TAXON 56 (2) • May 2007: 571–582 Garnock-Jones & al. • Southern Hemisphere Veronica Botanical names in Southern Hemisphere Veronica (Plantaginaceae): sect. Detzneria, sect. Hebe, and sect. Labiatoides Phil Garnock-Jones1, Dirk Albach2 & Barbara G. Briggs3 1 2 3 Centre for Biodiversity & Restoration Ecology, School of Biological Sciences, Victoria University of Wellington, P.O. Box 600, Wellington 6140, New Zealand. phil.garnock-jones@vuw.ac.nz (author for correspondence) Institut für Spezielle Botanik, Johannes Gutenberg-Universität Mainz, Bentzelweg 9b, 55099 Mainz, Germany Botanic Gardens Trust, Mrs Macquaries Road, Sydney 2000, Australia The classification of the Southern Hemisphere Veronica complex is discussed in the light of recent findings that these segregate genera are nested within the hitherto northern Veronica clade. In order to render Veronica monophyletic, we transfer Chionohebe, Derwentia, Detzneria, Hebe, Hebejeebie, Heliohebe, Leonohebe, and Parahebe to Veronica subgen. Pseudoveronica. Correct names are listed for all species in Veronica subgen. Pseudoveronica, according to the International Code of Botanical Nomenclature. Those species previously included in Derwentia together with other Australian species of Veronica are now classified in Veronica sect. Labiatoides, that in Detzneria is now classified in Veronica sect. Detzneria, and those in Chionohebe, Hebe, Heliohebe, Leonohebe, and Parahebe are now classified in Veronica sect. Hebe. Seventy-nine nomenclatural changes are provided: 61 new combinations (1 at section rank, 37 at species rank, 23 below species rank) and 18 new names (including 1 new name for a northern Veronica from the Caucasus). KEYWORDS: Australia, Chionohebe, Derwentia, Detzneria, Hebe, Heliohebe, Leonohebe, New Guinea, New Zealand, Parahebe, Plantaginaceae, Veronica subg. Pseudoveronica INTRODUCTION For over 150 years from their first collection by Banks & Solander in 1769, inclusion in Veronica of Southern Hemisphere plants now known as the Hebe complex (Heads, 1992) was not controversial. Although Hebe had been described as early as 1789, it received little acceptance until Pennell (1921), Oliver (1925), Cockayne & Allan (1926), and Cockayne (1929). During the years following acceptance of Hebe by New Zealand authors, the placement of several Veronica-like species was problematic. These plants have morphological characters that seem to place them close to Northern Hemisphere Veronica, but their chromosome numbers suggest a relationship with Hebe. Allan (1939) and Oliver (1944) placed greater weight on the cytological evidence, and Oliver (1944) erected a new genus Parahebe and resurrected Hooker’s Pygmea to accommodate some of these plants. Similarly the affinities of Veronica-like plants in Australia have also been unclear (Briggs & Ehrendorfer, 2006a). Overall, the close relationship of Southern Hemisphere plants to Veronica has never been questioned, although some authors have thought the Hebe complex ancestral to Veronica (Hong, 1994) whereas others have thought Veronica ancestral to Hebe (Raven, 1973; Garnock-Jones, 1975, 1993a). By the early 21st century, most taxonomic treatments placed the species of Southern Hemisphere Veronica under Chionohebe (a replacement name for Pygmea – Briggs & Ehrendorfer, 1976), Derwentia, Detzneria, Hebe, Heliohebe, Leonohebe, and Parahebe (Ashwin in Allan, 1961; Briggs & Ehrendorfer, 1976, 1992; van Royen, 1983; Garnock-Jones, 1993b; Garnock-Jones & Lloyd, 2004; Bayly & Kellow, 2006), but see Heads (1987, 1992, 1993, 1994a, b, c, 2003) for alternative circumscriptions. Recently, Albach & Chase (2001, 2004), Wagstaff & al. (2002), and Albach & al. (2004a, b) have shown with cladistic analyses of nucleic acid sequence data that recognition of Southern Hemisphere segregates renders Veronica paraphyletic. Veronica persica, for example, is more closely related to the Southern Hemisphere Hebe complex than it is to V. officinalis, the type of Veronica. Thus, Albach & al. (2004c) have demonstrated the necessity for a major realignment of generic circumscriptions in Veronica. SUMMARY OF RECENT PHYLOGENETIC TREATMENTS Since the first DNA-based phylogenetic analyses of Veronica (s.l.) by Wagstaff & Garnock-Jones (1998, 2000) and Albach & Chase (2001), several analyses addressed the circumscription of the genus (e.g., Wagstaff & al., 2002; Albach & Chase, 2004; Albach & al., 2004b, c, 571 Garnock-Jones & al. • Southern Hemisphere Veronica 2005b). Based on nuclear ribosomal ITS region and cpDNA sequences from the trnL-trnF region and the rps16 intron, these studies supported a circumscription of Veronica that includes Pseudolysimachion, the North American genus Synthyris (incl. Besseya—Hufford & McMahon, 2004), and the Australasian genera Derwentia, Detzneria, and Hebe and their relatives. Whereas such a circumscription of the Veronica clade seems to be unambiguous, the relationship of the major clades (subgenera) within Veronica remains elusive. All analyses support Veronica subgenera Beccabunga, Pseudolysimachium, Synthyris, and Veronica as early diverging clades. The Australasian genera form a monophyletic group nested among a group of Veronica subgenera that have a chromosome base number of x = 8, but other inferences depend on the marker used and taxon sampling. Sequencing low copy nuclear DNA regions is currently in progress to resolve these relationships (Albach, unpubl.) but is unlikely to give conclusive answers itself due to the problems of orthology in groups, such as the Southern Hemisphere Veronica, in which all subgenera have arisen from polyploid ancestors. GENERIC REALIGNMENTS There is widespread, but not universal (see Brummitt, 2002; 2006), agreement among taxonomists that recognised and named groups of species should be monophyletic. Only in a monophyletic group is every group member more closely related to every other group member than to any species classified in another group at the same rank. Additionally, it is argued that classifications that recognise only monophyletic groups enable better predictive inferences to be based upon them, compared with classifications that also accept paraphyletic and polyphyletic groups (Wiley, 1981; Judd & al., 2002). Further, although some (e.g., Cronquist, 1987; Brummitt, 2002, 2006) argue that distinctive paraphyletic groups should be recognised taxonomically, there are no objective criteria by which to decide which paraphyletic groups might be deemed acceptable. In contrast, monophyletic groups can be circumscribed objectively, although there are no agreed objective criteria for assigning ranks to them. It is also argued (e.g., Wiley, 1981; Judd & al., 2002) that phylogenetic classifications are more informationrich, but this is not universally so if groups become more inclusive and smaller groups within them are not recognised. Bayly & Kellow (2006) argued that including the Hebe complex within Veronica would lower information content, presumably because fewer and larger groups would be recognised. In addition to the quantity of information that can be inferred from a classification, it is also important to consider information quality. In all 572 TAXON 56 (2) • May 2007: 571–582 phylogenetic classifications, information content is based on inferred evolutionary relationships. Our preferred classification, presented below, treats recognisable clades at infrageneric rank, thus retaining high quality information content. As more information becomes available, further well-defined clades will also be identified and recognised in this way, but at present we lack information, especially within Hebe s.str. We can see four potential ways to configure the Southern Hemisphere members of the Veronica complex such that only monophyletic groups are recognised. These differ in the taxonomic rank at which these clades are treated. We set the alternatives out below, with a discussion of their advantages and disadvantages. (1) Retain Hebe with current circumscription. — Hebe is monophyletic if Leonohebe s.str. (i.e., L. cheesemanii, L. ciliolata, L. cupressoides, L. tetrasticha, and L. tumida) and the Australian species H. formosa are excluded (Wagstaff & al., 2002; Bayly & Kellow, 2006). In addition, a case could be made for widening its circumscription to include Heliohebe, which is sister group of the main Hebe clade (Wagstaff & al., 2002). In either case, such a circumscription of Hebe renders Parahebe paraphyletic (Wagstaff & al., 2002; Garnock-Jones & Lloyd, 2004), and would require the breakup of Parahebe into several small genera. Heads (2003) made a first step along this route by erecting Hebejeebie to include two species of Parahebe and one of Chionohebe. In such a solution, the numerous small southern segregate genera would mostly be retained, and the paraphyly of Veronica would need to be addressed by breaking it up into many small genera, the consequences of which are discussed below. Changes would be required to the generic classification of Australian species that have been retained in Veronica or referred to Parahebe, as indicated for option 2 below. (2) A single New Zealand genus. — The New Zealand Hebe complex is clearly monophyletic (Wagstaff & al., 2002) and could be a single genus, for which the correct name would be Hebe. Hebe, Parahebe, Heliohebe, Chionohebe, and Leonohebe would be included in this circumscription of Hebe. Australian Chionohebe, New Guinea Parahebe, and perhaps Detzneria would also need to be included in Hebe (Wagstaff & al., 2002; Albach & al., 2005b), but the Australian species Parahebe lithophila and Hebe formosa would be included in Derwentia. Wall (1929) and Allan (1939) took such a broader view of Hebe for New Zealand species, while most New Guinea species were first described under Hebe. This course of action would remove the difficulty of a paraphyletic Parahebe (Garnock-Jones & Lloyd, 2004) and a possibly polyphyletic Chionohebe (Wagstaff & al., 2002). In such a solution, the Australian segregate genus Derwentia would be retained. Again, the paraphyly of Veronica would need to be addressed by breaking it up into many small genera, discussed below. TAXON 56 (2) • May 2007: 571–582 (3) A single southern hemisphere genus. — In this approach, Hebe of option 2 above would be enlarged further to include its sister group the Australian genus Derwentia, including a number of Australian species that until now have been retained in Veronica. Most Australian species would require new combinations under the correct name of this larger southern genus, Hebe, and again the paraphyly of Veronica would need to be addressed by breaking it up into many small genera. All three options above do not directly address the paraphyly of Veronica when Southern Hemisphere groups are excluded. In our opinion, any of them could be applied only if the currently paraphyletic Veronica were to be broken up into smaller monophyletic genera. However, many of the clades that were found within Veronica by Albach & Chase (2001) and subsequent work cannot be recognised by morphological characters (Albach & al., 2004c) and species from different subgenera are often confused. Furthermore, most widely known species, all of the weedy species, and most species of horticultural importance would have to be transferred to new genera. Based on our best current phylogenetic hypothesis and species estimate, such a decision would require name changes for 193 (of 420) species and the creation of five entirely new genera, three of them for species widespread and common in Europe. Morphological cohesion of the Northern Hemisphere Veronica is such that splitting it would not be accepted. The name changes would affect all major floras of the world, even that of New Zealand (Webb & al., 1988), because none of the widespread weedy species V. persica, V. anagallis-aquatica or V. javanica would be retained in Veronica. If Veronica were divided, the name Veronica would apply to only one small clade that comprises the type V. officinalis and its close relatives, a total of 40–45 species (Albach & al., 2004c). Thus, any of the three options above would disrupt nomenclature in the Northern Hemisphere very much more than enlarging Veronica would disrupt it in the south. Shrubby New Zealand veronicas (usually treated as Hebe) are important in ornamental horticulture in both hemispheres and we are mindful of the effects of our proposed changes on familiar names. However, it took over 30 years for Hebe to be accepted as an alternative to Veronica in cultivation, e.g., Stearn (1956) treated New Zealand species under Veronica. The Hebe Society (see Hutchins, 1997) promotes the cultivation of New Zealand Veronica; our retention of Hebe at section rank, and perhaps also the use of hebe as a common name, will retain a link to previous names. Because cultivar names can be used with vernacular names (Brickell & al., 2004), names of familiar garden plants like Hebe ‘Inspiration’ only require minor typography changes to remain acceptable (e.g., hebe “Inspiration”). Garnock-Jones & al. • Southern Hemisphere Veronica The Hebe complex is widely believed in New Zealand and elsewhere to be morphologically quite different from Northern Hemisphere Veronica, leading to the view that a paraphyletic Veronica would be acceptable because of Hebe’s distinctiveness (Brummitt, 2006). However, not a single morphological character of those that have been used to delimit Hebe and its relatives is diagnostic for the Southern Hemisphere clade. Examples of such Hebe characteristics include (with non-conforming southern examples in parentheses): shrubby habit (V. lilliputiana, V. zygantha), axillary racemes (V. hulkeana, V. lilliputiana, V. raoulii ), entire leaves (V. derwentiana, V. hulkeana, V. macrantha), white flowers (V. benthamii, V. lilliputiana, V. perfoliata), long corolla tubes (V. decora, V. lyallii ), and latiseptate septicidal capsules (V. cheesemanii, V. notialis, V. zygantha). In other words, the morphological distinction between the Hebe clade and the Veronica grade is illusory, such that this is not simply an argument about acceptance or rejection of a recognisable paraphyletic group. While the crown clade of Hebe is recognisably different from northern Veronica, attempts to deal with a large basal grade of Veronica-like species in the Hebe complex (Oliver, 1944; Heads, 1987, 2003; Garnock-Jones, 1993b) have led to the profusion of small, segregate, sometimes non-monophyletic genera in recent years. (4) A greatly-enlarged Veronica. — This option involves the enlargement of Veronica to include not only American Synthyris and Besseya (Albach & al., 2004c) but also the Southern Hemisphere genera. The first three options (above) do not directly address the paraphyly of Veronica when Southern Hemisphere groups are excluded. Option 4 brings together a related group of species that share the following synapomorphies (mostly with several reversals within the genus): tetramerous calyx and corolla (pentamerous in a few New Zealand species), short to absent corolla tube (long in many sect. Hebe), subactinomorphic flowers, two exserted stamens, notched fruit apex (acute in many sect. Hebe and some subg. Stenocarpon), much compressed fruit (usually angustiseptate, but latiseptate in many sect. Hebe, and turgid in H. pareora and some Parahebe) and flattened seed (Albach & al., 2004c). In our opinion, such enlargement of Veronica is the most sensible and globally applicable solution, and we provide names in Veronica below for all the Southern Hemisphere species that we recognise. Inclusion of the Hebe complex within Veronica encourages comparisons of related species and thus makes patterns of character evolution, phytochemistry (Taskova & al., 2004; Albach & al., 2005a), biogeography (Albach & al., 2005b) and horticulture much more evident. Already, a broader concept of Veronica has made possible a revised understanding of the origin and relationships of several Australian species (Briggs & Ehrendorfer, 2006a). 573 Garnock-Jones & al. • Southern Hemisphere Veronica INFRAGENERIC NAMES The few names that are available between the ranks of genus and species have been discussed, and typified where necessary, by Bayly & Kellow (2004). It would be convenient to have a name at subgenus rank to refer to the entire Southern Hemisphere clade. The correct name under the ICBN must be chosen between V. subg. Pseudoveronica J.B. Armstr. and V. subg. Koromika J.B. Armstr. (Bayly & Kellow, 2004); both were published at the same time. In our opinion the former is inelegant and somewhat inappropriate, while the latter is an incorrect rendition of koromiko, the Maori name for most large leaved plants of New Zealand Veronica. We prefer Pseudoveronica to a miss-spelled Maori name that might have been appropriated into scientific nomenclature without approval of tangata whenua (New Zealand indigenous people). Veronica L., Sp. Pl.: 9. 1753 – Type: Veronica officinalis L. Synonyms based on Southern Hemisphere types: = Chionohebe B.G. Briggs & Ehrend. in Contr. Herb. Austral. 25: 1. 1976 ≡ Pygmea Hook. f., Handb. N. Zeal. Fl.: 217. 1864, nom. illeg. (non Pygmaea Stackh. in Mém. Soc. Imp. Naturalistes Moscou 2: 60, 95. 1809) – Lectotype (Oliver, 1944): P. ciliolata. = Derwentia Raf., Fl. Tellur.: 55. 1836 – Lectotype (Garnock-Jones & al., 1990): D. suaveolens (nom. illeg. ≡ D. derwentiana). = Detzneria Diels in Bot. Jahrb. Syst. 62: 490. 1929 – Type: D. tubata. = Hebe Juss. in Gen. Pl.: 105. 1789 – Type: H. magellanica. = Hebejeebie Heads in Newslett. Bot. Soc. Otago 36: 11. 2003 – Type: H. densifolia. = Heliohebe Garn.-Jones in New Zealand J. Bot. 31: 323. 1993 – Type: H. lavaudiana. = Leonohebe Heads in Newslett. Bot. Soc. Otago 5: 4. 1987 – Type: L. ciliolata. = Panoxis Raf. in Med. Fl. 2: 109. 1830 – Lectotype (here designated): Veronica salicifolia. [Of the three included species, V. catarractae does not match the protologue because its capsule is didymous, and V. salicifolia in its tubular corolla and oblong capsule matches the protologue more closely than does V. macrocarpa. No species combinations were made in Panoxis.] = Parahebe W.R.B. Oliv. in Rec. Domin. Mus. 1: 229. 1944 – Type: P. catarractae. Veronica subg. Pseudoveronica J.B. Armstr. in Trans. & Proc. New Zealand Inst. 13: 351. 1881 (as Pseudoveronica) – Lectotype (Bayly & Kellow, 2004): Veronica lycopodioides Hook. f. = Veronica subg. Koromika J.B. Armstr. in Trans. & Proc. 574 TAXON 56 (2) • May 2007: 571–582 New Zealand Inst. 13: 351. 1881 – Lectotype (Bayly & Kellow, 2004): Veronica pubescens Benth. We have corrected the orthography of Pseudoveronica by deleting the hyphen, in accordance with Art 60.9 and Ex. 16, especially noting the cited example Scirpus sect. Pseudoëriophorum. Below subgenus rank, more familiar names are available (Bayly & Kellow, 2004), and we advocate use of three of these to identify familiar clades: (1) V. sect. Detzneria, monotypic in New Guinea, (2) V. sect Hebe, for the entire New Zealand clade, including some species and groups that apparently resulted from dispersal from New Zealand to Australia, New Guinea, Rapa, and South America, and (3), V. sect. Labiatoides, for a smaller clade confined to Australia. Relationships within these clades are still unclear such that less inclusive monophyletic groups cannot yet be recognised with confidence. ITS sequences provide the most comprehensive dataset, but published trees have been poorly resolved within V. sect. Hebe (Wagstaff & Garnock-Jones, 1998, 2000; Wagstaff & al., 2002). More comprehensive species sampling (Low, 2005) generates ITS trees with nested polytomies that cannot easily be translated into infrageneric classifications of monophyletic groupings. ITS trees also may be incongruent with trees from chloroplast sequences in Veronica s.l. (Albach & Chase, 2004; Albach & al., 2005b), including V. sect. Hebe (Low, 2005). Veronica (subg. Pseudoveronica) sect. Detzneria (Diels) Albach, comb. nov. ≡ Detzneria Diels in Bot. Jahrb. Syst. 62: 490. 1929 – Type: D. tubata Diels. Veronica (subg. Pseudoveronica) sect. Hebe (Juss.) G. Don in Gen. Hist. 4: 570. 1838 – Lectotype (designated here): Veronica elliptica G. Forster; this species fits the description in the protologue, and maintains current usage. = Hebe sect. Glaucae Heads in Newslett. Bot. Soc. Otago 5: 11. 1987 – Type: Hebe pinguifolia (Hook. f.) Cockayne & Allan. = Hebe sect. Hebe [an autonym established by the publication of other section names in Hebe by Heads (1987)] – Type: Hebe magellanica J.F. Gmel. [= H. elliptica (G. Forster) Pennell]. = Hebe sect. Subdistichae Heads in Newslett. Bot. Soc. Otago 5: 11. 1987 – Type: Hebe diosmifolia (A. Cunn.) Cockayne & Allan. = Leonohebe sect. Apiti Heads in Newslett. Bot. Soc. Otago 5: 7. 1987 – Type: Leonohebe benthamii (Hook. f.) Heads. = Leonohebe sect. Aromaticae Heads in Newslett. Bot. Soc. Otago 5: 8. 1987 – Type: Leonohebe cupressoides (Hook. f.) Heads. TAXON 56 (2) • May 2007: 571–582 = Leonohebe sect. Buxifoliatae Heads in Newslett. Bot. Soc. Otago 5: 10. 1987 – Type: Leonohebe odora (Hook. f.) Heads. = Leonohebe sect. Connatae Heads in Newslett. Bot. Soc. Otago 5: 6. 1987 – Type: Leonohebe epacridea (Hook. f.) Heads. = Leonohebe sect. Flagriformes Heads in Newslett. Bot. Soc. Otago 5: 8. 1987 – Type: Leonohebe hectorii (Hook. f.) Heads. = Leonohebe Heads sect. Leonohebe [an autonym established by the publication of other section names in Leonohebe by Heads (1987)] – Type: Leonohebe ciliolata (Hook. f.) Heads. = Leonohebe sect. Salicornioides Heads in Newslett. Bot. Soc. Otago 5: 7. 1987 – Type: Leonohebe salicornioides (Hook. f.) Heads. = Veronica sect. Piritia G. Don in Gen. Hist. 4: 570. 1838 – Lectotype (designated here): Veronica speciosa A. Cunn. [The protologue listed three species. Of these V. diosmifolia does not match the protologue because its leaves are not entire, and the description of V. ligustrifolia does not mention fruits suggesting that Don did not have information about its capsules. The remaining species, V. speciosa, best matches the protologue.] = Veronica sect. Pygmaea (Hook.) Benth. & Hook. f., Gen. Pl. 2: 913. 1876. Veronica (subg. Pseudoveronica) sect. Labiatoides Wettst. in Engler & Prantl, Nat. Pflanzenfam., IV, 3b: 86. 1891 – Lectotype (designated here): V. perfoliata R. Br. Two species, V. perfoliata and V. labiata R. Br. (nom. illeg. = V. derwentiana) were included; V. perfoliata is a better fit to the protologue description “leaves fleshy”. NAMES AT SPECIES RANK AND BELOW In the conspectus that follows, most southern Veronica names have one of the following origins: • A combination has already been published in the genus Veronica and is still available in that genus. These names are listed at the beginning of the entry for each section (including the previously accepted name where this has a different epithet or where confusion between homonyms might arise). • The name was originally published in another genus (often Hebe) and a new combination in Veronica is based upon it because the epithet has not yet been used in Veronica. • The name was first legitimately published in another genus (often Hebe), although it might have been Garnock-Jones & al. • Southern Hemisphere Veronica illegitimately published earlier in Veronica, and the epithet is not available to be used in Veronica [e.g., Hebe divaricata (Cheeseman) Cockayne & Allan, non Veronica divaricata Boiss. & Balansa]. For these, new names have been provided. The species and infraspecific names are listed under their sectional names below. In making new names and new combinations, we have provided only basionyms; additional synonymy and typification is provided by Bayly & Kellow (2004, 2006) for names previously treated under Hebe and Leonohebe; Briggs & Ehrendorfer (1968, 1992, 2006b) for Australian species previously treated under Parahebe and Derwentia; Garnock-Jones (1993b) for names previously treated under Heliohebe; GarnockJones & Lloyd (2004) for New Zealand species previously treated under Parahebe; and van Royen (1972, 1983) and van Royen & Ehrendorfer (1970) both for New Guinea species previously treated under Parahebe and Detzneria. Typification of the names of species previously treated under Chionohebe (Briggs & Ehrendorfer, 1976) is provided herein. We have not included names that are thought to be based on hybrids, e.g., Veronica ×bidwillii. Veronica sect. Detzneria Veronica tubata (Diels) Albach, comb. nov. ≡ Detzneria tubata Diels in Bot. Jahrb. Syst. 62: 491. 1929. Veronica sect. Hebe In addition to the new combinations listed below, the following species and infraspecific taxa, the names of which are already published in Veronica, belong to this section: Veronica adamsii Cheeseman, V. albicans Petrie, V. annulata (Petrie) Cockayne ex Cheeseman, V. armstrongii Johnson ex J.B. Armstr., V. barkeri Cockayne, V. benthamii Hook. f., V. biggarii Cockayne, V. birleyi N.E. Br., V. bishopiana Petrie, V. bollonsii Cockayne, V. brachysiphon (Summerh.) Bean, V. breviracemosa W.R.B. Oliv., V. buchananii Hook. f., V. canterburiensis J.B. Armstr., V. carstensensis Wernham, V. catarractae G. Forster, V. chathamica Buchanan, V. cockayneana Cheeseman, V. colensoi Hook. f., V. cupressoides Hook. f., V. decumbens J.B. Armstr., V. densifolia (F. Muell.) F. Muell., V. dieffenbachii Benth., V. diosmifolia R. Cunn. ex A. Cunn., V. diosmoides Schltr., V. elliptica G. Forster, V. epacridea Hook. f., V. evenosa Petrie, V. gibbsii T. Kirk, V. glaucophylla Cockayne, V. haastii Hook. f., V. hookeriana Walp., V. insularis Cheeseman, V. lanceolata Benth., V. lavaudiana Raoul, V. leiophylla Cheeseman (Hebe gracillima), V. lendenfeldii F. Muell., V. ligustrifolia R. Cunn. ex A. Cunn., V. lilliputiana Stearn (Parahebe canescens), V. linifolia Hook. f., V. lyallii Hook. f., V. lycopodioides Hook. f., V. macrantha Hook. f., V. macrantha var. brachyphylla Cheesem., V. macrocarpa Vahl, V. obtusata Cheeseman, 575 Garnock-Jones & al. • Southern Hemisphere Veronica V. odora Hook. f., V. parviflora Vahl, V. petriei (Buchanan) T. Kirk, V. pinguifolia Hook. f., V. planopetiolata G. Simpson & J.S. Thomson, V. poppelwellii Cockayne, V. propinqua Cheeseman, V. pulvinaris (Hook. f.) Cheeseman, V. quadrifaria T. Kirk (Leonohebe cheesemanii), V. rakaiensis J.B. Armstr., V. rapensis F. Br., V. rupicola Cheeseman, V. salicifolia G. Forster, V. salicornioides Hook. f., V. spathulata Benth., V. speciosa R. Cunn. ex A. Cunn., V. subalpina Cockayne, V. tetrasticha Hook. f., V. thomsonii (Buchanan) Cheeseman, V. townsonii Cheeseman, V. traversii Hook. f., V. trifida Petrie, V. truncatula Colenso, V. tumida T. Kirk, V. vanderwateri Wernham, V. venustula Colenso, and V. vernicosa Hook. f. Veronica albiflora (Pennell) Albach, comb. nov. ≡ Hebe albiflora Pennell in J. Arnold Arbor. 24: 257. 1943. Veronica amplexicaulis J.B. Armstr. f. amplexicaulis, an autonym that is established by the combination V. amplexicaulis f. hirta (below). Veronica amplexicaulis f. hirta (Garn.-Jones & Molloy) Garn.-Jones, comb. nov. ≡ Hebe amplexicaulis f. hirta Garn.-Jones & Molloy in New Zealand J. Bot. 20: 395. 1983. Veronica angustissima (Cockayne) Garn.-Jones, comb. nov. ≡ Veronica salicifolia var. angustissima Cockayne in Trans. & Proc. New Zealand Inst. 50: 184. 1918. Veronica arganthera (Garn.-Jones, Bayly, W.G. Lee & Rance) Garn.-Jones, comb. nov. ≡ Hebe arganthera Garn.-Jones, Bayly, W.G. Lee & Rance in New Zealand J. Bot. 38: 380. 2000. Veronica baylyi Garn.-Jones, nom. nov. ≡ Veronica laevis var. carnosula Hook. f., Fl. Nov.-Zel. 1: 194. 1853 ≡ Veronica carnosula (Hook. f.) Hook. f., Handb. N. Zeal. Fl.: 210. 1864, nom. illeg. (non V. carnosula Lam., Tabl. Encycl. 1: 47. 1791). Etymology: named for Michael James Bayly in recognition of his contributions to knowledge of New Zealand Veronica (e.g., Bayly & Kellow, 2006). Veronica brassii (Pennell) Albach, comb. nov. ≡ Hebe brassii Pennell in Brittonia 2: 185. 1936. Veronica calcicola (Bayly & Garn.-Jones) Garn.-Jones, comb. nov. ≡ Hebe calcicola Bayly & Garn.-Jones in New Zealand J. Bot. 39: 57. 2001. Veronica carminea Albach, nom. nov. ≡ Hebe rubra Pennell in Brittonia 2: 184. 1936 [non Veronica rubra 576 TAXON 56 (2) • May 2007: 571–582 (Douglas ex Hook.) M.M. Mart. Ort. & Albach in Taxon 53: 443. 2004]. Etymology: a replacement epithet that follows the original in referring to the dark red flowers (van Royen, 1972). Veronica cheesemanii Benth. subsp. cheesemanii, an autonym that is established by the combination V. cheesemanii subsp. flabellata (below). Veronica cheesemanii subsp. flabellata (Garn.-Jones) Garn.-Jones, comb. nov. ≡ Parahebe cheesemanii subsp. flabellata Garn.-Jones in New Zealand J. Bot. 42: 197. 2004. Veronica chionohebe Garn.-Jones, nom. nov. ≡ Veronica thomsonii var. glabra Cheeseman, Man. New Zealand Fl.: 540. 1906 (non Veronica glabra Schrad., Comm. Veronic. Spic.: 25. 1803) – Lectotype (designated here): “Mount Pisa, D. Petrie”, AK No. 8335 (upper left hand piece). Typification: Cheeseman did not typify Veronica thomsonii var. glabra in the protologue, but listed several collections under the species that presumably included material of the variety: “Otago – Mt Alta, Buchanan and McKay ! Kurow Mountains, Mount St Bathan’s, Mount Pisa, Petrie ! 4500–6500 ft.” Ashwin (in Allan, 1961) cited just “A [i.e. AK] 8335, Mount Pisa, D. Petrie”, but there are several parts to this gathering and a further lectotypification is necessary. Hence I now select the upper left hand piece on the sheet AK 8335 as lectotype. Etymology: A reference to the genus Chionohebe Briggs & Ehrend. as a noun in apposition. The name means “snow hebe” and is particularly appropriate for this species which occupies snow hollows on mountains of central Otago, New Zealand. Veronica ciliolata (Hook. f.) Garn.-Jones, comb. nov. ≡ Pygmea ciliolata Hook. f., Handb. N. Zeal. Fl.: 217. 1864 ≡ Chionohebe ciliolata (Hook. f.) B.G. Briggs & Ehrend. in Contrib. Herb. Australiense 25: 2. 1976 – Lectotype (Ashwin, in Allan 1961, under var. ciliolata): “Type locality: Hopkins River. – Type: K, Haast.” There are two species of New Zealand Veronica for which the earliest available epithet is ciliolata, most recently known as Chionohebe ciliolata (Hook. f.) B.G. Briggs & Ehrend. and Leonohebe ciliolata (Hook. f.) Heads. Although both basionyms were published in the same work by J.D. Hooker, they are based on different types. In neither case has a combination in Veronica been previously validly published. The present species was first to bear the epithet ciliolata, and it has never been known by any other epithet. We choose to retain it in this TAXON 56 (2) • May 2007: 571–582 usage. The correct name for the other species is Veronica hookeri (see below). Veronica ciliolata var. ciliolata, an autonym established when the varieties below are published. Veronica ciliolata var. fiordensis (Ashwin) Garn.-Jones, comb. nov. ≡ Pygmea ciliolata var. fiordensis Ashwin in Allan, Fl. N. Zeal. 1: 874, 975. 1961. Veronica ciliolata var. pumila (Ashwin) Garn.-Jones, comb. nov. ≡ Pygmea ciliolata var. pumila Ashwin in Allan, Fl. N. Zeal. 1: 874, 975. 1961. Veronica colostylis Garn.-Jones, nom. nov. ≡ Parahebe linifolia subsp. brevistylis Garn.-Jones in New Zealand J. Bot. 14: 288–289. 1976 (non Veronica brevistylis Moris in Mem. Reale Accad. Sci. Torino Ser. 2, 2: 301. 1840, as brevistyla). Etymology: the epithet is an equivalent of the replaced epithet brevistylis, from the Greek kolos: docked, curtailed, shortened, or stunted (Brown 1956). Veronica corriganii (Carse) Garn.-Jones, comb. nov. ≡ Hebe corriganii Carse in Trans. & Proc. New Zealand Inst. 60: 573. 1930. Veronica cryptomorpha (Bayly, Kellow, G. Harper & Garn.-Jones) Garn.-Jones, comb. nov. ≡ Hebe cryptomorpha Bayly, Kellow, G. Harper & Garn.-Jones in New Zealand J. Bot. 40: 596. 2002. Veronica decora (Ashwin) Garn.-Jones, comb. nov. ≡ Parahebe decora Ashwin, in Allan, Fl. N. Zeal. 1: 877. 1961. Garnock-Jones & al. • Southern Hemisphere Veronica Veronica hectorii subsp. demissa (G. Simpson) Garn.Jones, comb. nov. ≡ Hebe demissa G. Simpson in Trans. & Proc. Roy. Soc. New Zealand 75: 193. 1945. Veronica hookeri (Buchanan) Garn.-Jones, comb. nov. ≡ Mitrasacme hookeri Buchanan in Trans. & Proc. New Zealand Inst. 14: 348 (text), plate 29, fig. 1. 1882. This species was previously known as Hebe ciliolata and Leonohebe ciliolata. Veronica hulkeana F. Muell. subsp. hulkeana, an autonym established by publication of subsp. evestita below. Veronica hulkeana subsp. evestita (Garn.-Jones) Garn.Jones, comb. nov. ≡ Heliohebe hulkeana subsp. evestita Garn.-Jones in New Zealand J. Bot. 31: 331. 1993. Veronica inflexa Albach, nom. nov. ≡ Hebe rigida Pennell in J. Arnold Arbor. 24: 259. 1943 (non Veronica rigida Turrill in Kew Bull. 1922: 186. 1922). Etymology: The replacement epithet refers, like the original, to the stiffly erect stems. Veronica ionantha Albach, nom. nov. ≡ Hebe ciliata Pennell in J. Arnold Arbor. 24: 258, pl. IV A. 1943 (non Veronica ciliata Fisch. in Mém. Soc. Imp. Naturalistes Moscou 3: 56. 1812, nec Schloss. & Vuk., Syll. Fl. Croat.: 90. 1857. Etymology: ion, mauve, anthos, flower, a reference to the flower colour (van Royen, 1983). Veronica flavida (Bayly, Kellow & de Lange) Garn.Jones, comb. nov. ≡ Hebe flavida Bayly, Kellow & de Lange, in Bayly & Kellow, Illustrated Guide New Zealand Hebes, 312. 2006. Veronica kellowiae Garn.-Jones, nom. nov. ≡ Hebe ramosissima G. Simpson & J.S. Thomson in Trans. & Proc. Roy. Soc. New Zealand 72: 29. 1943 (non Veronica ramosissima Boriss. in Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk. SSSR 17: 351. 1955). Etymology: named for Alison Valerie Kellow in recognition of her contributions to knowledge of New Zealand Veronica (e.g., Bayly & Kellow, 2006), in particular the “Connatae” to which this species belongs. Veronica hectorii Hook. f. subsp. hectorii, an autonym that is established by the publication of other subspecies combinations (below). Veronica macrocalyx J.B. Armstr. var. macrocalyx, an autonym established by publication of var. humilis (below). Veronica hectorii subsp. coarctata (Cheeseman) Garn.Jones, comb. nov. ≡ Veronica coarctata Cheeseman, Man. New Zealand Fl.: 531. 1906, pro parte. Veronica macrocalyx var. humilis (G. Simpson) Garn.Jones, comb. nov. ≡ Hebe macrocalyx var. humilis G. Simpson in Trans. & Proc. Roy. Soc. New Zealand 79: 428. 1952. Veronica dilatata (G. Simpson & J.S. Thomson) Garn.Jones, comb. nov. ≡ Hebe dilatata G. Simpson & J.S. Thomson in Trans. & Proc. Roy. Soc. New Zealand 73: 164. 1943. 577 Garnock-Jones & al. • Southern Hemisphere Veronica Veronica masoniae (L.B. Moore) Garn.-Jones, comb. nov. ≡ Hebe pauciramosa var. masoniae L.B. Moore, in Allan, Fl. N. Zeal. 1: 926. 1961. Veronica melanocaulon Garn.-Jones, nom. nov. ≡ Parahebe catarractae subsp. martinii Garn.-Jones in New Zealand J. Bot. 18: 295. 1980 ≡ Parahebe martinii (Garn.-Jones) Garn.-Jones in New Zealand J. Bot. 42: 217. 2004 (non V. martini H. Lév. in Bull. Soc. Agric. Sarthe, ser. 2, 39: 325. 1904). Etymology: the epithet is a reference to the dark purplish-black stems of most plants. Veronica mooreae (Heads) Garn.-Jones, comb. nov. ≡ Leonohebe mooreae Heads in Newslett. Bot. Soc. Otago 5: 10. 1987. Veronica murrellii (G. Simpson & J.S. Thomson) Garn.Jones, comb. nov. ≡ Hebe murrellii G. Simpson & J.S. Thomson in Trans. & Proc. Roy. Soc. New Zealand 73: 165. 1943. Veronica myosotoides (Ashwin) Garn.-Jones, comb. nov. ≡ Pygmea myosotoides Ashwin in Allan, Fl. N. Zeal. 1: 873, 975. 1961 – Holotype: Mt Pisa, Otago, 6000 ft., Petrie, AK 8334. Veronica notialis Garn.-Jones, nom. nov. ≡ Hebe pauciflora G. Simpson & J.S. Thomson in Trans. & Proc. Roy. Soc. New Zealand 73: 166. 1943 [non Veronica pauciflora Font Quer in Cavanillesia 4: 54. 1931 in obs., nec Link in Jahrb. Gewächsk. 1(3): 42. 1820]. Etymology: notialis, southern, a reference to its distribution in the far south-west of New Zealand. Veronica ochracea (Ashwin) Garn.-Jones, comb. nov. ≡ Hebe ochracea Ashwin in Allan, Fl. N. Zeal. 1: 936. 1961. Veronica papuana (P. Royen & Ehrend.) Albach, comb. nov. ≡ Parahebe papuana P. Royen & Ehrend. in Bot. Jahrb. Syst. 91: 395. 1972. Veronica pareora (Garn.-Jones & Molloy) Garn.-Jones, comb. nov. ≡ Hebe pareora Garn.-Jones & Molloy in New Zealand J. Bot. 20: 398. 1983. Veronica pauciramosa (Cockayne & Allan) Garn.-Jones, comb. nov. ≡ Hebe buxifolia var. pauciramosa Cockayne & Allan in Trans. & Proc. New Zealand Inst. 56: 27. 1926. Veronica pentasepala (L.B. Moore) Garn.-Jones, comb. 578 TAXON 56 (2) • May 2007: 571–582 nov. ≡ Hebe raoulii var. pentasepala L.B. Moore in Allan, Fl. N. Zeal. 1: 942, 972. 1961. Veronica perbella (de Lange) Garn.-Jones, comb. nov. ≡ Hebe perbella de Lange in New Zealand J. Bot. 36: 399. 1998. Veronica phormiiphila Garn.-Jones, nom. nov. ≡ Veronica salicifolia var. paludosa Cockayne in Trans. & Proc. New Zealand Inst. 48: 202. 1916 (non Veronica paludosa Lej., Fl. Spa 1: 22. 1812). Etymology: the name means Phormium-loving, refering to the habitat of this species in wet sites often associated with Phormium tenax. Veronica pimeleoides Hook. f. subsp. pimeleoides, an autonym established by the publication of subsp. faucicola below. Veronica pimeleoides subsp. faucicola (Kellow & Bayly) Garn.-Jones, comb. nov. ≡ Hebe pimeleoides subsp. faucicola Kellow & Bayly in New Zealand J. Bot. 41: 242. 2003. Veronica pubescens Banks & Sol. ex Benth. subsp. pubescens, an autonym established by publication of the subspecies below. Veronica pubescens subsp. rehuarum (Bayly & de Lange) Garn.-Jones, comb. nov. ≡ Hebe pubescens subsp. rehuarum Bayly & de Lange in New Zealand J. Bot. 41: 40. 2003. Veronica pubescens subsp. sejuncta (Bayly & de Lange) Garn.-Jones, comb. nov. ≡ Hebe pubescens subsp. sejuncta Bayly & de Lange in New Zealand J. Bot. 41: 42. 2003. Veronica punicea Garn.-Jones, nom. nov. ≡ Veronica speciosa var. brevifolia Cheeseman, Man. New Zealand Fl.: 500. 1906 (non V. brevifolia M. Bieb., Fl. Taur.-Caucas. 1: 6. 1808, nec Link, Enum. Hort. Berol. Alt. 1: 21. 1821). Etymology: a reference to the unusual purplish red corolla, in New Zealand otherwise seen only in V. speciosa. Veronica raoulii Hook. f. subsp. raoulii, an autonym established by publication of subsp. maccaskillii below. Veronica raoulii subsp. maccaskillii (Allan) Garn.Jones, comb. nov. ≡ Hebe raoulii var. maccaskillii Allan in Trans. & Proc. Roy. Soc. New Zealand 69: 273. 1940. TAXON 56 (2) • May 2007: 571–582 Garnock-Jones & al. • Southern Hemisphere Veronica Veronica rigidula Cheesem. var. rigidula, an autonym established by publication of var. sulcata below. var. hesperia Bayly & Garn.-Jones in New Zealand J. Bot. 38: 180. 2000. Veronica rigidula var. sulcata (Bayly & Kellow) Garn.Jones, comb. nov. ≡ Hebe rigidula var. sulcata Bayly & Kellow in New Zealand J. Bot. 40: 585. 2002. Veronica stenophylla var. oliveri (Bayly & Garn.-Jones) Garn.-Jones, comb. nov. ≡ Hebe stenophylla var. oliveri Bayly & Garn.-Jones in New Zealand J. Bot. 38: 182. 2000. Veronica rivalis Garn.-Jones, nom. nov. ≡ Veronica acutiflora Benth. in DC., Prodr. 10: 460. 1846, nom illeg. (non Lapeyr. ex Roem. & Schult., Syst. Veg. 1: 112. 1817). Etymology: rivalis, Latin for a neighbour sharing a river or brook (Brown, 1956), a reference to its habitat and to its narrow distribution. Veronica scopulorum (Bayly, de Lange & Garn.-Jones) Garn.-Jones, comb. nov. ≡ Hebe scopulorum Bayly, de Lange & Garn.-Jones in New Zealand J. Bot. 40: 586. 2002. Veronica scrupea Garn.-Jones, nom. nov. ≡ Heliohebe acuta Garn.-Jones in New Zealand J. Bot. 31: 326. 1993 (non Veronica acuta Mart., Pl. Hort. Erlang.: 12. 1814). Etymology: the epithet means “of sharp stones” (Brown, 1956) and is a reference to its habitat on cliffs of shattered greywacke. Veronica senex (Garn.-Jones) Garn.-Jones, comb. nov. ≡ Parahebe senex Garn.-Jones in New Zealand J. Bot. 42: 220. 2004. Veronica simulans Garn.-Jones, nom. nov. ≡ Hebe crenulata Bayly, Kellow & de Lange in New Zealand J. Bot. 40: 592. 2002 (non V. crenulata Hoffm., Phytogr. Bl. 1: 95. 1803, nec Sessé & Moç. in Naturaleza (Mexico City) ser. 2, 2(2): 5. 1892. Etymology: from Latin, simulans (similar), a reference to its close similarity to V. cryptomorpha. Veronica societatis (Bayly & Kellow) Garn.-Jones, comb. nov. ≡ Hebe societatis Bayly & Kellow in New Zealand J. Bot. 40: 576. 2002. Veronica spectabilis (Garn.-Jones) Garn.-Jones, comb. nov. ≡ Parahebe spectabilis Garn.-Jones in New Zealand J. Bot. 42: 223. 2004. Veronica stenophylla Steud. var. stenophylla, an autonym established by publication of the varieties below. Veronica stenophylla var. hesperia (Bayly & Garn.Jones) Garn.-Jones, comb. nov. ≡ Hebe stenophylla Veronica stricta Banks & Sol. ex Benth. var. stricta, an autonym established by publication of the varieties below. We include V. salicifolia var. atkinsonii Cockayne in its circumscription. Veronica stricta var. egmontiana (L.B. Moore) Garn.Jones, comb. nov. ≡ Hebe stricta var. egmontiana L.B. Moore in Allan, Fl. N. Zeal. 1: 907. 1961. Veronica stricta var. lata (L.B. Moore) Garn.-Jones, comb. nov. ≡ Hebe stricta var. lata L.B. Moore in Allan, Fl. N. Zeal. 1: 907, 1961. Veronica stricta var. macroura (Hook. f. ex Benth.) Garn.-Jones, comb. nov. ≡ Veronica macroura Hook. f. ex Benth. in DC., Prodr. 10: 459. 1846. Veronica strictissima (T. Kirk) Garn.-Jones, comb. nov. ≡ Veronica parviflora var. strictissima T. Kirk in Trans. & Proc. New Zealand Inst. 28: 527. 1896. Veronica strigosa Albach, nom. nov. ≡ Hebe tenuis Pennell in J. Arnold Arbor. 24: 259, pl. Vb. 1943 (non Veronica tenuis Ledeb., Fl. Altaic. 1: 38. 1829). Etymology: The replacement epithet has the same meaning as the original, and refers to the slender habit. Veronica subfulvida (G. Simpson & J.S. Thomson) Garn.Jones, comb. nov. ≡ Hebe subfulvida G. Simpson & J.S. Thomson in Trans. & Proc. Roy. Soc. New Zealand 73: 163. 1943. This species was previously known as Hebe divaricata (Cheeseman) Cockayne & Allan, non Veronica divaricata Boiss. & Balansa. Veronica tairawhiti (B.D. Clarkson & Garn.-Jones) Garn.-Jones, comb. nov. ≡ Hebe tairawhiti B.D. Clarkson & Garn.-Jones in New Zealand J. Bot. 34: 51. 1996. Veronica tetragona Hook. f. subsp. tetragona, an autonym established by publication of subsp. subsimilis below. Veronica tetragona subsp. subsimilis (Colenso) Garn.579 TAXON 56 (2) • May 2007: 571–582 Garnock-Jones & al. • Southern Hemisphere Veronica Jones, comb. nov. ≡ Veronica subsimilis Colenso in Trans. & Proc. New Zealand Inst. 31: 278. 1899. Veronica topiaria (L.B. Moore) Garn.-Jones, comb. nov. ≡ Hebe topiaria L.B. Moore in Allan, Fl. N. Zeal. 1: 917. 1961. Veronica treadwellii (Cockayne & Allan) Garn.-Jones, comb. nov. ≡ Hebe treadwellii Cockayne & Allan in Trans. & Proc. New Zealand Inst. 56: 27. 1926. Veronica urvilleana (W.R.B. Oliv.) Garn.-Jones, comb. nov. ≡ Hebe urvilleana W.R.B. Oliv. in Rec. Domin. Mus. 1: 212. 1944. Veronica wilhelminensis Albach, nom. nov. ≡ Hebe polyphylla Pennell in J. Arnold Arbor. 24: 257. 1943 (non Veronica polyphylla Colenso in Trans. & Proc. New Zealand Inst. 31: 277. 1899). Etymology: a replacement epithet that refers to the locality (Mt Wilhelmina) to which this species is restricted. Veronica zygantha Garn.-Jones, nom. nov. ≡ Veronica laxa G. Simpson & J.S. Thomson in Trans. & Proc. Roy. Soc. New Zealand 72: 32. 1942, nom. illeg. (non Veronica laxa Benth., Scroph. Ind.: 45. 1835). Etymology: zygos a yoke or pair, anthos a flower: a reference to the paired flowers in this species. Veronica derwentiana Andrews subsp. derwentiana, an autonym established by publication of the subspecies below. Veronica derwentiana subsp. anisodonta (B.G. Briggs & Ehrend.) B.G. Briggs, comb. nov. ≡ Derwentia derwentiana subsp. anisodonta B.G. Briggs & Ehrend. in Telopea 5: 273. 1992. Veronica derwentiana subsp. homalodonta (B.G. Briggs & Ehrend.) B.G. Briggs, comb. nov. ≡ Derwentia derwentiana subsp. homalodonta B.G. Briggs & Ehrend. in Telopea 5: 272. 1992. Veronica derwentiana subsp. maideniana (Gand.) B.G. Briggs, comb. nov. ≡ Veronica maideniana Gand. in Bull. Soc. Bot. France 66: 220. 1919. Veronica derwentiana subsp. subglauca (B.G. Briggs & Ehrend.) B.G. Briggs, comb. nov. ≡ Derwentia derwentiana subsp. subglauca B.G. Briggs & Ehrend. in Telopea 5: 275. 1992. Veronica lithophila (B.G. Briggs & Ehrend.) B.G. Briggs, comb. nov. ≡ Parahebe lithophila B.G. Briggs & Ehrend. in Telopea 5: 255. 1992. Veronica velutina (B.G. Briggs & Ehrend.) B.G. Briggs, comb. nov. ≡ Derwentia velutina B.G. Briggs & Ehrend. in Telopea 5: 276. 1992. Veronica sect. Labiatoides In addition to the new combinations listed below, the following species, the names of which are already published in Veronica, belong to this section: Veronica arenaria A. Cunn. ex Benth, Veronica brownii Roem. & Schult., Veronica calycina R. Br., Veronica continua B.G. Briggs, Veronica decorosa F. Muell., Veronica distans R. Br., Veronica formosa R. Br., Veronica gracilis R. Br., Veronica grosseserrata B.G. Briggs & Ehrend., Veronica hillebrandii F. Muell., Veronica nivea Lindl., Veronica notabilis F. Muell. ex Benth., Veronica novaehollandiae Poir., Veronica parnkalliana J.M. Black, Veronica perfoliata R. Br., Veronica plebeia R. Br., Veronica sobolifera B.G. Briggs & Ehrend., and Veronica subtilis B.G. Briggs & Ehrend. (Briggs & Ehrendorfer, 2006b). Veronica arcuata (B.G. Briggs & Ehrend.) B.G. Briggs, comb. nov. ≡ Derwentia arcuata B.G. Briggs & Ehrend. in Telopea 5: 279. 1992. Veronica blakelyi (B.G. Briggs & Ehrend.) B.G. Briggs, comb. nov. ≡ Derwentia blakelyi B.G. Briggs & Ehrend. in Telopea 5: 278. 1992. 580 Addendum In the course of preparing this manuscript, we noticed that the inclusion of Hebe in Veronica not only has nomenclatural effects in the Southern Hemisphere. Apparently, the valid publication of Veronica propinqua by Cheeseman (1906) was overlooked by Borissova (1955) whose V. propinqua was subsequently used in publications by Albach & al. (2004a, 2005a). Veronica vendettadeae Albach, nom. nov. ≡ Veronica propinqua Boriss. in Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk. SSSR 17: 351. 1955 (non V. propinqua Cheeseman, Man. New Zealand Fl.: 533. 1906) – Holotype: Dagestan. In pratis alpinis inter Zatanik & mont. Kanschta-balata July 1898, Prof. N. Kusnezow; holo: LE (n.v.), photo E (!); iso: TBI (!). Etymology: vendetta, revenge, deae, of the goddess. It seems to us like it is the revenge of goddess Hebe that the sinking of genus Hebe not only has nomenclatural effects on species in Hebe but also on species in the traditional genus Veronica. TAXON 56 (2) • May 2007: 571–582 ACKNOWLEDGEMENTS Mike Bayly and Alison Kellow most generously made available to us in advance of its publication (Bayly & Kellow, 2006) all the information they have accumulated on names and types in Veronica sect. Hebe. Friedrich Ehrendorfer contributed greatly to the delimitation and typification of the Australian species. We thank Steven Wagstaff, John McNeill, Elvira Hörandl, and an anonymous reviewer for sound advice and constructive criticism of the manuscript. PG-J thanks the New Zealand Foundation for Research, Science and Technology (contract MNZ601) for funding. LITERATURE CITED Albach, D.C. & Chase, M.W. 2001. Paraphyly of Veronica (Veroniceae; Scrophulariaceae): evidence from the internal transcribed spacer (ITS) sequences of nuclear ribosomal DNA. 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