Webbia 67(1): 65-91. 2012
Contribution to the Flora of Central-Southern Dhofar (Sultanate of Oman)
STEFANO MOSTI, MAURO RAFFAELLI, MARCELLO TARDELLI
Centro Studi Erbario Tropicale dell’Università
Via G. La Pira 4 - 50121 Firenze (Italia)
e-mail: stefano.mosti@unifi.it
Accepted 15 March 2012
Contributo alla Flora del Dhofar centro-meridionale (Sultanato dell’Oman) — Dal 2000 al 2009 sono state effettuate
10 missioni scientifiche dedicate all’esplorazione floristica del Dhofar centro meridionale (Oman). Le raccolte hanno
interessato i litorali sabbiosi, la pianura costiera e numerosi wadis che sboccano nell’Oceano Indiano (da Rachyut, Al
Mughsayl, Salalah, Khor Rori, Mirbat, Hadbin fino ad Hasik). Particolare attenzione è stata dedicata all’esplorazione dei
versanti monsonici delle montagne (soprattutto Jabal Qara) dove si concentra la maggiore ricchezza floristica e il maggior numero di taxa endemici. Raccolte sono state effettuate anche su Jabal Samhan, la montagna più alta del Dhofar,
e nell’area pre-desertica di Wadi Andur, alle spalle di Jabal Samhan. Sono state inoltre indagate quelle aree dove ancor
oggi sono presenti i maggiori nuclei di Boswellia sacra, l’albero dell’incenso, come Al Mughsayl, Adownib, Aluf, Uyun,
Wadi Dowkah e Hasik. Sporadiche esplorazioni sono state effettuate nella vasta area pre-desertica che, oltre Thumrayt, si
spinge verso nord fino al confine con l’Arabia Saudita, da Muddy, Al Mathafah, Ginan Bin Nawatish fino a Shisr (l’antica
Ubar), Sayh al Kharyat e Dawka. In totale sono state esaminate 233 stazioni di raccolta che nel loro complesso coprono
dettagliatamente l’area del Dhofar centro-meridionale. L’elenco floristico riporta 606 taxa di cui alcuni sono segnalati
per la prima volta o per l’Oman (17 taxa) o per il Dhofar (13 taxa). Numerose sono le specie endemiche rinvenute (14
endemismi del Dhofar, 35 endemismi regionali: Dhofar/Oman e Yemen) presenti soprattutto sui versanti monsonici
delle montagne costiere.
Key words: Flora, Dhofar, Oman.
Dhofar is a large region of the Sultanate of Oman
bordering Yemen on the west and extending eastward
for about 250 km. The northern boundary of Dhofar
spreads into the Ar Rub al Kali, the Saudi Arabian
desert, while its southern part faces the Arabian Sea.
Some different phytogeographical zones, each one
with its peculiar flora and vegetation, may be detected, as described by Radcliffe-Smith (1980), Miller &
Morris (1988) and Ghazanfar (1992b, 2003). Local
variations of flora and vegetation in Dhofar are the
response to two coexisting main factors: one is the
presence of a chain of mountains parallel to the coast
and extending eastwards for about 250 km from the
Yemen border until the Gulf of Hasik, and the other
are the monsoons.
From June to September during the southwest
monsoon, the mist and clouds full of rain approaching from the Arabian Sea encounter a natural obstacle
that precludes them from going further inland. As
a consequence the steep slopes of the coastal moun-
Centro Studi Erbario Tropicale, Università di Firenze – Pubbl. n. 122.
66
tains facing south (in particular from Jabal Sayk to
Jabal Qara) are covered with a luxuriant vegetation
of woods, shrubs and grassland. On the other side of
the mountains, instead, where the monsoon has little
or no effect, the vegetation is sparse with isolated
trees and shrubs. The northern sides of the coastal
chain slope down gradually to form a stony and arid
highland where many wadis have their source and
penetrate northward into a vast semi-desert level land
interrupted by rounded hillocks. Towards the border
with Saudi Arabia the rocky desert changes gradually into a sand desert with high more or less parallel
dunes where vegetation is scarce or absent.
The floristic exploration of Dhofar
A. Radcliffe-Smith (1980) was the first botanist to
produce a list of Dhofar plants; in 1977 he participated with other researchers to a scientific survey of flora
and fauna of Dhofar, supported by the Government
of Oman. In the eighties A.G. Miller and M. Morris
further enlarged the knowledge of the Dhofar flora
with the publication of an important illustrated work
(Miller & Morris, 1988) in which the authors treat
the plant species not only from the botanical point of
view but also describe the medicinal and economical
local practices.
New progress in the knowledge of the flora of
Oman is due to the work of S. A. Ghazanfar who
produced several monographic researches on critical families, genera and species and interesting studies on biodiversity and conservation. She is also the
author of the first Flora of Oman, a work in progress
of which the first two issues (2003 and 2007) have
been published.
Our researches on the flora of Dhofar
From 2000 to 2009 we carried out 10 scientific
missions to Dhofar; we explored several habitats, different from the ecological and altitudinal point of
view, such as seashores, coastal plains and the distal
parts of several wadis ending in the Arabian Sea (from
Rachyut, Al Mughasayl, Salalah, Khor Rori, Mirbat,
Sadh, Hadbin up to Hasik). In particular, many
plants were collected on the monsoon slopes of the
mountains of central Dhofar (with special reference
S. Mosti , M. Raffaelli, M. Tardelli
to Jabal Qara) and in areas still rich in frankincense
trees (Boswellia sacra): e.g. Al Mughsayl, Wadi Adownib, Aluf, Uyun, Wadi Dowkah and Hasik. We
also collected on the upper part of Jabal Samhan, the
highest mountain of Dhofar and on the semi-desert
area of Wadi Andur (beyond Jabal Samhan), and explored some other semi-desert environments north
to Thumrayt, i.e. Muddy, Al Mathafah, Ginan Bin
Nawatish, with sporadic excursions to Shirs (the ancient Ubar), Sayh al Kharyat and Dawkah.
This program has been carried out in cooperation
with IMTO (Italian Mission to Oman), supervised
by Prof. A. Avanzini (Pisa University) and supported by the Office of the Advisor for Cultural Affairs,
Sultanate of Oman (Muscat and Salalah). The Funds
PRIN (National Interest Research Projects) of the
years 2004 and 2007 and the Tropical Herbarium of
Florence University, offered the necessary financial
support to this research program.
The specimens collected are now conserved in the
Tropical Herbarium of Florence (FT); a complete set
of duplicates (about 350 specimens), mounted on
Herbarium sheets and identified, was presented to
the National Herbarium Natural History Museum
(ON) in Muscat.
Collection sites
The collection sites are grouped according to
their ecological and/or altitudinal similarities and are
marked with the following abbreviations: L = littorals
and beaches, CP = coastal plain and foothills of the
escarpment mountains, LH = low hills and grasslands
up to 500 m on monsoon affected escarpments, HH
= high hills and grasslands between 500 and 800 m on
monsoon affected escarpments, M = mountain slopes
and grassland above 800 m, sW = southwards draining wadis, nW = northwards draining wadis and semidesert areas, Ha = dry (no monsoon affected) hills
near Hasik, Wa = wet areas or mangrove vegetation.
Each abbreviation is followed by a progressive
number, which refers to the respective position in
the territory (from west to east). The geographical
coordinates define each locality and identify the sites
where the plants were collected. The maps of figure
1 and 2 show the sites of collection; when those sites
are very near to each other, they overlap on the maps
and therefore they are indicated in Fig. 1 and 2 with
67
Contribution to the Flora of Central-Southern Dhofar (Sultanate of Oman)
the same abbreviations and the same numbers, but
differentiated by a letter (a, b, c, etc.) both in the list
of the collection sites and the following floristic lists.
The collection sites are 233 in number, 139 of which
are indicated in Fig. 1 and 2. The number of sites explored was large enough to have exhaustive information on the local distribution of plant species.
List of the collection sites
L = littorals and beaches
L1
30.09.01. Rakhyut: beach. Alt. 2 m - 16°44.780’N,
53°25.420’E
L2
a: 24.04.01. Al Mughsayl: beach and rocky outcrops
near the sea-water spray. Alt. 10 m - 16°52.500’N,
53°45.980’E - b: 23.09.01. Al Mughsayl: beach in
front of the restaurant. Alt. 0-5 m - 16°52.740’N,
53°46.500’E - c: 02.09.02 (idem) - d: 10.09.02
(idem).
L3
a: 29.04.01. Khor Rhori: beach. Alt 0-5 m 17°1.830’N, 54°26.370’E - b: 09.09.02 (idem).
L4
17.09.02. Beach 4 km from Mirbat, towards Salalah. Alt. 0-5 m - 17°1.057’N, 54°40.208’E.
L5
a: 09.09.02. Mirbat: beach. Alt. 0-5 m 16°59.262’N, 54°41.193’E - b: 02.03.07 (idem).
L6
05.03.07. Hasik-Hadbin road: beach, 10 km from
Hasik, towards Hadbin. Alt. 0-5 m - 17°21.177’N,
55°17.415’E.
L7
14.03.05. Hasik: beach. Alt. 4 m - 17°26.770’N,
55°16.050’.
L8
a: 15.09.03. Hasik: at the base of the vertical cliffs
in front of the Ocean, 6 km east of the village. Alt.
5-10 m - 17°28.566’N, 55°13.806’E - b: 15.03.05
(idem) - c: 20.02.09 (idem).
CP7
CP8
CP9
CP10
CP11
CP12
CP13
CP14
CP = coastal plain and foothills of the escarpment
mountains
CP1 30.09.01. Woody deep valley behind Rakhyut. Alt.
60 m - 16°45.080’N, 53°24.590’E.
CP2 a: 27.04.01. Salalah-Al Mughsayl road: rocky hilly
slope on the left, 5 km to Al Mughsayl. Alt. 30100 m - 16°53.750’N, 53°48.950’E - b: 23.09.01
(idem) - c: 02.09.02 (idem).
CP3 19.03.04. Raysut-Al Mughsayl road: track to Adownib frankincense area, 2 km inland from the
crossroads. Alt. 80 m - 16°54.970’N, 53°50.420’E.
CP4 a: 04.10.01. Raysut: rocky coastal plain and grasslands on the hillocks west of the harbour. Alt. 10-30
m - 16°55.695’N, 53°58.650’E - b: 04.09.02 (idem
- 16°55.550’N, 53°59.150’E).
CP5 a: 13.09.03. Al Balid: inside the archaeological area:
sandy brackish ground. Alt. 10 m - 17°0.240’N,
54°7.720’E - b: 14.02.06 (idem - 17°0.315’N,
54°7.900’E).
CP6 a: 21.09.01. Salalah-Taqah road: hills near Ain Razat. Alt. 40-100 m - 17°7.580’N, 54°14.400’ - b:
CP15
CP16
CP17
CP18
CP19
CP20
07.09.02 (idem) - c: 24.03.04 (idem). - d: 28.02.09
(idem).
24.09.01. Salalah-Taqah road: at the crossroads to Ayn Hamran. Alt. 50 m - 17°3.330’N,
54°16.540’E.
a: 21.10.00. Khor Rori: rocky coastal plain between Sumhuram excavation and the Wadi Darbat
bed. Alt. 4-30 m - 17°2.730’N, 54°25.720’E - b:
11.09.03 (idem - 17°2.860’N, 54°26.470’E) - c:
13.09.03 (idem - 17°2.960N, 54°26.330’E) - d:
18.03.04 (idem).
21.10.00. Khor Rori: inside the archaeological site of
Sumhuram. Alt. 20 m - 17°2.330’N, 54°26.060’E
a: 11.09.03. Khor Rori: top of the hill on the left
of the ancient harbour. Alt. 20-25 m - 17°1.610’N,
54°26.620’E - b: 18.03.04 (idem).
20.03.04. Wadi Darbat: Ficus and Ziziphus thickets.
Alt 95 m - 17°5.240’N, 54°26.010’E.
22.04.01. Taqah-Mirbat road: rocky and sandy
coast, 9 km from Khor Rori. Alt. 5-10 m 17°2.020’N, 54°31.410’E - b: 29.09.01(idem).
29.09.01. Taqah-Mirbat road: near the first crossroads to Tawi Atayr. Alt. 50 m - 17°3.240N,
54°25.982’E.
12.02.06. Rocky hillocks between Mirbat and
Sadh, 11 km from Mirbat. Alt. 56 m - 17°1.220’N,
54°46.130’E.
a: 14.03.04. Mirbat-Sadh road: rocky hillocks with
Boswellia sacra, 5 km in the inland of Sadh. Alt.
70-87 m - 17°5. 640’N, 55°3.470’E - b: 22.03.04
(idem - 17°5.520’N, 55°3.550’E) - c: 18.03.05
(idem - 17°5.420’N, 55°3.700’E) - d: 20.02.09
(idem - 17°5.634’N, 55°3.479’E).
12.02.06. Rocky hillocks along the road between
Sadh and Hadbin, 13 km from Sadh. Alt. 13 m 17°7.410’N, 55°8.290’E.
26.04.01. Sadh-Hadbin track: 16 km from Sadh,
rocky plain. Alt. 60 m - 17°9.100’N, 55°8.720’E.
26.04.01. Rocky plain near Hadbin. Alt. 10 m 17°11.840’N, 55°13.140’E.
12.02.06. Hadbin-Hasik road: 1,8 km from
Hadbin, rocky cliffs at the roadside.Alt. 5 m 17°12.160’N, 55°13.860’E.
12.02.06. Hadbin-Hasik road: 10 km from Hadbin, rocky ground and cliffs at the roadside. Alt. 13
m - 17°16.270’N, 55°14.440’E.
68
S. Mosti , M. Raffaelli, M. Tardelli
Fig. 1 – Top: Central Southern Dhofar: the study area and the collection sites. Bottom: Collection sites in North
West Dhofar.
Contribution to the Flora of Central-Southern Dhofar (Sultanate of Oman)
Fig. 2 – Top: Collection sites in West Dhofar. Bottom: Collection sites in East Dhofar.
69
70
CP21 12.02.06. Hadbin-Hasik road: 15 km from Hadbin, rocky ground and cliffs at the roadside. Alt. 18
m - 17°19.260’N, 55°16.250’E.
LH: low hills and grasslands up to 500 m on monsoon
affected escarpments
LH1 21.04.01. Al Mughsayl-Ajdarawat road: rocky hilly
slopes facing the Ocean, 8 km from Al Mughsayl.
Alt. 500 m - 16°51.154’N, 53°43.089’E.
LH2 23.09.01. Grasslands, 3 km before Al Mughsayl
coming from Salalah. Alt. 200 m - 16°53.630’N,
53°45.600’E.
LH3 a: 26.09.01. Salalah-Thumrayt road: hilly slopes
beyond Salalah plain. Alt. 215 m - 17°6.140’N,
54°4.300’E - b: 02.03.07 (idem. Alt. 296 m 17°6.397’N, 54°4.053’E).
LH4 02.03.07. Hilly slopes above Salalah: Ain Garziz
valley. Alt. 172 m - 17°6.615’N, 54°4.767’E.
LH5 02.10.01. Grasslands on the foothills between
Salalah and Taqah. Alt. 160 m - 17°5.280’N,
54°19.380’E.
LH6 02.10.01. Hills between Salalah and Taqah. Alt.
330-550 m - 17°6.500’N, 54°18.570’E.
LH7 a: 02.10.01. Salalah-Taqah road: deep valley to
Athom Spring, Anogeissus dhofarica woods. Alt. 170310 m - 17°6.910’N, 54°21.860’E - b: 11.09.02
(idem. Alt. 170-300 m - 17°6.290’-54°21.130’E)
– c: 24.03.04 (idem. Alt 230 m - 17°6.830’N,
54°21.840’E) - d: 24.03.04 (idem: at the end of the
valley, near the spring. Alt. 235 m - 17°6.940’N,
54°21.800’E) - e: 13.02.06 (idem - 17°5.770’N,
54°20.380’E) - f: 13.02.06 (idem. Alt. 244 m 17°6.840’N, 54°21.840’E).
LH8 a: 11.09.03. Wadi Darbat: scrubland near the highter edge of the waterfall. Alt. 140 m - 17°4.200’N,
54°25.200’E - b: 13.09.03 (idem).
LH9 29.04.01. Wadi Darbat: above the waterfall, inside the valley along the wadi banks. Alt. 180 m 17°6.450’N, 54°27.210’.
LH10 06.09.02. Grasslands along the road to Tawi Atayr.
Alt. 400 m - 17°6.280’N, 54°29.660’E.
LH11 01.09.02. Salalah-Mirbat road: foothills on the
right, 7 km from the crossroads to Khor Rori. Alt.
130-150 m - 17°3.420’N, 54°30.020’E.
LH12 02.10.01. Taqah-Mirbat road: 13 km from Taqah,
along the track to Jabal as Shawr. Alt. 103-205 m 17°3.350’N, 54°31.510’E.
LH13 22.04.01. From Tawi Atayr, going down to the
Salalah-Mirbat road. Alt. 450 m - 17°3.900’N,
54°33.040’E
LH14 a: Salalah-Tawi Atayr road: small valley with baobab
trees, near Wadi Hinna. Alt. 330 m - 17°3.200’N,
54°36.540’E - b: 11.09.02 (idem).
S. Mosti , M. Raffaelli, M. Tardelli
LH15 16.09.02. Mirbat-Sadh road: 15 km from Mirbat,
climbing along the lower course of Wadi Ayn Hilf for
about 5 km. Alt. 50-400 m - 17°5.115’N, 54°49.460’E.
LH16 26.04.01. Mirbat-Sadh track: rocky slope 22 km to
Sadh. Alt. 125 m -17°3.850’N, 54°51.690’E.
LH17 26.04.01. Wadi banks along the track between
Mirbat and Sadh, 16 km to Sadh. Alt 305 m 17°5.100’N, 54°57.220’E.
HH: high hills and grasslands between 500- 800 m on
monsoon affected escarpments
HH1 30.09.01. From Rakhjut to the road n.° 47 to
Ajdarawat. Thick woodland. Alt. 400-800 m 16°48.200’N, 53°20.400’E.
HH2 30.09.01 Rakhiut-Ajdarawt road: along the track
to Ajdarawt, 8 Km from Rakhyut. Alt. 568 m 16°44.480’N, 53°22.040’E.
HH3 21.04.01. Salalah- Ajdarawt road: 13 km to
Ajdarawt. Alt. 600 m - 16°49.792’N, 53°39. 587’E.
HH4 a: 23.09.01. Al Mughsayl-Ajdarawt road: 11 km
from Al Mughsayl, going up towards Ajdarawt, rocky
slopes facing the Ocean. Alt. 568 m - 16°50.848’N,
53°41.650’E - b: 10.09.02 (idem. Alt. 600-900
m) - c: 27.02.07 (idem. Alt. 700 m) - d: 15.02.06
(idem. Alt. 476- 965 m - 16°51.305’-16°49.785’N,
53°43.275’- 53°39.690E) - e: 21.02.09 (idem).
HH5 a: 28.04.01. Salalah-Ghadow road: grassland and
scrubland 3 km to Ghadow, coming from Salalah.
Alt. 630 m - 17°6.780’N, 54°1.630’E - b: 26.09.01
(idem) - c: 26.09.01 (idem, 2 km to Ghadow. Alt.
700 m - 17°7.180’N, 54°0.330’E).
HH6 07.09.02. Grasslands along the road to Ashanhaib,
coming from Ain Razat. Alt. 600-700 m 17°15.030’N, 54°18.050’E.
HH7 22.04.01. Taqah-Tawi Atayr road: climbing to Tawi
Atayr, at the roadside - 17°7.130’N, 54°29.740’E
HH8 25.09.01. Sinkhole near Tawi Atayr. Alt 650 m 17°6.800’N, 54°33.550’E.
HH9 22.09.01. Salalah-Mirbat road: along the road to Tawi
Atayr (3° crossroads). Thick woodland on the top of
the hill. Alt. 600 m - 17°3.100’N, 54°36.400’E.
HH10 06.09.02. Salalah-Mirbat road: along the road to
Tawi Atayr (3° crossroads). Climbing from the
“baobab” valley to the plateau of Tawi Atayr. Alt.
500-700 m - 17°4.300’N, 54°36.580’E.
HH11 17.09.02. Mirbat-Sadh road, 15 km from Mirbat,
climbing along the bed of Wadi Ayn Hilf for 10 km,
up to the spring below Jabal Samhan. Alt. 500-600
m - 17°7.040’N, 54°48.120’E.
M: mountains slopes and grasslands above 800 m
M1 30.09.01. Al Mugsayl-Ajdarawat road: on the plateau along the road n.°47, near Ajdarawt. Alt. 1064
Contribution to the Flora of Central-Southern Dhofar (Sultanate of Oman)
M2
M3
M4
M5
M6
M7
M8
M9
M10
M11
M12
M13
M14
M15
m - 16°47.780’N, 53°33.320’E.
a: 10.09.02. Al Mugsayl-Ajdarawat road: 14 km
from Al Mughsayl, along the road n.°47, mountain
slope facing the Ocean, in front of the police station,
Dracaena scrubland. Alt. 965 m - 16°49.785’N,
53°40.006’E - b: 27.02.07 (idem).
13.09.02. Al Mugsayl-Ajdarawat road: grassland
and scrubland on the mountain slope facing the
Ocean, 15 km from Al Mughsayl. Alt. 800-900 m 16°49.690’N, 53°40.119’E.
21.04.01. Al Mughsayl -Ajdarawat road: 7 km from
Al-Mughsayl, rocky steep slope at the left roadside,
going up to the escarpment facing the Ocean. Alt.
1100 m - 16°51. 512’N, 53°43.381’E.
01.03.07. Qaftawt-Wanat road: along the mountain ridge, 16 km from Qaftwat, going down 4 km
towards the coast facing the Ocean. Alt. 1129 m 17°0.704’N, 53°44.767’E.
01.03.07. Qaftawt-Wanat road: along the mountain ridge, 16 km from Qaftwat, going down 2 km
towards the coast facing the Ocean. Alt. 1059 m 17°1.912’N, 53°44.960’E.
01.03.07. Qaftawt-Wanat road: along the mountain
ridge, 16 km from Qaftwat, going down 600-700 m
towards the coast facing the Ocean. Alt. 1063 m 17°1.990’N, 53°45.385’E.
01.03.07. Qaftawt-Wanat road: along the mountain ridge, 5 km from Ar Raydah. Alt 1237-1172 m
- 17°3.219’-17°2.587’N, 53°48.037’-53°46.618’E.
01.03.07. Salalah-Qaftawt road: 3 km from Qaftawt,
towards Ar Raydah. Alt. 1078 m - 17°5.223’N,
53°52.025’E.
a: 01.03.07. Teetaam-Gattawt road: going down
4 km from the mountain ridge. Alt. 933 m 17°4.169’N, 53°53.643’E - b: 18.02.06 (idem. Alt.
916 m - 17°4.166’N, 53°53.641’E).
a: 28.04.01. On the top of the plateau between
Ghadow and Teetaam, Olea woodland. Alt. 976
m - 17°8.170’N, 53°57.860’E - b: 01.10.01 (idem.
Alt. 986 m) - c: 19.03.05 (idem. Alt. 936-990 m 17°8.180’N, 53°57.890’E) - d: 10.02.06 (idem).
23.04.01. Teetaam- Ghadow road: on the ridge between the mountain escarpment and the pre-desert.
Alt. 810 m - 17°7.190’N, 53°57.930’E.
30.04.01. Grassland and scrubland between Ghadow and Teetaam, 6 km from Ghadow. Alt. 850 m
- 17°7.230’N, 53°58.640’E.
26.09.01. Deep woody valley before Ghadow,
coming from Salalah. Alt. 910 m - 17°7.260’N,
53°58.640’E.
08.09.02. Grassland on the plateau near Ghadow,
coming from Salalah. Alt. 810 m - 17°7.520’N,
53°59.860’E.
71
M16 28.02.07. Salalah-Thumrayt road: near the
crossroads to Zayk. Alt. 864 m. 17°15.447’N,
54°5.018’E.
M17 27.09.01. Grassland between Ashanhaib and
Zayk, 6 km to Zayk. Alt. 850 m - 17°16.910’N,
54°9.880’E.
M18 27.09.01. Rocky grassland on the plateau of Jibjat.
Alt. 950 m - 17°14.170’, 54°26.530’E.
M19 25.09.01. Grassland above Tawi Atayr. Alt. 920 m 17°7.150’N, 54°35.960’E.
M20 27.09.01. Climbing to Tawi Atayr, coming
from Wadi Hinna. Alt. 906 m - 17°3.920’N,
54°36.110’E.
M21 25.09.01. Track between Tawi Atayr and Jabal
Samhan, near the drinking trough. Alt. 950 m 17°6.810’N, 54°38.210’E.
M22 25.09.01. Jabal Samhan, going up from Tawi Atayr
to the radio station. Alt. 1214 m - 17°6.160N,
54°41.900E
M23 a: 25.09.01. Jabal Samhan: rocky plateau, going up
from the radio station (1214 m) to the mountain
ridge (1425 m) - 17°7.860’N, 54°44.000’E - b:
06.09.02 (idem. Alt. 1214-1425 m - 17°6.160’17°7.860N, 54°41.890-54°43.990E - c: 27.02.09
(idem - 17°6.158’-17°8.407’N, 54°41.885’54°45.456’E).
sW: south-draining wadis
sW1 a: 21.04.01. Al Mughsayl-Ajdarawat road: 7 km
from Al Mughsayl: Wadi Afal e W. Mughsayl (at the
end of the down going road): Boswellia sacra scrubland. Alt. 100 m - 16°52.024’N, 53°42.965’E - b:
27.04.01 (idem) - c: 23.09.01 (idem) - d: 10.09.02
(idem. Alt. 50-100 m - 16°53.800’N, 53°43.400’E)
- e: 10.09.02 (idem) - f: 19.03.04 (idem. Alt. 70
m - 16°51.900’N, 53°42.889’) - g: 11.03.05 (idem.
Alt. 30-100 m.) - h: 15.02.06 (idem. Alt 111 m 16°52.096’N, 53°42.932’E) - i: 27.02.07 (idem 16°52.103’N, 53°43.890’E).
sW2 a: 24.04.01. Al Mughsayl: from the lagoon going up
7 km along Wadi Ashawq. Alt. 70 m. 16°55.250’N,
53°44.320’E - b: 02.09.02 (idem. Alt. 50 m 16°55.240’N, 53°44.350’E).
sW3 19.03.04. Al Mughsayl: from the lagoon going up
1,7 km along Wadi Ashawq up to the pool of standing water and inside the left branch of the wadi. Alt.
50 m. 16°53.760’N, 53°46.390’E.
sW4 24.04.01. Al Mughsayl: right tributary of Wadi
Ashawq. Alt. 50 m - 16°53.880’N, 53°45.960’E.
sW5 02.09.02. Al Mughsayl: Wadi Ashawq: along the
right bank, 2,5 km from the crossroads near the
Al Mughsayl lagoon. Alt. 36 m - 16°53.840’N,
53°46.350’E.
72
sW6 a: 22.10.00. Salalah-Al Mughsayl road: rocky hills
of Adownib frankincense area (Boswellia sacra scrubland). Alt. 200 - 16°56.781N, 53°46.450’E - b:
05.09.02 (idem) - c: 11.03.05 (idem. Alt. 180
m - 16°56.910’N, 53°46.410’E) - d: 11.03.05
(idem. Alt 198 m, on a rocky slope - 16°57.190’N,
53°46.860’E) - e: 17.02.06 (idem. Alt. 161 m,
at the bottom of a little valley - 16°56.784’N,
53°46.431’E) - f: 03.03.07 (idem. Alt. 187 m)
sW7 03.03.07. From Adownib frankincense area going
up 3 km to the hilly slopes of Wadi Adownib. Alt.
371 m - 16°58.795’N, 53°46.655’E.
sW8 17.02.06. Wadi to the west of Salalah, 5 km behind
Raysut. Alt. 36 m - 16°59.568’N, 53°57.954’E.
sW9 16.09.03. Wadi to the west of Salalah, 4 km behind
Raysut. Alt. 30 m - 16°58.670’N, 53°58.500’E.
sW10 a: 09.02.06. Khor Rori: along the Wadi Darbat, between the excavations of Sumhuram and
the rocky slopes of the right bank. Alt. 5-16 m 17°3.080’-17°2.720’N, 54°25.530’- 54°25.730’E
- b: 26.02.07 (idem).
sW11 29.09.01. Khor Rori: coastal plain and bed of Wadi
Darbat, on the right of the excavations of Sumhuram. Alt. 5 m - 17°2.680’N, 54°25.740’E.
sW12 a: 01.09.02. Salalah-Khor Rori road: dry bed of
Wadi Darbat, on the right of the road, before the
crossroads to Khor Rori. Alt. 10 m -17°3.160’N,
54°25.600’E - b: 09.02.06 (idem. Alt. 13 m 17°3.080’, 54°25.530’E).
sW13 a: 20.10.00 Khor Rori-Mirbat road: wadi bed and
coastal plain 9 km east of Khor Rori towards Mirbat.
Alt. 1-10 m - 17°2.020’N, 54°31.326E - b: 01.09.02
(idem, Alt. 25 m - 17°2.220’N, 54°31.300’E - c:
26.02.07 (idem) - 17°2.012’N, 54°31.514’E) - d:
26.02.07 (idem. Alt. 1-10 m) 17°2.214’N, 54°31.
514’E.
sW14 22.04.01. Salalah-Mirbat road: wadi between
Khor Rhori and Mirbat, 15 km from Khor Rori.
17°2.460’N, 54°34.610’E.
sW15 22.04.01. Khor Rori-Mirbat road: bed of Wadi Hinna, 18 km from Khor Rori. Alt. 15 m - 17°2.053’N,
54°36.580’E.
sW16 a: 22.09.01. Khor Rori-Mirbat road: going up to
Tawi Atayr, baobab trees in the bed of Wadi Hinna. Alt 100 m - 17°2.440’N. 54°36.660’E) - b:
20.03.04. Wadi Hinna: Adenium scrubland on
the left bank of the wadi - Alt. 45 m - c: 13.03.05
(idem. Alt. 33 m) - d: 13.03.05 (idem. Alt. 40 m)
- e: 09.02.06 (idem) - f: 19.02.09 (idem. Alt. 18 m
-17°2.296’N. 54°36.430’E.
sW17 12.02.06. Wadi along the road Sadh-Hadbin, 5 km
to Hadbin. Alt. 50 m - 17°11.080’N, 55°10.360’E.
sW18 a: 12.02.06. Wadi along the road Hadbin-Hasik,
S. Mosti , M. Raffaelli, M. Tardelli
7 km from Hadbin. Alt. 18 m - 17°14.590’N,
55°15.640’E - b: 05.03.07 (idem) - c: 20.02.09
(idem).
sW19 18.03.05. Wadi along the road Hadbin-Hasik, 16
km from Hadbin. Alt. 20-100 m - 17°19.360’N,
55°16.690’E.
sW20 18.03.05. Wadi along the road Hadbin-Hasik, 20
km from Hadbin. Alt. 20-90 m - 17°21.940’N,
55°17.490’E.
sW21 12.02.06. Wadi along the road Habdin-Hasik,
10 Km to Hasik. Alt. 25 m. - 17°22.090’N,
55°17.530’E.
nW: north-draining wadis and semi-desert areas
nW1 13.09.02. Aybut-Hayrun road: semi-desert plateau
with rocky hillocks and gullies near Hayrun. Alt.
840 m - 17°4.928’N, 53°21.298’E.
nW2 13.09.02. Thumrayt-Mudday road: bed and banks
of Wadi Banar, 60 km from Thumrayt. Alt. 450 m
- 17°34.500’N, 53°28.500’E.
nW3 25.02.09. Thumrayt-Mudday road: bed of Wadi
Ghadur, 50 km from Thumrayt. Alt. 499m 17°37.857’N, 53°33.526’E.
nW4 28.09.01. Semi-desert area near the ancient citadel of Ubar (Shisr). Alt. 350 m - 18°15.230’N,
53°38.850’E
nW5 28.09.01. Semi-desert area near Ginan Bin Nawatish. Alt. 360 m - 18°4.687’N, 53°39.845’E.
nW6 a: 23.04.01. Uyun pool, wadi with permanent water. Alt 730 m - 17°14.800’N, 53°53.330’E - b:
01.10.01 (idem) - c: 19.03.05 (idem. Alt. 700 m
- 17°14.790’N, 53°53.320’E) - d: 10.02.06 (idem.
Alt. 697 m - 17°14.780’N, 53°53.320’E.
nW7 13.09.02. Thumrayt-Al Mathafah road: semi-desert
plateau, 14 km from Thumrayt. Alt. 400-450 m 17°39.055’N, 53°54.094’E.
nW8 30.04.01. Wadi between Ghadow and Teetan. Alt.
850 m. - 17°7.230’N, 53°58.640’E.
nW9 23.04.01. Salalah-Thumrayt road: along the track
to Haluf, rocky plateau, 1.7 km before Haluf. Alt.
700 m - 17°21.400’N, 53°58.880’E.
nW10 08.09.02. Salalah-Thumrayt road: along the
track Haluf-Uyun, rocky plateau. Alt. 600 m 17°22.310’-17°19.690’N, 54°0.480’-53°59.040’E.
nW11 10.02.06. Rocky slopes on the plateau Haluf-Uyun.
Alt. 718 m - 17°19.670’N, 53°59.050’E.
nW12 23.04.01. Salalah-Thumrayt road: semi-desert plain
20 Km before Thumrayt. Alt. 580 m - 17°28.178’N,
54°02.996’E.
nW13 12.09.02. Thumrayt-Dawkah road: semi-desert
plain along the road, 29 km to Dawkah. Alt. 220
m - 18°24.316’N, 54°3.740’E.
nW14 a: 23.10.00. Salalah-Thumrayt road: 42 km from
Contribution to the Flora of Central-Southern Dhofar (Sultanate of Oman)
Salalah, Wadi Dowkah Frankincense Park (numerous old Boswellia plants). Alt. 650-680 m 17°20.080’N, 54°4.500’E - b: 23.04.01 (idem) - c:
28.09.01 (idem) - d: 08.09.02 (idem - 17°20.170’17°20.080’N, 54°4.450’-54°4.500’E) - e:
14.09.02 (idem) - f: 12.09.02 (idem - 17°19.600’17°21.500’N, 54°3.010’- 54°5.000’E) - g: 10.09.03
(idem) - h: 12.09.03 (idem) - i: 21.03.04 (idem)
- l: 25.03.04 (idem - 17°20.010’N, 54°44.680’E)
- m: 12.03.05 (idem) - n: 11.02.06 (idem, inside
the transplanting area of the Park. 17.20.334’N,
54°.4.528’E) - o: 28.02.07 (idem - 17°20.085’N,
54°4.500’E)
nW15 10.02.06. North boundary of Wadi Dowkah Park.
Alt. 649 m - 17°21.200’N, 54°3.210’E.
nW16 04.03.07. 5 Km north of Wadi Dowkah Park, towards Thumrayt: rocky plain 1 km east of the road.
Alt. 630 m. - 17°23.018’N, 54°4.548’E.
nW17 23.10.00. Salalah-Thumrayt road: wadi bed and
rocky slopes 25 km north of Salalah. Alt. 700 m 17°13.060’N, 54°5.580’E.
nW18 a: 27.09.01 Salalah-Thumrayt road: gullies near
Aqabat al Hatab, 35 km from Salalah, rocky
slopes 3,5 km from the police station. Alt. 743 m
- 17°17.570’N, 54°5.200’E - b: 28.09.01 (idem)
- c: 03.09.02 (idem. Alt. 720 m - 17°17.600’N,
54°5.200’E) - d: 08.09.02 (idem) - e: 14.09.02
(idem. Alt. 730 m - 17°17.580’N, 54°5.210’E).
nW19 a: 23.04.01. Salalah-Thumrayt road: gullies near
Aqabat al Hatab, 33 km from Salalah, rocky slopes
2 km further on the police station. Alt. 700 m.
17°17.053’N, 54°5.610’E - b: 21.03.04 (idem. Alt.
715 m - 17°17.130’N, 54°5.490’E) – c: 11.02.06
(idem. Alt. 730 m - 17°17.120’N, 54°5.490’E).
nW20 a: 11.02.06. Salalah-Thumrayt road: 6 km northeast of Wadi Dowkah Park: rocky plain and hillocks near the ancient frankincense store at Wadi
Sahnut. Alt. 616-629 m - 17°23.080’N, 54°6.270’
- b: 04.03.07 (idem. Alt 617 m - 17°22.946’N,
54°6.291’E).
nW21 04.03.07. Salalah-Thumrayt road: 4 km north-east
of Wadi Dowkah Park, along the bed of Wadi Sahnut. Alt. 642 m - 17°21.759’N, 54°6.562’E.
nW22 23.10.00. Wadi along the track Thumrayt-Rabkut,
16 km south of Rabkut. Alt. 578 m - 17°24.400’N,
54°16.740’E.
nW23 14.09.03. Wadi Dhahabun: along the track to Jibjat,
17 km from Dhahabun. Alt. 600 m - 17°26.240’N,
54°21.890’E.
nW24 14.02.06. Wadi along the track Jibjat- Barbazum,
2 km before Jibjat. Alt. 942 m - 17°15.010’N,
54°30.876’E.
nW25 a: 14.09.03. From Barbazum to Wadi Andur, near
73
the pool of standing water of Wadi Andur. Alt.
500 m - 17°34.190’N, 54°42.230’E - b: 17.03.05
(idem).
nW26 17.03.05. Wadi Andur: rocky hills along a tributary
wadi of Wadi Andur, southwards for 2 km from the
pool of standing permanent water (nW25). Alt. 500
m ca. 17°33.630’N, 54°42.700’E.
nW27 14.02.06. Wadi Andur: rocky bed of a tributary
wadi of Wadi Andur, southwards for about 10
km from the pool of standing water(nW25a) . Alt
558 m - 17°34.224’-17°31.085’N, 54°42.171’54°45.050’E.
nW28 14.09.03. From the pool of standing water of Wadi
Andur (nW25a) for about 22 km to the northern
slopes of Jabal Samhan. Alt. 750 m - 17°24.342’N,
54°48.137’E.
Ha: dry (no monsoon affected) hills near Hasik (east
Dhofar)
Ha1 a: 15.03.05. Hadbin-Hasik road: rocky hills with
Boswellia sacra, 3 km before Hasik. Alt. 100-500 m 17°24.550’-17°24.860’N, 55°16.150’-55°16.830’E
- b: 16.02.06 (idem. Alt. 56-400 m - 17°24.710’17°24’870”N, 55°16.940’- 55°16’530”E) - c:
05.03.07 (idem. Alt. 10-350 m - 17°24.870’17°24.710’N, 55°16.530’-55°15.940’E).
Ha2 23.03.04. Hasik: rocky hills with Boswellia sacra, 9
Km east of the village. Alt. 100-475 m - 17°29.680’17°28.962’N, 55°12.345’-55°12.610’E.
Wa: wet areas or mangrove vegetation
Wa1 a: 04.10.01. Salalah: along the road to Raysut,
mangrove vegetation of Khawar Al Qurm Al Saghir
(near the Hilton Hotel) Alt. 0-3 m - 16°58.840’N,
54°0.784’E - b: 05.09.02 (idem).
Wa2 28.09.01. Thumrayt-Dawkah road: 1 km to
Dawkak, near a salty pool. Alt. 237 m - 18°39.610’N,
54°4.390’
Wa3 23.10.00. Wet area along the track Thumrayt-Dhahabun. Alt. 520 m - 17°30.014’N, 54°11.004’E.
Wa4 07.09.02. Salalah-Mirbat road: spring of Ain Razat.
Alt. 80 m - 17°7.800’N, 54°14.200’E.
Wa5 13.02.06. Mangrove vegetation along the SalalahKhor Rori road, 14 km from Salalah. Alt. 1-3 m
- 17°2.050’N, 54°17.020’E.
Wa6 23.10.00. Rocky semi-desert plateau north of
Ashanhaib: spring and wet area at the base of a rocky
cliff. Alt. 840 m - 17°18.150’N, 54°17.220’E.
Wa7 24.09.01. Wet area at Sahalnawt, near Salalah 17°8.182’N, 54°18.702’E
Wa8 02.10.01. Salalah-Mirbat road: track to Tawi Atayr
(3° crossroads): wet area inside the baobab valley of
Wadi Hinna. Alt. 330. - 17°3.216’N, 54°36.591’E.
74
S. Mosti , M. Raffaelli, M. Tardelli
FLORISTIC LIST
The list is subdivided in Pteridophyta, Gymnospermae, Angiospermae (Dicotyledones and Monocotyledones). To facilitate the consultation, families, genera,
species and subspecies are arranged in alphabetical order.
Abbreviations of the names of authors of plant names are
standardized according to Brummit & Powell (1992).
PTERIDOPHYTA
ADIANTHACEAE
Adiantum capillus-veneris L. - LH14a; HH11.
Adiantum philippense L. - LH(7b, 12b); HH(1, 10).
Negripteris scioana (Chiov.) Pic.-Serm. - L2d; CP6a;
LH3a; HH9; M14; sW(6b, 11); nW18d.
HYPODEMATIACEAE
Hypodematium crenatum (Forssk.) Kuhn - CP1; M(14, 19).
PARKERIACEAE
Ceratopteris cornuta (P.Beauv.) Le Prieur - LH14a; Wa7.
PTERIDACEAE
Pteris vittata L. - LH14a; HH11.
SELAGINELLACEAE
Selaginella imbricata (Forssk.) Spring ex Decne. - M2a.
GYMNOSPERMAE
EPHEDRACEAE
Ephedra foliata Boiss. ex C.A.Mey - nW(18d, 19c, 21);
Wa6.
ANGIOSPERMAE (DICOTYLEDONES-MAGNOLIOPSIDA)
ACANTHACEAE
Asystasia gangetica (L.) T.Anderson - Wa8.
Barleria acanthoides Vahl - LH17; sW(2b, 9).
Barleria hochstetteri Nees ex DC. - CP(2b, 6a); LH(9,
12, 14).
Barleria parviflora R.Br. ex T.Anderson - HH4b; M1;
sW(1d, 1e); Ha1a - First record from Oman.
Barleria proxima Lindau - L2d; CP(2b, 8b, 6a); HH(4b,
4e); nW(17, 18a, 18c); M23b; nW14m; Wa6.
Barleria almughsaylensis Mosti, Raffaelli & Tardelli CP2a; HH4e; M2b; sW1h (locus classicus) - Endemic to
Dhofar (Mosti et al., 2011). - (Fig. 3d).
Bentia fruticulosa Rolfe - CP(6a, 11); HH11 - Regional
endemic: Dhofar & Yemen (Hadramaut) (Miller &
Morris, 1988).
Blepharis dhofarensis A.G.Mill. - CP(2a, 2b); HH(4e, 8);
M13 - Regional endemic: Dhofar & SE Yemen (Thulin
et al., 2001).
Blepharis scindica T.Anderson - L5a; CP(4b, 10a, 14);
LH(11, 16); sW(2a, 2b, 6a, 6b); nW(14d, 14e, 25a,
25b, 27, 28); Ha1a - Syn.: B. linariifolia Pers.
Dyschoriste dalyi A.G.Mill. - HH(4b, 5a, 6, 9); M(1,
11b, 18, 17, 23b); nW(14d, 18d) - Endemic to Dhofar
(Miller & Morris, 1988; Ghazanfar, 1992a).
Hypoestes forskaolei (Vahl) Solander ex Roem. & Schult.
- LH7f; HH5c; M(1, 11a, 13).
Justicia calyculata Deflers - CP(6a, 11); HH11 - Syn.:
Bentia fruticulosa Rolfe.
Justicia diffusa Willd. - LH7f; HH(5c, 6, 11); M(3, 23b);
sW(2b, 6a, 6f ); nW(14i, 18b) - (Thulin et al., 2001:
also Yemen).
Justicia heterocarpa T.Anderson - LH3a; M18; Wa7.
Justicia odora (Forssk.) Lam. - HH11; M(14, 23b).
Lepidagathis calycina Hochst. ex Nees - LH(11, 14);
HH4c.
Megalochlamys violacea (Vahl) Vollesen - CP(2b, 2c);
LH(3a, 6, 8b, 14); M1; sW1a; nW18b; M1 - Syn.:
Ecbolium violaceum (Vahl) Hillc. & J.R.Wood.
Monechma debile (Forssk.) Nees - LH(5, 12, 14); sW16a
- Syn.: Justicia debilis (Forssk.) Vahl.
Neuracanthus robecchii (Lindau) C.B.Clarke subsp. atratus Bidgood & Brummitt - CP(2b, 4a, 4b, 8b); HH(4b,
4c); M23c; sW(3, 6a, 6b, 9, 16).
Peristrophe paniculata (Forssk.) Brummitt - CP2b; LH2.
Ruellia discifolia Oliv. - HH(1, 4a, 4b); M4; sW(1d, 2b,
6a, 8, 16a); nW6a - Syn.: R. longiflora Vahl.
Ruellia patula Jacq. s. l. - CP2c; LH(3a, 8b); HH1;
sW(1d, 2b).
Rungia pectinata Nees - LH(3a, 6, 7f ), M11d - Syn.: R.
parviflora Nees (Thulin et al., 2001: also Yemen).
Ruttya fruticosa Lindau - LH6; HH(8, 9); M14.
AIZOACEAE (INCL. MOLLUGINACEAE)
Aizoon canariense L. - L(2c, 5a); CP(4a, 10a, 12); LH12;
sW18c; nW(14d, 18c).
Corbichonia decumbens (Forssk.) Exell - LH 15; sW(19, 20).
Glinus lotoides L. - sW(12a, 12b); Wa4.
Trianthemum sp. - sW12a; nW28; Ha1a - Incomplete
specimens.
Zaleya pentandra (L.) C.Jeffrey - L1; CP(5a, 6a, 16) - Ruderal, disturbed areas.
AMARANTHACEAE
Achyranthes aspera L. - CP6a; LH(7b, 8a, 14); HH9;
M11b; sW12a; nW(14d, 18e) - Ruderal, disturbed areas.
Aerva artemisioides Viehr. & O.Schwartz subsp. batharitica A.G.Mill. & J.A.Nyberg - L8a; CP17; sW20;
nW(23, 27); Ha(1a, 1c, 2) - Endemic to Dhofar (Miller,
1996; Ghazanfar, 2003).
Contribution to the Flora of Central-Southern Dhofar (Sultanate of Oman)
Aerva javanica (Burm. f.) Juss. ex Schult. - CP(2b, 4a, 4b,
8b, 10b, 14, 15a, 15d); LH(2, 15, 17); sW(1a, 1b, 2a, 3,
6a, 6b, 6f, 9, 13a, 13b, 16a, 19, 20, 21); nW(14d, 18b,
25b, 26, 27).
Alternanthera pungens Kunth. - CP6a; LH2; M1; sW1d.
Amaranthus dubius Thell. - nW18c - First record from
Dhofar.
Amaranthus graecizans L. subsp. graecizans - M23b;
sW20; nW(14e, 18a); Ha1b.
Amaranthus graecizans L. subsp. silvestris (Vill.) Henkels
- CP4b; LH12; sW12a; nW(18a, 18e).
Amaranthus hybridus L. subsp. hybridus - sW12a,
nW14d.
Amaranthus viridis L. - sW12a.
Celosia trigyna L. - sW12a.
Digera muricata (L.) Mart. subsp. muricata - CP(1, 4b,
6a, 8b); LH8a; sW(5, 12a, 16); nW(14d, 14e, 18c).
Psilotrichum virgatum C.C.Towns. - sW(1b, 1c, 1f );
Ha1a.
Pupalia lappacea (L.) A.Juss. var. velutina (Moq.) Hook.
f. - CP(2c, 4b, 6a); LH8a; HH(1, 11); M2b; sW(1b, 2a,
2b, 8, 9).
ANACARDIACEAE
Lannea malifolia (Chiov.) Sacleux - CP13; LH12; M4;
sW(3, 6f ).
Pistacia chinensis Bunge subsp. falcata (Becc. ex Martelli)
Rech.f. - HH(4d, 4e); M 22.
Rhus somalensis Engl. - HH(4c, 4e); M(10a, 19, 20, 23b,
23c); nW8.
APIACEAE (UMBELLIFERAE)
Ammi majus L. - LH(6, 10); M(1, 17, 18, 19); sW(12a,
16); nW(14a, 14d, 17, 18c, 18e).
Adenosciadium arabicum (T.Anderson) H.Wolff - CP6a;
HH8; sW(1b, 1d, 12a, 20); nW19c; Ha1a.
Pimpinella schweinfurthii Asch. - LH(3a, 6, 10); HH(1,
5a, 10); M(11b, 18); sW16a.
Pycnocycla aucheriana Decne. ex Boiss. - Ha (1a, 2) Flowers white, calyx densely pubescent and bracts exceeding the umbel - First record from Dhofar.
APOCYNACEAE (INCL. ASCLEPIADACEAE)
Adenium obesum (Forssk.) Roem. & Schult. - CP(2c, 3);
HH4b; M23b; sW(16c, 16e, 16f ); nW17.
Carissa spinarum L. - HH5c; M23a - Syn.: C. edulis
(Forssk.) Vahl.
Calotropis procera (Aiton) W.T.Aiton - sW13a; nW2 Ruderal, disturbed areas.
Caralluma edulis (Edgew.) Benth. et Hook. - sW5 - Syn.:
Caudanthera edulis (Edgew.) Meve & Liede - Second
record from Oman.
Desmidorchis adenensis (Deflers) Meve & Liede - CP2a;
75
sW(1e, 1f, 1h) - Syn.: Caralluma adenensis (Deflers)
Berger - A rare, endangered species found only in a few
localities - Regional endemic: Oman & Yemen (Meve &
Liede, 2002; Pilbeam, 2010).
Desmidorchis flava (N.E.Br.) Meve & Liede - sW(6f, 16c,
16e, 16f ); nW(18a, 18c, 18e); Ha1a - Syn.: Caralluma
flava N.E.Br. - Regional endemic: Oman & Yemen
(Meve & Liede, 2002; Pilbeam, 2010).
Desmidorchis tardellii Mosti & Raffaelli - CP(4a, 4b);
LH17 (locus classicus); sW(8, 9) - Endemic to Dhofar
(Mosti & Raffaelli, 2004).
Desmidorchis cf. flava/tardellii - CP(8d, 14); sW(10a,
10b, 13a) - Specimens of doubtful identification: flowers lacking.
Echidnopsis scutellata A.Berger subsp. dhofarensis
Bruyns - nW(17, 18a, 18d, 19c); M9 - Syn.: Echidnopsis planiflora Bally - Endemic to Dhofar (Bruyns, 1988;
Pilbeam, 2010).
Glossonema varians (Stocks) Benth. ex Hook f. - CP10a;
M3; nW(14i, 18c).
Gomphocarpus fruticosus (L.) R.Br. - M(1, 11b, 19);
nW17.
Leptadenia pyrotechnica (Forssk.) Decne. - nW(5, 16,
19b).
Monolluma quadrangula (Forssk.) Plowes - LH11; M(2,
23a, 23b, 23c) - Syn.: Caralluma quadrangula (Forssk.)
N.E.Br. - Endemic to Oman, Saudi Arabia and Yemen
(Pilbeam, 2010).
Nerium oleander L. - CP8a - Culta.
Orbea luntii (N.E.Br.) Bruyns - M22 - Syn. Caralluma luntii N.E.Br., Pachycymbium luntii (N.E.Br.)
M.G.Gilbert - Regional endemic: Dhofar & S Yemen
(Raffaelli et al., 2008).
Orbea nardii Raffaelli, Mosti & Tardelli - M3 (locus classicus) - Endemic to Dhofar (Raffaelli et al., 2008; Mosti
& Raffaelli, 2011).
Pentatropis bentii (N.E.Br.) Liede - Ha(1b, 1c) - Regional
endemic: Dhofar & Yemen (Raffaelli et al., 2008).
Pentatropis nivalis (J.F.Gmel.) D.V.Field & J.R.I.Wood nW(20a, 20b) - Syn.: Pentatropis spiralis Auct. (non Decne.).
Pergularia daemia (Forssk.) Chiov. - CP6c, LH8b.
Pergularia tomentosa L. - L8a; CP16; LH15; sW(1b, 1c,
12b); nW7.
Periploca aphylla Decne. - nW16.
Periploca visciformis (Vatke) K.Schum. - HH4e; M(8,
22); sW(1b, 2a, 2b); nW(14d, 20a, 22).
Raphionacme arabica A.G.Mill. & Biagi - Wa8.
Rhazya stricta Decne. - nW(7, 13, 27).
Rhytidocaulon fulleri Lavranos & Mortimer - sW9 - A
rare, endangered species - Regional endemic: Oman &
Yemen (Lavranos & Mortimer, 1970; Pilbeam, 2010).
Sarcostemma viminale (L.) R.Br. - HH4e; nW(14a, 17,
19c).
76
ARISTOLOCHIACEAE
Aristolochia bracteolata Lam. - sW(8, 9, 11, 12a, 16d) Syn.: A. bracteata Retz.
ASTERACEAE (COMPOSITAE)
Atractylis kentrophylloides (Baker) F.G.Davies - M22b.
Bidens biternata (Lour.) Merr. & Scherff - CP6a; LH(7b,
8a); HH(5a, 6, 10); sW(2b, 8d); nW18c.
Blepharispermum hirtum Oliv. - CP6a; LH(6, 13, 14);
HH(1, 9) - Regional endemic: Dhofar & SE Yemen
(Thulin et al., 2001).
Blumea axillaris (Lam.) DC. - LH12; HH5c - Syn.: Conyza axillaris Lam., Blumea lacera (Burm. f.) DC. - First
record from Oman.
Centaurea dhofarica Baker - L2b.
Centaurea pseudosinaica Czerep. subsp. niebuhrii
Wagenitz - nW(14a, 14d, 14e, 20b).
Dicoma schimperi (DC.) Baill. ex O.Hoffm. - sW(1c, 1e,
1i, 5, 6a, 6f, 20); nW(1, 3); Ha(1a, 2) - Syn.: Hochstetteria schimperi DC.
Doellia bovei (DC.) Anderb. - HH11; nW(25a, 28) Syn.: Blumea bovei (DC.) Vatke.
Echinops spinosissimus Freyn - nW6a - First record from
Dhofar.
Eclipta prostrata (L.) L. - LH(8a, 9, 10); HH10; sW(11, 12a);
nW18c; Wa8 - Syn.: E. alba (L.) Hassk. (Beentje, 2006).
Erigeron bonariensis L. - M11b - Syn.: Conyza bonariensis
(L.) Cronq.
Erigeron trilobus (Decne.) Boiss. - HH(4e, 8a); M(1, 11a,
11b, 15); nW6c - Syn.: Conyza stricta Willd.
Helichrysum somalense Baker f. - LH(6, 11); HH(4b,
4e); M(1, 8, 11b, 23b); sW1d; nW(14l, 17, 18a, 18c,
18e); Ha2 - The specimens in LH6, M11b, nW(17,
18a) have leaves up to 25 mm long and 11 mm wide (vs
15 mm and 4,5 mm max according to Beentje, 2006);
the inflorescences have more than 10 heads.
Iphiona scabra DC. ex Decne. - sW(18a, 18c); nW(3, 7,
13, 14b, 14c, 14e, 27).
Iphiona senecionoides (Baker) Anderb. - L8c; CP14;
sW(1b, 1i, 2a, 2b, 12a, 13a, 13b); nW(10, 14d, 14m,
27); Ha1a - Syn.: Grantia senecionoides Baker.
Iphiona teretifolia Anderb. - nW(20a, 21) - First record
from Oman - Found up to now only in a restrict area of
Yemen (Anderberg, 1985) - Regional endemic: Dhofar
& Yemen - (Fig. 3a).
Kleinia odora (Forssk.) DC. - CP14; HH4e; M12;
nW(17, 18a, 19b); Ha1.
Kleinia saginata P.Halliday - M23b; nW(17, 19b) - Endangered species, known only from few localities - Endemic to Dhofar (Miller & Morris, 1988).
Launaea almahrahensis N.Kilian - nW14d.
Launaea castanosperma F.G.Davies - L3b; sW(1b, 1d,
13a); nW(10, 19b, 20b, 26).
S. Mosti , M. Raffaelli, M. Tardelli
Launaea crassifolia (Balf. f.) C.Jeffrey - CP(2b, 2c, 4b, 6a);
LH(3a, 6, 8b, 14); HH(1, 8a, 8b, 9, 14); sW(1d, 2b,
16a); nW(14m, 18c, 18d); Ha1a.
Launaea intybacea (Jacq.) Beauverd - CP5a.
Launaea massauensis (Fresen.) Sch.Bip. ex Kuntze - CP2c;
LH5; sW(1c, 1d); nW(14d, 18a, 18c, 19c).
Launaea sp. plur. - CP14, M18 - Incomplete specimens.
Parthenium hysterophorus L. - LH8a; M(11a, 11b);
sW9; nW6c - Pantropical.
Pegolettia senegalensis Cass. - M22b.
Pluchea arabica (Boiss.) Qaiser & Lack - LC8, CP(2a, 2b,
16, 17); LH13; M11c, sW(3, 12b, 19, 21); nW(1, 6a,
10, 11, 14f, 17, 18b, 19a, 20b, 25b, 27); Wa6.
Pluchea ovalis (Pers.) DC. - CP5a; HH(3, 4d); M(11a,
11b); sW1a.
Pseudoconyza viscosa (Mill.) D’Arcy - sW(3, 13a, 12b) Syn.: Blumea aurita (L. f.) DC.
Psiadia punctulata (DC.) Vatke - M(22, 23b, 23c) - Syn.:
P. arabica Jaub. et Spach.
Pulicaria argyrophylla Franch. var. oligophylla GamalEldin - CP2c; HH4b; M(1, 8, 11a, 11b, 22, 23b);
nW(14a, 14b, 14c, 14g, 14l, 17, 18b, 20a, 21).
Pulicaria cylindrica (Baker) O.Schwartz - nW(3, 7, 10,
12, 14h, 14l, 14o, 26, 27).
Pulicaria gamal-eldiniae N.Kilian & P.Hein - sW1i - First
record from Oman - Known from S Yemen (Kilian,
1999) - Regional endemic: Dhofar & S Yemen.
Pulicaria glutinosa (Boiss.) Jaub. et Spach subsp. glutinosa - L8a; M23c.
Pulicaria inuloides (Poir.) DC. - M11a.
Pulicaria nobilis Gamal-Eldin - M6 - A rare species, second record from Oman (Gamal-Eldin, 1981; Mosti &
Selvi, 2007).- (Fig. 3b).
Pulicaria cf. nobilis - M23c - Incomplete specimen: flowers lacking.
Pulicaria omanensis Gamal-Eldin subsp. omanensis LH15; sW(2b, 3, 6b, 20); nW(14b, 14c, 14d, 14e, 14g,
14o, 14n, 25b, 27); Ha(1a, 2) - Regional endemic: Dhofar & SE Yemen (Thulin et al., 2001).
Pulicaria undulata (L.) C.A.Mey. - CP(7, 11); sW(3, 12b);
nW(1, 3, 7, 12, 13, 27) - Syn.: P. crispa (Forssk.) Benth.
Sclerocarpus africanus Jacq. ex Murray - LH(8a, 14);
HH(5b, 8, 9).
Tridax procumbens L. - CP6d; sW(8, 9) - Pantropical,
collected in Salalah (Dhofar) - First record from Oman.
Tripteris vaillantii Decne. - nW(14e, 17, 18a, 18c, 18d,
19a) - Syn.: Osteospermum vaillantii (Decne.) Norl.
Vernonia arabica F.G.Davies - CP(2a, 2b, 4a, 8b); LH15;
HH(4b, 4e, 11); M(1, 22, 23a, 23b); sW(1b, 1c, 1d, 1e,
6a, 6b, 6f, 8, 9, 12a, 13a); nW(14a, 14d, 14i, 14o, 17,
18a, 19b, 20a, 25a, 27, 28).
Vernonia cinerascens Sch.Bip. - M 23b - First record from
Oman.
Contribution to the Flora of Central-Southern Dhofar (Sultanate of Oman)
Vernonia cinerea (L.) Less. - CP6a; LH(5, 7f, 8b, 10, 14);
HH9; M20; sW(12a, 16a); Wa(4, 8).
Volutaria sinaica (DC.) Wagenitz - M1; sW2b; nW(18c,
18d, 18e, 20a) - Syn.: V. lippii (L.) Cass. ex Maire.
BALSAMINACEAE
Impatiens balsamina L. - LH(3a, 7b); HH4b; M14;
sW16a - Native of tropical Asia.
BOMBACACEAE
Adansonia digitata L. - LH14b; sW(16a, 16b, 16c, 16d,
16e, 16f ) - Known only from two localities: Dalkut
(near the Yemen border) and Wadi Hinna (hills between
Khor Rori and Mirbat).
BORAGINACEAE
Arnebia hispidissima (Lehm.) DC. - L5a; CP(2c, 4b, 8b,
15c); LH15; M11c; sW(1d, 5); nW(1, 7, 10, 14d, 14e,
15, 18a, 18c, 25b, 26, 27); Ha1a.
Cordia monoica Roxb. - LH7d; M21 - Syn.: C. ovalis
Hochst. ex DC.
Cordia myxa L. - M11a; sW16f.
Cordia nevillii Alston - HH5a - Syn:. C. perrottettii Wight
(cf. Warfa, 1990; Thulin et al., 2001).
Echiochilon arabicum (O.Schwartz) I.M.Johnst. - sW(1f,
1i); sW(6a, 6b).
Echiochilon callianthum Lönn - sW6b.
Echiochilon jugatum I.M.Johnst. - L5a; CP10a.
Echiochilon kotschyi (Boiss. & Hohen.) I.M.Johnst. nW(14l, 14o, 21).
Echiochilon longiflorum Benth. - sW(1e, 6b, 6c).
Echiochilon persicum (Burm. f.) I.M.Johnst. - nW(1,
14d, 14n).
Echiochilon sp. plur. - LH15; sW(6b, 6f ); nW(1, 18e) Incomplete specimens.
Ehretia obtusifolia Hochst. ex DC. - CP6c.
Heliotropium bacciferum Forssk. - L(7, 8a, 8c); CP(14,
15d); LH7b; M(8, 22, 23a, 23b); sW(17, 18c, 20, 21);
nW(3, 4, 11, 14i, 18b, 18c, 20a, 21, 25b, 26, 27);
Ha(1a, 1b).
Heliotropium brevilimbe Boiss. - L(5a, 6); sW(17, 19);
nW(14a, 14d, 14e, 14o, 27); Ha1a - Syn.: H. calcareum
Stocks.
Heliotropium fartakense O.Schwartz - L(2a, 2c, 3b, 5a);
CP(2b, 4b, 5a, 8a, 8b, 10a, 12); LH(2, 15); HH11;
sW(1b, 1c, 2a, 2b, 3, 6a, 6b, 12a, 13a, 13d); nW(1, 7,
14b, 14m, 18a, 18c, 18d).
Heliotropium longiflorum (A.DC.) Jaub. & Spach subsp. longiflorum - CP(6a, 6b, 8d); LH(8b,12); HH5b;
Ha1a; M(1, 23b); sW(1d, 2b, 6b, 12a, 16a, 16d, 20);
nW(18c, 18d); Ha1a.
Heliotropium ophioglossum Stocks ex Boiss. - LH15;
M(23a, 23b); sW1d; nW18c.
77
Heliotropium ovalifolium Forssk. - LH8a; sW(3, 12a,
12b).
Heliotropium rariflorum Stocks - HH3; sW1d; nW26.
Heliotropium steudneri Vatke - CP4b; sW(1d, 2b, 6b,
9) - First record from Oman.
Heliotropium subspinosum Thulin & A.G.Mill. - M6;
sW6e.
Nogalia drepanophylla (Baker) Verdc. - CP14; LH15;
sW(10b, 11, 13a, 16a).
Trichodesma cinereum Mosti & Selvi - M6 (locus classicus), 23c; nW1 - Syn.: T. dhofaricum Ghaz. (Ghazanfar,
2010) - Endemic to Dhofar (Mosti & Selvi, 2007).(Fig. 3c).
Trichodesma hildebrandtii Gürke - CP(4b, 8b); M23b;
sW1d; nW(6d, 14o, 18e, 18b).
BRASSICACEAE
Diplotaxis harra (Forssk.) Boiss. - CP(2b, 2c, 6a, 6b);
sW(1a, 1d, 2b); nW(10, 18a, 18b, 18c, 18e, 19b, 19c,
25b).
Farsetia dhofarica Jonsell & A.G.Mill. - nW(26, 27) - Regional endemic: Oman & S Yemen (Miller, 1996).
Farsetia latifolia Jonsell & A.G.Mill. - CP4b; LH(2, 15);
HH4b; M3; sW(1a, 2b, 6b, 12a, 5, 9); nW(14d, 14e,
18a, 18c, 18d, 20a).
Farsetia longisiliqua Decne. - nW14e.
Farsetia sp. plur. - CP2c; LH(2, 15); M11b; sW(1d, 6b,
6c); nW(14o, 26, 27) - Incomplete specimens.
Moricandia sinaica (Boiss.) Boiss. - nW(14e, 19b, 19c).
BURSERACEAE
Boswellia sacra Flueck. - HH(4c, 4e, 11); sW(1a, 1f, 1i,
1g, 1h, 2a, 2b, 6a, 6b, 6c, 6d, 6e, 6f ); nW(1, 6b, 6c, 6d,
9, 14a, 14b, 14c, 14d, 14e, 14f, 14g, 14h, 14i, 14l, 14m,
14n, 14o, 15, 16, 21); Ha(1a, 1b, 1c, 2) - The localities in italics refer to specimens observed in the field but
not collected. - cf. Raffaelli et al. (2003, 2006); Raffaelli,
Tardelli et al. (2003). - (Fig. 3h).
Commiphora foliacea Sprague - CP(2a, 14, 15c, 17);
LH(15, 16); HH(4b, 4c); M22; sW(1c, 6a, 6b, 6c, 17,
19, 20, 21); Ha1a.
Commiphora gileadensis (L.) C.Chr. - CP(6b, 6c); LH(1,
3a, 7c, 8a, 8b, 16, 24); HH14; M(2, 3, 6, 22).
Commiphora kua (Royle) Vollesen - CP(4b, 6a, 6b,
8a, 11); LH(1, 6, 7b, 7c, 8a, 9); HH5b; M(20, 23a);
sW(13a, 16a); nW(17, 19b, 19c); Ha1a - Syn.: C. habessinica (O.Berg) Engl.
Commiphora wightii (Arn.) Bhandari - CP14; sW(1d,
6c).
CAPPARACEAE
Boscia arabica Pestal. - CP(6d, 9); LH1; sW(1a, 3, 13a).
Cadaba baccarinii Chiov. - sW6f.
78
Cadaba farinosa Forssk. - CP(5b, 8b, 8d, 10a, 10b); sW6f;
nW6b; Ha1c; Wa6.
Cadaba heterotricha Stocks ex Hook. - sW(4, 6a); Ha(1a,
1c, 2).
Capparis cartilaginea Decne. - CP(16, 20); HH11; M8;
sW(3, 20); nW(19a, 22).
Cleome albescens Franch. subsp. omanensis D.F.Chamb.
& Lamond - sW6b; nW(18e, 26) - Regional endemic:
Dhofar & SE Yemen (Thulin et al., 2001).
Cleome amblyocarpa Barratte & Murb. - CP2c.
Cleome austroarabica Chamb. & Lam. subsp. austroarabica - L3b; CP(8b, 15a, 16); LH15; HH11; sW(1a, 1d,
6a, 6d, 9, 12b, 13a, 18a, 18c, 19, 20); nW(14a, 14c,
14d, 14e, 15, 18e, 25a, 25b, 26, 27, 28); Ha2.
Cleome austroarabica Chamb. & Lam. subsp. muscatensis Chamb. & Lam. - nW25 - First record from Dhofar.
Cleome brachycarpa Vahl ex DC. - L5a; CP(4b, 8a, 8b, 10a,
10b, 16); LH15; HH4c; sW(1e, 1d, 2b, 3, 5, 6b, 12a, 19,
20, 21); nW(7, 10, 14d, 14e, 25a, 25b, 26, 28); Ha1a.
Cleome brevipetiolata Chamb. & Lam. - nW(2, 14c).
Cleome gynandra L. - CP2c - Syn.: Gynandropsis gynandra (L.) Briq.
Cleome noeana Boiss. subsp. noeana - sW(2b, 6a); nW1.
Cleome scaposa DC. - nW2 - First record from Dhofar.
Dhofaria macleishii A.G.Mill. - M(22, 23a); nW(19a,
19c) - Regional endemic: Dhofar & SE Yemen (Thulin
et al., 2001).
Dipterygium glaucum Decne. - nW(4, 7).
Maerua crassifolia Forssk. - LH17; sW1f; nW(6b, 6d, 9,
25a, 26, 28).
CARYOPHYLLACEAE
Cometes abyssinica R.Br. ex Wall. - L5a; CP(8b, 14,
15d); LH(15, 16); HH11; sW(13a, 21a); nW(14l, 25b);
Ha1a.
Gymnocarpos dhofarensis Petruss. & Thulin - nW(10,
14a, 14c, 14d, 20b) - Regional endemic: Dhofar & S
Yemen (Petrusson & Thulin, 1996).
Gypsophila montana Balf. f. subsp. somalensis (Franch.)
M.G.Gilbert - CP2b; HH(4b, 11); sW(1a, 1b, 1d, 1f, 8,
9, 13a, 18a, 18c); nW(18a, 25b); Wa6.
Herniaria maskatensis Bornm. - nW(3, 14d, 14h, 14i,
14n, 26) - Regional endemic: Oman & S Yemen (Miller,
1996).
Polycarpaea repens (Forssk.) Asch. & Schweinf. - nW(2,
25b).
Polycarpaea robbairea (Kuntze) Greuter & Burdet nW(18a, 18b, 18c, 18e).
Polycarpaea spicata Wight ex Arn. - L(4, 5a); CP(2b, 10a,
12); sW(1a, 1c, 5); nW(14d, 18a).
CELASTRACEAE
Maytenus dhofarensis Sebsebe - LH(6, 7d, 13, 7f );
S. Mosti , M. Raffaelli, M. Tardelli
HH(5a, 5b, 5c, 11); M(8, 11b, 11d, 12, 13, 19); nW6d
- Regional endemic: Dhofar & SE Yemen (Ghazanfar,
2007).
Maytenus senegalensis (Lam.) Exell - nW(21, 27).
CERATOPHYLLACEAE
Ceratophyllum demersum L. - CP8d; sW3; nW18e.
Ceratophyllum submersum L. - L3a; sW(1e, 13a) - First
record from Dhofar.
CHENOPODIACEAE
Arthrocnemum macrostachyum (Moric.) K.Koch - CP5a.
Atriplex farinosa Forssk. - CP5a.
Chenopodium murale L. - CP6b; HH4e; M23a; sW13a;
nW(18a, 18e).
Cornulaca amblyacantha Bunge - nW26.
Halotamnus bottae Jaub. & Spach - sW6f; nW(3, 14l,
21, 25b, 27).
Salsola imbricata Forssk. - CP(4a, 10a); LH15 - Syn.: S.
baryosoma (Roem. & Schult.) Dandy.
Salsola spinescens Moq. - CP(9, 10b); LH17; sW11.
Salsola rubescens Franch. - nW(14l, 14o, 27).
Suaeda aegyptiaca (Hasselq.) Zohary - CP16; nW(20a,
26, 27).
Suaeda vermiculata Forssk. ex J.F.Gmel. - CP(2b, 4b, 5a,
8a, 8b, 9, 10a); sW11; nW(18a, 18b, 18c, 18e) - Syn.:
S. fruticosa Forssk. ex J.F.Gmel.
CISTACEAE
Helianthemum citrinum Ghaz. - M6; sW1d; nW(14o,
18c, 18d, 18e, 19c, 25a, 28).
COMBRETACEAE
Anogeissus dhofarica A.J.Scott - LH(7a, 7b, 7c, 7d, 7e,
7f, 8a); HH(1, 2, 8, 10); M(1, 11a, 11b, 11d) - Regional
endemic: Dhofar & SE Yemen (Thulin et al., 2001).
CONVOLVULACEAE
Convolvulus glomeratus Hochst. ex Choisy - CP(2b, 2c,
6a, 6b, 8c); LH(3a, 5, 11, 15); HH2; M1; sW(1d, 6a,
6b, 20); nW14d; Ha1a.
Convolvulus hystrix Vahl - nW(6a, 6d, 7, 14c, 14d, 14e,
14o, 15, 22).
Convolvulus prostratus Forssk. - sW(6e, 18b); nW(6b,
14c, 14d).
Convolvulus ulicinus Boiss. - CP15b; sW18a; nW27;
Ha(1a, 2) - cf. Raffaelli et al. (2006).
Cressa cretica L. - L(2c, 3a); CP(8a, 8b, 8d, 12); sW(11,
13a).
Hildebrandtia africana Vatke - nW(17, 19c).
Hildebrandtia somalensis Engl. ex Peter - sW1g - First
record from Oman.
Ipomoea aitonii Lindley - CP6a; LH14b; HH(4b, 8).
Contribution to the Flora of Central-Southern Dhofar (Sultanate of Oman)
Ipomoea cairica (L.) Sweet - CP(2b, 2c); LH(3a, 5, 7f );
HH(1, 5a).
Ipomoea nil (L.) Roth - CP6a; LH(3a, 8a, 14).
Ipomoea pes-caprae (L.) R.Br. - L(1, 2c); CP(5a, 11);
LH9; HH7.
Ipomoea obscura (L.) Ker.-Gawl. - LH3a; HH1; M21;
nW19a.
Ipomoea sinensis (Desr.) Choisy - CP(2c, 6a); LH(3a, 11,
14); M1; sW16.
Ipomoea turbinata Lag. - HH8.
Merremia somalensis (Vatke) Hallier f. - sW(1d, 6a, 6b,
6e, 6f ); nW(6a, 6b, 10).
Seddera glomerata (Balf. f.) O.Schwartz - LH15; M3;
sW(1c, 1e); nW(18a, 18b).
Seddera latifolia Steud. & Hochst. - CP(8b, 10a).
CUCURBITACEAE
Citrullus colocynthis (L.) Schrad. - L5a; CP(7, 18).
Corallocarpus epigaeus (Rottler) Hook f. ex C.B.Clarke CP(4a, 4b); LH(6, 8b); M(11b, 17).
Corallocarpus cf. glomeruliflorus (Deflers) Cogn. - LH16;
sW6d - Insufficient materials: flowers and fruits lacking.
Cucumis prophetarum L. subsp. prophetarum - CP14;
LH(15, 17); M23a; sW(2b, 6e, 13a); nW(14a, 14c,
25b, 26); Wa6.
Cucumis prophetarum L. cf. subsp. dissectus (Naudin)
C.Jeffrey - nW(18c, 18d); M17 - Unripe fruits.
Cucumis pustulatus Hook. f. - M11a.
Cucumis sativus L. - LH(3a, 7b).
Luffa acutangula (L.) Roxb. - CP6a; LH(6, 7b, 8a, 10,
11); HH(2, 5a, 7, 10).
Mukia maderaspatana (L.) M.Roem. - LH3a; HH10;
sW(12a, 16a).
EBENACEAE
Euclea racemosa Murray subsp. schimperi (A.DC.) F.
White - LH7d; HH(1, 3); M11a.
EUPHORBIACEAE
Acalypha indica L. - CP(4b, 6a); LH(11, 15); sW(5, 12a);
Wa8.
Andrachne aspera Spreng. var. aspera - CP(6a, 6b); M11b;
sW(1e, 5); nW27.
Andrachne aspera Spreng. var. glandulosa A.Rich. - CP8b;
sW(6a, 6f ); nW(14a, 14c, 14e, 14o, 20a).
Andrachne telephioides L. - sW(1d, 1f, 2b, 6b, 18c);
nW(6b, 14d, 14l, 18c).
Chrozophora oblongifolia (Delile) A.Juss. ex Spreng. CP14; LH15; HH11; sW(1b, 1d, 2b, 6b, 8, 9, 13a);
nW(14d, 14e, 18c, 25b).
Croton confertus Baker - LH7e; M13 - Endemic to Dhofar, SE Yemen and N Somalia (Thulin et al., 2001).
Dalechampia scandens L. var. cordofana (Hochst. ex
79
Rich.) Müll.-Arg. - CP6b; LH5; HH(4b, 4c, 11);
sW(1b, 1c, 1d, 16d).
Euphorbia arabica Hochst. & Steud. ex Boiss. - sW(1d,
6b, 19).
Euphorbia balsamifera Ait. subsp. adenensis (Deflers)
Bally - M(1, 3, 14, 16, 22, 23b, 23c); nW17.
Euphorbia cactus Ehrenb. ex Boiss. - CP(2a, 2b, 2c, 6a,
6b, 6c); LH(13, 14); HH(3, 4b, 4c, 4e); M(22, 23b);
sW6f; nW(17, 18a, 18b, 18c, 18d, 18e, 19a, 19b, 19c) The localities in italics refer to specimens observed in the
field but not collected.
Euphorbia dhofarensis S.Carter - CP9; M5; nW14d;
Ha1a - Regional endemic: Dhofar & SE Yemen (Ghazanfar, 2007).
Euphorbia granulata Forssk. var. granulata - sW6b;
nW14d - First record from Oman.
Euphorbia granulata Forssk. var. glabrata (Gay) Boiss. M17; sW(12a, 16); Wa7.
Euphorbia hadramautica Baker - CP(3, 4a, 4b); nW(18a,
18c); M20; sW(5, 6b, 6f, 9) - A very variable species in
Dhofar, see Mosti et al. (2005).
Euphorbia hirta L. - CP8c; HH(4b, 4e); M1; sW(1d, 9);
nW18c - (cf. Raffaelli et al., 2003).
Euphorbia larica Boiss. - CP9; M5; nW14d; Ha1a.
Euphorbia orbiculifolia S.Carter - HH(4b, 4c); M23c
- Regional endemic: Dhofar & SE Yemen (Ghazanfar,
2007).
Euphorbia schimperi C.Presl - HH(4b, 4c, 7c, 9); M(3,
8, 23b); nW14l.
Euphorbia serpens Kunth - Wa2 - First record from Dhofar.
Euporbia smithii S.Carter - CP(2c, 6a, 6b); LH(3a, 6, 7b,
7e, 8b) - Regional endemic: Dhofar & SE Yemen (Thulin et al., 2001).
Euphorbia aff. uzmuk S.Carter & J.R.I.Wood - CP(2a,
14); HH4e; M11b; sW1i - (cf. Miller & Morris, 1988).
Jatropha dhofarica Radcl.-Sm. - CP(2a, 4b, 6a, 6b, 11);
LH(6, 7e, 9, 11); HH(1, 5c); M13; sW16a - Regional
endemic: Dhofar & SE Yemen (Thulin et al., 2001).
Jatropha pelargoniifolia Courbon var. sublobata (O.
Schwartz) Radcl.-Sm. - CP3; M7; sW(1h, 2b, 6b, 6c, 6e).
Meineckia phyllanthoides Baill. - CP6b; LH3a.
Phyllanthus maderaspatensis L. - M3; sW(1d, 1e).
Phyllanthus rotundifolius Willd. - CP6a; LH(7b, 8a).
Ricinus communis L. var. africana Müll.-Arg. - M23b;
sW6b.
FABACEAE (CESALPINIOIDEAE)
Cesalpinia erianthera Chiov. - sW(1i, 3, 6f ).
Cesalpinia bonduc (L.) Roxb. - Culta in Al Balid Botanical Garden (Salalah).
Delonix elata (L.) Gamble - CP(6a, 12); LH(3a, 13);
sW(14, 16a).
80
Senna holosericea (Fresen.) Greuter - CP(8a, 10a);
LH(15, 17); sW(1c, 1d, 2a, 6b, 8, 13a, 24); nW(14d,
14e, 25b, 26).
Senna italica Mill. - nW7.
Senna sophera (L.) Roxb. - nW19a - First record from
Dhofar.
Tamarindus indica L. - HH1; M20.
FABACEAE (FABOIDEAE)
Alysicarpus glumaceus (Vahl) DC. - LH6; HH10.
Alysicarpus longifolius (Rottl. ex Spreng.) Wight & Arn.
- CP6b.
Alysicarpus sp. - LH10 - Pods lacking.
Crotalaria aegyptiaca Benth. - CP15a; sW(19, 20);
nW(6a, 20a, 20b, 21, 27).
Crotalaria leptocarpa Balf. f. - nW14d.
Crotalaria medicaginea Lam. - LH(7b, 8b, 12); HH10;
Wa7.
Crotalaria persica (Burm. f.) Merr. - L5a; CP12; sW(10b,
13a, 13b); nW(7, 14d, 27).
Crotalaria retusa L. - LH7f; HH(2, 10); Wa7.
Indigofera articulata Gouan - M11c; sW(3, 6b, 13a, 18a);
nW(19a, 19b, 25b).
Indigofera coerulea Roxb. var. coerulea - CP(4b, 6a);
sW(8, 12b, 13a).
Indigofera coerulea Roxb. var. occidentalis J.B.Gillett &
Ali - CP6b; sW16a.
Indigofera nephrocarpa Balf. f. - CP2b; sW(1c, 20);
nW(14a, 14c).
Indigofera oblongifolia Forssk. - CP(4a, 5a, 17, 20);
nW(1, 6a, 6b, 6d, 25a, 28).
Indigofera rubromarginata Thulin - L5a; sW(19, 21).
Indigofera semitrijuga Forssk. - sW 21; nW(7, 14a, 14d,
14e, 14o, 26).
Indigofera trita L. f. var. maffeii (Chiov.) Ali - HH(5a,
10).
Indigofera volkensii Taub. - CP(2a, 2c, 4b, 8b); LH2;
HH4e; M1; sW(1d, 6b, 12a, 13b); nW18c.
Ormocarpum dhofarense Hillc. & J.B.Gillett - CP(6a,
6b); LH(3a, 8b, 14); HH(8, 10, 11); M14 - Regional
endemic: Dhofar & E Yemen (Thulin et al., 2001).
Psoralea corylifolia L. - M20l.
Rhynchosia minima (L.) DC. - CP(2c, 6b); LH(11, 14);
HH(4b, 4c, 9); M(11b, 14, 17).
Rhynchosia pulverulenta Stocks - CP8b; sW(1d, 5, 6b,
8, 20); Ha1a.
Rhynchosia schimperi Boiss. - CP6d.
Taverniera brevialata Thulin - CP(15d, 20); Ha1b.
Taverniera multinoda Thulin - CP(15a, 15d); nW(10,
14o).
Tephrosia pentaphylla (Roxb.) G.Don - Ha(1a, 1c) - First
record from Dhofar. - (Fig. 3g).
Tephrosia purpurea (L.) Pers. - CP(14, 17, 20); sW(3, 19,
S. Mosti , M. Raffaelli, M. Tardelli
20); nW(1, 14d, 25b, 27).
Tephrosia subtriflora Hochst. ex Baker - L5a; CP(2c,
6a); LH(7b, 8a, 14, 15); HH(9, 11); M(14, 17); sW16;
nW(18c, 18d) - cf. Raffaelli et al. (2006).
Tephrosia uniflora Pers. - CP4b; LH(16, 17); sW(1f, 2a,
3, 8, 9, 11).
Tephrosia sp. plur. - LH(11, 12, 15); sW(1f, 8, 20) - Incomplete specimens.
Teramnus repens (Taub.) Baker f. subsp. gracilis (Chiov.)
Verdc. - LH8a; HH10; sW12a.
FABACEAE (MIMOSOIDEAE)
Acacia ehrenbergiana Hayne - sW2a.
Acacia etbaica Schweinf. subsp. uncinata Brenan - CP1;
HH11.
Acacia gerrardii Benth. subsp. gerrardii - M21.
Acacia hamulosa Benth. - nW3.
Acacia laeta R.Br. ex Benth. - sW(3, 4, 6d).
Acacia nilotica (L.) Delile subsp. kraussiana (Benth.) Brenan - CP(8a, 11); LH(7d, 9, 13); HH10; M(11b, 20);
sW(12b, 16d); Wa8.
Acacia senegal (L.) Willd. - CP(1, 6a, 6c, 11); LH(3a, 3b,
7c, 7e, 15); HH11; M(10a, 13, 14); sW(12b, 16d, 17,
20); nW(19a, 27); Ha1c; Wa7.
Acacia tortilis (Forssk.) Hayne - CP(6c, 8a, 8d, 15b);
LH(15, 17); nW(6b, 7, 14c, 14o); Ha2.
Acacia sp. plur. - CP2b; sW1b; nW(4, 11) - Incomplete
specimens: flowers and pods lacking.
Prosopis cineraria (L.) Druce - sW16d.
Prosopis juliflora (Swartz) DC. - CP(6a, 8d).
GENTIANACEAE
Canscora concanensis C.B.Clarke - HH5a.
Enicostema axillare (Lam.) A.Raynal - L8a.
Exacum affine Balf. f. ex Regel - L8a - Syn.: E. gracilipes
Balf. f. - Endemic to Dhofar and Socotra (Thulin, 2001).
Exacum arabicum Thulin - CP2c; LH14b; HH(4b, 15);
M17 - Regional endemic: Dhofar & S Yemen (Thulin,
2001).
GERANIACEAE
Geranium biuncinatum Kokwaro - M19.
Geranium mascatense Boiss. - CP6b; HH10; M(14, 18,
19).
Geranium trilophum Boiss. - M14 - Syn.: G. yemense Deflers s. str.
Monsonia heliotropoides (Cav.) Boiss. - sW6b; nW(14d,
14e, 25b, 26).
LAMIACEAE (LABIATAE)
Becium dhofarense Sebald - LH8b; HH(1, 4b, 4c, 9);
M19 - Regional endemic: Dhofar & SE Yemen (Thulin
et al., 2001).
Contribution to the Flora of Central-Southern Dhofar (Sultanate of Oman)
Endostemon tenuiflorus (Benth.) M.Ashby - CP(2b, 6a,
6b, 8b); LH2; HH(4c, 11); M14; sW(1d, 5, 6b).
Lavandula dhofarensis A.G.Mill. subsp. dhofarensis CP(2b, 2c, 6a, 6b, 6c); LH(15, 16); M23b; sW(6f, 8,
16f ); Wa6 - Regional endemic: Dhofar & SE Yemen
(Thulin et al., 2001).
Lavandula dhofarensis A.G.Mill. subsp. ayunensis
A.G.Mill. - HH4e; sW(1d, 2b, 9); nW(6d, 19c, 20a) Endemic to Dhofar (Miller & Morris, 1988; Ghazanfar,
1992a).
Lavandula dhofarensis A.G.Mill. subsp. dhofarensis/ayunensis - M23b; nW1 - Specimens with intermediate
characters between the two subspecies.
Lavandula macra Baker - CP15a; sW(1a, 1f, 1i, 2a, 2b, 3,
4, 6a, 6e, 6f, 17, 18a, 21); nW27; Ha1a.
Leucas dhofarensis Hedge & Sebald - LH(2,14); HH4b;
M14; nW(17, 18a, 18d) - Regional endemic: Dhofar &
SE Yemen (Thulin et al., 2001).
Leucas urticifolia (Vahl) R.Br. - CP(2a, 2c, 4b, 6a, 6b);
LH11; sW12a.
Leucas inflata Benth. - sW(1d, 1e).
Ocimum forskaolii Benth. - CP(2c, 6a, 6b, 8c); LH(3a,
5); HH(1, 4a, 4b, 4e); M11a; sW(1d, 9, 16a, 16d);
nW(17, 18c, 18d).
Orthosiphon pallidus Royle ex Benth. - CP(2c, 4b, 6a,
6b); LH(3a, 8a, 11); M11a; sW(5, 9, 12a).
Orthosiphon sp. plur.- HH4b; sW8; Wa6 - Leaves hairy;
flowers lacking.
Plectranthus barbatus Andrews - LH7b; HH(5a, 5b);
M14.
Plectranthus cylindraceus Hochst. ex Benth. - sW1f;
nW(17, 18a).
Salvia macilenta Boiss. - nW(14c, 14h, 14o) - First record
from Dhofar.
Salvia sp. plur. - M(23a, 23b); nW18c - Insufficient and
incomplete materials.
Teucrium nummularifolium Baker - LH10; HH(4c, 9);
M(11b, 17, 18, 19); sW(1c, 1d, 12a, 8, 16a); nW17.
Teucrium sp. plur. - CP(3, 11); LH15; HH4b; M(2b, 3,
11a, 23b); sW(1c, 1f, 8, 9); nW19c - Incomplete specimens.
LORANTHACEAE
Plicosepalus acaciae (Zucc.) Wiens & Polhill - Wa6 - Syn.:
Loranthus acaciae Zucc.
LYTRACEAE
Lawsonia inermis L. - CP5b; sW(16b, 16d).
Woodfordia uniflora (A.Rich.) Koehne - HH11; sW1a;
nW(6a, 6d).
MALPIGHIACEAE
Acridocarpus orientalis A.Juss. - sW(1a, 1f, 2a).
81
MALVACEAE
Abelmoschus esculentus (L.) Moench - M20.
Abelmoschus manihot (L.) Medik. - M(11a, 20).
Abutilon bidentatum A.Rich. - LH8b; HH(4b, 4e).
Abutilon fruticosum Guill. & Perr. - CP(2b, 2c, 3, 4b,
6b); LH14b; HH4b; sW(6f, 12a, 16a, 20); nW1;
Ha(1a, 1b, 1c).
Abutilon hirtum (Lam.) Sweet - HH8.
Abutilon mauritianum (Jacq.) Medik. - Wa8.
Abutilon pannosum (G. Forst.) Schltdl. - CP4a; HH4b;
sW(6b, 19, 20); nW6d.
Cienfuegosia welshii (T. Anders.) Garcke - sW1e.
Gossypium hirsutum L. - CP5a - Culta.
Gossypium stocksii Mast.- CP(2a, 2b, 2c, 3, 4a, 8b);
sW(1a, 1b, 1d, 1g, 2b, 6a, 11, 13a).
Hibiscus micranthus L. f. - CP10b; HH11; M23b;
sW(1c, 1d, 8, 17c) - First record from Dhofar.
Hibiscus cf. scindicus Stocks - CP8b; sW8 - Flowers lacking.
Hibiscus vitifolius L. - M14.
Pavonia arabica Hochst. ex Steudl. - CP(2b, 2c, 4b, 6a,
6b); LH(3a, 5, 11); HH4b; M11b; sW1d; nW18d.
Pavonia pirottae (A.Terracc.) Chiov. - sW(1d, 5, 6a).
Pavonia schweinfurthii Ulbr. - HH4b; sW1e.
Pavonia sp. plur. - M11b; nW18c - Incomplete specimens.
Senra incana Cav. - L2d; CP(2b, 2c, 3, 5a, 8a, 16); LH16;
sW(11, 13a, 16a, 20, 21); Ha1a.
Sida cordata (Burm. f.) Borss.Waalk. - CP6a; LH(7a, 7f,
14); HH10.
Sida ovata Forssk. - CP(4b, 8b); LH(6, 15); HH(1, 2, 4b);
M(1, 19); sW(1e, 6b, 12a, 9, 16a).
Thespesia populnea (L.) Sol. ex Correa - L(8a, 8b) - First
record from Oman.
MENISPERMACEAE
Cocculus balfourii Schweinf. ex Balf. f. - LH1; HH4e;
M(3, 8); sW1a - Endemic to Dhofar, S Yemen and Socotra (Miller, 1996; Thulin at al., 2001).
Cocculus pendulus (J.B.Först. & G.Forst.) Diels - CP6c;
nW(6b, 14o, 20a, 20b, 21, 26, 27).
MORACEAE
Dorstenia foetida (Forssk.) Schweinf. - CP6a; LH(3a, 8b);
HH(4c, 9); nW(17, 18d, 19b, 19c).
Ficus cordata Thunb. subsp. salicifolia (Vahl) C.C.Berg CP(6a, 11, 20); LH9; sW(1b, 2a, 14, 15).
Ficus ingens (Miq.) Miq. - LH(4, 14); HH11; sW16a.
Ficus sycomorus L. - LH14b; Wa8.
Ficus vasta Forssk. - LH(6, 12, 13, 14); HH4c; Wa8.
MORINGACEAE
Moringa peregrina (Forssk.) Fiori - sW(1a, 3, 6a); nW6a.
82
NYCTAGINACEAE
Boerhavia diffusa L. - CP8b.
Boerhavia elegans Choisy subsp. elegans - LH17.
Boerhavia elegans Choisy subsp. stenophylla (Boiss.)
A.G.Mill. - nW(2, 25a, 28).
Commicarpus boissieri (Heimerl) Cufod. - CP(2b, 5a, 6a,
8b, 10a); LH14b; M(1, 18); sW(1a, 1c, 13a, 14, 16a,
20); nW(18b, 18d, 18e).
Commicarpus stenocarpus (Chiov.) Cufod. - CP10b;
LH17; sW(2, 6a, 9, 17, 20); nW(14d, 14e).
Commicarpus sp. plur. - CP6a; M(23b, 23c); sW1b - Incomplete specimens.
NYMPHAEACEAE
Nymphaea nouchali Burm. f. var. caerulea (Savigny)
Verdc. - CP6a - According to McKean (1996) probably
introduced in Yemen - First record from Oman.
OLEACEAE
Jasminum grandiflorum L. subsp. floribundum (R.Br.
ex Fresen.) P.S.Green - LH7f; HH(1, 9, 10); M(1, 10a,
11a, 19).
Olea europaea L. subsp. cuspidata (Wall. ex G.Don) Cif.
- M(11a, 11b, 12, 13, 14, 23b).
OROBANCHACEAE
Cistanche tubulosa (Schenk) Hook. f. - CP(2a, 5a); sW1b.
Cistanche rosea Baker - sW1e; nW(20a, 27).
Orobanche cernua Loefl. - M11b; nW(18a, 18e).
Orobanche dhofarensis M.J.Y.Foley - HH(1, 9); M14.
PAPAVERACEAE
Argemone mexicana L. - HH5a; sW(1b, 12a, 16) - Alien,
native of W Indies and Mexico.
PIPERACEAE
Peperomia pellucida (L.) Kunth - LH8a; HH9.
PLANTAGINACEAE
Plantago ovata Forssk. - M(1, 18); sW(1d, 2b); nW(14d,
14h, 18c).
PLUMBAGINACEAE
Dyerophytum indicum (Gibbs ex Wight) Kuntze - L8a;
CP(4b, 6a, 6c, 15c); HH(4e, 11); sW(1a, 1b, 6a, 13a,
19, 20, 21); nW6b.
Limonium axillare (Forssk.) Kuntze - L8a; CP(2a, 8a, 8d,
10a); LH16; sW(1b, 6c, 13a); nW(1, 14d, 20a, 26);
Ha1c.
Limonium milleri Ghaz. & J.R.Edm. - nW(18a, 18b).
Limonium sarcophyllum Ghaz. & J.R.Edm. - Ha1a - cf.
Raffaelli et al., (2006).
Plumbago zeylanica L. - LH(6, 14); HH(5b, 8, 9).
S. Mosti , M. Raffaelli, M. Tardelli
POLYGALACEAE
Polygala dhofarica Baker - nW(18a, 18d) - Regional endemic: Dhofar & Yemen (Wood, 1997).
Polygala erioptera DC. - CP10a; sW1d; nW(14e; 25b).
Polygala mascatensis Boiss. - sW(1b, 1c, 1i, 2b); Ha(1c, 2).
Polygala moggii Raffaelli, Mosti & Tardelli - sW(18a, 18b,
18c) - Endemic to Dhofar (Raffaelli et al., 2007).
Polygala senensis Klotzsch - CP6a; LH6; HH4b; M14,
sW1e.
Polygala sp. plur. - HH(4c, 4e); M3; sW1d - Incomplete
specimens.
POLYGONACEAE
Persicaria glabra (Willd.) M.Gómez - LH9.
Rumex vesicarius L. - sW(1c, 1d); sW(14e, 17b).
PORTULACACEAE
Portulaca dhofarica M.G.Gilbert - CP(4a, 4b, 10a); sW(6b,
6f); nW18a - Endemic to Dhofar.
Portulaca oleracea L. - sW1e.
Portulaca quadrifida L. - sW1e.
Talinum portulacifolium (Forssk.) Asch. ex Schweinf. sW11.
PRIMULACEAE
Anagallis arvensis L. - HH5a; M(11b, 14, 17); sW12a;
nW(18a, 18c).
RESEDACEAE
Ochradenus arabicus Chaudhary, Hillc. & A.G.Mill. CP(8a, 8c, 8d, 11); LH13; sW(1f, 12b, 8).
Ochradenus gifrii Thulin - nW11 - Regional endemic:
Dhofar & SE Yemen (Miller, 1996; Ghazanfar, 2003). –
(Fig. 3e).
Ochradenus sp. plur. - CP(8a, 8b); sW(9, 18a) - Incomplete
specimens: fruits lacking.
Reseda sphenocleoides Deflers - CP(2a, 2b, 2c); LH2;
sW(1a, 2b, 6b, 12b, 20); nW(6a, 14d, 14e); Ha1a; Wa6.
RHAMNACEAE
Rhamnus staddo A.Rich. - HH4c; M(13, 23c).
Ziziphus leucodermis (Baker) O.Schwartz - sW(1b, 2a,
13a); nW(6b, 25b, 27) - Regional endemic: Dhofar & S
Yemen (Miller & Morris, 1988).
Ziziphus spina-christi (L.) Desf. - CP(1, 11); HH7.
RUBIACEAE
Gaillonia aucheri (Guill.) Jaub. & Spach - M11c; nW(2, 3,
14e, 14o, 20a, 20b, 25a, 26, 27, 28); Ha2.
Kohautia caespitosa Schnizl. - CP(2c, 6a, 6b); LH(5, 8a);
M(1, 3, 11a, 14); sW(1d, 1e, 6b); nW18c.
Kohautia retrorsa (Boiss.) Bremek. - LH16; M6; sW(1c,
6a); nW(14a, 14b, 14i, 14o, 20a, 26).
Contribution to the Flora of Central-Southern Dhofar (Sultanate of Oman)
Oldenlandia corymbosa L. var. corymbosa - HH5a.
Oldenlandia corymbosa L. var. caespitosa (Benth.) Verdc. CP6b; LH(8a, 10); HH1; sW12a.
RUTACEAE
Haplophyllum tuberculatum (Forssk.) A.Juss. - CP(7, 8c);
sW(12a, 12b).
SALVADORACEAE
Azima tetracantha Lam. - LH4; HH(8, 11); M23c. - (Fig.
3f).
Salvadora persica L. var. persica - sW(11, 6a, 6f, 13a);
nW19a; Wa6.
Salvadora persica L. var. angustifolia Verdc. - nW(6a, 6b) First record from Oman - cf. Dahir (1999).
Salvadora persica L. var. cyclophylla (Chiov.) Verdc. - CP(9,
12); sW(13a, 14); nW6b - First record from Oman - cf.
Dahir (1999).
SAPINDACEAE
Allophylus rubifolius (Hochst. ex A.Rich.) Engl. - LH(7a,
7c, 7e, 8b); M(11a, 13, 14).
Dodonaea viscosa (L.) Jacq. - LH13; HH(2, 4e, 8b); M(2,
11a, 19, 23a, 23b, 23c); sW(16a, 16d) - Syn.: D. angustifolia L. f.
SCROPHULARIACEAE
Alectra parasitica Hochst. ex A.Rich. - HH5b.
Anticharis arabica Endl. - sW(2b, 6b).
Anticharis glandulosa Aschers. - CP15d; LH(15, 17);
M11c; sW(18c, 20); nW(2, 25b).
Bacopa monnieri (L.) Pennell - CP(5a, 8a, 6a); LH9; sW(11,
13a, 13d).
Campylanthus chascaniflorus A.G.Mill. - CP(3, 12);
HH4d; sW(8, 13a, 13d); nW(14a, 14e, 14g, 20a).
Campylanthus pungens O.Schwartz - CP2a; sW(1i, 6e, 6f);
nW(14f, 14l, 20a, 20b).
Camptoloma villosa Balf. f. - CP21; sW19.
Lindenbergia indica (L.) Vatke - CP(3, 4a, 8c, 10a); HH11;
M(11a, 11b); sW(1d, 12a, 12b, 8); nW(14d, 14e, 14l,
14o, 17, 18c, 19c, 25b); Wa6.
Lindenbergia muraria (Roxb ex D.Don) Brühl - CP(2b, 2c,
6b); LH3a; HH9; sW16a; Wa(7, 8).
Nanorrhinum hastatum (R.Br. ex Benth.) Ghebr. - CP2c;
LH(11, 12); HH4b; sW1d.
Nanorrhinum heterophyllum (Schousb.) Ghebr. - LH8b;
M11b; sW(2b, 8, 9, 12a); nW(14d, 14o, 18a, 18c, 19a,
19b); Wa6 - Syn.: Kickxia qaraica D.A.Sutton.
Nanorrhinum roseiflorum Mosti, Raffaelli & Tardelli M(5, 7); nW28 - Endemic to Dhofar (Mosti et al., 2008).
Schweinfurthia latifolia Baker ex Oliv. - nW(25a, 27)
- Regional endemic: Dhofar & S Yemen (Mosti et al.,
2006).
83
Schweinfurthia papilionacea (L.) Boiss. - sW19; nW(25a, 27).
Schweinfurthia pedicellata (T.Anderson) Balf. - sW(18b,
20) – cf. Raffaelli et al. (2006).
Schweinfurthia spinosa A.G.Mill., M.Short & D.A.Sutton
- CP4b; nW(14d, 14i, 14l, 14o) - Regional endemic:
Dhofar & SE Yemen (Thulin et al., 2001).
Striga gesnerioides (Willd.) Vatke - CP6a.
SOLANACEAE
Datura metel L. - CP(2a, 6a); LH8a - Syn.: D. fastuosa L.
Hyoscyamus gallagheri A.G.Mill. & Biagi - CP19; HH11.
Lycium shawii Roem. & Schult. - nW25b; Ha1a; nW(6b,
6c, 6d); Wa6.
Nicandra physalodes (L.) Gaertn. - LH(7a, 7b); HH1;
sW12a.
Nicotiana tabacum L. var. tabacum - CP6b.
Nicotiana tabacum L. var. lancifolia (Willd. ex Lehm.)
Comes - CP11; LH8b; M21; nW19a
Physalis minima L. - L1; LH(8a, 10); sW(6b, 12a).
Solanum incanum L. - CP(2a, 6b); LH(3a, 8a); HH7;
M(11a, 11b).
Solanum luteum Mill. - nW15.
Solanum nigrum L. - CP6b; LH8a; HH5a; M11b; nW18c.
Withania qaraitica A.G.Mill. & Biagi - HH(4a, 4b); M11b;
sW1d - Regional endemic: Dhofar & SE Yemen (Thulin
et al., 2001).
Withania somnifera (L.) Dunal - CP2a; LH8a; HH4e; M1;
sW(1b, 12a); nW14g.
STERCULIACEAE
Hermannia paniculata Franch. - CP8b; Ha2; sW(1e, 6b);
nW14e.
Hermannia testacea Vollesen - M18; sW1d.
Melhania muricata Balf. f. - CP8b; M23b; sW(1d, 5, 6b);
Ha1a.
Melhania ovata (Cav.) Spreng. - LH5.
Sterculia africana (Lour.) Fiori - LH(7e, 12); HH10.
TAMARICACEAE
Tamarix aphylla (L.) H.Karst. - HH11; sW2a.
Tamarix arabica Bunge - sW21.
Tamarix mascatensis Bunge - nW6a; Ha1c.
TILIACEAE
Corchorus aestuans L. - sW12a.
Corchorus depressus (L.) C.Chr. - L5a; CP4b; sW(5, 6b, 8).
Corchorus olitorius L. - LH10; HH9; sW1d.
Corchorus trilocularis L. - CP(2c, 4b, 6b, 8b); LH(7b, 11);
sW12a.
Grewia damine Gaertn. - sW(1b, 1c, 1d, 1f, 1i) - Syn.: G.
bicolor Juss.
Grewia erythraea Schweinf. - M(23b, 23c); sW(c, 1f, 1i,
6b); nW(17, 18a).
84
S. Mosti , M. Raffaelli, M. Tardelli
Fig. 3 – a) Iphiona teretifolia Anderb.; b) Pulicaria nobilis Gamal-Eldin; c) Trichodesma cinereum Mosti & Selvi; d) Barleria almughsaylensis Mosti, Raffaelli & Tardelli; e) Ochradenus gifrii Thulin, f ) Azima tetracantha Lam.; g) Tephrosia pentaphylla (Roxb.) G. Don;
h) Boswellia sacra Flueck. (Wadi Dowkah, sito UNESCO).
Contribution to the Flora of Central-Southern Dhofar (Sultanate of Oman)
Grewia mollis A.Juss. - HH11.
Grewia tenax (Forssk.) Fiori - HH11; sW5.
Grewia villosa Willd. - LH8b; sW(1b, 1c).
Triumfetta pentandra A.Rich. ex Guill. - CP6a; LH(8a, 11,
14); HH(1, 4b, 5a, 9, 10).
URTICACEAE
Forsskaolea tenacissima L. - sW(1a, 1b, 1c, 1d, 2b, 20);
nW(6a, 6d, 25a, 25b, 28); Ha1a.
Forsskaolea viridis Ehrenb. - CP(6a, 6b); sW(1d, 16a, 18b).
Laportea interrupta (L.) Chew - LH14b; HH10; Wa7.
Parietaria alsinifolia Delile - nW(14d, 14e) - First record
from Dhofar.
VERBENACEAE
Avicenna marina (Forssk.) Vierh. - Wa(1a, 1b, 5).
Chascanum sessilifolium (Vatke) Moldenke - CP6a; HH4b.
Lantana cf. rugosa Thunb. - M11b - Incomplete specimen.
Lantana viburnoides (Forssk.) Vahl - HH9; M(1,14, 17).
Phyla nodiflora (L.) Greene - L8a; LH9; sW12a; nW20.
Premna resinosa (Hochst.) Schauer - CP(6a, 6b).
Priva cordifolia (L. f.) Druce - HH11.
VIOLACEAE
Viola cinerea Boiss. var. cinerea - sW(1d, 5, 6b); nW(12,
14d, 14f, 14e) - Leaves hairy.
Viola cinerea Boiss. var. stocksii (Boiss.) W.Becker - CP(a,
6b); LH6; HH(5a, 6); M(11b, 14, 17) - Leaves glabrous
or hairy only near the base.
Viola cinerea var. cinerea/var. stocksii - sW(2b, 15) - Hairiness intermediate between the two taxa.
VITACEAE
Cissus quadrangularis L. - CP(6a, 11); LH(6, 9, 13); HH10;
sW(13a, 16f).
Cyphostemma ternatum (Forssk.) Desc. - LH14b; HH8;
M14; nW(14d, 18d).
ZYGOPHYLLACEAE
Fagonia bruguieri DC. - CP4a; sW(1f, 6a, 6c, 20); nW(1,
14d, 25b, 26).
Fagonia cf. bruguieri/schweinfurtii - CP(14, 15a, 15d);
nW(1, 26); Ha1a - Intermediate characters (leaves, spines,
etc.) between the two taxa.
Fagonia indica Burm. f. - sW1e; nW3 - See also F. paulayana.
Fagonia luntii Baker - CP(8b, 8d, 4a, 4b); LH(2, 15); HH4e;
sW(1f, 2b, 6b, 9, 12a, 16a, 16d); nW(18a, 18c, 20a).
Fagonia mahrana Beier - sW(1b, 1c, 1i, 6c, 6e) - Regional
endemic: Dhofar & SE Yemen (Beier, 2002; Ghazanfar,
2007).
Fagonia ovalifolia Hadidi - nW(2, 4, 7, 13, 26).
Fagonia paulayana Wagner & Vierh. - LH15 - Probably no
more than a glabrous form of F. indica (cf. Thulin, 1993;
85
Ghazanfar, 2007).
Fagonia schweinfurthii (Hadidi) Hadidi - CP(2b, 2c, 4b,
8b); sW(1d, 12a, 13a).
Fagonia sp. plur. - nW(7, 14c, 27); Ha2 - Incomplete specimens.
Kelleronia revoilii (Franch.) Chiov. - LH15; HH7a; sW(1d,
2a, 2b) - According to Thulin (1993) the previous records
of K. gillettii from Arabia are actually based on material of
K. revoilii.
Melocarpum robecchii (Engl.) Beier & Thulin - sW(2a,
2b); nW1.
Tetraena alba (L. f.) Beier & Thulin - L8c; sW(1f, 1i, 2a, 3,
6f); Ha2 - Syn.: Zygophyllum album L. f.
Tetraena decumbens (Delile) Beier & Thulin - M(5, 8);
nW(6b, 11, 14d, 14o, 18a, 18c, 20a) - Syn.: Zygophyllum
decumbens Delile.
Tetraena hamiensis (Schweinf.) Beier & Thulin - nW(13,
14d, 14o, 18a, 19b, 20a, 20b); Wa2 - Syn.: Zygophyllum
hamiense Schweinf.
Tetraena qatarensis (Hadidi) Beier & Thulin - nW(25b, 26,
27) - Syn.: Zygophyllum qatarense Hadidi.
Tetraena simplex (L.) Beier & Thulin - L(5a, 7, 8a); CP(4a,
4b, 8c, 8d, 10a, 12, 13, 15b); LH15; sW(11, 16a, 18c);
nW(7, 25b, 26); Ha1a - Syn.: Zygophyllum simplex L.
Tribulus terrestris L. - CP4b; M(1, 23a); sW(1d, 5, 6b,
12a); nW(14d, 18c, 25b).
Tribulus pentandrus Forssk. - nW2.
ANGIOSPERMAE (MONOCOTYLEDONES-LILIOPSIDA)
AGAVACEAE
Dracaena serrulata Baker - HH3; M3.
Sansevieria eherenbergii Schweinf. - CP12; sW(10, 13).
ALOACEAE
Aloe dhufarensis Lavranos - CP8a; LH17; Ha(1a, 1b, 2);
sW(1b, 4, 6a) - Endemic to Dhofar (Miller & Morris,
1988).
Aloe praetermissa T.A.McCoy & Lavranos - CP2a; HH4b;
M3 - (Miller & Morris, 1988: sub A. inermis Forssk.) Endemic to Dhofar, but probably present also beyond the
Oman-Yemen border (Carter et al., 2011).
ARACEAE
Arisaema flavum (Forssk.) Schott - M14.
Remusatia vivipara (Roxb.) Schott - CP6a; HH(5b, 9);
M14.
ARECACEAE (PALMAE)
Nannorrhops ritchieana (Griff.) Aitch. - nW(2, 14b, 20a,
20b, 21); Wa3 - Regional endemic: Dhofar & SE Yemen
(Miller & Morris, 1988; Thulin et al., 2001) - The localities in italics refer to plants observed in the field, of which
we have some photos.
86
Phoenix caespitosa Chiov. - sW1b; nW3; Ha(1b, 1c); Wa8
- First record from Oman.
ASPARAGACEAE
Asparagus africanus Lam. - sW(1a, 1b).
Asparagus racemosus Willd. - CP(2c, 6a); LH(3a, 8a, 14);
HH(1, 4b, 9); M(2, 10a, 10b).
COMMELINACEAE
Aneilema forskaolei Kunth - CP(6a, 6b); LH(3a, 6, 7b, 8a);
HH(9, 10); Wa7.
Commelina albescens Hassk. - CP6a.
Commelina forskaolii Vahl - CP(2c, 6a, 6b, 8b); LH(5, 8a);
sW12a.
Commelina ussilensis Schweinf. - CP2c; nW(18c, 18d).
Commelina sp. plur - CP2c; LH15a - Incomplete specimens.
CYPERACEAE
Cyperus bulbosus Vahl - CP4b; Wa1b - First record from
Dhofar.
Cyperus conglomeratus Rottb. - L(2a, 2c, 5a); CP(5a, 20);
sW(1c, 2b, 6a, 6b, 8, 13a, 13b); nW(14d, 14f).
Cyperus iria L. - CP6b; HH5a; sW12a.
Cyperus laevigatus L. - CP(8a, 8b); sW(11, 13a, 13d);
nW14c; Wa2 - Syn.: Juncellus laevigatus (L.) C.B.Clark.
Cyperus longus L. - LH8a; HH(5a, 5b, 10); M11b.
Cyperus nutans Vahl var. eleusinoides (Kunth) R.W. Haines
- CP6b.
Cyperus rotundus L. - nW25b.
Cyperus squarrosus L. - CP6b.
Cyperus wissmannii O.Schwartz - HH4b.
Cyperus sp. - CP8b.
Eleocharis geniculata (L.) Roem. & Schult. - nW25a.
Fimbristylis cymosa R.Br. subsp. spathacea (Roth) Kojama
- CP(6a, 6b).
Fimbristylis ferruginea (L.) Vahl. subsp. sieberiana (Kunth)
Lye - CP6a; Wa6.
Schoenoplectus subulatus (Vahl) Lye - CP8b; LH8a; sW(3,
11, 13a); nW25b - Syn.: S. litoralis (Shrad.) Palla.
IRIDACEAE
Gladiolus candidus (Rendle) Goldblatt - CP(4b, 6a);
LH(8d, 14b); HH(5a, 5b, 6, 9, 10); M(14, 17) - Syn. G.
ukambanensis (Baker) Marais.
JUNCACEAE
Juncus rigidus Desf. - CP8a; sW11; nW(25a, 25b); Wa6.
ORCHIDACEAE
Epipactis veratrifolia Boiss. & Hohen. - HH11.
Eulophia guineensis Lindl. var. purpurata Kotschy - LH14;
M8a.
S. Mosti , M. Raffaelli, M. Tardelli
Eulophia petersii (Rchb. f.) Rchb. f. - LH1; nW19a.
LILIACEAE
Asphodelus fistulosus L. - M23b; sW1d; nW(14d, 18a, 18c,
19c).
Chlorophytum laxum R.Br. - CP(4b, 6a); LH11.
Dipcadi sp. - CP10a - Incomplete specimen: flowers and
fruits lacking.
POACEAE (GRAMINEAE)
Acrachne racemosa (B. Heyne ex Roem. & Schult.) Ohwi
- CP(1, 6a).
Aeluropus lagopoides (L.) Trin. ex Thwaites - L8a; CP(4a,
5a); sW(1a, 6a, 11); Wa2.
Apluda mutica L. - CP(6a, 6b); LH(3a, 7a, 7b, 8a, 10, 14);
HH(5a, 5b, 8, 9, 10); Wa7.
Aristida abnormis Chiov. - sW(1e, 2b, 6b).
Aristida adscensionis L. - CP(6a, 6b); LH2; sW(1d, 5, 6b,
9); nW14e.
Aristida adscensionis L. var. pumila (Decne.) Coss. & Durieu - nW(14d, 18c).
Aristida funiculata Trin. & Rupr. - CP6a.
Aristida mutabilis Trin. & Rupr. - nW1.
Aristida triticoides Henrard - LH15; sW(1c, 2b, 6f).
Arthraxon cuspidatus (Hochst. ex A.Rich.) Hochst. ex
Hack. - HH(5a, 5b); M14.
Arthraxon hispidus (Thunb.) Makino - HH5a - Racemes
completely glabrous.
Arthraxon lancifolius (Trin.) Hochst. - CP6b; HH(2, 10);
sW1d.
Arthraxon pusillus Bor - CP6b; LH8b; HH(4b, 5a, 10);
sW12a.
Arthraxon prionodes (Steud.) Dandy - M(11b, 17).
Arundinella pumila (Hochst. ex A.Rich.) Steud. - LH7b;
HH(5b, 5c, 9, 10); M(11b, 14).
Brachiaria deflexa (Schumach.) C.E.Hubb. ex Robyns LH8a.
Brachiaria eruciformis (Sm.) Griseb. - M(1, 11b, 14).
Brachiaria ovalis Stapf - CP(2c, 6b); sW6a.
Capillipedium parviflorum (R.Br.) Stapf - M14.
Cenchrus pennisetiformis Hochst. & Steud. - CP(2c, 4b,
6a, 6b, 8b); LH(2, 11, 15, 17); HH9; sW(1a, 1d, 5, 6a,
6b, 9, 11, 12a, 13a, 19, 20); nW(6a, 18a, 18c, 18d, 25b);
Ha1a.
Cenchrus setigerus Vahl - CP(2b, 8b); LH2; HH4b; sW12a.
Chloris virgata Sw. - LH2; sW1d.
Chrysopogon macleishii Cope - nW27.
Chrysopogon plumulosus Hochst. - LH16, M23b; sW2b;
nW(12, 14a, 14d, 20a).
Cymbopogon commutatus (Steud.) Stapf - HH4b - Syn.:
C. schoenanthus (L.) Spreng. subsp. velutinus Cope (Cope,
2007).
Cymbopogon schoenanthus (L.) Spreng. - nW(9, 10, 14a).
Contribution to the Flora of Central-Southern Dhofar (Sultanate of Oman)
Cynodon dactylon (L.) Pers. - CP8a.
Dactyloctenium aegyptium (L.) Willd. - LH(2, 10); HH(5a,
9); sW9; nW14d.
Dactyloctenium aristatum Link - L2d; CP(4b, 8b, 10a, 12);
sW(12a, 13b).
Dactyloctenium robecchii (Chiov.) Chiov. - sW(1e, 2b).
Dactyloctenium scindicum Boiss. - LH2; M(11a, 11b);
sW1d; nW(14d, 17, 18a, 18d).
Dichanthium annulatum (Forssk.) Stapf -HH(2, 5a);
M11a.
Dichanthium foveolatum (Delile) Roberty - CP14; sW(1d,
6a, 6f); nW(14d, 25b).
Dichanthium micranthum Cope - CP(2b, 4b, 6a, 8b, 10a);
M3; sW9.
Digitaria ciliaris (Retz.) Koeler - CP6b; LH8a; HH11;
sW12a; nW18d.
Digitaria nodosa Parl. - CP2c; sW1d.
Digitaria stricta Roth ex Roem. & Schult. - HH5a.
Digitaria velutina (Forssk.) P.Beauv. - HH(9, 10).
Dimeria ornithopoda Trin. - HH5a.
Echinochloa colona (L.) Link - CP6b; LH8a; HH(5a, 5b).
Echinochloa pyramidalis (Lam.) Hitchc. & Chase - LH8a First record from Oman.
Enneapogon cenchroides (Roem. & Schult.) C.E.Hubb. HH11.
Eragrostis aspera (Jacq.) Nees - CP6b; HH11; sW12a.
Eragrostis barrelieri Daveau - sW1d; nW(14d, 18d)
Eragrostis cilianensis (All.) Vignolo ex Janch. - M14 - Specimens with contracted panicles and short spikelets. Leafblades with crateriform glands.
Eragrostis ciliaris (L.) R.Br. - CP6a; LH2; M11b.
Eragrostis minor Host - nW(2, 14d).
Eragrostis tenella (L.) P.Beauv. ex Roem. & Schult. - CP6b;
HH9; sW12a.
Eustachys paspaloides (Vahl) Lanza & Mattei - LH8a;
HH(9, 10); M11b.
Halopyrum mucronatum (L.) Stapf - L(5a, 5b).
Heteropogon contortus (L.) P.Beauv. ex Roem. & Schult. HH9; M14; sW1d.
Isachne globosa (Thunb. ex Murray) Kuntze - LH9.
Lasiurus scindicus Henrard - nW(2, 7, 26).
Leptothrium senegalense (Kunth) Clayton - CP10a; sW(1d,
2b, 6b).
Loudetia flavida (Stapf) C.E.Hubb. - HH(9, 11); M14;
sW1e - Glumes completely glabrous.
Oplismenus burmannii (Retz.) P.Beauv. - LH(7b, 14);
HH(9, 10); M14.
Panicum trichoides Sw. - LH(7b, 8a); HH9.
Panicum atrosanguineum Hochst. ex A.Rich. - CP(2c, 6b);
LH8a; sW6a.
Pennisetum divisum (J.F.Gmel.) Henrard - CP20; HH11;
87
nW(2, 25b).
Pennisetum setaceum (Forssk.) Chiov. - HH11.
Phragmites australis (Cav.) Trin. ex Steud. subsp. australis
- nW(6a, 6d).
Phragmites karka (Retz.) Trin. ex Steud. - CP20; nW25b;
Wa(2, 6) - First record from Oman.
Poa bulbosa L. var. vivipara Koeler - HH4c - First record
from Oman.
Rottboellia cochinchinensis (Lour.) Clayton - HH9.
Setaria intermedia Roem. & Schult. - LH7b; HH(5a, 9,
10).
Setaria pumila (Poir.) Roem. & Schult. - CP6b; LH(8a, 10);
HH(5a, 5b, 9); M14.
Setaria verticillata (L.) P.Beauv. - HH(9, 10); nW(17, 18d,
18e).
Setaria viridis (L.) P.Beauv. - CP(6a, 6b); HH5a; nW(14d,
18a, 18c).
Setaria cf. italica (L.) P.Beauv. - CP6b - Incomplete specimen.
Sorghum deccanense Stapf ex Raizada - HH9.
Sporobolus consimilis Fresen. - Wa6.
Sporobolus ioclados (Nees ex Trin.) Nees - CP(2b, 5a, 8a,
10a, 12); sW(11, 13a).
Sporobolus minimus Cope - sW(12a, 13b).
Sporobolus spicatus (Vahl) Kunth - sW13a; nW25b; Wa2.
Sporobolus virginicus (L.) Kunth - L(2c, 3a, 3b); CP(4a, 5a,
8a, 12); sW13a.
Stipagrostis hirtigluma (Steud. ex Trin. & Rupr.) De Winter
- HH11; sW1d.
Stipagrostis plumosa (L.) Munro ex T.Anderson - sW9;
nW14d.
Stipagrostis sokotrana (Vierh.) De Winter - nW(14d, 14e).
Stipagrostis sp. - nW14d - Incomplete specimen.
Tetrapogon tenellus (J.König ex Roxb.) Chiov. - CP(6a,
6b); LH2.
Tetrapogon villosus Desf. - sW(6f, 8).
Themeda quadrivalvis (L.) Kuntze - CP6b; HH(2, 5a, 9).
Tragus berteronianus Schult. - CP4b.
Tricholaena teneriffae (L. f.) Link - CP8a.
Urochloa panicoides P.Beauv. - LH8a; HH9; sW12a;
nW18c.
Urochondra setulosa (Trin.) C.E.Hubb. - L(2a, 2c, 3b);
CP(2b, 4a, 5a, 10a); sW(13b, 13d).
POTAMOGETONACEAE
Potamogeton nodosus Poir. - LH9; sW12a.
TYPHACEAE
Typha domingensis Pers. - nW25b; Wa6 - Syn.: T. angustata
Bory & Chaub.
88
S. Mosti , M. Raffaelli, M. Tardelli
Analysis of the flora
The amount of Families, Genera and Species of the present floristic contribution is reported below:
Pteridophyta
Gymnospermae
Dicotyledones
Monocotyledones
Total
Families
5
1
63
14
Genera
6
1
256
66
Species
7
1
476
122
83
323
606
The collected species correspond about to 69%
of the floristic richness of Dhofar (Ghazanfar, 2003).
The twelve families with the highest number of species are: Poaceae (85 taxa), Fabaceae (45), Asteraceae
(44), Euphorbiaceae (27), Apocynaceae incl. Asclepiadaceae (25), Acanthaceae (24), Boraginaceae (23),
Malvaceae (20), Capparaceae (17), Convolvulaceae
(17), Scrophulariaceae (17) and Zygophyllaceae (16).
The floristic list includes taxa found for the first
time in Oman (17 taxa) or Dhofar (13 taxa) and 6
species new to science published in previous works:
Desmidorchis tardellii, Trichodesma cinereum, Nanorrhinum roseiflorum, Polygala moggii, Orbea nardii and
Barleria almughsaylensis (see references cited in the
floristic list).
First records from Oman are: Barleria parviflora,
Blumea axillaris, Echinochloa pyramidalis, Euphorbia
granulata var. granulata, Heliotropium steudneri, Hildebrandtia somalensis, Iphiona teretifolia, Nymphaea
nouchali var.coerulea, Phragmites karka, Phoenix
caespitosa, Poa bulbosa var. vivipara, Pulicaria gamaledinae, Salvadora persica var.angustifolia and var. cyclophylla, Thespesia populnea, Tridax procumbens, Vernonia cinerascens.
First records from Dhofar are: Amaranthus dubius, Ceratophyllum submersum, Cleome austroarabica
subsp. muscatensis, Cleome scaposa, Cyperus bulbosus,
Echinops spinosissimus, Euphorbia serpens, Hibiscus micranthus, Parietaria alsinifolia, Pycnocycla aucheriana,
Salvia macilenta, Senna sophera, Tephrosia pentaphylla.
From the floristic point of view, the south slopes
of the coastal mountains, influenced by the SW monsoon, are the most interesting environments in Centre-south Dhofar. Plant richness and endemic taxa
are concentrated in this area. The local endemisms
(endemisms of Dhofar) are represented by 14 taxa,
while the regional endemics (distributed in Oman/
Dhofar and Yemen) are 35 taxa; in all the percentage
of endemisms is about 8,1% of the collected flora.
Local endemics (Dhofar) - 2,3% of the total flora
Aerva arthemisioides subsp. batharitica
Aloe dhufarensis
Aloe praetermissa Barleria almughsaylensis
Desmidorchis tardellii
Dyschoriste dalyi
Echidnopsis scutellata subsp. dhofarensis
Kleinia saginata
Lavandula dhofarensis subsp. ayunensis
Nanorrhinum roseiflorum
Orbea nardii
Polygala moggii
Portulaca dhofarica
Trichodesma cinereum
Regional endemics (Oman/Dhofar & Yemen) 5,8% of the total flora
Anogeissus dhofarica - Dhofar, Yemen (Mahrah region)
Becium dhofarense - Dhofar, SE Yemen (Mahrah region)
Bentia fruticulosa - Dhofar, Yemen (Hadramaut)
Blepharis dhofarensis - Dhofar, Yemen (Mahrah region)
Blepharispermum hirtum - Dhofar, SE Yemen (Mahrah
region)
Cleome albescens subsp. omanensis - Dhofar, Yemen
(Mahrah region)
Desmidorchis adenensis - Oman, Yemen
Desmidorchis flavus - Oman, Yemen
Dhofaria macleishii - Dhofar, Yemen (Mahrah region)
Euphorbia dhofarensis - Dhofar, SE Yemen
Euphorbia orbiculifolia - Dhofar, SE Yemen
Euphorbia smithii - Dhofar, Yemen (Mahrah region)
Exacum arabicum - Dhofar, S Yemen
Fagonia mahrana - Dhofar, SE Yemen
Farsetia dhofarica - Oman, S Yemen
Contribution to the Flora of Central-Southern Dhofar (Sultanate of Oman)
Gymnocarpos dhofarensis - Dhofar, S Yemen
Herniaria maskatensis - Oman, S Yemen
Iphiona teretifolia - Dhofar, Yemen
Jatropha dhofarica - Dhofar, Yemen (Mahrah region)
Lavandula dhofarensis subsp. dhofarensis - Dhofar, Yemen (Mahrah region)
Leucas dhofarensis - Dhofar, Yemen (Mahrah region)
Maytenus dhofarensis - Dhofar, SE Yemen
Nanorrhops ritchieana - Dhofar, SE Yemen
Ochradenus gifrii - Dhofar, SE Yemen
Orbea luntii - Dhofar & SE Yemen
Ormocarpum dhofarense - Dhofar, Yemen (Mahrah region)
Pentatropis bentii - Dhofar, Yemen
Polygala dhofarica - Dhofar, Yemen
Pulicaria gamal-eldinae - Dhofar, S Yemen
Pulicaria omanensis subsp. omanensis - Dhofar, Yemen
(Mahrah region)
Rhytidocaulon fulleri - Oman, Yemen
Scheiwenfurthia latifolia - Dhofar, S Yemen (Hadramaut)
Scheiwenfurthia spinosa - Dhofar, S Yemen (Hadramaut)
Ziziphus leucodermis - Dhofar, S Yemen
Whitania qaraitica - Dhofar, Yemen (Mahrah region)
From the phytogeographical point of view, most
of the plants found in Centre-south Dhofar belong
to the Somalia-Masai regional centre of endemism,
i.e. to the phytogeographical region (the Somalia-
89
Masai) that from East Africa spreads north-eastwards
to embrace the whole south Dhofar including the
coastal plain, the monsoon affected south mountain
sides and also the drier mountain slopes facing north
(see also Ghazanfar, 1992b, 2003). Only a few species
come from the pre-desert and desert environments,
which expand north of the coastal mountains. Here
the soil is scattered with stones and pebbles and is
cut by many wadis draining north towards the Saudi
Arabian border. Those species belong to the SaharoSindian phytochorion which includes the Arabianregional subzone that embraces the Centre-north
Dhofar and Centre Oman. Here the soil is very dry,
only a few species are present and the vegetation is
poor and formed by xerophytes and psammophytes. Among the plants sampled in this area there are
Centaurea pseudo-sinaica, Lasiurus scindicus, Rhazia
stricta, Suaeda aegyptiaca, Fagonia sp.pl., Polyacarpea
repens, Stipagrostis sp.pl., Tamarix sp.pl.
Acknowledgements
Thanks are due to the Office of the Adviser for Cultural Affairs, Sultanate of Oman, for the technical and
logistic assistance during our missions to Dhofar.- Our
thanks also to Dr. Milena Rizzotto who revised the English manuscript.
REFERENCES
ANDERBERG A., 1985 – The genus Iphiona (CompositaeInuleae). Nordic. J. Bot. 5(2): 169-194.
BEENTJE H., 2006 – Asteraceae. In: Thulin M. (Ed.), Flora
Somalia. 3. Royal Botanic Gardens, Kew.
BEIER B.-A., 2002 – Two new unifoliate species of Fagonia (Zygophyllaceae) from the Horn of Africa region, and
resurrection of Fagonia subinermis from Iran. Nordic J.
Bot. 21(5): 449-455. (2001)
BRUMMIT R.K., POWELL C.E., 1992 – Authors of plant
names. Royal Botanic Gardens, Kew.
BRUYNS V., 1988 – A revision of the genus Echidnopsis Hook.
F. (Asclepiadaceae). Bradleya 6: 1-48.
CARTER S., LAVRANOS J.J., NEWTON L.E. & WALKER
C.C., 2011 – Aloes. The Definitive Guide. Royal Botanic Gardens, Kew.
COPE T.A., 2007 – Flora of the Arabian Peninsula and Socotra. 5(1). Edinburgh University Press.
Dahir A.M., 1999 – Salvadoraceae. In: Thulin M. (Ed.),
Flora Somala. 2. Royal Botanic Gardens, Kew.
GAMAL-ELDIN E., 1981 – Revision der Gattung Pulicaria
(Compositae-Inuleae) fur Africa, Makaronesien und Arabien. Phanerogamarum Monographiae, Tomus XIV. J.
Cramer Ed. Vaduz.
GHAZANFAR S.A., 1992a – An annotated Catalogue of the
Vascular Plants of Oman and their Vernacular Names.
Scripta Botanica Belgica 2: 1-153.
GHAZANFAR S.A., 1992b – Quantitative and biogeographic analysis of the flora of the Sultanate of Oman. Glob.
Ecol. Biogeogr. Letters 2: 189-195.
GHAZANFAR S.A., 2003 – Flora of the Sultanate of Oman.
1 Piperaceae-Primulaceae. Meise. National Botanic Garden of Belgium. (Scripta Botanica Belgica 25, pp. 262).
GHAZANFAR S.A., 2007 – Flora of the Sultanate of Oman.
2 Crassulaceae-Apiaceae. Meise. National Botanic Garden of Belgium. (Scripta Botanica Belgica 36, pp. 220).
GHAZANFAR S.A., 2010 – Trichodesma dhofaricum (Bor-
90
aginaceae), a new species from the Sultanate of Oman.
Kew Bulletin 65: 97-100.
KILIAN N., 1999 – Studies on the Compositae of the Arabian Peninsula and Socotra - 1. Pulicaria gamal-eldinae
sp. nova (Inuleae) bridges the gap between Pulicaria and
former Sclerostephane (now sect. Sclerostephane). Willdenowia 29: 167-185
LAVRANOS J.J. & MORTIMER K.V., 1970 – Rhytidiocaulon
fulleri, a new Stapeliad from the Dhofar coast, Muscat
and Oman, Southerb Arabia. Nat. Cact. Succ. Journ.
25(1): 2-4.
MCKEAN D.R., 1996 – Nymphaeaceae. In: Miller A.G.,
Cope T.A. (Eds.), Flora of the Arabian Peninsula and
Socotra. 1. Edinburgh University Press.
MEVE U. & LIEDE S., 2002 – A molecular phylogeny and
generic rearrangement of the stapelioid Ceratopegieae
(Apocynaceae-Aslepiadoideae). Plant Syst. Evol. 234:
171-209.
MILLER A.G., 1996 – Amaranthaceae. In: MILLER A.G.,
COPE T.A. (Eds.), Flora of the Arabian Peninsula and
Socotra. 1. Edinburgh University Press.
MILLER A.G., 1996 – Brassicaceae. In: Miller A.G., COPE
T.A. (Eds.), Flora of the Arabian Peninsula and Socotra.
1. Edinburgh University Press.
MILLER A.G., 1996 – Menispermaceae. In: MILLER A.G.,
COPE T.A. (Eds.), Flora of the Arabian Peninsula and
Socotra. 1. Edinburgh University Press.
MILLER A.G., 1996 – Resedaceae. In: MILLER A.G., COPE
T.A. (Eds.), Flora of the Arabian Peninsula and Socotra.
1. Edinburgh University Press.
MILLER A.G. & MORRIS M., 1988 – Plants of Dhofar. The
Office of The Adviser for Conservation of the Environment, Diwan of Royal Court, Sultanate of Oman.
MOSTI S. & RAFFAELLI M., 2004 – Desmidorchis tardellii (Asclepiadaceae), a new species from Dhofar (Southern
Oman). Webbia 59(2): 285-291.
MOSTI S., RAFFAELLI M., 2011 – Orbea nardii and O.
wissmannii var. parviloba are different taxa. Nomenclatural rearrangement in Orbea wissmannii: subsp. wissmannii, stat. nov., subsp. parviloba, stat. nov. and subsp.
eremastrum, stat. nov. Asklepios 110: 13-15.
MOSTI S., RAFFAELLI M. & TARDELLI M., 2005 – Le Succulente del Dhofar (Oman meridionale).1. Cactus &
Co. vol. 9(4): 218-236.
MOSTI S., RAFFAELLI M. & TARDELLI M., 2006 – A contribution to the flora of Wadi Andur (Dhofar, Southern
Oman). Webbia 61(2): 253-260.
MOSTI S., RAFFAELLI M. & TARDELLI M., 2008 – Nanorrhinum roseiflorum (Scrophulariaceae), a new species
from Dhofar, Southern Oman. Webbia 63(2): 49-54.
MOSTI S., RAFFAELLI M. & TARDELLI M., 2011 – Flora
S. Mosti , M. Raffaelli, M. Tardelli
of Oman: Barleria almughsaylensis (Acanthaceae), a new
species from Dhofar (Southern Oman). Candollea 66(1):
1-7.
MOSTI S. & SELVI F., 2007 – Trichodesma cinereum (Boraginaceae), a new species from Oman. Candollea 62(2):
205-210.
PETRUSSON L. & THULIN M., 1996 – Gymnocarpos. In:
MILLER A.G., COPE T.A (Eds.), Flora of the Arabian
Peninsula and Socotra. 1. Edinburgh University Press.
PILBEAM J., 2010 – Stapeliads. A gallery of 686 pictures of
all 57 genera of this popular group of succulent plants. Pp.
207. British Cact. Succ. Soc. Wakefield, UK.
RADCLIFFE-SMITH A., 1980 – The vegetation of Dhofar.
In: Scientific Results of the Oman Flora and Fauna Survey 1977 (Dhofar). Jour. Oman Stud. special report no.
2: 58-86.
RAFFAELLI M., MOSTI S. & TARDELLI M., 2003 – The
Frankincense Tree (Boswellia sacra Flueck., Burseraceae)
in Dhofar, southern Oman:field-investigations on the
natural population. Webbia 58(1): 133-149.
RAFFAELLI M., MOSTI S. & TARDELLI M., 2006 – Boswellia sacra Flueck. (Burseraceae) in the Hasik area
(Eeastern Dhofar, Oman). Webbia 61(2): 245-251.
RAFFAELLI M., MOSTI S. & TARDELLI M., 2007 – Polygala moggii (Polygalaceae), a new species from Oman.
Webbia 62(2): 269-274.
RAFFAELLI M., MOSTI S. & TARDELLI M., 2008 – Apocynaceae from Oman: Orbea nardii, sp. nov.; Orbea luntii
and Pentatropis bentii, first finding. Webbia 63(2): 161167.
RAFFAELLI M., TARDELLI M. & MOSTI S., 2003 – The Wadi
Doka Frankincense Park in Dhofar, Oman. First steps towards the safeguard and improvement of Boswellia sacra
(Burseraceae). Centro Studi Erbario Tropicale, Università di Firenze, pubbl. n. 93.
THULIN M., 1993 – Zygophyllaceae. In: THULIN M. (Ed.),
Flora Somalia. 1. Royal Botanic Gardens, Kew.
THULIN M., 2001 – Exacum (Gentianaceae) on the Arabian
peninsula and Socotra. Nordic J. Bot. 21: 243-247.
THULIN M., AL.GIFRI A., HUSSEIN M.A. & GABALI S.,
2001 – Addition to the Yemen Flora - In: FRIIS IB, RYDING O. (Eds), Biodiversity Research in the Horn of Africa
Region. Proceedings of the Third International Symposium on the Flora of Ethiopia and Eritrea at the Carlsberg Academy, Copenhagen, August 25-27, 1999.
(Biologiske Skrifter 54: 137-153).
WARFA A.M., 1990 – Taxonomy and distribution of Cordia
sinensis and C. nevillii (Boraginaceae), a widespread species pair in Africa and Asia. Nordic J. Bot. 9: 649-656.
WOOD J.R.I., 1997 – A handbook of the Yemen Flora. Royal
Botanic Garden, Kew.
Contribution to the Flora of Central-Southern Dhofar (Sultanate of Oman)
91
Summary: From 2000 to 2009 we carried out 10 scientific missions to Central-Southern Dhofar with the aim of collecting plants. We eplored several habitats, such as littorals and seashores, coastal plains and many wadis ending in the
Arabian sea (from Rackhyt, Al Mughsayl, Salalah, Khor Rori, Mirbat, Hadbin up to Hasik). We paid particular attention
to the floristic exploration of the monsoon slopes of the coastal mountains (with special reference to Jabal Qara) where
plant richness and endemic taxa are mostly concentrated. Plants were also collected on the upper part of Jabal Samhan,
the highest mountain of Dhofar, in the pre-desertic area of Wadi Andur, beyond Jabal Samhan and in several areas where,
nowadays, large groups of Boswellia sacra, the frankincense tree, grow, i.e. Al Mughsayl, Adownib, Aluf, Uyun, Wadi
Dowkah and Hasik. Sporadic explorations were carried out on the vaste pre-desertic area that, from Thumrayt, spreads
northward to the Saudi Arabia border, from Muddy, Al Mathafah, Ginan Bin Nawatish to Shisr (the ancient Ubar), Sayh
al Kharyat and Dawkah.
The number of explored sites (233) was large enough to have exhaustive information on the local distribution of the
species. The collected taxa are 606; 17 of them are first records from Oman, 13 are first record from Dhofar. As regards
endemics, 14 taxa are endemic to Dhofar, while 35 are regional endemics, distributed in Oman/Dhofar & Yemen; most
of these are concentrated on the monsoon slopes of the coastal mountains.