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1a. Lip T-shaped, epichile lobes diverging at right angles to axis of lip .................................................................... 1. A. pingbianensis 1b. Lip Y-shaped, epichile lobes diverging at an acute angle to one another. 2a. Mesochile of lip with entire or occasionally shallowly notched flanges. 3a. Lip 1322 mm, yellow; spur 23.2 mm, apex obtuse ............................................................................... 2. A. burmannicus 3b. Lip ca. 10 mm, white; spur 46 mm, apex deeply bilobed ........................................................................ 3. A. koshunensis 2b. Mesochile of lip with pectinate or filamentous flanges. 4a. Flowers resupinate; ovary twisted. 5a. Mesochile flange composed of 48 teeth, each tooth ca. 1 mm .......................................................... 4. A. xingrenensis 5b. Mesochile flange 39-laciniate, each filament 28 mm. 6a. Ovary and pedicel ca. 8 mm; flange filaments of lip mesochile 23.5 mm ................................ 5. A. baotingensis 6b. Ovary and pedicel 1014 mm; flange filaments of lip mesochile 48 mm. 7a. Lip pale yellowish white; mesochile flanges 4- or 5-laciniate; spur 24 mm ........................ 6. A. formosanus 7b. Lip white; mesochile flanges 79-laciniate; spur ca. 8 mm .................................................... 7. A. hainanensis 4b. Flowers usually erect, not resupinate; ovary not twisted. 8a. Lip pinkish white, mesochile flanges 68-laciniate, each filament 2.56 mm ...................................... 8. A. roxburghii 8b. Lip white, mesochile flanges fimbriate, composed of 25 teeth, each tooth 1.53 mm. 9a. Epichile lobes 89 67 mm .......................................................................................................... 9. A. chapaensis 9b. Epichile lobes 56 25 mm. 10a. Leaves 0.72.6 0.62.1 cm; epichile lobes flabellate, ca. 5 mm wide ......................... 10. A. zhejiangensis 10b. Leaves 3.54 33.2 cm; epichile lobes oblong to oblanceolate, ca. 2 mm wide ............... 11. A. emeiensis 1. Anoectochilus pingbianensis K. Y. Lang, Acta Phytotax. Sin. 34: 556. 1996. ping bian jin xian lan Plants 1518 cm tall. Stem erect or ascending, 34 mm in diam., 4- or 5-leaved. Leaves abaxially pale green or pale red, adaxially dark green with gold venation, ovate, slightly oblique, 34.5 23 cm, apex acute; petiole-like base and tubular sheath 1.52 cm. Peduncle ca. 9 cm, with 2 or 3 pale red sheathing sterile bracts; rachis ca. 4 cm, few to 8-flowered;
floral bracts ovate-lanceolate, 1013 mm, ca. as long as ovary, outer surface pubescent, apex acuminate. Flowers resupinate; ovary and pedicel twisted, cylindric-fusiform, 1012 mm, densely pubescent. Sepals pink, outer surface pubescent, 1-veined; dorsal sepal ovate, cymbiform, ca. 6 4 mm, apex acute; lateral sepals spreading, oblong, slightly oblique, ca. 8 3.5 mm, apex acute. Petals white, ovate, slightly oblique, ca. 6 2 mm, 1-veined, apex acute; lip white, T-shaped, ca. 9 mm; hypochile ca. 2 mm; mesochile ca. 2.8 mm; flanges laciniate, composed of 5 or 6 narrow filaments; epichile transversely dilated, broader than long, 2-lobed; lobes diverging at right angles to axis of lip, narrowly oblong, entire, ca. 5.5 1.2 mm, apex obtuse; spur conic, ca. 3 mm, mouth ca. 2 mm in diam., apex obtuse, containing 2 narrowly elliptic, fleshy, stalked calli near entrance. Column ovoid, ca. 2.8 mm, with 2 lamellate wings; stigma lobes suborbicular. Fl. Oct.
Shaded humid places in forests; ca. 1500 m. SE Yunnan.
2. Anoectochilus burmannicus Rolfe, Bull. Misc. Inform. Kew 1922: 24. 1922. dian nan jin xian lan Plants 1630 cm tall. Roots fibrous, to 9 cm. Stem erect, 36-leaved. Leaves abaxially pinkish red, adaxially blackish green with limited bronze venation, broadly ovate to ovateelliptic, slightly oblique, 38 1.85 cm, apex acute; petiolelike base and tubular sheath 1.52.8 cm. Peduncle 914 cm, with 2 or 3 pale red sterile sheathing bracts; rachis 3.56 cm, 3 12-flowered; floral bracts ovate-lanceolate, often sheathing ovary at base, 914 mm, shorter than ovary, outer surface sparsely pubescent, apex acuminate. Flowers erect, not resupinate; ovary and pedicel narrowly fusiform, not twisted, 1115 mm, pubescent. Sepals pink, outer surface pubescent, 1-veined; dorsal sepal broadly ovate, cymbiform, 4.56 45 mm, apex subobtuse; lateral sepals spreading, oblong-elliptic, oblique, 6 7.5 35 mm, apex subacute. Petals yellowish white, broadly ovate-elliptic, strongly oblique, falcate, 56 34 mm, 1veined, apex obtuse; lip yellow, Y-shaped, 1322 mm; hypochile 23 mm; mesochile reflexed at right angle from apex of hypochile, 59 mm; flanges narrow, margin entire, terminating in small, slightly revolute auricles at apex of mesochile; epichile longitudinally dilated, 2-lobed; lobes diverging at an acute angle to one another, oblong, oblanceolate, or spatulate, 69 23 mm, entire, apex obtuse; spur conic, 23.2 mm, mouth 1.5 2.2 mm in diam., apex obtuse, containing 2 subelliptic, sessile calli near entrance. Column broadly ovoid, ca. 3 mm, with 2 lamellate wings; stigma lobes elliptic. Fl. SepDec.
Damp places in evergreen broad-leaved forests, valleys; 1000 2200 m. S Yunnan [Laos, Malaysia (Peninsular), Myanmar, Thailand].
cm, with 2 or 3 sheathing sterile bracts, reddish brown; rachis ca. 4.5 cm, 5- or 6-flowered; floral bracts pinkish, ovate to ovate-lanceolate, ca. 7 mm, shorter than ovary, outer surface pubescent. Flowers suberect, not resupinate, white; ovary and pedicel cylindric-fusiform, not twisted, ca. 12 mm, pubescent. Sepals reddish brown, pubescent on outer surface, 1-veined; dorsal sepal orbicular-ovate, cymbiform, 55.5 34 mm, apex acute to cuspidate; lateral sepals spreading, narrowly elliptic, oblique, 78 3.54 mm, apex acute. Petals white, ovate-elliptic, strongly oblique, falcate, 5.56 1.52 mm, 1-veined, apex cuspidate and incurved; lip white, Y-shaped, ca. 10 mm; hypochile 12 mm; mesochile reflexed at right angle from apex of hypochile, 34 mm; flanges oblong to linear-lanceolate, 45 ca. 2 mm, entire or sometimes shallowly notched at apex; epichile longitudinally dilated, 2-lobed; lobes diverging at an acute angle to one another, oblong, 79 3.53.7 mm, apex obtuse to slightly mucronate; spur tubular-conic, 46 mm, mouth ca. 2 mm in diam., apex deeply bilobed, containing 2 irregular, stalked calli. Column ovoid, 33.5 mm, with 2 lamellate wings; stigma lobes orbicular. Fl. JulOct.
Broad-leaved evergreen forests; 7002000 m. Taiwan.
4. Anoectochilus xingrenensis Z. H. Tsi & X. H. Jin, Acta Phytotax. Sin. 40: 84. 2002. xing ren jin xian lan Plants 1520 cm tall. Stem ascending, 23 mm in diam., 25-leaved. Leaves adaxially dark green with golden venation, ovate to orbicular-ovate, 25 23 cm, apex apiculate; petiolelike base and tubular sheath 1.52 cm. Peduncle ca. 12 cm, with 2 or 3 sterile sheathing bracts; rachis 35 cm, 35-flowered; floral bracts pink, ovate-lanceolate, 1012 mm, subequal in length to ovary. Flowers resupinate; ovary and pedicel twisted, 1013 mm, pubescent. Sepals pinkish red, outer surface pubescent; dorsal sepal ovate, concave, 57 35 mm, apex acute; lateral sepals suboblong, slightly oblique, 79 3.55 mm, apex acute. Petals white, ovate, oblique, 68 23 mm, 1veined, apex cuspidate; lip white, Y-shaped, 1318 mm; hypochile ca. 3 mm; mesochile ca. 3 mm; flanges pectinate, composed of 48 short teeth, each tooth ca. 1 mm; epichile longitudinally dilated, 2-lobed; lobes diverging at an acute angle to one another, flabellate, 67 2.53.5 mm, apical margin slightly incised; spur narrowly conic, 34 mm, mouth ca. 2 mm in diam., containing 2 irregular calli. Column ca. 1 mm. Fl. Aug.
Forests; ca. 1200 m. SW Guizhou (Xingren).
5. Anoectochilus baotingensis (K. Y. Lang) Ormerod, Taiwania 48: 87. 2003. bao ting jin xian lan Anoectochilus roxburghii (Wallich) Lindley var. baotingensis K. Y. Lang, Acta Phytotax. Sin. 34: 557. 1996. Plants 1517 cm tall. Stem erect, 2- or 3-leaved. Leaves purplish red, suborbicular, ca. 3 2.5 mm, apex acute; petiolelike base and tubular sheath ca. 0.6 cm. Peduncle ca. 8 cm, with 3 sheathing sterile bracts; rachis ca. 1 cm, 3-flowered; floral bracts ovate-lanceolate, 912 mm, ca. as long as to exceeding ovary, apex acute. Flowers resupinate, white; ovary and pedicel
3. Anoectochilus koshunensis Hayata, Icon. Pl. Formosan. 4: 104. 1914. heng chun yin xian lan Odontochilus koshunensis (Hayata) S. S. Ying. Plants ca. 20 cm tall. Stem ascending, 24-leaved. Leaves abaxially reddish purple, adaxially blackish green with white venation, ovate to orbicular-ovate, 34 23 cm, apex acute; petiole-like base and tubular sheath ca. 1 cm. Peduncle 710
twisted, cylindric-fusiform, ca. 8 mm, pubescent. Sepals 1veined, outer surface pubescent; dorsal sepal ovate, cymbiform, ca. 6.5 2.5 mm, apex acuminate; lateral sepals spreading, oblong-elliptic, ca. 7.5 3 mm, apex acute. Petals oblong-cuneate, strongly oblique, ca. 6.5 2.5 mm, 1-veined, apex ligulateacute; lip white, Y-shaped, ca. 17.5 mm; hypochile rectangular, ca. 5 mm, with lateral lobes ca. 1.5 mm wide; mesochile ca. 3 mm; flanges 3-laciniate, each filament 23.5 mm; epichile longitudinally dilated, 2-lobed; lobes diverging at an acute angle to one another, ligulate, ca. 8 2.5 mm, entire, apex obtuse; spur conic, 33.5 mm, apex obtuse, containing 2 large, circular, carunculate calli. Column ovoid, 56 mm, with 2 oblongelliptic wings; stigma lobes deltoid. Fl. Apr.
Soil-covered rocks in forests; 300400 m. Hainan.
als olive-green with white margins, elliptic-falcate, oblique, 8 8.5 ca. 3.6 mm, apex acuminate; lip white, Y-shaped, ca. 16 mm; hypochile ca. 2.5 mm; mesochile ca. 4.5 mm; flanges 79laciniate, each filament 48 mm; epichile longitudinally dilated, 2-lobed; lobes diverging at an acute angle to one another, ligulate-elliptic, ca. 9 3.5 mm, apex truncate; spur cylindric-conic, ca. 8 mm, mouth ca. 3.5 mm in diam., apex distinctly bilobed, containing 2 trapeziform calli. Column stout, ca. 4 mm, with 2 semicircular wings. Fl. Jan (in cultivation).
Damp shady places in montane forests. Hainan.
8. Anoectochilus roxburghii (Wallich) Lindley, Ill. Bot. Himal. Mts. 368. 1839. jin xian lan Chrysobaphus roxburghii Wallich, Tent. Fl. Napal. 37. 1826; Anoectochilus yungianus S. Y. Hu; Zeuxine roxburghii (Wallich) M. Hiroe. Plants 820 cm tall. Stem erect, 25-leaved. Leaves abaxially pale purplish red, adaxially dark green to blackish purple with pinkish golden reticulate venation, ovate to ovate-orbicular, 1.34.5 0.83.6 cm, apex acute; petiole-like base and tubular sheath 0.61.8 cm. Peduncle 715 cm, with 2 or 3 pinkish sterile bracts; rachis 15 cm, 210-flowered; floral bracts pinkish, ovate-lanceolate to lanceolate, 69 mm, shorter than ovary, apex acuminate. Flowers usually erect, not resupinate; ovary and pedicel narrowly cylindric, not twisted, 1015 mm, pubescent. Sepals pinkish, outer surface pubescent, 1-veined; dorsal sepal ovate, cymbiform, ca. 6 2.53 mm, apex acuminate; lateral sepals spreading, oblong to oblong-elliptic, 78 2.53 mm, apex subacute. Petals pinkish white, ovate-elliptic, strongly oblique, 78 22.5 mm, 1-veined, apex acuminate; lip pinkish white, Y-shaped, 1016 mm; hypochile 24 mm; mesochile reflexed at right angle from apex of hypochile, 45 mm; flanges 68-laciniate, each filament 2.56 mm; epichile longitudinally dilated, 2-lobed; lobes diverging at an acute angle to one another, obovate-oblong to spatulate, entire, 5.58 1.52.2 mm, apex obtuse; spur conic, 34.5 mm, apex shallowly bilobed, mouth ca. 1.5 mm in diam., containing 2 irregular, fleshy calli near entrance. Column ovoid, ca. 2.5 mm, with 2 broad, lamellate wings; stigma lobes elliptic. Fl. AugDec. 2n = 15 + 23B, 30.
Evergreen broad-leaved forests, damp valleys; 1001600 m. Fujian, Guangdong, Guangxi, Hainan, Hunan, Jiangxi, Sichuan, SE Xizang, Yunnan, Zhejiang [Bangladesh, Bhutan, India, Japan, Laos, Nepal, Thailand, Vietnam].
6. Anoectochilus formosanus Hayata, Icon. Pl. Formosan. 4: 101. 1914. tai wan yin xian lan Plants 1120 cm tall. Stem ascending, 24-leaved. Leaves abaxially reddish purple, adaxially dark green with white venation, ovate to ovate-orbicular, 25 13.5 cm, apex acute; petiole-like base and tubular sheath 0.81.6 cm. Peduncle 1015 cm, with 2 or 3 sheathing sterile bracts, reddish brown; rachis 1.55 cm, 35-flowered; floral bracts ovate-lanceolate, 810 mm, shorter than ovary, outer surface hairy, apex acuminate. Flowers resupinate, pale yellowish white; ovary and pedicel twisted, fusiform, 1014 mm, pubescent. Sepals reddish brown, pubescent on outer surface; dorsal sepal suborbicular, cymbiform, 67 56 mm, apex acute; lateral sepals narrowly elliptic, oblique, 810 56 mm, apex acute. Petals white, ovate, strongly oblique, falcate, ca. 8 2.6 mm, apex cuspidate; lip pale yellowish white, Y-shaped, 1216 mm; hypochile ca. 2 mm; mesochile 56 mm; flanges 4- or 5-laciniate, each filament 46 mm; epichile longitudinally dilated, 2-lobed; lobes diverging at an acute angle to one another, linear-oblong to rhombic-oblong, 67.5 1.53 mm, apex obtuse; spur conic, 24 mm, mouth ca. 3 mm in diam., apex shallowly bilobed, containing 2 carunculate calli. Column ovoid, ca. 2 mm. Fl. Oct Nov. 2n = 24, 40.
Broad-leaved evergreen forests, bamboo forests; 5001500 m. Taiwan [Japan (Ryukyu Islands)].
7. Anoectochilus hainanensis H. Z. Tian, F. W. Xing & L. Li, Ann. Bot. Fenn. 45: 220. 2008. hai nan kai chun lan Plants ca. 12 cm tall. Stem ascending, 3- or 4-leaved. Leaves abaxially lilac, adaxially velvety blackish green with dense golden reticulate venation, ovate, oblique, 33.5 1.8 2.2 cm, apex acute; petiole-like base and tubular sheath 1.21.5 cm. Peduncle 33.5 cm, with 2 or 3 sterile bracts, pubescent; rachis 34 cm, 46-flowered; floral bracts ovate-lanceolate, 69 mm, shorter than ovary, outer surface hairy, apex acuminate. Flowers resupinate, slightly fragrant, white; ovary and pedicel twisted, fusiform, ca. 14 mm, pubescent. Sepals green tinged reddish brown at apex and pubescent on outer surface; dorsal sepal ovate-elliptic, 910 ca. 6 mm, apex acuminate; lateral sepals elliptic, oblique, 1112 4.55 mm, apex subacute. Pet-
9. Anoectochilus chapaensis Gagnepain, Bull. Mus. Natl. Hist. Nat., sr. 2, 3: 679. 1931. dian yue jin xian lan Plants 1218 cm tall. Stem ascending or erect, 2.53 mm in diam., 4- or 5-leaved. Leaves abaxially pale green, adaxially blackish green with golden-red venation, ovate, oblique, 25 1.53.5 cm, apex acute; petiole-like base and tubular sheath 0.81.6 cm. Peduncle 811 cm, with 2 or 3 pale red sheathing sterile bracts; rachis 25 cm, 27-flowered; floral bracts pale red, ovate-lanceolate, 610 mm, shorter than ovary, outer sur-
face pubescent, apex acuminate. Flowers erect, not resupinate, white; ovary and pedicel narrowly cylindric, not twisted, 1013 mm, pubescent. Sepals pubescent on outer surface, 1-veined; dorsal sepal ovate, cymbiform, ca. 6 3.5 mm, apex acuminate; lateral sepals ovate-oblong, ca. 6 3 mm, apex obtuse. Petals ovate, strongly oblique, falcate, ca. 6 3 mm, 1-veined, apex caudate and incurved; lip white, Y-shaped, 1012 mm; hypochile short; mesochile 56 ca. 3 mm; flanges oblong, pectinate, divided into 3 or 4 narrow teeth; epichile longitudinally dilated, 2-lobed; lobes diverging widely, obovate, 89 67 mm, margin crenulate, apex truncate; spur conic, 45 mm, apex obtuse, containing 2 calli. Column short, with 2 broad lamellate wings. Fl. JulAug.
Shaded humid places in dense forests; 13001400 m. SE Yunnan [N Vietnam].
10. Anoectochilus zhejiangensis Z. Wei & Y. B. Chang, Bull. Bot. Res., Harbin 9(2): 39. 1989. zhe jiang jin xian lan Plants 816 cm tall. Stem ascending, pale reddish brown, 26-leaved. Leaves abaxially pale purplish red, adaxially dark greenish purple with golden-red venation, broadly ovate to ovate-orbicular, 0.72.6 0.62.1 cm, margin slightly undulate, apex acute; petiole-like base and tubular sheath ca. 0.8 cm. Peduncle 510 cm, with 2 sterile bracts; rachis 14 cm, 14flowered; floral bracts ovate-lanceolate, ca. 6.5 mm, subequal in length to ovary, outer surface pubescent, apex acuminate. Flowers erect, not resupinate; ovary and pedicel pale reddish brown, narrowly fusiform, not twisted, ca. 7 mm, white pubescent. Sepals pinkish red, outer surface pubescent; dorsal sepal ovate, cymbiform, ca. 5 3.5 mm, apex acute; lateral sepals spreading, oblong-obovate, slightly oblique, ca. 7 3 mm, apex obtuse. Petals white, oblanceolate, strongly oblique, ca. 6 2 mm, apex acute; lip white, Y-shaped, ca. 10 mm; hypochile ca. 2.5 mm; mesochile ca. 3 mm; flanges pectinate, composed of 25 narrow teeth, each tooth ca. 2 mm; epichile longitudinally dilated, 2-lobed; lobes diverging at an acute angle to one another, flabellate, slightly oblique, ca. 6 5 mm, entire, apex truncate; spur conic, slightly curved, ca. 5 mm, apex shallowly bilobed, containing 2 irregular, stalked calli near entrance. Column short. Fl. JulSep.
Damp places in dense forests, valleys; 7001200 m. C and N Fujian, N Guangxi, SW Zhejiang.
3-leaved. Leaves abaxially purplish red, adaxially blackish green with golden-red venation, ovate, 3.54 33.2 cm, apex acute; petiole-like base and tubular sheath 12 cm. Peduncle pale purplish red, ca. 12 cm, with 24 pale purplish red sterile bracts; rachis 22.5 cm, 3- or 4-flowered; floral bracts pale purplish red, lanceolate, ca. 6 mm, shorter than ovary, outer surface pubescent, apex acuminate. Flowers erect, not resupinate, white, foetid; ovary and pedicel cylindric-fusiform, not twisted, ca. 13 mm, pubescent. Sepals purplish red, outer surface sparsely pubescent, 1-veined; dorsal sepal ovate-elliptic, cymbiform, ca. 7 2 mm, apex acute; lateral sepals spreading, narrowly oblong-obovate, oblique, 78 ca. 3.2 mm, apex acute to shortly mucronate. Petals pinkish white, ovate, strongly oblique, falcate, ca. 7 3 mm, 1-veined, apex acute; lip white, Y-shaped, ca. 13 mm; hypochile ca. 3 mm; mesochile reflexed sharply downward from apex of hypochile, ca. 4 mm; flanges subsquare, 33.5 33.5 mm, margin fimbriate, divided into 5 teeth, each tooth ca. 1.5 mm; epichile longitudinally dilated, 2lobed; lobes diverging at an acute angle to one another, oblong to oblanceolate, ca. 5 2 mm, entire, apex obtuse; spur conic, ca. 6 mm, apex shallowly bilobed, containing 2 subcuneate calli. Column ca. 5.5 mm, stout, with 2 broad, oblong wings; stigma lobes elliptic. Fl. AugSep.
Forests along streams; ca. 900 m. SC Sichuan.
11. Anoectochilus emeiensis K. Y. Lang, Acta Phytotax. Sin. 20: 183. 1982. e mei jin xian lan Plants 1921 cm tall. Stem ascending, ca. 2 mm in diam.,