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The grass genera of the world

L. Watson, T.D. Macfarlane, and M.J. Dallwitz

Notodanthonia Zotov

From the Greek notos (south), thus the southern Danthonias.

~ Danthonia sensu lato; Rytidosperma

Type species: Type: N. unarede (Raoul) Zotov.

Excluding Rytidosperma

Habit, vegetative morphology. Perennial; caespitose. The flowering culms leafless. Culms 20–80 cm high; herbaceous; cylindrical; unbranched above; 2–7 noded. Culm nodes exposed; glabrous. Culm internodes solid. Young shoots extravaginal, or intravaginal. Leaves mostly basal; non-auriculate; without auricular setae. Sheath margins free. Leaf blades linear-lanceolate; neither leathery nor flimsy; narrow; 0.3–2 mm wide; flat; not pseudopetiolate; without cross venation; persistent. Ligule present; a fringe of hairs. Contra-ligule absent.

Reproductive organization. Plants bisexual, all with bisexual spikelets; with hermaphrodite florets. The spikelets all alike in sexuality; hermaphrodite. Not viviparous.

Inflorescence. Inflorescence many spikeleted; paniculate; open; more or less irregular; espatheate; not comprising ‘partial inflorescences’ and foliar organs. Spikelet-bearing axes solitary. Spikelets solitary; not secund; pedicellate.

Female-fertile spikelets. Spikelets morphologically ‘conventional’; cuneate; green; compressed laterally; disarticulating above the glumes; disarticulating between the florets; with conventional internode spacings. Rachilla prolonged beyond the uppermost female-fertile floret; hairless; the rachilla extension with incomplete florets. Hairy callus present. The callus hairs white. Callus short, or long (more or less equalling the rhachilla); blunt.

Glumes two; relatively large; more or less equal; exceeding the spikelets; long relative to the adjacent lemmas; hairless; scabrous (along veins, especially midrib); pointed; not subulate; awnless; carinate; similar. Lower glume much exceeding the lowest lemma; 3–7 nerved. Upper glume 3–7 nerved. Spikelets with incomplete florets. The incomplete florets distal to the female-fertile florets. The distal incomplete florets merely underdeveloped; awnless. Spikelets without proximal incomplete florets.

Female-fertile florets 2–7. Lemmas similar in texture to the glumes; smooth; not becoming indurated; white in fruit; incised; 2 lobed (tapering into slender setae); deeply cleft; awned. Awns 1, or 3; median (if solitary), or median and lateral (if three); from a sinus; geniculate (while the lateral are non-geniculate); hairless; much longer than the body of the lemma; entered by several veins; persistent. Awn bases twisted; flattened. Lemmas hairy. The hairs not in tufts (the upper margin of the indumentum with much longer hairs than on the body, which is generally shortly felted with hairs); not in transverse rows. Lemmas non-carinate; without a germination flap; 9 nerved; with the nerves confluent towards the tip. Palea present; relatively long; not convolute; entire, or apically notched; awnless, without apical setae; thinner than the lemma; not indurated; 2-nerved; 2-keeled. Palea back glabrous, or scabrous, or hairy. Palea keels wingless; scabrous. Lodicules present; 2; free; fleshy; ciliate; not or scarcely vascularized. Stamens 3. Anthers 0.5–1.5 mm long. Ovary apically glabrous. Styles free to their bases; free. Style bases widely separated. Stigmas 2; white.

Fruit, embryo and seedling. Disseminule a caryopsis enclosed in but free of the lemma and palea. Fruit free from both lemma and palea; small; golden-brown; obovate; compressed dorsiventrally; glabrous; smooth. Hilum short (less than 1/3 of caryopsis length). Pericarp thin; fused. Embryo large; waisted.

Transverse section of leaf blade, physiology. C3; XyMS+.

Cytology. Chromosome base number, x = 6. 2n = 24. 2 ploid, or 4 ploid.

Classification. Watson & Dallwitz (1994): Arundinoideae; Danthonieae. Soreng et al. (2015): Danthonioideae; Danthonieae. 5 species.

Distribution, phytogeography, ecology. Australia and New Zealand.

Not commonly adventive. Mesophytic; shade species (light shade); glycophytic.

References, etc. Morphological/taxonomic: Notodanthonia as re-circumscribed by H.P. Linder, who provided this morphological description (1997). Leaf anatomical: studied by us - N. alpicola, N. caespitosa, N. carphoides, N. setacea.

Illustrations. • Notodanthonia carphoides, as Danthonia: Turner, Australian Grasses (1895). • Notodanthonia caespitosa, as Rytidosperma: Linder et al, Ann. Miss. Bot. Gard. 97 (2010). • Notodanthonia lemmas (penicillata, pilosa = Austrodanthonia, semiannularis = Notodanthonia).


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Cite this publication as: ‘Watson, L., Macfarlane, T.D., and Dallwitz, M.J. 1992 onwards. The grass genera of the world: descriptions, illustrations, identification, and information retrieval; including synonyms, morphology, anatomy, physiology, phytochemistry, cytology, classification, pathogens, world and local distribution, and references. Version: 25th January 2024. delta-intkey.com’.

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