een 2000). Leucobryum comorense Müll.Hal. Without locality, leg. Rev. Rodriguez (Cardot 1904: 106, Renauld & Cardot 1915: 140). Leucobryum isleanum Besch. Without locality, leg. Duisabo (Renauld 1897). Leucobryum isleanum var. molle (Müll.Hal.) Cardot Without locality, leg. Rodriguez 1889 (Renauld 1897). Leucobryum mayottense Cardot [Ochrobryum boivinii Besch.] Sommet du Mont Cocotte alt. 740 m (Bizot 1968); Piton Grand Bassin 2336 ft., leg. Vaughan 1962; Mont Cocotte, northern slope (Een 1978). Leucobryum perrotii Renauld & Cardot Curepipe, leg. Rodriguez (Renauld 1897). Leucoloma amblyacron Müll.Hal in Besch. [Leucoloma brevifolia Dixon & P.de la Varde] Without locality, leg. Robillard (Bescherelle 1879: 313, Renauld 1889, LaFarge 2002); Rodriguez (Renauld 1891: 211). Leucoloma bifidum (Brid.) Brid. Without locality, leg. Bordas (Bescherelle 1879: 309–310, Renauld 1897); leg. Voeltzkow; Robillard (LaFarge 2002). Leucoloma candidulum Müll.Hal. in Besch. Without locality, leg. Robillard (Bescherelle 1879: 314, LaFarge 2002); Rodriguez (Renauld 1891: 211). Leucoloma cinclidiotioides Besch. Perrier Nature Reserve near Mare aux Vacoas (Een 1989, not mentioned in LaFarge 2002). Leucoloma dichelymoides (Müll.Hal.) A.Jaeger Pendulous on heath bushes, in upland scrubs, leg. Vaughan 1935 (Dixon 1938); Petrin, Philippia heath, leg. Vaughan (Dixon 1938); Petrin Heath Nature Reserve (Een 1989); without locality, leg. Coode; Barclay; Een; Guého; Vaughan; Crosby (LaFarge 2002); Black River National Park, Piton Savanne; Black River National Park, Les Mares südlich Petrin, leg. Frahm 2007. Leucoloma fuscifolium Besch. [Leucoloma dubyanum Besch.] Without locality, leg. Rodriguez 1889 (Renauld 1897); M. de Lecoultre (Bescherelle 1879: 16, Renauld 1897, LaFarge 2002), type of Leucoloma dubyanum; Chemin cheval near Grand Bassin, leg. Vaughan 1933 (Dixon 1938). Leucoloma grimmioides P.de la Varde Rivière de galets above '500 foot falls' leg. Guého (Tixier 1993: 41, as Leucoloma guehoi); Mont Pouce; Mt. Deux Mamelles; above La Grande case Royale, leg. Crosby, Pócs, Florens (LaFarge 2002). Leucoloma isleanum Besch. ex Paris Without locality, Voeltzkow s.n. 1903–4 (Renauld 1909: 98). Leucoloma lepervancheri Besch. Without locality, leg. Barclay (LaFarge 2002). Leucoloma longifolium (Brid.) Wijk & Margad. [Leucoloma sinuosum (Brid.) A.Jaeger, Leucoloma sinuosum var. setifolium Besch.] Curepipe, on tree, leg. Vaughan 1933; base of tree, Parc aux Cerfs, leg. Vaughan 1935 (Dixon 1938); plateau au nord du Mt. Cocotte; Macabé, upland forest 550 m alt.; cascade à 1 kilométre à l'ouest de la Plaine Champagne (Bizot 1968); Duisabo, leg. de Robillard ARCHIVE FOR BRYOLOGY 51 (2009) 14 Frahm, O´Shea, Ho (Bescherelle 1879, Renauld 1891: 211, 1897); Perrier Nature Reserve near Mare aux Vacoas; Mont Cocotte, northern slope (Een 1989). Leucoloma pallidulum Thér. Without locality, leg. Rodriguez as Leucoloma candidulum (LaFarge 2002). Leucoloma persecundum Müll.Hal. in Besch. Without locality, leg. Robillard (Bescherelle 1879: 314, LaFarge 2002); Macabé, upland forest 550 m alt. (Bizot 1968). Leucoloma rutenbergii var. porosis Thér. Without locality, leg. Aubert (LaFarge 2002). Leucoloma seychellense Besch. [Leucoloma crepinii Renauld & Cardot, Leucoloma bifidum var. orthothecioides Besch., Leucoloma acutum Müll.Hal.] Without locality, leg. Duisabo; leg. Rodriguez 1896 (type of Leucoloma crepinii, Renauld 1891: 211, 1897; LaFarge 2002); leg Bordas (in herb.) (Bescherelle 1879: 310); leg. Duisabo (Renauld 1897, LaFarge 2002); Le Pouce, sommet 800 m alt.; forest halfway to the top; versant Nord du Mont Cocotte alt. 700 m; réserve naturelle de la lande Pétrin (Bizot 1968); Petrin Heath Nature Reserve; Le Pouce (Een 1989); sur la montagne de Pouce, Darnty, juin 1874, no. 25 (Hb. Schimper) as Leucoloma bifidum var. orthothecioides (Bescherelle 1879: 310; LaFarge 2002); leg. Pike, type of Leucoloma acutum (LaFarge 2002). Leucoloma sinuosulum Müll.Hal. in Besch. Without locality, leg. Robillard (type, Bescherelle 1879: 315, LaFarge 2002), Rodriguez (Renauld 1891: 211), leg. Duisabo (Bescherelle 1879: 315). Leucophanes angustifolium Renauld & Cardot in Renauld [Leucophanes mayottense Cardot] Sommet du Mont Cocotte 740 m alt (Bizot 1968 as Leucophanes mayottense). Leucophanes hildebrandtii Müll.Hal. Without locality, leg. Commerson (Bescherelle 1879: 338, Renauld 1897, Salazar Allen 1993). Leucophanes molleri Müll.Hal. Salazar Allen (1993) only mentions the distribution by a map (p. 227), which appears to show a location on Mauritius. Leucophanes octoblepharioides Brid. Without locality and collector (Salazar Allen 1993). Black River Visitors Center, Mackabee trail, leg. Frahm 2007. Leucophanes renauldii Cardot Mont Cocotte, northern slope (Een 1978). Leucophanes rodriguezii Müll.Hal. Without locality, leg. Rodriguez 1889 (Renauld 1891: 211, 1897, Salazar Allen 1993); Le Pouce, forest halfway to the top (Een 1978). Lopidium struthiopteris (Brid.) M.Fleisch. Without locality and collector (Kruijer 1997), leg. ? Jussieu in hb. S (Een 1997); without locality, leg. Commerson (Kruijer 2002). Macrohymenium acidodon (Mont.) Dozy & Molk. Le Pouce 1800 ft. (Een 1997). Macromitrium calomicron Broth. Without locality or collector (Renauld 1909: 99). Macromitrium fimbriatum (P.Beauv.) Schwägr. Without locality, leg. Commerson; Du Petit-Thouars (Bescherelle 1879: 358, Renauld 1897). Macromitrium fimbriatum var. chloromitrium Besch. Montagne du Pavillon, leg. Boivin (Bescherelle 1879: 359, Renauld 1897). Macromitrium funicaule Schimp. in Besch. ISSN 0945-3466 Moss Flora of Mauritius 15 La Montagne Rouge aus Troix-Ilets, leg. Darnty 1874 (hb. S); Perrier Nature Reserve near Mare aux Vacoas; Mont Cocotte, on top (Een 1978); Sur les arbres de la montagne Rouge, aux Trois-Ilets, leg. Darnty, juin 1874 (Bescherelle 1879: 359) Macromitrium mauritianum Schwägr. Without locality, leg. Sieber; montagne de la Rivière Noire, leg. Boivin 847, Andersson, Robillard, (Bescherelle 1879: 361, Renauld 1891: 210–211, 1897), leg. Rodriguez (Renauld 1891: 210–211, 1897); leg. Voeltzkow 1904 (hb. S); Sommet du Mt. Pouce alt. 800 m; Le Pouce, sur les rochers minés pour faire passer la vieille route de Port Louis à St. Pierre, 550 m, leg. Een (Bizot 1968); Petrin Heath Nature Reserve; Le Pouce, on top; forest half way to the top; Perrier Nature Reserve 1750 ft., leg. Vaughan 1962. Macromitrium mauritianum var. viride Broth. Without details, only recorded from the type specimen (Brotherus 1908: 55). Macromitrium pallidum (P.Beauv.) Wijk & Margad. [Macromitrium aciculare Brid.] Without locality, leg. Aubert du Petit-Thouars; sur les rochers ombragés et bois morts, montagne de Pouce; anse Courtois, montagne de corps de garde, leg. Boivin (Bescherelle 1879: 360, Renauld 1897); Rock, Good End, leg. Vaughan 1935 (Dixon 1938); Le Pouce, halfway to the top (Een 1978); Black River National Park. Piton Savanne; Black River Visitor´s Center, Mackabee trail; Black River gorge viewpoint, trail to cascade, leg. Frahm 2007. Macromitrium voeltzkowii Broth. Without locality, Voeltzkow 1904 (S ex hb. Brotherus) (Renauld 1909: 99). Meiothecium madagascariense (Brid.) Broth. Without locality, leg. Rodriguez, det. Thériot in hb. S; Phoenix, garden on old stone walls and base of trees, leg. Vaughan 1953 (Een 1997). Mesonodon flavescens (Hook.) W.R.Buck [Campylodontium flavescens (Hook.) Bosch. & Sande Lac., Lepyrodon mauritianus Müll.Hal. ex Besch.] Without locality, leg. de Robillard (Bescherelle 1880: 261, Renauld 1891: 210, 1897). Mittenothamnium madagassum (Besch.) Cardot Curepipe, cave near Bassin Anglais (Bizot 1968; Een 1997). Mittenothamnium reptans (Hedw.) Cardot Curepipe, leg. Bijoux 1906 hb. S (Een 1997). Mitthyridium fasciculatum var. fasciculatum (Hook. & Grev.) H.Rob. [Syrrhopodon fasciculatus Hook. & Grev.] Without locality, leg. Robillard (Renauld 1891: 210, 1897). Neckeropsis lepineana (Mont.) M.Fleisch. [Neckera comorae Müll.Hal.] Without locality, Ryres (NY) (Touw 1962: 385); without locality, de Robillard (hb. Duby & Geheeb) as Neckera comorae (Bescherelle 1880: 272; Renauld 1891: 210); without details (Renauld & Cardot 1915: 372). Octoblepharum albidum Hedw. Without localities, leg. Commerson, Robillard, Rodriguez; sur les arbres pourris, leg. Richard; montagne de la Riviere Noire, leg.Boivin, (Bescherelle 1879: 338, Renauld 1891: 210–211, 1897); on Raphia palm, Curepipe Botanic Gardens, leg. Vaughan 1933 (Dixon 1938); Curepipe (Kanda 1987); Black River National Park, Black River Visitors Center, Mackabee trail, leg. Frahm 2007. Orthostichella longinervis (Renauld & Cardot) B.H.Allen & Magill [Orthostichopsis longinervis Renauld & Cardot] Without locality, Rodriguez s.n., det. Thériot (Hb. S) as Orthostichopsis longinervis; Black River, leg. Lorence 1587, 1599 (hb. MO); without locality, leg. Rodriguez (hb. PC) (Allen & Magill 2007); Macabé, upland climax forest, alt. 1800 ft, Een M513, 1962 (Een 1997). Orthostichella rigida (Müll.Hal.) B.H.Allen & Magill ARCHIVE FOR BRYOLOGY 51 (2009) 16 Frahm, O´Shea, Ho Plaines Wilhems, leg. Bijoux (hb. L) (Allen & Magill 2007) Orthostichidium pentastichum (Brid.) B.H.Allen & Magill [Orthostichella pentasticha (Brid.) W.R.Buck] Without locality, leg. Anderson 1853, hb. S; Le Pouce 1800 ft. (Een 1997); without localities, leg. Commerson (type, hb PC), leg. Danty 14 (hb. H-Bescherelle) (Allen & Magill 2007). Papillaria acinacifolia Besch. Parait frequent das l'ile, de Robillard (Renauld 1888, 1891, 1897). Papillaria africana (Müll.Hal.) A.Jaeger [Papillaria boiviniana Besch.] Montagne de la Rivière noire, leg. Boivin; without locality, leg. de Robillard (Bescherelle 1880: 264, Renauld 1888, 1891, 1897); without locality, leg. Rodriguez 1889 (Renauld 1888, 1891, 1897); Perrier Nature reserve near Mare aux Vacoas (Een 1978). Papillaria renauldii Besch. Without locality, de Robillard (Renauld 1888, 1891, 1897). Philonotis gracilescens Schimp. in C.H.Wright Leg. Ayres, hb. S; Mont Cocotte, northern slope (Een 1976). Philonotis mauritiana Ångstr. [Philonotis luteo-viridis Besch.] Without localities, leg. Boivin, Andersson, Robillard (Bescherelle 1880: 244, Renauld 1891: 210–211, 1897); Casacade à 1 kilométre à l'ouest de la Plaine Champagne (Bizot 1968); Le Pouce 1800 m (Een 1976); bord des ruisseaux pres de Plaisance, leg. Darnty juin 1874 as Philonotis luteo-viridis (Bescherelle 1880: 245). Pinnatella minuta (Mitt.) Broth. West flank of Mt. Laporte 800 ft., Barclay 1968 (Enroth 1994). Pogonatum. For keys and illustrations see De Sloover (1986), Hyvönen (1989). Pogonatum belangeri (Müll.Hal.) A.Jaeger Entre les pierres au mont Pouce, leg. Belanger (type, Bescherelle 1880, Renauld 1897, Hyvönen 1989); leg. Voeltzkow; Bel Ombre, Coode 1973; Mare aux Vacoas, Barclay 1972; Piton de la Rivière noire; Piton Grand Bassin, Crosby 1972; Curepipe, Onraedt 1973; Rivière du Pont, Onraedt 1971 (De Sloover 1986); road from Baye du Cap to Chamarel, leg. Frahm 2007. Pogonatum capense (Hampe) A.Jaeger Without locality, leg. Ayres (De Sloover 1986). Pogonatum convolutum (Hedw.) P.Beauv. [Pogonatum rutenbergii (Geh.) Paris] Without locality and collector (De Sloover 1986, Hyvönen 1989). Pogonatum gracifolium Besch. [Pogonatum rubenti-viride (Müll.Hal.) Paris] Without locality, leg. Ayres (De Sloover 1986); Black River National Park, Savanna trail, leg. Frahm 2007. Pogonatum marginatum Mitt. Without locality, leg. Howard s.n. (NY, hb. Mitten) (De Sloover 1986: 245); Steward s.n. (NY) (Touw 1986: 14). There is doubt about this species: Touw (1986: 14) comments: "Mitten's herbarium at NY contains an unidentified specimen labelled 'S.U.S., Mauritius, Mr Steward' (or Howard?) that J.L. de Sloover recognized as being P. marginatum. Though the occurrence of that species in Mauritius cannot be ruled out, mislabelling might have taken place, as British ships sailing between Europe and Sri Lanka may have called at Mauritius. During his work on African Polytrichaceae Dr. De Sloover (pers. comm.) did not find any more collections of P. marginatum." Polytrichastrum formosum (Hedw.) G.L.Sm. [Polytrichum formosum Hedw., Polytrichum calopogon Besch., Polytrichum mauritianum Müll.Hal. in Besch.] Without locality, leg. de Robillard, Rodriguez (Bescherelle 1880, Renauld 1891: 210– 211); leg. Commerson as Polytrichum calopogon (Bescherelle 1880: 256, De Sloover ISSN 0945-3466 Moss Flora of Mauritius 17 1986); Le Pouce, leg. Ayres 1857 (De Sloover 1986). Without localities, leg. Commerson, de Robillard (Bescherelle 1880: 255) Polytrichum. For keys and illustration see De Sloover (1986). Polytrichum commune Hedw. [Polytrichum elatum P.Beauv.; Polytrichum remotifolium P. Beauv.] In montibus ignivomis, du Petit Thouars (Duby 1876: 362); without locality, leg. Ängström as Polytrichum remotifolium, hb. S; au milieu des Sphagnum, leg. Commerson as Polytrichum elatum (Bescherelle 1880: 257, Renauld 1897, De Sloover 1986); Plaine Champagne, heath (Een 1993); Without localities, leg. Commerson, leg. Bory as Polytrichum remotifolium (Bescherelle 1880: 254) Polytrichum subpilosum P.Beauv. [Polytrichum subformosum Besch., fide De Sloover 1986]. Without locality, leg. Du Petit-Thouars, as Polytrichum appressum (Duby 1876: 362); leg. Commerson, de Robillard, Rodriguez 1888 as Polytrichum subformosum (Renauld 1897, De Sloover 1986); leg. Bory, Ayres, Thomas, Bojer, Voeltzkow; Le Pouce, Coode 1973; Curepipe, leg. Rodriguez; Riv. du Poste, Barclay 1967; Onraedt 1971, 1974; Coode (De Sloover 1986); without locality, leg. Erdtman (Een 2000); between Case Noyale and Black River National Park 250–700 m; Black River National Park, Bel Ombre trail, leg. Frahm 2007. Porotrichum. For keys and illustration see De Sloover (1983). Porotrichum elongatum (Welw. & Duby) A.Gepp [Porothamnium comorense (Müll.Hal.) Sim, Porotrichum robillardii Müll.Hal. in Besch. fide de Sloover 1983] Without locality, de Robillard 1876, lectotype of Porothamnium robillardii (Bescherelle 1880: 274, Renauld 1891: 210); Rodriguez (Renauld 1891: 211); Balfour; Voeltzkow 1887; Melville; le Pouce, Ayres 1862; Macabé, Onraedt 1971; Bassin anglais, leg. Vaughan 1933 (Dixon 1938); grotte près du bassin anglais, Onraedt 1981; (Bizot 1968); Le Pouce, forest at base (Een 1976); near Macabé Road turning to Brise Fer 1850 ft., leg. Vaughan 1962; Le Pouce 1800 ft. (Een 1978); St. Aubin, valley below the village, leg. Frahm 2007. Porotrichum madagassum Kiaer in Besch. Le Pouce, Ayres 1862 (De Sloover 1983). Porotrichum stipitatum (Mitt.) W.R.Buck [Thamnium hildebrandtii (Müll.Hal.) A.Jaeger] Without locality, associé au Rhacopilum, de Robillard (Bescherelle 1880: 276, Renauld 1897). Porotrichum usagarum Mitt. [Porotrichum molliculum Broth.] Without locality and collector (De Sloover 1983). Prionodon ciliatus Besch. Without locality or collector (Bizot & Pocs 1974: 431) Pyrrhobryum spiniforme (Hedw.) Mitt. [Rhizogonium spiniforme (Hedw.) Bruch; Rhizogonium mauritianum Hampe ex Besch.] Montagne du Pavillon, leg. Boivin 1847, Vesco 1850, de Robillard, Rodriguez (Bescherelle 1880: 242, Renauld 1891: 211, 1897); Macabé, upland forest 1800 ft.; Le Pouce, forest half way to the top (Bizot 1968, Een 1976); without locality, leg. Vaughan (Dixon 1938); Curepipe, Parc Hobell (Bizot 1968); Piton Grand Bassin 2336 ft., leg. Vaughan 1962 (Een 1978); Black River National Park, trail from road to Piton de la Petite Rivière Noire; Magdalena Falls, trail to Mt. Cocotte, leg. Frahm 2007. The same specimens were identified as Pyrrhobryum spiniforme by Bizot, but as P. mauritianum by Een. Although Een (1978) stressed the differences, Frahm et al. (2003) synonymized the species with the pantropical Pyrrhobryum spiniforme. Also Dixon (1938) had named Mauritian collections as Rhizogonium spiniforme. It could be that the plants from Mauritius form a different population; however, they cannot be differentiated on a species level. Racomitrium lanuginosum (Hedw.) Brid. [Racomitrium lanuginosum var. squalidum (Brid.) Brid.] ARCHIVE FOR BRYOLOGY 51 (2009) 18 Frahm, O´Shea, Ho Without locality, hb. Mus. Paris (Bescherelle 1879: 352 as Racomitrium lanuginosum var. squalidum, Renauld 1897). Racopilum africanum Mitt. Bassin Anglais, on rocks near cave, leg. Vaughan 1933 (Dixon 1938); Grotte près du bassin anglais de Curepipe, alt. 550 m (Bizot 1968); Phoenix, in garden on trees, leg. Vaughan 1962 (Een 1978, 1997); Le Pouce, forest half way to the top (Een 1997); St. Aubin, valley below the village, leg. Frahm 2007. *Racopilum ayresii Mitt. Montagne du Pouce, leg. Ayres; Rodriguez 1889 (Renauld 1897). Racopilum mauritianum Müll.Hal. ex Besch. Montagne de la Rivière noire, leg. Boivin 1847; Grateloup; Andersson; sur les bois morts, près de Curepipe et de Vacoa, leg. Darnty; de Robillard, (Bescherelle 1880: 325, Renauld 1891: 211–212, 1897), Rodriguez (Renauld 1891: 211–212, 1897); Curepipe, cave near Bassin Anglais (Een 1997). Racopilum tomentosum (Hedw.) Brid. Without locality, leg. N.J. Andersson, det. A.J. Ångström, Feb. 1853 (Åongström 1873). Radulina borbonica (Bél.) W.R.Buck [Trichosteleum borbonicum (Bél.) A.Jaeger; Rhaphidostegium borbonicum (Bél.) Besch.; Trichosteleum microdontum (Besch.) Renauld] Without locality, de Robillard (Bescherelle 1880, Renauld 1891: 210); sur les arbres morts, près du Grand bassin, Darnty 13 (Bescherelle 1880: 307, Renauld 1897), Rodriguez 1889 (Renauld 1897); without locality, leg. Balfour (O'Shea et al. 1997); Macabé, upland forest 1800 ft. (Een 1997, O'Shea 2006). Rhizofabronia persoonii var. persoonii (Schwägr.) M.Fleisch. [Fabronia persoonii Schwägr.] Without locality, leg. Bory, de Petit Thouars (Bescherelle 1880: 287, Renauld 1897). Rhodobryum commersonii (Schwägr.) Paris [Bryum commersonii (Schwägr.) Brid.] Sur la terre à Curepipe, leg. Darnty (Bescherelle 1880: 239, Renauld 1897). Rhynchostegium distans Besch. [Rhynchostegium homalobolax Müll.Hal. ex Besch.] Without locality, leg. Duisabo (Bescherelle 1880: 297), de Robillard (Bescherelle 1880: 297, Renauld 1891: 210, 1897). Rutenbergia prionodon (Besch.) Renauld [Leucoloma prionodon Besch.] Sur les arbres, bois de la Réunion, près de Curepipe, 21 mars 1877, leg. Darnty 26 (hb. Schimper), (Bescherelle 1879: 317; Renauld & Cardot 1915: 334). Schlotheimia angulosa (P.Beauv.) Dixon Trou aux Cerfs, leg. Vaughan; Perrier Nature Reserve near Mare aux Vacoas; Petrin Heath Nature Reserve 2200 ft. (Een 1978). Schlotheimia badiella Besch. Without locality, de Robillard, Rodriguez (Renauld 1891: 210–211); resérve naturelle de la lande Pétrin, alt. 650 m (Bizot 1968). Schlotheimia badiella var. brevifolia Renauld Without details, leg. Robillard (Renauld 1889: 82, Een 1993). Schlotheimia badiella var. helicophylla Besch. Without locality, leg. Robillard (Renauld 1897). *Schlotheimia ferruginosa (Hook. & Grev.) Brid. Black River National Park, savannah trail and Piton Savanne, leg. Frahm 2007. Schlotheimia fornicata Duby [Schlotheimia bursata Renauld & Cardot] Ad arbores in sylvis, leg. Robillard (Duby 1879: 5, Bescherelle 1879: 367); Curepipe, leg. Rodriguez 1889 (Renauld 1891: 211, 1897); Bassin Anglais near Curepipe, leg. Vaughan 1933 (Dixon 1938); Perrier Nature Reserve, leg. Vaughan; Petrin 2150 ft., leg. Vaughan (Een 1978); Resérve naturelle de la lande Pétrin, alt. 650 m (Bizot 1968); Perrier Nature Reserve near Mare aux Vacoas (Een 1978). Schlotheimia microphylla var. farlowii Renauld & Cardot ISSN 0945-3466 Moss Flora of Mauritius 19 Without details, hb. Farlow (Renauld 1909: 52). Schlotheimia robillardii Duby [Schlotheimia phaeochlora Besch.] Savane, leg. Boivin; Robillard (Duby 1879: 6, Bescherelle 1879: 370); Rodriguez; bois Descroitzilles, Savane, leg. Darnty (Bescherelle 1879: 370, Renauld 1891: 210–211, 1897); Perrier Nature reserve, leg. Vaughan 1962; Petrin heath, leg. Vaughan 1962, leg. Een (Een 1978); Le Pouce (Een 1978). Scorpiurium circinatum (Brid.) M.Fleisch. & Loeske Curepipe, Botanical Garden, concrete wall of bridge (Müller 2002). Sematophyllum aneuron (Duby) A.Jaeger Without locality, Lecoultre (Duby 1877: 93, 1879: 13). Sematophyllum corticola (Ångstr.) O'Shea [Plagiothecium corticola Ångstr.] Without locality and collector (Renauld 1897), hb. S; Le Pouce, forest, half way to the top, Een (1997). Sematophyllum crassiusculum (Brid.) Broth. [Rhaphidostegium crassiusculum Brid.] Without locality, Duisbao 1839; sur les vieilles souches, gros bois et montagne de la Rivière noire, Boivin 1841 (Bescherelle 1880: 300, Renauld 1897). Sematophyllum longinerve (Duby) A.Jaeger Without locality, Lecoultre (Duby 1877: 92, 1879: 9). Sematophyllum schimperi (Besch.) Broth. [Pterogoniella schimperi Besch.] Without locality, leg. Darnty (Bescherelle 1880: 259, Renauld 1897); on rocks, dry forest of the Trois Mamelles, leg. Vaughan 1935 (Dixon 1938). Sematophyllum subpinnatum (Brid.) E.Britton [Hypnum irrepens Duby; Hypnum lecoultriae Duby; Rhaphidostegium duisaboanum (Mont.) Paris; Sematophyllum caespitosum (Hedw.) Mitt.] Inter lichenes et alios muscos ad arbores, Robillard (Duby 1879: 10, as Hypnum robillardii Duby), longe inter Macromitria sese insinuens, cum Fabronia quadam semper sterili immixtum (Duby 1879: 14, as Hypnum irrepens); without locality, Lecoultre (Duby 1879: 8, as Hypnum lecoultriae); de Robillard (Renauld 1891: 210); Duisabo, sur les rochers à Plaisance et à la Montagne, Darnty, Commerson (Renauld 1897); Malherbe, leg. Vaughan 1935 (Dixon 1938); leg. Robillard as Rhaphidostegium duisaboanum, hb. S.; leg. Andersson as Sematophyllum spec. Without localities, leg. Duisabo, leg. Andersson, leg. de Robillard; sur les richer a Plaisance et a la Montagne fertile, leg. Darnty 10, 11, 27, 28 (Bescherelle 1880: 301 as Rhaphidostegium duisaboanum fo. virescens), without locality, leg. Commerson as Rhaphidostegium duisabonae var. lutescens, leg. Mad. Lecoultre as Rhaphidostegium duisabonae var. rigidiusculum, leg. Duisabo as Rhaphidostegium duisabonae var. rufulum, leg. Duisabo, Boivin Oct 1849 (Bescherelle 1880: 301-302) Sphagnum bordasii Besch. [Sphagnum truncatum var. bordasii (Besch.) A.Eddy, nom. illeg., Sphagnum mauritianum Warnst. (1911), Sphagnum oxycladum var. mauritianum (Warnst.) Warnst.]. Without locality, leg. Bordas 1879 (type, Bescherelle 1880: 330, Renauld 1897, Eddy 1985). Sphagnum perichaetiale Hampe [Sphagnum balfourii Warnst.] Without locality, leg. Balfour (type of Sphagnum balfourianum, Renauld 1897, Eddy 1985); in bog, north of Mt. Cocotte, leg. Onraedt 1971 (Eddy 1985); Mont Cocotte, Sphagnum fen; Petrin Heath Nature Reserve (Een 1993); without locality leg. Erdtman (Een 2000); Black River National Park, Les Mares südlich Petrin, leg. Frahm 2007. Sphagnum rutenbergii Müll.Hal. [Sphagnum mauritianum Warnst. (1891)] Without locality, leg. Ayres (type of Sphagnum mauritianum) (Renauld 1897, Eddy 1985). Sphagnum tumidulum Besch. var. tumidulum ARCHIVE FOR BRYOLOGY 51 (2009) 20 Frahm, O´Shea, Ho Les Mavers, leg. Vaughan 1935 (Dixon 1938); Tourbière à Philippia 600 m alt., leg. Onraedt 1971 (Eddy 1985). Sphagnum violascens Müll.Hal. [Sphagnum obtusiusculum Lindb. ex Warnst., Sphagnum purpureum Schimp. ex Warnst.] Without locality and collector sub Sphagnum obtusiusculum (Renauld 1897); without locality, leg. Blackburn as Sphagnum purpureum (Renauld 1897); without locality, leg. Bory; Rodriguez (Eddy 1985); without locality leg. Erdtman (Een 2000); Black River National Park, Les Mares südlich Petrin, leg. Frahm 2007. Stereophyllum limnobioides Renauld Without locality, leg. Robillard (Renauld 1891: 210, 1897). Probably a synonym of Stereophyllum radiculosum (Buck & Ireland 1985). Syntrichia rufa (Schimp. ex Besch.) O'Shea [Tortula rufa (Schimp. ex Besch.) Broth., hom. illeg., Barbula rufa Schimp. ex Besch.] Without locality, leg. Ayres (Bescherelle 1879: 344, Renauld 1897). Syrrhopodon apertifolius Besch. [Syrrhopodon glaucophyllus Renauld & Cardot], Syrrhopodon prolifer Schwaegr. var. acanthoneuros (Müll.Hal.)Müll.Hal. [S. apertifolius Besch., Syrrhopodon glaucophyllus Renauld & Cardot] auct. Savanne, leg. Boivin (Bescherelle 1879: 350, Renauld 1891: 211, 1897, Orbán 1981); leg. Rodriguez 1889 (Renauld 1891: 211, 1897, Orbán 1981); Resérve naturelle de la lande Pétrin, alt. 650 m (Bizot 1968); Plaine Champagne, heath (Een 1976). Savanne, leg. Boivin; leg. Rodriguez 1889 (Bescherelle 1879, Renauld 1897, Orban 1981, Ellis 2003, 2005); Without locality, leg. Rodriguez 1889 as S. glaucophyllus (type, Renauld 1897, Ellis 2003); leg. Balfour (Ellis 2005); Resérve naturelle de la lande Pétrin, alt. 650 m (Bizot 1968). Plaine Champagne, heath (Een 1976); Black River National Park, Plaine Champagne, Parakeet trail; Piton Savanne, leg. Frahm 2007 (as S. prolifer var. acanthoneuros) Syrrhopodon graminifolius Renauld & Cardot in Renauld Without locality, leg. Crosby (Reese 1991). Syrrhopodon hispidocostatus Renauld & Cardot in Renauld [Syrrhopodon prolifer var. hispidocostatus (Renauld & Cardot in Renauld) Orbán & W.D.Reese] Without locality, leg. Crosby (Reese 1991); (Ellis 2005). Syrrhopodon involutus Schwägr. Malenge, Deux Manelles, leg. Cook 1974 (Orbán 1981). Syrrhopodon mauritianus Müll.Hal. ex Åongstr. Without locality, leg. Andersson (type, Bescherelle 1879: 348, Renauld 1897, Orbán 1981, Reese 1996); leg. Voeltzkow; leg. Jitadorrou (Orbán 1981); without locality and collector (Orbán 1978); without locality leg. Voeltzkow (hb. H-Br) (Reese 1996). Syrrhopodon revolutus Dozy & Molk. Mt. Deuz Mamelles, leg. Lorence (hb. MO) (Fisher 2006). Taxithelium pseudo-amoenum (Bél.) Renauld Without locality, de Robillard (Renauld 1891: 210, 1897). Taxithelium lindbergii (A. Jaeger) Renauld & Cardot [Taxithelium nossianum, Besch.Taxithelium voeltzkowii Broth.] Without locality, leg. Voeltzkow s.n. (type of Taxithelium voeltzkowii) (Câmara 2008); without locality, leg. Balfour as Taxithelium nossianum (O'Shea et al. 1997); Le Pouce, leg. Onraedt 272 (hb. BR) (Câmara 2008) Thamniopsis pappeana (Hampe) W.R. Buck [Hookeriopsis darntyi (Schimp. in Besch.) Broth.; Lepidopilum darntyi Schimp. in Besch.] Sur les vieux arbres de la montagne du Pouce, leg. Darnty 19 13 Sep 1874 (Bescherelle 1880: 281, Renauld 1897). ISSN 0945-3466 Moss Flora of Mauritius 21 Thuidium pseudoinvolvens (Müll.Hal.) A.Jaeger [Cyrto-hypnum pseudoinvolvens (Müll. Hal.) W.R. Buck & H.A. Crum] Without locality, leg. Robillard (Touw 1976). Tortella humilis (Hedw.) Jenn. Garden at Phoenix, leg. Vaughan 1962 (Een 1978). Trachyphyllum inflexum (Harv.) A.Gepp in Hiern [Trachyphyllum fabronioides (Besch.) A.Gepp; Leptohymenium fabronioides Müll.Hal. ex Besch.] Without locality, leg. Robillard 1876, Rodriguez (Renauld 1891: 210–211, Buck 1979); without locality, de Robillard (Bescherelle 1880: 292, Renauld 1897); base of tree in dry forest, Trois Mamelles, leg. Vaughan 1935 (Dixon 1938). Trichosteleum microdontum (Besch.) Renauld Without locality, Balfour, det. H.N. Dixon (O'Shea et al. 1997: 178). Trichosteleum perrotii (Renauld & Cardot) Renauld & Cardot Without details (Renauld & Cardot 1915: 506 + tab. 104: 1). Trichostomum brachydontium var. brachydontium Bruch [Trichostomum ayresii Schimp. in Besch.; Trichostomum ayresianum Schimp. ex Besch.] Without locality, leg. Ayres, type of Trichostomum ayresianum (Bescherelle 1879: 342, Renauld 1897); Le Pouce 1800 ft., (Een 1976); garden at Phoenix (Een 1978 as Trichostomum ayresianum); Le Pouce, road from Port Louis to St. Pierre (Sollman & Een 1996). Road from Case Noyale to Black River National Park, leg. Frahm 2007; Black River Nationalpark, Black River Gorge viewpoint, trail cascade, leg. Frahm 2007. *Trichostomum crispulum Bruch Road from Case Noyale to Black River National Park, leg. Frahm 2007. Trichostomum tenuirostre var. tenuirostre (Hook. & Taylor) Lindb. Plaine des Cafres, G. de l'Isle, Lépervanche as Symblepharis circinata (Bescherelle 1879: 20). Ulota fulva Brid. Without locality, leg. Robillard (Bescherelle 1879: 353, Renauld 1897). Vesicularia alboviridis (Renauld) Broth. Without locality, leg. Robillard (Renauld 1891: 211, 1897). Vesicularia ayresii (Besch.) Broth. [Ectropothecium ayresii Schimp. ex Besch.] Without locality, leg. de Robillard (Renauld 1891: 211); sur les rochers de la cascade du Réduit, Darnty 24 Mai 1874 (Bescherelle 1880: 319, Renauld 1897). Vesicularia bescherellei (Renauld) Broth. Without locality, leg. Robillard 1876, hb. S, (Renauld 1891: 210, 1897; Een 1997); on rocks in stream near Curepipe, leg. Vaughan 1935 (Dixon 1938). Vesicularia galerulata (Duby) Broth. [Ectropothecium galerulata Duby in Besch., Ectropothecium sphaerocarpum (A.Jaeger) Besch., Ectropothecium virentipinnatum Müll.Hal. in Renauld] Without locality, de Robillard as Hypnum galerulatum (Duby 1879: 7, Bescherelle 1880: 321) and Ectropothecium galerulata; leg. Grateloup as Ectropothecium sphaerocarpum (Bescherelle 1880: 321, Renauld 1891:211, 1897); leg. Robillard as Ectropothecium virenti-pinnatum (Renauld 1891:211, 1897); Curepipe, cave near Bassin Anglais (Een 1997). Vesicularia lepervanchei (Besch.) Broth. Le Pouce, forest half way to the top (Een 1997). Warburgiella leptorrhyncha (A.Jaeger) Broth. [Trichosteleum leptorrhynchum (A.Jaeger) Renauld; Rhaphidostegium leptorrhynchum A.Jaeger] Without locality, Bory (Bescherelle 1880: 305, Renauld 1897). Weissia ayresii Schimp. ex Besch. [Hymenostomum ayresii (Schimp. ex Besch.) Broth.; Weissia mauritiana Schimp. ex Besch.] ARCHIVE FOR BRYOLOGY 51 (2009) 22 Frahm, O´Shea, Ho Without locality, leg. Ayres, type of Weissia ayresii; without locality, leg. Ayres, type of Weissia mauritiana Schimp. (Bescherelle 1879: 302, Dixon 1938, Een 1976); Plaine des Roches, leg. Vaughan 1935 (Dixon 1938, Een 1976). Zygodon conoideus var. latifolius Brid. Insula Franciae, Hook. (Bridel 1826: 592). Excluded and doubtful taxa Campylopus nivalis var. nivalis (Brid.) Brid. Een (1993b). The record of this alpine species on Mauritius is very unlikely, not in Frahm (1985). Campylopus pseudobicolor Müll.Hal. ex Renauld & Cardot Een (1993b), probably erroneous, not in Frahm (1985). Calyptrochaeta cristata (Hedw.) Desv. Reported for Mauritius, but the specimen came from Australia (Bridel 1826: 712). Campylopus filifolius (Hornsch.) Mitt. [Campylopus interruptulus Müll.Hal. in Besch.] Without locality, leg. Rodriguez; Curepipe, leg. Rodriguez 1889 (Renauld 1891: 211, 1897). This is a dubious record. The specimen (in PC) undoubtedly is a large representative of this species, however, Campylopus filifolius is a neotropic species and this would be the only record from the Palaeotropics. Records for Madagascar and Réunion are similarly suspected. leg. Robillard (Hb. C. Mueller) (Bescherelle 1879 as C. interruptulus). Cardotiella subappendiculata (Broth.) Vitt (Leiomitrium subappendiculatum (Broth.) Cardot) According to Goffinet (1996: 395), the sole specimen Een 413 reported from Mauritius, initially identified by Vitt (1981), has been proven to be Cardotiella appendiculata instead. Dicranum dichotomum P.Beauv. [Campylopus dichotomus (P.Beauv.) Brid., Dryptodon dichotomus (P.Beauv.) Brid.] This name appears in a list of Rodriguez collections from Mauritius (Renauld 1891: 211), but it is an ambiguous name and thus excluded. Leucobryum juniperoideum (Brid.) Müll.Hal. Reported for Mauritius, but no records are known: the earliest quoted record (Bescherelle 1879: 336) mentions only Réunion, and a more recent distribution list (Crundwell 1972: 4) also has no detail. Mittenothamnium saproadelphus (Broth.) Cardot Although this is indicated by Een (1993), quoting 'Dixon 1929', there is no reference to Mauritius, nor to this species. Pohlia wahlenbergii var. wahlenbergii (F.Weber & D.Mohr) A.L.Andrews Reported for Mauritius, but no records found. Phyllogonium viscosum (P.Beauv.) Mitt. Reported for Mauritius, but no records found. Physcomitrium aubertii Besch. Although reported for Mauritius, the only record probably comes from Tristan da Cunha rather than Mauritius (Bescherelle 1879: 374). Pyrrhobryum spiniforme var. acunhense (Besch.) Manuel. Mauritius? Leg. Peiti-Thouars (Bescherelle 1880: 242). The record of this taxon for Mauritius is probably erroneous: see O'Shea et al. (1997) Prionodon densus (Hedw.) Müll.Hal. [Prionodon rehmannii Mitt.] Doubtful record: Sim (1926) says "known in Mauritius". Syrrhopodon prolifer var. prolifer Schwägr. & Syrrhopodon prolifer var. acanthoneurus (Müll.Hal.) Müll.Hal. The collections recorded for these taxa belong to S. apertifolius (pers. com. L.T. Ellis). ISSN 0945-3466 Moss Flora of Mauritius 23 Acknowledgements We wish to thank C.C. Townsend for biographical data of British collectors and Jens Eggers for support with literature. Philip Sollman determined the Pottiaceae of the collections of the first author References Allen, B., Magill, R.E. 2007. A revision of Orthostichella (Neckeraceae). The Bryologist 110:1– 45. Allen, B.H., Crosby, M.R. & Magill, R.E. 1986. Jaegerina retrosquarrosa, species nova (Pterobryaceae: Musci) with comments on the species of Jaegerina from Mauritius. Cryptogamie: Bryologie, Lichénologie 7: 47–52. Ångström, J. 1873. Förteckning oc beskrifning öfver mossor samlade af Prof. N.J. Andersson onder Fregatten Eugenies verldsomsegling aren 1851–1853. Mossor fran Mauritius samlade I februari 1853. Öfversig af Förhandlingar: Kongl. Svenska Vetenskaps Akademien 30(5): 139–145. Argent, G.C.G. 1969. A Taxonomic Study of West African Mosses. The Families Pterobryaceae and Meteoriaceae. (Ph.D. thesis, Bangor): 1–150. Argent, G.C.G. 1973. A taxonomic study of African Pterobryaceae and Meteoriaceae I. Pterobryaceae. Journal of Bryology 7: 353–378. Arnell, S. 1965. Hepaticae collected by Mr. Gillis Een in Mauritius and Reunion in 1962, Svensk Botanisk Tidsskrift 59: 65–84. Bescherelle, E. 1879. Florule Bryologique de la Réunion, de Maurice et des autres Iles austroafricaines de l'océan indien. Annales des Sciences Naturelles, Botanique, sér. 6, 9: 291– 380 Bescherelle, E. 1880. Florule Bryologique de la Réunion, de Maurice et des autres Iles austroafricaines de l'océan indien (Deuxième partie). Annales des Sciences Naturelles, Botanique, sér. 6, 10: 233–332. Bescherelle, E. 1891. Selectio novorum muscorum. Journal de Botanique (Morot) 5: 142–350. Bizot, M. & Pócs, T. 1974. East African Bryophytes. I. Acta Academiae Paedagogicae Agriensis, Sect. Biol. 12: 383–449. Bizot, M. 1968. Mousses récoltées par Mr. Gillis Een dans les Iles Maurice et de la Réunion. Svensk Botanisk Tidsskrift 62: 471–481. Bizot, M. 1974. Enumeratio muscorum novarum. Revue Bryologique et Lichénologique 40: 101– 138. Bridel, S.E. 1812. Muscologiae recentiorum. Supplementum seu Species Muscorum. Pars II. Gothae: Ex Offic. Libr. Ettingeriana. Bridel-Brideri, S.E. 1826. Bryologia Universa seu systematica ad novam methodam disposito. Vol. 1. Brotherus, V.F. 1901–1909 Andreaeaceae; Bryales. Ed. Engler, A. and Prantl, K., Die natürlichen Pflanzenfamilien. Leipzig. Brotherus, V.F. 1925. Musci (Laubmoose) 2. Hälfte in Engler-Prantl, die Natürlichen Pflanzenfamilien. Leipzig. Buck, W.R. & Ireland, R.R. 1985. A reclassification of the Plagiotheciaceae. Nova Hedwigia 41: 89–125. Buck, W.R. 1979. A revision of the moss genus Trachyphyllum Gepp (Thuidiaceae). Brittonia 31(3): 379–394. ARCHIVE FOR BRYOLOGY 51 (2009) 24 Frahm, O´Shea, Ho Buck, W.R. 1980. A generic revision of the Entodontaceae. Journal of the Hattori Botanical Laboratory 48: 71–159. Câmara, P. E. A. S. 2008. A developmental, phylogenetic and taxonomic study on the moss genus Taxithelium Mitt. (Pylaisiadelphaceae). Unpublished Ph.D. dissertation, University of Missouri-St. Louis, Saint Louis, Missouri. 169pp. Cardot, J. 1904. Les Leucobryacees de Madagascar et des autres iles austro-africaines de l'ocean indien. Bulletin de l'Herbier Boissier, ser. 2, 4: 97–118. Crosby, M.R., Schultze-Motel, U. & W. 1983. Katalog der Laubmoose von Madagaskar und dem umliegenden Inseln. Willdenowia 13: 187–255.(2) Crundwell, A.C. 1972. Leucobryum juniperoideum (Brid.) C.Muell. in Britain. Journal of Bryology 7: 1–6. De Sloover, J.L. 1975. Note de bryologie africaine IV. Breutelia. Bulletin du Jardin Botanique National de Belgique 45: 237–271. De Sloover, J.L. 1982. Homaliodendron scalpellifolium (Mitt.) Fleisch. a l'isle Maurice. Bulletin du Jardin Botanique National de Belgique 52: 245. De Sloover, J.L. 1983. Note de bryologie africaine XII. – Porotrichum et Porothamnium. Bulletin du Jardin Botanique National de Belgique 53: 97–152. De Sloover, J.L. 1986. Note de bryologie africaine XIII. – Polytrichaceae. Bulletin du Jardin Botanique National de Belgique 56: 241–300. Demaret, F. & Potier de la Varde, R. 1951. Les especes africaines de genre Cyclodictyon Mitt. Bull. Jard. Bot. Etat Bruxelles 21: 11–59. Demaret, F. & Potier de la Varde, R. 1952. Les especes africaine du genre Callicostella (C.Muell.) Jaeg. Bulletin du Jardin Botanique de l'État Bruxelles 22: 323–396. Dirkse, G.M., Bruggeman-Nannenga, M.A., Bouman, A.C. 1991. Fissidens papillosus Lac. new to the Canary Islands. Cryptogamie: Bryologie, Lichénologie 12: 451–454. Dixon, H.N. 1938. Mosses collected in Mauritius by Dr. R.E. Vaughan. Revue Bryologique et Lichénologique 10: 121–127. Dorr, L.J. 1997. Plant collectors in Madagascar and the Comoro Islands. Kew. 524 pp. Duby, J.E. 1876. Choix de mousses exotiques nouvelles ou mal connues. Mémoirs de la Société de Physique et d'Histoire Naturelle de Genève 24: 361–374. Duby, J.E. 1877. Diagnoses muscorum novorum. Flora 60: 90–95 Duby, J.E. 1879. Choix de mousses exotiques nouvelles ou mal connues. Mémoirs de la Société de Physique et d'Histoire Naturelle de Genève 26: 1–14. Eddy, A. 1985. A revision of African Sphagnales. Bulletin of the British Museum (Natural History). Botany 12: 77–162.(5) Een, G. 1976. Mosses from the Mascarenes. Lindbergia 3: 213–217. Een, G. 1978. Mosses from the Mascarenes — 2. Lindbergia 4: 219–213. Een, G. 1989. Mosses from the Mascarenes — 3. Tropical Bryology 1: 55–62. Een, G. 1993a. Mosses from the Mascarenes — 4. Tropical Bryology 7: 47–54. Een, G. 1993b. Mosses from Africa 3 sensu Index Muscorum. (Manuscript on disk, dated 22/07/1993). Een, G. 1997. Mosses from the Mascarenes — 6. Tropical Bryology 13: 21–34. Een, G. 2000.Bryophytes collected in Mauritius by Gunnar Erdtman. Tropical Bryology 18: 87– 89. Ellis, L.T. 2003. A revised synonymy for Syrrhopodon trachyphyllus (Calymperaceae, Musci) and some related Old World taxa. Systematics and Biodiversity 1: 159–172. Ellis, L.T. 2005. A revision of some Old World moss taxa in the Syrrhopodon prolifer complex (Musci: Calymperaceae), and a new species from Malawi. Systematics and Biodiversity 3: 159–178. Enroth, J. 1994. A taxonomic monograph of the genus Pinnatella (Neckeraceae, Bryopsida). Acta Botanica Fennica 151: 1–90. ISSN 0945-3466 Moss Flora of Mauritius 25 Frahm, J.-P. 1985. Afrikanische Campylopus-Arten (Dicranaceae, Musci). Bryophytorum Bibliotheca 31: 1–216. Frahm, J.-P., Borchardt, P., Schramm, M., Sonntag, M. 2003. Synopsis der Gattung Rhizogonium (Musci, Rhizogoniaceae). Tropical Bryology 24: 129–137. Fransén, S. 2004. A taxonomic revision of extra-Neotropical Bartramia section Vaginella Müll.Hal. Lindbergia 29: 73–107. Goffinet, B. 1996. Cardotiella elimbata (Thér.) Goffinet, comb. nov., and C. appendiculata new to Mauritius (Musci, Orthotrichaceae). The Bryologist 99: 390–396. Grolle, R. 1995. The Hepaticae and Anthocerotae of the East African Islands. An annotated Catalogue. Bryophytorum Bibliotheca 48, 178 pp. Hyvönen, J. 1989. A synopsis of genus Pogonatum (Polytrichaceae, Musci). Acta Botanica Fennica 138: 1–87. Kanda, H. 1987. Catalog of moss specimens from Antarctica and adjacent regions. National Institute of Polar Research, Tokyo. 186pp. Kimura, M., Seto, K. 2003. Bryoxiphium norvegicum (Bryopsida: Bryoxiphiaceae), newly found in Mauritius. Bulletin of the Osaka Museum of Natural History 57: 33–37. Kruijer, J.D. 1997. British Bryological Society Expedition to Mulanje Mountain, Malawi. 6. Hypopterygiaceae (Musci). Journal of Bryology 19: 515–520. Kruijer, J.D. 2002. Hypopterygiaceae of the World. Blumea, Supplement 13: 1–388. LaFarge, C. 2002. Leucoloma I: A revision of subgenus Leucoloma (Dicranaceae, Bryopsida) in Africa and Madagascar. The Bryologist 105: 509–590. LaFarge, C. 2002. Leucoloma II: A revision of Leucoloma series Holomitrioidea (Dicranaceae). The Bryologist 105: 591–605. LaFarge, C. 2002. Leucoloma III: A species synopsis: typification, synonymy and excluded names. The Bryologist 105: 606–624. Mohamed, M.A.H. 1979. A taxonomic study of Bryum billardieri Schwägr. and related species. Journal of Bryology 10: 401–466. Müller, C. 1851. Synopsis Muscorum Frondosorum omnium hucusque cognitorum. Berolini sumptibus Alb. Foerstner. Müller, C. 1876. Musci Hildebrandtiani in Archipelago Comorensi et in Somalia littoris Africani anno 1875 ab I.M. Hildebrandt lecti. Linnaea 40: 225–326. Müller, F. 2002. Additions to the bryophyte floras of Réunion and Mauritius (East African Islands). Tropical Bryology 21: 47–50. Nishimura, N. 1985. A revision of the genus Ctenidium. Journal of the Hattori Botanical Laboratory 58: 1–82. Ochi, H. 1972. A revision of the African Bryoideae, Musci (first part). Journal of the Faculty of Education, Tottori University 23: 1–126. Orbán, S. 1978. Studies on African Calymperaceae II. Acta Botanica Academicae Scientiarum Hungaricae 24: 113–120. Orbán, S. 1981. Studies in African Calymperaceae III. Conspectus of the African species of Syrrhopodon Schwägr. Acta Botanica Academicae Scientiarum Hungaricae 27: 169–177. O'Shea, B.J. & Ochyra, R. 2000. Families and genera of mosses no longer believed to occur in sub-Saharan Africa. Tropical Bryology 18: 119–127. O'Shea, B.J. 2003. Checklist of the mosses of sub-Saharan Africa. Version 4. Tropical Bryology Research Reports No. 4, 1–185. O'Shea, B.J. 2006. A revision of Radulina W.R.Buck & B.C. Tan (Sematophyllaceae: Bryopsida). Tropical Bryology 27: 25–43. O'Shea, B. J. 2006. Checklist of the mosses of sub-Saharan Africa (version 5, 12/06). Tropical Bryology Research Report 6: 1–252. O'Shea, B.J., Eggers, J., Pursell, R.A., Sollman, P. & Stevenson, C.R. 1997. New bryophyte taxon records for tropical countries. 1. Tropical Bryology 13: 175–183. ARCHIVE FOR BRYOLOGY 51 (2009) 26 Frahm, O´Shea, Ho Pfeiffer, T., Kruijer, J.D., Frey, W. & Stech, M. 2000. Systematics of the Hypopterygium tamarisci complex (Hypopterygiaceae, Bryopsida): implications of molecular and morphological data. Journal of the Hattori Botanical Laboratory 89: 55–70. Pursell, R.A. 1987. A taxonomic revision of Fissidens subgenus Octodiceras (Fissidentaceae). Memoirs of the New York Botanical Garden 45: 639–660. Reese, W.D. 1991. Syrrhopodon armatissimus sp. nov. from Madagascar, with comments on the Calymperaceae from the Madagascar region. The Bryologist 94: 217–220. Renauld, F. 1888. Note sur une collection de mousses de l'ile Maurice. Revue Bryologique 15: 87–90. Renauld, F. 1889. Notice sur une collection de mousse de l'ile Maurice (Suite) (1). Revue Bryologique 16: 81–87. Renauld, F. 1891. Document nouveaux sur les Muscinées des iles Austro-Africaines de l'Océan indien. Revue de Botanique Mensuel 9: 209–229. Renauld, F. 1897. Prodrome de la flore bryologique de Madagascar, des Mascareignes et de Comores. Monaco. Renauld, F. 1909. Supplément au Prodrome de la Flore Bryologique de Madagascar des Mascareignes et des Comores. Supplement 1–139, pl. 1–23. Renauld, F. & Cardot, J. 1915, in Grandidier, A.G. Histoire physique, naturelle et politique de Madagascar (Mousses) 39: 1–560. Salazar Allen, N. 1993. A revision of the pantropical moss genus Leucophanes Brid. Bryophytorum Bibliotheca 46. Sim, T.R. 1926. The Bryophyta of South Africa. Transactions of the Royal Society of South Africa 15: 1–475. Snider, J.A. 1975. A revision of the genus Archidium (Musci). Journal of the Hattori Botanical Laboratory 39: 105–201. Sollman, P. & Een, G. 1996. Mosses from the Mascarenes — 5. Tropical Bryology 12: 1–4. Stech, M. 1999. A reclassification of Dicranaceae (Bryopsida) based on non-coding cpDNA sequence data. Journal of the Hattori Botanical Laboratory 86: 137–159. Thériot, I. 1923[1924]. Reliquiae Renauldianae. Bulletin du Muséum d'Histoire Naturelle 29: 456–461. Tixier, P. 1993. Bryophytes nouveaux pour la science et l'île Maurice. Proceedings of the Royal Society of Arts & Science of Mauritius 5: 49–54. Touw, A. 1962. Revision of the moss-genus Neckeropsis (Neckeraceae) I. Asiatic and Pacific species. Blumea 11: 373–425. Touw, A. 1976. A taxonomic revision of Thuidium, Pelekium and Rauiella in Africa South of the Sahara. Lindbergia 3: 135–195. Touw, A. 1986. A revision of Pogonatum sect. Racelopus, sect. nov., including Racelopus Dozy & Molk., Pseudoracelopus Broth. and Racelopodopsis Thér. Journal of the Hattori Botanical Laboratory 60: 1–33. Vitt, D.H. 1981. The genera Leiomitrium and Cardotiella gen. nova (Orthotrichaceae). Journal of the Hattori Botanical Laboratory 49: 93–113. Wigginton, M.J. 2009. Checklist and distribution of the liverworts and hornworts of sub-Saharan Africa, including the East African Islands (edition 3, January 2009). Tropical Bryology Research Reports 8: 1–114. ISSN 0945-3466