BIBLIOGRAPHY & ANNOTATED CHECKLIST of LIVING ORGANISMS FIRST NAMED from the G R A N D C A N Y O N and V I C I N I T Y (NORTHWESTERN ARIZONA) compiled and edited by EARLE E. SPAMER _____________________________________________________________________________________ COVER ILLUSTRATION Ostrya knowltonii Coville, 1894 (Betulaceae). Coville’s original illustration of the holotype (US) from the top of the Old Hance Trail, Grand Canyon, Arizona, gathered by J. W. Toumey, 10 July 1892. (See No. 84 herein, pp. 42–43.) BACK COVER General view of the type locality of O. knowltonii, looking downward at the top of the Old Hance Trail. Photo by Nancy Brian during the visit that rediscovered the locality, 23 May 1997. BIBLIOGRAPHY & ANNOTATED CHECKLIST of LIVING ORGANISMS FIRST NAMED from the G R A N D C A N Y O N and V I C I N I T Y (NORTHWESTERN ARIZONA) compiled and edited by EARLE E. SPAMER RAVE N ’ S P E RCH ME DIA PHILADELPHIA 1 2022 Copyright © Earle E. Spamer 2022 This publication is a resource manager’s and historian’s reference work documenting species-level taxa of living organisms that were first scientifically named based wholly or in part on collections made at, in, or near the Grand Canyon. This is not a work of systematics nor of taxonomic revision. Taxonomic combinations are those of the original authors or authorities; subsequent revisions of taxonomy and typification are not cited. Publications and pertinent taxa meeting the purpose of the checklist are as known to the compiler as of November 2022. RAVEN’S PERCH MEDIA, Philadelphia, Pennsylvania https://ravensperch.org Bibliographical and Historical Resources in the Grand Canyon and Lower Colorado River Regions of the United States and Mexico Published 11 DECEMBER 2022 REPRODUCTION AND FAIR USE • No commercial or for-profit use of this work is allowed without the author’s permission. Not-for-profit organizations, educational institutions, government agencies, and Indigenous communities may with credit use this work for purposes of resource management and interpretation, education, and public outreach; they also may download the entire document for cataloging and inclusion in their digital products collections, and they may freely allow users to copy this document under the conditions stated here. Reuse of this work, in whole or in part, for any purpose, must cite author, title, publisher, and notice of copyright. Academic or other individual researchers may copy selections for personal reference and may quote from this work following best practices of acknowledgment in scholastic research. This statement does not pertain to, nor supersedes, copyrights of items that still may be in effect among the items cited in the bibliography. 2 CONTENTS [In the PDF document, each line in the contents list is a hyperlink] Map of Approximate Area of Coverage For This Checklist 6 Introduction 7 Commemorative Names 9 Conventions and Styles Used in This Checklist 12 Note on Paleontological Names 16 BIBLIOGRAPHY AND ANNOTATED CHECKLIST 19 Serials Listed in the Bibliography and Annotated Checklist 114 Taxonomic List 117 Kingdom Animalia 119 Phylum Arthropoda 119 Phylum Chordata 128 Phylum Mollusca 130 Phylum Nematoda 131 Kingdom Plantae 132 Class Magnoliopsida 132 Class Pinopsida 137 Kingdom Fungi 138 Phylum (Division) Ascomycota 138 Phylum (Division) Basidiomycota 139 Kingdom Chromista (Phylum Apicomplexa) 140 Kingdom Protista (“Phylum” Hemimastigophora) 140 3 Contents (cont’d) Localities for Type Material from Grand Canyon National Park Map of Federally-Designated Boundaries of Grand Canyon National Park and Predecessor Units (1893–present) 141 142 “Grand Canyon” not discriminated 143 South Rim 144 North Rim 150 Inner Canyon 154 Colorado River Locales 161 Localities for Type Material from Near Grand Canyon 165 Cameron, Arizona, and vicinity 166 Arizona Strip 167 Grand Canyon vicinity undiscriminated 171 Grand Canyon–Parashant National Monument 172 Havasupai Indian Reservation 174 House Rock and House Rock Valley 174 Hualapai Indian Reservation 175 Hualapai Indian Reservation vicinity (off reservation) 177 “Kaibab Forest” (north) undiscriminated 177 “Kaibab Forest” (south) undiscriminated 177 Kaibab Indian Reservation 178 Kaibab National Forest, Tusayan Ranger District 178 Kaibab Plateau (including Kaibab National Forest, North Kaibab Ranger District) 179 Kanab Canyon 184 Lees Ferry 184 Marble Canyon [not on Colorado River] 184 Navajo Nation (westernmost) 185 Red Mountain 185 Index to Nominate Taxa 187 ___________________________________________ 4 5 APPROXIMATE AREA OF COVERAGE FOR THIS CHECKLIST UTAH Kaibab Indian Reservation A R I Z O N A Lees Ferry ↘ S T R I P Kaibab National Forest (Kaibab Ranger District) ARIZONA NEVADA (north and west of the Colorado River) House Rock Valley Kaibab Plateau Western extremity of Havasupai Indian Reservation the Navajo Nation Tusayan → (Tusayan Ranger District) Base map from Google, emended 6 INTRODUCTION INTRODUCTION This checklist provides federal and Native American resource managers and historians of the Grand Canyon region of northwestern Arizona with a census of living (neontological) organisms that had been scientifically named based on collections made wholly on, or in part from, the lands that these managers oversee. Their original scientific names are recorded in this checklist. It does not use updated or otherwise “current” names for taxa that have been scientifically reidentified (taxonomically synonymized) or that were systematically transfered to other supra-specific biological groups than those under which they were first named. Providing such information is beyond the scope of a historicallyand administratively-directed document such as this one; its inclusion would soon make the records herein out of date. For the same reasons no data regarding the biological attributes (descriptions) of the organisms are included herein. Federal administrative units to which this accounting may be of particular interest are: U.S. Department of the Interior National Park Service Grand Canyon National Park Glen Canyon National Recreation Area (in that area near where it shares a boundary with the Grand Canyon park at Lees Ferry) Bureau of Land Management Arizona Strip District Joint management by NPS and BLM Grand Canyon–Parashant National Monument U.S. Department of Agriculture U.S. Forest Service Kaibab National Forest (Tusayan and North Kaibab Ranger Districts only) Jurisdictions of Native American peoples addressed herein pertain to: Havasupai Tribe [south side of the Grand Canyon] Hualapai Indian Tribe [south side of the Grand Canyon] Kaibab Band of Paiute Indians [on the Arizona Strip] Navajo Nation [to the east of the Grand Canyon] 7 INTRODUCTION A few of the collections cited herein seem to have come from privately held lands that are inholdings within the federal units, and from within the boundaries of particular municipalities in Arizona. The bibliography/checklist includes 322 species-level taxa1 named on type specimens2 collected in the greater Grand Canyon region of northwestern Arizona; 215 of those taxa are based at least in part on specimens from within, or are with good probability from within, the current boundaries of Grand Canyon National Park. Five taxonomic kingdoms are represented among them—animals, plants, fungi, photosynthetic eukaryotes, and protists. Only the name-bearing types from within the geographical boundaries of this checklist are listed; referred specimens (those not selected to be types) are generally omitted. It is probable that there are additional taxa that can be included in this checklist, but they were not discovered during the course of the several decades of work that has contributed to it, although hardly little of that work was devoted to this specific purpose. This is an opportunistic work, condensing facts for an administrative document of interest to historians, too. There is far more beyond the geographical boundaries imposed here that might still be considered to be in the greater Grand Canyon region, which are ignored for the simple reason that there must be a limit. For example, there is an equal wealth of taxonomic neology from the lands best recognized now as the Navajo Nation, which does nudge briefly across this checklist’s eastern boundary; that area is accommodated herein. Then there is the wholly different zoo and garden of the Mohave Desert to the west, which, while it is decisively segregated from the Grand Canyon region by physiography and environment, does in fact sneak into the western part of the canyon along the lower elevations of the Colorado River corridor; so a few taxa more representative of those regions are included here. This bibliography and checklist finds its origin in my own researches in history, science and bibliography of the Grand Canyon vicinity in particular, thus the geographical bounds are the same. 1 2 “Species-level taxon” means any name that is at the taxonomic rank of species, subspecies, or variety. [The count includes No. 244a, added late.] “Type specimens,” more generally called “types”, are the very specimens that were selected by a species-level taxon’s author as bearing the formal characteristics that uniquely identify the taxon. There may be just one, or even hundreds, and they can be from different localities (even beyond the geographical area of interest in this checklist). 8 INTRODUCTION The checklist is also confined taxonomically to the original names of the organisms scientifically described for the first time by the authors cited in the bibliography. These names can disappear through taxonomic synonymy (and in many cases they have) when researchers in biological systematics have ascertained that one or another organism is actually the same as another, earlier named one, or if they move species-level taxa to different taxa than those in which they were originally placed. And in turn, genera can be forever “fiddled with”, moving them from one Family to another and often carrying its contained species with it (though this is a bit simplified); and similarly among the higher taxa, clear up to the Kingdom level. In some measure this is reflected in the highersystematics listed with the taxa herein, which information may not be the same as that originally assigned by the authors of these taxa. There is also a chance that some organisms can disappear actually, through extirpation or extinction; and this checklist thus would provide the original data that were communicated about their presence and scientific discovery. Ecological memory is ephemeral, but taxonomic memory—right, wrong, and revised—is forever, part of the resource manager’s treasury. Thus this checklist serves these managers as a documentary list of organisms first named from the lands that they oversee and the resources that have been used, and it serves as a contribution to the history of the Grand Canyon and adjacent areas of northwestern Arizona. Commemorative Names When a researcher names a new taxon they usually apply a “scientific” name that describes some feature or attribute of the organism to which the name is given. Other times, one or more persons are honored with the epithet, which customarily has one of the common Latin gender-specific suffixes, -i, -ii, -ae, or -orum. (For example, in this checklist see Ostrya knowltoni Coville, 1894 [No. 84 in the sequentially enumerated checklist, and also the cover illustration], an ironwood from the Old Hance Trail, named for Frank H. Knowlton.3 3 The epithet was later corrected to knowltonii. This was a grammatical correction only, which does not affect the author’s name. Grammatical corrections do not affect the epithet, being mandated alterations to the Latin suffix only, such as this change from -i to -ii, so the author’s name is not placed in parentheses. (When a species is moved to a different genus, or a subspecies to a different species, than the ones under which they were first named, the author’s name is placed within parentheses. An example of a transfer, from among the listings herein, is Crotalus confluentus abyssus Klauber, 1930 [No. 183], the “Grand Canyon Rattlesnake”, which was later redescribed as Crotalus viridis abyssus (Klauber), and has since also been placed in the genus Oreganus.) 9 INTRODUCTION Often, a place is commemorated, usually with the suffix -ensis or another, grammatically figured suffix like -ense; other formulations are used, too. A few examples from this checklist are: 238 Oreohelix yavapai angelica Pilsbry & Ferriss, 1911; a land snail from Bright Angel Trail 77 Sclerocactus havasupaiensis Clover, 1942; a cactus from “Havasupai Canyon” (Havasu Canyon) 158 Opuntia hualpaensis Hester, 1943; a cactus from Route 66 between Peach Springs and Hyde Park, the name denoting the Hualapai Indian Reservation 245 Panicum mohavense Reeder, 1991; a grass from Mohave County, on the Arizona Strip 109 Echinocereus triglochidiatus var. toroweapensis Fischer, 1991; a cactus from Toroweap Point A specially commemorative name may also acknowledge a regional feature closely associated with the collecting locality; in these two examples, the Colorado River 4: 179 Astragalus kentrophyta var. coloradoensis Jones, 1902; an herb from Lees Ferry 280 Helix (Arionta) coloradoensis Stearns, 1890, a land snail from the Old Hance Trail [the taxon is now Sonorella coloradoensis (Stearns)] In the case of the Grand Canyon itself, seven species-level taxa have been erected with epithets that commemorate the canyon or denote the canyon’s attribute as an “abyss”: 235 Rosa stellata Wooton subsp. abyssa Phillips, 1992; a rose from Twin Point, Shivwits Plateau 159 Opuntia abyssi Hester, 1943; a cactus from Peach Springs Canyon 183 Crotalus confluentus abyssus Klauber, 1930; a rattlesnake from Tanner Trail (the “Grand Canyon Rattlesnake”) 270 Melanoplus canonicus Scudder, 1897; a grasshopper from Grand Canyon (no specific locale) 54 Aphelonema convergens var. canyonensia Bunn, 1930; a planthopper from Grand Canyon (no specific locale) 267 Mentzelia canyonensis Schenk, Hodgson & Hufford, 2013; a loasa from Manzanita Canyon 166 Arenivaga grandiscanyonensis Hopkins, 2014; a sand cockroach from Colorado River Mile 211.5 Right 4 When searching for examples like these, bear in mind that such names are contextual. While Misumenops coloradensis Gertsch, 1933 is listed in this checklist [No. 113] for its paratype specimens from the Grand Canyon, it is not included in this list of examples because the epithet, coloradensis, was based on the primary type material having come from the State of Colorado. 10 INTRODUCTION The Kaibab Plateau is the most commonly commemorated place in this checklist, with 24 taxa named based on at least one type specimen. They are chiefly from the national forest, but a few are from within the boundary of Grand Canyon National Park. “Kaibab” is introduced into a Latinized epithet in various ways (not all of which may be in “good” Latin, depending upon the author’s decision or the context of the name). These are, in alphabetical order by species-level taxon: 117 Loxosceles kaiba Gertsch & Ennik, 1983 217 Agave kaibabensis McKelvey, 1949 253 Andrena (Scaphandrena) kaibabensis Ribble, 1974 48 Branchinecta kaibabensis Belk & Fugate, 2000 164 Catilleja kaibabensis N. Holmgren, 1973 172 Cicindela pussila kaibabensis Johnson, 1990 74 Cordylanthus wrightii kaibabensis Chuang & Heckard, 1986 108 Coryphantha vivipara var. kaibabensis Fischer, 1979 182 Enderleinellus kaibabensis Kim, 1966 227 Felis concolor kaibabensis E. Nelson & Goldman, 1931 258 Lesquerella kaibabensis Rollins, 1982 50 Mischocyttarus flavitaris var. kaibabensis Bequaert, 1932 42 Morchella kaibabensis Beug, T.A. Clem. & T.J. Baroni in Baroni et al., 2018 248 Morsea california kaibabensis Rehn & Grant, 1958 45 Papilio indra kaibabensis Bauer, 1955 115 Phidippus kaibabensis Gertsch, 1934b 155 Polistes canadensis var. kaibabensis Lynn, 1932 220 Sciurus kaibabensis Merriam, 1904 252 Scutellaria potosina var. kaibabensis Rhodes & Ayers, 2010 288 Suillus kaibabensis Thiers, 1976 123 Thomomys fossor kaibabensis Goldman, 1938 3 Tipula (Lunatipula) kaibabensis C. Alexander, 1946 223 Saxinis saucia kaibabiae Moldenke, 1970 180 Astragalus kaibensis Jones, 1902 11 INTRODUCTION Conventions and Styles Used in This Checklist The information given here is repeated at the heads of pertinent parts of this volume. Throughout the volume, Red Numbers indicate and cross-list the sequentially enumerated taxa in the bibliography. BIBLIOGRAPHY AND ANNOTATED CHECKLIST In this checklist the species-level taxa are the names of organisms are as originally erected by the authors. This is a historical record of nomenclature, not one of taxonomic revision. For convenience, the systematic levels of phylum, class, order, and family are listed for each enumerated taxon. Refer to the Taxonomic List herein for other details.  Arrow indicates taxa for which at least one locality reported for its types is in, or with good probabability within, Grand Canyon National Park. Refer to the bibliography for all details. Data were not often provided by the original authors in any concise or systematic fashion, thus direct quotations have been preferred in this checklist. By quoting, the user can grasp either the paucity or breadth of data that were conveyed in the original author’s locality and collecting information for the taxon there named. The information has not been editorially revised herein to create an artificial sense of conformity; doing so might obscure original precision, inconsistencies, or confusing data. Particular localities that lie within the geographical boundaries of this checklist are displayed in bold in order to bring attention to them, often amidst more detailed data. Refer to the respective separate lists of localities in this volume for other details. Data regarding the collectors and dates of collection are provided as published by the original author, usually as part of the quoted information. However, take note that many nominate taxa are not necessarily erected based on collections by the author themself, but on specimens sent to the researcher or are previously collected materials that were examined in museum collections. The published information thus records the data that appeared with the examined specimens—although this distinction is not always perfectly clear. Information regarding the type specimens themselves is generally as published by the taxon’s author. The particular codes of taxonomic nomenclature are followed here; there are independent codes for the animal and botanical kingdoms. In this checklist, zoological type specimens are indicated as holotype, paratype, allotype, and syntype; botanical type 12 INTRODUCTION specimens are indicated as holotype, isotype, syntype, and paratype. 5 Other terms (like topotype) are unessential and have no formal standing. Matters pertaining to lecto- or neotypifications and similar procedural acts, which are made after the original publication of a taxon, are beyond the scope of this checklist. In a few instances, older terminology was used by an author and is updated as needed herein as a parenthetical note; for example, cotype (= syntype in zoology, or used erroneously in botany to indicate an isotype or paratype). Genus and higher taxonomic levels have no type material, although older literature may indicate such terms as “genotype”, which refers to the “type species” (one species on which the description of a genus is based); the term has occasionally been used incorrectly to denote a particular specimen. When the checklist indicates only para- or isotypical material as having been collected at a locality within the bounds of the checklist, a note is usually prefixed to the quotation to indicate where the holotype (and if applicable, allotype) material was collected; so indicated for the informational value that indicates these primary types were collected elsewhere. In the checklist’s quoted data, certain interpretive remarks are inserted within [square brackets], introduced by the compiler. These usually clarify that the term “Type” or “the type” refers to the holotype specimen; and when other specimens are cited they may be interpreted to be (though not specifically mentioned as) material that is paratypical (in zoology) or iso- or paratypical (in botany). In older literature, prior to the establishment of formal codes of taxonomic nomenclature, later researchers are generally more liberal in the acceptance of type status for material that supplements the holotype, except in those cases where the original author may have precisely identified which specimens are types. 5 “Type specimens,” more generally called “types”, are the very specimens that were selected by a species-level taxon’s author as bearing the formally defined physical characteristics that uniquely identify the taxon. In older published descriptions, prior to the creation of the various international conventions for taxonomic nomenclature, a time when types were not always specially designated, it is the surviving elements known to have been used by the author in the original description that now represent the type material for a given taxon. “Holotype,” allotype,” and “isotype” are so-called primary types; other types supplement them in defining a taxon’s characteristics in such a way that characteristics not well shown in one specimen may be exhibited by another. “Paratype” has different meanings in zoology and botany: in zoology it is applied to one or more specimens that supplement the description of the name-bearing holotype; in botany it is applied to one or more specimens that were not formally specified to be type material but which were cited along with the type material. The latter procedure is different from that followed in zoology, where specimens not formally identified as type material (usually listed as “examined” or “referred” specimens) have no status at all as types. 13 INTRODUCTION Today the criteria are more rigorously applied; type material for new species-level taxa must be uniquely identified. When repository catalog numbers were published by the original author, they appear as part of the quoted material. However, since this is a nomenclatural checklist only, no attempt has been made to canvass repositories to identifiy either the continued existence of specimens or their catalog numbers. Standard repository acronyms are not spelled out. Throughout this checklist all text within [square brackets] has been inserted by the compiler, except where noted otherwise. TAXONOMIC LIST This is a nomenclatural record. Taxonomic combinations are as named by the original authors; later taxonomic reassignments, if any, are not generally noted herein. The checklist for the greater Grand Canyon region lists 322 taxa first named from collections in this area of northwestern Arizona, 215 of which were named from collections made within, or with good proabability within, the present boundaries of Grand Canyon National Park. [No. 244a was added late, thus the whole-number count stops at 321.] Higher systematics for the genus to which the nominate taxa were assigned are given here, as derived from the Integrated Taxonomic Information System (ITIS) online in 2022. Taxa not included in ITIS have had their higher systematics derived from other sources, including the National Center for Biotechnology Information Taxonomy Browser (National Library of Medicine) and the World Flora Online. The systematic placements listed here may not reflect the most current views, which are in any case bound to vary between researchers. Again, this is a resource manager’s checklist to nominate taxa, not one of systematics or revised nomenclature. In any case, for a species to be reassigned to another family does not happen often, except in significant works of systematic revision, especially those that erect new families. The taxonomic ranks cited in this list are: Phylum or Division: Class: Order: Family. Users of this bibliography and checklist may note that in some titles certain systematic groups do not correspond to the groups that are listed for each of the taxa cited from a given publication. This generally reflects a revised pespective of familial and higher groupings; again, as usually derived for this checklist from ITIS and other sources.  Arrow indicates taxa for which at least one locality reported for its types is within, or is with good probability from within, the boundaries of Grand Canyon National Park. Refer to the bibliography for all details. 14 INTRODUCTION The taxa are not listed in so-called “systematic order” because most users of this checklist will not likely be attuned to the scientifically proper ordering of taxa within each Phylum, Class, and so forth. To accommodate these more general users, who are more likely to be resource managers and not necessarily practicing biologists, all higher taxa are listed strictly in alphabetical order (the order of Kingdoms is arranged only for convenience). The nominate taxa within each category are listed in the order in which they appear in the checklist, arranged numerically to facilitate finding them in that bibliograph/checklist. Common names, mostly for the Families, are as listed by ITIS or other sources. They are not regularly adopted by all users and have been added here only a matter of convenience for those who might be more comfortable referring only to common names. LOCALITY LISTS Two lists of localities where type material was collected are provided herein; one for localities within the present boundaries of Grand Canyon National Park, the other for localities outside of the park and within the bounds defined for this checklist (see map). Localities for the collection of type material are as identified by the authors of the taxa. The bibliography/checklist quotes the authors’ data so as to record exactly how the data were reported. The arrangement of the localities in these lists are, however, in some measure modified for geographical and contextual sense; but specific determinants as communicated by the authors of the taxa are retained (for example, differences in altitude at the same general locality). In a few instances, sufficient historical or bibliographical data have made it possible to locate a place with greater precision, which information is appended to the basic locality data, and thus the data for a locality from which some type material came has been refined herein. Many localities are identified only to very broad geographical constraints—simply “Grand Canyon”, for example, as indicated mostly in older literature—which reflects the original data. In such cases no specifically identifiable type localities can be discriminated. At the most general, the only locality information is “Grand Canyon”, which does not distinguish either rim or inner-canyon locales. Except for the earliest years, it is unfortunate that the records of “Grand Canyon, Ariz.” have not distinguished whether a collection was made at Grand Canyon village (the place name, “Grand Canyon, Arizona”, which unofficially sometimes also capitalizes “Village”) or more generally the “Grand Canyon of Arizona”; thus taxa with this information for locality are separately listed. Only when the village area is specifically stated, or inferred from other information such as “Bright Angel, Ariz.” (which refers to the old Bright Angel Hotel), are taxa listed specifically from that locale. Similarly, potentially misleading data are recorded in localities cited as “North Rim, Grand Canyon” or similar arrangement, because there is the sometimes unofficial community name, “North 15 INTRODUCTION Rim”, which while it has been a postal address is otherwise the tourist conclave at Bright Angel Point on the North Rim. Another potentially confusing locale is “Vermilion Cliffs”, which may refer to the unofficial place name, a business called “Vermilion Cliffs”, which is along U.S. Route 89A along the base of the Vermilion Cliffs west of Marble Canyon. INDEX TO NOMINATE TAXA Names in bold indicate nominate taxa for which at least one locality reported for its types is within, or is with good probability from within, the present boundaries of Grand Canyon National Park. The names are original combinations, as published by the authors. The taxonomic phylum is appended to each name and author. Refer to the bibliography and annotated checklist for details. Note on Paleontological Names One may also ask about the record of new taxa named based on paleontological collections from the Grand Canyon and vicinity, the complement to the neontological record presented by this checklist. Those taxa are the intellectual antecedents of the living ark; in the Grand Canyon they are the lithologically embalmed life of the Proterozoic Eon to the Permian Period and the urologically indurated records of the Pleistocene, encroaching on the Recent Epoch (about 11,700 years before present). (There are also a few nearby records of fossils from the Triassic Period at Cedar Mountain, and a handful of Pliocene Epoch fossils from nearby Anita, south of the canyon.) These all have been accounted for in earlier bibliographies and checklists. Though now the publications are bit dated, lacking a comparatively small number of new taxa named since the early 1990s, they are itemized here for convenience and to alert interested users to their presence. A general bibliography of Grand Canyon paleontology (recently updated): Spamer, Earle E. 2019 Bibliography of Paleontology of the Grand Canyon Region and in the Stratigraphic Continuity of Grand Canyon Formations : compiled to commemorate the centennial of Grand Canyon National Park and National Fossil Day 2019. Raven’s Perch Media, Philadelphia, 120 pp. (PDF). Occasionally upated online at Raven’s Perch Media, https://ravensperch.org/bibliography-of-paleontology-of-the-grand-canyon-region/. 16 INTRODUCTION Historical overviews of paleontological research in Grand Canyon National Park: Spamer, Earle E. 1984 Paleontology in the Grand Canyon of Arizona: 125 years of lessons and enigmas from the late Precambrian to the present. The Mosasaur (Journal of the Delaware Valley Paleontological Society), Volume 2, pp. 45-128. 2020 History of paleontological work at Grand Canyon National Park. Up and down the long federal and NGO trails of paleontology in Grand Canyon National Park, 1858-2019. Chapter 2 in: Santucci, Vincent L., and Tweet, Justin S. (eds.), Grand Canyon National Park: centennial paleontological resource inventory (non-sensitive version). Fort Collins, Colorado: U.S. National Park Service, Natural Resource Stewardship and Science, pp. 11-44. (Volume: Natural Resource Report NPS/GRCA/NRR—2020/2103.) 2021 History of paleontological work at Grand Canyon National Park—up and down the long federal and NGO trails of paleontology in Grand Canyon National Park, 1858-2019. In: Santucci, Vincent L., and Tweet, Justin S. (eds.), Grand Canyon National Park centennial paleontological resource inventory; a century of fossil discovery and research. Utah Geological Association, Special Publication 1, pp. 11-32. [The volume is a reset and bound reprinting of the Natural Resource Report by Santucci and Tweet (2020).] Publications that contain information about paleontological taxa first named from collections made in the Grand Canyon and at nearby localities are listed below. These microform publications have since been made available as PDFs online by the Geological Society of America, https://pubs.geoscienceworld.org/books/. Spamer, Earle E. 1983 Geology of the Grand Canyon: An annotated bibliography, 1857-1982, with an annotated catalogue of Grand Canyon type fossils. (Foreword by Edwin D. McKee.) Geological Society of America, Microform Publication 13, 543 [544] pp. on six 98-frame fiche. 1984 Geology of the Grand Canyon: An annotated bibliography, with an annotated catalogue of Grand Canyon type fossils. Volume 2. Supplement to the annotated bibliography (1857-1983), Supplement and revisions to the annotated catalogue. Geological Society of America, Microform Publication 14, 229 pp. on three 98-frame fiche. [The PDF available online is absent pp. 17-48.] 1988 Geology of the Grand Canyon: An annotated bibliography, with an annotated catalogue of Grand Canyon type fossils. Volume 3. Second supplement (to 1987) with an annotated bibliography of the world literature on the Grand Canyon type fossil Chuaria circularis Walcott, 1899, an index fossil for the late Proterozoic. Geological Society of America, Microform Publication 17, 343 [355] pp. on four 98-frame fiche. 17 INTRODUCTION 1990 Geology of the Grand Canyon: An annotated bibliography, with an annotated catalogue of Grand Canyon type fossils. Volume 4. Third supplement (to 1989), with supplement to the annotated bibliography of the world literature on the Grand Canyon type fossil Chuaria circularis Walcott, 1899. Geological Society of America, Microform Publication 20, 178 pp. on two 98-frame fiche. 1992 Geology of the Grand Canyon: An annotated bibliography, with an annotated catalogue of Grand Canyon type fossils. Volume 5. Fourth supplement (to 1991) with second supplement to the annotated bibliography of the world literature on the Grand Canyon type fossil Chuaria circularis Walcott, 1899. Geological Society of America, Microform Publication 23, 234 pp. on three 98-frame fiche. 1992 The Grand Canyon fossil record; a source book on paleontology of the Grand Canyon and vicinity, northwestern Arizona and southeastern Nevada. Bibliography; indexes to taxa, stratigraphic records, localities, and repositories of type, figured, and cited specimens. Geological Society of America, Microform Publication 24, 1,008 pp. on eleven 98-frame fiche. 18 BIBLIOGRAPHY AND ANNOTATED CHECKLIST BIBLIOGRAPHY AND ANNOTATED CHECKLIST In this checklist the itemized species-level names of organisms are as originally erected by the authors. This is a historical record of nomenclature, not one of taxonomic revision. For convenience, the systematic levels of phylum/division, class, order, and family are listed for each enumerated taxon. Refer to the Taxonomic List herein for other details. Repository data for type material are as published by the original authors. The standard acronyms for these repositories, when used, are not spelled out (again, as published). Because locality data were not provided in any regular format by the original authors, with wide variances in the amount of data provided, direct quotations have been preferred in this checklist, which thus communicate the author’s (or publisher’s) style and the precision of data so conveyed. Some authors used the scientific symbols for male ( ♂) and female (♀), which uses are retained in the quotations. This information has not been editorially revised in the checklist; doing so would create an artificial sense of conformity that could obscure the original precision and overall quality of the data Throughout this checklist all text within [square brackets] has been inserted by the compiler, except where noted otherwise. ________________________________________________________________________________ FORMAT FOR THE ANNOTATED CHECKLIST Author(s) Year Bibliographical citation. SEQUENTIAL # * Taxonomic name [as published], TAXONOMIC LEVEL (page and illustration data, if from among many taxa in the publication) Taxonomic Phylum: Class: Order: Family [placed before the sequential number if there are multiple taxa under the same family] Data about types and localities, usually quoted from the original publication. Principal names of localities are highlighted in bold. (Refer to the respective separate lists of localities in this volume for other details.) Notes are added where appropriate. * An arrow  appears in this position if it is a taxon for which at least some of the type material recorded by the authors is from one or more localities that are within, or are with good probability from within, the present boundaries of Grand Canyon National Park. _________________________________________________________________________ _______ 19 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Adams, C. F. [Charles Frederick] 1903 Dipterological contributions. Kansas University Science Bulletin, [series 2], 2(2) (June): 21-47.  Ceratopogon dimidiatus, NEW SPECIES (p. 27); not illustrated 1 Arthropoda: Insecta: Diptera: Ceratopogonidae (biting midges) Syntypes: “Four specimens; Grand Canon, Arizona”; all females. ________________________________________________________________________________ Adams, James K., and Lafontaine, J. Donald 2009 A new species of Plagiomimicus Grote (Noctuidae: Stiriinae) from northern Arizona and southeastern Utah. Journal of the Lepidopterists’ Society, 63(3): 173-176, figs. 3, 4, 7, 8, 10. 2 Plagiomimicus kathyae, NEW SPECIES Arthropoda: Insecta: Lepidoptera: Noctuidae (owlet moths) Holotype male (CNC [Canadian National Collection]) and 18 paratypes (7 males, 11 females, CNC, personal collection of J. K. Adams, and USNM). Type material also includes specimens from Moab, Utah. “The precise location for collection of the Arizona specimens is the Cameron Trading Post, Cameron, AZ, at 35º 52’ 30” N, 111º 24’ 47” W, just south of the Little Colorado River and just W of Hwy. 89”. Holotype collected by J. K. Adams, 7 September 1995; paratypes collected by Adams, 7 September 1995, 1 September 1996. Text of species description (p. 174) indicates that holotype male is illustrated in fig. 4, but figure legends (p. 175) indicate holotype is in fig. 3 and a female paratype is in fig. 4. _________________________________________________________________________ _______ Alexander, Charles P. 1946 Records and descriptions of North American crane-flies (Diptera). Part VI. Tipuloidea of Arizona, New Mexico and Trans-Pecos Texas, 1. American Midland Naturalist, 35(2) (March): 484-531. 3  Tipula (Lunatipula) kaibabensis, NEW SPECIES (pp. 506-508, fig. 7 [p. 507]) Arthropoda: Insecta: Diptera: Tipulidae (crane flies) Holotype male from “Kaibab Plateau, north rim of the Grand Canyon, Arizona [undiscriminated], altitude 8,000 ft., June 18, 1942 (C. P. Alexander)”. Three male “paratopotypes [paratypes] June 17-18, 1942 (C. P. & M. M. Alexander)”. ________________________________________________________________________________ 20 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Al-Shehbaz, Ihsan Ali 1973 The biosystematics of the genus Thelypodium (Cruciferae). Contributions from the Gray Herbarium of Harvard University, (204) (October 1): 3-148.  Thelypodium integrifolium (Nuttall) Endlicher subsp. longicarpum, NEW SUBSPECIES 4 (pp. 111-112, Pl. 7 [p. 36]) Trachyophyta: Magnoliopsida: Brassicales: Brassicaceae [Cruciferae] (mustards) Holotype (G) from “ca. 0.5 mile below Indian Garden [Havasupai Gardens] on Bright Angel Trail, Grand Canyon National Park”, September 13, 1969, Ihsan & Mona Al-Shehbaz 6991; “isotypes to be distributed”. ________________________________________________________________________________ Atwood, N. Duane 2007 Six new species of Phacelia (Hydrophyllaceae) from Arizona and New Mexico. Novon, 17(4): 403-416. Trachyophyta: Magnoliopsida: Boraginales: Hydrophyllaceae (waterleafs) 5 Phacelia furnissii, NEW SPECIES (pp. 408-410, fig. 3 [p. 409]) Holotype and isotypes from “Mohave Co., Arizona Strip, Grand Canyon-Parashant National Monument, Andrus Canyon at rd. crossing, T32N, R10W, S6 & 7, 27 May 2003, N. D. Atwood & L. C. Higgins 29469 (holotype, BRY; isotypes, ARIZ, ASU, BRY [5], K, MO, NY, OSC, RM, TEX, UNLV, UNM, US)”, and paratypes from 22 locales on the Arizona Strip in the vicinities of Andrus Canyon, Trail Canyon, Whitmore Wash, Parashant Canyon, and Hack Canyon. 6 Phacelia higginsii, NEW SPECIES (pp. 410-412, fig. 4 [p. 411]) Holotype and isotypes from “Mohave Co., Arizona Strip, 12.5 mi. SW from head of Trail Canyon toward Andrus Canyon, 36°11.48′N, 113°19.10′W, 3 Sep. 2003, N. D. Atwood, B. Furniss & L. C. Higgsins 29729 (holotype, BRY; isotypes, ARIZ, ASU, BM, BRY [3], Dixie State College, K, MO, NY, RM, TEX, US)”, and paratypes from four locales on the Arizona Strip in the vicinities of Andrus Canyon, and Parashant Canyon. 7 Phacelia hughesii, NEW SPECIES (pp. 412-414, fig. 5 [p. 413]) Holotype and isotypes from “Mohave Co., Arizona Strip, T34N, R11W, S8, E of Salt Spring S base of Poverty Mtn., 20 Sep. 2001, N. D. Atwood & L. E. Hughes 27814 (holotype, BRY; isotypes, ARIZ, ASU, BRY [2], Dixie State College, MO, NY, US )”, and numerous paratypes from various locales on the Arizona Strip. ________________________________________________________________________________ 21 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Atwood, N. Duane, and Welsh, Stanley L. 2013 New plant taxa from Colorado, New Mexico, and Arizona. Western North American Naturalist, 73(1): 113-115. 8 Xylorhiza tortifolia (Torrey & Gray) Greene var. parashantensis, NEW VARIETY (p. 115); not illustrated Trachyophyta: Magnoliopsida: Asterales: Asteraceae (sunflowers) Holotype from “Grand Canyon–Parashant National Monument, Cunningham Canyon, T33N, R14W, S30/SE, Yucca, Acacia, Fouquieria, Ferrocactus community, at 900 m elevation, on limestone ledges and talus, N.D. Atwood & M. Madsen 26586, 29 March 2001. Holotype BRY!” Paratype from “Grand Canyon–Parashant National Monument, 1.5 mi due west of Cane Spring, Sandstone Canyon, with Lycium, Yucca, Agave, Rhus, and Fraxinus, at 1000 m elevation. N.D. Atwood & L.C. Higgins 26927, 12 April 2001, BRY!” ________________________________________________________________________________ Augustson, G. F. 1943 A new subspecies of Orchopeas sexdentatus (Baker) (Siphonaptera: Dolichopsyllidae). Bulletin of the Southern California Academy of Sciences, 42(1): 49-51, Pl. 5 (p. 51).  Orchopeas sexdentatus neotomae, NEW SUBSPECIES 9 Arthropoda: Insecta: Siphonaptera: Ceratophyllidae (rodent fleas) “Holotype: a ♀ from Neotoma lepida devia Goldman, collected by R. L. Rutherford, South Entrance Grand Canyon National Park, Coconino Co., Arizona, June 5, 1942. Deposited in the Allan Hancock Foundation, University of Southern California. Los Angeles, California. Allotype: a ♂ collected and deposited with the holotype as above.” ________________________________________________________________________________ Ball, Carleton R. 1921 10 Undescribed willows of the section Cordatae. Botanical Gazette, 71(6) (June): 426-437.  Salix lutea ligulifolia, NEW VARIETY (pp. 428-430); not illustrated Trachyophyta: Magnoliopsida: Malpighiales: Salicaceae (willows) “Referred specimens” [= isotypes] include “Grand Canyon, Indian Gardens [Havasupai Gardens], alt. 3800 ft., E. A. Goldman 2237, August 25, 1913 (N [U.S. National Herbarium], twig densely pubescent).” ________________________________________________________________________________ 22 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Ball, E. D. 1933 The genus Myndus Stal in North America (Homoptera Fulgoridae). Journal of the Washington Academy of Sciences (Washington, D.C.), 23(10) (October 15): 478-484. [Myndus Stål.] 11 Myndus yuccandus, NEW SPECIES (p. 482); not illustrated Arthropoda: Insecta: Hemiptera: Cixiidae (planthoppers) “Holotype ♂ and one paratype male taken from Yucca at the Grand Canyon Bridge, Ariz., Aug. 30, 1930 by the writer.” [Note: Kramer (1979) examined the type material, with the following observations regarding the locality, without further specifics: “Holotype male, Grand Canyon, Arizona, 4 August 1930, E.D. Ball. The information published with the original description is slightly different from that on the labels of the holotype; this is probably due to a lapsus by Ball. The previously published locality was given as ‘Grand Canyon Bridge’ and the date of collection ‘30 August 1930’. One paraty pe male with same data. Both are in the collection of the USNM.” (no further specifics by Kramer). Ball’s particular note of “Grand Canyon Bridge” is surely Navajo Bridge, by which name it was also known. The dates, however, both may be correct, and may reflect a bridge crossing on both dates as a part of his travels in the field, although it does not resolve the immediate problem with the dates ascribed to the collection of M. yuccandus. Compare data with Hysteropterum cornutum var. utahnum Ball, 1935, and with Yucanda miniata Ball, 1937, below.] [Reference: Kramer, James P., 1979, Taxonomic study of the planthopper genus Myndus in the Americas (Homoptera: Fulgoroidea: Cixiidae). American Entomological Society, Transactions, 105 (September): 301389. See Myndus yuccandus Ball (p. 328, figs. 54-56 [p. 327]; figure legend, p. 326, indicates illustrations from paratype specimen).] _______________________________________________________________ 1935 Some new Issidae, with notes on others—(Homoptera–Fulgoridae). Bulletin of the Brooklyn Entomological Society, 30(2) (April): 37-41. 12 Hysteropterum cornutum var. utahnum, NEW VARIETY (p. 38); not illustrated Arthropoda: Insecta: Hemiptera: Issidae (planthoppers) “Holotype ♀, allotype ♂, and 10 paratypes, taken by the writer near the Grand Canyon, August 1, 1930.” [Note: Regarding the locality and date of collection, compare the comments with Myndus yuccandus Ball, 1933, above, and with Yucanda miniata Ball, 1937, below, from which we glean that he was present at Navajo Bridge twice in August 1930. However, it is not possible to discern from these data whether “near the Grand Canyon” pertains to the North Rim or South R im, or in fact just where in the area the specimens were taken.] _______________________________________________________________ 23 BIBLIOGRAPHY AND ANNOTATED CHECKLIST 1937 Some new Fulgoridae from the western United States. Bulletin of the Brooklyn Entomological Society, Bulletin, 32(5) (December): 171-183. 13 Yucanda miniata, NEW SPECIES (p. 175); not illustrated Arthropoda: Insecta: Hemiptera: Dictyopharidae (planthoppers) “Holotype ♀, allotype ♂, and four paratypes, September 17, 1932, eight, August 4, 1930, and two, August 31, 1935, all taken by the writer from a small shrub that looks like a dwarf mesquite, at the Grand Canyon Bridge, Arizona.” [Note: Regarding the locality and date of collection, compare the comments with Myndus yuccandus Ball, 1933, and with Hysteropterum cornutum var. utahnum (above), from which we glean that he was present at Navajo Bridge twice in August 1930.] ________________________________________________________________________________ Ball, E. D., and Beamer, R. H. 1939 A revision of the genus Athysanella and some related genera (Homoptera-Cicadellidae). Kansas University Science Bulletin, 26(1) (October 1): 5-82 (including Pls. 1-12). Arthropoda: Insecta: Hemiptera: Cicadellidae (leafhoppers) 14  Athysanella fredonia, NEW SPECIES (pp. 12-13, Pl. II, fig. 3 [p. 71), Pl. IX, fig. 3 [p. 79]) “Holotype male, allotype female, and 3 males and 2 females, Fredonia, Ariz., August 6, 1930, E. D. Ball; paratypes as follows: . . . 7 males and 8 females, Grand Canyon, Ariz., E. D. Ball . . . .” [It is not clear whether “Grand Canyon, Ariz.” refers to the canyon or to Grand Canyon village.] 15  Athysanella globosa, NEW SPECIES (pp. 18-19, Pl. II, fig. 13 [p. 72], Pl. X, fig. 13 [p. 80]) “Holotype male, allotype female, 1 male and 3 female paratypes, Grand Canyon, Ariz., E. D. Ball, and in his collection.” [It is not clear whether “Grand Canyon, Ariz.” refers to the canyon or to Grand Canyon village.] 16  Athysanella (Gladionura) casa, NEW SPECIES (p. 48, Pl. VI, fig. 52 [p. 76], Pl. XI, fig. 52 [p. 81]) (Holotype, allotype, and paratypes are from Flagstaff, Ariz.) “. . . other paratypes as follows: 2 males, 3 females, Grand Canyon, Ariz., August, 1930, E. D. Ball . . . numerous specimens [nontype], Grand Canyon, Ariz., August 11, 1927, R. H. Beamer . . . .” [It is not clear whether “Grand Canyon, Ariz.” refers to the canyon or to Grand Canyon village.] ________________________________________________________________________________ 24 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Baral, Hans-Otto; Weber, Evi; and Marson, Guy 2020 Monograph of Orbiliomycetes (Ascomycota) based on vital taxonomy. Luxembourg: National Museum of Natural History Luxembourg, Parts I and II, pp. 1-952, 953-1752. Ascomycota (sac fungi): Orbiliomycetes: Orbiliales: Orbiliaceae This is a profusely documented and illustrated publication, chiefly comprising the descriptions of new species (date of publication 30 October 2020). This citation lists those taxa of interest to this checklist that are typified. Only page numbers are given; pertinent enumerated plates of illustrations are embedded within those page ranges so are not cited separately. Locality data are given here only in condensed format, documenting just the geographic locale, although each published description adds information such as altitude, host material, collector, date of collection, and repository and catalog number of the material. In Baral et al.’s publication, “included material” is indicated to be distinct from the primary types, thus in terms of botanical type nomenclature they are “ PARATYPES” (the term differs from the homonym used in zoological nomenclature). Take note as well that the authors also list material that is specifically “ not included ”; these references are omitted here. (The “not included” specimens “deviate from the established taxa only in some minor points . . . [though] mentioned under that taxon which appears to be most closely related, but they are not included in a description, nor are they proposed as a separate new species.” [p. 17].) All localities are specifically noted by Baral et al. as being in Arizona, although some locality references to “Grand Canyon” are, confusingly, neither within the physiographic canyon nor within the national park but are on adjacent national forest and other lands, sometimes at quite some distance from the canyon or the national park. In the tables of localities in the present volume, some interpretive statements have been included in order to clarify this; specifically, measurements have been made from a map that reveal that linear distances from geographic locales are mostly along the principal highways of the area, which information is added notationally to the original locality data. Also, a few localities northwest of Page, Arizona, when plotted on the map actually show that they are some distance into Utah, along U.S. Route 89; these records are thus omitted here as being extralimital to this checklist. The authorities (authors) of each species are as given in the publication; each thus is in turn “in” Baral, Weber & Marson, as so noted in the taxonomic and locality lists herein. Species are listed here in the same order as in which they appear in the publication. 17  Orbilia purshiae Baral & G. Marson, NEW SPECIES (pp. 466-468) Holotype from “Grand Canyon, Kaibab Plateau, 28 km ESE of Fredonia, 13 km NNW of Jacob Lake”; included material [paratype(s)] from “93 km SSE of Fredonia, S of North Rim”, “Coconino Plateau, 7.5 km ESE of Grand Canyon Village, close to South Rim”. 18  Orbilia ocellata Baral, G. Marson & E. Weber, NEW SPECIES (pp. 476-481) Included material [paratype(s)] from “Canyonlands, 45 km SW of Page, 26 km SW of Marble Canyon” and “Grand Canyon, Coconino Plateau, 3 km N of Tusayan”. 25 BIBLIOGRAPHY AND ANNOTATED CHECKLIST 19 Orbilia magnifica Baral & G. Marson, NEW SPECIES (pp. 509-511) Holotype and isotypes from “Grand Canyon, Kaibab Plateau, 37 km SE of Fredonia, S of Jacob Lake”; included material [paratype(s)] from “3 km N of Kaibab Lodge”. 20  Orbilia phanosoma Baral & G. Marson, NEW SPECIES (pp. 514-515) Included material [paratype(s)] from “Grand Canyon, Coconino Plateau, 3 km N of Tusayan” and “3 km S of Tusayan”. 21  Orbilia multiphanosoma Baral & G. Marson, NEW SPECIES (pp. 515-517) Holotype from “Grand Canyon, Coconino Plateau, 3 km N of Tusayan”. 22  Orbilia arizonensis Baral & G. Marson, NEW SPECIES (pp. 553-556) Included material [paratype(s)] from “Grand Canyon, Kaibab Plateau, 28 km ESE of Fredonia, 13 km NNW of Jacob Lake”, “93 km SSE of Fredonia, S of North Rim”, and “Coconino Plateau, 7.5 km ESE of Grand Canyon Village”. 23  Orbilia navajoana Baral & G. Marson, NEW SPECIES (pp. 571-573) Included material [paratype(s)] from “Grand Canyon, Coconino Plateau, 3.5 km N of Tusayan” and “3 km N of Tusayan”. 24  Orbilia spermoides Baral & G. Marson, NEW SPECIES (pp. 734-737) Holotype from “Coconino Plateau, 3 km N of Tusayan”; included material [paratype(s)] from “Grand Canyon, Kaibab Plateau, 28 km ESE of Fredonia, 13 km NNW of Jacob Lake”, “Coconino Plateau, 8.5 km NNE of Tusayan, Mather Point” and “3 km N of Tusayan”. 25  Orbilia ophiosoma Baral & G. Marson, NEW SPECIES (pp. 1050-1052) Included material [paratype(s)] from “Grand Canyon, Coconino Plateau, 3.5 km N of Tusayan” and “3 km S of Tusayan”. 26  Orbilia maeandrina Baral & G. Marson, NEW SPECIES (pp. 1075-1078) Included material [paratype(s)] from “Grand Canyon, Kaibab Plateau, 28 km ESE of Fredonia, 13 km NNW of Jacob Lake”, “93 km SSE of Fredonia, S of North Rim”, “3 km N of Kaibab Lodge”, “Coconino Plateau, 15 km ESE of Grand Canyon Village, Grandview Point”, and “8.5 km ESE of Grand Canyon Village, close to South Rim”. 27 Orbilia multimaeandrina Baral & G. Marson, NEW SPECIES (pp. 1078-1082) Included material [paratype(s)] from “Grand Canyon, Kaibab Plateau, 28 km ESE of Fredonia, 13 km NNW of Jacob Lake”. 28  Orbilia flexisoma Baral & G. Marson, NEW SPECIES (pp. 1095-1098) Holotype from “Coconino Plateau, 3 km S of Tusayan); included material [paratype(s)] from “Grand Canyon, Kaibab Plateau, 93 km SSE of Fredonia, S of North Rim”, “Coconino Plateau, 3.5 km N of Tusayan”, and “3 km N of Tusayan”. 26 BIBLIOGRAPHY AND ANNOTATED CHECKLIST  Orbilia delphinus Baral & G. Marson, NEW SPECIES (pp. 1105-1110) 29 Included material [paratype(s)] from “Grand Canyon, Kaibab Plateau, 93 km SSE of Fredonia, S of North Rim”, “Coconino Plateau, 15 km ESE of Grand Canyon Village, Grandview Point”, and “3 km N of Tusayan”.  Orbilia macrodelphinus Baral & G. Marson, NEW SPECIES (pp. 1113-1116) 30 Included material [paratype(s)] from “Grand Canyon, Kaibab Plateau, 28 km ESE of Fredonia, 13 km NNW of Jacob Lake” and “Coconino Plateau, 3.5 km N of Tusayan”. 31 Orbilia multitrapezoidea Baral & G. Marson, NEW SPECIES (pp. 1117-1120) Holotype from “Grand Canyon, Coconino Plateau, 3 km S of Tusayan”.  Orbilia calyptrata Baral & G. Marson, NEW SPECIES (pp. 1187-1189) 32 Included material [paratype(s)] from “Grand Canyon, Coconino Plateau, 3.5 km N of Tusayan” and “3 km S of Tusayan”.  Orbilia subovoidea Baral, Matočec & E. Weber, NEW SPECIES (pp. 1257-1260) 33 Included material [paratype(s)] from “Grand Canyon, Kaibab Plateau, 93 km SSE of Fredonia, S of North Rim”. ________________________________________________________________________________ Barber, H. G. [Harry Gardner] 1938 A new genus and species of the subfamily Triatominae (Reduviidae: Hempitera). Proceedings of the Entomological Society of Washington (Washington, D.C.), 40(4) (April): 104-105. Paratriatoma, 34 NEW GENUS  Paratriatoma hirsuta, NEW SPECIES; not illustrated Arthropoda: Insecta: Hemiptera: Reduviidae (assassin bugs) (The “type male” [holotype] is from Mojave, Calif., USNM 52747; disposition of other material not indicated.) “Paratypes: Two males, Phanton Ranch [Phantom Ranch is consistently misspelled in this paper], Grand Canyon, Ariz., May, 1929 (Vernon Bailey) . . . one nymph, Phanton Ranch, Grand Canyon, Ariz., May 1929 (Vernon Bailey) . . . .” ________________________________________________________________________________ 27 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Barneby, Rupert C. 1944 Alice Eastwood Semi-Centennial Publications. No. 3. Pugillus astragalorum alter. Proceedings of the California Academy of Sciences, Series 4, 25(3) (June 1): 147-170, Pl. 17.  Astragalus bryantii, NEW SPECIES (pp. 156-157, Pl. 17, figs. 10-18 [pp. 168-169]) 35 Trachyophyta: Magnoliopsida: Fabales: Fabaceae (legumes) Type [holotype] from “the head of Phantom Canyon in the Grand Canyon of the Colorado River, Coconino Co., 15 Dec. 1939. Collected by Dr. H. C. Bryant of the National Park Service in whose honor it is named. Type in Herb. Calif. Acad. Sci. No. 293940. Also collected [= paratype] in sand at the mouth of Hermit Creek in the Grand Canyon, 10 April 1917, Eastwood 5991 (G, F).” _______________________________________________________________ 1945 Pugillus astragalorum IV: The Section Diplocystium. Leaflets of Western Botany, 4(5) (March): 65-147 [including Pls. 1-4]. 36  Astragalus lentiginosus Dougl. var. oropedii, NEW VARIETY (pp. 135-137); not illustrated Trachyophyta: Magnoliopsida: Fabales: Fabaceae (legumes) “Type [holotype]. Kaibab Trail to Roaring Springs [North Kaibab Trail], Grand Canyon National Park, Coconino County, Arizona, 22 September 1938, fruct. Eastwood & Howell No. 7064 (Herb. Calif. Acad. Sci. No. 262056). Also ibid., 23 June 1933, flor., Eastwood & Howell No. 1054 (Herb. Calif. Acad. Sci. No. 211208, cotype).” [Barneby’s use of “cotype” is an older term often mistakenly used to indicate an isotype or paratype; in this case, the “ibid.” specimen is an isotype.] [Take note also of Jason Andrew Alexander (2009, The types of Astragalus Section Diphysi (Fabaceae), a complex endemic to western North America, Part I: Lectotypifications, epitypifications, and new combinations of several taxa. Botanical Research Institute of Texas, Journal, 3(1) (July): 211-218). Alexander indicates the following information regarding the type of A. l. var. oropedii from North Kaibab Trail: effective lectotypification later by Barneby (1989), epitype designated by Alexander from among Barneby’s syntypes. However, note that Barneby, 1945, did indeed designate the “type” [holotype] and that Alexander may have mistakenly interpreted Barneby’s “cotype” (an older, often misued term) to mean a syntype; thus the “effective lectotypification” by Barneby, as noticed by Alexander, likewise was a misinterpretation; see above).] [Barneby, 1989, is cited by Alexander as in Cronquist, A., Holmgren, A. H., Holmgren, N. H., Reveal, J. L., and Holmgren, P. K. (eds.), Intermountain Flora. Volume III, Part B (New York Botanical Garden, Bronx, New York).] _______________________________________________________________ 28 BIBLIOGRAPHY AND ANNOTATED CHECKLIST 1948 Pugillus astragalorum IX: Novelties in Batidophaca Rydb. Leaflets of Western Botany, 5(5) (February): 82-89.  Astragalus cremnophylax, NEW SPECIES (pp. 83-85); not illustrated 37 Trachyophyta: Magnoliopsida: Fabales: Fabaceae (legumes) [Sentry milkvetch; endemic and endangered.] Type locality “about 2 miles west of El Tovar, Coconino Co., alt. 7050 ft., 3 June 1947, fr., Ripley & Barneby No. 8473. Type [holotype] in Herb. Calif. Acad. Sci., No. 336060. Grand Canyon (‘at the end of the railroad on sandy ledges’), Jones in 1903, fl. (CAS, fragments, PO).” (p. 84). “In its type-locality, A. cremnophylax is confined to a narrow trip of limestone pavement immediately overlooking the Grand Canyon . . . Whence the name from the Greek, meaning watchman or overseer of the chasm. ” _______________________________________________________________ 1949 Pugillus astragalorum X: New species, varieties and combinations. American Midland Naturalist, 41(2) (March): 496-502. 38 Astragalus wootoni var. endopterus, NEW VARIETY (p. 498); not illustrated Trachyophyta: Magnoliopsida: Fabales: Fabaceae (legumes) From “sandy bed of the Little Colorado River, at Cameron, Coconino Co., alt. 3700 ft., 5 June 1947, fl. & fr., Ripley & Barneby No. 8491. Type in herb. Calif. Acad. Sci. No. 337308.” _______________________________________________________________ 1979 Dragma hippomanicum IV. New taxa of Astragalus sect. Humillimi. Brittonia, 31(4): 459-463. 39 Astragalus cremnophylax Barneby var. myriorrhaphis, NEW VARIETY (p. 463, fig. 2 [p. 462]) Trachyophyta: Magnoliopsida: Fabales: Fabaceae (legumes) “Type [holotype]: United States. Arizona. Coconino Co.: locally common in cracks and crevices of limestone cliff and rimrock, 1890 m (6200 ft), Buckskin Mt. [Kaibab Plateau] S of Honeymoon Trail, 28 km (17.5 mi) airline distance E of Fredonia (T41N, R2E, S29/30) [an area of Forest Service roads], 23 May 1979, N. H. Holmgren, P. K. Holmgren & R. C. Barneby 9145 (HOLOTYPE: NY; ISOTYPES: BRY, UTC ad others to be distributed).” ________________________________________________________________________________ 29 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Barnes, William, and Benjamin, F. H. Notes on diurnal Lepidoptera, with additions and corrections to the recent “List of Diurnal 1926 Lepidoptera”. Bulletin of the Southern California Academy of Sciences, 25(3) (September/December): 88-98.  Coenonympha fureae, NEW SPECIES (p. 90); not illustrated 40 Arthropoda: Insecta: Lepidoptera: Nymphalidae (admirals) “Type locality: Grand Canyon, Ariz.” “Holotype ♂, Allotype ♀, 3 ♀ Paratypes; no dates except on one female, 1-7 June.” [It is not clear whether “Grand Canyon, Ariz.” refers to the canyon or to Grand Canyon village.]  Cercyonis damei , NEW SPECIES (p. 90); not illustrated 41 Arthropoda: Insecta: Lepidoptera: Nymphalidae (admirals) “Type locality: Grand Canyon, Ariz.” “Holotype ♂, 1-7 June; 4 ♂Paratypes, one only dated 8- 15 June.” [It is not clear whether “Grand Canyon, Ariz.” refers to the canyon or to Grand Canyon village.] _______________________________________________________________________________ Baroni, Timothy J.; Beug, Michael W.; Cantrell, Sharon A.; Clements, Teresa A.; Iturriaga, Teresa; Læssøe, Thomas; Holgado Rojas, María E.; Aguilar, Frank M.; Quispe, Miguel O.; Lodge, D. Jean; and O’Donnell, Kerry 2018 Four new species of Morchella from the Americas. Mycologia, 110(6) (December): 12051221. 42 Morchella kaibabensis Beug, T.A. Clem. & T.J. Baroni, NEW SPECIES (pp. 1208-1213, figs. 3A- D Ascomycota: Pezizomycetes: Pezizales: Morchellaceae “Typification: USA. Arizona: Coconino County, along Forest Road 293, Kaibab Plateau, North Rim, Grand Canyon National Park, 36.40806, –112.26692, 2700 m, with Picea engelmannii and Picea pungens, 21 May 2017, T.A. Clements & D.M. Fulton TAC-1708 (KOD1793) (holotype ARIZ AN043595). Ex-type culture: NRRL 66753. GenBank: ITS = MH014728; TEF1 = MH014722; RPB1 = MH014733; RPB2_a = MH014738; RPB2-b = MH014744.” [Locality is ca. 7.3 miles west of Kaibab Lodge, on the Kaibab National Forest.] ________________________________________________________________________________ 30 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Barr, M. E. 1989 The Venturiaceae in North America: Revisions and additions. Sydowia, 41: 25-40.  Protoventuria arizonica, NEW SPECIES (pp. 35, 37, fig. 17 [p. 36]) 43 Ascomycota: Dothidiomycetes: Pleosporales: Venturiaceae “Holotypus in foliis emortuis Cercocarpi ledifolii NUTT. ‘North Rim of Grand Canyon, Arizona, 18 Aug. 1973’ a M. E. BARR n. 6096a lectis in Herb. MASS depositus.” [translation: Holotype from decaying leaves of Cercocarpi ledifolii Nutt., from “North Rim of Grand Canyon, Arizona, 18 Aug. 1973”, by M. E. Barr n. 6096a deposited in Herb. MASS.] [It is not certain whether “North Rim of Grand Canyon” refers to the North Rim of the canyon or to North Rim, Arizona.] ________________________________________________________________________________ Barr, William F. 1972 New species of North American Acmaeodera (Coleoptera: Buprestidae). Arquivos do Museu Boçage (Lisboa), Series 2, 3(7): 145-201.  Acmaeodera pletura, NEW SPECIES (pp. 164-166, fig. 12 [p. 199]) 44 Arthropoda: Insecta: Coleoptera: Buprestidae (jewel beetles) (Holotype from Lee Canyon, Clark Co., Nevada; California Academy of Sciences.) Paratypes include “. . . north rim, Grand Canyon National Park, Arizona, June 15, 1930 (C. Kelley) 1 specimen; south rim of Grand Canyon, Arizona, July 8, 1967, on blossom of Cowania stansburiana (W. F. Barr), 1 specimen. [All] Paratypes in the collections of the California Academy of Sciences, R. L. Westcott and W. F. Barr.” ________________________________________________________________________________ Bauer, David L. 1955 A new race of Papilio indra from the Grand Canyon region. The Lepidopterists’ News, 9(2/3): 49-54, plate (p. 52). 45  Papilio indra kaibabensis, NEW SUBSPECIES Arthropoda: Insecta: Lepidoptera: Papilionidae (swallowtails) “Holotype male: (expanse 76 mm.) Bright Angel Point, Grand Canyon, Coconino Co., Ariz., August 1953, leg. Ernst Christensen. Allotype female: (expanse 86 mm.) same locality and collector as holotype, 14 August 1953. Paratypes: 9 males and 3 females. Data as follows (all but one male from the Grand Canyon National Park): 1 male, North Rim, 3 August 1938, leg. Louis Schellback [sic, Schellbach]; 1 male, Yavapai Point, South Rim, 22 August 1944, leg. Louis Schellback; 1 male, Bright Angel Point, North Rim, 13 August 1951, leg. Ernst Christensen; 1 male, Bright Angel Point, North Rim, 7 August 1953, leg. Ernst Christensen; 1 male, Roaring Springs, North Rim, 4 August 1953, leg. Ernst Christensen; 1 male, near Ryan Ranger Station, Kaibab Plateau, Coconino Co., Ariz., 1 July 1952, leg. D. L. Bauer; 1 female, data same as for allotype; 1 female, data same as for holotype.” 31 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Allotype and holotype, respectively, are illustrated in the plate, the third pair of figures (upperside of specimens) and the fourth pair of figures (underside of specimens). “The holotype and allotype and four male paratypes will be returned to the Naturalist Work Shop Collection, Grand Canyon National Park; from there I believe the holotype and allot ype will be sent to the U. S. National Museum. One male and one female paratype will be returned to Dr. Rodeck and the University of Colorado Collection, one male paratype to the Yale Peabody Museum, and one male paratype to the Los Angeles County Museum. The remainder are in the author’s collection.” ________________________________________________________________________________ Baxter, E. M. 1933 Opuntia aurea—a new species. Cactus and Succulent Journal (Cactus and Succulent Society of America), 5: 489-490. 46 Opuntia aurea, NEW SPECIES ; illustrated Trachyophyta: Magnoliopsida: Caryophyllales: Cactaceae (cacti) “Type Locality: ½ mile north of Pipe Springs [Pipe Spring] on the Kaibab Indian Reservation, Arizona. Type Specimen [holotype]: Two joints collected in 1930 by Percy and Helen McCabe at the type lcoality and placed in the Dudley Herbarium at Stanford University as No. 213750, October, 1933.” [The illustration is a photo of a living plant, credited to “McCabe”; it is not indicated whether the photo depicts the source plant of the holotype. In addition to the type from Percy and Helen McCabe, Baxter also credits without further detail (p. 490) “Howard Gates for a specimen and much information on distribution.”] Discussion begins: “A new species, introduced first by the McCabe Gardens of San Diego, and later by Howard Gates of Anaheim, is here described for the first time. It has appeared in the McCabe catalog both under the name shown and as Opuntia basilaris aurea, and has been sold by Mr. Gates as ‘yellow-flowered basilaris’. The McCabe name is taken as the specific name for the species, denoting the gold color of the flower.” ________________________________________________________________________________ Beamer, R. H. 1948 Some new species of Delphacodes (Homoptera, Fulgoridae, Delphacinae); Part IV. Journal of the Kansas Entomological Society, 21(3) (July): 96-110. 47  Delphacodes apicata, NEW SPECIES (pp. 100-101, Pl. VIII, figs. 37, 37a, 37b [p. 107; legend p. 106]) Arthropoda: Insecta: Hemiptera: Delphacidae (planthoppers) (Under the heading, “Brachypterous Form” (pp. 100-101), Beamer designates a holotype, allotype, and two paratypes from “San Francisco Mt., Ariz.”; deposited in “Snow Entomological Collections and United States National Museum”.) Under the heading, “Macropterous Form” (p. 101), Beamer lists: “Holomorphotype , Grand Canyon, Ariz., Aug. 11, 1927, R. H. Beamer” with an “allomorphotype” and “paramorphotypes” from San Francisco Mt. and Flagstaff, Ariz. 32 It is BIBLIOGRAPHY AND ANNOTATED CHECKLIST unclear whether these may also be considered paratype material, based on provisions of earlier editions of the International Code of Zoological Nomenclature. The figures do not indicate which specimens are illustrated. [It is not clear whether “Grand Canyon, Ariz.” refers to the canyon or to Grand Canyon village.] ________________________________________________________________________________ Belk, Denton, and Fugate, Michael 2000 Two new Branchinecta (Crustacea: Anostraca) from the southwestern United States. Southwestern Naturalist, 45(2) (June): 111-117. Arthropoda: Branchiopoda: Anostraca: Branchinectidae (brine shrimp) 48 Branchinecta kaibabensis, NEW SPECIES (pp. 115-117, figs. 2b [p. 114], 3a-e [p. 116] “Holotype male (USNM 260875) and paratypes of both sexes (USNM 260876) from DB 232 are at the National Museum of Natural History, Smithsonian Institution, Washington, D.C.” “Type Locality—A natural pool 0.6 km east of Arizona Highway 67 just northeast of Forest Road 213 in the Kaibab National Forest . . . (36°30′38″N, 112°07′55″W). Forest Road 213 intersects Arizona Highway 67, 30.6 km south of the city of Jacob Lake.” “DB 232 pool in depression 0.6 km east of AZ 67 on Forest Road 213, Kaibab National Forest, Coconino Co., Arizona, collector D. Belk, 22 May 1974.” ________________________________________________________________________________ Benson, Bernard W. 1957 A new cactus from Arizona. Cactus and Succulent Journal (Cactus and Succulent Society of America), 29(5) (September/October): 136-137. 49 Pediocactus paradinei, NEW SPECIES ; illustrated Trachyophyta: Magnoliopsida: Caryophyllales: Cactaceae (cacti) Named as “Pediocactus (?) paradinei B. Benson, sp. nov.” From House Rock Valley (without further specific locale). Holotype, “Boyce Thompson Southwestern Arboretum Herbarium, No. Bwb 8-1956-1”. Isotype, “Herbarium of Pomona College, No. 286,120”. “Mr. N. A. Paradine collected the first plants in House Rock Valley, Coconino County, Arizona, in the spring of 1956. These were found under the protection of small sagebrush plants. Later others were found in clumps of perennial bunch grasses.” Benson adds: “In August of 1956, a description of this plant was sent to the editor of the Cactus and Succulent Journal for publication. In October of 1956, Mr. W. T. Marshall of the Desert Botanical Garden, Tempe, Arizona, described the same plant in the Saguaroland Bulletin 10: 8991. The author appreciates Mr. Marshall’s courtesy in stepping aside to permit the naming of Pediocactus paradinei.” The item thus cited is “A New Genus of Cactuses for Arizona”, unsigned but in Saguaroland Bulletin (W. Taylor Marshall, editor), 10(8) (October 1956): 89-91, wherein it 33 BIBLIOGRAPHY AND ANNOTATED CHECKLIST appears only as Pediocactus sp., based on gatherings in House Rock Valley by Gus and Sylvia Hermann. ________________________________________________________________________________ Bequaert, J. [Joseph C.] 1932 The Nearctic social wasps of the subfamily Polybiinae (Hymenoptera; Vespidae). Entomologica Americana, 13(3) (December): 87-149 (pagination includes Pls. 27-29).  Mischocyttarus flavitarsis var. kaibabensis, NEW VARIETY (p. 133); not illustrated [the 50 plates and text-figs. do not pertain to this new variety] Arthropoda: Insecta: Hymenoptera: Vespidae (hornets) “Bright Angel Trail, Grand Canyon, Cococino [sic] Co., one female, holotype (Cornell Univ.) and one female paratype (M.C.Z[.]); both collected by R. C. Shannon, on the Cornell Biological Expedition, August 4, 1917.” (The etymology of the varietal name is from its “perfect parallel in color (or ‘mimic’) of Polistes canadensis var. kaibabensis Hayward, which occurs in the same district.”) [Refer to Hayward (1932, no. 155 herein).] ________________________________________________________________________________ Beutenmüller, William 1907 Notes on a few North American Cynipidae, with descriptions of new species. Bulletin of the American Museum of Natural History, 23: 463-466, Pl. 37.  Andricus wheeleri, NEW SPECIES (p. 464, Pl. 37, figs. 7-9) 51 Arthropoda: Insecta: Hymenoptera: Cynipidae (gall wasps) “Coconino Forest, rim of the Grand Cañon, Arizona; altitude 7,000 feet. (William M. Wheeler.) Types [syntypes]. Coll. Am. Mus. Nat. Hist.” ________________________________________________________________________________ Bohart, R. M. 1958 A new Priononyx and a key to the North American species (Hymenoptera: Sphecidae). Bulletin of the Brooklyn Entomological Society, 53(4) (October): 90-93. 52  Priononyx subatrata, NEW SPECIES Arthropoda: Insecta: Hymenoptera: Sphecidae (sphecid wasps) (Holotype from Deep Springs, California.) Paratypes include specimens from “Grand Canyon South Rim (M. A. Evans), Kaibab Forest (M. Wasbauer)”. The “Kaibab Forest” specimen may or may not be from Kaibab National Forest units near Grand Canyon. ________________________________________________________________________________ 34 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Bromley, Stanley W. 1940 New U.S.A. robber flies (Diptera: Asilidae). Bulletin of the Brooklyn Entomological Society, 35(1) (February): 13-21.  Erax benedicti, NEW SPECIES (pp. 15-16, fig. 1 [p. 20]) 53 Arthropoda: Insecta: Diptera: Asilidae (robber flies) (Holotype from Winslow, Ariz.; allotype from Santa Rita Mountains, Ariz.; both in University of Kansas Collection.) Paratypes include “2 females, Grand Canyon, Ariz., June 12, 1937 (W. Benedict)”. No further data regarding these paratypes. [It is not clear whether “Grand Canyon, Ariz.” refers to the canyon or to Grand Canyon village.] ________________________________________________________________________________ Bunn, Ralph W. 1930 Notes on the genus Aphelonema Uhler with descriptions of new species (Homoptera, Fulgoridae). Journal of the Kansas Entomological Society, 3(3) (July): 73-77.  Aphelonema convergens var. canyonensia, NEW VARIETY (p. 76); not illustrated 54 Arthropoda: Insecta: Hemiptera: Caliscelidae (planthoppers) “Holotype; female, from vicinity of Grand Canyon, Arizona collected by Dr. [R. H.] Beamer August 11, 1927. Allotype; male, collected at the same time and place by Dr. Beamer. Paratypes; . . . one female from Grand Canyon, Arizona, taken August 11, 1927 by Dr. P. A. Readio; three females from Grand Canyon, Arizona taken August 11, 1927 by L. D. Anderson; and six females from the same locality and collected at the same time by Dr. Beamer. Types deposited in Snow Entomological Collection.” [It is not clear whether “Grand Canyon, Arizona” refers to the canyon or to Grand Canyon village.] ________________________________________________________________________________ Burke, H. E. 1907 A new buprestid enemy of Pinus edulis. (Melanophila pini-edulis, n. sp.) Proceedings of the Entomological Society of Washington (Washington, D.C.), 9(1/4) (March/December): 117-118. 55  Melanophila piniedulis, NEW SPECIES; fig. 6 Arthropoda: Insecta: Coleoptera: Buprestidae (jewel beetles) “Type [syntypes].—♀ and ♂, No. 11350, U. S. National Museum.” One syntype is “One female taken by Mr. E. A. Schwarz from pinyon (Pinus edulis) at Bright Angel Hotel (Grand Canyon P.O.), Arizona, on July 11, 1902.” ________________________________________________________________________________ 35 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Casey, Thomas L. 1895 Coleopterological notices. VI. Annals of the New York Academy of Sciences, 8 (July): 435-838. Arthropoda: Insecta: Coleoptera: Melyridae (soft-winged flower beetles) 56 Trichochrous incipiens, NEW SPECIES (pp. 489-490); not illustrated “Arizona (near the Grand Cañon of the Colorado). Dr. T. Mitchell Prudden. The single type [holotype] is a female . . . .” 57 Trichochrous reversus, NEW SPECIES (pp. 527-528); not illustrated “Arizona (near the Grand Cañon of the Colorado). The single type [holotype] is a female . . . ; it was collected and kindly given to me by Dr. T. Mitchell Prudden, of New York.” _______________________________________________________________ 1900 Review of the American Corylophidae, Cryptophagidae, Tritomidae and Dermestidae, with other studies. Journal of the New York Entomological Society, 8(2) (June): 51-172. Arthropoda: Insecta: Coleoptera: Dermestidae (dermestid beetles)  Cryptorhopalum pruddeni, NEW SPECIES (p. 156); not illustrated 58 “Arizona (Cañon of the Colorado River)—Dr. T. Mitchell Prudden.”  Orphilus aequalis, NEW SPECIES (p. 164); not illustrated 59 “Arizona (Cañon of the Colorado River)—Dr. Prudden.” _______________________________________________________________ 1907 Notes on Chalcolepidius and the Zopherini. The Canadian Entomologist (London), 39(2) (February): 29-46. Arthropoda: Insecta: Coleoptera: Tenebrionidae (darkling beetles) Zopherodes, NEW GENUS (p. 38)  Zopherodes lugubris, NEW SPECIES (p. 41); not illustrated 60 “Arizona (Grand Canyon of the Colorado).”  Zopherodes pruddeni, NEW SPECIES (p. 41); not illustrated 61 “Arizona (Grand Canyon of the Colorado), T. Mitchell Prudden.” _______________________________________________________________ 1908 A revision of the tenebrionid subfamily Coniontinæ. Proceedings of the Washington Academy of Sciences (Washington, D.C.), 10 (April 25): 51-166. Arthropoda: Insecta: Coleoptera: Tenebrionidae (darkling beetles) 62  Discodemus brevipennis, NEW SPECIES (p. 61); not illustrated “Arizona (Grand Canyon of the Colorado),—T. Mitchell Prudden.” _______________________________________________________________ 36 BIBLIOGRAPHY AND ANNOTATED CHECKLIST 1912 Studies in the Longicornia of North America. In: Casey, Thomas L., Memoirs on the Coleoptera. III. Lancaster, Pennsylvania: New Era Printing Co., pp. 215-386. Arthropoda: Insecta: Coleoptera: Cerambycidae (long-horned beetles)  Prionus spiculosus, NEW SPECIES (p. 240); not illustrated 63 “Arizona (Grand Cañon of the Colorado),—T. Mitchell Prudden.”  Prionus angustulus, NEW SPECIES (p. 241); not illustrated 64 “Arizona (Cañon of the Colorado River),—Prudden.”  Prionus terminalis, NEW SPECIES (pp. 243-244); not illustrated 65 “Arizona (Cañon of the Colorado River),—Prudden.”  Stenosphenus pruddeni, NEW SPECIES (pp. 346-347); not illustrated 66 “Arizona (Cañon of the Colorado),—T. Mitchell Prudden.” _______________________________________________________________ 1918 A review of the North American Bembidiinæ. In: Casey, Thomas L., Memoirs on the Coleoptera. VIII. 1918. Lancaster, Pennsylvania: New Era Printing Co., pp. 1-223. Arthropoda: Insecta: Coleoptera: Carabidae (ground beetles)  Bembidion (Cyclolopha) occultum, NEW SUBGENUS, NEW SPECIES (p. 144); not illustrated 67 “Arizona (Grand Cañon of the Colorado),—T. Mitchell Prudden.” ________________________________________________________________________________ Chamberlin, Ralph V. 1940 New American tarantulas of the family Aviculariidae. Bulletin of the University of Utah, 30(13) (May 6), 39 pp. Arthropoda: Euchelicerata: Areneae: Theraphosidae (tarantulas) 68  Aphonopelma behlei, NEW SPECIES (pp. 26-27); not illustrated “Locality: Arizona: Coconino Co., Grand Canyon Village, 7000 ftt. el., 15 Sept., 1939. Two males [syntypes] taken by Dr. W. H. Behle on a paved highway after a heavy rain.” (Additional syntypes from Aztec, New Mexico.) 69  Aphonopelma phasmus, NEW SPECIES (p. 28); not illustrated “Locality: Arizona: Grand Canyon, Phantom Ranch. 26 July. 1934. One male [holotype] collected by Dr. Lutz. American Museum collection.” _______________________________________________________________ 37 BIBLIOGRAPHY AND ANNOTATED CHECKLIST 1948 On some American spiders of the family Erigonidae. Annals of the Entomological Society of America, 41(4): 483-562. 70 Tapinocyba kesimba, NEW SPECIES (pp. 552-553, Pl. 11, figs. 117-121 [p. 515, legend p. 514]) Arthropoda: Euchelicerata: Areneae: Linyphiidae (dwarf weavers) “Type locality.—Arizona: Kaibab Forest (V. T. Ranch), September 4, 1931, 2 ♂ 6♀ [syntypes], R. V. Chamberlin.” (“Other records” include “Kaibab Forest, September 4, 1931, 2 ♂, 6♀, R. V. Chamberlin.) ________________________________________________________________________________ Chamberlin, Ralph V., and Ivie, Wilton 1942 A hundred new species of American spiders. Bulletin of the University of Utah, 32(13) (June 30) [Biological Series, 7(1)], 117 pp. 71 Neoantistea coconino, NEW SPECIES (pp. 28-29, figs. 59, 60 [p. 95, legend p. 94]) Arthropoda: Euchelicerata: Areneae: Hahniidae (hahniid spiders) “Type locality: W 112°:N 36°, Kaibab Forest, Arizona; ♂ holotype; June 14, 1934; W. Ivie and H. Rasmussen, collectors.” [The overly generalized coordinates lie within Grand Canyon National Park, just west of Grandview Point; in fact, by coincidence on a cliff face(!) just over the rim of the canyon (https://confluence.org/confluence.php?id=91). The specific epithet and indication of “Kaibab Forest” may point either to a locality on the nearby Kaibab National Forest or is only a generalized term for the South Rim forest, sometimes called the Coconino forest, on the Coconino Plateau.] ________________________________________________________________________________ Chandler, Peter 1993 The Holarctic species of the Mycetophila fungorum (De Geer) group (Diptera: Mycetophilidae). British Journal of Entomology and Natural History, 6: 5-11. 72  Mycetophila neofungorum, NEW SPECIES (pp. 7-9, figs. 7-9 [p. 8]) Arthropoda: Insecta: Diptera: Mycetophilidae (fungus gnats) “Holotype male USA, Arizona, Grand Canyon National Park (north rim), 15.vii.1954 (W. L. Downes, Natural History Museum, London).” [Based on a unique museum specimen.] ________________________________________________________________________________ 38 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Christy, Charlotte M. 1997 A new species of Mentzelia Section Bartonia (Loasaceae) from Arizona. Novon, 7(1): 2526. 73 Mentzelia collomiae, NEW SPECIES ; not illustrated Trachyophyta: Magnoliopsida: Cornales: Losaceae (loasa) (“[A]n edaphic endemic species restricted to cinder areas of the San Francisco Volcanic Field”, holotype and isotypes from “ca. 3.5 mi. due N of Sunset Crater National Monument".) Paratypes include various localities, the nearest of which pertinent to this bibliography (and at the limit of its coverage) is “Red Mountain, 35 mi. NW of Flagstaff on Hwy. 180”. ________________________________________________________________________________ Chuang, Tsan Iang, and Heckard, Lawrence R. 1986 Systematics and evolution of Cordylanthus (Scrophulariaceae—Pedicularieae) (including the taxonomy of subgenus Cordylanthus). Systematic Botany Monographs, Volume 10, 105 pp. 74 Cordylanthus wrightii kaibabensis, NEW SUBSPECIES (pp. 87-89, figs. 35i-n [p. 88]) Trachyophyta: Magnoliopsida: Lamiales: Orobanchaceae (broomrapes) “TYPE: U.S.A. Arizona: Coconino Co., openings in Juniperus monosperma, occasional Pinus ponderosa with Artemisia tridentata, 7.5 mi E of Jacobs Lake [Jacob Lake], 6400 feet, Heckard, Bacigalupi & Chuang 3585 (holotype: JEPS!; isotypes: ARIZ! ASU! BRY! COLO! F! GH! ISU! MO! NY! TEX! US! UTC!).” “Representative Specimens [= paratypes]. U.S.A. Arizona: Coconino Co., House Rock Valley, Smith s.n. (NY, PH); Kaibab Plateau near Jacobs Lake, Eastwood & Howell 6441 (DS, POM).” ________________________________________________________________________________ Clark, Curtis 1998 New names and combinations in Encelia frutescens sensu lato (Asteraceae: Heliantheae). Aliso (Rancho Santa Ana Botanic Garden, Claremont, California), 17(2): 201-202. 75  Encelia resinifera subsp. tenuifolia, NEW SUBSPECIES (p. 202); not illustrated Trachyophyta: Magnoliopsida: Asterales: Asteraceae (sunflowers) “TYPE: United States. Arizona. Coconino Co: 4 miles down Supai Trail, Havasupai Canyon [Havasu Canyon]. 3 June 1961. C. and C. Dailey s.n., MNA 2256/ B12,747. holotype: MNA!.” [Note: The precise gathering locality may be within the boundaries of the Havasupai Indian Reservation.] “Subspecies tenuifolia occurs almost entirely within the Grand Canyon of the Colorado River in northern Arizona, ranging from the Grand Wash Cliffs east to Marble Canyon, evidently following the 1200 m contour.” ________________________________________________________________________________ 39 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Clarke, J. F. Gates 1947 Notes on Oecophoridae, with descriptions of new species. Journal of the Washington Academy of Sciences (Washington, D.C.), 37(1): 2-18.  Depressaria schellbachi, NEW SPECIES (pp. 10, 13; figs. 6-6a [p. 7], 13 [p. 12]) 76 Arthropoda: Insecta: Lepidoptera: Oecophoridae (oecophorid moths) “Type.— U.S.N.M. No. 58013. Type locality.—Shoshone Point, Grand Canyon, Ariz., elev. 7,050 feet. Food plant.—Lomatium macdougali Coult. and Rose.” “Described from the ♂ type [holotype], five ♂ and three ♀ paratypes from the Grand Canyon (June dates, 1944 and 1946, Louis Schellbach, 3d). Paratypes in the U.S. National and British Museums and in the Grand Canyon National Park Collection.” ________________________________________________________________________________ Clover, Elzada U. 1942 A new species and variety of Sclerocactus from Arizona. American Journal of Botany, 29(2) (February): 172-173. Trachyophyta: Magnoliopsida: Caryophyllales: Cactaceae (cacti) 77  Sclerocactus havasupaiensis, NEW SPECIES; illustrated [fig. 1 (p. 172) shows “Actual type specimen photographed in natural habitat at Havasupai Canyon, Arizona (Clover 5229 in part: there are several cotypes”] “The type specimen [holotype] (fig. 1) was collected by William Belknap, Jr., April 26, 1941, on top of the Supai Formation in Havasupai Canyon [Havasu Canyon], Arizona, where the species grows abundantly. Specimens [= paratypes] not in flower were previously collected by the author in July, 1940, in Havasupai Canyon (Clover 5229) near Navajo Falls, and in Hualapai Canyon [tributary to Havasu Canyon] (Clover 5100) on talus of the Supai Formation. A few plants were seen on the floor of Cataract Canyon above Supai, September, 1941.” Note: The precise gathering localities may be within the boundaries of the Havasupai Indian Reservation. 78  Sclerocactus havasupaiensis var. roseus, NEW VARIETY; illustrated [figs. 3, 4 (p. 173) show “Actual type specimen [holotype] (Clover 6403) photographed while alive at Botanical Garden, University of Michigan, now dried and preserved as a herbarium specimen. Additional plants of the same collection (cotypes [= istotypes]) are in the living collection (Bot. Gard. no. 18079).”] “Specimen typicum [holotype] legit E. Clover & Wm. Belknap, Jr. (Clover 6403), in Havasupai Canyon [Havasu Canyon], Arizona, siccatum est in Herbario, Universitatis Michiganensis.” [comparable text in English not provided]. “This variety was found in association with S. havasupaiensis on top of the Supai Formation in the same location in May 1941, and are in the living collection (no. 18079) in the Botanical Gardens, University of Michigan.” Note: The precise gathering locality may be within the boundaries of the Havasupai Indian Reservation. ________________________________________________________________________________ 40 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Cockerell, T. D. A. [Theodore Dru Alison] 1905 New American bees. Proceedings of the Biological Society of Washington (Washington, D.C.), 18 (June 29): 177-184.  Triepeolus hopkinsi, NEW SPECIES (p. 184); not illustrated 79 Arthropoda: Insecta: Hymenoptera: Apidae (honey bees) Apparently named on a single male specimen [holotype]. “Grand Canyon of the Colorado, Arizona, August 3, 1904. (Webb). Received from Mr. Viereck, to whom it has been returned. It is named after Professor Hopkins [A. D. Hopkins?], who sent it to Mr. Viereck, and who has done good work in the region of the Grand Canyon.” ________________________________________________________________________________ Cole, A. C., Jr. 1936 Descriptions of seven new western ants. (Hymenop.: Formicidae.) Entomological News (Academy of Natural Sciences of Philadelphia, Entomological Section), 47 (May): 118121. 80 Myrmecocystus melliger subsp. semirufus Emery var. romainei, NEW VARIETY (p. 120). [Not valid as a quadrinomial taxon, and no identification of holotype; non Myrmecocystus romainei Snelling, 1975] Arthropoda: Insecta: Hymenoptera: Formicidae (ants) “Described from a series of 54 workers [= syntypes] taken by Miss Marjorie Romaine, at Cameron, Arizona. The holotype is in the author’s collection, and paratypes are to be deposited in the collections of the U. S. National Museum and of Dr. C. H. Kennedy.” ________________________________________________________________________________ Coquillett, D. W. 1902 New acalyptrate Diptera from North America. Journal of the New York Entomological Society, 10(4) (December): 177-191. 81  Trypeta varipennis, new species (p. 180); not illustrated Arthropoda: Insecta: Diptera: Tephritidae (fruit flies) “Habitat.—Bright Angel Hotel, brink of Grand Canyon, Coconino Co., Arizona. A female specimen collected July 10 by Mr. H. S. Barber. Type [holotype] No. 6635, U.S.N.M.” ________________________________________________________________________________ 41 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Coulter, John M. 1896 Preliminary revision of the North American species of Echinocactus, Cereus, and Opun tia. Contributions from the U.S. National Herbarium, 3(7): 355-462, I-IV [index]. 82 Echinocactus polycephalus xeranthemoides, NEW VARIETY (p. 358). Trachyophyta: Magnoliopsida: Caryophyllales: Cactaceae (cacti) “Type, Siler of 1881 and 1883 [= syntypes] in Herb. Mo. Bot. Gard.” [Herbarium of the Missouri Botanical Garden]. “Specimens examined: Utah (Siler of 1883, ‘Kanab Mts.’ [syntype]): Arizona (Siler of 1881, near the Colorado ‘on the Kanab wash’ [syntype]; Rusby 619, of 1883, at Peach Springs [paratype]; Evans of 1891, between Gila Bend and Yuma) [paratype].” ________________________________________________________________________________ Coulter, John M., and Rose, J. N. 1900 Monograph of the North American Umbelliferae. Contributions from the U.S. National Herbarium, 7(1): 9-256, i-vii. 83 Phellopterus multinervatus, NEW SPECIES (pp. 169-170); not illustrated Trachyophyta: Magnoliopsida: Brassicales: Apiaceae (parsleys) “Type locality, Peach Springs, northern Arizona; collected by Mr. and Mrs. J. G. Lemmon, May, 1884 (distributed as Cymopterus montanus); type specimen [holotype] in U.S. Nat. Herb.” (Paratypes indicated from San Francisco Mountains and Fort Huachuca, Arizona.) [Although there is no further data on the locality, the Lemmons visited Peach Springs Canyon, descending to Diamond Creek and the Colorado River and remained in the vicinity of Peach Springs for a month. See J. G. Lemmon (1888, Grand Cañon of the Colorado. Overland Monthly, New Series, 12(69) (September): 244-256). Thus the precise type locality for this species must be considered to be generalized. Also note that the gathering is likely from what is now the Hualapai Indian Reservation.] ________________________________________________________________________________ Coville, Frederick Vernon 1894 New or little-known plants. Ostrya Knowltoni, a new species of hop hornbeam. Garden and Forest, 7(317) (March 21): 114-116. 84  Ostrya knowltoni, NEW SPECIES, fig. 23. (Subsequent grammatical correction of specific epithet to Ostrya knowltonii.) Trachyophyta: Magnoliopsida: Fagales: Betulaceae (alders) “Type specimen in the United States National Herbarium, collected July 10th, 1892 (No. 272) in Yavapai County [today Coconino Co.], Arizona, within the Grand Cañon of the Colorado River, by J. W. Toumey.” Holotype illustrated in fig. 23 (p. 115). Type locality at top of Old Hance Trail, Hance Canyon (see below regarding its recovery). [The original holotype illustration and the recovered type locality are illustrated on the covers of the present checklist.] 42 BIBLIOGRAPHY AND ANNOTATED CHECKLIST “In the year 1889 Mr. Frank H. Knowlton was occupied in making a collection of the plants of San Francisco Mountain, in northern Arizona, and spent a portion of his time in a side trip to the Grand Cañon at a point known as Cañon Spring, having traversed the road from San Francisco Mountain by the way of Hull Spring and Red Horse Spring. [. . .] In the vicinity of Cañon Spring Mr. Knowlton obtained a specimen of a tree which subsequently proved to be an Ostrya, but which was not at that time in fruit. In 1892 Professor J. W. Toumey, of Tucson, Arizona, botanist of the State University, visited the same locality and obtained fruiting specimens of this tree, which in January, 1894, came into the writer’s hands.” “The following communication from Professor Toumey, under date of February 5th, 1894, will service to direct others to the extact station of the original tree, which is, with little doubt, the same specimen from which Mr. Knowlton obtained his material: “The Ostrya, which you inquire further in regard to, and specimens of wich were sent to the Department of Agriculture a year ago, as found north of Flagstaff, on the rim of the Grand Cañon of the Colorado River. It is growing at the left, a few rods after begining the descent to the river over John Hance’s trail.” Later researchers failed to locate the type locality, and the species was there presumed to have been extirpated. However, in 1997 Earle Spamer thought that the oversight was due to the misunderstanding of “John Hance’s trail,” portions of which were destroyed by landslides in 1895. Hance thereafter rerouted the upper part of his trail to descend into Red Canyon farther to the east, which is the “Hance Trail” as known today. The older trail at the head of Hance Canyon, if referred to at all, is called the “Old Hance Trail,” although it is hardly even a route today. Spamer asked Nancy Brian (Grand Canyon National Park) to investigate whether the type locality could be identified at the head of the Old Hance Trail. As reported by Nancy J. Brian and Earle E. Spamer (2000, Knowlton hop-hornbeam revisited (Ostrya knowltonii Cov.), Bartonia, no. 60: 49-56), p. 54: “On 23 May 1997, Brian located the head of the Old Hance Trail and descended into the canyon. She saw the distinctive dark yellow-green color of a deciduous tree nestled amongst the darker foliage of the surrounding Rocky Mountain Douglas-firs and white firs (Figs. 2A, B [of that paper]). Such a ‘window box’ habitat is frequently found just below the north-facing slope of the canyon wall. It is a shaded, moist, steep environment which allows forest-dwelling species to grow well below their usual elevational range. Scrambling down the steep slope, at ca. 6,800 feet (2,100 m) elevation, Brian located about 25 trees of Ostrya knowltonii. Other plants of the community include scattered shrubs such as snowberry and Utah serviceberry, along with an occasional pinyon pine, ponderosa pine, mutton grass, and phlox. No redbuds were in the immediate vicinity, as Tourney had described, although Gambel oak was found upslope on the canyon rim. Most of the Knowlton hop-hornbeams were shrub-like, with up to ten branches originating from ground level. A few trees were saplings, indicating a measure of reproductive success. Collections were made for the National Herbarium (US), the University of Arizona (ARIZ), the Desert Botanical Garden (DBG) in Phoenix, and the Academy of Natural Sciences of Philadelphia (PH; Fig. 3 [of Brian and Spamer]).” ________________________________________________________________________________ 43 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Cresson, Ezra T., Jr. 1919 Dipterological notes and descriptions. Proceedings of the Academy of Natural Sciences of Philadelphia, 71: 171-194.  Bombylius facialis, NEW SPECIES (pp. 187-188); not illustrated 85 Arthropoda: Insecta: Diptera: Bombyliidae (bee flies) “Type.—♂; Rim of Grand Canyon, Arizona, 7000 feet, alt., [sic] May 23, 1918, (F. M. Jones), [A. N. S. P. No. 6213].” [Square brackets around catalogue number data are part of quotation.] ________________________________________________________________________________ Crosswhite, Frank S. 1967 Revision of Penstemon Section Habroanthus (Scrophulariaceae). III. Series Virgati. American Midland Naturalist, 77(1) (January): 28-41. 86 Penstemon virgatus pseudoptutus, NEW SUBSPECIES (pp. 35-37) [the call-out for fig. 1 refers only to a map of distributions]. Trachyophyta: Magnoliopsida: Lamiales: Plantaginaceae (plantains) “Typus [holotype]: Mead 503 (US) from DeMotte Park, Kaibab Plateau, Coconino County, Arizona.” Other “specimens examined” [paratypes] include: “Bright Angel Point, N rim of Grand Canyon, Pilsbry s.n. (PH). 19 mi N of Bright Angel Point at V.T. Park, Richards s.n. (NY). V.T. Ranger Station, Kaibab Plateau, Collum s.n. (US) . . . Kaibab Plateau, Kraus s.n. (WIS). Kaibab Natl. Forest, Goodman & Hitchcock 1630 (F, NY, PH), Johnson s.n. (WIS). Dry Park, Kaibab N.F., Eggleston 10227 (US). Hwy. from Jacob Lake to Lodge, Kaibab Forest, Collom 944 (US). Roadside near Grand Canyon N.P., Kaibab Forest, Kearney & Peebles 13744 (NY). W of Jct. of Cape Royal & Point Imperial Rds., Kaibab Plateau, Collom s.n. (US) [i.e. in Grand Canyon National Park]. Marble Flat, N Rim of Grand Canyon, Collom 1049 (US).” ________________________________________________________________________________ 44 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Davis, William T. 1921 Records of cicadas from North America with descriptions of new species. Journal of the New York Entomological Society, 29(1) (March): 1-16.  Tibicen apache, NEW SPECIES (pp. 3-5, Pl. 1, figs. 4-6) 87 Arthropoda: Insecta: Hemiptera: Cicadidae (cicadas) (Holotype and allotype from Florence, Arizona; illustrated.) “The following typical material 6 [paratypes ; not illustrated] of apache from Arizona is in the writer’s collection: . . . Grand Canyon, Lower Bright Angel Trail, August 2, 1917, three males (Dr. Knight) . . . .” ________________________________________________________________________________ DeLong, Dwight M. 1964 A monographic study of the North American species of the genus Ballana (Homoptera: Cicadellidae). Ohio Journal of Science, 64(5) (September): 305-370. 88  Ballana basala, NEW SPECIES (p. 356, figs. 74 [p. 319], 193 [p. 327], 391 [p. 339]) Arthropoda: Insecta: Hemiptera: Cicadellidae (leafhoppers) (Holotype and allotype from Walsenburg, Colorado.) Paratypes include “. . . 5 males, 6 females, Grand Canyon, Arizona, July 28, 1936, R. H. Beamer . . .” Holotype, allotype, and paratypes (not specified) in DeLong collection; other paratypes (not specified) in U.S. National Museum, University of Kansas collection, and Ohio State University collection. ________________________________________________________________________________ 6 Whereas it seems that Davis’ use of the word “typical” may actually mean “usual”, and not being an implication of paratypical material, I have always preferred in preparing catalogs and checklists such as this one to err subjectively on the side of types when they appear +in older literature prior to the established codes of nomenclature, because occasionally it can be demonstrated through ancillary evidence, if not simply inferred, that such material is in fact of type status. The checklist entries can always be dismissed. —E.E.S. 45 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Dempster, Lauramay T., and Ehrendorfer, Friedrich 1965 Evolution of the Galium multiflorum complex in western North America. II. Critical taxonomic revision. Brittonia, 17 (October): 289-334.  Galium munzii Hilend & Howell subsp. ambivalens, NEW SUBSPECIES (pp. 309-310); not 89 illustrated Trachyophyta: Magnoliopsida: Gentianales: Rubiaceae (madders) “Type from ‘about Grand Canyon of the Colorado,’ MacDougal 157 (UC 138660. Isotypes, ARIZ, F, GH, NY, US)”; distribution notes “Grand Canyon of the Colorado River, and eastward on the Little Colorado River.” ________________________________________________________________________________ Doering, Kathleen C. 1939 A contribution to the taxonomy of the subfamily Issinae in America North of Mexico (Fulgoridae, Homoptera). University of Kansas Science Bulletin, 26(2) (October 1): 83167 (including Pls. 13-22).  Bruchomorpha bunni, NEW SPECIES pp. 119-121, Pl. 20, fig. 3, Pl. 21, fig. 15, Pl. 22, figs. 90 4, 4a Arthropoda: Insecta: Hemiptera: Issidae (planthoppers) “Holotype male, taken at Grand Canyon, Arizona, on August 11, 1927, by R. H. Beamer; allotype female, Grand Canyon, August 11, 1927, by P. A. Readio. Two female paratypes and nine male paratypes, same data.” (Other paratypes from Arizona and New Mexico.) “The types are in Snow entomological Collection at the University of Kansas.” ________________________________________________________________________________ Durham, Floyd E. 1952 A new pocket gopher from the North Rim of the Grand Canyon, Arizona. Journal of Mammalogy, 33(4) (November): 498-499. 91  Thomomys bottae boreorarius, NEW SUBSPECIES; not illustrated Chordata: Mammalia: Rodentia: Geomyidae (pocket gophers) “Type.—Adult female, skin and skull, number 1184, Hancock Foundation Collections, University of Southern California, taken at Swamp Point, 7522 ft. alt., 18½ mi. northwestward of Bright Angel Point, North Rim of the Grand Canyon, Coconino County, Arizona; July 10, 1947; Floyd E. Durham; original number, 2141.” ________________________________________________________________________________ 46 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Eastwood, Alice 1902 New western plants. Bulletin of the Torrey Botanical Club, 29(8) (August): 523-525.  Corydalis wetherillii, NEW SPECIES (p. 524); not illustrated 92 Trachyophyta: Magnoliopsida: Ranunculales: Papaveraceae (poppies) Listed under the section, “New Species from Colorado and Utah [sic].” “This peculiar species was collected near Bright Angel Creek, one of the branches of the Colorado River which comes into the river nearly opposite Hans’ Trail [Hance Trail]. This trail is that taken by tourists who go into the cañon from the Flagstaff side. It is named in honor of Mr. Alfred Wetherill who collected it in the summer of 1897.” “The type [holotype] is in the herbarium of the California Academy of Sciences.” _______________________________________________________________ 1903 New species of western plants. Bulletin of the Torrey Botanical Club, 30(9) (September): 483-502.  Fraxinus macropetala, NEW SPECIES (pp. 494-495); not illustrated 93 Trachyophyta: Magnoliopsida: Lamiales: Oleaceae (olives) “This was collected by E. O. Wooton, July 9, 1902, in the Grand Cañon of the Colorado, being no. 1102 of his collection; this specimen is in fruit. The flowers were collected at the same place [i.e., Grand Canyon], on the Bright Angel Trail, by Dr. C. Hart Merriam, the middle of May, 1903. These specimens [syntypes] are both in the Herbarium of the California Academy of Sciences.” _______________________________________________________________ 1937 94 New species of western plants. Leaflets of Western Botany, 2(1) (January): 7-9.  Oreocarya capitata, NEW SPECIES (p. 9); not illustrated Trachyophyta: Magnoliopsida: Boraginales: Boraginaceae (borage) “Type [holotype]: No. 232041, Herb. Calif. Acad. Sci., collected by the author, No. 5969, on the Hermit Trail, Grand Canyon of the Colorado, Arizona, April 9, 1917. This specimen is in flower. A fruiting specimen, Herb Calif. Acad. Sci. No. 232040 [paratype], was collected by the author on the same trail, June 18, 1916. It was collected June 23, 1933, on the north side of the Grand Canyon on the Kaibab Trail to Roaring Springs, Eastwood & Howell No. 1005, Herb. Calif. Acad. Sci. No. 232039 [paratype].” ________________________________________________________________________________ 47 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Ehrhorn, Edw. M. 1899 Five new Coccidæ. The Canadian Entomologist (London), 31(1) (January): 5-7. Arthropoda: Insecta: Hemiptera: Coccidae (soft scales)  Ripersia arizonensis, NEW SPECIES (p. 5); not illustrated 95 “Hab.—In ants’ nest on the roots of grass and Artemisia, sp., Camp Thurber [Hance Ranch], Grand Canyon of the Colorado, Arizona.”  Dactylopius formicarii, NEW SPECIES (pp. 6-7); not illustrated 96 “Hab.—In ants’ nests on the roots of Artemisia, sp. Thurber’s Camp [Hance Ranch], Grand Canyon of the Colorado, Arizona.” ________________________________________________________________________________ Ellis, Don E. 1946 Anthracnose of dwarf mistletoe caused by a new species of Septogloeum. Journal of the Elisha Mitchell Scientific Society, (June): 25-50, Pls. 4-6.  Septogloeum gillii, NEW SPECIES (pp. 30-38, text-fig. 2) 97 Ascomycota (sac fungi) incertae sedis “Parasititc on shoots of Aceuthobium campylopodum and A. douglasii in Western United States.” “The following specimens in the herbarium of the Division of Forest Pathology at Albuquerque, New Mexico, were examined: (Instances in which the disease was found on herbarium collections of the host are indicated by an asterick [sic] (*)).” “On A. campylopodum f. divaricatum (Engelmann) Gill . . . Grand Canyon, Gill, June 6, 1932 (68234)* . . . .” (Other host species examined from specimens gathered within the bounds of the present checklist did not exhibit the disease.) ________________________________________________________________________________ Ellis, J. B., and Everhart, B. M. 1900 New species of fungi from various localities with notes on some published species. Bulletin of the Torrey Botanical Club, 27(2) (February): 49-64. 98  Puccinia circinans, NEW SPECIES (p. 61); not illustrated Basidiomycota (club fungi): Pucciniomycetes: Pucciniales: Pucciniaceae “On Penstemon spectabilis, Grand Cañon of the Colorado (Prof. J. W. Toumey).” ________________________________________________________________________________ 48 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Esslinger, Theodore L. 2000 A key for the lichen genus Physconia in California, with descriptions for three new species occurring within the state. Bulletin of the California Lichen Society, 7(1) (Summer): 1-6.  Physconia isidiomuscigena, new species (pp. 5-6, figs. 5, 6) 99 Ascomycota (sac fungi): Lecanoromycetes: Teloschistales: Physciaceae “Type [holotype]: U.S.A. Arizona. Coconino Co.: Grand Canyon Natl. Park, Grandview Trail, 36°00′N, 111°59′W, 1980 m, Nash 30843 (ASU, holotype; TLE, isotype).” Figs. 5, 6 depict “part of holotype specimen”. [The geographic coordinates pinpoint a location on the upper part of Grandview Trail.] “Selected additional specimens examined ([designated] Paratypes): . . . Grand Canyon Natl. Park, N rim, junction of paved roads, ca. 6.5 km N of Kaibab Lodge, 36° 16′ N, 112° 03′ W [sic, Grand Canyon Lodge, coordinates pinpoint a location just east of Arizona Route 67 north of the Cape Royal Road jct], 2470 m, Nash 9943 (ASU, MIN, TLE); Grand Canyon Natl. Park, South Kaibab Trail, 36° 03′ 45″ N, 112° 03′ 30″ W, 1950 m, Nash 30819 (ASU, MIN). . . .” [In noting the paper’s title, Esslinger comments, “This species is presently known from only two collections in California and is apparently much more common in the southern Rocky Mountains.”] ________________________________________________________________________________ Fall, H. C. 1905 Revision of the Ptinidæ of boreal America. Transactions of the American Entomological Society, 31(2/3): 97-296, Pl. 7. Arthropoda: Insecta: Coleoptera: Ptinidae (spider beetles) 100  Petalium bistriatum Say var. arizonense, NEW VARIETY (p. 217); not illustrated. [P. bistriatum Say described, pp. 216-217.] “A small series [syntypes] collected by Hubbard or Schwarz at Oracle and Bright Angel, Arizona . . . .” 101  Catorama grande, NEW SPECIES (p. 236); not illustrated Species based on specimens [syntypes] seen from Texas and Arizona, including “Bright Angel”. 102  Catorama longulum, NEW SPECIES (p. 244); not illustrated “Described from four examples [syntypes] taken by Barber and Schwarz at Williams and Bright Angel. With these is included a nearly identical specimen taken by the same collectors Las Vegas, New Mexico.” 103  Catorama pingue, NEW SPECIES (p. 250); not illustrated Described on syntypes from “Arizona (Santa Rita Mountains, Oracle, Bright Angel, Pinal Mountains, Williams, Ash Fork)”. _______________________________________________________________ 49 BIBLIOGRAPHY AND ANNOTATED CHECKLIST 1909 Revision of the species of Diplotaxis of the United States. Transactions of the American Entomological Society, 35(1) (January): 1-97. 104  Diplotaxis conformis, NEW SPECIES (p. 34); not illustrated Arthropoda: Insecta: Coleoptera: Scarabaeidae (scarab beetles) Described on syntypes “from Glenwood Springs, Colorado (Fenyes), and from Prescott and Bright Angel, Arizona. Specimens in Mr. Dury’s collection are labeled ‘Northern Arizona.’ ” _______________________________________________________________ 1910 Miscellaneous notes and descriptions of North American Coleoptera. Transactions of the American Entomological Society, 36(2): 89-197. 105  Bruchus perplexus, NEW SPECIES (p. 177); not illustrated Arthropoda: Insecta: Coleoptera: Chrysomelidae (leaf beetles) “The type [holotype] is from Alburquerque [sic], New Mexico (Wickham). With it are associated specimens [paratypes] from Highrolls, New Mexico, Bright Angel, Arizona, Palm Springs, California, and Claremont, California.” ________________________________________________________________________________ Fernald, M. L. 1934 A new Primula from the Grand Canyon of the Colorado. Rhodora (Journal of the New England Botanical Club), 36 (April): 117-118, Pl. 282. 106  Primula (Section Farinosae) hunnewellii, NEW SPECIES; Pl. 282, figs. 1, 2 Trachyophyta: Magnoliopsida: Ericales: Primulaceae (primroses) “Arizona: limestone cliffs, North Rim, Grand Canyon, Coconino Co., August 19, 1928, Francis Welles Hunnewell, no. 10,883 (TYPE [holotype] in herb. F. W. Hunnewell, duplicate [isotype] in Gray Herb.).” (Fig. 3 of the plate illustrates a comparative species, P. specuicola Rydb. from its type locality, Bluff City, Utah.) ________________________________________________________________________________ Ferris, Roxana Stinchfield 1918 Taxonomy and distribution of Adenostegia. Bulletin of the Torrey Botanical Club, 45(10) (October): 299-423. 107  Adenostegia parviflora, NEW SPECIES (p. 409, Pl. 10, fig. 8; Pl. 11, fig. 4 [plate legends p. 422]) Trachyophyta: Magnoliopsida: Lamiales: Orobanchaceae (broomrapes) “TYPE LOCALITY: Grand Canyon of the Colorado River, near the San Francisco Mountains, Arizona, Knowlton 270. TYPE [holotype], No. 48859 of the U. S. National Herbarium.” ________________________________________________________________________________ 50 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Fischer, Pierre C. 1979 Coryphantha vivipara var. kaibabensis, a new variety from northern Arizona. Cactus and Succulent Journal (Cactus and Succulent Society of America), 51(6): 286-287. 108 Coryphantha vivipara var. kaibabensis, NEW VARIETY . [Illustrations depict living specimens.] Trachyophyta: Magnoliopsida: Caryophyllales: Cactaceae (cacti) “HOLOTYPE P.C. Fischer 4094, collected on 31/III/69 northwest of Jacob Lake, Coconino Co., Arizona. 1890 meters (6200 ft). Growing with Juniper, Pinyon Pine, Sagebrush (Artemisia tridentata) and Cliff Rose (Cowania sp.). Growing on dolomite and limestone. Deposited at the herbarium of the University of California (UC), Berkeley, CA.” _______________________________________________________________ 1991 Echinocereus triglochidiatus variety toroweapensis, a new variety from the Grand Canyon. Cactus and Succulent Journal (Cactus and Succulent Society of America), 63(4) (July/August): 194-195. 109  Echinocereus triglochidiatus var. toroweapensis, NEW VARIETY; [live specimens illustrated] Trachyophyta: Magnoliopsida: Caryophyllales: Cactaceae (cacti) “Holotype. P. C. Fischer 7196 collected April 7, 1990, Toroweap Point, Grand Canyon National Park, Mohave Co., Arizona; 1,340 meters (4,400 ft). [. . .] Deposited at the herbarium of the University of Arizona (Ariz.), Tucson.” ________________________________________________________________________________ Foissner, Ilse, and Foissner, Wilhelm 1993 Revision of the family Spironemidae Doflein (Protista, Hemimastigophora), with description of two new species, Spironema terricola n. sp. and Stereonema geiseri n. g., n. sp. Journal of Eukaryotic Microbiology, 40(4): 422-438. 110  Spironema terricola, NEW SPECIES (pp. 424-429, figs. 5-34, 59, 62, 65) Hemimastigophora: Hemimastigea: Hemimastigida: Spironemidae (hemimastigid protist) “Type location. Soil from the Grand Canyon, USA (upper entrance to Bright Angel Trail), 36° N, 112° E [sic]. Type specimens. One slide of holotype specimens and three slides of paratype specimens (all protargol-impregnated) have been deposited in the collection of microscope slides of the Oberösterreichische Landesmuseum in Linz, Austria. Accession numbers: 44, 45, 46, 47 /1992.” [Not only are the coordinates in error (“E” instead of “W”) but they are overly generalized, that geographical intersection being 8¾ miles southeast of the head of Bright Angel Trail.] ________________________________________________________________________________ 51 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Fox, William J. 1893 New species of fossorial Hymenoptera. The Canadian Entomologist (London), 25(5) (May): 113-117. 111  Gorytes dentatus, NEW SPECIES (pp. 116-117); not illustrated Arthropoda: Insecta: Hymenoptera: Crabronidae (crabronid wasps) Holotype male from “Grand Canyon, Arizona, ‘70 miles North of Flagstaff.’ (C. H. Tyler Townsend).” [Townsend visited the canyon in July 1892 at Hance Ranch and trail. For his general (non- scientific) account of the trip see Townsend, 1893, A wagon-trip to the Grand Cañon of the Colorado River, Appalachia, 7(1) (February): 48-63.] [Note: Townsend (1896, Notes on New Mexico and Arizona Hymenoptera, The Canadian Entomologist (London), 28(5) (May): 138-142), p. 139 there lists “Gorytes dentatus Fox., n. sp. —Grand Canyon, Arizona; Hance trail, July 11. One specimen. [. . .] Det., Fox.” It is of course not a new species therein but the record further substantiates the identification of the locality as the Old Hance Trail.] ________________________________________________________________________________ Garth, John S. 1949 Studies in Arizona Lepidoptera. I. A new subspecies of Speyeria atlantis (Edwards) from the Kaibab Plateau, Grand Canyon National Park. Bulletin of the Southern California Academy of Sciences, 48 (Part 1): 1-4. 112  Speyeria atlantis schellbachi, NEW SUBSPECIES; illustrated Arthropoda: Insecta: Lepidoptera: Nymphalidae (admirals) “Type material: Male holotype, AHF No. 471, and female allotype, AHF No. 471a, from Neal Spring, North Rim, Grand Canyon National Park, Coconino County, Arizona, 8,175 feet, July 5, 1947, collected by John S. Garth, Allan Hancock Foundation survey party. Twenty paratypes as follows: 1 female, North Rim, Grand Canyon, July 29, 1939, Louis Schellbach, collector; 1 female, North Rim, Grand Canyon, August 19, 1942, H. C. Bryant, collector; 1 male, 1 female, Two River Junction, Grand Canyon, July 28, 1945, Louis Schellbach, collector, the preceding four paratypes on loan from the Naturalist Workshop, Grand Canyon National Park; 1 male, 1 female, Neal Spring, North Rim, Grand Canyon, August 16, 1946, J. S. Garth, collector; 2 females, Kanabownits Spring, North Rim, Grand Canyon, August 22, 1946, J. S. Garth, collector; 7 males, 2 females, Neal Spring, North Rim, Grand Canyon, July 5 to 18, 1947, J. S. Garth, collector; 1 male, Robbers’ Roost Spring, North Rim, Grand Canyon, July 10, 1947, J. S. Garth, collector; 2 females, Swamp Lake and Swamp Ridge, North Rim, Grand Canyon, July 12 and 14, 1947, J. S. Garth, collector. The holotype, allotype, and ten paratypes are in the collection of the Allan Hancock Foundation, the University of Southern California. The remaining paratypes will be distributed as follows: one male and three females to the Naturalist 52 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Workshop, Grand Canyon National Park, one pair each to the United States National Museum, the American Museum of Natural History, and the Los Angeles County Museum.” [The type locality, Neal Spring, is by the junction of Cape Royal Road and Point Imperial Road.] ________________________________________________________________________________ Gertsch, Willis J. 1933 New genera and species of North American spiders. American Museum Novitates, (636), 28 pp. 113  Misumenops coloradensis, NEW SPECIES (p. 17, figs. 15 [p. 12], 46 [p. 24]) Arthropoda: Euchelicerata: Areneae: Thomisidae (crab spiders) (“Male holotype, female allotype, and paratypes of both sexes from Colorado.”) “Female paratypes from Grand Canyon (North Rim), Arizona, and Kanab, Utah.” _______________________________________________________________ 1934a Notes on American Lycosidae. American Museum Novitates, (693), 25 pp. 114  Arctosa mokiensis, NEW SPECIES (p. 8); not illustrated [but see note below] Arthropoda: Euchelicerata: Areneae: Lycosidae (wolf spiders) Holotype male. “Type Locality.—Male holotype from Indian Gardens [Havasupai Gardens], Grand Canyon, Arizona, May 26, 1905.” In synonymy is “Arctosa notuabanda Gertsch, 1933, American Museum Novitates, No. 637, p. 8, fig. 11 (not noctuabunda Montgomery)”, which was noted therein only by the name Arctosa noctuabunda Montgomery in the legend for fig. 11 and is not mentioned in that text. [It is not clear whether this represents a figure of the type of A. mokiensis.] _______________________________________________________________ 1934b 115 Further notes on American spiders. American Museum Novitates, (726), 26 pp. Phidippus kaibabensis, NEW SPECIES (pp. 13-14, fig. 19 [p. 17]) Arthropoda: Euchelicerata: Areneae: Salticidae (jumping spiders) “Male holotype from the Kaibab forest, near the north rim of the Grand Canyon, Arizona, July 8, 1931 (Gertsch).” _______________________________________________________________ 53 BIBLIOGRAPHY AND ANNOTATED CHECKLIST 1960 The fulva group of the spider genus Steatoda (Araneae, Theridiidae). American Museum Novitates, (1982), 48 pp. 116 Steatoda variata, (pp. 24-29, figs. 22-25 [p. 13], 34-44 [pp. 17, 26]) NEW SPECIES Arthropoda: Euchelicerata: Areneae: Theridiidae (cobweb weavers) “Type Locality: Male holotype from 25 miles north of The Gap, Navajo Indian Reservation, Arizona, July 21, 1949 (W. J. and J. W. Gertsch). (Non-type records include “Twenty-five males north of The Gap . . . male, many females.”) ________________________________________________________________________________ Gertsch, Willis J., and Ennik, Franklin 1983 The spider genus Loxosceles in North America, Central America, and the West Indies (Araneae, Loxoscelidae). Bulletin of the American Museum of Natural History, 175 (Article 3): 264-360. 117  Loxosceles kaiba, NEW SPECIES (p. 303, figs. 110-113 [p. 299], 133 [p. 300]) Arthropoda: Euchelicerata: Areneae: Sicariidae (recluse spiders) “Type Data: Male holotype, female and four immatures [paratypes] from Thunder Cave [Thunder River Cave], 3900 feet, near Monument Point, north rim of Grand Canyon National Park, Coconino County, Arizona, 15 September 1977 (S. Peck, D. Carlile), deposited in AMNH. Other Record [non-type?]: Grand Canyon National Park, Cameron Cave, 5000 ft., 7 Dec. 1954 (R. de Saussure), ♂.” The specific epithet, kaiba [sic], is named “for Kaibab Plateau on north face of Grand Canyon” (p. 303). ________________________________________________________________________________ Goldman, Edward A. 1924 Two new kangaroo rats from Arizona. Journal of the Washington Academy of Sciences (Washington, D.C.), 14(15) (September 19): 372-373. Chordata: Mammalia: Rodentia: Heteromyidae (kangaroo rats) 118 Dipodomys microps celsus, NEW SUBSPECIES (p. 372); not illustrated. (Virgin Valley Kangaroo Rat) “Type [holotype] from 6 miles north of Wolf Hole, Arizona (altitude 3,500 feet). No. 243101, ♂adult, U. S. National Museum (Biological Survey Collection) collected by E. A. Goldman, October 16, 1922. Original number 23411.” 119 Dipodomys ordii cupidineus, NEW SUBSPECIES (pp. 372-373); not illustrated. (Kaibab Kangaroo Rat) Type [holotype] from Kanab Wash, at southern boundary of Kaibab Indian Reservation, Arizona. No. 243093, ♂ adult, U. S. National Museum (Biological by E. A. Goldman, October 12, 1922. Original number 23384.” 54 Survey Collection), collected BIBLIOGRAPHY AND ANNOTATED CHECKLIST _______________________________________________________________ 1929 A new antelope squirrel from Arizona. Journal of the Washington Academy of Sciences (Washington, D.C.), 19: 435-436. 120  Ammospermophilus leucurus tersus, NEW SUBSPECIES; not illustrated. (Grand Canyon Antelope Squirrel) Chordata: Mammalia: Rodentia: Sciuridae (squirrels) “Type [holotype].—From lower end of Prospect Valley, Grand Canyon, Hualpai Indian Reservation [Hualapai Indian Reservation], Arizona (altitude 4,500 feet). No. 202645, ♂ young adult, U. S. National Museum (Biological Survey collection), collected by E. A. Goldman, October 3, 1913. Original number, 22269.” “Specimens examined.—Ten, all from the type locality.” [Examined specimens not itemized.] _______________________________________________________________ 1932 A new beaver from Arizona. Journal of Mammalogy, 13(3) (August): 266-267. 121  Castor canadensis repentinus, NEW SUBSPECIES; not illustrated. (Colorado River Beaver) Chordata: Mammalia: Rodentia: Castoridae (beavers) “Type [holotype] From Bright Angel Creek, Grand Canyon of the Colorado River, Arizona (altitude 4,000 feet); no. 161613, ♀ adult, U.S. National Museum (Biological Survey collection); collected by Clarence Birdseye, September 14, 1909; original number 72.” Two specimens examined from type locality (no specimen or collecting data published for the second specimen). “At the type locality the beavers inhabit Bright Angel Creek, which in short reaches descends the terraced north side, below the out rim, of the Grand Canyon. None are known from the Colorado River in that vicinity, and a measure of isolation would seem to be due to the rapids and heavy current through rock-bound gorges extending for many miles, and affording very few places suitable for beavers to establish homes.” _______________________________________________________________ 1937 A new canyon mouse of the genus Peromyscus from Arizona. Journal of Mammalogy, 18(1) (February): 92-93. 122  Peromyscus crinitus peridoneus, NEW SUBSPECIES; not illustrated. (Grand Canyon Mouse) Chordata: Mammalia: Rodentia: Cricetidae (deer mice) “Type [holotype].—From Bright Angel Trail, 4800 feet altitude, south side of Grand Canyon, Coconino County, Arizona. No. 202,424, ♂ adult, skin and skull, U. S. National Museum (Biol. Surv. coll.); collected by E. A. Goldman, August 19, 1913. Original number 22,145.” [Altitude on Bright Angel Trail indicates locality is at about the 2nd rest house descending.] “Specimens examined [non-type].—Total number, 36, as follows: Arizona: Fredonia, 1; Grand Canyon (near Bass Camp, 3000-5200 feet), 3; near Grand Canyon Spring, 4000-7500 feet, 9; Betatakin Canyon, 3; Bright Angel Creek, 3 miles above mouth, 1; Bright Angel Trail, south side, 4800-6500 feet, 2; Shinumo Creek, 3000 feet, 12; Nankoweap Valley, 1. near Fort Cameron, 3; Parowan, 1.” 55 Utah: Beaver River, BIBLIOGRAPHY AND ANNOTATED CHECKLIST _______________________________________________________________ 1938 New pocket gophers of the genus Thomomys from Arizona and Utah. Journal of the Washington Academy of Sciences (Washington, D.C.), 28(7) (July 15): 333-343. Chordata: Mammalia: Rodentia: Geomyidae (pocket gophers) 123 Thomomys fossor kaibabensis, NEW SUBSPECIES (pp. 333-334); not illustrated. (Common name given by the author, “Kaibab Pocket Gopher”) “Type [holotype].—From DeMotte Park, Kaibab Plateau, Coconino County, Arizona (altitude 9,000 feet). No. 262891, ♂ adult, skin and skull, U. S. National Museum (Biological Survey collection), collected by Luther C. Goldman, September 10, 1937. Original number 443.” “Specimens examined [non-type].—Total number, 23, all from Kaibab Plateau, Arizona, as follows: Bright Angel Spring, 3; DeMotte Park (type locality), 18; Greenland Spring, 1; Jacob Lake, 1.” 124 Thomomys bottae nicholi, NEW SUBSPECIES (pp. 337-338); not illustrated. (Common name given by the author, “Shivwits Plateau Pocket Gopher”) “Type [holotype].—From 20 miles south of Wolf Hole (road to Parashouts), Shivwits Plateau, Mohave County, Arizona (altitude 5,000 feet). No. 262864, ♂ adult, skin and skull, U. S. National Museum (Biological Survey Collection), collected by Luther C. Goldman, August 6, 1937. Original number 363.” “Specimens examined [non-type].—Total number, 19, as follows: Arizona: Diamond Butte (3 miles northwest), 1; Wolf Hole, 6 (20 miles south, 4; 6 miles north, 2). Utah: St. George, 12.” ________________________________________________________________________________ Gray, Asa 1878 Contributions to the botany of North America. Proceedings of the American Academy of Arts and Sciences, 13: 361-374. Trachyophyta: Magnoliopsida: Fabales: Fabaceae (legumes) 125 Astragalus subcinereus, NEW SPECIES (p. 366); not illustrated “Mokiak Pass in the northwestern part of Arizona, near the Utah boundary, Dr. E. Palmer, 1877.” 126 Astragalus scaposus, NEW SPECIES (p. 366); not illustrated “Among rocks in dry creek-bottoms, in Mokiak Pass, near the northeastern [sic] corner of Arizona, Dr. E. Palmer, 1877.” [Gray correctly cites the location of Mokiak Pass elsewhere.] 127 Astragalus mokiacensis, NEW SPECIES (p. 367); not illustrated “Rocky ravines, Mokiak Pass, on the borders of Utah and N. W. Arizona, Dr. Palmer, 1877.” 128 Astragalus artipes, NEW SPECIES (p. 370); not illustrated “Mokiak Pass, northwest corner of Arizona, in ravines, Dr. E. Palmer, 1877.” 56 BIBLIOGRAPHY AND ANNOTATED CHECKLIST 129 Actinella biennis, NEW SPECIES (p. 373); not illustrated Trachyophyta: Magnoliopsida: Asterales: Asteraceae (sunflowers) “S. Utah and Arizona, Mokiak Pass south of St. George, Palmer (no. 260 of coll. 1877)” [syntype; another from Richfield, Utah]. _______________________________________________________________ 1886 Contributions to American botany. Proceedings of the American Academy of Arts and Sciences, 21: 363-413. 130 Mirabilis bigelovii, NEW SPECIES (p. 413); not illustrated Trachyophyta: Magnoliopsida: Caryophyllales: Nyctaginaceae (four o’clocks) “Arizona, and perhaps in California on the Colorado. Common in the Grand Cañon of the Colorado, below Peach Spring [Peach Springs], collected May 5, 1885, A. G.” [indicating syntypes]. [Gray descended into Grand Canyon in Peach Springs Canyon to Diamond Creek and the Colorado River. Today the Grand Canyon localities “below Peach Spring” are in the Hualapai Indian Reservation.] ________________________________________________________________________________ Gray, Asa in Professors Gray, Torrey, Thurber, and Dr Engelmann [in Gray, Asa; Torrey, John; Thurber, George; and Engelmann, George] 1861 Botany. (“By Professors Gray, Torrey, Thurber, and Dr Engelmann”.) In: Ives, Joseph C., Report upon the Colorado River of the West, explored in 1857 and 1858 by Lieutenant Joseph C. Ives, Corps of Topographical Engineers, under the direction of the Office of Explorations and Surveys, A. A. Humphreys, Captain Topographical Engineers, in charge. By order of the Secretary of War. U.S. 30th Congress, 1st Session, House and Senate Executive Document 90. Washington, D.C.: U.S. Government Printing Office [1861], Part 4, 30 pp. [Botany section, with 1860 date printed on title leaf, apparently arranged or edited by John Strong Newberry; see p. 20, footnote.] The collecting localities noted here are now within the Hualapai Indian Reservation. 131 Phacelia (Coreanthus) ivesiana, NEW SPECIES (p. 21); not illustrated Trachyophyta: Magnoliopsida: Boraginales: Hydrophyllaceae (waterleafs) One of three localities noted is “Diamond river; April 3” (entire note) [indicating syntypes]. Not illustrated. 132 “Gilia dactylophyllum, (n. sp.?)” (p. 22); not illustrated Trachyophyta: Magnoliopsida: Ericales: Polemoniaceae (phlox) [It is unclear whether this entry refers to G. dactylophyllum Torr. (which itself is now in another genus), or whether Gray had questionably introduced the name or left his specimens open to 57 BIBLIOGRAPHY AND ANNOTATED CHECKLIST receive a new name in the future (see his remarks below); and so retained here with question as well.] “Mouth of Diamond river; April 3. The specimens [syntypes] are scarcely sufficient for description. A slender plant, about 3 inches high. Corolla twice as long as the calyz, white; ovules about 7 in each cell.” (entire entry). ________________________________________________________________________________ Greene, Edward Lee 1890 Bibliographical notes on well known plants.—X. Bulletin of the Torrey Botanical Club, 17(1) (January 15): 13-14. 133  Rhus canadensis Marsh. var. simplicifolia, NEW VARIETY, p. 13 (footnote); not Trachyophyta: Magnoliopsida: Sapindales: Anacardiaceae illustrated (cashews) “In the deep cañons of Northern Arizona which lead down to the Grand Cañon of the Colorado”. [Identity of these canyons uncertain and not resolved here based on ecological potential, which could well include the side canyons in which the more famous trails descend into the canyon—Hance Trail, Grandview Trail, Bright Angel Trail.] _______________________________________________________________ 1900 Studies in the Cruciferae.—III. Pittonia, 4 [1899-1901]: 187-207. Trachyophyta: Magnoliopsida: Brassicales: Brassicaceae [Cruciferae] (mustards) 134 Arabis eremophila, NEW SPECIES (pp. 194-195); not illustrated “Common about Peach Springs, northern Arizona . . . .” [No further details. Syntypic material from this locality and from Aztec, New Mexico.] 135 Arabis recondita, NEW SPECIES (p. 195); not illustrated “Diamond Creek Cañon, a tributary of the Colorado, in northern Arizona, N. C. Wilson, April 1893.” [Syntypic material from this locality and from Glenwood Springs, Colorado.] [It is unclear whether “Diamond Creek Cañon” refers to Diamond Creek or more generally to Peach Springs Canyon, to which Diamond Creek is a tributary close to its confluence with the Colorado River, the former being the long principal route into Grand Canyon from Peach Spring s.] _______________________________________________________________ 1905 Segregates of the genus Rhus. Leaflets of Botanical Observation and Criticism, 1: 114144. [Date of publication for pp. 129-144, November 29, 1905 (see p. 129).] Trachyophyta: Magnoliopsida: Sapindales: Anacardiaceae (cashews) 136  Schmaltzia cissodes, NEW SPECIES (p. 136); not illustrated “Grand Cañon of the Colorado, Ariz., near Indian Garden [Havasupai Gardens], Bright Angel Trail, C. H. Merriam, 10 May, 1903.” 58 BIBLIOGRAPHY AND ANNOTATED CHECKLIST 137  Schmaltzia hirtella, NEW SPECIES (p. 137); not illustrated “Grand Cañon of the Colorado, Ariz., 10 July, 1892, E. O. Wooton.” _______________________________________________________________ 1906 The genus Ptelea in the western and southwestern United States and Mexico. Contributions from the U.S. National Herbarium, 10(2) (July 16): 149-179. Trachyophyta: Magnoliopsida: Sapindales: Rutaceae (rues) 138 Ptelea palida, NEW SPECIES (p. 70); not illustrated “Arid rocky hills above Peach Springs, northern Arizona, collected by the writer July 3, 1889; type in the National Herbarium.” 139 Ptelea straminea, NEW SPECIES (p. 70); not illustrated “Rocky slopes of the Virgin Mountains in extreme northwestern Arizona, C. A. Purpus, 1898; his no 6165 as in the national Herbarium.” 140  Ptelea nitida, NEW SPECIES (p. 71); not illustrated “Species known to me only as in the Herbarium of the California Academy from somewhere in the Grand Canyon of the Colorado in Arizona, collected by E. O. Wooton, July 8, 1892.” 141  Ptelea argentea, NEW SPECIES (p. 71); not illustrated “This species . . . is known to me only as collected by Dr. F. H. Knowlton, somewhere in the Grand Canyon of the Colorado, September 10, 1889, the specimens filling two sheets in the national Herbarium (one the type).” 142  Ptelea triptera, NEW SPECIES (pp. 71-72); not illustrated “Grand Canyon of the Colorado, Arizona, excellent specimen in the National Herbarium, collected by J. W. Toumey, July 11, 1892.” [Very probably gathered near Hance Ranch; compare date and remarks with Ostrya knowltonii Coville, 1894.] 143  Ptelea lutescens, NEW SPECIES (p. 72); not illustrated “Species of northwestern Arizona, inhabiting canyons tributary to the Grand Canyon of the Colorado, the best specimen from Red Canyon Trail [Hance Trail], collected June 10, 1901, by Lester F. Ward (type [holotype] in the National Herbarium). Younger material, with fruit not mature, from Bright Angel Trail, by Dr. C. Hart Merriam, May 10, 1903. These last imperfect specimens have smaller relatively broader leaflets, and may possibly represent another species [paratypes, uncertainly].” 144  Ptelea elegans, NEW SPECIES (p. 72); not illustrated “Grand Canyon of the Colorado, Arizona, June 14, 1891, D. T. McDougal [sic], the shrub said to grow ‘in rich soil and in water.’ The only specimens seen are in the National Herbarium [syntypes].” 59 BIBLIOGRAPHY AND ANNOTATED CHECKLIST [Daniel Trembly MacDougal visited Hance Ranch in June 1891 and signed the Hance guest book. He likely also descended into the canyon on the Old Hance Trail, down which probably is the type locality for this species, in as much as there is no standing water on the rim of the canyon except at a reservoir maintained by John Hance for the use of his hotel and riding stock. (The upper portion of this trail was destroyed by landslides in 1895, after which Hance rerouted his trail into Red Canyon, which today is the “Hance Trail”, which should not be confused with the original trail, now barely not even a route and referred to as the Old Hance Trail.) See record of MacDougal’s guestbook signature in G[eorge]. K. Woods, 1899, Personal impressions of the Grand Cañon of the Colorado River near Flagstaff, Arizona, as seen through nearly two thousand eyes, and written in the private visitors’ book of the world-famous guide Capt. John Hance, guide, storyteller, and path-finder San Francisco: Whitaker and Ray Co., for G. K. Woods, Flagstaff, Arizona Territory. MacDougal added no comments to his signature.] 145  Ptelea confinis, NEW SPECIES (p. 72); not illustrated “. . . inhabiting the region of the Grand Canyon of the Colorado in northwestern Arizona and adjacent Utah. The material examined is on four sheets in the National Herbarium, among which I would name that on sheet 15254 as the type [holotype].” 146 Ptelea saligna, NEW SPECIES (p. 73); not illustrated “Nagle’s Ranch, Arizona, altitude 2,240 meters, M. E. Jones, September 15, 1894, no. 6048, as in the National Herbarium (type) and the Herbarium of the California Academy. [. . .] The locality is in northern Arizona, north of the Grand Canyon.” _______________________________________________________________ 1909 New plants from Arizona. Leaflets of Botanical Observation and Criticism, 2: 20-24. [Date of publication for pp. 1-24, 6 February 1909 (see p. 1).] 147 Senecio stygius, NEW SPECIES (p. 21); not illustrated Trachyophyta: Magnoliopsida: Asterales: Asteraceae (sunflowers) “Grand Cañon of the Colorado, J. G. Lemmon, May, 1884. In U. S. Herb., the sheet [holotype] not numbered.” [Lemmon, with his wife, spent a month in the Peach Springs area in May 1884. For a general (non-scientific) account of their visit, see J. G. Lemmon, 1888, Grand Cañon of the Colorado, Overland Monthly, new series, 12(69) (September): 244-256.] _______________________________________________________________ 1911 Some southwestern mulberries. Leaflets of Botanical Observation and Criticism, 2: 112121. [Date of publication for pp. 105-120, 6 October 1910 (see p. 105); date for pp. 121-152, 11 May 1911 (see p. 121).] 148 Morus grisea, NEW SPECIES (p. 121); not illustrated Trachyophyta: Magnoliopsida: Rosales: Moraceae (mulberries) “This fine specimen of this is on U. S. Herb. sheet 41506, and purports to have been collected forty years ago, by Dr. Edw. Palmer, at ‘Hell Cañon, Arizona’; I presume in the northwestern 60 BIBLIOGRAPHY AND ANNOTATED CHECKLIST part of the State, in some arm or tributary of the Grand Cañon.” [Date of publication for pp. 105 120, 6 October 1910 (see p. 105); date for pp. 121-152, 11 May 1911 (see p. 121).] [“Hell Canyon” could have been an informal name for Palmer’s locality, and it is not clear why Greene would have supposed it to be “in some arm or tributary of the Grand Cañon” other than through some familiarity with Palmer’s collecting activities. Consulting Will C. Barnes’ Arizona Place Names, revised and enlarged by Byrd H. Granger (University of Arizona Press, 1960), p. 346, the only “Hell Canyon” recognized therein is in the present Yavapai County, which notes, “this canyon was part of the route from the old Beale Trail into Chino Valley in the early days” (i.e. between Ash Fork and Prescott). Greene’s record is retained herein in the event that Palmer’s “Hell Canyon” is in fact elsewhere in northwestern Arizona.] ________________________________________________________________________________ Grijseels, Sietske; Nielsen, Jens Christian; Randelovic, Milica; Nielsen, Jens; Nielsen, Kristian Fog; Workman, Mhairi; and Frisvad, Jens Christian 2016 Penicillium arizonense, a new, genome sequenced fungal species, reveals a high chemical diversity in secreted metabolites. Scientific Reports (Springer Nature), 6(35112), doi:10.1038/srep35112, 13 pp. 149  Penicillium arizonense Frisvad, Grijseels & J.C. Nielsen, NEW SPECIES Ascomycota (sac fungi): Eurotiomycetes: Eurotiales: Trichocomaceae (saprobe fungi) “A new soil-borne species belonging to the Penicillium section Canescentia is described, Penicillium arizonense sp. nov. (type strain CBS 141311 T=IBT 12289T).” “Type: Herb. C-F-101845, cultures ex type IBT 12289=CBS 141311, from a sample of dry red soil, south rim of Grand Canyon, Grand Canyon Village, Arizona, USA (36° 3′ 22.31″ N; 112° 7′ 30.73″ W), Per V. Nielsen [Per Væggemose Nielsen], July 1990”; additional cultures “from same source as the culture ex type, but not of the same clone.” [Map coordinates pinpoint a locality just to the west of the footpath between Park Headquarters and the Rim Trail, at a point half-way between those two places (https://www.findlatitudeandlongitude.com/, accessed 28 October 2022).] ________________________________________________________________________________ Grissell, E. E. 1983 Boharticus, n. gen., with a review of Rhopalicus Foerster and Dinotiscus Ghesquiere (Hymenoptera: Pteromalidae). Pan-Pacific Entomologist, 59(1/4): 78-102. 150 Boharticus margaretae, NEW SPECIES (pp. 86-87, figs. 2 [p. 81], 13 [p. 85], 17 [p. 85]) Arthropoda: Insecta: Hymenoptera: Pteromalidae (pteromalid wasps) (Holotype female from Garden of the Gods, El Paso Co., Colorado; USNM 100593.) “Paratypes: 8 ♀, same data a holotype, 1 ♀, ARIZONA, Mohave Co., 8 miles SW Peach Springs, 4500′, 7 ♀ in British September 1964, C. W. O’Brien, ex Juniperus. All specimens in USNM, except 1 Museum (Natural History).” [The locality is in the vicinity of Truxton, Arizona.] 61 BIBLIOGRAPHY AND ANNOTATED CHECKLIST The type species of Boharticus Grissell, new genus, is designated as B. richardi Grissell, new species (which is not pertinent to this checklist). ________________________________________________________________________________ Hall, E. Raymond, and Davis, William B. 1934 Notes on Arizona rodents. Proceedings of the Biological Society of Washington (Washington, D.C.), 47(5): 51-56. 151  Thomomys bottae trumbullensis, NEW SUBSPECIES (pp. 51-53); not illustrated Chordata: Mammalia: Rodentia: Geomyidae (pocket gophers) “Type [holotype].—Male, adult, skin-and-skull; no. 58588, Mus. Vert. Zool.; three miles south of Nixon Spring, Mt. Trumbull, Mohave Co., Arizona; May 26, 1933; collected by Seth B. Benson, original no. 2005.” [The locality, “three miles south of Nixon Spring”, is just to the south of Mount Trumbull itself, today within the Grand Canyon–Parashant National Monument, about 2¾ miles north of the boundary of Grand Canyon National Park.] “Specimens examined [non-type]. — Total number, 31, as follows: From Mohave County, Arizona: Nixon Spring, 6250 ft., Mt. Trumbull, 8; three miles south of Nixon Spring (volcanic sand), Mt. Trumbull (type locality), 3; four miles south of Nixon Spring (volcanic sand), Mt. Trumbull, 13; head of Toroweap Valley, 2; six miles north of Wolf Hole, 4900 ft., 3; south side Virgin River, St. George, Washington County, Utah, 2.” ________________________________________________________________________________ Hall, Jack C., and Evenhuis, Neal L. 2003 Review of the subgenus Geron (Geron) Meigen in the Nearctic region (Diptera: Bombyliidae: Toxophorinae). Zootaxa, (181): 1-72. 152  Geron prosopidis, NEW SPECIES Arthropoda: Insecta: Diptera: Bombyliidae (bee flies) Paratypes include 11 taken at “S Rim Grand Canyon, 21-22.vii.1932 (R.H. Painter)”. ________________________________________________________________________________ Harvey, Mark S., and Wynne, J. Judson 2014 Troglomorphic pseudoscorpions (Arachnida: Pseudoscorpiones) of northern Arizona, with the description of two new short-range endemic species. Journal of Arachnology, 42(3): 205-219. Arthropoda: Euchelicerata: Pseudoscorpiones (pseudoscorpions): Chernetidae 153 Hesperochernes bradybaughi, NEW SPECIES (pp. 211-214, figs. 15-26) “Types. U.S.A.: Arizona: Mohave County: holotype male, PARA-1001 Cave, Grand CanyonParashant National Monument, ca. UTM 0264500 N, 4060700 E, Zone 12S, baited pitfall trap 3B, 20 August 2007, J.J. Wynne (MNA); 1 female, same data as holotype except baited pitfall 62 BIBLIOGRAPHY AND ANNOTATED CHECKLIST trap 5A (MNA); 1 female, same data as holotype except opportunistic, mid cave, 13 August 2005 (NMA [sic]).” 154 Tuberochernes cohni, NEW SPECIES (pp. 214-217) “Type: U.S.A.: Arizona: Mohave County: holotype male, PARA-1001 Cave, Grand CanyonParashant National Monument, ca. UTM 0264500 N, 4060700 E, Zone 12S, the deeper extent of the twilight zone (near the dark zone), opportunistic collecting, 13 August 2005, J.J. Wynne (MNA).” ________________________________________________________________________________ Hayward, C. Lynn 1932 The paper wasps of Utah; including a description of a new variety of Polistes canadensis Linn. Proceedings of the Utah Academy of Sciences, 9: 85–101 (pagination includes Pl. 9). 155  Polistes canadensis var. kaibabensis, NEW VARIETY (pp. 89-90, Pl. IX, figs. 4, 12, 14, 16 [p. 101, legend p. 100]) Arthropoda: Insecta: Hymenoptera: Vespidae (hornets) “HOLOTYPE: Female; Number 28, collection of Brigham Young University, Provo, Utah. Collected at Point Sublime, north rim of Grand Canyon of the Colorado River, Arizona, July 19, 1927 by Vasco M. Tanner. ALLOTYPE: Male; Number 29, collection of Brigham Young University, Provo, Utah. Collected at North Rim, Grand Canyon of the Colorado, Arizona, July 9, 1929 by Lowell Woodbury. PARATYPES: 22 females; all in collection of Brigham Young University, Provo, Utah; collected at Point Sublime, Rim of Grand Canyon of the Colorado River, Arizona, July 19, 1927 (Vasco M. Tanner, Clarence Cottom, Irvin Rasmussen, Anson Call). TYPE LOCALITY: Point Sublime, Rim of Grand Canyon of Colorado River, Arizona.” The author notes: “The variety kaibabensis is at present known only from the immediate vicinity of the type locality. I have had the privilege of examinng a considerable series from the collection of the University of Utah, Salt Lake City, which were taken in the type locality and could well be designated paratypes. I have also seen a very typical specimen from the San Diego Natural History Museum which was collected at El Tovar, South Rim of the Grand Canyon. This specimen is now in that museum.” [In consideration of the fact that the new variety is recorded only from the vicinity of Point Sublime, the reference to the allotype’s locality as “at North Rim, Grand Canyon of the Colorado” seems to imply that it is from the place, North Rim, as opposed to generically the canyon’s North Rim. It seems that the “considerable series” of specimens in the University of Utah should be excluded as paratypes; and the El Tovar specimen is a referred specimen only.] ________________________________________________________________________________ 63 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Henderson, Junius 1914 A new Sonorella from the Grand Canyon, Arizona. The Nautilus (Philadelphia), 27(11) (March): 122-124. 156  Sonorella betheli, NEW SPECIES; not illustrated Mollusca: Gastropoda: Stylommatophora: Helminthoglyptidae (shoulderbands) NOTE: The locality is in error, not of the Grand Canyon, but this species is included in this nomenclator to account for the published record of a new taxon named upon material ostensibly collected at Grand Canyon. The type specimens, though also subsequently reidentified taxonomically, still are the name-bearing types of the taxon Sonorella betheli. Refer to the information below. “Type [holotype] (in Univ. Colo. Museum) . . . . Cotype [paratype] (in Acad. Nat. Sci. Phila.) . . . . Type locality, Bright Angel Trail, Grand Canyon, Arizona.” Henderson: “Among some shells recently received from Mr. Ellsworth Bethel, of East Denver High School, were four dead specimens of Sonorella collected by him on Bright Angel Trail, at Grand Canyon, Arizona, in 1913. He was collecting fungi, and unfortunately did not note the exact locality of the snail find, but writes that he followed the trail closely, and thinks he got the shells ‘about one hundred yards west of the upper limit of the trail and not more than twenty feet below the top,’ though he cannot be certain and ‘may have gotten them as far down as the half way house.’ He supposed them to be common and made no note of the place. They are much larger than the common S. coloradoensis [refer to Helix (Arionta) coloradoensis Stearns, 1890, No. 280 in the present checklist] of that region, and differ in other respects. They did not seem to fit the description of any other species, but the finding of so large a species along a trail which has been searched by some of our ablest conchologists and most thorough collectors made me doubt that it could be new, so I sent two specimens to Dr. Pilsbry, who pronounced them undescribed. ” Under the description for Helminthoglypta traski (Newcomb), Henry A. Pilsbry (1939, Land Mollusca of North America (North of Mexico), Academy of Natural Sciences of Philadelphia, Monograph 3, Volume 1, Part 1, p. 174), made the following comments on this species: “Misled by a mistaken locality given by the collector, Junius Henderson described rather small specimens . . . as Sonorella betheli. In 1913 after visiting the Grand Canyon, where he thought that this shell was picked up, the late Prof. Ellsworth Bethel spent some time in Calfornia. His chief interest was botanical. Probably these shells were taken near Los Angeles; the type in University of Colorado Museum and a cotype, 109733 A.N.S.P., both of which I have seen, agree fully with traski from that vicinity.” ________________________________________________________________________________ 64 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Hershler, Robert; Liu, Hsiu-Ping; and Stevens, Lawrence E. 2016 A new springsnail (Hydrobiidae: Pyrgulopsis) from the lower Colorado River basin, northwestern Arizona. Western North American Naturalist, 76(1): 72-81. 157 Pyrgulopsis hualapaiensis, NEW SPECIES ; illustrated (including a color photo of the type locality, fig. 5) Mollusca: Gastropoda: Neotaenioglossa: Hydrobiidae (spring snails) “Types.—Holotype, USNM 1248611 (a dry shell), Upper Peach Springs, outflow just below concrete weir, Hualapai Indian Reservation, Mohave County, Arizona, 35.55786°N, 113.4314°W, 1280 m elevation, coll. RH and LS, 16 May 2014. Paratypes (from same lot), USNM 1266144 (approximately 250 dry shells and alcohol-preserved specimens).” “Referred material [non-type].—Arizona. Mohave County: topotypes, USNM 1247929 (dry shells), coll. LS, 28 May 2010; USNM 1248612 (dry shells and alcohol-preserved specimens), coll. LS, 9 January 2014; USNM 1259458 (dry shells and alcohol-preserved specimens), coll. LS, 24 July 2014.” ________________________________________________________________________________ Hester, J. Pinckney 1943 Two new Opuntias from the Grand Canyon region in Arizona. Cactus and Succulent Journal (Cactus and Succulent Society of America), 15: 191-194. Trachyophyta: Magnoliopsida: Caryophyllales: Cactaceae (cacti) 158 Opuntia hualpaensis, NEW SPECIES (pp. 191, 193, fig. 94, upper right [p. 192]) “Type [holotype]: Deposited in the Dudley Herbarium, Stanford University, No. 285575, collected along U. S.-Arizona Highway No. 66, 11 or 12 miles east of Peach Springs and 2 or 3 miles west of Hyde Park, Arizona, by J. Pinckney Hester, July, 1942.” [“Hyde Park”, near mile marker 117 on Historic Route 66, no longer exists. The gathering locality is near the present-day Grand Canyon Caverns tourist attraction.] 159 Opuntia abyssi, NEW SPECIES (pp. 193-194, fig. 94, upper left and lower [p. 192]) “Type [holotype]: Deposited in the Dudley Herbarium, Stanford University, No. 285624, collected in Peach Springs Canyon . . . by J. Pinckney Hester, in 1939.” ________________________________________________________________________________ 65 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Higgins, Larry C. 1974 Cryptantha atwoodii (Boraginaceae); a new species from Arizona. Southwestern Naturalist, 19(2) (July 26): 127-130. 160 Cryptantha atwoodii, NEW SPECIES ; illustrated Trachyophyta: Magnoliopsida: Boraginales: Boraginaceae (borage) “TYPE. Arizona: Coconino County, 7 mi N of junction of hwy 89 and 164 on hwy 89, 20 May 1970. N. D. Atwood 2624. Holotype deposited at BRY. Isotype at WTS.” “Apparently confined to the type locality near [T]he Gap . . . .” ________________________________________________________________________________ Hochstätter, Fritz 1995 The genera Pediocactus—Navajoa—Toumeya : revised : in the shadow of the Rocky Mountains. (Chris Holland, translator.) Mannheim, Germany: Fritz Hochstätter, 168 pp. [It is not determined here whether this title was originally published as this English translation, or if it was translated from another item thus far not identified.] 161 Sclerocactus whipplei (Englemann & Bigelow) Britton & Rose ssp. busekii, NEW SUBSPECIES (pp. 132-134) Trachyophyta: Magnoliopsida: Caryophyllales: Cactaceae (cacti) House Rock Valley, Arizona. ________________________________________________________________________________ Hodgson, Wendy C. 2001 Taxonomic novelties in American Agave (Agavaceae). Novon, 11(4) (Winter): 410-416. 162  Agave phillipsiana, NEW SPECIES (pp. 410-413, fig. 1 [“Drawn from Hodgson 8357B and Brian s.n.”]) Trachyophyta: Magnoliopsida: Asparagales: Asparagaceae (agaves) “TYPE: U.S.A. Arizona: Coconino County, Grand Canyon National Park, Clear Creek Canyon, ca [0].7 mi. upstream from camping area, 1140 m, 36°07.381′N, 112°00.568′W, 13 Sep. 1999, W. C. Hodgson 11861 (holotype, DES 44332; isotypes, GCNP, MO photo).” “Paratypes. U.S.A. Arizona: Coconino County, Grand Canyon National Park, Deer Creek Canyon, 36°23′52.2″N, 112°30′32″W, 731 m, 11 June 1994, Hodgson et al. 8357B (DES, GCNP, MO photo), 14 Aug. 1995, Hodgson 8357A and Brian (DES, GCNP), 26 Sep. 1995, Brian s.n. (ASU, DES, GCNP, MO); Grand Canyon National Park, Clear Creek Canyon, ca. 0.7-1 mi. upstream form camping area, 36°07.381′N, 112°00.568′W, 1140 m, 29 Apr. 1999, Hodgson et al. 11472 (DES, GCNP, MO photo); same area only 100 m further upstream, 37°[sic]07.592′N, 112°00.324′W, 1145 m, 29 Apr. 1999, Hodgson et al. 11473 (DES, GCNP, MO photo).” 66 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Note: The discovery of this species was first noted, without name, in Hodgson (1996, Field work and leg work; finding a new species, The Sonoran Quarterly (Desert Botanical Garden, Phoenix), 50(4) (December): 10-11). ________________________________________________________________________________ Holmgren, Arthur H., and Holmgren, Noel H. 1988 Euphorbia aaron-rossii (Euphorbiaceae), a new species from Marble and Grand Canyons of the Colorado River, Arizona. Brittonia, 40(4) (October/December): 357-362. 163  Euphorbia aaronrossii, NEW SPECIES; illustrated Trachyophyta: Magnoliopsida: Malpighiales: Euphorbiaceae (spurge) “Type: U.S.A. ARIZONA. Coconino Co.: Marble Canyon, along the Colorado River, 12 river miles below Lees Ferry, Salt Water Wash, among boulders, 930 m (3050 ft), 5 May 1971, A. Holmgren, N. Holmgren, P. Holmgren, D. Holmgren & A. Ross 15558 (HOLOTYPE: NY; ISOTYPES: ARIZ, ASU, BRY, F, TEX, UTC).” ________________________________________________________________________________ Holmgren, Noel H. 1973 Five new species of Castilleja (Scrophulariaceae) from the Intermountain Region. Bulletin of the Torrey Botanical Club, 100(2): 83-93. 164 Castilleja kaibabensis, NEW SPECIES (pp. 89-91, fig. 4 [p. 88]) Trachyophyta: Magnoliopsida: Lamiales: Scrophulariaceae (figworts) “Type: Arizona, Coconino County, Kaibab Plateau, V T Meadows, 6.5 km south of Kaibab Lodge, T.34 N., R.3 E., Sec. 7, elevation 2680 m, 7 August 1970, N. Holmgren & P. Holmgren 4682 (Holotype: NY; Isotypes: ARIZ, BRY, C, DAO, JEPS, KANU, NCU, RSA, TENN, US, UTC, WTU).” “Paratypes: All on the Kaibab Plateau, Coconino County, Arizona: Marble Flat, just north of North Rim Ranger Station, elevation 2500 m, 17 July 1941, Collom s.n. (US, GCNP [herbarium at Grand Canyon National Park Headquarters]); DeMotte Park, 7 August 1929, Mead 998 (GCNP); House Rock Canyon to Jacob Lake, elevation 2400 m, 14 August 1941, Kearney & Peebles 13648 (US); V T Ranger Station, elevation 2800 m, 12 July 1940, Collom s.n. (GCNP); V T Park, 30 km north of Bright Angel Point, 14 August 1938, Richards s.n. (NY); Ariz. Highway 67, 3.2 km north of Grand Canyon National Park boundary, T.34 N., R.3 E., Sec. 7, elevation 2650 m, 12 August 1966, Holmgren & Reveal 2995 (ARIZ, BRY, C, DAO, JEPS, KANU, NCU, NY, RSA, TENN, US, UTC, WTU); Lookout Canyon Road (422D), 5.6 km from (northwest of) Grand Canyon Highway (Ariz. 67), T.35 N., R.2 E., Sec. 15, elevation 2700 m, 6 August 1970, N. Holmgren & P. Holmgren 4668 (ARIZ, BRY, C, DAO, JEPS, KANU, NCU, NY, RSA, TENN, US, UTC, WTU).” ________________________________________________________________________________ 67 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Holmgren, Noel H., and Holmgren, Patricia K. 2002 New mentzelias (Loasaceae) from the Intermountain Region of western United States. Systematic Botany, 27(4) (October/December): 747-762. 165 Mentzelia memorabilis, new species (pp. 753-756; fig. 4) Trachyophyta: Magnoliopsida: Cornales: Losaceae (loasa) “TYPE: U.S.A. Arizona. Mohave Co., Uinkaret Plateau, just N of the Navajo Trail Road, 1.1 km W of the Clayhole Wash crossing, 27.5 km air distance SSW of Colorado City, 36°49′09″N, 113°11′21″W, T40N R8W S31, 1480 m, in partially barren gypsum-clay soil, 17 Sep 2001, N. H. Holmgren & P K. Holmgren 14580 (holotype: NY; isotypes: ASC, ASU, BRY, COLO, RM, RSA, UTC).” ________________________________________________________________________________ Hopkins, Heidi 2014 A review of the genus Arenivaga (Rehn) (Blattodea, Corydiidae), with descriptions of new species and key to the males of the genus. Sofia, Bulgaria; and Moscow, Russia: Pensoft Publishers, 256 pp. (ZooKeys, 384: 1-256 (Special Issue).) Arthropoda: Insecta: Blattodea: Polyphagidae (sand cockroaches) 166  Arenivaga grandiscanyonensis, NEW SPECIES (pp. 108-111, figs. 69, 70) [Note: “rm” indicates Colorado River Mile, by convention measured downstream from Lees Ferry, Arizona, with Right or Left bank indicated. Localities downstream from RM 164.7, on the left, could actually be within the boundaries of Hualapai Indian Reservation.] “Type locality. USA, Arizona, Mohave Co., Colorado River, Grand Canyon.” “Material examined. Holotype: ♂ in NAUF labeled ‘Mohave Co. AZ, Colorado R. GC, rm211.5R, 4/13/02, Coll. R.J.Delph, Ex: Light, Old High Water, blue label with “3”,’ ‘HOLOTYPE Arenivaga grandiscanyonensis Hopkins, 2012 [sic]’ [red label with black border].” [Square brackets are part of the quotation.] “Paratypes (2): USA: AZ, Coconino Co., Colorado River GC, 5/10/2001, J Rundall, rm160.5L, ex.light old high water, blue label ‘3’, 1 specimen—NAU 106 (2, NAUF). All paratypes labeled ‘Paratype Arenivaga grandiscanyonensis Hopkins 2012 [sic]’ [blue label with black border].” [Square brackets other than “sic” are part of the quotation.] Also 169 specimens examined (non-type). 167  Arenivaga impensa, NEW SPECIES (pp. 138-141, figs. 93, 94) (Holotype from Kingman, Arizona.) “Paratypes (14): . . . AZ, Indian Garden [Havasupai Gardens], Grand Canyon, 7/24/1934, 3800 ft., EL Bell & FE Lutz (2, AMNH); AZ, Coconino Co., Grand Canyon, 6/18/1954, M Cazier (1, AMNH) . . . . All paratypes labeled ‘Paratype Arenivaga impensa Hopkins 2012 [sic]’ [blue label with black border].” [Square brackets other than “sic” are part of the quotation.] 68 BIBLIOGRAPHY AND ANNOTATED CHECKLIST 168  Arenivaga pagana, NEW SPECIES (pp. 177, 182, figs. 120 [p. 180], 121 [p. 181]) “Type locality. USA, Arizona, Mohave Co., Colorado River, Grand Canyon.” “Material examined. Holotype: ♂ in NAUF labeled ‘Mohave Co. AZ, Colorado R. GC, blue label with “3”, rm202.5L, 9/4/03, Coll. R.J.Delph, Ex: Light, New High Water’ ‘HOLOTYPE Arenivaga pagana Hopkins, 2012 [sic]” [red label with black border].” [Square brackets other than “sic” are part of the quotation.] “Paratypes (9): USA: AZ, Mohave Co., Colorado River GC, 9/4/2003, RJ Delph, blue label with “3”, rm 202.5L, ex.light, new high water (2, NAUF); AZ, Mohave Co., Colorado River GC, 9/9/2001, J Rundall, blue label with “3”, rm 198.0R, ex.light, old high water (1, NAUF); AZ, Mohave Co., Colorado River GC, 9/9/2002, RJ Delph, blue label with “3”, rm 186.5L, ex.light, old high water, NAU 107 (1, NAUF); . . . AZ, Coconino Co., Colorado River GC, 9/1/2002, RJ Delph, blue label with “3”, rm37.3L, ex.light, old high water (1, NAUF); AZ, Coconino Co., Colorado River GC, 10/18/1982, 855 m, LE Stevens, blue label with “3”, M:53R, Nankowap [sic], sand dunes at night, Polyphagidae, Arenivaga det. D.Lightfoot (1, NAUF) . . . . All paratypes labeled ‘Paratype Arenivaga pagana Hopkins 2012 [sic]’ [blue label with black border].” [Square brackets other than “sic” are part of the quotation.] ________________________________________________________________________________ Horn, George H. 1893 The Galerucini of boreal America. Transactions of the American Entomological Society, 20(2) (April/June): 57-136. 169 Luperodes wickhami, NEW SPECIES (pp. 114-115); not illustrated Arthropoda: Insecta: Coleoptera: Chrysomelidae (leaf beetles) “Occurs in Arizona at Peach Springs (Wickham).” ________________________________________________________________________________ Hottes, F. C. 1956 Two new species of Cinara from northern Arizona with illustrations of hitherto unfigured species and notes on Schizolachnus flocculosa (Williams) (Aphidae). Proceedings of the Biological Society of Washington (Washington, D.C.), 69: 219-224. Arthropoda: Insecta: Hemiptera: Aphididae (aphids) 170  Cinara grande, NEW SPECIES (pp. 219-220; text-figs. on p. 223) “Host Abies concolor. Taken on the North Rim of the Grand Canyon, Arizona June 7, 1954. Collected by J. W. Bongberg. Holotype apterous viviparous female returned to the collection of The United States National Museum.” 69 BIBLIOGRAPHY AND ANNOTATED CHECKLIST 171 Cinara poketa, NEW SPECIES “Holotype apterous (pp. 220-221; text-figs. on p. 222) viviparous female. Host, Pinus edulis Aug. 24, 1956. Taken at point where road to Anita, Arizona branches from Highway 64 leading to Grand Canyon, Arizona.” [Locality is on the Kaibab National Forest, Tusayan Ranger District, ca. 35.861° N, 112.134° W.] ________________________________________________________________________________ Johnson, Walter N. 1990 A new subspecies of Cicindela pusilla Say from northern Arizona. Cicindela, 22(1) (March): 1-12. 172 Cicindela pusilla kaibabensis, NEW SUBSPECIES (description pp. 4-10, including right elytra of various subspecies illustrated pp. 8-9 [C. p. kaibabensis, figs. 16-21]) Arthropoda: Insecta: Coleoptera: Carabidae (ground beetles) “TYPE MATERIAL: Holotype (male) collected 8 miles north of Kaibab Lodge, Coconino Co., Arizona, on August 2, 1977, along with 11 male and 13 female paratypes, collected b y author (Walter N. Johnson). Allotype (female) collected 4 miles north of Kaibab Lodge, Coconino Co., Arizona, along with 4 male and 2 female paratypes on August 2, 1977, collected by author. Seven male and 2 female paratypes collected 8 miles north of Kaibab Lodge, Coconino Co., Arizona, on July 27, 1981, by author. Forty-eight male and 34 female paratypes collected 8 miles north of Kaibab Lodge, Coconino Co., Arizona, on July 22, 1987, by author. Thirty male and 14 female paratypes collected 8 miles north of Kaibab Lodge, Coconino Co., Arizona, on July 22, 1987, by Todd Lawton.” ________________________________________________________________________________ Johnston, M. Andrew 2019 Phylogenetic revision of the psammophilic Trogloderus Leconte (Coleoptera: Tenebrionidae), with biogeographic implications for the Intermountain Region. PeerJ, 7: e8039, doi:10.7717/peerj.8039, 45 pp. + Supplemental Information online. 173 Trogloderus skillmani, NEW SPECIES (pp. 24-26, figs. 6D [p. 16], 7F [p. 17]) Arthropoda: Insecta: Coleoptera: Tenebrionidae (darkling beetles) “Type material. Holotype: ‘USA: AZ: Mohave Co./6m E Colorado City/Rosy Canyon Road/1.5 m S UT state line/12-VII-2016/F.W. & S.A. Skillman,’ ‘ARTSYS0007053,’ bearing red holotype label. Deposited in the ASUHIC, catalog number ASUHIC0101565. Paratypes: A total of 920 specimens from the western regions of the Colorado Plateau around the Coral Pink Sand Dunes, Hurricane, and Toquerville Utah, bearing blue paratype labels (see Data S1 and S2 or SCAN for full specimen data).” The 920 paratypes are listed in the Supplemental Information online. ________________________________________________________________________________ 70 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Jones, Marcus E. 1895 Contributions to western botany. No. VII. Proceedings of the California Academy of Sciences, Series 2, 5: 611-732. 174 Astragalus humistratus var. tenerrimus, NEW VARIETY (p. 649); not illustrated Trachyophyta: Magnoliopsida: Fabales: Fabaceae (legumes) [Syntypes]: “No. 6052f. September 17, 1894, Buckskin Mountains, Arizona, 9000° [sic] alt., in gravel, under conifers. No. 6064. September 20, 1894, road to Nagle’s Ranch, Buckskin Mountains, Arizona, 9000° [sic] alt., in gravel. No. 6056gm. Same locality and date.” 175 Oenothera triloba var. ecristata, NEW VARIETY (pp. 681-682); not illustrated Trachyophyta: Magnoliopsida: Myrtales: Onagraceae (evening primroses) [Syntypes], all from Utah except for: “No. 6056ah. September 18, 1894, De Motte Park, Buckskin Mountains, Arizona, 9000° [sic] alt., in gravel.” 176 Bigelovia howardi var. attenuata, NEW VARIETY (pp. 691-692); not illustrated Trachyophyta: Magnoliopsida: Lamiales: Oleaceae (olives) [Syntypes], all from Utah except for: “No. 6052k. September 17, 1894, Buckskin Mountains, Arizona, 9000° [sic] alt., in gravel.” 177 Laphamia congesta, NEW SPECIES (p. 703); not illustrated Trachyophyta: Magnoliopsida: Asterales: Asteraceae (sunflowers) [Holotype]: “No. 6063. September 21, 1894, in clefts of rocks on the mesa below the Buckskin Mountains, Arizona, 7000° [sic] alt.” 178 Laphamia gracilis, NEW SPECIES (pp. 703-704) Trachyophyta: Magnoliopsida: Asterales: Asteraceae (sunflowers) [Holotype]: “No. 6050c. September 15, 1894, below Nagle’s Ranch, on edge of Buckskin Mountains, in crevices of limestone rocks, at 7000° [sic] alt., in very dry places.” _______________________________________________________________ 1902 Contributions to western botany. No. 10. Robinson, Utah: Mammoth Record Print, 90 pp.; [17], 16 plates. Trachyophyta: Magnoliopsida: Fabales: Fabaceae (legumes) 179 Astragalus kentrophyta var. coloradoensis, NEW VARIETY (p. 63, Pl. 1 [top row, 5th figure]) “Lee’s Ferry, Arizona, on the Colorado river, June 16, 1890.” 180 Astragalus kaibensis, NEW SPECIES (pp. 64-65, Pl. 2 [center row, 2nd figure]) “Gathered at House Rock, near Lee’s Ferry, northern Arizona, June 18, 1890, Jones.” ________________________________________________________________________________ 71 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Kavanaugh, David H. 2008 A new species of Nebria Latreille (Coleoptera: Carabidae: Nebriini) from the Grand Canyon, Arizona. Annals of the Carnegie Museum of Natural History, 77(1) (July): 1-5. 181  Nebria (Reductonebria) georgei, NEW SPECIES; illustrated Arthropoda: Insecta: Coleoptera: Carabidae (ground beetles) “Types.—HOLOTYPE, a male, deposited in MSBC, labeled: ‘USA AZ Coconino Co Grand Canyon Natl. Park, N36.102, W112.163 RM92.3L 06-07 May 2001 coll. Cobb, Brantley, Lightfoot’/ ‘shore zone pitfall trap’/ ‘COL CAR 001 006’/ ‘Nebria n. sp. ? det G.E. Ball 2004’/ ‘UNM-29’/ ‘HOLOTYPE Nebria georgei Kavanaugh n. sp. des. by D.H. Kavanaugh 2007 [sic]’ [red label]. One paratype, a female, deposited in CAS, labeled: “USA AZ Coconino Co Grand Canyon Natl. Park, N36.187, W113.107 RM180.8R 01-02 Sept 2003 coll. Cobb, Brantley, Lightfoot’/ ‘shore zone pitfall trap’/ ‘COL CAR 001 006’/ ‘UNM-30’/ ‘PARATYPE Nebria georgei Kavanaugh n. sp. des. by D. H. Kavanaugh 2007 [sic]’ [yellow label.” [Square brackets except for “sic” are part of the quotation.] “Type Locality.—Colorado River, Grand Canyon National Park, Coconino County, Arizona.” ________________________________________________________________________________ Kim, Ke Chung 1966 The species of Enderleinellus (Anoplura, Hoplopleuridae) parisitic on the Sciurini and Tamiasciurini. Journal of Parasitology, 52(5) (October): 988-1024. 182 Enderleinellus kaibabensis, NEW SPECIES (pp. 1000-1002, 1022; figs. 12 [p. 999, legend p. 998], 29 [p. 1001], 54-59 [p. 1005, legend p. 1004]) Arthropoda: Insecta: Psocodea: Enderleinellidae (sucking lice) “Type data: Holotype male, allotype female, and 6 paratypes (3 males and 3 females), ex Sciurus kaibabensis Merriam, Kaibab National Forest, Arizona, USA, Ferris Coll. 906 (USNM 168301), G.F.F. All type specimens except for 2 paratypes deposited in UCB; 1 male and 1 female paratype (on 1 slide) are deposited in UM.” [Inasmuch as the host is the endemic Kaibab squirrel, the type locality must be on the North Kaibab Ranger District.] ________________________________________________________________________________ Klauber, Laurence M. 1930 New and renamed subspecies of Crotalus confluentus Say, with remarks on related species. Transactions of the San Diego Society of Natural History, 6(3) (February 28): 95-144, Pls. 9-12. 183  Crotalus confluentus abyssus, NEW SUBSPECIES (pp. 114-117, Pl. 11, fig. 1 [legend p. 142]). (Common name given by the author, “Grand Canyon Rattlesnake”) Chordata: Reptilia: Squamata: Viparidae (rattlesnakes) “Type [holotype].—No. 2216 in collection of L.M.K. Captured alive on the Tanner Trail 300 ft. below the south rim of the Grand Canyon, Coconino County, Arizona; altitude approximately 7000 72 BIBLIOGRAPHY AND ANNOTATED CHECKLIST ft. Collected Sept. 15, 1929, by E. D. McKee.” The holotype is illustrated in Pl. 11, fig. 1. Subspecies also studied on two other living specimens and ten other specimens from the South Rim and the inner canyon [= paratypes]: “GCNP R37, Tonto Platform near Bright Angel Trail Alt. 4000 ft.; GCNP R38 [Coconino] Plateau near Navajo Point, Alt. 7000 ft.; GCNP R49 Phantom Ranch, north of Bright Angel Creek [sic] [seen as a skin]; GCNP R50 Desert View Point [specimen in poor condition]; GCNP R51 Desert View Point [specimen in poor condition]; LMK 2093 Roaring Springs Power Plant; LMK 2215 Roaring Springs Power Plant [specimen seen alive]; LMK 2272 Grand View Trail, 2000 ft. below South Rim [specimen seen alive]; USNM 32725 Grand Canyon [seen as a skin]; USNM 59747 Shimuno Creek [sic, Shinumo Creek] [seen as a skin]; USNM 78477 Bright Angel Canyon [seen as a skin]; USNM 78478 Burro Spring [North Rim; seen as a skin]. [Edwin D. McKee wrote of his 1929 sighting and capture of the holotype on Tanner Trail. (McKee, 1976, Discovery of the Grand Canyon Rattlesnake. Journal of Arizona History, 17(1) (Spring): 4749.)] ________________________________________________________________________________ Knight, Harry H. 1921 Monograph of the North American species of Deraeocoris (Heteroptera, Miridae). Minnesota State Entomologist, 18th Report to the Governor, pp. 77-210, Pl. 8. [Published June 18, 1921.] [Also published June 19, 1921 as University of Minnesota, Agricultural Experiment Station, Technical Bulletin 1, retaining original pagination, 77210.] Arthropoda: Insecta: Hemiptera: Miridae (jumping tree bugs) 184  Deraeocoris fulvus, NEW SPECIES (pp. 144-145; fig. 18) (Holotype and allotype from Williams, Arizona.) Paratypes include: “2 ♀ Aug 3 [1917], Grand View, Grand Canyon”. 185  Deraeocoris bullatus, NEW SPECIES (pp. 147-148; fig. 20) “Holotype: ♂ August 3 [1917], Grand View, Grand Canyon, Arizona (H. H. Knight); Cornell University collection. Allotype: taken with the type.” 186  Deraeocoris navajo, NEW SPECIES (pp. 155-156; fig. 24) “Holotype: ♂ August 3 [1917], Grand View, Grand Canyon, Arizona (H. H. Knight); Cornell University collection.” _______________________________________________________________ 73 BIBLIOGRAPHY AND ANNOTATED CHECKLIST 1925 Descriptions of thirty new species and two new genera of North American Miridae (Hemiptera). Bulletin of the Brooklyn Entomological Society, 20(1) (February): 33-58. Arthropoda: Insecta: Hemiptera: Miridae (jumping tree bugs) 187  Ceratocapsus clavicornis, NEW SPECIES (pp. 47-48) “Holotype: ♂ August 3, 1917, Grand View, Grand Canyon, Arizona (H. H. Knight); author’s collection. Allotype: same data as the type. Paratypes: 30 ♂♀, taken with the types on Cowania mexicana.” 188  Phytocoris mellarius, NEW SPECIES (pp. 56-57) “Holotype: ♂ August 3, 1917, Grand View, Grand Canyon, Arizona (H. H. Knight); author’s collection. Allotype: same data as the type. Paratypes: 3♂ 6♀, taken with the types. This species was probably collected on some conifer although I do not find the remarks in my notes.” _______________________________________________________________ 1926 Descriptions of seven new species of Pilophorus (Hemiptera, Miridae). Bulletin of the Brooklyn Entomological Society, 21(1/2) (February/April): 18-26. 189  Pilophorus fuscipennis, NEW SPECIES (pp. 23-24); not illustrated Arthropoda: Insecta: Hemiptera: Miridae (jumping tree bugs) (Holotype and allotype from near Trinidad, Colorado.) “Paratypes: . . . ♂ ♀, Aug. 3, 1917, Grand View, Grand Canyon, Arizona (H. H. Knight). Paratypes in collections of Iowa State college and Colorado Agricultural College.” _______________________________________________________________ 1928 New species of Phytocoris from North America (Hemiptera, Miridae). Bulletin of the Brooklyn Entomological Society, 23(1) (February): 28-46. 190  Phytocoris hesperius, NEW SPECIES (pp. 44-45); not illustrated Arthropoda: Insecta: Hemiptera: Miridae (jumping tree bugs) (Holotype from near Trinidad, Colorado; allotype from Chiricahua Mountains, Arizona.) P aratype specimens include “2♂ 1♀ Aug. 2, 1917, at top of Bright Angel trail, Grand Canyon (H. H. Knight) . . . .” _______________________________________________________________ 1929 Descriptions of five new species of Plagiognathus from North America (Hemip.: Miridae). Entomological News (Academy of Natural Sciences of Philadelphia, Entomological Section), 40 (March): 69-74. 191  Plagiognathus tenellus, NEW SPECIES (p. 73); not illustrated Arthropoda: Insecta: Hemiptera: Miridae (jumping tree bugs) “Holotype: ♂ August 2, 1917, top of Bright Angel trail, Grand Canyon, ARIZONA (H. H. Knight); ♂ ♀, taken with the types . . . .” author's collection. Allotype: taken with the type. Paratypes: 12 _______________________________________________________________ 74 BIBLIOGRAPHY AND ANNOTATED CHECKLIST 1934 Phytocoris Fallen—twelve new species from the western United States (Hemiptera, Miridae). Bulletin of the Brooklyn Entomological Society, 29(1) (February): 1-16. 192  Phytocoris varius, NEW SPECIES (pp. 9-11); not illustrated Arthropoda: Insecta: Hemiptera: Miridae (jumping tree bugs) ♂ September 6, 1931, Grand Canyon (H. H. Knight); author’s collection. Allotype: ♀, taken with the type. Paratypes: ♂, 3♀, taken with the types by beating on large cedar trees “Holotype: (Juniperus sp.) which were found growing behind the cabin camp located at the entrance gate of the Grand Canyon National Park.” (Additional paratypes from Durango, Colorado and Chiricahua Mountains, Arizona.) _______________________________________________________________ 1968 Taxonomic review: Miridae of the Nevada Test Site and the western United States. Brigham Young University Science Bulletin (Biological Series), 9(3), 282 pp. Arthropoda: Insecta: Hemiptera: Miridae (jumping tree bugs) 193  Parthenicus cowaniae, NEW SPECIES (pp. 148-149, fig. 211 [p. 147]) ♂ Aug. 3, 1917, Grand View, Grand Canyon, Arizona (H. H. Knight), taken on cliff rose, Cowania stansburiana, which is the host plant of the species. Allotype: ♀ taken with the type. Paratypes: “26 ♂ 20 ♀ taken with the types on cliff rose . . . ♂ ♀ Sept. 6, 1931, Grand View, Grand Canyon (H. H. Knight) . . . ♂ ♀ Aug. 1, 1914, Hermit Rim road, Grand Canyon, “Holotype: Arizona (L. C. Bradley).” 194 Bolteria juniperi, NEW SPECIES (pp. 202-203, fig. 253) (Holotype, allotype, and paratypes from Scipio, Utah.) Other paratypes include “. . . ♂ June 16, alt 8000 ft, near Grand Canyon, Arizona.” [Given the altitude data for this paratype, and the fact that the primary types are from Utah, it is presumably on the northern side of the canyon.] 195  Phytocoris flaviatus, NEW SPECIES (p. 241, fig. 297 [p. 242] “Holotype: ♂ Sept. 6, 1931, above Bright Angel Trail, Grand Canyon, Arizona (H. H. Knight).” [No other type material.] ________________________________________________________________________________ Knull, Josef N. 1957 Three new species of Aphricus with a note on Ctenicera (Coleoptera: Elateridae). Ohio Journal of Science, 57(4) (July): 200-202. 196 Aphricus knowltoni, NEW SPECIES (pp. 200-201); not illustrated Arthropoda: Insecta: Coleoptera: Elateridae (click beetles) (Holotype and paratypes from Washington, Utah.) Another paratype from “Cameron, Arizona, Aug. 19, 1950, D. J. and J. N. Knull, all in collection of author.” ________________________________________________________________________________ 75 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Kruidenier, Francis J., and Mehra, Krishna N. 1958 Wellcomia perognathi n. sp. (Nematoda: Oxyuridae) from pocket mice, Perognathus intermedius (Merriam, 1889) of the Grand Canyon, Arizona. Transactions of the Illinois State Academy of Science, 51(3/4) (July 10): 20-25. 197  Wellcomia perognathi, NEW SPECIES; illustrated Nematoda: Chromadorea: Ascaridida: Oxyuridae (pinworms) “Host: Perognathus intermedius (Merriam, 1889). Habitat: Intestine (cecum). Locality: Coconino County, Arizona (Grand Canyon National Park). Holotypes [sic]: U.S. Nat’l. Mus. Helm. Coll. No. 38306.” [Limited data, thus. Collection made in 1954; see note with no. 198, below.] _______________________________________________________________ 1959 Aspiculuris ackerti, n. sp, (Nematoda: Oxyruidae) from the wood rats of Arizona. Proceedings of the Helminthological Society, 26(2) (July): 147-150. 198  Aspiculuris ackerti, NEW SPECIES; illustrated Nematoda: Chromadorea: Ascaridida: Oxyuridae (pinworms) “Hosts: Neotoma albigula—U. of Ill. Museum #11650 [and] N. cinerea—U. of Ill. Museum #11688. Habitat: Intestine. Locality: Coconino County, Arizona. Holotype: U.S. Nat’l Mus. Helm. Coll. #56163.” “. . . recovered from a series of neotomid rats examined during a 1954 expedition to the Grand Canyon, Coconimo [sic] County, Arizona (see Kruidenier and Peebles, 1958).” The cited item does not appear in the Literature Cited section of this paper, but is identified (here) as the citation with no. 199, below, which there provides locality data only as “Grand Canyon National Park”. ________________________________________________________________________________ Kruidenier, Francis J., and Peebles, C. R. 1958 Gongylonema of rodents: G. neoplasticum (redefinition); G. dipodomysis n. sp.; and G. peromyski n. sp. Transactions of the American Microscopical Society, 77(3) (July): 307315. 199  Gongylonema peromysci, NEW SPECIES (pp. 310-312, Pl. 2, figs. 1-6 [p. 313]) Nematoda: Chromadorea: Spiurida: Gongylonematidae (spiurid gullet worms) “Host: Type: Peromyscus maniculatus rufinus (Merriam, 1890) Osgood, 1909. Univ. Ill. Mus. No. 10832 . . . . Habitat: Mucosa of cardiac stomach. Coconimo [sic] County, Arizona. Locality: Grand Canyon National Park, Holotypes [sic]: U.S. National Museum Helminthological Collection No. 38202.” [Collection made in 1954; see note with no. 198, above.] ________________________________________________________________________________ 76 BIBLIOGRAPHY AND ANNOTATED CHECKLIST LaBerge, Wallace E. 1967 A revision of the bees of the genus Andrena of the western hemisphere. Part I. Callandrena. (Hymenoptera: Andrenidae). Bulletin of the University of Nebraska State Museum, 7 (October): 1-318. 200  Andrena (Callandrena) utahensis, NEW SPECIES (pp. 258-260, figs. 292-296 [p. 314]) Arthropoda: Insecta: Hymenoptera: Andrenidae (andrenid bees) (Holotype and allotype from Duchesne, Utah.) Grand Canyon. 5 Additional paratypes include: “ARIZONA: . . . ♀♀, 1 ♂, June 5, 1940, G. E. Bohart . . . .” ________________________________________________________________________________ Levine, Norman D.; Ivens, Virginia; and Kruidenier, Francis J. 1957a New species of Eimeria from Arizona rodents. Journal of Protozoology, 4(2) (May): 8088. Apicomplexa: Conoidasida : Eucoccidiorida: Eimeriidae (coccid parasites) 201  Eimeria tamiasciuri, NEW SPECIES (pp. 80-81, fig. 1 [p. 82]) “Host: Tamiasciurus hudsonicus mogollonensis (U. of Ill. Mus. #10,432) (red or spruce squirrel). Location: Intestinal contents. Locality: The host animal was trapped by Mr. Woodrow Goodpaster on Aug. 7, 1954 on Road W-3 on the north rim of Grand Canyon near the entrance to Grand Canyon National Park, Arizona.” 202  Eimeria cutamiae, NEW SPECIES (pp. 82-83, fig. 3) “Host: Eutamias dorsalis dorsalis (U. of Ill. Mus. #10,386) (cliff chipmunk). Location: Intestinal contents. Locality: This species was found in one of two Eutamias d. dorsalis examined in the Grand Canyon National Park, Arizona. The host chipmunk was trapped by Mr. K. R. Nelson on July 4, 1954 at Hull Tank, on the south rim of Grand Canyon.” 203  Eimeria thomomysis, NEW SPECIES (p. 83, fig. 4 [p. 82]) “Host: Thomomys bottae fulvus (U. of Ill. Mus. #10,568 and #7990) (pocket gopher). Location: Intestinal contents. Locality: This species was found in two out of five T. bottae examined in Arizona. One animal was trapped by Mr. Donald Hoffmeister on July 4, 1954 at Hearst Ranch [Grandview] on the south rim of Grand Canyon in Grand Canyon National Park.” 204  Eimeria perognathi, NEW SPECIES (pp. 83-84, fig. 5 [p. 82]) “Host: Perognathus intermedius subsp. (U. of Ill. Mus. #10,640) (rock pocket mouse). Location: Intestinal contents. Locality: This species was found in one out of two P. intermedius examined from the Grand Canyon National Park, Arizona. The host animal was trapped by Dr. Donald F. Hoffmeister on July 23, 1954 along the Plateau Point Trail one mile below [sic] Indian Garden [Havasupai Gardens] inside Grand Canyon on the south side of the Colorado River.” 77 BIBLIOGRAPHY AND ANNOTATED CHECKLIST 205  Eimeria albigulae, NEW SPECIES (p. 84, fig. 6 [p. 85]) “Host: Neotoma albigula subsp. (U. of Ill. Mus. #11,650) (white-throated woodrat). Location: Intestinal contents. Locality: This species was found in a N. albigula in Grand Canyon National Park, Arizona. The host woodrat was collected by Mr. K. R. Nelson on July 11, 1954 on the east side of Cedar Mountain, on the south rim of Grand Canyon [sic].” [The precise collecting locality may be within the boundaries of the Navajo Indian Reservation.] 206  Eimeria operculata, NEW SPECIES (pp. 84-85); not illustrated “Host: Neotoma stephensi stephensi (U. of Ill. Mus. #11,636) (Stephens’ woodrat). Location: Intestinal contents. Locality: This species was found in one out of two N. s. stephensi examined in Grand Canyon National Park, Arizona. The host woodrat was collected by Mr. K. R. Nelson on July 5, 1954 at Zuni Point, on the south rim of Grand Canyon.” 207  Eimeria peromysci, NEW SPECIES (pp. 85-86, fig. 7) “Host: Peromyscus truei truei (U. of Ill. Mus. #11,249) (piñon mouse). Location: Intestinal contents. Locality: This species was found in one out of four P. t. truei examined in Grand Canyon National Park, Arizona. The host mouse was collected by Dr. Donald F. Hoffmeister on July 5, 1954 on the west side of Zuni Point, which is on the south rim of Grand Canyon.” 208  Eimeria arizonensis, NEW SPECIES (p. 86, fig. 8 [p. 85]) “Host: Peromyscus truei truei (U. of Ill. Mus. #11,220 and 11,249) (piñon mouse). Location: Intestinal contents. Locality: This species was found in two of four P. t. truei examined in Grand Canyon National Park, Arizona. The host mouse form which the larger series of oocysts was obtained was collected by Dr. Donald F. Hoffmeister on August 5, 1954 at Cape Royal on the north rim of Grand Canyon. The host mouse from which the smaller series of oocysts was obtained was collected by Dr. Hoffmeister on July 5, 1954 on the west side of Zuni Point, which is on the south rim of Grand Canyon. This same individual was infected with E. peromysci [see no. 207, above].” 209  Eimeria eremici, NEW SPECIES (pp. 86-87, fig. 9 [p. 85]) “Host: Peromyscus eremicus eremicus (U. of Ill. Mus. #11,354) (cactus mouse). Location: Intestinal contents. Locality: This species was found in one out of two P. eremiscus Grand Canyon National Park, Arizona. The host mouse was collected by Dr. Donald F. Hoffmeister on July 23, 1954 on the Tonto Plateau along the Plateau Point Trail one mile below [sic] Indian Garden [Havasupai Gardens]; this location is inside the Grand Canyon, on the south side of the Colorado River.” 210  Eimeria onychomysis, NEW SPECIES (p. 87, fig. 10 [p. 85]) “Host: Onychomys leucogster subsp. (U. of Ill. Mus. #10,666) (northern grasshopper mouse). Location: Intestinal contents. Locality: This species was found in one out of two O. leucogaster examined in Grand Canyon National Park, Arizona. The host mouse was collected by Dr. Donald F. Hoffmeister on July 19, 1954 along State Highway 64 at the southeast boundary of the park, south of the canyon.” _______________________________________________________________ 78 BIBLIOGRAPHY AND ANNOTATED CHECKLIST 1957b Isopora citolli n. sp. from the rock squirrel, Citellus variegatus utah. Journal of Protozoology, 4(3) (August): 143-144. 211  Isopora citolli, NEW SPECIES (p. 87, fig. 1) Apicomplexa: Conoidasida : Eucoccidiorida: Eimeriidae (coccid parasites) “The host rock squirrel was trapped by Dr. Hoffmeister on August 9, 1954 one-half mile from Swamp Point, near Powell Saddle, on the north rim of Grand Canyon at an elevation of 7500 feet. Its skin and skull were preserved by Dr. Hoffmeister, and are in the University of Illinois Natural History Museum (Museum No. 10220).” ________________________________________________________________________________ Lindsay, Dale R. 1938 New species of Norvellina (Homoptera, Cicadellidae). Journal of the Kansas Entomological Society, 11(4) (October): 113-123. 212  Norvellina bicolorata (Ball) var. inflata, NEW VARIETY (pp. 113-114); not illustrated Arthropoda: Insecta: Hemiptera: Cicadellidae (leafhoppers) (Holotype and allotype from White Sands, New Mexico.) “Numerous paratypes from the following localities: Grand Canyon, Ariz. . . . . Types and paratypes in the Snow Entomological Collection. Paratypes in collection of Dr. Ball and U. S. N. M.” ________________________________________________________________________________ Liu, Tong-Xian, and Kosztarab, Michael 1987 Two new species of Chionaspis (Homoptera: Coccoidea: Diaspididae) from North America. Florida Entomologist, 70(4) (December): 512-520. 213  Chionaspis gilli, NEW SPECIES (pp. 516-519, fig. 2) Arthropoda: Insecta: Hemiptera: Diaspididae (armored scales) (Holotype from Las Cruces, New Mexico.) “Paratypes: On Tamarix chinensis, 5 on 4 slides, Grand Canyon National Park, Colorado River, Coconino Co., Arizona (AZ), VII-23-1984 collected by L. E. Stevens, deposited in USNM . . . .” ________________________________________________________________________________ Marshall, William B. 1929 Three new land shells of the genus Oreohelix from Arizona. Proceedings of the U.S. National Museum, 76(2802), 3 pp. 214 Oreohelix yavapai vauxae, NEW SUBSPECIES (pp. 1-2, Plate1, figs. 1-3, 11). Mollusca: Gastropoda: Stylommatophora: Oreohelicidae (mountain snails) “The study of a collection of Oreohelix made by Mrs. Mary Vaux Walcott in the canyon at Supai, Coconino County, Ariz., in 1928, and presented by her to the United States National Museum, not 79 BIBLIOGRAPHY AND ANNOTATED CHECKLIST only proved that they belonged to a new subspecies, but their examination entailed a close scrutiny of forms long since contained in our collection but not previously adequately studied.” “The type [holotype] and 14 paratypes come from the Canyon at Supai, Coconino County, Ariz., and were collected and presented by Mrs. Charles D. Walcott, whose maiden name has been bestowed upon it.” A table provides measurements for the “type and those of the paratypes which are adult”, giving the USNM catalog numbers: 380687 (type), 380688 (eight adult paratypes in one lot; presumably the sub-adult paratypes are in the same lot). ________________________________________________________________________________ Massey, Calvin L. 1974 Biology and taxonomy of nematode parasites and associates of bark beetles in the United States. U.S. Forest Service, Agriculture Handbook 446, 233 pp. 215  Parasitorhabditis gracilis, NEW SPECIES (p. 69, fig. 41 [p. 70]) Nematoda: Chromadorea: Rhabditida : Rhabditidae (rhabditid roundworms) “Type habitat.—Associated with Pseudohylesinus grandis Sw. in white fir. Type locality.—Grand Canyon, Arizona. Type specimens.—Collection No. 84-J.” ________________________________________________________________________________ McAtee, W. L. [Waldo Lee] 1919 Notes on Nearctic Heteroptera. Bulletin of the Brooklyn Entomological Society, 14(1) (February): 8-15. 216  Chelinidea vittiger Uhler var. artuflava, NEW VARIETY (pp. 11-12); not illustrated Arthropoda: Insecta: Hemiptera: Coreidae (leaf-footed bugs) Syntypes include “Grand Cañon, Ariz., July 10 (U. S. N. M.)”. [No further data about this specimen.] ________________________________________________________________________________ McKelvey, Susan Delano 1949 A new Agave from Arizona. Journal of the Arnold Arboretum, 30(3) (July): 227-230, 2 plates. 217  Agave kaibabensis, NEW SPECIES (pp. 227-228, Pl. 1) “ARIZONA, Coconino County, Kaibab Plateau on the north side of the Grand Canyon of the Colorado River, McKelvey 4381 (Arnold Arboretum, TYPE [holotype]), 4381A (Arnold Arboretum) [paratype], May 15, 1934.” [Apparently in Grand Canyon National Park. See notes below, which suggest the author’s casualness with geographical and records concerning the gathering.] 80 BIBLIOGRAPHY AND ANNOTATED CHECKLIST McKelvey is not specific about the locality of the type, although she offers somewhat confusing additional data in her text from which one may infer that the type is from Grand Canyon National Park and not from the Kaibab National Forest north of the park. “On May 15, 1934, while visiting the Kaibab Plateau on the northern side of the Grand Canyon of the Colorado, Coconino County, Arizona, the writer first saw the Agave here described. Two plants had been collected by the Park authorities at some previous date, but their leaf-rosettes and long fruiting stalks were still intact. It was not possible to make the descent into the gorge, but a few scattered plants were discernible some distance below the rim. Flowers were promised at a later date but were never forthcoming, unfortunately. [¶]The two plants are shown in the included photograph (McKelvey 152-12); although one was considerably smaller than the other it was undoubtedly the same species. The dimensions are from the larger plant.” (p. 227). “The smaller plant of the picture [which is posed (!?) beside the type in Pl. I] is McKelvey 4381A. The plant, as noted [p. 227?], came originally from some distance below the rim on the north wall of the Canyon. The date of actual collection is not known; it must have been in the season of 1933; the writer made her specimens on May 15, 1934. Her photographs 152-11, 12, and 1531, 2 are of these plants; reproduced here is 153-2 (Pl. I)” (p. 229). Note that on p. 227 McKelvey indicates that the photograph of the two plants is her photo 152-12. [Pl. II of this paper depicts a comparative specimen of Agave utahensis Engelmann, gathered by McKelvey in 1929 “some 13 miles east of Peach Springs, Mohave County, Arizona, in the extreme northwestern corner of Yavapai County.”] ________________________________________________________________________________ Merriam, C. Hart [Clinton Hart] 1890 Results of a biological survey of the San Francisco Mountain region and the desert of the Little Colorado River, Arizona. U.S. Department of Agriculture, North American Fauna, (3), 136 pp., 13 plates, 5 maps. Regarding the localities noted at Grand Canyon, Merriam describes the trip to the canyon thusly (p. 4, in its entirety): “TRIP TO THE GRAND CAÑON OF THE COLORADO, SEPTEMBER 9 TO 16, INCLUSIVE. The usual road was followed from Little Spring to Hull Spring and Red Horse Tank, and thence to the tank known as Cañon Spring on the Cocanini [sic] Plateau, close to the cañon, which is here about 1,800 meters (6,000 feet) in depth. Mr. Bailey and myself climbed down into the cañon and remained in it two days and two nights.” This information is enough to identify the locale as John Hance’s accommodations at the head of the Old Hance Trail, down which Merriam and Bailey descended. 218  Hesperomys megalotis, NEW SPECIES (pp. 63-64, Pl. III, figs. 1-4; Pl. IV) [holotype illustrated] Chordata: Mammalia: Rodentia: Crecitidae (deer mice) (Type [holotype] from Black Tank, Desert of the Little Colorado River.) Paratype specimens include USNM 17930/24841 (original no. 469), “Grand Cañon, Arizona, Sept. 11, 1889, 81 ♀ ad.” BIBLIOGRAPHY AND ANNOTATED CHECKLIST 219  Spilogale gracilis, NEW SPECIES (pp. 83-84); not illustrated. (Common name given by the author, “Little Striped Skunk”) Chordata: Mammalia: Carnivora: Mephitidae (skunks) “Type [holotype] No. 17986/24897 ♂ ad. U. S. National Museum (Department of Agriculture collection). From Grand Cañon of the Colorado (altitude 3,500 feet), Arizona, north of San Francisco Mountain, September 12, 1889. Collected by C. Hart Merriam (Original number 451).” Also lists USNM 18598/25368 “♂ im.” [paratype] also from Grand Canyon. Merriam remarked (p. 83): “While asleep near a small spring in the Grand Canon of the Colorado, September 12, 1889, I was awakened at midnight by a sniffling noise about my head. Rising suddenly on my elbow, a small animal scampered hurriedly away over the rocks. His form was only dimly outlined in the dark, but a hasty shot left no doubt as to his identity, and a moment later I held in my hand the type of a new species of Little Striped Skunk. A day or two afterward a younger individual was captured among the cliffs at the top of the Cañon. The stomachs of both contained remains of the cliff mouse Hesperomys erenucus).” _______________________________________________________________ 1904 Two new squirrels of the aberti group. Proceedings of the Biological Society of Washington (Washington, D.C.) 17: 129-130. 220  Sciurus kaibabensis, NEW SPECIES (pp. 129-130); not illustrated. Chordata: Mammalia: Rodentia: Sciuridae (squirrels) “Type [holotype] from head of Bright Angel Creek, top of Kaibab Plateau, north side of Grand Canyon of Colorado [sic], Arizona. Adult male, No. 130,932, U. S. National Museum, Biological Survey Collection. December 1, 1903. John T. Stewart.” [This is the widely known “Kaibab Squirrel”.] ________________________________________________________________________________ Miller, Robert Rush 1946 Gila cypha, a remarkable new species of cyprinid fish from the Colorado River in Grand Canyon, Arizona. Journal of the Washington Academy of Sciences (Washington, D.C.), 36(12): 409-415. 221  Gila cypha, NEW SPECIES; illustrated Chordata: Teleostei: Cypriniformes: Cyprinidae (carps) “Holotype.—The holotype (U.S.N.M. no. 131389) is a specimen about 305 mm in standard length and was taken by N. N. Dodge near Phantom Ranch in the western end of Grand Canyon National Park [old boundaries], Arizona. It was caught in swift water on hook and line, presumably in the nearby Colorado River at or near the mouth of Bright Angel Creek.” [This is the widely known “Humpback Chub”.] ________________________________________________________________________________ 82 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Miller, Walter B. 1984 Three new species of Sonorella (Gastropoda: Pulmonata: Helminthoglyptidae) from Arizona. Proceedings of the Biological Society of Washington (Washington, D.C.), 97(4): 681-687. 222  Sonorella reederi, NEW SPECIES (pp. 681-684, figs. 1A-C, 2) Mollusca: Gastropoda: Stylommatophora: Helminthoglyptidae (shoulderbands) “Disposition of types.—Holotype: USNM 792406. Paratypes: ANSP 356004; CAS 033405; FMNH 206235; UTEP 9051; RLR 298; WBM 5241 & 6307.” “Type-locality.—Mohave Co., Arizona. Lower Granite Gorge of the Colorado River, in limestone rockslide just west and below Rampart Cave; elevation ca. 1700 ft., 36°06′N, 113°56′W.” ________________________________________________________________________________ Moldenke, Andrew R. 1970 A revision of the Clytrinae of North America north of the Isthmus of Panama (Coleoptera: Chrysomelidae). Stanford, California: Stanford University, 310 pp. 223 Saxinis saucia kaibabiae, NEW SUBSPECIES (pp. 182- ; illustrated) Arthropoda: Insecta: Coleoptera: Chrysomelidae (leaf beetles) Jacob Lake, Kaibab Forest. ________________________________________________________________________________ Monod, M. [Michel] 1983 Monographie taxonomique des Gnomoniaceae (ascomycès de l’ordre des Diaporthales). I. Beihefte zur Sydowia (Horn, Niederösterreich, Austria), (Annales Mycologici, Ser. II), IX. Beiheft, 315 pp. [This publication originally was Monod’s doctoral dissertation, Université de Lausanne”.] [Monograph is in French; species descriptions are in botanical Latin and in French.] 224  Gnomonia quercusgambellii, NEW SPECIES (pp. 98-99, fig. 13.5 [p. 278]) Ascomycota (sac fungi): Sordariomycetes: Diaporthales: Valsaceae “. . . sous Gnomonia fasciculata: Quercus gambellii: North Rim of Grand Canyon, Arizona, 18 août 1973, Barr, 6095 (MASS, Holotype).” ________________________________________________________________________________ 83 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Muchmore, William B. 1981 Cavernicolous species of Larca, Archeolarca, and Pseudogarypus with notes on the genera, (Pseudoscorpionida, Gaypidae and Pseudogarypidae). Journal of Arachnology, 9(1): 47-60. 225  Archeolarca cavicola, new species (pp. 55-56, figs. 11, 12 [p. 53]) Arthropoda: Euchelicerata: Pseudoscorpiones (pseudoscorpions): Larcidae “Holotype female (WM 5398.01001) from Cave of the Domes [Horseshoe Mesa], Grand Canyon National Park, Coconino County, Arizona, 15 April 1978 (W. Calvin Welbourn). The type is in the Florida State Collection of Arthropods, Gainesville.” ________________________________________________________________________________ Nabokov, Vladimir 1942 Some new or little known Nearctic Neonympha (Lepidoptera: Satyridae). Psyche, 49(3/4) (September/December): 61-80. Neonympha dorothea NEW SPECIES (pp. 62-63) 226  Neonympha dorothea dorothea, NEW SUBSPECIES (pp. 64-66); not illustrated Arthropoda: Insecta: Lepidoptera: Nymphalidae (admirals) “Male, holotype, female, allotype, and two males, paratypes, placed in the American Museum of Natural History. Taken during a brief visit to Grand Cañon, Ariz., South Rim, on June 9th, 1941 (bright cold morning after snow and rain). They were weakly fluttering beside the trail together with a few Coen[onympha]. tullia furcae almost no other butterflies about. Named in honor of Miss Dorothy Leuthold who kindly kicked up the first specimen. Female, paratype, labelled ‘Grand Cy., June 11th ’30’, ex Coll. of C.F. dos Passos, Am. Mus. Nat. Hist.” [Two additional subspecies of N. dorothea also named therein, which are not pertinent to the present checklist.] ________________________________________________________________________________ Nelson, Edward W., and Goldman, Edward A. 1931 Three new pumas. Journal of the Washington Academy of Sciences (Washington, D.C.), 21(10) (May 19): 209-212. 227  Felis concolor kaibabensis, NEW SUBSPECIES (pp. 209-210); not illustrated. (Common name given by the author, “Kaibab Mountain Lion”) Chordata: Mammalia: Carnivora: Felidae (cats) “Type [holotype].—From Powell Plateau, Grand Canyon National Park, Arizona (altitude 8,700 feet). No. 171186 ♂ adult, U. S. National Museum (Biological Survey collection), collected by J. T. Owens, April 15, 1911. X number 8432.” ________________________________________________________________________________ 84 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Nelson, G. H., and Westcott, R. L. 1995 Three new species of Acmaeodera Eschscholtz (Coleoptera: Buprestidae) from the United States and Mexico. The Coleopterists Bulletin, 49(1): 77-87. 228 Acmaeodera navajo, NEW SPECIES (pp. 82-84; figs. 2 [p. 78], 7 [p. 80]) Arthropoda: Insecta: Coleoptera: Buprestidae (jewel beetles) “The holotype ♀ (USNM) and 90 paratypes, Arizona: Coconino Co., 9 mi NW Page, 3-VI-70, R. L. Westcott, on either purple-flowered Opuntia or flowers Sphaeralcea. Other paratypes are as follows: . . . Arizona: Coconino Co., Vermillion [sic] Cliffs [Vermilion Cliffs], 9-VII-67, J. H. & J. M. Davidson, M. Cazier (1).” Non-type specimens examined include “Navajo Indian Reservation, Hwy. 89, 15 mi S Page, 26 VI-80, L. L. Lampert, on Linum lewisii; 8 mi. SW Cliff Dwellers Lodge, 11-VII-67, M. A. Cazier, J. H. and J. M. Davidson, on Sphaeralcea sp. . . . .” [It is unclear whether “Vermillion Cliffs” refers to the geographical feature or to the place name, Vermilion Cliffs, although note that the authors also specially mention nearby Cliff Dwellers Lodge.] _____________________________________________________________________________ ___ Oberholser, Harry C. 1937 Descriptions of three new screech owls from the United States. Journal of the Washington Academy of Sciences (Washington, D.C.), 27(8) (August 15): 354-357. 229  Otus asio mychophilus, NEW SUBSPECIES (pp. 356-357); not illustrated. (Common name given by the author, “Grand Canyon Screech Owl”) Chordata: Aves: Strigiformes: Strigidae (typical owls) “Type [holotype].—Adult female, No. 340593, U. S. National Museum, Biological Survey collection; south rim of Grand Canyon, 6,900 feet altitude, Grand Canyon Village, Arizona; January 28, 1935; Russell K. Grater, original number, 23.” ________________________________________________________________________________ Pape, Robert B.; Thomas, Donald B.; and Aalbu, Rolf L. 2007 A revision of the genus Eschatomoxys Blaisdell (Tenebrionidae: Pimeliinae: Edrotini) with notes on the biology. The Coleopterists Bulletin, 61(4): 519-540. 230  Eschatomoxys pholeter Thomas & Pape, NEW SPECIES (pp. 525-527, figs. 1 [p. 521], 5 [p. 523], 8A,B [p. 526]) Arthropoda: Insecta: Coleoptera: Tenebrionidae (darkling beetles) “Holotype. Male. ARIZONA: Mohave Co., Grand Canyon N.P., Bat Cave, 29-III-96, R.B. Pape coll. Allotype. Female. Labeled same data as holotype. Paratypes: Three males, one female, labeled same data as holotype; Four males, six females: AZ. Mohave Co., Bat Cave, Gr. Canyon, GRTCA Cave SCI-0043, 26-X-01, R.B. Pape (deposited CAS and DB Thomas collection). One female: AZ. Mohave Co., Eldel Cave [Grand Canyon–Parashant National Monument], twilight, IX- 85 BIBLIOGRAPHY AND ANNOTATED CHECKLIST 23-05, J. Judson Wynne col[l]. Three females: ARIZONA: Mohave Co., Grand Canyon N.P., Rampart Cave, teneb on 7,200 yr old sloth shit [sic!].” ________________________________________________________________________________ Parfitt, Bruce D. 1997 Opuntia pinkavae (Cactaceae), a new species from Arizona and Utah. Rhodora (Journal of the New England Botanical Club), 99(899) (Summer): 223-228. 231  Opuntia pinkavae, NEW SPECIES; illustrated Trachyophyta: Magnoliopsida: Caryophyllales: Cactaceae (cacti) “HOLOTYPE: USA, Arizona: Mohave Co., northwest of Bulrush Canyon south of Pipe Spring, 1400 m, n =44, 30 May 1980, B. D. Parfitt 2874 (ASU 111287!).” “PARATYPES: United States. Arizona: Coconino Co., House Rock Valley, N of Rock Canyon, Parfitt 2859 (ASU!); 1 mi N of Fredonia, Brown 651 & Parfitt (ASU!); Mohave Co., 1.2 mi N of hwy 389 on road to Moccasin, Brown 657 & Parfitt (ASU!); W of Kaibab Indian Reservation, Parfitt 3958 & Roberts (ASU!); Main Street Valley, W of Hurricane Cliffs, Palmer & Hodgson 4620 (ASU!, DES); Navaho Trail near Hurricane Rim, Gierisch 5132 (ASU!); 4.2 mi SW of Wolf Hole, Brown 851 & Parfitt (ASU!).” (Additional paratypes from southern Utah.) ________________________________________________________________________________ Peck, Stewart Blaine 1973 A systematic revision and the evolutionary biology of the Ptomaphagus (Adelops) beetles of North America (Coleoptera; Leiodidae; Catopinae), with emphasis on cave-inhabiting species. Bulletin of the Museum of Comparative Zoology, 145(2): 29-162. 232  Ptomaphagus cocytus, NEW SPECIES (p. 106, figs. 39, 40 [both p. 93, legend p. 92], 117 [p. 99, legend p. 98], 164 [p. 103, legend p. 102], 199 [p. 105, legend p. 104]) Arthropoda: Insecta: Coleoptera: Leiodidae (round fungus beetles) “Holotype male and allotype female in MCZ (no. 31893). Type locality: Arizona, Coconino County, Kaibab Plateau, Roaring Springs Cave. Type data: 1.i.1965, Gregory Lane leg. Paratypes, 2 females, same locality, 16.iv.1965, L. Ball and G. T. Lane leg., in SBP.” ________________________________________________________________________________ Peck, Stewart B., and Wynne, J. Judson 2013 Ptomaphagus parashant Peck and Wynne, new species (Coleoptera: Leiodidae: Cholevinae: Ptomaphagini): the most troglomorphic cholevine beetle known from western North America. The Coleopterists Bulletin, 67(3): 309-317. 233 Ptomaphagus parashant, NEW SPECIES ; illustrated Arthropoda: Insecta: Coleoptera: Leiodidae (round fungus beetles) “Type Specimens. Holotype female and allotype male deposited in Canadian Museum of Nature (CMN), Ottawa, Canada. Type locality: USA, Arizona, Mohave County, Parashant National 86 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Monument [Grand Canyon-Parashant National Monument], PARA-1001 cave, N36°39′, W113°37′. Type label data: female holotype; 2.ix.2011, J.J. Wynne, sweet potato bait #4, site 2, deep zone; and male allotype, same area but chicken liver bait #1. Our red label states ‘Ptomaphagus parashant Peck & Wynne, holotype.’ Paratypes, all with yellow paratype labels, seven with same locality data and five from site 2, direct intuitive searching, and two from ‘sweet potato bait #4’ are deposited at the CMN, the Los Angeles County Museum, California Academy of Sciences, and the U.S. National Museum of Natural History (Smithsonian Institution).” ________________________________________________________________________________ Pennell, Francis W. 1940 New species of Scrophulariaceae from Arizona. Notulae Naturae of the Academy of Natural Sciences of Philadelphia, (43) (May 15), 10 pp. 234  Cordylanthus tenuifolis, NEW SPECIES (pp. 9-10); not illustrated Trachyophyta: Magnoliopsida: Lamiales: Orobanchaceae (broomrapes) “Type [holotype].—Grand Canyon, Coconino County, Arizona, collected in flower August 24, 1919, by W. W. Eggleston, no. 15677a; in Herb. Academy of Natural Sciences of Philadelphia.”] ________________________________________________________________________________ Phillips, Arthur M., III 1992 A new subspecies of Rosa stellata (Rosaceae) from northwestern Arizona. Madroño, 39(1): 31-35. 235  Rosa stellata Wooton subsp. abyssa, NEW SUBSPECIES; illustrated Trachyophyta: Magnoliopsida: Rosales: Roasaceae (roses) “TYPE: USA, Arizona, Mohave Co.: SW edge of Shivwits Plateau, along W rim of Twin Point, 18 km S of Oak Grove, T30N R12W NW¼ sect. 7, 36°01′N, 113°37′W, 1823 m, in sandy to gravelly soils with limestone chips, derived form Kaibab limestone, in first 100 m from edge of plateau, open Great Basin conifer woodland, 15 Jun 1980, A. M. Phillips, III, 80-103 (holotype, ARIZ; isotypes, ASC, ASU, DES, MNA, MO, NY, UNLV, UNM, US, Lake Mead National Recreation Area herbarium).” [The locality is today within Grand Canyon-Parashant National Monument.] “Paratypes. USA, AZ, Mohave Co.: type locality . . . ; 27 Jul 1975, Holland 690 (UNLV); W rim of Kanab Canyon, gravelly soil, T38N R3W sect. 30, 1585 m., 2 Aug 1977, Gierisch 3978 (ASC, ASU); 8 Jun 1978, Gierisch 4388 (ARIZ, ASU, USDA Forest Service Herb., Albuquerque, NM); W rim of Kanab Canyon in small drainage 30 m from edge, restricted to Kaibab limestone conglomerate, T38N R3W NW¼ sect. 29, 1550 m, 13 Jun 1979, A. Phillips and B. Phillips 79624 (MNA); W rim of Kanab Canyon S of Water Canyon, in depression caused by breccia pipe collapse, T38N R3W NW¼ SW¼ sect. 8, 1525 m, 22 May 1980, A. Phillips 80-91 (ARIZ, ASC, ASU, MNA, UNLV, UNM, BLM Arizona Strip District herbarium); Grand Canyon National Monument (Park) [now in the enlarged Grand Canyon National Park], head of SB Trail, 1372 m, 24 May 1958, Riffey s.n. (UNLV, COLO); between SB Point and Hades Knoll, side canyon, 1675 m, 30 May 1978, Reichhardt 123 (MNA). Coconino Co.: Mesa Eremita [Eremita Mesa] S rim of Grand Canyon, 1980 m, 12 Jun 1935, Hawbecker s.n. (Grand Canyon National Park Study 87 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Collection); Dutton Point, N rim of Grand Canyon, dry ledge overlooking canyon, 2285 m, 17 Jul 1947, Bryant and Cooper s.n. (Grand Canyon National Park Study Collection, 2 specimens).” ________________________________________________________________________________ Pilsbry, Henry A., and Ferriss, James H. 1911 Mollusca of the southwestern states, V: The Grand Canyon and northern Arizona. Proceedings of the Academy of Natural Sciences of Philadelphia, 63: 174-199. 236  Oreohelix yavapai profundorum, NEW SUBSPECIES (pp. 182-184, text-fig. 4; Pl. XII, figs. 1-14 [legend p. 198]) Mollusca: Gastropoda: Stylommatophora: Oreohelicidae (mountain snails) “Grand Canyon, in ‘Spectacle Cove,’ Station A, the head of a recess in the cross-bed sandstone [Coconino Sandstone] south of where the Mystic Spring or Bass Trail [South Bass Trail] zigzags down, in a talus resting on the red sandstone forming the Le Conte Plateau. Elevation about 5,700 feet. Cotypes [syntypes], No. 103,234 A. N. S. P., collected by Ferriss and Pilsbry, October, 1906.” (Text-fig. 5 is a sketch view of “Spectacle Cove (Station A), from opposite side below Bass Trail. Type locality of Oreohelix yavapai profundorum on the mound at left end of talus slope.”) 237  Oreohelix yavapai extremitatis, NEW SUBSPECIES (pp. 184-185, Pl. XII, figs. 15-21 [legend p. 198]) “At Station 2, near Bass’s Trail [South Bass Trail], about 200 feet below the rim of the Grand Canyon [. . .] Cotypes [syntypes] are No. 103,236 A. N. S. P., collected by Ferriss and Pilsbry, 1906.” 238  Oreohelix yavapai angelica, NEW SUBSPECIES (pp. 185-186, Pl. XII, figs. 22-25 [legend p. 198]) “On the Bright Angel Trail, at Grand Canyon, from 100 to 400 feet below the rim [. . .] Cotypes [syntypes] are No. 103,239 A. N. S. P., collected by Ferriss and Pilsbry, 1906.” 239  Pupilla syngenes avus, NEW SUBSPECIES (p. 196, text-fig. 9) Mollusca: Gastropoda: Stylommatophora: Pupillidae (chrysalis snails) “Types [syntypes] No. 94,220 A. N. S. P., from upper slope of the Grand Canyon along the Mystic Spring or Bass Trail [South Bass Trail], about 200 feet below the rim, Station 2 . . . .” ________________________________________________________________________________ 88 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Pogue, Michael G. 2004 A new species of Schinia Hübner from riparian habitats in the Grand Canyon (Lepidoptera: Noctuidae: Heliothinae). Zootaxa, (788): 1-4. 240  Schinia immaculata, NEW SPECIES; illustrated Arthropoda: Insecta: Lepidoptera: Noctuidae (owlet moths) “Type material. HOLOTYPE: ♂, Arizona, Coconino Co., Colorado River, Grand Canyon, river mile 23.0 L, shore, 36.6003° N, 111.7668° W, May 2, 2003, R. J. Delph, genitalia slide USNM 47867, USNM ENT 00219964. The holotype is provisionally deposited at the USNM, Washington, DC, pending mutual resolution and agreement with the National Park Service regarding specimen deposition.” “PARATYPES (3°): U.S.A.: ARIZONA: COCONINO CO. 1° same data as holotype except: USNM ENT 00210120 (NAU); river mile 166.5 L, old high water, 36.2542 N, 112.8996 W, 14 Apr. 2003 (1°), R. J. Delph, USNM ENT 00219965 (USNM); river mile 202 R, new high water, 36.0526 N, 113.3489 W, 15 May 2001 (1°), J. Rundall, USNM ENT 00210119 (NAU). Paratypes deposited in the National Museum of Natural History, Washington, DC (USNM) and Northern Arizona University, Flagstaff, AZ (NAU).” paratypes may be a typographical error for [Note: The symbol ° as which appears in this list of ♂ (the female of the species being “unknown”); note as well that it is omitted from the geographical coordinates and that these symbols are correctly printed in the data for the holotype (above).] ________________________________________________________________________________ Poppius, B. [Bertil Robert] 1914 Übersicht der Pilophorus Arten nebst Beschreibung verwandter Gattungen (Hem. Het.). Annales de Société Entomologique de Belgique, 58: 237-254. [Article is in German.] [Volume 58, 1914-1919; signature 16 of this volume, which includes the two species listed here, is dated 20 August 1914.] Arthropoda: Insecta: Hemiptera: Miridae (jumping tree bugs) [The localities for the type specimens, cited peculiarly, refer to Williams, Arizona, and Bright Angel, Arizona (Grand Canyon village).] 241  Pilophorus americanus, NEW SPECIES (p. 243) “Ver. Staaten von N. Amerika : Williams, Ar.!, 21.VII; Br’t Angel, Ar.!, 12.VII, H. S. Barber, 2 ♂♂ (U. S. Nat. Mus.).” [Syntypes.] 242  Pilophorus exiguus, NEW SPECIES (pp. 246-247) “Vereinigte Staaten von N. Amerika : Br’t Angel, Ar.!, 10 VII, Barber, 1 ♀, 2 ♂♂ (U. S. Nat. Mus., Mus. Helsingf.)” [Syntypes.] ________________________________________________________________________________ 89 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Raven, Peter H. 1962 The systematics of Oenothera subgenus Chylismia. University of California Publications in Botany, 34(1): 1-122. Trachyophyta: Magnoliopsida: Myrtales: Onagraceae (evening primroses) 243  Oenothera confertiflora, NEW SPECIES (pp. 80-81); not illustrated “Type.—In cinder soil, on east side, base of Vulcan’s Throne, Toroweap Valley, at about 4,200 feet elevation, Grand Canyon National Monument [now in the enlarged Grand Canyon National Park], Mohave County, Arizona, April 30, 1952, E. McClintock 52-294 (CAS 373,481; isotypes, ARIZ, NY, WS).” “Known only from the type locality.” 244  Oenothera specuicola, NEW SPECIES (p. 87); not illustrated  Oenothera specuicola subsp. specuicola, NEW SUBSPECIES (p. 87); not illustrated “Type [holotype].—Debris slides and crevices of Redwall limestone, Kaibab Trail [South Kaibab Trail] at 5,300 feet elevation, south rim of Grand Canyon, Coconino County, Arizona, June 8, 1958, P. H. Raven 13119 (RSA; isotypes to be distributed).” 244a Oenothera specuicola subsp. hesperia, NEW SUBSPECIES (p. 88); not illustrated “Type [holotype].—Near Mooney Falls, Havasu Canyon, Coconino County, Arizona, May 23, 1950, J. T. Howell 26486 (CAS 359,774); isotype, ARIZ).” [Locality is in the Havasupai Indian Reservation.] [No. 244a added late to this checklist.] ________________________________________________________________________________ Reeder, John R. 1991 A new species of Panicum (Gramineae) from Arizona. Phytologia, 71(4) (October): 300303. 245 Panicum mohavense, NEW SPECIES ; illustrated Trachyophyta: Magnoliopsida: Poales: Poaceae (grasses) “TYPE: UNITED STATES. Arizona: Mohave Co., Arizona Strip: Main Street Valley, ca. 1.5 km S of the jct. of the Colorado City road with the Main Street Valley road. Low hills with limestone terraces, ca. 1525 m, 11 Oct 1990, J.R. & C.G. Reeder 8630 (HOLOTYPE: ARIZ; Isotypes: US,K,MO). ________________________________________________________________________________ 90 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Rehder, Alfred 1917 The genus Fraxinus in New Mexico and Arizona. Proceedings of the American Academy of Arts and Sciences, 53(2) (October): 199-212. Trachyophyta: Magnoliopsida: Lamiales: Oleaceae (olives) 246  Fraxinus cuspidata var. macropetala, NEW VARIETY (pp. 201-202); not illustrated [Syntypes include:] “Arizona. Coconino Co.: Grand Canyon, July 9, 1892, E. O. Wooton (U. S. Nat. Herb. No. 738265), July 13, 1892, J. W. Tourney (No. 273), June 1905, C. A. Purpus, June 13, 1913, A. E. Hitchcock (Nos. 76, 78); Red Canyon Trail [Hance Trail], June 10, 1901, L. F. Ward (U. S. Nat. Herb. No. 410119); Bright Angel Trail, alt. 1400-2200 m., 1909, E. W. Nelson (No. 108), May 31 and August 19, 1913, E. A. Goldman (Nos. 2067, 2223), alt. 1400-2000 m., July 19, 1914, A. Rehder (No. 106); Hermit Trail, June 18, 1916, Alice Eastwood (No. 5822); Grand View Trail, June 16, 1916, Alice Eastwood (No. 5693).” 247  Fraxinus lowellii Sargent, NEW SPECIES (p. 211); not illustrated (Type [holotype] from Oak Creek Canyon, Arizona.) [Paratype material includes:] “Mohave Co.: Peach Springs, 1884, J. G. Lemmon (No. 3242).” [This species description in Rehder is separately signed by C[harles]. S[prague]. Sargent, thus Fraxinus lowellii Sargent in Rehder, 1917.] [J. G. Lemmon, with his wife, spent a month in the Peach Springs area in May 1884. For a general (non-scientific) account of their visit, see J. G. Lemmon, 1888, Grand Cañon of the Colorado, Overland Monthly, new series, 12(69) (September): 244-256.] ________________________________________________________________________________ Rehn, James A. G., and Grant, Harold J., Jr. 1958 A revision of the genus Morsea (Orthoptera; Acridoidea; Eumastacidae). Transactions of the American Entomological Society, 84(3/4) (September/December): 217-259. 248 Morsea california kaibabensis, NEW SUBSPECIES ; (pp. 250-254, text-figs. 23-26) Arthropoda: Insecta: Orthoptera: Eumastacidae (monkey grasshoppers) “Type [holotype].— ♂; Northwest escarpment slope of Kaibab Plateau, Coconino County, Arizona. Elevation, 6700 feet. September 3, 1926. (Rehn and Hebard.) [Academy of Natural Sciences of Philadelphia, Type no. 5810].” [square brackets are part of quotation]. ________________________________________________________________________________ 91 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Reinhard, H. J. 1953 New muscoid Diptera from the western United States. Pan-Pacific Entomologist, 29(1) (January): 49-59. 249 Fabriciella evanida, NEW SPECIES (pp. 53-54); not illustrated Arthropoda: Insecta: Diptera: Tachinidae (tachinid flies) (Holotype and allotype from Lake City, Colorado.) Paratypes include: “1 ♂, S. Grand Canyon, Ariz., August 17, 1949 (P. R. Fitzgerald).” ________________________________________________________________________________ Reveal, James L. 1974a Two shrubby novelties in Eriogonum (Polygonaceae) from the deserts of Utah and Arizona. Brittonia, 26 (January/March): 90-94. 250 Eriogonum mortonianum, NEW SPECIES (pp. 91-92, fig. 1) Trachyophyta: Magnoliopsida: Caryophyllales: Polygonaceae (knotweeds) “Type: UNITED STATES: ARIZONA: Mohave Co.: along Arizona Hwy. 389, 4.5 mi SW of U.S. Hwy. 89A at Fredonia (Coconino Co.) and ca. 9 mi ENE of Pipe Springs [Pipe Spring], on red clay hills, associated with Eriogonum corymbosumn and Atriplex, at 4650 ft (1420 m), 16 Aug 1972, Reveal 2904 (HOLOTYPE: US!; ISOTYPES: ARIZ, ASU, BRY, CAS, GH, K, MO, NY, OKL, RM, RSA, SMU, TEX, UC, UTC, WTU, and elsewhere). _______________________________________________________________ 1974b Two new varieties of Eriogonum (Polygonaceae) from the Intermountain Region. Great Basin Naturalist, 34(3): 245-246. 251 Eriogonum thompsonae S. Wats. var. atwoodii, NEW VARIETY (pp. 245-246); not illustrated Trachyophyta: Magnoliopsida: Caryophyllales: Polygonaceae (knotweeds) “Typus: Arizona: Mohave Co.: Along Arizona Highway 389, 4.3 miles west of the junction of U.S. Highway 89A at Fredonia, on rolling reddish clay hills, associated with Atriplex, Ephedra, Stanleya, and Eriogonum mortonianum, at about 4700 feet elevation, 15 August 1973, Reveal & Reveal 5211. Holotypus, US! Isotypi, 15 duplicates to be distributed from US.” ________________________________________________________________________________ Rhodes, S. L., and Ayers, T. J. 2010 Two new taxa of Scutellaria Section Resinosa (Lamiaceae) from northern Arizona. Journal of the Botanical Research Institute of Texas, 4(1): 19-26. 252 Scutellaria potosina var. kaibabensis, NEW VARIETY (pp. 20, 24, figs. 2A-E, 3A-C) Trachyophyta: Magnoliopsida: Lamiales: Lamiaceae (mints) “Type: U.S.A. Arizona: Coconino Co.: East rim of the Kaibab Plateau, North Canyon Trail #4, 36°25′04″N 112°04′17″W (NAD 27), 2133 m (7000 ft), 9 Jul 1998, Suzanne Rhodes 9811 with 92 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Tina Ayes (holotype ASC; isotypes: ARIZ, ASU).” “Paratype: U.S.A. Arizona. Coconino Co.: Kaibab Plateau, East Rim View Point along Trail #7 ca. 2¼ mi down from trailhead, in North Canyon; 36°25′18″N, 112°04′22″W, 2133 m (7000 ft); 1 Jul 1993, W. Hodgson 7334 (DES, ASU).” ________________________________________________________________________________ Ribble, D. W. 1974 A revision of the bees of the genus Andrena of the western hemisphere; subgenus Scaphandrena. Transactions of the American Entomological Society, 100(2) (June): 101189. 253  Andrena (Scaphandrena) kaibabensis, NEW SPECIES (pp. 160-161); not illustrated Arthropoda: Insecta: Hymenoptera: Andrenidae (andrenid bees) “Type Material.—The holotype female (UCD) and 3 ♀♀ paratypes (UCS, INHS) were collected on the North Rim of Grand Canyon National Park, June 21, 1949, by W. H. Lang. One female paratype (AMNH) was collected 20 miles south of Jacob Lake, Arizona [in the Kaibab National Forest], June 19, 1958, by W. J. and J. W. Gertsch.” (Also another paratype from Red Canyon, Garfield Co., Utah.) ________________________________________________________________________________ Robinson, Benjamin Lincoln 1899 Three new Choripetalae from North America and Mexico. Botanical Gazette, 28(2) (August): 134-136. 254  Silene rectiramea, NEW SPECIES (p. 134); not illustrated Trachyophyta: Magnoliopsida: Caryophyllales: Caryophyllaceae (pinks) “Collected by Professor D. T. MacDougal about the Grand Cañon of the Colorado in Arizona, altitude 2150 m, 28 June, 1898, no. 181. Type in herb. Gray.” ________________________________________________________________________________ Robinson, Benjamin Lincoln, and Fernald, M. L. 1894 Contributions from the Gray Herbarium of Harvard University, New Series, No. VIII. New plants collected by Messrs. C. V. Hartman and C. E. Lloyd upon an archæological expedition to northwestern Mexico under the direction of Dr. Carl Lumholtz. Proceedings of the American Academy of Arts and Sciences, 30: 114-123. 255  Crossosoma parviflora, NEW SPECIES (p. 114); not illustrated Trachyophyta: Magnoliopsida: Crossosomatales: Crossosomataceae (crossomas) [Syntypes:] “Collected in the Grand Cañon of the Colorado, by Dr. Gray, February to May, 1885, but referred to Glossopetalon Nevadense, Gray; and at La Tinaja, Sonora, at 3,700 feet, by Mr. Hartman, 19 November, 1890 (no. 245).” 93 BIBLIOGRAPHY AND ANNOTATED CHECKLIST [In May 1885 Gray descended into Grand Canyon in Peach Springs Canyon to Diamond Creek and the Colorado River.] ________________________________________________________________________________ Robinson, Harold 1967 New species of Dolichopodidae from the United States and Mexico (Diptera). Proceedings of the Entomological Society of Washington (Washington, D.C.), 69(2) (June): 114-127. 256  Chrysotimus arizonicus, NEW SPECIES (p. 125, fig. 29 [p. 118]) Arthropoda: Insecta: Diptera: Dolichopodidae (longlegged flies) “Holotype ♂, allotype ♀, and 1♀ paratype, from Grand Canyon National Park (north rim), Arizona, July 15, 1954, W. L. Downes, Jr. Holotype (no. 69341) and allotype in USNM, paratype in the Downes collection at the Department of Entomology, University of Illinois.” ________________________________________________________________________________ Rollins, Reed C. 1953 A new Sisymbrium from Arizona. Leaflets of Western Botany, 7(1): 15-16. 257  Sisymbrium kearneyi, NEW SPECIES; not illustrated Trachyophyta: Magnoliopsida: Brassicales: Brassicaceae [Cruciferae] (mustards) “Type in the Gray Herbarium, collected along trail into S. B. Canyon, Grand Canyon National Monument [now in the enlarged Grand Canyon National Park], Arizona, May 8, 1952, Elizabeth McClintock 481a. Another collection (in flower) from the same locality but found growing under shelter of overhanging cliffs is Elizabeth McClintock 52-481 (OH). Duplicates in Herb. Calif. Acad. Sci.” _______________________________________________________________ 1982 Species of Draba, Lesquerella and Sibara (Cruciferae). Contributions from the Gray Herbarium of Harvard University, (211) (January 15): 107-113. 258 Lesquerella kaibabensis Rollins, NEW SPECIES (pp. 110-111); not illustrated Trachyophyta: Magnoliopsida: Brassicales: Brassicaceae [Cruciferae] (mustards) “Holotype in the Gray Herbarium, collected on a limestone-clay knoll, 18.6 miles south of Jacob Lake on road to the north entrance of Grand Canyon National Park, Coconino County, Arizona, June 8, 1979, Reed C. and Kathryn W. Rollins 79191. (Isotypes to be distributed.)” ________________________________________________________________________________ 94 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Rydberg, Per Axel 1909 Studies on the Rocky Mountain flora—XX. Bulletin of the Torrey Botanical Club, 36(12) (December): 675-698. 259  Penstemon coccinatus, NEW SPECIES (p. 691); not illustrated Trachyophyta: Magnoliopsida: Lamiales: Plantaginaceae (plantains) “Arizona: Grand Cañon of the Colorado, 1898, MacDougal 173 (type [holotype], in herb. N. Y. Bot. Gard.) . . . .” Paratype material includes: “Red Cañon Trail [Hance Trail], June 10, 1901, L. F. Ward”. (Other paratypes from Oak Creek, Arizona, and localities in Utah.) _______________________________________________________________ 1913 Studies on the Rocky Mountain flora—XXIX. Bulletin of the Torrey Botanical Club, 40(9) (September): 461-485. 260 Amsonia eastwoodiana, NEW SPECIES (p. 465) Trachyophyta: Magnoliopsida: Gentianales: Apocynaceae (dogbane) (Type [holotype] from Moab, Utah.) Paratype material includes “Lee’s Ferry, 1890, M. E. Jones”. (Other paratype material from east of Holbrook, Arizona, and from Willow Creek Canyon, Utah.) ________________________________________________________________________________ Rydberg, Per Axel, and Pennell, Francis Whittier 1919 North American flora. Volume 24, Part 1. (Rosales) Fabaceae, Psoraleae. Per Axel Rydberg. Eysenhardtia. Francis Whittier Pennell. New York: New York Botanical Garden, pp. 1-64. [Pennell’s contribution on the genus Eysenhardtia restricted to pp. 34-40.] Pediomelum Rydberg, 261 NEW GENUS (p. 17) Pediomelum retrorsum Rydberg, NEW SPECIES (p. 22); not illustrated Trachyophyta: Magnoliopsida: Fabales: Fabaceae (legumes) “Type [holotype] collected at Peach Spring [Peach Springs] near Big Bend of the Colorado, Arizona, June, 1884, J. G. Lemmon & wife (herb. Columbia Univ.).” [Lemmon, with his wife, spent a month in the Peach Springs area in May 1884. For a general (non-scientific) account of their visit, see J. G. Lemmon, 1888, Grand Cañon of the Colorado, Overland Monthly, new series, 12(69) (September): 244-256.] ________________________________________________________________________________ 95 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Sargent, C. S. [Charles Sprague] 1919 Notes on North American trees. IV. Botanical Gazette, 67(3): 208-242. 262  Juniperus utahensis var. megalocarpa, NEW VARIETY (p. 208); not illustrated Trachyophyta: Pinopsida: Pinales: Cupressaceae (cypress) [Syntypes include:] “Arizona: rim of the Grand Canyon, C. S. Sargent, September 9, 1904; Angel, near Flagstaff, Percival Lowell, September 4.” [Inasmuch as there seems to be no separate place name in Arizona by the simple name of “Angel”, Lowell’s gathering is presumably from “Bright Angel” (Grand Canyon village).] 263  Populus fremontii var. toumeyi, NEW VARIETY (p. 214); not illustrated Trachyophyta: Magnoliopsida: Malpighiales: Salicaceae (willows) (Type [holotype] from Tucson, Arizona.) Paratype material includes “Hermit Creek, Grand Canyon of the Colorado, Coconino County, Alice Eastwood, April 10, 1917 (no. 6002)”. ________________________________________________________________________________ Scarbrough, Aubrey G.; Stevens, Lawrence E.; and Nelson, C. Riley 2012 The albibarbis-complex of Efferia Coquillett, 1910 from the Grand Canyon region, southwestern U.S.A., with three new species and new distribution records (Diptera: Asilidae). Pan-Pacific Entomologist, 88(1) (January): 58-86. 264  Efferia tapeats, NEW SPECIES (pp. 75-78, figs. 25-30) Arthropoda: Insecta: Diptera: Asilidae (robber flies) [In this entry all square brackets are part of the quotation, with awkward typography thus. “GCNP” = Grand Canyon National Park. “HualIR/Peach Spr. Wash” = Hualapai Indian Reservation/ Peach Springs Wash.] “Types. Arizona: Coconino Co., GCNP/Clear Cr., 4 km up [from confluence with Colorado River]/12.ix.2005; 775 m/L.E. Stevens (holotype ♂, MNA). Paratypes: same data as holotype (1♀, MNA); GCNP/Colo. R. [Colorado River] Mi 108R [north bank] resting on Aster [Chloracantha] spinosus flower; 20.ix.2009; 670 m/L.E. Stevens (1♂, MNA); GCNP/S. [South] Bass Tr [Trail] in Supai Fmn. [Formation]/8.vii.2009; 1500 m/L.E. Stevens (1 ♀, MNA); GCNP/Colo. R. Mi 108R/Shinumo Cr. Mouth/21.ix.2009; 665 m/L.E. Stevens (1 ♀, MNA); GCNP/Colo. R. M. 108 L/12.vii.2007; 670 m/L.E. Stevens (1 ♂, MNA); Awatubi/Canyon (RM 58.2R) in Grand/Canyon Natl. Pk. 8.v.[19]92: M.E. Douglas, ASU (1 ♂, ASUT); GCNP/S. [South] Bass Trail, base of Tapeats (Fmn) [formation]/12.vii.2007; 900 m/L.E. Stevens (1 ♀, MNA); GCNP/Shinumo Cr. 3 km fr mouth/11.vii.2007; 810 m/L.E. Stevens (3♀, MNA); GCNP/S. [South] Bass Trail in Redwall Fmn/12.vii.2007; 1200 m/L.E. Stevens (1 ♀, MNA); same data except 13.vii.2007 (1♀, MNA); Mohave Co., HualIR/Peach Spr. Wash 4 km fr. CR [from Colorado River]/08.x.2008; 380 m/L.E. Stevens (1 ♀, MNA).” (Also a paratype from San Juan Spring, Glen Canyon National Recreation Area, Utah.) ________________________________________________________________________________ 96 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Schell, Dorothydean Viets 1943 The Ochteridae (Hemiptera) of the western hemisphere. Journal of the Kansas Entomological Society, 16(1) (January): 29-36, (2) (April): 37-47. 265  Ochterus barberi, NEW SPECIES (p. 32, Pl. 1, fig. 8 [p. 35]; pp. 41-42) Arthropoda: Insecta: Hemiptera: Ochteridae (velvety shore bugs) Species description and figure appear in the January issue; additional descriptive data and itemization of type materials appear in the April issue. “The holotype and allotype are deposited in the United States National Museum. The holotype, a male, was taken at Colorado Canyon, Arizona; the allotype at Hot Springs, Arizona. Another specimen in the National Museum bears the label, Orizaba, Mexico. Two specimens, paratypes, were received from the American Museum of Natural History, collected at Grand Canyon, Arizona. A series of 61 paratypes were collected at Indian Hot Springs and Castle Hot Springs, Arizona by Doctor R. H. Beamer and his collecting party in August, 1941. They are now in the Francis Huntington Snow collection.” [No further itemized data regarding type material; data quoted in their entirety here in order to embrace the disparate locality and collecting data as published.] ________________________________________________________________________________ Schenk, John J.; Hodgson, Wendy; and Hufford, Larry 2010 A new species of Mentzelia section Bartonia (Loasaceae) from the Grand Canyon, Arizona. Brittonia, 62(1): 1-6. 266  Mentzelia hualapaiensis, NEW SPECIES; illustrated Trachyophyta: Magnoliopsida: Cornales: Losaceae (loasa) “Type: U.S.A. Arizona: Mohave County, Grand Canyon National Park, along the Colorado River near the Mile 158 camp site on river right, 339451 E, 4018170 N, NAD 83, 36°, 17′, 42.6″ N; 112°, 47′, 16.6″ W, 990 m, 19 Apr 2008, L. Makarick, D. Hall, and J. Cipra 2008-13 (holotype: DES; isotype: GCNP).” _______________________________________________________________ 2013 Mentzelia canyonensis (Loasaceae), a new species endemic to the Grand Canyon, Arizona, U.S.A. Brittonia, 65(4) (December): 408-416. 267  Mentzelia canyonensis, NEW SPECIES; illustrated Trachyophyta: Magnoliopsida: Cornales: Losaceae (loasa) “Type: U.S.A. Arizona. Coconino Co.: Grand Canyon National Park, Manzanita Canyon, within ¼ mile of Bright Angel Creek and North Kaibab Trail, UTM 12S 4005015 N, 407532 E, 4830 ft, 27 Sep. 2010, W. Hodgson 25718 (holotype: DES; isotypes: GCNP, NY, WS).” ________________________________________________________________________________ 97 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Scudder, Samuel Hubbard 1878 Notice of the butterflies collected by Dr. Edward Palmer in the arid regions of southern Utah and northern Arizona during the summer of 1877. U.S. Geological and Geographical Survey of the Territories, Bulletin, 4: 253-258. 268  Neominois dionysus, NEW SPECIES (p. 254); not illustrated Arthropoda: Insecta: Lepidoptera: Nymphalidae (admirals) Syntype material comprises eight males and five females from “Juniper Mountains [extralimital to this checklist] June 4; Mount Trumbull, June 7-10”. 269 Synchloe thoosa, NEW SPECIES (p. 257); not illustrated Arthropoda: Insecta: Lepidoptera: Pieridae (orange-tips) “A single female [holotype] was taken at Mokiak Pass, April 28-30 or June 2.” _______________________________________________________________ 1897 Revision of the orthopteran group Melanopli (Acridiidæ), with special reference to North American forms. Proceedings of the U.S. National Museum, 20: 1-421, plates 1-26. [Family Acrididae spelled Acridiidae in title.] 270  Melanoplus canonicus, NEW SPECIES (pp. 300-301, Pl. 20, fig. 1) Arthropoda: Insecta: Orthoptera: Acrididae (short-horned grasshoppers) “One male, 1 female. Grand Canyon of the Colorado, Arizona, July 10 (L. Bruner).” ________________________________________________________________________________ Shaw, F. R. 1951 Some new Mycetophilidae from the western United States. Bulletin of the Brooklyn Entomological Society, 46(3) (June): 65-70. 271  Mycetophila denningi, NEW SPECIES (p. 67, Pl. 3, fig. 4) Arthropoda: Insecta: Diptera: Mycetophilidae (fungus gnats) “Described from one male [holotype] collected by D. G. Denning at Grand Canyon, Arizona on June 18, 1949.” ________________________________________________________________________________ 98 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Shear, William A.; Taylor, Steven J.; Wynne, J. Judson; and Krejca, Jean K. 2009 Cave millipeds of the United States. VIII. New genera and species of polydesmidan millipeds from caves in the southwestern United States (Diplopoda, Polydesmida, Macrosternodesmidae). Zootaxa, (2151): 47-65. Pratherodesmus Shear, 272 NEW GENUS Pratherodesmus voylesi Shear, (p. 50) NEW SPECIES (pp. 51-54, figs. 2, 6, 9, 11–14, 19 [p. 56], 33 [p. 61]) Arthopoda: Diplopoda: Polydesmida: Macrosternodesmidae (presumed troglobitic polydesmidan millipeds) “Types: Male holotype, five male and seven female paratypes from Millipede Cave, Mojave [sic] Co., Arizona, collected 16 February 2003 by K. Voyles, J. Jasper, M. Porter and K. Dittmar de la Cruz, deposited in FMNH. Four male and sixteen female paratypes from October Gyp Cave, collected 27 March 2004 by same collectors. All specimens deposited in FMNH and CPMAB.” ________________________________________________________________________________ Sinclair, B. J. 2006 A new species of Wiedemannia Zetterstedt from Grand Canyon National Park, with notes on additional Nearctic species (Diptera: Empididae). Journal of the Entomological Society of Ontario, 137: 25-30. 273  Wiedemannia digna, NEW SPECIES (pp. 27-29, fig. 1) Arthropoda: Insecta: Diptera: Empididae (balloon flies) “Type Material. Holotype male labelled: ‘USA:AZ:Grand Canyon NP/ Vasey’s Paradise/ 10.v.1998/ J.F. MacDonald’ (USNM). Paratype: Same data as holotype, ♂, USNM.” ________________________________________________________________________________ Small, John Kunkel 1906 Studies in North American Polygonaceae—II. Bulletin of the Torrey Botanical Club, 33(1) (January): 51-57. 274 Eriogonum fusiforme, NEW SPECIES (p. 56); not illustrated Trachyophyta: Magnoliopsida: Caryophyllales: Polygonaceae (knotweeds) (Type [holotype] from Grand Junction, Colorado.) Paratype material includes “ARIZONA: Peach Springs, June 28, I887, Tracy & Evans 315.” ________________________________________________________________________________ 99 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Smiley, Robert L., and Moser, John C. 1974 New tarsonemids associated with bark beetles (Acarina: Tarsonemidae). Annals of the Entomological Society of America, 67(4) (July): 639-665. 275  Heterotarsonemus nakaharai, NEW SPECIES (pp. 645-647, figs. 24, 25) Arthropoda: Euchelicerata: Trombidiformes: Tarsonemidae (tarsonemid mites) “Holotype.—Female, Canadian National Collection no. 13177, collected from Ips knausi Swaine in Pinus ponderosa Douglas at Grand Canyon, Arizona. The collector and date collected are unknown.” ________________________________________________________________________________ Stacey, J. W. 1937 Notes on Carex—VIII. Leaflets of Western Botany, 2(1) (January): 13-16. 276  Carex curatorium [sic], NEW SPECIES (pp. 13-14); not illustrated. Carex curatorum. [The species was named for the collectors, Alice Eastwood, curator, and J. T. Howell, assistant curator (Department of Botany, California Academy of Sciences); accordingly noticed thereafter in literature with the Latin suffix of the specific epithet grammatically corrected, as Carex curatorum Stacey.] Trachyophyta: Magnoliopsida: Poales: Cyperaceae (sedges) “Types: staminate plant, Herb. Calif. Acad. Sci., No. 204973, Eastwood and Howell No. 1101; pistillate plant, Herb. Calif. Acad. Sci., No. 204974, Eastwood and Howell No. 1100, collected June 23, 1933, on Kaibab Trail to Roaring Springs [North Kaibab Trail], Grand Canyon National Park, Arizona. Two other collections made with the type of the pistillate plant bear the numbers 1045 and 1089. Duplicates of some of these collections were distributed as C. pseudoscirpoidea Rydb., an early determination reported in the literature (Leafl. West. Bot. 1: 142, 1934).” ________________________________________________________________________________ Stahnke, Herbert L. 1940 277 The scorpions of Arizona. Iowa State College Journal of Science, 15 (October): 101-103. Vejovis [sic] aquilonalis, NEW SPECIES (p. 101); not illustrated. Vaejovis aquilonalis. Arthropoda: Euchelicerata: Scorpiones: Vaejovidae (vaejovid scorpions) “The specimen, a male, was taken 37 miles south of the Grand Canyon on highway 64.” [see note below]. The type locality is probably a typographical error, as clarified by W. David Sissom and Oscar F. Francke (1981), Scorpions of the genus Paruroctonus from New Mexico and Texas (Scorpiones, Vaejovidae), Journal of Arachnology, 9(1) (Winter): 93-108, under their discussion of Paruroctonus aquilonalis (Stahnke), p. 94: “Type data.—Holotype male from 30 miles south of the Grand Canyon, Arizona, 8 August 1938 (Kay Anderson). Deposited in the collection of H. L. Stahnke; examined. [. . .] The discrepancy in the type locality data seems to be a typographical error, for the label with the holotype clearly reads ‘30 mi. South.’ ” [Note: There is no indication 100 BIBLIOGRAPHY AND ANNOTATED CHECKLIST that the label with the specimen is a field label; if written for the collection drawer, it could itself be in error, obfuscating everything. —E.E.S.] ________________________________________________________________________________ Stallings, Don B., and Turner, J. R. 1957 Four new species of Megathymus (Lepidoptera, Rhopalocera, Megathymidae). Entomological News (Academy of Natural Sciences of Philadelphia, Entomological Section), 68(1) (January): 1-16. 278 Megathymus alliae, NEW SPECIES (pp. 1-5, Pl. 1) Arthropoda: Insecta: Lepidoptera: Hesperiidae (skippers) “Described from 62 specimens (35 males and 27 females) collected 15 miles west of Cameron, Ariz., along the canyon of the Little Colorado River, elevation 5000 ft. All ex-larvae or expupae emerging from Aug. 25 to Oct. 5, during 1953, 1954 and 1955. Collected by Dr. and Mrs. R. C. Turner, Dee, Jack, Don, and Viola Stallings.” [Quite probably the type locality is at or near the Little Colorado River Gorge Overlook, along Arizona Route 64.] “Holotype, female, Sept. 23, 1955, 15 miles west of Cameron, ARIZONA, el. 5000 ft. (Turner); allotype, male, Sept. 17, 1954, 15 miles west of Cameron, Ariz., el. 5000 ft. (Turner) are in the collection of the authors. Paratypes of both sexes will be placed in the following collections: H. A. Freeman, C. L. Remington, U. S. Nat. Museum, American Mus. of Nat. History, Los Angeles County Museum.” ________________________________________________________________________________ Standley, Paul C. 1909 The Allioniaceae of the United States with notes on Mexican species. Contributions from the U.S. National Herbarium, 12(Part 8): 303-389, plates. (As part of the Bulletin of the U.S. National Museum.) 279  Allionia pachyphylla, NEW SPECIES (p. 346); not illustrated Trachyophyta: Magnoliopsida: Caryophyllales: Nyctaginaceae (four o’clocks) “Type [holotype] U. S. National Herbarium no. 211717, collected in Arizona at the Grand Canyon, 1892, Toumey 485 [see note following]; cotype [isotype] in the herbarium of the University of Arizona. Other specimens seen [paratypes]: Arizona: Red Canyon Trail [Hance Trail], Grand Canyon, 1901, Ward; Grand Canyon, 1892, Wooton; Camp Verde, 1891, MacDougal.” [Toumey 485 is very likely from the Old Hance Trail, which locality he visited in 1892; see notes accompanying Coville (1894) in the present volume. Similarly, the Wooton gathering is likely from the same locale, inasmuch as Wooton accompanied Toumey and company on the trip to the Grand Canyon. For a general (non-scientific) account of the expedition, see C. H. Tyler Townsend (1893), A wagon-trip to the Grand Cañon of the Colorado River, Appalachia, 7(1) (February): 4863. Note that the Ward gathering in 1901 is from the newer Hance Trail, which replaced the Old Hance Trail that was destroyed by landslides in 1895.] ________________________________________________________________________________ 101 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Stearns, Robert E. C. 1890 Scientific results of explorations by the U.S. Fish Commissions [sic] steamer Albatross. No. XVII.—Descriptions of new west American land, fresh-water, and marine shells, with notes and comments. Proceedings of the U.S. National Museum, 13: 205-225. 280  Helix (Arionta) coloradoensis, NEW SPECIES (p. 206, Pl. XV, figs. 6, 8, 12 [not 6-8; see note below], legend p. 225) Mollusca: Gastropoda: Stylommatophora: Helminthoglyptidae (shoulderbands) “Grand Cañon of the Colorado, opposite the Kaibab plateau, at an elevation of 3,500 feet. (Mus. No. 104100.)” [The collection was made during the 1889 biological survey of C. Hart Merriam, thus the locality is the Old Hance Trail; also note that the elevation indicates the collection was made along the trail inside the canyon. See remarks with C. Hart Merriam in the present volume. The species was later reassigned to the helminthoglyptid genus Sonorella.] [Figures 7 and 12 of Pl. XV were transposed (fide H. A. Pilsbry, 1893, Manual of conchology : structural and systematic. Second series: Pulmonata. Vol. VIII, Part 4, Helicidæ, Vol. VI. Philadelphia: Academy of Natural Sciences of Philadelphia, Conchological Section, p. 226).] ________________________________________________________________________________ Stickel, William H. 1938 The snakes of the genus Sonora in the United States and Lower California. Copeia, 1938: 182-190. 281  Sonora semiannulata gloydi, NEW SUBSPECIES (pp. 186-187); not illustrated Chordata: Reptilia: Squamata: Colubridae (typical snakes) “Type [holotype].—MZUM 83754, collected by Mr. William Holzmark in 1936 on the Bright Angel Trail, Lower Sonoran level of the Grand Canyon, Grand Canyon National Park, Arizona. Paratypes.—Grand Canyon National Park, Arizona: Grand Canyon National Park collection 107, vicinity of Indian Gardens [Havasupai Gardens]; UCLA 32, mouth of Garden Creek; MVZ 1758, Bright Angel Trail.” ________________________________________________________________________________ Stutz, Howard C.; Chu, Ge-Lin; and Sanderson, Stewart C. 1994 Atriplex asterocarpa (Chenopodiaceae), a new species from southern Utah and northern Arizona. Madroño, 41(3): 199-204. 282  Atriplex asterocarpa, NEW SPECIES; illustrated Trachyophyta: Magnoliopsida: Caryophyllales: Amaranthaceae (pigweed) (Holotype from Garfield Co., Utah.) Paratypes include: “Arizona, Coconino Co., vic. Vermilion Lodge [Vermilion Cliffs], steep rocky south slope, 4200 ft elevation, 18 Apr 1978, R. K. Gierish 102 BIBLIOGRAPHY AND ANNOTATED CHECKLIST 4194 (ASC); Lee's Ferry, south-facing slope, Chinle Formation, 23 May 1993, H. C. Stutz 95792 (BRY).” ________________________________________________________________________________ Suksdorf, Wilhelm 1931 Untersuchungen in der Gattung Amsinckia. Werdenda (Beiträge zur Pflanzenkunde) (Bingen, Washington), 1(5/8) (December 31): 47-113. [In German.] Trachyophyta: Magnoliopsida: Boraginales: Boraginaceae (borage) 283  Amsinckia nana, NEW SPECIES (pp. 84-85); not illustrated Based on a single specimen (holotype): “Ariz.; Hermit Creek, Grand Canyon of the Colorado River, 10. April 1917, Alice Eastwood, nr. 6016 (C. A.)” 284  Amsinckia macrosepala, NEW SPECIES (p. 108); not illustrated Syntypes include: “Grand Canyon, 1885, Gray (G. H.)”. [Gray descended into Grand Canyon in Peach Springs Canyon to Diamond Creek and the Colorado River.] ________________________________________________________________________________ Tanner, Wilmer W. 1953 A study of taxonomy and phylogeny of Lampropeltis pyromelana Cope. Great Basin Naturalist, 13(1/2) (September 12): 47-66. 285  Lampropettis [sic] pyromelana Cope infralabials [sic], NEW SUBSPECIES (pp. 56-60, Pl. 2, fig. 1). (Common name given by the author, “Utah Ringed Snake”) Lampropeltis pyromelana infralabialis. Chordata: Reptilia: Squamata: Colubridae (typical snakes) (Holotype from Beaver Co., Utah.) Paratypes include: “Arizona: Coconino County, Grand Canyon Collection R3, R260-2, R291, R372, and UU 21 all from or near Bright Angel Point, North Rim of Grand Canyon.” [The misspelled generic and subspecific epithets are unfortunate and do not affect the taxonomy of this snake, in no way forcing a new taxon into existence with these spellings. The genus “Lampropettis” is simply a typographical error and is in fact spelled correctly (Lampropeltis) in the title of the paper and elsewhere. The subspecies “infralabials” is spelled correctly (infralabialis) in the key to subspecies (p. 60) and elsewhere throughout the paper.] ________________________________________________________________________________ 103 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Theroux, Michael E.; Pinkava, D. J.; and Keil, D. J. 1977 A new species of Flaveria (Compositae: Flaveriinae) from Grand Canyon, Arizona. Madroño, 24(1): 13-17. 286  Flaveria mcdougallii, NEW SPECIES; illustrated Trachyophyta: Magnoliopsida: Asterales: Asteraceae (sunflowers) “Type: United States, Arizona, Mohave Co., Grand Canyon National Park, Cove Canyon (174.2 mi below Lee’s Ferry), 27 Jan 1976, M. E. Theroux 1675 (Holotype: US; isotypes: ARIZ, ASC, ASU, DES, GCNP, GH, MNA, NY, RSA, SRSC, TEX, UC). Paratypes: United States, Arizona, Coconino Co., Grand Canyon National Park, Matkatamiba Canyon (148.8 [sic, 147.8] mi below Lee's Ferry), 1 Oct 1975, M. E. Theroux 1519 (MNA); Mohave Co., Grand Canyon National Park, Cove Canyon, 5 Oct 1975, M. E. Theroux 1567 (MNA).” ________________________________________________________________________________ Thiers, Harry D. 1976 Boletes of the southwestern United States. Mycotaxon, 3(2) (January/March): 261-273. [In this paper the type information is provided only within the portion written in botanical Latin. No English equivalent was published.] 287 Boletus barrowsii Thiers & Smith ex Thiers, NEW SPECIES (pp. 262-264); not illustrated. [Smith = Alexander H. Smith.] Basidiomycota (club fungi): Agaricomycetes: Boletales: Boletaceae “Holotypus (Thiers 27816) lectus prope Jacob Lake, Kaibab National Forest, Coconino County, Arizona, Aug. 21, 1971; in herbario San Francisco State University (SFSU) conservatus.” 288 Suillus kaibabensis Thiers, NEW SPECIES (pp. 266-268); not illustrated Basidiomycota (club fungi): Agaricomycetes: Boletales: Suillaceae “Holotypus (Thiers 27813) lectus prope Jacob Lake, Kaibab National Forest, Coconino County, Arizona, Aug. 21, 1971; in herbario San Francisco State University (SFSU) conservatus.” 289  Suillus occidentalis Thiers, NEW SPECIES (pp. 268-270); not illustrated Basidiomycota (club fungi): Agaricomycetes: Boletales: Suillaceae “Holotypus (Thiers 27775) lectus prope Point Sublime, North Rim, Grand Canyon National Park, Coconino County, Arizona, Aug. 19, 1971; in herbario San Francisco State University (SFSU) conservatus.” ________________________________________________________________________________ 104 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Timberlake, P. H. [Philip H.] 1928 Bees of the genus Perdita Smith in the American Museum of Natural History (Hymenoptera). American Museum Novitates, (321), 13 pp. 290  Perdita wheeleri, new species (pp. 5-7); not illustrated Arthropoda: Insecta: Hymenoptera: Andrenidae (andrenid bees) “1 ♂ (holotype) collected in May, 1905, in Indian Gardens [Havasupai Gardens] in the Grand Cañon, Arizona (W. M. Wheeler).” _______________________________________________________________ 1956 A revisional study of the bees of the genus Perdita F. Smith, with special reference to the fauna of the Pacific coast (Hymenoptera, Apoidea); Part II. University of California Publications in Entomology, 11(5): 247-350. 291 Perdita opacella, NEW SPECIES (p. 324) Arthropoda: Insecta: Hymenoptera: Andrenidae (andrenid bees) A single female, holotype, from Marble Canyon, near Lee’s Ferry, Arizona (collected by G. D. Butler, June 5, 1953, on Stanleya. _______________________________________________________________ 1962 A revisional study of the bees of the genus Perdita F. Smith, with special reference to the fauna of the Pacific coast (Hymenoptera, Apoidea); Part V. University of California Publications in Entomology, 28(1), 123 pp. [including 13 plates]. Arthropoda: Insecta: Hymenoptera: Andrenidae (andrenid bees) 292  Perdita inornata, NEW SPECIES (pp. 49-50, figs. 787, 788 [p. 115], 872 [p. 122]) (Holotype, allotype, and paratypes from Inyo Co., California.) “Additional paratypes [include] ARIZONA, Coconino Co.: 4 females, 14 males, South Rim of the Grand Canyon, July 26, 1934 (H. E. and M. A. Evans); 1 male, Grand Canyon, July 28, 1949 (W. H. Lange).” “Types at present in collection of the Citrus Experiment Station, Riverside; paratypes in collections of the University of Kansas, the University of California at Berkeley and Davis, and Cornell University.” _______________________________________________________________ 1968 A revisional study of the bees of the genus Perdita F. Smith, with special reference to the fauna of the Pacific coast (Hymenoptera, Apoidea); Part VII (including index to Parts I to VII). University of California Publications in Entomology, (49), 196 pp. 293 Perdita depressa, NEW SPECIES (pp. 48-50, plate figs. 1217, 1218 [p. 167], 1312 [p. 175]) Arthropoda: Insecta: Hymenoptera: Andrenidae (andrenid bees) (Holotype and allotype from Monument Valley, Utah.) Paratypes include: “1 female, Supai, 3,500 feet, Havasu Canyon, Grand Canyon National Park, Coconino Co., Arizona, Aug. 2, 1934 (F. C. 105 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Lutz).” “Types in collection of the University of California, the female paratype in collection of the American Museum of Natural History.” ________________________________________________________________________________ Townsend, C. H. Tyler [Charles Haskins Tyler] 1915 New western and southwestern Muscoidea. Journal of the New York Entomological Society, 23(4) (December): 216-234. [Item signed “Charles H. T. Townsend”.] Microsciasma NEW GENUS (p. 234) 294  Microsciasma minuta, NEW SPECIES (p. 234); not illustrated Arthropoda: Insecta: Diptera: Tachinidae (tachinid flies) “One male, Grand Canyon, Arizona, near Colorado River on Hance Trail [Old Hance Trail], about 3,500 ft., July 10, 1892 (Townsend).” “Holotype, No. 19579 U. S. N. M.” [Regarding the 1892 expedition see a general (non-scientific) account by Townsend (1893), A wagon-trip to the Grand Cañon of the Colorado River, Appalachia, 7(1) (February): 48-63.] ________________________________________________________________________________ Triplehorn, Charles A. 1975 A subgenus of Eleodes with three new cave-inhabiting species (Coleoptera: Tenebrionidae). The Coleopterists Bulletin, 29(1): 39-43. Caverneleodes, NEW SUBGENUS (p. 39) 295  Eleodes (Caverneleodes) leptoscelis, NEW SPECIES (pp. 42-43); not illustrated Arthropoda: Insecta: Coleoptera: Staphylinidae (rove beetles) “Types: holotype (female) and 2 paratypes: Arizona: Coconino County, Cave of Domes [Cave of the Domes, Horseshoe Mesa], 16-X-1953; 1 paratype, same data except 14-X-1954; 2 paratypes, Coconino County, Cave 68-Olje [Tse’an Olje, Cremation Canyon], 14-X-1954; 1 paratype, Coconino County, Tse-an-cho [Tse’an Cho, Horseshoe Mesa], 7-XI-1953. Holotype (USNM #73090) and paratypes in United States National Museum; paratypes in The Ohio State University Collection of Insects and Spiders. The following information on cave locations was supplied by Paul S. Bartholomew in 1957: Cave of Domes and Tse-an-Cho are both caves on Horseshoe Mesa, about 12 miles east of South Village, which is on the south rim of the Grand Canyon; Cave 68-Olje (also called Tse-an-Olje) is a cave east of South Village [Grand Canyon village]; the 3 caves are at approximately 4500 feet elevation (T. J. Spilman, in litt.). ” ________________________________________________________________________________ 106 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Triplehorn, Charles A., and Brown, Kirby W. 1971 A synopsis of the species of Asidina in the United States with description of a new species from Arizona (Coleoptera: Tenebrionidae). The Coleopterists Bulletin, 25(3): 73-86. 296  Asidina rugicollis, NEW SPECIES (pp. 76-77, figs. 1, 7) Arthropoda: Insecta: Coleoptera: Tenebrionidae (darkling beetles) “Holotype, male . . . Allotype, female . . . U. S. A., ARIZONA, COCONINO Co., Grand Canyon National Park, South Rim, 19-VII-68, W. E., C. A., and B. W. Triplehorn [Ohio State University Collection of Insects and Spiders]. Paratypes [include]: 33 specimens, all from ARIZONA: Coconino Co.: same data as holotype, 3 males, 2 females [OSUC]; 3 males, 3 females [KWBC]; 1 male, 1 female [USNM]; Grand Canyon, 26-27-VII-26, E. C. Van Dyke, 3 males, 5 females, [CASC]; 1 male, 1 female [ASUT]; 1 male, 1 female [BMNH]; 1 male, 1 female [MHNP] . . . northwest slope of Kaibab Plateau, 3-IX-26, 6700′, R. & H., 1 female [ANSP] . . . .” [Square brackets are part of the quotation.] ________________________________________________________________________________ Troubridge, J. T., and Crabo, L. G. 2002 A review of the Nearctic species of Hadena Schrank, 1802 (Lepidoptera: Noctuidae) with descriptions of six new species. Fabreries (Association des Entomologistes Amateurs du Québec), 27(2) (December): 109-154 (“Section en français”, pp. 135-145). [Article is principally in English, with an abbreviated section (not an abstract) in French.] 297 Hadena (Hadena) lafontainei, NEW SPECIES (pp. 114-117, figs. 3 [p. 146], 17 [p. 148], 33 [p. 153]) Arthropoda: Insecta: Lepidoptera: Noctuidae (owlet moths) (Holotype from Albany Co., Wyoming.) Paratype material of 85 specimens includes “2 ♀♀, Kaibab Plateau, nr. Jacob Lk., 8,300 ft [2,550 m], 18.VII.1980, D.C. Ferguson” (square brackets are part of quotation). [“nr. Jacob Lk.” = near Jacob Lake.] ________________________________________________________________________________ Tuthill, L. D. 1939 New species of Psyllidae from the western United States. Iowa State College Journal of Science, 13(2) (January): 181-186. 298  Trioza rubra, NEW SPECIES (pp. 185-186); not illustrated Arthropoda: Insecta: Hemiptera: Psyllidae (jumping plantlice) (Holotype female, allotype male, and nine paratypes from Creede, Colorado.) Additional paratypes include: “one male, Grand Canyon, Arizona, Aug. 11, 1927, R. H. Beamer”. Holotype and allotype in author’s collection; paratypes in author’s collecton and in Snow Collection, University of Kansas, and U.S. National Museum (not specified). ________________________________________________________________________________ 107 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Usinger, Robert L. 1931 A new species of Platylygus (Miridae, Hemiptera). Pan-Pacific Entomologist, 7(3) (January): 129-130. 299  Platylygus vanduzeei, NEW SPECIES; text-fig. Arthropoda: Insecta: Hemiptera: Miridae (jumping tree bugs) “Holotype, male. No. 2997, Mus. Calif. Acad. Sciences, collected by C. D. Duncan, June 17, 1921, at the Grand Canyon, Arizona, and allotype, female. No. 2998, Mus. Calif. Acad. Sciences, collected by the author in Grand Canyon National Park, Arizona, South Rim, June 29, 1930.” (Also 12 paratype females from Mesa Verde National Park, Colorado.) ________________________________________________________________________________ Van Duzee, M. C. 1927 North American species of Polymedon. (Diptera Dolichopodidæ). Annals of the Entomological Society of America, 20(1) (March): 123-126. Arthropoda: Insecta: Diptera: Dolichopodidae (longlegged flies) 300  Polymedon dilaticosta, NEW SPECIES (pp. 124-125); not illustrated “Described from nine males and three females, taken on the Bright Angel Trail, Grand Canon, Arizona, and one female taken in Oak Creek Canon, Arizona, August, at an elevation of 6,000 feet; the last one was taken by F. H. Snow. Type and allotype in the Canadian National Collection.” [Distinguishing data for primary types not indicated, thus all specimens are originally syntypes.] 301  Polymedon nitidus, NEW SPECIES (p. 125); not illustrated “Described from one female [holotype], taken on the Bright Angel Trail, Grand Canon, Arizona. Type in the Canadian National Collection.” ________________________________________________________________________________ Wang, Daqing, and Holsinger, John R. 2001 Systematics of the subterranean amphipod genus Stygobromus (Crangonyctidae) in western North America, with emphasis on species of the hubbsi group. Amphipacifica (Journal of Aquatic Systematic Biology), 3(2) (November 15): 39-147. 302  Stygobromus blinni, new species (pp. 74-76, fig. 21) Arthropoda: Malacostraca: Amphipoda: Crangonyctidae (crangonyctid amphipods) Female holotype and two female paratypes from “Roaring Springs Cave, on the north rim of the Grand Canyon near Bright Angel Trail and Bright Angel Creek” [sic] (p. 74), “about 1.6 km inside the cave” (p. 76); collected by Dean Blinn, 28 September 1994.] ________________________________________________________________________________ 108 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Watson, Sereno 1873 New plants of northern Arizona and the region adjacent. American Naturalist, 7(5) (May): 299-303. [Also published as offprint, with header, “Contributions to American Botany. I.”, “Printed at the Salem Press. May, 1873.”, 7 pp.] “In the botanical collections made in 1871-’72, mainly in the southern portion of the Great Basin, in northwestern Arizona and the adjacent desert section of California, by Mr. Ferd. Bischoff and others, under the direction of Lieut. G. M. Wheeler, Corps of Engineers, in the course of his exploration of that region, several new species have been found which are here described, by consent, in anticipation of the fuller report now in preparation. With these are given some others occurring in a small collection made by Mrs. Ellen P. Thompson near Kanab, Southern Utah, during the last summer while accompanying her brother, Maj. [John Wesley] Powell, in his survey of the Colorado.” [Below are listed only those specimens gathered by Thompson in “Northern Arizona”, inasmuch as these were clearly obtained while on relatively short journeys into the Arizona Strip from her base in Kanab, Utah.] 303 Dalea amoena, NEW SPECIES (p. 300); not illustrated Trachyophyta: Magnoliopsida: Fabales: Fabaceae (legumes) “Northern Arizona (Mrs. E. P. Thompson). In damp places; April.” 304 Oenothera (Chylisma) multijuga, NEW SPECIES (p. 300); not illustrated Trachyophyta: Magnoliopsida: Myrtales: Onagraceae (evening primroses) “Northern Arizona (Mrs. E. P. Thompson). In dry places; March.” 305 Cymopterus purpureus, NEW SPECIES (pp. 300-301); not illustrated Trachyophyta: Magnoliopsida: Brassicales: Apiaceae (parsleys) “Northern Arizona (Mrs. E. P. Thompson). In damp, shaded soil; March. 306 Peucedanum newberryi, NEW SPECIES (p. 301); not illustrated Trachyophyta: Magnoliopsida: Brassicales: Apiaceae (parsleys) “Northern Arizona, on stony soil (Mrs. E. P. Thompson). April.” 307 Brickellia (Clavigera) longifolia, NEW SPECIES (p. 301); not illustrated Trachyophyta: Magnoliopsida: Asterales: Asteraceae (sunflowers) “Northern Arizona (Mrs. E. P. Thompson). In a damp cañon; April.” 308 Calochortus flexuosus, NEW SPECIES (p. 303); not illustrated Trachyophyta: Magnoliopsida: Liliales: Liliaceae (lilies) “Southern Utah and Northern Arizona (Mrs. E. P. Thompson); April and May. The bulbs, as of other species, are eaten by the Indians.” 109 BIBLIOGRAPHY AND ANNOTATED CHECKLIST 309 Androstephium breviflorum, NEW SPECIES (p. 303); not illustrated Trachyophyta: Magnoliopsida: Asparagales: Asparagaceae (agaves) “Southern Utah and Northern Arizona (Mrs. E. P. Thompson); April and May. Bulbs also eaten.” _______________________________________________________________ 1888 Contributions to American botany. Proceedings of the American Academy of Arts and Sciences, 23(2): 249-287. 310 Lesquerella arizonica, NEW SPECIES (p. 254); not illustrated Trachyophyta: Magnoliopsida: Brassicales: Brassicaceae [Cruciferae] (mustards) Syntypic material includes “Arizona . . . at Peach Springs (4177 Lemmon, 64 Jones), and at Mokiak Pass (43 Palmer, 1877).” [J. G. Lemmon, with his wife, spent a month in the Peach Springs area in May 1884. For a general (non-scientific) account of their visit, see J. G. Lemmon, 1888, Grand Cañon of the Colorado, Overland Monthly, new series, 12(69) (September): 244-256.] ________________________________________________________________________________ Welsh, Stanley L., and Atwood, N. Duane 2001 New taxa and nomenclatural proposals in miscellaneous families—Utah and Arizona. Rhodora (Journal of the New England Botanical Club), 103: 71-95. 311 Astragalus lentiginosus Douglas ex Hook. var. trumbullensis, NEW VARIETY (pp. 81-82, fig. 3 [p. 83]) Trachyophyta: Magnoliopsida: Fabales: Fabaceae (legumes) [Localities are within the present boundaries of Grand Canyon-Parashant National Monument.] “Type: U.S.A. Arizona: Mohave Co., Mohave Strip, Andrus Canyon, 3 mi. W of Andrus Pt., T32N R10W S6, wash bottom with Mortonia, Purshia, Ephedra, Agave, etc., 26 Apr 1999, N. D. Atwood & B. Furniss 24293 (HOLOTYPE: BRY; six isotypes to be distributed).” “Additional specimens (paratypes; all BRY!): Arizona: Mohave Co., Parashant (Trail) Canyon, 11 mi. S of Mt. Trumbull Village, 26 Apr 1974, Atwood 6029, 6029a; ditto, Whitmore Canyon, 1.25 km (7.8 mi.) [sic] S of Mount Trumbull (abandoned town), T34N R9W S29, 25 May 1979, Holmgren & Barneby 9172; Daneill Canyon W of Andrus Canyon, 3 mi. W of Andrus Pt., T32N R11W S12, 26 Apr 1999, Atwood & Furniss 24300; ditto, Andrus Canyon, 1 mi. W of Andrus Pt„ T32N R10W S10, 26 Apr 1999, Atwood & Furniss 24302; ditto, Whitmore Canyon, T32N R9W S1, 27 May 1999, Higgins 20226; ditto, 2 mi. S of Mt. Trumbull school house, near Griffith Knoll, T34S R10W S1-2, 27 May 1999, Higgins 20277.” ________________________________________________________________________________ 110 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Welsh, Stanley L., and Licher, Max 2010 Pediomelum Rydberg (Leguminosae) in Arizona and two previously undescribed species. Western North American Naturalist, 70(1): 9-18. 312 Pediomelum pauperitense S.L. Welsh, M. Licher, & N. D. Atwood, NEW SPECIES (pp. 14-15, fig. 3). (Common name given by the author, “Poverty Mountain Breadroot”) Trachyophyta: Magnoliopsida: Fabales: Fabaceae (legumes) “Type.—USA, Arizona, Mohave Co., T35N, R12W, Sec. 35/36, SW of Poverty Mountain, near Dewdrop Spring; 1756 m, in gravelly lime- stone soil in pinyon-juniper community, L.C. Higgins 23135, 25 May 2001, holotype BRY, isotypes distributed previously as P. mephiticum.” “Paratypes.—Arizona, Mohave Co., T34N, R11W, Sec 4, SE of Poverty Mountain, Arizona Strip, SW of Mt. Trumbull, 5500 ft, Atwood & Franklin 18901, 16 May 1993, BRY! — . . . — BLM Road, S of Poverty Mountain and SW of Salt Spring, T34N, R11W [sic], 5000-5500 ft, Salywon et al. 1107, 21 May 2001, BRY!” [The abstract of this paper erroneously indicates P. pauperitense is from Coconino County, Arizona.] ________________________________________________________________________________ Welsh, Stanley L., and Thorne, K. H. 1977 Plants of Arizona: a new species of Astragalus from the Kaibab Plateau. Great Basin Naturalist, 37(1): 103-104. 313 Astragalus pinionis, NEW SPECIES ; illustrated Trachyophyta: Magnoliopsida: Fabales: Fabaceae (legumes) “Holotype: Arizona, Coconino County, 17 miles southeast of Fredonia Sawmill, along logging road to Ryan, pinyon-juniper-cowania-big sagebrush community, N. D. Atwood 6794, 15 June 1976 (BRY, isotypes to be distributed).” ________________________________________________________________________________ Wheeler, William Morton 1899 New species of Dolichopodidæ from the United States. Proceedings of the California Academy of Sciences, Series 3, Zoology, 2(1), 85 pp. [including 4 plates]. 314  Polymedon castus, NEW SPECIES, pp. 6-8, Pl. 1, fig. 8. Arthropoda: Insecta: Diptera: Dolichopodidae (longlegged flies) “One specimen labelled ‘Grand Cañon, Arizona,’ from the collections of the University of Kansas.” Specimen (holotype) is a female. _______________________________________________________________ 111 BIBLIOGRAPHY AND ANNOTATED CHECKLIST 1906 The ants of the Grand Cañon. Bulletin of the American Museum of Natural History, 22: 329-345. Arthropoda: Insecta: Hymenoptera: Formicidae (ants) Pheidole desertorum, NEW SPECIES (pp. 337-339). [Type material from Fort Davis, Texas.] 315  Pheidole desertorum var. maricopa, NEW VARIETY (pp. 339-340); not illustrated “Described from several soldiers and workers taken from a single colony under a stone in the Grand Cañon just above the Indian Garden [Havasupai Gardens] (3876 feet.)” [Syntypes?] 316  Formica moki, NEW SPECIES (pp. 343-344); not illustrated “Described from 25 workers taken from a small colony under a stone on the wall of the Grand Cañon at an altitude of about 5500 feet and 2 workers taken on the rim in the Kohonino [sic] Forest (7000 feet) about three miles west of the Bright Angel Hotel.” [The “wall of the Grand Cañon” locale is in all probability on Bright Angel Trail.] [Syntypes?] (Note: Wheeler, 1913, p. 343 [see citation below], placed the species Formica moki in Neoformica, NEW SUBGENUS [pp. 400-401].) _______________________________________________________________ 1913 A revision of the ants of the genus Formica (Linné) Mayr. Bulletin of the Museum of Comparative Zoology, 53(10): 379-565. 317  Formica fusca fusca var. gelida, NEW VARIETY (pp. 505-507); not illustrated. [Not valid as a quadrinomial taxon.] Arthropoda: Insecta: Hymenoptera: Formicidae (ants) (“Type locality.—[State of] Colorado: Ward, 9,000 ft. (T. D. A. Cockerell).”) Other material (= paratypes) from numerous, widespread continental localities, including: “Arizona: Coconino Forest, Grand Canyon, 7,000 ft. (Wheeler).” _______________________________________________________________ 1914 New and little known harvesting ants of the genus Pogonomyrmex. Psyche, (October): 149-157. 318  Pogonomyrmex californicus Buckley subsp. maricopa, NEW SUBSPECIES (pp. 155-156); not illustrated Arthropoda: Insecta: Hymenoptera: Formicidae (ants) [Based on syntypes although with the indication, “New Mexico: Alamogordo, type locality (G. von Krockow)”, without further correlation to specimens.] Other material noted from New Mexico, Arizona, California, Texas, and Mexico, including from Arizona “Grand Canyon (Wheeler)”. _______________________________________________________________ 112 BIBLIOGRAPHY AND ANNOTATED CHECKLIST 1915 Some additions to the North American ant-fauna. Bulletin of the American Museum of Natural History, 34: 389-421. 319  Camponotus acutirostris Wheeler clarigaster, NEW VARIETY (p. 420); not illustrated Arthropoda: Insecta: Hymenoptera: Formicidae (ants) “A single specimen [holotype] taken at an altitude of about 3000 ft. on the Bright Angel Trail in the Grand Cañon, Arizona.” ________________________________________________________________________________ Wickham, Henry Frederick 1899 The habits of American Cicindelidae. Proceedings of the Davenport Academy of Natural Sciences, 7: 206-228. 320  Cicindela rufiventris var. arizonae, NEW VARIETY (pp. 226-227); not illustrated Arthropoda: Insecta: Coleoptera: Carabidae (ground beetles) [Syntypes:] “Mr. Roland Hayward sent me specimens labelled ‘Grand Cañon of the Colorado’ and I have since received the same thing collected by Professor Townsend, who writes [underscored for emphasis in this checklist]: ‘I collected the species in the Grand Cañon during the first two weeks of July, 1892. It occurred on sandy places or along paths by a stream going down a side cañon, from 2500 feet above, down to the level of the Colorado River. Or from 5000 feet above sea (at Hance's stone cabin, 2500 feet below the rim and about two or two and a half miles down the trail from river) quite to the bottom of the cañon, which is here 2500 feet above the sea. The insect is not especially wary.’ ” [Regarding the 1892 expedition see a general (non-scientific) account by Townsend (1893), A wagon-trip to the Grand Cañon of the Colorado River, Appalachia, 7(1) (February): 48-63. The locality was on the Old Hance Trail, the upper portion of which was destroyed in 1895 by landslides after which John Hance re-routed his trail down Red Canyon (the present Hance Trail).] ________________________________________________________________________________ Wilcox, J. 1965 New Heteropogon Loew, with a key to the species and the description of a new genus (Diptera: Asilidae). Bulletin of the Southern California Academy of Sciences, 64(4): 207222. 321  Heteropogon stonei, NEW SPECIES (pp. 220-222); not illustrated Arthropoda: Insecta: Diptera: Asilidae (robber flies) (Holotype and allotype from Hualapai Mountains, Arizona.) “Paratypes: . . . one ♀, Grand Canyon, Arizona, 4 June 1963 (J. Wilcox) . . . .” ________________________________________________________________________________ 113 BIBLIOGRAPHY AND ANNOTATED CHECKLIST SERIALS LISTED IN THE BIBLIOGRAPHY AND ANNOTATED CHECKLIST Aliso (Rancho Santa Ana Botanic Garden, Claremont, California) American Journal of Botany American Midland Naturalist American Museum Novitates (American Museum of Natural History) American Naturalist Amphipacifica (Journal of Aquatic Systematic Biology) Annales de Société Entomologique de Belgique Annals of the Carnegie Museum of Natural History Annals of the Entomological Society of America Annals of the New York Academy of Sciences Arquivos do Museu Boçage (Lisboa) Beihefte zur Sydowia (Horn, Niederösterreich, Austria) Botanical Gazette Brigham Young University Science Bulletin (Biological Series) British Journal of Entomology and Natural History Brittonia Bulletin of the American Museum of Natural History Bulletin of the Brooklyn Entomological Society Bulletin of the California Lichen Society Bulletin of the Museum of Comparative Zoology Bulletin of the Southern California Academy of Sciences Bulletin of the Torrey Botanical Club Bulletin of the University of Nebraska State Museum Bulletin of the University of Utah Cactus and Succulent Journal (Cactus and Succulent Society of America) The Canadian Entomologist (London) Cicindela The Coleopterists Bulletin Contributions from the Gray Herbarium of Harvard University Contributions from the U.S. National Herbarium Contributions to Western Botany Copeia 114 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Entomologica Americana Entomological News (Academy of Natural Sciences of Philadelphia, Entomological Section) Fabreries (Association des Entomologistes Amateurs du Québec) Florida Entomologist Garden and Forest Great Basin Naturalist Iowa State College Journal of Science Journal of Arachnology Journal of Eukaryotic Microbiology Journal of Mammalogy Journal of Parasitology Journal of Protozoology Journal of the Arnold Arboretum Journal of the Botanical Research Institute of Texas Journal of the Elisha Mitchell Scientific Society Journal of the Entomological Society of Ontario Journal of the Kansas Entomological Society Journal of the Lepidopterists’ Society Journal of the New York Entomological Society Journal of the Washington Academy of Sciences (Washington, D.C.) Kansas University Science Bulletin Leaflets of Botanical Observation and Criticism Leaflets of Western Botany The Lepidopterists’ News Madroño Minnesota State Entomologist, Report to the Governor Mycologia Mycotaxon The Nautilus (Philadelphia) Notulae Naturae of the Academy of Natural Sciences of Philadelphi a Novon Ohio Journal of Science Pan-Pacific Entomologist PeerJ Phytologia Pittonia Proceedings of the Academy of Natural Sciences of Philadelphia Proceedings of the American Academy of Arts and Sciences 115 BIBLIOGRAPHY AND ANNOTATED CHECKLIST Proceedings of the Biological Society of Washington (Washington, D.C.) Proceedings of the California Academy of Sciences Proceedings of the Davenport Academy of Natural Sciences Proceedings of the Entomological Society of Washington (Washington, D.C.) Proceedings of the Helminthological Society Proceedings of the U.S. National Museum Proceedings of the Utah Academy of Sciences Proceedings of the Washington Academy of Sciences (Washington, D.C.) Psyche Rhodora (Journal of the New England Botanical Club) Scientific Reports (Springer Nature) Southwestern Naturalist Sydowia Systematic Botany Systematic Botany Monographs Transactions of the American Entomological Society Transactions of the American Microscopical Society Transactions of the Illinois State Academy of Science Transactions of the San Diego Society of Natural History University of California Publications in Botany University of California Publications in Entomology University of Kansas Science Bulletin U.S. Department of Agriculture, North American Fauna U.S. Forest Service Agriculture Handbook U.S. Geological and Geographical Survey of the Territories, Bulletin Werdenda (Beiträge zur Pflanzenkunde) (Bingen, Washington) Western North American Naturalist ZooKeys Zootaxa 116 TAXONOMIC LIST TAXONOMIC LIST This is a nomenclatural record. Taxa are as named by the original authors; later taxonomic reassignments, if any, are not generally noted herein. There are 322 taxa first named from collections made in the greater Grand Canyon region of northwestern Arizona, 215 of which have type material that is wholly or partly from within, or with good proabability within, the present boundaries of Grand Canyon National Park. [No. 244a was added late, thus the whole-number count stops at 321.] Higher systematics for the original genus are given here, as derived from the Integrated Taxonomic Information System (ITIS) online in 2022. Taxa not included in ITIS have had their higher systematics derived from other sources, including the National Center for Biotechnology Information Taxonomy Browser (National Library of Medicine) and the World Flora Online. The systematic placements listed here may not reflect the most current views, which are in any case bound to vary between researchers and over time. Again, this is a resource manager’s historical checklist to nominate taxa, not one of systematics or revised nomenclature. The taxonomic ranks cited in this list are: Phylum or Division: Class: Order: Family. The taxa in this list are not in so-called “systematic order” because some users of the checklist will not be accustomed to the methodical order of taxa within Phyla, Classes, and so forth. To accommodate these users, who are more likely to be resource managers and historically inclined users who are not necessarily scientists, all higher taxa are listed strictly in alphabetical order (the order of Kingdoms is arranged only for convenience based on the relative number of entries). The nominate taxa within each category are listed in the order in which they appear in the checklist, arranged numerically to facilitate finding them in the bibliography/ checklist. Common names are mostly as listed by ITIS or other sources. They should not be taken to be names regularly adopted by all users, who will understand other common names for these taxa. The names have been added here only a matter of convenience for those users, principally non-scientists, who prefer or need to refer in some way to common names.  Arrow indicates taxa for which at least one locality reported for its types is within, or is with good probability from within, the present boundaries of Grand Canyon National Park. Refer to the bibliography for all details. Red Numbers along the left margin of the Taxonomic List indicate and cross-list the sequentially enumerated taxa in the bibliography (they reflect the alphabetical order by publication author). 117 TAXONOMIC LIST TOTAL NUMBERS OF NOMINATE TAXA / NO. FROM GRAND CANYON NATIONAL PARK [In the PDF all lines below are hyperlinks to those categories in the Taxonomic List] Kingdom Animalia (animals) Phylum Arthropoda (arthropods) Total GRCA Page 172 133 119 141 107 119 Class Branchiopoda (branchiopod crustaceans) 1 119 Class Diplopoda (millipedes) 1 119 Class Euchelicerata (spiders/mites/scorpions/etc.) 14 7 119 124 99 121 1 1 128 18 14 128 1 1 128 13 9 128 Class Reptilia (reptiles) 3 3 129 Class Teleostei (fishes) 1 1 130 Class Insecta (insects) Class Malacostraca (malacostracan crustaceans) Phylum Chordata (vertebrates) Class Aves (birds) Class Mammalia (mammals) Phylum Mollusca (mollusks) 130 Class Gastropoda (snails) 9 7 130 4 5 131 111 49 132 110 48 132 1 1 137 27 22 138 23 20 138 4 2 139 Kingdom Chromista (photosynthetic eukaryotes) 11 11 140 Phylum Apicomplexa (parasitic alveolates) 11 11 140 Phylum Nematoda (roundworms) Class Chromadorea (chromadorid roundworms) Kingdom Plantae (plants) Phylum (Division) Trachyophyta (vascular plants) Class Magnoliopsida (flowering plants) Class Pinopsida (conifers) Kingdom Fungi (fungi) Phylum (Division) Ascomycota (sac fungi) Phylum (Division) Basidiomycota (club fungi) Kingdom Protista (protists) “Phylum” Hemimastigophora (single-celled eukaryotes) 118 1 140 1 140 TAXONOMIC LIST KINGDOM ANIMALIA (animals) Phylum ARTHROPODA (arthropods) Class Branchiopoda (branchiopod crustaceans) Arthropoda: Branchiopoda: Anostraca: Branchinectidae (brine shrimp) 48 Branchinecta kaibabensis Belk & Fugate, 2000 ________________________________________________________________________________ Class Diplopoda (millipedes) Arthopoda: Diplopoda: Polydesmida: Macrosternodesmidae (presumed troglobitic polydesmidan millipeds) 272 Pratherodesmus voylesi Shear in Shear, Taylor, Wynne & Krejca, 2009 ________________________________________________________________________________ Class Euchelicerata (spiders/mites/scorpions/etc.) Arthropoda: Euchelicerata: Areneae: Hahniidae (hahniid spiders) 71 Neoantistea coconino Chamberlin & Ivie, 1942 ________________________________________________________________________________ Arthropoda: Euchelicerata: Areneae: Linyphiidae (dwarf weavers) 70 Tapinocyba kesimba Chamberlin, 1948 ________________________________________________________________________________ Arthropoda: Euchelicerata: Areneae: Lycosidae (wolf spiders) 114  Arctosa mokiensis Gertsch, 1934a ________________________________________________________________________________ Arthropoda: Euchelicerata: Areneae: Salticidae (jumping spiders) 115 Phidippus kaibabensis Gertsch, 1934b ________________________________________________________________________________ 119 TAXONOMIC LIST Arthropoda: Euchelicerata: Areneae: Sicariidae (recluse spiders) 117  Loxosceles kaiba Gertsch & Ennik, 1983 ________________________________________________________________________________ Arthropoda: Euchelicerata: Areneae: Theraphosidae (tarantulas) 68 69  Aphonopelma behlei Chamberlin, 1940  Aphonopelma phasmus Chamberlin, 1940 ________________________________________________________________________________ Arthropoda: Euchelicerata: Areneae: Theridiidae (cobweb weavers) 116 Steatoda variata Gertsch, 1960 ________________________________________________________________________________ Arthropoda: Euchelicerata: Areneae: Thomisidae (crab spiders) 113  Misumenops coloradensis Gertsch, 1933 ________________________________________________________________________________ Arthropoda: Euchelicerata: Pseudoscorpiones (pseudoscorpions): Chernetidae 153 154 Hesperochernes bradybaughi Harvey & Wynne, 2014 Tuberochernes cohni Harvey & Wynne, 2014 ________________________________________________________________________________ Arthropoda: Euchelicerata: Pseudoscorpiones (pseudoscorpions): Larcidae 225  Archeolarca cavicola Muchmore, 1981 ________________________________________________________________________________ Arthropoda: Euchelicerata: Scorpiones: Vaejovidae (vaejovid scorpions) 277 Vaejovis aquilonalis Stahnke, 1940 ________________________________________________________________________________ Arthropoda: Euchelicerata: Trombidiformes: Tarsonemidae (tarsonemid mites) 275  Heterotarsonemus nakaharai Smiley & Moser, 1974 ________________________________________________________________________________ 120 TAXONOMIC LIST Class Insecta (insects) Arthropoda: Insecta: Blattodea: Polyphagidae (sand cockroaches) 166 167 168  Arenivaga grandiscanyonensis Hopkins, 2014  Arenivaga impensa Hopkins, 2014  Arenivaga pagana Hopkins, 2014 ________________________________________________________________________________ Arthropoda: Insecta: Coleoptera: Buprestidae (jewel beetles) 44 55 228  Acmaeodera pletura W. Barr, 1972  Melanophila piniedulis Burke, 1907 Acmaeodera navajo G. Nelson & Westcott, 1995 ________________________________________________________________________________ Arthropoda: Insecta: Coleoptera: Carabidae (ground beetles) 67 172 181 320  Bembidion (Cyclolopha) occultum Casey, 1918 Cicindela pusilla kaibabensis Johnson, 1990  Nebria (Reductonebria) georgei Kavanaugh, 2008  Cicindela rufiventris var. arizonae Wickham, 1899 ________________________________________________________________________________ Arthropoda: Insecta: Coleoptera: Cerambycidae (long-horned beetles) 63 64 65 66     Prionus spiculosus Casey, 1912 Prionus angustulus Casey, 1912 Prionus terminalis Casey, 1912 Stenosphenus pruddeni Casey, 1912 ________________________________________________________________________________ Arthropoda: Insecta: Coleoptera: Chrysomelidae (leaf beetles) 105 169 223  Bruchus perplexus Fall, 1910 Luperodes wickhami Horn, 1893 Saxinis saucia kaibabiae Moldenke, 1970 ________________________________________________________________________________ Arthropoda: Insecta: Coleoptera: Dermestide (dermestid beetles) 58 59  Cryptorhopalum pruddeni Casey, 1900  Orphilus aequalis Casey, 1900 ________________________________________________________________________________ 121 TAXONOMIC LIST Arthropoda: Insecta: Coleoptera: Elateridae (click beetles) 196 Aphricus knowltoni Knull, 1957 ________________________________________________________________________________ Arthropoda: Insecta: Coleoptera: Leiodidae (round fungus beetles) 232 233  Ptomaphagus cocytus Peck, 1973 Ptomaphagus parashant Peck & Wynne, 2013 ________________________________________________________________________________ Arthropoda: Insecta: Coleoptera: Melyridae (soft-winged flower beetles) 56 57 Trichochrous incipiens Casey, 1895 Trichochrous reversus Casey, 1895 ________________________________________________________________________________ Arthropoda: Insecta: Coleoptera: Staphylinidae (rove beetles) 295  Eleodes (Caverneleodes) leptoscelis Triplehorn, 1975 ________________________________________________________________________________ Arthropoda: Insecta: Coleoptera: Tenebrionidae (darkling beetles) 60 61 62 173 230 296  Zopherodes lugubris Casey, 1907  Zopherodes pruddeni Casey, 1907  Discodemus brevipennis Casey, 1908 Trogloderus skillmani Johnston, 2019  Eschatomoxys pholeter Thomas & Pape in Pape, Thomas & Aalbu, 2007  Asidina rugicollis Triplehorn & Brown, 1971 ________________________________________________________________________________ Arthropoda: Insecta: Coleoptera: Ptinidae (spider beetles) 100 101 102 103     Petalium bistriatum Say var. arizonense Fall, 1905 Catorama grande Fall, 1905 Catorama longulum Fall, 1905 Catorama pingue Fall, 1905 ________________________________________________________________________________ Arthropoda: Insecta: Coleoptera: Scarabaeidae (scarab beetles) 104  Diplotaxis conformis Fall, 1909 ________________________________________________________________________________ 122 TAXONOMIC LIST Arthropoda: Insecta: Diptera: Asilidae (robber flies) 53 264 321  Erax benedicti Bromley, 1940  Efferia tapeats Scarbrough, Stevens & Nelson, 2012  Heteropogon stonei Wilcox, 1965 ________________________________________________________________________________ Arthropoda: Insecta: Diptera: Bombyliidae (bee flies) 85 152  Bombylius facialis Cresson, 1919  Geron prosopidis Hall & Evenhuis, 2003 ________________________________________________________________________________ Arthropoda: Insecta: Diptera: Ceratopogonidae (biting midges) 1  Ceratopogon dimidiatus C. Adams, 1903 ________________________________________________________________________________ Arthropoda: Insecta: Diptera: Dolichopodidae (longlegged flies) 256 300 301 314     Chrysotimus arizonicus H. Robinson, 1967 Polymedon dilaticosta Van Duzee, 1927 Polymedon nitidus Van Duzee, 1927 Polymedon castus Wheeler, 1899 ________________________________________________________________________________ Arthropoda: Insecta: Diptera: Empididae (balloon flies) 273  Wiedemannia digna Sinclair, 2006 ________________________________________________________________________________ Arthropoda: Insecta: Diptera: Mycetophilidae (fungus gnats) 72 271  Mycetophila neofungorum Chandler, 1993  Mycetophila denningi Shaw, 1951 ________________________________________________________________________________ Arthropoda: Insecta: Diptera: Tachinidae (tachinid flies) 249 294 Fabriciella evanida Reinhard, 1953  Microsciasma minuta Townsend, 1915 ________________________________________________________________________________ Arthropoda: Insecta: Diptera: Tephritidae (fruit flies) 81  Trypeta varipennis Coquillett, 1902 ________________________________________________________________________________ 123 TAXONOMIC LIST Arthropoda: Insecta: Diptera: Tipulidae (crane flies) 3  Tipula (Lunatipula) kaibabensis C. Alexander, 1946 ________________________________________________________________________________ Arthropoda: Insecta: Hemiptera: Aphididae (aphids) 170 171  Cinara grande Hottes, 1956 Cinara poketa Hottes, 1956 ________________________________________________________________________________ Arthropoda: Insecta: Hemiptera: Cicadellidae (leafhoppers) 14 15 16 88 212      Athysanella fredonia Ball & Beamer, 1939 Athysanella globosa Ball & Beamer, 1939 Athysanella (Gladionura) casa Ball & Beamer, 1939 Ballana basala DeLong, 1964 Norvellina bicolorata (Ball) var. inflata Lindsay, 1938 ________________________________________________________________________________ Arthropoda: Insecta: Hemiptera: Cicadidae (cicadas) 87  Tibicen apache Davis, 1921 ________________________________________________________________________________ Arthropoda: Insecta: Hemiptera: Cixiidae (planthoppers) 11 Myndus yuccandus E. Ball, 1933 ________________________________________________________________________________ Arthropoda: Insecta: Hemiptera: Caliscelidae (planthoppers) 54  Aphelonema convergens var. canyonensia Bunn, 1930 ________________________________________________________________________________ Arthropoda: Insecta: Hemiptera: Coccidae (soft scales) 95 96  Ripersia arizonensis Ehrhorn, 1899  Dactylopius formicarii Ehrhorn 1899 ________________________________________________________________________________ Arthropoda: Insecta: Hemiptera: Coreidae (leaf-footed bugs) 216  Chelinidea vittiger Uhler var. artuflava McAtee, 1919 ________________________________________________________________________________ Arthropoda: Insecta: Hemiptera: Delphacidae (planthoppers) 47  Delphacodes apicata Beamer, 1948 ________________________________________________________________________________ 124 TAXONOMIC LIST Arthropoda: Insecta: Hemiptera: Diaspididae (armored scales) 213  Chionaspis gilli Liu & Kosztarab, 1987 ________________________________________________________________________________ Arthropoda: Insecta: Hemiptera: Dictyopharidae (planthoppers) 13 Yucanda miniata E. Ball, 1937 ________________________________________________________________________________ Arthropoda: Insecta: Hemiptera: Issidae (planthoppers) 12 90 Hysteropterum cornutum var. utahnum E. Ball, 1935  Bruchomorpha bunni Doering, 1939 ________________________________________________________________________________ Arthropoda: Insecta: Hemiptera: Miridae (jumping tree bugs) 184 185 186 187 188 189 190 191 192 193 194 195 241 242 299           Deraeocoris fulvus Knight, 1921 Deraeocoris bullatus Knight, 1921 Deraeocoris navajo Knight, 1921 Ceratocapsus clavicornis Knight, 1925 Phytocoris mellarius Knight, 1925 Pilophorus fuscipennis Knight, 1926 Phytocoris hesperius Knight, 1928 Plagiognathus tenellus Knight, 1929 Phytocoris varius Knight, 1934 Parthenicus cowaniae Knight, 1968 Bolteria juniperi Knight, 1968     Phytocoris flaviatus Knight, 1968 Pilophorus americanus Poppius, 1914 Pilophorus exiguus Poppius, 1914 Platylygus vanduzeei Usinger, 1931 ________________________________________________________________________________ Arthropoda: Insecta: Hemiptera: Ochteridae (velvety shore bugs) 265  Ochterus barberi Schell, 1943 ________________________________________________________________________________ Arthropoda: Insecta: Hemiptera: Psyllidae (jumping plantlice) 298  Trioza rubra Tuthill, 1939 ________________________________________________________________________________ Arthropoda: Insecta: Hemiptera: Reduviidae (assassin bugs) 34  Paratriatoma hirsuta Barber, 1938 ________________________________________________________________________________ 125 TAXONOMIC LIST Arthropoda: Insecta: Hymenoptera: Andrenidae (andrenid bees) 200 253 290 291 292 293 Andrena (Callandrena) utahensis LaBerge, 1967  Andrena (Scaphandrena) kaibabensis Ribble, 1974  Perdita wheeleri Timberlake, 1928 Perdita opacella Timberlake, 1956  Perdita inornata Timberlake, 1962  Perdita depressa Timberlake, 1968 ________________________________________________________________________________ Arthropoda: Insecta: Hymenoptera: Apidae (honey bees) 79  Triepeolus hopkinsi Cockerell, 1905 ________________________________________________________________________________ Arthropoda: Insecta: Hymenoptera: Crabronidae (crabronid wasps) 111  Gorytes dentatus Fox, 1893 ________________________________________________________________________________ Arthropoda: Insecta: Hymenoptera: Cynipidae (gall wasps) 51  Andricus wheeleri Beutenmüller, 1907 ________________________________________________________________________________ Arthropoda: Insecta: Hymenoptera: Formicidae (ants) 80 Myrmecocystus melliger subsp. semirufus Emery var. romainei Cole, 1936 [not valid as a quadrionomial taxon, and no identification of holotype; non Myrmecocstus romainei Snelling, 1975] 315 316 317 318 319      Pheidole desertorum var. maricopa Wheeler, 1906 Formica moki Wheeler, 1906 Formica fusca fusca var. gelida Wheeler, 1913 [not valid as a quadrinomial taxon] Pogonomyrmex californicus Buckley subsp. maricopa Wheeler, 1914 Camponotus acutirostris Wheeler clarigaster Wheeler, 1915 ________________________________________________________________________________ Arthropoda: Insecta: Hymenoptera: Pteromalidae (pteromalid wasps) 150 Boharticus margaretae Grissell, 1983 ________________________________________________________________________________ Arthropoda: Insecta: Hymenoptera: Sphecidae (sphecid wasps) 52  Priononyx subatrata Bohart, 1958 ________________________________________________________________________________ 126 TAXONOMIC LIST Arthropoda: Insecta: Hymenoptera: Vespidae (hornets) 50 155  Mischocyttarus flavitarsis var. kaibabensis Bequaert, 1932  Polistes canadensis var. kaibabensis Lynn, 1932 ________________________________________________________________________________ Arthropoda: Insecta: Lepidoptera: Hesperiidae (skippers) 278 Megathymus alliae Stallings & Turner, 1957 ________________________________________________________________________________ Arthropoda: Insecta: Lepidoptera: Noctuidae (owlet moths) 2 240 297 Plagiomimicus kathyae J. Adams & Lafontaine, 2009  Schinia immaculata Pogue, 2004 Hadena (Hadena) lafontainei Troubridge & Crabo, 2002 ________________________________________________________________________________ Arthropoda: Insecta: Lepidoptera: Nymphalidae (admirals) 40 41 112 226 268      Coenonympha fureae Barnes & Benjamin, 1926 Cercyonis damei Barnes & Benjamin, 1926 Speyeria atlantis Edwards schellbachi Garth, 1949 Neonympha dorothea dorothea Nabokov, 1942 Neominois dionysus Scudder, 1878 ________________________________________________________________________________ Arthropoda: Insecta: Lepidoptera: Oecophoridae (oecophorid moths) 76  Depressaria schellbachi Gates, 1947 ________________________________________________________________________________ Arthropoda: Insecta: Lepidoptera: Papilionidae (swallowtails) 45  Papilio indra kaibabensis Bauer, 1955 ________________________________________________________________________________ Arthropoda: Insecta: Lepidoptera: Pieridae (orange-tips) 269 Synchloe thoosa Scudder, 1878 ________________________________________________________________________________ Arthropoda: Insecta: Orthoptera: Acrididae (short-horned grasshoppers) 270  Melanoplus canonicus Scudder, 1897 ________________________________________________________________________________ 127 TAXONOMIC LIST Arthropoda: Insecta: Orthoptera: Eumastacidae (monkey grasshoppers) 248 Morsea california kaibabensis Rehn & Grant, 1958 ________________________________________________________________________________ Arthropoda: Insecta: Psocodea: Enderleinellidae (sucking lice) 182 Enderleinellus kaibabensis Kim, 1966 ________________________________________________________________________________ Arthropoda: Insecta: Siphonaptera: Ceratophyllidae (rodent fleas) 9  Orchopeas sexdentatus neotomae Augustson, 1943 ________________________________________________________________________________ Class Malacostraca (malacostracan crustaceans) Arthropoda: Malacostraca: Amphipoda: Crangonyctidae (crangonyctid amphipods) 302  Stygobromus blinni Wang & Holsinger, 2001 ________________________________________________________________________________ Phylum CHORDATA Class Aves (birds) Chordata: Aves: Strigiformes: Strigidae (typical owls) 229  Otus asio mychophilus Oberholser, 1937 ________________________________________________________________________________ Class Mammalia (mammals) Chordata: Mammalia: Carnivora: Felidae (cats) 227  Felis concolor kaibabensis E. Nelson & Goldman, 1931 ________________________________________________________________________________ Chordata: Mammalia: Carnivora: Mephitidae (skunks) 219  Spilogale gracilis Merriam, 1890 ________________________________________________________________________________ 128 TAXONOMIC LIST Chordata: Mammalia: Rodentia: Castoridae (beavers) 121  Castor canadensis repentinus Goldman, 1932 ________________________________________________________________________________ Chordata: Mammalia: Rodentia: Crecitidae (deer mice) 122 218  Peromyscus crinitus peridoneus Goldman, 1937  Hesperomys megalotis Merriam, 1890 ________________________________________________________________________________ Chordata: Mammalia: Rodentia: Geomyidae (pocket gophers) 91 123 124 151  Thomomys bottae boreorarius Durham, 1952 Thomomys fossor kaibabensis Goldman, 1938 Thomomys bottae nicholi Goldman, 1938  Thomomys bottae trumbullensis Hall & Davis, 1934 ________________________________________________________________________________ Chordata: Mammalia: Rodentia: Heteromyidae (kangaroo rats) 118 119 Dipodomys microps celsus Goldman, 1924 Dipodomys ordii cupidineus Goldman, 1924 ________________________________________________________________________________ Chordata: Mammalia: Rodentia: Sciuridae (squirrels) 120 220  Ammospermophilus leucurus tersus Goldman, 1929  Sciurus kaibabensis Merriam, 1904 ________________________________________________________________________________ Class Reptilia (reptiles) Chordata: Reptilia: Squamata: Colubridae (typical snakes) 281 285  Sonora semiannulata gloydi Stickel, 1938  Lampropeltis pyromelana Cope infralabialis Tanner, 1953 ________________________________________________________________________________ Chordata: Reptilia: Squamata: Viparidae (rattlesnakes) 183  Crotalus confluentus abyssus Klauber, 1930 ________________________________________________________________________________ 129 TAXONOMIC LIST Class Teleostei (fishes) Chordata: Teleostei: Cypriniformes: Cyprinidae (carp) 221  Gila cypha R. Miller, 1946 ________________________________________________________________________________ Phylum MOLLUSCA (mollusks) Class Gastropoda (snails) Mollusca: Gastropoda: Neotaenioglossa: Hydrobiidae (spring snails) 157 Pyrgulopsis hualapaiensis Herschler, Liu & Stevens, 2016 ________________________________________________________________________________ Mollusca: Gastropoda: Stylommatophora: Helminthoglyptidae (shoulderbands) 156  Sonorella betheli Henderson, 1914 [an erroneous account; see details in the bibliography herein] 222 280  Sonorella reederi W. Miller, 1984  Helix (Arionta) coloradoensis Stearns, 1890 ________________________________________________________________________________ Mollusca: Gastropoda: Stylommatophora: Oreohelicidae (mountain snails) 214 236 237 238 Oreohelix yavapai vauxae Marshall, 1929  Oreohelix yavapai profundorum Pilsbry & Ferriss, 1911  Oreohelix yavapai extremitatis Pilsbry & Ferriss, 1911  Oreohelix yavapai angelica Pilsbry & Ferriss, 1911 ________________________________________________________________________________ Mollusca: Gastropoda: Stylommatophora: Pupillidae (chrysalis snails) 239  Pupilla syngenes avus Pilsbry & Ferriss, 1911 ________________________________________________________________________________ 130 TAXONOMIC LIST Phylum NEMATODA (roundworms) Class Chromadorea (chromadorid roundworms) Nematoda: Chromadorea: Ascaridida: Oxyuridae (pinworms) 197 198  Wellcomia perognathi Kruidenier & Mehra, 1958  Aspiculuris ackerti Kruidenier & Mehra, 1959 ________________________________________________________________________________ Nematoda: Chromadorea: Rhabditida : Rhabditidae (rhabditid roundworms) 215  Parasitorhabditis gracilis Massey, 1974 ________________________________________________________________________________ Nematoda: Chromadorea: Spiurida: Gongylonematidae (spiurid gullet worms) 199  Gongylonema peromysci Kruidenier & Peebles, 1958 ________________________________________________________________________________ 131 TAXONOMIC LIST KINGDOM PLANTAE (plants) Phylum (Division) TRACHYOPHYTA (vascular plants) Class Magnoliopsida (flowering plants) Trachyophyta: Magnoliopsida: Asparagales: Asparagaceae (agaves) 162 217 309  Agave phillipsiana Hodgson, 2001  Agave kaibabensis McKelvey, 1949 Androstephium breviflorum Watson, 1873 ________________________________________________________________________________ Trachyophyta: Magnoliopsida: Asterales: Asteraceae (sunflowers) 8 75 129 147 177 178 286 307 Xylorhiza tortifolia (Torrey & Gray) Greene var. parashantensis Atwood & Welsh, 2013  Encelia resinifera subsp. tenuifolia Curtis, 1998 Actinella biennis A. Gray, 1878 Senecio stygius Greene, 1909 Laphamia congesta Jones, 1895 Laphamia gracilis Jones, 1895  Flaveria mcdougallii Theroux, Pinkava & Keil, 1977 Brickellia (Clavigera) longifolia Watson, 1873 ________________________________________________________________________________ Trachyophyta: Magnoliopsida: Boraginales: Boraginaceae (borage) 94 160 283 284  Oreocarya capitata Eastwood, 1937 Cryptantha atwoodii Higgins, 1974  Amsinckia nana Suksdorf, 1931  Amsinckia macrosepala Suksdorf, 1931 ________________________________________________________________________________ Trachyophyta: Magnoliopsida: Boraginales: Hydrophyllaceae (waterleafs) 5 6 7 131 Phacelia furnissii Atwood, 2007 Phacelia higginsii Atwood, 2007 Phacelia hughesii Atwood, 2007 Phacelia (Coreanthus) ivesiana A. Gray in Gray et al., 1861 ________________________________________________________________________________ 132 TAXONOMIC LIST Trachyophyta: Magnoliopsida: Brassicales: Apiaceae (parsleys) 83 305 306 Phellopterus multinervatus Coulter & Rose, 1900 Cymopterus purpureus Watson, 1873 Peucedanum newberryi Watson, 1873 ________________________________________________________________________________ Trachyophyta: Magnoliopsida: Brassicales: Brassicaceae [Cruciferae] (mustards) 4 134 135 257 258 310  Thelypodium integrifolium (Nuttall) Endlicher subsp. longicarpum Al-Shehbaz, 1973 Arabis eremophila Greene, 1900 Arabis recondita Greene, 1900  Sisymbrium kearneyi Rollins, 1953 Lesquerella kaibabensis Rollins, 1982 Lesquerella arizonica Watson, 1888 ________________________________________________________________________________ Trachyophyta: Magnoliopsida: Caryophyllales: Amaranthaceae (pigweed) 282  Atriplex asterocarpa Stutz, Chu & Sanderson, 1994 ________________________________________________________________________________ Trachyophyta: Magnoliopsida: Caryophyllales: Cactaceae (cacti) 46 49 77 78 82 108 109 158 159 231 Opuntia aurea Baxter, 1933 Pediocactus paradinei Benson, 1957  Sclerocactus havasupaiensis Clover, 1942  Sclerocactus havasupaiensis var. roseus Clover, 1942 Echinocactus polycephalus var. xeranthemoides Coulter, 1896 Coryphantha vivipara var. kaibabensis Fischer, 1979  Echinocereus triglochidiatus var. toroweapensis Fischer, 1991 Opuntia hualpaensis Hester, 1943 Opuntia abyssi Hester, 1943  Opuntia pinkavae Parfitt, 1997 ________________________________________________________________________________ Trachyophyta: Magnoliopsida: Caryophyllales: Caryophyllaceae (pinks) 254  Silene rectiramea Robinson, 1899 ________________________________________________________________________________ Trachyophyta: Magnoliopsida: Caryophyllales: Nyctaginaceae (four o’clocks) 130 279 Mirabilis bigelovii A. Gray, 1886  Allionia pachyphylla Standley, 1909 ________________________________________________________________________________ 133 TAXONOMIC LIST Trachyophyta: Magnoliopsida: Caryophyllales: Polygonaceae (knotweeds) 250 251 274 Eriogonum mortonianum Reveal, 1974a Eriogonum thompsonae S. Wats. var. atwoodii Reveal, 1974b Eriogonum fusiforme Small, 1906 ________________________________________________________________________________ Trachyophyta: Magnoliopsida: Cornales: Losaceae (loasa) 73 165 266 267 Mentzelia collomiae Christy, 1997 Mentzelia memorabilis N. Holmgren & P. Holmgren, 2002  Mentzelia hualapaiensis Schenk, Hodgson & Hufford, 2010  Mentzelia canyonensis Schenk, Hodgson & Hufford, 2013 ________________________________________________________________________________ Trachyophyta: Magnoliopsida: Crossosomatales: Crossosomataceae (crossomas) 255  Crossosoma parviflora B. Robinson & Fernald, 1894 ________________________________________________________________________________ Trachyophyta: Magnoliopsida: Ericales: Polemoniaceae (phlox) “Gilia dactylophyllum, (n. sp.?)”, A. Gray in Gray et al., 1861 [see note in bibliography herein] ________________________________________________________________________________ 132 Trachyophyta: Magnoliopsida: Ericales: Primulaceae (primroses)  Primula (Section Farinosae) hunnewellii Fernald, 1934 ________________________________________________________________________________ 106 Trachyophyta: Magnoliopsida: Fabales: Fabaceae (legumes) 35 36 37 38 39 125 126 127 128 174 179 180 261 303 311 312 313  Astragalus bryantii Barneby, 1944  Astragalus lentiginosus Dougl. var. oropedii Barneby, 1945  Astragalus cremnophylax Barneby, 1948 Astragalus wootoni var. endopterus Barneby, 1949 Astragalus cremnophylax Barneby var. myriorrhaphis Barneby, 1979 Astragalus subcinereus A. Gray, 1878 Astragalus scaposus A. Gray, 1878 Astragalus mokiacensis A. Gray, 1878 Astragalus artipes A. Gray, 1878 Astragalus humistratus var. tenerrimus Jones, 1895 Astragalus kentrophyta var. coloradoensis Jones, 1902 Astragalus kaibensis Jones, 1902 Pediomelum retrorsum Rydberg in Rydberg & Pennell, 1919 Dalea amoena Watson, 1873 Astragalus lentiginosus Douglas ex Hook. var. trumbullensis Welsh & Atwood, 2001 Pediomelum pauperitense S.L. Welsh, M. Licher, & N. D. Atwood in Welsh & Licher, 2010 Astragalus pinionis Welsh & Thorne, 1977 ________________________________________________________________________________ 134 TAXONOMIC LIST Trachyophyta: Magnoliopsida: Fagales: Betulaceae (birches) 84  Ostrya knowltonii Coville, 1894 ________________________________________________________________________________ Trachyophyta: Magnoliopsida: Gentianales: Apocynaceae (dogbane) 260 Amsonia eastwoodiana Rydberg, 1913 ________________________________________________________________________________ Trachyophyta: Magnoliopsida: Gentianales: Rubiaceae (madders) 89  Galium munzii Hilend & Howell subsp. ambivalens Dempster & Ehrendorfer, 1965 ________________________________________________________________________________ Trachyophyta: Magnoliopsida: Lamiales: Lamiaceae (mints) 252 Scutellaria potosina var. kaibabensis Rhodes & Ayers, 2010 ________________________________________________________________________________ Trachyophyta: Magnoliopsida: Lamiales: Oleaceae (olives) 93 176 246 247  Fraxinus macropetala Eastwood, 1903 Bigelovia howardi var. attenuata Jones, 1895  Fraxinus cuspidata var. macropetala Rehder, 1917  Fraxinus lowellii Sargent in Rehder, 1917 ________________________________________________________________________________ Trachyophyta: Magnoliopsida: Lamiales: Orobanchaceae (broomrapes) 74 107 234 Cordylanthus wrightii kaibabensis Chuang & Heckard, 1986  Adenostegia parviflora Ferriss, 1918  Cordylanthus tenuifolis Pennell, 1940 ________________________________________________________________________________ Trachyophyta: Magnoliopsida: Lamiales: Plantaginaceae (plantains) 86 259 Penstemon virgatus pseudoptutus Crosswhite, 1967  Penstemon coccinatus Rydberg, 1909 ________________________________________________________________________________ Trachyophyta: Magnoliopsida: Lamiales: Scrophulariaceae (figworts) 164 Castilleja kaibabensis N. Holmgren, 1973 ________________________________________________________________________________ Trachyophyta: Magnoliopsida: Liliales: Liliaceae (lilies) 308 Calochortus flexuosus Watson, 1873 ________________________________________________________________________________ 135 TAXONOMIC LIST Trachyophyta: Magnoliopsida: Malpighiales: Euphorbiaceae (spurge) 163  Euphorbia aaronrossii A. Holmgren & N. Holmgren, 1988 ________________________________________________________________________________ Trachyophyta: Magnoliopsida: Malpighiales: Salicaceae (willows) 10 263  Salix lutea var. ligulifolia C. Ball, 1921  Populus fremontii var. toumeyi Sargent, 1919 ________________________________________________________________________________ Trachyophyta: Magnoliopsida: Myrtales: Onagraceae (evening primroses) 175 243 244 244a 304 Oenothera triloba var. ecristata Jones, 1895  Oenothera confertiflora Raven, 1962  Oenothera specuicola subsp. specuicola Raven, 1962 Oenothera specuicola subsp. hesperia Raven, 1962 Oenothera (Chylisma) multijuga Watson, 1873 ________________________________________________________________________________ Trachyophyta: Magnoliopsida: Poales: Cyperaceae (sedges) 276  Carex curatorum Stacey, 1937 ________________________________________________________________________________ Trachyophyta: Magnoliopsida: Poales: Poaceae (grasses) 245 Panicum mohavense Reeder, 1991 ________________________________________________________________________________ Trachyophyta: Magnoliopsida: Ranunculales: Papaveraceae (poppies) 92  Corydalis wetherillii Eastwood, 1902 ________________________________________________________________________________ Trachyophyta: Magnoliopsida: Rosales: Moraceae (mulberries) 148 Morus grisea Greene, 1911 ________________________________________________________________________________ Trachyophyta: Magnoliopsida: Rosales: Roasaceae (roses) 235  Rosa stellata Wooton subsp. abyssa Phillips, 1992 ________________________________________________________________________________ 136 TAXONOMIC LIST Trachyophyta: Magnoliopsida: Sapindales: Anacardiaceae 133 136 137 (cashews)  Rhus canadensis Marsh. var. simplicifolia Greene, 1890  Schmaltzia cissodes Greene, 1905  Schmaltzia hirtella Green, 1905 ________________________________________________________________________________ Trachyophyta: Magnoliopsida: Sapindales: Rutaceae (rues) 138 139 140 141 142 143 144 145 146 Ptelea palida Green, 1906 Ptelea straminea Green, 1906       Ptelea nitida Green, 1906 Ptelea argentea Green, 1906 Ptelea triptera Green, 1906 Ptelea lutescens Green, 1906 Ptelea elegans Green, 1906 Ptelea confinis Green, 1906 Ptelea saligna Green, 1906 ________________________________________________________________________________ Class Pinopsida (conifers) Trachyophyta: Pinopsida: Pinales: Cupressaceae (cypress) 262  Juniperus utahensis var. megalocarpa Sargent, 1919 ________________________________________________________________________________ 137 TAXONOMIC LIST KINGDOM FUNGI (fungi) Phylum (Division) ASCOMYCOTA (sac fungi) Ascomycota: Dothidiomycetes: Pleosporales: Venturiaceae 43  Protoventuria arizonica M. Barr, 1989 ________________________________________________________________________________ Ascomycota: Eurotiomycetes: Eurotiales: Trichocomaceae (saprobe fungi) 149  Penicillium arizonense Frisvad, Grijseels & J.C. Nielsen in Grijseels et al., 2016 ________________________________________________________________________________ Ascomycota: Lecanoromycetes: Teloschistales: Physciaceae 99  Physconia isidiomuscigena Esslinger, 2000 ________________________________________________________________________________ Ascomycota: Orbiliomycetes: Orbiliales: Orbiliaceae 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33  Orbilia purshiae Baral & G. Marson in Baral, Weber & Marson, 2020  Orbilia ocellata Baral, G. Marson & E. Weber in Baral, Weber & Marson, 2020 Orbilia magnifica Baral & G. Marson in Baral, Weber & Marson, 2020        Orbilia phanosoma Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia multiphanosoma Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia arizonensis Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia navajoana Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia spermoides Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia ophiosoma Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia maeandrina Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia multimaeandrina Baral & G. Marson in Baral, Weber & Marson, 2020       Orbilia flexisoma Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia delphinus Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia macrodelphinus Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia multitrapezoidea Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia calyptrata Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia subovoidea Baral, Matočec & E. Weber in Baral, Weber & Marson, 2020 ________________________________________________________________________________ 138 TAXONOMIC LIST Ascomycota: Pezizomycetes: Pezizales: Morchellaceae 42 Morchella kaibabensis Beug, T.A. Clem. & T.J. Baroni in Baroni et al., 2018 ________________________________________________________________________________ Ascomycota: Sordariomycetes: Diaporthales: Valsaceae 224  Gnomonia quercusgambellii Monod, 1983 ________________________________________________________________________________ Ascomycota incertae sedis 97  Septogloeum gillii D. Ellis, 1946 ________________________________________________________________________________ Phylum (Division) BASIDIOMYCOTA (club fungi) Basidiomycota: Agaricomycetes: Boletales: Boletaceae 287 Boletus barrowsii Thiers & Smith in Thiers, 1976 ________________________________________________________________________________ Basidiomycota: Agaricomycetes: Boletales: Suillaceae 288 289 Suillus kaibabensis Thiers, 1976  Suillus occidentalis Thiers, 1976 ________________________________________________________________________________ Basidiomycota: Pucciniomycetes: Pucciniales: Pucciniaceae 98  Puccinia circinans J. Ellis & Everhart, 1900 ________________________________________________________________________________ 139 TAXONOMIC LIST KINGDOM CHROMISTA (photosynthetic eukaryotes) Phylum APICOMPLEXA (parasitic alveolates) Apicomplexa: Class : Eucoccidiorida: Eimeriidae (coccid parasites) 201 202 203 204 205 206 207 208 209 210 211            Eimeria tamiasciuri Levine, Ivens & Kruidenier, 1957a Eimeria cutamiae Levine, Ivens & Kruidenier, 1957a Eimeria thomomysis Levine, Ivens & Kruidenier, 1957a Eimeria perognathi Levine, Ivens & Kruidenier, 1957a Eimeria albigulae Levine, Ivens & Kruidenier, 1957a Eimeria operculata Levine, Ivens & Kruidenier, 1957a Eimeria peromysci Levine, Ivens & Kruidenier, 1957a Eimeria arizonensis Levine, Ivens & Kruidenier, 1957a Eimeria eremici Levine, Ivens & Kruidenier, 1957a Eimeria onychomysis Levine, Ivens & Kruidenier, 1957a Isopora citolli Levine, Ivens & Kruidenier, 1957b ________________________________________________________________________________ KINGDOM PROTISTA (protists) “Phylum” HEMIMASTIGOPHORA (single-celled eukaryotes) Hemimastigophora: Hemimastigea: Hemimastigida: Spironemidae (hemimastigid protist) 110 Spironema terricola Foissner & Foissner, 1993 ________________________________________________________________________________ 140 LOCALITIES FOR TYPE MATERIAL FROM GRAND CANYON NATIONAL PARK LOCALITIES FOR TYPE MATERIAL FROM GRAND CANYON NATIONAL PARK * [In the PDF document the lines immediately below are hyperlinks] page “Grand Canyon” not discriminated 143 South Rim 144 North Rim 150 Inner Canyon 154 Colorado River Locales 161 Locality data are as described in the original publications; thus some minor variances will be noted for specific sites or areas that are otherwise in very close proximity to each other. Refinements or special notes appear within [non-bold square brackets]. Red Numbers indicate and cross-list the sequentially enumerated taxa in the bibliography (thus they reflect the alphabetical order by publication author). _________________ * Regarding localities that are today within the Havasupai Indian Reservation or the Hualapai Indian Reservation, but which lie within the physiographic Grand Canyon, see pertinent entries in the LOCALITIES FOR TYPE MATERIAL FROM NEAR GRAND CANYON list. 141 LOCALITIES FOR TYPE MATERIAL FROM GRAND CANYON NATIONAL PARK Federally-Designated Boundaries of Grand Canyon National Park and Predecessor Units 1893–Present map by Stephanie Smith, Grand Canyon Trust, 2019 Grand Canyon Trust https://www.grandcanyontrust.org/grand-canyon-historic-boundaries-map non-commercial permission to use indicated on webpage; last accessed 29 November 2022 142 LOCALITIES FOR TYPE MATERIAL FROM GRAND CANYON NATIONAL PARK “GRAND CANYON” Grand Canyon or Colorado Canyon [without specific locale] 1 58 59 60 61 62 63 64 65 66 67 79 93 97 98 107 111 133 137 140 141 167 183 192 234 246 259 265 270 292 298 314 318 7 Ceratopogon dimidiatus Adams, 1903 Cryptorhopalum pruddeni Casey, 1900 Orphilus aequalis Casey, 1900 Zopherodes lugubris Casey, 1907 Zopherodes pruddeni Casey, 1907 Discodemus brevipennis Casey, 1908 Prionus spiculosus Casey, 1912 Prionus angustulus Casey, 1912 Prionus terminalis Casey, 1912 Stenosphenus pruddeni Casey, 1912 Bembidion (Cyclolopha) occultum Casey, 1918 Triepeolus hopkinsi Cockerell, 1905 Fraxinus macropetala Eastwood, 1903 Septogloeum gillii D. Ellis, 1946 Puccinia circinans J. Ellis & Everhart, 1900 Adenostegia parviflora Ferriss, 1918 Gorytes dentatus Fox, 1893 Rhus canadensis Marsh. var. simplicifolia Greene, 1890 Schmaltzia hirtella Green, 1905 Ptelea nitida Greene, 1906 Ptelea argente Greene, 1906 Arenivaga impensa Hopkins, 2014 Crotalus confluentus abyssus Klauber, 1930 Phytocoris varius Knight, 1934 Cordylanthus tenuifolis Pennell, 1940 Fraxinus cuspidata var. macropetala Rehder, 1917 Penstemon coccinatus Rydberg, 1909 Ochterus barberi Schell, 1943 Melanoplus canonicus Scudder, 1897 Perdita inornata Timberlake, 1962 Trioza rubra Tuthill, 1939 Polymedon castus Wheeler, 1899 Pogonomyrmex californicus Buckley subsp. maricopa Wheeler, 1914 _________________________________________________________________________________ 7 Most of these undiscriminated “Grand Canyon” records are likely based on collections on the far more usually visited South Rim but are listed thusly for their lack of further data. See also “Grand Canyon, Arizona” and (under the “South Rim” heading below) “Grand Canyon village”. Many taxa are represented by material collected prior to the creation of the national monument or park. For localities described as in the “vicinity of”, “near”, or “about” the Grand Canyon, see the LOCALITIES OF TYPE MATERIAL NEAR GRAND CANYON list. 143 LOCALITIES FOR TYPE MATERIAL FROM GRAND CANYON NATIONAL PARK “Grand Canyon National Park” [without specific locale] 197 199 Wellcomia perognathi Kruidenier & Mehra, 1958 Gongylonema peromysci Kruidenier & Peebles, 1958 _________________________________________________________________________________ “Grand Canyon, Arizona” [the canyon or Grand Canyon village not discriminated] 14 15 16 40 41 47 53 54 88 90 200 212 215 216 271 275 299 Athysanella fredonia Ball & Beamer, 1939 Athysanella globosa Ball & Beamer, 1939 Athysanella (Gladionura) casa Ball & Beamer, 1939 Coenonympha fureae Barnes & Benjamin, 1926 Cercyonis damei Barnes & Benjamin, 1926 Delphacodes apicata Beamer, 1948 Erax benedicti Bromley, 1940 Aphelonema convergens var. canyonensia Bunn, 1930 Ballana basala DeLong, 1964 Bruchomorpha bunni Doering, 1939 Andrena (Callandrena) utahensis LaBerge, 1967 Norvellina bicolorata (Ball) var. inflata Lindsay, 1938 Parasitorhabditis gracilis Massey, 1974 Chelinidea vittiger Uhler var. artuflava McAtee, 1919 Mycetophila denningi Shaw, 1951 Heterotarsonemus nakaharai Smiley & Moser, 1974 Platylygus vanduzeei Usinger, 1931 _________________________________________________________________________________ SOUTH RIM [Grand Canyon National Park] “Grand Canyon South Rim” [without specific locale] 44 52 152 226 292 Acmaeodera pletura W. Barr, 1972 Priononyx subatrata Bohart, 1958 Geron prosopidis Hall & Evenhuis, 2003 Neonympha dorothea dorothea Nabokov, 1942 Perdita inornata Timberlake, 1962 _________________________________________________________________________________ “Grand Canyon National Park, South Rim” [without specific locale] 299 Platylygus vanduzeei Usinger, 1931 _________________________________________________________________________________ 144 LOCALITIES FOR TYPE MATERIAL FROM GRAND CANYON NATIONAL PARK “Rim of Grand Canyon” [without specific locale] 85 262 Bombylius facialis Cresson, 1919 Juniperus utahensis var. megalocarpa Sargent, 1919 _________________________________________________________________________________ Arizona Route 64 at southeast boundary of Grand Canyon National Park [near Desert View] 210 Eimeria onychomysis Levine, Ivens & Kruidenier, 1957a _________________________________________________________________________________ Bright Angel, Arizona [Grand Canyon village] see also Grand Canyon village 100 101 102 103 104 105 241 242 262 Petalium bistriatum Say var. arizonense Fall, 1905 Catorama grande Fall, 1905 Catorama longulum Fall, 1905 Catorama pingue Fall, 1905 Diplotaxis conformis Fall, 1909 Bruchus perplexus Fall, 1910 Pilophorus americanus Poppius, 1914 Pilophorus exiguus Poppius, 1914 Juniperus utahensis var. megalocarpa Sargent, 1919 _________________________________________________________________________________ Bright Angel Hotel [Grand Canyon village] see also Grand Canyon village 55 81 Melanophila piniedulis Burke, 1907 Trypeta varipennis Coquillett, 1902 _________________________________________________________________________________ Bright Angel Trail (upper entrance) see also Bright Angel Trail under “Inner Canyon” listings 110 Spironema terricola Foissner & Foissner, 1993 _________________________________________________________________________________ Bright Angel Trail (above) 195 Phytocoris flaviatus Knight, 1968 _________________________________________________________________________________ 145 LOCALITIES FOR TYPE MATERIAL FROM GRAND CANYON NATIONAL PARK Bright Angel Trail (at top) 190 191 Phytocoris hesperius Knight, 1928 Plagiognathus tenellus Knight, 1929 _________________________________________________________________________________ Cataract Canyon, above Supai [The precise locality may be within the boundaries of the Havasupai Indian Reservation] 77 Sclerocactus havasupaiensis Clover, 1942 _________________________________________________________________________________ Cedar Mountain, east side [The precise locality may be within the boundaries of the Navajo Nation] 205 Eimeria albigulae Levine, Ivens & Kruidenier, 1957a _________________________________________________________________________________ Coconino Forest, Grand Canyon (7000 feet) 317 Formica fusca fusca var. gelida Wheeler, 1913 [not valid as a quadrinomial taxon] _________________________________________________________________________________ “on the rim in the Kohonino [Coconino] Forest (7000 feet) about three miles west of the Bright Angel Hotel” 316 Formica moki Wheeler, 1906 _________________________________________________________________________________ Coconino Forest, rim of Grand Canyon 51 Andricus wheeleri Beutenmüller, 1907 _________________________________________________________________________________ Desert View Point 183 Crotalus confluentus abyssus Klauber, 1930 _________________________________________________________________________________ El Tovar (ca. 2 miles west of) 37 Astragalus cremnophylax Barneby, 1948 _________________________________________________________________________________ Eremita Mesa 235 Rosa stellata Wooton subsp. abyssa Phillips, 1992 _________________________________________________________________________________ 146 LOCALITIES FOR TYPE MATERIAL FROM GRAND CANYON NATIONAL PARK Grand Canyon village see also listings under “Bright Angel, Arizona” and “Bright Angel Hotel” above 68 229 Aphonopelma behlei Chamberlin, 1940 Otus asio mychophilus Oberholser, 1937 _________________________________________________________________________________ Grand Canyon village, 36° 03′ 22.31″ N; 112° 07′ 30.73″ W [coordinates pinpoint a locality just to the west of the footpath between Park Headquarters and the Rim Trail, at a point half-way between those two places] 149 Penicillium arizonense Frisvad, Grijseels & J.C. Nielsen in Grijseels et al., 2016 _________________________________________________________________________________ Grand Canyon village (7.5 km east-southeast of), close to South Rim [Grand Canyon National Park, along Arizona Route 64] 17 22 Orbilia purshiae Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia arizonensis Baral & G. Marson in Baral, Weber & Marson, 2020 _________________________________________________________________________________ Grand Canyon village (8.5 km east-southeast of), close to South Rim [Grand Canyon National Park, along Arizona Route 64] 26 Orbilia maeandrina Baral & G. Marson in Baral, Weber & Marson, 2020 _________________________________________________________________________________ Grand View [Grandview] see also Hearst Ranch 184 185 186 187 188 189 193 Deraeocoris fulvus Knight, 1921 Deraeocoris bullatus Knight, 1921 Deraeocoris navajo Knight, 1921 Ceratocapsus clavicornis Knight, 1925 Phytocoris mellarius Knight, 1925 Pilophorus fuscipennis Knight, 1926 Parthenicus cowaniae Knight, 1968 _________________________________________________________________________________ Grandview Point 26 29 Orbilia maeandrina Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia delphinus Baral & G. Marson in Baral, Weber & Marson, 2020 _________________________________________________________________________________ 147 LOCALITIES FOR TYPE MATERIAL FROM GRAND CANYON NATIONAL PARK Hance Ranch see also Thurber’s Camp 142 Ptelea triptera Green, 1906 [type locality for this species refined here; see entry in bibliography] _________________________________________________________________________________ Hearst Ranch see also Grandview 203 Eimeria thomomysis Levine, Ivens & Kruidenier, 1957a _________________________________________________________________________________ Hermit Rim Road 193 Parthenicus cowaniae Knight, 1968 _________________________________________________________________________________ Hualapai Canyon (tributary of Havasu Canyon) [The precise gathering locality may be within the boundaries of the Havasupai Indian Reservation.] 77 Sclerocactus havasupaiensis Clover, 1942 _________________________________________________________________________________ Hull Tank 202 Eimeria cutamiae Levine, Ivens & Kruidenier, 1957a _________________________________________________________________________________ Mather Point 24 Orbilia spermoides Baral & G. Marson, in Baral, Weber & Marson, 2020 _________________________________________________________________________________ Navajo Point (near) (Coconino Plateau) 183 Crotalus confluentus abyssus Klauber, 1930 _________________________________________________________________________________ Shoshone Point 76 Depressaria schellbachi Gates, 1947 _________________________________________________________________________________ 148 LOCALITIES FOR TYPE MATERIAL FROM GRAND CANYON NATIONAL PARK South entrance to Grand Canyon National Park Orchopeas sexdentatus neotomae Augustson, 1943 9 _________________________________________________________________________________ “behind the cabin camp located at the entrance gate of the Grand Canyon National Park” 192 Phytocoris varius Knight, 1934 _________________________________________________________________________________ Thurber’s Camp [Hance Ranch] 95 96 Ripersia arizonensis Ehrhorn, 1899 Dactylopius formicarii Ehrhorn 1899 _________________________________________________________________________________ Tusayan (3.5 km north of) [Grand Canyon National Park, on Arizona Route 64]8 23 25 28 30 32 Orbilia navajoana Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia ophiosoma Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia flexisoma Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia macrodelphinus Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia calyptrata Baral & G. Marson in Baral, Weber & Marson, 2020 _________________________________________________________________________________ Tusayan (3 km north of) [Grand Canyon National Park, on Arizona Route 64]8 18 19 19 20 21 23 24 28 29 Orbilia ocellata Baral, G. Marson & E. Weber in Baral, Weber & Marson, 2020 Orbilia cucumispora Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia magnifica Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia phanosoma Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia multiphanosoma Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia navajoana Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia spermoides Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia flexisoma Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia delphinus Baral & G. Marson in Baral, Weber & Marson, 2020 _________________________________________________________________________________ 8 See the TYPE LOCALITIES NEAR GRAND CANYON list for localities south of Tusayan, Arizona. 149 LOCALITIES FOR TYPE MATERIAL FROM GRAND CANYON NATIONAL PARK Yavapai Point 45 Papilio indra kaibabensis Bauer, 1955 _________________________________________________________________________________ Zuni Point 206 Eimeria operculata Levine, Ivens & Kruidenier, 1957a _________________________________________________________________________________ Zuni Point, west side 207 208 Eimeria peromysci Levine, Ivens & Kruidenier, 1957a Eimeria arizonensis Levine, Ivens & Kruidenier, 1957a _________________________________________________________________________________ NORTH RIM [Grand Canyon National Park] Grand Canyon (North Rim of) [without specific locale] 44 72 170 224 253 256 Acmaeodera pletura W. Barr, 1972 Mycetophila neofungorum Chandler, 1993 Cinara grande Hottes, 1956 Gnomonia quercusgambellii Monod, 1983 Andrena (Scaphandrena) kaibabensis Ribble, 1974 Chrysotimus arizonicus H. Robinson, 1967 _________________________________________________________________________________ Grand Canyon (North Rim) [“Grand Canyon North Rim” or place name “North Rim, Arizona”, not discriminated] see also “North Rim” 43 45 106 112 113 Protoventuria arizonica M. Barr, 1989 Papilio indra kaibabensis Bauer, 1955 Primula (Section Farinosae) hunnewellii Fernald, 1934  Speyeria atlantis schellbachi Garth, 1949 Misumenops coloradensis Gertsch, 1933 _________________________________________________________________________________ Kaibab Plateau [within Grand Canyon National Park] 217 Agave kaibabensis McKelvey, 1949 [see entry in bibliography herein for notes concerning the locality] _________________________________________________________________________________ 150 LOCALITIES FOR TYPE MATERIAL FROM GRAND CANYON NATIONAL PARK Kaibab Plateau, North Rim of Grand Canyon [undiscriminated] 3 Tipula (Lunatipula) kaibabensis C. Alexander, 1946 _________________________________________________________________________________ Bright Angel Creek (head of), Kaibab Plateau 220 Sciurus kaibabensis Merriam, 1904 _________________________________________________________________________________ Bright Angel Point 45 86 Papilio indra kaibabensis Bauer, 1955 Penstemon virgatus pseudoptutus Crosswhite, 1967 _________________________________________________________________________________ Bright Angel Point (“from or near”) 285 Lampropeltis pyromelana Cope infralabialis Tanner, 1953 _________________________________________________________________________________ Burro Spring 183 Crotalus confluentus abyssus Klauber, 1930 _________________________________________________________________________________ Cape Royal 208 Eimeria arizonensis Levine, Ivens & Kruidenier, 1957a _________________________________________________________________________________ Cape Royal Road and Point Imperial Road, W of junction 86 Penstemon virgatus pseudoptutus Crosswhite, 1967 _________________________________________________________________________________ Grand Canyon National Park, North Rim, junction of paved roads, ca. 6.5 km north of Kaibab Lodge, 36° 16′ N, 112° 03′ W [actually Grand Canyon Lodge in Grand Canyon National Park; coordinates pinpoint a location just east of Arizona Route 67 north of the Cape Royal Road junction] 99 Physconia isidiomuscigena Esslinger, 2000 _________________________________________________________________________________ Dutton Point 235 Rosa stellata Wooton subsp. abyssa Phillips, 1992 _________________________________________________________________________________ 151 LOCALITIES FOR TYPE MATERIAL FROM GRAND CANYON NATIONAL PARK Mount Trumbull 268 Neominois dionysus Scudder, 1878 _________________________________________________________________________________ Neal Spring [junction of Cape Royal Road and Point Imperial Road] 112  Speyeria atlantis schellbachi Garth, 1949 _________________________________________________________________________________ North Rim [the place, “North Rim, Arizona”] 155  Polistes canadensis var. kaibabensis Lynn, 1932 [see note in bibliography] _________________________________________________________________________________ North Rim (S of) (93 km south-southeast of Fredonia) [Grand Canyon National Park, North Rim, Arizona] 17 22 26 28 29 33 Orbilia purshiae Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia arizonensis Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia maeandrina Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia flexisoma Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia delphinus Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia subovoidea Baral, Matočec & E. Weber in Baral, Weber & Marson, 2020 _________________________________________________________________________________ Phantom Canyon (head of) 35 Astragalus bryantii Barneby, 1944 _________________________________________________________________________________ Point Sublime 155 289 Polistes canadensis var. kaibabensis Lynn, 1932 Suillus occidentalis Thiers, 1976 _________________________________________________________________________________ Powell Plateau 227 Felis concolor kaibabensis E. Nelson & Goldman, 1931 _________________________________________________________________________________ Road W-3 on the north rim of Grand Canyon near the entrance to Grand Canyon National Park 201 Eimeria tamiasciuri Levine, Ivens & Kruidenier, 1957a _________________________________________________________________________________ 152 LOCALITIES FOR TYPE MATERIAL FROM GRAND CANYON NATIONAL PARK Robbers’ Roost Spring 112  Speyeria atlantis schellbachi Garth, 1949 _________________________________________________________________________________ S B Canyon, Grand Canyon National Monument [now in the enlarged Grand Canyon National Park] 257 Sisymbrium kearneyi Rollins, 1953 _________________________________________________________________________________ S B Point and Hades Knoll (side canyon between) 235 Rosa stellata Wooton subsp. abyssa Phillips, 1992 _________________________________________________________________________________ S B Trail (head of) 235 Rosa stellata Wooton subsp. abyssa Phillips, 1992 _________________________________________________________________________________ Swamp Lake 112  Speyeria atlantis schellbachi Garth, 1949 _________________________________________________________________________________ Swamp Point 91 112 Thomomys bottae boreorarius Durham, 1952  Speyeria atlantis schellbachi Garth, 1949 _________________________________________________________________________________ Swamp Point (½ mile from) (near Powell Saddle) 211 Isopora citolli Levine, Ivens & Kruidenier, 1957b _________________________________________________________________________________ Swamp Ridge 112  Speyeria atlantis schellbachi Garth, 1949 _________________________________________________________________________________ Toroweap Point 109 Echinocereus triglochidiatus var. toroweapensis Fischer, 1991 _________________________________________________________________________________ 153 LOCALITIES FOR TYPE MATERIAL FROM GRAND CANYON NATIONAL PARK Two River Junction 112  Speyeria atlantis schellbachi Garth, 1949 _________________________________________________________________________________ Vulcan’s Throne, Toroweap Valley 243 Oenothera confertiflora Raven, 1962 _________________________________________________________________________________ INNER CANYON [Grand Canyon National Park] Bat Cave [interior] 230 Eschatomoxys pholeter Thomas & Pape in Pape, Thomas & Aalbu, 2007 _________________________________________________________________________________ Bright Angel Canyon 183 Crotalus confluentus abyssus Klauber, 1930 _________________________________________________________________________________ Bright Angel Creek 121 Castor canadensis repentinus Goldman, 1932 _________________________________________________________________________________ Bright Angel Creek (near) 92 Corydalis wetherillii Eastwood, 1902 _________________________________________________________________________________ Bright Angel Trail [without specific locale] see also Bright Angel Trail under “South Rim” listings (above); and “Garden Creek” and “Indian Garden” (below) 50 93 143 156 281 300 301 Mischocyttarus flavitarsis var. kaibabensis Bequaert, 1932 Fraxinus macropetala Eastwood, 1903 Ptelea lutescens Greene, 1906 Sonorella betheli Henderson, 1914 [an erroneous account; see details in the bibliography] Sonora semiannulata gloydi Stickel, 1938 Polymedon dilaticosta Van Duzee, 1927 Polymedon nitidus Van Duzee, 1927 _________________________________________________________________________________ 154 LOCALITIES FOR TYPE MATERIAL FROM GRAND CANYON NATIONAL PARK Bright Angel Trail, 100-400 feet below rim 238 Oreohelix yavapai angelica Pilsbry & Ferriss, 1911 _________________________________________________________________________________ “wall of the Grand Cañon at an altitude of about 5500 feet” [in all probability on Bright Angel Trail] 316 Formica moki Wheeler, 1906 _________________________________________________________________________________ Bright Angel Trail, 4800 ft altitude [altitude indicates locality is at about the 2nd rest house descending] 122 Peromyscus crinitus peridoneus Goldman, 1937 _________________________________________________________________________________ Bright Angel Trail, 1400-2200 m altitude 246 Fraxinus cuspidata var. macropetala Rehder, 1917 _________________________________________________________________________________ Bright Angel Trail, 1400-2000 m altitude 246 Fraxinus cuspidata var. macropetala Rehder, 1917 _________________________________________________________________________________ Bright Angel Trail, about 3000 ft altitude 319 Camponotus acutirostris Wheeler clarigaster Wheeler, 1915 _________________________________________________________________________________ Bright Angel Trail, near Indian Garden [Havasupai Gardens] 136 Schmaltzia cissodes Greene, 1905 _________________________________________________________________________________ Bright Angel Trail, ca. 0.5 mile below Indian Garden [Havasupai Gardens] 4 Thelypodium integrifolium (Nuttall) Endlicher subsp. longicarpum Al-Shehbaz, 1973 _________________________________________________________________________________ Bright Angel Trail, lower 87 Tibicen apache Davis, 1921 _________________________________________________________________________________ 155 LOCALITIES FOR TYPE MATERIAL FROM GRAND CANYON NATIONAL PARK Bright Angel Trail, Lower Sonoran level 281 Sonora semiannulata gloydi Stickel, 1938 _________________________________________________________________________________ Cave of the Domes [interior] [on Horseshoe Mesa] 225 295 Archeolarca cavicola Muchmore, 1981 Eleodes (Caverneleodes) leptoscelis Triplehorn, 1975 _________________________________________________________________________________ Clear Creek, 4 km up from Colorado River confluence 264 Efferia tapeats Scarbrough, Stevens & Nelson, 2012 _________________________________________________________________________________ Clear Creek Canyon 162 Agave phillipsiana Hodgson, 2001 _________________________________________________________________________________ Deer Creek Canyon 162 Agave phillipsiana Hodgson, 2001 _________________________________________________________________________________ Garden Creek (mouth) = Colorado River Mile 88.9 Left 281 Sonora semiannulata gloydi Stickel, 1938 _________________________________________________________________________________ Grandview Trail 99 246 Physconia isidiomuscigena Esslinger, 2000 Fraxinus cuspidata var. macropetala Rehder, 1917 _________________________________________________________________________________ Grandview Trail, 2000 feet below rim 183 Crotalus confluentus abyssus Klauber, 1930 _________________________________________________________________________________ 156 LOCALITIES FOR TYPE MATERIAL FROM GRAND CANYON NATIONAL PARK Hance Trail, Red Canyon [not Old Hance Trail] 143 246 259 279 Ptelea lutescens Green, 1906 Fraxinus cuspidata var. macropetala Rehder, 1917 Penstemon coccinatus Rydberg, 1909 Allionia pachyphylla Standley, 1909 _________________________________________________________________________________ Havasu Canyon, 4 miles down “Supai Trail” [The precise gathering locality may be within the boundaries of the Havasupai Indian Reservation.] 75 Encelia resinifera subsp. tenuifolia Curtis, 1998 _________________________________________________________________________________ Havasu Canyon, near Navajo Falls [The precise gathering locality may be within the boundaries of the Havasupai Indian Reservation.] 77 Sclerocactus havasupaiensis Clover, 1942 _________________________________________________________________________________ Havasu Canyon, on top of Supai formation [The precise gathering locality may be within the boundaries of the Havasupai Indian Reservation.] 77 78 Sclerocactus havasupaiensis Clover, 1942 Sclerocactus havasupaiensis var. roseus Clover, 1942 _________________________________________________________________________________ Hermit Creek 263 283 Populus fremontii var. toumeyi Sargent, 1919 Amsinckia nana Suksdorf, 1931 _________________________________________________________________________________ Hermit Creek (mouth) 35 Astragalus bryantii Barneby, 1944 _________________________________________________________________________________ Hermit Trail 94 246 Oreocarya capitata Eastwood, 1937 Fraxinus cuspidata var. macropetala Rehder, 1917 _________________________________________________________________________________ 157 LOCALITIES FOR TYPE MATERIAL FROM GRAND CANYON NATIONAL PARK Indian Garden [Havasupai Gardens] 10 114 167 290 Salix lutea var. ligulifolia C. Ball, 1921 Arctosa mokiensis Gertsch, 1934a Arenivaga impensa Hopkins, 2014 Perdita wheeleri Timberlake, 1928 _________________________________________________________________________________ Indian Garden (just above) [Havasupai Gardens] 315 Pheidole desertorum var. maricopa Wheeler, 1906 _________________________________________________________________________________ Indian Garden (vicinity) [Havasupai Gardens] see also “Bright Angel Trail, near Indian Garden” 281 Sonora semiannulata gloydi Stickel, 1938 _________________________________________________________________________________ Manzanita Canyon, within ¼ mile of Bright Angel Creek and North Kaibab Trail, UTM 12S 4005015 N, 407532 E 267 Mentzelia canyonensis Schenk, Hodgson & Hufford, 2013 _________________________________________________________________________________ North Kaibab Trail 36 94 276 Astragalus lentiginosus Dougl. var. oropedii Barneby, 1945 Oreocarya capitata Eastwood, 1937 Carex curatorum Stacey, 1937 _________________________________________________________________________________ Old Hance Trail [not Hance Trail, Red Canyon] [the published type localities for these taxa are more precisely identified herein as Old Hance Trail; see main entries in bibliography] 84 144 218 219 279 280 294 320 Ostrya knowltonii Coville, 1894 Ptelea elegans Greene, 1906 Hesperomys megalotis Merriam, 1890 Spilogale gracilis Merriam, 1890 Allionia pachyphylla Standley, 1909 Helix (Arionta) coloradoensis Stearns, 1890 Microsciasma minuta Townsend, 1915 Cicindela rufiventris var. arizonae Wickham, 1899 _________________________________________________________________________________ 158 LOCALITIES FOR TYPE MATERIAL FROM GRAND CANYON NATIONAL PARK Phantom Ranch 34 69 183 Paratriatoma hirsuta Barber, 1938 Aphonopelma phasmus Chamberlin, 1940 Crotalus confluentus abyssus Klauber, 1930 _________________________________________________________________________________ Plateau Point Trail 204 209 Eimeria perognathi Levine, Ivens & Kruidenier, 1957a Eimeria eremici Levine, Ivens & Kruidenier, 1957a _________________________________________________________________________________ Rampart Cave [interior] 230 Eschatomoxys pholeter Thomas & Pape in Pape, Thomas & Aalbu, 2007 _________________________________________________________________________________ Rampart Cave (just west of and below) 222 Sonorella reederi W. Miller, 1984 _________________________________________________________________________________ Roaring Springs 45 Papilio indra kaibabensis Bauer, 1955 _________________________________________________________________________________ Roaring Springs Cave [interior] 232 Ptomaphagus cocytus Peck, 1973 _________________________________________________________________________________ Roaring Springs Cave, ca. 1.6 km inside 302 Stygobromus blinni Wang & Holsinger, 2001 _________________________________________________________________________________ Roaring Springs Power Plant 183 Crotalus confluentus abyssus Klauber, 1930 _________________________________________________________________________________ Shinumo Creek 183 Crotalus confluentus abyssus Klauber, 1930 _________________________________________________________________________________ 159 LOCALITIES FOR TYPE MATERIAL FROM GRAND CANYON NATIONAL PARK Shinumo Creek, 3 km from confluence with Colorado River 264 Efferia tapeats Scarbrough, Stevens & Nelson, 2012 _________________________________________________________________________________ South Bass Trail, about 200 feet below the rim 237 239 Oreohelix yavapai extremitatis Pilsbry & Ferriss, 1911 Pupilla syngenes avus Pilsbry & Ferriss, 1911 _________________________________________________________________________________ South Bass Trail (“the head of a recess in the cross-bed sandstone [Coconino Sandstone] south of where the Mystic Spring or Bass Trail [South Bass Trail] zigzags down, in a talus resting on the red sandstone forming the Le Conte Plateau”) [Pilsbry & Ferris 1911 also illustrate the appearance of the locality in a sketch] 236 Oreohelix yavapai profundorum Pilsbry & Ferriss, 1911 _________________________________________________________________________________ South Bass Trail, in Supai formation 264 Efferia tapeats Scarbrough, Stevens & Nelson, 2012 _________________________________________________________________________________ South Bass Trail, in Redwall Formation 264 Efferia tapeats Scarbrough, Stevens & Nelson, 2012 _________________________________________________________________________________ South Bass Trail, base of Tapeats Formation 264 Efferia tapeats Scarbrough, Stevens & Nelson, 2012 _________________________________________________________________________________ South Kaibab Trail, 5300 feet elevation, in debris slides and crevices of Redwall Limestone 244 Oenothera specuicola subsp. specuicola Raven, 1962 _________________________________________________________________________________ South Kaibab Trail, 36° 03′ 45″ N, 112° 03′ 30″ W 99 Physconia isidiomuscigena Esslinger, 2000 _________________________________________________________________________________ Tanner Trail 183 Crotalus confluentus abyssus Klauber, 1930 _________________________________________________________________________________ 160 LOCALITIES FOR TYPE MATERIAL FROM GRAND CANYON NATIONAL PARK Thunder River Cave [interior] 117 Loxosceles kaiba Gertsch & Ennik, 1983 _________________________________________________________________________________ Tonto Plateau, near Bright Angel Trail 183 Crotalus confluentus abyssus Klauber, 1930 _________________________________________________________________________________ Tse’an Cho [interior] [cave on Horseshoe Mesa] 295 Eleodes (Caverneleodes) leptoscelis Triplehorn, 1975 _________________________________________________________________________________ Tse’an Olje [interior] [cave in Cremation Canyon] 295 Eleodes (Caverneleodes) leptoscelis Triplehorn, 1975 _________________________________________________________________________________ COLORADO RIVER LOCALES [Grand Canyon National Park] Colorado River [without specific locale] 213 Chionaspis gilli Liu & Kosztarab, 1987 _________________________________________________________________________________ Marble Canyon, along the Colorado River, 12 river miles below Lees Ferry, Salt Water Wash 163 Euphorbia aaronrossii A. Holmgren & N. Holmgren, 1988 _________________________________________________________________________________ River Mile 23.0 Left 240 Schinia immaculata Pogue, 2004 _________________________________________________________________________________ Vasey’s Paradise [River Mile 31.8 Right] 273 Wiedemannia digna Sinclair, 2006 _________________________________________________________________________________ 161 LOCALITIES FOR TYPE MATERIAL FROM GRAND CANYON NATIONAL PARK River Mile 37.3 Left 168 Arenivaga pagana Hopkins, 2014 _________________________________________________________________________________ River Mile 53 Right, Nankoweap 168 Arenivaga pagana Hopkins, 2014 _________________________________________________________________________________ River Mile 58.2 Right, Awatubi Canyon 264 Efferia tapeats Scarbrough, Stevens & Nelson, 2012 _________________________________________________________________________________ Colorado River, at or near mouth of Bright Angel Creek [Bright Angel Creek confluence is at River Mile 87.7 Right] 221 Gila cypha R. Miller, 1946 _________________________________________________________________________________ River Mile 88.9 Left see “Garden Creek (mouth)” under “Inner Canyon” listings, above _________________________________________________________________________________ River Mile 92.3 Left 181 Nebria (Reductonebria) georgei Kavanaugh, 2008 _________________________________________________________________________________ River Mile 108 Right, Shinumo Creek (mouth) 264 Efferia tapeats Scarbrough, Stevens & Nelson, 2012 _________________________________________________________________________________ River Mile 108 Left 264 Efferia tapeats Scarbrough, Stevens & Nelson, 2012 _________________________________________________________________________________ River Mile 147.8 Left, Matkatamiba Canyon 286 Flaveria mcdougallii Theroux, Pinkava & Keil, 1977 _________________________________________________________________________________ River Mile 158 Right (campsite near), 339451 E, 4018170 N, NAD 83, 36°17′42.6″ N; 112°47′16.6″ W 266 Mentzelia hualapaiensis Schenk, Hodgson & Hufford, 2010 _________________________________________________________________________________ 162 LOCALITIES FOR TYPE MATERIAL FROM GRAND CANYON NATIONAL PARK River Mile 160.5 Left 166 Arenivaga grandiscanyonensis Hopkins, 2014 _________________________________________________________________________________ Note: Localities between River Miles 164.7 and 273.3, on the left, could be within the boundaries of the Hualapai Indian Reservation _________________________________________________________________________________ River Mile 166.5 Left 240 Schinia immaculata Pogue, 2004 _________________________________________________________________________________ River Mile 174.2 Right, Cove Canyon 286 Flaveria mcdougallii Theroux, Pinkava & Keil, 1977 _________________________________________________________________________________ River Mile 180.8 Right 181 Nebria (Reductonebria) georgei Kavanaugh, 2008 _________________________________________________________________________________ River Mile 186.5 Left 168 Arenivaga pagana Hopkins, 2014 _________________________________________________________________________________ River Mile 198.0 Right 168 Arenivaga pagana Hopkins, 2014 _________________________________________________________________________________ River Mile 202.5 Left 168 Arenivaga pagana Hopkins, 2014 _________________________________________________________________________________ River Mile 202 Right 240 Schinia immaculata Pogue, 2004 _________________________________________________________________________________ River Mile 211.5 Right 166 Arenivaga grandiscanyonensis Hopkins, 2014 _________________________________________________________________________________ 163 LOCALITIES FOR TYPE MATERIAL FROM GRAND CANYON NATIONAL PARK 164 LOCALITIES FOR TYPE MATERIAL FROM NEAR GRAND CANYON NATIONAL PARK LOCALITIES FOR TYPE MATERIAL FROM NEAR GRAND CANYON NATIONAL PARK See also various headings within this list for further refinements [In the PDF document the lines immediately below are hyperlinks] page Cameron, Arizona, and vicinity 166 Arizona Strip 167 Grand Canyon vicinity undiscriminated 171 Grand Canyon–Parashant National Monument 172 Havasupai Indian Reservation 174 House Rock and House Rock Valley 174 Hualapai Indian Reservation 175 Hualapai Indian Reservation vicinity (off reservation) 177 “Kaibab Forest” (north) undiscriminated 177 “Kaibab Forest” (south) undiscriminated 177 Kaibab Indian Reservation 178 Kaibab National Forest, Tusayan Ranger District 178 Kaibab Plateau (including Kaibab National Forest, North Kaibab Ranger District) 179 Kanab Canyon 184 Lees Ferry 184 Marble Canyon [not on Colorado River] 184 Navajo Nation (westernmost) 185 Red Mountain 185 165 LOCALITIES FOR TYPE MATERIAL FROM NEAR GRAND CANYON NATIONAL PARK Locality data are as described in the original publications; thus some minor variances will be noted for specific sites or areas that are otherwise in very close proximity to each other. Refinements or special notes appear within [non-bold square brackets]. Red Numbers indicate and cross-list the sequentially enumerated taxa in the bibliography (thus they reflect the alphabetical order by publication author). Some taxa listed here may have been collected from the physiographic Grand Canyon, but outside of the national park boundaries. Note: Baral et al. (2020) did not demonstrate a precise understanding of the geography of the Grand Canyon region. All of their localities are listed as being in Arizona (a few actually are in Utah), but some locality references to “Grand Canyon” are, confusingly, neither within the physiographic canyon nor within the national park but are on adjacent national forest and other lands, sometimes at quite some distance from the canyon or the national park. Some interpretive statements have been included in this locality list in order to clarify these discrepancies; specifically, measurements have been made from a map that reveal that linear distances from geographic locales are mostly along the principal highways of the area, which information is added notationally to the original locality data. CAMERON, ARIZONA and vicinity Cameron 80 Myrmecocystus melliger subsp. semirufus Emery var. romainei Cole, 1936 [not valid as a 196 Aphricus knowltoni Knull, 1957 quadrinomial taxon] _________________________________________________________________________________ Cameron Trading Post 2 Plagiomimicus kathyae J. Adams & Lafontaine, 2009 _________________________________________________________________________________ Little Colorado River (bed of) at Cameron 38 Astragalus wootoni var. endopterus Barneby, 1949 _________________________________________________________________________________ 166 LOCALITIES FOR TYPE MATERIAL FROM NEAR GRAND CANYON NATIONAL PARK Cameron (15 miles W of, along the canyon of the Little Colorado River) [quite probably at or near the Little Colorado River Gorge Overlook, along Arizona Route 64] 278 Megathymus alliae Stallings & Turner, 1957 _________________________________________________________________________________ ARIZONA STRIP 9 “Northern Arizona” [general vicinity nearest to Kanab, Utah; without specific locale] 303 304 305 306 307 308 309 Dalea amoena Watson, 1873 Oenothera (Chylisma) multijuga Watson, 1873 Cymopterus purpureus Watson, 1873 Peucedanum newberryi Watson, 1873 Brickellia (Clavigera) longifolia Watson, 1873 Calochortus flexuosus Watson, 1873 Androstephium breviflorum Watson, 1873 _________________________________________________________________________________ Arizona Route 389, 4.3 mi west of U.S. Route 89A [near Fredonia, Arizona] 251 Eriogonum thompsonae S. Wats. var. atwoodii Reveal, 1974b _________________________________________________________________________________ Arizona Route 389, 4.5 mi southwest of U.S. Route 89A [near Fredonia, Arizona] 250 Eriogonum mortonianum Reveal, 1974a _________________________________________________________________________________ Bulrush Canyon (northwest of, south of Pipe Spring) 231 Opuntia pinkavae Parfitt, 1997 _________________________________________________________________________________ Colorado City (6 miles east of; Rosy Canyon Road, 1.5 miles south of Utah state line) 173 Trogloderus skillmani Johnston, 2019 _________________________________________________________________________________ 9 Refer also separately in this list to: Grand Canyon–Parashant National Monument; House Rock Valley; Kaibab Indian Reservation; Kaibab National Forest; Kaibab Plateau; Kanab Canyon; Lees Ferry. 167 LOCALITIES FOR TYPE MATERIAL FROM NEAR GRAND CANYON NATIONAL PARK Fredonia, Arizona Athysanella fredonia Ball & Beamer, 1939 14 _________________________________________________________________________________ Fredonia (1 mile north of) 231 Opuntia pinkavae Parfitt, 1997 _________________________________________________________________________________ Hack Canyon (vicinity) Phacelia higginsii Atwood, 2007 6 _________________________________________________________________________________ “Hell Canyon” [see bibliography for comments on the locality] 148 Morus grisea Greene, 1911 _________________________________________________________________________________ Kaibab Indian Reservation (west of) 231 Opuntia pinkavae Parfitt, 1997 _________________________________________________________________________________ Millipede Cave [interior] [Bureau of Land Management lands] 272 Pratherodesmus voylesi Shear in Shear, Taylor, Wynne & Krejca, 2009 _________________________________________________________________________________ Mokiak Pass [20 miles southeast of St. George, Utah 125 126 127 128 129 269 310 10 ] Astragalus subcinereus A. Gray, 1878 Astragalus scaposus A. Gray, 1878 Astragalus mokiacensis A. Gray, 1878 Astragalus artipes A. Gray, 1878 Actinella biennis A. Gray, 1878 Synchloe thoosa Scudder, 1878 Lesquerella arizonica Watson, 1888 _________________________________________________________________________________ 10 Location of Mokiak Pass is as identified by S. H. Scudder (1878, Notice of the butterflies collected by Dr. Edward Palmer in the arid regions of southern Utah and northern Arizona during the summer of 1877, U.S. Geological and Geographical Survey of the Territories, Bulletin, 4). 168 LOCALITIES FOR TYPE MATERIAL FROM NEAR GRAND CANYON NATIONAL PARK Main Street Valley, ca. 1.5 km south of the junction of the Colorado City road with the Main Street Valley road 245 Panicum mohavense Reeder, 1991 _________________________________________________________________________________ Main Street Valley, west of Hurricane Cliffs 231 Opuntia pinkavae Parfitt, 1997 _________________________________________________________________________________ Navaho Trail, near Hurricane Rim 231 Opuntia pinkavae Parfitt, 1997 _________________________________________________________________________________ North Canyon (Kaibab Plateau, East Rim View Point along Trail #7 ca. 2¼ mi down from trailhead, in North Canyon; 36°25′18″N, 112°04′22″W) 252 Scutellaria potosina var. kaibabensis Rhodes & Ayers, 2010 _________________________________________________________________________________ North Canyon Trail #4, 36°25′04″N 112°04′17″W [east rim of Kaibab Plateau] 252 Scutellaria potosina var. kaibabensis Rhodes & Ayers, 2010 _________________________________________________________________________________ October Gyp Cave [interior] [Bureau of Land Management lands] 272 Pratherodesmus voylesi Shear in Shear, Taylor, Wynne & Krejca, 2009 _________________________________________________________________________________ Page (9 miles northwest of) 228 Acmaeodera navajo G. Nelson & Westcott, 1995 _________________________________________________________________________________ Pipe Spring (ca. 9 miles east-northeast of) see also Kaibab Indian Reservation 250 Eriogonum mortonianum Reveal, 1974a _________________________________________________________________________________ 169 LOCALITIES FOR TYPE MATERIAL FROM NEAR GRAND CANYON NATIONAL PARK Uinkaret Plateau, just north of the Navajo Trail Road, 1.1 km W of the Clayhole Wash crossing, 27.5 km air distance south-southweest of Colorado City, 36°49′09″N, 113°11′21″W, T40N R8W S31 165 Mentzelia memorabilis N. Holmgren & P. Holmgren, 2002 _________________________________________________________________________________ Vermilion Cliffs [geographical feature or place name undiscriminated11] 228 Acmaeodera navajo G. Nelson & Westcott, 1995 _________________________________________________________________________________ Vermilion Lodge (vicinity) [Vermilion Cliffs] 282 Atriplex asterocarpa Stutz, Chu & Sanderson, 1994 _________________________________________________________________________________ Virgin Mountains 139 Ptelea straminea Green, 1906 _________________________________________________________________________________ Wolf Hole 118 Dipodomys microps celsus Goldman, 1924 _________________________________________________________________________________ Wolf Hole (4.2 miles southwest of) 231 Opuntia pinkavae Parfitt, 1997 _________________________________________________________________________________ Wolf Hole (20 miles south of) 124 Thomomys bottae nicholi Goldman, 1938 _________________________________________________________________________________ 11 In their paper, Nelson & Westcott also specially mention “Cliff Dwellers Lodge”, which is not far from another business by the name of Vermilion Cliffs, thus it is unclear whether “Vermilion Cliffs” refers to that place, or to the natural geographical feature Vermilion Cliffs that align in this area along U.S. Route 89 Alternate. 170 LOCALITIES FOR TYPE MATERIAL FROM NEAR GRAND CANYON NATIONAL PARK GRAND CANYON VICINITY 12 undiscriminated “about Grand Canyon” or “region of the Grand Canyon” 89 145 254 Galium munzii Hilend & Howell subsp. ambivalens Dempster & Ehrendorfer, 1965 Ptelea confinis Greene, 1906 Silene rectiramea Robinson, 1899 _________________________________________________________________________________ Grand Canyon (“near”) 12 56 57 194 Hysteropterum cornutum var. utahnum E. Ball, 1935 Trichochrous incipiens Casey, 1895 Trichochrous reversus Casey, 1895 Bolteria juniperi Knight, 1968 _________________________________________________________________________________ Grand Canyon, Ariz. (“vicinity of”) Aphelonema convergens var. canyonensia Bunn, 1930 54 _________________________________________________________________________________ Grand Canyon (south of) 249 Fabriciella evanida Reinhard, 1953 _________________________________________________________________________________ Grand Canyon (30 miles south of) 277 Vaejovis aquilonalis Stahnke, 1940 _________________________________________________________________________________ 12 It is not clear whether materials collected with the indefinite locality descriptions here include among all specimens many, several, or one that may have come from the Grand Canyon itself. Because of this uncertainty, the taxa listed here are included in the “near Grand Canyon National Park” list of type specimens rather than in the national park list itself. 171 LOCALITIES FOR TYPE MATERIAL FROM NEAR GRAND CANYON NATIONAL PARK GRAND CANYON–PARASHANT NATIONAL MONUMENT 13 12.5 miles southwest from head of Trail Canyon toward Andrus Canyon, 36°11.48′N, 113°19.10′W Phacelia higginsii Atwood, 2007 6 _________________________________________________________________________________ Andrus Canyon, 1 mile west of Andrus Point, T32N R10W S10 311 Astragalus lentiginosus Douglas ex Hook. var. trumbullensis Welsh & Atwood, 2001 _________________________________________________________________________________ Andrus Canyon, 3 miles west of Andrus Point, T32N R10W S6, wash bottom 311 Astragalus lentiginosus Douglas ex Hook. var. trumbullensis Welsh & Atwood, 2001 _________________________________________________________________________________ Andrus Canyon at road crossing, T32N, R10W, S6 & 7 Phacelia furnissii Atwood, 2007 5 _________________________________________________________________________________ Cunningham Canyon Xylorhiza tortifolia (Torrey & Gray) Greene var. parashantensis Atwood & Welsh, 2013 8 _________________________________________________________________________________ Daneill Canyon, west of Andrus Canyon, 3 miles west of Andrus Pt., T32N R11W S12 311 Astragalus lentiginosus Douglas ex Hook. var. trumbullensis Welsh & Atwood, 2001 _________________________________________________________________________________ Eldel Cave [interior] 230 Eschatomoxys pholeter Thomas & Pape in Pape, Thomas & Aalbu, 2007 _________________________________________________________________________________ 13 Localities are as determined based on boundaries at the time of publication of this checklist. Prior to the creation of Grand Canyon–Parashant National Monument (in 2000), and depending upon original dates of collection, some material may originally be said to have been from Grand Canyon National Monument, Lake Mead National Recreation Area (that is, its northeastern section), or the Arizona Strip generally. 172 LOCALITIES FOR TYPE MATERIAL FROM NEAR GRAND CANYON NATIONAL PARK Mount Trumbull, 3 miles south of Nixon Spring [The locality, “three miles south of Nixon Spring”, is just to the south of Mount Trumbull itself, today within the Grand Canyon–Parashant National Monument, about 2¾ miles north of the boundary of Grand Canyon National Park.] 151 Thomomys bottae trumbullensis Hall & Davis, 1934 _________________________________________________________________________________ Mt. Trumbull school house (2 miles south of ), near Griffith Knoll, T34S R10W S1-2 311 Astragalus lentiginosus Douglas ex Hook. var. trumbullensis Welsh & Atwood, 2001 _________________________________________________________________________________ PARA-1001 Cave, ca. UTM 0264500 N, 4060700 E [interior] 153 154 233 Hesperochernes bradybaughi Harvey & Wynne, 2014 Tuberochernes cohni Harvey & Wynne, 2014 Ptomaphagus parashant Peck & Wynne, 2013 _________________________________________________________________________________ Parashant (Trail) Canyon, 11 miles south of Mount Trumbull Village 311 Astragalus lentiginosus Douglas ex Hook. var. trumbullensis Welsh & Atwood, 2001 _________________________________________________________________________________ Poverty Mountain (south base) 7 Phacelia hughesii Atwood, 2007 _________________________________________________________________________________ Poverty Mountain (southeast of), southwest of Mt. Trumbull, T34N, R11W, Sec 4 312 Pediomelum pauperitense S.L. Welsh, M. Licher, & N. D. Atwood in Welsh & Licher, 2010 _________________________________________________________________________________ Poverty Mountain (southwest of) near Dewdrop Spring, T35N, R12W, Sec. 35/36 312 Pediomelum pauperitense S.L. Welsh, M. Licher, & N. D. Atwood in Welsh & Licher, 2010 _________________________________________________________________________________ Poverty Mountain (BLM road south of) and southwest of Salt Spring, T34N, R11W [sic] 312 Pediomelum pauperitense S.L. Welsh, M. Licher, & N. D. Atwood in Welsh & Licher, 2010 _________________________________________________________________________________ Sandstone Canyon (1.5 mile due west of Cane Spring) 8 Xylorhiza tortifolia (Torrey & Gray) Greene var. parashantensis Atwood & Welsh, 2013 _________________________________________________________________________________ 173 LOCALITIES FOR TYPE MATERIAL FROM NEAR GRAND CANYON NATIONAL PARK Twin Point (west rim of) (18 km south of Oak Grove, T30N R12W NW¼ sect. 7, 36°01′N, 113°37′W) 235 Rosa stellata Wooton subsp. abyssa Phillips, 1992 _________________________________________________________________________________ Whitmore Canyon, T32N R9W S1 311 Astragalus lentiginosus Douglas ex Hook. var. trumbullensis Welsh & Atwood, 2001 _________________________________________________________________________________ Whitmore Canyon, 1.25 km (7.8 miles) [sic] south of Mount Trumbull (abandoned town), T34N R9W S29 311 Astragalus lentiginosus Douglas ex Hook. var. trumbullensis Welsh & Atwood, 2001 _________________________________________________________________________________ HAVASUPAI INDIAN RESERVATION See also entries under “Havasu” in the LOCALITIES FOR TYPE MATERIAL IN GRAND CANYON NATIONAL PARK list noted as possibly being within the Havasupai Reservation. These localities are within the physiographic Grand Canyon. “Canyon at Supai, Coconino County, Ariz.” 214 Oreohelix yavapai vauxae Marshall, 1929 _________________________________________________________________________________ Mooney Falls, near 244a Oenothera specuicola subsp. hesperia Raven, 1962 _________________________________________________________________________________ Supai, Havasu Canyon 293 Perdita depressa Timberlake, 1968 _________________________________________________________________________________ HOUSE ROCK and HOUSE ROCK VALLEY House Rock, near Lees Ferry 180 Astragalus kaibensis Jones, 1902 _________________________________________________________________________________ 174 LOCALITIES FOR TYPE MATERIAL FROM NEAR GRAND CANYON NATIONAL PARK House Rock Valley [without specific locale] 49 74 161 Pediocactus paradinei Benson, 1957 Cordylanthus wrightii kaibabensis Chuang & Heckard, 1986 Sclerocactus whipplei (Englemann & Bigelow) Britton & Rose ssp. busekii Hochstätter, 1995 _________________________________________________________________________________ House Rock Valley, north of Rock Canyon 231 Opuntia pinkavae Parfitt, 1997 _________________________________________________________________________________ 45 km southwest of Page, 26 km southwest of Marble Canyon [House Rock valley vicinity] 18 Orbilia ocellata Baral, G. Marson & E. Weber in Baral, Weber & Marson, 2020 _________________________________________________________________________________ HUALAPAI INDIAN RESERVATION These localities, with the exception perhaps of some collections from the “vicinity” of, or southerly from, Peach Springs are within the physiographic Grand Canyon. Diamond Creek Canyon 135 Arabis recondita Greene, 1900 _________________________________________________________________________________ Diamond River [Diamond Creek] 131 Phacelia (Coreanthus) ivesiana A. Gray in Gray et al., 1861 _________________________________________________________________________________ Diamond River [Diamond Creek] (mouth of) 132 “Gilia dactylophyllum, (n. sp.?)”, A. Gray in Gray et al., 1861 [see remarks in bibliography] _________________________________________________________________________________ Grand Cañon of the Colorado, below Peach Springs 130 Mirabilis bigelovii A. Gray, 1886 _________________________________________________________________________________ 175 LOCALITIES FOR TYPE MATERIAL FROM NEAR GRAND CANYON NATIONAL PARK Peach Springs 169 247 274 310 Luperodes wickhami Horn, 1893 Fraxinus lowellii Sargent in Rehder, 1917 Eriogonum fusiforme Small, 1906 Lesquerella arizonica Watson, 1888 _________________________________________________________________________________ Peach Springs (above) 138 Ptelea palida Green, 1906 _________________________________________________________________________________ Peach Springs (vicinity) 82 83 134 147 Echinocactus polycephalus var. xeranthemoides Coulter, 1896 Phellopterus multinervatus Coulter & Rose, 1900 Arabis eremophila Greene, 1900 Senecio stygius Greene, 1909 [type locality for this species refined here; see main entry in bibliography] 261 Pediomelum retrorsum Rydberg in Rydberg & Pennell, 1919 _________________________________________________________________________________ Peach Springs Canyon (Peach Springs Wash) 159 255 Opuntia abyssi Hester, 1943 Crossosoma parviflora B. Robinson & Fernald, 1894 [type locality for this species refined here; see entry in bibliography] 284 Amsinckia macrosepala Suksdorf, 1931 [type locality for this species refined here; see entry in bibliography] _________________________________________________________________________________ Peach Springs Wash, 4 km from confluence with Colorado River 264 Efferia tapeats Scarbrough, Stevens & Nelson, 2012 _________________________________________________________________________________ Prospect Valley (lower end) 120 Ammospermophilus leucurus tersus Goldman, 1929 _________________________________________________________________________________ Upper Peach Springs, outflow just below concrete weir, 35.55786°N, 113.4314°W 157 Pyrgulopsis hualapaiensis Herschler, Liu & Stevens, 2016 _________________________________________________________________________________ 176 LOCALITIES FOR TYPE MATERIAL FROM NEAR GRAND CANYON NATIONAL PARK HUALAPAI INDIAN RESERVATION vicinity (off reservation) Peach Springs (8 miles southwest of) [vicinity of Truxton, Arizona] 150 Boharticus margaretae Grissell, 1983 _________________________________________________________________________________ U.S.-Arizona Highway No. 66, 11 or 12 miles east of Peach Springs and 2 or 3 miles west of Hyde Park, Arizona [“Hyde Park”, near mile marker 117 on historic Route 66, no longer exists; the gathering locality is near the present-day Grand Canyon Caverns tourist attraction.] 158 Opuntia hualpaensis Hester, 1943 _________________________________________________________________________________ “KAIBAB FOREST” (north, undiscriminated) [see also “Kaibab Plateau”] “Kaibab forest, near the north rim of the Grand Canyon” 115 Phidippus kaibabensis Gertsch, 1934b _________________________________________________________________________________ “KAIBAB FOREST” (south, undiscriminated) [see also “Kaibab National Forest, Tusayan Ranger District”] “W 112°:N 36°, Kaibab Forest, Arizona” [The overly generalized coordinates lie within Grand Canyon National Park, just west of Grandview Point; the precise intersection is in fact, by coincidence, on a cliff face(!) just over the rim of the canyon (https://confluence.org/confluence.php?id=91). The specific epithet of the taxon noted here, and indication of “Kaibab Forest,” may point either to a locality on the nearby Kaibab National Forest or is only a generalized term for the South Rim forest, sometimes called the Coconino forest, on the Coconino Plateau.] 71 Neoantistea coconino Chamberlin & Ivie, 1942 _________________________________________________________________________________ 177 LOCALITIES FOR TYPE MATERIAL FROM NEAR GRAND CANYON NATIONAL PARK KAIBAB INDIAN RESERVATION Kanab Wash, at southern boundary of Kaibab Indian Reservation 119 Dipodomys ordii cupidineus Goldman, 1924 _________________________________________________________________________________ Pipe Spring (½ mile north of) Opuntia aurea Baxter, 1933 46 _________________________________________________________________________________ KAIBAB NATIONAL FOREST, TUSAYAN RANGER DISTRICT [see also “Kaibab Forest (south, undiscriminted)”] Arizona Route 64, junction of road to Anita [road junction coordinates ca. 35.861° N, 112.134° W] 171 Cinara poketa Hottes, 1956 _________________________________________________________________________________ Tusayan (3 km south of) [locality is on the Kaibab National Forest on Arizona Route 64]14 20 25 28 31 32 Orbilia phanosoma Baral & [G.] Marson in Baral, Weber & Marson, 2020 Orbilia ophiosoma Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia flexisoma Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia multitrapezoidea Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia calyptrata Baral & G. Marson in Baral, Weber & Marson, 2020 _________________________________________________________________________________ 14 See the LOCALITIES FOR TYPE MATERIAL IN GRAND CANYON NATIONAL PARK list for localities north of Tusayan that are within Grand Canyon National Park. 178 LOCALITIES FOR TYPE MATERIAL FROM NEAR GRAND CANYON NATIONAL PARK KAIBAB PLATEAU [also as Buckskin Mountains] including KAIBAB NATIONAL FOREST, NORTH KAIBAB RANGER DISTRICT [see also “Kaibab Forest (north, undiscriminated)”] Kaibab Plateau [without specific locale] 112 Speyeria atlantis schellbachi Garth, 1949 _________________________________________________________________________________ Kaibab National Forest [without specific locale but on North Kaibab Ranger District] 182 Enderleinellus kaibabensis Kim, 1966 _________________________________________________________________________________ Kaibab Plateau, North Rim of Grand Canyon [undiscriminated] 3 Tipula (Lunatipula) kaibabensis C. Alexander, 1946 _________________________________________________________________________________ Kaibab Plateau / Buckskin Mountains [without specific locale] 174 176 Astragalus humistratus var. tenerrimus Jones, 1895 Bigelovia howardi var. attenuata Jones, 1895 _________________________________________________________________________________ Buckskin Mountains (mesa below) 177 Laphamia congesta Jones, 1895 _________________________________________________________________________________ Kaibab Plateau (northwest escarpment slope, 6700 feet) 248 296 Morsea california kaibabensis Rehn & Grant, 1958 Asidina rugicollis Triplehorn & Brown, 1971 _________________________________________________________________________________ Arizona Route 67, 3.2 km north of Grand Canyon National Park boundary, T.34 N., R.3 E., Sec. 7 164 Castilleja kaibabensis N. Holmgren, 1973 _________________________________________________________________________________ 179 LOCALITIES FOR TYPE MATERIAL FROM NEAR GRAND CANYON NATIONAL PARK DeMotte Park 86 123 164 175 Penstemon virgatus pseudoptutus Crosswhite, 1967 Thomomys fossor kaibabensis Goldman, 1938 Castilleja kaibabensis N. Holmgren, 1973 Oenothera triloba var. ecristata Jones 1895 _________________________________________________________________________________ Dry Park 86 Penstemon virgatus pseudoptutus Crosswhite, 1967 _________________________________________________________________________________ Fredonia (28 km (17.5 miles) airline distance east of), T41N, R2E, S29/30 [an area of Forest Service roads] 39 Astragalus cremnophylax Barneby var. myriorrhaphis Barneby, 1979 _________________________________________________________________________________ Fredonia (28 km east-southeast of Fredonia, 13 km NNW of Jacob Lake) [along U.S. Route 89A] 17 22 24 26 27 30 Orbilia purshiae Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia arizonensis Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia spermoides Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia maeandrina Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia multimaeandrina Baral & G. Marson in Baral, Weber & Marson, 2020 Orbilia macrodelphinus Baral & G. Marson in Baral, Weber & Marson, 2020 _________________________________________________________________________________ Fredonia Sawmill (17 miles southeast of), along logging road to Ryan 313 Astragalus pinionis Welsh & Thorne, 1977 _________________________________________________________________________________ Hwy. from Jacob Lake to Lodge, Kaibab Forest [Arizona Route 67, between Jacob Lake and Kaibab Lodge] 86 Penstemon virgatus pseudoptutus Crosswhite, 1967 _________________________________________________________________________________ House Rock Canyon to Jacob Lake 164 Castilleja kaibabensis N. Holmgren, 1973 _________________________________________________________________________________ 180 LOCALITIES FOR TYPE MATERIAL FROM NEAR GRAND CANYON NATIONAL PARK Jacob Lake 223 287 288 Saxinis saucia kaibabiae Moldenke, 1970 Boletus barrowsii Thiers & Smith in Thiers, 1976 Suillus kaibabensis Thiers, 1976 _________________________________________________________________________________ Jacob Lake (near) 74 297 Cordylanthus wrightii kaibabensis Chuang & Heckard, 1986 Hadena (Hadena) lafontainei Troubridge & Crabo, 2002 _________________________________________________________________________________ Jacob Lake (northwest of) 108 Coryphantha vivipara var. kaibabensis Fischer, 1979 _________________________________________________________________________________ Jacob Lake (7.5 miles east of) 74 Cordylanthus wrightii kaibabensis Chuang & Heckard, 1986 _________________________________________________________________________________ Jacob Lake (south of), 37 km southeast of Fredonia [on Arizona Route 67] 19 Orbilia magnifica Baral & G. Marson in Baral, Weber & Marson, 2020 _________________________________________________________________________________ Jacob Lake (18.6 miles south of, on Arizona Route 67) 258 Lesquerella kaibabensis Rollins, 1982 _________________________________________________________________________________ Jacob Lake (20 miles south of) 253 Andrena (Scaphandrena) kaibabensis Ribble, 1974 _________________________________________________________________________________ Kaibab Forest, near Grand Canyon National Park 86 Penstemon virgatus pseudoptutus Crosswhite, 1967 _________________________________________________________________________________ Kaibab Lodge (3 km north of) [on Arizona Route 67] 26 Orbilia maeandrina Baral & G. Marson in Baral, Weber & Marson, 2020 _________________________________________________________________________________ 181 LOCALITIES FOR TYPE MATERIAL FROM NEAR GRAND CANYON NATIONAL PARK Kaibab Lodge (4 miles north of) 172 Cicindela pusilla kaibabensis Johnson, 1990 _________________________________________________________________________________ Kaibab Lodge (8 miles north of) 172 Cicindela pusilla kaibabensis Johnson, 1990 _________________________________________________________________________________ Kaibab National Forest, 0.6 km east of Arizona Highway 67 just northeast of Forest Road 213 (36°30′38″N, 112°07′55″W) [FR 213 intersects AZ 67 30.6 km south of Jacob Lake] 48 Branchinecta kaibabensis Belk & Fugate, 2000 _________________________________________________________________________________ Kaibab National Forest, Forest Road 293, 36.40806° N, 112.26692° W [ca. 7.3 miles west of Kaibab Lodge] 42 Morchella kaibabensis Beug, T.A. Clem. & T.J. Baroni in Baroni et al., 2018 _________________________________________________________________________________ Lookout Canyon Road (422D), 5.6 km from (northwest of) Grand Canyon Highway (Arizona Route 67), T.35 N., R.2 E., Sec. 15 164 Castilleja kaibabensis N. Holmgren, 1973 _________________________________________________________________________________ Marble Flat 86 Penstemon virgatus pseudoptutus Crosswhite, 1967 _________________________________________________________________________________ Marble Flat, just north of North Rim Ranger Station 164 Castilleja kaibabensis N. Holmgren, 1973 _________________________________________________________________________________ 182 LOCALITIES FOR TYPE MATERIAL FROM NEAR GRAND CANYON NATIONAL PARK Nagle’s Ranch15 146 Ptelea saligna Green, 1906 _________________________________________________________________________________ Nagle’s Ranch15 (below, on edge of Buckskin Mountains) 178 Laphamia gracilis Jones, 1895 _________________________________________________________________________________ Nagle’s Ranch15 (road to) 174 Astragalus humistratus var. tenerrimus Jones, 1895 _________________________________________________________________________________ Ryan Ranger Station (near) Papilio indra kaibabensis Bauer, 1955 45 _________________________________________________________________________________ VT Meadows, 6.5 km south of Kaibab Lodge, T.34 N., R.3 E., Sec. 7 164 Castilleja kaibabensis N. Holmgren, 1973 _________________________________________________________________________________ VT Park 86 164 Penstemon virgatus pseudoptutus Crosswhite, 1967 Castilleja kaibabensis N. Holmgren, 1973 _________________________________________________________________________________ VT Ranch Tapinocyba kesimba Chamberlin, 1948 70 _________________________________________________________________________________ VT Ranger Station 86 164 Penstemon virgatus pseudoptutus Crosswhite, 1967 Castilleja kaibabensis N. Holmgren, 1973 _________________________________________________________________________________ 15 “Nagle’s Ranch was about 60 miles south of Kanab, Utah, on the west slope of the Buckskin Mountains (i. e. the Kaibab Plateau), and was the first watering place on the old wagon road to the Grand Canyon from Kanab. [. . .] The present wagon road now ascends the plateau many miles farther north.” (Kearney, Thomas H., and Peebles, Robert H. (and collaborators), 1942, Flowering plants and ferns of Arizona, U.S. Department of Agriculture, Miscellaneous Publication 423, p. 986.) 183 LOCALITIES FOR TYPE MATERIAL FROM NEAR GRAND CANYON NATIONAL PARK KANAB CANYON Kanab Canyon (west rim, south of Water Canyon, in depression caused by breccia pipe collapse, T38N R3W NW¼ SW¼ sect. 8) 235 Rosa stellata Wooton subsp. abyssa Phillips, 1992 _________________________________________________________________________________ Kanab Canyon (west rim, in small drainage 30 m from edge, T38N R3W NW¼ sect. 29 235 Rosa stellata Wooton subsp. abyssa Phillips, 1992 _________________________________________________________________________________ Kanab Canyon (west rim, T38N R3W sect. 30) 235 Rosa stellata Wooton subsp. abyssa Phillips, 1992 _________________________________________________________________________________ Kanab Wash “near the Colorado” [Arizona] 82 Echinocactus polycephalus var. xeranthemoides Coulter, 1896 _________________________________________________________________________________ LEES FERRY Lees Ferry 179 260 282 Astragalus kentrophyta var. coloradoensis Jones, 1902 Amsonia eastwoodiana Rydberg, 1913 Atriplex asterocarpa Stutz, Chu & Sanderson, 1994 _________________________________________________________________________________ Lees Ferry (near) 291 Perdita opacella Timberlake, 1956 _________________________________________________________________________________ MARBLE CANYON Navajo Bridge 11 13 Myndus yuccandus E. Ball, 1933 Yucanda miniata E. Ball, 1937 _________________________________________________________________________________ 184 LOCALITIES FOR TYPE MATERIAL FROM NEAR GRAND CANYON NATIONAL PARK NAVAJO NATION (westernmost) See also entries for Cameron (above). See also “Cedar Mountain” in the LOCALITIES FOR TYPE MATERIAL IN GRAND CANYON NATIONAL PARK list, noted as possibly being within the boundaries of the Navajo Nation. The Gap 116 Steatoda variata Gertsch, 1960 _________________________________________________________________________________ The Gap (near) 160 Cryptantha atwoodii Higgins, 1974 _________________________________________________________________________________ RED MOUNTAIN Red Mountain, 35 miles northwest of Flagstaff on U.S. Route 180 73 Mentzelia collomiae Christy, 1997 _________________________________________________________________________________ 185 LOCALITIES FOR TYPE MATERIAL FROM NEAR GRAND CANYON NATIONAL PARK 186 INDEX TO NOMINATE TAXA INDEX TO NOMINATE TAXA Names in bold indicate nominate taxa for which at least one locality reported for its types is within, or is with good probability from within, the present boundaries of Grand Canyon National Park. The names are original combinations, as published by the authors. The taxonomic phylum is appended to each name and author. Refer to the bibliography and annotated checklist for details. Red numbers at left indicate and cross-list the sequentially enumerated taxa in the bibliography (thus they reflect the alphabetical order by publication author). 163 235 159 183 198 59 278 89 241 303 238 64 87 47 277 141 320 149 100 22 95 310 43 256 128 216 aaronrossii, Euphorbia A. Holmgren & N. Holmgren, 1988 (Trachyophyta) abyssa, Rosa stellata Wooton subsp. Phillips, 1992 (Trachyophyta) abyssi, Opuntia Hester, 1943 (Trachyophyta) abyssus, Crotalus confluentus Klauber, 1930 (Chordata) ackerti, Aspiculuris Kruidenier & Mehra, 1959 (Nematoda) aequalis, Orphilus Casey, 1900 (Arthropoda) alliae, Megathymus Stallings & Turner, 1957 (Arthropoda) ambivalens, Galium munzii Hilend & Howell subsp. Dempster & Ehrendorfer, 1965 (Trachyophyta) americanus, Pilophorus Poppius, 1914 (Arthropoda) amoena, Dalea Watson, 1873 (Trachyophyta) angelica, Oreohelix yavapai Pilsbry & Ferriss, 1911 (Mollusca) angustulus, Prionus Casey, 1912 (Arthropoda) apache, Tibicen Davis, 1921 (Arthropoda) apicata, Delphacodes Beamer, 1948 (Arthropoda) aquilonalis, Vaejovis Stahnke, 1940 (Arthropoda) argentea, Ptelea Green, 1906 (Trachyophyta) arizonae, Cicindela rufiventris var. Wickham, 1899 (Arthropoda) arizonense, Penicillium Frisvad, Grijseels & J.C. Nielsen in Grijseels et al., 2016 (Ascomycota) arizonense, Petalium bistriatum Say var. Fall, 1905 (Arthropoda) arizonensis, Orbilia Baral & G. Marson in Baral, Weber & Marson, 2020 (Ascomycota) arizonensis, Ripersia Ehrhorn, 1899 (Arthropoda) arizonica, Lesquerella Watson, 1888 (Trachyophyta) arizonica, Protoventuria M. Barr, 1989 (Ascomycota) arizonicus, Chrysotimus H. Robinson, 1967 (Arthropoda) artipes, Astragalus A. Gray, 1878 (Trachyophyta) artuflava, Chelinidea vittiger Uhler var. McAtee, 1919 (Arthropoda) 187 INDEX TO NOMINATE TAXA 282 176 160 251 46 239 asterocarpa, Atriplex Stutz, Chu & Sanderson, 1994 (Trachyophyta) 265 287 88 68 53 156 129 130 302 91 153 309 62 35 185 90 barberi, Ochterus Schell, 1943 (Arthropoda) 32 270 54 267 94 16 314 225 118 98 136 319 187 259 71 232 154 73 113 179 calyptrata, Orbilia Baral & G. Marson in Baral, Weber & Marson, 2020 (Ascomycota) attenuata, Bigelovia howardi var. Jones, 1895 (Trachyophyta) atwoodii, Cryptantha Higgins, 1974 (Trachyophyta) atwoodii, Eriogonum thompsonae S. Wats. var. Reveal, 1974b (Trachyophyta) aurea, Opuntia Baxter, 1933 (Trachyophyta) avus, Pupilla syngenes Pilsbry & Ferriss, 1911 (Mollusca) barrowsii, Boletus Thiers & Smith in Thiers, 1976 (Basidiomycota) basala, Ballana DeLong, 1964 (Arthropoda) behlei, Aphonopelma Chamberlin, 1940 (Arthropoda) benedicti, Erax Bromley, 1940 (Arthropoda) betheli, Sonorella Henderson, 1914 (Mollusca) biennis, Actinella A. Gray, 1878 (Trachyophyta) bigelovii, Mirabilis A. Gray, 1886 (Trachyophyta) blinni, Stygobromus Wang & Holsinger, 2001 (Arthropoda) boreorarius, Thomomys bottae Durham, 1952 (Chordata) bradybaughi, Hesperochernes Harvey & Wynne, 2014 (Arthropoda) breviflorum, Androstephium Watson, 1873 (Trachyophyta) brevipennis, Discodemus Casey, 1908 (Arthropoda) bryantii, Astragalus Barneby, 1944 (Trachyophyta) bullatus, Deraeocoris Knight, 1921 (Arthropoda) bunni, Bruchomorpha Doering, 1939 (Arthropoda) canonicus, Melanoplus Scudder, 1897 (Arthropoda) canyonensia, Aphelonema convergens var. Bunn, 1930 (Arthropoda) canyonensis, Mentzelia Schenk, Hodgson & Hufford, 2013 (Trachyophyta) capitata, Oreocarya Eastwood, 1937 (Trachyophyta) casa, Athysanella (Gladionura) Ball & Beamer, 1939 (Arthropoda) castus, Polymedon Wheeler, 1899 (Arthropoda) cavicola, Archeolarca Muchmore, 1981 (Arthropoda) celsus, Dipodomys microps Goldman, 1924 (Chordata) circinans, Puccinia J. Ellis & Everhart, 1900 (Basidiomycota) cissodes, Schmaltzia Greene, 1905 (Trachyophyta) clarigaster, Camponotus acutirostris Wheeler, 1915 (Arthropoda) clavicornis, Ceratocapsus Knight, 1925 (Arthropoda) coccinatus, Penstemon Rydberg, 1909 (Trachyophyta) coconino, Neoantistea Chamberlin & Ivie, 1942 (Arthropoda) cocytus, Ptomaphagus Peck, 1973 (Arthropoda) cohni, Tuberochernes Harvey & Wynne, 2014 (Arthropoda) collomiae, Mentzelia Christy, 1997 (Trachyophyta) coloradensis, Misumenops Gertsch, 1933 (Arthropoda) coloradoensis, Astragalus kentrophyta var. Jones, 1902 (Trachyophyta) 188 INDEX TO NOMINATE TAXA 280 243 145 104 177 193 37 119 276 221 coloradoensis, Helix (Arionta) Stearns, 1890 (Mollusca) 132 41 29 271 111 293 273 300 1 268 226 “dactylophyllum”, Gilia “(n. sp.?)” A. Gray in Gray et al., 1861 (Trachyophyta) 260 175 144 38 134 249 242 237 eastwoodiana, Amsonia Rydberg, 1913 (Trachyophyta) 85 195 28 308 96 14 184 40 5 189 274 facialis, Bombylius Cresson, 1919 (Arthropoda) confertiflora, Oenothera Raven, 1962 (Trachyophyta) confinis, Ptelea Green, 1906 (Trachyophyta) conformis, Diplotaxis Fall, 1909 (Arthropoda) congesta, Laphamia Jones, 1895 (Trachyophyta) cowaniae, Parthenicus Knight, 1968 (Arthropoda) cremnophylax, Astragalus Barneby, 1948 (Trachyophyta) cupidineus, Dipodomys ordii Goldman, 1924 (Chordata) curatorium, Carex Stacey, 1937 (Trachyophyta) cypha, Gila R. Miller, 1946 (Chordata) damei, Cercyonis Barnes & Benjamin, 1926 (Arthropoda) delphinus, Orbilia Baral & G. Marson in Baral, Weber & Marson, 2020 (Ascomycota) denningi, Mycetophila Shaw, 1951 (Arthropoda) dentatus, Gorytes Fox, 1893 (Arthropoda) depressa, Perdita Timberlake, 1968 (Arthropoda) digna, Wiedemannia Sinclair, 2006 (Arthropoda) dilaticosta, Polymedon Van Duzee, 1927 (Arthropoda) dimidiatus, Ceratopogon C. Adams, 1903 (Arthropoda) dionysus, Neominois Scudder, 1878 (Arthropoda) dorothea, Neonympha dorothea Nabokov, 1942 (Arthropoda) ecristata, Oenothera triloba var. Jones, 1895 (Trachyophyta) elegans, Ptelea Green, 1906 (Trachyophyta) endopterus, Astragalus wootoni var. Barneby, 1949 (Trachyophyta) eremophila, Arabis Greene, 1900 (Trachyophyta) evanida, Fabriciella Reinhard, 1953 (Arthropoda) exiguus, Pilophorus Poppius, 1914 (Arthropoda) extremitatis, Oreohelix yavapai Pilsbry & Ferriss, 1911 (Mollusca) flaviatus, Phytocoris Knight, 1968 (Arthropoda) flexisoma, Orbilia Baral & G. Marson in Baral, Weber & Marson, 2020 (Ascomycota) flexuosus, Calochortus Watson, 1873 (Trachyophyta) formicarii, Dactylopius Ehrhorn 1899 (Arthropoda) fredonia, Athysanella Ball & Beamer, 1939 (Arthropoda) fulvus, Deraeocoris Knight, 1921 (Arthropoda) fureae, Coenonympha Barnes & Benjamin, 1926 (Arthropoda) furnissii, Phacelia Atwood, 2007 (Trachyophyta) fuscipennis, Pilophorus Knight, 1926 (Arthropoda) fusiforme, Eriogonum Small, 1906 (Trachyophyta) 189 INDEX TO NOMINATE TAXA 317 181 213 97 15 281 178 215 219 101 170 166 148 gelida, Formica fusca fusca var. Wheeler, 1913 (Arthropoda) 77 244a 190 6 34 137 79 266 157 158 7 106 havasupaiensis, Sclerocactus Clover, 1942 (Trachyophyta) 240 167 56 212 285 292 99 131 immaculata, Schinia Pogue, 2004 (Arthropoda) 194 juniperi, Bolteria Knight, 1968 (Arthropoda) 117 217 253 48 164 172 74 kaiba, Loxosceles Gertsch & Ennik, 1983 (Arthropoda) georgei, Nebria (Reductonebria) Kavanaugh, 2008 (Arthropoda) gilli, Chionaspis Liu & Kosztarab, 1987 (Arthropoda) gillii, Septogloeum D. Ellis, 1946 (Ascomycota) globosa, Athysanella Ball & Beamer, 1939 (Arthropoda) gloydi, Sonora semiannulata Stickel, 1938 (Chordata) gracilis, Laphamia Jones, 1895 (Trachyophyta) gracilis, Parasitorhabditis Massey 1974 (Nematoda) gracilis, Spilogale Merriam, 1890 (Chordata) grande, Catorama Fall, 1905 (Arthropoda) grande, Cinara Hottes, 1956 (Arthropoda) grandiscanyonensis, Arenivaga Hopkins, 2014 (Arthropoda) grisea, Morus Greene, 1911 (Trachyophyta) hesperia, Oenothera specuicola subsp. Raven, 1962 (Trachyophyta) hesperius, Phytocoris Knight, 1928 (Arthropoda) higginsii, Phacelia Atwood, 2007 (Trachyophyta) hirsuta, Paratriatoma Barber, 1938 (Arthropoda) hirtella, Schmaltzia Green, 1905 (Trachyophyta) hopkinsi, Triepeolus Cockerell, 1905 (Arthropoda) hualapaiensis, Mentzelia Schenk, Hodgson & Hufford, 2010 (Trachyophyta) hualapaiensis, Pyrgulopsis Herschler, Liu & Stevens, 2016 (Mollusca) hualpaensis, Opuntia Hester, 1943 (Trachyophyta) hughesii, Phacelia Atwood, 2007 (Trachyophyta) hunnewellii, Primula (Section Farinosae) Fernald, 1934 (Trachyophyta) impensa, Arenivaga Hopkins, 2014 (Arthropoda) incipiens, Trichochrous Casey, 1895 (Arthropoda) inflata, Norvellina bicolorata (Ball) var. Lindsay, 1938 (Arthropoda) infralabialis, Lampropeltis pyromelana Cope Tanner, 1953 (Chordata) inornata, Perdita Timberlake, 1962 (Arthropoda) isidiomuscigena, Physconia Esslinger, 2000 (Ascomycota) ivesiana, Phacelia (Coreanthus) A. Gray in Gray et al., 1861 (Trachyophyta) kaibabensis, Agave McKelvey, 1949 (Trachyophyta) kaibabensis, Andrena (Scaphandrena) Ribble, 1974 (Arthropoda) kaibabensis, Branchinecta Belk & Fugate, 2000 (Arthropoda) kaibabensis, Castilleja N. Holmgren, 1973 (Trachyophyta) kaibabensis, Cicindela pusilla Johnson, 1990 (Arthropoda) kaibabensis, Cordylanthus wrightii Chuang & Heckard, 1986 (Trachyophyta) 190 INDEX TO NOMINATE TAXA 108 182 227 258 50 42 248 45 115 155 220 252 288 123 3 223 180 2 257 70 196 84 kaibabensis, Coryphantha vivipara var. Fischer, 1979 (Trachyophyta) 297 295 10 4 307 102 247 60 143 lafontainei, Hadena (Hadena) Troubridge & Crabo, 2002 (Arthropoda) 30 93 246 284 26 19 150 315 318 286 262 macrodelphinus, Orbilia Baral & G. Marson in Baral, Weber & Marson, 2020 (Ascomycota) kaibabensis, Enderleinellus Kim, 1966 (Arthropoda) kaibabensis, Felis concolor E. Nelson & Goldman, 1931 (Chordata) kaibabensis, Lesquerella Rollins, 1982 (Trachyophyta) kaibabensis, Mischocyttarus flavitarsis var. Bequaert, 1932 (Arthropoda) kaibabensis, Morchella Beug, T.A. Clem. & T.J. Baroni in Baroni et al., 2018 (Ascomycota) kaibabensis, Morsea california Rehn & Grant, 1958 (Arthropoda) kaibabensis, Papilio indra Bauer, 1955 (Arthropoda) kaibabensis, Phidippus Gertsch, 1934b (Arthropoda) kaibabensis, Polistes canadensis var. Lynn, 1932 (Arthropoda) kaibabensis, Sciurus Merriam, 1904 (Chordata) kaibabensis, Scutellaria potosina var. Rhodes & Ayers, 2010 (Trachyophyta) kaibabensis, Suillus Thiers, 1976 (Basidiomycota) kaibabensis, Thomomys fossor Goldman, 1938 (Chordata) kaibabensis, Tipula (Lunatipula) C. Alexander, 1946 (Arthropoda) kaibabiae, Saxinis saucia Moldenke, 1970 (Arthropoda) kaibensis, Astragalus Jones, 1902 (Trachyophyta) kathyae, Plagiomimicus J. Adams & Lafontaine, 2009 (Arthropoda) kearneyi, Sisymbrium Rollins, 1953 (Trachyophyta) kesimba, Tapinocyba Chamberlin, 1948 (Arthropoda) knowltoni, Aphricus Knull, 1957 (Arthropoda) knowltoni, Ostrya Coville, 1894 (Trachyophyta) leptoscelis, Eleodes (Caverneleodes) Triplehorn, 1975 (Arthropoda) ligulifolia, Salix lutea var. C. Ball, 1921 (Trachyophyta) longicarpum, Thelypodium integrifolium (Nuttall) Endlicher subsp. Al-Shehbaz, 1973 (Trachyophyta) longifolia, Brickellia (Clavigera) Watson, 1873 (Trachyophyta) longulum, Catorama Fall, 1905 (Arthropoda) lowellii, Fraxinus Sargent in Rehder, 1917 (Trachyophyta) lugubris, Zopherodes Casey, 1907 (Arthropoda) lutescens, Ptelea Green, 1906 (Trachyophyta) macropetala, Fraxinus Eastwood, 1903 (Trachyophyta) macropetala, Fraxinus cuspidata var. Rehder, 1917 (Trachyophyta) macrosepala, Amsinckia Suksdorf, 1931 (Trachyophyta) maeandrina, Orbilia Baral & G. Marson in Baral, Weber & Marson, 2020 (Ascomycota) magnifica, Orbilia Baral & G. Marson in Baral, Weber & Marson, 2020 (Ascomycota) margaretae, Boharticus Grissell, 1983 (Arthropoda) maricopa, Pheidole desertorum var. Wheeler, 1906 (Arthropoda) maricopa, Pogonomyrmex californicus Buckley subsp. Wheeler, 1914 (Mollusca) mcdougallii, Flaveria Theroux, Pinkava & Keil, 1977 (Trachyophyta) megalocarpa, Juniperus utahensis var. Sargent, 1919 (Trachyophyta) 191 INDEX TO NOMINATE TAXA 218 188 165 13 294 245 316 127 114 250 304 27 83 21 31 229 39 megalotis, Hesperomys Merriam, 1890 (Chordata) 275 283 228 186 23 72 9 306 124 140 301 nakaharai, Heterotarsonemus Smiley & Moser, 1974 (Arthropoda) 289 67 18 291 25 36 occidentalis, Suillus Thiers, 1976 (Basidiomycota) 279 168 138 49 233 8 107 pachyphylla, Allionia Standley, 1909 (Trachyophyta) mellarius, Phytocoris Knight, 1925 (Arthropoda) memorabilis, Mentzelia N. Holmgren & P. Holmgren, 2002 (Trachyophyta) miniata, Yucanda E. Ball, 1937 (Arthropoda) minuta, Microsciasma Townsend, 1915 (Arthropoda) mohavenss, Panicum Reeder, 1991 (Trachyophyta) moki, Formica Wheeler, 1906 (Arthropoda) mokiacensis, Astragalus A. Gray, 1878 (Trachyophyta) mokiensis, Arctosa Gertsch, 1934a (Arthropoda) mortonianum, Eriogonum Reveal, 1974a (Trachyophyta) multijuga, Oenothera (Chylisma) Watson, 1873 (Trachyophyta) multimaeandrina, Orbilia Baral & G. Marson in Baral, Weber & Marson, 2020 (Ascomycota) multinervatus, Phellopterus Coulter & Rose, 1900 (Trachyophyta) multiphanosoma, Orbilia Baral & G. Marson in Baral, Weber & Marson, 2020 (Ascomycota) multitrapezoidea, Orbilia Baral & G. Marson in Baral, Weber & Marson, 2020 (Ascomycota) mychophilus, Otus asio Oberholser, 1937 (Chordata) myriorrhaphis, Astragalus cremnophylax Barneby var. Barneby, 1979 (Trachyophyta) nana, Amsinckia Suksdorf, 1931 (Trachyophyta) navajo, Acmaeodera G. Nelson & Westcott, 1995 (Arthropoda) navajo, Deraeocoris Knight, 1921 (Arthropoda) navajoana, Orbilia Baral & G. Marson in Baral, Weber & Marson, 2020 (Ascomycota) neofungorum, Mycetophila Chandler, 1993 (Arthropoda) neotomae, Orchopeas sexdentatus Augustson, 1943 (Arthropoda) newberryi, Peucedanum Watson, 1873 (Trachyophyta) nicholi, Thomomys bottae Goldman, 1938 (Chordata) nitida, Ptelea Green, 1906 (Trachyophyta) nitidus, Polymedon Van Duzee, 1927 (Arthropoda) occultum, Bembidion (Cyclolopha) Casey, 1918 (Arthropoda) ocellata, Orbilia Baral, G. Marson & E. Weber in Baral, Weber & Marson, 2020 (Ascomycota) opacella, Perdita Timberlake, 1956 (Arthropoda) ophiosoma, Orbilia Baral & G. Marson in Baral, Weber & Marson, 2020 (Ascomycota) oropedii, Astragalus lentiginosus Dougl. var. Barneby, 1945 (Trachyophyta) pagana, Arenivaga Hopkins, 2014 (Arthropoda) palida, Ptelea Green, 1906 (Trachyophyta) paradinei, Pediocactus Benson, 1957 (Trachyophyta) parashant, Ptomaphagus Peck & Wynne, 2013 (Arthropoda) parashantensis, Xylorhiza tortifolia (Torrey & Gray) Greene var. Atwood & Welsh, 2013 (Trachyophyta) parviflora, Adenostegia Ferriss, 1918 (Trachyophyta) 192 INDEX TO NOMINATE TAXA 255 312 122 197 199 105 20 69 162 230 103 55 313 231 44 171 236 152 58 66 61 86 305 17 parviflora, Crossosoma B. Robinson & Fernald, 1894 (Trachyophyta) 224 quercusgambellii, Gnomonia Monod, 1983 (Ascomycota) 135 254 222 121 261 57 80 78 298 296 recondita, Arabis Greene, 1900 (Trachyophyta) 146 126 76 112 133 173 saligna, Ptelea Green, 1906 (Trachyophyta) pauperitense, Pediomelum S.L. Welsh, M. Licher, & N. D. Atwood in Welsh & Licher, 2010 (Trachyophyta) peridoneus, Peromyscus crinitus Goldman, 1937 (Chordata) perognathi, Wellcomia Kruidenier & Mehra, 1958 (Nematoda) peromysci, Gongylonema Kruidenier & Peebles, 1958 (Nematoda) perplexus, Bruchus Fall, 1910 (Arthropoda) phanosoma, Orbilia Baral & G. Marson in Baral, Weber & Marson, 2020 (Ascomycota) phasmus, Aphonopelma Chamberlin, 1940 (Arthropoda) phillipsiana, Agave Hodgson, 2001 (Trachyophyta) pholeter, Eschatomoxys Thomas & Pape in Pape, Thomas & Aalbu, 2007 (Arthropoda) pingue, Catorama Fall, 1905 (Arthropoda) piniedulis, Melanophila Burke, 1907 (Arthropoda) pinionis, Astragalus Welsh & Thorne, 1977 (Trachyophyta) pinkavae, Opuntia Parfitt, 1997 (Trachyophyta) pletura, Acmaeodera W. Barr, 1972 (Arthropoda) poketa, Cinara Hottes, 1956 (Arthropoda) profundorum, Oreohelix yavapai Pilsbry & Ferriss, 1911 (Mollusca) prosopidis, Geron Hall & Evenhuis, 2003 (Arthropoda) pruddeni, Cryptorhopalum Casey, 1900 (Arthropoda) pruddeni, Stenosphenus Casey, 1912 (Arthropoda) pruddeni, Zopherodes Casey, 1907 (Arthropoda) pseudoptutus, Penstemon virgatus Crosswhite, 1967 (Trachyophyta) purpureus, Cymopterus Watson, 1873 (Trachyophyta) purshiae, Orbilia Baral & G. Marson in Baral, Weber & Marson, 2020 (Ascomycota) rectiramea, Silene Robinson, 1899 (Trachyophyta) reederi, Sonorella W. Miller, 1984 (Mollusca) repentinus, Castor canadensis Goldman, 1932 (Chordata) retrorsum, Pediomelum Rydberg in Rydberg & Pennell, 1919 (Trachyophyta) reversus, Trichochrous Casey, 1895 (Arthropoda) romainei, Myrmecocystus melliger subsp. semirufus Emery var. Cole, 1936 (Arthropoda) roseus, Sclerocactus havasupaiensis var. Clover, 1942 (Trachyophyta) rubra, Trioza Tuthill, 1939 (Arthropoda) rugicollis, Asidina Triplehorn & Brown, 1971 (Arthropoda) scaposus, Astragalus A. Gray, 1878 (Trachyophyta) schellbachi, Depressaria Gates, 1947 (Arthropoda) schellbachi, Speyeria atlantis Garth, 1949 (Arthropoda) simplicifolia, Rhus canadensis Marsh. var. Greene, 1890 (Trachyophyta) skillmani, Trogloderus Johnston, 2019 (Arthropoda) 193 INDEX TO NOMINATE TAXA 244 24 63 321 139 147 52 125 33 specuicola, Oenothera Raven, 1962 (Trachyophyta) 264 191 174 75 234 65 110 120 269 109 263 142 311 151 tapeats, Efferia Scarbrough, Stevens & Nelson, 2012 (Arthropoda) 200 12 utahensis, Andrena (Callandrena) LaBerge, 1967 (Arthropoda) 299 116 81 192 214 272 vanduzeei, Platylygus Usinger, 1931 (Arthropoda) 92 51 290 169 wetherillii, Corydalis Eastwood, 1902 (Trachyophyta) 82 xeranthemoides, Echinocactus polycephalus var. Coulter, 1896 (Trachyophyta) 11 yuccandus, Myndus E. Ball, 1933 (Arthropoda) spermoides, Orbilia Baral & G. Marson in Baral, Weber & Marson, 2020 (Ascomycota) spiculosus, Prionus Casey, 1912 (Arthropoda) stonei, Heteropogon Wilcox, 1965 (Arthropoda) straminea, Ptelea Green, 1906 (Trachyophyta) stygius, Senecio Greene, 1909 (Trachyophyta) subatrata, Priononyx Bohart, 1958 (Arthropoda) subcinereus, Astragalus A. Gray, 1878 (Trachyophyta) subovoidea, Orbilia Baral, Matočec & E. Weber in Baral, Weber & Marson, 2020 (Ascomycota) tenellus, Plagiognathus Knight, 1929 (Arthropoda) tenerrimus, Astragalus humistratus var. Jones, 1895 (Trachyophyta) tenuifolia, Encelia resinifera subsp. Curtis, 1998 (Trachyophyta) tenuifolis, Cordylanthus Pennell, 1940 (Trachyophyta) terminalis, Prionus Casey, 1912 (Arthropoda) terricola, Spironema Foissner & Foissner, 1993 (Hemimastigophora) tersus, Ammospermophilus leucurus Goldman, 1929 (Chordata) thoosa, Synchloe Scudder, 1878 (Arthropoda) toroweapensis, Echinocereus triglochidiatus var. Fischer, 1991 (Trachyophyta) toumeyi, Populus fremontii var. Sargent, 1919 (Trachyophyta) triptera, Ptelea Green, 1906 (Trachyophyta) trumbullensis, Astragalus lentiginosus Douglas ex Hook. var. Welsh & Atwood, 2001 (Trachyophyta) trumbullensis, Thomomys bottae Hall & Davis, 1934 (Chordata) utahnum, Hysteropterum cornutum var. E. Ball, 1935 (Arthropoda) variata, Steatoda Gertsch, 1960 (Arthropoda) varipennis, Trypeta Coquillett, 1902 (Arthropoda) varius, Phytocoris Knight, 1934 (Arthropoda) vauxae, Oreohelix yavapai Marshall, 1929 (Mollusca) voylesi, Pratherodesmus Shear in Shear, Taylor, Wynne & Krejca, 2009 (Arthropoda) wheeleri, Andricus Beutenmüller, 1907 (Arthropoda) wheeleri, Perdita Timberlake, 1928 (Arthropoda) wickhami, Luperodes Horn, 1893 (Arthropoda) ____________________________________________________________________________________ 194