A checklist of Navicula

A Checklist of Navicula
(Class Bacillariophyceae)
of the Philippines
Milagrosa R. Martinez-Goss
Professor, Institute of Biological Sciences
and Curator, Museum of Natural History
University of the Philippines Los Baos
College, Laguna 4031, Philippines

ABSTRACT

A unified account of all known diatoms in the genus Navicula, class Bacillariophyceae, of the Philippines
from 1886 to 1987 is presented. A total of 226 taxa are listed. Included herewith are the references that
originally described or cited the taxon and the ecological/habitat records for each taxon. As much as
possible, a list of exsciccata materials is given together with their repository places.
Keywords: Navicula, Bacillariophyceae, Philippines, diatoms

INTRODUCTION
This compedium presents a unified account of all
known diatoms under the genus Navicula from 1886
to 1987 or within a span of over 100 years.
Referencing was based mainly on available literature
that deals in whole or in part on Philippine materials.
In some cases Philippine exsciccata materials
(diatom slides) that were found stored in the different
diatom herbaria in the Western World were also
noted. A wealth of these specimens were discovered
at the Diatom Section of the British Museum of
Natural History (BMNH) in London, England, which
include the collections of Adam, Bess, Comber, etc.
The collections of Mann from the USS Albatross
were also observed at the Smithsonian Institution of
the U.S. Museum of Natural History in Washington,
D.C., U.S.A.
I have not had a chance to see the holotypes of
Hustedt and Skvortzows collections. But according
to Dr. Charles W. Reimer of the Academy of Natural

Science Diliman (January-June 2001) 13:1, 1-32

Sciences of Philadelphia (ANSP), there are some

collections of Skvortzow that are in Hustedts
collection, and the latter are stored in the herbarium
of the Alfred Wegener Institute for Polar and Marine
Research in Bremerhaven, Germany.
The list includes 21 that are reported as new taxa
(Table 1) and four species found only in Mexico
(Campeche Bay) and in the marine waters of
the Philippines and nowhere else according to
Mann (1925), i.e., N. approximata var.
substauroneiformis Grun., N. carinefera Grun., N.
hennedyi var. californica (Grev.) Cl. and N.
margarita Schmidt (?).
There are 226 taxa recorded here that are recognized
by Patrick and Reimer (1966), Round et al., (1990),
and VanLandingham (1975). Unidentified specific
epithets are recorded but not enumerated. The scope
and use of the list is as in the previous paper
(Martinez-Goss, 1995).

Martinez-Goss

Table 1. New taxa of Navicula reported from the Philippines

Taxon
Navicula biggemata
N. caeca
N. clavata f. corpulenta (Mann) Hust
N. delecta Mann
N. elegantoides Hust.
N. funiculata Mann
N. glabrissima Mann
N. halophila f. tenuirostris Hust.
N. hustedtii f. philippina Skv.
N. imitans Mann
N. insignita Hust.
N. luzonensis Hust.
N. mendica Mann
N. mucicoloides Hust.
N. philippina Skv.
N. philippinarum Mann
N. pulverulenta (pulvulenta) Mann
N. pugio Mann
N. retrostauros Mann*
N. spiculifera Mann*
N. transluscens Mann

Source

Reference

Marine
Marine
Marine
Marine
Laguna de Bay; Camarines Sur, Buhi Lake
Marine
Marine
Rizal, Manila
Quezon City, Balara
Marine
Laguna de Bay, Rizal, Lyon (ponds)
Camarines Sur, Buhi Lake (rare)
Marine
Rizal, Lyon (ponds)
Quezon City, Balara
Marine
Marine
Marine
Marine
Marine
Marine

Mann, 1925
Mann, 1925
Hustedt, 1964
Mann, 1925
Hustedt, 1942a
Mann, 1925
Mann, 1925
Hustedt, 1942a
Skvortzow, 1937
Mann, 1925
Hustedt, 1942a
Hustedt, 1942a
Mann, 1925
Hustedt, 1942a
Skvortzow, 1937
Mann, 1925
Mann, 1925
Mann, 1925
Mann, 1925
Mann, 1925
Marine

*Not recognized by VanLandingham

THE LIST
NAVICULA Bory, p. 562, 1824; Patrick & Reimer, p.
439, 1966.
abrupta (Greg.) Donkin
Donkin, p. 13, 2/6, 1870;
Mann, p. 94, 1925.
Bottom marine samples taken near the islands
by the United States Steamer (USS) Albatross,
Mann (1.c.).
abruptoides Hust.
Hustedt, p. 515, f. 1557, 1964;
Podzorski & Hakansson, p. 79, 31/8, 1987.
LUZON: Palawan, Pto. Princesa (White
Beach mangrove, root scrapings from
Rhizophora), Podzorski & Hakansson (l.c.).
americana Ehr.
Ehrenberg, p. 129 (417), 1841 (1843);
Hustedt, p. 280, f. 464, 1930;
_______, p. 64, 1942a.

LUZON: Laguna, Los Baos (ponds, Laguna

de Bay); Camarines Sur, Buhi (river), Hustedt
(1942a).
MINDANAO: Lanao (Uyaan Lake), Hustedt
(1942a).
anglica Ralfs
Ralfs in Pritchard, p. 900, 1861;
Hustedt, p. 303, f. 530-531, 1930;
_______, p. 73, 1942a.
MINDANAO: Lanao (ponds), Hustedt
(1942a).
angusta Grun.
Grunow, 10:528, 5/19, 1860.
= Navicula cari var. angusta Grun.
Grunow in Van Heurck, 7/17, 1885;
Hustedt, p. 72, 1942a;
Patrick & Reimer, p. 514-515, 49/5, 1966.
LUZON: Laguna, Los Baos (ponds),
Hustedt, 1942a.

A checklist of Navicula

VISAYAS: Leyte, Danao Lake, Hustedt,

1942a.
approximata Grev.
Greville, p. 28, 4/4, 1859;
Mann, p. 94, 1925.
= N. lyra var. approximata (Grev.) Cl.
Cleve, p. 4 (1/1) 1878.
Bottom marine samples taken near the
islands by the USS Albatross, Mann (1.c.);
British Museum (BMNH), Adams slide
#45.1/2; TS 60/64.
_______ f. morei (moreii) (O Meara) Hust.
Hustedt p. 416, f. 1491, 1964.
= Navicula hennedyi var. niceaensis H. Perag.
Peragallo, p. 47, 5/39 e.p., 1888.
BMNH, Comber slide #31898.
VISAYAS: Cebu.
_______ var. substauroneiformis (Grun.) Cl.
Grunow in Schmidt, et al., 2/20, 21, 1874;
Mann, p. 94, 1925.
Bottom marine samples, Mann (l.c.).

LUZON: Manila (Reservoir); Laguna, Los

Baos; Camarines Sur, Buhi (lake), Hustedt
(1942a).
baileyana Grun.
Grunow in Schmidt, et al, p. 88, 1/31, 1874;
Castracane, p. 812, 1886.
VISAYAS: Cebu, Castracane (l.c.)
= Navicula polysticta Grev.
Greville, p. 247,5/2, 1859.
VISAYAS: Cebu, BMNH, Comber
# 31892.
barbitos Schmidt
Schmidt, et al., 129/5, 1874;
Mann, p. 95, 1925.
Bottom marine samples taken around the
islands by the USS Albatross, Mann (l.c.).
biformis (Grun.) Mann
Mann, p. 95, 20/6-7, 1925.
Bottom marine samples taken around the
islands by the USS Albatross, Mann (1.c.).
= Mastoneis biformis (Grun.) Cl.
Cleve, p. 194, 1894.

_______ var. ?
BMNH, Adams-Bess # TS497 3/2.

arvensis Hust.
Hustedt, p. 249, 20/19, 20, 1937b;
_______, p. 60, 1942a;
Patrick & Reimer, p. 483, 46/1-2, 1966.
LUZON: Batangas, Taal (Crater Lake),
Hustedt (1942a).
bacillum Ehr.
Ehrenberg, p. 130, 1838, (1839);
_________, p. 418 (130), 4/5, f. 8, 1841 (1843);
Hustedt, p. 64, 1942a.

LUZON: Manila, BMNH, Comber

# 31904.
VISAYAS: Cebu, BMNH, Comber
# 31895.
bigemmata Mann
Mann, pp. 95-96, 21/1, 1925.
Bottom marine samples taken around the
islands by the USS Albatross. Type-Cat. No. 43639
deposited in the United States National Museum
of Natural History (USNM), Mann (1.c.).

Martinez-Goss

bipunctata O Meara
O Meara, p. 286, 13/5, 1872;
De Toni, pp. 78-79, 1891.
MINDANAO: Brackishwaters coming from
the volcanic mountain of the island in Cagayan, Sulu
Archipelago (Chimmo), O Meara (l.c.); De Toni,
(l.c.).
borneoensis Hust.
Hustedt, p. 735, f. 1712, 1966;
Podzorski & Hakansson, p. 84, 33/7, 1987.
LUZON: Palawan, Taytay, Langen, Malapakan
Is. (algae encrusted on cove, marine), Podzorski
& Hakansson (1.c.).
brasiliensis Grun.*
Grunow, p. 152, 14/10, 1863;
Schmidt, Atlas, 6/19-21, 23/25, 31/33, 1874;
Mann, p. 97, 1925.
Bottom marine samples taken around the
islands by the USS Albatross, Mann (1.c.).
BMNH, Adams # F19.29, 40 2/18.883; TS
262 1/10-12, 6/7, TS 60/60.
VISAYAS: Cebu, BMNH, Combers #31898,
31895, 31894.
brasiliensis var. _______Cstr.
Castracane, p. 812, 1886.
BMNH, Adams-Bess #706, TS 60/66.

bruchii Grun.
Grunow in Cleve, p. 13, 1928;
Cleve, p. 36, 1895;
De Toni, pp. 177-178, 1891.
LUZON: Manila (marine), (Deby collection,
BMNH).
bryophila Peters.
Petersen, p. 388, f. 13, 1928;
Hustedt, p. 60, 1942a.
MINDANAO: Lanao Lakes of Dagiangan and
Uyaan, Hustedt (1.c.).
caeca Mann
Mann, p. 97, 21/3, 1925.
Bottom marine samples taken near the islands
by the USS Albatross. Type-Cat. No. 43641,
USNM, Mann (1.c.).
capitata Ehr.Ehrenberg, p. 185, 13/20, 1838;
Van Heurck, p. 187, 25/719, 1896;
Patrick & Reimer, pp. 536-537, 52/1-2, 1966;
Podzorski & Hakansson, p. 81, 32/10, 1987.
LUZON: Palawan, Taytay, Langen
(Malapakan Cove, algae encrusted on the cove,
marine), Podzorski & Hakansson (1.c.).
cari Ehr.
Ehrenberg, p. 83, 1836;
_________, p. 174, 1838;
Hustedt, p. 299, fig. 512, 1930;
_______, p. 72, 1942a.

VISAYAS: Cebu, Castracane (l. c.)

brekkaensis Peters.
Petersen, p. 389, f. 16, 1928;
Hustedt, p. 58, 1942a.
LUZON: Rizal, Pancipit River, Hustedt (l.c.).

Not recognized by VanLandingham, 1975.

LUZON: Rizal, Malabon (Dagatan swamp),

Hustedt (1942a).
MINDANAO: Uyaan Lake, Hustedt (1942a).
carinifera Grun.
Grunow in Schmidt, et al., 2/1, 1874;
Mann, p. 98, 1925;
Hustedt, p. 754, f. 1730, 1966.

A checklist of Navicula

Bottom marine samples taken near the islands

by the USS Albatross, Mann (1.c.).
cincta (Ehr.) Ralfs
Van Heurck, p. 178, 3/105a, 1896;
Schmidt, et al., 299/26-30, 1874;
Hustedt, p. 298, f. 510, 1930;
Pantastico, p. 194, 17/4, 1977;
Patrick & Reimer, p. 516, 49/8, 1966.
LUZON: Laguna, Los Baos, Mayondon (on
shoreline of Laguna de Bay), Pantastico (1.c.).
circumsecta (Grun.) Grun.
Grunow in Schmidt, et al., 3/26-27, 1874;
Cleve et Grunow, p. 42, 1880;
Mann, p. 98, 1925;
Hustedt, p. 409, f. 1487a-c, 1964.
Bottom marine samples taken near the islands
by the USS Albatross, Mann (1.c.).
= Navicula polysticta var. circumsecta (Grun.)
Grun.
Grunow in Schmidt, et al., p. 89, 1/36, 42, 1874.

LUZON: Manila (on shell), BMNH, Comber

# 30925, 31903.
VISAYAS: Cebu, BMNH, Comber # 31900,
31898.
= Navicula clavata var. caribaea Cl.
Cleve, p. 61, 1895.
VISAYAS: Cebu, BMNH, Comber
# 31895.
= Navicula caribaea Cl.
Cleve in Schmidt, et al., 2/17, 70/48, 1874;
Mann, pp. 99-100, 1925.
Bottom marine samples taken near the
islands by the USS Albatross, Mann (1.c.).
= Navicula wrightii O Meara
LUZON: Manila, BMNH, Comber
# B. 31903.
_______ f. corpulenta (Mann) Hust.
Hustedt, p. 445, f. 1510-1511, 1964.

BMNH, Adams # GC 2089.

circumtexta Meister
Meister in Schmidt, et al., 394/33-35, 1874;
Podzorski & Hakansson, pp. 81-82, 32/4, 1987.
LUZON: Palawan, Taytay, Langen
(Malapakan Cove, scrapings from sand at low tide,
marine), Podzorski & Hakansson (1.c.).
citrus Krasske
Krasske, p. 199, f. 14, 1923;
Hustedt, p. 303, f. 529, 1930;
_______, p. 66, 1942a.

= N. corpulenta Mann
Mann, p. 99, 22/4, 1925.
Bottom marine samples taken around the
islands by the USS Albatross. Type-Cat. No.
43642, USNM, Mann (1.c.).
_______ f. elliptica (Schmidt)Hust.
Schmidt, et al., 3/13, 1874;
Hustedt, p. 449, 1964.
LUZON: Manila, BMNH, Comber # 31903.
VISAYAS: Cebu, BMNH, Comber # 31898.

LUZON: Laguna de Bay, Hustedt (1942a).

clavata Greg.
Gregory, p. 46, 5/17, 1856.
BMNH, Adams # F83 1/2; 4/3; 4. 120, 3/2.5.

_______ var. indica (Grev.) Cl.

Cleve, p. 62, 1895;
Schmidt, et al., 204/12, 1874.
BMNH, Adams # GC 1865.6.1976; Bess
#920; TS #497.2/3.4.522.2/4, F885.

Martinez-Goss

= N. indica Grev. (non Ehr.)

Greville, p. 95, 9/13, 1862;
Cleve, p. 62, 1895;
Mann, p. 104, 1925.
Bottom marine samples taken near the
islands by the USS Albatross, Mann (1.c.).
BMNH, Adams # J 4201, M 68/4.
LUZON: Manila (marine), BMNH, Raes
collection.
VISAYAS: Cebu, BMNH, Comber
# 31898, Raes collection.
_______ var. producta ? sensu Adam
BMNH, Adams # F886.
LUZON: Manila, BMNH, Comber # 31904.

confervacea (Ktz.) Grun.

Grunow in Van Heurck, 14/36, 1880;
Woltereck, p. 37, 1941;
Hustedt, p. 63, 1942a;
Prowse, p. 41, 12/n-o, 1962;
Hustedt, p. 205, f. 1324a-d, 1962;
Podzorski & Hakansson, pp. 77-78, 1987.
LUZON: Benguet, Banaue (ponds), Woltereck,
(1942); Rizal, Malabon, Dagatan (swamp), Marulay
Water Reservoir, Lyon ponds; Laguna, Los Baos,
Laguna de Bay; Batangas, Taal (Taal Lake, YamboCrater Lake, Crater Lake); Camarines Sur, Buhi
(lake, river); Oriental Mindoro, Naujan (river),
Calapan Lake, Hustedt (1942a); Palawan, Roxas
(Caruray River, alt., c. 50 m, from rock surface),
Taytay, Lake Manguao, (algae growing on floating
grass roots, alt. c. 45 m), Podzorski & Hakansson,
(1987).
VISAYAS: Leyte, Danao Lake, Hustedt
(1942a).

_______ var. ?
MINDANAO: Lanao Lake, Hustedt (1942).
BMNH, Adams # TS60/61.657.
clipeiformis Knig
Knig, p. 51, 59, 2/3, 3/6, 1959;
Hustedt, p. 552, f. 1589, 1964;
Podzorski & Hakansson, p. 76, 31/9, 1987.
LUZON: Palawan, Taytay, Langen
(Malapakan Cove, surface scrapings from sand at
low tide, marine), Podzorski & Hakansson (1.c.).
cluthensis Greg.
Gregory, p. 478, 9/2, 1857;
Hustedt, p. 651, f. 1653a-d, 1966.
= N. erythraea Grun., p. 539, 5/17 (3/17), 1860;
Mann, p. 101, 1925.
Bottom marine samples taken near the
islands by the USS Albatross, Mann (1.c.).

= N. confervacea f. nipponica Skv.

Skvortzow, p. 34, 2/3, 4/23, 1936;
Hustedt, p. 61, f. 23, 1942a
LUZON: Camarines Sur, Buhi Lake,
Hustedt (l.c.).
consors Sch.
Schmidt, et al., 48/24-27, 1874;
Mann, p. 98, 1925.
Bottom marine samples taken near the islands
by the USS Albatross, Mann (1.c.).
contenta Grun.
Grunow in Van Heurck p. 109, 1885;
Van Heurck, p. 230, 5/239, 1896;
Hustedt, p. 277, f. 458a, 1930;
Skvortzow, p. 291, 1/40, 1937;
Hustedt, p. 58, 1942a.

A checklist of Navicula

LUZON: Batangas, Taal (Crater Lake),

Hustedt (1942a).
______ var. biceps (Arn. Ms. Grun. in V.H.) Cleve
Van Huerck, p. 230, 5/240, 1896.
= N. contenta f. biceps (Arn. Grun in V.H.) Hust.
Hustedt, p. 277, f. 458c, 1930;
_______, p. 58, 1942a.

cuspidata (Ktz.) Ktz.

Ktzing, p. 94, 3/24, 37, 1844;
W. Smith, p. 47, 16/131, 1853;
Van Heurck, p. 214, 4/190, 1896;
Mann, p. 99, 1925;
Hustedt, p. 268, f. 433, 1930.
Freshwater sample as detritus from the bottom
marine samples taken by the USS Albatross, Mann
(1.c.).

LUZON: Pancipit River, Hustedt (1942a).

_______ f. parallela (Peters.) Hust.
Hustedt, p. 277, f. 458b, 1930;
_______, p. 58, 1942a.
LUZON: Camarines Sur, Buhi (lake), Hustedt
(1942a).
cryptocephala Ktz.
Ktzing, p. 95, 3/20, 26, 1844;
Donkin, p. 37, 5/14, 1871;
Van Heurck, p. 180, 3/122, 1896;
Hustedt, p. 295, f. 496, 1930;
Skvortzow, p. 291, 2/3, 1937;
Hustedt, p. 67, 1942a;
Podzorski & Hakansson, p. 87, 1987.
LUZON: Rizal, Manila (Water Reservoir II),
ponds in the Bureau of Science grounds; Laguna,
Laguna de Bay; Camarines Sur, Buhi Lake; Oriental
Mindoro, Naujan Lake, Hustedt (1942a); Rizal,
Quezon City (Balara filters, Skvortzow, (l.c.);
Palawan, Brookes Point (scrapings from a
limestone rock in Tumarabong River, alt. c. 10 m),
Podzorski & Hakansson (1987).
cf. cryptolyra Brockmann
Brockmann, p. 19, 3/22, 23, 1950;
Hustedt, p. 534, f. 1570, 1964;
Podzorski & Hakansson, p. 79, 32/5, 1987.
LUZON: Palawan, Taytay, Langen, Malapakan
Is. (algae encrusted on the cove, marine), Podzorski
& Hakansson (1.c.).

_______ var. ambigua (Ehr.) Cl.

Cleve, p. 110, 1894;
Hustedt, p. 51, 1942a.
LUZON: Rizal (ponds in the Bureau of Science
grounds); Laguna, Los Baos, Laguna de Bay (lake
and ponds); Camarines Sur, Buhi Lake, Hustedt
(1942a); Palawan, Roxas, Pagdanan Range,
Ibangley Brookside Hill (streamside pool, alt. c. 40
m), Podzorski & Hakansson (1.c.).
MINDANAO: Cagayan de Oro, Singnan Lake;
Lanao (lake), Hustedt (1.c.).
cyclops Mann
Mann, pp. 99-100, 21/5, 1925.
Bottom marine samples taken near the islands
by the USS Albatross; frequently observed. TypeCat. No. 43643, USNM, Mann (1.c.).
delecta Mann
Mann, p. 100, 24/5-7, 1925.
Bottom marine samples taken around the
islands by the USS Albatross; frequently observed.
Type-Cat. No. 43644, USNM, Mann (1.c.).
BMNH, Adams # M58/31.
dicephala Ehr.
Ehrenberg, p. 45, 1837;
_________, p. 185, 1838;
Van Heurck, p. 188, 3/138, 1896;
Hustedt, p. 302, f. 526, 1930;
_______, p. 73, 1942a.

Martinez-Goss

LUZON: Laguna, Los Baos (freshwater

ponds), Hustedt (1942a).
_______ var. undulata str.
strup, p. 25, 3/33, 1918 (1920);
Hustedt, p. 73, 1942a.

_______ var. rhomboidea (Cstr.) Schmidt

Schmidt, et al., 129/2, 1874.
= Navicula bullata Norman var. rhomboides Cstr.
Castracane, p. 30, 30/8, 1886.
VISAYAS: Cebu, Castracane (l.c.).

LUZON: Camarines Sur, Buhi Lake, Hustedt

(1942a).
diffluens Schmidt
= Navicula diffusa Schmidt
Schmidt, et al., 2/15, 2/28, 1874;
Mann, p. 101, 1925.
Bottom marine samples taken near the
islands by the USS Albatross, Mann (l.c.).
directa (W. Sm.) Ralfs
Ralfs in Pritchard, p. 906, 1861.
LUZON: Manila, BMNH, Comber # 31903,
31904.
distans (W. Sm.) Ralfs
Ralfs in Pritchard, p. 907, 1861;
Schmidt, et al., 46/11-14, 1874.
LUZON: Manila (on shell), BMNH, Comber
# 30925, 30926.
durandii (durrandii) Kitt.
Kitton in Schmidt, et al., 129/1, 1874;
Castracane, p. 812, 1886;
Mann, p. 101, 21/6, 1925;
Hustedt, p. 485, f. 1540, 1964.
Bottom marine samples taken near the islands
by the USS Albatross, Mann (1.c.);
BMNH, Adams # TS522.1/1.
VanLandingham (1975) does not think the specimen
is in accordance to Kitton.
VISAYAS: Cebu, Castracane (l. c.)

_______ var ?
BMNH, Adams # M 65/5.
elegantoides Hust.
Hustedt, p. 76, f. 142, 1942a;
Woltereck, p. 52, 1941;
Prowse, p. 42, 8/d, 1962.
LUZON: Benguet, Banaue (ponds), Woltereck
(l.c.); Rizal, Manila (Lyon ponds, ponds in Bureau
of Science grounds); Laguna, Los Baos, Laguna
de Bay; Buhi Lake, Hustedt (1.c.).
exigua (Greg.) Grun.
Grunow in Van Heurck, 8/32, 1880;
Hustedt, p. 303, f. 538, 1930;
_______, p. 73, 1942a.
LUZON: Camarines Sur, Buhi Lake, Hustedt
(1942a).
MINDANAO: Uyaan Lake, Hustedt (1942a).
forcipata Grev.
Greville, p. 83, 6/10-11, 1859;
Schmidt, et al., 70/17, 1874;
Cleve, p. 65, 1895;
Van Heurck, p. 203, 4/163, 1896;
Peragallo & Peragallo, p. 130, 21/28, 1897-1908.
BMNH, Adams # J 413.
VISAYAS: Cebu, BMNH, Comber # 31898.
_______ var. densestriata Schmidt
Schmidt, et al., 70/14-16, 1874;
Mann, p. 102, 1925.

A checklist of Navicula

Bottom marine samples taken near the islands

by the USS Albatross, Mann (1.c.).

LUZON: Rizal, Manila (Tabaco River), sample

taken by the ship Princess Louise, Ehrenberg
(1854).

_______ cf. forcipata var. elongata Perag & Perag.

Peragallo & Peragallo, p. 131, 21/24, 1897-1908; granulata Bailey
Bailey, p. 10, f. 16, 1854 (1853);
Podzorski & Hakansson, p. 79, 33/1, 1987.
Cleve, p. 48, 1895;
Schmidt, et al., 244/7, 1874;
LUZON: Palawan, Pto. Princesa, Pto.
Hustedt, p. 702, f. 1696, 1966;
Princesa Bay (Table Head, on sandy bottom at 15
Podzorski & Hakansson, p. 84, 33/14, 1987.
m depth), Podzorski & Hakansson (1.c.).
_______ var. versicolor (Grun.) Grun.
Grunow in Cleve et Mller, no. 208-210, 1879.

LUZON: Palawan, Taytay, Langen, Malapakan

Is. (surface scrapings from the sand at low tide, by
the cove), Podzorski & Hakansson (1.c.).

VISAYAS: Cebu, BMNH, Comber # 31894,

cf. gregaria Donk.
31902.
Donkin, p. 10, 1/10, 1861;
Van Heurck, p. 181, 3/125, 1896;
_______ var ?
Hustedt, p. 269, f. 437, 1930;
Podzorski & Hakansson, p. 78, 1987.
BMNH, Adams # G-211.
funiculata Mann
Mann, p. 102, 22/1-2, 1925.
Bottom marine samples taken around the
islands by the USS Albatross. Type-Cat. No. 43645,
USNM, Mann (1.c.).
BMNH, Adams # TS 60/2.76.
genifera Schmidt
Schmidt, et al., 2/6, 1874;
Hustedt, p. 435, f. 1505, 1964.
BMNH, Adams # W 935.
glabrissima Mann
Mann, p. 103, 22/3, 1925.
MINDANAO: Jolo, Sulu, Sulu (bottom marine
sample, rare). Type-Cat. No. 43646, USNM, Mann
(1.c.).
gracilis Ehr.
Ehrenberg, p. 64, 69, 1830 (1832);
_________, p. 155, 1854;
Van Heurck, p. 179, 3/109, 1896;
Hustedt, p. 299, f. 514, 1930.

=Navicula granulata cf. gregaria Donk.

Donkin, p. 10, 1/10, 1861;
Van Heurck, p. 181, 3/125, 1896;
Hustedt, p. 269, f. 437, 1930;
Podzorski & Hakansson, p. 78, 1987.
LUZON: Palawan, Taytay, Langen,
Malapakan Is. (algae encrusted on the cove,
marine), Podzorski & Hakansson (1.c.).

aff. grevillei (Ag.) Heiberg

Heiberg, p. 83, 1863;
Van Heurck, p. 232, 5/243, 1896;
Prowse, p. 43, 12/6, 1962;
Podzorski & Hakansson, p. 77, 30/9, 1987.
LUZON: Palawan, Taytay, Langen, Malapakan
Is. (algae encrusted on the cove, marine), Podzorski
& Hakansson (1987).
grimmii (grimmei) Krasske
Krasske, p. 45, 1/14, 1925;
Woltereck, p. 52, 1941;
Hustedt, p. 274, f. 448, 1930;
Hustedt, p. 55, 1942a.

LUZON: Benguet, Banaue (ponds), Woltereck

(l.c.); Rizal, Lyon (ponds); Camarines Sur, Buhi
(river), Hustedt (1942a).
MINDANAO: Lanao (lake), Hustedt (1942a).
halophila (Grun.) Cl.
Cleve, p. 109, 1894;
Hustedt, p. 268, f. 436, 1930;
_______, p. 51, 1942a;
Woltereck, p. 52, 1941.

Van Heurck, p. 93, 9/14, 1880-1885;

Cleve, p. 57, 1895;
Mann, p. 104, 1925.
Bottom sediments taken near the islands by
the USS Albatross, Mann (1.c.); BMNH, Adams
# E 530, M 58/14, F119.1/14.3/3.
LUZON: Manila, BMNH, Comber #30925,
31903, 31904, 30926 (on shell).
VISAYAS: Cebu, BMNH, Comber #31900.

LUZON: Mt. Province, Banaue (ponds)

Woltereck (l.c.); Rizal, Lyon (ponds); Laguna, Los
Baos (pond), Hustedt (1942a).
MINDANAO: Lanao, Uyaan (lakes), Hustedt
(1942a).
_______ f. tenuirostris Hust.
Hustedt, pp. 51-52, f. 76, 1942a.
LUZON: Rizal, Manila (ponds in the Bureau
of Science grounds, rare), Hustedt (1.c.).
hamulifera Grun.
Grunow in Cleve and Grunow, p. 44, 1880;
Cleve, vol. 1, p. 154, 3/16-19, 1894;
Peragallo & Peragallo, p. 65, 8/16, 1897-1908;
Mann, p. 104, 1925;
Hustedt, p. 312, f. 1430, 1962.
Bottom sediments taken near the islands by
the USS Albatross, Mann (1.c.).
hassiaca Krasske
Krasske, p. 47, 2/26, 1925;
Hustedt, p. 279, f. 462, 1930;
_______, p. 216, f. 1332, 1962;
Podzorski & Hakansson, p. 81, 32/2, 1987.
LUZON: Palawan, Taytay, Lake Manguao
(Danao), Podzorski & Hakansson (l.c.).
hennedyi (henedii) W. Sm.
W. Smith, p. 93, 1856;
Gregory, p. 40, 5/3, 1856;

10

=Navicula hennedyi var. niceaensis

(niceaeensis) H. Perag.
H. Peragallo, p. 55, 5/39,1888.
BMNH, Adams # M 58/22-3.
______ f. bacillifera (Pantoc.) Cl.
Cleve, p. 58, 1895.
= Navicula rugosa Janisch
Janisch, 15/10, 11, 1888;
Cleve, p. 59, 1895.
BMNH, Adams # E 530.
_______ var. californica (Grev.) Cl.
Greville, p. 29, 4/5, 1859;
Cleve, p. 58, 1895;
Mann, p. 97, 1925.
= Navicula californica Grev.
Greville, p. 29, 4/5, 1859;
Mann, p. 97, 1925.
Bottom marine samples taken near the
islands by the USS Albatross, Mann (l.c.);
BMNH, Adams # M58/20-1; 24.30.
=Navicula californica var. campechiana Grun.
Grunow in Schmidt, et al., 3/19, 1874;
Mann, p. 97, 1925.
Bottom marine samples taken near the
islands by the USS Albatross, Mann (1.c.).

_______ var. furcata Perag. & Perag.

Peragallo & Peragallo, p. 141, 24/16, 17, 1897-1908.
BMNH, Adams # F 18 2/9.10.
_______ f. granulata (Grun.) Hust.
Grunow in Schmidt, et al., 3/3, 1874;
Hustedt, p. 458, f. 1519, 1964.
=Navicula hennedyi var. circumsecta Cl.
Cleve p. 58, 1895 (non Grunow, 1874 in
Schmidt, et al., 1874).
It means Cleve thought the specimen
is not the same as in Grunow in Schmidt, et al.,
but VanLandingham says they are the same.
BMNH, Adams # TS 519/3.
_______ var. neopolitana Cl.
Cleve, p. 58, 1895;
Podzorski & Hakansson, p. 80, 31/12, 1987.
LUZON: Palawan, Taytay (beach on washedashore seaweed), Podzorski & Hakansson (1.c.).
_______ f. ?
BMNH, Adams # F 120.3/7.
hennedyi Hust.*
Podzorski & Hakansson, p. 79, 30/1, 1987.
LUZON: Palawan, Pto. Princesa, Table Head,
South of Pto. Princesa Bay (from surface at 15 m
depth), Podzorski & Hakansson (1.c.).
howeana Hagelstein
Hagelstein, p. 385, 7/51, 1939;
Podzorski & Hakansson, pp. 76-77, 30/6, 1987.
LUZON: Palawan, Taytay, Langen, Malapakan
Is. (surface scrapings from the sand at low tide in
Malapakan Cove), Podzorski & Hakansson (1.c.).
hustedtii Krasske
Krasske, 3:198, f. 3, 1923;
Patrick & Reimer, p. 527, 50/8, 1966.

= Navicula hustedtii f. philippina Skv.

Skvortzow, p. 290, 1/44, 1937a.
LUZON: Rizal, Quezon City, Balara (water
filter no. 8), Skvortzow (l.c.).
imitans Mann
Mann, p. 104, 22/5, 6, 1925.
Bottom marine samples taken near the islands
by the USS Albatross. Type-Cat. No. 43643,
USNM, Mann (1.c.).
indicatrix VanLan.
VanLandingham, nom. nov.
= N. alpha Chol.
Cholnoky, p. 173, fig. 57, 1955 (non Cl. 1893).
LUZON: Manila, BMNH, Comber
#31903.
inexacta Mann
Mann, p. 105, 1925.
= N. (Alloioneis) gruendleri (Cl. et Grun.) Cl.
Mann, p. 104, 105, 22/7, 1925.
Bottom marine samples taken near the
islands by USS Albatross. No type designated,
Mann (1.c.).
inhalata Schmidt
Schmidt, et al., 2/30, 1874;
Cleve, p. 57, 1895;
Mann, p. 107, 23/2, 1925;
Hustedt, p. 390, f. 1475, 1964.
Bottom marine samples taken near the islands
by USS Albatross, Mann (1.c.); marine, Cleve
(l.c.).
insignita Hust.
Hustedt, p. 73, f. 126, 139-141, 1942a;
Woltereck, p. 52, 1941;
Patrick & Reimer, p. 447, 40/3, 1966.

11

Martinez-Goss

LUZON: Mt. Province, Bontoc, (Benguet,

Banaue, ponds); Lyon (ponds), Woltereck (1.c.),
Hustedt (1.c.); Rizal; Laguna, Laguna de Bay,
Hustedt (1.c.).
insociabilis Krasske
Krasske, p. 114, 3/17, 1932;
Schmidt, et al., 40/103-105, 1874;
Hustedt, p. 63, 1942a;
_______, p. 181, f. 1315a-h, 1962.
LUZON: Rizal, Pancipit River, Hustedt
(1942a).
iota (jota) Cl.
Cleve, p. 134, 5/22, 1894.

LUZON: Camarines Sur, Lake Buhi,

Hustedt (l.c.).
lagerheimii Cl.
Cleve, p. 131, 1894;
_____, p. 101, 7/1, 1894;
Woltereck, 41:37-176,1941;
Hustedt, in A. S. Atlas, T. 370, f. 19-21, 1874;
Hustedt, p. 54, 1942a.
LUZON: Mt. Province, Bontoc (Banaue,
ponds), Woltereck (l.c.); Laguna de Bay, Los
Baos (ponds), Pancipit River; Camarines Sur, Lake
Buhi, river, Hustedt (1942a).
MINDANAO: Lanao, Lanao Lake, Uyaan
Lake, Hustedt (1942a).

VISAYAS: Cebu, BMNH, Comber # 31898.

irrorata var. substauroneiformis (Grun.) Cl.
Cleve, p. 56, 1895
=Navicula approximata var substauroneiformis
(Grun.) Cl.
Grunow in Schmidt, et al., 2/20, 21, 1874;
Mann, p. 94, 1925.
Bottom marine samples taken near the
islands by the USS Albatross, Mann (l.c.).
jejuna Schmidt
Schmidt, (1876) in Schmidt, et al., 46/76, 1874;
Mann, p. 107, 1925;
Skvortzow, p. 280, 4/7, 1932.
Bottom marine samples taken near the islands
by the USS Albatross, Mann (1.c.).
laevissima Ktz.
Ktzing, p. 96, 21/14, 1844;
Patrick & Reimer, p. 497, 47/13, 1966.
BMNH, Adams # GC 1994.
= Navicula bacilliformis Grun.
Grunow in Cleve & Grunow, p. 44, 2/51, 1880;
Hustedt, p. 55, 1942a.

12

_______ f. intermedia Hust.

Hustedt, in A. S. Atlas, T. 370, f. 22, 1874;
LUZON: Oriental Mindoro, Calapan Lake,
Hustedt (l.c.).
lanceolata (Ag.) Ktz.
Ktzing, p. 94, 28/38, 30/48, 1844;
Van Heurck, p. 186, 3/139, 1896;
Hustedt, p. 305, f. 540, 1930;
_______, p. 72, 1942a.
MINDANAO: Lanao, Uyaan Lake, Hustedt
(1942a).
latissima Greg.
Gregory, p. 40, 5/4, 1856.
LUZON: Manila, BMNH, Comber #31904.
longa (Greg.) Ralfs
Ralfs in Pritchard, p. 906, 1861;
Donkin, p. 55, 8/3a-b, 1872;
Van Heurck, p. 185, 25/716, 1896;
Mann, p. 108, 1925.
Bottom marine samples taken near the islands
by the USS Albatross, Mann (1.c.).

A checklist of Navicula

lorenzii (Grun.) Hust.

Hustedt, p. 29, f. 1188, 1961;
Podzorski & Hakansson, p. 78, 3/1, 1a, 2, 2a, 1987.
= Stichodesmis australis Grev.
Greville, p. 535, 13/1-4, 1863.
Bottom marine samples taken near the
islands by the USS Albatross, Mann (l.c.).
LUZON: Palawan, Pto. Princesa, Table
Head (from sand surface at 15 m deep of Pto.
Princesa Bay), Podzorski & Hakansson (1.c.).
= Navicula scopulorum sensu Cl.
Cleve, p. 151, e.p., 1894.
VISAYAS: Cebu, BMNH, Comber
# 31892.
lunula Cl.
Cleve, 5/5,1894;
______, p. 51, 1895.
BMNH, Adams # G 211.
luzonensis Hust.
Hustedt, p. 59, f. 106, 1942a.
LUZON: Camarines Sur, Buhi Lake, Hustedt
(1.c.).
lyra Ehr.
Ehrenberg, p. 419, 1/1, f. 9a, 1841 (1843);
Bailey, p. 431, 1853;
Schmidt, et al., 2/16, 129/11-14, 1874;
Cleve, p. 63, 1895;
Mann, p. 108, 1925;
Podzorski & Hakansson, p. 80, 31/10, 1987.
Bottom marine samples taken near the islands
by the USS Albatross, Mann (1.c.).
BMNH, Adams-Bess #715, F 121, 2/7, M 58/
15.16, TS 497.2/2.

LUZON: Manila, BMNH, Comber #31904;

Palawan, Taytay, Langen, Malapakan Is. (algae
encrusted in Malapakan Cove, marine), Podzorski
& Hakansson (1.c.).
= Navicula gregoryana Grev.
Greville, p. 10, 3/7, 1857.
LUZON: Manila, (on sponge), BMNH,
Comber #33424.
VISAYAS: Cebu, BMNH, Comber
# 31894.
_______ var. cuneata Temp. & Perag.
Tempere & Peragallo, p. 473, No. 988, 1914.
VISAYAS: Cebu, BMNH, Comber # 31894,
31898.
_______ var. elliptica Schmidt
Schmidt in Schmidt, et al., 2/29, 2/34, 1874;
Cleve, p. 64, 1895;
Skvortzow, p. 272, 3/9, 1931;
Mann, p. 108, 1925.
Bottom marine samples taken near the islands
by the USS Albatross, Mann (1.c.); marine, Cleve
(l.c.).
BMNH, Adams #F 60.1/5.
VISAYAS: Cebu, BMNH, Comber #31898.
= Navicula lyra var. subelliptica Cl.
Cleve, p. 64, 1895.
Marine, Debys collection, Cleve (l.c.).
_______ var. elliptica f. bullata (Norman) Cl.
Cleve, p. 63, 1895;
Schmidt, et al., 3/8-9, 1874.
= N. bullata Norman
Norman, p. 8, 2/7, 1861;
Mann, p. 97, 1925.

13

Martinez-Goss

Bottom marine samples taken near the

islands by the USS Albatross, Mann (1.c.).
BMNH, Adams # F 40, 2/12.17.
VISAYAS: Cebu, BMNH, Comber
# 31894, 31897, 31898 (2), 31900.
_______ var. granulata Perag. & Perag.
Peragallo & Peragallo, 23/5, 1897;
Schmidt, et al., 258/1,1874___.
BMNH, Adams # F 45 1/10.
_______ var. indica, sensu Adam
BMNH, Adams # W 405.
_______ var. insignis Schmidt
Schmidt in Schmidt, et al., 2/27, 257/4, 1874;
Mann, p. 108, 1925.
Bottom marine samples taken near the islands
by the USS Albatross, Mann (1.c.).
= Navicula lyra var.? intermedia Perag. & Perag.
Peragallo & Peragallo, p. 136 (141), 23/6 (11),
24/18, 1897.

BMNH, Adams # F 40 2/15.45.1/9.60

1/8. TS 497 2/10.
_______ var. seductilis (Schmidt) OMeara
= Navicula seductilis Schmidt
Schmidt in Schmidt, et al., 2/35, 1874;
Mann, p. 119, 1925.
Bottom marine samples taken near the
islands by the USS Albatross, Mann (1.c.).
_______ var. signata Schmidt
Schmidt in Schmidt, et al., 2/4, 1874;
Castracane, p. 33, 30/13, 1886.
Bottom marine samples taken near the islands
by the USS Albatross, Mann (1.c.). Frustules found
in the vicinity of the Philippines, Castracane (l.c.).
VISAYAS: Cebu, BMNH, Comber # 31893,
31895.
_______ var. subtropica Schmidt
Schmidt in Schmidt, et al., 2/24.25, 1874;
Wolle, 16/14, 1890.
BMNH, Adams # F 40 2/15.45.1/9.60 1/8. TS
497 2/10.

BMNH, Adams # F 20.2/8.

_______ var. ?
_______ var. recta Grev.
Greville, p. 28, 4/3, 1859;
Mann, p. 108, 1925.
Bottom marine samples taken near the islands
by the USS Albatross, Mann (1.c.).
BMNH, Adams # TS 497-2/5.
= N. lyra var. recta f. typica Perag.
Peragallo, 22/8, 1891.
BMNH, Adams # TS 522 1/4.5.
= N. lyra var. recta f. subelliptica Perag. & Perag.
Peragallo & Peragallo, 22/7 (1897).

14

BMNH, Adams # F 20.2/7.45/12-14.119.3/8.

G211; TS 262.1/6-9.6/3.522 1/2.
maculata var. caribaea (Cl.) Cl.
Cleve, p. 46, 1895.
= Navicula caribaea Cl.
Cleve in Schmidt, et al., 6/10-12, 1874;
Mann, p. 97, 1925.
Bottom marine samples taken near the
islands by the USS Albatross, Mann (1.c.).
maculosa Donk.
Donkin, p. 25, 5/1, 1871;
Schmidt, et al., 259/25, 1874.

A checklist of Navicula

VISAYAS: Cebu, BMNH, Comber # 31899.

= Navicula diffusa Schmidt
Schmidt (1874) in Schmidt, et al., 2/28,
1874;
Mann, p. 101, 1925.
= Navicula madagascarensis Cl.
Cleve, p. 23, 4/2, 1891;
Mann, p. 108, 25/4, 1925.
Bottom marine samples taken near the
islands by the USS Albatross, Mann (1.c.).
malombensis Otto Mller*
Otto Mller, p. 84, 1/13, 1910.
= Navicula mammalis Cstr.
Castracane, p. 30, 20/2, 1886.
Bottom marine samples taken near
Philippine waters obtained by H.M.S.
Challenger, Castracane (1.c.).
mammalis Cstr.
Castracane, p. 30, 20/2, 1886.
Neighborhood of the Philippines, Castracane
(l.c.).
manokwariensis Choln.
Cholnoky, p. 181, f. 71, 1963.
= Navicula margarita Schmidt
Schmidt (1892) in Schmidt, et al., 174/17,
1874;
Mann, p. 108, 1925.
Bottom marine samples taken near the
islands by the USS Albatross, Mann (1.c.).
margaritifera Truan y Luard et Witt
Truan y Luard et Witt, p. 17, 4/10, 1888.

= Navicula marginata Lewis

Lewis, p. 64, 2/1, 1861;
Mann, p. 108, 1925.
Bottom marine samples taken near the
islands by the USS Albatross, Mann (1.c.).
mauriciana V.H.
Van Heurck, p. 11, 2/182, 1909.
= N. maxima Greg.
Gregory, p. 41, 4/19, 1855;
Mann, pp. 108-109, 1925.
Bottom marine samples taken near the
islands by the USS Albatross, Mann (1.c.).
mendica Mann
Mann, p. 109, 23/3, 1925.
Bottom marine sample taken near the islands
by the USS Albatross. Type-Cat. No. 43650,
USNM, Mann (1.c.).
menisculus Schum.
Schumann, p. 56, 2/33, 1867;
Skvortzow, p. 292, 2/8, 1937;
Hustedt, p. 70, 1942a.
LUZON: Rizal, Quezon City, Balara, Skvortzow
(l.c.); Manila (ponds, Bureau of Science grounds);
Batangas, Taal (lake); Camarines Sur, Buhi (lake);
Hustedt (1.c.).
minicans Hanna
Hanna, p. 200, 13/4, 1932.
= Navicula mimula Mann
Mann, p. 110, 23/4, 1925.
Bottom sediments taken near the islands
by the USS Albatross, Mann (1.c.).
minima Grun.
Grunow in Van Heurck, p. 107, 14/15, 1880-1885.

Not recognized by VanLandingham, 1975.

15

Martinez-Goss

= Navicula minima var. atomoides (Grun.) Cl.

Cleve, p. 128, 1894;
Hustedt, p. 56, 1942a.
LUZON: Rizal, Manila (ponds, Bureau of
Science grounds); Camarines Sur, Buhi Lake,
Hustedt (l.c.).
MINDANAO: Lanao (lake), Hustedt
(1.c.).
minuscula Grun.
Grunow in Van Heurck, 14/3, 1880;
Skvortzow, p. 291, 1/41, 1937;
Hustedt, p. 59, 1942a.
LUZON: Rizal, Quezon City (Balara Water
Filters), Skvortzow (1.c.).
MINDANAO: Lanao (lake, ponds), Hustedt
(1.c.).
mira Pant. & Greguss
Greguss in Greguss, p. 212, 6/8, 1913;
= Navicula mirabilis Leud.-Fortm.
Leuduger Fortmorel, p. 31, 2/21, 1879;
Mann, p. 110, 1925.
Bottom marine sediments taken near the
islands by the USS Albatross, rare, Mann (1.c.).
modica Hust.
Hustedt, p. 916, 41/21-23, 1945.

LUZON: Palawan, Taytay, Langen, Malapakan

Is. (surface scrapings from the sand in the cove, at
low tide); Podzorski & Hakansson (1.c.).
mucicoloides Hust.
Hustedt, p. 59, f. 98-102, 1942a.
LUZON: Benguet, Bontoc; Rizal, Lyon (ponds),
Hustedt (1.c.).
muralis Grun.
Grunow in Van Heurck, 14/27, 1880;
Hustedt, p. 59, 1942a.
LUZON: Camarines Sur, Buhi Lake, Hustedt
(1.c.).
mutica Ktz.
Ktzing, p. 93, 3/32, 1844;
Woltereck, p. 52, 1941;
Hustedt, p. 52, 1942a;
Podzorski & Hakansson, p. 85, 33/8, 1987.
LUZON: Benguet, Banaue (ponds), Woltereck
(1.c.); Laguna, Los Baos (ponds, Laguna de Bay);
Batangas, Taal Lake, Crater Lake; Camarines Sur,
Buhi Lake, Hustedt (1.c.); Palawan, Taytay,
Langen, Malapakan Is. (algae in the cove, marine);
Taytay, Lake Manguao (Danao), (alt. c. 45 m, at
the end of dry season, hence, lake at its shallowest);
Roxas, Pagdangan Range, Ibangley Brookside Hill
(alt. c. 40 m, on epiphyllous bryophyte beside a
river), Podzorski & Hakansson (1.c.).
VISAYAS: Leyte, Danao Lake, Hustedt (1.c.).

= Navicula molesta Mann

Mann, p. 110-111, 23/5, 1925.

MINDANAO: Lanao, Uyaan Lake, Hustedt

(1.c.).

BMNH, Adams # F 119, 7/3.

MINDANAO: Jolo, Sulu. Type-Cat. No.
43651, USNM, Mann (1.c.).
monilifera f. constricta (Perag. & Perag.) Hust.
Hustedt, p. 712, f. 1699B, 1966;
Podzorski & Hakansson, pp. 84-85, 34/1, 1987.

_______ var. cohnii (Hilse) Grun.

Grunow in Van Heurck, 10/17, 1880;
Hustedt, p. 52-53, 1942a.
LUZON: Rizal, Lyon; Laguna, Los Baos
(ponds), Hustedt (1.c.).
MINDANAO: Leyte, Uyaan Lake, Hustedt
(1.c.).

16

A checklist of Navicula

_______f. intermedia (Hust.) Hust.

Hustedt, p. 586, f. 1593, 1966;
Podzorski & Hakansson, p. 85, 33/9, 1987.
LUZON: Palawan, Roxas, Pagdangan Range,
(Ibangley Brookside Hill, from leaf surfaces),
Podzorski & Hakansson (1.c.).
= Navicula lagerheimii var. intermedia Hust.
Hustedt in Schmidt, et al., 370/22, 1874;
Hustedt, p. 54, 1942a.
LUZON: Oriental Mindoro, Calapan Lake,
Hustedt (1.c.).
_______ var. nivalis (Ehr.) Hust.
Hustedt, p. 290, 1911.
= Navicula mutica var. tropica Hust.
Hustedt, p. 233, 17/6, 1937b.
LUZON: Laguna, Los Baos (ponds);
Camarines Sur, Buhi Lake, Buhi River; Oriental
Mindoro, Naujan (river), Hustedt (1.c.).
nebulosa Greg.
Gregory, p. 480, 9/8, 1857;
Mann, p. 111, 1925.
Bottom sediments taken near the islands by
the USS Albatross, Mann (l.c.).
VISAYAS: Cebu, BMNH, Comber #31900.
nippon Skv.
Skvortzow, p. 277, 5/17, 1936.
= Navicula nitescens (Greg.) Ralfs*
Ralfs in Pritchard, p. 898, 1861;
Mann, p. 111, 1925.
Bottom sediments taken near the islands
by the USS Albatross, Mann (1.c.).

northumbricaeformis M. Perag
M. Pergallo in Heribaud, et al., p. 15, 1/19, 1920;
= Navicula notabilis Grev.
Greville, p. 18, 1/19, 1863;
Mann, p. 111, 1925.
Bottom sediments taken near the islands
by the USS Albatross, Mann, (l.c.).
nummularia Grev.
Greville, p. 249, 5/6, 1859;
De Toni, p. 96, 1891;
Mann, p. 111, 1925.
Bottom sediments taken near the islands by
the USS Albatross, Mann (1.c.).
= Navicula forcipata var. suborbicularis Grun.
Grunow in Van Heurck, 10/5, 1880.
Cleve, p. 66, 1895.
MINDANAO:Sulu Sea(Deby Coll.).
aff. ny Cl.
Cleve, p. 75, 1/24, 1894;
Podzorski & Hakansson, p. 75-76, 31/4, 1987.
LUZON: Palawan, Taytay, Langen, Malapakan
Is. (surface scrapings from sand at low tide in the
cove, marine), Podzorski & Hakansson (l.c.).
nympharum Hust.
Hustedt, 1936, in Schmidt, et al., 401/85-87,
1874.
= Navicula oamaruensis Grun.*
Grunow, 1888, in Schmidt, et al., 129/9, 1874__;
Mann, p. 111, 1925.
Bottom sediments taken near the islands
by the USS Albatross, Mann (l.c.).
obesa O. Mll.
O. Mller, p. 84, 2/27, 1910.

Not recognized by VanLandingham, 1975.

17

Martinez-Goss

= Navicula obesa (Grev.) Mann

Mann, pp. 111-112, 23/6, 24/1, 1925.
Bottom marine samples taken near the
islands by the USS Albatross. No type
designated, Mann (1.c.).
occulta Krasske
Krasske, p. 353, f. 12a-b, 1929.
= Navicula ocellata Mann*
Mann, p. 112, 24/2, 1925.
Bottom marine samples taken near the
islands by the USS Albatross. Type-Cat. No.
43652, USNM, Mann (1.c.).
oscitans Schmidt
Schmidt, 1875, in Schmidt, et al., 6/41, 1874;
Mann, p. 112, 1925.
Bottom marine samples taken near the islands
by the USS Albatross, Mann (1.c.).
______ var. subundulata Cl. & Grove
Grove, p. 67, 10/10, 1891.
BMNH, Adams # TS 523/5.8.
= Navicula suboscitans Mann
Mann, p. 122, 1925.
Bottom marine samples taken near the
islands by the USS Albatross, Mann (1.c.).
pacifica (Cstr.) Mann*
Castracane p. 23, pl. 20, f. a, 1886;
Mann, p. 113, 1925.
Bottom marine samples taken near the islands
by the USS Albatross, Mann (1.c.).
palpebralis var. semiplena (Greg.) Cl.
Cleve, p. 70, 1895.

Not recognized by VanLandingham, 1975.

18

= Navicula angulosa var. beta Greg.

Gregory, p. 42,5/8,1856;
Mann, p. 94, 1925.
Bottom sediments, Mann (l.c.).
palustris var. obesa Zanon
Zanon, p. 111, 2/10, 1949.
=Navicula pandura Breb.
Schmidt, et al., 11/1, 2, 4, 8, 9, 1874;
Mann, p. 113, 1925.
Bottom marine samples taken near the
islands by the USS Albatross, Mann (1.c.).
perrotettii (Grun.) Cl.
Cleve, p. 110, 3/12, 1894;
Hustedt, p. 51, 1942a.
Diatom samples from the Philippines of slightly
brackish waters derived from Dr. Raes collection,
Cleve (1.c.).
LUZON: Rizal, Manila (ponds, Bureau of
Science grounds); Laguna, Los Baos (Laguna de
Bay, ponds); Camarines Sur, Buhi (lake, river),
Hustedt (l.c.).
philippina Skv.
Skvortzow, p. 291, 1/25, 45, 1937a.
LUZON: Rizal, Quezon City, Balara (water
filter no. 8), Skvortzow (l.c.).
phillippinarum Mann
Mann, pp. 114-115, 25/1, 1925.
Bottom marine samples taken near the islands,
including those from the Sulu group, taken by the
USS Albatross. Type-Cat. No. 43655, USNM,
Mann (1.c.).
BMNH, Adams # F 83, 6/57; TS 523/9.
platyventris Meist.
Meister, p. 95, f. 33, 1934 (1935);
Podzorski & Hakansson, p. 81, 32/3, 1987.

A checklist of Navicula

LUZON: Palawan, Langen, Malapakan Is.

(surface scrapings from the sand at low tide in the
cove), Podzorski & Hakansson (1.c.).

VISAYAS: Cebu, BMNH, Comber # 31898,

31895, 31894.
_______ var. ?

plicatula Grun.
Cleve & Mller, slide #154-155;
Grunow Ms. in Cleve, p. 155, 3/28, 1894;
Mann, pp. 115-116, 24/8-9, 1925;
Podzorski & Hakansson, p. 78, 30/8, 1987.
Bottom marine samples taken near the islands
by the USS Albatross, Mann (1.c.).
LUZON: Palawan, Taytay, Langen, Malapakan
Is. (algae from the cove, marine), Podzorski &
Hakansson (1.c.).
VISAYAS: Cebu, BMNH, Comber # 31894.
plutonia O Meara*
OMeara, p. 286, 13/6, 1872;
De Toni, p.78, 1891.
MINDANAO: Sulu, Cagayan, (brackishwaters
from the volcanic mountain), De Toni (l. c.).
praetexta Ehr.
Ehrenberg, p. 214, 1840 (1841);
Castracane, p.812, 1886;
Mann, p. 116, 1925.
Bottom marine samples taken near the islands
by the USS Albatross, Mann (1.c.).
BMNH, Adams # GC 2094; F 119 3/9; TS
497.1/10; TS 523/11; TS 60/81.
LUZON: Manila (on shell), BMNH, Comber
# 30925.
VISAYAS: Cebu, Castracane (l.c.).
_______ var. abundans Schmidt
Schmidt, et al., 129/8, 1874;
Cleve, p. 55, 1895.

BMNH, Adams # F 45.2/16.

protracta (Grun.) Cl.
Cleve, p. 140, 1894;
Podzorski & Hakansson, p. 77, 33/13, 1987.
LUZON: Palawan, Taytay (Lake Manguao,
alt. c. 45 m., end of the dry season when it was at
its shallowest, surrounded by semi-deciduous
rainforest), Podzorski & Hakansson (1.c.).
pseudobryophila Hust.
Hustedt, p. 60, 1942a;
_______, p. 114, 1942b.
VISAYAS: Leyte (Danao Lake), Hustedt
(1942a).
pseudoclavata Mann
Mann, p. 116, 25/2, 1925.
Bottom marine samples taken near the islands
by the USS Albatross. Type-Cat. No. 43656,
USNM, Mann (1.c.).
pseudoforcipata Hust.
Hustedt, pp. 119-120 f. 11, 1942b;
Podzorski & Hakansson, p. 80, 31/67, 1987.
LUZON: Palawan, Taytay, Langen, Malapakan
Is. (from the cove, marine); Pto. Princesa (south
of Pto. Princesa Bay, from the rapidly deepening
zone of the Sulu Sea, hand collected from corals at
10 m depth), Podzorski & Hakansson (1.c.).
pseudony Hust.
Hustedt, p. 23, 8/11, 1955;
Podzorski & Hakansson, p. 80, 30/2, 1987.
LUZON: Palawan, Taytay, Langen, Malapakan
Is. (algae encrusted from the cove, marine),
Podzorski & Hakansson (1.c.).

Not recognized by VanLandingham, 1975.

19

Martinez-Goss

VISAYAS: Leyte (Danao Lake), Hustedt

(1.c.).

pugio Mann
Mann, p. 117, 26/4, 1925.

MINDANAO: Lanao (Uyaan Lake),

Hustedt (1.c.).

Bottom marine samples taken near the islands

by the USS Albatross, Mann (1.c.).
pulverulenta (pulvulenta) Mann
Mann, pp. 117-118, 25/3, 1925.
Bottom marine samples taken near the islands
by the USS Albatross. Type-Cat. No. 43659,
USNM, Mann (1.c.).
pupula Ktz.
Ktzing, p. 93, 30/40, 1844;
Woltereck, p. 52, 1941;
Hustedt, p. 64, 1942a;
Podzorski & Hakansson, p. 76, 30/3, 1987.
LUZON: Benguet, Banaue (ponds), Woltereck,
1941; Rizal, Malabon, Dagatan (swamp), Manila
(ponds in the Bureau of Science grounds, Pancipit
River, Reservoir II); Laguna, Los Baos (ponds,
Laguna de Bay), San Pablo (Bunot Lake, Bato
Lake); Batangas, Taal (Yambo Crater Lake, Crater
Lake); Camarines Sur, Buhi (lake, river); Payren
Lake; Oriental Mindoro, Naujan Lake, Hustedt
(1.c); Palawan, Taytay (Lake Manguao, alt. 45 m,
near Public Works resthouse, at the end of the dry
season, when lake was shallowest; small pool in
stream bed flowing into NW side of the lake, 1.8
km from Public Works resthouse, alt. 130 m),
Podzorski & Hakansson (1.c.).

_______ f. rostrata (Hust.) Hust.

Hustedt, p. 291, 3/39, 1911;
_______, p. 64, 1942a;
_______, p. 282, 1957.
LUZON: Laguna, Los Baos (Laguna de Bay,
pond); Batangas, Taal (Taal Lake); Camarines Sur,
Buhi (lake), Hustedt (1942a).
MINDANAO: Lanao (pond), Hustedt (1942a).
pygmaea Ktz.
Ktzing, p. 77, 1849;
Woltereck, p. 52, 1941;
Hustedt, pp. 66-67, 1942a;
Podzorski & Hakansson, p. 80, 31/11, 1987.
LUZON: Benguet, Banaue (ponds), Woltereck
(1.c.); Rizal, Manila (Bureau of Science grounds,
ponds), Pancipit River, Lyon (ponds); Laguna, Los
Baos (ponds, Laguna de Bay); Camarines Sur,
Buhi (river), Hustedt (1.c.); Palawan, Pto. Princesa
(south of Pto. Princesa Bay, hand collected from
sand surface at 15 m depth); Taytay, Langen,
Malapakan Is. (surface scrapings from sand at low
tide), Podzorski & Hakansson (1.c.).
MINDANAO: Lanao (Lanao Lake), Jolo, Sulu
(Seit Lake), Hustedt (1.c.).

MINDANAO: Lanao (Uyaan Lake, Lanao

Lake, river); Jolo, Sulu (Seit Lake), Hustedt (1.c.).
_______ var. rectangularis (Greg.) Cl. & Grun.
Cleve & Grunow, p. 45, 1880.

quarnerensoides Hust.*
Hustedt, p. 49, f. 1203, 1961; (doubtful-fide:
VanLandingham, p.2759,1975);
Podzorski & Hakansson, 78, 1987.

= Navicula f. rectangularis (Greg.) Cl. & Grun.

Hustedt, pp. 64-65, 1942a;
Podzorski & Hakansson, p. 76, 30/4, 1987.
LUZON: Palawan, Roxas, Pagdanan
Range (Ibangley Brookside Hill, alt. 40 m,
streamside pool), Podzorski & Hakansson
(1.c.).

20

LUZON: Palawan, Pto. Princesa, Table Head

(hand collected from sand surface at 15 m depth
of Sulu Sea), Podzorski & Hakansson (1.c.).

Not recognized by VanLandingham, 1975.

A checklist of Navicula

radiosa Ktz.
Ktzing, p. 91, 4/23, 1844;
Woltereck, p. 52, 1941;
Hustedt, pp. 69-70, 1942a;
Podzorski & Hakansson, p. 82, 33/2, 3.
LUZON: Benguet, Banaue (ponds); Rizal,
Malabon (Dagatan swamp), Manila (Water
Reservoir III, Ponds in Bureau of Science grounds,
Lyon ponds), Hustedt (1.c.); Brookes Point
(Tumarabong River over Oligocene-Miocene reef
limestones, alt. 10 m, scrapings from the surface
of a limestone rock), Podzorski & Hakansson (1.c.).
_______ var. tenella (Breb. ex Ktz.) Cl. et Mll.
(non Grun. fide VanLandingham, 1975);
Cleve et Mller slide no. 224, 1881;
Hustedt, pp. 69-70, 1942a;
Podzorski & Hakansson, p. 82, 33/4, 1987.
LUZON: Batangas, Taal (Volcano Lake),
Hustedt (1.c.); Palawan, Pto. Princesa (Irawan
River over mafic and mafic-plutonic rocks, alt. c.
150 m; scrapings from a bryophyte behind a
waterfall), Podzorski & Hakansson, (1.c.).
raena (Cstr.) De T.
De Toni, p. 30, 1891;
Mann, p. 118, 1925.
Bottom marine samples taken near the islands
by the USS Albatross. Type species from the
Philippines (fide: Castracane below), Mann (1.c.).
= Pinnularia raena Cstr.
Castracane, p. 25-26, 15/3, 1886.

retracta Meist.
Meister, p. 262, 4/6, 1937;
= Navicula retrostauros Mann*
Mann, p. 118, 25/2, 26/1, 2, 1925.
BMNH, Adams #M 59.1/8; TS 60/59.
Bottom marine samples taken near the
islands by the USS Albatross. Type-Cat. No.
43660 USNM, Mann (l.c.).
rhaphoneis (Ehr.) Grun.
Grunow, p. 19, 91/17, 1867;
Podzorski & Hakansson, pp. 82-83, 32/6, 7, 1987.
LUZON: Palawan, Taytay, Langen, Malapakan
Is. (algae scraped from the cove), Podzorski &
Hakansson (1.c.).
rhombica Greg.
Gregory, p. 40, 4/16, 1855;
Mann, p. 119, 1925.
Bottom marine samples taken near the islands
by the USS Albatross, Mann (1.c.).
_______var. japonica (Brun.) Cl.
Cleve, p. 152, 1894.
BMNH, Adams #TS 523/2.
rhynchocephala (rhynococephala) Ktz.
Ktzing, p. 152, 30/35, 1844;
Hustedt, p. 67, 72, 1942a;
delos Reyes, p. 59, pl. 4, f. 17, 1972.

VISAYAS: Cebu, from Dr. Raes

collection,
Castracane (1.c.).
ramosissima (Ag.) Cl.
Cleve, p. 26, 1895;
Podzorski & Hakansson, p. 82, 34/3, 1987.
LUZON: Palawan, Taytay, Langen, Malapakan
Is. (algae encrusted from the cove, marine),
Podzorski & Hakansson (1.c.).

LUZON: Laguna, Los Baos (Laguna de Bay);

Batangas, Taal (Crater Lake); Hustedt (1.c.).
VISAYAS: (Tigbao water supply), delos Reyes
(1.c.).
riparia Hust.
Hustedt, p. 52, f. 77-78, 1942a.

Not recognized by VanLandingham, 1975.

21

Martinez-Goss

LUZON: Rizal, Malabon (Dagatan Swamp),

Hustedt (1.c.).
robertsiana (robertsoniana) Grev.
Greville, p. 235, 3/9, 1865.
VISAYAS: Cebu, BMNH, Comber #31895.
_______ var. cuneata (Schmidt) Amosse
Amosse, p. 111, 1924.
= N. prodiga Mann
Mann, p. 252, 53/4, 1907;
______, p. 116, 1925.
Bottom marine samples taken near the
islands by the USS Albatross, Mann (1925).
ruttneri Hust.
Hustedt, 1936, in Schmidt, et al., 402/30-38, 49-52,
1874;
_______, p. 55, 1942a.
LUZON: Camarines Sur, Buhi (lake), Hustedt
(1942a).
_______ var. capitata Hust.
Hustedt, 1936, in Schmidt, et al., 402/47, 48,
1874;
_______, p. 55, 1942a.
LUZON: Manila (Reservoir II), Oriental
Mindoro, Naujan (river), Hustedt (l.c.).
_______ var. rostrata Hust.
Hustedt, 1936, in Schmidt, et al., 402/39-45,
1874;
______, p. 56, 1942a.
LUZON: Rizal, Pancipit River; Oriental
Mindoro, Naujan (Naujan Lake), Hustedt (1942a).

LUZON: Laguna, Los Baos, Mayondon,

Pantastico (1.c.).
schaarschmidtii Pantocsek
Pantocsek, p. 28, 14/121, 1886.
VISAYAS: Cebu, BMNH, Comber #31900.
schoenfeldii Hust.
Hustedt, p. 301, f. 520, 1930;
_______, p. 70, 1942a.
LUZON: Oriental Mindoro, Naujan (river),
Hustedt (1942a).
schroeteri Meist.
Meister, p. 38, f. 100, 1932;
Hustedt, p. 72, 1942a.
LUZON: Laguna, Los Baos (Laguna de Bay);
Batangas, Taal (Taal Lake); Oriental Mindoro,
Naujan (river), Hustedt (1.c.).
_______ var. escambica Patr.
Patrick, p. 104, 7/3, 1959;
Podzorski & Hakansson, p.83, 33/5, 1987.
LUZON: Palawan, Roxas, Pagdanan Range,
(Ibangley Brookside Hill streamside freshwater
pool, alt. c. 40 m.), Podzorski & Hakansson (l.c.).

scutelloides W. Sm.
Smith, p.9, 1856;
_____, in Gregory, p. 4, 1/15, 1856;
Hustedt, p. 66, 1942a.
LUZON: Batangas, Taal (Crater Lake);
Camarines Sur, Buhi (lake), Hustedt. (l.c.)
MINDANAO: Lanao (Lanao Lake, pond),
Hustedt (1.c.).

MINDANAO: Lanao (pond), Hustedt (1942a).

salinarum Grun.
Grunow in Cleve & Mller, slide no. 107, 1878;
Grunow in Cleve & Grunow, p. 33, 2/34, 1880;
Pantastico, p. 195, 17/3, 1977.

22

seminulum Grun.Grunow, p. 552, 4/3(2/3), 1860;

Hustedt, p. 56, 1942a.
LUZON: Camarines Sur, Buhi (lake);
Batangas, Taal (Crater Lake).

A checklist of Navicula

MINDANAO: Lanao (Lanao Lake, pond);

Hustedt (l.c.).
*

semistauros Mann
Mann, 119-120, 26/5, 1925.
Bottom marine samples taken near the islands
by the USS Albatross, Type-Cat. No. 43661,
USNM, Mann (l.c.).
sigma Brun
Brun, p. 39, 16/5, 1891;
Brun later homonym, Fide VanLandingham p. 2799,
1975;
Podzorski & Hakansson, p. 75, 29/12, 1987.
LUZON: Palawan, Pto. Princesa, Table Head
(from sand surface at 15 m depth of Sulu Sea),
Podzorski & Hakansson (1.c.).
spectabilis Greg.
Gregory, p. 481, 9/10, 1857;
Cleve, p. 60, 1895;
Mann, p. 121, 27/1-2, 1925;
Podzorski & Hakansson, pp. 80-81, 32/1, 1987.
Bottom marine samples taken near the islands
by the USS Albatross, Mann (1.c.).
Diatom samples (presumably from Dr. Raes
collection), Cleve, 1894, p. 60, 1895.
BMNH, Adams # J 4006; Bess 1709/3; G 211;
F 40.2/16.45.1/11.
LUZON: Palawan, Pto. Princesa, Table Head
(from the sand surface at 15 m depth of Sulu Deep
Sea), Podzorski & Hakansson (1.c.).

BMNH, Adams # TS 60/4.

_______ var. emarginata Cl.
Cleve p. 60, 1895.
= Navicula excavata var. ________ Schmidt
Schmidt in Schmidt, et al., 3/22-25, 1874.
VISAYAS: Cebu, BMNH, Comber
#31899.
_______ var. excavata (Grev.) Cl.
Cleve, p. 61, 1895;
= N. excavata Grev.
Greville, p. 130, 12/15, 1866;
Mann, p. 102, 1925.
Bottom marine samples taken near the
islands by the USS Albatross, Mann (1.c.).
BMNH, Adams # F 83.1/9.119.2/4; TS
522.2/2.
= N. oswaldii Janisch
Janisch, 1, 15/12, 1888;
Mann, p. 113, 1925.
Bottom marine samples taken near the
islands by the USS Albatross, Mann (1.c.).
_______ var. minuta Cl.
Mereschkowsky, p. 320, 2/9-12, 1902.
= Navicula hennedyi var. minuta Cl.
Cleve, p. 7, 1/15, 1881.
VISAYAS: Cebu, BMNH, Comber
#31894.

VISAYAS: Cebu, BMNH, Comber #318931.

_______ var. ?
_______ var. controversa (Schmidt) Cl.
Schmidt, et al., 3/5, 1874;
Cleve, p. 60, 1895.

BMNH, Adams # F 931; Bess 1710/3; TS 497.

2/7-9.
spiculifera Mann*
Mann, pp. 121-122, 26/8, 1925.

Not recognized by VanLandingham, 1975.

23

Martinez-Goss

Bottom marine samples taken near the islands

by the USS Albatross. Type-Cat. No. 4663, USNM,
Mann (1.c.).
spiralis O Meara*
De Toni, p. 74, 1891.
MINDANAO: Saline waters from the volcanic
crater, Cagayan, Sulu Archipelago, De Toni (l.c.).

LUZON: Rizal, Manila (Reservoir II), Hustedt

(1942a).
subforcipata Hust.
Hustedt, p. 533, f. 1569, 1964;
Podzorski & Hakansson, p. 75, 31/51, 1987.
LUZON: Palawan, Taytay, Langen, Malapakan
Is. (surface scrapings from sand at low tide),
Podzorski & Hakansson (1.c.).

studerri sensu Comber*

LUZON: Manila (sponge), BMNH, Comber #
30924.
subarvensis Hust.
Hustedt, p. 60, f. 107-108, 1942a.
LUZON: Camarines Sur, Buhi (Buhi Lake),
Hustedt (1.c.).
= N. lyra var. subcarinata Grun.
Grunow, 1874, in Schmidt, et al., 2/5, 1874;
Mann, p. 108, 1925.
Bottom marine samples taken near the
islands by the USS Albatross, Mann (1.c.).
BMNH, Bess 62; F 40 2/19; TS 497.2/6,
522, 2/6, 3/8.
LUZON: Manila, BMNH, Comber
#31903.
subdecipiens De T.
De Toni, p. 147, 1891.
= N. decipiens Cstr.
Castracane, p. 32, 27/17, 1886.
From Philippine marine diatom collection
of Dr. Rae, Castracane (l.c.).
subdecussis Hust.
Hustedt, pp. 75-76, f. 136-138, 1942a.

subrhynchocephala Hust.
Hustedt, p. 156, 1/11, 1935;
_______, p. 67, 1942a.
LUZON: Rizal, Manila (Reservoir II, Pancipit
River); Laguna, Los Baos (pond, Laguna de Bay,
Payren Lake); Batangas, Taal (Crater Lake);
Camarines Sur, Buhi (Buhi Lake, river), Hustedt
(1942a).
VISAYAS: Leyte (Danao Lake), Hustedt
(1942a.).
suluensis O Meara*
De Toni, p. 80, 1891.
MINDANAO: Brackishwaters in the crater
of a volcanic island, Cagayan, Sulu, De Toni (l.c.).
takoradiensis Hendey
Hendey, p. 67, 1/8, 1958;
Podzorski & Hakansson, p. 77, 30/7, 1987.
LUZON: Palawan, Pto. Princesa, Table Head
(from sand surface at 15 m depth of the Sulu Sea),
Podzorski & Hakansson (1.c.).
tenera Hust.
Hustedt, p. 259, 18/11-12, 1937a;
_______, p. 67, 1942a.
LUZON: Rizal, Manila (Pancipit River);
Batangas, Taal (Crater Lake); Hustedt (1942a).
terminata Hust.
Hustedt, p. 589, f. 1594, 1966;

Not recognized by VanLandingham, 1975.

24

A checklist of Navicula

= N. mutica var. tropica Hust.

Hustedt, 1936 in Schmidt, et al., 405/39-42,
1874;
_______, pp. 53-54, 1942a.
LUZON: Laguna, Los Baos (pond);
Camarines Sur (Buhi Lake); Oriental Mindoro,
Naujan (Naujan River), Hustedt (1942a).
translucens Mann
Mann, p. 123, 26/9, 1925.
Bottom marine samples taken near the islands
by the USS Albatross. Type-Cat. No. 43665,
USNM, Mann (1.c.).
tripunctata (O.F. Mll.) Bory*
Bory, p. 128, 1822;
Podzorski & Hakansson, p. 83, 33/6, 1987.
LUZON: Palawan, Taytay, Langen, Malapakan
Is. (scrapings from sand surface at low tide), Pto.
Princesa, Honda Bay (scraping from coral reefs at
the edge of an atoll), Podzorski & Hakansson (1.c.).
uniseriata strup
strup, p. 8, 1/9, 1913;
Podzorski & Hakansson, pp. 77, 33/12, 1987.
LUZON: Palawan, Taytay, Langen, Malapakan
Is. (algae encrusted from cove), Podzorski &
Hakansson (1.c.).
variostriata Krasske
Krasske, p. 197, f. 12, 1923;
Hustedt, p. 55, 1942a.
LUZON: Laguna (Laguna de Bay), Hustedt
(1.c.).
venusta Janisch
Janisch, 15/17, 1888;
Mann, p. 124, 1925.

Bottom marine samples taken near the islands

by the USS Albatross, Mann (1.c.).
venustissima Kitt.
Kitton, p. 101, 1889;
Kitton in Leuduger-Fortmorel., p. 17, 2/3, 1892.
BMNH, Adams # GC 2177.
viminea Hust.
Hustedt, 1934 in Schmidt, et al., 397/10-11,
1874;
Podzorski & Hakansson, p. 83, 32/8-9, 1987.
LUZON: Palawan, Taytay, Langen, Malapakan
Is. (surface scrapings from sand at low tide),
Podzorski & Hakansson (1.c.).
viridula (Ktz.) Ehr.
Ehrenberg, p. 53, 1836.
= Navicula viridula Ktz.
Ktzing, p. 23, 13/12, 1833;
Hustedt, p. 72, 1942a.
LUZON: Laguna, Los Baos (pond);
Camarines Sur, Buhi (Buhi Lake), Hustedt
(1.c.).
werestschangini (werestschangini) Skv. & Meyer
Skvortzow & Meyer, p. 20, 1/64, 1928;
Podzorski & Hakansson, p. 85, 34/5, 1987.
LUZON: Palawan, Taytay, Langen, Malapakan
Is. (surface scrapings from sand at low tide),
Podzorski & Hakansson (1.c.).
yarrensis (jarrensis) Grun.
Grunow 1876, in Schmidt, et al., 46/1-6, 1874;
Mann, p. 126, 1925;
Podzorski & Hakansson, p. 79, 31/3, 1987.
Bottom marine samples taken near the islands
by the USS Albatross, Mann (1.c.).
BMNH, Adams # GC 2190; TS 262.1/
4.5.18.6/1.2.89.

Not recognized by VanLandingham, 1975.

25

Martinez-Goss

LUZON: Palawan, Taytay, Langen, Malapakan

Is. (algae scraped from the cove), Podzorski &
Hakansson (1.c.).
zanzibarica Grev. var. zebuana Cstr.
Greville, p. 129, 12/22, 1866;
Castracane, p. 31, 28/8, 1886;
A. Schmidt, 1888 in A. Schmidt, et al., 129/4,
1874.
VISAYAS: Cebu, Castracane (1.c.).
zostereti (zosteretii) Grun.
Grunow, p. 528, 4/23, (2/23) 1860;
Mann, p. 125, 1925;
Podzorski & Hakansson, pp. 83-84, 33/10, 1987.
Bottom marine samples taken near the islands
by the USS Albatross, Mann (1.c.).
LUZON: Palawan, Taytay, Langen, Malapakan
Cove (scrapings from the cove), Pto. Princesa,
White Beach (on mud surface in a mangrove),
Honda Bay (coral scrapings from a bay), Podzorski
& Hakansson (1.c.).
sp. (miniscula)
Podzorski & Hakansson, p. 81, 19/11, 1987.
LUZON: Palawan, Taytay, Langen, Malapakan
Cove (scraping from sand surface at low tide),
Podzorski & Hakansson (1.c.).
sp. 1
Esguerra, p. 174, 1951;
Villadolid, p. 21, 1957.
sp. 2
BMNH, Adams-Bess # M64-66; TS 80; Bess 22/
6.714; TS 522.2/3.7.3/3.
VISAYAS: Cebu, Castracane, p. 812, 1886.
sp. 3
White chalk collected by the ship Princess Louise
to Manila, under Capt. Wendt (1830-34).
Identification by Dr. Meyer. Ehrenberg, Atlas, p.
177, pl. 1, 1854.

26

ACKNOWLEDGMENTS
This checklist was completed thru the financial
assistance of the Jessup-McHenry Fund Grant of the
Academy of Natural Sciences of Philadelphia (ANSP),
U.S.A. and the British Council of England; the kind
hospitality in making available their references and
specimens of the ANSP and the British Museum of
National History; the patience in encoding and revising
the paper by C.M. Nuez, M.V. Marasigan and W.F.
Fabelllore.

REFERENCES
Amosse, A., 1924. Diatomees de la cote orientale d
Afrique. Bulletin du Museum National d Histoire
Naturalle. t.20 (1924), p.109-116, 159-166, 247-254,
329-335.
Bailey, J.W., 1853. List of diatomaceae collected by
the United States Exploring Expedition under Capt.
Wilkes, U.S.N. Proc. Nat. Acad. Sci. Phila. 6:431-432.
Bory de Saint-Vincent, J.B.M, 1822-1831. Collaborator
in Dictionaire classique d Histoire Naturelle, 17 vol.
Paris. Atlas et Illustration des Planches t. 17, 1831.
(Diatomaceae are scattered throughout the work
alphabetically).
Brockman, C., 1950. Die Watt-Diatomeen der
schleswig-holsteinischen Westkste. Abhandlungen
d e r s e n c k e n b e rg i s c h e n N a t u r f o r s c h e n d e n
Gesellschaft. 478: 1-26, 6 Taf.
Brun, J., 1891. Diatomees especes nouvelles marines,
fossiles ou pelagiques, Geneve, Imprimerie AubertSchuchardt. 47p., 12 pl.
Castracane, C.A.F., 1886. Report on the Scientific
results of the Voyage of H.M.S. Challenger during the
years 1873-1876; Botany, London, Her Majestys
Stationary Office, vol. 2, 178p., 30 pl.
Cholnoky, B.J., 1955. Diatomeen aus salzhaltigen
Binnengen Binnengewsser der westlichen KaapProvincz in Sdafrika. Berichte der Dentschen
Botanischen Gesellschaft, Bd. 68, S. 11-23.

A checklist of Navicula

Cholnoky, B.J., 1963. Ein beitrag zur kenntnis der

Diatomeen flora von Hollan disch New Guinea. Nova
Hedwigia. 5(1-4): 157-198, 3 Taf.
Cleve, P.T., 1878. Diatoms erom the West Indian
Archipelago. Bihang till Kongl. Svenska VetenskapsAkadmiens Handlinger. 5(8):1-22, pl. 1-5.
Cleve, P.T., 1881. On some new and little known
Diatoms. Konglinga Svenska Vetenskaps-Akadmiens
Handlinger. 18(5): 1-28, pl. 1-6.

Ehrenberg, C.G., 1830 (1832). Beitrge zur Kenntniss

der Organization der Infusorien und iher geographischen
Verbreitung besonders in Sibirien. (Preussische
physikalische) Abhandlungen der Kniglichen
Akademie der Wissenschaften zu Berlin, S. 1-88, 8
Taf.
Ehrenberg, C.G., 1830 (1839). Uber die Bildung der
Kreidefelsen und des Kreidemergels durch unsichtbare
Organismen. Abhandlungen der Kniglichen Akademie
der Wissenschaften zu Berlin, S. 59-147, Taf. 1-4.

Cleve, P.T., 1894. Sur quel ques especes nouvelles ou

peu connues. Le Diatomiste. 2(17):99.

Ehrenberg, C.G., 1836. Nachrichten uber das

Vorkommen fossiler infusorien. Bericht die zur
Bekanntmachung geeigneten Verhandlungen der
koniglichen preuss. Akademie der Wissenschaften zu
Berlin, Erster Jahrgang, 1836, S. 83-86.

Cleve, P.T., 1895. Vetens Kaps-akademiens Handlingar,

Bd. 26, p. 1-194, pl. 1-5 (Part I (1894)); Bd. 27, p. 1219, pl. 1-4 (Part II (1895)).

Ehrenberg, C.G., 1837. Mittheilungen aus den

Verhandlungen der Gaselllschaft Naturforschender
Freunde zu Berlin 31 Marz 1837, etc.

Cleve, P.T. & J.D. Moller, (eds), 1877-1882. Diatoms

(Exsiccata). Upsala, Sweden, Esatas Edquists
Boktryckeri (This series of slides consists of 324 slides
and were all examined by A. Grunow).

Ehrenberg, C.G., 1838.Die Infusionsthierchen als

vollkommende Organismen. Ein Blick in das tiefere
Organiche Leben der Natur. S. I-XVII, 1-548, Taf. 164 (Atlas). Leopold Vass, Leipzig.

Cleve, P.T. & A. Grunow, 1880. Beitrage zur Kenntriss

der Artischen Diatomeen. Kongl. Svenska Veten.Akadem. Handl. Stockholm, Sweden, Kongl
Boktryckeriet. 17(2):121p., 7 pl.

Ehrenberg, C.G., 1841(1843sic).Verbreltung und

Einfluss des mikroskopischan Lebens in Sud-und
Nordamerika. Bericht uber die zur Bekanntmachung
gee igneten Verhandlungen der koniglichen Akademie
da Wissonschaften zer Berlin, Theil 1, s. 291-445, 4
Taf.

Cleve, P.T., 1891. Diatomees rares on nonvelles. Le

Diatomiste. 1: 75-78.

De los Reyes, P., 1972. The algae of the Tigbao water

supply. Leyte-Samar Studies 6:48-60.
De Toni, G.B., 1894. Sylloge algarum omnium
hucusque cognitarum, vol. II, Bacillarieae. Section I,
Raphideae, p. 1-490 (1891); Section II,
Pseudoraphideae, p. 491-817 (1892); Section III,
Cryptoraphideae, p. 818-1556.
Donkin, A. 1861. On marine Diatomeceae of
Northumberland with a description of several new species.
Quart. J. Microsc. Sci. 1 (New series): 1-15, pl. 1
Donkin, A. 1870-1873. The Natural History of the
British Diatomaceae. Part 1, p. 1-24, pl. 1-4 (1870);
Part 2, p. 25-48, pl. 5-8 (1871). London.

Ehrenberg, C.G., 1840 (1841). Uber ausgezeichnete

jetzt leberde peruaniche und mexikanische MeeresInfusorien, welche mit zur Erlauterung rathselhaften
fossilen Formen der Kreidebildung dienen. Bericht ber
die zur Bekanntmachung geeigneten verhandlungen der
knigl. Preuss. Akademie der wissenschaften zu Berlin,
S. 157-162.
Ehrenberg, C.G., 1854. Die systematische charakteristik
der neuen mikroskopichen Organismen des tiefen
Atlantischen Ocean fr den Monatsbericht zum Druck
zu bergeben, deren verzeichniss in Monat Februar
bereits mitgetheilt worden ist. Berucht ber die zur
Bechantmachung geeigneten verhandlungen der

27

Martinez-Goss

kniglichen preuss. Akademie der Wissenschaften zu

Berlin, S. 236-252.
Esguerra, R.S., 1951. Enumeration of algae in Philippine
bangos fishponds and in the digestive tract of the fish
with notes on conditions favorable for their growth. Phil.
J. Fish , 1: 175-196.
Gregory, W., 1855. On a Post-Tertiary Lacustrine
Sand containing Diatomaceous exuviae from
Glenshina near Inverness. Quart. J. Microsc. Sci.
3: 30-43, pl. 4.
Gregory, W., 1856. Notice of some new species of
British Freshwater Diatomaceae. Quart. J. of
Microsc. Sci. 4:1-14 (for the figure see: vol. IV, pl. 1,
f. 1-42).
Gregory, W., 1857. On new forms of marine
diatomaceae found in the Firth of Clyde and in Loch
Fyne. (drawn by R.K. Greville) Trans. Roy. Soc. of
Edinb. 21:473-542, pl. 9-14.

Greville, R.K., 1866. Descriptions of new and rare

diatoms. Series 20. Trans. Microsoc. Soc. 14
(N.S.):121-130, pl. 11 & 12, 25f.
Grove, E., (many dates). Collection (exsiccatae) of E.
Grove. Preston, Brighton.
Grunow, A., 1860. Weber neue oder ungeniigend
gekannte Algen. Aus den Verhandlungen der K.K.
Zoologisch-botanischen Gesellschaft in Wien
[Jahrgang 1860] besconclers abgedruckt. pp. 1-80, pl.
3-7.
Grunow, A., 1863. Uber einige neue und ungenugend
bekannte Arten und Gattungen von Diatomaceen.
Zweite Folge. Verhandl. der kaiserlich-konigl.
Zoologisch-botanischen Gesselschaft in Wien.
13:137-162, pls. 13-14.
Grunow, A., 1867. Diatomeen auf Sargassum von
Honduras, gesammelt von Lindig. Hedwigia. 6(1-3):
1-8, 17-37.

Greville, R.K., 1857. Description of some new

Diatomaceous forms from the West Indies. Quart. J.
Microsc. Sci. 5: 7-12, pl.3

Grunow, A., 1880. Vorlufige Bemerkungen zu einer

systematischen Anordung der Schizonema und
Berkeleya Arten, mit Bezug auf die in Van Heurchs
Diatocaceen- Flora von Belgien verffetlichten
Abbildungen der Frusteln auf Taf. XV, XVI, and XVII.
Botanisches Centralblatt. 4(47/48): 1506-1520, 15851598.

Greville, R.K., 1859. Descriptions of some new species

and varieties of Naviculae, etc., observed in California
guano. Edinb. New Philo. J. N.S. 10:25-30, pl. 4, 8
fig.

Hagelstein, R., 1938. The Diatomaceae of Puerto Rico

and the Virgin Islands. New York Academy of Sciences,
Survey of Porto Rico and the Virgin Islands. 8(3):
313-450, 9 plates.

Greville, R.K., 1862. Descriptions of new and rare

diatoms. Series VI. Trans. Microsc. Soc. 10:89-96, pl.
9, (pp. 1-7).

Hagelstein, R., 1938(1939). The Diatomaceae of Puerto

Rico and the Virgin Islands. New York Academy of
Science Survey of Puerto Rico and Virgin Islands.
8(3): 313-450, 9 pls.

Greguss, P. 1913. Die Kieselalgen der Meeraugen von

Surian. (A Surianitengerszemek kovamoszatai.)
Botanikai Kzlemenyek, Bd. 12, S. 202-225.

Greville, R.K., 1863. Descriptions of new genera and

species of diatoms from the South Pacific. Part I.
Edinb. New Philo. Journ. 18:34-43, pl. 1, 43 fig.
Greville, R.K., 1865. Description of new and rare
Diatoms. Trans. Microsc. Soc. 13 (N.S.): 1-10, 2 plates
(I-II), Series XV, p. 24-34, 2 plates (V-VI), Series
XVII< p. 97-105, 2 plates (VIII-IX).

28

Hanna, G.D., 1930. The dates and publication of

Tempere and Peragallos Diatomees du Monde entier,
Edition 2. J. Paleontology 4(3):296-297, 1-68p.
Hanna, G.D., 1932. Pliocene Diatoms of Wallace
Country, Kansas. University of Kansas Science
Bulletin 20(21): 369-394, pls. 31-34.

A checklist of Navicula

Heiberg, P.A.C., 1863. Conspectus criticus

Diatomacearum Danicarum. Wilhelm Priors Forlag,
Kjobenhavn: 135 p. 6 pls.
Hendey, N.I. 1958. Marine Diatoms from some West
African ports. Journ. of the Royal Microsc. Soc. 77(3):
28-85.
Heribaud, J., et al., 1920. Les Diatomees des travertins
d Auvergue. Bruxelles, Imprimerie de Scintifique: 206
p., 7 pl.
Hustedt, F., 1911. Beitrge zur Algenflora von Bremen.
IV. Bacillariaceen dus der Wumme. Abh. Naturw. ver.
Brem. 209(2): pp. 257-315, 3 pl.
Hustedt, F., 1930. Bacillariophyta (Diatomeae). In A.
Pascher Die Susswasser-Flora Mitteleuropas, Heft
10, 466 s., 875 fig., Jena, Gustav Fischa Verlag.
Hustedt, F., 1930 - 1966. Die Kieselalgen
Deutschlands, sterreichs und der Schweiz unter
Berucksichtigung der brigen Lnder Europas sowie
der angrenzenden Meeresgebiete. In: Rabenhorst L.,
Kryptogamen-Flora von Deutchland, sterreich
und der Schweiz, 7. Leipzig, Akademische
Verlagsgesellschaft.
Hustedt, F., 1935. Die fossile Diatomeenflora in den
Ablagerungen des Tobasees auf Sumatra. (Fossil
Diatoms Deposits in the Toba sea of Sumatra). Archiv
fr Hydrobiol. Suppl. Bd. XIV Tropische
Binnengewasser. VI: 143-192, Taf 5 (5 pl.).
Hustedt, F., 1937a. Systematische und kologische
Untersuchungen iiber die Diatomeen-flora von Java,
Bali und Sumatra. Archiv Fur Hydrobiologie. Suppl.
XV(I): 131-177, pl. 9-12.
Hustedt, F., 1937b. Systematische und kologische
Untersuchungen uber die Diatomeen-Flora von Java,
Bali und Sumatra. Archiv Fur Hydrobiologie. Suppl.
XV(2): 187-295, pl. 13-20.
Hustedt, F., 1942a. Ssswasser-Diatomeen des
indomalayischen Archipels und der Hawaii-Inseln.
Internationale Revue der gesamten Hydrobiologie und
Hydrographie, 42(1/3):1-252, 443 text figs.

Hustedt, F., 1942b. Beitrage zur Algenflora von Bremen.

V. Die Diatomeenflora einiger Sumpfwiesen bei
Bremen. Abhandl. Naturwissenschaftl. Verein zu
Bremen. 32,: 184-221.
Hustedt, F., 1955. Marine littoral diatoms of Beaufort,
North Carolina. Durham, North Carolina, Duke Univ.
Press: 67p., 16 pl.
Hustedt, F., 1957. Die Diatomeenflora des Flusssystems
der Weser im Gebiet der Hansestadt Bremen. Abh.
herausgegeben vom Naturwiss. Ver. zu Brem. 3(4):
181-440, 1 pl.
Janisch, C., 1888. Diatomaceen gesammelt auf der
Forschungs Reise der Gazelle in d. Jahren 18741876 bearbeitet durch C. Janisch. (Planches et
Phototypie d aprs les photographies de C. Janisch).
16 Lichtdruck-Tafeln (325 Figuren) mit TafelErklrungen von 16 Seiten.
Kitton, F., 1889. New Species of Navicula (N.
venustissima). Jour. Royal Microsc. Soc., p. 101.
Knig, D. 1959. Diatomeen der Bucht von Arcachon
(Dep. Gironde Frankreich). Zeitschrift der Deutschen
Geologischen Gesellschaft. 111: 33-61.
Krasske, G., 1923. Die Diatomeen des Casseler
Beckens und seiner Randgebirge, nebst einigen
wichtigen funden aus Niederhessen. Botanisches
Archiv. 3(4): 185-209.
Krasske, G., 1925. Die BacillariaceenVegetation
Niederhessens. Abhandalungen und Bericht. (56 des)
Veriens fr Naturkunde, zu casul, 84-849 Vereinsjahr
1919-1926, S. 1-119, 2 Taf.
Krasske, G., 1929. Beitrge zur Diatomeenflora
Sachsens. Botanisches Archiv. 27 (3/4): 348-380, 25
fig.
Krasske, G., 1932. Diatomeen aus dem Oberpliocn
von Willershausen. Archiv. fur Hydrobiol. 24(3): 430448, Taf. 16.
Ktzing, F. T., 1833. Synopsis Diatomacearum oder
Versuch einer systematischer Zusammenstellung der

29

Martinez-Goss

Diatomeen. Linnaea. 8: 529-620, Taf. 13-19. Berlin

(Sonderabdruck, s. 1-92, 1834).
Ktzing, F.T., 1844. Die Kieselschaligen Bacillarien
oder Diatomeen. Nordhausen, W. Kohne. 152p.,
30 pl.
Ktzing, F.T., 1849. Species Algarum. Lipsiae, F.A.
Brockhaus., 922s.
Leuduger-Fortmorel, G., 1879. Catalogue des Diatomees
de lile Ceylon. Mmoires de la Socit d Emulation
des Ctes du Nord.- Saint-Brieuc, Libraire Francisque
Guyon, Imprimeur, 73p., 9 pl.
Leuduger-Fortmorel, G., 1892. Diatomees de la
Malaisie. Annales du Jardin Botanique de
Buitenzorg. 11: 1-60, pl. 1-7.
Lewis, F.W., 1861. Notes on new and rarer species of
Diatomaceae of the United States Seaboard. Proc.
Acad. Nat. Sci. of Phila. 13:61-71, 2 pls. Also, Quart.
J. Microsc. Sci. 2( new series): 155.
Lewis, F.W., 1865. On extreme and exceptional
variations of diatoms in some white Mountain localities,
etc. Proc. Acad. Nat. Sci. Phila. 7-18, pl. 1-2.
Mann, A., 1907. Report on the diatoms of the Albatross
Voyages in the Pacific Ocean, 1888-1904. Contributions
from the United States National Herbarium, vol. 10,
part 5, pp. 221-419, 11 pl.
Mann, A., 1925. Marine Diatoms of the Philippine
Islands. Smithsonian Institution, United States
National Museum Bull. #100: Vol. 6, part 1, 182p.,
39 pl.
Martinez-Goss, M.R., 1995. A Checklist of Nitzschia
and Tryblionella (class Bacillariophyceae) of the
Philippines. Phil. J. Sci. 124(1):75-99.

Meister, F.R., 1934 (1935). Seltene und nene

Kieselalgen. Bericht der Schweizerische Botanische
Gessellschaft.44: 87-108.
Meister, F.R., 1937. Seltine und nene Kieselalgen. II.
Berichte der Schweiserischen Botanischen
Gessellschaft. 47: 258-276, Tafeln 3-13.
Mereschkowsky, C., 1902, Note sur quelques
Diatomes de la Mer Noire. Journal de Botanique
(Morot, Ed., Paris). 16: 319-324, 358-360, 416-430.
Muller, O., 1903-1910. Bacillariaceen aus dem
Nyassal and und einigen benachbarten Gebieten.
(Englers) botanische Jahrbuccher fur Systematik,
Pflanzengeschichte (Bd. 34, Heft1, 2 Taf.,3 Textfig.,
1903); II Folge, Discoideae-coscinodisceae.
Discoideae-Eupodisceae (Bd. 34, Heft 2, S. 255301, 4 Taft., 1904); III Folge, Naviculoideae,
Gomphoneminae, Nitzschioideae (Bd. 36, Heft ,
S. 137-206, 2 Taf., 1905); IV Folge, NaviculoideaeFragilarieae-Fragilarineae, FragilarioideaeFragilarieae-Eunotiinae (Bd. 45, Heft 1, S. 69-122,
Taf., 1910).
Norman, G., 1861. On some undescribed species of
Diatomaceae. Trans. Microsc. Soc. 9(N.S.): 5-9,
pl. 2.
OMeara, E., 1872. On some peculiar forms of
Navicula from the Sulu Archipelago. Quart. J.
Microsc. Sci. 12 (new series): 283-287, 13 pl.
strup, E., 1901. Freshwater-diatoms from the Faeroes.
In: Botany of the Faeroes, Part 1. Copenhagen, Det
Nordiske Forlar: p. 260-290.
strup, E., 1913. Diatomaceae ex Insulis Danicis
Indiae Occidentalis. Dansk Botanisk Ankiv. 1(1): 1-39,
pl. 1.

Meister, F.R., 1912. Die kieselalgen der Schweiz. In Beitrage

zur Kryptogamenflora der Schweiz. Band IV, Heft I. Bern,
Druck und Verlag von J.K. Wyss.: 254 s. 48 Taf.

strup, E., 1918. Freshwater Diatoms from Iceland,

In: Rosevinge, L.K. & E. Warming, Botany of Iceland,
vol. 2, part 1, no. 5, p. 1-96, 5 pls. (Arbejdan fraden
Botoniske i Kbenhavn, Nr. 86).

Meister, F.R., 1932. Keilalgen aus Asien. Berlin, Verlag

von Gebriider Borntraeger: 56 S., 19 Taf.

strup, E., 1918 (1920). Freshwater Diatoms from

Iceland. The Botany of Iceland by L. K. Rosenvinge

30

A checklist of Navicula

and E. Warming. Vol. 2, part 1, no. 5, p. 1-90, 5 pls.

Copenhagen.

Round, F.E., R.M. Crawford & D.G. Mann, 1990. The

Diatoms, biology and morphology of the genera.
Cambridge, University Press, 747p.

Pantastico, J. B. 1977. Taxonomy of the freshwater

algae of the Laguna de Bay and vicinity. Bicutan, Tagig,
Metro Manila, NRCP Bull. 61: 251p., 17 pl.

Schmidt, A., et al. 1874. Atlas der Diatomaceen-Kunde.

Leipzig, O.R. Reisland. 1-472 pl.

Pantocsek, J., 1886. Beitrge zur Kenntniss der fossilen

Baccilarien Ungarns. Teil I. Marine Bacillarien, 74s.,
30 Taf.

Schumann, J., 1862 (1863). Preussiche Diatomeen,

Schriften der koniglichen physikalisch-okonomischen
Gessellschaft zu Konigsberg. 3: 166-192, Taf. 8-9.

Patrick, R., 1959. New species and nomenclatural

changes in the genus Navicula (Bacillariophyceae).
Proc. Acad. Nat. Sci. Phila. 3:91-108, 2 pl.

Schumann, J., 1867. Die Diatomeen der Hohen Tatra.

Verhandlungaen kaiserlich-konigliche zoologischbotanische Gasellschaft in Wien.17: 1-102, 4 Taf.

Patrick, R. & C. W. Reimer, 1966. The Diatoms of the

United States exclusive of Alaska and Hawaii. vol. 1,
Monograph of the Acad. Nat. Sci. Phila. No.
13.Philadelphia, Acad. Nat. Sci. Phila. 688p., 64 pl.

Skvortzow, B.W., 1931. Pelagic diatoms of Korean Strait

of the Sea of Japan. Phil. J. Sci. 46(1):95-122, 10 pl.

Peragallo, H., 1888. Diatomes du Midi de la France.

Diatomes de la Baie de Villefranche (Alpes maritimes).
Bulletin Socit d Histoire Naturelle de Toulouse.
22: 13-100, pl. 1-6. (Separate pagination also.)
Peragallo, H. 1891. Monographic du genre Pleurosigma
et des genres, allies. Le Diatomiste. 1(4):1-35.
Peragallo, H. & M. Peragallo, 1897-1908. Diatomes
Marines de France et des Districts Maritimes Voisins.
Micrographe-diteur, Grez-sur-Loing (Siet-et-M.).
Atlas 137 pls. Text 491p. + 48p.
Petersen, J., 1928. Algefloraen I nogle Jordprover fra
Island. Dansk Botanisk Arkiv. 5(9): 1-23, 3 Fig.
Podzorski, A. C. & H. Hakansson, 1987. Freshwater
and marine diatoms from Palawan (a Philippine island).
Bibliotheca Diatomologica. Bd. 13, Berlin, J. Cramer,
134p., 55 pl.

Skvortzow, B.W., 1932. Marine littoral Diatoms from

environs of Vladivostok. Phil. J. Sci. 47(1): 129-150, 6
pls.
Skvortzow, B.W., 1936. Diatoms from Kizaki Lake,
Honshu Island, Nippon. Phil. J. Sci. 61(1):9-73, 16 pl.
Skvortzow, B.W., 1937. Diatoms from the Philippines,
I. Diatoms from the drinking water Balara, Rizal
Province. Phil. J. Sci. 64(3):287-298, 2pl.
Skvortzow, B.W. & C.I. Meyer, 1928. A contribution
to the Diatoms of Baikal Lake. Proc.of the Sungaree
River Biological Station, 1(5): 1-55, 2 pls.
Smith, W., 1853. A synopsis of the British Diatomaceae
with remarks on their structure, functions and
distribution and instructions for collecting and preserving
specimens. Vol. 1. London, John van Voorst: 89p., 31
pl.
Smith, W., 1856. A synopsis of the British Diatomaceae.
Vol. 2. London, John van Voorst: 107p., 62 pl.

Pritchard, A., 1861. A history of infusoria, living and

fossil. VI edition. Revised and enlarged by J. T. Arlidge,
W. Archer, J. Ralfs, W.C. Williamson & A. Pritchard.
London, Whittaker & Co., 968p., 40pl.

Tempere, J., & H. Peragallo, 1914. Diatomees du

Monde Entier, Edition 2; 30 fasc. Arcachon, Grezsur-Loing (S.-et-M.). Eascicule 1, 29-30, p. 449-480.

Prowse, G.A., 1962. Diatoms of Malayan freshwaters.

Singapore Gardens Bull. 19: 1-104, 2 fig., 24 pls.

Truan y Luard, A. & O.N. Witt, 1888. Diatomaceen

der Polycystinenkreide von Jermie in Haiti, Westindien,

31

Martinez-Goss

24S., 7 Taf. Friedlnder et Sohn, Berlin. Journal de

Micrographie. t 12, fasc. 5, 162 (Review.)
Van Heurck, H., 1880-1885. Synopsis des Diatomes
de Belgique. Atlas. Anvers, Edite Par Lauteur. Ducaju
et Cie. & J.F. Dieltjens, 132 pl.
Van Heurck, H., 1896. A treatise on the Diatomaceae.
Tr. by W.E. Baxter. London, William Wesley & Son:
558p., 35 pl.
Van Heurck, H., 1909. Expedition Antarctique Belge,
Resultate du Voyage du S. Y. Belgica en 1897-18981899., Botanique, Diatomees. Anvers, Imprm. J. E.
Buschmann: 129 p., 13 pl.
VanLandingham, S.L., 1975. Navicula Part V.
Catalogue of the fossil and recent genera and species
of diatoms and their synonyms. Germany, J. Cramer,
pp. 2386-2963.
Villadolid, D.V., 1957. The food habits of fry and
fingerlings of milkfish (Chanos chanos Forsk). Ar. J.
Agric., 4: 1-23.
Wolle, F., 1890. Diatomaceae of North America.
Bethlehem, Pa., Comenius Press: 47p., 112 pl.
Woltereck, R., 1941. Die seen und Inseln der
Wallacea Zwischen-region und ihre endemische
Tierwelt. Internat. Rev. der Ges. Hydrobiol. und
Hydrograph. 41:37-176.
Zanon, V., 1949. Diatomee di Buenos Aires (Argentina).
Atti della Accademia Nazionalc dei Lincei, Memoire
(Classe di Scienze fisiche, matematische e naturali) Ser.
8, sezione 3, t.2, (Botanica, zoologica fisiologica,
patologia), p.1-151-, 2 tav.

32