Classification
 Nomenclature
Scientific Name:
Veronica annulata (Petrie) Cheeseman, Man. New Zealand Fl., ed. 2, 819 (1925)
Synonymy:
  • Veronica armstrongii var. annulata Petrie, Trans. & Proc. New Zealand Inst. 45: 273 (1913)
  • Hebe annulata (Petrie) Andersen, Trans. New Zealand Inst. 56: 693 (1926)
  • Leonohebe annulata (Petrie) Heads, Bot. Soc. Otago Newsl. 5: 7 (1987)
Lectotype (designated by Ashwin, in Allan 1961): on rock face of northern slope of Takitimu Mts, at about 900 m altitude, L. Cockayne 7617, 13 Mar 1912, WELT 5347. Isolectotypes: K, CHR 328337, WELT 17493
Etymology:
The epithet annulata means ringed, a reference to the connate leaves that encircle the stem.
Vernacular Name(s):
whipcord hebe
 Description

Spreading low whipcord shrub to 0.5 m tall. Stems ascending or erect, glabrous except for narrow line of eglandular hairs at the connate leaf bases, and sometimes hairs bifarious or uniform at base of internodes. Leaf bud indistinct, its outer leaves fully grown, diverging. Leaves opposite-decussate, connate in pairs and encircling stem, appressed but not usually covering the obscure node above, scale-like; lamina coriaceous, broadly deltoid to rectangular, 1–2 mm long, 2–3 mm wide, more or less glossy green or yellowish-green above and beneath; veins not evident, although leaf sometimes longitudinally wrinkled when dry; surfaces glabrous; margin conspicuously shortly ciliate to ciliolate, entire; apex obtuse to truncate; base broad; petiole absent. Inflorescence a terminal spike, 2.5–7.0 mm long; flowers crowded, 4–10, all bisexual; bracts opposite-decussate and connate, broadly deltoid to semi-circular; pedicels absent. Calyx lobes 4, seemingly 3 because anterior pair fused, obtuse, 1.5–2.0 mm long, unequal, densely shortly eglandular-ciliate to -ciliolate. Corolla 4.5–6.0 mm diameter; tube white, 1.5–2.0 mm long, ≤ calyx, hairy inside; lobes 4, white, erecto-patent to recurved, unequal, elliptic or obovate, 2–3 mm long, obtuse to rounded; nectar guides absent. Stamen filaments white, 2.6–3.5 mm long; anthers pink to purple. Style glabrous, 2.8–5.0 mm long. Capsules latiseptate, obtuse to rounded, glabrous, 1.8–2.7 mm long, 1.5–1.9 mm at widest point. Seeds obovoid or irregular, weakly flattened, smooth, pale brown, 0.9–1.3 mm long.

 Recognition

Superficially, V. annulata plants resemble V. hectorii plants, but the latter differ in having their nodal joints concealed although well marked, and in having their anterior calyx lobes free. Several other whipcord hebes are characterised by fused anterior calyx lobes: V. salicornioides plants have longer internodes and a smooth surface to the leafy stem; V. armstrongii plants have more sub-acute to acuminate leaf apices, leaves not so strongly appressed, and more slender, leafy branches; V. ochracea plants have thicker, leafy stems, ovate to deltoid leaves, more yellowish to bronze coloration, and paler anthers, as well as a higher chromosome number and distribution in the northern South Island.

 Distribution

South Island: Otago (Criffel Range, Remarkables, Hector Mts); Southland (Takitimu Mountains). An old record from near Kurow, Waitaki Valley, has not been corroborated. Also, a cultivated specimen (CHR 331852), said to have originated at Hump Ridge, Southland, is not included in the distribution.

 Habitat

Sparse scrub and rock outcrops. Recorded elevations range from 710 to 1350 m.

 Biostatus
Indigenous (Endemic)
 Phenology

Flowers: October–January; fruits: December–March.

 Cytology

2n = 42 (see Bayly & Kellow 2006, as Hebe annulata).

 Notes

Veronica annulata is classified in V. subg. Pseudoveronica sect. Hebe and informally in the “Flagriformes” group, also known as “whipcord hebes” (Albach & Meudt 2010; Bayly & Kellow 2006). It is most likely related to V. armstrongii, V. salicornioides, and V. ochracea, similar whipcord hebes that have 2n = 42, 84 and the anterior calyx lobes fused together.

 Bibliography
Albach, D.C.; Meudt, H.M. 2010: Phylogeny of Veronica in the Southern and Northern Hemispheres based on plastid, nuclear ribosomal and nuclear low-copy DNA. Molecular Phylogenetics and Evolution 54: 457–471.
Andersen, J. 1926: Popular names of New Zealand plants. Transactions of the New Zealand Institute 56: 659–714.
Bayly, M.J.; Kellow, A.V. 2006: An Illustrated Guide to New Zealand Hebes. Te Papa Press, Wellington.
Cheeseman, T.F. 1925: Manual of the New Zealand Flora. Edition 2. Government Printer, Wellington.
de Lange, P.J.; Norton, D.A.; Courtney, S.P.; Heenan, P.B.; Barkla, J.W.; Cameron, E.K.; Hitchmough, R.; Townsend, A.J. 2009: Threatened and uncommon plants of New Zealand (2008 revision). New Zealand Journal of Botany 47: 61–96. [as Hebe annulata (Petrie) Cockayne & Allan] [Naturally uncommon]
de Lange, P.J.; Norton, D.A.; Heenan, P.B.; Courtney, S.P.; Molloy, B.P.J.; Ogle, C.C.; Rance, B.D. 2004: Threatened and uncommon plants of New Zealand. New Zealand Journal of Botany 42(1): 45–76. [as Hebe annulata (Petrie) Cockayne & Allan]
de Lange, P.J.; Rolfe, J.R.; Barkla J.W.; Courtney, S.P.; Champion, P.D.; Perrie, L.R.; Beadel, S.N.; Ford, K.A.; Breitwieser, I.; Schönberger, I.; Hindmarsh-Walls, R.; Heenan, P.B.; Ladley, K. 2018: Conservation status of New Zealand indigenous vascular plants, 2017. New Zealand Threat Classification Series. No. 22. [Naturally Uncommon]
de Lange, P.J.; Rolfe, J.R.; Champion, P.D.; Courtney, S.P.; Heenan, P.B.; Barkla, J.W.; Cameron, E.K.; Norton, D.A.; Hitchmough, R.A. 2013: Conservation status of New Zealand indigenous vascular plants, 2012. New Zealand Threat Classification Series 3. Department of Conservation, Wellington. [as Hebe annulata (Petrie) Andersen] [Naturally Uncommon]
Garnock-Jones, P.J. 2023: Veronica. In: Breitwieser, I. (ed.) Flora of New Zealand – Seed Plants. Fascicle 9. Manaaki Whenua Press, Lincoln.
Garnock-Jones, P.J.; Albach, D.; Briggs, B.G. 2007: Botanical names in Southern Hemisphere Veronica (Plantaginaceae): sect. Detzneria, sect. Hebe, and sect. Labiatoides. Taxon 56: 571–582.
Heads, M. 1987: New names in New Zealand Scrophulariaceae. Botanical Society of Otago Newsletter 5: 4–11.
Petrie, D. 1913: Descriptions of new species and varieties of native phanerogams. Transactions and Proceedings of the New Zealand Institute 45: 265–275.