DELTA home

Commercial timbers

H.G. Richter and M.J. Dallwitz

Sapium spp. - Old World species (Ludai, ebusok)

Nomenclature etc. EUPHORBIACEAE. Asia: Sapium baccatum, S. luzonicum, Shirakiopsis indica (Willd.) Esser (syn. S. indicum Willd.) Triadica cochinchinensis Lour. (syn. S. discolor); Africa: S. armatum, S. ellipticum. Trade and local names: Asia: ludai (MY); lelun (MM); bedi, ludahi (ID); balakat-sudat (PH); adamsali, larrna (IN). Africa: ebusok (CM); asep (GH); musasa (TZ). Not protected under CITES regulations.

Tree. Geographic distribution: India, Pakistan, Sri Lanka, Myanmar (Burma), Thailand, Laos, Vietnam, Cambodia, Indomalesia, tropical Africa.

General. Heartwood basically white or grey. Sapwood colour similar to heartwood colour. Density 0.25–0.43 g/cm³.

Vessels. Wood diffuse porous. Vessels arranged in no specific pattern, in multiples, commonly in short (2–3 vessels) radial rows or in radial rows of 4 or more. Average tangential vessel diameter 150–280 µm. Average number of vessels/mm² 2–10. Perforation plates simple. Intervessel pits alternate, average diameter (vertical) 10–14 µm. Vessel-ray pits with reduced borders or apparently simple, different from intervessel pits, rounded or angular. Tyloses present, thinwalled.

Tracheids and fibres. Fibres very thin-walled, or of medium wall thickness. Fibre pits mainly restricted to radial walls, simple to minutely bordered. Fibres non-septate.

Axial parenchyma. Axial parenchyma banded. Axial parenchyma bands forming a reticulate pattern with rays. Bands fine. Axial parenchyma paratracheal. Apotracheal axial parenchyma diffuse. Axial parenchyma as strands. Average number of cells per strand: 3–4.

Rays. Rays 12–20 per tangential mm, exclusively uniseriate. Height of large rays commonly 500 to 1000 µm, or commonly over 1000 µm. Rays composed of two or more cell types (heterocellular). Heterocellular rays with square and upright cells restricted to marginal rows, mostly 2–4 marginal rows of upright or square cells or with more than 4 marginal rows of upright or square cells. Some rays occasionally with biseriate central portion.

Storied structures. Storied structure absent.

Secretory structures. Oil and mucilage cells absent. Intercellular canals absent. Laticifers or tanniniferous tubes present, or absent. Latex tubes generally rare; in some Asian species forming large, macroscopically visible radial slits (latex traces).

Cambial variants. Included phloem absent.

Mineral inclusions. Crystals not observed. Silica present, as grains or in aggregates; in rays cells. Silica mostly as small particles in asian species, as large aggregates often filling the entire cell lumen in african species.

Illustrations. • Transverse section I. Sapium luzonicum. • Transverse section II. Sapium sp. (Africa). • Tangential section I. Sapium luzonicum. • Tangential section II. Sapium sp. (Africa). black dots in rays cells are large silica aggregates. • Radial section I. Sapium luzonicum. dark dots are small silica particles in ray cells. • Radial section II. Sapium sp. (Africa). • Silica. Sapium sp. (Africa). large silica aggregates in ray cells.


The interactive key allows access to the character list, illustrations, full and partial descriptions, diagnostic descriptions, differences and similarities between taxa, lists of taxa exhibiting specified attributes, summaries of attributes within groups of taxa, and geographical distribution.


Cite this publication as: ‘Richter, H.G., and Dallwitz, M.J. 2000 onwards. Commercial timbers: descriptions, illustrations, identification, and information retrieval. In English, French, German, Portuguese, and Spanish. Version: 9th April 2019. delta-intkey.com’.

Contents