Western Spotted Coralroot

Corallorhiza maculata var. occidentalis

Corallorhiza maculata var. occidentalis is an orchid with no chlorophyll and derives its carbohydrates from a parasitic relationship with fungi. The plant is reddish in color, fleshy and somewhat brittle with leaves that are reduced to mere scales. The flower petals are spotted with red dots (maculata = spotted).

Erect raceme of 6 to 50, ½-inch flowers. Flaring at the top is a central sepal flanked by a pair of petals, with two lateral spreading sepals just below them, the sepals slightly longer than the petals. The tips, edges and outer surface are a deep reddish brown to nearly red, consistent with the color of the ovaries and stem but more yellow on the inner surface, especially towards the base. A few darker purplish spots are scattered on the surfaces. The lower lip is white with purple spotting, the larger central lobe is an inverted egg shape (obovate), widest towards the tip, with a wrinkled surface and wavy edges, curling back along sides and tip. 2 short pointed oval lance lobes are at the base. The pollen sacs hug the inside of the curved center column.

The large conspicuous ovary and short stalk are subtended by a short lance-shaped bract up to 1/8 inch long that is often forked.

Leaves are reduced to several overlapping sheathes on the lower stem. Smooth throughout, leaves and stem range in color from paler brownish red to brighter reds. Stems are single to multiple clusters.

Ovaries develop into hanging elliptic capsules, often pinkish yellow at maturity.
Corallorhiza maculata var. occidentalis Corallorhiza maculata var. occidentalis flower detail. This plant was growing beneath Jack Pine (Pinus banksiana) and Black Spruce (Picea mariana) on a mesic site. There were hundreds of these orchids flowering beneath the trees. Corallorhiza maculata,Corallorhiza maculata var. occidentalis,Geotagged,Orchidaceae,Summer,United States,Western Spotted Coralroot,black spruce,coralroot,flower,forest,jack pine,orchid,pines

Appearance

Inflorescences: floral bracts averaging 1–2.8 mm, often forked. Flowers: middle lobe of lip distinctly expanded, ratio of width of dilated part to base of middle lobe greater than than 1.5; column curved, 3.5–7.8 × 1–1.3 mm at stigma; stigmatic surface oblong-pentagonal, 0.8 × 1.3 mm.
Corallorhiza maculata var. occidentalis Cluster of Corallorhiza maculata var. occidentalis flowering stalks growing beneath Jack Pine (Pinus banksiana) and Black Spruce (Picea mariana) on a mesic site. There were hundreds of these orchids flowering beneath the trees. Corallorhiza maculata,Corallorhiza maculata var. occidentalis,Geotagged,Orchidaceae,Summer,United States,Western Spotted Coralroot,black spruce,coralroot,flower,forest,jack pine,orchid,pines

Naming

Corallorhiza maculata var. occidentalis (Lindley) Ames, Enum. Orchids U.S. & Canada. 22. 1924.
Corallorhiza multiflora Nuttall var. occidentalis Lindley, Gen. Sp. Orchid. Pl., 534. 1840
Corallorhiza grab-hamii Cockerell
Corallorhiza leimbachiana Suksdorf
Corallorhiza maculata subsp. occidentalis (Lindley) Cockerell
Corallorhiza maculata var. fusca H. H. Bartlett
Corallorhiza maculata var. immaculata M. Peck
Corallorhiza maculata var. intermedia Farwell
Corallorhizamaculata var. punicea Bartlett
Corallorhiza multiflora var. flavida Peck
Corallorhiza maculata var. occidentalis in fruit Corallorhiza maculata var. occidentalis (Wester Coral-root Orchid) in a sunny opening in a mixed conifer/deciduous forest. Jack Pine (Pinus banksiana), Black Spruce (Piceae mariana), and Quaking Aspen (Populus tremuloides) were the most common trees in the canopy. This variety and C. m. var. maculata occur in Minnesota and other states as well. Also, var. maculata is yellowish and blooms about a month after var. occidentalis. Corallorhiza,Corallorhiza maculata,Corallorhiza maculata var. maculata,Corallorhiza maculata var. occidentalis,Geotagged,Minnesota,Summer,United States,Western Spotted Coralroot,coralroot,forest,orchid

Distribution

St. Pierre and Miquelon (variety unconfirmed); Alta., B.C., Man., N.B., Nfld. and Labr. (Nfld.), N.S., Ont., P.E.I., Que., Sask.; Ariz., Calif., Colo., Conn., Idaho, Ill., Maine, Mass., Mich., Minn., Mont., Nebr, Nev., N.H., N.Mex., N.Y., Oreg., Pa., S.Dak., Utah, Vt., Va., Wash., Wis., Wyo.

Behavior

Corallorhiza maculata var. occidentalis and other Corallorhiza are myco-heterotrophs parasitic upon fungi which supply them with the carbon compounds they need.

Habitat

Coniferous and mixed woods, and conifer plantations

Reproduction

By seeds.

Evolution

This variety and C. m. var. maculata occur in Minnesota and other states as well. Also, var. maculata is yellowish and blooms about a month after var. occidentalis.

References:

Some text fragments are auto parsed from Wikipedia.

http://www.efloras.org/florataxon.aspx?flora_id=1&taxon_id=242102227
https://www.minnesotawildflowers.info/flower/western-spotted-coralroot
https://www.calflora.org/cgi-bin/species_query.cgi?where-taxon=Corallorhiza+maculata+var.+occidentalis

Jonathan R Leake. Plants parasitic on fungi: Unearthing the fungi in myco-heterotrophs and debunking the 'saprophytic' plant myth. Mycologist 19(3):113-122. August 2005. https://www.researchgate.net/publication/222571975_Plants_parasitic_on_fungi_Unearthing_the_fungi_in_myco-heterotrophs_and_debunking_the_'saprophytic'_plant_myth
Taxonomy
KingdomPlantae
DivisionAngiosperms
ClassMonocots
OrderAsparagales
FamilyOrchidaceae
GenusCorallorhiza
SpeciesCorallorhiza maculata var. occidentalis