ACARORUM CATALOGUS IX ACARIFORMES Acaridida Petar Beron ACARORUM CATALOGUS Petar Beron ACARORUM CATALOGUS IX Acariformes Acaridida Schizoglyphoidea (Schizoglyphidae) Histiostomatoidea (Histiostomatidae, Guanolichidae) Canestrinioidea (Canestriniidae, Chetochelacaridae, Lophonotacaridae, Heterocoptidae) Hemisarcoptoidea (Chaetodactylidae, Hyadesiidae, Algophagidae, Hemisarcoptidae, Carpoglyphidae, Winterschmidtiidae) Petar Beron Pensoft National Museum of Natural History, Sofia Bulgarian Academy of Sciences Sofia–Moscow 2021 ACARORUM CATALOGUS IX Acariformes Acaridida Schizoglyphoidea (Schizoglyphidae) Histiostomatoidea (Histiostomatidae, Guanolichidae) Canestrinioidea (Canestriniidae, Chetochelacaridae, Lophonotacaridae, Heterocoptidae) Hemisarcoptoidea (Chaetodactylidae, Hyadesiidae, Algophagidae, Hemisarcoptidae, Carpoglyphidae, Winterschmidtiidae Petar Beron First published 2021 ISBN 978-619-248-057-8 (paperback) ISBN 978-619-248-058-5 (e-book) Pensoft Series Faunistica No 224 Beron P. 2021. Acarorum Catalogus IX. Schizoglyphoidea (Schizoglyphidae). Histiostomatoidea (Histiostomatidae, Guanolichidae). Canestrinioidea (Canestriniidae, Chetochelacaridae, Lophonotacaridae, Heterocoptidae). Hemisarcoptoidea (Chaetodactylidae, Hyadesiidae, Algophagidae, Hemisarcoptidae, Carpoglyphidae, Winterschmidtiidae. Pensoft & National Museum of Natural History, Sofia, 396 pp. © PENSOFT Publishers © National Museum of Natural History, Sofia Pensoft Publishers Prof. Georgi Zlatarski 12, Sofia 1700, Bulgaria info@pensoft.net www.pensoft.net Printed in Bulgaria, December 2021 Contents Introduction ............................................................................................................................... 9 Acariformes, Acaridida...........................................................................................................10 The Mite Superfamily Schizoglyphoidea ..............................................................................10 The Mite Family Schizoglyphidae .........................................................................................10 Catalogue of Schizoglyphidae ........................................................................................10 Index of Schyzoglyphidae ...............................................................................................10 Distribution of Schizoglyphidae ....................................................................................11 Hosts of Schizoglyphidae ...............................................................................................11 Bibliography of Schizoglyphidae ...................................................................................11 The Mite Superfamily Histiostomatoidea (Anoetoidea) ....................................................11 The Mite Family Histiostomatidae (Anoetidae)..................................................................11 Composition of Histiostomatidae (Anoetidae) ...........................................................12 Catalogue of Histiostomatidae ......................................................................................14 Index of Histiostomatidae ..............................................................................................98 Distribution of Histiostomatidae ................................................................................108 Hosts of Histiostomatidae ............................................................................................115 Bibliography of Histiostomatidae ................................................................................122 The Mite Family Guanolichidae Fain..................................................................................141 Composition of Guanolichidae ...................................................................................141 Catalogue of Guanolichidae .........................................................................................142 Index of Guanolichidae ................................................................................................143 Distribution of Guanolichidae .....................................................................................143 Hosts of Guanolichidae.................................................................................................143 Bibliography of Guanolichidae ....................................................................................143 The Mite Superfamily Canestrinioidea ...............................................................................144 The Mite Family Canestriniidae ..........................................................................................144 Composition of Canestriniidae ...................................................................................145 Catalogue of the Family Canestriniidae .....................................................................147 Index of Canestriniidae ................................................................................................213 Distribution of Canestriniidae .....................................................................................219 6 Acarorum Catalogus IX Hosts of Canestriniidae ................................................................................................225 Bibliography of Canestriniidae ....................................................................................236 The Mite Family Chetochelacaridae Fain ...........................................................................243 Index of Chetochelacaridae..........................................................................................244 Distribution of Chetochelacaridae ..............................................................................244 Hosts of Chetochelacaridae..........................................................................................244 Bibliography of Chetochelacaridae .............................................................................244 The Mite Family Lophonotacaridae Fain ...........................................................................244 Index of Lophonotacaridae ..........................................................................................245 Distribution of Lophonotacaridae...............................................................................245 Hosts of Lophonotacaridae ..........................................................................................245 Bibliography of Lophonotacaridae ..............................................................................245 The Mite Family Heterocoptidae Fain ................................................................................246 Composition of Heterocoptidae ..................................................................................246 Catalogue of Heterocoptidae .......................................................................................246 Index of Heterocoptidae ...............................................................................................253 Distribution of Heterocoptidae ...................................................................................254 Hosts of Heterocoptidae ...............................................................................................255 Bibliography of Heterocoptidae...................................................................................256 The Mite Superfamily Hemisarcoptoidea...........................................................................257 The Mite Family Chaetodactylidae .....................................................................................258 Composition of Chaetodactylidae ..............................................................................258 Catalogue of Chaetodactylidae ....................................................................................258 Index of Chaetodactylidae............................................................................................284 Distribution of Chaetodactylidae ................................................................................286 Hosts of Chaetodactylidae............................................................................................290 Bibliography of Chaetodactylidae ...............................................................................293 The Mite Family Hyadesiidae ..............................................................................................303 Composition of Hyadesiidae........................................................................................303 Catalogue of Hyadesiidae .............................................................................................304 Index of Hyadesiidae .....................................................................................................310 Distribution of Hyadesiidae .........................................................................................311 Bibliography of Hyadesiidae ........................................................................................312 The Mite Family Algophagidae Fain ...................................................................................315 Acarorum Catalogus IX 7 Composition of Algophagidae .....................................................................................316 Catalogue of Algophagidae ..........................................................................................316 Index of Algophagidae ..................................................................................................321 Distribution of Algophagidae ......................................................................................322 Hosts of Algophagidae ..................................................................................................323 Bibliography of Algophagidae .....................................................................................323 The Mite Family Hemisarcoptidae ......................................................................................326 Composition of Hemisarcoptidae ...............................................................................326 Catalogue of Hemisarcoptidae.....................................................................................326 Index of Hemisarcoptidae ............................................................................................332 Distribution of Hemisarcoptidae.................................................................................332 Hosts of Hemisarcoptidae ............................................................................................334 Bibliography of Hemisarcoptidae ................................................................................335 The Mite Family Carpoglyphidae........................................................................................339 Composition of Carpoglyphidae .................................................................................339 Catalogue of Carpoglyphidae ......................................................................................340 Index of Carpoglyphidae ..............................................................................................342 Distribution of Carpoglyphidae ..................................................................................342 Hosts of Carpoglyphidae ..............................................................................................343 Bibliography of Carpoglyphidae..................................................................................343 The Mite Family Winterschmidtiidae (Saproglyphidae) ..................................................344 Composition of Winterschmidtiidae ..........................................................................345 Catalogue of Winterschmidtiidae ...............................................................................346 Index of Winterschmidtiidae .......................................................................................377 Distribution of Winterschmidtiidae ...........................................................................380 Hosts of Winterschmidtiidae .......................................................................................384 Bibliography of Winterschmidtiidae...........................................................................387 Acknowledgements ...............................................................................................................396 Acarorum Catalogus IX 9 Introduction The present 9th volume of the series Acarorum Catalogus contains lists of mites of 13 families, 225 genera and 1268 species of the superfamilies Schizoglyphoidea, Histiostomatoidea, Canestrinioidea and Hemisarcoptoidea. Most of these mites live on insects or other animals (as parasites, phoretic or commensals), some inhabit rotten plant material, dung or fungi. Mites of the families Chetochelacaridae and Lophonotacaridae are specialised to live with Myriapods (Diplopoda). The peculiar aquatic or intertidal mites of the families Hyadesidae and Algophagidae are also included. 10 Acarorum Catalogus IX Acariformes, Acaridida The Mite Superfamily Schizoglyphoidea Contains so far only one family of mites, found in New Guinea on Coleoptera. The postnymphal stage remains unknown. The Mite Family Schizoglyphidae Schizoglyphidae Mahunka, 1978: 115 Monotypic genus of mites found on Tenebrionidae beetles. Catalogue of Schizoglyphidae Fam. Schizoglyphidae Schizoglyphidae Mahunka, 1978: 115 Type genus: Schizoglyphus Mahunka, 1978 Genus Schyzoglyphus Mahunka Schyzoglyphus Mahunka, 1978: 115 Type species: Schizoglyphus biroi Mahunka, 1978 Schizoglyphus biroi Mahunka Schizoglyphus biroi Mahunka, 1978: 115 Type locality: Geelvink [Bay] Distribution: Indonesia (Western New Guinea) Host: Tagalus tibialis (Coleoptera: Tenebrionidae) Index of Schyzoglyphidae (Names in bold are accepted as valid) biroi, Schizoglyphus 10 Schizoglyphidae 10 Schizoglyphus 10 Schizoglyphoidea 10 Acarorum Catalogus IX 11 Distribution of Schizoglyphidae Indonesia (Mahunka, 1978) – Schizoglyphus biroi [Western New Guinea] Hosts of Schizoglyphidae Insecta Coleoptera Fam. Tenebrionidae Tagalus tibialis – Schizoglyphus biroi Bibliography of Schizoglyphidae Mahunka S. 1978. Schizoglyphidae fam. n. and new taxa of Acaridae and Anoetidae (Acari: Acarida). – Acta Zoologica Academiae Scientiarum Hungaricae, 24(1–2): 115. The Mite Superfamily Histiostomatoidea (Anoetoidea) The Superfamily contains only one family - Histiostomatidae (Anoetidae). The Mite Family Histiostomatidae (Anoetidae) Histiostomatidae Berlese, 1897 Anoetidae Oudemans, 1904; Hughes & Jackson, 1958 Small or middle-sized mites (250–900 µm), living in rotten plant material, in the dung and fungi, some of them on animals (Hirudinea, Crustacea, Myriapoda, Insecta, Mammalia). Their study starts with the description of Hypopus feroniarum Dufour, 1839, Hypopus alicola Dujardin, 1842 and the first genera Anoetus Dujardin, 1842 and Histiostoma Kramer, 1876. Follow the many publications of Oudemans, who described in 1904 the family Anoetidae. However, OConnor (1984) insisted that the family name Histiostomatidae Berlese, 1897 antedates Anoetidae Oudemans, 1904, and since than the widely known name Anoetidae has been replaced. Vitzthum (1931) composed 12 Acarorum Catalogus IX the review of 10 genera of Anoetidae. Two revisions of the family took place almost simultaneously: of Scheucher (1957) and of Hughes & Jackson (1958). Until the revision of Scheucher (1957) only 12 valid genera have been described, seven of them by Oudemans: Anoetus Dujardin, 1842, Histiostoma Kramer, 1876, Zwickia Oudemans, 1924, Anoetoglyphus Vitzthum, 1927, Glyphanoetus Oudemans, 1929, Myianoetus Oudemans, 1929, Seliea Oudemans, 1929, Munduytia Oudemans, 1929, Cederhjelmia Oudemans, 1931, Creutzeria Oudemans, 1932, Chiropteranoetus Womersley, 1942, Prowichmannia Radford, 1950. From the 45 genera described after 1950 16 are due to A. Fain and co-authors and 20 to S. Mahunka and co-authors (most of them monospecific) . The other major contributors are Sevastyanov, OConnor, Southcott, Fashing, Woodring, Samšinák, Nesbitt, Ashfaq & Sarwar, Ashfaq, Chaudhri & Aheer, Eraky, and others. From the 58 genera 34 are monotypic. Only six genera include more than 10 sp. – Histiostoma (236, Anoetus (93), Myianoetus (48), Anoetoglyphus (15), Glyphanoetus (14), and Rhopalanoetus (11). Still exists the Anoetus – Histiostoma problem. According to Woodring & Moser (1970),”…there is sufficient distinction between the concept of Anoetus and Histiostoma to warrant the maintenance of the separate groups at the generic level. In this case the older name Anoetus has priority over Histiostoma, and Histiostoma Kramer, 1876 must be synonymized under Anoetus Dujardin, 1842, which is the type and name-bearing genus of the family Anoetidaе”. However, for Sevastianov (1975) Histiostoma is junior homonyme of Anoetus. Composition of Histiostomatidae (Anoetidae) Fam. Histiostomatidae (Anoetidae) - 58 gen., 522 sp. Subfam. Ceylanoetinae Mahunka, 1974–6 gen., 15 sp. Ceylanoetus Mahunka, 1974–1 sp. Chiropteranoetus Womersley, 1942–1 sp. Conglanoetus Mahunka, 1978 (= Comyianoetus Fain et Philips, 1979) – 3 sp. Capronomoia Mahunka, 1976–7 sp. Creutzeria Oudemans, 1932–2 sp. Curculanoetus Fain, 1974–1 sp. Subfam. Fibulanoetinae Fain, Camerik, Lukoschus et Kniest, 1980–3 gen., 18 sp. Fibulanoetus Mahunka, 1973–3 sp. Ghanoetus Mahunka, 1973–1 sp. Glyphanoetus Oudemans, 1929–14 sp. Subfam. Histiostomatinae Berlese, 1897 (Anoetinae Oudemans, 1904) - 46 gen., 489 sp. Ameranoetus Ide et Mahunka, 1978–1 sp. Acarorum Catalogus IX 13 Amyzanoetus Fain, 1976–1 sp. Ancyranoetus Fain et Santiago-Blay, 1993–1 sp. Anoetoglyphus Vitzthum, 1927–15 sp. Anoetus Dujardin, 1842–93 sp. (?)(= Nodipalpus Karpelles, 1893) Aphodanoetus Bongers, OConnor et Lukoschus, 1985–1 sp. Auricanoetus Fain et Zumpt, 1974–1 sp. Austranoetus Fain, 1976–1 sp. Bonomoia Oudemans, 1911–9 sp. Bothyanoetus Fain et Camerik, 1978–1 sp. Cederhjelmia Oudemans, 1931–1 sp. Chiloanoetus Fain, 1974–1 sp. Hauserania Mahunka, 1972–2 sp. Hexanoetus Fain et Philips, 1979–1 sp. Histiostoma Kramer, 1876–236 sp. Cavanoetus Mahunka, 1982 Histiostoma Kramer, 1876 Ichtanoetus Fain et Belpaire, 1985 Hormosianoetus Fain, 1980–1 sp. Hymenanoetus Mahunka, 1963–1 sp. Insulanoetus Sevastyanov, 1973–1 sp. Kanoetus Samšinák, 1966–3 sp. Kaszabanoetus Mahunka, 1976–1 sp. Loxanoetus Fain, 1970–2 sp. Munduytia Oudemans, 1929–1 sp. Nepenthacarus Fashing, 2002–1 sp. Otanoetus Fain et Zumpt, 1974–1 sp. Ovanoetus Fain et van Goethem, 1985–1 sp. Peripatetes Mahunka, 1976–1 sp. Porrhanoetus Mahunka, 1963–1 sp. Probonomoia Fain et Rack, 1987–9 sp. Prowichmannia Radford, 1950–3 sp. Psyllanoetus Fain et Beaucournu, 1973–3 sp. Pteranoetus Mahunka, 1978–4 sp. Rhaphidothrix Mahunka, 1967–3 sp. Rhopalanoetus Scheucher, 1957–11 sp. Richardanoetus Samšinák, 1989–1 sp. Sarraceniopus Fashing et OConnor, 1984–3 sp. Scheucheria Mahunka, 1969–1 sp. Scutanoetus Mahunka, 1969–1 sp. Seliea Oudemans, 1929–1 sp. 14 Acarorum Catalogus IX Semianoetus Mahunka, 1976–2 sp. Stercoranoetus Mahunka et Mahunka-Papp, 1991–2 sp. Synanoetus Mahunka, 1972–1 sp. Syringanoetus Fain, 1980–2 sp. Teinokyra Mahunka, 1973–1 sp. Traskorchestianoetus Fain et Colloff, 1990–2 sp. Xenanoetus Mahunka, 1969–5 sp. Zwickia Oudemans, 1924–3 sp. Subfam. Myianoetinae Scheucher, 1957–48 sp. Myianoetus Oudemans, 1929–48 sp. Subfam. Scolianoetinae Fain, 1974–5 sp. Scolianoetus Fain, 1974–5 sp. Subfam. Spinanoetinae Scheucher, 1957–4 sp Spinanoetus Scheucher, 1957–4 sp Catalogue of Histiostomatidae Family Histiostomatidae Berlese Histiostomatidae Berlese, 1897 Anoetidae Oudemans, 1904: 191. Type: Anoetus Dujardin, 1842 Nodipalpidae Oudemans, 1923: 206 Kaszabanoetinae Mahunka, 1976: 318 Type genus: Histiostoma Kramer, 1876 Subfamily Ceylanoetinae Mahunka Ceylanoetinae Mahunka, 1974c: 507 Type genus: Ceylanoetus Mahunka, 1974 Glyphanoetinae Mahunka, 1972: 371 Type genus: Glyphanoetus Oudemans, 1929 Genus Ceylanoetus Mahunka Ceylanoetus Mahunka, 1974c: 506 Type species: Ceylanoetus excavatus Mahunka, 1974 Ceylanoetus excavatus Mahunka Ceylanoetus excavatus Mahunka, 1974c: 507 Type locality: Central, Mahaweli Ganga, “7 Meilen flussabwärts von Kandy” Distribution: Sri Lanka Acarorum Catalogus IX 15 Genus Chiropteranoetus Womersley Chiropteranoetus Womersley, 1942: 92 Type species: Chiropteranoetus chalinolobus Womersley, 1942 Chiropteranoetus chalinolobus Womersley Chiropteranoetus chalinolobus Womersley, 1942: 92 Type locality: Australia Distribution: Australia Genus Conglanoetus Mahunka Conglanoetus Mahunka, 1978: 143 Comyianoetus Fain & Philips 1979: 147 – Type: Comyianoetus denticulatus Fain et Philips, 1979 Type species: Conglanoetus ornatofoveolatus Mahunka, 1978 Conglanoetus denticulatus (Fain et Philips) Comyianoetus denticulatus Fain & Philips, 1979: 147; Fain et al., 1993: 17 Conglanoetus denticulatus (Fain et Philips): Type locality: Syracuse, N.Y. Distribution: Norway, USA (N.Y.) Host: nest of Bubo bubo and on Trox scaber (Coleoptera: Trocidae) Conglanoetus ornatofoveolatus Mahunka Conglanoetus ornatofoveolatus Mahunka, 1978: 143 Type locality: Oriente 00°24’S, 76°36’W, Limoncocha Distribution: Ecuador Host: Labidus praedator (Hymenoptera: Formicidae, Dorylinae) Conglanoetus puteanus (Fain) Comyianoetus puteanus Fain, in Fain, Vangeluwe, Degreef & Wauthy,1993: 17 Conglanoetus puteanus (Fain): Type locality: Belgium Distribution: Belgium Genus Capronomoia Mahunka Capronomoia Mahunka, 1976b: 321 Type species: Bonomoia sphaerocerae Vitzthum, 1922 Capronomoia asper Ashfaq et Chaudhri Capronomoia asper Ashfaq & Chaudhri, 1988: 355 16 Acarorum Catalogus IX Copronomoia asper Ashfaq & Sarwar, 2001: 129 Type locality: 5 km N of Faisalabad Distribution: Pakistan Capronomoia bellulus Ashfaq et Chaudhri Capronomoia bellulus Ashfaq & Chaudhri, 1988: 352 Copronomoia bellulus: Ashfaq & Sarwar, 2001: 129 Type locality: 3 km W Murree Distribution: Pakistan Capronomoia gradatus Ashfaq et Sarwar Copronomoia gradatus Ashfaq & Sarwar, 2001: 127 Type locality: Quetta Distribution: Pakistan Capronomoia mahunkai Eraky Copronomoia mahunkai Eraky, 1999: 55; Abdelgayed et al., 2019: 32 Type locality: Assiut Distribution: Egypt Capronomoia pango Ashfaq et Chaudhri Capronomoia pango Ashfaq & Chaudhri, 1988: 350 Copronomoia pango Ashfaq & Sarwar, 2001: 129 Type locality: 5 km east of Faisalabad Distribution: Pakistan Capronomoia sphaerocerae (Vitzthum) Bonomoia sphaerocerae Vitzthum, 1922: 545; Scheucher, 1957: 365; Mahunka, 1970: 70 Capronomoia sphaerocerae (Vitzthum): Mahunka, 1976: 321 Type locality: Germany Distribution: Germany, Hungary, Saudi Arabia, Syria Hosts: Sphaerocera subsultans (Diptera: Sphaeroceridae [Syrphidae]); also on Borboridae, Scatophagidae and other Diptera. Capronomoia trossis Ashfaq et Sarwar Capronomoia trossis Ashfaq & Sarwar, 2001: 125 Type locality: Khanewal Distribution: Pakistan Genus Creutzeria Oudemans Creutzeria Oudemans, 1932: 207 Type species: Creutzeria tobaica Oudemans, 1932 Acarorum Catalogus IX 17 Creutzeria seychellensis Nesbitt Creutzeria seychellensis Nesbitt, 1979: 1202 Type locality: Seychelles Distribution: Seychelles Creutzeria tobaica Oudemans Creutzeria tobaica Oudemans, 1932: 207; Buitendijk, 1945: 363 Type locality: Haranggaol Distribution: Indonesia Genus Curculanoetus Fain Curculanoetus Fain, 1974: 66 Type species: Curculanoetus rhynchophorus Fain, 1974 Curculanoetus rhynchophorus Fain Curculanoetus rhynchophorus Fain, 1974: 66 Type locality: Lemba nr. Kinshasa Distribution: DR Congo Host: Rhynchophorus phoenicis (Coleoptera: Curculionidae) Subfamily Fibulanoetinae Fain, Camerik, Lukoschus et Kniest Fibulanoetinae Fain, Camerik, Lukoschus & Kniest, 1980: 39 Type genus: Fibulanoetus Mahunka, 1973 Genus Fibulanoetus Mahunka Fibulanoetus Mahunka, 1973a: 102 Type species: Fibulanoetus labiatus Mahunka, 1973 Fibulanoetus labiatus Mahunka Fibulanoetus labiatus Mahunka, 1973a: 102 Type locality: Meru Distribution: Tanzania Fibulanoetus longitarsus Fain, Camerik, Lukoschus et Kniest Fibulanoetus longitarsus Fain, Camerik, Lukoschus & Kniest, 1980: 42 Type locality: Thamaga Distribution: Botswana Host: Scarabaeus galenus (Coleoptera: Scarabaeidae) Fibulanoetus mahunkai Fain, Camerik, Lukoschus et Kniest Fibulanoetus mahunkai Fain, Camerik, Lukoschus & Kniest, 1980: 41 18 Acarorum Catalogus IX Type locality: Mahalapye Distribution: Botswana Host: Scarabaeus galenus (Coleoptera: Scarabaeidae) Genus Ghanoetus Mahunka Ghanoetus Mahunka, 1973a: 104 Type species: Ghanoetus marmoreus Mahunka, 1973 Ghanoetus marmoreus Mahunka Ghanoetus marmoreus Mahunka, 1973a: 104; 1975: 60 Type locality: Kumasi (Ghana) Distribution: Gambia, Ghana Genus Glyphanoetus Oudemans Glyphanoetus Oudemans, 1929: 450 Type species: Glyphanoetus fulmeki Oudemans, 1929 (= Hypopus phyllotrichus Berlese, 1881) Glyphanoetus africanus Mahunka Glyphanoetus africanus Mahunka, 1967a: 172 Type locality: Kindamba, Meya, Bangu Forest Distribution: Rep. of the Congo (Brazzaville) Glyphanoetus brevisetus Mahunka Glyphanoetus brevisetus Mahunka, 1963a: 38 Type locality: Cazombo (Alto Zambeze) Distribution: Angola Glyphanoetus eboris Fain Glyphanoetus eboris Fain, 1974: 88 Type locality: La Savane de Lamto Distribution: Ivory Coast Glyphanoetus flagellatus Mahunka Glyphanoetus flagellatus Mahunka, 1963d: 370 Type locality: Olkokola, nest of Tachyoryctes sp. Distribution: Tanzania Glyphanoetus incanus Ashfaq, Sarwar et Akbar Glyphanoetus incanus Ashfaq, Sarwar & Akbar, 1999: 21 Type locality: Lahore, Shimla Hill Acarorum Catalogus IX 19 Distribution: Pakistan Glyphanoetus leptos Ashfaq, Sarwar Akbar Glyphanoetus leptos Ashfaq, Sarwar & Akbar, 1999: 23 Type locality: Azad Kashmir, Kotli Distribution: Pakistan Glyphanoetus longus Mahunka Glyphanoetus longus Mahunka, 1969b: 371 Type locality: Mt. Meru, eastern slope Distribution: Tanzania Glyphanoetus mahunkai Eraky Glyphanoetus mahunkai Eraky, 1994: 218 Type locality: Egypt Distribution: Egypt Glyphanoetus micidus Chaudhri Glyphanoetus micidus Chaudhri, 1986: 212 Type locality: Pakistan Distribution: Pakistan Glyphanoetus nomiensis Cross Glyphanoetus nomiensis Cross, 1968: 329 Type locality: Wyoming Distribution: USA (Wyoming, Louisiana) Hosts: Nomia melanderi, N. nortoni (Hymenoptera: Halictidae) Glyphanoetus pappi Mahunka Glyphanoetus pappi Mahunka, 1982: 63 Type locality: Juan Fernandez Islands Distribution: Juan Fernandez Islands Glyphanoetus phyllotrichus (Berlese) Hypopus phyllotrichus Berlese, 1881: AMS Fasc.9, No 2 Anoetus phyllotrichus Oudemans, 1905b: 238 – from “Synchitodes crenatus” - Velp Glyphanoetus fulmeki Oudemans, 1929: 451 Glyphanoetus phyllotrichus (Berlese): Scheucher, 1957: 351; Samšinák, 1984: 62; Mahunka, 1970: 59; Mahunka & Mahunka-Papp: 1991: 286 Type locality: Padua (Italy) 20 Acarorum Catalogus IX Distribution: Austria, Germany, Holland, Hungary, Italy; Cabo Verde Hosts: Ips laricis, I. proximus (Coleoptera: Curculionidae, Scolytinae), Limosina talparum (Diptera: Sphaeroceridae) Glyphanoetus processum Eraky Glyphanoetus processum Eraky, 1994: 217 Type locality: Egypt Distribution: Egypt Glyphanoetus szunyoghyi Mahunka Glyphanoetus szunyoghyi Mahunka, 1961: 528 (as “Gylphanoetus”) Type locality: Arusha Distribution: Tanzania Glyphanoetus venustissimus (Berlese) Anoetus venustissimus Berlese, 1918: 115 Glyphanoetus venustissimus (Berlese): Scheucher, 1957: 354 Type locality: Florence Distribution: Germany, Italy Subfamily Histiostomatinae Scheucher Anoetinae Oudemans, 1904: 191 - Type genus: Anoetus Dujardin, 1842 Histiostomatini Scheucher, 1957: 287; Fain, 1974: 58 Histiostominae Scheucher: Fain, 1974: 63 Histiostomatinae Berlese, 1897; Fain, 1974: 58 Type genus: Histiostoma Kramer, 1876: 105 Genus Ameranoetus Ide et Mahunka Ameranoetus Ide & Mahunka, 1978: 47 Type species: Ameranoetus hematobii Ide et Mahunka, 1978 Ameranoetus hematobii Ide et Mahunka Ameranoetus hematobii Ide & Mahunka, 1978: 48 Type locality: USA Distribution: USA Host: Haematobia irritans (Diptera: Muscidae) Genus Amyzanoetus Fain Amyzanoetus Fain, 1976: 318 Type species: Amyzanoetus halophilus Fain, 1976 Acarorum Catalogus IX 21 Amyzanoetus halophilus Fain Amyzanoetus halophilus Fain, 1976: 318 Type locality: Kerguelen Island Distribution: Kerguelen Island Genus Ancyranoetus Fain et Santiago-Blay Ancyranoetus Fain & Santiago-Blay, 1993: 337 Type species: Ancyranoetus virkkii Fain et Santiago-Blay, 1993 Ancyranoetus virkkii Fain et Santiago-Blay Ancyranoetus virkkii Fain & Santiago-Blay, 1993: 339 Type locality: Puerto Rico Distribution: Puerto Rico Host: Lema nigripes (Coleoptera: Chrysomelidae, Criocerinae) Genus Anoetoglyphus Oudemans ] Anoetoglyphus Oudemans [Vitzthum in litt.], 1927: 75 Type species: Anoetoglyphus ateuchi Oudemans, 1927 Anoetoglyphus africanus Mahunka Anoetoglyphus africanus Mahunka, 1965: 448 Type locality: Lefinie Reservation, Nambouli River Distribution: Congo - Brazaville Anoetoglyphus antennatus Mahunka Anoetoglyphus antennatus Mahunka, 1975: 58 Type locality: Kumasi Distribution: Ghana Anoetoglyphus ateuchi Vitzthum Anoetoglyphus ateuchi Oudemans, 1927: 75 (Vitzthum, in litt.) Type locality: South Italy Distribution: Italy, Sicily Host: Scarabaeus semipunctatus (Coleoptera: Scarabaeidae) Anoetoglyphus bifastigatus Mahunka Anoetoglyphus (?) bifastigatus Mahunka, 1967: 9 Type locality: Dundo Distribution: Angola Host: Rhynchophorus phoenicis (Coleoptera: Curculionidae) 22 Acarorum Catalogus IX Anoetoglyphus copridis Samšinák Anoetoglyphus copridis Samšinák, 1962: 92 Type locality: Neusiedler See Distribution: Austria, Hungary Host: Copris lunaris (Coleoptera: Scarabaeidae, Coprinae) Anoetoglyphus ellipticus Mahunka Anoetoglyphus ellipticus Mahunka, 1973a: 114 Type locality: Nakpanduri Distribution: Ghana Anoetoglyphus globulus Mahunka Anoetoglyphus globulus Mahunka, 1975: 55 Type locality: Kumasi Distribution: Ghana Anoetoglyphus immaturus Mahunka Anoetoglyphus (?) immaturus Mahunka, 1965: 448 Type locality: Lefinie Reservation, Nambouli River Distribution: Republic of Congo - Brazaville Anoetoglyphus langi Mahunka Anoetoglyphus langi Mahunka, 1978: 131 Type locality: Lai Khe Distribution: Vietnam Hosts: Coleoptera (Scarabaeidae) Anoetoglyphus latus Mahunka Anoetoglyphus latus Mahunka, 1978: 56 Type locality: Ghana Distribution: Ghana Anoetoglyphus levis Mahunka Anoetoglyphus levis Mahunka, 1967d: 161 Type locality: Cuc-phuong, Bong Distribution: Vietnam Anoetoglyphus lootsi Mahunka Anoetoglyphus lootsi Mahunka, 1979: 140 Type locality: Transvaal, Nylsvley Res., 35 km NE from Pretoria Acarorum Catalogus IX 23 Distribution: South Africa Anoetoglyphus polygonatus Mahunka Anoetoglyphus polygonatus Mahunka, 1973a: 302 Type locality: Nakpanduri Distribution: Ghana Anoetoglyphus szunyoghyi Mahunka Anoetoglyphus szunyoghyi Mahunka, 1973a: 115; 1975: 59 Type locality: Banda-Kwanta (Ghana) Distribution: Gambia, Ghana Anoetoglyphus tunesiensis Mahunka Anoetoglyphus tunesiensis Mahunka, 1981: 135 Type locality: Tunisia Distribution: Tunisia Genus Anoetus Dujardin Anoetus Dujardin, 1842: 316 Nodipalpus Karpelles, 1893: 128 Type species: Hypopus alicola Dujardin, 1842 Anoetus aelleni Mahunka Anoetus aelleni Mahunka, 1974d: 508 Type locality: Central, Mahaweli Ganga, “7 Meilen flussabwärts von Kandy” Distribution: Sri Lanka Anoetus akodoni Mahunka Anoetus akodoni Mahunka, 1963c: 499 Type locality: El Bolsόn Distribution: Argentina Anoetus alicola (Dujardin) Hypopus alicola Dujardin, 1842: 316 Anoetus discrepans Oudemans, 1903 : 103. Type: Bremen, on Halictus 4-notatus; Oudemans, 1903: 103 Anoetus alicola (Dujardin): Buitendijk, 1945: 361; Scheucher, 1957: 374 Type locality: Hülden an de Aller Distribution: Germany Hosts: Halictus 4-notatus (Hymenoptera: Halictidae) 24 Acarorum Catalogus IX Anoetus angularis Mahunka Anoetus angularis Mahunka, 1974d : 511 Type locality: Uva, Diyaluma Waterfall Distribution: Sri Lanka Anoetus anoetopsis Mahunka Anoetus anoetopsis Mahunka, 1973a : 89 Type locality: Bui Camp Distribution: Ghana Host: Coleoptera Anoetus bradymeri Mahunka Anoetus bradymeri Mahunka, 1972c: 349 Type locality: Tulagi Distribution: Salomon Islands Hosts: Bradymerus leveri, Androsus greensladei (Coleoptera: Tenebrionidae) Anoetus bushlandi Hughes et Jackson Anoetus bushlandi Hughes & Jackson, 1958: 133 Type locality: Kerrville, Texas Distribution: USA (Texas) Hosts: on stable fly Anoetus cacuminatum Mahunka Anoetus cacuminatum Mahunka, 1969: 180 Type locality: Mongolia Distribution: Mongolia Anoetus caljubiensis Gad Hamada Hassan et Sevastianov Anoetus caljubiensis Gad Hamada Hassan & Sevastianov, in Radi & Sevastyanov, 1991: 48 Type locality: Egypt Distribution: Egypt Host: Cataglyphis bicolor (Hymenoptera: Formicidae) Anoetus capillatus Mahunka Anoetus capillatus Mahunka, 1967d: 151 Type locality: Giang Seo Distribution: Vietnam Anoetus castrii Mahunka Anoetus castrii Mahunka, 1972d: 306 Acarorum Catalogus IX 25 Type locality: Maipu, Quebrada, La Plata Distribution: Chile Anoetus ceylanensis Mahunka Anoetus ceylanensis Mahunka, 1974d : 511 Type locality: Central, Mahaweli Ganga, “7 Meilen flussabwärts von Kandy” Distribution: Sri Lanka Anoetus chilensis Mahunka Anoetus chilensis Mahunka, 1972d: 309 Type locality: Curacavi, Los Cerillos Distribution: Chile Anoetus cirrosus Mahunka Anoetus cirrosus Mahunka, 1972d: 308 Type locality: Maipu, Quebrada, La Plata Distribution: Chile Anoetus collis Mahunka Anoetus collis Mahunka, 1965: 445 Type locality: Lefinie Reservation, Nambouli River Distribution: Congo (Brazzaville) Anoetus congoensis Mahunka Anoetus congoensis Mahunka, 1967a: 166 Type locality: Kindamba, Meya, Bangu forest Distribution: Rep. of the Congo (Brazzaville) Anoetus coprophilus Mahunka Anoetus coprophilus Mahunka, 1965: 446 Type locality: Lefinie Reservation, Nambouli River Distribution: Rep. of Congo (Brazzaville) Anoetus debilis Woodring Anoetus debilis Woodring, 1973: 319 Type locality: Turrialba, Cartago Prov. Distribution: Costa Rica Anoetus decorus Mahunka Anoetus decorus Mahunka, 1967d: 152 Type locality: Cuc-phuong, Bong Distribution: Vietnam 26 Acarorum Catalogus IX Anoetus dolifer Mahunka Anoetus dolifer Mahunka, 1977: 94 Type locality: Switzerland, Wallis, Bagnes Distribution: France, Switzerland Hosts: Myrmica ruginodis, M. sabuleti (Hymenoptera: Formicidae - anthills) Anoetus domrowi (Fain) Anoetostoma domrowi Fain, 1968: 246 Type locality: Butemu Village, Finisterre Range, Madang, Central District Distribution: Papua New Guinea Host: Scoliophthalmus sp. near micans (Diptera: Chloropidae) Anoetus dudichi Mahunka Anoetus dudichi Mahunka, 1962: 431 Type locality: Jόsva, Nagyoldal Distribution: Hungary Anoetus endroedyyoungai Mahunka Anoetus endroedyyoungai Mahunka, 1973a : 92 Type locality: Bui Camp Distribution: Ghana Anoetus exornatus Sevastianov Anoetus exornatus Sevastianov, 1973: 849 Type locality: Roshtino Village, Leningradskaya Obl. Distribution: Russia Host: Formica rufa (Hymenoptera: Formicidae) Anoetus fetters Samšinák Anoetus fetters Samšinák, 1989: 97 Type locality: DR Congo Distribution: DR Congo Host: Metaborborus calcaratus (Diptera: Sphaeroceridae) Anoetus fibulatus Mahunka Anoetus fibulatus Mahunka, 1973a : 94 Type locality: Meru Distribution: Tanzania Anoetus ghanensis Mahunka Anoetus ghanensis Mahunka, 1973: 94 Acarorum Catalogus IX 27 Type locality: Ghana Distribution: Ghana Anoetus grapefruit Samšinák Anoetus grapefruit Samšinák, 1989: 95 Type locality: Centre Co. Fillmore Cap Distribution: USA Host: Diptera indet. Anoetus hauseri Mahunka Anoetus hauseri Mahunka, 1972: 1196; 1972e: 951 Type locality: Kephallinia, Aenos Distribution: Greece (Kephallinia) Anoetus halictonida (Woodring) Histiosoma halictonida Woodring, 1973: 312 Anoetus halictonida (Woodring) Type locality: Canada Distribution: Canada Anoetus helena Samšinak Anoetus helena Samšinak, 1989: 95 Type locality: Ecuador Distribution: Ecuador Host: Richardia (plant) Anoetus heliocopridis Vitzthum Anoetus heliocopridis Vitzthum, 1926: 182; Mahunka, 1974a : 141 Type locality: Meru (Tanzania) Distribution: Ghana, Tanzania Anoetus helobius Sevastianov Anoetus helobius Sevastianov, 1973: 851 Type locality: Hadjibeyskiy Liman nr. Odessa Distribution: Ukraine Anoetus horridus Mahunka Anoetus horidus Mahunka, 1972c: 352 Type locality: San Cristobal Distribution: Salomon Islands 28 Acarorum Catalogus IX Hosts: Leiochrodes suturalis, Scyophagus piceus (Coleoptera: Tenebrionidae) Anoetus hospitus Sevastianov Anoetus hospitus Sevastianov, 1973: 851 Type locality: Tchemerovtsi Village, Hmelnitskaya Obl. Distribution: Ukraine Host: anthip of Myrmica ruginodis (Hymenoptera: Formicidae) Anoetus humilis Fain Anoetus humilis Fain, 1974b: 86 Type locality: La Savane de Lamto Distribution: Ivory Coast Anoetus impressus Mahunka Histiostoma impressum Mahunka, 1963a : 31; 1963b: 59 Anoetus impressus (Mahunka): Mahunka, 1967a: 167; 1973a: 96 Type locality: Angola, Lumeje II, N of Santando Station of C.F.B., Poste de Bussaco (80 km E of Luso) Distribution: Angola, Rep. of the Congo (Brazaville), DR Congo, Ghana Anoetus indetonsus Sevastianov Anoetus indetonsus Sevastianov, 1973: 852 Type locality: Odessa area Distribution: Ukraine Host: Coleoptera (Carabidae) Anoetus indicus Oudemans Anoetus indicus Oudemans, 1911: 173; Hughes & Jackson, 1958: 80; Mahunka, 1963a: 32; 1973a : 96 Histiostoma indicus (Oudemans): Buitendijl, 1945; Hughes & Jackson, 1958: 80 Histiostoma indica (Oudemans): Mahunka, 1963b: 60 Type locality: Java, Banjoewangi Distribution: Angola, Ghana, Indonesia (Java) Host: Blattida fam., gen. sp. Anoetus insolita Woodring et Moser Anoetus insolita Woodring & Moser, 1970: 1237 Type locality: Elizabeth, La. Distribution: USA Host: from the boring dust of Dendroctonus frontalis and P. taeda (Coleoptera: Curculionidae) Acarorum Catalogus IX 29 Anoetus irinus Hassan et Sevastianov Anoetus irinus Hassan & Sevastianov, in Radi & Sevastyanov 1991: 45 Type locality: Egypt Distribution: Egypt Host: Iris tingitana (plant) Anoetus kaszabi Mahunka Anoetus kaszabi Mahunka, 1969: 181 Type locality: Mongolia Distribution: Mongolia Anoetus komarpictoris Samšinák Anoetus komarpictoris Samšinák, 1970: 33 Type locality: Kozákov Distribution: Czech Rep. Host: Formica lemani (Hymenoptera: Formicidae) Anoetus longipes (Oudemans) Anoetus longipes Oudemans, 1911: 171; Mahunka, 1967d: 154 Histiostoma longipes (Oudemans): Buitendijk, 1945: 362 Type locality: Banjoewangi Distribution: Indonesia (Java), Vietnam Host: Blattida fam. gen. sp. Anoetus longisetus Mahunka Anoetus longisetus Mahunka, 1967a: 167; 1973a : 96 Type locality: Kindamba, Meya, Louolo river Distribution: Republic of the Congo, Ghana Anoetus longitubus Mahunka Anoetus longitubus Mahunka, 1969b: 365 Type locality: Mt. Meru, eastern slope Distribution: Tanzania Anoetus lunulatus Mahunka Anoetus lunulatus Mahunka, 1971: 1198 Type locality: Kephallinia Distribution: Greece (Kephallinia) Anoetus machadoi Mahunka Anoetus machadoi Mahunka, 1963b: 63; 1967a: 168 30 Acarorum Catalogus IX Type locality: Dundo, Cundueji River (Angola) Distribution: Angola, Rep. of the Congo Anoetus macrosuctus Fain Anoetus macrosuctus Fain, 1974b: 86 Type locality: La Savane de Lamto Distribution: Ivory Coast Anoetus microti Mahunka Anoetus microti Mahunka, 1969: 182 Type locality: Mongolia Distribution: Mongolia Anoetus moshtohorensis Gad Hamada, Hassan et Sevastianov Anoetus moshtohorensis Gad Hamada Hassan & Sevastianov, in Radi & Sevastyanov 1991: 47 Type locality: Egypt Distribution: Egypt Host: Labidura riparia (Dermaptera: Labiduridae) Anoetus myrmicarum (Scheucher) Histiostoma myrmicarum Scheucher, 1957: 300; Mahunka, 1961: 445 Anoetus myrmicarum (Scheucher): Samšinak, 1961: 204; Kurosa, 1976: 62 Type locality: Erlangen, Germany Distribution: China, Czech Rep., Germany, Hungary, Malaysia Hosts: Coptotermes formosanus, Macrotermes malaccensis (Isoptera: Rhinotermitidae, Termitidae); ants (Hymenoptera, Formicidae: Myrmica, Lasius, Camponotus, Formica) Anoetus neglectus Oudemans Anoetus neglectus Oudemans, 1902: 46 Type locality: Bremen Distribution: Germany Host: Nicrophorus humator (Coleoptera: Silphidae) Anoetus notabilis Sellnick Anoetus notabilis Sellnick, 1940: 127 Type locality: Island Distribution: Island Anoetus opertus Sevastianov Anoetus opertus Sevastianov, 1973: 852 Type locality: Tchemerovtsi Village, Hmelnitskaya Obl. Acarorum Catalogus IX 31 Distribution: Ukraine Host: Myrmica rubra (Hymenoptera: Formicidae) Anoetus ornatissimus Mahunka Anoetus ornatissimus Mahunka, 1973a: 96 Type locality: Meru Distribution: Tanzania Anoetus ornatus Mahunka Anoetus ornatus Mahunka, 1967d: 154 Type locality: Tu-li Distribution: Vietnam Anoetus pannonicus Mahunka Anoetus pannonicus Mahunka, 1968: 251 Type locality: Hungary Distribution: Hungary Host: phoretic on Dryocoetes villosus (Coleoptera: Curculionidae) Anoetus parmatus Sevastianov Anoetus parmatus Sevastianov, 1973: 853 Type locality: Tchemerovtsi Village, Hmelnitskaya Obl. Distribution: Ukraine Anoetus pauliani Mahunka Anoetus pauliani Mahunka, 1967a: 168 Type locality: Mount Foari Reservation Distribution: Rep. of the Congo (Brazzaville) Anoetus pini (Scheucher) Histiostoma pini Scheucher, 1957: 317 Anoetus pini (Scheucher): Woodring & Mosaer, 1970: 1255 Type locality: Erlangen Distribution: Belgium, Hungary, Germany Host: phoretic on Hylastes ater and H. cunicularius (Coleoptera: Curculionidae) Anoetus pocsi Mahunka Anoetus pocsi Mahunka, 1967d: 156 Type locality: Giang Seo Distribution: Vietnam 32 Acarorum Catalogus IX Anoetus polynesiacus Vitzthum Anoetus polynesiacus Vitzthum, 1920: 64 Type locality: Fiji Distribution: Fiji Host: Eubactrus semiaeneus (Coleoptera: Brenthidae) Anoetus porosus Mahunka Anoetus porosus Mahunka, 1969: 183 Type locality: Mongolia Distribution: Mongolia Anoetus privorai Samšinák Anoetus přivorai Samšinák, 1962: 89 Type locality: Roccamena, Palermo (Sicily) Distribution: Sicily Host: Aphaenogaster testaceopilosa (Hymenoptera: Formicidae) Anoetus punctulatus Mahunka Anoetus punctulatus Mahunka, 1967a: 169 Type locality: Brazzaville, Orstrom Parc Distribution: Rep. of the Congo (Brazzaville) Anoetus reticulatus Mahunka Anoetus reticulatus Mahunka, 1969b: 366 Type locality: Mt. Meru, eastern slope Distribution: Tanzania Anoetus retractus Sevastianov Anoetus retractus Sevastianov, 1973: 853 Type locality: Jaralgach Island (Black Sea) Distribution: Jaralgach Isl., Crimea Hosts: Bledius bicornis, B. sarmaticus (Coleoptera: Staphylinidae, Oxythelinae) Anoetus samiensis Mahunka Anoetus samiensis Mahunka, 1972: 952 Type locality: Kephallinia: Sami Distribution: Greece (Kephalinia) Anoetus serratus (Mahunka) Histiostoma serrata Mahunka, 1962b: 429 Anoetus serratus (Mahunka), 1970: 26; Sevastianov, 1975: 400 Acarorum Catalogus IX 33 Type locality: Balatonfenyves Distribution: Hungary Host: Demetrias imperialis (Coleoptera: Carabidae) Anoetus shamyi Samšinák Anoetus shamyi Samšinák, 1970: 36 Type locality: Tzasweri Distribution: Russia, Caucasus Host: Formica lemani (Hymenoptera: Formicidae) Anoetus shanensis Mahunka Anoetus shanensis Mahunka, 1973a: 96 Type locality: Borbiri Mt Reserve Distribution: Ghana Anoetus sharae Gad Hamada Hassan et Sevastianov Anoetus sharae Gad Hamada Hassan & Sevastianov, in Radi & Sevastyanov 1991: 49 Type locality: Egypt Distribution: Egypt Anoetus sinuosus Mahunka Anoetus sinuosus Mahunka, 1973a : 299 Type locality: Sese Distribution: Ghana Anoetus solomoniensis Mahunka Anoetus solomoniensis Mahunka, 1972c: 355 Type locality: San Cristobal Distribution: Salomon Islands Host: Leiochrodes suturalis (Coleoptera: Tenebrionidae) Anoetus sordida Woodring et Moser Anoetus sordida Woodring & Moser, 1970: 1238 Type locality: Elizabeth, Louisiana Distribution: Canada, USA (Texas, Georgia, Louisiana) Host: boring dust in galleries of Dendroctonus frontalis, Ips avulsus and I. calligraphus (Col.: Curculionidae, Scolytinae), phoretic on Carcinops sp. (Histeridae) and Corticeus coynei (Tenebrionidae) Anoetus spinipes Mahunka Anoetus spinipes Mahunka, 1973a: 99 34 Acarorum Catalogus IX Type locality: Kumasi Distribution: Ghana Anoetus spinosus Mahunka Anoetus spinosus Mahunka, 1967d: 158 Type locality: Cuc-phuong, Bong Distribution: Vietnam Anoetus striatus Mahunka Histiostoma striatum Mahunka, 1963a: 36 Histiostoma striata Mahunka, 1963b: 62 Anoetus striatus (Mahunka): Mahunka, 1967a: 170 Type locality: Angola, Lumeje II, N of Santando Station of C.F.B., Poste de Bussaco (80 km E of Luso) Distribution: Angola, Rep. of the Congo (Brazzaville) Hosts: Dichaetodrilus incrassatus (Coleoptera: Carabidae), Rhynchophorus phoenicis (Col.: Curculionidae) Anoetus sumatrensis Oudemans Anoetus sumatrensis Oudemans, 1903: 103 Type locality: Sumatra Distribution: Indonesia Anoetus tauffliebi Mahunka Anoetus tauffliebi Mahunka, 1967a: 170 Type locality: Mount Foari Reservation Distribution: Republic of the Congo (Brazzaville) Anoetus topali Mahunka Anoetus ? topali Mahunka, 1963c: 497; 1972 : 311 Type locality: Argentina, Chubut Distribution: Argentina, Chile Anoetus toxici Mahunka Anoetus toxici Mahunka, 1972: 497; 1972 : 357 Type locality: Guadalcanal Distribution: Salomon Islands Host: Toxicum quadricorne rufipes (Coleoptera: Tenebrionidae) Anoetus turcastanae Oudemans Anoetus turcastanae Oudemans, 1917: 392 Acarorum Catalogus IX 35 Anoetus turcestanae Oudemans: Buitendijk, 1945: 361 Type locality: “Turkestan” Distribution: “Turkestan” Host: Homalocopris tmolus (Coleoptera: Scarabaeidae) Anoetus urcerosus Sevastianov Anoetus urcerosus Sevastianov, 1973: 855 Type locality: Tchemerovtsi Village, Hmelnitskaya Obl. Distribution: Ukraine Host: anthip of Formica rufa (Hymenoptera: Formicidae) Anoetus variolatus Mahunka Anoetus variolatus Mahunka, 1973a: 100 Type locality: Borbiri Mt. Res. Distribution: Ghana Anoetus vilhenaorum Mahunka Anoetus vilhenaorum Mahunka, 1963b: 65 Type locality: Dundo, Cundueji River Distribution: Angola Anoetus virginiana Woodring et Moser Anoetus virginiana Woodring & Moser, 1970: 1237; Kurosa, 1976: 62 Type locality: Virginia (USA) Distribution: Malaysia, USA Anoetus vitzthumi (Scheucher) Histiostoma vitzthumi Scheucher, 1957: 313 Anoetus vitzthumi (Scheucher): Woodring & Moser, 1970: 1256; Sevastianov, 1975: 400 Type locality: Schleswig-Holstein Distribution: Austria, Germany Host: Dendroctonus micans (Coleoptera: Curculionidae, Scolitinae) Anoetus wastis Gad Hamada Hassan et Sevastianov Anoetus wastis Gad Hamada Hassan & Sevastianov, in Radi & Sevastyanov 1991: 49 Type locality: Egypt Distribution: Egypt Host: Labidura riparia (Dermaptera) Genus Aphodanoetus Bongers, OConnor et Lukoschus Aphodanoetus Bongers, OConnor & Lukoschus, 1985: 33 36 Acarorum Catalogus IX Type species: Aphodanoetus teinophallus Bongers, OConnor et Lukoschus, 1985 Aphodanoetus teinophallus Bongers, OConnor et Lukoschus Aphodanoetus teinophallus Bongers, OConnor & Lukoschus, 1985: 36 Type locality: Nijmegen Distribution: Holland Host: Aphodius frater (Coleoptera: Scarabaeidae, Aphodiinae) Genus Auricanoetus Fain et Zumpt Auricanoetus Fain & Zumpt, 1974: 99 [as subgenus of Loxanoetus) Auricanoetus Fain & Zumpt: Fain 1983: 849 [as genus] Type species: Loxanoetus (Auricanoetus) longitarsus Fain et Zumpt, 1974 Auricanoetus longitarsus (Fain et Zumpt) Loxanoetus (Auricanoetus) longitarsus Fain et Zumpt, 1974: 99; Domrow & Ladds, 1984: 759 Auricanoetus longitarsus (Fain et Zumpt): Fain 1983: 849 Type locality: Karumu Game Reserve Distribution: Uganda Host: Syncerus caffer (Artiodactyla: Bovidae) Genus Austranoetus Fain Austranoetus Fain, 1976: 314 Type species: Austranoetus kerguelenensis Fain, 1976 Austranoetus kerguelenensis Fain Austranoetus kerguelenensis Fain, 1976: 314 Type locality: Kerguelen Distribution: Kerguelen Genus Bonomoia Oudemans Bonomoia Oudemans, 1911: 174 Type species: Bonomoia primitiva Oudemans, 1911 Bonomoia africana Mahunka Bonomoia africana Mahunka, 1967a: 171 Type locality: Mount Foari Reservation Distribution: Republic of the Congo Bonomoia certa Woodring et Moser Bonomoia certa Woodring & Moser, 1970: 1252 Type locality: Moskow Mtn., Moskow, Idaho Acarorum Catalogus IX 37 Distribution: USA Hosts: Dendroctonus brevicomis, D. pseudotsugae, Scolytus ventralis (Coleoptera: Curculionidae, Scolytinae) Bonomoia picturata Sevastianov Bonomoia picturata Sevastianov, 1973: 81 Type locality: Khmelnitskij Rayon Distribution: Ukraine Bonomoia primitiva Oudemans Bonomoia primitiva Oudemans, 1911: 174 Type locality: Banjoewangi, Java Distribution: Indonesia (Java) Host: Blattida fam., gen., sp. Bonomoia probata Sevastianov Bonomoia probata Sevastianov, 1973: 82 Type locality: Khmelnitskij Rayon Distribution: Ukraine Bonomoia recondita Sevastianov Bonomoia recondita Sevastianov, 1973: 83 Type locality: Mariyskaya ASSR, Orshanskiy Rayon Distribution: Russia Host: Myrmica ruginodis (Hymenoptera: Formicidae) Bonomoia reticulata Mahunka Bonomoia reticulata Mahunka, 1967d: 162 Type locality: Cuc-puong, Bong Distribution: Vietnam Bonomoia rugosa Mahunka Bonomoia rugosa Mahunka, 1973a: 301 Type locality: Kwandaso Distribution: Ghana Bonomoia zoheirii Metwali et Ahmad Bonomoia zoheirii Metwali & Ahmad, 1987: 85 Type locality: Sharkya Governorate Distribution: Egypt Host: ants (Formicidae) 38 Acarorum Catalogus IX Bonomoia sp. Hosts: Hybomitra affinis, H. epistates, H. illota, H. lasiophthalmus, H. nuda, Tabanus quinquestriatus (Canada)(Mullen et al., 1979) Genus Bothyanoetus Fain et Camerik Bothyanoetus Fain & Camerik, 1978: 170 Type species: Bothyanoetus pereirai Fain et Camerik, 1978 Bothyanoetus pereirai Fain et Camerik Bothyanoetus pereirai Fain & Camerik, 1978: 170 Type locality: Cotia, Sao Paulo Distribution: Brazil Host: Bothynus deiphobus (Coleoptera: Scarabaeidae) Genus Cederhjelmia Oudemans Cederhjelmia Oudemans, 1931: 261 Type species: Cederhjelmia quadriuncinata Oudemans, 1931 Cederhjelmia quadriuncinata Oudemans Cederhjelmia quadriuncinata Oudemans, 1931: 179 Type locality: South Brazil Distribution: Brazil Genus Chiloanoetus Fain Chiloanoetus Fain, 1974: 63 Type species: Chiloanoetus mayumbensis Fain, 1974 Chiloanoetus mayumbensis Fain Chiloanoetus mayumbensis Fain, 1974: 65 Type locality: Chiloango, Mayumbe Distribution: DR Congo Host: Prioscelis serrata (Coleoptera: Tenebrionidae) Genus Hauserania Mahunka Hauserania Mahunka, 1972e: 954 Type species: Hauserania ornata Mahunka, 1972 Hauserania graeca Mahunka Hauserania graeca Mahunka, 1972e: 956 Type locality: Kephalinia, Sami Distribution: Greece (Kephalinia) Acarorum Catalogus IX 39 Hauserania ornata Mahunka Hauserania ornata Mahunka, 1972e: 954 Type locality: Kephalinia, on the road Livadion - Argostolion Distribution: Greece (Kephalinia) Genus Hexanoetus Fain et Philips Hexanoetus Fain & Philips, 1979: 150 Type species: Hexanoetus conoidalis Fain et Philips, 1979 Hexanoetus conoidalis Fain et Philips Hexanoetus conoidalis Fain & Philips, 1979: 150 Type locality: Syracuse, N.Y. Distribution: USA (N.Y.) Genus Histiostoma Kramer Histiostoma Kramer, 1876: 105 Zschachia Oudemans, 1929: 451. Type: Zschachia laevia Oudemans, 1929 Type species: Hypopus julorum C.L. Koch, 1843 (= Histiostoma pectineum Kramer, 1876) Subgenus Histiostoma Kramer Histiostoma Kramer, 1876: 105 Type species: Histiostoma pectineum Kramer, 1876 (= Hypopus feroniarum Dufour, 1839) Histiostoma (H.) abietis Scheucher Histiostoma abietis Scheucher, 1957: 335 Anoetus abietis (Scheucher): Woodring & Moser, 1970: 1255 Type locality: Regensburg Distribution: Germany Host: Ips curvidens (Coleoptera: Curculionidae, Scolytinae) Histiostoma (H.) acrocini Mahunka Histiostoma acrocini Mahunka, 1982: 83 Type locality: Brazil Distribution: Brazil Host: phoretic on Acrocinus longimanus (Coleoptera: Cerambycidae) Histiostoma (H.) adarcuale Mahunka Histiostoma adarcuale Mahunka, 1979: 334 Type locality: Puerto Viejo Distribution: Costa Rica 40 Acarorum Catalogus IX Histiostoma (H.) adsimile Fain Histiostoma adsimile Fain, 1974b: 85 Type locality: La Savane de Lamto Distribution: Ivory Coast Histiostoma (H.) alphaticum Fain Histiostoma alphaticum Fain, 1974b: 82 Type locality: La Savane de Lamto Distribution: Ivory Coast Histiostoma (H.) alii Eraky Histiostoma alii Eraky, 2000: 8 Type locality: Sohag Distribution: Egypt Histiostoma (H.) amargynus Mahunka Histiostoma amargynus Mahunka, 1976b: 309 Type locality: Nondugl, 2200–2700 m Distribution: Papua New Guinea Host: Amarygmus [as “Amargynus”] cfr. capillatus (Coleoptera: Tenebrionidae) Histiostoma (H.) americanum Banks Histiostoma americanum Banks, 1906: 11 Type locality: USA Distribution: USA Histiostoma (H.) amieti Mahunka Histiostoma amieti Mahunka, 1979: 236 Type locality: Cameroon Distribution: Cameroon Histiostoma (H.) andrei Mahunka Histiostoma andrei Mahunka, 1963b: 57 Type locality: Nordeste: forest n. Cassai Distribution: Angola Histiostoma (H.) angolensis Mahunka Histiostoma angolensis Mahunka, 1963a: 27; 1963b: 58 Type locality: Cavuemba River, Alto Cuílo, Poste de Cacolo Distribution: Angola Acarorum Catalogus IX 41 Histiostoma (H.) antillense Mahunka Histiostoma antillense Mahunka, 1981: 350 Type locality: Castries, Vigie Distribution: St. Lucia, Windward Islands Histiostoma arcuata Mahunka Histiostoma arcuata Mahunka, 1963a: 28 Histiostoma arcuatus Mahunka: Type locality: Lumeje II, N of Santando Station of C.F.B., Poste de Bussaco (80 km E of Luso) Distribution: Angola, Egypt Histiostoma (H.) arenatus Negm Histiostoma arenatus Negm, 2007; Abdelgayed et al., 2019: 32 Type locality: Egypt Distribution: Egypt Histiostoma (H.) australiense Mahunka Histiostoma australiense Mahunka, 1975: 317; Tagami & Halliday, 2013: 218 Type locality: Australia Distribution: Australia Host: Titanolabis colossea (Dermaptera: Anisolabididae) Histiostoma (H.) bacilligerum Mahunka Histiostoma bacilligerum Mahunka, 1963a: 30 Type locality: Cazombo (Alto Zambeze) Distribution: Angola Histiostoma (H.) bakeri Hughes et Jackson Histiostoma bakeri Hughes & Jackson, 1958: 45 Type locality: Norris Dam, Tenn. Distribution: USA (Tenn.) Histiostoma (H.) baloghi Mahunka Histiostoma baloghi Mahunka, 1962: 517 Type locality: Bükk – Gebirge, Belko Distribution: Hungary Histiostoma (H.) banjuwangicus (Oudemans) Anoetus banjuwangicus Oudemans, 1911: 172 42 Acarorum Catalogus IX Histiostoma banjuwangicus (Oudemans): Buitendijk, 1945: 361; Hughes & Jackson, 1958: 49 Type locality: Banjoewangi Distribution: Indonesia (Java) Histiostoma (H.) bananai Eraky et Shoker Histiostoma bananai Eraky & Shoker, 1993: 53 Type locality: Egypt Distribution: Egypt Histiostoma (H.) berghii Jensen Histiostoma berghii Jensen, 1896: 72 Histiostoma berghi (Jensen): Buitendijk, 1945: 361 Type locality: Copenhagen Distribution: Denmark Host: in cocoons of Aulastoma gulo (Hirudinea) Histiostoma (H.) berndhauseri Mahunka Histiostoma berndhauseri Mahunka, 1992: 396 Type locality: Cascade de Faarumai Distribution: Tahiti Host: Diplopoda Histiostoma (H.) bernoni Mahunka Histiostoma bernoni Mahunka, 1979: 134 Type locality: Kenya Distribution: Kenya Host: Coleoptera (Scarabaeidae) Histiostoma (H.) bifoveolatum Mahunka Histiostoma bifoveolatum Mahunka, 1979: 135 Type locality: Poroto Mt. Distribution: Tanzania Histiostoma (H.) blomquisti Wirth et Moser Histiostoma blomquisti Wirth & Moser, 2010: 357 Type locality: Pineille and New Orleans Distribution: USA Host: Solenopsis invicta (Hymenoptera: Formicidae) Histiostoma (H.) brevimanum (Oudemans) Lipstorpia brevimana Oudemans, 1914: 73 Acarorum Catalogus IX 43 Anoetus brevimanus: Oudemans, 1917: 392 Histiostoma brevimanus (Oudemans): Buitendijk, 1945: 361; Hughes & Jackson, 1958: 52; Bongers, OConnor & Lukoschus, 1985: 6 Type locality: Sittard Distribution: Holland Histiostoma (H.) brevipodum Berlese Histiostoma brevipodum Berlese, 1910: 270 Type locality: Palermo Distribution: Sicily Habitat: “inter folia putria” Histiostoma (H.) brevipes Banks Histiostoma brevipes Banks, 1906: 11 Type locality: USA Distribution: USA Histiostoma (H.) calicifera Mahunka Histiostoma calicifera Mahunka, 1974b: 375; Eraky 2000: 15 Type locality: Banda-Nkwanta Distribution: Ghana Histiostoma (H.) camerunensis Mahunka Histiostoma camerunensis Mahunka, 1979: 238 Type locality: Cameroon Distribution: Cameroon Histiostoma (H.) campanula (Oudemans) Anoetus campanula Oudemans, 1914: 71 Histiostoma campanulae (Oudemans): Buitendijk, 1945: 361; Hughes & Jackson, 1958: 54 Type locality: Banjoewangi Distribution: Indonesia (Java) Histiostoma (H.) camphori Eraky Histiostoma camphori Eraky, 1999: 72; Abdelgayed et al., 2019: 32 Type locality: Upper Egypt (Assiut) Distribution: Egypt Histiostoma (H.) capillatum Mahunka Histiostoma capillata Mahunka, 1978: 143 Histiostoma capillatum Mahunka: Klimov & Khaustov, 2018 Type locality: Oriente, 00o24S, 76o36’W, Limoncocha 44 Acarorum Catalogus IX Distribution: Ecuador Host: Labidus praedator (Hymenoptera: Formicidae, Dorylinae) Histiostoma (H.) caro Ashfaq, Chaudhri et Aheer Histiostoma caro Ashfaq, Chaudhri & Aheer, 1985: 42 Type locality: Pakistan Distribution: Pakistan Histiostoma (H.) cataglyphi Yousef et Metwally Histiostoma cataglyphi Yousef & Metwally, 1974: 724 Type locality: Zagazig. Sharkia governorate Distribution: Egypt Histiostoma (H.) chilopophila Mahunka Histiostoma chilopophila Mahunka, 1979: 241 Type locality: Cameroon Distribution: Cameroon Histiostoma (H.) cirratum (Oudemans) Anoetus cirratus Oudemans, 1911: 173 Histiostoma cirratus (Oudemans): Buitendijk, 1945: 361 Type locality: Banjoewangi Distribution: Indonesia (Java) Host: Blattodea fam., gen. sp. Histiostoma (H.) clavatum Mahunka Histiostoma clavatum Mahunka, 1963b: 58 Type locality: Dundo Distribution: Angola Host: Brachytripes membranaceus (Orthoptera: Grylloidea) Histiostoma (H.) clavifer Mahunka Histiostoma clavifer Mahunka, 1978a: 146 Type locality: 4 km W of Villa Neily, 8o40’N, 82o56’W Distribution: Costa Rica Host: Eciton hamatum (Hymenoptera: Formicidae, Dorylinae) Histiostoma (H.) conclavicola (Oudemans) Anoetus conclavicola Oudemans, 1929: 461 Histiostoma (H.) conclavicola (Oud.) Type locality: Stanleyville (Kisangani) Acarorum Catalogus IX 45 Distribution: DR Congo Histiostoma (H.) conjunctum (Woodring et Moser) Anoetus conjuncta Woodring & Moser, 1970: 1250 Histiostoma conjuncta (Woodring et Moser): Woodring & Moser, 1975: 83 Histiostoma conjunctum (Woodring et Moser): Klimov & Khaustov, 2018: 2412 Type locality: Tegusigalpa, Honduras Distribution:Guatemala, Honduras, USA Hosts: galleries of Dendroctonus frontalis, Ips avulsus, I. calligraphus, I. cribricollis, I. grandicollis Histiostoma (H.) continuatum Mahunka Histiostoma continuata Mahunka, 1978a: 146 Type locality: Oriente, 00o24S, 76o36’W, Limoncocha Distribution: Ecuador Host: Labidus sp. (Hymenoptera: Formicidae, Dorylinae) Histiostoma (H.) coprae Chinniah et Mohanasundaram Histiostoma coprae Chinniah & Mohanasundaram, 1995: 73 Type locality: Tamil Nadu Distribution: India Host: Copris (Coleoptera) Histiostoma (H.) coronatum Mahunka Histiostoma coronatum Mahunka, 1979: 335 Type locality: Puerto Viejo Distribution: Costa Rica Histiostoma (H.) cossi Scheucher Histiostoma cossi Scheucher, 1957: 329 Type locality: Erlangen Distribution: Germany Hosts: Cossus cossus, C. ligniperda (Lepidoptera: Cossidae) Histiostoma (H.) crantor Ashfaq, Chaudhri et Aheer Histiostoma crantor Ashfaq, Chaudhri & Aheer, 1985: 22 Type locality: Pakistan Distribution: Pakistan Histiostoma (H.) crassipes (Oudemans) Anoetus crassipes Oudemans, 1912: 234; ? Vitzthum, 1923 46 Acarorum Catalogus IX Histiostoma crassipes (Oudemans): Klimov & Khaustov, 2018: 2413 Type locality: Tanzania Distribution: Tanzania, (?) Nepal Hosts: Platypus dispar, Ips stebbingi (Coleoptera: Curculionidae), Hololepta africanae (Col.: Histeridae) Histiostoma (H.) crypturgi Scheucher Histiostoma crypturgi Scheucher , 1957: 310; Klimov & Khaustov, 2018: 2413 Anoetus crypturgi (Scheucher): Woodring & Moser, 1970: 1255; Sevastianov, 1975: 400 Type locality: Erlangen and Regensburg Distribution: Germany, Poland, Slovakia Hosts: Crypturgus pusillus (Coleoptera: Curculionidae, Scolytinae), Rhagium sp. (Cerambycidae), Aphodius prodromus (Scarabaeidae) Histiostoma (H.) cubanum Mahunka Histiostoma cubanum Mahunka, 1978: 111 Type locality: La Habana Distribution: Cuba Host: Acanthoderes circumflexa (Coleoptera: Cerambycidae) Histiostoma (H.) cyrtandrae (Vitzthum) Anoetus cyrtandrae Vitzthum, 1931: 110 Histiostoma cyrtandrae (Vitzthum) Type locality: at Ngebel Lake, Middle Java Distribution: Indonesia Histiostoma (H.) dactyloscopicum Mahunka et Eraky Histiostoma dactyloscopicum Mahunka & Eraky, 1987: 130 Type locality: Hungary Distribution: Hungary Histiostoma (H.) darwishii Eraky Histiostoma darwishii Eraky, 1994: 220; Abdelgayed et al., 2019: 32 Type locality: Egypt Distribution: Egypt Histiostoma (H.) decemvirgae Jacot Histiostoma decemvirgae Jacot, 1939: 199 Type locality: Western North Carolina Distribution: USA Acarorum Catalogus IX 47 Histiostoma (H.) densus Ashfaq, Chaudhri et Aheer Histiostoma densus Ashfaq, Chaudhri & Aheer, 1985 18 Type locality: Pakistan Distribution: Pakistan Histiostoma (H.) diadematum Mahunka Histiostoma diademata Mahunka, 1978a: 148 Type locality: Oriente, 00o24S, 76o36’W, Limoncocha Distribution: Ecuador Host: Neivamyrmex gibbatus (Hymenoptera: Formicidae, Dorylinae) Histiostoma (H.) disiunctum Mahunka Histiostoma disiuncta Mahunka, 1962b: 428 Anoetus disiunctus Type locality: Balatonfenyves Distribution: Hungary Host: Demetrias imperialis (Coleoptera: Carabidae) Histiostoma (H.) distentosetum Mahunka Histiostoma distentoseta Mahunka, 1974b: 375 Type locality: Banda-Nkwanta Distribution: Ghana Histiostoma (H.) dryocoeti Scheucher Histiostoma dryocoeti Scheucher, 1957: 315 Anoetus dryocoeti (Scheucher): Woodring & Moser, 1970: 1255; Sevastianov, 1975: 397 Type locality: “Erlanger Umgebung und Oberpfalz” Distribution: Germany Hosts: Dryocoetus autographus (Coleoptera: Curculionidae, Scolytinae), Dorcus parallelopipedus (Coleoptera: Lucanidae) Histiostoma (H.) dugesi (Claparède) Hypopus dugesii Claparède, 1868: 506 Histiostoma dugesi (Clap.): Buitendijk, 1945: 361 Type locality: Giseh Distribution: Egypt Histiostoma (H.) duplex Mahunka Histiostoma duplex Mahunka, 1979: 337 Type locality: Puerto Viejo Distribution: Costa Rica 48 Acarorum Catalogus IX Histiostoma (H.) edax Ashfaq, Chaudhri et Aheer Histiostoma edax Ashfaq, Chaudhri & Aheer, 1985: 33 Type locality: Pakistan Distribution: Pakistan Histiostoma (H.) editum Mahunka Histiostoma edita Mahunka, 1974b: 378 Type locality: Kwandaso Distribution: Ghana Histiostoma (H.) egyptiacum Eraky et Shoker Histiostoma egyptiacum Eraky & Shoker, 1993: 51 Type locality: Egypt Distribution: Egypt Histiostoma (H.) eickworti Woodring Histiostoma eickworti Woodring, 1973: 316 Type locality: Costa Rica Distribution: Costa Rica Host: Augochlora spp. (Hymenoptera: Halictidae) Histiostoma (H.) elzingai Mahunka Histiostoma elzingai Mahunka, 1978a: 148 Type locality: Oriente, 00o24S, 76o36’W, Limoncocha Distribution: Ecuador Host: Eciton lucanoides (Hymenoptera: Formicidae, Dorylinae) Histiostoma (H.) endroedii (Mahunka) Anoetus endroedii Mahunka, 1973a: 90 Histiostoma endroedii: Mahunka, 1979: 128 Type locality: Abofour (Ghana) Distribution: Ghana, Gambia Host: Coleoptera Histiostoma (H.) ensifer (Oudemans) Anoetus ensifer Oudemans, 1912: 236 Histiostoma ensifer (Oudemans): Buitendijk, 1945: 361 Type locality: Tanganyika, Amani Distribution: Tanzania Histiostoma (H.) erugo Ashfaq, Chaudhri et Aheer Histiostoma erugo Ashfaq, Chaudhri & Aheer, 1985: 27 Acarorum Catalogus IX 49 Type locality: Pakistan Distribution: Pakistan Histiostoma (H.) farghali Eraky Histiostoma farghali Eraky, 2000: 13 Type locality: Egypt Distribution: Egypt Histiostoma (H.) feroniarum (Dufour) Hypopus feroniarum Dufour, 1839: 278 Histiostoma pectineum Kramer, 1876 Acarus mamillaris Canestrini, 1878: 140 Tyroglyphus rostro-serratus Mégnin, 1873: 372 Rhyzoglyphus rostro-serratus: Histiostoma rostro-serratum: Michael, 1901: Anoetus feroniarum (Dufour): Mahunka, 1971: 1195; 1972e: 951 Histiostoma feroniarum (Dufour): Scheucher, 1957: 290; Costa, 1967: 107; Caceres & Fain, 1975: 146; Chen et al., 1982; Shen et al. 1993; Qin & Rohitha 1996: 65; Fain, 1991: 79; Rebollero & Arroyo, 1996; Sanchez-Covisa et al., 1999 Type locality: France Distribution: Australia, Austria, China, France, Germany, Greece (Kephallinia, Zakynthos), Israel, Italy, New Zealand, Peru, Spain, Canary Is. (Tenerife) Hosts: Cormocephalus aurantiipes (Chilopoda: Scolopendridae), Hister merdarius, H. stercorarius, H. unicolor (Coleoptera: Histeridae); Spalax ehrenbergi (Mammalia: Rodentia) Histiostoma (H.) flabelliforme Leonardi Histiostoma flabelliforme Leonardi, 1899: 922 Type locality: Portici Distribution: Italy Histiostoma (H.) flaccus Sarwar Histiostoma flaccus Sarwar, 1998: 2 Type locality: Pakistan Distribution: Pakistan Histiostoma (H.) flagellifer Leonardi Histiostoma flagellifer Leonardi, 1897 Type locality: Italy Distribution: Italy 50 Acarorum Catalogus IX Histiostoma (H.) formosanum Philipsen et Coppel Histiostoma formosana Philipsen & Coppel, 1977: 497; Wang et al., 2002 Type locality: Taiwan Distribution: Taiwan, China Host: Coptotermes formosanus (Isoptera: Rhinotermitidae) Histiostoma (H.) formicarum Vitzthum Histiostoma formicarum Vitzthum, 1941: 315 Type locality: Tirol Distribution: Austria Host: Myrmica sulcinodis (Hymenoptera: Formicidae) Histiostoma (H.) fortis Ashfaq, Sarwar et Amiad Histiostoma fortis Ashfaq, Sarwar & Amiad, 2000: 37 Type locality: Rawalakot Distribution: Pakistan Host: from decomposing vegetable matter Histiostoma (H.) foveosimile Mahunka Histiostoma foveosimile Mahunka, 1975: 320 Type locality: Kiunga Distribution: Papua New Guinea Histiostoma (H.) fragosus Ashfaq, Chaudhri et Aheer Histiostoma fragosus Ashfaq, Chaudhri & Aheer, 1985: 30 Type locality: Pakistan Distribution: Pakistan Histiostoma (H.) fucatifer Mahunka Histiostoma fucatifer Mahunka, 1975b: 47 Type locality: Gambia Distribution: Gambia Host: Ploceus-nest Histiostoma (H.) fuliginosi Scheucher Histiostoma fuliginosi Scheucher, 1957: 336 Type locality: Erlangen Distribution: Germany Histiostoma (H.) furcata Mahunka Histiostoma furcata Mahunka, 1975b: 50 Type locality: Ghana Acarorum Catalogus IX 51 Distribution: Ghana Histiostoma (H.) galerus Sarwar, Ashfaq et Islam, Histiostoma galerus Sarwar, Ashfaq & Islam, 1998: 4 Type locality: Pakistan Distribution: Pakistan Histiostoma (H.) gambiense Mahunka Histiostoma gambiense Mahunka, 1975b: 53 Type locality: Gambia Distribution: Gambia Histiostoma (H.) gampsoceri Mahunka Histiostoma gampsoceri Mahunka, 1976b: 308 Type locality: “Formosa” (Taiwan) Distribution: Taiwan Host: Gampsocera mutata (Diptera: Chloropidae) Histiostoma (H.) genavensium Mahunka Histiostoma genavensium Mahunka, 1992: 396 Type locality: Viti Levu: entre Yanuca et Nadi Distribution: Fiji Host: Diplopoda Histiostoma (H.) gervaisi Oudemans Histiostoma gervaisi Oudemans, 1937: 2145 – nomen novum pro Hypopus ovalis Gervais, 1844 Type locality: Paris Distribution: France Host: Lithobius forficatus (Chilopoda: Lithobiidae) Histiostoma (H.) gladiger (Vitzthum) Anoetus gladiger Vitzthum, 1926a: 489; Woodring & Moser, 1970: 1255 Histiostoma gladiger Scheucher, 1957: 334; Cooreman, 1963: 39 Type locality: “Nikolajewsk an der Mundung des Amur” Distribution: Austria, Belgium, Bulgaria, France (incl. Corsica), Germany, Russia (Far East) Hosts: Ips cembrae, I. sexdentatus, I. subelongatus (Coleoptera: Curculionidae, Scolytinae) Histiostoma (H.) globotuberum Takami et Kuwahara Histiostoma globotuberum Takami & Kuwahara, 2005: 86 Type locality: Kyoto and Tsukuba Distribution: Japan 52 Acarorum Catalogus IX Histiostoma (H.) gordius (Vitzthum) Anoetus phyllotrichus Vitzthum, 1921a: Anoetus gordius Vitzthum, 1923a: 164; Mahunka, 1970: 27; Woodring & Moser, 1970: 1255; Sevastianov, 1975: 400 Anoetus julorum Vitzthum, 1926: Histiostoma gordius (Vitzthum): Scheucher, 1957: 311; Cooreman, 1963: 37; Hunter & Davis, 1963: 5; Distribution: Austria, Belgium, Bosnia & Herzegovina, Germany, Hungary, Romania Hosts: Ips laricis (Ipidae), Orthomicus [Ips] laricis, O. proximus (Coleoptera: Curculionidae, Scolytinae) Histiostoma (H.) gorgo Mahunka Histiostoma gorgo Mahunka, 1974b: 380 Type locality: Banda-Nkwanta Distribution: Ghana Histiostoma (H.) gracilipes Banks Histiostoma gracilipes Banks, 1906: 10 Type locality: USA Distribution: USA Histiostoma (H.) gracilis Ashfaq, Sarwar et Amiad Histiostoma gracilis Ashfaq, Sarwar & Amiad, 2000: 34 Type locality: Haripur Distribution: Pakistan Host: from gram (Cicer arietinum) Histiostoma (H.) granulatum Vitzthum Histiostoma granulatum Vitzthum, 1935: 93 Type locality: Hivaoa: Kaava Mt. Distribution: Marquesan Islands Host: Brachypeplus sp. (Coleoptera: Nitidulidae) Histiostoma (H.) guanophilum Mahunka Histiostoma guanophilum Mahunka, 1982: 293 Type locality: New Mexico Distribution: USA Histiostoma (H.) halicticolum Fain et Erteld Histiostoma halicticola Fain & Erteld, 1998: 48 Histiostoma halicticolum Fain & Erteld: Type locality: Berlin, Doberitzer Heide natural reserve Acarorum Catalogus IX 53 Distribution: Germany Host: Halictus sexcinctus (Hymenoptera: Halictidae) Histiostoma (H.) halictonida Woodring Histiostoma halictonida Woodring, 1973: 312; Engel & Fain, 2003: 649 Anoetus halictonida: Achtymichuk, 2017 Type locality: USA Distribution: Canada, Costa Rica, Panama, USA Hosts: Halictus rubicundus, Megalopta spp. (Hymenoptera: Halictidae) Histiostoma (H.) heinemanni Hill et Deahl Histiostoma heinemanni Hill & Deahl, 1978: 318 Type locality: Pennsylvania Distribution: USA Histiostoma (H.) herbali Eraky et al. Anoetus herbali Eraky et al., 2017: 183 Type locality: Assiut, Abu Teeg Distribution: Egypt Host: from Citrus orchards Histiostoma (H.) heriades Fain et Pauly Histiostoma heriades Fain & Pauly, 2001: 136 Type locality: Sainte Marie Island, Kalalao Forest Distribution: Madagascar Host: Heriades erythrosoma (Hymenoptera: Megachilidae) Histiostoma (H.) himalayae (Vitzthum) Anoetus himalayae Vitzthum, 1923: 162 Histiostoma himalayae (Vitzthum): Fain, Athias-Binche & Carton, 1993: 73; Type locality: Himalaya Distribution: Himalaya Host: Polygraphus minor (Coleoptera: Curculionidae, Scolytinae [Ipidae]) Histiostoma (H.) histeri Mahunka Histiostoma histeri Mahunka, 1976: 97 Type locality: Bükk-Gebirge, Czalajka völgy Distribution: Hungary, Romania Host: Platysoma compressum (Coleoptera: Histeridae) Histiostoma (H.) horridum Berlese Histiostoma horridum Berlese, 1913: 104 54 Acarorum Catalogus IX Type locality: Java Distribution: Indonesia (Java) Histiostoma (H.) huettleri Mahunka Histiostoma huettleri Mahunka, 1975: 45 Type locality: Gambia Distribution: Gambia Histiostoma (H.) humiditatis (Vitzthum) Anoetus humiditatis Vitzthum, 1927: 98 Histiostoma humiditatis (Vitzthum): Womersley, 1942; Hughes & Jackson, 1958: 78; Tseng, 1984 Type locality: Thaya, Lower Austria Distribution: Australia, Austria, Taiwan Host: Titanolabis colossea (Dermaptera: Anisolabididae) Histiostoma (H.) hungaricum Mahunka Histiostoma hungarica Mahunka, 1976: 100 Histiostoma hungaricum Mahunka: Klimov & Khaustov, 2018 Type locality: Slofok Distribution: Hungary Host: Carcinopus 14-striata (Coleoptera: Histeridae) Histiostoma (H.) imbecillus Ashfaq, Chaudhri et Aheer Histiostoma imbecillus Ashfaq, Chaudhri & Aheer, 1985: 39 Type locality: Pakistan Distribution: Pakistan Histiostoma (H.) incurvatum Mahunka Histiostoma incurvata Mahunka, 1978: 152 Type locality: Canal Zone, Barro Colorado Distribution: Panama Host: Eciton burchelli (Hymenoptera: Formicidae, Dorylinae) Histiostoma (H.) indetonsum (Sevastianov) Anoetus indetonsus Sevastianov, 1973: 852 Histiostoma indetonsum: Klimov & Khaustov, 2018: Type locality: Odesa Distribution: Ukraine Histiostoma (H.) inquilinum Cross Histiostoma inquilinus Cross, 1968: 333 Acarorum Catalogus IX 55 Type locality: Maryland Distribution: USA Host: Xenoglossa strenua (Hymenoptera: Apidae) Histiostoma (H.) insulare (Oudemans) Anoetus insularis Oudemans, 1914: 70; Mahunka, 1967: 15; 1970: 23; 1971: 1198 Histiostoma insulare: Scheucher, 1957: 292 Histiostoma insularis (Oudemans): Hughes & Jackson, 1958: 80 Type locality: Kephallinia Distribution: Germany, Greece (Kephalinia), Holland, Hungary, Mongolia Histiostoma (H.) ionicum Mahunka Histiostoma ionicum Mahunka, 1977: 123 Type locality: Greece Distribution: Greece Histiostoma irinum (Hassan et Sevastianov) Anoetus irinus Hassan & Sevastianov, 1991: 45 Histiostoma irinum (Hassan et Sevastianov): Klimov & Khaustov, 2018: Type locality: Low Egypt Distribution: Egypt Histiostoma (H.) janosi Mahunka Histiostoma janosi Mahunka, 1975: 319 Type locality: New Guinea Distribution: New Guinea Histiostoma (H.) julorum (C.L. Koch) Hypopus julorum C.L.Koch, 1843: H. 38, No20; Berlese, 1886: AMC, 29, 4 Phyllostoma pectineum Kramer, 1876: 36 Histiostoma pectineum Kramer, 1876: 105 Anoetus julorum (C.L. Koch): Vitzthum, 1926a: 480; Sevastianov, 1975: 398 Histiostoma julorum (C.L. Koch): Canestrini, 1888: 17; Canestrini & Kramer, 1899: 135; Haarløv, 1957:32 Type locality: Germany Distribution: Austria, Denmark, Germany, Italy Hosts: Julus mediterraneus, J. unilineatus (Diplopoda: Julida, Julidae) Histiostoma (H.) junctus Ashfaq, Chaudhri et Aheer Histiostoma junctus Ashfaq, Chaudhri & Aheer, 1985: 25 Type locality: Pakistan 56 Acarorum Catalogus IX Distribution: Pakistan Histiostoma (H.) karachiensis Khan Histiostoma karachiensis Khan, 1969: 207 Type locality: Karachi Distribution: Pakistan Histiostoma (H.) kerguelenense Fain Histiostoma kerguelenense Fain, 1976: 312 Type locality: Kerguelen Island Distribution: Kerguelen Island Histiostoma laevis (Oudemans) Zschachia laevis Oudemans, 1929: 451 Type locality: Valkeveen Distribution: Holland Histiostoma (H.) lekiensis Mahunka Histiostoma lekiensis Mahunka, 1979: 241 Type locality: Cameroon Distribution: Cameroon Histiostoma (H.) lineatula Mahunka Histiostoma lineatula Mahunka, 1976: 101 Type locality: Sopron (Hungary) Distribution: Hungary, Romania Host: Platysoma compressum (Coleoptera: Histeridae) Histiostoma (H.) lineosculpturatum Mahunka Histiostoma lineosculpturatum Mahunka, 1978: 121 Type locality: New Britain, Gazelle Pen., Mt. Sinewit Distribution: Papua New Guinea [New Britain] Host: Platolenes sp. (Coleoptera: Tenebrionidae) Histiostoma (H.) litorale (Oudemans) Anoetus litoralis Oudemans, 1914: 71; 1914 : 116 Histiostoma litorale (Oudemans): Scheucher, 1957: 333; Mahunka, 1976b: 294; Lotfollahi et al., 2010 Type locality: Holland (Terschelling I.) Distribution: Germany, Holland, Iran Host: Diplopoda, Coleoptera (Carabidae) Acarorum Catalogus IX 57 Histiostoma (H.) loebli Mahunka Histiostoma loebli Mahunka, 1979: 423 Type locality: Israel Distribution: Israel Histiostoma (H.) loksai Mahunka Histiostoma loksai Mahunka, 1961: 14 Type locality: Hungary Distribution: Hungary Histiostoma longisetum (Mahunka) Anoetus longisetus Mahunka, 1967: 167 Histiostoma longisetum: Klimov & Khaustov, 2018: Type locality: Congo Distribution: Congo Histiostoma (H.) lorentzi (Oudemans) Anoetus lorentzi Oudemans, 1905a: 223 Histiostoma lorentzi (Oudemans): Buitendijk, 1945: 362 Type locality: Perboeangan Distribution: Indonesia (Western New Guinea) Host: Coleoptera: Coccinellidae Histiostoma (H.) louisianae Woodring Histiostoma louisianae Woodring, 1963: 49 Type locality: Louisiana Distribution: USA Histiostoma (H.) luxus Ashfaq, Chaudhri et Aheer Histiostoma luxus Ashfaq, Chaudhri & Aheer, 1985: 36 Type locality: Pakistan Distribution: Pakistan Histiostoma (H.) magdolnae Mahuna et Eraky Histiostoma magdolnae Mahunka & Eraky, 1987: 130 Type locality: Hungary Distribution: Hungary Histiostoma (H.) mahunkai Fain Histiostoma mahunkai Fain, 1974b: 80; Tagami, 2013; 2016: 19 Type locality: Ivory Coast, la Savane de Lamto 58 Acarorum Catalogus IX Distribution: Ivory Coast, Japan Host: Anisolabella (Gonolabis) marginalis (Dermaptera: Anisolabididae) Histiostoma (H.) manicae Samšinák Histiostoma manicae Samšiṅák, 1958: 291 Type locality: Radhošt (Beskyden) Distribution: Czech Rep. Host: Manica rubida (Hymenoptera: Formicidae) Histiostoma (H.) mannai Eraky Histiostoma mannai Eraky, 1999: 53 Type locality: Aswan Distribution: Egypt Host: Associated with Amitermes desertorum (Insecta) Histiostoma (H.) mantelli Clark Histiostoma mantelli Clark, 2010: 10 Type locality: Tongaporutu, North Taranaki Distribution: New Zealand Host: Eudyptula minor (Aves: Spheniscidae) Histiostoma (H.) manuri Eraky Histiostoma manuri Eraky, 2000: 10 Type locality: Egypt Distribution: Egypt Histiostoma (H.) margaritata Mahunka Histiostoma margaritata Mahunka, 1976: 102 Histiostoma margaritatum Mahunka: Klimov & Khaustov, 2018 Type locality: Mecsek – Gebirge (Hungary) Distribution: Hungary, Romania Host: Platysoma compressum (Coleoptera: Histeridae) Histiostoma (H.) marianae Mahunka Histiostoma marianae Mahunka, 1979: 338 Type locality: Osa Pen., 2.5 mi SW of Rincon Distribution: Costa Rica Histiostoma (H.) maritimum (Oudemans) Anoetus maritimus Oudemans, 1914: 70; 1914: 110; Mahunka, 1970: 24 Histiostoma maritimum (Oudemans): Scheucher, 1957: 296; Mahunka, 1976b: 326 Type locality: Holland (Terschelling I.) Acarorum Catalogus IX 59 Distribution: Germany, Holland , Hungary Hosts: Julus unilineatus (Diplopoda: Julidae), Bembidium assimile, B. doris, B. tenellum, Elaphrus riparius, Platynus obscurus Coleoptera: Carabidae), Heterocerus marginatus (Col.: Heteroceridae), Hydrophilus caraboides (Col.: Hydrophilidae) Histiostoma (H.) matsakisi Mahunka Histiostoma matsakisi Mahunka, 1977: 125 Type locality: Greece Distribution: Greece Histiostoma medium (Woodring et Moser) Anoetus media Woodring & Moser, 1970: 1238 Histiostoma medium Klimov & Khaustov, 2018: Type locality: Pineville, Louisiana Distribution: USA (Louisiana, Florida) Host: galleries of Dendroctonus terebrans (Col.: Curculionidae, Scolytinae) Histiostoma (H.) melolonthae Scheucher Histiostoma melolonthae Scheucher, 1957: 335 Type locality: Erlangen Distribution: Germany Hosts: Melolontha hippocastani, Polyphylla fullo (Coleoptera: Scarabaeidae, Melolonthinae) Histiostoma (H.) meszarosorum Mahunka Histiostoma meszarosorum Mahunka, 1978: 113 Type locality: La Habana Distribution: Cuba Host: Acanthoderes circumflexa (Coleoptera: Cerambycidae) Histiostoma (H.) metwallii Fawzy et Mtewallii Histiostoma metwalliii Fawzy & Mtewallii, 2007: 125 (also metwllii) Type locality: Kafr El-Sheikh Distribution: Egypt Histiostoma (H.) minimum Mahunka Histiostoma minimum Mahunka, 1963a: 33 Type locality: Cazombo (Alto Zambeze) Distribution: Angola Histiostoma (H.) mirabile Mahunka Histiostoma mirabile Mahunka, 1981a: 353 Type locality: Micoud, Troumasée Estate 60 Acarorum Catalogus IX Distribution: Windward-Islands Histiostoma (H.) montanum Clark Histiostoma montanum Clark, 2010b: 193 Type locality: North Egmont, Mt Egmont, Taranaki Distribution: New Zealand Host: from shrub Brachyglottis elaeagnifolia litter at 1100–1300 m Histiostoma (H.) moralesi Mahunka Histiostoma moralesi Mahunka, 1978a: 152 Type locality: 4 km W of Villa Neily, 8o40’N, 82o56’W Distribution: Costa Rica Host: Eciton hamatum (Hymenoptera: Formicidae, Dorylinae) Histiostoma (H.) murchiei Hughes et Jackson Histiostoma murchiei Hughes & Jackson, 1958: 88 Type locality: Carp River, Emmet County, Michigan Distribution: USA Host: in cocoons of Allolobophora (Lumbricidae) Histiostoma (H.) muscae Chinniah et Mohanasundaram Histiostoma muscae Chinniah & Mohanasundaram, 1995: 74 Type locality: Tamil Nadu Distribution: India Host: Musca domestica (Diptera: Muscidae) Histiostoma (H.) nasseri Eraky Histiostoma nasseri Eraky, 1994: 159; Abdelgayed et al., 2019: 32 Type locality: Egypt Distribution: Egypt Histiostoma (H.) necopinus (Sevastianov) Anoetus necopinus Sevastianov, 1973: 852 Histiostoma necopinus (Sevastianov): Mahunka, 1976 Type locality: Hajibeyskiy Liman nr. Odessa Distribution: Ukraine Host: Galeruca sp. (Coleoptera: Chrysomelidae) Histiostoma (H.) negmi Eraky Histiostoma negmi Eraky, 2000: 5 Type locality: Sohag Distribution: Egypt Acarorum Catalogus IX 61 Histiostoma (H.) novamsterdamense Fain Histiostoma novamsterdamense Fain, 1976: 565 Type locality: Nouvelle Amsterdam, Pointe d’Entrecasteaux Distribution: Nouvelle Amsterdam Histiostoma onioni Eraky et Shoker Histiostoma onioni Eraky & Shoker, 1994; Abdelgayed et al., 2019: 32 Type locality: Sohag Distribution: Egypt Histiostoma (H.) onthophagae Chinniah et Mohanasundaram Histiostoma onthophagae Chinniah & Mohanasundaram, 1995: 74 Type locality: Tamil Nadu Distribution: India Host: Onthophagus pactolus (Coleoptera: Scarabaeidae) Histiostoma (H.) orientalis Mahunka Histiostoma orientalis Mahunka, 1974: 104 Type locality: unknown Distribution: unknown Host: Halictus tatrazonianellus (Hymenoptera: Halictidae) Histiostoma (H.) oudemansi Scheucher Histiostoma oudemansi Scheucher, 1957: 315 Anoetus oudemansi (Scheucher): Sevastianov, 1975: 400 Type locality: Erlangen Distribution: Germany Histiostoma (H.) ovale (J. Müller) Uropoda ovalis J. Müller, 1860 Histiostoma ovalis (J.Müller): Samšiṅák, 1958: 293; Mahunka, 1977b: 104; Moraza, Fernandez & Jurc, 2013: 597; Vissa et al., 2019 Histiostoma ovale (J. Müller): Klimov & Khaustov, 2018: 53 Type locality: Moravia (Slovakia) Distribution: Austria, Belgium, Bulgaria, Croatia, Czech Rep., France, Germany, Corsica, Hungary, Italy, Poland, Portugal, Romania, Slovakia, Slovenia, Ukraine Hosts: Ips sexdentatus (Coleoptera: Curculionidae - Scolytinae), Platysoma compressum (Col.: Histeridae) Histiostoma (H.) ovalis marani Samšiṅák Histiostoma ovalis f. marani Samšiṅák, 1958: 294 62 Acarorum Catalogus IX Type locality: Vizzavona Distribution: Corsica Host: Ips sexdentatus (Coleoptera: Curculionidae - Scolytinae) Histiostoma (H.) palustre Wirth Histiostoma palustre Wirth, 2003: 262 Type locality: nr. Teufelsee, Berlin/Grunewald Distribution: Germany Histiostoma (H.) papillatum Halliday et Collins Histiostoma papillata Halliday & Collins, 2002: 155 Type locality: Victoria, Deakin University, Warrnambool Distribution: Australia Host: Maccullochella peelii peelii (Pisces: Percichthyidae) Histiostoma (H.) pauciputeolum Fain et Philips Histiostoma pauciputeolum Fain & Philips, 1979: 147 Type locality: Syracuse, N.Y. Distribution: USA Host: In the nest of Otus asio (Aves, Strigiformes) Histiostoma (H.) perruptum Mahunka Histiostoma perruptum Mahunka, 1979: 339 Type locality: Osa Pen., 2.5 mi SW of Rincon Distribution: Costa Rica Histiostoma (H.) phyllophorum (Oudemans) Anoetus phyllophorus Oudemans, 1905b: 238; 1912: 236 Anoetus phyllophorum Oudemans: Mahunka, 1973a: 99 Histiostoma phyllophorum (Oudemans): Scheucher, 1957: 318 Histiostoma nichollsi Womersley, 1941: 484 - Australia Histiostoma phyllophorus (Oudemans): Mahunka & Mahunka-Papp: 1991: 286 Type locality: Leiden Distribution: Australia, Germany, Ghana, Holland, Tanzania Hosts: Oniscus asellus (Isopoda: Onyscidae), Oxythyrea funesta (Coleoptera: Scarabaeidae), Passalidae Histiostoma (H.) phytophaga Ewing Histiostoma phytophaga Ewing, 1911: 41 Type locality: USA Distribution: USA Acarorum Catalogus IX 63 Histiostoma (H.) piceae Scheucher Histiostoma piceae Scheucher, 1957: 304; Beron, 1973; Takov et al. 2009; Walter & Latonas, 2012 Anoetus piceae (Scheucher): Woodring & Moser, 1970: 1255 Type locality: Erlangen, Nurnberg, Regensburg, Harz Distribution: Bulgaria, Canada, Croatia, France, Germany, Japan, Slovakia, Sweden, Russia Host: Ips typographus (Coleoptera: Scolytidae) Histiostoma (H.) pickaxei Eraky et Shoker Histiostoma pickaxei Eraky & Shoker, 1995: 23; Negm & Alatawi, 2011: Type locality: Egypt Distribution: Egypt, Saudi Arabia Histiostoma (H.) piloseta Hughes et Jackson Histiostoma piloseta Hughes & Jackson, 1958: 100 Type locality: Ellerson, Va. Distribution: USA Host: from compost Histiostoma (H.) polaki (Oudemans) Anoetus polaki Oudemans, 1914: 72 Histiostoma polaki (Oudemans): Type locality: Tanzania (“Deutsch Oost Afrika”) Distribution: Spirostreptes sp. (Diplopoda: Spirostreptidae) Histiostoma (H.) polypori (Oudemans) Anoetus polypori Oudemans, 1914: 72 Histiostoma polypori (Oudemans): Buitendijk, 1945: 362; Behura, 1950: 1025; 1956; 1957: 51; Scheucher, 1957: 298; Chmielewski, 1984: 87; Lotfollahi et al., 2010; Tagami, 2016: 19 Type locality: Holland (Oosterbreek) Distribution: Germany, Great Britain, Holland, Hungary, Iran, Poland Hosts: Eugamasus cornutus (Acari: Gamasidae), Forficula auricularia (Dermaptera: Forficulidae) Histiostoma (H.) pontigerum Mahunka Histiostoma pontigerum Mahunka, 1963b: 60 Type locality: Dundo, nr. Cundueji River Distribution: Angola Histiosoma (H.) prodectoris (Vitzthum) Anoetus prodectoris Vitzthum, 1920: 62 64 Acarorum Catalogus IX Histiosoma prodectoris (Vitzthum): Type locality: “Bua-Kraeng in Süd-Celebes und Luzon Philippinen” Distribution: Indonesia (Sulawesi), the Philippines (Luzon) Hosts: Prodector fruhstorferi, Ectocemus cinnamomeus (Coleoptera: Brenthidae) Histiosoma (H.) prophthalmi (Vitzthum) Anoetus prophthalmi Vitzthum, 1920: 59 Histiosoma prophthalmi (Vitzthum): Hughes & Jackson, 1958: 107 Type locality: “Vorderindien, einschl. Assam” Distribution: India Host: Prophthalmus potens (Coleoptera: Brenthidae) Histiostoma (H.) protuberans Hughes et Jackson Histiostoma protuberans Hughes & Jackson, 1958: 109 Type locality: 8 mi west of Richmond, Va. Distribution: USA Histiostoma (H.) pseudolongipes Mahunka Histiostoma pseudolongipes Mahunka, 1992: 398 Type locality: Viti Levu: entre Yanuca et Nadi Distribution: Fiji Host: Diplopoda Histiostoma (H.) pulchrum Kramer Histiostoma pulchrum Kramer, 1886: 259; Scheucher, 1957: 304 Type locality: Germany Distribution: England, Germany Histiostoma (H.) punctaticeps Mahunka Histiostoma punctaticeps Mahunka, 1963b: 61 Type locality: Dundo, nr. Cundueji River Distribution: Angola Histiostoma (H.) pyriforme Michael Histiostoma pyriforme Michael, 1901: 221 Type locality: Great Britain Distribution: Great Britain, Italy Histiostoma (H.) radiferum Scheucher Histiostoma radiferum Scheucher, 1957: 324 Anoetus radiferum (Scheucher): Sevastianov, 1975: 386 Type locality: Erlangen Acarorum Catalogus IX 65 Distribution: Germany Histiostoma (H.) rakha Fawzy et Mtewallii Histiostoma rakha Fawzy & Mtewallii, 2007: 131 Type locality: Kafr El-Sheikh Distribution: Egypt Histiostoma (H.) reticulatissimum Mahunka Histiostoma reticulatissimum Mahunka, 1978: 123 Type locality: NE Wau, Morobe Distr. 1200 m Distribution: Papua New Guinea Histiostoma (H.) reticulofoveolatum Mahunka Histiostoma reticulofoveolatum Mahunka, 1978: 124 Type locality: Karubana, 1500 m Distribution: Papua New Guinea Host: Amarygmus [as “Amargynus”] cf. deformipes (Coleoptera: Tenebrionidae) Histiostoma (H.) rettenmeyerorum Mahunka Histiostoma rettenmeyerorum Mahunka, 1978a: 156 Type locality: Oriente, 00o24S, 76o36’W, Limoncocha Distribution: Ecuador Host: Labidus praedator (Hymenoptera: Formicidae, Dorylinae) Histiostoma (H.) rimosum Mahunka Histiostoma rimosa Mahunka, 1978: 156 Type locality: Oriente, 00o24S, 76o36’W, Limoncocha Distribution: Ecuador Host: Labidum sp. (Hymenoptera: Formicidae, Dorylinae) Histiostoma (H.) rizkii Eraky Histiostoma rizkii Eraky, 1994: 157; Abdelgayed et al., 2019: 32 Type locality: Egypt Distribution: Egypt Histiostoma (H.) rostratum Mahunka Histiostoma rostrata Mahunka, 1978a: 156 Type locality: 4 km W of Villa Neily, 8o40’N, 82o56’W Distribution: Costa Rica Host: Eciton hamatum (Hymenoptera: Formicidae, Dorylinae) 66 Acarorum Catalogus IX Histiostoma (H.) ruehmi Scheucher Histiostoma ruehmi Scheucher, 1957: 331 Anoetus ruehmi (Scheucher): Sevastianov, 1975: 400 Type locality: Nürnberg Distribution: Germany Host: larve of Melanotus rufipes (Coleoptera: Elateridae) Histiostoma (H.) rugosum Mahunka Histiostoma rugosa Mahunka, 1974b: 382 Type locality: Kumasi Distribution: Ghana Histiostoma (H.) sachsi Scheucher Histiostoma sachsi Scheucher, 1957: 319 Anoetus sachsi (Scheucher): Sevastianov, 1975: 400 Type locality: Erlangen Distribution: Germany Histiostoma (H.) sapromyzarum (Dufour) Hypopus sapromyzarum Dufour, 1839: 278 Hypopus filicum Dujardin, 1849: 261 Anoetus sapromyzarum: Oudemans, 1914: 107; Vitzthum, 1926a: 488; Mahunka, 1963c: 497; 1972e: 952 Histiostoma sapromyzarum (Dufour): Vizthum, 1932: 700; Theodorides, 1955; Scheucher, 1957: 302; Mahunka, 1963e: 369 Type locality: France Distribution: Argentina, Australia, Bolivia, Brazil, France, Germany, Greece, Holland, Hungary, Italy, Philippines, Syria Hosts: Melanotus rufipes (Coleoptera: Elateridae); Schizotrachelus bakeri (Brenthidae), Bubas bubalus (Scarabaeidae); Myodes [Clethrionomys] glareolus (Mammalia: Rodentia) Histiostoma (H.) sarrai Eraky et Shoker Histiostoma sarrai Eraky & Shoker, 1994: ? Type locality: Egypt Distribution: Egypt Histiostoma (H.) schauenbergi Mahunka Histiostoma schauenbergi Mahunka, 1978b: 274 Type locality: Indian Ocean Islands Distribution: Indian Ocean Islands Acarorum Catalogus IX 67 Histiostoma (H.) schoutedeni Fain et Elsen Histiostoma schoutedeni Fain & Elsen, 1972: 86 Type locality: Mangai on Kasai River Distribution: DR Congo Host: Glossina fuscipes quanzensis (Diptera: Glossinidae) Histiostoma (H.) scinditum Mahunka Histiostoma scindita Mahunka, 1978: 160; Eraky, 2000: 11 Type locality: Oriente, 00o24S, 76o36’W, Limoncocha Distribution: Ecuador Host: Eciton lucanoides (Hymenoptera: Formicidae, Dorylinae) Histiostoma (H.) sextoni Hughes et Jackson Histiostoma sextoni Hughes & Jackson, 1958: 116 Type locality: Chickahominy Swamp, near Ellerson, Va. Distribution: USA Host: in duck pen Histiostoma shiramba Klimov et Khaustov Histiostoma shiramba Klimov & Khaustov, 2018: 2375 Type locality: Sakhalin Island Distribution: Russia (Sakhalin, Primorskyi Kray) Host: in the galleries of Polygraphus proximus (Coleoptera: Curculionidae, Scolytinae) Histiostoma (H.) simile Mahunka Histiostoma simile Mahunka, 1963a: 34 Type locality: Cavuemba River, Alto Cuílo, Poste de Cacolo Distribution: Angola Histiostoma (H.) siphunculinae Chinniah et Mohanasundaram Histiostoma siphunculinae Chinniah & Mohanasundaram, 1995: 77 Type locality: Tamil-Nadu Distribution: India Host: Siphunculina sp. (Diptera: Chloropidae) Histiostoma (H.) solimani Eraky Histiostoma solimani Eraky, 1997: 107 Type locality: Assiut Distribution: Egypt Histiostoma (H.) spinitarsus (Oudemans) Anoetus spinitarsus Oudemans, 1917: 392 68 Acarorum Catalogus IX Histiostoma spinitarsus (Oudemans): Buitendijk, 1945: 363 Type locality: South America Distribution: South America Host: Phanaeus sp. (Coleoptera: Scarabaeidae) Histiostoma (H.) stammeri Scheucher Histiostoma stammeri Scheucher, 1957: 321 Anoetus stammeri (Scheucher): Sevastianov, 1975: 398 Type locality: Germany Distribution: Germany Hosts: Philonus, Ontolestes (Coleoptera: Staphylinidae) Histiostoma (H.) strenzkei Scheucher Histiostoma strenzkei Scheucher, 1957: 326 Type locality: Plön Distribution: Germany Histiostoma (H.) striatosimile Mahunka Histiostoma striatosimile Mahunka, 1975: 48 Type locality: Ghana Distribution: Ghana Histiostoma (H.) subsimile Mahunka Histiostoma subsimile Mahunka, 1963b: 62 Type locality: forest near Mussalomuca River Distribution: Angola Host: Col.: Carabidae Hirstiosoma (H.) sylvestre (Woodring et Moser) Anoetus gordius of Hughes & Jackson (1958), not A. gordius Vitzthum, 1923 Anoetus sylvestris Woodring & Moser, 1970: 1248 Hirstiosoma sylvestre (Woodring et Moser): Type locality: (Tirol)( Vitzthum, 1923), USA (Ellerson, Va., Hughes & Jackson,1958) Distribution: (Austria, Bosna & Herzegovina) (Vitzthum, 1923) USA (Hughes & Jackson, 1958: 70) Histiostoma (H.) szelenyii Mahunka Anoetus szelenyii Mahunka, 1974 : 101 Histiostoma (H.) szelenyii Mahunka: Fain & Erteld, 1998: 52 Type locality: Afghanistan Distribution: Afghanistan Acarorum Catalogus IX 69 Host: Halictus holtzi (Diptera: Halictidae) Histiostoma (H.) szentivanyi Mahunka Histiostoma szentivanyi Mahunka, 1978: 126 Type locality: NE Nondugl, 2200–2700 m Distribution: Papua New Guinea Host: Amarygmus [as “Amargymus”] cf. capillatus (Coleoptera: Tenebrionidae) Histiostoma (H.) tanypus Fain et Camerik Histiostoma tanypus Fain & Camerik, 1978: 171 Type locality: Cotia, Sao Paulu Distribution: Brazil Host: Hymenoptera: Vespidae gen. sp. Histiostoma (H.) tarsalis Banks (nomen dubium) Histiostoma tarsalis Banks, 1910: 12 Type locality: USA Distribution: USA Histiostoma (H.) telatum Mahunka Histiostoma telatum Mahunka, 1979: 137 Type locality: Tanzania Distribution: Tanzania Host: Coleoptera Histiostoma (H.) tesselata Jacot Histiostoma tesselata Jacot, 1939: 199 Type locality: Western North Carolina Distribution: USA Histiostoma (H.) timidum Fain Histiostoma timidum Fain, 1974b: 82 Type locality: La Savane de Lamto Distribution: Ivory Coast Histiostoma (H.) tinydorsalis Eraky Histiostoma tinydorsalis Eraky, 1999: 70 Type locality: Upper Egypt (Assiut) Distribution: Egypt Histiostoma (H.) titanolabi Tagami et Halliday Histiostoma titanolabi Tagami & Halliday, 2013: 222 70 Acarorum Catalogus IX Type locality: Australia Distribution: Australia Host: Titanolabis colossea (Dermaptera: Anisolabididae) Histiostoma (H.) toxopei (Oudemans) Anoetus toxopei Oudemans, 1927: 230 Histiostoma toxopei (Oudemans): Buitendijk, 1945: 363 Type locality: Wai Eno – Wai Temoen, Boeroe Distribution: Indonesia Histiostoma (H.) trichophorum (Oudemans) Anoetus trichophorus Oudemans, 1912: 235 Histiostoma trichophorus (Oudemans): Buitendijk, 1945: 363 Histiostoma trichophorum (Oudemans): Scheucher, 1957: 306 Type locality: Mkulumusi Distribution: Tanzania Hosts: Helops quisquilius, Hypophloeus bicolor (Coleoptera: Tenebrionidae); Platypus dispar (Col.: Platypodidae) Histiostoma (H.) ulmi Scheucher Histiostoma ulmi Scheucher, 1957: 308; Anoetus ulmi (Scheucher): Woodring & Moser, 1970: 1256; Sevastianov, 1975: 400 Type locality: Erlangen Distribution: Germany Host: Hypophloeus bicolor (Coleoptera: Tenebrionidae) Histiostoma (H.) umbrosa Mahunka Histiostoma umbrosa Mahunka, 1975b: 51 Type locality: Gambia Distribution: Gambia Histiostoma (H.) unidentatum Fain, Athias-Binche et Carton Histiostoma unidentatum Fain, Athias-Binche & Carton, 1993: 71 Type locality: Thailand Distribution: Australia, Thailand Host: Drosophila bipectinata (Diptera) imported from Thailand Histiostoma (H.) uvafer Mahunka Histiostoma uvafer Mahunka, 1974b: 383 Type locality: Kwandaso Distribution: Ghana Acarorum Catalogus IX 71 Histiostoma (H.) uxoria Mahunka Histiostoma uxoria Mahunka, 1974b: 385 Type locality: Tamale Distribution: Ghana Histiostoma (H.) validum Banks Histiostoma valida Banks, 1906: 141 Type locality: Canada Distribution: Canada Histiostoma (H.) varium (Woodring et Moser) Anoetus varia Woodring & Moser, 1970: 1243; Moser & Roton, 1971; Moser, 1976: 811; Histiostoma varia (Woodring et Moser): Stone & Simpson, 1991; Type locality: Elizabeth, La., USA Distribution: Australia, USA, Honduras, Mexico, Canada Hosts: boring dust of Ips avulsus, I. calligraphus, I. grandicollis, Dendroctonus frontalis (Coleoptera: Curculionidae, Scolytinae), phoretic on Carcinops sp. (Histeridae) and Corticeus coynei (Tenebrionidae) Histiostoma (H.) variabilis Mahunka Histiostoma variabilis Mahunka, 1979: 244 Type locality: Cameroon Distribution: Cameroon Histiostoma (H.) verruca Jacot Histiostoma verruca Jacot, 1939: 201 Type locality: Western North Carolina Distribution: USA Histiostoma (H.) verrucogenicula Jacot Histiostoma verrucogenicula Jacot, 1939: 200 Type locality: Western North Carolina Distribution: USA Hirstiosoma virginianum Woodring et Moser Hirstiosoma virginiana Woodring & Moser, 1970: 1255 Hirstiosoma virginianum Woodring & Moser: Type locality: Virginia, Florida, Tennessee Distribution: USA Histiostoma (H.) volgini Mahunka Histiostoma volgini Mahunka, 1979: 138 72 Acarorum Catalogus IX Type locality: Transvaal, Nylsvley Reservation Distribution: South Africa Histiostoma (H.) vulpis (Dujardin) Hypopus vulpis Dujardin, 1849: 263 Histiostoma vulpis (Dujardin): Type locality: Transvaal, Nylsvley Reservation Distribution: South Africa Histiostoma (H.) watanabei Kurosa Histiostoma watanabei Kurosa, 2002: 28 Type locality: Japan Distribution: Japan Host: Phoretic on Lathrobium (Lathrobium) (Coleoptera: Staphylinidae) Histiostoma (H.) wauense Mahunka Histiostoma wauense Mahunka, 1976b: 311 Type locality: Torricelli Mts. Distribution: Papua New Guinea Host: Platolenes femoratus (Coleoptera: Tenebrionidae) Histiostoma (H.) zaheri Fawzy et Mtewallii Histiostoma zaheri Fawzy & Mtewallii, 2007: 124 Type locality: Kafr El-Sheikh Distribution: Egypt Subgenus Cavanoetus Mahunka Cavanoetus Mahunka, 1982: 290 Type species: Histiostoma pierrestrinatii Mahunka, 1982 Histiostoma (Cavanoetus) pierrestrinatii Mahunka Histiostoma (Cavanoetus) pierrestrinatii Mahunka, 1982: 290 Type locality: New Mexico Distribution: USA Subgenus Ichtanoetus Fain et Belpaire Ichtanoetus Fain & Belpaire, 1985: 286 Type species: Histiostoma anguillarum Fain et Belpaire, 1985 Histiostoma (Ichtanoetus) anguillarum Fain et Belpaire Histiostoma (Ichtanoetus) anguillarum Fain & Belpaire, 1985: 287 Acarorum Catalogus IX 73 Type locality: Loire Distribution: France Host: from the gills of young eels (Anguilla Anguilla, Anguillidae, Anguilliformes) Histiostoma (Ichtanoetus) cyrtandrae (Vitzthum) Anoetus cyrtandrae Vitzthum, 1931: 110 Histiostoma (Ichtanoetus) cyrtandrae (Vitzthum): Fain & Belpaire, 1985: 287 Type locality: Java Distribution: Indonesia Host: in the axill fluid of Colocasia indica Histiostoma (Ichtanoetus) nigrellii Hughes et Jackson Histiostoma nigrellii Hughes & Jackson, 1958: 95 Histiostoma (Ichtanoetus) nigrellii: Fain & Belpaire, 1985: 287 Type locality: aquarium in New York Distribution: USA Host: goldfish tank Histiostoma (Ichtanoetus) ocellatum Fain et Lambrechts Histiostoma (Ichtanoetus) ocellatum Fain & Lambrechts, 1987: 95 Type locality: aquarium containing Discus fish, Antwerp Distribution: Holland Histiostoma (Ichtanoetus) piscium Fain et Lambrechts Histiostoma piscium Fain & Lambrechts, 1985: 121; Tagami, 2016: Histiostoma (Ichtanoetus) piscium: Fain & Belpaire, 1985: 287 Type locality: aquarium in Antwerpen (imported from Thailand) Distribution: Japan, Thailand Hosts: Pangasius sutchi (Pisces: Siluriformes, Pangasiidae); Periplaneta fuliginosa (Dermaptera: Blattidae) Genus Hormosianoetus Fain Hormosianoetus Fain, 1980: 753 Type species: Hormosianoetus aeschlimanni Fain, 1980 Hormosianoetus aeschlimanni Fain Hormosianoetus aeschlimanni Fain, 1980: 756; Eraky & Shoker 1993: 56 Type locality: Zurich Distribution: Switzerland Host: Drosophila sp. (Diptera: Drosophilidae) 74 Acarorum Catalogus IX Hormosianoetus laboratorium (Hughes) Histiostoma laboratorium Hughes, 1950: 177; Scheucher, 1957: 333 ? Histiostoma genetica (Stolpe, 1938) Hormosianoetus laboratorium (Hughes): Fain, Athias-Binche & Carton, 1993: 75 Type locality: Switzerland Distribution: Guadeloupe, Morocco, Switzerland Hosts: Drosophyla willstoni, D. subobscura (Diptera: Drosophylidae) Hormosianoetus mahunkai Eraky et Shoker Hormosianoetus mahunkai Eraky & Shoker, 1993: 53; Eraky & Mohamed, 2019: 2 Type locality: from Sohag governorate Distribution: Egypt Host: from Fusarium culture Hormosianoetus mallotae (Fashing) Mauduytia mallotae Fashing, 1973: 455 Hormosianoetus mallotae: Fain, 1980: 755 Type locality: Johnson County (Texas) Distribution: USA (Texas, Pennsylvania, Illinois) Hosts: Mallota posticata, M. bautius (Diptera: Syrphidae) Genus Hymenanoetus Mahunka Hymenanoetus Mahunka, 1963a: 39 Type species: Hymenanoetus nodosus Mahunka, 1963 Hymenanoetus nodosus Mahunka Hymenanoetus nodosus Mahunka, 1963a: 39 Type locality: Angola, Lumeje II, N of Santando Station of C.F.B., Poste de Bussaco (80 km E of Luso) Distribution: Angola Genus Insulanoetus Sevastyanov Insulanoetus Sevastyanov, 1973: 855 Type species: Insulanoetus brachypodus Sevastyanov, 1973 Insulanoetus brachypodus Sevastyanov Insulanoetus brachypodus Sevastyanov, 1973: 856 Type locality: Jaralgach Island (Black Sea) Distribution: Crimea Hosts: Bledius bicornis, B. sarmaticus (Coleoptera: Staphylinidae, Oxythelinae) Acarorum Catalogus IX 75 Genus Kanoetus Samšinák Kanoetus Samšinák, 1966: 126 Type species: Kanoetus cubanus Samšinák, 1966 Kanoetus australis Mahunka Kanoetus australis Mahunka, 1975a: 60 Type locality: Kpington Distribution: South Africa Kanoetus contrasetosus Mahunka Kanoetus contrasetosus Mahunka, 1974 : 388 Type locality: Banda-Nkwanta Distribution: Ghana Kanoetus cubanus Samšinák Kanoetus cubanus Samšinák, 1966: 126 Type locality: Cuba Distribution: Cuba Kanoetus samsinaki Mahunka Kanoetus samsinaki Mahunka, 1974: 386 Type locality: Kumasi Distribution: Ghana Genus Kaszabanoetus Mahunka Kaszabanoetus Mahunka, 1976b: 317 Type species: Kaszabanoetus ulomae Mahunka, 1976 Kaszabanoetus ulomae Mahunka Kaszabanoetus ulomae Mahunka, 1976b: 318 Type locality: Akcagoga Distribution: Turkey Host: Uloma culinaria (Coleoptera: Tenebrionidae) Genus Loxanoetus Fain Loxanoetus Fain, 1970: 173 Type species: Loxanoetus bassoni Fain, 1970 [Redescription, Fain 1983: 847] Loxanoetus bassoni Fain Loxanoetus bassoni Fain, 1970: 173; Fain & Zumpt, 1974: 99; Fain 1983: 847 Type locality: Kruger Park (South Africa) 76 Acarorum Catalogus IX Distribution: South Africa, Namibia, Uganda Hosts: Loxodonta africana (Mammalia, Proboscidea: Elephantidae), Syncerus caffer (Artiodactyla: Bovidae) Loxanoetus lenae Domrow et Ladds Loxanoetus lenae Domrow & Ladds, 1984: 759 Type locality: Townsville, Qeensland Distribution: from zoo in Australia Host: Elephas maximus (Mammalia, Proboscidea: Elephantidae) Genus Munduytia Oudemans Munduytia Oudemans, 1929: 450 [also “Mauduytia” in Scheucher, 1957] Type species: Anoetus tropicus Oudemans, 1911 Munduytia tropicus (Oudemans) Anoetus tropicus Oudemans, 1911a: 190 Munduytia tropicus (Oudemans): Oudemans, 1929: 450 Mauduytia tropicus: Scheucher, 1957: 379 Type locality: Java, Banjoewangi Distribution: Indonesia (Java) Host: Blattodea fam., gen. sp. Genus Nepenthacarus Fashing Nepenthacarus Fashing, 2002: 8 Type species: Nepenthacarus warreni Fashing, 2002 Nepenthacarus warreni Fashing Nepenthacarus warreni Fashing, 2002: 8 Type locality: Far North Queensland Distribution: Australia Genus Otanoetus Fain et Zumpt Otanoetus Fain & Zumpt, 1974: 100 [placement?] Type species: Otanoetus wetzeli Fain et Zumpt, 1974 Otanoetus wetzeli Fain et Zumpt Otanoetus wetzeli Fain & Zumpt, 1974: 100 Type locality: Karumu Game Reserve (Uganda) Distribution: Namibia, Uganda Hosts: Loxodontha africana (Proboscidea: Elephantidae), Syncerus caffer (Artiodactyla: Bovidae) Acarorum Catalogus IX 77 Genus Ovanoetus Fain et van Goethem Ovanoetus Fain & van Goethem, 1985: 227 Type species: Ovanoetus wautersi Fain et van Goethem, 1985 Ovanoetus wautersi Fain et van Goethem Ovanoetus wautersi Fain & van Goethem, 1985: 228 Type locality: St. Vincent Republic, Grenadines Isl., Bequia Id. (Windward Group) Distribution: St. Vincent Republic, Grenadines Isl., Bequia Id. (Windward Group) Host: Coenobita clypeata (Crustacea: Paguridae) Genus Peripatetes Mahunka Peripatetes Mahunka, 1976b: 315 Type species: Peripatetes acanthotarsus Mahnunka, 1976 Peripatetes acanthotarsus Mahunka Peripatetes acanthotarsus Mahunka, 1976b: 315 Type locality: Budafok Distribution: Hungary Host: Staphylinidae (Coleoptera) Genus Porrhanoetus Mahunka Porrhanoetus Mahunka, 1963a: 41 Type species: Porrhanoetus machadoi Mahunka, 1963 Porrhanoetus machadoi (Mahunka) Porrhanoetus machadoi (Mahunka): Mahunka, 1963a: 41; 1967: 168 Type locality: Angola, Cavuemba River, Alto Cuílo, Poste de Cacolo Distribution: Angola, Congo - Brazzaville Genus Probonomoia Fain et Rack Probonomoia Fain & Rack, 1987: 70 Type species: Probonomoia columbiana Fain et Rack, 1987 Probonomoia africana (Mahunka) Bonomoia africana Mahunka, 1967: 171 Probonomoia africana (Mahunka): Fain & Rack 1987: 71 Type locality: Mount Foari Reservation Distribution: Congo (Brazzaville) Probonomoia chrysis Dimitrieff et Fain Probonomoia chrysis Dimitrieff & Fain, 1994: 105 78 Acarorum Catalogus IX Type locality: Spain Distribution: Spain Host: Chrysis sp. (Hymenoptera: Chrysididae) Probonomoia columbiana Fain et Rack Probonomoia columbiana Fain & Rack, 1987: 71 Type locality: Paramo de Chisaca, 40 km south of Bogota Distribution: Columbia Host: from the flower of Espeletia grandiflora (Asteraceae) Probonomoia congoensis (Fain et Elsen) Bonomoia congoensis Fain & Elsen, 1972: 89 Probonomoia congoensis (Fain et Elsen): Fain & Rack, 1987: 71 Type locality: Mongati Distribution: DR Congo Host: Glossina fuscipes quanzensis (Diptera: Glossinidae) Probonomoia humprechti (Samšinák) Bonomoia humprechti Samšinák, 1956: 354 Probonomoia humprechti (Samšinák): Klimov & Khausov, 2018: 2411 Type locality: Czech Republic Distribution: Czech Republic, Hungary, Romania Hosts: Laphria flava (Diptera: Asilidae), Ips spp. (Coleoptera: Cuculionidae, Scolytinae), Acanthocinus aedilis (Col.: Cerambycidae) Probonomoia opuntiae Wirth Bonomoia opuntiae Wirth, 2005: 303 Probonomoia opuntiae (Wirth): Type locality: Sardinia Distribution: Sardinia, Corsica, Canary Islands (Gomera) Probonomoia pini (Scheucher) Bonomoia pini Scheucher, 1957: 366; Mahunka, 1970: 68; Woodring & Moser, 1970: 1256; Probonomoia pini (Scheucher): Fain & Rack 1987: 71; Khaustov et al., 2018: 62 Type locality: Erlangen (Germany) Distribution: Croatia, Czech Republic, England, Finland, Germany, Hungary, Japan, Poland, Russia Hosts: Hylaster ater, Ips acuminatus, I. cembrae, I. typographus (Coleoptera: Curculionidae, Scolytinae), Dorcus parallelipipedus (Coleoptera: Lucanidae), Acanthocinus aedilis (Col.: Cerambycidae), Helops quisquilius (Col.: Tenebrionidae) Acarorum Catalogus IX 79 Probonomoia sibirica (Wirth) Bonomoia sibirica Wirth, 2016: 99 Probonomoia sibirica: Khaustov et al., 2018: 64 Type locality: Tyumen Prov., East of Tyumen Distribution: Russia Hosts: Dendroctonus micans, Ips typographus (Coleoptera: Curculionidae, Scolytinae) Probonomoia spinifera (Scheucher) Bonomoia spinifera Scheucher, 1957: 367 Probonomoia spinifera (Scheucher): Fain & Rack, 1987: 71 Type locality: Erlangen Distribution: Germany, Slovakia Genus Prowichmannia Radford Wichmannia Oudemans, 1929: 450 [praeocc.] Prowichmannia Radford, 1950: 222 Type species: Histiostoma spiniferum Michael, 1901 Prowichmannia spiniferum (Michael) Histiostoma spinifera Michael, 1901: 215; Oudemans, 1903: Anoetus spiniferus Michael: Oudemans, 1902: 46 Wichmannia spiniferum (Michael): Oudemans, 1929: Wichmannia spiniferus (Michael): Buitendijk, 1945: 361 Wichmannia spinifera (Michael): Scheucher, 1957: 370 Prowichmannia spiniferum (Michael): Radford, 1950: [160]; Beron, 1973: 22 Prowichmannia spinifera (Michael): Mahunka, 1970: 72; Krivolucliy, 1975: 413 Type locality: England Distribution: Bulgaria, England, Germany, Hungary, Poland, Ukraine Hosts: Cetonia aurata (Coleoptera: Scarabaeidae); many species of Mammals (Soricomorpha and Rodentia) Prowichmannia stammeri (Scheucher)(new combination) Wichmannia stammeri Scheucher, 1957: 374 Prowichmannia stammeri: p.p. Type locality: Erlangen Distribution: Germany Prowichmannia talpae (Scheucher) Wichmannia talpae Scheucher, 1957: 372 Prowichmannia talpae: Zamec, 2014: 80 Acarorum Catalogus IX Type locality: England Distribution: Germany, Slovakia Host: nest of Talpa europaea (Soricomorpha: Talpidae) Genus Psyllanoetus Fain et Beaucournu Psyllanoetus Fain et Beaucournu , 1973: 516 Type species: Psyllanoetus Fain et Beaucournu , 1973 Psyllanoetus afropulicarius Fain et Beaucournu Psyllanoetus afropulicarius Fain & Beaucournu, 1973: 516 Type locality: Lemera, Kivu Distribution: DR Congo Hosts: Listropsylla dolosa (Siphonaptera: Hystrichopsyllidae), ex Praomys jacksoni (Rodentia: Muridae), Stivalius torvus (Siphonaptera), ex Hybomys univittatus (Rodentia), Dinopsyllus apistus (Siphonaptera), Ctenophthalmus devignati (Siphonaptera), ex H. univittatus (Rodentia) Psyllanoetus dinopsyllus Fain et Beaucournu Psyllanoetus dinopsyllus Fain et Beaucournu, 1973: 517 Type locality: Lemera, Kivu Distribution: DR Congo Hosts: Dinopsyllus apistus (Siphonaptera), ex Praomys jacksoni (Rodentia: Muridae), Ctenophthalmus debrauweri (Siphonaptera), ex Praomys jacksoni (Rodentia), Ct. calceatus cabirus (Siphonaptera), ex Otomys sp. (Rodentia) Psyllanoetus melesicola Fain et Beaucournu Psyllanoetus melesicola Fain & Beaucournu, 1973: 516 Type locality: Les Sieges, Yonne Distribution: France Host: Paraceras melis (Siphonaptera: Ceratophyllidae), ex Meles meles (Carnivora: Mustelidae) Genus Pteranoetus Mahunka Pteranoetus Mahunka, 1978: 128 Type species: Pteranoetus kaszabi Mahunka, 1978 Pteranoetus incertus Mahunka Pteranoetus incertus Mahunka, 1978a: 162 Type locality: Puerto Viejo, 10o26’N, 83o56’ Distribution: Costa Rica Host: Eciton burchelli (Hymenoptera: Formicidae, Dorylinae) Acarorum Catalogus IX 81 Pteranoetus kaszabi Mahunka Pteranoetus kaszabi Mahunka, 1978: 128 Type locality: NE Nondugl, 2200–2700 m Distribution: Papua New Guinea Pteranoetus latus Mahunka Pteranoetus latus Mahunka, 1978a: 162 Type locality: Oriente, 00o24S, 76o36’W, Limoncocha Distribution: Ecuador Pteranoetus rettenmeyeri Mahunka Pteranoetus rettenmeyeri Mahunka, 1978a: 166 Type locality: Oriente, 00o24S, 76o36’W, Limoncocha Distribution: Ecuador Host: Neivamyrmex gradualis (Hymenoptera: Formicidae, Dorylinae) Genus Rhaphidothrix Mahunka Rhaphidothrix Mahunka, 1967: 173 Type species: Rhaphidothrix gozmanyi Mahunka, 1967 Rhaphidothrix gozmanyi Mahunka Rhaphidothrix gozmanyi Mahunka, 1967a: 173 Type locality: Kindamba, Meya, Louolo river Distribution: Rep. of the Congo (Brazzaville) Rhaphidothrix punctolineatus Mahunka Rhaphidothrix punctolineatus Mahunka, 1973a: 106; 1979: 128 Type locality: Pretsea Distribution: Ghana Host: Coleoptera Rhaphidothrix simplex Mahunka Rhaphidothrix simplex Mahunka, 1974b: 390 Type locality: Kumasi Distribution: Ghana Genus Rhopalanoetus Scheucher Rhopalanoetus Scheucher, 1957: 362 Type species: Anoetus lanceocrinus Oudemans, 1914 82 Acarorum Catalogus IX Rhopalanoetus bifurcatus Mahunka Rhopalanoetus bifurcatus Mahunka, 1961b: 529; 1965: 449; 1969b: 371 Rhopalanoetus furcatus Mahunka: Samšiṅák, 1962 Type locality: Mt Meru, eastern slope (Tanzania) Distribution: Rep. of the Congo (Brazzaville), Tanzania, DR Congo Host: elephant excrement; Onitis sphinx (Coleoptera: Scarabaeidae) Rhopalanoetus chinensis Samšiṅák Rhopalanoetus chinensis Samšiṅák, 1962b: 188 Type locality: Kanton [Guandong] Distribution: China Host: Oniticellus cinctus (Coleoptera: Scarabaeidae) Rhopalanoetus endroedyyoungai Mahunka Rhopalanoetus endroedyyoungai Mahunka, 1979: 144 Type locality: Transvaal, Nylsvley Res., 35 km NE from Pretoria Distribution: South Africa Rhopalanoetus fimetarius (G. Canestrini et Berlese) Histiostoma fimetarium Canestrini & Berlese, 1881: 150 (not Histiostoma fimetarius sensu Hughes & Jackson, 1958) Rhopalanoetus fimetarius (G. Canestrini et Berlese): Samsinak, 1962: 96; Mahunka, 1970: 66; Bongers et al., 1985: 48 Anoetus lanceocrinus Oudemans, 1914: 72 Histiostoma lanceocrinus: Buitendijk, 1945: 362; Hughes & Jackson, 1958: 82 Rhopalanoetus lanceocrinus: Scheucher, 1957: 362; Mahunka, 1962: 117 Anoetus tienhoveni Oudemans, 1926: 123 – Type loc.: Naardelmeer, Holland Histiostoma tienhoveni: Buitendijk, 1945: 363 Type locality: Padova (Italy) Distribution: Holland, Hungary, Italy Rhopalanoetus gladifer Mahunka Rhopalanoetus gladifer Mahunka, 1969b: 372 Type locality: Mt Meru, eastern slope Distribution: Tanzania Rhopalanoetus heterotrichus Mahunka Rhopalanoetus heterotrichus Mahunka, 1973: 111 Type locality: Ghana Distribution: Ghana Acarorum Catalogus IX 83 Rhopalanoetus hirsutus Mahunka Rhopalanoetus hirsutus Mahunka, 1973a: 305 Type locality: Nakpanduri Distribution: Ghana Rhopalanoetus lanceocrinus (Oudemans) Anoetus lanceocrinus Oudemans, 1914: 72 Rhopalanoetus lanceocrinus (Oudemans): Scheucher, 1957: 362 Type locality: Holland Distribution: Germany, Holland Hosts: Aphodius fimetarius, A. fossor, A. merdarius, Geotrupes stercorarius (Coleoptera: Scarabaeidae), Sphaeridium sp. (Col.: Hydrophilidae) Rhopalanoetus robustus Mahunka Rhopalanoetus robustus Mahunka, 1973a: 112 Type locality: Meru Distribution: Tanzania Rhopalanoetus simplex Samšiṅák Rhopalanoetus simplex Samšiṅák, 1962b: 187 Type locality: Kanton [Guandong] Distribution: China Host: Oniticellus cinctus (Coleoptera: Scarabaeidae) Rhopalanoetus trogicola (Fain, Vangeluwe, Degreef et Wauthy) Histiostoma trogicola Fain, Vangeluwe, Degreef & Wauthy, 1993: 13 Rhopalanoetus trogicola (Fain, Vangeluwe, Degreef et Wauthy): Mašan, 2000: Type locality: near Namur (Belgium) Distribution: Belgium, Slovakia Hosts: Trox scaber, T. cadaverinus (Coleoptera: Trogidae) Genus Richardanoetus Samšinák Richardanoetus Samšinák, 1989: 99 Type species: Richardanoetus demel Samšinák, 1989 Richardanoetus demel Samšiṅák Richardanoetus demel Samšiṅák, 1989: 99 Type locality: Fuvo Distribution: Ecuador Host: Richardia sp. (plant) 84 Acarorum Catalogus IX Genus Sarraceniopus Fashing et OConnor Sarraceniopus Fashing & OConnor, 1984: 217 Type species: Zwickia gibsoni Nesbitt, 1954 Sarraceniopus darlingtoniae Fashing et OConnor Sarraceniopus darlingtoniae Fashing & OConnor, 1984: 219 Type locality: pitchers of Darlingtonia californica, Darlingtonia wayside, Lane County, Oregon Distribution: USA (Oregon) Sarraceniopus gibsoni (Nesbitt) Zwickia gibsoni Nesbitt, 1954: 454 Anoetus gibsoni (Nesbitt): Atyeo, 1978: 244 Sarraceniopus gibsoni (Nesbitt): Fashing & OConnor, 1984: 219 Type locality: Danford Lake Area, Quebec Distribution: Canada (N.S., Quebec, Ontario) Host: the pitchers of Sarracenia purpurea (Sarraceniaceae) Sarraceniopus hughesi (Hunter et Hunter) Anoetus hughesi Hunter & Hunter, 1964: 39 Sarraceniopus hughesi (Hunter & Hunter): Fashing & OConnor, 1984: 219; Tagami, 2004: 169 Type locality: Georgia and North Carolina (USA) Distribution: Japan (imported with plants), USA Host: Sarracenia plants Genus Scheucheria Mahunka Scheucheria Mahunka, 1969a: 184 Type species: Scheucheria mongolica Mahunka, 1969 Scheucheria mongolica Mahunka Scheucheria mongolica Mahunka, 1969a: 184 Type locality: Mongolia Distribution: Mongolia Genus Scutanoetus Mahunka Scutanoetus Mahunka, 1969b: 367 [placement?] Type species: Scutanoetus permixtus Mahunka, 1969 Scutanoetus permixtus Mahunka Scutanoetus permixtus Mahunka, 1969b: 367 Acarorum Catalogus IX 85 Type locality: Mt Meru, eastern slope Distribution: Tanzania Genus Sellea Oudemans Sellea Oudemans, 1929: 449 Type species: Histiostoma pulchrum Kramer, 1886 Sellea pulchrum (Kramer) Histiostoma pulchrum Kramer, 1886: 259; Li et al., 1992 Sellea pulchrum (Kramer): Oudemans, 1929: 449; Hughes & Jackson, 1958: 175 Type locality: Germany Distribution: England, Germany, China Genus Semianoetus Mahunka Semianoetus Mahunka, 1976b: 313 Type species: Semianoetus mrcziaki Mahunka, 1976 Semianoetus banksi Mahunka Semianoetus banksi Mahunka, 1979: 341 Type locality: Puntarenas, Monteverde, 1400 m Distribution: Costa Rica Semianoetus mrcziaki Mahunka Semianoetus mrcziaki Mahunka, 1976b: 313 Type locality: Budapest, Hüvõsvõlgy Distribution: Hungary Genus Spinanoetus Scheucher Spinanoetus Scheucher, 1957: 357 Type species: Spinanoetus pelznerae Scheucher, 1957 Spinanoetus heterotrichus Mahunka Spinanoetus heterotrichus Mahunka, 1962: 432; Kaluz, 1992: Type locality: Dunazuggebirge, Sikáros Distribution: Hungary, Slovakia Spinanoetus pelznerae Scheucher Spinanoetus pelznerae Scheucher, 1957: 358; Káluz, 1993; Haitlinger, 2004: Type locality: Erlangen Distribution: Germany, Hungary, Poland 86 Acarorum Catalogus IX Hosts: Necrophorus interruptus, Oeceoptoma thoracica, Thanatophilus rugosus, Th. sinuatus (Coleoptera: Silphidae) Spinanoetus variabilis Mahunka Spinanoetus variabilis Mahunka, 1973a: 109 Type locality: Nakpanduri Distribution: Ghana Spinanoetus weingaertnerae Scheucher Spinanoetus weingaertnerae Scheucher, 1957: 360; Haitlinger, 2004; Zamec, 2014 Type locality: Erlangen Distribution: Germany, Hungary, Poland, Slovakia Hosts: Emus hirtus, Ontolestes sp., Philontus sp. (Coleoptera: Staphylinidae) Genus Stercoranoetus Mahunka et Mahunka-Papp Stercoranoetus Mahunka & Mahunka-Papp, 1991: 285 Type species: Stercoranoetus harteni Mahunka et Mahunka-Papp, 1991 Stercoranoetus harteni Mahunka et Mahunka-Papp Stercoranoetus harteni Mahunka & Mahunka-Papp, 1991: 285 Type locality: Santiago: Santa Cruz Distribution: Cabo Verde Stercoranoetus woolleyi (Mahunka et Eraky) Histiostoma woolleyi Mahunka & Eraky, 1987: 135 Stercoranoetus woolleyi (Mahunka et Eraky): Mahunka & Mahunka- Papp, 1991: 285 Type locality: Hungary Distribution: Hungary Genus Synanoetus Mahunka Synanoetus Mahunka, 1972d: 311 Type species: Synanoetus baloghi Mahunka, 1972 Synanoetus baloghi Mahunka Synanoetus baloghi Mahunka, 1972d: 311 Type locality: Las Cardas (Prov. Coquimbo) Distribution: Chile Genus Syringanoetus Fain Syringanoetus Fain, 1980: 116 Type species: Syringanoetus vandenberghi Fain, 1980 Acarorum Catalogus IX 87 Syringanoetus faini Mahunka et Eraky Syringanoetus faini Mahunka & Eraky, 1987: 137 Type locality: Hungary Distribution: Hungary Syringanoetus vandenberghi Fain Syringanoetus vandenberghi Fain, 1980: 116; Mahunka & Eraky 1987: 137 Type locality: Rugege Forest, 2500 m, 75 km from Butare Distribution: Rwanda Genus Teinokyra Mahunka Teinokyra Mahunka, 1973a: 306 Type species: Teinokyra lineata Mahunka, 1973 Teinokyra lineata Mahunka Teinokyra lineata Mahunka, 1973a: 307 Type locality: Pretsea Distribution: Ghana Genus Tensiostoma Wurst et Kovac Tensiostoma Wurst & Kovac, 2003: 65 Type species: Tensiostoma veliaphilum Wurst et Kovac, 2003 Tensiostoma veliaphilum Wurst et Kovac, 2003 Tensiostoma veliaphilum Wurst & Kovac, 2003: 66 Type locality: Selangor, Darul Ehsan, Ulu Gombak Field Studies Centre, Univ. of Malaya Distribution: Malaysia (Selangor) Hosts: Lathriovelia rickmersi, L. capitata (Heteroptera: Veliidae) Genus Traskorchestianoetus Fain et Colloff Traskorchestianoetus Fain & Colloff, 1990: 667 Type species: Traskorchestianoetus spiceri Fain et Colloff, 1990 Traskorchestianoetus brevipes Fain et Colloff Traskorchestianoetus brevipes Fain & Colloff, 1990: 672 Type locality: Dianna Island, Dodger Sound, British Columbia Distribution: Canada Host: Traskorchestia traskiana (Crustacea: Talitridae); marine Traskorchestianoetus spiceri Fain et Colloff Traskorchestianoetus spiceri Fain & Colloff, 1990: 668 88 Acarorum Catalogus IX Type locality: Dianna Island, Dodger Sound, British Columbia Distribution: Canada Host: Traskorchestia traskiana (Crustacea: Talitridae); marine Genus Xenanoetus Mahunka Xenanoetus Mahunka, 1969: 359 Type species: Xenanoetus vestigialis Mahunka, 1969 Xenanoetus afrotropicalis Fain Xenanoetus afrotropicalis Fain, 1984: 281 Type locality: Kamobuga, Rugege Forest, 2500 m Distribution: Rwanda Host: elephant dung Xenanoetus grandiceps Mahunka Xenanoetus grandiceps Mahunka, 1973: 61 Type locality: Mongolia Distribution: Mongolia Xenanoetus kabrt Samšiṅák Xenanoetus kabrt Samšiṅák, 1989: 103 Type locality: DR Congo Distribution: DR Congo Host: Gymnometopina kibokoensis (Diptera: Sphaeroceridae) Xenanoetus kofrankova Samšiṅák Xenanoetus kofrankova Samšiṅák, 1989: 101 Type locality: Washington Distribution: USA Host: Coproica acutangula (Diptera: Sphaeroceridae) Xenanoetus vestigialis Mahunka Xenanoetus vestigialis Mahunka, 1969a: 359 Type locality: Finland Distribution: Finland Host: Sphaeridium sp. (Coleoptera: Hydrophiilidae) Genus Zwickia Oudemans Zwickia Oudemans, 1924: 310 [Diagnostic, Nesbitt 1979: 1205] Type species: Anoetus guentheri Oudemans, 1915 Acarorum Catalogus IX 89 Zwickia colocasiae (Vitzthum) Anoetus colocasiae Vitzthum, 1931: 118 Zwickia colocasiae (Vitzthum): Hughes & Jackson, 1958: 182 Type locality: Ngebal Lake Distribution: Indonesia (Java) Zwickia guentheri (Oudemans) Anoetus guentheri Oudemans, 1915: 242 Zwickia guentheri (Oudemans): Oudemans, 1924: 310; Vitzthum, 1931: 123; Hughes & Jackson, 1958: 177 Type locality: Sri Lanka Distribution: Sri Lanka Host: in pitchers of Nepenthes distillatoria Zwickia nepenthesiana Hirst Zwickia nepenthesiana Hirst, 1928: 19; Vitzthum, 1931: 131 Type locality: Singapore Distribution: Singapore Host: from the pitchers of Nepenthes ampullaria Subfamily Myianoetinae Scheucher Myianoetini Scheucher, 1957: 337 Myianoetinae: Fain, 1974: 58 Type genus: Myianoetus Oudemans, 1929 Genus Myianoetus Oudemans Myianoetus Oudemans, 1929: 499 Anoetostoma Womersley, 1941: 485 [Type is oudemansi; syn. by Fain & Domrow 1980: 41] Type species: Acarus muscarum Linnaeus, 1758 Myianoetus africanus Mahunka Myianoetus africanus Mahunka, 1979d: 141 Type locality: Kenya Distribution: Kenya Myianoetus antipodus Fain et Galloway Myianoetus antipodus Fain & Galloway, 1993: 106; Clark, 2009: 31; 2010a: 9 Type locality: Motunau Island Distribution: New Zealand Host: Eudyptula minor albosignata (Aves: Spheniscidae) 90 Acarorum Catalogus IX Myianoetus assiuti Eraky, Abdel-Aziz et Abo-Elmaged Myianoetus assiuti Eraky, Abdel-Aziz & Abo-Elmaged, 2020: 78 Type locality: Assiut, Upper Egypt Distribution: Egypt Myianoetus brevis Fain Myianoetus brevis Fain, 1984: 287 Type locality: Rugege forest, 75 km from Butare, 2300 m Distribution: Rwanda Host: elephant dung Myianoetus clavatus Mahunka Myianoetus clavatus Mahunka, 1963d: 369; 1970: 51 Type locality: Hortobágy-Máta Distribution: Hungary Host: Spermophilus [Citellus] citellus (Rodentia: Sciuridae) Myianoetus copromyzae Fain, Britt et Molyneux Myianoetus copromyzae Fain, Britt & Molyneux, 1980: 402 Type locality: Scotland Distribution: Scotland Host: Copromyza sp. (Diptera: Sphaeroceridae) Myianoetus digitiferus (Trägårdh) Histiostoma digitifera Trägårdh, 1904: 67 Anoetus digitiferus (Trägårdh): Vitzthum, 1925: 155 Myianoetus digitiferus (Trägårdh): Willmann, 1937: 183 Type locality: Greenland Distribution: Greenland, Germany Host: Heleomyza [Blepharoptera] serrata (Diptera: Helomyzidae) Myianoetus dionychus (Oudemans) Anoetus dionychus Oudemans, 1910: 50 Myianoetus diadematus Willmann, 1937: 182 Myianoetus dionychus (Oudemans): Scheucher, 1957: 339 Type locality: Holland (Maastricht) Distribution: France, Germany, Holland, Hungary, Romania, Slovakia Hosts: flies (Diptera: Heleomyzidae); Plecotus auritus (Chiroptera: Vespertilionidae) Myianoetus domrowi (Fain) Anoetostoma domrowi Fain, 1968: 246 Acarorum Catalogus IX 91 Myianoetus domrowi (Fain) Type locality: Butemu Village, Finisterre Range, Madang Central District Distribution: Papua New Guinea Myianoetus dycei Fain et Domrow Myianoetus dycei Fain & Domrow, 1980: 41 Type locality: Queensland, Rockhampton Distribution: Australia Host: Culicoides brevitarsis (Diptera: Ceratopogonidae) Myianoetus filicum (Dujardin) Hypopus filicum Dujardin, 1841: 261 Myianoetus filicum (Dujardin): Type locality: France Distribution: France, Hungary Myianoetus granatum Eraky Myianoetus granatum Eraky, 2020: 153 Type locality: Assiut Distribution: Egypt Myianoetus heterotrichus Mahunka Myianoetus heterotrichus Mahunka, 1968: 252 Type locality: Hungary Distribution: Hungary Myianoetus kaszabi Mahunka Myianoetus kaszabi Mahunka, 1967c: 19 Type locality: Chentej aimak, 7 km NE of Somon Mörön Distribution: Mongolia Myianoetus lili Eraky Myianoetus lili Eraky, 1993: 47; Negm & Alatawi, 2011: 97; Abdelgayed et al., 2019: 32 Type locality: Assiut, upper Egypt Distribution: Egypt, Saudi Arabia Host: manure Myianoetus longisetosus Mašan et Krištofik Myianoetus longisetosus Mašan & Krištofik, 1992: 84 Type locality: Slovakia Distribution: Slovakia 92 Acarorum Catalogus IX Host: Hydrotaea sp. (Diptera: Muscidae) Myianoetus manurei Eraky, Abdel-Aziz et Abo-Elmaged Myianoetus manurei Eraky, Abdel-Aziz & Abo-Elmaged, 2020: 76 Type locality: Assiut, Upper Egypt Distribution: Egypt Myianoetus micromaculatus Fain et Philips Myianoetus micromaculatus Fain & Philips, 1979: 151 Type locality: Syracuse, N.Y. Distribution: USA Host: in the nest of Otus asio (Aves: Strigiformes) Myianoetus microti Sevastianov et Vysotskaya Myianoetus microti Sevastianov & Vysotskaya, 1971: 41 Type locality: Transcarpatian Distribution: Ukraine Host: in the nest of rodents Myianoetus mlcoch Samšiṅák Myianoetus mlcoch Samšiṅák, 1989: 94 Type locality: Pullman, Washington Distribution: USA Host: Copromyza (Borborillus) sordidus (Diptera: Sphaeroceridae) Myianoetus muscarum (Linnaeus) Acarus muscarum Linnaeus, 1758: 617 Myianoetus muscarum (Linnaeus): Buitendijk, 1945: 361; Scheucher, 1957: 338; Nuorteva, 1971; Sevastianov, 1975: 401; Oconnor et al., 2015: 1; Pimsler et al., 2016: 290 Type locality: Europe Distribution: Belgium, Germany, England, Holland, Finland, ?France, Italy, Canada, USA Hosts: Synthesiomyia nudiseta and other flies (Diptera: Muscidae), sometime hosts not sure (Pimsler et al., 2016). Myianoetus mystacis Mahunka Myianoetus mystacis Mahunka, 1972b: 363 Type locality: Vértes-Geb., Fánivölgy Distribution: Hungary Host: Fannia sp. (Diptera: Fanniidae) Myianoetus nerminka (Eraky et Shoker) Anoetostoma nerminka Eraky & Shoker, 1995: 21 Acarorum Catalogus IX 93 Myianoetus nerminka (Eraky et Shoker): Type locality: Egypt Distribution: Egypt Myianoetus oudemansi (Womersley) Anoetostoma oudemansi Womersley, 1941: 485; Hughes & Jackson, 1958: 126 Myianoetus oudemansi (Womersley): Fain & Domrow 1980: 41 Type locality: Sidney, NSW Distribution: Australia Host: Musca domestica (Diptera: Muscidae) Myianoetus ovatus Mašan et Kristofik Myianoetus ovatus Mašan & Kristofik, 1992: 81 Type locality: Slovakia Distribution: Slovakia Host: Hydrotaea sp. (Diptera: Muscidae) Myianoetus paganus Sevastianov Myianoetus paganus Sevastianov, 1973: 856 Type locality: Tchemerovtsi Village, Hmelnitskaya Obl., Ukraine Distribution: Ukraine Host: horse dung Myianoetus pappi Mahunka Myianoetus pappi Mahunka, 1972b: 361 Type locality: Aranyosgadány, Kuhfladen Distribution: Hungary Host: Azelia parva (Diptera: Muscidae) Myianoetus parvus Mahunka Myianoetus parvus Mahunka, 1969a: 360; 1972b: 363 Type locality: Finland Distribution: Finland, Hungary, Slovakia Hosts: Ophyra leucostoma (Diptera: Muscidae), Borboropsis fulviceps (Diptera: Heleomyzidae) Myianoetus pseudomuscarum Sevastianov Myianoetus pseudomuscarum Sevastianov, 1973: 856 Type locality: Tchemerovtsi Village, Hmelnitskaya Obl., Ukraine Distribution: Russia (Sakhalin, Tatarstan, Bashkortistan), Ukraine Host: cow’s dung 94 Acarorum Catalogus IX Myianoetus reductus Mahunka Myianoetus reductus Mahunka, 1969d: 368 Type locality: Mt Meru, eastern slope Distribution: Tanzania Myianoetus rohaceki Samšiṅák Myianoetus rohaceki Samšiṅák, 1984: 57; 1989: 91 Type locality: Hrubý Jesenik Mts., Skřitek Distribution: Czech Rep., Hungary, USA Hosts: Limosina rufilabris, Coproica vistripennis, Lotophila atra (Diptera: Sphaeroceridae) Myianoetus rwandanus Fain Myianoetus rwandanus Fain, 1984: 283 Type locality: Kamobuga, Rugege Forest, 2500 m Distribution: Rwanda Host: elephant dung Myianoetus scheucherae Mahunka Myianoetus scheucheri Mahunka, 1972: 371 (as nom.nov. for the deutonymphs recorded as “M. muscarum” by Scheucher, 1957: 338) Myianoetus scheucherae Mahunka: Klimov & Khaustov, 2018 Type locality: Erlangen Distribution: Germany Myianoetus simplex Mahunka Myianoetus simplex Mahunka, 1972: 363; Bongers, OConnor & Lukoschus, 1985: 19 Type locality: Aranyosgadány (Hungary) Distribution: Czech Rep., France, Holland, Hungary Hosts: Coproica lugubris (Diptera: Sphaeroceridae), Hebecnema umbratica (Diptera: Muscidae), Sepsis thoracica (Diptera: Sepsidae) Myianoetus simunek Samšiṅák Myianoetus simunek Samšiṅák, 1989: 91 Type locality: Kiskunsagi N. P. Kunfehérto (lake) Distribution: Hungary Host: Coproica lugubris (Diptera: Sphaeroceridae) Myianoetus sovat Zamec Myianoetus sovat Zamec, 2014: 916 Type locality: Vlčie Diery Cave Distribution: Slovakia Host: phoretic on Gymnomus caesius (Diptera: Heleomyzidae) Acarorum Catalogus IX 95 Myianoetus szaboi Mahunka Myianoetus szaboi Mahunka, 1978: 109 Type locality: Kócs (Hortobágy Nat. Park) Distribution: Hungary Myianoetus szunyoghyi Mahunka Myianoetus szunyoghyi Mahunka, 1969d: 369 Type locality: Mt Meru, eastern slope Distribution: Tanzania Myianoetus thaleri Mahunka Myianoetus thaleri Mahunka, 1972: 374 Type locality: Unterengadin, Ramosch, Charbunera Distribution: Austria Myianoetus travei Fain Myianoetus travei Fain, 1976: 309 Type locality: Kerguelen Island Distribution: Kerguelen Island Myianoetus tuerkorum Scheucher Myianoetus tuerkorum Scheucher, 1957: 341; Mahunka, 1970: 48; Fain et al., 1993: 14 Type locality: Erlangen (Germany) Distribution: Belgium, Germany, Slovakia Host: bird’s nests Myianoetus tuomikoskii Mahunka Myianoetus tuomikoskii Mahunka, 1969: 361 Type locality: Finland Distribution: Finland, Slovakia, Romania, Mongolia Hosts: Hydrotaea occulta, H. tuberculata (Diptera: Muscidae) Myianoetus uncinatus Fain Myianoetus uncinatus Fain, 1984: 285 Type locality: Kamobuga, Rugege Forest, 2500 m Distribution: Rwanda Host: elephant dung Myianoetus undulatus Hughes et Jackson Myianoetus undulatus Hughes & Jackson, 1958: 158 Type locality: Chickahominy Swamp, Ellerson, Virginia Distribution: USA 96 Acarorum Catalogus IX Myianoetus vesparum Scheucher Myianoetus vesparum Scheucher, 1957: 343 Type locality: Erlangen Distribution: Germany Host: nests of Vespidae (Hymenoptera) Myianoetus virgatus Scheucher Myianoetus virgatus Scheucher, 1957: 345; Mahunka, 1970: 49 Type locality: Germany Distribution: Czech Rep., Germany, Slovakia Hosts: Diptera (Borboridae, Scatophagidae), Limosina clunipes (Diptera: Sphaeroceridae) Myianoetus zdena Samšiṅák Myianoetus zdena Samšiṅák, 1984: 60; 1989: 91 Type locality: Kremnicke pohoria, Mts Tuček, 850 m Distribution: Austria, Hungary, Slovakia Hosts: Elachisoma aterrima, Coproica lugubris, C. pallifrons, C. vagans (Diptera: Sphaeroceridae) Subfam. Scolianoetinae Fain Scolianoetinae Fain, 1974: 60 Type genus: Scolianoetus Fain, 1974 Genus Scolianoetus Fain Scolianoetus Fain, 1974: 60 Type species: Scolianoetus scheucherae Fain, 1974 Scolianoetus brasiliensis Fain Scolianoetus brasiliensis Fain, 1974: 62 Type locality: Novo Friburgo Distribution: Brazil Host: Spartocera cinnamomea (Heteroptera: Coreidae) Scolianoetus faini Mahunka Scolianoetus faini Mahunka, 1975a: 321 Type locality: Kiunga Distribution: Papua New Guinea Scolianoetus javanus Mahunka Scolianoetus javanus Mahunka, 1976b: 319 Type locality: Java, Pangerango Acarorum Catalogus IX 97 Distribution: Indonesia (Java) Host: Pelopides tridens (Lepidoptera: Hesperiidae) Scolianoetus ramozae Fain et Camerik Scolianoetus ramozae Fain & Camerik, 1978: 169 Type locality: Rio de Janeiro Distribution: Brazil Host: Passalus sp. (Coleoptera: Passalidae) Scolianoetus scheucherae Fain Scolianoetus scheucherae Fain, 1974: 62 Type locality: Lisala Distribution: DR Congo Host: Campsomeris mansueta (Hymenoptera: Scoliidae) Subfam. Spinanoetinae Scheucher Spinanoetini Scheucher, 1957: 357 Spinanoetinae: Fain, 1974: 58 Type genus: Spinanoetus Scheucher, 1957 Genus Spinanoetus Scheucher Spinanoetus Scheucher, 1957: 357 Type species: Spinanoetus pelznerae Scheucher, 1957 Spinanoetus heterotrichus Mahunka Spinanoetus heterotrichus Mahunka, 1962: 432; Kaluz, 1992: Type locality: Dunazuggebirge, Sikáros Distribution: Hungary, Slovakia Spinanoetus pelznerae Scheucher Spinanoetus pelznerae Scheucher, 1957: 358; Káluz, 1993; Haitlinger, 2004: Type locality: Erlangen (Germany) Distribution: Germany, Hungary, Poland Hosts: Nicrophorus interruptus, Oeceoptoma thoracica, Thanatophilus rugosus, Th. sinuatus (Coleoptera: Silphidae) Spinanoetus variabilis Mahunka Spinanoetus variabilis Mahunka, 1973a: 109 Type locality: Nakpanduri Distribution: Ghana 98 Acarorum Catalogus IX Spinanoetus weingaertnerae Scheucher Spinanoetus weingaertnerae Scheucher, 1957: 360; Haitlinger, 2004; Zamec, 2014 Type locality: Erlangen (Germany) Distribution: Germany, Hungary, Poland, Slovakia Hosts: Emus hirtus, Ontolestes sp., Philontus sp. (Coleoptera: Staphylinidae) Index of Histiostomatidae (Names in bold are accepted as valid) abietis, Histiostoma 39 abietis, Anoetus 39 acanthotarsus, Peripatetes 77 acrocini, Histiostoma 39 adarcuale, Histiostoma 39 adsimile, Histiostoma 40 aelleni, Anoetus 23 aeschlimanni, Hormosianoetus 73 africana, Bonomoia 36 africana, Probonomoia 77 africanus, Anoetoglyphus 21 africanus, Glyphanoetus 18 africanus, Myianoetus 89 afropulicarius, Psyllanoetus 80 afrotropicalis, Xenanoetus 88 akodoni, Anoetus 23 alicola, Anoetus 23 alicola, Hypopus 23 alii, Histiostoma 40 alphaticum, Histiostoma 40 amargynus, Histiostoma 40 Ameranoetus 20 americanum, Histiostoma 40 amieti, Histiostoma 40 Amyzanoetus 20 Ancyranoetus 21 andrei, Histiostoma 40 angolensis, Histiostoma 40 anguillarum, Histiostoma 72 angularis, Anoetus 24 Anoetidae 14 Anoetinae 20 Anoetoglyphus 21 anoetopsis, Anoetus 24 Anoetostoma 89 Anoetus 23 antennatus, Anoetoglyphus 21 antillense, Histiostoma 41 antipodus, Myianoetus 89 Aphodanoetus 35 arcuatus, Histiostoma 41 arcuata, Histiostoma 41 arenapus, Histiostoma 41 asper, Capronomoia 15 asper, Copronomoia 16 assiuti, Myianoetus 90 ateuchi, Anoetoglyphus 21 Auricanoetus 36 australiense, Histiostoma 41 australis, Kanoetus 75 Austranoetus 36 bacilligerum, Histiostoma 41 bakeri, Histiostoma 41 baloghi, Histiostoma 41 baloghi, Synanoetus 86 bananai, Histiostoma 42 banjuwangicus, Anoetus 41 banjuwangicus, Histiostoma 41 banksi, Semianoetus 85 bassoni, Loxanoetus 75 Acarorum Catalogus IX 99 bellulus, Capronomoia 16 bellulus, Copronomoia 16 berghi, Histiostoma 42 berghii, Histiostoma 42 berndhauseri, Histiostoma 42 bernoni, Histiostoma 42 bifastigatus, Anoetoglyphus 21 bifoveolatum, Histiostoma 42 bifurcatus, Rhopalanoetus 82 blomquisti, Histiostoma 42 Bonomoia 36 Bothyanoetus 38 brachypodus, Insulanoetus 74 bradymeri, Anoetus 24 brasiliensis, Scolianoetus 96 brevimanum, Histiostoma 42 brevimana, Lipstorpia 42 brevimanus, Anoetus 43 brevimanus, Histiostoma 42 brevipes, Histiostoma (nomen dubium) 43 brevipes, Traskorchestianoetus 87 brevipodum, Histiostoma 43 brevis, Myianoetus 90 brevisetus, Glyphanoetus 18 bushlandi, Anoetus 24 cacuminatus, Anoetus 24 calicifera, Histiostoma 43 caljubiensis, Anoetus 24 camerunensis, Histiostoma 43 campanula, Anoetus 43 campanula, Histiostoma 43 campanulae, Histiostoma 43 camphori, Histiostoma 43 capillatum, Histiostoma 43 capillata, Histiostoma 43 capillatus, Anoetus 24 Capronomoia 15 caro, Histiostoma 44 castrii, Anoetus 24 cataglyphi, Histiostoma 44 Cavanoetus 72 Cederhjelmia 38 certa, Bonomoia 36 ceylanensis, Anoetus 25 Ceylanoetinae 14 Ceylanoetus 14 chalinolobus, Chiropteranoetus 15 chilensis, Anoetus 25 Chiloanoetus 38 chilopophila, Histiostoma 44 chinensis, Rhopalanoetus Chiropteranoetus 15 chrysis, Probonomoia 77 cirratus, Anoetus 44 cirratum, Histiostoma 44 cirrosus, Anoetus 25 clavatum, Histiostoma 44 clavatus, Myianoetus 90 clavifer, Histiostoma 44 collis, Anoetus 25 colocasiae, Anoetus 89 colocasia, Zwickia 89 colocasiae, Zwickia 89 columbiana, Probonomoia 78 Comyianoetus 15 conclavicola, Anoetus 44 conclavicola, Histiostoma 44 Conglanoetus 15 congoensis, Anoetus 25 congoensis, Bonomoia 78 congoensis, Probonomoia 78 conjuncta, Anoetus 45 conjuncta, Histiostoma 45 conjunctum, Histiostoma 45 conoidalis, Hexanoetus 39 continuatum, Histiostoma 45 contrasetosus, Kanoetus 75 coprae, Histiostoma 45 copridis, Anoetoglyphus 22 copromyzae, Myianoetus 90 coprophilus, Anoetus 25 coronatum, Histiostoma 45 100 Acarorum Catalogus IX cossi, Histiostoma 45 crantor, Histiostoma 45 crassipes, Anoetus 45 crassipes, Histiostoma 45 Creutzeria 16 crypturgi, Anoetus 46 crypturgi, Histiostoma 46 cubanus, Kanoetus 75 cubanum, Histiostoma 46 Curculanoetus 17 cyrtandrae, Anoetus 46 cyrtandrae, Histiostoma 46 dactyloscopicum, Histiostoma 46 darlingtoniae, Sarraceniopus 84 darwishii, Histiostoma 46 debilis, Anoetus 25 decemvirgae, Histiostoma 46 decorus, Anoetus 25 demel, Richardanoetus 83 densus, Histiostoma 47 denticulatus, Comyianoetus 15 denticulatus, Conglanoetus 15 diademata, Histiostoma 47 diadematum, Histiostoma 47 diadematus Myianoetus 90 digitifera, Histiostoma 90 digitiferus, Anoetus 90 digitiferus, Myianoetus 90 dinopsyllus, Psylanoetus 80 dionychus, Anoetus 90 dionychus, Myianoetus 90 discrepans, Anoetus 23 disiunctus, Anoetus 47 disiuncta, Histiostoma 47 disiunctum, Histiostoma 47 distentoseta, Histiostoma 47 distentosetum, Histiostoma 47 dolifer, Anoetus 26 domrowi, Anoetostoma 26 domrowi, Anoetus 26 domrowi, Myianoetus 90 dryocoeti, Anoetus 47 dryocoeti, Histiostoma 47 dudichi, Anoetus 26 dugesi, Histiostoma 47 dugesii, Hypopus 47 duplex, Histiostoma 47 dycei, Myianoetus 91 eboris, Glyphanoetus 18 edax, Histiostoma 48 edita, Histiostoma 48 editum, Histiostoma 48 egyptiacum, Histiostoma 48 eickworti, Histiostoma 48 ellipticus, Anoetoglyphus 22 elzingai, Histiostoma 48 endroedii, Anoetus 48 endroedii, Histiostoma 48 endroedyyoungai, Anoetus 26 endroedyyoungai, Rhopalanoetus 82 ensifer, Anoetus 48 ensifer, Histiostoma 48 erugo, Histiostoma 48 excavatus, Ceylanoetus 14 exornatus, Anoetus 26 faini, Scolianoetus 96 faini, Syringanoetus 87 farghali, Histiostoma 49 feroniarum, Anoetus 49 feroniarum, Histiostoma 49 feroniarum, Hypopus 49 fetters, Anoetus 26 Fibulanoetinae 17 Fibulanoetus 17 fibulatus, Anoetus 26 filicum, Hypopus 91 filicum, Myianoetus 91 fimetarium, Histiostoma 82 fimetarius, Rhopalanoetus 82 flaccus, Histiostoma 49 flagellifer, Histiostoma 49 flagellatus, Glyphanoetus 18 Acarorum Catalogus IX 101 flabelliforme, Histiostoma 49 flagellifer, Histiostoma 49 formicarum, Histiostoma 50 formosana, Histiostoma 50 fortis, Histiostoma 50 foveosimile, Histiostoma 50 fragosus, Histiostoma 50 fucatifer, Histiostoma 50 fuliginosi, Histiostoma 50 fulmeki, Glyphanoetus 19 furcata, Histiostoma 50 furcatus, Rhopalanoetus 82 galerus, Histiostoma 51 gambiense, Histiostoma 51 gampsoceri, Histiostoma 51 genavensium, Histiostoma 51 genetica, Histiostoma 74 gervaisi, Histiostoma 51 ghanensis, Anoetus 26 Ghanoetus 18 gibsoni, Anoetus 84 gibsoni, Sarraceniopus 84 gibsoni, Zwickia 84 gladifer, Rhopalanoetus 82 gladiger, Anoetus 51 gladiger, Histiostoma 51 globotuberum, Histiostoma 51 globulus, Anoetoglyphus 22 Glyphanoetinae 14 Glyphanoetus 18 gordius, Anoetus 52 gordius, Histiostoma 52 gorgo, Histiostoma 52 gozmanyi, Rhaphidothrix 81 gracilipes, Histiostoma 52 gracilis, Histiostoma 52 gradatus, Capronomoia 16 gradatus, Copronomoia 16 graeca, Hauserania 38 granatum, Myianoetus 91 grandiceps, Xenanoetus 88 granulatum, Histiostoma 52 grapefruit, Anoetus 27 guanophilum, Histiostoma 52 guentheri, Anoetus 89 guentheri, Zwickia 89 halicticola, Histiostoma 52 halicticolum, Histiostoma 52 halictonida, Anoetus 27 halictonida, Histiostoma 27 halophilus, Amyzanoetus 21 harteni, Stercoranoetus 86 Hauserania 38 hauseri, Anoetus 27 heinemanni, Histiostoma 53 helena, Anoetus 27 heliocopridis, Anoetus 27 helobius, Acarus 27 helobius, Anoetus 27 hematobii, Ameranoetus 20 herbali, Histiostoma 53 heriades, Histiostoma 53 heterotrichus, Myianoetus 91 heterotrichus, Spinanoetus 85 Hexanoetus 39 himalayae, Anoetus 53 himalayae, Histiostoma 53 hirsutus, Rhopalanoetus 83 histeri, Histiostoma 53 Histiostoma 39 Histiostomatidae 14 Histiostomatinae 20 Histiostomatini 20 Histiostomatinae 20 Histiostomatoidea 11 Hormosianoetus 73 horridum, Histiostoma 53 horridus, Anoetus 27 hospitus, Anoetus 28 huettleri, Histiostoma 54 hughesi, Anoetus 84 hughesi, Sarraceniopus 84 102 Acarorum Catalogus IX humiditatis, Anoetus 54 humiditatis, Histiostoma 54 humilis, Anoetus 28 humprechti, Bonomoia 78 humprechti, Probonomoia 78 hungarica, Histiostoma 54 Hymenanoetus 74 Ichtanoetus 72 imbecillus, Histiostoma 54 immaturus, Anoetoglyphus 22 impressum, Histiostoma 28 impressus, Anoetus 28 incanus, Glyphanoetus 18 incertus, Pteranoetus 80 incurvata, Histiostoma 54 incurvatum, Histiostoma 54 indetonsum, Histiostoma 54 indetonsus, Anoetus 54 indica, Histiostoma 28 indicus, Anoetus 28 inquilinum, Histiostoma 54 insolita, Anoetus 28 insulare, Histiostoma 55 insularis, Anoetus 55 Insulanoetus 74 ionicum, Histiostoma 55 irinum, Histiostoma 55 irinus, Anoetus 55 janosi, Histiostoma 55 javanus, Scolianoetus 96 julorum, Anoetus 52 julorum, Histiostoma 55 julorum, Hypopus 55 junctus, Histiostoma 55 kabrt, Xenanoetus 88 Kanoetus 75 karachiensis, Histiostoma 56 Kaszabanoetinae 14 Kaszabanoetus 75 kaszabi, Anoetus 29 kaszabi, Myianoetus 91 kaszabi, Pteranoetus 81 kerguelenense, Histiostoma 56 kerguelenensis, Austranoetus 36 kofrankova, Xenanoetus 88 komarpictoris, Anoetus 29 labiatus, Fibulanoetus 17 laboratorium, Histiostoma 74 laboratorium, Hormosianoetus 74 laevis, Histiostoma 56 laevis, Zschachia (nomen dubium) 56 lanceocrinus, Anoetus 83 lanceocrinus, Rhopalanoetus 83 langi, Anoetoglyphus 22 latus, Anoetoglyphus 22 latus, Pteranoetus 81 lekiensis, Histiostoma 56 lenae, Loxanoetus 76 leptos, Glyphanoetus 19 levis, Anoetoglyphus 22 lili, Myianoetus 91 lineata, Teinokyra 87 lineatula, Histiostoma 56 lineosculpturatum, Histiostoma 56 Lipstorpia litorale, Histiostoma 56 litoralis, Anoetus 56 loebli, Histiostoma 57 loksai, Histiostoma 57 longipes, Anoetus 29 longipes, Histiostoma 29 longisetosus, Myianoetus 91 longisetum, Histiostoma 57 longisetus, Anoetus 29 longispina, Glyphanoetus longitarsus, Fibulanoetus 17 longitarsus, Auricanoetus 36 longitarsus, Loxanoetus (A.) 36 longitubus, Anoetus 29 longus, Glyphanoetus 19 lootsi, Anoetoglyphus 22 lorentzi, Anoetus 57 Acarorum Catalogus IX 103 lorentzi, Histiostoma 57 louisianae, Histiostoma 57 Loxanoetus 75 lunulatus, Anoetus 29 luxus, Histiostoma 57 machadoi, Anoetus 29 machadoi, Porrhanoetus 77 macrosuctus, Anoetus 30 magdolnae, Histiostoma 57 mahunkai, Capronomoia 16 mahunkai, Fibulanoetus 17 mahunkai, Glyphanoetus 19 mahunkai, Histiostoma 57 mahunkai, Hormosianoetus 74 mallotae, Hormosianoetus 74 mallotae, Mauduytia 74 manicae, Histiostoma 58 mannai, Histiostoma 58 mantelli, Histiostoma 58 manurei, Myuanoetus 92 manuri, Histiostoma 58 marani, Histiostoma ovalis 61 margaritata, Histiostoma 58 margaritatum, Histiostoma 58 marianae, Histiostoma 58 maritimus, Anoetus 58 maritimum, Histiostoma 58 marmoreus, Ghanoetus 18 matsakisi, Histiostoma 59 Mauduytia 76 mayumbensis, Chiloanoetus 38 media, Anoetus 59 medium, Histiostoma 59 melesicola, Psyllanoetus 80 melolonthae, Histiostoma 59 meszarosorum, Histiostoma 59 metwallii, Histiostoma 59 micidus, Glyphanoetus 19 micromaculatus, Myianoetus 92 microti, Anoetus 30 microti, Myianoetus 92 minimum, Histiostoma 59 mirabile, Histiostoma 59 mlkoch, Myianoetus 92 mongolica, Scheucheria 84 montanum, Histiostoma 60 moralesi, Histiostoma 60 moshtohorensis, Anoetus 30 mrcziaki, Semianoetus 85 Munduytia 76 murchiei, Histiostoma 60 muscae, Histiostoma 60 muscarum, Acarus 92 muscarum, Myianoetus 92 Myianoetinae 89 Myianoetini 89 Myianoetus 89 myrmicarum, Anoetus 30 myrmicarum, Histiostoma 30 mystacis, Myianoetus 92 nasseri, Histiostoma 60 necopinus, Anoetus 60 necopinus, Histiostoma 60 necrophori, Anoetus necrophori, Histiostoma necrophori, Hypopus neglectus, Anoetus 30 negmi, Histiostoma 60 Nepenthacarus 76 nepenthesiana, Anoetus 89 nepenthesiana, Zwickia 89 nerminka, Anoetostoma 92 nerminka, Myianoetus 92 nichollsi, Histiostoma 62 nigrellii, Histiostoma 73 Nodipalpidae 14 Nodipalpus 23 nodosus, Hymenanoetus 74 nomiensis, Glyphanoetus 19 notabilis, Anoetus 30 novamsterdamense, Histiostoma 61 ocellatum, Histiostoma 73 104 Acarorum Catalogus IX onioni, Histiostoma 61 onthophagae, Histiostoma 61 opertus, Anoetus 30 opuntiae, Bonomoia 78 opuntiae, Probonomoia 78 orientalis, Histiostoma 61 ornata, Hauserania 39 ornatissimus, Anoetus 31 ornatofoveolatus, Conglanoetus 15 ornatus, Anoetus 31 Otanoetus 76 oudemansi, Anoetostoma 93 oudemansi, Anoetus 61 oudemansi, Histiostoma 61 oudemansi, Myianoetus 93 ovalis, Hypopus 51 ovale, Histiostoma 61 ovalis, Histiostoma 61 ovalis, Uropoda 61 Ovanoetus 77 ovatus, Myianoetus 93 oxfordianum, Histiostoma paganus, Myianoetus 93 palustre, Histiostoma 62 pango, Capronomoia 16 pango, Copronomoia 16 pannonicum, Histiostoma pannonicus, Anoetus 31 papillata, Histiostoma 62 papillatum, Histiostoma 62 pappi, Glyphanoetus 19 pappi, Myianoetus 93 parmatus, Anoetus 31 parvus, Myianoetus 93 pauciputeolum, Histiostoma 62 pauliani, Anoetus 31 pectineum, Histiostoma 49 pelznerae, Spinanoetus 85 pereirai, Bothyanoetus 38 Peripatetes 77 permixtus, Scutanoetus 84 perruptum, Histiostoma 62 phyllophorum, Histiostoma 62 phyllophorus, Anoetus 62 phyllophorus, Histiostoma 62 phyllotrichus, Anoetus 19, 52 phyllotrichus, Glyphanoetus 19 phyllotrichus, Hypopus 19 phytophaga, Histiostoma 62 piceae, Anoetus 63 piceae, Histiostoma 63 pickaxei, Histiostoma 63 picturata, Bonomoia 37 pierrestrinatii, Histiostoma (Cavanoetus) 72 piloseta, Histiostoma 63 pini, Anoetus 31 pini, Bonomoia 78 pini, Histiostoma 31 pini, Probonomoia 78 piscium, Histiostoma 73 pocsi, Anoetus 31 polaki, Anoetus 63 polaki, Histiostoma 63 polygonatus, Anoetoglyphus 23 polynesiacus, Anoetus 32 polypori, Anoetus 63 polypori, Histiostoma 63 pontigerum, Histiostoma 63 porosus, Anoetus 32 Porrhanoetus 77 primitiva, Bonomoia 37 privorai, Anoetus 32 probata, Bonomoia 37 Probonomoia 77 processum, Glyphanoetus 20 prodectoris, Anoetus 63 prodectoris, Histiostoma 63 prophthalmi, Anoetus 64 prophthalmi, Histiostoma 64 protuberans, Histiostoma 64 Prowichmannia 79 Acarorum Catalogus IX 105 pseudolongipes, Histiostoma 64 pseudomuscarum, Myianoetus 93 Psyllanoetus 80 Pteranoetus 80 pulchrum, Histiostoma 85 pulchrum, Sellea 85 punctaticeps, Histiostoma 64 punctolineatus, Rhaphidothrix 81 punctulatus, Anoetus 32 puteanus, Comyianoetus 15 puteanus, Conglanoetus 15 pyriforme, Histiostoma 64 quadriuncinata, Cederhjelmia 38 radiferum, Histiostoma 64 rakha, Histiostoma 65 ramozae, Scolianoetus 97 recondita, Bonomoia 37 reductus, Myianoetus 94 reticulatissimum, Histiostoma 65 reticulatus, Anoetus 32 reticulata, Bonomoia 37 reticulofoveolatum, Histiostoma 65 retractus, Anoetus 32 rettenmeyeri, Pteranoetus 81 rettenmeyerorum, Histiostoma 65 Rhaphidothrix 81 Rhopalanoetus 81 rhynchophorus, Curculanoetus 17 Richardanoetus 83 rimosa, Histiostoma 65 rimosum, Histiostoma 65 rohaceki, Myianoetus 94 rizkii, Histiostoma 65 rostrata, Histiostoma 65 rostratum, Histiostoma 65 rostro-serratum, Histiostoma 49 rostroserratum, Histiostoma 49 rostro-serratus, Histiostoma 49 rostro-serratus, Rhizoglyphus 49 rostro-serratus, Tyroglyphus 49 ruehmi, Anoetus 66 ruehmi, Histiostoma 66 rugosa, Bonomoia 37 rugosum, Histiostoma 66 rwandanus, Myianoetus 94 sachsi, Anoetus 66 sachsi, Histiostoma 66 samiensis, Anoetus 32 sammari, Histiostoma samsinaki, Kanoetus 75 sapromyzarum, Anoetus 66 sapromyzarum, Histiostoma 66 sapromyzarum, Hypopus 66 Sarraceniopus 84 sarrai, Histiostoma 66 schauenbergi, Histiostoma 66 scheucherae, Myianoetus 94 scheucherae, Scolianoetus 97 scheucheri, Myianoetus 94 Scheucheria 84 schoutedeni, Histiostoma 67 scindita, Histiostoma 67 scinditum, Histiostoma 67 Scolianoetinae 96 Scolianoetus 96 Scutanoetus 84 Sellea 85 Semianoetus 85 serrata, Histiostoma 32 serratus, Anoetus 32 setipes, Anoetus 94 setipes, Hypopus 94 setipes, Myianoetus 94 sextoni, Histiostoma 67 seychellensis, Creutzeria 17 shamyi, Anoetus 33 shanensis, Anoetus 33 sharae, Anoetus 33 shiramba, Histiostoma 67 sibirica, Bonomoia 79 sibirica, Probonomoia 79 simile, Histiostoma 67 106 Acarorum Catalogus IX simplex, Myianoetus 94 simplex, Rhaphidothrix 81 simplex, Rhopalanoetus 83 simunek, Myianoetus 94 sinuosus, Anoetus 33 siphunculinae, Histiostoma 67 solimani, Histiostoma 67 solomoniensis, Anoetus 33 sordida, Anoetus 33 sovat, Myianoetus 94 sphaerocerae, Bonomoia 16 sphaerocerae, Capronomoia 16 spiceri, Traskorchestianoetus 87 Spinanoetinae 97 Spinanoetini 97 Spinanoetus 85 spinifera, Bonomoia 79 spinifera, Probonomoia 79 spinifera, Prowichmannia 79 spinifera, Wichmannia 79 spinifera, Histiostoma 79 spiniferum, Prowichmannia 79 spiniferus, Anoetus 79 spiniferus, Wichmannia 79 spinipes, Anoetus 33 spinitarsus, Anoetus 68 spinitarsus, Histiostoma 67 spinosus, Anoetus 34 stammeri, Anoetus 68 stammeri, Histiostoma 68 stammeri, Wichmannia 79 stammeri, Prowichmannia 79 Stercoranoetus 86 strenzkei, Histiostoma 68 striata, Histiostoma 34 striatosimile, Histiostoma 68 striatus, Anoetus 34 subsimile, Histiostoma 68 sumatrensis, Anoetus 34 sylvestris, Anoetus 68 sylvestre, Histiostoma 68 Synanoetus 86 Syringanoetus 86 szaboi, Myianoetus 95 szelenyii, Anoetus 68 szelenyii, Histiostoma 68 szentivanyi, Histiostoma 69 szunyoghyi, Anoetoglyphus 23 szunyoghyi, Glyphanoetus 20 szunyoghyi, Myianoetus 95 talpae, Wichmannia 79 talpae, Prowichmannia 79 tanypus, Histiostoma 69 tarsalis, Histiostoma (nomen dubium) 69 tauffliebi, Anoetus 34 Teinokyra 87 teinophallus, Aphodanoetus 36 telatum, Histiostoma 69 Tensiostoma 87 tesselata, Histiostoma 69 thaleri, Myianoetus 95 tibetana, Myianoetus (nomen dubium) tienhoveni, Anoetus 82 tienhoveni, Histiostoma 82 timidum, Histiostoma 69 tinydorsalis, Histiostoma 69 titanolabi, Histiostoma 69 tobaica, Creutzeria (nomen dubium) 17 topali, Anoetus 34 toxici, Anoetus 34 toxopei, Anoetus 70 toxopei, Histiostoma 70 Traskorchestianoetus 87 travei, Myianoetus 95 trichophorum, Histiostoma 70 trichophorus, Anoetus 70 trichophorus, Histiostoma 70 trogicola, Histiostoma 83 trogicola, Rhopalanoetus 83 tropicus, Anoetus 76 tropicus, Munduytia 76 trossis, Capronomoia 16 Acarorum Catalogus IX 107 tuerkorum, Myianoetus 95 tunesiensis, Anoetoglyphus 23 tuomikoskii, Myianoetus 95 turcastanae, Anoetus 34 ulmi, Anoetus 70 ulmi, Histiostoma 70 ulomae, Kaszabanoetus 75 umbrosa, Histiostoma 70 uncinatus, Myianoetus 95 undulatus, Myianoetus 95 unidentatum, Histiostoma 70 urcerosus, Anoetus 35 uvafer, Histiostoma 70 uxoria, Histiostoma 71 valida, Histiostoma 71 validum, Histiostoma 71 vandenberghi, Syringanoetus 87 varia, Anoetus 71 varia, Histiostoma 71 varium, Histiostoma 71 variabilis, Histiostoma 71 variabilis, Spinanoetus 86 variolatus, Anoetus 35 veliaphilum, Tensistoma 87 venustissimus, Anoetus 20 venustissimus, Glyphanoetus 20 verruca, Histiostoma 71 verrucogenicula, Histiostoma 71 vesparum, Myianoetus 96 vestigialis, Xenanoetus 88 vilhenaorum, Anoetus 35 virgatus, Myianoetus 96 virginiana, Anoetus 35 virginiana, Histiostoma 71 virginianum, Histiostoma 71 virkkii, Ancyranoetus 21 vitzthumi, Anoetus 35 vitzthumi, Histiostoma 35 volgini, Histiostoma 71 vulpis, Hypopus 72 vulpis, Histiostoma 72 warreni, Nepenthacarus 76 wastis, Anoetus 35 watanabei, Histiostoma 72 wauense, Histiostoma 72 wautersi, Ovanoetus 77 weingaertnerae, Spinanoetus 86 wetzeli, Otanoetus 77 Wichmannia 79 (praeoccup.) wooleyi, Histiostoma 86 woolleyi, Stercoranoetus 86 Xenanoetus 88 zaheri, Histiostoma 72 zdena, Myianoetus 96 zoheirii, Bonomoia 37 Zschachia 39 Zwickia 88 108 Acarorum Catalogus IX Distribution of Histiostomatidae EUROPE Austria (Vitzthum, 1926a, 1927; Scheucher, 1957; Mahunka, 1972b; Samšinák, 1958, 1962, 1989) – Anoetoglyphus copridis, Histiostoma feroniarum, H. gladiger, H. gordius, H. humiditatis, H. julorum, H. vitzthumi, Myianoetus thaleri, M. zdena Belgium (Cooreman, 1963; Fain et al., 1993) – Anoetus pini, Conglanoetus denticulatus, C. puteanus, Myianoetus tuercorum, Histiostoma gladiger, H. gordius, H. trogicola Bulgaria (Samšinák, 1958; Beron, 1973; Takov, Pilarska & Moser, 2009) – Histiostoma ovale (= gladiger), H. piceae, Prowichmannia spiniferum Croatia (Pernek et al. 2008; Pernek et al. 2012; Wirth & Pernek. 2012; Wirth et al., 2016) – Histiostoma piceae, H. ulmi Czech Rep. (Müller, 1860; Samšinák, 1956, 1958, 1984, 1989) – Capronomoia sphaerocerae, Histiostoma manica, H. ovale, Myianoetus simplex, M. virgatus, Probonomoia humprechti Denmark (Jensen, 1895; Haarløv, 1957) – Histiostoma berghii, H. julorum Greenland (Trägårdh, 1905) – Myianoetus digitiferus Finland (Mahunka, 1969, 1972b; Nuorteva, 1971; Penttinen et al., 2013) – Histiostoma sp., Myianoetus muscarum, M. parvus, M. tuomikoskii, Probonomoia pini, Xenanoetus vestigialis France (contin.) (Dufour, 1839; Gervais, 1844; Theodorides¸1955; Cooreman, 1954 ; Samšinák, 1958; Levieux et al., 1989; Fain & Beaucournu, 1973; Mahunka, 1972b, 1977) – Anoetus dolifer, Histiostoma feroniarum, H. gervaisi, H. gladiger, H. piceae, H. sapromysarum, Myianoetus dionychus [= diadematus], M. simplex, Psyllanoetus melesicola Corsica (Samšinák, 1958; Wirth, 2005) - Histiostoma ovale (= gladiger), Probonomoia opuntiae Germany (Koch, 1843; Kramer, 1876, 1886; Oudemans, 1902; Willmann, 1937; Scheucher, 1957; Fain & Erteld, 1998; Wirth, 2003) – Anoetus myrmicarum, neglectus, A. pini, Capronomoia sphaerocerae, Glyphanoetus phyllotrichus, G. venustissimus, Histiostoma crypturgi, H. gladiger, H. gordius, H. halicticolum, H. julorum, H. oudemansi, H. palustre, H. piceae, H. phyllophorum, H. polypori, H. sapromyzarum, H. stammeri, H. strenzkei, H. trichophorum, H. ulmi, H. vitzthumi, Myianoetus dionychus [= diadematus], Probonomoia pini, Prowichmannia spiniferum, Rhopalanoetus lanceocrinus, Sellea pulchrum, Spinanoetus pelznerae, S. weingaertnerae Great Britain (Michael, 1901; Fain, Britt & Molyneux, 1980; Behura, 1950; Hyatt, 1990) – Histiostoma polypori, Myianoetus copromyzae, Prowichmannia spiniferum, Selea pulchrum Greece (Mahunka, 1972e, 1977) – Anoetus hauseri, A. samiensis, Histiostoma feroniarum, H. sapromyzarum, H. ionicum, H. matsakisi Acarorum Catalogus IX 109 Kephallinia (Mahunka, 1971, 1972e) – Anoetus hauseri, A. lunulatus, A. sapromyzarum, A. samiensis, Hauserania graeca, H. ornata, Histiostoma feroniarum, H. insulare Zakynthos (Mahunka, 1972e) - Histiostoma feroniarum, Levkas (Mahunka, 1972e) - Histiostoma sapromyzarum, Holland (Oudemans, 1914; Bongers, OConnor & Lukoschus, 1985; Fain & Lambrechts, 1987) – Aphodanoetus teinophallus, Histiostoma brevimanum, H. insulare, H. ocellatum, H. phyllophorum, H. polypori, H. sapromyzarum, Myianoetus simplex, Rhopalanoetus lanceocrinus Hungary (Mahunka, 1961, 1962a, 1962b, 1963d, 1968, 1970, 1972b, 1976, 1977b, 1978; Mahunka & Eraky, 1987; Samšinák, 1989) – Anoetoglyphus copridis, Anoetus dudichi, A. myrmicarum, A. pannonicus, A. pini, Capronomoia sphaerocerae, Glyphanoetus longispina, Histiostoma baloghi, H. dactyloscopicum, H. disiunctum, H. gordius, H. histeri, H. hungaricum, H. lineatula, H. loksai, H. magdolnae, H. margaritatum, H. ovalis, H. polypori, H. sapromyzarum, H. serrata, Myianoetus clavatus, M. dionychus (= diadematus), M. heterotrichus, M. mystacis, M. pappi, M. parvus, M. rohaceki, M. simplex, M. simunek, M. szaboi, M. tuomikoskii, M. zdena, Peripatetes acanthotarsus, Probonomoia pini, P. humprechti, Prowichmannia spiniferum, Rhopalanoetus fimetarius, Semianoetus mrcziaki, Spinanoetus heterotrichus, S. pelznerae, S. weingaertnerae, Stercoranoetus woolleyi Iceland (Sellnick, 1940) – Anoetus notabilis Ireland (Curry & Momen, 1988) - Histiostoma feroniarum Italy (contin.) (Canestrini & Berlese, 1881; Berlese, 1881, 1883, 1918; Oudemans, 1927) – Anoetoglyphus ateuchi, Glyphanoetus phyllotrichus, G. venustissimus, Histiostoma feroniarum, H. julorum, H. pyriforme, Rhopalanoetus fimetarius Sicily (Berlese, 1910; Samšinák, 1962) – Anoetoglyphus ateuchi, Anoetus curtipes, A. privorai, Histiostoma brevipodum Sardinia (Wirth, 2005) – Probonomoia opuntiae Norway (Fain et al., 1993) – Conglanoetus denticulatus Poland (Chmielewski, 1984; Kiełczewski & Wiśniewski, 1980; Michalski & Ratajczak, 1989; Haitlinger, 2002, 2004; Witaliński, Rożej-Pabijan & Podkowa, 2014) – Histiostoma crypturgi, H. ovalis, H. piceae, H. polypori, Spinanoetus pelznerae, S. weingaertnerae, Probonomoia pini, P. spinifera, Portugal (cont.) (Vissa et al., 2019) – Histiostoma ovatus Romania (Cooreman, 1941; Mahunka, 1970, 1977b) – Histiostoma gordius, H. histeri, H. lineatula, H. margaritatum, H. ovalis, Myianoetus tuomikoskii, Probonomoia humprechti, Xenanoetus vestigialis Russia (all, continent.) (Vitzthum, 1026; Sevastianov, 1973; Krivoluckiy, 1975; Wirth, 1916; Khaustov, Trach & Bobylev, 2016; Khaustov et al., 2018) – Anoetus exornatus, Bonomoia recondita, Histiostoma gladiger, H. ovale, H piceae, H. pini, Probonomoia pini, P. sibirica Crimea (Krivoluckiy, 1975) 110 Acarorum Catalogus IX Sakhalin (Sevastianov, 1973) - Myianoetus pseudomuscarum Kuril Islands (Klimov, 1998) – Scheucheria sp. Slovakia (Müller, 1859; Mahunka, 1972b, Samšinák, 1984, 1989; Mangová et al., 2017; Kaluz, 1992; Mašan & Országhi, 1995; Mašan, 2000; Mašan & Krištofik, 1992; Zamec, 2013, 2014) – Bonomoia recondita, Capronomoia sphaerocerae, Histiostoma crypturgi, H. berghi, H. crypturi, H. dryocoeti, H. feroniarum, H. maritimus, H. myrmicarum, H. ovalis, H. pannonicus, H. piceae, H. pini, H. polypori, H. pulchrum, H. sapromyzarum, Myianoetus diadematus, M. dionychus, M. longisetosus, M. ovatus, M. parvus, M. sovat, M. tuerkorum, M. tuomikoskii, M. virgatus, M. zdena, Rhopalanoetus lanceocrinus, Spinanoetus heterotrichus, S. pelznerae, S. weingaertnerae Slovenia (Moraza, Fernandez & Jurc, 2013) – Histiostoma ovale Spain (cont.) (Dimitrieff & Fain, 1994; Rebolledo & Arroyo, 1996; Fernandez, Diez & Moraza, 2013) – Histiostoma feroniarum, Probonomoia chrysis Canary Islands (Sanchez-Covisa et al., 1999; Wirth, 2005) – Probonomoia opuntiae (Gomera), Histiostoma feroniarum (Tenerife) Sweden (Moser et al. 1989; Moser, Perry et al., 1989) – Histiostoma piceae Switzerland (Mahunka, 1977; Fain, 1980) – Anoetus dolifer, Hormosianoetus aeschlimanni Turkey (all) (Mahunka, 1976) – Kaszabanoetus ulomae Ukraine (Sevastianov, 1970, 1971, 1973; Sevastianov & Vysotskaya, 1971) – Anoetus helobius, A. hospitus, A. opertus, A. parmatus, A. retractus, Bonomoia picturata, B. probata, Histiostoma indetonsum, H. necopinum, Myianoetus microti, M. paganus, M. pseudomuscarum, Prowichmannia spinifera AFRICA Angola (Mahunka, 1963a, 1963b, 1967) – Anoetus machadoi, A. striatus, A. vilhenaorum, Anoetoglyphus (?) bifastigatus, Glyphanoetus brevisetus, Histiostoma andrei, H. angolensis, H. arcuatum, H. bacilligerum, H. clavatum, H. impressum, H. minimum, H. pontigerum, H. puncaticeps, H. simile, H. subsimile, Hymenanoetus nodosus, Porrhanoetus machadoi Botswana (Camerik, Lukoschus & Kniest, 1980) – Fibulanoetus longitarsus, F. mahunkai Cabo Verde (Mahunka & Mahunka-Papp, 1991) – Glyphanoetus phyllotrichus, Histiostoma phyllophorum, Stercoranoetus harteni Cameroon (Mahunka, 1979b) - Histiostoma amieti, H. camerunensis, H. chilopophila, H. variabile Congo Rep. of the (Brazzaville) (Mahunka, 1965, 1967a;) – Anoetoglyphus africanus, A. (?) immaturus, Anoetus collis, A. congoensis, A. coprophilus, A. impressus, A. longisetus, A. machadoi, A. pauliana, A. punctulatus, A. striatus, A. tauffliebi, Glyphanoetus africanus, Rhaphidothrix gozmanyi, Rhopalanoetus bifurcatus Congo DR (Oudemans, 1929; Fain & Elsen, 1972; Fain & Beaucournu, 1973; Fain, 1974; Samšinák, 1962, 1989) – Anoetus fetters, Chiloanoetus mayumbensis, Curculanoetus rhynchophorus, Histiostoma conclaviculum, H. schoutedeni, Probonomoia congoen- Acarorum Catalogus IX 111 sis, Psyllanoetus afropulicarius, Rhopalanoetus bifurcatus, Scolianoetus scheucherae, Xenanoetus kabrt Egypt (Yousef & Metwally, 1973; Metwali & Ahmad, 1987; Hassan & Sevastianov, 1991; Eraky & Shoker, 1993a, 1993b, 1994, 1995; Eraky & Nasser, 1993; Eraky, 1993, 1994a, 1994b, 1997, 1998, 1999a, 1999b, 1999c, 2000, 2003; Abu El-Maged, 1998; AbdelAziz, 1999; Eraky, Abdel-Galil & Bohibah, 2010; Fawzy & Mtewallii, 2007; Eraky, Abdelgayed, Negm, Helal & Moussa, 2017; Abdelgayed et al., 2019; Eraky, AbdelAziz & Abo-Elmaged, 2020; Desoky et al., 2020) – Anoetus caljubiensis, A. irinus, A. moshtohorensis, A. sharae, A. wastis, Bonomoia zoherii, Coprobonomoia mahunkai, C. sphaerocerae, Glyphanoetus mahunkai, G. omari, G. processum, Histiostoma alii, H. arcuatus, H. bananai, H. camphori, H. cataglyphi, H. darwishii, H. diadematum, H. egyptiacum, H. essami, H. farghali, H. herbali, H. irinum, H. mannai, H. manuri, H. nasseri, H. negmi, H. onioni, H. pickaxei, H. rizkii, H. sammari, H. sarrai, H. solimani, H. rizkii, H. tinydorsalis, H. zaheri, Hormosianoetus mahunkai, Myianoetus granatum, M. lili, M. manuri, M. nerminka Fiji (Mahunka, 1992) – Histiostoma pseudolongipes Gambia (Mahunka, 1975, 1979) - Anoetoglyphus szunyoghyi, Ghanoetus marmoreus, Histiostoma endroedii, H. fucatifer, H. gambiense, H. umbrosa Ghana (Mahunka, 1973a, 1973d, 1974a, 1975, 1979) – Anoetus heliocopridis, A. sinuosus, Anoetoglyphus antennatus, A. globulus, A. latus, A. polygonatus, Bonomoia rugosa, Histiostoma distentosetum, H. editum, H. endroedii, H. furcatum, H. gorgo, H. rugosum, H. striatosimile, Rhaphidothrix punctolineatus, Rh. simplex, Rhopalanoetus hirsutus, Spinanoetus variabilis, Teinokyra lineata Ivory Coast (Fain, 1974) – Anoetus humilis, A. macrosuctus, Histiostoma adsimile, H. alphaticum, Glyphanoetus eboris Kenya (Mahunka, 1979) – Histiostoma bernoni, Myianoetus africanus Madagascar (Fain & Pauly, 2001) – Histiostoma heriades Morocco (Fain, Athias-Binche & Carton, 1993) – Hormosianoetus laboratorium Namibia (Fain, 1974) – Loxanoetus bassoni Rwanda (Fain, 1980, 1984) – Myianoetus brevis, M. rwandanus, M. uncinatus, Syringanoetus vandenberghi, Xenanoetus afrotropicalis, Seychelles (Nesbitt, 1979) – Creutzeria seychellensis South Africa (Fain, 1970; Mahunka, 1979; ) – Anoetoglyphus lootsi, Histiostoma volgini, Loxanoetus bassoni, Rhopalanoetus endroedyyoungai Tanzania (Oudemans, 1912, 1914; Mahunka, 1961, 1969b, 1974a, 1979) – Anoetus heliocopridis, A. longitubus, A. reticulatus, Glyphanoetus flagellates, G. longus, G. szunyoghyi, Histiostoma bifoveolatum, H. phyllophorum, H. polaki, H. telatum, H. trichophorum, Myianoetus reductus, Rhopalanoetus bifurcates, Rh. gladifer Tunisia (Mahunka, 1981) – Anoetoglyphus tunesiensis Uganda (Fain, 1974; Fain & Zumpt, 1974) – Auricanoetus longitarsus, Loxanoetus bassoni, Otanoetus wetzeli 112 Acarorum Catalogus IX Indian Ocean Islands (Mahunka, 1978) – Histiostoma schauenbergi ASIA Afghanistan (Mahunka, 1974c) – Histiostoma szelenyii China (Samšinák, 1962b; Li et al., 1992; Wang et al., 2002; Fan, Chen & Wang, 2010) – Histiostoma feroniarum, H. formosanum, Rhopalanoetus chinensis, Rh. simplex, Seliea pulchrum Taiwan (Mahunka, 1976; Phillipsen, 1977; Tseng Y.-H., 1982, 1984) – Histiostoma formosanum, H. gampsoceri, H. humiditatis India (Vitzthum, 1926; Chinniah & Mohanasundaram, 1995) – Histiostoma coprae, H. himalayae, H. muscae, H. onthophagae H. siphunculinae Indonesia (Oudemans, 1903, 1911, 1914; Vitzthum, 1921, 1931, 1932; Mahunka, 1976) – Histiostoma banjuwangicus, H. campanula, H. cyrtandrae, H. indicus, H. prodectoris, Munduytia tropicus, Scolianoetus javanus, Zwickia colocasiae Iran (Lotfollahi, Haddad Irani Nejad, Bagheri, Valizadeh, 2010) – Histiostoma polypori, H. litorale Israel and Palestina (Costa, 1967; Mahunka, 1979a) – Histiostoma feroniarum, H. loebli Japan (Kurosa, 2000, 2002; Takami & Kuwahara, 2005; Tagami, 2016; Kurosa & Tagami, 2006) – Histiostoma globotuberum, H. mahunkai, H. piscium, H. watanabei, Probonomoia pini Malaysia (Vitzthum, 1921; Hirst, 1928; Kurosa, 1976; Wurst & Kovac, 2003; Azmiera, Mariana & Heo C.C., 2019) – Anoetus myrmicarum, A. virginiana, Histiostoma sp., Tensiostoma veliaphilum Mongolia (Mahunka, 1967c, 1969c, 1972b, 1973) – Anoetus cacuminatus, A. insulanus, A. kaszabi, A. microti, A. porosus, Histiostoma insulare, Myianoetus kaszabi, Myianoetus tuomikoskii, Scheucheria mongolica, Xenanoetus grandiceps Pakistan (Khan, 1969; Chauhdri, 1986; Ashfaq, 1986, 1988, 2001; Ashfaq, Chauhdri & Aheer, 1985; Ashfaq & Chauhdri, 1988; Sarwar, Ashfaq & Islam, 1998; Ashfaq, Sarwar & Akbar, 1999; Ashfaq, Sarwar & Ali, 2000; Ashfaq, Sarwar & Amiad, 2000; Ashfaq & Sarwar, 2001) – Capronomoia asper, C. bellulus, C. gradates, C. pango, C. trossis, Glyphanoetus incanus, G. leptos, G. micidus, Histiostoma caro, H. crantor, H. densus, H. edax, H. erigo, H. fortis, H. fragosus, H. galerus, H. gracilis, H. imbecillum, H. junctus, H. karachiensis, H. luxus, Philippines (Vitzthum, 1921) - Histiostoma sapromyzarum Saudi Arabia (Samšinák, 1979; Negm & Alatawi, 2011) – Capronomoia sphaerocerae, Histiostoma pickaxei, H. camphori, Myianoetus lili Singapore (Hirst, 1928) – Zwickia nepenthesiana Sri Lanka (Oudemans, 1915; Mahunka, 1974d) – Anoetus aelleni, A. angularis, A. ceylanensis, Ceylanoetus excavatus, Zwickia guentheri Syria () – Histiostoma [Anoetus] sapromysarum, Caprononomoia [Bonomoia] sphaerocerae Taiwan (Mahunka, 1976b) – Histiostoma gampsoceri Acarorum Catalogus IX 113 Thailand (Fain & Lambrechts, 1985; Fain, Athias-Binche & Carton, 1993; ) – Histiostoma piscium, H. unidentatum Vietnam (Mahunka, 1967d, 1978; Fain & Beaucournu, 1972) – Anoetoglyphus langi, Anoetus capillatus, A. decorus, A. longipes, A. ornatus, A. pocsi, A. spinosus, Bonomoia reticulata NORTH AMERICA Canada (Woodring & Moser, 1970; Lindquist et al., 1979; Mullen et al., 1989; Fain & Colloff, 1990; Walter & Latonas, 2012; OConnor et al., 2015; Achtymichuk, 2017; Baulieu et al., 2019) – Anoetus sordida, Bonomoia sp., Histiostoma halictonida, Myianoetus muscarum, Traskorchestianoetus brevipes, T. spiceri. According to Baulieu et al., 2019, in Canada are known 21 sp. of Histiostomatidae and the number expected is 80. Mexico (Woodring & Moser, 1970) – Histiostoma varium USA (Banks, 1906, 1910, Ewing, 1911; Hughes & Jackson, 1958; Cross, 1968; Woodring, 1971; Moser & Roton, 1971; Woodring & Moser, 1970, 1975; Moser, 1976; Hill & Deahl, 1978; Fain & Philips, 1979; Mahunka, 1982; Fashing & OConnor, 1984; Samšinák, 1989; Wirth & Moser, 2010; Uppstrom & Klompen, 2011; Oconnor et al., 2015; Pimsler et al., 2016; Achtymichuk, 2017) – Ameranoetus hematobii, Anoetus cossi, A. grapefruit, A. media, A. sordida, Capronomoia sphaerocerae, Glyphanoetus nomiensis, Histiostoma bakeri, H. blomquisti, H. conjunctum, H. gracilipes, H. brevipes, H. guanophilum, H. halictonida, H. heinemanni, H. inquilinus, H. murchiei, H. nigrellii, H. pauciputeolum, H. pilosetum, H. phytophaga, H. protuberans, H. sextoni, H. tarsalis, H. varium, H. virginianum, Myianoetus micromaculatus, M. mlkoch, M. muscarum, M. rohaceki, Sarraceniopus darlingtoniae, Xenanoetus kofrankova SOUTH and CENTRAL AMERICA Argentina (Mahunka, 1963c, 1963d) – Anoetus akodoni, Histiostoma [Anoetus] sapromysarum Bolivia (Vitzthum, 1921) – Histiostoma sapromyzarum Brazil (Vitzthum, 1926a; Fain, 1974; Fain & Camerik, 1978; Mahunka, 1982; Souza, Bernardi & Ferreira, 2018) – Bothyanoetus pereirai, Histiostoma acrocini, H. sapromyzarum, H. tanypus, Scolianoetus brasiliensis, S. ramozae, Spartocera cinammomea Chile (contin.) (Mahunka, 1963, 1972d) – Anoetus castrii, A. chilensis, A. cirrosus, Synanoetus baloghi Juan Fernandez Islands (Mahunka, 1982) – Glyphanoetus pappi Colombia (Fain & Rack, 1987) – Probonomoia columbiana Costa Rica (Woodring, 1973; Mahunka, 1976, 1978a, 1979) – Histiostoma adarcuale, H. claviger, H. coronatum, H. duplex, H. eickworthi, H. marianae, H. moralesi, H. perruptum, H. rostratum, Pteranoetus incertus, Semianoetus banksi Cuba (Samšinák, 1966, Mahunka, 1978) – Histiostoma cubanum, H. meszarosorum, Kanoetus cubanus 114 Acarorum Catalogus IX Ecuador (contin.) (Mahunka, 1978; Samšinák, 1989) – Anoetus helena, Conglanoetus ornatofoveolatus, Histiostoma capillatum, H. continuatum, H. diadematum, H. elzingai, H. rettenmeyerorum, H. rimosa, H. scinditum, Pteranoetus latus, P. rettenmayeri, Richardanoetus demel Guadeloupe (Fain, Athias-Binche & Carton, 1993) – Hormosianoetus laboratorium Guatemala (Woodring & Moser, 1975) – Histiostoma conjunctum Honduras (Woodring & Moser, 1970, 1975) – Histiostoma conjunctum, H. varium Panama (Mahunka, 1978a) – Histiostoma incurvatum Peru (Caceres & Fain, 1975) – Histiostoma ferroniarum Puerto Rico (Fain & Santiago-Blay, 1993) – Ancyranoetus virkkii St. Vincent Republic, Grenadines Isl., Bequia Id. (Windward Group)(Fain & Van Goethem, 1985) – Ovanoetus wautersi Windward Islands (St. Lucia) (Mahunka, 1981) – Histiostoma antillense, H. mirabile OCEANIA Australia (continent) (Womersley, 1941, 1942; Mahunka, 1975; Fain & Domrow, 1980; Halliday, 1998; Halliday & Collins, 2002; Tagami & Halliday, 2013) – Chiropteranoetus chalinolobus, Histiostoma australiense, H. feroniarum, H. humiditatis, H. papillata, H. phyllophorum (= nichollsi Womersley), H. sapromyzarum, H. titanolabi, H. unidentatum, H. varium, Loxanoetus lenae, Myianoetus dycei, M. oudemansi New Zealand (Fain & Galloway, 1993; Qin & Rohitha, 1996; Clark, 2009, 2010a, 2010b) – Histiostoma feroniarum, H. mantelli, Myianoetus antipodus Papua New Guinea (Fain, 1968; Mahunka, 1975, 1976, 1978) – Histiostoma amargynus, H. foveosimile, H. lineosculpturatum, H. reticulatissimum, H. reticulofoveolatum, H. szentivanyi, H. wauense, Myianoetus domrowi French Polynesia (Tahiti) (Mahunka, 1992) – Histiostoma berndhauseri French Polynesia (Hivaoa) (Vitzthum, 1935) – Histiostoma granulatum Fiji (Mahunka, 1995,1992) – Anoetus peloponesiacus, Histiostoma genavensium, H. pseudolongipes Marquesan Islands (Vitzthum, 1935) – Histiostoma granulatum Salomon Islands (Mahunka, 1972c) – Anoetus bradymeri, A. horridus, A. solomoniensis, A. toxici ANTARCTICA Kerguelen Islands (Fain, 1976) – Amyzanoetus halophilus, Austranoetus kerguelenensis, Histiostoma kerguelenense, Myianoetus travei Acarorum Catalogus IX 115 Hosts of Histiostomatidae Hirudinea Fam. Aulostomidae Aulastoma gulo (cocoons) – Histiostoma berghi Crustacea Amphipoda Fam. Talitridae Traskorchestia traskiana – Traskorchestianoetus brevipes Isopoda Fam. Oniscidae Oniscus asellus – Histiostoma phyllophorum Oniscus sp. – Histiostoma feroniarum Decapoda Fam. Paguridae Coenobita clypeata – Ovanoetus wautersi Arachnida Acari Fam. Gamasidae Eugamasus cornutus – Histiostoma polypori Myriapoda Diplopoda Fam. Iulidae Julus scandinavius - Histiostoma feroniarum, H. litorale, H. sapromyzarum Chilopoda Fam. Lithobiidae Lithobius forficatus – Histiostoma gervaisi Fam. Scolopendridae Cormocephalus aurantiipes - Histiostoma feroniarum Insecta Orthoptera Fam. Phalangopsidae Endecous (Endecous) alejomesai – Histiostoma sp. (cave) Coleoptera Fam. Carabidae Demetrias imperialis – Histiostoma disiunctum, H. serratum Dichaetodrilus incrassatus - Histiostoma striatum Fam. Brenthidae Ectocemus cinnamomeus – Histiostoma prodectoris Eubactrus semiaeneus – Anoetus polynesiacus 116 Acarorum Catalogus IX Prodector fruhstorferi – Histiostoma prodectoris Prophthalmus potens – Histiostoma prophthalmi Schizotrachelus bakeri – Histiostoma sapromyzarum Fam. Platypodidae Platypus dispar - Histiostoma crassipes Fam. Cerambycidae Acanthocinus aedilis – Probonomoia humprechti, P. pini Acrocinus longimanus – Histiostoma acrocini Rhagium sp. – Histiostoma crypturgi Fam. Chrysomelidae Galeruca sp. – Histiostoma necopinus Lema nigripes – Ancyranoetus virkkii Fam. Coccinellidae Gen. sp. – Histiostoma lorentzi Fam. Curculionidae Curculioninae s.l. Rhynchophorus phoenicis – Anoetoglyphus bifastigatus, Curculanoetus rhynchophorus, Histiostoma striatum Scolytinae Crypturgus pusillus – Histiostoma crypturgi Dendroctonus brevicomis - Bonomoia certa D. frontalis - Histiostoma conjunctum D. micans – Histiostoma vitzthumi D. pseudotsugae - Bonomoia certa Dryocoetus autographus – Histiostoma dryocoeti Dryocoetes villosus – Anoetus pannonicus Hylaster ater – Probonomoia pini Ips avulsus - Histiostoma conjunctum I. cembrae – Histiostoma oudemansi I. curvidens – Histiostoma abietis I. grandicollis – Histiostoma varium I. laricis – Histiostoma gordius I. sexdentatus – Histiostoma ovalis (= gladiger) I. typographus – Histiostoma piceae Mycetophilus piniperda – Probonomoia pini Polygraphus minor - Histiostoma himalayae Scolytus ventralis – Bonomoia certa Fam. Elateridae Melanotus rufipes – Histiostoma ruehmi, H. sapromyzarum Fam. Histeridae Carcinops 14-striata – Histiostoma hungaricum Carcinops sp. – Anoetus sordida Acarorum Catalogus IX 117 Hister merdarius – Histiostoma feroniarum H. stercorarius - Histiostoma feroniarum H. unicolor - Histiostoma feroniarum Hololepta africanae - Histiostoma crassipes Platysoma compressum – Histiostoma histeri, H. lineatula, H. margaritatum, H. ovale Fam. Hydrophilidae Hydrophilus caraboides – Histiostoma maritimus Sphaeridium sp. - Rhopalanoetus lanceocrinus, Xenanoetus vestigialis Fam. Lucanidae Dorcus parallelopipedus – Histiostoma dryocoeti, Probonomoia pini Fam. Nitidulidae Brachypeplus sp. – Histiostoma granulatum Fam. Passalidae Passalus sp. - Scolianoetus ramozae Fam. Scarabaeidae Aphodius fimetarius – Rhopalanoetus lanceocrinus A. fossor – Rhopalanoetus lanceocrinus A. frater – Aphodanoetus teinophallus A. merdarius – Rhopalanoetus lanceocrinus Bothynus deiphobus - Bothyanoetus pereirai Bubas bubalus - Histiostoma sapromyzarum Copris lunaris - Anoetoglyphus copridis Cetonia aurata – Prowichmannia spiniferum Geotrupes stercorarius - Rhopalanoetus lanceocrinus Melolontha hippocastani – Histiostoma melolonthae Oniticellus cinctus – Rhopalanoetus chinensis, Rh. simplex Onitis sphinx – Rhopalanoetus bifurcatus Onthophagus pactolus – Histiostoma onthophagae Oxythyrea funesta – Histiostoma phyllophorum Polyphyla fullo - Histiostoma melolonthae Scarabaeidae - Anoetoglyphus langi Fam. Staphylinidae Bledius bicornis – Insulanoetus brachypodus B. sarmaticus - Insulanoetus brachypodus Emus hirtus - Spinanoetus weingaertnerae Lathrobium (Lathrobium) sp. – Histiostoma watanabei Ontolestes sp. - Spinanoetus weingaertnerae Philontus sp. - Spinanoetus weingaertnerae Fam. Silphidae Nicrophorus interruptus - Spinanoetus pelznerae Oeceoptoma thoracica - Spinanoetus pelznerae 118 Acarorum Catalogus IX Thanatophilus rugosus - Spinanoetus pelznerae Th. sinuatus - Spinanoetus pelznerae Fam. Tenebrionidae Amarygmus [a “Amargynus”, or “Amargymus”] cf. capillatus - Histiostoma amargynum, H. szentivanyi Amarygmus [as “Amargynus”] cf. deformipes - Histiostoma reticulofoveolatum Androsus greensladei – Anoetus bradymeri Bradymerus leveri - Anoetus bradymeri Corticeus coynei – Anoetus sordida Helops quisquilius - Histiostoma ulmi Hypophloeus bicolor – Histiostoma ulmi Leiochrodes suturalis – Anoetus horridus, A. solomoniensis Platolenes femoratus – Histiostoma wauensis Platolenes sp. - Histiostoma lineosculpturatum Prioscelis serrata – Chiloanoetus mayumbensis Scyophagus piceus - Anoetus horridus Toxicum quadricorne rufipes – Anoetus toxici Uloma culinaria – Kaszabanoetus ulomae Fam. Trogidae Trox scaber – Rhopalanoetus trogicola T. cadaverinus - Rhopalanoetus trogicola Dermaptera Fam. Labiduridae Labidura riparia – Anoetus moshtohorensis, A. wastis, Histiostoma camphori Fam. Anisolabididae Anisolabella (Gonolabis) marginalis – Histiostoma mahunkai, H. piscium A. maritima – Histiostoma piscium Titanolabis colossea – Histiostoma australiense, H. humiditatis, H. titanolabi Fam. Forficulidae Forficula auricularia – Histiostoma polypori Heteroptera Fam. Coreidae Spartocera cinammomea - Scolianoetus brasiliensis Fam. Veliidae Lathriovelia rickmersi – Tensiostoma veliaphilum L. capitata - Tensiostoma veliaphilum Hymenoptera Fam. Megachilidae Heriades erythrosoma – Histiostoma heriades Fam. Apidae Xenoglossa strenua – Histiostoma inquilinus Acarorum Catalogus IX 119 Fam. Halictidae Augochlora spp. – Histiostoma eickworthi Halictus holtzi – Histiostoma szelenyii H. sexcinctus – Histiostoma halicticolum H. rubicundus - Histiostoma halictonida H. tatrazonianellus – Histiostoma orientalis Megalopta spp. - Histiostoma halictonida Nomia melanderi – Glyphanoetus nomiensis N. nortoni - Glyphanoetus nomiensis Fam. Formicidae Aphaenogaster testaceopilosa – Anoetus privorai Camponotus sp. - Histiostoma myrmicarum Eciton burchelli – Histiostoma incurvatum, Pteranoetus incertus E. hamatum – Histiostoma clavifer, H. moralesi, H. rostratum E. lucanoides – Histiostoma elzingai, H. scinditum Formica fusca - Histiostoma myrmicarum F. lemani – Anoetus komarpictoris, A. shamyi F. rufa – Anoetus exornatus Gymnometopina kibokoensis – Anoetus caljubiensis Labidus praedator - Conglanoetus ornatofoveolatus, Histiostoma capillatum, H. rettenmeyerorum Labidus sp. – Histiostoma continuatum, H. rimosum Lasius flavus - Histiostoma myrmicarum L. niger - Histiostoma myrmicarum Myrmica laevinodis - Histiostoma myrmicarum M. ruginodis – Anoetus dolifer, Bonomoia recondita, Histiostoma myrmicarum M. rubra - Anoetus opertus M. sabuleti – Anoetus dolifer Neivamyrmex gibbatus – Histiostoma diadematum N. gradualis – Pteranoetus rettenmeyeri Solenopsis invicta – Histiostoma blomquisti Fam. Scoliidae Campsomeris mansueta – Scolianoetus scheucherae Fam. Chrysididae Chrysis sp. – Probonomoia chrysis Fam. Vespidae Vespidae gen. sp. – Histiostoma tanypus Diptera Fam. Asilidae Laphria flava – Probonomoia humprechti Fam. Ceratopogonidae Culicoides brevitarsis – Myianoetus dycei 120 Acarorum Catalogus IX Fam. Chloropidae Gampsocera mutata - Histiostoma gampsoceri Scoliophthalmus sp. – Anoetus domrowi Siphunculina sp. – Histiostoma siphunculinae Fam. Drosophilidae Drosophyla bipectinata – Histiostoma unidentatum D. willstoni – Hormosianoetus laboratorium D. subobscura - Hormosianoetus laboratorium Drosophila sp. - Hormosianoetus aeschlimanni Fam. Glossinidae Glossina fuscipes quanzensis –Histiostoma schoutedeni, Probonomoia congoensis Fam. Muscidae Azelia parva – Myianoetus pappi Haematobia irritans – Ameranoetus haematobii Hebecnema umbratica – Myianoetus simplex Hydrotaea occulta – Myianoetus tuomikoskii H. tuberculata – Myianoetus tuomikoskii Hydrotaea sp. – Myianoetus longisetosus, M. ovatus Musca domestica – Histiostoma muscae Ophyra leucostoma – Myianoetus parvus Synthesiomyia nudiseta – Myianoetus muscarum Fam. Fanniidae Fannia sp. – Myianoetus mystacis Fam. Sepsidae Sepsis thoracica – Myianoetus simplex Fam. Heleomyzidae Amoebaleria cacsia - Myianoetus diadematus A. spectabilis - Myianoetus diadematus Borboropsis fulviceps - Myianoetus parvus Gymnomus caesius - Myianoetus sovat Helomyza brachypterna - Myianoetus diadematus H. serrata - Myianoetus diadematus Fam. Sphaeroceridae Borborillus sordidus - Myianoetus mlcoch Coproica acutangula – Xenanoetus kofrankova C lugubris – Myianoetus simplex, M. simunek, M. zdena C. pallifrons - Myianoetus zdena C. vagans - Myianoetus zdena C. viripennis – Myianoetus rohaceki Elachisoma aterrima - Myianoetus zdena Gymnometopina kibokoensis – Xenanoetus kabrt Acarorum Catalogus IX 121 G. magna - Anoetus fetters, Syringanoetus vandenberghi Ischiolepta pusilla - Capronomoia sphaerocerae Limosina clunipes – Myianoetus virgatus L. rufilabris – Myianoetus rohaceki L. talparum - Glyphanoetus phyllotrichus Lotobia pallidiventris - Myianoetus zdena Lotophila atra - Myianoetus rohaceki Metaborborus calcaratus – Anoetus fetters Philocoprella quadrispina - Myianoetus zdena Physiophora sp. - Capronomoia sphaerocerae Pilocoprella quadrispina - Myianoetus mlcoch Richardia sp. – Anoetus helena, Richardanoetus demel Spelobia clunipes - Myianoetus virgatus Sphaerocera curvipes - Capronomoia sphaerocerae S. subsultans - Capronomoia sphaerocerae Fam. Syrphidae Mallota bautius – Hormosianoetus mallotae M. posticata - Hormosianoetus mallotae Fam. Tabanidae Hybomitra affinis – Bonomoia sp. H. epistates - Bonomoia sp. H. illota - Bonomoia sp. H. lasiophthalmus - Bonomoia sp. H. nuda - Bonomoia sp. Tabanus quinquestriatus - Bonomoia sp. Lepidoptera Fam. Cossidae Cossus cossus – Histiostoma cossi, H. dryocoeti C. ligniperda - Histiostoma cossi Fam. Hesperiidae Pelopides tridens - Scolianoetus javanus Pisces Perciformes Fam. Percichthyidae Maccullochella peelii peelii – Histiostoma papillatum Siluriformes Fam. Pangasiidae Pangasius sutchi – Histiostoma piscium Anguilliformes Fam. Anguillidae Anguilla anguilla – Histiostoma anguillarum 122 Acarorum Catalogus IX Aves Sphenisciformes Fam. Spheniscidae Eudyptula minor albosignata – Histiostoma mantelli, Myianoetus antipodus Mammalia Rodentia Fam. Sciuridae Spermophilus [Citellus] citellus – Myianoetus clavatus Fam. 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Biologica, 82(1): 63–98. Youssef A.A., S.H. Metwally. 1973. A new species of the genus Histiostoma Kramer (Acarina, Anoetidae). – Acarologia, 15(4): 724–726. Yousef, A. A., E.A. El Badry, S.H. Metwally. 1978. Life history of the anoetid mite Histiostoma cataglyphi Yousef et Metwally, with a description of the immature stages (Acari, Astigmata, Anoetidae). - Journal of Applied Entomology, 87: 225–229. Zachvatkin A.A. 1941. Tyroglyphoidea. Fauna SSSR: Paukoobraznie, 6(1). Akademiya Nauk SSSR: Moscow-Leningrad. 475 pp. (in Russian, French abstract). Zamec R., V.Košel. 2014. A new species of mite (Acari: Histiostomatidae) phoretic on Gymnomus caesius (Diptera: Heleomyzidae) from Vlčie Diery Cave. – Biologia (Bratislava), 69(7): 916–918. The Mite Family Guanolichidae Fain Guanolichidae Fain, 1968: 199 Small family of four known species in Africa (Gabon) and America (Suriname, Panama and Mexico). These completely sclerotized mites live in the bat guano, some were found on bats (Noctilionidae, Natalidae). Composition of Guanolichidae Family Guanolichidae Fain, 1968–3 gen., 4 sp. Guanolichoides Fain, 1979–1 sp. Guanolichus Fain, 1968–1 sp. Neoguanolichus Fain, 1979–2 sp. 142 Acarorum Catalogus IX Catalogue of Guanolichidae Family Guanolichidae Fain Guanolichidae Fain, 1968: 199 Type genus: Guanolichus Fain, 1968 Genus Guanolichoides Fain Guanolichoides Fain, 1979: 117 Type species: Guanolichoides lukoschusi Fain, 1979 Guanolichoides lukoschusi Fain Guanolichoides lukoschusi Fain, 1979: 117 Type locality: Phedra Distribution: Suriname Host: Noctilio labialis (Chiroptera: Noctilionidae) Genus Guanolichus Fain Guanolichus Fain, 1968: 203 Type species: Guanolichus gabonensis Fain, 1968 Guanolichus gabonensis Fain Guanolichus gabonensis Fain, 1968: 203 Type locality: bat guano, cave Faucon, Belinga, Gabon Distribution: Gabon Substrate: bat guano Genus Neoguanolichus Fain Neoguanolichus Fain, 1979: 117 [Redescription, Fashing 1980: 109] Type species: Neoguanolichus mexicanus Fain, 1979 Neoguanolichus mexicanus Fain Neoguanolichus mexicanus Fain, 1979: 118 Type locality: Durango Distribution: Mexico Substrate: from a bat, Natalus stramineus ventanus (Chiroptera: Natalidae) Neoguanolichus panamensis Fashing Neoguanolichus panamensis Fashing, 1980: 109 Type locality: Cocle 1 Cave, 0.8 mile North of Penonomé Distribution: Panama Substrate: bat guano Acarorum Catalogus IX 143 Index of Guanolichidae (Names in bold are accepted as valid) gabonensis, Guanolichus 142 Guanolichidae 142 Guanolichus 142 lukoschusi, Guanolichoides 142 mexicanus, Neoguanolichus 142 Neoguanolichus 142 panamensis, Neoguanolichus 142 Distribution of Guanolichidae AFRICA Gabon (Fain, 1968) - Guanolichus gabonensis NORTH AMERICA Mexico (Fain, 1979) – Neoguanolichus mexicanus CENTRAL AMERICA Panama (Fashing, 1980) – Neoguanolichus panamensis SOUTH AMERICA Suriname (Fain, 1979) – Guanolichoides lukoschusi Hosts of Guanolichidae Chiroptera Fam. Natalidae Natalus stramineus ventatus – Neoguanolichus mexicanus Fam. Noctilionidae Noctilio labialis – Guanolichoides lukoschusi Bibliography of Guanolichidae Fain A. 1968. Deux nouveaux Acariens cavernicoles du Gabon (Sarcoptiformes). – Biologia Gabonica, 4(2): 195–205. 144 Acarorum Catalogus IX Fain A. 1979. Notes sur les Acariens astigmates cavernicoles. Description de deux genres nouveaux et de quatre espèces nouvelles. - Acarologia (Paris), 20(1): 116–126. Fashing N.J.1980. Neoguanolichus panamensis, a new species of Guanolichidae from bat caves in Panama (Acarina: Sarcoptiformes). – Acarologia (Paris), 1979, 21(1): 108–116. The Mite Superfamily Canestrinioidea Superfamily of four families (Canestriniidae, Chetochelacaridae, Lophonotacaridae and Heterocoptidae) of relatively large mites (113 genera, ca. 376 valid species) living as commensals or parasites on beetles (Coleoptera: Carabidae, Chrysomelidae, Elateridae, Scarabaeidae, Tenebrionidae, Passalidae and others), mostly under elytrae. One genus and species of Canestriniidae (Diplopodocoptes transkeiensis) and the mites of the families Chetochelacaridae and Lophonotacaridae are associated with Myriapods (Diplopoda). The Mite Family Canestriniidae Canestriniidae Berlese, 1884 As early as 1878 G. Canestrini described “Dermaleichus” [now Dicanestrinia] cerambycis. The study of the Canestriniidae (as of many other groups of mites) starts with some papers of Berlese (1881, 1882, 1884, 1910, 1911, 1920), with descriptions of the genera Canestrinia, Canestriniella, Coleopterophagus, Dicanestrinia and Percanestrinia and of the family Canestriniidae. Oudemans (1904) described the important genus Photia. Until the work of Cooreman (1950, 1955, 1958) and Lombardini (1950) have been described only five more genera (by Trägårdh, 1906, Banks, 1915, Lombardini, 1943, Turk, 1948) and 16 valid species. The publications of Cooreman, and especially the one of 1950, laid the base of the modern study of Canestriniidae (including nine new genera and several new species). In the period between 1950 and 1988 several other authors contributed to the study of these mites (Lavoipierre, Vomero, Fain, Nesbitt, Lindquist, Beron and others). Should be marked especially the results obtained by K. Samšinák (1963–1971) and Summers and Schuster (1979–1982). In the important paper of Samšinák (1971) have been described many new genera and new species from Carabidae. In five papers Summers and Schuster published the subfamily Melisiinae, 11 new genera and many new species. Until the beginning of the work of R. Haitlinger (1988, but including the paper of Fain, 1989) have been published 34 valid genera and 146 species of the family Canestriniidae. From the 100 valid genera almost half (42) are monotypic. In more than 450 papers since 1988 R. Haitlinger contributed most of all to the study of Canestriniidae. He described 53 of the 100 valid genera and 200 of the 348 valid species in the family (including the subfamily Vereoxiinae). The papers of Haitlinger deal with Acarorum Catalogus IX 145 material from more than 50 countries and changed the overall picture of the distribution and the list of hosts of these mites. Most of the new genera are so far monotypic. Recently some newer authors joint the research efforts of Haitlinger. Kim et al. (2006) described the new genus Haitlingeria, Trach, Eidelberg and Khaustov contributed to the study of Canestriniids of Ukraine, including descriptions of four new genera. Composition of Canestriniidae Fam. Canestriniidae - 100 gen., 348 sp. Subfam. Canestriniinae Acrotocarus Banks, 1915–3 sp. Afrocanestrinia Cooreman, 1955–3 sp. Albinorattia Haitlinger, 1989–1 sp. Amansiella Khaustov et Eidelberg, 2001–1 sp. Ambilohylla Haitlinger, 1990–1 sp. Amboinophela Haitlinger, 1993–1 sp. Anaspistes Summers et Schuster, 1982–1 sp. Apalotacarus Summers et Schuster, 1981–17 sp. Arraphosoma Summers et Schuster, 1981–1 sp. Arrunsithiana Haitlinger, 1992–10 sp. Athogavia Haitlinger, 1989–7 sp. Aurilossongia Haitlinger, 1989–6 sp. Averithiana Haitlinger, 1999–1 sp. Barbiangia Haitlinger, 1993–3 sp. Beeriphotia Haitlinger, 1994–1 sp. Bibulothiana Haitlinger, 1993–6 sp. Bircericola Haitlinger, 2000–1 sp. Boetophela Haitlinger, 1989–10 sp. Boleohylla Haitlinger, 1991–2 sp. Camirohylla Haitlinger, 1991–1 sp. Canestrinia Berlese, 1881–17 sp. Canestriniella Berlese, 1910–6 sp. Caraboecius Cooreman, 1950–1 sp. Cetonicola Cooreman, 1955–5 sp. Chelinochroa Summers et Schuster, 1982–1 sp. Cnecoderma Summers et Schuster, 1981–2 sp. Coleoglyphus Berlese, 1910b - 2 sp. Coleopterophagus Berlese, 1882–11 sp. Cooremania Nesbitt, 1976–2 sp. Danaithiana Haitlinger, 1991b – 4 sp. 146 Acarorum Catalogus IX Dicanestrinia Berlese, 1911–3 sp. Diplognathophilus Cooreman, 1955–1 sp. Diplopodocoptes Fain, 1987–1 sp. Donnelaphontia Lavoipierre, 1958–2 sp. Dorynothiana Haitlinger, 1999–2 sp. Ennodithiana Haitlinger, 1993–6 sp. Eunorithiana Haitlinger, 1999–2 sp. Gasuthiana Haitlinger, 1989–5 sp. Gioharattia Haitlinger, 1989–2 sp. Globosophotia Haitlinger, 1991–1 sp. Grandiellina Fain, 1989 (= Grandiella Lombardini, 1939 praeocc.) - 21 sp. Grandiellopsis Fain, 1989–7 sp. Haitlingeria Kim et al., 2006–1 sp. Hargeinella Haitlinger, 1989–1 sp. Hatohylla Haitlinger, 1990–4 sp. Hyllosihemia Haitlinger, 1994–1 sp. Hypopteryx Summers et Schuster, 198–1 sp. Ibizella Haitlinger et Šundić, 1918–3 sp. Irmongia Haitlinger, 1990b – 1 sp. Jowitella Haitlinger, 1990b – 1 sp. Jullongia Haitlinger, 1990b – 1 sp. Kahoorangia Haitlinger, 1991–1 sp. Lidiophela Haitlinger, 1990b – 7 sp. Megacanestrinia Trägårdh, 1906–2 sp. Subfam. Melisiinae Contramelisia Samšinák, 1969–1 sp. Melisia Lombardini, 1944–22 sp. Mesophotia Samšinak, 1971–1 sp. Metriothiana Haitlinger, 1999–1 sp. Mossongia Haitlinger, 1989–2 sp. Neophotia Samšinák, 1971–3 sp. Noemiphela Haitlinger, 1992–2 sp. Oceogavia Haitlinger, 1989–1 sp. Olafithiana Haitlinger, 1993–5 sp. Olgattia Haitlinger, 1990–1 sp. Ovarrothiana Haitlinger, 1993–6 sp. Paracanestrinia Haitlinger, 2017–1 sp. Paramansia Cooreman, 1950–3 sp. Paramelisia Summers et Schuster, 1982–1 sp. Paraphagella Cooreman, 1955–6 sp. Paraphotia Khaustov et Eidelberg, 2001–1 sp. Acarorum Catalogus IX 147 Passalophagus Nesbitt, 1976–1 sp. Percanestrinia Berlese, 1911–8 sp. Percanestriniella Khaustov et Eidelberg, 2001–4 sp. Peruthiana Haitlinger, 2008–1 sp. Phaleratus Nesbitt, 1976–2 sp. Phelliculophela Haitlinger, 1993–1 sp. Photia Oudemans, 1904–18 sp. Phriknodora Summers et Schuster, 1982–3 sp. Procericola Cooreman, 1950–2 sp. Pseudamansia Cooreman, 1950–2 sp. Pseudocanestrinia Khaustov et Eidelberg, 2001–5 sp. Pseudoparaphagella Haitlinger, 2010–1 sp. Rugoniphela Haitlinger, 1992–1 sp. Sabothiana Haitlinger, 1993–3 sp. Sajanophela Haitlinger, 1990–1 sp. Sandrophela Haitlinger, 1990–7 sp. Saniothiana Haitlinger, 1990–2 sp. Scarabiphotia Haitlinger, 1994–1 sp. Shillongia Haitlinger, 1990–1 sp. Sorbinophela Haitlinger, 1993–1 sp. Tamarangia Haitlinger, 1991–4 sp. Tamarophela Haitlinger, 1990–1 sp. Taseopus Summers et Schuster, 1982–1 sp. Teklathiana Haitlinger, 1999–3 sp. Teophyssa Haitlinger, 1990b – 3 sp. Thopia Summers et Schuster, 1982–6 sp. Uriophela Haitlinger, 1992–1 sp. Vandeinella Haitlinger, 1989–2 sp. Subfam. Vereoxiinae Vereoxia Haitlinger, 1995–1 sp. Catalogue of the Family Canestriniidae Family Canestriniidae Berlese Canestriniidae Berlese, 1884: 92 Type genus: Canestrinia Berlese, 1881 Subfamily Canestriniinae Berlese Canestriniidae Berlese, 1884: 92 Type genus: Canestrinia Berlese, 1881 148 Acarorum Catalogus IX Genus Acrotocarus Banks Acrotocarus Banks, 1915: 152 Type species: Acrotocarus mirabilis Banks, 1915 Acrotocarus alutaceus (Turk) Amansia alutacea Turk, 1948: 88 Acrotocarus alutaceus (Turk): Summers & Schuster 1979b: 480 Type locality: St. Augustine Distribution: Trinidad Host: Physonota alutacea (Coleoptera: Chrysomelidae, Cassidinae) Acrotocarus crataepus (Summers et Schuster) Acrotocarus crataepus Summers & Schuster, 1979b: 484 Type locality: unknown Distribution: unknown Host: Physonota dilata (Coleoptera: Chrysomelidae, Cassidinae) Acrotocarus mirabilis Banks Acrotocarus mirabilis Banks, 1915: 152; Summers & Schuster, 1979b: 476 Type locality: Cordoba (Mexico) Distribution: Costa Rica, Guatemala, Mexico Hosts: Physsonota alutacea, Physsonota sp. (Coleoptera: Chrysomelidae) Genus Afrocanestrinia Cooreman Afrocanestrinia Cooreman, 1955: 25 Type species: Afrocanestrinia straeleni Cooreman, 1955 Afrocanestrinia rufinae Haitlinger Afrocanestrinia rufinae Haitlinger, 1995a: 85 Type locality: Tanzania Distribution: Tanzania Host: Smaragdesthes oertzeni (Coleoptera: Scarabaeidae, Cetoniinae) Afrocanestrinia samsoni Haitlinger Afrocanestrinia samsoni Haitliriger, 1995a: 87 Type locality: Senegal Distribution: Senegal Host: Smaragdesthes africana (Coleoptera: Scarabaeidae, Cetoniinae) Afrocanestrinia straeleni Cooreman Afrocanestrinia straeleni Cooreman, 1955: 25; Haitlinger, 1995: 85 Acarorum Catalogus IX 149 Type locality: May-ya-Moto, Rutshuru Distribution: D.R. Congo Hosts: Smaragdesthes africana mutica, Dicronorhina micans (Coleoptera: Scarabaeidae, Cetoniinae) Genus Albinorattia Haitlinger Albinorattia Haitlinger, 1989: 588 Type species: Albinorattia igori Haitlinger, 1989 Albinorattia igori Haitlinger Albinorattia igori Haitlinger, 1989: 588 Type locality: Batjan Distribution: Indonesia Host: Ischipsophia olivacea (Coleoptera: Sarabaeidae, Cetoniinae) Genus Amansiella Khaustov et Eidelberg Amansiella Khaustov & Eidelberg, 2001: 41 Type species: Amansiella trilobata Khaustov et Eidelberg, 2001 Amansiella trilobata Khaustov et Eidelberg Amansiella trilobata Khaustov & Eidelberg, 2001: 41; Trach, 2006: 176; Trach & Khaustov, 2007: 268 Type locality: Crimea, Yalta Distribution: Crimea, Ukraine Host: Timarcha tenebricosa (Coleoptera: Chrysomelidae) Genus Ambilohylla Haitlinger Ambilohylla Haitlinger, 1990: 316 Type species: Ambilohylla Haitlinger, 1993 Ambilohylla favosa Haitlinger Ambilohylla favosa Haitlinger, 1990: 316 Type locality: Madagascar Distribution: Madagascar Host: Damatria formosus (Coleoptera: Tenebrionidae) Genus Amboinophela Haitlinger Amboinophela Haitlinger, 1993: 339 Type species: Amboinophela bernae Haitlinger, 1993 Amboinophela bernae Haitlinger Amboinophela bernae Haitlinger, 1993b: 340 150 Acarorum Catalogus IX Type locality: Moluccas, Amboina Is. Distribution: Indonesia, New Guinea Host: Metapodonotus bison, M. cinctus (Coleoptera: Lucanidae) Genus Anaspistes Summers et Schuster Anaspistes Summers & Schuster, 1982b: 43 Type species: Anapistes unguiculatus Summers et Schuster, 1982 Anaspistes unguiculatus Summers et Schuster Anaspistes unguiculatus Summers & Schuster, 1982b: 43 Type locality: Kisarawe Distribution: Tanzania Host: Sternotomis bohemanni (Coleoptera: Cerambycidae) Genus Apalotacarus Summers et Schuster Apalotacarus Summers & Schuster, 1981a: 28 Type species: Apalotacarus fusulus Summers et Schuster, 1981 Apalotacarus aristatus Summers et Schuster Apalotacarus aristatus Summers & Schuster, 1981a: 28 Type locality: Mucha Distribution: Taiwan, Indonesia (Java), India (Assam) Host: Aceraius grandis hirsutus (Coleoptera: Passalidae) Apalotacarus cidaris Summers et Schuster Apalotacarus cidaris Summers & Schuster, 1981a: 25 Type locality: Daru Distribution: Papua New Guinea Host: Protomocoelus australis (Coleoptera: Passalidae) Apalotacarus echinatus Summers et Schuster Apalotacarus echinatus Summers & Schuster, 1981a: 33 Type locality: 12 mi.W of Munnar, S. India Distribution: India Host: Pleurarius brachyphilus (Coleoptera: Passalidae) Apalotacarus fulgens Summers et Schuster Apalotacarus fulgens Summers & Schuster, 1981a: 31 Type locality: Batavia [Djakarta], Java Distribution: Indonesia (Java) Host: Pelopides tridens (Coleoptera: Passalidae) Acarorum Catalogus IX 151 Apalotacarus fusticulus Summers et Schuster Apalotacarus fusticulus Summers & Schuster, 1981a: 31 Type locality: Ulong, NSW Distribution: Australia (NSW) Host: Pharochilus nitidulus (Coleoptera: Passalidae) Apalotacarus fusulus Summers et Schuster Apalotacarus fusulus Summers & Schuster, 1981a: 20 Type locality: Finschhafen, Papua New Guinea Distribution: Papua New Guinea, Philippines (Mindoro) Host: Labienus plox (Coleoptera: Passalidae) Apalotacarus glaber Summers et Schuster Apalotacarus glaber Summers & Schuster, 1981a: 25 Type locality: Russel Island Distribution: Solomon Islands Host: Protomocoelus triumphator (Coleoptera: Passalidae) Apalotacarus gracilis Summers et Schuster Apalotacarus gracilis Summers & Schuster, 1981a: 33 Type locality: 40 mi. SE Walcha, Stop 418, NSW, Australia Distribution: Australia Host: Coleoptera (Passalidae) Apalotacarus luroris Summers et Schuster Apalotacarus luroris Summers & Schuster, 1981a: 33 Type locality: Ulong, NSW Distribution: Australia (NSW, Queensland) Hosts: Mastochilus australasicus, M. quaestionis, Pharochilus dilatatus, Ph. nitidulus (Coleoptera: Passalidae) Apalotacarus paxillus Summers et Schuster Apalotacarus paxillus Summers & Schuster, 1981a: 21 Type locality: Wau, Morobe Distribution: Papua New Guinea Host: Coleoptera (Passalidae) Apalotacarus petilus Summers et Schuster Apalotacarus petilus Summers & Schuster, 1981a: 23 Type locality: Finschhafen Distribution: Papua New Guinea, incl. Bougainville I. Host: Protomocoelus australis (Coleoptera: Passalidae) 152 Acarorum Catalogus IX Apalotacarus protensus Summers et Schuster Apalotacarus protensus Summers & Schuster, 1981a: 28 Type locality: Kurseong, Darjeeling Distribution: India Host: Macrolinus nicobaricus (Coleoptera: Passalidae) Apalotacarus rigescens Summers et Schuster Apalotacarus rigescens Summers & Schuster, 1981a: 26 Type locality: Wau, Morobe Distribution: Papua New Guinea Host: Coleoptera (Passalidae) Apalotacarus sajanae Haitlinger Apalotacarus sajanae Haitlinger, 1990b: 536 Type locality: Java Distribution: Indonesia (Java) Host: Eriocnemis tridens (Coleoptera: Passalidae) Apalotacarus scaurus Summers et Schuster Apalotacarus scaurus Summers & Schuster, 1981a: 30 Type locality: Kabasalan, Mindanao Distribution: Philippines (Mindanao) Host: Macrolinus weberi (Coleoptera: Passalidae) Apalotacarus scissus Summers et Schuster Apalotacarus scissus Summers & Schuster, 1981a: 37 Type locality: Anomali Hills, Cinchona, S. India Distribution: India Host: Plurarius brachyphyllus (Coleoptera: Passalidae) Apalotacarus trullus Summers et Schuster Apalotacarus trullus Summers & Schuster, 1981a: 23 Type locality: Hassensan Distribution: Taiwan Host: Leptaulax bicolor formosanus (Coleoptera: Passalidae) Genus Arraphosoma Summers et Schuster Arraphosoma Summers & Schuster, 1981b: 43 Type species: Arraphosoma minax Summers & Schuster, 1981 Arraphosoma minax Summers et Schuster Arraphosoma minax Summers & Schuster, 1981b: 44 Acarorum Catalogus IX 153 Type locality: New Guinea Distribution: New Guinea Host: (Coleoptera: Passalidae) Arraphosoma paraminax Summers et Schuster Arraphosoma paraminax Summers & Schuster, 1981b: 44 Type locality: Mollucan Islands Distribution: Indonesia Host: Labiens gigas (Coleoptera: Passalidae) Genus Arrunsithiana Haitlinger Arrunsithiana Haitlinger, 1991e: 392 Type species: Arrunsithiana artemoni Haitlinger, 1991 Arrunsithiana arcuata (Fain) Grandiella arcuata Fain, 1987: 109 Grandiellina arcuata (Fain): Fain, 1989: 67 Arrunsithiana arcuata (Fain): Haitlinger, 1991e: 397 Type locality: Bahia, Santarem Distribution: Brazil Hosts: Stolas decemguttata, S. esignata (Coleoptera: Chrysomelidae) Arrunsithiana artemoni Haitlinger Arrunsithiana artemoni Haitlinger, 1991e: 393 Type locality: Ecuador Distribution: Ecuador Host: Stolas latevittata (Coleoptera: Chrysomelidae) Arrunsithiana battosi Haitlinger Arrunsithiana battosi Haitlinger, 1991e: 396 Type locality: Bolivia, Coroica Distribution: Bolivia, Brazil, Colombia, Ecuador, Peru Hosts: Stolas pleurosticha, S. lenis, S. oblita, S. erichsoni, S. inexculta, S. huanocensis, S. cruentata, S. scoparia, S. funebris, S. niobe, S. ignita, S. conspersa, S. gibbosa, S. paranensis, S. chalybaea (Coleoptera: Chrysomelidae) Arrunsithiana erinnae Haitlinger Arrunsithiana erinnae Haitlinger, 1991e: 394 Type locality: Santa Inez Distribution: Ecuador Host: Stolas pectinata (Coleoptera: Chrysomelidae) 154 Acarorum Catalogus IX Arrunsithiana esacaudata (Lombardini) Grandiella esacaudata Lombardini, 1939: 118; 1950: 784; Summers & Schuster, 1979a: 335; Fain, 1987: 109 Grandiellina esacaudata (Lombardini): Fain, 1989: 67 Arrunsithiana esacaudata (Lombardini): Haitlinger, 1991e : 398 Type locality: Brazil Distribution: Bolivia, Brazil, Costa Rica, French Guyana, Guatemala, Mexico Hosts: Omoplata marginata (Coleoptera: Cerambycidae), Phaeneum sp., Stolas punicea (Coleoptera: Chrysomelidae) Arrunsithiana ilosi Haitlinger Arrunsithiana ilosi Haitlinger, 1991e: 395 Type locality: Bogota Distribution: Colombia Host: Cyrtonota dissecta (Coleoptera: Chrysomelidae) Arrunsithiana latipes (Fain) Grandiellina latipes Fain, 1989: 67 Arrunsithiana latipes (Fain): Haitlinger, 1991e: 398 Type locality: South America Distribution: South America Host: Stolas inaequalis (Coleoptera: Chrysomelidae) Arrunsithiana quadrata (Fain) Grandiella quadrata Fain, 1987: 106 Grandiellina quadrata (Fain): Fain, 1989: 67 Arrunsithiana quadrata (Fain): Haitlinger, 1991e: 397 Type locality: Brazil, Bolivia (Santa Cruz), Colombia (Muzo) Distribution: Colombia, Bolivia, Brazil Host: Stolas aenea, S. indigacea, S.conspersa, S. chalybaea, S. funebris, Eugenysa columbiana (Coleoptera: Chrysomelidae) Arrunsithiana spicantis (Summers et Schuster) Grandiella spicantis Summers & Schuster, 1979a: 341; Fain, 1987: 94 Grandiellina spicantis (Summers et Schuster): Fain, 1989: 67 Arrunsithiana spicantis (Summers et Schuster): Haitlinger, 1991e : 397 Type locality: Santa Catarina Distribution: Brazil Hosts: Canistra plagosa, Stolas chalybaea, S. conspersa (Coleoptera: Chrysomelidae) Arrunsithiana squarrosa (Summers et Schuster) Grandiella squarrosa Summers & Schuster, 1979a: 338; Fain, 1987: 94 Acarorum Catalogus IX 155 Grandiellina squarosa (Summers et Schuster): Fain, 1989: 67 Arrunsithiana squarrosa (Summers et Schuster): Haitlinger, 1991e : 398 Type locality: Chanchamayo Valley Distribution: Peru, French Guiana Host: Stolas sp. (Coleoptera: Chrysomelidae) Genus Athogavia Haitlinger Athogavia Haitlinger, 1989d: 337 Type species: Athogavia tanzanica Haitlinger, 1989 Athogavia arybbasi Haitlinger Athogavia arybbasi Haitlinger, 1989d: 592 Type locality: Papua New Guinea Distribution: Papua New Guinea Host: Dynastinae (Coleoptera: Scarabaeidae) Athogavia assamica Haitlinger Athogavia assamica Haitlinger, 1989d: 348 Type locality: Assam Distribution: India Host: Trichogomphus nartolani (Coleoptera: Scarabaeidae, Dynastinae) Athogavia borneoica Haitlinger Athogavia borneoica Haitlinger, 1989d: 346 Type locality: Borneo Distribution: Borneo Host: Oryctes rhinoceros (Coleoptera: Scarabaeidae, Dynastinae) Athogavia gamana Haitlinger Athogavia gamana Haitlinger, 1989d: 342 Type locality: Tanzania Distribution: Tanzania Host: Oryctes monoceros (Coleoptera: Scarabaeidae, Dynastinae) Athogavia nosiana Haitlinger Athogavia nosiana Haitlinger, 1989: 340 Type locality: Madagascar Distribution: Madagascar Host: Oryctes (Coleoptera: Scarabaeidae, Dynastinae) Athogavia scagana Haitlinger Athogavia scagana Haitlinger, 1989d: 342 156 Acarorum Catalogus IX Type locality: Borneo Distribution: Borneo Host: Oryctes sp. (Coleoptera: Scarabaeidae, Dynastinae) Athogavia tanzanica Haitlinger Athogavia tanzanica Haitlinger, 1989d: 338 Type locality: Tanzania Distribution: Tanzania Host: Oryctes monoceros (Coleoptera: Scarabeidae, Dynastinae) Genus Aurilossongia Haitlinger Aurilossongia Haitlinger, 1989a: 43 Type species: Aurilossongia hoaria Haitlinger, 1989 Aurilossongia bolanica Haitlinger Aurilossongia bolanica Haitlinger, 1989a: 47 Type locality: Bolan Mts. Distribution: Papua New Guinea Host: Xylotrupes asperulus (Coleoptera: Cerambycidae) Aurilossongia dioni Haitlinger Aurilossongia dioni Haitlinger, 1989: 593 Type locality: Indonesia Distribution: Indonesia Host: Chalcosoma atlas (Coleoptera: Scarabaeidae, Dynastinae) Aurilossongia fileasi Haitlinger Aurilossongia fileasi Haitlinger, 1989: 593 Type locality: Papua New Guinea Distribution: Papua New Guinea Host: Scarabaeidae, Dynastinae (Coleoptera) Aurilossongia gaurana Haitlinger Aurilossongia gaurana Haitlinger, 1989a: 48 Type locality: New Britain, Rabaul Distribution: Papua New Guinea Host: ?Xylotrupes sp. (Coleoptera: Cerambycidae, Dynastinae) Aurilossongia hoaria Haitlinger Aurilossongia hoaria Haitlinger, 1989a: 44 Type locality: Borneo (Kalimantan) Acarorum Catalogus IX 157 Distribution: Indonesia (Kalimantan) Host: Xylotrupes sp. (Coleoptera: Cerambycidae, Dynastinae) Aurilossongia norae Haitlinger Aurilossongia norae Haitlinger, 1989: 594 Type locality: Papua New Guinea Distribution: Papua New Guinea Host: Dynastes gideon (Coleoptera: Scarabaeidae, Dynastinae) Genus Averithiana Haitlinger Averithiana Haitlinger, 1999: 42 Type species: Averithiana anoni Haitlinger, 1999 Averithiana anoni Haitlinger Averithiana anoni Haitlinger, 1999: 42 Type locality: French Guyana Distribution: French Guyana Host: Zatrephina lineata (Coleoptera: Chrysomelidae, Cassidinae) Genus Barbiangia Haitlinger Barbiangia Haitlinger, 1993a: 5 Type species: Barbiangia alvari Haitlinger, 1993 Barbiangia alvari Haitlinger Barbiangia alvari Haitlinger, 1993a: 6 Type locality: Zanzibar Distribution: Tanzania (Zanzibar) Host: undetermined Cetoniinae (Coleoptera: Scarabaeidae) Barbiangia elongata Khaustov Barbiangia elongata Khaustov, 2006: 80 Type locality: Ethiopia Distribution: Ethiopia Host: Diplognatha gagates (Coleoptera: Scarabaeidae, Cetoniinae) Barbiangia ethiopica (Haitlinger) Diplognatophilus ethiopicus Haitlinger, 1990: 321 Barbiangia ethiopica (Haitlinger): Haitlinger, 1993a: 7 Type locality: Ethiopia Distribution: Ethiopia Host: Scarabaeidae, Cetoniinae (Coleoptera) 158 Acarorum Catalogus IX Genus Beeriphotia Haitlinger Beeriphotia Haitlinger, 1994a: 207 Type species: Beeriphotia volkeri Haitlinger, 1994 Beeriphotia volkeri Haitlinger Beeriphotia volkeri Haitlinger, 1994a: 207 Type locality: Guandong (Kanton) Distribution: China Host: Carabus angulicollis antaeus (Coleoptera: Carabidae) Genus Bibulothiana Haitlinger Bibulothiana Haitlinger, 1993e: 278 Type species: Grandiella brevispinata Summers et Schuster, 1979 Bibulothiana albertae Haitlinger Bibulothiana albertae Haitlinger, 1994: 82 Type locality: Turrialba Distribution: Costa Rica Host: Chelymorpha comata (Coleoptera: Chrysomelidae, Cassidinae) Bibulothiana brevispinata (Summers et Schuster) Grandiella brevispinata Summers & Schuster, 1979: 346 Bibulothiana brevispinata (Summers et Schuster): Haitlinger, 1993e: 278 Type locality: 9 km N .El Cope, Panama Distribution: Bolivia, Brazil, Colombia, Honduras, Panama Hosts: Stolas bifenestrata, S. bipustulata, S. coerulescens, S. ephippium, S. lineaticollis, S. plagiata (Coleoptera: Chrysomelidae, Cassidinae) Bibulothiana cyriaki Haitlinger Bibulothiana cyriaki Haitlinger, 1993e: 280 Type locality: Parana, Iguacu Distribution: Brazil, Mexico Hosts: Chelymorpha infecta, Ch. nigricollis, Ch. vermiculata (Coleoptera: Chrysomelidae, Cassidinae) Bibulothiana everittae Haitlinger Bibulothiana everittae Haitlinger, 1993e: 281 Type locality: Santa Catarina (Brazil) Distribution: Brazil, Peru Hosts: Chelymorpha inflate, Ch. rufoguttata (Coleoptera: Chrysomelidae, Cassidinae) Acarorum Catalogus IX 159 Bibulothiana oswaldi Haitlinger Bibulothiana oswaldi Haitlinger, 1993e: 279 Type locality: Bahia Distribution: Brazil Host: Stolas plagicollis (Coleoptera: Chrysomelidae, Cassidinae) Bibulothiana vatia (Summers et Schuster) Grandiella vatia Summers & Schuster, 1979: 333 Bibulothiana vatia (Summers et Schuster): Haitlinger 1993e: 281 Type locality: Lago, Catemaco, Vera Cruz Distribution: Mexico Host: Cyrtonota tristigma (Coleoptera: Chrysomelidae, Cassidinae) Genus Bircericola Haitlinger Bircericola Haitlinger, 2000: 69 Type species: Bicercicola bertrami Haitlinger, 2000 Bircericola bertrami Haitlinger Bircericola bertrami Haitlinger, 2000: 70 Type locality: Bir Bon Ragba, nr. Hammamet Distribution: Tunisia Host: undetermined small Tenebrionidae (Coleoptera) Genus Boetophela Haitlinger Boetophela Haitlinger, 1989: 589 Type species: Boetophela ginae Haitlinger, 1989 Boetophela cassandrae Haitlinger et Chmielewski Boetophela cassandrae Haitlinger & Chmielewski, 2004: 196 Type locality: Durban Distribution: South Africa Host: undetermined Cetoniinae (Coleoptera: Scarabeidae) Boetophela erhardi Haitlinger Boetophela erhardi Haitlinger, 1991d: 304 Type locality: Nyangassy Distribution: Cameroon Host: Mecynorrhina [Chelorrhina] savagei (Coleoptera: Scarabeidae, Cetoniinae) Boetophela ephraimi Haitlinger Boetophela ephraimi Haitlinger, 1993: 12 Type locality: Ashanti 160 Acarorum Catalogus IX Distribution: Ghana Host: undetermined Cetoniinae (Coleoptera: Scarabaeidae) Boetophela ginae Haitlinger Boetophela ginae Haitlinger, 1989: 591; 1993: 10 Type locality: Natal, Transvaal [described ex errore from Sumatra, Batocera rubus, Cerambycidae, corrected by Haitlinger, 1993a] Distribution: Republic of South Africa Host: Eudicella smithi (Coleoptera: Scarabaeidae, Cetoniinae) Boetophela robertsoni (Lavoipierre) Cetonicola robertsoni Lavoipierre, 1958: 616 Boetophela robertsoni: Haitlinger, 1993: 14 Type locality: Tanzania Distribution: Tanzania Host: Dicronorhina oberthuri (Coleoptera: Scarabaeidae, Cetoniinae) Boetophela silvanae Haitlinger Boetophela silvanae Haitlinger, 1991d: 303 Type locality: Tanzania Distribution: Tanzania Host: Geynodonta laeviplaga (Coleoptera: Scarabaeidae, Cetoniinae) Boetophela sybillae Haitlinger Boetophela sybillae Haitlinger, 1991d: 299 Type locality: “surroundings of Nyassa lake” Distribution: Tanzania Host: Goliathus alborignathus (Coleoptera: Scarabaeidae, Cetoniinae) Boetophela symeoni Haitlinger Boetophela symeoni Haitlinger, 1991d: 300 Type locality: probably Tanzania Distribution: probably Tanzania Host: Elasocheilon heterospila (Coleoptera: Scarabeidae, Cetoniinae) Boetophela werneri Haitlinger Boetophela werneri Haitlinger, 1993a: 14 Type locality: Bujumbura Distribution: Burundi Host: Plaesiorrhina mhondana (Coleoptera: Scarabeidae, Cetoniinae) Acarorum Catalogus IX 161 Genus Boleohylla Haitlinger Boleohylla Haitlinger, 1991a: 274 Type species: Boleohylla kiogana Haitlinger, 1991 Boleohylla garrica Haitlinger Boleohylla garrica Haitlinger, 1991a: 276 Type locality: Ghana Distribution: Ghana Host: Metallonotus metallicus (Coleoptera: Tenebrionidae) Boleohylla kiogana Haitlinger Boleohylla kiogana Haitlinger, 1991a: 274 Type locality: nr. Lake Kiega Distribution: Uganda Host: Psammodes carbonarius (Coleoptera: Tenebrionidae) Genus Camirohylla Haitlinger Camirohylla Haitlinger, 1991a: 273 Type species: Camirohylla feziana Haitlinger, 1991 Camirohylla feziana Haitlinger Camirohylla feziana Haitlinger, 1991a: 274; 2001: 25 Type locality: Morocco Distribution: Morocco, Spain (Ibiza, Formentera) Host: Blaps appendiculata (Coleoptera: Tenebrionidae) Camirohylla sp. Type locality: Alashtar, Lorestan Province Distribution: Iran Host: Dila sp. (Coleoptera: Tenebrionidae) Genus Canestrinia Berlese Canestrinia Berlese, 1881: 747 Type species: Canestrinia dorcicola Berlese, 1881 Canestrinia carabicola (Berlese) Coleopterophagus carabicola Berlese, 1882: 10 Canestrinia (Percanestrinia) sardica Vitzthum, 1924: 41 Percanestrinia carabicola: Samšinak, 1965: Canestrinia carabicola: Samšinak, 1971: 176 Type locality: Italy 162 Acarorum Catalogus IX Distribution: Italy (incl. Sardinia and Sicily), Corsica, Tunisia Host: Carabus (Macrothorax) morbillosus (Coleoptera: Carabidae) Canestrinia damiani Haitlinger Canestrinia damiani Haitlinger, 1989: 353 Type locality: Hungary Distribution: Hungary, Ukraine, Crimea Host: Pentodon idiota (Coleoptera) Canestrinia dorcicola Berlese Canestrinia dorcicola Berlese, 1881: 748; Canestrini, 1897: 910; Canestrini & Kramer, 1899: 22; Leonardi, 1900: 15; Buitendijk, 1945: 364; Turk, 1953: 82; Cooreman, 1954: 86; Haitlinger, 1988a; 2004: 13; 2012: 186; Khaustov & Edelberg, 2001: 40; Trach, 2006: 176 Type locality: Italy Distribution: Great Britain, Holland, Italy, Poland, Ukraine, Crimea Hosts: Oxythyrea funesta (Coleoptera: Scarabaeidae, Cetoniinae), Dorcus parallelopipedus (Coleoptera: Lucanidae) Canestrinia hispanica Samšinák Canestrinia hispanica Samšinák, 1971: 173 Type locality: “Hispania” Distribution: Morocco, Portugal, Spain Hosts: Carabus (Rhabdotocarabus) melancholicus, C. (Ctenocarabus) galicianus, C. (Oreocarabus) guadarramus (Coleoptera: Carabidae). Canestrinia illegalis Samšinák Canestrinia illegalis Samšinák, 1971: 162 Type locality: Hualien (Taiwan) Distribution: Taiwan Hosts: Carabus (Coptolabrus) hankotaizanus, C. (Coptolabrus) longipennis mandarinus (Coleoptera: Carabidae) Canestrinia kacperi Haitlinger Canestrinia kacperi Haitlinger, 1998: 3 Type locality: Guanzhou Distribution: China Host: Carabus subformosus (Coleoptera: Carabidae) Canestrinia kamelskyi Samšinák Canestrinia kamelskyi Samšinák, 1970: 75 Type locality: Tunisia Acarorum Catalogus IX 163 Distribution: Tunisia Host: Blaps sp. (Coleoptera: Tenebrionidae) Canestrinia macgillavryi Oudemans Canestrinia macgillavryi Oudemans, 1923: 204; Cooreman, 1954: 86 Type locality: Soember Pakel Station, Dampit, Probolingo Distribution: Indonesia (Java)Canestrinia manicata Berlese Canestrinia manicata Berlese, 1910: 269 Type locality: Brazil Distribution: Brazil Host: Doryphora tessellata (Coleoptera: Chrysomelidae) Canestrinia microdisca Berlese Canestrinia microdisca Berlese, 1910: 269 Type locality: Java Distribution: Indonesia Host: Dorcus saiga (Coleoptera: Lucanidae) Canestrinia nepalensis Berlese Canestrinia nepalensis Berlese, 1910: 269 Type locality: Java Distribution: Indonesia Host: Dorcus nepalensis (Coleoptera: Lucanidae) Canestrinia paavoi Haitlinger Canestrinia paavoi Haitlinger, 1994a: 208 Type locality: South China, Tschekiang (= Zhejiangsheng) Distribution: China Host: Carabus grandis (Coleoptera: Carabidae) Canestrinia pekingensis Samšinák Canestrinia pekingensis Samšinák, 1971: 164 Type locality: Pekin Distribution: China Hosts: Carabus (Acathaicus) brandti, C. (Cychrostomus) anchocephalus (Tibet, 3200 m), C. (Eupachys) glyptopterus, C. (Coptolabrus) lafossei saturatus, C. augustus, C. augustus ignigena, C. pustulifer, C. pustulifer pratti, C. p. querryi, C. p. szechwanensis, C. ignimetalla angusticollis, C. gemmifer (Coleoptera: Carabidae) 164 Acarorum Catalogus IX Canestrinia pentodontis Berlese Canestrinia dorcicola var. pentodontis Berlese, 1897: Canestrinia pentodontis Berlese: Vitzthum, 1925: 47 Type locality: Italy Distribution: Italy, Sicily Host: Pentodon (Coleoptera: Scarabaeidae, Dynastinae) Canestrinia pictura Samšinák Canestrinia pictura Samšinák, 1971: 171 Type locality: Mt Kurama near Kyoto Distribution: Japan Host: Carabus (Damaster) blaptoides (Coleoptera: Carabidae) Canestrinia ramoni Haitlinger Canestrinia ramoni Haitlinger, 1994a: 212 Type locality: South China, Fukien (= Fujiansheng) Distribution: China Host: Carabus coelestis giganteus (Coleoptera: Carabidae) Canestrinia remigans Berlese Canestrinia remigans Berlese, 1910: 387 Type locality: Africa Distribution: Africa Genus Canestriniella Berlese Canestriniella Berlese, 1910: 270 Type species: Canestriniella amplexans Berlese, 1910 Canestriniella amplexans Berlese Canestriniella amplexans Berlese, 1910: 270 Type locality: “Batavia” (Jakarta) Distribution: Indonesia (Java) Host: Batocera hector (Coleoptera: Cerambycidae) Canestriniella batocerae (Vishnupriya et Mohanasundaram) Grandiella batocerae Vishnupriya & Mohanasundaram, 1988: 253 Grandiellina batocerae (Vishnupriya et Mohanasundaram): Fain, 1989: 67 Type locality: Tamil Nadu Distribution: India Host: Batocera rufomaculata (Coleoptera: Cerambycidae, Lamiinae) Acarorum Catalogus IX 165 Canestriniella elae Haitlinger Canestriniella elae Haitlinger, 1989: 599 Type locality: Sumatra Distribution: Indonesia Host: Batocera ferruginea (Coleoptera: Cerambycidae) Canestriniella phoebinae Haitlinger Canestriniella phoebinae Haitlinger, 1995a: 89 Type locality: Java Distribution: Indonesia Host: Batocera albofasciata octomaculata (Coleoptera: Cerambycidae) Canestriniella physsana Haitlinger Canestriniella physsana Haitlinger, 1989: 596 Type locality: Java Distribution: Indonesia (Java, Sulawesi) Hosts: Batocera celebiana, B. humilidens (Coleoptera: Cerambycidae) Canestriniella togata Berlese Canestriniella togata Berlese, 1910: 270 Type locality: Java Distribution: Indonesia (Java) Host: Batocera lunulata (Coleoptera: Cerambycidae) Genus Caraboecius Cooreman Caraboecius Cooreman, 1950: 18 Type species: Caraboecius coriacei Cooreman, 1950 Caraboecius coriacei Cooreman Caraboecius coriacei Cooreman, 1950: 19 Type locality: Tangelanga Distribution: Belgium Host: Carabus (Procrustes) coriaceus (Coleoptera: Carabidae) Genus Cetonicola Cooreman Cetonicola Cooreman, 1955: 15 Type species: Cetonicola hispidus Cooreman, 1955 Cetonicola hispidus Cooreman Cetonicola hispidus Cooreman, 1955: 16 Type locality: May-ya-Moto (Riv. Rutshuru) 166 Acarorum Catalogus IX Distribution: D.R. Congo Hosts: Dicronorhina micans, Smaragdesthes africana mutica, Eudicella gralli var. meckowi (Coleoptera: Scarabaeidae, Cetoniinae) Cetonicola isadorae Haitlinger Cetonicola isadorae Haitlinger, 1991d: 296 Type locality: Nyangassy Distribution: Cameroon Host: Mecynorrhina [Chelorrhina] savagei (Coleoptera: Scarabaeidae, Cetoniinae) Cetonicola maritae Haitlinger Cetonicola maritae Haitlinger, 1992d: 297 Type locality: Distr. Kassai Distribution: Congo-Brazzaville Host: Stephanorrhina guttata (Coleoptera: Scarabaeidae, Cetoniinae) Cetonicola orestesi Haitlinger Cetonicola orestesi Haitlinger, 1992d: 294 Type locality: Liberia Distribution: Ghana, Liberia Host: Tmesorrhina iris (Coleoptera: Scarabaeidae, Cetoniinae) Cetonicola vatus Haitlinger Cetonicola vatus Haitlinger, 1990 : 323 Type locality: Tanzania Distribution: Tanzania Host: Poecilophila hebraea (Coleoptera: Scarabaeidae, Cetoniinae) Genus Chelinochroa Summers et Schuster Chelinochroa Summers & Schuster, 1982b: 41 Type species: Chelinochroa dictyophora Summers et Schuster, 1982 Chelinochroa dictyophora Summers et Schuster Chelinochroa dictyophora Summers & Schuster, 1982b: 41 Type locality: Wajir Distribution: Kenya Host: Coleoptera: Cerambycidae Genus Cnecoderma Summers et Schuster Cnecoderma Summers & Schuster, 1981b: 39 Type species: Cnecoderma ovalis Summers et Schuster, 1981 Acarorum Catalogus IX 167 Cnecoderma dolichopoda Summers et Schuster Cnecoderma dolichopoda Summers & Schuster, 1981b: 41 Type locality: Queensland Distribution: Australia Host: Passalidae (Coleoptera) Cnecoderma ovalis Summers et Schuster Cnecoderma ovalis Summers & Schuster, 1981b: 39 Type locality: Australia Distribution: Australia Host: (Coleoptera: Passalidae) Genus Coleogyphus Berlese Coleogyphus Berlese, 1910: 268 Type species: Coleogyphus fuscipes Berlese, 1910 Coleoglyphus fuscipes Berlese Coleoglyphus fuscipes Berlese, 1910: 268 Type locality: Africa Distribution: Africa Hosts: Tefflus “megerlei” (Coleoptera: Carabidae). Nymph on Copris hamadryas (Coleoptera: Scarabaeidae) Coleoglyphus simplex Berlese Coleoglyphus (?) simplex Berlese, 1910b: 387 Type locality: Ecuador Distribution: “Equatore” (Ecuador) Host: Mitrocephala humboldti () Genus Coleopterophagus Berlese Coleopterophagus Berlese, 1882: 9 Type species: Dermoglyphus megninii Berlese, 1881 Coleopterophagus albini Haitlinger Coleopterophagus albini Haitlinger, 1990d: 328; 2002: 69; 2012: 187; 2015: 239; Trach & Khaustov, 2011: 223 Type locality: Zaborze n. Polczyn Zdroj (Poland) Distribution: Austria, Czech Rep., Estonia, Germany, Norway, Poland, Romania, Slovakia, Ukraine Hosts: Protaetia (Cetonischema) [Potosia] aeruginosa, P. (Liocola) marmorata, P. metallica (Coleoptera: Scarabaeidae, Cetoniinae) 168 Acarorum Catalogus IX Coleopterophagus baali Haitlinger Coleopterophagus baali Haitlinger, 1990d: 330 Type locality: North Iran, Elburs Mts. Distribution: Iran Host: Potosia speciosa (Coleoptera: Scarabaeidae, Cetoniinae) Coleopterophagus belzebubi Haitlinger Coleopterophagus belzebubi Haitlinger, 1990d: 332 Type locality: Taihorim Distribution: Taiwan Host: Potosia aerata (Coleoptera: Scarabaeidae, Cetoniinae) Coleopterophagus blattophagus (Banks) Canestrinia blattophaga Banks, 1914: 61 Coleopterophagus blattophagus (Banks): Vitzthum, 1925: 48 Type locality: Panama Distribution: Panama Coleopterophagus dionizyi Haitlinger Coleopterophagus dionizyi Haitlinger, 1990d: 330 Type locality: Syria Distribution: Syria Host: Potosia cuprea (Coleoptera: Scarabaeidae, Cetoniinae) Coleopterophagus donaldi Haitlinger Coleopterophagus donaldi Haitlinger, 1990d: 332; Trach & Khaustov, 2011: 223 Type locality: Bolzano (Italy) Distribution: Italy, Ukraine Host: Protaetia (Eupotosia) [Potosia] affinis (Coleoptera: Scarabaeidae, Cetoniinae) Coleopterophagus maroni Haitlinger Coleopterophagus maroni Haitlinger, 1990d: 336; Trach & Khaustov, 2011: 220 Type locality: Syria Distribution: Romania, Syria, Ukraine Hosts: Protaetia (Cetonischema) [Potosia] aeruginosa, P. speciosa jousselini (Coleoptera: Scarabaeidae, Cetoniinae) Coleopterophagus megnini (Berlese) Dermoglyphus megnini Berlese, 1881: 750 Dermoglyphus megninii Berlese: Radford, 1950: 161 Acarorum Catalogus IX 169 Coleopterophagus megninii (Berlese): Berlese, 1882: A.M.S., fasc. I, No 9; Turk, 1953: 82; Oudemans, 1923 : 205; 1937: 2155; Buitendijk, 1945: 364; Haitlinger, 1988d: 535; 2002: 69; 2012: 186 Coleopterophagus megnini (Berlese): Trach & Khaustov, 2011: 214; Haitlinger, 2012: 186 Coleopterophagus cetoniae Oudemans, 1923; Buitendijk, 1945: 264; Cooreman, 1954: 85 Type locality: Italy Distribution: Croatia, Czech Rep., Great Britain, Holland, Hungary, Italy, Sicily, Poland, Serbia, Turkey, Ukraine Hosts: Cetonia aurata, Protaetia (Potosia) cuprina, Protaetia (Potosia) fieberi, Protaetia (Potosia) metallica, Protaetia cuprea cuprea, P. cuprea ignicollis, P. cuprea obscura, Protaetia (Eupotosia) affinis (Coleoptera: Scarabaeidae, Cetoniinae) Coleopterophagus neglectus Berlese Coleopterophagus neglectus Berlese, 1920: 147 Type locality: Africa Distribution: Africa Host: “Scarabeus” centaurus (Coleoptera: Scarabaeidae) Coleopterophagus pulcher Cooreman Coleopterophagus pulcher Cooreman, 1955: 23 Type locality: Rutshuru Distribution: D.R. Congo Host: Porphyronota cinnamonea (Coleoptera: Scarabaeidae, Cetoniinae) Coleopterophagus quadrisetosus Trägårdh Coleopterophagus quadrisetosus Trägårdh, 1904: 113 Type locality: Gebelein Distribution: Sudan Host: Coleoptera (Scarabaeidae, Cetoniinae) Coleopterophagus rudolfi Haitlinger Coleopterophagus rudolfi Haitlinger, 1990d: 334 Type locality: China Distribution: China Host: Potosia brevitarsis (Coleoptera: Scarabaeidae, Cetoniinae) Genus Cooremania Nesbitt Cooremania Nesbitt, 1976: 227 Type species: Cooremania wauensis Nesbitt, 1976 170 Acarorum Catalogus IX Cooremania kietaensis Haitlinger et Šundić Cooremania kietaensis Haitlinger & Šundić, 2020: 155 Type locality: Kieta Distribution: Papua New Guinea Host: Coleoptera (Lucanidae indet.) Cooremania wauensis Nesbitt Cooremania wauensis Nesbitt, 1976: 227 Type locality: Wau, Njau Distribution: Papua New Guinea Host: Eurytrachus arfakianus (Coleoptera: Lucanidae) Genus Danaithiana Haitlinger Danaithiana Haitlinger, 1991b: 253 Type species: Danaithiana chanchamayoica Haitlinger, 1991 Danaithiana archidonaica Haitlinger Danaithiana archidonaica Haitlinger, 1991b: 257 Type locality: Archidona (Ecuador) Distribution: Colombia, Ecuador Host: Semiotus furcatus (Coleoptera: Elateridae) Danaithiana ayraica Haitlinger Danaithiana ayraica Haitlinger, 1991b: 254 Type locality: Laguayra (Ecuador) Distribution: Colombia, Ecuador Host: Semiotus virgatus (Coleoptera: Elateridae) Danaithiana chanchamayoica Haitlinger Danaithiana chanchamayoica Haitlinger, 1991b: 254 Type locality: Chanchamayo (Peru) Distribution: Costa Rica, Cuba, Peru Host: Semiotus imperialis (Coleoptera: Elateridae) Danaithiana ecuadorica Haitlinger Danaithiana ecuadorica Haitlinger, 1991b: 257 Type locality: Archidona Distribution: Ecuador Host: Semiotus furcatus (Coleoptera: Elateridae) Acarorum Catalogus IX 171 Genus Dicanestrinia Berlese Dicanestrinia Berlese, 1911: 184 Type species: Dermaleichus cerambycis G.Canestrini, 1878 Dicanestrinia cerambycis (G. Canestrini) Dermaleichus cerambycis G. Canestrini, 1878: 69 Canestrinia (Dicanestrinia) cerambicis [sic]: Berlese, 1911: 184 Type locality: Italy Distribution: Italy Host: Cerambyx cerdo (Coleoptera: Cerambycidae) Dicanestrinia huberti Haitlinger Dicanestrinia huberti Haitlinger, 1994: 200; 2012: 186; Khaustov & Edelberg, 2001: 40 Type locality: Bystre n. Baligród (Bieszcady Mountains – voi. Krosno)(Poland) Distribution: Byelorussia, Poland Host: Carabus variolosus (Coleoptera: Carabidae) Dicanestrinia knobi Samšinák Dicanestrinia knobi Samšinák, 1971: 177; Haitlinger, 1988a: 535; 2012: 186; Trach, 2006: 175 Type locality: Carpathia, Užok (Ukraine) Distribution: Czech Rep., Bosnia and Herzegovina, Italy, Poland, Ukraine Host: Carabus variolosus, C.v. nodulosus (Coleoptera: Carabidae) Genus Diplognathophilus Cooreman Diplognathophilus Cooreman, 1955: 14 Type species: Diplognathophilus africanus Cooreman, 1955 (56?) Diplognathophilus africanus Cooreman Diplognathophilus africanus Cooreman, 1955: 14 Type locality: Ndeko (Katanda) Distribution: D.R. Congo Host: Rhabdotis sobrina (Coleoptera: Scarabaeidae, Cetoniinae) Genus Diplopodocoptes Fain Diplopodocoptes Fain, 1987: 167 Type species: Diplopodocoptes transkeiensis Fain, 1987 Diplopodocoptes transkeiensis Fain Diplopodocoptes transkeiensis Fain, 1987: 168 Type locality: Transkei, District Lusikisiki, Mouth of Riv. Mzimhlava Distribution: Kenya, South Africa 172 Acarorum Catalogus IX Hosts: millipede (Diplopoda: Glomeridae); paratypes from Odontopygidae (Diplopoda) from Kenya Genus Donnelaphontia Lavoipierre Donnelaphontia Lavoipierre, 1959: 611 Farahanella Haitlinger, 1990: 314; [Type is suatoha; Syn. Haitlinger 2001: 104] Type species: Donnelaphontia calostegis Lavoipierre, 1959 Donnelaphontia calostegis Lavoipierre Donnelaphontia calostegis Lavoipierre, 1959: 611 Type locality: Kumba Distribution: Cameroon Host: Calostega crassicornis (Coleoptera: Tenebrionidae) Donnelaphontia suatoha (Haitlinger) Farahanella suatoha: Haitlinger, 1990: 315 Donnelaphontia suatoha: Haitlinger, 2001: 104 Type locality: “Zaire” Distribution: DR Congo Host: Odontopezus cupreus (Coleoptera: Tenebrionidae) Genus Dorynothiana Haitlinger Dorynothiana Haitlinger, 1999: 43 Type species: Dorynothiana jurandi Haitlinger, 1999 Dorynothiana delmari Haitlinger Dorynothiana delmari Haitlinger, 2008: 66 Type locality: Brazil Distribution: Brazil Host: Dorynota sp. (Coleoptera: Chrysomelidae, Cassidinae) Dorynothiana jurandi Haitlinger Dorynothiana jurandi Haitlinger, 1999: 44 Type locality: Santa Catarina Distribution: Brazil Host: Dorynota pugionata (Coleoptera: Chrysomelidae, Cassidinae) Genus Ennodithiana Haitlinger Ennodithiana Haitlinger, 1993d: 66 Type species: Ennodithiana messalinae Haitlinger, 1993 Acarorum Catalogus IX 173 Ennodithiana maniusi Haitlinger Ennodithiana maniusi Haitlinger, 1993d: 68 Type locality: Brazil, Parana Distribution: Brazil, Mexico Hosts: Omaspides basilica, Psalidonota contemta, P. dorsoplagiata (Coleoptera: Chrysomelidae, Cassidinae) Ennodithiana medoni Haitlinger Ennodithiana medoni Haitlinger, 1993d: 71 Type locality: Brazil Distribution: Brazil Host: Acromis spinifex (Coleoptera: Chrysomelidae, Cassidinae) Ennodithiana megesi Haitlinger Ennodithiana megesi Haitlinger, 1993d: 71 Type locality: Goyaz Distribution: Brazil Host: Crambelea illuderis (Coleoptera: Chrysomelidae, Cassidinae) Ennodithiana messalinae Haitlinger Ennodithiana messalinae Haitlinger, 1993d: 66 Type locality: San Carlos Distribution: Costa Rica, Cuba Hosts: Psalidonota dorsopunctata, P. leprosa (Coleoptera: Chrysomelidae, Cassidinae) Ennodithiana penelopae Haitlinger Ennodithiana penelopae Haitlinger, 1993: 68 Type locality: Paramaribo (Suriname) Distribution: Brazil, Suriname Host: Acromis nebulosa (Coleoptera: Chrysomelidae, Cassidinae) Ennodithiana platysma (Summers et Schuster) Grandiella platysma Summers & Schuster, 1979: 335 Ennodithiana platysma (Summers & Schuster): Haitlinger, 1993d: 73 Type locality: Barro Colorado Isl. Distribution: Panama Host: Coptocycla sp. (Coleoptera: Chrysomelidae, Cassidinae) Genus Eunorithiana Haitlinger Eunorithiana Haitlinger, 1999: 46 Type species: Eunorithiana aidae Haitlinger, 1999 174 Acarorum Catalogus IX Eunorithiana aidae Haitlinger Eunorithiana aidae Haitlinger, 1999: 47; 2001: 103 Type locality: Rio Cantino Distribution: Colombia Host: Plagiometriona phoebe (Coleoptera: Chrysomelidae, Cassidinae) Eunorithiana heliodori Haitlinger Eunorithiana heliodori Haitlinger, 1999: 49 Type locality: Santa Cruz, Santa Catarina Distribution: Brazil Host: Plagiometriona ludiera, P. punctipennis (Coleoptera: Chrysomelidae, Cassidinae) Genus Gasuthiana Haitlinger Gasuthiana Haitlinger, 1989: 281 Type species: Gasuthiana abapoica Haitlinger, 1989 Gasuthiana abapoica Haitlinger Gasuthiana abapoica Haitlinger, 1989: 282 Type locality: Brazil Distribution: Brazil Host: Alurnus elysianus (Coleoptera: Chrysomelidae, Hispinae) Gasuthiana abufarica Haitlinger Gasuthiana abufarica Haitlinger, 1989: 287 Type locality: Bolivia Distribution: Bolivia Host: Meristomala nigripes (Coleoptera: Chrysomelidae, Hispinae) Gasuthiana coarica Haitlinger Gasuthiana coarica Haitlinger, 1989: 289 Type locality: Brazil Distribution: Brazil Host: Mecistomala marginatus (Coleoptera: Chrysomelidae, Hispinae) Gasuthiana melillae Haitlinger Gasuthiana melillae Haitlinger, 1995a: 83 Type locality: Brazil Distribution: Brazil Host: Tenebrionidae indet. (Coleoptera) Gasuthiana olareica Haitlinger Gasuthiana olareica Haitlinger, 1989: 284 Acarorum Catalogus IX 175 Type locality: Colombia Distribution: Colombia, French Guiana Hosts: Alurnus bipunctatus, A. boulardi (Coleoptera: Chrysomelidae, Hispinae) Genus Gioharattia Haitlinger Gioharattia Haitlinger, 1989a: 57 Type species: Gioharattia maharae Haitlinger, 1989 Gioharattia caroae Haitlinger Gioharattia caroae Haitlinger, 1989a: 59 Type locality: “South Ceylon” Distribution: Sri Lanka Host: Coryphocera sp. (Coleoptera: Scarabaeidae, Cetoniinae) Gioharattia maharae Haitlinger Gioharattia maharae Haitlinger, 1989a: 57 Type locality: Borneo Distribution: Borneo (?Indonesia) Host: Coglodera diaroi (Coleoptera: Scarabaeidae, Cetoniinae) Genus Globosophotia Haitlinger Globosophotia Haitlinger, 1991a: 279 Type species: Globosophotia bettinae Haitlinger, 1991 Globosophotia bettinae Haitlinger Globosophotia bettinae Haitlinger, 1991a: 279 Type locality: Morocco Distribution: Morocco Host: Pseudosteropus globosus (Coleoptera: Carabidae, Pterostichinae) Genus Grandiella Lombardini Grandiella Lombardini, 1939: 45 [considered by Fain, 1989 praeocc. from Williams 1928 Hymenoptera, but being nomen nudum, Grandiella Lombardini has been restored by OConnor, 2004] Grandiellina Fain, 1989: 67 Type species: Grandiella tetracaudata Lombardini, 1939 Grandiella cooremani Fain Grandiella cooremani Fain, 1987: 106; OConnor, 2004: 120 Grandiellina cooremani (Fain): Fain, 1989: 67 Type locality: Ecuador 176 Acarorum Catalogus IX Distribution: Ecuador Host: Canistra osculatii (Coleoptera: Chrysomelidae) Grandiella decemcaudata Lombardini Grandiella decemcaudata Lombardini, 1950: 289; OConnor, 2004: 120 Grandiellina decemcaudata (Lombardini): Fain, 1989: 67 Type locality: Bolivia, Zapirangai Distribution: Bolivia, Paraguay Hosts: Echoma [as Omoplata] marginata, Coptocycla contemta (Coleoptera: Chrysomelidae, Cassidinae) Grandiella ecuadorensis Fain Grandiella ecuadorensis Fain, 1987: 101; OConnor, 2004: 120 Grandiellina ecuadorensis (Fain): Fain, 1989: 67 Type locality: Ecuador Distribution: Ecuador Host: Canistra osculatii (Coleoptera: Chrysomelidae) Grandiella gambosa Summers et Schuster Grandiella gambosa Summers & Schuster, 1979: 343; Fain, 1987: 94 Grandiellina gambosa (Summers & Schuster): Fain, 1989: 67 Type locality: El Llano Carti Rd., 10 km N El Llano, Pina, Panama Distribution: Belize, Colombia, Panama Hosts: Discomorpha batesi, D. lanuginosa pubescens, D. depilata, Dolichotoma conspersipennis (Coleoptera: Chrysomelidae) Grandiella joliveti Fain Grandiella joliveti Fain, 1987: 97 Grandiellina joliveti (Fain): Fain, 1989: 67 Type locality: Vigosa, Minas Gerais Distribution: Brazil Host: Elytrosphaera xanthopyga (Coleoptera: Chrysomelidae) Grandiella kimseyi Summers et Schuster Grandiella kimseyi Summers & Schuster, 1979a: 349; Fain, 1987: 94; OConnor, 2004: 120 Grandiellina kimseyi (Summers et Schuster): Fain, 1989: 67 Type locality: El Llano Carti Rd., Pina (Panama) Distribution: Belize, Colombia, Panama Hosts: Discomorpha batesi, Dolichotoma conspersipennis (Coleoptera: Chrysomelidae) Acarorum Catalogus IX 177 Grandiella longini (Haitlinger) Grandiellina longini Haitlinger, 1991e: 389 Grandiella longini (Haitlinger): OConnor, 2004: 120 Type locality: Bolivia Distribution: Bolivia Host: Goniochenia (Baranosa) flavosparsa (Coleoptera: Chrysomelidae, Cassidinae) Grandiella marinae (Haitlinger) Grandiellina marinae Haitlinger, 1991e: 390 Grandiella marinae (Haitlinger): OConnor, 2004: 120 Type locality: Chiapas Distribution: Mexico Host: Ehoma confluens (Coleoptera: Chrysomelidae) Grandiella minor Fain Grandiella minor Fain, 1987: 106; OConnor, 2004: 120 Grandiellina minor (Fain): Fain, 1989: 67 Type locality: Santarem Distribution: Brazil Host: Canistra dohrni (Coleoptera: Chrysomelidae) Grandiella octocaudata Lombardini Grandiella octocaudata Lombardini, 1950: 288; OConnor, 2004: 120 Grandiellina octocaudata (Lombardini): Fain, 1989: 67 Type locality: Estado de Paraná, Bariqui Distribution: Brazil Host: Stolas chalybaea (Coleoptera: Chrysomelidae) Grandiella pentagona Lombardini Grandiella pentagona Lombardini, 1950: 286; OConnor, 2004: 120 Grandiellina pentagona (Lombardini): Fain, 1989: 67 Type locality: Distrito Federal, Cerro de Naiguatá Distribution: Venezuela Host: Coleoptera: Chrysomelidae, Cassidinae Grandiella rogeri (Haitlinger) Grandiellina rogeri Haitlinger, 1991e: 388 Grandiella rogeri (Haitlinger): OConnor, 2004: 120 Type locality: Chanchamayo Distribution: Peru Host: Goniochenia (Baranosa) heroldi (Coleoptera: Chrysomelidae, Cassidinae) 178 Acarorum Catalogus IX Grandiella rosascostai (Lombardini) Coleoglyphus rosascostai Lombardini, 1950: 279 Hatohylla rosascostai (Lombardini): Haitlinger 1991: 311 Grandiella rosascostai (Lombardini): OConnor, 2004: 120 Type locality: Distrito Federal Cerro del Naiguatá Distribution: Venezuela Host: Nyctobates sp. (Coleoptera: Tenebrionidae) Grandiella rugosita Summers et Schuster Grandiella rugosita Summers & Schuster, 1979: 343; OConnor, 2004: 120 Type locality: Panama Distribution: Panama Host: Coleoptera Grandiellina sicula Summers et Schuster Grandiella sicula Summers & Schuster, 1979a: 333; OConnor, 2004: 120 Grandiellina sicula (Summers & Schuster): Fain, 1989: 67; Haitlinger, 1991e: 392 Type locality: Depto. Del Cesar, 0.3 km N Castilla, Inez Ranch Distribution: Colombia Hosts: Coptocycla nigropunctata, C. nr. wagneri (Coleoptera: Chrysomelidae, Cassidinae) Grandiella soniae (Haitlinger) Grandiellina soniae Haitlinger, 1991e: 391 Grandiella soniae (Haitlinger): OConnor, 2004: 120 Type locality: Rio Negro Distribution: Colombia Hosts: Cyrtonota punicea, C. textilis, Stolas coalita, S. coerulescens, S. erichsoni, S. placida (Coleoptera: Chrysomelidae) Grandiella tacjannae (Haitlinger) Grandiellina tacjannae Haitlinger, 1991e: 383 Grandiella tacjannae (Haitlinger): OConnor, 2004: 120 Type locality: Brazil, Candido de Abreu Distribution: Argentina, Brazil, Paraguay, Peru Hosts: Anacassis fuscata, Botanochara angulata, B. bonariensis, B. convexiuscula, B. duodecimverrucata, B. impressa, B. macularia, B. nervosa, B. regina, B. ruforeticulata, B. sanguinea, B. sedecimpustulata, B. tesselata, B. texta (Coleoptera: Chrysomelidae, Cassidinae) Grandiella tertia Lombardini Grandiella tertia Lombardini, 1942: 46; OConnor, 2004: 120 Acarorum Catalogus IX 179 Grandiellina tertia (Lombardini): Fain, 1989: 67 Type locality:Brazil Distribution: Brazil Host: Cassids (Coleoptera: Chrysomelidae) Grandiella tetracaudata Lombardini Grandiella tetracaudata Lombardini, 1939: 45; 1950: 283; Summers & Schuster, 1979a: 330; Fain, 1987: 94; OConnor, 2004: 120 Grandiellina tetracaudata (Lombardini): Fain, 1989: 67; Haitlinger, 1991e : 383 Type locality: Brazil Distribution: Brazil, Colombia, Peru Hosts: [Botanochara sedecimpustulata, B. subnervosa], Coptocycla adamantina, Cyrtonota cyanea, C. thalassina, C. tigrina, Eumolpus fulgidus, Stolas chalybaea, S. subrugosa (Coleoptera: Chrysomelidae). According to Haitlinger (1992e), the mites recorded on genus Botanochara belong actually to Grandiellina tacjannae. Grandiella tetracaudata omoplata Fain Grandiella tetracaudata omoplata Fain, 1987 : 94; OConnor, 2004: 120 Grandiellina tetracaudata omoplata (Fain): Fain, 1989: 67 Type locality: Cayenne, St. Jean de Maroni, Gourdonville Distribution: French Guyana Host: Eichoma clypeata (= Omoplata marginata) (Coleoptera: Chrysomelidae, Cassidinae) Genus Grandiellopsis Fain Grandiellopsis Fain, 1989: 68 Type species: Grandiellopsis cyclosoma Fain, 1989 Grandiellopsis alani Haitlinger Grandiellopsis alani Haitlinger, 1992: 52 Type locality: Archidona Distribution: Ecuador Host: Canistra osculatii (Coleoptera: Chrysomelidae) Grandiellopsis ariani Haitlinger Grandiellopsis ariani Haitlinger, 1992: 53 Type locality: Campirias Distribution: Brazil Host: Canistra carbonaria (Coleoptera: Chrysomelidae) Grandiellopsis canistra Fain Grandiellopsis canistra Fain, 1989: 70 180 Acarorum Catalogus IX Type locality: Ecuador Distribution: Ecuador Host: Canistra osculatii (Coleoptera: Chrysomelidae) Grandiellopsis cyclosoma Fain Grandiellopsis cyclosoma Fain, 1989: 68 Type locality: Bas Maroni Distribution: French Guyana Host: Cyclosoma palliata (Coleoptera: Chrysomelidae) Grandiellopsis odiosi Haitlinger Grandiellopsis odiosi Haitlinger, 1992: 56 Type locality: Brazil Distribution: Brazil Host: Polychalca (P.) punctatissima (Coleoptera: Chrysomelidae) Grandiellopsis orsenae Haitlinger Grandiellopsis orsenae Haitlinger, 1992: 58 Type locality: Puerto Inca Distribution: Peru Host: Cyclosoma mirabilis (Coleoptera: Chrysomelidae) Grandiellopsis violae Haitlinger Grandiellopsis violae Haitlinger, 1992: 55 Type locality: Bolivia Distribution: Bolivia Host: Canistra cruentata (Coleoptera: Chrysomelidae) Genus Haitlingeria Wontae Kim et al. Haitlingeria Kim et al., 2006: 13 Type species: Haitlingeria longilobata Kim et al., 2006 Haitlingeria longilobata Kim et al. Haitlingeria longilobata Kim et al., 2006: 13 Type locality: South Korea, near the Bung-Dong Reservoir Seonbukcong Yuseongku Daejeon Distribution: Korea Host: Serrognathus platymelus (Coleoptera: Lucanidae) Genus Hargeinella Haitlinger Hargeinella Haitlinger, 1989a: 53 Acarorum Catalogus IX 181 Type species: Hargeinella poppana Haitlinger, 1989 Hargeinella poppana Haitlinger Hargeinella poppana Haitlinger, 1989a: 53 Type locality: Sumatra Distribution: Indonesia (Sumatra) Host: Trichogomphus milon (Coleoptera: Scarabaeidae, Dynastinae) Genus Hatohylla Haitlinger Hatohylla Haitlinger, 1990a: 307 1991 ? Type species: Hatohylla morrata Haitlinger, 1990 Hatohylla bahiana Haitlinger Hatohylla bahiana Haitlinger, 1990a: 308 Type locality: Santa Catarina Distribution: Brazil Host: Nyctobates maximus (Coleoptera: Tenebrionidae) Hatohylla bibiannae Haitlinger Hatohylla bibiannae Haitlinger, 1990a: 311 Type locality: Santa Catarina Distribution: Brazil Host: undeternined Tenebrionidae (Coleoptera) Hatohylla morrata Haitlinger Hatohylla morrata Haitlinger, 1990a: 308 Type locality: Amazonas Distribution: Brazil Host: Nyctobates gigas (Coleoptera: Tenebrionidae) Genus Hyllosihemia Haitlinger Hyllosihemia Haitlinger, 1994: 93 Type species: Hyllosihemia belerofoni Haitlinger, 1994 Hyllosihemia belerofoni Haitlinger Hyllosihemia belerofoni Haitlinger, 1994: 93 Type locality: Sumatra Distribution: Indonesia (Sumatra) Host: Coleoptera: Passalidae Genus Hypopteryx Summers et Schuster Hypopteryx Summers & Schuster, 1981b: 41 182 Acarorum Catalogus IX Type species: Hypopteryx collaris Summers et Schuster, 1981 Hypopteryx collaris Summers et Schuster Hypopteryx collaris Summers & Schuster, 1981b: 42 Type locality: Australia Distribution: Australia Host: Coleoptera Genus Ibizella Haitlinger et Šundić Ibizella Haitlinger & Šundić, 1916: 75 Type species: Ibizella balearica Haitlinger & Šundić, 1916 Ibizella asilahica Haitlinger et Šundić Ibizella asilahica Haitlinger et Šundić, 1918: 413 Type locality: Asilah (31 km south of Tangier) Distribution: Morocco Host: Coleoptera: Tenebrionidae indet. Ibizella balearica Haitlinger et Šundić Ibizella balearica Haitlinger & Šundić, 1916: 75 Type locality: Ibiza Distribution: Spain (Ibiza) Host: (Coleoptera, Tenebrionidae: Blaps sp.) Ibizella samsinaki (Beron) Canestrinia samsinaki Beron, 1975: 83; Haitlinger, 1993c: 87; 2001a: 26; Haitlinger & Šundić, 1916: 75 Ibizella samsinaki (Beron): Haitlinger & Šundić, 1916: 75 Type locality: Bulgaria, Sozopol Distribution: Bulgaria, Cyprus, Spain (Ibiza) Host: Blaps sp., Gnaptor sp. (Coleoptera: Tenebrionidae) Genus Irmongia Haitlinger Irmongia Haitlinger, 1990b: 547 Type species: Irmongia helgae Haitlinger, 1990 Irmongia helgae Haitlinger Irmongia helgae Haitlinger, 1990b: 547 Type locality: Tanzania Distribution: Tanzania Host: undetermined Passalidae (Coleoptera) Acarorum Catalogus IX 183 Genus Jowitella Haitlinger Jowitella Haitlinger, 1990b: 547 Type species: Jowitella barbarae Haitlinger, 1990 Jowitella barbarae Haitling Jowitella barbarae Haitlinger, 1990b: 547 Type locality: Santa Catarina Distribution: Brazil Host: undetermined Passalidae (Coleoptera) Genus Jullongia Haitlinger Jullongia Haitlinger, 1990b: 538 Type species: Jullongia izae Haitlinger, 1990 Jullongia izae Haitlinger Jullongia izae Haitlinger, 1990b: 539 Type locality: Stephansort Distribution: Papua New Guinea Host: undetermined Passalidae (Coleoptera) Genus Kahoorangia Haitlinger Kahoorangia Haitlinger, 1992: 288 Type species: Kahoorangia almae Haitlinger, 1992 Kahoorangia almae Haitlinger Kahoorangia almae Haitlinger, 1991d: 289 Type locality: ? South Africa Distribution: ? South Africa Host: Pachnoda carmelita (Coleoptera: Scarabaeidae, Cetoniinae) Genus Lidiophela Haitlinger Lidiophela Haitlinger, 1990b: 540 Type species: Lidiophela pelagiae Haitlinger, 1990 Lidiophela aruica Haitlinger Lidiophela aruica Haitlinger, 1990b: 542 Type locality: Aru Is. Distribution: Indonesia (Aru Is.) Host: Labienus ptox (Coleoptera: Passalidae) Lidiophela bognae Haitlinger Lidiophela bognae Haitlinger, 1990b: 543 184 Acarorum Catalogus IX Type locality: Moluccas, Batjan Is. Distribution: Indonesia (Moluccas, Batjan Is.) Host: undetermined Passalidae (Coleoptera) Lidiophela celinae Haitlinger Lidiophela celinae Haitlinger, 1990b: 541 Type locality: Moluccas, Ambon Is. Distribution: lndonesia, Moluccas (Ambon Is.) Host: undetermined Passalidae (Coleoptera) Lidiophela igae Haitlinger Lidiophela igae Haitlinger, 1990b: 542 Type locality: Moluccas, Batjan Is. Distribution: Indonesia (Batjan Is.) Host: Labienus gigas (Coleoptera: Passalidae) Lidiophela pauliana Haitlinger Lidiophela pauliana Haitlinger, 1991a: 276 Type locality: Australia Distribution: Australia Host: Amarygmus striatus (Coleoptera: Tenebrionidae) Lidiophela pecki (Nesbitt) Coleopterophagus pecki Nesbitt, 1976: 220 Lidiophela pecki: Haitlinger, 1990b: 540 Type locality: Baiyer River Sanctuary, Mt.Hagen, Papua New Guinea Distribution: Papua New Guinea, Solomon Islands (Gizo Is.), Hosts: Labienus sp., Obiensus sp., Protomocoelus australis (Coleoptera: Passalidae) Lidiophela pelagiae Haitlinger Lidiophela pelagiae Haitlinger, 1990b: 540 Type locality: Moluccas, Batjan Is. Distribution: Indonesia, Moluccas (Batjan Is.) Host: Labienus gigas (Coleoptera: Passalidae) Genus Megacanestrinia Trägårdh Megacanestrinia Trägårdh, 1906: 877 Type species: Megacanestrinia mucronata Trägårdh, 1906 Megacanestrinia beloniana Haitlinger Megacanestrinia beloniana Haitlinger, 1990e: 312 Type locality: Tanzania Acarorum Catalogus IX 185 Distribution: Tanzania Host: Tefflus purpuripennis (Coleoptera: Carabidae) Megacanestrinia mucronata Trägårdh Megacanestrinia mucronata Trägårdh, 1906: 877 Type locality: Umfolozi, Zululand Distribution: South Africa Host: «Auf einen grossen Carabide« (Coleoptera: Carabidae) Subfam. Melisiinae Summers et Schuster Melisiinae Summers & Schuster, 1982a: 23 Type genus : Melisia Lombardini, 1944 Genus Melisia Lombardini Melisia Lombardini, 1944: 2 Type species: Melisia melisii Lombardini, 1944 Melisia anomala Samšinák Melisia anomala Samšinák, 1963: 257 Type locality: Mts Singalang Distribution: Indonesia (Sumatra) Hosts: Epiphanus indicus, E. stoliczkae (Coleoptera: Passalidae) Melisia ariadnae Haitlinger Melisia ariadnae Haitlinger, 1989: 529 Type locality: Soekaranda (Sumatra); Borneo Distribution: lndonesia (Sumatra, Borneo) Host: Aceraius laevicollis (Passalidae, Coleoptera) Melisia baloghi Samšinák Melisia baloghi Samšinák, 1969: 407 Type locality: Cuc-phuong Bong Distribution: Vietnam Host: Coleoptera indet. Melisia gabrielae Haitlinger Melisia gabrielae Haitlinger, 1990: 531 Type locality: Soekaranda, Sumatra Distribution: Indonesia (Sumatra) Host: Aceraius laevicollis (Coleoptera: Passalidae) 186 Acarorum Catalogus IX Melisia helenae Haitlinger Melisia helenae Haitlinger, 1990b: 533 Type locality: North Vietnam Distribution: Vietnam Host: undetermined Passalidae (Coleoptera) Melisia hieronimae Haitlinger Melisia hieronimae Haitlinger, 1990b: 535 Type locality: holotype from North Vietnam; paratypes from Thailand and Sumatra Distribution: Indonesia (Sumatra), Thailand, Vietnam Host: undetermined Passalidae (Coleoptera) Melisia ingeborgae Haitlinger Melisia ingeborgae Haitlinger, 1990b: 535 Type locality: Soekaranda, Sumatra Distribution: Indonesia (Sumatra) Host: undetermined Passalidae (Coleoptera) Melisia irenae Haitlinger Melisia irenae Haitlinger, 1990b: 531 Type locality: Sinabong, Sumatra Distribution: Indonesia (Sumatra) Host: Aceraius grandis (Coleoptera: Passalidae) Melisia jadwigae Haitlinger Melisia jadwigae Haitlinger, 1990b: 532 Type locality: North Vietnam Distribution: Vietnam Host: undetermined Passalidae (Coleoptera) Melisia klapaleki Samšinák Melisia klapaleki Samšinák, 1963: 260 Type locality: India mer.: Coimbatore Distribution: India Host: Epiphenus stoliczkae (Coleoptera: Passalidae) Melisia leptinotarsa Summers et Schuster Melisia leptinotarsa Summers & Schuster, 1982a: 25 Type locality: Paluma, Stop 388, Queensland Distribution: Australia Host: Coleoptera Passalidae (gen. sp. Indet.) Acarorum Catalogus IX 187 Melisia lombardinii Samšinák Melisia lombardinii Samšinák, 1963: 255 Type locality: Mts Singalang Distribution: Indonesia (Sumatra) Host: Pelopides mniszecki (Coleoptera: Passalidae) Melisia mahunkai Samšinák Melisia mahunkai Samšinák, 1969: 409 Type locality: Cuc-phuong, Bong Distribution: Vietnam Host: Coleoptera indet. Melisia mariettae Haitlinger Melisia mariettae Haitlinger, 1990b: 528 Type locality: Sinabong, Sumatra Distribution: Indonesia (Sumatra) Host: Aceraius grandis (Coleoptera: Passalidae) Melisia melaniae Haitlinger Melisia melaniae Haitlinger, 1990b: 532 Type locality: Sumatra Distribution: Indonesia (Sumatra) Host: Aceraius mayeri (Coleoptera: Passalidae) Melisia melisii Lombardini Melisia melisii Lombardini, 1944: 2; Samšinák, 1963: 253 Type locality: America Distribution: America Host: Passalus sp. (Coleoptera: Passalidae) Melisia occidii Samšinák Melisia occidii Samšinák, 1969: 410; Haitlinger, 1990b: 532 Type locality: Cuc-phuong, Bong Distribution: Vietnam Host: Coleoptera indet. Melisia rutae Haitlinger Melisia rutae Haitlinger, 1990b: 533 Type locality: North Vietnam Distribution: Vietnam Host: undetermined Passalidae (Coleoptera) 188 Acarorum Catalogus IX Melisia salomeae Haitlinger Melisia salomeae Haitlinger, 1990b: 529 Type locality: Sinabong, Sumatra Distribution: Indonesia (Sumatra) Host: Aceraius grandis (Coleoptera: Passalidae) Melisia samsinaki Vomero Melisia samsinaki Vomero, 1972: 95 Type locality: Kon Techore Distribution: Cambodja Host: Aceraius helferi (Coleoptera: Passalidae) Melisia sentosa Summers et Schuster Melisia sentosa Summers & Schuster, 1982a: 27 Type locality: Ta Rau, Hainan Distribution: China (Hainan Isl.) Host: Passalidae gen. sp. indet. (Coleoptera) Melisia superba Summers et Schuster Melisia superba Summers & Schuster, 1982a: 27 Type locality: Musha, Taiwan Distribution: Taiwan Host: Aceraius grandis hirsutus (Coleoptera: Passalidae) Genus Mesophotia Samšinák Mesophotia Samšinák, 1971: 209 Type species: Mesophotia penicillata Samšinák, 1971 Mesophotia penicillata penicillata Samšinák Mesophotia penicillata penicillata Samšinák, 1971: 209 Type locality: Sevilla Distribution: Spain Host: Carabus (Rhabdotocarabus) melancholicus, C.m. costatus (Coleoptera: Carabidae) Mesophotia penicillata similis Samšinák Mesophotia penicillata similis Samšinák, 1971: 214 Type locality: Andalusia Distribution: Spain Host: Carabus (Rhabdotocarabus) melancholicus costatus (Coleoptera: Carabidae) Acarorum Catalogus IX 189 Mesophotia penicillata tertia Samšinák Mesophotia penicillata tertia Samšinák, 1971: 214 Type locality: Sevilla Distribution: Morocco, Spain Host: Carabus (Rhabdotocarabus) melancholicus (Coleoptera: Carabidae) Genus Metriothiana Haitlinger Metriothiana Haitlinger, 1999: 40 Type species: Metriothiana bolebori Haitlinger, 1999 Metriothiana bolebori Haitlinger Metriothiana bolebori Haitlinger, 1999: 40 Type locality: Tegusigalpa Distribution: Honduras Host: Metrionella bilimeki (Coleoptera: Chrysomelidae, Cassidinae) Genus Mossongia Haitlinger Mossongia Haitlinger, 1989a: 49 Type species: Mossongia ssavina Haitlinger, 1989 Mossongia bissaina Haitlinger Mossongia bissaina Haitlinger, 1989a: 52 Type locality: Bougainville, Kieta Distribution: Papua New Guinea Host: Scapanus solomonensis (Coleoptera: Scarabaeidae, Dynastinae) Mossongia ssavina Haitlinger Mossongia ssavina Haitlinger, 1989a: 50 Type locality: New Guinea Distribution: New Guinea Host: Scapanus australis (Coleoptera: Scarabaeidae, Dynastinae) Genus Neophotia Samšinák Neophotia Samšinák, 1971: 201 Type species: Neophotia drvotocka Samsinák, 1971 Neophotia drvotocka Samšinák Neophotia drvotocka Samšinák, 1971: 202 Type locality: San Ildefonso Distribution: Portugal, Spain Hosts: Carabus (Iniopachus) pyrenaeus, C. (Oreocarabus) amplipennis, C. (Oreocarabus) guadarramus (Coleoptera: Carabidae) 190 Acarorum Catalogus IX Neophotia jureceki Samšinák Neophotia jureceki Samšinák, 1971: 204 Type locality: Pyrenees, Pic du Midi Distribution: France Host: Carabus (Iniopachus) pyrenaeus (Coleoptera: Carabidae) Neophotia latissima Samšinák Neophotia latissima Samšinák, 1971: 206 Type locality: Guadarrama, Cercedilla Distribution: Spain Host: Carabus (Mesocarabus) lusitanicus (Coleoptera: Carabidae) Genus Noemiphela Haitlinger Noemiphela Haitlinger, 1992b: 536 Type species: Noemiphela izabelae Haitlinger, 1992 Noemiphela barryi Haitlinger Noemiphela barryi Haitlinger, 1992b: 538 Type locality: Sumatra Distribution: Indonesia Host: Eurytrachelus purpurescens (Coleoptera: Lucanidae) Noemiphela izabelae Haitlinger Noemiphela izabelae Haitlinger, 1992b: 537 Type locality: Celebes (Sulawesi) Distribution: Indonesia Host: Cladognathus giraffa (Coleoptera: Lucanidae) Genus Oceogavia Haitlinger Oceogavia Haitlinger, 1989: 586 Type species: Oceogavia ubaldi Haitlinger, 1989 Oceogavia ubaldi Haitlinger Oceogavia ubaldi Haitlinger, 1989: 586 Type locality: unknown Distribution: unknown Host: Papuana woodlarkiana (Coleoptera: Scarabaeidae, Dynastinae) Genus Olafithiana Haitlinger Olafithiana Haitlinger, 1993e: 274 Type species: Olafithiana adelinae Haitlinger, 1993 Acarorum Catalogus IX 191 Olafithiana adelinae Haitlinger Olafithiana adelinae Haitlinger, 1993e: 274 Type locality: Parana Distribution: Brazil Host: [Stolas] Anacassis cribrum (Coleoptera: Chrysomelidae, Cassidinae) Olafithiana foliacea (Fain) Grandiella foliacea Fain, 1987: 101 Grandiellina foliacea (Fain): Fain, 1989: 67 Olafithiana foliacea (Fain): Haitlinger 1993e: 277 Type locality: Brazil, Nova Teutonia Distribution: Argentina, Brazil, Panama Hosts: Stolas areolata, S. chalybea, S. conspersa, S. complexa, S. festiva, S. impluviata, S. lacunose, S. lenis, S. nudicollis, S. oblita, S. paranensis, S. subreticulata (Coleoptera: Chrysomelidae, Cassidinae) Olafithiana iwettae Haitlinger Olafithiana iwettae Haitlinger, 1994b: 83 Type locality: Costa Rica Distribution: Costa Rica, Venezuela Host: Charidotella (= Metriona) testudinacea (Coleoptera: Chrysomelidae, Cassidinae) Olafithiana joliveti (Fain) Grandiella joliveti Fain, 1987: 97 Olafithiana joliveti (Fain): Haitlinger 1993e: 278 Type locality: Viçosa, Minas Gerais Distribution: Brazil Host: Elytrosphaera xanthopyga (Coleoptera: Chrysomelidae) Olafithiana multifida (Fain) Grandiella multifida Fain, 1987: 101 Olafithiana multifida (Fain): Haitlinger, 1993e: 276 Type locality: Brazil Distribution: Argentina, Bolivia, Brazil, Ecuador Hosts: Stolas antiqua, S. aurosetosa, S. cruentata (Coleoptera: Chrysomelidae, Cassidinae) Genus Olgattia Haitlinger Olgattia Haitlinger, 1990b: 538 Type species: Olgattia useguaica Haitlinger, 1990 192 Acarorum Catalogus IX Olgattia useguaica Haitlinger Olgattia useguaica Haitlinger, 1990b: 538 Type locality: Usegua Distribution: Tanzania Host: Eumelasomus sansibaricus (Coleoptera: Passalidae) Genus Ovarrothiana Haitlinger Ovarrothiana Haitlinger, 1993d: 60 Type species: Ovarrothiana nikodemi Haitlinger, 1993 Ovarrothiana echinoderma (Summers et Schuster) Grandiella echinoderma Summers & Schuster, 1979: 346; Fain, 1987: 94 Grandiellina echinoderma (Summers & Schuster): Fain, 1989: 67 Ovarrothiana echinoderma (Summers & Schuster): Haitlinger, 1993d: 65 Type locality: Cerro Compana, Panama Distribution: Colombia, Panama Host: Eugenysa colombiana (Coleoptera: Chrysomelidae, Cassidinae) Ovarrothiana erazmi Haitlinger Ovarrothiana erazmi Haitlinger, 1993d: 61 Type locality: Cayenne Distribution: French Guyana Host: Eugenysa venosa (Coleoptera: Chrysomelidae, Cassidinae) Ovarrothiana leopoldi Haitlinger Ovarrothiana leopoldi Haitlinger, 1993d: 65 Type locality: Yungaz de La Paz Distribution: Bolivia Host: Miocalaspis gentilis (Coleoptera: Chrysomelidae, Cassidinae) Ovarrothiana ludomiri Haitlinger Ovarrothiana ludomiri Haitlinger, 1993d: 64 Type locality: Weise Distribution: Bolivia Host: Cyclosoma tristis (Coleoptera: Chrysomelidae, Cassidinae) Ovarrothiana nikodemi Haitlinger Ovarrothiana nikodemi Haitlinger, 1993d: 60 Type locality: Peru Distribution: Brazil, Peru Host: Eugenysa divalis (Coleoptera: Chrysomelidae, Cassidinae) Acarorum Catalogus IX 193 Ovarrothiana ozannae Haitlinger Ovarrothiana ozannae Haitlinger, 1993d: 63 Type locality: Peru, Chanchamayo Distribution: Brazil, Ecuador, Peru Hosts: Agenyssa caedemadens, A. crassicornis, Eugenysa andicola (Coleoptera: Chrysomelidae, Cassidinae) Genus Paracanestrinia Haitlinger Paracanestrinia Haitlinger, 2017: 658 Type species: Paracanestrinia denmarkica Haitlinger, 2017 Paracanestrinia denmarkica Haitlinger Paracanestrinia denmarkica Haitlinger, 2017: 659 Type locality: 11 km north of Hvidbjerg Distribution: Denmark Host: Carabus sp. (Coleoptera: Carabidae) Genus Paramansia Cooreman Paramansia Cooreman, 1950: 13 Type species: Paramansia menthastri Cooreman, 1950 Paramansia bicornis Khaustov et Eidelberg Paramansia bicornis Khaustov & Eidelberg, 2001: 43; Trach & Khaustov, 2007: 268 Type locality: Crimea, Yalta Distribution: Crimea, Ukraine Hosts: Chrysolina menthastri, Ch. polita, Ch. fastuosa (Coleoptera: Chrysomelidae) Paramansia giardi (Trouessart) Canestrinia giardi Trouessart, in Berlese, 1887: Paramansia giardi (Trouessart): Cooreman, 1950: 18 Type locality: ? Brazil Distribution: ? Brazil Host: ? Megistomela reticulata (Coleoptera: Chrysomelidae) Paramansia menthastri Cooreman Paramansia menthastri Cooreman, 1950: 14 Type locality: Bray-sous-Fay nr. Richelieu, Indre et Loire, France Distribution: France Host: Chrysolina menthastri (Coleoptera: Chrysomelidae) Genus Paramelisia Summers et Schuster Paramelisia Summers & Schuster, 1982a: 29 194 Acarorum Catalogus IX Type species: Paramelisia lacuna Summers & Schuster, 1982 Paramelisia lacuna Summers et Schuster Paramelisia lacuna Summers & Schuster, 1982a: 29 Type locality: Anomal Hills, Cinchona, S. India Distribution: India Host: Pleurarius brachyphyllus (Coleoptera: Passalidae) Genus Paraphagella Cooreman Paraphagella Cooreman, 1955: 28 Type species: Paraphagella princeps Cooreman, 1955 Paraphagella eudicellae Cooreman Paraphagella eudicellae Cooreman, 1955: 33 Type locality: Kibga, Sud Visoke, 2400 m; Rutshuru Distribution: D.R. Congo Hosts: Eudicella smithi var. bertherandi, E. gralli var. meckowi (Coleoptera: Chrysomelidae) Paraphagella indirae Haitlinger Paraphagella indirae Haitlinger, 1991d: 291 Type locality: Nyangassy Distribution: Cameroon Host: Mecynorrhina [Chelarrhina – recte Chelorrhina] savagei (Coleoptera) Paraphagella ingridae Haitlinger Paraphagella ingridae Haitlinger, 1991d: 290 Type locality: Liberia Distribution: Liberia Host: Tmesorrhina iris (Coleoptera: Scarabaeidae, Cetoniinae) Paraphagella minor Cooreman Paraphagella minor Cooreman, 1955: 36 Type locality: Kibga, Sud Visoke, 2400 m Distribution: D.R. Congo Host: Eudicella smithi var. bertherandi (Coleoptera: Chrysomelidae) Paraphagella odae Haitlinger Paraphagella odae Haitlinger, 1991d: 292 Type locality: Distr. Kassai Distribution: Congo (Brazzaville) Host: Stephanorrhina guttata (Coleoptera: Scarabaeidae, Cetoniinae) Acarorum Catalogus IX 195 Paraphagella princeps Cooreman Paraphagella princeps Cooreman, 1955: 29 Type locality: May-ya-Moto, riv. Rutshuru Distribution: D.R. Congo Hosts: Dicronorhina micans, Smaragdesthes africana mutica (Coleoptera: Chrysomelidae) Genus Paraphotia Khaustov et Eidelberg Paraphotia Khaustov & Eidelberg, 2001: 37 Type species: Paraphotia calosomae Khaustov et Eidelberg, 2001 Paraphotia calosomae Khaustov et Eidelberg Paraphotia calosomae Khaustov & Eidelberg, 2001: 39; Trach, 2006: 175 Type locality: Crimea, Opuk Mount. Distribution: Crimea Host: Calosoma auropunctatum (Coleoptera: Carabidae) Genus Passalophagus Nesbitt Passalophagus Nesbitt, 1976: 230 Type species: Passalophagus georgei Nesbitt, 1976 Passalophagus georgei Nesbitt Passalophagus georgei Nesbitt, 1976: 231 Type locality: Baiyer River Bird of Paradise Sanctuary, Mt. Hagen Area Distribution: Papua New Guinea Host: Labienus sp. (Coleoptera: Passalidae) Genus Percanestrinia Berlese Canestrinia (Percanestrinia) Berlese, 1911: 184 Percanestrinia Berlese: Samšinák, 1965 (syn. with Canestrinia); Haitlinger, 1992a: 535 (restored as genus) Type species: Alloptes blaptis Canestrini et Berlese, 1880 Percanestrinia blaptis (Canestrini et Berlese) Alloptes blaptis Canestrini & Berlese, 1880: 148 Percanestrinia blaptis: Berlese, 1911: 184; Turk, 1953: 82; Rack, 1968: 265; Haitlinger, 1992a: 541; 2012: 187; Trach, 2006: 176 Canestrinia blaptis: Berlese, 1884: A.M.S., fasc. XXIX, No1; Haitlinger, 1992a: 541 Coleopterophagus blaptis (Canestrini et Berlese): Oudemans, 1923: 187 Percanestrinia (Percanestrinia) blaptis: Cooreman, 1954: 85 Canestrinia blaptis rackae Samšinák, 1970: 72 (syn. by Haitlinger, 1992a) Type locality: ? „Italia septentrionalis“ 196 Acarorum Catalogus IX Distribution: Bulgaria, Georgia, Germany, Great Britain, Hungary, Italy (incl. Sardinia), Poland, Ukraine Hosts: Blaps gibbosa, B. lethifera, B. occulta, B. punctata (Coleoptera: Tenebrionidae), Sphodrus leucophthalmus (Coleoptera: Carabidae) Percanestrinia egypti El Bishlawi et Allam Percanestrinia egypti El Bishlawi & Allam, 2003: 267 Type locality: Egypt Distribution: Egypt Percanestrinia izabelae Haitlinger Percanestrinia izabelae Haitlinger, 1992a: 536 Type locality: Cyprus Distribution: Cyprus Host: Blaps gibbosa (Coleoptera: Tenebrionidae) Percanestrinia maroccana Cooreman Percanestrinia maroccana Cooreman, 1953: 6; Haitlinger, 1992a: 540 Type locality: Morocco Distribution: Corsica (France), Italy, Morocco Hosts: Blaps mucronata, B. pinguis, B. producta (Coleoptera: Tenebrionidae) Percanestrinia norodomi Haitlinger Percanestrinia norodomi Haitlinger, 1992a: 538 Type locality: Tashkent Distribution: Uzbekistan Host: Blaps hejconota cortosa (Coleoptera: Tenebrionidae) Percanestrinia sellnicki Samšinák Percanestrinia sellnicki Samšinák, 1965: 145; Beron, 1971: 205; Haitlinger, 1988d, 2012: 188; Canestrinia sellnicki: Samsinak, 1971: 155; Haitlinger, 1989; Trach, 2006: 176 Type locality: “Hungaria (Com. Bihar)” (now in Romania) Distribution: Czech Rep., Hungary, former Jugoslavia, Moldova, Poland, Romania, Slovakia, Ukraine Hosts: Carabus (Procrustes) coriaceus rugifer, C. c. coriaceus, C. coriaceus pseudorugifer, C. coriaceus banaticus, C. coriaceus subrugosus, C. coriaceus gressorius, C. coriaceus imminutus, C. violaceus (Coleoptera: Carabidae) Percanestrinia viviannae Haitlinger Percanestrinia viviannae Haitlinger, 1992a: 540 Type locality: China Acarorum Catalogus IX 197 Distribution: China Host: Tenebrionidae (Coleoptera) Genus Percanestriniella Khaustov et Eidelberg Percanestriniella Khaustov & Eidelberg, 2001: 34 Type species: Percanestriniella caucasica Khaustov et Eidelberg, 2001 Percanestriniella caucasica Khaustov et Eidelberg Percanestriniella caucasica Khaustov & Eidelberg, 2001: 34 Type locality: Caucasus, Mzymsha river bank near Kardyvach Distribution: Russia (Caucasus) Host: Carabus sp. (Coleoptera: Carabidae) Percanestriniella levis (Samšinák) Canestrinia levis Samšinák, 1971: 166 Percanestriniella levis: Khaustov & Eidelberg, 2001: 36 Type locality: ‘Turkestan’(Sussamyr – GB, Ketmen-Tjube), Buchara Distribution: Kirghizstan, Uzbekistan Hosts: Carabus (Pantophyrtus) turcomannorum, C.(P.) tanypedilus, C. (Cratocephalus) balassogloi, C. (C.) cicatricosus, C. (Ulocarabus) sturowskyi, C. (Cratocechenus) akinini (Coleoptera: Carabidae) Percanestriniella vlachiana (Samšinák) Canestrinia vlachiana Samšinák, 1971: 169 Percanestriniella vlachiana Khaustov & Eidelberg, 2001: 34 Type locality: Jalloori, Kulu (Kashmir) Distribution: India (Kashmir) Hosts: Carabus (Imaibius) boysi, C. (I.) stoliczkanus (Coleoptera: Carabidae) Genus Peruthiana Haitlinger Peruthiana Haitlinger, 2008: 69 Type species: Peruthiana briannae Haitlinger, 2008 Peruthiana briannae Haitlinger Peruthiana briannae Haitlinger, 2008: 69 Type locality: Peru Distribution: Peru Host: undetermined Hispinae (Coleoptera: Chrysomelidae) Genus Phaleratus Nesbitt Phaleratus Nesbitt, 1976: 233 Type species: Phaleratus fentoni Nesbitt, 1976 198 Acarorum Catalogus IX Phaleratus alatus Summers et Schuster Phaleratus alatus Summers & Schuster, 1982a: 29 Type locality: Anomali Hills, Madras Distribution: India Host: Pleurarius brachyphyllus (Coleoptera: Passalidae) Phaleratus fentoni Nesbitt Phaleratus fentoni Nesbitt, 1976: 233 Type locality: Baiyer River Bird of Paradise Sanctuary, Mt. Hagen Area Distribution: Papua New Guinea Host: Coleoptera Genus Phelliculophela Haitlinger Phelliculophela Haitlinger, 1993a: 9 Type species: Phelliculophela roaldi Haitlinger, 1993 Phelliculophela roaldi Haitlinger Phelliculophela roaldi Haitlinger, 1993a: 9 Type locality: Cameroon Distribution: Cameroon Host: Goliathus goliathus (Coleoptera : Scarabaeidae, Cetoniinae) Genus Photia Oudemans Photia Oudemans, 1904: 192 Type species: Canestrinia procustidis Berlese, 1881 Eucanestrinia Berlese, 1911: 184 Type species: “Canestrinia procrusti” Photia adolfinae Haitlinger Photia adolfinae Haitlinger, 1988 (sub “polymorpha”; 1994a: 202) Photia adolphinae Haitlinger: Khaustov & Edelberg, 2001: 30; Trach, 2006: 175 Type locality: Narewka, voi. Bialystok, and many other localities in Poland Distribution: Byelorussia, Poland, Crimea Hosts: Carabus (C.) glabratus, C. gyllenhali (Coleoptera: Carabidae) Photia bardoica Haitlinger Photia bardoica Haitlinger, 1988: 321; 1995b: 261; 2012: 189 Type locality: Poland Distribution: Austria, Poland Host: Carabus nemoralis (Coleoptera: Carabidae) Acarorum Catalogus IX 199 Photia bilkorum Samšinák Photia bilkorum Samšinák, 1971: 199; Haitlinger, 2012: 190 Type locality: Abhasia Distribution: Abhasia, Russia Hosts: Carabus (Tribax) protensis, C. (T.) komarovi, C. (T.) shamyli, C. (T.) plasoni (Coleoptera: Carabidae) Photia chrysocarabi Cooreman Photia chrysocarabi Cooreman, 1950: 34; Samšinák, 1971: 193; Haitlinger, 1988d; 1995b: 262; 2012: 188; Trach, 2006: 176 Type locality: Arlon (Belgium) Distribution: Austria, Belgium, Czech Rep., England, France, Germany, Poland, Slovakia, Ukraine Hosts: Carabus auratus, C. irregularis, C. nemoralis, C. scheidleri, Chrysocarabus auronitens (Coleoptera: Carabidae) Photia graeca Cooreman Photia graeca Cooreman, 1958: 1; Samšinák, 1965: 143; 1971: 189 Type locality: Philothée nr. Athens Distribution: Belgium, England, France, Greece (incl. Rhodos), Holland, Turkey Hosts: Carabus c. coriaceus, C. coriaceus foudrasi, C. c. kindermanni, C. c. impudicus, C. violaceus purpurascens (Coleoptera: Carabidae) Photia hejniana Samšinák Photia hejniana Samšinák, 1971: 193; Haitlinger, 1988d; 1995b: 261; 2012: 189; Trach, 2006: 176 Type locality: Sobotka Distribution: Austria, Croatia, Czech Rep., Moldova, Poland, Slovakia, Slovenia, Ukraine Hosts: Carabus alpestris hoppei, C. coriaceus, C. croaticus, C. violaceus, C. v. exasperates, C. v. fioris, C. v. obliquus, C. v. savinicus, C. v. styriensis (Coleoptera: Carabidae) Photia hermengildae Haitlinger Photia hermengildae Haitlinger, 1988: 319; 2012: 189 Type locality: Huta Szklana, Gory Swietokrzyskie, Voivodeship Kielce Distribution: Poland, Ukraine Host: Carabus linnaei (Coleoptera: Carabidae) Photia howdeni Nesbitt Photia howdeni Nesbitt, 1976: 223 Type locality: Baiyer River Bird of Paradise Sanctuary, Mt. Hagen Area Distribution: Papua New Guinea Host: Coleoptera 200 Acarorum Catalogus IX Photia javensis (Oudemans) Canestrinia javensis Oudemans, 1923b: 203 Photia javensis (Oudemans) Type locality: Java, Soember Pakel, Probolingo Distribution: Indonesia (Java) Host: Coleoptera Photia lopatini Khaustov et Eidelberg Photia lopatini Khaustov & Eidelberg, 2001: 26 Type locality: Sakhalin Is., Krasnoselskaya river bank Distribution: Russia (Sakhalin) Host: Carabus (Acoptolabrus) lopatini (Coleoptera: Carabidae) Photia lusitanica Samšinak Photia lusitanica Samšinak, 1971: 189 Type locality: Sierra de Queija, Prov. Orense, Spain Distribution: Portugal, Spain Hosts: Carabus (Oreocarabus) amplipennis, Carabus (Mesocarabus) lusitanicus breuningi, C. (M.) lusitanicus helluo, C.(M.) lusitanicus castilianus, C. (M.) lusitanicus logronicus, C. (M.) lusitanicus schaumi (Coleoptera: Carabidae) Photia melchiori Haitlinger Photia melchiori Haitlinger, 1998: 2; 2012: 191 Type locality: China Distribution: China Host: Carabus sp. (Coleoptera: Carabidae) Photia pacifica Khaustov et Eidelberg Photia pacifica Khaustov & Eidelberg, 2001: 28 Type locality: Kuril Is. (Kunashir Is., Golovnin volcano) Distribution: Russia (Kuril Is.) Host: Carabus (Megodontus) kolbei (Coleoptera: Carabidae) Photia polymorpha Samšinák Photia polymorpha Samšinak, 1971: 184; Khaustov & Eidelberg, 2001: 31; Trach, 2006: 175; Haitlinger, 2012: 191 Type locality: Ulbinskoe, Siberia Distribution: Japan, Korea, Russia (Siberia), Ukraine, Crimea Host: holotype from Carabus (Megodontus) imperialis (Coleoptera: Carabidae). Reported also from other species of the genus Carabus, but, according to Khaustov & Eidelberg (2001), the mites collected from them may represent other species. Acarorum Catalogus IX 201 Photia procera (Berlese) Canestrinia (Eucanestrinia) procera Berlese, 1911: 184 Photia procera procera (Berlese): Samšinák, 1964: 35 Photia procera (Berlese): Cooreman, 1950: 26; Beron, 1971: 205; Haitlinger, 2012: 192 Type locality: “Carniolia” (Slovenia) Distribution: Bulgaria, Slovenia (“Carniolia”), Italy Hosts: Carabus (Procerus) caucasicus, C. (P.) gigas, C. (P.) scabrosus, Carabus (Procrustes) coriaceus (Coleoptera: Carabidae) Photia procera adanensis Samšinák Photia procera adanensis Samšinák, 1964: 36 Type locality: Adana Distribution: Turkey Host: Carabus (Procerus) scabrosus transversalis (Coleoptera: Carabidae) Photia procera marani Samšinák Photia procera mařani Samšinák, 1964: 37 Type locality: Lebanon Distribution: Lebanon Host: Carabus (Procerus) syriacus (Coleoptera: Carabidae) Photia procera turcica Samšinák Photia procera turcica Samšinák, 1964: 35 Type locality: Turkey Distribution: Turkey Host: Carabus (Procerus) scabrosus caucasicus (Coleoptera: Carabidae) Photia procustidis (Berlese) Canestrinia procustidis Berlese, 1881: 749 Canestrinia procrusti Berlese, 1892; Canestrini, 1897 Photia procrusti (Berlese): Oudemans, 1904: 192 Photia procustidis (Berlese): Oudemans, 1904; 1923b: 203; Samšinák, 1965: 140; 1971; Haitlinger, 2012: 190 Eucanestrinia procrusti Berlese, 1911: 184 Eucanestrinia procrustidis Vitzthum, 1925: 48 Type locality: Italy Distribution: Albania, Cyprus, Greece (incl. Crete, Rhodes), Israel, Italy, Lebanon, Slovakia, Turkey, former Yugoslavia Hosts: Carabus anatolicus, C. coriaceus, C. impressus (Coleoptera: Carabidae) 202 Acarorum Catalogus IX Photia saetolata (Cooreman) Percanestrinia saetolata Cooreman, 1950: 42; 1954: 85 Photia saetolata (Cooreman): Samšinák, 1965: 145; 1971: 197 Type locality: Belgium Distribution: Belgium, Czech Rep., France, Germany, Great Britain, Holland Hosts: Carabus (Procrustes) c. coriaceus, C. violaceus purpurascens, C.v. exasperaus, C.v. bicinctus, C.v. fulgens (Coleoptera: Carabidae) Photia sibirica Khaustov et Eidelberg Photia sibirica Khaustov & Eidelberg, 2001: 23 Type locality: Novosibirskaya Obl., Togoochinskiy Reg., Kotorovo Village Distribution: Russia Host: Carabus (Megodontus) schoenherri (Coleoptera: Carabidae) Genus Phriknodora Summers et Schuster Phriknodora Summers & Schuster, 1982b: 43 Type species: Phriknodora pustulosa Summers & Schuster, 1982 Phriknodora avae Haitlinger Phriknodora avae Haitlinger, 1994b: 81 Type locality: Costa Rica Distribution: Costa Rica Host: Stolas lebesi (Coleoptera: Chrysomelidae) Phriknodora pustulosa Summers et Schuster Phriknodora pustulosa Summers & Schuster, 1982b: 43 Type locality: Oaxaca Distribution: Mexico, Panama Hosts: Brazilianus mexicanus, B. batus (Coleopera: Cerambycidae) Phriknodora scabra Summers et Schuster Phriknodora scabra Summers & Schuster, 1982b: 46 Type locality: South Africa Distribution: South Africa Host: Trachyderes succinctus (Coleoptera: Cerambycidae) Genus Procericola Cooreman Procericola Cooreman, 1950: 44 Type species: Procericola ichthyoides Cooreman, 1950 Procericola bourgognei (Oudemans) Photia bourgognei Oudemans, 1923b: 203 Acarorum Catalogus IX 203 Caraboecius coriacei Cooreman, 1950: 19 Caraboecius bourgognei Cooreman, 1954: 87 Procericola bourgognei (Oudemans): Samšinák, 1965: 138; 1971; Beron, 1971: 205; Lundquist, 1985: 381; Khaustov & Eidelberg, 2001: 39; Trach, 2006: 175 Type locality: France (Bourgogne) Distribution: Albania, Belgium, Bulgaria, Caucasus, former Jugoslavia, France, Greece, Hungary, Italy, Moldova, Romania, Russia, Slovakia, Sweden, Turkey, Ukraine Hosts: Carabus coriaceus, Carabus (Procrustes) ch. chevrolati, C. (P.) clypeatus clypeatus, C. (Megodontus) exaratus (Coleoptera: Carabidae) Procericola ichthyoides Cooreman Procericola ichthyoides Cooreman, 1950: 44 Procericola ichthyoides ichthyoides Cooreman: Samšinák, 1964: 38 Type locality: “Carniolia, Yougoslavie et Tolmezzo, Torrente Vinadia, Carnia” Distribution: Italy, Slovenia Host: Carabus (Procerus) gigas (Coleoptera: Carabidae) Procericola ichthyoides scabricola Samšinák Procericola ichthyoides scabricola Samšinák, 1964: 38 Type locality: Caucasus, Mekish Mt. Distribution: Bulgaria, Russia, Turkey Hosts: Carabus (Procerus) scabrosus transversalis (Coleoptera: Carabidae) Procericola ichthyoides transversicola Samšinák Procericola ichthyoides transversicola Samšinák, 1964: 40 Type locality: “Cilicien” Distribution: Turkey Hosts: Carabus (Procerus) scabrosus transversalis (Coleoptera: Carabidae) Genus Pseudamansia Cooreman Pseudamansia Cooreman, 1950: 5 Type species: Dermaleichus chrysomelinus Koch, 1841 Pseudamansia chrysomelinus (C.L. Koch) Dermaleichus chrysomelinus C.L. Koch, 1841: 4 Amansia chrysomelinus: Oudemans, 1923: Pseudamansia chrysomelinus (C.L. Koch): Cooreman, 1950: 5; Jolivet, 1952, 1954; Theodorides, 1955; Santiago-Blay & Fain, 1994: 412 Pseudamansia chrysomelina (C.L. Koch): Bernini, Type locality: various places in France (Neotype) 204 Acarorum Catalogus IX Distribution: Belgium, France, Algeria, Morocco, Mayorca Is., Italy Hosts: Timarcha atlantica, T. balearica, T. goettingensis, T. interstitialis, T. nicaeensis, T. pauperata, T. punctella, T. rugosa, T. tenebricosa (Coleoptera: Chrysomelidae) Pseudamansia decorata Trach et Khaustov Pseudamansia decorata Trach & Khaustov, 2007: 269 Type locality: Odessa and Vinnica regions Distribution: Ukraine Host: Timarcha goettingensis (Coleoptera: Chrysomelidae) Genus Pseudocanestrinia Khaustov et Eidelberg Pseudocanestrinia Khaustov & Eidelberg, 2001: 31 Type species: Pseudocanestrinia carabi Khaustov et Eidelberg, 2001 Pseudocanestrinia berndi (Haitlinger) Canestrinia berndi Haitlinger, 1995: 260 Pseudocanestrinia berndi (Haitlinger): Khaustov & Eidelberg, 2001: 34 Type locality: Jablanica (Herzegovina) Distribution: Bosna and Herzegovina, Crimea Hosts: Carabus latenulatus plassensis, C. gyllenhali (Coleoptera: Carabidae) Pseudocanestrinia carabi Khaustov et Eidelberg Pseudocanestrinia carabi Khaustov & Eidelberg, 2001: 33; Trach, 2006: 175 Type locality: Crimean mountains ‘Ay-Petri’ Distribution: Crimea Host: Carabus (C.) bessarabicus (Coleoptera: Carabidae) Pseudocanestrinia mahunkai (Samšinák) Canestrinia mahunkai Samšinák, 1971: 160; Haitlinger, 1988 Pseudocanestrinia mahunkai (Samšinák): Khaustov & Eidelberg, 2001: 32; Haitlinger, 2012: 186 Type locality: Balatonaracs (Hungary) Distribution: Hungary, Poland Host: Carabus coriaceus (Coleoptera: Carabidae) Pseudocanestrinia occidentalis (Samšinák) Canestrinia occidentalis Samšinák, 1971: 155 Pseudocanestrinia occidentalis (Samšinák): Khaustov & Eidelberg, 2001: 32 Type locality: La Lourde Distribution: France Host: Carabus (Chrysocarabus) auronitens (Coleoptera: Carabidae) Acarorum Catalogus IX 205 Pseudocanestrinia orientalis (Samšinák) Canestrinia orientalis Samšinák, 1971: 157 Pseudocanestrinia orientalis (Samšinák): Khaustov & Eidelberg, 2001: 32 Type locality: “Carniolia” Distribution: Slovenia Hosts: Carabus (Megodontus) caelatus, C. (M.) croaticus (Coleoptera: Carabidae) Genus Pseudoparaphagella Haitlinger Pseudoparaphagella Haitlinger, 2010: 58 Type species: Pseudoparaphagella gerardi Haitlinger, 2010 Pseudoparaphagella gerardi Haitlinger Pseudoparaphagella gerardi Haitlinger, 2010: 58 Type locality: Guinea Distribution: Guinea Host: Caelorrhina thoreyi (Coleoptera: Scarabaeidae, Cetoniinae) Genus Rugoniphela Haitlinger Rugoniphela Haitlinger, 1992b: 533 Type species: Rugoniphela marloni Haitlinger, 1992 Rugoniphela marloni Haitlinger Rugoniphela marloni Haitlinger, 1992b: 534 Type locality: holotype from Borneo, paratypes from Java and Sumatra Distribution: Indonesia (Borneo, Java, Sumatra) Host: Eurytrachelus bucephalus (Coleoptera: Lucanidae) Genus Sabothiana Haitlinger Sabothiana Haitlinger, 1993e: 269 Type species: Sabothiana riccardoi Haitlinger, 1993 Sabothiana boi Haitlinger Sabothiana boi Haitlinger, 1993e: 272 Type locality: Bahia Distribution: Brazil Host: Charidotis sp.(Coleoptera: Chrysomelidae, Cassidinae) Sabothiana reidari Haitlinger Sabothiana reidari Haitlinger, 1993e: 273 Type locality: Bocaina Distribution: Brazil Host: Charidotella kesseli (Coleoptera: Chrysomelidae, Cassidinae) 206 Acarorum Catalogus IX Sabothiana riccardoi Haitlinger Sabothiana riccardoi Haitlinger, 1993e: 271 Type locality: Brazil Distribution: Brazil Host: Charidotis furunculus (Coleoptera: Chrysomelidae, Cassidinae) Genus Sajanophela Haitlinger Sajanophela Haitlinger, 1990b: 545 Type species: Sajanophela alfredae Haitlinger, 1990 Sajanophela alfredae Haitlinger Sajanophela alfredae Haitlinger, 1990b: 545 Type locality: Papua New Guinea Distribution: Papua New Guinea Host: Protomocelus australis (Coleoptera: Passalidae) Genus Sandrophela Haitlinger Sandrophela Haitlinger, 1990: 315 Type species: Sandrophela kokodaica Haitlinger, 1990 Sandrophela amarae Haitlinger Sandrophela amarae Haitlinger, 1994a: 197 Type locality: East Java, Sumatra Distribution: Indonesia (Java, Sumatra) Host: Eurytrachelus eurycephalus, E. gypeatus (Coleoptera: Lucanidae) Sandrophela dittae Haitlinger Sandrophela dittae Haitlinger, 1992: 282 Type locality: Borneo Distribution: Indonesia Host: Colydium sp. (Coleoptera: Colydiidae) Sandrophela hugoni Haitlinger Sandrophela hugoni Haitlinger, 1994a: 194 Type locality: Sumatra Distribution: Indonesia (Sumatra) Host: Odontolabis wollastoni (Coleoptera: Lucanidae) Sandrophela kokodaica Haitlinger Sandrophela kokodaica Haitlinger, 1990e: 316 Type locality: Kokoda Acarorum Catalogus IX 207 Distribution: Papua New Guinea Host: Passandra trigemina (Coleoptera: Passandridae) Sandrophela rodioni Haitlinger Sandrophela rodioni Haitlinger, 1993b: 342 Type locality: North Vietnam, Monte Mouson Distribution: Vietnam Host: Eurytrachelus platymelus (Coleoptera: Lucanidae) Sandrophela sophiae Haitlinger Sandrophela sophiae Haitlinger, 1994a: 196 Type locality: India Distribution: India Host: Hemisodorcus chevrilati (Coleoptera: Lucanidae) Sandrophela spectanda (Berlese) Canestrinia spectanda Berlese, 1910: 269 Sandrophela spectanda (Berlese): Haitlinger, 1994a: 199 Type locality: Java Distribution: Indonesia (Java, Sumatra) Host: Dorcus bucephalus (Coleoptera: Lucanidae) Genus Saniothiana Haitlinger Saniothiana Haitlinger, 1990e: 317 Type species: Saniothiana pycnosa Haitlinger, 1990 Saniothiana barumbaica Haitlinger Saniothiana barumbaica Haitlinger, 1990e: 320 Type locality: Barumba Distribution: Cameroon Host: Odontopezus cupreus lucens (Coleoptera: Tenebrionidae) Saniothiana pycnosa Haitlinger Saniothiana pycnosa Haitlinger, 1990e: 318 Type locality: Africa Distribution: Africa Host: Pycnocerus sp. (Coleoptera: Tenebrionidae) Genus Scarabiphotia Haitlinger Scarabiphotia Haitlinger, 1994a: 205 Type species: Scarabiphotia vignoni Haitlinger, 1994 208 Acarorum Catalogus IX Scarabiphotia vignoni Haitlinger Scarabiphotia vignoni Haitlinger, 1994a: 205 Type locality: unknown Distribution: unknown Host: Scarabaeus variolosus (Coleoptera: Scarabaeidae) Genus Shillongia Haitlinger Shillongia Haitlinger, 1990b: 546 Type species: Shillongia marzenae Haitlinger, 1990 Shillongia marzenae Haitlinger Shillongia marzenae Haitlinger, 1990b: 546 Type locality: Assam, Shillong Distribution: India (Assam) Host: Leptaulus indicus (Coleoptera: Passalidae) Genus Sorbinophela Haitlinger Sorbinophela Haitlinger, 1993a: 7 Type species: Sorbinophela sandyi Haitlinger, 1993 Sorbinophela sandyi Haitlinger Sorbinophela sandyi Haitlinger, 1993a: 7 Type locality: nr. Lake Ngami Distribution: Botswana Host: Poecilophila maculatissima (Coleoptera : Scarabaeidae, Cetoniinae) Genus Tamarangia Haitlinger Tamarangia Haitlinger, 1991d: 284 Type species: Tamarangia nimfae Haitlinger, 1992 Tamarangia ethiopica Khaustov Tamarangia ethiopica Khaustov, 2006: 79 Type locality: Ethiopia Distribution: Ethiopia Host: Pachnoda marginata (Coleoptera: Scarabaeidae, Cetoniinae) Tamarangia fabiolae Haitlinger Tamarangia fabiolae Haitlinger, 1991d: 286 Type locality: Freetown Distribution: Sierra Leone Host: Pachnoda marginata (Coleoptera: Scarabaeidae, Cetoniinae) Acarorum Catalogus IX 209 Tamarangia flawiani Haitlinger Tamarangia flawiani Haitlinger, 1991d: 287 Type locality: Freetown Distribution: Sierra Leone Host: Pachnoda marginata (Coleoptera: Scarabaeidae, Cetoniinae) Tamarangia nimfae Haitlinger Tamarangia nimfae Haitlinger, 1991d: 284; Haitlinger & Chmielewski, 2004: 199 Type locality: Ethiopia Distribution: Ethiopia, South Africa Host: Pachnoda thoracica (Coleoptera: Scarabaeidae, Cetoniinae) Genus Tamarophela Haitlinger Tamarophela Haitlinger, 1990b: 537 Type species: Tamarophela batjanica Haitlinger, 1990 Tamarophela batjanica Haitlinger Tamarophela batjanica Haitlinger, 1990b: 537 Type locality: Moluccas, Batjan Is. Distribution: Indonesia (Moluccas, Batjan Is.) Host: undetermined Passalidae (Coleoptera) Genus Taseopus Summers et Schuster Taseopus Summers & Schuster, 1982a: 27 Type species: Taseopus platythrix Summers et Schuster, 1982 Taseopus platythrix Summers et Schuster Taseopus platythrix Summers & Schuster, 1982a: 27 Type locality: Queensland Distribution: Australia Host: Passalidae gen. sp. (Coleoptera) Genus Teklathiana Haitlinger Teklathiana Haitlinger, 1999: 36 Type species: Teklathiana onnae Haitlinger, 1999 Teklathiana conula (Summers et Schuster) Grandiella conula Summers & Schuster, 1979: 341 Grandiellina conula: Santiago-Blay & Fain, 1994: 210 Acarorum Catalogus IX Ennodithiana conula (Summers & Schuster): Haitlinger 1993: 72 Teklathiana conula (Summers et Schuster): Haitlinger, 1999: 40 Type locality: Panama (El Cope) Distribution: Panama Host: Omaspides bistriata (Coleoptera: Chrysomelidae, Cassidinae) Teklathiana onnae Haitlinger Teklathiana onnae Haitlinger, 1999: 37; 2001: 103 Type locality: holotype from Colombia, paratype from Bolivia Distribution: Bolivia, Colombia Host: Acromis sparsa (Coleoptera: Chrysomelidae, Cassidinae) Teklathiana rugosita (Summers et Schuster) Grandiella rugosita Summers & Schuster, 1979: 343 Teklathiana rugosita (Summers et Schuster): Haitlinger, 1999: 40 Type locality: Panama (Canal Zone and Panama City) Distribution: Panama Host: Acromis sparsa (Coleoptera: Chrysomelidae, Cassidinae) Genus Teophyssa Haitlinger Teophyssa Haitlinger, 1990b: 544 Type species: Teophyssa honoratae Haitlinger, 1990 Teophyssa beatae Haitlinger Teophyssa beatae Haitlinger, 1990b: 544 Type locality: Sumatra, Bekantiang Distribution: Indonesia (Sumatra) Host: undetermined Passalidae (Coleoptera) Teophyssa honoratae Haitlinger Teophyssa honoratae Haitlinger, 1990b: 544 Type locality: North Sumatra Distribution: Indonesia (Sumatra) Host: Gonatus noviculator (Coleoptera: Passalidae) Teophyssa teresae Haitlinger Teophyssa teresae Haitlinger, 1990b: 545 Type locality: Sumatra, Soekaranda Distribution: Indonesia (Sumatra) Host: undetermined Passalidae (Coleoptera) Acarorum Catalogus IX 211 Genus Thopia Summers et Schuster Thopia Summers & Schuster, 1982b: 35 Type species: Thopia tenax Summers et Schuster, 1982 Thopia amedae Haitlinger Thopia amedae Haitlinger, 1993e: 268 Type locality: Cayenne Distribution: French Guiana Host: Eugenysa venosa (Coleoptera: Chrysomelidae, Cassidinae) Thopia boguwolae Haitlinger Thopia boguwolae Haitlinger, 1993e: 266 Type locality: Brazil Distribution: Brazil Host: Polychalca punctatissima (Coleoptera: Chrysomelidae, Cassidinae) Thopia bulbocaudata Summers et Schuster Thopia bulbocaudata Summers & Schuster, 1982b: 37 Type locality: Panama Distribution: Panama Host: Neoptychodes trilineatus (Coleoptera: Cerambycidae, Lamiinae) Thopia corinnae Haitlinger Thopia corinnae Haitlinger, 1993e: 268 Type locality: Bahia Distribution: Brazil Host: Polychalca dentipennis (Coleoptera: Chrysomelidae, Cassidinae) Thopia pannucea Summers et Schuster Thopia pannucea Summers & Schuster, 1982b: 41 Type locality: Panama Distribution: Panama Host: Lagocheirus foveolatus (Coleoptera: Cerambycidae, Lamiinae) Thopia tenax Summers et Schuster Thopia tenax Summers & Schuster, 1982b: 37 Type locality: Canal Zone Distribution: Panama Host: Taeniodes scalaris (Coleoptera: Cerambycidae) Genus Uriophela Haitlinger Uriophela Haitlinger, 1992b: 539 212 Acarorum Catalogus IX Type species: Uriophela arieli Haitlinger, 1992 Uriophela arieli Haitlinger Uriophela arieli Haitlinger, 1992b: 540 Type locality: Amur District (Russia) Distribution: Russia (Far East) Host: Hemisodorcus [Macrodora] rubrofemoratus (Coleoptera: Lucanidae) Genus Vandeinella Haitlinger Vandeinella Haitlinger, 1989a: 54 Type species: Vandeinella adriasae Haitlinger, 1989 Vandeinella adriasae Haitlinger Vandeinella adriasae Haitlinger, 1989a: 54 Type locality: Sumatra Distribution: Indonesia (Sumatra) Host: Trichogomphus milon (Coleoptera: Scarabaeidae, Dynastinae) Vandeinella opressa Haitlinger Vandeinella opressa Haitlinger, 1989a: 55 Type locality: Borneo Distribution: ? Indonesia (Borneo) Host: Trichogomphus lunicollis (Coleoptera: Scarabaeidae, Dynastinae) Subfam. Vereoxiinae Haitlinger Vereoxiinae Haitlinger, 1995a: 80 Type genus: Vereoxia Haitlinger, 1995 Genus Vereoxia Haitlinger Vereoxia Haitlinger, 1995a: 80 Type species: Vereoxia bogeri Haitlinger, 1995 Vereoxia bogeri Haitlinger Vereoxia bogeri Haitlinger, 1995a: 80 Type locality: unknown Distribution: unknown Host: Lucanidae gen. sp. (Coleoptera) Acarorum Catalogus IX 213 Index of Canestriniidae (Names in bold are accepted as valid) abapoica, Gasuthiana 174 abufarica, Gasuthiana 174 Acrotocarus 148 adanensis, Photia procera 201 adelinae, Olafithiana 191 adolfinae, Photia 198 adolphinae, Photia 198 adriasae, Vandeleinella 212 africanus, Diplognathophilus 171 Afrocanestrinia 148 aidae, Eunorithiana 174 alani, Grandiellopsis 179 alatus, Phaleratus 198 albertae, Bibulothiana 158 albini, Coleoglyphus 167 Albinorattia 149 tacea, Amansia almae, Kahoorangia 183 alutacea, Amansia 148 alutacea, Grandiella 148 alutaceus, Acrotocarus 148 alvari, Barbiangia 157 Amansia Amansiella 149 amarae, Sandrophela 206 Ambilohylla 149 Amboinophela 149 amedae, Thopia 211 amplexans, Canestriniella 164 Anapistes 150 anomala, Melisia 185 anoni, Averithiana 157 Apalotacarus 150 archidonaica, Danaithiana 170 arcuata, Arrunsithiana 153 arcuata, Grandiella 153 arcuata, Grandiellina 153 ariadnae, Melisia 185 ariani, Grandiellopsis 179 arieli, Uriophela 212 aristatus, Apalotacarus 150 Arraphosoma 152 Arrunsithiana 153 artemoni, Arrunsithiana 153 aruica, Lidiophela 183 arybbasi, Athogavia 155 asahinai, Ibizella asilahica, Ibizella 182 assamica, Athogavia 155 Athogavia 155 Aurilossongia 156 Averithiana 157 avae, Phiknodorae 202 ayraica, Danaithiana 170 baali, Coleopterophagus 168 bahiana, Hatohylla 181 balearica, Ibizella 182 baloghi, Melisia 185 barbarae, Jovitella 183 barryi, Noemiphela 190 barumbaica, Saniothiana 207 batjanica, Tamarophela 209 batocerae, Canestriniella 164 batocerae, Grandiella 164 batocerae, Grandiellina 164 battosi, Arrunsithiana 153 beatae, Teophyssa 210 Beeriphotia 158 belerofoni, Hyllosihemia 181 beloniana, Megacanestrinia 184 belzebubi, Coleopterophagus 168 berlesei, Coleopterophagus 214 Acarorum Catalogus IX bernae, Amboinophela 149 berndi, Canestrinia 204 berndi, Pseudocanestrinia 204 bertrami, Bircericola 159 bettinae, Globosophotia 175 bibiannae, Hatohylla 181 Bibulothiana 158 bicornis, Paramansia 193 bilkorum, Photia 199 Bircericola 159 bissaina, Mossongia 189 blaptis, Alloptes 195 blaptis, Canestrinia 195 blaptis, Coleopterophagus 195 blaptis, Percanestrinia 195 blattophaga, Canestrinia 168 blattophagus, Coleopterophagus 168 Boetophela 159 bogeri, Vereoxia 212 bognae, Lidiophela 183 boguwolae, Thopia 211 boi, Sabothiana 211 bolanica, Aurilossongia 156 bolebori, Metriothiana 189 Boleohylla 161 borneoica, Athogavia 155 bourgognei, Caraboecius 203 bourgognei, Photia 202 bourgognei, Procericola 202 brevispinata, Bibulothiana 158 brevispinata, Grandiella 158 briannae, Peruthiana 197 bulbocaudata, Thopia 211 calosomae, Paraphotia 195 calostegis, Donnelafontia 172 Camirohylla 161 Canestrinia 161 Canestrinia (Dicanestrinia) 171 Canestrinia (Percanestrinia) 161 Canestriniella 164 Canestriniidae 147 Canestriniinae 147 canistra, Grandiellopsis 179 carabi, Pseudocanestrinia 204 carabicola, Canestrinia 161 carabicola, Coleopterophagus 161 carabicola, Percanestrinia 161 Caraboecius 165 caroae, Gioharattia 175 cassandrae, Boetophela 159 caucasica, Percanestriniella 197 celinae, Lidiophela 184 cerambicis, Canestrinia (Dicanestrinia) 171 cerambycis, Dermaleichus 171 cerambycis, Dicanestrinia 171 cetoniae, Coleopterophagus 169 Cetonicola 165 chanchamayoica, Danaithiana 170 Chelynochroa 166 chrysocarabi, Photia 199 chrysomelinus, Amansia 203 chrysomelinus, Dermaleichus 203 chrysomelinus, Pseudomansia 203 cidaris, Apalotacarus 150 Cnecoderma 166 coarica, Gasuthiana 174 Coleoglyphus 167 Coleopterophagus 167 collaris, Hypopteryx 181 conula, Ennodithiana 210 conula, Grandiella 209 conula, Grandiellina 209 conula, Teklathiana 209 cooremani, Grandiella 175 cooremani, Grandiellina 175 Cooremania 169 coriacei, Caraboecius 165 corinnae, Thopia 211 crataepus, Acrotocarus 148 cyclosoma, Grandiellopsis 180 cyriaki, Bibulothiana 158 damiani, Canestrinia 162 Danaithiana 170 Acarorum Catalogus IX 215 decemcaudata, Grandiella 176 decemcaudata, Grandiellina 176 decorata, Pseudamansia 204 delmari, Dorynothiana 172 denmarkica, Paracanestrinia 193 Dicanestrinia 171 dolichopoda, Cnecoderma 167 dictyophora, Chelinochroa 166 dioni, Aurilossongia 156 dionizyi, Coleopterophagus 168 Diplognathophilus 171 Diplopodocoptes 171 dittae, Sandrophela 206 dolichopoda, Cnecoderma 167 donaldi, Coleopterophagus 168 Donnelafontia 172 dorcicola, Canestrinia 162 Dorynothiana 172 drvotocka, Neophotia 189 echinatus, Apalotacarus 150 echinoderma, Grandiella 192 echinoderma, Grandiellina 192 echinoderma, Ovarrothiana 192 ecuadorica, Danaithiana 170 ecuadorensis, Grandiella 176 ecuadorensis, Grandiellina 176 egypti, Percanestrinia 156 elae, Canestriniella 165 elongata, Barbiangia 157 Ennodithiana 172 Ennorithiana 173 ephraimi, Boetophela 159 erazmi, Ovarrothiana 192 erhardi, Boetophela 159 erinnae, Arrunsithiana 154 escaudata, Arrunsithiana 154 esacaudata, Grandiella 154 esacaudata, Grandiellina 154 ethiopica, Barbiangia 157 ethiopicus, Diplopodophilus 157 ethiopica, Tamarangia 208 Eucanestrinia 198 eudicellae, Paraphagella 194 Eunorithiana 173 everittae, Bibulothiana 158 fabiolae, Tamarangia 208 favosa, Ambilohylla 149 fentoni, Phaleratus 198 feziana, Camirohylla 161 fileasi, Aurilossongia 156 flawiani, Tamarangia 209 foliacea, Grandiella 191 foliacea, Grandiellina 191 foliacea, Olafithiana 191 fuscipes, Coleoglyphus 167 fulgens, Apalotacarus 150 fusticulus, Apalotacarus 151 fusulus, Apalotacarus 151 gabrielae, Melisia 185 gamana, Athogavia 155 gambosa, Grandiella 176 gambosa, Grandiellina 176 garrica, Boleohylla 161 Gasuthiana 174 guarana, Aurilossongia 156 gentilis, Percanestrinia (Lombardiniella) georgei, Passalophagus 195 gerardi, Pseudoparaphagella 205 giardi, Canestrinia 193 giardi, Paramansia 193 ginae, Boetophela 160 Gioharattia 175 glaber, Apalotacarus 151 Globosophotia 175 gracilis, Apalotacarus 151 graeca, Photia 199 Grandiella 175 Grandiellina 175 Grandiellopsis 179 Haitlingeria 180 Hargeinella 180 Hatohylla 181 hejniana, Photia 199 helenae, Melisia 186 216 Acarorum Catalogus IX helgae, Irmongia 182 heliodori, Eunorithiana 173 hermengildae, Photia 199 hieronimae, Melisia 186 hispanica, Canestrinia 162 hispanica, Melisia hispidus, Cetonicola 165 hoaria, Aurilossongia 156 honoratae, Teophyssa 210 howdeni, Photia 199 huberti, Dicanestrinia 171 hugoni, Sandrophela 206 Hyllosihemia 181 Hypopteryx 181 Ibizella 182 ichtyoides, Procericola 203 igae, Lidiophela 184 igori, Albinorattia 149 illegalis, Canestrinia 162 ilosi, Arrunsithiana 154 indirae, Paraphagella 194 ingeborgae, Melisia 186 ingridae, Paraphagella 194 irenae, Melisia 186 Irmongia 182 isadorae, Cetonicola 166 iwettae, Olafithiana 191 izabelae, Noemiphela 190 izabelae, Percanestrinia 196 izae, Jullongia 183 jadwigae, Melisia 186 javensis, Canestrinia 200 javensis, Photia 200 joliveti, Grandiella 191 joliveti, Grandiellina 191 joliveti, Olafithiana 191 Jovitella 183 Jullongia 183 jurandi, Dorynothiana 172 jureceki, Neophotia 190 kacperi, Canestrinia 162 Kahoorangia 183 kamelskyi, Canestrinia 162 kietaensis, Cooremania 170 kimseyi, Grandiella 176 kimseyi, Grandiellina 176 kiogana, Boleohylla 161 klapaleki, Melisia 186 knobi, Dicanestrinia 171 kokodaica, Sandrophela 206 lacuna, Paramelisia 194 latipes, Arrunsithiana 154 latipes, Grandiellina 154 latissima, Neophotia 190 leopoldi, Ovarrothiana 192 leptinotarsa, Melisia 186 levis, Canestrinia 197 levis, Percanestriniella 197 Lidiophela 183 lombardinii, Melisia 187 longilobata, Haitlingeria 180 longini, Grandiella 177 longini, Grandiellina 177 lopatini, Photia 200 ludomiri, Ovarrothiana 192 luroris, Apalotacarus 151 lusitanica, Photia 200 macgillavryi, Canestrinia 163 maharae, Gioharattia 175 mahunkai, Canestrinia 204 mahunkai, Melisia 187 mahunkai, Pseudocanestrinia 204 manicata, Canestrinia 163 maniusi, Ennodithiana 173 marani, Photia procera 201 mariettae, Melisia 187 marinae, Grandiella 177 marinae, Grandiellina 177 maritae, Cetonicola 166 marloni, Rugoniphela 205 maroccana, Percanestrinia 196 maroni, Coleopterophagus 168 marzenae, Shillongia 208 medoni, Ennodithiana 173 Acarorum Catalogus IX 217 Megacanestrinia 184 megesi, Emmodithiana 173 megninii, Coleopterophagus 169 megnini, Coleopterophagus 169 megninii, Dermogliphus 168 megnini, Dermoglyphus 168 melaniae, Melisia 187 melchiori, Photia 200 melillae, Gasuthiana 174 Melisia 185 melisii, Melisia 187 Melisiinae 185 menthastri, Paramansia 193 Mesophotia 188 messalinae, Ennodithiana 173 Metriothiana 188 microdisca, Canestrinia 163 minax, Araphosoma 152 minor, Grandiella 177 minor, Grandiellina 177 minor, Paraphagella 194 mirabilis, Acrotocarus 148 morrata, Hatohylla 181 Mossongia 189 mucronata, Megacanestrinia 185 multifida, Grandiella 191 multifida, Olafithiana 191 neglectus, Coleopterophagus 169 Neophotia 189 nepalensis, Canestrinia 163 nikodemi, Ovarrothiana 192 nimfae, Tamarangia 209 Noemiphela 190 norae, Aurilossongia 157 norodomi, Percanestrinia 196 nosiana, Athogavia 155 occidentalis, Canestrinia 204 occidentalis, Pseudocanestrinia 204 occidii, Melisia 187 Oceogavia 190 octocaudata, Grandiella 177 octocaudata, Grandiellina 177 odae, Paraphagella 194 odiosi, Grandiellopsis 180 Olafithiana 190 olareica, Gasuthiana 174 Olgattia 191 omoplata, Grandiella tetracaudata 179 omoplata, Grandiellina tetracaudata 179 onnae, Teklathiana 210 opressa, Vandeleinella 212 orestesi, Cetonicola 166 orientalis, Canestrinia 205 orientalis, Pseudocanestrinia 205 orsenae, Grandiellopsis 180 oswaldi, Bubulothiana 159 ovalis, Cnecoderma 167 Ovarrothiana 192 ozannae, Ovarrothiana 193 paavoi, Canestrinia 163 pacifica, Photia 200 pannucea, Thopia 211 Paracanestrinia 193 Paramansia 193 Paramelisia 193 paraminax, Araphosoma 153 Paraphagella 194 Paraphotia 195 Passalophagus 195 pauliana, Lidiophela 184 paxillus, Apalotacarus 151 pecki, Coleopterophagus 184 pecki, Lidiophela 184 pekingensis, Canestrinia 163 pelagiae, Lidiophela 184 penelopae, Ennodithiana 173 penicilata p., Mesophotia 187 penicillata similis, Mesophotia 187 pentagona, Grandiella 177 pentagona, Grandiellina 177 pentodontis, Canestrinia 164 Percanestrinia 195 Percanestriniella 197 Peruthiana 197 218 Acarorum Catalogus IX petilus, Apalotacarus 151 Phaleratus 197 Phelliculophela 198 phoebinae, Canestriniella 165 Photia 198 Phriknodora 202 physsana, Canestriniella 165 pictura, Canestrinia 164 platysma, Ennodithiana 173 platysma, Grandiella 173 platythrix, Taseopus 209 polymorpha, Photia 200 poppana, Hargeinella 181 princeps, Paraphagella 195 procera, Canestrinia (Eucanestrinia) 201 procera, Photia 201 Procericola 202 procrusti, Eucanestrinia 201 procrusti, Photia 201 procrustidis, Eucanestrinia 201 procustidis, Canestrinia 201 procustidis, Photia 201 protensus, Apalotacarus 152 Pseudamansia 203 Pseudocanestrinia 204 Pseudoparaphagella 205 pulcher, Coleopterophagus 169 pustulosa, Phriknodora 202 pycnosa, Saniothiana 207 quadrata, Arrunsithiana 154 quadrata, Grandiella 154 quadrata, Grandiellina 154 quadrisetosus, Coleopterophagus 169 rackae, Canestrinia blaptis 195 ramoni, Canestrinia 164 reidari, Sabothiana 205 remigans, Canestrinia 164 riccardoi, Sabothiana 206 rigescens, Apalotacarus 151 roaldi, Pheliculophela 198 robertsoni, Cetonicola 160 robertsoni, Boetophela 160 rodioni, Sandrophela 207 rogeri, Grandiella 177 rogeri, Grandiellina 177 rosascostai, Coleoglyphus 178 rosascostai, Grandiella 178 rosascostai, Hatohylla 178 rosascostai, Donnelafontia 178 rudolfi, Coleopterophagus 169 rufinae, Afrocanestrinia 148 Rugoniphela 305 rugosita, Grandiella 210 rugosita, Teklathiana 210 rutae, Melisia 187 Sabothiana 205 saetolata, Percanestrinia 202 saetolata, Photia 202 sajanae, Apalotacarus 152 Sajanophela 206 salomeae, Melisia 188 samsinaki, Canestrinia 182 samsinaki, Ibizella 182 samsinaki, Melisia 188 samsoni, Afrocanestrinia 148 Sandrophela 206 sandyi, Sorbinophela 208 Saniothiana 206 sardica, Canestrinia (Percanestrinia) 161 scabra, Phriknodora 202 scabricola, Procericola ichthioides 203 scagana, Athogavia 155 Scarabiphotia 208 scaurus, Apalotacarus 152 scissus, Apalotacarus 152 sellnicki, Canestrinia 196 sellnicki, Percanestrinia 196 sentosa, Melisia 188 Shilongia 208 sibirica, Photia 202 sicula, Grandiella 178 sicula, Grandiellina 178 silvanae, Boetophela 160 simplex, Coleoglyphus 167 Acarorum Catalogus IX 219 soniae, Grandiella 178 soniae, Grandiellina 178 sophiae, Sandrophela 207 Sorbinophela 208 spectanda, Canestrinia 207 spectanda, Sandrophela 207 spicantis, Arrunsithiana 154 spicantis, Grandiella 154 spicantis, Grandiellina 154 squarrosa, Arrunsithiana 154 squarrosa, Grandiella 154 ssavina, Mossongia 189 straeleni, Afrocanestrinia 148 suatoha, Donelaphontia 172 suatoha, Farahanella 172 superba, Melisia 187 sybiellae, Boetophela 160 symeoni, Boetophela 160 tacjannae, Grandiella 178 tacjannae, Grandiellina 178 Tamarangia 208 Tamarophela 209 tanzanica, Athogavia 156 Taseopus 209 Teklathiana 209 tenax, Thopia 211 Teophyssa 210 teresae, Teophyssa 210 tertia, Grandiella 178 tertia, Grandiellina 179 tertia, Mesophotia penicillata 189 tetracaudata, Grandiella 179 tetracaudata, Grandiellina 179 Thopia 211 togata, Canestriniella 165 transkeiensis, Diplopodocoptes 171 transversicola, Procericola ichtihyoides 203 trilobata, Amansiella 149 trullus, Apalotacarus 152 turcica, Photia procera 201 ubaldi, Oceogavia 190 unguiculata, Anaspistes 150 Uriophela 211 useguaica, Olgattia 192 Vandeinella 212 vatia, Bibulothiana 159 vatia, Grandiella 159 vatus, Cetonicola 166 Vereoxia 212 Vereoxiinae 212 vietnamensis, Contramelisia vignoni, Scarabiphotia 208 violae, Grandiellopsis 180 viviannae, Percanestrinia 196 vlachiana, Canestrinia 197 vlachiana, Percanestriniella 197 volkeri, Beeriphotia 158 wauensis, Cooremania 170 werneri, Boetophela 160 Distribution of Canestriniidae ЕUROPE Albania (Samšinák, 1965) – Photia procustidis, Procericola bourgognei Austria (Samšinák, 1971; Haitlinger, 1990d, 1995b) - Coleopterophagus albini, Photia bardoica, Ph. chrysocarabi, Ph. hejniana Belgium (Cooreman, 1950; Fain, Noti & Dufrêne, 1995) - Photia chrysocarabi, Ph. saetolata, Procericola bourgognei, Pseudamansia chrysomelinus Bosna and Herzegovina (Samšinák, 1971; Haitlinger, 1995b) – Dicanestrinia knobi, Pseudocanestrinia berndi, P. carabi 220 Acarorum Catalogus IX Bulgaria (Samšinák, 1970; Beron, 1971, 1975) – Ibizella [Canestrinia] samsinaki, Percanestrinia blaptis, Procericola bourgognei, P. ichthyoides, Photia procera Byelorussia (Khaustov & Eidelberg, 2001) – Dicanestrinia huberti, Photia adolphinae Crimea (Khaustov & Eidelberg, 2001) – Amansiella trilobata, Canestrinia damiani, C. dorcicola, Paramansia bicornis, Paraphotia calosomae, Photia polymorpha, Pseudocanestrinia berndti, P. carabi Croatia (Samšinák, 1971; Haitlinger, 1990d ) - Coleopterophagus megnini, Photia hejniana Czech Rep. (Samšinák, 1965, 1971; Haitlinger, 1990d) - Coleopterophagus albini, C. megnini, Dicanestrinia knobi, Percanestrinia sellnicki, Photia chrysocarabi, Ph. hejniana, Ph. saetolata Denmark (Haitlinger, 2017) - Paracanestrinia denmarkica Estonia (Haitlinger, 2015) – Coleopterophagus albini France (contin.) (Oudemans, 1923; Cooreman, 1950; Jolivet, 1950, 1954; Samšinák, 1970, 1971) – N. jureceki, Paramansia menthastri, Percanestrinia blaptis, Photia chrysocarabi, Ph. saetolata, Procericola bourgognei, Pseudocanestrinia occidentalis, Pseudamansia chrysomelinus Corsica (Haitlinger, 1992a) – Canestrinia carabicola, Percanestrinia maroccana Georgia (Samšinák, 1971; Haitlinger, 1992a) – Percanestrinia blaptis, Photia bilkorum Germany (Poppe, 1907; Rack, 1968; Samšinák, 1965, 1970; Haitlinger, 1992a) – Coleopterophagus albini, Percanestrinia blaptis, Procericola bourgognei, Photia chrysocarabi, Ph. saetolata Great Britain (Turk, 1953; Evans, Sheals & Macfarlane, 1961; Samšinák, 1965, 1971) – Canestrinia dorcicola, Coleopterophagus megnini, Percanestrinia blaptis, Photia chrysocarabi, Ph. saetolata Greece (Cooreman, 1958; Samšinák, 1965) - Photia graeca, Ph. procustidis,, Procericola bourgognei Crete (Samšinák, 1965) – Photia procustidis Rhodos (Samšinák, 1965) - Photia procustidis Holland (Buitendijk, 1945; Cooreman, 1954; Samšinák, 1965; Siepel, Cremers & Vierbergen, 2016) – Canestrinia dorcicola, Coleopterophagus megnini, Photia saetolata Hungary (Samšinák, 1965; Haitlinger, 1989, 1990d; 1992a) – Canestrinia damiani, Coleopterophagus megnini, Percanestrinia blaptis, P. sellnicki, Procericola bourgognei, Italy (contin.) (Berlese, 1881; Lombardini, 1943; Cooreman, 1950; Haitlinger, 1990d, 1992a, 2004) – Canestrinia carabicola, C. dorcicola, Coleopterophagus donaldi, C. megninii, Dicanestrinia knobi, Percanestrinia blaptis, P. maroccana, Photia procera, Ph. procustidis, Procericola bourgognei, P. ichthyoides, Pseudamansia chrysomelinus Sicily (Haitlinger, 1990d) - Canestrinia carabicola, Coleopterophagus megnini, Sardinia (Canestrini & Berlese, 1880; Vitzthum,1924) – Canestrinia carabicola, Percanestrinia blaptis, P. sardica Moldova (Trach, 2006) – Percanestrinia sellnicki, Photia hejniana, Procericola bourgognei Norway (Haitlinger, 2015) - Coleopterophagus albini Poland (Haitlinger, 1988a, 1988b, 1988c, 1988d, 1989, 1990c, 1990d, 1991c, 1992a, 1994a, 2002, 2004, 2012) - Canestrinia dorcicola, Coleopterophagus albini, C. megnini, Di- Acarorum Catalogus IX 221 canestrinia huberti, D. knobi, Percanestrinia blaptis, P. sellnicki, Photia adolfinae (as “polymorpha”), P. hejniana, P. chrysocarabi, P. hermengildae, P. bardoica, Pseudocanestrinia mahunkai, Procericola bourgognei Portugal (cont.) (Samšinák, 1971) – Canestrinia hispanica, Neophotia drvotocka Romania (Samšinák, 1965; Haitlinger, 1990d) – Coleopterophagus albini, C. maroni, Percanestrinia sellnicki, Procericola bourgognei Russia (all, continent.)(Samšinák, 1971; Haitlinger, 1992b; Khaustov & Eidelberg, 2001) – Percanestriniella caucasica, Photia bilkorum, Ph. polymorpha, Ph. sibirica, Procericola bourgognei, Uriophela arieli Kunashir I. (Khaustov & Eidelberg, 2001) – Photia pacifica Sakhalin I. (Khaustov & Eidelberg, 2001) – Photia lopatini Serbia (Haitlinger, 19 ) – Coleopterophagus megnini Slovakia (Haitlinger, 1990 ; Samšinák, 1965, 1971; Zamec, 2014) – Coleopterophagus albini, Percanestrinia selnicki, Photia chrysocarabi, Ph. hejniana, Ph. procustidis, Procericola bourgognei Slovenia (Cooreman, 1950; Samšinák, 1971) – Photia hejniana, Ph. procera, Procericola ichthyoides, Pseudocanestrinia orientalis Spain (cont.) (Jolivet, 1954; Samšinák, 1971) - Canestrinia hispanica, Mesophotia p. penicillata, M. p. similis, Neophotia drvotocka, N. latissima, Photia lusitanica, Pseudamansia chrysomelinus Balearic Is. (Haitlinger, 1991a, 2001; Haitlinger & Šundić, 1918) - Camirohylla feziana [Ibiza, Formentera], Ibizella samsinaki [Ibiza], I. balearica [Ibiza], Pseudamansia chrysomelinus [Mayorca] Sweden (Lundquist, 1985) – Procericola bourgognei Turkey (all) (Samšinák, 1964, 1965, ; Haitlinger, 1990d) - Coleopterophagus megnini, Photia graeca, Ph. procera adanensis, Ph. procera turcica, Ph. procustidis, Procericola bourgognei, P. ichthyoides scabricola, P. i. transversicola Ukraine (Haitlinger, 1990, 2002; Khaustov & Eidelberg, 2001; Samšinák, 1971; Trach, 2006; Trach & Khaustov, 2007, 2011) – Amansiella trilobata, Canestrinia dorcicola, Coleopterophagus albini, C. donaldi, C. megnini, C. maroni, Dicanestrinia knobi, Paramansia bicornis, Paraphotia calosomae, Percanestrinia blaptis, P. sellnicki, Photia adolfinae, Ph. hejniana, Ph. hermengildae, Ph. polymorpha, Procericola bourgognei, Pseudamansia decorata, Pseudocanestrinia carabi, P. berndi AFRICA Algeria (Jolivet, 1954) - Pseudamansia chrysomelinus Botswana (Haitlinger, 1993) - Sorbinophela sandyi Burundi (Haitlinger, 1993) - Boetophila werneri Cameroon (Haitlinger, 1990, 1991, 1991d, 1993, 2011) - Boetophela erhardi, Cetonicola isadorae, Paraphagella grimaldi, P. indirae, P. odae, Pheliculophela roaldi, Saniothiana barumbaica Congo-Brazaville (Haitlinger, 1991d) – Cetonicola maritae, Paraphagella odae 222 Acarorum Catalogus IX Congo DR (Cooreman, 1953, 1955; Haitlinger, 1995) - Afrocanestrinia straeleni, Cetonicola hispidus, Coleopterophagus pulcher, Diplognatophilus africanus, Donnelaphontia suatoha Egypt (El-Bishlawy & Allam, 2003) – Percanestrinia egypti Ethiopia (Haitlinger, 1990, 1991d, 1993; Khaustov, 2006) – Barbiangia elongata, B. ethiopica, Diplognatophilus ethiopicus, Tamarangia ethiopica, T. nimfae Ghana (Haitlinger, 1991a, 1991d, 1993) - Boetophela ephraimi, Boleohylla garrica, Cetonicola orestesi Guinea (Haitlinger, 2010) - Pseudoparaphagella gerardi Kenya (Fain, 1987b) – Diplopodocoptes transkeiensis Liberia (Haitlinger, 1991d) - Cetonicola orestesi Madagascar (Haitlinger, 1989a, 1990) – Ambilohylla favosa, Athogavia nosiana Morocco (Cooreman, 1953; Samšinák, 1971; Haitlinger, 1991; Haitlinger & Šundić, 1918) – Canestrinia hispanica,Camirohylla feziana, Globosophotia bettinae, Ibizella asilahica Mesophotia penicillata tertia, Percanestrinia maroccana, Pseudamansia chrysomelinus Senegal (Haitlinger, 1995a) - Afrocanestrinia samsoni Sierra Leone (Haitlinger, 1992) - Tamarangia fabiolae South Africa (Trägårdh, 1906; Fain, 1987; Haitlinger, 1989, Haitlinger & Chmielewski, 2004) – Boetophela cassandrae, B. ginae, Diplopodocoptes transkeyensis, Megacanestrinia mucronata, Tamarangia nimfae Sudan (Trägårdh, 1904) – Coleopterophagus quadrisetosus Tanzania (cont.)(Lavoipierre, 1958; Haitlinger, 1989, 1990b, 1991, 1995a) - Afrocanestrinia rufinae, Athogavia gamana, A. tanzanica, Barbiangia alvari [Zanzibar], Megacanestrinia beloniana, Boetophela robertsoni, B. silvanae, B. sybillae, B. symeoni, Cetonicola vatus, Irmongia helgae, Olgattia useguaica Zanzibar (Haitlinger, 1993) - Barbiangia alvari Tunisia (Samšinák, 1970; Haitlinger, 2000) - Bircericola bertrami, Percanestrinia kamelskyi Uganda (Haitlinger, 1991) - Boleohylla kiogana “Africa” (Berlese, 1910, 1920; Haitlinger, 1990) – Canestrinia remigans, Coleopterophagus neglectus, Saniothiana pycnosa ASIA China (Samšinák, 1971; Haitlinger, 1990d, 1992a, 1994a, 1998; Fan, Chen & Wang, 2010) - Beeriphotia volkeri, Canestrinia kacperi, C. paavoi, C. pekingensis, C. ramoni, Coleopterophagus belzebubi, C. rudolfi, Gioharattia maharae, Percanestrinia melchiori, P. viviannae, Photia melchiori Hainan (Summers & Schuster, 1982a;) – Melisia sentosa Taiwan (Samšinák, 1971; Summers & Schuster, 1981a; Haitlinger, 1990d; Fan, Chen & Wang, 2010) – Apalotacarus aristatus, C. cristatus, A. trullus, Canestrinia illegalis, Coleopterophagus belzebubi, Melisia superba Cyprus (Samšinák, 1965; Haitlinger, 1992a) – Ibizella [Canestrinia] samsinaki, Percanestrinia blaptis, Photia procustidis Acarorum Catalogus IX 223 India (Samšinák, 19 1971; Summers & Schuster, 1981a; Haitlinger, 1990, 1994a; Vishnupriya & Mohanasundaram, 1988) - Apalotacarus aristatus, A. echinatus, A. protensus, A. scissus, “Coleopterophagus procerus”, Canestriniella batocerae (Tamil Nadu), Melisia anomala, M. klapaleki, Percanestriniella vlachiana (Kashmir), Shillongia marzenae (Assam), Sandrophela sophiae Indonesia (Berlese, 1910; Samšinák, 1963, 1944; Summers & Schuster, 1981a; Haitlinger, 1989a,1990b, 1991, 1992b, 1993b, 1994a, 1995) – Albinorattia igori [Batjan], Amboinophela bernae [Amboina, New Guinea], Apalotacarus aristatus [Java], A. fulgens [Java], A. sajanae [Java], Arraphosoma paraminax [Mollucas], Athogavia borneoica [Borneo, presumably Kalimantan], A. scagana [Borneo, presumably Kalimantan], Aurilossongia dioni, A. hoaria [Kalimantan], Canestriniella amplexans [Java], C. physsana [Java], C. elae, C. phoebinae [Java], C. togata phoebinae [Java], C. phoebinae [Java], Gioharattia maharae [Borneo, presumably Kalimantan], Hargeinella poppana [Sumatra], Hyllosihemia belerofoni [Sumatra], Lidiophela bognae, L. celinae, L. pelagiae, Melisia jadwigae [Moluccas], Melisia ariadnae, M. gabrielae, M. hieronimae, M. ingeborgae, M. irenae, M. lombardinii, M. mariettae, M. melaniae, M. salomeae, M. sumatrae, Noemiphela barryi [Sumatra], N. izabelae [Sulawesi], Rugoniphela marloni [Borneo, Java, Sumatra], Sandrophela amarae [Java, Sumatra], S. hugoni [Sumatra], S. spectanda [Java, Sumatra], Tamarophela batianica [Moluccas], Vandeinella adriasae [Sumatra], V. opressa [Borneo] Iran (Haitlinger, 1990c; Ardeshir et al., 2016) – Camirohylla sp., Coleopterophagus baali Israel and Palestina (Samšinák, 1965) - Dicanestrinia knobi, Photia procustidis Japan (Kishida, 1924, Samšinák, 1971; Haitlinger, 1990d) - Canestrinia pictura, Coleopterophagus belzebubi, Photia polymorpha Kambodia (Vomero, 1972) – Melisia samsinaki Kirghizstan (Khaustov & Eidelberg, 2001) - Percanestriniella levis Korea (Samšinák, 1971; Wontae Kim et al., 2006) - Haitlingeria longilobata, Photia polymorpha Lebanon (Samšinák, 1964, 1965) - Photia procera marani, Ph. procustidis Malaysia (cont.) Sarawak and Sabah (Haitlinger, 1991) - Rugoniphela marioni [Borneo] Philippines (Summers & Schuster, 1981a) - Apalotacarus fusulus [Mindoro], A. scaurus [Mindanao] Sri Lanka (Haitlinger, 1989a) – Gioharattia caroae Syria (Haitlinger, 1990d) – Coleopterophagus dionizyi, C. maroni, Taiwan (Summers & Schuster, 1981a) – Apalotacarus aristatus, A. trullus Thailand (Haitlinger, 1990) - Melisia hieronimae Uzbekistan (Samšinák, 1971) – Percanestrinia levis Vietnam (Samšinák, 1969; Haitlinger, 1990b, 1993b) – Melisia baloghi, M. helenae, M. jadwigae, M.hieronimae, M. mahunkai, M.occidii, M. rufae, Sandrophela rodioni Asia (Haitlinger, 1989) 224 Acarorum Catalogus IX NORTH AMERICA Mexico (Banks, 1915; Summers & Schuster, 1979a, 1982b; Haitlinger, 1992e, 1993d, 1993e) – Acrotacarus mirabilis, Arrunsithiana escaudata, Bibulothiana cyriaki, B. vatia,Grandiellina marinae, G. soniae, Ennodithiana maniusi, Phriknodora pustulosa SOUTH and CENTRAL AMERICA Argentina (Haitlinger, 1992e, 1993e) - Grandiella tacjannae, Olafithiana foliacea, O. multifida Belize (Fain, 1987a) - Grandiella gambosa, G. kimsleyi Bolivia (Lombardini, 1950; Fain, 1987a; Haitlinger, 1989, 1992e, 1993d, 1993e, 1999) – Arrunsithiana escaudata, Bibulothiana brevispinata, Grandiella decemcaudata, G. longini, G. quadrata, G. rosascostai, Olafithiana multifida, Ovarrothiana leopoldi, O. ludomiri, Teklathiana onnae Brazil (Lombardini, 1939, 1950; Summers & Schuster, 1979a; Fain, 1987a; Haitlinger, 1989, 1990a, 1991e, 1992e, 1993d, 1993e, 1995a, 2008) – Arrunsithiana arcuata, A. battosi, A. escaudata, A. quadrata, A. spicantis, Bibulothiana brevispinata, B. cyriaki, B. everittae, B. oswaldi, Dorynothiana delmari, D. jurandi, Ennodithiana maniusi, E. medoni, E. megesi, Ennorithiana heliodori, Gasuthiana abapoica, G. coarica, G. melillae, Grandiella minor, G. tacjannae, G. tetracaudata, Hatohylla bahiana, H. bibiannae, H. morrata, Olafithiana adelinae, O. multifida, O. foliacea, O. joliveti, Ovarrothiana erazmi, O. ozannae, Sabothiana boi, S. reidari, S. riccardoi, Thopia boguwolae, Th. corinnae Colombia (Summers & Schuster, 1979a; Fain, 1987; Haitlinger, 1997, 1989, 1991b, 1991e, 1993d, 1993e, 2001) - Arrunsithiana ilosi, A. battosi, A. quadrata, Bibulothiana brevispinata, Danaithiana ayraica, Ennorithiana aidae, Gasuthiana olareica, Grandiella gambosa, G. kimseyi, G. sicula, G. soniae, G. tetracaudata, Ovarrothiana echinoderma, Teklathiana onnae, Costa Rica (Haitlinger, 1991b, 1994b; Summers & Schuster, 1979b) – Acrotocarus mirabilis, Arrunsithiana escaudata, Bibulothiana albertae, Danaithiana chanchamayoica, Olafithiana iwettae, Phriknodora avae Cuba (Haitlinger, 1991b, 1993d) - Danaithiana chanchamayoica, Ennodithiana messalinae Ecuador (contin.) (Fain, 1987a; Haitlinger, 1991b, 1991e, 1993d, 1993e,) – Arrunsithiana artemoni, A. battoni, A. erinnae, Bibulothiana brevispinata, Danaithiana archidonaica; D. ayraica, D. ecuadorica, Grandiella cooremani, G. ecuadorensis, G. soniae, Olafithiana multifida, Ovarrothiana ozannae, French Guyana (Lombardini, 1950; Fain, 1987a; Haitlinger, 1993e, 1993d, 1999) – Averithiana anoni, Arrunsithiana esacaudata, A. quadrata, A. squarrosa, Gasuthiana olareica, Grandiella tetracaudata, G. t. omoplata, Ovarrothiana erazmi, Thopia amedae Guatemala (Summers & Schuster, 1979a, 1979b) - Acrotocarus mirabilis, Arrunsithiana esacaudata Honduras (Haitlinger, 1993e, 1999) - Bibulothiana brevispinata, Metriothiana bolebori Acarorum Catalogus IX 225 Panama (Banks, 1915; Summers & Schuster, 1979b, 1982b; Haitlinger, 1993d, 1993e, ) – Bibulothiana brevispinata, Coleopterophagus blattophagus, Ennodithiana platysma, Grandiella gambosa,G. kimseyi, Ovarrothiana echinoderma, Phriknodora pustulosa, Teklathiana conula, T. rugosita, Thopia tenax Paraguay (Lombardini, 1950; Haitlinger, 1992e) – Grandiella decemcaudata, G. tacjannae Peru (Summers & Schuster, 1979; Fain, 1987a; Haitlinger, 1991b, 1992e, 1993d, 1993e, 2008) – Bibulothiana everittae, Danaithiana chanchamayoica, Grandiella rogeri, G. soniae, G. tetracaudata, Ovarrothiana ozannae, Peruthiana briannae Suriname (Haitlinger, 1993) – Ennodithiana penelopae Trinidad and Tobago (Turk, 1948) – Acrotacarus alutaceus, Venezuela (Lombardini, 1950; Haitlinger, 1994b) - Olafithiana iwettae, Grandiella pentagona South America (Fain, 1989) – Arrunsithiana latipes OCEANIA Australia (continent) (Summers & Schuster, 1981a, 1981b, 1982a; Haitlinger, 1991) — Apalotacarus fusticulus, A. gracilis, A. luroris, Cnecoderma dolichopoda, C. ovalis, Hypopteryx collaris, Lidiophela pauliana, Melisia leptinotarsa, Taseopus platythrix Papua New Guinea (Nesbitt, 1976; Summers & Schuster, 1981a, b; Haitlinger, 1989a, 1989b, 1990a, 1990b, 1993b, Haitlinger & Šundić, 2020) – Apalotacarus cidaris, A. fusulus, A. paxillus, A. petilus, A. rigescens, Aurilossongia bolanica, A. fileasi, A. gaurana [New Britain], A. norae, Lidiophela [Coleopterophagus] pecki, Cooremania kietaensis [Bougainville], C. wauensis, Jullongia izae, Passalophagus georgei, Phaleratus fentoni, Photia howdeni, Sajanophela alfredae New Guinea (Summers & Schuster, 1981b) – Arraphosoma minax Salomon Islands (Nesbitt, 1976; Summers & Schuster, 1981a) – Apalotacarus glaber [Russel I.], Lidiophela [Coleopterophagus] pecki Hosts of Canestriniidae Myriapoda Diplopoda Glomerida Fam. Glomeridae – Diplopodocoptes transkeiensis Spirostrepsida Fam. Odontopygidae - Diplopodocoptes transkeiensis Insecta Coleoptera Fam. Carabidae 226 Acarorum Catalogus IX Calosoma (Campalita) auropunctatum – Paraphotia calosomae Carabus (Oreocarabus) amplipennis - Photia lusitanica C. (Cychrostomus) anchocephalus – Canestrinia pekingensis C. (Eupachys) glyptopterus - Canestrinia pekingensis C. (Coptolabrus) lafossei saturatus - Canestrinia pekingensis C. augustus - Canestrinia pekingensis C. augustus ignigena - Canestrinia pekingensis C. pustulifer - Canestrinia pekingensis C. pustulifer pratti - Canestrinia pekingensis C. p. querryi - Canestrinia pekingensis C. p. szechwanensis - Canestrinia pekingensis C. ignimetalla angusticollis - Canestrinia pekingensis C. gemmifer - Canestrinia pekingensis C. (Cratocephenus) akinini – Percanestriniella levis C. angulicollis antaeus – Beeriphotia volkeri C. (Tachypus) auratus – Photia chrysocarabi C. (Chrysocarabus) auronitens - Photia chrysocarabi, Pseudocanestrinia occidentalis C. (Cratocephalus) balassogloi – Percanestriniella levis C. (C.) bessarabicus – Pseudocanestrinia carabi C. (Damaster) blaptoides – Canestrinia pictura C. (Imaibius) boysi – Percanestriniella vlachiana C. (Acanthaicus) brandti – Canestrinia pekingensis C. (Megodontus) caelatus – Pseudocanestrinia orientalis C. (C.) cicatricosus - Percanestriniella levis C. (Procrustes) coriaceus - Photia procera, Percanestrinia saetolata, Procericola bourgognei C. (Megodontus) croaticus – Pseudocanestrinia orientalis C. coelestes giganteus – Canestrinia ramoni C. (Procrustes) a. anatolicus – Photia procustidis C. (Procrustes) auticollis – Procericola bourgognei C. (Procrustes) b. banoni – Photia procustidis C. (Procrustes) ch. chevrolati – Procericola bourgognei C. (Procrustes) clypeatus clypeatus - Procericola bourgognei C. (Procrustes) coriaceus banaticus - Percanestrinia sellnicki C. (Procrustes) c. coriaceus - Percanestrinia sellnicki, Photia saetolata C. (Procrustes) coriaceus foudrasi – Photia graeca C. (Procrustes) coriaceus gressorius - Percanestrinia sellnicki C. (Procrustes) coriaceus imminutus - Percanestrinia sellnicki C. (Procrustes) coriaceus impudicus - Photia graeca C. (Procrustes) coriaceus kindermanni - Photia graeca Acarorum Catalogus IX 227 C. (Procrustes) coriaceus pseudorugifer - Percanestrinia sellnicki C. (Procrustes) coriaceus rugifer - Percanestrinia sellnicki C. (Procrustes) coriaceus subrugosus - Percanestrinia sellnicki C. (Procrustes) coriaceus – Pseudocanestrinia mahunkai C. (Ctenocarabus) galicianus – Canestrinia hispanica C. (Procerus) gigas – Photia procera, Procericola ichthyoides C. (Procerus) scabrosus transversalis - Photia procera adanensis C. (Procerus) scabrosus caucasicus - Photia procera C. (Procerus) syriacus - Photia procera marani C. (C.) glabratus – Photia adolphinae C. grandis – Canestrinia paavoi C. (Oreocarabus) guadarramus - Canestrinia hispanica, Neophotia drvotocka C. (Megodontus) gyllenhali – Photia adolfinae, Pseudocanestrinia berndi C. (Coptolabrus) hankotaizanus – Canestrinia illegalis C. (Procrustes) impressus carmelita – Photia procustidis C. (Megodontus) kolbei – Photia pacifica C. (Tribax) komarovi – Photia bilkorum C. (Oreocarabus) linnaei – Photia hermengildae C. latenulatus plassensis - Pseudocanestrinia berndi C. (Coptolabrus) longipennis mandarinus - Canestrinia illegalis C. (Acoptolabrus) lopatini – Photia lopatini C. (Mesocarabus) lusitanicus – Neophotia latissima C. (M.) lusitanicus breuningi – Photia lusitanica C. (M.) lusitanicus castilianus – Photia lusitanica C. (M.) lusitanicus logronicus - Photia lusitanica C. (M.) lusitanicus schaumi - Photia lusitanica C. (Rhabdotocarabus) melancholicus – Canestrinia hispanica, Mesophotia p. penicillata C. (Macrothorax) morbillosus – Canestrinia carabicola C. (Procrustes) m. mulsatianus – Photia procustidis C. (Archicarabus) nemoralis – Photia bardoica, Ph. chrysocarabi C. (Procrustes) payafa - Procericola bourgognei C. (Tribax) plasoni – Photia bilkorum C. (Tribax) protensis – Photia bilkorum C. (Iniopachus) pyrenaeus – Neophotia drvotocka, N. jureceki C. (Procerus) scabrusus - Photia procera C. (Megodontus) schoenherri – Photia sibirica C. (Tribax) shamyli – Photia bilkorum C. (I.) stoliczkanus - Percanestriniella vlachiana C. (Ulocarabus) sturowskyi - Percanestriniella levis C. (P.) tanypedilus - Percanestriniella levis 228 Acarorum Catalogus IX C. (Pantophyrtus) turcomannorum - Percanestriniella levis C. (Hygrocarabus) variolosus – Dicanestrinia huberti, D. knobi C. (Megodontus) violaceus – Photia hejniana, Ph. saetolata C. v. bicinctus – Photia saetolata C. v. exasperatus – Photia hejniana, Ph. saetolata C. v. fioris – Photia hejniana C. v. fulgens – Photia saetolata C. v. obliquus - Photia hejniana C. v. purpurascens – Photia graeca, Ph. saetolata C. v. savinicus - Photia hejniana C. v. styriensis - Photia hejniana Pseudosteropus globosus – Globosophotia bettinae Sphodrus leucophthalmus - Percanestrinia blaptis Tefflus purpuripennis – Megacanestrinia beloniana Carabus sp. – Percanestriniella caucasica, Paracanestrinia denmarkica Carabidae gen. sp. – Megacanestrinia mucronata Fam. Cerambycidae Batocera albofasciata octomaculata – Canestriniella phoebinae B. celebiana - Canestriniella physsana B. ferruginea – Canestriniella elae B. hector – Canestriniella amplexans B. humilidens – Canestriniella physsana B. lunulata – Canestriniella togata B. procera - Canestriniella togata Brazilianus batus – Phriknodora pustulosa B. brazilianus - Phriknodora pustulosa Cerambyx cerdo – Dicanestrinia cerambycis Lagocheirus foveolatus – Thopia pannucea Neoptychodes trilineatus – Thopia bulbocaudata Sternotomis bohemanni – Anaspistes unguiculatus Taeniotes scalatus – Thopia tenax Trachyderes succinctus – Phriknodora scabra Xylotrupes asperulus – Aurilossongia bolanica Xylotrupes sp. – Aurilossongia gaurana, A. hoarica Cerambycidae gen. sp. – Chelinophora dictyophora Fam. Chrysomelidae Acromis nebulosa – Ennodithiana penelopae A. sparsa - Teklathiana onnae, T. rugosita A. spinifex – Ennodithiana medoni Agenyssa caedemadens – Ovarrothiana ozannae A. crassicornis - Ovarrothiana ozannae Acarorum Catalogus IX 229 Alurnus bipunctatus - Gasuthiana olareica A. boulsardi – Gasuthiana olareica A. clysianus – Gassuthiana abapoica Anacassis cribrum – Olafithiana adelinae A. fuscata - Grandiella tacjannae Botanochara angulata – Grandiella tacjannae B. bonariensis - Grandiella tacjannae B. convexiuscula - Grandiella tacjannae B. duodecimverrucata - Grandiella tacjannae B. impressa - Grandiella tacjannae B. macularia - Grandiella tacjannae B. nervosa - Grandiella tacjannae B. regina - Grandiella tacjannae B. ruforeticulata - Grandiella tacjannae B. sanguinea - Grandiella tacjannae B. sedecimpustulata - Grandiella tacjannae B. tesselata - Grandiella tacjannae B. texta - Grandiella tacjannae Canistra carbonaria – Grandiellopsis arriani C. cruentata – Grandiellopsis violae C. osculatii – Grandiellopsis alani, G. canistra C. plagosa – Arrunsithiana spicantis Charidotella kesseli – Sabothiana reidari Ch. (= Metriona) testudinacea – Olafithiana iwettae Charidotis furunculus - Sabothiana riccardoi Charidotis sp. – Sabothiana boi Chelymorpha comata – Bibulothiana albertae Ch. Infecta - Bibulothiana cyriaki Ch. inflata – Bibulothiana everittae Ch. nigricollis – Bibulothiana cyriaki Ch. rufoguttata - Bibulothiana everittae Ch. vermiculata - Bibulothiana cyriaki Chrysolina fastuosa – Paramansia bicornis Ch. mentastri – Paramansia bicornis, P. mentastri Ch. polita - Paramansia bicornis Coptocycla adamantina – Grandiella tetracaudata C. contenta - Grandiella decemcaudata C. nigropunctata - Grandiella sicula C. wagneri - Grandiella sicula Coptocycla sp. – Ennodithiana platysma Crambelea illuderis – Ennodithiana megesi 230 Acarorum Catalogus IX Cyclosoma mirabilis – Grandiellopsis orsenae C. paliata – Grandiellopsis cyclosoma C. tristis – Ovarothiana ludomiri Cyrtonota cyanea - Grandiella tetracaudata C. dissecta – Arrunsithiana ilosi C. latevittata – Arrunsithiana artemoni C. punicea – Grandiella soniae C. textilis - Grandiella soniae C. thalassina - Grandiella tetracaudata C. tigrina - Grandiella tetracaudata C. tristigma – Bibulothiana vatia Discomorpha batesi - Grandiella gambosa D. lanuginosa pubescens - Grandiella gambosa D. depilata - Grandiella gambosa Dolichotoma conspersipennis - Grandiella gambosa Dorynota pugionata – Dorynothiana jurandi Dorynota sp. – Dorynothiana delmari Echoma [as Omoplata] marginata – Arrunsithiana escaudata, Grandiellina decemcaudata E. clypeata – Grandiella tetracaudata E. confluens – Grandiella marinae Elytrosphaera xanthopyga – Olafithiana joliveti Eudicella gralli var. meckowi – Paraphagella eudicellae E. smithi var. bertherandi – Paraphagella eudicellae, P. minor Eugenysa andicola – Ovarrothiana ozannae E. bacchus - Ovarrothiana erazmi E. columbiana – Ovarrothiana echinoderma, Arrunsithiana quadrata E. delicata - Ovarrothiana nikodemi E. divalis – Ovarrothiana nikodemi E. grossa - Ovarrothiana erazmi, Arrunsithiana squarrosa E. venosa – Ovarrothiana erazmi, Thopia amedae Eumolpus fulgidus - Grandiella tetracaudata Goniochenia (Baranosa) decolor – Grandiella rogeri G. (B.) heroldi - Grandiella rogeri G. (B.) flavosparsa - Grandiella longini Mecistomala marginatus – Gassuthiana coarica M. nigripes – Gassuthiana abufarica Metriona testudinacea – Olafithiana iwettae Metrionella bilimeki - Metriothiana bolebori Myocalaspis gentilis – Ovarrothiana leopoldi Omaspides basilica - Ennodithiana messalinae O. bistriata – Teklathiana conula Acarorum Catalogus IX 231 Physonota alutacea – Acrotacarus alutaceus, A. mirabilis Ph. diluta – Acrotacarus crataepus Plagiometriona ludiera – Ennorithiana heliodori P. punctipennis - Ennorithiana heliodori Polychalca dentipennis – Thopia corinnae P. punctatissima – Grandiellopsis odiosi, Thopia boguwolae Psalidonota contemta – Ennodithiana maniusi P. dorsoplagiata - Ennodithiana messalinae P. dorsopunctata – Ennodithiana messalinae P. leprosa - Ennodithiana messalinae Pycnocerus sp. – Saniothiana pycnosa Stolas aenea - Arrunsithiana quadrata S. antiqua – Olafithiana multifida S. areolata - Olafithiana foliacea S. aurosetosa - Olafithiana multifida S. bifenestrata - Bibulothiana brevispinata S. bipunctata - Arrunsithiana quadrata S. bipustulata - Bibulothiana brevispinata S. chalybaea – Arrunsithiana spicantis, A. battosi, Grandiella tetracaudata, Olafithiana foliacea S. coalita - Grandiella soniae S. coerulescens – Grandiella soniae, Bibulothiana brevispinata S. conspersa - Olafithiana foliacea, Arrunsithiana battosi, A. spicantis S. cruentata - Olafithiana multifida, Arrunsithiana battosi S. decemguttata - Arrunsithiana arcuata S. demissa - Arrunsithiana quadrata S. ephippium - Bibulothiana brevispinata S. erichsoni - Grandiella soniae, Arrunsithiana battosi S. esignata - Arrunsithiana arcuata S. funebris - Arrunsithiana battosi, A. spicantis S. gibbosa - Arrunsithiana battosi S. huanocensis - Arrunsithiana battosi S. ignita - Arrunsithiana battosi S. imitans - Arrunsithiana quadrata S. implexa - Olafithiana foliacea S. impluviata - Olafithiana foliacea S. inaequalis – Arrunsithiana latipes, A. quadrata S. indigacea - Arrunsithiana quadrata S. inexculta - Arrunsithiana battosi S. lacunosa - Olafithiana foliacea S. latevittata – Arrunsithiana artemoni S. lebesi – Arrunsithiana escaudata, Phriknodora avae 232 Acarorum Catalogus IX S. lenis - Olafithiana foliacea, Arrunsithiana battosi S. lineaticollis - Bibulothiana brevispinata S. niobe - Arrunsithiana battosi S. nudicollis - Olafithiana foliacea S. oblita - Olafithiana foliacea, Arrunsithiana battosi S. paranensis - Olafithiana foliacea, Arrunsithiana battosi S. pectinata – Arrunsithiana erinnae S. placida - Grandiella soniae S. plagiata - Bibulothiana brevispinata S. plagicollis – Bibulothiana oswaldi S. pleurosticha - Arrunsithiana battosi S. punicea – Arrunsithiana escaudata, Grandiellina soniae S. redtenbacheri - Arrunsithiana quadrata S. scoparia - Arrunsithiana battosi S. subreticulata - Olafithiana foliacea S. subrugosa - Grandiella tetracaudata S. thoreyi - Arrunsithiana quadrata Stolas sp. – Arrunsithiana squarrosa Timarcha atlantica - Pseudamansia chrysomelinus T. balearica - Pseudamansia chrysomelinus T. goettingensis - Pseudamansia chrysomelinus, P. decorata T. interstitialis - Pseudamansia chrysomelinus T. nicaensis - Pseudamansia chrysomelinus T. pauperata - Pseudamansia chrysomelinus T. punctella - Pseudamansia chrysomelinus T. rugosa - Pseudamansia chrysomelinus T. tenebricosa – Amansiella trilobata, Pseudamansia chrysomelinus Zatrephina lineata – Averithiana anoni Hispinae gen. sp. – Peruthiana briannae Chrysomelidae gen. sp. – Acrotacarus mirabilis Fam. Colydiidae Colydium sp. – Sandrophela dittae Fam. Elateridae Semiotus furcatus - Danaithiana ecuadorica, D. archidonaica S. imperialis - Danaithiana chanchamayoica S. virgatus - Danaithiana ayraica Fam. Lucanidae Cladognathus giraffa - Noemiphela izabelae Dorcus bucephalus – Sandrophela spectanda D. paralellipipedus – Canestrinia dorcicola Eumelasomus sansibaricus -Olgattia useguaica Acarorum Catalogus IX 233 Eurytrachelus anthaeus – Rugoniphela marloni Eurytrachelus bucephalus – Rugoniphela marloni Eurytrachelus eurycephalus - Rugoniphela marloni, Sandrophela amarae Eurytrachelus gypeatus - Sandrophela amarae Eurytrachelus platymelus - Sandrophela rodioni Eurytrachelus purpurescens - Noemiphela barryi Hemisodorcus chevrilati – Sandrophela sophiae Hemisodorcus [Macrodora] rubrofemoratus - Uriophela arieli Metapodonotus bison - Amboinophela bernae Metapodonotus cinctus - Amboinophela bernae Odontolabis wollastoni – Sandrophela hugoni Serrognathus platymelus – Haitlingeria longilobata Telodorcus saiga – Canestrinia microdisca Lucanidae indet. - Vereoxia bogeri, Cooremania kietaensis Fam. Passalidae Aceraius grandis – Apalotacarus aristatus, Melisia mariettae, M. salomeae A. grandis hirsutus – Apalotacarus cristatus, Melisia superba A. helferi – Melisia samsinaki A. laevicollis – Melisia gabrielae A. mayeri – Melisia melaniae Epiphanus indicus – Melisia anomala E. stoliczkae - Melisia anomala, M. klapaleki Eriocnemis tridens – Apalotacarus sajanae Eumelasomus sansibaricus – Olgattia useguaica Gonatus noviculator - Teophyssa honoratae Labienus gigax – Lidiophela igae, L. pelagiae L. ptox – Apalotacarus fusulus, Lidiophela arnica Labienus sp. - Passalophagus georgei, Lidiophela [Coleopterophagus] pecki Leptaulax bicolor formosanus - Apalotacarus trullus L. indicus – Shillongia marzenae Mastochilus australasiens – Apalotacarus luroris M. quaestionis - Apalotacarus luroris Macrolinus nicobaricus – Apalotacarus protensus M. weberi – Apalotacarus scaurus Obiensus sp. – Lidiophela [Coleopterophagus] pecki Pelopides mniszecki – Melisia lombardinii P. tridens – Apalotacarus fulgens Pharochilus dilatus – Apalotacarus luroris Ph. nitidulus – Apalotacarus fusticulus, A. luroris Pleurarius brachyphyllus – Apalotacarus echinatus, A. scissus, Phaleratus alatus 234 Acarorum Catalogus IX Protomocoelus australis – Apalotacarus cidaris, A. petilus, Sajanophela alfredae, Lidiophela [Coleopterophagus] pecki P. triumphator - Apalotacarus glaber Passalidae gen. sp. – Apalotacarus gracilis, A. paxillus, A. rigescens, Arraphosoma minax, A. paraminax, Cnecoderma dolichopoda, C. ovalis, Hyllosihemia belerofoni, Jowitella barbarae, Jullongia izae, Lidiophela bognae, L. celinae, Melisia helenae, M. ingeborgae, M. irenae, M. jadwigae, M. leptinotarsa, M. rutae, M. sentosa, Tamarophela batjanica Fam. Passandridae Passandra trigemina – Sandrophela kokodaica Fam. Scarabaeidae Caelorrhina thoreyi – Pseudoparaphagella gerardi Cetonia aurata – Coleopterophagus megnini Coglodera diaroi – Gioharattia caroae Coryphocera sp. – Gioharattia caroae Dicronorhina micans – Afrocanestrinia straeleni, Cetonicola hispidus, Paraphagella princeps D. oberthuri – Boetophela robertsoni Diplognatha gagates – Barbiangia elongata Elasocheilon haterospila – Boetophela symeoni Eudicella smithi – Boetophela ginae E. gralli var. meckowi – Canestrinia hispidus Geynodonta laeviplaga – Boetophela silvanae Goliathus alborignathus – Boetophela sybillae Goliathus goliathus - Phelliculophela roaldi Ischipsophia olivacea – Albinorattia igori Mecynorrhina [Chelarrhina – recte Chelorrhina] savagei – Boetophela erhardi, Cetonicola isadorae, Paraphagella indirae Meristomala marginatus – Gasuthiana coarica M. nigriceps – Gasuthiana abufarica Oryctes monoceros – Athogavia gamana, A. tanzanica O. pyrrhus - Athogavia nosiana Oryctes sp. – Athogavia scagana Oxythirea funesta – Canestrinia dorcicola Pachnoda abyssinica – Tamarangia ethiopica P. carmelita – Kahoorangia almae P. marginata – Tamarangia ethiopica, T. fabiolae, T. flawiani P. thoracica – Tamarangia nimphae Papuana woodlarkiana – Oceogavia ubaldi Plaesiorrhina mhondana – Boetophela werneri Acarorum Catalogus IX 235 Poecilophila hebraea – Cetonicola vatus P. maculatissima – Sorbinophela sandyi Porphyroneta cinnamonea – Coleopterophagus pulcher Protaetia (Potosia) aerata - Coleopterophagus belzebubi Protaetia (Potosia) affinis – Coleopterophagus donaldi, C. megnini Protaetia (Potosia) brevitarsis - Coleopterophagus rudolfi Protaetia (Potosia) cuprea – Coleopterophagus dionizyi Protaetia (Potosia) cuprea ignicollis - Coleopterophagus megnini Protaetia (Potosia) cuprea cuprea - Coleopterophagus megnini Protaetia (Potosia) cuprea obscura - Coleopterophagus megnini Protaetia (Potosia) cuprina - Coleopterophagus megnini Protaetia (Potosia) fieberi - Coleopterophagus megnini Protaetia (Potosia) metallica - Coleopterophagus megnini, C. albini Protaetia (Potosia) speciosa – Coleopterophagus baali Protaetia (Potosia) speciosa jousselini – Coleopterophagus maroni Protaetia (Cetonischema) aeruginosa – Coleopterophagus albini, C. maroni Protaetia (Eupotosia) affinis – Coleopterophagus donaldi, C. megnini Protaetia (Liocola) marmorata - Coleopterophagus albini Rhabdotis sobrina – Diplognathophilus africanus Scarabaeus variolosus – Scarabiphotia vignoni Smaragdesthes africana mutica – Paraphagella princeps S. oertzeni – Afrocanestrinia rufinae Stephanorrhina guttata – Cetonicola maritae, Paraphagella odae Tmesorrhina iris – Cetonicola orestesi, Paraphagella ingridae Trichogomphus lunicollis – Vandeinella opressa T. milon – Hargeinella poppana, Vandeinella adriassae T. nartolani – Athogavia assamica “Scarabeus” centaurus – Coleopterophagus neglectus Scarabaeidae (Cetoniinae) – Babiangia alvari, B. ethiopica, Boetophela cassandrae, B. ephraimi, Coleopterophagus quadrisetosus Scarabaeidae (Dynastinae) – Athogavia arybbasi, Aurilossongia fileasi Scarabaeidae gen. sp. – Anaspistes unguiculatus Fam. Tenebrionidae Amarygmus striatus – Lidiophela pauliana Blaps appendiculata – Camirohylla feziana B. gibbosa – Percanestrinia blaptis, P. izabelae B. hejconota cortosa – Percanestrinia norodomi B. lethifera - Percanestrinia blaptis B. mucronata – Percanestrinia maroccana B. occulta - Percanestrinia blaptis B. pinguis - Percanestrinia maroccana B. polychresta – Percanestrinia egypti 236 Acarorum Catalogus IX B. producta - Percanestrinia maroccana B. punctata - Percanestrinia blaptis Blaps sp. – Ibizella [Canestrinia] samsinaki, Ibizella balearica Damatria formosus – Ambylohylla favosa Dila sp. - Camirohylla sp. Gnaptor sp. – Ibizella [Canestrinia] samsinaki Metallonotus metallicus – Boleohylla garrica Nyctobates gigas - Hatohylla morrata N. maximus – Hatohylla bahiana Nyctobates sp. – Grandiella [Hatohylla] rosascostai Odontopezus cupreus – Donnelaphontia cuatoha Odontopezus cupreus lucens - Saniothiana barumbaica Psammodes carbonarius – Boleohylla kiogana Tenebrionidae gen. sp. – Bicercicola bertrami, Gasuthiana melillae, Hatohylla bibiannae Coleoptera indet. – Ennodithiana platysma, Grandiella rugosita, Melisia baloghi, M. helenae, Phaleratus fentoni Bibliography of Canestriniidae Ardeshir F., I. Hasanvand, R. Haitlinger, S. Jafari. 2016. Camirochylla (Acari: Astigmata: Canestriniidae), a new mite genus to fauna of Iran. - Persian Journal of Acarology, 5(2): 159–160. Banks N. 1915. A new genus of Canestriniidae (Acari). – Entomological News, 26: 152–153. Beran F., S. Geiselhardt, G. Vargas, D. Windsor. 2014. 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Dicanestrinia knobi Samšiñák, 1971 i Coleopterophagus megnini (Berlese, 1881)(Acari, Astigmata, Canestriniidae) dwa gatunki roztoczy nowe dla fauny Polski. – Przegl. Zool., 32: 535–540. Haitlinger R. 1989a. New Canestriniid mites (Acari, Astigmata, Canestriniidae) connectes with beatles of the subfamilies Dynastinae and Cetoniinae (Insecta, Coleoptera, Scarabaeidae). – Wiadomości Parazytologiczne, 35(1): 43–60. Haitlinger R. 1989b. New canestriniid mites (Acari, Astigmata, Canestriniidae) associated with beetles of the subfamily Hispinae (Insecta, Coleoptera, Chrysomelidae). – Pol. Pismo ent., 59: 281–292. Haitlinger R. 1989c. New canestriniid mites (Acari, Astigmata, Canestriniidae) associated with Scarabaeidae and Cerambycidae (Insecta, Coleoptera) from the Indo – Australian region. – Wiadomości Parazytologiczne, 35: 585–601. Haitlinger R. 1989d. New Canestriniid mites (Acari, Astigmata, Canestriniidae) connectes with beatles of the genera Oryctes, Trichogomphus and Pentodon (Insecta, Coleoptera, Scarabaeidae, Dynarinae). - Wiadomości Parazytologiczne, 35: 337–355. Haitlinger R. 1990a. Hatohylla n. gen. (Acari, Astigmata, Canestriniidae) with description of three new species. – Rev. Ibér. Parasitol., 50(3–4): 307–312. Haitlinger R. 1990b. New Canestriniid Mites (Acari, Astigmata, Canestriniidae) associated with passalid beetles (Insecta, Coleoptera, Passalidae). – Bull. Entomol. Pologne, 59: 527–589. Haitlinger R. 1990c. The genus Coleopterophagus Berlese, 1882 (Acari, Astigmata, Canestriniidae) with descriptions of seven new species and with key for species determination. – Ann. zool., 43: 327–341. Haitlinger R. 1990d. New canestriniid mites (Acari, Astigmata, Canestriniidae) associated with beetles of the families Carabidae, Scarabaeidae, Tenebrionidae and Passandridae (Insecta, Coleoptera). – Ann. Zool., 43: 311–325. Haitlinger R. 1991a. New Canestriniid Mites (Acari, Astigmata, Canestriniidae) associated with some Tenebrionidae and Carabidae (Insecta, Coleoptera). – Zootechnika, 35(206): 273–281. Haitlinger R. 1991b. New Canestriniid Mites (Acari, Astigmata, Canestriniidae) connected with Elateridae (Insecta, Coleoptera). – Zeszyty naukowe Akademii Rolnicziej we Wroclawiu, Zootechnika, 35(206): 253–263. Haitlinger R. 1991c. List of mites occuring on insects in Poland. – Wiadomości Parazytologiczne, 37(1): 85–90. Acarorum Catalogus IX 239 Haitlinger R. 1991d. Fifteen new castriniid mites (Acari, Astigmata, Canestriniidae) associated with beetles of the Colydiidae and Cetoniinae (Scarabaeidae) (Insecta, Coleoptera). – Wiadomosci Parazytologiczne, 37: 281–305. Haitlinger R. 1991e. Canestriniid mites (Acari, Astigmata, Canestriniidae) occurring on Chrysomelidae (Insecta, Coleoptera) in Neotropical Region. I. Genera Grandiellina Fain, Arrunsithiana n.gen. - Wiadomosci Parazytologiczne, 37(3): 381–399. Haitlinger R. 1992a. The genus Percanestrinia Berlese, 1911 (Acari, Astigmata, Canestriniidae) with description of three new species. – Zool. Jb. Syst., 119: 535–542. Haitlinger R. 1992b. Rugoniphela n. gen., Noemiphela n. gen. and Uriophela n. gen. three new genera of Canestrinid mites (Acari, Astigmata) from Asia. – Redia, 74 [1991]: 533–542. Haitlinger R. 1992c. A preliminary description of new species of Grandiellopsis Fain, 1989 (Acari, Astigmata, Canestriniidae) associated with Cassidinae (Insecta, Coleoptera, Chrysomelidae) from Neotropical Region. – Boll. Mus. Reg. Sci. nat. Torino, 10(1): 51–61. Haitlinger R. 1993a. New genera and species of Afrotropical Canestriniidae (Acari, Astigmata). – Spixiana, 16(1): 5–17. Haitlinger R. 1993b. New genus and two new species of Canestriniid Mites (Acari, Astigmata) associated with Lucanidae (Insecta, Coleoptera). – Mitt. Zool. Mus. Berl., 69(2): 339–344. Haitlinger R. 1993c. Acari (Arachnida) and Anoplura (Insecta) collected on small mammals, reptiles and insects in Greece and Cyprus. – Biologia Gallo-hellenica, 20: 83–88. Haitlinger R. 1993d. Canestriniid mites (Acari, Astigmata, Canestriniidae) occurring on Chrysomelidae (Insecta, Coleoptera) in Neotropical Region. II. Genera Ovarrothiana n.gen., Ennodithiana n.gen. - Wiadomosci Parazytologiczne, 39(1): 59–73. Haitlinger R. 1993e. Canestriniid mites (Acari, Astigmata, Canestriniidae) occurring on Chrysomelidae (Insecta, Coleoptera) in Neotropical Region. III. Genera Thopia Summers et Schuster, Sabothiana n.gen., Olafithiana n.gen., Bibulothiana n.gen. - Wiadomosci Parazytologiczne, 39(3): 265–283. Haitlinger R. 1994a. New species mites (Acari, Astigmata, Canestriniidae) associated with Lucanidae, Carabidae and Scarabaeidae (Insecta, Coleoptera). – Wiadomości Parasytologiczne, 40(2): 193–213. Haitlinger R. 1994b. New Canestriniid mites (Acari, Astigmata) from Costa Rica. – Brenesia, 41–42: 81–93. Haitlinger R. 1995a. New species of mites collected on Tenebrionidae, Scarabaeidae, Cerambycidae and Lucanidae with description of new subfamily Vereoxiinae (Acari, Astigmata, Canestriniidae). – Wiadomości Parasytologiczne, 41(1): 79–91. Haitlinger R. 1995b. New mites (Acari, Astigmata: Canestriniidae; Prostigmata: Erythraeidae, Trombidiidae, Microtrombidiidae) for the fauna of Austria, Germany and Hercegovina with descriptions of four new species. – Linzer biol. Beitr., 27(1): 259–272. Haitlinger R. 1998. Four new species of canestriniid and heterocoptid mites from China (Acari, Astigmata, Canestriniidae, Heterocoptidae). – Spixiana, 21: 1–10. Haitlinger R. 1999. New Canestriniid Mites (Acari, Astigmata, Canestriniidae) from Neotropical region. – Zootechnika, 45(362): 35–56. 240 Acarorum Catalogus IX Haitlinger R. 2000. A new canestriniid mite Bircericola bertrami n. g., n. sp. 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The relations between mites and insects. Int. Meeting of Acarologists, Warszawa, II. Samšinák K. 1969. Zoologische Ergebnisse der Forschung von Dr. T. Pócs in der Demokratischen Republik Vietnam. – Zool. Anzeiger, 182: 417–416. Samšinák K. 1970a. Die auf Blaps (Col., Tenebrionidae) lebenden Milben der Gattung Canestrinia Berlese, 1881 (Acari). – Entomol. Mitt. Zool. Mus. Hamburg, 4(68): 71–78. Samšinák K. 1970b. Vikariirende Milben unter den Symphylen der Käfer Mitteleuropas. – Polskie pismo entomologiczne, 40(3): 601–603. Samšinák K. 1971. Die auf Carabus – Arten (Coleoptera, Adephaga) der palearktischen Region lebende Milben der Unterordnung Acariformes (Acari); ihre Taxonomie und Bedeutung für die Lösung zoogeographisher, entwicklungsgeschichtlicher und parasitophyletischer Fragen. – Entomol. Abhandlungen, 38(6): 1–25. Santiago-Blay J.A., A. Fain. 1994. Phoretic and ectoparasitic mites (Acari) of the Chrysomelidae. In: Jolivet P.H., Cox M.L., Petitpierre E. (Eds). Novel aspects of the biology of Chrysomelidae, Series Entomologica book series (SENT, volume 50): 407–417. Siepel H., H. Cremers, B. Vierbergen. 2016. Provisional checklist of the astigmatic mites of the Netherlands (Acari: Oribatida: Astigmatina). – Nederlandse Faunistische Mededelingen, 47: 49–88. Summers F.M., R.O. Schuster. 1979a. Eleven new species of Grandiella Lombardini (Acari, Canestriniidae). – Intern. J. of Acarology, 5(4): 325–350. Summers F.M., R.O. Schuster. 1979b. Revision of Acrotacarus Banks (Acari: Canestriniidae). - Proc. Entom. Soc. Washington, 81: 475–485. Summers F.M., R.O. Schuster. 1981a. Apalotacarus, a new genus of canestriniid mites from passalid beetles. – Internat. J. of Acarology, 7(1–4): 17–38. Summers F.M., R.O. Schuster. 1981b. Three new genera of canestriniid mites associated with passalid beetles. - Internat. J. of Acarology, 7(1–4): 39–46. Summers F.M., R.O. Schuster. 1982a. The Melisiinae, new subfamily of mites in the Canestriniidae associated with passalid beetles. - Internat. J. of Acarology, 8(1): 23–31. Summers F.M., R.O. Schuster. 1982b. New canestriniid mites from beetles of the family Cerambycidae. – Int. J. Acarol., 8(1): 33–46. Théodoridès J. 1955. Contribution à l’étude des parasites et phorétiques de coléoptères terrestres. – Actualités scientifiques et industrielles 1217, Paris, Hermann & Cie, Eds. - Vie et Milieu, Supplements 4: 1–310. Acarorum Catalogus IX 243 Trach V.A. 2006. The host-parasite associations of the mites family Canestriniidae (Acari: Astigmata) in Ukraine and Moldova. – Vestnik odesskogo nat. universiteta. Biologiya, 11(9): 174–180 (in Ukrainian, sum. Russ., Engl.). Trach V.A., A.A. Khaustov. 2007. A review of mites of the family Canestriniidae (Acari, Astigmata) associated with the leaf beetles (Coleoptera, Chrysomelidae) in Ukraine with description of a new species of the genus Pseudamansia. – Vestnik Zoologii, 41(3): 267–272. Trach V.A., A.A. Khaustov. 2011. A review of the genus Coleopterophagus Berlese, 1882 (Acari: Astigmata: Canestriniidae) of Ukraine. – Acarina, 19(2): 213–230. Trägårdh I. 1906. Neue Acariden aus Natal and Zululand. – Zool. Anzeiger, 30: 870–887. Turk F. A. 1948. Insecticolous Acari from Trinidad, B. W. I. – Proc. Zool. Soc. London, 118, art 1: 82–125. Turk F.A. 1953. A synonymic catalogue of British Acari: Part II. – Annals and Magazine of Nat. History, 12 ser., 62, 6: 81–99. Vishnupriya R., M. Mohanasundaram. 1988. Mites associated with insects in Tamil Nadu, India. – Entomon, 13(3–4): 247–257. Vitzthum H.G. 1925. Acarologische Beobachtungen. 8. Reihe. – Arch. Naturgesch., 90 (1924): A(10): 1–86. Vomero V. 1972. Una nuova specie del genere Melisia Lombardini, 1944, associata a coleotteri passalidi (Acarina – Sarcoptiformes). – Acarologia, 14(1): 95–108. The Mite Family Chetochelacaridae Fain Chetochelacaridae Fain, 1987: 161 One genus and species of African mites associated with Diplopoda. Fam. Chetochelacaridae Fain Chetochelacaridae Fain, 1987: 161 Type genus: Chetochelacarus Fain, 1987 Genus Chetochelacarus Fain Chetochelacarus Fain, 1987: 162 Type species: Chetochelacarus mamillatus Fain, 1987 Chetochelacarus mamillatus Fain Chetochelacarus Fain, 1987: 162 Type locality: Kivu, Djuma Valley Distribution: D.R. Congo (former Zaire) Host: millipede (Diplopoda: Julidae) 244 Acarorum Catalogus IX Index of Chetochelacaridae (Names in bold are accepted as valid) Chetochelacaridae 243 Chetochelacarus 243 mamillatus, Chetochelacarus 243 Distribution of Chetochelacaridae AFRICA D.R. Congo (Fain, 1987) – Chetochelacarus mamillatus Hosts of Chetochelacaridae Myriapoda Diplopoda Julida Fam. Julidae Gen. sp. indet. - Chetochelacarus mamillatus Bibliography of Chetochelacaridae Fain A. 1987. Notes on mites associated with Myriapoda 1. Three new astigmatic mites from Afrotropical Myriapoda (Acari, Astigmata). – Bull. Inst. R. Sci. Nat. de Belgique, Entomologie, 57: 161–172. The Mite Family Lophonotacaridae Fain Lophonotacaridae Fain, 1987: 163 One genus and species of African mites associated with Diplopoda. Family Lophonotacaridae Fain Lophonotacaridae Fain, 1987: 163 Type genus: Lophonotacarus Fain, 1987 Acarorum Catalogus IX 245 Genus Lophonotacarus Fain Lophonotacarus Fain, 1987: 164 Type species: Lophonotacarus minutus Fain, 1987 Lophonotacarus minutus Fain Lophonotacarus minutus Fain, 1987: 164 Type locality: Transkei, District Lusikisiki, Mouth of Riv. Mzimhlava Distribution: South Africa Host: millipede (Diplopoda: Glomeridae) Index of Lophonotacaridae (Names in bold are accepted as valid) Lophonotacaridae 244 Lophonotacarus 245 minutus, Lophonotacarus 245 Distribution of Lophonotacaridae AFRICA South Africa (Fain, 1987) – Lophonotacarus minutus Hosts of Lophonotacaridae Myriapoda Diplopoda Glomerida Fam. Glomeridae Gen. sp. indet. - Chetochelacarus mamillatus Bibliography of Lophonotacaridae Fain A. 1987. Notes on mites associated with Myriapoda 1. Three new astigmatic mites from Afrotropical Myriapoda (Acari, Astigmata). – Bull. Inst. R. Sci. Nat. de Belgique, Entomologie, 57: 161–172. 246 Acarorum Catalogus IX The Mite Family Heterocoptidae Fain Heterocoptidae Fain, 1961: 91 Mites associated with chrysomelid beetles (Cassidinae and Hispinae), other Coleoptera (Lycidae, Erotylidae, Passandridae) and Hymenoptera (Apidae, Sphecidae), mostly in the Afritropical and Indomalayan Regions. The first genus and species (Heterocoptes tarsii Fain, 1967) has been found on Tarsius (Mammalia: Primates)(chance host ?). Haitlinger described since 1990 10 of the 12 genera, known so far, and most of the species from Africa, Madagascar, South Asia, Peru, Australia, New Guinea and other tropical areas. Composition of Heterocoptidae Fam. Heterocoptidae Fain, 1967–13 gen., 39 sp. Abboticoptes Haitlinger, 1996–1 sp. Amarucoptes Haitlinger, 2001–1 sp. Bahrucoptes Haitlinger, 1999 – 1 sp. Cassiocoptes Haitlinger, 1996–1 sp. Conchycoptes Haitlinger, 2002–1 sp. Erotylocoptes Fain, 1987–7 sp. Gascaricoptes Haitlinger, 2002–1 sp. Heterocoptes Fain, 1967–15 sp. Honiarrea Haitlinger, 1990–1 sp. Kamerucoptes Haitlinger, 2002–5 sp. Nolaecoptes Haitlinger, 1996–2 sp. Tannuicoptes Haitlinger, 1999–2 sp. Uvallicoptes Haitlinger, 1999–1 sp. Catalogue of Heterocoptidae Family Heterocoptidae Fain Heterocoptidae Fain, 1967: 91 Type genus: Heterocoptes Fain, 1967 Genus Abboticoptes Haitlinger Abboticoptes Haitlinger, 1996: 988 Type species: Abboticoptes eddae Haitlinger, 1996 Acarorum Catalogus IX 247 Abboticoptes eddae Haitlinger Abboticoptes eddae Haitlinger, 1996: 988 Type locality: Philippines Distribution: Philippines Host: Aspidomorpha quadrilobata (Coleoptera: Chrysomelidae, Cassidinae) Genus Amarucoptes Haitlinger Amarucoptes Haitlinger, 2001: 112 Type species: Amarucoptes eddae Haitlinger, 2001 Amarucoptes busannae Haitlinger Amarucoptes busannae Haitlinger, 2001: 113 Type locality: 30 km east from Puerto Maldonado, at river Madre de Dios Distribution: Peru Host: Coleoptera: Passalidae indet. Genus Bahrucoptes Haitlinger Bahrucoptes Haitlinger, 1999: 60 Type species: Bahrucoptes amaliae Haitlinger, 1999 Bahrucoptes amaliae Haitlinger Bahrucoptes amaliae Haitlinger, 1999: 60 Type locality: Kota Bahru Distribution: Malaysia Host: Aspidomorpha elevata (Coleoptera: Chrysomelidae, Cassidinae) Genus Cassiocoptes Haitlinger Cassiocoptes Haitlinger, 1996: 986 Type species: Cassiocoptes mikki Haitlinger, 1996 Cassiocoptes mikki Haitlinger Cassiocoptes mikki Haitlinger, 1996: 986 Type locality: Soekaranda (Sumatra) Distribution: Indonesia (Sumatra) Host: Lasiochila fallax (Coleoptera: Chrysomelidae, Hispinae) Genus Conchycoptes Haitlinger Conchycoptes Haitlinger, 2002: 25 Type species: Conchycoptes tangaicus Haitlinger, 2002 Conchycoptes tangaicus Haitlinger Conchycoptes tangaicus Haitlinger, 2002: 25 248 Acarorum Catalogus IX Type locality: Tanga Distribution: Tanzania Host: Conchyloctenia hybrida (Coleoptera: Chrysomelidae, Hispinae) Genus Erotylocoptes Fain Erotylocoptes Fain, 1987: 244 Type species: Erotylocoptes linodesmus Fain, 1987 Erotylocoptes anthophora Fain Erotylocoptes anthophora Fain, 1987: 251 Type locality: Mobwasa Distribution: DR Congo [Zaire] Host: Anthophora acraensis (Hymenoptera, Apidae) Erotylocoptes helenae Haitlinger Erotylocoptes helenae Haitlinger, 1996: 980 Type locality: ?Radburg (RSA) Distribution: Mozambique, South Africa Hosts: Laccoptera rugosicollis (Coleoptera: Chrysomelidae, Cassidinae) Erotylocoptes linodesmus Fain Erotylocoptes linodesmus Fain, 1987: 244 Type locality: Mediakoko Distribution: DR Congo [Zaire] Host: Linodesmus coccus apicalis (Coleoptera, Erotylidae) Erotylocoptes luluensis Fain Erotylocoptes luluensis Fain, 1987: 252 Type locality: Lulua-Kapanga Distribution: DR Congo [Zaire] Host: Chlorion (Isodontia) simoni (Hymenoptera, Sphecidae) Erotylocoptes taorettae Haitlinger Erotylocoptes taorettae Haitlinger, 1996: 981 Type locality: ?Pugu Distribution: Tanzania Host: Laccoptera aurosa (Coleoptera: Chrysomelidae, Cassidinae) Erotylocoptes verenae Haitlinger Erotylocoptes verenae Haitlinger, 1996: 982 Type locality: Congo (Brazaville) Distribution: Congo (Brazaville) Acarorum Catalogus IX 249 Host: Laccoptera caduca (Coleoptera: Chrysomelidae, Cassidinae) Genus Gascaricoptes Haitlinger Gascaricoptes Haitlinger, 2002: 22 Type species: Gascaricoptes analevelonaicus Haitlinger, 2002 Gascaricoptes analevelonaicus Haitlinger Gascaricoptes analevelonaicus Haitlinger, 2002: 22 Type locality: Analavelona Distribution: Madagascar Host: Cassida fuscomacula (Coleoptera: Chrysomelidae, Cassidinae) Genus Heterocoptes Fain Heterocoptes Fain, 1967: 94 Type species: Heterocoptes tarsii Fain, 1967 Heterocoptes barbarae Haitlinger Heterocoptes barbarae Haitlinger, 2009: 334 Type locality: New Britain, Gazelle Peninsula Distribution: Papua New Guinea (New Britain) Host: Aspidomorpha aurata (Coleoptera: Chrysomelidae, Cassidinae) Heterocoptes floresiensis Haitlinger Heterocoptes floresiensis Haitlinger, 2009: 62 Type locality: Flores Distribution: Indonesia (Flores) Host: Aspidimorpha indica (Coleoptera: Chrysomelidae, Cassidinae) Heterocoptes hannae Haitlinger Heterocoptes hannae Haitlinger, 1998: 5 Type locality: Shanghai Distribution: China Host: Basipronota bisignata (Coleoptera: Chrysomelidae, Cassidinae) Heterocoptes ingeri Haitlinger Heterocoptes ingeri Haitlinger, 1999: 67 Type locality: South Luzon Distribution: Philippines Host: Laccoptera tredecimguttata (Coleoptera: Chrysomelidae, Cassidinae) Heterocoptes kamillae Haitlinger Heterocoptes kamillae Haitlinger, 1998: 8 250 Acarorum Catalogus IX Type locality: ? Solim-Ho, Yunnan Distribution: China (Yunnan) Host: Aspidimorpha miliaris (Coleoptera: Chrysomelidae, Cassidinae) Heterocoptes lombokensis Haitlinger Heterocoptes lombokensis Haitlinger, 2009: 326 Type locality: Lombok, Senggigi Distribution: Indonesia (Lombok, Bali, Java) Hosts: Aspidimorpha sanctaecrucis, A. miliaris (Coleoptera: Chrysomelidae, Cassidinae) Heterocoptes lottae Haitlinger Heterocoptes lottae Haitlinger, 1996: 985; 1999: 72 Type locality: North Vietnam Distribution: Malaysia, Vietnam Host: Laccoptera hospita, Aspidomorpha elevata (Coleoptera: Chrysomelidae, Cassidinae) Heterocoptes marcellae Haitlinger Heterocoptes marcellae Haitlinger, 2009: 336 Type locality: South Sulawesi Distribution: Indonesia (Sulawesi) Host: Aspidimorpha amabilis (Coleoptera: Chrysomelidae, Cassidinae) Heterocoptes nolae Haitlinger Heterocoptes nolae Haitlinger, 1996: 983 Type locality: Andaman Is. Distribution: India (Andaman Is.) Host: Aspidomorpha inquinata (Coleoptera: Chrysomelidae, Cassidinae) Heterocoptes rukaensis Haitlinger Heterocoptes rukaensis Haitlinger, 1999: 70 Type locality: Ruka Distribution: Papua New Guinea Host: Aspidimorpha punctum (Coleoptera: Chrysomelidae, Cassidinae) Heterocoptes ruperti Haitlinger Heterocoptes ruperti Haitlinger, 2009: 64 Type locality: ? Liangages, Sumatra Distribution: Indonesia (Sumatra) Host: Platypria echinogale (Coleoptera: Chrysomelidae, Hispinae) Heterocoptes samueli Haitlinger Heterocoptes samueli Haitlinger, 2009: 332 Acarorum Catalogus IX 251 Type locality: “New Territory”, Darwin Distribution: Australia Host: Aspidimorpha quadriradiata (Coleoptera: Chrysomelidae, Cassidinae) Heterocoptes simonettae Haitlinger Heterocoptes simonettae Haitlinger, 1999: 70 Type locality: Papua New Guinea Host: Aspidimorpha punctata (Coleoptera: Chrysomelidae, Cassidinae) Heterocoptes sumatrensis Haitlinger Heterocoptes sumatrensis Haitlinger, 2009: 329 Type locality: Sumatra, Soekaranda Distribution: Indonesia (Sumatra) Host: Basipronota decemmaculata (Coleoptera: Chrysomelidae, Cassidinae) Heterocoptes tarsii Fain Heterocoptes tarsii Fain, 1967: 94; Haitlinger, 1996: 985; 2009: 69 Type locality: Sarawak (Malaysia) Distribution: India (Assam), Indonesia (Bali, Sanur, Lombok, Java, Sumatra, Ceram), Malaysia (Sarawak, Perak), Philippines Hosts: Aspidomorpha elevata, A. sanctaecrucis (Coleoptera: Chrysomelidae, Cassidinae); described from Tarsius sp. (Primates: Tarsiidae) Genus Honiarrea Haitlinger Honiarrea Haitlinger, 1990: 343 Type species: Honiarrea lagossa Haitlinger, 1990 Honiarrea lagossa Haitlinger Honiarrea lagossa Haitlinger, 1990: 343 Type locality: Hainan Is. Distribution: China (Hainan Is.) Host: Passandra heros (Coleoptera: Passandridae) Genus Kamerucoptes Haitlinger Kamerucoptes Haitlinger, 2002: 8 Type species: Kamerucoptes doubeicus Haitlinger, 2002 Kamerucoptes doubeicus Haitlinger Kamerucoptes doubeicus Haitlinger, 2002: 8 Type locality: Doube Distribution: Cameroon 252 Acarorum Catalogus IX Host: Aspidimorpha obovata (Coleoptera: Chrysomelidae, Cassidinae) Kamerucoptes lycius Haitlinger Kamerucoptes lycius Haitlinger, 2002: 13 Type locality: D.R. Congo Distribution: D.R. Congo Host: Lycidae gen. sp. (Coleoptera) Kamerucoptes ruvenzoricus Haitlinger Kamerucoptes ruvenzoricus Haitlinger, 2002: 13 Type locality: Ruvenzori, Mutwanga Distribution: Uganda Host: Aspidimorpha dissentonea (Coleoptera: Chrysomelidae, Cassidinae) Kamerucoptes sibuticus Haitlinger Kamerucoptes sibuticus Haitlinger, 2002: 16 Type locality: N’Chari, Fort Sibut Distribution: Tshad Host: Laccoptera (Patrisma) murrayi (Coleoptera: Chrysomelidae, Cassidinae) Kamerucoptes tsaratananaicus Haitlinger Kamerucoptes tsaratananaicus Haitlinger, 2002: 19 Type locality: Mts Tsaratanana, 2000 m Distribution: Madagascar Host: Cassida pretiosa (Coleoptera: Chrysomelidae, Cassidinae) Genus Nolaecoptes Haitlinger Nolaecoptes Haitlinger, 1996: 987 Type species: Nolaecoptes vonettae Haitlinger, 1996 Nolaecoptes tanahlotions Haitlinger Nolaecoptes tanahlotions Haitlinger, 2009: 324 Type locality: Bali, Tanahlot Distribution: Indonesia (Bali) Nolaecoptes vonettae Haitlinger Nolaecoptes vonettae Haitlinger, 1996: 987 Type locality: Sikkim Distribution: India (Sikkim) Host: Aspidomorpha sanctaecrucis (Coleoptera: Chrysomelidae, Cassidinae) Acarorum Catalogus IX 253 Genus Tannuicoptes Haitlinger Tannuicoptes Haitlinger, 1999: 62 Type species: Tannuicoptes ulfi Haitlinger, 1999 Tannuicoptes jeroni Haitlinger Tannuicoptes jeroni Haitlinger, 1999: 65 Type locality: Indonesia Distribution: Indonesia Host: Corynodes compressicornis (Coleoptera: Chrysomelidae, Eumolpinae) Tannuicoptes ulfi Haitlinger Tannuicoptes ulfi Haitlinger, 1999: 62 Type locality: Sikkim Distribution: India (Sikkim) Host: Aspidomorpha dorsata (Coleoptera: Chrysomelidae, Cassidinae) Genus Uvallicoptes Haitlinger Uvallicoptes Haitlinger, 1999: 62 Type species: Uvallicoptes peeteri Haitlinger, 1999 Uvallicoptes peeteri Haitlinger Uvallicoptes peeteri Haitlinger, 1999: 62 Type locality: ? Small Gela, Florida Is., Solomon Islands Distribution: Solomon Islands, Indonesia [Sumatra] Host: Aspidomorpha adhaerens (Coleoptera: Chrysomelidae, Cassidinae) Index of Heterocoptidae (Names in bold are accepted as valid) Abboticoptes 246 amaliae, Bahrucoptes 247 Amarucoptes 247 analevelonaicus, Gascaricoptes 249 anthophora, Erotylocoptes 248 Bahrucoptes 247 barbarae, Heterocoptes 249 busannae, Amarucoptes 247 Cassiocoptes 247 Conchycoptes 247 doubeicus, Kamerucoptes 251 eddae, Abboticoptes 247 Erotylocoptes 248 floresiensis, Heterocoptes 249 Gascaricoptes 249 hannae, Heterocoptes 249 helenae, Erotylocoptes 248 Heterocoptes 249 Heterocoptidae 246 Honiarrea 251 254 Acarorum Catalogus IX ingeri, Heterocoptes 249 jeroni, Tannuicoptes 253 Kamerucoptes 251 kamillae, Heterocoptes 249 lagossa, Honiarrea 251 linodesmus, Erotylocoptes 248 lombokensis, Heterocoptes 250 lottae, Heterocoptes 250 luluensis, Erotylocoptes 248 lycius, Kamerucoptes 252 marcellae, Heterocoptes 250 mikki, Cassiocoptes 247 nolae, Heterocoptes 250 Nolaecoptes 252 peeteri, Uvallicoptes 252 rukaensis, Heterocoptes 250 ruperti, Heterocoptes 250 ruvenzoricus, Kamerucoptes 252 samueli, Heterocoptes 250 sibuticus, Kamerucoptes 252 simonettae, Heterocoptes 251 sumatrensis, Heterocoptes 251 tanahlotions, Nolaecoptes 252 tangaicus, Conchycoptes 247 Tannuicoptes 253 taorettae, Erotylocoptes 248 tarsii, Heterocoptes 251 tsaratananaicus, Kamerucoptes 252 ulfi, Tannuicoptes 253 Uvallicoptes 253 verenae, Erotylocoptes 248 vonettae, Nolaecoptes 252 Distribution of Heterocoptidae AFRICA Cameroon (Haitlinger, 2002) – Kamerucoptes doubeicus Congo (Brazzaville) (Haitlinger, 1996) - Erotylocoptes verenae Congo DR (Fain, 1987; Haitlinger, 2002) – Erotylocoptes anthophora, E. linodesmus, E. luluensis, Kamerucoptes lycius Madagascar (Haitlinger, 2002) - Gascaricoptes analevelonaicus, Kamerucoptes tsaratananaicus Mozambique (Haitlinger, 1996) - Erotylocoptes helenae South Africa (Haitlinger, 1996) – Erotylocoptes helenae Tanzania (Haitlinger, 1996, 2002) – Conchycoptes tangaicus, Erotylocoptes taorettae Tshad (Haitlinger, 2002) - Kamerucoptes sibuticus Uganda (Haitlinger, 2002) - Kamerucoptes ruvenzoricus ASIA China (Haitlinger, 1998; Fan, Chen & Wang, 2010) – Heterocoptes hannae, H. kamillae Hainan (Haitlinger, 1990) - Honiarrea lagossa India (Fain, 1967; Haitlinger, 1996 , 1999) – Heterocoptes tarsii, Nolaecoptes vonettae, Tannuicoptes ulfi Andaman Is. (Haitlinger, 1996) – Heterocoptes nolae Indonesia (Haitlinger, 1996, 1999; 2009a) – Cassiocoptes mikki [Sumatra], Heterocoptes floresiensis [Flores], H. lombokensis [Lombok, Bali, Java], H. marcellae [Sulawesi], Acarorum Catalogus IX 255 H. ruperti [Sumatra], H. sumatrensis [Sumatra], H. tarsii (Bali, Sanur, Lombok, Sumatra, Java, Ceram), Nolaecoptes tanahloticus [Bali], Uvallicoptes peeteri [Sumatra], Tannuicoptes jeroni Malaysia (Fain, 1967; Haitlinger, 1999) – Bahrucoptes amaliae, Heterocoptes tarsii Sarawak and Sabah (Fain, 1967) – Heterocoptes tarsii Philippines (Haitlinger, 1996, 1999) - Abboticoptes eddae, Heterocoptes ingeri, H. tarsi, Vietnam (Haitlinger, 1996) – Heterocoptes lottae NORTH AMERICA Canada (Beaulieu et al., 2019) – one unidentified sp. SOUTH and CENTRAL AMERICA Peru (Haitlinger, 2001) – Amarucoptes busannae OCEANIA Australia (Haitlinger, 2009a) – Heterocoptes samueli Papua New Guinea (Haitlinger, 1999, 2009) - Heterocoptes simonettae, H. rukaensis New Britain (Haitlinger, 2009a) – Heterocoptes barbarae New Ireland - Uvallicoptes peeteri Solomon Islands (Haitlinger, 1999) - Uvallicoptes peeteri Hosts of Heterocoptidae Insecta Coleoptera Fam. Chrysomelidae Subfam. Hispinae Conchyloctenia hybrida - Conchycoptes tangaicus Lasiochila sp. – Cassiocoptes mikki Platypria echinogale - Heterocoptes ruperti Subfam. Cassidinae Aspidimorpha adhaerens – Uvallicoptes peeteri Aspidimorpha amabilis – Heterocoptes marcellae Aspidimorpha dissentonea - Kamerucoptes ruvenzoricus Aspidimorpha dorsata – Tannuicoptes ulfi Aspidimorpha elevata – Bahrucoptes amaliae, Heterocoptes tarsii Aspidimorpha fuscopunctata - Uvallicoptes peeteri Aspidimorpha inquinata – Heterocoptes nolae Aspidimorpha miliaris – Heterocoptes kamillae, H. lombokensis Aspidimorpha obovata - Kamerucoptes doubeicus Aspidimorpha punctum – Heterocoptes simonettae 256 Acarorum Catalogus IX Aspidimorpha quadrilobata – Abboticoptes edda Aspidimorpha quadriradiata – Heterocoptes samueli Aspidimorpha sanctaecrucis – Heterocoptes lombokensis, H. tarsii, Nolaecoptes tanahloticus, N. vonettae Aspidimorpha socia – Heterocoptes barbarae Basipronota bisignata - Heterocoptes hannae Basipronota decemmaculata - Heterocoptes sumatrensis Cassida fuscomacula - Gascaricoptes analevelonaicus Cassida pretiosa - Kamerucoptes tsaratananaicus Laccoptera aurosa - Erotylocoptes taorettae Laccoptera caduca – Erotylocoptes verenae Laccoptera hospita – Heterocoptes lottae Laccoptera (Patrisma) murrayi - Kamerucoptes sibuticus Laccoptera rugosicollis - Erotylocoptes helenae Laccoptera tredecimguttata – Heterocoptes ingeri Subfam. Eumolpinae Corynodes compressicornis - Tannuicoptes jeroni Fam. Erotylidae Linodesmus coccus apicalis – Erotylocoptes linodesmus Fam. Lycidae gen. sp. - Kamerucoptes lycius Fam. Passandridae Passandra heros – Honiarrea lagossa Fam. Passalidae gen.sp. - Amarucoptes busannae Hymenoptera Fam. Apidae Anthophora acraensis - Erotylocoptes anthophora Fam. Sphecidae Chlorion (Isodontia) simoni – Erotylocoptes luluensis Mammalia Primates Fam. Tarsiidae Tarsius sp. - Heterocoptes tarsii Bibliography of Heterocoptidae Beaulieu et al. 2019. Acari of Canada. – ZooKeys, 819: 77–168. Fain A. 1967. Un Acarien rémarquable recolté sur un Tarsier (Heterocoptidae f. n.: Sarcoptiformes). – Zool. Anzeiger, 178 (1/2): 90–94. Fain A. 1987. Nouveau taxa dans la famille Heterocoptidae Fain, 1967 (Acari, Astigmata). – Bull. Annls Soc. r. belge Ent., 123: 243–253. Acarorum Catalogus IX 257 Fan Qing-Hai, Yan Chen, Zi-Qing Wang. 2010. Acaridia (Acari: Astigmatina) of China: A review of research progress. – p.225–259. In: Zhang Z.-Q., Hong X.-Y. & Fan Q.-H. (eds) Xin Jie-Lin Centenary: Progress in Chinese Acarology. Zoosystema 4, 1–345. Haitlinger R. 1990. Honiarrea gen. nov., a new genus of the family Heterocoptidae (Acari, Astigmata). – Annales Zoologici, Warszawa, 43(16): 343–345. Haitlinger R. 1996. New heterocoptid mites (Acari, Astigmata, Heterocoptidae) associated with Cassidinae and Hispinae (Coleoptera, Chrysomelidae) from Africa and Asia. – Linzer Biol. Beitr., 28: 979–998. Haitlinger R. 1998. Four new species of canestriniid and heterocoptid mites from China (Acari, Astigmata, Canestriniidae, Heterocoptidae). – Spixiana, 21: 1–10. Haitlinger R. 1999. Six new heterocoptid mites (Acari, Astigmata, Heterocoptidae) associated with Cassidinae and Eumolpinae (Coleoptera, Chrysomelidae) from Indo-Australian Region. – Misc. Zool., 22: 59–73. Haitlinger R. 2001. Amarucoptes, a new genus of the family Heterocoptidae (Acari, Astigmata). – Syst. Appl. Acarol., 6: 111–114. Haitlinger R. 2002. Seven new Heterocoptid mites (Acari: Astigmata: Heterocoptidae) from Africa. – Zeszyty naukowe Akademii Rol. we Wroclawiu, Zootechnika XLIX, 447: 7–29. Haitlinger R. 2009a. New species of mites (Acari: Astigmata: Heterocoptidae) from IndoAustralian region. – Zesz. Nauk. UP Wroc., Biol. Hod. Zwierz., 58(572): 61–72. Haitlinger R. 2009b. Nine new species and new records of mites (Acari: Astigmata: Heterocoptidae) from tropical Africa. – Revista Ibérica de Aracnología, 17: 97–115. Haitlinger R. 2009c. New species of mites (Acari: Astigmata: Heterocoptidae) from the Australian and Oriental regions, with a description of the female of Uvalicoptes peeteri Haitlinger, 1999. – Biologia, 64(2): 324–339. OConnor B.M. 1979. A review of the family Heterocoptidae. – Proc. V Int. Congr. Acar. (August 6–12, 1978, East Lansing). Recent Advances of Acarology II: 429–433. The Mite Superfamily Hemisarcoptoidea According to O’Connor (2009), the superfamily contains the families Chaetodactylidae, Hyadesiidae, Carpoglyphidae, Algophagidae, Hemisarcoptidae and Winterschmidtiidae, or in total 51 genera and ca. 345 species. Some of them (Chaetodactylidae) live on Hymenoptera, some (Hyadesiidae, Algophagidae) prefer the sea shore and the saline habitats. Hemisarcoptidae are found to live as parasites on Insects (Coccodea, Coleoptera – Chrysomelidae, Coccinellidae ) and some of them are used for the pest control. Carpoglyphidae and Winterschmidtiidae are inhabitants of rotten materials, sometimes found also on insects. 258 Acarorum Catalogus IX The Mite Family Chaetodactylidae Glyciphagidae (Chaetodactylinae) Zachvatkin, 1941: 347 Chaetodactylidae Zachvatkin: Baker, 1962: 229 (part.); Klimov & OConnor, 2008: 98 Obligate associates of long-tongued bees (OConnor & Klimov, 2009). From the five known genera in the family the first (Chaetodactylus Rondani) has been described as early as in 1866, but remained the only one until Oudemans (1905) described (based on Pediculus cerambycinus Scopoli, 1763) the large genus Sennertia, revised much later by Fain (from 1966 to 2001). The thorough study of the family starts with the results of B. OConnor and P. Klimov, who published generic revision of the group (OConnor, 1993) and very detailed monograph (Klimov & OConnor, 2008), containing the history of the research on Chaetodactylidae until this time. As early as in 1903 Tschitscherin separated these mites in a group Chaetodactylini. Zachvatkin (1941) considered Chaetodactylinae as subfamily of Glycyphagidae and Baker (1962) raised them in a family rank. Meanwhile some other authors contributed to the taxonomy and/or the biology of Chaetodactylidae (Womersley, 1941, Vitzthum, 1941, Elbadry, 1971, Delfinado & Baker, 1976, Sherbef & El-Duweini, 1980, Baker & Delfinado-Baker, 1983, Haitlinger, 1999, 2000, Kurosa, 1987, 2003, Ramaraju & Mohanasundaram, 2001, and others). Composition of Chaetodactylidae Fam. Chaetodactylidae Zachvatkin, 1941–5 genera, 109 sp. Achaetodactylus Fain, 1981–3 sp. Centriacarus Klimov et OConnor, 2007–2 sp. Chaetodactylus Rondani, 1866–26 sp. Roubikia OConnor, 1993–4 sp. Sennertia Oudemans, 1905–72 sp. Catalogue of Chaetodactylidae Fam. Chaetodactylidae Zachvatkin Glycyphagidae (Chaetodactylinae) Zachvatkin, 1941: 347 Chaetodactylidae Zachvatkin; Baker, 1962: 229 (par.); Klimov & OConnor, 2008 : 98 Type genus: Chaetodactylus Rondani, 1866 Genus Achaetodactylus Fain Achaetodactylus Fain, 1981a: 2 (as subgenus) Acarorum Catalogus IX 259 Achaetodactylus Fain: Klimov et al., 2007a: 1370 (as genus) Chaetodactylus (Ochaetodactylus) Fain, 1981b: 2. Type species Chaetodactylus decellei Fain, 1974 Type species: Chaetodactylus leleupi Fain, 1974 Achaetodactylus ceratinae (Fain) Chaetodactylus ceratinae Fain, 1974a: 214 Chaetodactylus (Achaetodactylus) ceratinae Fain, 1981a: 4 Achaetodactylus ceratinae (Fain): Klimov et al., 2007a: 1371; Klimov & OConnor, 2008: 215 Type locality: Nairobi Distribution: Kenya, South Africa, Tanzania Hosts: Ceratina apaca, Ceratina (Hirashima) lativentris, C. (H.) nigriceps (Hymenoptera: Apidae, Xylocopinae) Achaetodactylus decellei (Fain) Chaetodactylus decellei Fain, 1974: 215 Chaetodactylus (Ochaetodactylus) decellei: Fain, 1981a: 8 Ochaetodactylus decellei: Klimov et al., 2007a: 1371 Achaetodactylus decellei (Fain): Klimov & OConnor, 2008: 216 Type locality: Ngurdoto Nat. Park, Tanzania Distribution: Cameroon, DR Congo, Tanzania Hosts: Ceratina aereola, C. excavata, C. spilota (Hymenoptera: Apidae, Xylocopinae) Achaetodactylus leleupi (Fain) Chaetodactylus leleupi Fain, 1974: 214 Chaetodactylus (Achaetodactylus) leleupi: Fain, 1981a: 4; Fain & Pauly, 2001: 131 Achaetodactylus leleupi (Fain): Klimov et al., 2007a: 1371; Klimov & OConnor, 2008: 215 Type locality: Karen, Nairobi (Kenya) Distribution: Cameroon, DR Congo, Kenya, Tanzania Hosts: Ceratina (Hirashima) lativentris, Ceratina diloloensis, C. ruwenzorica, C. spilota (Hymenoptera: Apidae, Xylocopinae) Genus Centriacarus Klimov et OConnor Centriacarus Klimov & OConnor, 2007: 816 Type species: Centriacarus turbator Klimov et OConnor, 2007 Centriacarus guahibo Klimov et OConnor Centriacarus guahibo Klimov & OConnor, 2007: 817; Klimov et al., 2007a: 1371; 2008: 216 Type locality: Territorio Amazonas Distribution: Venezuela Host: Centris sp. (Hymenoptera: Apidae, Apinae) 260 Acarorum Catalogus IX Centriacarus turbator Klimov et OConnor Centriacarus turbator Klimov & OConnor, 2007: 816; Klimov et al., 2007a: 1371; Klimov & OConnor, 2008: 216 Type locality: Mato Grosso de Sul, Aquidauana (Brazil) Distribution: Brazil, Colombia, Mexico, Panama Host: Centris (Heterocentris) vittata (Hymenoptera: Apidae, Apinae) Genus Chaetodactylus Rondani Chaetodactylus Rondani, 1866: 19 Type species: Trichodactylus osmiae Dujardin, 1839 Subgenus Chaetodactylus Rondani Chaetodactylus Rondani, 1866: 19 Type species: Trichodactylus osmiae Dujardin, 1839 Chaetodactylus abditus Klimov et OConnor Chaetodactylus abditus Klimov & OConnor, 2004: 175; 2008: 124 Type locality: Arizona, Pima Co., Continental Distribution: Mexico, USA (Arizona) Hosts: Lithurgus (Lithurgopsis) echinocacti, L. (Lithurgopsis) planifrons (Hymenoptera: Megachilidae) Chaetodactylus anthidii (Oudemans) Trichotarsus anthidii Oudemans, 1911a: 165; Vitzthum, 1919: 31 Chaetodactylus anthidii: Oudemans, 1924: 328; Zachvatkin, 1941: 398; Klimov & OConnor, 2008: 110 Chaetodactylus (Chaetodactylus) anthidii: Fain, 1981a: 2; OConnor, 1993a: 62 Type locality: Tunisia Distribution: Tunisia Host: Rhodanthidium sticticum (as Anthidium sticticum)(Hymenoptera: Megachilidae) Chaetodactylus antillarum Klimov et OConnor Chaetodactylus antillarum Klimov & OConnor, 2008: 113 Type locality: Jamaica, Hellshire Hills Distribution: Dominican Republic, Jamaika Host: Lithurgus antilleorum antilleorum (Hymenoptera: Megachilidae) Chaetodactylus azteca Klimov et OConnor Chaetodactylus azteca Klimov & OConnor, 2008: 127 Type locality: Hidalgo: Tepeapulco Distribution: Mexico Acarorum Catalogus IX 261 Host: Osmia (Diceratosmia) azteca (Hymenoptera: Megachilidae) Chaetodactylus birulai Zachvatkin Chaetodactylus birulai Zachvatkin, 1941: 396; Samšinák, 1973: 404; Fain, 1974a: 213; Klimov & OConnor, 2008: 216 Chaetodactylus (Chaetodactylus) birulai Zachvatkin: Fain, 1981a: 3; Kurosa, 1987: 373 Chaetodactylus poetae Samšinák, 1973: 401; Fain, 1981a: 2. Type: Czech Republic: Sobotka; syn. by Klimov & OConnor, 2008: 109 Chaetodactylus (Chaetodactylus) poetae: Kurosa, 1987: 373 Type locality: Kirovskaya Oblast’ (Russia) Distribution: Czech Rep., Holland, Russia Host: Chelostoma florisomne (Hymenoptera: Megachilidae), also found phoretic on cleptoparasites: Sapyga quinquepunctata, Sapyga clavicornis (Hymenoptera: Sapygidae) Chaetodactylus chrysidis Fain et Baugnée Chaetodactylus chrysidis Fain & Baugnée, 1996: 23; Van Asselt, 2000: 225; Klimov & OConnor, 2008: 110 Chaetodactylus chrysidis aurulenticola Fain & Baugnée,1996: 28. Type: Belgium, TreignesRivelottes, ex Osmia aurulenta; syn. by Klimov & OConnor, 2008: 110; Van Asselt, 2000: 225 Type locality: Treignes-Saumières Distribution: Belgium Host: Chrysura trimaculata (Hymenoptera: Chrysididae), Osmia aurulenta and O. bicolor (Hymenoptera: Megachilidae) Chaetodactylus claudus Klimov et OConnor Chaetodactylus claudus Klimov & OConnor, 2008: 133 Type locality: Utah, Cache Co., Green Canyon Distribution: USA (Utah, Texas) Host: Osmia (O.) rubiflorus (Hymenoptera: Megachilidae) Chaetodactylus claviger Oudemans Chaetodactylus claviger Oudemans, 1924: 328; Zachvatkin, 1941: 396; Klimov & OConnor, 2008: 111 Chaetodactylus (Spinodactylus) claviger: Fain, 1981a: 2 (part., excluding Chaetodactylus krombeini) Chaetodactylus osmiae: Abou Senna, 1997: 667 (misidentification, Egypt, from Apis melifera) Type locality: Promontorio del Gargano (“Monte Gargano, Apulië”), Italy Distribution: Egypt, France, Italy 262 Acarorum Catalogus IX Hosts: Osmia tricornis (Hymenoptera: Megachilidae), Apis melifera (Hymenoptera: Apidae) Chaetodactylus dalyi Fain Chaetodactylus dalyi Fain, 1974: 213; Klimov & OConnor, 2007: 826; 2008: 213 Chaetodactylus (Chaetodactylus) dalyi: Fain, 1981a: 6 Type locality: Natal Distribution: Mozambique, South Africa, Zimbabwe Host: Ceratina (Pithitis) turneri (Hymenoptera: Apidae, Xylocopinae) Chaetodactylus dementjevi Zachvatkin Chaetodactylus dementjevi Zachvatkin, 1941: 399; Klimov & OConnor, 2008: 103 Chaetodactylus (Achaetodactylus) dementjevi: Fain, 1981a: 2 Type locality: Uzbekistan: Toshkent (Gora Aktash) Distribution: Kazakhstan, Russia, Uzbekistan Hosts: Megachile bombycina, M. ligniseca (Hymenoptera: Megachilidae), also on Vespula germanica (Vespidae) Chaetodactylus furunculus Klimov et OConnor Chaetodactylus furunculus Klimov & OConnor, 2008: 116 Type locality: California, San Bernardino Co., Cedar Canyon Distribution: USA (California) Host: Lithurgus listrotus (Hymenoptera: Megachilidae) Chaetodactylus gibbosi Klimov et OConnor Chaetodactylus gibbosi Klimov & OConnor, 2004: 178; 2008: 124 Type locality: Florida, Liberty Co. Distribution: USA (Florida) Host: Lithurgus gibbosus (Hymenoptera: Megachilidae) Chaetodactylus hirashimai Kurosa Chaetodactylus (Chaetodactylus) hirashimai Kurosa, 1987: 374 Saproglyphus sp.: Hirashima, 1957: 200 (Host: Osmia excavata) Chaetodactylus sp.: Krombein, 1962: 239; Host: Osmia excavata, Japan Chaetodactylus hirashimai: Qu et al., 2002: 121; Klimov & OConnor, 2008: 110 Type locality: Japan: Honshu, Nara Pref., Nara-shi, Furuichi-chô Distribution: Japan (Honshu, Kyushu) Hosts: Osmia (Osmia) excavata, O. (O.) cornifrons, O. (O.) pedicornis, Osmia (Helicosmia) imaii (Hymenoptera: Megachilidae) Acarorum Catalogus IX 263 Chaetodactylus hopliti Klimov et OConnor Chaetodactylus hopliti Klimov & OConnor, 2008: 137 Type locality: New York, Onondaga Co., Syracuse Distribution: Canada, USA Hosts: Hoplitis (Alcidamea) producta (type host); Hoplitis (Alcidamea) pilosifrons; Hoplitis (Monumetha) spoliata; Megachile (Megachile) relativa; Osmia (Melanosmia) brevis; Osmia (Centrosmia) bucephala; Osmia (Chenosmia) inermis (Megachilidae); Isodontia mexicana (Hymenoptera: Sphecidae) Chaetodactylus kouboy Klimov et OConnor Chaetodactylus kouboy Klimov & OConnor, 2008: 121 Type locality: New Mexico, Colfax Co., Cimarron Canyon Distribution: USA (New Mexico) Host: Lithurgus (Lithurgopsis) apicalis (Hymenoptera: Megachilidae) Chaetodactylus krombeini Baker Chaetodactylus krombeini Baker, 1962: 229; Klimov & OConnor, 2008: 141; McKinney & Park, 2012: Chaetodactylus (Spinodactylus) krombeini Baker: Fain, 1981 : 2; OConnor, 1993a: 362 Type locality: Maryland Distribution: USA Hosts: Osmia cornifrons, O. rubifloris, O. r. biedermanni, O. bucephala, O. chlybaea, O. lignaria propinqua, O. montana, O. californica, O. grindeliae (Hymenoptera: Magachilidae) Chaetodactylus lassulus Klimov et OConnor Chaetodactylus lassulus Klimov & OConnor, 2007: 824; 2008: 111 Type locality: Coquimbo Distribution: Chile Hosts: Trichothurgus dubius, T. herbsti (Hymenoptera: Magachilidae) Chaetodactylus lithurgi Klimov et OConnor Chaetodactylus lithurgi Klimov & OConnor, 2004: 173; 2008: 121 Type locality: New Mexico, Colfax Co., Cimarron Canyon Distribution: USA (New Mexico, Idaho, Texas, Arizona) Hosts: Lithurgus apicalis, L. gibbosus, L. littoralis (Hymenoptera: Megachilidae) Chaetodactylus ludwigi (Trouessart) Trichotarsus Ludwigi Trouessart, 1904a: 234; 1904b: 365; Ludwig, 1904: 216; Vitzthum, 1912d: 292; 1919; Knülle, 1959: 385 264 Acarorum Catalogus IX Chaetodactylus ludwigi (Trouessart): Zachvatkin, 1941: 42; Baker, 1962a: 229; 1987: 65; OConnor, 1993a: 353; Pauly & Munzinger, 2003: 160; Klimov & OConnor, 2007: 826; Klimov et al., 2007a: 1371; Klimov & OConnor, 2008: 111 Chaetodactylus (Chaetodactylus) ludwigi: Fain & Pauly, 2001: 128 Type locality: Carolines Islands: Ponapé Distribution: Federated States of Micronesia: Pohnpei Is. (“Carolines Islands: Ponapé”); New Caledonia; French Polynesia: Moorea Is. Nr. Taiti; India, Indonesia (Java), Madagascar Hosts: Lithurgus (Lithurgus) atratus (as Megachile lonalap), nest; Lithurgus scabrosus (Hymenoptera: Megachilidae) Chaetodactylus melitomae Klimov et OConnor Chaetodactylus melitomae Klimov & OConnor, 2007: 821; 2008: 113 Type locality: Mexico: Tabasco, Cardenas Distribution: Honduras, Mexico Host: Melitoma sp., Melitoma segmentaria (Hymenoptera: Apidae, Apinae) Chaetodactylus micheneri Klimov et OConnor Chaetodactylus micheneri Klimov & OConnor, 2008: 127 Type locality: California, Fresno Co., Mt. Crocker, 11800 ft. Distribution: USA (California) Host: Osmia subaustralis (Hymenoptera: Megachilidae) Chaetodactylus nipponicus Kurosa Chaetodactylus sp.: Yamada et al., 1971: 32 Chaetodactylus (Chaetodactylus) nipponicus Kurosa, 1987: 377 Chaetodactylus nipponicus: Van Asselt, 2000: 221; Bosh & Kemp, 2001: 62; Qu et al., 2002: 121; Qu et al., 2003: 55; Klimov & OConnor, 2008: 110 Type locality: Tsugaru-gun (Tsushima Is.) Distribution: Japan (Tsushima Is., Honshu) Hosts: Osmia (Osmia) excavata, O. (O.) cornifrons, O. (O.) taurus, O. (O.) pedicornis (Hymenoptera: Megachilidae) Chaetodactylus osmiae (Dufour) Trichodactylus osmiae Dufour, 1839: 276; Gervais, 1844: 266 (part.); Murray, 1877: 252 (part.); Mégnin, 1880: 147; Canestrini, 1888a: 395; 1888b: 23 Chaetodactylus osmiae: Rondani, 1866: 183; Zachvatkin, 1941: 394 (part.); Turk, 1953: 82; Türk & Türk, 1957: 207; Lith, 1957: 197; Baker, 1962a: 229; 1987: 65; Krombein, 1962: 238; Fain, 1966: 249; Elbadry, 1971: 88; De Jong et al., 1982: 245 (Apis); Bosch, 1992: 77; Chmielewski, 1993: 133; Fain & Baugnée, 1996: 23; Van Asselt, 2000: 221; Acarorum Catalogus IX 265 Fain & Pauly, 2001: 127; Qu et al., 2003: 59; Krunic´ et al., 2005: 143; Klimov et al., 2007a: 1371; Klimov & OConnor, 2008: 110 Trichotarsus osmiae: Berlese, 1897: 105; Canestrini & Kramer, 1899: 149; Giard, 1900: 377; Oudemans, 1900: 117; 1901: 81; 1903a: 147; 1905a: 22; Oudemans, 1905b: 80; 1911a: 165; Michael, 1903: 17; Ludwig, 1904: 216; Trouessart, 1904a: 235; 1904b: 365; Vitzthum, 1912d: 291; 1919: 38; Popovici-Baznosanu, 1913: 32; Knülle, 1959: 385 Chaetodactylus mahunkai Samsinák, 1973: 404. Type: Hungary; synonymized by Fain, 1981b. Chaetodactylus (Chaetodactylus) osmiae: Fain, 1981b: 4; Kurosa, 1987: 373; OConnor, 1993a:353 Misidentifications (acc. to Klimov & OConnor, 2008): Donnadieu, 1868 (Chaetodactylus claviger), Banks, 1902: 176 (Chaetodactylus krombeini or Ch. rozeni), Abou Senna, 1997 (Chaetodactylus claviger), Fain et al., 1992 (Chaetodactylus chrysidis aurulenticola) Type locality: France, Département des Landes (Gervais) Distribution: Belgium, Croatia, England, France, Georgia, Germany, Holland, Hungary, Italy, Luxemburg, Poland, Romania, Russia Hosts: Osmia bicornis, O. cornuta, O. fronticornis, O. kohlii, O. niveata, O. rufa, O. rufiventris, O. tricornis (Hymenoptera: Megachilidae); Andrena spp., Megachile sp. (Berlese); Anthophora sp. Chaetodactylus reaumuri (Oudemans) Trichotarsus reaumuri Oudemans, 1905b: 81; 1905a: 22; 1911a: 165; Vitzthum, 1919: 31 Chaetodactylus reaumuri (Oudemans): Oudemans, 1924: 328; Vitzthum, 1929: 77; Zachvatkin, 1941: 396; Türk & Türk, 1957: 210; Fain & Baugnée, 1996: 28; Klimov & OConnor, 2008: 110 Chaetodactylus (Chaetodactylus) reaumuri: Fain, 1981b: 2; Kurosa, 1987: 373 (reamuri); OConnor, 1993a: 362 (reaumerii) Type locality: Kolin (Czech Rep.) Distribution: Czech Rep., Georgia, Germany, Greece (Corfu), ? Poland (“Odran, Oostenrijksch Silezië”); Crimea. Hosts: Osmia rufiventris, O. brevicornis, O. leucogastra (as Osmia panzeri)(Hymenoptera: Megachilidae) Chaetodactylus rozeni Klimov et OConnor Chaetodactylus rozeni Klimov & OConnor, 2008: 132 Type locality: North Carolina, Swain Co., Bryson City Distribution: USA (North Carolina, Texas, Tennessee) Hosts: Osmia georgica (Hymenoptera: Megachilidae); Chrysis coerulans (Hymenoptera: Chrysididae) 266 Acarorum Catalogus IX Chaetodactylus zachvatkini Klimov et OConnor Chaetodactylus zachvatkini Klimov & OConnor, 2008: 110 (nom. nov. pro Chaetodactylus osmiae sensu Zachvatkin, 1941(part., non Dufour, 1839) Type locality: Italy Distribution: Italy, Spain, Tunisia Hosts: Osmia cornuta, O. latreillei, O. niveata, O. tricornis (Hymenoptera: Megachilidae) Genus Roubikia OConnor Roubikia OConnor, 1993: 347 Type species: Chaetodactylus panamensis Baker, Roubik et Delfinado-Baker, 1987 Roubikia imberba Klimov et OConnor Roubikia imberba Klimov & OConnor, 2007: 819; 2008: 221 Type locality: Tucumán, Salta Distribution: Argentina Host: Tetrapedia sp. (Hymenoptera: Apidae, Apinae) and its cleptoparasites Coelioxoides waltheriae and C. exulans Roubikia latebrosa Klimov et OConnor Roubikia latebrosa Klimov & OConnor in Klimov et al., 2007b: 118; Klimov et al., 2007a: 1371; Klimov & OConnor, 2008: 103 Type locality: Loreto Distribution: Peru Host:Tetrapedia sp. (Hymenoptera: Apidae, Apinae) Roubikia officiosa Klimov et OConnor Roubikia officiosa Klimov & OConnor, 2007: 821; 2008: 107 Type locality: Jalisco Distribution: Mexico Host: Tetrapedia maura (Hymenoptera: Apidae, Apinae) Roubikia panamensis (Baker, Roubik et Delfinado-Baker) Chaetodactylus panamensis Baker, Roubik & Delfinado-Baker, 1987: 67; Roubik, 1987: 75 Roubikia panamensis (Baker, Roubik et Delfinado-Baker): OConnor, 1993 : 345; Van Asselt, 2000: 225; Klimov & OConnor, 2007: 819; Klimov et al., 2007a: 1371; 2007b: 116; Klimov & OConnor, 2008: 103 Type locality: Panama, Curundú Distribution: Bolivia, Brazil, French Guiana, Mexico, Panama Hosts: Tetrapedia diversipes, T. peckoltii (Hymenoptera: Apidae, Apinae), also phoretic on Coelioxoides waltheriae (cleptoparasite of Tetrapedia diversipes) Acarorum Catalogus IX 267 Genus Sennertia Oudemans Sennertia Oudemans, 1905: 21 Type species: Pediculus cerambycinus Scopoli, 1763 Subgenus Afrosennertia Fain Afrosennertia Fain, 1981b: 147 Asiosennertia Fain, 1981b: 147, type species: Sennertia (Afrosennertia) delfinadoae Fain, 1981,synonymized by Klimov & OConnor, 2008: 147 Type species: Sennertia monicae Fain, 1971 Sennertia (Afrosennertia) basilewskyi Fain Sennertia basilewskyi Fain, 1974: 215; Klimov & OConnor, 2008: 223 Sennertia (Afrosennertia) basilewskyi: Fain, 1981b: 180; Fain, 1982: 67 Type locality: Bambesa, Haut-Congo Prov. Distribution: DR Congo Hosts: Hymenoptera (Megachilidae); Chedrion nigrihirtum, Mesotrichia africana Sennertia (Afrosennertia) delfinadoae Fain Sennertia (Asiosennertia) delfinadoae Fain, 1981b: 172 Sennertia delfinadoae Fain: Klimov & OConnor, 2008: 227 Sennertia bakeri Ramaraju & Mohanasundaram, 2001: 109 (Type: Tamil Nadu Coimbatore; Klimov & OConnor, 2008: 227) Type locality: India Distribution: India Host: Xylocopa (Nodula) amethystina (Hymenoptera: Apidae, Xylocopinae) Sennertia (Afrosennertia) jeanalexi Fain Sennertia jeanalexi Fain, 1971: 269; Fain, 1974a: 216; Klimov & OConnor, 2008: 230 Sennertia (Afrosennertia) jeanalexi: Fain, 1981b: 178; Fain, 1982: 67; OConnor, 1993a: 362 Type locality: Bubulu river, near Mvungu, Mayumbe Distribution: DR Congo Hosts: Xylocopa (Koptortosoma) nigrita, X. (K.) lepeletieri (Hymenoptera: Apidae, Xylocopinae) Sennertia (Afrosennertia) monicae Fain Sennertia monicae Fain, 1971: 269; 1974a: 215; Klimov & OConnor, 2008: 233 Sennertia (Afrosennertia) monicae: Fain, 1981b: 178; 1982: 67 Type locality: Kivu, Uvira Distribution: DR Congo Host: Xylocopa (Xenoxylocopa) inconstans (as Mesotrichia inconstans) (Hymenoptera: Apidae, Xylocopinae) 268 Acarorum Catalogus IX Sennertia (Afrosennertia) oudemansi Zachvatkin Sennertia oudemansi Zachvatkin, 1941: 390; Turk, 1948: 84; Klimov & OConnor, 2008: 233 Sennertia (Asiosennertia) oudemansi: Fain, 1981b: 176 Type locality: Japan Distribution: Japan Host: Xylocopa (Biluna) nasalis (as Xylocopa dissimilis) (Hymenoptera: Apidae, Xylocopinae) Sennertia (Afrosennertia) queenslandica Womersley Sennertia queenslandica Womersley, 1941: 479; Klimov & OConnor, 2008: 234 Sennertia (Asiosennertia) queenslandica: Fain, 1981b: 176 Sennertia (Afrosennertia) queenslandica: Fain, 1982: 68 Type locality: Queensland, «Moa Is., Torres Straits” Distribution: Australia Host: Xylocopa (Koptortosoma) bryorum (as Mesotrichia bryorum) (Hymenoptera: Apidae, Xylocopinae) Sennertia (Afrosennertia) vitzthumi Fain Sennertia (Asiosennertia) vitzthumi Fain, 1981b: 172 Sennertia vitzthumi Fain: Klimov & OConnor, 2008: 237 Type locality: Unknown Distribution: Unknown (probably Oriental) Host: Unknown Subgenus Amsennertia Fain Amsennertia Fain, 1981a: 147 Type species: Sennertia frontalis Vitzthum, 1941 Sennertia (Amsennertia) americana Delfinado et Baker Sennertia americana Delfinado & Baker, 1976: 84; OConnor, 1988: 341; 1993b: 164; Ramaraju & Mohanasundaram, 2001: 107; Klimov & OConnor, 2008: 173 Sennertia (Amsennertia) americana (Delfinado et Baker): Fain, 1981b: 147; Alzuet & Abrahamovich, 1987: 350; Lombert et al., 1987: 113; OConnor, 1993a: 362 Type locality: New York (Albany Co., Albany) Distribution: USA Host: Xylocopa (Xylocopoides) virginica, X. (Xylocopoides) virginica texana (Hymenoptera: Apidae, Xylocopinae) Sennertia (Amsennertia) faini Baker et Delfinado-Baker Sennertia faini Baker & Delfinado-Baker, 1983: 117; Alzuet & Abrahamovich, 1987: 350; Ramaraju & Mohanasundaram, 2001: 107 (Senertia); Klimov & OConnor, 2008: 170 Acarorum Catalogus IX 269 Type locality: Sololá, San Lucas Tolimán Distribution: Guatemala Host: Apis mellifera (Hymenoptera: Apidae, Apinae) Sennertia (Amsennertia) frontalis Vitzthum Sennertia frontalis Vitzthum, 1941: 308; Baker & Delfinado-Baker, 1983: 119; Ramaraju & Mohanasundaram, 2001: 109; Klimov & OConnor, 2008: 179 Sennertia (Amsennertia) frontalis Vitzthum : Fain, 1981b: 176; Alzuet & Abrahamovich, 1987: 346; 1990: 630; OConnor, 1993a: 362 Type locality: Argentina, Entre Rios Distribution: Argentina, Brazil, Honduras, Mexico, Panama, Peru, Suriname, Venezuela Hosts: Xylocopa (Neoxylocopa) frontalis, X. (N.) nautlana (Hymenoptera: Apidae, Xylocopinae) Sennertia (Amsennertia) ignota Delfinado et Baker Sennertia ignota Delfinado & Baker, 1976: 85; Baker & Delfinado-Baker, 1983: 119; Klimov & OConnor, 2008: 211 Senertia ignota [sic !]: Ramaraju & Mohanasundaram, 2001: 107 Sennertia (Amsennertia) ignota: Fain, 1981a: 178; Alzuet & Abrahamovich, 1987: 346 Type locality: Peru: Piura, Talara Distribution: Peru Host: Xylocopa sp. (Hymenoptera: Apidae, Xylocopinae) Sennertia (Amsennertia) longipilis Alzuet et Abrahamovich Sennertia (Amsennertia) longipilis Alzuet & Abrahamovich, 1987: 346 Sennertia longipilis Alzuet & Abrahamovich, 1989: 115; Klimov & OConnor, 2008: 232; Montalva & Allendes, 2010: 55 Type locality: Misiones Prov. (Argentina) Distribution: Argentina, Chile Host: Xylocopa (Schonnherria) splendidula splendidula (Hymenoptera: Apidae, Xylocopinae) Sennertia (Amsennertia) splendidulae Alzuet et Abrahamovich Sennertia (Amsennertia) brevipilis Alzuet & Abrahamovich, 1987: 345 (praeoccup. from Sennertia vanderhammeni brevipilis Fain,1974) Sennertia splendidulae Alzuet & Abrahamovich, 1989: 236; 1990: 630 Sennertia (Amsennertia) splendidulae Alzuet & Abrahamovich, 1991: 1; OConnor, 1993a. 362 Type locality: La Plata, Buenos Aires Prov. Distribution: Argentina 270 Acarorum Catalogus IX Host: Xylocopa (Schonnherria) splendidula splendidula (Hymenoptera: Apidae, Xylocopinae) Subgenus Sennertia Oudemans Sennertia Oudemans, 1905: 21 (Fain, 1981b: 146) Eosennertia Kurosa, 2003: 25. Type species: Sennertia (Eosennertia) bifida Kurosa, 2003, synonymized by Klimov & OConnor, 2008 Type species: Pediculus cerambycinus Scopoli, 1763 Sennertia (Sennertia) aldeodadi Haitlinger Sennertia (Sennertia) aldeodadi Haitlinger, 2000: 18 Sennertia aldeodadi : Klimov & OConnor, 2008: 222 Type locality: Mohebourg Distribution: Mauricius Host: undetermined Anthophoridae (Hymenoptera) Sennertia (Sennertia) alfkeni (Oudemans) Trichotarsus alfkeni Oudemans, 1900: 115; 1901: 82; 1903a: 147; Vitzthum, 1912c: 233 ~part., only specimens from Xylocopa circumvolans – see Klimov & OConnor (2008). Sennertia alfkeni Oudemans, 1905a: 22; 1924: 329; Vitzthum, 1914: 323; 1919: 31; Vitzthum, 1941: 308; Zachvatkin, 1941: 389; Fain, 1974b: 229; Delfinado & Baker, 1976: 85; Okabe & Makino, 2002: 73; Klimov & OConnor, 2008: 222; Kontschán et al., 2016: 3 Trichotarsus japonicus Oudemans, 1900: 117, synonymized by Zachvatkin, 1941; Oudemans, 1901: 83; 1903a: 147; Trägårdh, 1904: 156 Sennertia japonica (Oudemans): Oudemans, 1905a: 22; Vitzthum, 1914: 323; 1919: 43; 1941: 308; Okabe & Makino, 2002: 73 Sennertia japonicus: Fain, 1974b: 224 Sennertia (Sennertia) japonicus: Fain, 1981b: 163 Sennertia (Sennertia) alfkeni: Fain, 1981b: 163; Fain, 1982: 70; OConnor, 1993a: 362 Sennertia ?bifilis: Womersley, 1941: 480 (after Fain, 1982); from Xylocopa (Koptortosoma) bryorum (as “Mesotricha bryorum“), Australia: Queensland Type locality: Japan Distribution: Australia, China, Japan, Korea Host: Xylocopa (Alloxylocopa) circumvolans [in Zachvatkin, 1941 as “X. kalinovskii] (Hymenoptera: Apidae, Xylocopinae) Misidentifications (after Klimov & OConnor, 2008): Trichotarsus Alfkeni: Vitzthum, 1912c: 233, from Xylocopa (Koptortosoma) aestuans (East Sumatra), Xylocopa (Koptortosoma) caerulea (Java) Sennertia (Sennertia) antarctica (Trägårdh) Trichotarsus antarcticus Trägårdh, 1907: 12 Acarorum Catalogus IX 271 Chaetodactylus antarcticus: Pugh, 1993: 373 Sennertia antarctica (Trägårdh): Klimov & OConnor, 2008: 222 Type locality: Booth Is. (as Wandel Is.) Distribution: Antarctica (The geographic locality is, most likely, erroneous because carpenter bees, the only hosts of Sennertia, do not occur in Antarctica - Klimov & Oconnor, 2008) Host: marin algae Sennertia (Sennertia) augustii Alzuet et Abrahamovich Sennertia augustii Alzuet & Abrahamovich, 1990: 628; Klimov & OConnor, 2008: 192 Type locality: Buenos Aires, Berazategui Distribution: Argentina, Mexico, Uruguay Hosts: Xylocopa (Neoxylocopa) augusti, X. fimbriata (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) benoiti Fain Sennertia benoiti Fain, 1974: 218; 1980: 988; Klimov & OConnor, 2008: 223 Sennertia (Sennertia) benoiti Fain: Fain, 1981a: 157; Haitlinger, 2000: 17 Type locality: Bafing, 7 km ESE Touba (Ivory Coast) Distribution: DR Congo, Ivory Coast Host: Ceratina sp. (type host), Ceratina atopura (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) bifida Kurosa Sennertia (Eosennertia) bifida Kurosa, 2003: 26 Sennertia (Sennertia) bifida: Klimov & OConnor, 2008: 224 Type locality: Hitsujigaoka, Sapporo, Hokaido Distribution: Japan (Hokkaido, Honshu and Ryukyus) Hosts: Ceratina (Ceratinidia) japonica, C. flavipes, C. okinawana (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) bifilis (Canestrini) Trichotarsus bifilis Canestrini, 1897: 474; Canestrini & Kramer, 1899: 149; Giard, 1900: 377; Oudemans, 1901: 83; 1903: 147; 1905a: 22 Sennertia bifilis (Canestrini): Vitzthum, 1919: 61; Zachvatkin, 1941: 42; Fain, 1981b: 180; 1982: 70; Lombert et al., 1987: 113; Klimov & OConnor, 2008: 224 Type locality: Madang: Astrolabe Bay (Erima) Distribution: Papua New Guinea Host: Xylocopa (Koptortosoma) combinata (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) caffra Vitzthum Sennertia caffra Vitzthum, 1919: 53; 1941: 310; Zachvatkin, 1941: 42; Klimov & OConnor, 2008: 224 Sennertia (Sennertia) caffra: Fain, 1981b: 159; Lombert et al., 1987: 113; Haitlinger, 2000: 17 272 Acarorum Catalogus IX Type locality: Eastern Cape (Willowmore) Distribution: South Africa Host: Xylocopa (Koptortosoma) caffra (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) cantabrica Zachvatkin Sennertia cantabrica Zachvatkin, 1941: 385; Lombert et al., 1987: 113; Klimov & OConnor, 2008: 224 Sennertia (Sennertia) cantabrica: Fain, 1981b: 163; Haitlinger, 2000: 17 Type locality: Algeria Distribution: Algeria Host: Xylocopa (X.) cantabrita (Hymenoptera: Apidae, Xylocopinae) Sennertia (Senneria) capensis Fain Sennertia capensis Fain, 1971: 266; Klimov & OConnor, 2008: 225 Sennertia (Sennertia) capensis: Fain, 1981b: 157; Haitlinger, 2000: 17 Type locality: Eastern Cape (Willowmore) Distribution: South Africa Host: Xylocopa (Gnathoxylocopa) sicheli (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) cerambycina (Scopoli) Pediculus cerambycinus Scopoli, 1763: 386 Trichodactylus osmiae: Gervais, 1844: 266 (part.) Trichodactyle Xylocopae Donnadieu, 1868: 84 Trichodactylus xylocopae: Murray, 1877: 252 (part.); Canestrini & Fanzago, 1878: 205; Mégnin, 1880: 147; Berlese, 1885: XVIII, No 1; Canestrini & Berlese, 1885: 206 Trichotarsus xylocopae: Dugès,1834; Canestrini, 1888a: 394; 1888b: 23; 1897: 474; Berlese, 1892: fasc. LXV, n 1; 1897: 105; 1903a: 322; Canestrini & Kramer, 1899: 149; Giard, 1900: 377; Oudemans, 1900: 116; 1901: 81; 1902: 44; 1903a: 322; Trägårdh, 1907: 12; Popovici-Baznosanu, 1913: 32; Vitzthum, 1912c: 232; 1912d: 292 Trichotarsus cerambycinus : Oudemans, 1905b: 80 Sennertia cerambycina (Scopoli): Oudemans, 1905: 21; Vitzthum, 1919: 31; 1929: 77; 1933: 152; Vitzthum, 1941: 307; Zachvatkin, 1941: 382; Eyndhoven, 1941: 325; 1952: 34; Skaife, 1952: 76; Fain, 1971: 265; Elbadry, 1971: 87; Vicidomini & Meloni, 1999: 138; Klimov & OConnor, 2008: 226 Sennertia xylocopae: Türk & Türk, 1957: 210 Sennertia (Sennertia) cerambycina (Scopoli): Fain, 1981b: 152; Lombert et al., 1987: 113; OConnor, 1993a: 362; Vicidomini, 1996: 71 Type locality: “Carniola” (now Slovenia) Acarorum Catalogus IX 273 Distribution: France, Germany, Greece, Holland, Italy, Slovenia, Spain, China; “All southern, central, and eastern Europe, up to 55, 8 N; West and Central Asia up to Tibet and Gashun Gobi Desert China, on the east and up to northern Iran to the south”. Hosts: Xylocopa valga, X. varentzowi, X. violacea, Xylocopa (Proxylocopa) nitidiventris, Xylocopa (P.) przewalskyi, Xylocopa (Copoxyla) turanica (Hymenoptera: Apidae, Xylocopinae). Also phoretic on Polochrum repandum (Hymenoptera, Sapygidae, cleptoparasite of Xylocopa, occasionally found on Hoplitis princeps (as Osmia princeps), Osmia (Monosmia) apicata (as Osmia macroglossa), Megachile (Eutricharaea) leachella (as Megachile argentata), and Anthophora (Paramegilla) balassogloi(as Anthidium christofi) Sennertia (Sennertia) ceratinarum Fain Sennertia ceratinarum Fain, 1974 : 216; 1980: 983; Klimov & OConnor, 2008: 226 Sennertia (Sennertia) ceratinarum Fain: Fain, 1981b: 156; Haitlinger, 2000: 17 Type locality: Salt Rock, Natal Distribution: Cameroon, D.R. Congo, South Africa Hosts: Ceratina atopura, C. acutipyga, C. congoensis, C. nilotica, C. turneri, Pithitis viridis (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) congoicola Fain Sennertia congoicola Fain, 1971: 265; Klimov & OConnor, 2008: 227 Sennertia (Sennertia) congoicola: Fain, 1981b: 161; Haitlinger, 1999: 59; 2000: 17 Type locality: Moanda Distribution: DR Congo Hosts: “Mesotrichia striata”, Xylocopa (Mesotrichia) torrida, Xylocopa (Koptortosoma) imitator (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) dalyi Fain Sennertia dalyi Fain, 1980: 990; Klimov & OConnor, 2008: 227 Sennertia (Sennertia) dalyi: Fain, 1981b: 163; Haitlinger, 2000: 17 Type locality: 3.5 km S Tabarka Distribution: Tunisia Hosts: Ceratina (Euceratina) dallatorreana, C.(E.) chalybea, C. (E.) mocsaryi, C. (E.) callosa (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) dalyi nilotica Fain Sennertia dalyi nilotica Fain, 1980: 990; Klimov & OConnor, 2008: 227 Sennertia (Sennertia) dalyi nilotica: Fain, 1981b: 163 Type locality: Al QalyuS bıyah, Banha (“Benha, Nile Delta”) Distribution: Egypt Host: Ceratina (Pithitis) tarsata (Hymenoptera: Apidae, Xylocopinae) 274 Acarorum Catalogus IX Sennertia (Sennertia) devincta Klimov et OConnor Sennertia devincta Klimov & OConnor, in Klimov et al., 2007: 121; Klimov & OConnor, 2008: 148 Type locality: Huanuco Distribution: Peru Host: Ceratina sp. (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) dissimilis Zachvatkin Sennertia dissimilis Zachvatkin, 1941: 385; Fain, 1981b: 182; Klimov & OConnor, 2008: 227 Type locality: Japan Distribution: Japan Host: Xylocopa (Biluna) nasalis (as Xylocopa dissimilis) (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) duweinii Sherbef et Duweini Sennertia duweinii Sherbef & Duweini, 1980: 245; Klimov & OConnor, 2008: 228 Type locality: not indicated Distribution: ? Egypt Host: Xylocopa (Koptortosoma) aestuans (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) egyptiaca Elbadry Sennertia egyptiaca Elbadry, 1971: 87; Sherbef & Duweini, 1980: 246; Fain, 1981b: 182; Klimov & OConnor, 2008: 228 Type locality: Shoubra Elkheima, Cairo Distribution: Egypt, Sudan Host: Xylocopa (Koptortosoma) aestuans (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) elseni Fain Sennertia elseni Fain, 1971: 266; Klimov & OConnor, 2008: 228 Sennertia (Sennertia) elseni: Fain, 1981b: 161; OConnor, 1993a: 362; Haitlinger, 1999: 59; 2000: 17; Fain & Pauly, 2001: 132 Type locality: Bas-Congo (Muanda) Distribution: DR Congo, Madagascar Hosts: Xylocopa (Koptortosoma) caffra (as Mesotrichia olivacea) (Hymenoptera: Apidae, Xylocopinae); on Madagascar: except Ranomafana: Melanempis sp.; Ranomafana: cleptoparasitic bee of the family Nomadinae Acarorum Catalogus IX 275 Sennertia (Sennertia) flabellifera Oudemans Sennertia flabellifera Oudemans, 1924: 331; Lombert et al., 1987: 113; Klimov & OConnor, 2008: 228 Type locality: Bogor (Java) Distribution: Indonesia (Java) Host: Xylocopa (Koptortosoma) sp., nest (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) frontalis Vitzthum Sennertia frontalis Vitzthum, 1941: 308; Klimov & OConnor, 2008: 228 Type locality: Entre Rios, Argentina Distribution: Argentina, Brazil, Mexico, Peru, Panama, Suriname, Venezuela Hosts: Xylocopa frontalis, X. nautlana (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) gargantua Zachvatkin Sennertia gargantua Zachvatkin, 1941: 385; Klimov & OConnor, 2008: 228 Sennertia (Sennertia) gargantua: Fain, 1981b: 154 Type locality: Turkey Distribution: Tajikistan, Turkey, Uzbekistan Hosts: Xylocopa (Nodula) punctilabris, X. (X.) valga (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) greeni (Oudemans) Hericia greeni Oudemans, 1917: 345 Sennertia greeni (Oudemans): Oudemans, 1924: 330; Zachvatkin, 1941: 42; Lombert et al., 1987: 113; Klimov & OConnor, 2008: 228 Type locality: Eton Estate, Punduloga Distribution: Sri Lanka Host: Xylocopa (Mesotrichia) tenuiscapa (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) haustrifera Klimov et OConnor Sennertia haustrifera Klimov & OConnor, 2008: 151 Type locality: Jalisco, Estacion de Biologia Chamela Distribution: Mexico (Jalisco, Chiapas) Host: Ceratina captiosa (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) herminae Haitlinger Sennertia (Sennertia) herminae: Haitlinger, 1999: 57; 2000: 17 Sennertia herminae: Klimov & OConnor, 2008: 229 Type locality: Antananarivo Distribution: Madagascar Host: Xylocopa sp. (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) hipposideros (Oudemans) 276 Acarorum Catalogus IX Trichotarsus hipposideros Oudemans, 1902: 44; 1903: 145 Trichotarsus hipposiderus Oudemans, 1903a: 145 (unjust. emend.); Trägårdh, 1904: 156; 1907: 12; Vitzthum, 1912d: 290 Sennertia hipposiderus (Oudemans): Oudemans, 1905a: 22; Vitzthum, 1941: 310 (hipposideruns); Fain, 1974b: 226; OConnor, 1993b: 161; Krantz, 1998: 298 Sennertia hipposidera: Vitzthum, 1919: 43 (unjust. emend.) Sennertia hipposideros: Zachvatkin, 1941: 42; Fain, 1974 : 226; Klimov & OConnor, 2008: 229 Sennertia (Sennertia) hipposiderus (Oudemans): Fain, 1981b: 156 Type locality: India Distribution: India, Malaysia Hosts: Xylocopa (Platynopoda) latipes, X. (Mesotrichia) tenuiscapa (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) horrida (Vitzthum) Trichotarsus horridus Vitzthum, 1912: 290 Sennertia horrida (Vitzthum): Vitzthum, 1919: 57; 1941: 308; Zachvatkin, 1941: 387; Klimov et al., 2007b: 124; Klimov & OConnor, 2008: 229 Sennertia (Sennertia) horrida: Fain, 1981b: 170 Sennertia sumatrensis Oudemans, 1924: 329; Fain, 1974b: 228 (synonymized by Fain, 1981b) Sennertia leucothorae Ramaraju & Mohanasundaram, 2001: 107 (Type: Tamil Nadu, Coimbatore; synonymized by Klimov et al., 2007b) Type locality: Jakarta (Java) Distribution: India, Indonesia (Java, Sumatra), Japan, Taiwan Hosts: Xylocopa (Biluna) nasalis (as Xylocopa (Koptorthosoma?) dissimilis), Xylocopa (Koptortosoma) aestuans (as Xylocopa leucothorae), X. auripennis, X. fallax, X. iridipennis, X. mcgregori, X. mimetica (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) hurdi Klimov et OConnor Sennertia hurdi Klimov & OConnor, 2008: 164 Type locality: Oaxaca, Presa Benito Juárez, nr. Jalapa de Marquez Distribution: Mexico Hosts: Xylocopa t. tabaniformis, X. tabaniformis azteca (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) indica Delfinado et Baker Sonnertia [sic !] indica Delfinado & Baker, 1976: 87 Seneria [sic !] indica: Ramaraju & Mohanasundaram, 2001: 107 Sennertia indica: Fain, 1981b: 182 (inadequately described); Klimov & OConnor, 2008: 230 Type locality: Hissar Acarorum Catalogus IX 277 Distribution: India Host: Ceratina binghami (as Pithitis binghami) (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) koptorthosomae (Oudemans) Trichotarsus koptorthosomae Oudemans, 1901: 82; 1903a: 47; Vitzthum, 1912d: 290 (part.) Trichotarsus coptorthosomae Trägårdh, 1904: 156 (unjust. emend.) Sennertia coptorthosomae: Oudemans, 1905a: 22 (unjust. emend.) Sennertia koptorthosomae (Oudemans): Oudemans, 1924: 329; Zachvatkin, 1941: 42; Fain, 1974: 219; OConnor 1993b: 161; Krantz, 1998: 298; Klimov & OConnor, 2008: 192 Sennertia (Sennertia) koptorthosomae (Oudemans): Fain, 1981b: 156 Type locality: Java (Bogor) Distribution: India, Indonesia (Java, Sumatra), Malaysia, Sri Lanka Hosts: nest of Xylocopa (Mesotrichia) latipes, X. (M.) tenuiscapa [Koptorthosoma] (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) latipilis Fain Sennertia latipilis Fain, 1974: 217; 1980: 986; Klimov & OConnor, 2008: 231 Sennertia (Sennertia) latipilis: Fain, 1981a: 156; Haitlinger, 2000: 17 Type locality: Malindi Distribution: Kenya Host: Ceratina (Ctenoceratina) penicilligera (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) lauta Klimov and OConnor Sennertia lauta Klimov & OConnor, in Klimov et al., 2007b: 125; Klimov & OConnor, 2008: 231 Type locality: Davao (Philippines) Distribution: Malaysia (Sarawak), Philippines (Mindanao, Palawan) Hosts: Xylocopa (Zonohirsuta) fuliginata, X. (Z.) dejeanii (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) leclercqi Fain Sennertia leclercqi Fain, 1971: 268; Klimov & OConnor, 2008: 232 Sennertia (Sennertia) leclercqi: Fain, 1981b: 157; Haitlinger, 2000: 17 Type locality: Sud-Kivu, Uvira Distribution: DR Congo Host: Xylocopa (Xenoxylocopa) inconstans (as Mesotrichia inconstans) (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) leei Fain Sennertia (Sennertia) leei Fain, 1982: 67; OConnor, 1993a: 362 Sennertia leei Fain: Klimov & OConnor, 2008: 150, 192 278 Acarorum Catalogus IX Type locality: Ku-rin-gai, New South Wales Distribution: Australia (N.S.W., Queensland, SA) Hosts: Xylocopa (Lestis) bombylans, X. aerata (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) loricata Klimov et OConnor Sennertia loricata Klimov & OConnor, 2008: 181 Type locality: Florida, Alachua Co., Gainesville Distribution: Mexico (Chiapas, Jalisco), USA (Florida, Louisiana, Puerto Rico, Texas, Virginia) Hosts: Xylocopa b. barbatella, X. micans, X. mordax, X. muscaria (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) lucrosa Klimov and OConnor Sennertia lucrosa Klimov & OConnor, 2008: 164 Type locality: California, Los Angeles Co., Los Angeles, Crenshaw Distr. Distribution: Mexico, USA Hosts: Xylocopa californica, X. clarionensis, X. guatemalensis, X. mexicanorum, X. varipuncta (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) madagascarensis Fain Sennertia madagascarensis Fain, 1971: 270; Fain & Pauly, 2001: 125; Klimov et al., 2007b; Klimov & OConnor, 2008: 232 Sennertia (Sennertia) madagascarensis: Fain, 1981b: 170; OConnor, 1993a: 362 Type locality: Antananarivo (“Tananarive”) Distribution: Madagascar Hosts: Xylocopa (Prosopoxylocopa) mirabilis, Xylocopa (Koptortosoma) calens (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) mesotrichia Fain Sennertia mesotrichia Fain, 1971: 265; Klimov & OConnor, 2008: 232 Sennertia (Sennertia) mesotrichia: Fain, 1981b: 154; Haitlinger, 2000: 17 Type locality: Bambesa, Uele Distribution: D.R. Congo Hosts: Xylocopa (Koptortosoma) africana (as Mesotrichia africana), X. (K.) lepeletieri, Xylocopa (Xylomelissa) tarsata (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) micheli Fain Sennertia micheli Fain, 1971: 268; Klimov & OConnor, 2008: 232 Sennertia (Sennertia) micheli: Fain, 1981b: 164 Type locality: Java Distribution: Indonesia (Java) Host: Bembix borrei (Hymenoptera: Sphecidae) Acarorum Catalogus IX 279 Sennertia (Sennertia) morstatti Vitzthum Sennertia morstatti Vitzthum, 1914: 323; 1919: 42; Zachvatkin, 1941: 42; Klimov & OConnor, 2008: 233 Sennertia (Sennertia) morstatti: Fain, 1981b: 164; Lombert et al., 1987: 113; Sherbef & Duweini, 1980: 245 Sennertia moandensis Fain, 1971: 267 (synonymized by Fain, 1981b). Type: Moanda, DR Congo Sennertia tanythrix Fain, 1971: 267 (part., specimens from X. nigrita) Type locality: Tanzania, Amani Distribution: DR Congo, Sudan, Tanzania Hosts: Xylocopa (Koptortosoma) nigrita (Hymenoptera: Apidae, Xylocopinae), Megachilidae Sennertia (Sennertia) perturbans Vitzthum Sennertia perturbans Vitzthum, 1919: 48; Zachvatkin, 1941: 42; Fain, 1981b: 145; Lombert et al., 1987: 113; Klimov & OConnor, 2008: 233 Type locality: Amani Distribution: Tanzania Host: Xylocopa (Koptortosoma) nigrita (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) pirata Klimov et OConnor Sennertia pirata Klimov & OConnor, 2008: 144 Type locality: Bahamas, Bimini, South Bimini Island Distribution: Bahamas, British Virgin Islands (Guana Is.), Cuba, Virgin Islands of the United States, Puerto Rico Hosts: Xylocopa cubaecola, X. mordax (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) potanini Zachvatkin Sennertia potanini Zachvatkin, 1941: 390; Klimov & OConnor, 2008: 233 Sennertia (Sennertia) potanini: Fain, 1981b: 170; Klimov et al., 2007b: 125 Type locality: Sichuan, Yazhou Distribution: China Host: Xylocopa (Biluna) tranquebarorum or Xylocopa (Bomboixylocopa) rufipes (as Xylocopa pictifrons) (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) ratiocinator Klimov et OConnor Sennertia ratiocinator Klimov et OConnor, in Klimov et al., 2007b: 128; Klimov & OConnor, 2008: 234 Type locality: NW Sri Lanka Distribution: Sri Lanka 280 Acarorum Catalogus IX Hosts: Xylocopa (Zonohirsuta) bhowara, X. (Z.) dejeanii, Xylocopa (Nodula) amethystina (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) recondita Klimov et OConnor Sennertia recondita Klimov & OConnor, 2008: 151 Type locality: Quintana Roo, Vallarta, 17 km W Pto. Morelos Distribution: Mexico Host: Ceratina (Calloceratina) exemia (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) robusta Delfinado et Baker Sennertia robusta Delfinado et Baker, 1976: 87; Klimov & OConnor, 2008: 234 Sennertia (Sennertia) robusta: Fain, 1981b: 157; Haitlinger, 2000: 18 Sennertia carpenteri Ramaraju & Mohanasundaram, 2001: 109 – Type: Tamil Nadu, Coimbatore Type locality: Hissar Distribution: India Hosts: «Megachilid bee«, Xylocopa (Nodula) amethystina (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) roepkei Oudemans Sennertia roepkei Oudemans, 1924: 330; Lombert et al., 1987: 113; Klimov & OConnor, 2008: 234 Type locality: Java (Bogor) Distribution: Indonesia (Java) Host: On eggs of Horia (as “Cissites)”(Coleoptera: Meloidae) in nest of Xylocopa sp. (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) sayutara Klimov et OConnor Sennertia sayutara Klimov & OConnor, in Klimov et al., 2007: 122; 2008: 161 Type locality: San José Distribution: Costa Rica Host: Ceratina laticeps (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) scutata Fain Sennertia scutata Fain, 1974: 217; 1980: 986; Klimov & OConnor, 2008: 234 Sennertia (Sennertia) scutata: Fain, 1981b: 159; Haitlinger, 2000: 17 Type locality: Cape Town Distribution: South Africa, El Salvador, Guatemala Hosts: Ceratina truncata, Ceratina (Pithitis) nasalis (as Ceratina viridior) (Hymenoptera: Apidae, Xylocopinae) Acarorum Catalogus IX 281 Sennertia (Sennertia) segnis Klimov et OConnor Sennertia segnis Klimov & OConnor, 2008: 170 Type locality: California, Tulare Co., 2 mi N Hartland Distribution: Mexico, USA (California, Arizona) Hosts: Xylocopa c. californica, X. californica arizonensis, X. cyanea, X. tabaniformis orpifex (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) shimanukii Baker et Delfinado-Baker Sennertia shimanukii Baker & Delfinado-Baker, 1983: 117; Klimov & OConnor, 2008: 176 Type locality: Sololá, San Lucas Tolimán (Guatemala) Distribution: Guatemala, Mexico, USA Hosts: Xylocopa (Neoxylocopa) fimbriata, X. (N.) mexicanorum, X. (N.) sonorina, X. (N.) varipuncta, Xylocopa (Xylocopoides) californica arizonensis (Hymenoptera: Apidae, Xylocopinae); Apis mellifera (Hymenoptera: Apidae, Apinae)(type host, accidental) Sennertia (Sennertia) simplex (Trägårdh) Trichotarsus simplex Trägårdh, 1904: 156 Sennertia simplex (Trägårdh): Vitzthum, 1919: 32; 1941: 308; Zachvatkin, 1941: 42; Klimov & OConnor, 2008: 235 Sennertia (Sennertia) simplex: Fain, 1981b: 159; Haitlinger, 2000: 17 Type locality: Cameroon Distribution: Cameroon Host: Xylocopa (Koptortosoma) nigrita (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) sodalis Klimov et OConnor Sennertia sodalis Klimov & OConnor, 2008: 156 Type locality: San Juan de la Punta (Veracruz) Distribution: Costa Rica, Mexico, Panama Host: Ceratina sp., Ceratina (Calloceratina) eximia (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) spinifera Fain Sennertia ceratinarum spinifera Fain, 1974: 217 Sennertia (Sennertia) spinifera Fain, 1980: 984; 1981b: 156; Haitlinger, 2000: 17 Sennertia spinifera Fain: Klimov & OConnor, 2008: 235 Type locality: N. E. Lusambo Distribution: D.R. Congo Host: Ceratina (Propithitis) aereola (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) surinamensis Fain et Lukoschus Sennertia surinamensis Fain & Lukoschus, 1971: 270; Klimov et al., 2007a: 1371; Klimov & OConnor, 2008: 237 282 Acarorum Catalogus IX Sennertia (Sennertia) surinamensis (Fain et Lukoschus): Fain, 1981b: 170; Alzuet & Abrahamovich, 1987: 348 Type locality: Paramaribo Distribution: French Guiana, Panama, Suriname Host: Ceratina (Calloceratina) chloris (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) tanythrix Fain Sennertia tanythrix Fain, 1971: 267; Klimov & OConnor, 2008: 236 Sennertia (Sennertia) tanythrix: Fain, 1981b: 167; Haitlinger, 2000: 17 Type locality: Uele, Haut-Congo Prov. Distribution: DR Congo Hosts: Xylocopa (Mesotrichia) torrida, Xylocopa (Koptortosoma) imitator, X. (K.) nigrita (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) tunisiana Fain Sennertia tunisiana Fain, 1980: 991; Klimov & OConnor, 2008: 236 Sennertia (Sennertia) tunisiana: Fain, 1981b: 163; Haitlinger, 2000: 17 Type locality: Silya nah Maktar Distribution: Tunisia Hosts: Ceratina (Ceratina) cucurbitina, Ceratina (Euceratina) albosticta, Ceratina (Euceratina) mocsaryi, Ceratina dallatorreana, C. chalybea (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) vaga Klimov et OConnor Sennertia vaga Klimov & OConnor, 2008: 150 Type locality: California, Los Angeles Co., Rustic Canyon Distribution: Mexico, USA Hosts: Xylocopa (Notoxylocopa) tabaniformis orpifex, X. (N.) tabaniformis androleuca, X. (Xylocopoides) californica (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) vanderhammeni Fain Trichotarsus koptorthosomae Oudemans, 1901: 82 (part.) Sennertia vanderhammeni Fain, 1974: 222; Klimov & OConnor, 2008: 236 Sennertia (Sennertia) vanderhammeni: Fain, 1981b: 156 Type locality: Peradeniya Distribution: India, Indonesia (Java), Sri Lanka Host: Xylocopa (Koptorthosoma) tenuiscapa (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) vanderhammeni brevipilis Fain Trichotarsus koptorthosomae Oudemans, 1901: 82 (part.) Acarorum Catalogus IX 283 Sennertia vanderhammeni brevipilis Fain, 1974: 224; 1981b: 156; Klimov & OConnor, 2008: 236 Sennertia (Sennertia) vanderhammeni brevipilis: Fain, 1981b: 156; Alzuet & Abrahamovich, 1989: 236; 1990: 319 Type locality: Java Distribution: Indonesia (Java) Host: Xylocopa (Koptorthosoma) tenuiscapa (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) varicosa Fain . Sennertia varicosa Fain, 1971: 268; Klimov & OConnor, 2008: 237 Sennertia (Sennertia) varicosa: Fain, 1981b: 167; Haitlinger, 2000: 17 Type locality: Katanga – Monts Kundelungu (DR Congo) Distribution: Angola, DR Congo Hosts: Xylocopa (Mesotrichia) sp., X. (Xenoxylocopa) inconstans (as Mesotrichia flavescens inconstans) (Hymenoptera: Apidae, Xylocopinae) Sennertia (Sennertia) zhelochovtsevi Zachvatkin Sennertia zhelochovtsevi Zachvatkin, 1941: 388; Turk, 1948: 84; Klimov et al., 2007a: 1371; Klimov & OConnor, 2008: 237; Nazan et al., 2019 Sennertia (Sennertia) zhelochovtsevi Zachvatkin: Fain, 1981b: 148 Type locality: Greece Distribution: Armenia, Greece, Iran, Turkey, Uzbekistan Hosts: Xylocopa olivieri, X. rufa (Hymenoptera: Apidae, Xylocopinae) Subgenus Spinosennertia Fain Spinosennertia Fain, 1981b: 147; Klimov & OConnor, 2008: 147 Type species: Sennertia argentina Vitzthum, 1941 Sennertia (Spinosennertia) argentina Vitzthum Sennertia argentina Vitzthum, 1941: 309; Klimov & OConnor, 2008: 156 Sennertia (Spinosennertia) argentina Vitzthum: Fain, 1981b: 147 Type locality: Argentina Distribution: Argentina, Brazil, Costa Rica, Guatemala, Mexico, Nicaragua, Panama, Peru, Suriname Hosts: Xylocopa (Neoxylocopa) fimbriata, X. (N.) frontalis, X. (N.) mexicanorum, X. (N.) nasica, X. (N.) nautlana (Hymenoptera: Apidae, Xylocopinae) Sennertia (Spinosennertia) donaldi Turk Sennertia donaldi Turk, 1948: 84; Klimov & OConnor, 2008: 148 Sennertia (Spinosennertia) argentina: Fain, 1981b: 176 (part.); Alzuet & Abrahamovich, 1987: 350 (part.); Haitlinger, 1999: 59 (part.) 284 Acarorum Catalogus IX Sennertia (Spinosennertia) donaldi: Klimov et al., 2007b: 130 Type locality: Piarco Distribution: Brazil, Trinidad and Tobago, Venezuela Hosts: Xylocopa (Neoxylocopa) fimbriata, X. (N.) frontalis (Hymenoptera: Apidae, Xylocopinae) Speciae inquirendae and Taxa Incorrectly attributed to Chaetodactylidae – see in Klimov & OConnor, 2008 Index of Chaetodactylidae (Names in bold are accepted as valid) abditus, Chaetodactylus 258 Achaetodactylus 257 Afrosennertia 265 aldeodadi, Sennertia 268 alfkeni, Trichotarsus 268 alfkeni, Sennertia 268 americana, Sennertia 267 Amsennertia 267 antarctica, Sennertia 269 antarcticus, Chaetodactylus 269 antarcticus, Trichotarsus 269 anthidii, Chaetodactylus 269 anthidii, Trichotarsus 269 antillarum, Chaetodactylus 269 argentina, Sennertia 282 Asiosennertia 265 augustii, Sennertia 269 aurulenticola, Chaetodactylus chrysidis 259 azteca, Chaetodactylus 259 bakeri, Sennertia 265 basilewskyi, Sennertia 265 basilewskyi, Sennertia (Afrosennertia) 283 benoiti, Sennertia 269 bifida, Sennertia 269 bifida, Sennertia (Eosennertia) 269 bifilis, Sennertia 270 bifilis, Trichotarsus 270 birulai, Chaetodactylus 259 brevipilis, Sennertia vanderhammeni 281 caffra, Sennertia 270 cantabrica, Sennertia 270 capensis, Sennertia 270 carpenteri, Sennertia 277 Centriacarus 258 cerambycina, Sennertia 270 cerambycinus, Pediculus 270 cerambycinus, Trichotarsus 271 ceratinae, Achaetodactylus 257 ceratinae, Chaetodactylus 257 ceratinarum, Sennertia 271 Chaetodactylidae 257 Chaetodactylinae 257 Chaetodactylini 256 Chaetodactylus 257 chrysidis, Chaetodactylus 259 claviger, Chaetodactylus 260 claviger, Chaetodactylus (Spinodactylus) 260 claudus, Chaetodactylus 260 congoicola, Sennertia 271 coptorthosomae, Sennertia 275 dalyi, Chaetodactylus 260 dalyi, Sennertia 272 decellei, Achaetodactylus 257 Acarorum Catalogus IX 285 decellei, Chaetodactylus (Ochaetodactylus) 257 decellei, Ochaetodactylus 257 delfinadoae, Sennertia 265 dementjevi, Chaetodactylus 260 devincta, Sennertia 272 dissimilis, Sennertia 272 donaldi, Sennertia 282 duweinii, Sennertia 272 egyptiaca, Sennertia 272 elseni, Sennertia 273 Eosennertia 268 faini, Sennertia 267 flabellifera, Sennertia 273 frontalis, Sennertia 267 furunculus, Chaetodactylus 260 gargantua, Sennertia 273 gibbosi, Chaetodactylus 261 greeni, Sennertia 273 greeni, Hericia 273 guahibo, Centriacarus 258 haustrifera, Sennertia 274 herminae, Sennertia 274 hipposidera, Sennertia 274 hipposideros, Sennertia 274 hipposiderus, Sennertia 274 hipposideros, Trichotarsus 274 hipposiderus, Trichotarsus 274 hirashimai, Chaetodactylus 261 hopliti, Chaetodactylus 261 horrida, Sennertia 274 horridus, Trichotarsus 274 hurdi, Sennertia 275 ignota, Sennertia 267 imberba, Roubikia 264 indica, Sennertia 275 japonica,Sennertia 268 japonicus, Sennertia 269 japonicus, Trichotarsus 269 jeanalexi, Sennertia 266 koptorthosomae, Sennertia 275 koptorthosomae, Trichotarsus 275 kouboy, Chaetodactylus 261 krombeini, Chaetodactylus 261 krombeini, Chaetodactylus (Spinodactylus) 261 lassulus, Chaetodactylus 262 latebrosa, Roubikia 264 latipilis, Sennertia 275 lauta, Sennertia 276 leclercqi, Sennertia 276 leei, Senertia 276 leleupi, Achaetodactylus 258 leleupi, Chaetodactylus 258 leucothorae, Sennertia 274 lithurgi, Chaetodactylus 262 longipilis, Sennertia 268 loricata, Sennertia 276 lucrosa, Sennertia 276 ludwigi, Chaetodactylus 262 ludwigi, Trichotarsus 262 madagascarensis, Sennertia 277 mahunkai, Chaetodactylus 261 melitomae, Chaetodactylus 262 mesotrichia, Sennertia 277 micheli, Sennertia 277 micheneri, Chaetodactylus 262 moandensis, Sennertia 277 monicae, Sennertia 266 morstatti, Sennertia 277 nilotica, Sennertia dalyi 272 nipponicus, Chaetodactylus 263 Ochaetodactylus 257 officiosa, Roubikia 265 osmiae, Chaetodactylus 263 osmiae, Trichodactylus 263 osmiae, Trichotarsus 263 oudemansi, Sennertia 266 panamensis, Chaetodactylus 265 panamensis, Roubikia 265 perturbans, Sennertia 277 pirata, Sennertia 278 poetae, Chaetodactylus 259 potanini, Sennertia 278 286 Acarorum Catalogus IX queenslandica, Sennertia 266 ratiocinator, Sennertia 278 reaumuri, Trichotarsus 263 reaumuri, Chaetodactylus 263 recondita, Sennertia 277 robusta, Sennertia 277 roepkei, Sennertia 279 Roubikia 264 rozeni, Chaetodactylus 264 sayutara, Sennertia 279 scutata, Sennertia 279 segnis, Sennertia 279 Sennertia 265 shimanukii, Sennertia 279 simplex, Sennertia 279 simplex, Trichotarsus 279 sodalis, Sennertia 280 spinifera, Sennertia 280 Spinodactylus Spinosennertia 282 splendidulae, Sennertia 268 sumatrensis, Sennertia 274 surinamensis, Sennertia 280 tanythrix, Sennertia 280 Trichotarsus tunisiana, Sennertia 280 turbator, Centriacarus 258 vaga, Sennertia 281 vanderhammeni, Sennertia 281 varicosa, Sennertia 281 vitzthumi, Sennertia 266 xylocopae, Sennertia 271 xylocopae, Trichodactylus 270 xylocopae, Trichotarsus 271 zachvatkini, Chaetodactylus 264 zhelochovtsevi, Sennertia 281 Distribution of Chaetodactylidae EUROPE Armenia (Zachvatkin, 1941) – Sennertia zhelochovtsevi Belgium (Fain & Baugnée, 1996 ; Van Asselt, 2000; Klimov & OConnor, 2008) – Chaetodactylus chrysidis, Ch. osmiae Bulgaria (Beron, 2010) – Sennertia sp. Croatia (Klimov & OConnor, 2008) – Chaetodactylus osmiae Czech Rep. (Oudemans, 1905) – Chaetodactylus birulai (= poetae), Ch. reaumuri France (contin.) (Dufour, 1839;) – Chaetodactylus anthidii, Ch. claviger, Ch. osmiae, Sennertia cerambycina Georgia (Klimov & OConnor, 2008) - Chaetodactylus osmiae, Ch. reaumuri, Germany (Oudemans, 1905) – Chaetodactylus chrysidis, Ch. osmiae, Ch. reaumuri, Sennertia cerambycina Great Britain (Michael, 1901; Turk, 1953) – Chaetodactylus osmiae Greece (Zachvatkin, 1941) – Ch. reaumuri, Sennertia cerambycina, S. zhelochovtsevi Holland (Eyndhoven, 1952; Siepel, Cremers & Vierbergen, 2016) – Chaetodactylus birulai, Ch. osmiae, Sennertia cerambycina Hungary (Samšinak, 1973; Klimov & OConnor, 2008) – Chaetodactylus osmiae (= mahunkai) Acarorum Catalogus IX 287 Italy (contin.) (Oudemans, 1924, 1928; Fain, 1981b; Klimov & OConnor, 2008) – Chaetodactylus claviger, Ch. osmiae, Ch. reaumuri, Ch. zachvatkini, Sennertia cerambycina Luxemburg - Chaetodactylus osmiae Poland (Chmielewski, 1993) – Chaetodactylus osmiae Romania (Popovici-Baznosanu, 1913; Klimov & OConnor, 2008) – Chaetodactylus osmiae Russia (all, continent.) (Zachvatkin, 1941) – Chaetodactylus birulai, Ch. dementjevi, Ch. osmiae Crimea (Zachvatkin, 1941) - Chaetodactylus reaumuri Serbia (Krunić et al., 2005) – Chaetodactylus osmiae Slovenia (Scopoly, 1763) – Sennertia cerambycina Spain (cont.) (Fain, 1981b; Bosch, 1992) – Chaetodactylus osmiae, Ch. reaumuri, Ch. zachvatkini, Sennertia cerambycina Turkey (all) (Zachvatkin, 1941; Fain, 1981b) – Ch. reaumuri, Sennertia gargantua, S. zhelochovtsevi Western Palaearctic (Zachvatkin, 1941) – Chaetodactylus dementjevi AFRICA Algeria (Zachvatkin, 1941) – Sennertia cantabrica Angola Sennertia varicosa Cameroon (Trägårdh, 1904; Klimov & OConnor, 2008) – Achaetodactylus decellei, A. leleupi, Sennertia af. basilewskyi, S. simplex Congo DR (Fain, 1981b) – Achaetodactylus decellei, A. leleupi, Sennertia basilewskyi, S. benoiti, S. ceratinarum, S. elseni, S. jeanalexi, S. leclercki, S. mesotrichia, S. monicae, S. morstatti, S. spinifera, S. tanythrix, S.varicosa Egypt (Elbadry, 1971; Sherbef & El-Duweini, 1980; Faini, 1981a) – Chaetodactylus anthidii, Ch. claviger, Sennertia dalyi nilotica, S. duweinii, S. egyptiaca Ivory Coast (Fain, 1974) – Sennertia benoiti Kenya (Fain, 1974) – Achaetodactylus ceratinae, A. leleupi, Sennertia latipilis Madagascar (Fain, 1971; Haitlinger, 1999; Fain & Pauly, 2001) – Chaetodactylus dalyi, Ch. ludwigi, Sennertia elseni, S. herminae, S. madagascarensis Mauricius (Haitlinger, 2000) - Sennertia aldeodadi Mozambique (Fain, 1974) – Chaetodactylus dalyi South Africa (Fain, 1974; Klimov & OConnor, 2008) – Achaetodactylus ceratinae, Chaetodactylus dalyi, Senertia caffra, S. capensis, S. ceratinarum, S. scutata Sudan – Sennertia egyptiaca, S. morstatti Tanzania (Vitzthum, 1914, 1919) – Achaetodactylus ceratinae, A. decellei, A. leleupi, Sennertia morstatti, S. perturbans Tunisia (Oudemans, 1911; Fain, 1980; ) – Chaetodactylus anthidii, Ch. reaumuri, Ch. zachvatkini, Sennertia dalyi, S. tunisiana Zimbabwe (Fain, 1974) – Chaetodactylus dalyi 288 Acarorum Catalogus IX ASIA China (Zachvatkin, 1941) – Sennertia alfkeni, S. cerambycinus, S. potanini Taiwan (Kawazoe et al., 2010) – Sennertia horrida India (Oudemans, 1902 ; Ramaraju & Mohanasundaram, 2001; Fain & Pauly, 2001) – Chaetodactylus ludwigi, Sennertia delfinadoae, S. hipposideros, S. horrida, S. indica, S. koptorthosomae, S. robusta (= carpenteri), S. vanderhammeni Indonesia (Vitzthum, 1912; Oudemans, 1924; Fain, 1971, 1981b; ) - Chaetodactylus ludwigi, Sennertia flabellifera, S. horrida (= sumatrensis), S. koptorthosomae, S. micheli, S. roepkei, S. vanderhammeni, S. vanderhammeni brevipilis Japan (Oudemans, 1901 ; Hirashima, 1957; Krombein, 1962; Fain, 1974 ; Kurosa, 1987, 2003; Qu et al., 2002; Okabe & Makino, 2002, 2005; Okabe, Makino & Endo, 2008; Kawazoe et al., 2010) – Chaetodactylus hirashimai, Ch. nipponicus (Tsushima Is., Honshu), Sennertia alfkeni, S. horrida, S. japonicus, S. oudemansi Kazakhstan (Zachvatkin, 1941) – Chaetodactylus dementjevi Korea (Kontschán et al., 2016) – Sennertia alfkeni Malaysia (Fain, 1981b; Klimov & OConnor, 2007) – Sennertia hipposideros, S. kopthortosomae, S. lauta Philippines (Klimov & OConnor, 2007) - Sennertia lauta Sri Lanka (Oudemans, 1917; Klimov et OConnor, 2007) – Sennertia greeni, S. kopthortosomae, S. ratiocinator, S. vanderhammeni Tajikistan (Zachvatkin, 1941) - Sennertia gargantua Uzbekistan (Zachvatkin, 1941) – Chaetodactylus dementjevi, Sennertia gargantua, S. zhelochovtsevi NORTH AMERICA Canada (Klimov & OConnor, 2008) – Chaetodactylus hopliti, Ch. krombeini, Ch. micheneri Mexico (Klimov & OConnor, 2007, 2008) – Centriacarus turbator, Chaetodactylus abditus, Ch. melitomae, Roubikia officiosa, R. panamensis, Sennertia argentina, S. augustii, S. frontalis, S. haustrifera, S. hurdi, S. loricata, S. lucrosa, S. recondita, S. segnis, S. shimanukii, S. sodalis, S. vaga USA (Osborn, 1893; Baker, 1962; Klimov & OConnor, 2008; McKinney & Park, 2012) – Chaetodactylus abditus, Ch. azteca, Ch. claudus, Ch. furunculus, Ch. gibbosi, Ch. hopliti, Ch. kouboy, Ch. krombeini, Ch. lithurgi, Ch. micheneri, Ch. rozeni, Sennertia americana, S. loricata, S. lucrosa, S. segnis, S. shimanukii, S. vaga Puerto Rico (Klimov & OConnor, 2008) - Sennertia loricata, S. pirata Virgin Islands of the USA (Klimov & OConnor, 2008) - Sennertia pirata SOUTH and CENTRAL AMERICA Argentina (Vitzthum, 1941; Alzuet & Abrahamovich, 1989) – Roubikia imberba, Sennertia argentina, S. augustii, S. frontalis, S. longipilis, S. splendidulae Bahamas (Klimov & OConnor, 2008) - Sennertia pirata Acarorum Catalogus IX 289 Belize (Klimov & OConnor, 2008) – Sennertia recondita Bolivia (Klimov & OConnor, 2008) – Roubikia panamensis Brazil (Klimov & OConnor, 2008) – Centriacarus turbator, Roubikia panamensis, Sennertia argentina, S. donaldi, S. frontalis British Virgin Islands (Klimov & OConnor, 2008) - Sennertia pirata Chile (contin.) (Klimov & OConnor, 2007; Montalva & Allendes, 2010) – Chaetodactylus lassulus, Sennertia longipilis Colombia (Klimov & OConnor, 2008) – Centriacarus turbator Costa Rica (Klimov & OConnor, 2007, 2008) – Sennertia argentina, S. sayutara, S. sodalis Cuba (Klimov & OConnor, 2008) – Sennertia pirata Dominican Republic (Klimov & OConnor, 2008) – Chaetodactylus antillarum El Salvador (Klimov & OConnor, 2008) – Sennertia scutata French Guiana (Klimov & OConnor, 2008) – Roubikia panamensis, Sennertia surinamensis Guatemala (Baker & Delfinado-Baker, 1983; ) – Sennertia faini, S. scutata S. shimanukii Honduras (Klimov & OConnor, 2007) – Chaetodactylus melitomae, Sennertia frontalis Jamaica (Klimov & OConnor, 2008) – Chaetodactylus antillarum Nicaragua (Klimov & OConnor, 2008) – Sennertia argentina Panama (Baker, Roubik & Delfinado-Baker, 1987; Klimov & OConnor, 2008) – Centriacarus turbator, Roubikia panamensis, Sennertia argentyina, S. frontalis, S. sodalis, S. surinamensis Peru (Delfinado & Baker, 1976; Klimov & OConnor, 2007, 2008) – Roubikia latebrosa, Sennertia argentina, S. devincta, S. frontalis, S. ignota Suriname (Fain & Lukoschus, 1971) - Sennertia argentina, S. frontalis, S. surinamensis Trinidad (Turk, 1948) – Sennertia donaldi Uruguay (Klimov & OConnor, 2008) – Sennertia augustii Venezuela (Klimov & OConnor, 2007) – Centriacarus guahibo, Sennertia donaldi, S. frontalis Virgin Islands of the USA (Klimov & OConnor, 2008) - Sennertia pirata OCEANIA Australia (continent) (Womersley, 1941; Fain, 1982; Houston, 1992) – Sennertia queenslandica, S. alfkeni, S. leei Papua New Guinea (Canestrini, 1897) – Sennertia bifilis New Caledonia (Klimov & OConnor, 2008; Fain & Pauly, 2001) – Chaetodactylus ludwigi Micronesia (Klimov & OConnor, 2008) - Chaetodactylus ludwigi French Polynesia (Klimov & OConnor, 2008; Fain & Pauly, 2001) - Chaetodactylus ludwigi ? ANTARCTICA (Trägårdh, 1907) - Sennertia antarctica 290 Acarorum Catalogus IX Hosts of Chaetodactylidae Insecta Hymenoptera Fam. Apidae Subfam. Xylocopinae Ceratina aereola - Achaetodactylus decellei Ceratina apaca – Achaetodactylus ceratinae Ceratina cucurbitina – Sennertia tunisiana Ceratina diloloensis – Achaetodactylus leleupi Ceratina excavata – Achaetodactylus decellei Ceratina exemia – Sennertia recondita Ceratina lativentris – Achaetodactylus ceratinae Ceratina nigriceps - Achaetodactylus ceratinae Ceratina penicilligera – Sennertia latipilis Ceratina ruwenzorica - Achaetodactylus leleupi Ceratina spilota - Achaetodactylus leleupi, A. decellei Ceratina turneri – Chaetodactylus dalyi, Sennertia ceratinarum Ceratina sp. – Sennertia benoiti Xylocopa aerata - Sennertia leei Xylocopa aestuans – Sennertia duweinii, S. egyptiaca, S. horrida Xylocopa amethystina – Sennertia delfinadoae (= bakeri), S. ratiocinator, S. robusta Xylocopa auripennis - Sennertia horrida Xylocopa b. barbatella – Sennertia loricata Xylocopa bhowara - Sennertia ratiocinator Xylocopa bombylans – Sennertia leei Xylocopa calens – Sennertia elseni Xylocopa californica - Sennertia lucrosa Xylocopa californica arizonensis - Sennertia segnis Xylocopa clarionensis - Sennertia lucrosa Xylocopa cubaecola – Sennertia pirata Xylocopa dejeanii - Sennertia ratiocinator Xylocopa fallax - Sennertia horrida Xylocopa fimbriata - Sennertia argentina, S. donaldi Xylocopa frontalis – Sennertia argentina, S. donaldi Xylocopa guatemalensis - Sennertia lucrosa Xylocopa imitator - Sennertia tanythrix Xylocopa inconstans – Sennertia leclercqi Xylocopa iridipennis - Sennertia horrida Acarorum Catalogus IX 291 Xylocopa latipes - Sennertia hipposideros Xylocopa lepeletieri - Sennertia jeanalexi Xylocopa mcgregori - Sennertia horrida Xylocopa mexicanorum – Sennertia argentina, S. lucrosa Xylocopa micans – Sennertia loricata Xylocopa mimetica - Sennertia horrida Xylocopa mirabilis - Sennertia madagascarensis Xylocopa mordax – Sennertia loricata, S. pirata Xylocopa muscaria – Sennertia loricata Xylocopa nasalis – Sennertia dissimilis, S. horrida, S. oudemansi Xylocopa nautlana – Sennertia argentina Xylocopa nigrita - Sennertia jeanalexi, S. perturbans, S. simplex, S. tanythrix Xylocopa nitidiventris - Sennertia cerambycina Xylocopa olivieri - Sennertia zhelochovtsevi Xylocopa przewalskyi - Sennertia cerambycina Xylocopa punctilabris - Sennertia gargantua Xylocopa rufa - Sennertia zhelochovtsevi Xylocopa scioensis (= Mesotrichia bryorum) – Sennertia queenslandica Xylocopa sonorina - Sennertia shimanukii Xylocopa splendidula splendidula Xylocopa tenuiscapa - Sennertia greeni, S. hipposideros, S. koptorthosomae, S. vanderhammeni Xylocopa torrida - Sennertia tanythrix Xylocopa tranquebarorum – Sennertia potanini Xylocopa turanica - Sennertia cerambycina Xylocopa valga – Sennertia cerambycina, S. gargantua Xylocopa varentzowi – Sennertia cerambycina Xylocopa varipuncta - Sennertia lucrosa Xylocopa violacea – Sennertia cerambycina Subfam. Apinae Apis mellifera – Chaetodactylus claviger, Sennertia faini Melitoma segmentaria – Chaetodactylus melitomae Tetrapedia diversipes – Roubikia panamensis T. maura - Roubikia officiosa T. peckoltii - Roubikia panamensis Tetrapedia sp. – Roubikia imberba, R. latebrosa Fam. Megachilidae Chelostoma florisomne – Chaetodactylus birulai Hoplitis pilosifrons - Chaetodactylus hopliti H. producta - Chaetodactylus hopliti 292 Acarorum Catalogus IX Lithurgus antilleorum – Chaetodactylus antillarum L. apicalis – Chaetodactylus kouboy, Ch. lithurgi L. atratus – Chaetodactylus ludwigi L. echinocacti – Chaetodactylus abditus L. gibbosus – Chaetodactylus gibbosi, Ch. lithurgi L. littoralis - Chaetodactylus lithurgi L. planifrons – Chaetodactylus abditus L. scabrosus - Chaetodactylus ludwigi Megachile bombycina – Chaetodactylus dementjevi M. liguseca - Chaetodactylus dementjevi Megachile sp. – Chaetodactylus osmiae Osmia aurulenta – Chaetodactylus chrysidis O. azteca – Chaetodactylus azteca O. bicolor – Chaetodactylus chrysidis O. bicornis – Chaetodactylus osmiae O. brevicornis – Chaetodactylus reaumuri O. brevis - Chaetodactylus hopliti O. bucephala - Chaetodactylus hopliti O. cornifrons – Chaetodactylus hirashimai, Ch. krombeini, Ch. nipponicus O. cornuta – Chaetodactylus osmiae, Ch. zachvatkini O. excavata - Chaetodactylus hirashimai, Ch. nipponicus O. fronticornis – Chaetodactylus osmiae O. georgica – Chaetodactylus rozeni O. imaii - Chaetodactylus hirashimai O. inermis - Chaetodactylus hopliti O. kohlii – Chaetodactylus osmiae O. latreillei – Chaetodactylus zachvatkini O. leucogastra – Chaetodactylus reaumuri O. niveata - Chaetodactylus osmiae, Ch. zachvatkini O. pedicornis - Chaetodactylus hirashimai, Ch. nipponicus O. rubriflorus – Chaetodactylus claudus O. rufa – Chaetodactylus osmiae O. rufiventris – Chaetodactylus osmiae, Ch. reaumuri O. subaustralis – Chaetodactylus micheneri O. taurus - Chaetodactylus nipponicus O. tricornis - Chaetodactylus claviger, Ch. osmiae, Ch. zachvatkini Rhodanthidium sticticum – Chaetodactylus anthidii Trichorhurgus dubius - Chaetodactylus lassulus Fam. 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Abteilung: Arachnoidea. 5. Buch. Acarina. 1011. Leipzig: Akademische Verlagsgesellschaft, Becker & Erler Kom.-Ges. White J.B., Y.L. Park, T.P. West, P.C. Tobin. 2009. Assessment of potential fumigants to control Chaetodactylus krombeini (Acari: Chaetodactylidae) associated with Osmia cornifrons (Hymenoptera: Megachilidae). – J. Econ. Entomol., 102(6): 2090–2095. Womersley H. 1941. Studies in Australian Acarina. (2) Tyroglyphidae (s. l.). – Records of the South Australian Museum, 6: 451–488. Acarorum Catalogus IX 303 Yamada M. 1990. Control of Chaetodactylus mite, Chaetodactylus nipponicus Kurosa, an important mortality agent of hornfaced bee, Osmia cornifrons Radoszkowski. - Bulletin of the Aomori Apple Experiment Station, 26: 39–77. Walter D. E., J.J. Beard, K.L. Walker, K. Sparks. 2002. Of mites and bees: A review of mite–bee associations in Australia and a revision of Raymentia. Yamada M., N. Oyama, N. Sekita, S. Shirasaki, C. Tsugawa. 1971. Preservation and utilization of natural enemies and useful insects in apple orchards III. The ecology of the megachilid bee, Osmia cornifrons (Radoszkowski) (Hym.: Apidae) and its utilization for apple pollination. - Bulletin of the Aomori Apple Experiment Station, 15: 1–721 (In Japanese with English Summary). Zachvatkin A. A. 1941. Tyroglyphoid mites Tyroglyphoidea. Fauna SSSR: Paukoobraznuye, 6(1). Akademiya Nauk SSSR: Moscow-Leningrad. 475 pp. (in Russian, French abstract). Zachvatkin A. A. 1953. Investigations on the morphology and postembryonic development of tyroglyphids (Sarcoptiformes, Tyroglyphoidea). In: V. B. Dubinin and A. B. Lange (eds). Zachvatkin A. A.: Sbornik Nauchnykh rabot. Zachvatkin A.A. 1953. Collection of Scientific Works. Moscow State University, Moscow: 19–120. (In Russian). The Mite Family Hyadesiidae Hyadesiidae Halbert, 1915: 53 The mites of the genera Hyadesia and Amhyadesia (total of 47 species, see Proches and Marshall, 2001), inhabit marine intertidal to littoral habitats, occurring on all coastlines and often associated with barnacles, algal mats. The Hyadesiids feed primarily on marine algae, but other organic debris may also be consumed. (Pugh & King, 1985a). Deutonymphs are never formed in the family, and dispersal is accomplished by feeding forms. The start of the studies of Hyadesiidae belongs to Mégnin (1891), who described the genus Hyadesia, to Michael (1893) and to Halbert (1915) – the author of the family. Major taxonomic contributions are due also to Fain, Fain & Schuster, Fain & Ganning, Marshall & Nunkumar, Luxton, Lohmann, Marshall & Ugrasen, Viets, Womersley and others. Important biological data are due to Schuster, Pugh, Matshall and other researchers of coastal habitats. Composition of Hyadesiidae Family Hyadesiidae Halbert, 1915–2 gen., 47 sp. Amhyadesia Fain et Schuster, 1984–15 sp. Hyadesia Mégnin, 1891–32 sp. 304 Acarorum Catalogus IX Catalogue of Hyadesiidae Fam. Hyadesiidae Halbert Hyadesidae Halbert, 1915: 53 Hyadesiidae: Fain, 1981: 47 Lentungulidae Berlese, 1897 Type genus: Hyadesia Mégnin, 1889 Genus Amhyadesia Fain et Schuster Amhyadesia Fain & Schuster, 1984: 15 Type species: Amhyadesia californica Fain et Schuster, 1984 Amhyadesia atlantica Fain et Schuster Amhyadesia atlantica Fain & Schuster, 1983: 420 Type locality: Penhurst Park, Main Island Distribution: Bermudas Amhyadesia austrafricana Marshall et Nunkumar Amhyadesia austrafricana Marshall & Nunkumar, 1999: 173 Type locality: Sardinia Bay near Port Elizabeth Distribution: South Africa Amhyadesia bartschae Luxton Amhyadesia bartschae Luxton, 1992: 229 Type locality: Hong Kong Distribution: China Amhyadesia bermudana Fain et Schuster Amhyadesia bermudana Fain & Schuster, 1983: 418 Type locality: Crawl Hill, Main Island Distribution: Bermudas Amhyadesia brasiliensis Fain et Schuster Amhyadesia brasiliensis Fain & Schuster, 1984b: 274 Type locality: south of São Sebastião (São Paulo State) Distribution: Brazil Amhyadesia bursaria Fain et Schuster Amhyadesia bursaria Fain & Schuster, 1984a: 106 Type locality: Noumea, between Baie des Citrons and Anse Vata Acarorum Catalogus IX 305 Distribution: New Caledonia Amhyadesia californica Fain et Ganning Amhyadesia californica Fain & Ganning, 1979: 139 Type locality: California Distribution: USA Amhyadesia costaricensis Fain et Schuster Amhyadesia costaricensis Fain & Schuster, 1984: 271 Type locality: Pacific coast, northern part of the Nat. park “Manuel Antonio”, Quepos Distribution: Costa Rica Amhyadesia elizabethensis Marshall et Nunkumar Amhyadesia elizabethensis Marshall & Nunkumar, 1999: 176 Type locality: Sardinia Bay near Port Elizabeth Distribution: South Africa Amhyadesia heteromorpha Luxton Amhyadesia heteromorpha Luxton, 1992a: 232 Type locality: Hong Kong Distribution: China Amhyadesia heterophallus Fain et Schuster Amhyadesia heterophallus Fain & Schuster, 1984a: 103; Marshall & Nunkumar, 1999: 177 Type locality: Villingili Island, Male Atoll Distribution: Maledives, Philippines, Mozambique, South Africa Amhyadesia longipillis Fain et Schuster Amhyadesia longipillis Fain & Schuster, 1984a: 109 Type locality: Pang-an Island Distribution: Philippines Amhyadesia madeirensis Fain et Schuster Amhyadesia madeirensis Fain & Schuster, 1986: 4 Type locality: East of Caniçal Distribution: Madeira (incl. Porto Santo) Amhyadesia pacifica Fain et Schuster Amhyadesia pacifica Fain & Schuster, 1984a: 109 Type locality: Coconut Island, Camotes Sea Distribution: Philippines 306 Acarorum Catalogus IX Amhyadesia punctulata Luxton Amhyadesia punctulata Luxton, 1989a: 73; 1990: 374 Type locality: Auckland Distribution: New Zealand Genus Hyadesia Mégnin Hyadesia Mégnin, 1891: 53 Lentungula Michael, 1893: 262 Type species: Hyadesia uncinifer Mégnin, 1891 Subgenus Hyadesia Mégnin Hyadesia Mégnin, 1891: 53; made subgenus by Fain, 1981 Parahyadesia (unavailable name? No type species fixation) Type species: Hyadesia uncinifer Mégnin, 1891 Hyadesia (H.) agulhensis Marshall et Ugrasen Hyadesia (Parahyadesia) agulhensis Marshall & Ugrasen, 2000: 88 Type locality: South Africa Distribution: South Africa Hyadesia (H.) alexi Marshall et Chown Hyadesia alexi Marshall & Chown, 2003: 120 Type locality: Gough Island Distribution: Gough Island (nr. Tristan da Cunha) Hyadesia (H.) algivorans (Michael) Lentungula algivorans Michael, 1893: 262 Hyadesia algivorans (Michael): Turk, 1953: 82; Fain, 1981: 55 Type locality: Cornwall, Land’s End Distribution: Great Britain, Hyadesia (H.) arabica Fain et Schuster Hyadesia (H.) arabica Fain & Schuster, 1985: 68 Type locality: Red Sea at Al Ghardaqa Distribution: Egypt Hyadesia (H.) australiana Fain et Synnot Hyadesia australiana Fain & Synnot, 1981: 195 Type locality: Gunnamatta Beach, Mornington Peninsula, nr. Melbourne, Victoria Distribution: Australia (Victoria) Acarorum Catalogus IX 307 Hyadesia (H.) benguelensis Marshall et Ugrasen Hyadesia (Parahyadesia) benguelensis Marshall & Ugrasen, 2000 Type locality: Angola Distribution: Angola, South Africa Hyadesia (H.) curassaviensis Viets Hyadesia curassaviensis Viets, 1936: 426; Fain, 1981: 53 Type locality: Curaçao, Riftwal nr. Willemstad Distribution: Curaçao Hyadesia (H.) fusca (Lohmann) Lentungula fusca Lohmann, 1894: 85; Canestrini & Kramer, 1899: 136; Halbert, 1915: 108 Hyadesia fusca (Lohmann): Halbert, 1920: 137; Willmann, 1937: 167; Sellnick, 1949: 127; Turk, 1953: 82 Hyadesia (H.) fusca (Lohmann): Fain & Ganning, 1989: 163 Hyadesia furcillipes Benard, 1961: 81. Type: France (syn. by Fain, 1981) Type locality: Helgoland Island (Germany) Distribution: Belgium, France, Germany, Great Britain, Finland, Holland, Ireland, Norway, Sweden, USA Hyadesia (H.) glynni Manson Hyadesia glynni Manson, 1963: 163 Type locality: Hopkins Marine Station, Pacific Grove, California Distribution: USA (California) Hyadesia (H.) halophila Fain Hyadesia (Hyadesia) halophila Fain, 1974: 699 Type locality: Kerguelen Island Distribution: Kerguelen Island Hyadesia (H.) heteromorpha Marshall et Ugrasen Hyadesia (Parahyadesia) heteromorpha Marshall & Ugrasen, 2000: 87 Type locality: South Africa Distribution: South Africa Hyadesia (H.) maxima Fain, Sømme et Block Hyadesia maxima Fain, Sømme & Block, 1983: 171 Type locality: South Georgia, Maiviken and Sooty Bluff Distribution: South Georgia 308 Acarorum Catalogus IX Hyadesia (H.) microseta Luxton Hyadesia (H.) microseta Luxton, 1989a: 81; 1990: 376 Type locality: Auckland Distribution: New Zealand Hyadesia (H.) mollis Luxton Hyadesia (Parahyadesia) mollis Luxton, 1989a: 81; 1990: 378 Type locality: Auckland Distribution: New Zealand (Auckland) Hyadesia (H.) nearctica Fain et Ganning Hyadesia (H.) nearctica Fain & Ganning, 1978: 215 Type locality: from Saint John, New Brunswick, Canada throughout the coast of Main, New Hampshire to Cape Ann, Massachusetts, USA Distribution: Canada, USA Hyadesia (H.) pakistanensis Fain et Schuster Hyadesia pakistanensis Fain & Schuster, 1985: 73 Type locality: Paradise Point near Karachi Distribution: Pakistan Hyadesia (H.) paulensis Fain Hyadesia (H.) paulensis Fain, 1975b: 157 Type locality: St. Paul Is. Distribution: St. Paul Is. Hyadesia (H.) plicata Luxton Hyadesia (Parahyadesia) plicata Luxton, 1989a: 84; 1990: 378 Type locality: Chatham Islands Distribution: New Zealand (Chatham Islands) Hyadesia (H.) reticulata Luxton Hyadesia (Parahyadesia) reticulata Luxton, 1989a: 88; 1990: 378 Type locality: Kaikoura Distribution: New Zealand Hyadesia (H.) sanjuanensis Fain et Ganning Hyadesia (H.) sanjuanensis Fain & Ganning, 1989: 162 Type locality: San Juan Archip., Friday Harbor Marine Lab. Acarorum Catalogus IX 309 Distribution: USA (State of Washington) Hyadesia (H.) sellai Viets Hyadesia sellai Viets, 1937: 3; Fain, 1981: 54 Type locality: Rovinj, Split Distribution: Croatia Hyadesia (H.) subantarctica Fain Hyadesia (Hyadesia) subantarctica Fain, 1974: 702 Type locality: Kerguelen Island Distribution: Kerguelen Island Hyadesia (H.) tenuipilis Fain Hyadesia fusca tenuipilis Fain, 1979: 131 Type locality: Abo Distribution: Finland Hyadesia (H.) tesselata Luxton Hyadesia (Parahyadesia) tesselata Luxton, 1989a: 91; 1990: 378 Type locality: Dunedin Distribution: New Zealand Hyadesia (H.) travei Fain Hyadesia (H.) travei Fain, 1975b: 153 Type locality: Saint-Paul Isl. Distribution: Saint-Paul and New Amsterdam Islands Hyadesia (H.) tumida Benard Hyadesia tumida Benard, 1961: 71 Type locality: Roscoff, Bloscon (Brittany), Cotentin (Normandy) Distribution: France Hyadesia (H.) uncinifer Mégnin Hyadesia uncinifer Mégnin, 1889: 51 Type locality: Tierra del Fuego Distribution: Tierra del Fuego Hyadesia (H.) verrucosa Fain et Schuster Hyadesia verrucosa Fain & Schuster, 1985: 70 Type locality: Rovinj Distribution: Croatia 310 Acarorum Catalogus IX Hyadesia (H.) vietsi Womersley Hyadesia vietsi Womersley, 1961: 204; Fain, 1979: 127 Type locality: Biak Is. Distribution: Indonesia (Papua) Hyadesia (H.) womersleyi Luxton Hyadesia womersleyi Luxton, 1989b: 407 Type locality: Macquarie Island Distribution: Macquarie Island Hyadesia (H.) zelandica Luxton Hyadesia zelandica Luxton, 1984: 293; 1990: 376 Type locality: Auckland Distribution: New Zealand, Auckland Subgenus Hyadesiella Fain Hyadesiella Fain, 1974: 693 Type species: Hyadesia kerguelensis Lohmann, 1907 Hyadesia (Hyadesiella) kerguelensis Lohmann Hyadesia kerguelensis Lohmann, 1907: 369 Hyadesia (Hyadesiella) kerguelensis Lohmann: Fain, 1974: 693 Type locality: Kerguelen Island Distribution: Kerguelen Island Index of Hyadesiidae (Names in bold are accepted as valid) agulhensis, Hyadesia 306 alexi, Hyadesia 306 algivorans, Hyadesia 306 algivorans, Lentungula 306 Amhyadesia 304 arabica, Hyadesia 306 atlantica, Amhyadesia 304 austrafricana, Amhyadesia 304 australiana, Hyadesia 306 bartschae, Amhyadesia 304 benguelensis, Hyadesia 307 bermudana, Amhyadesia 304 brasiliensis, Amhyadesia 304 bursaria, Amhyadesiа 304 californica, Amhyadesia 305 costaricensis, Amhyadesia 305 curassаviensis, Hyadesia 307 elizabethensis, Amhyadesia 305 furcillipes, Hyadesia 307 fusca, Hyadesia 307 fusca, Lentungula 307 glynni, Hyadesia 307 halophila, Hyadesia 307 heteromorpha, Amhyadesia 305 Acarorum Catalogus IX 311 heteromorpha, Hyadesia 307 heterophallus, Amhyadesia 305 Hyadesia 306 Hyadesiella 310 Hyadesiidae 304 Hyadesidae 304 kerguelensis, Hyadesia 310 Lentungula 306 Lentungulidae 304 longipilis, Amhyadesia 305 madeirensis, Amhyadesia 305 maxima, Hyadesia 307 microseta, Hyadesia 308 mollis, Hyadesia 308 nearctica, Hyadesia 308 pacifica, Amhyadesia 305 pakistanensis, Hyadesia 308 Parahyadesia 306 paulensis, Hyadesia 308 plicata, Hyadesia 308 punctulata, Amhyadesia 306 reticulata, Hyadesia 308 sanjuanensis, Hyadesia 308 sellai, Hyadesia 309 subantarctica, Hyadesia 309 tenuipilis, Hyadesia 309 tenuipilis, Hyadesia fusca 309 tesselata, Hyadesia 309 travei, Hyadesia 309 tumida, Hyadesia 309 uncinifer, Hyadesia 309 verrucosa, Hyadesia 309 vietsi, Hyadesia 309 womersleyi, Hyadesia 310 zelandica, Hyadesia 310 Distribution of Hyadesiidae EUROPE Belgium (Fain, 1979) – Hyadesia fusca Croatia (Viets, 1937; Fain & Schuster, 1985) – Hyadesia (H.) sellai, H. (H.) verrucosa Finland (Fain, 1979) – Hyadesia fusca, H. tenuipilis France (contin.) (Benard, 1961) – Hyadesia fusca (= furcillipes), H. tumida Germany (Lohmann, 1894; Willmann, 1937) – Hyadesia fusca Great Britain (Michael, 1893; Turk, 1953) – Hyadesia algivorans, H. fusca Holland (Oudemans, 1927; Pugh & King, 1985) – Hyadesia fusca Ireland (Halbert, 1920) – Hyadesia fusca Madeira (Fain & Schuster, 1986) – Amhyadesia madeirensis, Hyadesia (H.) verrucosa Sweden (Sellnick, 1949; Ganning, 1970) – Hyadesia fusca AFRICA Angola (Marshall & Ugrasen, 2000) - Hyadesia benguelensis Egypt (Fain & Schuster, 1985) – Hyadesia (H.) arabica South Africa (Marshall & Nunkumar, 1999; Marshall & Ugrasen, 2000) – Amhyadesia austrafricana, A. elizabethensis, Hyadesia agulhensis, H. benguelensis ASIA China (Luxton, 1992a) – Amhyadesia bartschae, A. heteromorpha [Hong Kong] 312 Acarorum Catalogus IX Indonesia (Womersley, 1961) – Hyadesia vietsi [Papua, Biak Isl.] Maledive Islands (Fain & Schuster, 1984a) - Amhyadesia heterophallus Pakistan (Fain & Schuster, 1985) – Hyadesia (H.) pakistanensis Philippines (Talker, Alberti & Sotto, 1981; Fain & Schuster, 1984a) - Amhyadesia heterophallus, A. longipilis, A. pacifica NORTH AMERICA Bermuda Islands (Fain & Ganning, 1989; Fain & Schuster, 1983) – Amhyadesia atlantica, A. bermudana Canada (Fain & Ganning, 1978, 1979, 1989) – Hyadesia nearctica USA (Manson, 1963; Fain & Ganning, 1979, 1989) – Amhyadesia californica, A. glynni, Hyadesia fusca, H. sanjuanensis, H. nearctica SOUTH and CENTRAL AMERICA Argentina (Megnin, 1891) – Hyadesia uncinifer (Tierra del Fuego) Brazil (Fain & Schuster, 1984b) – Amhyadesia brasiliensis Costa Rica (Fain & Schuster, 1984b) – Amhyadesia costaricensis Curaçao (Viets, 1936) – Hyadesia curassaviensis ATLANTIC OCEAN Gough Island (Marshall & Chown, 2003) – Hyadesia alexi Saint Paul and New Amsterdam Islands (Fain, 1975a, b) – Hyadesia paulensis, H. travei OCEANIA Australia (continent) (Fain & Synnot, 1981) – Hyadesia australiana New Zealand (Luxton, 1984, 1989, 1990) – Amhyadesia punctulata, Hyadesia microseta, H. mollis, H. reticulata, H. tesselata, H. zelandica New Caledonia (Fain & Schuster, 1984a) - Amhyadesia bursaria ANTARCTICA South Georgia (Fain, Sømme & Block, 1983) – Hyadesia maxima Kerguelen Island (Lohmann, 1907; Fain, 1974) – Hyadesia halophila, H. subantarctica, H. (Hyadesialla) kerguelensis Bibliography of Hyadesiidae André M. 1931. Sur le genre Hyadesia Megnin, 1889. – Bull. Mus. Hist. Nat. Paris, (2) 3: 496–506. André M. 1933. Contribution à l’étude des acariens marins des Iles Kerguelen et Saint-Paul. – Ann. Inst. Océanographique, N. Sér. 13: 137–161. Acarorum Catalogus IX 313 André M. 1947. Croisière du Bougainville aux îles australes françaises. Acariens. – Mém. Mus. Hist. Nat. Paris, Sér. 2(20) : 65–100. Benard F. 1961. Sur deux nouvelles espèces d’Acariens marins Hyadesia tumida et Hyadesia furcillipes : sous-ordre des Sarcoptiformes, famille des Hyadesidae. – Cahiers de Biologie Marine, 2: 71–96. Berlese A. 1892–1897. Acari, Myriopoda et Scorpiones hucusque in Italia reperta. Ordo Cryptostigmata. Portici : 99–100. Fain A. 1974a. Acariens récoltés par le Dr. J. Travé aux îles subantarctiques I. Familles Saproglyphidae et Hyadesidae (Astigmates). – Acarologia, 16(4): 684–708. Fain A. 1974b. Notes on the Hyadesia species (Acari, Astigmata, Hyadesidae) of the Oudemans Collection, with a redescription of Hyadesia vietsi Womersley, 1961. – Zool. Meded., 54(10): 127–131. Fain A. 1975a. Acariens récoltés par le Dr. J. Travé aux îles subantarctiques. I. Familles Saproglyphidae et Hyadesidae (Astigmates). – Acarologia, 16(4): 684–708. Fain A. 1975b. Deux nouvelles espèces du genre Hyadesia récoltées par le Dr. Travé aux îles SaintPaul et Nouvelle-Amsterdam (Astigmates: Hyadesiidae). – Acarologia, 17(1): 153–159. Fain A. 1979. Notes on the Hyadesia species (Acari, Astigmata, Hyadesiidae) of the Oudemans Collection, with a redescription of Hyadesia vietsi Womersley,1961. – Zool. Mededelingen, Leiden, 54(10): 127–131. Fain A. 1981. Notes on the Hyadesiidae Halbert, 1915 and Algophagidae Fain, 1974, nov. tax. (Acari, Astigmata) with a redescription of Hyadesia curassaviensis Viets, 1936 and H. sellai Viets, 1937. – Acarologia, 22(1): 47–61. Fain A., B. Ganning. 1978. Hyadesia (Hyadesia) nearctica sp.n. (Acari, Astigmata, Hyadesidae) from the East Coast of U.S.A. – Acta Zoologica et Pathologica Antwerpiensia, 73: 215–219. Fain A., B. Ganning. 1979. Amhyadesia californica g.n., sp.n. (Acari, Hyadesiidae) from the Southern Californian Coast. – Int. J. Acarol., 5: 139–142. Fain A., B. Ganning. 1989. Notes on hyadesiid mites (Acari, Hyadesiidae) from North America with description of a new species. - Bull. Inst. r. Sci. nat. Belg., 59: 161–164. Fain A., R. Schuster. 1983. New astigmatic mites from the coastal area of Bermuda Islands (Acari: Hyadesiidae, Saproglyphidae, Acaridae). – Entomol. Mitt. Zool. Mus. Hamburg, 7(119): 417–434. Fain A., R. Schuster. 1984a. Four new species of the genus Amhyadesia Fain & Ganning, 1979 from the intertidal zone of several Indo-Pacific islands (Acari: Hyadesidae). - International Journal of Acarology, 10(2): 103–111. Fain A., R. Schuster. 1984b. Two new species of the family Hyadesiidae (Acari, Astigmata) from Costa-Rican and Brasilian Coasts. – Acarologia, 25: 271–279. Fain A., R. Schuster. 1985. New observations on the Hyadesiidae (Acari, Asigmata) description of three new species of the genus Hyadesia Mégnin, 1891. – Acarologia, 26(1): 67–77. Fain A., R. Schuster. 1986. First record of the mite family Hyadesiidae (Acari, Astigmata) from the Madeira Archipelago. – Bocagiana, Madeira, 99: 1–10. Fain A., L. Sømme, W. Block. 1983. Hyadesia maxima sp. n. (Acari, Hyadesiidae) from South Georgia. – Bull. Ann. Soc. r. belge Ent., 119: 171–176. 314 Acarorum Catalogus IX Fain A., R. Synnot. 1981. Hyadesia australiana sp.n. (Astigmata, Hyadesiidae) from southeastern Australia. - Bulletin et Annales de la Societe royal Belgique (Entomologie), 117(7–9): 195–201. Ganning B. 1970. Population Dynamics and Salinity Tolerance of Hyadesia fusca (Lohmann) (Acarina, Sarcoptiformes) from Brakish Water Rockpools, with notes on the Microenvironment inside Enteromorpha Tubes. – Oecologia (Berl.), 5: 127–137. Halbert J.N. 1915. Clare Island Survey. Part 39 Acarinida. Section II. Terrestrial and marine Acarina. – Proc. R. Irish Acad., 39(2): 45–136. Halbert J.N. 1920. The Acarina of the seashore. – Proc. R. Irish Acad., 35B(7): 106–152. Hughes A.M., B.J.A. Goodman. 1969. Neohyadesia signyi (Hyadesidae: Acarina). A new genus and species from Signy Island, South Orkney Islands. – Bull. Antarct. Surv. Bull., 22: 39–48. Lohmann H. 1894. Lentungula fusca nov. spec., eine marine Sarcoptide. – Wissenschaftliche Untersuchungen, 1: 84–90. Lohmann H. 1906. Die Meeresmilben der Deutschen Sudpolar-Expedition 1901–1903. Berlin 9, Zool., 1(5): 361–413. Lohmann H. 1907. Ueber einige faunistische Ergebnisse der Deutschen Südpolar-Expedition. Besonderer Berucks der Meeresmilben. – Schrift. Naturw. Vereins Schleswig-Holstein, 14(1): 1–14. Lohmann H. 1908. Die Meeresmilben der Deutschen Südpolar-Expedition 1901–1903. Deutsche Südpolar-Expedition 1901–1903. Berlin, 9. Zool. 1(5): 361–413. Luxton M. 1984. More marine littoral mites (Acari) from New Zealand. - New Zealand Journal of Marine and Freshwater Research, 18(3): 291–302. Luxton M. 1989a. Mites of the family Hyadesiidae (Acari: Astigmata) from New Zealand. Zoological Journal of the Linnean Society, 95(1): 71–95. Luxton M. 1989b. New taxa of intertidal mites (Acari). - Journal of Natural History, 23(2): 407–428. Luxton M. 1990. The marine littoral mites of the New Zealand region. – J. of the Royal Society of New Zealand, 20(4): 367–418. Luxton M. 1992. Hong Kong hyadesiid mites (Acari: Astigmata). – Pp. 229–236. In: The Marine Flora and Fauna of Hong Kong and southern China, 3(1): 3–33. Manson D.C.M. 1963. A new species of Hyadesia (Acarina: Carpoglyphidae). – Proc. Entomol. Soc. Washington, 65: 163–167. Marshall D.J., S.L. Chown. 2003. Marine hyadesiid mites from Gough Island. – Hydrobiologia, 495: 119–126. Marshall D.J., Sh. Nunkumar. 1999. First records and descriptions of the marine littoral mite genus Amhyadesia Fain and Ganning 1979 (Acari: Astigmata: Hyadesiidae) from southern Africa. – South African J. of Zoology, 34(4): 173–179. Marshall D.J., K. Ugrasen. 2000. Three new species of the marine littoral mite Hyadesia (Parahyadesia) (Astigmata: Hyadesiidae) from southern Africa. – African Zoology, 35(1): 85–92. Mégnin P. 1889. Note sur un Acarien de la Terre de Feu (Hyadesia uncinifer n.g., n.sp.). - Mission scientifique du Cap Horn, 1882–1883 (Zool.), 6(3): L. 51 – L.53. Acarorum Catalogus IX 315 Mégnin P. 1891. Note sur un Acarien de la Terre de Feu (Hyadesia uncinifer n.g., n.sp.). – Mission Scient. Cap Horn 1882–1883, Paris, 6(2): 51–53. Michael A.D. 1893. On a new genus species of Acari found in Cornwall. – Proc. Zool. Soc. London: 262–267. Michael A.D. 1901. British Tyroglyphidae, 1: 1–291. Oudemans A.C. 1927. [Mededeling over Acari]. – Tijdschr. Ent., 70: lxx-lxxvi. Proches S., D.J. Marshall. 2001. Global Patterns of Non-Halacarid Marine Intertidal Mites: Implications for Their Origins in Marine Habitats. - Journal of Biogeography, 28(1): 47–58. Pugh P. J. A. 1993. A synonymic catalogue of the Acari from Antarctica, the subAntarctic Islands and the Southern Ocean. - Journal of Natural History, 27: 323–421. Pugh P.J.A., P.E. King. 1985. Feeding in intertidal Acari. – Journal of Experimental Marine Biology and Ecology, 94: 269–280. Schuster R. 1962. Das marine Litoral als Lebensraum terrestrischer Kleinarthropoden. – Int. Revue ges. Hydrobiol., 47(3): 359–412. Schuster R. 1979. Soil mites in the marine environment. – Recent Advances in Acarology, 1: 593–602. Schuster R. 1988. Transoceanic distribution of air-breathing litoral mites. – Progress in Acarology, 1: 355–362. Sellnick M. 1949. Milben von der Küste Schwedens. – Entomologisk Tidskrift, 1949, 123–135. Talker H.E., G. Alberti, F.B. Sotto. 1981. Mites (Acari, Arachnida) in intertidal habitats off Cebu. – The Philippine Scientist, 18: 27–44. Viets K. 1936. Eine neue Hyadesia (Sarcoptiformes, Acari) von Curaçao. – Zool. Jahrb. Syst., 67: 425–428. Viets K. 1937. Eine neue Hyadesia-Art aus der Adria (Sarcoptiformes, Acari). – Not. Ist. Biol. Rovigno, 2(9): 1–10. Willmann C. 1937. Beitrag zur Kenntnis der Acarofauna der ostfriesischen Inseln. – Abh. Naturwiss. Verein zu Bremen, 30(1/2): 152–169. Womersley H. 1961. New species of Acarina from the Intertidal zone in Netherlands New Guinea. – Zool. Mededelingen, 37(12): 203–209. The Mite Family Algophagidae Fain Algophaginae Fain, 1974: 703 Algophagidae Fain, 1981: 49 The small family Algophagidae (eight genera, 17 species) was proposed by A. Fain, first as subfamily (1974), than as family (1981). The first genera (Hericia G. Canestrini, 1888, Fusohericia Vitzthum, 1931, Algophagus Hughes, 1955) and species have been described much earlier, but only with the intensifying of the Antarctic and subantarctic research many new taxa (mainly of Algophaginae) were discovered by Fain & Johnston (1975), 316 Acarorum Catalogus IX Hughes & Goodman (1969), Cruz & Socarras (1992 - Cuba), Fashing, OConnor & Kitching (2000), Marshall, OConnor & Pugh (2001, 2003), and others. These peculiar mites are aquatic, with various species inhabiting sap flows on trees (Hericia, Fusohericia), water filled tree holes {Lamingtonacarus), rivers and lakes (Algophagopsis), or other aquatic or subaquatic habitats (O’Connor, 2009) up to 100 m (Anderson & Krantz, 1993). Species of Algophagus and Neohyadesia are generally restricted to subantarctic islands. They range from coastal brackish pools to inland tussock grass soils, where they feed primarily on algae (Goddard 1982; Marshall, OConnor & Pugh, 2003, generalized by O’Connor, 2009). Genital papillae are strongly reduced in Algophagidae, but the uniquely developed axillary organs between legs I—II characterize the family. The sap inhabiting taxa of the algophagid subfamily Hericiinae form deutonymphs for dispersal on sap frequenting Coleoptera (Nitidulidae, Nosodendridae) and Lepidoptera (O’Connor, 2009). The algophagine genus Lamingtonacarus oreillyorum is known to form deutonymphs, but these have not been collected from any insect host. OConnor and Moser (1985) proposed generic phylogenetic analysis. Composition of Algophagidae Fam. Algophagidae Fain, 1981–9 gen., 24 sp. Subfam. Algophaginae Fain, 1974–14 sp. Algophagopsis Fain et Johnston, 1975–1 sp. Algophagus Hughes, 1955–6 sp. Lamingtonacarus Fashing, OConnor et Kitching, 2000–2 sp. Neohyadesia Hughes et Goodman, 1969–3 sp. Terraphagus Clark et Andrews, 2012–1 sp. Thalassophagacarus J. de la Cruz et Socarras, 1992–1 sp. Subfam. Hericiinae OConnor et Moser, 1985–10 sp. Fusohericia Vitzthum, 1931–3 sp. Hericia G. Canestrini, 1888–6 sp. Prohericia OConnor et Moser, 1985–1 sp. Catalogue of Algophagidae Fam. Algophagidae Fain Algophaginae Fain, 1974: 703 Algophagidae Fain: Fain, 1981: 49 Type genus: Algophagus Hughes, 1955 Subfam. Algophaginae Fain Algophaginae Fain, 1974: 703 Acarorum Catalogus IX 317 Type genus: Algophagus Hughes, 1955 Genus Algophagopsis Fain et Johnston Algophagopsis Fain & Johnston, 1975: 67 Type species: Algophagopsis pneumatica Fain et Johnston, 1975 Algophagopsis pneumatica Fain et Johnston Algophagopsis pneumatica Fain & Johnston, 1975: 67 Type locality: Kings River, California Distribution: USA (California) Genus Algophagus Hughes Algophagus Hughes, 1955: 1 Type species: Algophagus antarcticus Hughes, 1955 Algophagus antarcticus Hughes Algophagus antarcticus Hughes, 1955: 1; OConnor & Pugh, 2003: 39 Type locality: Heard Island Distribution: Heard Island, Kerguelen Island Algophagus brachytarsus OConnor et Pugh Algophagus brachytarsus OConnor & Pugh, 2003: 41 Type locality: Prince Edward Islands Distribution: Prince Edward Islands Algophagus laticollaris Fain Algophagus antarcticus laticollaris Fain, 1974: 704 Algophagus laticollaris Fain: OConnor & Pugh, 2003: 39 Type locality: Kerguelen I. Distribution: Kerguelen I. Algophagus macrolithus OConnor et Pugh Algophagus macrolithus OConnor & Pugh, 2003: 43 Type locality: Prince Edward Islands Distribution: Prince Edward Islands Algophagus pennsylvanicus Fashing et Wiseman Algophagus pennsylvanicus Fashing & Wiseman, 1980: 79 Type locality: Cook Forest State Park, Forest County, Pennsylvania Distribution: USA 318 Acarorum Catalogus IX Algophagus semicollaris Fain Algophagus semicollaris Fain, 1974: 704; Fedorova, Fedorov, Stavsky, Matyukhina & Voronin 1980: 82 Type locality: Kerguelen Island Distribution: Kerguelen Island Genus Lamingtonacarus Fashing, OConnor et Kitching Lamingtonacarus Fashing, OConnor & Kitching, 2000: 592 Type species: Lamingtonacarus oreillyorum Fashing, OConnor et Kitching, 2000 Lamingtonacarus oreillyorum Fashing, OConnor et Kitching Lamingtonacarus oreillyorum Fashing, OConnor & Kitching, 2000: 592 Type locality: Lamington National Park, Box Forest; Queensland Distribution: Australia (Queensland) Lamingtonacarus posidonis Fashing, OConnor et Kitching Lamingtonacarus posidonis Fashing, OConnor & Kitching, 2000: 599 Type locality: Lamington National Park, Box Forest; Queensland Distribution: Australia (Queensland) Genus Neohyadesia Hughes et Goodman Neohyadesia Hughes & Goodman, 1969: 39 Type species: Neohyadesia signyi Hughes et Goodman, 1969 Neohyadesia microtricha Marshall, OConnor et Pugh Neohyadesia microtricha Marshall, OConnor & Pugh, 2001: 101 Type locality: Marion Island (Prince Edward Islands, South Georgia, Trypot beach) Distribution: Indian Ocean islands Neohyadesia minor Clark et Andrews Neohyadesia minor Clark & Andrews, 2012: 1478 Type locality: Lake Forsyth/Wairewa, Canterbury Distribution: New Zealand Neohyadesia signyi Hughes et Goodman Neohyadesia signyi Hughes & Goodman, 1969: 39 Type locality: Signy Island Distribution: South Orkney Islands Neohyadesia signyi punctulata Fain Neohyadesia signyi punctulata Fain, 1974: 706 Acarorum Catalogus IX 319 Type locality: Kerguelen Island Distribution: Kerguelen Island Genus Terraphagus Clark et Andrews Terraphagus Clark & Andrews, 2012: 1476 Type species: Terraphagus antipodus Clark et Andrews, 2012 Terraphagus antipodus Clark et Andrews Terraphagus antipodus Clark & Andrews, 2012: 1476 Type locality: Antipodes Islands Distribution: Antipodes Islands Host: from grey petrel Procellaria cinerea burrow soil Genus Thalassophagacarus J. de la Cruz et Socarras Thalassophagacarus J. de la Cruz & Socarras, 1992: 1 Type species: Thalassophagacarus faime J. de la Cruz et Socarras, 1992 Thalassophagacarus faime J. de la Cruz et Socarras Thalassophagacarus faime J. de la Cruz & Socarras, 1992: 2 Type locality: “playa cercana al Faro Roncali, Cabo de San Antonio, Pinar del Rio” Distribution: Cuba Subfam. Hericiinae OConnor et Moser Hericiinae OConnor & Moser, 1985: 786 Type genus: Hericia G. Canestrini, 1888 Genus Fusohericia Vitzthum Fusohericia Vitzthum, 1931: 98 Type species: Fusohericia incredibilis Vitzthum, 1931 Fusohericia incredibilis Vitzthum Fusohericia incredibilis Vitzthum, 1931: 98 Type locality: South Sumatra Distribution: Indonesia (Sumatra) Fusohericia heliconiae Fashing et Glist Fusohericia heliconiae Fashing & Glist, 2012: 591 Type locality: La Selva Biological Station and Reserve, Heredia Province Distribution: Costa Rica 320 Acarorum Catalogus IX Fusohericia lawrencei Baker et Crossley Fusohericia lawrencei Baker & Crossley, 1964: 4 Type locality: treehole, trunk of yellow poplar, Liriodendron tulipijera, Roane County, Tennessee Distribution: USA (Tennessee) Genus Hericia G.Canestrini Hericia G. Canestrini, 1888: 6 Type species: Glyciphagus hericius Robin, 1868 Hericia fermentationis Vitzthum Hericia fermentationis Vitzthum, 1931: 85 Type locality: Sumatra Distribution: Indonesia (Sumatra) Hericia georgei Michael Hericia georgei Michael, 1903: 38; Samšinák, 1972: 65; Haitlinger, 2002: 68; Fashing, 2008: 5 Type locality: Great Britain Distribution: Great Britain, Finland, France, Poland, Russia Hosts: Cetonia aurata, Potosia cuprea metallica (Coleoptera: Scarabaeidae), Catocala sponsa, C. nupta, C. adultera, C. elocata, C. neonympha, C. promissa, C. pacta, C. fraxini (Lepidoptera: Noctuidae), Cossus cossus (Lep.: Cossidae), Nymphalis antiopa, N. atalanta, Polygonia c-album, Acronicta [Apatele] psi, Europis occulta (Nymphalidae), Scoliopteryx libatrix (Erebidae) Hericia hericia (Robin) Glyciphagus hericius Robin, 1868: 607 Hericia robini Michael, 1903 (synonymy 1868–1903) Hericia hericia (Robin): Zachvatkin, 1941 Type locality: France Distribution: England, France, Germany, Italy Hericia janehenleyi Fashing Hericia janehenleyi Fashing, 2008: 7 Type species: Newport News, Eastern Virginia Distribution: USA Host: Glischrochilus obtusus (Coleoptera, Nitidulidae). Hericia paradoxa Türk et Türk Hericia (?) paradoxa Türk & Türk, 1957: 180 Type species: Erlangen Distribution: Germany Acarorum Catalogus IX 321 Hericia sanukiensis Fashing Hericia sanukiensis Fashing & Okabe, 2006: 3 Type locality: Takamatsu City, Kagawa Prefecture Distribution: Japan Host: Librodor japonicus (Coleoptera: Nitidulidae) Genus Prohericia OConnor et Moser Prohericia OConnor & Moser, 1985: 783 Type species: Prohericia longipes OConnor et Moser, 1985 Prohericia longipes OConnor et Moser Prohericia longipes OConnor & Moser, 1985: 783 Type locality: Germany Distribution: Germany Index of Algophagidae (Names in bold are accepted as valid) Algophagidae 316 Algophaginae 316 Algophagopsis 317 Algophagus 317 antarcticus, Algophagus 317 antipodus, Terraphagus 319 brachytarsus, Algophagus 317 faime, Thalassophagacarus 319 fermentationis, Hericia 320 Fusohericia 319 georgei, Hericia 320 heliconiae, Fusohericia 319 Hericia 320 hericia, Hericia 320 Hericiinae 319 hericius, Glyciphagus 320 incredibilis, Fusohericia 319 janehenleyi, Hericia 320 Lamingtonacarus 318 laticollaris, Algophagus 317 laticollaris, Algophagus antarcticus 317 lawrencei, Fusohericia 320 longipes, Prohericia 321 macrolithus, Algophagus 317 microtricha, Neohyadesia 318 minor, Neohyadesia 318 Neohyadesia 318 oreillyorum, Lamingtonacarus 318 paradoxa, Hericia 320 pennsylvanicus, Algophagus 317 pneumatica, Algophagopsis 317 posidonis, Lamingtonacarus 318 Prohericia 321 punctulata, Neohyadesia signyi 318 robini, Hericia 320 sanukiensis, Hericia 321 semicollaris, Algophagus 318 signyi, Neohyadesia 318 Terraphagus 319 Thalassophagacarus 319 322 Acarorum Catalogus IX Distribution of Algophagidae EUROPE Finland (Samšinák, 1972) – Hericia georgei France (contin.) (Türk & Türk, 1957) – Hericia georgei, H. hericia Germany (Türk & Türk, 1957; OConnor & Moser, 1985) – Hericia hericia, H. paradoxa, Prohericia longipes Great Britain (Michael, 1903) – Hericia georgei, H. hericia Italy (Bernini et al., 1995) – Hericia hericia Poland (Haitlinger, 2002, 2004) – Hericia georgei Russia (all, continent.) (Samšinák, 1972) – Hericia georgei ASIA Indonesia (Vitzthum, 1931) - Hericia fermentationis (Sumatra), Fusohericia incredibilis (Sumatra) Japan (Fashing & Okabe, 2006; Hayashi, Ichikawa & Yasui, 2010) – Hericia sanukiensis NORTH AMERICA USA (Baker & Crossley, 1964; Fain & Johnston, 1975; Fashing, 2008) – Algophagopsis pneumatica, Hericia janehenleyi, Fusohericia lawrencei SOUTH and CENTRAL AMERICA Costa Rica (Fashing, 2012) – Fusohericia heliconiae Cuba (Cruz & Socarras, 1992) – Thalassophagacarus faime OCEANIA Australia (continent) (Fashing, OConnor & Kitching, 2000) – Lamingtonacarus oreillyorum, L. posidonis New Zealand (Clark & Andrews, 2012) - Neohyadesia minor Antipodes Islands, New Zealand (Clark & Andrews, 2012) – Terraphagus antipodus ANTARCTICA Heard Island (Hughes, 1955) - Algophagus antarcticus Kerguelen Island (Fain, 1974) - Algophagus antarcticus, A. laticollaris, A. semicollaris, Neohyadesia signyi punctulata South Orkney Islands (Hughes & Goodman, 1969) – Neohyadesia signyi Marion island (Marshall, OConnor & Pugh, 2001) - Neohyadesia microtricha South Georgia (Marshall, OConnor & Pugh, 2001) - Neohyadesia microtricha Prince Edward Islands (Marshall et al., 1999; Marshall, OConnor & Pugh, 2003) - Algophagus brachytarsus, A. macrolithus Acarorum Catalogus IX 323 Hosts of Algophagidae Insecta Coleoptera Fam. Scarabaeidae Cetonia aurata - Hericia georgei Potosia cuprea metallica - Hericia georgei Fam. Nitidulidae Glischrochilus obtusus – Hericia janehenleyi Librodor japonicus – Hericia sanukiensis Lepidoptera Fam. Noctuidae Catocala sponsa – Hericia georgei C. nupta - Hericia georgei C. adultera - Hericia georgei C. elocata - Hericia georgei C. neonympha - Hericia georgei C. promissa - Hericia georgei C. pacta - Hericia georgei C. fraxini - Hericia georgei Nymphalis antiopa - Hericia georgei N. atalanta - Hericia georgei Polygonia c-album - Hericia georgei Acronicta [Apatele] psi - Hericia georgei Fam. Nymphalidae Europis occulta - Hericia georgei Fam. Erebidae Scoliopteryx libatrix - Hericia georgei Bibliography of Algophagidae Baker E.W., D.A. Crossley. 1964. A new species of mite, Fusohericia lawrencei, from an artificial tree-hole (Acarina: Glycyphagidae). – Annals of the Natal Museum, 16: 4. Bernini F., M. Castagnoli, R. Nannelli. 1995. Checklist delle specie fauna d’Italia. Fascicolo 24 Arachnida: Acari. Ed. Calderini. Canestrini G. 1888. Prospetto dell’Acarofauna Italiana. Famiglie: Tarsonemini e Tiroglofini e Appendice ogli Analgesini. Vol.3: 309–425. Clark J.M.; N. Andrews. 2012: New mites Terraphagus antipodus gen. n., sp. n. and Neohyadesia minor sp. n. (Acarina: Astigmata: Algophagidae) from islands of the Southern Ocean. Polar biology, 35(10): 1473–1485 324 Acarorum Catalogus IX Cruz J. de la, A.A. Socarras. 1992. Nuevos genero y especie de acaro litoral (Acarina: Algophagidae) de Cuba. – Poeyana, 412: 1–4. Fain A. 1974. Acariens récoltés par le Dr. J. Travé aux îles subantarctiques I. Familles Saproglyphidae et Hyadesidae (Astigmates). – Acarologia, 16(4): 684–708. Fain A. 1981. Notes on the Hyadesiidae Halbert, 1915 and Algophagidae Fain, 1974, nov. tax. (Acari, Astigmata), with a redescription of Hyadesia curassaviensis Viets, 1936 and H. sellai Viets, 1937. - Acarologia, 22(1): 47–61. Fain A., D. Johnston. 1975. A new algophagid mite Algophagopsis pneumatica n.g., n.sp. living in a river (Astigmata: Hyadesidae). – Bull. Ann. Soc. r. belge Ent., 111: 67–70. Fashing N. J. 1991. Deutonymphal dimorphism in the genus Hericia (Astigmata: Algophagidae). In: Dusbabek, F. & Bukva, V. (eds.) Modern Acarology Vol. 2. Prague, Academia and The Hague, SPB Academic Publishing bv. pp. 287–291. Fashing N. J. 1992. Observations on the feeding biology of Algophagus pennsylvanicus (Astigmata: Algophagidae), a mite restricted to water-filled treeholes. - International Journal of Acarology, 18(2): 77–81. Fashing N.J. 2001. Morphological adaptations associated with mating behaviour in the genus Hericia (Algophagidae: Astigmata). In R. B. Halliday, D. E. Walter, H. C. Proctor, R. A. Norton and M. J. Colloff (eds.) Proceedings of the 10th International Congress of Acarology. CSIRO Publishing, Collingwood, Victoria, Australia. pp. 176–179. Fashing N. J. 2008. Biology of the genus Hericia (Algophagidae: Astigmata), with the description of a new species from the eastern United States. - Systematic and Applied Acarology, 13(1): 3–25. Fashing N. J. 2016. Fine structure of the axillary organs of Hericia janehenleyi Fashing (Algophagidae: Astigmatina : Sarcoptiformes) with a discussion of a potential correlation between axillary organ size and habitat. - Journal of the Acarological Society of Japan, 25(S1): 3–17 Fashing N. J., A. Glist. 2012. A new Fusohericia (Acari: Astigmata: Algophagidae) from Costa Rica. - Systematic and Applied Acarology, 17(3): 591–606. Fashing N. J., K. Okabe. 2006. Hericia sanukiensis, a new species of Algophagidae (Astigmata) inhabiting sap flux in Japan. - Systematic and Applied Acarology Special Publications, 22(1): 1–14. Fashing N. J., L.L. Wiseman. 1980. Algophagus pennsylvanicus a new species of Hyadesiidae from water-filled treeholes. – Internat. J. Acarology, 6(1): 79–84. Hayashi K., T. Ichikawa, Y. Yasui. 2010. Life history of the newly discovered Japanese tree sap mite, Hericia sanukiensis (Acari, Astigmata, Algophagidae). – Exp. Appl. Acarol., 50(1): 35–49. Haitlinger R. 2002. Mites (Acarina) associated with Cetoniinae and Trichiinae (Insecta: Coleoptera: Scarabaeidae) in Poland. – Postępy polskiej acarologii: 63–73. Haitlinger R. 2004. Mites (Acari) occuring on some Coleoptera (Insecta) in Poland. – Polish Journal of Entomology, 73: 3–24. Acarorum Catalogus IX 325 Hughes A.M. 1955. A new genus and species of hyadesid mites – Algophagus antarcticus from Heard Island. – A.N.A.R.E. Rep., B1 (Zool.): 1–19. Hughes A.M., B.J.A. Goodman. 1969. Neohyadesia signyi (Hyadesidae: Acarina): a new genus and species from Signy Island, South Orkney Islands. - British Antarctic Survey Bulletin, 22: 39–48. Marshall D. J., N.G.M. Gremmen, L. Coetzee, B.M. OConnor, P.J.A. Pugh, P.D. Theron, E.A. Ueckermann. 1999. New records of Acari from the sub-antarctic Prince Edward Islands. – Polar Biology, 21: 84–89. Marshall D.J., B.M. OConnor, Ph.J.A. Pugh. 2006. Neohyadesia microtricha (Acari: Astigmata: Algophagidae): A new species from the sub-Antarctic. - Polar Biology, 24(2): 101–104. Marshall D.J., B.M. OConnor, Ph.J.A. Pugh. 2003. Algophagus mites (Astigmata: Algophagidae) from the sub-Antarctic Prince Edward Islands: habitat-related morphology and taxonomic descriptions. - Journal of Zoology, 259(01): 31–47 Michael A.D. 1901–1903. British Tyroglyphidae. – Adlard and Son, London, 1901–1: 1–291; 1903–2: 1–183. OConnor B.M. 2009. The mite community associated with Xylocopa latipes (Hymenoptera: Anthophoridae: Xylocopinae) with description of a new type of acarinarium. – International Journal of Acarology, 19(2): 159–166. OConnor M., J.C. Moser. 1985. Phylogenetic relationships of the Algophagidae (Acari: Astigmata), with descriptions of a new subfamily, genus, and species. - Annals of the Entomological Society of America, 78(6): 783–789. Robin M.C. 1868. Sur une espèce nouvelle de Sarcoptides du genre Glyciphage. Journal of Anatomy and Physiology (Paris), 5: 603–625. Samšinák K. 1972. Redescription of Hericia georgei Michael, 1903 (Acarina, Tyroglyphydae) phoretic on Lepidoptera. – Ann. Zool. Fenn., 9: 65–68. Schenker R. 1986. Biological and ecological notes on Neohyadesia signyi (Acari: Astigmata). - British Antarctic Survey Bulletin, 70: 85–91. Türk E., F. Türk. 1957. Systematik und Ökologie der Tyroglyphiden Mitteleuropas in H. J. Stammer: Beiträge zur Systematik und Ökologie Mitteleuropäischer Acarina. – Akad. Verlagsges., Leipzig, 1, Teil 1, Abh. 1: 1–231. Vazquez I., M.G. Lopez-Campos, M. G. 1996. Prostigmatid mites (Prostigmata) from a littoral zone in Veracruz, Mexico. - International congress of acarology, 575–578. Vitzthum H. 1931. Terrestrische Acarinen (unter Ausschluße der Oribatiden und Ixodiden) der Deutschen Limnologischen Sunda-Expedition. - Archiv für Hydrobiologie (Supplement), 9(1): 59–134. 326 Acarorum Catalogus IX The Mite Family Hemisarcoptidae Hemisarcoptidae Oudemans, 1908: 60 Hemisarcoptoid mites are common inhabitants of various arboreal or aquatic habitats, with many species specialized as inquilines in nests of Hymenoptera. Linobia coccinellae is ectoparasite (hematophagous) of Chrysomela populi and Ch. saliceti (Coleoptera: Chrysomelidae). Other Linobia species have been described on the plant Eichhornia crassipes in Egypt (Soliman, Hussein & Ramadan, 1991). According to OConnor (1982), “Deutonymphs of Hemisarcoptes are known to be phoretic only on coccinellid beetles of the genus Chilocorus, which feeds exclusively on scale insects ”. Hemisarcoptes species are used for control of scale insects (Coccodea), as they are predaceous on the eggs and adults of these insects (Gerson, 1967). The peculiar Espeletiacarus andinus Fain, 1987 is known from the high Andes in Colombia on the flowers of Espeletia incana. Composition of Hemisarcoptidae Fam. Hemisarcoptidae - 8 gen., 24 sp. Divilia Sevastjanov, 1969–3 sp. Espeletiacarus Fain, 1987–1 sp. Hemisarcoptes Lignières, 1893–5 sp. Huronopus OConnor et Houck, 1989–1 sp. Linobia Berlese, 1884–4 sp. Nanacarus Oudemans, 1902–8 sp. Pasohopus OConnor et Houck, 1989–1 sp. Superioropus OConnor et Houck, 1989–1 sp. Catalogue of Hemisarcoptidae Family Hemisarcoptidae Oudemans Hemisarcoptidae Oudemans, 1908: 60 Nanacaridae Oudemans, 1923: 206 Linobiidae Oudemans, 1908 Type genus: Hemisarcoptes Lignières, 1893 Genus Divilia Sevastjanov Divilia Sevastjanov, 1969: 447 [Diagnostic, Fain, Hurst, Tweddle, Lachlan, Majerus & Britt 1995: 103] Acarorum Catalogus IX 327 Type species: Divilia oculata Sevastjanov, 1969 Divilia exigua Fain, Hurst, Fassotte, Webberley, Sloggett et Majerus Divilia exigua Fain, Hurst, Fassotte, Webberley, Sloggett & Majerus, 1997: 91 Type locality: Torun Distribution: Poland Host: Vibidia duodecimguttata (Coleoptera: Coccinellidae) Divilia occidentalis Fain, Hurst, Tweddle, Lachlan, Majerus et Britt Divilia occidentalis Fain, Hurst, Tweddle, Lachlan, Majerus & Britt, 1995: 103 Type locality: England Distribution: England Host: Chilocorus renipustulatus (Coleoptera: Coccinellidae, Chilocorinae) Divilia oculata Sevastjanov Divilia oculata Sevastjanov, 1969: 447 Type locality: Ukraine Distribution: Russia, Ukraine Hosts: Lasius fuliginorus, Formica polyctena, F. rufa (Hymenoptera: Formicidae) Divillia sp. Hosts: Chrysops univittatus, Hybomitra epistates, H. frontalis, H. lasiophthalma, H. nuda, Tabanus quinquevittatus (Canada) Genus Espeletiacarus Fain Espeletiacarus Fain, 1987: 37 Type species: Espeletiacarus andinus Fain, 1987 Espeletiacarus andinus Fain Espeletiacarus andinus Fain, 1987: 38 Type locality: Páramo La Rusia, ca. 200 km NNW of Bogotá, 10 km N of Duitame, alt. 3550 m Distribution: Colombia Host: from a flower of Espeletia incana [Genus Giardius Perraud, 1896; placement? Nanacaridae] [Giardius vitis Perraud, 1896] Genus Hemisarcoptes Lignières Hemisarcoptes Lignières, 1893: 16; [Diagnostic: Fain, 1988: 127] Subgenus Coleovidia H. A. Thomas 328 Acarorum Catalogus IX Vidia (Coleovidia) H. A. Thomas, 1961: 461 [syn. of Hemisarcoptes by Gerson 1967: 637]. Type species: Vidia cooremani H. A. Thomas, 1961 Type species: Acarus malus Shimer, 1868 Hemisarcoptes budensis Fain et Ripka Hemisarcoptes budensis Fain & Ripka, 1998: 35 Type locality: Budapest Distribution: Hungary Hosts: Chilocorus renipustulatus (Coleoptera: Coccinellidae) & Coccoidea (Hemiptera) Hemisarcoptes coccophagus Meyer Hemisarcoptes coccophagus Meyer, 1962: 414; Charles, Hill & Allan, 1995: 297; Houk, 1999: 100 Type locality: Letaba, Transvaal, South Africa Distribution: New Zealand, South Africa Hemisarcoptes cooremani (Thomas) Vidia (Coleovidia) cooremani Thomas, 1961: 461 Hemisarcoptes cooremani (Thomas): syn. by Gerson, 1967: 637; Houck & OConnor, 1990: 870 Type locality: Weslaco, Texas Distribution: USA Host: Chilocorus cacti (Coleoptera: Coccinellidae) Hemisarcoptes dzhashii K. Dzhibladze Hemisarcoptes dzhashii K. Dzhibladze, 1969: 435; Rekk, 1976: 40 Type locality: Zelenyy Cape Distribution: Georgia Hemisarcoptes malus (Shimer) Acarus malus Shimer, 1868: 368 Hemisarcoptes malus (Shimer): André, 1942: 173; Cooreman, 1951: 30; Redescription: Vacante 1985: 750; Abdelgayed et al., 2019: 32 Hemisarcoptes coccisugus Lignières, 1893: 75 Type locality: Mt. Carroll, Illinois Distribution: Algeria, Canada, Egypt, Italy, Morocco, USA Hosts: Chrysomphalus dictyospermi, Lepidosaphes beckii, L. gloveri, Parlatoria blanchardi, P. zizyphi (Coccodea: Diaspididae) Genus Huronopus OConnor et Houck Huronopus OConnor & Houck, 1989b: 6 Type species: Huronopus michiganensis OConnor et Houck, 1989 Acarorum Catalogus IX 329 Huronopus michiganensis OConnor et Houck Huronopus michiganensis OConnor & Houck, 1989b: 6 Type locality: Michigan Distribution: USA Genus Linobia Berlese Linobia Berlese, 1884: 29 Type species: Acarus coccinellae Scopoli, 1763 Linobia coccinellae (Scopoli) Acarus coccinellae Scopoli, 1763: 389 Berlese, 1884: 29; Théodoridès, 1955: 71; Haitlinger, 1999: 181; Bernini et al., 1995; Tarasi, Saboori & Sadeghi, 2001; Trach, 2016; Pohlink, Georgi & Müller, 2020: 723 Type locality: Europe Distribution: Czech Rep., France, Germany, Great Britain, Iran, Italy, Poland, Ukraine Hosts: Chrysomela populi, C. saliceti, Leptinotarsa decemlineata (Coleoptera: Chrysomelidae) Linobia eichhornia Soliman, Hussein et Ramadan Linobia eichhornia Soliman, Hussein & Ramadan, 1991: 22 Type locality: Egypt Distribution: Egypt Host: Eichhornia crassipes (plant) Linobia nilotica Soliman, Hussein et Ramadan Linobia nilotica Soliman, Hussein & Ramadan, 1991: 17 Type locality: Egypt Distribution: Egypt Host: Eichhornia crassipes (plant) Linobia replacementica Soliman, Hussein et Ramadan Linobia replacementica Soliman, Hussein & Ramadan, 1991: 19 Type locality: Egypt Distribution: Egypt Host: Eichhornia crassipes (plant) Genus Nanacarus Oudemans Nanacarus Oudemans, 1902: 7, 45 [Diagnostic, Fain 1988: 127; Redescription, Fain, 1985: 280] Froweinia Oudemans, 1923: 183 Congovidiella Fain, 1971: 282] [type is hieroglypha; syn. Fain 1988: 126] Nanacaroides Volgin & Mironov, 1979: 93 [type is elongatus; syn. Purvis & Evans 1982: 820] 330 Acarorum Catalogus IX Type species: Hypopus minutus Oudemans, 1901 Nanacarus collembolicola (Fain et Johnston) Congovidiella collembolicola Fain & Johnston, 1974: 418 Nanacarus collembolicola (Fain et Johnston) Type locality: England, Domleo, Spinney, Sutton Bonington Distribution: Great Britain Host: Sminthurus fuscus (Collembola, Symphypleona: Sminthuridae) Nanacarus elongatus (Volgin et Mironov) Nanacaroides elongatus Volgin & Mironov, 1979: 93 Nanacarus elongatus (Volgin et Mironov): Purvis & Evans, 1982: 820 Type locality: Habarovskiy Kray, Bolshe-Hehcirskiy Zapovednik (the Far East of Russia) Distribution: Russia Nanacarus hieroglypha (Fain et Elsen) Congovidiella hieroglypha Fain et Elsen, in Fain, 1971: 282 Nanacarus hieroglypha (Fain et Elsen): Type locality: Haut Lopori Distribution: D.R. Congo Host: Glossina fuscipes fuscipes (Diptera: Glossinidae) Nanacarus hungaricus Halmai et Mahunka Nanacarus hungaricus Halmai & Mahunka, 1980: 266 [mammalian host] Type locality: Hungary Distribution: Hungary Nanacarus longisetosus Volgin et Mironov Nanacarus longisetosus Volgin & Mironov, 1979: 91 Type locality: Buryatian ASSR, Gremyachenskiy Rayon, Istok Distribution: Russia Nanacarus manus Purvis et Evans Nanacarus manus Purvis & Evans, 1982: 815 Type locality: S.E. Ireland Distribution: Ireland Nanacarus minutus (Oudemans) Hypopus minutus Oudemans, 1901: 85 Nanacarus minutus (Oudemans): Krivoluckiy, 1975: 474; Fain 1985: 280; Fain et al., 1997: 94; Li et al., 1992: 23 Acarorum Catalogus IX 331 Type locality: Indonesia (Java) Distribution: China, Germany, Indonesia (Java, Buru), Poland, Russia Hosts: Adalia sp., Calvia duodecimguttata, Exochomus quadripustulus (Coleoptera: Coccinellidae); Xylocopa (Koptorthosoma) tenuiscapa (Hymenoptera: Apidae, Xylocopinae), Eptesicus serotinus, Pipistrellus pipistrellus, Sorex araneus (Mammalia), Megapodius reinwardt buruensis (Aves: Megapodiidae) Nanacarus nominis Woodring Nanacarus nominis Woodring, 1966: 82 Type locality: Holden, Utah Distribution: USA Host: «from the cells of the alkali bee Nominia melanderi«(Hymenoptera: Halictidae, Nomininae) Nanacarus pragensis (Samšinák et Vobrazkova) Congovidiella pragensis Samšinák & Vobrazkova, 1983: 119 Nanacarus pragensis (Samšinák & Vobrazkova): Khaustov et al., 2018: 48 Type locality: Prague Distribution: Czech Rep. Genus Pasohopus OConnor et Houck Pasohopus OConnor & Houck, 1989a: 17 Type species: Pasohopus labrynthus OConnor et Houck, 1989 Pasohopus labrynthus OConnor et Houck Pasohopus labrynthus OConnor & Houck, 1989a: 17 Type locality: Pasoh Forest Reserve Distribution: Malaysia Genus Superioropus OConnor et Houck Superioropus OConnor & Houck, 1989b: 2 Type species: Superioropus huronmontanus OConnor & Houck, 1989 Superioropus huronmontanus OConnor et Houck Superioropus huronmontanus OConnor & Houck, 1989b: 2; OConnor, 1991; Khaustov et al., 2018: 49 Type locality: the Huron Mountains of northern Michigan Distribution: Russia, USA Hosts: Exeristes comstockii (Hymenoptera: Ichneumonidae), galleries of Ips typographus (Coleoptera: Curculionidae, Scolytinae) 332 Acarorum Catalogus IX Index of Hemisarcoptidae (Names in bold are accepted as valid) andinus, Espeletiacarus 327 budensis, Hemisarcoptes 328 coccinellae, Acarus 329 coccinellae, Linobia 329 coccisugus, Hemisarcoptes 328 coccophagus, Hemisarcoptes 328 Coleovidia 327 collembolicola, Congovidiella 330 collembolicola, Nanacarus 330 Congovidiella 329 cooremani, Hemisarcoptes 328 cooremani, Vidia (Coleovidia) 328 Divilia 326 dzhashii, Hemisarcoptes 327 eichhornia, Linobia 327 elongatus, Nanacarus 330 elongatus, Nanacaroides 328 Espeletiacarus 326 exigua, Divilia 325 Froweinia 329 Hemisarcoptes 327 Hemisarcoptidae 326 hieroglypha, Congovidiella 330 hieroglypha, Nanacarus 330 hungaricus, Nanacarus 330 huronmontanus, Superioropus 331 Huronopus 328 labrynthus, Pasohopus 331 Linobia 329 Linobiidae 326 longisetosus, Nanacarus 330 malus, Acarus 328 malus, Hemisarcoptes 328 manus, Nanacarus 330 michiganensis, Huronopus 329 minutus, Hypopus 330 minutus, Nanacarus 330 Nanacaridae 326 Nanacaroides 329 Nanacarus 329 nilotica, Linobia 329 nominis, Nanacarus 331 occidentalis, Divilia 327 oculata, Divilia 327 Pasohopus 331 pragensis, Congovidiella 331 pragensis, Nanacarus 331 replacementica, Linobia 329 Superioropus 331 Distribution of Hemisarcoptidae EUROPE Belgium (Fain, Hurst et al. 1997) – Nanacarus minutus Czech Rep. (Samšinak & Vobrazkova, 1983) – Nanacarus pragensis France (contin.)(Théodoridès, 1955) – Linobia coccinellae Georgia (Dzhibladze, 1969) – Hemisarcoptes dzhashii Germany (Pohlink, Georgi & Müller, 2020) – Linobia coccinellae, Nanacarus minutus Acarorum Catalogus IX 333 Great Britain (Türk & Türk, 1952; Fain & Johnston, 1974; Fain, Hurst, Tweddle et al., 1995; Fain, Hurst, Fassotte et al. 1997) – Congovidia coccinellidarum, Linobia coccinellae, Nanacarus (= Congovidiella) collembolicola Hungary (Halmai & Mahunka, 1980; Fain & Ripka, 1998) – Hemisarcoptes budensis, Nanacarus hungaricus Ireland (Purvis & Evans, 1982) – Nanacarus manus Italy (contin.) (Bernini et al., 1995) – Hemisarcoptes malus, Linobia coccinellae Sicily (Bernini et al., 1995) – Hemisarcoptes malus Poland (Fain et al., 1997) – Divilia exigua, Nanacarus minutus Russia (all, continent.) (Sevastjanov, 1969; Krivoluckiy, 1975: 474; Volgin & Mironov, 1979) – Divilia oculata, Divilia aff. occidentalis, Hemisarcoptes sp., Nanacarus longisetosus, N. elongatus, N. minutus, Superioropus huronmontanus Ukraine (Sevastjanov, 1969; Trach, 2007) - Divilia oculata, Linobia coccinellae AFRICA Congo DR (Fain, 1971) – Nanacarus hieroglypha Egypt (Soliman, Hussein & Ramadan, 1991; Abdelgayed et al., 2019) – Hemisarcoptes malus, Linobia eichhornia, L. nilotica, L. replacementica Morocco (Cooreman, 1951) – Hemisarcoptes malus South Africa (Meyer, 1962) – Hemisarcoptes coccophagus ASIA China (Fan, Chen & Wang, 2010; Ji, L.-Z., Gerson & Izraylevich, 1994; Li, L.-S., Xuan, J.-Y. & Fan, Q.-H., 1992) – Hemisarcoptes sp., Nanacarus minutus Indonesia (Oudemans, 1901, 1928) – Nanacarus minutus Iran (Tarasi, Saboori & Sadeghi, 2001) – Linobia coccinellae Israel and Palestina (Gerson, 1964, 1967; Gerson & Schneider, 1981, 1982; Izraylevich & Gerson, 1993) – Hemisarcoptes coccophagus Malaysia (OConnor & Houck, 1989a) – Pasohopus labrynthus NORTH AMERICA Canada (Mullen et al., 1989; Tothill, 1918; Baulieu et al., 2019) – Hemisarcoptes malus and two other sp., Divilia sp. USA (Shimer, 1868; Woodring, 1966; OConnor, 1991, OConnor & Houck, 1989b; Luck, Jiang & Houck, 1999) – Huronopus michiganensis, Nanacarus nominis, Superioropus huronmontanus SOUTH and CENTRAL AMERICA Colombia (Fain, 1987) – Espeletiacarus andinus 334 Acarorum Catalogus IX OCEANIA Australia (continent) (Gerson, 1994) – Hemisarcoptes 2 spp. New Zealand (Charles, Hill & Allan, 1995a, 1995b; Hill et al., 1993) – Hemisarcoptes coccophagus Hosts of Hemisarcoptidae Insecta s.l. Collembola Fam. Sminthuridae Sminthurus fuscus – Nanacarus collembolicola Coleoptera Fam. Chrysomelidae Chrysomela populi – Linobia coccinellae C. saliceti - Linobia coccinellae Fam. Coccinellidae Adalia sp. - Nanacarus minutus Chilocorus cacti – Hemisarcoptes cooremani Ch. renipustulatus - Hemisarcoptes budensis Exochomus quadripustulus - Nanacarus minutus Vibidia duodecimguttata - Nanacarus minutus Homoptera Coccodea Fam. Diaspididae Chrysomphalus dictyospermi – Hemisarcoptes malus Hemiberlesia lataniae – Hemisarcoptes coccophagus Lepidosaphes beckii – Hemisarcoptes malus L. gloveri - Hemisarcoptes malus Parlatoria blanchardi – Hemisarcoptes malus P. zizyphi - Hemisarcoptes malus Hymenoptera Fam. Apidae Xylocopa (Koptorthosoma) tenuiscapa – Nanacarus minutus Fam. Halictidae Nominia melanderi – Nanacarus nominis Fam. Formicidae Lasius fuliginorus – Divilia oculata Formica polyctena - Divilia oculata F. rufa - Divilia oculata Fam. Ichneumonidae Exeristes comstockii - Superioropus huronmontanus Acarorum Catalogus IX 335 Diptera Fam. Glossinidae Glossina fuscipes fuscipes – Nanacarus hieroglypha Aves Fam. Megapodiidae Megapodius reinwardt buruensis – Nanacarus minutus Mammalia Soricomorpha Fam. Soricidae Sorex araneus – Nanacarus minutus Chiroptera Fam. Vespertilionidae Eptesicus serotinus – Nanacarus minutus Pipistrellus pipistrellus - Nanacarus minutus Bibliography of Hemisarcoptidae Abdelgayed S. A., N. M Abd El-Wahed, A. M. Ali, S. Eraky. 2019. Species Composition and Diversity of Mites Inhabiting Pomegranate Orchards at Assiut, Upper Egypt. - Acarines, 13: 29–36. André M. 1942. Sur l’Hemisarcoptes malus Shimer (= coccisugus Lignières). – Bull., Mus. Paris, 2e sér., 14(3): 173–180. Bernini F., M. Castagnoli, R. Nannelli. 1995. Checklist delle specie fauna d’Italia. Fascicolo 24 Arachnida: Acari. Ed. Calderini. Charles J.G., M.G. Hill, D.J. Allan. 1995a. Releases and recoveries of Chilocorus spp. (Coleoptera: Coccinellidae) and Hemisarcoptes spp. (Acari: Hemisarcoptidae) in kiwifruit orchards: 1987–93. – New Zealand J. Zool., 22: 319–324. Charles J.G., M.G. Hill, D.J. Allan. 1995b. Persistence of the predatory mite, Hemisarcoptes coccophagus Meyer (Hemisarcoptidae), on low populations of Hemiberlesia lataniae (Signoret) (Diaspididae) in New Zealand. - Israel Journal of Entomology, 29: 297–300. Cooreman J. 1951. 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Gr. Lakes Entomol., 22: 1–10. Oudemans A.C. 1901. Notes on Acari: Third series. - Tijdschrift der Nederlandsche Dierkundige Vereeniging. Leiden, 7: 50–87. Oudemans A.C. 1902. Acarologische Aanteekeningen. – Ent. Berichten, 1(7): 43–45. Oudemans A.C. 1908. Notes on Acari. XVth series. (Parasitidae, Acaridae). – Tijdschrift voor entomologie, 51: 28–88. Oudemans A.C. 1923. Acarologische Aanteekeningen LXXX. - Ent. Berichten, 6(130): 145–155. Oudemans A.C. 1928. Fauna Buruana Acari. - Treubia, 7: 37–100. Pohlink K., R. Georgi, M. Müller. 2020. Occurrence and impact of the parasitic Mite Linobia coccinellae on its host beetle Chrysomela populi: implications for its potential as a biological control agent. – Entomologia experimentalis et aplicata, 168: 723–731. Purvis G., G.O. Evans. 1982. Astigmatic mites of the genera Schwiebea Oudms. and Nanacarus Oudms. from S.E.Ireland. - Journal of Natural History, 16(6): 815–821. Rekk G.F. 1976. [Catalogue of the Acarofauna of Georgian SSR]. Ed. “Mezniereba”, 127 pp. Samšinák K. 1957. Einige Bemerkungen zur Faunistik der in Gesellschaft von Insekten lebenden Acari. – Acta faun. Ent. Mus. Nat. Pragae, 2: 109–114. Samšinák K., E. Vobrazkova. 1983. Mites from the city pavement. - Věstník Československé Zoologické Společnosti, 47: 118–121. Sevastjanov V.D. 2003. Определитель родов клещей семейства Saproglyphidae фауны мира по самкам и самцам. - Вісник Одеського національного університету, 8(6): Biology, 138–143 [Key for identification of mites of the family Saproglyphidae of the world’s fauna – males and females. – Visnik Odeskogo Nacionalnogo universitetu]. Shimer H. 1868. Notes on the “Apple Bark-Louse” Lepidosaphes conchiformis (Gmelin) with a decription of a supposed new Acarus. – Trans. Am. Entomol. Soc., 1: 361–374. Smirnoff W. 1957. - La cochenille du palmier dattier (Parlatoria blanchardi Targ.) en Afrique du Nord, comportement, importance économique, prédateurs et lute biologique. – Entomophaga, 2: 1–99. Soliman F. E.-S., Hussein, A.A. Ramadan. 1991. Description of four water mite species belonging to genus Linobia (family Linobiidae, Canestrinioidea, Astigmata, Acarina) from the Nile. - Bul. Fac. Sci. Assiut Univ., E Zool., 20(2): Acarorum Catalogus IX 339 Tarasi J., A. Saboori, S.E. Sadeghi. 2001. Report of an ectoparasitic mite on adults of Melasoma populi L. from Iran. - Journal of Entomological Society of Iran, 21(2): 107–108. Théodoridès J. 1955. Contribution a l’étude des parasites et phorétiques de coléoptères terrestres. - Vie et Milieu, Supplements 4: 1–310. Thomas H. A. 1961. Vidia (Coleovidia) cooremani, new subgenus and new species, and notes on the life history (Acarina: Saproglyphidae). - Ann. Ent. Soc. Amer., 54: 461–463. Tothill J.D. 1918. The predaceous mite Hemisarcoptes malus Shimer and its relation to the natural control of the oyster-shell scale, Lepidosaphes ulmi. – Agricultural Gazette, Canada, 5: 234–239. Trach V.A. 2007. On the record of Linobia coccinellae (Astigmata, Hemisarcoptidae) in Ukraine. -Vestnik Zoologii, 41: 510. Türk P., S. Türk. 1952. LV.—Studies of Acari.—7th series: “Records and descriptions of mites new to the British fauna, together with short notes on the biology of sundry species”. Annals and Magazine of Natural History, 5: 475– 506. Türk P., S. Türk. 1957. Systematik und Ökologie der Tyroglyphiden Mitteleuropas in H.- J. Stammer: Beiträge zur Systematik und Ökologie Mitteleuropäischer Acarina. – Akad. Verlagsges., Leipzig, 1, Teil 1, Abh. 1: 1–231. Volgin V. I., S.V. Mironov. 1979. New species and new genus of mites of the family Saproglyphidae (Acarina, Acaroidea). – Trudy Zool. Inst. Akad. Nauk U.S.S.R., 85: 91–98 (in Russian). The Mite Family Carpoglyphidae Carpoglyphidae Oudemans, 1923: 206 The best known representative of the Carpoglyphidae is the dried fruit mite, Carpoglyphus lactis (L.). This species is a cosmopolitan pest of stored materials with a high sugar content, such as dried fruit, powdered milk, and honey, as well as of other stored food substrates (Chmielewski 1971; Hughes 1976). Deutonymphs have been collected from Lepidoptera and Coleoptera (Türk & Türk, 1957). Composition of Carpoglyphidae Fam. Carpoglyphidae Oudemans, 1923–4 gen., 12 sp. Carpoglyphus Robin, 1869–9 sp. Coproglyphus Türk et Türk, 1957–1 sp. Dichotomiopus Fain et Camerik, 1978–1 sp. Pullea Canestrini, 1884–1 sp. 340 Acarorum Catalogus IX Catalogue of Carpoglyphidae Family Carpoglyphidae Oudemans Carpoglyphidae Oudemans, 1923: 206 Type genus: Carpoglyphus Robin, 1869 Genus Carpoglyphus Robin Carpoglyphus Robin, 1869: 197 Type species: Acarus lactis Linnaeus, 1758 Carpoglyphus afzali Khan Carpoglyphus afzali Khan, 1967: 398 Type locality: Pakistan Distribution: Pakistan Carpoglyphus alienus Banks Carpoglyphus alienus Banks, 1905: 42 Type locality: “Batavia” (Java) Distribution: Indonesia (Java) Carpoglyphus ganzhouensis Jiang Carpoglyphus ganzhouensis Jiang, 1991: 82 Type locality: Jiangxi Distribution: China Carpoglyphus lactis (Linnaeus, 1758) [ 1767?] Acarus lactis Linnaeus, 1767: 1024 Carpoglyphus lactis (Linnaeus): Jeleva & Angelkova, 1964: Carpoglyphus passularum Robin, 1869 Glycyphagus anonymus Haller, 1882 Type locality: Europe Distribution: Argentina, Australia, Bulgaria, China, England, Germany, Italy Carpoglyphus munroi Hughes Carpoglyphus munroi Hughes, 1962: 168 Type locality: Imperial College, Field Station, Silwood Park, Ascot, Surrey Distribution: England Carpoglyphus nidicolous Hubard et Fashing Carpoglyphus nidicolous Hubard & Fashing, 1996: 215 Acarorum Catalogus IX 341 Type locality: Oregon Distribution: USA Host: Petrochelidon pyrrhonota (Aves) nests Carpoglyphus sturmi Fain et Rack Carpoglyphus sturmi Fain & Rack, 1987: 12 Type locality: Páramo de Montserrate, 3250 m Distribution: Colombia Host: Espeletiopsis corymbosa (plant) Carpoglyphus vagabundus Berlese Carpoglyphus vagabundus Berlese, 1910: 267 Type locality: Rosignano (Pisa) Distribution: Italy Host: nest of Polystes gallicus (Hymenoptera: Vespidae) Carpoglyphus wardleorum Clark Carpoglyphus wardleorum Clark, 2010: 454 Type locality: Ashley Gorge, Canterbury Distribution: New Zealand Host: from sooty mould fungi on the bark of black beech, Nothofagus solandri. Genus Coproglyphus Türk et Türk Coproglyphus Türk & Türk, 1957: 181 Type species: Coproglyphus stammeri Türk et Türk, 1957 Coproglyphus stammeri Türk et Türk Coproglyphus stammeri Türk & Türk, 1957: 181; Li, 1999: 72 Type locality: Erlangen (Germany) Distribution: China, Germany Genus Dichotomiopus Fain et Camerik Dichotomiopus Fain & Camerik, 1978: 172 Type species: Dichotomiopus biaggioi Fain et Camerik, 1978 Dichotomiopus biaggioi Fain et Camerik Dichotomiopus biaggioi Fain & Camerik, 1978: 173 Type locality: Cotia, Sao Paulo Distribution: Brazil Host: Dichotomius anaglypticus (Coleoptera: Scarabaeidae) 342 Acarorum Catalogus IX Genus Pullea Canestrini Pullea Canestrini, 1884: 720 Type species: Pullea discoidalis Canestrini, 1884 Pullea discoidalis Canestrini Pullea discoidalis Canestrini, 1884: 720 Type locality: Australia Distribution: Australia Index of Carpoglyphidae (Names in bold are accepted as valid) afzali, Carpoglyphus 340 alienus, Carpoglyphus 340 anonymus, Glyciphagus 340 biaggioi, Dichotomiopus 341 Carpoglyphidae 340 Carpoglyphus 340 Coproglyphus 341 Dichotomiopus 341 discoidalis, Pullea 342 ganzhouensis, Carpoglyphus 340 lactis, Acarus 340 lactis, Carpoglyphus 340 munroi, Carpoglyphus 340 nidicolous, Carpoglyphus 340 passularum, Carpoglyphus 340 Pullea 342 stammeri, Coproglyphus 341 sturmi, Carpoglyphus 341 vagabundus, Carpoglyphus 341 wardleorum, Carpoglyphus 341 Distribution of Carpoglyphidae EUROPE Bulgaria (Jeleva & Angelkova, 1964) – Carpoglyphus lactis Germany (Türk & Türk, 1957) – Carpoglyphus lactis, Coproglyphus stammeri Great Britain (Hughes, 1962) – Carpoglyphus lactis, C. munroi Holland (Siepel, Cremers & Vierbergen, 2016) - Carpoglyphus lactis Italy (contin.) (Berlese, 1910) – Carpoglyphus lactis, C. vagabundus Turkey (Turgu & Kumral, 2019) - Carpoglyphus lactis ASIA China (Jiang, 1991; Li, 1999; Fan, Chen & Wang, 2010) – Carpoglyphus ganzhouensis, C. lactis, Coproglyphus stammeri Indonesia (Banks, 1905) – Carpoglyphus alienus Pakistan (Khan, 1967) – Carpoglyphus afzali Acarorum Catalogus IX 343 NORTH AMERICA USA (Hubard & Fashing, 1996) – Carpoglyphus nidicolous SOUTH and CENTRAL AMERICA Argentina (Hughes, 1978) – Carpoglyphus lactis Brazil (Fain & Camerik, 1978) – Dichotomiopus biaggioi Colombia (Fain & Rack, 1987) – Carpoglyphus sturmi OCEANIA Australia (continent) (Womersley, 1941) – Carpoglyphus lactis, Pullea discoidalis New Zealand (Clark, 2010) – Carpoglyphus wardleorum Hosts of Carpoglyphidae Hymenoptera Fam. Vespidae Polystes gallicus - Carpoglyphus vagabundus Bibliography of Carpoglyphidae Baker E.W., M.D. Delfinado. 1978. Notes on dried fruit mite Carpoglyphus lactis (Acarina: Carpoglyphidae) infesting honeybee combs. - Jour. Agr. Res., 17: 52–54. Banks N. 1905. An alleged parasitic Tyroglyphid. - Proceedings of the Entomological Society of Washington, 7(1): 40–42. Berlese A. 1910. Lista di nuove specie e nuovi generi di Acari. – Redia, 6(2): 242–271. Chmielewski W. 1971. Morfologia, biologia i ekologia Carpoglyphus lactis (L., 1758) (Glycyphagidae, Acarina). - Prace Nauk. Inst. Ochr. Rostlin, 13: 63–166. Chmielewski W. 1999. Bionomics of Carpoglyphus lactis (Acari: Carpoglyphidae) on honey. - Ecology and Evolution of the Acari, Series Entomologica book series (SENT, volume 55), 423–424 Clark J.M. 2010. A new species of Carpoglyphus (Astigmatina: Carpoglyphidae) from the bark of black beech (Nothofagus) honeydew in New Zealand. – Int. J. of Acarology, 36(5): Fain A., A.M. Camerik. 1978. New hypopi phoretic on Brazilian Coleoptera and Hymenoptera (Acari: Astigmata). – Intl. J. Acar., 4(3): 169–178. Fain A., G. Rack. 1987. Notes on the mites living in the flowers of Espeletia spp. (Asteraceae) in Colombia. 1. Carpoglyphus sturmi sp. n. (Acari, Carpoglyphidae). - Entomologische Mitteilungen aus dem Zoologischen Staatsinstitut und Zoologischen Museum Hamburg, 9(130): 9–19. Fan Qing-Hai, Yan Chen, Zi-Qing Wang. 2010. Acaridia (Acari: Astigmatina) of China: A review of research progress. – p.225–259. In: Zhang Z.-Q., Hong X.-Y. & Fan Q.-H. (eds) Xin Jie-Lin Centenary: Progress in Chinese Acarology. Zoosystema 4, 1–345. 344 Acarorum Catalogus IX Hubard C.K., N.J. Fashing. 1996. Carpoglyphus nidicolus - a new species of Carpoglyphidae (Acarina: Astigmata) inhabiting the nests of swallows. -International Journal of Acarology, 22(3): 215–220. Hughes A.M. 1952. A new glycyphagid mite - Carpoglyphus munroi. - Journal of the Linnean Society of London, Zoology, 42(284): 168–175. Hughes A.M. 1976. The mites of stored food and houses (No. 2nd edition). Her Majesty’s Stationery Office, London, 400 pp. Jeleva M., E.B. Angelkova. 1964. Contribution to the study of the Tyroglyphid fauna in Bulgaria. – Ann. Univ. Sofia, 56, Fac. Biol., Géol., Géogr., Livre 1 – Biologie (Zoologie): 141–146 (in Bulg., summ. Russ., Engl.). Jiang Z.-T. 1991. A new species of Carpoglyphidae from China (Acarina: Acaroidea). - Journal of Nanchang University (Natural Science), 15(1): 82–86. Li C.P. 1999. Preliminary study of the acaridid mites breeding in stored Chinese traditional medical materials. – Chinese Journal of Parasitic Diseases Control, 12(1): 72–73. ODonnovan W.J. 1920. Carpoglyphus Passularum causing Dermatitis. - Proc R Soc Med. 1920; 13 (Dermatol Sect): 150–151. Oudemans A.C. 1923. Studie over de sedert 1877 ontworpen systemen der Acari; nieuwe classificatie; phylogenetische beschouwingen. - Tijdschrift voor entomologie, 66: 49—85. Robin C. 1869. Note sur un nouveau genre d‘acariens. - Journal de l‘Anatomie et de la Physiologie Normales et Pathologiques de l‘Homme et des Animaux, 6: 197–204, plates 7–8. Siepel H., H. Cremers, B. Vierbergen. 2016. Provisional checklist of the astigmatic mites of the Netherlands (Acari: Oribatida: Astigmatina). – Nederlandse Faunistische Mededelingen, 47: 49–88. Tomaševič B. 1956. Carpoglyphus lactis L. na suvim sljivama. - Zastita Bilja, 33: 57–65. Turgu V., N.A. Kumral. 2019. Alternative control agents of the dried fruit mite, Carpoglyphus lactis (L.) (Acari: Carpoglyphidae) on dried apricots. - Acarological Studies, 1(2): 59–64. Türk E., F. Türk. 1957. Systematik und Ökologie der Tyroglyphiden Mitteleuropas in H. J. Stammer: Beiträge zur Systematik und Ökologie Mitteleuropäischer Acarina. – Akad. Verlagsges., Leipzig, 1, Teil 1, Abh. 1: 1–231. Womersley H. 1941. Studies in Australian Acarina. (2) Tyroglyphidae (s.l.). – Records of the South Australian Museum, 6(4): 481–487. The Mite Family Winterschmidtiidae (Saproglyphidae) Winterschmidtiidae Oudemans, 1923 Saproglyphinae Oudemans, 1924 Winterschmidtiidae (= Saproglyphidae) is the largest family in the Hemisarcoptoidea, with 28 genera and 139 known species with a wide range of ecological associations. Acarorum Catalogus IX 345 Species of Saproglyphus are fungivores in decaying materials, bark crevices, and woody fungi, and deutonymphs are dispersed by a wide variety of insecthosts. Other species are common fungivores on leaf surfaces. In temperate regions, the tritonymph is a resistant overwintering stage. Species of Winterschmidtia are also predators of the eggs of bark beetles (Scolytinae). Some taxa are known from deutonymphs associated with various families of subcortical Coleoptera. Other winterschmidtiid genera are found in various types of decaying material such as dung, carrion and in vertebrate nests. Procalvolia zacheri (Oudemans) and Acalvolia squamata (Oudemans) occasionally infest stored food products, P. zacheri was associated with severe dermatitis in a horse. The subfamily Ensliniellinae includes obligate associates of Hymenoptera. Some taxa are associated with sphecid wasps (Crabrovidia and others), most species are associates of Vespidae. Other genera are associated with solitary wasps of the subfamilies Masarinae (pollen wasps) and Eumeninae. A partial list of associations and a phylogeny for the subfamily was provided by OConnor and Klompen (1999). Some species such as Monobiacarus quadridens Baker and Kurosaia jiju OConnor et Okabe feed on hemolymph of paralyzed arthropod prey provisioned by the female wasp. Other species, in the genera Ensliniella, Kennethiella, and Vespacarus feed directly on hemolymph of the wasp host larva or pupa. Many female (or rarely male) wasp hosts have more or less elaborated acarinaria in different locations on the body that harbor dispersing deutonymphs. Based on their phylogeny, OConnor and Klompen (1999) concluded that acarinaria have evolved independently in many host groups. For many years was used the family name Saproglyphidae Oudemans, 1924, but OConnor (1984) insisted to use the antedating name Winterschmidtiidae Oudemans, 1923 and was followed by other researchers. Now both names are used on subfamily level. Composition of Winterschmidtiidae Fam. Winterschmidtiidae (Saproglyphidae) - 28 gen., 139 sp. Subfam. Ensliniellinae Vitzthum, 1924–71 sp. Crabrovidia Zakhvatkin, 1941–20 sp. Ensliniella Vitzthum, 1925–7 sp. Kennethiella Cooreman, 1954–1 sp. Kurosaia Okabe et OConnor, 2002–1 sp. Macroharpa Mostafa, 1970–4 sp. Monobiacarus Baker et Cunliffe, 1960–3 sp. Sphexicozela Mahunka, 1970–1 sp. Vespacarus Baker, 1960–10 sp. Vidia Oudemans, 1905–2 sp. Zethacarus Mostafa, 1971–6 sp. Zethovidia Mostafa, 1970–16 sp. Subfam. Saproglyphinae Oudemans, 1924–18 sp. Acalvolia Fain, 1971–2 sp. 346 Acarorum Catalogus IX Congovidia Fain et Elsen, in Fain, 1971–6 sp. Gambacarus Mahunka, 1975–1 sp. Neocalvolia Hughes, 1970–3 sp. Neosuidasia Ranganath et Channabasa,1983–1 sp. Neottiglyphus Volgin, 1974–1 sp. Saproglyphus Berlese, 1890–4 sp. Subfam. Oulenziinae Oudemans, 1928– 7 sp. Oulenzia Radford, 1950–1 sp. Oulenziella Fan et al., 2015–2 sp. Procalvolia Fain, 1971–3 sp. Trypetacarus Fain, 1971–1 sp. Subfam. Winterschmidtiinae Oudemans, 1923–50 sp. Allocalvolia Fain et Rack, 1987–2 sp. Calvolia Oudemans, 1911–34 sp. Czenspinskia Oudemans, 1927–2 sp. Parawinterschmidtia Khaustov, 2000–1 sp. Riemia Oudemans, 1925–1 sp. Winterschmidtia Oudemans, 1923 (= Afrocalvolia Fain et Elsen, 1971) – 10 sp. Catalogue of Winterschmidtiidae Fam. Winterschmidtiidae Oudemans Winterschmidtiidae Oudemans, 1923: 206 Saproglyphidae Oudemans, 1924 Czenspinskiidae Oudemans, 1927 Oulenziidae Oudemans, 1928 Calvoliinae Turk & Turk, 1957: 155 Type genus: Winterschmidtia Oudemans, 1923 Subfam. Ensliniellinae Vitzthum Genus Crabrovidia Zakhvatkin Vidia (Crabrovidia) Zakhvatkin, 1941: 371 Crabrovidia Zachvatkin: Fain, 1971: 284 Euvidia Delfinado & Baker, 1976: 81] [Type is utahensis; syn. OConnor & Eickwort 1988: 172] Type species: Vidia (Crabrovidia) gussakovskii Zachvatkin, 1941 Crabrovidia africana Fain Crabrovidia africana Fain, 1973: 183 Acarorum Catalogus IX 347 Type locality: Kibombo Distribution: DR Congo Host: Sphex hemorrhoidalis (Hymenoptera: Sphecidae) Crabrovidia brasiliensis Fain Crabrovidia brasiliensis Fain, 1973: 169 Type locality: Merretes, Alto de Sierra, Sta Catarina, Nova Teutonia Distribution: Brazil Host: Ectemnius cubiceps (Hymenoptera: Crabronidae) Crabrovidia concellaria (Cooreman) Vidia concellaria Cooreman, 1948a: 1 Crabrovidia concellaria Fain, 1973: 179 Type locality: Uccle (Bruxelles) Distribution: Belgium, France Host: Cerceris arenaria (Hymenoptera: Crabronidae); ? accidental host: Hedichrum nobile (Chrysididae) Crabrovidia corsicana Fain Crabrovidia corsicana Fain, 1973: 183 Type locality: Bonifacio Distribution: Corsica Host: Crossocercus elongatus (Hymenoptera: Crabronidae) Crabrovidia crossocerus Fain Crabrovidia crossocerus Fain, 1973: 167 Type locality: Sta Catarina, Nova Teutonia Distribution: Brazil Host: Crossocerus porexus (Hymenoptera: Crabronidae) Crabrovidia cubensis Fain Crabrovidia cubensis Fain, 1973: 170 Type locality: Guantanamo Distribution: Cuba Host: Lestica cubensis (Hymenoptera: Crabronidae) Crabrovidia dasyproctus Fain Crabrovidia dasyproctus Fain, 1973: 177 Type locality: Taiherin (Taiwan) Distribution: India, Philippines, South Africa (Natal), Taiwan 348 Acarorum Catalogus IX Hosts: Dasyproctus agilis orientalis, D. bipunctatus lugubris, D. buddha, D. cevirus, D. westermanni (Hymenoptera: Crabronidae) Crabrovidia ectemnii Fain Vidia gussakovskii: Cooreman, 1948: 8 (nec Zachvatkin, 1941) Crabrovidia ectemnii Fain, 1973: 163 Type locality: La Panne (Belgium) Distribution: Argentina, Austria, Belgium, Bolivia, Brazil, Colombia, Costa Rica, Croatia, Cyprus, France, Greece (Crete), Guiana, Honduras, Jamaica, Japan [Okinawa], Mexico, Morocco, Puerto Rico, Spain, Switzerland Hosts: Ectemnius carinatus, E. cavifrons, E. centralis, E. continuus, E. craesus, E. guttatus, E. hypsae, E. impressus, E. laevigatus, E. lapidaries, E. schettereri, E. semipunctatus, E. singularis, E. sonorensis, E. sexcinctus (Hymenoptera: Crabronidae) Crabrovidia gussakovskii (Zachvatkin) Vidia (Crabrovidia) gussakovskii Zachvatkin, 1941: 270 Crabrovidia gussakovskii (Zachvatkin): Fain, 1973: 160; Klimov, 1998: 25 Type locality: Crimea (Karadag) Distribution: Russia, Crimea, Kuril Isl. (Iturup) Hosts: Crabro solenius, Ectemnius (Metacrabro) fossorius, E. (Hypocrabro) impressus, E. (Hypocrabro) continuus (= vagus auct.)(Hymenoptera: Crabronidae) Crabrovidia hirsuta (OConnor et Eickwort) Vidia hirsuta OConnor & Eickwort, 1988: 161 Crabrovidia hirsuta (OConnor et Eickwort): Type locality: New Mexico, Sandoval Co., Jemez Springs Distribution: USA Host: Megachile (Sayapis) fidelis (Hymenoptera: Megachilidae) Crabrovidia julianii (Sevastianov) Vidia julianii Sevastianov, in Kadzhaya & Sevastianov, 1967: 199 Crabrovidia julianii (Sevastianov): OConnor & Eickwort, 1988: 172 Type locality: Odessa Distribution: Ukraine Crabrovidia latimanus (OConnor et Eickwort) Vidia latimanus OConnor & Eickwort, 1988: 160 Crabrovidia latimanus (OConnor et Eickwort): Type locality: Nova Scotia, Cape Breton Highlands National Park, Ingonish Beach Distribution: Canada Acarorum Catalogus IX 349 Host: Megachile (Xanthosaurus) latimanus (Hymenoptera: Megachilidae) Crabrovidia leclercqi Fain Crabrovidia leclercqi Fain, 1973: 179 Type locality: Yermasoyia Distribution: Cyprus Host: Cerceris rubida (Hymenoptera: Crabronidae) Crabrovidia ninae (Sevastianov) Vidia ninae Sevastianov, in Kadzhaya & Sevastianov 1967: 198 Crabrovidia ninae (Sevastianov): OConnor & Eickwort 1988: 172 Type locality: Kherson Prov. Distribution: Ukraine Host: Pseudovespa sp. (Hymenoptera: Vespidae) Crabrovidia oudemansi Fain Crabrovidia oudemansi Fain, 1971: 284; 1972: 244; 1973: 175; Nazan et al., 2019 Type locality: Corfu (Greece) Distribution: Greece (Corfu), Iran Host: Cerceris sp. (Hymenoptera: Crabronidae) Crabrovidia popovi (Zachvatkin) Vidia (Crabrovidia) popovi Zachvatkin, 1941: 375 Crabrovidia popovi (Zachvatkin): Fain, 1973: 161 Type locality: Krasnograd (Ukraine) Distribution: Abhasia, Croatia, Italy, Spain, Turkey, Ukraine, Uzbekistan Hosts: Lestica (Clypeocrabro) clypeata, L. (Ceratocolus) alata, L. (Clypeocrabro) wollmanni, Ectemnius impressus (Hymenoptera: Crabronidae) Crabrovidia rubi (OConnor et Eickwort) Vidia rubi OConnor & Eickwort, 1988: 148 Crabrovidia rubi (OConnor et Eickwort): Fain, 1971: 284 Type locality: Georgia, Liberty Co., St. Catherine‘s Island Distribution: USA (Georgia) Host: Megachile (Xeromegachile) rubi (Hymenoptera: Megachilidae) Crabrovidia texana (OConnor et Eickwort) Vidia texana OConnor & Eickwort, 1988: 162 Crabrovidia texana (OConnor et Eickwort): Type locality: New York, Albany Co., Rensselaerville, Edmund Niles Huyck Preserve Distribution: USA Host: Megachile (Litomegachile) texana (Hymenoptera: Megachilidae) 350 Acarorum Catalogus IX Crabrovidia thomasi (Delfinado et Baker) Vidia thomasi Delfinado & Baker, 1976: 81 (nom. nom. for V. cooremani Baker) Crabrovidia thomasi (Delfinado et Baker): OConnor & Eickwort 1988: 172 (nom. nov. for Vidia cooremani Baker, 1964 nec V. cooremani Thomas, 1961) Type locality: Plummers Island, Maryland Distribution: USA (Maryland) Crabrovidia utahensis (Delfinado et Baker) Vidia (Euvidia) utahensis Delfinado & Baker, 1976: 81 Crabrovidia utahensis (Delfinado et Baker): OConnor & Eickwort 1988: 172 Type locality: Beaver, Utah Distribution: USA Host: Hymenoptera (Sphecidae) Genus Ensliniella Vitzthum Ensliniella Vitzthum, 1925 [1924?]: 289 Type species: Ensliniella aegyptiana Baker et Cunliffe, 1960 Ensliniella aegyptiana Baker et Cunliffe Ensliniella aegyptiana Baker & Cunliffe, 1960: 214; Klompen & OConnor 1995: 126 Type locality: Abu Suttan Distribution: Egypt Hosts: Allodynerus koenigi, A. vinciquerrae (Hymenoptera: Vespidae, Eumeninae) Ensliniella asiatica Abe et Okabe Ensliniella asiatica Abe & Okabe, 2006: 929 Type locality: Sakurazawa, Yorii town, Osato District, Saitama Prefecture, central Honshu, 36°07’49”N, 139°12’28”E; 102 m Distribution: Japan Host: Allodynerus mandschuricus (Hymenoptera: Vespidae, Eumeninae) Ensliniella dignotus Klompen et OConnor Ensliniella dignotus Klompen & OConnor, 1995: 126 Type locality: Turkey Distribution: Turkey Host: Allodynerus dignotus (Hymenoptera: Vespidae, Eumeninae) Ensliniella floricola Klompen et OConnor Ensliniella floricola Klompen & OConnor, 1995: 127 Type locality: Pula (Croatia) Distribution: Croatia, Austria, Bulgaria, France, Greece, Holland, Turkey Hosts: Allodynerus floricola, A. rossii (Hymenoptera: Vespidae, Eumeninae) Acarorum Catalogus IX 351 Ensliniella koenigi Baker et Cunliffe Ensliniella koenigi Baker & Cunliffe, 1960: 216; Klompen & OConnor 1995: 127 Type locality: Ijonkak Distribution: Morocco Host: Allodynerus koenigi (Hymenoptera: Vespidae, Eumeninae) Ensliniella kostylevi Zakhvatkin Ensliniella kostylevi Zakhvatkin, 1941: 267; Klompen & OConnor, 1995 Type locality: Byalovezh (Belarus) Distribution: Belarus, Crimea, Holland, Algeria, Austria, Belgium, Bulgaria, France, Greece, Turkey Host: Allodynerus rossii (Hymenoptera: Vespidae, Eumeninae) Ensliniella parasitica Vitzthum Ensliniella parasitica Vitzthum, 1925: 289; Cooreman, 1942; Baker & Cunliffe, 1960: 214; Krivoluckiy, 1975: 476; Klompen & OConnor 1995: 128 Type locality: Germany (Bayern, Nürnberg) Distribution: Armenia, Italy, Germany, Holland, Slovakia, Uzbekistan, Belgium, Czech Rep., Spain, Gibraltar, Tadjikistan, Kyrghizstan, Kazakhstan Hosts: Allodynerus delphinalis, A. floricola (Hymenoptera: Vespidae, Eumeninae; Melittobia acasta (Eulophidae) Genus Kennethiella Cooreman Kennethiella Cooreman, 1954: 2 Type species: Ensliniella trisetosa Cooreman, 1942 Kennethiella trisetosa (Cooreman) Ensliniella trisetosa Cooreman, 1942: 1; Cooper, 1955 Kennethiella trisetosa (Cooreman): Cooreman, 1954; Baker & Cunliffe, 1960: 218 Type locality: USA Distribution: Finland, USA Host: Ancistrocerus antilope (Hymenoptera: Vespidae, Eumeninae) Genus Kurosaia Okabe et OConnor Kurosaia Okabe & OConnor, 2002: 69 Type species: Kurosaia jiju Okabe et OConnor, 2002 Kurosaia jiju Okabe et OConnor Kurosaia jiju Okabe & OConnor, 2002: 69 Distribution: Japan Host: Anterhynchium flavomarginatum (Hymenoptera, Vespidae) 352 Acarorum Catalogus IX Genus Macroharpa Mostafa Macroharpa Mostafa, 1970b: 395 Type species: Macroharpa amazonica Mostafa, 1970 Macroharpa amazonica Mostafa Macroharpa amazonica Mostafa, 1970b: 397 Type locality: Faro, Estado de Para Distribution: Brazil Host: Zethus biglumis ferrugineus (Hymenoptera: Vespidae, Zethinae) Macroharpa arota Mostafa Macroharpa arota Mostafa, 1970b: 399 Type locality: Amazonas, Manaus Distribution: Brazil Host: Zethus arotus (Hymenoptera: Vespidae, Zethinae) Macroharpa pachymapoda Mostafa Macroharpa pachymapoda Mostafa, 1970b: 398 Type locality: Obidos, Estado de Para Distribution: Brazil Host: Zethus lobulatus (Hymenoptera: Vespidae, Zethinae) Macroharpa palpognatha Mostafa Macroharpa palpognatha Mostafa, 1970b: 400 Type locality: Tefe Distribution: Brazil Host: Zethus roridus (Hymenoptera: Vespidae, Zethinae) Genus Monobiacarus Baker et Cunliffe Monobiacarus Baker & Cunliffe, 1960: 210 Type species: Monobiacarus quadridens Baker et Cunliffe, 1960 Monobiacarus funebris Baker et Cunliffe Monobiacarus funebris Baker & Cunliffe, 1960: 211 Type locality: Santa Rosa, Veracruz Distribution: Mexico Host: Monobia apicalipennis (Hymenoptera: Vespidae) Monobiacarus insularis Baker et Cunliffe Monobiacarus insularis Baker & Cunliffe, 1960: 212 Type locality: San Rafael, Jicoltepee Distribution: Mexico Acarorum Catalogus IX 353 Host: Monobia angulosa (Hymenoptera: Vespidae) Monobiacarus quadridens Baker et Cunliffe Monobiacarus quadridens Baker & Cunliffe, 1960: 211 Type locality: Kill Devis Hills, North Carolina Distribution: USA Host: Monobia quadridens (Hymenoptera: Vespidae) Genus Sphexicozela Mahunka Sphexicozela Mahunka, 1970: 77 Type species: Sphexicozela connivens Mahunka, 1970 Sphexicozela connivens Mahunka Sphexicozela connivens Mahunka, 1970: 77; Rusina &· Orlova, 2011: 685 Type locality: Budapest Distribution: Hungary, Ukraine, Crimea Hosts: Polistes dominula, P. gallica, P. nimpha (Hymenoptera: Vespidae, Polistinae) Genus Vespacarus Baker et Cunliffe Vespacarus Baker & Cunliffe, 1960: 220 Type species: Vespacarus rufovestis Baker et Cunliffe, 1960 Vespacarus anacardivorus Baker et Cunliffe Vespacarus anacardivorus Baker & Cunliffe, 1960: 224 Type locality: Paradise Key, Everglades National Park, Florida Distribution: USA (Florida) Host: Stenodynerus (Parancystrocerus) perennis anacardovora (Hymenoptera: Vespidae, Eumeninae) Vespacarus fulvipes Baker et Cunliffe Vespacarus fulvipes Baker & Cunliffe, 1960: 222 Type locality: Kill Devil Hills, North Carolina Distribution: USA Host: Stenodynerus (Parancystrocerus) f. fulvipes (Hymenoptera: Vespidae, Eumeninae) Vespacarus histrio Baker et Cunliffe Vespacarus histrio Baker & Cunliffe, 1960: 222 Type locality: Kill Devil Hills, North Carolina Distribution: USA Hosts: Stenodynerus (Parancystrocerus) histrio, S. (P.) f. fulvipes (Hymenoptera: Vespidae, Eumeninae) 354 Acarorum Catalogus IX Vespacarus minimoferus Mostafa Vespacarus minimoferus Mostafa, 1967: 618 Type locality: California Distribution: USA Host: Parancystrocercus minimoferus (Hymenoptera: Eumenidae) Vespacarus pedestris Baker et Cunliffe Vespacarus pedestris Baker & Cunliffe, 1960: 226 Type locality: Derby, New York Distribution: USA Host: Stenodynerus (Parancystrocerus) p. pedestris (Hymenoptera: Vespidae, Eumeninae) Vespacarus rufovestis Baker et & Cunliffe Vespacarus rufovestis Baker & Cunliffe, 1960: 220 Type locality: Lake Placid, Florida Distribution: USA Host: Stenodynerus (Parancystrocerus) fulvipes rufovestis (Hymenoptera: Vespidae, Eumeninae) Vespacarus saecularis Baker et Cunliffe Vespacarus saecularis Baker & Cunliffe, 1960: 224 Type locality: Lake Placid, Florida Distribution: USA Host: Stenodynerus (Parancystrocerus) saecularis (Hymenoptera: Vespidae, Eumeninae) Vespacarus tigris Baker et Cunliffe Vespacarus tigris Baker & Cunliffe, 1960: 228 Type locality: Arlington, Virginia Distribution: USA Host: Ancistrocerus t. tigris (Hymenoptera: Vespidae, Eumeninae) Vespacarus toltecus Baker et Cunliffe Vespacarus toltecus Baker & Cunliffe, 1960: 226 Type locality: Chisos Mts., Brewster Co., Texas Distribution: USA (Texas) Host: Stenodynerus (Parancystrocerus) toltecus (Hymenoptera: Vespidae, Eumeninae) Vespacarus vagus Baker et Cunliffe Vespacarus vagus Baker & Cunliffe, 1960: 228 Type locality: Toronto Distribution: Canada Host: Stenodynerus v. vagus (Hymenoptera: Vespidae, Eumeninae) Acarorum Catalogus IX 355 Genus Vidia Oudemans Vidia Oudemans, 1905: 22 [OConnor & Eickwort 1988] Type species: Vidia undulata Oudemans, 1905 Vidia lineata Oudemans Vidia lineata Oudemans, 1917: 391; Zachvatkin, 1941: 373; Buitendijk, 1945: 356; Fain, 1972: 240; Nazan et al., 2019 ? Vidia stoeckherti Türk & Türk, 1957: 152 – Type: Erlangen, Germany. Type locality: Bozen Distribution: Belgium, Croatia, Germany, Greece (Corfu), Holland, Iran, Italy, Russia Host: Megachile pilicrus (Hymenoptera: Megachilidae) Vidia undulata Oudemans Vidia undulata Oudemans, 1905: 22; Zachvatkin, 1941: 374; Fain, 1972: 239 Type locality: Maggianico Distribution: Italy Host: Hylaeus (Prosopis) conformis (Hymenoptera: Colletidae) Genus Zethacarus Mostafa Zethacarus Mostafa, 1971: 566 Type species: Zethacarus boharti Mostafa, 1971 Zethacarus andinus Mostafa Zethacarus andinus Mostafa, 1971: 572 Type locality: R.A. Cordoba, Totoral Distribution: Argentina Host: Zethus andinus (Hymenoptera: Vespidae, Zethinae) Zethacarus argentinae Mostafa Zethacarus argentinae Mostafa, 1971: 571 Type locality: Chaco de Santiago del Estro Bords du Rio Salado env. d’Icano Distribution: Argentina Host: Zethus haywardi (Hymenoptera: Vespidae, Zethinae) Zethacarus boharti Mostafa Zethacarus boharti Mostafa, 1971: 567 Type locality: Tamaulipas Distribution: Mexico Host: Zethus gurreroi mayorum (Hymenoptera: Vespidae, Zethinae) Zethacarus ellipsus Mostafa Zethacarus ellipsus Mostafa, 1971: 575 356 Acarorum Catalogus IX Type locality: Orizaba, Ver. Distribution: Mexico Host: Zethus orizabae (Hymenoptera: Vespidae, Zethinae) Zethacarus miscogaster Mostafa Zethacarus misco gaster Mostafa, 1971: 569 Type locality: 38 mi. W Jiquilpan Mich. Distribution: Mexico Host: Zethus miscogaster (Hymenoptera: Vespidae, Zethinae) Zethacarus paranicus Mostafa Zethacarus paranicus Mostafa, 1971: 574 Type locality: Parana Distribution: Brazil Host: Zethus smithii (Hymenoptera: Vespidae, Zethinae) Genus Zethovidia Mostafa, 1970 Zethovidia Mostafa, 1970a: 178 (subgenus of Vidia) Zethovidia Mostafa: Fain, 1972: 245 (genus) Type species: Vidia (Zethovidia) diminuta Mostafa, 1970 Zethovidia acaris (Mostafa) Vidia (Zethovidia) acaris Mostafa, 1970b: 385 Zethovidia acaris (Mostafa) Type locality: Maroni Distribution: French Guiana Host: Zethus carbonarius (Hymenoptera: Vespidae, Zethinae) Zethovidia arizonensis (Mostafa) Vidia (Zethovidia) arizonensis Mostafa, 1970a: 187 Zethovidia arizonensis (Mostafa) Type locality: San Simeon, Cochise Co., Arizona Distribution: USA Host: Zethus guerreroi arizonensis (Hymenoptera: Vespidae, Zethinae) Zethovidia brasilensis (Mostafa) Vidia (Zethovidia) brasilensis Mostafa, 1970a: 389 Zethovidia brasilensis (Mostafa) Type locality: Chapa Distribution: Brazil Host: Zethus coeruleopennis (Hymenoptera: Vespidae, Zethinae) Acarorum Catalogus IX 357 Zethovidia brevapodemis (Mostafa) Vidia (Zethovidia) brevapodemis Mostafa, 1970b: 387 Zethovidia brevapodemis (Mostafa) Type locality: Suriname Distribution: Suriname Host: Zethus magnus (Hymenoptera: Vespidae, Zethinae) Zethovidia compactilis (Mostafa) Vidia (Zethovidia) compactilis Mostafa, 1970a: 188 Zethovidia compactilis (Mostafa) Type locality: 3 mi. N.W. Petlalcingo, Puebla Distribution: Mexico Host: Zethus guerreroi guerreroi (Hymenoptera: Vespidae, Zethinae) Zethovidia diminuta (Mostafa) Vidia (Zethovidia) diminuta Mostafa, 1970a: 181 Zethovidia diminuta (Mostafa) Type loc/ality: San Bernardino Distribution: Brazil, Paraguay Host: Zethus diminutus diminutus, Z. diminutus santaremae (Hymenoptera: Vespidae, Zethinae) Zethovidia floridae (Mostafa) Vidia (Zethovidia) floridae Mostafa, 1970b: 392 Zethovidia floridae (Mostafa) Type locality: Royal Palm Park, Florida Distribution: USA Host: Zethus slossonae (Hymenoptera: Vespidae, Zethinae) Zethovidia fusca (Mostafa) Vidia (Zethovidia) fusca Mostafa, 1970b: 394 Zethovidia fusca (Mostafa) Type locality: Barro Colorado, Canal Zone Distribution: Panama Host: Zethus fuscus (Hymenoptera: Vespidae, Zethinae) Zethovidia laevapodemis (Mostafa) Vidia (Zethovidia) laevapodemis Mostafa, 1970a: 183 Zethovidia laevapodemis (Mostafa) 358 Acarorum Catalogus IX Type locality: 23 mi. S. Matias Romero, Oaxaca Distribution: Mexico Host: Zethus menkei (Hymenoptera: Vespidae, Zethinae) Zethovidia mexicana (Mostafa) Vidia (Zethovidia) mexicana Mostafa, 1970a: 184 Zethovidia mexicana (Mostafa) Type locality: 30 mi. W. Acayucan, Veracruz Distribution: Mexico Host: Zethus parkeri (Hymenoptera: Vespidae, Zethinae) Zethovidia oculabsensa (Mostafa) Vidia (Zethovidia) oculabsensa Mostafa, 1970a: 191 Zethovidia oculabsensa (Mostafa) Type locality: Chapada de Aunr Mato Grosso Distribution: Brazil Host: Zethus inconstans (Hymenoptera: Vespidae, Zethinae) Zethovidia paraguayensis (Mostafa) Vidia (Zethovidia) paraguayensis Mostafa, 1970a: 182 Zethovidia paraguayensis (Mostafa) Type locality: San Bernardino Distribution: Paraguay Host: Zethus diminutus diminutus (Hymenoptera: Vespidae, Zethinae) Zethovidia parvula (Mostafa) Vidia (Zethovidia) parvula Mostafa, 1970b: 392 Zethovidia parvula (Mostafa) Type locality: Corrientes Distribution: Argentina Host: Zethus parvulus (Hymenoptera: Vespidae, Zethinae) Zethovidia promina (Mostafa) Vidia (Zethovidia) promina Mostafa, 1970a: 388 Zethovidia promina (Mostafa) Type locality: Chapa Distribution: Brazil Host: Zethus prominens (Hymenoptera: Vespidae, Zethinae) Zethovidia reticulata (Mostafa) Vidia (Zethovidia) reticulata Mostafa, 1970b: 384 Acarorum Catalogus IX 359 Zethovidia reticulata (Mostafa) Type locality: 42 km SE Maturin Monagas Distribution: Venezuela Host: Zethus romandinus (Hymenoptera: Vespidae, Zethinae ) Zethovidia rugocoxa (Mostafa) Vidia (Zethovidia) rugocoxa Mostafa, 1970a: 189 Zethovidia rugocoxa (Mostafa) Type locality: 3 mi. N.W. Petlalcingo, Puebla Distribution: Mexico Host: Zethus guerreroi guerreroi (Hymenoptera: Vespidae, Zethinae ) Zethovidia salvadorensis (Mostafa) Vidia (Zethovidia) salvadorensis Mostafa, 1970a: 186 Zethovidia salvadorensis (Mostafa) Type locality: 4 mi. N. Quezaltepeque Distribution: El Salvador Host: Zethus irwini (Hymenoptera: Vespidae, Zethinae ) Subfamily Saproglyphinae Oudemans Saproglyphinae Oudemans, 1924 Type genus: Saproglyphus Berlese, 1890 Genus Acalvolia Fain Acalvolia Fain, 1971: 283 Type species: Vidia squamata Oudemans, 1909 Acalvolia americana Fan Qing-Hai, George & Kumarasinghe Acalvolia americana Fan Qing-Hai, George & Kumarasinghe, 2010: 42 Type locality: intercepted in New Zealand from oranges from the USA Distribution: USA Acalvolia squamata (Oudemans) Vidia squamata Oudemans, 1909: 317; Zachvatkin, 1941: 371 Acalvolia squamata (Oudemans): Fain, 1971: 283; 1972: 234; Fain & Knülle, 1981: 140; Barbosa & de Moraes, 2021 Type locality: Semarang (Java) Distribution: Germany, Holland, (?) Indonesia (probably an error), Brazil Host: Mus jerdoni (Rodentia: Muridae); house dust Genus Congovidia Fain et Elsen, in Fain, 1971 360 Acarorum Catalogus IX Congovidia Fain & Elsen, in Fain 1971: 282; Fain, Hurst, Tweddle, Lachlan, Majerus & Britt 1995: 100; see also Fain & Camerik, 1977 description of adult] Type species: Congovidia glossinae Fain, 1971 Congovidia benelux Fain, Hurst, Fassotte, Webberley, Sloggett et Majerus Congovidia benelux Fain, Hurst, Fassotte, Webberley, Sloggett & Majerus, 1997: 89 Type locality: Wageningen Distribution: Holland Host: Adalia bipunctata (Coleoptera: Coccinellidae) Congovidia brasiliensis Fain et Camerik Congovidia brasiliensis Fain & Camerik, 1977: 46; Fain, Hurst, Tweddle, Lachlan, Majerus & Britt 1995: 103 Type locality: Itatiaia (State of Rio de Janeiro) Distribution: Brazil Host: in the nest of Trypoxylon (Trypargilum) aestivale (Hymenoptera: Sphegidae) Congovidia coccinellidarum Fain, Hurst, Tweddle, Lachlan, Majerus et Britt Congovidia coccinellidarum Fain, Hurst, Tweddle, Lachlan, Majerus & Britt, 1995: 100 Type locality: England Distribution: England, Belgium Host: Halyzia sedecimguttata (Coleoptera: Coccinellidae) Congovidia glossinae Fain Congovidia glossinae Fain & Elsen , 1971: 282; 1972: 77; Fain, Hurst, Tweddle, Lachlan, Majerus & Britt, 1995: 103 Type locality: Luputa, Mabenga, Benzali Distribution: DR Congo Host: Glossina fuscipes quanzensis (Diptera: Glossinidae) Congovidia lomamiensis Fain et Elsen Congovidia lomamiensis Fain & Elsen, 1972: 81; Fain, Hurst, Tweddle, Lachlan, Majerus & Britt, 1995: 103 Type locality: Luputa Distribution: DR Congo Host: Glossina fuscipes quanzensis (Diptera: Glossinidae) Congovidia servalina Fain et Elsen Congovidia servalina Fain & Elsen, 1972: 83; Fain, Hurst, Tweddle, Lachlan, Majerus & Britt, 1995: 103 Type locality: Lundu, Mayumbe Distribution: DR Congo Acarorum Catalogus IX 361 Host: Glossina palpalis palpalis (Diptera: Glossinidae) Congovidia sp. Hybomitra epistates H. lasiophthalma, H. microcephala, H. nuda, H. sodalist, Tabanus lineata, T. reinwardii (Canada) Genus Gambacarus Mahunka Gambacarus Mahunka, 1975: 43 Type species: Gambacarus huettleri Mahunka, 1975 Gambacarus huettleri Mahunka Gambacarus huettleri Mahunka, 1975: 43 Type locality: Gambia Distribution: Gambia Genus Neocalvolia Hughes Neocalvolia Hughes, 1970: 153 Type species: Neocalvolia claggi Hughes, 1970 Neocalvolia claggi Hughes Neocalvolia claggi Hughes, 1970: 153 Type locality: grass, Bird I., South Georgia Distribution: South Georgia Neocalvolia coperminensis Rosas et Casanueva Neocalvolia coperminensis Rosas & Casanueva, 1993: 66 Type locality: South Shetlands Distribution: South Shetlands Neocalvolia inermis Fain et Schuster Neocalvolia inermis Fain & Schuster, 1983: 421 Type locality: Harrington Sound, Main Island Distribution: Bermudas Genus Neosuidasia Ranganath et Channabasa, in Ranganath, Channabasavanna & Krishna-Rao Neosuidasia Ranganath & Channabasa, in Ranganath, Channabasavanna & Krishna-Rao, 1983: 63 Type species: Neosuidasia faini Ranganath et Channabasa, in Ranganath, Channabasavanna & Krishna-Rao, 1983 362 Acarorum Catalogus IX Neosuidasia faini Ranganath et Channabasa, in Ranganath, Channabasavanna & Krishna-Rao Neosuidasia faini Ranganath & Channabasa, in Ranganath, Channabasavanna & KrishnaRao, 1983: 64 Type locality: unknown (India?) Distribution: on imported animal feed by Vegetal Quarantine of Cuba Genus Neottiglyphus Volgin Neottiglyphus Volgin, 1974: 223 Type species Neottiglyphus alatavicus Volgin, 1974 Neottiglyphus alatavicus Volgin Neottiglyphus alatavicus Volgin, 1974: 224 Type locality: Kazakhstan Distribution: Kazakhstan Genus Saproglyphus Berlese Saproglyphus Berlese, 1890: 6 Type species: Saproglyphus neglectus Berlese, 1890 (Diagnostic, Volgin 1980: 161) Saproglyphus cocciphagus Womersley Saproglyphus cocciphagus Womersley, 1941: 471 Type locality: Australia Distribution: Australia Saproglyphus neglectus Berlese Saproglyphus neglectus Berlese, 1890: 6 [Diagnostic, Volgin 1980: 161] Type locality: Sahalin Distribution: Russia (Sahalin), Italy Saproglyphus setosus Volgin Saproglyphus setosus Volgin, 1980: 159 Type locality: Yuzhno-Sakhalinsk (Sakhalin Island) Distribution: Russia (Far East) Host: on birch bark Saproglyphus waldorfae (Fain et Johnston) Calvolia waldorfae Fain & Johnston, 1974: 416 Saproglyphus waldorfae (Fain et Johnston): Khaustov et al., 2018: 53 Type locality: Domleo, Spinney, Sutton Bonington Acarorum Catalogus IX 363 Distribution: England, Russia (Tyumen Region) Host: Sminthurus fuscus (Collembola: Sminthuridae), galleries of Ips typographus (Coleoptera: Curculionidae, Scolytinae) Subfam. Oulenziinae Oudemans Oulenziidae: Oudemans, 1928 Type genus: Oulenzia Radford, 1950 Genus Oulenzia Radford Lenzia Oudemans, 1928: 327 [praeocc.!] Oulenzia Radford, 1950: 152; [syn. by Hughes 1962: 60; Type is arboricola; nomen nova for Lenzia?; restaured by Fan et al., 2012] Type species: Lenzia arboricola Oudemans, 1928 Oulenzia arboricola (Oudemans) Lenzia arboricola Oudemans, 1928: 327 Calvolia arboricola (Oudemans): Hughes, 1962: 61 Oulenzia arboricola (Oudemans): Radford, 1950: 222; Fan, George & Kumarasinghe, 2012: 334; Barbosa & de Moraes, 2021 Type locality: Medan, Deli, Sumatra Distribution: Indonesia (Sumatra), Brazil Host: Hevea leaves Genus Oulenziella Fan, George, Li et Zhang Oulenziella Fan, George, Li & Zhang, 2015: 191 Type species: Calvolia bakeri Hughes, 1962 Oulenziella bakeri (Hughes) Calvolia bakeri Hughes, 1962: 59 (as nom. nov.); Woodring, 1966: 80 Oulenzia bakeri (Hughes): Radford, 1950 Oulenziella bakeri (Hughes): Fan et al., 2015: 191; Barbosa & de Moraes, 2021 Oulenzia gossypii Meyer & Rodrigues, 1965: (Mozambique, Kaidundjua, on Gossypium sp., syn. by Fan et al., 2015) Type locality: India Distribution: India, Mozambique, Brazil, Indonesia (Sumatra) Oulenziella longiseta Barbosa et de Moraes Oulenziella longiseta Barbosa & de Moraes, 2021: Type locality: Brazil Distribution: Brazil 364 Acarorum Catalogus IX Genus Procalvolia Fain Procalvolia Fain, 1971: 283 Type species: Calvolia zacheri Oudemans, 1929 Procalvolia aviophilus Sevastianov et Hassan Procalvolia aviophilus Sevastianov & Hassan, 1989: 141 Type locality: Ukraine Distribution: Ukraine Host: Pica pica (Aves: Corvidae) Procalvolia coccinella Sevastianov et Hassan Procalvolia coccinella Sevastianov & Hassan, 1989: 141 Type locality: Ukraine Distribution: Ukraine Host: Coccinella (Coleoptera: Cocinellidae) Procalvolia hamadi Sevastianov et Hassan Procalvolia hamadi Sevastianov & Hassan, 1989: 141 Type locality: Egypt Distribution: Egypt Host: Labidura riparia (Dermaptera: Labiduridae) Procalvolia zacheri (Oudemans) Calvolia zacheri Oudemans, 1929: 36; Zachvatkin, 1941: 361 Calvolia romanovae Zachvatkin, 1941: 345; Hughes, 1962: 54 (syn. by Fain, 1972)(type from Moskow); Angelkova, 1988: Calvolia tarsoinfracta Türk & Türk, 1957: 167 (syn. with Calvolia romanovae by Hughes, 1962) Procalvolia zacheri (Oudemans): Fain, 1971: 283; 1972: 235; Rack, 1980: 304 Type locality: Berlin Distribution: Bulgaria, Germany, New Zealand (cheese), Russia Genus Psylloglyphus Fain Psylloglyphus Fain, 1966: 159 Type species: Psylloglyphus u. uilenbergi Fain, 1966 Subgenus Psylloglyphus (s. str.) Fain Psylloglyphus Fain, 1966: 159 Type species: Psylloglyphus u. uilenbergi Fain, 1966 Acarorum Catalogus IX 365 Psylloglyphus (Psylloglyphus) australiensis Fain, Bartholomaeus, Cooke et Beaucournu Psylloglyphus (Psylloglyphus) australiensis Fain, Bartholomaeus, Cooke & Beaucournu, 1990: 97 Type locality: Forrest (Victoria) Distribution: Australia Hosts: Pygiopsylla hoplia, Bibikovana rainbowi, Acanthopsylla r. rothschildi, Macropsylla hercules (Siphonaptera: Pygiopsyllidae) Psylloglyphus (Psylloglyphus) crenulatus Fain Psylloglyphus (Psylloglyphus) crenulatus Fain & Beaucournu, 1984: 100 Type locality: Ile de la Possession, Pointe Lieutard, alt. 50 m, Arch. Crozet Distribution: Arch. Crozet Psylloglyphus (Psylloglyphus) parapsyllus Fain et Galloway Psylloglyphus (Psylloglyphus) parapsyllus Fain & Galloway, 1993a: 160 Type locality: Motunau Island, Plateau Region Distribution: New Zealand Host: fleas in nests of Eudyptula minor and Pachyptila turtur Psylloglyphus (Psylloglyphus) maculatus Fain et Beaucournu Psylloglyphus (Psylloglyphus) maculatus Fain & Beaucournu, 1976: 184 Type locality: Ulugombak, Selangor, Malaysia Distribution: Malaysia Host: Medwayella robinsoni (Siphonaptera: Pygiopsyllidae) Psylloglyphus (Psylloglyphus) reticulatus Fain et Beaucournu Psylloglyphus reticulatus Fain & Beaucournu, 1976: 182 Type locality: Buhengere, Kivu, DR Congo Distribution: DR Congo Host: Ctenophthamus calceatus cabirus (Siphonaptera) Psylloglyphus (Psylloglyphus) u. uilenbergi Fain Psylloglyphus (Psylloglyphus) uilenbergi Fain, 1966: 162; Fain & Beaucournu, 1976: 181 Type locality: Madagascar, Iwato Distribution: Madagascar Hosts: Paractenophyllus petiti, Synopsyllus fonquerniei (Siphonaptera from tenrecs - Microgale and Setifer setosus) Psylloglyphus (Psylloglyphus) uilenbergi kivuensis Fain et Beaucournu Psylloglyphus (Psylloglyphus) uilenbergi kivuensis Fain & Beaucournu, 1976: 182; Fain & Schwan, 1976: 634 366 Acarorum Catalogus IX Type locality: Kivu (DR Congo) Distribution: DR Congo, Gabon, Kenya Hosts: Ctenophthalmus cabirus, Dinopsyllus apistus, D. echinus, D. lypusus, Listropsylla dolosa, Stivalius sitianus, Xiphiopsylla levis (Siphonaptera) Psylloglyphus (Psylloglyphus) vietnamensis Fain et Beaucournu Psylloglyphus (Psylloglyphus) vietnamensis Fain & Beaucournu, 1972: 529 Type locality: Dalat, South Vietnam Distribution: Vietnam Host: Stivalius aporus rectodigitus (Siphonaptera) from Rattus rattus (Rodentia) Subgenus Psylloglyphus (Hemimeropus) Fain et Beaucournu Psylloglyphus (Hemimeropus) Fain & Beaucournu, 1976: 184 Type species: Psylloglyphus (Hemimeropus) hemimerus Fain et Beaucournu, 1976 Psylloglyphus (Hemimeropus) hemimerus Fain et Beaucournu Psylloglyphus (Hemimerus) hemimerus Fain & Beaucournu, 1976: 184 Type locality: Butare Distribution: Rwanda Host: on Hemimerus bouvieri (Dermaptera: Hemimeridae), parasite on Cricetomys gambianus (Rodentia: Nesomyidae) Subgenus Psyllobia Fain, Bartholomaeus, Cooke et Beaucournu Psyllobia Fain, Bartholomaeus, Cooke et Beaucournu, 1990: Type species: Psylloglyphus (Psylloglyphus) foveolatus Fain et Mason, 1989 Psylloglyphus (Psyllobia) foveolatus Fain et Mason Psylloglyphus (Psylloglyphus) foveolatus Fain & Mason, 1989: 43; Fain et al., 1990: Type locality: 12 km S of Launceston, Tasmania Distribution: Australia (Tasmania, Victoria) Hosts: phoretic on Pygiopsylla hoplia from Rattus rattus (Muridae), Bibikovana rainbowi, Acanthopsylla r. rothschildi (Siphonaptera: Pygiopsyllidae) Subgenus Tetrapsyllopus Fain et Beaucournu Psylloglyphus (Tetrapsyllopus): Fain & Beaucournu, 1986: 257 Type species: Psylloglyphus (Tetrapsyllopus) micronychus Fain et Beaucournu, 1986 Psylloglyphus (Tetrapsyllopus) chiliensis Fain et Beaucournu Psylloglyphus (Tetrapsyllopus) chiliensis Fain & Beaucournu, 1989: 139 Type locality: Isla Quila (Chiloë) Distribution: Chile Acarorum Catalogus IX 367 Hosts: Sphinctopsylla ares (Stephanocircidae), Ctenoparia inopinata (Hystrichopsyllidae) (Siphonaptera) Psylloglyphus (Tetrapsyllopus) micronychus Fain et Beaucournu Psylloglyphus (Tetrapsyllopus) micronychus Fain & Beaucournu, 1986: 258 Type locality: Galletué (Prov. Malleco) Distribution: Chile Genus Trypetacarus Fain Trypetacarus Fain, 1971: 283 Type species: Calvolia trypetidarum Cooreman, 1942 Trypetacarus trypetidarum (Cooreman) Calvolia trypetidarum Cooreman, 1942: 1 Trypetacarus trypetidarum (Cooreman): Fain, 1971: 283 Type locality: Kutu. District des Bangala Distribution: DR Congo Host: Celidodacus fenestratus (Diptera: Trypetidae) Subfamily Winterschmidtiinae Oudemans Winterschmidtiidae Oudemans, 1923: 206 Calvoliinae Türk & Türk, 1957: 155 – Type: Calvolia Oudemans, 1911 Type genus: Winterschmidtia Oudemans, 1923 Genus Allocalvolia Fain et Rack Allocalvolia Fain & Rack, 1987: 310 Type species: Allocalvolia habrocytus Fain et Rack, 1987 Allocalvolia bisculpturata (Mahunka) Calvolia bisculpturata Mahunka, 1974a: 137; Eraky 1998: 248 Allocalvolia bisculpturatus (Mahunka): Fain & Rack 1987: 311 Type locality: Amedzofe Distribution: Ghana Allocalvolia habrocytus Fain et Rack Allocalvolia habrocytus Fain & Rack, 1987: 311 Type locality: Eastern Austria Distribution: Austria Host: Habrocytus elevatus (Hymenoptera: Pteromalidae) 368 Acarorum Catalogus IX Genus Calvolia Oudemans Calvolia Oudemans, 1911: 187 Type species: Calvolia hagensis Oudemans, 1911 Calvolia antarctica Hughes Calvolia antarctica Hughes, 1966: 45 Type locality: South Sandwich Islands Distribution: South Sandwich Islands Calvolia astoma Türk et Türk Calvolia astoma Türk & Türk, 1957: 160; Hughes, 1962: 61 Type locality: Erlangen (Germany) Distribution: Germany, Slovakia Hosts: Formica rufa, Lasius fuliginosus (Hymenoptera: Formicidae) Calvolia bulgarica Štorkan Calvolia bulgarica Štorkan, 1935: 22; Zachvatkin, 1941: 257; Hughes, 1962: 61 Type locality: Bulgaria, Botev Summit (= Yumrukchal) Distribution: Bulgaria, Russia Host: Aphthona euphorbiae (Coleoptera: Chrysomelidae, Alticinae) Calvolia calliphorae Vitzthum Calvolia calliphorae Vitzthum, 1922: 539; Zachvatkin, 1941: 256; Hughes, 1962: 61 Type locality: Weimar Distribution: Germany Host: Calliphora vomitoria (Diptera: Calliphoridae) Calvolia domicola Oshima Calvolia domicola Oshima, 1979: 240 Type locality: Minami-ku, Yokohama Distribution: Japan Host: house dust Calvolia elisabethae Zachvatkin Calvolia elisabethae Zachvatkin, 1941: 254 Type locality: Staraya Bukhara (Uzbekistan) Distribution: Georgia, Uzbekistan Hosts: Ellobius talpinus, Prometheomys shaposhnikovi (Mammalia: Rodentia) Calvolia elongata Volgin Calvolia elongata Volgin, 1980: 167 Acarorum Catalogus IX 369 Type locality: Alma-Atinskaya Obl., Kaskelenskiy Rayon Distribution: Kazakhstan Host: manure Calvolia fraxini Türk et Türk Calvolia fraxini Türk & Türk, 1957: Type locality: Munich Distribution: Germany Host: Leperisinus orni (Coleoptera: Curculionidae) Calvolia furnissi Woodring Calvolia furnissi Woodring, 1966: 76 Type locality: Moskow, Idaho Distribution USA Host: from the galleries and frass: “from the galleries and frass therein of Dendroctonus pseudotsugae in Douglas fir” Calvolia fusiformis Zachvatkin Calvolia fusiformis Zachvatkin, 1941: 259; Hughes, 1962: 61 Type locality: Klinskiy Rayon, Moskovskaya Obl. Distribution: Russia Host: Blastophagus minor (Coleoptera: Curculionidae, Scolytinae) Calvolia glabra Womersley Calvolia glabra Womersley, 1941: 474 Type locality: Adelaide Zoo Distribution: Australia Host: “from a branchium of a Boa” Calvolia goetzae Türk et Türk Calvolia goetzae Türk & Türk, 1957: 163; Hughes, 1962: 61 Type locality: Moskow Distribution: Russia, Slovakia Hosts: Aphodius minor, A. prodromus (Coleoptera: Scarabaeidae) Calvolia hagensis Oudemans Calvolia hagensis Oudemans, 1911: 187; Vitzthum, 1920: 58; Zachvatkin, 1941: 359; Hughes, 1962: 61; Fain, 1972: 229 Type locality: Hague Distribution: Holland 370 Acarorum Catalogus IX Calvolia hebeclinii (Sicher) Glycyphagus hebeclinii Sicher, in Canestrini, 1899: 971 Calvolia hebeclinii (Sicher): Zachvatkin, 1941: 253; Hughes, 1962: 61 Type locality: Sicily Distribution: Italy (Sicily) Host: on plants Calvolia heterocoma (Michael) Tyroglyphus heterocomus Michael, 1903: 106 Czenspinskia heterocoma: Oudemans, 1927: Calvolia heterocoma (Michael): Zachvatkin, 1941: 248; Hughes, 1962: 61 Type locality: New Forest, Hampshire Distribution: England, Bulgaria, Slovakia Calvolia kamenskii Zachvatkin Calvolia kamenskii Zachvatkin, 1941: 254; Hughes, 1962: 61 Type locality: Bashkortistan Distribution: Russia Calvolia longipes Türk et Türk Calvolia longipes Türk & Türk, 1957: 170 Type locality: Erlangen (Germany) Distribution: Germany, Slovakia Host: Chelidura acanthopygia (Dermaptera: Forficulidae) Calvolia longireticulatae Ramaraju et Mohanasundaram Calvolia longireticulatae Ramaraju & Mohanasundaram, 1999: 47 Type locality: Chidambaram, Tamil Nadu Distribution: India Host: Coleoptera, Meloidae gen. sp. Calvolia marie Samšinák Calvolia marie Samšinák, 1989: 105 Distribution: DR Congo Host: Gymnometopina magna (Diptera: Sphaeroceridae) Calvolia minima Türk et Türk Calvolia minima Türk & Türk, 1957: 170; Hughes, 1962: 61 Type locality: Erlangen Distribution: Germany Acarorum Catalogus IX 371 Host: Scolytus intricatus (Coleoptera: Curculionidae, Scolytinae) Calvolia miniopteri Dusbábek Calvolia miniopteri Dusbábek, 1964: 228 Type locality: Ciertova diera Distribution: Slovakia Host: Miniopterus schreibersi (Chiroptera: Miniopteridae) Calvolia priapus Volgin Calvolia priapus Volgin, 1980: 161; Faradonbeh et al., 2018: 12 Type locality: Semirabad (Azerbaidzhan) Distribution: Azerbaijan, Iran, Ukraine (Odessa) Host: bird’s nests Calvolia reticulata Zachvatkin Calvolia reticulata Zachvatkin, 1941: 254; Hughes, 1962: 61 Type locality: Sussillon (Switzerland) Distribution: Slovakia, Switzerland Host: Osmia (Anthocopa) dalmatina (Hymenoptera: Calvolia romanovae Zachvatkin Calvolia romanovae Zachvatkin, 1941: 252 Calvolia tarsoinfracta Türk & Türk, 1957: 167 (syn. by Hughes, 1962) Type locality: Moskow (Russia) Distribution: Russia, Germany, New Zealand Calvolia ruehmi Türk et Türk Calvolia ruehmi Türk & Türk, 1957: 162; Hughes, 1962: 61; Woodring, 1966: 80 Type locality: Erlangen Distribution: Germany Host: Dryocoetes villosus (Coleoptera: Curculionidae, Scolytinae) Calvolia solimani Eraky Calvolia solimani Eraky, 1999: 215 Type locality: Aswan, Botanical Island Distribution: Egypt Host: Phoretic on Amitermes desertorum (Insecta) Calvolia striata Vitzthum Calvolia striata Vitzthum, 1914: 324; Zachvatkin, 1941: 260; Hughes, 1962: 61 Type locality: Ivory Coast 372 Acarorum Catalogus IX Distribution: Italy (?), Ivory Coast Hosts: Taphrorhynchus sp. (?), Pityogenes lepidus (Coleoptera: Curculionidae, Scolytinae) Calvolia summersi Mostafa Calvolia summersi Mostafa, 1971: 577 Type locality: Amazonas, Manaus Distribution: Brazil Host: Zethus binodis (Hymenoptera: Vespidae, Eumeninae) Calvolia thraca Vitzthum Calvolia thraca Vitzthum, 1922: 542; Zachvatkin, 1941: 255; Hughes, 1962: 61 Type locality: Bulgaria, Sofia Distribution: Bulgaria, Russia Host: Diptera Calvolia tuberculata Zachvatkin Calvolia tuberculata Zachvatkin, 1941: 248; Hughes, 1962: 61; Woodring, 1966: 80 Type locality: Georgia Distribution: Georgia, Slovakia Calvolia wichmanni Türk et Türk Calvolia wichmanni Türk & Türk, 1957: 164 Type locality: Erlangen Distribution: Germany Host: Pyrrhidium sanguineum (Coleoptera: Cerambycidae) Calvolia zachvatkini Volgin Calvolia zachvatkini Volgin, 1980: 165 Type locality: Russia (Orichevsky Rayon, Kirovskaya Obl.) Distribution: Kazakhstan, Russia Host: hay and rodent’s nests Calvolia zacheriana nom. nov. Eraky Calvolia zaheri Eraky, 1998: 246; 1999: 217 Type locality: 47 km North of El-Kharga city, New Valley Distribution: Egypt Host: Psammotermes hypostoma (Isoptera: Rhinotermitidae) Calvolia sp. Hosts: Hybomitra affinis, H. epitates, H. lasiophthalma, H. nuda, Tabanus atratus (Canada) Acarorum Catalogus IX 373 Genus Czenspinskia Oudemans Genus Czenspinskia Oudemans, 1923 [syn. with Calvolia by Hughes 1962: 60; separate by OConnor] Donndorffia Oudemans, 1931: 203; [syn. by Hughes 1962: 60; Type is transversostriata] Type species: Tyroglyphus heterocomus Michael, 1903 Czenspinskia transversostriata (Oudemans) Donndorffia transversostriata Oudemans, 1931: 203 Czenspinskia transversostriata (Oudemans): Ripka et al., 2002: 278; Barbosa & de Moraes, 2021 Czenspinskia lordi Nesbitt, 1946: syn. by Hughes, 1962 Calvolia transversostriata (Oudemans): Hughes 1962: 48 Type locality: “Mares niet gevonen” Distribution: Croatia, Holland, France, Hungary, Italy, Switzerland, Spain, Canada, USA, Brazil, Australia Genus Parawinterschmidtia Khaustov Parawinterschmidtia Khaustov, 2000: 57 Type species: Calvolia kneissli Krausse, 1919 Parawinterschmidtia kneissli (Krausse) Calvolia kneissli Krausse, 1919: 123; Zachvatkin, 1941: 255; Türk & Türk, 1957: 157; Samšinák, 1957; Hughes, 1962: 61; Cooreman, 1963: 24; Woodring, 1966: 80 Parawinterschmidtia kneissli (Krausse): Khaustov, 2000: 57; Khaustov et al., 2018: 49 Type locality: Germany Distribution: Belgium, Germany; “Czechoslovakia”, Poland, Crimea, Russia, Slovakia Hosts: Lithobius forficatus (Chilopoda, Lithobiomorpha: Lithobiidae); Raphidia sp. (larva) (Raphidioptera: Raphidiidae); Ips [Orthomicus] laricis, Brachyderes incanus Hylobius abietis, Pissodes pini, P. piniphilus (Coleoptera: Curculionidae, Scolytinae), Thanasimus formicarius (Coleoptera: Cleridae); Mesosa nebulosi, Rhagium sycophanta, R. inquisitor, Plagionotus detritus, Monochamus galloprovincialis, Spondylis buprestoides (Col.: Cerambycidae), Crypticus quisquilius (Col.: Tenebrionidae), Myzia oblongoguttata (Col.: Coccinellidae), Acanthocinus aedilis, Spondylis buprestoides, Sericus brunneus (Elateridae) Tomicus piniperda, Hylastes attenuates, Crossocerus vagabundus, C. leucostoma, Ectemnius cavifrons, E. cephalotes (Crabronidae), Sceliphron deforme (Sphecidae), Astata kashmirensis (Astatinae) Parawinterschmidtia sp. – Finland (Penttinen, Viiri & Moser, 2013) 374 Acarorum Catalogus IX Genus Riemia Oudemans Riemia Oudemans, 1925: 90 Type species: Riemia hesperidum Oudemans, 1925 Riemia hesperidum Oudemans Riemia hesperidum Oudemans, 1925: 90 Type locality: Arnhem Distribution: Holland Genus Winterschmidtia Oudemans Winterschmidtia Oudemans, 1923: 207 Afrocalvolia Fain & Elsen, 1971: 281 [Type is glossinarum; Syn. Khaustov 2000: 50] Type species of Winterschmidtia Oudemans: Suidasia hamadryas Vitzthum, 1923 Winterschmidtia athiasae (Fain) Afrocalvolia athiasae Fain, 1974: 70 Type locality: La Savane de Lamto Distribution: Ivory Coast Winterschmidtia brenyi (Cooreman) Calvolia brenyi Cooreman, 1963: 32 Winterschmidtia brenyi (Cooreman): Khaustov 2000: 53 Type locality: Gembloux, Prov. de Namur) Distribution: Belgium, Crimea Host: Pteleobius vittatus (Coleoptera: Curculionidae, Scolytinae) Winterschmidtia chaetoptelii Khaustov Winterschmidtia chaetoptelii Khaustov, 2000: 55 Type locality: Crimea, Alushta Distribution: Crimea Host: Phoretic on Chaetoptelius vestitus (Coleoptera: Curculionidae) Winterschmidtia circumspectans (Vitzthum) Calvolia circumspectans Vitzthum, 1920: 56; Zachvatkin, 1941: 258; Hughes, 1962: 61 Afrocalvolia circumspectans (Vitzthum): Fain, 1974: 70 Type locality: Himalaya, Nilam-Dshong, NW Abhang .des Gaurisankar Distribution: Tibet Host: Ips stebbingi (Coleoptera: Curculionidae, Scolytinae) Winterschmidtia crassisetosa Willmann Winterschmidtia crassisetosa Willmann, 1941: 65 Acarorum Catalogus IX 375 Type locality: Italy Distribution: Italy Host: Phloeotribus scarabaeoides (Coleoptera: Curculionidae, Scolytinae) Winterschmidtia elongata Türk et Türk Winterschmidtia elongata Türk & Türk, 1957: 154 Type locality: Erlangen Distribution: Germany Winterschmidtia glossinarum (Fain et Elsen) Afrocalvolia glossinarum Fain & Elsen, in Fain 1971: 281; Fain & Elsen, 1972: 74 Winterschmidtia glossinarum (Fain et Elsen): [Khaustov, 2000] Type locality: Benzali nr. Kinshasa Distribution: DR Congo Hosts: Glossina fuscipes quanzensis, G. palpalis palpalis (Diptera: Glossinidae) Winterschmidtia hamadryas (Vitzthum) Suidasia hamadryas Vitzthum, 1923: 156 Winterschmidtia hamadryas (Vitzthum): Oudemans, 1923: 206; 1927: 246; Zachvatkin, 1941: 240; Khaustov, 2000: 51; Ahadyiat & Ostovan, 2005 Type locality: Germany Distribution: Germany, Crimea, Iran Host: burrows of Scolytus rugulosus; on Scolytus amyggdali (Coleoptera: Curculionidae, Scolytinae) Winterschmidtia lamtoensis (Fain) Afrocalvolia lamtoensis Fain, 1974: 72 Type locality: La Savane de Lamto Distribution: Ivory Coast Winterschmidtia nataliae (Zachvatkin) Calvolia nataliae Zachvatkin, 1941: 262; Cooreman, 1963: 28 Calvolia elliptica Zachvatkin, 1941: 262; Hughes, 1962: 61 – Type: Sviblovo near Moskow, Russia, on Ips acuminatus (Coleoptera: Curculionidae, Scolytinae) Calvolia fraxini Turk & Turk, 1957: 1957: 166; Hughes, 1962: 61 (syn. by Cooreman, 1963) Winterschmidtia nataliae (Zachvatkin): Khaustov, 2000: 52 Type locality: Alekseevskoe Village near Moskow Distribution: Belgium, Germany, Poland, Russia, Crimea Hosts: Leperesinus [Hylesinus] fraxini, L. omi (Coleoptera: Curculionidae, Scolytinae) Winterschmidtia tsetse (Fain et Elsen) 376 Acarorum Catalogus IX Afrocalvolia tsetse Fain & Elsen, 1972: 75 Type locality: Kaniama, Lomami Distribution: DR Congo Hosts: Glossina fuscipes quanzensis (Diptera: Glossinidae) Winterschmidtia villifronsi Khaustov Winterschmidtia villifronsi Khaustov, 2000: 56 Type locality: Crimea, Yaltinskiy Gornolesnoy reserve Distribution: Crimea Host: Phoretic on Taphrorychus villifrons (Coleoptera) Winterschmidtia zachvatkini Khaustov Winterschmidtia zachvatkini Khaustov, 2000: 57 Type locality: Yalta Distribution: Crimea Host: Phoretic on Pityogenes calcaratus (Coleoptera: Curculionidae, Scolytinae) Acarorum Catalogus IX 377 Index of Winterschmidtiidae (Names in bold are accepted as valid) Acalvolia 359 acaris, Vidia (Zethovidia) 356 acaris, Zethovidia 356 aegyptiana, Ensliniella 350 africana, Crabrovidia 346 Afrocalvolia 374 alatavicus, Neottiglyphus 362 Allocalvolia 367 amazonica, Macroharpa 352 americana, Acalvolia 359 anacardivorus, Vespacarus 353 andinus, Zethacarus 355 antarctica, Calvolia 368 arboricola, Calvolia 363 arboricola, Lenzia 363 arboricola, Oulenzia 363 argentinae, Zethacarus 355 arizonensis, Vidia (Zethovidia) 256 arizonensis, Zethovidia 356 arota, Macroharpa 352 asiatica, Ensliniella 350 astoma, Calvolia 368 athiasae, Afrocalvolia 374 athiasae, Winterschmidtia 374 australiensis, Psylloglyphus 365 aviophilus, Procalvolia 364 bakeri, Calvolia 363 bakeri, Oulenzia 363 bakeri, Oulenziella 363 benelux, Congovidia 360 bisculpturata, Calvolia 367 bisculpturata, Allocalvolia 367 bisculpturatus, Allocalvolia 367 boharti, Zethacarus 355 brasiliensis, Congovidia 360 brasiliensis, Crabrovidia 347 brasilensis, Vidia (Zethovidia) 356 brasilensis, Zethovidia 356 brenyi, Calvolia 375 brenyi, Winterschmidtia 375 brevapodemis, Vidia (Zethovidia) 357 brevapodemis, Zethovidia 357 bulgarica, Calvolia 368 calliphorae, Calvolia 368 Calvolia 368 Calvoliinae 346 chaetoptelii, Winterschmidtia 375 chiliensis, Psylloglyphopus 366 circumspectans, Afrocalvolia 375 circumspectans, Calvolia 375 circumspectans, Winterschmidtia 375 claggi, Neocalvolia 361 coccinella, Procalvolia 364 coccinellidarum, Congovidia 360 cocciphagus, Saproglyphus 362 compactilis, Vidia (Zethovidia) 357 compactilis, Zethovidia 357 concellaria, Crabrovidia 347 concellaria, Vidia 347 Congovidia 360 connivens, Sphexicozela 353 cooremani, Vidia 350 coperminensis, Neocalvolia 361 corsicana, Crabrovidia 347 Crabrovidia 346 crassisetosa, Winterschmidtia 375 crenulatus, Psylloglyphus 365 crossocerus, Crabrovidia 347 cubensis, Crabrovidia 347 Czenspinskia 373 Czenspinskiidae 346 dasyproctus, Crabrovidia 347 378 Acarorum Catalogus IX dignotus, Ensliniella 350 diminuta, Vidia (Zethovidia) 257 diminuta, Zethovidia 357 domicola, Calvolia 367 Donndorffia 373 ectemnii, Crabrovidia 348 elisabethae, Calvolia 368 ellipsus, Zethacarus 355 elliptica, Calvolia 375 elongata, Calvolia 368 elongata, Winterschmidtia 375 Ensliniella 350 Ensliniellinae 346 Euvidia 346 faini, Neosuidasia 359 floricola, Ensliniella 350 floridae, Vidia (Zethovidia) 357 floridae, Zethovidia 357 foveolatus, Psylloglyphus 366 fraxini, Calvolia 369 fulvipes, Vespacarus 353 funebris, Monobiacarus 352 furnissi, Calvolia 369 fusca, Vidia (Zethovidia) 357 fusca, Zethovidia 357 fusiformis, Calvolia 369 Gambacarus 361 glabra, Calvolia 369 glossinae, Congovidia 360 glossinarum, Afrocalvolia 375 glossinarum, Winterschmidtia 375 goetzae, Calvolia 369 gossypii, Oulenzia 363 gussakovskii, Crabrovidia 348 gussakovskii, Vidia (Crabrovidia) 348 habrocytus, Allocalvolia 367 hagensis, Calvolia 369 hamadi, Procalvolia 364 hamadryas, Suidasia 375 hamadryas, Winterschmidtia 375 hebeklinii, Calvolia 370 hebeklinii, Glycyphagus 370 Hemimeropus 366 hemimerus Psylloglyphus 366 hesperidum, Riemia 374 heterocoma, Calvolia 370 heterocoma, Czenspinskia 370 heterocomus, Tyroglyphus 370 hirsuta, Crabrovidia 348 hirsuta, Vidia 348 histrio, Vespacarus 353 huettleri, Gambacarus 361 inermis, Neocalvolia 361 insularis, Monobiacarus 352 jiju, Kurosaia 351 juliani, Crabrovidia 348 juliani, Vidia 348 kamenskii, Calvolia 367 Kennethiella 351 kivuensis, Psylloglyphus uilenbergi 362 kneissli, Calvolia 373 kneissli, Parawinterschmidtia 373 koenigi, Ensliniella 351 kostylevi, Ensliniella 351 Kurosaia 351 laevapodemis, Vidia (Zethovidia) 357 laevapodemis, Zethovidia 357 lamtoensis, Afrocalvolia 375 lamtoensis, Winterschmidtia 375 latimanus, Crabrovidia 348 latimanus, Vidia 348 leclercqi, Crabrovidia 349 Lenzia (praeoccup.) 363 lineata, Vidia 355 lomamiensis, Congovidia 360 longipes, Calvolia 370 longireticulatae, Calvolia 370 longiseta, Oulenziella 363 lordi, Czenspinskia 373 Macroharpa 352 maculatus, Psylloglyphus 365 marie, Calvolia 370 mexicana, Vidia (Zethovidia) 358 mexicana, Zethovidia 358 Acarorum Catalogus IX 379 micronychus, Psylloglyphus (Tetrapsyllopus) 367 minimoferus, Vespacarus 354 miniopteri, Calvolia 371 miscogaster, Zethacarus 355 Monobiacarus 352 nataliae, Calvolia 375 nataliae, Winterschmidtia 375 neglectus, Saproglyphus 362 Neocalvolia 361 Neosuidasia 361 Neottiglyphus 362 ninae, Crabrovidia 349 ninae, Vidia 349 oculabsensa, Vidia (Zethovidia) 358 oculabsensa, Zethovidia 358 oudemansi, Crabrovidia 349 Oulenzia 363 Oulenziella 363 Oulenziidae 346 Oulenziinae 363 pachymapoda, Macroharpa 352 palpognatha, Macroharpa 352 paraguayensis, Vidia (Zethovidia) 358 paraguayensis, Zethovidia 358 paranicus, Zethacarus 356 parapsyllus, Psylloglyphus 365 parasitica, Ensliniella 351 Parawinterschmidtia 373 parvula, Vidia (Zethovidia) 358 parvula, Zethovidia 358 pedestris, Vespacarus 351 popovi, Vidia (Crabrovidia) 349 popovi, Crabrovidia 349 priapus, Calvolia 371 Procalvolia 364 promina, Vidia (Zethovidia) 358 promina, Zethovidia 358 Psyllobia 366 Psylloglyphus 364 quadridens, Monobiacarus 353 reticulata, Calvolia 371 reticulata, Vidia (Zethovidia) 358 reticulata, Zethovidia 358 reticulatus, Psylloglyphus 365 Riemia 374 romanovae, Calvolia 364, 371 rubi, Crabrovidia 349 rubi, Vidia 349 ruehmi, Calvolia 371 rufovestis, Vespacarus 354 rugocoxa, Vidia (Zethovidia) 359 rugocoxa, Zethovidia 359 saecularis, Vespacarus 354 salvadorensis, Vidia (Zethovidia) 359 salvadorensis, Zethovidia 359 Saproglyphidae 346 Saproglyphinae 359 Saproglyphus 362 servalina, Congovidia 360 setosus, Saproglyphus 362 solimani, Calvolia 371 Sphexicozela 353 squamata, Acalvolia 359 squamata, Vidia 359 stoeckherti, Vidia 355 striata, Calvolia 371 summersi, Calvolia 372 tarsoinfracta, Calvolia 364 Tetrapsyllopus 366 texana, Crabrovidia 349 texana, Vidia 349 thomasi, Crabrovidia 350 thomasi, Vidia 350 thraca, Calvolia 372 tigris, Vespacarus 354 toltecus, Vespacarus 354 transversostriata, Calvolia 373 transversostriata, Czenspinskia 373 transversostriata, Donndorffia 370 trisetosa, Ensliniella 351 trisetosa, Kennethiella 351 Trypetacarus 367 trypetidarum, Calvolia 367 380 Acarorum Catalogus IX trypetidarum, Trypetacarus 367 tsetse, Afrocalvolia 376 tsetse, Winterschmidtia 376 tuberculata, Calvolia 372 uilenbergi, Psylloglyphus 365 undulata, Vidia 355 utahensis, Crabrovidia 350 utahensis, Vidia (Euvidia) 350 vagus, Vespacarus 354 Vidia 355 vietnamensis, Psylloglyphus 366 villifronsi, Winterschmidtia 376 waldorphae, Calvolia 362 waldorphae, Saproglyphus 362 wichmanni, Calvolia 372 Winterschmidtia 374 Winterschmidtiidae 346 Winterschmidtiinae 367 zachvatkini, Calvolia 372 zachvatkini, Winterschmidtia 376 zaheri, Calvolia Oud. 364 zaheri, Calvolia Eraky (praeoccup.) 372 zaheri, Procalvolia 364 zaheriana, Calvolia 372 (nom. nov.) Zethacarus 352 Zethovidia 353 Distribution of Winterschmidtiidae EUROPE Armenia (Klompen & OConnor, 1995) - Ensliniella parasitica Austria (Fain, 1973; Fain & Rack, 1987; Klompen & OConnor, 1995; Moser et al., 2005) - Allocalvolia habrocytus, Crabrovidia ectemnii, Ensliniella kostylevi Azerbaidjan (Volgin, 1980) - Calvolia priapus Belarus (Vitzthum, 1925) – Ensliniella kostylevi Belgium (Cooreman, 1948, 1963; Fain, 1972, 1973; Fain, Hurst et al. 1997; Klompen & OConnor, 1995) – Congovidia coccinellidarum, Crabrovidia concellaria, C. ectemnii, Ensliniella floricola, E. kostylevi Winterschmidtia brenyi, W. nataliae Bulgaria (Vitzthum, 1922; Štorkan, 1935; Angelkova, 1988; Klompen & OConnor, 1995) – Calvolia bulgarica, C. heterocoma, C. thraca, C. tuberculata, Ensliniella kostylevi, Procalvolia zacheri (= Calvolia romanovae) Croatia (Fain, 1972, 1973; Klompen & OConnor, 1995; Ripka et al., 2002; ) – Crabrovidia ectemnii, C. popovi, Czenspinskia transversostriata, Ensliniella floricola Finland (Penttinen, Viiri & Moser, 2013; Linnakoski et al., 2016) – Probonomoia pini France (Fain, 1972, 1973; Klompen & OConnor, 1995) – Crabrovidia concellaria, C. ectemnii, Ensliniella floricola, Е. kostylevi Corsica (Fain, 1973) – Crabrovidia corsicana Georgia (Zachvatkin, 1941) – Calvolia elisabethae, C. tuberculata Germany (Vitzthum, 1922, 1923, 1925; Türk & Türk, 1957) – Calvolia calliforae,C. fraxini, C. minima, C. romanovae (= tarsoinfracta), Ensliniella parasitica, Procalvolia zacheri, Winterschmidtia hamadryas, W. nataliae Great Britain (Fain & Johnston, 1974; Fain, Hurst et al., 1997; Fain & Greenwood, 1991) – Calvolia waldorfae, Congovidia coccinellidarum, Probonomoia pini Greece (Klompen & OConnor, 1995) - Ensliniella floricola, Е. kostylevi Acarorum Catalogus IX 381 Crete (Fain, 1973) - Crabrovidia ectemnii Corfu (Fain, 1971) – Crabrovidia oudemansi, Vidia lineata Holland (Oudemans, 1909, 1911, 1917, 1925, 1931; Klompen & OConnor, 1995) – Acalvolia squamata, Calvolia hagensis, Czenspinskia transversostriata, Ensliniella kostylevi, E. parasitica, Myianoetus dionychus, M. simplex, Riemia hesperidum, Vidia lineata Hungary (Mahunka, 1970; Ripka et al, 2002) - Sphexicozela connivens, Czenspinskia transversostriata Italy (Oudemans, 1905; Fain, 1973; Baker & Cunliffe, 1960) – Calvolia hebeklinii, C. striata, Crabrovidia popovi, Czenspinskia transvertostriata, Ensliniella parasitica, Saproglyphus neglectus, Vidia lineata, V. undulata Sicily (Sicher, in Canestrini, 1899) – Calvolia hebeclinii Sardinia - Calvolia hebeclinii Poland (Fain, Hurst et al., 1997; Kiełczewski, Moser & Wiśniewski, 1983; Kiełczewski & Wiśniewski, 1971, 1980; Haitlinger, 1904; Michalski & Ratajczak, 1989) – Divilia exigua, Probonomoia pini Russia (Zachvatkin, 1941; Türk & Türk, 1957; Krivoluckiy, 1975; Bugrov, 1997; Klimov, 1998; Ermilov & Mokrousov, 2008, 2010; Khaustov et al., 2018) – Calvolia elisabethae, C. goetzae, C. kamenskii, C. romanovae, C. thraca, Crabrovidia gussakovskii, Parawinterschidtia kneissli, Probonomoia pini, P. sibirica, Winterschmidtia nataliae Crimea (Zachvatkin, 1941; Khaustov, 2000) - Crabrovidia gussakovskii, Parawinterschidtia kneissli, Winterschmidtia brenyi, W. chaetoptelii, W. villifronsi, W. zachvatkini Kuril Islands (Klimov, 1998) - Crabrovidia gussakovskii (Iturup) Slovakia (Dusbabek, 1964; Kaluz & Žuffa, 1986; Sidor, 1986; Zamec, 2013, 2014) – Calvolia astoma, C. fraxini, C. goetzae, C. heterocoma, C. kneissli, C. longipes, C. miniopteri, C. reticulata, C. tarsoinfracta, Ensliniella parasitica Spain (Fain, 1973, Klompen & OConnor, 1995) - Crabrovidia ectemnii, C. popovi, Ensliniella parasitica Switzerland (Fain, 1973) - Crabrovidia ectemnii Turkey (Fain, 1973; Klompen & OConnor, 1995) - Crabrovidia popovi, Ensliniella dignotus, E. floricola, Е. kostylevi Ukraine (Zachvatkin, 1941; Khaustov, 2000; Rusina & Orlova, 2011) - Crabrovidia popovi, Ensliniella kostylevi, Sphexicozela connivens, Winterschmidtia hamadryas, W. natalyae, AFRICA Algeria (Klompen & OConnor, 1995) – Ensliniella kostylevi DR Congo (Cooreman, 1942; Fain, 1971; Fain & Elsen, 1972, 1973; Fain & Beaucournu, 1976; Samšinák, 1989) – Calvolia marie, Congovidia glossinae, Congovidiella hieroglypha, Psylloglyphus uilenbergi kivuensis, Winterschmidtia (= Afrocalvolia) glossinarum, Trypetacarus trypetidarum Egypt (Baker & Cunliffe, 1960; Sevastianov & Gad-Hamada-Hassa, 1989; Hassan & Sevastianov, 1989; Eraky, 1998; El-Kawas & Negm, 2018) – Calvolia solimani, C. zaheri, Ensliniella aegyptiaca, Procalvolia hamadi 382 Acarorum Catalogus IX Gambia (Mahunka, 1975) - Gambacarus huettleri Ghana (Mahunka, 1974) – Allocalvolia bisculpturata Ivory Coast (Fain, 1974) – Winterschmidtia (= Afrocalvolia) athiasae, W. lamtoensis Madagascar (Fain, 1966; Fain & Beaucournu, 1976) - Psylloglyphus uilenbergi Kenya (Fain & Schwan, 1976) – Psylloglyphus uilenbergi kivuensis Morocco (Fain, 1972, 1973; Baker & Cunliffe, 1960) – Crabrovidia ectemnii, Ensliniella koenigi Mozambique (Meyer & Rodrigues, 1965) – Oulenziella bakeri (syn. Oulenzia gossypii) Rwanda (Fain & Beaucournu, 1976) - Psylloglyphus (Hemimerus) hemimerus South Africa (Fain, 1973) - Crabrovidia dasyproctus ASIA China (Chang et al., 2017, 2020) - Winterschmidtiidae Taiwan (Fain, 1973) – Crabrovidia dasyproctus Cyprus (Fain, 1973) - Crabrovidia ectemnii, C. leclercqi India (Hughes, 1962; Fain, 1973) - Crabrovidia dasyproctus, Calvolia bakeri Indonesia (Oudemans, 1928; Hughes, 1962) – Oulenzia arboricola, Calvolia bakeri Iran (Ahadiyat & Ostovan, 2006; Faradonbeh et al., 2018; Nazari, Monfared, Nemati & Azhari, 2019) – Calvolia priapus, Crabrovidia oudemansi, Vidia lineata, Winterschmidtia hamadryas Japan (Fain, 1973; Okabe & OConnor, 2002; Abe & Okabe, 2006) – Crabrovidia ectemnii [Okinawa], Ensliniella asiatica, Kurosaia jiju Kazakhstan (Volgin, 1974) - Neottiglyphus alatavicus Malaysia (Fan et al., 2012) – Oulenzia sp. Philippines (Fain, 1973) - Crabrovidia dasyproctus Singapour (Fain, 1973) – Crabrovidia dasyproctus Tadjikistan (Klompen & OConnor, 1995) - Ensliniella parasitica Uzbekistan (Zachvatkin, 1941) - Crabrovidia popovi, Ensliniella parasitica Vietnam (Fain & Beaucournu, 1972) – Psylloglyphus vietnamensis NORTH AMERICA Bermudas (Fain & Schuster, 1983) – Neocalvolia inermis Canada (Nesbitt, 1946; Baker & Cunliffe, 1960; OConnor & Eickwort, 1988; Mullen, Leprince & OCoonor, 1989) – Crabrovidia latimanus, Vespacarus vagus, Vidia sp. A and B, Calvolia sp. Mexico (Baker & Cunliffe, 1960; Mostafa, 1970b, 1971; Fain, 1972, 1973) – Crabrovidia ectemnii, Monobiacarus funebris, M. insularis, Zethacarus boharti, Z. ellipsus, Z. miscogaster, Zethovidia floridae USA (Baker & Cunliffe, 1960; Baker, 1964; Woodring, 1966; Mostafa, 1967; Delfinado & Baker, 1976; Fain & Philips, 1978; OConnor & Eickwort, 1988; Fan, George & Kumarasinghe, 2010) – Acalvolia americana, Calvolia furnissi, Crabrovidia rubi, C. Acarorum Catalogus IX 383 texana, C. thomasi, C. utahensis, Monobiacarus quadridens, Sapracarus tuberculatus, Vespacarus anacardivorus, V. histrio, V. minimoferus, V. pedestris, V. tigris, V. toltecus SOUTH AND CENTRAL AMERICA Argentina (Mostafa, 1971; Fain, 1972, 1973) - Crabrovidia ectemnii, Zethacarus andinus, Z. argentinae, Z. haywardi, Zethovidia parvula Bolivia (Fain, 1972, 1973) - Crabrovidia ectemnii Brazil (Mostafa, 1970b; 1971; Fain, 1972, 1973; Fain & Camerik, 1978; Barbosa & de Moraes, 2021) – Acalvolia squamata, Calvolia summersi, Crabrovidia brasiliensis, C. crossocerus, C. ectemnii, Czenspinskia transversostriata, Dichotomiopus biiaggioi, Macroharpa amazonica, M. arota, M. pachymapoda, M. palpognatha, Oulenzia arboricola, Oulenziella bakeri, O. longiseta, Zethacarus paranicus, Z. smithii, Zethovidia oculabsena, Z. brasilensis, Z. diminuta, Z. promina Chile (Fain & Beaucournu, 1986, 1989) – Psylloglyphus (Tetrapsyllopus) chiliensis, P. (T.) micronychus Colombia (Fain, 1972, 1973) - Crabrovidia ectemnii Costa Rica (Fain, 1972, 1973) - Crabrovidia ectemnii Cuba (Fain, 1973) - Crabrovidia cubensis El Salvador (Mostafa, 1970a) - Zethovidia salvadorensis French Guiana (Mostafa, 1970b) - Zethovidia acaris Guiana (Fain, 1972, 1973) - Crabrovidia ectemnii Honduras (Fain, 1972, 1973) - Crabrovidia ectemnii Jamaica (Fain, 1973) - Crabrovidia ectemnii Panama (Mostafa, 1970b) – Zethovidia fusca Paraguay (Mostafa, 1970a) - Zethovidia diminuta, Z. paraguayensis Puerto Rico (Fain, 1973) - Crabrovidia ectemnii Suriname (Mostafa, 1970b) - Zethovidia brevapodemis Venezuela (Mostafa, 1970b) - Zethovidia reticulata OCEANIA Australia (Womersley, 1941; Fain, 1989; Fain, Bartholomaeus, Cooke & Beaucornu, 1990) – Calvolia glabra, Czenspinskia transversostriata, Psylloglyphus australiensis, P. foveolatus [Tasmania, Victoria], Saproglyphus cocciphagus New Zealand (Fain & Galloway, 1973a, 1973b) - Psylloglyphus (Psylloglyphus) parapsyllus ANTARCTICA South Georgia (Hughes, 1970) – Neocalvolia claggi South Shetlands (Rosas & Casanueva, 1993) - Neocalvolia coperminensis South Sandwich Islands (Hughes, 1966) - Calvolia antarctica 384 Acarorum Catalogus IX Hosts of Winterschmidtiidae Insecta Coleoptera Fam. Scarabaeidae Aphodius minor – Calvolia goetzae prodromus - Calvolia goetzae Cetonia aurata – Vidia sp. Fam. Curculionidae Subfam. Scolytinae Hylesinus fraxini – Winterschmidtia nataliae Ips stebbingi – Winterschmidtia circumspectans Leperisinus omi – Winterschmidtia nataliae, Calvolia fraxini Orthomicus laricis – Parawinterschmidtia kneissli Pityogenes calcaratus - Winterschmidtia zachvatkini Pteleobius vittatus – Winterschmidtia brenyi Scolytus amygdali – Winterschmidtia hamadryas S. intricatus – Calvolia minima Fam. Coccinellidae Adalia bipunctata - Congovidia benelux Calvia duodecimguttata – Divilia exigua Chilocorus renipustulatus – Divilia occidentalis Halyzia sedecimguttata - Congovidia coccinellidarum Fam. Cerambycidae Mesosa nebulosa - Parawinterschmidtia kneissli Dermaptera Fam. Labiduridae Chelidura acanthopygia – Calvolia longipes Labidura riparia - Procalvolia hamadi Hymenoptera Fam. Megachilidae Megachile pilicrus - Vidia lineata Megachile (Litomegachile) texana – Crabrovidia texana Megachile (Sayapis) fidelis - Crabrovidia hirsuta Megachile (Xeromegachile) rubi – Crabrovidia rubi Fam. Crabronidae Crabro solenius - Crabrovidia gussakovskii Cerceris arenaria – Crabrovidia concellaria C. rubida – Crabrovidia leclercqi Cerceris sp. - Crabrovidia oudemansi Acarorum Catalogus IX 385 Crossocerus elongatus – Crabrovidia corsicana C. porexus - Crabrovidia crossocerus Dasyproctus agilis orientalis – Crabrovidia dasyproctus D. bipunctatus lugubris - Crabrovidia dasyproctus D. buddha - Crabrovidia dasyproctus D. cevirus - Crabrovidia dasyproctus D. westermanni - Crabrovidia dasyproctus Ectemnius carinatus - Crabrovidia ectemnii E. cavifrons - Crabrovidia ectemnii E. centralis - Crabrovidia ectemnii E. continuus [= Crabro vagus auct.] - Crabrovidia ectemnii, C. gussakovskii E. craesus - Crabrovidia ectemnii E. cubiceps – Crabrovidia brasiliensis E. [Crabro] fossorius - Crabrovidia gussakovskii E. guttatus - Crabrovidia ectemnii E. hypsae - Crabrovidia ectemnii E. impressus - Crabrovidia ectemnii, C. gussakovskii, C. popovi E. laevigatus - Crabrovidia ectemnii E. lapidaries - Crabrovidia ectemnii E. schettereri - Crabrovidia ectemnii E. semipunctatus - Crabrovidia ectemnii E. singularis - Crabrovidia ectemnii E. sonorensis - Crabrovidia ectemnii E. sexcinctus - Crabrovidia ectemnii Lestica alata - Crabrovidia popovi L. clypeata – Crabrovidia popovi L. cubensis – Crabrovidia cubensis L. wollmanni - Crabrovidia popovi Sphex hemorrhoidalis – Crabrovidia africana Fam. Pteromalidae Habrocytus elevatus – Allocalvolia habrocytus Fam. Colletidae Hylaea (Prosopis) conformis – Vidia undulata Fam. Vespidae Subfam. Eumeninae Allodynerus floricola - Ensliniella floricola A. delphinalis – Ensliniella parasitica A. mandschuricus – Ensliniella asiatica Subfam. Zethinae Zethus andinus – Zethacarus andinus Z. carbonarius – Zethovidia acaris 386 Acarorum Catalogus IX Z. coeruleopennis – Z. brazilensis Z. d. diminutus – Z. diminuta Z. dimunutus santaremae – Z. diminuta Z. haywardi – Zethacarus argentinae Z. guerreroi arizonensis – Zethovidia arizonensis Z. guerreroi mayorum – Zethacarus boharti Z. orizabae – Z. ellipticus Z. miscogaster – Z. miscogaster Z. smithii – Z. paranicus Z. arotus – Macroharpa arota Z. biglumis ferrugineus – Macroharpa amazonica Z. lobulatus – Macroharpa pachymapoda Z. orizabae – Zethacarus ellipsus Z. roridus – Macroharpa palpognatha Subfam. Polystinae Polystes dominula - Sphexicozela connivens P. gallica - Sphexicozela connivens P. nimpha - Sphexicozela connivens Fam. Eulophidae Melittobia acasta – Ensliniella parasitica Diptera Fam. Trypetidae Celidodacus fenestratus - Trypetacarus trypetidarum Fam. Glossinidae Glossina fuscipes quanzensis - Winterschmidtia glossinarum, W. tsetse, Congovidia glossinae Glossina palpalis palpalis - Winterschmidtia glossinarum Fam. Sphaeroceridae Gymnometopina magna - Calvolia marie Fam. Tabanidae Hybomitra epistates – Vidia sp. A, Calvolia sp. H. affinis – Calvolia sp. H. lasiophthalmus - Calvolia sp. H. microcephala - Vidia sp. A H. nuda - Calvolia sp. Tabanus atratus - Calvolia sp. T. quinquevittatus - Vidia sp. B Siphonaptera (phoretic acari on parasites) Notiopsylla kerguelenensis – Notiopsyllopus segermanae, Psylloglyphus crenulatus Synopsyllus fonquerniei – Psylloglyphus u. uilenbergi Acarorum Catalogus IX 387 Paractenopsyllus petiti - Psylloglyphus u. uilenbergi Medwayella robinsoni – Psylloglyphus maculatus Stivalius aporus rectodigitus – Psylloglyphus vietnamensis Pygiopsylla hoplia – Psylloglyphus foveolatus Bibliography of Winterschmidtiidae Abe W., K. Okabe. 2006. 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Acari, Myriapoda et Scorpiones hucusque in Italia reperta 57, 17 text pages. Bernini F., M. Castagnoli, R. Nannelli. 1995. Checklist delle specie fauna d’Italia. Fascicolo 24 Arachnida: Acari. Ed. Calderini. Bugrov S.A. 1997. [Free-living Astigmata (Acariformes) of the Moscow District fauna]. -Zoologicheskii Zhurnal, 76: 147–156. Buitendijk A.M. 1945. Vorloopige catalogus van de Acari in de Collectie – Oudemans. – Zool. Meded., 24: 281–391. Chang R., T.A. Duong, S.J. Taerum, M.J. Wingfield, Zhou X, Z.W. de Beer. 2017. Ophiostomatoid fungi associated with conifer-infesting beetles and their phoretic mites in Yunnan, China. – MycoKeys, 28: 19–64. Chang Runlei, T. A. Duong, S. J. Taerum, M. J. Wingfield, Xu Dong Zhou, Z. W. de Beer. 2020. Ophiostomatoid fungi associated with mites phoretic on bark beetles in Qinghai, China. - IMA Fungus, 11: 15. Chmielewski W. 1997. Dane biologiczne roztoczka suszowego, Carpoglyphus lactis (L.) 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Soil astigmatine (Acari: Astigmatina) mites from birdsʼ nests in Slovakia. Acta Societatis. - Zoologicae Bohemicae, 77: 165–172. Zamec R., V. Košel. 2014. A new species of mite (Acari: Histiostomatidae) phoretic on Gymnomus caesius (Diptera: Heleomyzidae) from Vlčie Diery cave. - · Biologia, 69(7): 916–919. Acknowledgements This volume required the support of many distinguished specialists, to mention only a few: A.Fain, J.Cooreman, R.Haitlinger, S.Mahunka, N.J.Fashing, K.Samšinák, A.A.Khaustov, U. Gerson, B.M.OConnor, P.B.Klimov, J.de la Cruz, and others. To all of them the author is immensely grateful. ACARORUM CATALOGUS IX The ninth volume of this series "Acarorum Catalogus" is devoted to the Superfamilia Shizoglyphoidea (Schizoglyphidae). Histiostomatoidea (Histiostomatidae, Guanolichidae). Canestrinioidea (Canestriniidae, Chetochelacaridae, Lophonotacaridae, Heterocoptidae). Hemisarcoptoidea (Chaetodactylidae, Hyadesiidae, Algophagidae, Hemisarcoptidae, Carpoglyphidae, Winterschmidtiidae). About 1268 species of 225 genera are listed therein, with synonymies and citations in various papers. The book also contains a list of by-country distributions, type localities, hosts of the parasitic species and lists of authors who published on any known species of these families. The last Xth volume of the same series is under way.