Catalogo de Decapodos

Carideorum Catalogus:
The Recent Species of the Dendrobranchiate, Stenopodidean,

Procarididean and Caridean Shrimps (Crustacea: Decapoda)
S. De Grave & C.H.J.M. Fransen*
De Grave, S. & C.H.J.M. Fransen. Carideorum catalogus: the recent species of the dendrobranchiate,
stenopodidean, procarididean and caridean shrimps (Crustacea: Decapoda).
Zool. Med. Leiden 85(9): 30.ix.2011: 195-589 fgs 1-59. ISSN 0024-0672, ISBN 978-90-6519-200-4.
S. De Grave, Oxford University Museum of Natural History, Parks Road, Oxford, OX1 3PW, United
Kingdom (sammy.degrave@oum.ox.ac.uk).
C.H.J.M. Fransen, Department of Marine Zoology, Netherlands Centre for Biodiversity Naturalis, P.O.
Box 9517, 2300 RA Leiden, The Netherlands (charles.fransen@ncbnaturalis.nl).
* The order of authorship is purely alphabetical, with both authors having equally contributed to this
work.
Key words: Crustacea; Decapoda; Dendrobranchiata; Stenopodidea; Procarididea; Caridea; checklist.
A checklist of recent species of dendrobranchiate, stenopodidean, procarididean and caridean
shrimps including synonyms and type localities. Also listed are unavailable names, larval names,
nomina dubia and nomina nuda. A complete list of references to original descriptions of taxa listed is
provided.
Contents
Introduction ............................................................................................................................................................ 196
The higher classifcation of shrimp .................................................................................................. 196
Fossil taxa ........................................................................................................................................................ 197
How many shrimp species are there? ............................................................................................ 197
Structure of the list ............................................................................................................................................. 197
Notes on type locality ....................................................................................................................................... 199
Notes on dating of some papers ................................................................................................................ 200
Notes on author names .................................................................................................................................... 201
Notes on authorship of taxa ......................................................................................................................... 202
Notes on some papers of nomenclatorial signifcance ................................................................. 203
Notes on the spelling of some names ..................................................................................................... 204
Notes on the gender of generic names and gender agreement in species names ........ 204
Acknowledgements ........................................................................................................................................... 205
Checklist ................................................................................................................................................................... 206
Suborder Dendrobranchiata ................................................................................................................ 206
Suborder Pleocyemata ............................................................................................................................. 248
Infraorder Procarididea .......................................................................................................................... 248
Infraorder Stenopodidea ........................................................................................................................ 249
Infraorder Caridea ..................................................................................................................................... 254
Unavailable names ..................................................................................................................................... 471
Penaeoidea Larvata .................................................................................................................................... 471
Penaeoidea Nomina dubia .................................................................................................................... 471
196 De Grave & Fransen. Carideorum catalogus (Crustacea: Decapoda). Zool. Med. Leiden 85 (2011)
Penaeoidea Nomina nuda ..................................................................................................................... 472
Sergestoidea Larvata ................................................................................................................................. 472
Sergestoidea Nomina dubia ................................................................................................................. 472
Sergestoidea Nomina nuda .................................................................................................................. 473
Caridea Larvata ............................................................................................................................................ 473
Caridea Nomina dubia ............................................................................................................................ 475
Caridea Nomina nuda ............................................................................................................................. 476
References ................................................................................................................................................................ 478
Systematic index to currently valid families and genera ............................................................ 585
Introduction
The higher classifcation of shrimp
Over the last decade or so, much has been writen on the classifcation of Decapoda,
fuelled by a surge in molecular phylogenetic studies, as well as close scrutiny of inter-
nal and external morphological characteristics. As discussed by Fransen & De Grave
(2009), such studies on shrimps are still somewhat thin on the ground, at least com-
pared to the more extensive work done on the Brachyura and Anomura. At a higher
level in decapod classifcation it has long been recognised that three distinct lineages of
shrimps can be distinguished: Dendrobranchiata, Stenopodidea and Caridea, a system
which has not been seriously challenged by recent studies.
The internal classifcation of Dendrobranchiata and Stenopodidea alike has been
stable for some time, with the only major addition being the family Macromaxilloca-
rididae Alvarez, Ilife & Villalobos (2006) to the Stenopodidea in recent years.
A diferent picture has emerged for Caridea very recently with Bracken et al. (2009)
and Chan et al. (2010), both drawing atention to the non-monophyletic status of certain
superfamilies and families. Further, we are aware of work currently in progress (some
by the authors of this compilation) corroborating the hypothesis that the current clas-
sifcation of Caridea is unnatural, lines of study which will lead to the resurrection of
certain family names as well as further refnement to other families. As one of our objec-
tives for the current efort was to link this compilation of species level information
with the earlier work by Chace (1992) for families and Holthuis (1993a) for genera, we
have elected to largely follow the classifcation outlined by De Grave et al. (2009) which
builds upon this earlier work. As such, it was deemed advisable to include the recently
resurrected family Acanthephyridae Spence Bate, 1888 in the superfamily Oplopho-
roidea, rather than in this catalogue to create a new superfamily, which would perhaps
be more congruent with the results in Chan et al. (2010).
Although we follow herein the classifcation scheme of De Grave et al. (2009), two
recent changes have been implemented. The clarifcation of the status of Galatheacaris
abyssalis Vereshchaka, 1997a, as the megalopal stage of Eugonatonotus chacei Chan & Yu,
1991a, by De Grave et al. (2010) resulted in the removal of the family Galatheacarididae
and superfamily Galatheacaridoidea in the current listing. Bracken et al. (2010) clarifed
the status of the family Procarididae, resulting in the recognition of a fourth group of
shrimp, Infraorder Procarididea.
De Grave & Fransen. Carideorum catalogus (Crustacea: Decapoda). Zool. Med. Leiden 85 (2011) 197
Fossil taxa
The current listing is restricted to extant genera, species and subspecies only. For
readers interested in how the current classifcation of extant taxa compares with those
of fossil taxa, we refer to De Grave et al. (2009) who employed a suprageneric classifca-
tion of Decapoda across both fossil and extant taxa, as well as Schweitzer et al. (2010)
who list all species (and their classifcation) of fossil species. Schweitzer et al. (2010) list
97 fossil species of Dendrobranchiata in 31 genera, two fossil species in Stenopodidae
(each in its own genus) and 52 species of fossil Caridea in 33 genera. Their placement of
the genus Udora Mnster, 1839, within the superfamily Procaridoidea (herein consid-
ered at infraordinal level) is interesting, as this group is currently only known from six
species, restricted to anchialine habitats. If the fve known species of Udora are indeed
correctly placed, the fossil record of the groups extends back to the Middle Jurassic, an
interesting observation as the frst extant species was only discovered in 1971, drawing
parallels with the discovery of coelacanths, one of the beter known living fossils.
How many shrimp species are there?
Fransen & De Grave (2009) working from a preliminary version of the present
catalogue estimated the species/subspecies richness of shrimps as follows: Dendro-
branchiata (505), Stenopodidea (58) and Caridea (circa 3108), with those fgures slightly
adjusted in De Grave et al. (2009). Since then numerous descriptions have appeared,
synonymies corrected and species placed in synonymy. The current tally stands as fol-
lows: Dendrobranchiata (68 genera, 533 species), Procarididea (2 genera, 6 species),
Stenopodidea (12 genera, 71 species) and Caridea (389 genera, 3438 species). Although
the fgures listed in De Grave et al. (2009) for the other decapod taxa have clearly been
adjusted by now, the Caridea remain the second most species-rich group within the
Decapoda, with approximately half as many species as Brachyura, the dominant taxon
within Decapoda.
On a more inclusive level, the Caridea are dominated by Palaemonidae (981 spe-
cies), followed by Alpheidae (663), Atyidae (469), Hippolytidae (338) and Crangonidae
(219). Under the traditional classifcation employed here, Palaemonidae comprises two
subfamilies, with Pontoniinae being considerably richer in species (602) than Palae-
moninae (379). In contrast to those species-rich families, are those 12 families which are
monogeneric, some only consisting of a single species, e.g. Physetocarididae.
On a generic level, the most speciose genera are Caridina (Atyidae, 290 species), Al-
pheus (Alpheidae, 286), Macrobrachium (Palaemoninae, 243), Synalpheus (Alpheidae, 159)
and Periclimenes (Pontoniinae, 152). As doubt has been cast on the monophyletic status
of these mega-genera in recent years, this ranking may substantially alter when phylo-
genetic studies progress.
Structure of the list
As discussed above, we recognise herein four major groups of shrimp: Infraorder
Dendrobranchiata and suborders Procarididea, Stenopodidea and Caridea. For each
of those higher categories we list the currently recognized superfamilies and families,
following De Grave et al. (2009) with the changes noted above. As the focus of the cur-
rent compilation is on genera and species, for the sake of brevity we have not provided
the historical information for superfamilies and families, nor have we listed their syno-
nyms. These can be found in Holthuis (1993a) for Stenopodidae and Caridea (including
Procarididae) and Prez Farfante & Kensley (1997) for Dendrobranchiata. For each ge-
nus, we provide the reference to the original description, as well as on type species (and
the way in which they were designated) as well as their gender. In addition, we list all
their known synonyms. For species and subspecies, we list their original generic desig-
nation, in original orthography for both genus and species, as well as all known syno-
nyms. For each, we provide the page number on which the species description starts, as
well as a reference to all fgures and/or plates on which the species is illustrated, as well
as their type locality (see below). Lectotype and neotype designations are included, as
well as additional comments when required. Additional comments concern dating is-
sues, as well as fgures and/or further descriptions by the same author (ofen again as
sp. nov.) in a later work. For instance, Parapasipha gilesii was illustrated in Wood-Mason
(1892), thus validating the name, but without any accompanying textual description
which appeared later in Wood-Mason & Alcock (1893). Such cases have been annotated
to provide clarity for future researchers.
All species names are given in their original orthography. This also includes usage of
the Latin ligatures, and when present in a species name, such as in Palmon longi-
cornis Olivier, 1811, or dipus gramineus Dana, 1852a. Although we are aware that the
ICZN Code does not diferentiate this orthography, we felt this to be more appropriate in
view of the long history of taxonomy. For instance we list Pelias Niloticus Roux, 1833 (now
Caridina nilotica) and Pal.[mon] Swainsonii White, 1847a (a synonym of Macrobrachium
acanthurus), as writen in their original descriptions. The square brackets in the later case
signifying an addition on our part, to make the generic designation clearer.
As a general rule the latest taxonomic or systematic treatment for each species were
followed, listing species as valid or as a synonym accordingly. This does not take into
account the uncertainties regarding the true status of some taxa, which in reality may
be valid species instead of a synonym and vice versa, but does refect our current un-
derstanding of species richness. One notable exception is that the sergestid genera
erected by Judkins & Kensley (2008) were not recognised in a more recent revision by
Vereshchaka (2009), who instead recognised several species groups in Sergestes sensu
lato. As these species groups largely correspond to the new genera proposed by Judkins
& Kensley (2008), we have elected to continue to treat them as genera, necessitating
some generic rearrangements to new species proposed in Vereshchaka (2009).
Nomina dubia and nomina nuda have been listed as synonyms under currently
valid names if such information was available, if not they have been listed as a separate
category. We were aided herein by the Holthuis card catalogue, who listed many such
names even under appropriate family headings. As Holthuiss logic was not always
transparent and other names surfaced or were assigned diferently by other workers,
we have not followed his family level assignations, but have retained them under three
categories: Penaeoidea, Sergestoidea and Caridea.
The scientifc literature abounds with misspellings of species names due to printing
and typographical errors. Listing all of these would enlarge the current volume many-
fold. Thus a minor error, such as Palaemon pandaliformes instead of Palaemon pandali-
formis (Stimpson, 1871) in Tavares (1993) we do not list. However, if the magnitude of
the error is such that it could possibly be interpreted as a diferent name we have in-
cluded those, to avoid further confusion. For instance, Burukovsky (1974) used Artis-
teus viritli for Aristeus virilis Spence Bate, 1881, or Acanthephyra parvirostris in Coutire
(1911a), an erroneous spelling of A. brevirostris Spence Bate, 1888.
During the early years of ocean exploration, numerous larval forms were described,
for instance by Spence Bate (1888) and Ortmann (1893). Although on occasion these
have been linked to adult forms, as is the case for Embryocaris stylicauda Ortmann, 1893,
which is a larval stage of Stenopus hispidus (Olivier, 1811), many remain unassigned to
adult forms at present. Those which have not been linked are herein listed under the
headings of Penaeoidea, Sergestoidea and Caridea Larvata.
The present catalogue includes the binomial names for shrimps from Linnaeus
(1758) onwards, up to 1st June 2011. We have not considered pre-Linnean names, even
though sometimes these have been mentioned in later literature, for instance in the
works by Holthuis (1991), Tavares (1993) and others. The cut-of date for inclusion of
names was set at 1st June 2011, although we are acutely aware that a number of names
are currently in press and some of these will have appeared in the scientifc literature
before the present work sees the light of day.
Notes on type locality
For all species we have listed their type locality in one of two ways. On the whole
we have done this by reference to the original type series, but restricted to holotype
locality if a holotype is indicated, as is the norm in recent papers. For instance, for Peri-
climenes cannaphilus Komai & J.N. Kim, 2010, we only list the locality of Kasuga 2 Sea-
mount where the holotype originated from and not the locality of the 15 paratypes from
Nikko and Daikoku Seamounts. If no holotype was indicated in the original publica-
tion, then we have included all syntypic localities in the listed type locality, resulting in
upwards of 10 stations being listed for some species from the early expeditions (e.g.
Spence Bate, 1888). If a later nomenclatorial act selected a lectotype from the syntypic
series, we have only listed the locality details for that particular specimen. If a neotype
was selected, then the information for the neotype only features, and not the original
type series. For example, Cai et al. (2006) selected neotypes for the six Japanese species
of Atyidae described by Stimpson (1860a), the original type series being lost in the great
Chicago fre of 1871. Thus the type locality information is only given for the neotypes,
and the reader is referred to the discussion in Cai et al. (2006) or the original publication
by Stimpson (1860a) for locality information for the original (now lost) syntypes. For
the majority of taxa we have elected to adhere to the original language of the descrip-
tion in our listing of type locality, as well as the original orthography and names of lo-
calities. For example, Stimpson (1860a) describes Alpheus pachychirus and other species
from ad insulam Loo Choo, currently known as the Ryukyu Islands in southern Ja-
pan. We accept the potential criticism that this may make this part of the current listing
somewhat difcult to work with for non-taxonomists, but in our opinion it accurately
refects the rich tapestry of species descriptions, and correctly emphasizes the historical
heritage and multinational aspect of alpha-level taxonomy. By necessity, we had to ac-
cept our own linguistic limitations and have had (by other taxonomists who are more
profcient in such maters) localities originally listed in Russian, Chinese and Vietnam-
ese translated. These are simply listed in English in the present compilation.
In many earlier publications no type localities are listed, bar a reference to a certain
expedition station or a vague collection locality but with a specifc station included. For
instance, Rathbun (1902a) described 50 taxa from the coast of Alaska southwards to San
Diego, based on samples which had accumulated for many years in the Smithsonian as
a result of the Albatross expeditions. The majority of these have vague localities for the
syntypic series in the actual publication, but do include a specifc reference to one or
more Albatross stations. By reference to various published lists, we have added such
information between brackets in the herein listed type locality information. These may
need to be corrected once the material and original labels are re-examined as part of
future revisions. Several sources were consulted for this, notably Anon. (1914) for Inves-
tigator stations, Murray & Hjort (1912) for the Michael Sars expedition, as well as Sewell
(1935) for species in Calman (1939) and others. For some taxa, such information could
not be directly obtained from published station lists or the original description, but
later authors have tracked down such information. For example, Crosnier & Forest
(1973) provided accurate locality information for Gennadas talismani Bouvier, 1906a, and
Metapenaeus perlarum Nobili, 1905a. In such instances, we have always included a refer-
ence as to where the information came from. On a few occasions, the actual type de-
scription nor any later paper discussed the exact type locality details. For instance
Macpherson (1984) when describing Nematocarcinus gladius listed three stations with-
out further details with the material being deposited in the South African Museum
(now IZIKO). In those instances we made direct enquiries with the museums involved
and provide here the information on the specimen labels and their catalogues.
Notes on dating of some papers
A very important aspect of nomenclature is the accurate dating of nomenclatorial
acts, something which is not always understood by colleagues in other biological disci-
plines. For an extreme example of this issue we refer to Low & Guinot (2010) who dis-
cuss a case of 48 hour precedence. Dating issues have been ofen reported for older
references and are most prominent in early works, such as Gurin-Mneville (1829-
1838) and Leach (1815-1875) to name but two. In those cases where dating issues have
been reported in the literature, discrepancies were noted in various draf versions of the
current catalogue or have been brought to our atention by our colleagues, we have
endeavoured to decide upon the correct date of publication. For this we have relied on
a variety of sources, such as Rathbun (1897) for Herbst (1791-1796), Gordon (1959) for
Bell (1844-1853), dErasmo (1949) for O.G. Costa & A. Costa (1838-1871), Holthuis (1947a)
for Sowerby (1804-1806) and Holthuis (1961a) for Gurin-Mneville (1829-1838). In ad-
dition, we followed Harrison & Smith (2008) for dating of all of Leachs publications.
For dating of individual issues of certain journals with a known history of confusion,
we followed Duncan (1937) and Dickinson (2005) for Proceedings of the Zoological Society
in its numerous guises as well as their Transactions, Evenhuis (2003) for the Annals and
Magazine of Natural History and others.
Contrary to popular belief, such dating issues are not just restricted to the earlier
taxonomic works and are almost as ubiquitous in more recent decades, although usually
much easier to solve. For example, the description of Plesionika williamsi Forest, 1964 is
in the volume of the Bulletin du Musum national dHistoire naturelle (2me srie) for 1963,
but this particular issue was printed on the 19th June 1964 (R. Clva, pers. comm.). A
peculiar case are the two pontoniine genera Exoclimenella Bruce, 1995, and Periclime-
nella Bruce, 1995. As already pointed out by Holthuis (1996) the descriptions of these
two genera in Bruce (1995) is valid under the current ICZN Code and thus predates the
descriptions (as gen. nov.) in uri & Bruce (1995). However, this assessment was
based on a publication date of 30th November 1994, as indicated on the frontispiece.
Per Koeltz (Koeltz Scientifc Books) has confrmed that the actual publication date of
Bruce (1995) was 16th March 1995, which curiously does not afect the seniority of
those descriptions as the uri & Bruce (1995) article appeared in the May/June issue
of Journal of Natural History, which according to Evenhuis (2003) was published on 31st
May 1995.
In those cases where we identifed a discrepancy between the date usually associ-
ated with a publication and the real publication date determining priority, we have
annotated the references following the conventions used by Schweitzer et al. (2010).
Some of the early descriptions appear in multi-year works, such as Gurin-Mneville
(1829-1838) and H. Milne Edwards (1836-1844). For these, we follow the quotation style
in De Grave et al. (2009). For example, the original description of Xiphocaris elongata
appears in Gurin-Mneville (1855-1856), as Hippolyte elongatus with 1855 as the pub-
lication date of the actual nomenclatorial act. This is thus listed as Hippolyte elongatus
Gurin-Mneville, 1855 [in Gurin-Mneville, 1855-1856] to distinguish the nomen-
clatorial act from the bibliographical reference.
Notes on author names
The numerous works by Charles Spence Bate (1819-1889) have usually been cited in
modern literature under the name of Bate, C.S., although invariably Charles himself
used the name Spence Bate, C.. It is not clear when Charles started using Spence
Bate as a surname, nor the exact reasons why, nor indeed if he just adopted this name
in publications and continued to use Bate in his private life. According to the Oxford
National Dictionary of Biography his mother was Harriet Spence (1788-1879) and we
postulate here that he added Spence to his surname to distinguish himself from his fa-
ther Charles Bate (1789-1872) who was also a dentist in Cornwall. We have checked all
his carcinological papers, as well as most of his dentistry ones and fnd that indeed he
invariably signed of in all his papers as C. Spence Bate. This is also the surname
by which his contemporaries refer to him in their own publications (e.g. Semper, 1868;
Miers, 1884a; A. Milne-Edwards, 1891, etc.), although sometimes abbreviated to Sp.
Bate. In the two editorial notes by John Murray in Spence Bate, 1888 it is stated: il-
lustrate the text of Mr. Spence Bates report as well as: The Report on the Crustacea
Macrura, by C. Spence Bate, Esq.. Within the text of Spence Bate (1888) itself the
author always (with one exception) refers to his own, older work as Spence Bate, of-
ten shortened in the synonymy listings as Sp. B. The exception being when he refers
to Bate & Westwood (1863, A History of British Sessile-eyed Crustacea). We interpret that
here as a lapsus, as the title page of that work also identifes the authors as C. Spence
Bate and J.O. Westwood.
The present case mirrors the consistent usage of H. Milne Edwards in publications
by that author, whose surname was Edwards, but who added Milne to distinguish him-
self from his brother William Edwards (see Forest, 1996). As we can see no reason why
the name Charles Spence Bate should be treated diferent to Henri Milne Edwards, we
here accept the arguments put forward by Dubois (2008) in that authors are signatures
and not persons, and thus have elected to use the name C. Spence Bate.
For the father and son team of Henri Milne Edwards and Alphonse Milne-Edwards,
we herein follow Forest (1996) in the spelling of their surnames, and have added, as is
customary in carcinology, their initial to all citations. Issues surrounding the use of
Gurin and Gurin-Mneville were discussed in Ng et al. (2008) and we follow here
their lead in atributing names afer 1836 onwards to Gurin-Mneville and before 1836
to Gurin.
Ng et al. (2008) also provide a more in depth discussion on the spelling of author
names and how these should be cited in the references. As we herein adhere to the same
rules, we refer the reader to that treatise.
In those instances where diferent taxa have been described by diferent authors
who share the same surname, we generally have adopted the same rules as outlined in
Ng et al. (2008). In brief, we do not add the authors initials to the person having de-
scribed signifcantly more taxa, but add those to the author having described fewer.
For instance, for taxa described by Peter K.L. Ng we do not add P.K.L. in front of the
surname, whilst we do so for Ngan Kee Ng (thus N.K. Ng). As in Ng et al. (2008) we
do not include initials if the authors were not contemporaries, the intervening time lag
making this sufciently obvious. For instance, J.Y. Johnson (1863, 1868) and D.S. John-
son (1961, 1962, 1967, 1973). A special case is the citation of frst names for workers of
Chinese descent or nationality, where fashion has dictated how names are used rather
than bibliographical accuracy. For instance, the Chinese freshwater carcinologist Guo
Zhao Liang has published under the names Guo Zhaoliang (abbreviated to Guo, Z.),
Guo Zhao-Liang (Guo, Z.-L.) and Guo Zhao Liang (Guo, Z.L.). As it is evident that in
all three cases this is the same worker, we have elected not to include his initials in the
citations, others, more purists perhaps, may disagree. In the case of the team of A.H.
Hank and D.M. Dora Banner, who described numerous alpheid taxa, ofen jointly,
they stated several times in their larger publications (e.g. Banner & Banner, 1978; Ban-
ner & Banner, 1981a) that they list their own papers in strict chronological order, with-
out reference to seniority of authorship. We have herein interpreted this as a desire to
refect a joint efort with order of authorship, perhaps somewhat arbitrarily decided
upon and a refection of equal share in the description. As such we have elected to not
add initials to citations in the checklist, although order of authorship in the references
is as given in the actual papers to facilitate bibliographical searches. All this makes for
less cumbersome citations, however in other cases, we had no option but to include
initials.
Notes on authorship of taxa
As extensively discussed by Ng et al. (2008) atributing the correct author to a cer-
tain taxon can be confusing, particularly (but not exclusively) in the earlier literature.
Ofen in the earlier literature another carcinologists name is cited behind the species
name in the work of a diferent author. Such reference is usually to an unpublished
manuscript, a personal leter or even a museum label and many examples abound in,
for instance, H. Milne Edwards (1834-1840). An interesting example is the name Lys-
mata nilita as published in Hope (1851) as a nomen nudum, with Risso MS featuring
behind the name. Clearly Hope must have had access to unpublished manuscripts by
Risso, as is evident from Monod (1931). Fitingly, when Dohrn & Holthuis (1950) de-
scribed the second species of Lysmata from the Mediterranean, they adopted the name
Lysmata nilita.
As in Ng et al. (2008) we herein strictly adhere to Article 50 of the ICZN and atribute
authorship only to those who are directly responsible for the name and for satisfying the
criteria of availability. For example, we thus list Caridina cavalerieioides Liu & Liang in Li-
ang (2004), but in contrast in the case of Stenopus devaneyi we use Goy (1984) rather than
Goy & Randall (1984), as was used in Goy & Randall (1986). Observant readers will thus
notice a discrepancy between the current list and Holthuis (1993a) for certain names at-
tributed to Wood-Mason & Alcock in the The Annals and Magazine of Natural History. This
series of papers was discussed in Ng (1998) and we herein follow that interpretation.
Notes on some papers of nomenclatorial signifcance
Confusion exists in the literature regarding the type description of the rare alpheid
Betaeus jousseaumei (now Amphibetaeus jousseaumei), which Holthuis (1993a) atributed
to Coutire (1897a) but therein dated as 1896. Coutire himself in his magnifcent opus
on Alpheidae (Coutire, 1899a) refers instead to Coutire (1896). We have not been able
to accurately date either of those publications and herein follow Coutire (1899a) in
considering Coutire (1896) as the type description, as well as following him in consid-
ering the publication year of Coutire (1897a) as 1897 and not 1896 as indicated in
Holthuis (1993a), as indeed do Banner & Banner (1981a) and Anker & Jeng (2006).
Gurin-Mneville authored the Crustacea section in Ramon de la Sagras work
Histoire Physique, Politique et Naturelle de lle de Cuba, which also almost contempora-
neously appeared in a Spanish language version Historia fsica politica y natural de la Isla
de Cuba. Although nothing is known about the actual publication dates for the Crusta-
cea section, it is known that for the Mollusca the French text is the original version,
which appeared in livraisons and was aferwards translated into Spanish (G. Rosen-
berg, pers. comm.). Curiously for Caridea, the French edition has several more shrimp
names in the text, which do not feature in the Spanish version. Perhaps this can be ex-
plained by the fact that these additional species did not originate from Cuba, and thus
perhaps were deleted from a Spanish translation, targeted at a more local audience. It
is also known that the dates appearing on the frontispiece of the Mollusca part are in-
correct. For the French edition it is the date the text was concluded, even though the
work appeared in livraisons over several years, whilst for the Spanish version it is an
intermediate date (G. Rosenberg, pers. comm.). In the absence of any frm evidence re-
garding the exact publication date of the crustacean parts, both text and Plates, we have
no other course of action available than to follow the dates on the frontispiece, even
though future research will almost certainly prove these to be wrong, this task is now
taken up by P. Clark. Thus, for the species mentioned in both the French and Spanish
editions, for instance Atya Poeyi Gurin-Mneville, 1855 [in Gurin-Mneville, 1855-1856]
we employ 1855 as their description date, corresponding to the date on the frontispiece
of Tomo VIII (Atlas de Zoologia), with the corresponding text in Tomo VII (Crustceos,
Aragnides Insectos) with 1856 on the frontispiece. For the additional taxa listed in the
French edition, for example Alpheus Rouxii Gurin-Mneville, 1857, we use 1857 as the
date, corresponding to what is writen on the frontispiece of that volume.
Notes on the spelling of some names
In general, we have listed the species in their original spelling, unless a justifed
emendation needed to be made, for example to comply with Art. 31.2 in relation to
gender agreement between generic and species names.
When revising the genus Pandalus, Komai (1999) proposed that the species usu-
ally quoted as Pandalus propinquus in the majority of the taxonomic and ecological
literature, should revert to its original spelling in the new combination of Atlantopan-
dalus propinqvus, as used (Pandalus propinqvus) in the type description by G.O. Sars
(1870). Under Art. 32.1 we herein follow Komai (1999) and continue to list the species
as A. propinqvus.
A similar case exists for the commercially important species, Plesionika narval
(Fabricius, 1787), which was originally described by Fabricius (1787) under the name
Astacus narual. Under Art. 33.3.1 we herein maintain the species as P. narval, and do
not revert back to the original spelling.
The species Lebbeus microceros (Kryer, 1841) has been variable listed as L. microceros
and L. microceras in older literature. Kryer (1841) originally described the species un-
der the name Hippolyte microceras (Kryer, 1841, p. 578), but a note has been inserted in
the journal afer the table of content, in which the name is corrected to Hippolyte micro-
ceros. Under Art. 32.5.1.1 the original spelling thus has to be treated as an inadvertent
error and corrected to L. microceros.
Acanthephyra eximia Smith, 1884 has ofen been listed as A. eximea Smith, 1884. Both
names were used by Smith (1884). Smith himself used A. eximia in his publication of
1886 (pp. 189, 190, 192). In between no other author had used both names in an article
and selected a correct spelling as First Reviser, thus here Art. 24.2.4 holds and Smith
himself becomes the First Reviser.
Notes on the gender of generic names and gender agreement in species names
As already discussed by Ng et al. (2009) there has been confusion about the gender
of generic names, as well as the correct ending of species names, if adjectival in nature.
If the original genus description stated gender, we have simply followed this. Care
needs to be taken when assuming gender though. For instance, Eumanningia Crosnier,
2000, may sound feminine, but as clearly stated by Crosnier (2000), the name has to be
treated as masculine. In the majority of older descriptions the gender was not stated, in
all those cases we follow Holthuis (1983) and Prez Farfante & Kensley (1997) for the
gender of the genus, with one exception. The gender of the genus Petalidium Spence
Bate, 1881, was treated as masculine by Prez Farfante & Kensley (1997). We herein treat
it as neuter, in recognition of the name of the type species, Petalidium foliaceum, as well
as the other species assigned to this genus.
Confusion exists about the correct ending of some species names, as it is not always
clear from the type description if a species name has to be treated as a noun or an adjec-
tive, especially in older descriptions when derivations were not commonly included.
For some, the adjectival nature of the name is abundantly clear, and the necessary
changes have been introduced in the present listing. For others, our own limited knowl-
edge of the classical languages has prevented us from forming a defnite conclusion
on this mater. In those few cases, we have been fortunate that Lipke Holthuis, a mas-
ter of classical names, used these names before and we have simply followed suit. For
instance, even though the genus Halocaridinides is masculine, Holthuis (1982) used H.
trigonophthalma, efectively treating it as a noun, instead of the masculine form trigo-
nophthalmus.
Acknowledgements
Over the long gestation period of this catalogue, numerous people provided assis-
tance with tracking down references, translated relevant papers, advised on nomencla-
torial issues, spelling of author names and dating issues, provided us with copies of
their own work and checked our listing in their special families. We are immensely
grateful to them all. We list them here in alphabetical order: A. Anker, C.W. Ashelby,
A.J. Bruce, R.N. Burukovsky, Y. Cai, T.-Y. Chan, P. Clark, P.C. Dworschak, C. dUdekem
dAcoz, J.W. Goy, L.Y.D. Goulding, R. Gulledge, K.-I. Hayashi, M.E. Hendrickx, S.
Ivanenko, W. Klotz, T. Komai, R. Lemaitre, X. Li, M.E.Y. Low, D. Mann, I. Marin, T.
Naruse, P.K.L. Ng, K. Pocklington, Nguyen Thanh Son, K. Reed, J. Poupin, S.H. Tan,
A.L. Vereshchaka, M.K. Wicksten and K. Wouters, whilst ofering our apologies to
those who assisted along the way and are not listed. We are also grateful to the librari-
ans of the Koninklik Belgisch Instituut voor Natuurwetenschappen (KBIN-IRSNB),
Netherlands Centre for Biodiversity Naturalis, University College Cork, the Hope Li-
brary (OUMNH) and the Radclife Science Library, Oxford who provided references
and expedited the necessary inter-library loans. For the many photographs used we
thank A. Anker, A.J. Bruce, T.-Y. Chan, N. Ciccarese, L. Harris, O. Helker, J. Hoover, T.M.
Ilife, T. Komai, G.A. Lovrich, C. Lukhaup, K. Neely, T. Sakihara, E. del Solar, J. Tomas,
and C. dUdekem dAcoz. We are especially grateful to A. Anker, C. dUdekem dAcoz,
P.K.L. Ng and P.C. Dworschak, who went through a near complete, fnal version of the
catalogue and corrected numerous errors. Any remaining errors are the authors re-
sponsibility. Lastly, we must pay homage to L.B. Holthuis (1921-2008) who kept meticu-
lous records of species names of shrimps and erroneous spellings, many never reported
again in the scientifc literature. Without access to his fles, this catalogue would have
been far less complete. Although sadly Lipke is no longer with us and able to see the
fnished product, we are grateful for his discussions on a preliminary version, during
which he recommended the inclusion of type localities. To his memory, we dedicate this
catalogue.
Checklist
Suborder Dendrobranchiata Spence Bate, 1888
Superfamily PENAEOIDEA Rafnesque, 1815
Family ARISTEIDAE Wood-Mason in Wood-Mason & Alcock, 1891a
Aristaeomorpha Wood-Mason in Wood-Mason & Alcock, 1891a
= Aristomorpha Wood-Mason in Wood-Mason & Alcock, 1891a (type species Aristeus rostridentatus
Spence Bate, 1881, a junior subjective synonym of Penaeus foliacea Risso, 1827, by original designa-
tion, gender feminine)
Aristaeomorpha foliacea (Risso, 1827) (Fig. 1)
= Penaeus foliacea Risso, 1827: 69; Plate 2, fg. 6. [lEurope Mridionale, environs de Nice et des Alpes
Maritimes, grandes profondeurs]
= Aristeus rostridentatus Spence Bate, 1881: 189. [near the Fii Islands, 300 fms; according to Spence
Bate, 1888: Challenger stn 173, 1909'35"S 17941'50"E]
= Aristaeomorpha Giglioliana Wood-Mason, 1892: Plate 2, fg. 2. [Type locality not indicated]
= Aristaeomorpha mediterranea Adensamer, 1898: 627; unnumbered text fgure. [Pola stns 128, Nord-
kste von Afrika, 725 m; 192, Sdkuste von Kreta (3633'N 2859'E, 1242 m); 204, Meer von Kandia
(3625'N 242'E, 808 m)]
= Penaeus Meridionalis Hope, 1851: 19. [Nizza; nomen nudum]
= Aristeus japonicus Yokoya, 1933: 3; Fig. 1. [Sy-Maru stns 383, E of Owase, Mie-ken, Japan, 353 m;
399, near Owase, Mie-ken, 61 m]
Aristaeomorpha woodmasoni Calman, 1925: 8. [Of Port Blair in the Andaman Sea, 271 fms]
Aristaeopsis Wood-Mason in Wood-Mason & Alcock, 1891a
= Aristopsis Wood-Mason in Wood-Mason & Alcock, 1891a (type species Penaeus edwardsianus
Johnson, 1868, by original designation, gender feminine)
Aristaeopsis edwardsiana (Johnson, 1868)
= Penaeus edwardsianus Johnson, 1868: 897. [of Madeira, Northeast Atlantic Ocean]
= Aristeus coralinus Spence Bate, 1888: xxxii; fg. X. [Talisman expedition, exact locality not indicated]
= Aristeus splendens Richard, 1900: 89. [Talisman expedition stn 1442, Azores; Talisman expedition stn
1883, of Morocco; Talisman Expedition, of Cape Spartel, according to Davie, 2002: 119]
Aristeus Duvernoy, 1840
= Aristeus Duvernoy, 1840 (type species Penaeus antennatus Risso, 1816, by original designation,
gender masculine)
Aristeus alcocki Ramadan, 1938: 40; Figs 1, 2a, 3a. [John Murray Expedition, stns 34, Gulf of Aden, 1022
m (1305'36''N 4624'42''E); 35, Gulf of Aden, 450-550 m (1314'24''N 4614'12''E to 1313'24''N
4610'00''E); 176, Gulf of Aden, 650-730 m (1204'06''N 5038'36''E); 177, Gulf of Aden, 270-730 m
(1201'54''N 5039'12''E); 193, Gulf of Aden, 1051 m (1306'12''N 4624'30''E to 1303'00''N 4621'42''E)]
Aristeus antennatus (Risso, 1816)
= Penaeus Antennatus Risso, 1816: 96; Plate 2, fg. 6. [Nice]
= Sicyonia duvernoii Risso, 1844: 95. [Nice] [nomen nudum]
= Penaeus antemarius Costes, 1890: 558. [nomen nudum]
Aristeus antillensis A. Milne-Edwards & Bouvier, 1909: 201; Plate 1, fgs 8-13. [Nevis Island, Antilles]
Aristeus mabahissae Ramadan, 1938: 43; Figs 2b, 3b, 4a-c. [John Murray Expeditions, stns 143 (515'48''N
7322'48''E to 513'42''N 7323'36''E), vicinity of the Maldives, 795 m; 145 (448'42"S 7316'24"E), vi-
cinity of the Maldives, 510 m]
Aristeus pallidicauda Komai, 1993: 23; Figs 1-4. [Of Miyako, Japan, 600-700 m]
Aristeus semidentatus Spence Bate, 1881: 189. [south of the Philippine Islands]
? = Aristeus occidentalis Faxon, 1893: 215. [Albatross stns 3403 (of Galapagos Islands, 058'30"S
8917'00"W, 384 fms); 3410 (of Galapagos Islands, 019'00"N 9034'00"W, 331 fms)]
Aristeus varidens Holthuis, 1952a: 71; Figs 17-18. [716'S 1202'E, 53 M.W. Ambrizete, 440 m; 539'S
1125'E, 47 M.WbyS. Cabinda, 480 m; 1045'S 1310'E, 35 M.W. Cap Morro, 340 m; 1045'S 1317'E,
40 M.W. Cap Morro, 400-500 m; 1153'S 1320'E, 28 M. WbyN. Egito, 500 m; 625'S 1129'E, 50 M
WSW. Moita Seca, 430 m]
Aristeus virilis (Spence Bate, 1881)
= Hemipenaeus virilis Spence Bate, 1881: 187. [near the Philippine Islands, 255 fms; according to
Spence Bate, 1888: Challenger stn 200, near the Philippine Islands, 647'N 12228'E, 250 fms]
= Aristeus tomentosus Spence Bate, 1881: 189. [south of the Philippine Islands]
= Aristeus viritli Burukovsky, 1974: 48. [Erroneous spelling]
Austropenaeus Prez Farfante & Kensley, 1997
= Austropenaeus Prez Farfante & Kensley, 1997 (type species Plesiopenaeus nitidus Barnard, 1947, by
original designation, gender masculine)
Austropenaeus nitidus (Barnard, 1947)
= Plesiopenaeus nitidus Barnard, 1947: 383. [Of Cape Point, South Africa, 475-630 fms]
= Aristeus crosnieri Burukovsky, 1975: 779; Figs 1-4. [SE Atlantic, 2611'S 0602'E, 1150 m]
Hemipenaeus Spence Bate, 1881
= Hemipenaeus Spence Bate, 1881 (type species Hemipenaeus spinidorsalis Spence Bate, 1881, desig-
nated by Faxon, 1895, gender masculine)
Hemipenaeus carpenteri Wood-Mason & Alcock, 1891b
= Hemipenus Carpenteri Wood-Mason & Alcock, 1891b: 189. [Investigator stn 97 (Bay of Bengal,
1826'N 852'E), 1310 fms]
= Hemipenaeus triton Faxon, 1893: 215. [Albatross stns 3360 (of Panama, 617'00"N 8205'00"W, 1672
fms); 3374 (of Panama, 235'00"N 8353'00"W, 1823 fms); 3381 (456'00"N 8052'30"W, 1772 fms)]
Hemipenaeus spinidorsalis Spence Bate, 1881: 186. [South Atlantic, near the Island of Tristan dAcunha,
1900 fms; corresponding to Spence Bate, 1888: Challenger stn 133, near Tristan da Cunha, 3541'S
2055'W, 1900 fms]
Fig. 1. Aristaeomorpha foliacea (Risso, 1827). Photo by Tin-Yam Chan.
Hepomadus Spence Bate, 1881
= Hepomadus Spence Bate, 1881 (type species Hepomadus glacialis Spence Bate, 1881, designated by
Fowler, 1912, gender masculine)
Hepomadus glacialis Spence Bate, 1881: 190. [mid South Atlantic, 1875 fms; corrected in Spence Bate, 1888
to Challenger stn 237, near Yokohama, 3437'N 14032'E, 1875 fms]
Hepomadus inermis Spence Bate, 1881: 190. [middle of South Pacifc, 2550 fms; according to Spence Bate,
1888 Challenger stn 289, South Pacifc, 3941'S 13123'W, 2550 fms]
Hepomadus tener Smith, 1884: 409; Plate 9, fgs 7-8. [Albatross stn 2099, NW Atlantic, of eastern USA,
3712'20"N 6930'W, 2949 fms]
Parahepomadus Crosnier, 1978
= Parahepomadus Crosnier, 1978 (type species Parahepomadus vaubani Crosnier, 1978, by original des-
ignation and monotypy, gender masculine)
Parahepomadus vaubani Crosnier, 1978: 48; Figs 20-22. [Vauban CH 113, 2218'S 4359.7'E, 990-1010 m]
Plesiopenaeus Spence Bate, 1881
= Plesiopenaeus Spence Bate, 1881 (type species Aristeus armatus Spence Bate, 1881, designated by
Faxon, 1895, gender masculine)
Plesiopenaeus armatus (Spence Bate, 1881)
= Aristeus armatus Spence Bate, 1881: 188. [among the islands of the Australasian archipelago, in the
North Pacifc, and South Atlantic, 1900-2050 fms; according to Spence Bate, 1888: Challenger stns
133, South Atlantic Ocean, near Tristan da Cunha, 3541'S 2055'W, 1900 fms; 184, near Torres Strait,
128'S 14510'E, 1400 fms; 213, near the Philippines, 547'N 1241'E, 2050 fms; 237, of Japan, 3437'N
14032'E, 1875 fms; 246, Mid-Pacifc, 3610'N 17800'E, 2050 fms; 276, the Low Archipelago, 1328'S
14930'W, 2350 fms; 323, east of Buenos Ayres, 3539'S 5047'W, 1900 fms]
Plesiopenaeus coruscans (Wood-Mason in Wood-Mason & Alcock, 1891a)
= Aristeus coruscans Wood-Mason in Wood-Mason & Alcock, 1891a: 280; Fig. 6. [Investigator stn 112
(Bay of Bengal, 1347'30"N 9236'E), 561 fms]
Pseudaristeus Crosnier, 1978
= Pseudaristeus Crosnier, 1978 (type species Aristus crassipes Wood-Mason in Wood-Mason &
Alcock, 1891a, by original designation, gender masculine)
Pseudaristeus crassipes (Wood-Mason in Wood-Mason & Alcock, 1891a)
= Aristus crassipes Wood-Mason in Wood-Mason & Alcock, 1891a: 281; Fig. 7. [Investigator stn 116,
Andaman Sea, 1125'05"N 9247'06"E, 405 fms; lectotype designation by Prez Farfante, 1987]
Pseudaristeus gracilis (Spence Bate, 1888)
= Hemipenaeus gracilis Spence Bate, 1888: 302; Plate 44, fg. 2. [Challenger stn 207, of Tablas Island,
Philippines, 1221'N 12215'E, 700 fms]
Pseudaristeus kathleenae Prez Farfante, 1987: 314; Figs 1-3, 4C, 5-9. [Albatross stn 5657, Teluk Bone,
Sulawesi (Celebes), Indonesia, 319'40"S 12036'30"E, 900 m]
Pseudaristeus protensus Prez Farfante, 1987: 327; Figs 4D, 9, 14. [Investigator stn 370, W of Everal Gujarat,
India (Arabian Sea), 1951'30"N 6907'30"E, 1569 m]
Pseudaristeus sibogae (De Man, 1911a)
= Hemipenaeus Sibogae De Man, 1911a: 6, 25. [Siboga stn 52, Savu Sea, Indonesia, 93.4'S 11956.7'E,
1000 m]
Pseudaristeus speciosus (Spence Bate, 1881)
= Hemipenaeus speciosus Spence Bate, 1881: 186. [the Atlantic of the coast of South America, 2650
fms; according to Spence Bate, 1888: Challenger stn 325, east of Buenos Ayres, 3644'S 4616'W, 2650
fms]
Family BENTHESICYMIDAE Wood-Mason in Wood-Mason & Alcock, 1891a
Altelatipes Crosnier & Vereshchaka, 2008
= Altelatipes Crosnier & Vereshchaka, 2008 (type species Altelatipes falkenhaugae Crosnier & Veresh-
chaka, 2008, by original designation and monotypy, gender masculine)
Altelatipes brevirostris (Kikuchi & Nemoto, 1991)
= Benthesicymus brevirostris Kikuchi & Nemoto, 1991: 75; fgs 10-11. [2308.4'N 15004.8'E, 0-1300 m]
Altelatipes carinatus (Smith, 1884)
= Benthesicymus carinatus Smith, 1884: 396; Plate 10, fgs 6-7. [Albatross stn 2094, of Long Island, NW
Atlantic, 3944'30"N 714'W, 1022 fms]
= Benthesicymus expansus Kensley, 1977: 22; Figs 4-5. [Meiring Naude stn 107 (South Africa, of Natal,
2837.8'S 3238.4'E, 1200-1000 m)]
Altelatipes falkenhaugae Crosnier & Vereshchaka, 2008: 402; Figs 1-5, 6A-C, 7A-C. [ride mdio-atlan-
tique, 5304'N 3717'W, 1478-665 m]
Bentheogennema Burkenroad, 1936a
= Bentheogennema Burkenroad, 1936a (type species Gennadas intermedius Spence Bate, 1888, by origi-
nal designation, gender feminine)
Bentheogennema borealis (Rathbun, 1902a)
= Gennadas borealis Rathbun, 1902a: 887. [Albatross stn 3783, of Copper Island, Kamchatka, 1567
fms]
= Gennadas calmani Kemp, 1909: 724; Plate 74, fgs 5-11; Plate 95, fgs 4-5. [Challenger stns 232, S of
Japan, 3511'N 13928'E, 345 fms; 236, S of Japan, 347'N 138E, 565 fms; 237, near Yokohama,
3437'N 14032'E, 1875 fms]
Bentheogennema burkenroadi Krygier & Wasmer, 1975: 737; Figs 1-6. [Northeastern Pacifc, 5126'N
13828'W]
Bentheogennema intermedia (Spence Bate, 1888)
= Gennadas intermedius Spence Bate, 1888: 343; Plate 58, fg. 3. [Challenger stn 106, 147'N 2426'W,
1850 fms, of Sierra Leone; between Bermuda and Azores, surface; 137, 3559'S, 134'E, surface]
= Gennadas Alicei Bouvier, 1906a: 748. [Type locality not indicated; according to Bouvier, 1908a, the
syntypic series came from Princesse-Alice stns 1306, a 300 milles louest de S.-Miguel, 4275 m; 1715,
Canaries, 0-1000 m; 1749, entre Madre et les Canaries, 0-2500 m; 1768, Canaries, 0-3000; 1794, entre
les Canaries et les Aores, 0-3000 m; 1797, entre les Canaries et les Aores, 0-3000 m; 2016, sud-ouest
du cap Spartel, 0-1800 m; 2022, a lest du banc Seine, 0-4000 m; 2113, Mer Sargasses, 0-1500 m; 2138,
mer des Sargasses (rgion orientale), 0-2500 m; 2153, au sud des Aores, 0-2000 m; 2168, au sud de
S.-Miguel, 0-2000 m; 2168, au sud de S.-Miguel, 0-2000 m; 2212, a louest de Flores, 0-1200 m; 2244,
au sud de S.-Miguel, 0-3000 m]
Bentheogennema pasithea (De Man, 1907a)
= Gennadas Pasithea De Man, 1907a: 146. [Siboga stn 230, 358'S 12820'E, 0-2000 m]
= Gennadas praecox Kemp, 1910: 176; Plate 13, fgs 1-4. [Investigator stn 320, of C. Comorin, 723'N
7544'E, 1053 fms]
= Gennadas caini Tirmizi, 1960: 368; Figs 40g, 48g, 84. [John Murray Expedition, stn 171 (907'06''N
5527'06''E to 908'48''N 5531'48''E), central part of Arabian Sea, 3840-3872 m]
Bentheogennema stephenseni Burkenroad, 1940: 37. [Dana stn 3624 I (SE of New Caledonia, 2817.6'N
17701'E, 5000 m wire out)]
Benthesicymus Spence Bate, 1881
= Benthesicymus Spence Bate, 1881 (type species Benthesicymus crenatus Spence Bate, 1881, desig-
nated by Spence Bate, 1888, gender masculine)
= Bentheocetes Smith, 1884 (type species Benthesicymus bartleti Smith, 1882; by monotypy; gender
masculine)
Benthesicymus altus Spence Bate, 1881: 191. [between Australia and Japan, 350-1400 fms]
Benthesicymus armatus MacGilchrist, 1905: 235. [Investigator stn 287 (Arabian Sea, 218'30''N 6547'E),
1506 fms]
Benthesicymus bartleti Smith, 1882: 82; Plate 14, fgs 1-7. [Blake stn 343, 3945'40''N 7055'W, 732 fms]
= Benthesicymus pleocanthus Spence Bate, 1888: 334; Fig. 48; Plate 57, fg. 2. [Challenger stns 23, of
Sombrero Island, 1824'N 6328'W, 450 fms; 205, Philippine Islands, 1642'N 11922'E, 1050 fms; 250,
North Pacifc Ocean, 3749'N 16647'W, 3050 fms]
Benthesicymus brasiliensis Spence Bate, 1881: 191. [Atlantic and Pacifc Oceans; according to Spence Bate,
1888: Challenger stns 323, east of Buenos Ayres, 3539'S 5047'W, 1900 fms; 168, of New Zealand,
4029'S 17743'E, 1100 fms; 173, of Matuku, Fii Islands, 199'35"S 17941'50"E, 315 fms; 181, be-
tween Australia and the Solomon Islands, 1350'S 15149'E, 2440 fms; 184, near Torres Strait, 128'S
14510'E, 1400 fms; 285, South Pacifc, 3236'S 13743'W, 2375 fms]
= Benthesicymus moratus Smith, 1886a: 90. [Albatross stns 2042 (3933'00"N 6826'45"W, 1555 fms);
2174 (3815'00"N 7203'00"W, 1594 fms); 2222 (3903'15"N 7050'45"W, 1537 fms); 2575 (4107'00"N
6526'30"W, 1710 fms)]
Benthesicymus cereus Burkenroad, 1936a: 30; Figs 6, 11-12, 19, 24, 28, 35, 42, 47. [Pawnee stn 54, 2115'40"N
7117'06"W, 900-945 fms]
Benthesicymus crenatus Spence Bate, 1881: 190. [mid Pacifc, 2600 fms; according to Spence Bate, 1888:
Challenger stns 272, north of the Low Archipelago, 348'S 15256'W, 2600 fms; 276, near the Low
Archipelago, 1328'S 14930'W, 2350 fms]
Benthesicymus investigatoris Alcock & Anderson, 1899: 282. [Investigator stns 222 (Andaman Sea, 1327'N
9314'30''E), 400-200 fms (200-405 fms?); 228 (Andaman Sea, 137'N 9444'15''E), 640 fms; 234 (Anda-
man Sea, 1315'30''N 9326'E), 498 fms; 235 (Andaman Sea, 1413'N 9340'E), 370-419 fms]
Benthesicymus iridescens Spence Bate, 1881: 191. [near the island of Tristan dAcunha, 1900 fms; according
to Spence Bate, 1888: Challenger stn 133, near Tristan da Cunha, 3541'S 2055'W, 1900 fms]
= Benthesicymus mollis Spence Bate, 1888: 339; Plate 58, fg. 2. [Challenger stn 133, near Tristan da
Cunha, 3541'S 2055'W, 1900 fms]
= Benthesicymus longipes Bouvier, 1906a: 747. [Princesse-Alice stn 1150, au sud-ouest des les du Cap-
Vert, 3890 m; according to Bouvier, 1908a]
Fig. 2. Benthesicymus tanneri Faxon, 1893. Photo by E. del Solar.
Benthesicymus laciniatus Rathbun, 1906: 906; Fig. 59; Plate 19, fg. 3. [Albatross stn 4018, vicinity of Kauai
Island, Hawaii, 804-724 fms]
= Benthesicymus Hjorti Sund, 1920: 30; Fig. 48; Plate 11, fg. 4. [Michael Sars stn 35 (2727'N 1452'W),
S of Canary Islands, 2603 m]
= Gennadas pectinatus Schmit, 1921: 25; Fig. 12; Plate 11, fg. 1. [Albatross stn 4390, of Santa Catalina
Island, California, 1350-2182 m]
Benthesicymus seymouri Tirmizi, 1960: 328; Figs 14-23. [John Murray Expedition, stn 118, 405'54"S
4110'12"E, 1789 m]
Benthesicymus strabus Burkenroad, 1936a: 45; Figs 10, 23, 27, 34, 41, 46, 49. [Challenger stn 285, 3236'S
13743'W]
Benthesicymus tanneri Faxon, 1893: 215. [Albatross stns 3358 (of Panama, 630'00"N 8144'00"W, 555 fms);
3362 (of Panama, 556'00"N 8510'30"W, 1175 fms); 3363 (of Panama, 542'00"N 8550'00"W, 978
fms); 3364 (of Panama, 530'00"N 8608'30"W, 902 fms); 3365 (of Panama, 531'00"N 8631'00"W,
1010 fms); 3366 (of Panama, 530'00"N 8645'00"W, 1067 fms); 3376 (of Panama, 309'00"N
8208'00"W, 1132 fms); 3377 (356'00"N 8140'15"W, 764 fms); 3380 (of Panama, 403'00"N
8131'00"W, 899 fms); 3384 (of Panama, 731'30"N 7914'00"W, 458 fms); 3393 (of Panama,
0715'00"N 7936'00"W, 1020 fms); 3400 (of Galapagos Islands, 036'00"S 8646'00"W, 1322 fms);
3403 (of Galapagos Islands, 058'30"S 8917'00"W, 383 fms); 3407 (of Galapagos Islands, 004'00"S
9024'30"W, 885 fms); 3410 (of Galapagos Islands, 019'00"N 9024'00"W, 331 fms); 3411 (of Galapa-
gos Islands, 054'00"N 9109'00"W, 1189 fms); 3418 (of Mexico, 1633'00"N 9952'30"W, 660 fms);
3424 (of Mexico, 2115'00"N 10623'00"W, 676 fms); 3425 (of Mexico, 2119'00"N 10624'00"W, 680
fms); 3435 (Gulf of California, 2648'00"N 11045'20"W, 859 fms); 3436 (2703'40"N 11053'40"W, 905
fms)] (Fig. 2)
Benthesicymus tirmiziae Crosnier, 1978: 16; Figs 7a-b, 8a-b, 11a. [Vauban stn CH 132, Madagascar, 1343.8'S
4729.0'E, 1950-2150 m]
Benthesicymus howensis Dall, 2001
= Benthesicymus urinator howensis Dall, 2001: 428; Figs 13A-D. [Lord Howe Rise, Tasman Sea, 2844'S
16154'E, 1325 m]
Benthesicymus urinator Burkenroad, 1936a: 29; Figs 4, 5, 8-9, 17-18, 22, 32, 33, 39-40, 45. [Challenger stn 184,
near Torres Strait, Australia, 128'S 14510'E, 1400 fms]
Benthonectes Smith, 1885a
= Benthonectes Smith, 1885a (type species Benthonectes flipes Smith, 1885a, by monotypy, gender
masculine)
= Hapalopoda Filhol, 1885a (type species Hapalopoda investigator Filhol, 1885a, by monotypy, gender
feminine)
Benthonectes flipes Smith, 1885a: 509. [Albatross stns 2181, 3929'00''N 7146'00''W, 603 fms; 2206,
3935'00''N 7124'30''W, 1043 fms; 2210, 3937'45''N 7118'45''W, 991 fms; 2235, 3912'00''N
7203'30''W, 707 fms]
= Hapalopoda investigator Filhol, 1885a: 229; Fig. 2. [Crevete prise 1900 mtres de profondeur,
bord du Talisman (339'N 1158'W (of Paris) (= 938'W of Greenwich))]
Gennadas Spence Bate, 1881
= Gennadas Spence Bate, 1881 (type species Gennadas parvus Spence Bate, 1881, by monotypy, gender
masculine)
= Amalopenaeus Smith, 1882 (type species Amalopenaeus elegans Smith, 1882, by monotypy, gender
masculine)
= Pasiphodes Filhol, 1885b (type species, Pasiphodes purpureus Filhol, 1885b, by monotypy, gender
masculine)
Gennadas barbari Vereshchaka, 1990: 131; Fig. 1. [stn 1925, 2539'S 8527'W]
Gennadas bouvieri Kemp, 1909: 726; Plate 74, fgs 1-4; Plate 75, fgs 6-7. [Challenger stns 206, W of Manila,
1754'N 11714'E, 2100 fms; 220, N of New Guinea, 042'S 147E, 1100 fms]
= Gennadas alcocki Kemp, 1910: 174 (?partim, males only); Plate 13, fgs 5-6. [Investigator stns 111, Bay
of Bengal, 1250'N 9052'E, 1644 fms; 103, Bay of Bengal, 1514'N 819'E, 1260 fms; 108, of C. Co-
morin, 74'N 7634'15"E, 1043 fms; 309, near the Andaman Islands, 109'N 932'15"E, 765 fms; see
Crosnier, 1978 for discussion of this problem]
Gennadas brevirostris Bouvier, 1905a: 748. [provient des parages de Sainte-Lucie o elle fut trouve par le
Blake sur des fonds de 221 brasses]
= Gennadas similis Stephensen, 1923: 6, 12; Fig. 1. [Atlantic, 3935'N 945'W, 1150 m]
= Gennadas chiasmifera Stephensen, 1923: 6, 13; Figs 2-3. [Atlantic, 3935'N 945'W, 1150 m]
Gennadas capensis Calman, 1925: 5; Plate 1, Figs 1-2. [SS Pickle stn 87, of Cape, South Africa, 1014 fms]
Gennadas crassus Tirmizi, 1960: 346, 371; Figs 40h, 85. [John Murray Expedition, stn 121 (539'00''N
3938'30''E to 540'30''N 3943'00''E), Zanzibar area, botom at 925 m, net apparently not on bot-
tom]
Gennadas elegans (Smith, 1882)
= Amalopenaeus elegans Smith, 1882: 87; Plate 14, fgs 8-14; Plate 15,fgs 1-5. [Blake stns 323, 3319'0"N
7612'30"W, 457 fms; 324, 3327'20"N 7553'30"W, 1386 fms; 325, 3335'20"N 760'0"W, 647 fms; 328,
3428'25"N 7522'50"W, 1632 fms; 330, 3141'0"N 7435'0"W, 1047 fms; 343, 3945'40"N 7055'0"W,
732 fms; 893, 3952'20"N 7058'0"W, 372 fms; 935, 3945'0"N 6944'45"W, 770 fms; 952, 3955'0"N
7028'0"W, 388 fms]
Gennadas gilchristi Calman, 1925: 6; Plate 1, fgs 3-4. [SS Pickle stns 77 (755 fms), 84 (500 fms), 87 (1014
fms), 357 (900 fms), of Cape, South Africa]
Gennadas incertus (Balss, 1927)
= Amalopenaeus incertus Balss, 1927: 265; fgs 24-29. [Valdivia stns 232, 326'S 5834'E, 0-1500 m; 235,
434'S 5342'E, 0-2000 m; 237, 445'S 4858'E, 0-2000 m]
= Amalopenaeus Gardineri Balss, 1927: 267; Fig. 31. [Valdivia stn 236, 438'S 5116'E, 0-2000 m; 4 miles
N.W. of Desroches Atoll, 750-0 fms]
Gennadas kempi Stebbing, 1914a: 283; Plate 27. [Scotia stn 468, Cape Point, South Africa, 3948'S 233'E,
2772 fms]
Gennadas parvus Spence Bate, 1881: 192. [of Japan, 2425 fms]
Gennadas propinquus Rathbun, 1906: 907; Fig. 61a-b. [Between Erben Bank and Kaiwi Channel, Hawaiian
Islands]
= Gennadas clavicarpus De Man, 1907a: 144 (partim). [Siboga stns 128, 427'N 12525.7'E, 0-700 m; 141,
10.4'S 12725.3'E, 0-1500 m; 230, 358'S 12820'E, 0-2000 m; see Burkenroad, 1936a]
= Gennadas scutatus indicus Kemp, 1913a: 62. [Investigator stns 108, of C. Comorin, 74'N 7634'15"E,
1043 fms; 109, of C. Comorin, 71'N 7821'E, 738 fms]
Gennadas scutatus Bouvier, 1906a: 748. [lHirondelle stn 156, entre les Aores et Terre-Neuve, surface ac-
cording to Bouvier, 1908a]
= Gennadas scutatus Bouvier, 1906b: 690. [nomen nudum]
Gennadas sordidus Kemp, 1910: 177; Plate 14, fgs 1-3. [Investigator stns 193, N of the Laccadive Islands,
1511'N 7228'45"E, 931 fms; 194, of the Laccadive Islands, 1347'N 723'45"E, 891 fms; 198, NE of
Ceylon, 855'N 8117'30"E, 764 fms]
Gennadas talismani Bouvier, 1906a: 749. [Talisman stn 105, 1638'N 1824'W, 3200 m, according to Crosnier
& Forest, 1973]
Gennadas tinayrei Bouvier, 1906c: 10; Figs 2-4, 14. [Mer des Sargasses, Aores, Cap Spartel (Princesse-Alice
stn 2264, 3730'N 2239'W, 0-3000 m, according to Davie, 2002: 127)]
Gennadas valens (Smith, 1884)
= Amalopenaeus valens Smith, 1884: 402; Plate 10, fg. 2. [Albatross stn 2003, 3716'30"N 7420'36"W,
640 fms]
= Pasiphodes purpureus Filhol, 1885b: Plate 3. [Type locality not indicated; Holthuis, 1980a refers to
A. Milne-Edwards original colour sketch of the species in the Paris Museum, probably from Talis-
man stn 38, of Cap Ghir, Morocco, 2210 m]
= Gennadas bidentata Stephensen, 1923: 6, 14; Figs 4, 5. [Atlantic, 5747'N 1133'W, 1985 m]
Family PENAEIDAE Rafnesque, 1815
Alcockpenaeopsis Sakai & Shinomiya, 2011
= Alcockpenaeopsis Sakai & Shinomiya, 2011 (type species Parapenaeopsis hungerfordi Alcock, 1905, by
original designation, gender feminine)
Alcockpenaeopsis hungerfordii (Alcock, 1905)
= Parapenaeopsis hungerfordii Alcock, 1905: 522, 530. [Hongkong]
Arafurapenaeopsis Sakai & Shinomiya, 2011
= Arafurapenaeopsis Sakai & Shinomiya, 2011 (type species Parapenaeopsis arafurica Racek & Dall,
1965, by original designation, gender feminine)
Arafurapenaeopsis arafurica (Racek & Dall, 1965)
= Parapenaeopsis arafurica Racek & Dall, 1965: 102; Fig. 16; Plate 8, fgs 6-7; Plate 13, fg. 5. [of the Fly
River, Papua New Guinea]
Artemesia Spence Bate, 1888
= Artemesia Spence Bate, 1888 (type species Artemesia longinaris Bate, 1888, by monotypy, gender
feminine)
Artemesia longinaris Spence Bate, 1888: 281; Plate 90. [Challenger stn 321, of Montevideo, Uruguay,
3502'S 5515'W, 13 fms]
= Artemesia brevinaris Nobili, 1901a: 1; Figs 1-2. [Mar del Plata, Argentina]
Atypopenaeus Alcock, 1905
= Atypopenaeus Alcock, 1905 (type species Penaeus compressipes Henderson, 1893, by original desig-
nation, gender masculine)
= Miyadiella Kubo, 1949a (type species Miyadella pedunculata Kubo, 1949a, by original designation,
gender feminine)
Atypopenaeus bicornis Racek & Dall, 1965: 85; Fig. 14; Plate 7, fgs 5-6; Plate 12, fg. 6. [Of Fly River mouth,
Papua New Guinea]
Atypopenaeus compressipes (Henderson, 1893)
= Penaeus compressipes Henderson, 1893: 450. [Gulf of Martaban, India]
= Parapenaeopsis brevirostris Kubo, 1936a: 55; Plate 61. [Inland Sea of Japan]
Atypopenaeus dearmatus De Man, 1907a: 135. [Siboga stns 302, 1017.9'S 12328.7'E, 216 m; 306, 827'S
12254.5'E, 247 m; 312, Saleh bay, N coast of Sumbawa, Indonesia, 819'S 11741'E, 274 m]
= Miyadiella ornata Holthuis, 1955a: 76; Figs 1-2. [Of NE Java, 733'S 11436'E, 200 m]
Atypopenaeus formosus Dall, 1957: 199; Fig. 21A-H. [5 km E of Sandgate, Moreton Bay, Queensland,
Australia, 2-3 fms]
Atypopenaeus stenodactylus (Stimpson, 1860a)
= Penaeus stenodactylus Stimpson, 1860a: 43. [Portu Hong Kong]
= Penaeus podophthalmus Stimpson, 1860a: 43. [Portu Hong Kong]
= Miyadella pedunculata Kubo, 1949a: 264; Figs 7N, 23G, H, 58O, 74C, I, 79E, 104, 105. [Osaka-wan,
Japan]
Batepenaeopsis Sakai & Shinomiya, 2011
= Batepenaeopsis Sakai & Shinomiya, 2011 (type species Penaeus tenellus Spence Bate, 1888, by origi-
Batepenaeopsis tenella (Spence Bate, 1888)
= Penaeus tenellus Spence Bate, 1888: 270. [Challenger stn 235, Bay of Kobe, Japan, 3439'N 13514'E,
8 fms]
= Penaeus crucifer Ortmann, 1890: 451; Plate 56, fg. 5. [Maizuru, Japan]
Batepenaeopsis venusta (De Man, 1907a)
= Parapeneopsis venusta De Man, 1907a: 134. [Siboga expedition, station of Pulu Jedan, E coast of Aru
Islands, shallow water]
Farfantepenaeus Burukovsky, 1997
= Penaeus (Farfantepenaeus) Burukovsky, 1997 (type species Penaeus [as Pne] brasiliensis Latreille,
1817, by original designation, gender masculine)
= Penaeus (Farfantepenaeus) Burukovsky, 1972 [no type species indicated]
Farfantepenaeus aztecus (Ives, 1891)
= Penaeus Brasiliensis var. Aztecus Ives, 1891: 190. [Veracruz, Mexico; lectotype designated by Prez
Farfante, 1969]
Farfantepenaeus brasiliensis (Latreille, 1817)
= Penaeus [as Pne] brasiliensis Latreille, 1817: 156. [ctes du Brsil]
Farfantepenaeus brevirostris (Kingsley, 1878a)
= Penaeus brevirostris Kingsley, 1878a: 98. [Estero at Realio, W. coast of Nicaragua]
Farfantepenaeus californiensis (Holmes, 1900)
= Penaeus californiensis Holmes, 1900: 218. [Santa Monica, California; neotype designation by Burken-
road, 1938]
Farfantepenaeus duorarum (Burkenroad, 1939)
= Penaeus duorarum Burkenroad, 1939: 31; Figs 18-19, 23, 25-27. [Atlantis stn 2813, of Alabama, Gulf
of Mexico, 19 fms]
Farfantepenaeus notialis (Prez Farfante, 1967)
= Penaeus duorarum notialis Prez Farfante, 1967: 94; Figs 4a-d. [Oregon stn 5664, of Las Piedras, Gulf
of Venezuela, 1144'N 7022'W, 26 fms]
= Penaeus duorarum var. cameronensis Rossignol, 1962 [unavailable name under Art. 15.2]
Farfantepenaeus paulensis (Prez Farfante, 1967)
= Penaeus (Melicertus) paulensis Prez Farfante, 1967: 84; Figs 1a-d. [Santos, So Paulo, Brazil]
Farfantepenaeus subtilis (Prez Farfante, 1967)
= Penaeus aztecus subtilis Prez Farfante, 1967: 89; Figs 2a-b, 3a-c. [Oregon stn 5685, of Gallinas Point,
Departemento de la Guajira, Colombia, 1229'N 7154'W, 95 fms]
Fenneropenaeus Prez Farfante, 1969
= Fenneropenaeus Prez Farfante, 1969 (type species Penaeus indicus H. Milne Edwards, 1837 [in H.
Milne Edwards, 1834-1840], by original designation, gender masculine)
Fenneropenaeus chinensis (Osbeck, 1765)
= Cancer chinensis Osbeck, 1765: 151. [Bocca Tiger, auf Chinesisch Pho-hao, der Tigermund, oder Pho-
munn, die Tigersfnung. Fischer kamen mit verschiedenen Fisharten zu uns, .... und chinesischen
Krabben: Cancer chinensis]
= Penaeus orientalis Kishinouye, 1917: 79. [Chosen, Kantoshu and Chintao]
Fenneropenaeus indicus (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840])
? = Palmon longicornis Olivier, 1811: 662. [Type locality not indicated]
= Penaeus indicus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 415. [les ctes de Coro-
mandel]
= Penaeus indicus longirostris De Man, 1892a: 511; Plate 29, fg. 53. [Celebes, Makassar]
Fenneropenaeus konkani Chanda & Bhatacharya, 2003: 229; Fig. 1. [Mirkarwada Fish landing centre, Ra-
tangiri, Maharstra, West Coast of India]
Fenneropenaeus merguiensis (De Man, 1888a)
= Penaeus merguiensis De Man, 1888a: 287; Plate 18, fg. 8; Plate 19, fg. 1. [Burma, Mergui Archipelago]
Fenneropenaeus penicillatus (Alcock, 1905)
= Penaeus indicus penicillatus Alcock, 1905: 515, 525. [Of the Orissa coast; at Bombay; Karchi;
Gangetic Delta; Mergui]
Fenneropenaeus silasi (Muthu & Motoh, 1979a)
=Penaeus silasi Muthu & Motoh, 1979a: 64; Figs 1-2. [Tawau, Sabah, 36 m]
Funchalia Johnson, 1868
= Funchalia Johnson, 1868 (type species Funchalia woodwardi Johnson, 1868, by monotypy, gender
feminine)
= Grimaldiella Bouvier, 1905b (type species Grimaldiella richardi Bouvier, 1905b, by monotypy, gender
feminine)
= Hemipenaeopsis Bouvier, 1905b (type species Hemipenaeopsis villosus Bouvier, 1905b, by monotypy,
gender feminine)
Funchalia danae Burkenroad, 1940: 36. [Dana stn 4017 VII (Canary Islands, 2913'N 1412'W, 1000 m wire
out)]
Funchalia sagamiensis Fujino, 1975: 200; Figs 1-3. [Amadai-ba, Sagami Bay, central Japan, upper water
layer]
Funchalia taaningi Burkenroad, 1940: 36. [Dana stn 3920 III (N of Seychelles, 106'N 6225'E, 3000 m wire
out)]
Funchalia villosa (Bouvier, 1905b)
= Hemipenaeopsis villosus Bouvier, 1905b: 981 (pro parte). [capture par la Princesse-Alice entre les les
Canaries et les Aores]
= Funchalia vanhfeni Lenz & Strunck, 1914: 306; Plate 17, fgs 20-21. [nrdlich von Tristan da Cun-
ha]
Funchalia woodwardi Johnson, 1868: 895. [Of Madeira, NE Atlantic Ocean]
= Peneus Genianus Risso, 1841 [nomen nudum]
= Sycionia geny Risso, 1844: 95. [Nice] [nomen nudum]
= Sicyonia Genyana Hope, 1851: 19. [nomen nudum]
= Grimaldiella Richardi Bouvier, 1905b: 981, 982. [capture par la Princesse-Alice dans se dernire
campagne, au large de lle Saint-Michel, en plein groupe des Aores]
= Peneus genyanus Monod, 1931: 107. [nomen nudum]
Ganjampenaeopsis Sakai & Shinomiya, 2011
= Ganjampenaeopsis Sakai & Shinomiya, 2011 (type species Parapenaeopsis uncta Alcock, 1905, by
Ganjampenaeopsis uncta (Alcock, 1905)
= Parapenaeopsis uncta Alcock, 1905: 522, 528. [Ganjam coast, 10-11 fms]
= Parapenaeopsis probata Hall, 1961: 96; Plate 19, fgs 11-13. [South of Bedok, 24 fms; South of Singa-
pore, 24 and 45 fms]
Heteropenaeus De Man, 1896
= Heteropenaeus De Man, 1896 (type species Heteropenaeus longimanus De Man, 1896, by monotypy,
gender masculine)
Heteropenaeus longimanus De Man, 1896: 111. [Java-See]
Holthuispenaeopsis Sakai & Shinomiya, 2011
= Holthuispenaeopsis Sakai & Shinomiya, 2011 (type species Parapenaeopsis atlantica Balss, 1914a, by
Holthuispenaeopsis atlantica (Balss, 1914a)
= Parapenaeopsis atlantica Balss, 1914a: 593. [Victoria, Kamerun]
= Trachypenaeus constrictus var. africana Balss, 1916: 17; Figs 3-4. [Goldkste: Kap Coast Castle, 13 m;
Addah, 11 m; Acra, 9 m. Togo, Lome, 13 m. Dahomey, Whyda, Cabinda, Landana, 7 m. Franzsisch-
Kongo: Nyanga-Flu, 13 m; Loango, 10 m. Angola: Ambriz, 13 m; Kinsembo, 13 m]
Kishinouyepenaeopsis Sakai & Shinomiya, 2011
= Kishinouyepenaeopsis Sakai & Shinomiya, 2011 (type species Penaeus cornutus Kishinouye, 1900, by
Kishinouyepenaeopsis amicus (V.C. Nguyn, 1971)
= Parapenaeopsis amicus V.C. Nguyn, 1971: 46; Fig. 1. [West Tonkin Gulf]
= Parapenaeopsis sinica Liu & Wang, 1986: 214, 215. [Wailuo, Xuwen County, Guangdong, South
China Sea]
Kishinouyepenaeopsis cornuta (Kishinouye, 1900)
= Penaeus cornutus Kishinouye, 1900: 23; unnumbered text fg. [Bay of Ariake, Kushu]
Kishinouyepenaeopsis incisa (Liu & Wang, 1986)
= Parapenaeopsis incisa Liu & Wang, 1986: 214, 215. [Wailuo, Xuwen County, Guangdong, South
China Sea]
Kishinouyepenaeopsis maxillipedo (Alcock, 1905)
= Parapenaeopsis maxillipedo Alcock, 1905: 522, 527. [Bombay, Madras, Arakan coast, India]
Litopenaeus Prez Farfante, 1969
= Litopenaeus Prez Farfante, 1969 (type species Penaeus vannamei Boone, 1931, by original designa-
tion, gender masculine)
Litopenaeus occidentalis (Streets, 1871a)
= Penaeus occidentalis Streets, 1871a: 243. [Isthmus of Panama]
Litopenaeus schmiti (Burkenroad, 1936b)
= Penaeus schmiti Burkenroad, 1936b: 315; Figs 1-3. [Kingston Bay, Jamaica]
Litopenaeus setiferus (Linnaeus, 1767)
= Cancer setiferus Linnaeus, 1767: 1054. [Indiis]
= Pneus orbignyanus Latreille, 1817: 155. [ctes maritimes du dpartement de la Vende]
= P.[enaeus] fuviatilis Say, 1818: 236. [North America]
= Penaeus gracilirostris Thallwitz, 1892: 3; Plate 1, fg. 5. [Nord-Celbes]
Litopenaeus stylirostris (Stimpson, 1871)
= Penaeus stylirostris Stimpson, 1871: 134. [Panama]
Litopenaeus vannamei (Boone, 1931)
= Penaeus vannamei Boone, 1931: 173; Fig. 16. [Golfo de Panama]
Macropetasma Stebbing, 1914b
= Macropetasma Stebbing, 1914b (type species Parapenaeus africanus Balss, 1913a, by monotypy, gen-
der neuter)
Macropetasma africana (Balss, 1913a)
= Parapenaeus africanus Balss, 1913a: 105. [Swakopmund, Namibia]
Marsupenaeus Tirmizi, 1971a
= Marsupenaeus Tirmizi, 1971a (type species Penaeus canaliculatus var. japonicus Spence Bate, 1888, by
monotypy, gender masculine)
Marsupenaeus japonicus (Spence Bate, 1888)
= Penaeus canaliculatus japonicus Spence Bate, 1888: 245; Plates 31; 32, fg. 4; 37, fg. 2. [Japan]
= Penaeus pulchricaudatus Stebbing, 1914b: 14; Plate 67. [Great Fish Point Lighthouse, Cape province,
South Africa]
Megokris Prez Farfante & Kensley, 1997
= Megokris Prez Farfante & Kensley, 1997 (type species Penaeus granulosus Haswell, 1879, by origi-
nal designation, gender masculine)
Megokris akademik Shinomiya & Sakai, 2006a: 1220; Figs 1-2. [R.V. Akademik stn PG-13-Ku, Persian
Gulf, 2913.587'N 4953.955'E, 41 m]
Megokris ghamrawyi Shinomiya & Sakai, 2006a: 1223; Figs 3-6. [R.V. Akademik stn PG-13-Ku, Persian
Gulf, 2913.587'N 4953.955'E, 41 m]
Megokris gonospinifer (Racek & Dall, 1965)
= Trachypenaeus gonospinifer Racek & Dall, 1965: 89; Fig. 15; Plate 7, fgs 7-8; Plate 12, fg. 7. [S. of Port
Romilly, Papua New Guinea, 755'S 14448'E, 7 fms]
Megokris granulosus (Haswell, 1879)
= Penaeus granulosus Haswell, 1879: 41. [Darney Island; lectotype designation by Grifn, 1970]
= Trachypeneus salaco De Man, 1907a: 135. [Siboga stns 205, Lohio bay, Buton Strait, Indonesia, 22 m;
258, Tual Anchorage, Kai Islands, 22 m]
= Trachypeneus furcilla Hall, 1961: 102; Plate 20, fgs 16-17. [Outer Shoal, Malaysia, 6 fms]
Megokris halli Shinomiya & Sakai, 2006b: 1255; Figs 1-2. [Singapore Fisheries Research stn B 146, Perma-
nent Buoy 10, Angler Buoy, 9 m]
Megokris manihine Shinomiya & Sakai, 2006b: 1259; Figs 3-4. [Singapore Fisheries Research stn c5/15,
512'N 11459'E, 37 m]
Megokris motohburiorum Shinomiya & Sakai, 2006b: 1261; Fig. 5. [Iloilo, Philippines]
Megokris pescadoreensis (Schmit, 1931a)
= Trachypeneus pescadoreensis Schmit, 1931a: 265; Plate 32, fgs 2-4. [Pescadores Islands, Formosa]
Megokris sedili (Hall, 1961)
= Trachypeneus sedili Hall, 1961: 100; Plate 20, fgs 18-19. [Malaysia, S of Bedok, 24 fms]
Melicertus Rafnesque, 1814
= Melicertus Rafnesque, 1814 (type species Melicertus tigrinus Rafnesque, 1814, by monotypy, gen-
der masculine)
Melicertus canaliculatus (Olivier, 1811)
= Palmon canaliculatus Olivier, 1811: 660. [Indonesia, N Sumatra, Pulu Weh; neotype designated
by Prez Farfante, 1976]
Melicertus hathor (Burkenroad, 1959)
= Penaeus latisulcatus hathor Burkenroad, 1959: 80; Figs 7, 8. [Mersa Thlemel, Gulf of Suez]
Melicertus kerathurus (Forskl, 1775)
= Cancer kerathurus Forskl, 1775: 95. [Smirn & Alexandri]
= Palmon sulcatus Olivier, 1811: 661. [Elle se trouve aux environs de Smyrne & dAlexandrie]
= Alpheus Trisulcatus Leach, 1814 [in Leach, 1813-1814]: 431. [Anglesea]
= Melicertus tigrinus Rafnesque, 1814: 22. [Sicily]
= Alpheus caramote Risso, 1816: 90. [Nice]
= Peneus mars Risso, 1816: 47; Plate 2, fg. 5. [Nice, se tient une grande profondeur]
= Alpheus punctulatus Risso, 1822: 247. [mer de Nice, rgions sablonneuses]
= Peneus cristatus Risso, 1827: 67. [environs de Nice et des Alpes Maritimes; grandes profondeurs]
= Penaeus fasciatus Hope, 1851: 19. [nomen nudum]
Melicertus latisulcatus (Kishinouye, 1896)
= Penaeus latisulcatus Kishinouye, 1896: 372. [Kagoshima, Tokyo Bay, Japan]
Melicertus longistylus (Kubo, 1943)
= Penaeus longistylus Kubo, 1943: 200. [of South-East coast of Hainan Island]
= Penaeus jejunus Hall, 1956: 75; Plate 10, fgs 9-10. [probably from the neighbourhood of Singapore]
= Penaeus caesius Dall, 1957: 143; Fig. 2A-G. [12 miles NNE of Bowen, Queensland, Australia, 19-25 fms]
Melicertus marginatus (Randall, 1840) (Fig. 3)
= P.[enus] marginatus Randall, 1840: 146. [Sandwich Islands]
= Penaeus teraoi Kubo, 1949a: 288; Figs 7F, 20F, 24N-P, 49K, 58G, 73D, J, 77S, 112. [Miya, Aiti Prefec-
ture, Japan]
Melicertus plebejus (Hess, 1865)
= Penaeus plebejus Hess, 1865: 168; Plate 7, fg. 19. [Sydney]
= Penaeus canaliculatus var. australiensis Spence Bate, 1888: 248; Plate 32, fg. 3. [Port Jackson, Aus-
tralia, 2-10 fms]
= Penaeus maccullochi Schmit, 1926a: 359, 370; Plate 66, fgs 1-3. [21 km NE of North Reef, Queens-
land, Australia]
Melicertus similis Chanda & Bhatacharya, 2002: 495; Figs 1-6. [Port Blair, Andaman Is., Bay of Bengal]
Metapenaeopsis Bouvier, 1905b
= Metapenaeopsis Bouvier, 1905b (type species Metapenaeopsis pubescens Bouvier, 1905b, by mono-
typy, gender feminine)
= Archipenaeopsis Bouvier, 1905b (type species Archipenaeopsis vesitus Bouvier, 1905b, by monotypy,
gender feminine)
= Leptopenaeus Kishinouye, 1929 (type species Penaeus philippii Spence Bate, 1888, designated by
Prez Farfante & Kensley, 1997, gender masculine)
= Ceratopenaeus Kishinouye, 1929 (type species Parapenaeus mogiensis Rathbun, 1902b, designated by
= Erythropenaeus Kishinouye, 1929 (type species Parapenaeus akayebi Rathbun, 1902b, designated by
Metapenaeopsis acclivis (Rathbun, 1902b)
= Parapenaeus acclivis Rathbun, 1902b: 41; Figs 12-14. [Mogi, Japan]
Metapenaeopsis aegyptia Galil & Golani, 1990: 229; Figs 1a-b, 2a-c, 3a. [of Palmahim, Israel, 3156'N
3435'E, 50 m]
= Metapenaeopsis judaensis Por, 1989: 211. [nomen nudum]
Metapenaeopsis andamanensis (Wood-Mason in Wood-Mason & Alcock, 1891a)
= Metapenaeus philippinensis var. andamanensis Wood-Mason in Wood-Mason & Alcock, 1891a: 271.
[N of Port Blair, Andaman Sea, 112-244 fms]
Metapenaeopsis angusta Crosnier, 1987a: 441; Figs 16b, 18b, 19c-d. [MUSORSTOM I, stn 5, 1401.5'N
12023.5'E, 215-200 m]
Metapenaeopsis assimilis (De Man, 1920a)
= Penaeopsis assimilis De Man, 1920a: 105. [Of Pulu Weh, N Sumatra, Indonesia; lectotype designa-
tion by Crosnier, 1991]
= Metapenaeopsis raceki Starobogatov, 1972: 405, 409; Plate 10, fg. 127a-b. [nomen novum for Meta-
penaeopsis distincta sensu Racek & Dall, 1965 nec De Man, 1907a]
Metapenaeopsis barbata (De Haan, 1844 [in De Haan, 1833-1850])
= Penaeus barbatus De Haan, 1844 [in De Haan, 1833-1850]: Plate 46, fg. 3. [Japan]
= Parapenaeus akayebi Rathbun, 1902b: 39. [Mogi, Japan]
Metapenaeopsis batei (Miers, 1884b)
= Penus batei Miers, 1884b: 296; Plate 32, fg. D. [Albany Island, 3-4 fms]
Metapenaeopsis beebei (Burkenroad, 1938)
= Penaeopsis (Metapenaeopsis) beebei Burkenroad, 1938: 74; Figs 18-19. [Templeton Crocker Expedition
stn 136, dredge 30, Arena Bank, Gulf Of California, 2327'N 10924'W, 35 fms]
Fig. 3. Melicertus marginatus (Randall, 1840). Photo by Tin-Yam Chan.
Metapenaeopsis ceylonica Starobogatov, 1972: 406, 408 (key); Plate 11, fg. 159a-b; nomen novum for M.
hilarulus sensu De Bruin, 1965 nec De Man, 1911a. [Ceylon, W coast, W of Mannar, 6 fms]
Metapenaeopsis commensalis Borradaile, 1899: 1001, Plate 63, fg. 1-1b. [Ellice Islands, Rotuma, Fii]
= Penaeopsis Borradailei De Man, 1911a: 73. [Siboga stn 131, anchorage of Beo, Karakelong Island,
Talaud Archipelago, Indonesia; lectotype designation by Crosnier, 1991]
Metapenaeopsis coniger (Wood-Mason in Wood-Mason & Alcock, 1891a)
= Metapenaeus coniger Wood-Mason in Wood-Mason & Alcock, 1891a: 272. [Investigator stns 119 (Bay
of Bengal, 1220'N 858'E), 95 fms; of Mahnaddi Delta, 68 fms; 96 (Bay of Bengal, 1830'N 8446'E),
98-102 fms]
Metapenaeopsis costata Crosnier, 1991: 275; Figs 83, 84. [MUSORSTOM 3, stn CP 142, Philippines, 1147'N
12301.5'E, 26 m]
Metapenaeopsis crassissima Racek & Dall, 1965: 26; Fig. 2C; Plate 2, fg. 1; Plate 4, fgs 5-6; Plate 9, fg. 3.
[Shark Bay, Western Australia]
Metapenaeopsis dalei (Rathbun, 1902b)
= Parapenaeus dalei Rathbun, 1902b: 40; Figs 9-11. [Mogi, Japan]
? = Metapenaeopsis incomptus Kubo, 1949a: 424; Figs 8D, 46A, 76L, R, 80N, 148F. [East China Sea]
Metapenaeopsis difcilis Crosnier, 1991: 255; Figs 65-67. [MUSORSTOM 3, stn CP 134, Philippines,
1201.1'N 12157.3'E, 92-95 m]
Metapenaeopsis distincta (De Man, 1907a)
= Metapeneus distinctus De Man, 1907a: 132. [Siboga stn 184, Anchorage of kampong Kelang, south
coast of Manipa-island, 36 m; female specimen illustrated by De Man, 1913; designated as lectotype
by Crosnier, 1991]
Metapenaeopsis dura Kubo, 1949a
= Metapenaeopsis durus Kubo, 1949a: 421; Figs 8A, 18G-L, 22O, 46D, 64C-C', 76E, J, 80L, 148C, 149.
[Tokyo fsh market, Miya, Aichi Prefecture, Japan]
Metapenaeopsis erythraea Crosnier, 1987a: 443; Figs 16c, 18c, 19e-f. [Valdivia stn 122, Mer Rouge, 2122'N
3904'E, 383-363 m]
Metapenaeopsis evermanni (Rathbun, 1906)
= Metapenaeus evermanni Rathbun, 1906: 904; Fig. 58. [Albatross stn 3849, south coast of Molokai Is-
land, Hawaii Islands, 79-134 m]
Metapenaeopsis faouzii (Ramadan, 1938)
= Penaeopsis (Metapenaeus) faouzii Ramadan, 1938: 72; Fig. 14d. [John Murray Expedition stn 161,
Maldives, 504'48"N 7250'30"E, 46 m; lectotype designation by Crosnier, 1991]
Metapenaeopsis fusca R.J.G. Manning, 1988: 91; Fig. 1A-D. [Singleton, south Western Australia (3227'S
11544'E)]
Metapenaeopsis gaillardi Crosnier, 1991: 200; Figs 25-27. [Lagon stn 314, Iles des Pins, Nouvelle-Caldonie,
27 m]
Metapenaeopsis gallensis (Pearson, 1905)
= Parapenaeus gallensis Pearson, 1905: 72; Plate 1, fgs 3, 3a-b. [S of Galle, Ceylon]
Metapenaeopsis gerardoi Prez Farfante, 1971a: 20; Figs 11-12, 13C. [Oregon stn 5440, of Mayagez, Puerto
Rico, 1808.5'N 6723'W, 22 m]
Metapenaeopsis goodei (Smith, 1885b)
= Parapenaeus Goodei Smith, 1885b: 176. [Bermuda; Bay of Panama]
= Archipenopsis vestitus Bouvier, 1905a: 747. [captur par le Blake dans la mer des Antilles, la faible
profondeur de 37 brasses (Blake stn 11, NW of Dry Tortugas Islands, 2443'N 8325'W, 37 fms, see A.
Milne-Edwards & Bouvier, 1909 and Prez Farfante, 1971a)]
= Parapenaeopsis Rathbuni Bouvier, 1905a: 748. [dans la mer des Antilles, 17 brasses de profondeur
(2616'25''N 8250'10''W, west of Florida; see Prez Farfante, 1971a)]
Metapenaeopsis hilarula (De Man, 1911a)
= Penaeopsis sp. (provisional name hilarulus) De Man, 1911a: 70 (partim). [Siboga stn 240, Banda-an-
chorage, Indonesia, 9-36 m; lectotype designation by Crosnier, 1991]
Metapenaeopsis hobbsi Prez Farfante, 1971a: 24; Figs 13D, 14-17. [Pillsbury stn 737, NW of Cabo Cordera,
Venezuela, 1044'N 6607'W to 1045'N 6608'W, 60-73 m]
Metapenaeopsis incisa Crosnier, 1991: 277; Figs 85-87. [BENTHEDI stn 101D, Iles Glorieuses, 1125.7'N
4719.5'E, 26 m]
Metapenaeopsis ivanovi Crosnier, 1994a: 340; Figs 1-3. [au large du Kenya]
Metapenaeopsis kishinouyei (Rathbun, 1902c)
= Parapenaeus kishinouyei Rathbun, 1902c: 288; Plate 12, fgs 13-15. [Tagus Cove, reef N of Tagus Hill,
Albemarle Island]
Metapenaeopsis kuboi Ivanov & Hassan, 1976a: 1302; Figs 4, 5d, 6f. [Western Indian ocean, 2326'S 3331'E,
410 m]
Metapenaeopsis kyushuensis (Yokoya, 1933)
= Leptopenaeus Kyushuensis Yokoya, 1933: 5; Fig. 2. [Sy-Maru stns 296, E of Tanegasima Island, Ja-
pan, 219 m; 317, coast of Miyazaki-ken, Japan, 97 m; 428, W of Amakusa Islands, Japan, 119 m
Metapenaeopsis lamellata (De Haan, 1844 [in De Haan, 1833-1850])
= Penaeus lamellatus De Haan, 1844 [in De Haan, 1833-1850]: Plate 46, Figs 4-5. [Japan; lectotype
designated by Yamaguchi & Baba, 1993]
Metapenaeopsis lata Kubo, 1949a
= Metapenaeopsis latus Kubo, 1949a: 434; Figs 8G, 46G, 76N, T, 81B, 148H, 150. [Of Nagashima, Mie
Prefecture, Japan, circa 350 m]
? = Leptopenaeus Kyushuensis Yokoya, 1933: 5; Fig. 2. [Sy-Maru stns 296, east of Tanegasima Isl., 219
m; 317, coast of Miyazaki-ken, 97 m; 428, west of Amakusa Isls., 119 m]
Metapenaeopsis laubieri Crosnier, 1991: 238; Figs 51-53. [Albatross 1907-1910, stn 5160, 2.75 mi SW de lle
Tinakta Island, Malaisie, 512'40"N 11955'10"E, 22 m]
Metapenaeopsis lindae R.J.G. Manning, 1988: 97; Fig. 2A-D. [Bell Buoy, NW Rotnest Island (3159'S
11534'E), south Western Australia]
Metapenaeopsis liui Crosnier, 1987a: 434; Figs 3d, 10b-d, 13d-f, 14a, 15. [MUSORSTOM I, stn 40, Philip-
pines, 1357.4'N 12027.8'E, 287-265 m]
Metapenaeopsis mannarensis De Bruin, 1965: 88; Fig. 2a-d. [Ceylon, of Mannar, 6 fms]
Metapenaeopsis manningi Crosnier, 1994a: 344; Figs 4-6. [Anton Bruun, Cruise 9, stn 499, Somalie, 1003'N
5115'E, 31-39 m]
Metapenaeopsis marquesas Crosnier, 1991: 244; Figs 57-59. [Nat. Geogr. Marquesas Exped. stn UP II/Haul
5, cte ouest de Ua Pou, Iles Marquises, Polynsie, 48-51 m]
Metapenaeopsis martinella Prez Farfante, 1971a: 16; Figs 9-10, 13B. [Oregon stn 4252, of Acara, Cear,
Brazil, 210'S 3952'W, 137 m]
Metapenaeopsis menoui Crosnier, 1991: 180; Figs 6c-d, 8-9. [Reves 2 stn 19, Seychelles, 554.5'S 5619.4'E,
30-35 m]
Metapenaeopsis miersi (Holthuis, 1952a)
= Penaeopsis miersi Holthuis, 1952a: 80; Fig. 19. [nomen novum for Penus velutinus sensu Miers,
1881a nec Dana, 1852a and nomen novum for Metapenopsis pubescens Bouvier, 1905b nec Penus
pubescens Stimpson, 1871]
= Metapenopsis pubescens Bouvier, 1905b: 981; nec Penus pubescens Stimpson, 1871. [les du Cap
Vert]
Metapenaeopsis mineri Burkenroad, 1934a: 25; Figs 8-10. [Conception Bay, Lower California, USA]
Metapenaeopsis mogiensis complanata Crosnier, 1991: 219; Figs 37o-r, 38d. [Soela Cruise 682, stn 128, cte
nord-ouest de Australie, 1907.9'S 11906.9'E, 78 m]
Metapenaeopsis mogiensis consobrina (Nobili, 1904)
= Metapenaeus consobrinus Nobili, 1904: 230. [Djibouti; lectotype designation by Crosnier, 1991]
= Metapenaeus perlarum Nobili, 1905a: 158. [ctes dArabie, dragages entre 2510'N 5510'E 2455'N
5440'E, 16-24 m, according to Crosnier, 1991]
= Metapenaeopsis bruini Starobogatov, 1972: 405, 408; Plate 11, fg. 158. [Sri Lanka, E of Mullaitivu
lighthouse, of Mannar, 6 fms; nomen novum for M. mogiensis sensu De Bruin, 1965 nec Rathbun,
1902b]
Metapenaeopsis mogiensis intermedia Crosnier, 1991: 222; Figs 37s-w, 38e. [MUSORSTOM 3, stn CP 142,
Philippines, 1147'N 12302'E, 26-27 m]
Metapenaeopsis mogiensis mogiensis (Rathbun, 1902b)
= Parapenaeus mogiensis Rathbun, 1902b: 39; Figs 6-8. [Mogi, Japan; lectotype designation by Cros-
nier, 1991]
Metapenaeopsis novaeguineae (Haswell, 1879)
= Penaeus Novae-Guineae Haswell, 1879: 43. [Chevert Expedition, Katow, Papua New Guinea]
Metapenaeopsis palmensis (Haswell, 1879)
= Penaeus Palmensis Haswell, 1879: 43. [Palm Island, Queensland, Australia]
= Metapenaeopsis barbeensis Hall, 1962: 32; Figs 118-118f. [35 miles ESE of St. Barbe (= Pejantan),
007'S 10746'E, 20 fms (37 m)]
Metapenaeopsis parahilarula Crosnier, 1991: 232; Figs 45e, 47. [MUSORSTOM 3, stn CP 121, Philippines,
1208.3'N 12117.3'E, 73-84 m]
Metapenaeopsis parapalmensis Crosnier, 1994b: 313; Figs 37-38. [Albatross stn 5159, archipel de Sulu,
groupe des Tawitawi, le Tinakta, 511'50"N 11954'E, 18-22 m]
Metapenaeopsis persica Crosnier, 1991: 210; Figs 32, 33a-d. [Dammam, district dHasa, Arabie Saoudite,
Golfe Persique, 2625'N 5006'E]
Metapenaeopsis philippii (Spence Bate, 1881)
= Penaeus Philippii Spence Bate, 1881: 181. [Challenger stn 201, 73'N 12148'E, 150 m; lectotype des-
ignation by Crosnier, 1987a]
= Penaeus philippinensis Spence Bate, 1888: 261; Plate 35, fgs 2-2", 3-3". [Challenger stn 201, 73'N
12148'E, 150 m; lectotype designation by Crosnier, 1987a]
Metapenaeopsis propinqua Crosnier, 1991: 186; Figs 12-13. [Patate bayonnaise, lagon est, Nouvelle-Cal-
donie, 20 m]
Metapenaeopsis provocatoria longirostris Crosnier, 1987a: 435, Figs 1d, 7g-j, 8m-o. [Soela Cruise 184 stn 08,
Australie, 1920.2'S 11544.1'E, 306-308 m]
Metapenaeopsis provocatoria provocatoria Racek & Dall, 1965: 48; Fig. 10A-D. [Challenger stn 31, NNE of
Cape Moreton, Queensland, Australia, 80-90 fms]
Metapenaeopsis provocatoria owstoni Shinomiya & Sakai, 2000: 123; Figs 7, 8a-f, 9a-m, 11, 12. [Okinose,
Sagami Bay, Japan, ca. 35N 13935'E]
Metapenaeopsis proxima Crosnier, 1991: 258; Figs 68-70, 78a. [REVES 2 stn 12, Seychelles, 513.5'S 5608.5'E,
60 m]
Metapenaeopsis quadrilobata Crosnier, 1991: 199; Figs 23-24. [Nosy Be (Pointe Lokobe), Madagascar, 10 m]
Metapenaeopsis quinquedentata (De Man, 1907a)
= Metapeneus quenquedentatus De Man, 1907a: 133. [Siboga stn 109, of Tongkil Island, Sulu Archi-
pelago, Philippines, 13 m; lectotype designation by Crosnier, 1991]
Metapenaeopsis richeri Crosnier, 1991: 280; Figs 88-89. [CORAIL 2, stn DW 147, lles Chesterfeld,
1936.87'S 15813.52'E, 25 m]
Metapenaeopsis rosea Racek & Dall, 1965: 29; Figs 2D, 3; Plate 1, fg. 4; Plate 4, Figs 7, 8; Plate 9, fg. 4.
[Mackay, Queensland, Australia, 4 fms]
Metapenaeopsis scoti Champion, 1973: 195; Figs 3A, 4A-D. [South Africa, Natal, of Durban, 160 fms]
Metapenaeopsis sibogae (De Man, 1907a)
= Metapeneus sibogae De Man, 1907a: 131. [Siboga stns 306, 827'S 12254.5'E, 247 m; 312, Saleh-bay,
north coast of Sumbawa, Indonesia, 819'S 11741'E, 274 m]
Metapenaeopsis sinica Liu & Zhong, 1988: 3, 224, 270; Fig. 136. [Western Guangdong, South China Sea, 64 m]
= Metapenaeopsis sinensis Liu & Zhong, 1988: 24, 216 (key). [nomen nudum]
Metapenaeopsis sinuosa Dall, 1957
= Metapenaeopsis sinuosus Dall, 1957: 176; Fig. 14A-F. [Of shore island (Kennedy Sound), Cumber-
land Group, NE coast of Australia, 16 m]
Metapenaeopsis smithi (Schmit, 1924a)
= Penaeopsis smithi Schmit, 1924a: 62; Figs 1b-c, 2a-c. [Caracas Baai, Curaao]
Metapenaeopsis spatulata Crosnier, 1991: 224; Figs 40-41. [MUSORSTOM 3, stn CP 142, Philippines,
1147'N 12302'E, 26-27 m]
Metapenaeopsis spiridonovi Crosnier, 1991: 268; Figs 77a-f, 78d. [REVES 2 stn 33, Seychelles, 425.9'S
5439.0'E, 45-60 m]
Metapenaeopsis stokmani Burukovsky, 1990: 189; Fig. 1A. [2539'S 8524'W, 160-192 m]
Metapenaeopsis stridulans (Alcock, 1905)
= Metapenaeus stridulans Alcock, 1905: 518, 526. [from Orissa to Palk Strait, 20-35 fms; Gulf of Marta-
ban, 20 fms; various places in the Andamans, 20 fms]
? = Metapenaeopsis tchekunovae Starobogatov, 1972: 402, 413; Plate 9, fg. 107. [Arabian Sea of Paki-
stan, of Indus delta, 2348.5' - 2347.7'N 6717.5' - 6718.7'E]
Metapenaeopsis tarawensis Racek & Dall, 1965: 46; Fig. 9A-D. [Tarawa Group, Gilbert Islands]
Metapenaeopsis tenella Liu & Zhong, 1988: 242; Fig. 146. [East of Hainan Island, South China Sea, 270 m]
Metapenaeopsis toloensis Hall, 1962
= Metapenaeopsis tolnsis Hall, 1962: 33; Figs 119-119d. [Central part of South China Sea, 613'N
10749'E, 40 fms; 551'N 10753'E, 38 fms]
Metapenaeopsis vaillanti (Nobili, 1904)
= Metapenaeus vaillanti Nobili, 1904: 229. [Suez; Mer Rouge]
? = Metapenaeopsis philippii var. Ataqa Al-Kholy & El-Hawary, 1970: 361. [Gulf of Suez till El-Adabi-
ah, 5-8 fms]
Metapenaeopsis velutina (Dana, 1852a)
= Penaeus velutinus Dana, 1852a: 27. [Albatross 1902, stn 3853, cte sud de lle Molokai, Hawaii, 210-
245 m; neotype designation by Crosnier, 1991]
= Metapenaeopsis insona Racek & Dall, 1965: 41; Fig. 6A-B; Plate 2, fg. 3; Plate 10, fg. 2. [7 miles of
Long Reef, New South Wales, Australia, 73 m]
= Metapenaeopsis caliper Liu & Zhong, 1988: 238, 269; Fig. 145. [S of mouth of Pearl River, South
China Sea, 138 m]
Metapenaeopsis wellsi Racek, 1967: 251; Plates 12-13. [Exmouth Gulf, Western Australia]
Metapenaeus Wood-Mason in Wood-Mason & Alcock, 1891a
= Mangalura Miers, 1878 (type species Penus dobsoni Miers, 1878, by monotypy, gender feminine;
name placed on the Ofcial List of Generic Names in Zoology in Opinion 864 in 1969: ruled under
the plenary powers not to be given precedence over Metapenaeus Wood-Mason, 1891)
= Metapenaeus Wood-Mason in Wood-Mason & Alcock, 1891a (type species Penaeus afnis H. Milne
Edwards, 1837 [in Milne Edwards, 1834-1840], by original designation, gender masculine)
Metapenaeus afnis (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840])
= Penaeus afnis H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 416. [cte de Malabar]
= Penaeus mutatis Lanchester, 1901: 572; Plate 34, fg. 6. [Type locality not indicated; lectotype desig-
nation by Miquel, 1982]
= Metapenaeus necopinans Hall, 1956: 83; Plate 11, fg. 15; Plate 12, fg. 16. [Jurong, Singapore]
= Metapenaeus ivanovi Hassan, 1978: 385; Figs 1-2. [Arabian Gulf, 29145'N 4930'E, 25 m]
Metapenaeus alcocki M.J. George & Rao, 1968: 146; Fig. 1. [Gulf of Kutch, Northwest coast of India,
2248'N 7003'E, 3-12 m]
Metapenaeus anchistus (De Man, 1920a)
= Penaeopsis intermedia anchista De Man, 1920a: 5; Plate 1, Figs 3-3d. [Ternate, Indonesia; lectotype
designation by Miquel, 1982]
Metapenaeus arabicus Hassan, 1978: 387; Figs 3-5. [Arabian Gulf, 29145'N 4930'E, 25 m]
Metapenaeus bennetae Racek & Dall, 1965: 74. [Lake Budgewoi, Tuggerah Lakes, New South Wales, Aus-
tralia]
Metapenaeus brevicornis (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840])
= Penaeus brevicornis H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 417. [ctes de lInde]
= Penaeus avirostris Dana, 1852a: 27. [Singapore]
Metapenaeus conjunctus Racek & Dall, 1965: 64; Fig. 12; Plate 5, fgs 8-9; Plate 11, fg. 1. [Shallow brackish
water at mouth of Tuaran river, Sabah, N Borneo, Malaysia, 4-5 fms]
Metapenaeus dalli Racek, 1957: 4; Plate 1, fgs 1-3; Plate 2, fgs 1-2; Plate 3, fg. 3. [Peel Inlet, Murray River,
West Australia]
Metapenaeus demani demani (Roux, 1921)
= Penaeopsis demani Roux, 1921: 599; Plate 16, fgs 4-12. [Varen and Lorentz Rivers, New Guinea]
Metapenaeus demani stephani Miquel, 1982: 74; Figs 24, 25a, c, 26a-e. [Port Moresby, Golfe de Papoua,
Nouvelle-Guine]
Metapenaeus dobsoni (Miers, 1878)
= Penus dobsoni Miers, 1878: 302; Plate 17, fg. 2. [Mangalur (Mangalore), west coast of India] [as
Mangalura dobsoni on page 303]
= Metapenaeus dobsoni choprai Holthuis, 1980b: 22. [nomen nudum]
Metapenaeus eboracensis Dall, 1957: 193; Fig. 19a-g. [Mouth of Norman River, Gulf of Carpentaria,
Queensland, Australia, 2-4 m]
Metapenaeus elegans De Man, 1907a: 130. [Siboga stn 121, Manado, Clbes, Indonesia, 55 m]
= Metapenaeus singaporensis Hall, 1956: 84; Plate 12, fgs 17-19. [Jurong, Singapore]
Metapenaeus endeavouri (Schmit, 1926a)
= Penaeopsis endeavouri Schmit, 1926a: 329; Plate 59, fgs 1-3; Plate 68, fg. 4. [FIS Endeavour expedi-
tion, S coast of Queensland, Australia]
Metapenaeus ensis (De Haan, 1844 [in De Haan, 1833-1850])
= Penoeus Ensis De Haan, 1844 [in De Haan, 1833-1850]: Plate 46, fg. 2. [Japan; lectotype designation
by Miquel, 1982]
= Penaeus Mastersii Haswell, 1879: 42. [80 km WNW of Darwin, Northern Territory, Australia]
= Penaeus incisipes Spence Bate, 1888: 257; Plate 34, fg. 2. [Challenger stns 190, Arafura Sea, south of
Papua, 856'S 1365'E, 49 fms; 203, of Panay, Philippines, 116'N 1239'E, 20 fms]
= Metapenaeus ensis var. baramensis Hall, 1962: 23. [30 miles NNE of Tanjong Baram, 458'N 11358'E,
77 m]
= Metapenaeus philippinensis Motoh & Muthu, 1979: 1351; Figs 1-2. [Himamaylan, Philippines, 10-
15 m]
Metapenaeus insolitus Racek & Dall, 1965: 69; Fig. 13A-C; Plate 6, fgs 3, 4; Plate 11, fg. 4. [Chambers Bay,
Northern Territory, Australia, 30 m]
Metapenaeus intermedius (Kishinouye, 1900)
= Penaeus intermedius Kishinouye, 1900: 21, unnumbered textfg. [province of Tosa]
Metapenaeus joyneri joyneri (Miers, 1880)
= Penaeus Joyneri Miers, 1880: 458; Plate 15, fgs 8-10. [Yokohama, Japan]
= Penaeus pallidus Kishinouye, 1897: 275. [Tokyo Bay, Ise area, Seto Inland Sea; Kagoshima Bay]
Metapenaeus joyneri formosus Lee & Yu, 1977: 101; Fig. 71. [Donggang, Pingtong County, SW Taiwan]
Metapenaeus krishnatrii Silas & Muthu, 1976: 645; Plate 1; fg. 1. [Corbyn Cove, Port Blair, Andaman Is-
lands]
Metapenaeus kutchensis P.C. George, M.J. George & Rao, 1963: 284; Fig. 1. [Gulf of Kutch, N.W. India,
3-12 m]
Metapenaeus lysianassa (De Man, 1888a)
= Penaeus lysianassa De Man, 1888a: 290. [Mergui Archilepago, Burma]
= Metapenaeus lysianassa var. malaccnsis Hall, 1962: 24. [Kuala Sanglang, 1 m]
Metapenaeus macleayi (Haswell, 1879)
= Penus macleayi Haswell, 1879: 40. [Port Jackson, New South Wales, Australia]
= Penaeus haswelli Phillips, 1925: 3; Plate 2, fg. 1. [shallow waters of New South Wales]
Metapenaeus monoceros (Fabricius, 1798)
= Penaeus monoceros Fabricius, 1798: 409. [Type locality restricted by Holthuis & Gotlieb, 1958 to
Tranquebar, Madras Province, India]
? = Penaeopsis spinulicauda Stebbing, 1914b: 17; Plate 68. [Durban Bay]
? = Metapenaeus cognatus Nobili, 1904: 229. [Djibouti, Mer Rouge]
? = Metapenaeus Deschampsi Nobili, 1903a: 452; Plate 2, fg. 1. [Pondichry, Foce dellArian Koupur;
Mah]
Metapenaeus motohi Shinomiya & Sakai, 2009: 1068; Fig. 1. [Tigbauan Panay I., Philippines]
Metapenaeus moyebi (Kishinouye, 1896)
= Penaeus moyebi Kishinouye, 1896: 373. [Japan; Makassar, Celebes, Indonesia]
= Metapenaeus burkenroadi Kubo, 1954: 92; Fig. 1A-D. [Japan]
Metapenaeus papuensis Racek & Dall, 1965: 66; Plate 3, fg. 3; Plate 6, fgs 1-2; Plate 11, fg. 2. [Mouth of
Panaroa river, Gulf of Papua, Papua New Guinea]
= Metapenaeus bengalensis Tirmizi, 1971b: 242; Figs 1-3. [Bay of Bengal]
Metapenaeus stebbingi Nobili, 1904
= Metapenaeus Stebbingi Nobili, 1904: 229. [Mer Rouge; Suez; full description in Nobili, 1906a]
Metapenaeus suluensis Racek & Dall, 1965: 61; Fig. 11A-B; Plate 5, fgs 6-7; Plate 10, fg. 8. [Philippines,
Sulu Sea, 20 fms]
Metapenaeus tenuipes Kubo, 1949a: 348; Figs 7R, T, 22B, 31G, H, 47G, 62B-B', 74N-M, S-T, 81H, K, 125G,
127, 129F, 130. [S coast of Borneo, Indonesia]
= Metapenaeus spinulatus Kubo, 1949a: 355; Figs 7T, 47G, 74M, S, 81K, 129F, 130. [Bangkok, Thailand]
Mierspenaeopsis Sakai & Shinomiya, 2011
= Mierspenaeopsis Sakai & Shinomiya, 2011 (type species Penaeus sculptilis Heller, 1862a, by original
designation, gender feminine)
Mierspenaeopsis cultrirostris (Alcock, 1906)
= Parapenaeopsis sculptilis var. cultrirostris Alcock, 1906: 39; Plate 7, fg. 23. [India: Orissa and Ganjam;
Sunderbunds]
Mierspenaeopsis hardwickii (Miers, 1878)
= Penus hardwickii Miers, 1878: 300; Plate 17, fgs 1-1a. [Indian Seas?]
Mierspenaeopsis indica (Muthu, 1972)
= Parapenaeopsis indica Muthu, 1972: 174; fgs 1-6. [Kakinada Bay, Kakinada, east coast of India,
1656.5'N 8216.5'E, 1-4 m]
Mierspenaeopsis sculptilis (Heller, 1862a)
= Penaeus sculptilis Heller, 1862a: 528. [Java]
Parapenaeopsis Alcock, 1901
= Parapeneopsis Alcock, 1901 (type species Penaeus styliferus H. Milne Edwards, 1837 [in H. Milne
Edwards, 1834-1840], by original designation, gender feminine; name emended under the plenary
powers of the ICZN to Parapenaeopsis and placed in that emended spelling on the Ofcial List of
Generic Names in Zoology in Opinion 864 in 1969)
Parapenaeopsis acclivirostris Alcock, 1905: 522, 530. [Persian Gulf; in Palk Strait; at Madras; of the Vizaga-
patam and Ganjam coasts]
Parapenaeopsis aroaensis Hall, 1962
= Parapenopsis aroaensis Hall, 1962: 27; Figs 109-109a. [20 mls NNE of Aroa Islands, 320'N 10039'E,
51 m]
Parapenaeopsis balli Burkenroad, 1934a: 64; Fig. 17. [Acajutla, El Salvador, 1336'N 8950'W]
Parapenaeopsis gracillima Nobili, 1903b: 4; Fig. 1. [Buntal, Borneo]
Parapenaeopsis longirostris Chandra & Bhatacharya, 2004: 23; Fig. 1. [Ongaria Ghat, Baleshwar, Orissa]
Parapenaeopsis nana Alcock, 1905: 522, 529. [Of Ganjam and Orissa coasts up to 68 fms; at Madras]
Parapenaeopsis stylifera (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840])
= Penaeus styliferus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 418. [les environs de
Bombay]
= Parapeneopsis stylifera var. coromandelica Alcock, 1906: 37; Plate 7, fgs 20, 20a-c. [Madras, Coroman-
del Coast; lectotype designation by Ravindranath, 1989]
= Parapenaeopsis stylifera var. cochinensis M.J. George, 1974: 421; Fig. 1b, d. [unavailable name under
Art. 15.2]
Parapenaeus Smith, 1885b
= Parapenus Smith, 1885b (type species Penaeus longirostris Lucas, 1846, by original designation,
gender masculine)
= Neopenopsis Bouvier, 1905a (type species Neopenopsis paradoxus Bouvier, 1905a, by original des-
ignation and monotypy, gender feminine)
Parapenaeus americanus Rathbun, 1901: 102; Plate 2. [Mayaguez Harbour, 220-225 fms]
Parapenaeus australiensis Dall, 1957: 179; Fig. 15A-F. [Of Newcastle, New South Wales, Australia]
= Parapenaeus australiensis forma nodosus Crosnier, 1986a: 313, 339. [types not indicated; unavailable
under Art. 15.2]
Parapenaeus cayrei Crosnier, 2005: 262; Figs 3, 4A-C. [BORDAU 2, stn CP 1541, les Tonga, 2115'S
17514'W, 319-333 m]
Parapenaeus fssuroides erythraeus Crosnier, 1986a: 329; Figs 5i, 6k, m, 8c-d. [Pola stn 117, Mer Rouge,
2016'N 3733'E, 638 m]
Parapenaeus fssuroides fssuroides Crosnier, 1986a: 321; Figs 5e-f, 6d-f, 7c, 8a. [Hong Kong, 1922.5'N
11407.5'E, 700-795 m]
Parapenaeus fssuroides indicus Crosnier, 1986a: 325; Figs 5g, h, 6g-j, l, 7d, f-g, 8b. [Vauban stn CH 80,
Madagascar, 2502.7'S 4705.8'E, 65-70 m]
Parapenaeus fssurus (Spence Bate, 1881)
= Penaeus fssurus Spence Bate, 1881: 180. [south of New Guinea, 50 fms]
= Penaeus fssurus Spence Bate, 1888: 263; Plate 36, fg. 1. [Challenger stns 190, 856'S 1365'E, 49 fms;
204A, 1243'S 1229'E, of Tablas Island, 100 fms; 204B, 1246'S 12210'E, of Tablas Island, 115 fms;
209, 1014'N 12354'E, of Zebu, 95-100 fms]
Parapenaeus investigatoris Alcock & Anderson, 1899: 279. [Investigator stns 233 (Andaman Sea, 1317'15''N
9310'25''E), 185 fms; 235 (Andaman Sea, 1413'N 9340'E), 370-419 fms; 166 (Bay of Bengal,
1334'55''N 8032'12''E), 133 fms]
Parapenaeus kensleyi Crosnier, 2005: 258; Figs 1-2. [HALIPRO I, stn CP 853, Nouvelle-Caldonie, 2145'S
16637'E, 241-250 m]
Parapenaeus lanceolatus Kubo, 1949a: 405; Figs 4C, 7D', 21C, 28G-I, 61C, C', 75C-I, 78I, 143C, G, 144. [Heta,
Shizuoka Prefecture and Kumano-nada, of Owase, Mie Prefecture, Japan, 300-350 m]
Parapenaeus longipes Alcock, 1905: 520, 525. [of Malabar coast (Mangalore), 21-26 fms; of Orissa and
Ganjan coasts, 20-68 fms; of Vizagapatam coast, 7,5-23 fms; S coast of Ceylon, 20 fms]
= Parapenaeus longipes forma denticulatus Crosnier, 1986a: 311, 312, 347. [Indonsie et Philippines,
exact type locality not indicated, see Crosnier, 1986a; unavailable under Art. 15.2]
Parapenaeus longirostris (Lucas, 1846)
= Peneus Cocco Prestandrea, 1833: 6. [Mari di Messina; suppressed under the plenary powers for the
purposes of the Principle of Priority but not those of the Principle of Homonymy in Opinion 611 in
1961]
= Penaeus longirostris Lucas, 1846: 46; Plate 4, fg. 6. [Algerie, Alger, aux environs du cap Matifou]
= Penus bocagei Johnson, 1863: 255. [mouth of the Tagus]
= Penaeus lividus Filhol, 1885b: Plate 1. [Type locality not indicated, 500-600 m]
Parapenaeus murrayi Ramadan, 1938: 74; Fig. 15a-c. [John Murray Expedition stn 105 (36'12''N
3913'12''E), Zanzibar area, 238-293 m]
Parapenaeus perezfarfantae Crosnier, 1986a: 339; Figs 12a-b, 13a-b, 14a. [Albatross expedition stn 5183,
Philippines, 1032'48"N 12226'E, 176 m]
Parapenaeus politus (Smith, 1881)
= Penaeus politus Smith, 1881: 444. [Fish Hawk stn 878 (3955'N 7054'W), 142 fms, of the eastern end
of Long Island, known as the Block Island]
= Neopenopsis paradoxus Bouvier, 1905a: 747. [dans la mer des Antilles par 84 et 91 brasses de pro-
fondeur]
Parapenaeus ruberoculatus Hall, 1962: 31; Figs 115-115b. [30 Miles NNE of Tanjong Baram, 504'N 11348'E,
183 m]
Parapenaeus sextuberculatus Kubo, 1949a: 403 (partim); Figs 7F', 21B, 28D-F, 61A-A', 75B, H, 78H, 143B, F
(nec Fig. 47R = P. fssuroides fssuroides Crosnier, 1986a). [Heta and Maisaka, Shizuoka Prefecture,
Japan, circa 330 m]
Pelagopenaeus Prez Farfante & Kensley, 1997
= Pelagopenaeus Prez Farfante & Kensley, 1997 (type species Penaeus balboae Faxon, 1893, designated
by Prez Farfante & Kensley, 1997, gender masculine)
Pelagopenaeus balboae (Faxon, 1893)
= Penaeus balboae Faxon, 1893: 211. [Albatross stn 3371 (of Isla del Coco, Costa Rica, 526'20"N
8655'00"W), 770 fms]
= Penaeus meridionalis Lenz & Strunck, 1914: 298, Plate 15, fgs 1-15. [Deutsche Sdpolar-Expedition,
3021'S 142'W, 10 m]
Penaeopsis Spence Bate, 1881
= Penaeopsis Spence Bate, 1881 (type species Penaeopsis serratus Spence Bate, 1881, designated by
Bouvier, 1905b, gender feminine)
Penaeopsis balssi Ivanov & Hassan, 1976b: 1; Figs 1-2. [of central Mozambique, 2526'S (not 2326'S as
stated), 3331'E, 410 m]
Penaeopsis challengeri De Man, 1911a: 76. [nomen novum for Penaeus serratus Spence Bate, 1881]
= Penaeus serratus Spence Bate, 1881: 182. [Challenger stn 173, of Matuku, Fii Islands, 1909'35"S
17941'50"E, 315 fms, lectotype designation by Prez Farfante, 1980a]
Penaeopsis eduardoi Prez Farfante, 1977a: 172; Figs 1-4. [Albatross stn 5116, Balayan Bay, Luzon Island,
Philippines, 1341'00"N 12047'05"E, 200 fms]
Penaeopsis jerryi Prez Farfante, 1979: 209; Figs 1-3. [John Murray Expedition stn 16, of Berbera, Somalia,
Gulf of Aden, 1029'48"N 4501'48"E, 186 m]
Penaeopsis mclaughlinae Crosnier, 2006: 334; Figs 2-5. [CHALCAL 2 stn CC 1, Nouvelle-Caldonie,
2454.96'S 16821.91'E, 500 m]
Penaeopsis rectacuta (Spence Bate, 1881)
= Penaeus rectacutus Spence Bate, 1881: 180. [among the Philippine Islands, about 100 fms; according
to Prez Farfante, 1980a: Challenger stn 209, between Bohol and Cebu, Philippines, 1014'N 12354'E,
95 fms]
Penaeopsis serrata Spence Bate, 1881
= Penaeopsis serratus Spence Bate, 1881: 183. [Gulf of Mexico; according to Prez Farfante, 1980a:
Blake stn 275, of Barbados, Gulf of Mexico, 1258'33"N 5936'45"W, 218 fms; name placed on the
Ofcial List of Specifc Names in Zoology in Opinion 864 in 1969]
= Parapenaeus megalops Smith, 1885b: 172. [Albatross stns 2125, S of Curaao, 1143'N 6909'30"W, 208
fms; 2143, Golfo de Urab, 930'45"N 7625'30"W, 155 fms]
= Artemesia talismani Bouvier, 1905b: 982. [le Talisman captura en quantit assez grande au large des
ctes du Maroc et du Sahara; Talisman stn 72, of Guerguerat, Western Sahara, 2541'N 1556'W (of
Greenwich) 1816'W (of Paris) on label accompanying specimen), 410 m, see Prez Farfante, 1980a]
= Penaeopsis serratus var. antillensis A. Milne-Edwards & Bouvier, 1909: 226; Plate 3, fg. 10; Plate 4,
fg. 5. [Blake stn 148, of St Kits, 208 fms]
Penaeus Fabricius, 1798
= Penaeus Fabricius, 1798 (type species Penaeus monodon Fabricius, 1798, designated by Latreille,
1810, gender masculine)
Penaeus esculentus Haswell, 1879: 38. [Port Jackson, New South Wales, Australia]
Penaeus monodon Fabricius, 1798: 408. [Bay of Batavia [=Jakarta], Indonesia; neotype designation by
Holthuis, 1949a]
= Penaeus carinatus Dana, 1852a: 27. [Singapore]
= Penaeus tahitensis Heller, 1862a: 528. [Taiti]
= Penaeus semisulcatus exsulcatus Hilgendorf, 1879: 843. [Quelimane, Mozambique]
= Penaeus coeruleus Stebbing, 1905: 77; Plates 21-21bis. [Nahoon River on E coast near East London,
South Africa]
? = Penaeus durbani Stebbing, 1917a: 442; Plate 22. [Durban Bay]
= Penaeus bubulus Kubo, 1949a: 296; Figs 1G, 7B, 15U-B', 24K-M, 37, 49C, 53, 58D, 67N-Q, 73F, L, 77P,
113, 114. [Hatazawa, Chiba Prefecture: Miya; Aichi Prefecture; Tainan, Formosa; Palau; Batavia;
Menado, Celebes]
Penaeus semisulcatus De Haan, 1844 [in De Haan, 1833-1850]: Plate 46, fg. 1. [Japan]
? = Penaeus Manilensis de Proc, 1822: 134. [Manille]
= Penaeus ashiaka Kishinouye, 1900: 14; Plate 3; Plate 7, fg. 4. [southern half of our coast]
= Penaeus semisulcatus paucidentatus Parisi, 1919: 65; Plate 5, fg. 5. [Misaki, Baia di Sagami]
= Penaeus monodon manillensis Villaluz & Arriola, 1938: 39; Plate 3, fg. 1. [Manila Bay, Philippines]
Protrachypene Burkenroad, 1934a
= Protrachypene Burkenroad, 1934a (type species Protrachypene precipua Burkenroad, 1934a, by
monotypy, gender feminine)
Protrachypene precipua Burkenroad, 1934a: 43. [Bella Vista Beach, Panama City, Panama]
Rimapenaeus Prez Farfante & Kensley, 1997
= Rimapenaeus Prez Farfante & Kensley, 1997 (type species Trachypenaeus faoe Obarrio, 1954, by
Rimapenaeus byrdi (Burkenroad, 1934a)
= Trachypeneus (Trachysalambria) byrdi Burkenroad, 1934a: 51; Fig. 13. [Panama City Market, Panama]
Rimapenaeus constrictus (Stimpson, 1871)
= Penaeus constrictus Stimpson, 1871: 135. [Beaufort, N.C., 4 fms; Charleston Harbour]
= Penaeopsis agassizii Bouvier, 1905a: 748. [provient de Sombrero]
Rimapenaeus faoe (Obarrio, 1954)
= Trachypenaeus faoe Obarrio, 1954: 3. [Golfo de Panam, en el Pacfco]
Rimapenaeus fuscina (Prez Farfante, 1971b)
= Trachypenaeus fuscina Prez Farfante, 1971b: 637; Figs 1-2, 3A, 4A-F, 5A, 6. [Of Cocodrillo Chiapas,
Pacifc coast of Mxico, 22 m]
Rimapenaeus pacifcus (Burkenroad, 1934a)
= Trachypeneus (Trachysalambria) similis pacifcus Burkenroad, 1934a: 50. [Pearl Islands, Gulf of Pana-
ma, Panama, 829'40"N 7852'30"W, 19-24 fms]
Rimapenaeus similis (Smith, 1885b)
= Parapenaeus constrictus similis Smith, 1885b: 175. [Albatross stn 2121, Gulf of Paria, 1037'40"N
6142'40"W, 31 fms]
Tanypenaeus Prez Farfante, 1972
= Tanypenaeus Prez Farfante, 1972 (type species Tanypenaeus caribaeus Prez Farfante, 1972, by orig-
inal designation, gender masculine)
Tanypenaeus caribaeus Prez Farfante, 1972: 187. [Pillsbury stn 788, of Punta Faro, Colombia, 1108'N
7447'W, 155-157 m]
Trachypenaeopsis Burkenroad, 1934a
= Trachypeneopsis Burkenroad, 1934a (type species Metapenaeus mobilispinis Rathbun, 1915, by origi-
= Trachypenaeopsis [emendment of Trachypeneopsis Burkenroad, 1934a, under the plenary powers of
the ICZN in Opinion 684 in 1969]
Trachypenaeopsis mobilispinis (Rathbun, 1915)
= Metapenaeus mobilispinis Rathbun, 1915: 117. [Cave Round Bay, Saba, Netherlands Antilles, 7 m]
Trachypenaeopsis minicoyensis Thomas, 1972: 117; Fig. 1. [western lagoon of Minicoy island, Laccadive
Archipelago, 3 m]
Trachypenaeopsis richtersii (Miers, 1884a)
= Penus richtersii Miers, 1884a: 564; Plate 52, fg. A. [Cerf Island, 10 fms]
Trachypenaeus Alcock, 1901
= Trachypeneus Alcock, 1901 (type species Penaeus anchoralis Spence Bate, 1881, by original designa-
= Trachypenaeus [emendment of Trachypeneus Alcock, 1901, under the plenary powers of the ICZN in
Opinion 864 in 1969]
Trachypenaeus anchoralis (Spence Bate, 1881)
= Penaeus anchoralis Spence Bate, 1881: 181. [south of New Guinea, 28 fms]
= Penaeus anchoralis Spence Bate, 1888: 258; Plate 35, fg. 1. [Challenger stns 188, 959'S 13942'E,
Arafura Sea, south of Papua, 28 fms; 190, 856'S 1365'E, Arafura Sea, south of Papua, 49 fms; of
Yokohama, 5-20 fms; between stns 190-191, near the Arrou Islands in the Arafura Sea]
Trachysalambria Burkenroad, 1934a
= Trachypeneus (Trachysalambria) Burkenroad, 1934a (type species Penaeus curvirostris Stimpson,
1860a, by original designation, gender feminine)
Trachysalambria albicoma (Hayashi & Toriyama, 1980)
= Trachypenaeus albicomus Hayashi & Toriyama, 1980: 69; Figs 1-2. [Japan, Tosa Bay]
Trachysalambria aspera (Alcock, 1905)
= Trachypenaeus asper Alcock, 1905: 523, 531. [fully described and illustrated in Alcock, 1906; type
locality, Ganjam coast, 20-35 fms, according to Alcock, 1906]
Trachysalambria brevisuturae (Burkenroad, 1934a)
= Trachypeneus (Trachysalambria) brevisuturae Burkenroad, 1934a: 55; Fig. 14. [Acajutla, El Salvador]
Trachysalambria curvirostris (Stimpson, 1860a)
= Penaeus curvirostris Stimpson, 1860a: 44. [Tosa Bay, Kochi, Japan, 3330'N 13330'E, 35 m; neotype
designation by Sakaji & Hayashi, 2003]
= Metapenaeus palaestinensis Steinitz, 1932: 161; Figs 1-3. [aus der Bucht von Haifa]
Trachysalambria fulva (Dall, 1957)
= Trachypenaeus fulvus Dall, 1957: 206; Fig. 23A-G. [Townsville, Queensland, Australia]
= Trachypeneus unicus Hall, 1961: 102. [S of Singapore, Malaysia, 35 fms]
Trachysalambria longipes (Paulson, 1875)
= Penaeus longipes Paulson, 1875: 125; Plate 19, fgs 1-1a. [Red Sea]
Trachysalambria malaiana (Balss, 1933)
= Trachypenaeus curvirostris malaiana Balss, 1933: 234. [Gier 16 Exp. 2, Sdl. Sumatra, 642'S 103E;
Gier 2 Exp. 3, Bai von Batavia; Gier 12 Exp. 4, Javasee vor Borneo, 342'S 11430'E; Gier 4, vor Toeban
(Nordjava); Gier 9 Exp. 20, nrdl. Sumatra, 120'S 10443'E; Gier 17 Exp. 4, westlich Borneo, 136'S
10946'E; Gier 6 Exp. 21, Bangkastrasse, 23'S 10548,5'E; Gier 4 Exp. 20, Javasee, 651'S 11256'E;
Gier 4 Exp. 11, vor Pekalongan (Nordjava)]
Trachysalambria nansei Sakaji & Hayashi, 2003: 162; Figs 7-9. [Japan, Tosa Bay, of Kochi, 3330'N, 13330'E,
75 m]
Trachysalambria starobogatovi (Ivanov & Hassan, 1976a)
= Trachypenaeus starobogatovi Ivanov & Hassan, 1976a: 1300; Figs 2, 3a. [Western Indian Ocean,
1903'S 3629'E, 25 m]
Trachysalambria villaluzi (Muthu & Motoh, 1979b)
= Trachypenaeus villaluzi Muthu & Motoh, 1979b: 57; Figs 1-2. [Tigbauan, Philippines, 7 m]
Xiphopenaeus Smith, 1869
= Xiphopeneus Smith, 1869 (type species Xiphopeneus hartii Smith, 1869, by monotypy, gender mas-
culine)
= Xiphopenaeus [emendment of Xiphopeneus Smith, 1869, under the plenary powers of the ICZN in
Opinion 864 in 1969]
Xiphopenaeus kroyeri (Heller, 1862b)
= Penaeus Kroyeri Heller, 1862b: 425; Plate 2, fg. 51. [Rio Janeiro]
= Xiphopeneus Hartii Smith, 1869: 27, 40; Plate 1, fg. 1. [Caravelas, Estado da Bahia, Brazil]
= Xiphopeneus Riveti Bouvier, 1907: 113; Fig. 1. [achete sur le march de Pata]
Family SICYONIIDAE Ortmann, 1898
Sicyonia H. Milne Edwards, 1830
= Sicyonia H. Milne Edwards, 1830 (type species Sicyonia sculpta H. Milne Edwards, 1830, designat-
ed by Desmarest, 1858, gender feminine)
= Ruvulus De Natale, 1850 (type species Sicyonia sculpta H. Milne Edwards, 1830, by monotypy,
gender masculine)
= Synhimantites Boeck, 1864 (type species Synhimantites typicus Boeck, 1864, by monotypy, gender
masculine)
= Eusicyonia Stebbing, 1914b (substitute name for Sicyonia H. Milne Edwards, 1830, gender femi-
nine)
Sicyonia abathophila Crosnier, 2003: 321; Fig. 98. [Albatross, Philippines Expedition, 1908-1909, Camp
Overton, Iligan Bay, Mindanao, Philippines, 815,24'N 12407,18'E, 2-4 m]
Sicyonia adunca Crosnier, 2003: 265; Figs 42-44, 107F. [BATHUS 2, stn DW 749, au large de la Nouvelle-
Caldonie, 2233,4'S 16626'E, 233-258 m]
Sicyonia afnis Faxon, 1893: 209. [Albatross stns 3367, of Isla del Coco, Costa Rica, 531'30"N 8652'30"W,
183 m; 3369, of Isla del Coco, Costa Rica, 532'45"N 8655'20"W, 95 m; 3378, W of Isla de Malpelo,
Colombia, 358'20"N 8136'00"W, 205 m; 3379, W of Isla de Malpelo, Colombia, 359'40"N 8135'00"W,
95 m]
Sicyonia aliafnis (Burkenroad, 1934a)
= Eusicyonia aliafnis Burkenroad, 1934a: 92; Fig. 24. [Pawnee, Pacifc coast of southern Mexico,
1448'40"N 9254'40"W, 19-30 fms]
Sicyonia altirostrum Crosnier, 2003: 297; Figs 73-75, 109G. [MUSORSTOM 6, stn CP 464, les Loyaut,
2102,3'S 16731,6'E, 430 m]
Sicyonia australiensis Hanamura & Wadley, 1998: 701; Figs 1-4. [Southern Surveyor, cruise SS6/96, stn 183,
SE coast of Australia, 3718.7'S, 15016.7'E, 123 m]
Sicyonia benthophila De Man, 1907a: 143. [Siboga stn 253, near Kai Islands, Indonesia, 304 m]
Sicyonia bispinosa (De Haan, 1844 [in De Haan, 1833-1850])
= Hippolyte bispinosus De Haan, 1844 [in De Haan, 1833-1850]: Plate 45, fg. 9. [Japan]
Sicyonia brevirostris Stimpson, 1871: 132. [S Florida coast]
Sicyonia burkenroadi Cobb, 1971: 104; Figs 1-2. [Oregon stn 1083, Gulf of Mexico of Port Isabel, Texas,
2613'N 9645'W, 42 m]
Sicyonia carinata (Brnnich, 1768)
= Cancer carinatus Brnnich, 1768: 102. [Mari Adriatico, Dalmatinis Skilla, in fnu maris juxta Salo-
nas]
= Cancer pulchellus Herbst, 1796 [in Herbst, 1791-1796]: 175; Plate 43, fg. 3. [das Veterland ist mir
unbekannt]
= Sicyonia sculpta H. Milne Edwards, 1830: 340; Plate 9, fgs 1-8. [Baie de Naples]
= Sicyonia foresti Rossignol, 1962: 145; 2 unnumbered Figs [Ile du Prince, 27 m]
Sicyonia curvirostris Balss, 1913b: 235. [Fukuura, Sagami Bay, Japon, 150 m]
Sicyonia dejouanneti Crosnier, 2003: 293; Figs 70-72, 108G, H, 109A-F. [MUSORSTOM 4, stn CP 194, Nou-
velle-Caldonie, 1852,8'S 16321,7'E, 545 m]
Sicyonia disdorsalis (Burkenroad, 1934a)
= Eusicyonia disdorsalis Burkenroad, 1934a: 96; Figs 25, 36. [Pawnee, Pearl Islands (Archipelago de las
Perlas), Gulf of Panama, 829'40"N 7852'30"W, 19-24 fms; Golfo de Panam; W coast of Central
America]
Sicyonia disedwardsi (Burkenroad, 1934a)
= Eusicyonia disedwardi Burkenroad, 1934a: 86; Figs 23, 29, 34. [Pawnee, Conception Bay, Lower Cali-
fornia]
Sicyonia disparri (Burkenroad, 1934a)
= Eusicyonia disparri Burkenroad, 1934a: 83; Fig. 27. [Pawnee, Bahia San Luis Gonzaga, Lower Cali-
fornia, Mexico]
Sicyonia dorsalis Kingsley, 1878a: 97. [Fort Jeferson, Florida]
Sicyonia fallax De Man, 1907a: 141. [Siboga stn 105, Sulu Sea, Indonesia, 608'N 12119'E, 275 m]
Sicyonia furcata Miers, 1878: 310; Plate 17, fg. 4. [Sulu Archipelago, Philippines]
= Sicyonia formosa Chan & Yu, 1985: 99; Fig. 1; Plates 5A, B, 6E-F. [Taiwan, 100 m]
Sicyonia galeata Holthuis, 1952a: 84. [Mbizi Expedition stns A.S.182, Angola, Luanda, 13 miles WSW of
Cape Ledo, 947'S 1311'E, 35 m; A.S.170, Congo, 10 miles W of Pointe Noire, 448'S 1141'E, 49 m]
Sicyonia infexa (Kubo, 1949a)
= Eusicyonia infexa Kubo, 1949a: 458; Figs 8O, 48D, 77C, I, 79G, 159, 160A-F. [Of Atsumi, Aichi
Prefecture, Japan, ca. 300 m; and Kumanonada, of Owase, Mie Prefecture, Japan, ca. 400 m]
Sicyonia ingentis (Burkenroad, 1938) (Fig. 4)
= Eusicyonia ingentis Burkenroad, 1938: 88; Figs 31-34. [Zaca stn 127D-1, of E coast of Cedros Island
(Isla Cedros), 2805'N 11509'W, Baja California, Mexico]
Sicyonia japonica Balss, 1914b
= Sicyonia lancifer var. japonica Balss, 1914b: 16; Fig. 9. [Dzushi, 50-100 m; Misaki, 200-300 m; Fukuu-
ra; Yagoshima, 150 m; zwischen Ito und Hatsushima, 150 m]
Sicyonia komai Crosnier, 2003: 327; Figs 103-104. [Thailande, cte ouest, Ao Tang Knen, Phuket, zone in-
tertidale]
Sicyonia laevigata Stimpson, 1871: 131. [Charleston]
= Sicyonia sculpta var. americana De Man, 1907b: 450. [Of Bahia]
Sicyonia laevis Spence Bate, 1881: 173. [north of New Guinea, 150 fms; fully described and illustrated in
Spence Bate, 1888; type locality according to Spence Bate, 1888: Challenger stn 219, north of New
Guinea, 154'S 14639'40"E, 150 fms]
= Eusicyonia nebulosa Kubo, 1949a: 454; Figs 8N, 48C, 77B, H, 79K, 156G, 157. [Of Heta, Shizuoka
Prefecture, Japan, ca. 350 m]
Sicyonia lancifer (Olivier, 1811)
= Palmon lancifer Olivier, 1811: 664. [CORINDON stn 295, Indonesia, Makassar, Strait, 126,5'S,
11702,1'E, 51-54 m; neotype selection by Crosnier, 2003; emendation to S. lancifera by Holthuis,
1980b is an unjustifed emendation]
= Hippolyte cristatus De Haan, 1844 [in De Haan, 1833-1850]: Plate 45, fg. 10. [Japan; lectotype des-
ignation by Yamaguchi & Baba, 1993]
Fig. 4. Sicyonia ingentis (Burkenroad, 1938). Photo by Sammy De Grave.
Sicyonia longicauda Rathbun, 1906: 908; Plate 20, fg. 6 (partim, nec specimens of stn 4002 = S. truncata
Kubo, 1949; see Crosnier, 2003). [Albatross stn 3865, Pailolo Channel, Hawaii Islands, 2109'20"N
15635'10"W, 468-517 m]
Sicyonia longicornis Crosnier, 2003: 276; Figs 55-56. [KARUBAR, stn CP 20, les Kai, Indonsie, 515'S
13259'E, 769-809 m]
Sicyonia martini Prez Farfante & Boothe, 1981: 424; Figs 1-4. [Pillsbury, stn 556, SW of Punta Ana Maria,
Golfo de Panama, 750'30"N 7849'00"W, 58 m]
Sicyonia metavitulans Crosnier, 2003: 304; Figs 79-81. [Port Jackson, cte est de lAustralie]
Sicyonia mixta Burkenroad, 1946: 3; Figs 1-4. [St. Joseph (probably San Jos, Lower California), Swedish
Eugenie Expedition #818]
Sicyonia nasica Burukovsky, 1990: 193, 211; Figs 2A(1-6), 6. [2504'S 9728'W, 267-280 m]
Sicyonia ocellata Stimpson, 1860a: 43. [Portu Hong Kong; in mari Sinensi quoque, lat. bor. 24; ad prof.
20 org]
= Sicyonia ommanneyi Hall, 1961: 110; Plate 21, fgs 26-27. [Malaysia, S of Singapore, 45 fms]
Sicyonia olgae Prez Farfante, 1980b: 775; Figs 1-3. [Oregon stn 2277, Of Parimaribo, Suriname, 0637'N
5536'W, 35 m]
Sicyonia parafallax Crosnier, 1995: 193; Figs 3-4. [Mer Rouge, dtroit de Bab el Mandeb, 1243,7'N
4315,0'E, 228 m]
Sicyonia parajaponica Crosnier, 2003: 226; Figs 10-12; 106E-F. [MUSORSTOM 1, stn 7, Philippines, dans
louest de Luon, 1401,0'N, 12020,0'E, 185-200 m]
Sicyonia parri (Burkenroad, 1934a)
= Eusicyonia parri Burkenroad, 1934a: 80; Fig. 22. [Crooked Island, Bahamas]
Sicyonia parvula (De Haan, 1844 [in De Haan, 1833-1850])
= Hippolyte parvulus De Haan, 1844 [in De Haan, 1833-1850]: Plate 45, fg. 6. [Japan]
Sicyonia penicillata Lockington, 1878a: 164. [Bolinas Bay, Lower California, 14 fms; Angeles Bay, Gulf of
California]
Sicyonia picta Faxon, 1893: 210. [Albatross stns 3387 (of Golfo de Panam, 740'00"N 7917'50"W, 127 fms);
3355 (of Punta Mariato, Panam, 712'20"N 8055'00"W, 182 fms]
Sicyonia rectirostris De Man, 1907a: 141. [Siboga stn 139, Sanana bay, E coast of Sula Besi, Indonesia, 22 m]
Sicyonia robusta Crosnier, 2003: 323; Figs 99-102, 110C, D. [CORAIL 2, stn DW 166, les Chesterfeld,
1941,49'S 15825,24'E, 56 m]
Sicyonia rocroi Crosnier, 2003: 280; Figs 60-62. [Mutu One, Hatutaa, les Marquises, 751,7'S 14030,6'W,
416-460 m]
Sicyonia rotunda Crosnier, 2003: 290; Figs 68-69, 108E-F. [SMIB 5, stn DW 84, Nouvelle-Caldonie,
2220,8'S 16843,1'E, 290 m]
Sicyonia stimpsoni Bouvier, 1905a: 748. [Blake stn 273, of Barbados, 1303'05"N 5936'18"W, 103 fms; lec-
totype designation by Holthuis, 1959a]
Sicyonia taiwanensis Crosnier, 2003: 282; Figs 63-64, 108B. [au large de Tai-Chi, Taiwan]
Sicyonia trispinosa De Man, 1907a: 142. [Siboga stn 37, Pulau Sailus ketjil, Paternoster Islands, Indonesia,
27 m]
Sicyonia truncata (Kubo, 1949a)
= Eusicyonia truncata Kubo, 1949a: 456; Figs 8M, 48F, 77A, G, 79J, 156E, 158. [Kumanonada, of
Owase, Mie Prefecture, Japan, 350 m]
Sicyonia typica (Boeck, 1864)
= Synhimantites typicus Boeck, 1864: 189. [Norge]
= Palmon carinatus Olivier, 1811: 667; nec Cancer carinatus Brnnich, 1768. [il se trouve sur les ctes
de la Nouvelle-Hollande]
= Sicyonia Edwardsii Miers, 1881a: 367. [nomen novum for Palmon carinatus Olivier, 1811]
Sicyonia vitulans (Kubo, 1949a)
= Eusicyonia vitulans Kubo, 1949a: 448; Figs 8Q, 48G, 77F, L, 79I, 154, 156F. [Kumanonada, of Owase,
Mie Prefecture, Japan, ca. 350 m]
Sicyonia wheeleri Gurney, 1943: 1; Figs 1-13. [Bermuda; description of adult in Burkenroad, 1946 as Sicyo-
nia wheeleri sp. nov.]
Family SOLENOCERIDAE Wood-Mason in Wood-Mason & Alcock, 1891a
Cryptopenaeus de Freitas, 1979
= Cryptopenaeus de Freitas, 1979 (type species Cryptopenaeus catherinae de Freitas, 1979, by original
designation, gender masculine)
= Crassipenaeus Liu & Zhong, 1983 (type species Crassipenaeus sinensis Liu & Zhong, 1983, by origi-
nal designation and monotypy, gender masculine)
Cryptopenaeus brevirostris Hayashi in Baba, Hayashi & Toriyama, 1986: 39; Plate 1; fg. 17. [Bungo Strait,
Japan, 360-380 m]
Cryptopenaeus catherinae de Freitas, 1979: 125; Fig. 1A-I. [Of Cape Santa Maria, southern Mozambique,
2606'S 3308'E, 350 m]
Cryptopenaeus clevai Crosnier, 1985: 26; Figs 1a-b, 2a-e, 3a. [CORINDON IV stn IV 1 (0328.3'N 12824.8'E
0330.0'N 12823.0'E), 400-300 m]
Cryptopenaeus crosnieri Prez Farfante & Kensley, 1985: 281; Figs 1-2. [R/V Kapala stn 78-05-07, NE of
North Solitary Island, New South Wales, Australia, 2947-49'S 15341'E, 234 m]
Cryptopenaeus sinensis (Liu & Zhong, 1983)
= Crassipenaeus sinensis Liu & Zhong, 1983: 171; Fig. 1. [North-Eastern Guangdong near Shantou
(Swatow), 261 m]
Gordonella Tirmizi, 1960
= Gordonella Tirmizi, 1960 (type species Gordonella polyarthra Tirmizi, 1960, by monotypy, gender
feminine)
Gordonella kensleyi Crosnier, 1988a: 597; Figs 2e, 12b, 15h-i, 16g. [BIOCAL stn CP 23, Nouvelle-Caldonie,
2246'N 16620'E, 2040 m]
Gordonella paravillosa Crosnier, 1988a: 589; Figs 2d, 3c, 12a, 13-14, 15a-e, 16a-f. [Cidaris I stn 18-1, cte est
de lAustralie, 1719.58'S 14747.61'E, 1147-1132 m]
Gordonella villosa (Alcock & Anderson, 1894)
= Haliporus villosus Alcock & Anderson, 1894: 146. [Investigator stns 121 Laccadive Sea, 1435'15"N,
1140 fms; 127, Laccadive Sea, of the Island of Minnikoy, 1200 fms]
= Gordonella polyarthra Tirmizi, 1960: 373, Figs 86-96. [John Murray Expedition stn 135, southern
Arabian Sea, 447'42"N 7235'36"E, 2727 m]
Hadropenaeus Prez Farfante, 1977b
= Hadropenaeus Prez Farfante, 1977b (type species Hymenopenaeus modestus Smith, 1885b, by origi-
Hadropenaeus afnis (Bouvier, 1906d)
= Haliporus afnis Bouvier, 1906d: 4. [capturs par le Talisman aux les du Cap Vert, sur des fonds de
100 et de 410 mtres; un spcimen provient des Barbades o il fut pris par le Blake 82 brasses de
profondeur." = Talisman stn 116, of Cape Verde Islands, 1653'N 2510'W, 410-460 m; uncited Talis-
man stn, of Cape Verde Islands, 100 m; Blake stn 273, of Barbados, 1303'05"N 5936'18"W, 188 m
according to Prez Farfante, 1977b]
Hadropenaeus lucasii (Spence Bate, 1881)
= Solenocera Lucasii Spence Bate, 1881: 185. [south of New Guinea, about 130 fms; according to
Spence Bate, 1888 (as Philonicus lucasii): Challenger stn 192, of the Ki Islands, south of Papua,
549'15"S 13214'15"E, 140 fms]
= Haliporus malhaensis Borradaile, 1910: 258; Plate 16, fg. 2. [of Saya de Malha, Indian Ocean, 145
fms]
Hadropenaeus modestus (Smith, 1885b)
= Hymenopenaeus modestus Smith, 1885b: 183. [Fish Hawk stn 1047, of Bethany Beach, Delaware,
3831'N 7321'W, 156 fms]
Hadropenaeus spinicaudatus Liu & Zhong, 1983: 174; Fig. 2. [Eastern Guangdong, of shore (1959.9'N
11324.0'E), 140 m]
Haliporoides Stebbing, 1914b
= Haliporoides Stebbing, 1914b (type species Haliporoides triarthrus Stebbing, 1914b, by monotypy,
gender masculine)
= Parahaliporus Kubo, 1949a (type species Haliporus sibogae De Man, 1907a, by original designation,
gender masculine)
Haliporoides cristatus Kensley, Tranter & Grifn, 1987: 265; Figs 1, 2, 5G-L. [NE of Point Danger, Queens-
land, Australia]
Haliporoides diomedeae (Faxon, 1893)
= Peneopsis diomedeae Faxon, 1893: 212. [Albatross stns 3384 (of Golfo de Panam, 731'30"N
7914'00"W, 458 fms); 3395 (SW of Golfo de Panam, 730'36"N 7839'00"W, 730 fms); 3394 (SE of
Golfo de Panam, 721'N 7935'W, 511 fms); 3393 (of Punta Mala, Panama, 715'N 7936'W, 1020
fms); 3353 (of Punta Mariato, Panama, 706'15"N 8034'00"W, 695 fms); 3358 (S of Pennsula de
Azuero, 630'N 8144'W, 555 fms)]
Haliporoides sibogae australiensis Kensley, Tranter & Grifn, 1987: 269; Figs 3, 4, 5A-F. [E of Broken Bay,
New South Wales, Australia]
Haliporoides sibogae madagascariensis Crosnier, 1978: 102; Figs 36b, 37a-e, 38a-c, 39b, 40b, 41b, 42b. [Vauban
CH 5, Madagascar, 1244.8'S 4810.6'E, 570-563 m]
Haliporoides sibogae sibogae (De Man, 1907a)
= Haliporus sibogae De Man, 1907a: 138. [Siboga stns 38, 735.4'S 11728.6'E, 521 m; 74, 53.5'S 1190'E,
450 m; 212, 554.5'S 12019.2'E, 462 m; 256, 526.6'S 13232.5'E, 397 m]
Haliporoides triarthrus triarthrus Stebbing, 1914b
= Haliporoides triarthrus Stebbing, 1914b: 21. [East London NW. 1/2N., 18 miles, (Cape Colony), 250-
300 fms]
Haliporoides triarthrus vniroi (Crosnier, 1978)
= Hymenopenaeus triarthrus vniroi Crosnier, 1978: 103; Figs 36c, 38e. [Van Gogh stn 264, Mozambique,
2528'S 3331'E, 410 m]
Haliporus Spence Bate, 1881
= Haliporus Spence Bate, 1881 (type species Haliporus curvirostris Spence Bate, 1881, designated by
Haliporus curvirostris Spence Bate, 1881: 185. [mid Pacifc, 2375 fms]
Haliporus taprobanensis Alcock & Anderson, 1899: 280. [Investigator stn 219, Gulf of Manaar and Cape
Comorin, southern India, 550 fms)]
= Hymenopenaeus kannemeyeri Kensley, 1977: 16, 27; Fig. 7. [Meiring Naude stn 38, of South Africa,
2821.9'S 3234.6'E, 775-825 m]
Haliporus thetis Faxon, 1893: 214. [Albatross stn 3413 (of the Galapagos Islands, 234'00"N 9206'00"W,
1360 fms)]
Hymenopenaeus Smith, 1882
= Hymenopenaeus Smith, 1882 (type species Hymenopenaeus debilis Smith, 1882, by monotypy, gender
masculine)
Hymenopenaeus aphoticus Burkenroad, 1936a: 112; Figs 62, 65-67. [Pawnee stn 54, Turks Island Passage,
2115'40"N 7117'06"W, 900-945 fms]
Hymenopenaeus chacei Crosnier & Forest, 1969: 545; Figs 1, 2a-b. [Ombango stn 406, au large du plateau
continental du Gabon, 835'S 1251'E, 545-555 m]
Hymenopenaeus debilis Smith, 1882: 91; Plate 15, fgs 6-11; Plate 16, fgs 1-3. [Blake stns 317, SE of Savannah
Beach, Georgia, 3157'00"N 7818'35"W, 333 fms; 323, SE of Cape Fear, North Carolina, 3319'00"N
7612'30"W, 457 fms; 326, E of Cape Fear, North Carolina, 3342'15"N 7600'50"W, 464 fms]
= Hymenopenaeus debilis var. africanus Bouvier, 1908a: 83. [Type locality not indicated; according to
Crosnier & Forest, 1973: Talisman stn 21, 3346'N 902'W, 1319 m]
Hymenopenaeus doris (Faxon, 1893)
= Haliporus doris Faxon, 1893: 214. [Albatross stns 3414 (of Cabo Velas, Costa Rica, 1014'N 9628'W,
2232 fms); 3415 (S of Punta Maldonado, Guerrero, Mexico, 1446'N 9840'W, 1879 fms)]
Hymenopenaeus equalis (Spence Bate, 1888)
= Haliporus equalis Spence Bate, 1888: 285; Plate 41, fg. 1. [Challenger stn 200, 647'N 12228'E, be-
tween the Philippine Islands and Borneo, 250 fms]
Hymenopenaeus fallax Crosnier & Dall, 2004: 10; Figs 6-8. [Albatross stn 4106, Kaiwi Channel, 335-350 fms]
Hymenopenaeus fatahi Ramadan, 1938: 60; Fig. 8. [John Murray Expedition stn 54 (2158'00''N 6219'42''E
to 2158'36''N 6221'24''E), South Arabian coast, 1046 m]
Hymenopenaeus furici Crosnier, 1978: 127; Figs 39f, 40f, 42f, 43d, 46f-h. [Vauban CH 133, Madagascar,
1302'S 4802'E, 1000-1525 m]
Hymenopenaeus halli Bruce, 1966a: 216; Figs 1-2. [Cape St. Mary Cruise 1/64, stn 26, South China Sea,
1922.5'N 11407.5'E to 1922.0'N 11411.0'E, 400-435 fms]
Hymenopenaeus laevis (Spence Bate, 1881)
= Haliporus laevis Spence Bate, 1881: 185. [mid Atlantic, 2500 fms]
= Hymenopenaeus microps Smith, 1884: 413; Plate 10, fg. 1. [Albatross stns 2076, E of Georges Bank, Mas-
sachusets, 4113'00"N 6000'50"W, 906 fms; 2037, of New Jersey, 3850'00"N 6923'30"W, 1731 fms]
= Haliporus androgynus Bouvier, 1906e: 253. [Talisman stns 101, entre Dakar et la Praya, 1638'N
2044'W, 3200 m; 102, entre Dakar et la Praya, 1548'N 2023'W, 3655 m]
Hymenopenaeus methalli Crosnier & Dall, 2004: 17; Figs 11-13, 14B, C. [MUSORSTOM 5, stn 384, 1942.40'S
15850.80'E, 756-772 m]
Hymenopenaeus neptunus (Spence Bate, 1881)
= Haliporus neptunus Spence Bate, 1881: 185. [among the Celebes Islands, about 600 fms; according
to Spence Bate, 1888: Challenger stns 191, of the Arrou Islands, 541'S 1344'30"E, 800 fms; 196, near
the Philippines, 048'30"S 12658'30"E, 825 fms]
Hymenopenaeus nereus (Faxon, 1893)
= Haliporus nereus Faxon, 1893: 213. [Albatross stns 3366 (S of Cabo Blanco, Costa Rica, 530'N
8645'W, 1067 fms); 3353 (S of Morro de Puercos, Panama, 706'15"N 8034'00"W, 695 fms); 3382 (of
Pen de Azuero, Panama, 621'N 8041'W, 1793 fms); 3413 (NW of Galapagos Islands, Ecuador,
234'N 9206'W, 1360 fms); 3398 (NW of Punta Galera, Ecuador, 107'N 8021'W, 1573 fms); 3399
(NW of Punta Galera, Ecuador, 107'N 8104'W, 1740 fms); 3400 (E of Galapagos Islands, Ecuador,
0036'N 8646'W, 1322 fms); 3407 (Galapagos Islands, Ecuador, 0004'00"S 9024'30"W, 885 fms)]
Hymenopenaeus obliquirostris (Spence Bate, 1881)
= Haliporus obliquirostris Spence Bate, 1881: 186. [of Kermadec Island; according to Spence Bate,
1888: Challenger stn 170, of the Kermadec Islands, 2955'S 17814'W, 520 fms]
Hymenopenaeus propinquus (De Man, 1907a)
= Haliporus propinquus De Man, 1907a: 140. [Siboga stns 80, Pulu Kaniungan Ketjil, Indonesia, 11 m;
178, 240'S 12837.5'E, 835 m; 316, 719.4'S 11649.5'E, 538 m]
Hymenopenaeus sewelli Ramadan, 1938: 58; Figs 7a-e. [John Murray Expedition stn 156 (444'30''N
7246'00''E to 441'12''N 7242'48''E), Maldive area, 1828 m]
Hymenopenaeus tuerkayi Crosnier, 1995: 189; Figs 1-2. [Golfe dAden central, 1256,7'N 4747,0'E-1255,9'N
4747,4'E, 2276-2282 m]
Mesopenaeus Prez Farfante, 1977b
= Mesopenaeus Prez Farfante, 1977b (type species Parartemesia tropicalis Bouvier, 1905a, by original
Mesopenaeus brucei Crosnier, 1986b: 20; Figs 1a-f, 2a-b. [Soela Cruise 184 stn 55, Australie, 1741.2'S
11842.5'E, 354-360 m]
Mesopenaeus mariae Prez Farfante & Ivanov, 1982: 303; Figs 1-7. [Professor Mesystsev haul 135, SE of Saya
de Malha Bank, Indian Ocean, 1127'36"S 6137'36"E, 158-160 m]
Mesopenaeus tropicalis (Bouvier, 1905a)
= Parartemesia tropicalis Bouvier, 1905a: 748. [de la mer des Antilles o elle a t prise par le Blake par
des profondeurs de 80 175 brasses]
= Solenocera weymouthi Lindner & Anderson, 1941: 181; Fig. 1a-e. [Pelican stn 137, 2928'N 8730'W,
46 fms]
Pleoticus Spence Bate, 1888
= Philonicus Spence Bate, 1888 (type species Philonicus mlleri Spence Bate, 1888, designated by
= Pleoticus Spence Bate, 1888 (nomen novum for Philonicus Spence Bate, 1888, a junior homonym of
Philonicus Loew, 1849 (Diptera))
= Parartemesia Bouvier, 1905a (type species Parartemesia carinata Bouvier, 1905a, designated by Fowler,
1912, gender feminine)
= Faxonia Bouvier, 1905b (type species Penaeopsis [sic] ocularis Faxon, 1895, designated by Fowler,
Pleoticus muelleri (Spence Bate, 1888)
= Philonicus mlleri Spence Bate, 1888: 275; Plate 39, fgs 1-2. [Challenger stn 321, of Montevideo,
Uruguay, 3502'N 5515'W, 13 fms]
= Parartemesia carinata Bouvier, 1905a: 748. [capture par le Hassler.. ..au large de Montevideo, par
7 et 44 brasses de profondeur.; of mouth Ro de la Plata, 3542'S 5620'W, 44 fms (80 m); see Prez
Farfante, 1977b]
Pleoticus robustus (Smith, 1885b)
= Hymenopenaeus robustus Smith, 1885b: 180. [Albatross stn 2125, S of Curaao, 1143'00"N 6909'30"W,
208 fms]
= Peneopsis ocularis Faxon, 1895: 187. [I have examined with some care a specimen in this Museum
[Museum of Comparative Zoology, Harvard] from the Blake collection, labelled Peneopsis ocula-
ris by A. Milne-Edwards]
Pleoticus steindachneri (Balss, 1914c)
= Haliporus Steindachneri Balss, 1914c: 135. [Pola stns 9, 2321'N 3737'E, 791 m; 20, 2320'N 3620'E,
780 m; 47, 2341'N 389'E, 610 m; 59, 2543'N 3610'E, 780 m; 61, 2435'N 3651'E, 828 m; 66, 268'N
3527'E, 1168 m; 107, 2027'N 3818'51"E, 748 m; 109, 2119'N 3739'E, 890 m; 110, 217'N 3728'E,
635 m; 114, 1938'N 3755'E, 535 m; 117, 2016'N 3733'E, 638 m; 120, 1921'N 3829'E, 560 m; 121,
1851'N 395'E, 690 m; 143, 177'N 3955'E, 212 m; 156, 2251'N 382'E, 712 m; 170, 272'N 3517'E,
690 m; 178, 2619'N 3424'E, 720 m; full description and list of syntypes in Balss, 1915]
Solenocera Lucas, 1849
= Solenocera Lucas, 1849 (type species Solenocera Philippii Lucas, 1849, by monotypy, gender femi-
nine; name placed on the Ofcial List of Generic Names in Zoology in Opinion 611 in 1961)
= Parasolenocera Wood-Mason in Wood-Mason & Alcock, 1891a (type species Parasolenocera annec-
tens Wood-Mason in Wood-Mason & Alcock, 1891a, by monotypy, gender feminine)
= Transolenocera Burkenroad, 1934b (type species Solenocera (Parasolenocera) maldivensis Borradaile,
1910, by monotypy, gender feminine)
Solenocera acuminata Prez Farfante & Bullis, 1973: 8; Figs 1c, 2, 4-6, 18, 19. [Oregon stn 2022, N of Rivire
Organabo, French Guiana, 0715'N 5335'W, 210 m]
Solenocera africana Stebbing, 1917b: 32; Plate 93A. [Sebastian Bluf NW. 3/4W., 8 miles, South Africa, 34
fms]
? = Solenocera membranacea capensis Heegaard, 1966: 32; Figs 15-21, 23. [Discovery stns 100B, 3320'S-
3346'S 1508'E 1518'E, 5-0 m; 102, 2529'30"S 1833'40"E, 50-0 m; 260, 3306'30"S 1745'15"E, 100-0
m; 277, 144'00"S 838'00"E, 63 m; 89, 3405'15"S 1600'45"E, 50-0 m; 99A, 3320'00"S 1717'00"E, 5-0
m; 99E, 3311'00"S 1726'00"E, 5-0 m; 276, 554'00"S 1119'00"E, 150 m]
Solenocera agassizii Faxon, 1893: 211. [Albatross stns 3389 (of Panama, 716'45"N 7956'30"W, 210 fms);
3391 (of Panama, 733'40"N 7943'20"W, 153 fms)
Solenocera alfonso Prez Farfante, 1981: 631; Figs 1-5. [of Capitancillo Island, W of Leyte, Philippines]
= Solenocera alfonso forma inermis Crosnier, 1989: 56; Figs 3e-f, 5f-h, 6a, e. [many locations in the
Philippines, 175-200 m; unavailable under Art. 15.2]
Solenocera algoensis Barnard, 1947
= Solenocera algoense Barnard, 1947: 383. [E portion of Algoa Bay, South Africa, 50 fms]
= Solenocera ramadani Ivanov & Hassan, 1976c: 242; Figs 1-2. [Van Gogh stn 269, of E Africa, 2931'S
3442'E, 285-305 m]
Solenocera alticarinata Kubo, 1949a: 227; Figs 8W, 45E, 72P, V, 80F, 93, 94A-C, 100. [Tainan, Formosa]
Solenocera annectens (Wood-Mason in Wood-Mason & Alcock, 1891a)
= Parasolenocera annectens Wood-Mason in Wood-Mason & Alcock, 1891a: 276. [Investigator stn 116
(Andaman Sea, 1125'5"N 9247'6"E), 405 fms]
Solenocera atlantidis Burkenroad, 1939: 10; Figs 5-10. [Atlantic stn 2813, of Mobile Bay, Alabama, 2945'N
8811'W, 19 fms]
Solenocera australiana Prez Farfante & Grey, 1980: 422; Figs 1-7. [Apache cruise 21, haul 3, N of Groote
Eylandt, Gulf of Carpentaria, Northen Territory, Australia, 1334'S 13630'E, 22 m]
Solenocera barunajaya Crosnier, 1994c: 355; Figs 1a-c, 2a, 3a-c. [KARUBAR stn CP 83, Iles Tanimbar Is-
land, Indonsie, 923'S 13100'E, 285-297 m]
Solenocera bedokensis Hall, 1962: 13; Figs 78-78c. [Of Bedok Village, Singapore, 17 fms]
Solenocera bifurcata Dall, 1999: 574; Fig. 14A-F. [Of Cape Moreton, SE Queensland, Australia]
Solenocera burukovskyi Timofeev, 1993: 37; Fig. 1. [Gulf of Aden, 1219'4N 4421'7E, 470-475 m]
Solenocera choprai Nataraj, 1945: 91, Figs 1-4. [Arabian Sea, 1727'N 7141'E, 56-58 fms]
Solenocera comata Stebbing, 1915
= Solenocera comatum Stebbing, 1915: 67; Plates 13-14. [Of East London, E coast of South Africa]
= Solenocera novae-zealandiae Borradaile, 1916: 79; Fig. 1. [7 miles E of North Cape, of New Zealand,
70 fms]
= Solenocera brevipes Kubo, 1949a: 246; Figs 1S, 8X, 20N, 27F-H, 45D, 66I-J, 72Q, W, 80A, 98H-J, 99,
100. [Komanonada, of Owase, Mie Prefecture, Japan, about 300 m]
Solenocera crassicornis (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840])
? = Penaeus planicornis Fabricius, 1798: 409. [in Oceano Indico]
= Penaeus crassicornis H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 418. [ctes de lInde]
= Solenocera sinensis Y, 1937: 112; Figs 1-5. [Amoy, Tinghai]
= Solenocera indicus Nataraj, 1945: 94; Figs 5-8. [Cocanada Bay, Bay of Bengal]
= Solenocera subnuda Kubo, 1949a: 255; Figs 8T, 20O, 27I-J, 45C, 66C, D, 72M, S, 80D, 100, 102D-G,
103. [S coast of Borneo, Indonesia]
= Solenocera kuboi Hall, 1956: 69; Plate 9, Figs 2-3. [Of Tanjong Stapa, Malaysia, 6-7 fms]
Solenocera faxoni De Man, 1907a: 136. [Siboga stn 254, 540'S 13226'E, 310 m]
Solenocera forea Burkenroad, 1938: 64; Text-Figs 6-7. [Pearl Island, Gulf of Panama, 829'40"N 7852'30"W,
19-24 fms]
Solenocera geiskesi Holthuis, 1959a: 56; Figs 4-5. [Surinam, 20 miles of the coast, between mouths of
Nickerie and Coppename Rivers, 27 m]
Solenocera gurjanovae Starobogatov, 1972: 365; Plate 3, fg. 13. [Gulf of Tonkin, Vietnam, 50 m]
Solenocera halli Starobogatov, 1972: 412; Plate 2, fg. 7. [Andaman Sea, of India, 605.8' 602'N 9859'
9900.5'E]
Solenocera hextii Wood-Mason & Alcock, 1891b: 188. [Investigator stns 81 (24 miles SE of Gapalpur, Gan-
jam Coast, 89-93 fms); 96 (Bay of Bengal, 1830'N 8446'E, 98-102 fms), 89-102 fms; of Chitagon, 65
fms; of the Mahnadi Delta, 68 fms; of the Godvari Delta, 70 fms]
Solenocera koelbeli De Man, 1911a: 45, 48, 50, 51. [Japan]
= Solenocera depressa Kubo, 1949a: 237; Figs 8R, 27T-V, 45B, 66E-F, 72O, U, 80G, 96, 98A-D, 100.
[Komanonada, of Owase, Mie Prefecture, Japan]
= Solenocera vietnamensis Starobogatov, 1972: 363, 384; Plate 2, fg. 6a-c. [Pelamida, stn 39, Tonkin
Gulf, Vietnam]
Solenocera maldivensis Borradaile, 1910
= Solenocera (Parasolenocera) maldivensis Borradaile, 1910: 258; Plate 16, fg. 1. [Funadu Velu, Miladu-
madulu Atoll, Maldive Islands]
Solenocera mascarensis Burukovsky, 1993a: 23; Fig. 1.3-1.7. [Sala-de-Malya Bank, 1111'1''S 6237'E, 2020-
2000 m]
Solenocera melantho De Man, 1907a: 137. [Siboga stns 302, 1017.9'S 12328.7'E, 216 m; 306, 827'S
12254.5'E, 247 m; 312, Saleh bay, N coast of Sumbawa, Indonesia, 819'S 11741'E, 274 m]
= Solenocera prominentis Kubo, 1949a: 231; Figs 8V, 14C, 16C, 20M, 27Q-S, 45A, 66G, H, 72F, L, 80E,
94D-G, 95, 100. [Komanonada, of Owase, Mie Prefecture, Japan, about 350 m]
Solenocera membranacea (Risso, 1816)
= Peneus membranaceus Risso, 1816: 98. [Prsident Thodore Tissier stn J.376, France, Ligurian Sea be-
tween Corsica and Nice, 4256.5'N 937.2'E, 310-415 m; neotype designation by Holthuis, 1961b]
= Peneus siphonoceros Philippi, 1840: 190; Plate 4, fg. 3. [Neapel]
= Solenocera philippii Lucas, 1849: 300. [baie dAlger]
= Penaeus distinctus De Haan, 1849 [in De Haan, 1833-1850]: 194. [Japan; erroneous locality, see
Yamaguchi & Baba, 1993]
Solenocera moosai Crosnier, 1985: 37; Figs 5a, 6a, 7c,d, h, i. [CORINDON II stn 273 (0156.0'S 11916.0'E),
120-200 m]
Solenocera mutator Burkenroad, 1938: 61; Text-Figs 2-5. [Templeton Crocker Expedition stn 150, from Gorda
Banks, tip of Lower California, 2301'30"N 10930"W, 40-100 fms]
Solenocera necopina Burkenroad, 1939: 7; Figs 1-4. [Atlantis stn 2377, of Mobile Bay, Alabama, 2916'N
8754'W, 125 fms]
Solenocera pectinata (Spence Bate, 1888)
= Philonicus pectinatus Spence Bate, 1888: 279; Plate 38. [Challenger stn 188, Arafura Sea, south of
Papua, 959'S 13942'E, 28 fms]
= Philonicus cervicalis Zehntner, 1894: 210; Plate 9, fg. 26. [Amboine]
Solenocera pectinulata Kubo, 1949a: 251; Figs 8S, 27A-B, 66 K-L, 72N-T, 83B, 101, 102A-C. [Komanonada,
of Owase, Mie Prefecture, Japan, about 350 m]
? = Solenocera utinomii Kubo, 1951: 263; Fig. 4. [Of Kii Peninsula, Japan, ca. 150 m]
Solenocera phuongi Starobogatov, 1972: 366; Plate 3, fg. 12a-b. [Pelamida stn 9, Tonkin Gulf, 113 m]
Solenocera rathbuni Ramadan, 1938: 57 (partim, nec Fig. 6 = S. algoensis Barnard, 1947). [Albatross stn 3987,
Kauai Island, Hawaii, 50-55 fms; lectotype designation by Crosnier, 1978] (Fig. 5)
Solenocera spinajugo Hall, 1961: 81; Plate 17, fgs 1-3. [N of the Malacca Strait, 120 miles NW of Penang,
41 fms]
Solenocera vioscai Burkenroad, 1934b: 65; Figs 1-4. [About 5 miles of Pass la Loutre, Louisiana, USA, 15
fms]
Solenocera waltairensis M.J. George & Muthu, 1970: 292; Figs 1-4. [Waltair, east coast of India, 1743'N
8323'E, 20 m]
Solenocera zarenkovi Starobogatov, 1972: 367, Plate 3, fg. 14a-b. [Orlik, stn 18, Tonkin Gulf]
Fig. 5. Solenocera rathbuni Ramadan, 1938. Photo by Tin-Yam Chan.
Superfamily SERGESTOIDEA Dana, 1852a
Family LUCIFERIDAE De Haan, 1849 [in De Haan, 1833-1850]
Lucifer J.V. Thompson, 1829 (Fig. 6)
= Lucifer J.V. Thompson, 1829 (type species Leucifer typus H. Milne Edwards, 1837 [in H. Milne Ed-
wards, 1834-1840], by subsequent indication under Article 68b in H. Milne Edwards, 1834-1840,
gender masculine)
= Leucifer H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840] [incorrect subsequent spelling
of Lucifer]
Lucifer chacei Bowman, 1967: 266; Figs 1-4. [Eniwetok Atoll, Marshall Islands, lagoon 4.8 km W of Parry
Island, 3 m]
Lucifer faxoni Borradaile, 1915a: 228, 230. [of Chesapeake Bay, U.S.A.; lectotype designation by Holthuis,
1959a]
= Lucifer afnis Borradaile, 1915a: 228, 230. [Floridastrom, J.N., 56, N.E. of Bermuda, roughly 35N
60W; lectotype designation by Holthuis, 1959a]
Lucifer hanseni Nobili, 1905a
= Lucifer Hanseni Nobili, 1905a: 395. [Djibouti]
= Lucifer inermis Borradaile, 1915a: 229. [Melbourne Harbour, Australia]
Lucifer intermedius Hansen, 1919: 57; Plate 4, fgs 8a-b; Plate 5, fgs 1a-g. [Siboga stns 7, 755'.5S 11426'E,
15 m and more; 35, 80'.3S 11659'E, 1310 m, surface; 37, Sailus ketjil, Paternoster-islands, 27 m and
less, surface; 66, bank between islands of Bahuluwang and Tambolungan, south of Saleyer, 8-10 m;
75, 457'.4S 1192'.8E, from 11 m to surface; 81, Pulu Sebangkatan, Borneo-bank, 34 m; 106, Anchor-
age of Kapul-island, Sulu-archipelago, 13 m; 109, Anchorage of Pulu Tongkil, Sulu-archipelago, 13
m; 117a, 115'N 12337'E; ?132, about 556'.7N 12625'E; 136, Ternate anchorage, 23 m - surface; 138,
Anchorage on the east coast of Kajoa-island, 66 m - surface; 141, 10'.4S 12725'.3E, from 1500 m;
144, Anchorage North of Salomakice-(Damar)island, 45 m; 157, 032'.9S 13014'.6E; 177a, 230'S
12928'E; 189a, 222'S 12646'E]
Lucifer orientalis Hansen, 1919: 55; Plate 4, fgs 7a-g. [Siboga stns 37, Sailus Ketjil, Paternoster-islands, 27
m and less; 66, bank between islands of Bahuluwang and Tambolungan, S of Saleyer, 8-10 m; 81,
Pulu Sebangkatan, Borneo-bank, 34 m; 104, Sulu-harbour, Sulu-island, 14 m; 106, anchorage of
Fig. 6. Lucifer spec. Photo by Arthur Anker.
Kapul-island, Sulu-archipelago, 13 m; 133, anchorage of Lirung, Salibabu-island, depth up to 36 m;
136, Ternate anchorage, 23 m - surface; 138, anchorage on the E coast of Kajoa-island, 66 m - surface;
140, Bay of Batjan, 13 m; 141, 10'.4S 12725'.3E, 1500 m - surface; 143, 14'.5S, 12752'.6E, 1000 m -
surface; 148, 017'.6S 12914'.5E, 1000 m - surface; 157, 032'.9S 13014'.6E, 45 m; 177a, 230'S
12928'E; 189a, 222'S 12646'E; 194-197, 153'.5-145'.3S 12639-1278'.3E; 203, 332'.5S 12415'.5E,
1500 m - surface; 220, anchorage of Pasir Pandjang, W coast of Binongka, 278 m - surface; 225, N.
279E, from Southpoint of South-Lucipara-island, 894 m; 243.430'.2S 12925'E, 1000 m - surface;
245.416'.5S 13015'.8E, surface; 276, 647'.5S 12840'.5 E, 750 m - surface; near stn 300, between
Timor and Lomblen]
Lucifer penicillifer Hansen, 1919: 59; Plate 5, fgs 2a-k. [Siboga stns 7, 755'.5S 11426'E, 25 m and more; 16,
Bay of Kankamaran, S coast of Kangeang, 22 m; 35, 80'.3S 11659'E, surface; 37, Sailus ketjil, Pater-
noster-islands, 27 m and less; 40, anchorage of Pulu Kawassang, Paternoster-islands, 12 m; 66, bank
between Bahuluwang and Tambolungan, S of Saleyer, 8-10 m; 81, Pulu Sebangkatan, Borneo-bank,
34 m; 93, Pulu Sanguisiapo, Tawi-Tawi-islands, Sulu-archipelago, 12 m; 96, SE side of Pearl-bank,
Sulu-archipelago; 98-99, 69'-67'.5N 12021'-12026'E; 99, 67'.5N 12026'E, 16-23 m - surface; 105,
68'N 12119'E, 275 m; 106, anchorage of Kapul-island, Sulu-archipelago, 13 m; 107, 61'.5N.
12128'E; 109, anchorage of Pulu Tongkil, Sulu-archipelago, 13 m; 112, 31'N 1222'E; 117
a
, 115'N
12337'E; 125, anchorage of Sawan, Siau-island, 27 m; 128, 427'N 12525'.7E, from 700 m to surface;
136, Ternate anchorage, 23 m; 138, anchorage on the E coast of Kajoa-island, 66 m; 140, Bay of Bat-
jan, 13 m; 141, 10'.4S 12725'.3E, from 1500 m to surface; 143, 14'.5S 12752'.6E, from 1000 m to
surface; 144, anchorage north of Salomakie- (Damar-) island, 45 m; 146, 036'S 12832'.7E, 512 m;
157, 032'.9S 13014'.6E, 45 m; 165, anchorage on NE side of Daram-island, E coast of Misool, 49 m;
168, anchorage N of Sabuda-island, 63 m; 174, Waru-bay, N coast of Ceram, 18 m; 177
a
, 230'S
12928'E; 184, anchorage of Kampong Kelang, S coast of Manipa-island, 36 m; 185, 320' 12722'.9E,
from 1536 m to surface; 189
a
, 222'S 12646'E; 194, 153' 5S 12639'E; 194-197, 153'.5-14S'.3S 12639'-
1278'.3E; 203, 332'5S 12415'.5E, from 1500 m to surface; 206, Buton-strait, surface; 220, anchorage
of Pasir Pandjang, W coast of Binongka, 278 m; 225, 5700 m N 279E from Southpoint of South-
Lucipara-island, 894; 243, 430'.2S 12925'E, from 1000 m to surface; 245, 416'.5S 13015'.8E; near stn
300, between Timor and Lomblen; 315, anchorage E of Sailus Besar, Paternoster-islands, surface]
Lucifer typus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]
= Leucifer typus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 469. [Type locality not
indicated]
= Leucifer Reynaudii H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 469. [trouv dans
lOcean indien]
= Lucifer acestra Dana, 1852b: 671. [Pacifc Ocean, 630'S 177E, near Shersons Island]
= Lucifer acicularis Dana, 1852b: 674. [Harbour of Rio de Janeiro]
= Lucifer Zybrantsii Philippi, 1857: 323. [im Atlantischen Ocean unter 25N.B. und 2250'W.L. von
Greenwich]
= Leucifer uracanthus Steindachner, 1861: 365. [E mari atlantico]
= Lucifer bonitensis Borradaile, 1915a: 228, 230. [toutes les mers]
= Lucifer clausi Borradaile, 1915a: 228, 230. [Messina]
= Lucifer batei Borradaile, 1915a: 228, 230. [Plankton-Expedition, Floridastrom, J.N. 62, Bermuda;
lectotype designation by Holthuis, 1959a]
Family SERGESTIDAE Dana, 1852a
Acetes H. Milne Edwards, 1830
= Acetes H. Milne Edwards, 1830 (type species Acetes indicus H. Milne Edwards, 1830, by original
Acetes americanus americanus Ortmann, 1893: 39; Plate 2, fg. 2. [Plankton-Expedition der Humbolt-Stif-
tung, Mndung des Tocantins JN 239 (0.6S 48.1W, 0 m), 240 (1.6S 49.2W, 0-35 m), 241 (1.6S
49.2W, 0-35 m), 243 (0.7S 48.2W, 0-13 m), Pl. 106 (0.7S 48.2W, 12 m) and 107 (1.6S 49.2W, 35 m)]
= Acetes brasiliensis Hansen, 1919: 45; Figs 1-7. [Brazil]
= Acetes americanus limonensis Burkenroad, 1934a: 99; Figs 37-38. [Mouth of the Sweetwater River,
Limon Bay, Canal Zone, Panama]
Acetes americanus carolinae Hansen, 1933
= Acetes carolinae Hansen, 1933: 31; Figs 1-8. [near the Sea Buoy of Beaufort Inlet, North Carolina]
Acetes binghami Burkenroad, 1934a: 101; Figs 39-40. [Bella Vista Beach, Panama City]
Acetes chinensis Hansen, 1919: 41; Plate 4, fg. 3a-b. [3310'N 12918'E, 40 fms; Formosa Channel, 25 fms]
Acetes erythraeus Nobili, 1905a: 393; Fig. 1. [Mer Rouge et Prim; Djibouti, dans la cavit dune Bero;
Abdelkader, prs de Massaouah, et Massaouah]
Acetes indicus H. Milne Edwards, 1830: 351; Plate 11, fgs 1-9. [Le Gange]
= Acetes spiniger Hansen, 1919: 35, 43; Plate 4, fg. 5. [Surabaya, Java, Indonesia; 10 miles of Ran-
goon; Indo-Chinese Sea]
Acetes intermedius Omori, 1975: 40; Figs 14, 16, 17. [Of Tungkiang, Taiwan]
Acetes japonicus Kishinouye, 1905: 167; 2 unnumbered Figs [the Bay of Ariake, the Bay of Kojima, of
Nagasu in Buzen, Japan; of Mokpho, Korea]
= Acetes dispar Hansen, 1919: 39; Plate 3, fgs 5a-f; Plate 4, fg. 1a. [Cheribon, Java, Indonesia; Lem
Ngob, Gulf of Siam]
= Acetes cochinensis Rao, 1970: 298; Figs 1-10. [Inshore sea of Cochin, southwest coast of India, 3.5-
11 m]
Acetes johni Nataraj, 1949: 139; Fig. 1. [Travancore coast, India]
Acetes marinus Omori, 1975: 49; Figs 8, 21, 25. [Rio Tocantins, near its junction with Rio Para, Brazil]
Acetes natalensis Barnard, 1955: 2, 43. [Durban Bay, South Africa]
Acetes paraguayensis Hansen, 1919: 46; Figs 8-14. [Lagoon at Rio Paraguay near its junction with Rio
Parana, Paraguay, outlet of Riacho del Oro in Rio de la Plata, Paraguay]
Acetes serrulatus (Kryer, 1859)
= Sergestes serrulatus Kryer, 1859: 268; Plate 4, fg. 12. [nordligste Kategat]
= Acetes insularis Kemp, 1917a: 54; Figs 1f-g, 2c, 3c, 4c, 5b, e, 7c. [Mouth of Rajang R., Sarawak, Bor-
neo]
Acetes sibogae sibogae Hansen, 1919: 38; Plate 3, fgs 4a-h. [Siboga stns 47, Bay of Bima, near south fort, 55
m; 323, Sangkapura roads, Bawean Island, 12 m]
= Acetes sibogalis Achuthankuty & M.J. George, 1973: 139; Figs 1-20. [Aroor region of the Cochin
backwaters]
= Acetes orientalis Achuthankuty & Ayyappan Nair, 1976: 233; Figs 1-19. [near fsh jety in Mandovi
estuary, Arabian Sea, Goa, India]
Acetes sibogae australis Colefax, 1940
= Acetes australis Colefax, 1940: 341, 345; Figs 1-19. [Australia: Homebush Bay, Port Jackson (Sydney
Harbour); Folly Point, Middle Harbour, Port Jackson; Clarence River, N coast of New South Wales;
Tuggerah Lake, 50 miles N of Port Jackson]
Acetes vulgaris Hansen, 1919: 35; Plate 3, fgs 2a-r. [Siboga stns 4, anchorage of Djangkar (Java), 742'S
11412.6'E, 9 m; 47, Bay of Bima, near south fort, 55 m; 311, Sapeh Bay, E coast of Sumbawa, 0-36 m]
Allosergestes Judkins & Kensley, 2008
= Allosergestes Judkins & Kensley, 2008 (type species Sergestes sargassi Ortmann, 1893, by original
Allosergestes index (Burkenroad, 1940)
= Sergestes (Sergestes) index Burkenroad, 1940: 41. [Dana stn 3630 II (New Zealand, Auckland, 3424'S
17842.5'E), 3000 m wire out)]
Allosergestes nudus (Illig, 1914)
= Sergestes nudus Illig, 1914: 366; Figs 26-28. [Atlantischen Ozean, 32S 8W, 1000 m]
Allosergestes oleseni (Vereshchaka, 2009) comb. nov.
= Sergestes oleseni Vereshchaka, 2009: 78; Figs 35, 38. [Dana stn 3602-2 (south western Pacifc Ocean,
between Fii and New Caledonia, 2000'S 17429'E, 600 m wire out)]
Allosergestes pectinatus (Sund, 1920)
= Sergestes pectinatus Sund, 1920: 24; Figs 42-43. [Michael Sars stns 23 (3532'N 77'W, 1215 m); 34
(2852'N 1416'W, 2170 m); 52 (3124'N 3447'W); 45 (2842'N 200'W); 49 (296'N 252'W); 51
(3120'N 357'W, 3886 m); 53 (3459'N 331'W, 2615-2865 m); 56 (3653'N 2917'W, 3239 m); 62
(3652'N 3955'W); 64 (3444'N 4752'W); 67 (4017'N 5039'W)]
Allosergestes pestafer (Burkenroad, 1937)
= Sergestes pestafer Burkenroad, 1937: 318; Figs 1-3. [Templeton Crocker Expedition stn 165 T-3, 145 miles
N of Clarion Island, Revillagigido Islands, Lower California, E Pacifc, 2036'N 11507'W, 500 m]
Allosergestes sargassi (Ortmann, 1893)
= Sergestes sargassi Ortmann, 1893: 34; Plate 3, fg. 1. [Plankton-Expedition der Humbolt-Stifung,
Floridastrom JN 50 (39.4N 57.8W, 0-200 m); Sargasso-See JN 94 (31.5N 45.6W, 0-400 m), 99
(31.7N 43.6W, 0-400 m), 102 (31.7N 42.7W, 0-400 m), 110 (30.3N 37.9W, 0-400 m), 113 (29.8N
36.8W, 0-400 m), 117 (28.3N 34.3W, 0-400 m), 127 (24.6N 31.0W, 0-400 m), 262 (23.7S 36.0W, 0
m), 263 (25.6S 34.9W, 0-400 m), Pl. 58 (25.1N 31.5W, 200 m); Nrdl. Aequatorialstrom JN 135
(18.9N 26.4W, 0-400 m), 260 (20.4N 37.8W, 0-400 m), Pl. 64 (16.1N 23.1W, 200 m); Sdl. Aequa-
torialstrom Pl. 76 (0.1S 15.2W, 200 m)]
Allosergestes verpus (Burkenroad, 1940)
= Sergestes (Sergestes) verpus Burkenroad, 1940: 40. [Dana stn 3739 IX (Celebes Sea, 320'N 12350'E),
2000 m wire out); according to Vereshchaka, 2009, the holotype label indicates that the specimen
was collected at Dana stn 3844-6 (Indo-West Pacifc, 1205'S 9645'E, 600 m wire out)]
Allosergestes vinogradovi (Vereshchaka, 2009) comb. nov.
= Sergestes vinogradovi Vereshchaka, 2009: 90; Figs 35, 44. [Dana stn 3902-2, Indo-West Pacifc, 605'N
9530'E, (600 m wire out)]
Deosergestes Judkins & Kensley, 2008
= Deosergestes Judkins & Kensley, 2008 (type species Sergestes curvatus Crosnier & Forest, 1973, by
Deosergestes coalitus (Burkenroad, 1940)
= Sergestes (Sergestes) coalitus Burkenroad, 1940: 39. [Dana stn 3737 I (Mindanao Sea, 723'N 12129'E,
1000 m wire out)]
= Sergestes (Sergestes) erectus Burkenroad, 1940: 38. [Dana stn 3579 I (Rarotonga, 2056'S 16003'W),
1000 m wire out)]
Deosergestes corniculum (Kryer, 1855)
= Sergestes corniculum Kryer, 1855: 30. [fra Atlanterhavets tropiske Deel, 4.5N 21.5W; according to
Kryer, 1859]
= Sergestes laciniatus Kryer, 1859: 274, 282; Plate 5, fg. 15. [fra Kategatet]
? = Pasiphae rubrogutata Filhol, 1885b: Plate 5. [Type locality not indicated]
= Sergestes longirostris Spence Bate, 1888: 415; Plate 75, fg. 3. [Mid Atlantic]
= Sergestes (Sergestes) curvatus Crosnier & Forest, 1973: 315; Figs 105i-k, 107c-d, f-g. [au large de
lAfrique du Sud, 3542'S 2440'E, 500 m]
Deosergestes disjunctus (Burkenroad, 1940)
= Sergestes (Sergestes) disjunctus Burkenroad, 1940: 38. [Dana stn 3630 IV (New Zealand, Auckland,
3424'S 17842.5'E), 1000 m wire out)]
Deosergestes henseni (Ortmann, 1893)
= Sergia henseni Ortmann, 1893: 38; Plate 3, fg. 3. (partim, see Crosnier & Forest, 1973). [Plankton-
Expedition der Humboldt-Stifung, Nrdl. Aequatorialstrom JN 146 (12.3N 22.3W); Vertikalnetz
0-400 m; Guineastrom JN 158 (7.5N 21.3W), circa 4000 m]
Deosergestes paraseminudus (Crosnier & Forest, 1973)
= Sergestes (Sergestes) paraseminudus Crosnier & Forest, 1973: 313; Figs 105d, 106c-d, f. [Ombango stn
309-GS 20 bis, 155'S 830'E, 0-350 m]
Deosergestes pediformis (Crosnier & Forest, 1973)
= Sergestes (Sergestes) pediformis Crosnier & Forest, 1973: 313; Figs 105e-h, 107a-b, e. [Ombango stn
302-GS 19, 447'S 1042'E, 0-725 m]
Deosergestes rubrogutatus (Wood-Mason in Wood-Mason & Alcock, 1891c)
= Sergestes rubrogutatus Wood-Mason in Wood-Mason & Alcock, 1891c: 354; Fig. 10. [Investigator
stns 107 (Laccadive Sea, 823'N 7547'E), 738 fms; 109 (S of C. Comorin, 741'N 7821'E), 738 fms; 110
(Bay of Bengal, 934'N 8543'15''E), 1997 fms; 117 (Bay of Bengal, 1158'N 8852'17''E), 1748 fms]
Deosergestes seminudus (Hansen, 1919)
= Sergestes seminudus Hansen, 1919: 18; Plate 1, fgs 7a-c; Plate 2, fgs 1a-f. [Siboga stns 66, Bank be-
tween islands of Bahuluwang and Tambolungan, S of Salayer, Indonesia, 8-10 m; 144, Anchorage N
of Salomakie (Damar-) Island, 45 m; 148, 017.6' S 12914.5'E, 0-1000 m; 157, 032.9'S 13014.6'E, 45
m; 167, Anchorage N of Sabuda Island, 63 m; 177a, 230'S 12928'E; 185, 320'S 12722.9'E, Manipa
Strait, 0-1536 m; 189a, 222'S 12646'E; 194-197, 155' 145.3'S 12639'- 1278.3'E; 203, 332.5'S
12415.5'E 0-1500 m; 223, 544.7'S 12627.3'E, surface; 230, 358'S 12820'E 0-2000 m; 276, 647.5'S
12840.5'E, 0-750 m]
= Sergestes nipponensis Yokoya, 1933: 13; text-fg. 4. [Sy-Maru stns 21, Japan, SE of Siwoya-zaki, 209
m; 25, Japan, E of Siwoya-zaki, 525 m; 253, Japan, Sagami Bay, 452 m]
Eusergestes Judkins & Kensley, 2008
= Eusergestes Judkins & Kensley, 2008 (type species Sergestes arcticus Kryer, 1855, by original desig-
Eusergestes antarcticus (Vereshchaka, 2009) comb. nov.
= Sergestes antarcticus Vereshchaka, 2009: 57; Figs 25, 26. [Dana stn 3975-2 (South Atlantic, Cape
Town, 3542'S 1837'E, 2500 m wire out)]
Eusergestes arcticus (Kryer, 1855)
= Nika Sinuolata Risso, 1816: 87. [Nice; status of name discussed by Holthuis, 1977a]
= Sergestes arcticus Kryer, 1855: 27. [fra Grnland, according to Kryer, 1859]
= Sergestes Rinkii Kryer, 1855: 33. [Atlanterhav p 58-59 n. Br., according to Kryer, 1859]
= Sergestes Meyeri Metzger, 1875: 302; Plate 6, fg. 7. [Kiel]
= Sergestes magnifcus Chun, 1888: 33; Plate 4, fgs 4-5. [Ischia, 800 m]
Eusergestes similis (Hansen, 1903)
= Sergestes similis Hansen, 1903: 60; Plate 11, fg. 6. [Challenger stn 232, of Japan, 3511'N 13928'E,
345 fms]
? = Sergestes nasidentatus Spence Bate, 1888: 398; Plate 72, fg. 2. [Pacifc Ocean between Valparaiso
and Juan Fernandez, 0-200 fms; Holthuis (pers. comm.)]
? = Sergestes laeviventralis Spence Bate, 1888: 425; Plate 67, fg. 3. [north of New Guinea]
? = Sergestes longicaudatus Stimpson, 1860a: 46. [Oceano Pacifco, lat. Bor. 40, long occ. 155]
? = Sergestes afnis Hansen, 1919: 7. [nomen nudum]
Neosergestes Judkins & Kensley, 2008
= Neosergestes Judkins & Kensley, 2008 (type species Sergestes edwardsii Kryer, 1855, by original
Neosergestes brevispinatus (Judkins, 1978)
= Sergestes brevispinatus Judkins, 1978: 13; Figs 2d, 5d-f,h,k, 6-8, 216. [Eastern tropical Pacifc, 1102'S
8141'W, ca. 200 m at night]
Neosergestes consobrinus (Milne, 1968)
= Sergestes consobrinus Milne, 1968: 26; Figs 5-9. [Brown Bear cruise 199, haul 227, 3344'N 12453'W,
120 m]
Neosergestes edwardsii (Kryer, 1855)
= Sergestes Edwardsii Kryer, 1855: 28. [nrheden af Linien, 3S to 10N, according to Kryer, 1859]
= Sergestes oculatus Kryer, 1855: 28. [tropiske Atlanterhav, 4.5N 21.5W, according to Kryer, 1859]
= Sergestes brachyorrhos Kryer, 1859: 272, 281; Plate 3, Figs 5a-f. [i Atlanterhavet, efer Angivelse paa
omtrent 30N 33W]
= Sergestes intermedius Spence Bate, 1888: 383. [Of Luzon, China Sea]
= Sergestes ventridentatus Spence Bate, 1888: 431. [N of the Sandwich Islands]
Neosergestes orientalis (Hansen, 1919)
= Sergestes orientalis Hansen, 1919: 22; Plate 2, fg. 2a-q. [Siboga stns 37, Sailus Ketjil, Paternoster Is-
lands, 0-27 m; 40, Anchorage of Pulu Kawassang, Paternoster Islands, 12 m; 66, Bank between is-
lands of Bahuluwang and Tambolungan, S of Salayer, Indonesia, 8-10 m; 96, SE side of Pearl bank,
Sulu Archipelago, 0-15 m; 106, anchorage of Kapul Island, Sulu Archipelago, 13 m; 118, 138'N
12428.2'E, 0-900 m; 125, anchorage of Sawan, Siau Island, 27 m; 128, 427'N 12525.7'E, 0-700 m;
131-133, about 556.7'N 12625'E; 141, 10.4'S 12725.3'E, 0-1500 m; 143, 14.5'S 12752.6'E, 0-1000 m;
144, Anchorage N of Salomakie (Damar-) Island, 45 m; 146, 036'S 12832.7'E, surface; 148, 017.6'
S 12914.5'E, 0-1000 m; 157, 032.9'S 13014.6'E, 45 m; 177a, 230'S 12928'E; 168, anchorage N of
Sabuda Island, surface; 172, Gisser, anchorage between this island and Ceram Island, 18 m; 177a,
230'S 12928'E, surface; 185, 320'S 12722.9'E, Manipa Strait, 0-1536 m; 189a, 222'S 12646'E;
194-197, 155'-145.3'S 12639'-1278.3'E; 194-197, 153.5'-145.3'S 12639'-1278.3'E; 203, 332.5'S
12415.5'E, 0-1500 m; 206 or 207, Buton Strait, surface; 220, anchorage of Pasir Pandjang, W coast of
Binongka, surface; 225, N. 279E from Southpoint of South Lucipara Island, 894 m; 230, 358'S
12820'E, 0-2000 m; 245, 416.5'S 13015.8'E, surface]
= Sergestes geminus Judkins, 1978: 25; Figs 2a-c, 7, 16f-j, 17, 18, 21a. [Eastern Tropical Pacifc, 448'N
8338'W, ca. 200 m at night]
= Sergestes gibbilobatus Judkins, 1978: 27; Figs 2g, 4c, 7, 19a-h, 20, 21a. [Equatorial Pacifc, 00'S
16542'W, 2550 m wire out, at night]
Neosergestes semissis (Burkenroad, 1940)
= Sergestes (Sergestes) semissis Burkenroad, 1940: 42. [Dana stn 3905 I (Bay of Bengal, 444'N 8805.5'E),
1000 m wire out]
Neosergestes tantillus (Burkenroad, 1940)
= Sergestes (Sergestes) tantillus Burkenroad, 1940: 42. [Dana stn 3556 II (Gulf of Panama, 252'N
8736'W), 2500 m wire out)]
Parasergestes Judkins & Kensley, 2008
= Parasergestes Judkins & Kensley, 2008 (type species Sergestes armatus Kryer, 1855, by original
Parasergestes armatus (Kryer, 1855)
= Sergestes armatus Kryer, 1855: 31. [737'n.Br. og N 22.5 v. Lgd., according to Kryer, 1859]
= Sergestes incertus Hansen, 1896: 962. [3450'S 430'W and 404'S 5320'W, surface]
= Sergestes extensus Hanamura, 1983: 64; Figs 7-8. [Kaiyo Maru stn MT 9-B, Seamount 350, of Baja
California, 23-05.4N 124-56.9W, 0-1236 m]
Parasergestes cylindricus (Vereshchaka, 2009) comb. nov.
= Sergestes cylindricus Vereshchaka, 2009: 96; Figs 47, 48. [Dana stn 3639-1, Southeast Pacifc, 3919'S
17918'E (300 m wire out)]
Parasergestes diapontius (Spence Bate, 1881)
= Sergestes diapontius Spence Bate, 1881: 194. [Atlantic]
= Sergestes penerinki Spence Bate, 1888: 418; Plate 76, fg. 3. [North Atlantic Ocean]
= Sergestes fermerinkii Spence Bate, 1888: 419; Plate 76, fg. 4. [Pacifc Ocean, 24S 148W]
= Sergestes semiarmis Spence Bate, 1888: 423; Plate 67, fg. 1. [Challenger stn 354, Mid North Atlantic
Ocean, 3241'N 366'W, surface]
Parasergestes halia (Faxon, 1893)
= Sergestes halia Faxon, 1893: 217. [Albatross stn 3388 (of Panama, 706'N 7948'W, surface to 400 fms)]
Parasergestes sirenkoi (Vereshchaka, 2009) comb. nov.
= Sergestes sirenkoi Vereshchaka, 2009: 103; Figs 51, 52. [Dana stn 3593-8, Southwest Pacifc, 1727'S
17933'E (300 m wire out)]
Parasergestes stimulator (Burkenroad, 1940)
= Sergestes (Sergestes) stimulator Burkenroad, 1940: 41. [Dana stn 3656 VIII (of Sydney, Australia,
3326'S 15702'E), 300 m wire out)]
Parasergestes vigilax (Stimpson, 1860a)
= Sergestes vigilax Stimpson, 1860a: 45. [Oceano Atlantico prope insulas Azores]
= Sergestes macrophthalmus Stimpson, 1860a: 46. [Oceano Pacifco, lat. bor. 27,5, long. orient. 138,5;
etiam lat. bor. 35, long. occ. 155]
= Sergestes parvidens Spence Bate, 1888: 409; Plate 74, fg. 3. [the tropical part of the Atlantic; Pacifc
Ocean, north of the Sandwich Islands; of Sydney and Wellington, Australia]
? = Sergestes spiniventralis Spence Bate, 1888: 426; Plate 67, fg. 5. [North Pacifc Ocean]
? = Sergia Clausi Knig, 1895: 10; Plate 1, fgs 1-7. [stliches Mitelmeer, 3612'N 2854'E, Ober-
fchenfang]
Peisos Burkenroad, 1945
= Peisos Burkenroad, 1945 (type species Peisos petrunkevitchi Burkenroad, 1945, by original designa-
Peisos petrunkevitchi Burkenroad, 1945: 554; Plates 1-2. [Montevideo, 5-6 fms]
Petalidium Spence Bate, 1881
= Petalidium Spence Bate, 1881 (type species Petalidium foliaceum Spence Bate, 1881, by monotypy,
gender neuter)
Petalidium foliaceum Spence Bate, 1881: 194. [taken in South Indian Ocean at a depth of about 2100 fms;
according to Spence Bate, 1888: Challenger stn 159, South of Australia, 4725'S 13022'E, 2150 fms]
Petalidium obesum (Kryer, 1855)
= Sergestes obesus Kryer, 1855: 31. [4.5N Br. i Atlanterhavet, according to Kryer, 1959]
= Sergestes sanguineus Chun, 1889: 538; Plate 3, fg. 1. [Canarischen Inseln]
Petalidium suspiriosum Burkenroad, 1937: 325; Text-Figs 1-6. [Templeton Crocker Expedition stn 165, 145
miles N of Clarion Island, Revillagigido Islands, Lower California, E Pacifc, 500 fms]
Sergestes H. Milne Edwards, 1830
= Sergestes H. Milne Edwards, 1830 (type species Sergestes atlanticus H. Milne Edwards, 1830, by
= Acheles Cocco, 1832 (type species Acheles arachnipodus Cocco, 1832, by monotypy, gender feminine)
Sergestes atlanticus H. Milne Edwards, 1830: 349; Plate 10, fgs 1-9. [lOcan atlantique, une grande
distance des ctes]
= Sergestes Frisii Kryer, 1855: 26. [Atlanterhav, 13N 27.5W and 20N 36W, according to Kryer,
1859]
= Sergestes ancylops Kryer, 1855: 32. [Atlanterhav 13N 27.5'W, according to Kryer, 1859]
? = Sergestes pacifcus Stimpson, 1860a: 45. [Oceano Pacifco, lat. bor. 27,5, long. orient. 138]
= Sergestes ovatoculus Spence Bate, 1888: 408; Plate 74, fg. 2. [N Atlantic Ocean]
Sergestes cornutus Kryer, 1855: 29. [Atlanterhavet omtrent 4.5N for Linien; indtil 10 nord for Linien og
undtil 8 syd for Linien, according to Kryer, 1859]
= Sergestes longispinus Spence Bate, 1888: 417; Plate 76, fg. 2. [Challenger stn 106, Mid Atlantic Ocean,
147'N 2426'W, within 40 fms of the surface]
Sergestes formosensis Yokoya & Shibata, 1965: 2; Fig. 2. [Philippine Sea, 2142'N 12305'E, 100 m]
Sergestes grandipes Yokoya & Shibata, 1965: 1; Fig. 1. [Philippine Sea, 1357'N 12529'E, 100 m]
Sergestes hamifer Alcock & Anderson, 1894: 148. [Investigator stn 126, Laccadive Sea, 849'0"N 7318'45"E,
1370 fms]
Sergestes latirostris Yokoya & Shibata, 1965: 3; Fig. 3. [Philippine Sea, 1303'N 12550'E, 40 m]
Sergia Stimpson, 1860a
= Sergia Stimpson, 1860a (type species Sergia remipes Stimpson, 1860a, by monotypy, gender femi-
nine; the type species was determined to be a late mastigopus stage by Burkenroad, 1945, who
identifed it with the group of species in what was then the subgenus Sergia)
Sergia bigemmea (Burkenroad, 1940)
= Sergestes (Sergia) bigemmeus Burkenroad, 1940: 49. [Dana stn 3570 VI (NE of Tahiti, 309'N
12609.3'E), 2150 m]
Sergia bisulcata (Wood-Mason & Alcock, 1891b)
= Sergestes bisulcatus Wood-Mason & Alcock, 1891b: 190. [Investigator stns 100 (Bay of Bengal,
1655'41"N, 8321'18"E), 840 fms; 105 (Arabian Sea, 1502'N, 7234'E), 740 fms]
Sergia burukovskii Vereshchaka, 2000: 121; Figs 31-33. [Dana stn 3980-1, South Atlantic, 2326'S, 0356'E,
1000 m wire out]
Sergia challengeri (Hansen, 1903)
= Sergestes challengeri Hansen, 1903: 61; Plate 12, fg. 2a-n. [Challenger stn 173, Western Pacifc, of
Matuku, Fii Islands, 199'35"S, 17941'50"E, 315 fms]
Sergia crosnieri Vereshchaka, 2000: 196; Figs 85-87. [Dana stn 3809-4, of Indonesian islands, 622'S,
10512'E, 50 m wire out]
Sergia erythraeensis Iwasaki & Couwelaar, 2001: 92; Figs 1-3. [Central Red Sea, 2123.89'N 3803.62'E, 150
m]
Sergia extenuata (Burkenroad, 1940)
= Sergestes (Sergia) extenuates Burkenroad, 1940: 45. [Dana stn 3999 II (Saint Helena, S Atlantic, 345'S
1000'E, 1000 m wire out); see discussion in Vereshchaka, 2000]
Sergia flicta (Burkenroad, 1940)
= Sergestes (Sergia) flictum Burkenroad, 1940: 52. [Dana stn 3549 IV (Gulf of Panama, 716'N 7930'W),
wire out 600 m]
Sergia foresti Kensley & Judkins, 2008: 151; Fig. 1. [Albatross stn 5241, Pujada Bay, Mindanao, 650'45"N
12614'38"E, 393 m]
Sergia fulgens (Hansen, 1919)
= Sergestes fulgens Hansen, 1919: 17; Plate 1, fg. 6a-g. [Siboga stn 312, Indonesia, Saleh Bay, Sum-
bawa, 0819'S 11741'E, 174 m]
Sergia gardineri (Kemp, 1913a)
= Sergestes gardineri Kemp, 1913a: 55; Plate 7, fgs 2-5. [S by E of Farquhar, 1027'S, 5117'E; NE of
Madagascar, between Providence and Alphonse Islands, 816'S, 5126'E; 5 miles of Desroches Atoll]
Sergia grandis (Sund, 1920)
= Sergestes grandis Sund, 1920: 16; Figs 22-26. [Michael Sars stns 34, (2852'N 1416'W, 2170 m); 49
(296'N, 252'W); 51 (3120'N 357'W, 3886 m); 52 (3124'N, 3447'W]
Sergia hansjacobi Vereshchaka, 1994: 91; Figs 23-24. [Dana stn 1198-2, 1743'N, 6456'W]
= Sergia hanseni Vereshchaka, 1994: 91. [nomen nudum]
Sergia inequalis (Burkenroad, 1940)
= Sergestes (Sergia) inequalis Burkenroad, 1940: 51. [Dana stn 3768 (N of NW New Guinea, 120'S
13842'E), 2500 m wire out)]
Sergia inoa (Faxon, 1893)
= Sergestes inous Faxon, 1893: 216. [Albatross stn 3380 (East Pacifc, of Malpelo Island, Colombia,
0403'N, 8131'W, 899 fms)]
Sergia japonica (Spence Bate, 1881)
= Sergestes japonicus Spence Bate, 1881: 194. [south of Japan; according to Spence Bate, 1888: Chal-
lenger stn 232, western Pacifc of southern coast of Japan, 3511'N, 13928'E, 345 fms]
= Sergestes mollis Smith, 1884: 419. [Albatross stns 2002, 3720'42"N 7417'36"W, 641 fms; 2018,
3712'22"N 7417'36"W, 788 fms; 2040, 3835'13"N 5816'00"W, 2226 fms; 2045, 4004'20"N 6843'50"W,
373 fms; 2051, 3941'00"N 6920'20"W, 1106 fms; 2083, 4026'40"N 6705'15"W, 959 fms; 2093,
3942'50"N 7101'20"W, 1000 fms; 2094, 3944'30"N 7104'00"W, 1022 fms; 2097, 3756'20"N
7057'30"W, 1917 fms; 2099, 2712'20"N 6939'00"W, 2949 fms; 2100, 3922'00"N 6834'30"W, 1628
fms; 2101, 3918'30"N 6824'00"W, 1686 fms; 2103, 3847'20"N 7237'00"W, 1091 fms; 2104, 3848'00"N
7240'30"W, 991 fms; 2105, 3750'00"N 7303'50"W, 1395 fms; 2110, 3512'10"N 7457'15"W, 516 fms;
2116, 3545'23"N 7431'25"W, 888 fms]
= Sergestes profundus Spence Bate, 1888: 428 (partim). [Challenger stn 300, southeastern Pacifc west
of Valparaiso, Chile, 3342'S, 7818'W, 1375 fms; lectotype designation by Hansen, 1903]
Sergia jeppeseni Vereshchaka, 2000: 179; Figs 67, 74-75; Plate 5F. [Dana stn 3943-1, Western Indian Ocean
of Mombasa, 530'S, 4040'E, 500 m wire out]
Sergia kensleyi Vereshchaka, 2000: 110; Figs 17, 24-25; Plate 4D. [Dana stn 3970-1, Western Indian Ocean
of Mozambique, 3409'S, 2738'E]
Sergia laminata (Burkenroad, 1940)
= Sergestes (Sergia) laminatus Burkenroad, 1940: 53. [Dana stn 3933 I (N of Madagascar, 1118'S
5003'E), 4000 m wire out]
= Sergestes (Sergia) guineensis Crosnier & Forest, 1973: 343; Fig. 118. [Ombango stn 394-8, 552'S
1000'E, 0-2500 m]
Sergia lucens (Hansen, 1922)
= Sergestes lucens Hansen, 1922: 12, 38. [Suruga Bay; see discussion in Vereshchaka, 2000]
Sergia manningorum Froglia & Gramito, 2000: 72; Figs 1-3. [Atlantis II, cruise 60 stn RH2281, of Congo,
1123'S 1055'E, 190-200 m]
Sergia maxima (Burkenroad, 1940)
= Sergestes (Sergia) maximus Burkenroad, 1940: 47. [Dana stn 3933 I (N of Madagascar, 1118'S
5003'E, 4000 m wire out)]
Sergia oksanae Vereshchaka, 2000: 182: Figs 73, 76-77; Plate 5E. [Dana stn 3736-3, Mindanao Sea, 917'N,
12358'E]
Sergia phorca (Faxon, 1893)
= Sergestes phorcus Faxon, 1893: 217. [Albatross stns 3382 (of Panama, 621'N 8041'W, 1793 fms);
3388 (of Panama, 706'N 7948'W, 1168 fms); 3401 (of Galapagos Islands, 050'S 8858'30"W, 395
fms); 3386 (of Panama, 733'12"N 7917'15"W, 242 fms); 3437 (Gulf of California, 2739'40"N
11100'30"W, 628 fms)]
Sergia plumea (Illig, 1927)
= Sergestes plumeus Illig, 1927: 295; Figs 30-32. [Valdivia stn 268, Indischer Ozean, sdlich von Ras
Hafun, 96'N, 5341'E, im Vertikalnetzfang von 1500 m Tiefe]
Sergia potens (Burkenroad, 1940)
= Sergestes (Sergia) potens Burkenroad, 1940: 48. [Dana stn 3975 VII (Agulhas Bank, South Africa,
3542'S 1837'E, 600 m wire out)]
Sergia prehensilis (Spence Bate, 1881)
= Sergestes prehensilis Spence Bate, 1881: 193. [of Japan, 500 fms; according to Spence Bate, 1888:
Challenger stn 236, of Japan, 3458'N, 13929'E, 775 fms]
= Sergestes gloriosus Stebbing, 1905: 84; Plates 22-23. [of Sandy Point, 800 fms]
= Sergestes fujiyamaensis Nakazawa, 1932: 32. [Type locality not indicated]
Sergia profunda (Spence Bate, 1888)
= Sergestes profundus Spence Bate, 1888: 428. [Challenger stn 137, 3559'S 134'E, 2550 fms; 300, W of
Valparaiso, 3342'S 7818'W, 1375 fms]
Sergia regalis (Gordon, 1939)
= Sergestes regalis Gordon, 1939: 498; Figs 1-4. [Discovery stn 81, South Atlantic, 3245'S, 847'W]
= Sergestes (Sergia) creber Burkenroad, 1940: 44. [Dana stn 3766 XVIII (N of NW new Guinea, 113'S
13842'E), 2900 m wire out]
Sergia remipes Stimpson, 1860a: 46. [Oceano Pacifco, lat bor. 27 log. orient. 138]
Sergia robusta (Smith, 1882)
= Sergestes robustus Smith, 1882: 97; Plate 16, fg. 5-8b. [Blake stn 328, 3428'25''N 7522'50''W, 1632
fms; Albatross stns 893 (3717'N 7321'W), 372 fms; 952 (3428'50"N 7522'50"W), 388 fms]
= Sergestes dissimilis Spence Bate, 1888: 437. [St. Vincent, Cape Verde Island, at the surface]
= Sergestes mediterraneus Hansen, 1896: 954. [Ragusa and Lesina, Adriatic Sea]
Sergia scintillans (Burkenroad, 1940)
= Sergestes (Sergia) scintillans Burkenroad, 1940: 43. [Dana stn 3622 I (SE of New Caledonia, 2554'S
17236.9'E, 300 m wire out)]
Sergia splendens (Sund, 1920)
= Sergestes splendens Sund, 1920: 14; Figs 16-18. [Michael Sars stns 29 (3510'N 755'W) 440 m wire
out; 42 (282'N 1417'W), 300 m wire out; 45 (2842'N 200'W), 100, 200, 300, 2000, & 3000 m wire
out; 49 (296'N 252'W), 3000 m wire out; 51 (3120'N 357'W), 200, 300 & 1000 m wire out; 52
(3124'N 3447'W), 100 m wire out; 53 (3459'N 331'W), 100, 300, 600 & 2600 m wire out; 56
(3653'N 2917'W), 200 & 300 m wire out; 62 (3652'N 3955'W), 2000 m wire out; 64 (3444'N
4752'W), 100 & 2000 m wire out; 66 (3930'N 4942'W), 1000 m wire out; 67 (4017'N 5039'W),
1200 & 2000 m wire out)]
= Sergestes Richardi Hansen, 1920: 482. [nomen novum for Sergestes splendens Sund, 1920, considered
a junior homonym of Sergestes splendens Hansen, 1919 by Hansen, 1920]
= Sergestes crassus Hansen, 1922: 98; Plate 5, fg. 4. [nomen novum for Sergestes splendens Sund, 1920,
considered a junior homonym of Sergestes splendens Hansen, 1919 by Hansen, 1922]
Sergia stellata (Burkenroad, 1940)
= Sergestes (Sergia) stellatus Burkenroad, 1940: 44. [Dana stn 3908 I (SE of Sri Lanka, 428'N 8213'E,
1000 m wire out)]
Sergia talismani (Barnard, 1947)
= Sergestes splendens Hansen, 1919: 18. [nomen nudum]
= Sergestes splendens Hansen, 1920: 480; nec Sund, 1920. [Talisman Expedition, stn 113, canal de Saint-
Vincent, 1652'N 2730 - 2731'W, 550-760 m]
= Sergestes talismani Barnard, 1947: 384. [nomen novum for Sergestes splendens Hansen, 1919 nec
Sund, 1920]
Sergia tenuiremis (Kryer, 1855)
= Sergestes tenuiremis Kryer, 1855: 30. [2N, 21W, according to Kryer, 1859]
= Sergestes Kryeri Spence Bate, 1881: 193. [of Kermadec Island, about 500 fms; according to Spence
Bate, 1888: Challenger stn 170, of the Kermadec Islands, 2955'S 17814'W, 520 fms]
= Sergestes junceus Spence Bate, 1888: 416; Plate 76, fg. 1. [South Pacifc Ocean]
= Sergestes longicollis Spence Bate, 1888: 421; Fig. 1. [Challenger near stn 131, South Atlantic Ocean,
2935'S 289'W; stn 295, South Pacifc Ocean, 387'S 944'W, 1500 fms]
= Sergestes tropicus Sund, 1920: 18; Figs 27, 28, 30-33. [Michael Sars stns 29 (3510'N 755'W); 45
(2842'N 200'W); 49 (296'N 252'W); 51 (3120'N 357'W, 3886 m); 52 (3124'N 3447'W); 53 (3459'N
331'W, 2615-2865 m); 56 (3653'N 2917'W, 3239 m); 64 (3444'N 4752'W)]
Sergia umitakae Hashizume & Omori, 1995: 72; Figs 1-4. [South of Sri Lanka]
Fig. 7. Sicyonella af. maldivensis Borradaile, 1910. Photo by Tin-Yam Chan.
Sergia vityazi Vereshchaka, 2000: 157; Figs 51, 57-58. [Dana stn 3601-1, southwestern Pacifc, 1821'S,
17821'E]
Sergia wolf Vereshchaka, 1994: 88; Figs 19-21. [Dana stn 1217-1, 1850'N, 7907'W]
Sicyonella Borradaile, 1910
= Aphareus Paulson, 1875 (type species Aphareus inermis Paulson, 1875, by monotypy, gender mas-
culine; invalid junior homonym of Aphareus Cuvier & Valenciennes, 1830 (Pisces))
= Sicyonella Borradaile, 1910 (type species Sicyonella maldivensis Borradaile, 1910, by monotypy, gen-
der feminine)
= Aphareocaris Calman, 1913 (nomen novum for Aphareus Paulson, 1875, gender feminine)
Sicyonella antennata Hansen, 1919: 30; Plate 2, fg. 5a-c; Plate 3, fg. 1a-f. [Siboga stn 258, Tual anchorage,
Kei Islands, 22 m]
Sicyonella inermis (Paulson, 1875)
= Aphareus inermis Paulson, 1875: 117; Plate 18, fg. 3. [Red Sea]
= Aphareocaris elegans Calman, 1913: 219; Plate 16, fgs 1-16. [Thursday Island, Torres Straits]
Sicyonella maldivensis Borradaile, 1910: 259; Plate 16, fgs 3-3a. [North Male Atoll, Maldives; lectotype
designation by Fukuoka, Tamaki & Kikuchi, 2005] (Fig. 7)
Suborder PLEOCYEMATA Burkenroad, 1963
Infraorder PROCARIDIDEA Felgenhauer & Abele, 1983
Family PROCARIDIDAE Chace & Manning, 1972
Procaris Chace & Manning, 1972
= Procaris Chace & Manning, 1972 (type species Procaris ascensionis Chace & Manning, 1972, by
Procaris ascensionis Chace & Manning, 1972: 6; Figs 4-9. [coral pool back of Shelly Beach, Ascension Is-
land]
Procaris chacei C.W.J. Hart & Manning, 1986: 408; Figs 1-26. [Bermuda, Hamilton Parish, Green Bay Cave
(North Shore passage), 3219'N 6444'W]
Procaris hawaiana Holthuis, 1973a: 12; Figs 4-6. [near Nukuele Point, Cape Kinau, Maui, Hawaiian Is-
lands, in lava pool] (Fig. 8)
Procaris mexicana von Sternberg & Schote, 2004: 514-522; Figs 1-3. [Mxico, Cueva Quebrada, Chanka-
naab Park, Cozumel, Quintana Roo]
Procaris noelensis Bruce & Davie, 2006: 23-33; Figs 1-3. [Runaway Cave, c. 1.5 km south from the North
East Point of Christmas Island]
Fig. 8. Procaris hawaiana Holthuis, 1973. Photo by Troy Sakihara.
Vetericaris Kensley & Williams, 1986
= Vetericaris Kensley & Williams, 1986 (type species Vetericaris chaceorum Kensley & Williams, 1986,
by monotypy, gender feminine)
Vetericaris chaceorum Kensley & Williams, 1986: 419, Figs 2-7. [Lua o Palahemo, 150 m north of shoreline
at Ka Lae (South Point), Hawaii Island, 1855'N 15542'W, 33 m]
Infraorder STENOPODIDEA Spence Bate, 1888
Family MACROMAXILLOCARIDIDAE Alvarez, Ilife & Villalobos, 2006
Macromaxillocaris Alvarez, Ilife & Villalobos, 2006
= Macromaxillocaris Alvarez, Ilife & Villalobos, 2006 (type species Macromaxillocaris bahamaensis
Alvarez, Ilife & Villalobos, 2006, by original designation and monotypy, gender feminine)
Macromaxillocaris bahamaensis Alvarez, Ilife & Villalobos, 2006: 370; Figs 2-7. [Oven Rock Cave, Great
Guana Cay, Exuma Cays, Bahamas] (Fig. 9)
Family SPONGICOLIDAE Schram, 1986
Engystenopus Alcock & Anderson, 1894
= Engystenopus Alcock & Anderson, 1894 (type species Engystenopus palmipes Alcock & Anderson,
1894, by monotypy, gender masculine)
Engystenopus palmipes Alcock & Anderson, 1894: 149; Plate 9, fg. 1. [Investigator stn 172, Bay of Bengal,
of Trincomallee, 200-350 fms]
Globospongicola Komai & Saito, 2006
= Globospongicola Komai & Saito, 2006 (type species Globospongicola nudibranchus Komai & Saito,
Globospongicola nudibranchus Komai & Saito, 2006: 268; Figs 1-6. [Kai Islands, Banda Sea, Indonesia,
605'S 13244'E, 268-210 m]
Globospongicola spinulatus Komai & Saito, 2006: 276; Figs 7-12. [Vanuatu, 1750.35'S 16839.33'E, 437-504 m]
Fig. 9. Macromaxillocaris bahamaensis Alvarez, Ilife & Villalobos, 2006. Photo by T.M. Ilife.
Microprosthema Stimpson, 1860a
= Microprosthema Stimpson, 1860a (type species Microprosthema valida Stimpson, 1860a, by mono-
typy, gender neuter)
= Stenopusculus Richters, 1880 (type species Stenopusculus crassimanus Richters, 1880, designated by
Holthuis, 1955, a junior subjective synonym of Microprosthema valida Stimpson, 1860a, gender mas-
culine)
Microprosthema emmiltum Goy, 1987: 717; Figs 1-4. [116'36''S 9029'42''W, of Black Beach, Isla Santa Ma-
ria, Galapagos Islands; rocky shores]
Microprosthema fujitai Saito & Okuno, 2011: 84; Figs 1-7. [Maeda-misaki, Okinawa Island, Ryukyu Is-
lands, 3 m]
Microprosthema granatense Criales, 1997: 538; Figs 1-4. [Granate Bay, Caribbean coast of Colombia,
1118'N 749'W, 23 m]
Microprosthema inornatum Manning & Chace, 1990: 26; Figs 12-13. [of North Point, Ascension Island, 18
m]
Microprosthema jareckii Martin, 2002: 109; Figs 1-5. [Monkey Point, Guana Island, British Virgin Islands,
10 m]
Microprosthema looensis Goy & Felder, 1988: 1286; Figs, 5-7. [near study fnger number 3, Looe Key, Flor-
ida, USA, 4-5 m]
Microprosthema lubricum Saito & Okuno, 2011: 93; Figs 8-11. [Agenashiku Islet, the Kerama Islands, the
Ryukyu Islands, 3m]
Microprosthema manningi Goy & Felder, 1988: 1277; Figs 1-4. [1729'N 8810'W, Stann Creek District, Carrie
Bow Cay, Belize, near shore in dead conch shell]
Microprosthema plumicorne (Richters, 1880) (Fig. 10)
= Stenopusculus plumicornis Richters, 1880: 167; Plate 18, fgs 16-26. [Fouquets, Mauritius]
Microprosthema scabricaudatum (Richters, 1880)
= Stenopusculus scabricaudatus Richters, 1880: 168; Plate 18, fgs 30-32. [Fouquets, Mauritius]
Microprosthema semilaeve (von Martens, 1872)
= Stenopus semilaevis von Martens, 1872: 144. [Cuba]
= Stenopusculus spinosus Pocock, 1890: 523. [Fernando de Noronha]
Fig. 10. Microprosthema plumicorne (Richters, 1880). Photo by Arthur Anker.
Microprosthema validum Stimpson, 1860a
= Microprosthema valida Stimpson, 1860a: 45. [in sinu insulae Ousima, sublitoralis, in locis lapillosis
algosisque]
= Stenopusculus crassimanus Richters, 1880: 168; Plate 18, fgs 27-29. [Fouquets, Mauritius]
= Stenopus robustus Borradaile, 1910: 260; Plate 16, fg. 4. [Salomon Atoll, Chagos Archipelago]
Paraspongicola de Saint Laurent & Cleva, 1981
= Paraspongicola de Saint Laurent & Cleva, 1981 (type species Paraspongicola pusilla de Saint Laurent
& Cleva, 1981, by original designation and monotypy, gender masculine)
Paraspongicola acantholepis Komai, 2011: 305; Figs 12-15. [R/V Tansei-maru, KT07-31, stn L3-200, Isu Is-
lands, N of Toshima Island, 3434.04'N 13918.37'E to 3433.56'N 13917.81'E, 187-261 m]
Paraspongicola infatus de Saint Laurent & Cleva, 1981
= Spongicola infata de Saint Laurent & Cleva, 1981: 179; Fig. 13. [1402.2'N 12017.7'E, 193-184 m]
Paraspongicola pusillus de Saint Laurent & Cleva, 1981
= Paraspongicola pusilla de Saint Laurent & Cleva, 1981: 181; Figs 14-15. [1356.3'N 12016.2'E, 150-
159 m]
Spongicola De Haan, 1844 [in De Haan, 1833-1850]
= Spongicola De Haan, 1844 [in De Haan, 1833-1850] (type species Spongicola venusta De Haan, 1844
[in De Haan, 1833-1850], by monotypy, gender masculine)
Spongicola andamanicus Alcock, 1901
= Spongicola andamanica Alcock, 1901: 148; Plate 2, fg. 2. [Andaman Sea, 170 and 238-290 fms]
= Spongicola henshawi Rathbun, 1906: 901; Plate 24, fg. 8. [south coast of Molokai Island, Hawaii,
169-182 fms]
= Spongicola henshawi spinigera de Saint Laurent & Cleva, 1981: 174; Figs 10b, 11c, g-h. [847'N
12331.2'E, 333 m]
= Spongicola holthuisi de Saint Laurent & Cleva, 1981: 177; Fig. 12. [1359.8'N 12018.6'E, 192-188 m]
Spongicola cubanicus Ortiz, Gmez & Lalana R., 1994
= Spongicola cubanica Ortiz, Gmez & Lalana R., 1994: 189; Figs 2-7. [Baha Honda, costa norte de la
Provincia de Pinar del Rio, 300 m, asociado a la esponja silcea Dactylocalyx pumiceus]
Spongicola depressus Saito & Komai, 2008: 103; Figs 8-10. [Loyalty Islands, depth unknown]
Spongicola goyi Saito & Komai, 2008: 107; Figs 11-15. [2203.60'S 16727.00'E, 850 m]
Spongicola japonicus Kubo, 1942a
= Spongicola japonica Kubo, 1942a: 90; Figs 1-2. [Kumanonada, of Mie Prefecture, about 300 m, com-
mensal with Euplectella marshalii]
Spongicola levigatus Hayashi & Ogawa, 1987
= Spongicola levigata Hayashi & Ogawa, 1987: 367; Figs 1-4. [East China Sea, 3044.7'N 12748.3'E,
about 200 m]
Spongicola parvispinus Zarenkov, 1990
= Spongicola parvispina Zarenkov, 1990: 218. [Sala-y-Gomez ridges, southeastern Pacifc, 470-485 m]
Spongicola robustus Saito & Komai, 2008: 118; Figs 20-24. [Mauritius, 1029.7'S 6112.4'E, 115-110 m]
Spongicola venustus De Haan, 1844 [in De Haan, 1833-1850]
= Spongicola venusta De Haan, 1844 [in De Haan, 1833-1850]: Plate 46, fg. 9 (1844); 194, Plate P (1849).
[Japan; lectotype designation by Yamaguchi & Baba, 1993]
Spongicoloides Hansen, 1908
= Spongicoloides Hansen, 1908 (type species Spongicoloides profundus Hansen, 1908, by monotypy,
gender masculine)
Spongicoloides evolutus (Bouvier, 1905b)
= Spongicola evoluta Bouvier, 1905b: 983. [au large des ctes du Sahara, 882 m]
Spongicoloides galapagensis Goy, 1980: 760; Figs 1-4. [Galapagos Islands, 0029' 8954'30''W, 717 m]
Spongicoloides hawaiiensis Baba, 1983: 477; Figs 1-2. [French Frigate Shoals, north-western Hawaiian Is-
lands, 640 m, associated with glass sponge Euplectella sp.]
Spongicoloides iheyaensis Saito, Tsuchida & Yamamoto, 2006: 224; Figs 3-8. [the North Knoll of the Iheya
Ridge, Ryukyu Islands, 2747.17'N 2653.91'E, 988 m]
Spongicoloides inermis (Bouvier, 1905a)
= Richardina inermis Bouvier, 1905a: 749. [dans les parages de Sainte-Lucie, 22-423 brasses]
Spongicoloides koehleri (Caullery, 1896)
= Spongicola Khleri Caullery, 1896: 382; Plate 16, fgs 1-9. [Golfe de Gascogne, 4557'N 621'E, 1410
m]
Spongicoloides novaezelandiae Baba, 1979: 311; Figs 1-2. [Chatham Rise, 4444.0'S 17542.0'E, 990-1110 m]
Spongicoloides profundus Hansen, 1908: 45; Plate 3, fgs 5a-k; Plate 4, fgs 1a-l. [78 miles of the southwest
coast of Iceland, 6037'N 2752'W, 1480 m; lectotype designation by Goy, 1982]
Spongicoloides tabachnicki Burukovsky, 2009: 498; Figs 1-3. [Atlantic Ocean, Florida Straits, 2354'N
8127'W, 1003-1336 m]
Spongiocaris Bruce & Baba, 1973
= Spongiocaris Bruce & Baba, 1973 (type species Spongiocaris semiteres Bruce & Baba, 1973, by original
Spongiocaris hexactinellicola Berggren, 1993: 784; Figs 1-5. [2430'N 7428'W, Tartar Bank, Cat Island, Ba-
hamas, 60-610 m; in the internal cavity of Euplectella jovis]
Spongiocaris goyi Ortiz, Lalana & Varela, 2007: 26; Figs 1-2. [sur de la Bahia de Cienfuegos, 297-351 m, en
la esponja Pheronema annae ?]
Spongiocaris semiteres Bruce & Baba, 1973: 155; Figs 1-6. [13 miles S.E. of Durban, South Africa, 230 fms,
from hexactinellid sponge]
Spongiocaris yaldwyni Bruce & Baba, 1973: 163; Figs 7-10. [15 miles N, 50E of Plate Island in the Bay of
Plenty, northeast of North Island, New Zealand, 320-340 fms, from Regadrella okinoseana]
Family STENOPODIDAE Claus, 1872
Juxtastenopus Goy, 2010
= Juxtastenopus Goy, 2010 (type species Engystenopus spinulatus Holthuis, 1946a, by original designa-
tion and monotypy, gender masculine)
Juxtastenopus spinulatus (Holthuis, 1946a)
= Engystenopus spinulatus Holthuis, 1946a: 45; Plate 4, fgs a-b. [Lobetobi Strait, E. of Flores, 827'S
12254.5'E, 247 m]
Odontozona Holthuis, 1946a
= Odontozona Holthuis, 1946a (type species Stenopus ensiferus Dana, 1852a, by original designation,
gender feminine)
Odontozona addaia Pretus, 1990: 343; Figs 1-6. [4001'57''N 411'46''E, litoral marine cave near Port
Addaia Bay, NE coast of Minorca, Balearic Islands, Spain]
Odontozona anaphorae Manning & Chace, 1990: 29; Figs 14-15. [of North Point, Ascension Island, 10 m]
Odontozona crinoidicola Saito & Fujita, 2009: 124; Figs 1-7. [Mizugama, Okinawa islands, Ryukyu Islands,
3.2 m, associated with Phanogenia gracilis]
Odontozona edwardsi (Bouvier, 1908b)
= Richardina Edwardsi Bouvier, 1908b: 888. [au large des ctes marocaines et soudanaises]
Odontozona ensifera (Dana, 1852a)
= Stenopus ensiferus Dana, 1852a: 27. [archipelago Viti]
Odontozona fasciata Okuno, 2003: 167; Figs 1-6. [Shimoji-shima Island, Miyako Group, Ryukyu Islands,
2449.6'N 12508.2'E, 25 m]
Odontozona foresti Hendrickx, 2002: 406; Figs 1-3. [southern Gulf of California, 2543.5'N 10953.7'W,
1240-1270 m]
Odontozona libertae Gore, 1981: 153; Figs 4-5. [Elbow Reef, of Key Largo, Monroe County, Florida]
Odontozona minoica Dounas & Koukouras, 1989: 341; Figs 1-4. [3529'10''N 2417'20''E, of north-western
coast of Crete, 330 m]
Odontozona rubra Wicksten, 1982: 130; Figs 1-2. [Isla Blanca, of Guaymas, Sonora, Mexico, approx.
2752'N 11052'W, 6-9 m]
Odontozona sculpticaudata Holthuis, 1946a: 37; Plate 2, fg. F; Plate 4, fg. C. [Sape Strait, east of Soem-
bawa, 70 m]
Odontozona spinosissima Kensley, 1981: 66; Figs 4-5. [3159'S 2922'E, 150-200 m]
Odontozona spongicola (Alcock & Anderson, 1899)
= ?Richardina spongicola Alcock & Anderson, 1899: 291. [Andaman Sea, 498 fms, from hexactinellid
sponge]
Odontozona striata Goy, 1981: 843; Figs 1-5. [Gulf of Mexico, West of Cabo San Antonio, Cuba]
Richardina A. Milne-Edwards, 1881a
= Richardina A. Milne-Edwards, 1881a (type species Richardina spinicincta A. Milne-Edwards, 1881a,
by monotypy, gender feminine; name placed on the Ofcial List of Generic Names in Zoology in
Opinion 712 in 1964)
Richardina fredericii Lo Bianco, 1903: 250; Plate 8, fgs 27-28. [a circa 9 chilometri da Punta Campanella,
direzione S.E., 1100 m; a circa 6.5 chilometri dai Galli di Positano, direzione E., 950 m; a circa 9
chilometri dai Galli di Positano, direzione O.S.O., 1100 m; a circa 10 chilometri da Punta Carena,
direzione E.N.E., 100 m]
Richardina ohtsukai Saito & Komatsu, 2009: 31; Figs 1-6. [Kerama Islands, N of Nagan-nu Island,
2623.15'N 12730.09'E, 730-728 m]
Richardina parvioculata Saito & Komatsu, 2009: 39; Figs 7-9. [of Tosa Bay, western Japan, 3256.45'N
13330.52'E, 517 m]
Richardina spinicincta A. Milne-Edwards, 1881a: 933. [4448'30''N 70'30''E, Golfe de Gascogne, 5100 m]
Stenopus Latreille, 1819
= Byzenus Rafnesque, 1814 (type species Byzenus scaber Rafnesque, 1814, by monotypy, an invalid
senior synonym of Stenopus spinosus Risso, 1827, gender masculine; name of genus and of its type
species suppressed for the purposes of the Principle of Priority but not for those of the Principle of
Homonymy, placed on the Ofcial Index of Rejected and Invalid Generic (resp. Specifc) Names in
Zoology in Opinion 522 in 1958)
= Stenopus Latreille, 1819 (type species Palmon hispidus Olivier, 1811, by monotypy, gender mascu-
line; name placed on the Ofcial list of Generic Names in Zoology in Opinion 522 in 1958)
= Embryocaris Ortmann, 1893 (type species, Embryocaris stylicauda Ortmann, 1893, by monotypy, a
junior subjective synonym of Palmon hispidus Olivier, 1811, gender feminine)
Stenopus chrysexanthus Goy, 1992: 80; Figs 1-4. [Heron Island, Capricorn Group, Queensland, Australia,
ca. 2325'S 15155'E, south west reef fat, algal crest]
Stenopus cyanoscelis Goy, 1984: 116. [Broadhurst Reef, Of Townsville, Queenland]
Stenopus devaneyi Goy, 1984: 117. [French Polynesia, Marquesas Islands, Nuka Hiva Island, outer portion
of Taiohae Baie, W side of islet, Sentinelle de lEst, 6.2 m, according to Goy & Randall, 1986]
Stenopus earlei Goy, 1984: 117. [of Makua, Oahu Island, Hawaiian Islands, 39.6 m, according to Goy &
Randall, 1986]
Stenopus goyi Saito, Okuno & Chan, 2009: 111; Figs 1-5. [Donggang fshing port, Pingtung County, Taiwan,
commercial trawler]
Stenopus hispidus (Olivier, 1811) (Fig. 11)
= Squilla Groenlandica Seba, 1759: 54; Plate 21, fgs 6-7. [Type locality not indicated; see Davie, 2002
for status of the older synonyms]
= Astacus muricatus Olivier, 1791: 346. [mer du Gronland]
= Cancer (Astacus) longipes Herbst, 1793 [in Herbst, 1791-1796]: 90; Plate 31, fg. 2. [Grnland]
= Penus borealis Latreille, 1802: 250. [mers du Nord]
= Palmon hispidus Olivier, 1811: 666. [Type locality not indicated]
= Palmon ? asper Latreille, 1818: 3; Plate 293, fg. 3. [Type locality not indicated]
= Embryocaris stylicauda Ortmann, 1893: 85; Plate 6, fg. 4. [Plankton-Expedition der Humboldt-Stif-
tung, sdl. Aequatorialstrom JN 235 (0.1S 44.2W, 0-400 m)]
Fig. 11. Stenopus hispidus (Olivier, 1811). Photo by Arthur Anker.
Stenopus pyrsonotus Goy & Devaney, 1980: 781; Figs 1-6. [of Pokai Bay, Oahu Island, Hawaii, 22.9 m]
Stenopus scutellatus Rankin, 1898: 242; Plate 24, fg. 3. [Silver Cay, New Providence]
Stenopus spinosus Risso, 1827: 66; Plate 3, fg. 8. [golfe de Nice, rgions profondes]
= Byzenus scaber Rafnesque, 1814: 23. [Sicile; name suppressed for the purposes of the Principle of
Priority but not for those of the Principle of Homonymy, placed on the Ofcial Index of Rejected
and Invalid Specifc Names in Zoology in Opinion 522 in 1958]
Stenopus tenuirostris De Man, 1888b: 567; Plate 22a, fg. 5. [Amboina]
Stenopus zanzibaricus Bruce, 1976a: 90; Figs 1-5. [546.6'S 3923.5'E, Mwenba Island, of N.E. Zanzibar
Island, 0.5 m]
Infraorder CARIDEA Dana, 1852b
Superfamily PASIPHAEOIDEA Dana, 1852a
Family PASIPHAEIDAE Dana, 1852a
Alainopasiphaea Hayashi, 1999
= Alainopasiphaea Hayashi, 1999 (type species Pasiphaea nudipedia Burukovsky, 1993b, by original
designation and monotypy, gender feminine)
Alainopasiphaea australis (Hanamura, 1989)
= Pasiphaea australis Hanamura, 1989: 59; Figs 5-7. [Maria Island area, of east coast of Tasmania,
southern Australia, 4242.0'S 14824.0'E, 345-350 m]
Alainopasiphaea nudipeda (Burukovsky, 1993b)
= Pasiphaea nudipeda Burukovsky, 1993b: 35; Figs 1(8-13). [2528'-2534'5"S 3508'-3500'E, 535-490 m]
Eupasiphae Wood-Mason in Wood-Mason & Alcock, 1893
= Eupasipha Wood-Mason in Wood-Mason & Alcock, 1893 (type species Parapasipha (Eupasipha)
Gilesii Wood-Mason, 1892, designated by Holthuis, 1955b, gender feminine)
Eupasiphae gilesii (Wood-Mason, 1892)
= Parapasipha Gilesii Wood-Mason, 1892: Plate 3, fg. 8; accompanying text in Wood-Mason in
Wood-Mason & Alcock, 1893: 166 (as Parapasipha (Eupasiipha) Gilesii). [of Cinque Island, Anda-
man Sea, 650 fms]
= Eupasiphae rhinocerata Burukovsky, 1977: 473; Figs A-C. [of Morocco, near Meteor Shoal, 3000'N
2843'W, 1400-2060 m]
Eupasiphae latirostris (Wood-Mason & Alcock, 1891b)
= Parapasipha latirostris Wood-Mason & Alcock, 1891b: 196. [Investigator stn 105 (Arabian Sea,
152'N 7234'E), 740 fms]
Eupasiphae paucidentata Crosnier, 1988b: 789; Figs 2c, 3-4, 5b-c. [Madagascar, N.O., 2126.5S 4311'E, 810-
1020 m]
Eupasiphae serrata (Rathbun, 1902a)
= Parapasiph serrata Rathbun, 1902a: 904. [of Cortez Bank, California, 984 fms]
= Parapasiphae Grimaldii Coutire, 1911a: 157. [Princesse-Alice stn 3036, of Spain and Portugal,
3606'40"N, 1018'W, 0-4740 m, according to Crosnier, 1988b]
Glyphus Filhol, 1884
= Glyphus Filhol, 1884 (type species Glyphus marsupialis Filhol, 1884, by monotypy, gender mascu-
line)
= Sympasiphaea Alcock, 1901 (type species Sympasiphaea annectens Alcock, 1901, by monotypy, gen-
der feminine)
Glyphus marsupialis Filhol, 1884: 231. [Talisman stn 75 (2539'N 1602'W)]
= Sympasiphaea annectens Alcock, 1901: 63. [Arabian Sea, south-west of Cape Comorin, 487 fms]
= Sympasiphaea imperialis Terao, 1922: 110; unnumbered fgure. [purchased at the market of Uchiura,
Prov. Idzu]
Leptochela Stimpson, 1860a
= Leptochela Stimpson, 1860a (type species Leptochela gracilis Stimpson, 1860a, designated by Kemp,
= Proboloura Chace, 1976 (type species Leptochela carinata Ortmann, 1893, by original designation
and monotypy, gender feminine)
Leptochela (Leptochela) aculeocaudata Paulson, 1875
= Lep.[tochela] aculeocaudata Paulson, 1875: 100; Plate 16, fgs 1-1s. [Red Sea]
Leptochela (Leptochela) bermudensis Gurney, 1939a
= Leptochela bermudensis Gurney, 1939a: 427; Figs 1-10. [about 7 miles of the south shore, between
1000 m and surface]
? = Leptochela carinata Ortmann, 1893: 41 (partim, not Plate 4, fg. 1). [Plankton-Expedition der Hum-
boldt-Stifung, Kstenbank vor der Tocantins-Mndung, JN 236 (0.1S 45.2W), 50-100 m]
Leptochela (Leptochela) chacei Hayashi, 1995: 85; Figs 2-4. [Viet-Nam, Cai Dua]
Leptochela (Leptochela) crosnieri Hayashi, 1995: 89; Figs 5-7. [New Caledonia, st. Balabio, 2003.6'S
16407.7'E, 13 m]
Leptochela (Leptochela) gracilis Stimpson, 1860a
= Leptochela gracilis Stimpson, 1860a: 42. [Sinu Kagosima, in profundis]
= Leptochela pellucida Boone, 1935: 105; Plates 26-27. [Near Equator, to the South Brothers Island,
south entrance of Durian Straits, Dutch East Indies, 29'N by E 10447'E, 14 fms]
Leptochela (Leptochela) hawaiiensis Chace, 1976: 15; Figs 11-13. [Hawaii, south coast of Oahu, Diamond
Head Light, 62S 3.9'E]
Leptochela (Leptochela) irrobusta Chace, 1976: 19; Figs 14-18. [Bikini Atoll, northeast end of lagoon at
Bowditch anchorage, surface light at night]
Leptochela (Leptochela) japonica Hayashi & Miyake, 1969
= Leptochela japonica Hayashi & Miyake, 1969: 1; Figs 1-2. [Chiiwa Bay, northwestern Kyushu, 65-66 m]
Leptochela (Leptochela) papulata Chace, 1976: 26; Figs 22-24. [east of Cape Lookout, North Carolina,
3435'30"N 7545'30"W, 59 m]
Leptochela (Leptochela) pugnax De Man, 1916
= Leptochela pugnax De Man, 1916: 148. [Bay of Bima, near south fort, 55 m; Tual-anchorage, Kei-is-
lands, 22 m; Elat, west coast of Great-Kei-island, 27 m]
Leptochela (Leptochela) robusta Stimpson, 1860a
= Leptochela robusta Stimpson, 1860a: 43. [Mari sinensi, prof. 20 org; prope insulam Loo Choo
quoque]
Leptochela (Leptochela) serratorbita Spence Bate, 1888 (Fig. 12)
= Leptochela serratorbita Spence Bate, 1888: 859; Plate 139, fg. 1. [Saint Thomas, Virgin Islands, shal-
low water]
Leptochela (Leptochela) sydniensis Dakin & Colefax, 1940
= Leptochela sydniensis Dakin & Colefax, 1940: 153; Figs 245-246. [New South Wales]
? = Leptochela hainanensis Yu, 1936: 87; Figs 1-3. [Hai-kiu-sche]
Leptochela (Proboloura) carinata Ortmann, 1893
= Leptochela carinata Ortmann, 1893: 41 (partim); Plate 4, fg. 1. [Plankton-Expedition der Humboldt-
Stifung, Kstenbank vor der Tocantins-Mndung, JN 236 (0.1S 45.2W), 50-100 m]
Leptochela (Proboloura) soelae Hanamura, 1987: 21; Figs 6-8. [north-west Australian shelf, 1905.0'S
11853.8'E, 82 m]
Parapasiphae Smith, 1884
= Parapasipha Smith, 1884 (type species Parapasipha sulcatifrons Smith, 1884, designated by Fowler,
1912, gender feminine; name placed in its corrected spelling on the Ofcial List of Generic Names
in Zoology in Opinion 470 in 1957)
= Orphania Spence Bate, 1888 (type species Orphania tenuimana Spence Bate, 1888, a junior subjective
synonym of Parapasipha sulcatifrons Smith, 1884, gender feminine)
= Dantecia Caullery, 1896 (type species Dantecia Caudani Caullery, 1896, a subjective synonym of
Parapasipha sulcatifrons Smith, 1884, by monotypy, gender feminine)
Parapasiphae compta Smith, 1884
= Parapasipha compta Smith, 1884: 389. [Albatross stn 2039, 3819'26"N 6820'20"W, 2369 fms]
Fig. 12. Leptochela (Leptochela) serratorbita Spence Bate, 1888. Photo by Arthur Anker.
Parapasiphae cristata Smith, 1884
= Parapasipha cristata Smith, 1884: 388; Plate 5, fg. 3. [Albatross stn 2100, 3922'N 6834'30"W, 1628
fms]
= Parapasipha macrodactyla Chace, 1939: 33. [S.S.E. of Bahia de Guantanamo, Oriente Province,
Cuba, 1940'N 7503'W, 1920 fms]
Parapasiphae kensleyi Wasmer, 2005: 165; Figs 1-3. [31N 159W, 4200 m]
Parapasiphae sulcatifrons Smith, 1884
= Parapasipha sulcatifrons Smith, 1884: 384; Plate 5, fg. 4; Plate 6, fgs 1-7. [Albatross stns 2002,
3702'42''N 7417'36''W, 641 fms; 2034, 3927'10''N 6956'20''W, 1346 fms; 2037, 3853'00''N 6923'30''W,
1731 fms; 2072, 4153'00''N 6535'00''W, 858 fms; 2074, 4143'00''N 6521'50''W, 1309 fms; 2094,
3944'30''N 7104'00''W, 1022 fms; 2097, 3756'20''N 7057'30''W, 1917 fms; 2099, 3712'20''N
6939'00''W, 2949 fms; 2105, 3750'00''N 7303'50''W, 1395 fms; 2110, 3512'10''N 7457'15''W, 516 fms]
= Orphania tenuimana Spence Bate, 1888: 872; Plate 141, fg. 4. [Challenger stn 45, 3834'N 7210'W,
south-east of New York, 1240 fms]
= Dantecia Caudani Caullery, 1896: 372, Plate 14, fgs 1-11. [Golfe de Gascogne, 4628'N 7W, 1710 m]
= Pasiphaea metriomma Dohrn, 1908: 9. [Valdivia Expedition stn 26 (ca. 1000 mls E of Madeira, 3159'N
55'W, 1800 m)]
Pasiphaea Savigny, 1816
= Pasiphaea Savigny, 1816 (type species A.[lpheus] sivado Risso, 1816, by monotypy, gender feminine)
= Pasipha (Phye) Wood-Mason, 1892 (type species Pasipha (Phye) Parapasipha Alcocki Wood-Mason
& Alcock, 1891b, by monotypy, gender feminine)
Pasiphaea acutifrons Spence Bate, 1888
= Pasipha acutifrons Spence Bate, 1888: 871; Plate 141, fg. 3. [Challenger stns 311, 5245'30''S
7346'0''W, of Port Churruca, Patagonia, 245 fms; 236, 3458'N 13929'E, south of Japan, 77 fms]
= Pasipha forceps A. Milne-Edwards, 1891: 51; Plate 6, fgs 2-2e. [dtroit de Magellan, au sud-est de
Port-Famine, 326 m]
Pasiphaea afnis Rathbun, 1902a
= Pasipha afnis Rathbun, 1902a: 905. [near Cortez Bank, California, 984 fms]
Pasiphaea alcocki Wood-Mason & Alcock, 1891b
= Parapasipha Alcocki Wood-Mason & Alcock, 1891b: 196. [Bay of Bengal, 1611'15"N 8230'30"W,
922 fms]
Pasiphaea americana Faxon, 1893
= Pasiphaeia (sic) cristata americana Faxon, 1893: 208. [Gulf of Panama, 732'36"N 7916'W, 523 m;
lectotype designation by Hayashi, 2004]
Pasiphaea amplidens Spence Bate, 1888
= Pasipha amplidens Spence Bate, 1888: 870; Plate 141, fg. 2. [Challenger stn 3458'N 13929'E, 775
fms]
= Pasiphaea vereschhaka Burukovsky, 1993b: 33; Fig. 1(1-7). [1222'S 5302'E, 1045-1050 m]
Pasiphaea arabica Timofeev, 1997: 142; Fig. 1. [Arabian Sea, 1632'2''N 6425'5''E, collecting horizon 110-
100 m, botom at 3600 m]
Pasiphaea balssi Burukovsky & Romensky, 1987: 52; Figs 1, 2(9), 4. [South Atlantic, 4155'S, 000'E, 910 m]
Pasiphaea barnardi Yaldwyn, 1971: 86 [Cook Strait, 4142.5'S 1759'E, ca. 500 fms]
Pasiphaea berentsae Kensley, Tranter & Grifn, 1987: 294; Figs 7-9. [New South Wales, northeast of Norah
Head, 3311'S 15224'E to 3309'S 15225'E, 720 m]
Pasiphaea burukovskyi Wasmer, 1993: 79; Figs 22-24. [4022'S 16825'E to 4026'S 16830'E, 600-750 m]
Pasiphaea chacei Yaldwyn, 1962: 18; Figs 1-17. [3336'11"N 11832'13"W 3330'52"N 11822'48"W, 475
fms]
Pasiphaea corteziana Rathbun, 1902a
= Pasipha corteziana Rathbun, 1902a: 905. [near Cortez Bank, California, 776 fms]
Pasiphaea cristata Spence Bate, 1888
= Pasipha cristata Spence Bate, 1888: 865; Plate 140; Plate 141, fg. 1. [Challenger stn 173, 199'35"S
17941'50"E, of Matuka, Fii Islands, 315 fms]
Pasiphaea crosnieri Hayashi, 2004: 334; Figs 7-9. [Philippines, 1340.7'N 12030.0'E, 685-757 m]
Pasiphaea debitusae Hayashi, 1999: 281; Figs 8-10. [Indonesia, 0515'S 13301'E, 605-576 m]
Pasiphaea diaphana Burukovsky & Romensky, 1980: 1097. [South-East Atlantic, 3148'S 0218'E, 1100 m]
Pasiphaea dofeini Schmit, 1932a: 333; Fig. 1. [Punta Arenas, now Magelhanes, Chile]
Pasiphaea ecarina Crosnier, 1969: 537; Figs 19, 23-36. [Congo, 508'S 1122'E, 795-805 m]
Pasiphaea emarginata Rathbun, 1902a
= Pasipha emarginata Rathbun, 1902a: 905. [Gulf of California, 857 fms]
Pasiphaea faxoni Rathbun, 1904: 22; nomen novum for Pasiphaea acutifrons sensu Faxon, 1895 nec Spence
Bate, 1888 [Albatross stns 3402 (057'30"S 8903'30"W), 421 fms; 3403 (058'30" 8917'00"W) 384 fms
and 3406 (016'00"S 9021'30"W), 551 fms]
Pasiphaea fagellata Rathbun, 1906
= Pasipha fagellata Rathbun, 1906: 928; Fig. 78; Plate 23, fg. 2. [Kaiwi Channel, Hawaii, 411-442
fms]
Pasiphaea fragilis Hayashi, 1999: 285; Fig. 11. [Loyalty Islands, 2023.0'N 16620.1'E, 780 m]
Pasiphaea gelasinus Hayashi & Yaldwyn, 1998: 512; Figs 1-3. [South Indian Ocean, Seamounts of Saint
Paul and Amsterdam Islands, 3737.8'S, 7751.8'E, 730-905 m]
Pasiphaea gracilis Hayashi, 1999: 290; Figs 15-17. [New Caledonia, 2308'S 16651'E, 850 m]
Pasiphaea grandicula Burukovsky, 1976: 17; Fig. 1. [South Atlantic, 2611'S, 0602'8"E, 1150 m]
Pasiphaea hoplocerca Chace, 1940: 124; Figs 4-5. [Bermuda Oceanographic Expedition Nets 713, 753 and
778 (3212'N 6436'W), 700 fms]
Pasiphaea japonica Omori, 1976: 250; Figs 1-2. [Of Iwase, Toyama Bay, 0-300 m]
Pasiphaea kapala Kensley, Tranter & Grifn, 1987: 297; Figs 10-12. [New South Wales, northeast of Norah
Head, 3308'S 15227'E to 3310'S 15224'E, 580 m]
Pasiphaea kaiwiensis Rathbun, 1906
= Pasipha kaiwiensis Rathbun, 1906: 927; Fig. 76; Plate 23, fg. 4. [Kaiwi Channel, 343-337 fms]
Pasiphaea korzuni Burukovsky, 1995: 121; Figs 1-5. [Aden Bay, near Ras-Fartak Island, depth unknown]
Pasiphaea laevis Hayashi, 1999: 287; Figs 12-14. [Indonesia, Moluccas, 0608'S 13245'E, 390-502 m]
Pasiphaea ledoyeri Hayashi, 2006a: 215; Figs 6-8. [South Indian Ocean, 4633.8'S 4123.7'E, 660-911 m, in
stomach of Dissostichus eleginoides]
Pasiphaea levicarinata Hanamura, 1994: 167; Figs 1-2. [north-western Australia, 1830'S 11723'E, 530 m]
Pasiphaea liocerca Chace, 1940: 122; Figs 2-3. [Bermuda Oceanographic Expeditions Net 1160 (3212'N
6436'W), 800 fms]
Pasiphaea longitaenia Kensley, Tranter & Grifn, 1987: 301; Figs 13-14. [New South Wales, east of Long
Reef Point, 3343'S 15156'E to 3339'S 15158'E, 626-637 m]
Pasiphaea magna Faxon, 1893: 209. [Albatross stn 3384 (of Panama, 731'30"N, 7915'00"W), 458 fms]
Pasiphaea major Hayashi, 2006a: 219; Figs 9-11. [Norfolk Ridge, 2509'S 16853'E, 1173-1340 m]
Pasiphaea marisrubri Iwasaki, 1989: 178; Figs 1-3. [Central Red Sea, 2114.80'N 3715.40'E 2118.20'N
3712.00'E, 0-220 m]
Pasiphaea mclaughlinae Hayashi, 2006b: 342; Figs 1-3. [SE coast of Taiwan, 2213.8'N 12101.8'E, 1222-1226 m]
Pasiphaea meiringnaudei Kensley, 1977: 34-37; Figs 11-14. [Meiring Naude stn 107 (South Africa, of Natal
coast, 2837.8'S 3238.4'E, 1200-1000 m)]
Pasiphaea merriami Schmit, 1931b
= Pasipha merriami Schmit, 1931b: 391. [south of the Dry Tortugas, 253-283 fms]
= Pasiphaea nishiei Iwasaki, 1990: 190; Figs 1-2. [of Jamaica, 1734.3'N, 7603.3'W, 1490-1504 m]
Pasiphaea multidentata Esmark, 1866: 259. [Bjrumsforden omtrent 1/8 Miil fra Namsos]
= P.[asipha] norvegica M. Sars, 1866a: 260. [nomen nudum]
= Pasipha norvegica M. Sars, 1866b: 314. [Christianiaforden; fully illustrated in M. Sars, 1868]
= Pasipha (Phye) sicula Riggio, 1896 [in Riggio, 1895-1896]: 41; Plate 1 (1865, no legend), Figs 2a-b.
[proviene dal mare di Augusta]
Pasiphaea natalensis Burukovsky & Romensky, 1982: 1797; Figs 1-6. [south-western part of the Indian
Ocean, 3348.5'S 4430.8'E, 900-910 m]
Pasiphaea notosivado Yaldwyn, 1971: 86. [Cook Strait, 4132'S 17456'E, 50-100 fms]
Pasiphaea orientalis Schmit, 1931a: 267; Plate 32, fgs 1, 5. [Ryukyusho, Takao, Formosa]
Pasiphaea oshoroae Komai & Amaoka, 1993: 367; Figs 1-3. [Of Aleuten Islands, 4959.9'N 17655.4'W,
0-400 m]
Pasiphaea pacifca Rathbun, 1902a
= Pasipha pacifca Rathbun, 1902a: 905. [of Point Sur, California, 328 fms]
Pasiphaea philippinensis Hayashi, 1999: 277; Figs 5-7. [Philippines, 1339'N 12043'E, 520-550 m]
Pasiphaea planidorsalis Hayashi, 2004: 348; Figs 13-14. [New Caledonia, 1942.4'S 15850.8'E, 772-756 m]
Pasiphaea poeyi Chace, 1939: 31. [Bahia de Cochinos, Santa Clara Province, Cuba, 2207'N 8108'W, 220-
275 fms]
Pasiphaea princeps Smith, 1884
= Pasipha princeps Smith, 1884: 381; Plate 5, fg. 2. [Albatross stn 2095, 3929'N 7058'40"W, 1451 fms]
Pasiphaea propinqua De Man, 1916: 147. [0611'N 12037.5'E, of the Sulu Islands, 450 m]
Pasiphaea pseudacantha Hayashi, 2004: 353; Figs 16-18. [Kai Islands, Indonesia, 0608'S 13245'E, 390-502 m]
Pasiphaea rathbunae (Stebbing, 1914a)
= Phye rathbun Stebbing, 1914a: 295; Plate 31. [Scotia stn 450, 4800'S 950'W, 1332 fms]
Pasiphaea romenskyi Burukovsky, 1995: 123; Figs 6-11. [Ecuador Seamount, 0023'S, 5602'E, 200-235 m]
Pasiphaea scotiae (Stebbing, 1914a)
= Phye scoti Stebbing, 1914a: 294; Plate 30. [Scotia stns 417, 7122'S 1634'W, 1410 fms; 422, 6832'S
1249'W, from surface to 600 fms]
= Pasiphaea longispina Lenz & Strunck, 1914: 315; Plate 19, fgs 1-11. [6515'S 8019'E, also in der Nhe
des Eisrandes, im Magen eines Pinguins, Aptenodytes forsteri, gefunden]
Pasiphaea semispinosa Holthuis, 1951a: 9; Fig. 1. [of Angola, 755'S 1238'E, 235-460 m]
Pasiphaea sinensis Hayashi & Miyake, 1971: 39; Fig. 1. [East China Sea, 2927.2'N 12816.2'E 2933.3'N
12823.5'E, 1065-1075 m]
Pasiphaea sirenkoi Burukovsky, 1987: 37; Figs 1-5. [SE Indian Ocean, 1127.5'N 10956.2'E, 600 m]
Pasiphaea sivado (Risso, 1816)
= A.[lpheus] sivado Risso, 1816: 93; Plate 3, fg. 4. [sur la plage de Nice]
= P.[asiphaea] Savignyi H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 426. [Patrie in-
connue]
= Pasipha brevirostris H. Milne Edwards, 1837 [in H. Milne Edwards, 1836-1844]: Plate 54bis, fg. 2.
[Type locality not indicated]
? = Pasipha distincta Gurin-Mneville, 1844 [in Gurin-Mneville, 1829-1844]: 16, Plate 22, fg. 3.
(as Pasipha sivado). [la Mditerrane]
= Pasiphaea neapolitana Hope, 1851: 44. [Type locality not indicated]
Pasiphaea tarda Kryer, 1845: 453. [Type locality not indicated]
= Pasipha principalis Sund, 1913: 6; Figs 5-7, 9a-f. [Sogneford; xsund (E. of the Lofoten isles, about
67.5N)
Pasiphaea telacantha Hayashi, 2004: 363; Figs 22-24. [New Caledonia, 2308'S 16651'E, 850 m]
Pasiphaea timofeevi Burukovsky, 1993b: 35; Fig. 1(14-19). [Aden Bay, Socotra Island, 1220'N 5309'W, 515-
480 m]
Pasiphaea truncata Rathbun, 1906
= Pasipha truncata Rathbun, 1906: 928; Fig. 7; Plate 23, fg. 5. [vicinity of Modu Manu, 293-800 fms]
Pasiphaea unispinosa Wood-Mason, 1892
= Pasipha unispinosa Wood-Mason, 1892: Plate 3, fg. 7; accompanying text as Wood-Mason in
Wood-Mason & Alcock, 1893: 163. [7 miles SE by S of Ross Island, Andaman Sea, 265 fms]
Pasiphaea westindica Tchesunov, 1984: 993; Fig. 1. [Atlantic Ocean, 1248'N 6207'W, 1500 m]
Psathyrocaris Wood-Mason in Wood-Mason & Alcock, 1893
= Psathyrocaris Wood-Mason in Wood-Mason & Alcock, 1893 (type species Psathyrocaris fragilis
Wood-Mason in Wood-Mason & Alcock, 1893, by monotypy, gender feminine)
Psathyrocaris fragilis Wood-Mason in Wood-Mason & Alcock, 1893: 171; Plates 10-11. [1556'50"N
8130'30"E, 240 fms]
= Psathyrocaris infrma var. atlantica Caullery, 1896: 374; Plate 14, fgs 12-15. [Golfe de Gascogne,
4439'N, 210'W, 800 m]
Psathyrocaris hawaiiensis Rathbun, 1906: 928; Fig. 79. [vicinity of Modu Manu, 876 fms]
Psathyrocaris infrma Alcock & Anderson, 1894: 159. [Investigator stn 116, Andaman Sea, 1125'5''N
9247'6''E, 405 fms]
Psathyrocaris platyophthalmus Alcock & Anderson, 1894: 158. [Investigator stn 124, Laccadive Sea,
1047'45"N 7240'20"E, 705 fms]
Psathyrocaris plumosa Alcock & Anderson, 1894: 159. [Investigator stn 128, of Ceylon, 658'N 7726'50''E,
902 fms]
Superfamily OPLOPHOROIDEA Dana, 1852a
Family ACANTHEPHYRIDAE Spence Bate, 1888
Acanthephyra A. Milne-Edwards, 1881b (Fig. 13)
= Ephyra Roux, 1831 (an invalid junior homonym of Ephyra Peron & Leseur, 1810 (Coelenterata),
type species A.[lpheus] Pelagicus Risso, 1816, designated by Kingsley, 1880, gender feminine; name
placed on the Ofcial Index of Rejected and Invalid Generic Names in Zoology in Opinion 359 in
1955)
= Miersia Kingsley, 1880 (nomen novum for Ephyra Roux, 1831, gender feminine; name suppressed
for the purposes of the Principle of Priority but not for those of the Principle of Homonymy, placed
on the Ofcial Index of Rejected and Invalid Generic Names in Zoology in Opinion 359 in 1955)
= Acanthephyra A. Milne-Edwards, 1881b (type species Acanthephyra armata A. Milne-Edwards,
1881b, by original designation, gender feminine; name placed on the Ofcial List of Generic Names
= Bentheocaris Spence Bate, 1888 (type species Bentheocaris stylorostratis Spence Bate, 1888, desig-
nated by Holthuis, 1955b, gender feminine)
= Acanthephyropsis Riggio, 1895 [in Riggio, 1895-1896] (type species Acanthephyra pulchra A. Milne-
Edwards, 1890, a junior subjective synonym of Acanthephyra eximia Smith, 1884, gender feminine)
= Hoplocaricyphus Coutire, 1907a (type species Hoplocaricyphus similis Coutire, 1907a, probably a
subjective synonym of A.[lpheus] Pelagicus Risso, 1816, gender masculine)
Fig. 13. Acanthephyra spec. Photo by Tin-Yam Chan.
Acanthephyra acanthitelsonis Spence Bate, 1888: 745; Plate 125, fg. 3. [Challenger stns 106, 147'N 2426'W,
Atlantic, south-west of Sierra Leone, 1850 fms; 107, 122'N, 2636'W, Atlantic, south-west of Sierra
Leone, 1500 fms]
Acanthephyra acutifrons Spence Bate, 1888: 749 (partim); Plate 126, fg. 3. [of Kepulauan Aru, Indonesia,
541'S 13404'30"E, 1463 m; type locality restricted by Kemp, 1906a]
Acanthephyra armata A. Milne-Edwards, 1881b: 12. [prs de Sainte-Lucie, 422 brasses]
Acanthephyra brevicarinata Hanamura, 1984: 65; Figs 1-2. [1328.7'N, 11953.9'W, 1617-0 m]
Acanthephyra brevirostris Smith, 1885a: 504. [Albatross stns 5448, 3712'20"N 6939'00"W, 2949 fms; 5449,
3922'00"N, 6834'30"W, 1686 fms; 5673, 3750'00"N, 7303'50"W, 1395 fms]
= Hymenodora duplex Spence Bate, 1888: 843; Plate 136, fg. 3. [Challenger stn 147, 4616'S 4827'E, of
Marion Island, 1600 fms]
Acanthephyra carinata Spence Bate, 1888: 748; Plate 126, fg. 2. [Challenger stn 310, 5127'30"S 7403'00"W,
Sarmiento channel, Patagonia, 400 fms]
= Acanthephyra approxima Spence Bate, 1888: 755; Plate 126, fg. 8. [Challenger stn 310, 5127'30"S
7403'00"W, Sarmiento Channel, Patagonia, 400 fms]
Acanthephyra chacei Krygier & Forss, 1981: 96; Figs 1-2. [4445.2'N 12744.0'W, 0-2600 m]
Acanthephyra cucullata Faxon, 1893: 206. [Albatross stn 3381 (North Pacifc Ocean, Colombia, 456'00"N,
8052'30"W), 1772 fms]
Acanthephyra curtirostris Wood-Mason & Alcock, 1891b: 195. [Investigator stns 100 (1112'47"N 7425'30"E),
840 fms; 104 (1655'41"N 8321'18"E), 1000 fms]
= Acanthephyra acutifrons Spence Bate, 1888: 749 (partim) (nec Plate 126, fg. 3)
Acanthephyra eximia Smith, 1884: 376. [Albatross stn 2111, 3509'50"N 7457'40"W, 938 fms]
= Acanthephyra angusta Spence Bate, 1888: 737; Plate 124, fg. 6. [Challenger stn 194, 434'00"S
12957'30"E, of Banda Island, 200 fms]
= Acanthephyra edwardsii Spence Bate, 1888: 747; Plate 126, fg. 1. [Challenger stn 126, 1046'S 3608'W,
south of Pernambuco, 770 fms]
= Acanthephyra brachytelsonis Spence Bate, 1888: 753; Plate 126, fg. 7. [Challenger stns 170, 2955'S
17814'W, of the Kermadec Islands, 520 fms; 170A, 2945'S 17811'W, north of the Kermadec Is-
lands, 630 fms; 171, 2833'S 17750'W, north of the Kermadec Islands, 600 fms; 194, 434'N
12957'30''E, of Banda Island, 200 fms; 214, 433'N 1276'E, south of the Philippine Islands, 500 fms;
232, 3511'N 13928'E, Japan, 345 fms; 236, 3458'N, 13929'E, south of Japan, 775 fms; 318, 4232'S
5629'W, north of the Falkland Islands, 2040 fms]
= Acanthephyra pulchra A. Milne-Edwards, 1890: 163. [au large de Monaco, 1650 m]
= Acanthephyra frontieri Crosnier, 1987b: 699; Fig. 1. [Madagascar, 1302'S 4802'E, 1000-1525 m]
Acanthephyra faxoni Calman, 1939
= Acanthephyra Faxoni Calman, 1939: 191; Fig. 1. [John Murray Expedition stn 184, Gulf of Aden
(1436'06''N 5100'18''E - 1438'42''N 5057'42''E), 1270 m]
Acanthephyra fmbriata Alcock & Anderson, 1894
= Acanthephyra armata var. fmbriata Alcock & Anderson, 1894: 156. [Bay of Bengal [of Madras,
1250'N 8130'E], 475 fms; of the Malabar coast [of Goa, 1529'N 7241'E], 559 fms; illustrated in
Wood-Mason, 1892: Plate 3, fg. 1. (as Acanthephyra armata var.)]
Acanthephyra indica Balss, 1925a: 264; Figs 34-35. [Valdivia stn 215, 701'N 8556'E, 0-2500 m, zwischen
Sumatra und Ceylon]
Acanthephyra kingsleyi Spence Bate, 1888: 751; Plate 126, fg. 4. [Challenger stn 104, 225'N 201'W, Atlan-
tic, south-west of Sierra Leone, 2500 fms]
= Acanthephyra sexspinosa Kemp, 1939: 575. [Central and South Atlantic from 17N to 18S]
Acanthephyra media Spence Bate, 1888: 736; Plate 124, fg. 5. [Challenger stn 207, 1221'N 12215'E, of
Tablas Island, 700 fms]
= Acanthephyra media var. obliquirostris De Man, 1916: 150. [017.6'S 12914.5'E, Halmaheira Sea, 1855 m]
Acanthephyra pelagica (Risso, 1816)
= A.[lpheus] Pelagicus Risso, 1816: 91; Plate 2, fg. 7. [la mer de Nice; neotype listing by Fransen,
Holthuis & Adema, 1997 not valid]
= Ephyra Haeckelii von Martens, 1868: 52; Plate 1, Figs 7a-b. [Messina]
= Acanthephyra Agassizzi mediterranea Riggio, 1900: 20. [Mare di Messina]
= Acanthephyra rectirostris Riggio, 1900: 20. [Mare di Messina]
= Acanthephyra purpurea var. multispina Coutire, 1905a: 10. [Princesse-Alice stns 1797 (3218'N
2358'W), 0-2000 m; 1851 (4615'N 709'W), 0-3000 m; 1991 (4253'N 822'E), 0-2000 m; 1583 (4736'N
738'W, 1490 m) and 1639 (3617'N 2853'W, 0-3000 m)]
= Acanthephyra parva multidens Coutire, 1905a: 15; Fig. 5. [Princesse-Alice stns 1856 (3646'N
2641'W), 0-3250 m; 1834 (3728'N 2553'30"W), 0-1000 m]
Acanthephyra prionota Foxton, 1971: 35; Figs 1-2. [East African coast, 0539'S 4622'E, 1900-1850-(0) m]
Acanthephyra purpurea A. Milne-Edwards, 1881a: 933. [au large des les Berlingues par 2.590 mtres]
= Miersia Agassizii Smith, 1882: 67; Plate 11, Figs 5-7; Plate 12, Figs 1-4. [Blake stns 305, 4123'15''N
6551'25''W, 810 fms; 323, 3319'0''N 7612'30''W, 457 fms; 330, 3141'0''N 7435'0''W, 1047 fms]
= Acanthephyra abyssorum Filhol, 1885b: Plate 7. [Type locality not indicated, 4000 m]
= Acanthephyra parva paucidens Coutire, 1905a: 16; Fig. 5(3). [Princesse-Alice stn 1856 (3617'N,
2853'W), 0-3250 m]
= Hoplocaricyphus similis Coutire, 1907a: 7; Fig. 1. [2804'N, 1649'30"W, 0-1000 m]
Acanthephyra quadrispinosa Kemp, 1939: 576. [Indo-Pacifc from the E. African coast to 163W, and from
25N to 42S; South Atlantic from 32S to 40S]
= Acanthephyra batei Stebbing, 1905: 107; Plate 24B; nec Faxon, 1895. [Cape Point Lighthouse, S 83E,
35.5 miles, 360 fms]
Acanthephyra rostrata (Spence Bate, 1888)
= Hymenodora rostrata Spence Bate, 1888: 846; Plate 136, fg. 4. [Challenger stns 184, 128'S 14510'E,
near Torres Strait, 1400 fms; 205, 1642'N 11922'S, Philippine Islands, 1050 fms; 245, 3623'N
17431'E, North Pacifc, 2775 fms]
Acanthephyra sanguinea Wood-Mason in Wood-Mason & Alcock, 1892: 358. [Laccadive Sea, 953'34''N
7516.5'E, 1091 fms; Laccadive Sea, 823'N 7547'E, 738 fms; Bay of Bengal, 1258'N 8852'17''E,
1748 fms]
= Acanthephyra kempii Balss, 1914a: 595. [7001'N 8556'E, 0-2500 m]
Acanthephyra sibogae De Man, 1916
= Acanthephyra (Meningodora) sibogae De Man, 1916: 149. [526'S 12118'E, entrance of Gulf of Boni,
1944 m]
Acanthephyra sica Spence Bate, 1888: 739; Plate 125, fg. 1. [Challenger stns 168, 4028'S 17743'E, of
New Zealand, 100 fms; 40, 3451'N 6830'W, north-west of Bermuda, 2675 fms; 159, 4725'S
13022'E, south of Australia, 2150 fms; 169, 3734'S 17922'E, near New Zealand, 700 fms; 170,
2955'S 17814'W, of the Kermadec Islands, 520 fms; 170A, 2945'S 17811'W, near the Kermadec
Islands, 630 fms; 181, 1350'S 15149'E, between Australia and the Solomon Islands, 2440 fms; 194,
434'0''S 12957'30''E, of Banda Island, 200 fms; 230, 2629'N 13757'E, south of Japan, 2425 fms;
235, 347'N 1380'E, south of Japan, 565 fms; 318, 4232'S 5629'W, north of the Falkland Islands,
2040 fms]
Acanthephyra smithi Kemp, 1939: 577. [Indo-Pacifc from the East African coast to mid-Pacifc in 131W;
at the western end of its range it extends to 14S and at the eastern end from 20N to 24S]
Acanthephyra stylorostratis (Spence Bate, 1888)
= Bentheocaris stylorostratis Spence Bate, 1888: 726; Fig. 72; Plate 123, fg. 4. [Challenger stn 13, 2138'N
4439'W, Mid North Atlantic, 1900 fms]
? = Bentheocaris exuens Spence Bate, 1888: 724; Plate 123, fg. 3. [Challenger stn 285, 3236'S 13743'W,
South Pacifc Ocean, 2357 fms]
Acanthephyra tenuipes (Spence Bate, 1888)
= Tropiocaris tenuipes Spence Bate, 1888: 836; Plate 136, fg. 2. [Challenger stn 184, 128'S 14510'E, 1400
fms]
= Acanthephyra gracilipes Chace, 1940: 149; Figs 26-27. [Bermuda Oceanographic Expedition Net 689
or 734 (approx. 3212'N 6436'W), 1000 fms]
Acanthephyra trispinosa Kemp, 1939: 577. [west coast of Central America from 7N to 4S, extending
westwards to 116W]
Ephyrina Smith, 1885a
= Ephyrina Smith, 1885a (type species Ephyrina Benedicti Smith, 1885a, by monotypy, gender femi-
nine)
= Calymarina Spence Bate, 1888 (nomen nudum, gender feminine)
= Tropirinus Spence Bate, 1888 (nomen nudum, gender masculine)
= Tropiocaris Spence Bate, 1888 (type species Tropiocaris planipes Spence Bate, 1888, a junior subjective
synonym of Ephyrina Benedicti Smith, 1885a, gender feminine)
Ephyrina benedicti Smith, 1885a
= Ephyrina Benedicti Smith, 1885a: 506. [Albatross stn 2083, 4026'40''N 675'15''W, 959 fms]
= Tropiocaris planipes Spence Bate, 1888: 835; Plate 136, fg. 1. [Challenger stn 230, 2629'N 13757'E,
2425 fms]
Ephyrina bifda Stephensen, 1923: 58; Fig. 18. [4537'N 703'W, 4300 m of water]
Ephyrina childressi Chace, 1986: 34; Fig. 17. [Halmahera Sea, Indonesia, 033'42"S 12852'06"E, 950-1200 m]
Ephyrina fgueirai fgueirai Crosnier & Forest, 1973
= Ephyrina fgueirai Crosnier & Forest, 1973: 73; Figs 20b, 21g-h, 22c-d, 23. [Bay of Biscay, 4712'30"N
648'W, 2120-2350 m]
Ephyrina fgueirai spinicauda Lin & Chan, 2001: 184; Figs 1, 2a, d-e, 3. [N.E. Taiwan, I-Lan County, fshing
port at Tai-Shi, about 600 m]
Ephyrina hoskynii Wood-Mason & Alcock, 1891b
= Ephyrina Hoskynii Wood-Mason & Alcock, 1891b: 194. [Investigator stn 105 (Arabian Sea, 152'N
7234'E), 740 fms]
Ephyrina ombango Crosnier & Forest, 1973: 68; Figs 20a, 21a-f, 22a-b. [of So Tom, Gulf of Guinea,
030'N 630'E, 0-1000 m in total depth of 2900 m]
Heterogenys Chace, 1986
= Heterogenys Chace, 1986 (type species Acanthephyra microphthalma Smith, 1885a, by original desig-
nation and monotypy, gender feminine)
Heterogenys microphthalma (Smith, 1885a)
= Acanthephyra microphthalma Smith, 1885a: 502. [Albatross stn 2224, 3616'30"N 6821'00"W, 2574 fms]
= Acanthephyra longidens Spence Bate, 1888: 735; Plate 124, fg. 4. [Challenger stns 198, 255'N 12453'E,
near the Philippine Islands, 2150 fms; 285, 3236'S 13743'W, South Pacifc Ocean, 2375 fms]
Heterogenys monnioti Crosnier, 1987b: 704; Fig. 3. [2950.9'S 4835.5'E, 3668-3800 m]
Hymenodora G.O. Sars, 1877
= Hymenodora G.O. Sars, 1877 (type species Pasipha glacialis Buchholz, 1874, by monotypy, gender
feminine)
Hymenodora acanthitelsonis Wasmer, 1972: 87; Figs 1-8. [4518.0'N 12543.2'W to 4517.2'N 12548.3'W,
0-2560 m]
Hymenodora frontalis Rathbun, 1902a: 904. [west of Unalaska, 322 fms]
Hymenodora glacialis (Buchholz, 1874)
= Pasipha glacialis Buchholz, 1874: 279; Plate 1, Figs 2-2g. [an der Oberfche des Meeres, also in der
Nhe des 74. nrdl. Br. in betrchtlicher Entfernung von der Grenze des Packeises]
= Hymenodora glauca Spence Bate, 1888: 847; Plate 137, fg. 1. [Challenger stns 159, 4725' 13022'E,
south of Australia, 2150 fms; 215, 419'N 13015'E, south of the Philippines, 2550 fms]
= Hymenodora mollicutis Spence Bate, 1888: 848; Plate 137, fg. 2. [Challenger stns 104, 225'N 201'W,
Atlantic, 2500 fms; 87, 2549'N 2055'W, near the Canary Islands, 1675 fms; 133, 3541'S 2055'W,
near Tristan da Cunha, 1900 fms; 156, 6226'S 9544'S, 1975 fms; 157, 5355'S 10835'S (sic), 1950 fms;
318, 4232'S 5629'W, 2040 fms]
Hymenodora gracilis Smith, 1886a: 680; Plate 12, fg. 6. [Albatross stns 2036, 3852'40''N 6924'40''W, 1735
fms; 2083, 4026'40''N 6705'15''W, 959 fms; 2095, 3929'00''N 7058'40''W, 1342 fms; 2099, 3712'20''N
6936'00''W, 2949 fms; 2100, 3922'00''N 6834'430''W, 1628 fms; 2101, 3918'30''N 6824'00''W, 1686
fms; 2116, 3545'23''N 7431'25''W, 888 fms; 2182, 3925'30''N 7144'00''W, 861 fms; 2193, 3944'30''N
7010'30''W, 1122 fms]
Kemphyra Chace, 1986
= Kemphyra Chace, 1986 (type species Notostomus corallinus A. Milne-Edwards, 1883, by original
Kemphyra corallina (A. Milne-Edwards, 1883)
= Notostomus corallinus A. Milne-Edwards, 1883: Plate 32. [Travailleur dragage 20 (4130'N 1140'W),
2080 m]
= Acanthephyra valdiviae Balss, 1914a: 595. [306'S 8750'E, 0-2070 m]
Meningodora Smith, 1882
= Meningodora Smith, 1882 (type species Meningodora mollis Smith, 1882, by monotypy, gender fem-
inine)
Meningodora compsa (Chace, 1940)
= Notostomus compsus Chace, 1940: 156; Figs 31-32. [Bermuda Oceanographic Expedition Net 748
(3212'N 6436'W), 900 fms]
Meningodora longisulca Kikuchi, 1985: 191; Figs 1-14. [2308.1'N 14933.8'E, 0-975 m]
Meningodora marptocheles (Chace, 1940)
= Notostomus marptocheles Chace, 1940: 158; Figs 33-34. [N.E Providence Channel, Bahamas, 2529'N
7718'W, 875 fms of wire]
Meningodora miccyla (Chace, 1940)
= Notostomus miccylus Chace, 1940: 161; Figs 35-36. [north of Litle Inagua Island, Bahamas, 2144'N
7243'W, 1167 fms of wire]
Meningodora mollis Smith, 1882: 74; Plate 11, Figs 8-9; Plate 12, Figs 5-9. [Blake stn 328, 3428'25"N
7522'50"W, 1632 fms]
= Notostomus fragilis Faxon, 1893: 207. [Albatross stn 3371 (North Pacifc Ocean, Costa Rica, 526'20"N,
8655'00"W), 770 fms]
Meningodora vesca (Smith, 1886a)
= Notostomus viscus Smith, 1886b: 189. [nomen nudum]
= Notostomus vescus Smith, 1886a: 676; Plate 12, fg. 5. [Albatross stn 2099, 3712'20"N 6939'00"W,
2949 fms]
= Acanthephyra brevirostris Spence Bate, 1888: 751; Plate 126, Figs 5-6; nec Smith, 1885a. [Challenger
stn 107, 122'N 2636'W, Atlantic, south-west of Sierra Leone, 1500 fms]
= Acanthephyra batei Faxon, 1895: 167. [nomen novum for A. brevirostris Spence Bate, 1888]
= Acanthephyra parvirostris Coutire, 1911a: 157. [erroneous spelling of A. brevirostris]
Notostomus A. Milne-Edwards, 1881b
= Notostomus A. Milne-Edwards, 1881b (type species Notostomus gibbosus A. Milne-Edwards, 1881b,
by monotypy, gender masculine)
Notostomus auriculatus Barnard, 1950: 670; Fig. 124h-i. [of Cape Point, 800 fms]
Notostomus crosnieri Macpherson, 1984: 54; Figs 6a, 7a, 8a-b. [0233.5'S, 0543.2'E, 4088 m]
Notostomus distirus Chace, 1940: 166; Figs 39-40. [Bermuda Oceanographic Expedition Net 1281 (3212'N
6436'W), 1000 fms]
Notostomus elegans A. Milne-Edwards, 1881b: 8. [la mer des Antilles, 2436'N 8405'W, 955 brasses]
= Notostomus patentissimus Spence Bate, 1888: 826; Plate 133; Plate 134, Figs 1-2. [Challenger stn 198,
255'N 12453'E, south of the Philippine Islands, 2150 fms]
= Notostomus longirostris Spence Bate, 1888: 833; Plate 135; fg. 4. [Challenger stn 195, 421'S 1297'E,
of Banda Island, 1425 fms]
= Notostomus westergreni Faxon, 1893: 208. [Albatross stn 3399 (North Pacifc Ocean, Ecuador, 107'N,
8104'W), 1740 fms]
= Notostomus atlanticus Lenz & Strunck, 1914: 330. [2041'N 3153'W, 3000 m, Westich von den Kap-
verden und Kanarischen Inseln]
Notostomus gibbosus A. Milne-Edwards, 1881b: 7. [Grenade, 626 brasses]
= Notostomus perlatus Spence Bate, 1888: 831; Plate 135, fg. 2. [Challenger stn 198, 255'N 12453'E,
near the Philippine Islands, 2150 fms]
= Notostomus brevirostris Spence Bate, 1888: 832; Plate 135, fg. 3. [Challenger stn 120, 837'S 3428'W,
of Pernambuco, 675 fms]
Notostomus japonicus Spence Bate, 1888: 830; Plate 135, fg. 1. [Challenger stn 235, 347'N 1380'E, south of
Japan, 565 fms]
Notostomus murrayi Spence Bate, 1888: 829; Plate 134, fg. 3. [Challenger stn 133, 3541'S 2055'W, near
Tristan da Cunha, 1900 fms]
Notostomus robustus Smith, 1884: 377; Plate 7, fg. 2. [Albatross stns 2042, 3933'N 6826'45"W, 1555 fms;
2074, 4143'N 6521'50"W, 1309 fms]
= Notostomus beebei Boone, 1930a: 39; Fig. 6. [fve miles south of Nonsuch Island, Bermuda, 3216'N
6436'W, 900 fms]
Notostomus sparsidenticulatus Wasmer, 1986: 44; Fig. 9. [4440'S 14526'W, 2050-0 m]
Family OPLOPHORIDAE Dana, 1852a
Janicella Chace, 1986
= Janicella Chace, 1986 (type species Oplophorus spinicauda A. Milne-Edwards, 1883, by original des-
Janicella spinicauda (A. Milne-Edwards, 1883)
= Oplophorus spinicauda A. Milne-Edwards, 1883: Plate 30. [of Casablanca, Morocco, 3413'30"N
743'W, 636 m]
= Oplophorus foliaceus Rathbun, 1906: 922; Fig. 72; Plate 20, fg. 8. [Kaiwi Channel, 337-442 fms]
= Acanthephyra anomala Boone, 1927: 104; Fig. 21. [north of Glover Reef, 484 fms]
Oplophorus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]
= Oplophorus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840] (type species O.[plophorus]
typus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840], by monotypy, gender masculine)
Oplophorus gracilirostris A. Milne-Edwards, 1881b: 6. [la Dominique, 118 brasses] (Fig. 14)
= Oplophorus longirostris Spence Bate, 1888: 765; Plate 127, fg. 2. [Challenger stn 174c, 1907'50"S
17819'35"E, of Kandavu, Fii Islands, 610 fms]
Fig. 14. Oplophorus gracilirostris A. Milne-Edwards, 1881. Photo by Tin-Yam Chan.
= Hoplophorus Smithii Wood-Mason & Alcock, 1891b: 194 [Investigator stns 62 (1644'30"N, 8819-32'E),
1439 fms; 103 (1515'N, 8109'E), 1260 fms]
= Oplophorus okitsuensis Yokoya, 1922: 302. [Okitsu, Suruga Bay]
Oplophorus novaezeelandiae (De Man, 1931)
= Hoplophorus (Oplophorus) nov-zeelandi De Man, 1931: 369; Figs 1-20. [of Kaikoura, on the east
coast of the South Island of New Zealand, 2 fms]
Oplophorus spinosus (Brull, 1839)
= Palmon spinosus Brull, 1839: 18; vignete on page 3 (as Palmon pineux). [les Canaries]
= Hoplophorus Grimaldii Coutire, 1905a: 1; Fig. 1. [full description in Coutire, 1905a; 3218'N
2358'W, 0-2000 m]
Oplophorus typus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]
= O.[plophorus] typus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 424. [Nouvelle-
Guine]
= Oplophorus brevirostris Spence Bate, 1888: 766; Plate 127, fg. 3. [Challenger stn 207, 1221'N 12215'E,
of Tablas Island, 700 fms]
Systellaspis Spence Bate, 1888
= Systellaspis Spence Bate, 1888 (type species Systellaspis lanceocaudata Spence Bate, 1888, by original
= Hoplopasiphaea Yokoya & Shibata, 1965 (type species Hoplopasiphaea philippinensis Yokoya & Shi-
bata, 1965, by monotypy, gender feminine)
Systellaspis braueri braueri (Balss, 1914a)
= Acanthephyra Braueri Balss, 1914a: 594. [026'N 632'W, 0-4000 m; 056'N 434'W, 0-4000 m]
? = Systellaspis echinurus Coutire, 1911a: 158. [au large des ctes dEspagne et du Portugal entre les
parallles 36 et 45, de part et d'autre du 11
e
degr W]
= Systellaspis densispina Stephensen, 1923: 57; Fig. 17. [4630'N 700'W, > 4000 m]
Systellaspis braueri paucispinosa Crosnier, 1987c: 953; Fig. 3a-b. [3159'08"N, 15804'04"E, entre 1685 m de
profondeur et la surface]
Systellaspis cristata (Faxon, 1893)
= Acanthephyra cristata Faxon, 1893: 206. [Albatross stns 3361 (North Pacifc Ocean, Panama,
0610'00''N 08306'00''W), 1471 fms; 3381 (North Pacifc Ocean, Colombia, 456'00"N, 8052'30"W),
1772 fms]
Systellaspis curvispina Crosnier, 1987b: 711; Figs 6a-b, 8a-b. [Madagascar, 1322'S, 4717'38"E, 2000-0 m]
Systellaspis debilis (A. Milne-Edwards, 1881b)
= Acanthephyra debilis A. Milne-Edwards, 1881b: 13. [trouve une profondeur de 500 brasses dans
le canal de Bahama]
= Miersia gracilis Smith, 1882: 70, Plate 1, fgs 4-4d. [Blake stn 328, 3428'25"N 7522'50"W, 1632 fms]
= Acanthephyra debilis var. Europa A. Milne-Edwards, 1883: Plate 33, fg. 2. [Type locality not indi-
cated]
= Systellaspis Bouvieri Coutire, 1905a: 8; Fig. 3. [Princesse-Alice stn 1856, 3646'N 2641'W, 0-3250 m]
= Systellaspis debilis var. indica De Man, 1916: 151. [110.5'S 13009'E, Halmaheira Sea, 798 m]
= Hoplopasiphaea philippinensis Yokoya & Shibata, 1965: 4; Figs 4, 5. [1317'N 12550'E, 100 m]
Systellaspis eltanini Wasmer, 1986: 52; Figs 13-14. [South Atlantic, 3412'S 1635'E, 1550-0 m]
Systellaspis guillei Crosnier, 1987b: 718; Fig. 11. [Madagascar, 1304'S 4756'E, 1500-0 m]
Systellaspis intermedia Crosnier, 1987c: 948; Figs 1b, 2, 3d. [Ouest-Pacifque, au nord des les Marquises,
0-140E, chalutage plagique]
Systellaspis lanceocaudata Spence Bate, 1888: 758; Plate 124, fg. 7. [Challenger stn 232, 3511'N 13928'E, of
Japan, 345 fms]
Systellaspis pellucida (Filhol, 1884)
= Acanthephyra pellucida Filhol, 1884: 199. [2620'N 1453'W, 782 m, lectotype designation by Cros-
nier & Forest, 1973]
= Acanthephyra afnis Faxon, 1896: 162; Plate 2, fgs 1-3. [Blake stn 258 (Of Granada, 1203'15"N,
6146'25"W), 159 fms]
Superfamily ATYOIDEA De Haan, 1849 [in De Haan, 1833-1850]
Family ATYIDAE De Haan, 1849 [in De Haan, 1833-1850]
Antecaridina Edmondson, 1954
= Mesocaris Edmondson, 1935a (type species Mesocaris lauensis Edmondson, 1935a, by monotypy,
gender feminine; invalid junior homonym of Mesocaris Ortmann, 1893 (Crustacea Larvata))
= Antecaridina Edmondson, 1954 (nomen novum for Mesocaris Edmondson, 1935a, gender feminine)
Antecaridina lauensis (Edmondson, 1935a)
= Mesocaris lauensis Edmondson, 1935a: 13; Fig. 4. [cave containing brackish water, Island of Na-
muka, Lau islands, Fii]
Archaeatya Villalobos F., 1960a
= Archaeatya Villalobos F., 1960a (type species Archaeatya chacei Villalobos F., 1960a, by original des-
Archaeatya chacei Villalobos F., 1960a: 332; Figs 1-25. [Isla de Cocos, 8730'W 530'N]
Atya Leach, 1816a
= Atys Leach, 1816b (type species Atys scaber Leach, 1816b, by monotypy, gender masculine; name
placed on the Ofcial List of Rejected and Invalid Generic Names in Zoology in Opinion 470 in
1957)
= Atya Leach, 1816a (type species Atys scaber Leach, 1816b, gender feminine; name placed on the
Ofcial List of Generic Names in Zoology in Opinion 470 in 1957)
= Evatya Smith, 1871 (type species Evatya crassa Smith, 1871, by monotypy, gender feminine)
Atya abelei Felgenhauer & Martin, 1983: 333; Figs 1-4. [Panama, El Valle, Cocle Province, unnamed tribu-
tary of Rio Anton, 600 m elevation]
Atya africana Bouvier, 1904a: 138. [Samkita, dans la rivire Ogoou]
Atya brachyrhinus H.H.Jr. Hobbs & C.W.J. Hart, 1982: 35; Figs 1c, 12g, 15. [Coles Cave, Barbados, West
Indies, 1310'40"N 5934'3"W]
Atya crassa (Smith, 1871)
= Evatya crassa Smith, 1871: 95. [Fresh water streams, Polvon, and the Rio Fulva, two and a half
miles northwest of Realejo; both localities in the Occidental Department of Nicaragua]
Atya dressleri Abele, 1975a: 51; Figs 1-2. [Panam, Pacifc drainage, Veraguas Province, Ro Santa Mara
drainage, headwaters of Ro San Juan, about 15 km above Calobre, 566 m elevation]
Atya gabonensis Giebel, 1875: 52. [Gabon]
= Euatya sculptilis Koelbel, 1884: 317; Plate 2, fg. 8; Plate 3, fgs 1-8. [Orinoco]
= Atya sculptata Ortmann, 1890: 465. [Afrika, vielleicht aus West Afrika]
Atya innocous (Herbst, 1792)
= Cancer (Astacus) Innocous Herbst, 1792 [in Herbst, 1791-1796]: 62; Plate 28, fg. 3. [das vaderland is
unbekannt; considered to be Oceano Americano ad Martinicam, see H.H.Jr. Hobbs & C.W.J. Hart,
1982]
= Astacus 988 Gronovius, 1764: 231; Plate 17, fg. 6. [in Oceano Americano ad Martinicam; included
in a work rejected for nomenclatorial purposes in Opinion 261 in 1955]
= Astacus Nasoscopus Meuschen, 1778: 86. [nomen novum for Astacus 988 Gronovius, 1764; included
in a work rejected for nomenclatorial purposes in Opinion 260 in 1954]
= A.[tya] occidentalis Newport, 1847: 159. [West India Islands]
= Atya robusta A. Milne-Edwards, 1864: 148; Plate 3, fg. 1. [Nouvelle-Caldonie]
= Atya tenella Smith, 1871: 94, 95. [fresh water streams, Polvon, Occidental Department, Nicaragua]
Atya intermedia Bouvier, 1904a: 137. [Afrique occidentale, le Saint-Thomas]
Atya lanipes Holthuis, 1963a: 61; Figs 1-2. [St. Thomas, Virgin Islands]
Atya limnetes Holthuis, 1986a: 438; Figs 1-3. [Rio Anchicaya, Departemento del Valle del Cauca]
Atya margaritacea A. Milne-Edwards, 1864
= Atya Margaritacea A. Milne-Edwards, 1864: 148; Plate 3, fg. 2-2c. [Nouvelle-Caldonie (evidently
erroneous, most certainly originates from East American waters, see Holthuis, 1966)]
= Atya rivalis Smith, 1871: 94. [fresh water streams Polvon, Occidental Department, Nicaragua]
Atya ortmannioides Villalobos F., 1956: 459; Figs 1-6. [Ro de las Truchas, La Mira, 52.5 km S.SE de Ar-
teaga, Michoacn, 2.5 km de la costa, en la vertiente del Pacifco]
Atya scabra (Leach, 1816b)
= Atys scaber Leach, 1816b: 345. [Misantla, Estado de Veracruz, Mexico, 1956'N 9650'W; neotype
designation by H.H.Jr. Hobbs & C.W.J. Hart, 1982]
= Atya mexicana Wiegmann, 1836: 145. [Misantla]
? = A.[tya] sulcatipes Newport, 1847: 159; Plate 8, fg. 1. [in fresh water, San Nicolao, Cape Verd Islands]
= Atya punctata Kingsley, 1878a: 91, 92. [Hayti]
= Atya margaritacea var. claviger Aurivillius, 1898a: 14; Plate 3, Figs 5-8. [Kamerun, Etome, in Bchen]
= Atya margaritaria clavipes Holthuis, 1966: 234. [lapsus for Atya margaritacea var. claviger Aurivillius,
1898a]
Atyaephyra de Brito Capello, 1867
= Symethus Rafnesque, 1814 (type species Symethus fuviatilis Rafnesque, 1814, an invalid senior
subjective synonym of Hippolyte Desmarestii Millet, 1831, gender masculine; name suppressed for
the purposes of the Principle of Priority but not for those of the Principle of Homonymy and placed
= Acilius Rafnesque, 1815 (nomen novum for Symethus Rafnesque, 1814, gender masculine; name
suppressed for the purposes of the Principle of Priority for those of the Principle of Homonymy and
1958 and Opinion 619 in 1961)
= Symaethus Agassiz, 1846 [in Agassiz, 1842-1846] (invalid emendation of Symethus Rafnesque,
= Atyaephyra de Brito Capello, 1867 (type species Atyaephyra Rosiana de Brito Capello, 1867, by
monotypy, gender feminine; name placed on the Ofcial List of Generic Names in Zoology in Opin-
ion 522 in 1958)
Atyaephyra desmarestii (Millet, 1831)
= Hippolyte Desmarestii Millet, 1831: 56; Plate 1, fgs 1-1b. [les eaux de la Mayenne, de la Sarthe, du
Loir, du Thouet et du Layon]
= Symethus fuviatilis Rafnesque, 1814: 23. [dans les ruisseaux et les mares; name suppressed under
the plenary powers for the purposes of the Principle of Priority but not for those of the Principle of
Homonymy in Opinion 522 in 1958]
= Atyaphyra Desmaresti var. occidentalis Bouvier, 1913a: 72. [le Nord de lAfrique depuis la Tunisie,
tout le Sud de lEurope depuis la Macdonie]
= Atyaephyra rosiana de Brito Capello, 1867: 6; Plate 1, fgs 1-1e. [So Barnabe River, Algarve, Portu-
gal; neotype selection by Anastiadou, Kitsos & Koukouras, 2008]
Atyaephyra orientalis Bouvier, 1913a
= Atyaphyra Desmaresti var. orientalis Bouvier, 1913a: 72. [Syrie]
= Atyaephyra desmarestii mesopotamica Al-Adhub, 1987: 1; Fig. 1. [Shat Al-Arab river]
Atyaephyra stankoi Karaman, 1972
= Atyaephyra desmarestii stankoi Karaman, 1972: 82; Figs 3, 6, 9-10. [Mazedonien, Vardar Flussystem]
Atydina Cai, 2010a
= Atydina Cai, 2010a (type species Caridina atyoides Nobili, 1900a, by original designation, gender
feminine)
Atydina atyoides (Nobili, 1900a)
= Caridina atyoides Nobili, 1900a: 478. [Si Oban; Sioban, Pulau Sipura, Kepulauan Mentawei, of west
coast of Sumatra, Indonesia, according to Cai, 2010a]
Atyella Calman, 1906a
= Atyella Calman, 1906a (type species Atyella brevirostris Calman, 1906a, by original designation, gen-
der feminine; name placed on the Ofcial List of Generic Names in Zoology in Opinion 470 in 1957)
Atyella brevirostris Calman, 1906a: 201; Plate 14, fgs 57-64. [Mbete; near mouth of Lofu; Kala; rock-pool
at Kasakalawe (all Lake Tanganyika)]
Atyella longirostris Calman, 1906a: 202; Plate 14, fgs 65-72. [Mbete; Kala (both Lake Tanganyika)]
Atyoida Randall, 1840
= Atyoida Randall, 1840 (type species A.[tyoida] bisulcata Randall, 1840, by monotypy, gender femi-
nine)
= Ortmannia Rathbun, 1901 (type species Ortmannia henshawi Rathbun, 1901, a junior subjective
synonym of A.[tyoida] bisulcata Randall, 1840, gender feminine)
= Pseudatya Roux, 1928a (type species Pseudatya beauforti Roux, 1928a, a junior subjective synonym
of A.[tya] pilipes Newport, 1847, by monotypy, gender feminine)
= Vanderbiltia Boone, 1935 (type species Vanderbiltia rosamondae Boone, 1935, a junior subjective syn-
onym of A.[tya] pilipes Newport, 1847, by original designation and monotypy, gender feminine)
Atyoida bisulcata Randall, 1840
= A.[tyoida] bisulcata Randall, 1840: 140; Plate 5, fg. 5. [Sandwich Islands]
= Ortmannia henshawi Rathbun, 1901: 120. [Kaiwiki, Hilo, Hawaii, 1800f altitude, 3 miles from the sea]
Atyoida pilipes (Newport, 1847)
= A.[tya] pilipes Newport, 1847: 160. [Apia, Upoln, New Zealand]
= Atyoida tahitensis Stimpson, 1860a: 28. [in aquis dulcibus insulae Tahiti]
= Caridina acuminata Stimpson, 1860a: 29. [stream of Naka-kaigan coast, 60-70 m above sealevel,
Chichi-jiam Island, Ogasawar Islands; neotype designation by Cai et al., 2006]
= Caridina brevirostris Stimpson, 1860a: 29. [River Aritsu, Kume-jima Island; neotype designation by
Cai et al., 2006]
= Atya brevirostris De Man, 1892a: 360; Plate 21, fgs 21-21d. [Fluss Wukur bei Sikka; Fluss Lella bei
Sikka; Fluss bei Mbawa unterhalbs des Wasserfalls (all Flores); Fluss Koinino bei Kupang (Timor)]
= Pseudatya beauforti Roux, 1928a: 209; Figs 1-9. [Batjan]
= Vanderbiltia rosamondae Boone, 1935: 160; Plates 41-42. [Venus Point Reef, Tahiti, Society Islands, in
coral]
Fig. 15. Atyopsis moluccensis (De Haan, 1849). Photo by Tin-Yam Chan.
= Vanderbiltia mirabilis Holthuis, 1953a: 114. [manuscript name published as a synonym]
Atyoida serrata (Spence Bate, 1888)
= Atya serrata Spence Bate, 1888: 699; Plate 119, fg. 2. [Valley of San Antonia, San Iago, Cape Verde
Islands from a fresh-water stream; probably erroneous, see Chace, 1983a]
= Atya serrata mutation Alluaudi Bouvier, 1904b: 448. [un torrent de la montagne dAmbre, Mada-
gascar; lle Bourbon; dans les ravines des montagnes de Salasie et dHelbour; lle Maurice]
Atyopsis Chace, 1983a
= Atyopsis Chace, 1983a (type species A.[tya] spinipes Newport, 1847, by original designation, gender
feminine)
Atyopsis moluccensis (De Haan, 1849 [in De Haan, 1833-1850]) (Fig. 15)
= Atya moluccensis De Haan, 1849 [in De Haan, 1833-1850]: 186; Plate O. [Moluccas, Indonesia; lec-
totype designation by Holthuis, 1993b]
= Atya armata A. Milne-Edwards, 1864: 149; Plate 3, fgs 3-3a. [Batavia (p. 145, 152), not les les Phil-
ippines (p. 149), see Chace, 1983a]
= Atya gustavi Ortmann, 1890: 467; Plate 36, fg. 9a-c. [Sumatra, Indrapura-Fluss]
= Atya lineolata De Man, 1892a: 357 (footnote). [manuscript name atributed to Kulh, cited as syno-
nym of Atya moluccensis]
Atyopsis spinipes (Newport, 1847)
= A.[tya] spinipes Newport, 1847: 159. [Philippine Islands]
= Atya dentirostris Thallwitz, 1891: 101. [Nord-Celebes; fully described in Thallwitz, 1892]
? = Atya brevirostris var. De Mani Nobili, 1900a: 475, fg. 1. [Fiume Sereinu, Isole Mentawei]
Australatya Chace, 1983a
= Australatya Chace, 1983a (type species Atya striolata McCulloch & McNeill, 1923, by original des-
Australatya striolata (McCulloch & McNeill, 1923)
= Atya striolata McCulloch & McNeill, 1923: 55; Plate 9, fgs 3-4. [Nortons Basin, Nepean River, New
South Wales]
Caridella Calman, 1906a
= Caridella Calman, 1906a (type species Caridella cunningtoni Calman, 1906a, by original designation,
gender feminine; name placed on the Ofcial List of Generic Names in Zoology in Opinion 470 in
1957)
Caridella cunningtoni Calman, 1906a: 199; Plate 13, fgs 45-52. [Kala, on rocks, shallow water; Kirando, 8
fms; Utinta, 15 fms (all Lake Tanganyika)]
Caridella minuta Calman, 1906a: 200; Plate 13, fgs 53-56. [near mouth of Lofu, on rocks, shallow water;
Karema, tow-neting, surface (both Lake Tanganyika)]
Caridella paski Calman, 1928: 739; Figs 1-3. [Kigoma, east side of Lake Tanganyika]
Caridina H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]
= Caridina H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840] (type species Caridina typus H.
Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840], by monotypy and by indication under
Article 68c of the ICZN, gender feminine; name placed on the Ofcial List of Generic Names in
Caridina ablepsia Guo, Jiang & Zhang, 1992: 4; Figs 1-9. [cave river, Wangcun Village, 10956'E 2846'N,
Yongshun County, Hunan Province]
Caridina acuta Liang, Chen & W.-X. Li, 2005: 530; Figs 15-29. [caves of Libo County (2541'N 10783'E),
Maolan Nature Sanctuary of Guizhou]
Caridina acutirostris Schenkel, 1902: 496; Plate 8, fgs 3a-c. [Sden des Posso-See]
Caridina africana Kingsley, 1883: 127; Plate 1, fgs 3-3a. [Zulu Mission, South Africa]
= Caridina africana forme typica Bouvier, 1925: 213; Fig. 470. [pays des Zoulous]
= Caridina wyckii var. paucipara Weber, 1897: 168. [River Umhlasine, Natal; lectotype designation by
Richard & Clark, 2010a]
Caridina alba J. Li & S. Li, 2010: 19; Figs 2-5. [Tenglongdong cave, Lichuan city, Enshi Autonomous Pre-
fecture, Hubei province, China]
Caridina alphonsi Bouvier, 1919: 330. [localit inconnue; austro-malaise according to Bouvier, 1925]
Caridina amnicolizambezi Richard & Clark, 2009: 45; Figs 22-23. [Angola, Rio Lucoge, branch of the Chi-
capu River, 7336'S 2031'E]
Caridina amoyensis Liang & Yan, 1977: 219; Figs 1-4. [Xiamen, suburabm rivulet, Fujian Province]
Caridina angulata Bouvier, 1905c: 84; Fig. 8. [rivire Ranofotsy, prs de Fianarantsoa]
Caridina angustipes Guo & Liang, 2003: 45; Figs 1A-C. [Wentang Village, Xinhua County, Hunan Prov-
ince, 400 m]
Caridina anislaq Cai, Choy & Ng, 2009: 82; Fig. 11. [Spring Anislaq, outside Cave Valencia, Valencia]
Caridina annandalei Kemp, 1918a: 96; Plate 25, fgs 6-15. [Inl Lake]
Caridina apodosis Cai & N.K. Ng, 1999: 1627; Figs 14-17. [mountain stream near Taitong village, New
Territory, Hong Kong]
Caridina appendiculata Jalihal & Shenoy, 1998: 128. [River Bari, Flores; lectotype designation by Cai & Ng,
2007]
Caridina aruensis Roux, 1911: 82. [ruisseau Matora, Soungi Manoumbai, Arou; ruisseau Panoua Bori,
Soungi Manoumbai, Arou]
Caridina babaulti babaulti Bouvier, 1918
= Caridina Babaulti Bouvier, 1918: 388; Figs 4-6. [Mukhi; Majghaon; Rajadhar]
Caridina babaulti basrensis Al-Adhub & Hamzah, 1987: 225; Fig. 1. [small channels connected to Shat al-
Arab River, Iraq]
Caridina babaultioides babaultioides Y, 1938
= Caridina babaultioides Y, 1938: 301; Figs 13-14. [small mountain stream, at an altitude of about 910
m, at Shui-tuan-pa near Yen-tsing city, Yunnan Province]
Caridina babaultioides angustifolia Cai & Yuan, 1996
= Caridina angustifolia Cai & Yuan, 1996: 385; Figs 14-15. [Wafang stream, near Jinsha village]
Caridina babaultioides emeica Liang, 2004: 218; Fig. 105. [Emeishan, Sichuan]
Caridina babaultioides phyllopoda Huang, 1984: 1; Figs 1-8. [Kwan Shien, Sichuan Province]
= Caridina phyllopoda Huang, 1984: 1; Figs 1-8. [Kwan Shien, Sichuan Province]
Caridina bakoensis Ng, 1995a: 192; Figs 8-10. [Middle stretch of Sungai Serait]
Caridina bamaensis Liang & Yan, 1983a: 252; Figs 1-10. [Bama County, Guangxi, Southern China]
Caridina baojingensis Guo, He & Bai, 1992: 609; Figs 1-9. [China, Hunan Province, Baojing County, Qing-
shui village]
Caridina batuan Cai, Choy & Ng, 2009: 80; Fig. 10. [Cave Castigio, Batuan]
Caridina belazoniensis Richard & Clark, 2009: 33; Figs 14-15. [Kenya, Mbelazoni, Lower Ahti River, Sa-
baki Estuary]
Caridina boholensis Cai, Choy & Ng, 2009: 75; Figs 7-8. [Quila Cave, Nueva Vida Norte, Batuan]
Caridina brachydactyla De Man, 1908a
= Caridina nilotica var. brachydactyla De Man, 1908a: 269; Plate 20, fgs 8a-c. [Indonesia, by Reo, Flo-
res; lectotype designation by Richard & Clark, 2010a]
= Caridina nilotica var. natalensis De Man, 1908a: 262; Plate 20, fgs 3-3b [South Africa, Umgeni River,
Durban, Natal; lectotype designation by Richard & Clark, 2010a]
= Caridina nilotica var. brevidactyla Roux, 1920: 320. [Aru-Inseln: Bach Matora, am Sungi Manumbai;
Bach Panua Bori am Sungi Manumbai; Bach bei Papakula, Kobroor; Wokamar, Wokam; Samang;
Seltuti, Kobroor; Dobo, Wammer; Pobdietur, Terangan; Udjir]
Caridina breviata N.K. Ng & Cai, 2000: 171; Figs 4-6. [Zhapu village, Yangjiang County, Guangdong Prov-
ince, southern China]
Caridina brevicarpalis De Man, 1892a: 397; Plate 24, fgs 30-30d. [Celebes, aus einem Flusse bei Palopo,
Luwu]
Caridina brevispina Liang & Yan, 1986a: 203; Fig. 5. [Tongren County, Guizhou Province]
Caridina bruneiana Choy, 1992: 49; Figs 1-4. [Negara Brunei Darussalam, on the upper reaches of Tembu-
rong River at Batang Duri, 0436'05''N 11506'45''E, altitude approx. 33 m]
Caridina buehleri Roux, 1934a
= Caridina bhleri Roux, 1934a: 219; Figs 1-5. [Bimoun, cte occidentale Nouvelle-Irlande]
Caridina buergersi Karge, von Rintelen & Klotz, 2010: 146; Figs 5-6. [Papua New Guinea, former
Deutsch-Neuguinea, near Manderberg]
Caridina buhi Cai & Shokita, 2006a: 253; Figs 3-4. [Binahugan River, Buhi, Camarines Sur, Luzon, Philip-
pines]
Caridina bunyonyiensis Richard & Clark, 2005: 715; Figs 5-6. [Uganda, Lake Bunyonyi]
Caridina burmensis Cai & Ng, 2000: 941; Fig. 7. [Myitkyina, Myanmar]
Caridina caerulea von Rintelen & Cai, 2009: 428; Figs 51-53. [Lake Poso, west shore, Cape Bancea,
0159.023'S 12035.108'E, on rocks in shallow water]
Caridina calmani Bouvier, 1919
= Caridina Calmani Bouvier, 1919: 334. [Ambatoubavara, Madagascar]
= Caridina Bouvieri Roux, 1929: 312; Figs 7-12. [Manjakatompo, altit. 1940 m, ruisseau descendant de
lAnkaratra]
Caridina camaro Cai, Choy & Ng, 2009: 82; Fig. 12. [Cave Camaro, Batuan]
Caridina cantonensis Y, 1938: 290; Figs 7-8. [Qingao vilage, Nanao County, Guangdong Province, Chi-
na; neotype designation by Cai & N.K. Ng, 1999]
= Caridina mutata Cai & N. K. Ng, 1999: 1624; Figs 12-13. [mountain stream near Fangcheng Town,
Fangcheng County, Guangxi Province, China]
Caridina caobangensis S.-Q. Li & Liang, 2002: 711; Figs 3-4. [PacBo Village, District Ha Quang, Cao Bang
province, Vietnam]
Caridina carli Roux, 1931a: 38; Figs 4-11. [Anaimalais, environs de Valparai, dans la rivire Naduar et ses
afuents]
Caridina cavaleriei cavaleriei Bouvier, 1919
= Caridina Cavalerii Bouvier, 1919: 332. [Gan-chouen-fou (Kouy-Tchou)]
Caridina cavaleriei industana Roux, 1931a: 35; Figs 1-3. [Aliyar Riv., prs de Malayandi Patanam, 6 milles
au Sud de Pollachi]
Caridina cavalerieioides Liu & Liang in Liang, 2004: 207; Fig. 100. [Malin Village, near Guiyang City,
Guizhou Province, China]
Caridina caverna Liang, Chen & W.-X. Li, 2005: 529; Figs 1-14. [caves of Libo County (2541'N 10783'E),
Maolan Nature Sanctuary of Guizhou]
Caridina cavernicola Liang & Zhou, 1993: 232; Figs 2(1)-2(2). [Lenggu Cave, Duan County, Guangxi Prov-
ince]
Caridina cebuensis Cai & Shokita, 2006a: 250; Figs 1-2. [spring water to Sagay River, Cebu Island, Philip-
pines]
Caridina celebensis De Man, 1892a
= Caridina serratirostris var. celebensis De Man, 1892a: 385; Plate 23; fgs 28f-h. [Celebes, Fluss bei
Palopo, Luwu]
= Caridina serratirostris koterai Kamita, 1951: 75; Figs A-J. [Shimoko, Iwami province, SE Hunshu,
Japan]
Caridina celestinoi Blanco, 1939a: 392; Plate 3, Figs 8-10. [mountain stream, Helosig, Leyte]
Caridina chauhani Chopra & Tiwari, 1949
= Caridina nilotica var. chauhani Chopra & Tiwari, 1949: 219; Figs 2-3 [tanks at Sale bhata, Chandan-
bhati, Bolangir, Salepali and Titilagarh; Ang River at Salebhata]
= Caridina williamsoni Jalihal, Shenoy & Sankolli, 1984: 1; Figs 1-3. [Sadhankeri tank, Dharwar]
Caridina chishuiensis Cai & Yuan, 1996: 379; Figs 8-10. [stream near Binan village]
= Caridina euryphylla Cai & Yuan, 1996: 382; Figs 11-13. [Qintan village, near Dongping village]
Caridina clavipes Guo & Liang, 2003: 47; Figs 1D-F. [Wentang Village, Xinhua County, Hunan Province]
Caridina clinata Cai, X.Q. Nguyn & Ng, 1999: 531; Figs 1-2. [ditch at Cuc Phuong National Park, Ninh
Binh Province, Northern Vietnam]
Caridina cognata De Man, 1915a: 397; Plate 28, fgs 3-3g, 4-4b [in einem kleinen Bache von sssem Wasser
in der Ksten-gegend zwischen der Humboldt-Bai und dem unteren Laufe des Tami-Flusses; in
Bchlein zwischen den kleinen Flssen Tjah and Jasa; aus dem Tjah-Flusse; aus dem kleinen
Tjano-Flusse bei Njao; Zoutbron]
Caridina confusa Choy & Marshall, 1997: 27; Figs 1a, 2i-j, 3b-c, 4a-e. [Thiaki Creek at Seamark Road cross-
ing, 1723.5'S 14532.5'E]
Caridina congoensis Richard & Clark, 2009: 48; Figs 24-25. [Congo, Bakou, in river, in total darkness, 30
m]
Caridina cornuta Liang & Yan, 1986a: 200; Fig. 3. [Zhiin County, Guizhou Province]
Caridina crassipes Liang, 1993: 23; Fig. 2. [Xianshui Cave, Pangshi village, Hunan]
Caridina crurispinata Gurney, 1984: 574; Figs 4-7. [Dark zone, la Grote dAntsatrobonko, Ankarana Mas-
sif, 40 km north of Ambilob, 60 km south of Diego Suarez, northern province of Diego Suarez]
Caridina cucphuongensis ng, 1980
= Caridina serrata cucphuongensis ng, 1980: 404; Fig. 230. [streams in Cucphuong area, Ninh Binh
Province, Northern Vietnam]
Caridina curta Liang & Cai, 2000: 179; Fig. 2. [Longquan spring in Huanglong (Yellow Dragon) Temple,
Jianshui County, Yunnan]
Caridina demani Roux, 1911: 94. [Tawarin]
Caridina demenica Cai & Li, 1997: 315; Fig. 1. [Demen Cave, near Yongkang village, Libo County of
Guizhou Province, southern China]
Caridina dennerli von Rintelen & Cai, 2009: 355; Figs 4-6. [Lake Matano, east shore, just at entrance to
outlet bay, 0231.54'S 12127.00'E]
Caridina dentifrons N.K. Ng & Cai, 2000: 167; Figs 1-3. [Baiin village, Huishui County, Guizhou Province,
southern China]
Caridina devaneyi Choy, 1991: 348; Figs 2-4. [stream at 8-mile Point near Suva, Viti Levu, Fii Islands]
Caridina dianchiensis Liang & Yan, 1985a: 196; Fig. 1. [Songhuaba Reservoir- Panlong River, Kunming,
Jiangchuan County, Yunnan province]
Caridina disjuncta Cai & Liang, 1999: 77; Fig. 4. [Shilin (Stone forest) village, Lunan County, central Yun-
nan]
Caridina disparidentata Liang, Yan & Wang, 1984: 8; Figs 1-12. [mountain stream, Qujing County, Yunnan
Province]
= Caridina heterodentata Liang & Yan, 1985a: 196. [unjustifed emendation of Caridina disparidentata
Liang, Yan & Wang, 1984; see Cai & Ng, 2001a]
Caridina ebuneus Richard & Clark, 2009: 39; Figs 18-19. [Ivory Coast, vicinity Abengourou]
Caridina edulis Bouvier, 1904a: 135. [Anantsahalankely]
Caridina elisabethae Karge, von Rintelen & Klotz, 2010: 142; Figs 3-4. [Papua New Guinea, Morobe, Her-
zog Mts., Bundun, 700-800 m, 0651.598'S 14637.07'E]
Caridina elliptica Cai & Yuan, 1996: 376; Figs 5-7. [stream near Panlong town]
Caridina elongapoda Liang & Yan, 1977
= Caridina nilotica elongapoda Liang & Yan, 1977: 220; Figs 5-8. [Xingzai, Zhangpu County, Fujian
Province]
Caridina endehensis De Man, 1892a
= Caridina brevicarpalis var. endehensis De Man, 1892a: 399; Plate 24, fg. 30e. [aus dem Flusse Ba bei
Endeh; Fluss bei Mbawa (both Flores)]
Caridina ensifera Schenkel, 1902: 490; Plate 8, fgs 1-11e. [Lake Poso, Celebes (Sulawesi), Indonesia; lecto-
type designation by Cai & Wowor, 2007]
Caridina evae Richard & Clark, 2009: 28; Figs 11-13. [Nigeria, Chubra division]
Caridina excavata Kemp, 1913b: 306; Plate 20, fgs 32-35; Plate 21, fgs 36-37. [backwater of the Rowta
River, Brahmaputra drainage system]
Caridina excavatoides Johnson, 1961: 127; Figs 3-11. [stream running between rubber plantations and rice-
felds, about nine miles from Alor Star, Kedah, on the Pokok Sena Road]
Caridina fasciata Hung, Chan & Yu, 1993: 489; Figs 1E, 6. [Ping Tong County, Heng Chun]
Caridina fecunda Roux, 1911: 95. [Lac Jamour]
Caridina feixiana Cai & Liang, 1999: 74; Figs 1-2. [stream in Feixia cave, Geju County, Yunnan, China]
Caridina fernandoi Arudpragasam & Costa, 1962: 8; Fig. 1. [shallow streamlets at Warakapola, and Seel-
angama, and also from shallow sheltered areas of the Maha Oya at Mawanella and the Deduru Oya
and the Magura Oya at Kurunegala, Sri Lanka]
Caridina fjiana Choy, 1983: 147; Fig. 1. [Fii, Viti Levu, Nadarivatu, Nukunuku Creek, 1735'40''S
17757'25''E, 640 m]
Caridina favilineata ng, 1975: 70; Fig. 5. [Nam Ha Province]
Caridina formosae Hung, Chan & Yu, 1993: 487; Figs 1D, 5. [Keelung, Pa Chih Men]
Caridina fossarum Heller, 1862b
= C.[aridina] fossarum Heller, 1862b: 411. [Schiraz]
? = Caridina syriaca Bouvier, 1904a: 132. [Syrie]
Caridina gabonensis Roux, 1927a: 239; Figs 1-7. [Lambarn, Gabon]
Caridina ghanensis Richard & Clark, 2009: 35; Figs 16-17. [Republic of Ghana, Pond Vume]
Caridina glaubrechti von Rintelen & Cai, 2009: 359; Figs 7-9. [Lake Towuti, west shore, south of Cape
Timbalo, 0242.91'S 12126.78'E]
Caridina glossopoda Liang, Guo & Gao, 1993: 41; Fig. 1. [Ala, Fenghuang County, Hunan, 2795'N
10965'E]
Caridina gordonae Richard & Clark, 2005: 717; Figs 7-8. [Uganda, Bufundi, Lake Bunyonyi]
Caridina gortio Cai & Anker, 2004: 247; Figs 8, 9a-f. [Can Gortio Cave, 1159'00''N 12453'00''E, altitude ca
200 m, Matalud, West Samar, Philippines]
Caridina gracilipes De Man, 1892a
= Caridina Wyckii var. gracilipes De Man, 1892a: 387; Plate 24, fgs 29e-k [aus dem Flusse von Maros;
Makassar; aus einem kleinen mit dem Meere nicht in Verbindung stehenden Bache zu Balangnipa;
Palima, aus dem Tjenrana; Pampanua, aus dem Tjenrana (all Celebes); Saleyer, aus dem Flusse
Bona]
= Caridina nilotica var. bengalensis De Man, 1908a: 265; Plate 20, fgs 6-6b. [Port Canning an Dhappa,
Calcuta]
Caridina gracilirostris De Man, 1892a: 399; Plate 25, fg. 31-31d. [river near Maros, Sulawesi (Celebes),
Indonesia; lectotype designation by Cai & Ng, 2007]
= Caridina pseudogracilirostris Thomas, V.K. Pillai & N.N. Pillai, 1976: 871; Fig. 1. [Cochin backwater]
Caridina gracillima Lanchester, 1901: 560; Plate 34, fg. 1. [Malay Peninsula; lectotype designation by Cai
& Ng, 2007]
Caridina grandirostris Stimpson, 1860a: 28 [upper stream of Tima River, about 4 km from river mouth,
Okinawa Island, Ruykyu Islands; neotype designation by Cai et al., 2006]
Caridina guangxiensis Liang & Zhou, 1993: 234; Figs 3(1)-3(2). [Dawangshan cave, Guilin City, Guangxi
Province]
Caridina gueryi Marquet, Keith & Kalfatak, 2009: 160; Figs 1-3. [unnamed river, Santo Island (Vanuatu),
15.29590 S 167.16 E]
Caridina guiyangensis Liang, 2002a: 119; Fig. 2. [China, Guizhou Province, Guiyang County]
Caridina gurneyi Jalihal, Shenoy & Sankolli, 1984: 29; Figs 12-14. [Malaprabha river, near old bridge,
Khanapur (Belgaum District)]
Caridina hainanensis Liang & Yan, 1983b: 211; Fig. 1. [Wencang County, Hainan Island, China]
Caridina hanshanensis Tan, 1990: 278; Figs 1-9. [Hanshan County, 3145'N 1187'E, Anhui Province]
Caridina harmandi Bouvier, 1906f
= Caridina Harmandi Bouvier, 1906f: 483. [Japon]
Caridina hodgarti Kemp, 1913b: 309; Plate 20, Figs 29-31; Plate 21, fgs 38-39. [Kobo, Abor county]
Caridina holthuisi von Rintelen & Cai, 2009: 364; Figs 10-12. [Lake Matano, south shore, canal between
island and mainland, 0228.46'S 12115.83'E]
Caridina hongyanensis Cai & Yuan, 1996: 373; Figs 2-4. [Yantanggou stream near Hongyan village]
Caridina hova Nobili, 1905c: 499. [Fort Dauphin]
Caridina huananensis Liang, 2004: 147; Fig. 69. [Guangdong]
Caridina hubeiensis Liang & S.-Q. Li, 1993: 213; Figs 1-9. [Badong, Hubei Province]
Caridina hunanensis Liang, Guo & Gao, 1993: 44; Fig. 3. [Zhonglian, Lengshuiiang City, Hunan, 277'N
1114'E]
Caridina imitatrix Holthuis, 1970: 100; Fig. 3. [Rivire Bleue near the bridge near Ounarou (Mont des
Sources); Marsh of Rivire Blanche, Ounarou region; Side branch of Rivire Blanche, near the for-
est road leading from Ounarou forestry station to the forestry log cabin on the western slope of Mt.
Poudihi, Ounarou region; same branch, near the log cabin; Thir River near the confuence with a
side branch, granite area near St. Louis, S.E. of Nouma; Rivire des Lacs, which empties in the Lac
en Huit, near highway to Nouma]
Caridina indistincta indistincta Calman, 1926
= Caridina indistincta Calman, 1926: 244; Fig. 3. [St. George District, Queensland]
Caridina indistincta sobrina Riek, 1953: 119; Fig. 9. [Rocky Creek, Fraser Island, Queensland]
Caridina isaloensis isaloensis Coutire, 1899b
= Caridina typus var. Isaloensis Coutire, 1899b: 383. [Fleuve Onilahy, cte Ouest]
= Caridina madagascariensis Bouvier, 1904a: 134. [rivire Fandramanona]
= Caridina brevirostris var. brevipes Bouvier, 1925: 230 (partim). [baie Amposindova, Madagascar]
Caridina isaloensis grandidieri Bouvier, 1904a
= Caridina Grandidieri Bouvier, 1904a: 133. [rivire Fandiamanana, Madagascar]
Caridina jalihali Mariappan & Richard, 2006: 47; Figs 22-24. [River Arani at Periapalayam; lake at Then-
neri; ponds at Maduranmangalam, Manimangalam, Redipalayam, RoshanNagar and Singaperu-
mal Koli]
= Caridina gurneyi lonavalensis Kadrekar & Sankolli, 1987: 60. [nomen nudum]
Caridina jeani Cai, 2010b: 80; Figs 1-2; nomen novum for Caridina typus var. brevirostris Roux, 1911. [Elat,
Kei Islands, Moluccas, Indonesia; lectotype designation by Cai, 2010b]
Caridina jiangxiensis Liang & Zheng, 1985
= Caridina pingi jiangxiensis Liang & Zheng, 1985: 319; Fig. 1(1b-7b). [Shixi stream, Fengxin County,
Jiangxi Province]
Caridina johnsoni Cai, Ng & Choy, 2007: 301; Figs 15-16. [Singapore, Lower Peirce Reservoir, north arm]
Caridina kaombefutilis Richard & Clark, 2010a: 331; Figs 14-15. [Malawi, Kaombe River]
Caridina kempi Jalihal, Shenoy & Sankolli, 1984: 13; Figs 6-8. [Hosayellapur tank (Hirekeri), Dharwar]
Caridina kilimae Hilgendorf, 1898: 35. [Marangu am Kilimandscharo]
Caridina kunmingensis Z.-Z. Wang & Liang, 2001: 303; Fig. 1. [Kunming, Yunnan Province]
Caridina kunnathurensis Richard & Chandran, 1994: 250; Figs 4, 5A. [Kunnathur pond, Madras, India]
Caridina laevis Heller, 1862b
= C.[aridina] laevis Heller, 1862b: 411. [Java]
Caridina lamiana Holthuis, 1965a: 39; Fig. 14. [Ambodivoangy near Maroantsetra, Antongil Bay, NE
Madagascar; Forest between 20 and 30 km NW of Ambahoab, Simianona River, N of Soanierana,
east coast of Madagascar]
= Caridina brevirostris var. brevipes Bouvier, 1925: 230 (partim). [baie Amposindova, Madagascar]
Caridina lanceifrons Yu, 1936: 89; Figs 4-7. [near the light house at Hai-kiu-sche in the salt water]
Caridina lanceolata Woltereck, 1937: 224; Fig. 1.7a-c; Table 1. [Lake Mantano, Mengonuwai, Sulawesi;
neotype designated by Cai, Wowor & Choy, 2009]
Caridina lanzana Holthuis, 1980c: 2; Figs 1-3. [small well named Bog Der, 0835'54''N 4846'30''E, Nogal
Valley, northern Somalia]
Caridina laoagensis Blanco, 1939a: 390; Plate 2. [?Laoag River, Laoag, Ilocos Norte Province, Luzon]
Caridina leclerci Cai & Ng, 2009: 1108; Fig. 9. [Pange, cave at Tallasa, Maros, Sulawesi Selatan]
Caridina leucosticta Stimpson, 1860a: 28. [13856.43'E 3441.83'E, Inouzawa River, Shimoda City, Izu Pen-
insula; neotype designation by Cai et al., 2006]
Caridina leytensis Blanco, 1939a: 391; Plate 3, Figs 1-7. [Helosig, Leyte]
Caridina liangi Jiang, Guo & Zhang, 2002: 220; Fig. 1. [small stream at an elevation of 450 m near Yong-
ping Village, Yuanling County]
Caridina liaoi Cai, Choy & Ng, 2009: 72; Figs 5-6. [Bilar River, Bilar]
Caridina lima Liang, Guo & Gao, 1993: 43; Fig. 2. [Buermen, Yongshun County, Hunan, 29N 1098'E]
Caridina linduensis Roux, 1904: 541; Plate 9, fgs 1-4. [Lac Lindu, rgion centrale de Clbes (part. Occid.),
altitude 980 m]
Caridina lineorostris Richard & Clark, 2009: 51; Figs 26-27. [Gabon, Keri, forest stream, 20 km North of
Lambarene]
Caridina lingkonae Woltereck, 1937
= Caridina Lingkonae Woltereck, 1937: 218; Fig. 1.1; Table 1. [Lake Towuti, west shore, at entrance to
outlet bay, Cape Larona, 0248.43'S 12124.75'E; neotype designated by von Rintelen & Cai, 2009]
Caridina lipalmaria Richard & Clark, 2010b: 639; Figs 1-2. [Bestiboka River Basin, Beharena River, Antan-
iditra, 4802'55''E 18 25'43''S, 13730 asl]
Caridina liui Liang & Yan, 1986a: 202; Fig. 4. [Tongzi County, Guizhou Province]
Caridina lobocensis Cai, Choy & Ng, 2009: 68; Figs 2-4. [tributuary of Loboc River, Loboc]
Caridina loehae Woltereck, 1937
= Caridina Lohae Woltereck, 1937: 222; Fig. 1.5a-d; Table 1. [Lake Towuti, about 3 km south of Timam-
pu, estuary of Sungei Batuopa, Sulawesi, Indonesia; neotype designated by Cai, Wowor & Choy, 2009]
Caridina longa Liang & Yan, 1985a: 200; Fig. 3. [spring water, Huanglongsi, Jianshui, Yunnan Province]
Caridina longiacuta Guo & Wang, 2005: 14; Fig. 1. [near Yuanliping village, Yizhang County, Hunan Prov-
ince (ca. 2525'N 11257'E)]
Caridina longicarpus Roux, 1926a
= Caridina weberi var. longicarpus Roux, 1926a: 212; Figs 37-39. [Nouvelle-Caldonie, au-dessus
dOubatche, pentes du Mt. Ignambi, env. 600 m daltitude]
Caridina longidigita Cai & Wowor, 2007: 317; Figs 4-5. [west coast of Lake Poso at Taipa area, Kab. Poso,
Sulawesi Tengah]
Caridina longifrons Cai & Ng, 2007: 1599; Fig. 6. [Kabupaten Maros, stream above Bantimurung waterfall,
Sulawesi, Indonesia]
Caridina longirostris H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]
= C.[aridina] longirostris H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 363. [la rivire
de la Macta, prs dOran [probably erroneous]]
= Caridina nilotica var. meridionalis Roux, 1926a: 207. [Pemboa; Haute Tiouaka; Kon; Coula-Borar;
La Foa; Coind, Ruisseua Katiramona, entre Nouma et Pata (all Nouvelle-Caldonie)]
= Caridina acuticaudata ng, 1975: 70; Fig. 4. [Boi River, Hoa Binh Province]
Caridina lovoensis Roth-Woltereck, 1955: 197; Figs 1-2. [Lovo B., 20 km von Thysville (Bas-Congo),
Kalkhhlen]
Caridina lufengensis Cai & Duan, 1998: 330; Figs 1-3. [pond near Sizhi Town, Lufeng County, Yunnan,
China, 25.07N, 102.10E, elevation 1800 m]
Caridina lumilympha Richard & Clark, 2010a: 328; Figs 12-13. [Kenya, Lumi River]
Caridina macrodentata Cai & Shokita, 2006b: 2156; Figs 13-14. [12351.84'E 2423.65'N, fast fowing water,
about 200 m from sea, Omia River, Iriomote Island, Ryukyu Islands]
Caridina macrophora Kemp, 1918b
= Caridina nilotica var. macrophora Kemp, 1918b: 277; Fig. 9. [Tale Sap, Peninsular Siam]
= Caridina subnilotica ng, 1975: 69; Fig. 3. [ponds in Hanoi]
Caridina maculata L. Wang, Liang & F. Li, 2008: 49; Figs 1-2. [Yingde County, Guangdong Province,
N2354.5' E11313.6']
Caridina mahalona Cai, Wowor & Choy, 2009: 29; Figs 8-9. [Lake Mahalona, at southwest coast]
Caridina malayensis Cai, Ng & Choy, 2007: 305; Fig. 17. [stream at Nee Soon Swamp, Singapore]
Caridina malawensis Richard & Clark, 2009: 69; Figs 38-39. [Malawi, rocky shore of Lake Malawi]
Caridina masapi Woltereck, 1937
= Caridina Masapi Woltereck, 1937: 223; Fig. 1.6a-h; Table 1. [southern shore of Lake Masapi, South
Sulawesi, 250.84'S 12121.09'E; neotype designated by Cai, Wowor & Choy, 2009]
Caridina mathiassi Silas & Jayachandran, 2010: 1; Figs 1-5, 6J, 7. [Mahendragiri Estate, part of southern
Western Ghats, Kanyakumari District, Tamil Nadu, India, 8 27'32.2''N 7723'32.5''E]
Caridina mauritii Bouvier, 1912a
= Caridina Mauritii Bouvier, 1912a: 298. [Maurice]
Caridina mccullochi Roux, 1926b: 249. [Pallal, Horton River, near Bingara]
Caridina medifolia Cai & Yuan, 1996: 388; Figs 16-18. [Xiaojin village near Daqun town]
Caridina mengae Liang, 1993: 22; Fig. 1. [Xianshui Cave, Panshi village, Songtao County (28.19N
109.20E), Guizhou Province]
Caridina mengaeoides Guo & Suzuki, 1996: 97; Figs 1-2. [Yuanling County, Hunan Province, China, ca.
2818'N 11020'E, altitude ca. 750 m]
Caridina menghaiensis Cai & Dai, 1999: 217; Figs 5-6. [reservoir in Menghai County]
Caridina meridionalis L. Wang, Liang & F. Li, 2008: 56; Figs 7-8. [Heyuan County, Guangdong Province]
Caridina mertoni Roux, 1911: 84. [Elat, le de Grand-Kei; entre Elat et Ohinangan, le de Grand-Kei; En-
raalan, le de Grand-Kei; Warka, le de Grand-Kei]
Caridina mesofuminis Richard & Clark, 2009: 66; Figs 36-37. [Cameroon, Mess stream]
Caridina mindanao Cai & Shokita, 2006a: 259; Figs 7-9. [Lake Mainit, Mindanao, Philippines]
Caridina minidentata Cai & Anker, 2004: 250; Figs 10-11. [Can Gortio Cave, 1159'00''N 12453'00''E, alti-
tude ca 200 m, Matalud, West Samar]
Caridina minnanica Liang, 2002a: 121; Fig. 3. [China, Fujian Province, Yunxiao County]
Caridina modiglianii Nobili, 1900a
= Caridina Modiglianii Nobili, 1900a: 477. [Kifa-juc]
Caridina moeri Roth-Woltereck, 1984: 102; Figs 1 (1a-b, 2, 3 (1a-e). [Mwerusee]
Caridina mongziensis Liang, Yan & Z.-Z. Wang, 1987: 133; Figs 1-9. [Mongzi County, 2320'N 10325'E,
Yunnan Province]
Caridina multidentata Stimpson, 1860a: 29. [stream in Tenno-ura, Chichi-jima Island, Ogasawara Islands;
neotype designation by Cai et al., 2006]
= Caridina japonica De Man, 1892b: 261; Plate 9, Figs 7-7a. [Kagar Hayagana, Japan]
= Caridina Voeltzkowi Lenz, 1910a: 569. [Alaotra-See (Madagaskar)]
= Caridina japonica sikokuensis Kubo, 1938a: 91; Fig. 20. [Rygad, Kti Prefecture]
Caridina nanaoensis Cai & N. K. Ng, 1999: 1617; Figs 6a, f, 8. [Xiaoliao stream near Xishan Town, Nanao
County, Guangdong Province, China]
Caridina natalensis Bouvier, 1925
= Caridina africana forme natalensis Bouvier, 1925: 214. [South Africa, Umholti River, Verulum, Natal
(according to Richard & Clark, 2009)]
Caridina natarajani Tiwari & R.S. Pillai, 1968: 163; Fig. 1. [Trivandrum, Kerala]
Caridina neglecta Cai & Ng, 2007: 1595; Figs 4-5. [Sungai Batang, 13 km on road from Palopo to Wotu,
Sulawesi, Indonesia]
Caridina nguyeni S.-Q. Li & Liang, 2002: 709; Figs 1-2. [PacBo Village, District Ha Quang, Cao Bang
Province,Vietnam]
Caridina nilotica (Roux, 1833)
= Pelias Niloticus Roux, 1833: 73; Plate 7, fg. 1. [les eaux du Nil]
= Caridina nilotica var. typica Bouvier, 1925: 146; Fig. 308. [Caire]
Caridina norvestica Holthuis, 1965a: 35; Fig. 12. [River north of Majunga; near Majunga, small brook at
km 530 of the highway to Tananarive; Mahajambe lake, Madagascar]
Caridina novaecaledoniae Roux, 1926a
= Caridina novae-caledoniae Roux, 1926a: 214; Figs 40-46. [Tchalabel; Valle du Diahot, chemin du Col
Poraris, 150 m; versant Est du Col Poraris, 100 m; Pemboa; Haute Tiouaka (all Nouvelle-Caldonie)]
Caridina nudirostris Choy, 1984: 288; Fig. 1. [above Vuwa Falls, Wainisavulevu Creek, Nadrau Plateau,
Central Viti Levu, Fii, 1750'30''S 17801'30''E, altitude 800 m; below Vuwa Falls, Wainisavulevu
Creek, 1748'10''S 17805'30''E, altitude 160 m; above Monasavu Falls, Nanuku Creek, Nadrau Pla-
teau, Central Viti Levu, Fii; 1744'30''S 17802'30''E, altitude 700 m]
Caridina okiamnis Richard & Clark, 2009: 54; Figs 28-29. [Cameroon, Okia stream]
Caridina okinawa Cai & Shokita, 2006b: 2154; Fig. 12. [Ingue Cave at Yomitan Village, Okinawa Island,
Ryukyu Islands, Japan]
Caridina oligospina Liang, Guo & Tang, 1999: 72; Fig. 3. [China, Hunan Province, Fengfang County, Muli
village]
Caridina opaensis Roux, 1904: 547; Plate 9, fgs 8-10. [Lac Opa. Bras S.-E. de lle, region centrale, alt. 30 m]
Caridina palawanensis Cai & Shokita, 2006a: 256; Figs 5, 6A-B. [Panitian River, upper stream, ca. 30 km
from river mouth, Palawan Island, Philippines]
Caridina panikkari Jalihal, Shenoy & Sankolli, 1984: 9; Figs 4-5. [Hosayellapur tank (Hirekeri), Dharwar]
Caridina papuana Nobili, 1905d
= Caridina Weberi var. papuana Nobili, 1905d: 481; Plate 12, fg. 1-1b. [Nova Guinea Tedesca]
Caridina paracornuta Cai & Yuan, 1996: 391; Figs 19-21. [Chuangfengao stream, Hongyan village]
Caridina pareparensis De Man, 1892a: 379; Plate 22, Figs 25-25b. [river near Pare Pare, Sulawesi, Indonesia;
lectotype designated by Cai & Ng, 2009]
Caridina parvidentata Roux, 1904
= Caridina pareparensis var. parvidentata Roux, 1904: 545; Plate 9, fgs 5-7. [Malawa Quelle, Celebes (=
Sulawesi); lectotype designation by Cai & Ng, 2009]
Caridina parvirostris De Man, 1892a: 375; Plate 22, fg. 24. [Flores, Fluss bei Bombang]
Caridina parvocula Gurney, 1984: 569; Figs 1-3. [Dark zone, la Grote dAntsatrobonko, Ankarana Massif,
40 km north of Ambilob and 60 km south of Diego Suarez, northern province of Diego Suares]
Caridina parvula von Rintelen & Cai, 2009: 392; Figs 27-29. [Lake Towuti, southwest shore, Cape Sioloya,
0250.7'S 12126.32'E]
Caridina paucidentata L.-Q. Wang & Liang, 2005: 748; Figs 1-15. [Mongzi County, Yunnan Province]
Caridina pedicultrata Guo & Choy, 1994: 123; Figs 1-2. [stream near Huaihua City, at Xuefeng Mountain,
Hunan, c. 2740'N 11011'E]
Caridina peninsularis Kemp, 1918b
= Caridina brachydactyla peninsularis Kemp, 1918b: 279; Fig. 10. [Botanical Garden, Penang, Malaysia;
lectotype designation by Cai, Ng & Choy, 2007]
Caridina petiti Roux, 1929
= Caridina Petiti Roux, 1929: 108; Figs 13-18. [Ambila, lagunes orientales]
Caridina pingi Y, 1938: 294; Figs 9-10. [Amoy, Fukien]
Caridina pingioides Y, 1938: 298; Figs 11-12. [locality being unkown to the writer]
Caridina plicata Liang, 2004: 282; Fig. 138. [Xita, Jiangxi]
Caridina prashadi Tiwari & R.S. Pillai, 1971: 87; Figs 3-4. [Aberdeen (Port Blair), South Andaman, from a
freshwater stream]
= Caridina sakishimensis Fujino & Shokita, 1975: 99; Fig. 5. [Yonaguni I.: Arakawabana River; Spring
of Tendabanata; Tabaru River; Ishigaki I.: Miyara River; Yonehara River; Sakutara River; Fukido
River; Miyako I.: Izaga Cave; Bora Rover; Kume I.: Shirase River]
Caridina pristis Roux, 1931b: 63; Figs 1-3. [alentours de Perdenyia; prs de Kandy, Ceylan]
= Caridina pristis cruszi de Silva, 1982: 131; Fig. 3. [stream in Sinharaja forest]
Caridina profundicola von Rintelen & Cai, 2009: 396; Figs 30-32. [Lake Towuti, Loeha Island, southwest
shore, 0245.58'S 12131.14'E, on boulders in deeper water]
Caridina propinqua De Man, 1908b: 227; Plate 19, Figs 6-6f. [Dhappa, near Calcuta]
= Caridina blancoi Chace, 1997: 6; Fig. 2. [Philippines, near mouth of Tyabas River, Luzon, 1354'N
12136'E]
Caridina pseudodenticulata Hung, Chan & Yu, 1993: 498; Figs 9D, 13. [Yun Lin County, Ku Keng]
Caridina pseudonilotica Richard & Clark, 2005: 722; Figs 9-10. [Uganda, Bufundi, Lake Bunyonyi]
Caridina pseudoserrata ng & , 2007: 1; Figs 1-2. [Bang River, Cao Bang Province]
Caridina qingyuanensis Guo & He, 2007: 47; Figs 1-19. [Baishikeng Village, Qingyuan City, Guangdong-
Province (ca. 2335'N 11310'E)]
Caridina rajadhari Bouvier, 1918
= Caridina Rajadhari Bouvier, 1918: 386; Figs 1-3. [Rajadhar, dans ltat de Kawarda, massif monta-
gneux situ dans les provinces centrales, entre Jubbulpoor et Nagpoor; Majghaon, non loin de Ra-
jadhar; Mukhi, mme rgion]
Caridina rangoona Cai & Ng, 2000: 939; Fig. 6. [1709.97'N 9699.20'E, Win Paw Hta River, near border
between Pegu (Bago) and Yangon, Yangon State, Myanmar]
Caridina rapaensis Edmondson, 1935b: 12; Fig. 5a-h. [freshwater stream, Rapa]
Caridina richtersi Thallwitz, 1892
= Caridina serrata Richters, 1880: 163; Plate 17, Figs 24-27; nec Stimpson, 1860a. [Botan. Garten Pam-
plemousses; Black river]
= Caridina richtersii Thallwitz, 1892: 27. [nomen novum for Caridina serrata Richters, 1880]
= Caridina apiocheles Bouvier, 1904a: 134. [peut-tre des Seychelles]
= Caridina apiocheles mutation Edwardsi Bouvier, 1904a: 134. [peut-tre des Seychelles]
= Caridina Richtersi forma typica Bouvier, 1925: 203. [Type locality not indicated]
Caridina roubaudi Bouvier, 1925
= Caridina africana forme Roubaudi Bouvier, 1925: 217; Fig. 477. [Congo, Brazzaville; lectotype desig-
nated by Richard & Clark, 2009]
Caridina rouxi De Man, 1915a: 387; Plate 27, fgs 1-11. [in einem kleinen, schnell fiessenden Flusse im
Bougainville-Gebirge, 500 m]
Caridina rubella Fujino & Shokita, 1975: 102; Fig. 6. [Miyako I.: Izaga Cave, Morikaga Cave; Nikadori,
Hirara City, in well]
Caridina rubropunctata ng & , 2007: 3; Figs 3-4. [streams in Van Lang, Dong Hy District, Thai Nguy-
en Province]
Caridina samar Cai & Anker, 2004: 240; Fig. 4-5. [Sulpan Cave, 1202'00''N 12455'30''E, altitude 100 m,
Matalud, West Samar, Philippines]
Caridina sarasinorum Schenkel, 1902: 492; Plate 8, Figs 2a-e. [Lake Poso, central Sulawesi, Indonesia; lec-
totype designation by Cai & Wowor, 2007]
Caridina schenkeli von Rintelen & Cai, 2009: 442; Figs 60-62. [small stream, west of Lake Poso, 022.613'S
12037.311'E, on macrophytes]
Caridina semiblepsia Guo, Choy & Gui, 1996: 66; Figs 1-4. [Dongpaoshan Cave (c. 2844'N 10939'E), Hu-
nan Province, China]
Caridina serrata Stimpson, 1860a: 29. [hill above Bekhers, Hong Kong island, Hong Kong; neotype desig-
nation by Cai & N. K. Ng, 1999]
Caridina serratirostris De Man, 1892a: 382; Plate 23, Figs 28-28e. [Bangkalanvir River, Saleyer; lectotype
designation by Richard & Clark, 2010a]
Caridina shenoyi Jalihal & Sankolli in Jalihal, Shenoy & Sankolli, 1984: 21; Figs 9-11. [Malaprabha river
near old bridge, Khanapur (Belgaum District), India]
Caridina shilinica Liang & Cai, 2000: 177; Fig. 1. [Shilin (Stone forest) Tour Resort in Lunan County Yun-
nan Province, southern China]
Caridina similis Bouvier, 1904a: 135. [Mah, Seychelles; lectotype designation by Marquet & Keith, 2008]
= Caridina brevirostris forma typica Bouvier, 1912b: 916. [Seychelles]
= Caridina brevirostris forma Gardineri Bouvier, 1912b: 916. [Seychelles]
Caridina simoni Bouvier, 1904a
= Caridina Simoni Bouvier, 1904a: 131. [Kandee, lle de Ceylan]
= Caridina costai de Silva, 1982: 126; Fig. 2. [three streams in Sinharaja forest, Sri Lanka]
Caridina sodenensis Richard & Clark, 2009: 42; Figs 20-21. [Cameroon, Lake Soden, West Cameroons]
Caridina solearipes Guo & De Grave, 1997: 127; Figs 2-3. [small stream near Dabaozi, Jingzhou County,
Hunan province, circa 2641'N 0925'E, altitude between 400-900 m]
Caridina songtaoensis Liang, 2004: 270; Fig. 32. [Songtao, Guizhou]
Caridina spathulirostris Richters, 1880: 163; Plate 17, fg. 28. [Botanischer Garten Pamplemousses]
Caridina spelunca Choy, 1996: 103; Fig. 1. [Old Napier Downs Cave, Western Australia, Australia, 1714'S
12439'E]
Caridina sphyrapoda Liang & Zhou, 1993: 236; Figs 4(1)-4(2). [Longdong River in Laibing County,
Guangxi, southern China]
Caridina spinalifrons Guo & De Grave, 1997: 124; Fig. 1. [stream near Sangzhi city, Hunan Province, circa
2910'N 11016'E, 650 m altitude]
Caridina spinata Woltereck, 1937: 221; Fig. 1.3; Table 1. [Lake Towuti, estuary of Sungei Batuopa, about 2
km south of Timampu, Sulawesi, Indonesia; neotype designated by Cai, Wowor & Choy, 2009]
Caridina spinipoda Liang, Hong & Yang, 1990: 161; Figs 1-9. [Mianzhu County, 3120'N 10411'E, Sichuan
Province]
Caridina spinosipes Liang, Guo & Tang, 1999: 71; Fig. 2. [China, Hunan Province, Fengfang County, Muli
village]
Caridina spinula Choy & Marshall, 1997: 31; Figs 1c, 2g-h, 4f-q. [east fowing frst order tributary of
Leo Creek, near crossing of Leo Creek Mine Road, 1344.6'S 14321.5'E, Nesbit River catchment,
McIlwraith Range, Cape York]
Caridina spongicola Zitzler & Cai, 2006: 271; Figs 1-3. [Indonesia, Sulawesi Selatan, Lake Towuti, west
shore, outlet bay, west of Cape Tokaluku, 0247.261'S 12123.17'E]
Caridina steineri Cai, 2005: 314; Figs 2-3. [Lakata Zafera, western Madagascar, 1945.075'S 4511.436'E,
stream pool at far end of cave]
Caridina striata von Rintelen & Cai, 2009: 410; Figs 39-41. [Indonesia, Sulawesi Selatan, Lake Towuti,
north shore, 0238.56'S 12127.82'E, on rocks]
Caridina subventralis Richard & Clark, 2005: 725; Figs 11-12. [Uganda, Bufundi, Lake Bunyonyi]
Caridina sulawesi Cai & Ng, 2009: 1103; Figs 6-8. [Sungai Beru, Kappang, Maros, Sulawesi Selatan]
Caridina sumatianica Cai & Yuan, 1996: 395; Figs 22-23. [Sumatian stream, near Hongyan village]
Caridina sumatrensis De Man, 1892a
= Caridina Weberi var. sumatrensis De Man, 1892a: 375; Plate 22, fg. 23g. [fssen des unteren Bat-
taklandes bei Deli, an der Ostkste von Sumatra]
Caridina sundanella Holthuis, 1978a: 32; Figs 11-12. [Waikamburu Brook, 4 km N. of Waimangura, West
Sumba, about 250 m above sea level]
Caridina susurufabra Richard & Clark, 2009: 60; Figs 32-33. [South Africa, Eoshowe, Zululand]
Caridina temasek Choy & Ng, 1991: 266; Figs 2-5. [freshwater stream near Sime road, MacRitchie catch-
ment area, Singapore, ca. 0120'14''N 10348'47''E]
Caridina tenuirostris Woltereck, 1937: 224; Fig. 1.8; Table 1. [Lake Towuti, southwest shore, west of Cape
Tetetu, 0254.13'S 12123.78'E; neotype designation by von Rintelen & Cai, 2009]
= Caridina Towutensis Woltereck, 1937: 220; Fig. 1.2; Table 1. [Towuti-Sd]
Caridina thambipillai Johnson, 1961: 138; Figs 25-35. [Sungei Putat near the pumping station, Malacca]
Caridina thermophila Riek, 1953: 120; Fig. 10. [Mutaburra, western Queensland]
Caridina thomasi von Rintelen, Karge & Klotz, 2008: 2247; Figs 2-3. [Indonesia, Central Sulawesi, Banggai
islands, Peleng, west peninsula, east of Alani, river with lake-like extension, 0128.315'S 12252.473'E]
Caridina timorensis De Man, 1893: 300; Plate 8, fg. 6. [freshwater lake of Nefo on the island of Timor]
Caridina togoensis Hilgendorf, 1893a: 156. [Togo, Bismarksburg; lectotype designation by Richard &
Clark, 2009]
= Caridina togonsis var. stuhlmanni Hilgendorf, 1898: 35. [Undussuma und Bach bei Undussuma]
= Caridina togoensis var. Decorsei Bouvier, 1904a: 131. [Congo franais, rivire prs de Mpoko; rgion
Chari-Tchad, Krbdje (Fort-Sibut) et Bingutou, rivire Gribingui; Bangoran]
= Caridina togoensis var. breviatus Lenz, 1910b: 131. [Urwaldbache N.W. von Beni]
= Caridina africana forme gyptiaca Bouvier, 1925: 214. [Caire]
= Caridina togoensis var. Kasaiensis De Man, 1925: 5; Figs 2
n1
-2
n3
. [Kamaiembe; Kondu]
= Caridina togoensis var. Kwamouthensis De Man, 1925: 9; Fig. 2
o1
-2
012
. [Kwamouth au confuent du
feuve Congo et de la rivire Kasa]
= Caridina togoensis var. Schoutedeni De Man, 1925: 20; Figs 2
q1
-2
q21
. [les rivires du bassin du Chiloan-
go; Buto-Polo dans la rivire Mapanga]
Caridina tonkinensis Bouvier, 1919: 331. [Tonkin]
Caridina trifasciata Yam & Cai, 2003: 277; Figs 1, 3-5. [Tsak Yue Wu, New Territories, Hong Kong]
Caridina troglodytes Holthuis, 1978b: 214; Figs 3-4. [Danmin Cave near Konogusgus, New Ireland, in
subterranean stagnant pool, 0.3-0.6 m]
Caridina troglophila Holthuis, 1965a: 37; Fig. 13. [Ambovonomby cave, Namoroka, NW Madagascar]
Caridina tumida L. Wang, Liang & F. Li, 2008: 54; Figs 5-6. [Yingde County, Guangdong Province,
N2354.4' E11315.1']
Caridina typus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]
= C.[aridina] typus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 363; Plate 25bis, Figs
4-5. [Type locality unknown, probably lle Maurice according to Bouvier, 1925]
= Caridina exilirostris Stimpson, 1860a: 29. [Okuma River, Okinawa Island, Rykyu islands; neotype
designation by Cai, Ng, Shokita & Satake, 2006]
= Caridina siamensis Giebel, 1863: 329. [Siam]
? = Caridina Spencebatei De Man, 1892a: 371. [?Cap Verdischen Inseln]
= Caridina typus forme typica Bouvier, 1925: 250. [lle Maurice]
= Caridina typus forme caledonica Bouvier, 1925: 253; Figs 296-297. [Nouvelle-Caldonie]
Caridina uminensis ng & , 2007: 6; Figs 5-6. [U Minh Thuong wetland, Ca Mau Province]
Caridina umtatensis Richard & Clark, 2009: 63; Figs 34-35. [South Africa, Kraal Dam, Umtata]
Caridina unca Gurney, 1984: 579; Figs 8-11. [Dark zone, Grote dAntsatrobonko, Andarana Massif, north-
ern province of Diego Suarez]
Caridina valencia Cai, Choy & Ng, 2009: 78; Fig. 9. [spring Anislaq, outside Valencia]
Caridina venusta L. Wang, Liang & F. Li, 2008: 52; Figs 3-4. [Yingde County, Guagndon Province,
N2354.3' E11315.8']
Caridina vietriensis ng & , 2007: 9; Figs 7-8. [confuence of rivers in Viet Tri City, Phu Tho Province]
Caridina villadolidi Blanco, 1939a: 389; Plate 1. [Laoag River, Laoag, Ilocos Norte Province, Luzon]
= Caridina typus var. longirostris De Man, 1892a: 370; Plate 22, fg. 22f-i; nec C.[aridina] longirostris H.
Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]. [Flores, as einem Flusse bei Reo; auch auf
Celebes und Saleyer]
Caridina vitiensis Borradaile, 1899: 1003; Plate 63, fgs 3-3a. [Suva, Tamavua River, Viti Levu, Fii]
= Caridina vitiensis var. canacorum Roux, 1926a: 199; Figs 32-36. [Nouvelle-Caldonie, Canala]
Caridina weberi De Man, 1892a
= Caridina Weberi De Man, 1892a: 371; Plate 22, fgs 23-23g. [Koting; Fluss bei Mbawa, oberhalb des
Wasserfalls; Fluss bei Bombang (all Flores); Fluss bei Palopo, Luwu; Wasserfall bei Bantimurong
unweit Maros; Fluss bei Pare-pare (all Celebes); Saleyer, Fluss Bangkalan]
= Caridina weberi var. typica Bouvier, 1925: 243; Figs 562-566. [Flores, Koting, environs de Mbawa,
de Bombang; Java; Kifa-juc, Ile Engano]
? = Caridina weberi var. keiensis Roux, 1911: 85. [Elat, le de Grand-Kei; ruisseau entre Elat et Ohi-
nangan; Warka, le de Grand-Kei; ruisseau entre Elat et Ohilim, le de Grand-Kei]
Caridina williamsi Cai & Ng, 2000: 933; Figs 4-5. [1748.11'N 9609.20'E, Balar stream, 12 km Yangon to
Mandalay highway, Htauk Kyant, Yangon State, Myanmar]
Caridina woltereckae Cai, Wowor & Choy, 2009: 19; Fig. 3. [Cape Larona, near Sungai Larona outlet, Lae
Towuti, Sulawesi, Indonesia]
Caridina wumingensis Cai & N.K. Ng, 1999: 1632; Figs 18-19. [cave near Wuming, Guangxi Province,
China]
Caridina wyckii (Hickson, 1888)
= Atya Wyckii Hickson, 1888: 358; Plates 13-14. [Lake Tondano, Minahassa, North Celebes]
= Caridina nilotica var. minahassae De Man, 1902: 895. [Minahassa, Celebes]
Caridina xiangnanensis X.-Y. Liu, Guo & Yu, 2006: 44; Fig. 1. [near Lingxiu Village, Rucheng County, Hu-
nan Province (ca. 2533'N 11340'E)]
Caridina xiphias Bouvier, 1925
= Caridina nilotica var. xiphias Bouvier, 1925: 149; Figs 310-312. [district dAmbatondrazaka, Mada-
gascar]
= Caridina nilotica var. stylirostris Bouvier, 1925: 148; Fig. 309. [Madagascar]
= Caridina gladiifera Roux, 1929: 306; Figs 1-6. [Prinet, fort, Madagascar]
Caridina yilong Cai & Liang, 1999: 76; Fig. 3. [Yilong Lake, Shiping County, Yunnan]
Caridina yulinica Cai & N.K. Ng, 1999: 1620; Figs 9-11. [Niuwo cave near Kuiyang Town, Yulin County,
Guangxi Province, China]
Caridina yunnanensis Y, 1938: 304; Figs 14-16. [Mountain stream at Hsn-tien, Yunnan Province]
= Caridina impensa Cai & Ng, 2001a: 223; Figs 8-9. [Xilong Tang (Pond), Chengjiang County, Yunnan]
Caridina zebra Short, 1993: 62; Figs 1-3. [OLeary Creek, 1750.7'S 14537.7'E, altitude ca. 750 m]
Caridina zeylanica Arudpragasam & Costa, 1962: 17; Fig. 4. [Nawala, suburb of Colombo]
= Caridina nilotica var. veliensis R.S. Pillai, 1964: 44. [Veli Lake, Trivandrum]
Caridina zhejiangensis Liang & Zheng, 1985
= Caridina pingi zhejiangensis Liang & Zheng, 1985: 321; Fig. 1(1c-7c). [Baisha Village, Jiande County,
Zhejiang Province, China]
Caridina zhongshanica Liang, 2004: 182; Fig. 88. [Zhongshan, Guangdong]
Caridinides Calman, 1926
= Caridinides Calman, 1926 (type species Caridinides wilkinsi Calman, 1926, by monotypy, gender
masculine)
Caridinides wilkinsi Calman, 1926: 242; Figs 1-2. [Olive River, Temple Bay, east coast of Cape York Penin-
sula]
Caridinopsis Bouvier, 1912c
= Caridinopsis Bouvier, 1912c (type species Caridinopsis Chevalieri Bouvier, 1912c, by monotypy,
1957)
Caridinopsis chevalieri Bouvier, 1912c
= Caridinopsis Chevalieri Bouvier, 1912c: 300; Figs 1-4. [rgion du Haut Niger, Sampouyara, bassin
de la Makowa]
= Caridinopsis brevinaris Holthuis, 1956a: 56. [Garrigues Cave near Sougouta, French Guinea]
Dugastella Bouvier, 1912d
= Dugastella Bouvier, 1912d (type species Dugastella marocana Bouvier, 1912d, by monotypy, gender
feminine; name placed on the Ofcial List of Generic Names in Zoology in Opinion 470 in 1957)
Dugastella marocana Bouvier, 1912d: 993. [la source de Setat]
Dugastella valentina (Ferrer Galdiano, 1924)
= Atyaephira valentina Ferrer Galdiano, 1924: 210; Figs 1, 3. [Laguna de Almenara (Castelln) y Al-
bufera de Valencia]
= Dugastella marocana var. hispanica Balss, 1925b: 206; Figs 1-4. [Silla, bei Valencia, Acequia com-
munera, 1.5 km v. Albufera]
Edoneus Holthuis, 1978b
= Edoneus Holthuis, 1978b (type species Edoneus atheatus Holthuis, 1978b, by original designation
and monotypy, gender masculine)
Edoneus atheatus Holthuis, 1978b: 220; Figs 5-6. [the Philippines, Luzon, Quirino province, Aglipay Mu-
nicipality, Barrio Palasian, cave near Sitio Disiluad; see Balete & Holthuis, 1992]
Edoneus erwini Cai & Husana, 2009: 54; Figs 2-3. [Bantakay Cave, station 2, Luzon, the Philippines]
Edoneus marulas Cai & Husana, 2009: 60; Figs 6-7. [at middle of Marulas Cave, Luzon, the Philippines]
Edoneus sketi Cai & Husana, 2009: 57; Figs 4-5. [small pool in Bantakay cave, Luzon, the Philippines]
Gallocaris Sket & Zakek, 2009
= Gallocaris Sket & Zakek, 2009 (type species Troglocaris Schmidti inermis Fage, 1937, by original
Gallocaris inermis (Fage, 1937)
= Troglocaris Schmidti inermis Fage, 1937: 215; Figs 1-6. [grote de Cambous, prs Saint-Hippolyte-du-
Fort, dpartement du Gard]
Halocaridina Holthuis, 1963b
= Halocaridina Holthuis, 1963b (type species Halocaridina rubra Holthuis, 1963b, by monotypy, gen-
der feminine)
Halocaridina palahemo Kensley & Williams, 1986: 429; Figs 9-11. [Hawaii Island, Ka Lae, Lua o Palahemo,
lava tube pool, 1855'N 15542'W]
Halocaridina rubra Holthuis, 1963b: 262; Fig. 1. [pool in the base of Lohena Rock, between the deserted
villages of Wai-o-ahu-kini and Kaili-kii, W. of Ka Lae or South Point, Kau district, island of Hawaii]
Halocaridinides Fujino & Shokita, 1975
= Halocaridinides Fujino & Shokita, 1975 (type species Halocaridina (Halocaridinides) trigonophthalma
Fujino & Shokita, 1975, by monotypy, gender masculine)
= Palauatya C.W.J. Hart, 1980 (type species Palauatya dasyomma C.W.J. Hart, 1980, by original desig-
Halocaridinides fowleri Gordon in Gordon & Monod, 1968
= Parisia (?) fowleri Gordon in Gordon & Monod, 1968: 514; Figs 30-31. [subterranean lake, Kufle,
Zanzibar]
Halocaridinides trigonophthalma (Fujino & Shokita, 1975)
= Halocaridina (Halocaridinides) trigonophthalma Fujino & Shokita, 1975: 106; Figs 7-8. [Okinawa Is-
land: Kaneshi, Nakiin, in well, 10 m; Aja, Naha City, in well]
= Palauatya dasyomma C.W.J. Hart, 1980: 481; Figs 1-31. [anchialine lake on Anguar Island, Palau,
Caroline Islands, 654'49''N 13408'12''E]
Jolivetya Cals, 1986
= Jolivetya Cals, 1986 (type species Jolivetya foresti Cals, 1986, by monotypy, gender feminine)
Jolivetya foresti Cals, 1986: 387; Fig. 1. [Nouvelle-Bretagne, la grote de Pogo Ngim, lieu daccs au rseau
souterrain dArais, dans un karst dolines, quelques dizaines de kilomtres du litoral de la Nou-
velle-Bretagne]
Jonga C.W.J. Hart, 1961
= Jonga C.W.J. Hart, 1961 (type species Ortmannia Serrei Bouvier, 1909a, by original designation and
Jonga serrei (Bouvier, 1909a)
= Ortmannia Serrei Bouvier, 1909a: 332. [Cuba]
Lancaris Cai & Bahir, 2005
= Lancaris Cai & Bahir, 2005 (type species Caridina singhalensis Ortmann, 1894, by original designa-
Lancaris kumariae (de Silva, 1990)
= Caridina kumariae de Silva, 1990: 9; Fig. 1. [small stream near railway station at Rozella, Sri Lanka,
656'-657'N 8033'-8035'E]
Lancaris singhalensis (Ortmann, 1894)
= Caridina singhalensis Ortmann, 1894: 11; Plate 1, fg. 2. [Ceylon, Newera Elya]
Limnocaridella Bouvier, 1913b
= Limnocaridella Bouvier, 1913b (type species Limnocaridina Alberti Lenz, 1910b, by monotypy, gen-
der feminine; name placed on the Ofcial List of Generic Names in Zoology in Opinion 470 in
1957)
Limnocaridella alberti (Lenz, 1910b)
= Limnocaridina Alberti Lenz, 1910b: 132; Plate 3, Figs 6-9. [Albert-See]
Limnocaridina Calman, 1899a
= Limnocaridina Calman, 1899a (type species Limnocaridina tanganyik Calman, 1899a, by monotypy,
1957)
Limnocaridina iridinae Roth-Woltereck, 1958: 188; Figs 1-5. [Tanganjika-See bei Mpulungu (Nordrho-
desien, Abercorn District), 3 m, aus dem Kiemenraum einer Iridina spekei]
Limnocaridina latipes Calman, 1906a: 196; Plate 12, fgs 23-29. [Mbete, shallow water amongst rocks; near
mouth of Lofu, on rocks, shallow water; Kalambo, tow-neting; Tembwi, on rocks, shallow water
(all Lake Tanganyika)]
Limnocaridina parvula Calman, 1906a: 193; Plate 11, fgs 9-14. [Kasawa, tow-neting; Kalambo, tow-net-
ting; Karema tow-neting, surface (all lake Tangaynika)]
Limnocaridina retiarius Calman, 1906a: 192; Plate 11, fgs 2-8. [Lake Tanganyika, Mbete, taken in shrimp-
net, shore wading]
Limnocaridina similis Calman, 1906a: 195; Plate 12, fgs 15-22. [Lake Tanganyika, Kalambo, tow-neting;
Rusisi River, close to Tanganyika]
Limnocaridina socius Calman, 1906a: 196; Plate 12, fgs 30-37. [Niamkolo Harbour, 3 fms; Utinta, 10 fms;
Kirando, about 8 fms (all Lake Tangyanika)]
Limnocaridina spinipes Calman, 1906a: 197; Plate 13, fgs 38-44. [Niamkolo Harbour, 3 fms; Kirando, 8
fms; Utinta, about 15 fms (all Lake Tanganyika)]
Limnocaridina tanganyikae Calman, 1899a
= Limnocaridina tanganyik Calman, 1899a: 704; Plate 39, fgs 1-2, 4-9a; Plate 40, fgs 10-19. [Lake
Tanganyika, shallow water]
Mancicaris Liang, Guo & Tang, 1999
= Mancicaris Liang, Guo & Tang, 1999 (type species Mancicaris sinensis Liang, Guo & Tang, 1999, by
original designation and monotypy, gender feminine)
Mancicaris sinensis Liang, Guo & Tang, 1999: 70; Fig. 1. [Tianxinpu village, Tianxin district, Lanshan
County, Hunan Province, China]
Marosina Cai & Ng, 2005
= Marosina Cai & Ng, 2005 (type species Marosina brevirostris Cai & Ng, 2005, by original designa-
Marosina brevirostris Cai & Ng, 2005: 131; Fig. 1. [Gua Salukkan, Kallang, Kappang, Maros, Sulawesi
Selatan, Indonesia]
Marosina longirostris Cai & Ng, 2005: 133; Figs 2-4. [Gua Salukkan, Kallang, Kappang, Maros, Sulawesi
Selatan]
Micratya Bouvier, 1913b
= Calmania Bouvier, 1909b (type species Atya Poeyi Gurin-Mneville, 1855 [in Gurin-Mneville,
1855-1856], by monotypy, gender feminine; an invalid junior homonym of Calmania Laurie, 1906
(Crustacea Brachyura) and Calmania Nobili, 1907 (a junior subjective synonym of Brachycarpus
Spence Bate, 1888); name placed on the Ofcial Index of Rejected and Invalid Generic Names in
= Micratya Bouvier, 1913b (type species Atya Poeyi Gurin-Mneville, 1855 [in Gurin-Mneville,
1855-1856], by monotypy, gender feminine; name placed on the Ofcial List of Generic Names in
= Balssiola Strand, 1922 (nomen novum for Calmania Bouvier, 1909b, gender feminine; name placed
Micratya poeyi (Gurin-Mneville, 1855 [in Gurin-Mneville, 1855-1856])
= Atya Poeyi Gurin-Mneville, 1855 [in Gurin-Mneville, 1855-1856]: xviii; Plate 2, Figs 7, 7a-b.
[Cuba]
Neocaridina Kubo, 1938a
= Neocaridina Kubo, 1938a (type species Hippolyte denticulatus De Haan, 1844 [in De Haan, 1833-
1850], by original designation, gender feminine)
Neocaridina anhuiensis (Liang, Zhu & Xiong, 1984)
= Caridina denticulata anhuiensis Liang, Zhu & Xiong, 1984: 251; Figs 1-7. [Taiping County, Anhui
Province]
Neocaridina bamana Liang, 2004: 112; Fig. 55. [Bama, Guangxi]
Neocaridina brevidactyla Liang, Chen & W.-X. Li, 2005: 532; Figs 3-43. [caves of Libo County (2541'N
10783'E), Maolan Nature Sanctuary of Guizhou]
Neocaridina curvifrons (Liang, 1979)
= Caridina curvifrons Liang, 1979: 118; Figs 1-10. [Baisha, Zhejiang Province]
Neocaridina denticulata davidi (Bouvier, 1904a)
= Caridina Davidi Bouvier, 1904a: 133. [Inkiafou, Chensi mridonale]
Neocaridina denticulata denticulata (De Haan, 1844 [in De Haan, 1833-1850])
= Hippolyte denticulatus De Haan, 1844 [in De Haan, 1833-1850]: Plate 45, fg. 8. [Japan]
= Caridina denticulata sinensis Kemp, 1918b: 287; Fig. 11c-d. [creeks and irrigation channels at the
edge of the Tai Hu lake in Kiangsu province]
Neocaridina denticulata moganica Liang, 2004: 82; Fig. 41. [Mogashan, Zhejiang]
Neocaridina euspinosa Cai, 1996: 150; Figs 15-16. [Yanshan mountain near Cuilin City (2520'N 11018'E),
Guangxi Zhuang Autonomous Region, China]
Neocaridina fukiensis (Liang & Yan, 1977)
= Caridina spinosa fukiensis Liang & Yan, 1977: 222; Figs 9-11. [Gutian, Shnaghang County (rivulet);
Xinquan, Liancheng County (pool); Luokou and Ansha, Yongan County (pool) (all Fujian Province)]
Neocaridina gracilipoda Liang, 2004: 88; Fig. 44. [Junlian, Sichuan]
Neocaridina heteropoda heteropoda Liang, 2002b
= Neocaridina heteropoda Liang, 2002b: 167; Fig. 1a-h. [Baisha Village, Jiande County, Zhejiang Prov-
ince, China]
Neocaridina heteropoda koreana Kubo, 1938a
= Neocaridina denticulata koreana Kubo, 1938a: 81; Figs 5c, 7g-m, 7g-m, 12. [Huzan, Korea]
Neocaridina heteropoda luoyangensis Cai, 1996
= Neocaridina denticulata luoyangensis Cai, 1996: 139, Figs 7-8. [from a mountain stream near Luoyan
City (3440'N 11220'E), Henan Province, China]
Neocaridina hofendopoda (Shen, 1948)
= Caridina hofendopoda Shen, 1948: 122; Plate 13, Figs f-l. [Kweilin]
Neocaridina homospina Liang, 2002b
= Neocaridina euspinosa homospina Liang, 2002b: 169; Fig. 2. [China, Hunan Province, Shaodong County]
Neocaridina iriomotensis Naruse, Shokita & Cai, 2006: 26; Figs 1, 2. [upper reaches of the Nishifunatsuki,
Nakama River, Iriomote Island]
Neocaridina ishigakiensis (Fujino & Shokita, 1975)
= Caridina denticulata ishigakiensis Fujino & Shokita, 1975: 95; Figs 3-4. [Yonehara River; Miyara
River; Sakutara River, Arakawa River (all Ishigaki Island)]
Neocaridina ketagalan Shih & Cai, 2007: 687; Figs 5-6, 7D-E. [Sihih, Taipei Co., Taiwan]
Neocaridina keunbaei (H.S. Kim, 1976)
= Caridina denticulata keunbaei H.S. Kim, 1976: 155; Figs 1-3. [upper stream of Cheonji fall, Jeju Is-
land]
Neocaridina linfenensis Cai, 1996
= Neocaridina denticulata linfenensis Cai, 1996: 137; Figs 5-6. [near Linfen City, 365'N 11130'E, Shanxi
Province, China]
Neocaridina longipoda (Cai, 1995a)
= Caridina longipoda Cai, 1995a: 166; Figs 1-15. [Jinbianxi stream, Zhangjiajie National Forest Park,
Hunan Province]
Neocaridina palmata bosensis Cai, 1996: 145; Figs 11-12. [Longlin County (2515'N 10522'E), in Guangxi
Zuang Autonomous Region, China]
Neocaridina palmata luodianica Liang, 2004: 110; Fig. 54. [Luodian, Guizhou]
Neocaridina palmata meridionalis Liang, 2004: 108; Fig. 53. [Luoding, Guangdong]
Neocaridina palmata palmata (Shen, 1948)
= Caridina palmata Shen, 1948: 120; Plate 12. [Sha-ping-pa Chungking]
= Caridina elongata Shen, 1948: 121; Plate 13, fgs a-e. [Kweilin]
= Caridina denticulata vietnamensis ng, 1967: 157; Fig. 4. [Cao Bang province; Lang Son province]
Neocaridina saccam Shih & Cai, 2007: 682; Figs 2-4, 7A-B. [mountain stream at Longci, Tainan Co.,
Taiwan]
Neocaridina spinosa (Liang, 1964)
= Caridina spinosa Liang, 1964: 187; Figs 1-12. [mountain stream, Lantang, Tzechin Hsien, Kwangtung]
Neocaridina xiapuensis Zheng, 2002: 19; Fig. 1. [Xiapu County, 2655'N 120E, Fujian Province, China]
Neocaridina zhangjiajiensis Cai, 1996: 147; Figs 13-14. [Zhangjiajie National Forest Park in Dayong City,
Hunan Province of China]
Neocaridina zhoushanensis Cai, 1996
= Neocaridina denticulata zhoushanensis Cai, 1996: 142; Figs 9-10. [Dinhai County in Zhoushan Island
(3002'N 12208'E), Zhejiang Province, China]
Palaemonias Hay, 1902
= Palmonias Hay, 1902 (type species Palmonias ganteri Hay, 1902, by monotypy, gender masculine;
name placed on the Ofcial List of Generic Names in Zoology in Opinion 470 in 1957)
Palaemonias alabamae Smalley, 1961: 127; Fig. 1. [Shelta Cave, Huntsville, Madison County, Alabama]
Palaemonias ganteri Hay, 1902
= Palmonias ganteri Hay, 1902: 179. [Roaring River passage, Mammoth Cave]
Paracaridina Liang, Guo & Tang, 1999
= Paracaridina Liang, Guo & Tang, 1999 (type species Caridina longispina Guo, He, Xu & Gui, 1992,
by original designation and monotypy, gender feminine)
Paracaridina chenxiensis Guo & De Grave, 2004: 203; Figs 2-3. [small stream at an elevation of 450 m near
Houmachong village, Chenxi County, ca. 2747'N 11016'E]
Paracaridina guizhouensis (Liang & Yan, 1986a)
= Caridina guizhouensis Liang & Yan, 1986a: 199; Fig. 2. [Maopo, Yuping County, Guizhou]
Paracaridina leptocarpa (Liang & Zheng, 1988)
= Caridina leptocarpa Liang & Zheng, 1988: 15; Figs 1-9. [Fuzhou, 26N 119E, Min River, Fujian]
Paracaridina longispina (Guo, He, Xu & Gui, 1992)
= Caridina longispina Guo, He, Xu & Gui, 1992: 717; Figs 1-9. [Yuelu Hills, Changsha, China, 11256'E
2810'N]
Paracaridina ziinica Liang, 2002b: 171; Fig. 3. [Ziin, Guangdong, southern China]
Paratya Miers, 1882
= Paratya Miers, 1882 (type species Ephyra compressa De Haan, 1844 [in De Haan, 1833-1850], by
monotypy, gender feminine; name placed on the Ofcial List of Generic Names in Zoology in Opin-
ion 470 in 1957 and in Direction 85 in 1957)
= Xiphocaridina Bouvier, 1909b (type species Ephyra compressa De Haan, 1844 [in De Haan, 1833-
1850], designated by Holthuis, 1955b, gender feminine; name placed on the Ofcial Index of Re-
jected and Invalid Names in Zoology in Opinion 470 in 1957)
= Xiphatyoida Roux, 1915 (established without included nominal species; type species Paratya
(Xiphatyoida) typa Roux, 1926a, designated by Roux, 1926a, gender feminine)
Paratya annamensis Balss, 1924: 45. [Annam, Phuc Son]
Paratya australiensis Kemp, 1917b: 303; Fig. 5. [Seven Hills, near Sydney; neotype designation by W.D.
Williams & Smith, 1979]
= Paratya australiensis arrostra Riek, 1953: 114, Fig. 2. [Terrors Creek, Dayboro, Queensland]
= Paratya atacta Riek, 1953: 114; Fig. 3. [Upper Nerang River, south Queensland]
= Paratya atacta adynata Riek, 1953: 115; Fig. 4. [small creek in upper reaches of Middle Harbour,
Sydney, N.S.W.]
= Paratya tasmaniensis Riek, 1953: 115; Fig. 5. [small stream at Kingston, Tasmania]
Paratya boninensis Satake & Cai, 2005: 306; Figs 1-2. [Oku-ohotaki river, a tributary of the west inlet
stream of Shigure Dam, 210-220 m above sealevel, Chichiima Island, Ogasawara Islands, Japan]
Paratya borealis Volk, 1938: 123; Figs 1-4. [Uluncha River, southern part of the Peter the Great Basin, Japan
Sea]
Paratya bouvieri Roux, 1926a
= Paratya (Xiphatyoida) bouvieri Roux, 1926a: 189; Figs 1-5. [Nouvelle-Caldonie, Rivire du Mt. Pani]
= Paratya (Xiphatyoida) bouvieri ngoiensis Roux, 1926a: 190; Figs 6-8. [Nouvelle-Caldonie, Valle de
Ngo, 200 m alt.]
Parataya caledonica Roux, 1926a
= Parataya (Xiphatyoida) caledonica Roux, 1926a: 192; Figs 9-16. [Nouvelle-Caldonie, Lac en 8]
= Paratya caledonica var. magna Roux, 1926a: 194; Figs 17-19. [Nouvelle-Caldonie, Lac en 8]
Paratya compressa (De Haan, 1844 [in De Haan, 1833-1850])
= Ephyra compressa De Haan, 1844 [in De Haan, 1833-1850]: Plate 46, fg. 7 (1844); 186, Plate O (1849).
Paratya improvisa Kemp, 1917b
= Paratya compressa improvisa Kemp, 1917b: 299; Figs 2-3. [Lake Haruna, near Ikao, altitude about
3000f]
Paratya curvirostris (Heller, 1862a)
= Caridina curvirostris Heller, 1862a: 525. [Aukland]
= Palmon (Leander) fuviatilis Thomson, 1879: 231. [Waikato River; Taieri River; lagoons in Taieri
plain]
Paratya howensis Roux, 1926b
= Paratya (Xiphatyoida) howensis Roux, 1926b: 244. [Big Creek, Lord Howe Island]
Paratya intermedia Roux, 1926a
= Paratya caledonica var. intermedia Roux, 1926a: 195; Figs 20-22. [Nouvelle-Caldonie, Plaine des
Lacs, 200 m alt.]
Paratya martensi Roux, 1925: 146. [Insel Adenare bei Flores]
Paratya norfolkensis Kemp, 1917b
= Paratya australiensis norfolkensis Kemp, 1917b: 305; Fig. 6. [west side of Norfolk Island]
Paratya typa Roux, 1926a
= Paratya (Xiphatyoida) typa Roux, 1926a: 196; Figs 23-31. [Nouvelle-Caldonie, La Madeleine, Plaine
des Lacs, 200 m]
Parisia Holthuis, 1956a
= Parisia Holthuis, 1956a (type species Caridina microphthalma Fage, 1946, by original designation,
gender feminine)
Parisia dentata Gurney, 1984: 584; Figs 12-14. [Dark zone, Grote d'Andafabe, Ankarana Massif, 40 km
north of Ambilob and 60 km south of Diego Suarez, northern province of Diego Suarez]
Parisia deharvengi Cai & Ng, 2009: 1110; Fig. 10. [Gua (cave) Tanete, Kappang, Maros, Sulawesi]
Parisia edentata Holthuis, 1956a: 55. [southern part of Antsingly Mts., near Bekopaka, Mahilaka Prov-
ince, W Madagascar]
Parisia gracilis W.D. Williams, 1964: 94; Figs 1, 2A-H. [caves 16 miles south of of Katherine, Northern
Territories (approx. 1430'S 132E)]
Parisia holthuisi Cai, 2010c: 174; Figs 1-2. [Tigibi, Tari subdistrict, Southern Highlands, Territory of Papua
Australian New Guinea (= Papua New Guinea), 1600 m alt.]
Parisia macrophora Cai & Anker, 2004: 253; Figs 12-14. [Nalubog Cave, Quezon Province, Luzon, Philip-
pines]
Parisia macrophthalma Holthuis, 1956a: 55. [Grote des Fanihy, Ankarana Mts., N of Ambilob, N.-W.
Madagascar]
Parisia microphthalma (Fage, 1946)
= Caridina microphthalma Fage, 1946: 324; Figs 1-2. [grote des Fanihys, Ankarana]
Parisia unguis W.D. Williams, 1964: 99; Figs 2L-S, 3. [caves 16 miles south of of Katherine, Northern Ter-
ritories (approx. 1430'S 132E)]
Potimirim Holthuis, 1954
= Potimirim Holthuis, 1954 (type species Caridina mexicana de Saussure, 1857a, by original designa-
Potimirim americana (Gurin-Mneville, 1855 [in Gurin-Mneville, 1855-1856])
= Caridina americana Gurin-Mneville, 1855 [in Gurin-Mneville, 1855-1856]: xix; Plate 2, Figs 13-
13a. [la Isla de Cuba]
Potimirim glabra (Kingsley, 1878a)
= Atyoida glabra Kingsley, 1878a: 93. [Polvon and Corcuera, west coast of Nicaragua]
= Potimirim brasiliana Villalobos F., 1960b: 275; Figs 1-5. [Ro Arir, Angra dos Reis, Edo. De Ro de
Janeiro, Brasil]
Potimirim mexicana (de Saussure, 1857a)
= Caridina mexicana de Saussure, 1857a: 505. [Vera-Cruz]
Potimirim potimirim (Mller, 1881)
= Atyoida Potimirim Mller, 1881: 117; Figs 1, 8-15. [Itajahy]
Puteonator Gurney, 1987
= Puteonator Gurney, 1987 (type species Puteonator iraqiensis Gurney, 1987, by original designation
Puteonator iraqiensis Gurney, 1987: 162; Figs 1-4. [southern Iraq, Samawa, anchialine habitat at 160 m]
Pycneus Holthuis, 1986b
= Pycneus Holthuis, 1986b (type species Pycneus morsitans Holthuis, 1986b, by original designation
Pycneus morsitans Holthuis, 1986b: 105; Figs 1-2. [cave at Munjingerra, Gibson Desert, Western Australia,
at about 2230'S 12410'E]
Pycnisia Bruce, 1992a
= Pycnisia Bruce, 1992a (type species Pycnisia raptor Bruce, 1992a, by original designation and mono-
Pycnisia bunyip Suzuki & Davie, 2003: 447; Figs 1-4. [Forbes Inferno Cave, Riversleigh, Lawn Hill Na-
tional Park, northwestern Queensland, in shallow subterranean pools]
Pycnisia raptor Bruce, 1992a: 554; Figs 1-6. [Cave 8MD26, Katherine Region, Northern Territory, 1355'S
13228'E, 33.5 m]
Sinodina Liang & Cai, 1999
= Sinodina Liang & Cai, 1999 (type species Caridina gregoriana Kemp, 1923, by original designation,
gender feminine)
Sinodina acutipoda (Liang, 1989)
= Caridina acutipoda Liang, 1989: 282; Figs 1-10. [Lugu Lake, 2743'N 10046'E]
Sinodina angulata Liang, 2002a: 118; Fig. 1. [Longtan stream, Jiangchuan County, Yunnan Province, China]
Sinodina banna (Cai & Dai, 1999)
= Caridina banna Cai & Dai, 1999: 214; Figs 3-4. [Runjinghong, Hinghong County]
= Sinodina bannica Liang, 2004: iv. [Erroneous spelling]
Sinodina bispinosa (Liang, 1990)
= Caridina bispinosa Liang, 1990: 221; Fig. 3. [Yunnan Province, Chuxiong (Jiulongdian)]
Sinodina dianica Liang & Cai, 1999: 583; Figs 3-4. [Malipo village, Wenshan County, south Yunnan]
Sinodina gregoriana (Kemp, 1923)
= Caridina gregoriana Kemp, 1923: 437; Figs 1-2. [Tali-fu (Erhai Lake), near Dali city, Yunnan; lecto-
type designation by Liang & Cai, 1999]
Sinodina heterodactyla (Liang & Yan, 1985a)
= Caridina heterodactyla Liang & Yan, 1985a: 198; Fig. 2. [rivulet, Shizong, Yunnan Province]
Sinodina leptopropoda Liang, 1990
= Caridina leptopropoda Liang, 1990 (sensu Y, 1938 (partim)): 220; Fig. 2. [several localities in Yun-
nan]
Sinodina liiang Liang & Cai, 1999: 587; Fig. 7. [Liiang County, Yunnan Province]
Sinodina wangtai Liang & Cai, 1999: 585; Figs 5-6. [Wangtai village, Liing County]
Sinodina yongshengica Chen & Liang, 2002: 240; Fig. 1. [Taoyuan, Yongsheng County, Yunnan Province,
2675'N 10070'E, 1100 m]
Sinodina yui (Liang & Yan, 1985a)
= Caridina yui Liang & Yan, 1985a: 200; Fig. 4. [Shuanglongyin, Qiubei County, Yunnan Province]
Stygiocaris Holthuis, 1960
= Stygiocaris Holthuis, 1960 (type species Stygiocaris lancifera Holthuis, 1960, by original designation,
gender feminine)
Stygiocaris lancifera Holthuis, 1960: 48; Figs 1-2. [Kuddumurra Well, Yardie Creek Station, North-West
Cape Peninsula, W. Australia]
Stygiocaris stylifera Holthuis, 1960: 54; Fig. 3. [Kuddumurra Well, Yardie Creek Station, North-West Cape
Peninsula, W. Australia]
Syncaris Holmes, 1900
= Syncaris Holmes, 1900 (type species Miersia pacifca Holmes, 1895, by original designation, gender
Syncaris pacifca (Holmes, 1895)
= Miersia pacifca Holmes, 1895: 577; Plate 21, fgs 27-28. [Sonoma County]
Syncaris pasadenae (Kingsley, 1897)
= Caradina pasaden Kingsley, 1897: 98; Plate 3, fgs 1-7. [streams about Pasadena, California]
= Syncaris Trewi Holmes, 1900: 213; Plate 4, fg. 63. [small stream near San Gabriel, Los Angeles
County, California]
Troglocaris Dormitzer, 1853
= Troglocaris Dormitzer, 1853 (type species Troglocaris Schmidtii Dormitzer, 1853 (a junior subjective
synonym of Palaemon anophthalmus Kollar, 1848), by monotypy, gender feminine; name placed on
the Ofcial List of Generic Names in Zoology in Opinion 470 in 1957)
= Troglocaridella Babi, 1922 (type species Troglocaridella hercegovinensis Babi, 1922, by monotypy,
gender feminine)
= Xiphocaridinella Sadowsky, 1930 (type species Xiphocaridinella kutaissiana Sadowsky, 1930, by
= Spelaeocaris Matjai, 1956 (type species Spelaeocaris pretneri Matjai, 1956, by monotypy, gender
feminine)
Troglocaris (Troglocaridella) hercegovinensis (Babi, 1922)
= Troglocaridella hercegovinensis Babi, 1922: 303; Fig. 5. [Hhle Vjeternica bei Zavala (Hercegovina)]
= Troglocaris hercegovinensis Absolon, 1916: 609. [nomen nudum]
Troglocaris (Troglocaris) anophthalmus anophthalmus (Kollar, 1848)
= Palaemon anophthalmus Kollar, 1848: 137. [aus den unterirdischen Gewssern von Krain]
= Troglocaris Schmidtii Dormitzer, 1853: 87. [Grote von Kumpole in Unterkrain]
= Troglocaris Schmidti Schmidti forma typica Fage, 1937: 226. [Type locality not indicated]
Troglocaris (Troglocaris) anophthalmus intermedia Babi, 1922
= Troglocaris schmidti var. intermedia Babi, 1922: 300; Figs 1-4. [Mikainovis Hhle (in der Nhe von
Gornje Dubrave), ungefhr 6 km von Ogulin]
Troglocaris (Troglocaris) bosnica Sket & Zakek, 2009: 799; Figs 7-8. [Bosnia and Herzegovina, Bosanka
Krajina, Luci polje, Suvaja cave]
Troglocaris (Troglocaris) planinensis Birstein, 1948
= Troglocaris schmidti planinensis Birstein, 1948: 5; Fig. 2. [Planina Cave, 9 km from Postoina Cave,
Rieka River, Yugoslavia]
Troglocaris (Spelaeocaris) kapelana Sket & Zakek, 2009: 806; Fig. 11Ska. [Croatia, Jezerane, Obajdini, Obajdi-
nova peina cave]
Troglocaris (Spelaeocaris) neglecta Sket & Zakek, 2009: 808; Figs 11Sne, 12. [Croatia, Novigrad, Karinica
cave near Karin]
Troglocaris (Spelaeocaris) prasence Sket & Zakek, 2009: 804; Figs 9-10, 11Spra. [Bosnia and Herzegovina,
Fatniko polje, Obod cave]
Troglocaris (Spelaeocaris) pretneri (Matjai, 1956)
= Spelaeocaris pretneri Matjai, 1956: 66; Figs 1-2. [einer Hhle aus der sdstlichen Herzegowina (in
der Nhe von Bilee)]
Troglocaris (Xiphocaridinella) ablaskiri Birstein, 1939
= Troglocaris schmidti ablaskiri Birstein, 1939: 961; Figs 1-2. [Abkhasia, cave near Achkshe-Tyz-Gua]
Troglocaris (Xiphocaridinella) fagei Birstein, 1939
= Troglocaris schmidti fagei Birstein, 1939: 964; Figs 3-5. [Abkhasia, cave near Psyrtzkha]
Troglocaris (Xiphocaridinella) jusbaschjani Birstein, 1948
= Troglocaris schmidti jusbaschjani Birstein, 1948: 4; Fig. 1. [Russian Federation, Krasnodarskiy Krai,
Sochi area, cave at Matzesta]
Troglocaris (Xiphocaridinella) kutaissiana (Sadowsky, 1930)
= Xiphocaridinella kutaissiana Sadowsky, 1930: 95. [unterirdische Gewsser in Karsthhlen bei Kutais
(West-Georgien)]
Troglocaris (Xiphocaridinella) osterlof Juzbajan , 1940: 73. [Shakuran]
Typhlatya Creaser, 1936
= Typhlatya Creaser, 1936 (type species Typhlatya pearsei Creaser, 1936, by monotypy, gender femi-
nine)
Typhlatya arfeae Jaume & Brhier, 2005: 390; Figs 3-18. [Rsurgence de Font Estramar, Salses-le-chteau,
Pyrnes-orientales, France]
Typhlatya campecheae H.H.III Hobbs & H.H.Jr. Hobbs, 1976: 6; Figs 3-4. [Grutas de Xtacumbilxunam,
Bolonchenticul, Campeche, Mexico]
Typhlatya consobrina Botoneanu & Holthuis, 1970: 123; Figs 1-2. [Cueva del Agua, Sierra de Cubitas, at
the foot of Cerro Tuabaquei, very near to Finca la Entrada (NE from the town Camagey, prov.
Camagey, in a subterranean lake]
Typhlatya dzilamensis Alvarez, Ilife & Villalobos, 2005: 83; Figs 2-3. [Cenote Buya Uno, Dzilam de Bravo,
Yucatan, Mexico]
Typhlatya elenae Juarrero, 1994: 58; Figs 1-10. [Cueva Perico Sanchez, 5 km al N de Jaguey Grande, Pro-
vincia de Matanzas]
Typhlatya galapagensis Monod & Cals, 1970: 70; Figs 1-46, 50, 53, 55, 58, 60, 63-64. [Ile de Santa Cruz,
versant S-E, eau courante peine sodique, dans une crevasse profonde de 15-20 m, 50 m daltitude
et 2 km de la cte]
Typhlatya garciadebrasi Juarrero de Varona & Ortiz, 2000: 46; Figs 1-3. [Cueva Chicarrones, Bolondrn,
Procincia de Matanzas]
Typhlatya garciai Chace, 1942a: 99; Plate 29. [Potrero del Molino Cave, Las Cuatrocientas, Banes, Oriente
Province, Cuba]
Typhlatya ilifei C.W.J. Hart & Manning, 1981: 447; Figs 29-55. [Tuckers Town cave, Bermuda]
Typhlatya kakuki Alvarez, Ilife & Villalobos, 2005: 86; Figs 4-5. [Shrimp Hole, Salinas Point, Acklins Is-
land, Bahamas]
Typhlatya miravetensis Sanz & Platvoet, 1995: 80; Figs 1-4. [Cave Ullal de la Rambla de Miravet between
the towns of Cabanes and Orpesa, province of Castelln, eastern Spain]
Typhlatya mitchelli H.H.III Hobbs & H.H.Jr. Hobbs, 1976: 2; Figs 1-2. [Cenote Kabahchen, Man, Yucatn,
Mexico]
Typhlatya monae Chace, 1954: 318; Fig. 1. [Mona Island, Puerto Rico, from well 30f deep at El Molino,
about 1 mile southeast of NYA camp at Sardinera]
Typhlatya pearsei Creaser, 1936: 128; Figs 31-41. [Balam Canche Cave, 4.8 km east 0.8 km south of Chichen
Itza, Yucatan]
Typhlatya rogersi Chace & Manning, 1972: 14; Figs 10-11. [Marl Pool, back of Shelly Beach, Ascension Is-
land]
Typhlatya taina Estrada & Gmez, 1987: 3; Figs 2-5. [Cueva de la India, 2 km al E de Puerto Escondido,
Santa Cruz del Norte, La Habana, Cuba]
Typhlatya utilaensis Alvarez, Ilife & Villalobos, 2005: 89; Figs 6-7. [George Gaberels Cave, Utila, Bay Is-
lands, Honduras]
Typhlocaridina Liang & Yan, 1981
= Typhlocaridina Liang & Yan, 1981 (type species Typhlocaridina lanceifrons Liang & Yan, 1981, by
Typhlocaridina lanceifrons Liang & Yan, 1981: 32; Figs 1-19. [Daji Cave, Wuming County, Guangxi]
Typhlocaridina liui Liang & Zhou, 1993: 232; Figs 1(1-3). [Paobingyan Cave, Lingui County, Guangxi
Province]
Typhlocaridina semityphlata Cai, 1995b: 157; Fig. 1. [cave in Guilin, Guangxi Autonomous Region, South
China]
Typhlopatsa Holthuis, 1956a
= Typhlopatsa Holthuis, 1956a (type species Typhlopatsa pauliani Holthuis, 1956a, by original designa-
tion and monotypy, gender feminine)
Typhlopatsa pauliani Holthuis, 1956a: 53. [Mitoho Cave, N.-E. corner of Tsimanampetsotsa Lake, Maha-
faly Province, S.-W. Madagascar]
Superfamily BRESILIOIDEA Calman, 1896
Family AGOSTOCARIDIDAE C.W.J. Hart & Manning, 1986
Agostocaris C.W.J. Hart & Manning, 1986
= Agostocaris C.W.J. Hart & Manning, 1986 (type species Agostocaris williamsi C.W.J. Hart & Man-
ning, 1986, by monotypy, gender feminine)
Agostocaris acklinsensis Alvarez, Villalobos & Ilife, 2004: 370; Figs 2-4. [Jumby Hole Cave, Snug Corner,
Acklins Island, Bahamas]
Agostocaris bozanici Kensley, 1988: 688; Figs 1-3 [Xcan-ha Cenote (Cenote Roja), Cozumel Island, Quin-
tana Roo, Mexico, 80-100 feet]
Agostocaris williamsi C.W.J. Hart & Manning, 1986: 412; Figs 27-47. [Bahamas, 2427'N, 7530'W, Big
Fountain Blue Hole] (Fig. 16)
Fig. 16. Agostocaris williamsi C.W.J. Hart & Manning, 1986. Photo by Thomas M. Ilife.
Family ALVINOCARIDIDAE Christofersen, 1986
Alvinocaridinides Komai & Chan, 2010a
= Alvinocaridinides Komai & Chan, 2010a (type species Alvinocaridinides formosa Komai & Chan,
2010a, by original designation and monotypy, gender feminine)
Alvinocaridinides formosa Komai & Chan, 2010a: 26; Figs 1C, 7-10. [Gueishandao, Yilan County, Taiwan,
2451.231'N 12159.204'E, 252-275 m]
Alvinocaris Williams & Chace, 1982
= Alvinocaris Williams & Chace, 1982 (type species Alvinocaris lusca Williams & Chace, 1982, by
Alvinocaris alexander Ahyong, 2009: 777; Figs 1-3. [Rumble V Seamount, 3608.27-07.96'S 17811.74-
11.70'E, 485-415 m]
Alvinocaris brevitelsonis Kikuchi & Hashimoto, 2000: 136; Figs 1-3. [Depression C of the Minami-Ensei
Knoll, 2823.35'N 12738.38'E, 705 m]
Alvinocaris chelys Komai & Chan, 2010a: 16; Figs 1A-B, 2-6. [Gueishandao, Yilan County, Taiwan,
2449.682'N 1220.254'E, 300-276 m] (Fig. 17)
Alvinocaris dissimilis Komai & Segonzac, 2005a: 1158; Figs 25-26, 29. [Depression C, Minami-Ensei Knoll,
2823.35'N 12738.38'E, 705 m]
Alvinocaris komaii Zelnio & Hourdez, 2009: 55; Figs 1-6. [Kilo Moana, 209'S 7612'E, 2620 m, Eastern Lau
Spreading Center, Lau Basin, southwest Pacifc]
Alvinocaris longirostris Kikuchi & Ohta, 1995: 772-778, Figs 1-7. [Iheya Ridge, Clam Site, Okinawa Trough,
2732.70'N 12658.20'E, 1360 m]
Alvinocaris lusca Williams & Chace, 1982: 137; Figs 1-7. [Galapagos Rif Rose Garden area, 048.25'N
8613.48'W, maximum of 2450 m]
Alvinocaris markensis Williams, 1988: 264; Figs 1-2, 7l. [Mid-Atlantic Rif Valley about 70 km south of
Kane Fracture Zone, 2322.09'N 4457.12'W, 3437 m]
Fig. 17. Alvinocaris chelys Komai & Chan, 2010. Photo by Tin-Yam Chan.
Alvinocaris methanophila Komai, Shank & Van Dover, 2005: 29; Figs 1-4. [ODP site 996, Blake Ridge Dia-
pir, 3229.623'N 7611.467'W, 2155 m]
Alvinocaris muricola Williams, 1988: 268; Figs 3-4, 7e-k. [West Florida Escarpment, 2601'N 8454.61'W,
3277 m]
Alvinocaris niwa Webber, 2004: 5; Figs 1-4. [Rumble V, 368.63-8.57'S 17811.77-11.50'E, 877-655 m]
Alvinocaris stactophila Williams, 1988: 272; Figs 5-6, 7a-d. [north central Gulf of Mexico about 129 km S of
Louisiana, 2746.94'N 9130.34'W, 534 m]
Alvinocaris williamsi Shank & Martin, 2003: 159; Figs 1-3. [Menez Gwen hydrothermal feld, North Atlan-
tic Ocean, 3750.5'N 3131.3'W, 850 m]
Chorocaris Martin & Hessler, 1990
= Chorocaris Martin & Hessler, 1990 (type species Chorocaris vandoverae Martin & Hessler, 1990, by
Chorocaris chacei (Williams & Rona, 1986)
= Rimicaris chacei Williams & Rona, 1986: 455; Figs 8-10. [TAG Hydrothermal Field, Mid-Atlantic
Ridge, 2608.3'N 4449.6'W, 3620-3650 m]
Chorocaris paulexa Martin & Shank, 2005: 186; Figs 1-8. [Homer Vent (347OC black smoker), 1737.220'S
11315.123'W, 2595 m, southern East Pacifc Rise]
Chorocaris vandoverae Martin & Hessler, 1990: 2; Figs 1-2, 3a-e. [Alice springs vent feld, Mariana Back-
Arc Basin, 1812.599'N 14442.431'E, 3640 m]
Mirocaris Vereshchaka, 1997b
= Mirocaris Vereshchaka, 1997b (type species Mirocaris keldyshi Vereshchaka, 1997b (junior subjec-
tive synonym of Chorocaris fortunata Martin & Christiansen, 1995), by monotypy, gender femi-
nine)
Mirocaris fortunata (Martin & Christiansen, 1995)
= Chorocaris fortunata Martin & Christiansen, 1995: 221; Figs 1-3. [Vent site 3, Lucky Strike hydro-
thermal vent, Azores, 3717.6'N 3216.5'W, 1624 m]
= Mirocaris keldyshi Vereshchaka, 1997b: 431; Figs 1-5. [TAG location, 2609'N 4450'W, 3650 m]
Mirocaris indica Komai, Martin, Zala, Tsuchida & Hashimoto, 2006: 110; Figs 1-5. [Central Indian Ridge,
Kairei Field, 2519.2'S 7002.4'E, 2422 m]
Nautilocaris Komai & Segonzac, 2004
= Nautilocaris Komai & Segonzac, 2004 (type species Nautilocaris saintlaurentae Komai & Segonzac,
Nautilocaris saintlaurentae Komai & Segonzac, 2004: 1179-1188; Figs 2-6. [North Fii Basin, White Lady
site, 1659.50'S 17355.47'E, 2000 m]
Opaepele Williams & Dobbs, 1995
= Opaepele Williams & Dobbs, 1995 (type species Opaepele loihi Williams & Dobbs, 1995, by mono-
Opaepele loihi Williams & Dobbs, 1995: 229; Figs 1-3. [Loihi Seamount, Hawaii, 1855'N 15516'W, 980 m]
Opaepele susannae Komai, Giere & Segonzac, 2007: 240; Figs 2-7. [Lilliput, southern Mid-Atlantic Ridge,
0932.845'S 1312.546'W, 1500 m, mussel feld with difuse vent fuids]
Opaepele vavilovi Lunina & Vereshchaka, 2010: 70; Figs 1-2. [Mid-Atlantic Ridge, Broken Spur vent site,
stn 4797]
Rimicaris Williams & Rona, 1986
= Rimicaris Williams & Rona, 1986 (type species Rimicaris exoculata Williams & Rona, 1986, by origi-
= Iorania Vereshchaka, 1996 (type species Iorania concordia Vereshchaka, 1996 (junior subjective syn-
onym of Rimicaris exoculata Williams & Rona, 1986), by monotypy, gender feminine)
Rimicaris exoculata Williams & Rona, 1986: 448; Figs 5-7. [TAG Hydrothermal Field, Mid-Atlantic Ridge,
2608.3'N 4449.6'W, 3620-3650 m]
= Iorania concordia Vereshchaka, 1996: 954; Figs 1-5. [TAG location, 2609N 4450W, point D, 3650 m]
= Rimicaris aurantiaca Martin, Signorovitch & Patel, 1997: 400; Figs 1-5. [Snake Pit hydrothermal vent
feld, Mid-Atlantic Ridge, 2322.1'N 4457.0'W]
Rimicaris kairei Watabe & Hashimoto, 2002: 1168; Figs 1-4. [The Central Indian ridge, Indian Ocean, the
Kairei Field, 2519.16'S 7002.40'E, 2454 m]
Shinkaicaris Komai & Segonzac, 2005a
= Shinkaicaris Komai & Segonzac, 2005a (type species Alvinocaris leurokolos Kikuchi & Hashimoto,
Shinkaicaris leurokolos (Kikuchi & Hashimoto, 2000)
= Alvinocaris leurokolos Kikuchi & Hashimoto, 2000: 141; Figs 4-7. [Depression C of the Minami-
Ensei Knoll, 2823.35'N 12738.38'E, 705 m]
Family BRESILIIDAE Calman, 1896
Bresilia Calman, 1896
= Bresilia Calman, 1896 (type species Bresilia atlantica Calman, 1896, by monotypy, gender feminine;
Bresilia antipodarum Bruce, 1990a: 855; Figs 6-10. [Tasman Sea, of Cape Freycinet, Tasmania, 422.20'S
14838.70'E, 800 m]
Bresilia atlantica Calman, 1896: 7; Plate 1; Plate 2, fgs 15-18. [south west of Ireland, 750 fms]
Bresilia briankensleyi Bruce, 2005a: 176; Figs 1-3. [Egyptian Red Sea, 2285.2'N 3645.9'E to 2234'N
3646.2'E, 750-753 m]
Bresilia corsicana Forest & Cals, 1977: 551; Figs 1-20. [canal de Corse, 545 milles dans le 285 du cap Trat-
toja, sud-ouest de lle Capraia, 450 m]
Bresilia gibbosa Komai & Yamada, 2010: 42; Figs 1-5. [Apo-gama Cave, Onna Village, Okinawa Island,
Ryukyu Islands, 30 m]
Bresilia plumifera Bruce, 1990b: 1; Figs 1-4. [Taupo Sea Mount, Western Tasman Sea, 3314.21'S 5610.68'E,
133 m]
Bresilia rufoculus Komai & Yamada, 2011: 72; Figs 1-6. [Ohoba Cave, Ie Island, Okinawa Islands] (Fig. 18)
Fig. 18. Bresilia rufoculus Komai & Yamada, 2011. Photo by Tomoyuki Komai.
Bresilia saldanhai Calado, Chevaldonn & dos Santos, 2004: 191; Figs 1-4. [underwater cave at Ponta ga-
rajau, south of Madeira island, eastern Atlantic, 32.63N 16.85W, 15 m]
Encantada Wicksten, 1989a
= Encantada Wicksten, 1989a (type species Encantada spinoculata Wicksten, 1989a, by monotypy, gen-
der feminine)
Encantada spinoculata Wicksten, 1989a: 667; Figs 1-2. [Beagle Island, east of Jervis Island, Galapagos,
026'S 9040'W, 55-92 m]
Family DISCIADIDAE Rathbun, 1902c
Discias Rathbun, 1902c (Fig. 19)
= Anisocaris Ortmann, 1893 (type species Anisocaris dromedarius Ortmann, 1893 (probably an invalid
senior subjective synonym of Discias atlanticus Gurney, 1939b), by monotypy, gender feminine;
name suppressed for the purposes of the Principle of Priority, but not for those of the Principle of
Homonymy and placed on the Ofcial Index of Rejected and Invalid Generic Names in Zoology in
= Discias Rathbun, 1902c (type species Discias serrifer Rathbun, 1902c, by monotypy, gender mascu-
line)
Discias atlanticus Gurney, 1939b: 388; Figs 1-13. [in plankton at night in the shallow waters of the Reach
close to the Bermuda Biological Station]
? = Anisocaris dromedarius Ortmann, 1893: 74; Plate 4, fgs 3-3z. [Plankton Expedition, Nrdl.
Aequa torialstrom, JN 148 (10.2N 22.2W, 0-400 m); Sdl. Aequatorialstrom, JN 218 (3.8S 32.6W,
0-400 m)]
Discias brownae Kensley, 1983: 8; Figs 6-9. [Green Point, Port Jackson, New South Wales, 3350'S 15119'E,
9.8 m]
Fig. 19. Discias spec. Photo by Leslie Harris.
Discias exul Kemp, 1920: 138; Figs 1-3. [Port Blair, Andaman Islands, on the reef at the N end of Ross Is-
land, low water]
= Discias mvitae Bruce, 1976b: 119; Figs 1-5. [Fort Jesus, Mombasa, 404.0'S 3942.18'E, 1 m]
Discias musicus Holthuis, 1981: 787; Figs 1-2. [Lagoon west of Saipan, Marianas Islands, immediately
shoreward of barrier reef fat]
= Discias musicus Holthuis, 1953b: 52. [nomen nudum]
Discias pascuensis Fransen, 1987: 501; Figs 1-3. [Tahai, west coast of Easter Island, Chile, 39 m]
Discias serratirostris Lebour, 1949a: 1107; Figs 1-2. [about three miles of Castle Roads, Bermuda, near
botom, at about 100f or more]
Discias serrifer Rathbun, 1902c: 290; Figs 1-4. [Tagus Cove, Albemarle Island, on the reef north of Tagus
Hill]
Discias vernbergi Boothe & Heard, 1987: 506; Figs 1-3. [65 nautical miles W of Egmont Key, Florida,
2737'N 8358'W, 55 m]
Kirnasia Burukovsky, 1988
= Kirnasia Burukovsky, 1988 (type species Kirnasia nesisi Burukovsky, 1988, by original designation,
gender feminine)
Kirnasia nesisi Burukovsky, 1988: 457; Fig. 1. [4239'N, 2845'W, 850-1200 m]
Kirnasia siedlecki Burukovsky, 1988: 458; Fig. 2. [3618'N, 1950'W, 874 m]
Lucaya Chace, 1939
= Lucaya Chace, 1939 (type species Lucaya bigelowi Chace, 1939, by monotypy, gender feminine)
Lucaya bigelowi Chace, 1939: 34. [east of Great Abaco Island, Bahamas, 2612'N, 7626'W, 2610 fms]
Tridiscias Kensley, 1983
= Tridiscias Kensley, 1983 (type species Tridiscias transkei Kensley, 1983, by monotypy, gender mascu-
line)
Tridiscias transkei Kensley, 1983: 18; Figs 15-17. [of Transkei, South Africa, 3159'S 2922.5'E, 150-200 m]
Family PSEUDOCHELIDAE De Grave & Moosa, 2004
Pseudocheles Chace & Brown, 1978
= Pseudocheles Chace & Brown, 1978 (type species Pseudocheles enigma Chace & Brown, 1978, by
Pseudocheles chacei Kensley, 1983: 22; Figs 18-22. [Looe Key, Florida, 6 m]
Pseudocheles enigma Chace & Brown, 1978: 757; Figs 1-4. [west side of lagoon, Lizard Island, Queensland,
Australia, 1-15 m]
Pseudocheles neutra De Grave & Moosa, 2004: 1; Figs 1-4. [Kaledupa Reef, Kaledupa, Tukangbesi Archi-
pelago, south-eastern Sulawesi, Indonesia, 10 m]
Superfamily NEMATOCARCINOIDEA Smith, 1884
Family EUGONATONOTIDAE Chace, 1937a
Eugonatonotus Schmit, 1926a
= Gonatonotus A. Milne-Edwards, 1881b (type species Gonatonotus crassus A. Milne-Edwards, 1881b,
by monotypy, gender masculine; invalid junior synonym of Gonatonotus Adams & White, 1847
(Crustacea Brachyura); name placed on the Ofcial Index of Rejected and Invalid Generic Names in
= Eugonatonotus Schmit, 1926a (nomen novum for Gonatonotus A. Milne-Edwards, 1881b, type spe-
cies therefore Gonatonotus crassus A. Milne-Edwards, 1881b, gender masculine; name placed on the
= Gomphonotus Chace, 1936 (nomen novum for Gonatonotus A. Milne-Edwards, 1881b, gender mas-
culine; name placed on the Ofcial Index of Rejected and Invalid Generic Names in Zoology in
= Galatheacaris Vereshchaka, 1997a (type species Galatheacaris abyssalis Vereshchaka, 1997a, by origi-
nal designation and monotypy, gender feminine)
Eugonatonotus crassus (A. Milne-Edwards, 1881b) (Fig. 20)
= Gonatonotus crassus A. Milne-Edwards, 1881b: 10. [Grenade par 262 brasses de profondeur]
= Gonatonotus crassus var. longirostris A. Milne-Edwards, 1883: Planche 34. [Of Sand Key]
Eugonatonotus chacei Chan & Yu, 1991a: 144; Fig. 1; Plate 1A. [Ta-Shi, I-Lan County, Taiwan]
= Galatheacaris abyssalis Vereshchaka, 1997a: 365; Figs 1-9. [Celebes Sea, 0150'N 11920'E, 4940-4970 m]
Family NEMATOCARCINIDAE Smith, 1884
Lenzicarcinus Burukovsky, 2005
= Lenzicarcinus Burukovsky, 2005 (type species Lenzicarcinus struncki Burukovsky, 2005, by original
designation and monotypy, gender masculine)
Lenzicarcinus struncki Burukovsky, 2005: 572; Figs 1-2. [Atlantic Ocean, south of Ascension Island, 1211'S
6W, 2000 m]
Lipkius Yaldwyn, 1960
= Lipkius Yaldwyn, 1960 (type species Lipkius holthuisi Yaldwyn, 1960, by original designation and
Lipkius holthuisi Yaldwyn, 1960: 16; Fig. 1. [4139'30''S 17517'E, between 50 and 200 fms]
Nematocarcinus A. Milne-Edwards, 1881b (Fig. 21)
= Nematocarcinus A. Milne-Edwards, 1881b (type species Nematocarcinus cursor A. Milne-Edwards,
1881b, by monotypy, gender masculine; name placed on the Ofcial List of Generic Names in Zool-
ogy in Opinion 470 in 1957)
= Eumiersia Smith, 1882 (type species Eumiersia ensifera Smith, 1882, by monotypy, gender feminine)
= Stochasmus Spence Bate, 1888 (type species Stochasmus exilis Spence Bate, 1888, by monotypy,
gender masculine)
Nematocarcinus africanus Crosnier & Forest, 1973: 101; Figs 29a, 30a-e, 31a-b. [905'N 1510'W, 310-380 m]
Fig. 20. Eugonatonotus crassus (A. Milne-Edwards, 1881). Photo Pillsbury Expedition sta 936b.
Nematocarcinus agassizii Faxon, 1893: 204. [Albatross stn 3358 (0630'N 08144'W, 555 fms); lectotype des-
ignation by Burukovsky, 2001]
Nematocarcinus batei Burukovsky, 2000a: 284; Figs 1b-c. [Challenger stn 237, 3437'N 14032'E, near Yo-
gama, Japan, 3470 m]
Nematocarcinus bituberculatus Chace, 1986: 69; Fig. 37. [west of Halmahera, Indonesia, 012'15"N
12729'30"E, 527 m]
Nematocarcinus chacei Burukovsky, 2002a: 5; Fig. 1. [Indian Ocean, Gulf of Aden, 1131'40''N 4246'40''E,
335 m]
Nematocarcinus challengeri Burukovsky, 2006a: 896; Fig. 1. [Challenger stn 250, Pacifc Ocean, 3749'N
16647'W, 5477 m]
Nematocarcinus combensis Burukovsky, 2000b: 1038; Fig. 2. [south-west part of Indian Ocean, Comb Bank,
1215'S 17728'W, 800-810 m]
Nematocarcinus crosnieri Burukovsky, 2000c: 666; Fig. 3. [New Caledonia, 2319'S 16759'E 850-920 m]
Nematocarcinus cursor A. Milne-Edwards, 1881b: 14. [la Mer des Antilles, 500 brasses]
Nematocarcinus ensifer (Smith, 1882)
= Eumiersia ensifera Smith, 1882: 77; Plate 13, Figs 1-9. [Blake stns 305, 4133'15''N 6551'25''W, 810
fms; 308, 4124'45''N 6535'30''W, 1242 fms; 330, 3141'0''N 7435'0''W, 1047 fms; 339, 3816'45''N
7310'30''W, 1186 fms; 340, 3925'30''N 7058'40''W, 1394 fms; 341, 3938'20''N 7056'0''W, 1241
fms]
Nematocarcinus evansi Burukovsky, 2000d: 1291; Fig. 2. [south-west Australia, 2016.3'S 11313.5'E, 913-
916 m]
Nematocarcinus exilis (Spence Bate, 1888)
= Stochasmus exilis Spence Bate, 1888: 823; Plate 132, fg. 14. [Challenger stn 84, 3038'N 185'W, of the
Canary Islands]
Nematocarcinus faxoni Burukovsky, 2001: 1429; Figs 1-2. [Eastern Pacifc, 1633'N 9952'W, 1208 m]
Fig. 21. Nematocarcinus spec. Photo by Tin-Yam Chan.
Nematocarcinus gladius Macpherson, 1984: 67; Figs 13-14, 15a-b. [3437'S 1703'E, 1580-1620 fms; exact
locality not indicated in publication, information from IZIKO collection label]
Nematocarcinus gracilipes Filhol, 1884: 232; Fig. 1. [pch 850 mtres de profondeur; no further indica-
tion of type locality]
Nematocarcinus gracilis Spence Bate, 1888: 815; Plate 132, fg. 8. [Challenger stn 174c, 197'50"S 17819'35"E,
of Kandavu, Fii Islands, 610 fms; lectotype designation by Burukovsky, 2007a]
Nematocarcinus hanamuri Burukovsky, 2000d: 1290; Fig. 1. [south-west Australia, 2016.3'S, 11313.5'E,
913-916 m]
Nematocarcinus hiatus Spence Bate, 1888: 821; Plate 132, fg. 12. [Challenger stn 169, 3734'S 17922'E, of
New Zealand, 700 fms]
Nematocarcinus kaiensis Burukovsky, 2000b: 1040; Fig. 3. [Indonesia, Selan-Timur Archipelago, 0515'S
13359'E, 769-800 m]
Nematocarcinus lanceopes Spence Bate, 1888: 804; Plate 131. [Challenger stn 152, 6052'S 8020'E, Antarctic
Sea, 1260 fms]
= Acanthephyra antarctica Bage, 1938: 6; Plate 4, fg. 1. [6313'S 10142'E, 870 fms]
Nematocarcinus longirostris Spence Bate, 1888: 806; Plate 132, fg. 2. [Challenger stn 237, 3437'N 14032'E,
near Yokohama, Japan, 1875 fms]
Nematocarcinus machaerophorus Burukovsky, 2003: 116; Fig. 33. [Pacifc Ocean, Eiao Island, 0752'S
14039'W, 1000 m]
= Nematocarcinus machaerophorus Burukovsky, 2004: 1181; Figs a-e. [Pacifc Ocean, Eiao Island,
0752'S 14039'W, 1000 m]
Nematocarcinus manningi Burukovsky, 2003: 118; Fig. 34. [North Pacifc Ocean, 3506'45''N 13919'45''E,
1138 m]
Nematocarcinus novaezealandicus Burukovsky, 2006b: 441; Fig. 1. [Chatham Rise, 4251.7'-4249.5'S
17523.3'-17546.1'W, 1057 m]
Nematocarcinus nudirostris Burukovsky, 1991a: 41; Fig. 1(5-8). [933'5"-935'S 5952'7"-5958'8"E, 830-430 m]
= Nematocarcinus rectirostris Burukovsky, 1991a: 39; Fig. 1(1-4). [2528'-2529'S 3514'-3511'E, 715-
700 m]
Nematocarcinus ovalis Komai & Segonzac, 2005b: 355; Figs 8-10. [East Pacifc Rise, 1245.50'N 10355.80'W,
2558-2619 m (c. 120 m above botom)]
Nematocarcinus paucidentatus Spence Bate, 1888: 816; Plate 132, fg. 9. [Challenger stn 174c, of Kandavu,
Fii Islands, 197'50''S 17819'35'', 610 fms]
= Nematocarcinus alisae Burukovsky, 2000e: 903; Fig. 4. [New Caledonia, 2105'S 16550'E, 960-1100 m]
Nematocarcinus parvus Burukovsky, 2000f: 1163; Fig. 7. [Madagascar, 1226'-1227'S 4208'5''-4813'E, 600-
705 m]
Nematocarcinus poupini Burukovsky, 2007b: 1; Fig. 1. [Austral Islands, East of Rapa Island, 2738.2'S
14414.4'W, 840-1200 m]
Nematocarcinus productus Spence Bate, 1888: 810; Plate 132, fg. 5. [Challenger stns 205, 1642'N 11922'E,
of Luzon, Philippine Islands, 1050 fms; 195, 421'S 1297'E, of Banda Island, 1425 fms; 237, 3437'N
14032'E, near Yokohama, Japan, 1875 fms; 176, 1830'S 17352'E, of the New Hebrides, 1450 fms]
= Nematocarcinus tenuipes Spence Bate, 1888: 812 (partim); Plate 132, fg. 6. [Challenger stn 218, 233'S
1444'E, of New Guinea, 1070 fms ]
= Nematocarcinus intermedius Spence Bate, 1888: 821; Plate 132, fg. 13. [Challenger stn 218, 233'S
1444'E, of New Guinea, 1070 fms]
Nematocarcinus proximatus Spence Bate, 1888: 808; Plate 132, fg. 3. [Challenger stn 146, 3342'S 7818'W,
west of Valparaiso, 1375 fms; lectotype designation by Burukovsky, 2003]
Nematocarcinus pseudocursor Burukovsky, 1990: 194; Fig. 2B. [2507'S 9926'W, 780-790 m]
Nematocarcinus pseudogracilis Burukovsky, 2007a: 807; Figs 5-7. [Pacifc Ocean, Austral Islands, Tubuai,
2312.3'S 14934.4'W, 750-1000 m]
Nematocarcinus richeri Burukovsky, 2000f: 1155; Figs 1-3. [Indonesia, Kai Islands, 0515'S 13301'E, 605-
576 m]
Nematocarcinus romenskyi Burukovsky, 2000a: 288; Fig. 5. [Atlantic Ocean, Discovery Bank, 423'8''S
0003'W, 1030-1050 m]
Nematocarcinus rotundus Crosnier & Forest, 1973: 103; Figs 29c, 30f-i, 31e-f. [2843'N 8714'30''W, 960 m]
Nematocarcinus safari Burukovsky, 2000c: 662; Fig. 1. [south-west part of Indian Ocean, Madagascar sink,
3042'9''S 4820'6''E, 4297 m]
Nematocarcinus serratus Spence Bate, 1888: 819; Plate 132, fg. 11. [Challenger stn 169, 3734'S 17922'E, of
New Zealand, 700 fms]
Nematocarcinus sigmoideus Macpherson, 1984: 63; Figs 10-12 [area del Cabo entre 840 y 2200 m; Banco
Valdivia, entre 754 y 940 m; costas de Namibia (2819'S 1418,6'E)]
Nematocarcinus subtegulisfactus Burukovsky, 2000c: 664; Fig. 2. [Philippines, 1205'N 12214'E, 1404 m]
Nematocarcinus subtilis Burukovsky, 2000b: 1042; Fig. 4. [Philippines, 1158'N 12106'E, 448-466 m]
Nematocarcinus tenuipes Spence Bate, 1888: 812 (partim); Plate 132, fg. 6. [Challenger stns 235, 3407'N
1380'E, south of Japan, 565 fms; 218, 233'S 14404'E, near the Admiralty Islands, 1070 fms; 232,
3511'N 13928'E, Japan, 345 fms]
= Nematocarcinus parvidentatus Spence Bate, 1888: 814; Plate 132, fg. 7. [Challenger stn 237, 3437'N
14032'E, near Yokohama, Japan, 1875 fms]
= Nematocarcinus serratirostris Burukovsky, 1991a: 41; Fig. 1(9-11). [2458'-2505'S 3540'-3534'E,
1010-910 m]
Nematocarcinus tenuirostris Spence Bate, 1888: 817; Plate 132, fg. 10. [Challenger stns 174c, 197'50"S
17819'35"E, of Kandavu, Fii Islands, 610 fms; 214, 433'N 1276'E, south of the Philippine Islands,
500 fms]
= Nematocarcinus tenuirostris var. sibogae De Man, 1917: 279. [1048'.6S 12323'.1E, 918 m]
Nematocarcinus tuerkayi Burukovsky, 2005: 573; Fig. 3. [Indian Ocean, Gulf of Aden, 1252.5'N 4553.3'E,
1185 m]
Nematocarcinus undulatipes Spence Bate, 1888: 801; Plate 130. [Challenger stn 200, 647'N 12228'E, of
Sibago, Philippine Islands, 250 fms; lectotype designation by Burukovsky, 2002a]
Nematocarcinus webberi Burukovsky, 2006b: 444; Fig. 2. [Of New Zealand, 4241.7'S 17428'E, 1723-1549 m]
Nematocarcinus yaldwyni Burukovsky, 2006b: 445; Fig. 3. [NE of Ritchie Bane, New Zealand, 3947.9'S
17817.9'E, 832-788 m]
Nigmatullinus Burukovsky, 1991a
= Nigmatullinus Burukovsky, 1991a (type species Nematocarcinus acanthitelsonis Pequegnat, 1970, by
original designation and monotypy, gender masculine)
Nigmatullinus acanthitelsonis (Pequegnat, 1970)
= Nematocarcinus acanthitelsonis Pequegnat, 1970: 69; Figs 4-3, 4-4. [Southwestern Gulf of Mexico,
2336'N 9357'W, 2045 fms]
Segonzackomaius Burukovsky, 2011
= Segonzackomaius Burukovsky, 2011 (type species Nematocarcinus burukovskyi Komai & Segonzac,
Nematocarcinus burukovskyi Komai & Segonzac, 2005b: 346; Figs 2-7. [South East Pacifc Rise, Garret Seg-
ment, Wormwood site, 1734.91'S 11314.68'W, 2595 m]
Nematocarcinus altus Spence Bate, 1888: 809; Plate 132, fg. 4. [Challenger stn 198, 255'N 12453'E, south
of the Philippine Islands, 2150 fms]
Family RHYNCHOCINETIDAE Ortmann, 1890
Cinetorhynchus Holthuis, 1995
= Rhynchocinetes (Cinetorhynchus) Holthuis, 1995 (type species Rhynchocinetes rigens Gordon, 1936,
by original designation, gender masculine)
Cinetorhynchus brucei Okuno, 2009a: 940; Figs 1-3. [Imazuni, Kume-jima Island, Rykyu Islands, Japan,
2621.8'N 12649.6'E, 18 m]
Cinetorhynchus concolor (Okuno, 1994a)
= Rhynchocinetes concolor Okuno, 1994a: 66; Figs 1-2, 3A, 4A-D. [2611.2'N 12716.8'E, Hizushi-hama,
Aka-jima Islet, Kerama Group, Ryukyu Islands, 3 m]
Cinetorhynchus erythrostictus Okuno, 1997a: 36; Plate 1A-B; Figs 2A, 3, 4A-C. [Ryukyu Islands, Aka Har-
bor, Aka-jima Islet, Kerama Group, 2611.2'N 12717.1'E, 3 m]
Cinetorhynchus fasciatus Okuno & Tachikawa, 1997: 16; Figs 1-4. [Tenno-ura beach, Chichi-jima Island,
Ogasawara Islands, Japan, 5 m]
Cinetorhynchus hawaiiensis Okuno & Hoover, 1998: 33; Figs 1-3, 4A-B, 5A. Hawaiian Islands, 1939.0'N
15600.0'W, Kailua Harbor, Hawaii, 8 m]
Cinetorhynchus hendersoni (Kemp, 1925)
= Rhynchocinetes hendersoni Kemp, 1925: 265; Figs 3-7. [Pamban and Kilakarai, Gulf of Manaar]
= Rhynchocinetes intermedius Edmondson, 1952: 72; Fig. 3. [of the southwest coast of Oahu, at about
16 fms]
= Rhynchocinetes marshallensis Edmondson, 1952: 75; Figs 4-6. [Ebenina, Eniwetok, Marshall Islands]
Cinetorhynchus hiati (Holthuis & Hayashi, 1967)
= Rhynchocinetes hiati Holthuis & Hayashi, 1967: 162; Figs 1-2. [Formosa, Kosho Bay, southern ex-
tremity of Formosa, 3 or 4 m]
= Rhynchocinetes hiati Holthuis, 1953b: 54. [nomen nudum]
Cinetorhynchus manningi Okuno, 1996a: 725; Figs 1-2. [Caribbean Sea, Virgin Islands, Eagle Shoal, 10.5
m] (Fig. 22)
Cinetorhynchus reticulatus Okuno, 1997a: 49; Plate 1G-H; Figs 10-11, 12A-C. [Loyalty Islands, Banya Islet,
Uvea Island, 2035.8'S 16616.7'E, 27 m]
Cinetorhynchus rigens (Gordon, 1936)
= Rhynchocinetes rigens Gordon, 1936: 76; Figs 1-4, 5e. [Potinha Bay, Madeira]
Cinetorhynchus striatus (Nomura & Hayashi, 1992)
= Rhynchocinetes striatus Nomura & Hayashi, 1992: 199; Figs 1-4. [Kadena Port, Okinawa Island, 1-10 m]
Rhynchocinetes H. Milne Edwards, 1837
= Rhynchocinetes H. Milne Edwards, 1837 (type species Rhynchocinetes typus H. Milne Edwards,
Rhynchocinetes albatrossae Chace, 1997: 28; Figs 15-16. [Surigao Strait, east of Leyte, 1027'30"N 12518'E,
123 m]
Rhynchocinetes australis Hale, 1941: 270; Fig. 8. [Edithburgh, South Australia]
Fig. 22. Cinetorhynchus manningi Okuno, 1996. Photo by Arthur Anker.
Rhynchocinetes balssi Gordon, 1936: 85; Fig. 7a-b; nomen novum for Rhynchocinetes typus sensu Balss,
1922 nec H. Milne Edwards, 1837. [Masatierra]
Rhynchocinetes brucei Okuno, 1994b: 29; Figs 1-3, 4a-b; Plate 1. [Long Ke Wan, Hong Kong, 8 m]
Rhynchocinetes conspiciocellus Okuno & Takeda, 1992: 64; Figs 1-3, 4A-D; Plate 1A-B. [Sokodo Beach,
Hachio-jima Island, Izu Islands, Japan, 3307'N 13949'E, 1 m]
Rhynchocinetes durbanensis Gordon, 1936: 83; Figs 5b-b, 7c-d; nomen novum for Rhynchocinetes typus
sensu Stebbing, 1917c nec H. Milne Edwards, 1837. [Durban]
Rhynchocinetes enigma Okuno, 1997b: 13; Figs 1-3. [3417'S 13242'E, Great Australian Bight, approx. 15
km west-south-west of Pearson Islands, 140-160 m]
Rhynchocinetes holthuisi Okuno, 1997c: 43; Figs 1-3, 4a-e. [Aqaba, Gulf of Aqaba, Jordan]
Rhynchocinetes ikatere Yaldwyn, 1971: 87. [of Mayor Island, Bay of Plenty, 3715'S 17612'E, 80-120 fms]
Rhynchocinetes kuiteri Tiefenbacher, 1983: 121; Figs 1-3. [before Portsea at the mouth of Port Phillip Bay,
Victoria]
Rhynchocinetes rathbunae Okuno, 1996b: 309; Figs 1-3. [Hawaiian Islands, Oahu, Waimea Bay, 2138.0'N
1584.0'W]
Rhynchocinetes serratus (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840])
= H.[ippolyte] serratus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 377. [la baie de
Jarvis]
= Rhynchocinetes rugulosus Stimpson, 1860a: 36. [in portu Jacksoniensi Australiae]
Rhynchocinetes typus H. Milne Edwards, 1837: 168; Plate 4c. [Type locality not indicated]
Rhynchocinetes uritai Kubo, 1942b: 30; Figs 1-3. [Kominato, Tiba Prefecture; lectotype designation by
Okuno & Takeda, 1992]
Family XIPHOCARIDIDAE Ortmann, 1895
Xiphocaris von Martens, 1872
= Xiphocaris von Martens, 1872 (type species Hippolyte elongatus Gurin-Mneville, 1855 [in Gurin-
Mneville, 1855-1856], by monotypy, gender feminine); name placed on the Ofcial List of Generic
Names in Zoology in Opinion 470 in 1957)
Fig. 23. Xiphocaris elongata (Gurin-Mneville, 1855). Photo by Chris Lukhaup.
Xiphocaris elongata (Gurin-Mneville, 1855 [in Gurin-Mneville, 1855-1856]) (Fig. 23)
= Hippolyte elongatus Gurin-Mneville, 1855 [in Gurin-Mneville, 1855-1856]: xx; Plate 2, Figs 16-
16a. [costas de Cuba]
= Oplophorus americanus de Saussure, 1858: 56; Fig. 31. [embouchure des rivires de lIle de Hati]
= Xiphocaris gladiator Pocock, 1889: 18; Plate 2, fg. 6. [Laiou]
= Xiphocaris gladiator var. intermedia Pocock, 1889: 19; Plate 2, fg. 7. [Laiou]
= Xiphocaris brevirostris Pocock, 1889: 20; Plate 2, fgs 5-5a. [Dominica]
= Xiphocaris elongata typica Ortmann, 1895: 400. [Type locality not indicated]
Xiphocaris gomezi Juarrero de Varona, 1993: 2; Figs 2-4. [Agua Rev, Sierra Maestra, provincia Santiago
de Cuba, Cuba, altura aproximada de 1000 m snm]
Superfamily PSALIDOPODOIDEA Wood-Mason [in Wood-Mason & Alcock, 1892]
Family PSALIDOPODIDAE Wood-Mason [in Wood-Mason & Alcock, 1892]
Psalidopus Wood-Mason [in Wood-Mason & Alcock, 1892]
= Psalidopus Wood-Mason [in Wood-Mason & Alcock, 1892] (type species Psalidopus Huxleyi Wood-
Mason [in Wood-Mason & Alcock, 1892] designated by Holthuis, 1955b, gender masculine; name
placed on the Ofcial List of Generic Names in Zoology in Opinion 470 in 1957)
Psalidopus barbouri Chace, 1939: 36. [Nicholas Channel, south of Cay Sal Bank, 2321'N 7958'W, 300-315
fms]
Psalidopus huxleyi Wood-Mason [in Wood-Mason & Alcock, 1892] (Fig. 24)
= Psalidopus Huxleyi Wood-Mason [in Wood-Mason & Alcock, 1892]: 273; Plate 14, fgs 1-2, 7. [7.5
miles E of N Cinque Island, Andaman Sea, 490 fms]
= Psalidopus spiniventris Wood-Mason [in Wood-Mason & Alcock, 1892]: 274; Plate 14, fgs 3-6a, 8;
Plate 15. [8 miles SE of Cinque Island, Andaman Sea, 500 fms]
= Psalidopus japonensis Kubo, 1952: 91; Figs 1-2; Plate 5. [of Dai-zaki, Kii Peninsula, ca. 530 m]
Psalidopus tosaensis Toriyama & Horikawa, 1993: 1; Figs 1-3, 5. [of Muroto Cape, Kochi Pref., Japan,
3213'12''N 13401'36''E, 2765-2881 m]
Fig. 24. Psalidopus huxleyi Wood-Mason [in Wood-Mason & Alcock, 1892]. Photo by Tin-Yam Chan.
Superfamily STYLODACTYLOIDEA Spence Bate, 1888
Family STYLODACTYLIDAE Spence Bate, 1888
Bathystylodactylus Hanamura & Takeda, 1996
= Bathystylodactylus Hanamura & Takeda, 1996 (type species Stylodactylus bathyalis Cleva, 1994, by
Bathystylodactylus bathyalis (Cleva, 1994)
= Stylodactylus bathyalis Cleva, 1994: 56; Fig. 2. [Bligh Canyon, Coral Sea, 1223'S 14608'E. 3515-
3502 m]
Bathystylodactylus echinus Wicksten & Martin, 2004: 377; Figs 1-5. [basin of Magdalena Bay, Baja Califor-
nia, Mexico, 2435'N 11325'W, 3563-3621 m]
Bathystylodactylus infatus Hanamura & Takeda, 1996: 930; Figs 1-3. [of east coast of Taiwan, 2342'03''N
12345'08''E, 3436-3452 m]
Neostylodactylus Hayashi & Miyake, 1968a
= Neostylodactylus Hayashi & Miyake, 1968a (type species Stylodactylus amarynthis De Man, 1902, by
Neostylodactylus afnis Hayashi & Miyake, 1968a: 605; Fig. 7. [Korea Strait, 3334.9'N 12825.2'E, 120 m]
Neostylodactylus amarynthis (De Man, 1902)
= Stylodactylus amarynthis De Man, 1902: 897; Plate 27, fgs 64-64b. [Ternate]
Neostylodactylus investigatoris (Kemp, 1925)
= Stylodactylus investigatoris Kemp, 1925: 260; Figs 1-2. [2 miles of Great Torres Island, Mergui 40 fms]
Neostylodactylis litoralis Okuno & Tachikawa, 2000: 39; Figs 1-4. [Takinoura, Ani-jima Island, Ogasawara
Islands, 5 m]
Neostylodactylus sibogae (De Man, 1918a)
= Stylodactylus Sibogae De Man, 1918a: 159. [Siboga Expedition stn 95, 543'.5N 11940'E, Sulu Sea,
522 m]
Parastylodactylus Figueira, 1971
= Parastylodactylus Figueira, 1971 (type species Stylodactylus bimaxillaris Spence Bate, 1888, by origi-
Parastylodactylus bimaxillaris (Spence Bate, 1888)
= Stylodactylus bimaxillaris Spence Bate, 1888: 855; Plate 138, fg. 3. [Challenger stn 219, 154'0"S
14639'40"E, of the Admiralty Islands, 150 fms]
Parastylodactylus hayashii (Komai, 1997)
= Neostylodactylus hayashii Komai, 1997a: 125 (in part); Figs 1-3. [R/V Tansei-maru, KT95-5 cruise, stn
TB18-2, Okinoyama, of Sunosaki, Boso Peninsula, 3459'N 13939'E, 105-113 m]
Parastylodactylus longidactylus Cleva, 1990: 125; Figs 12d, 15a-b, 16a-d. [Philippines, 1355.2'N 12030.5'E,
320-318 m]
Parastylodactylus moluccensis Cleva, 1997: 398; Fig. 3. [Indonsie, 0517'06''S 13251'19''E, 315-348 m]
Parastylodactylus richeri Cleva, 1990: 127; Figs 15c, 16e-h. [Nouvelle-Caldonie, 2345'S 16712'E, 380
m]
Parastylodactylus semblatae Cleva, 1990: 122; Fig. 12c, 14, 18c. [Nouvelle-Caldonie, 2216.5'S 16716.5'E,
475-500 m]
Parastylodactylus tranterae Cleva, 1990: 119; Figs 11b, 12b, 13. [Nouvelle-Caldonie, 2211.3'S 16715.0'E,
495-550 m]
Stylodactyloides Cleva, 1990
= Stylodactyloides Cleva, 1990 (type species Stylodactyloides crosnieri Cleva, 1990, by monotypy, gen-
der masculine)
Stylodactyloides crosnieri Cleva, 1990: 129; Figs 17, 18d-e. [Iles Chesterfeld, 1953.20'S 15839.50'E, 370-
400 m]
Stylodactylus A. Milne-Edwards, 1881b
= Stylodactylus A. Milne-Edwards, 1881b (type species Stylodactylus serratus A. Milne-Edwards,
1881b, by monotypy, gender masculine; name placed on the Ofcial List of Generic Names in Zool-
Stylodactylus brevidactylus Cleva, 1990: 106; Fig. 8a-g. [Philippines, 1340.5'N 12053.7'E, 230-204 m]
Stylodactylus brucei Cleva, 1994: 54; Fig. 1A, C-F. [south-west Pacifc, Wallis Island, 1337'S 17956'E, 820-
840 m]
Stylodactylus discissipes Spence Bate, 1888: 851; Plate 138, fg. 1. [Challenger stn 171, 2833'S 17750'W,
north of the Kermadec Islands, 600 fms]
= Stylodactylus orientalis Spence Bate, 1888: 854; Plate 138, fg. 2. [Challenger stn 171, 2833'S 17750'W,
north of the Kermadec Islands, 600 fms]
Stylodactylus gracilis Cleva, 2008b: 68. [nomen novum for Stylodactylus gracilis Cleva, 2008a]
= Stylodactylus gracilis Cleva, 2008a: 32; Figs 1, 3A. [Philippine Islands, Panglao Island, 849.3'N
1231.9'E, 2149-2217 m; not validly published name according to Art. 16.4.2]
Stylodactylus kauaiensis Figueira, 1971: 3. [Kauai Island, Hawaii, 230-253 fms]
Stylodactylus laurentae Cleva, 1990: 96; Figs 6, 19. [Nouvelle-Caldonie, 2455.48'S 16821.29'E, 500 m]
Stylodactylus libratus Chace, 1983b: 12; Fig. 5. [Indonesia, Selat Butung, Celebes, 529'06"S 12236'06"E,
377 m]
Stylodactylus licinus Chace, 1983b: 14; Fig. 6. [Philippines, Palawan Passage, 1057'45"N 11838'15"E, 686 m]
Stylodactylus macropus Chace, 1983b: 16; Fig. 7. [Philippines, north of Samar, 1244'42"N 12459'50"E,
700 m]
Stylodactylus major Hayashi & Miyake, 1968a: 590; Figs 2-3. [east China Sea, 3053.2'N 12726.0'E
3044.0'N 12729.0'E, 122-124 m]
Stylodactylus multidentatus multidentatus Kubo, 1942b (Fig. 25)
= Stylodactylus multidentatus Kubo, 1942b: 34; Figs 4-5. [Kumanonada, of Mie Prefecture, about 300 m]
= Stylodactylus brevidactylus Cleva, 1990: 106; Fig. 8a-g. [Philippines, 1340.5'N 12053.7'E, 230-204 m]
Fig. 25. Stylodactylus multidentatus multidentatus Kubo, 1942. Photo by Tin-Yam Chan.
Stylodactylus multidentatus robustus Cleva, 1990: 105; Fig. 8n-s. [Madagascar, 2220.5'S 4306.1'E, 400 m]
Stylodactylus profundus Cleva, 1990: 85; Fig. 2. [Nouvelle-Caldonie, 2419'S 16708'E, 1530-1480 m]
Stylodactylus pubescens Burukovsky, 1990: 198; Fig. 3B. [2509'S 9618'W, 545-800 m]
Stylodactylus rectirostris A. Milne-Edwards, 1883: Plate 35. [of Dominica, 1518'12''N 6126'32''W, 542
fms]
Stylodactylus serratus A. Milne-Edwards, 1881b: 11. [prs de la Dominique, 524 brasses]
Stylodactylus stebbingi Hayashi & Miyake, 1968a: 595; Fig. 4. [Of Bufalo River, NW W, 19 miles, East
London, Cape Colony, 300 fms]
Stylodactylus tokarensis Zarenkov, 1968a: 58; Figs 2-3. [Vitjaz stn 3768, Tokara Strait, East China Sea, 820
m]
Superfamily CAMPYLONOTOIDEA Sollaud, 1913
Family BATHYPALAEMONELLIDAE De Saint Laurent, 1985
Bathypalaemonella Balss, 1914a
= Bathypalaemonella Balss, 1914a (type species Bathypalaemonella zimmeri Balss, 1914a, by monotypy,
gender feminine)
Bathypalaemonella adenensis Cleva, 2001: 770; Fig. 6. [Golfe dAden, 1156.5'N 4334.5'E, 1600-1400 m]
Bathypalaemonella delsolari Wicksten & Mndez, 1983: 225; Figs 1-4. [SW of Lobos de Tierra, Peru, 631'S
8101'W, 712-714 m]
Bathypalaemonella hayashii Komai, 1995a: 40; Figs 1-4. [southeast of Amami-oshima Island, 2802.5'N
12934.5'E / 2802.8'N 12934.9'E, 815-789 m]
Bathypalaemonella humilis Bruce, 1966b: 277; Figs 1-3. [1555.7'N 10928.5'E to 1557.9'N 10931.5'E, 172-
142 fms]
Bathypalaemonella pandaloides (Rathbun, 1906)
= Palaemon pandaloides Rathbun, 1906: 924; Fig. 73; Plate 22, fg. 4. [vicinity of Kauai Island, 528 fms]
Bathypalaemonella serratipalma Pequegnat, 1970: 77; Figs 4.5-4.6. [Southwestern Gulf of Mexico, 2144'N
9646'W, 1774 m] (Fig. 26)
Bathypalaemonella texana Pequegnat, 1970: 81; Figs 4.7-4.8. [Northwestern Gulf of Mexico, 2701.6'N
9442'W, 1463 m]
Bathypalaemonella zimmeri Balss, 1914a: 598. [618'N 4932'E, 1079 m]
Fig. 26. Bathypalaemonella serratipalma Pequegnat, 1970. Photo Gilliss Expedition stn G-1111.
Bathypalaemonetes Cleva, 2001
= Bathypalaemonetes Cleva, 2001 (type species Bathypalaemonella brevirostris Bruce, 1986a, by original
Bathypalaemonetes brevirostris (Bruce, 1986a)
= Bathypalaemonella brevirostris Bruce, 1986a: 252; Figs 1-5. [1352.1'S 12300.4'E, 306-308 m]
Bathypalaemonetes chani Cleva, 2004: 508; Figs 4-5, 6h. [220.98'N 1206.73'E, 690-700 m]
Bathypalaemonetes pilosipes (Bruce, 1986a)
= Bathypalaemonella pilosipes Bruce, 1986a: 257; Figs 6-10. [1333.8'S 12253.4'E, 390-394 m]
Family CAMPYLONOTIDAE Sollaud, 1913
Campylonotus Spence Bate, 1888
= Campylonotus Spence Bate, 1888 (type species Campylonotus semistriatus Spence Bate, 1888, desig-
nated by Holthuis, 1955b, gender masculine; name placed on the Ofcial List of Generic Names in
= Anchistiella A. Milne-Edwards, 1891 (type species Anchistiella Hyadesi A. Milne-Edwards, 1891
(junior subjective synonym of Campylonotus vagans Spence Bate, 1888), designated by Holthuis,
1955b, gender feminine)
Campylonotus arntzianus Thatje, 2003: 243; Figs 2-4. [Saunders Island, South Sandwich Islands, Scotia
Sea, Antarctica, 5740.31'S 2627.81'W, 475-589 m]
Campylonotus capensis Spence Bate, 1888: 773; Plate 128, fg. 3. [Challenger stns 145, 4643'S 384'30''E, of
Marion Island, 140 fms; 122, 95'S 3450'W, of Pernambuco, 350 ms]
Campylonotus rathbunae Schmit, 1926a
= Campylonotus rathbun Schmit, 1926a: 373; Plate 67, fgs 1-5. [South of Eucla, Great Australian
Bight, 12928'E, 250-450 fms]
Fig. 27. Campylonotus vagans Spence Bate, 1888. Photo by Karen Neely.
Campylonotus semistriatus Spence Bate, 1888: 768; Plate 128, fgs 1-2. [Challenger stns 309, 5056'S 7415'W,
Puerto Bueno, Patagonia, 40 fms; 305A, 4748'30''S 7447'0''W, Messier Channel, 125 fms; 306A,
4827'S 7430'W, Messier Channel, 345 fms; 307, 4924'30''S 7423'30''W, of Port Grappler, 140 fms;
308, 508'30''S 7441'0''W, of Tom Bay, 175 fms; 310, 5127'30''S 743'0''W, Sarmiento Channel, 400
fms; 311, 5245'30''S 7346'0''W, of Port Churruca, 245 fms]
Campylonotus vagans Spence Bate, 1888: 775; Plate 122, fg. 3. [Challenger stn 308, 508'30''S 7441'0''W, of
Tom Bay, Patagonia, 175 fms] (Fig. 27)
= Anchistiella Hyadesi A. Milne-Edwards, 1891: 38; Plate 4, fgs 1-1n. [ lentre est du canal de Fran-
klin, entre Herschel et Wollaston, 51 m; dans la baie Orange, 26-28 m; dans la baie du Naturaliste,
35 m; dans le dtroit de Magellan, 18 m; lle de Wollaston, 95 m]
= Anchistiella Hahni A. Milne-Edwards, 1891: 41; Plate 4, fgs 2-2d. [ lle Wollaston dans la baie
Greton, Lajaifwaya, 30 m; aux Murray narrows, 200 m; 10 milles au sud-est de la baie Seogat,
816 m]
= Anchistiella Seneuilli A. Milne-Edwards, 1891: 42; Plate 3, fgs 2-2d. [dans le dtroit de Magellan,
en face de Punta Arenas, 143 m]
Superfamily PALAEMONOIDEA Rafnesque, 1815
Family ANCHISTIOIDIDAE Borradaile, 1915b
Anchistioides Paulson, 1875
= Anchistioides Paulson, 1875 (type species Anchistioides compressus Paulson, 1875, by monotypy,
gender masculine; name placed on the Ofcial List of Generic Names in Zoology in Opinion 470 in
1957)
= Palaemonopsis Borradaile, 1900 (type species Palaemonopsis willeyi Borradaile, 1900, by monotypy,
gender feminine)
= Amphipalaemon Nobili, 1901a (nomen novum for Palaemonopsis Borradaile, 1900, gender mascu-
line)
Anchistioides antiguensis (Schmit, 1924b) (Fig. 28)
= Periclimenes antiguensis Schmit, 1924b: 84. [English Harbour, Antigua, electric light]
= Periclimenes barbadensis Schmit, 1924b: Plates 3-4. [English Harbour, Antigua, electric light]
Anchistioides compressus Paulson, 1875
= Anch.[istioides] compressus Paulson, 1875: 115; Plate 19, fgs 5-5n. [Red Sea]
= Amphipalaemon Seurati Nobili, 1906b: 259. [Tearia, Tuamotu Islands according to Bruce, 1967a]
Fig. 28. Anchistioides antiguensis (Schmit, 1924). Photo by Arthur Anker.
Anchistioides willeyi (Borradaile, 1900)
= Palaemonopsis willeyi Borradaile, 1900: 410; Plate 36, fgs 7-7b; Plate 37, fgs 7c-e. [Ralun, New
Britain]
= Amphipalaemon cooperi Borradaile, 1915b: 209. [S. Nilandu Atoll, Maldive Islands]
= Amphipalaemon gardineri Borradaile, 1915b: 209. [N. Mal Atoll, Maldive Islands]
= Amphipalaemon australiensis Balss, 1921a: 11; Figs 3-6. [Cape Jaubert, 45 meilen WSW, 66 Fuss Tiefe]
Family DESMOCARIDIDAE Borradaile, 1915b
Desmocaris Sollaud, 1911a
= Desmocaris Sollaud, 1911a (type species Palaemonetes trispinosus Aurivillius, 1898a, by monotypy,
1957)
Desmocaris trispinosa (Aurivillius, 1898a) (Fig. 29)
= Palaemonetes trispinosus Aurivillius, 1898a: 29; Plate 4, fgs 1-2. [Kamerun, Kita, in einem Bache]
Desmocaris bislineata Powell, 1977: 667; Fig. 5. [River Ethiope at Adagbrassa-Amukpe, 551'N 545.5'E]
Family EURYRHYNCHIDAE Holthuis, 1950a
Euryrhynchina Powell, 1976
= Euryrhynchina Powell, 1976 (type species Euryrhynchina edingtonae Powell, 1976, by monotypy,
gender feminine)
Euryrhynchina edingtonae Powell, 1976: 895; Figs 5-7. [swamp forest along Ogba River, 1 km south of
Ogba village and approx. 8 km southwest of Benin City centre, 616.5'N 535'E]
Euryrhynchoides Powell, 1976
= Euryrhynchoides Powell, 1976 (type species Euryrhynchoides holthuisi Powell, 1976, by monotypy,
gender masculine)
Euryrhynchoides holthuisi Powell, 1976: 886; Figs 1-4. [Sierra Leone: River Taja (= River Jong) at Njala,
806'N 1204'W]
Euryrhynchus Miers, 1877
= Euryrhynchus Miers, 1877 (type species Euryrhynchus wrzesniowskii Miers, 1877, by monotypy, gen-
der masculine; placed on the Ofcial List of Generic Names in Zoology in Opinion 518 in 1958)
= Euryrhynchella Balss, 1955 (nomen novum for Euryrhynchus Miers, 1877, gender feminine)
Euryrhyncus amazoniensis Tiefenbacer, 1978: 183; Fig. 2a-b. [A15-1, Rio Tupana, am 2. Vermessungs-
punkt]
Euryrhynchus burchelli Calman, 1907: 297; Figs 2-8. [Par]
Euryrhynchus pemoni Pereira, 1985: 619; Fig. 5. [Creek in La Gran Sabana road, between Sn. Rafael Town
and Kama Fall, Venezuela, 615'N 601'W]
Euryrhynchus tomasi De Grave, 2007a: 194; Figs 1-7. [Crique Timothy, near the N2 road from Rgina to St
Georges de lOyapock, 20 km NW of St Georges de lOyapock, French Guyana, approx. 400'N
5152'W] (Fig. 30)
Euryrhynchus wrzesniowskii Miers, 1877: 662; Plate 67, fgs 2-2b. [Cayenne]
Family GNATHOPHYLLIDAE Dana, 1852a
Gnathophylleptum dUdekem dAcoz, 2001
= Gnathophylleptum dUdekem dAcoz, 2001 (type species Gnathophylleptum tellei dUdekem dAcoz,
2001, by monotypy, gender neuter)
Gnathophylleptum tellei dUdekem dAcoz, 2001: 114; Figs 1-7. [Canary Islands, Gran Canaria, Sardina del
Norte, Caulerpa racemosa meadow, 15 m]
Fig. 29. Desmocaris trispinosa (Aurivillius, 1898). Photo by Chris Lukhaup.
Fig. 30. Euryrhynchus tomasi De Grave, 2007. Photo by J. Tomas & O. Helker.
Gnathophylloides Schmit, 1933
= Gnathophylloides Schmit, 1933 (type species Gnathophylloides mineri Schmit, 1933, by monotypy,
gender masculine)
Gnathophylloides mineri Schmit, 1933: 7; Fig. 3. [Ballena Point, Ensenada, Puerto Rico]
Gnathophylloides robustus Bruce, 1973a: 17; Figs 1-7. [of Point Moore, Geraldton, Western Australia,
2847.0'S 11434.2'E, 3 m]
Gnathophyllum Latreille, 1819
= Gnatophyllum Latreille, 1819 (type species A.[lpheus] elegans Risso, 1816, designated by H. Milne
Edwards, 1837 [in H. Milne Edwards, 1836-1844], gender neuter; name emended under the plenary
power of the ICZN to Gnathophyllum and placed in that emended spelling on the Ofcial List of
= Drimo Risso, 1827 (type species A.[lpheus] elegans Risso, 1816, gender masculine; name placed on
the Ofcial Index of Rejected and Invalid Generic Names in Zoology in Opinion 470 in 1957)
Gnathophyllum americanum Gurin-Mneville, 1855 [in Gurin-Mneville, 1855-1856]: xx; Plate 2, fg. 14.
[Cuba]
= Gnathophyllum fasciolatum Stimpson, 1860a: 28. [Rose Bay, Port Jackson, NSW; neotype designation
by Ahyong, 2003]
= Gnathophyllum zebra Richters, 1880: 161; Plate 17, Figs 18-20. [Fouquets]
= Gnathophyllum pallidum Ortmann, 1890: 537. [Tahiti]
= Gnathophyllum tridens Nobili, 1906b: 259. [Rikitea]
= Gnathophyllum minuscularium Armstrong, 1940: 9; Fig. 4C-K. [in the Reach, St. George Island, Ber-
muda]
Gnathophyllum ascensione Manning & Chace, 1990: 11; Figs 5-6. [Northeast Bay, Ascension Island]
Gnathophyllum circellum Manning, 1963: 54; Figs 3-4. [300 yards southwest of Alligator Reef Light, Mon-
roe Co., Florida, rock ledge at 15-20 feet]
Gnathophyllum elegans (Risso, 1816)
= Peneus quadricolor Rafnesque, 1814: 22. [Type locality not indicated; suppressed under the plenary
power for the purposes of the Principle of Priority but not for those of the Principle of Homonymy
in Opinion 522 in 1958]
= A.[lpheus] elegans Risso, 1816: 92; Plate 2, fg. 4. [environs de Nice, dans les profondeurs rocail-
leuses]
= Gnathophyllum elegans var. brevirostris Gourret, 1887a: 1034. [lestaque jusqu la Corbire; prs de
Somaty; Mourepiano; Roucas blanc]
Gnathophyllum modestum Hay, 1917: 72. [about 20 miles S.W. of Beaufort, N.C., in about 15 fms]
Gnathophyllum panamense Faxon, 1893: 198. [Panama] (Fig. 31)
Gnathophyllum precipuum Titgen, 1989: 203; Figs 1-3. [Oahu, Mahaka, 12 m]
Gnathophyllum splendens Chace & Fuller, 1971: 493; Figs 1-5. [Puerto Yabucoa, one-half mile east of Playa
de Guayans, Municipio de Yabucoa, Puerto Rico]
Gnathophyllum taylori Ahyong, 2003: 237; Figs 1-2. [S side of Long Bay, Malabar, New South Wales,
3358'S 15115'E, 4 m, under Centrostephanus rogersi]
Levicaris Bruce, 1973a
= Levicaris Bruce, 1973a (type species Coralliocaris mammilatus Edmondson, 1931, by monotypy, gen-
der feminine)
Levicaris mammillata (Edmondson, 1931)
= Coralliocaris mammillata Edmondson, 1931: 5; Fig. 2; Plate 1. [Waikiki Reef, Oahu, associated with
echinoid Heterocentrotus mamillatus]
Pycnocaris Bruce, 1972a
= Pycnocaris Bruce, 1972a (type species Pycnocaris chagoae Bruce, 1972a, by monotypy, gender feminine)
Pycnocaris chagoae Bruce, 1972a: 54; Figs 1-7. [East Point, Diego Garcia, Chagos Archipelago] (Fig. 32)
Fig. 31. Gnathophyllum panamense Faxon, 1893. Photo by Arthur Anker.
Fig. 32. Pycnocaris chagoae Bruce, 1972. Photo by Tin-Yam Chan.
Family HYMENOCERIDAE Ortmann, 1890
Hymenocera Latreille, 1819
= Hymenocera Latreille, 1819 (type species Hymenocera picta Dana, 1852b, designated under the ple-
nary power of the ICZN, gender feminine; name placed on the Ofcial List of Generic Names in
= Nematophyllum Bleeker, 1856 (type species Hymenocera picta Dana, 1852b, designated by Holthuis,
1955b, gender neuter; invalid junior hononym of Nematophyllum H. Milne Edwards & Haime,
1850 (Coelenterata); name placed on the Ofcial List of Rejected and Invalid Names in Zoology in
Hymenocera picta Dana, 1852b: 593 (1852); Plate 39, fgs 3a-c (1855). [coral reefs of Raraka, Paumotu Is-
lands] (Fig. 33)
= H.[ymenocera] elegans Heller, 1861: 25. [Zu Tor zwischen Korallen]
= H.[ymenocera] elegans Heller, 1862c: 264; Plate 3, fgs 9-14. [Tor]
= Hymenocera latreillii Sharp, 1893: 119 [Agalega, Seychelles]
Phyllognathia Borradaile, 1915b
= Phyllognathia Borradaile, 1915b (type species Hymenocera (?) ceratophthalma Balss, 1913b, by mono-
typy, gender feminine; name placed on the Ofcial List of Generic Names in Zoology in Opinion
470 in 1957)
Phyllognathia ceratophthalma (Balss, 1913b) (Fig. 34)
= Hymenocera (?) ceratophthalma Balss, 1913b: 236. [Satsuma]
Phyllognathia simplex Fujino, 1973a: 90; Figs 1-3. [near Shuragane, Sagami Bay, central Japan, 40-50 m deep]
Fig. 33. Hymenocera picta Dana, 1852. Photo by Arthur Anker.
Family KAKADUCARIDIDAE Bruce, 1993a
Calathaemon Bruce & Short, 1993
= Calathaemon Bruce & Short, 1993 (type species Palaemonetes holthuisi Strenth, 1976, by monotypy,
gender masculine)
Calathaemon holthuisi (Strenth, 1976)
= Palaemonetes holthuisi Strenth, 1976: 3, Fig. 1. [subterranean waters of Ezells Cave in the city of San
Marcos, Hays County, Texas]
Kakaducaris Bruce, 1993a
= Kakaducaris Bruce, 1993a (type species Kakaducaris glabra Bruce, 1993a, by monotypy, gender fem-
inine)
Kakaducaris glabra Bruce, 1993a: 28; Figs 1-11, 12A. [Lightening Dreaming Creek, Nourlangi, Kakadu
National Park, Arnhem Land, Northern Territory, 1255.3'S 13255.8'E, 90 m alt.] (Fig. 35)
Leptopalaemon Bruce & Short, 1993
= Leptopalaemon Bruce & Short, 1993 (type species Leptopalaemon gagadjui Bruce & Short, 1993, by
Leptopalaemon gagadjui Bruce & Short, 1993: 75; Figs 1-6. [Barramundi Creek, Kakadu National Park,
Northern Territory, 1320'S 13227'E]

Family PALAEMONIDAE Rafnesque, 1815
Subfamily PALAEMONINAE Rafnesque, 1815
Arachnochium Wowor & Ng, 2010a
= Arachnochium Wowor & Ng, 2010a (type species Palaemon mirabilis Kemp, 1917c, by original des-
ignation, gender neuter)
Arachnochium kulsiense (Jayachandran, Lal Mohan & Raji, 2007)
= Macrobrachium kulsiense Jayachandran, Lal Mohan & Raji, 2007: 43; Figs 1-3. [Kulsi River]
Arachnochium mirabilis (Kemp, 1917c)
= Palaemon mirabilis Kemp, 1917c: 227; Plate 10. [Pazudaung and Dala Creeks, Rangoon]
Fig. 34. Phyllognathia ceratophthalma (Balss, 1913). Photo by Tin-Yam Chan.
Brachycarpus Spence Bate, 1888
= Brachycarpus Spence Bate, 1888 (type species Brachycarpus savignyi Spence Bate, 1888 (junior sub-
jective synonym of Brachycarpus biunguiculatus Lucas, 1846), by original designation, gender mas-
culine; name placed on the Ofcial List of Generic Names in Zoology in Opinion 470 in 1957)
= Retrocaris Ortmann, 1893 (type species Retrocaris spinosa (junior subjective synonym of Palmon
biunguiculatus Lucas, 1846), designated by Holthuis, 1955b, gender feminine)
= Calmania Nobili, 1907 (type species Palmon biunguiculatus Lucas, 1846, by original designation
and monotypy, gender feminine; invalid junior homonym of Calmania Laurie, 1906 (Crustacea
Brachyura))
Brachycarpus biunguiculatus (Lucas, 1846)
= Palmon biunguiculatus Lucas, 1846: 45; Plate 4, fgs 4-4a. [Oran et Bne]
= Brachycarpus savignyi Spence Bate, 1888: 795; Plate 124, fg. 4. [Bermuda]
= Brachycarpus neapolitanus Cano, 1890: 38; Plate 4, fg. 1. [Golfo di Napoli]
= Retrocaris spinosa Ortmann, 1893: 84; Plate 6, fg. 2. [Plankton-Expedition der Humboldt-Stifung,
Sargasso-See JN 62 (33.2N 63.8W, 0-400 m) and Pl. 34 (32.1N 63.4W, 200 m); Sdl. Aequatorial-
strom JN 249 (5.6S 44.0W, 0 m)]
= Brachycarpus advena Nobili, 1905b: 395. [Mer Rouge]
= Palmonella rathbunensis Borradaile, 1917: 358. [Hawaiian Is.]
Brachycarpus crosnieri Bruce, 1998a: 158; Figs 1-4. [New Caledonia, Loyalty Islands, Uvea, Passe de la
Meurthe, 6-10 m]
Brachycarpus holthuisi Fausto Filho, 1966: 123; Figs 1-11. [Acara, Cear]
Coutierella Sollaud, 1914
= Coutierella Sollaud, 1914 (type species Coutierella tonkinensis Sollaud, 1914, by monotypy, gender
feminine)
Coutierella tonkinensis guangdongensis (Liu, Liang & Yan, 1990a)
= Palaemonetes tonkinensis guangdongensis Liu, Liang & Yan, 1990a: 250; Fig. 41. [Guangzhou]
Fig. 35. Kakaducaris glabra Bruce, 1993. Photo by A.J. Bruce.
Coutierella tonkinensis tonkinensis Sollaud, 1914
= Coutierella tonkinensis Sollaud, 1914: 318; Figs 2A, 3B, 4. [delta du feuve Rouge, Tonkin]
Creaseria Holthuis, 1950a
= Creaseria Holthuis, 1950a (type species Palmon morleyi Creaser, 1936, by original designation,
gender feminine)
Creaseria morleyi (Creaser, 1936)
= Palmon morleyi Creaser, 1936: 126; Figs 25-30. [San Isidro Cave, Salar Colony, Merida, Yucatan]
Cryphiops Dana, 1852a
= Cryphiops Dana, 1852a (type species Cryphiops spinuloso-manus Dana, 1852a (junior subjective syn-
onym of Cancer caementarius Molina, 1782), by monotypy, gender masculine)
= Bithynis Philippi, 1860 (type species Bithynis longimana Philippi, 1860 (junior subjective synonym
of Cancer caementarius Molina, 1782), by monotypy, gender feminine)
= Bithynops Holthuis, 1973b (type species Bythinops luscus Holthuis, 1973b, by original designation
Cryphiops (Bithynops) brasiliensis (Gomes Corra, 1973)
= Cryphiops brasiliensis Gomes Corra, 1973: 169; Figs 1-21. [riacho da Granja do Ip, em Braslia,
distrito Federal]
Cryphiops (Bithynops) luscus (Holthuis, 1973b)
= Bythinops luscus Holthuis, 1973b: 136; Figs 1-2. [Grutas de lArco, near San Raphael de lArco, La
Trinitaria, about 1610'N 9201'W, Chiapas State, Mexico, altitude 1470 m]
Cryphiops (Bithynops) perspicax (Holthuis, 1977b)
= Bythinops perspicax Holthuis, 1977b: 182; Figs 3-4. [Cenote la Cueva, Lagunas de Montebello, La
Trinitaria, Chiapas, altitude 1480 m]
Cryphiops (Bithynops) sbordonii Baldari, Meja-Ortiz & Lpez-Meja, 2010: 48; Figs 2-4. [Cueva Cham-
burro, Las Margaritas, Chiapas, Mexico, 1625'57''N 9156'40''W]
Cryphiops (Bithynops) villalobosi Villalobos Hiriart, Nates Rodriguez & Cant Diaz Barriga, 1989: 166;
Figs 1-5, 7b-d, 8c. [arroyo La Laja, km. 140 carretera Palenque-Ocosingo, a 5 km al NO del poblado
de Ocosingo. Mpio. de Ocosingo, Chiapas]
Cryphiops (Cryphiops) caementarius (Molina, 1782) (Fig. 36)
= Cancer cmentarius Molina, 1782: 208. [Chili]
= P.[almon] Gaudichaudii H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 400. [Chili]
= Cryphiops spinuloso-manus Dana, 1852a: 26. [in fuminibus Chilensibus mari remotis]
= Bithynis longimana Philippi, 1860: 164. [im Flusse la Ligua, Chile]
= Macrobrachium africanum Spence Bate, 1868a: 366; Plate 31, fg. 3. [Tambo river; according to Sem-
per, 1868 on the west coast of South America, near Islay in Peru]
Exopalaemon Holthuis, 1950a
= Exopalaemon Holthuis, 1950a (type species P.[alemon] styliferus H. Milne Edwards, 1840, by origi-
Exopalaemon annandalei (Kemp, 1917c)
= Leander annandalei Kemp, 1917c: 211; Fig. 1-4. [China, Whangpoo River, between Shanghai and
Woosung, 5.5-7.5 m]
= Leander annandalei var. stylirostris Yu, 1930a: 460; Fig. 4C-P. [Tangku]
Exopalaemon carinicauda (Holthuis, 1950a)
= Palaemon (Exopalaemon) carinicauda Holthuis, 1950a: 48; Fig. 9. [nomen novum for Leander longiro-
stris var. carinatus Ortmann, 1890]
= Leander longirostris var. carinatus Ortmann, 1890: 521. [China]
Exopalaemon guangdongensis Guo, Wang & Zhang, 2005: 840; Figs 1-2. [Zhujiang river, near Tangjiawan,
Zhuhai City, circa 2236'N 11336'E]
Exopalaemon hainanensis Liang, 2000: 278; Figs 11-19. [fshery market, Haikou, Hainan Island]
Exopalaemon mani (Sollaud, 1914)
= Leander Mani Sollaud, 1914: 315; Figs 1, 2B. [delta du feuve Rouge, Tonkin]
Exopalaemon modestus (Heller, 1862a)
= L.[eander] modestus Heller, 1862a: 527. [Schanghai]
= Leander modestus Heller, 1865: 111; Plate 10, fg. 6. [Shanghai]
= Leander czerniavskyi Branikov, 1907: 176. [Amurskiy Liman]
= Leander modestus var. sibirica Czerniavsky, 1878; 24. [nomen nudum]
= Leander macrogenitus Yu, 1930b: 559; Figs 1A-C. [Hangchow]
= Leander czerniavsky lacustris Buldovsky, 1933: 50; Plate 2, fgs 21, 23. [Lake Khanka, Soviet Far East]
Exopalaemon orientis (Holthuis, 1950a)
= Palaemon (Exopalaemon) orientis Holthuis, 1950a: 49. [nomen novum for Leander longirostris var.
japonicus Ortmann, 1890]
= Leander longirostris var. japonicus Ortmann, 1890: 519; Plate 37, fgs 14, 14z. [Japan, Tokiobai]
Exopalaemon styliferus (H. Milne Edwards, 1840 [in H. Milne Edwards, 1834-1840])
= P.[alemon] styliferus H. Milne Edwards, 1840 [in H. Milne Edwards, 1834-1840]: 638. [nomen no-
vum for P.[alemon] longirostris H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 394 nec
P.[alemon] longirostris H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 392]
= P.[alemon] longirostris H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 394. [lembouchure
du Gange]
Exopalaemon vietnamicus Nguyn, 1992: 25; Figs 4, 12 (part). [Can Gi, 15 km west of Vu T]
Exopalaemon xinjiangensis Liang, 2000: 277; Figs 1-10. [Talimu River, Xinjiang, northwest China]
Leander Desmarest, 1849
= Leander Desmarest, 1849 (type species Leander erraticus Desmarest, 1849 (junior subjective syno-
nym of P.[alaemon] tenuicornis Say, 1818), by original designation and monotypy, gender masculine;
= Cryptoleander Gurney & Lebour, 1941 (type species P.[alaemon] tenuicornis Say, 1818, by monotypy,
gender masculine)
Leander kempi Holthuis, 1950a: 31; Fig. 3. [Siboga stn 121, Menado anchorage, shore exploration, 55 m;
Snellius Expedition, Beo, Talaud Islands]
Fig. 36. Cryphiops (Cryphiops) caementarius (Molina, 1782). Photo by Arthur Anker.
Leander manningi Bruce, 2002a: 71; Figs 1-4. [BP Oil Refnery Jety, Kwinana, Cockburn Sound, Western
Australia]
Leander paulensis Ortmann, 1897: 192; Plate 1, fg. 14. [canal entre o continente e a ilha de So Sebastio,
Estado de So Paulo]
Leander plumosus Bruce, 1994a: 39; Figs 1-5, 6A-B; Plate 2. [Ari Atoll, Maldive Islands]
Leander tenuicornis (Say, 1818)
= P.[almon] tenuicornis Say, 1818: 249. [Banks of Newfoundland]
= Astacus locusta Fabricius, 1781: 513; nec Astacus locusta Pennant, 1777. [Type locality not indicated]
? = Penaeus punctatissimus Bosc, 1802: 109; Plate 14, fg. 3. [dans sa traverse dEurope en Amrique,
sur les fucus nageans]
? = Penaeus adspersus Tilesius, 1819: 4; Plate 21, fg. 1. [suppressed under the Plenary Power for the
Purpose of the Principle of Priority but not for those of the Principle of Homonymy in Opinion 564
in 1959]
= P.[alemon] natator H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 393. [dans locan
Indien, sur du fucus nageans]
= Palaemon latirostris De Haan, 1844 [in De Haan, 1833-1850]: Plate 45, fg. 12 (1844); 170, Plate P
(1849). [Japan; lectotype designation by Yamaguchi & Baba, 1993]
= Leander erraticus Desmarest, 1849: 92; unnumbered fgure. [Ocan Atlantique, cent ou cent cin-
quante lieues des ctes de la Guadeloupe]
= P.[alaemon] torensis Paulson, 1875: 116; Plate 17, fg. 3. [Red Sea]
= P.[alaemon] torensis var. A Paulson, 1875: 116; Plate 17, fg. 4. [Red Sea]
= P.[alaemon] torensis var. B Paulson, 1875: 116; Plate 17, fg. 5. [Red Sea]
? = Palaemon mauritii Spamer & Bogan, 1992: 163. [nomen nudum]
? = Palaemon mexicanus Spamer & Bogan, 1992: 163. [nomen nudum]
Leandrites Holthuis, 1950a
= Leandrites Holthuis, 1950a (type species Leander celebensis De Man, 1881 by original designation,
gender masculine)
Leandrites celebensis (De Man, 1881)
= Leander celebensis De Man, 1881: 141. [on the road of Macassar (Celebes)]
= Palaemonetes hornelli Kemp, 1925: 318; Figs 14-15. [Silavathuri Lagoon, Tuticorin, S. India]
= Leander wieneckii Holthuis, 1950a: 36. [nomen nudum, manuscript name for Leander celebensis De
Man, 1881]
Leandrites deschampsi (Nobili, 1903c)
= Leander Deschampsi Nobili, 1903c: 8. [Singapore]
Leandrites indicus Holthuis, 1950a: 37; Fig. 5. [Of Makassar]
Leandrites stenopus Holthuis, 1950a: 40; Fig. 6. [Madoera Strait, 725'S 11316'E, 56 m]
Leptocarpus Holthuis, 1950a
= Leptocarpus Holthuis, 1950a (type species Leander fuminicola Kemp, 1917c, by original designation,
gender masculine)
Leptocarpus fuminicola (Kemp, 1917c)
= Leander fuminicola Kemp, 1917c: 223; Plate 9, fg. 2. [River Ganges, Mirzapur, United Provinces]
Leptocarpus kempi Jayachandran, 1992: 129; Fig. 2. [Cochin backwaters at Panangad]
Leptocarpus potamiscus (Kemp, 1917c)
= Leander potamiscus Kemp, 1917c: 225; Fig. 7. [Patani River, below the town of Patani, Siamese Ma-
lay States]
Macrobrachium Spence Bate, 1868a
= Macrobrachium Spence Bate, 1868a (type species Macrobrachium americanum Spence Bate, 1868a,
designated by Fowler, 1912, gender neuter; name placed on the Ofcial List of Generic Names in
= Eupalaemon Ortmann, 1891 (type species Palaemon (s.s.) dolichodactylus Hilgendorf, 1879 (junior
subjective synonym of Palaemon scabriculus Heller, 1862a), designated by Holthuis, 1955b, gender
masculine)
= Parapalaemon Ortmann, 1891 (type species Palaemon (s.s.) dolichodactylus Hilgendorf, 1879 (junior
subjective synonym of Palaemon scabriculus Heller, 1862a, designated by Holthuis, 1955b, gender
masculine)
= Macroterocheir Stebbing, 1908 (type species Palaemon (s.s.) lepidactylus Hilgendorf, 1879, by mono-
typy, gender masculine)
Macrobrachium acanthochirus Villalobos F., 1967: 168; Plates 1-2. [Ro Valdefores, Valdefores de Tona-
meca, Pochutla, Estado de Oaxaca, Mexico]
Macrobrachium acanthurus (Wiegmann, 1836)
= P.[alaemon] acanthurus Wiegmann, 1836: 150. [Kste Brasiliens]
= P.[alemon] forceps H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 397. [Rio-Janeiro]
= Pal.[mon] Swainsonii White, 1847a: 78. [nomen nudum]
= Palmon mexicanus de Saussure, 1857a: 504. [Cuba et Mexique]
= Macrobrachium longidigitum Spence Bate, 1868a: 365; Plate 31, fg. 2. [Type locality not indicated]
= Palmon dasydactylus Streets, 1871b: 225; Plate 2, Figs 3-3a. [tide-water of the Coatzacoalcos River,
Isthmus of Tehuantepec]
= Palmon sexdentatus Streets, 1871b: 226; Plate 2, Figs 4-4a. [tide-water of the Coatzacoalcos River,
Isthmus of Tehuantepec]
? = Palaemon Potiet Mller, 1892: 184. [nomen nudum]
Macrobrachium acherontium Holthuis, 1977b: 188; Figs 6-7. [Grutas del Cocona, 3 km NE of Teapa, Tabas-
co, altitude 60 m above sea level]
= Macrobrachium coconaensis Guzman, Cabrera & Kensler, 1977: 208. [nomen nudum]
Macrobrachium aemulum (Nobili, 1906b)
= Palaemon (Parapalaemon) aemulus Nobili, 1906b: 258. [Gatavake, 180 mtres daltitude]
Macrobrachium agwi Klotz, 2008: 49; Figs 1-3. [India, West Bengal, Alipurduar District, Barobisha, ap-
prox. 800 km north of Calcuta]
Macrobrachium ahkowi Chong & Khoo, 1987a: 562. [nomen novum for Macrobrachium johnsoni Chong &
Khoo, 1987b]
= Macrobrachium johnsoni Chong & Khoo, 1987b: 360; Figs 1-3; nec Macrobrachium johnsoni Ravin-
dranath, 1979. [frst landing, Gunong Palai waterfall stream, Johore, ca. 200 m altitude]
Macrobrachium altifrons altifrons (Henderson, 1893)
= Palmon altifrons Henderson, 1893: 444; Plate 40, Figs 4-6. [Delhi; River Jumna; Lahore]
Macrobrachium altifrons ranjhai Tiwari, 1964a
= Macrobrachium altifrons ranjhai Tiwari, 1964a: 237; Figs 2A-F, 5-6. [Kabul River at Nowshera, Pesha-
war District (West Pakistan)]
Macrobrachium amazonicum (Heller, 1862b)
= P.[alaemon] amazonicus Heller, 1862b: 418; Plate 2, fg. 45. [im Amazonenstrome]
= Palmon ensiculus Smith, 1869: 26; Plate 1, fgs 2-2c. [Par, Brazil]
= Palaemon Dieperinkii De Man, 1879: 167. [Suriname]
Macrobrachium americanum Spence Bate, 1868a: 363; Plate 30. [Lake of Amatitlan, Guatemala]
Macrobrachium amplimanus Cai & Dai, 1999: 231; Figs 13-15. [forest stream near Mengla County]
Macrobrachium andamanicum (Tiwari, 1952)
= Palaemon andamanicum Tiwari, 1952: 30. [Andaman Islands]
Macrobrachium aracamuni Rodrguez, 1982: 379; Fig. 2. [Cerro Aracamuni, Territorio Federal Amazonas,
680 m above sea level]
Macrobrachium asperulum (von Martens, 1868)
= Palaemon asperulus von Martens, 1868: 43; Plate 1, fg. 5. [Shanghai, aus dem Fischmarkt]
= Palaemon asperulus var. brevirostris Yu, 1931a: 287; Fig. 4. [Ichang, Changsha, Soochow]
= Macrobrachium pinguis Dai, 1984: 245; Figs 1-5. [Longhai County, Fujian Province]
= Macrobrachium anhuiense Tan, 1991: 286; Figs 1-4. [Wuwei County, (3120'N 11754'E), Anhui
Province]
Macrobrachium assamense assamense (Tiwari, 1958)
= Palaemon assamensis assamensis Tiwari, 1958: 298. [Someswari River, near Siu, Garo Hills, Assam]
Macrobrachium assamense peninsulare (Tiwari, 1958)
= Palaemon assamensis peninsularis Tiwari, 1958: 298. [Nerbudda River at Khetgaon, 22.50N 81.20E,
Mandla District, Madhya Pradesh]
Macrobrachium atabapense Pereira S., 1986: 202; Figs 4, 5, 6A. [Atabapo River, Sta. Cruz, Territorio Fed-
eral Amazonas, Venezuela, 320'N 6729'W]
Macrobrachium auratum Short, 2004: 71; Figs 27-28, 36C. [Flame Tree Creek between Airlie Beach and
Shute Harbour, 2016'S 14845'E]
Macrobrachium australe (Gurin-Mneville, 1838 [in Gurin-Mneville, 1829-1838])
= Palaemon australis Gurin-Mneville, 1838 [in Gurin-Mneville, 1829-1838]: 37. [le de Tati]
? = Palaemon Danae Heller, 1865: 120; Plate 11, fg. 3. [Sidney; considered erroneous by Short, 2004]
= Palaemon dispar von Martens, 1868: 41. [Insel Adenare, umweit Flores]
= Palaemon alphonsianus Hofman, 1874: 33; Plate 9, fgs 63-65. [lle de la Runion]
= Palaemon parvus Hofman, 1874: 35; Plate 7, fg. 59. [lle de Nossy-Faly]
= Palaemon Malliardi Richters, 1880: 166; Plate 18, fgs 1-3. [Creole river, Black river]
= P.[almon] (Eupalmon) ustulatus Nobili, 1899: 241. [Rigo]
= Leander lepidus De Man, 1915a: 410; Plate 28, fgs 6-6d. [Mndung des kleinen Flusses zu Oinak,
einem Dorfe an der Kste stlich von der Humboldt-Bai]
Macrobrachium australiense Holthuis, 1950a: 174. [nomen novum for Palaemon australis Ortmann, 1891]
= Palaemon australis Ortmann, 1891: 709; nec Palaemon australis Gurin-Mneville, 1838. [Gayndah,
Rockhampton (both Queensland); Peak Downs, Ost-Australien]
= Macrobrachium adscitum adscitum Riek, 1951: 363; Fig. 3. [Queensland, Accomodation Creek,
Oakey]
= Macrobrachium adscitum subsp. Riek, 1951: 363. [Angus River, Strathalbyn, south of Adelaide,
South Australia]
= Macrobrachium atactum atactum Riek, 1951: 364; Fig. 5. [Conondale, Mary River, Queensland]
= Macrobrachium atactum ischnomorphum Riek, 1951: 364; Fig. 6. [Elimbah Creek, Elimbah, Queens-
land]
= Macrobrachium atactum sobrinum Riek, 1951: 364; Fig. 7. [Mutaburra, Queensland]
= Macrobrachium australiense crassum Riek, 1951: 366; Fig. 10. [Cairns, Queensland]
= Macrobrachium australiense cristatum Riek, 1951: 366; Fig. 9. [Pallal, Horton River, near Bingara,
New South Wales]
= Macrobrachium australiense eupharum Riek, 1951: 365; Fig. 8. [Burdekin River, Macrossan, Queens-
land]
Macrobrachium banjarae (Tiwari, 1958)
= Palaemon banjarae Tiwari, 1958: 299. [Banjar River, of Aonrai Forest Village, Baihar Tehsil, Dist.
Balaghat, M.P.]
Macrobrachium bariense (De Man, 1892a)
= Palaemon (Macrobrachium) bariensis De Man, 1892a: 496; Plate 29, fgs 50-50d. [Flores, aus dem
Flusse bei Bari]
Macrobrachium birai Lobo, Melo & Fernandes, 1986: 50. [Rio Branco, afuente do Itapitangui, Canania,
SP, 2454'44''S 475'30''W; fully described in Melo, Lobo & Fernandes, 1988]
Macrobrachium birmanicum Schenkel, 1902
= Palaemon spinipes var. birmanicus Schenkel, 1902: 503; Plate 9, fgs 8-8c. [Mandalay and Bhamd;
lectotype designated by Cai & Ng, 2002]
Macrobrachium bombajense Almelkar & Sankolli, 2006: 187; Figs 1-2. [Maharasthra State Fisheries Depart-
ment fsh farm Aarey, Goregaon Mumbai, Maharasthra, India]
= Macrobrachium bombayensis Indulkar & Shirgur, 1995: 40. [nomen nudum]
Macrobrachium borellii (Nobili, 1896)
= Palmon Borellii Nobili, 1896: 2. [San Lorenzo (Jujuy) e et diversa della provincia di San Luis]
Macrobrachium brasiliense (Heller, 1862b)
= P.[alaemon] brasiliensis Heller, 1862b: 419; Plate 2, fg. 46. [in einem Bache zu Camaroes, Brasilien]
= Palaemon appuni var. aequatorialis Ortmann, 1891: 731; Plate 47, fg. 6. [Ecuador]
Macrobrachium brevicarpum Tan & Dong, 1996: 287; Figs 1-4. [Qimen County (2952'N 11743'E), Anhui
Province]
Macrobrachium bullatum Fincham, 1987: 351; Fig. 1. [Magela Creek, Northern Territory, Australia]
Macrobrachium cacharense (Tiwari, 1952)
= Palaemon hendersoni cacharensis Tiwari, 1952: 32. [Cachar and Silchar districts, Assam]
Macrobrachium caledonicum (Roux, 1926a)
= Palaemon (Macrobrachium) caledonicus Roux, 1926a: 224; Figs 52-54. [Bond; Tao; Canala; Kon et
environs; Bopope; Coula-Borar; Ni; La Foa; Coind (all Nouvelle-Caldonie)]
Macrobrachium callirrhoe (De Man, 1898)
= Palaemon (Macrobrachium) callirrho De Man, 1898: 152; Plate 8. [Mandai river at Nanga Raoen;
Ketoengau river]
Macrobrachium canarae (Tiwari, 1958)
= Palaemon canarae Tiwari, 1958: 298. [Sitanadi River, near Ghats, South Kanara, Madras]
Macrobrachium carcinus (Linnaeus, 1758)
= Cancer Carcinus Linnaeus, 1758: 631. [Americ fuvis]
= Cancer (Astacus) Jamaicensis Herbst, 1792 [in Herbst, 1791-1796]: 57; Plate 27, fg. 2. [Jamaica, in
Flussen]
= Palaemon brachydactylus Wiegmann, 1836: 148. [Ostkste Mexiko]
= P.[alemon] punctatus Randall, 1840: 146. [?East Indies and West Indies]
= Palemon brevicarpus De Haan, 1849 [in De Haan, 1833-1850]: 172. [Japan; lectotype designation by
Yamaguchi & Baba, 1993; specimens considered to be from Suriname, see Yamaguchi & Baba, 1993]
= Palmon aztecus de Saussure, 1857a: 504. [Vera-Cruz]
? = Palmon Montezum de Saussure, 1857a: 504. [Vera-Cruz]
= P.[alaemon] laminatus von Martens, 1869: 24. [Caracas]
= Palaemon ornatus Torralbas, 1917: 616; Figs 56, 57; nec Olivier, 1811. [corrientes de agua dulce,
Cuba]
= Periclimenes portoricensis Schmit, 1933: 3; Fig. 2. [Porto Rico]
Macrobrachium catonium H.H.III Hobbs & H.H.Jr. Hobbs, 1995: 50; Fig. 1. [Lake in Actun Chapat, Cayo
District, Belize]
Macrobrachium cavernicola (Kemp, 1924)
= Palaemon cavernicola Kemp, 1924: 42; Plate 3, fgs 1-4. [Siu Cave, Garo Hills, Assam]
Macrobrachium chevalieri (Roux, 1935a)
= Palaemon chevalieri (Macrobrachium) Roux, 1935a: 193; Figs 1-2. [Iles du Cap-vert, Saint Antonio
Paul]
Macrobrachium clymene (De Man, 1902)
= Palaemon (Macrobrachium) clymene De Man, 1902: 794; Plate 25, fgs 50-50h. [aus dem Baramfusse,
Borneo]
Macrobrachium cocoense Abele & W. Kim, 1984: 951; Figs 1-2. [Isla del Coco, Costa Rica, stream on east
side of Wafer Bay]
Macrobrachium cortezi Rodrguez, 1982: 383; Fig. 3. [Tobogn, 30 km S of Puerto Ayacucho, Ro Orinoco]
Macrobrachium cowlesi Holthuis, 1950a: 257. [Manila water supply, Luzon, Philippines; described and
illustrated by Cowles, 1914 as Palaemon sp.]
Macrobrachium crebrum Abele & W. Kim, 1989: 6; Fig. 2. [Republic of Panama, Panama Canal, Mirafores
Third Locks Lake]
Macrobrachium crenulatum Holthuis, 1950b: 95. [Pejebobo River. E. Panama]
Macrobrachium dalatense Nguyn, 2003: 454; Figs 1-3. [Mountain stream near the village of Krean at about
1250 m altitude, 1152'N 10829'E, south of Dalat]
Macrobrachium dayanum (Henderson, 1893)
= Palaemon Dayanus Henderson, 1893: 443; Plate 40, fgs 7-13. [Orissa; Jubbulpore; Calcuta; Beer-
bhoom; Debroo; Delhi; Roorkee; Hurdwar; Loodiana; River Jumna; Lahore]
Macrobrachium denticulatum Ostrovski, Da Fonseca & Da Silva-Ferreira, 1996: 361; Figs 4-5. [Brazil, bor-
der between the states of Alagoas and Sergipe, So Francisco river, 937'30''S 3746'W]
Macrobrachium depressimanum Pereira S., 1993
= Macrobrachium depressimanus Pereira S., 1993: 339; Figs 1, 2A. [Per, Amazonas Department, San-
tiago River, La Poza, 401'S 7747'W]
Macrobrachium dienbienphuense ng & B.Y. Nguyn, 1972: 4; Fig. 3. [Nam Rom River, Dien Bien Phu, Lai
Chau Province]
= Macrobrachium longidigitum Dai, 1984: 248; Figs 18-22; nec Spence Bate, 1868a. [Ganlanba, Lancang
River, Yunnan Province, China]
= Macrobrachium eriocheirum Dai, 1984: 247; Figs 13-17. [Jingsan, Xishuangbanna Dai Aut. Pref., Yun-
nan Province]
Macrobrachium dierythrum Pereira S., 1986: 204; Figs 7-8, 12C. [Aguaro River, Paso Garzerito, Edo. Guarico,
Venezuela; 810'N 6625'W]
Macrobrachium digitus Abele & W. Kim, 1989: 8; Figs 3-4. [Republic of Panama, Panama Canal, Mirafores
Locks]
Macrobrachium digueti (Bouvier, 1895)
= Palemon Digueti Bouvier, 1895: 159; Fig. 2. [rivire Mulege et canaux dirrigation issus de cete
rivire]
Macrobrachium dolatum Cai, Naiyanetr & Ng, 2004: 598; Figs 6-7. [Trang, South Thailand]
Macrobrachium dolichodactylus (Hilgendorf, 1879)
= Palaemon (s.s.) dolichodactylus Hilgendorf, 1879: 840; Plate 4, fg. 18. [Tete]
Macrobrachium duri Wowor & Ng, 2010b: 279; Figs 1-2. [Banten, Banten Province, Java]
Macrobrachium dux (Lenz, 1910b)
= Palaemon (Eupalaemon) dux Lenz, 1910b: 129; Plate 3, fgs 2-5. [Avakubi am Ituri]
= Palaemon (Eupalaemon) Lenzii De Man, 1911b: 225. [Congo River, probably near Boma]
= Palaemon (Eupalaemon) dux var. congoensis De Man, 1912a: 416. [la rivire Kole, afuent du Lohali
(Aruwimi), nord-est du Congo Belge]
= Palmon (Eupalmon) dux var. congoensis De Man, 1912b: 229; Plate 4, fgs 6-6a. [la rivire Kole,
afuent du Lohali (Aruwimi), nord-est du Congo Belge]
= Palaemon (Eupalaemon) dux var. tenuicarpus De Man, 1925: 47; Fig. 12k, g, l, h-j . [Kikada, Congo
Belge]
Macrobrachium edentatum Liang & Yan, 1986b: 107; Figs 1-4. [Junlian County, Sichuan Province]
Macrobrachium elatum Jayachandran, 1987: 370; Fig. 2. [Ashtamudi estuary at Perumon, Quilon District,
Kerala]
Macrobrachium elegantum Pan, Hou & S. Li, 2010: 86; Figs 2-4. [unnamed cave in Xiaorui Village, Ludong
Town, Jingxi County (23.1N 106.4E), Guangxi Zhuang Autonomous Region, China]
Macrobrachium empulipke Wowor, 2010: 717; Figs 1-2. [sg. Cibogo, Sukabumi Regency, W. Java]
Macrobrachium equidens (Dana, 1852a)
= Palmon equidens Dana, 1852a: 26. [in mari prope portum Singapore]
= Palaemon sundaicus var. bataviana De Man, 1897: 784. [Fluss zu Batavia]
= P.[almon] (Eupalmon) sundaicus var. brachydactyla Nobili, 1899: 238. [Amboina]
= P.[almon] sundaicus var. De Mani Nobili, 1899: 239. [Atjeh; Amboina]
= P.[almon] (Eupalmon) acanthosoma Nobili, 1899: 242. [Katau]
= Palaemon (Eupalaemon) sundaicus var. baramensis De Man, 1902: 770. [aus dem Baramfusse]
= Palaemon (Eupalaemon) nasutus Nobili, 1903c: 9; unnumbered fg. [Singapore]
= Palaemon delagoae Stebbing, 1915: 74; Plate 16. [mouth of river fowing into Delagoa Bay]
= Urocaridella borradailei Stebbing, 1923: 8; Plate 14. [Umhlotzu River, Natal]
Macrobrachium esculentum (Thallwitz, 1891)
= Palaemon esculentus Thallwitz, 1891: 98. [Nord-Celebes; fully described in Thallwitz, 1892]
= Palaemon dulcis Thallwitz, 1891: 99. [Nord-Celebes; fully described in Thallwitz, 1892]
Macrobrachium faustinum (de Saussure, 1857a)
= Palmon Faustinus de Saussure, 1857a: 505. [Haiti]
= Palaemon cubanus Sharp, 1893: 123. [Cuba; nomen nudum]
= P.[alemon] spinimanus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 399 (partim); nec
Latreille, 1818. [les Antilles et les ctes du Brsil]
Macrobrachium felicinum Holthuis, 1949b: 183. [Catumbela near Benguella, Angola]
Macrobrachium ferreirai Kensley & Walker, 1982: 4; Figs 5-6, 12b. [Igarap near Castanhal, Aripuan]
Macrobrachium feunteuni Keith & Vigneux, 2002: 130; Photo 25-33. [Nuku Hiva, les Marquises, Polynsie
Franaise, rivire Hakaui]
Macrobrachium foai (Coutire, 1902a)
= P.[alaemon] (Eupalmon) Foai Coutire, 1902a: 517. [Haut Congo]
Macrobrachium forcipatum Ng, 1995b: 251; Figs 2-4. [Tasik Temengor, south of Banding, Sungai Halong]
Macrobrachium formosense Spence Bate, 1868a: 364; Plate 31, fg. 1. [River Tamsuy, Island of Formosa]
= Palemon longipes De Haan, 1849 [in De Haan, 1833-1850]: 171; nec Olivier, 1811. [Japan; lectotype
designation by Yamaguchi & Baba, 1993]
= Palmon similis Yu, 1931a: 281; Fig. 2. [Amoy]
? = Palaemon riukiuensis Kubo, 1940a: 21; Figs 12-13; Plate 1B. [Okinawa-zima (Riu-Kiu)]
Macrobrachium fukienense Liang & Yan, 1980: 30; Figs 1-7. [Fujian Province: Xiamen (Bantou, Dongfu),
Tongan (Dingxi), Anxi, Haicang (Jiaomei), Yongchun, Zhangpu (Shiliu)]
Macrobrachium gallus Holthuis, 1952b: 67; Fig. 1. [Rio Peripa, Ecuador]
Macrobrachium gangeticum Spence Bate, 1868a: 365. [Ganges River near Duferin Bridge at Rajghat, Ba-
naras India; neotype selection by Tiwari & Holthuis, 1996]
= Palaemon choprai Tiwari, 1949a: 333; Figs 1-2. [Ganges River near Duferin Bridge at Rajghat, Ba-
naras India; lectotype selection by Tiwari & Holthuis, 1996]
Macrobrachium glabrum Holthuis, 1995: 139; Figs 1-2. [Kotomay, sous-prfecture Mitsinjo, prov. Majunga,
N.W. Madagascar]
Macrobrachium gracilirostre (Miers, 1875)
= Palaemon gracilirostris Miers, 1875: 343. [Samoa Islands, Upolu]
= Palaemon (Parapalaemon) modestus De Man, 1892a: 469; Plate 27, fgs 43-43d; nec Heller, 1862a.
[Flores, Fluss bei Wukur unweit Sikka]
= Palaemon (Parapalaemon) modestus brevimanus Roux, 1934a: 228; Figs 9- 10. [Bimun, Nouvelle-
Irlande]
= Macrobrachium sophronicum Holthuis, 1950a: 198; Fig. 40. [nomen novum name for Palaemon (Para-
palaemon) modestus De Man, 1892a]
Macrobrachium grandimanus (Randall, 1840)
= P.[alemon] grandimanus Randall, 1840: 142. [Sandwich Islands]
= P.[alemon] gracilimanus Randall, 1840: 143. [Sandwich Islands]
= Palmon acutirostris Dana, 1852a: 26. [insulis Hawaiensibus]
Macrobrachium gua Chong, 1989: 32; Figs 1-2. [stream at resurgence from Gomantong Hill, ca. 533'N
11806'E, Sabah, Borneo]
Macrobrachium guangxiense Liang & Yan, 1981: 33; Figs 20-21. [Longzhou County, Guangxi Province]
Macrobrachium gurudeve Jayachandran & Raji, 2005: 1180; Figs 1-2, 6. [Bhavani River (at Thavalam), an
east fowing mountain river, Palakkad District of Kerala State]
Macrobrachium hainanense (Parisi, 1919)
= Palaemon (Parapalaemon) hainanense Parisi, 1919: 87; Plate 2, fg. 1; Plate 6, fgs 1, 7. [Fiume Ken-
Kong, Isla di Hainan]
Macrobrachium hancocki Holthuis, 1950b: 96. [Esparta, Rio Barranca, Costa Rica]
Macrobrachium handschini (Roux, 1933)
= Palaemon (Macrobrachium) handschini Roux, 1933: 345. [Katherine River, Northern Territory, Aus-
tralia; lectotype designation by Bruce, 1992b]
= Macrobrachium glypticum Riek, 1951: 363; Fig. 4. [Coen, north Queensland]
Macrobrachium hendersodayanum (Tiwari, 1952)
= Palaemon henderso-dayanus Tiwari, 1952: 29. [Western Ghats, from Satara District up to Mysore State]
Macrobrachium hendersoni (De Man, 1906)
= Palmon (Parapalmon ?) Hendersoni De Man, 1906: 405. [Darjeeling, Bengal, at a height of 2500
feet]
= Palaemon yunnanensis Y, 1936: 308; Figs 3, 4. [Mann-Tchi-Pan]
Macrobrachium heterochirus (Wiegmann, 1836)
= Palaemon heterochirus Wiegmann, 1836: 149. [Ostkste Mexiko]
= Palaemon Appuni von Martens, 1869: 31; Plate 2, fg. 5. [Porto Cabello in Venezuela]
Macrobrachium heterorhynchos Guo & He, 2008: 12; Figs 1-2, 7A-B. [Xiiang river near Jiangmen City, ca.
2233'N 11308'E]
Macrobrachium hildebrandti (Hilgendorf, 1893b)
= Bithynis? hildebrandti Hilgendorf, 1893b: 244. [Central-Madagaskar]
Macrobrachium hirsutimanus (Tiwari, 1952)
= Palaemon hirsutimanus Tiwari, 1952: 31. [forest stream 95 km north from Nan Town, Nan province,
about 15-20 km north from Ban Pon, Nam Gae, north of Ban Sala, near Laos, North Thailand; neo-
type selection by Cai, Naiyanetr & Ng, 2004]
Macrobrachium hirtimanus (Olivier, 1811)
= Palmon hirtimanus Olivier, 1811: 663. [la mer des Indes]
Macrobrachium hobbsi Villalobos Hiriart & Nates Rodriguez, 1990: 7; Fig. 3. [Mxico, Chiapas, Ro El
Naranjo, aprox. a 8 km al NO de Piiiapan, carretera Tonal-Piiipan]
Macrobrachium holthuisi Genofre & Lobo, 1978: 273; Fig. 1. [Guaec river of So Sebastio, State of So
Paulo, Brazil]
Macrobrachium horstii (De Man, 1892a)
= Palaemon (Parapalaemon) Horstii De Man, 1892a: 460; Plate 27, fgs 39-39c. [Celebes, aus dem Flusse
bei Palopo, Luwu]
= Palaemon (Parapal.) horsti brevidigitus Roux, 1930: 358. [Bali]
Macrobrachium idae (Heller, 1862b)
= P.[alaemon] Idae Heller, 1862b: 416; Plate 2, fgs 40-41. [Borneo; lectotype selection by Cai & Ng,
2001b]
= Palaemon (Eupalaemon) ritsemae De Man, 1897: 774; Plate 37, fgs 70-70c. [Atjeh]
= Palmon (Eupalmon) Id var. subinermis Nobili, 1899: 237. [Innawi, Fiume San Guiseppe]
= Palmon (Eupalmon) Mari Coutire, 1900a: 1266. [Rivire Ivaloina, prs Tamatave]
= Palaemon (Eupalaemon) robustus De Man, 1902: 771; Plate 24, fg. 48. [Tabelo, Halmahera; lectotype
selection by Cai & Ng, 2001b]
= Macrobrachium palawanensis Johnson, 1962: 307; Fig. 1. [Palawan]
Macrobrachium idella idella (Hilgendorf, 1898)
= Palaemon (Eupalaemon) idae var. idella Hilgendorf, 1898: 29; Fig. A. [Ungu, Teich bei Matomondo;
Usaramo]
= Palmon (Eupalmon) multidens Coutire, 1900a: 1266. [Riv. Kotofotsy, Bras de lOnilahy]
Macrobrachium idella georgii Jayachandran & Joseph, 1985a: 130; Fig. 1. [Pallickal, Pamba, Manimala and
Meenachil rivers]
Macrobrachium iheringi (Ortmann, 1897)
= Palaemon iheringi Ortmann, 1897: 211; Plate 1, fgs 7-8. [Alto da Serra and Rio Tiet, Estado de So
Paulo]
Macrobrachium inca Holthuis, 1950b: 93. [Rio Moch, near Salavery, N. Peru]
Macrobrachium indicum Jayachandran & Joseph, 1986: 217; Figs 1-4. [Vellayani Lake, 824'09"-86'30"N
7659'08"-7659'47"E]
Macrobrachium infatum Liang & Yan, 1985b: 254; Fig. 3. [Kunshan County, 3120'N 12058'E, Jiangsu
Province]
Macrobrachium inpa Kensley & Walker, 1982: 6; Figs 7-9, 12c. [Igarap da Cachoeira]
Macrobrachium insulare (Parisi, 1919)
= Palaemon (Parapalaemon) insularis Parisi, 1919: 85; Fig. 7; Plate 3, fgs 2-3; Plate 6, fg. 12. [Formosa]
Macrobrachium jacobsoni Holthuis, 1950a: 227; Fig. 47. [Lugu, Sinabang Bay, Simalur, of the west coast of
Sumatra]
Macrobrachium japonicum (De Haan, 1849 [in De Haan, 1833-1850])
= Palaemon japonicus De Haan, 1849 [in De Haan, 1833-1850]: 172. [Japan; lectotype designation by
= Palaemon boninensis Stimpson, 1860a: 41. [insulis Bonin, in rivulis montanis]
Macrobrachium jaroense (Cowles, 1914)
= Palaemon jaroensis Cowles, 1914: 385; Plate 3, fgs 8-8k. [Hibucawan River, near Jaro, Leyte]
Macrobrachium jayasreei Jayachandran & Raji, 2005: 1186; Figs 3-4. [Kabbini River (at Mananthavadi), an
east fowing mountain river, Wynad District of Kerala State]
Macrobrachium jelskii (Miers, 1877)
= Palmon jelskii Miers, 1877: 661; Plate 67, fg. 1-1b. [Guiana, Oyapock]
Macrobrachium jiangxiense Liang & Yan, 1985b: 256; Fig. 4. [Boyang County, 29N 11640'E, Jiangxi
Provinces]
Macrobrachium johnsoni Ravindranath, 1979: 184; Figs 1-2. [Guntur fsh market]
Macrobrachium joppae Holthuis, 1950a: 233; Fig. 48. [Nias]
Macrobrachium kelianense Wowor & Short, 2007: 82; Figs 3-4. [Sg. Kelian confuence with Sg. Ketang,
Makaham River basin, E. Kalimantan]
Macrobrachium kempi (Tiwari, 1949b)
= Palaemon kempi Tiwari, 1949b: 330. [small stream between Chitagong and Sultan Bagu Bastan]
Macrobrachium kistnense (Tiwari, 1952)
= Palaemon kistnensis Tiwari, 1952: 28. [Vindhyan Satpuran range of mountains, beginning from the
Tanda Falls in the Mirzapur District, U.P.; Western Ghats; Ceylon, Aravalli Hills]
Macrobrachium kiukianense (Yu, 1931a)
= Palmon kiukianensis Yu, 1931a: 279; Fig. 1. [Kiukiang]
Macrobrachium koombooloomba Short, 2004: 61; Figs 23-24, 36A. [Koombooloomba area, creek on Koom-
booloomba road near Koombooloomba Dam turnof, 750'S 14534'E]
Macrobrachium kulkarnii Almelkar & Sankolli, 2006: 190; Figs 3-4. [fsh farm of M/s Tata Hydro-Electric
Company, Walvan Dam, Lonavala (Pune District), Maharasthra State, India]
Macrobrachium kunjuramani Jayachandran & Raji, 2005: 1188; Fig. 5. [Kabbini River (at Chundal, Kalpet-
ta, Vythiri, Thalapuzha, Noolpuzha, Kenipuzha), east fowing mountain river, Wynad District of
Kerala State]
Macrobrachium lamarrei lamarrei (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840])
= P.[alemon] Lamarrei H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 397. [ctes du Ben-
gale]
Macrobrachium lamarrei lamarroides (Tiwari, 1952)
= Palaemon lamarrei lamarroides Tiwari, 1952: 28. [Logtak Lake, Manipur, Assam]
Macrobrachium lanatum Cai & Ng, 2002: 72; Figs 13-16. [southern Myanmar]
Macrobrachium lanceifrons (Dana, 1852a)
= Palmon lanceifrons Dana, 1852a: 26. [insula Luzon, archiepelagi Phillipinsis, prope portum
Manila]
= Palaemon lanceifrons var. montalbanensis Cowles, 1914: 371; Plate 2, fgs 6-6i. [city of Manila at Mon-
talban]
Macrobrachium lanchesteri (De Man, 1911c)
= Pal.[aemon] (Eupalaemon) Lanchesteri De Man, 1911c: 264. [nomen novum for Palmon paucidens
Lanchester, 1901]
= Palmon paucidens Lanchester, 1901: 568; Plate 33, fg. 4; nec De Haan, 1844, nec Hilgendorf, 1893a.
[Singora]
Macrobrachium lar (Fabricius, 1798)
= Palaemon Lar Fabricius, 1798: 402. [in India]
= Palaemon longimanus Fabricius, 1798: 402. [in India orientali]
= Palmon ornatus Olivier, 1811: 660. [Nouvelle-Hollande]
= Palmon tridens White, 1847a: 78. [nomen nudum]
= P.[alaemon] vagus Heller, 1862b: 417; Plate 2, fgs 42-43. [Amboina]
= Palaemon spectabilis Heller, 1862a: 527. [Taiti]
= Palaemon ruber Hess, 1865: 165; Plate 7, fg. 20. [Viti-Inseln]
= Palaemon spectabilis Heller, 1865: 113; Plate 10, fg. 8. [Taiti]
= Palaemon longimanus Hofman, 1874: 34; Plate 9, fgs 68-69; nec Fabricius, 1798. [lle de la Ru-
nion]
= Palaemon mayotensis Hofman, 1874: 32; Plate 9, fgs 61-62. [lle de Mayote; lle de Nossy-Faly]
= Palaemon reunionnensis Hofman, 1874: 33; Plate 9, fgs 66-67. [lle de la Runion]
= Palaemon madagascariensis Hofman, 1874: 35; Plate 7, fg. 58. [lle de Nossy-Faly]
= Leander dionyx Nobili, 1905d: 482; Plate 12, fg. 2. [Stephansort]
= Cancer teatae Curtiss, 1938: 162. [Vaitapiha River, Tautira township, Tahiti]
? = Macrobrachium ornatus Jayachandran & Raji, 2004: 41; Figs 1-2. [small rivulet of Muvatupuzha
river at Pothanikkad, Kerala]
Macrobrachium latidactylus (Thallwitz, 1891)
= Palaemon latidactylus Thallwitz, 1891: 97. [Nord-Celebes; fully described in Thallwitz, 1892]
= Palaemon (Eupalaemon) endehensis De Man, 1892a: 465; Plate 27, fgs 42-42h. [Endeh, aus dem Nan-
ga Ba; aus dem Flusse bei Reo; aus dem Flusse bei Bari; Mbawa, dicht am Meere; Sungei Nargi bei
Konga (all localities from Flores)]
= Palaemon (Macrobrachium) lampropus De Man, 1892a: 493; Plate 29, fgs 49-49c. [aus einem Flusse
bei Palopo; aus dem Wasserfalle Bantimurong unweit Maros; aus einem Flusse bei Pare-Pare (all
from Celebes); Timor, aus dem Flusse Koinino bei Kupang]
Macrobrachium latimanus (von Martens, 1868)
= Palaemon latimanus von Martens, 1868: 44. [Philippinen, bei Loquilocun auf der Insel Samar]
= Palaemon euryrhynchus Ortmann, 1891: 738; Plate 47, fg. 12. [Fidji-Levu; Fidji-Ins.]
= Palmon (Macrobrachium) singalangensis Nobili, 1900a: 487. [Aier Mantcior]
Macrobrachium lepidactyloides (De Man, 1892a)
= Palaemon (Macrobrachium) lepidactyloides De Man, 1892a: 497; Plate 29, fgs 51-51c. [Flores, aus dem
Flusse bei Mbawa, oberhalb des Wasserfalls]
Macrobrachium lepidactylus (Hilgendorf, 1879)
= Palaemon (s.s.) lepidactylus Hilgendorf, 1879: 838; Plate 4, fgs 14-16. [Quellimane; Tete]
= Palmon (Macrobrachium) Hilgendorf Coutire, 1899b: 382. [Madagascar, Cte Est, region des
grandes forts]
Macrobrachium leptodactylus (De Man, 1892a)
= Palaemon pilimanus var. leptodactylus De Man, 1892a: 476; Plate 28, fg. 44i-l. [Buitenzorg, Java, In-
donesia; lectotype selection by Cai, Naiyanetr & Ng, 2004]
Macrobrachium leucodactylus Wowor & Choy, 2001: 274; Figs 4-6. [tributary of Sg. Temburong-Machang,
Temburong District, altitude 580 m]
Macrobrachium lingyunense J. Li, Cai & Clarke, 2006: 277; Figs 1-3. [Shadong (Sand Cave), Guancang Vil-
lage, Sicheng Town, Lingyun County, Guangxi Province, southern China, 10623'-10655'33''E
2405'-2437'N]
Macrobrachium lopopodus Wowor & Choy, 2001: 270; Figs 2-3. [Ulu Sg. Sawatan, Kimanis, Papar District,
Sabah, Malaysia]
Macrobrachium lorentzi (Roux, 1921)
= Palaemon (Parapalaemon) lorentzi Roux, 1921: 596; Plate 16, fgs 1-3. [Kllof-Biwak; Alkmaar; Went-
Geb; Sabang (all Irian Jaya)]
Macrobrachium lucifugum Holthuis, 1974
= Macrobrachium faustinum lucifugum Holthuis, 1974: 233; Figs 2-3. [Cueva del Agua de Yara, on the
territory of the bario Yara, E. of Baracoa, Oriente Province, Cuba]
= Macrobrachium crybelum Chace, 1975: 30; Figs 1-4. [Cave at Ciudad del Caribe, 1858'N 7023'W,
Santo Domingo, D.N., Dominican Republic]
Macrobrachium lujae (De Man, 1912a)
= Palaemon (Eupalaemon) Luj De Man, 1912a: 415. [Kondu, dans le district de Kasa, sur la rivire
Sankuru, dans les parages de Lusambo]
= Palmon (Eupalmon) Luj De Man, 1912b: 215; Plate 2, fgs 3-3b; Plate 3, fgs 3-3e. [Kondu, dans
le district de Kasa, sur la rivire Sankuru, dans les parages de Lusambo]
Macrobrachium macrobrachion (Herklots, 1851)
= Palemon macrobrachion Herklots, 1851: 15. [Boutry] [species name spelled mabrobrachion on page 15
and macrobrachion on page 25; former treated as an inadvertent spelling mistake by Holthuis, 1951a]
= Palmon africanus Kingsley, 1883: 107. [West coast of Africa]
Macrobrachium maculatum Liang & Yan, 1980: 31; Figs 8-14. [Fujian Province: Yongan (Jiulongxi),
Liancheng (Xianshuitang)]
Macrobrachium madhusoodani Unnikrishnan, P.M. Pillai & Jayachandran, 2011: 123; Fig. 1. [Ithikkara River,
856'42''N 7656'20''E, Kerala, South India]
= Macrobrachium madhusoodani Unnikrishnan, P.M. Pillai & Jayachandran, 2010: 1116; Figs 1-2. [upper
reaches of the Ithikkara River, 856'42''N 7656'20''E, Kerala, South India; unavailable under Art. 16.4.2]
Macrobrachium malayanum (Roux, 1935b)
= Palmon (Macrobrachium) pilimanus malayanus Roux, 1935b: 32. [Lasah, Plus Valley, East Perak,
Malay Peninsula]
= Macrobrachium geron Holthuis, 1950a: 258; Fig. 52. [Banka, E. of Sumatra]
Macrobrachium malcolmsonii kotreeanum Johnson, 1973: 279. [Kotree, Indus River, Pakistan]
Macrobrachium malcolmsonii malcolmsonii (H. Milne Edwards, 1844)
= Palemon Malcolmsonii H. Milne Edwards, 1844: 8; Plate 3. [eaux douces prs de Naguepore, une
distance denviron 200 lieues de la mer]
Macrobrachium mammillodactylus (Thallwitz, 1892)
= Palaemon idea var. mammillodactylus Thallwitz, 1892: 15. [Nord-Celbes; Luzon]
= Palaemon (Eupalaemon) Wolterstorf Nobili, 1900b: 1. [Srabaia-Giava]
= Palaemon philippinensis Cowles, 1914: 340; Plate 2, fgs 2-2m. [San Juan River; Pasig River in the city
of Manila]
? = Palmon talavar Blanco, 1939b: 168; Plate 2. [Lake Sampaloc, San Pablo, Laguna province, Luzon]
Macrobrachium manipurense (Tiwari, 1952)
= Palaemon manipurensis Tiwari, 1952: 30. [Manipur, Assam]
Macrobrachium manningi Pereira & Lasso, 2007: 134; Figs 1-3. [Cucurital River, afuent of Caroni River,
Orinoco Basin, Cainama National Park, Bolivar State, Venezuela, 600'N 6247'W, 380 m a.s.l.]
Macrobrachium meridionale Liang & Yan, 1983b
= Macrobrachium meridionalis Liang & Yan, 1983b: 213; Fig. 2. [Jongjiang River, Chengmai County,
Hainan Island, China]
Macrobrachium michoacanus Villalobos Hiriart & Nates Rodriguez, 1990: 2; Fig. 2. [Mxico, Michoacn,
Ro Mexcalhuacn, aprox. 40 km al NO de Playa Azul, carretera Azul-Caleta de Campos]
Macrobrachium microps Holthuis, 1978b: 210; Figs 1-2. [Danmin Cave, near Konogusgus, New Ireland]
Macrobrachium mieni ng, 1975: 68; Fig. 2. [streams in Kim Boi, Hoa Binh Province]
Macrobrachium minutum (Roux, 1917)
= Palaemon minutus Roux, 1917: 599; Plate 27, Figs 1-3. [Lac Sentani]
Macrobrachium miyakoense Komai & Fujita, 2005: 14; Figs 1-6. [anchialine cave located at southern coast
of Miyako Island, Ryukyu Islands]
Macrobrachium moorei (Calman, 1899a)
= Palaemon moorei Calman, 1899a: 709; Plate 40, fgs 20-24. [Lake Tanganyika, 50f]
Macrobrachium naso (Kemp, 1918a)
= Palaemon naso Kemp, 1918a: 91; Plate 25, fgs 1-5. [Sizon, Tang-do cirle of Yawnghwe State in the
river that runs south from the Inl Lake]
Macrobrachium natereri (Heller, 1862b)
= P.[alaemon] Natereri Heller, 1862b: 414; Plate 2, fgs 36-37. [Brasilien, im Rio Negro]
Macrobrachium natulorum Holthuis, 1984a: 164; Figs 2-3. [Jawej River near Tigi Lake]
Macrobrachium neglectum (De Man, 1905)
= Palaemon (Eupalaemon) neglectus De Man, 1905: 201; Plate 15, fg. 6. [lower Batak landen, near Deli,
North Sumatra; lectotype designation by Wowor & Ng, 2010b]
Macrobrachium nepalense Kamita, 1974: 10; Fig. 4; Plate 2, fg. 2. [Tarahara Farm (Birat.) of the Breeding
and Distribution Centre]
Macrobrachium niloticum (Roux, 1833)
= Palmon Niloticus Roux, 1833: 73; Plate 7, fg. 2. [les eaux de Nil]
Macrobrachium niphanae Shokita & Takeda, 1989: 148: Figs 1-2; Plate 1. [Nang Rong waterfall stream,
Thailand]
Macrobrachium nipponense (De Haan, 1849 [in De Haan, 1833-1850])
= Palaemon nipponense De Haan, 1849 [in De Haan, 1833-1850]: 171. [Japan; lectotype designation by
= Palaemon asper Stimpson, 1860a: 41; nec Latreille, 1818. [in fuvii et rivulis Sinenses prope urbem
Canton]
= Palaemon sinensis Heller, 1862a: 528. [Schanghai]
= Palaemon sinensis Heller, 1865: 119; Plate 10, fg. 11. [Shanghai]
= Macrobrachium obtusifrons Dai, 1984: 246; Figs 6-12. [Guanting Reservoir, Miyun County, Beiing]
= Macrobrachium meishanense Tan & Lu, 1992: 169; Figs 1-4. [rivulet from Meishan Reservoir, Jinzhai
County, west Anhui Province]
Macrobrachium nobilii (Henderson & Mathai, 1910)
= Palaemon nobilii Henderson & Mathai, 1910: 295; Plate 17, fgs 6a-e. [Walajabad, Chingleput Dis-
trict]
Macrobrachium novaehollandiae (De Man, 1908c)
= Pal. (Eupal.) nov-hollandi De Man, 1908c: 370; Plate 16. [neighbourhood of Sydney]
Macrobrachium occidentale Holthuis, 1950b: 95. [Rio de los Esclavos, just S of Cuilapa, Guatemala]
Macrobrachium oenone (De Man, 1902)
= Palaemon (Macrobrachium) oenone De Man, 1902: 784; Plate 25, fgs 49-49c. [Kau and Saluta, Halma-
hera]
= Palaemon (Macrobrachium) oenone papuana Roux, 1927b: 324; Fig. 2. [Pionierbovak, Fl. Mamberamo,
Nouvelle-Guine]
Macrobrachium ohione (Smith, 1874)
= Palmon Ohionis Smith, 1874: 640. [Ohio River at Cannelton, Indiana]
= P.[alaemon] sallei Kingsley, 1883: 108. [Mississippi]
Macrobrachium olfersii (Wiegmann, 1836)
= Palaemon Olfersii Wiegmann, 1836: 150. [Kste Brasiliens]
= P.[alemon] spinimanus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840] nec Latreille,
1818: 399 (partim). [les Antilles et les ctes du Brsil]
= Palmon consobrinus de Saussure, 1857a: 504. [Vera-Cruz]
= Palaemon Desausuri Heller, 1862b: 420; Plate 2, fg. 47. [Neu-Granada]
= Palaemon Potiporanga Mller, 1880: 152. [Itajahy]
Macrobrachium oxyphilus Ng, 1992: 442; Figs 1A, C, E, G; 2A, C; 3A, C. [39 km road marker, Sungai Besar
to Tanjong Malim trunk road, north Selangor peat swamp forest, Peninsular Malaysia]
Macrobrachium panamense Rathbun, 1912a
= Macrobrachium acanthurus panamense Rathbun, 1912a: 1. [Rio Calabre, Republic of Panama]
Macrobrachium patheinense Phone & Suzuki, 2004: 524; Figs 1-2. [Mayan Creek, Thayet Kone village,
Pathein City, Ayeyawaddy Division]
Macrobrachium patsa (Coutire, 1899b)
= Palmon (Parapalmon) Patsa Coutire, 1899b: 382. [Madagascar, Rivire Mahanara (Cte Est);
Bras de lOnilahy (Cte Ouest)]
Macrobrachium pectinatum Pereira S., 1986: 200; Figs 2, 3, 6B. [Atabapo River, Sta. Cruz, Territorio Fed-
eral Amazonas, Venezuela, 320'N 6729'W]
Macrobrachium peguense (Tiwari, 1952)
= Palaemon peguensis Tiwari, 1952: 27. [Burma]
Macrobrachium pentazona He, Gao & Guo, 2009: 39; Figs 1-3. [Beiiang River, near Qingyuan City (ca.
2342'N 11301'E)]
Macrobrachium petersii (Hilgendorf, 1879)
= Palaemon (s.s.) Petersii Hilgendorf, 1879: 841; Plate 44, fg. 19. [Tete]
Macrobrachium petiti (Roux, 1934b)
= Palaemon (Macrobrachium) Petiti Roux, 1934b: 537; Figs 1-2. [Vatomandry]
Macrobrachium petronioi Melo, Lobo & Fernandes, 1986: 51. [Rio Branco, afuente do Itapitangui,
Canania, SP, 2454'44''S 4758'30''W; fully described in Melo, Lobo & Fernandes, 1988]
Macrobrachium pilimanus (De Man, 1879)
= Palaemon pilimanus De Man, 1879: 181. [Moearalboeh, West Sumatra, Indonesia; lectotype selec-
tion by Cai, Naiyanetr & Ng, 2004]
= Palaemon (Macrobrachium) pygmaeus Roux, 1928b: 222; Figs 1-4. [Telaga Kastobo, Bawean]
Macrobrachium pilosum Cai & Dai, 1999: 226: Figs 10-12. [mountain stream near Mengban village, Meng-
la County]
Macrobrachium placidulum (De Man, 1892a)
= Palaemon (Macrobrachium) placidulus De Man, 1892a: 489; Plate 28, fgs 48-48g. [Fluss bei Mbawa,
oberhalb des Wasserfalls; aus dem Flusse Dona bei Endeh; Fluss Ba bei Endeh; Fluss Lella bei Sikka,
aus dem Flusse Wukur bei Sikka (all from Celebes); Saleyer, aus dem Flusse Bangkalan; Celebes,
Fluss bei Palopo; Timor, aus dem Flusse Koinino bei Kupang]
? = Palmon spinimanus Latreille, 1818: 5; Plate 319, fg. 1. [Type locality not indicated]
Macrobrachium placidum (De Man, 1892a)
= Palaemon (Macrobrachium) placidus De Man, 1892a: 483; Plate 28, fgs 46-46c. [Sumatra, Fluss bei
Kaju-tanam]
Macrobrachium platycheles Ou & Yeo, 1995: 300; Figs 1-3. [stream leading from Nee Soon swamp forest,
150 m upstream from pipeline, Singapore]
Macrobrachium platyrostris (Tiwari, 1952)
= Palaemon hendersoni platyrostris Tiwari, 1952: 32. [Type locality not indicated]
Macrobrachium poeti Holthuis, 1984b: 143; Fig. 1. [Luwang Jurangjero, grid 668115, ca 100 m below entrance]
Macrobrachium potiuna (Mller, 1880)
= Palaemon Potiuna Mller, 1880: 152. [aus dem Itajahy; fully described and illustrated in Mller, 1892]
Macrobrachium praecox (Roux, 1928c)
= Palaemon (Eupalaemon) praecox Roux, 1928c: 43. [El Mene; El Pozon (both Prov. Falcon, Vnzuela);
Santander, Colombie]
Macrobrachium pumilum Pereira S., 1986: 208; Figs 11, 12B. [Aguaro River, Cachimbo pass, Edo. Guarico,
Venezuela; 810'N 6635'W]
Macrobrachium purpureamanus Wowor, 1999: 34; Figs 1A-B, 2A, C, E-F, 3A, C, E, 4. [Sungai Nibong B,
nearby Kampung Baru, Kundur Island, Riau Province, Indonesia, 046'46.8''N 10328'05.4''E]
Macrobrachium quelchi (De Man, 1900)
= Palmon (Macrobrachium) Quelchi De Man, 1900: 57; Plate 6, fgs 1-8. [Upper Mazaruni River,
2500f; Mt. Roirama range, 3500f]
Macrobrachium raridens (Hilgendorf, 1893c)
= Palaemon (Eupalaemon?) paucidens Hilgendorf, 1893a: 155; nec De Haan, 1844 [in De Haan, 1833-
1850]. [Adeli bei Bismarckburg, Togoland]
= Palaemon (Eu.[palaemon]) raridens Hilgendorf, 1893c: 181. [nomen novum for Palaemon (Eupalae-
mon?) paucidens Hilgendorf, 1893a]
Macrobrachium rathbunae Holthuis, 1950b: 94. [Hog Creek Valley, San Jos Island, Archipielago de las
Perlas, Gulf of Panama]
Macrobrachium reyesi Pereira S., 1986: 198; Figs 1, 6C. [Quebrada Corral de Piedra, El Limn, Maracay,
Edo. Aragua, Venezuela, 1015'N 6735'W]
Macrobrachium rhodochir Ng, 1995a: 182; Fig. 1. [small stream parallel to Tajur Waterfall, Sungaj Tajur]
Macrobrachium rodriguezi Pereira S., 1986: 206; Figs 10, 12A. [Caris River, El Tigre, Edo. Anzoategui,
Venezuela, 845'N 6450'W]
Macrobrachium rogersi (Tiwari, 1952)
= Palaemon rogersi Tiwari, 1952: 31. [Arakan and Pegu Yomas, Burma]
Macrobrachium rosenbergii (De Man, 1879)
= Palaemon Rosenbergii De Man, 1879: 167. [Batavia (Jakarta), Java, Indonesia; neotype designated by
Wowor & Ng, 2008; see Anon., 2010]
= Palaemon dAcqueti Sunier, 1925: cxvii. [Batavia, Java, Indonesia; lectotype designation by Wowor
& Ng, 2007]
= Palaemon whitei Sharp, 1893: 122. [Bombay]
= Macrobrachium rosenbergii schenkeli Johnson, 1973: 274, 277. [Tavoy, Burma]
Macrobrachium rostratum X. Wang, 1997: 460; Fig. 2. [Jishou, Hunan Province]
Macrobrachium rude (Heller, 1862a)
= Palaemon rudis Heller, 1862a: 527. [Ceylon]
= Palaemon rudis Heller, 1865: 114. [Ceylon]
= Palaemon (s.s.) Mossambicus Hilgendorf, 1879: 839; Plate 4, fg. 17. [Quellimane; Moambique]
= Palaemon (Eupalaemon) Alcocki Nobili, 1903a: 9; Plate 2, fg. 5. [Pondichry]
Macrobrachium sabanus Ng, 1994: 76; Figs 4-5. [waterfall stream, of Safoda Oli Palm Plantation, between
5 and 10 km marks]
Macrobrachium saigonense Nguyn, 2006a: 236; Figs 1, 2a-b, 3a-j, 4-6. [near Hoa An Bridge, on the Dong
Nai river, Bien Hoa province, about 30 km northwest of Ho-Chi-Minh City, 1055'N 10650'E]
Macrobrachium sankolli Jalihal & Shenoy in Jalihal, Shenoy & Sankolli, 1988: 11; Figs 3-4. [Malaprabha
River, near old bridge, Khanapur]
Macrobrachium santanderensis Garcia-Perez & Villamizar, 2009: 62; Figs 2-4. [70 km on an east to west
road connecting the cities of Bucaramanga and Barrancabermeja (Department of Santander, Central
East of Colombia, South America), 900 m above sea level, 0706'15''N 7324'44''W]
Macrobrachium sbordonii Meja-Ortz, Baldari & Lpez-Meja, 2008: 50; Figs 2-4. [terminal lake in the
Sistema de la Lucha, Chipas, Mexico, 1703'40''N 9353'23''W]
Macrobrachium scabriculum (Heller, 1862a)
= Palaemon scabriculus Heller, 1862a: 527. [Ceylon]
= Palaemon scabriculus Heller, 1865: 117; Plate 10, fg. 9. [Ceylon]
= P.[alaemon] dubius Henderson & Mathai, 1910: 300; Plate 18, fgs 9a-d. [Walajabad, Saidapet and
other localities in the Chingleput District]
Macrobrachium scorteccii Maccagno, 1961: 336; Plates 17-18. [Cl Gallon, Somalia]
Macrobrachium shaoi Cai & Jeng, 2001: 276; Figs 1-2. [stream of tributary of Shuangchi river in Taipei
County, northern Taiwan]
Macrobrachium shokitai Fujino & Baba, 1973: 101; Figs 1-4. [River head, Urauchi River, Iriomote Island,
Ryukyu Islands]
Macrobrachium sintangense (De Man, 1898)
= Palaemon (Eupalaemon) sintangensis De Man, 1898: 138; Plate 6. [Sintang]
= Palaemon (Eupalaemon) elegans De Man, 1892a: 440; Plate 26, fgs 36-36d; nec Rathke, 1837. [Buiten-
zorg; Sinagar (Java)]
Macrobrachium sirindhorn Naiyanetr, 2001: 610; Figs 1-2; Plate 1. [Pong Nam Dung Waterfall, Mae Soon,
Amphoe Fang, Chiang Mai Province, northern Thailand]
Macrobrachium siwalikense (Tiwari, 1952)
= Palaemon siwalikensis Tiwari, 1952: 28. [base of Simla Hills]
Macrobrachium sollaudii (De Man, 1912a)
= Palaemon (Eupalaemon) Sollaudii De Man, 1912a: 413. [dans la rivire Otenge, prs de Banzyville,
dans le Congo Belge]
= Palmon (Eupalmon) Sollaudii De Man, 1912b: 205; Plate 1, fgs 2-2i. [dans la rivire Otenge, prs
de Banzyville, dans le Congo Belge]
Macrobrachium spinipes (Schenkel, 1902)
= Palaemon spinipes Schenkel, 1902: 501; Plate 9, fgs 7-7b; nec Desmarest, 1817. [Kema, Minahasa; see
Ng & Wowor, 2011]
= Macrobrachium wallacei Wowor & Ng, 2008 (nomen novum for Macrobrachium rosenbergii sensu
Wowor & Ng, 2007): 291. [Laloki River, ca. 30 miles from Port Moresby, Papua New Guinea]
Macrobrachium spinosum Cai & Ng, 2001b: 676; Figs 8-9. [Sungai Dadago, Halmahera]
Macrobrachium srilankense Costa, 1979: 60; Fig. 6; Plate 1D. [Negombo lagoon; Kelni river at Kelaniya;
Panadura]
Macrobrachium striatum N.N. Pillai, 1991
= Macrobrachium striatus N.N. Pillai, 1991: 249; Figs 1-3. [Cochin backwater around Thevara area,
Kerala State, South India]
Macrobrachium sulcatus (Henderson & Mathai, 1910)
= Palaemon sulcatus Henderson & Mathai, 1910: 289; Plate 16, fg. 4a-g. [Cochin]
Macrobrachium sulcicarpale Holthuis, 1950a: 220; Fig. 45. [Bangkalan River, Salajar, SE of Celebes]
Macrobrachium sundaicum (Heller, 1862b)
= P.[alaemon] sundaicus Heller, 1862b: 415; Plate 2, fgs 38-39. [Java, Indonesia; lectotype designated
by Wowor & Ng, 2010b]
= P.[alaemon] javanicus Heller, 1862b: 421; Plate 2, fg. 48. [Java, Indonesia; lectotype designation by
Wowor & Ng, 2010b]
= Palaemon (Parapalaemon) Trompii De Man, 1898: 144; Plate 7. [Ketungau river, Sintang Regency, W.
Kalimantan; lectotype designation by Wowor & Ng, 2010b]
= Palaemon (Parapalaemon) thienemanni Roux, 1932: 565, 570; Figs a-b. [Sg. Musi near Muara Klingi,
S. Sumatra; lectotype designation by Wowor & Ng, 2010b]
= Palmon (Parap.) trompi armatus Roux, 1936: 30. [Gunong Pulai, Johore]
Macrobrachium suongae Nguyn, 2003: 460; Figs 4-5. [Stream near An Phu village, 10 km south of Pleiku
city (altitude about 880 m), 1357'N 10802'E, centre of Vietnam]
Macrobrachium superbum (Heller, 1862a)
= Palaemon superbus Heller, 1862a: 528. [Schanghai]
= Palaemon superbus Heller, 1865: 118; Plate 10, fg. 10. [Schanghai]
Macrobrachium surinamicum Holthuis, 1948: 1112. [Plantation Geyersvlit near Paramaribo, Surinam]
Macrobrachium tenellum (Smith, 1871)
= Palmon tenellus Smith, 1871: 98. [Polvon, Occidental Department of Nicaragua]
= Palaemon longipes Lockington, 1878a: 161. [Mulege River, West Coast of Gulf of California]
Macrobrachium tenuirostrum X. Wang, 1997: 459; Fig. 1. [Tongren, Guizhou Province]
Macrobrachium thai Cai, Naiyanetr & Ng, 2004: 605; Figs 8-9. [Nong Khai, north-est Thailand]
Macrobrachium therezieni Holthuis, 1965b: 281; Fig. 1. [Fnrive district, Tamatave province, eastern Mad-
agascar]
Macrobrachium thuylami Nguyn, 2006b: 1; Figs 1-3. [local market in Duc Lap, a district in the highlands
of South Vietnam, at 1226'N 10739'E (about 300 km northeast of Ho Chi Minh City)]
Macrobrachium thysi Powell, 1980: 318; Figs 1-3. [Cte dIvoire, Banco fort]
Macrobrachium tiwarii Jalihal, Shenoy & Sankolli, 1988: 27; Figs 8-9. [Malaprabha River, near old bridge,
Khanapur]
Macrobrachium tolmerum Riek, 1951: 362; Fig. 1. [Black River, Macrossan, Queensland, Australia; consid-
ered erroneous by Short, 2004]
Macrobrachium totonacum Meja, Alvarez & Hartnoll, 2003: 79; Figs 2-3. [spring of the San Antonio River,
Oaxaca, Mexico,1828'8"N 9638'6"W, 90 m altitude]
Macrobrachium transandicum Holthuis, 1950b: 94. [Rio Telembi, near San Lorenzo, S.W. Colombia]
Macrobrachium tratense Cai, Naiyanetr & Ng, 2004: 595; Figs 3-4, 5A. [Khlong Fuai, road 3271, 1223'44.8''N
10239'34.5''E, Trat Province]
Macrobrachium trichodactylum Liang, Liu & Chen in Li, Liu, Liang & Chen, 2007: 136; Fig. 54. [Qiong-Hai
city, Hainan]
Macrobrachium tuxtlaense Villalobos & Alvarez, 1999: 746; Figs 1-3. [small stream 1 km S of Coyame,
1826'N 957'W, tributary of lake Catemaco, Municipio de Catemaco, Veracruz, Mxico]
Macrobrachium unikarnatakae Jalihal, Shenoy & Sankolli, 1988: 21; Figs 6-7. [Hosayellapur tank (Hirekeri),
Dharwad]
Macrobrachium urayang Wowor & Short, 2007: 78; Figs 1-2. [Sg. Enggeng Iut, a tributary of Sg. Bahau,
Kayan Basin, East Kalimantan]
Macrobrachium veliense Jayachandran & Joseph, 1985b: 185; Figs 1-2. [Veli Lake, 0830'-0832'N 7653'-
7655'E]
Macrobrachium venustum (Parisi, 1919)
= Palaemon (Eupalaemon) venustus Parisi, 1919: 82 (as P. vernustus); Plate 4, fg. 1; Plate 6, fgs 5-13 (as
P. venustus). [Isla di Hainan]
Macrobrachium vicconi Romn, Ortega & Meja, 2000: 186-192; Figs 1-12. [Ro Perlas, Municipio de
Ocosingo, state of Chiapas, Mxico, 1647'78N, 912929W]
Macrobrachium vietnamiense ng in ng & B.Y. Nguyn, 1972: 2; Fig. 2. [Ky Phu stream, Bac Thai
Province]
Macrobrachium villalobosi H.H.Jr. Hobbs, 1973a: 77; Fig. 3. [Cueva del Naciemento del Ro San Antonio,
10 km SSW Acatln, Oaxaca, Mxico]
Macrobrachium villosimanus (Tiwari, 1949b)
= Palaemon villosimanus Tiwari, 1949b: 329. [Calcuta, Pulta Waterworks]
Macrobrachium vollenhoveni (Herklots, 1857)
= Palaemon Vollenhovenii Herklots, 1857: 96. [cte de Guine]
= P.[alaemon] jamaicensis var. africanus Bouvier, 1895: 160. [les rivires de lAfrique occidentale]
= Palmon (Macrobrachium) jamaicensis var. angolensis De Man, 1904: 314; Plate 19, fgs 39-45, 47;
Plate 20, Figs 46, 48-53. [river at Catumbella, near Benguella, Angola]
= Palmon (Macrobrachium) jamaicensis var. Herklotsii De Man, 1912b: 239. [Mayumba, Bas-Congo]
Macrobrachium walvanense Almelkar, Jalihal & Sankolli, 1999: 328; Figs 1-19. [fsh rearing ponds of
Messrs. Tata Hydro-Electric Companys fsh farm, Walvan Dam, Lonavala (Pune district), Maha-
rashtra State, India, 540 m above mean sea level]
Macrobrachium wannanense Dai & Tan, 1993: 301; Figs 1-4. [Shitai County, 3011'N 11727'E, Anhui Province]
Macrobrachium weberi (De Man, 1892a)
= Palaemon (Eupalaemon) Weberi De Man, 1892a: 421; Plate 25, fgs 33-33h. [See von Tempe; See von
Sidenreng; aus dem Flusse Tjenara bei Pampanua (all from Celebes)]
Macrobrachium yui Holthuis, 1950a: 211. [nomen novum for Palaemon brevicarpus var. heterochirus Y, 1936]
= Palaemon brevicarpus var. heterochirus Y, 1936: 305; Figs 1-2; nec Palaemon heterochirus Wiegmann,
1836. [ning-Erh]
= Macrobrachium yeti ng, 1975: 67; Fig. 1. [streams in Son La Province]
Macrobrachuim zariquieyi Holthuis, 1949b: 178; Figs 1-2. [Fernando Poo, Rio Consul, 1 km from its mouth]
Nematopalaemon Holthuis, 1950a
= Nematopalaemon Holthuis, 1950a (type species Leander tenuipes Henderson, 1893, by original des-
ignation, gender masculine)
Nematopalaemon colombiensis (Squires & Mora L., 1971)
= Palaemon (Nematopalaemon) colombiensis Squires & Mora L., 1971: 102; Fig. 1. [of Pizarro, Colom-
bia, 509'N 7728'W, 9.1 m]
Nematopalaemon hastatus (Aurivillius, 1898a)
= Palaemon (Leander) hastatus Aurivillius, 1898a: 27; Plate 4, fgs 3-6. [Kamerun, im Meere bei Beticka
ba Mallale]
Nematopalaemon karnafuliensis (Ali Azam Khan, Fincham & Mahmood, 1980)
= Palaemon (Nematopalaemon) karnafuliensis Ali Azam Khan, Fincham & Mahmood, 1980: 85; Figs 1-2.
[Karnafuli Estuary, Chitagong, Bangladesh]
Nematopalaemon schmiti (Holthuis, 1950b)
= Palaemon schmiti Holthuis, 1950b: 97. [mouth of the Surinam River, near Resolutie, Dutch Guiana]
Nematopalaemon tenuipes (Henderson, 1893)
= Leander tenuipes Henderson, 1893: 440; Plate 40, Figs 14-15. [Bombay; Gulf of Martaban; Madras]
= Palmon luzonensis Blanco, 1939c: 201; Plate 1. [Aparri, Cagayan Province, Luzon]
Neopalaemon H.H.Jr. Hobbs, 1973b
= Neopalaemon H.H.Jr. Hobbs, 1973b (type species Neopalaemon nahuatlus H.H.Jr. Hobbs, 1973b, by
Neopalaemon nahuatlus H.H.Jr. Hobbs, 1973b: 26, Figs 1-2. [Cueva del Guano, 10 km NE Valle Nacional,
Oaxaca, Mxico]
Palaemon Weber, 1795
= Palaemon Weber, 1795 (type species Palaemon adspersus Rathke, 1837, by direction under the ple-
nary power of the ICZN, gender masculine; name placed on the Ofcial List of Generic Names in
= Palaemon Fabricius, 1798 (type species Cancer Squilla Linnaeus, 1758, designated by Latreille, 1810,
gender masculine; name placed on the Ofcial Index of Rejected and Invalid Names in Zoology in
= Palaeander Holthuis, 1950a (type species Palaemon elegans Rathke, 1837, by original designation,
gender masculine)
Palaemon adspersus Rathke, 1837
= Palaemon adspersus Rathke, 1837: 368; Plate 4, fg. 4. [an den vom schwarzen Meere besplten
Ksten der Krym]
= Cancer Squilla Linnaeus, 1758: 632. [Mare Balthico, Oceano Europo]
= Palaemon communis Anslin, 1826: 262. [nomen nudum]
= Palaemon Fabricii Rathke, 1843: 6. [Christiansund]
= Palaemon rectirostris Zaddach, 1844: 1. [maris baltici in litore Gedanensi]
= Palaemon Leachii Bell, 1851 [in Bell, 1844-1853]: 307; unnumbered text fgure. [Poole Harbour]
? = Palaemon imbellis Fischer, 1872: 422. [Bassin dArcachon, dans les Zostera (1-2 brasses)]
= P.[alaemon] rectirostris var. octodentata Neumann, 1878: 37. [Palam de Mallorka]
= Leander rectirostris var. transitans Czerniavsky, 1884: 41. [sinus Sevastopol, portu Streletzkaja ad
litor.]
? = Leander Brandti Czerniavsky, 1884: 43; Plate 3, fg. 8. [Sevastopol]
= Palaemon foliirostris Phan, 1971: 35; Figs 1-5. [Caspian Sea]
Palaemon afnis H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]
= P.[alemon] afnis H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 391. [Nouvelle-Z-
lande]
= P.[alemon] Quoianus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 393. [Nouvelle-
Zlande]
Palaemon capensis (De Man in Weber, 1897)
= Leander capensis De Man in Weber, 1897: 174; Plate 15, fgs 3-3g. [Cap-Colonie, im Knysna-Fluss, in
sssem Wasser]
Palaemon concinnus Dana, 1852a
= Palmon concinnus Dana, 1852a: 26. [archipelago Viti]
= Palmon exilimanus Dana, 1852a: 26. [archipelago Viti]
= Leander longicarpus Stimpson, 1860a: 40. [Portu Hong Kong Sinensi]
= Palmon lagdaoensis Blanco, 1939b: 167; Plate 1. [Cagayan River]
Palaemon curvirostris Nguyn, 1992: 34; Figs 7-8, 12 (part). [small fshery centre near Thanh An, a village
20 km north-west of Vung Tu ( = Cap St. Jacques)]
Palaemon debilis Dana, 1852a
= Palmon debilis Dana, 1852a: 26. [insulas Hawaiensibus]
= Palmon debilis var. Dana, 1852a: 26. [insulas Hawaiensibus]
= Palmon debilis var. , atenuatus Dana, 1852a: 26. [insulas Hawaiensibus]
= Leander gardineri Borradaile, 1901: 98. [fresh-water kuli in Ekasdu, Miladummadulu Atoll]
= Leander beauforti Roux, 1923: 18; Figs 1-2. [Cram: Kairatoe, eau saumtre]
= Palaemonetes pacifcus Gurney, 1939c: 145; Plates 5-6. [brackish canal at Charanka in the island of
Saipan, Marianne Islands]
Palaemon dolospinus Walker & Poore, 2003
= Palaemon dolospina Walker & Poore, 2003: 251; Figs 1E, 2F-J, 3C, 6-7. [Margate Beach, North West
Bay, Tasmania, in Zostera muelleri and Heterozostera tasmanica on sand, 0.2-1 m]
Palaemon elegans Rathke, 1837
= Palaemon elegans Rathke, 1837: 370; Plate 4, fg. 5. [bei Sudagh und am Cap Porthenion]
= Palmon minans Norman, 1861: 279; Plate 14, fgs 1-2. [Guernsey]
= Pal.[aemonella] gracilis Paulson, 1875: 117; Plate 17, fgs 6-6f. [Red Sea]
= Leander squilla var. prototypa Czerniavsky, 1884: 50. [sinus Sevatopol, in portu Streletzk, ad litor.]
= Leander squilla var. brevidigitata Czerniavsky, 1884: 51. [promontorium Santa, prope Theodosi-
am]
= Leander squilla var. intermedia De Man, 1915b: 127; Plate 10, fgs 10-1v. [Oosterschelde, near Ye-
rseke; Goesche Sas (Sluis van Goes), Oosterschelde; of Kats, Oosterschelde; Oosterschelde; Plym-
outh; Brixham, Tor Bay; Le Portel, Straits of Calais; Straits of Calais]
Palaemon foridanus Chace, 1942b: 80; Plate 23. [tide pool or sinkhole in back of beach at south end of
Captiva Island, Florida]
Palaemon gladiator Holthuis, 1950b: 96. [Academy Bay, Indefatigable Island, Galpagos Archipelago]
Palaemon gracilis (Smith, 1871)
= Leander gracilis Smith, 1871: 97. [Estero at Realejo, west coast of Nicaragua]
Palaemon gravieri (Yu, 1930b)
= Leander Gravieri Yu, 1930b: 555, 564; Fig. 3. [Tientsin, Tangkou]
Palaemon guangdongensis Liu, Liang & Yan, 1990a: 234; Fig. 27. [Guanghai, Taishan County, Guangdong]
Palaemon hancocki Holthuis, 1950b: 97. [Guayas River, Ecuador]
Palaemon intermedius (Stimpson, 1860a)
= Leander intermedius Stimpson, 1860a: 41. [in portu Jacksoniensi Australiae, prof. 2 org]
Palaemon khori De Grave & Al-Maslamani, 2006: 38; Figs 1-4. [creek on the northern side of Al-Khor, east
coast of Qatar, 5133'30''E 2541'30''N]
Palaemon litoreus (McCulloch, 1909)
= Leander litoreus McCulloch, 1909: 308; Fig. 16. [rock-pools on the coast near Sydney]
Palaemon longirostris H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]
= P.[alemon] longirostris H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 392. [lembouchure
de la Garonne, prs de Bordeaux]
= Astacus albescens Pennant, 1812: 25. [coasts of Kent]
= Palaemon Edwardsii Heller, 1863a: 265. [Mitelmeer, Corsika]
= Palaemon longipes Fischer, 1872: 421. [nomen nudum]
= Leander edwardsi var. brevidigitata forma similis Czerniavsky, 1884: 54. [sinus Sevastopol; promon-
torium Santa, prope Theodosiam]
= Leander edwardsi var. brevidigitata forma simplicior Czerniavsky, 1884: 55. [sinus Sevastopol; prom-
ontorium Santa]
= Leander edwardsi forma intermedia Czerniavsky, 1884: 55. [litus Tauriae]
= Leander edwardsi var. prototypa Czerniavsky, 1884: 55. [sinus Sevastopol]
= Leander longirostris var. robusta De Man, 1925b: 4. [mouth of the river Adour; Hollandsch Diep]
= Palaemon garciacidi Zariquiey lvarez, 1968: 167; Figs 69c, 70. [ro Gaudalquivir, Ilegando hasta
Sevilla; ro Francol, en Tarragona; ro Fluvi, en el Golfo de Rosas; Portugal]
Palaemon macrodactylus Rathbun, 1902b
= Palmon macrodactylus Rathbun, 1902b: 52; Fig. 24. [Aomori, Rikuoku]
= Leander serrifer var. longidactylus Yu, 1930b: 570 (partim); Fig. 4B'-C' (nec text fgures). [Yangmatoa,
Peitaiho, Tangkou, Chefoo]
Palaemon maculatus (Thallwitz, 1892)
= Leander maculatus Thallwitz, 1892: 19; Plate 1, fg. 4. [Ogow, West-Afrika]
Palaemon miyadii (Kubo, 1938b)
= Leander miyadii Kubo, 1938b: 538; Fig. 1A, C-F, M-P. [Liao-yang]
Palaemon northropi (Rankin, 1898)
= Leander northropi Rankin, 1898: 245; Plate 30, fg. 4. [Nassau, New Providence]
= Palmon brachylabis Rathbun, 1900: 154; Plate 8, fg. 10. [Rio Parahyba do Norte, on mangroves]
Palaemon ogasawaraensis Kato & Takeda, 1981: 101; Figs 2-5. [stream mouth, a stream fowing into Buta
beach (Minami-fukurosawa), Chichi-jimi Island, Ogasawara Islands]
Palaemon ortmanni Rathbun, 1902b
= Palmon ortmanni Rathbun, 1902b: 53. [nomen novum for Leander longipes Ortmann, 1890]
= Leander longipes Ortmann, 1890: 519; Plate 37, fg. 13; nec Olivier, 1811. [Japan, Sagamibai]
Palaemon pacifcus (Stimpson, 1860a)
= Leander pacifcus Stimpson, 1860a: 40. [ad insulas Hong Kong et Hawaii etiam in portu Si-
moda]
? = Leander okiensis Kamita, 1950: 216; Fig. 2. [innermost coast of Suwa Bay]
Palaemon paivai Fausto Filho, 1967
= Palaemon (Palaemon) paivai Fausto Filho, 1967: 19; Figs 1-12. [Praia de Meireles, Fortaleza, Cear,
Brasil]
Palaemon pandaliformis (Stimpson, 1871)
= Leander pandaliformis Stimpson, 1871: 130. [found by Prof. T. Gill in fresh-water streams near the
sea, either in Barbados or Trinidad, he is uncertain which]
? = Leander Potitinga Mller, 1892: 181. [nomen nudum]
= Palmonetes cubensis Hay, 1903: 433; Fig. 3. [Palacio, Cuba]
Palaemon paucidens De Haan, 1844 [in De Haan, 1833-1850]
= Palemon paucidens De Haan, 1844 [in De Haan, 1833-1850]: Plate 45, fg. 11 (1844); 170 (1849). [Ja-
pan; lectotype designation by Yamaguchi & Baba, 1993]
Palaemon peringueyi (Stebbing, 1915)
= Leander peringueyi Stebbing, 1915: 75; Plate 17. [3349'S 2556'E]
= Leander gilchristi Stebbing, 1915: 76; Plate 18. [East London wood]
Palaemon peruanus Holthuis, 1950b: 97. [La Palisada, near Tumbes, N. Peru]
Palaemon powelli Ashelby & De Grave, 2009: 33; Figs 3-8. [Oguck, km from entrance to Bonny River,
Niger Delta, Nigeria, 0439'40''N 0709'20''W]
Palaemon riteri Holmes, 1895
= Palmon Riteri Holmes, 1895: 579; Plate 21, fgs 29-35. [San Diego]
Palaemon semmelinkii (De Man, 1881)
= Leander semmelinkii De Man, 1881: 137. [on the road of Makassar, Celebes]
Palaemon serenus Heller, 1862a
= Leander serenus Heller, 1862a: 527. [Sidney; fully described in Heller, 1865]
Palaemon serratus (Pennant, 1777)
= Astacus Serratus Pennant, 1777: 15; Plate 16, fg. 28. [Type locality not indicated]
= M.[elicerta] Triliana Risso, 1816: 111; Plate 3, fg. 6. [environs de Nice, dans les moyennes profon-
deurs]
= Cancer captivus Nardo, 1847: 6. [golfo tra gli Asprei]
= Leander Latreillianus forma gigantea Czerniavsky, 1884: 44. [nomen novum for Melicerta Triliana
Risso, 1816]
= Leander Latreillianus forma typica Czerniavsky, 1884: 45. [sinus Sevastopol, portu Streletzkaja ad
lit.; sinus Novorossisk, 6-10 m]
= Leander Latreillianus forma transitans Czerniavsky, 1884: 46. [sinus Suchum, 1-1.5 m]
= Leander Latreillianus var. intermedia Czerniavsky, 1884: 47. [Sevastopol]
= Leander Latreillianus var. aberrans Czerniavsky, 1884: 47. [lacus salinus prope Sevastopol]
= Palaemon rostratus Gimenez, 1922: 141. [nomen nudum]
= Palemon Oratelli Monod, 1931: 133. [nomen nudum]
Palaemon serrifer (Stimpson, 1860a)
= Leander serrifer Stimpson, 1860a: 41. [Portu Hong Kong et sinibus insulae Ousima, litoralis]
= Leander Fagei Yu, 1930b: 555, 561; Fig. 2. [Pninsule de Shantong]
= Leander serrifer var. longidactylus Yu, 1930b: 570 (partim); text fgures (nec Fig. 4B-C). [Yangmatoa,
Peitaiho, Tangkou, Chefoo]
Palaemon sewelli (Kemp, 1925)
= Leander sewelli Kemp, 1925: 299; Figs 9-10. [of Betim Point, opposite Nova Goa, Portuguese India]
Palaemon tenuidactylus Liu, Liang & Yan, 1990a: 238; Fig. 31; Plate 1, Figs 2-4. [Changjiang (Yangtze)
River estuary]
Palaemon vicinus Ashelby, 2009: 826; Figs 1-40. [Boa Vista, W coast, NW coast of Ilhu de Sal rei, 1610'N
2258'E, intertidal (rockpool) and shallow sublitoral]
Palaemon xiphias Risso, 1816
= P.[alemon] Xiphias Risso, 1816: 102. [environs de Nice, dans les zostres du rivage]
? = P.[alemon] Trisetaceus Risso, 1816: 103. [environs de Nice, dans les fucus, loin du rivage]
= Palaemon crenulatus Risso, 1827: 60. [environs de Nice, rgion des algues]
= Palaemon Sogiontii Monod, 1931: 133. [nomen nudum]
Palaemon yamashitai Fujino & Miyake, 1970a
= Palaemon (Palaemon) yamashitai Fujino & Miyake, 1970a: 244; Figs 1-2. [3314.3'N 12216.9'E, 26 m]
Palaemonetes Heller, 1869
= Palaemonetes Heller, 1869 (type species Palaemon Varians Leach, 1813 [in Leach, 1813-1814], by
monotypy, gender masculine; name placed on the Ofcial List of Generic Names in Zoology in
= Palaemonopsis Stimpson, 1871 (type species Palaemonopsis carolinus Stimpson, 1871 (junior subjec-
tive synonym of P.[almon] vulgaris Say, 1818), designated by Holthuis, 1955b, gender feminine)
= Allocaris Sollaud, 1911b (type species Allocaris sinensis Sollaud, 1911b, by monotypy, gender femi-
nine)
= Alaocaris Holthuis, 1949c (type species Palaemonetes antrorum Benedict, 1896, by monotypy, gender
feminine)
Palaemonetes africanus Balss, 1916: 27; Fig. 9. [Nigeria, Old Calabar]
Palaemonetes antennarius (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840])
= P.[alemon] antennarius H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 391. [la mer
Adriatique]
= Palaemon lacustris von Martens, 1857: 183; Plate 10, Figs 1-9. [Albanersee]
= P.[elias] migratorius Heller, 1862b: 409; Plate 2, fg. 35. [aus dem Adriatischen Meere, aus dem
Gardasee und aus gypten]
= Palaemonetes varians var. thermajophilus Garbini, 1879: 187. [nomen nudum]
= Palaemon varians var. termaiophilus Garbini, 1881: 108. [nomen dubium]
= Palaemonetes varians var. macrogenitor Boas, 1889: 800; Figs (partim) 2 (N1-5), 3 (N1-5), 4 (N1-5);
Plate 23, fgs 2, 4, 6. [im sdlichen Europa, in den Mitelmeerlndern]
Palaemonetes antrorum Benedict, 1896
= Palmonetes antrorum Benedict, 1896: 615. [artesian well, San Marcos, Texas]
Palaemonetes argentinus Nobili, 1901a: 3. [Buenos-Ayres; Tigre, Rio de la Plata]
= Leander brasiliensis Ortmann, 1890: 524; Plate 37, fg. 16. [Rio Grande do Sul]
Palaemonetes atrinubes Bray, 1976: 76; Figs 23-42. [Canning Bridge, lower Swan River]
Palaemonetes australis Dakin, 1915: 572; Plate 1. [Upper Swan Bridge; neotype designation by Bray, 1976]
Palaemonetes camranhi Nguyn, 1997: 85; Fig. 1. [Ba Ngoi, Cam Ranh, a coastal district 50 km south of
Nha Trang, 1154'N 1909'E]
Palaemonetes carteri Gordon, 1935: 324; Fig. 12. [creek near Camp 1, Upper Cuyuni]
Palaemonetes cummingi Chace, 1954
= Palaemonetes (Palaemonetes) cummingi Chace, 1954: 319; Fig. 2. [Squirrel Chimney, Alachua County,
Florida]
Palaemonetes hiltoni Schmit, 1921: 36; Plate 12, fg. 5. [San Pedro, California]
Palaemonetes hobbsi Strenth, 1994: 292; Fig. 1. [Nacimiento del Ro Mante, 8 km southwest of Ciudad
Mante, Tamaulipas, Mxico]
Palaemonetes intermedius Holthuis, 1949c
= Palaemonetes (Palaemonetes) intermedius Holthuis, 1949c: 94; Fig. 2j-l. [Box Iron Bay, Chincoteague
Bay, Virginia; see Holthuis, 1952b]
Palaemonetes ivonicus Holthuis, 1950b: 98. [Ivon at the Beni River, N. Bolivia]
Palaemonetes kadiakensis Rathbun, 1902a
= Palmonetes kadiakensis Rathbun, 1902a: 903. [Kadiak Island, Alaska, under stones at low water]
Palaemonetes karukera Carvacho, 1979
= Palaemonetes (Palaemonetes) karukera Carvacho, 1979: 449; Figs 2-3. [Rivire Lzarde, 300 m de
lembouchure]
Palaemonetes lindsayi Villalobos Figueroa & H.H.Jr. Hobbs, 1974
= Palaemonetes (Palaemonetes) lindsayi Villalobos Figueroa & H.H.Jr. Hobbs, 1974: 9; Figs 4-7, 8e.
[springs and canals in the watershed of Laguna de la Media Luna, 4.8 miles south of Ro Verde (on
the orad to Pedro Montoya) and 2.5 miles to the west on a road to the Mina El Refugio, San Luis
Potos, Mexico]
Palaemonetes mercedae Pereira S., 1986
= Palaemonetes (Palaemonetes) mercedae Pereira S., 1986: 209; Fig. 13. [Atabapo River at Chamuchina,
Territorio Federal Amazonas, Venezuela, 320'N 6729'W]
Palaemonetes mesogenitor Sollaud, 1912
= Palmonetes varians mesogenitor Sollaud, 1912: 1270. [Gabs, dans loued qui arrose cet oasis]
= Palaemonetes punicus Sollaud, 1923: 589; Figs 23-24. [Tunisie]
Palaemonetes mesopotamicus Pesta, 1913
= Palaemonetes mesopotamicus Pesta, 1913: 23; Figs 6-7, 9. [Hsitsche (= Heseke) am Khabur]
Palaemonetes mexicanus Strenth, 1976: 7; Fig. 3. [small river 15 road miles (24 km) west of Ciudad Valles,
San Luis Potos, Mexico]
Palaemonetes octaviae Chace, 1972
= Palaemonetes (Palaemonetes) octaviae Chace, 1972: 22; Figs 3-4. [Guadeloupe, Pointe--Pitre, be-
tween let Monroux and let Rat, sandy mudfats]
Palaemonetes paludosus (Gibbes, 1850)
= Hippolyte caroliniana Gibbes, 1848: 16. [nomen nudum]
= Hippolyte paludosa Gibbes, 1850: 197. [in fresh water ponds, St. Andrews Parish, South Carolina]
= Palaemonetes exilipes Stimpson, 1871: 130. [freshwater at Somerville, South Carolina]
Palaemonetes pugio Holthuis, 1949c
= Palaemonetes (Palaemonetes) pugio Holthuis, 1949c: 95; Fig. 2m-o. [Lagoon near Cove Point Light,
Chesapeake Bay; see Holthuis, 1952b]
Palaemonetes schmiti Holthuis, 1950b: 99. [Upper Chamber, east side, Mirafores Locks, Panama Canal]
Palaemonetes sinensis (Sollaud, 1911b)
= Allocaris sinensis Sollaud, 1911b: 50; Figs 1-2. [prs de Pkin]
= Palaemonetes chankensis Buldovsky, 1933: 43; Plate 1; Plate 2, fgs 12-16, 18-19. [First river (Jerick) to
the south of the Village of Astrachanka, lake Khanka, Far East]
= Palaemonetes venephicus Birstein & Vinogradov, 1934: 45; Fig. 3. [in der Nhe von Daubiche (Bassin
des Flusses Ussuri)]
Palaemonetes sutkusi Smalley, 1964: 229; Fig. 1. [Rio Grande drainage, 24 miles north of Monclova,
Coahuila Province, Mexico]
Palaemonetes texanus Strenth, 1976: 5; Fig. 2. [San Marcos River, within the city limits of San Marcos,
Hays County, Texas]
Palaemonetes turcorum Holthuis, 1961c: 14; Figs 6-8. [Sakarya River near the bridge of the highway from An-
kara to Eskiehir, about halfway between Sivrihisar and Haymana, about 95 km S.W. of Ankara, Turkey]
Palaemonetes varians (Leach, 1813 [in Leach, 1813-1814])
= Palaemon Varians Leach, 1813 [in Leach, 1813-1814]: 401. [Devonshire and Glamorgan coasts]
= Palaemon variabilis Bouchard-Chanteraux, 1829: 15. [les fosss deau saumtre de Capcure]
= Palaemonetes varians var. microgenitor Boas, 1889: 800; Figs (partim) 2 (A1-5), 3 (A1-5), 4 (A1-5);
Plate 23, fgs 1, 3, 5. [nrdliches Europa]
= Palaemonetes varians occidentalis Sollaud, 1923: 547; Figs 12-13; Plate 18, fgs 5-7. [eaux saumtres le
long des ctes occidentales de lEurope et de lAfrique]
Palaemonetes vulgaris (Say, 1818)
= P.[almon] vulgaris Say, 1818: 248. [bays and estuaries]
= Palaemonetes carolinus Stimpson, 1871: 129. [harbor of Charleston, South Carolina; Great Egg Har-
bor, New Jersey; harbor of Beaufort, North Carolina]
Palaemonetes zariquieyi Sollaud, 1938
= Palmonetes Zariquieyi Sollaud, 1939: 638; Fig. 1c-d. [lAlbufera (lac) de Valence; les lagunes de la
Almenara, les canaux dAmposta (delta de lEbre), une petit tang litoral au cap Salou (prs de
Tarragone)]
Pseudopalaemon Sollaud, 1911c
= Pseudopalaemon Sollaud, 1911c (type species Pseudopalaemon Bouvieri Sollaud, 1911c, by monotypy,
1957)
Pseudopalaemon amazonensis Ramos-Porto, 1979: 693. [arredores de Manaus, Amazonas]
Pseudopalaemon bouvieri Sollaud, 1911c
= Pseudopalmon Bouvieri Sollaud, 1911c: 12; Figs 1, 2a. [environs de Montvido]
= Pseudopalaemon Iheringi Sollaud, 1911d: 285; Fig. 1. [Arroyo del Bellaco, Brsil]
Pseudopalaemon chryseus Kensley & Walker, 1982: 16; Figs 18-19. [Rio Tarumazinho]
Pseudopalaemon funchiae Garca-Dvila & Magalhes, 2004: 677; Figs 28-38. [Loreto, Provncia de Re-
quena: distrito Jenaro Herrera, Jenaro Herrera, cocha Carahuaite, afuente da cocha Supay, bacia do
rio Ucayali]
Pseudopalaemon gouldingi Kensley & Walker, 1982: 18; Figs 20-21. [Rio Negro, Ilha de Cumuru, Lago da
Ilha]
Pseudopalaemon iquitoensis Garca-Dvila & Magalhes, 2004: 679; Figs 39-50. [Provncia de Maynas,
distrito de Iquitos, Iquitos, quebrada Paujil, km 22 da Estrada Iquitos-Nauta, afuente do rio
Itaya]
Pseudopalaemon nigramnis Kensley & Walker, 1982: 20; Figs 22-23. [Rio Marania, Rio Negro basin]
Rhopalaemon Ashelby & De Grave, 2010
= Rhopalaemon Ashelby & De Grave, 2010 (type species Leander belindae Kemp, 1925, by original
Rhopalaemon belindae (Kemp, 1925)
= Leander belindae Kemp, 1925: 308; Figs 12-13. [Kilakarai, Gulf of Manaar]
Tenuipedium Wowor & Ng, 2010a
= Tenuipedium Wowor & Ng, 2010a (type species Macrobrachium palaemonoides Holthuis, 1950a, by
original designation and monotypy, gender neuter)
Tenuipedium palaemonoides (Holthuis, 1950a)
= Macrobrachium palaemonoides Holthuis, 1950a: 136; Fig. 31. [Laut Tawar, Laulo Lake, N. Simalur, of
west coast of Sumatra]
Troglindicus Sankolli & Shenoy, 1979
= Troglindicus Sankolli & Shenoy, 1979 (type species Troglindicus phreaticus Sankolli & Shenoy, 1979,
Troglindicus phreaticus Sankolli & Shenoy, 1979: 86; Figs 1-2. [northernmost old freshwater well (13
th

century?) in the fort area of Ratnagiri, west coast of India]
Troglocubanus Holthuis, 1949c
= Troglocubanus Holthuis, 1949c (type species Palmonetes eigenmanni Hay, 1903, by original desig-
Troglocubanus calcis (Rathbun, 1912b)
= Palaemonetes calcis Rathbun, 1912b: 451; Plate 1, fgs 1-5. [Cuba, pool in a cave between Madruga
and Aguacate]
Troglocubanus eigenmanni (Hay, 1903)
= Palmonetes eigenmanni Hay, 1903: 431; Fig. 2. [cavern at Ashton, Cuba]
Troglocubanus gibarensis (Chace, 1943)
= Palaemonetes gibarensis Chace, 1943: 28; Plate 7. [a well, 29 yards deep, entering an underground
stream, Aguada del Montaes, en el Jobal, Barrio de Cupeysillo, Termino de Gibara, Oriente prov-
ince, Cuba]
Troglocubanus inermis (Chace, 1943)
= Palaemonetes inermis Chace, 1943: 26; Plate 6. [pool in a cave situated on the calzada from Madruga
to Aguacate, about one kilometer from the junction with the calzada from Madruga to Matanzas,
Havana Province, Cuba]
Troglocubanus jamaicensis Holthuis, 1963a: 67; Fig. 3. [stream in limestone cave near Lucky Hill Coopera-
tive farm, near Goshen, Jamaica]
Troglomexicanus Villalobos, Alvarez & Ilife, 1999
= Troglomexicanus Villalobos, Alvarez & Ilife, 1999 (type species Troglocubanus perezfarfantae Villalo-
bos Figueroa, 1971, by original designation, gender masculine)
Troglomexicanus huastecae Villalobos, Alvarez & Ilife, 1999: 117; Figs 4-5. [Manantial de San Rafael de los
Castros, 2246'N 9903'W, Municipio de Ciudad Mante, San Luis Potos]
Troglomexicanus perezfarfanteae (Villalobos Figueroa, 1971)
= Troglocubanus perezfarfanteae Villalobos Figueroa, 1971: 1; Figs 1-23, 24 (partim). [Stano de Tinaja,
11.7 km E. NE. Valles, San Luis Potos, Mxico]
Troglomexicanus tamaulipasensis Villalobos, Alvarez & Ilife, 1999: 113; Figs 2-3. [Cueva del Nacimiento
del Ro Fro, 2301'N 9904'W, Municipio de Gmez Fars, Tamaulipas]
Urocaridella Borradaile, 1915b
= Urocaridella Borradaile, 1915b (type species Urocaridella gracilis Borradaile, 1915b (invalid senior
synonym of Urocaridella urocaridella (Holthuis, 1950a), by monotypy, gender feminine)
Urocaridella antonbruunii (Bruce, 1967b)
= Periclimenes antonbruunii Bruce, 1967b: 45; Figs 19-22. [Pamanzi Island reef, Dzaoudzi, Ile Mayote,
Archipel des Comores]
= Leandrites cyrtorhynchus Fujino & Miyake, 1969a: 143; Figs 1-3. [T-shima Island, Tanabe Bay, Wa-
kayama Prefecture]
= Leandrites longipes Liu, Liang & Yan, 1990b: 127; Fig. 24. [Sanya, Hainan Isl., China; additional
fgure in Liu, Liang & Yan, 1990a]
Urocaridella pulchella Yokes & Galil, 2006: 749; Figs 2A-B, 3. [Turkey, Gvercin Ada, Ka, 12 m]
Urocaridella urocaridella (Holthuis, 1950a)
= Leander urocaridella Holthuis, 1950a: 28. [nomen novum for Urocaridella gracilis Borradaile, 1915b]
= Urocaridella gracilis Borradaile, 1915b: 210. [Maldive Islands]
Urocaridella vestigialis Chace & Bruce, 1993: 45; Fig. 17. [Indonesia, Selat Butung, Celebes, 431'40"S
12249'42"E, 68 m]
Subfamily PONTONIINAE Kingsley, 1879
Allopontonia Bruce, 1972b
= Allopontonia Bruce, 1972b (type species Allopontonia iani Bruce, 1972b, by original designation and
monotypy, gender feminine; a junior synonym of Anchistia Brockii De Man, 1888b)
Allopontonia brockii (De Man, 1888b)
= Anchistia Brockii De Man, 1888b: 548; Plate 22a, fgs 3-3d. [Amboina]
? = Periclimenes (Periclimenes) signatus Kemp, 1925: 322; Figs 16-17. [Andamans]
= Allopontonia iani Bruce, 1972b: 7; Figs 1-4. [Zanzibar Harbour, 609.5'S 3910.2'E]
= Periclimenes priodactylus Bruce, 1992c: 53; Figs 1, 7-11. [Lizard island, Queensland, 1440'S 14528'E,
northern entrance to Blue Lagoon, 3.5 m]
Allopontonia alastairi Bruce, 2010a: 33; Fig. 1. [North-western Australia, Imperieuse L23 transect,
1825'31''S 1200555''E to 1826'12''S 12006'30''E, 105-103 m]
Altopontonia Bruce, 1990c
= Altopontonia Bruce, 1990c (type species Altopontonia disparostris Bruce, 1990c, by original designa-
Altopontonia disparostris Bruce, 1990c: 192; Figs 25-33, 39k. [New Caledonia, 2303'S 16719'E, 503 m]
Amphipontonia Bruce, 1991a
= Amphipontonia Bruce, 1991a (type species Amphipontonia kanak Bruce, 1991a, by original designa-
Amphipontonia kanak Bruce, 1991a: 382; Figs 58-63. [Loyalty Islands]
Anapontonia Bruce, 1966c
= Anapontonia Bruce, 1966c (type species Anapontonia denticauda Bruce, 1966c, by monotypy, gender
feminine)
= Anapontonia Bruce, 1967b (type species Anapontonia denticauda Bruce, 1967b, by monotypy, gender
feminine)
Anapontonia denticauda Bruce, 1966c: 596. [Pange Reef, Zanzibar; fully described in Bruce, 1967b]
Anchiopontonia Bruce, 1992d
= Anchiopontonia Bruce, 1992d (type species Pontonia hurii Holthuis, 1981, by original designation
Anchiopontonia hurii (Holthuis, 1981)
= Pontonia hurii Holthuis, 1981: 796; Fig. 4. [Arno Atoll, Marshall Islands, from mantle cavity of rock
clam, Spondylus spec.]
= Pontonia hurii Holthuis, 1953b: 57. [nomen nudum]
Anchistus Borradaile, 1898
= Anchistus Borradaile, 1898 (type species Harpilius Miersi De Man, 1888a, by original designation,
gender masculine; name placed on the Ofcial List of Generic Names in Zoology in Opinion 470
in 1957)
= Tridacnocaris Nobili, 1899 (nomen novum for Anchistus Borradaile, 1898, gender feminine)
= Marygrande Pesta, 1911 (type species Marygrande mirabilis Pesta, 1911, by monotypy, gender
feminine)
= Ensiger Borradaile, 1915b (type species Anchistia aurantiaca Dana, 1852a (junior subjective syno-
nym of Cancer custos Forskl, 1775), being the frst species subsequently placed in this subgenus,
which was described without any nominal species, frst placed in Ensiger by Borradaile, 1917)
Anchistus australis Bruce, 1977a
= Anchistus australis forma typica Bruce, 1977a: 56; Figs 7-9. [Capre Cay, Swains Reefs; unavailable
under Art. 15.2]
= Anchistus australis forma dendricauda Bruce, 1977a: 62; Fig. 10. [West Cay, Diamond Islets; unavail-
able under Art. 15.2]
Anchistus custoides Bruce, 1977a: 50; Figs 4-6. [north-west end, Gillet Cay, Great Barrier Reef, 2143'S
15225'E, in bivalve mollusk Atrina vexillum according to Springthorpe & Lowry, 1994]
Anchistus custos (Forskl, 1775)
= Cancer custos Forskl, 1775: 94. [Lohaj, intra Pinna nigras]
= Pontonia infata H. Milne Edwards, 1840 [in H. Milne Edwards, 1834-1840]: 633. [latinization of the
French vernacular name Pontonie enfe H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]:
360. [Ceylan et sur les ctes de Vanicoso]
= Anchistia aurantiaca Dana, 1852a: 25. [archipelago Viti]
= Harpilius inermis Miers, 1884b: 291; Plate 32, fg. B. [Port Molle, in the interior of a shell of a species
of Pinna]
= Pontonia pinnae Ortmann, 1894: 16; Plate 1, Figs 3-3n; nec Lockington, 1878a. [Dar-es-Salaam,
Chokirbank, in den Schalen einer Pinna-Art]
= Marygrande mirabilis Pesta, 1911: 573; Figs 1-5 (partim). [Samoa]
= Pontonia spinax Dawydof, 1952: 136. [nomen nudum]
Anchistus demani Kemp, 1922: 256; Figs 86-88. [Aberdeen, Port Blair, Andamans, low tide]
Anchistus gravieri Kemp, 1922: 252; Figs 82-84. [Vanikoro, Santa Cruz Islands, Polynesia]
Anchistus miersi (De Man, 1888a)
= Harpilius Miersi De Man, 1888a: 274; Plate 17, fgs 6-10. [Elphinstone Island]
= Marygrande mirabilis Pesta, 1911: 571, Figs 1-5 (partim). [Samoa]
Anchistus pectinis Kemp, 1925: 327; Figs 19-20. [Octavia Bay, Nancowry Harbor, Nicobars]
= Anchistus misakiensis Yokoya, 1936: 136; Fig. 5. [vicinity of the Misaki Marine Biological Station,
inside of a bivalve, Amusium japonicum]
Ancylomenes Okuno & Bruce, 2010
= Ancylomenes Okuno & Bruce, 2010 (type species Periclimenes venustus Bruce, 1989a, by original
Ancylomenes adularans (Bruce, 2003a)
= Periclimenes adularans Bruce, 2003a: 116; Fig. 2. [Cape Flatery Service Jety, 2 m]
Ancylomenes aesopius (Spence Bate, 1863)
= Anchistia sopia Spence Bate, 1863: 502; Plate 41, Figs 5-5z. [Gulf of St. Vincent]
Ancylomenes amirantei (Bruce, 2007a)
= Periclimenes amirantei Bruce, 2007a: 52; Figs 1-4. [Amirante Islands, Seychelle Islands, station E.15,
70 m]
Ancylomenes aqabai (Bruce, 2008a)
= Periclimenes aqabai Bruce, 2008a: 27; Figs 1-4. [Aqaba, Jordan, 35-45 m]
Ancylomenes grandidens (Bruce, 2005b)
= Periclimenes grandidens Bruce, 2005b: 350; Figs 8-10, 22C. [Loloata Island, 16 m]
Ancylomenes holthuisi (Bruce, 1969a)
= Periclimenes holthuisi Bruce, 1969a: 258. [Lung Ha Wan, N.T., Hong Kong, 2218.5'N 11418.2'E,
2 fms] (Fig. 37)
Ancylomenes kobayashii (Okuno & Nomura, 2002)
= Periclimenes kobayashii Okuno & Nomura, 2002: 84; Figs 1-5. [of Akazawa, Ito, Izu Peninsula, Hon-
shu, Japan, 3451.2'N 13905.5'E, 32 m]
Ancylomenes kuboi Bruce, 2010b: 170; Figs 1-2. [Northwest Shelf, Western Australia, 1905.3'S 11853.8'E,
80 m]
Ancylomenes longicarpus (Bruce & Svoboda, 1983)
= Periclimenes longicarpus Bruce & Svoboda, 1983: 13; Figs 4-8. [Aqaba, Jordan, 15 m]
Ancylomenes lucasi Chace, 1937b
= Periclimenes (Ancylocaris) lucasi Chace, 1937b: 133; Fig. 8. [San Lucas Bay, 2253'N 10954'W, 3-9 fms]
Fig. 37. Ancylomenes holthuisi (Bruce, 1969). Photo by Charles H.J.M. Fransen.
Ancylomenes luteomaculatus Okuno & Bruce, 2010: 88; Figs 1-5, 6A-B. [of Gima Harbor, Kume-jima Is-
land, Ruykyu Islands, Japan, 2619.7'N 12645.6'E, 32 m, associated with Pachycerianthus sp.]
Ancylomenes magnifcus (Bruce, 1979a)
= Periclimenes magnifcus Bruce, 1979a: 195; Figs 1-5; Plate 1A-C. [Wistari Reef, Heron Island, Capri-
corn Islands, Queensland, Australia, 85-95 feet]
Ancylomenes okunoi Bruce, 2010b: 172; Fig. 3. [1905.0'S 11850.5'E, Northwest Shelf, Western Australia,
83 m]
Ancylomenes pedersoni (Chace, 1958)
= Periclimenes (Periclimenes) pedersoni Chace, 1958: 125; Figs 1-17. [Lyford Cay, New Providence Is-
land, Bahamas]
= Periclimenes (Periclimenes) anthophilus Holthuis & Eibl-Eibesfeldt, 1964: 185; Figs 1-4. [Whalebone
Bay, Bermuda, near the shore, 2-3 m, on sea anemones; considered valid species by Okuno & Bruce,
2010]
Ancylomenes sarasvati (Okuno, 2002)
= Periclimenes sarasvati Okuno, 2002: 212; Figs 1-4, 6A-B. [of Hateno-hama, Kume-jima Island,
2625.9'N 12746.5'E, 26 m]
Ancylomenes speciosus (Okuno, 2004)
= Periclimenes speciosus Okuno, 2004: 866; Figs 1-5, 6A-B. [Hira-ne, Hasama, Tateyama, Boso Penin-
sula, Honshu, Japan, 3458.6'N 13947.1'E, 18 m]
Ancylomenes tenuirostris (Bruce, 1991b)
= Periclimenes tenuirostris Bruce, 1991b: 247; Figs 13-16. [New Caledonia, 2235.1'S 16659.5'E, Grand
Rcif Sud, 82 m]
Ancylomenes tosaensis (Kubo, 1951)
= Periclimenes (Ancylocaris) tosaensis Kubo, 1951: 268; Figs 7-8. [of Usa, Takaoka-gun, Kchi Pre-
fecture]
Ancylomenes venustus (Bruce, 1989a)
= Periclimenes venustus Bruce, 1989a: 178. [Nailon]
= Periclimenes venustus Bruce, 1990d: 230; Figs 1-6. [Table Head, Port Essington, 3 m]
Anisomenaeus Bruce, 2010c
= Anisomenaeus Bruce, 2010c (type species Periclimenaeus spinimanus Bruce, 1969b, gender mascu-
line)
Anisomenaeus spinimanus (Bruce, 1969b)
= Periclimenaeus spinimanus Bruce, 1969b: 165. [North-western Indian Ocean, of Ras Asir, 1137'N
5127'E 1138'N 5127'E, 37-40 fms]
Apopontonia Bruce, 1976c
= Apopontonia Bruce, 1976c (type species Apopontonia falcirostris Bruce, 1976c, by original designa-
Apopontonia falcirostris Bruce, 1976c: 303; Figs 1-5. [NW coast of Madagascar, 1244.5'S 4825.2'E, 73 m]
Araiopontonia Fujino & Miyake, 1970b
= Araiopontonia Fujino & Miyake, 1970b (type species Araiopontonia odontorhyncha Fujino & Miyake,
1970b, by original designation and monotypy, gender feminine)
Araiopontonia odontorhyncha Fujino & Miyake, 1970b: 2; Figs 1-4. [Koniya, Amami O Shima Island, Ryukyu
Islands, Japan]
Ascidonia Fransen, 2002
= Ascidonia Fransen, 2002 (type species P.[ontonia] favomaculata Heller, 1864, by original designa-
Ascidonia californiensis (Rathbun, 1902a)
= Pontonia californiensis Rathbun, 1902a: 902. [Of Santa Cruz Island, California, 30 fms]
Ascidonia favomaculata (Heller, 1864)
= P.[ontonia] favomaculata Heller, 1864: 51. [aus dem Adriatischen Meere, in der Mantelhhle von
Phallusia mamillata]
= Alciope heterochelus Rafnesque, 1814: 24. [Sicile; suppressed under the plenary power or the purposes
of the Principle of Priority but not for those of the Principle of Homonymy in Opinion 522 in 1958]
= Pontonia Phallusiae Marion in de Folin & Prier, 1879: 226. [nomen nudum; les eaux de Marseille]
= Pontonia Diazona Joliet, 1882: 118 (as Pontonia Diazon on page 120). [Menton]
= Pontonia elegans Sarato, 1887: 2. [les eaux de Saint-Jean, prs de Nice] [nomen nudum]
Ascidonia miserabilis (Holthuis, 1951b)
= Pontonia miserabilis Holthuis, 1951b: 148; Plate 47, fgs d-i. [of Vieques Island, Porto Rico, Culebritas
light house, NE 3/8 E 7 miles, 16 fms]
= Pontonia grayi Rathbun, 1901: 122 (partim; nec fg. 25). [between San Antonio Bridge and San
Geronimo, San Juan, Puerto Rico]
Ascidonia pusilla (Holthuis, 1951b)
= Pontonia pusilla Holthuis, 1951b: 142; Plate 45. [Ecuador, Salango Island, 3 fms]
= Pontonia spighti Fujino, 1972: 293; Figs 1-3. [Playas del Coco, Pacifc coast of Costa Rica, shallow
water of sublitoral zone]
Ascidonia quasipusilla (Chace, 1972)
= Pontonia quasipusilla Chace, 1972: 41; Fig. 10. [Charlote Point, English Harbour, Antigua Island]
Balssia Kemp, 1922
= Balssia Kemp, 1922 (type species Amphipalaemon Gasti Balss, 1921b, by monotypy, gender femi-
Balssia antipodarum Bruce, 2004a: 367; Figs 1-3. [North Norfolk Ridge, north of Norfolk Island, 2854.39'S
16741.05'E, 111-115 m]
Balssia gasti (Balss, 1921b)
= Amphipalaemon Gasti Balss, 1921b: 524; Figs 1-8. [Golf von Neapel, von Corallium rubrum]
? = Brachycarpus Antonini Magri, 1926 [lapsus for Brachycarpus Audouini but considered a synonym
as here listed by L.B. Holthuis (pers. comm.)]
Balssia noeli Bruce, 1998b: 604; Figs 1-4. [Mediterranean Sea, vicinity of Banyuls]
Blepharocaris Mitsuhashi & Chan, 2007
= Blepharocaris Mitsuhashi & Chan, 2007 (type species Blepharocaris panglao Mitsuhashi & Chan,
2007, by original designation and monotypy, gender feminine)
Blepharocaris panglao Mitsuhashi & Chan, 2007: 2; Figs 1-4. [937.6'N 12347.3'E, Bingag, Panglao Island,
Philippines, 20 m]
Brucecaris Marin & Chan, 2006
= Brucecaris Marin & Chan, 2006 (type species Periclimenes tenuis Bruce, 1969a, by original designa-
Brucecaris tenuis (Bruce, 1969a)
= Periclimenes tenuis Bruce, 1969a: 272. [Chukwani, Zanzibar Island, 615.1'S 3912.7'E, 1f]
Bruceonia Fransen, 2002
= Bruceonia Fransen, 2002 (type species Pontonia ardeae Bruce, 1981a, by original designation and
Bruceonia ardeae (Bruce, 1981a)
= Pontonia ardeae Bruce, 1981a: 113; Figs 1-8. [Heron Island, Wistari Reef, 2327.5'S 15155.0'E, 60-70
feet]
Cainonia Bruce, 2005b
= Cainonia Bruce, 2005b (type species Pontonia medipacifca Edmondson, 1935a, by original designa-
Cainonia medipacifca (Edmondson, 1935a)
= Pontonia medipacifca Edmondson, 1935a: 6; Fig. 2. [Midway Island, in shallow water]
Carinopontonia Bruce, 1988a
= Carinopontonia Bruce, 1988a (type species Carinopontonia paucipes Bruce, 1988a, by original desig-
Carinopontonia paucipes Bruce, 1988a: 1264; Figs 1-3. [1904.4'S 11847.55'E, 83 m]
Chacella Bruce, 1986b
= Chacella Bruce, 1986b (type species Dasycaris kerstitchi Wicksten, 1983a, by original designation
Chacella kerstitchi (Wicksten, 1983a)
= Dasycaris kerstitchi Wicksten, 1983a: 16; Fig. 2. [Punta Doble, San Carlos, Sonora, Mexico, 2756'N
11102'W, 30 m]
Chernocaris Johnson, 1967
= Chernocaris Johnson, 1967 (type species Chernocaris placunae Johnson, 1967, by monotypy, gender
feminine)
Chernocaris placunae Johnson, 1967: 500; Figs 1-12. [Telok Paku, Singapore, in Placuna sella taken at low
tide]
Climeniperaeus Bruce, 1995
= Climeniperaeus Bruce, 1995 (type species Periclimenaeus truncoides Chace & Bruce, 1993, by original
= Climeniperaeus Bruce, 1996 (type species Periclimenaeus truncoides Chace & Bruce, 1993, by original
Climeniperaeus orbitospinatus (Bruce, 1969b)
= Periclimenaeus orbitospinatus Bruce, 1969b: 160. [Gulf of Carpentaria, northern Australia, 10-15
fms]
= Apopontonia tridentata Bruce, 1988a: 1270; Figs 4-7. [1941.9'S 1757.15'E, 54 m]
Climeniperaeus truncatus (Rathbun, 1906)
= Coralliocaris truncata Rathbun, 1906: 920; Fig. 70; Plate 24, fg. 2. [South coast of Molokai, 23-24 fms]
Climeniperaeus truncoideus (Chace & Bruce, 1993)
= Periclimenaeus truncoideus Chace & Bruce, 1993: 93. [2.3 miles [3.7 km] N, 63W from north point
of Kai Besar, Kepulauan Kai, Indonesia, 536'5'S 13255.2'E, 90 m]
Colemonia Bruce, 2005b
= Colemonia Bruce, 2005b (type species Colemonia litodactylus Bruce, 2005b, by original designation
Colemonia litodactylus Bruce, 2005b: 335; Figs 1-4, 22A. [Milne Bay, 12 m]
Conchodytes Peters, 1852
= Conchodytes Peters, 1852 (type species Conchodytes tridacnae Peters, 1852, designated by Hilgen-
dorf, 1879, gender masculine; name placed on the Ofcial List of Generic Names in Zoology in
Conchodytes biunguiculatus (Paulson, 1875)
= P.[ontonia] biunguiculata Paul'son, 1875: 111; Plate 15, fg. 1-1n. [Red Sea]
= Conchodytes kempi Bruce, 1989a: 183; Fig. 3B-E. [Moalboal, in Isognomon isognomon]
Conchodytes maculatus Bruce, 1989b: 182; Figs 1-6. [Pao Yuan No. 1, 1634.5'S 12127.0'E, 40 m]
Conchodytes meleagrinae Peters, 1852: 594. [Sdostkste Africas, Ibo, im 12S Br., zwischen den Mantel-
lapen von Melaegrina margaritifera]
Conchodytes monodactylus Holthuis, 1952c: 200; Figs 96-98. [Takao, Southern Taiwan (in bivalve mollusk,
Pinna); Lesser Sunda Island]
Conchodytes nipponensis (De Haan, 1844 [in De Haan, 1833-1850])
= Hymenocera niponensis De Haan, 1844 [in De Haan, 1833-1850]: Plate 46, fg. 8 (1844); 180 (1849, as
Pontonia nipponensis). [Japan; incorrect original spelling corrected in Opinion 383 in 1956]
Conchodytes philippinensis Bruce, 1996: 212; Figs 6, 29b. [Philippines, few meters]
Conchodytes pteriae Fransen, 1994: 97; Figs 38-58. [N of dArros Island, 524'S 5319'E, 45-55 m, in small
Pteria loveni (Dunker, 1872)]
Conchodytes tridacnae Peters, 1852: 594. [Sdostkste Africas, bei Ibo, im 12 S Br., zwischen den Mantel-
lappen von Tridacna squamosa]
Coralliocaris Stimpson, 1860a
= Oedipus Dana, 1852a (type species Oedipus superbus Dana, 1852a, designated by Kingsley, 1880,
gender masculine; invalid junior homonym of Oedipus Berthold, 1827 (Orthoptera), Oedipus Tschudi,
1838 (Amphibia) and Oedipus Lesson, 180 (Mammalia); name placed on the Ofcial Index of Rejected
and Invalid Generic Names in Zoology in Opinion 470 in 1957)
= Coralliocaris Stimpson, 1860a (nomen novum for Oedipus Dana, 1852a, gender feminine; name
Coralliocaris brevirostris Borradaile, 1898: 386. [Ellice Islands]
Coralliocaris graminea (Dana, 1852a)
= dipus gramineus Dana, 1852a: 25. [archipelago Viti]
= Coralliocaris inaequalis Ortmann, 1890: 510; Plate 36, fg. 21. [Japan, Kagoshima and Samoa]
Coralliocaris labyrintha Mitsuhashi & Takeda, 2008: 16; Figs 1E-F, 6E-F, 11-14. [Urasoko Bay, Ishigaki-jima
Island, Rykyu Islands, coral reef, 2 m, from Acropora sp.]
Coralliocaris macrophthalma (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840])
= P.[ontonia] macrophthalma H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 359. [les
mers dAsie]
Coralliocaris nudirostris (Heller, 1861)
= O.[edipus] nudirostris Heller, 1861: 27. [im rothen Meere]
= O.[edipus] nudirostris Heller, 1862c: 279; Plate 3, fg. 25. [im rothen Meere]
= Coralliocaris tahitoei Boone, 1935: 180; Fig. 12; Plate 49. [Venus Point reef, Tahiti, Society Islands]
Coralliocaris sandyi Mitsuhashi & Takeda, 2008: 11; Figs 1C-D, 6C-D, 8-10. [of Kuro-shima Island, Rykyu
Islands, 8 m, from Acropora sp.]
Fig. 38. Coralliocaris viridis Bruce, 1974. Photo by Arthur Anker.
Coralliocaris superba (Dana, 1852a)
= dipus superbus Dana, 1852a: 25. [insula Tongatabu]
= Oed.[ipus] dentirostris Paulson, 1875: 112; Plate 14, Figs 7-7d. [Red Sea]
Coralliocaris taiwanensis Fujino & Miyake, 1972: 92; Figs 1-3. [Herngchuen, Shiangtiau Bay, southern
Taiwan, 2-5 m]
= Coralliocaris pavonae Bruce, 1972c: 77; Figs 8-11. [fringing reef at Singatoka, Viti Levu, Fii]
Coralliocaris tridens Mitsuhashi, Fujino & Takeda, 2001: 944; Figs 1-3. [Kabira Bay, 2426'N 1248'E, Ishi-
gaki Island, Ryukyu Islands, southwestern Japan, from tabular coral (Acropora sp)]
Coralliocaris venusta Kemp, 1922: 274; Figs 100-101. [N.E. Tholayiram Paar, Gulf of Manaar]
Coralliocaris viridis Bruce, 1974a: 222; Fig. 1A-B. [seaward reefs of Mombasa Island, 404.5'S 3940.5'E]
(Fig. 38)
Coutierea Nobili, 1901b
= Coutirea Nobili, 1901b (type species Coralliocaris Agassizi Coutire, 1901, by monotypy, gender
feminine; name placed in the corrected spelling (Coutierea) on the Ofcial List of Generic Names in
Coutierea agassizi (Coutire, 1901)
= Coralliocaris Agassizi Coutire, 1901: 115, unnumbered text fg. [Barbades, 94 brasses]
Crinotonia Marin, 2006
= Crinotonia Marin, 2006 (type species Crinotonia anastasiae Marin, 2006, by original designation,
gender feminine)
Crinotonia anastasiae Marin, 2006: 322; Figs 1-6. [South China Sea, Vietnam, Nha Thrang Bay, Mun Island,
10-12 m]
Crinotonia atenuatus (Bruce, 1971a)
= Periclimenes atenuatus Bruce, 1971a: 533; Figs 1-5. [Waterhouse Cove, Burukuk, Duke of York Is-
land, 47.3'S 15227.3'E, 1-2 m]
Ctenopontonia Bruce, 1979b
= Ctenopontonia Bruce, 1979b (type species Ctenopontonia cyphastreophila Bruce, 1979b, by original
Ctenopontonia cyphastreophila Bruce, 1979b: 425; Figs 1-6. [Enewetak Atoll, Marshall Islands, 5-15 m]
Cuapetes Clark, 1919
= Cuapetes Clark, 1919 (nomen novum for Falciger Borradaile, 1915b, gender masculine)
= Falciger Borradaile, 1915b (type species Periclimenes (Falciger) nilandensis Borradaile, 1915b, desig-
nated by Holthuis, 1955b, gender masculine)
= Kemponia Bruce, 2004b (type species Anchistia grandis Stimpson, 1860a, by original designation,
gender feminine)
Cuapetes agag (Kemp, 1922)
= Periclimenes (Ancylocaris) agag Kemp, 1922: 197; Figs 47-50; Plate 7, fg. 9. [Ross Channel, Port Blair,
Andamans, 4-8 fms]
Cuapetes akiensis (Kubo, 1936b)
= Periclimenes (Ancylocaris) akiensis Kubo, 1936b: 47; Plate 14. [Simokamogari-mura, Province
Aki]
Cuapetes americanus (Kingsley, 1878a)
= Anchistia americana Kingsley, 1878a: 96. [Key West, Florida]
= Periclimenes (Ancylocaris) bermudensis Lebour, 1949a: 1115; Fig. 6. [Mangrove Lake, Bermuda]
= Periclimenes (Ancylocaris) rhizophorae Lebour, 1949b: 605. [nomen novum for Periclimenes (Ancylo-
caris) bermudensis Lebour, 1949a nec Periclimenes (Periclimenaeus) bermudensis Armstrong, 1940]
Cuapetes amymone (De Man, 1902)
= Periclimenes amymone De Man, 1902: 829; Plate 25, Figs 53-53g. [Ternate]
Cuapetes anacanthus (Bruce, 1988b)
= Periclimenes anacanthus Bruce, 1988b: 105; Figs 1-5. [Polka Point, Dunwich North Stradbroke Is-
land, Moreton Bay, Queensland, 2729'S 15324'E, 0.5-1.0 m below LWS]
Cuapetes andamanensis (Kemp, 1922)
= Periclimenes (Ancylocaris) andamanensis Kemp, 1922: 204; 54-57. [Ross Channel, Port Blair, Anda-
mans, 4-8 fms]
Cuapetes calmani (Tatersall, 1921)
= Periclimenes calmani Tatersall, 1921: 385; Plate 27, fg. 11; Plate 28, fgs 14-15. [Sudan coast]
Cuapetes darwiniensis (Bruce, 1987a)
= Periclimenes darwiniensis Bruce, 1987a: 29; Figs 1-5. [Weed Reef, Darwin Harbour, 1231.6'S
13047.3'E, LWS]
Cuapetes demani (Kemp, 1915)
= Periclimenes demani Kemp, 1915: 279; Fig. 27; Plate 13, fg. 10. [Ennur backwater and Adar river
(both neighbourhood of Madras); Chilka Lake]
Cuapetes edwardsii (Paulson, 1875)
= Anch.[istia] Edwardsii Paulson, 1875: 114; Plate 17, fg. 2-2b. [Red Sea]
Cuapetes elegans (Paulson, 1875)
= Anch.[istia] elegans Paulson, 1875: 113; Plate 17, fg. 1-1h. [Red Sea]
= Periclimenes (Falciger) dubius Borradaile, 1915b: 211. [Minikoi]
Cuapetes ensifrons (Dana, 1852a)
= Anchistia ensifrons Dana, 1852a: 25. [in freto Balabac]
Cuapetes grandis (Stimpson, 1860a)
= Anchistia grandis Stimpson, 1860a: 39. [ad insulam Ousima]
= Periclimenes vitiensis Borradaile, 1898: 383. [Fii]
Cuapetes johnsoni (Bruce, 1987b)
= Periclimenes johnsoni Bruce, 1987b: 115; Figs 1-5. [Pasir Laba, Singapore, 121'N 10338'E]
Cuapetes kororensis (Bruce, 1977b)
= Periclimenes kororensis Bruce, 1977b: 33; Figs 1-4. [Koror, Palau Islands]
Cuapetes lacertae (Bruce, 1992c)
= Periclimenes lacertae Bruce, 1992c: 46; Figs 1, 2-6. [of Chinamans Ridge, Mrs Watsons Bay, Lizard
Island, 22-33 m]
Cuapetes longirostris (Borradaile, 1915b)
= Palmonella longirostris Borradaile, 1915b: 210. [Fardifolu Atoll, Maldive Islands]
= Periclimenes (Falciger) afnis Borradaile, 1915b: 211. [Salomon Island]
= Periclimenes (Ancylocaris) proximus Kemp, 1922: 201; Figs 51-53. [Ross Channel, Port Blair, Anda-
mans, 4-8 fms]
Cuapetes nilandensis (Borradaile, 1915b)
= Periclimenes (Falciger) nilandensis Borradaile, 1915b: 211. [S. Nilandu Atoll, Maldive Islands]
Cuapetes paulsoni (Bruce, 2003a)
= Periclimenes paulsoni Bruce, 2003a: 118; Fig. 3A-P. [Cape Flatery, inner wharf pile, 7 m]
Cuapetes platycheles (Holthuis, 1952c)
= Periclimenes (Harpilius) platycheles Holthuis, 1952c: 85; Fig. 33. [of Fau Island, west coast of Gebe
Island (31 m); of Atiationin, west coast of New Guinea (to 57 m)]
Cuapetes rapanui (Fransen, 1987)
= Periclimenes rapanui Fransen, 1987: 519; Figs 13-15. [Tahai, W coast of Easter Island, Chile, 39 m]
Cuapetes seychellensis (Borradaile, 1915b)
= Periclimenes (Falciger) seychellensis Borradaile, 1915b: 212. [Praslin, Seychelles]
Cuapetes suvadivensis (Borradaile, 1915b)
= Periclimenes (Falciger) suvadivensis Borradaile, 1915b: 212. [Suvadiva Atoll, Maldive Islands]
Cuapetes tenuipes (Borradaile, 1898)
= Periclimenes tenuipes Borradaile, 1898: 384. [New Britain]
= Periclimenes (Falciger) kolumadulensis Borradaile, 1915b: 213. [Kolumadulu Atoll, Maldive Islands]
= Periclimenes borradailei Rathbun, 1904: 34. [nomen novum for Periclimenes tenuipes Borradaile,
1898]
Cuapetes ungujaensis (Bruce, 1969a)
= Periclimenes ungujaensis Bruce, 1969a: 275. [Unguja Ukuu, Zanzibar Island, 618.8'S 3921.1'E,
0.5 f]
Dactylonia Fransen, 2002
= Dactylonia Fransen, 2002 (type species Pontonia ascidicola Borradaile, 1898, by original designation,
gender feminine)
Dactylonia anachoreta (Kemp, 1922)
= Pontonia anachoreta Kemp, 1922: 264; Figs 93-95. [Of Madras coast, 20 fms]
Dactylonia ascidicola (Borradaile, 1898)
= Pontonia ascidicola Borradaile, 1898: 389. [New Britain]
Dactylonia borradalei Bruce, 2005b: 375. [Neokumbi Reef, New Caledonia, 20-40 m]
Dactylonia carinicula Bruce, 2006a: 4; Figs 2-5. [Rhyi di Hamri, E of Hawlaf, N coast, 1240.429'N
5411.731'E, 7-9 m]
Dactylonia franseni Bruce, 2003b: 299; Figs 1-4. [Nyali, Old Port, Mombasa, Kenya, 2 m, in Ascidia sp.]
Dactylonia holthuisi Fransen, 2002: 295; Figs 194-201; Plate 13A. [Ambon, S coast, Seri Bay, 0345'S
12809'E, 25 m, in colony of Plurella spec.]
Dactylonia monnioti (Bruce, 1990c)
= Pontonia monnioti Bruce, 1990c: 183; Figs 21-24, 38e-h, 39i-j. [Chesterfeld Islands, 2446.6'S
15940.3'E, 285 m]
Dactylonia okai (Kemp, 1922)
= Pontonia okai Kemp, 1922: 261; Figs 89-92. [of Cape Negrais, Burma, 1525'N 9345'E, 40-49 fms]
Dasella Lebour, 1945
= Dasia Lebour, 1938 (type species Dasia herdmani Lebour, 1938, by monotypy, gender feminine;
invalid junior homonym of Dasia Gray, 1839 (Reptilia))
= Dasella Lebour, 1945 (nomen novum for Dasia Lebour, 1938, gender feminine)
Dasella ansoni Bruce, 1983a: 22; Figs 1-5. [Arafura Sea, 1058'S 13210'E, 27 m; see Bruce & Coombes,
1995]
Dasella brucei Berggren, 1990: 558. [Heron Island, Queensland, Australia, 15 m]
Dasella herdmaniae (Lebour, 1938)
= Dasia herdmani Lebour, 1938: 650; Plate 1. [Tuticorin, Gulf of Manaar, Madras]
Dasycaris Kemp, 1922
= Dasycaris Kemp, 1922 (type species Dasycaris symbiotes Kemp, 1922, by monotypy, gender femi-
Dasycaris ceratops Holthuis, 1952c: 176; Figs 87-88. [Borneo Bank, 225'S 11743'E, 34-0 m]
Dasycaris doederleini (Balss, 1924)
= Dasygius doederleini Balss, 1924: 49; Fig. 2. [Dzushi, Sagamibai, 130 m]
Dasycaris symbiotes Kemp, 1922: 240; Figs 76-77; Plate 9. [2.5 miles ESE of Santapilli Lt., near Vizagapa-
tam, Madras Coast, 15-17 fms and 3 miles ESE of Kabusa Island, Mergui, 1244'30''N 9755'30''E, 35
fms]
Dasycaris zanzibarica Bruce, 1973b: 247; Figs 1-5. [Changu Island reefs, of west coast of Zanzibar, 606.2'S
3908.9'E, 8 m]
Diapontonia Bruce, 1986c
= Diapontonia Bruce, 1986c (type species Diapontonia maranulus Bruce, 1986c, by original designation
Diapontonia maranulus Bruce, 1986c: 126; Figs 1-5. [of Wood Cay, West End, Grand Bahama Island,
2642.55'N 7901.72'W, 244-309 m]
Epipontonia Bruce, 1977c
= Epipontonia Bruce, 1977c (type species Epipontonia spongicola Bruce, 1977c, by original designation
Epipontonia anceps Bruce, 1983b: 19; Figs 1-10. [Heron Island, Queensland, 2328.0'S 15159.2'E, 18 m]
Epipontonia hainanensis Li, 1999: 357; Figs 1-3. [Yezhu Island, Yalong Bay, Hainan Island, 6-9 m, from
dead corals]
Epipontonia spongicola Bruce, 1977c: 308; Figs 1-5. [Wasini Channel, Kenya, 439.4'S 3922.2'E, 6 fms]
Epipontonia tahitiensis Bruce, 2004c: 280; Figs 1-4. [French Polynesia, Tahiti]
Eupontonia Bruce, 1971b
= Eupontonia Bruce, 1971b (type species Eupontonia noctalbata Bruce, 1971b, by original designation
Eupontonia noctalbata Bruce, 1971b: 227; Figs 1-5. [Anse Etoile, Mah, Seychelle Islands, 0435'12"S
5527'48"E, coral reef fats, 0.3 m]
Eupontonia oahu Bruce, 2010d: 406; Figs 1-5. [Kahe Point, Oahu, Hawaiian Islands]
Exoclimenella Bruce, 1995
= Exoclimenella Bruce, 1995 (type species Periclimenes Petithouarsi var. denticulata Nobili, 1906b, by
= Exoclimenella uri & Bruce, 1995 (type species Periclimenes Petithouarsi var. denticulata Nobili,
1906b, by original designation, gender feminine)
Exoclimenella denticulata (Nobili, 1906b)
= Periclimenes Petithouarsi var. denticulata Nobili, 1906b: 257. [Gatavake]
Exoclimenella maldivensis uri & Bruce, 1995: 622; Figs 1-5. [Genego Islet, North Nilandu Atoll, Maldive
Islands, outer reef slope, 20 m]
Exoclimenella sibogae (Holthuis, 1952c)
= Periclimenes (Harpilius) sibogae Holthuis, 1952c: 73; Figs 28-29. [Banda anchorage, 9-36 m]
Exoclimenella sudanensis uri & Bruce, 1995: 631; Figs 6-8. [coral reef near Port Sudan, Sudan, Red Sea,
in coral head of Stylophora pistillata]
Exopontonia Bruce, 1988c
= Exopontonia Bruce, 1988c (type species Exopontonia malleatrix Bruce, 1988c, by original designation
Exopontonia malleatrix Bruce, 1988c: 123; Figs 1-5. [Ashmore Reef, Timor Sea, 1216'S 12302'E, intertidal]
Fennera Holthuis, 1951b
= Fennera Holthuis, 1951b (type species Fennera chacei Holthuis, 1951b, by original designation and
Fennera chacei Holthuis, 1951b: 171; Plate 54. [Secas Islands, Bay of South Island, shallow water]
Fennera holthuisi Marin, 2011: 34; Figs 5-7, 8d-f. [French Polynesia, Moorea, outer reef slope, 172902S
1495209W, 10 m, in Pocillopora]
Hamiger Borradaile, 1916
= Hamiger Borradaile, 1916 (type species Periclimenes (Hamiger) novae-zealandiae Borradaile, 1916, by
Hamiger novaezealandiae (Borradaile, 1916)
= Periclimenes (Hamiger) novae-zealandiae Borradaile, 1916: 87; Fig. 4. [seven miles E. of North Cape,
New Zealand, 128 m]
Hamodactyloides Fujino, 1973b
= Hamodactyloides Fujino, 1973b (type species Hamodactylus incompletus Holthuis, 1958, by original
Hamodactyloides incompletus (Holthuis, 1958)
= Hamodactylus incompletus Holthuis, 1958: 11; Fig. 4. [Sharam a Sheikh, Sinai Peninsula]
= Hamodactyloides ishigakiensis Fujino, 1973b: 174; Figs 1-3. [Kabira Bay, Ishigaki-jima Island, Ryukyu
Islands, 1 m]
Hamodactylus Holthuis, 1952c
= Hamodactylus Holthuis, 1952c (type species Hamodactylus boschmai Holthuis, 1952c, by original
Hamodactylus aqabai Bruce & Svoboda, 1983: 26; Figs 10-14. [Aqaba, Jordan]
Hamodactylus boschmai Holthuis, 1952c: 209; Figs 102-104. [Ternate, of Halmahera, and Djedan, Kepu-
lauan Aru, Indonesia, 2-13 m]
Hamodactylus noumeae Bruce, 1970a: 539; Fig. 2. [between Isle aux Canards and Islt Matre, near
Noumea, New Caledonia, 25 m]
Hamopontonia Bruce, 1970b
= Hamopontonia Bruce, 1970b (type species Hamopontonia corallicola Bruce, 1970b, by original desig-
Hamopontonia corallicola Bruce, 1970b: 41; Figs 1-4. [Kat O Chau, Mirs Bay, New Territories, Hong Kong,
2232.1'N 11417.95'E]
Hamopontonia essingtoni Bruce, 1986d: 158; Figs 1C, 11-14, 15D-G. [Coral Bay, Port Essington, 1111.05'S
13203.4'E, 6 m]
Harpiliopsis Borradaile, 1917
= Harpiliopsis Borradaile, 1917 (type species Palmon Beaupressi Audouin, 1826, by original designa-
Harpiliopsis beaupresii (Audouin, 1826)
= Palmon beaupresii Audouin, 1826: 91 (accompanying fgures in Savigny, 1826). [Egypt]
= Pontonia (Harpilius) dentata Richters, 1880: 165; Plate 17, Figs 36-38. [Fouquets]
Harpiliopsis depressa (Stimpson, 1860a)
? = Anchistia gracilis Dana, 1852a: 25. [in mari Suluensi]
= Harpilius depressus Stimpson, 1860a: 38. [ad insulam Hawaii, inter madreporas]
= Pelias notatus Heller, 1862a: 526. [Nicobaren]
= Anchistia notata Heller, 1865: 109; Plate 10, fg. 3. [Nicobaren]
= Periclimenes pusillus Rathbun, 1906: 921; Fig. 71; Plate 24, fg. 7. [south coast of Oahu, surface]
Harpiliopsis spinigera (Ortmann, 1890)
= Anchistia spinigera Ortmann, 1890: 511; Plate 36, fg. 23-23a. [Samoa]
= Harpilius depressus var. gracilis Kemp, 1922: 234; Fig. 71. [Andamans]
Harpilius Dana, 1852a
= Harpilius Dana, 1852a (type species Harpilius lutescens Dana, 1852a, by monotypy, gender mascu-
line; name placed on the Ofcial List of Generic Names in Zoology in Opinion 712 in 1964)
Harpilius bayeri (Holthuis, 1981)
= Periclimenes bayeri Holthuis, 1981: 792; Fig. 3. [Ine village, Arno Atoll, Marshall Islands, outer edge
of sea reef, on coral, from head of Pocillopora spec. according to K. Reed (pers. comm.)]
= Periclimenes (Harpilius) bayeri Holthuis, 1953b: 56. [nomen nudum]
Harpilius consobrinus De Man, 1902: 836; Plate 26, fg. 54-54c. [Ternate]
Harpilius lutescens Dana, 1852a
= Harpilius lutescens Dana, 1852a: 25. [insula Tongatabu]
= Periclimenes (Ancylocaris) amamiensis Kubo, 1940b: 44; Figs 11, 12. [sima, Riu-Kiu Islands]
Harpilius spinifer Bruce, 2006b: 272; Figs 1-5. [Ilt Maitre, New Caledonia, 4 m]
Holthuisaeus Anker & De Grave, 2010
= Holthuisaeus Anker & De Grave, 2010 (type species Periclimenes (Periclimenaeus) bermudensis Arm-
strong, 1940, by original designation and monotypy, gender masculine)
Holthuisaeus bermudensis (Armstrong, 1940)
= Periclimenes (Periclimenaeus) bermudensis Armstrong, 1940: 4; Figs 2, 3A-F. [The Reach, St. George
Island, Bermuda]
Ischnopontonia Bruce, 1966c
= Ischnopontonia Bruce, 1966c (type species Philarius lophos Barnard, 1962, by original designation
Ischnopontonia lophos (Barnard, 1962)
= Philarius lophos Barnard, 1962: 242; Fig. 2. [Inhaca Island, Delagoa Bay, U.W.]
Isopontonia Bruce, 1982a
= Isopontonia Bruce, 1982a (type species Isopontonia platycheles Bruce, 1982a, by original designation
Isopontonia platycheles Bruce, 1982a: 55; Figs 1-4. [North Cay, Bennet Island, Chesterfeld Islands, 1948.0'S
15817.0'E, 15 m]
Izucaris Okuno, 1999a
= Izucaris Okuno, 1999a (type species Izucaris masudai Okuno, 1999a, by original designation and
Izucaris crosnieri Li, 2008: 207; Figs 1-4, 5A. [Marquesas Islands, Eiao, 759.2'S 14044.2'W, 100 m]
Izucaris masudai Okuno, 1999a: 399; Figs 1-6. [3452.7'N 3908.2'E, Izu Oceanic Park, Izu Peninsula, Hon-
shu, Japan, 27 m]
Jocaste Holthuis, 1952c
= Jocaste Holthuis, 1952c (type species Coralliocaris lucina Nobili, 1901c, by monotypy, gender femi-
nine)
Jocaste japonica (Ortmann, 1890)
= Coralliocaris superba var. japonica Ortmann, 1890: 50; Plate 36, fg. 22. [Japan, Kagoshima]
? = Cavicheles kempi Holthuis, 1952c: 205; Figs 99-101. [Ternate, 4 m]
Jocaste lucina (Nobili, 1901c)
= C.[oralliocaris] lucina Nobili, 1901c: 5. [Eritrea]
? = Coralliocaris lamellirostris Stimpson, 1860a: 38. [ad insulam Loo Choo, inter corallia ad prof 2 org]
Jocaste platysoma Fransen, 1994: 111; Figs 65-84; Plate 2C. [St. Joseph Atoll, S rim, 527'S 5321'E, 10 m, on
tabular Acropora]
Laomenes Clark, 1919
= Corniger Borradaile, 1915b (type species Periclimenes (Corniger) ceratophthalmus Borradaile, 1915b,
designated by Borradaile, 1917, gender masculine; invalid junior homonym of Corniger Agassiz,
1831 (Pisces) and Corniger Boehm, 1879 (Pycnogonida))
= Laomenes Clark, 1919 (nomen novum for Corniger Borradaile, 1915b)
= Parapontonia Bruce, 1968a (type species Parapontonia nudirostris Bruce, 1968a, by original designa-
Laomenes amboinensis (De Man, 1888b)
= Anchistia amboinensis De Man, 1888b: 546; Plate 22a, fgs 2-2b. [Amboina]
Laomenes ceratophthalmus (Borradaile, 1915b)
= Periclimenes (Corniger) ceratophthalmus Borradaile, 1915b: 211. [Mal Atoll, Maldive Islands, on
crinoid]
Laomenes clarki Marin, 2009a: 13; Figs 13-19. [Vietnam, Nhatrang Bay, Nok Island, 15 m, on crinoid Zy-
gometra cf. punctata]
Laomenes cornutus (Borradaile, 1915b)
= Periclimenes (Corniger) cornutus Borradaile, 1915b: 211. [Mal Atoll, Maldive Islands, on crinoid]
Laomenes holthuisi Marin & Okuno, 2010: 462; Figs 1-6. [Pacifc Ocean, Japan, Izu Islands, Hachio Island,
Aozumi, 15 m]
Laomenes jackhintoni (Bruce, 2006c)
= Periclimenes jackhintoni Bruce, 2006c: 23; Figs 1-5. [Nukualofa, Tongatapu, Tonga, 1.5 m]
Laomenes nudirostris (Bruce, 1968a)
= Parapontonia nudirostris Bruce, 1968a: 1149; Figs 1-5. [Tiar Bay, Noumea, New Caledonia, 2210'S
16615'E]
Laomenes pardus Marin, 2009a: 19; Figs 15-18. [Vietnam, Nhatrang Bay, Mun Island, 5 m, on crinoid
Comantheria briareus]
Laomenes tigris Marin, 2009a: 25; Figs 20-24. [Mun Island, Nhatrang Bay, Vietnam, 15-20 m, on crinoid
Stephanometra sp.]
Leptomenaeus Bruce, 2007b
= Leptomenaeus Bruce, 2007b (nomen novum for Leptomenes Bruce, 2006d, gender masculine)
= Leptomenes Bruce, 2006d (type species Periclimenes dolichosternum Okuno & Mitsuhashi, 2003, by
original designation and monotypy, gender masculine; invalid junior homonym of Leptomenes Soi-
ka, 1939 (Insecta))
Leptomenaeus dolichosternum (Okuno & Mitsuhashi, 2003)
= Periclimenes dolichosternum Okuno & Mitsuhashi, 2003: 487-496; Figs 1-5. [Ou-shima Harbor,
Kume Island, Ryukyu Islands, 2620.1'N 12649.2'E, 1.5 m]
Leptomenaeus nhatrangensis uri & Hork, 2008: 46; Figs 1-5. [Hon Chong Beach, N of Nhatrang Bay
(South China Sea), Vietnam, 1217'45.8''N 10914'05.7''E, 1 m]
Lipkebe Chace, 1969
= Lipkebe Chace, 1969 (type species Lipkebe holthuisi Chace, 1969, by original designation and mono-
Lipkebe holthuisi Chace, 1969: 263; Figs 8-9. [Gulf of Mexico, west-northwest of Dry Tortugas, Florida,
2513'N 8355'W, 119 m]
Lipkemenes Bruce & Okuno, 2010
= Lipkemenes Bruce & Okuno, 2010 (type species Periclimenes (Periclimenes) lanipes Kemp, 1922, by
Periclimenes lanipes Kemp, 1922
= Periclimenes (Periclimenes) lanipes Kemp, 1922: 156; Plate 4, fg. 4. [Mergui Archipelago, 1248'N
9816'10"E, 24 fms]
Manipontonia Bruce, Okuno & Li, 2005
= Manipontonia Bruce, Okuno & Li, 2005 (type species Urocaris psamathe De Man, 1902, by original
Manipontonia persiana Marin, 2010: 64; Figs 3-4. [Indian Ocean, Persian Gulf, 2849.661'N 4948.506'E, 51
m depth]
Manipontonia psamathe (De Man, 1902)
= Urocaris psamathe De Man, 1902: 816; Plate 25, fg. 51. [Ternate]
Margitonia Bruce, 2007c
= Margitonia Bruce, 2007c (type species Periclimenes insolitus Bruce, 1974b, by original designation
Margitonia insolitus (Bruce, 1974b)
= Periclimenes insolitus Bruce, 1974b: 293; Figs 1-8. [Waikiki Beach, Oahu, Hawaii, 2115.9'N
15750.5'W]
Mesopontonia Bruce, 1967b
= Mesopontonia Bruce, 1967b (type species Mesopontonia gorgoniophila Bruce, 1967b, by original des-
Mesopontonia brevicarpus Li & Bruce, 2006: 652; Figs 7-8. [La Runion, 2051.5'S 5536.87'E, 270 m]
Mesopontonia brucei Burukovsky, 1991b: 38; Figs 1.8-1.18. [3316'S 4353'E, 415-460 m]
Mesopontonia gorgoniophila Bruce, 1967b: 13; Figs 5-9. [2147.7'N 11628.5'E to 2143.3'N 11628.0'E, 64-72 fms]
Mesopontonia gracilicarpus Bruce, 1990c: 202; Figs 34-37, 39l-m. [New Caledonia, 2256'N 16714'E, 398-
410 m]
Mesopontonia monodactylus Bruce, 1991a: 392; Figs 65-69. [Loyalty Islands, 2035.0'S 16654.0'E, Uva, 460
m]
Mesopontonia verrucimanus Bruce, 1996: 216; Figs 8a-n, 29c. [Indonesia, Tanimbar Islands, 759'S 13302'E,
184-186 m]
Metapontonia Bruce, 1967b
= Metapontonia Bruce, 1967b (type species Metapontonia fungiacola Bruce, 1967b, by original designa-
Metapontonia fungiacola Bruce, 1967b: 24: Figs 10-12. [Pamanzi Reef, Mayote Island, Comore Archipelago]
Miopontonia Bruce, 1985a
= Miopontonia Bruce, 1985a (type species Miopontonia yongei Bruce, 1985a by original designation
Miopontonia yongei Bruce, 1985a: 168; Figs 1-5. [1904.3'S 11815.5'E, 80 m]
Neoanchistus Bruce, 1975a
= Neoanchistus Bruce, 1975a (type species Neoanchistus cardiodytes Bruce, 1975a, by original designa-
Neoanchistus cardiodytes Bruce, 1975a: 151; Figs 1-6. [Nosy B, Madagascar]
Neoanchistus nasalis Holthuis, 1986c: 264; Figs 1-2. [artifcial reef near Raysut Harbour, Dhofar, southern
Oman, 1657'45''N 5400'E, in Chlamys townsendi]
Neoclimenes Mitsuhashi, Li & Chan, 2010
= Neoclimenes Mitsuhashi, Li & Chan, 2010 (type species Neoclimenes holthuisi Mitsuhashi, Li &
Chan, 2010, by original designation, gender masculine)
Neoclimenes holthuisi Mitsuhashi, Li & Chan, 2010: 518; Figs 1-6. [2140.2'N 11743.4'E, 513-509 m]
Neopericlimenes Heard, Spote & Bubucis, 1993
= Neopericlimenes Heard, Spote & Bubucis, 1993 (type species Neopericlimenes thornei Heard, Spote
& Bubucis, 1993, by original designation and monotypy, gender masculine)
Neopericlimenes thornei Heard, Spote & Bubucis, 1993: 795; Figs 1-4. [fringing reef of north shore of Pine
Cay, Turks and Caicos Islands, British West Indies, 40 m, approx. 2153'N 7206'W]
Neopontonides Holthuis, 1951b
= Neopontonides Holthuis, 1951b (type species Periclimenes beaufortensis Borradaile, 1920, by original
Neopontonides beaufortensis (Borradaile, 1920)
= Periclimenes beaufortensis Borradaile, 1920: 132. [Beaufort, North Carolina]
Neopontonides brucei Fransen & de Almeida, 2009: 838; Figs 1-4. [Boquiro beach, Ubatuba, NE So Paulo,
Brazil, 32 m, mud]
Neopontonides chacei Heard, 1986: 472; Figs 1A, 2-3, 4B-D. [reef just south of Marigot Bay, St. Lucia Island,
West Indies, 4-6 m]
Neopontonides dentiger Holthuis, 1951b: 193; Plate 61. [Ecuador, of Cape San Francisco, 2 fms]
Neopontonides henryvonprahli Ramos, 1995: 231; Figs 1-3. [Punta Alta, Baha Mlaga, simbionte de Muri-
cea robusta, 5 m]
Nippontonia Bruce & Bauer, 1997
= Nippontonia Bruce & Bauer, 1997 (type species Nippontonia minirostris Bruce & Bauer, 1997, by
Nippontonia minirostris Bruce & Bauer, 1997: 1711; Figs 1-8. [Nishihama, Aka-jima, Kerama group, Ryukyu
Islands, Japan, 15 m]
Notopontonia Bruce, 1991c
= Notopontonia Bruce, 1991c (type species Notopontonia platycheles Bruce, 1991c, by original designa-
Notopontonia platycheles Bruce, 1991c: 608; Figs 1-6, 14E. [northwest of Robe, South Australia, 3653'S
13953'E, 64 m]
Odontonia Fransen, 2002
= Odontonia Fransen, 2002 (type species Pontonia katoi Kubo, 1940b, by original designation, gender
feminine)
Odontonia compacta (Bruce, 1996)
= Pontonia compacta Bruce, 1996: 242; Figs 15-18. [New Caledonia, 2236.3'S 16714.2'E, 10-60 m]
Odontonia katoi (Kubo, 1940b)
= Pontonia katoi Kubo, 1940b: 55; Figs 21-23. [Of Simoda, Siduoka Prefecture]
Odontonia maldivensis Fransen, 2006a: 718; Figs 2-5. [Maldives, S Mal Atoll, ocean side North reef,
0407.54'N 7330.55'E, 10 m, in Polycarpa cryptocarpa]
Odontonia rufopunctata Fransen, 2002: 352; Figs 229-234; Plate 17. [SW Sulawesi, Spermonde Archipela-
go, Kudingareng Keke, 20 m, in ascidian]
Odontonia seychellensis Fransen, 2002: 361; Figs 235-240; Plate 18. [Seychelles, Bird Island, of N coast,
342'S 5512'E, 20 m, in Ascidiidae]
Odontonia sibogae (Bruce, 1972d)
= Pontonia sibogae Bruce, 1972d: 182; Fig. 1. [Curtis Channel, Port Curtis, Queensland, Australia, 23
fms]
Odontonia simplicipes (Bruce, 1996)
= Pontonia simplicipes Bruce, 1996: 247; Figs 19-20, 29i. [Chesterfeld Islands, 1915.15'S 15847.73'E, 71 m]
Onycocaridella Bruce, 1981b
= Onycocaridella Bruce, 1981b (type species Onycocaridella prima Bruce, 1981b, by original designa-
Onycocaridella monodoa (Fujino & Miyake, 1969b)
= Onycocaridella monodoa Fujino & Miyake, 1969b: 405; Figs 1-5. [Kasari-zaki, Amami-oshima Island,
Ryukyu Islands, Japan, 1 m]
Onycocaridella prima Bruce, 1981b: 243; Figs 1-6. [Wistari Reef, Heron Island, Capricorn Islands, Queens-
land, 2325.7'S 15155.0'E, 12 m]
Onycocaridella stenolepis (Holthuis, 1952c)
= Onycocaris stenolepis Holthuis, 1952c: 148; Figs 66-68. [southeast side of Pearl Bank, Sulu Archi-
pelago, 15 m]
Onycocaridites Bruce, 1987c
= Onycocaridites Bruce, 1987c (type species Onycocaridites anomodactylus Bruce, 1987c, by original
Onycocaridites anomodactylus Bruce, 1987c: 772; Figs 1-4. [Arafura Sea, 1040'S 13350'E, 60 m]
Onycocaris Nobili, 1904
= Onycocaris Nobili, 1904 (type species Coralliocaris (Onycocaris) aualitica Nobili, 1904, designated by
Holthuis, 1952c, gender feminine)
Onycocaris amakusensis Fujino & Miyake, 1969b: 413; Figs 6, 8a-c, 9a-c. [Tsuji-shima Islet, Amakusa Is-
land, in sponge collected at low tide level]
Onycocaris aualitica (Nobili, 1904)
= Coralliocaris (Onycocaris) aualitica Nobili, 1904: 233. [Djibouti]
Onycocaris balssi Bruce, 2011b: 478; Figs 1-5. [Table Head, Port Essington, Northern Territory, Australia,
1113.5'S 13210.5'E, 3 m, in unidentifed sponge]
Onycocaris bocki Bruce, 1992e: 335; Figs 4-6. [Tuvalu, Niue, leeward reef]
Onycocaris callyspongiae Fujino & Miyake, 1969b: 422; Figs 10-12. [Tomioka, Amakusa Island, in sponges]
Onycocaris fujinoi Bruce, 2011b: 487. [Terasaki, Yoron-jima, Ryukyu Islands, Japan]
Onycocaris furculata Bruce, 1979c: 324; Figs 1-4. [La Saline, La Runion, approximately 2120'S 5500'E, 20 m]
Onycocaris longirostris Bruce, 1980a: 15; Figs 6-10. [Ilot Matre, Nouma, New Caledonia, 20 m]
Onycocaris nieli Bruce, 2011a: 320; Figs 1-5. [Heron Island, Capricorn Islands, Queensland, reef fat]
Onycocaris oligodentata Fujino & Miyake, 1969b: 415; Figs 7, 8d-f, 9d-f. [Tomioka, Amakusa Island, in
sponge fshed up entangling with gill net, 35 m]
Onycocaris profunda Bruce, 1985b: 241; Figs 8-11. [1333.0'N 12210.1'E, 81-84 m]
Onycocaris quadratophthalma (Balss, 1921a)
= Pontonia quadratophthalma Balss, 1921a: 15; Fig. 7. [Cape Jaubert]
Onycocaris seychellensis Bruce, 1971c: 208; Figs 1-6. [Anse Etoile, Mah, Seychelle Islands]
Onycocaris spinosa Fujino & Miyake, 1969b: 429; Figs 13-15. [Terasaki, Yoron-jima Island, Ryukyu Is-
lands, in sponge, 1 m]
Onycocaris stradbrokei Bruce, 1998c: 390; Figs 2-3. [Myora Reef, North Stradbroke Island, Queensland,
2729'S 15325'E, 9 m]
Onycocaris temiri Marin, 2005: 113; Figs 1-8. [Mung Island, 16 m, inside boring sponge Kallipilidion]
Onycocaris trullata Bruce, 1978a: 269; Figs 36-41. [Tany-Kely, Madagascar, 1328'S 4812'E, 28 m]
Onycocaris zanzibarica Bruce, 1971d: 293; Figs 1-2. [channel between Chumbe Island and mainland of
Zanzibar Island, 616.0'S 3912.6'E, 10 fms]
Onycomenes Bruce, 2009a
= Onycomenes Bruce, 2009a (type species Onycocaridella antokha Marin, 2007a, by original designa-
Onycomenes antokha (Marin, 2007a)
= Onycocaridella antokha Marin, 2007a: 221; Figs 5-9. [South China Sea, Vietnam, Nhatrang Bay, Nok
Island, 10 m]
Orthopontonia Bruce, 1982b
= Orthopontonia Bruce, 1982b (type species Periclimenaeus ornatus Bruce, 1970c, by original designa-
Orthopontonia ornata (Bruce, 1970c)
= Periclimenaeus ornatus Bruce, 1970c: 313. [Heron Island, Capricorn Group, Great Barrier Reef, Aus-
tralia, litoral]
Palaemonella Dana, 1852a
= Palaemonella Dana, 1852a (type species Palaemonella tenuipes Dana, 1852a, designated by Kingsley,
1880, gender feminine; name placed on the Ofcial List of Generic Names in Zoology in Opinion
470 in 1957)
Palaemonella aliska Marin, 2008a: 376; Figs 1-5, 9a. [South China Sea, Vietnam, Nhatrang Bay, Tre Island,
Tre Bay, mangrove litoral]
Palaemonella asymmetrica Holthuis, 1951b: 19; Plate 5. [Sullivan Bay, James Island]
Palaemonella atlantica Holthuis, 1951a: 152; Fig. 31. [So Pedro Bay, So Vicente, Cape Verde Islands,
1650'N 2504'W]
Palaemonella burnsi Holthuis, 1973a: 24; Figs 8-9. [Cape Kinau Peninsula, Maui, Hawaiian Islands; small
lava pool near coast of Keoneoio (= La Perouse) Bay, at the extreme east end of the Cape]
Palaemonella crosnieri Bruce, 1978a: 210; Figs 2-4. [Iles Glorieuses, 1128.1'S 27[sic]21.1'E, 20 m]
Palaemonella dionesae Bruce, 2010e: 151; Fig. 1. [Rocky head, Enderby Island, Dampier Archipelago,
2040'14''S 11627'96''E, 12.5 m]
Palaemonella disalvoi Fransen, 1987: 511; Figs 7-12. [Tahai, west coast of Easter Islands, 35 m]
Palaemonella dolichodactylus Bruce, 1991b: 232; Figs, 6f-l, 7. [2214.5'S 16702.0'E, Lagon Est, 65-70 m]
Palaemonella foresti Bruce, 2002b: 279; Figs 1-4. [BP Oil Refnery Jety, Cockburn Sound, Western Aus-
tralia, 3213.9'S 11545.14'E, 7 m]
Palaemonella hachio Okuno, 1999b: 739; Figs 1-3. [3303.5'N 13947.9'E, southern Japan, Izu Islands,
Hachio-jima Island, Occho-gahama, submarine cave, 20 m]
Palaemonella holmesi (Nobili, 1907)
= Periclimenes Holmesi Nobili, 1907: 5. [nomen novum for Anchista tenuipes Holmes, 1900]
= Anchista tenuipes Holmes, 1900: 216. [Santa Catalina Island]
Palaemonella komaii Li & Bruce, 2006: 657; Figs 9-11. [Tonga, Vavau group, 1837'S 17403'W, 327-360
m]
Palaemonella lata Kemp, 1922: 127; Figs 3-6. [Aberdeen, Port Blair, Andamans, rock pool at low tide]
Palaemonella longidactylus Hayashi, 2009: 97; Figs 5-7. [Beppu Bay, Ooita Prefecture]
Palaemonella maziwi Bruce, 2002c: 15; Fig. 1. [Maziwi Island, Pangani, Tanganyika, 30.6'S 3904.06'E, 2 m]
Palaemonella meteorae Bruce, 2008b: 64; Figs 1-2. [Red Sea, of Port Sudan, 1933.3'N 3716.4'E - 1933.9'N
3716.2'E, 519-544 m]
Palaemonella potsi (Borradaile, 1915b)
= Periclimenes (Falciger) potsi Borradaile, 1915b: 212. [Torres Straits, on Comanthus]
Palaemonella pusilla Bruce, 1975b: 169; Figs 1-5. [Kisiti Island, near Wasin, Kenya, 443.3'S 3922.15'E, on
sheltered coral reef at LWS]
Palaemonella rotumana (Borradaile, 1898)
= Periclimenes rotumanus Borradaile, 1898: 383. [Rotuma]
= Palaemonella vestigialis Kemp, 1922: 123; Figs 1-2; Plate 3, fg. 2. [Port Blair, Andamans]
Palaemonella spinulata Yokoya, 1936: 135; Fig. 4. [Sagami Bay, of Misaki to Yokosuka, Miura Peninsula,
3510.53'N 13934.56'E to 3910.53'N 13934.53'E, 73-62 m; neotype designation by Hayashi, 2009]
Palaemonella tenuipes Dana, 1852a
= Palmonella tenuipes Dana, 1852a: 25. [in mari Suluensi]
= Palmonella tridentata Borradaile, 1899: 1007; Plate 64, Figs 8-8c. [Funafuti]
= Palmonella elegans Borradaile, 1915b: 210. [Salomon Island]
Paraclimenaeus Bruce, 1988d
= Paraclimenaeus Bruce, 1988d (type species Periclimenaeus fmbriatus Borradaile, 1915b, by original
Paraclimenaeus dentimanus (Mitsuhashi & Chan, 2008)
= Apopontonia dentimanus Mitsuhashi & Chan, 2008: 32; Figs 3-7. [North of Doljo, Panglao Island,
935.9'N 12344.5'E, 24 m]
Paraclimenaeus fmbriatus (Borradaile, 1915b)
= Periclimenus fmbriatus Borradaile, 1915b: 213. [Makalu Atoll, Maldive Islands; holotype designa-
tion by Bruce, 1988d invalid]
Paraclimenaeus seticauda (Bruce, 2008c)
= Apopontonia seticauda Bruce, 2008c: 3; Figs 2-5, 14A. [1827'37''S 12008'41''E, 80 m]
Paraclimenaeus spinicauda (Bruce, 1969b)
= Periclimenaeus spinicauda Bruce, 1969b: 164. [South China Sea, 2057.5'N 11555.0'E 2057.5'N
11558.6'E, 35-36 fms]
= Apopontonia dubia Bruce, 1981c: 225; Figs 1-3. [Shag Rock, E of North Stradbroke Island, Queens-
land, Australia, 2725'S 15332'E, 20 m]
Paraclimenes Bruce, 1995
= Paraclimenes Bruce, 1995 (type species Periclimenes franklini Bruce, 1990e, by original designation,
gender masculine)
Paraclimenes franklini (Bruce, 1990e)
= Periclimenes franklini Bruce, 1990e: 55; Figs 1-5. [1721.7'S 14648.52'E, 296-302 m]
Paraclimenes gorgonicola (Bruce, 1969a)
= Periclimenes gorgonicola Bruce, 1969a: 257. [South China Sea, 2147.7'N 11628.5'E to 2143.3'N
11628.0'E, 60-70 fms]
Paraclimenes setirostris (Bruce, 1991a)
= Periclimenes setirostris Bruce, 1991a: 364; Figs 45-49. [Chesterfeld Islands, 2532.8'S 15946.1'E, 300 m]
Paranchistus Holthuis, 1952c
= Paranchistus Holthuis, 1952c (type species Anchistus biunguiculatus Borradaile, 1898 (junior subjec-
tive synonym of Pontonia armata H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840], by
Paranchistus armatus (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840])
= Pontonia armata H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 359. [prs de ctes de
la Nouvelle-Irlande]
= Anchistus biunguiculatus Borradaile, 1898: 387. [British New Guinea]
= Anchistus oshimai Kubo, 1949b: 26; Figs 1-2. [Helen Atoll]
Paranchistus liui Li, Bruce & Manning, 2004: 535; Figs 21-23. [21N 108E, 30 m]
Paranchistus nobilii Holthuis, 1952c: 100; Figs 41-42. [Arzana Island, Rakas Zakoum Bank, Persian Gulf,
out of Spondylus gaederopus]
Paranchistus ornatus Holthuis, 1952c: 97; Figs 39-40. [Mozambique]
Paranchistus pycnodontae Bruce, 1978b: 233; Figs 1-5; Plate 39. [Heron Island, Capricorn Group, Queens-
land, 3 m]
= Paranchistus serenei Bruce, 1983c: 890; Figs 7H-I, 9. [Seleman Bay, Seram Island]
Paranchistus spondylis Suzuki, 1971: 15; Figs 8-9. [Shiraiso, near the Manazuru M.B.L.]
Paratypton Balss, 1914d
= Paratypton Balss, 1914d (type species Paratypton siebenrocki Balss, 1914d, by monotypy, gender
masculine; name placed on the Ofcial List of Generic Names in Zoology in Opinion 470 in 1957)
Paratypton siebenrocki Balss, 1914d: 84; Fig. 1. [Senafr, Koseir, Mersa Scheikh; Jaluit, Samoa]
Patonia Mitsuhashi & Chan, 2006
= Patonia Mitsuhashi & Chan, 2006 (type species Patonia mclaughlinae Mitsuhashi & Chan, 2006, by
Patonia mclaughlinae Mitsuhashi & Chan, 2006: 390; Figs 1-6. [Taiwan northeastern coast, 2434.71'N
1224.02'E, 209-280 m]
Periclimenaeus Borradaile, 1915b
= Periclimenus Borradaile, 1915b (type species Periclimenus robustus Borradaile, 1915b, designated
by Borradaile, 1917, gender masculine; name placed on the Ofcial List of Generic Names in Zool-
Periclimenaeus arabicus (Calman, 1939)
= Periclimenes (Periclimenus) arabicus Calman, 1939: 210; Fig. 4. [Khali al Masirah, eastern Oman,
1922.6'N 5753.0'E, 13.5 m, from surface of sponge]
= Periclimenaeus ohshimai Miyake & Fujino, 1967: 275; Fig. 1. [Takamatsu, Amakusa Islands, Kyushu]
Periclimenaeus ardeae Bruce, 1970c: 310. [Heron Island, Capricorn Group, Great Barrier Reef, Queens-
land, Australia, litoral]
Periclimenaeus arthrodactylus Holthuis, 1952c: 122; Figs 51-53. [Sailus-ketjil, Paternoster Islands, up to 18
m]
Periclimenaeus ascidiarum Holthuis, 1951b: 80; Plate 22, fgs g-l; Plate 23. [Bird Key Reef, Tortugas]
Periclimenaeus atlanticus (Rathbun, 1901)
= Coralliocaris atlantica Rathbun, 1901: 122; Fig. 26. [Of St. Thomas 20-23 fms]
Periclimenaeus aurae dos Santos, Calado & Arajo, 2008: 156; Figs 1-4, 5C, K. [Garajau, south of Madeira
Island, eastern Atlantic, 32.36N 16.85W, 6-7 m]
Periclimenaeus bidentatus Bruce, 1970c: 305. [Heron Island, Capricorn Group, Great Barrier Reef, Queens-
land, Australia, litoral]
Periclimenaeus bouvieri (Nobili, 1904)
= Typton Bouvieri Nobili, 1904: 233. [Djibouti]
Periclimenaeus bredini Chace, 1972: 26; Fig. 5. [Isla Mujeres]
Periclimenaeus brucei Cardoso & Young, 2007: 302; Figs 20-25. [Rocas Atoll, outer reef, 20 m, in Ircinia sp.]
Periclimenaeus caraibicus Holthuis, 1951b: 110; Plate 32, fgs h-j; Plate 34. [British West Indies, Tobago,
Buccoo Reef]
Periclimenaeus chacei Abele, 1971: 38; Figs 1-2. [Gulf of Mexico, of the West coast of Florida, 2831'N
8416'W, 26 m]
Periclimenaeus colodactylus Bruce, 1996: 222; Figs 9-10. [New Caledonia, Lagoon, Uatio Islet, 20-25 m]
Periclimenaeus crassipes (Calman, 1939)
= Periclimenes (Ancylocaris) crassipes Calman, 1939: 211; Fig. 5. [Ghubbat Sawquirah, southeastern
Oman, 18025.5'N 57025'E, 38 m; animals pulled of calcareous sponges or removed from the de-
bris in the jar containing them]
Periclimenaeus creef Bruce, 2010f: 29; Figs 7-11. [Heron island reef, north-eastern side, 2325.990'S
15155.601'E, 10 m, dead coral heads]
Periclimenaeus crosnieri Cardoso & Young, 2007: 309; Figs 26-31. [Rocas Atoll, outer reef, 15-20 m, in
sponge]
Periclimenaeus devaneyi Bruce, 2010g: 380; Figs 1-5. [Kahe Point, Oahu, Hawaiian Islands]
Periclimenaeus diplosomatis Bruce, 1980b: 39; Figs 1-6. [Heron Island, Capricorn Islands, Queensland,
Australia, 2326.9'S 15155'E, low water]
Periclimenaeus djiboutensis Bruce, 1970c: 307. [Jibouti, French Somaliland]
Periclimenaeus fawatu Bruce, 2006e: 33; Figs 1-6. [of Fungu Fawatu, W coast of Unguja, Zanzibar, 33-36.5 m]
Periclimenaeus garthi Bruce, 1976d: 443; Figs 2-4. [Dunidu Island, Male Atoll]
Periclimenaeus gorgonidarum (Balss, 1913b)
= Periclimenes gorgonidarum Balss, 1913b: 236. [full description in Balss, 1914b, type locality therein
as Sagamibai, bei Miski, an Gorgoniden, 20-30 m Tiefe]
Periclimenaeus hancocki Holthuis, 1951b: 97; Plate 29. [Pias Bay, Panama, 32 fms]
Periclimenaeus hebedactylus Bruce, 1970c: 308. [Makunduchi, Zanzibar, 50 fms]
Periclimenaeus hecate (Nobili, 1904)
= Coralliocaris hecate Nobili, 1904: 232. [Djibouti]
Periclimenaeus heronensis Bruce, 2010f: 21; Figs 1-5. [Heron Island reef, north-eastern side, 2329.990'S
15155.601'E, 12 m, dead Acropora]
Periclimenaeus holthuisi Bruce, 1969b: 159. [Banda, Moluccas, Indonesia, 9 fms]
Periclimenaeus jeancharcoti Bruce, 1991a: 371; Figs 50-55. [New Caledonia, 2131'S 16621'E, 375-450 m]
Periclimenaeus kotae Bruce, 2005c: 325; Figs 1-3. [Australia, Western Australia, Ashmore Reef, outer
slope, 6-16 m]
Periclimenaeus leptodactylus Fujino & Miyake, 1968: 90; Figs 3-5. [Kasari-cho, Amami-oshima Island]
Periclimenaeus lobiferus Bruce, 1978a: 260; Figs 30-35. [Mozambique Channel, 1521.7'S 4612.6'E, 80-85 m]
Periclimenaeus manihinei Bruce, 1976e: 138; Figs 29-30. [Baie St. Anne, Praslin]
Periclimenaeus matherae Bruce, 2005c: 331; Figs 4-8. [Australia, Western Australia, Ashmore Reef, West
Laggon entrance, 2013.26'S 122059.28'E, 4-7 m]
Periclimenaeus maxillulidens (Schmit, 1936)
= Periclimenes maxillulidens Schmit, 1936: 371; Plate 13. [Bonaire, in the Lac (entrance), about 4f]
Periclimenaeus minutus Holthuis, 1952c: 134; Figs 57-59. [Banda anchorage, 18-36 m]
Periclimenaeus mortenseni Bruce, 1993b: 829; Figs 1-2. [Toeal, Kei-Doelah Islands, 2 m]
Periclimenaeus myora Bruce, 1998c: 394; Figs 4-5. [Myora Reef, North Stradbroke Island, Queensland,
2729'S 15325'E, 3 m]
Periclimenaeus nobilii Bruce, 1975c: 1577; Figs 13F, 14. [Red Sea]
Periclimenaeus nielbrucei Bruce, 2006f: 2; Figs 1-6. [Wreck Island, Capricorn Islands, Queensland]
Periclimenaeus nufu uri, Hork & Hoc, 2009: 454; Figs 1-5. [Bai Tr site, Lon Island, 1236'25''N
10919'58''E, Van Phong Bay, Vietnam, South China Sea, 5 m, coral rubble]
Periclimenaeus orbitocarinatus Fransen, 2006a: 732; Figs 13-15. [Loyalty Islands, 2022.25'S 16610.00'E, 21
m, in Lissoclinum verrilli]
Periclimenaeus orontes Bruce, 1986d: 151; Figs 1B, 6-10. [Orontes Reef, Port Essington, 1103.6'S 13205.0'E,
3 m]
Periclimenaeus pachydentatus Bruce, 1969b: 162. [Great Barrier Reef, Australia, 1412'N 14248'E, 19 fms]
Periclimenaeus pachyspinosus Marin, 2007a: 230-235; Figs 12-15. [South China Sea, Vietnam, Nhatrang
Bay, Nok Island, 15-20 m]
Periclimenaeus pacifcus Holthuis, 1951b: 85; Plate 25. [Pias Bay, Panama, 32 fms]
Periclimenaeus palauensis Miyake & Fujino, 1968: 417; Fig. 5. [Ngadark Reef]
Periclimenaeus pearsei (Schmit, 1932b)
= Coralliocaris pearsei Schmit, 1932b: 123; Fig. 1. [in the vicinity of the Tortugas Marine Laboratory
of the Carnegie Institution, from Spongia ofcinalis]
Periclimenaeus pectinidactylus uri, Hork & Sandford, 2009: 32; Figs 1-4, 5B-E. [of Southwater Caye,
Belizean Barrier Reef, SE of Dangriga, Belize, Caribbean Sea, 16.4 m, from an unidentifed sponge
(Smenospongia sp.?)]
Periclimenaeus perlatus (Boone, 1930b)
= Coralliocaris perlatus Boone, 1930b: 45; Fig. 8. [Gonave Bay, Haiti]
Periclimenaeus quadridentatus (Rathbun, 1906)
= Coralliocaris quadridentata Rathbun, 1906: 920; Fig. 69; Plate 24, fg. 1. [Auau Channel, 28-43 fms]
= Coralliocaris rathbuni Borradaile, 1917: 385. [nomen novum for Coralliocaris quadridentata Rathbun,
1906]
Periclimenaeus rastrifer Bruce, 1980a: 27; Figs 12, 13a-b. [Ilot Matre, Nouma, 20 m]
Periclimenaeus rhodope (Nobili, 1904)
= Coralliocaris (Onycocaris) rhodope Nobili, 1904: 233. [Djibouti]
Periclimenaeus robustus Borradaile, 1915b
= Periclimenus robustus Borradaile, 1915b: 213. [Amirante Islands]
Periclimenaeus serrula Bruce & Coombes, 1995: 111; Figs 3-5. [Orontes Reef, 114.5'S 1324.8'E, 11-12 m]
Periclimenaeus schmiti Holthuis, 1951b: 90; Plate 27. [Tortugas, Florida] (Fig. 39)
Periclimenaeus solitus Bruce & Coombes, 1995: 115; Figs 6-7. [Orontes Reef, 1103.6'S 13205.0'E, 3 m]
Fig. 39. Periclimenaeus schmiti Holthuis, 1951. Photo by Arthur Anker.
Periclimenaeus spinosus Holthuis, 1951b: 113; Plate 35. [Costa Rica, near South Viradores Islands, Port
Culebra, shallow water]
Periclimenaeus spongicola Holthuis, 1952c: 137; Figs 60-62. [Java Sea, 441'S 1132'E, 28-32 m, in sponge]
Periclimenaeus storchi Bruce, 1989a: 181; Fig. 5. [Cuaming Island, N of Bohol Island, Visayan Sea]
Periclimenaeus stylirostris Bruce, 1969b: 167. [South China Sea, 2034.0'N 11330.5'E 2030.3'N 11329.0'E,
49-50 fms]
Periclimenaeus tchesunovi uri, 1990a: 615; Figs 1-2. [Genego Islet, North Nilandu Atoll, Maldive Is-
lands, 20 m]
Periclimenaeus tridentatus (Miers, 1884b)
= Coralliocaris ? tridentata Miers, 1884b: 294; Plate 32, fg. C. [Thursday Island, 4-6 fms]
Periclimenaeus trispinosus Bruce, 1969b: 169. [Mkokotoni, N. end of Zanzibar Island, East Africa, 14-15 fms]
Periclimenaeus tuamotae Bruce, 1969b: 170. [Mururoa Island, Tuamotu Archipelago, Polynesia]
Periclimenaeus uropodialis Barnard, 1958: 18; Fig. 6. [Delagoa Bay]
Periclimenaeus usitatus Bruce, 1969b: 172. [Indian Ocean, 746'48"S 3942'36"E, 11 fms]
Periclimenaeus wilsoni (Hay, 1917)
= Coralliocaris wilsoni Hay, 1917: 71. [20 miles of Beaufort, N. C., in 10 to 20 fms]
Periclimenaeus wolf Bruce, 1993b: 833; Figs 4-5. [2320'N 11830'E, southern Tai-Wan Hai-Hsia, 31 m]
Periclimenaeus zanzibaricus Bruce, 1969b: 174. [Uroa, east coast of Zanzibar Island, East Africa, litoral]
Periclimenaeus zarenkovi uri, 1990a: 620; Figs 3-4. [Genego Islet, North Nilandu Atoll, Maldive Islands;
0.7 m depth]
Periclimenella Bruce, 1995
= Periclimenella Bruce, 1995 (type species Palmon Petithouarsii Audouin, 1826, by original designa-
= Periclimenella uri & Bruce, 1995 (type species Palmon Petithouarsii Audouin, 1826, by original
Periclimenella petithouarsii (Audouin, 1826)
= Palmon Petithouarsii Audouin, 1826: 91 (accompanying fgures in Savigny, 1826). [Egypt]
= A.[nchistia] inaequimana Heller, 1861: 28. [im rothen Meere; fully described in Heller, 1862c]
Periclimenella spinifera (De Man, 1902)
= Periclimenes petithouarsii var. spinifera De Man, 1902: 824. [Ternate; Pulo Edam; Bai von Batavia;
Amboina; Tahiti]
Periclimenes O.G. Costa, 1844a
= Pelias Roux, 1831 (type species A.[lpheus] amethystea Risso, 1827, designated by Holthuis, 1955b,
gender masculine; a junior invalid homonym of Pelias Merrem, 1820 (Reptilia))
= Periclimenes O.G. Costa, 1844a (type species Periclimenes insignis O.G. Costa, 1844a, by mono-
typy, gender masculine; name placed on the Ofcial List of Generic Names in Zoology in Opinion
470 in 1957)
= Anchistia Dana, 1852a (type species Anchistia gracilis Dana, 1852a, designated by Kingsley, 1880,
gender feminine)
= Dennisia Norman, 1861 (type species Dennisia sagitifera Norman, 1861, by monotypy, gender
feminine)
= Ancylocaris Schenkel, 1902 (type species Ancylocaris brevicarpalis Schenkel, 1902, by monotypy,
gender feminine)
= Cristiger Borradaile, 1915b (type species Periclimenes (Cristiger) commensalis Borradaile, 1915b, des-
ignated by Holthuis, 1955b, gender masculine)
Periclimenes acanthimerus Bruce, 2006g: 704; Figs 1-6. [New Caledonia, 2152.3'S 16647.9'E, 174 m]
Periclimenes aegylios Grippa & dUdekem dAcoz, 1996
= Periclimenes sagitifer aegylios Grippa & dUdekem dAcoz, 1996: 406; Fig. 1, 2b; Plate 1b. [west-
ern Italy, Giglio Island, Punta della Secca, 4221'N 1054'E, 10 m, associated with Anemonia vi-
ridis]
Periclimenes afnis (Zehntner, 1894)
= Palmonella afnis Zehntner, 1894: 208. [Amboine, sur Actinometra]
= Periclimenes (Falciger) brocketi Borradaile, 1915b: 212. [Mal Atoll, Maldive Islands, on brown
crinoid; lectotype designation by Bruce, 1978c]
Periclimenes albatrossae Chace & Bruce, 1993: 100; Fig. 20. [Philippines, South China Sea of western Lu-
zon, 1633'52"N 11952'54"E, 315 m]
Periclimenes albolineatus Bruce & Coombes, 1997: 305; Figs 1-2. [Lee Point, Darwin Harbour, 1219.0'S
13052.5'E, ELWST]
Periclimenes alcocki Kemp, 1922
= Periclimenes (Periclimenes) alcocki Kemp, 1922: 154; Figs 21-24. [Laccadive Sea, 934'57"N 7536'30"E,
406 fms]
Periclimenes aleator Bruce, 1991a: 315; Figs 10-14. [Loyalty Islands, 2053.0'S 16717.0'E, 570-610 m]
Periclimenes alexanderi Li, 2008: 219; Figs 7-10. [Marquesas Islands, Nuku Hiva, 845.1'S 14015.1'W, 108-
112 m]
Periclimenes amethysteus (Risso, 1827)
= A.[lpheus] amethystea Risso, 1827: 77; Plate 4, fg. 16. [golfe de Nice, rochers peu profonds]
= Periclimenes insignis O.G. Costa, 1844a: 291. [Napoli]
Periclimenes andresi Macpherson, 1988: 52; Figs 1-4. [1715'S 1127'E, 185 m]
Periclimenes antipathophilus Spote, Heard & Bubucis, 1994: 214; Figs 1-6. [Pine Cay, Turks and Caicos
Islands, near 2153'N 7205'W, 37-40 m, from the antipatharian Antipathes gracilis]
Periclimenes batei (Borradaile, 1917)
= Palmonella batei Borradaile, 1917: 358. [Philippine Is.]
Periclimenes boucheti Li, Mitsuhashi & Chan, 2008: 386; Fig. 1C, 2-4, 5A-B, 16B-D. [842.4'N 12313.4'E,
123 m]
Periclimenes bowmani Chace, 1972: 32; Figs 6-7. [Saint Lucia Island, reef south of Marigot Harbour, 2-3 m]
Periclimenes brevicarpalis (Schenkel, 1902)
= Ancylocaris brevicarpalis Schenkel, 1902: 563; Plate 13, fgs 21a-m. [Makassar]
= Palmonella amboinensis Zehntner, 1894: 206; Plate 9, fgs 27-27a. [Amboine]
= Palaemonella aberrans Nobili, 1904: 234. [Djibouti]
= Harpilius latirostris Lenz, 1905: 380; Plate 47, fgs 14-14b. [Zanzibar, Kokotoni, Bawi]
= Periclimenes potina Nobili, 1905a: 159. [Ctes dArabie, sur une algue brune fotante]
= Periclimenes hermitensis Rathbun, 1914: 655; Plate 1, fgs 1-3. [Hermite, under rock]
Periclimenes brevinaris Nobili, 1906c: 42; Plate 3, fg. 7-7a. [nomen novum for Periclimenes Borradailei No-
bili, 1905a nec Rathbun, 1904]
Periclimenes brevirostris Bruce, 1991a: 322; Figs 15-20. [2205.8'S 16710.3'E, Isle of Pines, 500-550 m]
Periclimenes brucei uri, 1990b: 1; Figs 1-2. [Indian Ocean, the Maldive Islands, North Nilandu Atoll,
Genego Island, 52 m]
Periclimenes burrup Bruce, 2007d: 113; Figs 5-7. [Burrup Peninsula, 2031.59'S 11652.09'E, 11.0 m]
Periclimenes calcaratus Chace & Bruce, 1993: 104; Fig. 21. [Philippines, Albay Gulf, east of southern Lu-
zon, 1312'N 12349'18"E, 267 m]
Periclimenes canalinsulae Bruce & Coombes, 1997: 309; Figs 3-4. [Channel Island, 1233.4'S 13052.3'E,
ELWS]
Periclimenes cannaphilus Komai, Nemoto & Tsuchida, 2010: 800; Figs 2-6. [Kasuga 2 Seamount, 2136.574'N
14238.175'E, 392 m]
Periclimenes carinidactylus Bruce, 1969a: 254. [Botle and Glass Rocks, Port Jackson, Sydney Harbour,
Australia, 20 feet]
Periclimenes chacei Li, Bruce & Manning, 2004: 542; Figs 27-28. [Xincun, Hainan Island, 1-3 m, with living
corals]
Periclimenes colemani Bruce, 1975d: 488; Figs 1-7. [Wistari Reef, Heron Island, Capricorn Group, Queens-
land, Australia, 12 m]
Periclimenes colesi De Grave & Anker, 2009: 141; Figs 1-26. [Black Coral Wall dive site, Utila, Bay Islands,
Honduras, 1605.027'N 8654.944'W, 15 m, inside osculum of Callyspongia vaginalis]
Periclimenes commensalis Borradaile, 1915b
= Periclimenes (Cristiger) commensalis Borradaile, 1915b: 211. [Torres Strait, on Comanthus annulatus]
Periclimenes compressus Borradaile, 1915b
= Periclimenes (Falciger) compressus Borradaile, 1915b: 212. [Saya de Malha]
Periclimenes coriolis Bruce, 1985b: 234; Figs 4-7. [1401.0'N 12017.1'E, 186-184 m]
Periclimenes crinoidalis Chace, 1969: 251; Figs 1-2. [Jan Thiel Beach, Curaao, Netherlands Antilles, 38 m,
associated with Nemaster grandis]
Periclimenes cristimanus Bruce, 1965: 487; Figs 1-2. [112.7'N 10343.65'E, Pulau Sudong, near Pulau Salu,
Singapore]
Periclimenes crosnieri Li & Bruce, 2006: 681; Figs 16-18. [Indonesia, Tanimbar Islands 754'S 13247'E, 302-
305 m]
Periclimenes curvirostris Kubo, 1940b
= Periclimenes (Periclimenes) curvirostris Kubo, 1940b: 35; Figs 3-5. [Kumano-nada, of Mie Prefecture,
about 170 fms]
Periclimenes dardanicola Bruce & Okuno, 2006: 368; Figs 1-4. [Japan. Kuryo, Numazu, Izu Peninsula, Su-
ruga Bay, Honshu, 3501.2'N 13849.9'E, 21 m]
Periclimenes delagoae Barnard, 1958: 14; Fig. 4-B. [Delagoa Bay, in coral]
Periclimenes dentidactylus Bruce, 1984: 7; Figs 1-6. [031.4'N 11750.1'E, 592-595 m]
Periclimenes difcilis Bruce, 1976e: 111; Figs 15-17. [Baie St. Anne, Praslin]
Periclimenes digitalis Kemp, 1922
= Periclimenes (Ancylocaris) digitalis Kemp, 1922: 224; Figs 65; Plate 8, fg. 12. [of Viper Island, Port
Blair, Andamans, 3-5 fms]
Periclimenes diversipes Kemp, 1922
= Periclimenes (Ancylocaris) diversipes Kemp, 1922: 179; Figs 36-39. [Kilakarai, Gulf of Manaar]
Periclimenes elefherioui Koukouras & Trkay, 1996: 136; Figs 1-3. [Aegean Sea, between Aspronisi and
Thera, 3622'20''N 2521'20''E, 40-73 m depth]
Periclimenes exederens Bruce, 1969a: 255. [South China Sea, 2036.0'N 11354.2'E 2038.8'N 11357.8'E,
47-48 fms]
Periclimenes fenneri Bruce, 2005d: 5; Figs 2-4, 8C. [West Norfolk Ridge, 3437.20'S 15859.03'E, 521-539 m]
Periclimenes fnlayi Chace, 1972: 35; Fig. 8. [Of Marigot Bay, St. Lucia, Windward Islands, 165 m, mollusk
traps]
Periclimenes forcipulatus Bruce, 1991a: 330; Figs 21-25. [Loyalty Islands, 2035.0'S 16654'E, 460 m]
Periclimenes foresti Bruce, 1981d: 201; Figs 10-11, 17c. [1400.0'N 12018.0'E to 1401.7'N 12020.2'E, 11
miles N. of Ambil island, 189-209 m]
Periclimenes forgesi Li & Bruce, 2006: 686; Figs 19-20. [New Caledonia, 2302.90'S 16657.97'E, 500-514 m]
Periclimenes foveolatus Bruce, 1981d: 196; Figs 6-9, 17a-b, 18b, e. [1401.0'N 12015.8'E to 1359.2'N
12018.8'E, 10 miles N. of Ambil Island, 191-188 m]
Periclimenes fujinoi Bruce, 1990c: 161; Figs 8-11, 39a-b. [Chesterfeld Islands, 2206.9'S 15924.6'E, 487-610 m]
Periclimenes gonioporae Bruce, 1989c: 149; Figs 1-3, 4A. [Ras Iwatine, Mombasa, 401.15'S 3943.78'E]
Periclimenes granulatus Holthuis, 1950c
= Periclimenes (Periclimenes) granulatus Holthuis, 1950c: 4; Fig. 1, Plate 1. [le travers de Bou-Haroun
(Dpart. DAlger), environ 100 m]
Periclimenes granulimanus Bruce, 1978a: 237; Figs 16-19. [Tany-Kely, NW coast near Nosy B, on grey
sand with foraminifera]
Periclimenes granuloides Hayashi in Baba, Hayashi & Toriyama, 1986: 103; Fig. 18; unnumbered plate.
[Tosa Bay, Japan, 130 m]
Periclimenes guarapari De Grave, 2008: 15; Figs 1-5. [Ilha Escalvada, in front of Guarapri City, Espirito
Santo, Brazil, approx. 2042'S 0424'W, 17 m, from Muricea fammea]
Periclimenes harringtoni Lebour, 1949a
= Periclimenes (Periclimenes) harringtoni Lebour, 1949a: 1110; Fig. 3. [Harrington Sound]
Periclimenes hertwigi Balss, 1913b: 235. [Sagamibai, 120 m]
= Periclimenes (Ancylocaris) gracilirostris Kubo, 1940b: 41; Figs 8-10. [Kumano-nada, of Mie Prefec-
ture, about 170 fms]
Periclimenes hongkongensis Bruce, 1969a: 259. [Rocky Harbour, Hong Kong, 2220.0'N 11421'E, 14
fms]
Periclimenes hydrothermophilus Hayashi & Ohtomi, 2001: 161; Figs 1-4. [Kagoshima Bay, 90-100 m, associ-
ated with Lamellibranchia satsuma (Polychaeta)]
Periclimenes imperator Bruce, 1967b: 53; Figs 23-25. [Tumbatu Island, west coast of Zanzibar Island, on
nudibranch, 3 f below low water spring tide level]
Periclimenes incertus Borradaile, 1915b
= Periclimenes (Cristiger) incertus Borradaile, 1915b: 210. [Maldive Islands]
= Periclimenes (Periclimenes) impar Kemp, 1922: 147; Figs 16, 17; Plate 3, fg. 1. [Port Blair, Andamans,
5 fms]
Periclimenes infraspinis (Rathbun, 1902a)
= Urocaris infraspinis Rathbun, 1902a: 903. [Gulf of California in Concepcion Bay, Lower California]
Periclimenes ingressicolumbi Berggren & Svane, 1989: 432; Figs 1-5. [San Salvador Island, 2402.91'N
7432.7'W, 1900f]
Periclimenes inornatus Kemp, 1922
= Periclimenes (Ancylocaris) inornatus Kemp, 1922: 191; Figs 43-46. [Port Blair, Andamans]
= Periclimenes paraornatus Bruce, 1979a: 207. [nomen nudum]
Periclimenes investigatoris Kemp, 1922
= Periclimenes (Periclimenes) investigatoris Kemp, 1922: 160; Fig. 26-27; Plate 5, fg. 6. [Persian Gulf,
2920'N 4847'E, 13 fms]
Periclimenes involens Bruce, 1996: 234; Figs 13, 28h. [Philippines, of Mindoro, 1231'N 12039'E, 92-97 m]
Periclimenes iridescens Lebour, 1949a
= Periclimenes (Periclimenes) iridescens Lebour, 1949a: 1112; Figs 4-5. [tow-net about 10 miles outside
Castle Roads over about 8-100 fms]
Periclimenes ischiospinosus Bruce, 1991b: 240; Figs 3b, 9-12. [New Caledonia, 2144'S 16632'E, 50 m]
Periclimenes josephi Li, 2008: 224; Figs 11-14. [Austral Islands, Neilson Reef, 2703.4'S 14604'W, 130-
140 m]
Periclimenes jugalis Holthuis, 1952c
= Periclimenes (Harpilius) jugalis Holthuis, 1952c: 67; Fig. 26. [anchorage of Djedan Island, east coast
of the Aru Islands, 13 m]
Periclimenes kallisto Bruce, 2008d: 89; Figs 1-6. [Keeper Reef, Queensland, 1844'S 14716'E, 38 m]
Periclimenes kempi Bruce, 1969a: 260. [Hurghada, Red Sea, Egypt, 2714'N 3850'E, 0.5 fms]
Periclimenes kornii (Lo Bianco, 1903)
= Anchistia Kornii Lo Bianco, 1903: 250; Plate 7, fg. 13. [a circa 12.5 chilometri da Punta Carena, dir-
ezone S.E., 1100 m; a circa 8.5 chilometri da Punta Tragera, direzione O.S.O., 1000 m; a circa 10
chilometri da Punta Carena, direzione E.N.E., 1000 m]
Periclimenes laccadivensis (Alcock & Anderson, 1894)
= Palmonella laccadivensis Alcock & Anderson, 1894: 157. [Investigator stn 124, Laccadive Sea,
1047'45''N 7240'20''E, 705 fms]
Periclimenes laevimanus uri, 2010: 107; Figs 1-6. [Vietnam, Nhatrang Bay, 1210'06''N 10917'45''E, 14 m,
on hydroid cf. Lytocarpia sp.]
Periclimenes latipollex Kemp, 1922
= Periclimenes (Periclimenes) latipollex Kemp, 1922: 150; Fig. 18; Plate 4, fg. 3. [Mergui Archipelago,
1215'20"N 9710'10"E, 62 fms]
Periclimenes lepidus Bruce, 1978a: 244; Figs 20-24. [NW coast of Madagascar near Nosy B, 40 m]
Periclimenes leptodactylus Bruce, 1991a: 338; Figs 26-30. [Loyalty Islands, 2037.8'S 16702.7'E, Lifou/Uva,
825-370 m]
Periclimenes leptopus Kemp, 1922
= Periclimenes (Ancylocaris) leptopus Kemp, 1922: 173; Figs 31-33. [Brigade Creek, Port Blair, Anda-
mans, 2-5 fms]
Periclimenes leptunguis Li, Mitsuhashi & Chan, 2008: 393; Figs 6-8, 17B. [849.9'N 12334.9'E, 437-443 m]
Periclimenes longimanus (Dana, 1852a)
= Anchistia longimana Dana, 1852a: 25. [Type locality not indicated]
Periclimenes longipes (Stimpson, 1860a)
= Urocaris longipes Stimpson, 1860a: 39. [insulam Ousima, prof 20 org]
Periclimenes loyautensis Li & Bruce, 2006: 700; Fig. 24 [Loyalty Islands, 2102.30'S 16731.60'E, 430 m]
Periclimenes macrophthalmus Fujino & Miyake, 1970a
= Periclimenes (Harpilius) macrophthalmus Fujino & Miyake, 1970a: 250; Figs 3-5. [3236.7'N 12742.8'E,
145 m]
Periclimenes madreporae Bruce, 1969a: 262. [Erskine Island, Capricorn Group, Great Barrier Reef, Queens-
land, Australia, 3-6 fms]
Periclimenes magnus Holthuis, 1951b
= Periclimenes (Harpilius) magnus Holthuis, 1951b: 52; Plate 15. [Gulf of Mexico, of Aransas, Texas,
2740'N 9634'W, 27.5 fms]
Periclimenes mahei Bruce, 1969a: 263. [North West Bay, Mah, Seychelles Islands, 436'15"S 5526'01"E,
1-2 fms]
Periclimenes maldivensis Bruce, 1969a: 264. [Suvadiva Atoll, Maldive Islands]
Periclimenes manihine Bruce, 2006h: 46; Figs 1-4. [of Ras Ngomeni, Kenya, 301'S 4022.5'E to 259.5'N
4022.6'E, 245-256 m]
Periclimenes mclaughlinae Fransen, 2006b: 380; Figs 1-6. [Philippines, Cebu Strait, W of Bohol, NW side of
Cabilao Island, La Estrella Dive Resort, 953.20'N 12345.53'E, 5 m, on Phyllacanthus imperialis]
Periclimenes mclellandi Heard & Spote, 1997: 39; Figs 1-5. [forereef slope west of Pine Cay, Turks and
Caicos Islands, British West Indies, on gorgonian Pseudopterogorgia americana, 29 m]
Periclimenes meyeri Chace, 1969: 255; Figs 3-4. [Jan Thiel Beach, Curaao, Netherlands Antilles, 24 m, as-
sociated with Nemaster?]
Periclimenes milleri Bruce, 1986e: 637; Figs 1-5. [2402.75'N 7432.53'W, 527 m]
Periclimenes murcielagensis Vargas, 2000: 17; Figs 1-5. [Isla San Pedrito, Archipilago de las Islas Mur-
cilago, Guanacaste, Costa Rica, 25 m]
Periclimenes nevillei Bruce, 2010h: 45; Figs 1-6. [Espiritu Santo, Vanuatu, 8 m]
Periclimenes ngi Li, Mitsuhashi & Chan, 2008: 397; Figs 9-11, 17C-D. [927.4'N 12349.4'E, 273-302 m]
Periclimenes nomadophila Berggren, 1994a: VII.3; Figs 1-5. [Inhaca Island, Mocambique]
= Periclimenes nomadophila Berggren, 1994b: 783; Figs 1-5. [Inhaca Island, Mocambique]
Periclimenes novafnis Bruce & Coombes, 1997: 315; Fig. 6. [of East Point, Darwin Harbour, 1225.92'S
13048.72'E, 8 m]
Periclimenes obscurus Kemp, 1922
= Periclimenes (Periclimenes) obscurus Kemp, 1922: 144; Figs 14-15. [Springhaven, Madras Harbor]
Periclimenes ordinarius Bruce, 1991a: 344; Figs 31-35. [New Caledonia, 1804.0'S 16327.5'E, 260 m]
Periclimenes ornatellus Bruce, 1979d: 219; Figs 4-6; Plate 1C-E. [Enewetak Atoll]
Periclimenes ornatus Bruce, 1969a: 266. [Lung Ha Wan, N.T., Hong Kong, 2218.5'N 11418.2'E, 2 fms]
Periclimenes paivai Chace, 1969: 259; Figs 5-7. [Canania, Estado de So Paulo, Brazil]
Periclimenes pandionis Holthuis, 1951b
= Periclimenes (Periclimenes) pandionis Holthuis, 1951b: 41; Plate 11. [Gulf Stream of Key West,
Florida, 2421'55"N 8158'25"W, 98 fms]
Periclimenes panglaonis Li, Mitsuhashi & Chan, 2008: 402; Figs 12-13, 14A-D, 17E. [938.8'N 12345.9'E,
418-477 m]
Periclimenes paralcocki Li & Bruce, 2006: 707; Fig. 27. [Tuvalu, Banc Bayonnaise, 1154'S 17932'W, 600 m]
Periclimenes paraleator Li & Bruce, 2006: 711; Fig. 28. [New Caledonia, 1856.3'S 16312.9'E, 520 m]
Periclimenes paraparvus Bruce, 1969a: 267. [South China Sea, 2028.2'N 11246.5'E to 2023.3'N 11252.2'E,
46-48 fms]
Periclimenes parvispinatus Bruce, 1990c: 154; Figs 3-6, 39d. [New Caledonia, S.W. Rcif Jouan, 200 m]
Periclimenes parvus Borradaile, 1898: 384. [New Britain]
Periclimenes patae Heard & Spote, 1991: 40; Figs 1-4. [fringing reef northwest of Pine Cay, Turks and
Caicos Islands, British West Indies, near 2153'N 07205'W, on gorgonian, 3-4 m]
Periclimenes pauper Holthuis, 1951b
= Periclimenes (Harpilius) pauper Holthuis, 1951b: 50; Plate 14. [Venezuela, Isla Cubagua, rocky
shore]
Periclimenes pectiniferus Holthuis, 1952c
= Periclimenes (Periclimenes) pectiniferus Holthuis, 1952c: 48; Figs 15-16. [Kabala dua Island, Borneo
Bank, 22 m]
Periclimenes pectinipes Bruce, 1991a: 351; Figs 36-40, 75. [New Caledonia, 2341.2'S 16800.5'E, Norfolk
Ridge, 280 m]
Periclimenes perlucidus Bruce, 1969a: 268. [South China Sea, 1606.5'N 11441.5'E to 1605.8'N 11438.2'E,
43-44 fms]
Periclimenes perryae Chace, 1942b
= Periclimenes (Periclimenes) perryae Chace, 1942b: 82; Plate 24. [of Sanibel Island, Florida, about 5.5
fms, on Astrophyton muricatum]
Periclimenes perturbans Bruce, 1978a: 253; Figs 25-26. [NW coast of Madagascar, near Nosy B, 40 m]
Periclimenes pholeter Holthuis, 1973a: 30; Figs 10-11; Plate 1, fg. 1. [Ras Muhammad Crack, Ras Muham-
mad, southern tip of Sinai Peninsula, in pool in crack in coral rock about 150 m from the sea]
Periclimenes platydactylus Li, 2008: 231; Figs 16-19. [Marquesas Islands, Nuku Hiva, 845.1'S 14015.1'W,
108-112 m]
Periclimenes platyrhynchus Bruce, 1991a: 358; Figs 41-44. [New Caledonia, 1904.0'S 16327.0'E, 260 m]
Periclimenes polynesiensis Li, 2008: 236; Figs 20-23. [Marquesas Islands, Nuku Hiva, 845.1'S 14015.1'W,
108-112 m]
Periclimenes poriphilus Bruce, 2010i: 53; Figs 2-6. [Heron Island reef, north-eastern side, 2325.990'S
15155.601'E, 10 m]
Periclimenes poupini Bruce, 1989d: 852; Figs 1-5, 6A. [Tubuai, 2319'S 14222'W, 430-520 m]
Periclimenes pseudalcocki Li & Bruce, 2006: 716; Fig. 30. [Indonesia, Kai Islands, 517'S 13250'E, 315-349 m]
Periclimenes rathbunae Schmit, 1924a: 70; Figs 5-6. [Spanish Port]
Periclimenes rectirostris Bruce, 1981d: 204; Figs 12-15. [1353.1'N 12008.9'E to 1353.3'N 12010.7'E, 3
miles NE. of town of Lubang, 134-129 m]
Periclimenes rex Kemp, 1922
= Periclimenes (Periclimenes) rex Kemp, 1922: 158; Fig. 25; Plate 5, fg. 5. [Ross Channel, Port Blair,
Andamans, 8 fms]
Periclimenes richeri Bruce, 1990c: 181; Figs 20, 39f. [New Caledonia, 2454.5'S 16823.3'E, 527 m]
Periclimenes ruber Bruce, 1982c: 197; Figs 3-5, 8f. [Bribie Passage, Pumicestone Channel, Queensland,
Australia, in 8 m depth, from crinoid host Zygometra microdiscus]
Periclimenes sagitifer (Norman, 1861)
= Dennisia sagitifer Norman, 1861: 278; Plate 13, fgs 8-13. [St. Catherines Bay, Jersey, 4 fms]
Periclimenes sandybrucei Mitsuhashi & Chan, 2009: 920; Figs 1-4. [2430.55'N 1225.78'E, 399-397 m]
Periclimenes sandyi De Grave, 2009: 830; Figs 1-4. [Crawl Cay, Bocas del Toro Archipelago, Caribbean
Panama, 0915.028'N 8207.897'W, 10 m, from unidentifed sponge]
Periclimenes sarkanae Bruce, 2007e: 61; Figs 1-5. [Fisherman Island, near mouth of Brisbane River, More-
ton Bay, 2722'S 15310'E, 0.2-0.5 m]
Periclimenes scriptus (Risso, 1822)
= A.[lpheus] scriptus Risso, 1822: 247. [mer de Nice]
? = Periclimenes elegans Gourret, 1884: 15; nec Anch.[istia] elegans Paulson, 1875. [nomen nudum]
= Urocaris de Mani Balss, 1916: 29; Fig. 10. [Franzsisch-Kongo, Set Cama, salzwasser]
Periclimenes sibogae Holthuis, 1952c
= Periclimenes (Harpilius) sibogae Holthuis, 1952c: 73; Figs 28-29. [Banda anchorage, 9-36 m]
Periclimenes sinensis Bruce, 1969a: 270. [Hong Kong]
= Periclimenes (Periclimenes) setoensis Fujino & Miyake, 1969a: 149; Figs 4, 5. [Shis-jima Island, Tan-
abe Bay, Wakayama Prefecture, 5 m]
Periclimenes soror Nobili, 1904: 232. [Djibouti]
? = Periclimenes parasiticus Borradaile, 1898: 384. [British New Guinea]
= Periclimenes (Cristiger) frater Borradaile, 1915b: 210. [Seychelles]
= Periclimenes bicolor Edmondson, 1935a: 10; Fig. 3. [Kaneohe Bay, Oahu, on Linckia multifora in shal-
low water]
Periclimenes tangeroa Bruce, 2005d: 12; Figs 5-7, 8D. [South Norfolk Ridge, 3409.14'S 17127.95'E, 242-254 m]
Periclimenes tenellus (Smith, 1882)
= Anchistia tenella Smith, 1882: 55; Plate 9, fgs 1-1b. [Blake stn 316, 3207'N 7837'05"W, 229 fms]
Periclimenes terangeri Bruce, 1998c: 387; Fig. 1. [Polka Point, North Stradbroke Island, Queensland,
2730'S 15323'E, 0.5 m]
Periclimenes thermohydrophilus Hayashi & Ohtomi, 2001: 161; Figs 1-4. [Kagoshima Bay, 90-100 m]
Periclimenes toloensis Bruce, 1969a: 273. [Ap Chau, Tolo Channel, Hong Kong, 5-15 fms]
Periclimenes tonga Bruce, 1988e: 23; Figs 1-5. [Nuapapa Island (southside), Vavau Group, Tonga, 5-50 f]
Periclimenes uniunguiculatus Bruce, 1990c: 167; Figs 12-15, 39e. [New Caledonia, 2306'S 16747'E, 540-
600 m]
Periclimenes vanellus Fransen, 2010: 242; Figs 1-6. [Indonesia, of Halmahera mainland, Teluk Dodinga,
Karang Galiasa Besar E, 050'45.6''N 12735'7.4''E, on Ianthella basta, 15 m]
Periclimenes vaubani Bruce, 1990c: 174; Figs 16-9, 38a-d, 39g. [New Caledonia, 2338'S 16742'S, 470 m]
Periclimenes veleronis Holthuis, 1951b
= Periclimenes (Harpilius) veleronis Holthuis, 1951b: 67; Plate 20, fgs a-h. [of beach at La Libertad, 4 fms]
Periclimenes vicinus Li, 2008: 242; Figs 24-27. [Austral Islands, Tubuai, 2321.4'S 14934.2'W, 600-1200 m]
Periclimenes watamuae Bruce, 1976f: 16; Figs 5-6. [Watamu Park, Kenya, 322.0'S 4000.5'E, 2 m, from al-
cyonarian; information from NCB-Naturalis collection label]
Periclimenes wirtzi dUdekem dAcoz, 1996a: 133; Figs 1-7. [Madeira, small seamount in front of the bay
of Machico, from Antipathes bushes at 30-40 m]
Periclimenes yaldwyni Holthuis, 1959b
= Periclimenes (Harpilius) yaldwyni Holthuis, 1959b: 197. [nomen novum for Periclimenes batei
Holthuis, 1950a]
= Brachycarpus audouini Spence Bate, 1888: 798; Plate 129, fg. 5. [Challenger stn 167A, 414'S 17419'E,
of New Zealand, 10 fms]
= Periclimenes batei Holthuis, 1950a: 22. [nec Palmonella batei Borradaile, 1917]
Periclimenes yucatanicus (Ives, 1891)
= Palmonella Yucatanica Ives, 1891: 183; Plate 5, fg.8. [of Progreso, 20f]
Periclimenes zanzibaricus Bruce, 1967b: 62; Figs 26-29. [Fawatu Reef, of west coast of Zanzibar Island, on
echinoid at low water spring tide level on sand fats]
Periclimenes zevinae uri, 1990b: 4; Figs 3-4. [Indian Ocean, the Maldive Islands, North Nilandu Atoll,
Genego Island, 50 m]
Periclimenoides Bruce, 1990f
= Periclimenoides Bruce, 1990f (type species Periclimenaeus odontodactylus Fujino & Miyake, 1968, by
Periclimenoides odontodactylus (Fujino & Miyake, 1968)
= Periclimenaeus odontodactylus Fujino & Miyake, 1968: 85; Figs 1-2. [Ushibuka, Amakusa Island,
west of Kyushu, Japan]
Periclimenoides socotrae Bruce, 2006a: 22; Figs 6-9. [1240.264'N 5327.204'E, SW of Qualansiyah, NW-
coast, 5-7 m]
Philarius Holthuis, 1952c
= Philarius Holthuis, 1952c (type species Harpilius Gerlachei Nobili, 1905a, by original designation,
gender masculine)
Philarius albimaculatus Marin & Anker, 2011: 22; Figs 16-18, 19E-F. [Madagascar, Nosy-B W side of Saka-
tia, 13.2989S 48.1472E, 3-8 m]
Philarius gerlachei (Nobili, 1905a)
= Harpilius Gerlachei Nobili, 1905a: 160. [Golfe Persique]
Philarius imperialis (Kubo, 1940c)
= Harpilius imperialis Kubo, 1940c: 1; Figs 1-3. [Nankin-Hama, Haha-zima, Bonin Islands]
Philarius lifuensis (Borradaile, 1898)
= Periclimenes lifuensis Borradaile, 1898: 384. [Loyalty Islands]
Philarius minor Marin & Anker, 2011: 13; Figs 13-15, 19C-D. [Australia, Great Barrier Reef of S Queens-
land, Heron Island, 23.47262S 151.95988E, 14-16 m]
Philarius polynesicus Marin & Anker, 2011: 6; Figs 4-8, 9A-E. [French Polynesia, Society Islands, Moorea,
S of Vaiare Pass, 17.5303S 149.7621W, 5-6 m]
Philarius rufus Marin & Anker, 2011: 13; Figs 10-14, 19A-B. [Australia, Great Barrier Reef of S Queens-
land, Heron Island, 23.45322S 151.90045E, 24-26 m]
Phycomenes Bruce, 2008e
= Phycomenes Bruce, 2008e (type species Phycomenes zostericola Bruce, 2008e, by original designation
Phycomenes cobourgi (Bruce & Coombes, 1995)
= Periclimenes cobourgi Bruce & Coombes, 1995: 125; Figs 10-11. [Table Head, 3 m, on gorgonian
Rumphella aggregata]
Phycomenes indicus (Kemp, 1915)
= Urocaris indica Kemp, 1915: 275; Fig. 26; Plate 13, fg. 9. [Chilka Lake; Ennur and Adyar River near
Madras; fringing coral-reef at Kilakarai, northern end of Gulf of Manaar]
Phycomenes siankaanensis (Martnez-Mayn & Romn-Contreras, 2006)
= Periclimenes siankaanensis Martnez-Mayn & Romn-Contreras, 2006: 33; Figs 1-3. [Cayo Cule-
bras, Baha de la Ascensin, Quintana Roo, Mxico, 1 m]
Phycomenes sulcatus (uri, Hork & Marin, 2008)
= Periclimenes sulcatus uri, Hork & Marin, 2008: 36; Figs 1-6. [S of Nhatrang Bay, Vietnam,
1209'42.0''N 10912'13.1''E, 0.1-0.2 m]
Phycomenes zostericola Bruce, 2008e: 221; Figs 1-7. [Loders Creek, Labrador, Queensland]
Fig. 40. Platypontonia hyotis Hipeau-Jacquote, 1971. Photo by Arthur Anker.
Pinnotheronia Marin & Paulay, 2010
= Pinnotheronia Marin & Paulay, 2010 (type species Pinnotheronia rumphiusi Marin & Paulay, 2010, by
Pinnotheronia rumphiusi Marin & Paulay, 2010: 38; Figs 1-6. [Pacifc Ocean, Palau, Koror Island, Iwayama
Bay, in mantle cavity of Periglypta crispata, 0-10 m]
Platycaris Holthuis, 1952c
= Platycaris Holthuis, 1952c (type species Platycaris latirostris Holthuis, 1952c, by monotypy, gender
feminine)
Platycaris latirostris Holthuis, 1952c: 173; Figs 85-86. [Ende, Flores]
Platypontonia Bruce, 1968b
= Platypontonia Bruce, 1968b (type species Pontonia ? brevirostris Miers, 1884a, by original designa-
Platypontonia brevirostris (Miers, 1884a)
= Pontonia? brevirostris Miers, 1884a: 562; Plate 51, fgs Bb-b'''. [Seychelles, 12 fms, in clamp shells
(bivalves?)]
Platypontonia hyotis Hipeau-Jacquote, 1971: 126; Figs 1-7. [Madagascar, S of Tular, outer slope of Banc
de Belosa, in Pycnodonta hyotis; according to Fransen, Holthuis & Adema, 1997] (Fig. 40)
= Platypontonia pterostreae Suzuki, 1971: 5; Figs 3-4; Plate 3. [Hatsu-shima, Sagami Bay]
Plesiomenaeus Bruce, 2009b
= Plesiomenaeus Bruce, 2009b (type species Plesiomenaeus poorei Bruce, 2009b, by original designation
Plesiomenaeus poorei Bruce, 2009b: 26; Figs 1-7. [Chukwani, Unguja, Zanzibar, 613'60''S 3013'00''E, at
low water spring tide]
Plesiopontonia Bruce, 1985b
= Plesiopontonia Bruce, 1985b (type species Plesiopontonia monodi Bruce, 1985b, by original designa-
Plesiopontonia monodi Bruce, 1985b: 250; Figs 13-17. [1349.6'N 12051'E, 299-320 m]
Pliopontonia Bruce, 1973c
= Pliopontonia Bruce, 1973c (type species Pliopontonia furtiva Bruce, 1973c, by original designation
Pliopontonia furtiva Bruce, 1973c: 99; Figs 1-5; Plate 1. [Ras Iwatine, Mombasa, Kenya, 400.55'S 3944.17'E,
1 m]
Pliopontonia harazakii Okuno, 2009b: 106; Figs 1-4, 5A-B, 6. [Onoaida, Yaku-shima Island, Osumi Islands,
Japan, 3014.1'N 13033.2'E, 3 m]
Pontonia Latreille, 1829
= Alciope Rafnesque, 1814 (type species Alciope heterochelus Rafnesque, 1814, gender feminine; an
invalid senior subjective synonym of P.[ontonia] favomaculata Heller, 1864; name suppressed for the
purposes of the Principle of Priority, but not for those of the Principle of Homonymy, placed on the
Ofcial Index of Rejected and Invalid Generic Names in Zoology in Opinion 522 in 1958)
= Pontonia Latreille, 1829 (type species Palaemon pinnophylax Oto, 1821, gender feminine; designated
under the plenary power of the ICZN and placed on the Ofcial List of Generic Names in Zoology
in Opinion 378 in 1956)
Pontonia chimaera Holthuis, 1951b: 125; Plate 39. [Panama, Archipielago de las Perlas, north side of Pedro
Gonzalez Island, west of Cocal]
Pontonia domestica Gibbes, 1850: 196. [coast of South Carolina]
= Pontonia occidentalis Gibbes, 1848: 16. [South Carolina; nomen nudum]
Pontonia longispina Holthuis, 1951b: 128; Plate 40. [Lower California, Mexico, Angel de la Guardia Is-
land, Puerto Refugio, shore]
Pontonia manningi Fransen, 2000: 101; Figs 1-3, 4a-f. [Caribbean Sea, 25 miles N of Margarita Island, 36.5 m]
Pontonia margarita Verrill, 1869: 245. [Bay of Panama, in Pearl Oyster (Margaritophora fmbriata)]
= Coralliocaris Camerani Nobili, 1901b: 3. [Isola Flamenco]
Pontonia mexicana Gurin-Mneville, 1855 [in Gurin-Mneville, 1855-1856]: xix; Plate 2, fg. 12. [Mexico;
see Holthuis, 1951b]
= Pontonia grayi Rathbun, 1901: 122 (partim); Fig. 5. [between San Antonio Bridge and San Geronimo,
San Juan, Porto Rico]
Pontonia panamica Marin & Anker, 2008: 503; Figs 1-7. [Panama, Playa Venao, 10 km west of Panama City,
lower rocky intertidal, in ascidian, Ascidia cf. interrupta]
Pontonia pilosa Fransen, 2002: 147; Figs 91-99; Plate 4. [East Atlantic, Cape Verde Islands, Sao Tiago Is-
land, Tarrafal, from Pseudochama radians]
Pontonia pinnae Lockington, 1878a: 163. [interior of the shell of the common Pinna, Gulf of California,
Angelas Bay, Mulege Bay, San Jos Island] (Fig. 41)
Pontonia pinnophylax (Oto, 1821)
= Palaemon pinnophylax Oto, 1821: 12. [Neapoli in Pinna nobili marem; fully described in Oto, 1828
as Alpheus pinnophylax]
= A.[lpheus] Tyrhenus Risso, 1816: 94; Plate 2, fg. 2. [environs de Nice, dans les valves du jambon-
neau marin]
= Pontonia parasitica Roux, 1831: 26. [sur les cotes de More, dans linterieur de la Pinna Nobilis]
= Pontonia custos Gurin, 1832: 36; Plate 37, fg. 1. [Sapience dans la Pinne marine (Pinna nobilis)]
= Pontonia heterochelis Gurin, 1832: 37. [see Holthuis, 1947b]
Pontonia simplex Holthuis, 1951b: 135; Plate 42. [Jalisco, Mexico, Tenacatita Bay, lagoon]
Fig. 41. Pontonia pinnae Lockington, 1878. Photo by Arthur Anker.
Pontonides Borradaile, 1917
= Pontonides Borradaile, 1917 (type species Pontonia maldivensis Borradaile, 1915b, by monotypy, gen-
der masculine; name placed on the Ofcial List of Generic Names in Zoology in Opinion 470 in 1957)
Pontonides ankeri Marin, 2007b: 3; Figs 1-3, 10B, 11F-G. [Vietnam, Nhatrang Bay, Nok Island, 15 m]
Pontonides asperulatus Bruce, 2005b: 359; Figs 13-16, 22E. [Horseshoe Reef, Loloata Island, 16 m]
Pontonides loloata Bruce, 2005b: 367; Figs 17-21, 22F. [Loloata Island]
Pontonides maldivensis (Borradaile, 1915b)
= Pontonia maldivensis Borradaile, 1915b: 213. [Fadifolu Atoll, Maldive Islands]
Pontonides sibogae Bruce, 2005b: 377. [Sape Strait, Indonesia, 2330'S 1194.6'E, 70 m]
Pontonides tatianae Marin, 2007b: 13; Fig. 8. [Vietnam, Nhatrang Bay, Dun Island, 40 m]
Pontonides unciger Calman, 1939: 213; Figs 6-7. [Red Sea, 1331'00''N 4231'00''E, 55 m]
Pontoniopsides Bruce, 2005e
= Pontoniopsides Bruce, 2005e (type species Pontoniopsis paulae Gore, 1981, by original designation
Pontoniopsides paulae (Gore, 1981)
= Pontoniopsis paulae Gore, 1981: 139; Fig. 1. [Carysfort Reef, of Key Largo, Monroe County, Florida,
2510.30'N 8012.82'W, 62.5 m]
Pontoniopsis Borradaile, 1915b
= Pontoniopsis Borradaile, 1915b (type species Pontoniopsis comanthi Borradaile, 1915b, by monotypy,
1959)
Pontoniopsis comanthi Borradaile, 1915b: 213. [Torres Straits, on Comanthus]
Poripontonia Fransen, 2003
= Poripontonia Fransen, 2003 (type species Poripontonia dux Fransen, 2003, by original designation
Poripontonia dux Fransen, 2003: 130; Figs 1-6. [Tanjung Benoa, Loloan Benoa, 0845'46''S 11514'01''E, to 25 m]
Poripontonia cornuta Marin, 2007a: 235; Figs 16-18. [South China Sea, Vietnam, Nhatrang Bay, Nok Is-
land, 10 m]
Propontonia Bruce, 1969c
= Propontonia Bruce, 1969c (type species Propontonia pellucida Bruce, 1969c, by original designation
Propontonia pellucida Bruce, 1969c: 142; Figs 1-5. [Remire Island, Amirante Islands, Seychelles, 0504'S
5322'E, 1.5 m]
Pseudoclimenes Bruce, 2008c
= Pseudoclimenes Bruce, 2008c (type species Pseudoclimenes holthuisi Bruce, 2008c, by original desig-
nation and monotypy, gender masculine)
Pseudoclimenes holthuisi Bruce, 2008c: 17; Figs 10-13. [Jurien Bay, 2948'25''S 11425'52''E to 2948'53''S
11425'55''E, 113-114 m]
Pseudocoutierea Holthuis, 1951b
= Pseudocoutirea Holthuis, 1951b (type species Pseudocoutirea elegans Holthuis, 1951b, by original
Pseudocoutierea antillensis Chace, 1972: 43; Fig. 11. [Saba Bank, 1728'N 6313'W, dredged in 13 m]
Pseudocoutierea conchae Criales, 1981: 174; Figs 6-9. [Baha Concha, Colombia, 1118'N 7410'W, 15 m]
Pseudocoutierea dotae De Grave, 2007b: 30; Figs 1-5. [Hospital Point, Cayo Solarte, Bocas Province, Pana-
ma, 0920.016'N 08213.133'W, 15 m, on Stichopathes lutkeni]
Pseudocoutierea edentata Criales, 1981: 168; Figs 2-5. [Baha Concha, Colombia, 1118'N 7410'W, 18 m]
Pseudocoutirea elegans Holthuis, 1951b: 182; Plate 57. [southern California, Santa Catalina Island, mile
E of Long Point, 45-50 fms]
Pseudocoutierea wirtzi dUdekem dAcoz, 2000: 70; Figs 1-12. [Cape Verde Islands, So Tiago Island, 20-30
m, on Leptogorgia gaini]
Pseudopontonia Bruce, 1992d
= Pseudopontonia Bruce, 1992d (type species Pontonia minuta Baker, 1907, by original designation
Pseudopontonia minuta (Baker, 1907)
= Pontonia minuta Baker, 1907: 189; Plate 24, Figs 9-12. [unidentifed locality in South Australia; see
Bruce, 1992d]
Pseudopontonides Heard, 1986
= Pseudopontonides Heard, 1986 (type species Neopontonides principes Criales, 1980, by original desig-
Pseudopontonides plumosus Sniders & Fransen, 2010: 7; Figs 1-6. [south coast, Otrobanda, harbour, Supe-
rior Producer, Curaao, Lesser Antilles, 1205'21.5''N 06856'35.5''W, on Antipathes spec., depth 29
m]
Pseudopontonides principis (Criales, 1980)
= Neopontonides principis Criales, 1980: 75; Figs 25-29. [Awa di Oostpunt, Curaao, 18 m]
Pseudoveleronia Marin, 2008b
= Pseudoveleronia Marin, 2008b (type species Veleronia laevifrons Holthuis, 1951b, by original designa-
Pseudoveleronia laevifrons (Holthuis, 1951b)
= Veleronia laevifrons Holthuis, 1951b: 199; Plate 63, fgs f-m. [Galapagos Islands, Ecuador, Gardner
Bay, Hood Island, 4 fms]
Rapipontonia Marin, 2007c
= Rapipontonia Marin, 2007c (type species Rapipontonia paragalene Marin, 2007c, by original designa-
Rapipontonia galene (Holthuis, 1952c)
= Periclimenes (Harpilius) galene Holthuis, 1952c: 62; Fig. 24. [Amboina, pier, 0-2 m; Islet near Menado]
Rapipontonia hydra Marin, 2009b: 10; Figs 5-7. [Pacifc Ocean, Vietnam, Nhatrang Bay, on pile under pier
of MMA, depth 2 m, on hydroid Lytocarpia sp.]
Rapipontonia paragalene Marin, 2007c: 777; Figs 1-6. [Vietnam, South China Sea, Nhatrang Bay, on pile
under pier of Marine Military Academy, 2 m, on hydroid Lytocarpia sp.]
Rapipontonia platalea (Holthuis, 1951a)
= Periclimenes (Harpilius) platalea Holthuis, 1951a: 157; Fig. 32. [of French Guinea, 923'N 1507'W,
30-34 m]
Rostronia Fransen, 2002
= Rostronia Fransen, 2002 (type species Pontonia stylirostris Holthuis, 1952c, by original designation
Rostronia stylirostris (Holthuis, 1952c)
= Pontonia stylirostris Holthuis, 1952c: 169; Figs 82-84. [between Misool and New Guinea, 142.5'S
13047.5'W, 32 m]
Sandimenes Li, 2009
= Sandimenes Li, 2009 (type species Periclimenes hirsutus Bruce, 1971e, by original designation and
Sandimenes hirsutus (Bruce, 1971e)
= Periclimenes hirsutus Bruce, 1971e: 91; Figs 1-6. [Nukulau Island, Laucala Bay Suva, Viti Levu, Fii]
Sandyella Marin, 2009c
= Sandyella Marin, 2009c (type species Chacella tricornuta Hendrickx, 1990a, by original designation,
gender feminine)
Sandyella bicornuta (Li & Poupin, 2009)
= Chacella bicornuta Li & Poupin, 2009: 581; Figs 1-4. [Clipperton, 1019.22'N 10913.38'W, 55 m,
outer reef]
Sandyella mclaughlinae (Li, 2006a)
= Chacella mclaughlinae Li, 2006a: 360; Figs 1-4. [Clipperton Island, 1019.22'N 10913.38'W, 55 m]
Sandyella sexicornuta (Li & Poupin, 2009)
= Chacella sexicornuta Li & Poupin, 2009: 587; Figs 9-12. [Clipperton, 1018.03'N 10913.77'W, 54 m,
outer reef]
Sandyella tricornuta (Hendrickx, 1990a)
= Chacella tricornuta Hendrickx, 1990a: 326; Figs 1-3. [Mexico, Nayarit, Islas Tres Marias, east coast
of Isla Maria Madre, 2138'N 10632'W, 30-40 m]
Sandyella quadricornuta (Li & Poupin, 2009)
= Chacella quadricornuta Li & Poupin, 2009: 584; Figs 5-8. [Clipperton, 1019.22'N 10913.38'W, 55 m,
outer reef]
Stegopontonia Nobili, 1906b
= Stegopontonia Nobili, 1906b (type species Stegopontonia commensalis Nobili, 1906b, by monotypy, gen-
Stegopontonia commensalis Nobili, 1906b: 258. [lagon de Hao, commensal de Echinothrix turcarum]
Tectopontonia Bruce, 1973d
= Tectopontonia Bruce, 1973d (type species Tectopontonia maziwiae Bruce, 1973d, by original designa-
Tectopontonia maziwiae Bruce, 1973d: 172; Figs 1-4. [Maziwi Island, of Pangani, Tanganyika, 530.0'S
3904.1'E]
Thaumastocaris Kemp, 1922
= Thaumastocaris Kemp, 1922 (type species Thaumastocaris streptopus Kemp, 1922, by original desig-
Thaumastocaris streptopus Kemp, 1922: 244; Figs 78-80. [Noumea, New Caledonia]
Tuleariocaris Hipeau-Jacquote, 1965
= Tuleariocaris Hipeau-Jacquote, 1965 (type species Tuleariocaris holthuisi Hipeau-Jacquote, 1965, by
Tuleariocaris holthuisi Hipeau-Jacquote, 1965: 248; Plates 1-5. [Tular, Madagascar, on echinoids Echi-
nometra and Stomopneustes; see Chace & Bruce, 1993]
Tuleariocaris neglecta Chace, 1969: 266; Figs 10-11. [Bellairs Research Institute of McGill University, St.
James, Barbados, on Diadema antillarum]
Tuleariocaris sarec Berggren, 1994a: VIII.19; Figs 10-14. [coral reef of Ilha des Portugueses, 500 m west of
the northern half of Inhaca Island, approx. 26S 33E, Moambique]
Tuleariocaris sarec Berggren, 1994b: 794; Figs 10-14. [coral reef of Ilha des Portugueses, 500 m west of the
northern half of Inhaca Island, approx. 26S 33E, Moambique]
= Tuleariocaris sarec Berggren, 1994b: 794; Figs 10-14. [coral reef of Ilha des Portugueses, 500 m west
of the northern half of Inhaca Island, approx. 26S 33E, Moambique]
Tuleariocaris zanzibarica Bruce, 1967b: 33; Figs 13-18. [Mtoni, Zanzibar, from an echinoid collected at
L.W.S.]
Typton O.G. Costa, 1844a
= Typton O.G. Costa, 1844a (type species Typton spongicola O.G. Costa, 1844a, by monotypy, gender
masculine; name placed on the Ofcial List of Generic Names in Zoology in Opinion 470 in 1957)
= Pontonella Heller, 1856 (type species Pontonella glabra Heller, 1856 (junior subjective synonym of
Typton spongicola O.G. Costa, 1844a), by monotypy, gender feminine)
Typton ascensionis Manning & Chace, 1990: 10; Fig. 4. [McArthur Point, Ascension Island]
Typton australis Bruce, 1973e: 254; Figs 1-4. [Chinamans Reef, between Opal and Rudder Reefs, Great
Barrier Reef, Queensland, Australia, 10f]
Typton bawii Bruce, 1972e: 243; Figs 1-5. [South of Bawi Island, of west coast of Zanzibar, 69.7'S 398.3'E,
18-25 m]
Typton capricorniae Bruce, 2000: 94; Figs 6-8. [Heron Island, Capricorn Islands, Queensland]
Typton carneus Holthuis, 1951b: 162; Plate 51. [Tortugas, Florida]
Typton crosslandi Bruce, 1978d: 294; Figs 1-3. [Of Onslow Island, near Charles (= Floreana, or Santa Ma-
ria Island), Galapagos Archipelago, 4 fms]
Typton distinctus Chace, 1972: 49; Figs 13-14. [Los Arroyos, Provincia de Pinar del Rio, in sponge]
Typton gnathophylloides Holthuis, 1951b: 159; Plate 50. [Tortugas, about 13 miles S of No. 2 Red Buoy, up
to 45 fms]
Typton hephaestus Holthuis, 1951b: 159; Plate 49, fgs o-p. [southern part of the Gulf of California, 2412'N
10955'W, 9.5 fms]
Typton holthuisi De Grave, 2010: 210; Figs 1-2. (Ascension Island, Comfortless Cove, 0754.615'S
01424.142'W, 2 m)
Typton manningi Bruce, 2000: 87; Figs 1-5. [Abbot Point, Queensland, Australia, 1953'S 14805'E]
Typton prionurus Holthuis, 1951b: 165; Plate 52. [Tortugas, Florida, in channel between Middle Ground
and White Shoal, 10 fms]
Typton serratus Holthuis, 1951b: 167; Plate 53. [Galapagos Islands, Ecuador, Albemarle Island, Tagus
Cove, shallow water]
Typton spongicola O.G. Costa, 1844a: 289. [Bay of Naples, Italy; neotype designated by Bruce, 2009c]
= Pontonia pulsatrix Nardo, 1847: 6. [le lagune e golfo Veneto]
= Pontonella glabra Heller, 1856: 634; Plate 9. [Meereskste von Zara]
= Typton spongiosus Spence Bate, 1868b: 119; Plate 11, Figs 1-1z. [Type locality not indicated]
? = Pontonia Vagans Gourret, 1887b: 1132. [nomen nudum]
? = Pontonia Vagans Gourret, 1888: 39. [Fonds vaseux entre lle de Tiboulen et la calanque de Mjean,
par 35 brasses]
Typton tortugae McClendon, 1911
= Typton tortug McClendon, 1911: 57; Plate 1, fg. 2. [Tortugas, Florida]
Typton vulcanus Holthuis, 1951b: 157; Plate 49, Figs a-n. [south of Tortugas, Florida]
Typton wasini Bruce, 1977d: 272; Figs 1-6. [Wasini Island Channel, Kenya, 439.4'S 3922.2'E, 6 fm]
= Typton nanus Bruce, 1987d: 49; Figs 1-5. [Australian North-West Shelf, 1634'S 12127'E, 40-46 m]
Typtonoides Bruce, 2010j
= Typtonoides Bruce, 2010j (type species Typtonoides neili Bruce, 2010j, by original designation and
Typtonoides neili Bruce, 2010j: 69; Figs 1-6. [1953'S 15819'E, Long Island, Chesterfeld Islands, New
Caledonia, 15 m]
Typtonychus Bruce, 1996
= Typtonychus Bruce, 1995 (unavailable name; see Holthuis, 1996)
= Typtonychus Bruce, 1996 (type species Typtonychus crassimanus Bruce, 1996, by original designa-
Typtonychus anomalus (Bruce, 1979e)
= Onycocaris anomala Bruce, 1979e: 69; Figs, 1-4. [between North and South Shell Islands, Port Dar-
win, Northern Australia, 3-7 fms]
Typtonychus crassimanus Bruce, 1996: 253; Figs 22-27. [New Caledonia, Anaa Reef, 2122.70'S 16601.50'E, 38 m]
Typtonychus dentatus (Fujino & Miyake, 1969c)
= Typton dentatus Fujino & Miyake, 1969c: 80; Figs 1-2. [Ukachi, Yoron-jima Island, Ryukyu Islands,
Japan]
Typtonychus dimorphus (Bruce, 1986f)
= Typton dimorphus Bruce, 1986f: 278; Figs 1-4. [Ashmore Reef, Timor Sea, 1215.0'S 12300.0'E, outer
slope of NW reef, 5 m]
Unguicaris Marin & Chan, 2006
= Unguicaris Marin & Chan, 2006 (type species Unguicaris panglaonis Marin & Chan, 2006, by origi-
Unguicaris alegrias (Bruce, 1986d)
= Periclimenes alegrias Bruce, 1986d: 143; Figs 1A, 2-5, 15A-C. [Coral Bay, Port Essington, 1111.2'S
13202.8'E, 2-4 m]
Unguicaris novaecaledoniae (Bruce, 1968a)
= Periclimenes novaecaledoniae Bruce, 1968a: 1157; Figs 6-9. [Ilot Matre, Noumea, New Caledonia,
2220'S 11625'E]
Unguicaris panglaonis Marin & Chan, 2006: 530; Figs 6-13. [Philippines, 929.4'N 12356.0'E, 6-37 m]
Unguicaris pilipes (Bruce & Zmarzly, 1983)
= Periclimenes pilipes Bruce & Zmarzly, 1983: 644; Figs 1-5. [southern tip of Medren Islet, Enewetak
Atoll, Marshall Islands, 1124'N 16222'E, 3 m]
Urocaris Stimpson, 1860a
= Urocaris Stimpson, 1860a (type species Urocaris longicaudata Stimpson, 1860a, by original selection,
gender feminine)
Urocaris longicaudata Stimpson, 1860a: 39. [in litoribus Carolinensibus]
Veleronia Holthuis, 1951b
= Veleronia Holthuis, 1951b (type species Veleronia serratifrons Holthuis, 1951b, by original selection,
gender feminine)
Veleronia serratifrons Holthuis, 1951b: 196; Plate 62; Plate 63, fgs a-e. [Ecuador, La Libertad, 4 fms]
Veleronia sympathes (De Ridder & Holthuis, 1979)
= Pontonides sympathes De Ridder & Holthuis, 1979: 101; Figs 1-3. [Punta Pit, north-east coast of San
Cristobal Island, Galapagos Archipelago, Ecuador, 8 m, on Antipathes galapagensis]
Veleroniopsis Gore, 1981
= Veleroniopsis Gore, 1981 (type species Veleroniopsis kimallynae Gore, 1981, by original designation
Veleroniopsis kimallynae Gore, 1981: 147; Fig. 2. [Elbow Reef, of Key Largo, Monroe County, Florida,
2507.70'N 8015.90'W, 18.3 m]
Vir Holthuis, 1952c
= Vir Holthuis, 1952c (type species Palaemonella orientalis Dana, 1852a, by original designation and
Vir colemani Bruce, 2003c: 119; Figs 1-6. [Loloata Island, Papua New Guinea, 15 m]
Vir euphyllius Marin & Anker, 2005: 118; Figs 1-5. [Vietnam, Nha Trang Bay, Tre Island, southern bay,
near lighthouse, 15 m, on Euphyllia cf. divisa]
= Vir pareuphyllius Marin & Anker, 2005: 123; Figs 6-8. [Vietnam, Nha Trang Bay, Tre Island, northern
Bay Tre, 7 m, on Euphyllia cf. parancora]
Vir longidactylus Marin, 2008a: 383; Figs 6-8, 9c. [South China Sea, Vietnam, Nhatrang Bay, Tre Island, Tre
Bay, on Physogyra lichtensteini, 5 m]
Vir orientalis (Dana, 1852a)
= Palaemonella orientalis Dana, 1852a: 26. [in mari Suluensi]
Vir philippinensis Bruce & Svoboda, 1984: 87; Figs 1-4. [island of Cebu, associated with Plerogyra sinuosa]
Vir smiti Fransen & Holthuis, 2007: 101; Figs 1-27, 32-34. [Philippines, Cebu Strait, W of Bohol, N side of
Cabilao Island, NE of Looc, 953.59'N 12346.92'E, 15 m, on Physogyra lichtensteini]
Waldola Holthuis, 1951b
= Waldola Holthuis, 1951b (type species Waldola schmiti Holthuis, 1951b, by original designation
Waldola schmiti Holthuis, 1951b: 186; Plate 58; Plate 59, Figs a-f. [Tepic, Mexico, of Isabel Island, 10-25
fms]
Yemenicaris Bruce, 1997a
= Yemenicaris Bruce, 1997a (type species Yemenicaris trullicauda Bruce, 1997a, by original designation
Yemenicaris trullicauda Bruce, 1997a: 1215; Figs 1-4. [Khalf, Yemen, 1435'N 4910'E, to 9 m]
Zenopontonia Bruce, 1975e
= Zenopontonia Bruce, 1975e (type species Periclimenes (Periclimenes) noverca Kemp, 1922, by original
Zenopontonia noverca (Kemp, 1922)
= Periclimenes (Periclimenes) noverca Kemp, 1922: 162; Figs 28-30. [New Caledonia]
Family TYPHLOCARIDIDAE Annandale & Kemp, 1913
Typhlocaris Calman, 1909
= Typhlocaris Calman, 1909 (type species Typhlocaris galilea Calman, 1909, by monotypy, gender fem-
inine; name placed on the Ofcial List of Generic Names in Zoology in Opinion 470 in 1957)
Typhlocaris ayyaloni Tsurnamal, 2008: 490; Figs 1-4. [Ayyalon Cave, Israel]
Typhlocaris galilea Calman, 1909: 94; Plate 19. [small pond near the town of Tiberias]
Typhlocaris lethaea Parisi, 1921: 241; Figs 1-5. [Grota del Lete (Giok-Kebir), 10 km d Est di Bengasi, Cire-
naica]
Fig. 42. Typhlocaris salentina Caroli, 1923. Photo by Ninni Ciccarese.
Typhlocaris salentina Caroli, 1923: 265. [grota presso Castro, in Terra dOrtanto; full description in Caro-
li, 1924; type locality corrected therein to grote la Zinzulusa, costa della Penisola Salentina copreso
tra il Capo dOtranto e il Capo di Leuca] (Fig. 42)
Superfamily ALPHEOIDEA Rafnesque, 1815
Family ALPHEIDAE Rafnesque, 1815
Acanthanas Anker, Poddoubtchenko & Jeng, 2006
= Acanthanas Anker, Poddoubtchenko & Jeng, 2006 (type species Acanthanas pusillus Anker, Pod-
doubtchenko & Jeng, 2006, by original designation and monotypy, gender masculine)
Acanthanas pusillus Anker, Poddoubtchenko & Jeng, 2006: 342; Figs 1-5. [north of Doljo, Panglao Island,
Philippines, 935.9'N 12344.5'E, 24 m]
Alpheopsis Coutire, 1897b
= Alpheopsis Coutire, 1897b (type species Betaeus trispinosus Stimpson, 1860a, designated by
Holthuis, 1955b, gender feminine; name placed on the Ofcial List of Generic Names in Zoology in
Alpheopsis aequalis Coutire, 1897b
= Alpheopsis qualis Coutire, 1897b: 382. [mer Rouge; ocan Indien]
= Alpheopsis equalis var. truncatus Coutire, 1903: 89; Figs 37-38. [Goifufahendu Atoll]
= Alpheopsis equalis var. truncatus Coutire, 1905b: 869; Fig. 139. [Goifufahendu Atoll]
= Alpheopsis consobrinus De Man, 1910a: 305. [825.2'S 12718.4'E, anchorage between Nusa Besi and
the NE-point of Timor, 27-54 m]
Alpheopsis africana Holthuis, 1952a
= Alpheopsis africanus Holthuis, 1952a: 45; Fig. 11 (as A. africana). [830'S 13E, 20 m W. Pointa do
Dand, 150 m]
Alpheopsis allanhancocki Wicksten, 1992: 51; Figs 1-2. [Isla Candelero, Sonora, Gulf of California, Mexico,
2756'N 11103'W, 10 m]
Alpheopsis azorica Anker, dUdekem dAcoz & Poddoubtchenko, 2005: 98; Figs 1-3. [Monte da Guia near
Horta, Faial Island, Azores, in large marine cave, 35 m]
Alpheopsis biunguiculata Banner, 1953
= Alpheopsis biunguiculatus Banner, 1953: 18; Fig. 5. [Halape, Kau Coast, Hawaii, from Pocillopora
meandrina]
Alpheopsis chalciope De Man, 1910a
= Alpheopsis Chalciope De Man, 1910a: 306. [07.2'N 13025.5'E, 83-59 m]
Alpheopsis chilensis Coutire, 1897b: 382. [Chili]
Alpheopsis cortesiana Wicksten & Hendrickx, 1986: 196, Fig. 1. [Of Ro Fuerte, Sinaloa, Mexico, 2548'N
10934'W, 90 m, mud]
Alpheopsis diabolus Banner, 1956: 325 (as A. diabilus); Fig. 3. (as A. diabolus). [Saipan, Mariana Islands;
name corrected to A. diabolus by Banner & Banner, 1964 under First Reviser Principle]
Alpheopsis equidactylus (Lockington, 1877a)
= Alpheus equidactylus Lockington, 1877a: 35. [Monterey]
Alpheopsis garricki Yaldwyn, 1971: 87. [of Rangitoto Island, Cook Strait, 4044'S 1740'E, 40-50 fms]
Alpheopsis harperi Wicksten, 1984a: 97; Fig. 1. [of Freeport, Texas, 2843'41''N 9514'10''W, 21 m]
Alpheopsis idiocarpus Coutire, 1908: 194. [Providence, 50-78 brasses]
Alpheopsis labis Chace, 1972: 55; Fig. 15. [Antigua, Charlote Point, English Harbour]
Alpheopsis shearmii (Alcock & Anderson, 1899)
= Alpheus Shearmii Alcock & Anderson, 1899: 283. [Investigator stn 232 (Laccadive Sea, 717'30''N
7654'30''E), 430 fms]
Alpheopsis tetrarthri Banner, 1956: 328; Fig. 4. [north side of passage into Saipan harbor, 40-45f]
Alpheopsis trigona (Rathbun, 1901)
= Jousseaumea trigona Rathbun, 1901: 111; Fig. 21. [of Vieques, 6 fms]
Alpheopsis trispinosa (Stimpson, 1860a)
= Betaeus trispinosus Stimpson, 1860a: 32. [Green Cape, New South Wales, 30 fms; neotype selection
by Banner & Banner, 1973a]
Alpheopsis undicola Banner & Banner, 1973a: 340; Fig. 15. [Opal Reef, outermost margin of Great Barrier
Reef, 5f]
Alpheopsis vietnamensis Tiwari, 1964b: 314. [Anchorage of Itu Aba; fully described in Tiwari, 1965, as
Alpheopsis vietnami]
Alpheopsis yaldwyni Banner & Banner, 1973a: 344; Fig. 17. [One Tree Island, Capricorn Group] (Fig. 43)
Alpheus Fabricius, 1798
= Crangon Weber, 1795 (type species Astacus Malabaricus Fabricius, 1775, by monotypy, gender fem-
inine; name suppressed under the plenary power of the ICZN for both the Principle of Priority and
that of Hononymy, and placed on the Ofcial Index of Rejected and Invalid Names in Zoology in
= Alpheus Fabricius, 1798 (type species Alpheus avarus Fabricius, 1798, designated by Latreille, 1810,
gender masculine; junior homonym of Alpheus Weber, 1795 (Crustacea Brachyura); name conserved
by the suppression for the Principle of Priority and that of Homonymy under the plenary powers
of the ICZN of the generic name Alpheus Weber, 1795; name placed on the Ofcial List of Generic
= Cryptophthalmus Rafnesque, 1814 (type species Cryptophtalmus ruber Rafnesque, 1814 (junior sub-
jective synonym of Cancer glaber Olivi, 1792), by monotypy, gender masculine)
= Autonoma Risso, 1816 (type species A.[utonoma] Olivii Risso, 1816 (junior subjective synonym of
Cancer glaber Olivi, 1792), by monotypy, gender feminine)
= Asphalius Roux, 1831 (type species Palmon brevirostris Olivier, 1811, by monotypy, gender mascu-
line)
= Diencia Westwood, 1835a (type species Hipplyte rbra Hailstone, 1835a (junior objective syno-
nym of Hipplyte macrochles Hailstone, 1835b), by monotypy, gender feminine)
= Nauplius Risso, 1844 (type species N.[ika] Variegata Risso, 1816 (invalid senior subjective synonym
of Alpheus dentipes Gurin, 1832), designated by Holthuis, 1977, gender masculine)
= Phleusa Nardo, 1847 (type species Phleusa cynea Nardo, 1847 (junior subjective synonym of Cancer
glaber Olivi, 1792), gender feminine)
Fig. 43. Alpheopsis yaldwyni Banner & Banner, 1973. Photo by Arthur Anker.
= Halopsych de Saussure, 1857b (type species Halopsych lutaria de Saussure, 1857b, a junior subjec-
tive synonym of A.[lpheus] heterochlis Say, 1818, by monotypy, gender feminine)
= Alpheoides Paulson, 1875 (type species A.[lpheus] insignis Heller, 1861, designated by Holthuis,
1955b, gender masculine)
= Paralpheus Spence Bate, 1888 (type species Palmon diversimanus Olivier, 1811, by monotypy, gen-
der masculine)
= Thunor Armstrong, 1949 (type species Crangon rathbunae Schmit, 1924b (junior subjective syno-
nym of Alpheus Simus Gurin-Mneville, 1855 [in Gurin-Mneville, 1855-1856), by monotypy, gen-
der masculine)
Alpheus acutocarinatus De Man, 1909a: 104. [725'S 11316'E, Madura Strait, 56 m; 844'.5S 1162'.5E, Bay
of Labuan Tring, west coast of Lombok, 18-27 m; 058'.5N 12242'.5E, west of Kwandang-bay-en-
trance 72 m; Sapeh-bay, east coast of Sumbawa, up to 36 m]
Alpheus acutofemoratus Dana, 1852a
= Alpheus acuto-femoratus Dana, 1852a: 22. [in freto Balabac]
= Alpheus parabrevipes Coutire, 1898a: 151; Figs 2-2c. [Samoa]
Alpheus adamastor Coutire, 1908
= Alpheus Adamastor Coutire, 1908: 209. [Salomon, Chagos]
Alpheus aequus W. Kim & Abele, 1988: 55; Fig. 23. [Playa Blancas, Costa Rica]
Alpheus agilis Anker, Hurt & Knowlton, 2009: 11; Figs 4, 5F. [So Tom, 200 m west of Lagoa Azul, rocky
intertidal]
Alpheus agrogon Ramos, 1997: 395; Figs 1-3. [La Ventana, Gorgona Island, 256'10''N 7812'05''W]
Alpheus albatrossae (Banner, 1953)
= Crangon albatrossae Banner, 1953: 60; Fig. 18. [Penguin Bank, west of Molokai, 27 fm]
Alpheus alcyone De Man, 1902: 870; Plate 27, fg. 61. [Ternate]
= Alpheus aculeipes Coutire, 1905b: 892; Plate 79, fgs 31-31g. [South Nilandu Atoll; Hulule Male
Atoll; North Male Atoll; Milandumadulu Atoll, Naifaro Reef; Djibouti; Fernando-Veloso]
Alpheus alpheopsides Coutire, 1905b: 901; Plate 83; fgs 40-40d. [Hulule Male Atoll]
Alpheus amblyonyx Chace, 1972: 59; Fig. 16. [Dominica, north end of Woodbridge Bay]
Alpheus amirantei Coutire, 1908
= Alpheus Amirantei Coutire, 1908: 205. [Amirante, 25-80 brasses]
Alpheus anchistus De Man, 1920a: 108. [of Ambon, 54 m]
Alpheus angulosus McClure, 2002: 368. [nomen novum for Alpheus angulatus McClure, 1995; nec Alpheus
strenuus var. angulatus Coutire, 1905b]
= Alpheus angulatus McClure, 1995: 85; Figs 1-2; nec Alpheus strenuus var. angulatus Coutire, 1905b.
[South Padre Island, Texas, Laguna Madre just north of Brazos-Santiago Pass]
Alpheus angustilineatus Nomura & Anker, 2005: 111; Figs 3-4, 13C-D. [Fuki, Kuro-shima, Yaeyama Is-
lands, southern Ryukyu Archipelago, Japan, 3 m]
Alpheus antepaenultimus W. Kim & Abele, 1988: 103; Fig. 44. [Amador, Panama]
Alpheus architectus De Man, 1897: 726; Plate 34, fgs 60-60f. [Atjeh]
? = Alpheus bullatus Barnard, 1955: 45; fg. 22. [Delagoa Bay; St. Lucia Bay]
Alpheus arenensis (Chace, 1937b)
= Crangon arenensis Chace, 1937b: 119; Fig. 4. [of Arena Bank, 2.5 fms, in coral]
Alpheus arenicolus Banner & Banner, 1983: 14; Fig. 1. [Madagascar]
Alpheus arethusa De Man, 1909a
= Alpheus Arethusa De Man, 1909a: 100. [Haingsisi, Samau island, Timor, 36 m]
Alpheus armatus Rathbun, 1901: 108; Fig. 20. [Ponce]
Alpheus armillatus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]
= A.[lpheus] armillatus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 354. [les Antilles]
= Alpheus lancirostris Rankin, 1900: 541; Plate 17, fg. 5. [Baileys Bay, at low tide]
= Crangon verrilli Schmit, 1924b: 77; Plate 2, fgs 7-10. [Barbados]
Alpheus arnoa Banner, 1957: 199; Fig. 5. [lagoon reef, Arno Atoll, Marshall Islands, 4-60 f]
Alpheus astrinx Banner & Banner, 1982a: 35; Fig. 5. [Garden Island, near Perth, Western Australia, 100
yards ofshore]
Alpheus australiensis Banner & Banner, 1982a: 256; Fig. 79. [Caloundra, Queensland]
Alpheus australosulcatus Banner & Banner, 1982a: 83; Fig. 21. [North Head, Flinders, Victoria]
Alpheus baccheti Anker, 2010a: 50; Figs 1-5. [French Polynesia, Tuamotu Islands, Makemo, lagoon, reef
fat, around 0.5 m at low tide]
Alpheus bahamensis Rankin, 1898
= Alpheus hippotho var. bahamensis Rankin, 1898: 247; Plate 30, fg. 5. [New Providence and Nassau,
N. P.]
Alpheus balaenodigitus Banner & Banner, 1982a: 223; Fig. 70. [Port Walcot, Western Australia, 2039'S
11710'E, 8 fms]
Alpheus bannerorum Bruce, 1987e: 61; Figs 1-6; Plate 1. [Dudley Point, Darwin, Northern Territory]
Alpheus barbadensis (Schmit, 1924b)
= Crangon barbadensis Schmit, 1924b: 70; Plate 2, fgs 1-3. [Of the Castel, E side Barbados, 1-4 fms]
Alpheus barbatus Coutire, 1897c: 235. [Djibouti] (Fig. 44)
Alpheus batesi Banner & Banner, 1964: 94; nomen novum for Alpheus leviusculus sensu Spence Bate, 1888;
nec Dana, 1852b. [Challenger stn 203, 116'N 1239'E, of Panay, Philippine Islands, 20 fms]
Alpheus belli Coutire, 1898a
= Alpheus Belli Coutire, 1898a: 149; Figs 1-1a. [Fernando-Noronha]
Alpheus bellimanus Lockington, 1877a: 34. [San Diego, among kelp]
Alpheus bellulus Miya & Miyake, 1969: 308; Figs 1-2. [Ezura, Tanabe Bay, Wakayama Prefecture, burrow-
ing on gravely sand, 5 m]
Alpheus bicostatus De Man, 1908d: 102. [Sialus Ketjil, Paternoster-islands, 27 m and less; 67'.5N 12026'E,
anchorage of Nort-Ubian, 16-23 m; Buton-strait, between foating seaweed]
Alpheus bidens (Olivier, 1811)
= Palmon bidens Olivier, 1811: 663. [ctes de la Nouvelle-Hollande]
= Alpheus tridentatus Zehntner, 1894: 204; Plate 8, fg. 24. [Amboine]
= Alpheus dissodontonotus Stebbing, 1915: 83; Plate 22. [3350'S 2546'E, 20 fms]
Alpheus bisincisus De Haan, 1849 [in De Haan, 1833-1850]
= Alpheus bis-incisus De Haan, 1849 [in De Haan, 1833-1850]: Plate 45, fg. 3. (1844); 179 (1849, as
Alpheus avarus). [Japan]
Fig. 44. Alpheus barbatus Coutire, 1897. Photo by Arthur Anker.
= Alpheus bis-incisus var. Malensis Coutire, 1905b: 910; Plate 86, fgs 48-48d. [South Male Atoll; Su-
vadiva Atoll; South Nilandu Atoll; Mulaku Atoll]
= Alpheus bis-incisus var. stylirostris Coutire, 1905b: 911; Plate 86, fgs 49-49a. [Mulaku Atoll]
? = Alpheus bisincisus var. variabilis De Man, 1909a: 109. [Makassar, up to 32 m]
Alpheus blachei Crosnier & Forest, 1965a: 358; Fig. 3. [lieudit Pointe Indienne, situ dans les environs de
Pointe-Noire, Congo]
Alpheus bouvieri A. Milne-Edwards, 1878
= Alpheus Bouvieri A. Milne-Edwards, 1878: 231. [toutes les les du Cap-Vert]
Alpheus brachymerus (Banner, 1953)
= Crangon brachymerus Banner, 1953: 58; Fig. 17. [between Maui and Lanai Islands, 21-28 fms]
Alpheus bradypus Coutire, 1905b: 891; Plate 78, fgs 30-30a; Plate 79, fgs 30b-30e. [Minikoi]
Alpheus brevicristatus De Haan, 1844 [in De Haan, 1833-1850]: Plate 45, fg. 1. (1844); 177 (1849, as Alpheus
malabaricus Fabricius) [Japan; lectotype designation by Yamaguchi & Baba, 1993]
= Alpheus kingsleyi Miers, 1879: 54. [357'N 13655'E, 3 fms]
Alpheus brevipes Stimpson, 1860a: 30. [ad insulas Hawaiensis, inter ramos madreporarum]
Alpheus brevirostris (Olivier, 1811)
= Palmon brevirostris Olivier, 1811: 664. [ctes de la Nouvelle-Hollande]
= Alpheus brevirostris var. angustodigitus De Man, 1911d: 385. [Balikpapan, east coast of Borneo]
= A.[lpheus] dispar Randall, 1840: 141. [Manilla]
Alpheus brucei Banner & Banner, 1982b: 359; Fig. 1. [Heron Island reef fat, 0.5 m from Porites andrewsi]
Alpheus bucephaloides Nobili, 1905e: 238. [Golfe Persique, Banc au N.E. dle Arzana et pche ctires
dans les polypiers; lectotype selection by Banner & Banner, 1981a]
Alpheus bucephalus Coutire, 1905b: 890; Plate 78, fgs 29-29f. [Hulule Male Atoll; Minikoi; Cap; Fernan-
do-Veloso; Mah; Djibouti]
= Alpheus consobrinus De Man, 1908d: 101 [Lumu-Lum-shoal, Borneo Bank, reef; South-east side of
Pearl-bank, Sulu-archipelago, 15 m; East side of Pajunga-island, Kwandang-bay, reef exploration;
Anchorage of Sawan, Siau-island, 27 m; Saleyer-anchorage and surroundings, up to 36 m; 1052'.4S
1231'.1E, Boeka- or Cyrus-bay, South coast of Roti-island, up to 36 m; Anchorage east of Sailus
Besar, Paternoster-islands, up to 36 m]
Alpheus buchanorum Banner & Banner, 1983: 19; Fig. 2. [Round Island, of Victoria, Mah, Seychelles]
Alpheus bunburius Banner & Banner, 1982a: 213; Fig. 66. [Bunbury, Western Australia]
Alpheus californiensis Holmes, 1900: 186; Plate 2, fg. 42; Plate 3, fgs 43-44. [San Pedro, California, dredged
in the harbour]
Alpheus canaliculatus Banner & Banner, 1968: 141; Fig. 1. [2005'N 11511'E, 137 fms]
Alpheus candei Gurin-Mneville, 1855 [in Gurin-Mneville, 1855-1856]
= Alpheus Candei Gurin-Mneville, 1855 [in Gurin-Mneville, 1855-1856]: xix; Plate 2, fgs 9-9a.
[costas de la Isla de Cuba]
Alpheus chacei Carvacho, 1979: 455; Figs 4-6. [entre les canaux Belle Plaine et Perrin, prs de la mangrove]
= Alpheus maxilliplanus Christofersen, 1979: 318; Figs 11-13. [Balnerio de Atalaia, Sergipe, Brazil]
Alpheus chamorro Banner, 1956: 349; Fig. 14. [Saipan]
Alpheus chilensis Lenz, 1902
= Alpheus bouvieri var. chilensis Lenz, 1902: 732. [Calbuco]
Alpheus chiragricus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]
= A.[lpheus] chiragricus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 354. [les mers
dAsie]
Alpheus christoferseni Anker, Hurt & Knowlton, 2007a: 3; Figs 1-6, 11a-c, 12b-c. [Brazil, Atol das Rocas,
Ilha do Cemitrio]
Alpheus clamator Lockington, 1877b: 43. [Santa Barbara Island]
= Alpheus transverso-dactylus Kingsley, 1878b: 196. [Santa Barbara and San Diego, California; Bermu-
das]
? = Alpheus barbara Lockington, 1878b: 471. [Santa Barbara, California]
Alpheus clypeatus Coutire, 1905b: 897; Plate 81, fgs 36-36; Plate 82, fgs 36a-36g. [Minikoi; Hulule Male
Atoll]
Alpheus coetivensis Coutire, 1908
= Alpheus Coetivensis Coutire, 1908: 210. [Coetivy; Salomon]
= Crangon edmondsoni Banner, 1953: 78; Fig. 26. [Kawela Bay, Oahu]
Alpheus collumianus Stimpson, 1860a: 30. [ad insulas Bonin, inter corallia vientia ad prof. 1 org]
= Alpheus Seurati Coutire, 1905b: 881; Plate 75, fgs 20-20e. [lle Marutea, archipel Gambier]
= Alpheus Malhaensis Coutire, 1908: 205. [Saya de Malha, 29 brasses]
= Alpheus collumianus probabilis Banner, 1956: 338; Fig. 10. [about 1.4 miles ofshore, north of Puntan
Muchot, Saipan, 8-9f]
= Alpheus collumianus medius Banner, 1956: 340; Fig. 11. [Saipan]
= Alpheus collumianus inermis Banner, 1956: 342; Fig. 12. [about 1 mile of Muchot Point, near en-
trance to Saipan harbor, 22f]
Alpheus colombiensis Wicksten, 1988: 1; Fig. 1. [Baha Malaga, Pacifc coast of Colombia, 40'N 7715'W,
under rocks]
= Alpheus hamus W. Kim & Abele, 1988: 91; Fig. 38. [Golfto, Costa Rica]
Alpheus compressus Banner & Banner, 1981b: 227; Fig. 3. [1359.8'N 12018.6'E, 188-192 m]
Alpheus confusus Carvacho, 1989: 59; Fig. 1. [Isla San Jos]
Alpheus coutierei De Man, 1909a
= Alpheus Coutierei De Man, 1909a: 107. [South-east side of Pearl-bank, Sulu-archipelago, 15 m;
825'.2S 12718'.4E, Anchorage between Nusa-Besi and the N.E.-point of Timor, 27-54 m]
Alpheus crinitus Dana, 1852a: 21. [in freto Balabac]
Alpheus cristatus Coutire, 1897d: 303. [Thursday-Island]
Alpheus cristulifrons Rathbun, 1900: 152. [Brazil, Fernando de Noronha; lectotype selection by Anker,
Hurt & Knowlton, 2008a]
Alpheus crockeri (Armstrong, 1941)
= Crangon crockeri Armstrong, 1941: 8; Figs 2-3. [eastern reef, Mataatu Harbor, Savaii, shallow water]
= Crangon tuthilli Banner, 1953: 63; Fig. 19. [of southwest Oahu, 40-350 f]
Alpheus cyanoteles Yeo & Ng, 1996: 46; Figs 4-7. [Sungei Tementang, Kota Tinggi, Johore, 152'0.81''N
10355'49.2''E]
Alpheus cylindricus Kingsley, 1878b: 196. [Pearl Island, Bay of Panama; neotype designation by Anker,
Hurt, Jara & Knowlton, 2008 invalid, as holotype extant]
Alpheus cythereus Banner & Banner, 1966a: 123; Fig. 45. [Patong Beach, Phuket, Thailand, 2 m]
Alpheus dasycheles Coutire, 1908: 211. [Seychelles, 37 brasses]
Alpheus davaoensis Chace, 1988: 21; Fig. 2. [Philippines, Davao Gulf, Mindanao, 704'48''N 12539'38''E,
51 m]
Alpheus dentipes Gurin, 1832: 39; Plate 27, fg. 3. [Sapience; cap Tnare; golfe de Gnes]
= N.[ika] Variegata Risso, 1816: 86. [le golfe de Beaulieu, dans les algues profondes; name placed on
the Ofcial Index of Rejected and Invalid Names in Zoology in Opinion 606 in 1961]
= Alpheus streptochirus Stimpson, 1860a: 30. [ad insulas Cape de Verdes, inter nulliporas ad prof.
20 org]
= Crypt.[ophthalmus] Costa O.G. Costa, 1871 [in O.G. Costa & A. Costa, 1838-1871]: 2; Plate 7, fg. 2.
[vive nel Mediterraneo egualmente che nello Ionio]
= Alpheus similis Czerniavsky, 1884: 30. [Portu Suchum, circa 2 m]
= Alpheus cristidigitus Spence Bate, 1888: 546; Plate 97, fg. 3. [Challenger stn 93C, 1657'15''N 251'W,
St. Vincent, Cape Verde Islands, 52 fms]
= Alpheus Gabrieli Gourret, 1887a: 1033. [pointe rouge de Montredon]
Alpheus deuteropus Hilgendorf, 1879: 834; Plate 4, fgs 8-10. [Zanzibar]
Alpheus diadema Dana, 1852a: 23. [ad insulam Maui Hawaiensem]
= A.[lpheus] insignis Heller, 1861: 24. [im rothen Meere; fully described in Heller, 1862c]
Alpheus digitalis De Haan, 1844 [in De Haan, 1833-1850]: Plate 45, fg. 4 (1844); 178 (1849). [Japan]
= Alpheus distinguendus De Man, 1909b: 155; Plate 7, Figs 9-14. [Japan]
Alpheus distinctus W. Kim & Abele, 1988: 95; Fig. 40. [Punta Paitilla, Panama]
Alpheus djeddensis Coutire, 1897e: 202. [Djeddah]
Alpheus djiboutensis De Man, 1909b: 160; Plate 8, Figs 17-24. [Djibouti]
Alpheus dolerus Banner, 1956: 362; Fig. 21. [Saipan]
Alpheus edamensis De Man, 1888b
= Alpheus Hippotho var. edamensis De Man, 1888b: 518. [Pulo Edam; Amboina]
= Alpheus acanthomerus Ortmann, 1890: 474; Plate 36, fg. 12. [Tahiti]
Alpheus edwardsii (Audouin, 1826)
= Athanas Edwarsii Audouin, 1826: 91 (lapsus for Athanas Edwardsii, justifed emendation in Au-
douin, 1827; accompanying fgures in Savigny, 1826: Plate 10, fg. 1). [Suez; neotype selection by
Banner & Banner, 1973b]
= Alpheus Audouini Coutire, 1905b: 911; Plate 87, Figs 52a-52a. [Miladumadulu Atoll, Goidu Atoll,
Hulule Male Atoll, Reef Naifaro, Suvadiva Atoll, Minokoi]
Alpheus ehlersii De Man, 1909c: 663; Plate 70. [Edam, Bay of Batavia]
Alpheus estuariensis Christofersen, 1984: 191; Figs 1-2. [Rio Potengi estuary, Natal, State of Rio Grande
do Norte, Brazil]
Alpheus euchirus Dana, 1852a: 21. [in freto Balabac]
Alpheus euphrosyne De Man, 1897: 745; Plate 36, fgs 64-64d. [Java-See]
= Alpheus eurydactylus De Man, 1920a: 109. [Java]
Alpheus exilis W. Kim & Abele, 1988: 105; Fig. 45. [Sullivan Bay, James Island, Galapagos Islands]
Alpheus explorator Boone, 1935: 139; Fig. 11; Plate 36. [Flores Strait, near Larantuka village, Flores Island,
Dutch East Indies, 140 fms]
Alpheus eulimene De Man, 1909a
= Alpheus Eulimene De Man, 1909a: 101. [07'.2N 13025'.5E, 83 m]
Alpheus facetus De Man, 1908d: 100. [Anchorage of Pulu Jedan, east coast of Aru-islands (Pearl-banks),
13 m]
Alpheus fagei Crosnier & Forest, 1965b: 603; Fig. 1. [So Tom, 020'N 647'E, 10 m, algues calcaires]
Alpheus fasciatus Lockington, 1878b: 478. [Port Escondido, Gulf of California]
Alpheus fasqueli Anker, 2001: 59; Figs 1c-d, 2b, 3f-j, 4e-g, 5i-n, 6l-n. [imported from Sri Lanka]
Alpheus felgenhaueri W. Kim & Abele, 1988: 40; Fig. 16. [outer Miramar Bay on Cape Haro, north of Guay-
mas, Sonora, Mexico]
Alpheus fenneri Bruce, 1994b: Figs 1-2, 5A. [Manado, Sulawesi, Indonesia, of Nusantara diving Centre, 6 m]
Alpheus frmus W. Kim & Abele, 1988: 93; Fig. 39. [Mirafores Locks, Panama]
Alpheus foridanus Kingsley, 1878b: 193. [Fort Jeferson, Florida]
= Alpheus foridanus var. africana Balss, 1916: 21; Fig. 5. [Elfenbein-Kste, Wappu, 40 m]
= Alpheus platycheirus Boone, 1927: 131; Figs 29-30. [Siguanea Bay, Isle of Pines, 12 fms]
Alpheus foresti Banner & Banner, 1981b: 229; Fig. 4. [1402.7'N 12020.3'E, 191-200 m]
Alpheus formosus Gibbes, 1850: 196. [USA, Florida, Big Pine Key, Bahia Honda State Park, shallow subtid-
al (less than 1.5 m), under rocks; neotype designation by Anker, Hurt & Knowlton, 2008b]
= Alpheus Poeyi Gurin-Mneville, 1855 [in Gurin-Mneville, 1855-1856]: xix; Plate 2, fg. 10. [Cuba]
Alpheus frontalis H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]
= A.[lpheus] frontalis H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 356. [les ctes de la
Nouvelle-Hollande]
= Alpheus latifrons A. Milne-Edwards, 1873: 87. [Upolu]
= Betaeus utricola Richters, 1880: 164; Plate 17, Figs 34-35. [Insel Mauritius und Seychellen]
Alpheus funafutensis Borradaile, 1899: 1013; Plate 65, fg. 10. [Funafuti]
= Alpheus acanthomerus var. inermis Lanchester, 1901: 564. [Kelantan]
Alpheus fujitai Nomura & Anker, 2005: 115; Figs 5-6, 13E-F. [Mizugama, Kadena, Okinawa-jima, Ok-
inawa Islands, central Ryukyu Archipelago, Japan, submarine cave, 5-8 m]
Alpheus fushima Nomura, 2009: 115; Figs 1-3. [Nishinohama, Kuro-shima Island, Yaeyama Islands,
southern Ryukyu Archipelago, Japan, 5 m, internal canals of large colony of an unidentifed
sponge]
Alpheus galapagensis Sivertsen, 1933
= Alpheus strenuus var. galapagensis Sivertsen, 1933: 3; Plate 1, fgs 1-5. [Floreana, Post Ofce Bay,
Ranvik]
= Alpheus canalis W. Kim & Abele, 1988: 72; Fig. 30. [North Bay, San Francisco Island, Baja Califor-
nia]
? = Alpheus cryptodentatus Christofersen & Ramos, 1988: 61; Figs 1- 2. [Isla Playa Blanca, Ensenada
de Utra, Choc, Pacifc coast of Colombia, 602'N 7720'W]
Alpheus georgei Banner & Banner, 1982a: 200; Fig. 61. [40 miles W of Cape Jaubert, 23 fms, from sponge]
Alpheus glaber (Olivi, 1792)
= Cancer glaber Olivi, 1792: 51; Plate 3, fg. 4. [la Golfo e delle Lagune di Venezia]
= Cryptophthalmus ruber Rafnesque, 1814: 23. [Sicile]
= A.[utonoma] Olivii Risso, 1816: 166. [golfe de Nice]
= Phleusa cynea Nardo, 1847: 6. [le lagune e golfo Veneto]
Alpheus gracilipes Stimpson, 1860a: 31. [French Polynesia, Tahiti; neotype selection by Nomura & Ank-
er, 2005]
Alpheus gracilis Heller, 1861
= A.[lpheus] gracilis Heller, 1861: 27. [im rothen Meere]
= Alpheus gracilis var. Alluaudi Coutire, 1905b: 882. [Mah]
= Alpheus gracilis var. luciparensis De Man, 1911d: 337. [5700 m N, 279 E from South Point of South-
Lucipara-island, reef]
= Crangon gracilis var. simplex Banner, 1953: 75; Fig. 25. [Waikiki Reef, Oahu]
Alpheus grahami Abele, 1975b: 72; Fig. 29 A-C, E-J. [southeastern side of Malpelo Island, Colombia, 10
m]
Alpheus haanii Ortmann, 1890
= Alpheus Haanii Ortmann, 1890: 472. [nomen novum for Alpheus minor De Haan, 1844 [in De Haan,
1833-1850]]
= Alpheus minor De Haan, 1844 [in De Haan, 1833-1850]: Plate 45, fg. 6 (1844); 180 (1849). [Japan]
Alpheus hailstonei Coutire, 1905b
= Alpheus Hailstonei Coutire, 1905b: 879; Plate 74, fgs 18-18c. [North Male Atoll; Mahlos Atoll;
Hulule Male Atoll]
= Alpheus Hailstonei var. laetabilis De Man, 1908d: 98. [823'.5S 1194'6E, Sapeh-Strait, 70 m; 70'S
12034'5E, 400-120 m; 07'2N 13025'5E, 83-59 m; Banda-Anchorage, 9-36 m; 825'2S 12718'4E,
anchorage between Nusa Besi and the N.E.-point of Timor, 27-54 m; 839'.1S 1274'.4E, anchorage
south of Timor, 34 m; Haingsis, Samau-island, up to 36 m; 830'S 1197'.5E, Sapeh-Strait, 73 m]
= Alpheus Hailstonei var. assimulans De Man, 1908d: 99. [536'.5S 13255'.2E, 2.3 miles N, 63W from
the North Point of Nuhu Jaan, Kei-islands, 90 m; 825'2S 12718'.4E, anchorage between Nusa Besi
and the N.E.-point of Timor, 27-54 m; Mid-channel in Solor-strait of Kampong Menanga, 113 m]
= Crangon hailstonei var. paucispinata Banner, 1953: 51; Fig. 6. [of Kauai, 68-90 fms]
Alpheus halesii Kirk, 1887: 194; Plate 6d. [East Coast of Wellington Province]
Alpheus hebes W. Kim & Abele, 1988: 62; Fig. 26. [James Island, Galapagos Islands]
? = Alpheus bouvieri var. Bastardi Coutire, 1898b: 133 (partim); Fig. 1a. [specimen from Panama only]
Alpheus heeia Banner & Banner, 1975a: 433; Fig. 5. [Kaneohe Bay, Oahu, Hawaii, 2126'55''N 15747'48''W]
Alpheus heronicus Banner & Banner, 1982a: 220; Fig. 69. [Heron Island, Capricorn Group, Queensland,
probably intertidal]
Alpheus heterocarpus (Yu, 1935)
= Crangon heterocarpus Yu, 1935: 63; Figs 3-4. [Peitaio]
Alpheus heterochaelis Say, 1818
= A.[lpheus] heterochlis Say, 1818: 243. [Fort Saint George Inlet, 8 miles south of Amelia Island, Du-
val County, Florida, intertidal; neotype selection by McClure, 1995]
= Halopsych lutaria de Saussure, 1857b: 100. [les ctes de Cuba]
= Alpheus andronyx Christofersen, 1998: 358. [nomen nudum]
Alpheus heurteli Coutire, 1897d
= Alpheus Heurteli Coutire, 1897d: 305. [baie de Fernando-Velosa]
Alpheus hippothoe De Man, 1888a
= Alpheus Hippotho De Man, 1888a: 268; Plate 17, Figs 1-5. [Sullivan Island and King Island Bay]
Alpheus holthuisi Ribeiro, 1964: 1; Figs 1-11. [St. Anto, Baha de Porto-Novo, 12 m; So Vicente, Baa de
Porto Grande, 3-5-11 m]
Alpheus homochirus (Yu, 1935)
= Grangon [sic] homochirus Yu, 1935: 61; Figs 1-2. [Weihaiwei]
Alpheus hoonsooi W. Kim & Abele, 1988: 18; Fig. 7. [Academy Bay, Indefatigable Island, Galapagos Is-
lands]
Alpheus hoplocheles Coutire, 1897e: 197. [China, Amoy; lectotype designation by Fransen, Holthuis &
Adema, 1997]
Alpheus hortensis Wicksten & McClure, 2003: 26; Figs 1-3. [Stetson Bank, 2810'N 9417'W, 21-24 m]
Alpheus hululensis Coutire, 1905b
= Alpheus Bouvieri var. Hululensis Coutire, 1905b: 908; Plate 85, fgs 46-46b. [Hulule Male Atoll]
Alpheus hutchingsae Banner & Banner, 1982a: 191; Fig. 58. [Lizard Island, Queensland]
Alpheus hyeyoungae W. Kim & Abele, 1988: 75; Fig. 31. [northeast beach of Isla Jaltimba, of Rincon de los
Guayabitas, Nayarit, Mexico]
Alpheus hyphalus Chace, 1988: 31; Figs 4-5. [Philippines, Verde Island Passage, 1338'45''N 12101'12''E,
296 m]
Alpheus idiocheles Coutire, 1905b: 883; Plate 75, Figs 21-21c. [Minikoi]
= Alpheus baculifer Coutire, 1908: 16. [Ile du Coin, Chagos]
Alpheus immaculatus Knowlton & Keller, 1983: 354; Figs 1-4. [East Discovery Bay, Jamaica, near the cen-
tral harbor buoy at approx. 10 m, from the anemone Bartholomea annulata]
Alpheus inca Wicksten & Mndez G., 1981: 137; Figs 1-16. [Baha de los Mejilones, Chile, 228'S 70.5'W]
Alpheus inopinatus Holthuis & Gotlieb, 1958: 42; Figs 8-9. [Herzliya, litoral, Israel]
Alpheus intrinsecus Spence Bate, 1888: 557; Plate 100, fg. 1. [Challenger (no stn), of Bahia, 7-20 fms]
Alpheus japonicus Miers, 1879: 53. [346'N 13615'E, 11 fms; 357'N 13655'E, 3 fms; holotype designation
by Banner & Banner, 1984 not valid]
= Alpheus longimanus Spence Bate, 1888: 551; Plate 98, fg. 4. [Challenger (no stn), of Yokoska, Japan,
5-20 fms; Challenger stns 233, (3439'0"N 13514'0"E) of Kob, Japan, 8 fms; 233a (3438'0"N
1351'0"E), of Kob, Japan, 50 fms]
Alpheus javieri Anker, Hurt & Knowlton, 2009: 7; Figs 2, 5C-D. [Panama, Pacifc Coast, Ro Mar, rocky
intertidal, under rocks and from crevices in rocky reef platform]
Alpheus kagoshimanus Hayashi & Nagata, 2000: 1111; Figs 1-5. [Of Tarumizu, Kagoshima Bay, 223-
225 m]
Alpheus kuroshimensis Nomura & Anker, 2005: 119; Figs 7-8, 13G-H. [Cyan, Kuro-shima, Yaeyama Is-
Alpheus labis Banner & Banner, 1982a: 127; Fig. 35. [Albany Passage area, Torres Straits, Queensland]
Alpheus lacertosus W. Kim & Abele, 1988: 68; Fig. 28. [Algarrobo, CA, 7 km south of Valparaiso, Val-
paraiso Province, Chile]
Alpheus ladronis Banner, 1956: 360; Fig. 20. [Saipan]
Alpheus lanceloti Coutire, 1905b
= Alpheus Lanceloti Coutire, 1905b: 900; Plate 83, fg. 39-39e. [North Male Atoll; Hulule Male Atoll;
Goidu Atoll]
Alpheus lanceostylus Banner, 1959: 136; Fig. 3. [Pearl and Hermes Reef, Hawaiian Archipelago, 27N
146W]
Alpheus latus W. Kim & Abele, 1988: 88; Fig. 37. [west side of Panama Canal, Panama, in mangroves]
Alpheus lentiginosus Anker & Nizinski, 2011: 50; Figs 1-3. [North Atlantic Ocean, Gulf of Mexico, 188.8
km south of Louisiana, 2811'41"N 8947'59"W, depth 434-438 m, base of Lophelia pertusa]
Alpheus lepidus De Man, 1908d: 106. [Madura-bay and other localities in the southern part of Molo-strait,
54-90 m; 058'.5N 12255'E, Kwandang-bay-entrance, 75 m]
Alpheus leptocheles Banner & Banner, 1975b: 261; Fig. 1. [mouth of Sepik River, New Guinea, between
Cape Girgir and Kaup, 1.5-10 fms]
Alpheus leptochiroides De Man, 1909a: 110. [536'5S 13255'.2E, 2.3 miles N, 63W from the north point of
Nuhu Jaan, Kei-islands, 90 m]
Alpheus leptochirus Coutire, 1905b: 914; Plate 87, Figs 54-54e. [North Male Atoll, Mahlosmadulu Atoll]
Alpheus leviusculus Dana, 1852b
= Alpheus Edwardsii var. leviusculus Dana, 1852b: 543; Plate 34, Figs 3a-f. [Wakes Island, north Pacifc]
= Alpheus Bouvieri var. Bastardi Coutire, 1898b: 133 (partim); Fig. 1a. [cte Ouest, Nossi-Lava, baie
de Bombtock (Madagascar); Mascate; Djibouti; nec specimen from Panama (? = Alpheus hebes W.
Kim & Abele, 1988)]
Alpheus lobidens De Haan, 1849 [in De Haan, 1833-1850]
= Alpheus lobidens De Haan, 1849 [in De Haan, 1833-1850]: 179. [Japan]
= Alpheus crassimanus Heller, 1862a: 526. [Nicobaren]
= Alpheus crassimanus Heller, 1865: 107; Plate 10, fg. 2. [Nicobaren]
= Alpheus lobidens polynesica Banner & Banner, 1975a: 429; Fig. 3A-H, J-L. [outer side of Heeia Fish
Pond, Kaneohe Bay, Oahu, Ahwaii, 2126'23''N 15748'29''W]
Alpheus longecarinatus Hilgendorf, 1879: 833; Plate 4, Figs 3-7. [Zanzibar]
Alpheus longichaelis Carvacho, 1979: 459; Figs 7-8. [Grand-Cul-de-Sac]
Alpheus longiforceps Hayashi & Nagata, 2002: 79; Figs 3-6. [Japan, Kii Strait, Wakanoura Bay, Wakayama
Prefecture, 35 m]
Alpheus longinquus W. Kim & Abele, 1988: 65; Fig. 27. [Braithwaite Bay, Socoro Island, Mexico]
Alpheus lotini Gurin, 1829 [in Gurin-Mneville, 1829-1838]
= Alpheus Lotini Gurin, 1829 [in Gurin-Mneville, 1829-1838]: Plate 3, fg. 3 (1829); 38 (1838).
[Nouvelle-Irlande]
= Cancer (Astacus) sublucanus Forskl, 1775: 94. [Djidd; suppressed under the plenary powers for
the purposes of the Principle of Priority but not those of the Principle of Homonymy in Opinion
1367 in 1987]
= A.[lpheus] ventrosus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 352. [les ctes de
lIle-de-France]
= A.[lpheus] lvis Randall, 1840: 141. [Sandwich Islands]
= Alpheus Thetis White, 1847a: 75. [nomen nudum]
= Alpheus Rouxii Gurin-Mneville, 1857: 50. [Bombay]
= Crangon latipes Banner, 1953: 82; Fig. 27. [of Waikiki Reef, Oahu, 20 f]
Alpheus lutosus Anker & De Grave, 2009: 812; Figs 1-18. [Kuwait, Shat-Al-Arab delta, Bubiyan Island,
Ras Al-Barshah and area bordering Khor Al-Subbiya, approx. 2936'N 48188'E, muddy intertidal,
in burrow with gobies]
Alpheus macellarius Chace, 1988: 35; Figs 6-7. [vicinity of Cebu City, Cebu, Philippines]
Alpheus macrocheles (Hailstone, 1835b)
= Hipplyte rbra Hailstone, 1835a: 272; Fig. 31. [of Hastings]
= Hipplyte rbra Westwood, 1835b: 274. [of Hastings]
= Hipplyte macrochles Hailstone, 1835b: 395. [nomen novum for Hipplyte rbra Hailstone, 1835a
nec Cryptophthalmus ruber Rafnesque, 1814]
= A.[lpheus] Edwardsii H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 352. [la Mditer-
rane]
= Alphaeus latimanus Hope, 1851: 43. [Type locality not indicated]
= Alpheus afnis Guise, 1854: 280; Figs 1-6. [Herm, Channel Islands, tide pool at low water]
= Alpheus Milnei Gurin-Mneville, 1857: 49. [nomen novum for A.[lpheus] Edwardsii H. Milne Ed-
wards, 1837 [in H. Milne Edwards, 1834-1840 [nec Athanas Edwarsii Audouin, 1826]
= Alpheus platyrhynchus Heller, 1862b: 400; Plate 1, Figs 21-24. [im adriatischen Meere, Lissa und
Lesina]
= Alpheus megacheles Norman, 1868: 175. [lapsus for Hipplyte macrochles Hailstone, 1835b]
= Nika platyura Fischer, 1872: 419. [Cap Breton (Landes), 28 brasses]
Alpheus macroskeles Alcock & Anderson, 1899: 153. [Investigator stn 129, Bay of Bengal, of Godvari
Delta, 270 fms; 162, of Pulicat, 1351'12''N 8028'12''E, 145-250 fms, the Swatch, 193 fms; Andaman
Sea, 193 fms]
Alpheus maindroni Coutire, 1898b
= Alpheus Maindroni Coutire, 1898b: 133; Figs 2-2. [Mascate; Dibouti]
Alpheus malabaricus (Fabricius, 1775)
= Astacus Malabaricus Fabricius, 1775: 415. [ad litora Malabarica]
= Alpheus forceps White, 1847a: 75. [nomen nudum]
= Alpheus dolichodactylus Ortmann, 1890: 473; Plate 36, fg. 11. [Japan, Tokiobai]
= Alpheus macrodactylus Ortmann, 1890: 473; Plate 36, fg. 10, 10l. [Sydney]
= Alpheus dolichodactylus var. leptopus De Man, 1910a: 289. [715'S 11515'E, 289 m; Bay of Labuan
Tring, west coast of Lombok, 844'.5S 1162'.5E, 18-27 m; Saleyer-anchorage and surroundings, 18-
45 m; Sapeh-bay, East coast of Sumbawa, up to 36 m]
= Alpheus malabaricus mackayi Banner, 1959: 149; Fig. 12. [Wailupe fsh pond, Oahu, Hawaii]
= Alpheus malabaricus songkla Banner & Banner, 1966a: 147; Fig. 56. [Lake Songkla, 1 m]
= Alpheus malabaricus trefzae Banner & Banner, 1982a: 207; Fig. 64. [Brammo Bay, Dunk Island, near
Tully, northern Queensland, 0 tide level]
Alpheus malleator Dana, 1852a: 23. [in portu Rio Janeiro?]
= Alpheus pugilator A. Milne-Edwards, 1878: 229. [toutes les les du Cap-Vert]
? = Alpheus tuberculosus Osorio, 1892: 201. [Iogo-Iogo]
= Alpheus malleator var. edentatus Zimmer, 1913: 387; Figs G-M. [Barbados (Rif)]
Alpheus malleodigitus (Spence Bate, 1888)
= Betus malleodigitus Spence Bate, 1888: 565; Plate 101, fg. 5. [Levuka, Fii Islands, reefs]
= Alpheus obesomanus var. japonicus Ortmann, 1890: 478. [Japan, Tokiobai]
= Alpheus Danae Coutire, 1905b: 887; Plate 72, fgs 26-26d. [Hulule Male Atoll]
= Alpheus phrygianus Coutire, 1905b: 886; Plate 72, fgs 25-25e. [Hulule Male Atoll; Suvadiva Atoll;
Madagascar; lle Tague]
= Alpheus persicus Nobili, 1905e: 238. [Golfe Persique]
= Alpheus malleodigitus var. gracilicarpus De Man, 1909a: 99. [Bank between islands of Bahuluwang
and Tambolungan, south of Saleyer, 8-10 m; Pulu Sanguisiapo, Tawi-Tawi-islands, Sulu archipela-
go, 12 m; 67'.5N 12026'E, anchorage of North-Ubian, 16-23 m; 825'.2S 12718'.4E, anchorage be-
tween Nusa-Besi and the N.E. point of Timor, 27-54 m]
Alpheus martini W. Kim & Abele, 1988: 98; Fig. 41. [Venado Beach, Panama]
Alpheus mazatlanicus Wicksten, 1983a: 46; Figs 7-8. [Laguna Caimaero, Sinaloa, Mxico]
Alpheus microrhynchus De Man, 1897
= Alpheus microrhynchus De Man, 1897: 754; Plate 36, Figs 65-65b (as Alpheus sp.). [Pontianak, West-
kste von Borneo]
Alpheus microscaphis (Banner, 1959)
= Thunor microscaphis Banner, 1959: 151; Fig. 13. [Likiep Atoll, Marshall Islands]
Alpheus microstylus (Spence Bate, 1888)
= Betus microstylus Spence Bate, 1888: 566; Plate 101, fg. 6. [Albany Island, Cape York]
= Alpheus microstylus var.? De Man, 1911d: 345. [Anchorage of Pulu Kawassang, Paternoster-is-
lands, coralreef]
Alpheus miersi Coutire, 1898c
= Alpheus rapax var. Miersi Coutire, 1898c: 166; Figs 1-1b. [Port-Molle; Ceylan; Japon, 3249'N
12854'E]
? = Alpheus gracilipes var. serratus Coutire, 1898a: 166. [Type locality not indicated]
Alpheus migrans Lewinsohn & Holthuis, 1978: 76; Figs 1-2. [Mediterranean, of the north coast of Sinai
Peninsula, 3126'N 3411'E, 36.6 m]
Alpheus millsae Anker, Hurt & Knowlton, 2007b: 50; Figs 3-4, 5C, 6C, 7G-H, 8C. [Panama, Panama City,
of Casco Viejo, near extreme low tide mark]
Alpheus mitis Dana, 1852a: 22. [in freto Balabac]
Alpheus moretensis Banner & Banner, 1982a: 177; Fig. 54. [2710'S 15321'E, 8 fms]
Alpheus naos Anker, Hurt & Knowlton, 2007a: 12; Figs 7-9, 11d-e. [Panama, Pacifc coast, Amador cause-
way near Panama City, Punta Culebra, rocky shore, under large mud-covered rocks on muddy
sand, in burrow of Listriolobus sp., extreme low tide]
Alpheus nipa Banner & Banner, 1985: 21; Fig. 2. [420'N 9854'E, Straits of Malacca, roughly 60 km from
shore in the vicinity of Medan, Sumatera [sic], from foating frond of nipa palm]
Alpheus nobili Banner & Banner, 1966b: 172; nomen novum for Alpheus sp. Nobili, 1906c. [bancs dhutres
perlires entre 2510'-2455'N 5510'-5440'E]
Alpheus nonalter Kensley, 1969: 172; Fig. 15. [2934'S 3139'E, 118 m]
Alpheus normanni Kingsley, 1878a: 93. [nomen novum for Alpheus afnis Kingsley, 1878b]
= Alpheus afnis Kingsley, 1878b: 195; nec Guise, 1854. [Panama]
Alpheus notabilis Stebbing, 1915: 80; Plates 20- 21. [Delagoa Bay]
Alpheus novaezealandiae Miers, 1876a
= Alpheus nov-zealandi Miers, 1876a: 224. [New Zealand; illustrated in Miers, 1876b; lectotype
selection by Yaldwyn, 1956]
Alpheus nutingi (Schmit, 1924b)
= Crangon nutingi Schmit, 1924b: 78; Plate 2, Figs 4-6. [Pelican Island, shallow]
Alpheus oahuensis (Banner, 1953)
= Crangon oahuensis Banner, 1953: 64; Fig. 20. [of Waikiki Reef, Oahu, 20f]
Alpheus obesomanus Dana, 1852a
= Alpheus obeso-manus Dana, 1852a: 21. [in archipelago Viti]
= Alpheus Lutini Coutire, 1905b: 885; Plate 76, Figs 24-24e. [Hulule Male Atoll; Samoa; lle Tague]
Alpheus ovaliceps Coutire, 1905b: 888; Plate 77, fgs 27-27e. [Minikoi]
Alpheus pachychirus Stimpson, 1860a: 30. [ad insulam Loo Choo]
Alpheus pacifcus Dana, 1852a: 21. [Kalama Park, about 18 miles southeast of Lahaina, Maui, 3f; neotype
selection by Banner, 1953, as Crangon pacifca]
= Alpheus gracilidigitus Miers, 1884b: 287. [Fii Islands; Toyota; Sandwich Islands]
Alpheus packardii Kingsley, 1880: 417. [Key West]
= Alpheus bermudensis Spence Bate, 1888: 547; Plate 98, fg. 3. [Bermuda, St. Thomas, West Indies]
= Alpheus beanii Verrill, 1922: 81; Fig. 7; Plate 22, fg. 1; Plate 32, fgs 1a-u. [Challenger Bank, 24 fms]
Alpheus paludicola Kemp, 1915: 303; Fig. 33; Plate 13; fgs 11-13. [Outer Channel, Lake Cilka, Orissa, In-
dia; lectotype selection by Yeo & Ng, 1996]
Alpheus panamensis Kingsley, 1878b: 192. [Acajutla, Central America; Panama]
Alpheus papillosus Banner & Banner, 1982a: 260; Fig. 80. [Sandgate, Moreton Bay, Queensland]
Alpheus paracrinitus Miers, 1881a: 365; Plate 16, fg. 6. [Goree Bay, Senegambia, 9-15 fms]
= Alpheus ascensionis Ortmann, 1893: 45. [Plankton-Expedition der Humboldt-Stifung, Ascension,
JN 200 (7.9S 14.4W) 20 m]
= Alpheus paracrinitus var. Bengalensis Coutire, 1905b: 901; Plate 82, fgs 37-37f. [Minikoi]
= Crangon togatus Armstrong, 1940: 2; Fig. 1. [Piedra Prieta Reef, Barahone, St. Domingo; Grazburys
Island, Bermuda]
Alpheus paradentipes Coutire, 1905b: 880; Plate 74, fgs 17-17d. [Miladumadulu Atoll; North Male Atoll]
Alpheus paraformosus Anker, Hurt & Knowlton, 2008b: 12; Figs 5-6, 7f-h. [Bocas del Toro, San Cristbal,
Punta Coco, shallow subtidal (1-1.5 m), from rock and coral rubble crevices]
Alpheus paralcyone Coutire, 1905b: 895; Plate 80, fgs 34-34e; Plate 81, fgs 34f-h. [South Nilandu Atoll;
Miladumadulu Atoll; North Male Atoll; Mahlos Atoll]
= Alpheus Providencei Coutire, 1908: 208. [Providence, 50-78 brasses]
= Crangon laysani Edmondson, 1925: 17; Fig. 3. [Laysan Island, shallow water]
Alpheus paralpheopsides Coutire, 1905b: 902; Plate 83, fgs 41-41e. [North Male Atoll, Suvadiva Atoll]
Alpheus parasocialis Banner & Banner, 1982a: 72; Fig. 17. [Palm Beach, Rockingham, Western Australia,
from jety piles]
Alpheus pareuchirus pareuchirus Coutire, 1905b
= Alpheus pareuchirus Coutire, 1905b: 906; Plate 84, fgs 43-43d. [North Male Atoll]
= Alpheus pareuchirus var. Leucothea De Man, 1909a: 105. [725'S 11316'E, Madura-Strait, 56 m;
823'.5S 1194'.6E, Sapeh-Strait, 70 m; Anchorage of Dongala, Palos-bay, Celebes, 36 m; South-east
side of Pearl-bank, Sulu-archipelago, 15 m]
Alpheus pareuchirus imitatrix De Man, 1909a
= Alpheus pareuchirus var. imitatrix De Man, 1909a: 106. [03'8N 13024'3E, 141 m; Between Loslos
and Broken-islands, west-coast of Salawati, 18 m; 142'.5S 13047'.5E, 32 m; Mid-channel in Solor-
strait of Kampog Menanga, 113 m]
Alpheus parvimaculatus Nomura & Anker, 2005: 122; Figs 9-10, 14A-C. [Fuki, Kuro-shima, Yaeyama Is-
Alpheus parvirostris Dana, 1852a
= Alpheus parvi-rostris Dana, 1852a: 22. [in freto Balabac]
= Alpheus lineifer Miers, 1875: 343. [Samoa Islands]
= Alpheus euchiroides Nobili, 1906b: 257. [Marutea]
= Alpheus braschi Boone, 1935: 131; Fig. 10; Plate 34. [Pago Pago, Samoa]
Alpheus parvus De Man, 1909a: 102. [Anchorage of Lirung, Salibabu island, 36 m]
Alpheus peasei (Armstrong, 1940)
= Crangon peasei Armstrong, 1940: 1. [Castle Harbor Reefs, Bermuda]
Alpheus percyi Coutire, 1908
= Alpheus Percyi Coutire, 1908: 211. [Cargados Carajos, 30 brasses]
Alpheus perezi Coutire, 1908
= Alpheus Perezi Coutire, 1908: 212. [Golfe Persique]
Alpheus perplexus Banner, 1956: 347; Fig. 13. [1.2 miles ofshore, immediately south of uncovering reef SE
of Magaha Islet, 24f]
Alpheus philoctetes De Man, 1909a
= Alpheus Philoctetes De Man, 1909a: 103. [Banda, 9-36 m]
Alpheus platydactylus Coutire, 1897d: 306. [Travailleur stn 52, au large de Madre (3230' 31N 1851'E
(of Paris)), 100 m; lectotype designation by Forest, 1965]
Alpheus platyunguiculatus (Banner, 1953)
= Crangon platyunguiculata Banner, 1953: 130; Fig. 47. [Waialua Bay, Oahu, 6f]
Alpheus polystictus Knowlton & Keller, 1985: 894; Fig. 1. [Morrocoy, Venezuela, 1-10 m]
Alpheus polyxo De Man, 1909a
= Alpheus Polyxo De Man, 1909a: 108. [Banda-anchorage, 9-45 m]
Alpheus pontederiae de Rochebrune, 1883
= Alpheus Pontederi de Rochebrune, 1883: 174. [Marigots de Leybar, Thiank, Dakar-Bango]
= Crangon langi Schmit, 1926b: 20; Fig. 63. [Banana]
? = Alpheus heterochaelis var. orientalis Vilela, 1949: 53; Fig. 3. [canal de Mantambua, porto de Pass;
rio Baboc, Canchungo; Ilha de Rubane; foz de rio Grande (all Guin Portuguesa)]
Alpheus pouang Christofersen, 1979: 324; Figs 14-15. [2339'S 4337'W, of the State of So Paulo, 120-
121 m]
Alpheus praedator De Man, 1908d: 103. [Ambon, reef-exploration]
Alpheus proseuchirus De Man, 1908d: 111. [058'.5N 12255'E, Kwandang-bay-entrance, 75 m; 058'.5N
12242'.5E, West of Kwandang-bay-entrance, 72 m]
Alpheus pseudopugnax (Banner, 1953)
= Crangon pseudopugnax Banner, 1953: 122; Fig. 44. [Kalama Park, southeast Maalea Bay, Maui, 8f]
Alpheus puapeba Christofersen, 1979: 328; Figs 16-18. [3505'S 5233'W, of the province of Buenos Aires,
115 m]
Alpheus pubescens De Man, 1908d: 109. [Anchorage of Pulu Sarassa, Postillon-islands, up to 30 m;
Haingsisi, Samau-island, Timor, 36 m; Makassar, up to 32 m; 327'S 11736'E, Borneo-bank, 59 m;
between Loslos and Broken-islands, west-coast of Salawati, 18 m; Anchorage of Pulu Jedan, East
coast of Aru-islands, 13 m]
Alpheus pugnax Dana, 1852a: 23. [ad insulam Maui Hawaiensem]
Alpheus pustulosus Banner & Banner, 1968: 143; Fig. 2. [21N 114E, 30-40 fms]
Alpheus quasirapacida Chace, 1988: 50; Fig. 12. [Of Tawitawi, Sulu Archipelago, Philppines, 511'50''N
11954'E, 18 m]
Alpheus randalli Banner & Banner, 1980: 401; Figs 1-3. [Nuka Hiva, Marquesas Islands, northwest side
Sentinelle de lEst, 18 m commensal with Amblyeleotris sp.]
Alpheus rapacida De Man, 1908d: 105. [Ruma-Kuda-bay, Roma-island, 36 m; Anchorage east of Dangar
Besar, Saleh-bay, up to 36 m]
Alpheus rapax Fabricius, 1798: 405. [in India orientali]
Alpheus rectus W. Kim & Abele, 1988: 16; Fig. 6. [Bahia Honda, Panama]
Alpheus ribeiroae Anker & Dworschak, 2004: 48; Figs 1-27. [tidal fat at Moia-Moia, NE coast of Ilha do
So Tiago, Cabo Verde]
Alpheus richardsoni Yaldwyn, 1971: 88. [of Mariot Id., Waikare River Inlet, Bay of Islands, 1-3 fms]
Alpheus rudolphi Almeida & Anker, 2011: 1; Figs 1-22. [REVIZEE Program, Score Nordeste III stn 171, of
Alagoas, Brazil, 955.11'S 3532.73'W, 49 m]
Alpheus romensky Burukovsky, 1990: 197; Fig. 3A.1-6. [2534'S 8904'W, 540-560 m]
Alpheus roquensis Knowlton & Keller, 1985: 897; Fig. 1. [Los Roques, Venezuela, 3-16 m]
Alpheus roseodigitalis Nomura & Anker, 2005: 126; Figs 10-11, 14D-F. [Aragusuku-jima, Yaeyama Islands,
southern Ryukyu Archipelago, Japan, intertidal]
Alpheus rostratus W. Kim & Abele, 1988: 51; Fig. 21. [Darwin Bay, Tower Island, Galapagos Islands]
Alpheus rugimanus A. Milne-Edwards, 1878: 230. [les les du Cap-Vert]
Alpheus samoa Banner & Banner, 1966b: 174. [Lions Head, Tutuila, American Samoa]
Alpheus savuensis De Man, 1908d: 110. [Anchorage of Seba, Savu]
Alpheus saxidomus Holthuis, 1980d: 47; Figs 1-2. [Playa Tamarindo, Nicoya Peninsula, Pacifc coast of
Costa Rica, about 1019'N 8553'W, boring in basaltic rock in the surf zone]
Alpheus schmiti Chace, 1972: 70; Figs 21-22. [Antigua Island, Falmouth Harbour, beach north of Blacks
Point, 2-3 f]
Alpheus scopulus W. Kim & Abele, 1988: 77; Fig. 32. [Whorehouse reef tidepools, Panama]
Alpheus serenei Tiwari, 1964b: 314. [St. Cauda, 3-4 m; fully described in Tiwari, 1965]
Alpheus sibogae De Man, 1908d: 107. [823'.5S 1194'.6E, Sapeh-strait, 70 m; Madura-bay and other lo-
calities on the southern part of Moho-strait, 54-90 m; Mid-channel in Solor-strait of Kampong Me-
nanga, 113 m]
Alpheus simus Gurin-Mneville, 1855 [in Gurin-Mneville, 1855-1856]
= Alpheus Simus Gurin-Mneville, 1855 [in Gurin-Mneville, 1855-1856]: xix; Plate 2, fg. 11. [Cuba]
= Crangon rathbunae Schmit, 1924b: 74; Plate 1; nec Alpheus Rathbuni Coutire, 1900b. [Needhams
Point, Barbados]
Alpheus sizou Banner & Banner, 1967
= Alpheus amirantei sizou Banner & Banner, 1967: 265. [north side of Bora Bora, outer reef platform, 1f]
Alpheus socialis Heller, 1862a
= Alpheus socialis Heller, 1862a: 526. [Aukland]
= Alpheus socialis Heller, 1865: 106; Plate 10, fg. 1. [Auckland; Sidney]
= Alpheus Doto White, 1847a: 75 [nomen nudum]
Alpheus soelae Banner & Banner, 1986: 21; Fig. 1. [1841'S 12007'E, 430 m]
Alpheus soror Bruce, 1999: 454; Figs 1A, 2-3. [of Trincomalee, Sri Lanka]
Alpheus spatulatus Banner & Banner, 1968: 146; Fig. 3. [1917'N 11281'E, 52 fms]
Alpheus splendidus Coutire, 1897c: 236. [Djibouti]
= Alpheus pomatoceros Banner & Banner, 1966a: 93; Fig. 32. [Lam Chongklan, Koh Samui, Thailand,
outer reef front, 2-3 m]
Alpheus spongiarum Coutire, 1897c: 236. [Djibouti]
= Alpheus paraculeipes Coutire, 1905b: 894; Plate 79, fgs 32-32b'; Plate 80, fgs 32c-32g. [South Ni-
landu Atoll; Mahlosmadulu Atoll; North et South Male Atoll]
Alpheus stanleyi Coutire, 1908
= Alpheus Stanleyi Coutire, 1908: 207. [Amirante, 30 brasses]
= Alpheus Stanleyi var. dearmatus De Man, 1910a: 287. [Wunoh-bay, N.W. coast of Waigeu-island,
reef; Banda-anchorage, 9-36 m]
= Alpheus cloudi Banner, 1956: 352; Fig. 16. [about 0.6 mile of Muchot Point, between reef clusters at
south side of Saipan harbor, 5-15f]
Alpheus stantoni Banner & Banner, 1986: 24; Fig. 2. [Heron Island, Great Barrier Reef, dug out of coral
rubble, 13 m]
Alpheus staphylinus Coutire, 1908: 204. [Salomon Island, Chagos]
Alpheus stephensoni Banner & Smalley, 1969: 43; Fig. 2. [Moreton Bay, Queensland, 4-5 fms]
Alpheus strenuus strenuus Dana, 1852a: 21. [insula Tongabatu]
= Alpheus Alope White, 1847a: 74. [nomen nudum]
= Alpheus Rhode White, 1847a: 74. [nomen nudum]
= Alpheus Doris White, 1847a: 75. [nomen nudum]
= Alpheus strenuus var. angulatus Coutire, 1905b: 914. [Hulule Male Atoll]
Alpheus strenuus cremnus Banner & Banner, 1982a: 229; Fig. 72. [Minnie Waters, near Grafon, New South
Wales, intertidal rock platform]
Alpheus styliceps Coutire, 1905b: 889; Plate 78, fgs 28-28d. [Naifaro Reef]
Alpheus sudara Banner & Banner, 1966a: 153; Fig. 59. [Phuket, Thailand, outer reef edge, 2 m]
Alpheus sulcatus Kingsley, 1878b: 193. [Bay of Panama; Zorritas, Peru]
= Alpheus macrochirus Richters, 1880: 164; Plate 17, fgs 31-33. [Fouquets]
= Alpheus luci Barnard, 1947: 389. [St. Lucia Bay, Zululand]
Alpheus suluensis Chace, 1988: 57; Figs 15-16. [near Siasi, Sulu Archipelago, 541'40''N 12047'10''E, 38 m]
Alpheus supachai Banner & Banner, 1966a: 89; Fig. 30. [Koh Samui, west side of Gulf of Thailand, 3 m]
Alpheus superciliaris Coutire, 1905b: 896; Plate 81, fgs 35-35g. [Reef Naifaro]
Alpheus talismani Coutire, 1898d
= Alpheus Talismani Coutire, 1898d: 31; Figs 3-4. [les du Cap-Vert, 410-450 m]
Alpheus tasmanicus Banner & Banner, 1982a: 215; Fig. 67. [Tasmania]
Alpheus tenuicarpus De Man, 1908d: 104. [058'.5N 12255'E, Kwandang-bay-entrance, 75 m; Sapeh-bay,
East coast of Sumbawa up to 36 m]
Alpheus tenuipes De Man, 1910a: 288. [420'S 12258'E, between islands of Wowoni and Buton, northern
entrance of Buton-Strait, 75-94 m]
Alpheus tenuis W. Kim & Abele, 1988: 79; Fig. 33. [Farfan Point, Panama]
Alpheus thomasi Hendrix & Gore, 1973: 415; Figs 1-3. [Virginia Beach, Virginia Key, Miami, Florida, 1 m]
Alpheus tirmiziae Kazmi, 1974: 170; Figs 1-4. [Bay of Bengal, 2005'-2030'N 9150'-9050'E]
Alpheus tricolor Anker, 2001: 58; Figs 1a-b, 2a, 3a-e, 4a-d, 5a-h, 6a-k. [Bali, Indonesia]
Alpheus triphopus Nobili, 1906b
= Alpheus aculeipes var. triphopus Nobili, 1906b: 257. [Marokau]
Alpheus tungii Banner & Banner, 1966b: 160. [south coast of Tongatabu, Tonga, outer edge of fringing
reef near surge channel]
Alpheus umbo W. Kim & Abele, 1988: 84; Fig. 35. [Isabel Island, Sinaloa, Mexico]
Alpheus utriensis Ramos & von Prahl, 1989: 477; Figs 1-2. [Utria Sound]
Alpheus vanderbilti Boone, 1930c: 163; Plate 58, fg. 5. [south of Sand Key, Key West, Florida, 20 fms]
Alpheus villosus (Olivier, 1811)
= Palmon villosus Olivier, 1811: 664. [la mer des Indes]
= Palmon diverfmanus Olivier, 1811: 663. [ctes de la Nouvelle-Hollande]
Alpheus villus W. Kim & Abele, 1988: 82; Fig. 34. [spit north of mill site, Angeles Bay, Baja California,
Mexico]
Alpheus viridari (Armstrong, 1949)
= Crangon viridari Armstrong, 1949: 8; Fig. 2. [inside North end of El Cayo, Dominican Republic]
Alpheus waltervadi Kensley, 1969: 175; Fig. 16. [Walters Shoal, 3313'S 4351'E, 38-46 m]
Alpheus websteri Kingsley, 1880: 416. [Key West]
= Alpheus Ridleyi Pocock, 1890: 518. [Fernando de Noronha]
= Alpheus nigro-spinatus Rankin, 1898: 249; Plate 30, fg. 6. [New Providence]
Alpheus wickstenae Christofersen & Ramos, 1987: 333; Figs 1-3. [Rampa de los Suecos, Baha Mlaga,
Pacifc coast of Colombia, 359'N 7720'W]
Alpheus williamsi Bruce, 1994b: 19; Figs 3, 5C. [north of Charles Point, Northern Territory, Australia,
1217.18'S 3040.06'N, 18-24 m]
Alpheus xishaensis Liu & Lan, 1980: 102; Fig. 13. [Xisha Island, coral reef]
Alpheus xanthocarpus Anker, Hurt & Knowlton, 2008a: 556; Figs 6, 7E-F. [So Tom, Ilha Santana, 15-20 m]
Alpheus zimmermani Anker, 2007a: 242; Figs 1-3. [Caribbean Sea, British Virgin Islands, Guana island,
near end of Long Point, 1.5-11 m]
Alpheus zulfaquiri Kazmi, 1982: 137; Figs 1-2. [Manora Island, coastal waters of Karachi, northern Ara-
bian Sea]
Amphibetaeus Coutire, 1897b
= Amphibetus Coutire, 1897b (type species Betus Jousseaumei Coutire, 1896, by monotypy, gen-
der masculine)
Amphibetaeus jousseaumei (Coutire, 1896)
= Betus Jousseaumei Coutire, 1896: 313; Figs 1-12. [Perim; Tadjourah]
= Amphibetus Jousseaumei Coutire, 1897b: 384. [nomen novum for Betus Jousseaumei Coutire, 1896]
Arete Stimpson, 1860a
= Arete Stimpson, 1860a (type species Arete dorsalis Stimpson, 1860a, by monotypy, gender feminine;
Arete acanthocarpus (Miya & Miyake, 1968a)
= Athanas acanthocarpus Miya & Miyake, 1968a: 157; Figs 10D, H, J, 11C, E, 12E, 13. [Kamiyama-jima,
near Naha City, Okinawa-jima, Ryukyu Islands]
Arete dorsalis Stimpson, 1860a: 32. [Cape DAguilar, Hong Kong Island, 1 m; neotype designation by
Bruce, 1990g, as Athanas dorsalis]
? = Athanas mascarenicus Richters, 1880: 164. [Insel Mauritius und Seychellen]
= Arete dorsalis var. Pacifcus Coutire, 1903: 85; Figs 31-34. [Hong-Kong (?); Nouvelle Caldonie;
Samoa; Amrique Centrale; Amboine (??)]
= Arete Maruteensis Coutire, 1905b: 864. [Marutea]
= Arete maruteensis var. salibabuensis De Man, 1910a: 313. [Anchorage of Lirung, Salibabu-island, up
to 36 m]
Arete amboinensis De Man, 1910b: 25; Figs 1-6. [Insel Amboina]
Arete indicus Coutire, 1903
= Arete dorsalis var. Indicus Coutire, 1903: 84; Figs 25-29. [Djibouti; Hulule Male Atoll; Masca-
reignes (?)]
= Arete Indicus Coutire, 1905b: 863; Figs 134-135. [Hulule Male Atoll]
= Arete Iphianassa De Man, 1910a: 312. [Anchorage of Sawan, Siau-island, reef]
= Arete intermedius Yu, 1931b: 513; Fig. 1. [?Amoy]
? = Athanas kominatoensis Kubo, 1942c: 82; Figs 1-2. [Kominato, Prov. Awa (Bsy)
Aretopsis De Man, 1910a
= Aretopsis De Man, 1910a (type species Aretopsis amabilis De Man, 1910a, by monotypy, gender
feminine)
Aretopsis amabilis De Man, 1910a: 311. [Pulu Kaniungan ketjil, reef]
= Aretopsis aegyptiaca Ramadan, 1936: 16; Plate 1, fgs 9-10; Plate 2, fgs 9-17. [Ghardaga, Red Sea]
Aretopsis manazuruensis Suzuki, 1971: 19; Figs 10-12. [Shikkake, near the Manazuru M.B.L., intertidal]
Athanas Leach, 1814 [in Leach, 1813-1814]
= Athanas Leach, 1814 [in Leach, 1813-1814] (type species Palaemon nitescens Leach, 1813 [in Leach,
1813-1814], by monotypy, gender masculine; name placed on the Ofcial List of Generic Names in
Athanas ahyongi Anker & Komai, 2010: 46; Figs 1-6. [Madagascar, Nosy-B, west of Hellville, 1341.56'S
4824.65'E, 0.5-1 m]
Athanas amazone Holthuis, 1951a: 111; Fig. 23. [of Nigeria, 401'N 712'E, 52 m]
Athanas anatidactylus Anker & Marin, 2007: 163; Figs 1-2, 3a. [Vietnam, Nhatrang Bay, Tre Island, 7 m, on
unidentifed crinoid]
Athanas areteformis Coutire, 1903: 79; Figs 17-18. [Naifaro Reef; Hulule Male Atoll]
= Athanas Naifaroensis Coutire, 1903: 77; Figs 14-16. [Naifaro Reef; Hulule Male Atoll]
= Athanas Naifaroensis Coutire, 1905b: 859; Fig. 131. [Naifaro Reef; Hulule Male Atoll]
= Athanas aretiformis Coutire, 1905b: 860; Fig. 132. [Naifaro Reef; Fadifolu Atoll; Hulule Male Atoll]
? = Athanas crosslandi Tatersall, 1921: 372; Plate 27, fgs 13-17. [Khor Dongonab, Engineer Island,
2050'N, reef fat]
= Athanas erythraeus Ramadan, 1936: 13; Plate 1, fg. 1. [Ghardaga, Red Sea]
= Athanas dubius Banner, 1956: 322; Fig. 2. [Saipan]
Athanas daviei Anker, 2011a: 5; Figs 3-4, 8E-F. [Australia, Queensland, Great Barrier Reef, Lizard Island,
Watsons Bay, 1439'50.4''S 14527'46.8''E, 1 m]
Athanas dentirostris Anker, Jeng & Chan, 2001: 1049; Figs 1-3. [Cua Luc, N Vietnam, together with Upo-
gebia af. takaoensis]
Athanas dimorphus Ortmann, 1894: 12; Plate 1, fg. 1-1z. [Dar-es-Salaam, Upanga-Rif]
? = A.[lpheus] monoceros Heller, 1862c: 274. [im rothen Meere]
? = Athanas transitans var. longispina Czerniavsky, 1884: 25. [nomen novum for A.[lpheus] monoceros
Heller, 1862c]
= Athanas solenomerus Coutire, 1897b: 381. [mer Rouge]
= Athanas leptocheles Coutire, 1897b: 381. [mer Rouge]
= Athanas dispar Coutire, 1897c: 233. [mer Rouge]
= Athanas setoensis Kubo, 1951: 265; Figs 5-6. [vicinity of Seto Marine Biological Laboratory of Kyoto
University]
= Athanas dimorphus seedang Banner & Banner, 1966a: 28; Fig. 4. [Koh Krarard, Trad., southwest side
of island, 1.5 m]
Athanas djiboutensis Coutire, 1897c
= Athanas Djiboutensis Coutire, 1897c: 234. [Djibouti]
= Athanas sulcatipes Borradaile, 1899: 1011; Plate 65, fgs 9-9i. [Funafuti]
Athanas gracilipes Banner & Banner, 1978: 234; Fig. 3a-g. [1202'N 11249'E, 365-385 m]
Athanas gracilis Boone, 1935: 122; Plate 32. [coral at Teviatoa reef, Raiatea Island, Society Islands]
Athanas granti Coutire, 1908
= Athanas Granti Coutire, 1908: 192. [South Adelade Coast]
Athanas grimaldii Coutire, 1911b
= Athanas Grimaldii Coutire, 1911b: 1; Figs 1-6. [Belle Ile, 155 m; Iles du Cap Vert, 16 m + 91 m]
Athanas hasswelli Coutire, 1908
= Athanas Hasswelli Coutire, 1908: 192. [South Adelade Coast; emendation to A. haswelli by Banner
& Banner, 1973 is an unjustifed emendation]
Athanas hongkongensis Bruce, 1990f: 624; Figs 7-8. [Long Ke Wan, low water]
Athanas iranicus Anker, Naderloo & Marin, 2010: 54; Figs 1-4. [Iran, Hormozgan, Qeshm Island north of
Strait of Hormuz, northern coast, 2 km east of desalination center, 2656'N 5547'E, intertidal]
Athanas ivoiriensis Anker & Ahyong, 2007: 23; Figs 3-4. [Cte dIvoire, Grand Bassam, 20 m]
Athanas japonicus Kubo, 1936c: 43; Plate 8. [Mitaziri, Yamaguti Prefecture]
= Athanas lamellifer Kubo, 1940d: 102; Figs 3-5. [Kominato (Matuga-hana), Tiba Prefecture, near low
water mark]
Athanas jedanensis De Man, 1910a: 313. [Anchorage of Pulau Jedan, east coast of Aru-islands (Pearl-
banks)]
Athanas locincertus Banner & Banner, 1973a: 311; Fig. 5. [Panchoran Buoy]
Athanas marshallensis Chace, 1955: 17; Fig. 8. [Eniwetok Atoll, Bogombogo Island, intertidal]
? = Athanas esakii Kubo, 1940e: 93; Fig. 13. [Kusaie, Caroline Islands]
Athanas minikoensis Coutire, 1903: 76; Figs 9-11. [Minikoi Atoll]
= Athanas Minikoensis Coutire, 1905b: 858; Fig. 130. [Minikoi Atoll]
Athanas naga Banner & Banner, 1966a: 26; Fig. 3. [Gulf of Thailand, 1012'48''N 10332'30''E, 27-30 m]
Athanas nitescens (Leach, 1813 [in Leach, 1813-1814])
= Palaemon nitescens Leach, 1813 [in Leach, 1813-1814]: 401. [southern coast of Devonshire]
= P.[alemon] Lvirhincus Risso, 1816: 108. [environs de Nice, sur les bas fonds]
= Cancer listellus Nardo, 1847: 6. [Trovato nel golfo, tra gli Asprei]
= Alpheus ? vitatus Nardo, 1847: 6. [Trovato nel golfo, tra gli Asprei]
= Arete Diolectiana Heller, 1862b: 404; Plate 1, Figs 28-33. [Pirano]
= Athanas veloculus Spence Bate, 1888: 529; Plate 96, fg. 1. [Cape Verde Islands]
= Athanas transitans var. pontica Czerniavsky, 1884: 25. [Black Sea]
= Athanas nitescens forma rotundicauda Czerniavsky, 1884: 24; Plate 2, fgs 5A-D. [promontorium
Ivan-baba (prope Theodosia, Strembitzki)]
= Athanas alpheoides Czerniavsky, 1884: 26. [Portus Suchum, zona litor., 1-2 m]
Athanas nouvelae Holthuis, 1951a: 104; Fig. 22. [of So Pedro Bay, So Vicente, Cape Verde Islands,
22 m]
Athanas orientalis Pearson, 1905: 88; Plate 2, fgs 10-10b. [Cheval Paar, 6.5-13 fms; across Mutuvaratu
Paar, 6-9 fms; south of Adams bridge, Gulf of Manaar, 4-40 fms; northern end of Periya Paar, 12-
14 fms]
Athanas ornithorhynchus Banner & Banner, 1973a: 319; Fig. 8. [Chambers Bay, Van Diemen Gulf, Northern
Territory]
Athanas ohsimai Yokoya, 1936: 129; Fig. 1. [tide pools at Aburatubo]
Athanas parvus De Man, 1910a: 315. [839.1'S 1274.4'E, anchorage south coast of Timor, 34 m]
= Athanas Sibogae De Man, 1910a: 314. [Anchorage of Pulu Jedan, east coast of Aru-islands (pearl-
banks), 13 m]
Athanas phyllocheles Banner & Banner, 1983: 152; Figs 13-14. [of Runion, 450 m]
Athanas polymorphus Kemp, 1915: 295; Figs 31-32. [outer channel of Satpara and Barhampur Island,
Chilka Lake, 6-10 f]
Athanas rhothionastes Banner & Banner, 1960a: 142; Fig. 3. [Canton Island, Phoenix group]
Athanas shawnsmithi Anker, 2011a: 2; Figs 1-2, 8A-D. [Australia, Queensland, Great Barrier Reef, Lizard
Island, about 50 m east of Lumus Beach (in front of marine station), 1440'92.6''S 14526'95.2''E, less
than 0.5 m]
Athanas squillophilus Hayashi, 2002: 396; Figs 1-3. [Ishikari Bay, of Takashima Port, Otaru, Hokkaido, 10
m, associated with Oratosquilla oratoria]
Athanas stebbingii De Man, 1920a
= Athanas Stebbingii De Man, 1920a: 106. [Sangkapoera-roads, Bawean-island, 12 m]
Athanas sydneyensis Anker & Ahyong, 2007: 18; Figs 1-2. [mid-stream between Juno Head and Hungry
Beach, Hawkesbry River, New South Wales, Australia, 3334'S 15116'E, 10 m]
Athanas tenuipes De Man, 1910a: 316. [058.5'N 12242.5'E, west of Kwandang-by-entrance, 72 m]
Athanopsis Coutire, 1897f
= Athanopsis Coutire, 1897f (type species Athanopsis platyrhynchus Coutire, 1897f, by monotypy,
gender feminine)
Athanopsis australis Banner & Banner, 1982a: 9; Fig. 1. [Beaumaris, Port Phillip Bay, Victoria, 8 m]
Athanopsis brevirostris Banner & Banner, 1981a: 45; Fig. 5. [Cundabilu Island, Dahlak Archipelago, south-
ern Red Sea, 0.3 m]
Athanopsis dentipes Miya, 1980: 118; Figs 1-3. [in front of the Aquaculture Research Laboratory, Nagasaki
Prefectural Institute of Fisheries, Nomo Bay, Nomozaki, Nagasaki Peninsula, LWS, in burrow of
Thalassema ?mucosa]
Athanopsis platyrhynchus Coutire, 1897f: 301. [Djibouti]
Athanopsis rubricinctuta Berggren, 1991: 166; Figs 1-6. [Inhaca Island, Moambique]
Athanopsis saurus Anker, 2011a: 9; Figs 5-7, 8G-H. [Australia, Queensland, Great Barrier Reef, Lizard Is-
land, Casuarina Beach near marine station, 1440'31.4''S 14526'39.5''E, 1 m]
Automate De Man, 1888b
= Arethusa De Man, 1887 (nomen nudum)
= Automate De Man, 1888b (type species Automate dolichognatha De Man, 1888b, by monotypy, gen-
Automate anacanthopus De Man, 1910a: 317. [Kwadang-bay entrance, 058'.5N 12255'E, 75 m; Sanana-
bay, East coast of Sula Besi, 22 m]
Automate branchialis Holthuis & Gotlieb, 1958: 34; Figs 5-6. [of Gaza]
Automate dolichognatha De Man, 1888b: 529; Plate 22, fg. 5-5i. [Insel Noordwachter]
= Automate Gardineri Coutire, 1902b: 337. [Minikoi Atoll; Madhu Atoll; Goidu Atoll; Hulule Atoll]
= Automate kingsleyi Hay, 1917: 72. [Beaufort, North Carolina]
= Automate haight Boone, 1931: 184; Fig. 22. [north shore of Tabogilla Island, Pearl Islands, Bay of
Panama]
= Automate johnsoni Chace, 1955: 13; Fig. 7. [Bikini Atoll]
Automate evermanni Rathbun, 1901: 112; Fig. 22. [of Aguadilla, 137 fms]
Automate hayashii Anker & Komai, 2004: 1897; Figs 1-4. [tidal fat at Kamiiso, Hakodate Bay, southern
Hokkaido, Japan, intertidal]
Automate rectifrons Chace, 1972: 75; Fig. 24. [Antigua Island, north side of Bird Island, Nonsuch Bay, 1-4 f]
Automate rugosa Coutire, 1902b: 341. [844N 7909W, baie de Panama, 30 fms]
Automate salomoni Coutire, 1908
= Automate Salomoni Coutire, 1908: 192. [Chagos, Salomon Island]
Automate talismani Coutire, 1902b: 340. [Puerto-Grande, Aores, 20 m]
Bannereus Bruce, 1988f
= Bannereus Bruce, 1988f (type species Bannereus anomalus Bruce, 1988f, by original designation and
Bannereus anomalus Bruce, 1988f: 140; Figs 1-6. [Marian Plateau, Coral Sea, 2235.3'S 15346.7'E to
2236.3'S 15350.0'E, 345-350 m]
Batella Holthuis, 1955b
= Cheirothrix Spence Bate, 1888 (type species Cheirothrix parvimanus Spence Bate, 1888, by monotypy,
gender feminine; invalid junior homonym of Cheirothrix Pictet & Humbert, 1866 (Pisces))
= Batella Holthuis, 1955b (nomen novum for Cheirothrix Spence Bate, 1888, gender feminine)
Batella leptocarpus Chace, 1988: 65; Fig. 17. [Philippines, western Mindanao Sea, 847'15''N 12335'00''E,
296 m]
Batella parvimanus (Spence Bate, 1888)
= Cheirothrix parvimanus Spence Bate, 1888: 533; Plate 96, fg. 2-2b. [Challenger stn 186, 1030'S
14218'E, of Cape York, 8 fms]
= Batella bifurcata Miya & Miyake, 1968b: 116; Figs 2-4. [near Dajo Group, Nagasaki Prefecture, Ja-
pan, 3214.0'N 12750.4'E, 156 m]
Batella praecipua De Grave, 2004a: 504; Figs 1-3. [New Caledonia, 2253'8"S 16713'9"E, 425-440 m, dans
une ponge avec des stnopodides]
Bermudacaris Anker & Ilife, 2000
= Bermudacaris Anker & Ilife, 2000 (type species Bermudacaris harti Anker & Ilife, 2000, by original
Bermudacaris australiensis Anker & Komai, 2004: 1903; Figs 5-7. [north-west shelf, Western Australia,
1954.0'S 11752.2'E, 38 m]
Bermudacaris britayevi Anker, Poddoubtchenko & Marin, 2006: 1677; Figs 1-4. Vietnam, Nha Trang Bay,
Tre Island, Dam Bay, intertidal]
Bermudacaris harti Anker & Ilife, 2000: 762; Figs 1-4. [Bermuda Island, Christies Cave]
Betaeopsis Yaldwyn, 1971
= Betaeopsis Yaldwyn, 1971 (type species Betus quimanus Dana, 1852a, by original designation,
gender feminine)
= Hamalpheus Bruce & Ilife, 1991 (type species Hamalpheus acanthops Bruce & Ilife, 1991, by original
Betaeopsis acanthops (Bruce & Ilife, 1991)
= Hamalpheus acanthops Bruce & Ilife, 1991: 584; Figs 1-5. [Tosua-Tolesua lave tube cave, near Lotof-
aga village, south coast of Upolu Island, Western Samoa]
Betaeopsis aequimanus (Dana, 1852a)
= Betus quimanus Dana, 1852a: 23. [in portu Bay of Islands, ad insulas Black Rocks, Novi-
Zealandi]
Betaeopsis indica (De Man, 1910a)
= Betaeus indicus De Man, 1910a: 309. [Anchorage of Labuan Pandan, Lombok, 18 m]
Betaeus Dana, 1852a
= Betaeus Dana, 1852a (type species Betus truncatus Dana, 1852a, designated by Fowler, 1912, gen-
der masculine; name placed on the Ofcial List of Generic Names in Zoology in Opinion 470 in
1957)
Betaeus australis Stimpson, 1860a: 31. [Long Reef, Collaroy, New South Wales, intertidal rock platform;
neotype selection by Banner & Banner, 1973a]
Betaeus emarginatus (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]) (Fig. 45)
= A.[lpheus] emarginatus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 357. [Type local-
ity not indicated]
= Alpheus lvigatus Gurin-Mneville, 1838 [in Gurin-Mneville, 1829-1838]: 38. [lle de Timor]
= Betus scabro-digitus Dana, 1852a: 23. [juxta urbem Valparaiso Chilensem]
Betaeus ensenadensis Glassell, 1938: 416; Plate 28. [Estero de la Punta Banda, Ensenada, Baja California,
Mexico, low tide]
Betaeus gelasinifer Nomura & Komai, 2000: 45; Figs 1-5. [Kominato, Boso Peninsula, intertidal]
Betaeus gracilis J.F.L. Hart, 1964: 453; Figs 50-51, 56, 65-67, 77-78. [Laguna Beach]
Betaeus granulimanus Yokoya, 1927: 173; Plate 7, fgs 17-22. [Bay of Kagoshima]
= Betaeus yokoyai Kubo, 1936b: 50; Plate 15. [near low tide mark at Kominato]
= Betaeus murayamai Yokoya, 1936: 132; Fig. 2. [vicinity of the Misaki Marine Biological Station]
Betaeus harfordi (Kingsley, 1878b)
= Alpheus harfordi Kingsley, 1878b: 198. [Santa Barbara and Catalina Island, California, under the
mouth of Haliotis rufescens]
= Betus equimanus Lockington, 1877b: 43. [Catalina Island, under the mantle of Haliotis rufescens]
= Alpheus lvigatus Nicolet, 1849: 215. [costas de Chile]
= Alpheus qualis Kingsley, 1878b: 199. [nomen novum for Betus equimanus Lockington, 1877b]
Betaeus harrimani Rathbun, 1904
= Betus harrimani Rathbun, 1904: 108; Fig. 49. [Sitka]
Betaeus jucundus Barnard, 1947
= Betus jucundus Barnard, 1947: 388. [Keurbooms River estuary, Pletenberg Bay]
Betaeus levifrons Vinogradov, 1950: 198; Plate 10, fg. 32. [Zolotoi Rog Bay, near Vladivostok region, Rus-
sian Far East]
Betaeus lilianae Boschi, 1966: 84; Figs 1-3. [Quequn, provincia de Buenos Aires, entre algas en la playa]
Betaeus longidactylus Lockington, 1877a: 35. [San Diego, sandy mud fat, between tides]
Betaeus macginitieae J.F.L. Hart, 1964: 451; Figs 48-49, 55, 62-64, 75-76. [California, Corona del Mar, of
Ladder Rock, 4 fms]
Betaeus pingi Yu, 1930a
= Betaeus Pingi Yu, 1930a: 454; Figs 1-3. [Chefoo?]
Betaeus setosus J.F.L. Hart, 1964: 455; Figs 52-53, 57, 68-72, 79-80; Plate 2. [British Colombia, Vancouver
Island, west coast, Clayoquot Sound]
Betaeus truncatus Dana, 1852a
= Betus truncatus Dana, 1852a: 23. [in mari prope insulam Hermite]
= Alpheus Sinuosus Gurin-Mneville, 1857: 51. [Callao, au Chili]
Bruceopsis Anker, 2010b
= Bruceopsis Anker, 2010b (type species Bruceopsis projectus Anker, 2010b, by original designation,
gender feminine)
Bruceopsis projectus Anker, 2010b: 392; Figs 1A-B, 2-5. [Mariana Islands, Guam, Agat Bay north of Alu-
tom Islet, under rocks, 3-7 m]
Bruceopsis guamensis Anker, 2010b: 399; Figs 1C, 6-8. [Marian Islands, Guam, Agat Bay north of Alutom
Islet, under rocks, 3-7 m]
Coronalpheus Wicksten, 1999
= Coronalpheus Wicksten, 1999 (type species Coronalpheus natator Wicksten, 1999, by original desig-
Coronalpheus natator Wicksten, 1999: 106; Figs 1-4. [Devils Crown, Isla Onslow, north of Isla Floreana, 15 m]
Coutieralpheus Anker & Felder, 2005
= Coutieralpheus Anker & Felder, 2005 (type species Coutieralpheus setirostris Anker & Felder, 2005,
by original designation and monotypy, gender masculine)
Coutieralpheus setirostris Anker & Felder, 2005: 42; Figs 1-6. [south margin of Fort Pierce Inlet channel,
beside US Highway A1A South Causeway, Indian River Lagoon, St. Lucie County, Florida, 2727.7'N
8018.7'W, intertidal sand fat]
Deioneus Dworschak, Anker & Abed-Navandi, 2000
= Deioneus Dworschak, Anker & Abed-Navandi, 2000 (type species Deioneus sandizelli Dworschak,
Anker & Abed-Navandi, 2000, by original designation and monotypy, gender masculine)
Deioneus sandizelli Dworschak, Anker & Abed-Navandi, 2000: 304; Figs 1-26. [Cape Verde Islands, Ilha
do So Tiago, NE-coast, tidal fat at Moia-Moia, with Corallianassa intesi]
Fenneralpheus Felder & Manning, 1986
= Fenneralpheus Felder & Manning, 1986 (type species Fenneralpheus chacei Felder & Manning, 1986,
Fenneralpheus chacei Felder & Manning, 1986: 498; Figs 1-3. [Florida, St. Lucie County, Indian River la-
goon, Fort Pierce Inlet, 2727.7'N 8018.7'W]
Harperalpheus Felder & Anker, 2007
= Harperalpheus Felder & Anker, 2007 (type species Harperalpheus pequegnatae Felder & Anker, 2007,
Harperalpheus pequegnatae Felder & Anker, 2007: 457; Figs 1-4. [northwestern Gulf of Mexico, southeast
of Galveston Island, Texas, USA, near 2918.5'N 9440.0'W, about 5 km ofshore, 11.5-12 m]
Fig. 45. Betaeus emarginatus (H. Milne Edwards, 1837). Photo by Arthur Anker.
Jengalpheops Anker & Dworschak, 2007a
= Jengalpheops Anker & Dworschak, 2007a (type species Jengalpheops rufus Anker & Dworschak,
2007a, by original designation and monotypy, gender masculine)
Jengalpheops rufus Anker & Dworschak, 2007a: 291; Figs 1-7. [Philippines, Panglao Island, Sungcolan
Bay, 938.3'N 12349.6'E, tidal fat, from Glypturus armatus mound]
Leptalpheus Williams, 1965
= Leptalpheus Williams, 1965 (type species Leptalpheus forceps Williams, 1965, by original designa-
Leptalpheus axianassae Dworschak & Coelho, 1999: 477; Figs 1-30. [Praia do Ara, So Sebastio, So
Paulo, Brazil]
Leptalpheus denticulatus Anker & Marin, 2009: 92; Figs 1-6, 7a-b, 8. [Vietnam, Nhatrang Bay, Dam Bay,
intertidal sand fat, from burrow of Glypturus cf. armatus]
Leptalpheus dworschaki Anker & Marin, 2009: 100; Figs 9-12. [Philippines, Panglao Island, Sungcolan
Bay, 938.3'N 12349.6'E, intertidal fat, from burrow of Glypturus sp.]
Leptalpheus felderi Anker, Vera Caripe & Lira, 2006: 688; Figs 1-5, 6A-B. [Venezuela, Estado Nueva Es-
parta, Isla Margarita, southern shore near Guamache, 0.5-1 m, from burrow of Upogebia omissa]
Leptalpheus forceps Williams, 1965: 194; Figs 1-2. [Gallants Point, Newport River, Carteret County,
North Carolina]
Leptalpheus mexicanus Ros & Carvacho, 1983: 306; Figs 1-3. [estuary of Muleg River, Baja California
Sur, Mexico, 2654'N 11159'W]
Leptalpheus pacifcus Banner & Banner, 1975a: 424; Fig. 1. [reef in Kaneohe Bay, Oahu, 2127'00''N
15747'16''W, low intertidal]
Leptalpheus pierrenoeli Anker, 2008: 783; Figs 1-2, 3A-B. [Panama, Caribbean coast, Isla Grande, south-
ern shore, village, near Cabaas Super Jackson, from burrow, 0.5-1.0 m]
Fig. 46. Leslibetaeus coibita Anker, Poddoubtchenko & Wehrtmann, 2006. Photo by Arthur Anker.
Leptathanas De Grave & Anker, 2008
= Leptathanas De Grave & Anker, 2008 (type species Leptathanas powelli De Grave & Anker, 2008,
Leptathanas powelli De Grave & Anker, 2008: 45; Figs 1-5. [Nigeria, Niger Delta, Eagle Island, Bonny
River system, from burrows of Upogebia furcata]
Leslibetaeus Anker, Poddoubtchenko & Wehrtmann, 2006
= Leslibetaeus Anker, Poddoubtchenko & Wehrtmann, 2006 (type species Leslibetaeus coibita An-
ker, Poddoubtchenko & Wehrtmann, 2006, by original designation and monotypy, gender mas-
culine)
Leslibetaeus caribbeaus Anker, 2011b: 64; Figs 1-2. [Trinidad & Tobago, Tobago Island, Sandy Bay, 1 m]
Leslibetaeus coibita Anker, Poddoubtchenko & Wehrtmann, 2006: 30; Figs 1-5. [Isla Coibita, on side fac-
ing Isla Coiba, north side of STRI beach house, frst small island connected by causeway, 738'80''N
8142'159''W, Isla Coiba National Park, Pacifc Coast of Panama] (Fig. 46)
Metabetaeus Borradaile, 1899
= Metabetaeus Borradaile, 1899 (type species Betaeus minutus Whitelegge, 1897, by monotypy, gen-
der masculine)
Metabetaeus lohena Banner & Banner, 1960b: 299; Fig. 1. [Lohena pool, a few hundred yards from the
shore between the deserted villages of Wai-o-ahu-kini and Kaili-kI, immediately west of Lae
(South Point), Hawaii]
Metabetaeus mcphersonae Anker, 2010c: 40; Figs 1-7. [French Polynesia, Society Islands, Moorea, in front
of Hilton hotel, 149.84247 W 17.47564 S, 13-17 m]
Metabetaeus minutus (Whitelegge, 1897)
= Betaeus minutus Whitelegge, 1897: 147; Plate 7, fgs 4-4b. [Funafuti, under stones and in sponges
in the mangrove swamp]
Metalpheus Coutire, 1908
= Metalpheus Coutire, 1908 (type species Alpheus rostratipes Pocock, 1890, designated by Shelford,
Metalpheus hawaiiensis (Edmondson, 1925)
= Crangon hawaiiensis Edmondson, 1925: 14; Fig. 2. [Lisiansky Island, shallow water]
Metalpheus paragracilis (Coutire, 1897d)
= Alpheus paragracilis Coutire, 1897d: 304. [Ile Tague]
? = Alpheus aglaopheniae Borradaile, 1900: 417; Plate 38, fgs 13a-f. [Engineer Group, British New
Guinea]
Metalpheus rostratipes (Pocock, 1890)
= Alpheus rostratipes Pocock, 1890: 522. [Fernando de Noronha]
= Alpheus sp.?, Metalpheus n. gen. Coutire, 1908: 213.
= Crangon hawaiiensis clippertoni Schmit, 1939: 11. [Clipperton Island, on rocks to south of landing
place, on north-eastern shore of atoll, shore collecting]
= Crangon nanus Banner, 1953: 90; Figs 30-31; nec Kryer, 1843. [Hanauma Bay, Oahu, 10f]
= Alpheus huikau Banner, 1959: 139; Fig. 5. [nomen novum for Crangon nanus Banner, 1953]
Mohocaris Holthuis, 1973c
= Mohocaris Holthuis, 1973c (type species Mohocaris bayeri Holthuis, 1973c, by original designation
Mohocaris bayeri Holthuis, 1973c: 490; Figs 1-2. [Of French Guiana, 658'N 5310'W to 701'N 5315'W,
127-131 m]
Nennalpheus Banner & Banner, 1981b
= Nennalpheus Banner & Banner, 1981b (type species Alpheopsis Sibogae De Man, 1910a, by original
Nennalpheus inarticulatus Banner & Banner, 1981b: 221; Fig. 1. [1359.2'N 12020.3'E, 208-222 m]
Nennalpheus sibogae (De Man, 1910a)
= Alpheopsis Sibogae De Man, 1910a: 307. [823.5'S 1194.6'E, Sapeh-strait, 70 m]
Notalpheus Mndez G. & Wicksten, 1982
= Notalpheus Mndez G. & Wicksten, 1982 (type species Notalpheus imarpe Mndez G. & Wicksten,
Notalpheus imarpe Mndez G. & Wicksten, 1982: 709; Figs 1-2. [of Paita, Peru, 503'S 8120'W, 143 m]
Orygmalpheus De Grave & Anker, 2000
= Orygmalpheus De Grave & Anker, 2000 (type species Orygmalpheus polites De Grave & Anker, 2000,
Orygmalpheus polites De Grave & Anker, 2000: 250; Figs 1-4. [Liang Island Lagoon, about 410'30"S
14452'47"E, Hansa Bay, Madang Province, Papua New Guinea, 10 m]
Parabetaeus Coutire, 1897b
= Parabetus Coutire, 1897b (type species Parabetus Culliereti Coutire, 1897b, by monotypy, gen-
der masculine)
= Neoalpheopsis Banner, 1953 (type species Neoalpheopsis hiati Banner, 1953, by original designation,
gender feminine)
Parabetaeus culliereti Coutire, 1897b
= Parabetaeus Culliereti Coutire, 1897b: 383. [Papeete]
Parabetaeus euryone (De Man, 1910a)
= Alpheopsis? Euryone De Man, 1910a: 308. [anchorage of Kawio- and Kamboling-islands, Karkaro-
long-group, reef]
= Neoalpheopsis hiati Banner, 1953: 21; Fig. 6. [Hanauma Bay, Oahu, 20f]
Parabetaeus hummelincki (Schmit, 1936)
= Alpheopsis hummelincki Schmit, 1936: 364; Plate 11, fgs 1a-h. [Kralendik (de Hoop) Island, Bo-
naire, about 3f]
Pomagnathus Chace, 1937b
= Pomagnathus Chace, 1937b (type species Pomagnathus corallinus Chace, 1937b, by original designa-
Pomagnathus corallinus Chace, 1937b: 124; Fig. 5. [of Arena Bank, 2.5 fms in coral]
Potamalpheops Powell, 1979
= Potamalpheops Powell, 1979 (type species Potamalpheops pylorus Powell, 1979, by original designa-
Potamalpheops amnicus Yeo & Ng, 1997: 165; Figs 1-2. [151'46.5''N 10400'31.0''E, Sungei Selangi, Kota
Tinggi, Johore]
Potamalpheops darwiniensis Bruce, 1993c: 698; Figs 1-3. [Hudsons Creek, Darwin Harbour, Northern Ter-
ritory, Australia, 1228.75'S 13055.67'E, intertidal]
Potamalpheops galle Anker, 2005: 32; Fig. 1. [SW Sri Lanka, Lelkada near Dodangoda, Galle District, Gib
River basin, 6 km from the sea, altitude 5 m, 0608'N 8010'E, 0.5-1 m]
Potamalpheops hanleyi Bruce, 1991d: 629; Figs 1-4. [Creek H, East Arm, Darwin Harbor, Northern Terri-
tory, 1232.6'S 13056.6'E, low water spring tide]
Potamalpheops haugi (Coutire, 1906)
= Alpheopsis Haugi Coutire, 1906: 377, 378; Figs 1-2. [Ngm, petit lac deau douce des bords de
lOgou, Gabon]
Potamalpheops johnsoni Anker, 2003a: 290; Figs 5-7, 20c-d. [Sungei Buloh mangrove forest, Singapore]
Potamalpheops miyai Yeo & Ng, 1997: 175; Figs 3-4. [Sungei Lagoi, bridge leading to Kampung Lagoi,
downstream of reservoir outlet, Pulau Bintan, Indonesia]
Potamalpheops monodi (Sollaud, 1932)
= Alpheopsis monodi Sollaud, 1932: 377; Figs 1-2. [Cameroun]
Potamalpheops palawanensis Cai & Anker, 2004: 259; Figs 15-16. [St Paul Cave subterranean river, near
cave entrance, Palawan]
Potamalpheops pininsulae Bruce & Ilife, 1992: 231; Figs 1-53. [Grote de la Troisime (Paradise Cave), Ile
de Pins, New Caledonia, 0-6 m]
Potamalpheops pylorus Powell, 1979: 120; Figs 1, 3. [Nigeria, mangrove creek at southwest corner of Col-
lege of Science and Technology, Port Harcourt, approx. 447'15''N 658'45''E]
Potamalpheops stygicola (H.H.Jr. Hobbs, 1973a)
= Alpheopsis stygicola H.H.Jr. Hobbs, 1973a: 73; Figs 1-2. [Cueva del Nacimineto del Ro San Antonio,
10 km SSW Acatln, Oaxaca, Mxico]
Potamalpheops tigger Yeo & Ng, 1997: 182; Figs 5-6. [Sungi Buloh mangrove, Singapore]
Prionalpheus Banner & Banner, 1960c
= Prionalpheus Banner & Banner, 1960c (type species Prionalpheus triarticulatus Banner & Banner,
1960c, by original designation and monotypy, gender masculine)
Prionalpheus brachytomeus Banner & Banner, 1971: 267. [outer edge of fringing reef at Korolevu, Viti-
levu, Fii]
Prionalpheus fssipes Coutire, 1908
= Alpheopsis fssipes Coutire, 1908: 193. [Providence, 50-78 brasses]
Prionalpheus gomezi Martnez-Iglesias & Carvacho, 1991: 85; Figs 1-12. [rcif Juan Garca, Golfe de Bata-
ban, SW de Cuba, 10 m]
Prionalpheus mortoni Bruce, 1990f: 647; Fig. 18. [Tolo Channel, 5 m; full description as Prionalpheus sp. af.
triarticulatus Bruce, 1990f: 628; Figs 10-12]
Prionalpheus nayaritae Alvarez, Camacho & Villalobos, 1996: 719; Fig. 4. [Punta de Mita, Nayarit, 2046'N
10531'W]
Prionalpheus sulu Banner & Banner, 1971: 268; Fig. 2. [eastern end of Big Santa Cruz Island, Zamboanga,
Mindanao, 10f]
Prionalpheus triarticulatus Banner & Banner, 1960c: 293; Fig. 1. [Korolevu, Vitilevu, Fii, reef fat on fring-
ing reef]
Pseudalpheopsis Anker, 2007b
= Pseudalpheopsis Anker, 2007b (type species Pseudalpheopsis guana Anker, 2007b, by original desig-
Pseudalpheopsis guana Anker, 2007b: 430; Figs 1-8. [Caribbean Sea, British Virgin Islands, Guana Island,
North Bay, North Beach, 9 m]
Pseudathanas Bruce, 1983d
= Pseudathanas Bruce, 1983d (type species Pseudathanas darwiniensis Bruce, 1983d, by original desig-
Pseudathanas darwiniensis Bruce, 1983d: 464; Figs 1-5. [Dudley Point Reef, East Point, Darwin, Australia,
1225.0'S 3049.1'E, LWS]
Pterocaris Heller, 1862b
= Pterocaris Heller, 1862b (type species Pterocaris typica Heller, 1862b, by monotypy, gender femi-
nine)
Pterocaris typica Heller, 1862b: 398; Plate 1, fgs 7-18. [Amboina]
Racilius Paulson, 1875
= Racilius Paulson, 1875 (type species R.[acilius] compressus Paulson, 1875, by monotypy, gender
masculine)
Racilius compressus Paulson, 1875
= R.[acilius] compressus Paulson, 1875: 107; Plate 14, fg. 2. [Red Sea]
Richalpheus Anker & Jeng, 2006
= Richalpheus Anker & Jeng, 2006 (type species Richalpheus palmeri Anker & Jeng, 2006, by original
Richalpheus dahabensis Anker & Dworschak, 2007b: 2333; Figs 1-3. [Egypt, Red Sea, Dahab, Laguna, 13 m,
from mound of Glypturus sp.]
Richalpheus palmeri Anker & Jeng, 2006: 381; Figs 1-6. [Philippines, Panglao Island, Pontod Islet lagoon,
933.1'N 12344.0'E, 3-4 m]
Rugathanas Anker & Jeng, 2007
= Rugathanas Anker & Jeng, 2007 (type species Athanas verrucosus Banner & Banner, 1960a, by origi-
Rugathanas borradailei (Coutire, 1903)
= Arete Borradailei Coutire, 1903: 80; Figs 19-24. [Hulule Male Atoll]
= Arete Borradailei Coutire, 1905b: 861; Fig. 133. [Hulule Male Atoll]
= Arete ghardaqensis Ramadan, 1936: 15; Plate 1, fgs 2-8. [Ghardaga, Red Sea]
= Athanas polynesia Banner & Banner, 1966b: 152; Fig. 7. [Alofau, Tutuila, 3-10f]
Rugathanas verrucosus (Banner & Banner, 1960a)
= Athanas verrucosus Banner & Banner, 1960a: 147; Fig. 4. [ocean (windward) reef, Parry Island, Eni-
wetok]
Salmoneus Holthuis, 1955b
= Jousseaumea Coutire, 1897b (type species Jousseaumea serratidigitus Coutire, 1897b, designated by
Holthuis, 1955b, gender feminine; invalid junior homonym of Jousseaumia Sacco, 1894 (Mollusca);
name placed on the Ofcial Index of Rejected and Invalid Names in Zoology in Opinion 673 in
1963)
= Salmoneus Holthuis, 1955b (nomen novum for Jousseaumea Coutire, 1897b, gender masculine;
Salmoneus alpheophilus Anker & Marin, 2006: 310; Figs 12-13. [South China Sea, Vietnam, Nhatrang Bay,
Tre Island, Dam Bay, 1210'20''N 10915'30''E, low tide, in burrow of goby-associated Alpheus sp.]
Salmoneus armatus Anker, 2010d: Figs 3-4, 14a. [Panama, Caribbean coast, Isla Grande, southern shore,
100 m east of western-most point, under rock on muddy sand, 0.5 m depth]
Salmoneus arubae (Schmit, 1936)
= Jousseaumea arubae Schmit, 1936: 366; Plate 12, Figs 2a-g. [Aruba, Punta Braboe (Oranjestad),
Schorrenfche, about 1 f]
Salmoneus auroculatus Anker & Marin, 2006: 300; Figs 4-6, 7a. [South China Sea, Vietnam, Nhatrang Bay,
Tre Island, 6-8 m]
Salmoneus babai Miyake & Miya, 1966: 133; Fig. 1. [Suno coral reef, Kasari, Amami-oshima]
Salmoneus brevirostris (Edmondson, 1930)
= Jousseaumea brevirostris Edmondson, 1930: 7; Fig. 3. [Waikiki reef, Oahu, in shallow water]
Salmoneus brucei Komai, 2009: 870; Figs 1-5. [sand fat in Uehara Port, Iriomote Island, Yaeyama Islands,
southern Rykyu Islands, intertidal burrow of Lepidophthalmus tridentatus]
Salmoneus bruni Banner & Banner, 1966a: 42; Fig. 11. [Gulf of Thailand, 28 m]
Salmoneus caboverdensis Dworschak, Anker & Abed-Navandi, 2000: 309; Figs 27-41. [Cape Verde Islands,
Ilha do So Tiago, NE-coast, tidal fat at Moia-Moia, with Neocallichirus pachydactylus]
Salmoneus camaroncito Anker, 2010d: 184; Figs 5-6, 14b. [Panama, Caribbean coast, Isla Grande, western
point, under rock on coarse sand mixed with rubble, 0.5 m depth]
Salmoneus carvachoi Anker, 2007c: 27; Fig. 3. [Guadeloupe, estuary of Rivire Lzarde, dredge between
estuary and canal]
Salmoneus cavicolus Felder & Manning, 1986: 503; Figs 4-6. [Florida, St. Lucie County, Indian River la-
goon, south side of Fort Pierce Inlet, 2727.7'N 8018.7'W]
Salmoneus colinorum De Grave, 2004b: 41; Figs 1-4. [Ngerbeched mangrove, SW of Malakal Harbour,
Palau, 0719.703'N 13428.611'E]
Salmoneus cristatus (Coutire, 1897c)
= Jousseaumea cristata Coutire, 1897c: 234. [Djibouti]
Salmoneus degravei Anker, 2010d: 188; Figs 7-9, 14c. [Panama, Caribbean coast, Isla Grande, southern
shore, about 100 m east of western point and Hotel Isla Grande, from burrow of unknown host,
0.5-1 m depth] (Fig. 47)
Salmoneus erasimorum Dworschak, Anker & Abed-Navandi, 2000: 313; Figs 42-56. [Adria, Kvarner,
Opatja, Hotel Beach, ca. 1.5 m, with Callianassa tyrrhena]
Salmoneus falcidactylus Anker & Marin, 2006: 304; Figs 8-9. [South China Sea, Vietnam, Nhatrang Bay,
about 2 km SW of Mung Island, 40 m]
Salmoneus gracilipes Miya, 1972: 38; Plate 3. [Amakusa Islands, Shino-shima Island, Tomioka, low tide mark]
Salmoneus hilarulus (De Man, 1910a)
= Jousseaumea hilarula De Man, 1910a: 304. [142.5'S 13047.5'E, 32 m]
Salmoneus hispaniolensis Anker, 2010d: 178; Figs 1-2. [Dominican Republic, Boca Chica, sandy lagoon
with artifcial barrier in front of hotel, 1 m]
Salmoneus jarli (Holthuis, 1951a)
= Jousseaumea jarli Holthuis, 1951a: 94; Fig. 20. [of Nigeria, 412'N 705'E, 19 m]
Salmoneus kekovae Grippa, 2004: 45; Figs 1-2. [Kekova, southwest coast of Turkey]
Salmoneus komaii Anker, 2011c: 73; Figs 5-7. [Mariana Islands, Guam, west coast of Guam, south of Orote
Point, Barracuda Rock, less than 10 m]
Salmoneus latirostris (Coutire, 1897b)
= Jousseaumea latirostris Coutire, 1897b: 382. [Mer Rouge]
Salmoneus mauiensis (Edmondson, 1930)
= Jousseaumea mauiensis Edmondson, 1930: 5; Fig. 2. [island of Maui, shallow water]
Salmoneus nhatrangensis Anker & Marin, 2006: 296; Figs 1-3. [South China Sea, Nhatrang Bay, Tre Island,
8-10 m]
Salmoneus ortmanni (Rankin, 1898)
= Athanas ortmanni Rankin, 1898: 251; Plate 30, fg. 7. [near Nassau, New Providence, along shore]
Salmoneus paulayi Anker, 2011c: 68; Figs 1-4. [Mariana Islands, Guam, west coast of Guam, north of Apra
Harbor, between Piti Channel and Cabras, reef fat, 0.5-1.5 m]
Salmoneus poupini Anker, 2011c: 78; Figs 8-11. [French Polynesia, Society Islands, Moorea, Moorea la-
goon, public beach between Temae point and Softel, 0.5-2 m]
Salmoneus pusillus Anker & Marin, 2006: 307; Figs 10-11. [South China Sea, Vietnam, Nhatrang Bay,
Pyramides, 14-16 m]
Fig. 47. Salmoneus degravei Anker, 2010. Photo by Arthur Anker.
Salmoneus rocas Anker, 2007c: 29; Fig. 4. [Brazil, Atol das rocas, east of Laguna Interna, in calcareous al-
gae, 1 m]
Salmoneus rostratus Barnard, 1962: 240; Fig. 1. [Nosi B, U. W.; lectotype designation by De Grave &
Wilkins, 1997]
Salmoneus serratidigitus (Coutire, 1897b)
= Jousseaumea serratidigitus Coutire, 1897b: 382. [Mer Rouge]
Salmoneus seticheles Anker, 2003b: 102; Figs 1-5. [mouth of Ludmilla Creek, Darwin, 1224.8'S 13051.7'E,
LWS]
Salmoneus setosus Manning & Chace, 1990: 17; Fig. 9. [South West Bay, Ascension Island]
Salmoneus sibogae De Man, 1910a
= Jousseaumea Sibogae De Man, 1910a: 303. [5700 M. N. 279E from south point of South-Lucipara-
island, reef-exploration]
Salmoneus singaporensis Anker, 2003a: 284; Figs 1-2, 3a-b. [Tanjong Penuru, Jurong, Singapore, mudfats]
Salmoneus sketi Fransen, 1991a: 171; Figs 1-29. [underwater cave, of the N coast of the small island Laer-
naka, Kornati Archipelago, Middle Dalmatia, Adriatic]
Salmoneus tafaongae Banner & Banner, 1966b: 155. [3 miles east of Apia, Upolu, Western Samoa, 1f below
low water]
Salmoneus teres Manning & Chace, 1990: 20; Fig. 10. [South West Bay, Ascension Island]
Salmoneus tricristatus Banner, 1959
= Salmoneus tricristata Banner, 1959: 131; Fig. 1. [Tomil Harbor, Yap island, Caroline Archipelago]
Salmoneus wehrtmanni Anker, 2010d: 193; Figs 10-11, 12a, 13b, 14d. [Panama, Caribbean coast, Isla
Grande, southern shore, under rocks and rubble on sand, near seagrass, 0.5-1 m depth]
Stenalpheops Miya, 1997
= Stenalpheops Miya, 1997 (type species Stenalpheops anacanthus Miya, 1997, by original designation
= Cavipelta Hayashi, 1998 (type species Cavipelta yamashitai Hayashi, 1998 (junior subjective syno-
nym of Stenalpheops anacanthus Miya, 1997), by original designation and monotypy, gender femi-
nine)
= Chelomalpheus W. Kim, 1998 (type species Chelomalpheus koreanus W. Kim, 1998 (junior subjective
synonym of of Stenalpheops anacanthus Miya, 1997), by original designation and monotypy, gender
masculine)
Stenalpheops anacanthus Miya, 1997: 149; Figs 1A, 2-5. [Seto Inland Sea, Hiroshima Prefecture, Miyajima
Island, intertidal zone, found in a burrow of Upogebia (Upogebia) major]
= Chelomalpheus koreanus W. Kim, 1998: 141; Figs 1-3. [Daecheon, the Yellow Sea, Korea, 3619'05''N
12630'25''E, from pool dug in mud-sand beach]
= Cavipelta yamashitai Hayashi, 1998: 231; Figs 1-4. [Seto Inland Sea, tidal fat of Miyajima, Horishi-
ma Prefecture, living in tube of Upogebia major]
Stenalpheops crangonus (Anker, Jeng & Chan, 2001)
= Chelomalpheus crangonus Anker, Jeng & Chan, 2001: 1053; Figs 4-8. [Chung-Hua County, W Tai-
wan, mudfats, collected together with Upogebia edulis]
Synalpheus Spence Bate, 1888
= Homaralphus Spence Bate, 1876 (nomen nudum)
= Homaralpheus Spence Bate, 1888 (type species A.[lpheus] minus Say, 1818, designated by Holthuis,
1955b, gender masculine)
= Synalpheus Spence Bate, 1888 (type species Synalpheus falcatus Spence Bate, 1888 (junior subjective
synonym of Alpheus Comatularum Haswell, 1882), by monotypy, gender masculine)
= Alpheinus Borradaile, 1900 (type species Alpheinus tridens Borradaile, 1900 (junior subjective syno-
nym of Alpheus Stimpsonii De Man, 1888b), gender masculine)
= Zuzalpheus Ros & Dufy, 2007 (type species Zuzalpheus kensleyi Ros & Dufy, 2007, by original
Synalpheus albatrossi Coutire, 1909: 89; Fig. 54. [Laysan Island, 10-19 fms]
Synalpheus agelas Pequegnat & Heard, 1979: 110; Figs 1-4. [2752'N 9348'W, West Flower Garden Bank,
Gulf of Mexico, 25 m]
Synalpheus amabilis De Man, 1910a: 295. [Banda, 9-36 m]
Synalpheus anasimus Chace, 1972: 82; Figs 25-28. [Baha de lAscensin, Punta Solimn to 300 m south-
west, 5f]
Synalpheus anceps Banner, 1956: 334; Fig. 8. [Saipan]
Synalpheus ancistrorhynchus De Man, 1909a: 124. [Anchorage of Pulu Jedan, east cost of Aru-islands
(pearl-banks), 13 m]
Synalpheus androsi Coutire, 1909: 82; Fig. 50. [Andros Island, Bahamas]
Synalpheus antenor De Man, 1910a
= Synalpheus Antenor De Man, 1910a: 293. [07'.2N 13025'.5E, 83-59 m; 142'.5S 13047'.5E, 32 m;
Banda-anchorage, 9-36 m]
Synalpheus apioceros apioceros Coutire, 1909
= Synalpheus apioceros Coutire, 1909: 27; Fig. 9. [Marco, Florida]
Synalpheus apioceros desterroensis Coutire, 1909: 31; Fig. 13. [Desterro]
Synalpheus apioceros leiopes Coutire, 1909: 30; Fig. 12. [Venezuela]
Synalpheus apioceros mayaguensis Coutire, 1909: 30; Fig. 11. [Mayaguez, Porto Rico, on coral reef]
Synalpheus arostris Wicksten, 1989b: 78; Fig. 1. [Morro de Los Agujeros, Baha de Malaga, Colombia,
355'N 7720'W, rocky intertidal zone]
Synalpheus bannerorum Abele, 1975b: 79; Fig. 32. [southeastern side of Malpelo Island, Colombia, 10 m]
Synalpheus barahonensis Armstrong, 1949: 20; Fig. 7. [behind Piedra Prieta Reef, Dominican Republic,
from Agaricia agaricites]
Synalpheus belizensis Anker & Tth, 2008: 19; Figs 12-13, 14I. [Belize, Carrie Bow Cay, approx. 12 m, in
cryptic sponge]
Synalpheus bispinosus De Man, 1910a: 302. [Bay of Pidjot, Lombok, 9-22 m; Banda-anchorage, 9-36 m]
Synalpheus bituberculatus De Man, 1910a: 294. [Haingsisi, Samau Island, Timor, 36 m; Lumu-lumu-shoal,
Borneo-bank, reef, 142'.5S 13047'.5E, 32 m; Anchorage of Pulu Jedan, east coast of Aru-islands
(Pearl-banks), 13 m; 825'.2S 12718'.4E, anchorage between Nusa Besi and the N.E. point of Timor,
27-54 m; Anchorage East of Sailus Besar, Paternoster islands, up to 36 m]
Synalpheus biunguiculatus (Stimpson, 1860a)
= Alpheus biunguiculatus Stimpson, 1860a: 31. [ad insulas Hawaiensis, inter madreporas]
Synalpheus bocas Anker & Tth, 2008: 17; Figs 10-11, 14E-H. [Panama, Caribbean coast, Bocas del Toro,
STRI Bay, in Xestospongia rosariensis, 5-15 m]
Synalpheus bousfeldi Chace, 1972: 86; Figs 29-30. [Virgin Gorda, Colquhoun Reef at entrance to North
Sound]
Synalpheus bradleyi Verrill, 1922: 108; Plate 33, fg. 1. [Pearl Islands, Bay of Panama]
Synalpheus brevicarpus (Herrick, 1891)
= Alpheus saulcyi var. brevicarpus Herrick, 1891: 384 (?partim); Plate 4, fgs 1-3; Plate. 21, fgs 1-4, 8, 9;
Plate 22, fgs 1, 2, 4-10, 12-16; Plate 23, fgs 1-8; Plate 24, fgs 1, 3. [Bahamas, assumed to be Nassau,
New Providence]
= Synalpheus brevicarpus guerini Coutire, 1909: 52; Fig. 30. [Key West]
Synalpheus brevidactylus Anker & Tth, 2008: 14; Figs 8-9, 14C-D. [Panama, Caribbean coast, Isla Grande,
between village and west point, in cryptic sponge, 1-3 m]
Synalpheus brevifrons Chace, 1972: 89; Figs 31-32. [Dominica, north end of Woodbridge Bay]
Synalpheus brevispinis Coutire, 1909
= Synalpheus townsendi brevispinis Coutire, 1909: 34; Fig. 16. [Lower California]
Synalpheus brooksi Coutire, 1909: 69; Fig. 41. [Sugar Loaf Key, Florida]
= Synalpheus brooksi strepsiceros Coutire, 1909: 73; Fig. 42. [St. Thomas]
= Synalpheus brooksi eleuther Coutire, 1909: 73; Fig. 43. [The Current, Eleuthera Island, Bahamas]
Synalpheus carinatus (De Man, 1888b)
= Alpheus carinatus De Man, 1888b: 508; Plate 22, fg. 2-2c. [Amboina]
= Synalpheus carinatus var. binongcensis De Man, 1909a: 111. [Anchorage of Pasir Pandjang, west
coast of Binongka, 278 m]
= Synalpheus carinatus var. ubianensis De Man, 1909a: 111. [67'5.N 12026'E, anchorage of North-
Ubian, 16-23 m; Anchorage of Pasir Pandjang, west coast of Binongka, reef]
Synalpheus carpenteri MacDonald & Dufy, 2006: 2; Figs 1-7. [Long Reef, Belize, from canals of Agelas
clathrodes]
Synalpheus chacei Dufy, 1998: 389; Figs 1-5. [Carrie Bow Cay, Belize, from Agelas dispar, 3 m]
Synalpheus charon (Heller, 1861)
= A.[lpheus] charon Heller, 1861: 25. [im rothen Meere; fully described in Heller, 1862c]
= Alpheus prolifcus Spence Bate, 1888: 556; Plate 99, fg. 4. [of Honolulu, Sandwich Islands, 18 fms]
= Synalpheus Helleri De Man, 1911d: 246. [Nicobar Islands]
= Synalpheus charon obscurus Banner, 1956: 329; Fig. 5. [reef fat on the SE side of Unai Obyan, Saipan]
Synalpheus comatularum (Haswell, 1882)
= Alpheus Comatularum Haswell, 1882: 762. [Albany Passage, near Cape York, Queensland, few fms]
= Synalpheus falcatus Spence Bate, 1888: 574; Plate 103. [Challenger stn 186, 1030'S 14218'E, between
Cape York and the Arrou Islands, 8 fms]
Synalpheus corallinus MacDonald, Hultgren & Dufy, 2009: 16; Figs 5-9; Plates 2D, 3A. [Jamaica, Dairy
Bull Reef, 1828.083'N 7723.289'W, from canals of Hyatella intestinalis]
Synalpheus coutierei Banner, 1953: 36. [nomen novum for Synalpheus biunguiculatus sensu Coutire, 1898e
nec Stimpson, 1860a]
? = Synalpheus biunguiculatus var. exilipes Coutire, 1905b: 874; Plate 71, fgs 10-10a. [Haddumati
Atoll; North Male Atoll; South Nilandu Atoll]
Synalpheus cretoculatus Banner & Banner, 1979: 30; Fig. 2. [1037'N 9734'E, Andaman Sea, of southern
tip of Burma, 75-80 m]
Synalpheus crosnieri Banner & Banner, 1983: 92; Fig. 11. [northwest coast of Madagascar, 1244'S 4825.2'E,
73 m]
Synalpheus curacaoensis Schmit, 1924a
= Synalpheus curaaoensis Schmit, 1924a: 66; Fig. 3. [Spanish Bay, between stones in the surf]
Synalpheus dardeaui (Ros & Dufy, 2007)
= Zuzalpheus dardeaui Ros & Dufy, 2007: 18; Figs 3-6; Plate 2. [Twin Cays, Belize, from Lissodendoryx
colombiensis, 1 m]
Synalpheus demani Borradaile, 1900: 416. [Lifu, Loyalty Islands]
= Alpheus triunguiculatus De Man, 1888b: 504; Plate 22, fg. 1-1e; nec Paulson, 1875. [Amboina]
= Synalpheus Brockii Nobili, 1901c: 2. [Amboina]
Synalpheus digueti Coutire, 1909: 48; Fig. 28a-a, c-c, k-m. [Lower California]
? = Alpheus lviusculus Lockington, 1878b: 474. [Port Escondido, Mulege Bay and other points on the
Californian shore of the Gulf of California]
= Synalpheus digueti ecuadorensis Coutire, 1909: 50; Fig. 28aa, cc. [St.Helena, Ecuador]
Synalpheus disparodigitus Armstrong, 1949: 17; Fig. 6. [quiet water behind Piedra Pietra Reef, Dominican
Republic]
Synalpheus dominicensis Armstrong, 1949: 23; Fig. 8. [quit water inside Piedra Prieta Reef]
Synalpheus dorae Bruce, 1988g: 843; Figs 1-6. [1905.0'S 11857.8'E, 82 m]
Synalpheus dufyi Anker & Tth, 2008: 8; Figs 4-5, 14A, 15A-B. [Panama, Caribbean coast, Isla Grande,
southern shore facing La Guaira, between village and west point, in cryptic sponge, 1-1.5 m]
Synalpheus echinus Banner & Banner, 1975c: 374; Fig. 27. [Trigg Island, near Perth, Western Australia]
Synalpheus elizabethae (Ros & Dufy, 2007)
= Zuzalpheus elizabethae Ros & Dufy, 2007: 23; Figs 7-10; Plate 2. [The Pinnacles (Sand Bores), SW of
Carrie Bow Cay, Belize, in Hymeniacidion caerulae, 2 m]
Synalpheus flidigitus Armstrong, 1949: 15; Fig. 5. [halfway between the Piedra Prieta Reef of Barahona
Harbour and the shore, Dominican Republic]
Synalpheus fossor (Paulson, 1875)
= Alph.[eus] fossor Paulson, 1875: 103; Plate 13, fgs 5-5g. [Red Sea]
= Synalpheus trionyx Coutire, 1908: 196. [Saya de Malha, 26 + 29 fms]
= Synalpheus Bakeri Coutire, 1908: 199. [South-Adelade]
= Synalpheus fossor var. propinqua De Man, 1909a: 121. [South-east side of Pearl-bank, Sulu-archipel-
ago, 15 m; 142'.5S 13047'.5E, 32 m; Anchorage of Pulu Jedan, East coast of Aru-islands (Pearl-
banks), 13 m; 825'2.S 12718'.4E, anchorage between Nusa Besi and the N.E. point of Timor, 27-54
m; Anchorage East of Sailus Besar, Paternoster-islands, up to 36 m]
= Synalpheus Bakeri var. Stormi De Man, 1911d: 253. [Balikpapan, east coast of Borneo; Atjeh]
Synalpheus fritzmuelleri Coutire, 1909
= Synalpheus fritzmlleri Coutire, 1909: 35; Fig. 18. [Marco, Florida]
= Synalpheus fritzmlleri elongatus Coutire, 1909: 37; Fig. 19. [South Carolina, Mouth of Bull Creek]
= Synalpheus fritzmuelleri var. cariba Verrill, 1922: 97; Fig. 8; Plate 39, fgs 3a-c. [Dominica Island]
= Synalpheus fritzmuelleri var. carolinensis Verrill, 1922: 99; Plate 22, fg. 6; Plate 39, fgs 1-1d. [Fort
Macon, North Carolina]
Synalpheus gambarelloides (Nardo, 1847)
= Alpheus gambarelloides Nardo, 1847: 6. [trovato di rado nella valle Gambarelli vicino Chioggia]
= Cancer glaber var. Olivi, 1792: 51; Plate 3, fg. 5; nec Cancer glaber Olivi, 1792. [la Golfo e delle La-
gune di Venezia]
= Alphaeus spincicerus Hope, 1851: 43. [mari neapolitano]
= Alpheus laevimanus Heller, 1862b: 401; Plate 1, fgs 25-27. [Sicilien, sowie von Pirano aus dem adri-
atischen Meere]
Synalpheus goodei Coutire, 1909: 58; Fig. 33. [Bermuda]
Synalpheus gracilirostris De Man, 1910a: 291. [825.2'S 12718.4'E, anchorage between Nusa Besi and the
N.E. point of Timor, 27-54 m]
Synalpheus haddoni Coutire, 1900c
= Synalpheus laevimanus var. Haddoni Coutire, 1900c: 411, unnumbered fgures. [Saibai Channel,
Torres Strait]
Synalpheus harpagatrus Banner & Banner, 1975c: 311; Fig. 9. [5 miles west of North Beach, Perth, Western
Australia, 50f]
Synalpheus hastilicrassus Coutire, 1905b: 875; Plate 72, fgs 12-12d. [South Nilandu Atoll; North Male
Atoll; Felidu Atoll; Kendikolu Reef; Miladumadulu Atoll]
? = Synalpheus acanthitelsonis Coutire, 1905b: 875; Plate 72, fgs 13-13c. [Hulule Male Atoll; Fadifolu
Atoll; Addu Atoll; Mulaku Atoll; North male Atoll]
= Synalpheus hastilicrassus var.? De Man, 1911d (partim): 264. [mid-channel in Solor-Strait of Kam-
pong Menanga, 113 m; 830'S 1197'.5E, Sapeh Strait, 73 m]
= Synalpheus hastilicrassus var. acanthitelsoniformis De Man, 1920a: 108. [east side of Pajunga island,
Kwandang-bay, on the reef]
Synalpheus heardi Dardeau, 1984: 47; Figs 23-26. [Florida Middle ground, 2836'N 8416'W, 30-34 m]
Synalpheus hemphilli Coutire, 1909: 39; Fig. 20. [West coast of Florida, 21-28 fms] (Fig. 48)
= Synalpheus hemphilli longicornis Coutire, 1909: 39; Fig. 39. [West coast of Florida]
Synalpheus herdmaniae Lebour, 1938
= Synalpheus herdmani Lebour, 1938: 651; Plate 2, fgs 21-34. [Tuticorin, Gulf of Manaar, 5-12 fms]
Synalpheus heroni Coutire, 1909: 42; Fig. 24. [Djibouti, Hron]
Synalpheus herricki Coutire, 1909: 74; Fig. 44. [Anclote, Florida; lectotype selection by Dardeau, 1984]
= Synalpheus herricki angustipes Coutire, 1909: 76; Fig. 45. [Anclote Section, Florida, 12.5 fms]
= Synalpheus herricki dimidiatus Coutire, 1909: 77; Fig. 46. [Anclote Section, Florida, 12.5 fms]
= Synalpheus tanneri Coutire, 1909: 78; Fig. 47. [Gulf of Mexico, 2915'30''N 8529'30''W, 27 fms]
Synalpheus hilarulus De Man, 1910a: 290. [Wunoh-bay, N.W. coast of Waigeu-island, reef]
Synalpheus hoetjesi Hultgren, MacDonald & Dufy, 2010: 231; Figs 3-9; Plate 4B-D. [Caracas Baai,
Curaao, 1204'11.64''N 6851'43.59''W, from the canals of Hyatella intestinalis]
Synalpheus idios (Ros & Dufy, 2007)
= Zuzalpheus idios Ros & Dufy, 2007: 36; Figs 13-17; Plate 3. [Carrie Bow Cay, in Hymeniacidion am-
philecta, 6 m]
Synalpheus iphinoe De Man, 1909a
= Synalpheus Iphino De Man, 1909a: 116. [823'.5S 1194'.6E, Sapeh-strait, 70 m; Madura-bay and
other localities in the southern part of Molo-strait, 54-90 m; Banda-anchorage, 9-36 m; Rumah-ku-
da-bay, Roma-island, 36 m]
Synalpheus iocasta De Man, 1909a
= Synalpheus Iocasta De Man, 1909a: 119. [Makassar and surroundings, up to 32 m; 058'.5N
12242'.5E, west of Kwadang-bay-entrance, 72 m; Anchorage north of Salomakie (Damar) is-
land, 45 m; 142'.5S 13047'.5E, 32 m; 420'S 12258'E, between islands of Wowoni and Buton,
northern entrance of Buton-strait, 75-94 m; Banda-anchorage, 9-36 m; Anchorage of Pulu Jedan,
east coast of Aru-islands (Pearl-banks), 13 m; 528'.2S 13453'.9E, 57 m; 825'.2S 12718'.4E, an-
chorage between Nusa Besi and the N.E. point of Timor, 27-54 m; 839'.1 1274'.4E, anchorage
south coast of Timor, 34 m; Mid-channel in Solor-strait of Kampong Menanga, 113 m; 830'S
1197'.5E, 73 m]
Synalpheus irie MacDonald, Hultgren & Dufy, 2009: 25; Figs 11-16; Plate 3C-D. [fore-reef (near M1 chan-
nel marker), 1828.083'N 7723.289'W, from canals of Auleta cf. sycinularia]
Synalpheus jedanensis De Man, 1909a: 117. [Anchorage of Pulu Jedan, east coast of Aru-islands (Pearl-
banks), 13 m]
Synalpheus kensleyi (Ros & Dufy, 2007)
= Zuzalpheus kensleyi Ros & Dufy, 2007: 41; Figs 18-22; Plate 3. [The Pinnacles (Sand Bores), SW of
Carrie Bow Cay, Belize, 6 m]
Synalpheus kuadramanus Hultgren, MacDonald & Dufy, 2010: 240; Figs 10-13; Plate 5A. [Piscadera Baai
east, 1207'15.96''N 6858'11.64''W, in Xestospongia subtriangularis]
Synalpheus kusaiensis Kubo, 1940e: 87; Fig. 10. [Kusaie, Malem, Caroline Islands]
Synalpheus lani Hermoso & Alvarez, 2005: 522; Figs 1-2. [Punta de Mita, Nayarit, Mxico]
Synalpheus laticeps Coutire, 1905b: 874; Plate 72, fgs 11-11d. [North Male Atoll]
Synalpheus lockingtoni Coutire, 1909: 21; Fig. 1; nomen novum for Alpheus leviusculus Lockington, 1878b
nec Alpheus edwardsi var. leviusculus Dana, 1852b. [Port Escondido, Mulege Bay, and other points on
the Californian shore of the Gulf of California; Albatross stn 4421, eastern point of San Nicolas Is-
land, N. 26 W. 3.8 miles, 229-298 fms]
Fig. 48. Synalpheus hemphilli Coutire, 1909. Photo by Arthur Anker.
Synalpheus longicarpus (Herrick, 1891)
= Alpheus saulcyi var. longicarpus Herrick, 1891: 383; Plate 21, fgs 5-7; Plate 22, fgs 3, 11, ? 14, 17;
Plate 24, ? fgs 2, 4-9 (partim). [Bahamas, assumed to be Nassau, New Providence]
= Synalpheus longicarpus approxima Coutire, 1909: 57; Fig. 32. [Gulf of Mexico, 26 fms]
Synalpheus lophodactylus Coutire, 1908: 197. [Lagon de Diego]
Synalpheus macromanus Edmondson, 1925: 9; Fig. 1. [Lisiansky Island]
Synalpheus mcclendoni Coutire, 1910: 487; Fig. 3. [Dry Tortugas, Florida]
Synalpheus merospiniger Coutire, 1908: 195. [Amirante, 20-44 brasses]
Synalpheus mexicanus Coutire, 1909
= Synalpheus townsendi mexicanus Coutire, 1909: 34; Fig. 17. [southern part of Gulf of California, 9.5 fms]
Synalpheus microneptunus Hultgren, MacDonald & Dufy, 2011: 8; Figs 3-6. [Cement Factory, 1317'21.84''N
5939'27.72''W]
Synalpheus minus (Say, 1818)
= A.[lpheus minus] Say, 1818: 245. [coasts of the southern states, and of East Florida]
= Synalpheus morinus Coutire, 1898f: 189. [nomen nudum, lapsus for Synalpheus minus (Say, 1818)]
= Synalpheus minus bahiensis Coutire, 1909: 45; Fig. 26. [Brazil, Plataforma, Bahia]
= Synalpheus minus antillensis Coutire, 1909: 46; Fig. 27. [St. Thomas, 20-23 fms]
= Synalpheus minus var. somersi Verrill, 1922: 108; Plate 33, fgs 4-4a; Plate 34, fgs 1-1u; Plate 36, fgs
1-1e, 2. [Bermuda, on the coral reefs]
Synalpheus modestus De Man, 1909a: 115. [Banda, reef-exploration]
Synalpheus mortenseni Banner & Banner, 1985: 47; Fig. 5. [Kei Islands, Ambon, 13-18 m]
Synalpheus mulegensis Ros, 1992: 9; Fig. 6. [Baha Concepcin, 22 m]
Synalpheus mushaensis Coutire, 1908
= Synalpheus Mushaensis Coutire, 1908: 202. [Iles Musha]
Synalpheus neomeris (De Man, 1897)
= Alpheus neomeris De Man, 1897: 734; Plate 35, fg. 61. [Atjeh]
= Synalpheus Gravieri Coutire, 1905b: 870; Plate 70, fgs 2-2c. [North Male Atoll]
= Synalpheus Bougainvillei Coutire, 1905b: 871 [lapsus for Synalpheus Gravieri Coutire, 1905b]
= Synalpheus miscellaneus De Man, 1909a: 118. [825'.2S 12718'.4E, anchorage between Nusa Besi
and the N.E. point of Timor, 27-54 m]
Synalpheus neptunus neptunus (Dana, 1852a)
= Alpheus neptunus Dana, 1852a: 22. [Tictabon Island, near Zamboanga, Philippines, Sulu Sea, scat-
tered coral heads, 4-20 f; neotype selection by Banner & Banner, 1972a]
Synalpheus neptunus germanus Banner & Banner, 1975c: 321; Fig. 12. [Cotesloe, Western Australia]
Synalpheus nilandensis Coutire, 1905b
= Synalpheus Nilandensis Coutire, 1905b: 871; Plate 70, fgs 4-4d. [North Male Atoll; Suvadiva Atoll;
South Male Atoll; South Nilandu Atoll]
= Synalpheus Nilandensis var. oxyceros Coutire, 1905b: 871; Plate 70, Figs 5-5a. [South Nilandu Atoll]
= Synalpheus Nilandensis var. bandaensis De Man, 1909a: 121. [823'.5S 1194'6E, Sapeh-strait, 70 m;
Banda, 9-36 m]
Synalpheus nobilii Coutire, 1909: 40; Fig. 22. [St. Helena, Ecuador]
Synalpheus obtusifrons Chace, 1972: 99; Figs 37-38. [Baha de la Ascensin, just behind center of Arrecife
Nicchehabin, 1-5 f]
Synalpheus occidentalis Coutire, 1909
= Synalpheus goodei occidentalis Coutire, 1909: 60; Fig. 34. [Bermudas]
Synalpheus odontophorus De Man, 1909a: 113. [very near Siboga stn 65, 70'S 12034'.5E, 120-400 m; 536'.5S
13255'.2E, 2.3 miles N, 63W from the north point of Nuhu Jaan, Kei-islands, 90 m; 90'.3S
12624'.5E, 112 m]
Synalpheus orapilosus Hultgren, MacDonald & Dufy, 2010: 246; Figs 14-17, Plate 5D-E. [Caracas Baai,
Curaao, 1204'11.64''N 6851'43.56''W, in an unidentifed white web-like sponge embedded in Ma-
dracis sp. rubble]
Synalpheus osburni Schmit, 1933: 1; Fig. 1. [inside Cayo Maria Langa, near Guayanilla Harbor, Porto
Rico]
Synalpheus otiosus Coutire, 1908: 195. [Coevity]
Synalpheus pachymeris Coutire, 1905b
= Synalpheus biunguiculatus var. pachymeris Coutire, 1905b: 873; Plate 71, fgs 9-9a. [South Nilandu
Atoll]
= Synalpheus pachymeris var. Cargadosi Coutire, 1921: 417; Plate 61, fg. 10. [Cargados Carajos]
Synalpheus pandionis Coutire, 1909: 67; Fig. 39. [St. Thomas, 20-23 fms]
= Synalpheus pandionis extentus Coutire, 1909: 69; Fig. 40. [St. Thomas, 20-23 fms]
= Synalpheus grampusi Coutire, 1909: 62; Fig. 36. [Gulf of Mexico, 2633'N 8310'W, 28 fms]
Synalpheus paradoxus Banner & Banner, 1981a: 68; Fig. 10. [Negus Salomon, 1537'N 4043'E, 15 fms]
Synalpheus paralaticeps Banner & Banner, 1982a: 299; Fig. 92. [Rudder Reef, of Port Douglas, Queens-
land, outer reef fat, 200 yards from edge]
Synalpheus paraneomeris Coutire, 1905b: 872; Plate 71, fgs 7-7e. [Goidu Atoll, Minikoi]
= Synalpheus paraneomeris Coutire, 1899a: 456, 466, 486, 501. [nomen nudum]
= Synalpheus paraneomeris oxyceros Coutire, 1909: 9. [nomen nudum]
= Synalpheus paraneomeris prolatus Coutire, 1909: 9. [nomen nudum; see discussion in Banner, 1953]
= Synalpheus paraneomeris var. halmaherensis De Man, 1909a: 122. [Wunoh-bay, NW coast of Waigeu-
island]
= Synalpheus paraneomeris var. praedabunda De Man, 1909a: 123. [Anchorage of Labuan Pandan,
Lombok, 18 m; Sailus Keyjil, Paternoster-islands, 27 m; Anchorage of Kawio-and Kamboling-is-
land, Karkarolong-group, reef; Anchorage of Kilsuin, west-coast of Kur-island, reef-exploration]
= Synalpheus paraneomeris var. prolatus De Man, 1911d: 241. [Wunoh-bay, NW coast of Waigeu-is-
land, reef; Anchorage of the south point of Kabana-island, reef]
= Synalpheus Sluiteri De Man, 1920a: 107. [Bay of Batavia]
= Synalpheus paraneomeris var. praslini Coutire, 1921: 415; Plate 61, fg. 6. [Seychelles, Praslin Reef]
= Synalpheus paraneomeris var. seychellensis Coutire, 1921: 415; Plate 61, fg. 7. [Seychelles]
Synalpheus paraneptunus Coutire, 1909: 86; Fig. 52 (partim). [Panama, Caribbean coast, of Golfo de
Morrosquillo, 0930'N 7620'W, 77 m]
Synalpheus parfaiti (Coutire, 1898f)
= Synalpheus laevimanus var. Parfaiti Coutire, 1898f: 191; Figs 3-3a. [Annobon]
Synalpheus paulsoni paulsoni Nobili, 1906c
= Synalpheus Paulsoni Nobili, 1906c: 28; Fig. 1. [Banc au NE de lle Arzana]
Synalpheus paulsoni kurracheensis Coutire, 1908
= Synalpheus Paulsoni Kurracheensis Coutire, 1908: 203. [Kurrachee]
Synalpheus paulsoni liminaris Coutire, 1908
= Synalpheus Paulsoni liminaris Coutire, 1908: 201. [Djibouti; Golfe Persique]
Synalpheus paulsoni rameswarensis Coutire, 1908
= Synalpheus Paulsoni Rameswarensis Coutire, 1908: 201. [Rameswaran]
Synalpheus paulsonoides Coutire, 1909
= Synalpheus paulsonodes Coutire, 1909: 24; Fig. 6. [island of San Jos, Lower California]
Synalpheus pectiniger Coutire, 1907b: 611. [Albatross stn 2413 (26N 08257'30''W), golfe de Mexico,
banc de la Floride, 24 brasses; Curaao; full description in Coutire, 1909]
= Alpheus Praecox Herrick, 1888: 34. [nomen oblitum; see De Grave & Anker, 2010]
Synalpheus pescadorensis Coutire, 1905b
= Synalpheus Pescadorensis Coutire, 1905b: 877; Plate 73, fgs 15-15e. [Miladumadulu Atoll; South
Nilandu Atoll; North Male Atoll, Pescadores]
Synalpheus peruvianus Rathbun, 1910
= Synalpheus townsendi peruvianus Rathbun, 1910: 563; Plate 53, fg. 4. [oysterbeds of Matapalo, near
Capon]
Synalpheus plumosetosus MacDonald, Hultgren & Dufy, 2009: 36; Figs 17-21. [Jamaica, Dairy Bull Reef,
1828.083'N 7723.289'W, from canals of Auleta cf. sycinularia]
Synalpheus pococki Coutire, 1898c
= Synalpheus neomeris var. Pococki Coutire, 1898c: 167; Figs 2-2a. [Holothuria Bank, NW Australie;
Macclesfeld Bank, Arafura See]
Synalpheus quadriarticulatus Banner & Banner, 1975c: 297; Fig. 5. [between Hammond and Waivea Is-
lands, Torres Straits, 3 m]
Synalpheus quadrispinosus De Man, 1910a: 298. [823'.5S 1194'.6E, Sapeh-Strait, 70 m; 142'.5S 13047'.5E,
32 m; Anchorage of Pulu jedan, East coast of Aru-islands (Pearl-banks), 13 m; 830'S 1197'.5E,

73
m; Anchorage East of Sailus Besar, Paternoster-islands, up to 36 m]
= Synalpheus quadridens De Man, 1910a: 299. [825'.2S 12718'.4E, anchorage between Nusa besi and
the N.E. point of Timor, 27-54 m]
Synalpheus quinquedens Tatersall, 1921: 376; Plate 28, fgs 1-5. [Suez, among coral; Khor Dongonab,
among coral on reef; Mersa Ar-rakiya, 2015'N, among coral in 1f; Suakin Harbour, 198'N, washed
from sponges]
Synalpheus rathbunae Coutire, 1909: 84; Fig. 51. [St. Thomas, 20-30 fms]
Synalpheus readi Banner & Banner, 1972b: 137; Fig. 1. [Malakal Harbor, Urukthapel, Palau Island, 5 m]
Synalpheus recessus Abele & W. Kim, 1989: 15; Figs 7- 8. [Republic of Panama, Panama Canal, Mirafores
Locks]
Synalpheus redactocarpus Banner, 1953: 29; Fig. 8. [Keaoi Island, Halape, Kau Coast, Hawaii, intertidal]
Synalpheus regalis Dufy, 1996: 564; Figs 1-5. [outer reef ridge at Carrie Bow Cay, Belize, 1648'N 8805'W,
15 m, from Xestospongia cf. subtriangularis]
Synalpheus riosi Anker & Tth, 2008: 11; Figs 6-7. [Dominica, Prince Rupert Bay, 1.5 m]
Synalpheus ruetzleri MacDonald & Dufy, 2006: 8; Figs 8-13. [Sand Bores, Belize, from canals of Hymeni-
acidon cf. caerulea]
Synalpheus sanctithomae Coutire, 1909: 61; Fig. 35. [St. Thomas, 20-30 fms]
Synalpheus sanjosei Coutire, 1909
= Synalpheus apioceros sanjosei Coutire, 1909: 29; Fig. 10. [San Jos, Lower California]
Synalpheus sanlucasi Coutire, 1909: 41; Fig. 23. [Cape St. Lucas, Lower California]
Synalpheus scaphoceris Coutire, 1910
= Synalpheus townsendi scaphoceris Coutire, 1910: 486; Fig. 2. [Dry Tortugas, Florida]
Synalpheus sciro Banner & Banner, 1975c: 304; Fig. 7. [2740'S 11320'E, northwest of Bluf Point, Western
Australia, 7.5 fms]
Synalpheus senegambiensis Coutire, 1908
= Synalpheus Paulsoni Senegambiensis Coutire, 1908: 202. [Cap Vert]
Synalpheus septemspinosus De Man, 1910a: 297. [Anchorage east of Sailus Besar, Paternoster-islands, up
to 36 m]
Synalpheus sladeni Coutire, 1908
= Synalpheus Sladeni Coutire, 1908: 198. [Cargados Carajos, 30 brasses]
Synalpheus somalia Banner & Banner, 1979: 31; Fig. 3. [1111'N 5114'E, of Somalia, 47-49 m]
Synalpheus spinifrons (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840])
= A.[lpheus] spinifrons H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 355. [ctes de
Chili]
= Synalpheus latastei Coutire, 1909: 25; Fig. 7. [Chile; ?New Holland]
Synalpheus spiniger (Stimpson, 1860a)
= Alpheus spiniger Stimpson, 1860a: 31. [ad insulas Amakirrima prope Loo Choo]
Synalpheus spongicola Banner & Banner, 1981a: 78; Fig. 12. [Marsa Murach, from inside sponge]
Synalpheus stimpsonii (De Man, 1888b)
= Alpheus Stimpsonii De Man, 1888b: 513; Plate 22, fg. 3a-b. [Amboina]
= Alpheus amboin Zehntner, 1894: 202; Plate 8, fgs 23-23b. [Amboine]
= Alpheinus tridens Borradaile, 1900: 415; Plate 38, Figs 12a-g. [Sandal Bay, Lifu, Loyalty Islands]
= Alpheus stimpsoni Var.? Schenkel, 1902: 567; Plate 13, fgs 22a-c. [Kema]
= Synalpheus Stimpsoni var. Maldivensis Coutire, 1905b: 878; Plate 73, fgs 16-16d. [Hulule Male
Atoll]
= Synalpheus consobrinus De Man, 1909a: Moearas-reef, inner side, east coast of Borneo, up to 54 m;
Saleyer-anchorage and surroundings, reef-exploration; 825'.2S 12718'.4E, anchorage between
Nusa Besi and the NE point of Timor, 27-54 m; Mid-channel in Solor-strait of Kampong Menanga,
113 m]
= Synalpheus brucei Pots, 1915: 76; Figs 1-3; Plate 1, fg. 2. [Murray Island]
= Synalpheus striatus Kubo, 1938c: 89; Figs 1-2. [sima Island, Wakayam Prefecture]
Synalpheus streptodactylus Coutire, 1905b
= Synalpheus neomeris var. streptodactylus Coutire, 1905b: 870; Plate 70, fg. 1. [Haddumati Atoll;
Suvadiva Atoll; South Nilandu Atoll]
= Synalpheus streptodactyloides De Man, 1909a: 114. [825'.2S 12718'.4E, anchorage between Nusa
Besi and the NE point of Timor, 27-54 m]
= Synalpheus metaneomeris Coutire, 1921: 414; Plate 60, fg. 4. [Amirante Bank, 20-80 fms; Saya de
Malha, 47 and 90 fms; Cargados carajos, 28-33 fms; Providence, 58 fms]
= Synalpheus streptodactylus hadrungus Banner & Banner, 1966b: 158. [nomen novum for S. metaneom-
eris streptodactylus Coutire, 1921]
Synalpheus stylopleuron Hermoso Salazar & Hendrickx, 2006: 1109; Figs 3-4. [of Lobos Island, 2313'49"N
10627'43"W, Sinaloa, Mexico, in sponge Mycale parishii, 3-4 m]
Synalpheus superus Abele & W. Kim, 1989: 18; Figs 9-10. [Republic of Panama, Panama Canal, Mirafores
Locks]
Synalpheus tenuispina Coutire, 1909
= Synalpheus latastei tenuispina Coutire, 1909: 26; Fig. 8. [Desterro, Brazil]
Synalpheus thai Banner & Banner, 1966a: 61; Fig. 19. [of Koh Samet, Rayong, Thailand, 2 m]
Synalpheus thele MacDonald, Hultgren & Dufy, 2009: 43; Figs 22-27; Plate 5C-D. [Jamaica, Columbus
Park, Discovery Bay, 1827.955'N 7724.843'W, from canals of Agelas cf. clathrodes]
Synalpheus theano De Man, 1910a
= Synalpheus Theano De Man, 1910a: 296. [142.5'S 13047.5'E, 32 m]
Synalpheus tiou Banner & Banner, 1982a: 296; Fig. 91. [Tiou Reef, Lizard Island, with a crinoid]
Synalpheus townsendi Coutire, 1909: 32; Fig. 14. [Gulf of Mexico (2914'00''N 08529'15''W, 25 fms)]
= Synalpheus townsendi productus Coutire, 1909: 33; Fig. 15. [Gulf of Mexico (2846'00''N 08449'00''W,
26 fms)]
Synalpheus triacanthus De Man, 1910a: 301. [90.3'S 12624.5'E, 112 m, living in Solenocaulon]
Synalpheus tricuspidatus (Heller, 1861)
= A.[lpheus] tricuspidatus Heller, 1861: 24. [im rothen Meere]
= A.[lpheus] tricuspidatus Heller, 1862c: 267; Plate 3, fg. 15. [im rothen Meere]
Synalpheus tridentulatus (Dana, 1852a)
= Alpheus tridentulatus Dana, 1852a: 22. [in portu Rio Janeiro?]
Synalpheus trispinosus De Man, 1910a: 300. [Madura-bay and other localities in the southern part of
Molo-strait, 54-90 m]
Synalpheus triunguiculatus (Paulson, 1875)
= Alph.[eus] triunguiculatus Paulson, 1875: 103; Plate 14, fgs 1-1g. [Red Sea]
= Synalpheus physocheles Coutire, 1908: 200. [Djibouti]
Synalpheus tropidodactylus Banner & Banner, 1975c: 286; Fig. 2a-k, o. [west of Geraldton, 2814'S 11314'E,
60 fms]
Synalpheus tumidomanus tumidomanus (Paulson, 1875)
= Alph.[eus] tumido-manus Paulson, 1875: 101; Plate 13, fgs 2-2i. [Red Sea]
= Alph.[eus] tumido-manus Var. Alph.[eus] gracili-manus Paulson, 1875: 102; Plate 13, fgs 3-3c. [Red
Sea]
= Synalpheus Hululensis Coutire, 1908: 202. [Maldives]
= Synalpheus Mac-Cullochi Coutire, 1908: 203. [Cte S.W. dAustralie]
= Synalpheus tumidomanus var. exilimanus Coutire, 1909: 10. [lapsus for Alph.[eus] tumido-manus
Var. Alph.[eus] gracili-manus Paulson, 1875]
= Synalpheus Theophane De Man, 1910a: 292. [Lumu-lumu-shoal, Borneo-bank, reef; Anchorage
north of Salomakiee-(Damar)-island, reef; 825'.2S 12718'.4E, anchorage between Nusa-Besi and
the N.E. point of Timor]
= Synalpheus anisocheir Stebbing, 1915: 86; Plate 87. [Gordons Bay, False Bay]
= Synalpheus japonicus Yokoya, 1936: 133; Fig. 3. [vicinity of the Misaki Marine Biological Station]
Synalpheus tumidomanus africanus Crosnier & Forest, 1965b
= Synalpheus hululensis africanus Crosnier & Forest, 1965b: 607; Fig. 2. [So Tom, Morro Peixe, 2-6
m]
Synalpheus tumidomanus congoensis Crosnier & Forest, 1965b
= Synalpheus hululensis congoensis Crosnier & Forest, 1965b: 608; Fig. 3. [Djno, prs de Pointe-
Noire (Congo)]
Synalpheus tuthilli Banner, 1959: 133; Fig. 2. [Tomil Harbor, Yap Island, Caroline Archipelago]
Synalpheus ul (Ros & Dufy, 2007)
= Zuzalpheus ul Ros & Dufy, 2007: 63; Figs 27-30; Plate 5. [Sandbores, Belize, in Hymeniacidion
caerulea, 2 m]
Synalpheus wickstenae Hermoso Salazar & Hendrickx, 2006: 1101; Figs 1-2. [of Altata (2435'N
10755'W), Sinaloa, Mexico, in sponges, 10 m]
Synalpheus williamsi Ros & Dufy, 1999: 542; Figs 1-6. [Carrie Bow Cay, Belize, from Hymeniacidon
caerulea, 18 m]
Synalpheus yano (Ros & Dufy, 2007)
= Zuzalpheus yano Ros & Dufy, 2007: 69; Figs 31-34; Plate 5. [Twin Cays, Belize, in Lissodendoryx
colombiensis, 1-2 m]
Thuylamea Nguyn, 2001
= Thuylamea Nguyn, 2001 (type species Thuylamea camelus Nguyn, 2001, by original designation
Thuylamea camelus Nguyn, 2001: 218; Figs 1-22. [Donghoa, a coastal community of the Can Gio dis-
trict, 21 km northwest of Vung Tau (Cap St Jacques), Vietnam, 1020'N 10652'E, in an estuarine
area formed by the confuence of two large rivers, 5-9 m]
Triacanthoneus Anker, 2010e
= Triacanthoneus Anker, 2010e (type species Triacanthoneus toro Anker, 2010e, by original designa-
Triacanthoneus alacranes Anker, 2010e: 57; Figs 7-9, 10G. [Mexico, Yucatn, Arrecife Alacranes, La An-
egada, 2227'37''N 8936'40.6''W, 0.5 m]
Triacanthoneus pacifcus Anker, 2010e: 54; Figs 5-6, 10D-F. [Panama, Pacifc coast, Playa Venao, inter-
tidal]
Triacanthoneus toro Anker, 2010e: 49; Figs 1-4, 10A-C. [Panama, Caribbean coast, Bocas del Toro, Isla
Coln, Punta Caracol, 1.5 m]
Vexillipar Chace, 1988
= Vexillipar Chace, 1988 (type species Vexillipar repandum Chace, 1988, by original designation and
monotypy, gender neuter)
Vexillipar repandum Chace, 1988: 91; Figs 23-25. [Of Murcielago Bay, Mindanao, Philippines, 847'15''N
12335'00''E, 296 m]
Yagerocaris Kensley, 1988
= Yagerocaris Kensley, 1988 (type species Yagerocaris cozumel Kensley, 1988, by original designation
Yagerocaris cozumel Kensley, 1988: 693; Figs 4-6. [Areolito Cenote, Cozumel, Quintana Roo, Mexico, 40 f]
Family BARBOURIIDAE Christofersen, 1987
Barbouria Rathbun, 1912b
= Barbouria Rathbun, 1912b (type species Barbouria poeyi Rathbun, 1912b (junior subjective synonym
of Hippolyte cubensis von Martens, 1872), by original designation and monotypy, gender feminine;
Barbouria cubensis (von Martens, 1872)
= Hippolyte Cubensis von Martens, 1872: 136; Plate 5, fg. 14. [Cuba]
= Barbouria poeyi Rathbun, 1912b: 455; Plates 2-5. [Cave near seashore, between Morro Castle and
Cojimar]
Barbouria yanezi Meja, Zarza & Lpez, 2008: 663; Figs 1-2. [Cenote Tres Potrillos, Cozumel Island, Mex-
ico, 2027'05''N 8659'15''W, 26 m]
Janicea Manning & C.W.J. Hart, 1984
= Janicea Manning & C.W.J. Hart, 1984 (type species Barbouria antiguensis Chace, 1972, by original
Janicea antiguensis (Chace, 1972)
= Barbouria antiguensis Chace, 1972: 107; Figs 40-41. [Antigua Island, English Harbour]
Parhippolyte Borradaile, 1900
= Parhippolyte Borradaile, 1900 (type species Parhippolyte uveae Borradaile, 1900, by monotypy, gen-
der feminine)
= Somersiella C.W.J. Hart & Manning, 1981 (type species Somersiella sterreri C.W.J. Hart & Manning,
= Koror J. Clark, 1989 (type species Koror misticius J. Clark, 1989, by original designation and mono-
Parhippolyte cavernicola Wicksten, 1996a: 201; Figs 1-4. [San Diego Reef, west of San Diego Island (north
of San Jose Island), Baja California Sur, Mexico, 2520'N 11040'W, 6 m, in sea cave]
Parhippolyte misticia (J. Clark, 1989) (Fig. 49)
= Koror misticius J. Clark, 1989: 446; Figs 1-4. [South Point Cave, Koror, Ngermeuangel, Palau,
718'32"N 13430'05"E]
Parhippolyte rukuensis Burukovsky, 2007c: 3; Fig. 2. [Japan, Ryukyu Islands, Ie-jima, W of Okinawa, in
sea-connected cave, 30 m]
Fig. 49. Parhippolyte misticia (J. Clark, 1989). Photo by John Hoover.
Parhippolyte sterreri (C.W.J. Hart & Manning, 1981)
= Somersiella sterreri C.W.J. Hart & Manning, 1981: 442; Figs 1-28. [Tuckers Town Cave, Tuckers
Town, Bermuda]
Parhippolyte uveae Borradaile, 1900: 414; Plate 38, fgs 11a-g. [Uvea, Loyalty Islands]
Family HIPPOLYTIDAE Spence Bate, 1888
Alcyonohippolyte Marin, Chan & Okuno, 2011
= Alcyonohippolyte Marin, Chan & Okuno, 2011 (type species Alcyonohippolyte dossena Marin, Chan
& Okuno, 2011, by original designation, gender feminine)
Alcyonohippolyte commensalis (Kemp, 1925)
= Hippolyte commensalis Kemp, 1925: 331; Figs 21-22. [Coral reef of Reed Point, Nancowry Island,
Nicobars, on a compound actinian]
Alcyonohippolyte dossena Marin, Chan & Okuno, 2011: 34; Figs 1-4, 16A. [Izu Islands, Hachio-jima Island,
Yaene fshing port, 5 m]
Alcyonohippolyte maculata Marin, Chan & Okuno, 2011: 46; Figs 12-13, 14D-F, 15B-H, 16C. [Taiwan, Kee-
lung, Badouzih, 5-6 m]
Alope White, 1847b
= Alope White, 1847b (type species Alope palpalis White, 1847b (junior subjective synonym of Hip-
polyte spinifrons H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]), by monotypy, gender
= Hetairocaris De Man, 1890 (type species Hetairocaris orientalis De Man, 1890, by monotypy, gender
feminine)
Alope orientalis (De Man, 1890)
= Hetairocaris orientalis De Man, 1890: 122; Plate 6, fg. 16-16c. [Ponap]
= Hippolyte ponapensis Ortmann, 1890: 502; Plate 36, fgs 20-20d. [Karolinen, Ponap]
= Alope australis Baker, 1904: 154; Plate 30, fgs 1-7. [Smiths Bay, Kangaroo Island, shallow water]
Alope spinifrons (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840])
= Hippolyte spinifrons H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 377. [les ctes de la
Nouvelle-Zlande]
= Alope palpalis White, 1847b: 124. [New Zealand]

Bathyhippolyte Hayashi & Miyake, 1970
= Bathyhippolyte Hayashi & Miyake, 1970 (type species Bathyhippolyte yaldwyni Hayashi & Miyake,
Bathyhippolyte yaldwyni Hayashi & Miyake, 1970: 42; Figs 1-16. [Chatham Rise, New Zealand, 4444'S
17542'E, 995-1110 m]
Birulia Branikov, 1903
= Birulia Branikov, 1903 (type species Birulia sachalensis Branikov, 1903, by monotypy, gender
= Paraspirontocaris Yokoya, 1930 (type species Paraspirontocaris kishinouyei Yokoya, 1930, by mono-
Birulia kishinouyei (Yokoya, 1930)
= Paraspirontocaris kishinouyei Yokoya, 1930: 536; Plate 16, fgs 1-6. [on the line between Kanita and
Yokohama, of shima Isl.; of Tairadate]
Birulia sachalinensis Branikov, 1903: 44. [southern Sakhalin Island, Russian Far East]
Bythocarides Sokolov, 2002
= Bythocarides Sokolov, 2002 (type species Bythocarides menshutkinae Sokolov, 2002, by original des-
Bythocarides menshutkinae Sokolov, 2002: 139; Figs 1-3. [Laptev Sea, 8110.8'N 14147.1'E, 945 m]
Bythocaris G.O. Sars, 1870
= Bythocaris G.O. Sars, 1870 (type species Bythocaris simplicirostris G.O. Sars, 1870, by monotypy;
Bythocaris akidopleura Fransen, 1993: 588; Figs 41-62. [Azores, West of Formigas, 3717'N 2514'W, 2070-
2120 m]
Bythocaris biruli Kobjakova, 1964
= Bythocaris leucopis biruli Kobjakova, 1964: 326; Fig. 1. [Arctic Ocean, 8118'N 936'E, 1300 m]
Bythocaris cosmetops Holthuis, 1951a: 135; Fig. 30. [of Sierra Leone, 729'N 1338'W, 74-78 m]
Bythocaris cryonesus Bowman & Manning, 1972: 189; Fig. 1. [Arctic Ocean, 8133.9'N 15712.5'W, 3803
m]
Bythocaris curvirostris Kobjakova, 1957: 363; Figs 1-2. [Arctic Basin, eastern sector, 3255 m]
Bythocaris elegans Bryazgin, 1982: 603; Figs 1-7. [Arctic Basin, western fank of Franz-Victoria Trough,
8151'N 3620'E, 420 m]
Bythocaris foridensis Abele & Martin, 1989: 29; Fig. 1. [Florida, east of Jacksonville, 3109'N 7933'30"W,
644 m]
Bythocaris gorei Abele & Martin, 1989: 38; Fig. 2. [Florida, East of St. Augustine, 2941'N 7955'W, 682 m]
Bythocaris gracilis Smith, 1885a: 497. [Albatross stns 2116, 355'23''N 7431'25''W, 888 fms; 2206,
3935'00''N 7124'30''W, 1043 fms]
Bythocaris grumanti Burukovsky, 1966: 538; Fig. 2. [south end of Spitzbergen, Strait between islands
edge and Nadejda, 50 m]
Bythocaris irene Retowsky, 1946: 298; Fig. 1. [IS G.Sedov stn 35/71, Kara Sea, 520 m according to
Sokolov, 2000]
Bythocaris kobjakovae Sokolov, 2000: 461; Figs 37-39. [Laptev Sea; 1368 m]
Bythocaris leucopis G.O. Sars, 1879: 427. [71.59N 11.40W, 1110 fms]
Bythocaris miserabilis Abele & Martin, 1989: 41; Fig. 3. [Southeast Florida, 2711'N 7930'W, 677 m]
Bythocaris nana Smith, 1885a: 499. [Albatross stns 865, 4005'00''N 7023'00''W, 65 fms; 872, 4005'39''N
7023'52''W, 86 fms; 874, 4000'00''N 7057'00''W, 85 fms; 878, 3955'00''N 7054'15''W, 143 fms (all
of Marthas Vineyard)]
Bythocaris payeri (Heller, 1875a)
= Hippolyte Payeri Heller, 1875a: 609. [full description in Heller, 1875b: 26; Plate 1, fgs 1-4; type lo-
cality therein as Nordpol, 182 m; corrected in Fransen, 1993 to S of Franz Josef Land, 790.4'N
6229.7'E, 182 m]
Bythocaris simplicirostris G.O. Sars, 1870: 149. [Lofoten, 250 orgyarum]
= Hippolyte Panschii Buchholz, 1874: 277; Plate 1, fg. 1. [Nordshannon-Insel, 30 faden]
= Bythocaris spinipleura Squires, 1990: 158; Figs 82-83. [of Bonavista Bay, Newfoundland, 4849'N
5130'W, 309 m, from cod stomachs]
Calliasmata Holthuis, 1973a
= Calliasmata Holthuis, 1973a (type species Calliasmata pholidota Holthuis, 1973a, by original desig-
Calliasmata nohochi Escobar-Briones, Camacho & Alcocer, 1997: 733; Figs 2-4. [Crack House Cave, No-
hoch Nah Chich system, Quintana Roo, Mexico]
Calliasmata pholidota Holthuis, 1973a: 37; Figs 12-13; Plate 1, fg. 2; Plate 2, fg. 1. [Ras Muhammad
Crack, near Ras Muhammad, southern tip of Sinai Peninsula, in pool in narrow crack in elevated
coral rock, about 150 m from the sea]
Calliasmata rimolii Chace, 1975: 37; Figs 5-7. [cave 4 km from town of Estero Hondo (1951'N 7111'W),
Provincia de Puerto Plate, northern Dominican Republic]
Caridion Gos, 1864
= Doryphorus Norman, 1861 (type species Hippolyte Gordoni Spence Bate, 1858, by monotypy, gender
masculine; invalid junior homonym of Doryphorus Cuvier, 1829 (Reptilia); name placed on the Of-
fcial Index of Rejected and Invalid Generic Names in Zoology in Opinion 470 in 1957)
= Caridion Gos, 1864 (nomen novum for Doryphorus Norman, 1861, gender neuter; name placed on
the Ofcial List of Generic Names in Zoology in Opinion 470 in 1957)
Caridion gordoni (Spence Bate, 1858)
= Hippolyte Gordoni Spence Bate, 1858: iv, 52; unnumbered fgures. [Moray Firth]
Caridion steveni Lebour, 1930: 185; Plate 1; Plate 4, fg. 2. [rocky pools under stones at low water and be-
tween tide-marks in bays around Plymouth, Wembury Bay, Rum Bay, Cawsand Bay, Drakes Island,
below Mount Edgcumbe]
Chorismus Spence Bate, 1888
= Chorismus Spence Bate, 1888 (type species Chorismus tuberculatus Spence Bate, 1888, by monotypy,
1957)
Chorismus antarcticus (Pfefer, 1887)
= Hippolyte antarctica Pfefer, 1887: 51; Plate 1, fgs 22-27. [Sd-Georgien, 7-9 Faden]
= Hippolyte Romanchei A. Milne-Edwards, 1891: 45; Plate 5, fgs 1-1d. [Punta Arenas, 25 m]
Chorismus tuberculatus Spence Bate, 1888: 617; Plate 110, fg. 2. [Challenger stn 145A, 4641'S 3810'E, of
Cryptocheles G.O. Sars, 1870
= Cryptocheles G.O. Sars, 1870 (type species Cryptocheles pygmaea G.O. Sars, 1870, by original desig-
nation and monotypy, gender feminine; name placed on the Ofcial List of Generic Names in Zool-
Cryptocheles pygmaea G.O. Sars, 1870: 150. [Lofoten, 120-800 orgyarum]
= Cryptocheles abyssicola M. Sars, 1869: 262, 275. [nomen nudum]
Eualus Thallwitz, 1892
= Eualus Thallwitz, 1892 (type species Eualus obses Thallwitz, 1892 (junior subjective synonym of
Hippolyte Gaimardii H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840), gender masculine;
= Helia Thallwitz, 1892 (type species Hippolyte Fabricii Kryer, 1841, by monotypy, gender feminine;
invalid junior homonym of Helia Hbner (Lepidoptera))
? = Vianellia Nardo, 1847 (type species Vianellia dorsioculata Nardo, 1847, by monotypy, gender femi-
nine; name suppressed for the purposes of the Principle of Priority but not for those of the Principle
of Homonomy under the plenary power of the ICZN, and placed on the Ofcial Index of Rejected
and Invalid Generic Names in Zoology, in Opinion 671, in 1963)
= Lysippe Kinahan, 1858a (type species Hippolyte Cranchii Leach, 1817 [in Leach, 1815-1875], by
monotypy, gender masculine; generic name suppressed for the purposes of both the Principle of
Priority and the Principle of Homonomy under the plenary power of the ICZN, and placed on the
Ofcial Index of Rejected and Invalid Generic Names in Zoology, in Opinion 671, in 1963)
= Spirontocarella Branikov, 1907 (type species Hippolyte macilenta Kryer, 1841, by monotypy, gen-
der feminine)
= Thoralus Holthuis, 1947a (type species Hippolyte Cranchii Leach, 1817 [in Leach, 1815-1875], by
original designation, gender masculine; name placed on the Ofcial List of Generic Names in Zool-
Eualus avinus (Rathbun, 1899)
= Spirontocaris avina Rathbun, 1899: 557. [North of Unalaska Island, 5400'45"N 16653'50"W, 351 fms]
Eualus barbatus (Rathbun, 1899)
= Spirontocaris barbata Rathbun, 1899: 556. [5618'N 16038'W, 86 fms]
Eualus berkeleyorum Butler, 1971: 1616; Figs 1-2. [of Gabriola Island, Strait of Georgia, 4909.0'N
12332.6'W, 384 m]
Eualus biunguis (Rathbun, 1902a)
= Spirontocaris biunguis Rathbun, 1902a: 899. [of Cape St. James, Queen Charlote Islands, British
Columbia, 876 fms]
Eualus bulychevae Kobjakova, 1955: 238; Figs 4-5. [South Kurile Straits, Kuril Islands, max depth 54 m]
Eualus butleri Jensen, 2004: 463; Figs 1-2. [Gastineau Channel, Juneau, Alaska, 5817'N 13424'W, 30 m,
with sponges]
Eualus cteniferus (Barnard, 1950)
= Spirontocaris ctenifera Barnard, 1950: 696; Fig. 129c-k. [Algoa Bay; Durban]
Eualus cranchii (Leach, 1817 [in Leach, 1815-1875])
? = P.[alemon] Microramphos Risso, 1816: 104. [dans les rochers du rivage, environs de Nice]
= Hippolyte Cranchii Leach, 1817 [in Leach, 1815-1875]: Plate 38, Figs 17-21. [southern point of the
Saltstone, in the Kingsbridge Estuary]
? = Hippolyte Crassicornis H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 375. [St Malo,
Britany, France]
= Hippolyte mutila Kryer, 1841: 573. [Norges Vestkyst]
= Hippolyte Bunseni Neumann, 1878: 36. [Palma de Mallorka]
? = Hippolyte Lovenii Rathke, 1843: 13. [Molde]
? = Vianellia dorsioculata Nardo, 1847: 8. [Lagune e Golfo Veneto]
? = Hippolyte Yarrellii W. Thompson, 1853: 112; Plate 6, fgs 2-2b. [Weymouth Bay, 5-7 fms]
Eualus dozei (A. Milne-Edwards, 1891)
= Hippolyte Dozei A. Milne-Edwards, 1891: 46; Plate 5, fgs 3-3c. [ lle Grevy, 65 m]
Eualus drachi Nol, 1978: 23; Figs 1-2. [Banyuls-sur-mer, Mditerrane, 95 m]
Eualus fabricii (Kryer, 1841)
= Hippolyte Fabricii Kryer, 1841: 571. [Grnland]
Eaulus gaimardii belcheri (Bell, 1855)
= Hippolyte belcheri Bell, 1855: 402; Plate 34, fg. 1. [between Beechey Island and Northumberland
Sound]
Eualus gaimardii gaimardii (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840])
? = Cancer nautilor Herbst, 1796 [in Herbst, 1791-1796]: 175; Plate 43, fg. 4.
= Hippolyte Gaimardii H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 378. [les mers
dIslande]
= Hippolyte gibba Kryer, 1841: 572. [Spitsbergen og Grnland]
= Hippolyte recurvirostris Rathke, 1843: 12. [Christiansund]
= Hippolyte lentiginosa Rathke, 1843: 14. [bei Molde und Christiansund]
= Hippolyte Retzii Rathke, 1843: 16. [Meerbusen von Drontheim]
= H.[ippolyte] gracilis Liljeborg, 1851: 83. [Troms]
= Hippolyte pandaliformis Bell, 1851 [in Bell, 1844-1853]: 294; unnumbered vignete. [Loch Fyne,
about 20 fms]
= Eualus obses Thallwitz, 1892: 23. [Grnland]
= Spirontocaris recurvirostris Molander, 1914: 1; Plate 1, fg. 1. [Vaigatet, Greenland, 315 m]
Eualus gracilipes Crosnier & Forest, 1973: 163; Fig. 50. [San Tiago, Cape Verde Islands, 275-150 m]
Eualus gracilirostris (Stimpson, 1860a)
= Hippolyte gracilirostris Stimpson, 1860a: 34. [portu Hakodadi, in regione laminarium]
Eualus horii Komai & Hayashi, 2002: 382; Figs 1-5. [Of Hayatsuki-gawa Rivermouth, Toyama Bay, 80 m]
Eualus kikuchii Miyake & Hayashi, 1967: 261; Figs 6-7. [Tomioka Bay, Amakusa Island]
Eualus kinzeri Tiefenbacher, 1990: 117; Fig. 1. [Weddell Sea, 7106.2'S 1253.8'W, 771 m]
Eualus kuratai Miyake & Hayashi, 1967: 253; Fig. 3. [between Rebun I. and Rishiri I., Hokkaido, 100-
150 m]
Eualus lebourae Holthuis, 1951a: 124; Fig. 26. [of French Guinea, 1049'N 1639'W, 42 m]
Eualus leptognathus (Stimpson, 1860a)
= Hippolyte leptognatha Stimpson, 1860a: 34. [sinu Hakodadi, vulgaris in fundis algoso-arenoso,
prof. 2-6 org.]
= Spirontocaris japonica Yokoya, 1930: 533; Fig. 3. [Between Yuno-Shima isl. and Asamushi, 5-6 fms]
= Spirontocaris fabricii var. minuta Urita, 1942: 25; Fig. 6. [Otomari, 2-3 fms] [nec Spirontocaris minuta
Yokoya, 1930]
Eualus lindbergi Kobjakova, 1955: 240; Fig. 6. [near Levenorn, of Se Sakhalin, 110 m]
Eualus lineatus Wicksten & Butler, 1983: 3; Figs 1-2. [1.5 mi. southwest of Gull Island, of Santa Cruz Is-
land, California, 3356'00"N 11950'55"W, 89 m]
Eualus macilentus (Kryer, 1841)
= Hippolyte macilenta Kryer, 1841: 574 [Grnland]
= Spirontocaris stoneyi Rathbun, 1902a: 899. [Bering Sea, 6215'N 16748'W, 20.5 fms]
Eualus macrophthalmus (Rathbun, 1902a)
= Spirontocaris macrophthalma Rathbun, 1902a: 900. [of Tawhit Head, Washington, 178 fms]
Eualus middendorf Branikov, 1907: 165; Fig. 23. [Sea of Okhotsk, of South Sakhalin Island]
Eualus occultus (Lebour, 1936)
= Spirontocaris occulta Lebour, 1936: 96; Plate 1; Plate 2, fg. 2. [Plymouth]
? = Hippolyte Crassicornis H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 375. [la rade de
Saint-Malo]
? = Hippolyte Lovenii Rathke, 1843: 13. [Molde]
? = Vianellia dorsiocullata Nardo, 1847: 8. [Lagune e Golfo Veneto]
? = Hippolyte Yarrellii W. Thompson, 1853: 112; Plate 4, fg. 2. [Weymouth Bay]
Eualus pax (Stebbing, 1915)
= Spirontocaris pax Stebbing, 1915: 91; Plate 24. [3411'N 1831'E, 20 fms; of Bufels Bay (False Bay),
30 fms]
Eualus pectiniformis Hanamura, 2008: 88; Figs 1-3. [3808.3'S 14931.9'E, 150-189 m]
Eualus pusiolus (Kryer, 1841)
= Hippolyte pusiola Kryer, 1841: 576. [Norges Vestkyst]
= Hippolyte subula Rathke, 1843: 9. [bei Molde]
= Hippolyte vitata Rathke, 1843: 10. [Christiansund]
= Hippolyte Barleei Spence Bate, 1852: 357; Plate 5B(1). [of the Haaf, Shetlands]
= H.[ippolyte] Korenii Danielssen, 1859: 6. [Vadse paa Leerbund, 60 fms]
= Hippolyte Andrewsii Kinahan, 1860a: 48. [Dalkey Sound]
Eualus ratmanovi Makarov, 1941: 125. [Bering Sea, stns 40, 51 (1932); 10, 16, 18 (1933)]
Eualus sinensis (Yu, 1931b)
= Spirontocaris sinensis Yu, 1931b: 514; Fig. 2. [Chefoo]
Eualus sollaudi (Zariquiey Cenarro, 1936)
= Thor Sollaudi Zariquiey Cenarro, 1936: 242; Figs 17-21. [Cadaqus, Arenys]
Eualus spathulirostris (Yokoya, 1933)
= Spirontocaris spathulirostris Yokoya, 1933: 28; Fig. 10. [Sy-Maru stn 70, near middle between Siri-
ya-zaki and Todo-saki, 285 m; 474, north-west of Is. Tusima, 219 m; 652, Tugaru Strait, 110 m]
Eualus suckleyi (Stimpson, 1864)
= Hippolyte Suckleyi Stimpson, 1864: 154. [Puget Sound, circumlitoral zone]
Eualus subtilis Carvacho & Olson, 1984: 61; Figs 1-2. [Punta Banda, extremo sur de Baha de Todos San-
tos, Baja California, 30 m]
Eualus townsendi (Rathbun, 1902a)
= Spirontocaris townsendi Rathbun, 1902a: 897. [Washington Sound, Straits of Fuca, 48 fms]
Eumanningia Crosnier, 2000
= Eumanningia Crosnier, 2000 (type species Eumanningia pliarthron Crosnier, 2000, by original desig-
Eumanningia pliarthron Crosnier, 2000: 110; Figs 1-4. [South-West Pacifc, Wallis Island, 1411.0'S
17617.3'W, 890-915 m]
Exhippolysmata Stebbing, 1915
= Exhippolysmata Stebbing, 1915 (type species Hippolysmata ensirostris Kemp, 1914, designated by
Holthuis, 1955b, gender feminine)
Exhippolysmata ensirostris ensirostris (Kemp, 1914)
= Hippolysmata ensirostris Kemp, 1914: 113, 118; Plate 7, fgs 1-4. [Colombo]
Exhippolysmata ensirostris punctata (Kemp, 1914)
= Hippolysmata ensirostris var. punctata Kemp, 1914: 120; Plate 7, fgs 5-7. [Sandheads, Ganges delta;
Green Isl., Amherst, Tenasserim; Thongwa, Burma]
Exhippolysmata hastatoides (Balss, 1914a)
= Mimocaris hastatoides Balss, 1914a: 596. [Victoria, Cameroon, Reuse (Geringe tiefe)]
Exhippolysmata oplophoroides (Holthuis, 1948)
= Hippolysmata (Exhippolysmata) oplophoroides Holthuis, 1948: 1106; Figs 2-3. [Mouth of Surinam
River, near Resolutie]
Exhippolysmata tugelae Stebbing, 1915: 94; Plate 89. [Of South Head, Tugela River, 12 fms; Cape Hender-
son, NW 2.5 miles, 26 fms]
Gelastocaris Kemp, 1914
= Gelastocaris Kemp, 1914 (type species Latreutes Paronae Nobili, 1905f, by monotypy, gender femi-
Gelastocaris paronae (Nobili, 1905f)
= Latreutes Paronae Nobili, 1905f: 2. [isola di Zanzibar]
Gelastreutes Bruce, 1990h
= Gelastreutes Bruce, 1990h (type species Gelastreutes crosnieri Bruce, 1990h, by original designation
Gelastreutes crosnieri Bruce, 1990h: 139; Figs 1-4. [New Caledonia, 1908'30"S 16329'30"E, 65-120 m]
Heptacarpus Holmes, 1900
= Heptacarpus Holmes, 1900 (type species Hippolyte palpator Owen, 1839, by original designation,
1957)
Heptacarpus acuticarinatus Komai & Ivanov, 2008: 9; Figs 5-9. [Sagami Bay, 3507.858'N 13933.698'E, 100-
101 m]
Heptacarpus brachydactylus (Rathbun, 1902a)
= Spirontocaris brachydactyla Rathbun, 1902a: 898. [of Santa Cruz Island, 266 fms]
Heptacarpus brevirostris (Dana, 1852a)
= Hippolyte brevirostris Dana, 1852a: 24. [in freto de Fuca, juxta portum Dungeness]
Heptacarpus camtschaticus (Stimpson, 1860a)
= Hippolyte camtschatica Stimpson, 1860a: 33. [Type locality not indicated]
Heptacarpus carinatus Holmes, 1900: 202; Plate 3, fg. 60. [Monterey Bay, shallow water]
Heptacarpus commensalis Hayashi, 1979: 14; Figs 1-2. [just north of Seto Marine Biological Laboratory,
Shirahama, from Acropora sp.]
Heptacarpus decorus (Rathbun, 1902a)
= Spirontocaris decora Rathbun, 1902a: 896. [of Santa Cruz Island, California, 150 fms]
Heptacarpus fexus (Rathbun, 1902a)
= Spirontocaris fexa Rathbun, 1902a: 896. [north of Bird Island, Shumagins, Alaska, 21 fms]
Heptacarpus franciscanus (Schmit, 1921)
= Spirontocaris franciscana Schmit, 1921: 60; Plate 12, fgs 8-9. [San Francisco Bay]
Heptacarpus fuscimaculatus Wicksten, 1986: 47; Figs 1-2. [Big Fishermans Cove, Santa Catalina Island,
California, 3327'N 11828'W, among low-growing algae on foating dock]
Heptacarpus futilirostris (Spence Bate, 1888)
= Nauticaris futilirostris Spence Bate, 1888: 606; Plate 109, fg. 1. [Challenger stn 233A, 3438'N
13501'E, of Japan, 50 fms]
Heptacarpus geniculatus (Stimpson, 1860a)
= Hippolyte geniculata Stimpson, 1860a: 34. [Sinu Hakodadi, inter lapides ad prof 2 org.]
= Spirontocaris alcimede De Man, 1906: 404. [Inland sea of Japan]
Heptacarpus grebnitzkii (Rathbun, 1902b)
= Spirontocaris grebnitzkii Rathbun, 1902b: 44; Fig. 18. [Mororan, Hokkaido]
Heptacarpus herdmani (Walker, 1898)
= Spirontocaris herdmani Walker, 1898: 277; Plate 16, fg. 2. [Puget Sound]
Heptacarpus igarashii Hayashi & Chiba, 1989: 71; Figs 1-3. [Toni Bay, Kamaishi City, Iwate Prefecture]
Heptacarpus jordani (Rathbun, 1902b)
= Spirontocaris jordani Rathbun, 1902b: 44; Fig. 17. [Hakodata, Hokkaido]
Heptacarpus kincaidi (Rathbun, 1902a)
= Spirontocaris kincaidi Rathbun, 1902a: 899. [of Santa Cruz, California, 21 fms]
Heptacarpus longirostris (Kobjakova, 1936a)
= Eualus geniculata var. longirostris Kobjakova, 1936a: 211; Fig. 38. [Peter the Great Bay, Sea of Japan;
fully described in Kobjakova, 1937]
Heptacarpus maxillipes (Rathbun, 1902a)
= Spirontocaris maxillipes Rathbun, 1902a: 898. [Of Segouam, Aleutians, 283 fms]
Heptacarpus minutus (Yokoya, 1930)
= Spirontocaris minuta Yokoya, 1930: 531; Fig. 2. [Of Arito, 19 fms]
Heptacarpus moseri (Rathbun, 1902a)
= Spirontocaris moseri Rathbun, 1902a: 897. [of Segouam, Aleutians, 283 fms]
Heptacarpus palpator (Owen, 1839)
= Hippolyte palpator Owen, 1839: 89; Plate 28, fg. 3. [Monteray, California]
= Hippolyte? Hemphillii Lockington, 1877a: 35. [San Diego]
Heptacarpus paludicola Holmes, 1900: 201; Plate 3, fgs 56-57. [Humboldt Bay; Shelter Cove; Bodega Bay]
Heptacarpus pandaloides (Stimpson, 1860a)
= Hippolyte pandaloides Stimpson, 1860a: 34. [Sinu Hakodadi, inter lapides ad prof 2 org.]
= Spirontocaris propugnatrix De Man, 1906: 404. [Inland Sea of Japan, 6 fms]
Heptacarpus pugetensis Jensen, 1983: 314; Figs 1-3. [Alki Point, Seatle, Washington, 4734'N 12225'W,
low intertidal, under rock]
Heptacarpus rectirostris (Stimpson, 1860a)
= Hippolyte rectirostris Stimpson, 1860a: 33. [Portu Hakodadi Japoni borealis, in locis profundis
maris]
Heptacarpus sitchensis (Brandt, 1851)
= Hippolyte sitchensis Brandt, 1851: 116; Plate 5, fgs 18-18c. [Insel Sitcha]
= Hippolyte picta Stimpson, 1871: 125. [Monterey, Cal.]
= Heptacarpus litoralis Butler, 1980: 220; unnumbered fgure; Fig. 17. [Bunsby Islands, 2-9 m]
Heptacarpus stimpsoni Holthuis, 1947a: 13. [nomen novum for Hippolyte cristata Stimpson, 1860a]
= Hippolyte cristata Stimpson, 1860a: 33, nec De Haan, 1844 [in De Haan, 1833-1850]. [Portu San
Francisco Californiae, fundo arenoso prof. 5-10 org.]
Heptacarpus stylus (Stimpson, 1864)
= Hippolyte stylus Stimpson, 1864: 154. [Straits of De Fuca]
= Hippolyte esquimaltiana Spence Bate, 1864: 666. [Esquimalt Harbor]
Heptacarpus taylori (Stimpson, 1857)
= Hippolyte taylori Stimpson, 1857: 500. [Monterey]
Heptacarpus tenuissimus Holmes, 1900: 203. [Monterey]
= Hippolyte gracilis Stimpson, 1864: 155; nec Liljeborg, 1851. [Puget Sound, deep water]
= Hippolyte amabilis Lenz, 1901: 432; Plate 32, fgs 2-3. [Bare Island]
Heptacarpus tridens (Rathbun, 1902a)
= Spirontocaris tridens Rathbun, 1902a: 896. [Admiralty Inlet, Puget Sound, 40 fms]
Heptacarpus yaldwyni Wicksten, 1984b: 241; Figs 1-3. [of Salina Cruz, Mexico, 1447'N 9619'W
1450.5'N 9613'W, 1052-1145 m]
Hippolyte Leach, 1814 [in Leach, 1813-1814]
= Hippolyte Leach, 1814 [in Leach, 1813-1814] (type species Hippolyte Varians Leach, 1814 [in Leach,
= Nectoceras Rafnesque, 1817 (type species Nectoceras pelagica Rafnesque, 1817, by monotypy, gen-
der neuter, a junior subjective synonym of Astacus coerulescens Fabricius, 1775; name placed on the
Ofcial Index of Rejected and Invalid Names in Zoology in Opinion 522 in 1958)
= Virbius Stimpson, 1860a (type species Hippolyte acuminatus Dana, 1852a (junior subjective syno-
nym of Astacus coerulescens Fabricius, 1775), designated by Kingsley, 1880, gender masculine)
= Bellidia Gosse, 1877 (type species Bellidia Huntii Gosse, 1877, by monotypy, gender feminine)
Hippolyte acuta (Stimpson, 1860a)
= Virbius acutus Stimpson, 1860a: 35. [ad insulam Loo Choo, litoralis in rupibus algosis]
Hippolyte australiensis (Stimpson, 1860a)
= Virbius australiensis Stimpson, 1860a: 35. [in portu Jacksoniensi Australiae, inter algas ad prof. org. 2]
= Caradina cincinnuli Spence Bate, 1863: 500; Plate 40, fgs 3-3z. [Gulf St. Vincent, 4.5 fms]
Hippolyte bifdirostris (Miers, 1876a)
= Virbius bifdirostris Miers, 1876a: 224. [New Zealand; illustrated in Miers, 1876b]
Hippolyte californiensis Holmes, 1895: 576; Plate 20, fgs 21-26. [Bodega Bay]
Hippolyte caradina Holthuis, 1947a: 14. [nomen novum for Caradina tenuirostris Spence Bate, 1863 nec
Hippolyte tenuirostris H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]]
= Caradina tenuirostris Spence Bate, 1863: 501; Plate 40, fgs 4-4k. [Gulf St. Vincent, 4.5 fms]
Hippolyte catagrapha dUdekem dAcoz, 2007: 185; Figs 1-4. [South Africa, False Bay, 6-8 m, from the cri-
noid Tropiometra carinata]
Hippolyte clarki Chace, 1951: 37; Fig. 1f-p. [Friday Harbor, Washington, in eel grass]
Hippolyte coerulescens (Fabricius, 1775)
= Astacus coerulescens Fabricius, 1775: 414. [Pelago inter Tropicos frequens]
= Palaemon pelasgicus Bosc, 1802: 105; Plate 14, fg. 2. [la haute mer, sur fucus nageans]
= Nectoceras pelagica Rafnesque, 1817: 41. [Atlantic Ocean, in the gulph stream on the Fucus natans]
= Hippolyte tenuirostris H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 374. [en haute
mer, prs de Aores]
= Hippolyte acuminatus Dana, 1852a: 24. [in mari Atlantico cum Sargasso lat. bor. 3607' - 407', long.
occid. 2043' - 7136']
= Hippolyte bidentatus Spence Bate, 1888: 591; Plate 105, fgs 1-2. [Atlantic Ocean, on gulf-weed; be-
tween 3207'-3529'N and 5053'-5232'W]
= Hippolyte Martiali A. Milne-Edwards, 1891: 47; Plate 6, fgs 1-1g. [canal du Beagle, en vue de
Loupataya, 198 m]
Hippolyte edmondsoni Hayashi, 1981a: 185; Figs 1-3. [Waimanalo, Oahu]
Hippolyte garciarasoi dUdekem dAcoz, 1996b: 16; Figs 5-6, 8-10. [Banyuls, between algae on rocks at the
laboratory Arago, 4229'N 0308'E, 0-1 m]
Hippolyte holthuisi Zariquiey lvarez, 1953: 104; Figs 3-4, 6, 8. [Cadaqus, junto al Cabo de Creus, 22-60 m]
Hippolyte inermis Leach, 1816b: 347. [Cornwall (United Kingdom), Plymouth, 5013'N 0410'W; neotype
selection by dUdekem dAcoz, 1996b]
? = Carida viridis Rafnesque, 1814: 24. [Sicile; nomen nudum]
= P.[alemon] Olivieri Risso, 1816: 107. [dans les fucus et les conferves, environs de Nice]
= P.[alemon] Margaritaceus Risso, 1816: 108. [dans les endroits pierreux, environs de Nice]
= Alpheus viridis Oto, 1821: 12. [in mari circa Nicaeam; fully described in Oto, 1828]
= A.[lpheus] elongatus Risso, 1827: 77. [golfe de Nice, dans les fucus]
= Hippolytus Brullei Gurin, 1832: 41; Plate 27, fg. 2. [Grce]
= Hippolyte virescens H. Milne Edwards, 1837 [in H. Milne Edwards, 1836-1844]: Plate 53, fgs 3-3b
(as Hippolyte viridis). [Type locality not indicated]
= Hippolyte mauritanicus Lucas, 1846: 42; Plate 4, fg. 3. [rade de Mers-el-Kebir; Lager, Stora, Bne]
= Hippolyte Whitei W. Thompson, 1853: 110; Plate 6, fgs 1-1a. [Weymouth Bay, 4-6 fms]
= Hippolyte Whitei var. ensis W. Thompson, 1853: 111; Plate 6, fg. 1b. [Weymouth Bay]
= Hippolyte Whitei var. falcatus W. Thompson, 1853: 112; Plate 6, fg. 1c. [Weymouth Bay]
= Hippolyte Mitchelli W. Thompson, 1853: 114; Plate 6, fgs 4-4a. [Weymouth Bay]
= Hippolyte producta Norman, 1861: 275; Plate 13, fg. 5. [Lamlash Bay]
Hippolyte jarvinensis Hayashi, 1981a: 190; Figs 4-5. [Jarvis Islands, the Line Islands]
Hippolyte kraussiana (Stimpson, 1860a)
= Virbius Kraussianus Stimpson, 1860a: 36. [in sinu Simons Bay prope promontorium Bonae Spei]
= Virbius capensis Lenz & Strunck, 1914: 319; Plate 20, fgs 1-4. [Simonsbai]
Hippolyte lagarderei dUdekem dAcoz, 1995: 494; Fig. 1. [Maroc, cte Atlantique, Tmara, 3355'N
0655'W]
Hippolyte leptocerus (Heller, 1863a)
= Virbius leptocerus Heller, 1863a: 289; Plate 10, Figs 5-6. [Hafen von Genua]
= V.[irbius] gracilis Heller, 1862b: 399; nec H.[ippolyte] gracilis Liljeborg, 1851. [aus dem adriatischen
Meere]
= Virbius gracilis varietas intermedia Czerniavsky, 1868: 68; Plate 5, fgs 2-7. [sinu Jaltensi]
= Virbius gracilis var. longirostris Czerniavsky, 1868: 68; Plate 5, fgs 8-25. [Black Sea]
= Virbius gracilis forma typica Czerniavsky, 1884: 14. [Black Sea]
= Virbius gracilis var. articulirostris Czerniavsky, 1884: 15; Plate 1, fg.1a. [Black Sea]
= Virbius gracilis var. brevirostris Czerniavsky, 1884: 15; Plate 1, fgs 1p-s. [Black Sea]
= Virbius Brullei var. elongata Czerniavsky, 1884: 18. [sinus Jaltensis, zona litoralis; sinus Sevastopol]
= Virbius Brullei forma fortior Czerniavsky, 1884: 19; Plate 2, fgs 3a-n. [sinus Jaltensis, zona litoralis,
1-1.5 m; sinus Suchum, zona litoralis, 1-1.5 m]
= Virbius tenuirostris Czerniavsky, 1884: 20; Plate 2, fgs 4a-g. [sinus Suchum, zona litoralis, 1-1.5 m]
= Virbius rectifrons Czerniavsky, 1884: 21; Plate 1, fg. 2. [sinus Suchum, zona litoralis, 1-1.5 m]
= Hippolyte longirostris armoricana Sollaud in Bourdon, 1965: 6, 39. [ctes bretonnes]
Hippolyte leptometrae Ledoyer, 1969: 343; Plate 2. [ctes de Provence, 110 m]
Hippolyte longiallex dUdekem dAcoz, 2007: 193; Figs 7-9. [West Africa, Gulf of Guinea, Principe Island,
Pedra da Gal, 35 m, from Muriceops tuberculata]
Hippolyte multicolorata Yaldwyn, 1971: 90. [Island Bay, Wellington, from intertidal algae]
Hippolyte nicholsoni Chace, 1972: 113; Figs 46-47. [Milford Bay, between Pigeon Point and Crown Point,
Tobago, 9-12 m]
Hippolyte niezabitowskii dUdekem dAcoz, 1996b: 58; Figs 25-28, 29a. [Ionian coast of Greece, Drepanos,
western part, 3930'N 2015'E, small seagrasses on non muddy sand]
Hippolyte obliquimanus Dana, 1852a: 24. [in portu Rio Janeiro]
= Hippolyte exilirostratus Dana, 1852a: 24. [in portu Rio Janeiro]
= Virbius gracilis var. brasiliensis Czerniavsky, 1884: 14. [Type locality not indicated]
= Hippolyte curaaoensis Schmit, 1924a: 68; Fig. 4. [West Punt, Curaao]
Hippolyte palliola Kensley, 1970: 183; Figs 1-2. [4 or 5 miles south of the Kunene River mouth, South West
Africa, in brown algae growing on intertidal rocks]
Hippolyte pleuracantha (Stimpson, 1871)
= Virbius pleuracanthus Stimpson, 1871: 127. [harbor of Norfolk, Va.; at Somers Point, in Great Egg
Harbor, N. J.]
Hippolyte prideauxiana Leach, 1817 [in Leach, 1815-1875]
= Hippolyte Prideauxiana Leach, 1817 [in Leach, 1815-1875]: Plate 38, fgs 1, 3-5. [Near Bantham, on
the southern coast of Devon]
= Hippolyte Moorii Leach, 1817 [in Leach, 1815-1875]: Plate 38, fg. 2. [Plymouth Sound]
= Bellidia Huntii Gosse, 1877: 313; Plate 10. [of the Shag Rock, northern end of Torbay, 6 fms]
Hippolyte proteus (Paulson, 1875)
= V.[irbius] Proteus Paulson, 1875: 109, Plate 16, fgs 2-5, Plate 18, fgs 1-1k. [Red Sea]
Hippolyte sapphica dUdekem dAcoz, 1993: 56; Figs 1, 5, 7, 8-9. [20 m lest du port de Skala Kallonis,
herbiers de Zostracees, 0.5-1 m]
= Hippolyte sapphica forma A dUdekem dAcoz, 1996b: 79; Figs 38-39. [Gulf of Kallonis, 20 m at East
of Skala Kallonis harbour, 3912'N 2615'E, 0.5-1 m]
= Hippolyte sapphica forma B dUdekem dAcoz, 1996b: 84; Figs 40-43. [Gulf of Amvrakikos]
Hippolyte varians Leach, 1814 [in Leach, 1813-1814]
= Hippolyte Varians Leach, 1814 [in Leach, 1813-1814]: 432. [rocky shores of Devon]
= Hippolyte smaragdina Kryer, 1841: 570. [Norges Vestkyst og Kategatet]
= Hippolyte fascigera Gosse, 1853: 153. [Weymouth Bay, a few miles from land]
= Hippolyte Grayana W. Thompson, 1853: 113; Plate 6, fgs 3-3a. [Weymouth Bay, 4 fms]
= Caradina tenuis Spence Bate, 1866: 28; Plate 2, fg. 1. [Plymouth]
Hippolyte ventricosa H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]
= Hippolyte ventricosus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 371. [mers dAsie;
India according to dUdekem dAcoz, 1999]
= H.[ippolyte] orientalis Heller, 1862c: 277. [im rothen Meere]
? = Virbius Mossambicus Hilgendorf, 1879: 836; Plate 4, fg. 1. [Zambeze]
Hippolyte williamsi Schmit, 1924c: 163; Fig. 40. [Eden, rock pools]
= Hippolyte mexicana Chace, 1937b: 127; Fig. 6. [Santa Inez Bay, 2650'45''N 11154'20''W, 1 fm]
Hippolyte zostericola (Smith, 1873)
= Virbius zostericola Smith, 1873: 550; Plate 3, fg. 11. [among eel-grass about Vineyard Sound]
= Hippolyte pleuracantha bermudensis Gurney, 1936: 27; Plate 1, fgs 4-12; Plate 2, fgs 13-21. [Zostera-
bed in Walsingham Bay, on the west of Castle Harbour; in Sargassum growing on rocks at Tobacco
Bay on the north shore]
Latreutes Stimpson, 1860a
= Cyclorhynchus De Haan, 1849 [in De Haan, 1833-1850] (type species Hippolyte planirostris De Haan,
1844 [in De Haan, 1833-1850], by monotypy, gender neuter; invalid junior homonym of Cyclorhyn-
chus Kaup, 1829 (Aves), Cyclorhynchus Sundevall, 1836 (Aves) and Cyclorhynchus Macquart, 1841
(Diptera))
= Latreutes Stimpson, 1860a (type species Hippolyte ensiferus H. Milne Edwards, 1837 [in H. Milne
Edwards, 1834-1840], designated by Kingsley, 1880, gender masculine; name placed on the Ofcial
List of Generic Names in Zoology in Opinion 470 in 1957)
= Rhynchocyclus Stimpson, 1860a (nomen novum for Cyclorhynchus De Haan, 1849 [in De Haan, 1833-
1850], gender masculine; invalid junior homonym of Rhynchocyclus Cabanis & Heine, 1859 (Aves))
= Concordia Kingsley, 1880 (type species Concordia gibberosus Kingsley, 1880 (junior subjective syno-
nym of Rhynchocyclus parvulus Stimpson, 1871), by monotypy, gender feminine)
= Platybema Spence Bate, 1888 (nomen novum for Cyclorhynchus De Haan, 1849 [in De Haan, 1833-
1850] and Rhynchocyclus Stimpson, 1860a, gender neuter)
Latreutes acicularis Ortmann, 1890: 506; Plate 37, fg. 6. [Japan, Kadsiyama]
Latreutes anoplonyx Kemp, 1914: 104; Plate 4, fgs 3-5. [Bombay]
Latreutes antiborealis Holthuis, 1952d: 62; Fig. 14. [Golgo de Ancud, Canal San Antonio, inner part,
4144'10"S 7315'15"W, 15 m]
Latreutes compressus (Stimpson, 1860a)
= Rhynchocyclus compressus Stimpson, 1860a: 28. [in portu Jackson Australiae, f. algoso p. 2 org.]
= Caradina truncifrons Spence Bate, 1863: 499; Plate 40, fgs 2-2u. [Gulf St. Vincent, 4.5 fms]
Latreutes foliirostris Kobjakova, 1935: 91; Fig. 4. [Bucht Peters des Groen]
Latreutes fucorum (Fabricius, 1798)
= Palaemon fucorum Fabricius, 1798: 404. [in Oceani Fuco natante]
= Hippolyte ensiferus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 374. [haute mer,
prs de Aores]
Latreutes inermis Chace, 1972: 122; Figs 51-52. [Reef just south of Marigot Harbour, Saint Lucia Island, 4-6 m]
Latreutes laminirostris Ortmann, 1890: 506; Plate 37, fg. 5. [Japan, Tanagava]
Latreutes mucronatus (Stimpson, 1860a)
= Rhynchocyclus mucronatus Stimpson, 1860a: 27. [in freto Ly-I-moon prope Hong Kong, f. con-
choso p. 25 org]
= Latreutes Gravieri Nobili, 1904: 231. [Djibouti]
= Latreutes mucronatus var. multidens Nobili, 1905b: 394. [Mer Rouge, Java]
= Latreutes natalensis Lenz & Strunck, 1914: 320; Plate 21, fgs 1-11. [Port Natal]
Latreutes parvulus (Stimpson, 1871)
= Rhynchocyclus parvulus Stimpson, 1866: 48. [nomen nudum]
= Rhynchocyclus parvulus Stimpson, 1871: 124. [St. Joseph I., Texas]
= Concordia gibberosa Kingsley, 1880: 414. [Fort Macon (Beaufort Inlet), North Carolina, U.S.A.]
Latreutes phycologus Nobili, 1905a: 159; unnumbered fgure. [Ctes dArabie, sur une algue brune fot-
tante]
Latreutes planirostris (De Haan, 1844 [in De Haan, 1833-1850])
= Hippolyte planirostris De Haan, 1844 [in De Haan, 1833-1850]: Plate 45, fg. 7. [Japan]
= Latreutes dorsalis Stimpson, 1860a: 27. [in sinu Hakodadi Japoniae, prof. 8 org.]
Latreutes planus Spence Bate, 1888: 584; Plate 89, fg. 5. [Of Sibago Island, Samboangan, Philippine Is-
lands]
Latreutes porcinus Kemp, 1916a: 397; Fig. 3; Plate 36, fg. 3. [jety on Ross Island, Port Blair, Andamans,
among weeds]
Latreutes pristis (Nobili, 1899)
= Platybema pristis Nobili, 1899: 233. [Beagle Bay, sulle secche fra Beagle e Volverine Passage]
Latreutes pymoeus Nobili, 1904: 231. [Djibouti; original spelling maintained under Art. 32.1]
Latreutes unidentatus Spence Bate, 1888: 586; Plate 89, fg. 6. [Of Sibago, Samboangan, Philippine Is-
lands]
Lebbeus White, 1847a
= Lebbeus White, 1847a (type species Lebbeus orthorhynchus White, 1847a (junior subjective synonym
of Alpheus Polaris Sabine, 1824), by monotypy, gender masculine; name conserved under the ple-
nary power of the ICZN, and placed on the Ofcial List of Generic Names in Zoology in Opinion
671 in 1963)
= Hetairus Spence Bate, 1888 (type species Alpheus Polaris Sabine, 1824, designated under the ple-
nary powers of the ICZN, gender masculine; name placed on the Ofcial List of Rejected and Inva-
lid Names in Zoology in Opinion 671 in 1963)
= Birulaecaris Dons, 1915 (type species Hippolyte mysis Birulya, 1898 (junior subjective synonym of
Alpheus Polaris Sabine, 1824), gender feminine)
Lebbeus acudactylus Jensen, 2006: 34; Figs 4D, 7-9. [Aleutian Islands, Alaska, 51.86N 174.92W, 155 m]
Lebbeus africanus Fransen, 1997a: 231; Figs 1-29. [Mauritania, of Banc dArguin, 1943'N 1730'W, 1500
m]
Lebbeus antarcticus (Hale, 1941)
= Spirontocaris antarcticus Hale, 1941: 267; Figs 5-6. [Of Adelie Coast, Wilkes Land, Antarctica,
6621'S 13828'E, 640 m]
Lebbeus balssi Hayashi, 1992: 112; Figs 1-3. [East China Sea, 3359.4'N 12848.0'E, 102 m]
Lebbeus bidentatus Zarenkov, 1976: 13; Fig. 5. [R.V. Akademik Kurchatov stn 276, of Peru, 1680 m]
Lebbeus brandti (Branikov, 1907)
= Hetairus brandti Branikov, 1907: 157; Fig. 20. [near Cape Terpeniya, East coast of Sakhaln Island,
Sea of Okhotsk]
Lebbeus brevicornis Komai, 2011: 291; Figs 5-8. [R/V Rinkai-maru, Sagami Bay, of Misaki, 351-338 m]
Lebbeus brevirostris Chang, Komai & Chan, 2010: 727; Figs 1-3, 10A-B. [2452.33'N 12203.10'E, 600 m]
Lebbeus carinatus Zarenkov, 1976: 9; Fig. 2. [R.V. Akademik Kurchatov stn 290, of Peru, 1850 m]
Lebbeus catalepsis Jensen, 1987: 89; Figs 1-3. [Strait of Juan de Fuca between Sekiu and Neah Bay, Washing-
ton, 4819'N 12428'W, low intertidal]
Lebbeus clarehannah McCallum & Poore, 2010: 127; Figs 1C, 2-3. [Western Australia, of Bald Island,
3512.49'S 11839.04'E - 3512.14'S 11840.08'E, 408-431 m]
Lebbeus comanthi Hayashi & Okuno, 1997: 49; Figs 1-2, 3(2), 4(3). [Izu Ocean Park, Sagami Bay, 15 m, from
Oxycomanthus japonicus]
Lebbeus compressus Holthuis, 1947a: 9, 40. [nomen novum for Spirontocaris gibberosa Yokoya, 1933 sensu
Balss, 1914b]
= Spirontocaris gibberosa Yokoya, 1933: 25; Fig. 8. [Sy-Maru stn 26, north-east of Siwoya-zaki, 232 m]
Lebbeus cristagalli McCallum & Poore, 2010: 131; Figs 1A-B, 4. [Western Australia, north-west Australia,
near Ashmore Reef, 1228.53'S 12325.04'E 1229.58'S 12325.00'E, 397-405 m]
Lebbeus cristatus Ahyong, 2010: 341; Figs 1-2, 3A. [3654.76'S 16732.10'E, Challenger Plateau, New Zea-
land, 1231-1226 m]
Lebbeus curvirostris Zarenkov, 1976: 12; Fig. 4. [R.V. Akademic Kurchatov stn 290, of Peru, 1680-1860 m]
Lebbeus elegans Komai, Hayashi & Kohtsuka, 2004: 103; Figs 1-3. [of Togi, Hakui District, Ishikawa Pre-
fecture, west coast of Noto Peninsula, Sea of Japan, 400 m]
Lebbeus eludus Jensen, 2006: 24; Figs 1-3, 4A-B. [Burrows Channel, Anacortes, Washington, 48.29.4N
122.41.7W, 14 m]
Lebbeus fasciatus (Kobjakova, 1936a)
= Hetairus fasciata Kobjakova, 1936a: 222; Fig. 17. [Okhotsk Sea; fully described in Kobjakova, 1937]
= Hetairus zebra Makarov, 1935: 319; Fig. 1; nec Leim, 1921. [Bering-Insel, Bucht Nikolskaja, sandiges
Ufer am Ende der Bucht; Bering-Insel, Fazies der Steine und Felsen im Litoral nahe dem Dorf Nokol-
skaja; Bering-Meer, 5953'N 17026'E, 32 m; Awatsch-Golf, nahe der Bucht Betschewinskaja, 30 m]
= Hetairus zebra Makarov, 1941: 123. [Karaginsky Island, Bering Island, Mednyi Island, Avanchisky
Gulf (near Becheviskaya Bay), Bering Sea stn 85]
= Spirontocaris makarof Urita, 1942: 18. [nomen novum for Hetairus zebra Makarov, 1935]
Lebbeus formosus Chang, Komai & Chan, 2010: 732; Figs 4-5, 10C. [2206.3345'N 12108.2237'E, 1260-1275
m]
Lebbeus grandimanus (Branikov, 1907)
= Hetairus grandimana Branikov, 1907: 152; Fig. 18. [coasts of Kamtchatka, Sea of Okhotsk; Sea of
Japan, 19-118 m]
Lebbeus groenlandicus (Fabricius, 1775)
= Astacus Groenlandicus Fabricius, 1775: 416. [in mari groenlandico]
= Cancer aculeatus O. Fabricius, 1780: 239. [Naularnak]
= Hippolite armata Owen, 1839: 88; Plate 27, fg. 2. [shores of Kamtschatka]
= Hippolite cornuta Owen, 1839: 89; Plate 28, fg. 2. [Type locality not indicated]
Lebbeus heterochaelus (Kobjakova, 1936a)
= Hetairus heterochaela Kobjakova, 1936a: 222; Figs 18-19. [Okhotsk Sea, 165 m; fully described in
Kobjakova, 1937]
Lebbeus indicus Holthuis, 1947a: 40; Figs 1-3. [Bali Sea, 728.2'S 11524.6'E, 1018 m]
Lebbeus kuboi Hayashi, 1992: 123; Figs 6-8. [central Sea of Japan, of Namerikawa, Toyama Prefecture, 200
m]
Lebbeus laevirostris Crosnier, 1999a: 454; Figs 1-3. [Indonsie (dtroit de Makassar), 0237.6'S 11810.9'E,
798-779 m]
Lebbeus lagunae (Schmit, 1921)
= Spirontocaris lagunae Schmit, 1921: 57; Fig. 35; Plate 12, Figs 10-11. [Laguna Beach, California, 12-15
fms]
Lebbeus laurentae Wicksten, 2010: 196; Figs 1-4; nomen novum for Lebbeus carinatus de Saint Laurent, 1984
nec Zarenkov, 1976. [Northeastern Pacifc, 1249'N 10357'W, 2630 m]
Lebbeus longidactylus (Kobjakova, 1936a)
= Hetairus longidactyla Kobjakova, 1936a: 222; Figs 12-13. [Okhotsk Sea, 440-504 m; fully described in
Kobjakova, 1937]
Lebbeus longipes (Kobjakova, 1936a)
= Hetairus longipes Kobjakova, 1936a: 222; Fig. 16. [Peter the Great Bay and Tatar Strait, Sea of Japan;
fully described in Kobjakova, 1937]
Lebbeus manus Komai & Collins, 2009: 29; Figs 1-3. [Manus Basin, 0379'S 14609'E, 1575 m]
Lebbeus microceros (Kryer, 1841)
= Hippolyte microceras Kryer, 1841: 578. [Grnland; microceras is an incorrect original spelling under
Art. 33.3.1 and corrected to microceros]
= Spirontocaris zebra Leim, 1921: 133; Plates 2-3. [Passamquoddy bay, 30 m; Joes Point, St. Croix
River; Head Harbour, Campobello Island, low tide and 10 m (all New Brunswick); St. Marys Bay
(Nova Scotia)]
Lebbeus miyakei Hayashi, 1992: 127; Figs 10-11. [Sea of Genkainada, Orono-shima Island, Fukuoka Pre-
fecture, 30-40 m]
Lebbeus mundus Jensen, 2006: 30; Figs 4C, 5-6. [Willis Point, Vancouver Island, British Columbia, Canada,
48.34.601N 123.29.319W, 9-12 m]
Lebbeus nudirostris Komai & Takeda, 2004: 78; Figs 1A, 2-3. [west of Taibusa-misaki, Boso Peninsula,
3504.68'N 13945.32'E, 250 m, trap for scampi]
Lebbeus polaris (Sabine, 1824)
= Alpheus polaris Sabine, 1824: ccxxxviii; Plate 2, fgs 5-8. [on the coast of Melville Island, 50 fms]
= Hippolite borealis Ross, 1835: lxxxiv; Plate B, fg. 3. [of Elizabeth Harbour, 80 fms]
= Lebbeus orthorhynchus White, 1847a: 76. [Type locality not indicated]
= Hippolyte St. Pauli Brandt, 1851: 118; Plate 5, fg. 19. [Beeringschen Meere bei Pauls-Insel]
= Hippolyte cultellata Norman, 1867: 200. [the Minch]
= Hippolyte incerta Buchholz, 1874: 272. [Ostgrnland]
= Hippolyte Amazo Pfefer, 1886: 46; Plate 1, fgs 7a-b. [Cumberland-Sund, Bafns Land, 6635'40''N
6719'15''W]
= Hetarius debilis Spence Bate, 1888: 615; Plate 109, fg. 4. [Challenger stn 49, 433'N 6339'W, 85 fms]
= Hetairus tenuis Spence Bate, 1888: 613; Plate 109, fg. 3. [Challenger stn 49, 433'N 6339'W, south of
Halifax, Nova Scotia, 85 fms]
= Hippolyte projecta Spence Bate, 1888: 594; Plate 105, fg. 3. [Challenger stn 49, 433'N 6339'W, south
of Halifax, Nova Scotia, 85 fms]
= Hippolyte mysis Birulya, 1898: 184; Plate 1, fgs 1-15. [White Sea, SW coast of Kandalak Bay, Kovda]
Lebbeus polyacanthus Komai, Hayashi & Kohtsuka, 2004: 109; Figs 4-6. [of Togi, Hakui District, west
coast of Noto Peninsula, 400 m]
Lebbeus profundus (Rathbun, 1906)
= Spirontocaris profunda Rathbun, 1906: 914; Plate 24, fg. 10. [vicinity of Modu Manu, 762-1000 fms]
Lebbeus rubrodentatus Bruce, 2010k: 75; Figs 1-2. [935'S 12928'E, Timor Sea, NW Australia, 360-396 m]
Lebbeus saldanhae (Barnard, 1947)
= Spirontocaris saldanhae Barnard, 1947: 385. [of Saldanha Bay, 145 fms]
Lebbeus schrencki (Branikov, 1907)
= Hetairus schrenki Branikov, 1907: 161; Fig. 21. [East coast of Sakhalin Island, Sea of Okhotsk, 43-100 m]
Lebbeus scrippsi Wicksten & Mndez G., 1982: 106; Plates 1-2. [Of Arica, Chile, 1840.5'S 7036.0'W to
1832.2'S 7029.8'W, 768-968 m]
Lebbeus similior Komai & Komatsu, 2009: 540; Figs 10-12. [of Soh-ma, Fukushima Prefecture, 3747.6'N
14237.1'E to 3747.4'N 14237.2'E, 1196-1196 m]
Lebbeus speciosus (Urita, 1942)
= Spirontocaris makarof speciosa Urita, 1942: 19; Fig. 4. [Otomari, 2-3 fms]
= Lebbeus possjeticus Kobjakova, 1967: 235; Fig. 4. [Minosok Bay, Possjet Gulf, Sea of Japan, Russian
Far East]
Lebbeus spinirostris (Kobjakova, 1936a)
= Hetairus spinirostris Kobjakova, 1936a: 222; Fig. 10. [Okhotsk Sea; fully described in Kobjakova,
1937]
Lebbeus splendidus Wicksten & Mndez G., 1982: 110; Plates 3-5. [Southwest of Lobos de Tierra, Peru,
631'S 8101'W, 712-744 m]
Lebbeus spongiaris Komai, 2001a: 57; Figs 1-4. [W of Izu-Oshima Island, Izu Islands, 3444.538'N
13919.723'E, 257-263 m]
Lebbeus tosaensis Hanamura & Abe, 2003: 17; Figs 1-5. [3317'34''8N 13413'07''E, 344 m]
Lebbeus unalaskensis (Rathbun, 1902a)
= Spirontocaris unalaskensis Rathbun, 1902a: 895. [north of Unalaska, 350 fms]
= Hetairus brevipes Kobjakova, 1936a: 222; Figs 9a-c. [Okhotsk Sea; fully described in Kobjakova,
1937]
= Hetairus unalaskensis var. japonica Kobjakova, 1936a: 222; Fig. 14. [Tatar Straits, Sea of Japan; fully
described in Kobjakova, 1937]
= Hetairus unalaskensis var. ochotensis Kobjakova, 1936a: 222; Fig. 15. [Okhotsk Sea; fully described in
Kobjakova, 1937]
Lebbeus unguiculatus Chang, Komai & Chan, 2010: 737; Figs 7-9, 10D. [2209.556'N 12107.174'E, 1257-
1262 m]
Lebbeus ushakovi (Kobjakova, 1936a)
= Hetairus ushakovi Kobjakova, 1936a: 222; Fig. 11. [Okhotsk Sea, 165 m; fully described in Kobjako-
va, 1937]
Lebbeus vicinus vicinus (Rathbun, 1902a)
= Spirontocaris vicina Rathbun, 1902a: 895. [north of Unalaska Island, 309 fms]
Lebbeus vicinus montereyensis Wicksten & Mndez G., 1982: 114; Plate 6. [West of Punta Banda, Baja Cali-
fornia, Mexico, 3118'N 11736'W, 2068-2086 m]
Lebbeus vinogradowi Zarenkov, 1960: 346; Fig. 3. [Sea of Okhotsk, 5657.5'N 14557'E, 204 m]
Lebbeus washingtonianus (Rathbun, 1902a)
= Spirontocaris washingtoniana Rathbun, 1902a: 895. [of Sea Lion Rock, Washington, 685 fms]
Lebbeus wera Ahyong, 2009: 786; Figs 5-7. [Brothers Caldera, 3452.93'S 17904.13'E, 1208-1209 m]
Lebbeus yaldwyni Kensley, Tranter & Grifn, 1987: 304; Figs 15-17; frontispiece (B). [New South Wales,
east of Sydney, 3343'S 15151 53'E, 450 m]
Leontocaris Stebbing, 1905
= Leontocaris Stebbing, 1905 (type species Leontocaris paulsoni Stebbing, 1905, by monotypy, gender
= Problemacaris Stebbing, 1921a (type species Problemacaris spinetum Stebbing, 1921a (junior subjec-
tive synonym of Leontocaris paulsoni Stebbing, 1905), by monotypy, gender feminine)
Leontocaris alexander Poore, 2009: 958; Figs 2-4. [Seamount Hill U, 14710.8'S 4419.6'E, 1200-1300 m]
Leontocaris amplectipes Bruce, 1990i: 121; Figs 1-5. [Victoria, South of Point Hicks, 3821.9'S 14920.0'E,
1000 m]
Leontocaris bulga Taylor & Poore, 1998: 59; Figs 2-4. [Tasmania, 94.5 km SSE of Southeast Cape, V sea-
mount, 4424.0'S 14709.0'E, 1400-1650 m]
Leontocaris lar Kemp, 1906b: 299. [outside the Porcupine Bank, 5307'N 1450'W, 500 fms]
? = Problemacaris boschmai Gordon, 1964: 337; Figs 3-6, 9B. [4803'N 904'W, 500-0 m]
Leontocaris pacifca Zarenkov, 1976
= Leontocaris pacifcus Zarenkov, 1976: 10; Fig. 3. [of Chile, 680-700 m]
Leontocaris paulsoni Stebbing, 1905: 99; Plate 26. [25 miles of Lions Head, N. 67E, 131-136 fms]
? = Problemacaris spinetum Stebbing, 1921a: 626. [about 300 fms, Table Mountain E. by S. 40 miles
according to Stebbing, 1924]
Leontocaris vanderlandi Fransen, 2001: 57; Figs 1-26. [Seychelles, N of Plate Island atoll, 0548'S 5522'E,
600 m]
Leontocaris yarramundi Taylor & Poore, 1998: 64; Figs 5-7. [Tasmania, 82.8 km SSE of Southeast Cape, U
seamount, 4419.2'S 14707.2'E, 1083-1448 m]
Ligur Sarato, 1885
= Lybia Risso, 1844 (type species P.[alaemon] Ensiferus Risso, 1816, by monotypy, gender feminine;
invalid junior homonym of Lybia H. Milne Edwards, 1834 (Crustacea Brachyura))
= Ligur Sarato, 1885 (type species Ligur Edwardsii Sarato, 1885 (junior subjective synonym of
P.[alaemon] Ensiferus Risso, 1816), by monotypy, gender masculine; name placed on the Ofcial List
of Generic Names in Zoology in Opinion 470 in 1957)
Ligur ensiferus (Risso, 1816)
= P.[alaemon] Ensiferus Risso, 1816: 106. [dans la rgion de coraux, environs de Nice]
= Hippolyte Carneus Roux, 1831: 28. [nomen nudum]
= Hippolytus Incarnatus Hope, 1851: 18. [nomen nudum]
= Palemon Vedianti Monod, 1931: 133. [nomen nudum]
= Ligur Edwardsii Sarato, 1885: 2. [mers de Saint-Jean et dEza]
Lysmata Risso, 1816
= Aglaope Rafnesque, 1814 (type species Aglaope striata Rafnesque, 1814 (invalid senior subjective
synonym of M.[elicerta] Seti Caudata Risso, 1816), by monotypy, gender feminine; invalid junior
homonym of Aglaope Latreille, 1809 (Lepidoptera); name placed on the Ofcial Index of Rejected
and Invalid Names in Zoology in Opinion 522 in 1958)
= Niphea Rafnesque, 1815 (nomen novum for Aglaope Rafnesque, 1814, gender feminine; name sup-
pressed for the purposes of the Principle of Priority but not for those of the Principle of Homonymy
under the plenary powers of the ICZN, placed on the Ofcial Index of Rejected and Invalid Names
= Melicerta Risso, 1816 (type species M.[elicerta] Seti Caudata Risso, 1816, designated by H. Milne
Edwards, 1837 [in H. Milne Edwards, 1836-1844], gender feminine; invalid junior homonym of
Melicerta Schrank, 1803 (Rotifera) and Melicerta Peron & Leseur, 1810 (Colenterata); name placed on
the Ofcial Index of Rejected and Invalid Names in Zoology in Opinion 522 in 1958)
= Lysmata Risso, 1816 (nomen novum for Melicerta Risso, 1816, gender feminine; name placed on the
? = Opithiocheirus Leach, 1830a (type species Opithiocheirus Chrysophthalmus Leach, 1830a, by mono-
? = Usterocheirus Leach, 1830a (type species Usterocheirus Macropocoilium Leach, 1830a, designated
by Holthuis, 1993a, gender masculine)
= Arno Roux, 1831 (nomen novum for Aglaope Rafnesque, 1814, gender feminine)
= Eretmocaris Spence Bate, 1888 (type species Eretmocaris stylorostris Spence Bate, 1888, designated
by Holthuis, 1955b, gender feminine)
= Hippolysmata Stimpson, 1860a (type species Hippolysmata vitata Stimpson, 1860a, gender femi-
Lysmata amboinensis (De Man, 1888b)
= Hippolysmata vitata var. amboinensis De Man, 1888b: 495. [Amboina]
Lysmata anchisteus Chace, 1972: 125; Figs 53-54. [Grenada, Point Saline, rocks at southwest end of frst
beach on lee coast]
Lysmata ankeri Rhyne & Lin, 2006: 179; Figs 7-9; Plate 1C. [Pompano Beach, Florida, southeast of inlet, 4 m]
Lysmata argentopunctata Wicksten, 2000a: 3; Figs 1-3. [Morro Colorado, Sonora, 2820'N 11118'W, 5-10 m]
Lysmata bahia Rhyne & Lin, 2006: 191; Figs 16-18; Plate 1F. [Bahia, Salvador, Brazil, seawall at harbor,
purchased from a pet store]
Lysmata boggessi Rhyne & Lin, 2006: 183; Figs 10-12; Plate 1D. [northwest of Hernando Beach, Florida,
seagrass bed, 4 m]
Lysmata californica (Stimpson, 1866)
= Hippolysmata californica Stimpson, 1866: 48. [San Diego, Cal.]
= Hippolyte lineata Lockington, 1877a: 35. [San Diego, between tides]
Lysmata chica Wicksten, 2000a: 8; Figs 5-7. [Baha Cartago, Isla Isabela (Albemarle Island), Galapagos,
034'N 9058'W, shore]
Lysmata debelius Bruce, 1983e: 115; Figs 1-9. [Polillo Island, east of Luzon, Philippines, 28 m]
Lysmata dispar Hayashi, 2007: 85; Figs 1-3. [Goodwyn I., 2032.000'S 11632.420'E, intertidal]
Lysmata galapagensis Schmit, 1924c: 165; Fig. 41. [Northeast of Eden, 7 fms]
Lysmata grabhami (Gordon, 1935)
= Hippolysmata grabhami Gordon, 1935: 319; Figs 10, 11a-b. [Funchal, Madeira]
Lysmata gracilirostris Wicksten, 2000b: 208; Figs 1-3. [Golfo de Papagayo, Costa Rica, 1042.4'N 8555'W,
8-86 m]
Lysmata guamensis Anker & Cox, 2011: 205; Figs 1-3. [Mariana Archipelago, Guam, Glass Breakwater
near mouth of Apra Harbor, 3-6 m]
Lysmata hochi Baeza & Anker, 2008: 149; Figs 1-2. [Panama, Caribbean coast, Bocas del Toro, Isla Colon,
Playa Bluf, 0.5-1 m]
Lysmata holthuisi Anker, Baeza & De Grave, 2009: 684; Figs 1-5. [Taboga I., intertidal, low tide, under
rocks and coral rubble]
Lysmata intermedia (Kingsley, 1878a)
= Hippolysmata intermedia Kingsley, 1878a: 90. [Fort Jeferson; Tortugas, Florida]
Lysmata kempi Chace, 1997: 53. [nomen novum for Hippolysmata dentata Kemp, 1914 nec Palaemon denta-
tus De Haan, 1844]
= Hippolysmata dentata Kemp, 1914: 117; Plate 6, fg. 5. [of M. of Irrawaddy R, 1520'N 9455'E, 20
fms]
Lysmata kuekenthali (De Man, 1902)
= Hippolyte kkenthali De Man, 1902: 850; Plate 26, fg. 56; nomen novum for Merhippolyte orientalis
sensu De Man, 1892a nec Spence Bate, 1888. [Flores; Ternate]
= Hippolysmata marleyi Stebbing, 1919: 120; Plate 18. [Isezela, Natal]
Lysmata lipkei Okuno & Fiedler, 2010: 599; Figs 1-4. [Ubara-jima Islet, Katsuura, Boso Peninsula, Honshu,
Japan, 3507.7'N 14016.7'E, 15 m]
Lysmata moorei (Rathbun, 1901)
= Hippolysmata moorei Rathbun, 1901: 115; Fig. 23. [Playa de Ponce]
Lysmata morelandi (Yaldwyn, 1971)
= Hippolysmata (Hippolysmata) morelandi Yaldwyn, 1971: 90. [Bay of Islands, North Auckland, subtid-
al algal beds on rocky substrate to a depth of about 20 f]
Lysmata multiscissa (Nobili, 1904)
= Hippolysmata multiscissa Nobili, 1904: 232. [Djibouti]
Lysmata nayaritensis Wicksten, 2000a: 14; Figs 9-11. [Playa Mantac, about 5 miles SE of San Blas, Nayarit,
Mexico, 2133'N 10519'W]
Lysmata nilita Dohrn & Holthuis, 1950: 339; Fig. 1; Plate 9. [Western half of Bay of Naples]
= Lysmata Nilita Hope, 1851: 17. [nomen nudum]
Lysmata olavoi Fransen, 1991b: 63; Figs 1-34. [Azores, Pico, Ponto da Ilha, 3825'00"N 2759'10"W, 135
m]
Lysmata pederseni Rhyne & Lin, 2006: 187; Figs 13-15, Plate 1E. [southeast of Sombrero Reef, Marathon,
Florida, in tube sponge, 15 m]
Lysmata philippinensis Chace, 1997: 53, 75; Figs 19-20. [Albay Gulf, east of southern Luzon, 1312'N
12349'18"E, 267 m]
Lysmata porteri (Rathbun, 1907)
= Hippolysmata porteri Rathbun, 1907: 49; Plate 3, fg. 4. [Bay of Valparaiso]
Lysmata rafa Rhyne & Anker, 2007: 292; Figs 2-4. [Florida, Key west Lakes, west of Key West, under
ledge, about 1 m]
Lysmata rathbunae Chace, 1970: 59; Figs 1-3. [Of Boynton Beach, Florida, 2631'N 8001'W, 55-64 m]
Lysmata rauli Laubenheimer & Rhyne, 2010: 299; Figs 1-3. [Salvador, Bahia, Brazil, under rocky ledge in
tide pool, 2 m depth]
Lysmata seticaudata (Risso, 1816)
= M.[elicerta] Seti Caudata Risso, 1816: 110; Plate 2, fg. 1. [environs de Nice, dans les eaux profondes]
= Aglaope striata Rafnesque, 1814: 24. [Name suppressed under the plenary powers for the pur-
poses of the Principle of Priority but not for those of the Principle of Homonymy in Opinion 522 in
1958]
= P.[alemon] Cognetii Risso, 1816: 106. [environs de Nice, dans la rgion des madrpores]
= Lysmata aberrans Czerniavsky, 1884: 63; Plate 3, Figs 7A-K. [Portus Suchum, 1-1.5 m]
= Miersia clavigera Chun, 1888: 34; Plate 4, fg. 6. [Type locality not indicated, presumed to be around
Gulf of Naples, 0-600 m]
Lysmata splendida Burukovsky, 2000g: 223; Figs 1-3. [Maldives, Ari-Atoll, inner reef Maayafushi, in
caverns on reef roof wall, 6-35 m]
Lysmata stenolepis Crosnier & Forest, 1973: 177; Figs 55, 56a-c. [Of Sao Tiago, 275-150 m]
Lysmata ternatensis De Man, 1902
= Palaemon dentatus De Haan, 1844 [in De Haan, 1833-1850]: Plate 45, fg. 13 (1844); 176 (1849, as
Lysmata seticaudata Risso). [Japan] [nec Palaemon dentatus Roemer, 1841]
= Lysmata seticaudata var. ternatensis De Man, 1902: 846. [Ternate]
= Hippolysmata acicula Rathbun, 1906: 912; Plate 24, fg. 6. [vicinity of Kauai Island, 7-148 fms]
= Lysmata afnis Borradaile, 1915b: 209. [Minikoi, Peros Banhos, Salomon, Seychelles]
Lysmata trisetacea (Heller, 1861)
= H.[ippolyte] trisetacea Heller, 1861: 29. [im rothen Meere]
= Lysmata pusilla Heller, 1862c: 287; Plate 3, fg. 26. [im rothen Meere]
= Lysmata chiltoni Kemp, 1914: 110; Plate 6, fgs 1-4. [Meyer Island, Kermadec group]
= Hippolysmata paucidens Rathbun, 1906: 913; Plate 24, fg. 4. [Waikiki beach]
Lysmata udoi Baeza, Bolaos, Hernandez & Lpez, 2009: 62; Figs 1-3. [Caribbean Coast, Cubagua Island,
Venezuela, from dens of the sapo bocn toadfsh Amphichthys criptocentrus, 2-4 m]
Lysmata uncicornis Holthuis & Maurin, 1952: 198; Figs 1-2. [port de Casablanca, 4-5 m]
Lysmata vitata (Stimpson, 1860a)
= Hippolysmata vitata Stimpson, 1860a: 26. [in portu Hong Kong]
= Nauticaris unirecedens Spence Bate, 1888: 608; Plate 110, fg. 1. [Hong Kong]
= Hippolysmata vitata var. subtilis Thallwitz, 1892: 22. [Ceb]
= Hippolysmata durbanensis Stebbing, 1921b: 20; Plate 5. [cast up on the beach in Durban Bay]
Lysmata wurdemanni (Gibbes, 1850)
= Hippolyte Wurdemanni Gibbes, 1850: 197. [Key West Lakes, west of Key West, Florida, 1 m; neotype
selection by Rhyne & Lin, 2006]
Lysmata zacae Armstrong, 1941: 10; Fig. 4. [Mataatu Harbor, Savaii, 8f]
Lysmatella Borradaile, 1915b
= Lysmatella Borradaile, 1915b (type species Lysmatella prima Borradaile, 1915b, by monotypy, gen-
der feminine)
Lysmatella prima Borradaile, 1915b: 209. [Maldive Islands]

Merguia Kemp, 1914
= Merguia Kemp, 1914 (type species Hippolyte oligodon De Man, 1888a, by monotypy, gender femi-
Merguia oligodon (De Man, 1888a)
= Hippolyte oligodon De Man, 1888a: 277; Plate 18, fgs 1-6. [Elphinstone Island]
Merguia rhizophorae (Rathbun, 1900)
= Hippolysmata rhizophor Rathbun, 1900: 153; Plate 8, fg. 9. [Rio Parahyba do Norte, on mangroves]
Merhippolyte Spence Bate, 1888
= Merhippolyte Spence Bate, 1888 (type species Merhippolyte agulhasensis Spence Bate, 1888, by origi-
nal designation, gender feminine; name placed on the Ofcial List of Generic Names in Zoology in
Merhippolyte agulhasensis Spence Bate, 1888: 619; Plate 110, fg. 4. [Challenger stn 142, 3504'S 1837'E,
south of the Cape of Good Hope, 150 fms]
Merhippolyte americana Holthuis, 1961d: 1; Fig. 1. [Yucatan Channel, 2059'30"N 8623'45"W, 130 fms]
Merhippolyte ancistrota Crosnier & Forest, 1973: 167; Figs 52-53. [les du Cap Vert, 1534.5'N 2311.5'W,
185 m]
Merhippolyte calmani Kemp & Sewell, 1912: 20; Plate 1, fgs 1-4. [of the S.W. coast of India, 914'10"N
7545'E, 237 fms]
Merhippolyte chacei Kensley, Tranter & Grifn, 1987: 309; Figs 18-19. [New South Wales, 3343-37'S 15155'
15202'E, 686 m]
Merhippolyte kauaiensis (Rathbun, 1906)
= Spirontocaris kauaiensis Rathbun, 1906: 913; Plate 24, fg. 5. [vicinity of Kauai Island, 55-362 fms]
Merhippolyte peroni Ledoyer, 1990: 48; Fig. 3A. [Sud de lle Saint-Paul]
Mimocaris Nobili, 1903b
= Mimocaris Nobili, 1903b (type species Mimocaris heterocarpoides Nobili, 1903b, by monotypy, gen-
Mimocaris heterocarpoides Nobili, 1903b: 6; Fig. 2. [Pulu Burong]
Nauticaris Spence Bate, 1888
= Nauticaris Spence Bate, 1888 (type species Nauticaris marionis Spence Bate, 1888, designated by
Calman, 1906b, gender feminine; name placed on the Ofcial List of Generic Names in Zoology in
Nauticaris brucei Stebbing, 1914a: 292; Plate 24. [Scotia stn 461, Gough Island, 100 fms]
Nauticaris magellanica (A. Milne-Edwards, 1891) (Fig. 50)
= Hippolyte magellanicus A. Milne-Edwards, 1891: 46; Plate 5, fgs 2-2i. [dans la baie Orange, 17 m; au
nord-ouest de lle Grvy (cap Hall), 65 m]
= Hippolyte consobrinus A. Milne-Edwards, 1891: 47; Plate 5, fg. 4. [ terre dans la baie Orange]
= Nauticaris marionis var. chilensis Dofein & Balss, 1912: 30. [Port Stanley, Falkland-Inseln; Magal-
haens-Strae]
Nauticaris marionis Spence Bate, 1888: 603; Plate 108. [Challenger stns 144A, 4648'0''S 3749'30''E, of
Marion Island, 69 fms; 145, 4643'0''S 384'30''E, of Prince Edward Island, 140 fms; 315, 5140'S
5750'W, of the Falkland Islands, 12 fms]
= Hippolyte stewarti Thomson, 1889: 259; Plate 13, fg. 1. [Paterson Inlet, Stewart Island]
= Merhippolyte australis Hodgson, 1902: 233; Plate 29 (as Hippolyte australis). [Auckland Island, 10
fms]
Paralatreutes Kemp, 1925
= Paralatreutes Kemp, 1925 (type species Paralatreutes bicornis Kemp, 1925, by original designation
and monotypy, gender masculine; name placed on the Ofcial List of Generic Names in Zoology in
Paralatreutes bicornis Kemp, 1925: 334; Figs 23-24. [Ross Channel, Port Blair, Andamans, 3-4 fms]
Paralebbeus Bruce & Chace, 1986
= Paralebbeus Bruce & Chace, 1986 (type species Paralebbeus zotheculatus Bruce & Chace, 1986, by
Paralebbeus zotheculatus Bruce & Chace, 1986: 238; Figs 1-6. [1730.1'S 11828.9'E, 505-506 m]
Paralebbeus zygius Chace, 1997: 82; Figs 25-28. [Selat Butung, Sulawesi (Celebes), Indonesia, 535'00"S
12220'00''E, 1023 m]
Fig. 50. Nauticaris magellanica (A. Milne-Edwards, 1891). Photo by Arthur Anker.
Phycocaris Kemp, 1916a
= Phycocaris Kemp, 1916a (type species Phycocaris simulans Kemp, 1916a, by original designation
and monotypy, gender feminine; name placed on the Ofcial List of Generic Names in Zoology in
Phycocaris simulans Kemp, 1916a: 392; Fig. 2; Plate 36, fg. 2. [near Ross Island, Port Blair, Andaman Is-
lands, 2-4 fms]
Saron Thallwitz, 1891
= Saron Thallwitz, 1891 (type species Hippolyte gibberosus H. Milne Edwards, 1837 [in H. Milne Ed-
wards, 1834-1840], by original designation and monotypy, gender masculine; name placed on the
Saron inermis Hayashi in Debelius, 1983: 117 (partim, see Hayashi, 1989). [Indonesien]
Saron marmoratus (Olivier, 1811)
= Palmon marmoratus Olivier, 1811: 665. [Nouvelle-Hollande]
= Hippolyte gibberosus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 378. [les ctes de
la Nouvelle-Hollande]
= Hippolyte Leachii Gurin-Mneville, 1838 [in Gurin-Mneville, 1829-1838]: 37. [ctes de lle de
Strong ou Oualan]
= H.[ippolyte] Hemprichii Heller, 1861: 29. [im rothen Meere]
= Hyppolite Kraussii Bianconi, 1869: 209; Plate 1, fg. 2a. [Mosambico]
= Nauticaris grandirostris Pearson, 1905: 79; Plate 1, fgs 6-6c. [entrance to Galle Harbour, 4.5-7 fms]
Saron neglectus De Man, 1902: 854; Plate 26, fgs 58-58b. [Ternate; Pulo Edam in der Bai von Batavia;
Kagoshima, Japan]
Saron rectirostris Hayashi, 1984: 116. [Indonesia]
Spirontocaris Spence Bate, 1888
= Sowerbyus Hoek, 1887 (type species Sowerbyus spinus Hoek, 1887, probably a later combination of
Cancer Spinus Sowerby, 1805 [in Sowerby, 1804-1806], by monotypy, gender masculine; name place
= Spirontocaris Spence Bate, 1888 (type species Cancer Spinus Sowerby, 1805 [in Sowerby, 1804-1806],
by monotypy, gender feminine; name placed on the Ofcial List of Generic Names in Zoology in
Spirontocaris arcuata Rathbun, 1902a: 893. [Washington Sound, Straits of Fuca, 48 fms]
Spirontocaris arcuatoides Kobjakova, 1962: 244; Fig. 2. [Ostrov Zelenii (Green Island), 14 m; Shitokan Is-
land, 10-4 m]
Spirontocaris brashnikowi Kobjakova, 1936a: 221. [Tatar Strait and Peter the Great Bay, Sea of Japan; no-
men novum for Spirontocaris dalli sensu Branikov, 1907 nec Rathbun, 1902a; fully described in
Kobjakova, 1937]
= Spirontocaris brashnikovi Makarov, 1941: 121. [Avachinsk Bay; nomen nudum]
Spirontocaris brevidigitata Kobjakova, 1935: 88; Fig. 3. [von der Bucht Peters des Groen nach Norden bis
zur Bucht Nelma, 75-1380 m]
Spirontocaris dalli Rathbun, 1902a: 894. [Coal Harbor, Unga Island, Alaska, 9-8 fms]
Spirontocaris gurjanovae Kobjakova, 1955: 238; Fig. 3. [4th Kuril Straits, Kurile Islands, 100 m]
Spirontocaris holmesi Holthuis, 1947a: 8. [nomen novum for Spirontocaris bispinosus Holmes, 1900 nec
Hippolyte bispinosa De Haan, 1844 [in De Haan, 1833-1850]]
= Spirontocaris bispinosus Holmes, 1900: 207. [Puget Sound]
Spirontocaris intermedia Kobjakova, 1936a
= Spirontocaris spina intermedia Kobjakova, 1936a: 221. [Okhotsk Sea; fully described in Kobjakova,
1937]
= Spirontocaris spina intermedia Makarov, 1941: 117; Figs 1-2. [Bering Sea and Chukchi Sea, 30-100 m]
Spirontocaris lamellicornis (Dana, 1852a)
= Hippolyte lamellicornis Dana, 1852a: 24. [in freta de Fuca Oregoni, juxta portum Dungeness]
Spirontocaris lilljeborgii (Danielssen, 1859)
? = Astacus histrio Fabricius, 1775: 415. [Groenlandia]
= H.[ippolyte] Lilljeborgii Danielssen, 1859: 5. [Slotholem (Lofoten) paa sandig Leerbund, 40 fms]
= Hippolyte securifrons Norman, 1862: 151. [of the Shetland Isles; full description in Norman, 1863]
Spirontocaris microdentata Kobjakova, 1962
= Spirontocaria [sic] microdentatus Kobjakova, 1962: 245; Fig. 4. [Ushishir Island, about 18 m]
Spirontocaris murdochi Rathbun, 1902a: 893. [of Robben Island, east coast of Sghalin, Okhostk Sea]
Spirontocaris ochotensis (Brandt, 1851)
= Hippolyte ochotensis Brandt, 1851: 120; Plate 5, Figs 17-17b. [Type locality not indicated]
= Spirontocaris mororani Rathbun, 1902b: 43; Fig. 16. [Mororan, Hokkaido]
= Spirontocaris makarovi Kobjakova, 1936a: 221. [Okhotsk Sea; fully described in Kobjakova, 1937]
= Spirontocaris makarovi spatula Kobjakova, 1936a: 221. [Okhotsk Sea; fully described in Kobjakova,
1937]
= Spirontocaris onagawaensis Yokoya, 1939: 268; Fig. 5. [Takasiro, 7.5 m; Nonohama, 4.6 m]
Spirontocaris pectinifera (Stimpson, 1860a)
= Hippolyte pectinifera Stimpson, 1860a: 35. [sinu Hakodadi f. conchoso org 8.]
= Spirontocaris crassirostris Kubo, 1951: 274; Figs 11-12. [of Heta, Izu peninsula, ca. 300 m]
Spirontocaris phippsii (Kryer, 1841)
= Hippolyte Phippsii Kryer, 1841: 575. [Spitsbergen, Norges Vestkyst (og Grnland ?)]
= Hippolyte turgida Kryer, 1841: 575. [Spitsbergen, Norges Vestkyst, Grnland]
= Hippolyte vibrans Stimpson, 1871: 125. [Massachusets Bay]
Spirontocaris prionota (Stimpson, 1864)
= Hippolyte prionota Stimpson, 1864: 153. [Hales Passage, 10 fms; of Lummi Island, 8-12 fms, near
San Juan Island, 2-4 fms (all Puget Sound)]
= Spirontocaris macrodonta J.F.L. Hart, 1930: 102; Plate 1. [Gonzales Point, tide pool; False Narrows,
near the waters edge; Departure Bay in about 10 fms (all Vancouver Island)]
Spirontocaris sica Rathbun, 1902a: 894. [Santa Barbara Channel, 265 fms]
Spirontocaris snyderi Rathbun, 1902a: 894. [Monterey Bay, California]
Spirontocaris spinus (Sowerby, 1805 [in Sowerby, 1804-1806])
? = Astacus histrio Fabricius, 1775: 415. [Groenlandia]
= Cancer spinus Sowerby, 1805 [in Sowerby, 1804-1806]: 47; Plate 23. [among oysters on the Scotish
coast]
= Hippolyte Sowerbi Leach, 1817 [in Leach, 1815-1875]: Plate 39, fgs 1-10. [Newhaven, in the Frith
of Forth]
= Spirontocaris spina laevidens Kobjakova, 1936a: 221. [Tatar Strait and Peter the Great Bay, Sea of Ja-
pan; fully described in Kobjakova, 1937]
Spirontocaris truncata Rathbun, 1902a: 894. [Heceta Bank, Oregon, 50 fms]
Spirontocaris urupensis Kobjakova, 1962: 243; Fig. 1. [Urup Island, 5-7 m]
Thinora Bruce, 1998d
= Thinora Bruce, 1998d (nomen novum for Thorina Bruce, 1997b, gender feminine)
= Thorina Bruce, 1997b (type species Thor maldivensis Borradaile, 1915b, by original designation
and monotypy, gender feminine; invalid junior homonym of Thorina Stephensen, 1944 (Amphi-
poda))
Thinora maldivensis (Borradaile, 1915b)
= Thor maldivensis Borradaile, 1915b: 208. [Minikoi, Maldives; Salomon]
Thor Kingsley, 1878a
= Thor Kingsley, 1878a (type species Thor foridanus Kingsley, 1878a, by monotypy, gender mascu-
line; name placed on the Ofcial List of Generic Names in Zoology in Opinion 470 in 1957)
= Paschocaris Nobili, 1905b (type species H.[ippolyte] paschalis Heller, 1862c, by original designation
Thor algicola Wicksten, 1987: 27; Figs 1-3. [Baha Bacochibampo, Guaymas, Sonora, Mexico, 2757'N
11102'W, 5 m, among Sargassum]
Thor amboinensis (De Man, 1888b)
= Hippolyte amboinensis De Man, 1888b: 535. [Amboina]
= Thor discosomatis Kemp, 1916a: 388; Fig. 1; Plate 36, fg. 1. [Port Blair, Andamans]
Thor cocoensis Wicksten & Vargas, 2001: 139; Figs 1-4. [Bajo Alcyone, Isla del Coco, Costa Rica (Pacifc),
35 m]
Thor cordelli Wicksten, 1996b: 295; Fig. 1. [Sulphur Bay, Clarion Island, Mexico, shallow]
Thor dobkini Chace, 1972: 133; Fig. 57. [Florida, Punta Rassa, 2 m]
Thor foridanus Kingsley, 1878a: 95. [Key West, Florida]
Thor intermedius Holthuis, 1947a: 51; Figs 4-6. [Sissie near Misool, shore and reef]
Thor manningi Chace, 1972: 137; Figs 59-60. [English Harbour, Antigua, from botom of yacht anchored
for several months]
Thor marguitae Bruce, 1978e: 159; Figs 1-6. [reef fat, Heron Island, Capricorn Islands, Queensland, Aus-
tralia, 2325.25'S 15155.20'E]
Thor paschalis (Heller, 1862c)
= H.[ippolyte] paschalis Heller, 1862c: 276; Plate 3, fg. 24. [im rothen Meere]
Thor spinipes Bruce, 1983f: 2; Figs 1-6. [Burford Island, Cobourg Peninsula, Northern Territory, Australia,
1129.3'S 13157.5'E, muddy reef fat pools at LWS tide]
Thor spinosus Boone, 1935: 192; Plate 52. [Tmukus Roads, Bali, Dutch East Indies, in coral]
Thorella Bruce, 1982d
= Thorella Bruce, 1982d (type species Thorella cobourgi Bruce, 1982d, by original designation and
Thorella cobourgi Bruce, 1982d: 452; Figs 1-5. [Black Point, Port Essington, Cobourg Peninsula, Northern
Territory, 1109.0'S 13208.2'E, 1-2 m, in Sargassum]
Tozeuma Stimpson, 1860a
= Tozeuma Stimpson, 1860a (type species Tozeuma lanceolatum Stimpson, 1860a, by monotypy, gen-
der neuter)
= Angasia Spence Bate, 1863 (type species Angasia pavonina Spence Bate, 1863, by monotypy, gender
feminine)
Tozeuma armatum Paulson, 1875
= Toz.[euma] armata Paulson, 1875: 99; Plate 15, fg. 2. [Red Sea]
= Angasia Stimpsonii Henderson, 1893: 437; Plate 40, fgs 18-20. [Gulf of Martaban]
Tozeuma carolinense Kingsley, 1878a (Fig. 51)
= Tozeuma carolinensis Kingsley, 1878a: 90. [Fort Macon, North Carolina]
Fig. 51. Tozeuma carolinense Kingsley, 1878. Photo by Arthur Anker.
Tozeuma cornutum A. Milne-Edwards, 1881b: 16. [prs de Barbades, 40 brasses]
Tozeuma elongata (Baker, 1904)
= Angasia elongata Baker, 1904: 147; Plate 27, fgs 1-4. [south Australian coast, 15 fms; Port Victor]
Tozeuma erythraeum Nobili, 1904
= Tozeuma erythrum Nobili, 1904: 231. [Mer Rouge]
Tozeuma kimberi (Baker, 1904)
= Angasia kimberi Baker, 1904: 149; Plate 27, fg. 5. [Port Willunga, 4 fms]
Tozeuma lanceolatum Stimpson, 1860a: 27. [in portu Hong Kong, in fundo limoso prof. sex. org. sat
vulgaris]
Tozeuma novaezealandiae Borradaile, 1916
= Tozeuma novae-zealandiae Borradaile, 1916: 86; Fig. 3. [7 miles E of North Cape, New Zealand, 128
m]
Tozeuma pavoninum (Spence Bate, 1863)
= Angasia pavonina Spence Bate, 1863: 498; Plate 90, fgs 1-1z. [St. Vincents Gulf, 4.5 fms]
= Angasia robusta Baker, 1904: 150; Plate 28, fgs 1-8. [St.Vincent Gulf, 10-12 fms]
Tozeuma serratum A. Milne-Edwards, 1881b: 16. [prs de Barbades, 56 brasses]
Tozeuma tomentosum (Baker, 1904)
= Angasia tomentosa Baker, 1904: 152; Plate 29, fgs 1-4. [south Australian coast, about 20 fms]
Trachycaris Calman, 1906b
= Trachycaris Calman, 1906b (type species Platybema rugosus Spence Bate, 1888, by original designa-
tion and monotypy, gender feminine; name placed on the Ofcial List of Generic names in Zoology
Trachycaris restricta (A. Milne-Edwards, 1878)
= Hippolyte restrictus A. Milne-Edwards, 1878: 231. [les du Cap-Vert]
Trachycaris rugosa (Spence Bate, 1888)
= Platybema rugosus Spence Bate, 1888: 579; Plate 104, fg. 2. [Challenger stn 24, 1838'30''N 655'30''W,
of Culebra island, West Indies, 390 fms]
Family OGYRIDIDAE Holthuis, 1955b
Ogyrides Stebbing, 1914b
= Ogyris Stimpson, 1860a (type species Ogyris orientalis Stimpson, 1860a, by monotypy, invalid jun-
ior homonym of Ogyris Westwood, 1851 (Lepidoptera), gender feminine; name placed on the Of-
cial Index of Rejected and Invalid Names in Zoology in Opinion 470 in 1957)
= Ogyrides Stebbing, 1914b (nomen novum for Ogyris Stimpson, 1860a, gender masculine; name
Ogyrides alphaerostris (Kingsley, 1880)
= Ogyris alphrostris Kingsley, 1880: 420; Plate 14, fg. 7. [Northampton County, Virginia, Eastern
shore, Atlantic side]
= Ogyris occidentalis Ortmann, 1893: 46; Plate 3, fg. 4. [Plankton-Expedition der Humbolt-Stifung:
Tocantins-Mndung JN 239 (0.6S 48.1W, 0 m); Pl. 106 (0.7S 48.2W, 12 m)]
= Ogyrides yaquiensis Armstrong, 1949: 3; Fig. 1. [Rio Yaqui del Sul, Dominican Republic, 1-3 f]
= Ogyrides limicola Williams, 1955: 57; Fig. 1. [mouth of Far Creek at Ebgelhard, Hyde Count, North
Carolina, 1-8 f]
Ogyrides delli Yaldwyn, 1971: 89. [Lowry Bay, Wellington Harbour, 4 fms]
Ogyrides hayi Williams, 1981: 145. [of Bogue Bank, west of Fort Macon, North Carolina, ca. 3.5 m] (Fig. 52)
Ogyrides mjoebergi (Balss, 1921a)
= Ogyris Mjbergi Balss, 1921a: 7; Figs 1-2 [Cape Jaubert, 45 meilen WSW, 54 fuss]
Ogyrides orientalis (Stimpson, 1860a)
= Ogyris orientalis Stimpson, 1860a: 36. [mari Sinensi, et in sinu Kagosima, 5-25 org]
Ogyrides rarispina Holthuis, 1951a: 119; Fig. 25. [anchorage of Monrovia, Liberia, 11 m]
Ogyrides saldanhae Barnard, 1947
= Ogyrides saldanh Barnard, 1947: 387. [Saldanha Bay, 10 fms]
Ogyrides sibogae (De Man, 1910a)
= Ogyris Sibogae De Man, 1910a: 318. [64.1'N 12044'E, 535 m]
Ogyrides striaticauda Kemp, 1915: 284; Figs 28-30. [Chilka Lake]
Ogyrides tarazonai Wicksten & Mendez G., 1988: 622; Fig. 1. [Ventanilla, Peru, 1150'S, shallow water, 50
m from tide line]
Superfamily PROCESSOIDEA Ortmann, 1896a
Family PROCESSIDAE Ortmann, 1896a
Ambidexter Manning & Chace, 1971
= Ambidexter Manning & Chace, 1971 (type species Ambidexter symmetricus Manning & Chace, 1971,
Ambidexter panamensis Abele, 1972: 373; Figs 4-5. [Panama, Canal Zone, Pacifc coast, Ft. Amador, Naos
Island, from burrows on sand-mud beach adjacent to Smithsonian Tropical Marine Laboratory]
Ambidexter swifi Abele, 1972: 366; Figs 1-3. [Panama, Panama Province, Pacifc coast, Punta Paitilla, 0.5
m]
Ambidexter symmetricus Manning & Chace, 1971: 3-7; Figs 1-2. [Florida, Dade County, Miami, Biscayne
Bay, Matheson Hammock Wading Beach, push nets on grass fats]
Clytomanningus Chace, 1997
= Clytomanningus Chace, 1997 (type species Processa molaris Chace, 1955, by original designation,
gender masculine)
Clytomanningus coutierei (Nobili, 1904)
= Processa Coutierei Nobili, 1904: 234. [Djibouti]
Clytomanningus molaris (Chace, 1955)
= Processa molaris Chace, 1955: 11; Fig. 5. [Rongelap Atoll, Burok Island, intertidal coral]
Hayashidonus Chace, 1997
= Hayashidonus Chace, 1997 (type species Nika japonica De Haan, 1844 [in De Haan, 1833-1850], by
Fig. 52. Ogyrides hayi Williams, 1981. Photo by Arthur Anker.
Hayashidonus japonicus (De Haan, 1844 [in De Haan, 1833-1850])
= Nika japonica De Haan, 1844 [in De Haan, 1833-1850]: Plate 46, fg. 6 (1844); 184, Plate N (1849).
Nikoides Paulson, 1875
= Nikoides Paulson, 1875 (type species Nik.[oides] Danae Paulson, 1875, by monotypy, gender mas-
culine; name placed on the Ofcial List of Generic Names in Zoology in Opinion 470 in 1957)
Nikoides boraboraensis Burukovsky, 2002b: 1389; unnumbered fgure. [French Polynesia, Bora-Bora Atoll,
near Motutagu, 1-1.5 m, on coral boulder]
Nikoides danae Paulson, 1875
= Nik.[oides] Danae Paulson, 1875: 98; Plate 14, Figs 5-5d.
Nikoides gurneyi Hayashi, 1975a: 58; Figs 3-4. [Kikambala, Kenya, weedy pools inner reef fat at LWS]
Nikoides longicarpus Nol, 1986: 264; Figs 1-8. [Philippines, 1359'N 12010'E, 164-150 m]
Nikoides maldivensis Borradaile, 1915b: 209. [Amirante island, 25-28 f; see Chace, 1997]
= Processa Jacobsoni De Man, 1921: 95. [Sinabang, island of Simalur, near the west coast of Atjeh,
Sumatra]
Nikoides multispinatus Hayashi, 1981b: 53; Figs 1-3. [Heron Island, Great Barrier Reef, Australia, 35f]
Nikoides plantei Burukovsky, 2007c: 1; Fig. 1. [Madagascar, Antseranana, 47 m]
Nikoides schmiti Manning & Chace, 1971: 8; Figs 3-5. [Florida, Monroe County, Tortugas, near black
buoy, 1.25 km south of Garden Key, 26 m]
Nikoides sibogae De Man, 1918a
= Nikoides Sibogae De Man, 1918a: 160. [Siboga Expedition stns 71, Makassar and surroundings, up
to 32 m; 274, 528'.2S 13453'.9E, east coast of Aru-islands, 57 m; 282, 825'.2S 12718'.4E, anchorage
between Nusa Besi and the N.E.-point of Timor, 27-54 m]
Nikoides steinii (Edmondson, 1935a)
= Processa steinii Edmondson, 1935a: 3; Fig. 1. [shoal water reef of Maui, in a coral head]
= Nikoides nanus Chace, 1955: 8; Fig. 4. [Eniwetok Atoll, Runit island, intertidal]
Processa Leach, 1815 [in Leach, 1815-1875]
= Thalassalpes Bosc, 1813 (type species N.[ika] Edulis Risso, 1816, designated by Holthuis, 1955b,
gender masculine; name suppressed for the purposes of the Principle of Priority, but not those of
the Principle of Homonymy, placed on the Ofcial Index of Rejected and Invalid Generic Names in
= Processa Leach, 1815 [in Leach, 1815-1875] (type species Processa canaliculata Leach, 1815 [in Leach,
= Nika Risso, 1816 (type species N.[ika] Edulis Risso, 1816, designated by H. Milne Edwards, 1837 [in
H. Milne Edwards, 1836-1844], gender feminine)
= Velocina Gistel, 1848 (nomen novum for Processa Leach, 1815 [in Leach, 1815-1875], gender femi-
nine)
? = Chiereghina Nardo, 1869 (type species Cancer pellucidus Nardo, 1847 (probably a junior subjective
synonym of N.[ika] Edulis Risso, 1816), gender feminine)
= Hectarthropus Spence Bate, 1888 (type species Hectarthropus expansus Spence Bate, 1888, designated
by Holthuis, 1955b, gender masculine)
Processa acutirostris Nouvel & Holthuis, 1957: 23; Figs 81-107. [prs de Monaco]
? = Cancer longipes Nardo, 1847: 6. [trovato di rado nel golfo]
Processa aequimana (Paulson, 1875)
= N.[ika] aequimana Paulson, 1875: 97; Plate 14, fgs 6-6a. [Red Sea]
Processa afnis Hayashi, 1975a: 85; Fig. 12. [Sanana Bay, east coast of Sanana, Soela Islands, 22 m]
Processa australiensis Baker, 1907: 185; Plate 25, fgs 2-2e. [south Australian coast]
Processa austroafricana Barnard, 1947: 386. [Agulhas Bank]
Processa bermudensis (Rankin, 1900)
= Nika bermudensis Rankin, 1900: 536; Plate 17, fgs 2, 2a-b. [Harrington Sound, 1fm]
Processa borboronica Holthuis, 1951a: 43; Fig. 7. [of Gold Coast, 537'N 038'E, 50 m]
Processa brasiliensis Christofersen, 1979: 364; Figs 34-35. [815'S 3442'W, of Pernambuco, 33 m]
Processa canaliculata Leach, 1815 [in Leach, 1815-1875]: Plate 41. [Torcross, southern coast of Devon]
= Nika cannelata Grifth & Pidgeon, 1833: Plate 17, fg. 2. [Type locality not indicated]
= Nika Couchii Bell, 1847 [in Bell, 1844-1853]: 278; unnumbered vignete. [coast of Cornwall]
= Nika edulis var. britannica Czerniavsky, 1884: 59. [nomen novum for Nika Couchii Bell, 1847 [in Bell,
1844-1853]]
= Nika mediterranea Parisi, 1915: 65. [Nizza]
= Processa prostatica Zariquiey Cenarro, 1941: 345; Figs 17-35. [Rgion Tramontana]
Processa compacta Crosnier, 1971: 577; Fig. 3. [Congo, baie de Pointe-Noire, 10 m]
= Processa barnardi Hayashi, 1975a: 92; Figs 15-16. [South Africa, 3350'S 2546'E, 20 fms]
Processa crosnieri Nol, 1986: 274; Figs 9-12. [Ile de la Runion, radiale B au large de St Pierre, 2120'7[S]
5526'9[E], 73-77 m]
Processa demani Hayashi, 1975a: 98; Figs 19-20. [Elat, west coast of Great Kei Islands, 27 m]
Processa dimorpha Hayashi, 1975a: 102; Fig. 21. [Japan, Sea of Genkai, of Shingu, Fukuoka Prefecture,
weed belts]
Processa edulis crassipes Nouvel & Holthuis, 1957: 16; Figs 28-37. [herbier de lle Verte, Roscof, France,
Finistre]
Processa edulis edulis (Risso, 1816)
= N.[ika] Edulis Risso, 1816: 85; Plate 3, fg. 3. [dans la rgion des algues, environs de Nice]
? = Cancer pellucidus Nardo, 1847: 6. [trovato nel golfo, tra gli Asprei]
? = Nikoides pontica Sowinsky, 1882: 226. [mer Noire]
= Nika edulis var. mediterranea forma typica Czerniavsky, 1884: 60.
= Processa edulis arcassonensis Nouvel & Holthuis, 1957: 18; Figs 38-52. [basin dArcachon]
Processa elegantula Nouvel & Holthuis, 1957: 37; Figs 174-204. [baie de Cadaqus, 30-40 m]
Processa famelica Manning & C.W.J. Hart, 1991: 317; Figs 1-3. [Hungry Bay, Bermuda, 3217'N 6446'W]
Processa fmbriata Manning & Chace, 1971: 19; Figs 8-10. [Florida, Monroe County, Tortugas, of East Key,
3 m] (Fig. 53)
Processa foresti Nol, 1986: 280; Fig.13. [Philippines, 1359'N 12018'E, 187-205 m]
Processa gracilis Baker, 1907: 187; Plate 25, Figs 3-3c. [South Australia coast]
Processa guyanae Holthuis, 1959a: 115, Figs 18-19. [NW of the Coppename River, 654'N 5614'W, 49 m]
= Processa tenuipes Manning & Chace, 1971: 31; Figs 17-18. [Gulf of Mexico, of west coast of Florida,
2912'N 8422'W, 31 m]
Processa hawaiensis (Dana, 1852a)
= Nika Hawaiensis Dana, 1852a: 20. [prope insulam Maui Hawaiensem]
= Processa paucirostris Edmondson, 1930: 3; Fig. 1. [Kahana Bay, Oahu, in shallow water]
Processa hemphilli Manning & Chace, 1971: 23; Figs 11-12. [Florida, Collier County, Marco, 2-6 m]
Processa indica Nol, 1986: 282; Figs 14-17. [Inde, cte N-E (baie du Bengale), proximit de Visakhapat-
nam, 40-60 m]
Processa intermedia Holthuis, 1951a: 38; Figs 5-6. [of French Guinea, 957'N 1522'W, 25 m]
Processa kotiensis (Yokoya, 1933)
= Nika kotiensis Yokoya, 1933: 30; Fig. 13. [Sy-Maru stn 322, west of Muroto-zaki, 110 m]
Processa longirostris Hayashi, 1975a: 118; Figs 26-27. [South Viet Nam, Bay of Nha Trang, 5 m]
Processa macrodactyla Holthuis, 1952a: 30; Fig. 8. [Baie de Gorri, au S. dAngra de Cintra, Rio de Oro]
Processa macrognatha (Stimpson, 1860a)
= Nica macrognatha Stimpson, 1860a: 26. [in portu Hong Kong f. conchoso, p. 8 org.]
Processa macrophthalma Nouvel & Holthuis, 1957: 27; Figs 108-133. [prs de Monaco]
Processa moana Yaldwyn, 1971: 91. [of Cuvier Id, Bay of Plenty, 30-40 fms]
Processa modica modica Williamson in Williamson & Rochanaburanon, 1979: 12; Figs 1, 2a-e, 3f-g, 4-5, 9d.
[5219'N 422'E, about 25 km north of The Hague, The Netherlands]
Processa modica carolii Williamson in Williamson & Rochanaburanon, 1979: 21; Figs 2f-h, 3h-i, 6a, d, g.
[Tunisia, Carthage-Salammb, from of Baths of Antoninus to INSTOP, 4-5 m]
Processa namibiensis Macpherson, 1983: 60; Figs 31-32. [norte de Namibia, 1744'S 1134'E, 70 m]
Processa neglecta Hayashi, 1975a: 127; Figs 31-32. [South Viet Nam, Bay of Nha Trang, 11 m]
Processa nouveli nouveli Al-Adhub & Williamson, 1975: 694; Figs 1a-b, 2a-b, 3. [Monaco]
Processa nouveli holthuisi Al-Adhub & Williamson, 1975: 701. [20 km north-west of Port Erin, Isle of Man,
90 m]
Processa packeri Manning & Chace, 1990: 24; Fig. 11. [Ascension Island, McArthur Point, northern edge
of South West Bay]
Processa parva Holthuis, 1951a: 47; Fig. 8. [of Bathurst, Gambia, 18 m]
Processa peruviana Wicksten, 1983a: 29; Figs 4-6. [Isla Manuelita, Costa Rica, 536'09''N 8701'14''W, 146 m]
Processa philippinensis Nol, 1986: 288; Fig. 18. [Philippines, 353'N 12008'E, 134-129 m]
Processa pippinae Wicksten & Mndez G., 1985: 16; Figs 1-4. [E of Isla ngel de la Guardia, Gulf of Cali-
fornia, Mexico, 2920'N 11311.5'W, 265-284 m]
Processa processa (Spence Bate, 1888)
= Nika processa Spence Bate, 1888: 527. (nec Plate 95, fg. 1) [Amboina, 15 fms]
Processa profunda Manning & Chace, 1971: 25; Figs 13-15. [Gulf of Mexico, of west coast of Florida,
2836'N 8533'30''W, 202 m]
Processa pygmaea Burukovsky, 1990: 191; Fig. 1B. [2504'S 9729'W, 267-280 m]
Processa riveroi Manning & Chace, 1971: 28; Fig. 16. [Puerto Rico, Lajas, La Parguera, east side of Maguey
Island, on Thalassia fats]
Processa robusta Nouvel & Holthuis, 1957: 19; Figs 53-80. [prs de Monaco, Baie de Monte-Caro, fond de
Posidonies, 15 m]
Processa sulcata Hayashi, 1975a: 134; Fig. 34. [Japan, Sea of Genkai, Ainoshima Island, Fukuoka Prefec-
ture]
Fig. 53. Processa fmbriata Manning & Chace, 1971. Photo by Arthur Anker.
Processa vicina Manning & Chace, 1971: 34; Figs 19-20. [of north Carolina, 3435'30''N 7545'30''W, 59 m]
Processa vossi Manning, 1992: 552; Figs 1-2. [Florida, Atlantic Ocean, Martin County, about 1 mile north
of St. Lucie Inlet, 2711'N 8009.5'W, Bathtub Beach]
Processa wheeleri Lebour, 1941: 403; Figs 1-9, 11-27. [5 miles from Bermuda, 150 m]
Processa zostericola Hayashi, 1975a: 137; Fig. 35. [Japan, Tomioka Bay, Amakusa Islands, Kumamoto Pre-
fecture, Zostera belt]
Superfamily PANDALOIDEA Haworth, 1825
Family PANDALIDAE Haworth, 1825
Anachlorocurtis Hayashi, 1975b
= Anachlorocurtis Hayashi, 1975b (type species Anachlorocurtis commensalis Hayashi, 1975b, by origi-
Anachlorocurtis commensalis Hayashi, 1975b: 175; Figs 1-3. [of Andonohana, Shionomisaki, southern Kii
Peninsula, Japan, 8 m, associated with Antipathes sp.]
Atlantopandalus Komai, 1999
= Atlantopandalus Komai, 1999 (type species Pandalus propinqvus G.O. Sars, 1870, by original desig-
Atlantopandalus propinqvus (G.O. Sars, 1870)
= Pandalus propinqvus G.O. Sars, 1870: 148. [Lofoten near Skrava, Guldbrandsy, 360-540 m; lecto-
type designation by Komai, 1999; spelling maintained as propinqvus under Art. 32.1]
Austropandalus Holthuis, 1952d
= Austropandalus Holthuis, 1952d (type species Hippolyte Grayi Cunningham, 1871, by original des-
Austropandalus grayi (Cunningham, 1871)
= Hippolyte Grayi Cunningham, 1871: 496; Plate 59, fg. 8. [Port Otway]
= Pandalus paucidens Miers, 1881b: 74; Plate 7, fgs 6-7. [Tom Bay, on a botom of rock, kelp, and mud;
Trinidad Channel, 30 fms, on a sandy botom]
= Nothocaris spiniserratus Spence Bate, 1888: 663; Plate 132, fg. 1. [Challenger stn 304, 4653'15''S
7512'0''W, Port Otway, Messier Channel, 45 fms]
Bitias Fransen, 1990
= Bitias Fransen, 1990 (type species Bitias stocki Fransen, 1990, by original designation and mono-
Bitias brevis (Rathbun, 1906)
= Pandalus brevis Rathbun, 1906: 916; Fig. 65; Plate 21, fg. 3. [vicinity of Niihau Island, 426-417 fms]
Bitias stocki Fransen, 1990: 68; Figs 1-3. [Azores, S of Pico, 3809'N 2831'W, 1320-1350 m]
Calipandalus Komai & Chan, 2003
= Calipandalus Komai & Chan, 2003 (type species Calipandalus elachys Komai & Chan, 2003, by orig-
inal designation and monotypy, gender masculine)
Calipandalus elachys Komai & Chan, 2003: 880-889; Figs 1-4. [2218.59'N 12129.39'E, 267-302 m]
Chelonika Fransen, 1997b
= Chelonika Fransen, 1997b (type species Chelonika macrochela Fransen, 1997b, by original designa-
Chelonika macrochela Fransen, 1997b: 179, Figs 1-7. [New Caledonia, 2221.5'S 16614.6'E, 144-155 m]
Chlorocurtis Kemp, 1925
= Chlorocurtis Kemp, 1925 (type species Chlorocurtis miser Kemp, 1925 (junior subjective synonym of
Virbius (?) jactans Nobili, 1904), by original designation and monotypy, gender masculine; name
Chlorocurtis jactans (Nobili, 1904)
= Virbius (?) jactans Nobili, 1904: 230. [Djibouti]
= Chlorocurtis miser Kemp, 1925: 280. [Port Blair, Andamans, low water]
Chlorotocella Balss, 1914b
= Chlorotocella Balss, 1914b (type species Chlorotocella gracilis Balss, 1914b by monotypy, gender fem-
inine; name placed on the Ofcial List of Generic Names in Zoology in Opinion 470 in 1957)
Chlorotocella gracilis Balss, 1914b: 33; Figs 16-22. [Type locality restricted to Sagami Bay (Japan) by
Hayashi & Miyake, 1968b]
Chlorotocella spinicaudus (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840])
= Hippolyte spinicaudus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 378. [les ctes de
la Nouvelle Hollande]
= Pandalus leptorhynchus Stimpson, 1860a: 38; nec Kinahan, 1858b. [Portu Jacksoniensi Australiae, ad
litora arenosa et algosa]
= Pandalus (Parapandalus) spinicauda var. gibber Hale, 1924: 68; Plate 4, fgs 6-7. [St. Vincent Gulf]
Chlorotocus A. Milne-Edwards, 1882
= Chlorotocus A. Milne-Edwards, 1882 (type species Chlorotocus gracilipes A. Milne-Edwards, 1882,
by monotypy, gender masculine; name placed on the Ofcial List of Generic Names in Zoology in
Chlorotocus crassicornis (A. Costa, 1871) (Fig. 54)
= Pandalus crassicornis A. Costa, 1871: 89; Plate 2, fg. 2. [golfo di Napoli]
= Chlorotocus gracilipes A. Milne-Edwards, 1882: 14. [Alboran Sea of Mediterranean coast of Mo-
rocco, 3521'30"N 433'05"W 3524'20"N 432'45'W, 322-432 m; see Chace, 1985]
= Palemon chlorotocus Filhol, 1885b: Plate 1. [Type locality not indicated]
? = Chlorotocus incertus Spence Bate, 1888: 674; Plate 116, Figs 1-2. [Challenger stn 142, 354'S 1837'E,
Agulhas Bank, of the Cape of Good Hope, 150 fms]
Fig. 54. Chlorotocus crassicornis (A. Costa, 1871). Photo by Tin-Yam Chan.
= Chlorotocus gracilipes var. andamanensis Alcock & Anderson, 1899: 284. [Andaman Sea, east of
North Andaman Island, 1317'15"N 9310'15"E, 185 fms]
Chlorotocus novaezealandiae (Borradaile, 1916)
= Thalassocaris novae-zealandiae Borradaile, 1916: 84; Fig. 2. [7 miles E of North Cape, New Zealand,
128 m]
Dichelopandalus Caullery, 1896
= Dichelopandalus Caullery, 1896 (type species Dichelopandalus Bonnieri Caullery, 1896, by monotypy,
1957)
Dichelopandalus bonnieri Caullery, 1896
= Dichelopandalus Bonnieri Caullery, 1896: 379; Plate 15, fgs 7-15. [4530'N 553'W, 1200 m; 4518'N
523'W, 180 m; 4518'N 523'W, 300-400 m; 4713'N 556'W, 110 m (all Golfe de Gascogne)]
Dichelopandalus leptocerus (Smith, 1881)
= Pandalus leptocerus Smith, 1881: 437. [Fish Hawk stns 870 (of Block Island, R.I., 4002'36"N
7022'58"W, 155 fms); 873 (of Newport, R.I., 4002'N 7057'W, 100 fms); 878 (of Block Island, R.I.,
3955'N 7054'15"W, 142 fms)]
= Pandalus falcipes Spence Bate, 1888: 668; Plate 115, fg. 2. [Challenger stn 49, 4303'N 6339'W, south
of Halifax, Nova Scotia, 85 fms]
Dorodotes Spence Bate, 1888
= Dorodotes Spence Bate, 1888 (type species Dorodotes refexus Spence Bate, 1888, designated by
Holthuis, 1955b, gender masculine; name placed on the Ofcial List of Generic Names in Zoology
Dorodotes refexus Spence Bate, 1888: 678; Plate 116, fg. 3. [Challenger stns 195, 421'S 12907'E, near
Banda Island, 1425 fms; 205, 1642'N 11922'E, Philippine Islands, 1050 fms]
Heterocarpus A. Milne-Edwards, 1881b
= Heterocarpus A. Milne-Edwards, 1881b (type species Heterocarpus ensifer A. Milne-Edwards, 1881b,
by monotypy, gender masculine; name placed on the Ofcial List of Generic Names in Zoology in
= Atlantocaris Ortmann, 1893 (type species Atlantocaris gigas Ortmann, 1893 (junior subjective synonym
of Heterocarpus ensifer A. Milne-Edwards, 1881b), designated by Holthuis, 1955b, gender feminine)
Heterocarpus abulbus Yang, Chan & Chu, 2010: 207; Figs 1, 3A-B, 4A-D, 5A. [Taiwan, Dasi fshing port,
Yilan County, commercial trawler]
Heterocarpus afnis Faxon, 1893: 204. [Albatross stns 3418 (1630'00''N 09952'30''W), 660 fms; 3424
(2115'00''N 10623'00''W), 676 fms; 3425 (2119'00''N 10624'00''W), 680 fms]
Heterocarpus agassizi Allen & Butler, 1994: 431; Figs 7-9. [Mid-Pacifc Ocean, Darwin Guyot, 17136.0'E
2207.7'N, 1281 m]
Heterocarpus amacula Crosnier, 1988c: 72; Fig. 4a; Plate 1b; Plate 2h. [Polynsie franaise, Tuamotu
(Mururoa), 250-600 m]
Heterocarpus calmani Crosnier, 1988c: 59; Fig. 1a-h. [Madagascar, 1252'S 4810.3'E, 420-428 m]
Heterocarpus chani Li, 2006b: 362; Figs 1-4. [Philippines, 852.1'N 12337.1'E, 569-597 m depth]
Heterocarpus corona Yang, Chan & Chu, 2010: 210; Figs 2, 3D-F, 4G-H, 5C-D. [Vanuatu, 1541.1'S
16700.5'E, 509-659 m]
Heterocarpus cutressi Monterrosa, 1988: 633; Figs 1-3. [southwest of Puerto Rico, 1751.7'N 6715.4'W, 777
m]
Heterocarpus dorsalis Spence Bate, 1888: 630; Plate 112. [Challenger stn 194, 434'00"S 12957'30"E, of Ban-
da Island, 200 fms]
= Heterocarpus alphonsi Spence Bate, 1888: 632; Plate 112, fg. 1. [433'N 1276'E, south of the Philip-
pines, 500 fms; 3511'N 13928'E, of Japan, 345 fms]
= Heterocarpus afnis Borradaile, 1915b: 208; nec Faxon, 1893. [Saya de Malha, 300-500 fms]
Heterocarpus ensifer A. Milne-Edwards, 1881b: 8. [prs des Barbades 218 brasses]
= Pandalus carinatus Smith, 1882: 63; Plate 10, fgs 2-2f; Plate 11, fgs 1-3. [Blake stn 327, 340'30"N
7610'30"W, 178 fms]
= Atlantocaris gigas Ortmann, 1893: 80; Plate 5, fg. 3. [Plankton-Expedition der Humboldt-Stifung,
Sdl. Aequatorialstrom, JN 209 (5.7S 26.5W, 0-400 m); Pl. 88 (6.6S 24.5W, 200 m)]
= Procletes (?) atlanticus Lenz & Strunck, 1914: 334; Plate 22, fgs 9-4. [nhe der Inseln Ascension,
843'S 1155'W, 3000 m]
Heterocarpus gibbosus Spence Bate, 1888: 634; Plate 112, fg. 2. [Bohol Sea, 931.5'N 12341.7'E, 336 m;
neotype selection by Li et al., 2007]
Heterocarpus grimaldii A. Milne-Edwards & Bouvier, 1900
= Heterocarpus Grimaldii A. Milne-Edwards & Bouvier, 1900: 58. [entre Pico et So Jorge, 1330 m]
Heterocarpus hayashii Crosnier, 1988c: 81; Figs 4b, 5b; Plate 1, fg. d; Plate 3, fgs c-e. [les Chesterfeld Is-
lands, 1940.86'S 15846.0'E, 615-600 m] (Fig. 55)
Heterocarpus hostilis Faxon, 1893: 204. [Albatross stns 3353 (0706'15''N 08034'00''W), 695 fms; 3363
(0543'00''N 08550'00''W), 978 fms; 3364 (0530'00''N 08608'30''W), 902 fms; 3371 (0526'20''N
08655'00''W), 770 fms; 3380 (0403'00''N 08131'00''W), 899 fms; 3393 (0715'00''N 07936'00''W),
1020 fms; 3395 (0730'36''N 07839'00''W), 730 fms]
Heterocarpus inopinatus Tavares, 1999: 673; Fig. 1. [1938'S 3843'W, 960 m]
Heterocarpus intermedius Crosnier, 1999b: 346; Fig. 1. [MUSORSTOM 5, stn CP 363(1947'53.9874''S
15844'17.9874''E), Chesterfeld Islands, Coral Sea, 685-700 m]
Heterocarpus laevigatus Spence Bate, 1888
= Heterocarpus lvigatus Spence Bate, 1888: 636; Plate 112, fg. 3. [434'0"S 12957'30"E, of Banda Is-
land, 200 fms]
Heterocarpus lepidus De Man, 1917: 282. [West 1000 m distant from North point of Kabia-island reef,
Flores Sea, 500 m; 553'.8S 13248'8E, Kei-islands, 560 m]
Heterocarpus longirostris MacGilchrist, 1905: 237. [Investigator stn 310, Bay of Bengal, 960 fms (1329'30"N,
9529'E)]
= Heterocarpus facetus Zarenkov & Khodkina, 1981: 83; Fig. 1. [central part of the Marcus-Nekker
seamount chain, 1270-1320 m]
Fig. 55. Heterocarpus hayashii Crosnier, 1988. Photo by Tin-Yam Chan.
Heterocarpus nasicus Timofeev, 1993: 40; Fig. 2. [Gulf of Aden, 1131'3N 5240'1E, 490-500 m]
Heterocarpus oryx A. Milne-Edwards, 1881b: 10. [2436'N 8405'W, 955 brasses]
Heterocarpus parvispina De Man, 1917
= Heterocarpus ensifer var. parvispina De Man, 1917: 282. [735'.4S, 11728'.6E, Bali Sea, 521 m; 68'N
12119'E, north of Sulu Island, 275 m; 540'S 13226'E, of the Kei-islands, 310 m]
Heterocarpus reedi Bahamonde, 1955: 106; Figs 1-4. [entres los Puertos de Valparaiso y Quinteros, Chili,
329 m]
Heterocarpus sibogae De Man, 1917: 283. [715'S 11515'.6E, Bali Sea, 289 m; 735'.4S 11728'.6E, Bali Sea,
521 m; 53'.5S 1190'E, southern entrance of Strait of Makassar, 450 m; 011'S 12725'E, North of
Batjan, 397 m; 554'.5S 12019'.2E, West of Saleyer, 462 m; 526'.6S 13232'.5E, Kei-islands, 397 m;
553'.8S 13248'.8E, Kei-islands, 560 m; 719.'4S 11649'.5E, Bali Sea, 538 m]
Heterocarpus signatus Rathbun, 1906: 918; Plate 21, fg. 6. [West coast of Hawaii Island, 382-253 fms]
Heterocarpus tenuidentatus Cleva & Crosnier, 2006: 61; Figs 1-3. [Solomon Islands, New Georgia Sound,
841.5'S 15738.2'E, 814-980 m]
Heterocarpus tricarinatus Alcock & Anderson, 1894: 154. [Investigator stn 122, Laccadive Sea, 1205'35"N
7135'50"E, 880 fms]
= Heterocarpus tricarinatus angustus Crosnier, 1988c: 84; Plate 4, Figs g-h. [539'S 12212'E, au nord de
lle Muna, 1886 m]
Heterocarpus vicarius Faxon, 1893: 203. [Albatross stns 3385 (0732'36''N 07916'00''W), 286 fms; 3386
(0733'12''N 07917'15''W), 242 fms; 3389 (0716'45''N 07956'30''W), 210 fms; 3396 (0732'00''N
07836'30''W), 259 fms]
Heterocarpus woodmasoni Alcock, 1901
= Heterocarpus Wood-masoni Alcock, 1901: 108. [Andaman Sea, 265 fms]
Miropandalus Bruce, 1983g
= Miropandalus Bruce, 1983g (type species Miropandalus hardingi Bruce, 1983g, by original designa-
Miropandalus hardingi Bruce, 1983g: 483; Figs 1-5. [Enewetak Island, Marshall Islands, 23 m]
Notopandalus Yaldwyn, 1960
= Notopandalus Yaldwyn, 1960 (type species Pandalus magnoculus Spence Bate, 1888, by original des-
Notopandalus magnoculus (Spence Bate, 1888)
= Pandalus magnoculus Spence Bate, 1888: 667; Plate 115, fg. 1. [3850'S 16920'E, near New Zealand,
275 fms; 3932'S 17148'E, of the west coast of New Zealand, 150 fms]
Pandalina Calman, 1899b
= Pandalina Calman, 1899b (type species Pandalus brevirostris Rathke, 1843, by original designation
and monotypy, gender feminine; name placed on the Ofcial List of Generic Names in Zoology in
Pandalina brevirostris (Rathke, 1843)
= Pandalus brevirostris Rathke, 1843: 17. [Molde]
= Hippolyte Thompsoni Bell, 1851 [in Bell, 1844-1853]: 290; unnumbered vignete [?Belfast]
= Pandalus Rathkii Heller, 1863b: 441; Plate 3, fg. 31. [Lissa und Lesina in 20-30 Faden Tiefe]
Pandalina modesta (Spence Bate, 1888)
= Pandalus modestus Spence Bate, 1888: 670; Plate 114, fg. 4. [354'S 1837'E, at the Agulhas Bank, of
the Cape of Good Hope, 150 fms]
Pandalina nana Burukovsky, 1990: 201; Fig. 4A. [2503'S 9727'W, 271 m]
Pandalina profunda Holthuis, 1946b: 281; Fig. 1a-c. [Barents Sea; Bergen, Norway]
Pandalina spinicauda Komai & Chan, 2010b: 626; Figs 1-2, 7A. [Mozambique Channel, MAINBAZA stn
CP 3130, 2553'S 3307'E, 112-127 m]
Pandalopsis Spence Bate, 1888
= Pandalopsis Spence Bate, 1888 (type species Pandalopsis amplus Spence Bate, 1888, by monotypy,
1959)
Pandalopsis aleutica Rathbun, 1902a: 901. [of Segouam, Aleutians, 283 fms]
Pandalopsis ampla Spence Bate, 1888
= Pandalopsis amplus Spence Bate, 1888: 671; Plate 115, fg. 3. [3717'S 5352'W, of Monte Video,
600 fms]
Pandalopsis coccinata Urita, 1941: 12; Figs 1-2. [of Totori-mura, near Kushiro, Hokkaido, Japan]
Pandalopsis dispar Rathbun, 1902a: 903. [Chernofski Harbor, Unalaska, 109 fms]
Pandalopsis gibba Komai & Takeda, 2002: 91; Figs 1-5. [Sagami Bay, 3504.68'N 13945.32'E, 250 m]
Pandalopsis glabra Kobjakova, 1936b
= Pandalopsis aleutica var. glabra Kobjakova, 1936b: 192; Fig. 4. [Ochotsk Meer, 515 and 1076 m]
Pandalopsis japonica Balss, 1914b
= Pandalopsis dispar var. japonica Balss, 1914b: 32. [Etschiu]
Pandalopsis lamelligera (Brandt, 1851)
= Pandalus lamelligerus Brandt, 1851: 124; Plate 5, fgs 20-20b. [von der Chantarischen Breninsel; im
Ochotskischen Meer]
Pandalopsis longipes Komai, 1994: 545; Figs 4-5. [of Nemuro, Hokkaido]
Pandalopsis longirostris Rathbun, 1902a: 902. [of Iliuliuk Harbor, Unalaska, 309 fms]
Pandalopsis lucidirimicola Jensen, 1998: 82; Figs 1-3. [Point Atkinson, Burrard Inlet, British Colombia,
Canada, 4919'N 12315'W, 18 m]
Pandalopsis miyakei Hayashi in Baba, Hayashi & Toriyama, 1986: 123; Fig. 19; unnumbered plate. [Tosa
Bay, 700 m]
Pandalopsis multidentatus Kobjakova, 1936b: 188; Fig. 1. [Japanisches Meer von der Peter-des-Groen-
Bucht nrdlich bis zum Kap Terpenjia (Tataren-Sund), 64-500 m]
Pandalopsis ochotensis Kobjakova, 1936b: 190; Fig. 2. [Ochotsk Meer, 500-504 m]
Pandalopsis pacifca (Dofein, 1902)
= Pandalus pacifcus Dofein, 1902: 619; Plate 4, fg. 1. [Nemuro, Nordspitze der Insel Yesso]
= Pandalopsis mitsukurii Rathbun, 1902b: 48; Figs 22-23. [Mororan, Hokkaido]
Pandalopsis profundus Zarenkov, 1971: 189; Fig. 3(11-20). [south of Japan, 820 m]
Pandalopsis punctata Kobjakova, 1936b
= Pandalopsis punctatus Kobjakova, 1936b: 191; Fig. 3. [Ochotsk Meer, 165 m]
Pandalopsis rubra Komai, 1994: 553; Figs 1, 9-10. [eastern Hokkaido, 4246.0'N 14519.5'E, 1080-1120 m]
Pandalopsis spinosior Hanamura, Khono & Sakaji, 2000: 27; Figs 1-3. [Urup (or Etorofu) Strait, around
400 m]
Pandalopsis zarenkovi Ivanov & Sokolov, 2001: 160; Figs 1-3. [Bering Sea, 6143.4'N 17737.8'W, 362 m]
Pandalus Leach, 1814 [in Leach, 1813-1814]
= Pandalus Leach, 1814 [in Leach, 1813-1814] (type species Pandalus Montagui Leach, 1814 [in Leach,
1813-1814], by monotypy, gender masculine; name placed on the Ofcial List of Generic Names in
Zoology in Opinion 104 in 1928 and in Direction 47 in 1956)
= Dyma Kryer, 1861 (type species Dymas typus Kryer, 1861, by monotypy and by indication under
Art. 68c of the ICZN)
= Boreocaris Ortmann, 1893 (type species Boreocaris mbiusi Ortmann, 1893 (probably a junior subjec-
tive synonym of Pandalus Montagui Leach, 1814 [in Leach, 1815-1875]), gender feminine)
Pandalus borealis Kryer, 1838: 254. [Sydprven, Julianehaab, Fisksset, &c]
= Dymas typus Kryer, 1861: 63. [Grnland]
= Pandalus borealis v. edenticulatus Retowsky, 1946: 300; Figs 2a-d. [Arctic Ocean]
Pandalus chani Komai, 1999: 1337; Figs 33-35, 44B. [Su-Aou, I-Lan County, northeast of Taiwan, circa
400 m]
Pandalus curvatus Komai, 1999: 1302; Figs 16-17. [southern Japan, Amakusa-nada, western Kyushu,
3208.6'N 12941.2'E, 501 m]
Pandalus danae Stimpson, 1857
= Pandalus dan Stimpson, 1857: 87; Plate 21, fgs 6-7. [opposite Fort Townsend, Puget Sound]
= Pandalus franciscorum Kingsley, 1878a: 94. [San Francisco, California]
Pandalus eous Makarov, 1935
= Pandalus borealis var. eous Makarov, 1935: 321; Figs 2, 3b. [im Bering-Meere nahe dem Nawarin-
Kap, 6133'N 17909'W, 128 m; Bering-Meer bei dem Kap Oljutorsky, 5948'N 17043'E, 142 m; nahe
der Bering-Insel, 5225'N 16544'E, 234 m]
Pandalus formosanus Komai, 1999: 1342; Figs 36-37, 44C. [Su-Aou, I-Lan County, NE Taiwan, about
500 m]
Pandalus goniurus Stimpson, 1860a: 36. [in sinu Avatska Kamtschatkae, in fundo limoso prof. 10 org]
= Pandalus dapifer Murdoch, 1884: 519. [of Point Franklin, 13.5 fms]
Pandalus gracilis Stimpson, 1860a: 37. [Hakodate Bay, Hokkaido Island, Japan; lectotype designation by
Holthuis, 1976a]
= Pandalus robustus Stimpson, 1860a: 37. [sinu Hakodadi, in profundis]
= Pandalus stimpsoni Thallwitz, 1892: 3. [Japan oder China]
Pandalus gurneyi Stimpson, 1871
= Pandalus Gurneyi Stimpson, 1871: 128. [Monterey, Cal.]
Pandalus hypsinotus Brandt, 1851: 125. [Unalaschka]
Pandalus ivanovi Komai & Eletskaya, 2008: 47; Figs 1-6. [of Eastern Sakhalin, Sea of Okhotsk, 46N
145E, 150-200 m]
Pandalus jordani Rathbun, 1902a: 900. [of Santa Cruz Island, California, 155 fms]
Pandalus latirostris Rathbun, 1902b: 46; Figs 20-21. [Mororan, Hokkaido]
= Pandalus Kessleri Czerniavsky, 1878: 23. [nomen nudum]
Pandalus montagui Leach, 1814 [in Leach, 1813-1814]
= Pandalus Montagui Leach, 1814 [in Leach, 1813-1814]: 432. [Zetland, Scotland; lectotype designa-
tion by Holthuis, 1956b]
= Astacus maculatus Leach, 1814 [in Leach, 1813-1814]: 432. [nomen nudum]
= Pandalus annulicornis Leach, 1815 [in Leach, 1815-1875]: Plate 40. [Zetland, Scotland; lectotype
designation by Holthuis, 1956b]
? = Pandalus levigatus Stimpson, 1854: 58. [Grand Manan]
= P.[andalus] Leptorhynchus Kinahan, 1858b: 41; unnumbered fgures. [Sandycove, prope Dublin]
= P.[andalus] Leptorhynchus Kinahan, 1860b: 80; unnumbered fgures. [Sandycove, prope Dublin]
? = Boreocaris mbiusi Ortmann, 1893: 84; Plate 6, fg. 3; Plate 7, fg. 1. [Plankton-Expedition der
Humboldt-Stifung, Golfstrom, JN 4 (59.2N 11.8W, 0-400 m)]
Pandalus nipponensis Yokoya, 1933: 16; Fig. 5. [of Tohtomi, Suruga Bay, 100 m; lectotype designation by
Komai, 1999]
Pandalus platyceros Brandt, 1851: 123. [Insel Unalaschka]
= Pandalus pubesculentus Dana, 1852a: 24. [in freta de Fuca Oregoni, juxta portum Dungeness]
Pandalus prensor Stimpson, 1860a: 37. [sinu Hakodadi, fundo conchoso, prof. 8 org.]
= Pandalus hypsinotus var. meridionalis Balss, 1914b: 29. [Nagasaki bis Nemuro und Wladiwostok]
Pandalus stenolepis Rathbun, 1902a: 901. [Straits of Fuca, 40 fms]
Pandalus teraoi Kubo, 1937a: 98; Figs 4-6. [of yama, Prefecture Aiti]
Pandalus tridens Rathbun, 1902a
= Pandalus montagui tridens Rathbun, 1902a: 901. [of North Head, Akutan Island, Alaska, 72 fms]
Pantomus A. Milne-Edwards, 1883
= Pantomus A. Milne-Edwards, 1883 (type species Pantomus parvulus A. Milne-Edwards, 1883, by
Pantomus afnis Chace, 1937b: 116; Fig. 3a-b. [Santa Inez Bay, 2656'30''N 11148'30''W, 60 fms]
Pantomus parvulus A. Milne-Edwards, 1883: Plate 26. [North part of Yucatan Bank, 2313'N 8916'W, 84
brasses]
Peripandalus De Man, 1917
= Peripandalus De Man, 1917 (type species Pandalus serratus A. Milne-Edwards, 1873, by monotypy,
gender masculine)
Peripandalus serratus (A. Milne-Edwards, 1873)
= Pandalus serratus A. Milne-Edwards, 1873: 87. [Upolu]
Plesionika Spence Bate, 1888
= Nisea Risso, 1844 (type species Nisea formosa Risso, 1844 (junior subjective synonym of Astacus
Narual Fabricius, 1787), gender feminine; nomen nudum, see Holthuis, 1977)
= Plesionika Spence Bate, 1888 (type species Plesionika uniproducta Spence Bate, 1888 (junior subjec-
tive synonym of Acanthephyra ensis A. Milne-Edwards, 1881b), designated by Alcock, 1901, gender
= Nothocaris Spence Bate, 1888 (type species Nothocaris rostricrescentis Spence Bate, 1888, designated
by Fowler, 1912, gender feminine)
= Parapandalus Borradaile, 1900 (type species Pandalus (Parapandalus) serratifrons Borradaile, 1900,
designated by Alcock, 1901, gender masculine; name placed on the Ofcial List of Generic Names
= Parapandalopsis Dong, Chen & Wang, 1986 (type species Plesionika spinipes Spence Bate, 1888, by
monotypy, gender feminine; see Holthuis, 1993a)
Plesionika acanthonotus (Smith, 1882)
= Pandalus acanthonotus Smith, 1882: 61; Plate 13, fgs 10-11. [Blake stn 321, 3243'25"N 7720'30"W,
233 fms]
= Pandalus Parfaitii A. Milne-Edwards, 1883: Plate 21. [East Atlantic, 444'30"N 925'W (of Paris),
119 m]
= Pandalus geniculatus A. Milne-Edwards, 1883: Plate 25. [East Atlantic, 444'30"N 925'W (of Paris),
119 m]
Plesionika acinacifer Chace, 1985: 51; Figs 21-23. [Between southern Cebu and Siquior, Philippines,
927'15"N 12331'48"E, 790 m]
Plesionika adensameri (Balss, 1914c)
= Parapandalus Adensameri Balss, 1914c: 134. [Rote Meer, zwischen 18 und 26 n. Br., von 800 bis 1300
m; fully described in Balss, 1915]
Plesionika albocristata Chan & Chuang, 2002: 611; Figs 1-2. [Taiwan, NE coast, Su-Aou fshing port, I-Lan
County, about 350 m]
Plesionika alcocki (Anderson, 1896)
= Pandalus alcocki Anderson, 1896: 92. [934'57"N 7536'30"E, 406 fms]
= Plesionika adenensis Timofeev, 1993: 44; Fig. 4. [Gulf of Aden, 1218'7N 4428'E, 610 m]
Plesionika alexandri (A. Milne-Edwards, 1883)
= Heterocarpus Alexandri A. Milne-Edwards, 1883: Plate 28. [north of Havana, 2314'00''N 8225'00''W,
805 fms]
Plesionika antigai Zariquiey lvarez, 1955: 11. [Barcelona, barcas de arrastre, ms de 300 m]
Plesionika assimilis De Man, 1917: 280. [Madura-bay and other localities in the southern part of Molo-
strait, 54-90 m]
Plesionika beebei Chace, 1937b: 114; Fig. 2. [13 miles northeast by north of San Ildefonso Island, 2648'40''N
11120'30''W, 300 fms]
Plesionika bifurca Alcock & Anderson, 1894: 155. [Investigator stn 177, Laccadive Sea, 1347'49"N 7307'E,
636 fms]
Plesionika bimaculata Chan, 2004: 295; Figs 1, 10. [Vanuatu, 2018.87'S 16953.12'E, 250-315 m]
Plesionika binoculus (Spence Bate, 1888)
= Nothocaris binoculus Spence Bate, 1888: 656; Plate 114, fg. 2. [856'S 13605'E, Arafura Sea, south of
New Guinea, 49 fms]
Plesionika brevipes (Crosnier & Forest, 1968)
= Parapandalus brevipes Crosnier & Forest, 1968: 1136; Fig. 5. [au large de Pointe-Noir, Congo, par
500'S 1122'E et 506'S 1126'E, 190-355 m]
Plesionika brevirostris Spence Bate, 1888: 650; Plate 113, fg. 5. [647'N 12228'E, between the Philippine
Islands and Borneo, 250 fms]
Plesionika carinata Holthuis, 1951a: 59; Fig. 12. [of Angola, 755'S 1238'E, 235-460 m]
Plesionika carinirostris Hendrickx, 1990b: 38; Figs 2-3. [of Isla San Lorenzo, 2846'N 11254'W, 360-380 m]
Plesionika carsini Crosnier, 1986c: 369; Fig. 3. [Iles Tuamotu, Mururoa, 450 m]
Plesionika chwitchii Burukovsky, 1978: 1729; Figs 1-4 [south-west Atlantic, 3326'S 5058'W, 130 m]
Plesionika costelloi (Yaldwyn, 1971)
= Parapandalus costelloi Yaldwyn, 1971: 91. [of Astrolabe Reef, Bay of Plenty, 65 fms]
Plesionika crosnieri Chan & Yu, 1991b: 546; Figs 1, 3a. [Ta-Chi, I-Lan County, northeastern Taiwan, fsh
market]
? = Plesionika dentirostris Tung, Wang & Li, 1988: 20; Fig. 1. [East China Sea, 400 m]
Plesionika curvata Chan & Crosnier, 1991: 440; Figs 10b, 11d-f, 33. [Tubuai, French Polynesia, 2340'S
14940'W, 200 m]
Plesionika echinicola Chan & Crosnier, 1991: 416; Figs 1a, 2a, 3a-b, 19-20. [New Caledonia, 2447.0'S
16809.4'E, 274 m]
Plesionika edwardsii (Brandt, 1851)
= Pandalus Guerinii Risso, 1844: 95. [nomen nudum]
= Pandalus (Pontophilus) Edwardsii Brandt, 1851: 122; nomen novum for Pandalus narval H. Milne
Edwards, 1841 [in H. Milne Edwards, 1836-1844] nec Astacus Narual Fabricius, 1787. [Mediterrane-
an, type locality designated by Chan & Yu, 1991b]
= Pandalus (Parapandalus) longirostris Borradaile, 1900: 413; Plate 37, fg. 10a-h. [New Britain]
Plesionika ensis (A. Milne-Edwards, 1881b)
= Acanthephyra ensis A. Milne-Edwards, 1881b: 14. [prs de Barbades, 237 brasses]
= Plesionika uniproducta Spence Bate, 1888: 641 (partim); Plate 113, fg. 1. [95'S 3450'W, of Barra
Grande, 350 fms]
Plesionika erythrocyclus Chan & Crosnier, 1997: 214; Figs 16, 34-35. [Tuamotu Islands, Mururoa, French
Polynesia, 350-600 m]
Plesionika exigua (Rathbun, 1906)
= Pandalus exiguus Rathbun, 1906: 916; Plate 21, fg. 2. [vicinity of Kauai, Hawaii Islands, 233-40 fms]
Plesionika fenneri Crosnier, 1986d: 691. [nomen novum for Plesionika chacei Crosnier, 1986c]
= Plesionika chacei Crosnier, 1986c: 363; Figs 1a-b, 2a-h; nec Hayashi in Baba, Hayashi & Toriyama,
1986. [Iles Australes, Tubuai, 700 m]
Plesionika fmbriata Chace, 1985: 63; Fig. 27. [Samar Sea east of Masbate, Philippines, 1213'15"N
12405'03"E, 216 m]
Plesionika favicauda Chan & Crosnier, 1991: 438; Figs 10a, 11a-c, 29-32. [Rurutu, French Polynesia,
2227.8'S 15122.9'W, 240-260 m]
Plesionika gigliolii (Senna, 1902)
= Pandalus Gigliolii Senna, 1902: 315; Plate 16, Figs 5-16. [3915'37''3'''N 926'37''7'''E, presso il capo
Carbonara, Sardegna, 508 m]
= Pandalus subtilirostris Riggio, 1905 [in Riggio, 1904-1905]: 283. [mare di Messina]
Plesionika gracilis (Zarenkov, 1971)
= Parapandals (sic) gracilis Zarenkov, 1971: 187; Fig. 3.1-3.10. [South China Sea, of Vietnam, 38 m]
Plesionika grahami Kensley, Tranter & Grifn, 1987: 315; Figs 20-21. [New South Wales, north-east of Port
Jackson, 3344'S 15149'E to 3342'S 15150'E, 405 m]
Plesionika grandis Dofein, 1902
= Plesionika spinipes var. grandis Dofein, 1902: 618; Plate 3, fgs 3-5. [Sagamibai]
Plesionika heterocarpus (A. Costa, 1871)
= Pandalus heterocarpus A. Costa, 1871: 89; Plate 2, fg. 3. [golfo di Napoli]
= Pandalus sagitarius A. Milne-Edwards, 1883: Plate 23. [383'N 1132'E (of Paris), 370 m]
= Pandalus longicarpus A. Milne-Edwards, 1883: Plate 25. [44N 917'E (of Paris), 156 m]
Plesionika holthuisi Crosnier & Forest, 1968: 1141; Fig. 7b-c. [Ombango stn 394, of Pointe Noir, Congo,
500-505 m]
Plesionika hsuehyui Chan, 2004: 300; Figs 3, 12-13. [Taiwan, Tai-Shi fshing port, NE coast, I-Lan County]
Plesionika hypanodon Dofein, 1902: 615; Plate 3, fg. 1. [Sagamibai]
Plesionika indica De Man, 1917
= Plesionika longipes var. indica De Man, 1917: 279. [540'S 13226'E, near the Kei-islands, 310 m]
Plesionika intermedia Chace, 1985: 72; Fig. 33. [Taon Straight, east of Negros, Philippines, 1029'45"N
12331'15"E, 472 m]
Plesionika izumiae Omori, 1971: 242; Figs 1-2; Plate 1. [3454.1'N 13824.7'E, 45 m]
Plesionika kensleyi Chace, 1985: 77; Figs 35-36. [Mindanao Sea, of Murcielagos Bay, Mindanao Philip-
pines, 845'30"N 12333'45"E, 309 m]
Plesionika laevis (A. Milne-Edwards, 1883)
= Heterocarpus laevis A. Milne-Edwards, 1883: Plate 28. [of Martinique, 1443'48''N 6111'25''W, 169
fms]
Plesionika laurentae Chan & Crosnier, 1991: 431; Figs 6, 24. [les Chesterfeld, 1925.49'S 15837.96'E: 215-
217 m]
Plesionika longicauda (Rathbun, 1901)
= Pandalus longicauda Rathbun, 1901: 117; Fig. 24. [Gulf of Mexico, 2842'30''N 8529'W, 88 fms]
Plesionika longidactylus Li & Komai, 2003: 262; Figs 2-5. [northern South China Sea, 1730'N 10930'E,
106 m]
Plesionika longipes (A. Milne-Edwards, 1881b)
= Pandalus longipes A. Milne-Edwards, 1881b: 15. [prs de Barbades, 204 brasses; lectotype designa-
tion by Forest & Holthuis, 1997]
Plesionika lophotes Chace, 1985: 81; Fig. 37. [Samar Sea between southeastern Masbate and Almagro Is-
land, Philippines, 1157'27"N 12410'42"E, 245 m]
Plesionika macropoda Chace, 1939: 37. [Bahia de Cochinos, Santa Clara Province, Cuba, 2207'N 8108'W,
195-235 fms]
Plesionika martia (A. Milne-Edwards, 1883)
= Pandalus martius A. Milne-Edwards, 1883: Plate 21. [Atlantic Ocean, roughly between 34 and
44N, 512-1290 m; see Forest & Holthuis, 1997]
= Plesionika uniproducta Spence Bate, 1888: 641 (partim); Plate 113, fg. 1. [95'S 3450'W, of Barra
Grande, 350 fms]
= Plesionika (Pandalus) sicherii Riggio, 1900: 20. [mare Siculo]
Plesionika miles (A. Milne-Edwards, 1883)
= Pandalus miles A. Milne-Edwards, 1883: Plate 18. [Dominica, 98 fms; see Forest & Holthuis, 1997]
Plesionika minor Calman, 1939: 199; Fig. 2. [John Murray Expedition, stn 75 (2510'48''N 5647'30''E to
2509'48''N 5647'30''E), Gulf of Oman, 210 m]
Plesionika multispinosa (Zarenkov, 1971)
= Parapandalus multispinosus Zarenkov, 1971: 185; Fig. 2(11-20). [of Easter Island, 80-150 m]
Plesionika narval (Fabricius, 1787)
= Astacus Narual Fabricius, 1787: 331. [in mari mediterraneo]
= P.[alemon] Pristis Risso, 1816: 105. [sur les fonds rocailleux, environs de Nice]
= Palaemon tarentinum O.G. Costa, 1844b: 9. [Mare di Taranto]
= Nisea formosa Risso, 1844: 95. [nomen nudum]
= Pandalus escatilis Stimpson, 1860a: 37. [prope insulam Madeirae, in profundis]
? = Pandalus stylopus A. Milne-Edwards, 1883: Plate 19. [3411'30''N 959'E (of Paris), 530 m]
Plesionika neon Komai & Chan, 2010b: 631; Figs 3-4, 7B. [Mozambique Channel, MAINBAZA st CP 3130,
2553'S 3307'E, 112-127 m]
Plesionika nesisi (Burukovsky, 1986)
= Heterocarpus nesisi Burukovsky, 1986: 62; Fig. 1. [East Pacifc sea-mount, 1334'N 12033'W, 800 m]
Plesionika ocellus (Spence Bate, 1888)
= Nothocaris ocellus Spence Bate, 1888: 657; Plate 114, fg. 3. [703'N 12148'E, near Samboangan,
Philippine Islands, 82 fms]
= Plesionika chacei Hayashi in Baba, Hayashi & Toriyama, 1986: 129; Fig. 20; unnumbered plate. [of
Kagoshima Prefecture, 107-110 m]
Plesionika orientalis Chace, 1985
? = Plesionika cotei Kote, 1903: 622; Plates 23-26. [im Indischen Ocean, von der Suahelikste, 630 m]
= Plesionika martia orientalis Chace, 1985: 84; Figs 38-39. [Sulu Archipelago, between Jolo and Ta-
witawi Islands, 548'00"N 12933'45"E, 490 m]
= Plesionika semilvis Spence Bate, 1888: 644. (partim)
Plesionika ortmanni Dofein, 1902: 616; Plate 3, fg. 2. [Sagamibai]
Plesionika pacifca Edmondson, 1952
= Plesionika pacifcus Edmondson, 1952: 67; Fig. 1. [of the north Kona coast of Hawaii, litle more
than 100 fms, from stomach of fsh, the opakapaka (Pristipomoides sp.)]
Plesionika parvimartia Chace, 1985: 93; Figs 42-43. [Samar Sea south of Tagapula Island, 1157'27"N
12410'42"E, 245 m]
Plesionika payeni Chan & Crosnier, 1997: 221; Figs 19, 39. [Tuamoto Islands, Mururoa, French Polynesia,
2147.6'S 13855.7'W, 200 m]
Plesionika persica (Kemp, 1925)
= Parapandalus persicus Kemp, 1925: 273; Fig. 8. [Gulf of Oman, 2344'30"N 5852'15"E, 261 fms]
= Parapandalus flipes Calman, 1939: 202; Fig. 3. [John Murray Expedition, Red Sea, stns 5 (1859'00''N
3913'30''E to 1849'18''N 3823'42''E), 500-0 m; 76B (2413'54''N 5903'30''E to 2416'42''N 5906'06''E),
South Arabian coast, 600-2500 m]
Plesionika philippinensis Chace, 1985: 97; Fig. 44. [Northern Surigao Strait between Cabugan Grande and
Hibuson Islands, Philippines, 1027'30"N 12519'15"E, 135 m]
Plesionika picta Chan & Crosnier, 1997: 223; Figs 20, 40. [Rurutu, Tubuai Islands, French Polynesia,
2227.8'S 15123.0'W, 570 m]
Plesionika polyacanthomerus Pequegnat, 1970: 97; Figs 4-13, 4-14. [Northeastern Gulf of Mexico, 2851'N
8847.5'W, 472-289 fms]
Plesionika poupini Chan & Crosnier, 1997: 189; Figs 1-2, 22. [Tubuai Islands, Raevavae, 2355.0'S
14740.0'W, 450 m]
Plesionika protati Chan & Crosnier, 1997: 219; Figs 16, 38. [Tahuata, Marquesas Islands, French Polynesia,
954.5'S 13907.9'W, 190 m]
Plesionika pumila Chace, 1985: 100; Figs 45-46. [Western part of Basilan Strait, northwest of Basilan Island,
Sulu Archipelago, 644'45"N 12148'E, 46 m]
Plesionika quasigrandis Chace, 1985: 104; Figs 47-48. [Davao Gulf about 2 miles (3.2 km) of Davao, Mind-
anao, Philippines, 702'N 12538'45"E, 247 m]
= Parapandalus yugniroi Timofeev, 1993: 42; Fig. 3. [Gulf of Aden, 1453'2N 5032'9E, 240-250 m]
Plesionika refexa Chace, 1985: 108; Fig. 49. [East of southern Luzon, northeast of San Bernardino Strait,
Philippines, 1256'24"N 12425'24"E, 494 m]
Plesionika rossignoli Crosnier & Forest, 1968: 1139; Fig. 6b. [Ombango stn 323 (of St. P. de Loanda, Angola,
9S 9E)]
Plesionika rostricrescentis (Spence Bate, 1888)
= Nothocaris rostricrescentis Spence Bate, 1888: 653; Plate 114, fg. 1. [549'15"S 13214'15"E, of the Ki
Islands, south of New Guinea, 140 fms]
Plesionika rubrior Chan & Crosnier, 1991: 433; Figs 7, 25-28. [Maiao, French Polynesia, 1738.6'S
15039.0'W, 320 m]
Plesionika rufomaculata Chan, 2004: 309; Figs 7, 16. [Loyalty Islands, 2041.8'S 1670.2'E, 282 m]
Plesionika sanctaecatalinae Wicksten, 1983b: 138; Figs 1-3. [7.3 miles, 17 true from Long Point Light, San-
ta Catalina Island, California, 3333'29''N 1820'42''W 3330'12''N 11817'28''W, 886 m]
Plesionika scopifera Chan, 2004: 311; Figs 8-9, 17. [Chesterfeld Islands, 1948.6'S 15829.1'E, 260-270 m]
Plesionika semilaevis Spence Bate, 1888
= Plesionika semilvis Spence Bate, 1888; Plate 113, fg. 3. [Moro Gul east of Basilan Strait, Philip-
pines, 647'N 12228'E, 457 m; lectotype designation by Chace, 1985]
Plesionika serratifrons (Borradaile, 1900)
= Pandalus (Parapandalus) serratifrons Borradaile, 1900: 411; Plate 37, fgs 8a-8d. [Blanche Bay, New
Britain, 50-100 fms]
= Pandalus (Parapandalus) tenuipes Borradaile, 1900: 412; Plate 37, fg. 9; nec Smith, 1881. [Blanche
Bay, New Britain; DEntrecasteaux Group, British New Guinea]
Plesionika simulatrix Chace, 1985: 125; Fig. 57. [Samar Sea, east of Masbate, Philippines, 1213'15"N
12405'03"E, 216 m]
Plesionika sindoi (Rathbun, 1906)
= Pandalus sindoi Rathbun, 1906: 915; Plate 21, fg. 4. [Albatross stn 3998, vicinity of Kauai Island
(2156'25''N 15948'35''W), 235-228 fms]
Plesionika spinensis Chace, 1985: 128; Figs 58-59. [Western Mindanao Sea, of Murcielagos Bay, Philip-
pines, 845'30"N 12333'45"E, 309 m]
Plesionika spinidorsalis (Rathbun, 1906)
= Pandalus spinidorsalis Rathbun, 1906: 917; Plate 21, fg. 5. [Albatross stn 3986 (2204'35''N
15918'40''W, 55-362 fms), Vicinity of Kauai Island]
Plesionika spinipes Spence Bate, 1888: 646; Plate 113, fg. 2. [154'0"S 14639'40"E, north of New Guinea,
150 fms]
Plesionika sufusa Chan, 2004: 305; Figs 5, 14. [New Caledonia, 1857.8'S 16314'E, 425 m]
Plesionika taiwanica Chan & Yu, 2000: 120; Figs 1-3. [Taiwan, southwestern coast, fshing port at Tung-
kang, Pingtong County, about 150 m]
Plesionika tenuipes (Smith, 1881)
= Pandalus tenuipes Smith, 1881: 441. [Fish Hawk stns 870 (4002'36''N 7022'58''W, 155 fms), 871
(4002'54''N 7023'40''W, 115 fms), 873 (4002'00''N 7057'00''W, 100 fms), 877 (3956'00''N 7054'18''W,
126 fms), 880 (3948'30''N 7054'00''W, 252 fms); all of Block Island, 100-252 fms]
Plesionika trispinus Squires & Barragan, 1976: 113; Figs 1-2. [of Pizarro, Colombia, 0510'N 7725'W, 257-
275 m]
Plesionika unicarinatus (Borradaile, 1915b)
= Heterocarpus unicarinatus Borradaile, 1915b: 208. [Providence Island, 637-665 fms]
Plesionika unidens Spence Bate, 1888: 648; Plate 113, fg. 4. [154'00"S 14639'48"E, north of New Guinea,
150 fms]
= Plesionika afnis Alcock & Anderson, 1899: 285. [Investigator stn 236 (Andaman Sea, 1359'N 93E),
172-303 fms]
Plesionika williamsi Forest, 1964: 620; Figs 1-4. [439'N 246'W, 300-400 m]
? = Plesionika dentirostris Tung, Wang & Li, 1988: 20; Fig. 1. [East China Sea, 400 m]
= Plesionika crosnieri Burukovsky, 1992: 145; Figs 1-4; nec Chan & Yu, 1991b. [2539'S 8651'E, 583-
600 m]
= Plesionika alaini Burukovsky, 1993c: 18. [nomen novum for Plesionika crosnieri Burukovsky, 1992]
Plesionika willisi (Pequegnat, 1970)
= Parapandalus willisi Pequegnat, 1970: 87; Figs 4.9-4.10. [Northeastern Gulf of Mexico, 2927.6'N
8646.5'W, 210 fms]
Plesionika yui Chan & Crosnier, 1991: 428; Figs 4b, 5c, e, g-h, 23. [Tong-Kong, Ping-Tong County, SW
Taiwan]
Procletes Spence Bate, 1888
= Procletes Spence Bate, 1888 (type species Procletes biangulatus Spence Bate, 1888 (junior subjective
synonym of Dorodotes levicarina Spence Bate, 1888), designated by Holthuis, 1955b, gender mascu-
line)
= Heterocarpoides De Man, 1917 (type species Dorodotes levicarina Spence Bate, 1888, by monotypy,
1957)
Procletes levicarina (Spence Bate, 1888) (Fig. 56)
= Dorodotes levicarina Spence Bate, 1888: 680; Plate 112, fg. 5. [959'S 13942'E, Arafura Sea, south of
Papua, 28 fms]
= Procletes biangulatus Spence Bate, 1888: 884; Plate 127, fg. 4. [Challenger stn 191, 541'0''S 1344'30''E,
of the Arrou islands, 800 fms]
? = Heterocarpus (Heterocapoides) [sic] glabrus Zarenkov, 1971: 193; Fig. 4(16-27). [South-China Sea, of
S Vietnam, 75 m]
Pseudopandalus Crosnier, 1997
= Pseudopandalus Crosnier, 1997 (type species Pseudopandalus curvirostris Crosnier, 1997, by original
Pseudopandalus curvirostris Crosnier, 1997: 172; Figs 1-3. [Nouvelle-Caldonie, 2258.9'S 16720.2'E, 508-
514 m]
Stylopandalus Coutire, 1905c
= Stylopandalus Coutire, 1905c (type species Pandalus (Stylopandalus) Richardi Coutire, 1905c, by
Stylopandalus richardi (Coutire, 1905a)
= Pandalus (Stylopandalus) Richardi Coutire, 1905c: 1115. [full description in Coutire, 1905a; Prin-
cesse-Alice stns west of Madeira, 3218'N 2358'W, 2000-0 m; and Canary Islands, 2743'N 1828'W,
3000-0 m; see Chace, 1985]
= Plesionika nana Murray & Hjort, 1912: 585. [Spain to Newfoundland; nomen nudum]
= Parapandalus zur strasseni Balss, 1914a: 597. [1008'S 9714'E, 0-2400 m; 324'S 5838'E, 0-2000 m]
= Pandalus (Plesionica) gracilis Borradaile, 1915b: 208; nec Stimpson, 1860a. [Western Indian Ocean,
200 fms]
Family THALASSOCARIDIDAE Spence Bate, 1888
Chlorotocoides Kemp, 1925
= Chlorotocoides Kemp, 1925 (type species Chlorotocus spinicauda De Man, 1902, by original designa-
tion and monotypy, gender masculine; name placed on the Ofcial List of Generic Names in Zool-
Chlorotocoides spinicauda (De Man, 1902)
= Chlorotocus spinicauda De Man, 1902: 856; Plate 26, fg. 59 [Ternate]
= Thalassocaris maldivensis Borradaile, 1915b: 208. [Maldive Islands]
Fig. 56. Procletes levicarina (Spence Bate, 1888). Photo by Tin-Yam Chan.
Thalassocaris Stimpson, 1860a
= Regulus Dana, 1852a (type species Regulus lucidus Dana, 1852a, designated by Kingsley, 1880; in-
valid junior homonym of Regulus Cuvier, 1880 (Aves); name placed on the Ofcial List of Rejected
and Invalid Generic Names in Zoology in Opinion 470 in 1957)
= Thalassocaris Stimpson, 1860a (nomen novum for Regulus Dana, 1852a, gender masculine; name
Thalassocaris crinita (Dana, 1852a) (Fig. 57)
= Regulus crinitus Dana, 1852a: 27. [in mari Suluensi]
= Thalassocaris afnis Borradaile, 1915b: 208. [Maldives; Saya de Malha; lectotype designation by
Gopala Menon & Williamson, 1971]
= Thalassocaris crinita var.? De Man, 1920b: 100; Plate 10, fgs 23-23c. [Saleyer-anchorage and sur-
roundings, including Pulu Pasi Tanete, near the North point of Saleyer-island, up to 36 m]
Thalassocaris lucida (Dana, 1852a)
= Regulus lucidus Dana, 1852a: 27. [in mari Pacifco, prope insulas Ladrone]
= Thalassocaris dan Spence Bate, 1888: 683; Plate 117, fg. 1. [Fii Islands, at the surface]
= Thalassocaris stimpsoni Spence Bate, 1888: 684; Plate 117, fg. 2. [Japan, near Yokohama, surface; Fii
Islands, surface]
Thalassocaris obscura Gopala Menon & Williamson, 1971: 36; Figs 1c, 4, 5c, 6e [Arabian Sea, 0900'N
7520'E, 200-0 m]
Superfamily PHYSETOCARIDOIDEA Chace, 1940
Family PHYSETOCARIDIDAE Chace, 1940
Physetocaris Chace, 1940
= Physetocaris Chace, 1940 (type species Physetocaris microphthalma Chace, 1940, by monotypy, gen-
der feminine)
Physetocaris microphthalma Chace, 1940: 196, Figs 62-63. [Bermuda Oceanographic Expedition Net 798
(3212'N 6436'W), 600 fms]
Fig. 57. Thalassocaris crinita (Dana, 1852). Photo by Arthur Anker.
Superfamily CRANGONOIDEA Haworth, 1825
Family CRANGONIDAE Haworth, 1825
Aegaeon Agassiz, 1846 [in Agassiz, 1842-1846]
= Egeon Bosc, 1813 (type species Cancer cataphractus Olivi, 1792, by monotypy, gender masculine;
invalid junior homonym of Egon de Montfort, 1808 (Protozoa); name placed on the Ofcial Index of
Rejected and Invalid Generic Names in Zoology in Opinion 434 in 1956)
= Aegaeon Agassiz, 1846 [in Agassiz, 1842-1846] (nomen novum for Egeon Bosc, 1813, gender mascu-
line; see Holthuis, 1993a)
Aegaeon boschii (Christofersen, 1988)
= Pontocaris boschii Christofersen, 1988: 49; Figs 2-3. [37S 56W, Argentina, province of Buenos
Aires]
Aegaeon cataphractus (Olivi, 1792)
= Cancer cataphractus Olivi, 1792: 50; Plate 3, fg. 1. [la Golfo e delle Lagune di Venezia]
= E.[geon] loricatus Risso, 1827: 58; Plate 1, fg. 3. [golfe de Nice, sur les fonds rocailleux]
Aegaeon lacazei (Gourret, 1887a)
= Crangon Lacazei Gourret, 1887a: 1033. [rgion nord-ouest du golfe de Marseille, 70-80 m]
= geon Brendani Kemp, 1906b: 299. [about 50 miles W. by N. of Inishtearaght Lighthouse, Co.
Kerry, 320-370 fms]
= Pontocaris habereri Dofein, 1902: 620; Fig. A; Plate 1, fgs 4-5. [Sagamibai]
Aegaeon orientalis Henderson, 1893
= geon orientalis Henderson, 1893: 446; Plate 40, fgs 16-17. [Gulf of Martaban]
= geon rugulosum Borradaile, 1915b: 210. [Western Indian Ocean]
Aegaeon rathbuni De Man, 1918b
= Aegeon Rathbuni De Man, 1918b: 304. [Pulu Kaniungan Ketjil, 11 m]
Argis Kryer, 1843
= Argis Kryer, 1843 (type species Crangon lar Owen, 1839, by monotypy, gender feminine; name
= Nectrocrangon Brandt, 1850 (type species Crangon lar Owen, 1839, by monotypy, gender feminine)
= Nectocrangon Brandt, 1851 (type species Crangon lar Owen, 1839, by monotypy, gender feminine;
name placed on the Ofcial Index of Rejected and Invalid Generic Names in Zoology in Opinion
470 in 1957)
Argis alaskensis (Kingsley, 1883)
= Nectocrangon alaskensis Kingsley, 1883: 128. [Marmot Island, Kodiac Archipelago, Alaska]
Argis californiensis (Rathbun, 1902a)
= Nectocrangon californiensis Rathbun, 1902a: 892. [of Santa Catalina Island, California, 80 fms]
Argis crassa (Rathbun, 1899)
= Nectocrangon crassa Rathbun, 1899: 556. [574'N 17024'W, 26 fms]
= Nectocrangon sadoensis Yokoya, 1933: 40; Fig. 21. [Sy-Maru stn 600, north of Is. Sado, 168 m]
Argis dentata (Rathbun, 1902a)
= Nectocrangon dentata Rathbun, 1902a: 892. [of Sitkalidak Island, Alaska, 69 fms]
= Argis argilicola Kryer, 1843: 256. [nomen nudum]
Argis hozawai (Yokoya, 1939)
= Nectocrangon hozawai Yokoya, 1939: 276; Fig. 9. [about 5 miles of Ozaki, 130 fms]
= Nectocrangon lar kobjakovi Vinogradov, 1950: 221. [Sea of Japan, southern Sea of Okhotsk and
southern Kuril Islands]
Argis lar (Owen, 1839)
= Crangon lar Owen, 1839: 88; Plate 28, fg. 1. [between Bristol Bay and the Pribilof islands, 25.5 fms;
neotype selection by Squires, 1964]
Argis levior (Rathbun, 1902a)
= Nectocrangon levior Rathbun, 1902a: 892. [Admiralty Inlet, Puget Sound, 40 fms]
Argis ochotensis ochotensis Komai, 1997b
= Argis ochotensis Komai, 1997b: 149; Figs 12-15. [Okhotsk Sea, Kitami, Yamato Bank, 4439.1'N
14418.3'E, 815-820 m]
Argis ochotensis kamtschatica Sokolov, 2001: 1056; Figs 3-5, 6f-h. [Ochotsk Sea, 5245'N 15443'E, 245 m]
Argis ovifer (Rathbun, 1902a)
= Nectocrangon ovifer Rathbun, 1902a: 892. [of the Trinity Islands, Alaska, 159 fms]
Argis toyamaensis (Yokoya, 1933)
= Nectocrangon toyamaensis Yokoya, 1933: 39; Fig. 20. [Sy-Maru stn 591, Toyama Bay, 311 m]
Crangon Fabricius, 1798
= Crangon Fabricius, 1798 (type species Cancer Crangon Linnaeus, 1758, by absolute tautonymy,
gender feminine; junior homonym of Crangon Weber, 1795 (invalid senior synonym of Alpheus Fab-
ricius, 1798); name conserved under the plenary powers of the ICZN and placed on the Ofcial List
of Generic Names in Zoology in Opinion 334 in 1955)
= Crago Lamarck, 1801 (type species Cancer Crangon Linnaeus, 1758, by monotypy, gender mascu-
line; name placed on the Ofcial Index of Rejected and Invalid Generic Names in Zoology in Opin-
ion 334 in 1955)
= Crangonus Rafnesque, 1815 (nomen novum for Crangon Fabricius, 1798, gender masculine; name
1955)
= Steiracrangon Kinahan, 1862 (type species Crangon Allmanni Kinahan, 1860c, designated by Fow-
ler, 1912, gender feminine)
Crangon afnis De Haan, 1849 [in De Haan, 1833-1850]: 183. [Bay of Kob, Japan, 3439'N 13514'E, 8 fms;
neotype designation by Hayashi & J.N. Kim, 1998]
= Crangon consobrinus De Man, 1906: 401. [Inland Sea of Japan, deep water]
Crangon alaskensis Lockington, 1877a
= Crangon Alaskensis Lockington, 1877a: 34. [Mutiny Bay, Alaska]
= Crangon alaskensis elongata Rathbun, 1902a: 888. [of Santa Barbara, California, 29 fms]
Crangon alba Holmes, 1900: 174. [Monterey]
Crangon allmanni Kinahan, 1860c
= Crangon Allmanni Kinahan, 1860c: 28; unnumbered text Figs [Zonam corallinam Maris Hibernici
juxta, Bray, comitat: Wicklow]
Crangon amurensis Branikov, 1907
= Crangon septemspinosa f. amurensis Branikov, 1907: 68. [Amuriskiy Liman]
= Crangon septemspinosa f. anivensis Branikov, 1907: 68. [Aniva Bay]
Crangon capensis Stimpson, 1860a: 24. [in sinu Simons Bay, promont. Bonae Spei, prof. 12 org]
Crangon cassiope De Man, 1906: 402. [Inland Sea of Japan, deep water]
Crangon crangon (Linnaeus, 1758)
= Cancer Crangon Linnaeus, 1758: 632. [Mare Balthico]
= Crangon vulgaris Fabricius, 1798: 410. [mari balthico]
= Crangon Rubro punctatus Risso, 1816: 83. [environs de Nice, dans les endroits sabloneux]
= Cancer schillinus Nardo, 1847: 7. [Trovato nel golfo in estate]
= Crangon maculosus Rathke, 1837: 366. [Cap Porthenion und bei Sevastopol]
= Crangon maculosus forma typica Czerniavsky, 1884: 71. [Sevastopol; Odessa; Theodosia]
= Crangon maculosus forma brevirostris Czerniavsky, 1884: 72. [sinus Sevatopol et Odessa]
= Crangon maculosus var. suchumica Czerniavsky, 1884: 72. [sin. Suchum]
= Steiracrangon orientalis var. longicauda forma intermedia Czerniavsky, 1884: 74. [sinus Sevastopol;
Orianda]
= Steiracrangon orientalis var. brevicauda Czerniavsky, 1884: 75. [sinus Sevastopol; Orianda]
= Crangon crangon typicus Dofein, 1900: 325. [Type locality not indicated]
= Crangon crangon f. mediterranea Branikov, 1907: 70. [Naples]
= Crangon crangon f. typica Branikov, 1907: 70. [White Sea; Norway]
Crangon dalli Rathbun, 1902a: 889. [Bering Sea, of Cape Seniavin, Alaska, 30 fms]
Crangon franciscorum franciscorum Stimpson, 1856
= Crangon Franciscorum Stimpson, 1856: 89. [sandy coves around the bay (San Francisco)]
Crangon franciscorum angustimana Rathbun, 1902a: 889. [of Chuckanuts Island, Bellingham Bay, Wash-
ington, 11 fms]
Crangon hakodatei Rathbun, 1902b: 42; Fig. 15. [Hakodate, Hokkaido]
Crangon handi Kuris & Carlton, 1977: 543; Figs 1, 2a, d, g. [Horseshoe Cove, Bodega Head, Sonoma
County, 3818'30''N 1234'W]
Crangon holmesi Rathbun, 1902a: 888. [Of Wilmington, California, 27 fms]
Crangon lockingtonii Holmes, 1904
= Crangon Lockingtonii Holmes, 1904: 312; Plate 35, fgs 14-16. [Magdalen Bay, Lower California]
Crangon nigricauda Stimpson, 1856: 89. [California, deeper water]
Crangon nigromaculata Lockington, 1877a: 34. [San Diego, 6 fms]
Crangon propinquus Stimpson, 1860a: 25. [Lake Saroma, Hokkaido; neotype selection by Hayashi & J.N.
Kim, 1999]
= Steiracrangon orientalis var. longicauda forma pacifca Czerniavsky, 1884: 74. [Sachalin (ocean. Pa-
cifc); colonia Dui (Itere Amurensi)]
= Crangon vulgaris var. Shidlovskii Ostroumov, 1896: 81. [Vladivostok]
Crangon septemspinosa Say, 1818
= C.[rangon] septemspinosus Say, 1818: 246. [bay shores and inlets of the sea]
Crangon uritai Hayashi & J.N. Kim, 1999: 86; Figs 13-16. [Dadaepo, Pusan, Krea, 1 m]
Lissocrangon Kuris & Carlton, 1977
= Lissocrangon Kuris & Carlton, 1977 (type species Crangon stylirostris Holmes, 1900, by original
Lissocrangon stylirostris (Holmes, 1900)
= Crangon stylirostris Holmes, 1900: 174. [Trinidad, Humboldt County, California]
Lissosabinea Christofersen, 1988
= Lissosabinea Christofersen, 1988 (type species Sabinea tridentata Pequegnat, 1970, by original des-
ignation, gender feminine)
Lissosabinea armata Komai, 2006a: 44; Figs 7-9. [New Caledonia, 2444.24'S 17008.01'E, 770-830 m]
Lissosabinea arthuri Taylor & Collins, 2009: 177; Figs 3-4. [Australia, Vic., 3809.80'S 14941.71'E -
3810.11'S 14941.01'E, 260-265 m]
Lissosabinea beresfordi Taylor & Collins, 2009: 176; Figs 1-2, 8A-B. [Australia, Southern Ocean, south of
Tasmania, 4359.5'S 14732.76'E - 4359.7'S 14733.80'E, 370-410 m]
Lissosabinea ecarina Komai, 2006a: 49; Figs 10-12. [Indonesia, Kai Islands, 0530'N 13252'E, 336-346 m]
Lissosabinea indica (De Man, 1918b)
= Sabinea indica De Man, 1918b: 304. [70'S 12034'.5E, North of Tanah Djampeah Island, 400 m]
Lissosabinea lynseyae Taylor & Collins, 2009: 186; Figs 15-6, 8C. [Australia, WA, of Bunbury, 3300.5'S
11459.26'E - 3300.11'S 11434.50'E, 421-414 m]
Lissosabinea tridentata (Pequegnat, 1970)
= Sabinea tridentata Pequegnat, 1970: 115; Figs 4-16, 4-17. [Southeastern Gulf of Mexico, 2458'N
8417'W, 214 fms]
Lissosabinea unispinosa Komai, 2006a: 53; Figs 13-16. [New Caledonia, 1854.8'S 16322.2'E, 465 m]
Mesocrangon Zarenkov, 1965
= Mesocrangon Zarenkov, 1964 (unavailable name under Art. 13.3)
= Mesocrangon Zarenkov, 1965 (type species Crangon intermedius Stimpson, 1860a, by original desig-
Mesocrangon intermedia (Stimpson, 1860a)
= Crangon intermedius Stimpson, 1860a: 25. [mari Beringiano prope promontorium Chepoonski,
ad prof. 40 org]
= Crangon tenuifrons Kingsley, 1883: 128; Plate 1, fg. 10. [Marmot Island, Kodiac Archipelago, Alaska]
= Sclerocrangon intermedius var. bidentata Balss, 1914b: 65. [Sagamibai]
Mesocrangon munitella (Walker, 1898)
= Crangon munitellus Walker, 1898: 275; Plate 16, fg. 1. [Puget Sound, pacifc coast of North America]
Mesocrangon volki (Birstein & Vinogradov, 1953)
= Sclerocrangon volki Birstein & Vinogradov, 1953: 217; Fig. 1. [near Bering Island, Bering Sea, 93-
112 m]
Metacrangon Zarenkov, 1965
= Metacrangon Zarenkov, 1964 (unavailable name under Art. 13.3)
= Metacrangon Zarenkov, 1965 (type species Crangon variabilis Rathbun, 1902a, by original designa-
Metacrangon acclivis (Rathbun, 1902a)
= Crangon acclivis Rathbun, 1902a: 890. [of Santa Cruz Island, California, 266 fms]
Metacrangon agassizii (Smith, 1882)
= Cheraphilus Agassizii Smith, 1882: 32; Plate 7, fgs 4-5a. [Blake stns 317, 3157'0''N 7818'35''W, 333
fms; 326, 3342'15''N 760'50''W, 464 fms; 329, 3439'40''N 7514'40''W, 603 fms; 332, 3545'30''N
7448'0''W, 263 fms; of Block Island, 3946'N 7110'W, 500 fms]
Metacrangon australis Komai & Taylor, 2010: 46; Figs 1-3. [Australia, Tasmania, southwestern Pacifc,
west of Macquarie Island, 5442.42'S 15845.12'E 5441.36'S 15843.12'E, 700-900 m]
Metacrangon bahamondei Retamal & Gorny, 2003: 85; Fig. 1. [5544'00''S 6614'05''W, sur de la Isla Nueva,
Tierra del Fuego, 746 m]
Metacrangon bellmarleyi (Stebbing, 1914b)
= Sclerocrangon bellmarleyi Stebbing, 1914b: 29; Plate 10. [Cape Natal N by E 24 miles, Natal, 440 fms]
Metacrangon cornuta Komai & Komatsu, 2009: 515; Figs 2-4, 19G. [of Kushi-ro, eastern Hokkaido,
4230.3'N 14450.5'E to 4230.6'N 1442.2'E, 1535-1543 m]
Metacrangon crosnieri Komai, 1997c: 662; Figs 1c, 5-6. [Madagascar, of Cape Ambre, 1302'S 4802'E,
1000-1525 m]
Metacrangon haona Komai & Ahyong, 2011: 78; Figs 1-3. [Pukaki Rise, Campbell Plateau, 4921.50'S
17153.00'E, 353 m]
Metacrangon hikurangi Komai & Ahyong, 2011: 83; Figs 4-6. [Hikurangi Margin, east coast of North Is-
land, New Zealand, 4002.22-02.14'S 17748.06-47.97'E, 1171-1172 m]
Metacrangon holthuisi Komai, 2010a: 370; Figs 1-2. [west of Sunosaki, Tateyama, Sagami Sea, 3456.46'N
13933.12'E, 1039-1300 m]
Metacrangon jacqueti (A. Milne-Edwards, 1881a)
= Pontophilus Jacqueti A. Milne-Edwards, 1881a: 933. [golfe de Gascogne, 4448'30''N 70'30''W,
5100 m]
Metacrangon knoxi (Yaldwyn, 1960)
= Sclerocrangon knoxi Yaldwyn, 1960: 35; Fig. 7. [Chatham Rise, 4340'S 17928'E, 220 fms]
Metacrangon laevis (Yokoya, 1933)
= Crangon laevis Yokoya, 1933: 36; Fig. 16. [Sy-Maru stn 638, west of Tubaki-zaki, Aomori-ken,
73 m]
Metacrangon longirostris (Yokoya, 1933)
= Crangon longirostris Yokoya, 1933: 36; Fig. 17. [Sy-Maru stn 381, east of Owase, Mie-ken, 329 m]
Metacrangon miyakei J.N. Kim, 2005: 246; Figs 5-6. [Sea of Japan, of Taiza, Tango Peninsula, 110 m]
Metacrangon monodon (Birstein & Vinogradov, 1951)
= Sclerocrangon monodon Birstein & Vinogradov, 1951: 357; Figs 1-3. [4th Kuril Straits, Kuril Islands,
NW Pacifc, 630 m]
Metacrangon munita (Dana, 1852a)
= Crangon munitus Dana, 1852a: 20. [in freto Pugetensi]
Metacrangon nipponensis (Yokoya, 1933)
= Crangon nipponensis Yokoya, 1933: 38; Fig. 19. [Sy-Maru stn 22, east of Siwoya-zaki, 539 m]
Metacrangon ochotensis (Kobjakova, 1955)
= Sclerocrangon ochotensis Kobjakova, 1955: 235; Fig. 1. [Ochotsk Sea, Kuril Island area, of Ekaterina
Straits, 2850 m]
Metacrangon procax (Faxon, 1893)
= Sclerocrangon procax Faxon, 1893: 199. [Albatross stns 3380 (0403'00''N 8131'00''W, 899 fms); 3418
(1633'00''N 9952'30''W, 660 fms); 3435 (2648'00''N 11045'20''W, 859 fms); 3436 (2703'40''N
11053'40''W, 905 fms]
= Crago lomae Schmit, 1921: 100; Plate 12, fgs 3-4. [Of Point Loma, California, 525-541 fms]
Metacrangon proxima J.N. Kim, 2005: 242; Figs 1-3. [Pacifc coast of central Japan, Amadai-ba, Sagami Bay,
280 m]
Metacrangon poorei Komai & Taylor, 2010: 49; Figs 4-5. [Australia, of southeastern Victoria, 3953.76'S
14903.39'E, 1608 m]
Metacrangon rau Komai & Ahyong, 2011: 91; Figs 7-10. [west of Northland, North Island, 3519.99'S
17219.99'E, 1029-1074 m]
Metacrangon richardsoni (Yaldwyn, 1960)
= Sclerocrangon richardsoni Yaldwyn, 1960: 39; Fig. 8. [4142'30''S 1759'E, circa 550 fms]
Metacrangon robusta (Kobjakova, 1935)
= Sclerocrangon robustus Kobjakova, 1935: 85; Figs 1-2. [Japanischen Meer von der Bucht Peters des
Groen nach Norden bis zu 50 n. Br., 70-1380 m; Ochotskisches Meer im nordwestlichen Teil, in
der Umgegend von der St.-Jonas-Insel, 50-515 m]
Metacrangon similis Komai, 1997c: 675; Figs 1h, 11-12. [Japan, of Owase, Kumano-nada, 3405.8'N
13635.9'E, 369-412 m]
Metacrangon sinensis Fujino & Miyake, 1970a: 269; Figs 10-15. [3701.5'N 12357.9'E, 74 m]
Metacrangon spinidorsalis Komai & Taylor, 2010: 52; Figs 6-10. [Australia, Western Australia, of Point
Hillier, 3522.54'S 11712.25'E 3522.54'S 11712.25'E, 539 m]
Metacrangon spinirostris (Rathbun, 1902a)
= Crangon spinirostris Rathbun, 1902a: 891. [North of Unalaska, 399 fms, Albatross stn 3329
(5356'50''N 16708'15''W)]
Metacrangon spinosissima (Rathbun, 1902a)
= Crangon spinosissima Rathbun, 1902a: 891. [Of Point Arena, California, 51 fms]
Metacrangon teina Komai & Ahyong, 2011: 95; Figs 11-15. [Challenger Plateau, 3932.61'S 16942.87'E,
636-634 m]
Metacrangon trigonorostris Yokoya, 1933
= Crangon trigonorostris Yokoya, 1933: 37; Fig. 18. [Sy-Maru stn 73, south of Siwoya-zaki, 390 m]
Metacrangon variabilis variabilis (Rathbun, 1902a)
= Crangon variabilis Rathbun, 1902a: 890. [Of North Head, Akutan Island, Alaska, 72 fms]
Metacrangon variabilis asiaticus (Kobjakova, 1955)
= Sclerocrangon variabilis asiaticus Kobjakova, 1955: 236; Fig. 2. [SE of Shikotan Island, Kuril Islands,
136-414 m]
Neocrangon Zarenkov, 1965
= Neocrangon Zarenkov, 1964 (unavailable name under Art. 13.3)
= Neocrangon Zarenkov, 1965 (type species Crangon communis Rathbun, 1899, by original designa-
Neocrangon abyssorum (Rathbun, 1902a)
= Crangon abyssorum Rathbun, 1902a: 890. [Bering Sea, southwest of Pribilof Islands, 1771 fms]
Neocrangon communis (Rathbun, 1899)
= Crangon communis Rathbun, 1899: 556. [574'20''N 17052'30''W, 51 fms]
Neocrangon geniculata (Yokoya, 1933)
= Crangon geniculata Yokoya, 1933: 35; Fig. 15. [Sy-Maru stn 213, Kii Strait, 353 m]
Neocrangon joloensis (De Man, 1929)
= Crangon joloensis De Man, 1929: 131; Figs 9-9b. [of the Jolo Islands, 16 fms]
Neocrangon resima (Rathbun, 1902a)
= Crangon resima Rathbun, 1902a: 889. [Of San Diego, California, 124 fms]
= Crago zacae Chace, 1937b: 136; Fig. 9. [east of Cedros Island, 2813'N 11507'W, 44 fms]
Neocrangon sagamiensis (Balss, 1913b)
= Crangon (Crangon) sagamiense Balss, 1913b: 237. [Fukuura, Sagamibai]
= Neocrangon orientalis Han & Li, 2009: 65; Figs 1-2. [East China Sea, 3031.0'N 12756.5'E, 365-395 m]
Notocrangon Coutire, 1900d
= Notocrangon Coutire, 1900d (type species Crangon antarcticus Pfefer, 1887, by monotypy, gender
Notocrangon antarcticus (Pfefer, 1887) (Fig. 58)
= Crangon antarcticus Pfefer, 1887: 45; Plate 1, Figs 1-21. [Sd-Georgien]
= Crangon (Notocrangon) antarcticus var. gracilis Borradaile, 1916: 89. [Terra Nova Expedition stns 294
(7425'S 16647'W, 0-1750 m); 314 (5 miles N of Inaccesible Island, McMurdo Sound, 406-411 m); 316
(of Glacier Tongue, about 8 miles N of Hut Point, McMurdo Sound, 348-457 m); 338 (7713'S
16418'E, 379 m); 339 (775'S 16417'E, 256 m); 348 (of Barne Glacier, McMurdo Sound, 366 m); 355
(7746'S 1668'E, 547 m)]
Paracrangon Dana, 1852a
= Paracrangon Dana, 1852a (Paracrangon echinatus Dana, 1852a, by monotypy, gender feminine;
name placed on the Ofcial List of Generic Names in Zoology in 470 in 1957)
Paracrangon abei Kubo, 1937b: 3; Figs 2-3. [Kumano-nada, of Nagasima, Mie Prefecture, about 170 fms]
Paracrangon areolata Faxon, 1893: 200. [Albatross stns 3424 (2115'00''N 10623'00''E), 676 fms; 3425
(2119'00''N 10624'00''E), 680 fms]
Paracrangon australis Hanamura, Wadley & Taylor, 1999: 311; Figs 1-3. [Tasmania, approx. 84 km SSE of
South East Cape, 4416'S 14720'E, 987 m]
Paracrangon echinata Dana, 1852a
= Paracrangon echinatus Dana, 1852a: 20. [in freto Pugetensi]
Paracrangon furcata Kubo, 1937b: 7; Figs 4-5. [Kumano-nada, of Nagasima, Mie Prefecture, about 170
fms]
Paracrangon okutanii Oh & Takeda, 1986: 76; Figs 2-3. [3504'N 13928.4'E, 770-930 m]
Paracrangon ostlingos Komai & J.N. Kim, 2004: 531; Figs 1-13. [Pacifc coast of Japan, of Taito-saki, Boso
Peninsula, 3509.4'N 14049.0'E, 311-323 m]
Fig. 58. Notocrangon antarcticus (Pfefer, 1887). Photo by Cedric d'Udekem d'Acoz.
Parapontocaris Alcock, 1901
= Parapontocaris Alcock, 1901 (type species Crangon bengalensis Wood-Mason in Wood-Mason &
Alcock, 1891c, designated by Holthuis, 1955b, gender feminine)
Parapontocaris andamanensis (Wood-Mason in Wood-Mason & Alcock, 1891c)
= Crangon andamanensis Wood-Mason in Wood-Mason & Alcock, 1891c: 360. [Andaman Sea,
1131'40''N 9246'40''E, 188-220 fms]
Parapontocaris aspera Chace, 1984: 30; Figs 9-11. [Albay Gulf, east of southern Luzon, Philippines,
1310'21''N 12359'54'', 368 m]
Parapontocaris bengalensis (Wood-Mason in Wood-Mason & Alcock, 1891c)
= Crangon bengalensis Wood-Mason in Wood-Mason & Alcock, 1891c: 360. [Bay of Bengal, 1556'50''N
8130.5'E, 240-276 fms]
Parapontocaris caribbaea (Boone, 1927)
= Aegeon caribbaeus Boone, 1927: 125; Fig. 28. [north of Glover Reef, 484 fms]
Parapontocaris levigata Chace, 1984: 34; Figs 12-14. [Balayan Bay, southern Luzon, Philippines, 1351'30''N
12050'30''E, 291 m]
Parapontocaris vicina (Dardeau & Heard, 1983)
= Pontocaris vicina Dardeau & Heard, 1983: 10; Figs 2g, 4-7. [Straits of Florida, 2335'N 8022'W, 457 m]
Parapontophilus Christofersen, 1988
= Parapontophilus Christofersen, 1988 (type species Pontophilus gracilis Smith, 1882, by original des-
ignation, gender masculine)
Parapontophilus abyssi (Smith, 1884)
= Pontophilus abyssi Smith, 1884: 363. [Albatross stns 7023/7025, 3756'20''N 7057'30''W, 1917 fms;
7024, 3740'30''N 7037'30''W, 2221 fms]
= Pontophilus gracilis Spence Bate, 1888: 487 (partim); Plate 87; nec Smith, 1882. [Challenger stn 133,
3541'S 2055'W, near Tristan da Cunha, 1900 fms only]
= Pontophilus challengeri Ortmann, 1893: 49. [nomen novum for Pontophilus gracilis Spence Bate,
1888]
= Pontophilus batei Faxon, 1893: 200. [nomen novum for Pontophilus gracilis Spence Bate, 1888]
Parapontophilis caledonicus Komai, 2008: 313; Figs 24-25. [New Caledonia, 2035.9'S 16458.3'E, 400-
500 m]
Parapontophilus cornutus Komai, 2008: 292; Figs 10-12. [Austral Islands, Tubuai Island, 2321.3'S
14933.9'W, 500-200 m]
Parapontophilus cyrton Komai, 2008: 295; Fig. 13. [New Caledonia, 2352'S 16758'E, 1220-1225 m]
Parapontophilus demani (Chace, 1984)
= Pontophilus demani Chace, 1984: 48. [Selat Roti, between Timor and Pulau Roti, Lesser Sunda Is-
lands, Indonesia, 1039'S 12340'E, 520 m]
Parapontophilus difcilis Komai, 2008: 297; Fig. 6C, 14-16. [Vanuatu, 1600.73'S 16639.94'E, 1014-1050 m]
Parapontophilus geminus Komai, 2008: 302; Figs 17, 20F. [Japan, Hyuga Basin, 322.58'N 13230.54'E, 1644-
1657 m]
Parapontophilus gracilis (Smith, 1882)
= Pontophilus gracilis Smith, 1882: 36; Plate 7, Figs 2-3a. [Blake stn 315, 3218'20''N 7843'W, 225 fms;
Albatross stn 1029, of Marthas Vineyard, 458 fms]
Parapontophilus junceus (Spence Bate, 1888)
= Pontophilus junceus Spence Bate, 1888: 491; Plate 88, Figs 2-4. [Challenger stn 200, 647'N 12228'E,
between the Philippines and Borneo, 250 fms]
= Pontophilus occidentalis var. indica De Man, 1918a: 161. [Siboga Expedition stns 45, 724'S 11815'.2E,
Flores Sea, 794 m; 88, 034'.6N 1198'.5E, northern part of the Strait of Makassar, 1301 m; 178, 240'S
12837'.5E, Ceram Sea, 835 m; 211, 540'.7S 12045'.5E, E of Saleyer Island, 1158 m; 300, 1048'.6S
12323'.1E, S of Roti island, 918 m; 316, 719'.4S 11649'.5E, Bali Sea, 538 m]
Parapontophilus juxta Komai, 2008: 316; Figs 26-27. [Western Indian Ocean, of Runion, 2103.6'S
5509.8'E, 412-460 m]
Parapontophilus longirostris Komai, 2008: 305; Figs 18-19, 21A. [Austral Islands, Tubuai island, 2321.3'S
14933.9'W]
Parapontophilus modumanuensis (Rathbun, 1906)
= Pontophilus modumanuensis Rathbun, 1906: 910; Fig. 63. [vicinity of Modu Manu, Hawaii Islands,
293-800 fms]
Parapontophilus occidentalis (Faxon, 1893)
= Pontophilus occidentalis Faxon, 1893: 200. [0234.0'N 9206.0'W, 2448 m; 0456.0'N 8052.30'W, 3243
m; 0543.0'N 8550.0'W, 1760 m; 0610.0'N 8306.0'W, 2648 m; 0621.0'N 8041.0'W, 3281 m;
0705.30'N 7940.0'W, 2286 m (all Gulf of Panama); of Ecuador, 0107.0'N 8021.0'W, 2831 m; of
Costa Rica, 1014.0'N 9628.0'W, 4018 m; of Guatemala, 1446.0'N 9840.0'W, 3190 m]
Parapontophilus profundus (Spence Bate, 1888)
= Pontophilus profundus Spence Bate, 1888: 490; Plate 88, fg. 1-1z. [Challenger stn 165, 3450'S 15528'E,
of Sydney, 2600 fms]
Parapontophilus psyllus Komai, 2008: 319; Figs 28-29. [Japan, of Nago, Okinawa Island, Ryukyu Islands,
2632.90'N 12743.94'E, 396-407 m]
Parapontophilus sibogae Komai, 2008: 322; Figs 30-31. [Indonesia, Pulu Kaniungan Ketjil, 11 m]
Parapontophilus stenorhinus Komai, 2008: 324; Figs 32-33. [Tonga, 2102'S 17519'W, 555-581 m]
Parapontophilus talismani (Crosnier & Forest, 1973)
= Pontophilus talismani Crosnier & Forest, 1973: 245; Fig. 80a-d. [1638'N 1824'W, 3200 m]
Philocheras Stebbing, 1900
= Mesapus Rafnesque, 1814 (type species Mesapus fasciatus Rafnesque, 1814 (invalid senior subjec-
tive synonym of Crangon Fasciatus Risso, 1816), gender masculine; name of genus and its type spe-
cies suppressed for the purposes of the Principle of Priority, but not those of the Principle of Ho-
monymy; placed on the Ofcial Index of Rejected and Invalid Generic (resp. Specifc) Names in
= Philocheras Stebbing, 1900 (type species Crangon nanus Kryer, 1843, selected by Holthuis, 1955b,
gender masculine)
Philocheras acutirostratus (Yaldwyn, 1960)
= Pontophilus acutirostratus Yaldwyn, 1960: 41; Fig. 9. [Chatham Rise, 4342'S 17955'E, 280 fms]
Philocheras aglyptus (Crosnier, 1971)
= Pontophilus aglyptus Crosnier, 1971: 581; Figs 4a-b, 5. [Congo, baie de Point-Noire, senne de plage]
Philocheras angustirostris (De Man, 1918a)
= Pontophilus angustirostris De Man, 1918a: 163. [Siboga Expedition stns 7, 755'.5S 11426'E, reef of
Batjulmani (Java); 50, Bay of Bajo, W coast of Flores, up to 40 m; 164, 142'.5S 13047'.5E, between
Misool and New Guinea, 32 m; 273, anchorage of Pulu Jedan, E coast of Aru-islands, 13 m]
Philocheras anthonyi Taylor, 2010: 159; Figs 1-2. [Australia, Western Australia, 1735.98'S 1185908'E to
1738.56'S 11901.26'E, 222 m]
Philocheras australis (Thomson, 1879)
= Crangon australis Thomson, 1879: 231; Plate 10, Fig. A1a-c. [Cook Straits, Dunedin; Stewart Island]
Philocheras bidens (Holthuis, 1951a)
= Pontophilus bidens Holthuis, 1951a: 165; Fig. 33. [of Portuguese Guinea, 1154'N 1714'W, 55-80 m]
Philocheras bidentatus (De Haan, 1844 [in De Haan, 1833-1850])
= Crangon bidentatus De Haan, 1844 [in De Haan, 1833-1850]: Plate 45, fg. 14 (1844); 183 (1849). [Ja-
pan]
Philocheras bispinosus bispinosus (Hailstone, 1835a)
= Pontphilus bispinsus Hailstone, 1835a: 271; Fig. 30. [of Hastings]
= Pontphilus bispinsus Westwood, 1835c: 273. [of Hastings]
= Crangon nanus Kryer, 1843: 231; Plate 4, Figs 15-28. [i det sydligste Kategat (udfor Hornbet, paa
en halv Snees Favnes Dybbe og paa Leerbund)]
= Pontophilus mbizi Holthuis, 1952a: 64; Fig. 16. [1557'S 1140'E, 5 M.SW. Pointa Albina]
Philocheras bispinosus neglectus (G.O. Sars, 1883)
= Ceraphilus neglectus G.O. Sars, 1883: 45; Plate 1, fg. 7. [ad oras meridionales et occidentalis Nor-
vegi in prof 2-6 orgyarum]
Philocheras brevifagella Komai, 2001b: 419; Figs 1-3. [Ala Moana, Oahu, Hawaii, subtidal]
Philocheras brucei Komai, 2004a: 665; Figs 1-4. [Cairns, northeast Queensland, 1655'S 14546'E, 10 m]
Philocheras candidus (Kemp, 1916b)
= Pontophilus candidus Kemp, 1916b: 365; Fig. 3; Plate 8, fg. 3. [Port Blair, Andamans]
Philocheras carinicauda (Stimpson, 1860a)
= Crangon carinicauda Stimpson, 1860a: 25. [portu Sinensi Hong Kong]
Philocheras chiltoni (Kemp, 1911)
= Pontophilus chiltoni Kemp, 1911: 5; Plate 2, Figs 6-10. [New Zealand]
Philocheras coralliophilus Komai & J.N. Kim, 2010: 12; Figs 1-4. [Uehara Beach, Iriomote Island, Yaeyama
Islands, Ryukyus, subtidal] (Fig. 59)
Philocheras dimorphus (Fujino & Miyake, 1971)
= Pontophilus dimorphus Fujino & Miyake, 1971: 27; Figs 1-2. [Sagami Bay, Amadai-ba, Maruyama-
dashi, 80 m]
Philocheras echinulatus (M. Sars, 1862)
= Crangon echinulatus M. Sars, 1862: 186. [Norges Vestkyst, Manger ved Bergen]
? = Cheraphilus Patersonii Kinahan, 1860d: 130. [at the Gobbins, full description in Kinahan, 1862]
= Crangon serratus Norman, 1862: 151. [of the Shetland Isles]
Philocheras fasciatus (Risso, 1816)
= Mesapus fasciatus Rafnesque, 1814: 23. [Type locality not indicated; name suppressed for the pur-
pose of the Principle of Priority, but not those of the Principle of Homonymy; placed on the Ofcial
Index of Rejected and Invalid Specifc Names in Zoology in Opinion 522 in 1958]
= Crangon Fasciatus Risso, 1816: 82; Plate 3, fg. 5. [environs de Nice, sure les bas fond sabloneux]
Philocheras findersi (Fulton & Grant, 1902)
= Pontophilus findersi Fulton & Grant, 1902: 67; Plate 10, fg. 3. [of Shoreham, Western Port, about
4 fms]
Philocheras fujinoi J.N. Kim & Hayashi, 2000: 694; Figs 4-5. [of Shantou, northern South China Sea, 80 m]
Philocheras gaillardi (Crosnier, 1971)
= Pontophilus gaillardi Crosnier, 1971: 588; Figs 7-8. [Congo, au large de Pointe-Noire]
Philocheras gemmaceus J.N. Kim & Hayashi, 2000: 688; Figs 1-2, 3a-e. [of Kaminoseki, Seto Inland Sea,
Japan, 50 m]
Philocheras gorei (Dardeau, 1980)
= Pontophilus gorei Dardeau, 1980: 563; Figs 1-4. [2624'N 8322'W, 55 m]
Philocheras hamiltoni (Yaldwyn, 1971)
Fig. 59. Philocheras coralliophilus Komai & Kim, 2010. Photo by Arthur Anker.
= Pontophilus hamiltoni Yaldwyn, 1971: 92. [of Portobello Marine Biological Station wharf, Otago
Harbour, 2-4 fms]
Philocheras hendersoni (Kemp, 1915)
= Pontophilus hendersoni Kemp, 1915: 261; Fig. 25; Plate 13, fg. 8. [outer channel of Chilka Lake,
1-2 f]
Philocheras incisus (Kemp, 1916b)
= Pontophilus incisus Kemp, 1916b: 357; Fig. 1; Plate 8, fg. 1. [Andamans, 20 fms; Port Blair, Anda-
mans, 2-12 fms]
Philocheras intermedius (Spence Bate, 1863)
= Crangon intermedius Spence Bate, 1863: 503; Plate 41, fgs 6-6l; nec Stimpson, 1860a. [St. Vincent
Gulf, 4.5 fms]
= Crangon batei Kingsley, 1883: 129. [nomen novum for Crangon intermedius Spence Bate, 1863]
Philocheras japonicus (Dofein, 1902)
= Pontophilus japonicus Dofein, 1902: 621; Fig. B; Plate 3, fg. 6. [Sden der Sagamibai]
Philocheras kempii (De Man, 1918a)
= Pontophilus Kempii De Man, 1918a: 165. [very near Siboga Expedition stn 65 (70'S 12034'.5E), be-
tween the islands of Saleyer and Tanah Djampeah, 300-400 m]
Philocheras lapillus Wicksten, 1989b: 80; Figs 2-4. [Of Gardner Bay, Hood Island (Isla Espaola), Galpa-
gos (Archipelago de Coln), 122'S 9040'W, 46-65 m]
Philocheras lowisi (Kemp, 1916b)
= Pontophilus lowisi Kemp, 1916b: 361; Fig. 2; Plate 8, fg. 2. [Port Blair, Andamans, 3-12 fms]
Philocheras magnioculus Komai & Chan, 2007: 159; Figs 1-3. [between Libaong and Pamilacan, 931.3'N
12351.4'E, 145-163 m]
Philocheras megalocheir Stebbing, 1915: 71; Plate 15. [near East London, Cove Rock, NE 2 miles, 25 fms;
3313'S 2739'E, 37 fms]
Philocheras modestus (De Man, 1918a)
= Pontophilus modestus De Man, 1918a: 162. [Siboga Expedition stn 253, 548'.2S 13213'E, W of Kei-
islands, 304 m]
Philocheras monacanthus (Holthuis, 1961c)
= Pontophilus monacanthus Holthuis, 1961c: 26; Fig. 9. [harbour of Mersin, south-east coast of Turkey,
0.5-1.5 m]
Philocheras nikiforovi (Burukovsky, 1990)
= Pontophilus nikiforovi Burukovsky, 1990: 209; Fig. 5B. [2503'S 9727'W, 271 m]
Philocheras obliquus (Fulton & Grant, 1902)
= Pontophilus obliquus Fulton & Grant, 1902: 63; Plate 10, fg. 1. [of Shoreham, Western Port, about 5
fms]
Philocheras opici (Crosnier, 1971)
= Pontophilus opici Crosnier, 1971: 585; Figs 4c-d, 6. [Ile Annobon, 126'30''S 535'30''E, 50 m]
Philocheras parasculptus Burukovsky, 1991b: 40; Figs 2.4-2.9. [3316'S 4353'E, 415-460 m]
Philocheras parvirostris (Kemp, 1916b)
= Pontophilus parvirostris Kemp, 1916b: 372; Fig. 6; Plate 8, fg. 6. [Kilakarai, Ramnad district, S. India]
Philocheras pilosoides (Stephensen, 1927)
= Pontophilus pilosoides Stephensen, 1927: 299; Fig. 1. [Auckland Island, Carnley Harbour, 45 fms;
Campbell Island, Perseverance Harbour, 10-20 fms]
Philocheras pilosus (Kemp, 1916b)
= Pontophilus pilosus Kemp, 1916b: 367; Fig. 4; Plate 8, fg. 4. [Kilakarai, Ramnad district, S. India]
Philocheras planoculminus Bruce, 1994c: 753; Fig. 3. [Flat Top Bank, Timor Sea, 1235'S 12935'E, about
30 m]
Philocheras plebs (Kemp, 1916b)
= Pontophilus plebs Kemp, 1916b: 370; Fig. 5; Plate 8, fg. 5. [Port Blair, Andamans, 2 fms]
Philocheras prionolepis (Holthuis, 1952a)
= Pontophilus prionolepis Holthuis, 1952a: 61: Fig. 15. [606'S 1036'E, 35 M. W. Moita Seca]
Philocheras quadrispinosus (Yaldwyn, 1971)
= Pontophilus quadrispinosus Yaldwyn, 1971: 93. [of Cape Reinga, North Auckland, approx. 3421.5'S
17249'E, 58 m]
Philocheras sabsechota (Kemp, 1911)
= Pontophilus sabsechota Kemp, 1911: 6; Plate 2, Figs 11-14. [S. Sentinel Island, Andamans]
Philocheras sculptus (Bell, 1847 [in Bell, 1844-1853])
= Crangon sculptus Bell, 1847 [in Bell, 1844-1853]: 263, unnumbered text fgure. [Weymouth]
Philocheras triangulus Komai, 2006b: 31; Figs 1-4. [West Alligator Head, Kakadu National Park, Northern
Territory, 129.725'S 13213.959'E, 7.3 m]
Philocheras trispinosus (Hailstone in Hailstone & Westwood, 1835)
= Pontphilus trispinsus Hailstone in Hailstone & Westwood, 1835: 261; Fig. 25. [Hastings; see
Holthuis, 1976b for discussion of authorship]
Philocheras vanderbilti (Boone, 1935)
= Pontophilus vanderbilti Boone, 1935: 200; Plate 55. [Flores Strait, near Larantuka village, Flores Is-
land, 140 fms]
Philocheras vestigialis (Fujino & Miyake, 1971)
= Pontophilus vestigialis Fujino & Miyake, 1971: 32; Figs 3-4. [Kagoshima Bay, Kagoshima Prefecture]
Philocheras victoriensis (Fulton & Grant, 1902)
= Pontophilus victoriensis Fulton & Grant, 1902: 65; Plate 10, fg. 2. [Altona Bay and Beaumaris, Port
Phillip, about 5 fms]
Philocheras wilkinsae De Grave, 2000: 49; Figs 1-4. [north-western side of Laing Island lagoon, Hansa Bay,
Madang Province, Papua New Guinea, 410'30''S 14452'47''E, 7 m]
Philocheras wolf (Holthuis, 1951a)
= Pontophilus wolf Holthuis, 1951a: 170; Fig. 34. [of French Guinea, 927'N 1448'W, 50 m]
Philocheras yaldwyni (Zarenkov, 1968b)
= Pontophilus yaldwyni Zarenkov, 1968b: 165; Fig. 10. [southwest of New Zealand, 334 m]
Placopsicrangon Komai & Chan, 2009
= Placopsicrangon Komai & Chan, 2009 (type species Placopsicrangon formosa Komai & Chan, 2009, by
Placopsicrangon formosa Komai & Chan, 2009: 256; Figs 1-6. [2226.44'N 12221.18'E, 4824-4807 m]
Pontocaris Spence Bate, 1888
= Pontocaris Spence Bate, 1888 (type species Pontocaris propensalata Spence Bate, 1888, designated by
Holthuis, 1947b, gender feminine; name placed on the Ofcial List of Generic Names in Zoology in
= Pontocheras Bruce, 1988h (type species Pontocheras arafurae Bruce, 1988h, by original designation
Pontocaris afnis afnis (Alcock, 1901)
= Aegeon afne Alcock, 1901: 118. [of Bombay, 56-58 fms]
Pontocaris afnis allodactylus Chan, 1996: 303; Fig. 12. [Red Sea, north of Massawa Channel]
Pontocaris arafurae (Bruce, 1988h)
= Pontocheras arafurae Bruce, 1988h: 213; Figs 1-6, 7A-D. [Arafura Sea, 1104.5'S 13204.5'E, 15-22
m]
Pontocaris hilarula (De Man, 1918b)
= Aegeon propensalata var. hilarula De Man, 1918b: 301. [Bay of Bima, near south fort, 55 m]
Pontocaris laurentae Chan, 1996: 307; Fig. 14. [Indonesia, Makassar Strait, 156.6'S 11916.7'E, 134-186 m]
Pontocaris major Chan, 1996: 304, Fig. 13. [Philippines, 140.3'N 12017'E, 187-210 m]
Pontocaris pennata Spence Bate, 1888: 499; Plate 91. [Challenger stn 190, 856'S 1365'E, Arafura Sea, south
of Papua, 49 fms]
= Aegeon obsoletum Balss, 1914b: 70; Plate 1, fg. 3. [bei Enoshima, 80 m; Dzushi, 50-100 m; Sagamibai;
Sagamibai, zwischen Ito und Hatsushima, 150 m]
Pontocaris profundior Chan, 1996: 294; Fig. 8. [Red Sea, of Sudan, 2133'N 3821'E, 753-804 m]
Pontocaris propensalata Spence Bate, 1888: 496; Plate 86, fg. 5; Plate 90, fgs 2-3. [Challenger stn 192,
549'15''S 1324'15''E, of the Ki Islands, south of Papua, 140 fms]
= Pontocaris media Alcock & Anderson, 1899: 282. [Andamans, 55 fms]
Pontocaris sibogae (De Man, 1918b)
= Aegeon Sibogae De Man, 1918b: 302. [72'.6S 11523'.6E, Bali Sea, south of Kangeang, 100 m]
= Aegeon Sibogae var. intermedia De Man, 1918b: 303 [1027'.9S 12328'.7E, between the islands of
Roti and Timor, 216 m]
Pontocaris spinifera Chan, 1996: 317; Fig. 18. [Indonesia, Saleh Bay, Sumbawa, Flora Sea, 819'S 11741'E,
274 m]
Pontophilus Leach, 1817 [in Leach, 1815-1875]
= Pontophilus Leach, 1817 [in Leach, 1815-1875] (type species Crangon spinosus Leach, 1816b, by
= Cheraphilus Kinahan, 1862 (nomen novum for Pontophilus Leach, 1817 [in Leach, 1815-1875], gen-
der masculine)
Pontophilus brevirostris Smith, 1881: 435. [Fish Hawk stns 865 (4005'00''N 7023'00''W, 65 fms); 866
(4005'18''N 7022'18''W, 65 fms); 867 (4005'42''N 7022'06''W, 64 fms); 870 (4002'36''N 7022'58''W,
155 fms); 871 (4002'54''N 7023'40''W, 11 fms); 872 (4005'39''N 7023'52''W, 86 fms); 873 (4002'00''N
7057'00''W, 100 fms); 874 (4000'00''N 7057'00''W, 85 fms); 877 (3956'00''N 7054'18''W, 126 fms);
878 (3955'00''N 7054'15''W, 142.5 fms)]
Pontophilus norvegicus (M. Sars, 1861)
= Crangon norvegicus M. Sars, 1861: 248. [Floren; Manger; Bundeforden ved Christiana, 30-50 m]
? = Hippolyte costata Leuckart, 1847: 89. [Helgoland]
Pontophilus spinosus (Leach, 1816b)
= Crangon spinosus Leach, 1816b: 346. [in Danmoni australi mari; apud Plymouth Sound]
Prionocrangon Wood-Mason in Wood-Mason & Alcock, 1891c
= Prionocrangon Wood-Mason in Wood-Mason & Alcock, 1891c (type species Prionocrangon omma-
tosteres Wood-Mason in Wood-Mason & Alcock, 1891c, by monotypy, gender feminine; name placed
on the Ofcial List of Generic Names in Zoology in Opinion 470 in 1957)
Prionocrangon curvicaulis Yaldwyn, 1960: 46; Fig. 10. [Chatham Rise, 4340'S 17928'E, 220 fms]
Prionocrangon demani J.N. Kim & Chan, 2005: 1617; Fig. 9. [Sulu Sea, 1000.70'-1001.62'N 12054.72'-
12055.27'E, 1482-1488 m]
Prionocrangon dofeini Balss, 1913b: 238. [Sagamibai, 350-600 m]
Prionocrangon formosa J.N. Kim & Chan, 2005: 1615; Figs 8, 11B. [2448.86'N 1225.31'E, 365-447 m]
Prionocrangon ommatosteres Wood-Mason in Wood-Mason & Alcock, 1891c: 362. [Andaman Sea,
1125'5"N 9247'6"E, 405 fms]
Prionocrangon paucispina J.N. Kim & Chan, 2005: 1620; Figs 10, 11C. [220.54'N 11927.94'E, 2334-2543 m]
Prionocrangon pectinata Faxon, 1896: 157; Plate 2, fgs 4-7. [of Martinique, 565 fms]
Pseudopontophilus Komai, 2004b
= Pseudopontophilus Komai, 2004b (type species Pseudopontophilus serratus Komai, 2004b, by original
Pseudopontophilus serratus Komai, 2004b: 76; Figs 1-6. [New Caledonia, Sponge Bank, Norfolk Ridge,
2455.1'S 16821.6'E, 510 m]
Rhynocrangon Zarenkov, 1965
= Rhynocrangon Zarenkov, 1964 (unavailable name under Art. 13.3)
= Rhynocrangon Zarenkov, 1965 (type species Crangon (Sclerocrangon) sharpi Ortmann, 1896b, by
Rhynocrangon alata (Rathbun, 1902a)
= Sclerocrangon alata Rathbun, 1902a: 891. [Admiralty Inlet, Puget Sound, 40 fms]
Rhynocrangon rugosa Komai & Komatsu, 2008: 184; Figs 1-5. [of Kuji, Iwate prefecture, northeastern Ja-
pan, 4014.98'N 14206.64'E to 4013.31'N 1427.38'E, 156 m]
Rhynocrangon sharpi (Ortmann, 1896b)
= Crangon (Sclerocrangon) sharpi Ortmann, 1896b: 178. [Alaska, Kodiak Archipelago, Marmot Isle, 45
fms]
Sabinea Ross, 1835
= Sabinea Ross, 1835 (type species Crangon septemcarinata Sabine, 1824, by monotypy; name placed
on the Ofcial List of Generic Names in Zoology in Opinion 470 in 1957)
= Myto Kryer, 1845 (type species Myto Gaimardii Kryer, 1845 (junior subjective synonym of Cran-
gon septemcarinata Sabine, 1824), by monotypy, gender masculine)
Sabinea hystrix (A. Milne-Edwards, 1881b)
= Paracrangon hystrix A. Milne-Edwards, 1881b: 6. [prs de la Guadeloupe, 734 brasses]
= Sabinea princeps Smith, 1882: 38; Plate 8, Figs 1-1b. [Blake stns 312, 3950'45''N 7011'0''W, 466 fms;
326, 3342'15''N 760'50''W, 464 fms; 337, 3820'8''N 7323'20''W, 740 fms; Albatross stns 892, of Block
Island, 3946'N 715'W, 487 fms; 893, 3952'20''N 7058'W, 372 fms]
Sabinea sarsii Smith, 1879
= Sabinea Sarsii Smith, 1879: 59; Plate 11, Figs 6-8. [St. Georges Banks, 60 fms; Gulf of Maine, about
east-southeast from Cape Sable, Nova Scotia, 4240'N 6658'W, 112 fms; Le Have Bank, 60 fms; Lo-
foten Islands, coast of Norway]
Sabinea septemcarinata (Sabine, 1824)
= Crangon septemcarinata Sabine, 1824: ccxxxvi; Plate 2, fgs 11-13. [west coast of Davis Strait]
? = Myto gaimardi Kryer, 1845: 470. [Spitsbergen]
Sclerocrangon G.O. Sars, 1883
= Sclerocrangon G.O. Sars, 1883 (type species Cancer Boreas Phipps, 1774, by monotypy, gender femi-
Sclerocrangon atrox Faxon, 1893: 199. [Albatross stns 3418 (1633'00''N 9952'30''W, 660 fms); 3424
(2115'00''N 10623'00''W, 676 fms)]
Sclerocrangon boreas (Phipps, 1774)
= Cancer Boreas Phipps, 1774: 190; Plate 12, fg. 1. [stomach of a seal, caught near the coast of Spitz-
bergen]
= Cancer homaroides O. Fabricius, 1780: 241. [Umitak]
= Cancer arctica Dewhurst, 1834: 254. [Northern and Greenland Ocean]
Sclerocrangon derjugini Kobjakova, 1936a: 224. [Okhotsk Sea, 182-664 m; fully described in Kobjakova,
1937]
Sclerocrangon ferox (G.O. Sars, 1877)
= Cheraphilus ferox G.O. Sars, 1877: 239. [6210.2N 459.6E, 417 fms]
Sclerocrangon igarashii Komai & Amaoka, 1991: 27; Figs 1-4. [of Urup Island, Kurile Islands, 4517'2N
14928'2E, 450 m]
Sclerocrangon salebrosa (Owen, 1839)
= Crangon salebrosus Owen, 1839: 88; Plate 27, fg. 1. [shores of Kamtschatka]
= Sclerocrangon gasuyebi Yokoya, 1933: 41; Fig. 22. [Sy-Maru stns 602, north of Is. Sado, 205 m; 628,
south-west of Oga, 148 m; 652, Tugaru Strait, 110 m]
Sclerocrangon unidentata Komai & Takeda, 1989: 77, Figs 1-5. [of Miyako, Iwate Pref., northeastern Hon-
shu, 600-700 m]
Sclerocrangon zenkevitchi Birstein & Vinogradov, 1953: 218; Figs 2-5. [Bering Sea, 2995-3940 m]
Syncrangon J.N. Kim & Hayashi, 2003
= Syncrangon J.N. Kim & Hayashi, 2003 (type species Crangon angusticauda De Haan, 1844 [in De
Haan, 1833-1850], by original designation, gender feminine)
Syncrangon angusticauda (De Haan, 1844 [in De Haan, 1833-1850])
= Crangon angusticauda De Haan, 1844 [in De Haan, 1833-1850]: Plate 45, fg. 15 (1844); 183 (1849).
[Japan]
Syncrangon dentata (Balss, 1914b)
= Crangon (Sclerocrangon) angusticauda var. dentata Balss, 1914b: 65; Fig. 40 (as Sclerocrangon angusti-
cauda dentala). [Negishi Harbour bei Yokohama, 45 m, and Dzushi, 130 m]
Vercoia Baker, 1904
= Vercoia Baker, 1904 (type species Vercoia gibbosa Baker, 1904, by monotypy, gender feminine)
Vercoia gibbosa Baker, 1904: 158; Plate 31. [South Australia, 36-54 m; see uri, 1992]
Vercoia interrupta J.N. Kim & Fujita, 2004: 2014; Figs 1-7. [Cape Maeda, Okinawa island, Ryukyu Islands,
Japan, 8.5 m]
Vercoia japonica Komai, 1995b: 123; Figs 1-4. [Omurodashi, of Izu-Ohshima Island, 3431.7'N 13923.2'E,
138-167 m]
Vercoia socotrana uri, 1992: 1448; Figs 7-11. [Socotra Island, Gulf of Aden, western Indian Ocean,
1221'08''N 5428'00''E, 80 m]
Family GLYPHOCRANGONIDAE Smith, 1884
Glyphocrangon A. Milne-Edwards, 1881b
= Thalascaris Spence Bate, 1878 (type species Glyphocrangon rimapes Spence Bate, 1888, designated by
Holthuis, 1971, gender feminine; name suppressed for the purposes of the Principle of Priority but
not for those of the Principle of Homonymy, placed on the Ofcial Index of Rejected and Invalid
= Glyphocrangon A. Milne-Edwards, 1881b (type species Glyphocrangon spinicauda A. Milne-Ed-
wards, 1881b, by original designation, gender feminine; name placed on the Ofcial List of Generic
= Rhachocaris Smith, 1882 (type species Rhachocaris Agassizii Smith, 1882 (junior subjective synonym
of Glyphocrangon aculeatum A. Milne-Edwards, 1881b), designated by Fowler, 1912, gender feminine)
= Plastocrangon Alcock, 1901 (type species Glyphocrangon ccescens Wood-Mason in Wood-Mason &
Alcock, 1891c, designated by Fowler, 1912, gender feminine)
Glyphocrangon aculeata A. Milne-Edwards, 1881b
= Glyphocrangon aculeatum A. Milne-Edwards, 1881b: 5. [ Saint-Vincent, 593 brasses]
= Rhachocaris Agassizii Smith, 1882: 43; Plate 5, fg. 2; Plate 6, fg. 2. [Blake stn 326, 3342'15''N
760'50''W, 464 fms]
Glyphocrangon acuminata Spence Bate, 1888: 522; Plate 94, Figs 2-3. [Challenger stn 175, Fii Islands, 1902'S
17710'E, 2468 m; lectotype designation by Komai, 2004c]
Glyphocrangon africana Komai, 2010b: 84; Figs 1-4. [of Cape Point, South Africa, 2269 m]
Glyphocrangon alata Faxon, 1893: 201. [Albatross stns 3395 (North Pacifc Ocean, 0730'36''N 07839'00''W),
730 fms; 3418 (North Pacifc Ocean, 1633'00''N 09852'30''W), 660 fms; ?]
Glyphocrangon albatrossae Komai, 2004c: 468; Figs 40-41. [Philippines, South China Sea, of W Luzon,
1558.15'N 11940.20'E, 1719 m]
Glyphocrangon alispina Chace, 1939: 39. [of Bahia Cardenas, Matanzas Province, Cuba, 2324'N
8100'30''W, 370-605 fms]
Glyphocrangon amblytes Komai, 2004c: 561; Figs 89-90. [Madagascar, 2225.3'N 4304.9'E, 525-550 m]
Glyphocrangon andamanensis Wood-Mason in Wood-Mason & Alcock, 1891c
= Glyphocrangon investigatoris var. andamanensis Wood-Mason in Wood-Mason & Alcock, 1891c: 356;
accompanying illustration in Wood-Mason, 1894: Plate 6, fgs 2-2ba. [Andaman Sea, 1131.40'N
9246'40''E, 188-220 fms]
Glyphocrangon arduus Komai, 2007: 566; Figs 1-3. [French Polynesia, Austral Islands, of Tubuai Island,
2321.4'S 14934.2'W, 600-1200 m]
Glyphocrangon armata Komai, 2004c: 570; Figs 95-96. [New Caledonia, 2310.41'S 16649.16'E, 870-1000 m]
Glyphocrangon assimilis De Man, 1918b: 294. [719'.4S 11649'.5E, 538 m; lectotype designation by Komai,
2004c]
Glyphocrangon atlantica Chace, 1939
= Glyphocrangon (Plastocrangon) caecescens atlantica Chace, 1939: 40. [south of Santa Clara Province,
Cuba, 2047'N 8024'W, 2125 fms]
Glyphocrangon aurantiaca Holthuis, 1971: 303; Fig. 8. [Of Tobago, 1137.3'N 6059.4'W to 1141'N
6101.3'W, 400-700 fms]
Glyphocrangon boletifera Komai, 2004c: 558; Figs 87-88. [Gulf of Aden, 1306.12'N 4624.30'E, 1061-1080
m]
Glyphocrangon brevis Komai, 2006c: 250; Figs 1A, 2A, 3. [Madagascar, 1740'S 4312'E, 1075-1175 m]
Glyphocrangon caeca Wood-Mason in Wood-Mason & Alcock, 1891c
= Glyphocrangon cca Wood-Mason in Wood-Mason & Alcock, 1891c; 358 accompanying illustration
in Wood-Mason, 1894: Plate 7, fgs 1-1a. [Bay of Bengal, 1347'30''N 9236'E, 561 fms]
Glyphocrangon caecescens Wood-Mason in Wood-Mason & Alcock, 1891c
= Glyphocrangon ccescens Wood-Mason in Wood-Mason & Alcock, 1891c: 357; accompanying il-
lustration in Wood-Mason, 1894: Plate 7, fgs 5-5a. [Bay of Bengal, 1158'N 8852'17''E, 1748 fms]
Glyphocrangon cerea Alcock & Anderson, 1894: 151. [Investigator stn 150, Laccadive Sea, of the Northern
Maldive Atoll, 705'45''N 7504'E, 719 fms]
Glyphocrangon chacei Komai, 2004c: 534; Figs 73-74. [Philippines, between Bohol and Siquior Islands,
Mindanao Sea, 0922.30'N 12347.40'E, 719 m]
Glyphocrangon confusa Komai, 2004c: 597; Figs 110, 112A-D. [Indonesia, Halmahera, 0015.00'N
12724.35'E, 545 m]
Glyphocrangon conodactylus Komai, 2004c: 555; Figs 85-86. [New Caledonia, Mount K, Loyalty Islands,
2444.55'S 17007.00'E, 790-825 m]
Glyphocrangon cornuta Komai, 2004c: 472; Figs 42-43. [New Caledonia, 2356'S 16641'E, 2660-2750 m]
Glyphocrangon crosnieri Komai, 2004c: 431; Figs 22-23, 115. [Madagascar, 2226'S 4303'E, 540 m]
Glyphocrangon demani Komai, 2006c: 253; Figs 1B, 2B, 4. [Indonesia, between Ceram and New Guinea,
0337.7'S 13126.4'E, 924 m]
Glyphocrangon dentata Barnard, 1926
= Glyphocrangon gilesii var. dentata Barnard, 1926: 128. [2559'S 3331'E, 540 m]
Glyphocrangon dimorpha Komai, 2004c: 459; Figs 37-38. [Norfolk Ridge, 2356'S 16828'E, 1846-1862 m]
Glyphocrangon elephas Komai, 2005: 253; Figs 1-3. [Southern Australia, Great Australian Bight, 3317'S
12832'E, 360 m]
Glyphocrangon faxoni De Man, 1918b
= Glyphocrangon (Plastocrangon) Faxoni De Man, 1918b: 298. [715'S 11515'.6E, Bali Sea, 289 m;
1039'S 12340'E, between the islands of Roti and Timor, 520 m]
Glyphocrangon ferox Komai, 2004c: 506; Figs 59-60. [Madagascar, 1316'S 4733'E, 1490-1600 m]
Glyphocrangon fmbriata Komai & Takeuchi, 1994: 458; Figs 1-3. [Sio Guyot, Mid-Pacifc Mountains,
1816.05'N-1815.87'N 17120.99'E-17122.01'E, 1300-1312 m]
= Glyphocrangon joani Allen & Butler, 1994: 435; Figs 10-14. [Mid-Pacifc Ocean, 17115.0'E 1814.0'N,
1281-1651 m]
Glyphocrangon formosana Komai, 2004c: 594; Figs 108, 109E-H. [Taiwan, of Su-Aou, I-Lan County, NE of
Taiwan, 300-400 m]
Glyphocrangon gilesii Wood-Mason & Alcock, 1891b
= Glyphocrangon Gilesii Wood-Mason & Alcock, 1891b: 193; accompanying illustration in Wood-
Mason, 1894: Plate 7, Figs 4-4a. [eight miles south-east of Cinque Island, in the Andaman Sea, 500
fms]
Glyphocrangon grandis Komai & Chan, 2008: 41; Figs 1-5, 12D. [321.32'N 12412.26'E, 1037-1100 m]
Glyphocrangon granulosis Spence Bate, 1888: 507; Plate 92; Plate 93, fg. 1. [Challenger stn 218, 0233'S
14404'E, between New Guinea and Admiralty Islands, 1070 fms]
Glyphocrangon haematonotus Holthuis, 1971: 315; Figs 6c, 7c. [Straits of Florida, 2634'N 7943'W, 270 fms]
Glyphocrangon hakuhoae Takeda & Hanamura, 1994: 24; Figs 11-13. [Flores Sea, 0547.3'N 11935.4'E -
0546.9'N 11934.6'E, 280 m]
Glyphocrangon hastacauda Spence Bate, 1888: 519; Plate 93, fg. 5. [Challenger stn 232, 3511'N 13928'E, of
Japan, 345 fms]
Glyphocrangon holthuisi Kensley, Tranter & Grifn, 1987: 321; Figs 22, 25C. [New South Wales, south-east
of Cape Byron, 2841'S 15352'E, 153 m]
Glyphocrangon humilis Komai, 2006c: 254; Figs 1C, 2C, 5. [Japan, Tosa Bay, 3300'N 13335'E, 660-700 m]
Glyphocrangon indonesiensis Komai, 2004c: 408-412; Figs 10-11. [Indonesia, Tanimbar Islands, 0816'S
13159'E, 769-809 m]
Glyphocrangon investigatoris Wood-Mason & Alcock, 1891b: 191; accompanying illustration in Wood-
Mason, 1894: Plate 6, Figs 3-3b. [1935'N 9224'E, 272 fms]
Glyphocrangon isos Komai & Chan, 2008: 54; Figs 9-11, 15. [1455'N 1238'E, 922-909 m]
Glyphocrangon kapala Komai, 2004c: 428; Figs 20-21. [Eastern Australia, East of Broken Bay, New South
Wales, 3335'S 15200'E, 810 m]
Glyphocrangon lineata Komai, 2004c: 545; Figs 79-80. [Indonesia, Kai Islands, 0515'S 13259'E, 769-809 m]
Glyphocrangon longipes Komai, 2004c: 567; Figs 93-94. [Philippines, North of Palawan Passage, 1057.45'N
13838.15'E, 686 m]
Glyphocrangon longirostris (Smith, 1882)
= Rhachocaris longirostris Smith, 1882: 51; Plate 5, fg. 1; Plate 6, fg. 1. [Blake stn 330, 3141'N 7435'W,
1047 fms]
Glyphocrangon longleyi Schmit, 1931b: 393. [south of Tortugas, 18-220 fms]
Glyphocrangon loricata Faxon, 1895: 140; Plate 38bis. [Albatross stns 3402 (South Pacifc Ocean, 0057'30''S
08903'30''W), 421 fms; and 3410 (North Pacifc Ocean, 0019'00''N 09034'00''W), 331 fms]
Glyphocrangon lowryi Kensley, Tranter & Grifn, 1987: 322; Figs 23, 25D. [Queensland, north-east of Dan-
ger Point, 2803'S 15404'E, 720 m]
Glyphocrangon mabahissae Calman, 1939: 217, fg. 8. [John Murray Expedition stn 115 (0505.18'S
3922.12'E), Zanzibar area, 640-658 m]
Glyphocrangon major Komai, 2004c: 514; Figs 63-64. [Indonesia, N Makassar Strait, 034.6'N 11908.5'E,
1301 m]
Glyphocrangon megalophthalma De Man, 1918b: 296. [422'.1S 11816'.9E, Strait of Makassar, 2029 m; lec-
totype designation by Komai, 2004c]
Glyphocrangon musorstomia Komai, 2006c: 256; Figs 1D-E, 6-7. [Wallis and Futuna Islands, Tuscarora
Bank, 1147.4'S 17825.3'E, 900 m]
Glyphocrangon neglecta Faxon, 1896: 159; Plate 1, Figs 5-6. [of Grenada, 340 fms; of Montserrat, 303 fms;
of Grenada, 291 fms]
Glyphocrangon nobilis A. Milne-Edwards, 1881b
= Glyphocrangon nobile A. Milne-Edwards, 1881b: 5. [prs de la Dominique, 1131 brasses; neotype
designation by Holthuis, 1971; see discussion in Forest & Holthuis, 1997]
Glyphocrangon novaecastellum Kensley, Tranter & Grifn, 1987: 324; Figs 24, 25E. [New South Wales,
south-east of Newcastle, 3308'S 15227'E, 720 m]
Glyphocrangon panglao Komai & Chan, 2008: 47; Figs 6-8, 13B1-B2. [851'N 12310'E, 982-1040 m]
Glyphocrangon parva Komai, 2004c: 412; Figs 12-13. [Philippines, Sulu Sea, East of Palawan, 0913'N
11851.15'E, 2021 m]
Glyphocrangon parviocullus Komai, 2006c: 259; Figs 1F, 2E, 8. [New Caledonia, 2310'S 16649'E, 870-
1000 m]
Glyphocrangon perplexa Komai, 2004c: 579; Figs 101-102. [East China Sea, 2921.5'N 12728.7'E, 543-600 m]
Glyphocrangon podager Spence Bate, 1888: 516; Plate 93, fg. 2. [Challenger stn 146, 4646'S 4531'E, near
Glyphocrangon priononota Wood-Mason & Alcock, 1891b: 192; accompanying illustration in Wood-Ma-
son, 1894: Plate 6, fgs 1-1a. [Investigator stn 104 (Laccadive Sea, 1112'47''N 7425'30''E), 1000 fms]
Glyphocrangon proxima Komai, 2004c: 416; Figs 14-15. [Indonesia, Kai Islands, 0514'S 13300'E, 688-694
m]
Glyphocrangon pugnax De Man, 1918b: 293. [1039'S 12340'E, between the islands of Roti and Timor, 520
m; lectotype designation by Komai, 2004c]
Glyphocrangon punctata Komai, 2004c: 475; Figs 44-45. [Philippines, South of Mindoro, 0606.78'N
12558.91'E, 2820-2770 m]
Glyphocrangon regalis Spence Bate, 1888: 517; Plate 93, fgs 3-4. [Challenger stn 194, 434'S 12957'30"E, of
Banda Island, 200 fms; see Komai, 2004c]
= Glyphocrangon juxtaculeata Chace, 1984: 15; fgs 3-4. [Indonesia, of southern Buru, 347.15'S
12623.40'E, 946 m]
Glyphocrangon richeri Komai, 2004c: 495; Figs 54-55. [Wallis and Futuna Islands, SW slope of Combe
Bank, 1234'S 17811'W, 1280-1300 m]
Glyphocrangon rimapes Spence Bate, 1888: 523; Plate 94, fg. 4. [Challenger stns 300, 3342'S 7818'W, near
Juan Fernandez, 1375 fms; 237, 3437'N 14032'E, near Yokohama, 1875 fms; 331, 3747'S 3020'W,
South Atlantic, between Buenos Ayres and Tristan de Cunha, 1715 fms]
Glyphocrangon robusta Komai, 2004c: 526; Figs 69-70. [Philippines, South of Mindoro, 1159'N 12113'E,
320-337 m]
Glyphocrangon rubricinctuta Komai, 2004c: 599; Figs 111, 112E-H. [Wallis and Futuna Islands, Bayonnaise
Bank, 1154.2'S 17931.4'W, 597-600 m]
Glyphocrangon rudis Komai, 2006c: 261; Figs 1G, 2F, 9. [Solomon Islands, 0831.2'S 16037.7'E, 1036-
1138 m]
Glyphocrangon runcinata Komai, 2004c: 420; Figs 16-17. [East China Sea, East of Kuchiminose Bank,
2921.5'N 12728.7'E, 543-600 m]
Glyphocrangon saintlaurentae Komai, 2004c: 487; Figs 50-51. [Runion Island, 2128.1'S 5632.4'E, 4030 m]
Glyphocrangon sculpta (Smith, 1882)
= Rhacocharis sculpta Smith, 1882: 49; Plate 5, fg. 3; Plate 6, fgs 3-3d. [Blake stn 339, 3816'45''N
7310'30''W, 1186 fms]
Glyphocrangon sibogae De Man, 1918b
= Glyphocrangon Sibogae De Man, 1918b: 295. [Flores Sea, 0724'S 11815.2'E, 794 m; lectotype desig-
nation by Komai, 2004c]
Glyphocrangon sicaria Faxon, 1893
= Glyphocrangon sicarius Faxon, 1893: 202. [Albatross stn 3382 (North Pacifc Ocean, 0621'00''N
08041'00''W), 1793 fms]
Glyphocrangon similior Komai, 2004c: 590; Figs 107, 109A-D. [Vanuatu, of Epi, 1633'S 16755'E, 602-
620 m]
Glyphocrangon smithii Wood-Mason in Wood-Mason & Alcock, 1891c
= Glyphocrangon Smithii Wood-Mason in Wood-Mason & Alcock, 1891c: 357; accompanying illustra-
tion in Wood-Mason, 1894: Plate 7, fgs 3-3a. [Bay of Bengal, 1347'30''N 9236'E, 561 fms]
Glyphocrangon speciosa Komai, 2004c: 500; Figs 56-58. [New Caledonia, 2504'S 16846'E, 1093-1193 m]
Glyphocrangon spinicauda A. Milne-Edwards, 1881b: 3. [prs de St-Kits, 250 brasses]
Glyphocrangon spinossisima Brand & Takeda, 1996: 264; Figs 1-4. [Sulu Sea, 0831.6'S 11835.7'E, 1712-
1840 m]
Glyphocrangon spinulosa Faxon, 1893: 202. [Albatross stns 3353 (North Pacifc Ocean, 0706'15''N
08034'00''W), 695 fms; 3418 (North Pacifc Ocean, 1633'00''N 09952'30''W), 660 fms; 3419 (North
Pacifc Ocean, 1634'30''N 10003'00''W), 672 fms; 3424 (North Pacifc Ocean, 2115'00''N
10623'00''W), 676 fms; 3425 (North Pacifc Ocean, 2119'00''N 08034'00''W), 695 fms; 3435 (North
Pacifc Ocean, 2648'00''N 11045'20''W), 859 fms]
Glyphocrangon stenolepis Chace, 1984: 22; Figs 5-6. [South China Sea, southeast of Hong Kong, 2031'N
11549'E, 485 m]
Glyphocrangon taludensis Hendrickx, 2010: 359; Figs 1-3. [1826'45''N 10416'10''W, 1858-1879 m]
Glyphocrangon tasmanica Komai, 2004c: 574; Figs 97-98. [Lord Howe Ridge, 400 m]
Glyphocrangon unguiculata Wood-Mason & Alcock, 1891b: 193; accompanying illustration in Wood-Ma-
son, 1894: Plate 7, fgs 2-2a. [Investigator stn 105 (Arabian Sea, 152'N 7234'E), 740 fms]
Glyphocrangon vicaria Faxon, 1896: 159. [054'N 919'W, 1189 fms]
Glyphocrangon wagini Burukovsky, 1990: 206; Fig. 4B (as G. wagigi), 5A. [2456'S 8831'W, 580-564 m]
Unavailable names
Macrobrachium divakarani Jayachandran, 2001: 91; Fig. 21. [unavailable under Art. 16.4.1]
Macrobrachium josephi Jayachandran, 2001: 123; Fig. 32. [Veli Lake, Kulathoor rivulet (Kerala, India);
unavailable under Art. 16.4.1]
Allobrachium Jayachandran, 2001 [unavailable under Art. 13.3]
Penaeoidea Larvata
Cerataspis afnis Kishinouye, 1926: 66; Fig. 2. [Between Kyushyu and the Ryukyu Archipelago, Japan]
Cerataspis longiremis Dohrn, 1871: 360; Plates 28-29. [dessen Herkunf mir aber entfallen is]
Cerataspis monstruosus Gray, 1828: 8; Plate 6, fg. 5. [Found in stomach of a delphin of the coast of Brazil]
Cerataspis Petitii Gurin-Mneville, 1844 [in Gurin-Mneville, 1829-1844]: 18; Plate 22. [trouv en pleine
mer dans lOcan Indien]
Cryptopus Defrancii H. Milne Edwards, 1837 [in H. Milne Edwards, 1836-1844]: 149; Plate 54bis, fg. 4. [la
Mditerrane]
Euphema armata H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 421. [dans locan Atlantique
austral]
Euphema polyacantha Ortmann, 1893: 77. [Plankton-Expedition der Humbolt-Stifung, JN 252 (9.4S
41.9W, 0-400 m), Guineastrom]
Hoplites longirostris Philippi, 1857: 329; Plate 14, fgs 1a-i. [Atlantischen Ocean in 25N.B. und 2250'W.L.]
Loxopis tridens Dana, 1852b: 661 (1852); Plate 44, fg. 4 (1855). [Sooloo Archipelago]
Ophthalmeryon transitionalis Spence Bate, 1889: 6; Plate 9. [swallowed by a dolphin, and therefore been
afected somewhat by the gastric juices of the fshs stomach]
Opisthocaris mlleri Ortmann, 1893: 77; Plate 4, fg. 5. [Plankton-Expedition der Humbolt-Stifung, Flori-
dastrom, JN 45 (41.6N 56.3W, 0-200 m) and 47 (40.4N 57.0W, 0-300 m); Sdl. Aequatorialstrom
JN 218 (3.8S 32.6W, 0-400 m), 228 (1.8S 38.1W, 0-600 m), 231 (1.5S 39.2W, 0-400 m), 232 (0.4S
42.4W, 0-400 m), 235 (0.1S 44.2W, 0-400 m), 246 (0.4S 46.6W, 0-400 m)]
Platysacus crenatus Spence Bate, 1888: 363; Plate 63. [Challenger stn 352, on the surface of the Atlantic
Ocean, Sierra Leone, 1055'N 1746'W]
Rachitia spinalis Dana, 1852b: 667 (1852); Plate 44, fg. 8 (1855). [Atlantic, of the harbour of Rio de
Janeiro]
Solenocera sp. larva barbata Heegaard, 1966: 89; Figs 181-192. [Great Barrier Reef stn 46, 1432'S 14532'E,
inside Cooks Passage, 33 m; Great Barrier Reef Expedition, no further data; Discovery stn 276,
554'00"S 1119'00"E, 110-0 m]
Solenocera sp. larva danae Heegaard, 1966: 53; Figs 66-108. [Dana stns 3922-II, 345'S 5633'E, 600 m wire
out; 3903-II, III, IV, 530'N 9338'E, 600, 300, 100 m wire out resp.; 3860-XX, XXI, 257'S 9936'E, 600,
300 m wire out resp.]
Solenocera sp. larva elongata Heegaard, 1966: 85; Figs 65, 170-180. [Terra Nova stn 43, 2206'S 3940'W,
surface]
Solenocera sp. larva nodulosa Heegaard, 1966: 81; Figs 154-169. [Dana stn 3921-III, 336'S 5819'E 300 m
wire out]
Solenocera sp. larva sumatransis Heegaard, 1966: 66; Figs 109-153. [Dana stn 3903-III, 530'N 9328'E, 300
m wire out]
Penaeoidea Nomina Dubia
Hemipenaeus dubius Spence Bate, 1881: 187. [Among Philippine Islands, less than 20 fms]
Pandalus tenuicornis Rankin, 1900: 544. [Type locality not indicated]
Penaeus amazonicus Kingsley, 1883: 106. [Upper Amazon]
Penaeus gracilis Dana, 1852a: 27. [in mari Suluensi; Burkenroad, 1934a considers this an immature Metap-
enaeopsis species]
Penaeus pubescens Stimpson, 1871: 133. [St. Thomas]
Penaeus telsodecacanthus Spence Bate, 1881: 182. [in the channels of the Japanese Islands, 8-10 fms]
Penaeus tenuis Dana, 1852a: 27. [in mari Atlantico prope portum Rio Negro Patagoniae]
Penaeus villosus Gurin-Mneville, 1838 [in Gurin-Mneville, 1829-1838]: 36. [Nouvelle-Irlande; quite
probably Metapenaeus monoceros or M. mastersii, according to Burkenroad, 1934a]
Penacus (Protosolenocera) kessleri Czerniavsky, 1878: 23. [Hankou, Yan-Tse-Kian, north of Nankin, China]
Sergestes caudatus Kryer, 1859: 270; Plate 5, fg. 14. [nordlige Kategat]
Penaeoidea Nomina Nuda
Metapenaeus tweedii Morris & Bennet, 1952: 165.
Sergestoidea Larvata
Acanthosoma brevitelsonis Spence Bate, 1888: 367; Fig. 51; Plate 64, fg. 1. [Western Pacifc Ocean]
Acanthosoma dohrni Gurney, 1924: 92; Fig. 31. [Terra Nova Expedition stn 129, of Three Kings Island, New
Zealand; 311, Atlantic, 3529'S 5026'W]
Acanthosoma dorsispinalis Spence Bate, 1888: 370: Plate 65, fg. 1. [Western Pacifc Ocean]
Acanthosoma hispida Gurney, 1924: 90; Figs 29-30. [Terra Nova Expedition stn 129, of Three Kings Island,
New Zealand]
Acanthosoma laevirostratis Spence Bate, 1888: 374; Plate 65, fg. 2. [North of Admiralty Islands, between
Challenger stations 221 and 222]
Acanthosoma longitelsonis Spence Bate, 1888: 371; Plate 44, fg. 3. [South Pacifc Ocean]
Acanthosoma macrotelsonis Spence Bate, 1888: 373; Plate 66, fg. 1. [Samboangan, Philippines]
Acanthosoma tynitelsonis Spence Bate, 1888: 369; Plate 64, fg. 2. [Western Pacifc Ocean]
Elaphocaris crassus Spence Bate, 1888: 362; Plate 61, fg. 4. [South Atlantic, between Montevideo and
Tristan da Cunha, 32S 45W]
Elaphocaris dohrni Spence Bate, 1888: Plate 62. [South Atlantic, 32S 45W, between Monte Video and
Tristan da Cunha]
Elaphocaris hispida Gurney, 1924: 85; Figs 24-28. [Terra Nova Expedition: stns 65, Atlantic, 2328'N 3445'W;
66, Atlantic, 2535'N 3410'W; 109, of New Zealand, 3415'S 172E; 120, of New Zealand, 3426'S
17214'E]
Elaphocaris ortmanni Gurney, 1924: 84; Figs 21-23. [Terra Nova Expedition stn 17, Atlantic, 2617'N
1814'W]
Elaphocaris suhmi Spence Bate, 1888: 353; Figs 49-50; Plate 61, fgs 1-3. [South Atlantic, between Montevi-
deo and Tristan da Cunha, 32S 45W]
Erichthina demissa Dana, 1852b: 634; Plate 42, fg. 3. [Near eastern entrance of Straits of Sunda]
Mastigopus acetiformis Spence Bate, 1888: 376. [Western Pacifc Ocean]
Mastigopus crassus Spence Bate, 1888: 381; Figs 52-53. [Taken on the voyage from Hilo to Tahiti, 50'N]
Mastigopus dorsispinalis Spence Bate, 1888: 375; Plate 65, fg. 3. [Pacifc Ocean, NW of the Admiralty Is-
lands, 1N 146E]
Mastigopus spiniventralis Spence Bate, 1888: 379; Plate 67, fg. 4. [Western Pacifc Ocean]
Mastigopus suhmi Spence Bate, 1888: 378; Plate 66, fg. 2. [North Atlantic Ocean]
Mastigopus tenuis Spence Bate, 1888: 428; Plate 65, fg. 4. [Western Pacifc Ocean]
Sceletina armata Dana, 1852b: 663. [Atlantic, 030'S 1730'W]
Sceletina laticeps Dana, 1852b: 664. [Pacifc, 30 miles W of Assumption Island, one of the Ladrones]
Sceletina orientalis Dana, 1852b: 665. [Sooloo Sea, SW of Panay]
Sciacaris telsonis Spence Bate, 1888: 438; Plate 78, fg. 1. [N of New Guinea]
Sergestoidea Nomina Dubia
Acheles arachnipodus Cocco, 1832: 204; unnumbered Plate, fg. 1. [Messina, Sicily]
Lucifer pacifcus Dana, 1852b: 673 (1852); Plate 45, fg. 2. (1855). [Pacifc, 1520'S 148W]
Mastigopus spinosus Leuckart, 1853: 258. [Type locality not indicated]
Phasmatocarcinus discophthalmus Tilesius, 1819: 7; Plate 21a, fg. 10. [placed on the Ofcial Index of Re-
jected and Invalid Works in Zoological Nomenclature in Opinion 864 in 1969]
Phasmatocarcinus glaucus Tilesius, 1819: 6; Plate 21a, fg. 9. [placed on the Ofcial Index of Rejected and
Invalid Works in Zoological Nomenclature in Opinion 864 in 1969]
Prionorhynchus Apus Tilesius, 1819: 6; Plate 21a, fg. 7. [placed on the Ofcial Index of Rejected and Inva-
lid Works in Zoological Nomenclature in Opinion 864 in 1969]
Sergestes dorsispinalis Spence Bate, 1888: 394; Plate 72, fg. 1. [S of Australia; possibly a juvenile stage of
Sergestes arcticus T. Kikuchi (in lit.)]
Sergestes hanseni Dennell, 1955: 400. [Bermuda area]
Sergestes laterodentatus Spence Bate, 1888: 395. [S of Australia; possibly a mastigopus stage of Sergestes
arcticus (T. Kikuchi (in lit.))]
Sergestes oculatus Perrier, 1886: 81. [Le Challenger et le Talisman, la faune des Sargasses]
Sergestes praecollus Spence Bate, 1888: 423; Plate 77, fg. 2. [North Pacifc Ocean]
Sergestes sundi Dennell, 1955: 400. [Bermuda area]
Sergestes tenuis Spence Bate, 1888: 233, Figs 43-45. [North of New Guinea, and on the voyage along the
Marianne Islands]
Sergestes utrinquedens Spence Bate, 1888: 433. [North Pacifc Ocean]
Sergestes Colosii Cecchini, 1933: 4; Plate 1, fg. 1. [Ammiraglio Magnaghi stns 138, a sud ouest di Aden, 125'
- 127'N 4447' - 4445'E, 388 m; 150, a nord est di Porto Sudan, 2047'N 3822'30"E, 1200 m]
Symphysopus hirtus Tilesius, 1819: 9; Plate 21b, fg. 19. [placed on the Ofcial Index of Rejected and Inva-
lid Works in Zoological Nomenclature in Opinion 864 in 1969]
Sergestoidea Nomina Nuda
Sergestes forins Bakus, 1989: 36.
Caridea Larvata
Amphiplectus depressus Spence Bate, 1888: 623; Plate 110, fg. 3. [Of Recife, Brazil, 905'S, 3450'W, 6640
m]
Anebocaris ancylifer Coutire, 1907c: 407; unnumbered fgure. [Gorgie du Sud]
Anebocaris quadroculus Spence Bate, 1888: 722; Plate 123, fg. 1. [Of Port Jackson, Australia, surface; of
Sibago, Samboangan, Philippine Islands; of Basilan Strait; China Sea, of Luzon; Zebu Harbour,
Philippine Islands]
Anisocaris puerilis Coutire, 1907c: 410; unnumbered fgure. [Gorgie du Sud]
Astacus cylindrus Fabricius, 1793: 483. [in Oceani Indico]
Astacus planatus Fabricius, 1798: 482. [Type locality not indicated]
Atlantocaris longirostris Ortmann, 1893: 80; Plate 5, fg. 3. [Plankton-Expedition der Humboldt-Stifung,
JN 153 (7.9N 21.4W), im Guineastrom]
Camptocaris maxima Ortmann, 1893: 81; Plate 5, fg. 4. [Plankton-Expedition der Humboldt-Stifung, Sar-
gasso-See, JN 60 (35.0N 62.1W, 0-300 m); Sdl Aequatorialstrom, JN 188 (2.6S 14.6W, 0-400 m)]
Caricyphus acutus Coutire, 1905a: 21; Fig. 7. [Princesse-Alice stn 1834 (3728'N 2553'30"W), 0-1000 m]
Caricyphus angulatus Spence Bate, 1888: 718; Plate 121, fg. 6. [4646'S 4531'E, near Marion Island, sur-
face]
Caricyphus bigibbosus Coutire, 1905c: 1114. [fully described in Coutire, 1905a: 26; Fig. 8]. [Princesse-Al-
ice stn 1851 (4615'N 709'W), 0-3000 m]
Caricyphus cornutus Spence Bate, 1888: 712; Plate 121, fg. 2. [near Samboangan, Philippine Islands]
Caricyphus edentulus Ortmann, 1893: 74; Plate 2, fg. 3. [Plankton-Expedition der Humboldt-Stifung,
Sargasso-See, JN 72 (31.5N 59.0W, 0 m); 102 (31.7N 42.7W, 0-400 m); Pl.55 (28.3N 34.3W, 200 m,
in table as Nrdl. Aequatorialstrom und Canarienstrom)]
Caricyphus gibberosus Spence Bate, 1888: 716; Plate 121, fg. 4. [near the Sandwich Islands, Pacifc Ocean,
surface]
Caricyphus serramarginis Spence Bate, 1888: 714; Plate 121, fg. 3. [Kandavu, Fii Islands, surface]
Caricyphus targidus Spence Bate, 1888: 717; Plate 121, fg. 5. [of Luzon, China Sea, surface]
Caulurus pelagicus Stimpson, 1860a: 42. [in Oceano Pacifco, lat. Bor. 34 long. orient. 126]
Copiocaris messinensis Thiele, 1905: 454; Plate 15, Figs 17-30. [Strae von Messina]
Coronocaris brevis Ortmann, 1893: 82; Plate 5, fg. 5. [Plankton-Expedition der Humboldt-Stifung, Sdl.
Aequatorialstrom JN 234 (0.4S 42.4W, 0 m); 235 (0.1S 44.2W, 0-400 m); 249 (5.6S 44.0W, 0 m);
Brasilianische Kstenbank, Pl. 112 (0.4S 46.6W, 207 m); Pl. 113 (0.4S 46.6W, 200 m)]
Coronocaris gracilis Ortmann, 1893: 81; Plate 6, fg. 1. [Plankton-Expedition der Humboldt-Stifung,
Nrdl. Aequatorialstrom JN 145 (13.3N 22.7W, 0-400 m); 146 (12.3N 22.3W, 0-400 m); 148 (10.2N
22.2W, 0-400 m); 150 (10.2N 22.2W, 0-1000 m); Sdl. Aequatorialstrom JN 232 (0.4S 42.4W, 0-400
m); 234 (0.4S 42.4W, 0 m); 235 (0.1S 44.2W, 0-400 m); 246 (0.4S 46.6W, 0-400 m); Pl. 103 (0.4S
42.4W, 200 m); Brasilianische Kstenbank, Pl. 112 (0.4S 46.6W, 207 m)]
Coronocaris humilis Coutire, 1907a: 58; Fig. 21. [Aores, 0-1000 m]
Diaphoropus longidorsalis Spence Bate, 1888: 688; Plate 117, fg. 4. [Cape Verde Islands]
Diaphoropus versipellis Spence Bate, 1888: 687; Plate 107, fg. 3. [Of Cape Horn, Australia, surface]
Eretmocaris corniger Spence Bate, 1888: 900; Plate 145, fg. 4. [Cape Verde]
Eretmocaris dolichops Ortmann, 1893: 79; Plate 5, fg. 1. [Plankton-Expedition der Humboldt-Stifung, bei
Boavista, JN 141 (16.1N 23.1W, 0-500 m)]
Eretmocaris longicaulis Spence Bate, 1888: 897; Plate 145, fg. 2. [1729'N 141'31E, south of Japan, sur-
face]
Eretmocaris stylorostris Spence Bate, 1888: 898; Plate 145, fg. 3. [Of Cape Verde Islands, surface.]
Falcicaris tenuis Ortmann, 1893: 74; Plate 3, fg. 6. [Plankton-Expedition der Humboldt-Stifung, Flori-
dastrom JN 55 (37.9 N 59.1W, 0-400 m); Sdl. Aequatorialstrom JN 213 (5.7S 26.5W, 0-440 m) and
218 (3.8S 32.6W, 0-400 m)]
Hectarthropus compressus Spence Bate, 1888: 891; Plate 144, fg. 3. [Pacifc, between Api and Cape York,
surface]
Hectarthropus exilis Spence Bate, 1888: 889; Plate 144, fg. 2. [of Basilan Strait, Philippine Islands, surface]
Hectarthropus expansus Spence Bate, 1888: 893; Plate 144, fgs 4-5. [of Basilan Strait, Philippine Islands,
surface]
Hectarthropus nikiformis Coutire, 1907a: 34; Fig. 11. [SAS Le Prince de Monaco stn 1715, Canary Islands,
prs de Tnrife, 0-1000 m]
Hectarthropus tenuis Spence Bate, 1888: 893; Plate 144, fg. 6. [North Atlantic, probably near the Cape
Verde Islands, surface]
Icotopus amplissimus Coutire, 1907a: 23; Fig. 7. [SAS Le Prince de Monaco stns 1715, Canary Islands,
0-1000 m; 1760, Canary Islands, 0-3000 m; 2276, au voisinage des Balares, 1-1700 m]
Icotopus amproxima Coutire, 1907a: 27; Fig. 8. [SAS Le Prince de Monaco stn 1715, Canary Islands,
0-1000 m]
Icotopus arcurostris Spence Bate, 1888: 886; Plate 144, fg. 1. [of Cape Howe, Australia, surface]
Kyptocaris oligodon Coutire, 1907a: 29; Fig. 9. [SAS Le Prince de Monaco stn 1874, Aores, 0-2000 m]
Kyptocaris stylofrontalis Spence Bate, 1888: 690; Plate 121, fg. 1. [of Sibago, Samboangan, Philippine Is-
lands, surface]
Mesocaris recurva Ortmann, 1893: 82; Plate 5, fg. 6. [Plankton-Expedition der Humboldt-Stifung,
Brasilianische Kstenbank, JN 246 (0.4S 46.6W, 0-400 m)]
Odontolophus serratus Spence Bate, 1888: 665. [of Malta, Mediterranean]
Oligocaris bispinosa Ortmann, 1893: 85; Plate 7, fg. 2. [Plankton-Expedition der Humboldt-Stifung,
Irminger-See, JN 9 (60.2N 22.7W), JN 15 (60.3N 27.0W)]
Oligocaris (?) brevirostris Coutire, 1907a: 32; Fig. 10. [SAS Le Prince de Monaco stn 1781, Aores,
0-5000 m]
Pandacaricyphus pandaliformis Coutire, 1907a: 21; Fig. 6. [SAS Le Prince de Monaco stn 1834, rgion de
Aores, 0-1000 m]
Posydon depressus Fabricius, 1798: 417. [in Oceano Indico]
Procletes ellioti Spence Bate, 1888: 885. [of Waltair, on the coast of Coromandel]
Retrocaris antarcticus Coutire, 1907c: 411; unnumbered fgure. [Gorgie du Sud]
Retrocaris contraria Ortmann, 1893: 83; Plate 5, fg. 7. [Plankton-Expedition der Humboldt-Stifung, Sdl.
Aequatorialstrom JN 226 (2.4S 36.4W, 0 m); 235 (0.1S 44.2W, 0-400 m); Brasilianische Ksten-
bank, Pl. 112 (0.4S 46.6W, 207 m)]
Retrocaris serrata Nobili, 1906c: 54; Plate 4, fg. 12. [Port dAden]
Rhomaleocaris hamulus Spence Bate, 1888: 720: Plate 123, fg. 2. [on the passage from Api, New Hebrides,
to Cape York, Pacifc, surface]
Caridea Nomina Dubia
Alpheus avarus Fabricius, 1798: 404. [in India orientali]
Alpheus bispinosus Streets, 1871a: 242. [Isthmus of Panama]
Alpheus favescens Latreille, 1806: 53. [in Indi Orientalis Oceano]
Alpheus galathea Miers, 1874: 5; Plate 4, fg. 4. [Port Essington]
= Alpheus Galathea White, 1847a: 75. [Port Essington; nomen nudum]
Alpheus hoplites Nobili, 1906b: 257. [les Touamotou]
Alpheus Jourdainii Gurin-Mneville, 1857: 51. [Type locality not indicated]
Alpheus parvimanus Kingsley, 1878b: 195. [Panama]
Alpheus Saulcyi Gurin-Mneville, 1855 [in Gurin-Mneville, 1855-1856]: xviii; Plate 2, fg. 8. [Cuba]
Alpheus Savignyi Gurin-Mneville, 1857: 49. [nomen novum for Athanas nitescens Audouin, 1826 nec
Leach, 1813 [in Leach, 1813-1814]]
Alpheus sculptimanus Gurin-Mneville, 1857: 49. [De Janaon]
Alpheus setimanus Cano, 1888: 180. [provenienza ?]
Alpheus spinicaudus Lockington, 1878b: 476. [Port Escondido, Gulf of Cal.]
Alpheus tamulus Fabricius, 1798: 405. [in Oceano Indico]
Alpheus tenuimanus Lockington, 1878b: 473. [Port Escondido, Gulf of California]
Alphous [sic] Gundlach Torralbas, 1917: 72; Figs 50-51. [Cuba]
Anchistia antennata Hansen, 1925: 140. [Type locality not indicated]
Anchistia brachiata Stimpson, 1860a: 39. [Portu Lloyd ad insulas Bonin]
Anchistia Danae Stimpson, 1860a: 39. [Ins. Tahiti, in corrallis]
Astacus carinatus Meuschen, 1778: 86. [Type locality not indicated]
Astacus ferratus Fabricius, 1793: 486. [in Oceano Norwegico]
Astacus Gibbosus Pennant, 1812: 32. [Type locality not indicated]
Astacus Squilla major Meuschen, 1778: 86. [Type locality not indicated]
Astacus tetigonus Fabricius, 1775: 417. [Islandia]
Betaeus vladivostokiensis Vinogradov, 1950: 198; Plate 10, fg. 31. [Zolotoi Bay, near Vladivostok]
Caridina africana forma brevis Roth-Woltereck, 1984: 111. [Itimbiri]
Caridina africana forma longa Roth-Woltereck, 1984: 111. [Mwerusee]
Caridina africana nigerdeltae Roth-Woltereck, 1984: 117. [Nigerdelta]
Caridina barombi Roth-Woltereck, 1984: 110. [Barombisee, Kamerun]
Caridina powelli Roth-Woltereck, 1984: 110. [Nigerdelta]
Caridina weberi var. celebensis Schenkel, 1902: 499. [Tabela Bach, System des Kalaena, Luwu]
Cr.[yptophthalmus] ventricosus Costa, 1871 [in O.G. Costa & A. Costa, 1838-1871]: 4; Plate 7, fgs 3A-B.
[trovato nel Golfo di Taranto e sul faro di Messina]
Crangon marginatus Fabricius, 1798: 410. [Isle de France]
Crangon monopodium Bosc, 1802: 96; Plate 13, fg. 2. [la mer des Indes]
Harpilius spinuliferus Miers, 1884b: 203. [from the interior of Tridacna without precise indication of locality]
Hippolite afnis Owen, 1839: 90; Plate 27, fg. 4. [Monterey, California]
Hippolyte gracilipes Randall, 1840: 142. [Sandwich Islands]
Hippolyte ignobilis Kinahan, 1858c: 131. [Port Philip]
Hippolyte Marioni Gourret, 1887a: 1033. [graviers et sables vaseux qui stendent depuis lle de Riou
jusquau del de Planier, 108 m]
Hymenodora parva Coutire, 1905c: 1114. [Princesse-Alice stns 1834 (3728'N 2553'30"W), 1000-0 m; 1851
(4615'N 709'W), 3000-0 m]
Latreutes ceylonensis Pearson, 1905: 81; Plate 2, fg. 7. [10-12 miles north of Cheval Paar and 12 miles west
of Vankali Church, 7.5-9 fms]
Leander distans Heller, 1862a: 526. [Nicobaren]
Leander hammondii Kingsley, 1883: 108; Plate 1, fg. 2. [Bakers Island, North Pacifc]
Leander indicus Heller, 1862b: 413. [Java und Borneo]
Leptalpheus petronii Ramos-Porto & de Souza, 1994: 22. [Ilha de Itamaraca, Pernambuco]
Macrobrachium parasintangense Nguyn, 2004: 24. [Tri An Lake]
Merhippolyte orientalis Spence Bate, 1888: 621. [541'0"S, 1344'30"E, of New Guinea, 800fms]
Odontocerus lutescens Leach, 1830b: 170. [shores of Guinea]
Opithiocheirus Chrysophthalmus Leach, 1830a: 172. [Atlantic ocean near the river Congo]
Palaemon (Eupalaemon) cognatus Roux, 1927b: 322; Fig. 1. [Pionierbivak, Fl. Mamberamo]
Palaemon (Parapalaemon) stresemanni Roux, 1918: 113; Figs 1-2. [Tjelukan Bawang, N.-O. Bali]
Palaemon adriaticus O.G. Costa & A. Costa, 1871 [in O.G. Costa & A. Costa, 1838-1871]: 7. [Golfo di
Napoli]
Palaemon Audouini Heller, 1861: 26. [im rothen Meere]
Palaemon brevimanus Fabricius, 1798: 403. [in India orientali]
Palaemon brevirostris Andrzeiowksi, 1839: 22. [Type locality not indicated]
Palaemon coromandelianus Fabricius, 1798: 403. [in India orientali]
Palmon favescens Olivier, 1811: 667. [Nouvelle-Hollande]
Palmon fuvialis Streets, 1871b: 227; Plate 2, fgs 5-5a. [Tributary of the Coatzacoalcos River among the
Cordilleras]
Palaemon noctilucus Tilesius, 1819: 5; Plate 21a, fg. 2. [placed on the Ofcial Index of Rejected and Invalid
Works in Zoological Nomenclature in Opinion 864 in 1969]
Palaemon parvulus O.G. Costa & A. Costa, 1871 [in O.G. Costa & A. Costa, 1838-1871]: 7. [Golfo di Napoli]
Palaemon parvus Olivier, 1811: 666. [dans la Mditerrane]
Palaemon rosalesi Rodrguez de la Cruz, 1965: 100; Plate 7. [Ciudad del Carmen, arroyo de agua salobre
(cerca Av. Ultimo Paseo), Estado de Campeche]
Palaemon tranquebaricus Fabricius, 1798: 403. [in India orientalis]
Palaemonetes (Palaemonetes) granulosus Rodrguez de la Cruz, 1965: 89; Plate 6. [Laguna Las Palomas,
Tampico, Tam.]
Palaemonetes natalensis Stebbing, 1915: 78; Plate 19. [Cape Natal N. by E. 24 miles; depth 440 fms]
Pandalus bipennus Poisson, 1947: 18. [lile Maurice]
Pandalus quadridentata A. Milne-Edwards, 1883: Plate 19. [Travailleur dragage 54, 3240'20''N 1854'30''W,
400 m]
Plesionika capreensis Lo Bianco, 1903: 186. [vicinanze di Capri]
Pontonia maculata Stimpson, 1860a: 38. [ad insulas Bonin, in Tridacnis]
Pontonia unidens Kingsley, 1880: 422; Plate 14, fg. 9. [Key West, Fla.]
Prionorhynchus Cranchianus Leach, 1830b: 171. [Atlantic Ocean Lat. 2413' Long. 1851'15'', on the surface
of the sea]
Spirontocaris makrognathus Stebbing, 1921b: 19; Plate 4. [Durban waters]
Synalpheus brachyceros Nobili, 1906b: 256. [Makatea]
Typhlocaridina lingyunensis W.-X. Li & Luo, 2001: 72; Fig. 1. [cave in Shadong, Lingyun County, Guangxi]
Usterocheirus Brachypocoilion Leach, 1830a: 174. [the Atlantic, adhering to fucus natans]
Usterocheirus Macropocoilium Leach, 1830a: 173. [the Atlantic, adhering to fucus natans]
Zuphanusa Smithiana Leach, 1830a: 175. [the Atlantic, adhering to fucus natans]
Caridea Nomina Nuda
Acanthephyra pleuracantha Coutire, 1914: 886.
Alpheus albertei Kazmi & Kazmi, 1979: 154.
Alpheus Amphitrite White, 1847a: 74.
Alpheus Doris White, 1847a: 75.
Alpheus Faxoni Coutire, 1900b: 356.
Alpheus forceps White, 1847a: 75.
Alpheus intermedius Stimpson, 1860b: 444.
Alpheus isodactylus Afzal, Javed & Barkati, 1986: 339.
Alpheus isthmalleator Hurt, Anker & Knowlton, 2009: 526.
Alpheus manorensis Afzal, Javed & Barkati, 1986: 339.
Alpheus miyakei Miya, 1974: 106.
Alpheus ochrostriatus Karplus, Szlep & Tsurnamal, 1981: 5.
Alpheus pseudoedwardsii Afzal, Javed & Barkati, 1986: 338.
Alpheus purpurilenticularis Karplus & Ben Tuvia, 1979: 226.
Alpheus Rathbuni Coutire, 1900b: 356.
Alpheus Rhode White, 1847a: 74.
Alpheus rubromaculatus Karplus, Szlep & Tsurnamal, 1981: 5.
Alpheus Triton White, 1847a: 74.
Athanas arabicus Afzal, Javed & Barkati, 1986: 339.
Brachycarpus dentatus Nobili, 1907: 4.
Caricyphus acutus Coutire, 1905c: 1114.
Caridion monctoni Citarella, 1993: 15. [of Buctouche (4632'N 6432'30''W)]
Crangon cymodocea Price, 1971: 166.
Egeon tricuspidatum O.G. Costa, 1844c: 72.
Gasconella parvula de Saint Laurent, 1985: 473.
Hippolyte burunsis Bleeker, 1856: 37.
Hippolyte Hasseltii Bleeker, 1856: 64.
Hippolyte lygdamis White, 1847a: 76.
Hippolyte Macandreae Bell, 1847: 81.
Hippolyte metis White, 1847a: 76.
Hippolyte rubrosignata Wagner, 1885: 60.
Macrobrachium honnaense Jayachandran, 2010: 47.
Palmon Abbotii White, 1847a: 78.
Palmon aciculatus White, 1847a: 79.
Palaemon Afnis Hope, 1851: 17.
Palaemon afnis O.G. Costa & A. Costa, 1840 [in O.G. Costa & A. Costa, 1838-1871]: 4.
Palaemon Bipunctatus Hope, 1851: 17.
Palmon Brasiliensis White, 1847a: 79.
Palmon Colombicus White, 1847a: 78.
Palmon Creusa White, 1847a: 78.
Palaemon cubanus Spamer & Bogan, 1992: 150.
Palmon Cydippe White, 1847a: 78.
Palaemon Delaserii Hope, 1851: 17.
Palmon Electra White, 1847a: 78.
Palmon fuvialis Spence Bate, 1876: 378.
Palmon Glauce White, 1847a: 79.
Palmon Hypsa White, 1847a: 78.
Palmon Latreillei White, 1847a: 79.
Palmon Nicippe White, 1847a: 79.
Palmon Procles White, 1847a: 79.
Palemon pusillum Rafnesque, 1814: 24.
Palaemon Rathki Aurivillius, 1898b: 35.
Palaemon recticornis Sherborn, 1933: 785.
Palaemon seriatus Suckling, 1876: 270.
Palaemon Siamensis von Martens, 1868: 30.
Palaemon Splendens Hope, 1851: 17.
Palaemon splendens O.G. Costa & A. Costa, 1840 [in O.G. Costa & A. Costa, 1838-1871]: 4.
Palaemon walkeri OHara & Barmby, 2000: 43, 46.
Palemon Inermis Roux, 1831: 15.
Tizeuma Stimpsoni Perrier, 1886: 81.
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Bruce, A.J., 1967b. Notes on some Indo-Pacifc Pontoniinae III-IX. Descriptions of some new genera and
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Bruce, A.J., 1969a. Preliminary descriptions of sixteen new species of the genus Periclimenes Costa, 1844
(Crustacea, Decapoda Natantia, Pontoniinae). Zoologische Mededelingen 43: 253-278.
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1915 (Crustacea, Decapoda Natantia, Pontoniinae). Zoologische Mededelingen 44: 159-176.
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a new pontoniid shrimp from the Amirante Islands. Crustaceana 17: 141-150.
Bruce, A.J., 1970a. Report on some commensal pontoniid shrimps (Crustacea: Palaemonidae) associated
with an Indo-Pacifc gorgonian host (Coelenterata: Gorgonacea). Journal of Zoology, London 160:
537-544.
Bruce, A.J., 1970b. Notes on some Indo-Pacifc Pontoniinae. XV. Hamopontonia corallicola gen. nov., sp.
nov., a new pontoniid shrimp from Hong Kong. Crustaceana 18: 37-48.
Bruce, A.J., 1970c. Further preliminary descriptions of new species of the genus Periclimenaeus Borra-
daile, 1915, (Crustacea, Decapoda Natantia, Pontoniinae). Zoologische Mededelingen 44: 305-
315.
Bruce, A.J., 1971a. Periclimenes atenuatus sp. nov. (Crustacea Decapoda, Natantia, Pontoniinae), a new
commensal shrimp from the Duke of York Islands. Pacifc Science 25: 533-544.
Bruce, A.J., 1971b. Notes on some Indo-Pacifc Pontoniinae. XVII. Eupontonia noctalbata gen. nov., sp.
nov., a new pontoniid shrimp from Mah, the Seychelle islands. Crustaceana 20: 225-236.
Bruce, A.J., 1971c. Notes on some Indo-Pacifc Pontoniinae. XVI. Onycocaris seychellensis sp. nov., a new
species of shrimp from Mah. Crustaceana 20: 208-218.
Bruce, A.J., 1971d. Onycocaris zanzibarica sp. nov., a new pontoniid shrimp from East Africa. Journal of
Natural History 5: 293-298.
Bruce, A.J., 1971e. On a new commensal shrimp Periclimenes hirsutus sp. nov. (Crustacea, Decapoda
Natantia, Pontoniinae) from Fii. Pacifc Science 25: 91-99.
Bruce, A.J., 1972a. Pycnocaris chagoae gen. nov., sp. nov., a new shrimp from the Chagos Archipelago
(Decapoda Natantia, Gnathophyllidae). Crustaceana 23: 50-64.
Bruce, A.J., 1972b. Notes on some Indo-Pacifc Pontoniinae. XIX. Allopontonia iani gen nov., sp. nov., a
new echinoid associate from Zanzibar. Crustaceana 22: 1-12.
Bruce, A.J., 1972c. A report on a small collection of pontoniid shrimps from Fii, with the description of
a new species of Coralliocaris Stimpson (Crustacea, Decapoda, Natantia, Pontoniinae). Pacifc Sci-
ence 26: 63-86.
Bruce, A.J., 1972d. Notes on some Indo-Pacifc Pontoniinae, XX. Pontonia sibogae sp. nov., a new species
of Pontonia from eastern Australia and Indonesia (Decapoda Natantia, Palaemonidae). Crusta-
ceana 23: 182-186.
Bruce, A.J., 1972e. Notes on some Indo-Pacifc Pontoniinae, XXI. Typton bawii sp. nov., the frst occur-
rence of the genus Typton Costa in the Indian Ocean (Decapoda Natantia, Palaemonidae). Crus-
taceana 23: 243-254.
Bruce, A.J., 1973a. Gnathophylloides robustus sp. nov., a new commensal gnathophylloid shrimp from
Western Australia, with the designation of a new genus Levicaris (Decapoda, Caridea). Crusta-
ceana 24: 17-32.
Bruce, A.J., 1973b. Notes on some Indo-Pacifc Pontoniinae, XXIV. Dasycaris zanzibarica sp. nov. from the
western Indian Ocean, with remarks on other species of Dasycaris Kemp, 1922 (Decapoda Natan-
tia). Crustaceana 24: 247-260.
Bruce, A.J., 1973c. Notes on some Indo-Pacifc Pontoniinae. XXII. Pliopontonia furtiva gen. nov., sp. nov.,
a new shrimp associated with a corallimorph zoantharian. Crustaceana 24: 97-109.
Bruce, A.J., 1973d. Notes on some Indo-Pacifc Pontoniinae, XXIII. Tectopontonia maziwiae gen. nov., sp.
nov., a new coral associate from Tangaynika (Decapoda, Palaemonidae). Crustaceana 24: 169-180.
Bruce, A.J., 1973e. Typton australis sp. nov., a new pontoniid shrimp from the Great Barrier Reef, Aus-
tralia. Records of the Australian Museum 28: 253-263.
Bruce, A.J., 1974a. Coralliocaris viridis sp. nov., a preliminary note (Decapoda Natantia, Pontoniinae).
Bruce, A.J., 1974b. Periclimenes insolitus sp. nov. (Decapoda Natantia, Pontoniinae), a new commensal
shrimp from Waikiki Beach, Oahu, Hawaii. Crustaceana 26: 293-307.
Bruce, A.J., 1975a. Notes on some Indo-Pacifc Pontoniinae. XXVI. Neoanchistus cardiodytes gen. nov., sp.
nov., a new mollusc-associated shrimp from Madagascar (Decapoda, Palaemonidae). Crusta-
ceana 29: 149-165.
Bruce, A.J., 1975b. Further observations on the Indo-West Pacifc species of the genus Palaemonella Dana,
1852 (Decapoda Natantia, Pontoniinae). Crustaceana 29: 169-185.
Bruce, A.J., 1975c. Observations upon some specimens of the genus Periclimenaeus Borradaile (Decapoda
Natantia, Pontoniinae) originally described by G. Nobili. Bulletin du Musum national dHistoire
naturelle (3), Zoologie 258: 1557-1583.
Bruce, A.J., 1975d. Periclimenes colemani sp. nov., a new shrimp associate of a rare sea urchin from Heron
Island, Queensland (Decapoda Natantia, Pontoniinae). Records of the Australian Museum 29:
486-502.
Bruce, A.J., 1975e. Notes on some Indo-Pacifc Pontoniinae, XXV. Further observations upon Periclimenes
noverca Kemp, 1922, with the designation of a new genus Zenopontonia, and some remarks upon
Periclimenes parasiticus Borradaile (Decapoda Natantia, Palaemonidae). Crustaceana 28: 275-285.
Bruce, A.J., 1976a. Studies on Indo-West Pacifc Stenopodidea, 1. Stenopus zanzibaricus sp. nov., a new
species from East Africa. Crustaceana 31: 90-102.
Bruce, A.J., 1976b. Discias mvitae sp. nov., a new sponge associate from Kenya (Decapoda Natantia, Dis-
ciadidae). Crustaceana 31: 119-130.
Bruce, A.J., 1976c. Notes on some Indo-Pacifc Pontoniinae, XXVII. Apopontonia falcirostris gen. nov., sp.
nov., from Madagascar. Crustaceana 31: 301-311.
Bruce, A.J., 1976d. A report on a small collection of pontoniine shrimps from the Northern Indian
Ocean. Journal of the Marine Biological Association of India 16 [for 1974]: 437-454.
Bruce, A.J., 1976e. A report on some pontoniid shrimps collected from the Seychelle Islands by the F.R.V.
Manihine, 1972, with a review of the Seychelles pontoniid shrimp fauna. Zoological Journal of the
Linnean Society 59: 89-153.
Bruce, A.J., 1976f. A report on a small collection of shrimps from the Kenya National Marine Parks at
Malindi, with notes on selected species. Zoologische Verhandelingen 145: 1-72.
Bruce, A.J., 1977a. Pontoniine shrimps in the collections of the Australian Museum. Records of the
Bruce, A.J., 1977b. Periclimenes kororensis n. sp., an unusual shrimp associate of the fungiid coral, Helio-
fungia actiniformis. Micronesica 13: 33-43.
Bruce, A.J., 1977c. Notes on some Indo-Pacifc Pontoniinae, XXIX. Epipontonia spongicola gen. nov., sp.
nov., from Wasin Island, Kenya. Crustaceana 32: 304-315.
Bruce, A.J., 1977d. Notes on some Indo-Pacifc Pontoniinae, XXVIII. Typton wasini sp. nov., from Wasin
Island, Kenya. Crustaceana 32: 272-285.
Bruce, A.J., 1978a. A report on a collection of pontoniine shrimps from Madagascar and adjacent seas.
Zoological Journal of the Linnean Society 62: 205-290.
Bruce, A.J., 1978b. Paranchistus pycnodontae sp. nov., a new pontoniine shrimp associated with an ostreid
bivalve host. Memoirs of the Queensland Museum 18: 233-243, Plate 39.
Bruce, A.J., 1978c. The re-examination of some pontoniine shrimp types frst described by L.A. Borra-
daile (Decapoda, Palaemonidae). Crustaceana 34: 251-268.
Bruce, A.J., 1978d. Typton crosslandi sp. nov., a new pontoniine shrimp, from the Galapagos Islands.
Bruce, A.J., 1978e. Thor marguitae sp. nov., a new hippolytid shrimp from Heron Island, Australia.
Bruce, A.J., 1979a. Notes on some Indo-Pacifc Pontoniinae, XXXI. Periclimenes magnifcus sp. nov., a
coelenterate associate from the Capricorn Islands (Decapoda, Palaemonidae). Crustaceana Sup-
pl. 5: 195-208, Plate 1.
Bruce, A.J., 1979b. Ctenopontonia cyphastreophila, a new genus and species of coral associated pontoniine
shrimp from Eniwetok Atoll. Bulletin of Marine Science 29: 423-435.
Bruce, A.J., 1979c. Onycocaris furculata sp. nov., a new pontoniine shrimp from La Runion. Cahiers de
lIndo-Pacifque 1: 323-334.
Bruce, A.J., 1979d. A report on a small collection of pontoniine shrimps from Eniwetok Atoll. Crusta-
ceana Supplement 5: 209-230.
Bruce, A.J., 1979e. Onycocaris anomala sp. nov., a new pontoniine shrimp from the Northern Territory,
Australia. Records of the Australian Museum 32: 69-79.
Bruce, A.J., 1980a. On some Pontoniine shrimps from Noumea, New Caledonia. Cahiers de lIndo-
Pacifque 2: 1-39.
Bruce, A.J., 1980b. Notes on some Indo-Pacifc Pontoniinae, XXXIII. Periclimenaeus diplosomatis sp. nov.,
an ascidian associate from Heron Island, Australia. Crustaceana 39: 39-51.
Bruce, A.J., 1981a. Notes on some Indo-Pacifc Pontoniinae, XXXVI. Pontonia ardeae sp. nov., a new bi-
valve associate from the Capricorn Islands (Decapoda, Natantia). Crustaceana 40: 113-126.
Bruce, A.J., 1981b. Onycocaridella prima, new genus, new species, a new pontoniine sponge-associate
from the Capricorn Islands, Australia (Decapoda, Caridea, Pontoniinae). Journal of Crustacean
Biology 1: 241-250.
Bruce, A.J., 1981c. Notes on some Indo-Pacifc Pontoniinae, XXXVIII. Apopontonia dubia sp. nov., from a
southern Queensland sponge host. Crustaceana 41: 225-232.
Bruce, A.J., 1981d. Results of the MUSORSTOM expeditions. I. Philippines (18-28 March 1976). Deca-
pod Crustacea: Pontoniinae. Memoirs ORSTOM 91: 189-215.
Bruce, A.J., 1982a. Notes on some Indo-Pacifc Pontoniinae, XXXIX. Isopontonia platycheles gen. nov., sp.
nov., from the Chesterfeld Islands, New Caledonia (Decapoda, Caridea). Crustaceana 42: 54-64.
Bruce, A.J., 1982b. Notes on some Indo-Pacifc Pontoniinae, XLI. Orthopontonia, a new genus proposed
for Periclimenaeus ornatus Bruce. Crustaceana 43: 163-176.
Bruce, A.J., 1982c. The shrimps associated with Indo-West Pacifc echinoderms, with the descriptions of
a new species in the genus Periclimenes Costa, 1844 (Crustacea: Pontoniinae). Memoirs of the
Bruce, A.J., 1982d. Thorella cobourgi, new genus, new species, a hippolytid shrimp from the Northern
Territory, Australia. Journal of Crustacean Biology 2: 451-458.
Bruce, A.J., 1983a. A second species of the pontoniine shrimp genus Dasella Lebour, D. ansoni sp. nov.,
from the Arafura Sea. The Beagle, Records of the Northern Territory Museum of Arts and Sci-
ences 1: 21-29.
Bruce, A.J., 1983b. Epipontonia anceps n. sp., a sponge-associated pontoniine shrimp from Heron Island,
Queensland (Crustacea: Decapoda: Palaemonidae). Records of the Australian Museum 35: 19-28.
Bruce, A.J., 1983c. Expdition Rumphius II (1975). Crustacs parasites, commensaux, etc. (Th. Monod
d). IX. Crustacs Dcapodes (1
ere
partie: Natantia Pontoniinae). Bulletin du Musum national
dHistoire naturelle (4), section A, Zoologie, Biologie et cologie animales 5: 871-902.
Bruce, A.J., 1983d. Pseudathanas darwiniensis, new genus, new species, an alpheid shrimp from the
Northern territory, Australia. Journal of Crustacean Biology 3: 463-471.
Bruce, A.J., 1983e. Lysmata debelius new species, a new hippolytid shrimp from the Philippines. Revue
franaise dAquariologie 9 [for 1982]: 115-120.
Bruce, A.J., 1983f. Thor spinipes sp. nov., a new hippolytid shrimp from the Cobourg Peninsula, northern
Australia. The Beagle, Records of the Northern Territory Museum of Arts and Sciences 1: 1-10.
Bruce, A.J., 1983g. Miropandalus hardingi, new genus, new species, a bizarre commensal pandalid shrimp
from the Marshall Islands. Journal of Crustacean Biology 3: 482-490.
Bruce, A.J., 1984. Periclimenes dentidactylus, a new deep water pontoniine shrimp from Makassar Strait,
Indonesia. Marine Research in Indonesia 24: 7-17.
Bruce, A.J., 1985a. Notes on some Indo-Pacifc Pontoniinae, XLII. Miopontonia yongei gen. nov., sp. nov.,
from the Australian north west shelf (Decapoda, Caridea). Crustaceana 48: 167-178.
Bruce, A.J., 1985b. Decapod Crustacea: Pontoniinae (MUSORSTOM II). Mmoires du Musum na-
tional dHistoire naturelle (A) Zoologie 133: 229-260.
Bruce, A.J., 1986a. Two new species of Bathypalaemonella Balss (Crustacea, Decapoda, Campylonotidae)
from the Australian Northwest shelf. Zoologica Scripta 15: 251-264.
Bruce, A.J., 1986b. Chacella, a new palaemonid shrimp genus proposed for Dasycaris kerstitchi Wicksten,
1983 (Crustacea: Decapoda: Natantia). Journal of Crustacean Biology 6: 485-490.
Bruce, A.J., 1986c. Diapontonia maranulus, new genus, new species, a pontoniine shrimp associate of a
deep-water echinoid. Journal of Crustacean Biology 6: 125-133.
Bruce, A.J., 1986d. Three new species of commensal shrimps from Port Essington, Arnhem Land, north-
ern Australia (Crustacea: Decapoda: Palaemonidae). The Beagle, Records of the Northern Terri-
tory Museum of Arts and Sciences 3: 143-166.
Bruce, A.J., 1986e. Periclimenes milleri new species, a bathyal echinoid-associated pontoniine shrimp
from the Bahamas. Bulletin of Marine Science 39: 637-645.
Bruce, A.J., 1986f. Notes on some Indo-Pacifc Pontoniinae, XLIII. A new species of Typton from Ashmore
Reef, Timor Sea (Decapoda, Palaemonidae). Crustaceana 50: 278-286.
Bruce, A.J., 1987a. Notes on some Indo-Pacifc Pontoniinae, XLIV. Periclimenes darwiniensis sp. nov. from
the Northern Territory, Australia (Decapoda, Caridea). Crustaceana 52: 29-39.
Bruce, A.J., 1987b. Periclimenes johnsoni sp.nov., a new species of shrimp from Singapore (Crustacea:
Decapoda: Palaemonidae). Indo-Malayan Zoology 4: 113-126.
Bruce, A.J., 1987c. Onycocaridites anomodactylus, new genus, new species (Decapoda: Palaemonidae), a
commensal shrimp from the Arafura Sea. Journal of Crustacean Biology 7: 771-779.
Bruce, A.J., 1987d. Typton nanus sp. nov., a new commensal shrimp (Crustacea: Decapoda: Palaemoni-
dae) from the Australian north-west shelf. The Beagle, Records of the Northern Territory Muse-
um of Arts and Sciences 4: 49-56.
Bruce, A.J., 1987e. A new species of alpheid shrimp, Alpheus bannerorum from northern Australia. The
Beagle, Records of the Northern Territory Museum of Arts and Sciences 4: 61-72.
Bruce, A.J., 1988a. Two new palaemonid shrimps (Crustacea: Decapoda) from the Australian North
West shelf. Journal of Natural History 22: 1263-1276.
Bruce, A.J., 1988b. A new palaemonid shrimp from the Zostera-beds of Moreton Bay, Queensland, Aus-
tralia (Decapoda: Palaemonidae). The Beagle, Records of the Northern Territory Museum of Arts
and Sciences 5: 105-114.
Bruce, A.J., 1988c. Exopontonia malleatrix, new genus, new species, a palaemonid shrimp from Ashmore
Reef, Timor Sea. Journal of Crustacean Biology 8: 122-130.
Bruce, A.J., 1988d. A redescription of Periclimenaeus fmbriatus Borradaile, 1915, with the designation of
a new genus (Crustacea: Decapoda: Palaemonidae). Zoological Journal of the Linnean Society 94:
219-232.
Bruce, A.J., 1988e. Periclimenes tonga sp. nov., a commensal shrimp associated with a scyphozoan host
from Tonga (Crustacea: Decapoda: Palaemonidae). Micronesica 21: 23-32.
Bruce, A.J., 1988f. Bannereus anomalus, new genus, new species, a deep-sea alpheid shrimp from the
Coral Sea. Pacifc Science 42: 139-149.
Bruce, A.J., 1988g. Synalpheus dorae, a new commensal alpheid shrimp from the Australian northwest
shelf. Proceedings of the Biological Society of Washington 101: 843-852.
Bruce, A.J., 1988h. A new crangonid shrimp, Pontocheras arafurae gen. et sp. nov., from the Arafura Sea.
Zoologica Scripta 17: 213-221.
Bruce, A.J., 1989a. A report on some coral reef shrimps from the Philippine Islands. Asian Marine Biol-
ogy 6: 173-192.
Bruce, A.J., 1989b. Notes on some Indo-Pacifc Pontoniinae, XLV. Conchodytes maculatus sp. nov., a new
bivalve associate from the Australian northwest shelf. Crustaceana 56: 182-192.
Bruce, A.J., 1989c. Periclimenes gonioporae sp. nov. (Crustacea: Decapoda: Palaemonidae), a new coelen-
terate-associated shrimp. The Beagle, Records of the Northern Territory Museum of Arts and
Sciences 6: 149-156.
Bruce, A.J., 1989d. Periclimenes poupini sp. nov., a new anemone-associated shrimp from deep-water
traps (Crustacea, Decapoda, Palaemonidae). Bulletin du Musum national dHistoire naturelle,
4me srie, section A, Zoologie, Biologie et cologie animales 11: 851-863.
Bruce, A.J., 1990a. Two deep-sea shrimps new to the Australian fauna, Psathyrocaris hawaiiensis Rathbun
(Pasiphaeidae) and Bresilia antipodarum, sp. nov. (Bresiliidae), with remarks on Encantada spinocu-
lata Wicksten (Bresiliidae). Invertebrate Taxonomy 4: 847-866.
Bruce, A.J., 1990b. A second species of Bresilia, B. plumifera sp. nov., new to the Australian fauna (Crus-
tacea: Decapoda: Bresiliidae). The Beagle, Records of the Northern Territory Museum of Arts and
Sciences 7: 1-8.
Bruce, A.J., 1990c. Crustacea Decapoda: Deep-sea palaemonid shrimps from New Caledonian waters.
In: Crosnier, A. (ed.), Rsultats des Campagnes MUSORSTOM, vol. 6. Mmoires du Musum
national dHistoire naturelle (A) Zoologie 145: 149-215.
Bruce, A.J., 1990d. A new cnidarian-associated palaemonid shrimp from Port Essington, Cobourg Pen-
insula. Indo-Malayan Zoology 6 [for 1989]: 229-243.
Bruce, A.J., 1990e. Periclimenes franklini sp. nov., a new deep-sea shrimp from the Coral Sea (Crustacea:
Decapoda: Palaemonidae). The Beagle, Records of the Northern Territory Museum of Arts and
Sciences 7: 55-64.
Bruce, A.J., 1990f. Additions to the marine shrimp fauna of Hong Kong. In: Morton, B. (ed.), Proceedings
of the second international marine biological workshop: The marine fauna and fora of Hong Kong
and southern China, Hong Kong, 1986: 611-648. Hong Kong University Press, Hong Kong.
Bruce, A.J., 1990g. Redescriptions of fve Hong Kong carideans frst described by William Stimpson,
1860. In: Morton, B. (ed.), Proceedings of the second international marine biological workshop: The
marine fauna and fora of Hong Kong and southern China, Hong Kong, 1986: 569-610. Hong Kong
University Press, Hong Kong.
Bruce, A.J., 1990h. Crustacea Decapoda: Gelastreutes crosnieri gen. nov., sp. nov. (Hippolytidae) from
New Caledonia. In: Crosnier, A. (ed.), Rsultats des Campagnes MUSORSTOM, vol. 6. Mmoires
du Musum national dHistoire naturelle (A) Zoologie 145: 137-147.
Bruce, A.J., 1990i. Leontocaris amplectipes sp. nov. (Hippolytidae), a new deep-water shrimp from south-
ern Australia. Memoirs of the Museum of Victoria 51: 121-130.
Bruce, A.J., 1991a. Crustacea Decapoda: Further deep-sea palaemonoid shrimps from New Caledonian
waters. In: Crosnier, A. (ed.), Rsultats des Campagnes MUSORSTOM, vol. 9. Mmoires du Mu-
sum national dHistoire naturelle (A) Zoologie 152: 299-411.
Bruce, A.J., 1991b. Shallow-water palaemonoid shrimps from New Caledonia (Crustacea: Decapoda).
In: Richer de Forges, B. (ed.), Le benthos des Fonds Meubles des Lagons de Nouvelle-Caldonie, 1:
221-279. ORSTOM, Paris.
Bruce, A.J., 1991c. Notopontonia platycheles, new genus, new species (Decapoda: Pontoniinae) from South
Australia, with remarks on Pontonia pinnophylax (Oto), the type species of Pontonia Latreille.
Bruce, A.J., 1991d. The African shrimp genus Potamalpheops in Australia, with the description of
P.hanleyi, new species (Decapoda: Alpheidae). Journal of Crustacean Biology 11: 629-638.
Bruce, A.J., 1992a. Pycnisia raptor, a new genus and species of predatory troglobitic shrimp (Crustacea:
Decapoda: Atyidae) from northern Australia. Invertebrate Taxonomy 6: 553-566.
Bruce, A.J., 1992b. A re-description of Macrobrachium handschini (Roux, 1933) (Crustacea: Decapoda: Pal-
aemonidae). Hydrobiologia 231: 131-139.
Bruce, A.J., 1992c. Two new species of Periclimenes (Crustacea: Decapoda: Palaemonidae) from Liz-
ard Island, Queensland, with notes on related taxa. Records of the Australian Museum 44:
45-84.
Bruce, A.J., 1992d. Designation of two new pontoniine shrimp genera (Decapoda: Palaemonidae).
Journal of Natural History 26: 1273-1282.
Bruce, A.J., 1992e. Additions to the marine caridean fauna of Hong Kong, with a description of a new
species of Onycocaris (Crustacea: Decapoda: Palaemonidae) from Tuvalu. In: Morton, B. (ed.), The
marine fauna and fora of Hong Kong and southern China III. Proceedings of the Fourth Interna-
tional Marine Biological Workshop. The Marine Fauna and Flora of Hong Kong and southern Chi-
na: 329-343. Hong Kong University Press, Hong Kong.
Bruce, A.J., 1993a. Kakaducaris glabra gen. nov., sp. nov., a new freshwater shrimp from the Kakadu Na-
tional Park, Northern Territory, Australia, Crustacea: Decapoda: Palaemonidae with the designa-
tion of a new subfamily Kakaducaridinae. Hydrobiologia 268: 27-44.
Bruce, A.J., 1993b. Pontoniine shrimps from the Zoological Museum, Copenhagen. Journal of Natural
History 28: 829-840.
Bruce, A.J., 1993c. Potamalpheops darwiniensis (Crustacea: Decapoda: Alpheidae), the third Indo-West
Pacifc species. Proceedings of the Biological Society of Washington 106: 698-704.
Bruce, A.J., 1994a. Leander plumosus sp. nov., a new palaemonine shrimp (Crustacea: Palaemonidae)
from the Maldive Islands. The Beagle, Records of the Museums and Art Galleries of the Northern
Territory 11: 39-51.
Bruce, A.J., 1994b. Alpheus fenneri sp. nov. and A. williamsi sp. nov., two new Indo-West Pacifc alpheid
shrimps of the brevirostris species group. The Beagle, Records of the Museums and Art Galleries
of the Northern Territory 11: 15-28.
Bruce, A.J., 1994c. Shrimps from Flat-Top Bank, Timor Sea (Crustacea: Decapoda: Caridea). Rafes
Bulletin of Zoology 42: 743-756.
Bruce, A.J., 1995. A synopsis of the Indo-West Pacifc genera of the Pontoniinae (Crustacea: Decapoda:
Pontoniinae). Theses Zoologicae 25: 1-172 [imprint 1994, published 16st March 1995].
Bruce, A.J., 1996. Crustacea Decapoda: Palaemonoid shrimps from the Indo-West Pacifc region mainly
from New Caledonia. In: Crosnier, A. (ed.), Rsultats des Campagnes MUSORSTOM, vol. 15. M-
moires du Musum national dHistoire naturelle 168: 197-267.
Bruce, A.J., 1997a. A new pontoniine shrimp genus (Crustacea: decapoda) from the Yemen, with a note
on other species. Journal of Natural History 31: 1213-1222.
Bruce, A.J., 1997b. A new genus of hippolytid shrimp (Crustacea: Decapoda: Hippolytidae) for Thor
maldivensis Borradaile. Memoirs of the Queensland Museum 42: 13-23.
Bruce, A.J., 1998a. A new species of the genus Brachycarpus (Decapoda, Caridea, Palaemonidae) from
New Caledonia. Zoosystema 20: 157-165.
Bruce, A.J., 1998b. A second species of the genus Balssia Kemp, 1922 (Crustacea, Decapoda, Pontonii-
nae). Zoosystema 20: 603-611.
Bruce, A.J., 1998c. Pontoniine shrimps from Moreton Bay, Queensland (Crustacea: Decapoda: Pontonii-
nae). Memoirs of the Queensland Museum 42: 387-398.
Bruce, A.J., 1998d. A new name, Thinora, proposed for the preoccupied name Thorina Bruce (Crustacea:
Decapoda: Hippolytidae). Memoirs of the Queensland Museum 42: 398.
Bruce, A.J., 1999. Alpheus soror, a new snapping shrimp cryptospecies from Sri Lanka (Crustacea: De-
capoda: Alpheidae). Rafes Bulletin of Zoology 47: 453-463.
Bruce, A.J., 2000. Typton manningi and T. capricorniae, new species, new pontoniine shrimps from North-
ern Queensland, with a review of the Indo-West Pacifc species of Typton Costa (Decapoda: Palae-
monidae). Journal of Crustacean Biology 20 (Special number 2): 87-100.
Bruce, A.J., 2002a. Leander manningi, a new palaemonine shrimp from Western Australia (Crustacea,
Decapoda, Palaemonidae), with a review of the Indo-West Pacifc species of the genus Leander E.
Desmarest, 1849. Records of the Western Australian Museum 21: 71-81.
Bruce, A.J., 2002b. Notes on some Indo-Pacifc Pontoniinae, XLVI. Palaemonella foresti sp.nov., a new
pontoniine shrimp from Western Australia (Decapoda, Palaemonidae), with a review of the Indo-
West Pacifc species of the genus Palaemonella Dana, 1852. Crustaceana 75: 277-298.
Bruce, A.J., 2002c. A new species of Palaemonella (Crustacea: Decapoda: Pontoniinae) from East Africa.
The Beagle, Records of the Museums and Art Galleries of the Northern Territory 18: 15-18.
Bruce, A.J., 2003a. Periclimenes species (Crustacea: Decapoda: Pontoniinae) from far North Queens-
land. Memoirs of the Queensland Museum 49: 115-122.
Bruce, A.J., 2003b. A new species of Dactylonia Fransen (Crustacea: Decapoda: Pontoniinae) from East
Africa. Cahiers de Biologie Marine 44: 299-306.
Bruce, A.J., 2003c. Vir colemani sp. nov., a new commensal pontoniine shrimp (Crustacea: decapoda:
Palaemonidae) from Papua New Guinea. The Beagle, Records of the Museums and Art Galleries
of the Northern Territory 19: 119-124.
Bruce, A.J., 2004a. Balssia antipodarum sp. nov., the frst occurrence of the genus Balssia Kemp in the Indo-
West Pacifc region (Crustacea: Decapoda: Pontoniinae). Cahiers de Biologie Marine 45: 365-372.
Bruce, A.J., 2004b. A partial revision of the genus Periclimenes Costa, 1844 (Crustacea: Decapoda: Palae-
monidae). Zootaxa 582: 1-26.
Bruce, A.J., 2004c. A new pontoniine shrimp from Tahiti, French Polynesia (Crustacea, Decapoda, Palae-
monidae). Zoosystema 26: 279-289.
Bruce, A.J., 2005a. Bresiliid shrimps from the Red Sea (Crustacea: Decapoda: Caridea) with the descrip-
tion of a new species. Proceedings of the Biological Society of Washington 118: 176-182.
Bruce, A.J., 2005b. Pontoniine shrimps from Papua New Guinea, with designation of two new genera,
Cainonia and Colemonia (Crustacea: Decapoda: Palaemonidae). Memoirs of the Queensland Mu-
seum 51: 333-383.
Bruce, A.J., 2005c. New species of Periclimenaeus Borradaile (Crustacea: Decapoda: Pontoniinae) from
Ashmore reef, North Western Australia. Records of the Western Australian Museum 22: 325-342.
Bruce, A.J., 2005d. Pontoniine shrimps from the 2003 NORFANZ Expedition, 10 May-16 June (Crusta-
cea: Decapoda: Palaemonidae). Zootaxa 981: 1-20.
Bruce, A.J., 2005e. A new genus Pontoniopsides, for Pontoniopsis paulae Gore, 1981 (Crustacea: Decapoda:
Pontoniinae). Zootaxa 826: 1-4.
Bruce, A.J., 2006a. Pontoniine shrimps (Decapoda: Palaemonidae) from the island of Socotra, with de-
scriptions of new species of Dactylonia Fransen, 2002 and Periclimenoides Bruce, 1990. Zootaxa
1137: 1-36.
Bruce, A.J., 2006b. A new species of the pontoniine shrimp genus Harpilius Dana, 1852, H. spinifer (Crus-
tacea: Decapoda: Palaemonidae), from New Caledonia. Cahiers de Biologie Marine 47: 271-280.
Bruce, A.J., 2006c. Periclimenes jackhintoni sp. nov. (Crustacea: Decapoda: Palaemonidae), a new pontoni-
ine shrimp and crinoid associate from Tonga. The Beagle, Records of the Museums and Art Gal-
leries of the Northern Territory 22: 23-29.
Bruce, A.J., 2006d. A new genus Leptomenes, for the pontoniine shrimp Periclimenes dolichosternum Oku-
no & Mitsuhashi, 2003 (Crustacea: Decapoda: Palaemonidae). Cahiers de Biologie Marine 47:
223-225.
Bruce, A.J., 2006e. Periclimenaeus fawatu spec. nov. (Crustacea: Decapoda: Pontoniinae), from Zanzi-
bar. Zoologische Mededelingen 84: 33-43.
Bruce, A.J., 2006f. Periclimenaeus nielbrucei sp. nov. (Crustacea: Decapoda: Pontoniinae), a new sponge
associate from the Capricorn Islands, Queensland, with notes on related Periclimenaeus species.
Zootaxa 1224: 1-22.
Bruce, A.J., 2006g. An unusual new Periclimenes (Crustacea, Decapoda, Palaemonidae) from New Cale-
donia. Zoosystema 28: 703-712.
Bruce, A.J., 2006h. Periclimenes manihine sp. nov., a new pontoniine shrimp of the P.alcocki species group
(Crustacea: Decapoda: Palaemonidae). Zootaxa 1309: 45-54.
Bruce, A.J., 2007a. Re-examination of Borradailes Urocaris longicaudata specimens from the 1905 J.S.
Gardiner Collection (Crustacea: Decapoda: Pontoniinae). Zootaxa 1644: 51-57.
Bruce, A.J., 2007b. Leptomenaeus gen. nov., a new name for Leptomenes Bruce, 2006 (Crustacea: Decapoda:
Pontoniinae). Cahiers de Biologie Marine 48: 411-412.
Bruce, A.J., 2007c. A re-defnition of the genus Periclimenes Costa, 1844 and the designation of a new
genus Margitonia (Crustacea: Decapoda: Pontoniinae). Cahiers de Biologie Marine 48: 403-406.
Bruce, A.J., 2007d. Palaemonoid shrimps from the Dampier Archipelago (Crustacea: Decapoda), with a
review of the Western Australian pontoniine shrimp fauna. Records of the Western Australian
Museum Supplement 73: 97-129.
Bruce, A.J., 2007e. Periclimenes sarkanae sp. nov. a new pontoniine shrimp from Moreton Bay, Queens-
land (Crustacea: Decapoda: Palaemonidae). Zootaxa 1393: 61-68.
Bruce, A.J., 2008a. Periclimenes aqabai sp. nov., a further new species of the aesopius species group
(Caridea: Pontoniinae) from the Red Sea. Zootaxa 1682: 27-32.
Bruce, A.J., 2008b. A new species of Palaemonella Dana, 1852, (Crustacea: Decapoda; Pontoniinae) from
the Red Sea. Zootaxa 1844: 63-68.
Bruce, A.J., 2008c. Palaemonoid shrimps from the Australian north west shelf. Zootaxa 1815: 1-24.
Bruce, A.J., 2008d. A new pontoniine shrimp from the Great Barrier Reef (Crustacea. Decapoda. Palae-
monidae). Memoirs of the Queensland Museum 52: 89-98.
Bruce, A.J., 2008e. Phycomenes zostericola gen. nov., sp. nov., a new pontoniine shrimp (Crustacea: De-
capoda: Palaemonidae) from Moreton Bay, Queensland. Memoirs of the Queensland Museum 54:
219-232.
Bruce, A.J., 2009a. Comments on the generic position of Typton australis Bruce, 1973, and some related
taxa (Crustacea: Decapoda: Pontoniinae). Zootaxa 2076: 60-62.
Bruce, A.J., 2009b. Plesiomenaeus poorei gen. nov., sp. nov., (Crustacea: Decapoda: Pontoniinae) from Zan-
zibar. Memoirs of Museum Victoria 66: 25-34.
Bruce, A.J., 2009c. A re-description of Typton spongicola Costa, 1844, the type species of the genus Typton
Costa, 1844 (Crustacea: Decapoda: Pontoniinae). Cahiers de Biologie Marine 50: 382-394.
Bruce, A.J., 2010a. Allopontonia alastairi sp. nov., a second species of the genus Allopontonia Bruce, 1972
(Crustacea: Decapoda: Pontoniinae), from the Australian Northwest Shelf. In: De Grave, S. &
C.H.J.M. Fransen (eds.), Contributions to shrimp taxonomy. Zootaxa 2372: 33-36.
Bruce, A.J., 2010b. Ancylomenes kuboi and A.okunoi spp. nov. (Decapoda: Pontoniinae), from the Austral-
ian Northwest Shelf, Vietnam and the Philippines. In: De Grave, S. & C.H.J.M. Fransen (eds.), Con-
tributions to shrimp taxonomy. Zootaxa 2372: 169-176.
Bruce, A.J., 2010c. A revision of the systematic position of Periclimenaeus spinimanus Bruce, 1969 (Crusta-
cea: Decapoda: Pontoniinae) and the designation of Anisomenaeus gen. nov. In: De Grave, S. &
Bruce, A.J., 2010d. Eupontonia oahu sp. nov., a second species of the genus Eupontonia Bruce, 1971, (Crus-
tacea: Decapoda: Pontoniinae) from Oahu, Hawaiian Islands. In: De Grave, S. & C.H.J.M. Fransen
(eds.), Contributions to shrimp taxonomy. Zootaxa 2372: 405-414.
Bruce, A.J., 2010e. Palaemonella dionesae sp. nov. (Crustacea: Decapoda: Pontoniinae) from Western Aus-
tralia. In: De Grave, S. & C.H.J.M. Fransen (eds.), Contributions to shrimp taxonomy. Zootaxa
2372: 151-156.
Bruce, A.J., 2010f. More pontoniine shrimps (Crustacea: Decapoda: Palaemonidae) from the CReefs 2009
Heron Island expedition. Zootaxa 2604: 20-36.
Bruce, A.J., 2010g. Periclimenaeus devaneyi sp. nov., from Oahu, Hawaii (Crustacea: Decapoda: Pontonii-
nae). In: De Grave, S. & C.H.J.M. Fransen (eds.), Contributions to shrimp taxonomy. Zootaxa
2372: 379-388.
Bruce, A.J., 2010h. Periclimenes nevillei sp. nov. (Crustacea: Decapoda: Pontoniinae) from Vanuatu.
Zootaxa 2510: 45-54.
Bruce, A.J., 2010i. Pontoniine shrimps (Crustacea: Decapoda: Palaemonidae) from the CReefs 2009 Her-
on Island Expedition, with a review of the Heron island pontoniine fauna. Zootaxa 2541: 50-68.
Bruce, A.J., 2010j. Typtonoides nieli gen. nov., sp. nov., a new pontoniine shrimp (Crustacea: Decapoda:
Palaemonidae) from the Chesterfeld Island. The Beagle, Records of the Museums and Art Galler-
ies of the Northern Territory 26: 69-74.
Bruce, A.J., 2010k. Lebbeus rubrodentatus sp. nov. (Crustacea: Caridea: Hippolytidae) from the Australian
North West Shelf. The Beagle, Records of the Museums and Art Galleries of the Northern Terri-
tory 26: 75-78.
Bruce, A.J., 2011a. Notes on some Indo-Pacifc Pontoniinae, XLVIII. Onycocaris nieli sp. nov., a new pon-
toniine shrimp from Heron Island, Queensland. Crustaceana 84: 319-330.
Bruce, A.J., 2011b. Notes on some Indo-Pacifc Pontoniinae, XLIX. Onycocaris balsii sp. nov., from north-
ern Australia, with the designation of O. fujinoi sp. nov. Crustaceana 84: 477-490.
Bruce, A.J. & K. Baba, 1973. Spongiocaris, a new genus of stenopodidean shrimp from New Zealand and
South African waters, with a description of two new species (Decapoda Natantia, Stenopodidea).
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Holthuis, L.B., 1977b. Cave shrimps (Crustacea Decapoda, Natantia) from Mexico. Quaderno del Ac-
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Holthuis, L.B., 1978a. A collection of Decapod Crustacea from Sumba, Lesser Sunda islands, Indone-
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Holthuis, L.B., 1978b. Zoological results of the British speleological expedition to Papua New Guinea
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Holthuis, L.B., 1980d. Alpheus saxidomus new species, a rock boring snapping shrimp from the Pacifc
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Holthuis, L.B., 1982. Notes on Indo-West Pacifc Crustacea Decapoda I and II. Crustaceana 42: 26-36.
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Holthuis, L.B., 1984b. Freshwater prawns (Crustacea Decapoda: Natantia) from subterranean waters of
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Holthuis, L.B., 1986a. Fresh-water shrimps of the family Atyidae (Crustacea: Decapoda) from western
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Holthuis, L.B., 1986b. A new genus and species of subterranean shrimp from Western Australia (Crus-
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Received: 16.v.2011
Accepted: 15.vi.2011
Edited: L.P. van Ofwegen
Acanthanas ............................. 376
Acanthephyra ......................... 260
Acanthephyridae ................ 260
Acetes ...................................... 239
Aegaeon ................................... 454
Agostocarididae .................. 291
Agostocaris ............................. 291
Alainopasiphaea ..................... 254
Alcockpenaeopsis ................... 213
Alcyonohippolyte ................... 414
Allopontonia ........................... 339
Allosergestes ........................... 240
Alope ....................................... 414
Alpheidae .............................. 376
Alpheopsis ............................... 376
Alpheus ................................... 377
Altelatipes ............................... 209
Altopontonia ........................... 339
Alvinocarididae ................... 292
Alvinocaridinides ................... 292
Ambidexter ............................. 436
Amphibetaeus ......................... 391
Amphipontonia ...................... 339
Anachlorocurtis ..................... 440
Anapontonia ........................... 340
Anchiopontonia ...................... 340
Anchistioides .......................... 308
Anchistioididae ................... 308
Anchistus ................................ 340
Ancylomenes .......................... 341
Anisomenaeus ........................ 342
Antecaridina ........................... 267
Apopontonia ........................... 342
Arachnochium ........................ 314
Arafurapenaeopsis ................. 213
Araiopontonia ........................ 342
Archaeatya .............................. 267
Arete ........................................ 391
Aretopsis ................................. 391
Argis ........................................ 454
Aristaeomorpha ...................... 206
Aristaeopsis ............................ 206
Aristeidae .............................. 206
Aristeus ................................... 206
Ascidonia ................................ 342
Athanas ................................... 391
Athanopsis .............................. 391
Atlantopandalus .................... 440
Atya ......................................... 267
Atyaephyra ............................. 268
Atydina ................................... 268
Atyella ..................................... 268
Atyidae .................................. 267
Atyoida .................................... 269
Atyopsis .................................. 270
Atypopenaeus ......................... 213
Australatya ............................. 270
Austropandalus ...................... 440
Austropenaeus ........................ 207
Automate ................................ 393
Balssia ..................................... 343
Bannereus ............................... 394
Barbouria ................................ 412
Barbouriidae ......................... 412
Batella ...................................... 394
Batepenaeopsis ....................... 213
Bathyhippolyte ....................... 414
Bathypalaemonella ................. 306
Bathypalaemonellidae ....... 306
Bathypalaemonetes ................ 307
Bathystylodactylus ................ 304
Bentheogennema .................... 209
Benthesicymidae ................. 209
Benthesicymus ....................... 209
Benthonectes ........................... 211
Bermudacaris .......................... 394
Betaeopsis ................................ 394
Betaeus .................................... 395
Birulia ..................................... 414
Bitias ....................................... 440
Blepharocaris .......................... 343
Brachycarpus .......................... 315
Bresilia .................................... 294
Bresiliidae ............................. 294
Brucecaris ............................... 343
Bruceonia ................................ 343
Bruceopsis ............................... 395
Bythocarides ........................... 414
Bythocaris ............................... 415
Cainonia .................................. 343
Calathaemon ........................... 313
Calipandalus ........................... 440
Calliasmata ............................. 415
Campylonotidae .................. 307
Campylonotus ........................ 307
Caridella .................................. 270
Caridina .................................. 270
Caridinides ............................. 282
Caridinopsis ........................... 282
Caridion .................................. 415
Carinopontonia ...................... 344
Chacella ................................... 344
Chelonika ................................ 440
Chernocaris ............................. 344
Chlorocurtis ............................ 440
Chlorotocella ........................... 441
Chlorotocoides ........................ 452
Chlorotocus ............................. 441
Chorismus ............................... 416
Chorocaris ............................... 293
Cinetorhynchus ...................... 300
Climeniperaeus ...................... 344
Clytomanningus .................... 436
Colemonia ............................... 344
Conchodytes ........................... 344
Coralliocaris ........................... 345
Coronalpheus .......................... 396
Coutieralpheus ....................... 396
Coutierea ................................. 346
Coutierella .............................. 315
Crangon .................................. 455
Crangonidae ......................... 454
Creaseria ................................. 316
Crinotonia ............................... 346
Cryphiops ................................ 316
Cryptocheles ........................... 416
Cryptopenaeus ....................... 232
Ctenopontonia ........................ 346
Cuapetes .................................. 346
Dactylonia .............................. 348
Dasella ..................................... 348
Dasycaris ................................ 348
Deioneus ................................. 396
Deosergestes ........................... 241
Desmocarididae .................. 309
Desmocaris ............................. 309
Diapontonia ............................ 348
Dichelopandalus .................... 442
Disciadidae ........................... 295
Discias ..................................... 295
Dorodotes ................................ 442
Dugastella ............................... 282
Edoneus ................................... 282
Encantada ............................... 295
Engystenopus ......................... 249
Ephyrina ................................. 263
Epipontonia ............................ 349
Eualus ..................................... 416
Eugonatonotidae ................. 296
Eugonatonotus ....................... 296
Eumanningia .......................... 418
Eupasiphae .............................. 255
Eupontonia ............................. 349
Systematic index to currently valid families and genera
Euryrhynchidae ................... 309
Euryrhynchina ....................... 309
Euryrhynchoides .................... 309
Euryrhynchus ........................ 309
Eusergestes ............................. 242
Exhippolysmata ...................... 418
Exoclimenella ......................... 349
Exopalaemon .......................... 316
Exopontonia ............................ 349
Farfantepenaeus ..................... 214
Fennera ................................... 349
Fenneralpheus ........................ 396
Fenneropenaeus ...................... 214
Funchalia ................................ 214
Gallocaris ................................ 282
Ganjampenaeopsis ................. 215
Gelastocaris ............................ 419
Gelastreutes ............................ 419
Gennadas ................................ 211
Globospongicola ..................... 249
Glyphocrangon ....................... 467
Glyphocrangonidae ............ 467
Glyphus ................................... 255
Gnathophylleptum ................. 309
Gnathophyllidae ................. 309
Gnathophylloides ................... 311
Gnathophyllum ...................... 311
Gordonella .............................. 232
Hadropenaeus ......................... 232
Haliporoides ............................ 233
Haliporus ................................ 233
Halocaridina ........................... 282
Halocaridinides ...................... 283
Hamiger .................................. 349
Hamodactyloides .................... 349
Hamodactylus ........................ 350
Hamopontonia ........................ 350
Harperalpheus ........................ 396
Harpiliopsis ............................ 350
Harpilius ................................. 350
Hayashidonus ......................... 436
Hemipenaeus .......................... 207
Hepomadus ............................. 208
Heptacarpus ........................... 419
Heterocarpus .......................... 442
Heterogenys ............................ 263
Heteropenaeus ........................ 215
Hippolyte ................................ 420
Hippolytidae ........................ 414
Holthuisaeus ........................... 351
Holthuispenaeopsis ................ 215
Hymenocera ............................ 313
Hymenoceridae ................... 313
Hymenodora ........................... 263
Hymenopenaeus ..................... 233
Ischnopontonia ....................... 351
Isopontonia ............................. 351
Izucaris .................................... 351
Janicea ..................................... 413
Janicella ................................... 265
Jengalpheops ........................... 397
Jocaste ...................................... 351
Jolivetya .................................. 283
Jonga ........................................ 283
Juxtastenopus ......................... 252
Kakaducarididae ................. 314
Kakaducaris ............................ 314
Kemphyra ................................ 264
Kirnasia ................................... 296
Kishinouyepenaeopsis ........... 215
Lancaris ................................... 283
Laomenes ................................. 351
Latreutes ................................. 423
Leander .................................... 317
Leandrites ............................... 318
Lebbeus .................................... 424
Lenzicarcinus ......................... 297
Leontocaris .............................. 427
Leptalpheus ............................. 397
Leptathanas ............................ 398
Leptocarpus ............................ 318
Leptochela ............................... 255
Leptomenaeus ......................... 352
Leptopalaemon ....................... 314
Leslibetaeus ............................. 398
Levicaris .................................. 311
Ligur ........................................ 427
Limnocaridella ........................ 283
Limnocaridina ........................ 283
Lipkebe .................................... 352
Lipkemenes ............................. 352
Lipkius ..................................... 297
Lissocrangon .......................... 456
Lissosabinea ............................ 456
Litopenaeus ............................. 216
Lucaya ..................................... 296
Lucifer ..................................... 238
Luciferidae ............................ 238
Lysmata ................................... 427
Lysmatella ............................... 430
Macrobrachium ...................... 318
Macromaxillocarididae ..... 249
Macromaxillocaris ................. 249
Macropetasma ........................ 216
Mancicaris .............................. 284
Manipontonia ......................... 352
Margitonia .............................. 352
Marosina ................................. 284
Marsupenaeus ........................ 216
Megokris ................................. 216
Melicertus ............................... 217
Meningodora .......................... 264
Merguia ................................... 430
Merhippolyte .......................... 430
Mesocrangon .......................... 456
Mesopenaeus .......................... 234
Mesopontonia ......................... 353
Metabetaeus ............................ 398
Metacrangon .......................... 456
Metalpheus ............................. 398
Metapenaeopsis ...................... 217
Metapenaeus ........................... 222
Metapontonia ......................... 353
Micratya ................................. 284
Microprosthema ..................... 250
Mierspenaeopsis ..................... 224
Mimocaris ............................... 430
Miopontonia ........................... 353
Mirocaris ................................. 293
Miropandalus ......................... 444
Mohocaris ............................... 398
Nauticaris ............................... 430
Nautilocaris ............................ 293
Nematocarcinidae ............... 297
Nematocarcinus ..................... 297
Nematopalaemon ................... 332
Nennalpheus ........................... 398
Neoanchistus .......................... 353
Neocaridina ............................ 284
Neoclimenes ............................ 353
Neocrangon ............................ 458
Neopalaemon .......................... 332
Neopericlimenes ..................... 353
Neopontonides ........................ 353
Neosergestes ........................... 242
Neostylodactylus .................... 304
Nigmatullinus ........................ 300
Nikoides .................................. 437
Nippontonia ........................... 354
Notalpheus .............................. 399
Notocrangon ........................... 459
Notopandalus ......................... 444
Notopontonia .......................... 354
Notostomus ............................ 264
Odontonia ............................... 354
Odontozona ............................ 252
Ogyrides ................................. 435
Ogyrididae ............................ 435
Onycocaridella ....................... 354
Onycocaridites ....................... 354
Onycocaris .............................. 354
Onycomenes ........................... 355
Opaepele .................................. 293
Oplophoridae ....................... 265
Oplophorus ............................. 265
Orthopontonia ........................ 355
Orygmalpheus ........................ 399
Palaemon ................................. 332
Palaemonella ........................... 355
Palaemonetes .......................... 336
Palaemonias ............................ 286
Palaemonidae ....................... 314
Pandalidae ............................ 440
Pandalina ................................ 444
Pandalopsis ............................. 445
Pandalus ................................. 445
Pantomus ................................ 446
Parabetaeus ............................. 399
Paracaridina ........................... 286
Paraclimenaeus ...................... 356
Paraclimenes .......................... 356
Paracrangon ........................... 459
Parahepomadus ...................... 208
Paralatreutes .......................... 431
Paralebbeus ............................. 431
Paranchistus ........................... 357
Parapasiphae ........................... 256
Parapenaeopsis ....................... 224
Parapenaeus ........................... 224
Parapontocaris ....................... 460
Parapontophilus ..................... 460
Parasergestes .......................... 243
Paraspongicola ....................... 251
Parastylodactylus .................. 304
Paratya .................................... 286
Paratypton .............................. 357
Parhippolyte ........................... 413
Parisia ..................................... 287
Pasiphaea ................................ 257
Pasiphaeidae ........................ 254
Patonia .................................... 357
Peisos ....................................... 244
Pelagopenaeus ........................ 225
Penaeidae .............................. 213
Penaeopsis ............................... 226
Penaeus ................................... 226
Periclimenaeus ....................... 357
Periclimenella ......................... 360
Periclimenes ........................... 360
Periclimenoides ...................... 366
Peripandalus ........................... 447
Petalidium .............................. 244
Philarius ................................. 366
Philocheras .............................. 461
Phycocaris ............................... 432
Phycomenes ............................ 367
Phyllognathia ......................... 313
Physetocarididae ................. 453
Physetocaris ............................ 453
Pinnotheronia ......................... 368
Placopsicrangon ..................... 464
Platycaris ................................ 368
Platypontonia ......................... 368
Pleoticus .................................. 235
Plesiomenaeus ........................ 368
Plesionika ................................ 447
Plesiopenaeus ......................... 208
Plesiopontonia ........................ 368
Pliopontonia ........................... 368
Pomagnathus .......................... 399
Pontocaris ............................... 464
Pontonia .................................. 368
Pontonides .............................. 370
Pontoniopsides ....................... 370
Pontoniopsis ........................... 370
Pontophilus ............................ 465
Poripontonia ........................... 370
Potamalpheops ....................... 399
Potimirim ................................ 287
Prionalpheus ........................... 400
Prionocrangon ........................ 465
Procarididae ......................... 248
Procaris ................................... 248
Processa ................................... 437
Processidae ........................... 436
Procletes .................................. 451
Propontonia ............................ 370
Protrachypene ........................ 227
Psalidopodidae .................... 303
Psalidopus ............................... 303
Psathyrocaris .......................... 259
Pseudalpheopsis ..................... 400
Pseudaristeus ......................... 208
Pseudathanas .......................... 400
Pseudocheles ........................... 296
Pseudochelidae .................... 296
Pseudoclimenes ...................... 370
Pseudocoutierea ...................... 370
Pseudopalaemon ..................... 337
Pseudopandalus ..................... 452
Pseudopontonia ...................... 371
Pseudopontonides .................. 371
Pseudopontophilus ................. 465
Pseudoveleronia ..................... 371
Pterocaris ................................ 400
Puteonator .............................. 288
Pycneus ................................... 288
Pycnisia ................................... 288
Pycnocaris .............................. 311
Racilius ................................... 400
Rapipontonia .......................... 371
Rhopalaemon .......................... 338
Rhynchocinetes ...................... 301
Rhynchocinetidae ............... 300
Rhynocrangon ........................ 465
Richalpheus ............................ 401
Richardina .............................. 253
Rimapenaeus .......................... 227
Rimicaris ................................. 293
Rostronia ................................. 371
Rugathanas ............................. 401
Sabinea .................................... 466
Salmoneus ............................... 401
Sandimenes ............................. 371
Sandyella ................................. 372
Saron ....................................... 432
Sclerocrangon ......................... 466
Segonzackomaius ................... 300
Sergestes .................................. 244
Sergestidae ............................ 239
Sergia ....................................... 244
Shinkaicaris ............................ 294
Sicyonella ................................ 248
Sicyonia ................................... 229
Sicyoniidae ............................ 229
Sinodina .................................. 288
Solenocera ............................... 235
Solenoceridae ....................... 232
Spirontocaris .......................... 432
Spongicola ............................... 251
Spongicolidae ....................... 249
Spongicoloides ........................ 251
Spongiocaris ........................... 252
Stegopontonia ......................... 372
Stenalpheops ........................... 403
Stenopodidae ....................... 252
Stenopus ................................. 253
Stygiocaris .............................. 289
Stylodactylidae .................... 304
Stylodactyloides ..................... 304
Stylodactylus .......................... 305
Stylopandalus ......................... 452
Synalpheus ............................. 403
Syncaris .................................. 289
Syncrangon ............................ 466
Systellaspis ............................. 266
Tanypenaeus ........................... 227
Tectopontonia ......................... 372
Tenuipedium ........................... 338
Thalassocarididae ............... 452
Thalassocaris .......................... 453
Thaumastocaris ...................... 372
Thinora .................................... 433
Thor ......................................... 433
Thorella ................................... 434
Thuylamea .............................. 412
Tozeuma .................................. 434
Trachycaris ............................. 435
Trachypenaeopsis ................... 227
Trachypenaeus ........................ 227
Trachysalambria ..................... 228
Triacanthoneus ....................... 412
Tridiscias ................................. 296
Troglindicus ............................ 338
Troglocaris .............................. 289
Troglocubanus ........................ 338
Troglomexicanus .................... 339
Tuleariocaris ........................... 372
Typhlatya ................................ 290
Typhlocarididae .................. 375
Typhlocaridina ....................... 291
Typhlocaris ............................. 375
Typhlopatsa ............................. 291
Typton ..................................... 372
Typtonoides ............................. 373
Typtonychus ........................... 373
Unguicaris .............................. 374
Urocaridella ............................ 339
Urocaris .................................. 374
Veleronia .................................. 374
Veleroniopsis ........................... 374
Vercoia ..................................... 467
Vetericaris ............................... 249
Vexillipar ................................. 412
Vir ........................................... 374
Waldola ................................... 375
Xiphocarididae .................... 302
Xiphocaris ............................... 302
Xiphopenaeus ......................... 228
Yagerocaris .............................. 412
Yemenicaris ............................. 375
Zenopontonia ......................... 375

Catalogo de Decapodos

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Catalogo de Decapodos

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Carideorum Catalogus:

The Recent Species of the Dendrobranchiate, Stenopodidean,

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