Classification
 Nomenclature
Scientific Name:
Veronica scopulorum (Bayly, de Lange & Garn.-Jones) Garn.-Jones in Garnock-Jones et al., Taxon 56: 579 (2007)
Synonymy:
  • Hebe scopulorum Bayly, de Lange & Garn.-Jones in Bayly et al., New Zealand J. Bot. 40: 586 (2002)
Holotype: New Zealand, North Island, South Auckland, Kawhia region, Rock Peak, 520 m, limestone bluffs around summit, M. J. Bayly 1444, P. J. Garnock-Jones & P. J. de Lange, 10 Oct 2000, WELT 82488/A. Isolectotypes: AK, CHR 549429
Etymology:
The epithet scopulorum means of the crags, a reference to the distinctive habitat characteristic of this species.
Vernacular Name(s):
Awaroa koromiko
 Description

Small, bushy shrub to 0.7 m tall. Stems erect to ascending, eglandular-pubescent; hairs bifarious. Leaf bud distinct, its leaves appressed at margins until fully grown; sinus acute. Leaves opposite-decussate to sub-distichous, erecto-patent to spreading; lamina sub-coriaceous, linear-elliptic to elliptic to narrowly oblanceolate, 14–55 mm long, 4–16 mm wide, glossy green to dark green above, dull glaucous beneath, midrib evident; surfaces glabrous except for  eglandular hairs along midrib above; margin glabrous except for eglandular hairs on petiole, minutely papillate, entire; apex acute to obtuse and plicate-mucronate, sometimes weakly; base cuneate; petiole 2–5 mm long. Inflorescence a lateral raceme, usually tripartite or sometimes simple or compound, 15–46 mm long; flowers crowded, 7–40, all bisexual; bracts opposite-decussate below, becoming alternate above, linear to lanceolate or rarely ovate, slightly < to slightly > pedicels; pedicels erecto-patent, 0.5–4.0 mm long, eglandular-hairy all around. Calyx lobes 4, sometimes the anterior pair fused in lower ⅓, acute, 1.5–2.3 mm, sub-equal, eglandular-ciliolate or mixed glandular- and eglandular-ciliolate. Corolla 6–8 mm diameter; tube white, 3.0–4.2 mm long, > calyx, glabrous; lobes 4, pale purplish or white, sub-erect to spreading, sub-equal, lanceolate to elliptic to oblong, 2.5–3.5 mm long, sub-acute; nectar guides absent. Stamen filaments white, 3–5 mm long; anthers pale purplish or white. Style glabrous, 6–9 mm long. Capsules latiseptate, acute, glabrous, 3.2–4.5 mm long, 2–3 mm at widest point. Seeds ellipsoid to ovoid or oblong, flattened, smooth, pale brown, 1.1–1.4 mm long.

 Recognition

Veronica scopulorum is one of only two North Island species of Veronica that are characterised by glaucous leaves; the other is V. colensoi. V. colensoi plants differ in having leaves that are glaucous on both surfaces, corolla tubes about equalling the calyx, and glabrous calyx margins.

V. scopulorum leaves are deeply folded along the midrib to give a very deep groove above and a very prominent midrib beneath, and each edge of the lamina is curved to make the leaf m-shaped in cross-section. For this reason, the leaf bud is tetragonous with concave faces in section, and this contrasts with the similar V. rigidula of Nelson, which has buds that are also tetragonous but with convex faces.

The bark of older stems of V. scopulorum plants is thick and corky, and on young stems the swelling below the leaf is transversely wrinkled.

 Distribution

North Island: Auckland (West Waikato near Kawhia, on outcrops along the ridges east and west of Awaroa Valley).

 Habitat

Outcrops and cliffs of limestone on ridges in forest. Recorded elevations range from 335 to 525 m.

 Biostatus
Indigenous (Endemic)
 Phenology

Flowers: September–November; fruits: September–April.

 Cytology

2n = 40 (see Bayly & Kellow 2006, as Hebe scopulorum).

 Notes

Veronica scopulorum is classified in V. subg. Pseudoveronica sect. Hebe and the informal group “Apertae” (small-leaved) (Albach & Meudt 2010; Bayly & Kellow 2006). ITS sequence data suggest V. scopulorum and V. colensoi are sister species; this is supported by their close similarity and North Island distributions. Beyond that, their relationships are likely to be with other species that are characterised by glaucous leaves, all of which occur in the South Island, such as V. rigidula .

 Bibliography
Albach, D.C.; Meudt, H.M. 2010: Phylogeny of Veronica in the Southern and Northern Hemispheres based on plastid, nuclear ribosomal and nuclear low-copy DNA. Molecular Phylogenetics and Evolution 54: 457–471.
Bayly, M.J.; Kellow, A.V. 2006: An Illustrated Guide to New Zealand Hebes. Te Papa Press, Wellington.
Bayly, M.J.; Kellow, A.V.; Mitchell, K.A.; Markham, K.R.; de Lange, P.J.; Harper, G.E.; Garnock-Jones, P.J.; Brownsey, P.J. 2002: Descriptions and flavonoid chemistry of new taxa in Hebe sect. Subdistichae (Scrophulariaceae). New Zealand Journal of Botany 40(4): 571–602.
de Lange, P.J.; Heenan, P.B.; Given, D.R.; Norton, D.A.; Ogle, C.C.; Johnson, P.N.; Cameron, E.K. 1999: Threatened and uncommon plants of New Zealand. New Zealand Journal of Botany 37: 603–628. [as Hebe aff. rigidula (WAIK 5822; Awaroa Valley); Hebe scopulorum Bayly, de Lange & Garn.-Jones]
de Lange, P.J.; Norton, D.A.; Courtney, S.P.; Heenan, P.B.; Barkla, J.W.; Cameron, E.K.; Hitchmough, R.; Townsend, A.J. 2009: Threatened and uncommon plants of New Zealand (2008 revision). New Zealand Journal of Botany 47: 61–96. [as Hebe scopulorum Bayly, de Lange & Garn.-Jones] [Naturally uncommon]
de Lange, P.J.; Norton, D.A.; Heenan, P.B.; Courtney, S.P.; Molloy, B.P.J.; Ogle, C.C.; Rance, B.D. 2004: Threatened and uncommon plants of New Zealand. New Zealand Journal of Botany 42(1): 45–76. [as Hebe scopulorum Bayly, de Lange & Garn.-Jones]
de Lange, P.J.; Rolfe, J.R.; Barkla J.W.; Courtney, S.P.; Champion, P.D.; Perrie, L.R.; Beadel, S.N.; Ford, K.A.; Breitwieser, I.; Schönberger, I.; Hindmarsh-Walls, R.; Heenan, P.B.; Ladley, K. 2018: Conservation status of New Zealand indigenous vascular plants, 2017. New Zealand Threat Classification Series. No. 22. [Declining]
de Lange, P.J.; Rolfe, J.R.; Champion, P.D.; Courtney, S.P.; Heenan, P.B.; Barkla, J.W.; Cameron, E.K.; Norton, D.A.; Hitchmough, R.A. 2013: Conservation status of New Zealand indigenous vascular plants, 2012. New Zealand Threat Classification Series 3. Department of Conservation, Wellington. [as Hebe scopulorum Bayly, de Lange & Garn.-Jones] [Naturally Uncommon]
Garnock-Jones, P.J. 2023: Veronica. In: Breitwieser, I. (ed.) Flora of New Zealand – Seed Plants. Fascicle 9. Manaaki Whenua Press, Lincoln.
Garnock-Jones, P.J.; Albach, D.; Briggs, B.G. 2007: Botanical names in Southern Hemisphere Veronica (Plantaginaceae): sect. Detzneria, sect. Hebe, and sect. Labiatoides. Taxon 56: 571–582.