l*l 331:!

y"

Checklist of

of Canada and Alaska

 Checklist of

BEETLES
of Canada and Alaska

Edited by

Yves Bousquet
Biosystematics Research Centre
Ottawa, Ontario

Research Branch
Agriculture Canada
Publication 186UE

t99l
oMinister of Supply anrl Services Canada 1991

Available in Canada through

Assoeiated Bookstores
and other borksellers
or by mail fr,rm

Canada Com-nunication Group Publishing

Ottawa, Canada KlA0S9
-

Cat. no. A43-1861/1991E

ISBN 0-660-13?67-4

Canadian Cataloguing in Publication Data

Checklist c f beetles of Canada and Alaska
(Publication ; 18'61/E)

Includes bibliographical references.

Cat. no. A43. 1861i1991E
ISBN 0-660- r3767-4

1. Beetles-(lanada-Nomenclature. 2. Beetles-
Alaska-Nomr:nclature. I. Bousquet, Yves. II.
Canada. Agriculture Canada. Research Branch.
III. Series: .?ublication (Canada. Agriculture
Canada). English ; 1861/E.

QI581.C4 1991 595.76 C91-099101-4

Stafr Editor
Frances Smith
 CONTENTS
List ofcontributors v SuperfamilyScarabaeoidea 143
Acknowledgments v Family Lucanidae (J. McNamara) 143
Summary vi Family Passalidae (J. McNamara) 145
R6sum6 vi Family Scarabaeidae (J. McNamara) 145
Synoptic table 1 SuperfamilyByrrhoidea 158
Introduction 4 Family Byrrhidae (P.J. Johnson) 158
Format 4 SuperfamilyBuprestoidea 160
Subsequent revisions ofthe checklist 5 Family Buprestidae (A. Davies) 160
Abbreviations and symbols 5
SuperfamilyDryopoidea 169
ORDERCOLEOPTERA 6 Family Ptilodactylidae (L. LeSage) 169
SuborderArchostemata 6 Family Heteroceridae (L. LeSage) 169
Family Cupedidae (J.M. Campbell) 6 Family Limnichidae (L. LeSage) 171
Family Micromalthidae (J.M. Campbell) 6 Family Dryopidae (L. LeSage) 771
Family Elmidae (L. LeSage) 172
SuborderAdephaga 7 Family Psephenidae (L. LeSage) 174
Family Rhysodidae (Y. Bousquet) 7
Family Trachypachidae (Y. Bousquet) 7 SuperfamilyElateroidea L74
Family Carabidae (Y. Bousquet) 8 Family Artematopidae (Y. Bousquet) 174
Family Haliplidae (R.E. Roughley) 60 Family Elateridae (Y. Bousquet) 175
Family Amphizoidae (Y. Bousquet) 61 Family Throscidae (Y. Bousquet) 186
Family Noteridae (R.E. Roughley) 61 Family Eucnemidae (Y. Bousquet) 186
Family Dy'tiscidae (D.J. Larson and R.E. Superfamily Cantharoidea 188
Roughley) 62 Family Lycidae (J. McNamara) 188
Family Gyrinidae (R.E. Roughley) 72 Family Phengodidae (J. McNamara) 190
Suborder Polyphaga 74 Family Lampyridae (J. McNamara) 190
Family Cantharidae (J. McNamara) 192
SuperfamilyStaphylinoidea 74
Family Hydraenidae (R.E. Roughley) 74 SuperfamilyDermestoidea 195
Family Ptiliidae (J.M. Campbell) 75 Family Derodontidae (D.E. Bright) 195
Family Limulodidae (J.M. Campbell) 76 Family Nosodendridae (J.M. Campbell) 196
Family Agyrtidae (J.M. Campbell) 77 Family Dermestidae (Y. Bousquet) 196
Family Leiodidae (S.B. Peck) 77 SuperfamilyBostrichoidea 199
Family Scydmaenidae (A. Davies) 82 Family Bostrichidae (J. McNamara) 199
Family Micropeplidae (J.M. Campbell) 84 Family Lyctidae (J. McNamara) 20L
Family Silphidae (J.M. Campbell) 84 Family Anobiidae (J. McNamara) 201
Family Staphylinidae (J.M. Campbell and A. Family Ptinidae (Y. Bousquet) 205
Davies) 86 Superfamily Lymexyloidea 206
Family Scaphidiidae (J.M. Campbell) 124 Family Lymexylidae (J.M. Campbell) 206
Family Pselaphidae (A. Davies) 125 SuperfamilyCleroidea 206
SuperfamilyHydrophiloidea 130 Family Trogossitidae (Y. Bousquet) 206
Family Hydrophilidae (R.E. Roughley) 130 Family Cleridae (J. McNamara) 208
Family Georyssidae (R.E. Roughley) 135 Family Melyridae (D.E. Bright) 2LI
Family Sphaeritidae (R.E. Roughley) 135 Superfamily Cucujoidea 213
Family Histeridae (A. Davies) 135 Family Sphindidae (J.M. Campbell) 213
SuperfamilyEucinetoidea I4I Family Nitidulidae (J. McNamara) 2I4
Family Eucinetidae (J.M. Campbell) I4I Family Rhizophagidae (Y. Bousquet) 2I8
Family Clambidae (J.M. Campbell) I4l Family Cucujidae (Y. Bousquet) 219
Family Scirtidae (J.M. Campbell) 142 Family Cryptophagidae (Y. Bousquet) 22I
SuperfamilyDascilloidea 143 Family Languriidae (J.M. Campbell) 223
Family Dascillidae (J.M. Campbell) 143 Family Erotylidae (J.M. Campbell) 225
Family Rhipiceridae (J.M. Campbell) I43 Family Phalacridae (J.M. Campbell) 226

Ill
 Family Cerylonidae (J.M. Campbell) 227 Family Anthicidae (Y. Bousquet) 27t
Family Corylophidae (J.M. Campbell) 228 Family Pedilidae (Y. Bousquet) 274
Family Coccinellidae (J. McNamara) 229 Family Aderidae (Y. Bousquet) 275
Family Endomychidae (J.M. Campbell) 287 Family Scraptiidae (L. LeSage) 275
Family Lathridiidae (Y. Bousquet) 239 SuperfamilyChrysomeloidea 277
Family Biphyllidae (J.M. Campbell) 247 Family Cerambycidae (J. McNamara) 277
Family Byturidae (J.M. Campbell) 24I Family Bruchidae (J. McNamara) 300
SuperfamilyTenebrionoidea 242 Family Chrysomelidae (L. LeSage) 3C,1
Family Mycetophagidae (Y. Bousquet) 242 SuperfamilyCurculionoidea 323
Family Tetratomidae (L. LeSage) 243 Family Nemonychidae (J. McNamara) 323
Family Ciidae (J. McNamara) 244 Family Anthribidae (J. McNamara) 324
Family Melandryidae (L. LeSage) 245 Family Ithyceridae (J. McNamara) 32:,5
Family Mordellidae (J. McNamara) 248 Family Attelabidae (J. McNamara) 3tl6
Family Rhipiphoridae (J.M. Campbell) 250 Family Rhynchitidae (J. McNamara) 326
Family Colydiidae (Y. Bousquet) 251 Family Brentidae (J. McNamara) 327
Family Zopheridae (J.M. Campbell) 252 Family Apionidae (J. McNamara) 328
Family Tenebrionidae (Y. Bousquet and J.M. Family Curculionidae (J. McNamara) 329
Campbell) 253 Family Scolytidae (J. McNamara) 35;
Familr' Lagriidae (J.M. Campbell) 261 Family Platypodidae (J. McNamara) :]65
Famils Cephaioidae (J.M. Campbell) 262
Famil r Meloidae (J.M. Campbell) 269 ORDERSTREPSIPTERA 366
Famil r Oedemeridae (J.M. Campbell) 266 SuborderStylopidia 366
Famil r' Prostomidae (J.M. Campbell) 267 Family Corioxenidae (S.B. Peck) 366
Famil Synchroidae (L. LeSage) 267 Family Halictophagidae (S.B. Peck) 336
Familr'z Mycteridae (J.M. Campbell) 267 Family Stylopidae (S.B. Peck) 367
Famil,r Boridae (J.M. Campbell) 268
Famil r Pythidae (J.M. Campbell) 268 References 368
Famil r Pyrochroidae (J.M. Campbell) 269 Index 397
Famil,r Salpingidae (J.M. Campbell) 270

iv
 LIST OF CONTRIBUTORS
Y. Bousquet, Biosystematics Research Centre, D.J. Larson, Department of Biology, Memorial
Agriculture Canada, Ottawa, Ontario K1A 0C6 University of Newfoundland, St. John's,
D.E. Bright, Biosystematics Research Centre, Newfoundland AlB 3X9
Agriculture Canada, Ottawa, Ontario K1A 0C6 L. LeSage, Biosystematics Research Centre,
J.M. Campbell, Biosystematics Research Centre, Agriculture Canada, Ottawa, Ontario K1A 0C6
Agriculture Canada, Ottawa, Ontario K1A 0C6 J. McNamara, Biosystematics Research Centre,
A. Davies, Biosystematics Research Centre, Agriculture Canada, Ottawa, Ontario KlA 0C6
Agriculture Canada, Ottawa, Ontario K1A 0C6 S.B. Peck, Department of Biology, Carleton
P.J. Johnson, Department of Entomology, University, Ottawa, Ontario KlS 586
University of Wisconsin, Madison, Wisconsin R.E. Roughley, Department of Entomology,
53706 University of Manitoba, Winnipeg, Manitoba
R3T 2N2

ACKNOWLEDGMENTS
We wish to thank the following persons for Moldenke (Clytrinae); H.G. Nelson (Dryopoidea);
reviewing sections of the checklist: R.L. Aalbu A.F. Newton Jr. (Leiodidae, Staphylinidae); S.
(Tenebrionidae); R.S. Anderson (Silphidae, Oygur (Gyrinidae); J. Pakaluk (Corylophidae); J.D.
Curculionidae); F.G. Andrews (Lathridiidae); LS. Pinto (Meloidae); D. Pollock (Mycteridae, Boridae,
Askevold (Chrysomelidae); F.M. Atton (Gyrinidae); Pythidae, Salpingidae); V. Puthz (Steninae); E.G.
E.U. Balsbaugh (Chrysomelidae); R. Baranowski Riley (Cassidinae); W.T. Schultz (Eumolpinae); T.N.
(Leiodidae); W.F. Barr (Cleridae); E.C. Becker Seeno (Chrysomelidae); R.B. Selander (Meloidae);
(Elateroidea); H.P. Brown (Dryopoidea); D. S.A. Slipinski (Cerylonidae); C.L. Staines
Chandler (Anthicidae, Pselaphidae, Melandryidae, (Hispinae); W.E. Steiner Jr. (Phalacridae); J.B.
Tetratomidae, Synchroidae, Scraptiidae); J. Cicero Stribling (Ptilodactylidae); M. Thayer
(Lampyridae); W. Connell (Nitidulidae); M. (Staphylinidae); M.C. Thomas (Cucujidae,
Daccordi (Chrysomelinae); J.G. Edwards Prostomidae); C.A. Triplehorn (Tenebrionidae); R.E.
(Synetinae); D.G. Furth (Chrysomelidae); R.E. White (Anobiidae, Chrysomelidae); D.P. Wooldridge
Gordon (Coccinellidae): L.H. Herman (Limnichidae); D.K. Young (Scirtidae,
(Staphylinidae); A.T. Howden (Curculionidae); H. Pyrochroidae); P. Zwick (Leiodidae).
Howden (Scarabaeidae); C. Johnson (Ptiliidae); J.B. Thanks are also extended to B.F. Carr, J.L.
Karren (Chlamisinae); J.M. Kingsolver (Bruchidae); Carr, and D.W. Langor, for providing several
J.L. Krysan (Chrysomelidae); S. Laplante (Cleridae, provincial records; to D. Beauchamp, B. Walter, L.
Cerambycidae); J.F. Lawrence (Melandryidae, Robillard, and R. Hutchinson, for typing the check-
Tetratomidae, Synchroidae, Scraptiidae); I. Lobl list; and to A. Smetana, for checking the termina-
(Scaphiidae); W.V. Miller (Heteroceridae); A.R. tion of all species-group names.
 SUMMARY
All species and subspecies of beetles known to in each province
presence or absence of the taxa
occur in C.anada and Alaska are listed along with and territory, in Labrador, and in Alaska.
their synonyms; these names are arranged alpha- Introduced and holarctic species are highlighted.
betically t,nder the respective genus-group names. References to pertinent identifrcation keys are indi-
Valid genus-group and family-group names are cated under the corresponding supraspecific taxon.
listed in p aylogenetic order. No new taxa, combina- The work also includes a brief introduction for
tions, or synonyms are proposed. Species distribu- every family treated and an index to all supraspe-
tions are presented in tabular form, indicating the cifrc names.

RESUME
L'ouvrage r6pertorie toutes les espdces et sous- tabulaire, indiquant la pr6sence ou l'absl:nce des
espdces dt,: Col6optdres connues du Canada et de taxa dans chaque province et territoire, le
l'Alaska arnsi que leurs synonymes; ces noms sont Labrador et l'Alaska. On met en relief les espbces
6num6r6s par ordre alphab6tique d la suite du nom introduites ou holarctiques et on mentionne, sous
du groupe-genre correspondant. Les noms valides le taxon suprasp6cifique correspondra.nt, les
des group':-genre et groupe-famille sont pr6sent6s r6f6rences contenant des tableaux de d61;ermina-
par ordre phylog6n6tique. Cette liste ne propose tion utiles. L'ouvrage comprend aussi une courte
aucun nouveau taxon, combinaison ou synonyme. introduction sur chaque famille consid6r'6e ainsi
La r6partibion des espdces est pr6sent6e sous forme qu'un index des noms suprasp6cifrques.

vi
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 INTRODUCTION
Beetles represent the most diverse group of No comprehensive checklist of Coleoptera is
living organisms on earth. More than 250 000 available for Canada and Alaska. Leng's catalogue
species have been described, and the actual number (1920) and its supplements (Leng and Mutchler
may easily exceed a million. Given such staggering 1927, 1933; Blackwelder 1939a; Blackwelder and
frgures, it is not surprising that Coleoptera can be Blackwelder 1948) as well as the world-based
found neally everywhere, from the tropics to the Coleopterorum Catalogus list most of the species
Arctic and from caves to the summit of high moun- that occur in Canada and Alaska. However, these
tains. Coleoptera species are particularly abundant publications are out of date for many groups, and
on vegetation, inside rotten wood, under the bark of their distribution data are incomplete. Beaulne
dead trees, and in soil, ground litter, fungi, dung, (1913-1929)started a checklist of the beetles of
carrion, rodent burrows, and nests. Several groups Canada but never completed it, and as pointed out
are adapted to life in fresh or brackish water. by Lindroth (1966) for the ground beetles, many of
Although some dytiscids can venture into salt water Beaulne's distributional records are inaccurate.
and some species do complete all stages of their life This checklist is intended to provide the correct
cycle in the intertidal zone where they are periodi- names of the beetles that occur in Canada and
cally immersed, no beetles have become completely Alaska, give a brief survey of the distribution of the
adapted to marine life. Beetles are important to taxa in the area covered, and lead the user's to the
humans because many species are beneficial in the proper publications for the identification of the
biological control of pest arthropods and weeds or taxa the checklist of Strepsiptera has been added
are major pests of stored products, fabrics, forests, at the end simply because the group has often been
wood prodructs, and crops. included in the Coleoptera.

FORMAT
With some exceptionsreflecting the author's J.L. LeConte, G. Horn, and J.C. Fabriciusr respec-
personal perspective, the concept of the families tively. An asterisk [*] placed after a species-group
used in this checklist follows that of Lawrence name indicates that the taxon is Holarctic, a dagger
(1982a). N,najor changes concern the trachypachids, ttl that it is introduced in North Americra, and a
limulodids, scaphidiids, lyctids, lagriids, pedilids, double dagger t{l that it is probably introduced in
rhynchitids, scolytids, and platypodids, which are North America.
treated as distinct families, and the leptinids, AII synonyms based on material collected in
trogids, and geotrupids, which are included respec- North America north of Mexico should be included
tively within the Leiodidae and the Scarabaeidae and are listed in italics. Synonyms of Hola:nctic and
for the last two groups. The superfamily concepts introduced species based on material frorri outside
and the listing of the families follow Lawrence North America have been included occasionally,
(1982a). particularly when the name has been in use in
The classification and information for each North American literature. Misidentifications,
family are the responsibility of the respective spelling errors, and invalid emendations are not
authors. No new taxa, combinations, or synonyms llited. The ending of all species-group names has
are proporied. Variety and form names published been checked to conform with the gender of their
before 1961 are given subspecifrc rank unless they respective genera.
were deemed to be synonyms or their status has Valid genus-group names are in capital letters,
been changed (article +5 (g) of the International their synonyms in italics. All synonyms crf genus-
Code of Zoological Nomenclature, 3rd ed.). group names whose type species are indigengu-s to
Varieties and forms published after 1960 are North America north of Mexico should be included.
omitted. New distributional records are listed but Some of the synonyms whose type species are not
not identified as such. indigenous to the continent are included also when
The arLthors of species-group names are listed in the name has been used in North American litera-
parentheses when the original combination differs ture.
from the one used in the checklist. To avoid confu- References to keys are indicated under the cor-
sion, initials are included for several authors, such responding supraspecific taxon. It should be noted
as C.R. Sahlberg, R.F. Sahlberg. J.R. Sahlberg, J. that only references including keys useful for the
MiiIIer, O.F. Miiller, W. Horn, J.E. LeConte, and O. determination of all or most Canadian taxa of the
Fabricius. LeConte, Horn, and Fabricius stand for group treated are considered. Keys published
before the 1930s are usually out of date, and most of from other collections have been included. Doubtful
them are therefore omitted. records from the literature that apparently are
An abbreviation of two letters is used to indi- based on misidentifications or mislabeled specimens
cate Canadian provinces and Alaskan records (see are rejected.
section entitled "Abbreviations and symbols"). A question mark (?) inserted in a geographic
Labrador is considered to be a distinct geographic column indicates that the taxon was recorded in the
entity. The distributional citation in the checklist is area, but the record is questionable. The absence of
from west to east. a geographic entry after a species-group name indi-
The distributional records are based on pub- cates that the taxon was recorded in Canada. but
lished records and on specimens in the Canadian the province was not specified.
National Collection. For some families. records

SUBSEQUENT REVISIONS OF THE CHECKLIST

This publication is set as a data base that can sions to their attention. We would also welcome
be easily updated to maintain an accurate summary receiving specimens that represent new provincial
of our current knowledge of the beetles of Canada or Alaskan records. Specialists and other users
and Alaska. The authors intend to maintain a cor- may send material to the Biosystematics Research
rected version of the checklist and to republish it Centre for confirmation, and with the sender's per-
whenever we have accumulated enough changes. mission, for incorporation as voucher specimens in
The authors would appreciate the assistance of all the Canadian National Collection.
users of this checklist in calling errors and omis-

AB B REV *TI
r.rl' O N S AND SYMB O L S
AB Alberta NT NorthwestTerritories
AK Alaska ON Ontario
BC British Columbia PE Prince Edward Island
LB Labrador PQ Quebec
MB Manitoba SK Saskatchewan
NB New Brunswick YK Yukon Territorv
NF Newfoundland H Holarctic
NS Nova Scotia I Introduced in North America

* Holarctic f Probably introduced in North America

t Introduced in North America $ No subgeneric name available
Micromalthidae

Order Coleoptera
Suborder Archostemata
FamilyCUPEDIDAE reticulatedbeetles
J.M. Campbell

Three species of Cupedidae occur in Canada. Neboiss (1984) the species of Cupes. Ponornarenko
This small, primitive family of beetles contains only (1969) discussed the fossil origins of the Coleoptera
30 extant species around the world, but it is well with particular reference to the srrborder
represented by fossils dating back to the early Archostemata and the family Cupedidae, rerviewing
Triassic. l,ittle is known about the biology of the the fossil and extant species of the family. Atkins
species-the larvae apparently bore into wood, and (1979) cataloged the North American taxa of the
the adults iue collected from foliage or under bark. family.
The farnily Cupedidae is well known taxonomi-
cally. Atk.ns (1963) revised the world species, and BC (1); oN (2);PQ (1)

Subfamily CUPEDINAE
Tribe Priacmini
Genus PRIAC MA LeConte
P. serrata (LeConte) BC

Tribe Cupedini
Genus CUPEII Fabricius
C. capitatus tr abricius ON

Genus TENO T{ERGA Neboiss

T. concolor (Si estwood) ON PQ
cinerea (flay)
oculata (Oasey)
t r iI ineat c; (Melsheimer)

Family MICROMAL'THIDAE telephone-pole beetles

J.M. Campbell

This fz: mily contains only one species, which known to occur in Coleoptera. The larvae are pae-
was descri ced from the eastern United States but dogenetic, producing either living larvae or a single
has subserluently become established in western egg that develops through a different larval type to
Canada, C uba, Brazil, Hawaii, Hong Kong, and produce an adult male (Pringle 1938).
South Afrir:a. The larvae are wood borers and are Atkins (1982) cataloged the species for North
found in decaying oak logs and in moist, creosote- America.
treated timbers and poles. The life cycle of
Micromalthus debilis is one of the most complex BC (1)

Genus MICROMALTHUS LeConte

M. debilis Le(jonte BC
 Trachvoachidae

Suborder Adephaga
Family RIIYSODIDAE wrinkled bark beetles
Y. Bousquet

The family Rhysodidae includes only two and larvae live under the bark of dead trees and in
speciesin Canada. The adults are cylindrical, elon- rotting wood.
gate beetles resembling in habitus some species of Bell (1970) revised the North American species.
the families Colydiidae or Cucujidae. Both adults eC (1);ON (1)

Genus OMOGLYMMIUS Ganglbauer

Key to species: Bell and Bell (1982)
(Subgenus BOREOGLYMMIUS Bell & Bell)
O. americanus (Castelnau) - ON
orolas (Newman)
e ra r at u s (Audinet-Serville)

Genus CLINIDIUM Kirby

Key to North American species: Bell and Bell (1975)
(Subgenus ARCTOCLINIDIUM Bell)
C. calcaratum LeConte - BC

Family TRAC IIYPAC HIDAE

Y. Bousquet

This family includes two genera: Trachypachus, decades, however, some authors have highlighted
which has three species in western North America many character states that suggest that
and one species in Eurasia, and Systolosorno Solier, Trachypachus and Sysfolosoma are more closely
which has two species in Chile. Members of related to Hydradephaga than to Carabidae (see
Trachypachus are usually found on dry, more or Kavanaugh (1986) for a different point of view).
Iess shaded soil, often under dead leaves. Nothing Lindroth (1961) has redescribed the two species
is known about their feeding habits. that inhabit Canada.
This group has been included within the
Carabidae for a long time. During the last two aK (1);YK (1); NT (1); BC (2); aB (1); SK (1)

Genus TRACHYPACHUS Motschulsky

T. holmbergi Mannerheim AK YK .\T BC AB SK
inermis Motschulsky
oregonus Casey
specularis Casey
T. gibbsii LeConte _BC
alticola Casey
c alifor nicu s Motschulsky
Carabidae

Family CARABIDAE ground beetles

Y. Bousquet

With 9,t6 species and subspecies in Canada and in the family. In recent years, some authc,rs haYe
Alaska, thr,: family Carabidae is an important ele- included the rhysodids within Carabidae, but based
ment of thr: Coleoptera fauna in the northern part on larval characters I prefer to exclude thenL; on the
of North A:nerica. In Canada and Alaska members other hand, some others consider the cicinclelids to
of these species occur in most terrestrial habitats as be a distinct family, a view with which I do not
well as in .marshes and bogs; a few species live on agree. Thanks to the work of Wallis (1961) lor tiger
trees. Adults of most species are carnivorous or beetles and Lindroth (1961-1969), the carabid fauna
omnivorous,; only those of a few groups seem to be of Canada and Alaska is well known taxonomically.
primarily phytophagous; larvae of most species are Bell (1990) recently published a key to the North
probably ca.rnivorous, but those of a few groups are American genera.
known to be phytophagous or parasitic.
The Carabidae of North America are here AK(237); YK (164); NT (207); BC (a83); AB (386);
grouped in 36 tribes. The classification used is basi- SK (314); MB (350); ON (503); PQ (451); NB (2a3);
cally that of Lindroth (1961-1969), with a few modi- NS (24a); PE (137); LB (.94); NF (175) ' II (84); I
fications re,flecting recent taxonomic developments (46)

Tribe Cicindelini
Genus OMUS Eschscholtz
Leptontut Casey
Megomus Casey
O. audouini audouini Reiche BC
aequicorr,.is Casey
audens Casey
borealis ()asey
delicatuhn Casey
distans Casey
oregonentiis Casey
paruulus Casey
rugipenn,.s Casey
tacomae ()asey
thoracicus Casey
uandykei W. Horn
O. dejeani Rei:he BC
foueatus l)asey
robustus Casey

Genus CICIN.)ELA Linn6

(Subgenus CICINDELA s. str.)
(Pachydetn Rivalier)
(Tribonia Rivalier)
C. ancocisconr nsis Harris PQ
carolinae Casey
dowiana Casey
eriensis (iasey
C. decemnotata Say YK BC SK
albertina Casey
lantzi lJ.a.rcts
C. depressula depressula Casey AK BC
scapularis Casey
C. duodecimgrrttata Dejean BC AB SK MB ON PQ NB NS PE LB NF
bucolica tlasey
edmontor'.ensis Can:r
hudsonic,z Casey
proteus Birby
C. formosa folmosa Say AB SK
fletcheri Criddle
luxuriosa Casey
manitoba Lelrg
 Carabidae

C. formosa generosa Dejean SK MB ON PQ

latecincta Godd
C. formosa gibsoni Brown SK
C. fulgida westbournei Calder AB SK lln- :
elegans Calder
C. hirticollis athabascensis Graves AB bt\
C. hirbicollis couleensis Graves BC AB
C. hirticollis rhodensis Calder _ONPQNBNS_LB NF
nigrita Davis
shermani Casey
C. hirticollis shelfordi Graves AB SK MB
C. lengi versuta Casey _BCABSKMB
C. limbalis limbalis KIug NT BC AB SK MB oN pe -NS-- NF
arnoena LeConte
awemeana Casey
eldorensis Casey
m il ita r is Varas-Arangua
spreta LeConte
C. limbata hyperborea LeConte NT_ABSKMB LB
C. lirnbata nympha Casey AB SK MB
C. longilabris longilabris Say AK YK NT BC AB SK MB oN pe NB NS PE LB NF
albilabris Kirby
nouaterrae Leng
terracensis Casey
C. nebraskana nebraskana Casey BC AB SK MB
calgaryana Casey
canadensis Casey
chamberlaini Knars
montdna LeConte
splssllorsis Casey
uteana Casey
C. oregona guttifera LeConte AK YK NT BC
audax Casey
oregonella Casey
sterope Casey
C. oregona oregona LeConte BC
oualipennis Casey
quadripennis Casey
sonoma Casey
C. parowana wallisi Calder BC
azurea Cald,er
C. patruela patruela Dejean ON PQ
C. pugetana Casey BC
nigerrimoides Hatch
C. purpurea audubonii LeConte BC AB SK MB
auguralis Casey
graminea Schaupp
nigerrina Leng
C. purpurea purpurea Olivier BC ON PQ NS
marginalis Fabricius
ramosa Gistel
C. repanda novascotiae Vaurie PQ NB NS PE
C. repanda repanda Dejean NT BC AB SK MB ON PQ NB NS PE LB NF
baltimorensis Hetbst
d.uncaniKnats
hoosieri Mares
maehleri Robinson
unijuncta Casey
C. scutelaris criddlei Casey _MB
C. scutelaris lecontei Haldeman
C. scutelaris scutelaris Say _ABSK -MBONPQ
C. sexguttata denikei Brown _MBON
C. sexguttata sexguttata Fabricius oN PQ NB NS
guttata Emmons
harrisii Leng
illinoensis Mates
kansanal(nans
leuettei Casey
quadriguttata Davis
tridens Casey
uarians Ljungh
uiolacea Fabricius
Carabidae

C. tranqueba'ica borealis Harris NT BC

C. tranqueba:ica kirbyi LeOonte NT_ABSK nae
tranquebartca roguensis Harris -BCABsx
C.
C. tranqueba ica tranquebarica MB ON pe Ns NS PE NF
Herbst
crinifron.s Casey
horiconezsis Leng
minor Lt:ng
obliquat':.t Dejean
turbulenta Casey
u ir idula Varas-Arangua
uulgaris Say
uulgaris;,ninor Haris
wichitan.a Casey
C. tranquebar:ica vibex Horn BC
(Subgenus CI CINDELIDIA Rivalier)
C. punctulatri punctulata Olivier AB SK MB ON PQ NB
boulderensis Casey
chihuah:ne BaLes
fontinar,ra Casey
jenisoni llistel
prolixa (.)asey
(Subgenus C' T,INDERA Westwood)
(Plectogratr. ha Rivalier)
C. tericola cinctipennis LeConte YK _AB SK
cyanella LeOonte
C. terricola irrperfecta LeConte
C. terricola tcrricola Say
Bcy SK MB
pusilla E ay
sayanellt Casey
(Subgenus EI,LIPSOPTERA Dohktourow)
C. cuprascenr, LeConte MB
amnicohL Casey
mundukr Casey
C. lepida Dejean AB SK MB ON PQ
insomnil: Casey
C. nevadica k.nausi Leng AB SK MB

Tribe Carabini
Genus CALOSOMA Weber
Key to Norbh American species: Gidaspow (1959)
(Subgenus CitLOSOMA s. str.)
(Callipa'o Motschulsky)
(S y nc alc so ma Breuning)
C. lrigidum l.jrby BC SK MB ON PQ NB NS PE
Ieuettei Casey
(Subgenus CTTLODREPA Motschulsky)
C. scmlator (Fabricius.t _ONPQ
C. wilcoxi Letlonte _ONPQ
(Subgenus C;T LLITROPA Motschulsky)
(Paratrooa Vacher de Lapouge)
C. exlernum Sayt ON
longipenne Dejean
(Subgenus CI{RYSOSTIGMA Kirby)
(Lyperostenia Vacher de Lapouge)
(Tapinosthenes Kolbe)
C. calidum (Fabricius) NT SK MB ON PQ NB NS NF
comes Casey
concretum Casey
expansu,n Casey
laticolle Casey
stellatur,z Casey

10
 Carabidae

C. cancellatum Eschscholtz BC
aenescens LeConte
esuriens Casey
praestans Casey
rectilaterum Casey
sagax Casey
transuersum Casey
C. lepidum LeConte AB SK MB
C. obsoletum Say AB SK MB
microstictum Casey
C. tepidum LeConte BC
caelator Casey
cogitans Casey
indigens Casey
irregulare Walker
pellar Casey
semicupreum Casey
(Subgenus CAILISTHENES Fischer von Waldheim)
(Callistenia Vacher de Lapouge)
(Isostenia Vacher de Lapouge)
C. luxatum Say BC SK
debile (Casey)
diffractum (Casey)
etcaratum (Casey)
opacum (G6hin)
parowanunr (Casey)
pimelioides (Walker)
pustulosum (Casey)
reflexum (Casey)
semotum (Casey)
striatulum LeConte
striatum Breuning
tahoense Van Dyke
tegulatum (Casey)
utense (Casey)
uiator (Casey)
(?) zimtnermanl (LeConte)
C. moniliatum ( Leconte) BC AB SK
bicolor (Walker')
concinnum (Casey)
laqueatum LeConte
uancouuericum (Csjkl)
C. monticola I Casey)
neuadense (Casey)
C. subaeneum Chaudoir
C. wilkesii (Leconre)

Genus CARABUS Linn6

(Subgenus CARABUS s. str.)
(Cryocarabus Vacher de Lapouge)
(H omoeocar abus Reitter)
(.Lic hnocarab us Reitter)
(Neocarabus Hatchl
C. chamissonis Fischer AK YK NT_AB SK MB ON PQ LB
von Waldheim
brachyderus Wiedemann
groenlandicus Dejean
washingtoni Casey
C.goryt Dejean ON
clarkei Blwnenthal
limbatus Say
C. granulatus granulatus Linn6t _BCAB MB ON PQ
C. granulatus hibernicus Lindrotht _NBNSNF
C. maeander maeander Fischer AK NT BC AB SK MB ON PQ NB NS PE LB NF
von Waldheimx
atlanticus Vacher de Lapouge
excatenatus Kraatz
excostatus Kraatz
h udsonicus Motschulsky

11
Carabidae

lapilayi Castelnau
lecontei G6hin
simoni I{eyden
C. nemoralis O.F. Mullert BC AB SK oN PQ NB NS PE NF
canadensis Vacher de Lapouge
C. vinctus W':ber ON
carinatu s Dejean
georgiae Cslki
interruptus Say
ligatus Getrnat
(Subgenus DIOCARABUS Reitter)
C. taedatus a-gassii LeConte AK tiK BC SK MB ON PQ
canadic,ts Roescke
coloradensis Breuning
gladiatcr Motschulsky
montant:.cus Casey
oregonetrcis LeConte
patulicotlis Casey
stocktonensis Casey
C. taedatus taedatus Fabricius
bacciuorus Fischer von Waldheim
bicanaliceps Casey
francisutnus Casey
rainieri Van Dyke
seriatus Wiedemann
C. truncatico llis truncaticollis AK tT( NT
Eschscholtz
ala s ken :,:i s Basilewsky
lutshniA ianus Basilewsky
(Subgenus H 0MICARABUS G6hin)
C. serratus Say BC AB SK MB ON PQ NB NS PE NF
canadensis LeConte
lineato p unct atzs Dej ean
lolumi l4otschulsky
uegasensis Casey
(Subgenus T,\NAOCARABUS Reitter)
C. sylvosus s.ylvosus Say ON PQ
lhermin:eri Dejean
(Subgenus MEGODONTUS Solier)
C. vietinghol hi vietinghoffii AK YK NT
Adarr:s*

Tlibe Cychrini
Genus SPHAIRODERUS Dejean
S. canadensiri canadensis Chaudoir oN PQ NS
blancha,di Leng
palpalis Motschulsky
S. Iecontei Ie:ontei Dejean MB ON PQ NB NF
bicarinatus LeConte
diffractt.ts Casey
niagarensis Castelnau
S. nitidicoliis brevoorti LeConte SK MB ON PQ NS
granulosus Chaudoir
S. nitidicollis nitidicollis PQ NF
Chevlolat

Genus SCAI'HINOTUS Dejean

(Subgenus STIAPHINOTUS s. str.)
(Anabol'rs Gishel')
(Irichroa Newman)
(Megaliridia Casey)
S. elevatus coloradensis Van Dyke
S. viduus (Dejean)
yu_ pe NB
ir r eg ula r i s Beutenmuller
leonard:'.i (Harris)

t2
 Carabidae

(Subgenus NOMARETUS LeConte)

Key to species: Gidaspow (1973)
S. bilobus (Sayt MB ON PQ NB NS
(Subgenus PSEUDONOMARETUS Roeschke)
Key to species: Gidaspow (1973)
S. merkelii(Horn) - BC
idahoensis (Webb)
S. regularis (LeContet - BC
S. relictus (Horn) - BC
(Subgenus STENOCANTHARUS Gistel)
(Pemphus Motschulsky)
Key to species: Gidaspow (1973)
S. angusticollis (Fischer AK BC
von Waldheim)
longipes Casey
nigripennis Roeschke
opacus Casey
uelutinus M6n6tries
(Subgenus NEOCYCHRUS Roeschke)
Key to species: Gidaspow (1973)
S. angulatus (Harris) - BC
rnaritimus Van Dyke
(Subgenus BRENNUS Motschulsky)
Key to species: Gidaspow (1968)
S. johnsoni Van Dyke - BC
hlahowyae Perrault
S. marginatus (Fischer AK BC AB
von Waldheim)
columbianus (Casey)
confusus (Casey)
cupripennis (Casey)
/ollor (Roeschke)
fulleri (Horn)
gracilis (G6hin)
insularis (Casey)
montanicus (Casey)
oregonus (Casey)
wrangelli (Casey)

Genus CYCHRUS Fabricius

Key to North American species: Gidaspow (1973)
C. rickseckeri LeConte - BC
C. tuberculatus Harris - BC
pustulosus Casey

Tlibe Nebriini
Genus LEISTUS Friilich
Key to North American species: Erw.in (1970a)
(Subgenus LEISTUS s. str.)
L. fernrgineus (Linn6)f - NF
(Subgenus NEOLEISTUS Erwin)
L. ferruginosus Mannerheim AK BC AB
ferrugineus Dejean
nigropiceus Casey

Genus PELOPHILA Dejean

P. borealis (Paykull)* AK YK NT BC AB SK MB ON PQ - LB NF
californica Motschulsky
e s c hs c holtzi (Mannerheim)

shermani Casey
ulkei Horn
P. rudis (LeConte) NT BC AB - MB ON - NF

13
Carabidae

Genus NEBRIA Latreille

N. acuta acuta Lindroth
N.
N.
N.
arkansana edwardsi Kavanaugh
brevicollis (Fabricius)t
campbelli Kavanaugh
_v
AK BC
BC

BC
v_ PQ

N. charlottae Lindroth BC
N. crassicorn is crassicornis BC
Van fryke
N. crassicornis intermedia Van Dvke BC AB
N. diversa LeConte BC
liuidaLeConte
townsendi Casey
N. frigida R.I'. Sahlberg* BC
paruula J. Sahlberg
reducta'Casey
uiridis E:lorn
N. gebleri cai;cadensis Kavanaugh _BC
N. gebleri gebleri Dejean AK BC AB
melanariaHatch
N. gouleti Kavanaugh ?
N. gregaria Fischer von Waldheim AK
cuneata casey
N. gyllenhali castanipes (Kirby)* AK YK NT BC AB SK MB ON PQ LB NF
curtulata Casey
(?) elias Motschulsky
labradorica Casey
moesta leCorrte
prominens Casey
N. haida Karanaugh _BC
N. hudsonica LeConte
N. kincaidi k ncaidi Schwarz
_YKNTBCABSK
AK BC
yu_
columbi'ma Casey
N. lacustris I lcustris Casev MB ON PQ NB
expansa Casey
N. lituyae Kavanaugh AK
N. Iouiseae Kavanaugh -BC
N. mannerheimii Fischer AK BC
von S'aldheim
coruallis Casey
hippisleyi Casey
oregona Casey
N. meanyi meanyi Van Dyke AKYK_BC
N. meanyi sylvatica Kavanaugh -BC
N. metallica Fischer von Waldheim AK BC
pacifica Chaudoir
_PQ
N. nivalis gar;pesiana Kavanaugh
N. nivalis nir alis (Paykull)* AK YK I.TT _ _PQ
LB
v
bifaria llIannerheim
mollis l!:lotschulsky
N. obliqua obliqua LeOonte AKYK_BC SK
incerta Casey
obtusa I.,eConte
testaceipes Casey
texana Casey
NB PE
N. pallipes Say
N. piperi Van Dyke AK--eC 9* PQ NS

N. sahlbergii sahlbergii Fischer AKYK-BCAB

von Waldheim
aleuta\anDyke
uiolacea Motschulsky
N. schwarzi schwarzi Van Dvke AB
N. suturalis l.eConte :" ON PQ LB
longula LeConte
nimbosa Casey
N. virescens .florn BC
breuis Casey

T4
 Carabidae

Tlibe Notiophilini
Genus NOTIOPHILUS Dum6ril
N. aeneus (Herbst) oN PQ NB NS
porrectus Say
N. aquaticus (Linn6)x BC SK MB ON PQ NB NS LB NF
hardyi Putzeys
N. biguttatus (Fabricius)t BC NB NS NF
N. borealis Harris* AK YK NT BC AB MB ON PQ _LB NF
N. directus Casey BC AB
lanei lJatc]l
N. intermedius Lindroth AK BC MB PQ LB NF
N. nitens LeConte BC
N. semistriatus Say AK YK- BC AB MB ON PQ NS LB
americanus }{atris
coloradensis Casey
confusus LeConte
punctatus LeConte
N. simulator FalI BC AB
euanescens Casey
N. sylvaticus Eschscholtz AK BC

Tribe Loricerini
Genus LORICERA Latreille
Key to species: Ball and Erwin (1969)
(Subgenus LORICERA s. str.)
L. decempunctata Eschscholtz AK _BC
L. pilicornis congesta Mannerheimx AK
L. pilicornis pilicornis
(Fabricius)*
AK YK NT BC AB SK MB ON PQ NB NS is NT

californica LeConte
neoscotica LeConte
semip unct ata Eschscholtz
sierraeYanDyke
uteana Casey

Tribe Opisthiini
Genus OPISTHIUS Kirby
O. richardsoni Kirby AK YK NT BC AB SK

TYibe Elaphrini
Genus DIACHEILA Motschulsky
Arctobia C.G. Thomson
D. arctica amoena (Faldermann)* LB
americana Motschulskv
subpolaris LeOonte
D. polita (Faldermann)* AK YK NT

Genus BLETHISA Bonelli

HelobiumLeach
Rhaphiona Fischer von Waldheim
Key to species: Goulet and Smetana ( 1983)
B. catenaria W.J. Brown* AKYKNT_MB
B. inexspectata Goulet and Smetana AK
B. julii LeConte NT_ABSKMBONPQNBNSNF
B. multipunctata aurata AK YK NT BC AB SK MB ON PQ NB NS PE - NF
Fischer von Waldheimx
hudsonica Casey
B. oregonenis LeConte BC
acutangula Chaudoir
columbica Casey
B. quadricollis Haldeman NT BC AB SK MB ON PQ NB NS PE LB NF
arnericana Hatris

15
Carabidae

Genus ELAI'HRUS Fabricius

Key to species: Goulet (1983)
(Subgenus ARCTELAPHRUS Semenov)
Gyllenhal*
E. lapponicus lapponicus AK YK NT BC AB - MB ON PQ - LB
obscurit.tr I{trby
E. lapponicus obliteratus AK
ManrLerheim
(Subgenus NEOELAPHRUS Hatch)
E. cicatricosrrs LeConte PQ
rhodear; us Casey
E. clairuillei KJrby AK YK NT BC AB SK MB ON PQ NB NS PE LB NF
/rosll H.ppisley
lynni Pierce
politus ,eCorrte
torreyer,rsis Tanner
E. fuliginosus Say AB - MB ON PQ
E. olivaceus LeOonte NT BC AB SK MB ON PQ NB NS PE - NF
(Subgenus ELAPHRUS s. str.)
(.Trichel tphrus Semenov)

E. americanrrs americanus Dejean AK YK NT BC AB SK MB ON PQ NB NS PE LB NF

bitubercsus Casey
gratiost^s Mannerheim
interme lizs Kirby
punctat ssjmzs LeConte
sinuatu i LeConte
E. americanrrs syivanus Goulet BC AB
E. californicrLs Mannerheim AK - NT BC AB SK MB ON PQ NB NS PE
hesperh'.s Casey
similis LeCorrte
E. lecontei Crotch - NT BC AB SK MB ON
deuinctt:s Casey
spisslco''nls Casey
E. parviceps Van Dyke* AK - NT - MB ON
E. ruscarius Say ON PQ
texanus Casey
E. tubercular us Maklin* AK YK NT
(Subgenus E LAPHROTERUS Semenov)
(Elaphriatus Semenov)
E. angustico lis angusticollis AX YK NT
R.F. liahlberg*
E. purpuranr' Hausen AK YK NT BC AB
pallipes Hotr'

Tribe Omophronini
Genus OMO.?HRON Latreille
Key to North American species: Benschoter and Cook (1956)
O. americanus Dejean AB SK MB ON PQ NB NS PE - NF
fontinal;.s Casey
illustris Casey
iridescens Casey
locustri'; Casey
lengi Casey
proximus Chandler
soil Kirby
tanneri Chandler
texanus Casey
O. Iabiatus (Fabricius) NS
O. ovalis Horn - BC AB SK
concinnits Casey
frater Casey
gemma'Jasey
O. robustus llorn AB ON
breuipen.nis Casey
decoloratus Fall

16
 Carabidae

O. tesselatus Say AB SK MB ON PQ NB NS PE
ellipticus Casey
lecontei Dejean

Tribe Promecognathini
Genus PROMECOGNATHUS Chaudoir
P. crassus LeOonte BC
contractus Casey
corpulentus Casey
grandiceps Casey

Tribe Scaritini
Genus SCARITES Fabricius
S. subterraneus Fabricius ON
beckwithii Stephens
denticollis Chaudoir
interruptus (Herbst)
spinipes (Sulzer)
subtemeus Bonelli

Genus PASIMACHUS Bonelli

P. elongatus LeConte AB SK MB
angustulus Casey
euanescens Casey
pirnalis Casey
uegasensis Casey
uernicatus Casey

Tribe Clivinini
Genus DYSCHIRIUS Bonelli
Key to North American species: Bousquet (1g88)
(Subgenus AKEPHORUS LeConte)
D. obesus LeConte - BC
(Subgenus DYSCHIRIUS s. str.)
D. affinis Fall ON PQ
duplicatus Fall
D. alticola Lindroth -BC
D. aratus LeConte _BCAB MB
D. brevispinus LeConte -ONPQ
D. campicola Lindroth AB MB
D. compactus Lindroth
D. criddlei FaIl
D. er5rthrocerus LeConte
_:rf MB
-oNPQ
_ONPQ
D. foigidus Mannerheim AK YK NT BC LB
aeneolus LeConte
D. globulosus (Say) AK YK -BCAB SK MB ON PQ NB NS NF
paruus LeConte
D. haemorrhoidalis (Dejean)
D. hiemalis Bousquet AK NT BC AB -oN
MB ON PQ _LB
D. integer LeOonte AK NT BC AB SK MB ON PQ NB NS PE NF
apicalis LeOonte
basalis LeConte
nigripes LeConte
tr an s m arin u s Mannerheim
D. interior Fall _AB SK MB
arizonicus Van Dyke
D. laevifasciatus Horn BC AB
_NF
_v
D. larochellei Bousquet NB NS
D. longulus LeConte BC AB SK MB_PQ
D. montanus LeConte BC AB MB ON PQ
thompsoni Hatcll

L7
Carabidae

D. nigricornis Motschulsky* AK YK NT
secretus Fall
D. pacificus .l,indroth _BC
D. pallipenn:s (Say) AB SK ON PQ
D. perversus Fall AB MB
desertut Fall
D. pilosus LrConte MB ON PQ
hispidu,; LeConte
D. planatus ,indroth AB SK MB
D. politus (Dejean)* AK YK _BCABSKMB ON PQ NF
aureolu.; Notman
D. quadrima :ulatus Lindroth AB SK MB
D. sellatus LeConte PQ NB NS PE NF
D. setosus LcConte AK nc AB SK MB ON PQ NB NS PE
alternatus Hatch
D. sextonj Bousquet ON
D. sphaerico lis tSayr BC AB sr me ON PQ NB NS po NF
subpunt:tatus Hatch
D. subarcticus Lindroth AK YK NT
D. tenuispin rs Lindroth AB
D. timidus Lindroth
D. truncatus LeConte AK Nr sc AB
-me
SK MB
Genus CLIVINA Latreille
(Subgenus S 0MICLN'INA Kult)

C. dentipes.l)ejean ON
coruina Putzeys
confusa LeConte
fissipes Putzeys
georgiana LeConte
(Subgenus PARACLIVINA Kult)
C. bipustula' a (Fabrici us) ON
quadrintaculafo (Palisot de Beauvois)
(Subgenus CLIVINA s. str.)
C. americana Dejean ON PQ NS
acuducta Haldeman
cordata Putzeys
ludouiciana Putzeys
C. collaris tIlerbst)t BC MB ON PQ
elongatt Randall
randalli LeConte
C. fossor il,inn6)t _BCAB ON PQ NB NS PE NF
C. impressifrons LeConte
C. oregona Fal) -BC _-ONPQ
Genus SCHIZOGENIUS Putzeys
Key to North American species: Whitehead (1972)
(Subgenus SCHIZOGENIUS s. str.)
S. amphibius (Haldeman) PQ
frontalis (LeConte)
S. depressus LeConte BC
S. ferrugineus Putzeys sx ON pe
sulcatu;; (LeConte)
S. lineolatus {Sayr ON PQ
S. litigiosus Fall ec
S. sulcifrons Putzeys ON PQ NB

TYibe Brachininr
Genus BRA(IHINUS Weber
Key to North American species: Erwin (19706)
(Subgenus NEOBRACHINUS Erwin)
B. cordicollis Dejean - ON PQ NB
gracilis Blatchley
leptocerus Chaudoir
uelox LeConte

18
 Carabidae

B. cyanipennis Say MB ON PQ NB
cephalotes Dejean
rejectus LeConte
B. cyanochroaticus Erwin - BC - SK MB ON PQ
B. fumans (Fabriciust - BC AB - MB ON PQ
affinis LeConte
amplipennis Bates
atbarae Stehr
cyanopterus LeConte
suffians LeConte
tabasconus Bates
B. janthinipennis (Dejean t - ON PQ
pumilio LeConte
B. medius Harris - BC - ON PQ
minutus }J.arris
B. ovipennis LeConte - ON PQ
B. quadripennis Dejean - BC AB SK MB ON
stygicornis Say
I schern ihhii Mannerheim
B. tenuicollis LeConte - ON PQ
ballistarius LeConte
similis LeConte
B. texanus Chaudoir AB

Tribe Paussini
Genus METRIUS Eschscholtz
M. contractus contractus - BC
Eschscholtz

Tribe Gehringiini
Genus GEHRINGIA Darlinston
G. olympica Darlington NT BC AB

Tribe Broscini
Genus BROSCUS Panzer
B. cephalotes (Linn6tf NS PE

Genus MISCODERA Eschscholtz

Leiochiton Curtis
M. arctica (Paykull)x AK YK NT BC AB SK MB ON PQ - NS PE LB NF
americana Mannerheim
hardyi Chaudoir

Genus BROSCODERA Lindroth

B. insignis tMannerheim) AK BC

Genus ZACOTUS LeConte

Z. matthewsii LeConte AK BC
angustus Casey
fre deric ki Nunenmacher
subopacus Hopping

Tlibe Trechini
Genus BLEMUS Dejean
Las iotrec hu s Ganglbauer

B. discus (Fabriciustf - ON PQ NB

Genus TRECHUS Clairville

(Subgenus TRECHUS s. str.)

19
Carabidae

T. apicalis Motschulskyx AK YK NT BC AB SK MB ON PQ NB NS LB NF
borealis Schaeffer
brumalts Casey
canader'.sis Putzeys
fuluus (",eCorte)
micans .LeConte)
pallescens Casey
puritan,ts Casey
rhodensis Casey
T. chalybeus Dejean* AK BC AB
brachycierus Jeannel
califor n ic us Motschulsky
tahoensi.s Casey
T.
T.
crassiscapus Lindroth
obtusus Erichsont ---ec-- _ :.T _
NS
_:uIu
T. oregonensis Hatch -BCAB
T.
T.
ovipennis Motschulsky
quadristriatus (Schrank)f
y_ _ BC_ _ _ONPQ
pe- y
T. rubens (Fabricius)t PQN eNs
T. tenuiscapr.rs Lindroth _YK_BCAB

Tribe Bembidiini
Genus ASAI HIDION des Gozis
Tachypi:.ts Dejean
A. alaskanurn Wickham AK YK NT
A. yukonens'r Wickham AKYK_BCAB
Genus BEM.IIIDION Latreille
(Subgenus CHRYSOBRACTEON Netolitzky)
(Parabrzcteon Notman)
B. alaskense Lindroth AK
B. balli Lindroth
B. carinula ( jhaudoir
---ABix
YK NT BC AB SK ff,fS On PQ NB NF
tubercu o/um Notman
B. co1villenstr Lindroth .AK
B. foveum Motschulsky* AKYKNTBC_SK MB
bryanti Carr
(.?) glabr iusculum Motschulsky
grahamiHatch
B. hesperium Fall BC
binariu;n Casey
B. inaequale inaequale Say MB ON PQ NB NS PE
arenari,r,m Dejean
lacustre LeConte
B. inaequale opaciceps Casey AK BC AB SK
B. lapponicum Zetterstedt* AK_NT BC AB SK
pugetanumFall
B. levettei Casey AK YK NT BC SK MB oN PQ NB NS LB NF
carrianlm Casey
B. lorquinii (lhaudoir BC
tacomar Clssy
B. punctatostriatum Say BC AB SK oN PQ NB
(?) sigiliare Say
stigmat:icum Dejean
B. zephyrurr Fall BC
marginosum Casey
tristicuium Casey
(Subgenus BRACTEON Bedel)
B. bowditchii LeConte BC
(Subgenus ODONTIUM LeConte)
B. confusum Hay'ward AB SK MB ON PQ
aeneorubrum Casey
marquettense Casey
nitidult:m Dejean

20
 Carabidae

B. coxendix Say NT_AB SK MB

snowi Casey
unicum Casey
uenator Casey
uigilans Casey
ueesi Hatch & Ortenburger
(Subgenus PSEUDOPERYPHUS Hatch)
(B r a cteomimus Lindroth)
B. chalceum Dejean BC AB SK MB ON PQ NB NS NF
antiquum Dejean
dilatatum (LeConte)
B. honestum Say ON PQ NB
basale (LeConte)
platyderum Chaudoir
(Subgenus OCHTHEDROMUS LeConte)
B. americanum Dejean
illini Casey
-oN PQ

B. bifossulatum (LeOonte) BC AB SK MB ON PQ
cheyennense Casey
ferreum Casey
nuperum Casey
regestum Casey
suffiatum Casey
(Subgenus EURYTRACHELUS Motschulsky)
(Eudromus l\trby)
(P la ty t r a c hel us Motschulsky)

B. interventor Lindroth AK YK NT BC AB SK MB ON PQ
B. nitidum (Kirby) AK YK NT BC AB SK MB ON PQ
edolatum Casey
(Subgenus HYDRIUM LeConte)
B. levigatum Say AB
delawarense Casey
(Subgenus METAILINA Motschulsky)
B. dyschirinum LeOonte AK YK BC AB
agitabile Casey
aleneanum Casey
atrolucens Casey
heechelum }Iatch
perturbatum Casey
speculinum Casey
B. lampros tHerbsilf BC NF
B. properans ( Stephens tt PQ NB NS PE NF
(Subgenus PI{YLA Motschulsky)
B. obtusum Audinet-Servillet oN PQ
(Subgenus LIONEPFIA Casey)
Key to species: Erwin and Kavanaugh (1981)
B. castum Casey AK BC
brumale Casey
nescium Casey
serenum Casey
uaciuum Casey
B. disjunctum Lindroth BC
B. erasum LeConte BC AB
lasciuum Casey
lubricum Casey
probatum Casey
B. lindrothellum Erwin & Kavanaugh AK BC
B. lummi Erwin & Kavanaugh BC
B. sequoiae Lindroth BC
(Subgenus ACTEDIUM Motschulsky)
B. lachnophoroides Darlington AB SK
(Subgenus TRECHONEPHA Casey)
B. iridescens I LeConte) BC AB

2I
Carabidae

atnicum Casey
deceptor Casey
fabrum Oasey
impium Casey
obliuiost:.m Casey
p ar allel c'colle (Motschulsky)
repens Casey
uolatile t)asey
R
J

B. simplex Hayrvard PQ
essexense Casey
$

B. kuprianov:.i Mannerheim AK YK NT BC AB
adultum Casey
biimp r es s u m Mannerheim
bucolicum Casey
dilutum Casey
funereurz LeConte
maehlint Hayward
ouipennt (Motschulsky)
(Subgenus PI,ATAPHODES Ganglbauer)
B. arcticum Lindroth* AK YK
B. brachytho:'ax Lindrothx AK YK NT BC
B. complanul om (Mannerheim) AK BC AB
paruulutn Notman
B. compressum Lindroth* AK YK NT
B. farrarae Ilatch AK BC
B. haruspex l.)asey _BC
B. incertum ( Motschulsky) AKYK?BC AB
ampliatum Casey
blanditum Casey
(Motschulsky)
(?') breue
improuit,um Casey
liuidulu n Casey
saturatum Casey
testatun Casey
tetr ag ly 1 t u m (Mannerheim)
B. manninge:rse Lindroth AK YK NT BC AB
ochropu:; Lindroth
AK _AB PQ NB NS LB NF
B. occultator Notman
B. quadrifoveolatum Mannerheim AK YK BC AB
SK
9*
iller Casey
B. rosslandicrm Lindroth BC AB
B. viator Casey BC
(Subgenus P.I,ATAPHUS Motschulsky)
(.Melomt'lus Casey)
(M icromelomalus Casey)
(Trachelonepho Casey)
B. basicorne Notman _PQ_NS
B. carolinense Casey _PQNBNS NF
heeneanzm Casey
B. curtuiaturn Casey AK YK BC AB
decrepitu.m Casey
effetum Casey
flebile Casey
timefactum Casey
B. gebleri turbatum Casey YK BC AB
conflictt:m Casey
B. gordoni Lindroth BC
B. gratiosum Casey AK YK BC AB
pallidumHatch
B. hyperboraeorum Munster* AK YK NT
B. kalumae Lindroth BC
B. planatum (LeConte) AK YK NT BC AB--oNpeNnNs--Nr
adjutor Casey
aequalis: (Walker)
solutum Casey

22
 Carabidae

B. planiusculum Mannerheim AK BC
B. rufrnum Lindroth AK YK - BC .AB
B. rusticum lenensoides Lindroth AK YK - BC
B. nrsticum rusticum Casey - ON PQ NB NS - LB NF
notmani Casey
B. siemicola Casey - BC
B. stillaguamish Hatch - BC
B. sulcipenne hyperboroides AK YK - BC
Lindroth
B. sulcipenne prasinoides Lindroth - BC .AB - MB - PQ NB LB NF
(Subgenus BLEPHAROPLATAPHUS Netolitzky)
B. hastii C.R. Sahlbergx AK YK NT BC MB ON PQ - LB
(Subgenus TRICHOPLATAPHUS Netolitzky)
B. planum (Haldeman) - ON PQ NB NS
filicorne Casey
guexii Chatdo\r
uulsum Casey
B. rolandi Fall - ON PQ NB NS
(Subgenus HIRMOPLATAPHUS Netolitzky)
B. concolor (Kirby) AK YK NT BC .AB SK MB ON PQ NB NS NF
longulum (LeConte)
subaeneum (LeConte)
B. nigrum Say - ON PQ NB NS PE
facile Casey
morosum Casey
B. quadrulum LeConte - YK - BC .AB
aegrotum Casey
callidum Casey
porrectum Casey
tartareurn Casey
tritum Casey
uiridinigrum Casey
B. recticolle LeConte - BC AB
oblatum Casey
pertinax Casey
Chaudoir
tetr ag onoderum
umbraticola Casey
B. salebratum (LeOonte) NT BC .AB SK MB ON PQ NB NS NF
consessor Casey
inopinum Casey
mackinacense }latc}r
purpurascens (LeConte)
(Subgenus HYDRIOMICRUS Casey)
B. quadratulum Notman PQ NB NS NF
proximum Notman
B. semistriatum (Haldeman) - ON PQ - NS
(Subgenus BEMBIDIONETOLITZKYA Strand)
(Daniela Netolitzky)
B. mckinleyi carneum Lindroth - BC AB
B. mckinleyi mckinleyi FalI AK YK
$
B. lenae Csiki* AK YK NT
intermedium Poppius
$

B. commotum Casey _BCAB

seclusum Casey
B. nebraskense LeConte _BCAB
denueranum Casey
gouanicum Casey
tractabile Casey
$
B. actuosum Casey - BC AB
debilicolle Casey
urgens Casey

23
Carabidae

B. nevadense Illke BC
uiaticun.: Casey
B. plagiatum (Zimmermann) ON
B. striola (LeConte) BC
angustit:r Casey
modulatum Casey
shastani.cum Casey
uancouu zri Casey
(Subgenus PiIRYPHANES Jeannel)
B. dauricum Motschulskyx AK YK NT MB
B. grapii Gyl enhal* AK YK NT BC AB SK MB ON PQ NB NS is NF
nitens Casey
picipes (t{irby)
scrutatum Casey
seducturn Casey
B. lacunariurn (Zimmerrnann) ON PQ
histricurn Casey
militare Casey
B. platynoides Hayward _BC
insopita:zs Casey
merens li)asey
optatum Casey
sedulutr, Casey
B. stephensi lrotch'l -oN PQ NB NS PE NF
canadense Hayward
B. texanum ( lhaudoir) MB
cogitans Casey
inquietum Casey
B. yukonum lall* AK YK NT BC MB PQ

$
B. bimaculat rm (Kirby) AK YK NT BC AB SK MB ON
B. postremur:r Say -PQ NB
B. sordidum rKirby) AI< YK NT BC AB SK MB oN PQ LB
B. umiatenser Lindroth AK
(Subgenus PIIRYPHUS Stephens)
B. bruxellense Wesmaelt -PQNBNSPE NF
B. femoratunr Sturm* NB NS
B. obscurellu m (Motschulsky)x AK YK NT BC AB sx lle ONPQNB_PE NF
albidipezne Casey
caducunt Casey
fuscicrus (Motschulsky)
lucidum (Mannerheim)
mixtum LeConte
parowar.um Casey
petulans Casey
(?) subinflatum (Motschulsky)
B. petrosum ilttuense Lindroth AK
B. petrosum'letrosum Gebler* AK YK Nr ec AB SX ON PQ Nn lls pr LB NF
castaliu;n Casey
exiguiceps Casey
lepusculum Casey
lucidum (LeConte)
substric,um (LeConte)
wenatchee Hatch
B. poppii sch,rlleri Lindroth AK
B. rupicola (lGrby) AK NT BC AB sr l.rn oN-
auersan:; Casey
B. sejunctum sejunctum Casey
B. sejuncturn semiaureum Fall
NT
_BC
AB SK
_:a YNS LB NF

fenderi llatch
B. tetracolum Sayt _BC oN PQ NB NS NF
nactum Casey
$

B. transverszrle Dejean YK_BC SK MB ON PQ NB NS LB NF

acoman,tm casey
animatu.m Casey

24
 Carabidae

(?) erosum (Motschulsky)

ercursum Casey
haplogonum Chaudoir
ma nner heimii (LeConte)
marinicum Casey
perspicuum (LeConte)
sarpedon Casey
tuolumnae Casey
$

B. scopulinum (Kirby)x AK YK NT BC AB SK MB ON PQ NB NS PE LB NF
bellulum Casey
gelidum (LeConte)
(Subgenus LIOCOSMIUS Casey)
B. mundum (LeConte) BC
b ifa s c iat um (Motschulsky)
hilare Casey
(Subgenus PERYPHODES Casey)
B. ephippigerum (LeConte) _BC
B. salinarium Casey _BCAB SK MB
$

B. constricticolle Hayward _AB

B. nudipenne Lindroth _AB SK MB
(Subgenus EUPETEDROMUS Netolitzky)
B. graciliforme Hayward -oNPQ
B. immaturum Lindroth _ONPQNBNSNF
B. incrematum LeConte AK YK Nt EC AB sr MB ON PQ NB NS PE LB NF
arcuatum LeConte
fortunatum Casey
gulosum Casey
/ezgi Notman
mobile Casey
nubiferum Casey
oblectans Casey
sernotum Casey
(Subgenus LEUCIIYDRIUM Casey)
B. tigrinum LeConte -BC
$
B. consimile Hayward ,AB SK MB
B. flohri Bates BC .AB SK MB
henshawi Hayward
B. obtusidens Fall ,AB SK MB
B. scudderi LeConte
spissicorne Casey
--nc ,AB SK MB

B. dejectum Casey ,AB SK MB

aberti}].atch
fidele Casey
tencenti Hatch
B. mormon Hayward _BC
B. obtusangulum LeConte _ BC ,{B SK MB
cornix Casey
(Subgenus NOTAPHUS Stephens)
B. aeneicolle (LeConte) NT BC .q,B SK MB
B. castor Lindroth _ ,A,B SK MB ON PQ NB NS NF
B. coloradense Hayward NT BC .{B SK MB
albertanum Casey
amplipenne Casey
imperitum Casey
prociduum Casey
B. cordatum (LeConte) MB ON
placabile Casey
B. debiliceps Casey BC

25
Carabidae

B. dorsale Say _BCAB SK MB

B, graphicum Casey NT BC AB SK MB ON
lassulutn. Casey
B. indistinctum Dejean AK BC
barbarae Casey
caudex (;asey
conse nt a neum (LeConte)
curiosun,, Casey
derisor C.iasey
deuinctum Casey
erpositurn Casey
extricatum Casey
formale l}asey
franciscc:,num Casey
ornatellt'.m Casey
reconditum Casey
B. insulatum (LeConte) NT BC AB SK MB
caliginosum Casey
B. intermeditLm (Kirby) AK BC AB SK MB ON PQ
B. nigripes (IrJrby)x AK YK NT BC AB SK MB ON PQNB_PELB NF
concitatt.Lm Casey
quadrah colle (Mannerheim)
retracturn Casey
B. patruele Dejean NT BC AB SK MB ON PQ NB NS PE NF
contristans Casey
editum Oasey
exclusunt Casey
fenisex (.asey
fraternun LeOonte
imitator Casey
marcidum Casey
mediocn: Casey
monstra,'.um Casey
plectile Oasey
pos ticun,'. (Haldeman)
subexiguum Casey
B. rapidum (LeConte) SK MB ON PQ NB
docile Ctsey
fugitans Casey
negligeni Casey
sociale ( asey
B. semipunct etum (Donovan)* AK YK BC AB SK ON
accuratt'.m Casey
prouoanitm Casey
B. umbratum (LeConte) NT BC AB SK
u ar iolos u m ( Motschulsky)

B. variegaturn Say ON PQ
postfascntum Hamilton
B. versutum LeConte oN PQ NB
$

B. constricturn (LeConte) NB NS PE
ciuile Ca.sey
festinans Casey
uernula Casey
B. contracturr Say _PQ NB NS PE NF
B. viridicolle (LaFert6-S6nectdre) NT AB SK MB ON
hamiferum Chaudoir
particeps Casey
R

B. oberthueri Hayward AB SK MB ON PQ NB
(Subgenus FIJRCACAMPA Netolitzky)
B. affine Say ON
decipiens Dejean
fallax Dejearr
thespis Casey
R
J

B. impotens Casey BC SK MB ON NB

26
 Carabidae

aestiuum Casey
frugale Casey
gratuitum Casey
imbelle Casey
ind,igens Casey
pictum (LeConte)
pullulum Casey
uirgatulum Casey
B. mimus Hay'ward - SK MB ON PQ NB NS PE - NF
pellax Casey
rotundiceps Casey
B. timidum (LeConte) NT BC AB SK MB ON
oregonum Casey
sord.idulum Chaudoir
B. versicolor (LeConte) AK - NT BC AB SK MB ON PQ NB NS PE LB NF
terracense Casey
tolerans Casey
wisconsinium Casey
(Subgenus EMPHANES Motschulsky)
(Omala Motschulsky)
B. diligens Casey - BC AB SK MB
parabile Casey
uilescans Casey
B. vile (LeConte.) - BC
(Subgenus BEMBIDION s. str.)
(Lopfto Stephenst
B. mutatum Gemminger & Harold AK YK NT BC AB SK MB ON PQ - NS NF
arillare (LeConte)
uegetum Casey
B. praecinctum LeConte AB SK MB
alutaceum Hatch
ueridicum Casey
B. quadrimaculatum dubitans AK YK NT BC AB SK MB
(LeConte)
caseyi Letg
crurale (LeConte)
gregale Casey
pugetanurn Casey
sapphicum Casey
tenax Casey
B. quadrimaculatum oppositum Say AB SK MB ON PQ NB NS PE - NF
(Subgenus SEMICAMPA Netolitzky)
B. convexulum Hayrvard AK YK - BC AB
nouellum Casey
B. morulum LeConte AK YK NT BC AB SK MB NF
browni Lindroth
B. muscicola Hayward - SK MB ON PQ NB NS PE
B. praticola Lindroth - BC - SK - ON PQ NB
B. roosevelti Pic AB SK MB
blaisdelli Casey
concinnum Blaisdell
p er conc innunt Blaisdell
B. semicinctum Notman - ON PQ NB NS
(Subgenus DIPLOCAMPA Bedel)
(Paralopha Casey)
B. transparens (Gebler)* AK YK NT BC AB SK MB ON pQ NB NS pE LB NF
edmontonense Casey
sulcatum (LeConte)
trepidum (LeConte)
(Subgenus TREPANEDORIS Netolitzky)
B. acutifronsLeConte AK - NT BC AB SK MB
m ic u la u m Hatch
r o r e t ic t
B. canadianum Casey NT - AB SK MB ON PQ
B. concretum Casey AK YK NT BC AB SK MB ON PQ NB NS PE - NF
congruens Casey
habile Casey

27
Carabidae

B. connivens (LeConte) BC
digressum Casey
elizabetl:ae Hatch
B. fortestriatrm (Motschulsky) AK YK NT BC SK MB ON PQ NB NS PE NF
cdutum .LeConte)
concurre'.ns Fall
peregrinum Casey
utnbrati:um Casey
B. frontale (l.eOonte) _BC_SKMBONPQ NS PE
B. pseudocaubum Lindroth _BCABSKMBONPQ )u
B. siticum Cz.sey _BC
adolescew Casey
(Subgenus AIvIERIZUS Chaudoir)
B. oblonguloiJes Lindroth _BC
B. oblongulurn (Mannerheim) AK BC
crassicorne (Casey)
heeni (C*ey')
B. spectabile (Mannerheim) AK BC
longicor.rc (Casey)
B. wingatei Filand oN PQ NB NS LB NF

Genus PHR\PEUS Casey

P. ricksecker (Hayward) BC
rutilinut Casey
Genus ELAP HROPUS Motschulsky
Barytac,rys Chaudoir
Tachyur a Motschulsky
E. anceps (Lt.Conte) NT AB SK MB ON PQ
YNS
E. capax (Le0onte) -oN
E. dolosus (LeConte) -oN
apacheanus { Casey)
audax (l,eConte)
rapar (I eConte)
E. granarius (Dejean) -oN PQ _NS
glossem'r (Casey)
E. incurwus ( Jay) BC SK MB ON PQ NB NS PE
parallel 1s (Casey)
rhodens'.s (Cslki)
E. parvulus (Dejean)t BC
E. saturatus (Casey) ---oNFeNeNs--
_ONPQNBNS
E. tripunctatus (Say)
barnesi (Stehr)
laredoanus (Casey)
seruzs ({lasey)
E. vernicatus, (Casey) SK MB ON PQ NB
laetificus (Casey)
unionis (Csiki)
E. vivax (LeConte) oN PQ
mendar (LeConte)
E. xanthopur (Dejean) ON NS
ancilla rCasey)
famelia.Ls (Casey)
loaipennis (Casey)

Genus TACFiYTA Kirby

Key to species: Erwin (1975)
BC SK MB ON NS
T. angulata Casey
T. falli (Hayrvard)
YK
BC
AB PQ
fu
T. nana inornata (Say) BC ON PQ
arizonica Casey
californ;;,ca Casey
collaris Casey
debilicol,lis Casey
picipes lktby
ulari s (Motschulsky)
riu
T. nana kirb ri Casey BC AB ON PQ

28
 Carabidae

Genus POROTACHYS Netolitzky

P. bisulcatus tNicolaitt - ON PQ NB
frontalis (Hayward)
Genus POLYDERIS Motschulsky
Microtachys Casey
P. laevis {Sayt - ON PQ - NS
congesta (Casey)
flumenalis (Casey)
recta (Casey)
troglodytes (Dejean)
unistriata (Casey)
unistriolata (Cslki)

Genus TACHYS Stephens

Isotachys Casey
T. bryanti Lindroth ,\B
T. halophilus Lindroth ,\B SK MB
T. vittiger LeConte - BC
marginellus LeConte

Genus PARATACHYS Casey

P. edax (LeConte) - BC
P. oblitus {Casey) - ON PQ
appalachius (Casey)
cuneatus (Casey)
gentilis (Casey)
iowensis (Casey)
obliquus (Casey)
P. proximus tSayt - ON PQ
nubifer {Casey)
P. rhodeanus (Caseyr - ON PQ
P. scitulus (LeConte) - ON PQ NB
pallescens (Casey)
p aIIidiu sc u Iu s (C sikl)
Genus MIOPTACIfYS Bates
T ac hy menis Motschulsky
M. flavicauda (Say) - BC r\B ON PQ
ma rg inicollis (Motschulsky)
r eflexicoll is (Motschulsky)

Tlibe Nomriini
Genus NOMIUS Castelnau
Haplochile LeConte
N. pygmaeus (Dejeant - BC AB - MB ON PQ

Genus PSYDRUS LeConte

Monillopatrobzs Hatch
P. piceus LeConte - BC AB SK MB ON PQ
punctdtus (Hatch)

Tribe Patrobini
Genus PLATYPATROBUS Darlington
P. lacustris Darlington NT - l\B SK MB ON PQ NB

Genus PATROBUS Dejean

Geo p a t r ob u s Darlington
N eo pat r ob u s Darlington
P. lossifrons (Eschscholtz)x AK BC l\B
dimorp hic us Darlington
/uluus Mannerheim
latiusculus Chaudoir
long i uen I ri s Mannerheim

29
Carabidae

P. foveocollis (Eschscholtz)* AK YK NT BC AB SK MB ON PQ - LB NF
ang usticollis Mannerheim
insularis Casey
Iaeuiceps Casey
obt usiu sc u I us Chaudoir
tenuis (l'eConte)
P. lecontei Chaudoir - BC AB SK MB - PQ NB - NF
canadensis Casey
grauidus Darlington
ruftpes LeConte
p. tongicbrnis (Say) NT BC AB SK MB ON PQ NB NS PE - NF
americanus Dejean
P. septentrionis Dejlan* AK YK NT BC AB SK MB ON PQ - LB NF
hyperboreus Dejean
Iabradorinus Casey
minuen.; Casey
tritus Casey
P. stygicus Chaudoir AK YK NT BC AB SK MB ON PQ - LB NF

Genus DIPLOUS Motschulskv

Platidiu.s Chaudoir
D. aterrimus, (Dejean) AK YK NT BC AB
breuiceps (Casey)
coloradensis (Casey)
fulcratus (LeConte)
reflexus (Casey)
tenuitarsis (Casey)
D. californic rs (Motschulsky) - BC
breuius;ylas (Casey)
lnciszs (Casey)
Iatipenris (Casey)
rectus (Jasey)
sietan es (Casey)
strenuus (Casey)
troc han te ricus (LeConte)
D. frlicornis iCasey) - BC
D. rugicollis (Randall) PQ NB NS
longipalpis (Notman)

Genus PLAI]IDIOLUS Chaudoir

Patroboidea Van Dyke
P. vandykei Kurnakow AK YK - BC AB
rufus (Van Dyke)

Tlibe Pterostichini
Genus LOK\NDRUS LeConte
Key to Ncrth American species: Allen (1972)
L. minor (Chaudoir) - ON
inquietrc Casey
L. velocipes Casey - ON

Genus STOMIS Clairville

S. pumicatus (Panzer)t NS

Genus LOP}{OGLOSSUS LeConte

L. scrutator (LeConte) - ON PQ
canadensis (Chaudoir)

Genus POECILUS Bonelli

Feronio Latreille
Leconteus Lutshnik
(Subgenus POECILUS s. str.)
P. chalcites (Say) - ON PQ
micans Chaudoir
soyi (BruII6)

30
 Carabidae

P. corvus (LeContet AB SK MB
P. lucublandus (Say) NT BC AB SK MB ON pe NB NS pE
castanipes Kirby
conue*icollis (Say)
dilatatus LeConte
elucens Casey
fraternus Say
manhattanis (Casey)
planifer Casey
P. scitulus LeConte - BC AB SK MB
(Subgenus DERUS Motschulsky)
(D er ulus Tschitsch6rine)
P. nearcticus (Lindroth.t* NT
Genus STEREOCERUS Kirby
Boreobi a Tschitscherine
S. haematopus (Dejean)* AK YK NT BC AB SK MB ON pe - LB NF
similis I\trby
S. rubripes (Motschulsky)* AK
hry z hanou s kii (Lindroth)
Genus MYAS Dejean
Key to North American species: Allen (1980)
M. coracinus (Say) - ON
M. cyanescens Dejean - ON pe NB
foueatus LeConte

Genus PTEROSTICHUS Bonelli

(Subgenus CYLINDROCHARIS Casey)
Key to species: Barr (1971)
P. rostratus (Newman) - ON pe NB
grandiceps (LeConte)
piceatus (Casey)
sulcatulus (Casey)
(Subgenus HOLCIOPHORUS LeConte)
P. lama (M6netri6st - BC
o/er (Dejean)
at e r r imu s (Motschulsky)
cephalus (Casey)
domitor (Casey)
pollens (Casey)
uancouueri (Casey)
(Subgenus FIYPHERPES Chaudoir)
(B r ac hy s tilu s Chaudoir)
(Gonoderus Motschulsky)
(H ap locoelus Chaudoir)
P. adoxus(Say) - ON pe NB NS
rejectus LeConte
sufflatus (Casey)
P. a)gidus LeConLe AK BC
anthrat (Casey)
innatus (Casey)
responsor (Casey)
ualidus (Dejean)
P. amethystinus Mannerheim AK BC
jejunus Casey
metlakatlae Casey
nouellus Casey
stoicus (Casey)
P. castaneus (Dejean) AK BC
brunneus (Dejean)
terracensis (Casey)
wrangelli Casey
P. crenicollis LeConte AK BC
pugetanus Casey
rectilatus Casey

31
Carabidae

P. ecarinatur, Hatch BC AB
P. herculane,rs Mannerheim BC
scenicus Casey
P. neobrunneus Lindroth BC
P. nigrocoerrleus Van Dyke BC
P. protractus LeConte BC AB
fontinalis Casey
inornah.ts Bland
prouenss (Casey)
utensis t.Casey)
uiuax (Casey)
P. tristis (De"iean) ON PQ NB NS
interfector (Newman)
subarcuztus LeConte
sustentu s LeConte
tetriculc Casey
zephyru; Casey
(Subgenus L.DPTOFERONIA Casey)
Key to speries: Hacker (1968)
P. pumilus Casey BC
longicol'.is (Casey)
longicol'.is LeConte
oregonui (Chaudoir)
oregonis Cslki
P. sphodrinus LeConte BC AB
idahoanus (Casey)
(Subgenus M ONOFERONIA Casey)
P. diligendus (Chaudoir) PQ
apalachius Horn
osculan:, Casey
(Subgenus G A.STRELLARIUS Casey)
P. honestus (Say) ON PQ NB NS
america ous (Castelnau)
atronite * (Casey)
deficien.t (Casey)
fastiditu s (Dejean)
scolopateus (Casey)
(Subgenus S'IEROPUS Dejean)
(Refonic: Casey)
(.Sterode rus Motschulsky)
P. circulosus Lindroth AK YK
(Subgenus B')THRIOPTERUS Chaudoir)
(Dysidit.s Chaudoir)
(.Pararg;ttor Casey')
P. adstrictus Eschscholtz+ AK YK NT BC AB SK MB ON PQ NB NS PE LB NF
angusticollis (Casey)
commixius Chaudoir
latebricola (Casey)
latescans (Casey)
laxicolli.; (Casey)
luczotii .Dejean)
maehlini LeConte
oblong ius c ulus (Motschul sky)
oblongotzotatus (Say)
ob t u sa n,7 ulu s (Motschulsky)
saxatilis (Casey)
sericeus (Casey)
seriepun ctatus Mannerheim
sex,p unc :atus (Mannerheim)
shastan.rs (Casey)
P. lustrans L,eConte BC
atrolucens (Casey)
p uncticollis (Motschulsky)
P. mutus (Say) BC MB ON PQ NB NS PE NF
carbonarius (Dejean)
egens (Casey)

32
 Carabidae

rnlorosus (Dejean)
picicornis (Kirby)
politus (Harris)
puluinatus Hausen
stenopus Hausen
P. oregonus LeConte BC
colligatus (Walker)
P. pensylvanicus LeConte NT BC AB SK MB ON PQ NB NS PE NF
(Subgenus MORPHNOSOMA Lutshnik)
(Euferonia Casey)
(Omaseidius Jeannel)
P. coracinus (Newman) oN PQ NB NS PE LB NF
adjunctus LeConte
erebeus Casey
flebilis LeConte
lacustris (Casey)
ludibundus (Casey)
moerens (Newrnan)
(?) monedulus (Newman)
roanicus (Casey)
rufitarsis (Nicolay & Weiss)
strigosulus (Casey)
uenator (Casey)
uashingtonensis (Nicolay & Weiss)
P. Iachrymosus (Newman) ON PQ NB
P. melanarius (IlligerX _BCAB SKMBONPQNBNSPE-NF
P. novus Straneo -MBONPQ
P. relictus (Newman) ON
protensus LeConte
P. stygicus (Say) oN PQ
bisigillatus (Harris)
(?) picipes (Newman)
probus Casey
quadrifer (Casey)
rugicollis (Haldeman)
subaequalis (Casey)
umbonatus (Casey)
uapidus Casey
(Subgenus ABACIDUS LeConte)
(Peristethus LeConte)
P. atratus (Newman) ON
P. permundus (Say) ON
planifer (Casey)
(Subgenus MELANIUS Bonelli)
(Ly p ero som us Motschulsky)
(Lyperus Chaudoir)
(M e tame I ani us Tschitsch6rine )
(Omaseus Dejean)
P. castor Goulet & Bousquet ON PQ
P. caudicalis (Say) NT BC AB SK MB ON PQ NF
agrestis Bland
breuibasis (Casey)
P. corvinus (Dejean) NT BC AB SK MB ON PQ NB
aequalis (Casey)
subpunctatus (Harris)
tenebrosus (Chaudoir)
P. ebeninus (Dejean) ON
acutangulus (Chaudoir)
(Subgenus PSEUDOMASEUS Chaudoir)
P. luctuosus (Dejean) BC AB SK MB ON PQ NB NS PE NF
abjectus LeConte
confluens (Casey)
hamatus (Harris)
P. tenuis (Casey) AB MB ON PQ NB NS PE NF
testaceus (Casey)
(Subgenus LAGARUS Chaudoir)
(Pseudargutor Casey)
(P se udol ag ar us Lutshnik)

33
Carabidae

P. Ieconteianus (Lutshnik) BC SK MB ON PQ NB NS
erythropus (Dejean)
nitidus iKirby)
(Subgenus ARGUTOR Dejean)
(Americ cmasezzs Csiki)
(Micronnseus Casey)
(.O ma se :rlu s Lutshnik)

P. corruscultLs LeConte ON PQ
P. femoralis iKirby) AB SK MB oN PQ
aequicoi.Iis (Casey)
desidiosus LeConte
P. patruelis (Dejean) AK BC AB SK MB ON PQ NB NS PE LB NF
bicolor (Kitby)
linearis (Mannerheim)
P. strenuus (Panzer)f BC NF
(Subgenus CRYOBIUS Chaudoir)
Key to No th American species: Ball (1966)
P. arcticola (Chaudoir) AK YK NT MB PQ LB
I ab r ad.a rens is ( Chaudoir)
P. auriga Ball AK
P. barryorun Ball AK NT AB
P. biocryus liall AK
P. brevicornis (Kirby)* AK YK NT BC nfg ON P6-LB NT
delicatus (Casey)
ep ipl e ur alis (J. Sahlberg)
fa st idiosus (Mannerheim)
P. bryanti (\ian Dyke) NT
P. bryantoid:s Ball AK
P. cacumenir; Ball AI{
P. caribou Brrll AK YK NT _MB
P. chipewyarr Ball NT BC AB _MBON
P. empetricola (Dejean)* AKYK_BC
globicollis Csikt
r ot undi :olli s ( Mannerheim)
r ufi coll,:. s (Mannerheim)
P. gerstlensis Ball AK
P. haftorni I.,indroth AK
P. hudsonicr:.s LeConte AK YK NT BC
P. kotzebuei Ball AK
P. mandibulrryoides Ball AK YK NT BC
P. nivalis {R F. Sahlbergt+ AK
P. parasimil:s BaIl* AK YK
pl:)
P. pinguedineus (Eschscholtz)x AK YK I'ff - - rg
alaskensis (Poppius)
beringi (Casey)
diplogrnus (Chaudoir)
holmbergi (Poppius)
incogni;lus (Poppius)
mandibularis (Kirby)
montanellus Poppius
splendidus (J. Sahlberg)
s ub nititlul u s (Poppius)

s ub s inuos us (Chaudoir)
washingtoni (Casey)
P. planus (J. Sahlberg) AK
blaisdeili Van Dyke
P. riparius ( Dejean) AK YK NT BC AB
breuiusculus (Casey)
caseyi Csiki
/ofuus (Mannerheim)
fus coae,zeus ( Chaudoir)
laeuilatus Notman
patulus (Casey)
P. similis Mannerheim* AK
q uadr icollis ( Mannerheim)
P. soperi BalI AK NT
P. stantonersis BaIl NT

34
 Carabidae

P. surgens LeConte AB
P. tareumiut Ballx AK YK NT
P. tiliaceoradix Ball AK
P. ventricosus tEschscholtz)x AI{ YK NT
aleutorum (Lutshnik)
b reu icoll is (Tschitsch6rine)
hy perboreus (Mannerheim)
otariidinus (Casey)
r ufi sca p us (Mannerheim)
r ug ifer us ( Tschitsch6rine)
s ubcaudatus ( Mannerheim)
s ubexarat us (Mannerheim)
u indic atus (Mannerheim)

(Subgenus LYPEROPHERUS Motschulsky)

Key to species: Budarin (1976)
P. agonus Horn* AK YK NT
P. costatus (Men6tri6s)* AK YK NT
P. punctatissimus (Randall) - YK NT BC AB SK MB ON PQ NB NS - LB NF
P. sublaevis (J. Sahlberg)* AI{ YK NT
rufofemoralis Van Dyke
P. vermiculosus (M6n6tri6s)x AK YK NT
innuitorum (W.J. Brown)

Genus CYCLOTRACHELUS Chaudoir

Key to species: Freitag (1969)
(Subgenus EVARTHRUS LeConte)
C. sodalis sodalis (LeConte) - ON
corax (LeConte)
fatuus (LeConte)
inflatipennis (Casey)
I icinoide s ( Motschul sky)
retractus (Casey)
sulcatus (Casey)
uagans (.LeConte')

Genus ABAX Bonelli

A. parallelepipedus NS NF
(Piller & Mitterpacher)f
aler (Villers)
s/riolo (Fabricius)

Tribe Zabrini
Genus DISAMARA Lindroth
Pseud,amara Lindrotn
D. arenaria (LeConte) _ONPQNBNSPE
Genus AMARA Bonelli
(Subgenus CURTONOTUS Stephens)
(Leirus Zimmermann)
A. alpina (Paykull)* AKYKNTBCAB-MBONPQ-LBNF
alaskana Cslki
angustata J. Sahlberg
arguta (Casey)
borealis (Chaudoir)
brunnipennis Dejean
deficiens (Casey)
inanis (Casey)
obtusa LeConte
rubripennis (Casey)
subtilis (Casey)
A. aulica (Panzer)t
A. blanchardi Hay'ward ___sc____PQIUNS__NF
A. bokori Csiki* AK YK NT BC MB
sahlbergi Poppius
A. carinata (LeOonte) _BCABSKMBONPQNB

35
Carabidae

adstricta (Putzeys)
catenulata (Casey)
giluipes (Casey)
laticollis (LeConte)
A. hyperborea Dejean* AK YK NT BC AB SK MB ON PQ LB NF
canadensis (Putzeys)
elongatr.r (LeConte)
imperfe':ta (W.J. Brown)
tristis (Prtzeys)
A. lacustris l,eConte AK YK NT BC AB SK MB ON NS
manitobensis (Casey)
A. pennsylva-nica Hayward -oN PQ NS
fuluipes (Putzeys)
A. thoracica Hayward BC AB SK
A. torrida (Panzer)* AK YK NT BC AB SK MB ON PQ Ne NS pn LB NF
albertana (Casey)
biarcuala (Casey)
breuilat'ris (Kirby)
breuipennis (Casey)
consueta Fall
cylindrica LeConte
dura (Casey)
holnrbergi (Putzeys)
hudsonica Hayward
labradarensis (Casey)
melanollastrica Dejean
reflexa rPrtzeys)
rufimat a (Kirby)
scrutat( (Casey)
somnoh nta (Putzeys)
(Subgenus B RADYTUS Stephens)
(P seuda brady tzs Csiki)
A. apricaria Paykull)t BC AB SK MB ON PQ NB NS LB NF
putzeystana Csiki
putzeysti Hotn
A. avida (Sa d BC SK MB ON PQ NB NS PE NF
confinis Dejean
A. browni Li rdroth -YK NT
A. colvillensi s Lindroth
A. exarata Dejean
AK NT -ms-- ON
curta ((:asey')
furtiua;Jay
grauida (Casey)
stygiali:; (Casey)
A. fulva (O.F. Mriller)i _PQ NB NS PE LB NF
A. glacialis ( Mannerheim)* AK YK Nr sc SK oN PQ _LB
nainensis Casey
A. Iatior (Kirby) _BC SK MB oN PQ NB NS NF
decepta (Casey)
humphreysi (Casey)
laeuistri.ata (Putzeys)
Iibera LeConte
oregona LeConte
relicta (Casey)
A. schwarzi.{ayward YK NT AB MB ON PQ
septentrionalis (LeConte)
(Subgenus PERCOSIA Zimmermann)
Key to species: Hieke (1978)
A. obesa (Say) YK NT BC SK MB ON PQ NB NS PE NF
dffinis (LeConte)
extensa (Casey)
latissima (Casey)
sulcatuta (Casey)
uentricosa (Casey)
(Subgenus CELIA Zimmermann)
(.Isopleurus l{rtby)
A. aurata De;jean
euanida (Casey)

36
 Carabidae

fragilis (Casey)
gouanensis (Casey)
irnitatrix Horn
proba (Casey)
A. bifrons (Gyllenhal)t PQ NB NS PE LB NF
A. brunnea (Gyllenhal)* AK
am pl ico ll is (Mannerheim)
A. californica Dejean BC
mericana Dejean
perspecta Casey
pugetana (Casey)
A. chalcea Dejean ON
coruina (Casey)
maneei (Casey)
pinorunr (Casey)
schotti (Casey)
sodalis (Casey)
A. discors Kirby AK YK NT BC AB SK MB ON PQ NS LB NF
gibba (LeConte)
lugubris (Casey)
merula (Casey)
neuadica (Casey)
obsolescens (Casey)
paganica (Casey)
paroudna (Casey)
scolopax (Casey)
spaldingi (Casey)
sphaerops (Casey)
wahelandi Hatcft
A. hicksi Lindroth* NT
A. idahoana (Casey) _BC AB
exlineae Minsk & Hatch
A. musculis (Say) _BC AB SK MB ON PQ I\D
breuitarsis (Casey)
contempta (LeConte)
crenulata (Casey)
curticeps (Casey)
fluminea (Casey)
limbalis (Casey)
lyncea (Casey)
minnesotana (Casey)
nugator (Casey)
paulula (Casey)
scintilla (Casey)
shoemaheri (Casey)
uegrandis (Casey)
A. pseudobrunnea Lindroth AKl'K-BCABSKMB-PQ LB
A. quenseli (Schbnherr)x AK YK NT BC AB SK MB ON PQ NB NS pe LB NF
brumalis (Casey)
brunnescens (Casey)
ceruicalis (Casey)
defecta (Casey)
definita (Casey)
docilis (Casey)
eldorensis (Casey)
erplanatula (Casey)
exposita (Casey)
femoralis Hotn
fontinalis (Casey)
hornis Cslki
i ndi st inc ta (Mannerheim)
laxicollis (Casey)
mimica (Casey)
obligata (Casey)
piperi (.Casey)
relucens (Mannerheim)
remota (Zimmermann)
remotestriata Dejean
tahomae (Casey)
washoeana (Casey)
A. rubrica Haldeman ON PQ

37
Carabidae

ferruginea Casey
liquida (Casey)
lubrica (Casey)
lucina (Oasey')
nigripennis (Casey)
pallida (Casey)
piceonitt:ns (Casey)
politissitna (Casey)
proditor (Casey)
A. sinuosa (Casey) AK YK NT BC AB SK MB ON PQ NB PE LB NF
elusa (Casey)
hospes (Casey)
nitida (Yitby)
nupta (Casey)
subaenett (LeConte)
subaenescens Csiki
thoracicu (Casey)
A. spuria Lindroth AK NT
(Subgenus AIIARA s. str.)
A. aenea (DeGeer)f oN PQ NB NS PE NF
deuincta Casey
A. aeneopolit.r Casey NT_ABSKMB oN PQ NB_ LB NF
A. coelebs Hayward _BCABSKMB
leuiceps Jasey
nebrask<,.na Casey
oblonguta Casey
rustica ()asey
A. communis (Panzer)t
A. conflata Lt:Conte BC Iu_
diffidenr Casey
leydeni l.)asey
semota (Jasey)
A. confusa Lt Conte AK YK BC SK MB PQ
acomant'. Casey
ebenina Casey
impeditu Casey
oblongifrrmis Casey
protenso Putzeys
subarctica Casey
subpunctata LeConte
uiridula Casey
A. convexa L+:Conte BC AB SK MB ON PQ
breuiforrnis Casey
cockerell,i Casey
frugalis (Casey)
lecontei,Jsiki
ouiformis Casey
piceola (.Jasey
polita LeConte
wingatet Casey
A. cupreolata Putzeys NT BC SK MB ON PQ NB NS PE
eneruis Oasey
haywarc!.i Csiki
inflaticol.lis Casey
neoscotica Casey
paruiceps Hayward
tenax Ctrsey
A. ellipsis (C;rsey) BC AB SK MB ON PQ
w innipe;1en sis (Casey)
A. emancipata Lindroth BC
A.
A.
erratica (Duftschmid)x
eurynota (Panzer)t
AK NT BC -s^ Ite - pe -iuNr NF
A. familiaris (Duftschmid)t AK BC AB -oN PQ NB NS PE_NF
humilis Casey
A. farcta LeConte YK BC AB SK MB
albertae (Casey)
formalis (Casey)
funebris (Casey)
margino.tella (Casey)
olympia (Casey)

38
 Carabidae

parallela (Casey)
sho.stanica (Casey)
solita (casey)
subdepressa (Casey)
subsimilis (Casey)
uancouueri (Casey)
A. impuncticollis (Say) SK MB ON PQ NB NS PE NF
(? )hracina Haldeman
a nt
arcuata Casey
dfficilis LeConte
edor Notman
otiosa Casey
wadei Casey
A. interstitialis Dejean* AK YK NT
A. Iaevipennis Kirby AK_NTBCABSK ms oN PQ NB NS PE NF
aeneolucens Casey
A. littoralis Mannerheim AK YK NT BC AB SK MB ON PQ NB NS PE NF
conuergens Casey
fallaxLeConte
hesperia Casey
inepta LeConte
heeni Casey
lacustrina Casey
laurana Casey
microcephala Casey
mystica Casey
oodiformis Casey
pullmani Casey
recticollis Casey
(?) teres Notman
turbata Casey
A. iunicollis Schiddtex AK YK BC AB SK MB ON PQ NB NS LB NF
carriana Casey
marquettensis Casey
A. patruelis Dejean* AK YK NT BC AB SK MB ON PQ NB NS PE NF
columbiana (Casey)
inaequalis Kirby
reducta (Casey)
splendida Haldeman
A. sanjuanensis Hatch BC
(Subgenus ZEZEA Csiki)
(Pseudotriaena Minsk & Hatch)
(Triaena LeConte)
A. angustata (Say) SK MB ON PQ
flebilis (Casey)
indistincta Haldeman
A. Iongula LeConte BC
afoueolata Hayward
alaxnoguia Minsk & Hatch
atrichata Minsk & Hatch
glabrata Minsk & Hatch
profuga (Casey)
A. pallipes Kirby BC AB SK MB ON PQ NB NS
depressa (LeConte)
h,incaidi Minsk & Hatch
lawrenciana (Casey)
parallela (Casey)
sherrnani (Casey)
A. scitula Zimmermann BC AB SK MB
irregularis (Casey)
obliqua Casey
prouana Casey
uinta (Casey)
uapida (.Casey)

Tribe Harpalini
Genus CRATACANTHUS Dejean
C. dubius (Palisot de Beauvois) AB SK
bisectus Casey

39
Carabidae

litoreus Jasey
pensylucinicus Dejean

Genus PIOS0MA LeConte

P. setosum L-'Conte - BC AB SK
breuipenne Casey

Genus EURYDERUS LeConte

Nothopus LeConte
Key to species: Bail (1960)
E. grossus (S,ryt - BC AB SK MB ON
arizonic,ts (Casey)
obtusus .Casey)
priuatus (Casey)
ualens ((.)asey)
zabroides LeConte

Genus HARF ALOBRACIIYS Tschitsch6rine

H. leiroides t vlotschulskyrx AK - NT - A-B

Genus HARPAI-US Latreille

(Subgenus OPHONUS Dejean)
H. puncticeps (Stephensti - ON PQ - NS
H. rufibarbis t Fabricius rt PQ
(Subgenus PSiEUDOPHONUS Motschulsky)
Key to Norbh American species: BaIl and Anderson (1962)
H. bicolor tFz bricius t MB ON PQ - NS
admissu; Casey
compar .,eConte
excubant: Casey
feroculut Casey
nactus Casey
H. ery.thropu r Dejean MB ON PQ - NS
abstrusus Casey
cupiens t)asey
deludent Casey
effetus Casey
fenisex Casey
rufopicet.ts Casey
H. faunus Sa.r MB ON PQ
badius Ltejean
conuiuus LeConte
H. longicollis LeConte - ON PQ
dolosus ()asey
haldemoni Casey
latescanr Casey
uagans L,eConte
H. paralus Cusey AB SK
spaldingi Casey
H. pennsylva:ricus (DeGeer) - BC - SK MB ON PQ NB NS PE
inmixtus Casey
longior \:Jtby
mormoni.cus Casey
H. rufrpes (DeGeer)t PQ NB NS PE - NF
(Subgenus M IGAPANGUS Casey)
H. caliginosu; tFabricius) MB ON PQ
dux Casey
(Subgenus PI,ECTRAI-IDUS Casey)
Key to species: El-Moursy (1958)
H. erraticus Siay MB ON PQ
caudalis Casey
(Subgenus PIIARAJ-US Casey)
H. indianus C siki - ON
testaceus LeConte

40
 Carabidae

(Subgenus IIARPAI-US s. str.)

H. affinis (Schranktt -BCABONPQNBNSPELBNF
aenescens Casey
oenezs (Fabricius)
assimilis Dejean
canonicus Casey
conuictor Casey
lustralis Casey
uiridiaeneus Palisot de Beauvois
uiridis Say
(Subgenus HARPALOMERUS Casey)
H. amputatus Say AK YK NT BC AB SK MB
ancillaris Casey
bracatus Casey
cuculus Casey
cupreolatus Casey
papagonalis Casey
rotundicollis I\ttby
stephensi l\ttby
transuersus Casey
(Subgenus EUHARPALOPS Casey)
H. animosus Casey _BCABSK
montanus LeConte
montuosus Csiki
H. egregius Casey AK YK NT BC AB SK MB ON PQ _ NS PE LB NF
alienus LeConte
baergi Csiki
cascadiensis Hatch
instructus Casey
rufimanus LeConte
H. fraternus LeConte
nimius Casey
-BCAB
occidenlalis Chaudoir
praestans Casey
wadei (Casey)
H. funerarius Csiki AB SK MB ON PQ
funestus LeConte
reuersus Casey
H. hoppingi Lindroth _BC
H. laticeps LeConte AK YK NT BC AB SK MB ON PQ NB NS - LB NF
solutus Casey
H. lewisii LeConte NT-ABSKMBONPQNBNS
aesopus Casey
H. viduus LeConte _ONPQ
prouidens Casey
$
H. indigens Casey - ON PQ NB
beatulus Casey
piperi (Casey)
H. ventralis LeConte AB SK MB
electus Casey
$

H. fulvilabris Mannerheim AK YK NT BC AB SK MB ON PQ NB NS PE LB NF
H. megacephalus LeConte MB ON
(Subgenus CORDOHARPALUS Hatch)
H. cordifer Notman AK BC
w ashingtoniensls Van Dyke

$
H. carbonatus LeConte NT BC AB SK
H. fallax LeConte - ON PQ NB NS
amiculus Casey
recisus Casey
H. herbivagus Say - BC AB SK MB ON PQ NB NS PE
ossenszs Casey

4L
Carabidae

foueicollis LeConte
mutabilis (Haldeman)
H. intactus Casey _BC
uintanus Casey
H. nigritarsis C.R. Sahlbergx AK YK NT BC AB MB ON PQ LB NF
curtatus Mannerhein
proximus LeConte
recensus Casey
H. pleuriticus Kirby AK BC AB SK MB ON PQ NB NS PE LB NF
aequabilis Casey
lasciuus tJasey
liuidulus Casey
niualis Casey
perspicuus Casey
placidus Casey
pumilio Casey
H. seclusus Casey -BCAB
parowanus Casey
H. solitaris D,:jean NT BC AB SK MB ON PQ LB NF
fuliginos zs (Duftschmid)
opicus Ctl.sey
H. somnulentrs Dejean BC
blanditu; Casey
contactut'Casey
pacificus FalI
H. uteanus Cr,Lsey AB SK MB
oppositut Casey
A
x
H. alaskensis Lindroth AK
H. innocuus LeConte NT sc AB oN pe
fugitans Casey
intactus Jasey
persolus Casey
$

H. cautus Dej:an BC
aduena I eConte
(?) alb ior, icus Mannerheim
crenatell n Casey
columbianus Casey
(?) defix.us Walker
oregonensis Casey
H. ellipsis Le(lonte
H.
H.
obnixus CzLsey
ochropus FJrby
---scTsK-------
AK BC AB SK MB ON PQ
H. opacipennis (Haldeman) AK YK NT BC AB SK MB ON PQ
lacustris Casey
leuiceps Oasey
oodioides Chaudoir
H. plenalis Casey NT BC MB ON PQ NB NS NF
latebricoi,a Casey
modulatus Casey
$
H. desertus LeConte BC AB SK
captiosu:t Casey
couleensts Hatch
dulciculus Casey
lucidus LeConte
lustrans Casey
metuens Casey
uaciuus Oasey
R
Y

H. fulgens Csiki ON
nitidulus Chaudoir

Genus IIARP ALELLUS Lindroth

H. basilaris (Kirby) AK YK NT BC AB SK MB ON PQ NS
durescans (Casey)

42
 Carabidae

er.tensus (Walker)
furuiculus (Casey)
obesulus (LeConte)
oblongus (Casey)
renoicus (Casey)
sejunctus (Casey)
stocktonensis (Casey)
subenormis (Casey)
uaricornis (LeConte)
uentricosus (Casey)

Genus TRICHOTICHNUS Morawitz

(Subgenus TRICHOTICHNUS s. str.)
(Episcopellus Casey)
(Pteropalus Casey)
T. autumnalis (Say) ON
nitescens (Casey)
T. dichrous (Dejean) ON
fluuialis (Casey)
uersutulus (Casey)
T. vulpeculus (Say) ON PQ
nigripennis (Dejean)

Genus SELENOPHORUS Dejean

(Subgenus SELENOPHORUS s. str.)
(Gynandropus Dejean)
(Hemisopalus Casey)
S. gagatinus Dejean ON PQ NB
m(turus Haldeman
uiridescens LeConte
S. hylacis (Say) ON
americanus (Dejean)
S. opalinus (LeConte) oN PQ
iripennis LeConte
S. pedicularius Dejean ON
aereus LeConte
puellus Putzeys
troglodjttes Dejean
S. planipennis LeConte BC AB SK MB ON
( Subgenus CELIAMORPHUS Casey)
S. ellipticus Dejean ON
currens (Casey)
opaculus (Casey)
oualis Dejean

Genus DISCODERUS LeConte

Selenalius Casey
D. parallelus (Haldeman) BC ON
amer icanus (Motschulsky)
gener Casey
hesperus Casey
paruuliceps Casey

Genus GEOPINUS LeConte

G. incrassatus (Dejean) AB SK MB ON PQ
fluuiaticus Casey
Genus ANISODACTYLUS Dejean
(Subgenus ANADAPTUS Casey)
A. discoideus Dejean
A. nivalis Horn
sK yu oN PQ
BC
paruulus (Casey)
A. pitychrous LeConte _ABSKMB
A. sanctaecrucis (Fabricius) BC AB SK MB ON PQ NB lls PE
baltimoriensis (Say)
uteanus (Casey)

43
Carabidae

A. viridescenr LeConte
chalceus LeConte
idahoensis (Hatch)
Iecontei Gemminger & Harold
(Subgenus G\T{ANDROTARSUS LaFert6-S6nectbre)
(Triplect:us LeConte)
Key to Norl;h American species: Noonan (1973)
A. dulcicollis LaFert6-S6nectdre) ON
ellipticu: LeConte
elongatu.; Chaudoir
modicus (Casey)
A. merula Ge'mar MB ON PQ
aethiops (Casey)
crassus LeConte
grauidus LeConte
hetnpi (Casey)
margina.lus (Casey)
pinguis l.,eConte
sulcipennis (Casey)
uolcotti iCasey)
A. ovularis {Casey}
semirubilus (Casey)
-oN
A. rusticus t S ry r MB ON PQ NB NS PE
oblongus (Casey)
lrjslls Dr jean
(Subgenus AIIISODACTYLUS s. str.)
(.Cephaloqyna Casey)
A. binotatus ( Fabricius tt _BC
A. californicu r Dejean _BC
aleneanus Casey
angustus Casey
comes Ct.sey
confusus LeConte
humeral,s Casey
maestus Casey
obsolescens Casey
oregonus Casey
paganict s Casey
sinuatus Casey
A. carbonariu s t Say t oN PQ
breuior (t laseY)
luctuosu,:; Dejean
rufipenn,'.s LeConte
A. furvus LeC onte -oN
MB ON NS-
A. harrisii Le'lonte BC AB SK PQ
Iacertosus Casey
A. kirbyi Lincroth BC MB ON PQ NB NS
(?) Iecontei Chaudoir
A. melanopus (Haldeman) ON
A. nigerrimus {Dejean) oN PQ NN NS NF
Iaticollis (Kirby)
opacus (()asey)
punctulatus LeOonte
A. nigrita Dejean BC AB SK MB ON PQ NB NS PE
inte rp un.:t at u s (Kirby)
A. similis Le(lonte BC
incertus Jasey
incisus C. asey
punclico,lis Chaudoir
semipunctdtus LeConte
sericatus Casey
solidus Casey
(Subgenus AI'LOCENTRUS LeConte)
A. amaroides LeConte BC
(Subgenus PS,EUDAPLOCENTRUS Noonan)
A. laetus Dejr an ON
gemmeus (Casey)

44
 Carabidae

(Subgenus SPONGOPUS LeConte)

A. verticalis (LeConte) SK MB ON PQ

Genus AMPHASIA Newman

(Subgenus AMPI{ASIA s. str.)
A. interstitialis { Say) ON PQ
fuluicollis Newman
mollis Casey
obscuripennis (Dejean)
(Subgenus PSEUDAMPHASIA Casey)
A. sericea (Harris) SK oN PQ
femorata (Dejean)

Genus DICHEIRUS Mannerheim

Key to species: Noonan (1968)
D. piceus (Menetriest BC
alutaceus Casey
angustulus Casey
australinus Casey
decoloratus Casey
facilis Casey
incidens Casey
iteg ul ar is Motschulsky
insularis Casey
parallelus LeConte
rupimontis Casey
sodalis Casey
uillosus Motschulsky

Genus XESTONOTUS LeConte

X. lugubris tDejeant oN PQ NS
manhattanis (Casey)
tioganus (Casey)

Genus NOTIOBIA Perty

(Subgenus ANISOTARSUS Chaudoir)
(Diaphoromerus Chaudoir)
(Eurytrichus LeConte)
(Stilbolidus Casey)
Key to North American species: Noonan (1973)
N. nitidipenni s (LeConte) oN PQ
agitabilis (Casey)
cephala (Casey)
conspectd (Casey)
delicata (Casey)
tenuitarsis (Casey)
N. picea tLeConte) oN PQ
soyi (Blatchley)
N. terminata tSayl ON PQ NB NS
agilis (Dejean)
angusticollis (Casey)
conniuens (Casey)
florid,ana (Casey)
foueicollis (Bates)
fuscipennis (Casey)
hebes (Casey)
inerrans (Casey)
ocreata (Say)
parallela (Casey)
similis (Say)
suboualis (Casey)
subuirens (Casey)
testacea (Haldeman)
uernicata (Casey)

45
Carabidae

Genus TRIC JOCELLUS Ganglbauer

(Subgenus T RICHOCELLUS s. str.)
T. cognatus (Gyllenhal)x AK YX NT BC AB SK MB ON PQ NB NS PE LB NF
axillar i s (Mannerheim)
boreellus Casey
conflag r at us ( Mannerheim)
lateralis Casey
longius<. ul us (Mannerheim)
montico.ta Casey
nitens (l,eConte)
punctipr,:nnis Casey
ruficrus (Kirby)
(Subgenus REOXENUS Tschitsch6rine)
O

T. mannerheimii (R.F. Sahlberg)* AK YK NT BC MB - PQ

porsildi W.J. Brown

Genus AMEIIINUS Casey

A. linearis (I.eConte) - ON

Genus BRAITYCELLUS Erichson

(Subgenus L OCELLUS Motschulsky)
(Glyceri n Casey)
B. politus tF:rllt BC
(Subgenus B RADYCELLUS s. str.)
B. harpalinu; tAudinet-Serviller* - BC
(Subgenus C \THARELLUS Casey)
B. lecontei C;iki AK YK NT BC AB SK MB ON PQ NB NS PE LB NF
cordicol'is (LeConte)
(Subgenus S' IENOCELLUS Casey)
B. congener tl,eContet - BC AB SK MB ON PQ
alutacet.s (Casey)
B. nigerrimu; Lindroth AB SK
B. nubifer Lt Conte - BC AB
nubicollis (Casey)
uentrali; LeConte
B. rupestris r Say) - ON PQ NB NS
antennolis (Casey)
(.?) cinct;rs (Say)
(?) debilipes (Say)
elong att":lus (Dej ean)
flauipes (Kirby)
parallel,^rs Chaudoir
A

B. insulsus (Oasey)
B. neglectus il,eConte) _ -oN
BC AB SK MB ON PQ NB NS PE _ NF
B. nigriceps LeConte MB ON PQ
occultus (Casey)
$

B. californicr.s (LeContet - BC
B. tantillus (Dejean) - ON
dfficilis (Dejean)
(Subgenus LIOCELLUS Tschitsch6rine)
B. nigrinus (Dejean) AK BC AB SK MB ON PQ NB NS PE - NF
quadricollis (LeConte)
tibi,alis (Kirby)
B. semipubescens Lindroth AB ON PQ NB
(Subgenus TRILIARTHRUS Casey)
B. atrimedeus (Say) MB ON PQ NB
B. badipennis (Haldeman) - ON PQ
curticoliis (Casey)
properus (Casey)

46
 Carabidae

B. conformis (Fall) BC
tetricus (Casey)
B. georgei Lindroth AB
B. kirbyi (Horn) oN PQ
B. lugubris (LeConte) AB _MBONPQ NB NS PE NF
frosti (Fall)
protractus (Casey)

Genus STENOLOPHUS Stephens

(Subgenus STENOLOPHUS s. str.)
S. anceps LeConte BC
fidelis Casey
rolu nd i colli s Motschul sky
S. carbonarius (Dejean) ON NS
S. fuliginosus Dejean BC AB SK MB ON PQ NU NS PE N"
fuscipennis (LeConte)
uersicolor Kirby
S. fuscatus Dejean ON PQ
S.
S.
humidus Harnilton
incultus Casey BC
ON PQ_ fr_ _ _
consors Casey
debiliceps Casey
S. limbalis LeConte BC
indistinctus Motschulsky
longitarsis Casey
S. megacephalus Lindroth oN PQ
S. ochropezus (Say) _MBONPQ NB NS
conuexicollis LeConte
floridanus Casey
(?) gracilis Casey
testaceicollis Casey
S. plebejus Dejean ON PQ
(?) lugubris (Haldeman)
riuularis Casey
(Subgenus AGONODERUS Dej ean)
S. comma (Fabricius) BC SK MB ON PQ NB NS PE NF
dorsalis (LeConte)
gracilitarsis Casey
obliqulus Casey
similis (I\trby)
S. lecontei (Chaudoir) ON PQ
plagiatus Casey
S. lineola (Fabricius) AB SK MB ON PQ
furcatus (Fabricius)
s
S. conjunctus (Say) BC SK MB ON PQ NB NS PE
captiosus Casey
humilis (Dejean)
immunis (Kirby)
misellus (Dejean)
moquinus Casey
thoracicus Casey
S. rotundatus LeConte -oN pe-
S. rotundicollis (Haldeman)
incitatus Casey
-oN
scitulus Casey

Genus ACUPAJ-PUS Latreille

(Subgenus PHILODES LeConte)
(Goniolophus Casey)
A. alternans (LeConte) oN PQ
testaceus (LeConte)
A. rectangulus Chaudoir oN PQ
(Subgenus ACUPALPUS s. str.)
A. canadensis Casey
A. carus (LeConte) _ __ NC_:"MB3Ni3NBN3Ff;_ Nil

47
Carabidae

curlipet nis Casey

expertus Casey
subrectu,.s Casey
triuialis Casey
A. meridiant s ll-inn6t+ - BC AB - PQ
A. nanellus (lasey - ON PQ NB NS
A. pumilus I indroth - ON PQ - NS
(Subgenus T A.CHISTODES Casey)
A. indistinct rs Dejean - ON
obscurus (Casey)
A. partiariu: tSayt ON PQ
conuerg tns (Casey)
Iyratus Casey')
A. pauperculus Dejean ON PQ NB NS PE
consimi'.is Dejean
A. testaceus Dejean - ON PQ
fusciceps (Casey)
micros LeConte

Tlibe Licinini
Key to North American species: Ball (1959)
Genus DIPL ]CHEILA Brull6
Rembus Dejean
(Subgenus If IOREMBUS Jeannel)
D. assimilis rl-eConter ON PQ
modesta Casey
D. impressic',llis rDejeanr ON PQ
cliens C*ey
foueata Jasey
laticollis (LeConte)
planulala Casey
D. major ma. or (Leconte ) - ON
expansa Casey
oblonga Casey
procera Casey
D. obtusa (LeConte) NT BC AB SK - ON PQ NB NS PE
parallelt Casey
D. oregona tllatcht BC AB SK MB
D. striatopurLctata (LeConte) NT BC AB SK MB ON PQ
alternar.s Casey
amplipennis Casey
breuicoli,is Casey
D. undulata larr AB - MB

Genus DICAELUS Bonelh

(Subgenus P.ARADICA-ELUS Ball)
D. dilatatus lilatatus Say PQ
carolinensis Casey
dejeanii Dejean & Boisduval
planicollis LeConte
D. elongatus Bonelli - ON PQ
osieuill rnsis Casey
debilice;ts Casey
obscuru.; LeConte
simplex Dejean
D. politus Dt jean - ON PQ
angustus Casey
leonardli Hatis
D. sculptilis ntricatus LeConte - ON
ocellatus Blatchley
D. sculptilis rpioides Ball - SK MB ON
D. teter Bonelli ON PQ
ouipenn'.s Casey
(Subgenus DICAELUS s. str.)
D. purpuratus purpuratus Bonelli * ON
uiolaceus Bonelli

48
 Carabidae

(Subgenus LIODICAELUS Casey)

D. laevipennis laevipennis LeOonte AB
eudnescens (Casey)

Genus BADISTER Clairville

(Subgenus BADISTER s. str.)
B. lernrgineus Dejean - BC
anthracinus LeConte
B. neopulchellus Lindroth NT BC AB SK MB ON PQ - NS PE
B. notatus Haldeman - ON PQ
angustus Casey
giluip.es Casey
terminalis LeConte
B. obtusus LeConte NT - AB SK MB ON PQ - NS
B. pulchellus LeConte - ON
(Subgenus BAUDIA Ragusa)
B. grandiceps Casey - BC AB - MB ON PQ - NS
B. transversus Casey MB ON PQ
B. ocularis Casey - ON PQ NB NS PE
B. parviceps Ball - BC MB ON PQ
B. reflexus LeConte - ON
seclusus Blatchley

Tribe Oodini
Genus OODES Bonelli
O. amaroides Dejean - ON
O. americanus Dejean - ON PQ
fluuialis LeConte
O. brevis Lindroth - ON

Genus LACHNOCREPIS LeConte

L. parallela tSayt MB ON PQ

Genus STENOCREPIS Chaudoir

(Subgenus STENOUS Chaudoir)
S. cuprea tChaudoirt - ON
leucodacty Ia (LaFert6-S6nectdre)

Tribe Panagaeini
Genus PANAGAEUS Latreille
P. fasciatus Say -oN
Tlibe Callistini
Genus CHLAXNIUS Bonelli
Key to North American species: Bell (1960)
(Subgenus EURYDACTYLUS LaFert6-S6nectdre)
(Gly ptoder us LaFert6-S6nectdre)
C. tomentosus tomentosus (Say) AB SK MB ON PQ
amplus LeConte
lacustrinus Casey
luctuosus Germar
(Subgenus ANOMOGLOSSUS Chaudoir)
C. emarginatus Say - ON PQ NB NS
delectans (Casey)
grauis (Casey)
semotus (Casey)

49
Carabidae

C. pusillus Say - ON
elegant,ilus Dejean
nanulu,; (Casey)
(Subgenus CHLAENIUS s. str.)
C. aestivus liay - ON
cobaltir'.us Dejean
congener LeConte
C. cordicollir, Kirby MB ON PQ NB
C. erlrthropr.. s Germar - ON
oblongi;cennis Casey
rufilabris Dejean
umbritursis Casey
C. laticollis :iay ON
brachyclerus Chaudoir
breuicol.lis LeConte
rufipes Dejean
C. sericeus sericeus (Forster) - BC AB SK MB ON PQ NB NS PE - NF
peruirict.is LeConte
uteanus Casey
C. solitarius Say AB
chlorophanus Dejean
(Subgenus C HLAXNIELLUS Reitter)
C. brevilabr-s LeCon[e - ON
(.?) cons tnilis LeConte
texaneltus Casey
C. impunctilrons Say MB ON PQ
C. nebraske.'rsis LeConte AB SK
sedulus Casey
C. nemoralir Say - ON
Iongico,lis Chaudoir
C. pensylvarLicus pensylvanicus Say BC AB SK MB ON PQ NB NS PE - NF
(?) fulgtceps Newman
impunctifrons l\ttby
pubescc'ns l{arlis
uicinus Dejean
C. tricolor Dejean - BC AB SK MB ON PQ NB NS NF
atripen,'tis LeConte
chaly be ipennis Chevrolat
jacinto Casey
quadricollis Kirby
uigilant Say
(Subgenus F,RACFM,OBUS Chaudoir)
C. Iithophilus lithophilus Say NT BC AB SK MB ON PQ NB NS PE - NF
sma ra g' dig e r Motschulsky
uiridanus Dejean
(Subgenus A GOSTENUS Motschulsky)
C. alternatus Horn AK - NT BC AB SK MB ON PQ NB NS NF
alberta,tus Casey
C. harpalinr s Eschscholtz - BC AB
crestonensis W.J. Browrr
Horn
C. interrupt rs - BC
C. niger Rardall NT BC AB SK MB ON PQ NB NS PE - NF
ludouici.anus Leng
$

C. purpuric< llis Randall NT BC AB SK MB ON PQ

frostii Carr
punctipennis Casey

Tribe Pentagonicini
Genus PENIIAGONICA Schmidt-Goebel
Didetut LeConte
Rhomb tdera Reiche
P. picticornis Bates -ONPQ

50
 Carabidae

Tribe Perigonini
Genus PERIGONA Castelnau
Trechicus LeConte
P. nigriceps tDejeant - ON PQ
umbripennis (LeConte)
P. pallipennis (LeConte) - ON

Tribe Platynini
Genus CALATHUS Bonelli
Key to North American species: Ball and Ndgre (1972)
(Subgenus CALATHUS s. str.)
(Fuscocalathus Ndgre)
C. fuscipes (Goeze)* - BC
(Subgenus NEOCAJ-ATHUS BalI & Nbgre)
C. gregarius (Sayt - ON PQ NB NS
distinguendus LeConte
C. ingratus Dejean AK YK NT BC AB SK MB ON PQ NB NS PE - NF
acomanus Casey
aquilus Casey
beringi Casey
calator Casey
coloradensis Casey
confusus LeConte
incommodus Mannerheim
labrad,orinus Casey
nanulus Casey
planifer Casey
reductus Casey
C. opaculus LeConte - ON PQ
alutaceus Casey
appalachius Casey
obesulus Casey
sonoricus Casey
uentricosus Casey
C. ruficollis grandicollis Casey - BC
(Subgenus PROCALATHUS Jedlicka)
C. advena(LeConte) AK BC AB SK - ON PQ - LB NF
binarius (Casey)
b reu ius cul u s (Mannerheim)
b runne s cen s (Mannerheim)

dzlcis (Mannerheim)
lenis (Mannerheim)
mollis (Eschscholtz)
scolopax (Casey)

Genus LAEMOSTENUS Bonelli

Key to species: Casale (1988)
(Subgenus LAEMOSTENUS s. str.)
L. complanatus (Dejean )* - BC
(Subgenus PRISTOI.IYCHUS Dej ean)
L. terricola (Herbst)f - BC PQ NB NS PE - NF

Genus SYNUCHUS Gyllenhal

Pristodactyla Dejean
Key to species: Lindroth (1956)
S. irnpunctatus (Say) - BC AB SK MB ON PQ NB NS PE - NF
americanus (Dejean)
americanus (LeConte)
conuexu.s (Casey)
coruinus (LeConte)

Genus OLISTHOPUS Dejean

Odontonyx Stephens

51
Carabidae

O. micans LcConte -oNPQ

_ONPQ
O. parmatus (Say)
cinctus )ay
iterans t)asey
pictus C'asey

Genus SERI{IODA Kirby

Agonodromius Reitter
S. bembidioides Kirby AK YK NT BC AB SK MB
cicatricosa Motschulsky
colonica Casey
montico,!,a Casey
S. bogemannii (Gyllenhal)* AK YK _ BC .AB
inuidiosa Casey
str ig icolle (Mannerheim)
tacomae Casey
S. obsoleta ([iay) NT BC AB SK MB ON PQ NB NS _ LB NF
consimi.e (LeConte)
insulina Casey
luctuosc (Dejean)
uicina (()emminger & Harold)
S. quadripun ctata (DeGeer)* AK YK NT BC AB SK MB ON PQ NB NS _ LB NF
octocola (Mannerheim)
stigmost.t, (LeConte)

Genus ANCIIOMENUS Bonelli

Tetraleucus Casey
A. picticornis, Newman -oN
Genus PSEL DANCHUS Casey
P. aeneolus (LeConte) -BCAB
bjorkmcnae Gray & Hatch
P. quadratus (LeConte) -BC
dilleyanus (Casey)
metuen: (Casey)

Genus ATRANUS LeConte

A. pubescens (Dejean) _ONPQNB
obconia.rs (Haldeman)
pallesce.zs Casey

Genus RHADINE LeConte

Comstockia Van Dyke
R. Iindrothi llan AB SK

Genus PARANCHUS Lindroth

P. albipes (F rbricius)'l _NBNSNF
clemens (LeConte)

Genus OXYI'SELAPHUS Chaudoir

Anchus LeConte
O. puncticeps (Casey) -BC_SKMBONPQ_NSPE
pusillus (LeConte)

Genus AGOI{UM Boneili

(Subgenus E UROPHILUS Chaudoir)
A. anchomenoides Randall AK _ NT BC AB SK MB ON PQ NB NS PE - NF
lanellum Gray
A. canadenser Goulet MB ON PQ NB NS
A. consimile (GyllenhaI)x AK YK NT BC AB _ MB ON PQ - NS - LB NF
fragile (Mannerheim)
inualidum Casey
A. darlingtorri Lindroth
A. exaratum (Mannerheim)*
A. galvestonicum (Casey)
AK5*rec _ _ r,rn:x:a]"]'_ _ _

52
 Carabidae

A. gratiosum (Mannerheim) AK YK NT BC AB SK MB ON PQ NB NS PE LB NF
antiquum (Notman)
carri (.Casey)
properans (Casey)
ruficorne (LeConte)
symmetricum (Casey)
A. lutulentum (LeConte) NT BC AB SK MB ON PQ NB NS PE
adustum (Casey)
atriceps (Casey)
elegantulum (Casey)
A. palustre Goulet
A. picicornoides Lindroth
A. retractum LeConte
- - -NT scABsxrunBNiSNsNspr-
BC AB SK MB ON PQ NB NS PE
collusor (Casey)
dil uti p e nne (Motschulsky)
elongatulum Haldeman
facile (Casey)
serenum (Casey)
A. simile Kirby AKYK_BCABSKMB
A. sordens Kirby
ineptum (Casey)
AK YK NT BC AB SK MB oN pe Ns Ns pr LB NF

frosti (Casey)
picicorne (LeConte)
A. superioris Lindroth AK-NT-AB-MBONPQNB
A. thoreyi Dejeanx AK YK NT BC AB SK MB ON PQ NB NS PE _ NF
gemellum (LeConte)
(?) lenum Dejean
picipenne Kirby
(Subgenus PLAT\' IOMICRUS Casey)
A. bicoior (Dejean)* AK YK NT BC AB
ma r g inell un'L (LeConte)
A. fer"ruginosum (Dejean) AK _BC AB SK
erasutn (LeConte)
marcidum (Casey)
uadei (.Casey)
A. nigriceps LeConte* AK NT BC AB SK MB ON PQ NB PE NF
fragilissimum (Casey)
A. piceolum (LeOonte) _BC AB SK MB ON PQ NF
d.issensum (Casey)
fractum (Casey)
frigidulum (Casey)
lasciuum (Casey)
tritum (Casey)
(Subgenus AGONUM s. str.)
(Melanagonum Casey)
(Paragonum Casey)
(Punctagonum Casey)
(Taphranchus Casey)
A. affrne Kirby AK YK NT BC AB SK MB ON PQ NB NS PE NF
cor6o (LeConte)
A. albicrus Dejean -oN
A. belleri (Hatch) BC
A. brevicolle Dejean BC
frater (LeConte)
A. corvus (LeConte) BC AB SK MB ON PQ
debiliceps Casey
hyslopi Casey
A. cupreum Dejean NT BC AB SK MB ON PQ NB
borealinum Casey
chalceum LeConte
crassicolle (LeConte)
cupreolucens Casey
esuriale Casey
Iongulum Casey
marquettense Casey
oualicauda Casey
parallelum Casey
protractum (LeConte)
seminitidum Kirby

53
Carabidae

A. cupripenne (Say) NT BC AB SK MB ON PQ NB NS PE - NF
gemmeum Casey
nitidulunt Dejean
A. deceptivurn (LeContet NS
A. excavatum Dejean - ON PQ NB
ontarioni; (Casey)
trinariunt (Casey)
A. fidele Case, - ON PQ NB NS PE
laeue (Let)onte)
molestwn (LeConte)
subinflat,tm Casey
A. harrisii LeConte - BC AB SK MB ON PQ NB NS PE
aethiops l)asey
mordax Casey
A. melanariun Dejean - BC AB SK MB ON PQ NB NS PE
maurum Haldeman
militare l)asey
(?) scuteliiare (.Say)
A. metallescer:rs (LeConte) NT BC AB SK MB ON PQ NB NS NF
lacustre Oasey
terracens: Casey
A. moerens Dr jean MB ON PQ
A. muelleri (Eerbst)f - BC - ON PQ NB NS PE - NF
hardyi (LeConte)
A. mutatum ((lemminger & Harold) AK BC - SK MB ON PQ NB NS PE LB NF
dtratum t LeConte)
A. octopunctat.um (Fabricius) - ON PQ NB NS PE
A. placidum (fiay) - BC AB SK MB ON PQ NB NS PE LB NF
amplior (.)asey
aztecanu;n Casey
citatum (iasey
morosum Dejean
rhodeant.m Casey
A. propinquurr (Gemminger & Harold) AK YK NT BC AB SK MB ON PQ NB NS PE - NF
amens Ci:tsey
fratercuhLm (LeConte)
humile Crcey
insuetum Casey
piceum LeConte
A. quinquepu:rctatum Motschuisky* AK YK NT - AB SK MB - PQ - LB
perforatum (LeConte)
A. subsericeurn (Leconte) - BC AB SK
latiusculum Casey
suffusum Casey
uteanum Casey
uirid.issintum Casey
A. tenue (Lec')nte) - ON PQ NB NS PE
A. trigeminunr Lindroth MB ON PQ NB NS PE
(Subgenus MICRAGONUM Casey)
(Circinal;:dia Casey)
A. aeruginosum Dejean - ON PQ NB NS
undulatum (Casey)
A. nutans (Sayr - ON PQ
femoratun Dejean
A. striatopunr.tatum Dejean - ON
breuiceps (Casey)
decipiens LeConte
luculentum (Casey)
A. variolatum (LeConte) - BC
coronadiizum (Casey)
(?) limba:um Motschulsky
shastanicum (Casey)
(Subgenus CIRCINALIA Casey)
A. crenistriatum (LeConte) - oN PQ - NS PE
liticola (Casey)
politissimum (Casey)
roticolle (Casey)
statenense (Casey)
A. punctiforme t Sayt 2 -

54
 Carabidae

foueicolle Chaudoir
ludouicianum (Casey)
A. rufrpes Dejean ON
rubripes (Zimmermann)
(Subgenus STEREAGONUM Casey)
A. errans (Say) BC SK MB ON PQ
erythropus Kirby
subcordatum (LeConte)
A. ferreum Haldeman ON
ocreatum Haldeman
quadrulum (Casey)
solid.ulum (Casey)
(Subgenus STICTANCHUS Casey)
Key to species: Liebherr (1986)
A. decorum (Say) BC SK MB ON PQ
arenarium (Casey)
californicum (Dejean)
charmis (Bates)
ftornii (Hausen)
itnpictum (Casey)
irruptum (Casey)
luxatum (Casey)
obscurum (LeConte)
sitnplex (LeConte)
solutum (Casey)
syrdcusense (Hatch)
tepidum (Casey)
t e stac eonot um (Hausen)
thoracicum (Dejean)
uteanum (Casey)
uinnulum (Casey)
A. extensicolle (Say) SK MB ON PQ NB NS
clientulum (Casey)
(? ) cy ane sce ns (Motschulsky)
gaudens (Casey)
lecontei LeConte
ob s c urat utn (Chaudoir)
proximum (Harris)
uigilans (Casey)
uiride (LeConte)
(Subgenus DERATANCHUS Casey)
A. quadrimaculatum (Horn) AB

Genus PLATYNUS Bonelli

(Subgenus PLATYNUS s. str.)
P. brunneomarginatus (Mannerheim) BC
bicolor LeConte
bicoloratus Gemminger & Harold
cinctellus LeConte
ma r ginat u s (M6n6tries)
rugiceps (Mannerheim)
tenebricosus Gemminger & Harold
P. cincticollis (Say)
(?)
oN PQ NS
blandus Germar
coruinus (Dejean)
deplanatus (Chaudoir)
(?) maculifrons (Say)
marginalis (Haldeman)
P. decentis (Say) AK NT BC AB SK MB ON PQ NB NS LB NF
albertanus (Casey)
aleneanus (Casey)
coracinus (LeConte)
d.epressus (Haldeman)
gagates (Dejean)
iowanus (Casey)
missourianus (Casey)
oblongipennis (Casey)

DD
Carabidae

pacatus ( Casey)
sinuatus (Dejean)
turbidus (Casey)
P. mannerhei:nii (Dejean)* AK YK NT BC AB SK MB ON PQ NB NS PE LB NF
inter s t itiali s (Motschulsky)
octofoueo atus (Maklin)
t

stygicus ,-eConte
P. opaculus L':Conte - ON PQ
6oopls (Casey)
inquisitor (Casey)
P. tenuicollis LeConte) - ON PQ NB NS
distinguendus (Casey)
margina;:us (LeOonte)
pennsylu znicus (Casey)
reflexus -l,eConte
(Subgenus PI ATYNIDIUS Casey)
P. hypolithos Sayr - ON
erythrop'.ts Dejean
ontarien::is (Casey)
P. ovipennis (Mannerheim) - BC
arachnoides (Casey)
maurus (Motschulsky)
rotundip znnis (Motschulsky)
similatut: (Casey)

Tribe Ctenodactylini
Genus LEPT( )TRACHELUS Latreille
Sphaera':ra Say
L. dorsalis (F.rbriciusj - ON

Tribe Odacanthini
Genus COLL. URIS DeGeer
(Subgenus C()SNANIA Dejean)
(.O dacan,, hella Liebke)
C. pensylvanica (Linn6r ON PQ

Tlibe Masoreini
Genus TETR,,\GONODERUS Dejean
T. lasciatus (llaldemanj ON
undulatt'.s LeConte

Tribe Lebiini
Genus LEBIA. Latreille
Key to Nor:,h American species: Madge (1967)
(Subgenus LOXOPEZA Chaudoir)
L. atriceps LeConte AB SK MB
nanulinc: (Casey)
L. atriventris Say - SK MB ON PQ
enormis .Casey)
L. grandis Hrntz - ON PQ
magister (Casey)
majuscui.a (Chaudoir)
riuularis (Casey)
L. tricolor Sa., - ON PQ
(Subgenus LIMPRIAS Bonelli)
L. divisa LeC rnte AB SK MB
concinna LeConte
(Subgenus LI)BIA s. str.)
(Ap helog enia Chaudoir)

56
 Carabidae

(D ianchomena Chaudoir)
(Metabola Chaudoir)
L. analis Dejean ON
anchora Chevrolat
ap p endic ulata Chaudoir
bonellii Putzeys
(? ) s ubfr.g ur ata Motschulsky

L. cyanipennis Dejean BC AB SK
barbarae Casey
montanaHorn
melaenaHatch
ruficollis LeConte
L. fuscata Dejean BC MB ON PQ NB NS
canonica Casey
L. guttula LeConte BC
metuens Casey
pacifica Casey
L. lobulata LeConte
(? ) b r unnicol lis Motschulsky
-oN
L. moesta LeConte BC SK MB ON PQ NB NS LB NF
cynica Casey
L. ornata Say
(?) apicalis (Haldeman)
-oN PQ

asheuillensis Casey
axillaris Dejean
brunnea Haldeman
erythrocephala Dejean
fl au iu entris Motschulsky
fluuiatilis Casey
frigida Chaudoir
marginella Dejean
nigripennis Dejean
reperta Casey
uirginica Casey
L. perita Casey _BC
L. pleuritica LeConte _SK ON
L. pulchella Dejean AB SK MB ON
rufopyga (Chaudoir)
tahoensis Casey
uiuida (Bates)
L. pumila Dejean BC AB SK MB ON PQ NB NS
frugalis Casey
illini Casey
ludouiciana Casey
maculicornis LeConte
quadrata Casey
rhodopa Schwarz
tertiaria Casey
L. solea Hentz SK MB ON PQ NS
(.?
) fl au olineata Motschulsky
scapularis Dejean
websteri Casey
L. viridis Say YK NT BC SK MB ON PQ NB NS PE
adolescens Casey
bracata Casey
castigata Casey
cobaltina Casey
(?) cyanella (Motschulsky)
duluthiana Casey
euoluta Casey
histrica Casey
incitata Casey
magica Casey
papago Casey
planifera Casey
prominens Casey
smaragd.ula Dejean
subaffinis Casey
subopaca Schaeffer
trajecta Casey

57
Carabidae

truckeens s Casey
uermiculi za Casey
uiridis D< jean
L. vittata (Fabricius) - BC AB SK MB ON PQ - NS
amnicola Casey
conjunget:s LeConte
connecta tlChaudoir)
debiliceps Casey
depicta Horn
flauouittata Chevrolat
furcata LeConte
sonomae l)asey
spraguei (}{orlr)
tempeana Casey

Genus DROM. US Bonelli

(Subgenus DRIMIUS s. str.)
(Dinodrortius Casey)
D. piceus Dejean - BC AB - MB ON PQ NB NS
quadricollis LeConte

Genus APRIS''IUS Chaudorr

A. constrictus Jasey BC AB
A. latens (LeCrntet - SK MB ON PQ - NS
(.?)
fuscipt nnis Motschulsky
A. pugetanus l)asey - BC
A. iubsulcatus (Dejean) - ON PQ NB NS
cordicolli, ; (LeConte)

Genus MICRC LESTES Schmidt-Goebel

Blechrus Motschulskv
Bomius L eConte
M. brevilobus Lindroth - ON
M. curtipennis (Casey) AB SK
M. linearis (Lt,Conte) - BC AB SK MB ON PQ - NS
anEustus (LeConte)
prominul rs (Casey)
M. major Lindroth - BC
M. nigrinus (N'[annerheim) - BC
fretus (.Ca,sey)
obispinus (Casey)
M. pusio (Leconle) - ON

Genus SYNTC,MUS Hope

M etable tt.. s Schmidt-Goebel
s. americanus (Dejean) AK \T( NT BC AB sK MB ON PQ NB NS PE LB NF
borealis ( 4immermann)

Genus AXINOPAIPUS LeConte

Variopalpis Solier
A. biplagiatus tDejeant - BC AB SK - ON PQ

Genus CALLtr.IDA Dejean

(Subgenus CAI,LEIDA s. str.)
(Spongoloba Chaudoir)
C. punctata LeConte MB ON PQ
C. purpurea tsiayt IIB
cyanipennis Chaudoir
smaragdina Dejean
(Subgenus PHILOPHUGA Motschulsky)
Key to species: Larson (1969)
C. viridis amorna (LeConte) - BC AB SK MB
canora (.Casey)
obscura ( lasey)
puella (C*ey)

58
 Carabidae

Genus TECNOPHILUS Chaudoir

Key to species: Larson (1969)
T. croceicollis peigani Larson AB SK

Genus CYMINDIS Latreille

(Subgenus PINACODERA Schaum)
(Pla nes us Motschuisky)
C. limbata Dejean - ON PQ - NS
(?) fuscicollis (Motschulsky)
(?) laeuigata (Motschulsky)
C. platicollis tSayt - ON PQ
fuscata Dejean
uirescens Drrty
(Subgenus CYMINDIS s. str.)
C. americana Dejean - ON PQ
continens Casey
uenator Dejean
C. borealis LeConte - BC AB SK MB ON PQ NB NS NF
C. cribricollis Dejean - YK NT BC AB SK MB ON PQ NB NS - LB NF
abstrusa LeConte
acomana Casey
alternans Casey
alticola Casey
coseyi Liebke
hirbyi Casey
marginata Kirby
obliqua Casey
planifera Casey
reflexa LeConte
rupimontis Casey
sinuata Casey
tardaLiebke
C. interior Lindroth AB SK MB
C. neglecta Haldeman AB SK MB ON PQ NB
C. pilosa Say - ON PQ
cribrata LeConte
pubescens Dejean
C. planipennis LeConte NT BC AB SK MB - PQ
bipartita Casey
b reu i p enni s Zimmermann
directa Casey
gouanica Casey
C. unicolor Kirby AK YK NT BC AB - MB ON PQ - LB NF
hudsonica LeConte
parowana Casey

Genus APENES LeConte

(Subgenus APENES s. str.)
(Nominus Motschulsky)
A. sinuatus (Say) - ON
pustulatus (Dejean)

Tlibe Zuphiini
Genus ZUPHIUM Latreille
Z. americanum Dejean -oN
Tfibe Galeritinr
Genus GAJ-ERITA Fabricrus
Galeritina Jeannel
Galeritula Strand,
Key to New World species: Reichardt (1967)
(Subgenus PROGAIERITINA Jeannel)
Key to species: BalI & Nimmo (1983)

59
Haliplidae

G. janus (Fablicius) MB ON PQ
ahasueru; Liebke
americana (Fabricius)
borealis Oastelnau
caseyi Lel'g
caseyi Lit:lske
cordicollis Chaudoir
cyanipeniis Dejean
intermed o Casey
occipitali.; Casey
thoracica Casey

Family HALIPLIDAE crawling water beetles

R.E. Roughley

Haliplidae is a family of aquatic beetles that dent invasion of the water (for alternative views,
includes 3 7 species in Canada and Alaska. The see Kavanaugh 1986). There is no recetrt taxo-
adults are distinguished among aquatic beetles by nomic revision of the North American fipecies.
their enlarged hind coxal plates suspended below Matheson (1912) and Roberts (1913) provided the
the abdominal sterna. Adult haliplids swim slowly last comprehensive treatments. Proper species
or crawl o r submerged vegetation. Members of identification often requires examination of male
Brychius are found in erosional portions of streams. genitalia. Therefore, previously widespread species
Those of other genera use a wide variety of freshwa- are found to represent species complexes. A num-
ter habitats but are most often found in permanent, ber of regional taxonomic works, such as Gordon
clean, nutrient-rich ponds and lakes or in slow and Post (1965), Matta (1976), Brigham (1982),
backwaters of streams. Larvae feed on periphyton Hilsenhoff and Brigham (1978), and Gunderson
or algae. Adults are primarily algophagous but may and Otremba (1988), are useful for interpreting the
be facultative scavengers. Canadian fauna.
The hr'liplids have often been placed near
Dytiscidae in classifications. This position is main- AK (3); YK (4); NT (9); BC (23); AB (15); liK (10);
tained by lleutel and Roughley (1988), who, how- MB (13); ON (17); PQ (18); NB (3); NS (3); LB (2);
ever, sugg(, st that haliplids represent an indepen- NF(4) . H(1)

Genus BRYCI'IIUS C.G. Thomson

Key to North American species: Carr (1928)
B. albertanus Carr _AB SK MB
B. hornii Crotr:h BC
B. hungerfordi Spangler -oN
B. pacificus Carr BC

Genus HAJ-IPLUS Latreille

Key to North Arnerican species: Wallis (1933)
(Subgenus IL.aI-IPLUS s. str.)
H. allisonae Brigham BC
H. apicalis C.(i. Thomson* YI< Nr BC AB SK MB
strigatus Roberts ^K
H. blanchardi Roberts
H. distinctus ''Vallis BC _-PQ
H. dorsomacu, atus Zimmermann BC -NF
H. falli Mank A]{ YK NT MB_PQ
H. immaculicollis Harris AK }'K NT BC AB SK MB ON PQ NB \Tq DF LB NF
americanus Aub6.
impressus Kirby
H. longulus LeConte NT BC AB MB ON PQ
hoppingiWallis
H. robertsi Zirnmermann BC
pallidus loberts
H. stagninus lL,eech BC AB SK MB
(Subgenus PARAIIAPHLUS Wallis)

60
 Noteridae

H. borealis LeConte _ABSKMBONPQ

H. connexus Matheson -MBONPQ NB NS
H. fasciatus Aub6 ON PQ
H. leopardus Roberts -PQ
H. pantherinus Aub6 oN PQ
H. triopsis Say ON PQ
(Subgenus LIAPHLUS Guignot)
H. canadensis Wallis _BCAB SK MB ON PQ
H. columbiensis Wallis NT BC
_LB
H. cribrarius LeConte
H. leechi carteri Leech
H. leechi leechi Wailis
-YKNTBCAB
_BC
_BCAB
:" yu oN PQ NB NF

H. nitens LeConte NT ON
H. salinarius Wallis _BC
H. salmo Wallis _BCAB
H. subguttatus Crotch NT BC AB SK MB ON PQ _NS NF
H. ungularis Wallis _BC
H. vancouverensis Matheson _BCAB
Genus PELTODYTES R6gimbart
Cnemidotus Erichson
P. callosus (LeConte) BC AB
P. duodecimpunctatus (Say) oN PQ
P. edentulus (LeConte) BC AB SK MB ON PQ
P. lengi Roberts ON PQ
P. muticus (LeConte) oN PQ
P. torbulosus Roberts BC AB SK MB oN PQ

Family AMPHIZOIDAE trout-stream beetles

Y. Bousquet

Amphizoidae is a small monogeneric family con- waterfalls. They apparently feed on living stonefly
taining the genus Amphizoa, which has four nymphs.
species: three occur in western North America and Kavanaugh (1986) recently revised the species
one in the People's Republic of China, wesrern of this family.
Szechwan Province. Both larvae and adults are
found crawling over submerged rocks, debris, or
wood in rivers, fast-flowing mountain streams, or aK (1); YK (2); BC (3); AB (2)

Genus AMPHIZOA LeOonte

Dysmalhes Mannerheim
A. insolens LeConte AKYK_BC
josephi Matthews
sah I ber g ii (Mannerheim)
A. lecontei Matthews YK_BC
carinata Edwards
planata Van Dyke
A. striata Van Dyke BC

Family NOTERIDAE burrowing water beetles

R.E. Roughley

Noteridae is a family of aquatic beetles that larvae of a European species burrow in the mud and
includes only two species in Canada. Adults and take air from submerged plant tissue. Most North
larvae are found in standing water and are associ- American larvae crawl on the substrate or pond
ated with dense masses of aquatic vegetation. The vegetation. Adults are predaceous, and larvae may

61
Dytiscidae

be omnivorous. A detailed study of the life history known (Beutel and Roughley 1987), and ther system-
of any North American species is needed. atic position of the family near Dytiscidae is well
Taxonomically, Nearctic noterids are well supported (Beutel and Roughley 1988, Kavanaugh
known bet:ause all genera have been revised. No 1986).
additional species are expected for the Canadian
fauna. TlLe classification of noterids is fairly well oN (2);PQ (1)

Genus HYDFi,OCANTHUS Say

Key to Nevr World species: Young (1985)
(Subgenus IIIDROCANTHUS s. str.)
H. iricolor Say ON PQ
similato- Zimmermann

Genus SUPIJISELLUS Crotch

Key to North American species: Young (1979a)
S. puncticolli; (Crotch r - ON

Family DYTISCIDAE predaceous diving beetles

D.J. Larson and. R.E. Roughley

In Car'ada and Alaska 262 species of Dytiscidae especially within the larger genera. Identifrcation
are known, making this the largest family of of species may be difficult, as differences between
aquatic beetles in the region. These beetles occur in species are often subtle. There is therefore a need
a wide valiety of shallow-water habitats. The rich- for additional analysis to provide better species defi-
est fauna is in the emergent vegetation zone along nitions and characters for their recognition.
the edges of pools, small ponds, and small, slow Reviews and revisions of many genera hirrve been
streams. .t\. number of species are characteristic of prepared or are in progress, but no single publica-
cold-wate: habitats, such as small seepages and iion treats the entire fauna. The regional works of
springs, as weII as bog, forest, alpine, and arctic Hatch (1953) and Larson (1975) together ccver most
pools. Clear, gravelly streams and saline ponds of the western and boreal species. Wallis (1973)
each contain distinctive groups ofspecies. and Larson and Colbo (1983) have publisled fairly
Adults are predators or scavengers, and all lar- complete faunal lists for Manito ba and
vae are apparently predaceous. The range of prey Newfoundland, respectively. Leech and Chandler
taken probably is determined by what prey of suit- (1956) and D.S. White et al. (1984) provide good
able size , s available, but some hydroporines feed keys to adults and larvae of most North l\merican
largely olL microcrustacea, whereas some colym- genera. A number of works treating the fauna of
betines prey mainly on culicid larvae. As far as is various states have been published in the United
known or can be inferred, aII species in the fauna States. Some of these works are especially valuable
are univo itine or semivoltine. Most species over- in interpreting the Canadian fauna: California
winter as adults, but time of breeding and occur- (Leech and Chandler 1956), Florida (Young 1954),
rence of larval stages vary greatly among species. Maine (Malcolm 1971), Nevada (La Rivers 1951),
Represenuative larvae of most genera have been North Dakota (Gordon and Post 1965), North and
described, but larvae are known for only a small South Carolina (Brigham 1982), and Utah (R.D.
percentagr: of species. Anderson 7962). Zaitzev (1953) outlijned the
The feLmily is relatively well known taxonomi- Palaearctic distribution of Holarctic species.
cally. The principal taxonomic problems include
relating the northern Nearctic fauna with that of
the Palaezrrctic; analyzing and interpreting patterns AK (92); YK (111); NT (118); BC (167); AB {153); SK
of geogral,hical variation, which in many species is (r24); MB (145); ON (155); pQ (147); NB (82); NS
complex; aind developing suprasp ecifrc classifrcation, (77):PE (31);LB (7a); NF (78) ' H (45)

Subfamily COPELATINAE
TYibe Copelatini
Genus COPELATUS Erichson
Key to No:r'th American species: Young (1963o)

62
 Dytiscidae

C. chewolati Aub6 - ON
renouatus Guiglot
C. glyphicus (Say) - oN pe - NS pE - NF

Subfamily AGABETINAE
Tlibe Agabetini
Genus AGABETES Crotch
A. acuductus (Harris) - ON pe

Subfamily LACCOPHILINAE
T?ibe Laccophilini
Genus LACCOPHILUS Leach
Key to North American species: Zimmerman (1970)
L. biguttatus Kirbyx AK YK NT BC AB SK MB ON pe NB LB
inconspicuus Fall
stroehmi C.G. Thomson
L. maculosus Say - BC AB SK MB ON pe NB NS pE
cal ifornic u s Motschulsky
descipiens LeConte
fusculus Sharp
shermaniLeech
trunco tus Mannerheim
L. proximus Say - ON
americanus Au.b6
confusus Sharp
L. undatus Aub6 - ON

Subfamily FfYDROPORINAE
Tlibe Hydrovatini
Genus HYDROVATUS Motschulsky
Key to North American species: Young (1g56, 1g63b)
H. pustulatus (Melsheimer) - ON pe
compressus Shatp
cuspidatus auct. amer.

Tribe Methlini
Genus CELINA Aub6
Key to North American species: Young (1979b)
C. hubbelli Young - ON pe

Tlibe Hyphydrini
Genus DESMOPACHRIA tsabinston
D. convexa (Aub6) NT BC AB SK MB ON PQ NB NS PE

Tlibe Bidessini
Key to genera: Young (1967)
Genus BIDESSONOTUS R6gimbart
Key to North American species: Young (1g54)
B. inconspicuus (LeConte] - ON pe - NS
B. pulicarius (Aub6)
- ON

Genus LIODESSUS Guignot

Key to North American species: Larson and Roughley (1990)
L. affinis (Say) AK yK NT BC AB SK MB ON pe NB NS pE - NF
charlotti (Clark)
emilianus (Clark)
e ry t h r ostotn us (Mannerheim)

63
Dytiscidae

macularis (LeOonte)
microrei.iculatzs (Hatch)
nanus (l\tb6)
nigrinut (Casey)
obscurellus (LeConte)
L. cantralli (''loung) .AB SK MB
L. fl avicollis r LeConte) _MBONPQ
L. fuscatus (Crotch) SKMBONPQ_NS_LB
Genus NEOCLYPEODYTES Young
N. ornatellus (Fali)

Genus IJVAIiiUS Guignot

U. granarius (Aub6) MB ON PQ NB NS
U. lacustris (Say) oN______
U. subplicatt.s (Hatch)

Tribe Hydroporini
Genus HYGIIOTUS Stephens
Coelamitus C.G. Thomson
Key to North American species: Anderson (1971, 1976, 1983)
H. acaroides lLeconte) MB
winnipe,g (Wallis)
H. bruesi (Fzll) AB
H. compar (tr all) AK _BC AB SK MB ON PQ NB LB
H. dissimilis Gemminger & Haroid _BC AB_MBONPQ
dispar (. -eConte)
H. falli tWallis ) - _ONPQ NF
H. farctus (L rConte) ABSKMB_PQNB
H. impressol unctatus (Schaller)x AK \-I( NT BC AB SK MB ON PQ NB NS PE LB NF
similis (.Kjtby)
H. infuscatur (Sharp) - YK NT BC AB SK MB
dentiger (FalI)
oregonu.; (FalI)
H. intermedi -rs iFall,1 _BC
H. laccophili rus t LeContet NT_ABSKMBONPQNBNS
H. lutescens ileConte) AK _BC
impress /rons Motschulsky
infacetus (Clark)
mediali:: (LeConte)
H. marklini (Gyllenhal)x AK YK NT AB SK MB PQ NF
canadensis (Fa11)
uirgo (F:rll)
H. masculinr s (Crotchl BC AB SK MB
fastidiotus (Fa11')
H.
H.
nigrescen r tFallt
novemlinc,atus hudsonicus (Fa11)* AK YK NT
BC
MB
-pe
H. nubilus tI eConte) oN PQ NN NS
H. obscurepl rgiatus t Fall t sc
columbi znus (W.J. Brown)
H. patruelis ileConte) AK YK NT BC SK MB ON PQ LB NF
borealis (Fall)
discoideus (LeConte)
quebecensis (W.J. Brown)
H. picatus (Iriirby) AK YK NT BC AB SK MB ON PQ NB NS PE LB NF
H. punctilintatus tFall t YK BC AB SK MB ON
H. salinarius lWallist _NT _ABSKMB
H. sal Balfour-Browne YK NT BC AB SK MB ON PQ NB NS PE LB NF
punctatus (Say)
H. sellatus (LeConteJ NT _ABSKMBON
H. semivittarus tFall) BCAB_MB
H. suturalis rl,eConte) AK YK NT BC AB SK MB ON PQ in lre
ouoideur,; (LeConte)
H. sylvanus ,Fallt ON PQ
H. tumidiver tris (Fallt _NT BC AB SK MB ON
H. turbidus (LeOonte) AK YK NT BC AB SK MB ON PQ NSpn--
H. unguicularis (Crotch)* AX YK NT BCABSKMB-PQ LB NF
mongoli:us (Jakovlev)

64
 Dytiscidae

Genus FIYDROPORUS Clairville

(Subgenus HETEROSTERNUTA Strand)
(H eter o ster nu s Zimmermann)
(H etero s teth u s Falkenstrom)
Key to species: Matta and Wolfe (1981)
H. alleghenianus Matta & Wolfe PQ NB
H. cocheconis Fall PQ
H. pulcher LeConte _ONPQNBNS-LBNF
H. wickhami Zaitzev -oNPQNB
concinnus LeConte
(Subgenus SANFILIPPODYTES Franciscolo)
H. browni Wallis BC
H. brumalis W.J. Brown PQ
H. compertus W.J. Brown BC AB
H. edwardsi Wallis BC AB
H. falsificus W.J. Brown BC
H. pacil-rcus Fall BC
H.
H.
planiusculus Fall
terminalis Sharp
BC
BC
AB sx y" oN pe LB NF

- - - pq-
H. acadianus Balfour-Browne -
H. persimilis Crotch N3 LB NF
aequus Fall
H. paugus Fall AK YK NT BC AB SK MB ON PQ NB NS LB NF
H. rubyae Larson YK NT BC AB SK MB ON PQ NB LB NF
H. stagnalis Gemminger & Harold _NT_AB SK MB ON PQ NB NS DD
rD -NF
collaris LeConte
(Subgenus NEOPORUS Guignot)
(C ir cinopo ru s Guignot)
Key to species: Fa]] (1923) and Wolfe (1984)
H. blanchardi Sherman _ONPQ
H. carolinus Fall -oNPQNBNS-LBNF
H. clypealis Sharp _ONPQNBNSNF
anticus Shato
H. dilatatus Fall NB
H. dimidiatus Gemminger & Harold _AB
semirufus LeConte
H. mellitus LeConte ON PQ NB
H. mixtus LeConte ON PQ
H. solitarius Sharp SK MB ON PQ NB NS pn ie NF
H. spurius LeConte ON PQ NB
peltatus Shatp
H. sulcipennis Fall _ONPQ NB
H. superioris Balfour-Browne - Nr ec AB SK MB ON
sericeus LeConte
H. tennetum Wolfe
H. undulatus Say NT BC AB SK
-oN
MB ON PQ NB NS PE LB NF
anticus Sharp
consimilis LeConte
fasciatus Haris
placatus Fall
pubipennis Aub6
p un ctati s sirn u s Attb6.
scrutator Sharp
uelutinus Aub6
H. vittatus LeConte AB SK MB ON
H. vitiosus LeConte oN PQ
(Subgenus IIIaDROPORUS s. str.)
Key to North American species: Fall (1923)
H. appalachius Sherman YK NT BC AB SK MB ON PQ
signatus Sharp
H. badiellus Fall AK YK NT BC AB SK MB ON PQNBNS_LBNF
H. brevicornis Fall PQNBNS_LBNF
H. carri Larson AB
H. columbianus Fall YK NT BC AB s* ms ON PQ NB

65
Dy'tiscidae

H. dentellus Fall AK YK NT BC AB SK MB ON PQ NB NS PE LB
H. despectus Sharp - YK NT BC AB - MB ON PQ - NS NF
rusticut Sharp
H. dichrous Melsheimer - ON PQ
discicol',is Say
H. fatigus G rrdon - BC
H. fortis Le('onte - BC
H. funestus Fall - BC - SK
H. fuscipennis Schaumx AK YK NT BC AB SK MB ON PQ NB LB NF
criniticrtxis Larson
piceus ,cittrrn
puberut'us Mannerheim
H. lapponunL Gyllenhal AK YK NT BC MB ON PQ - LB NF
labradc rensis Fall
[L larsoni NLlsson - YK NT BC AB SK MB - PQ - LB
lL longiusculus Gemminger & Harold AK BC
oblongus Aub6
H. mannerhr:imi Balfour-Browne AK YK NT BC AB
humerolis Alb6
H. melsheimeri Fall ON PQ
H. morio Aur6* AK YK NT BC AB SK MB ON PQ - LB NF
caligintsus LeConte
me lanor:ep ha I us Gyllenhal
H. nigellus l{annerheim* AK YK NT BC AB SK MB ON PQ NB LB NF
H. niger Say - ON PQ NB NS PE - NF
latifrons Sharp
modestus Attb6
H. notabilis LeConte AK YK NT BC AB SK MB ON PQ NB NS PE LB NF
arcticut C.G. Thomson
H. obscurus Sturm* AK YK NT BC AB SK MB ON PQ NB LB NF
hockin1i Larson
H. occidentalis Sharp - YK - BC AB SK
H. pervicinus Fall - BC AB SK MB
H. polaris Fr,Ll1 AK YK NT MB - PQ
subuiret;cens Fall
H. puberulur LeContex AK YK NT BC AB - MB ON PQ NB LB NF
leuanderi J. Sahlberg
H. rectus Fall AK YK NT BC AB SK MB ON PQ NB LB NF
H. rufinasus Mannerheim AK YK NT BC AB SK MB ON PQ - NS
H. sibiricus,,I. Sahlbergx AK YK NT
boreus llordon
H. signatus \{annerheim AK YK NT BC AB SK MB ON PQ NB NS PE LB NF
inornat'"ts Sharp
youngi k,rdorl
H. similaris Fall - BC
H. sinuatipe* Fall - BC
H. striola (Gyllenhal)x AK YK NT BC AB SK MB ON PQ NB NS - LB NF
ambiguus Aub6
subtonsus LeConte
uittula .Eic]l.son
H. tademus Leech - BC AB
H. tartaricus LeConte* AK YK NT BC AB - MB ON PQ
cam ina., iu s Motschulsky
lugubri: Motschulsky
mel an c nolicu s Motschulsky
H. tenebrosus LeConte AK YK NT BC AB SK MB ON PQ - NS NF
luridiptnnis LeConte
lutulen: us LeConte
H. transpun-'tatus H.P. Chandler - BC AB
H. tristis (Pr,rykull)* AK YK NT BC AB SK MB ON PQ NB NS - LB NF
elongat:ilus Schiodte
ru fi co p, ll u s Mannerheim
uarians LeOonte

Genus POTITIVIONECTES Zimmermann

Deronectes auct. amer.
P otarno dy te s Zimmermann
Key to North American species: Zimmerman and Smith (1975a)
P. aequinoctialis (Clarkt AB
P. deceptus Fallr - BC

66
 Dy'tiscidae

P. depressus t Fabricius)* NT_AB_MB

P. elegans (Panzer)8 _BCABSKMBON
P. griseostriatus (DeGeer)x AK YK NT BC AB SK MB ON PQ NB NS pn in Nr
catascopium (Say)
coloradensis (FalIt
mathiasi (Hatch)
parallelus (Say)
suffusus (Sharp)
P. rotundatus ( LeConte t ON PQ NB NS PE LB NF
P. spenceri (Leech) - YK _BCABSKMB
P. striatellus (LeConte) - YK NT BC AB SK MB
infaustus (Clark)
pulcher (Motschulsky)

Genus OREODYTES Seidlitz

Key to North American species: Zimmerman (1985)
O. alaskanus (Fall) AK YK BC AB
p r o d u c t o t r u n c at u s (Hatch)
rainieri (Hatch)
recticollis (Fall)
O. angustior (Hatcht _BC
O. congruus (LeContet _BCAB
O. crassulus iFall; _BCAB
O. laevis (Kirby)
duodecimlinealzs (LeConte)
AK YK NT BC AB MB ON PQ -iuNu
hortensis (Hatch)
semiclarus (.Fall)
yukonensis (Fall)
O. leechi Zimmerman AK YK
O. obesus cordillerensis Larson - YK _BCAB
O. picturatus (Horn) _BC
O. quadrimaculatus (Horn) _BCAB
brodei (.Gellerrnan)
O. sanmarkii (C.R. Sahlberg)* - YK NT
riuolis (Gyllenhal)
O. scitulus {LeConte) NT BC AB SK MB ON PQ NB NS NF
bisulcatus (Fa],\)
O. snoqualmie (Hatchl BC AB

Genus LACCORNIS des Gozis

Agaporus Zimmermann
Key to species: Wolfe and Roughley (1990)
L. conoideus (LeConte) - YK NT BC AB SK MB ON PQ NB _NF
L. deltoides (Fall) ON
L. latens (Fall) ON PQ NB
L. oblongus t Stephens tx NT
L. pacificus Leech BC AB

Subfamily COLYMBETINAE
Tribe Agabini
Key to North American genera: Leech (1942)
Genus AGABINUS Crotcl'
Key to species: Leech (1941)
A. glabrellus (Motschulsky) _BC
A. sculpturellus Zimmermann _BC
Genus AGABUS Leach
Allonychus Zaitzev
Anisornera LeConte
Apator Semenov
Arctodytes C.G. Thomson
Gaurodytes C.G. Thomson
Heteronychus Seidlitz
Ilybiosoma C'rotch
Key to North American species: Fa\l (1922b), Larson (1989)
A. adpressus Aub6* AK YK NT BC
solas Leech

67
Dytiscidae

A. aeruginosrs Aub6 -oN

disposit'ts Guignot
A. ajax Fal1 YK NT BC AB SK MB ON PQ LB
fuscipennis auct. amer.
A. ambiguus (Say) YK _BC AB SK MB ON PQ NB NS PE LB NF
fimbriatus LeConte
reticulaLus A,e.b6
A. antennatus Leech AK YK NT BC AB SK MB ON PQ LB
clauatus LeConte
A. anthracin:rs Mannerherm AK YK NT BC AB SK MB ON PQ NB NS LB NF
scapuloris Mannerheim
A. approximatus FaIl NT BC AB SK
A. arcticus (['aykull)x AK YK NT BC AB SK ms oN pe LB NF
reticulaius (Kirby)
subfasctttus LeConte
A. audeni Wl1lis AK YK NT BC AB SK MB
A. austinii S.rarp AK YK NT BC AB
A. bicolor (K:rby) AK YK NT BC AB SK MB oN- NS-
mutus S.harp
A. bifarius (F.irby)* AK YK NT BC AB SK MB oN PQ NB NS LB
hessleri Hochhuth
A. bjorkmanare Hatch BC AB
rectus (LeConte)
A. canadensir Fall YK_BCABSK MB
A. clavicorniu Sharp* AK YKNTBC_SK MB
uerzs W,J. Brown
A. col)rynbus :eech YK NT MB
A. confertus ,eOonte BC
A. confrnis (( iyllenhal)* AK YK NT BC AB SK MB oN pe - NS- LD NE
longulu: (LeConte)
ouoideut (Crotch)
AK YK NT BC
A. coxaiis Shrrpx
A. discolor (F.arris)
A. discors Lelonte
AK YK NT BC
_BC
AB SK MB ON PQ NB NS PE ie-
A. disintegrabus (Crotch)
A. elongatus (Gyllenhal)* AK YK NT BC AB SX
-oN
MB
PQ

bryanti Jan
A. erichsoni tlemminger & Haroldx AK YK NT BC AB SK MB ON PQ NB NS NF
lutosus t Crotch)
nigroaer,,eus Erichson
sobrinus Motschulsky
A. erythropterus (Say) ON PQ NB NS NF
A. falli (Zimrnermann) AB SK MB oN PQ
semtpunctatus auct. amer.
sharpi F all
A. fuscipenni.s (Paykull)* AK YK NT BC AB SK MB ON
ontarior,,is FalI
A. gagates Arrb6 -oN PQ NB NS
A. hypomelar; Mannerheim AK BC AB
i rregula 'is Mannerheim

A. infuscatus Aub6 AKYKNTBCAB_MBONPQ _LBNF

A. inscriptus (Crotch) AK YK NT BC AB SK MB ON PQ _NS _LBNF
A. leptapsis (LeConte) _YK_BCABSKMBONPQ NB NS _LBNF
A. lugens LeConte -BC
perplexus Sharp
suturalis (Crotch)
A. lutosus LeConte BC AB SK
discolor LeConte
lecontei lCrotch)
mimusLeech
A. margaretae Larson AB SK
A. moestus (Ourtis)* AK YK NT _MBONPQ
borealis Sharp
nigripalcis (J. Sahlberg)
A. nectris Leech -BC
_BC
AB SK
A. oblongulus Fall
A. obtusatus (Say) oN PQ f.fn NS
nitidus lSay)
A. opacus Aub6* AK \'K NT BC AB SK MB ON PQ NB LB NF
gelidus )!a71
pseudoc' tn fertus Wallis
sharpi Jacobson

68
 Dytiscidae

A. phaeopterus (Kirby) AK YK NT BC AB SK MB ON PQ NB NS PE LB NF
A. pisobius Leech _BCAB
A. punctulatus Aub6 _YK AB SK MB ON PQ NB LB
aeneolus (Crotch)
A. semipunctatus (Kirby) AK YK NT BC AB SK MB ON PQ NB NS LB NF
stridulator Shatp
A. semivittatus LeConte ON PQ
spilotus LeConte
texanus Sharp
A. seriatus (Say) YK NT BC AB SK MB ON PQ NB NS NF
arcfzs Melsheimer
intersectus Ctotch
parallelus LeConte
striatus Aub6,
A. smithi W.J. Brown _BCAB
A. strigulosus (Crotch) AK YK NT BC AB SK MB
nanus (.LeConte)
A. subfuscatus Sharp MB ON PQ NB NS PE
A. thomsoni (J. Sahlberg)* AKYKNTBCABSKMB-PQ LB NF
A. tristis Aub6x AK YK NT BC AB SK PQ LB NF
atrdtus Mannerheim
crotchi Zaitzev
dubizs Mannerheim
piceus Zaitzev
piceolus Zaitzev
subopocus Mannerheim
A. triton Fall SK MB
A. vancouverensis Leech AK_-SCT
A. velox Leech AK YK NT _MR
A. verisimilis W.J. Brown _BCAB
A. wasastjernae (C.R. Sahlberg)* AK YK NT BC AB SK MB ON pe- _LBNF
henaiensis Fall
palustris Wallis
A. zattzewi Poppius* AK YK NT MB
hudsonicus Leech
A. zetterstedti C.G. Thomson* AK YK NT MB
browni Leech

Genus CARRI{YDRUS Fall

C. crassipes Fall YK NT BC SK MB ON

Genus ILYBIUS Erichson

Key to North American species: Larson (1987)
I. angustior (Gylienhal)* NS PE LB
I. biguttulus (Germar)
laramaeus LeConte
AK
YNTBCYSKMB3NiS NB
NB NS
NF
NF
L churchillensis Wallis
I. confusus Aub6
deniheiWallis
f_ I'_ _MB
ON NS

I. discedens Sharp AK YK NT BC AB SK MB ON PQ NB NS _LBNF

L fraterculus (LeConte)
I. ignarus (LeConte)
I. incarinatus Zimmermann
--NTBCY:"YuoNpe
-oNPQ
NS __I"
I. oblitus Sharp
I. picipes (Kirby)* AK Nr sc AB !x nrn 3N pe
"K NT
_ \'K -isNr
L pleuriticus (LeConte) BC AB MB SK ON PQ NB NS PE LB NF
inuersus Sharp
I. quadrimaculatus Aub6 AK BC AB
I. subaeneus (Erichson)* AK YK NT BC AB SK MB ON PQ LB NF
uiridianeus Crctch
I. vittiger (Gyllenhal)* AK YK NT BC MB

Tribe Colymbetini
Genus COLYMBETES Clairville
Cymatopterus Dejean
Key to North American species: Zimmerman (1981)
C. dahuricus Aub6x AK YK NT BC AB SK MB ON PQ
obscuratus Mannerheim
seminiger LeConte

69
Dytiscidae

C. densus LeConte AK YK - BC AB SK MB
inaequalLs Horn
C. doiabratus (Paykull)* AK YK NT BC MB - PQ - LB NF
drewsonl. LeConte
groenlanCicus At:.b6
margina'.is (Fabricius)
thomsont. Sharp
triseriatu.s I{trby
C. exaratus LeConte - YK NT BC AB SK MB
i Zimmerman
ncog nitr"r s
rugipenn.is Sharp
C. paykulli Erichsonx - YK NT BC AB SK MB ON PQ NB NS NF
longulus LeConte
C. sculptilis Ftarris AK YK NT BC AB SK MB ON PQ NB NS PE LB NF

Genus NEOS'IUTOPTERUS Balfour-Browne

P seudo sc utop te r u s Hatch
Key to species: Larson (1975)
N. angustus (LeConte) - YK - BC AB SK MB ON PQ NB NS - LB NF
N. hornii (Crc,tch) AK YK NT BC AB SK MB ON PQ NB LB
coriaceut: Hotn
Genus RHANTUS Dejean
Key to Nor',h American species: Zimmerrnan and Smith (1975b)
(Subgenus NI,\RTUS Zaitzev)
(Rantogilon des Gozis)
R. sinuatus (l,eConte) NT BC AB SK MB ON PQ - NS
(Subgenus RIIANTUS s. str.)
R. binotatus (Harris) AK YK NT BC AB SK MB ON PQ NB NS PE LB NF
aequalis ll.atc]r
assimilit (Kirby)
diuiszs ( {ub6)
flauogriseus Crotch
longipes Sharp
obscurus, Shatp
plebeius Shatp
R. consimilis Motschulsky NT BC AB SK MB ON PQ
disceden; Shatp
loslzs (LeConte)
R. frontalis (l{arsham)* AK YK NT BC AB SK MB ON PQ
fla u icoll t s E schscholtz
gyllenhat i (Castelnau)
notatus (Fabricius)
punctatus (Hope)
sericozs Sharp
suturalk (Boisduval & Lacordaire)
R. gutticollis Sayt - BC AB
dominguensis Gemminger & Harold
hoppingt.Wallis
hubbelliHatch
maculiccllis (Arb6)
mexican, t s (Castelnau)
R. suturellus (Harris) AK YK NT BC AB SK MB ON PQ - NS - LB NF
subopacus (Mannerheim)
zimmermanni WalTis
R. wallisi Hatch AK YK NT BC AB SK MB ON PQ NB NS - LB NF
suturellus anct.

Tribe Coptotomini
Genus COPTJTOMUS Say
Key to eastern North American species: Hilsenhoff(1980)
C. longulus LeConte - BC AB SK MB ON PQ NB NS PE
lenticus Hilsenhoff
C. loticus Hil;enhoff - ON PQ

70
 Dytiscidae

Tlibe Matini
Genus MATUS Aub6
Key to species: Young (1953)
M. bicarinatus (Say) ON
M. ovatus ovatus Leech ON PQ

Subfamily DYTISCINAE
TYibe Dytiscini
Genus DYTISCUS Linn6
Leionotus l\trby
Macrodytes C.G. Thomson
Key to species: Roughley (1990)
D. alaskanus Balfour-Browne AK YK NT BC AB SK MB ON PQ LB NF
paruulus Motschulsky
D. circumcinctus Ahrens* AK YK NT BC AB SK MB ON PQ
arrrizs Mannerheim
ooligbukiilirby
D. cordieri Aub6 AK NT BC AB SK MB ON PQ NB
sublimbatus LeConte
D. dauricus Gebler* AK YK NT BC AB SK MB ON PQ NB NS PE LB NF
confluens Say
franklinii l{trby
frontalis Motschulsky
uentralis Motschulsky
uexatus Sharp
D. fasciventris Say _YKNTBCABSK MB ONPQNBNS_LB
D. harrisii Kirby AK YK NT BC AB SK MB ONPQNBNS_LB NF
D.
D.
hatchi WaIIis
hybridus Aub6
-BCABsx MB ON PQ NB
compar (Melsheimer)
D. marginicollis LeConte NT BC AB SK MB
alb ioni c u s Motschulsky
fu scostr iat us Motschulsky
D. verticalis Say MB ON PQ NB NS PE

Tlibe Hydaticini
Genus HYDATICUS Leach
Key to North American species: Roughley and Pengelly (1982)
(Subgenus I{YDATICUS s. str.)
H. aruspex Clark* AK YK NT BC AB SK MB ON PQ NB NS PE LB NF
americanus Sharp
laeuipennis C.G. Thomson
modestus S}:'atp
H. oiceus LeConte AB SK MB ON PQ NS

Tribe Aciliini
Genus ACILIUS Leach
Key to North American species: Hilsenhoff(1975)
A. abbreviatus Mannerheim AK _BCAB
oregonensis Ctotch
MB
A. athabascae Larson
A. mediatus (Say) _ ICT:" -oN
ON PQ
PQ NB NS
maccullochii (Kirby)
A. semisulcatus Aub6
A. sylvanus Hilsenhoff
AK YK y:'T:" MB
MB
ON
ON
PQ
PQ
NB NS PE LB NF

Genus GRAPHODERUS Dejean

Key to North American species: Wallis (1939)
G. fascicollis (Harris) oN PQ

7T
Gytininae

G. liberus (Sr,Ly) NT BC SK MB ON PQ NB NS NF
brunnipznnis Aub6
rugicolli.s I{ttby
thoracic ^ts Haris
G. manitober.sis Wallis
G. occidentalis Horn _YKN,scABs^MBoNpe
G. perplexus Sharp AK YK NT BC AB SK MB ON PQ NB Ns po ie Nr
elatus S tarp

Genus THERMONECTUS Dejean

T. basilaris (]{arris) -oN
Tribe Cybistrini
Genus CYBIfITER Curtis
C. frmbriolatrrs (Say) MB ON NS
crotchi \{ilke
oliuieri (.)totc}'

Family GYRINIDAE whirligig beetles

R.E. Roughley

Adult lyrinids form a group of beetles wonder- Dineutus; few changes have been required since
fully adalrted for life on the water surface; 36 that publication. Fall's work (1922a) is the lastest
species ocsur in Canada and Alaska in streams, revision of the North American species of G'yriruus; a
rivers, pords, and lakes. They can be found quite taxonomic revision of Nearctic Gyrin zs is in
far from s rore but are more common closer to the progress and should help ease the diffrculty of iden-
shore. Thc adults are readily distinguished by their tifying specimens in this genus. Morrisette (1979)
highly mo<1ifred, short and flat middle and hind legs provides good regional works on gyrinids for
and their horizontally separated eyes. Adults glide Quebec, Gordon and Post (1965) for North Dakota,
on the surlace of water, cluster together, and when Sanderson (1982) for North and South Carolina,
disturbed lnove rapidly in circles and scatter widely and Ferkinhoff and Gundersen (1983) for Minnesota
or dive. Such aggregations often include a number and environs. The general classification of ryrinids
of species, Adults produce defensive secretions, is now well understood (Folkerts 1979). Beutel and
which discourage predation by aquatic vertebrates, Roughley (1988) suggest that gyrinids are l;he most
and feed on dead and dying insects trapped on the primitive family of the suborder Adephaga end that
water surf;lce. Larvae are aquatic and respire cuta- they represent a very early and independent use of
neously wibh the aid of lateral abdominal filaments. aquatic habitats by beetles (see also Kavanaugh
Pupation occurs on emergent plants along the 1986).
shoreline.
There is no recent taxonomic treatment of the aK (7); YK (10); NT (14); BC (18);AB (21); SK (23);
North American species of gyrinids. Roberts (1895) MB (22); ON (27); PQ (30); NB (14);xS (101; pE (5);
prepared the latest revision of the species of LB (8);NF (10) . H (4)

Subfamily GYRININAE
Key to North American genera: Folkerts (1979)
Tlibe Enhvdrini
Genus DINEIJTUS Macleay
Cyclinus Kirby
D. assimilis (.iGrby) BC AB SK MB ON PQ PE
americanus (Say)
D. discolor ArLb6
labratus (Melsheimer)
-oN PQ NS

D. horni Roberts _SK MB ON

D. nigrior Rolrerts MB ON PQ

72
 Gyrininae

Tribe Gyrinini
Genus GYRINUS O.F. Miiller
(Subgenus GYRINULUS Zaitsev)
G. minutus Fabricius* AK YK NT BC SK MB ON PQ NB NS
(Subgenus GYRINUS s. str.)
G. aeneolus LeConte AB SK MB ON PQ NB
G. aeratus Stephensx AX YK NT BC AB -oN PQ
instabilis Fall
G. affrnis Aub6 NT BC AB SK MB ON PQ NB NS LB NF
canadensis R6gimbart
G. analis Say MB ON PQ _NS
G. aquiris LeConte AB_ MB ON PQ NB PE NT
G. bifarius Fall NT BC AB SK MB ON PQ NB NS LB NF
G. cavatus Atton -YK NT BC AB MB
MB
-NS
NB
G. confinis LeConte NT BC AB SK ON PQ NS LB NF
G. consobrinus LeConte BC AB SK
G. dichrous LeConte _SK MB ON PQ
G. dubius Wallis _SK MB PQ NF
G. fraternus Couper MB ON PQ -NS PE :u
G. frosti Fall _:* PQ
G. gehringi Chamberlain -oN PQ NF
G. hoppingi Leech NT BC AB SK
G. impressicollis Kirby mn on PQ
G. latilimbus Fall _YK BC AB SK -oN PQ NN NS LB NF
G. lecontei Fall MB ON PQ pe
G. maculiventris LeConte NT BC AB ;K yu oN PQ Tu_
G. marinus Gyllenhalx AK YK
G. marginellus FalI PQ NB
G. opacus C.R. Sahlberg* AK YK NT BC AB SK MB ON PQ LB NF
hoppingiLeech
G. pectoralis LeConte _YK NT BC AB SK MB ON PQ NB
G. piceolus Blatchley PQ
G. picipes Aub6 AK YK NT BC AB SK MB ON PQ
G. pleuralis Fall AK YK NT BC ABSK_ON PQ lln -
G. plicifer LeConte BC AB
G. pugionis Fall _SKMBON PQ NB NS
G. punctellus Ochs BC AB SK
G. sayi Aub6 _YK NT BC AB SK MB ON PQ NB NS pt LB NF
lugens LeConte
G. ventralis Kirby NT ABSK_ON PQ NB-
G. wallisi Fail AK YK NT BC AB SK MB ON PQ

73
Hydraenidae

Suborder Polyphaga
Superfamily Staphylinoidea
Family FIYDRAENIDAE minute moss beetles
R.E. Roughley

Twenty.ssyen species of Hydraenidae are found such as wing venation and the absence of tire basal
in Canada and Alaska. The family includes minute piece on the aedeagus, leave little doubt about its
aquatic beeitles, most of which are under 2 mm long. place in the superfamily Staphylinoidea. Perkins
Adults are found along the edges of aquatic habi- (1980) recently revised the Canadian and .Alaskan
tats, particularly streams; some also live in semi- species in his monograph of the Hydraenidae of the
aquatic habitats. Adults and larvae feed on peri- Western Hemisphere.
phy'ton.
The farnily is sometimes assigned to the super- NT (a); BC (21);AB (10); SK (4); UB (5); ON (5);PQ
family Hyclrophiloidea; however, several features, (7);NB (3);NS (1);NF (1)

Subfamily IIYDRAENINAE
Tribe Hydraenini
Genus HYDR\XNA Kugelann
H. anguiicollil Notman NT_AB-MB ON PQ NB NF
H. atlantica F erkins PQ NS
H. circulata Perkins _BCAB__ IU
H. nigra Hatcr -BC
H. occidentali r Perkins _BC
H. pacifica Pe.:kins _BCAB
H. pennsylvar.ica Kiesenwetter ON PQ_
H. petila Perk ins _BC
H. vandykei d'Orchynont _BC
Genus LIMNEBIUS Leach
Limnocharis Horn
L. alutaceus (Casey) _BCAB
L. borealis Pelkins _BC

Subfamily OCHTHEBIINAE
Genus GYMNOCHTHEBIUS d'Orchwnont
G. nitidus (Le,lonte) NT oN PQ

Genus OCHTII{EBIUS Leach

(Subgenus OCHTHEBIUS s. str.)
(Enicocerus Stephens)
O. benefossus LeConte oN PQ
O. bisinuatus Perkins BC AB__
O. borealis Pe:rkins BC
O. intermptus LeOonte BC
aberti }Ia.tc}r
O. kaszabi Janssens NT BC AB SK MB ON PQ NB
O. lecontei Perkins BC
O. lineatus LeConte BC AB SK MB
milleriHatch
O. marinus PnLykull BC AB SK MB
holmbergi Maklin
O. rectus LeConte BC
(Subgenus ASIOBATES C.G. Thomson)
(H omaloc ht hebizs Kuwert)
.(Trymoch thebizs Kuwert)
O. brevipennir; Perkins BC

74
 Ptiliidae

O. cribricollis LeConte _BCABSK MB_ PQ

O. discretus LeConte _BC
O. hibernus Perkins NT
O. mimicus W.J. Brown _BCAB
Genus NEOCHTHEBIUS d'Orchymont
N. vandykei (Knisch)
lapid.icola (Van Dyke)

Family PTILIIDAE feather-winged beetles

J.M. Campbell

Members of Ptiliidae, a family of very small bee- willbe found, particularly in eastern Canada.
tles, have fringed wings with long filaments, which Hatch (1957) studied the western species. Bistrom
give them a feather-like appearance. The family and Silfverberg (1979) examined many of the
comprises about 560 species around the world, of generic type species and frxed the generic names.
which 32 are known to occur in Canada and Alaska. Dybas and Dybas (1990) recently published a key to
Most feather-winged beetles are found in associa- the North American genera. Many Canadian
tion with moist organic matter, such as leaf litter, records require confrrmation, as the identification of
decaying wood, and fungi. most species has not been checked by specialists.
Ptiliidae is probably the most poorly known
family of beetles in North America. Undoubtedly, aK (5);BC (16);MB (1); oN (2); PQ (10); NB (5); NS
when the family is revised, many additional species (1) . H(1);I(11)
Subfamily PTILIINAE
Genus MOTSCHULSKIUM Matthews
M. sinuatocolle Matthews BC

Genus PTENIDIUM Erichson

P. nitidum (Heer)t NB
P. pullum Maklin _BC
P. pusillum (Gyllenhal)t _BCPQ _NS
apicale Erichson
terminale Holdeman

Genus ACTIDIUM Matthews

A. crotchianum Matthews BC

Genus PTILIOLA Haldeman

P. brevicollis (Matthews)t PQ
P. collani Mannerheim
canadensis LeConte
-oN
Genus PTILIOLUM Flach
P. columbianum (Matthews) _BC
P. obscurum (Matthews) _BC
P. sharpi (Matthews) _BC
P. spencei (A]libert)t PQ

Genus PTILIUM Gyllenhal

Millidium Motschulsky
P. minutissimum (Ljungh)t PQ

75
Limuiodidae

Subfamily ACROTRICHIINAE
Genus NEPHANES C.G. Thomson
N. titan (Nev'man)f PQ

Genus PTILI IPYCNA Casey

P. moerens (ldatthews) MB ON PQ
quadratr.t (Casey)

Genus ACROTRICHIS Motschulsky

Macdonuldium Abdullah & Abdullah
Trichopteryx Kirby & Spence
A. castanea (lvlatthews) BC
A. cognata (Niatthews)f BC Ne-
platonoffi Renkonen
pruinosc Hatdn
A. dilfinis (Mrtthews) BC
A. fasciculari ; (Herbst)i NB
A. fungi (Abdullah & Abdullah) PQ
A. grandicolli s (Mannerheim)f PQ
A. henrici (M,rtthews) BC
fratercuia (Matthews)
A. insularis (.v{aklin) AK BC
A. intermediz (Gillmeister) AK
A. josephi (M,rtthews) BC
A. laticollis N aklin AK
A. parallelopi peda (Matthews) BC
A. sericans (Iteer)i PQ NB
A. sitkaensis (Motschulsky)* AK
A. thoracica ( Waltl)t PQ
anthracina (Matthews)
A. vicina (Ma bthews) BC
A. volans (Mctschulsky)i NB
fennica l:lenkonen
A. xanthocer:. (Matthews) BC

Family LIMULODIDAE horse-shoe crab beetles

J.M. Campbell

Members of this small family are obligate reported in Canada, but one species of Litnulodes,
guests of ants. The family comprises about 30 which I have tentatively identifred as L. pa,radoxus,
species distributed in North and South America and occurs in southern Ontario. Seevers antl Dybas
Australia, lnly one of which is found in Canada. (1943) revised the classification of the farnily and
Recent workers have placed the limulodids in included a key to all genera and most North
the family Ptiliidae (e.g. Lawrence 1982o), but I American species.
prefer to nraintain the group as a separate family,
on basis of papers by Seevers and Dybas (1943) and
Dybas (19?6). This family has not been previously oN (1)

Genus LIMU -ODES Matthews

(Subgenus LIMULODES s. str.)
L. paradoxus Matthews ON

76
 Leiodidae

Family AGYRTIDAE agyrtid beetles

J.M. Campbell

Eleven species of agyrtids are known in However, these studies have shown that agyrtids
America north of Mexico, of which eight are found are more closely related to species of the family
in Canada and Alaska. Adults are scavengers of Leiodidae than to Silphidae and therefore must be
decaying organic material. Larval feeding habits treated as a separate family. The North American
are probably similar to those of the adults species are well known taxonomically. The species
(Anderson and Peck 1985). of Canada and Alaska were treated bv Anderson
Until recent papers by Lawrence (1982o) and and Peck (1985).
Lawrence and Newton (1982), species of this family
had been included in the family Silphidae. AK (4); NT (1); BC (6);AB (3); ON (1)

Genus NECROPHILUS Latreille

N. hydrophiloides Gu6rin-M6neville AK BC
a/er Motschulsky
N. pettiti Horn ON

Genus IPELATES Reitter

Pelates }Iotn
Pelatines Cockerell
I. latus (Mannerheim) AK BC

Genus AGYRTES Frrtlich

(Subgenus AGYRTECANUS Reitter)
A. longulus (LeConte) AK BC
politus (Casey)

Genus LYROSOMA Mannerheim

L. opacum Mannerheim AK
Genus PTEROLOMA Gyllenhal
P. nebrioides W.J. Brown

Genus APTEROLOMA Hatch

A. caraboides (Fall) _BC
A. tenuicorne (LeConte) _BCAB

Family LEIODIDAE small scavenger beetles

S.B. Peclz

In Canada and Alaska 128 species of Leiodidae various decaying organic materials in usually moist
are known. Those grouped here have been placed forest or freld habitats. The two biggest subfamilies
under many family names, but all share the derived are Leiodinae and Cholevinae. Members of the
(but diffrcult to see) character of a nearly enclosed Leiodinae subfamily (= Anisotomidae, Agathidiidae),
gutter that circles the antennal articulation on the or round fungus beetles, are often globose and com-
dorsal face of segments 7,9, and 10 or 8-10 in 11- pact in body shape, and many species can contract
segmented antennae and segments 7-9 in 10-seg- into a ball-like form. Some species are specialized
mented antennae, with internal sensory vesicles in for digging in soil, where they feed on subterranean
some segments. The higher classification here lists fungi; others are specialized predators of slime
five subfamilies and follows recent advances molds; and still others feed on soft fungi or are more
reported by Lawrence (1982o), Lawrence and generally saprophagous (Wheeler 1984). The feed-
Newton (1982), and Newton (1984). ing habits of both adult and larval Coloninae (=
Members of Leiodidae are, for the most part, Colonidae) are not known. Members of the
scavengers, both as larvae and adults, feeding on Cholevinae subfamily (= Catopidae, Leptodiridae),

77
Leiodidae

or small carrion beetles, are more elliptical in body The leiodid fauna of Canada and Alaska is only
shape and feed on carrion or dung. Some species are partly known. Several genera have been reviewed
specializecl occupants of ant nests, caves (but not in and revised recently, and others are being prepared.
Canada), rr the nests and burrows of vertebrates. No single publication treats the entire fauna, but a
Members irf the Leptininae family (= Leptinidae), or generic review of the North American fauna is
mammal rrest beetles and beaver beetles, are nest available (Peck 1990). The largest faunal works
scavengers and ectoparasites on small mammal (Hatch 1933, 1957; Jeannel 1936) are no longer
hosts or on beavers. completely reliable. Distributions and bionomics of
Collec;ing Leiodidae usually requires a concen- the species are usually incompletely known, and
tration on the particular habits or habitats of the many species in the northern United States may
genera or species sought. They can be collected in eventually be found in southern Canada.
the following ways: in baited pitfall traps; by sifting
and Berlese extraction of litter or fungi; by sweeping AK (36);YK (18); NT (16); BC (84);AB (39)t SK (18);
forest vegc,tation on still, warm, and humid evenings; MB (26); oN (57); PQ (53); NB (1e);NS (15); t B (7);
and by use,, of large-area flight intercept traps. NF(10) . H(6);I(1)
Subfamily LEIODINAE
Tribe Hydnobiini
Genus HYDIIOBIUS Schmidt
H. contortus Hatch -BC
H. crestonensis Hatch _BCAB
H. femoratus Hatch BC
H. longulus l,eConte _BC
longidets LeConte
H. luggeri H rtch AK
H. matthewsii Crotch
H. obtusus L,eConte
--ec-- SK
H. pumilus L,eConte ON PQ
hiseri Hatch
latidens LeConte
H. simulator W.J. Brown _BC
H. stacesmit ri Hatch _BC
H. strigilatu r Horn _BC
H. substriatrLs LeOonte AB pe NS
curuid.ens LeConte
H. validus \[.J. Brown PQ

Genus TRIAIRTHRON Markel

T. lecontei Horn _BC

cedonuli'.i Schaufuss

Tribe Leiodini
Genus ANO(]DUS LeConte
Neocyrtlsa W.J. Brown
Key to species: Daffner (1988)
A. fusciclavus (Fall t BC
sternitu; (Hatch)
A. insolitus tW.J. Brownt BC SK
A. obsoletus Melsheimer) oN--
blandis :; ima (Zimmermann)
A. potens tW.J. Brownt BC _ONPQ
A. puritanus {Fallt _ONPQ
A. secretus (W.J. Brown) -oNPQ
A. superans Fall) AB_MBON
Genus LIONOTHUS W.J. Brown
Key to No;th American species: Daffner (1988)
L. forticornis Daffner -oN
L. ulkei W.J Brown -oN PQ

78
 Leiodidae

Genus CYRTUSA Erichson

Caenocyrta W.J. Brown
Key to North American species: Daffner (1988)
C. subtestacea (Gyllenhal)* BC oN PQ
picipennis (LeConte)

Genus LIOCYRTUSA Daffner

Key to North American species: Daffner (1988)
L. luggeri (Hatch) _BCABSKMBONPQNB
L. nigriclavis (Hlisnikovsky)* _BC_ONPQ
Genus ECARINOSPHAERULA Hatch
E. ecarinata (Horn) BC

Genus ISOPLASTUS Horn

I. fossor Horn oN PQ
uncus Wheeler

Genus LEIODES Latreille

L. assimilis (LeConte)
L. bradorata W.J. Brown
yyNrBC AB SK MB ON PQ
PQ
NB NS LB NF

L. canadensis (W.J. Brown) SK MB

L. collaris (LeConte) AKYK_BC AB MB ON PQ NB LB NF
L. conferta (LeConte)
L. curvata (Mannerheim) AK BC AB _ 9*PQ
morula LeConte
L. diffrcilis (Horn)
L. dilatata Hatch
L. grossa Hatch
ncy
- - --BCAB
L. impersonata W.J. Brown -oNpe NB NS
L. lateritia (Mannerheim) AKBC_
L. lostine (Hatch) _BCAB
L. merkeliana (Horn) NT BC
L. mirabilis (Hatch) _BC
L. paludicola (Crotch) -BC
L. puncticollis (C.G. Thomson) AKYK_BCAB SK nfn ON Pq
L. punctostriata Kirby AK YK NT BC AB MB ON PQ NB NS LB NF
indistincta (LeConte)
laefo (Mannerheim)
punctatostriato Horn
punctistriata LeConte
L. punctulata (GyIlenhal) AKYKNTBCAB_MBONPQ
L. rhaetica (Erichson) AK
L. rufrpes (Gebler) AK YK NT BC AB SK MB ON
L. serripes Hatch _BCABSK
L. strigata (LeConte) AKBCABSK-ONPQ NB
L. tenuis Hatch _BC
L. triepkei (Schmidt) -YKNT-AB-MBONPQ NB
L. valida (Horn) NTBCAB_MBONPQ NB

Genus ZEADOLOPUS Broun

Apheloplastus W.J. Brown
Key to North American species: Daffer (1988)
Z. egenus (LeConte) oN PQ
impubis (Zimmermann)

Tlibe Scotocryptini
Genus AGLYPTINUS Cockerell
A. laevis (LeConte) oN PQ

Tribe Pseudoliodini
Genus COLENIS Erichson
C. impunctata LeConte BC MB ON PQ NB NS

79
Leiodidae

Tribe Agathidini
Genus ANIS()TOMA Panzer
Key to North American species Wheeler (1979)
A. amicaW.J. Brown BC oN PQ
A. basalis (Lc,Conte) _ONPQ_NS
dichroa .LeConte)
A. blanchard' (Horn)
A. confusa (Horn)
interstrialis Hatch
BC AB _9* PQ

A. discolor (l\'lelsheimer)
piceo (Melsheimer)
BC -oN PQ

A. errans W..1. Brown AK BC AB SK -oN PQ NS

A. expolita S .J. Brown
polita (LeConte)
-oN
A. geminata r.Horn)
A. globososa {atch BC AB_
-oN
MB ON
PQ
PQ
globosa LeConte)
A. horni Whe,eler
A. humeralis (Fabricius)i
-oN PQ NB
BC
A. inops W.J. Brown
A. nevadensi:i W.J. Brown BC
-oN
MB
PQ NB

bicolor (..Jorn)

Genus AGAT'HIDIUM P anzer

Key to North American species: FaIl (1933)
A. alutaceum Fa11 AK
A. athabasca:rum Fali AB_
A. angulare Idannerheim AK _BC
A. californicu.m Horn _BC
A. cavisternum Fa1l -BC
_BC
A. columbianum Fall
A. concinnuni Mannerheim AK -BC
_BC
A. conjuncturn W.J. Brown
A. contiguun Fall _BC
A. depressun. Fall AK _BC AB PQ
A. difforrnis (LeConte)
canadense W.J. Brown
-oN
A. effluens Mannerheim AK
A. jasperanurn Fall YK
A. maculosurr W.J. Brown
A. obtusum llatch
A. pulchrurn LeConte AK
rilatum Mannerheim
ma ndib
A. rotundulu.n Mannerheim AK
hincaidi Hatch

Subfamily CATOPOCERINAE
Genus CATOPOCERUS Motschulsky
Homoeosoma ArsLin
Pinodytr,'s Hotn
Typhloleiodes Hatch
C. cryptophagoides (Mannerheim) AK BC
nolilzs l[otschulskv

Subfamily COLONINAE
Genus COLCN Herbst
(Subgenus CiJRVIMANON Fleischer)
C. bidentaturn (C.R. Sahlberg)*
C. mannerheimi Szl'rnczakowski
-BCABONPQ
ON Iu_ NF

(Subgenus COLON s. str.)

C. femorale I'Iatch _BC
C. serripoides Hatch _BC
C. tibiale Har,ch _BC

80
 Leiodidae

(Subgenus MYLOECHUS Latreille)

C. aedeagosum Hatch _BC
C. celatum Horn _BC
C. clavatum Mannerheim AK _BC
C. cornplicatum Hatch -BC
C. dentatum LeConte -oN PQ
C. discretum Hatch -BC
C. elongatum Notman
C. inerme Mannerheim AK _BC -oN
C. kincaidi Hatch -BC
C. magnicolle Mannerheim AK NT BC AB MB ON PQ NB LB NF
C. nevadense Horn _BC
C. pribilof Hatch AK
C. pusillum Horn _YK pe

Subfamily CHOLEVINAE
Tribe Ptomaphagini
Genus PTOMAPHAGUS Illiger
Key to North American species: Peck (1973)
P. breviorJeannel ON PQ
P. nevadicus Horn BCAB_MB
densus Hatch
piperi Hatc}r
thomomysiHatch

Tribe Nemadini
Genus NEMADUS C.G. Thomson
N. brachyderus (LeConte) NS
N. decipiens (Horn) --ec--- _MB
9* PQ

N. gracilicornis Fall
N. parasitus (LeConte) _MB ON PQ
N. pusio (LeConte) BC

T?ibe Cholevini
Genus CATOPS Paykuli
C. alpinus sitkanus Jeannel AK YK NT BC
C. alsiosus (Horn)x AK YK NT BC AB MB ON PQ NB NS LB
C. americanus Hatch AK YK NT BC AB SK MB ON PQ NB NS
clauicornis LeConte
C. basilaris Say AK YK NT BC AB SK MB ON PQ NB NS LB NF
co d o ueri n u s Mannerheim
spencianus (Kirby)
C. egenus (Horn) AK YK NT BC AB SK ON
C. gratiosus (Blanchard) _BCAB oN PQ NS
C. luridipennis Mannerheimx AK YK NT BC AB SK MB ON PQ
b r unnip e nni s Mannerheim
C. mathersi Hatch -YK_BCAB
C. simplex Say AK BC AB SK_ONPQ NB NS_LBNF
Genus SCIODREPOIDES Hatch
Sciodrepa C.G. Thomson
S. fumatus terminans (LeConte) AK _NTBCABSKMBONPQ
S. watsoni hornianus (Blanchard) BC AB SK MB ON PQ YNS NF

Genus CATOPTRICHUS Murray

C. frankenhaeuseri (Mannerheim) AK BC

Genus PRIONOCHAXTA Horn

P. opaca (Say) ON PQ NS

81
Scydmaenidr,Le

Subfamily LEPTININAE
Genus LEPTINILLUS Horn
L. validus (Horn) AK l.K NT BC AB SK MB ON PQ NB NF

Genus PLAI'YPSYLLUS Ritsema

Platy psitllus Westwood
P. castoris Ritsema* AK BC MB ON PQ
castorinus Westwood

Genus LEPTINUS P.W.J. Muller

Key to North American species: Peck (1982)
L. occidentarnericanus Peck AK YK BC

Family SCYDMAENIDAE antlike stone beetles

A. Dauies

Forty-eight species of Scydmaenidae have been Scydmaenidae, but these works included rlany syn-
recorded in Canada and Alaska. Improvements in onyms, and the higher classification was confused.
the current taxonomic status and in the methods Marsh (1957) made major improvements for the
and frequency of collection of the species should western North America fauna, and recently Franz
greatly in:rease this number. Scydmaenidae tend (1985) viewed Casey's North American gemera in a
to be common locally, possibly following distribu- worldwide perspective. The last publlication,
tions of the mites on which both the adults and lar- together with recent work by Suter and Newton
rrae feed. They are found chiefly in wet mosses, (personal communication), forms the basis of the
deep forer;t litter, decaying logs and tree stumps, present arrangement.
tree holes, sawdust piles, and compost; some species
are associirted with ants and caves.
LeCorrte (1852) and Casey (1897) described AK (5); BC (12);AB (1); SK (1); MB (3); ON (2e); PQ
most of the North American species of (14);NB (8);NS (6);LB (1)

Subfamily SCYDMAENINAE
Tlibe Lophioderini
Genus LOPF:IIODERUS Casey
L. biformis (.iweklin) AK BC
L. insignis Ni.arsh BC
L. myops Ca;ey _BC
L. similis Mr.rsh -BC

Tribe Euconnini
Genus DRASITOPHUS Casey
D. laevicollis Casey BC

Genus EUCONNUS C.G. Thomson

(Subgenus EUCONNUS s. str.)
E. clavipes (Say) MB ON PQ
pilosico Jlis (LeConte)
E. lecontei (Sichaufuss) -oN
E. semiruber Casey AB SK MB
E. ventralis ilasey _NS
(Subgenus CONNOPHRON Casey)
E. ater (Casr:y) ON
E. brevicornis (Say) ON
E. comptus (Casey) ON
E. debilitans (Casey) oN PQ NB
E. elongatus (Casey) ON
E. flavitarsis (LeConte) oN PQ NB
E. gaudens (Casey) ON NB

82
 Scydmaenidae

E. illustris (Casey) ON
E. novellus (Casey) -PQ
E. optatus (Casey) ON PQ
E. oreophilus (Casey) ON
E. pertinax (Casey) ON
E. pumilus (Casey) ON
E. tenebrosus (Casey) ON
E. testaceipes (Casey) ON PQ
E. trifrdus (Casey) ON
(Subgenus SCOPOPHUS Casey)
E. affrnis Casey ON
(Subgenus PSOMOPHUS Casey)
(Xestophus Casey)
E. debilis Casey ON
E. fatuus (LeConte) ON PQ NB NS
E. salinator (LeConte) _MBON_NB
(Subgenus PYCNOPHUS Casey)
(Nudatoconnus Franz)
E. rasus (LeConte) ON PQ
$

E. longiceps Fall AK BC

Tribe Stenichnini
Genus MICROSCYDMUS Saulcy & Croissandeau
Opresus Casey
M. misellus (LeConter oN PQ NB

Genus STENICHNUS C.G. Thomson

(Subgenus CYRTOSCYDMUS Motschulsky)
S. badius (Casey) ON _NS
S. ovipennis {Casey) _BC
S. perforatus (Schaumr -oN
S. turbatus tCaseyt PQ NB NS
(Subgenus BRACHYCEPSIS Brendel)
S. adjutor Fall AK
S. pacificus (Casey)
S. rothi (Marsh)
S. subpunctatus (LeOonte) ON PQ _NS_LB
(Subgenus PARASCYDMUS Casey)
S. corpusculus (Casey) oN PQ NB
(Subgenus TAPHROSCYDMUS Casey)
S. californicus Motschulskv AK

Tfibe Eutheiini
Genus EUTHEIA Stephens
E. morae Marsh BC
E. scitula Meklin AK

Genus \IERAPHIS Casey

V. lacuna Marsh BC

Tribe Leptoscydmini
Genus LEPTOSCYDMUS Casey
L. caseyi (Brendel) ON PQ

Tlibe Scydmaenini
Genus SCYDMAENUS Latreille
Acholerops Casey

83
Silphidae

S. retrusus (Casey) PQ
S. zimmermz:.nni Schaum oll
punctatus Casey

Family MICROPEPLIDAE micropeplid beetles

J.M. Campbell

Micropeplidae is a small family of beetles with the feeding habits of the species; however, both lar-
10 species known in Canada and Alaska. They are vae and adults are assumed to feed ott fungal
found in a variety of habitats, including deciduous spores.
and conifr,,rous leaf litter, inside beaver lodges, in The family is well known in North .t\merica.
Carex clurnps growing in shallow water, on freshly Campbell (1968) revised the New World spr;cies and
cut stumps of fir and cedar and possibly associated published a catalog of the North Americatr species
with the fungus Pyrobema, which is found in forests (1984c).
after a firr:. Matthews (1970) described two species,
M. hoogervdorni and M. hopkinsi, from Pliocene AK (a); YK (1); NT (1); BC (8); AB (a); SK. (2); MB
deposits irr western Alaska. Little is known about (1); ON (5);PQ (a) ' H(1)

Tribe Micropeplini
Genus KALI|jSUS LeConte
K. nitidus Lc Conte BC

Genus MICROPEPLUS Latreille

Anhenooeplus Koch
Key to North American species: Campbell (1973b)
M. browni Cr'Lmpbell -oNPQ
M. brunneus Maklin AK BC
M. cribratus LeConte AB -oN
_ONPQ
M. laticollis li{eklin AK YK BC AB SK
M. punctatus LeConte AK BC
costatus Maklin
oregonensis Hatch'
M. robustus ()ampbell BC
M. sculptus l,eConte BC AB sx ON PQ
obliquus LeConte
M. smetanai Campbell BC
M. tesserula Curtis* AK NT BC AB MB ON PQ
costatus LeConte
costipenzis Makiin

Family SILPHIDAE carrion beetles

J.M. Campbell

The ca.rrion beetles belong to a family of about The family name Silphidae is userl in the
175 species found primarily in the Northern restricted sense of Lawrence (I982a), Lawrence and
Hemisphere, with a few species having ranges Newton (1982), and Anderson and Peck (198{5). These
extending into South America, southeast Asia, and authors exclude the agyrtids, treating them es a sepa-
Australia. Twenty-five species are known to occur in rate family, Agyrtidae. The Canadian and Alaskan
Canada and Alaska. Carrion beetles are common, species are well known taxonomically, based on a
and because oftheir large size and often bright colors handbook ofthe species by Anderson and Peck (1985).
they are frequently collected. Both adults and Iarvae
ofmost species feed on carcasses ofvertebrates; other
species are scavengers, and some species of Aclypea AK (8); YK (6); NT (9); BC (13); AB (17); SK (18);
are phy'tophagous and may be considered to be pests MB (18); ON (18); PQ (15); NB (10); NS (11); PE (7);
ofcultivated crops such as spinach. LB (1); NF (6) . H (4)

84
 Silphidae

Subfamily SILPHINAE
Genus NECRODES Leach
Proton ec rod es Portevin
N. surinamensis (Fabricius) - BC AB SK MB ON PQ NB NS PE - NF
Genus THANATOPHILUS Leach
Philas Portevin
T. coloradensis (Wickham) AK BC
obalskii (Portevin)
T. lapponicus (Herbst)* AK YK NT BC AB SK MB ON PQ NB NS - LB NF
cal ifor ni cus (Mannerheim)
caudatus (Say)
tuberculatus (Germar)
T. sagax (Mannerheim) AK YK NT BC AB SK MB ON PQ
T. trituberculatus (Kirby) AK - NT - AB SK MB ON
Genus OICEOPTOMA Leach
O. inaequale tFabricius) - ON PQ
O. noveboracense (Forster) AB SK MB ON PQ NB NS PE
marginale (Fabricius)
marginata Kirby

Genus NECROPHILA Kirby & Spence

N. americana {Linn6} MB ON PQ NB NS
affinis (I{ttby)
canadensis (Kirby)
peltata (Catesby)
terminata (KJrby)

Genus HETEROSILPHA Portevin

H. ramosa {Say) NT BC AB SK MB ON
ceruaria (Mannerheim)

Genus ACLYPEA Reitter

Blitophaga Reitter
A. bituberosa {Leconte) NT BC AB SK MB
A. opaca tl-innet* AX YK NT

Subfamily NICROPHORINAE
Genus NICROPHORUS Fabricius
Neonicrophorus Hatch
N. americanus Olivier - ON pe
grandis Fabricius
uirginicus Fr6lich
N. carolinus (Linn6J AB
med.iatus Fabricius
mysticallis Angell
N. defodiens Mannerheim AK YK NT BC AB sK MB oN PQ NB NS pE - NF
binotatus Portevin
conuersator Walker
hadjakensis Portevin
lateralis Porte.rin
mannerheimi Portevin
plagiatus Motschulsky
pollinctor LeConte
N. guttula Motschulsky - BC AB SK
hecate Bland
immaculosis }{atch
punctostriatus Pietce
N. hybridus Hatch & Angell - BC AB SK MB ON
N. investigator Zetterstedt* AK YK NT BC AB SK MB oN Pe - NS pE - NF
aleuticus Gistel
confossor LeConte

85
Staphylinidae

infodiens Mannerheim
mar it imu,s Gu6rin-M6neville
melsheimeri lltrby
particeps Fischer von Waldheim
pollinctor Mannerheim
N. marginatus, Fabricius BC AB SK MB ON PQ
cordiger Portevin
labreae Pterce
latifrons Pierce
mchittrictei Pierce
m ontez u r,' ta e Matthews
obtusiscu lellum Pier ce
requiescalor Gistel
N. nigrita Marrnerheim BC
alphaPierce
ruficornit Motschulsky
N. obscurus K.rby AB SK MB
melsheimeri LeOonte
N. orbicollis S,ry SK MB ON PQ NB NS PE
hallii \{trby
quadrisignatus Castelnau
N. pustulatus Herschel AB SK MB ON PQ NB NS
bicolon N ewman
fasciatus Portevin
lordzs Mrnnerheim
unicolor l)ortevin
N. sayi Castel rau AB SK MB ON PQ NB NS NF
lunatus I eOonte
luniger Harold
lunulatus Gistel
N. tomentosur Weber SK MB ON PQ NB NS PE
angustifosciolzs Portevin
aurigaster Portevin
uelutinus Fabricius
N. vespilloider, Herbst* AK 1T< NT BC AB SK MB ON PQ NB NS PE NF
hebes I\trby
pygmaeu.r; I{trby

Family STAPFIfl INIDAE rove beetles

J.M. Campbell and A. Dauies

The role beetles form one of the largest families they can also be found in a variety of habitats
of insects in the world, with over 32 000 described around human dwellings.
species. ,[n Canada, ll29 species have been Several authors are currently investigating the
reported, b,ut hundreds more remain to be either higher classifrcation of the family. The system fol-
described or recorded. Most rove beetles are preda- Iowed in this checklist has been extensively modi-
tors of oth,:r insects and live on or in the soil, in fied from the conventional system used in earlier
ground litt*:r, moss, or in decomposing organic mat- papers. The system is based primarily on a paper
ter. Some r:ove beetles prey on aphids, bark beetles, by Lawrence and Newton (1982), but will uindoubt-
or other ir.sects on trees and herbaceous plants, edly be modifred further as more phylogene bic stud-
whereas others are specialized feeders on pollen or ies are completed. Adults of most genera of rove
are myceto rhagous. Rove beetles prefer moist habi- beetles (except the genera of Aleocharinae) can be
tats along t,he margins of streams, lakes, and snow- identifred with the use of keys by Arnett (1968) and
frelds, and in decomposing vegetation, manure, and Moore and Legner (1974 and 1979) or with keys
carcasses. They are common in all biomes, ranging based on the European fauna such as those ofLohse
from tropical forests to tundra well above arctic and (1964) and Lohse et al. (1974). Most larvae have
alpine timberlines. Some species are associated not yet been described. Newton (1990) provides keys
with mammal burrows, ant and termite nests, to the North American genera of larvae and adults
intertidal and coastal habitats. and saline flats: that inhabit soil litter. Other Nearctic genera can

86
 Staohvlinidae

be identified using the keys by Kasule (1966), based type specimens. In recent years a few genera of
on the rove beetle larvae of Britain. North American Aleocharinae have been revised,
Taxonomically, the North American fauna of and their species can now be identified. These
many tribes and subfamilies is now well known: include the genera Aleochara, Myllaena, Gymnusa,
Quediini and Xantholininae (Smetana Lg7l, 1982); and Deinopsls by Klimaszewski (1979, 1982, 1984)
most Tachyporinae (various generic revisions by and the genus Falagriini by Hoebeke (1985). Ashe
Campbell between 1973 and 1988); Oxyporinae (1984,1986) revised adults and larvae ofthe genera
(Campbell f969); and Pseudopsinae (Herman 1975, of the aleocharine subtribe Gyrophaenina. Other
Newton 1982). The large subfamilies Steninae and subfamilies with most species poorly known in
Staphylininae are becoming well known among the Canada include Paederinae and Oxytelinae, except
Canadian fauna, although comprehensive revisions for the genus Bledius, which was revised by
of most genera are not yet available. However, Herman (7972a-f986).
except for a few genera, Aleocharinae, the largest Moore and Legner (1975a) published a catalog
subfamily of Canadian rove beetles, still remains of the North American Staphylinidae.
almost totally unknown. Although Seevers (1978)
provides a key and a redescription ofthe genera and AK (308);YK(r72); NT (155);BC (581);AB (265); SK
tribes of the subfamily, identification even to genus (126); MB (278); ON (aa1); PQ (a60); NB (167); NS
remains difficult or impossible without access to (195);PE (19); LB (63);NF (171) ' H (88);I (99)

Subfamily EMPELINAE
Genus EMPELUS LeConte
E. brunnipennis (Mannerheim) BC

Subfamily PROTEININAE
Tribe Proteinini
Genus MEGARTHRUS Curtis
M. americanus Sachse MB ON PQ NS PE
M. arcuatus Hatch AI{ BC IU
M. atratus Maklin AK BC AB
M.
M.
excisus LeConte
pictus Motschulsky AK BC
-oN
M. sinuaticollis Boisduval & AK YK NT BC AB
Lacordaire*
angulicollis Maklin

Genus PROTEINUS Latreille

P. atomarius Erichsonf BC ON
paruulus LeConte
P. basalis Makiin AK BC AB
P. brachypterus (Fabricius)* AK BC
P. limbatus Maklin*
maekliniFavel
AK BC -oN

Subfamily OMALIINAE
Tribe Eusphalerini
Genus EUSPHALERUM Kraatz
Anthobium auct.
(Subgenus EUSPHALERUM s. str.)
E. convexum (Fauvel) ON PQ NB NS NF
E. fenyesi (Bernhauer) SC AB _MBONPQ NB NS NF
E. pothos (Mannerheim) AI{ BC AB ON PQ NB NS NF
dirn idiat um (Melsheimer)
E. rugulosum (Meklin) AK
E. segmentarium (Meklin) AK
E. torquatum ( Marsham)t NF
(Subgenus ONIBATHUM Tottenham)

87
Staphylinida I

E. californicum (Fauvel) _BC

E. farrarae Hatch _BC
E. grayae Ha:ch AK BC AB
E. swauki Ha.tch _BC

Tribe Omaliini
Genus ACRLTLIA C.G. Thomson
A. crenulata lHatch) AK _BC
A. tumidula (Maklin) AK _BC
Genus PYCNOGLYPTA C.G. Thomson
Key to Norbh American species: Campbell (19836)
P. aptera Carnpbell MB_PQ NB NS_LB NF
P. lurida (Gy lenhal)* AK YK Nr--ix MB ON PQ NB NS

Genus ELONIUM Leach

Acrolocha C.G. Thomson
E. diffusum ( :'auvel,) -oN PQ NB NS NF
E.
E.
E.
exsculptun tMiiklin
fractum {Fauvel)
helferi (St.elt
r BC

RC
v_ MB ON

E. leechi tHa chj BC

E. minutum r Olivierl'i BC
striatum (Gravenhorst)
E. rugosum t . {atch r BC

Genus HAPALARAEA C.G. Thomson

Phyllodr epa C.G. Thomson
H. angustior Hatcht BC
H. caseyi (Be'nhauer & Schubert) BC
pacifica .Casey)
H. floralis tPr.ykull)+ - BC AB_MBONPQ NS NF
H. hamata (Fauvel) BC MB ON PQ NS
H. humerosa tFauvel)
H. humilis (Ntaklin) AK BC ___9*PQ_
H. longula (Nl,iklin) AK BC
H. megarthrcides (Fauvel) BC -AB
H. strigipennLs (Maklin) AK
Genus OMALIUM Gravenhorst
Ochthen'nus Motschulsky
O. algarum Casey BC
fucicola Jasey
O. callosum llaklin AK
O. excavaturr Stephenst PQ
O. flavidum Ilamilton
O. foraminosum Meklinx AK YK NT BC AB SK
-o^
MB ON
PQ
PQ
O. oxyacanth;le Gravenhorstt - ?-
O. plagiatum Mannerheim AK
O. repandum Erichson pe-
O. rivulare (Faykull)t AK _BC -o^ PQ NS NF

Genus PHLOEONOMUS Heer

(Subgenus PIILOEOSTIBA C.G. Thomson)
P. iapponicus (Zetterstedt)E AK YK NT BC AB SK MB ON PQ NB NS
argus Lt.Conte
(Subgenus PHLOEONOMUS s. str.)
(Distemrnus LeConte)
P. flavipennis (Maklin) AK
P. pusillus (Gravenhorst)* AK BC AB -oN PQ Ns-:Ne
laesicollis (Miklin)
P. suffusus (Casey) AK
Genus MICRI\J-YMMA Westwood
Key to species: Bdcher (1988)

88
 Staphylinidae

M. brevilingue Schipdte* AK YK NT
dicksoni Maklin
M. marinum (Strctm)* PQ NB - NF
stimpsoni LeConte

Tribe Coryphiini
Key to North American species: Campbell (1978c)
Genus CORYPHIUM Stephens
Occlephelinus Hatch
C. arizonense tBernhauer, - BC
C. brunneum (Hatch) - BC
C. hyperboreum (Maklint* AK YK
C. nigrum Campbell PQ - NS

Genus SUBHAIDA Hatch

S. ingratatHatcht AI( BC AB
S. monticola Campbell AB

Genus HOLOBOREAPHILUS Campbell

H. nordenskioeldi (Maklin)x AK YK NT - MB - PQ
nearcticus (B\ait)

Genus BOREAPHILUS C.R. Sahlberg

B. henningianus C.R. Sahlbergx AK YK AB SK MB ON PQ NB
americanus Notman

Tribe Anthophagini
Genus BRATHINUS LeConte
Key to species: Frank et al. (1987)
B. nitidus LeConte PQ NB NS NF
B. varicornis LeConte - ON PQ NB
Genus ACIDOTA Stephens
Key to North American species: Campbell (7982a)
A. crenata (Fabricius)* AK BC AB SK MB ON PQ NB NS PE LB NF
seriata LeConte
A. quadrata (Zetterstedt)x AK YK NT BC AB - MB ON PQ - LB
fr a nkenh ae u s eri Miiklin
patruelis Leconte
A. subcarinata Erichson - ON PQ NB NS NF
Genus OLOPHRUM Erichson
Lathrium LeConte
Key to North American species: Campbell (1983o)
O. boreale (Paykull)x AK YK NT BC AB - MB - PQ
helleni Scheerpeltz
nig r op ic eum Motschulsky
quesneliHatch
O. consimile (Gyllenhal)* AK YK NT BC AB SK MB ON PQ NB NS NF
Iimbatum Maklin
marginatum (Kirby)
paruulum Meklin
O. latum Maklin* AK YK NT
O. obtectum Erichson - ON PQ
emarginatum Erichson
rotundicolle (Say)
O. rotundicoile (C.R. Sahlberg)* AK YK NT BC AB SK MB ON PQ - NS NF
conuexicolle LeConte
conuercum Meklin
Genus ARPEDIIIM Erichson
Key to North American species: Campbell (19840)
A. cribratum Fauvel - BC MB ON PQ NB NS - LB NF

89
Staphylinidae

angulare Fanvel
columbier,.se Hatch

Genus EUCNIICOSUM Reitter

Reuelstoh,:a Hatch
Key to Nort.r American species: Campbell (19840)
E. brachyptenLm (Gravenhorst)* AK YK NT MB
beringense (Van Dyke)
E. brunnescens (J. Sahlberg)* AK YK NT BC AB MB ON PQ LB NF
hoppingi (Hatch)
E. tenue (LeConte)* AK YK NT BC AB SK MB oN PQ .B NF

Genus AMPHICHROUM Kraatz

Staphygrophis Horn
A. floribundum LeConte BC
flauicorne'Casey
A. macuiatum (Horn) BC
immaculctum Hatch
A. maculicolle (Mannerheim) AK

Genus PELEC OMALIUM Casey

Heterops -Vlannerheim
P. laevicolle (LeConte)
P. puberulum tFauvel)
P. testaceum (:Vlannerheim) AK
_BCAB
BC AB
_ :a_ I'_ . _

binotatun',. Casey
nigrwn Hatch

Genus LESTE VA Latreille

Zeslo Bla:kweider
Paralestetn Casey
Pseudolesteua Casey
Teuales C*ey
L. pallipes Le( jonte ON PQ NB NS NF
biguttata LeConte
picescens LeConte

Genus UNAMIS Casey

U. columbiensis Hatch _BC
U. fulvipes Fall _BC
Genus PHLAEOPTERUS Motschulsky
Phloeopte,.us LeConte
Tilea Fatvel
P. castaneus (Oasey) BC AB
cascadien;is Hatc}'
P. cavicollis (Fauvel) AK BC AB
breuipennis Casey
longipalp;rs Hatch
P. frosti Hatch AK BC
P. fusconiger trlotschulsky AK BC
P. houkae Hat,:h AK BC AB
P. lagrandeuri Hatch _BC
P. loganensis llatch _BCAB
Genus VELLICA Casey
V. longipennis Casey BC

Genus OROBANUS LeConte

O. falli Mank _BC
O. rufipes Cas,:y _BC
O. simulator LeConte _BC
O. tarsalis Hatch _BC
Genus ARTOCHIA Casey
A. productifrorrs Casey AK BC

90
 Staphylinidae

Genus XYLODROMUS Heer

X. concinnus (Marsham)t AK BC AB SK
Iacustris Casey
X. depressus (Gravenhorst)t AK BC

Genus ANTHOBIUM Leach

Lathrimaeum Erichson
A. atrocephalum (Gyllenhal)* AK
A. clarki Hatch _BC
A. fimetarium (Mannerheim) AK _BC
A. occiduum (Casey) _BC AB
A. reflexicolle Casey _BC
A. sinuosum Hatch _BC
Genus DEINOPTEROLOMA Jansson
Mathrilaeum Moore
Key to species: Smetana (19856)
D. pictum tFauvel) -BC
_BC
D. subcostatum (Maklin) AK
humerale (Casey)

Genus TANYRHINUS Mannerheim

T. singularis Mannerheim AK BC

Genus TRIGONODEMUS LeConte

T. fasciatus Leech BC
T. striatus LeConte ON PQ NS

Genus PORRHODITES Kraatz

Key to North American species: Campbell (1984b)
P. fenestralis (Zetterstedt)x AK YK NT BC AB MB PQ LB
breuicollis Maklin
P. inflatus tHatcht _BCAB -oN PQ

Genus OROCHARES Kraatz

Paradeliphrum Hatc}:
Key to North American species: Campbell (1984b)
O. tumidus (Hatch) BC

Genus GEODROMICUS Redtenbacher

G. brunneus (Say) PQ
coeszs Erichson
G. fauveli Casey _BC
G. integer Casey _BCAB
G. rufrpennis Van Dyke AK
G. schultzi Fauvel -BC
G. strictus Fauvel o" pe NF
nigrita Farel
G. temporalis Casey BC
G. verticalis (Say) AK YK AB MB ON PQ NB NF
ouipennis LeConte

Genus MICROEDUS LeConte

M. austinianus LeConte AK YK _BCAB_PQ
M. fenderi Hatch _BC
M. laticollis (Mannerheim) AK _BC
M. porterae Hatch _BCAB
M. rogersi Hatch _BC
Genus HAIDA Keen
Key to species: Campbell (19786)
H. keeni Keen AK BC

91
Staphylinidar,,

Subfamily OLISTHAERINAE
Genus OLIST HAXRUS Dejean
O. megacephz,.lus (Zetterstedt)* AX YK NT AB _PQ
laticeps i,eConte
O. substriatur; (Gyllenhal)x YK NT AB oN PQ
nitidus I eConte

Subfamily PIESTINAE
Tribe Piestini
Genus SIAGCTNIUM Kirby & Spence
Prognatl, us Berthold
Key to Nort,h American species: Moore (1975)
S. americanurn Melsheimer AB NF
conuergeizs Sachse
S. punctatum LeConte AB SK MBONPQ_NS NF
S. stacesmith: Hatch _BC AB SK MBONPQ_LB

Tlibe Trigonurini
.Genus TRIG(,NURUS Mulsant
Key to Nor h American species Blackwelder (1941)
T. crotchii Lellonte BC
lecontei flharp
T. dilaticollis Van Dyke BC
T. sharpi Blat.kwelder BC
T. subcostatu,' Meklin AK

Subfamily OSORIINAE
Tribe Osoriini
Genus HOLO IROCHUS Erichson
Neotroch,^ts Blackwelder
H. laevicaudus (LeConte) MB

Tlibe Lispinini
Genus CLAVILISPINUS Bernhauer
C. prolixus (L,-.Conte) MB PQ
C. rufescens (,:Iatch) BC AB

Genus THOR,\COPHORUS Motschulsky

Glyptomtt Erichson
T. costalis (Erichson) MB ON PQ

Tlibe Eleusinini
Genus RENAIDIA Motschulsky
Eumalus Sharp
R. canadensis Horn
R. nigrella (Lt:Conte)
jubilaea Motschulsky
---ecAB- _ 9*_ _ _ _ _ _

Subfamily OXYTELINAE
Key to genera: Herman (1970)
Tribe Coprophilini
Genus DELEi\STER Erichson
D. concolor LcConte BC
D. dichrous (Ciravenhorst)t oN PQ LB NF

92
 Staphylinidae

Genus COPROPHILUS Latreille

Zonoptilus Solsky
Key to North American species: Campbell (1979a)
C. castoris Campbell NB
C. sexualis Leech - 9*PQ
Genus SYNTOMIUM Curtis
Eunonia Casey
S. confragosum Meklin AKBCAB_PQ
S. grahami Hatch AK_PQ
S. keenianum (Casey) _BC )"
S. malkini Hatch AK BC

Genus MITOSYNUM Campbell

M. vockerothi Campbell NB

Genus APLODERUS Stephens

Phloeonaeus Erichson
A. linearis LeConte

Genus BLEDIUS Leach

Microbledius Herman
Psamathobledizs Herman
Key to North American species: Herman (I972a,1976, 1983, 1986)
B. albonotatus Miiklin AK BC
alb idi p ennis Bernhauer
ornatus LeConte
B. analis LeConte AB_MB ON PQ
B. annularis LeConte AK YK NT BC AB SK MB ON PQ NB LB NF
Ianguidus Casey
B. aquilonarius Herman AK YK NT BC AB oN PQ
B. assimilis Casey _MB ON PQ
B. basalis LeConte -NS---
ignauus Casey
misellus Casey
B. confusus LeConte AK YK NT BC AB SK _ONPQ
B. coulteri Hatch BC SK MB
B. emarginatus (Say) -oNPQ
troglodytes Erichson
B. flavipennis LeConte BC SK MB
agonus Casey
cuspidatus LeConte
nelsoniHatch
tenuis Casey
B. foraminosus Casey BC
deceptiuus FalI
relictus Fall
B. fumatus LeConte . MB ON
B. gravidus Casey NT BC AB SK MB ON
missionensis Hatch
B. habrus Herman AK YK BC
B. ineptus Casey AB SK MB ON
B. longipennis M,iklin AK
B. mandibularis Erichson SK MB
breuidens LeConte
B. melanocephalus (Say) MB
confinis Fall
B. monstratus Casey BC
B. mysticus Fall BC
B. nardus Herman BC
B. neglectus Casey NB NS NF
B. nitidicoilis LeConte _MB
B. omega Herrnan AB _MB oN PQ
B. opacifrons LeConte nc _MB
condonensis Hatc}'
lectus Casey
piceus Fall
regularis Fall
specularis Fall

93
Staphylinidr,Le

B. opaculus LeConte _NS PE _NF

B. philadelphicus Fall -oNPQ NB _NF
dickersoni Notman
dissimit'.is Fall
folli Wendeler
mirlas.Notman
B. politus Erichson NB
nigriceJts Notman
B. rotundicollis LeConte MB
B. rubiginosus Erichson MB
B. ruficornis LeConte
B. semiferrugineus LeConte
canaliailatus Notman
_:c_:"yuo^rF8_ _ _ _ _

B. strenuus tlasey BC AB SK MB ON
arizonensis Fall
furtiuus Casey
B. suturalis LeConte AK YK NT BC AB SK ON
luteiper nis LeConte
medialis Fall
oregonensis Hatch
pleuralt.s LeConte
B. tarandus Herman AK YK NT BC AB SK MB ON PQ NF
diuisus LeConte
B. tau LeCorrte MB ON PQ NB NS NF
B. turgidus t.)asey AK YK NT BC AB SK MB ON PQ NF
borealis Blatchley
bowronensis Hatch
B. viriosus lr.erman AK YK NT BC
B. washingtonensis Hatch _BC
AB SK
_9*PQ
B. zophus H rrman _YK_BC
Genus OCH'|HEPHILUS Mulsant & Rey
Ancyrol horus Kraatz
O. biimpressus (Meklin)*
O. columbierLsis (Hatch)
O. planus (LeOonte)
AK
AKYK_BC
AKBC_ONPQ
BC
_v
Genus THIN ODROMUS Kraatz
T. corvinus (Casey)
T. diffusus (tlasey) _ nc _ _ _ g*PQ _ -_ _ _ _

Genus CARI'ELIMUS Leach

P a rat r a,go p hlo e us Hatch
Trogophloeus Mannerheim
C. bilineatus Stephenst BC
C. congener Jasey BC
C. conjunctus (Casey) BC
C. diffusus (tlasey) BC
C. harneyi H.atch BC
C. nanuloides Hatch BC
C. obesus (Kiesenwetter)f -PQ
fo s s ula t u s Motschulsky
mancus (Casey)
spectatl;.s (Casey)
C. vancouvelensis Hatch -BC
_BC
C. volans (Notman)

Genus THIN OBIUS Kiesenwetter

Torrentornus Bietig
T. alaskanus (Fall) .AK
T. frizzelli Hatch _BC
T. teres (Fall) AI{

Tlibe Oxytelini
Genus PLA1 YSTETHUS Mannerheim
P. americamrs Erichson
P. degener Mulsant & Reyt _:'T:" MB ON
-oN
PQ

94
 Staphylinidae

Genus OXYTELUS Gravenhorst

O. fuscipennis Mannerheim AKYK-BC ABSKMBONPQ_NS
O. Iaqueatus (Marsham)t _BC
montanus Casey BC AB ON PQ NB
O.
O. nimius Casey T_ _ _ONPQ
_ONPQ
O. pennsylvanicus Erichson
O. sculptus Gravenhorstf _BC _ONPQ
Genus APOCELLUS Erichson
A. sphaericollis (Say) AB MB ON PQ
amabilis Sachse

Genus ANOTYLUS C.G.Thomson

Key to species: Hammond (1976)
A. insecatus (Gravenhorst)t -oN
A. insignitus (Gravenhorst)
A. niger (LeConte) _BC MB
A. nitidulus (Gravenhorst)f _BC
b or eal is (Motschulsky)
punctatus (LeOonte)
rugulosus (Say)
_BC
A.
A.
A.
rugosus (Fabricius)t
sobrinus (LeConte)
suspectus (Casey)
_BC AB SK
MB
MB
MB
ON PQ
IU
NS
__Iu
A. tetracarinatus (Block)t _BC PQ

Subfamily PSELJDOPSINAE
Key to genera: Newton (1982)
Genus PSEUDOPSIS Newman
Key to species: Herman (1975)
P. montoraria Herman _BC
P. sagitta Herman
P. subulata Herman T NF

Genus ZALOBIUS LeConte

Key to species: Herman (1977)
Z. spinicollis LeConte BC

Genus NANOBIUS Herman

Key to species: Herman (1977)
N. serricollis (LeConte) BC

Genus ASEMOBIUS Horn

Key to species: Herman (1977)
A. caelatus Horn BC

Subfamily PHLOE OCIIARINAE

Key to genera: Smetana and Campbell (1980)
Genus CHARIfYPHUS Sharp
Chapmania Bernhauer
u s is Bernhauer
P seude le
Siberia Blackwelder
Triga Farel
Trigites Handlirsch
Key to species: Herman (I972b)
C. picipennis (LeConte) oN PQ NS

Genus VICELVA Moore & Leqner

V. vandykei (Hatch) AK YK BC

95
Staphylinidu e

Subfamily TACI{YPORINAE
Tribe Tachyporini
Genus TACFIINUS Gravenhorst
Key to North American species: Campbell (1973o), (1983)
(Subgenus T.ACHINUS s. str.)
T. addendus Horn _MB ON PQ _NS
T. angustatus Horn - YK _BC AB
T. apterus M:iklin AK
T. basalis Erichson*
circumcinctus Maklin
AK YK _BC AB ix me ON PQ NN NS

T. beckeri Ci,Lmpbell - YK _BC

T. brevipennis J. Sahlberg* AK YK NT
T. canadensis Horn
_BC
oN pe
T. contortus Hatch AB
uarians Hatch
T. corticinus Gravenhorstt PQ
T. crotchii H.rrn BC
T. dilatatus l{atch BC AB
T. elongatus Gyllenhal* AK YK NT BC AB SK MB ON PQ LB NF
parallelus Hot:n
T. fi mbriatus, Gravenhorst ON PQ_NS
T. frigidus E ichson AI( YK NT BC AB SK MB ON PQ_NS TII NF
propinq tus Mannerherm
T. fumipennis tSayt AB _MB ON PQ NB NS
T. horni Can pbell ON PQ_NS
T. instabilis \{tiklin* AK YK BC
T. jacuticus j rcuticus Poppiusx AK l-I( NT
nearctic'ts Campbell
T. jacuticus r Lllrichi Campbell PQ LB NF
T. lanei Hatch BC AB
T. limbatus itlelsheimer ON PQ NB NS
T. luridus Er ichson _MB ON PQ NB NS
T. maculicoll s Maklin ec- _MB
T. memnonil s Gravenhorst ON PQ NB NS
batychrous Gravenhorst
rulzzs Sz.chse
T. minimus (lampbell ON PQ
T. nigricornil; Mannerheim AK BC AB
T. picipes Er chson _ONPQNBNSPE
T. quebecensis Robert AK YK NT BC AB SK MB ON PQ NB NS NT
acutus Ilatch
T. rainieri H rtch BC
T. rufipes (D rCeert+ BC -PQNBNS NF
T. schwarzi Ilorn _PQ-NS
T. scrutator tlemminger & Harold _AB MB ON PQ NB NS
discoideus Melsheimer
repandu:.s Hotn
T. semirufus Horn AK BC
T. smeLanai tlampbell BC AB
T. stacesmitl i Campbell BC
T. subterranlus t Linne rt BC
T. tachyporoides Horn NT BC AB SX ff,fS
_MB
oN pe Nn Ns ie
-_LBNF lrr
T. thruppi H;rtch - YK BC AB ON PQ NB NS
T. vergatus ( ampbell oN PQ
(Subgenus T.,\CHINODERUS Motschulsky)
T. debilis Ho'n BC

Genus COPROPORUS Kraatz

Erchom'ts Motschulsky
Key to North American species: Campbell (1975)
C. ventriculus (Say) - NT BC AB SK oN PQ NB NS
acuducttrs (Kirby)
affinis (I{trby)
flauidus (Casey)
gibbulus (Erichson)
maneei liicheerpeltz
punctul<t tus (Melsheimer)
politus (.Manee)

96
 Staphylinidae

Genus CILEA Jacquelin du Val

Key to North American species: Campbell (1975)
C. silphoides ( Linn6tt BC AB oN PQ NB

Genus TACHYPORUS Gravenhorst

Key to North American species: Campbell (1979b)
(Subgenus PAI-PORUS Campbell)
T. nitidulus (Fabricius)* AK YK NT BC SK MB ON PQ LB NF
faber Say
(Subgenus TACHYPORUS s. str.)
T. abdominalis (Fabricius)* AK YK NT BC AB MB ON PQ NB NF
T. borealis Campbell AK YK NT BC AB SK MB ON PQ NS
T. browni Campbeil PQ NB
T. californicus Horn _BC
tehamoe Blackwelder
T. canadensis Campbeli -\'K-BC _BC SK MB ON PQ NB NF
T. dimorphus Campbell
T. dispar (Paykull)t _BC _ONPQ NB NS NF
chrysomelinus arct.
T. elegans Horn -ONPQ
T. flavipennis Campbell AK YK NT BC AB SK MB ON PQ NB NF
MB ON
T. inornatus Campbell
T. jocosus Say
acaudus Say
AK YK NT BC
v SK
MB ON
PQ NB

arduus Erichson
infuscatus Bernhauer
maculipennis LeConte
snyderi Blackwelder
T. lecontei Campbell BC AB SK MB ON PQ
T. maculicollis LeConte BCABSKMBONPQ_NS
T. mexicanus Sharp YK_ BCABSKMB_PQ_NSNF
alleni Blackwelder
oregonus Blackwelder
temacus Blackwelder
T. nanus Erichson -oNPQ
T. nimbicola Campbell AK YK NT BC AB PQ NB NS -LB NF
ornatus Campbell MB_PQ
T.
T. pacifrcus Campbell BC T SK

T. pulchrus Blatchley MB ON PQ
T. rulomoides Campbell MB ON PQ NB
T.
T.
T.
rulomus Blackwelder
stacesmithi Campbell
transversalis Gravenhorstf
TY NT BC
BC
AB MB ON
_ONPQ
PQ
_:u NT

Genus SEPEDOPHILUS Gistel

Key to North American species: Campbell (1976)
S. basalis (Erichson) ON
pulicarius (Sachse)
setifer (Casey)
S. bisigratus (Horn) BC
S. cinctulus (Erichson) ON PQ
corticola (Casey)
S. crassus (Gravenhorst) ON PQ
moestus (Say)
S. ctenidialis Campbell ON
S. Iittoreus (Linn6)t BC AB MB ON PQ NN NS NF
carissimus Flatch
hnoxii (LeConte)
S. marshami (Stephens)t -PQ _NS
S. opicus (Say) -PQ
S. scriptus (Horn) oN PQ
S. testaceus (Fabricius)t -ABSK ON PQ NB NS I\TF

elongatus (Blatchley)
imbricatus (Casey)
limuloides (Casey)
S. velocipes (Casey) NS
uirginiae (Casey)
S. wickhami Campbeil ON PQ

97
Staphylinidr.e

Tlibe Bolitobiini
Genus LORDITHON C.G. Thomson
Key to No:th American species: Campbell (1982b)
(Subgenus LORDITHON s. str.)
L. anticus (Forn) ON PQ NB NS
L. appalachi,rnus Campbell MB ON PQ NB NS
L. axillaris {,lravenhorst) PQ
L. cascadensis iMalkin) BC
hatchi (llalkin)
L. diffrcilis Campbell BC
L. facilis {Caseyt -oN PQ NB NS_ _NF
L. niger (Grevenhorstt -oN PQ
L. poecilus (J,{annerheim t BC AB
L. scutellarii Campbeli -oN PQ NB NS _NF
L. thoracicus thoracicus AK YK NT BC AB MB ON PQ NB NS LB NF
(Fabricius)x
cincticoilis (Say)
occiduus (Casey)
(Subgenus B OLITOBUS Tottenham)
L. bimaculat rs (Coupen BC AB MB ON PQ NB NS
kelleyi l[.alkin
L. cinctus (Gravenhorsl)
atricautt.atus (.Say)
-oN PQ

gentilis iLeConte)
L. fungicola ilampbell
L. idahoae Hatch
L. longiceps lleConte)
AK YK NT BC AB
_BCAB
AB
SK MB ON PQ NB NS PE
:uy
elelas (Elernhauer)
AK -oN PQ NS

L. notabilis (:ampbell PQ
L. obsoletus Sayr NS
sellolzs Sachse
L. oregonus l)ampbell BC
L. quaesitor Hornt
rostratus (LeConte)
-oN PQ NS

Genus CARI'HACIS des Gozis

Key to North American species: Campbell (1980)
C. intrusus ( lornJ -MBONPQ
C. nepigoner:sis (Bernhauer) - BC MB ON PQ NB NS _NF
rz6er (1\lalkin)
wallisi (Malkin)
Genus BOLI'IOBIUS Leach
B. analis tPaykullt* NT AB SK MB ON PQ NS
cingulai.us arct.
B. kremeri tldalkin] _BC
Genus BRYCITPORUS Kraatz
B. rubescens (Hatch) _BC
B. rufescens LeConte _BC -oN peNe- LB
paruulu.s Casey
rubidus Leconte
testaceui LeConte
B. varians (F atch) _BC
Genus MYCIITOPORUS Mannerheim
(Subgenus ISCHNOSOMA Stephens)
M. pictus Horn _BCABSK MB ON PQ NB NS NF
M. splendidus (Gravenhorst)* AK YK NT BC AB SK MB ON PQ NS NF
elegans Matthews
inquisiti^ts Casey
pallidus Mannerheim
(Subgenus MYCETOPORUS s. str.)
M. american rs Erichson MB ON PQ NB
lucidulus LeConte

98
 Staphylinidae

M. brunneus (Marsham) _BC

M. consors LeConte MB ON PQ NB NS
M. discalis Hatch -BC
M. inquisitus Casey
M. insignitus Maklin
M. neotomae Fall
AK___f:"yu9*PQIU___
_BCAB
M. punctatissimus Hatch _BC
M. punctulatus Hatch _BC
M. rusosus Hatch AK YK NT BC AB SK MB ON PQ - NS

Subfamily TRICHOPIIYINAE
Genus TRICHOPHYA Mannerheim
Eumitocerus Casey
T. piiicornis (Gyllenhal)f BC ONPQ_NS

Subfamily HABROCERINAE
Genus HABROCERUS Erichson
H. capillaricornis (Gravenhorst)t BC-ONPQ _NF
H. magnus LeConte ON PQ _NF
H. schwarzi Horn _AB_MBONPQ

Subfamily ALE O CHARINAE

T?ibe Oxypodini
Key to North American genera: Seevers (1978)
Genus DEVIA Blackwelder
Dasyglossa Kraatz
D. congruens (Casey) AK_NT MB

Genus GNATHUSA Fenyes

G. caribou Lohse AK YK NT
G. tenuicornis Fenyes _BC
Genus OCYUSAl{raatz
O. canadensis Lohse AK YK
Genus ILYOBATES Kraalz
Gennadota Casey
L canadensis (Casey) -oN
Genus MOLUCIBA Casey
M. g:randipennis Casey _BC
Genus OCAJ-EA Erichson
lsoglosso Casey
O. vancouveri Casey _BC
Genus OXYPODA Mannerheim
Hylota Casey
O. bradorensis Lohse
O. chillcotti Lohse YTNT_ _ _ MB
PQ

O. demissa Casey
O. egestosa Casey
O. famula Casey
AK
_BC
BC _ 9N_ _

O. frigida Bernhauer _BC

O. glenorae Casey _BC
O. hemingi Lohse AK
O. hiemalis Casey
O. impressa Casey AK BC
-oN--
O. irrasa Meklin AK
O. lacustris Casey
O. lassula Casey AK
-oN--
O. lata Lohse MB

99
Staphylinida e

O. leechi Lohse _YK

O. madgei Lc,hse AK
O. manitobae Casey MB
O. nelsoni Lchse AK__-
O. optiva Casey _BC
O. parafunebris Lohse AK YK NT MB
O. regressa Clasey _BC
O. subpolaris Casey AK

Genus PENT ANOTA Bernhauer

P. meuseli Bernhauer* AK

Genus BRACIHYUSA Mulsant & Rey

Tetralinz Casey
B. americana (Fenyes) _BC
B. helenae (C.asey) AK .\T
Genus GNYI:ETA C.G. Thomson
Euliusa Casey
G. brevicornis Casey BC
G. caerula (C.R. Sahlberg)'l NF
G. manitobae Casey _MB
Genus MER()NERA Sharp
M. venustula (Erichson) -oNPQ
Tribe Athetini
Genus ACROTONA C.G. Thomson
A. adjuvans Oasey
A. onthophik Lohse AKYK_ _ _ _ 9"_ _ _ _ _ _

Genus HYDTrOSMECTA C.G. Thomson

H. pseudodior;ica Lohse YK

Genus CANASTOTA Casey

C. canadensis; (Casey) -oN
Genus DATOMICRA Mulsant & Rey
Hilarina:. Casey
Micromt:ta Casey
Oligomia Casey
Taxicere.tla Casey
D. surgens (Casey) BC
D. wrangeli (tlasey) AK
Genus DIME'IROTA Mulsant & Rey
Dalotia ()asey
Dimetrotina Casey
Engamotla Casey
D. alaskana ( Oasey)
D. altaica (Berrnhauer)*
D. bradorensrs Lohse
D. burwelli L,rhse
yv_ -
AK BC

_ pa:)
DN
!q
LB

D. campbelli -ohse AK \'K

D. caribou Lohse _YK
D. dempsterensis Lohse -YK oat
!q
D. moesta Miiklin AK
maeklini (Fenyes)
D. munsteri (Bernhauer)* AK YK NT MB
D. nearctica Lohse AK YK NT
D. omissa Casey _BC
D. picipennis (Mannerheim)x AK BC
carlottae (Casey)
D. prudhoensis Lohse AK
D. regissalmonis Lohse AK

100
 Staphylinidae

D. resplendens Casey _BC

D. retrusa Casey -BC
D. smetanai Lohse AK YK NT MB pe LB
D. stercoris (Fenyes) _BC
columbica (Bernhauer)
D. venti Lohse YK
D. vicaria Casey BC

Genus PSEUDOTA Casey

P. cornicula Casey _BC
P. formalis Casey _BC
P. irrupta Casey _BC
P. nanulina Casey _BC
P. nescia Casey _BC
P. vana Casey _BC
Genus AMISCHA C.G. Thomson
Colposura Casey
A. analis (Gravenhorst)t NF

Genus GEOSTIBA C.G. Thomson

Sibiota Casey
G. circellaris (Gravenhorst)t NF
impressula Casey
G. parvipennis (Bernhauer) AB

Genus BOREOSTIBA Lohse

B. campbelliana Lohse AK YK
B. hudsonica Lohse
B. lagunae Lohse -YK----PQ-----
_YKPQ
B. lamellifera Lohse
B. sibirica (Maklin)* AKYKNT_PQ_LB
Genus ADOTA Casey
Panalota Casey
A. comparabilis (Maklin) -AK
A. maritima (Mannerheim) AK BC
massettensis Casey

Genus ANEPSIOTA Casey

A. quadricollis Casey _BC
A. terminalis Casey _BC
renominata Bernhauer & Scheerpeltz

Genus ATHETA C.G. Thomson

Elytrusa Casey
Stonalota Mannerheim
A. circulicollis Lohse PQ
A. granulata (Mannerheim) AK YK NT MB-PQ-LB
nomadica Casey
A. martini Lohse - I't(
Genus ATHETALIA Casey
A. metlakatlana (Bernhauer) _BC
A. nimia (Casey) _BC
Genus ATHETOTA Casey
A. atriventris Casey BC
A. wickhami (Casey) BC

Genus LAMIOTA Casey

L. achromata (Casey) BC
L. concessa (Casey) BC
L. keeni (Casey) BC
L. profecta (Casey) BC

101
Staphylinidae

Genus LIOGLUTA C.G. Thomson

P s e udon; eg ist a Bernhauer
L. aloconotoic.es Lohse LB
L. granulosa Lohse AK
L. insolens (Casey) _BC
L. nigropolita (Bernhauer) _YK PQ LB
L. nitens (Maklin) AKYK-BC
L. trapezicollis Lohse _YK
L. vasta (Maklin) AK \.K
Genus MOCITA Mulsant & Rey
M. amblystegii (Brundin tx AK YK NT MB:
M. breviuscula (Maklin) AK BC AB
prudens (Casey)

Genus AI-AO.BIA C.G. Thomson

A. sparreschr eideri (Munster)+ AK YK

Genus SCHISiTOGLOSSA Kraatz

S. aubeiodes .llrundin -AK
S. holmbergi Brundin .AK

Genus PSEUIfOUSIPAJ-IA Lohse

P. microptera. Lohse AK YK

Genus BORE fPHILIA Benick

B. caseyi Loh;le AK YK NT
B. caseyiana..,ohse _YK
B. chillcotti Lrhse MB
B. eremita (R re)* AK-MB LB
B. fusca (C.R. Sahlberg)x AK_NT
B. gelida (J. Sahlberg) AKYKNT_MB PQ
B. hyperborei,. (Brundin)* AK_NT
R. insecuta (F ppelsheim)* AK YK MB
B. islandica (.iftaatz)x AK YK NT
B. manitobensis Lohse AK_MB
B. nearctica l,ohse AK YK MB PQ- LB
B. nomensis (Casey) AK
B. subplana tJ. Sahlbergt* AK_NT
Genus CLUSIOTA Casey
C. claviventris Casey BC

Genus DINA-IiA-EA C.G. Thomson

D. borealis Lchse
D. planaris (I\Ieklin) AKYKPQ__-__
Genus BESSOBIA C.G. Thomson
B. cryptica Lchse PQ

Genus HOMIIOTUSA Casey

H. manitobae (Casey) MB

Genus MICRODOTA Mulsant & Rey

Hilara Mrlsant & Rey
Sonomota Casey
M. globicollis (Bernhauer) ON
M. holmbergi (Bernhauer) AK

Genus PARAI)ILACRA Bernhauer

P. glenorica ({)asey) BC

Genus PHILI{YGRA Mulsant & Rey

P. awemeana (Casey) MB
P. botanicaru.m Muonaf N"

L02
 Staphylinidae

P. criddlei (Casey)
P. erudita (Casey)
P. falcifera Lohse
P. junii Lohse _YK
P. laevicollis (Meklin) AK
P. leechi Lohse NT
P. malleoides Lohse AK YK NT
P. polaris (Bernhauer)x AK YK NT
P. prognata (Casey) BC
P. pseudoboreostiba Lohse _YK
P. ripicoloides Lohse
P. rostrifera Lohse _ 5NT
P. satanas (Bernhauer)
P. stylifera Lohse
P. surrufa (Casey)
-T
P. varula (Casey)

Genus XENOTA Mulsant & Rey

Delphota Casey
Donesia Casey
H alob r e cthina Bernhauer
Nemota Casey
Roualida Casey
X. apposita (Casey) _BC
X. brumalis (Casey) _BC
X. cursor (Meklin) AK
X. districta (Casey) _BC
X. fanatica (Casey) _BC
X.
X.
X.
fungi (Gravenhorst)t
mollicula (Casey)
mordax (Casey)
_BO-_
_BC
v
X. pallitarsis (Kirby) MB
X. postulans (Casey) _BC
X. pratensis (Maklin) AK
X. pugnans (Fenyes) AK
militaris (Casey)
X. relicta (Casey) _BC
X.
X.
remulsa (Casey)
resplendens (Casey)
-BC
_BC
X. rurigena (Casey) _BC
X. subintima (Casey) _BC
X. sumpta (Casey)
X. tenuicula (Casey)
X. wrangelica (Casey)
_:r_ _ MB
AK

Genus LYPOGLOSSA Fenyes

M eg acrotona Scheerpeltz
L. angularis (Maklin) AK YK -BC-ONPQ NF
L. fenyesi (Bernhauer) _BC
Genus PONTOMALOTA Casey
P. Iuctuosa Casey BC

Genus STROPHOGASTRA Fenyes

S. penicillata Fenyes MB

Genus TARPHIOTA Casey

T. debilicollis Casey BC
T. fucicola (Maklin) AK
T. geniculata (Maklin) AK BC
insolita Casey
T. seditiosa Casey BC

Genus TRICHIUSA Casey

T. columbica Casey BC
T. postica Casey ON

103
Staphylinidae

Tribe Aleocharini
Genus AJ-ECiCHARA Gravenhorst
Key to North American species: Klimaszewski (1984)
(Subgenus CI)PROCHARA Mulsant & Rey)
(.Eucharina Casey)
(Funda Blackwelder)
A. bilineata Gyllenhalt - BC AB - MB ON PQ - NS PE - NF
anthomviae Sprague
ontarionis (Casey)
A. bimaculata Gravenhorst NT BC AB SK MB ON PQ - LB
deserticc'la (Casey)
innocua (Casey)
obsolescens (Casey)
recta (Casey)
A. suffusa (Casey)* AK BC AB - PQ
acumincta (Casey)
perturbc,ns Fenyes
A. sulcicollis Mannerheim AK BC
cylindre']a (Casey)
debilicornis (Casey)
rugosa (Jasey)
tibialis ( Casey)
A. verna Say': AK YK - BC AB SK MB ON PQ - NS PE LB NF
(Subgenus XINOCHARA Mulsant & Rey)
A. castaneipcnnis Mannerheim AK YK - BC AB ON PQ
acomantl (Casey)
concurr(,ns (Casey)
eludens .Casey)
glenorar.a (Casey)
insulant. (Casey)
mannertteimi (Casey)
oregona Hatch
rotundicollis (Casey)
A. fumata Gravenhorsti - YK - BC AB - MB ON PQ NB NS PE
affiuens (Casey)
defecta (Casey)
languidu Sachse
A. inexspectata Klimaszewski - - PQ - NS
A. lacertina Sharp - BC AB SK MB ON PQ NB NS
den s iu er, t r is Bernhauer
idonea ('t)asey)
imbricata (Casey)
salicola lCasey)
A. lanuginosaLGravenhorstf - BC AB ON PQ NF
(Casey)
A. sculptivenbris - ON PQ NB - NF
A. tristis Gravenhorstf PQ
(Subgenus Cr\I-OCHARA Casey)
(.OreochcLra Casey)

A. laramiens s (Casey) AB SK
A. rubricalis Caseyt BC - ON
A. rubripenn s (Casey) MB ON PQ
rubripes Blatchley
wallawallae Hatc}r
A. speculicollis Bernhauer AB ON PQ
A. villosa Mannerheimf AK BC AB
(Subgenus AI,EOCHARA s. str.)
A. curtula (Greze)t - BC - ON PQ NB NS NF
bugnioni Fa:uel
fuscipes (Linn6)
A. gracilicornis Bernhauer* NT BC - SK MB ON PQ - NS
americata Casey
ellipsicoi.lis Casey
kansana Casey
pleuralis Casey
A. Iata Gravr nhorst+ - ON PQ
A. assiniboin Klimaszewski - YK - BC - SK MB ON -
r ufo nig r z Klimaszewski

t04
 Staphylinidae

A. tahoensis Casey YK NT BC AB SK MB ON NS
montanica Casey
postpicta Casey
A. thoracica Casey
collusor Casey
-oN PQ

A.' sekanai Klimaszewski AK YK AB MB ON

unicolor Klimaszewskr
(Subgenus ECHOCHARA Casey)
A. ocularis Kiimaszewski MB ON PQ
(Subgenus EMPLENOTA Casey)
(Polistoma Casey)
(Polystoma Stephens)
A. curtidens Klimaszewskr _BC
A. litoralis (Miiklin) AK BC PQ NB NS NF
arenaria (Casey)
longiceps (Casey)
maritima (Casey)
quadrifer (Casey)
trilimbata (Casey)
A. pacifica (Casey) BC

Genus PHLOEOPORA Erichson

P. arctica Lohse NT

Genus PAROCAI-EA Bernhauer

P. nearctica Lohse AK YK NT
P. pseudobaicalica Lohse AKYKNT_PQ

Tribe Hoplandriini
Genus HOPLANDRIA Kraatz
Key to species: G6nier (1989)
(Subgenus HOPLANDRIA s. srr.r
H. lateralis (Melsheimer) ON PQ
arizonica Casey
carinata Casey
ochracea Ktaatz
texana Casey
(Subgenus LOPHOMUCTER Notman)
H. klimaszewskii G6nier _PQ
H. Iaevicollis (Notman) -PQ

Tlibe Falagriini
Key to North American species: Hoebeke (1985)
Genus CORDALIA Jacobs
C. obscura (Gravenhorst)f ON PQ

Genus MYRMECOCEPHALUS Macleay

Lorinota Casey
Stenagria Sharp
M. arizonicus (Casey) BC
acomanus (Casey)
fontinalis (Casey)
M. cingulatus (LeConte) oN PQ NS
sinuosus Casey
tenuicornis Casey
M. gatineauensis Hoebeke PQ NB
Genus FAI-AGRIA Leach
F. dissecta Erichson _BCABSKMBONPQNB NS
angulata Casey
ery t hr optera Melsheimer

105
Staphylinidae

globosa l{elsheimer
iowana (iasey
ithacana Casey
sterilis Casey
subsimilis Casey
tetana Casey
F. sulcata (Paykull)t AB ON PQ
sicula Jekel

Genus ALEODORUS Say

Chitalia Sh.ary
A. bilobatus (llay) ON PQ
canadent:is (Casey)
nigresceris (Casey)
turbatus (Casey)
A. intricatus (Casey) AB SK
A. scutellaris iLeConte) ON
granulos ts (Casey)
illustris (Casey)

Tribe Myrmedoniini
Genus DRUS;LLA Leach
D. cavicollis ((lasey) AK PQ

Genus XENO DUSA Wasmann

Pseudolomechuso Mann
Key to Nonh American species: Hoebeke (1976)
X. cava (LeCo nte) ON
lobata Cr,sey
probata l)asey
X. reflexa (W:,Llker) BC SK MB ON PQ

Genus ZYRAfi Stephens

Platyusa Casey
Z. criddlei (Ce.sey) MB
Z. loricatus Casey -oN
Tlibe Gyrophaenini
Subtribe Gyrophaenina
Key to North American genera: Ashe (1984)
Key to North American species: Seevers (1951)

Genus GYRO PIIAENA Mannerheim

(Subgenus G\ROPHAENA s. str.)
G. affinis C.R. Sahlbergf BC MB PQ NB
lacustris Casey
inconspir.:ua Casey
subpunctata Casey
G. antennalis Casey NB_
G. californica Casey ec-----
G. caseyi Seevers
_MB -PQ
G. criddlei Casey
G. egena Casey ON PQ
exilis Ca,:ey
G. flavicornis Melsheimer oN PQ
G. gaudens C,rsey oN PQ
G. geniculata Meklin AK
G. gilvicollis Casey ; ---
G. insolens Casey ON-NB
G. keeni Casey YK BC AB ON PQ NB
G. modesta Casey NB
G. nana (Paykull)* AK YK BC AB MB ON
perpolita Casey
G. nanoides Seevers ON PQ
G. sculptipennis Casey _PQ

106
 Staphylinidae

G. uteana Casey PQ
pacifica Casey
G. vitrina Casey - ON PQ
dttonsa Casey
(Subgenus PHAENOGYRA Seevers)
G. gracilis Seevers - 2
G. subnitens Casey MB ON

Genus EUMICROTA Casey

E. socia lErichsont PQ

Subtribe Bolitocharina
Genus BOLITOCHARA Mannerheim
Ditropalia Casey
Pleurotobia Casey
Silusid.a Casey
Stictalia Casey
Venusa Casey
B. arcuata (Casey) _BC
B. carlottae (Casey) _BC
B.
B.
densicollis (Casey)
notata M:iklin
-BC
AK
Genus LEPTUSA Kraatz
Dianusa Casey
Eucryptusa Casey
Ulitusa Casey
L. fontana (Casey) -oN
Subtribe Homalotina
Genus HOMALOTA Mannerheim
H. depressiuscula Mannerheim AK
Genus PLACUSA Erichson
P. turbata Casey - BC

Subtribe Cyphina
Genus CYPHEA Fauvel
C. wallisi Fenyes MB

Tlibe Autaliini
Genus AUTAI-IA Leach
A. brevicornis Casey _BC
A. truncatula Casey _BC

TYibe Phytosini
Key to western Norbh American species: Moore (1956)
Genus AMBLOPUSA Casey
A. borealis Casey AK BC
A. brevipes Casey AK BC
pallida Casey

Genus DIAULOTA Casey

D. densissima Casey AK BC
insolita Casey

107
Staphyiinida:

Genus LIPAIiOCEPHALUS Maklin

L. brevipennis Miiklin AK
L. cordicollis LeConte BC

Genus THIN JSA Casey

T. fletcheri Casey AK BC
diuergens Casey
nigra Ct.sey
robustulz Casey

Tribe Gymnusini
Key to North American species: Klimaszewski (1979)
Genus GYMIIUSA Gravenhorst
G. atra Casel'* AK YK NT BC MB ON PQ NB NS _LBNF
G. brevicollis (Paykull)t PQ _LB
G. campbelli Klimaszewski AK YK NT -oN PQ NB NF
G. grandicepr; Casey MB ON PQ lrs NF
G. konopacki. Klimaszewski AK YK NT MB
G. pseudovariegata Klimaszewski AKYKNT_AB MB ON PQ NB NS _ I,B NF
G. smetanai .l(] imaszewski AK YK NT MB ON NF

Tribe Deinopsini
Kev to North American species: Klimaszewski (1979)
Genus DEIN,JPSIS Matthews
D. canadensi,i Klimaszewski -oN NS
D. harringtorLi Casey -oNPQ NB NF
D. rhadina K limaszewski -oN
Tribe Myllaenini
Kev to North American species: Klimaszewski (1982)

Genus MYLI.AENA Erichson

M. arcana Casey AB ON PQ NB
intmunci.a Casey
M. audax Car;ey NT BC oN PQ NB
abdita Casey
dissimuLans Casey
molesta Casey
uegeta Casey
M. decreta Casey BC AB
impellen.s Casey
scobinel't.a Casey
umbra Qasey
M. fenyesi Bcrnhauer BC
esuriens Casey
M. insomnis ilasey
M. Iudifrcans Casey
AK yl'BCysKyuSN PQ
PQ
NB NS

breuiues;lis Casey
M. procidua Casey PQ
M. vulpina B:rnhauer NS

Tlibe Oligotini
Genus CYPHA Leach
Hypocyphtus Gyllenhal
C. crotchi Horn BC AB

108
 Staphylinidae

Subfamily OXYPORINAE
Key to North American species: Campbeil (1969), (19780)
Genus OXYPORUS Fabricius
(Subgenus PSEUDOXYPORUS Nakane & Sawada)
O. lateraiis Gravenhorst MB ON PQ NB NS
breuis Melsheimer
O. occipitalis Fauvel
O. quinquemaculatus LeConte _ T_:"ysKy" oN PQ
oNPQ-NS
(Subgenus OXYPORUS s. str.)
O. femoralis femoralis Gravenhorst ON PQ
pulcher Ziegler
O. kiteleyi Campbell PQ
O. rufipennis LeConte
O. stygicus Say
-oN PQ_NS
PQ_NS
O. vittatus vittatus Gravenhorst MB ON PQ NB NS
cinclzs Gravenhorst
dimidiatus Melsheimer
fasciatus Melsheimer

Subfamily STENINAE
Key to North American species: Casey (1884)
Genus DIANOUS Samouelle
D. chalybaeus LeConte AK NT oN PQ NB NF
c oer u les cen s Blatchley
D. nitidulus LeConte AK YK BC AB SK -PQ _NS NF
zephyrus Casey

Genus STENUS Latreille

Laestris Melsheimer
(Subgenus "HEMISTENUS" Motschulsky)
S. croceatus (Casey) oN PQ
S. kryzhanovskii Puthzx - Nr
-AK I'K ec AB _ix Yu - - - - -
S. niveus Fauvels NT MB

_v
uislo Sanderson
S. pubescens fraternus (Casey) AK YK NT BC AB SK MB ON
S. pubescens sandersonianus Puthz AK YK
S. retrusus (Casey) AK _BC
S. umbratilis (Casey)x AK YK NTBCAB-MB -PQ LB NF
(Subgenus IIYPOSTENUS Rey)
(Arezs Casey)
S. advena (Casey) NT-MBONPQ NB NS
S.
S.
S.
aiexanderi Puthz*
annularis Erichson
arculus Erichson
-BCAB-MBONPQ
_ _ _:"TSKyuSNPQ NB
NB
NS _T"
S. cubensis Bernhauer
dissentiens (Casey) AK BC AB SK MB ON
-oN lls
S.
hubbardi (Casey)
PQ NB - -Ne
leuiceps (Casey)
lucidus (Casey)
politulus (Casey)
politus Sanderson
simiolus (Casey)
S. flavicornis Erichson BC SK MB ON PQ NB NS
geniculatus Say
S. fulvicornis Stephensf BC
S. insperatus Puthz
S. lugens (Casey)
lrn oN PQ NB
NB
perbreuis L. Benick
S. pinguis (Casey) BC AB SK MB
trajectus (Casey)
utensis (Casey)
S. plicipennis (Casey) AK YK NT BC AB MB
agnatus (Casey)
/orzs L. Benick

109
Staphylinidae

S. pollens (Casey) AK NT BC AB SK MB
patens (Casey)
S. punctatus Erichson
artus ((.asey)
-oN PQ

dilutus lCasey)
obsoletus (Casey)
S. quebecensis Puthz AK_NTBCABSK MB ON PQ NB NS NF
S. reconditus breviusculus Puthz AX BC
S. reconditus, reconditus (Casey) AK_NTBCAB MB ON PQ NB
canader.sis (Casey)
propinquus (Casey)
S. rossi Sanc erson AX YK NT BC AB SK MB ON PQ NB ft PE NF
S. varipes (Casey) -MBONPQ
(Subgenus P ARASTENUS Heyden)
S. pterobrachys Gemminger & Harold AK _BC
breuipermis Maklin
S. sibiricus J. Sahlberg* AK NT BC AB MB ON PQ NS LB NF
alpicolo Sanderson
confusus Casey
deducto" (Casey)
exaspentlus L. Benick
montan'"ts Casey
noctiua1lus Casey
wenzeli Prt]az
(Subgenus SIENUS s. str.)
S. ageus Caseyx AK YK NT AB MB ON PQ NB NS LB NF
rigidus Casey
S. alveolatus Casey _BC AB
S. angustus'Jasey MB ON PQ NB NS NF
indigen.; Casey
mendan Casey
pettiti C.asey
placidu.; Casey
tutnicol,'is Casey
S. assequens Rey* AK YK NT BC MB ON PQ NF
nanulu:, Casey
pusio Clsey
S. austini Cr.sey AK YK NT BC SK MB ON PQ NS LB NF
amplifir::atus L. Benick
S. bilentigat rs Casey -oN PQ
murphyanus Bernhauer
S. bilineatus J. Sahlberg* AK NT BC AB SK MB ON PQ
coruus Casey
tacomar: Casey
S. brivioi Pu,hz AK YK NT AB SK MB oN PQ NB NS
:u
NF
S. callosifror.s Puthz ON
S. canaliculatus Gylienhal* AK YK NT BC AB MB oN PQ NB NS NF
congener Maklin
S. cariniceps Miiklin AK BC
S. carinicollis Casey ON PQ -NS-- NF
S. caseyi Puthz ON PQ NB NS
S. clarostigma Puthz AB
S. clavicornis (Scopoli)t _PQ NB
S. colon Say ON
carolintts Bernhauer
S. colonus E:ichson SK MB ON PQ NB NS PE
arizonae Casey
championi Sharp
chapini Blackwelder
floridan,us Casey
inermis Sharp
integer Dasey
interpres L. Benick
militaris Casey
obstrusus Casey
suspectus Blatchley
weiseri Betnhaner
S. columbianus Sanderson BC

110
 Staphylinidae

S. comrna LeConte* YK NT BC AB SK MB ON PQ
punctiger Casey
S. coriaceus Puthz* AK-NT
S. dolosus Casey AKYKNT_ABSKMBONPQNB
jejunus Casey
uillosus Casey
S.
S.
S.
egenulus Puthz
egenus Erichson
eilipticus Casey
TY NT BC

BC
AB SK MB ON
-MBONPQ
PQ LB NF

S. erythropus Melsheimer ON PQ NSNS_

S. exploratus Fall* AK YK BC AB
incanoides Prt}:rz
S. fasciculatus J. Sahlberg* AK YK NT BC AB SK MB ON PQ NB NF
S. femoratus Say MB ON PQ
S. fulvoguttatus Notman -oNPQ
S. geminorum Puthz BC
S. hyperboreus J. Sahlberg* AKYK NT BC AB sr MB ON PQ LB
falli Scheetpeltz
frigid,us Fall
S. insularis Casey AK BC
S. intrusus Casey* AK YK NT SK MB ON PQ NB NS
S. jacuticus Poppius* NT
S. juno Paykull* AK YK NT BC AB SK MB ON PQ NB NS PE LB NF
S. kamtschaticus Motschulskyx AK YK
aquilonius L. Benick
S. koebelei Puthz BC
raniermontis Ptt]az
S. Iabilis Erichsonx AK YK NT
S. laccophilus Cdsey AK_NTBCAB SK MB ON PQ NB LB
anastamozans Casey
edar Notman
rugifer Casey
uerticosus Casey
S. latipennis J. Sahlberg* AK YK NT
S. latissimus hatchi Sanderson
S. lustrator Erichsont
AKYK_BCAB -pe
S. mammops Casey*
humilis Blatchley
AK \'K NT BC AB - ftfS oN PQ Ne Ns Fe is Nr
rulomus Blackwelder
S. maritimus Motschulsky
S. melanarius Stephens*
S. melanopus (Marsham)l
S. mendosus Puthz
S. morio Gravenhorst*
vvv_
AK

AK YK NT BC
BC AB
_

AB
_MB

s^ me
oNpeNs--
ON
ON
ON PQ NB
NF

diues Casey
enod.is Casey
haplus Casey
indistinctus Casey
luculentus Casey
subgriseus Casey
terricola Casey
S. mundulus Casey _BC
S. neglectus Casey MB ON PQ_
S. occidentalis Casey AK-NTBCABSKMB
S. parallelopipedus Mdklin AK BC AB
adspector Maklin
animatus Casey
conuictor Casey
S. paululus L. Benick AK YK
campbelliPrthz
S. pauperculus Casey MB ON PQ
brumalis Casey
S. planatus Puthz
S. pluto Casey
BC -oN
MB ON PQ
S. pudicus Casey
enodis Casey
-oN
juuencus Casey
minor Casey
sibyllinus L. Benick
S. pumilio Erichson* AK YK NT MB ON PQ
atomarius Casey

111
Staphylinid re

S. pusillimus Puthz MB ON PQ NB NS
S. scabiosus Casey AK YK rVr ec AB ix MB ON PQ NB NS NF
S. schwarzi Oasey MB ON PQ NB NS:u_
S. scrupeus Casey AK _NTBCABSK MB ON PQ
S. sculptilis Casey BC
S. sectator Oasey BC
_NTBCABSK lo*peNn
_ *t_ y
S. semicolon LeConte AK
S.
S.
sordidus .l)uthz
sphaeroprir Casey
S. strangula.tus Casey
MB ON PQ
_ONPQNB
_ONPQNB
NS v
S. stygicus Say AK \'K NT BC AB SK MB ON PQ NB NS NF
S. tenuis Casey _ABSK _ONPQ
dfficilis Casey
eriensis Casey
ingratus Casey
inornatus Casey
paralle,-us Casey
pauper Casey
subtilis Casey
S. venustus Casey _BC
S. vespertinrrs Casey _BC
incultu:; Casey
luctuos;ts Casey
S. vexatus C.asey _BCAB MB
S. vicinoides Puthz _BCAB SK MB
S. vicinus Crrsey ON
simplet Casey
tricuspidatus L. Benick
S. vinnulus ilasey* AK YK NT AB MB ON PQ NB NS LB
confusoLdes Renkonen
(Subgenus lESNUS Rey)
(MutintLs Casey)
S. californicrrs Casey BC
S. curtus Casey _NT_AB_MBONPQ
S. formicetorurn Mannerheim+ YKAB_MB_PQ
borealis M,iklin
S. gratiosus Casey
hirsutu; Casey
-oN PQ NS

S. immargin atus Miiklin* AK YK NT BC AB MB PQ NB NS

monticrla Casey
uariabi,;.is Prthz
S. tahoensis Casey BC
pugetensis Casey
uacuus Casey

Subfamily EUAESTHETINAE
Genus NOR.lf ENSKIOELDIA J. Sahlberg
Key to species: Puthz (197h)
N. columbiana Puthz _BC
Genus STIC'IOCRANIUS LeConte
Key to species: Puthz (7974b)
S. puncticeps LeConte ON

Genus EUAIISTHETUS Gravenhorst

E. americanus Erichson
E. brevipennis Casey
E. floridae Casey
T_ I'- T_ MB ON PQ NB NS
ON PQ NB
_NS
NF

E. ganglbaueri Bernhauer ____ oN PQ

E. iripennis Casey oNPQ-NS
E. laeviusculus Mannerheim
E. pacificus Jlall
E. pugetensis Hatch
AK l'K
AK
NT
_BC
BC T:" MB oNPQ-NS
oNPQNBNS--

t12
 Staphylinidae

Subfamily LEPTOTYPHLINAE
Genus CHIONOTYPHLUS Smetana
C. alaskensis Smetana 4K

Subfamily PAEDERINAE
Key to genera: Blackwelder (1939b)
Tlibe Pinophilini
Genus PAJ-AMINUS Erichson
P. luteus Casey -oN
Tlibe Paederini
Subtribe Medonina
Genus LITHOCHARIS Deiean
Metaxyodonta Casey
(Subgenus PSEUDOMEDON Mulsant & Rey)
(Ramona Casey)
L. capitula {Casey) - BC
L. obsoleta tNordmann)t - BC
L. thoracica {Casey) MB ON PQ NB
clarescens (Casey)
ruficollis (Casey)
(Subgenus LITHOCHARIS s. str.)
L. ochracea (Gravenhorst)'l - BC - ON PQ NB
alutacea Casey
quadricollis Casey

Genus ACHENOMORPHUS Motschulsky

(Subgenus ADEROCHARIS Sharp)
A. corticinus tGravenhorst) MB ON PQ
millepunctatus (Say)
rufescens Nordmann

Genus SUNIUS Curtis

Medonella Casey
Micromedon Casey
Oligopterus Casey
Xenocharis Bierig
(Subgenus CAI-ODERMA Casey)
S. rugithorax Hatch - BC
(Subgenus IIYPOMEDON Mulsant & Rey)
(Asteria Fauvel)
(Hemimedon Casey)
(Lena Casey)
S. melanocephalus (Fabriciustf PQ
S. peregrinus (Casey) - BC
discolor Casey
S. quadripennis Casey - BC
(Subgenus TRACHYSECTUS Casey)
S. confluentus rSay) MB ON PQ NB NS
confluens Erichson

Genus MEDON Stephens

Oxymedon Casey
(Subgenus MEDON s. str.)
M. americanus Casey PQ
M. fusculus (Mannerheimt+ - - PQ
M. guppyi Hatch - BC

113
Staphylinidr.e

(Subgenus PLATYMEDON Casey)

(Paramzdon Casey)
M. capitalis Flatch _BC
M. convergens (Casey) AK
M. insularis Casey -BC
M. laticollis iCasey) MB
M. mcleodi llatch _BCAB
M. pugetensis Hatch _BC
M. shastanic,us (Casey) _BC
M. truncatu'r Hatch BC
M. vancouve,ri (Casey) _BC
borealil Casey
(Subgenus X'IEDONODONTA Casey)
M. ctenopho:um Hatch _BC

Subtribe Scopaeina
Genus ORUII Casey
(Subgenus FYCNORUS Casey)
Key to North American species: Herrnan (1965)
O. dentiger (LeConte) AK - MB ON PQ
humilis (Notman)
iowanus Casey
(Subgenus C RUS s. str.)
Key to North American species: Herman (1965)
O. distinctu, Casey BC
O. punctatu,r Casey BC
filius C*ey
pallidus Casey
sonomae Casey

Genus SCOI'AEUS Erichson

Leptorus Casey
Pseudo"us Casey
(Subgenus SCOPAXOMA Casey)
S. brunniper LeConte - BC AB
(Subgenus SCOPAXUS s. str.)
S. notangulrLs Casey PQ
(Subgenus -cCOPAEOPSIS Casey)
S. ventralis Casey) - ON

Subtribe Stilicina
Genus PACI{YSTILICUS Casey
P. hanhami (Wickham) _BCAB_MBNB
Genus RUG.[LUS Leach
Stilicosoma Casey
R. biarrnatus (LeConte) MB ON PQ NB
abbreuiellus Casey
R. dentatus Say -oN
luculentus Casey
minusculus Casey
nigrolucens Casey
R. fragilis (Gravenhorst)t _ONPQ_NS
dngustatus (Fourcroy)
s cutella t us Motschulsky

R. latiusculus (Casey)
R. oregonus (Casey)
R. rufrpes Germarf
AK- - sc- - - il- - - - - -
Genus MEGASTILICUS Casey
M. formicar:us Casey -oN

rt4
 Staphylinidae

Subtribe Lathrobiina
Genus LATHROBIUM Gravenhorst
(Subgenus APTERAI-IUM Casey)
L. sibiricum Fauvelx AK
(Subgenus ABLETOBIUM Casey)
L. shermani Fall ON
(Subgenus LATHROBIUM s. str.)
(Litolathra Casey)
L. amplipenne Casey ON
deceptiuum Casey
L. arrnatum Say oN PQ
procerum Casey
L. confusum LeConte ON PQ
L. divisum LeConte BC
L. fauveli Duvivier -ABSKMB oN PQ NB NS PE NF
simplex LeConte
L. fulvipenne (Gravenhorst)t - BC AB NF
p unct ulat u m Mannerheim
L. hesperum (Casey) BC
L. leconteanum Scheerpeltz NF
concolor LeConte
L. nigrolineum (Casey) _MB
L. obtusum Casey _MB
L. shoshonicum (Caseyr -BC
L. simile LeConte -MBONPQ-NS--
_MBONPQNBNS
L. sparsellum Casey AB
L. spissicorne Casey ON PQ PE
L. vancouveri Casey AK _BC
L. washingtoni Casey AK NT BC AB SKMBONPQ_NS NT
(Subgenus LATHROBIOMA Casey)
L. othioides LeConte ON PQ
L. scolopaceum (Caseyt
L. tenue LeConte nc - - MnoNFS f"NS:
(Subgenus LATHROLEPTA Casey)
L. debile LeConte ON PQ
(Subgenus TETARTOPEUS Czwalina)
(Deratopeus Casey)
Key to North American species: Watrous (1980)
L. angulare LeConte PQ NB NS
nigrescens (Casey)
semirubrum (Casey)
L. convolutum Watrous -ONPQ
L. furvulum {Casey) NT AB SK MB ON PQ NB NS PE LB N"
L. Iacustre (Caseyt MB ON PQ NB NS
L. nigrum LeConte AK YK NT BC AB SK MB ON PQ NB NS PE NF
finitimum LeConte
stibium (Casey)
L. nitidulum LeConte ON_NBNS NF
bicolor LeConte
bicoloratutn Bernhauer & Schubert
paruipenne (Casey)
L. punctulatum LeConte AK NT BC AB SK MB ON PQ
capitosum (Casey)
hebes Casey
L. rubripenne tCasey) ON PQ
agitans (Casey)
callidwn (Casey)
L. sollicitum Fall AK YK AB
L. tetricum (Caseyt ON PQ
canoaensis (Bierig)
nigriceps Notman
$
L. puncticolle Kirby MB

115
Staphylinidr:e

Genus LOBIiATHIUM Mulsant & Rey

(Subgenus EULATHROBIUM Casey)
(Lathro.iropis Casey)
L. grande (LeConte)
L. subseriatum (LeOonte) ___sc___g*PQ_fr___
ustulatum Casey
(Subgenus LOBRATHIUM s. str.)
(Lathro,Siella Casey)
(Lathro;laxis Casey)
L. caseyianum ( Scheerpeltz) _BC
bipartitum Casey
L. collare (Erichson) MB ON PQ NS
L. coloraden,;e Casey AK NT BC
L. impressinLum Hatch _BC
L. longiuscu um (Gravenhorst)f
multipunc,tum (Gravenhorst)t
-ox
L. -BO
_BC
L. rubricolle (Casey)
L. sewardi (Fall) AK
(Subgenus PSEUDOLATHRA Casey)
(Linolathra Casey)
(Microluthra Casey)
(.Parala::hra Casey)
L. ambiguunr (LeConte) _ONPQ
L. dimidiatum (Say)
horni Duvivier
-oN
rufoparlitum Fairmaire & Germain
L. famelicunr (Casey) _PQ
L. lineiformt, (Casey)
L. oregonense (Casey) BC 9*_
Subtribe Paederina
Genus PAEI)ERUS Fabricius
Leucopt.ederus Casey
N eo p ae t:.le r us Blackwelder
Paederillus Casey
P. littorarius Gravenhorst NT BC SK MB ON PQ NB NS NF
pugetensis Casey

Subtribe Cryptobiina
Genus HOMAEOTARSUS Hochhuth
(Subgenus GASTROLOBIUM Casey)
H. badius (Gravenhorst) ON
H. bicolor (Gravenhorst) oN PQ
Dr\
H. carolinus (Erichson) - rq

(Subgenus HESPEROBIUM Casey)

H. californicus (LeConte) BC
pacificus Casey
uancout'eri Casey
H. cinctus (Say) BC AB oN PQ NS
latebricola Nordmann
H. cribratus (LeConte) -oNPQ
H. pallipes (Gravenhorst) -oNPQ
grauenhorsti I\trby
H. sellatus (]-eConte) oN PQ

Genus OCHI|HEPHILUM Stephens

(Subgenus OCHTHEPHILUM s. str.)
(C ry p to b ium Mannerheim)
O. fracticorne (Paykull)+ BC ON PQ NS
brachelytratum Korge
furcaticcrne Grimmer

116
 Staphylinidae

Subtribe Astenina
Genus ASTENUS Dejean
A. americanus (Casey)
-oN
A. binotatus (Say)
A. brevipennis (Austin)
-oN
MB ON
A. cinctus (Say) _BCAB PQ
A. discopunctatus (Say)
centralis Ars.stin
AB -oNPQ NB
A. linearis (Erichson)
A. longiusculus (Mannerheim) BC
(?') t risig natu s Boheman

Subfamily STAPFIYLININAE
Tribe Philonthini
Genus NEOBISNIUS Ganglbauer
Key to New World species: Frank (1981)
N. jucundus (Horn t - BC AB ON
N. lathrobioides (Baudi )* PQ
Frank
N. occidentoides AB - MB
N. senilis tHornt - BC
N. sobrinus (Erichsont ON PQ NB I\TF
arduus (Sharp)
ocreatus (Hotn)
simulator Smetana
N. terminalis ( LeConte) PQ NS
adustus (Sharp)
delicatulus (Sharp)
N. viliosulus t Stephenslt ON PQ
fuluicornis (Notman)

Genus ERICHSONIUS Fauvel

Actobius Fatvel
Key to North American species: Frank (192b)
E. alumnus Frank _ONPQ
E. civicus Frank
E. fraterculus (Horn)
-oN
E. inutilis tHorn.l
-oNPQ
E. nanus (Horn) AK
-oNPQ
_ONPQ
E. patella {Horn
- NT BC NB NS NF
E. pusio tHorn )
r

Ilrr
-oNPQ
*oNPQ
E. rosellus Frank
-oNPQ
Genus PHILONTHUS Curtis
P. aequalis Horn
P. agilis{Gravenhorsilt - BC
nlger Melsheimer
P. albionicus Mannerheim AK BC
P. asper Horn NF
P. aurulentus Horn AK BC
P. blandus ( Gravenhorst) ON PQ-NS
paederinus Sachse
pulchellus Melsheimer
P. brunneus (Gravenhorst)
breuis Melsheimer
oN PQ

dimidiatus (Say)
picatus l\ttby
P. bucephalus Horn BC
P. carbonarius (Gravenhorst)t AK BC Of,l PQ NB NS NF
uorlzs (GyllenhaI)
P. caurinus Horn BC
P. cephalicus Casey _AB - l,ff -
P. cephalotes (Gravenhorst)t - yuoN
P. chlorinus Tottenham
uiridanus Hotn
BC
BC _ PQ NS
NS
NF

P. cognatus Stephensf BC ON PQ NS NF
fu sci p enni s (Mannerheim)

r17
Staphylinidar,r

P. concinnus t Gravenhorst)f BC AB SK MB ON PQ NS PE NF
oc hro pus ( Gravenhorst)
P. confertoides Hatch BC
P. confertus f,eOonte
P. couleensis Hatch NS__Nr
P. crestonensrs Hatch BC
P. crotchi Horn BC
P. cruentatus (Gmelin)t BC AB _MB ON PQ -NF
P. cunctans F.orn NS NF
P. cyanipennis (Fabricius)* BC AB _ IYID ON PQ NB NS
coe r ule i7:, ennis (Mannerheim)
P. debilis (Gravenhorst)t BC ON PQ NF
P. dimidiatus C.R. Sahlbergf AB
P. distans Horn BC
P. duplicatus Bernhauer & Schubert AK YK BC AB
basalis LIotn
P. ferreipenn:s Horn _BC_
P. fi metarius (Gravenhorst)t NF
P. finitimus Cjasey PQ
P.
P.
flavibasis Oasey
forcipatus .Flatch
---ec- _BC
P. fraternus Fall -YK pe
P. furvus Nordmann NT BC ON NB NS -LB NT
ca li forn i : us Mannerheim
comptus Haldeman
P. fusiformis Melsheimer
P. gaudens Tlrttenham
P. grandicollir Horn
P. hepaticus ltrrichson PQ
cinctulu:: Melsheimer
(?) nanu;; Melsheimer
uaricolor Boheman
P.
P.
hudsonicurr Horn
hyperboreus alaskensis Smetana
AK
AK_NT v
P. instabilis }Iorn _YK ecABi*Ms--
_ABSKMBONPQ
P. lacustrinuir Scheerpeltz NT
lacustris Casey
P. Iecontei Hr.rrn BC
P. leechensis Hatch AK BC ie NF
P. lindrothi Smetana -NS NF
P. lomatus Erichson BC_ONPQ _NS NF
georgianus Sachse
P. Iongicornis Stephenst NF
P. micans (Glavenhorst)t
P. neonatus Simetana -NS IU
P. nigritoides Smetana AK
P. occidentalis Horn
P. opacipennis Notman
P. palliatus (Gravenhorst)
-:"y -NS
_NS
NF

P. perforatus Fauvel
P. perversus Horn -sc- fu
P. pettiti Horn _BC
P. picicornis lJorn _BC
P. politus (Lirrn6)t AK_ _BCAB SK MB oN PQ we Ns NI'
horrisi Melsheimer
mandibularis Kirby
P. protervus Casey -MB
P. puberulus Horn BC
P. pubes Hor:r BC
_MB
P.
P.
pumilio Ct;.sey
quadricollis Horn BC AB _Fe
_PQ
P. quediinus Horn -AB
P. rainieri Hatch BC
P. rectangulus Sharpt AK_ BC SK MB ON PQ NF
tetr ag onoce p halus Notman
P. sanguinolentus (Gravenhorst)t BCi
P. septentrionis Fall _YK
P. sericinus [Iorn
P. siegwaldii Mannerheim AK BC
-"e
ON PQ _LBNF
P. sordidus (Gravenhorst)t BC AB SK MB ON PQ NB NS \TET

long i pen nis Provancher

pachycephalus Nordman

118
 Staphylinidae

P. spiniformis Hatch _BC

P. stacesmithi Hatch _BC
P. stictus Hausen PQ
P. subvirescens C.G. Thomson* AI( YK NT BC AB PQ
P. theveneti Horn AK BC
P. triangulum Horn _BC
P. umbratilis (Gravenhorst)t -BC NB NS
P. umbrinus (Gravenhorst) NS
P. varians (Paykull)f NT BC ON PQ NB NF
P. vulgatus Casey AK NS

Genus GABRONTHUS Tottenham

G. thermarum (Aub6)t oN PQ

Genus GABRIUS Stephens

G. bidentatus (Horn) BC
G. brevipennis (Horn) AK_N' BC AB SK MB ON PQ NB NS T\TF

G. decipiens (Horn) BC
G. downiei (Hatch) BC
G. linearis (Casey) BC
G. malkini (Hatch) AK__ NF
G. microphthalmus (Horn) BC
G. nigritulus (Gravenhorst)t BC AB - lle oN pe
pumilus Mannerheim
G. oceanus (Hatch) BC
G. ottawensis (Casey) ON
G.
G.
picipennis (Maklin)
seattlensis (Hatch)
AK YK NT BC
BC
AB SK MB ON PQ NE NS -inNr NF
G. subnigritulus (Reitter)+ BC _PQ NF
G. virilis (Horn) BC
G. wawawai (Hatch) BC _-__
Genus CAFIUS Stephens
(Subgenus EUREMUS Bierig)
C. bistriatus Erichson NB NF
bilineatus Erichson
rufifrons Bierig
C. luteipennis Horn BC
(Subgenus BRYONOMUS Casey)
C. canescens Miiklin AK _BC
C. seminitens Horn _BC
(Subgenus REMUS Holme)
C. femoralis Meklin AK BC
mutatus Gemminger & Harold
(Subgenus PSEUDOREMUS Koch)
C. nudusjohnsoni Fall BC

Genus BELONUCHUS Nordmann

B. rufrpennis (Fabricius) ON PQ
for mos us ( Gravenhorst)
pallipes Melsheimer

Genus LAETULONTHUS Moore & Legrer

L. Iaetulus (Say) oN PQ

Genus HESPERUS Fauvel

H. apicialis (Say) ON PQ
haernaturus (Erichson)

119
Staphylinide:e

Tribe Staphylinini
Genus CREOPHILUS Samouelle
Saprophilus Streubel
C. maxillosuii (Linn6)t AK YK NT BC AB SK MB ON PQ NB NS PE LB NF
arcticus Erichson
uillosus (Gravenhorst)

Genus ONTIIOLESTES Ganglbauer

O. cingulatus (Gravenhorst) NT BC AB SK MB ON PQ NB NS _LBNF
O. murinus (Linn6)t NF

Genus HADIIOTES Meklin

H. crassus (I{annerheim) AK BC
extensu! LeConte

Genus THINOPINUS LeConte

Tri c hon n t h us Motschulsky
r

T. variegatus (Motschulsky) BC

Genus PLATYDRACUS C.G. Thomson

P.
P.
cinnamopterus (Gravenhorst)
comes (Letlonte)
-oNPQ ?
P. exulans (Lrichson) -oN
P. fossator ((|ravenhorst) MB ON PQ NB
immaculatus (Horn)
P. maculosui (Gravenhorst) ON PQ
ery thr otrennis (Mannerheim)
uiduatu r (Fabricius)
P. mysticus ( 0richson)
_BC -ONPQ-____
P. rutilicaudrr (Horn)
P. tarsalis (N lannerheim) AK _BC
sub meta: llicus (Mannerheim)
P. tomentosus (Gravenhorst) ON
P. violaceus ( Gravenhorst) ON PQ NS
cupriper',nis (Melsheimer)
P. viridanus Horn) ON PQ NS
P. vulpinus ( \ordmann) ON PQ

Genus STAP.FIYLINUS Linn6

S. aeneocephllus DeGeerf
S. ater Gravenhorstt
BC
BC Io*pe NS-- NT
S. badipes LcConte ix MB ON PQ NB NS
carbona',us LeConte
S. capitatus 131and
^B _ONPQ
BC AB NB NF
S. erythropterus Linn6f MB ON PQ NS
S. melanarius Heert _ONPQ
S. nigrellus Ilorn
S. pleuralis L,eConte
S. saphyrinus LeConte
S. winkleri Bernhauert
BC
BC
BC
BC
v-
Tlibe Xanthopygini
Genus TYMF'ANOPHORUS Nordmann
Diaphoetes Waterhouse
T. puncticollis (Erichson) BC AB oN PQ NB NF
borealis (Hatch)

Tribe Quediini
Key to North American species: Smetana (1971)
Genus BEERIA Hatch
B. nematocera (Casey) BC

t20
 Staphylinidae

Genus HETEROTHOPS Stephens

H. asperatus Smetana _BC
H. carbonatus Fall _BC
H. conformis Smetana _BC
H. fraternus Smetana _BC
H. fusculus LeConte _BC AB SK MB ON PQ _NS
californicus LeConte
fumigatus LeConte
H. marmotae Smetana
H. minor Smetana N,ecAB-tu3NPe-ns--wr
_BC_ONPQ
H. pusio LeConte
H. sordidus Smetana AK_ NT BC

Genus QUEDIUS Stephens

(Subgenus MEGAQUEDIUS Casey)

Q. explanatus LeConte BC
Q. manitobensis (Casey) AB -ll"-
(Subgenus MICROSAURUS Stephens)
(Anastictodera Casey)
(Quediochrus Casey)
Q. campbelli Smetana SK oN PQ
Q. canadensis (Casey) oN PQ Ne-
Q. caseyi caseyi Scheerpeltz --Nt- AB SK MB oN PQ
curtipennis (Casey)
Q. caseyi divergens (Casey)
_BC AB
paruipennis Bernhauer
uteanus (Casey)
Q. charactus Hatch -BC
_BC
Q. crescenti Hatch
Q. criddlei (Casey) AK YK NT BC AB SK MB
hoodiHatch
Q. erythrogaster Mannerheim
_BC AB SK ON PQ
neotomae Hatc}'
Q. fulgidus (Fabricius)t
_BC _MB
iracundus (Say)
Q. griffinae Hatch BC
Q. incisus Hatch BC AB
Q. Iimbifer Horn BC
_NS
Q. mesomelinus (Marsham)t
Q. mordax Smetana
Q. pellax Smetana
A"-
_YK
BC
BC
BC
AB
AB -y" oN PQ LB NF

Q. peregrinus (Gravenhorst) ON PQ -NS

siluicola Casey
Q. pullmani Hatch
Q. rainieri Hatch
Q. spelaeus spelaeus Horn
v_ _
BC
BC
BC
AB

AB
SK
SK
MB
_NS
quadriceps Casey
Q. tahomae Hatch BC
Q. terminatus Melsheimer
Q. velox Smetana AK BC AB
(Subgenus QUEDIUS s. str.)

Q. curtipennis Bernhauert _BC

parallelus Hatch
Q. horni Hatch _BC
Q. labradorensis insularis Smetana AK. pe---ie
Q. labradorensis labradorensis AK NT BC AB SK MB ON NF
Smetana
Q. lanei Hatch _BC
Q. laticollis (Gravenhorst) ON
marginatus Korge
neomolochinus Korge
Q. molochinus (Gravenhorst)t
_PQ NF
nigribasis Kotge
Q. molochinoides molochinoides AKYK_BC oN PQ NF
Smetana
(Subgenus QUEDIONUCHUS Sharp)

Q. plagiatus Mannerheimx AI{ YK NT BC AB MB ON PQ LB NF

r2l
Staphyiinida:

laeuigati ts (Gyllenhal)
I ongipen n is Mannerheim

ruftpennis Maklin
(Subgenus Dl STICHALIUS Casey)

Q. capucinus (Gravenhorst) ON PQ
ater (Zie1let)
bordzs Melsheimer
inuersus (.Say)
seeuersi Korge
Q. frater Smetana _BCAB
Q. montivagus Smetana _BC
Q. nevadensis (Casey) _YK _BCAB
Q. pediculus (Nordmann) AK _BCAB
Q. tetricus Smetana _BC
Q. transparerLs Motschulsky AK _BCAB
marginalis Mdklin
pugetensis Hatc}l
(Subgenus RI\PHIRUS Stephens)
(.Quediel'.us Casey)

Q. adnexus Snetana _NF

Q. aenescens Maklin AK BC AB
agnatus iCasey)
Q. breviceps (Casey) AK BC
Q. brunnipen nis Mannerheim AKYKNTBCABONPQ_LBNF
Q. densiventris (Casey) AKYK-BCAB-PQNF
alutaceu; Hatch
bonapari:ei }{atc}i'
Q. elevatus Hatch _BC
Q. felimani (Zetterstedt) AK YK NT BC AB SK MB PQ LB NF
hyperboreus Erichson
Q. ftigidus Srretana AK-NT_AB ON NF
Q. fulvicollis r Stephens)* AKYK_BCAB _MBONPQ_LB NF
pugetant.rs (Casey)
raphiroides Hatch
Q. impar Smctana BC AB
Q. nanulus (C asey) _BCAB
helenae ( Casey)
humilis rtCasey)
Q. oculeus (Crrsey) AK BC
sparsus (Casey)
Q. prostans Horn BC
rupimonlis (Casey)
Q.
Q.
Q.
rusticus Snetana
seriatus H,rrn
simulator iJmetana
_BCAB_MB_PQ
_BC
AKYKNT_ABSKMBONPQ
v_ - in-
Q. sublimbatr.rs Maklinx AKYKNTBCABSKMB_PQ _LBNF
orbiceps ICasey)
solitarius (Casey)
Q. vilis SmetzLna BC
(Subgenus P/.RAQUEDIUS Casey)

Q. puncticeps Horn BC

Genus HEMIiIUEDIUS Horn

H. ferox (LeC,rnte) ON PQ NS

Genus ANAQIIEDIUS Casey

A. vernix (LeConte) ON PQ NB NS

Genus ACYL()PHORUS Nordmann

(Subgenus AI{ACYLOPHORUS Smetana)
A. pratensis l,eConte ON PQ NF
(Subgenus ACYLOPHORUS s. str.)
A. alternans Smetana -oN _NS
A. caseyi Leni; -oNPQ
A. flavicollis Sachse -oN
A. pronus Eri,chson AK NT BC MB ON PQ NB NS NF

r22
 Staphylinidae

Subfamily XANTHOLININAE
Key to North American species: Smetana (1982)
Tribe Diochini
Genus DIOCHUS Erichson
D. schaumi Kraatz _ONPQ
breuipennis Casey
pallidiceps Casey
thoracicus Casey

T?ibe Othiini
Genus ATRECUS Jacquelin du Val
A. americanus (Casey) - ON PQ - NS
A. macrocephalus (Nordmann) AK - NT BC AB ON PQ NB NS NF
A. quadripennis (Caseyt - BC AB
Genus PAROTHIUS Casey
P. californicus (Mannerheim.t - BC
P. punctatus Smetana - BC

Tribe Xantholinini
Genus GAUROPTERUS C.G. Thomson
G. fulgidus {Fabricius)* - ON

Genus NIIDOBIUS C.G. Thomson

N. cephalus (Say) AK - NT BC AB SK MB ON PQ NB NS NF
assimilis (Kirby)
consentane us (Nordmann)
flauipennis (Nordmann)
giluipennis (Casey)
hiesenwetteri (Sachse)
nubipennis Casey
p alliat us (Melsheimer)
N. pugetanus Casey - BC
corticallis Casey
debilis Casey
limbalis Casey

Genus LEPTACINUS Erichson

Leptacinodes Casey
Xa nt ho p hius Motschul sky
L. batychrus tGyllenhal)+ - BC MB ON PQ
seriatus LeConte
L. intermedius Donisthorpet - BC AB SK MB ON PQ NF
L. pusillus (Stephens)f - ON PQ
nigritulus LeConte

Genus PHACOPHAILUS Coiffait

P. parumpunctatus tGyllenhal)* - ON PQ
cephalicus (LeConte)

Genus STENISTODERUS Jacquelin du Val

S. rubripennis tleConte) - ON

Genus LITHOCIIARODES Sharo

Nematolinus Casey
Oligolinus Casey
L. longicollis (LeContet - ON PQ

Genus LINOHESPERUS Smetana

(Casey)
L. borealis - BC
Smetana
L. cuspifer - BC AB
L. subhamaLus Smetana AB

t23
Scaphidiidae

Genus STIC'IOLINUS Casey

S. flavipes (leConte) oN PQ NS
scolopaceus Casey

Genus HESF;EROLINUS Casey

Leiolinus Casey
H. parcus (LeConte)
tarsalis (Casey)
uteonus (Casey)

Genus GYRCTIIYPNUS Samouelle

G. angustatus Stephenst _BC_ONPQ NB NS NF
G. campbelli Smetana MB ON PQ
G. fracticorn:.s (O.F. Mulier)t _BCAB_MBONPQ NB NS NF'
obsidiar us (Melsheimer)

Genus NEOI|YPNUS Coiffait & Saiz

N. beckeri Srretana ON PQ
N. emmesus,'Gravenhorst) oN PQ
luteiuen; ris (Casey)
sang uinolent us (Melsheimer)
N. fragilis (C,rsey) _BC ON
N. hamatus ( Say) NT BC AB SK IYI D ON PQ NS
macilenius (Casey)
N. lineatus (lVendeler) ON
linearis Blatchley)
N. melanops .Casey) oN PQ
N. monstrosr- s Smetana ON PQ
N. obscurus (Orichson)
oregonul: (Casey)
---ecABix MB oN PQ N" NS NF
protractils (Casey)
N. picipennis (LeConte) _BC
Genus OXYBLEPTES Smetana
O. kiteieyi Slretana ON PQ NB

Genus XANTHOLINUS Dejean

X. linearis (Olivier)t R11 NS

Genus HYPNOGYRA Casey

H. gularis (Lr,rConte) ON
uernicaht Casey

Genus XEST()LINUS Casey

X. abdominalis Casey SK MB ON PQ
frontalis Hatc}:
indianeri sis Blatchley
ouiceps Casey

Family SCAPHIDIIDAE shining fungus beetles

J.M. Campbell

Scaphidiidae is a small family of fungus-feeding consensus among researchers on their proper place-
beetles that comprises over 1200 species around the ment. The family Scaphidiidae is poorly known in
world, but; only about 50 species occur in North North America, and species recognition of most
America: cif these. 17 are recorded in Canada and Canadian taxa is not possible. Casey (1893, 1900)
Alaska. IVIost species are found in fungi, but some has completed the most recent revision of the North
live in rottrng wood, leaf litter, and under bark. American shining fungus beetles.
Althorrgh many recent authors (Lawrence
1982a; Lawrence and Newton 1982) have placed the aK (1); YK (1); NT (1); BC (4); AB (1); SK (2); MB
scaphidiids in the Staphylinidae family, there is no (3); oN (13);PQ (10);NF (1)

t24
 Pselaphidae

Tribe Scaphidiini
Genus SCAPHIUM Kirby
S. castanipes Kirby AK YK NT BC AB MB ON PQ

Genus SCAPHIDIUM Olivrer

S. obliteratum LeConte -oN
S. piceum Melsheimer -oN
_ONPQ
S. quadriguttatum Say
q uadr inota tu m Castelnau
quadripustulatum Say

T?ibe Scaphisomini
Genus SCAPHISOMA Leach
S. castaneum Motschulsky
S. convexum Say
S. lacustris Casey
| -:'- - yuBN pe_
S. repandum Casey oN
S. rubens Casey _i PQ

Genus TOXIDIUM LeOonte

T. compressum Zimmermann
T. gammaroides LeConte
ON
ON pe-
Genus BAEOCERA Erichson
Eubaeocera Cornell
Key to Norbh American species: Cornell (1967)
B. apicalis LeConte -oN PQ
distincta (Blatchley)
B. congenera Casey BC SK MB ON PQ
speculifera Casey
B. deflexa Casey D'\
rq
-
B. lalsata Achard SK_ONPQ NF
concolor auct.
youngi (Corr]ell)
B. humeralis FaIl
B. nana Casey __BC___ oN Fe
rubriuentris Casey

Family PSELAPHIDAE short-winged mold beetles

A. Dauies

Eighty-frve species of Pselaphidae are recorded J.L. LeConte, E. Brendel, and T.L. Casey described
in Canada and Alaska: however. the members of most North American species in the mid to late
this family have been poorly collected because of nineteenth century; their work was subsequently
their minute size and the obscurity of their habi- brought together by Bowman (1934). The present
tats. They are found chiefly in moist habitats, arrangement of the supraspecific taxa is based on
including the edges of bogs and marshes, under the recent world catalog of genera by Newton and
bark of dead trees, and especially in leaf litter and Chandler (1989). Chandler (1990o) also published a
rotten wood; a few are myrmecophilous. Adults of key to the North American genera.
most Pselaphidae are predatory on small inverte-
brates such as nematodes, mites, and collembolans;
the larvae are seldom seen, and their habits are
equally unknown. AK (10); YK (1); NT (3); BC (23); AB (16); SK (9);
Jeannel (1955) divided Pselaphidae into six sub- MB (16); ON (56); PQ (38);NB (15);NS (11); pE (2);
families, all of which are represented in Canada. LB (1);NF (3) ' l(2)

125
Pselaphidae

Subfamilv FARONINAE
Tribe Faronini
Genus MEGARAFONUS Casey
Key to Ncrrth American species: Schuster and Marsh (1958)
(Subgenus IIEGARAI'ONUS s. str.)
M. ventralis Casey AK BC

Genus SONTfMA Casey

Rafonus Casey
Key to North American species: Marsh and Schuster (1962)
S. margemina Park & Wagner AK BC
S. parviceps (Maklin) AK BC

Subfamilv EUPLECTINAE
Trib; Tlogastrini
Genus CON,CPLECTUS Brendel
Hexirherius Grigarick & Schuster
Prorhet:ius Raffray
C. canaliculttus (LeConte) PQ
syluatit.us (Raffray)

Genus ORO ?US Casey

Key to Nc'rth American species: Schuster & Grigarick (1960)
O. striatus Oasey BC

Genus TETIIASCAPIIA Schuster & Marsh

T. dentata Park & Wagner BC

Genus FOVIIOSCAPHA Park & Wagner

F. terracola Park & Wagner BC

Tlibe Euplectini
Genus THE|IIUM Casey
Apothinus Shatp
T. cavifrons (LeConte) ON

Genus EUP],ECTUS Leach

Key to North American species: Wagner (1975)
E. acomanus Casey PQ
E. confluens LeConte PQ
E. duryi Casey PQ NB NS
barri P;trk & Wagner
E. elongatus Brendel ON
E. karsteni (Reichenbach)t
iowensis Casey
--s^- ON PQ PE

E. longicollis Casey ON
E. signatus tReichenbach)t SK MB ON pa:)

E. silvicolus Chandler NS

Genus PYCI{OPLECTUS Casey

Key to Ncrrth American species: Wagner (1975)
P. congener (Casey) ON
tenellus Casey
P. interruptrrs (LeConte) ON
hudsonicus (Casey)
P. linearis (LeOonte) ON
irnpressiceps (Casey)

Genus RAMECIA Casey

Liniolis Grigarick & Schuster
R. crinitus (tsrendel) PQ NS

126
 Pselaphidae

Genus TRIMIOPLECTUS Brendel

Key to North American species: Chandler (1990b)
T. obsoletus Brendel - ON PQ

Genus BIBLOPLECTUS Reitter

Key to North American species: Chandler (19906)
B. integer (LeConte) MB ON PQ NB NS
B. ruficeps tleContet - ON

Genus ACTIUM Casey

Proplectus Raffray
Key to North American species: Grigarick and Schuster ( 1971)
A. barri Park & Wagner - BC AB
A. retractum Casey AK BC
hatchiPatk & Wagner
A. wawonaensis Grigarick & Schuster - BC

Genus ACTIASTES Casey

Key to North American species: Grigarick and Schuster (1971)
A. foveicollis (LeConte.l - BC - ON PQ
costalis (Brendel)
pennsy lu anicus (Bowman)
A. globiferum (LeConte) PQ
blandurn (Casey)

Genus ACTIZONA Chandler

A. borealis Chandler - BC AB

Genus CUPILA Casey

Key to North American species: Grigarick and Schuster (1968)
(Subgenus CUPILA s. str.)
C. clavicornis (Maklinl AK BC

Genus MELBA Casey

Key to eastern North American species: Chandler ( 1990b)
(Subgenus MELBA s. str.)
M. parvr:la tl,eConte) - ON

Subfamily BATRISINAE
Tlibe Batrisini
Genus BATRISODES Reitter
Key to western North American species: Grigarick and Schuster (1962)
Key to eastern North American species: Park (1947)
(Subgenus BABNORMODES Park)
B. riparius (Sayt - ON PQ
(Subgenus EMPINODES Park)
B. albionicus (Aub6) AK BC
B. denticauda (Caseyt - BC
(Subgenus EXCAVODES Park)
B. frontalis tl,eConte) AB - MB ON PQ
B. globosus (LeConte) AB ON PQ NB NS
B. scabriceps (LeConte) - ON PQ
harringtoni (Casey)
B. sinuatifrons (Brendelt - ON
(Subgenus PUBIMODES Park)
B. striatus tl,eConte) - ON PQ
aterrimus (Casey)
cephalotes (Casey)
psotai Park
sirnplex (LeConte)

I27
Pselaphidae

Subfamily GONIACERINAE
Tribe Bythinini
Genus MAC HAXRODES Brendel
M. carinatus (Brendel) ON

Genus TYCI{OBYTHINUS Ganslbauer

Bythinopsis Raffray
T. bythinioides (Brendel) ON PQ NS

Tlibe Brachyglutini
Genus RYB,I\XIS Saulcy
Bryaxis Leach
R. clavata tlirendel) ON PQ NB
brendeii (LeConte)
R. conjunctr tLeContet ON PQ NB NS
R. mystica ( asey AB -MBONPQ NB NS
R. transversa Fall AK - NT -AB SK MB ON PQ NB NF
R. varicornir, t Brendel) ON PQ NB

Genus BRAI.)HYGLUTA Thomson

Nisa Ci,Lsey
B. arguta (Casey) ON
B. terebrata (Casey, SK ON PQ
intricatt (Casey)
perfora o (Brendel)

Genus REIC HENBACHIA Leach

Key to western North American species: Grigarick and Schuster (1967)
R. albionica Morschulskyr AK
americt:..na Raffray
compar (LeConte)
francistana Casey
fundata Casey
R. arcilera C asey _PQ
R. binodiferi,L Casey AK BC AB
R. borealis Casey AB - MB ONPQNB_PE
R. canadens.s (Brendel) ON
R. congener Brendelr _PQ
R. corporalis Casey oN PQ
R. cyiindrana tBrendel r ON
R. distans C,rsey ON
R. facilis Ca rey ON
R. fusLicorni s Casey BC AB
R. kapones (lrigarick & Schuster - BC AB
R. peregrinaLor Casey MB oN----
R. propinqu:. rleconte) MB ON PQ NB NS
R. rubicundr. (Aubet ON
R. scabra (B'endelt ON
R. spatulifer Casey - \aK - BC AB SK MB ON PQ NB LB NF
Genus NISAXIS Casey
N. tomentos..r tAub6, ON
cincinnLtta Casey
minuta (Brendel)

Genus EUTIiICHITES LeOonte

E. zonatus (llrendel t ON PQ
zimmer,nanni LeOonte

Genus DECTTiRTHRON Brendel

Key to eastern North American species: Park (1958)
(Subgenus DECARTHRON s. str.)
D. abnorme (LeConte)
D. formiceti il,eConte)
D. inusitaturn Park
_ yl"T:"yu ON PQ
ON
_PQ

I28
 Pselaphidae

D. laurenticum Casey ON
D. Iongulurn (LeConte) oN PQ
D. scarifrcatum Brendel
D. stigrnosum Brendel UN

Tribe Tychini
Genus LUCIFOTYCHUS Park & Wagner
Key to western North American species: Park and Wagner (1962)
Key to eastern North American species: Park (1956)
L. cogrratus (LeConte)
L. testaceus (Casey) T---BC--nneoNpeNs--- NF

Subfamily PSELAPHINAE
Tribe Pselaphini
Genus PSELAPHUS Herbst
(Subgenus PSELAPHUS s. str.)
P. bellax Casey BC AB SK MB ON PQ NB NS
P. erichsoni LeConte
_AB -oN
MB ON PQ NB NS
P. fustifer Casey

Tribe Tyrini
Genus TYRUS Aub6
Pytna Casey
T. corticinus (Casey) BC AB
T. humeralis tAub6t NT nfn OW Pe Ne--
compdr LeOonte

Genus CEDIUS LeConte

Key to North American species: Park (1949)
(Subgenus SINISTROCEDIUS Park)
C. ziegleri LeConte ON

Tlibe Ceophyllini
Genus CEOPHYLLUS LeConte
C. monilis LeConte MB ON

Tlibe Tmesiphorini
Genus TMESIPHORUS LeConte
T. costalis LeConte ON

Tribe Ctenistini
Genus PILOPIUS Casey
P. piceus (LeConte) ON PQ
lacustris Casey
saginatus Casey

Subfamily CLAVIGERINAE
Tribe Adranini
Genus ADRANES LeConte
A. lecontei Brendel _ABSK MB ON
A. taylori Wickharn BC_SK

t29
Hydrophilicae

Superfamily Hydrophiloidea
FamilyIIYDROPHILIDAE waterscavengerbeetles
R.E. Roughley

Hydr'tphilidae forms a large family of mostly species, both aquatic and terrestrial, are scav-
aquatic t'eetles; 149 species are known within the engers.
Canadiar' and Alaskan borders. The family can be Smetana (1988) has recently revised 1;he taxon-
divided into two biologically different groups: one omy of the Canadian species, and his contribution
aquatic lnd the other terrestrial. The aquatic was used as the basis for this checklist.
species or:cur in a wide range of habitats, but most
prefer shallow standing water. The terrestrial AJ{Q0; YK (20); NT (36); BC (81);AB (65); SK (52);
species (subfamily Sphaeridiinae) live in most MB (6a); oN (e2); PQ (86);NB (38);NS (33); PE (2);
kinds of lecaying organic matter. Adults of most LB (5);NF (24) ' H (11);I (18)

Subfamily IfYDROC HINAE

Genus HYjI'ROCHUS Leach
H. brevitars is Knisch
H. currani IV.J. Brown _BCABSX -oN
MB ON
PQ
PQ
H. granulatns Blatchley _BC MB ON PQ
ouellet, Leech
H. neosquarnifer Hellman
H. pseudoscuamifer D.C. Miller BC_
-oN
MB ON
PQ
PQ
H. scabratus Mulsant :" PQ
gibbost.ts Melsheimer
H. squamife r LeConte BC AB SK MB ON PQ NB
laticollis Notman
H. subcupreus Randall PQ

Subfamily HELOPHORINAE
Key to North American species: Smetana ( 19854)
Genus HEL OPHORUS Fabricius
(Subgenus ]IELOPHORUS s. str.)
H. g'randis I lliger* ON PQ NB NS
(Subgenus ITHOPALELOPHORUS Kuwert)
H. angustic,rllis d'Orchymont SK MB ON PQ NB NS NF
H. auricollisi Eschscholtzx AK YK BC PQ NF
angust tIus Mannerheim
consimilis Mannerheim
inquinttus Mannerheim
H. browni Angusx YK NT
H. columbia.nus McCorkle _NTBCABSK MB
H. electus d'Orch).rnont _NT_ABSK MB
corri d'Orchymont
H. frosti Srr.etana PQ NB
H. furius Srnetana NT BC MB AB SK ON PQ
frater ,t\rr:.etana
[I. inflectus McCorkle BC MB ON PQ
[I. lacustris LeConte AK YK NT BC AB SK MB ON PQ NU lls _LBNF
lL ledatus rl'Orchymont BC .AB
H. leechi M,:Corkle BC AB SK MB
H. linearis ..-eConte _ABON
oblitus Shatp
pallent Casey
H. linearoioes d'Orchlnnont
H. lineatus Say _ _ _ :cy:"yuoN pe NB
o b solet e s ulcat us Motschulsky

uentratis Motschulsky
H. marginicollis Smetana
H. nitiduloicles d'Orchymont AK YK NT BC AB s^
-oN
MB ON PQ NB NF

130
 Hydrophilidae

H. nitidulus LeConte BC AB SK MB
gregolis d'Orchymont
H. oblongus LeConte* AK YK NT BC AB SK MB ON PQ
H. orchymonti Smetana
_NTBCAB SK
-oN
MB ON NB
H. orientalis Motschulskyx PQ NS
H. parasplendidus parasplendidus YK NT MB
Angus
H. sempervarians Angus AK YK NT BC AB SK MB ON PQ NB NS LB NF
H. splendidus J. Sahlberg* YK NT
(Subgenus GEPHELOPHORUS Sharp)
H. sibiricus (Motschulsky)x AK YK NT
(Subgenus CYPHELOPHORUS Kuwert)
H. tuberculatus Gyllenhal* AK YK BC AB SK MB ON PQ NS
scaber LeConte
(Subgenus ORPHELOPHORUS d'Orchyrnont)
,,
H. arcticus W.J. Brown* NT MB ON

Subfamily BEROSINAE
Genus BEROSUS Leach
Enoplurus Hope

B. fraternus LeConte _BC AB SK MB ON PQ

cal ifor nicus Motschulsky
B. hatchi D.C. Miller NT BC AB SK MB ON PQ
B. maculosus Mannerheim AK
te sselatu s Motschulsky
B. oregonensis D.C. Miller AB SK
B. peregrinus (Herbst) ON PQ
auritus Melsheimer
imma c uI at u s Zimmermann
subsignatus LeConte
B. striatus (Say) NT BC AB SK MB ON PQ
B. stylifer Horn _AB -oN
falcatus Shatp

Subfamily CHAETARTHRIINAE
Genus CHA.ETARTHRIA Stephens
Key to northwestern species: Miller (1974)
C. atra (LeContet _PQ_NS
C. nigrella (LeConte) - BC
,,2
C. pallida tl-eContet - SK
nigriceps (LeConte)

Subfamily IIYDROBIINAE
Tlibe Hydrobiini
Genus LACCOBIUS Erichson
Key to Canadian species: Gentili ( 1985)
L. agilis (RandaII)
L. borealis Cheary
L. carri d'Orchymont
NT
AK YK NT
BC
BC
AB_MB
AB
AB SK
ON
ON
ON
PQ
v_ NF

pacificus D.C. Miller

L. cinereus columbianus D.C. Miller AK YK NT BC AB SK MB -PQ
L. minutoides d'Orchymont ON PQ NB
L. reflexipenis Cheary NT MB ON PQ NB NS
reflexipenis Malcolm
L. spangleri Cheary SK MB ON PQ NB \TIT
spangleri Malcolm
L. teneralis Cheary
L. truncatipenis D.C. Miller NT AB SK
-oN
MB ON
Genus IfYDROBIUS Leach

131
Hydrophili<Lae

NT BC AB SK MB ON PQ NB NS _ LB
H. fuscipes llinn6)*
H. melaenus (Germar)
globostts (Say)
AK
5 -oNPQNBNS
NF

Genus SPERCHOPSIS LeConte

S. tessellata (Zie gler) ON PQ NS

Genus AMFITOR Semenov

A. latus (Hcrn) _BC
A. scabrosus (Horn) .4K BC AB

Genus PAR.{CYMUS C.G. Thomson

P. confusus Wooldridge _BC
P. despectur; (LeConte) AB SK MB ON PQ
P. subcupreus (Say) _YKNTBCAB SK MB ON PQ NB
P. tarsalis t).C. Miller _BC
Genus CRENITIS Bedel
Crenip.lilus Ilorn
C. alticola (.lall) BC
C. digesta (n,eConte) _AB sK y" 9* PQ NB NS
C. maculifrc ns W.J. Brown YK_BCAB
C. malkini l).C. Miller BC AB
C. monticol,L (Horn) -PQ NS
C. morata (l{orn) AK YK NT BC AB -PQ _NF
C. paradign.a ( d'Orchymont) BC AB
paracy noides (d'Orchymont)
C. rufiventris (Horn) BC
C. snoqualnLie D.C. Miller BC

Genus ANACAENA C.G. Thomson

Crenitu.lus Winters
A. limbata ( Fabricius)t BC AB SK MB ON PQ NS NF

Tribe Helocharini
Genus ENOCHRUS C.G. Thomson
Key to Ncrth American species: Gundersen (1978)
(Subgenus I'INOCHRUS s. str.)
E. piceus pir:eus D.C. Miller BC
carinalus (Horn)
(Subgenus IIETHYDRUS Rey)
E. blatchleyr tFallr MB
E. californicus (Horn) _BC
E. cinctus tSay) _SK MB ON PQ NB
limbalis (Melsheimer)
E. consors {l,econte) PQ
E. consortus Green MB ON
E. ochraceus (Melsheimer) Nr ec AB sr MB ON PQ NB NS NF
lacustris (LeConte)
simplet: (LeConte)
E. perplexurr t LeContet MB ON PQ NS
fimb r iatu s (Melsheimer)
E. pygrnaeus nebulosus (Say) MB ON PQ
E. sayi Gunrlersen -oN
(Subgenus L,UMETUS Zaitsev)
E. diffusus (LeConte) NT BC AB SK MB ON
E. hamiltoni (Horn) NT BC AB MB ON PQ NS NF
collinus W.J. Brown
conjunr:tus (Fall)
horniLeech
liuidus (Walker)
pacificus Leech
pyretus Leech

r32
 Hydrophilidae

E. reflexipennis (Zimmermann) PQ NB NS
angustulus (Casey)

Genus CYMBIODYTA Bedel

Key to species: Smetana (1974)
C. acuminata Fall - YK NT BC AB SK MB ON PQ - NS
C. blanchardi Horn - ON PQ
C. chamberlaini Smetana - ON
C. columbianaLeech AK BC
C. minima Notman - BC MB ON PQ
C. pacifica Leech - BC
hatchi D.C. Millet
C. semistriata (Zimmermann) - ON PQ NB
C. toddi Spangler MB ON
C. vindicata Fall NT BC AB SK MB ON PQ NB NS - LB NF
Genus HELOCOMBUS Horn
H. bifidus {LeContet - ON PQ NB

Subfamilv IIYDROPHILINAE
Genus TROPISTERNUS Solier
T. columbianus W.J. Brown - BC AB SK - ON
T. ellipticus (Leconte) - BC
ofizis Motschulsky
concolor Sharp
T. glaber lHerbst) - ON PQ - NS
T. lateralis humeralis Motschulsky - BC
T. lateralis marginatus Motschulsky - BC AB
binotatus Walker
limbalis (LeConte)
T. lateralis nimbatus (Say) AB - MB ON Pe - NS
T. mixtus (Leconte) MB ON pQ - NS
T. natalor d'Orchyrnont - ON PQ
Genus HYDROPHILUS O.F. Miiller
H. triangularis Say - BC AB SK - ON pe
subsulcatus LeConte

Genus DIBOLOCELUS Bedel

D. ovatus (Gemminger & Haro1d) - ON pQ
oualis (Ziegler)

Genus HYDROCHARA Berthold

Key to species: Smetana (1980)
H. brevipalpis Smetana - ON
H. obtusata {Say) NT BC AB SK MB ON pQ
H. simula Hilsenhoff& Tracy - ON

Subfamily SPHAERIDIINAE
Tlibe Sphaeridiini
Key to North American species: Smetana (1978)
Genus PHAENONOTUM Sharp
P. exstriatum (Say)
dubium S}ratp
-oN
sem ig I obos um (Zimmermann)

Genus SPHAERIDIUM Fabricius

S. bipustulatum Fabriciust _BCABSK_ONPQNBNS
S. lunatum Fabriciust _BCABSKMBONPQNBNSNF
S. scarabaeoides (Linn6)t _BCABSKMBONPQNBNSNF
crendtum Palisot de Beauvois

133
Hydrophilidae

Tribe Cercyonini
Genus CERCYON Leach
(Subgenus PROSTERCYON Smetana)
C. roseni Knlsch NT BC AB SK MB ON PQ NB NS
opacellus Fall
(Subgenus CERCYON s. str.)
C. adumbrat us Mannerheim AX BC
unicolor' (Motschulsky)
C. assecla Slnetana
C. atricapilh.Ls (Marsham)t
AB
AB
-oN PQ
PQ
NB NS
NS
mundut Melsheimer
C. cinctus Slretana NT BC AB SK MB ON PQ
C.
C.
connivens Fall
erraticus i]metana _BC -oN
MB ON PQ
C. fimbriatus Mannerheim AK _BC
C. haemorrhoidalis (Fabricius)t -BC -oN PQ NB NS PE NF
basillaris Notman
nigricol,t.is (Say)
C. herceus frigidus Smetana AK YK NT BC AB SK MB ON PQ
C. herceus h':rceus Smetana BC AB SK MB ON PQ
C. impressus (Sturm)t BC
C. indistinct,rs Horn _PQNB
C. lateralis ( Vlarsham)-i NT BC AB ON PQ NB NS NT
incrematus Notman
C. limbatus . vlannerheim AK YK NT BC AB
ful uiper n i s Mannerhei m
C.
C.
C.
litoralis (( iyllenhal)x
luniger Ma.nnerheim
marinus (j.G. Thomson*
AK BC
AK 1T( NT BC AB SK MB ON
PQ

PQ
NB NS
__I"
C. matthewsi Smetana AK_AB
C. mendax Smetana -oN
C. nevadanus Knisch
C. praetextat;us (Say) MB ON PQ NS
testaceus Blatchley
C. pygmaeus (Illiger)+ BC AB SK MB ON PQ NB NS NF
apicalis (Say)
C. quisquilius (Linn6)t BC AB SK MB ON PQ NB NS NF
C. terminatus { Marsham;f
nozzs llelsheimer
-oN PQ

C. tolfrno Habch AK BC
C. unipunctu tus (Linn6)l BC AB SK MB ON PQ NB NS
(Subgenus DICYRTOCERCYON Ganglbauer)
C. ustulatus ( Preyssler)* PQ
(Subgenus P.ARACERCYON Seidlitz)
C. analis tPaykullt* NT BC AB SK MB ON PQ NB NS NF
falliWi:fterc
maculalus Melsheimer
C. minusculus Melsheimer BC SK MB ON PQ NB NF

Genus TECT OSTERNUM Balfour-Browne

Genyon Smetana
T. naviculare (Zimmermann) - ON PQ

Genus DELI'OSTETHUS Sharo

Ictinosternum Smetana
,,
D. columbier.sis ( Hatcht

Tribe Megasternini
Genus MEGASTERNUM Mulsant
M. obscurum (Marsham)t _BC
M. posticaturn (Mannerheim) AK BC
pugetense Halch

134
 Histeridae

Genus CRYPTOPLEURUM Mulsant

C. minutum (Fabricius)t _BCAB MB ON PQ NB NS PE -NF
C. subtile Sharpf _BCAB SK MB ON PQ _NF

Family GEORYSSIDAE minute mud-loving beetles

R.E. Roughley

Georyssidae includes minute beetles, usually mud attached to the body; they are apparently algal
under 2 mm long; two species are known for the feeders.
Nearctic region, one of which occurs in Canada.
Adults are found in mud or sand at the edges of
streams and are often concealed by sand grains or BC (1);AB (1);PQ (rl

Genus GEORYSSUS Latreille

G. ousillus LeConte BC AB PQ

Family SPHAERITIDAE false clown beetles

R.E. Roughley

Sphaeritidae contains only one genus, organic material (e.g. fungi) and on trees when the
Sphaerites, with two species in the Holarctic sap runs.
region; one of them occurs in Canada and Alaska.
Adults are found in dung and other decaying eK (1); BC (1);AB (1)

Genus SPHAERITES Duftschmid

S. politus Mannerheim AK BC

Family HISTERIDAE hister beetles

A. Dauies

In Canada and Alaska 118 species of Histeridae treatments of the North American Histeridae. The
have been recorded. The adults and larvae are chiefly classification used here is based on Arnett (1968),
predators of other insects and their larvae. Most whereas the status of the species is based on Mazur
species are associated with decaying organic material, (1984) and on work by Helava (personal communica-
dung, and carrion; some are found in sandy areas, in tion).
mammal burrows, under bark, and in bark-beetle
galleries; a few are myrrnecophilous. Although some
species are associated with stored products, they most
likely feed on other insects that occur there. AK (1); YK (2); NT (2); BC (57); AB (42); SK (25);
LeConte (1845), Marseul (1853-1857, 1860-1862), MB (43); oN (65);PQ (58);NB (13);NS (12);PE (3);
and Horn (1873) have conducted major taxonomic NF(7) . H(1);I(10)
Subfamily ABRAEINAE
T?ibe Abraeini
Genus ABRAEUS Leach
A. bolteri LeConte BC

135
Histeridae

Tribe Plegaderini
Genus PLEGADERUS Erichson
P. comonfor.i Marseul - BC
uergran.dis Casey
P. consors Horn - BC
egenus Casey
P. fraternus Horn - BC
P. molestus Casey - BC
obesus Casey
P. nitidus Horn - BC
P. sayi Mars,eul - BC - ON PQ NB NS
P. setulosus Ross BC
P. transversus (Say) ON PQ NF

Tribe Acritini
Genus ACR.ITUS LeConte
A. exiguus I lrichson I - ON PQ
acicula ius (J.E. LeConte)
cribriptnnis Marseul
elliptictrc Casey
natchez Marseul
obliquu s (J.E. LeConte)
subdep,'essus (Blatchley)
A. nigricorn s lHoffman r-l' - ON PQ

Genus AELIITES Horn

A. politus tleContet - ON PQ
A. simplex t,l.E. LeConter - ON PQ
acupict ts (Marseul)

Tlibe Teretriini
Genus TERIITRIUS Erichson
T. american rs J.E. LeConte
dissonans Marseul
-oNPQ
picipes J.E. LeConte
T. montanus Horn _BC

Subfamily SAPRININAE
Genus SAPIIINUS Erichson
S. discoidalir LeConte - BC
amplus Casey
latiforn,.is Casey
mayhewi Papp
S. impressusr J.E. LeConte - BC AB ON
infaustus LeConte
piceus ., .8. LeConte
S. lugens Errchson NT BC AB SK MB ON PQ
co I iforn icus Mannerheim
consimilis Waiker
disting uendoides Hatch
oregonensoides Hatch
sus Dahl gren
p se udoclete r
sejunctoides }Jatch
S. oregonensis J.E. LeConte - BC AB SK MB ON PQ
distinguendus Marseul
lecontecides Hatch
p se u do ing u e ndu s Hatch
cl i s t
p se udo s ej unc toide s Hatch
pseudosejunctzs Hatch
sejunctus Marseul
spurcus LeConte
S. pensylvar-"icus tPaykull) MB ON PQ
profusoi.des Hatch

136
 Histeridae

S. sedakovii Motschulsky* YK

Genus EUSPILOTUS Lewis

(Subgenus HESPEROSAPRINUS Wenzel)
E. assimilis (Paykull) MB ON PQ NB
extraneus (Illiger)
sardous (Flach)
semisulcus (Hatch)
simulatus (Blatchley)
E. conformis (J.E. LeOonte) -oN
ouiformis (Blatchley)
E. cribrum (Casey) BC AB SK MB
carri (Hatch)
E. insertus (LeConte) BC AB SK
obductus (LeOonte)
opacellus (Casey)
uinctus (LeConte)
(Subgenus NEOSAPRINUS Bickhardt)
E. scrupularis (J.E. LeConte) BC

Genus XEROSAPRINUS Wenzel

X. fimbriatus (LeOonte) NT BC AB SK
desertorum (Marseul)
X. lubricus (LeConte) _BC AB SK MB
rotundiftons (Marseul)

Genus GEOMYSAPRINUS Ross

(Subgenus PRISCOSAPRINUS Wenzel)
G.
G.
castanipennis (Fall)
cheyennensis (Casey) _BCAB
AB SK yu_
G. copei (Horn) _BCAB
G. lanei (Hatch & McGrath) _BC
G. laramiensis (Casey) -BCAB
G.
G.
obscurus (LeOonte)
obsidianus (Casey)
moniliatus (Casey)
_ _ T_ -oN pr])

(Subgenus GEOMYSAPRINUS s. str.)

G. rugosifrons (FaIl) MB

Genus HYPOCACCUS C.G. Thomson

H. aeneipunctatus (Horn) -BC
H. bigemmeus (LeOonte)
H. bigener (J.E. LeConte)
eriensis (Hatch)
- - - :'T 3* MBor.pe NS:
illinoensis Wolcott
impunctellus ( Casey)
lakensis Blatchley
michiganensis (Hatch)
ohioensis (Hatch)
ontarioensis (Hatch)
sphaeroides (J.E. LeConte)
H. estriatus (LeConte) BC AB
H. frtchi (Marseul) PQ
H. fraternus (Say) BC AB SK MB ON PQ NS PE NF
H. iris (Fall) AB SK MB ON PQ
alutiger Wenzel
H. mancus (Say) MB
H. omissus (Casey) AB SK MB
H. patruelis (J.E. LeConte) AB_MBONPQ
jaueti (Marceul)
H. seminitens (LeConte) MB ON PQ

Genus BAECKMANNIOLUS Reichardt

B. dimidiatipennis (J.E. LeConte) PQ NB NS NF
deserticola (Marseul)
desertorum (Marseul)
palmatus (Say)

L37
Histeridae

Genus NEOI'ACHYLOPUS Reichardt

N. sulcilrons tMannerheim.l - BC

Genus GNA'IIHONCUS Jacquelin du Val

G. rossi Hat< h - BC
G. rotundatus (Kugelann)f - BC AB SK MB ON PQ NF
communis (Marseul)
deletus (J.E. LeConte)
nanus (Scriba)

Subfamilv DENDROPHILINAE
Tri6e Dendrophilini
Genus DENI )ROPHILUS Leach
D. punctatus (Herbsu+ - BC MB ON PQ
punctulatus (Say)
D. xavieri M rrseult - BC - ON
sexstria;lus Hatc}a

Tlibe Bacaniini
Genus BACFNIUS LeOonte
B. punctifon:ris (LeConte) _ONPQ

Tribe Paromalini
Genus CARC INOPS Marseul
Carcinus Marseul
C. pumilio (Etrichson)t - BC AB SK MB ON PQ NF
nanus Ql.E. LeConte)
q uatuordecimsfrjolzs (Stephens)

Genus XESTIPYGE Marseul

X. conjunctu n tSayt - ON
X. geminaturn tJ.E. LeConte) - ON

Genus PLATYLOMAIUS Cooman

P. aequalis (iiayt - BC AB - MB ON PQ
affinis (,1.8. LeConte)
estriatu.t (J.E. LeConte)

Genus PARC,MALUS Erichson

P. bistriatus Erichson AB ON PQ NB - NF
P. mancus C,lsey - BC AB
P. seeversi (\Venzel; - ON
P. teres LeConte - ON PQ NB - NF

Subfamilv TRIBALINAE
Genus EPIEIRUS Erichson
E. pulicarius Erichson - ON PQ
minor J,8. LeOonte
E. regularis iPalisot de Beauvois) - ON
nigrellus (Say)
uicinus LeConte

Genus ONTIIOPHILUS Leach

O. deflectus .l{elava - ON

Genus CAEF|OSTERNUS LeConte

C. americanus U.E. LeConte) - ON

138
 Histeridae

Subfamily HISTERINAE
Tribe Histerini
Genus ATHOLUS C.G. Thomson
Peranus Lewis
A. americanus {Paykull ) AB SK MB ON PQ
diffractus (Casey)
exard.tus (J.E. LeConte)
perplexus (LeConte)
A. bimaculatus ( Linn6)t BC AB -oN PQ
obliquus (.Say)
A. falli tBickhardtt BC AB SK MB ON PQ
gagates Fatl
A. sedecimstriatus { Sayl MB ON PQ

Genus MARGARINOTUS Marseul

(Subgenus MARGARINOTUS s. str.)
M. guttifer Horn _AB
(Subgenus PTOMISTER Houlbert & Monnot)
M. brunneus (Fabricius;f ON PQ NB NS PE
cadau e r inu s (Ho{frnann)
impressus (Fabricius)
obtusatus (Harris)
uirginiae (Casey)
M. egregius {Casey) -MBONPQ
M. foedatus (J.E. LeConte) NB NS
M. fractifrons {Caseyt -:nc- _yuoNPQ
fidelis (Casey)
planifrons (Lewis)
M. harrisii (Kirbyt BC AB SK MB ON PQ
repletus (J.E. LeConte)
M. hudsonicus (Caseyt MB ON PQ NB
M. immunis (Erichson) BC MB ON PQ ols---
M. intermptus (Palisot de Beauvois) -oNPQ NB
M. merdarius (Hoffmann)t nc _ONPQ NS
memnonius (Say)
M. pluto (Casey) BC AB MB
albertensis (Hatch)
carri (}{atc}r)
M. rectus (Caseyt BC
M. umbrosus (Casey) BC
umbratilis (Casey)
(Subgenus STENISTER Reichardt)
M. obscurus (Kugelann)f BC
semisculptus (LeConte)
stercorarius (Hoffmann)
(Subgenus PARAIISTER Bickhardt)
M. cognatus (J.E. LeConte) MB ON PQ
unicus (Casey)
M. lecontei Wenzel SK MB ON PQ NB NS PE
M. purpurascens (Herbst)t --nc-
(Subgenus PROMETHISTER Kryzhanovsky)
M. confusus Wenzel
M. marginicollis (J.E. LeConte)
-oN
MB ON
PQ
PQ

Genus HISTER Linn6

H. abbreviatus Fabricius AK BC AB SK MB ON PQ NB
bifidus Say
coloradensis Casey
crenicollis Flach
subhernisphaericus Palisot de Beauvois
H. albertanus Casey AB
H. curtatus J.E. LeConte PQ NS
haldemanni Marseul

139
Histeridae

H. defectus .f.E. LeConte

permixlus Zimmermann
SK -oN PQ

H. depuratol Say BC AB SK MB ON PQ NB NS
paykulliKirby
perbreu.'.s Casey
spretus J.E. LeConte

Genus SPILODISCUS Lewis

S. arcuatus (Say) NS
S. militaris (Horn) -BC
electus (Casey)
oregonus (Casey)
simpliupes (.Fa\l)
S. sellatus (LeOonte) BC
senneui lei (Marseul)
S. semiruber (Casey) BC AB SK MB
soloris t Carnochan)

Genus PSILr)SCELIS Marseul

P. abnormal s Mann SK MB
P. carri Hatch _AB
P. corrosa Casey _AB
P. planipes (LeConte) MB ON PQ_NS
harrisi:J.8. LeConte)
P. subopaca LeConte) YK BC AB MB

Tribe Platysomatini
Genus PLA1YSOMA Leach
(Subgenus PLATYSOMA s. str.)
P. aequum (l,eConte) ON PQ
P. aurelianu n (Horn) PQ
P. lecontei Nlarseul BC AB _MBON PQ
d.epress tm J.E. LeConte
tabellurt Casey
(Subgenus EURYLISTER Bickhardt)
P. carolinum (Paykull) oN PQ
cinnamomeum (A. White)
sordidum (Say)
(Subgenus CYLISTER Cooman)
P. coarctaturn J.E. LeConte
P. punctigemm (LeConte)
-BCAB
_BCAB _ yuoNPQ
Genus CYLISTIX Marseu.
C. cylindrica (Palisot de Beauvois)
C. deficiens Casey
C. gracilis (J.E. LeConte) _ _ rsxy"giliH _ _ _ _ _

Tribe Hololeptini
Genus HOLOLEPTA Paykull
H. aequalis Say BC AB MB ON PQ
fossularis Say

Tribe Exosternini
Genus PHEL,ISTER Marseul
P. subrotunius (Say) oN PQ
P. vernus (Say) ON
saunieri Marseul

I40
 Clambidae

Subfamily HETAERIINAE
Tribe Hetaeriini
Genus HETAERIUS Erichson
H. brunneipennis (Randall)
H. exiguus Mannerheim BC
-oN
nitidus Martin
H. tristriatus Horn BC AB SK MB

Superfamily Eucinetoidea
Family EUCINETIDAE plate-thigh beetles
J.M. Campbell

Eucinetidae is a small family that contains Horn (1880) last reviewed the species of this
about 30 species throughout the world; seven of family, as part of Dascillidae; they are poorly known
them, placed in one genus, are known to occur in taxonomically.
Canada. These beetles are associated with fungal
decay and are found in leaf litter or under bark of vK (1); NT (2); BC (5); eB (3); SK (3); MB (4); ON
fungus-infested logs. (a);PQ (a);NB (2);NS (2) . I (1)

Genus EUCINETUS Germar

E.
E.
E.
haemorrhoidalis (Gerrnar)t
infumatus LeConte
morio LeConte
_v BC
BC
AB SK MB oN
ON
PQ

PQ
NB NS
_NS
E. oviformis LeConte MB
E. punctulatus LeConte YK_ SC AB sx MB ON PQ
E. terminalis LeConte BC
E. testaceus LeConte _NT BC AB SK MB oN PQ NB

Family CLAMBIDAE minute beetles

J.M. Campbell

Clambidae forms a family of small beetles that decaying vegetation, where they probably feed on
are rarely collected because of their size and habi- spores of fungi and slime molds.
tat. The family contains five genera and about 50 The American species were revised by Endrridy-
species in the world. Only two genera comprising Younga (1981) and are well-known taxonomically.
seven species are known to occur in Canada and
Alaska, but more are likely to be found. Adults are
usually collected during their evening flight. They AK (1); BC (3); AB (3); oN (1); PQ (6) o I1[
can also be collected by sifting leaflitter and piles of (1); I (1)

Genus CALYPTOMERUS Redtenbacher

C. oblongulus Mannerheim AK BC

Genus CLAMBUS Fischer von Waldheim

Sternuchus LeConte
C. armadillo armadillo (DeGeer)t PQ
niger Hatch
C. gibbulus (LeContetf, BC AB PQ
r adul a Endr }dy-Younga
C. howdeni Endrddy-Younga ON PQ

r47
Scirtidae

C. pubescens Redtenbacher* BC AB PQ
puberu.lus LeConte
C. smetanai Endrcidy-Younga PQ
C. vulneratus LeConte PQ

Family SCIRTIDAE marsh beetles

J.M. Campbell

The lamily Scirtidae contains about 600 species sion. Horn (1880) last reviewed the species of this
throughout the world, but most of them are found in family, as part of Dascillidae. Klausnitzer (1976),
temperai;e zones. Six genera and 26 species are however, reviewed several species occurring in
known tc occur in Canada and Alaska. Adults are Canada; he (1980) also segregated some species for-
collected, often in large numbers, on vegetation in merly assigned to Elodes to form the new genus
wet habitats, particularly marshes, bogs, and Flauohelodes.
swamps; they are attracted to light. Larvae are
aquatic and feed on fungi, algae, diatoms, and other
organic nratter.
Most Canadian species are well known except AK (1); YK (1); NT (2); BC (10); AB (a); SK (3); MB
those of the genus Cyphon, which is in need of revi- (3); oN (15);PQ (15);NB (7);NS (3);NF (5)

Genus ELCTDES Latreille

Hemic;tphon LeConte
Sacodt:s LeConte
E. angusta Hatch _BC
E. apicalis 1,eConte _BC
E. emargin;r.ta Hatch _BC
E. maculicc'llis Horn PQ_NS NF

Genus FLAVOHELODES Klausnitzer

F. fuscipennis (Gu6rin-M6neville) ON
F. pulchella (Gu6rin-M6neville) ON
F. thoracica. (Melsheimer) ON

Genus MICROCARA C.G. Thomson

M. explanat;a LeConte BC SK MB ON PQ NB

Genus CYPHON Paykull

C. brevicoll:.s (LeConte) BC
C. collaris (rGu6rin-M6neville) NB
C.
C.
concinnus (LeOonte)
confusus W.J. Brown
BC AB- - - :a
_ONPQ NF
horiont Klausnitzer
C. exiguus :1orn BC
C. modestu'; (LeConte)
nebulosus (LeConte)
--pe NF'
C. -oNPQ
_ONPQ
C. neovaria bilis Klausnitzer NS-
_ONPQ
C.
C.
obscurus (Gu6rin-M6neville)
ochreatus Klausnitzer yu oN PQ fu Ns NF

C. punctatr:.s (LeConte) BC
C. pusillus,.LeConte) _NT BC AB SK
C. rufrcollis (Say) _ONPQ
C. variabilis (Thunberg) AK YK NT BC AB SK MB ON PQ lle Ns NF

Genus PRIONOCYPHON Redtenbacher

P. discoideus (Say) oN PQ
P. limbatus LeConte oN PQ NB

Genus SCIIITES Illiger

S. orbiculatus (Fabricius) oN PQ
S. tibialis Gu6rin-M6neville AB ON PQ NB

t42
 Lucanidae

Superfamily Dascilloidea
Family DASCILLIDAE soft-bodied plant beetles
J.M. Campbell

The status of Dascillidae has changed consider- occur throughout the world. The adults are usually
ably since Horn reviewed it (1880). Its size has found on foliage; they feed on plants or are carnivo-
been greatly reduced by the r_emoval of species now rous. The larvae live in soil or are aquatic; they feed
included in the families Scirtidae, Eucinetidae, on vegetative matter such as roots or leaf mold.
Artematopidae, and Ptilodactylidae, so that only
one species is now known to occur in Canada.
Fifteen genera and about 80 species are known to BC (1)

Subfamily DASCILLINAE
T?ibe Cneoglossini
Genus ARAIOPIDIUS Cockerell
Araeopus LeConte
A. monachus (LeConte) BC

Family RHIPICERIDAE cedar beetles

J.M. Campbell

Rhipiceridae contains five genera and about 50 The adults are often found in mating congregations
species throughout the world. Only one genus with on the trunks of trees. with many males grouped
one species is known to occur in southern Ontario. around one female.
The larvae of this species live on the roots of trees,
where they are parasitic on the nymphs of cicadas. oN (1)

Genus SANDAI-US Knoch

S. niger Knoch ON
hnochi (Gt6in-M6neville)
rubidus Melsheimer
r uftpennis (Latreille)
scabricollis Haldeman

Superfamily Scarabaeoidea
Family LUCANIDAE stag beetles
J. McNq.mara

Lucanidae includes 14 species in Canada. The Howden and Lawrence (1974) published a key
larvae of all species breed in or beneath the decay- of the North American genera, and since the genera
ing wood of logs or stumps and therefore are of little occurring in Canada include few species, specific
economic importance. The adults are believed to identifrcation is relativelv easv.
feed on honeydew or sap. They lay their eggs in
bark crevices, especially near the roots. The adults
are attracted to light and are most commonly found vK (1); NT (2); gC (7); AB (2); SK (2); MB (3); ON
in wooded areas. (8);PQ (5); NB (2);NS (3);NF (1)

r43
Lucanidae

Subfamilv SYNDESINAE
Tribe Sinodendrini
Genus SINODENDRON Hellwig
S. rugosum Mannerheim - BC

Tlibe Ceruchini
Genus CERUCHUS Macleay
C. piceus (V/eber) MBONPQ-NS
bolbi (t.lastelnau)
uirginiznsis Casey
C. punctatus LeConte BC
C. striatus LeConte _BC

Subfamily NICAGINAE
T?ibe Nicagini
Genus NIC,\GUS LeConte
N. obscurus tleConter ON PQ

Subfamily LUCANINAE
Tribe Lucanini
Genus LUC ANUS Scopoli
(Subgenus LUCANUS s. str.)
L. elaphus I'abricius - ON
carleng:i Angell
(Subgenus I'SEUDOLUCANUS Hope & Westwood)
L. capreolus tl.inndt - ON
domo l:'abricius
muticu.; Thunberg
nigricephalus (Benesh)
trigonu:.s Thunberg
L. placidus iay - ON
lentus \)astelnat

Tribe Dorcini
Genus DORCUS Macleay
D. parallelu* {Sayt - ON PQ
carnocl,,ani Angell
costatu,s LeConte
nanus Casey
oblongus (Charpentier)
uoeti (Schonherr)

TYibe Platycerini
Genus PLA',YCEROPSIS Benesh
P. keeni (Casey) BC
pedicelt aris (Moilenk)

Genus PLAIIYCERUS Geoffroy

Key to species: Benesh (1946)
P. marginalis Casey NT BC AB SK MB NB NS
P. oregonensis Westwood - BC
chalybo:eus Casey
coerulescens LeConte
P. piceus (K,rby) - \'K NT BC AB SK MB ON PQ NB - NF
depressus LeConte

144
 Scarabaeidae

virescens (Fabricius) oN PQ NS
angustus casey
iowanus Casey
peregrinus Casey
quercus (Weber)
securidens (.Say)

Family PASSALIDAE peg and b,ess beetles

J. McNamara

Only one species of Passalidae occurs in Reyes-Castillo (1970) published key to the
Canada. These beetles, which live in rotten logs American genera of this family.
and stumps, are not economically important. The
adults and larvae communicate with each other by
stridulation. The adults prepare food for the larvae
by chewing the wood and mixing it with saliva. MB (1);ON (1)

Tribe Proculini
Genus ODONTOTAINIUS Kuwert
Passalotaenius Kuwert
O. disjunctus (Illiger) NtB ON
cornutus (Fabricius)

Family SCARABAEIDAE scarErb beetles

J. McNamara

Scarabaeidae is one of the largest families of Mating taker; place during the summer. The eggs
beetles in the world; 248 species are known to occur are laid in the soil in late July and early August;
in Canada and Alaska. Beetles of this family are they hatch immediately, and the larvae begin to
popular with collectors because of their bright colors feed. When bhe weather turns cold the larvae bur-
and the large horns of some groups. The size of row deeper into the soil, below the frost line, where
these beetles varies greatly, from 1.9 mm for they hibernate. In the spring when the weather is
Pleurophorus longulus to 15 cm for Goliathus, warmer, they move up near the surface of the
which is probably the largest insect known based on ground and continue to feed until they pupate in
mass. The scarabs contain some of the most June. The adults emerge about the first of July,
destructive beetles, such as the Japanese beetle depending on the area. Many variations occur in
(Popillia japonica), the Asiatic garden beetle the life cycle of scarabs. The Phyllophoga species
(Maladera castanea), and the European chafer cycle ranges firom 2 to 4 years, depending largely on
(Amphirnallon majalis). These belong to a group of the latitude.
scarabs that feed primarily on Iive plant matter as John Cooper is currently revising the scarabs of
adults, and usually on plant roots or rotting wood as Canada and Alaska, and his work is expected to be
larvae. The other group of scarabs are mainly dung published as part of The Insects and Arachnids of
feeders or scavengers. They feed primarily on dung Canada, a serries published by Agriculture Canada.
or decaying plant and animal matter as adults, and The classifrcation used here is taken almost entirely
in many cases the larvae feed on the same material from his manuscript.
as it is supplied to them by the adults. Because of
their economic importance, the life cycle of many AK (13); YK (9); NT (17); BC (88); AB (96); SK (58);
species is fairly well known. That of the Japanese MB (85); ON (136); PQ (109); NB (a1); NS (a7); PE
beetle is probably typical of the family in general. (11);LB (7);ltIF (20) . H (1); I (21)

145
Scarabaeidae

Subfamilv SCARABAEINAE
Tribl Scarabaeini
Genus CAIITHON Hoffmansegg
Key to North American species: Robinson (1948)
(Subgenus CANTHON s. str.)
C. chalcites (Haldemanr ON
C. pilularir:s {Linn6) - AB
hudsonias (Forster)
laeuis i.Dtury)
obtusi'fens (Ziegler)
uiridetcens Horn
uoluen; (Fabricius)
C. vigilans LeConte - ON
(Subgenus BOREOCANTHON Halflter)
C. praticok LeConte BC A-B SK MB
C. simplex Leconte - BC AB
(Subgenus ILAPHYROCANTHON Martinez)
C. viridis 1l)alisot de Beauvois) - ON
metallicus Stutrn
obsoletus (.Say'1
uiridicatus (Say)

Tribe Coprini
Genus COF'RIS O.F. Miiller
Key to New World species: Matthews (1961)
C. lricator lricator rFabricius) - ON PQ
anagllpticus Say
tullius (O\ivter)
C. minutus (Drury) PQ
ammo:l, Fabricius
lar Fa.:ricius
reflexu:.s Panzer
silenzs Fabricius

Tlibe Onthophagini
Genus ONTHOPHAGUS Latreille
Key to Nrrth American species: Howden and Cartwright (1963)
O. hecate hecate (Panzer) - BC AB SK MB ON PQ NB NS
furcicollis Dejean
hastator (Fabricius)
lamal)ejean
latebrosus (Fabricius)
obtectt:s (Palisot de Beauvois)
soyi C;rstelnau
scabricollis l{rtby
O. nuchicornis (Linn6)t - BC AB - MB ON PQ NB NS NF
r hinoc'zro s Mel sheimer
xiphias LeConte
O. orpheus canadensis lFabriciusr - ON PQ
O. orpheus pseudorpheus MB
Hou'den & Cartwright
O. pennsyl' anicus Harold - ON PQ

Subfamily APHODIINAE
Tribe Aegialiini
Key to North American species: Gordon and Cartwright (1988)
Genus MICRAIGIAJ-IA W.J. Brown
M. pusilla (Horn) - BC AB - MB

I46
 Scarabaeidae

Genus AXGIALIA Latreille

(Subgenus LEPTAIGIALIA W.J. Brown)
A. humeralis W.J. Brown
_BCAB -oN PQ
A. montana W.J. Brown
A. rufescens Horn _BCABSK MB ON PQ NB
(Subgenus PSAMMOPORUS Thomson)
A. criddlei W.J. Brown AK BC AB SK MB ON PQ NF
A. cylindrica (Eschscholtz) AK BC
A. lacustris LeConte AK YK NT BC AB SK MB ON PQ NB NS
A. nana W.J. Brown
_BCAB
PQIU___ )u
A. opaca W.J. Brown
A. terminalis W.J. Brown AK YK NT BC AB SK MB LB NF
(Subgenus RHYSOTHORAX Bedel)
(Anomalaeg ial ia W J . Brown)
.

A. spissipes LeContef oN PQ NS NF
rzlo (Fabricius)
rufina Sllfvergerg
(Subgenus AXGIAJ-IA s. str.)
A. arenaria (Fabricius)t NS
A. blanchardi Horn _BC PQ
A. carri Gordon & Cartwrieht AB
NtrD r\I\T
A.
A.
conferta Horn
crassa LeConte _ _ _ ecf:" PQ

insularis W.J. Brown

A. opifex Horn oN PQ NS PE

Tribe Aphodiini
Genus APHODIUS illiger
Key to eastern North American species: Gordon (1983)
A. aenictus Cooper & Gor'lon ON
A. albertanus W.J. Brown NT BC AB
A. aleutus Eschscholtz _BCAB
ursinus Motschulsky
A. alternatus Horn BC SK MB
inermis Robinson
moclipsiHaLch
A. anthracinus LeConte _
A. aquilonarius W.J. Brown BCTSKIo*pe-----
^AB

A. badipes Melsheimer
oblongus Say
A. bicolor Say
A. bidentatus Schmidt
bidens LeConte
BC AB __9* PQ

A. borealis Gyllenhalx BC AB _MBON PQ NB NS LB NF

errons W.J. Brown
A. brevicollis LeConte
A. browni Hinton
smithiW.J.Brcwn
-BC
AB
_ y"_
A. bryanti W.J. Brown AB SK
A. carri W.J. Brown AB SK
A. coloradensis Horn AB
A. congregatus Mannerherm AK ec AB
arcticus Harold
A. consentaneus LeConte NT AB SK MB
A. cormptor W.J. Brown AK \'K BC AB SK MB
A. cribratulus Schmidt BC
cribratus LeConte
A.
A.
criddlei W.J. Brown
decipiens Horn ___sc AB
AB :" yu_
A. denticulatus Haldeman AB
A. distinctus (O.F. Miiller)t _BC AB SK MB ON pe NS
inquinatus (Herbst)
m a eu I i pe n n i s Mel sheimer
A. duplex LeConte BC AB SK MB
A. edmontonus W.J. Brown AB

I47
Scarabaeic ae

A. erraticus (Linn6)t MB ON PQ NB NS NF
pensuullensis Melsheimer
A. fimetarius (Linn6)-f BC AB SK MB ON PQ NB NS LB NF
nodifrons Randall
A. fossor (Linn6)t AK BC AB SK MB ON PQ NB NS PE LB NF
A. granarirLs (Linn6)t _BCAB _MB ON PQ NB NS
aterr i;' n u s Melsheimer
inutilt:s Horn
metalticus Haldemarr
spretu.s Haldeman
A. guttatus Eschscholtz AKYK_BCAB _MB PQ _NS
A. haemorrhoidalis (Linn6)i _BCAB SK MB ON PQ NB NS _LB NF
A. hamatus hamatus Say _BCAB SK MB PQ
occidentalis Horn
sagittarius Harold
A. hirsutus W.J. Brown BC
A. hyperboreus LeConte NT BC AB SK MB ON PQ NB NS NF
anguloris LeConte
A. iowensis Wickham AB SK MB
A. lentus Eorn ON PQ
A. leopardr.Ls Horn AK YK NT BC AB SK MB ON PQ NN NS
A. leptotarr;is W.J. Brown AB TU
A. manitobensis W.J. Brown AB _MB ON PQ _NS
A. omissus omissus LeConte
A. omissus torpidus Horn
A. opacus l,eConte
YK
YK f' BC
BC
BC
AB
AB
AB
SK MB
SK MB
ON

colum,tiensis Wickham
A. pardaiis LeConte BC
A. pectoral:s LeConte BC AB
A. peculios rs Schmidt _MB
awemeanus W.J. Brown
longitursis Fall
tenuihr,rsis Reitter
A. phaeopterus LeConte BC AB
canadensis Garnett
(?) rus;elli Hatch
A. piceatus Robinson AK BC AB
rainer', Hatch
A. pinguell rs W.J. Brown BC AB SK MB
A. pinguis1:Ialdeman NT ? AB SK MB oN pe NF
ualidus Hotn
A. prodromus (Brahm)t _BC
_BC
SK MB oN PQ NB NS
A. richards'tnae Hatch
A. rogersi Ilatch
A. rubeolus (Palisot de Beauvois) _:r_ ON
c op r on.y tn u s Mel sheimer

A. rubripennis Horn oN PQ NB NS
A. rufrpes (.Linn6)t ON PQ
A. ruricola Melsheimer AB -MB oN PQ _NS NF
aureliunus Harold
curtus Haldeman
A. scabriceps LeConte -AB SK
A. scrofa (Fabricius)t -PQ NB NF
A. sigmoideus Van Dyke BC
A. socialis \V.J. Brown _AB _MB
A. stercoros,us Melsheimer oN pe
A. subaeneits LeConte BC?
gentne.i }{atch
perpoli,tus Gordon
A. subterraneus (Linn6)t ON PQ NB NS
A. talpoides,i W.J. Brown
A. tenellus Say AK I'K NT BC AB
-ms
_MB ON PQ
A. thomomJrsi W.J. Brown _MB
A. vittatus Say _BCAB SK MB oN PQ NB NS _LB
A. walshii }Iorn AB SK MB
Genus OXYOMUS Stephens
O. silvestris (Scopoli)'l BC PQ
opacifrons }Iorn
porcatus (Fabricius)

I48
 Scarabaeidae

Tlibe Eupariini
Genus ATAENIUS Harold
Key to North American species: Cartwright (1974)
A. abditus {Haldeman.t ON PQ NB
attenuator Harold
A. gracilis ( Melsheimer t ON PQ
A. imbricatus (Melsheimer) ON
sordidus Harold
A. spretulus (Haldemant ON
cozsors Fall
falliHinton
A. strigatus (Sayt oN PQ

Genus DIAJ-YTES Harold

D. criddlei W.J. Brown MB
D. striatulus tSayt MB ON PQ NB NS
cribrosus (LeConte)
D. truncatus (Melsheimer) ON PQ
coruinus (Haldeman)
D. ulkei Horn ON PQ

Genus DIAI-YTELLUS W.J. Brown

Key to species: Brown (1929)
D. dialytoides tFall r
D. humeralis (Leconte)
-oNPQ
-oN
Tribe Psammodiini
Genus PSAMMODIUS Fallen
Key to New World species: Cartwright (1955)
P. caelatus (LeContet BC
P. laevipennis Costa+
aegialioides Haldeman
-oN
P. oregonensis Cartwright BC
P. sulcicollis (Illigertt * Dr'l
rq
Genus RHYSSEMUS Mulsant
Key to New World species: Gordon and Cartwright (1980)
R. californicus Horn AB
R. germanus tLinnet*
puncticollis W.J. Brown
-oN pa:)

Brown
R. neglectus W.J. AB
R. sonatus LeConte AB SK MB

Genus PLEUROPHORUS Mulsant

Key to New World species: Cartwright (1948)
P. atlanticus Cartwright ON
P. caesus (Creutzer)t - BC ON PQ

Subfamily OCHODAEINAE
Genus OCHODAXUS Audinet-Serville
Cadocera Eschscholtz
Psephus I\ttby
Stornphax Fischer von Waldheim
O. luscinus Howden _BCAB
O. musculus (Say)
americanus Westwood
-oN
O. opacus LeConte MB
O. simplex LeOonte AB SK MB

t49
Scarabaeidae

Subfamily GEOTRUPINAE
Key to North American species: Howden (1955)
Tribe Bolboceratini
Genus BOL,IOCEROSOMA Schaeffer
B. bruneri f.tawson & McColloch MB

Genus EUC.ANTHUS Westwood

Key to Ncrth American species: Howden (1964)
E. greeni Rcbinson AB - MB
E. lazarus iltabricius; - ON
melibot us (Fabricius)

Genus BOLTIOCERAS Kirby

Odontaeus Fischer von Waldheim
Key to Ncrth American species: Wallis ( 1928)
B. falli (wallis) - SK MB ON PQ
B. filicornis Sayt - SK MB
B. liebecki l ilallisr - ON PQ - NS
B. obesus (L.econte) - BC AB

Tlibe Geotrupini
Genus GEOIRUPES Latreille
(Subgenus (IEOTRUPES s. str.)
G. stercorarius (Linn6)t PQ NB - PE - NF
(Subgenus INOPLOTRUPES Jekel)
(M ela notr u p e s Blanchard)
G. ball Jek:l MB ON PQ NB NS
similis Jekel
G. hornii Bl rnchard - ON PQ NF
(Subgenus ()NYCHOTRUPES Jekel)
G. semiopacus Jekel MB ON PQ
b la chb ur nii Melsheimer
melsheimeri Jekel
oualipennis Jekel
G. splendidus miarophagus Say - 'ON PQ
G. splendidrrs splendidus - - NS
(Fab:ricius)
gilnichi Jekel
mixtus Ilorn
starki .tekel

Subfamily GLAPFIYRINAE
Genus LICI INANTHE Burmeister
Dasyde'ra LeConte
Key to sp':cies: Carlson (1980)
L. rathvoni LeConte) - BC
canina Ilorn
edwara'siHorn
Subfamily TROGINAE
Genus GLARESIS Erichson
Key to Nt,rth American species: Gordon (1970c)
G. canadens,is W.J. Brown AB - MB
G. medialis Gordon - BC

Genus TRO.K Fabricius

Omorgus Erichson
Pseudotrox Robinson
Key to North American species: Vaurie (1955)
T. aequalis Say MB ON PQ NB NS
T. atrox LeConte - BC AB SK - ON

150
 Scarabaeidae

T. capillaris Say ON
T. fascifer LeConte _BC
T. foveicollis Harold ON
aciculeatus Robinson
T. hamatus Robinson ON
T. robinsoni Vaurie AB SK MB
T. scaber (Linn6) -ec AB _MBON PQ NB NS-
T. scabrosus Palisot de Beauvois ON
T. sonorae LeConte NT AB SK
alternans LeConte
lecontei Harold
T. sordidus LeConte ON PQ
T. spinulosus spinulosus Robinson ON
T. striatus Melsheimer ON
T. unistriatus Palisot de Beauvois AB SK MB ON pe NB NS ID NF
porcatus Say
T. variolatus Melsheimer ON PQ NB NS
erinaceus LeOonte
monticola Robinson

Subfamily MELOLONTHINAE
T?ibe Sericini
Genus SERICA Macleay
Camptorhina Kirby
Stilbolemma Harris
S. anthracina LeConte BC AB SK
crdssata Walker
frontalis LeConte
robusta LeConte
ualidaHarcld.
S. atracapilla (Kirby) _BC MB ON PQ NB NS
cucullata Dawson
S. curvata LeConte
S. elusa Dawson
S. georgiana Leng
NT BC
T:^ MB

-oN
PQ
PQ NB NS PE
lecontei Dawson
S. intermixta Blatchley NT BC AB SK MB ON PQ
S. parallela Casey
_SK -oN
MB ON
PQ
S. sericea (Illiger) NT BC PQ
S. sponsa Dawson -oN
MB ON PQ NB NS PE
S. tristis LeConte

Tribe Melolonthini
Key to North American genera: Hardy (1974)
Genus DIPLOTAXIS Kirby
Alobus LeConte
Diazus LeConte
Orsonyx LeConte
Key to species: Vaurie (1958; 1960)
D. atlantis Fall ON
nigra Caftwight
D. brevicollis LeConte BC AB
nigropicea (Walker)
D. haydenii LeConte AB SK
impressifrons Caziet
innoxia LeConte
leuicoxa FalI
D. obscura LeOonte AK BC AB SK MB
cornpacta Fall
consequens (Walker)
protninens Fall
D. sordida (Say) ON PQ
carbonaria Burmeister
rugosioides Schaeffer
D. subangulata LeConte BC
californica Schaeffer
D. tenebrosa Fall BC

151
Scarabaeid,re

D. tristis K rby MB ON pe NB NS pE
c or p ul e n t a Burrneis ter

Genus PHYLLOPHAGA Harris

Ancylonycha Blanchard
Endrosa LeConte
Gymnis LeConte
Lachnr>sterna Hope
Triche:;thes Erichson
Key to North American species: Luginbilt and Painter (1g53)
P. anxia (LeConte) NT BC AB SK MB ON pe NB NS pE - NF
alpina (Linell)
breuiccllis (Blanchard)
cephal,.ca (LeConte)
dubia:.Smittr)
inspert:ta (Smith)
p u nc t i r::ol li s (Blanchard)
uninot:tta (Walker)
P. balia , Sa/r ON
comola (Burmeister)
P. crenulatit tFrrilichr ON
georgicana (Gyllenhal)
P. drakii (Kirby) AB SK MB ON pe NB NS pE
consimilis (LeConte)
grandi.; (Smith)
P. ephilida 'Sayt
burmasteri (LeConte)
P. fervidatl'abricius)
arcuatut (Smith)
quercir a (Knoch)
P. lraterna Jarris ON pQ
cognaflt (Burmeister)
P. fusca (Fr tlich) - BC - oN pe - NS NF
feruens (Gyllenhal)
P. futilis Lt.Conte)
(
- ON PQ NB
decidutt (LeConte)
gibbosc,. (Burmeister)
serricornis (LeConte)
P. gracilis tllurmeisterr - ON
inana (LeConte)
uoluulo. (LeConte)
P. hirsuta tl(nocht - ON pe
P. hirticula Knochr - ON PQ
P.ilicistKn,rcht - ON
ciliata LeConte)
fimb r ia ta (Burmeister)
ybnesi [ianderson
porcina (Hentz)
P. inversa tllorn r ON
P. lanceolat.t tSayt MB ON
arizoncte von Bloeker
cazieri von Bloeker
grisian,z von Bloeker
P. longispin,r tSmithr - ON PQ
P. marginal s (LeConter ON PQ
P. nitida t Lt Conte) AB - MB ON PQ
innominata (Smith)
limula lHorn)
P. rugosa tl\lelsheimert AB - MB ON pQ
P. tristis tFa,,bricius) - BC - ON
pilosicctllis (Knoch)
P. vilifrons tl.eConter ON pe
hirticep,s (LeConte)

Genus POL'IPHYLLA Harris

MacrorLoria CroLch
Key to Ncrth American species: Young (1988)
P. crinita Le Conte - BC
incolum.is Casey
mystica Casey

L52
 Scarabaeidae

ona von Bloeker

pacifica Casey
peruersa Casey
santarosae von Bloeker
P. decemlineata (Say) BC AB SK
castanea Casey
com stockiana von Bloeker
laticauda Casey
matrona Casey
oregona Casey
parilis Casey
peruersa Casey
potsiana Casey
reducta Casey
ruficollis Casey
squamotecta Casey
P. hammondi LeConte AB SK MB
acomana Casey
bisinuata Casey
diffusa Casey
impigra Casey
latifrons Casey
molesta Casey
oblita Casey
oklahomensis }].atc}r
pimalis Casey
proba Casey
rufescenta Tanner
sejuncta Casey
speciosa Casey
squatnicauda Casey
subuittata LeConte
uerecunda Casey
P. variolosa (Hentz) ON PQ

Genus AMPHIMAILON Berthold

A. majalis (Razoumowsky)f ON PQ

Tlibe Pachydeniini
Genus PHOBETUS LeConte
Key to species: Hardy (1978)
P. comatus LeConte BC
leachi Banett

Tribe Macrodactylini
Genus DICHELONYX Harris
Dichelonycha lltrby
Key to North American species: Saylor (1945)
Key to eastern North American species: Brown (1946)
D. albicollis (Burmeister) ON PQ NB NS
D. backii (Kirby) _NTBCABSKMB oN PQ
arizonica Barrett
D. canadensis (Horn) oN PQ lrD
D. diluta (FalI) ON PQ NB NS-
_PE
D. elongatula (Schtinherr)
D. fulgida (LeConte) _BCAB _:a)u
crotchi (Horn)
mormona (Fall)
oregona (Van Dyke)
D. kirbyi W.J. Brown AB SK MB
D. linearis (Gyllenhal) MB ON PQ
hexagona (Germar)
testacea (Kirby)
D. subvittata (LeConte) MB ON PQ NB NS
D. truncata (LeConte) sc AB SK

153
Scarabaeida':

D. vicina (Fall) - BC
columbinna Hopping
deserta Hopping

Genus MAC I,ODACTYLUS Latreille

Stenothyax Haris
M. subspinor,us (Fabricius) oN PQ
barbatu:,s Fitct'
elongatus (Herbst)
poly p ha. g us (Burmeister)

Tribe Hopliini
Genus HOPLIA Illiger
Echyra Erichson
Key to No:th American species: Hardy (1977)
H. callipyge LeConte BC
cazieri \loyer
conuexul,a LeConte
desertic'tla Boye'r
humbol,.ltensis Boyet
irrorata LeConte
lecontei Dalla Torre
mutata Gemminger & Harold
oregona LeConte
pubicollis LeConte
utahensis Boyer
H. Iaticollis LeConte _MB?_
H. modesta llaldeman ON
barbata Blatchley
heluola Melsheimer
monticota Melsheimer
singularis Burmeister
H. sackenii LeConte BC AB
H. trifasciate. Say MB ON PQ NB NS
ferrisi E:,oyet
primarit Burmeister
lrlsfis N.,elsheimer

Subfamily RUTELINAE
Tribe Anomalini
Genus ANOUIALA Samouelle
Anomalepta Casey
Anomalopus Casey
Hemispilota Casey
Oliganomala Casey
Paranornala Casey
Rhombanalia Casey
Spilota Burmeister
Key to North American species: Potts (1977)
A. binotata {'}yllenhal t ON
compacia Casey
maculata (Castelnau)
marginella LeConte
unifascifia Say
A. lucicola iF abricius) ON
olrolo (llabricius)
cinctello Burmeister
moerens (Fabricius)
nigritula (Castelnau)
thoides Nonfried
p hy I lope' r
pinicola Melsheimer
q uadr im.ac ulat a Dr apiez
A. undulata .Vlelsheimer ON
canadensis Casey
dubia BlatchTey

r54
 Scarabaeidae

floridana Casey
nigropicta Casey
saginatula Casey
uarians Burmeister

Genus STRIGODERMA Burmeister

Alamona Casey
Strigodermella Casey
S. arboricola (Fabricius) ON

Genus POPILLIA Audinet-Serville

P. japonica Newman-i ON PQ NS

Tribe Rutelini
Genus PELIDNOTA Macleay
Key to North American species: Hardy (1975)
(Subgenus PELIDNOTA s. str.)
(Pelidnotidia Casey)
P. punctata (Linn6) MB ON PQ
breuicollis Casey
breuis Casey
debiliceps Casey
hudsonica Casey
lutea (Olivier)
oblonga Casey
pallidipes Casey
ponderella Casey
strenua Casey
tarsalis Casey
texensis Casey

Genus COTAI-PA Burmeister

Ciocotalpa Saylot
C. lanigera (Linn6) SK MB ON
molaris Casey
fou Wickham
uernicata Casey

Subfamily DYNASTINAE
Key to species: Endrodi (1985)
Tribe Cyclocephalini
Genus CYCLOCEPHALA Latreille
Dichromina Casey
Ochrosidia Casey
Spilosota Casey
C. borealis Ar-rolv NS
uillosa Burmeister
C. longula LeConte BC
abrupta (Casey)
ambiens (Casey)
californica Atrow
marcida (Casey)
modulata (Casey)
obesula (Casey)
oblongula (Casey)
phasma (Casey)
prona (Casey)
reflexa (Casey)
regulifrons (Casey)
rustica (Casey)

155
Scarabaeidae

Tribe Pentodontinr
Genus LIGf RUS Burrneister
(Subgenus L,IGYRODES Casey)
L. relictus (Say) MB ON PQ NB
clypealis (Casey)
dawsoni (Casey)
paruice,:s (Casey)
quadriT ennis (Casey)
uernicicollis (Casey)
(Subgenus L,IGYRUS s. str.)
(Anagrylius Casey)
(.Ligyretilus Casey)
L. gibbosus 1,;ibbosus (DeGeer) AB ON
bicorniculatus Casey
breuiusculus Casey
curtiper'.nis Casey
farctus Jasey
juuencus (Fabricius)
lacustri; Casey
laetulu:, Casey
Iaticaut:ta Casey
Iongulu; Casey
lucubla tdus Casey
morio (LeConte')
parallelus Casey
punctiutuda Casey
remotut: Casey
rubidus Casey
texanus Casey
uariolos us (Burmeister)
uirginic'"ts Casey

Genus APHONUS LeConte

Key to No:th American species: Gill and Howden (1985)
A. castaneus (Melsheimer) PQ
cubiforr zis Casey
obesus (Burmeister)
saginat,"rc Casey
trapezictllis Casey
A. tridentatts (Say) ON PQ
aterrim;ts Casey
congestt,ts Casey
elongatus Casey
frater L.:Conte
modula,lus Casey
politus l)asey
scutella:is Casey

Tribe Oryctini
Genus XYLC.,RYCTES Hope
X. jamaicensis (Drury) ON PQ
americanus (Palisot de Beauvois)
faunus \)asey
hebes Casey
lacustri:; Casey
obsolescens Casey
sotyrus (Fabricius)
tenuicornutus Casey

156
 Scarabaeidae

Subfamily CETONIINAE
Tlibe Cetoniini
Key to New World genera: Hardy (1988)
Genus EUPHORIA Burrneister
Erirhipidia Casey
Erirhipis Burmeister
Euphorhipis Casey
Euphoriaspis Casey
Haplophoria Casey
Isorhipina Casey
Rhipiphorina Casey
E. fulgida (Fabricius) _ONPQ
E. inda (Linn6) AB SK MB ON PQ
barbata (Say)
brunnea (Gory & Percheron)
marilandica (Frdlich)
nigripennis l{lages
E. rufobrunnea Casey BC

Tribe Cremastocheilini
Genus CREMASTOCHEILUS Knoch
Key to North American species: Potts (1945)
(Subgenus CREMASTOCHEILUS s. str.)
C. armatus Walker BC
cribripennis Casey
densicollis Casey
maritimus Casey
montanus Casey
obliquus Casey
pilosicollis Horn
C. canaliculatus
hentzi Haris
Kirby -oNPQ
C. castaneus Knoch AB SK MB NS
breuisetosus Casey
lecontei Westwood
pocularis Casey
C. crinitus LeConte BC AB
bifoueatus Van Dyke
pugetanus Casey
C. knockii LeConte SK MB
areolatus Casey
crenicollis Westwood
gracilipes Casey
(Subgenus ANATRINODIA Casey)
C. wheeleri LeConte SK MB

Subfamily TRICHIINAE
Key to North American species: Howden (1968)
Tlibe Osmodermini
Genus OSMODERMA LePeletier & Audinet-Serville
Gymnodus Kirby
O. eremicola (Knoch)
O. scabra (Palisot de Beauvois)
-oNPQ
_ONPQ NB
NS
NS
beauuoisii Bainbridge
coseyi Schenkling
cauiceps Casey
delicatula Casey
foueatus (Kirby)
gracilipes Casey
lacustrina Casey
rugosa (Kirby)
O. subplanata Casey AB MB ON
montana Wickham

t57
Byrrhidae

Tribe Trichiini
Genus TRITIHIOTINUS Casey
T. affinis (C'ory & Percheron) - ON PQ
paruuius Casey
uentricosus Casey
T. assimilis, (Kirby) NT BC AB SK MB ON PQ NB NS
bistriga (Newman)
uariabilis (Burmeister & Schaum)
T. piger (Fabriciust ON
drumrzondi (Gory & Percheron)
reductus Casey
r ot unc:,'.icol I is (Kirby)
T. viridans tKirby) - ON

Genus GNORIMELLA Casey

G. maculosr tKnocht - ON PQ
bigs6i, (Kirby)
dissiniilis (Gory & Percheron)

Superfamily Byrrhoidea
Family BYRRHIDAE pill beetles
P.J. Johnson

The lamily Byrrhidae is represented in Canada Taxonomically, the family is reasonably well
and Alaska by 26 of the 34 North American species, known in North America. Johnson (-t987) has
compare(l with about 280 species around the world. treated the bionomics, larval taxonomy, and bio-
The greatest diversity of Canadian byrrhids is geography of the North American generur, and has
found in the montane provinces. All species in provided a key to the North American taxa (1991b).
Canada irre bryophagous and express microhabitat
specifrcity. Many species are associated with the
moss Cerotodon purpureus on disturbed sandy-
gravelly soils. Species of the coastal forests of
British C.iolumbia and Alaska are closely associated aK (14); yK (9); NT (13); BC (15); AB (1:2); SK (5);
with certain forest communities and are more MB (7); ON (10); PQ (12);NB (2);NS (a); l,B (5);NF
polyphagous in their host mosses. (4) . H (3); I (2)
Subfamilv SYNCALYPTINAE
Genus CHIiETOPHORA Stephens
C. spinosa Rossi t+ - BC - ON PQ

Genus CURIMOPSIS Ganglbauer

Key to North American species: Johnson (1986)
C. albonotata (LeConter - YK AB
grisea LeConte
C. echinata (LeOonte) AK - NT BC AB SK PQ
breuicr'tllis Casey
C. moosilar.ike Johnson AK - NT - AB - PQ - LB
C. seLulosa (Mannerheim) AK - NT

Subfamilv BYRRHINAE
Tribl Byrrhini
Genus BYRRHUS Linn6
Key to North American species: El-Moursy (1970)
B. americanus LeConte MB ON PQ - NS - LB
angustulus Casey
brunnescens Casey

158
 B1a-rhidae

criddlei Casey
imperitus Casey
manitobae casey
neglectus Casey
B. concolor Kirby AK YK NT SK
laramiensis Casey
rigens Casey
B. cyclophorus Kirby YK NT AB SK MB ON PQ LB
canterius Casey
dfficilis Casey
fasciatus atct.
fuluouestitus Casey
stolidus Casey
torpidus Casey
B. eximius LeConte AK YK NT BC
callidus Casey
centralis Casey
explicatus Casey
B. geminatus LeContex I'K PQ
pacificus Casey
pettiti }{otn
B. kirbyi LeConte AK NT BC AB MB ON PQ NB NS LB NF
consuetus Casey
egenus Casey
perditus Casey
picipes Casey
uafer Casey
wickhami Casey
B. pilula (Linn6)x NT

Genus PORCINOLUS Mulsant & Rey

P. undatus (Melsheimer) AB SK MB ON PQ
crescentifer Casey
glabellus Melsheimer
hystris Casey
murinus Fabricius

Tlibe Simplocariini
Genus SIMPLOCARIA Stephens
S. elongata J.Sahlberg* - YK NT MB ON
remotaW.J.Btown
S. metallica (Sturm) NT BC AB PQ NF
colurnbica Casey
subnuda Casey
tesselata LeConte
S. semistriata ( Fabricius)t BC ON PQ NS NF

Genus LIOLIGUS Casey

L. nitidus (Motschulsky) AK BC
aequabilis Casey
heeni Casey
striolatus Casey

Genus LIOON Casey

L. simplicipes (Mannerheim) AK BC
puncticeps Casey
inflatum LeConte
speculare Casey

Genus EXOMELLA Casey

Key to North American species: Johnson (1985)
E. pleuralis (Casey) AK BC

159
Buprestida:

Tlibe Pedilophorini
Genus EUSOMALIA Casey
E. lecontei tWickham) BC

Genus LISIIEMUS Casey

L. acuminabus (Mannerheim) AK BC
formosus Casey
satelles Casey

Genus MOIIYCHUS Erichson

M. aeneolur; (LeConte) AK NT BC AB MB ON
lateralis (.Fall1
albertt:. nus (W.J. Brown)
szbseloszs (Fall)
hesper rc Wickham
M. oblongur; (LeConte) AK BC AB
subcup,reus (Fall)
insulsus Casey

Tribe Tylicini
Genus TYLICUS Casey
T. subcanur, (LeConte) \T( NT BC AB SK ON PQ

Genus CYT I LUS Erichsor

C. alternatrLs (Say) MB ON PQ NB NS LB NF
nigrans Casey
C. mimicus Casey NT BC AB
longultLs Casey

Superfamily Buprestoidea
Family BUPRESTIDAE metallic wood-boringbeetles
A. Dauies

One hundred and fifty-two species of his handbook has been used as the basis for this
Buprestidae have been recorded in Canada and checklist, with updated distributiorLal data.
Alaska. The adults are usually found resting or Further revision of the species in such groups as
ovipositi eg on the bark and foliage of trees and Anthaxiinae is required before a proper subgeneric
shrubs, and feed chiefly on pollen-rich flowers; they arrangement can be established. It should be noted
are also ;active fliers. The larvae develop mainly that the number of species recorded in the northern
under ther bark of the host plants, which are usually regions does not reflect the number of species pre-
fairly specific, although a few are leafminers, gall- sent but rather only the lack of collecting in those
makers, and coniferous cone borers. regions.
Major taxonomic treatments of the North
America.n buprestids include those of LeConte AK (11);YK (13); NT (18); BC (75);AB (57); SK (a1);
(1860) and Chamberlin (1926). Bright (1987) pro- MB (63); ON (99);PQ (80);NB (36);NS (22);pE (7);
vides keys and descriptions ofthe Canadian species; LB (4);NF (5) . H (1); I (2)

160
 Buprestidae

Subfamily POLYCESTINAE
Tribe Polycestini
Genus CHRYSOPHANA LeConte
C. placida (LeConte) BC AB
cupreola Van Dyke
strandi Obenberger

Subfamily ACMAEODERINAE
Tlibe Acmaeoderini
Genus ACMAEODERA Eschscholtz
A. idahoensis Barr BC
A. pulchella (Herbst) AB -llsoN-
arizonaeHotn
flauosignata Gory
imtnaculata Horn
mixtaLeConte
ornata Olivier
uoluulus Castelnau
A. tubulus (Fabricius) ON
culta (Weber)
geranii (Harris)
q uattuor de cimg uttata (F abicirs)
A. vandykei Fall
A. variegata LeConte
distincta Kerremans
Iucia FalI

Genus ANAMBODERA Barr

A. gemina (Horn) BC
nebulosa (}lorn)

Subfamily CHALCOPHORINAE
T?ibe Chalcophorini
Genus CHALCOPHORA Solier
C. flortis LeConte MB ON PQ NB
cupreola Casey
Iaurentica Casey
C. liberta (Germar) MB ON PQ
borealis (Gory & Castelnau)
paruiceps Casey
C. virginiensis (Drury) - BC AB SK MB ON PQ NB NS
angulicollis (LeConte)
antennalis Casey
breuicollis Casey
c upr eomac ulata (Goeze)
ingens Casey
lacustris LeConte
melanotum Muttkowski
montana Casey
nouaebo racensis (Fitch)
obliterata Casey
obscurata (FiLch)
oregonensis (Filc}r)
pallida Kerremans
prominens Casey

Genus TEXANIA Casey

T. campestris (Sayt ON
substrigosa (Gory & Castelnau)

161
Buprestidae

Subfamily BUPRESTINAE
Tlibe Dicercini
Genus SPE CTRAJ-IA Casey
S. gracilipes t Melsheimer) ON

Genus DICERCA Eschscholtz

Argante Gistel
Stenuris I{ttby
Key to North American species: Nelson (1975)
D. callosa callosa Casey AX YK NT BC AB SK MB ON PQ
pertirutx Casey
rigida Casey
tetrica Casey
D. callosa f rosti Nelson _BCAB
D. caudata LeOonte NT_ABSKMBONPQ NB NS
abrup;a Casey
biangulata Casey
cupreola Casey
filiola Casey
inflatula Casey
longicu.ud.a Casey
pisciformis Casey
D. crassico.llis LeConte BC AB
californica Crotch
hesperica Casey
D. divarica;a (Say) MB ON PQ NB NS
aestiuo Casey
angus icauda Casey
aurich alcea Melsheimer
dabio Melsheimer
incisa Casey
lirnulo Casey
nigra l)asey
parumpunctata Melshermer
rustict:,, Casey
subaet,ualis Casey
uancotlueri Casey
D. dumolini(Castelnau & Gory) SK MB ON PQ
consobrina Melsheimer
D. hesperoboreaiis Hatch & Beer _BCAB
D. horni Crotch _BC
ampliuta Casey
D. lugubris LeConte NT SK MB ON PQ NB
auster't Casey
lacustt'is LeConte
morio Casey
D. lurida (tr'abricius) oN PQ
erosa Casey
floridae Casey
gaudens Casey
gracilis Casey
indis tinc ta Mel sheimer
indura,ta Casey
innocua Casey
leuette,;. Casey
porcat,'tla Casey
pruinasa Gory
regula,is Casey
sagor lasey
seriato. Casey
truncata Casey
D. pectorosa LeConte _BC_SK
D. pugionata (Germar) ON
D. punctulata (Schtinherr) oN PQ
pinorum Casey
transuersa Say
D. sexualis Crotch _BC
D. tenebrica (Kirby) AK YK NT BC AB SK MB ON PQ NB NS
longipennis Casey
prolongata LeConte

162
 Buprestidae

seuera casey
subargentea Casey
subcuprea Casey
sulcatula Casey
D. tenebrosa (Kirby) AK YK NT BC AB SK MB ON PQ NB NS PE LB
acerba Casey
bifoueata LeConte
chrysea Melsheimer
distinguenda (Castelnau & Go.y)
montana Casey
mystica Casey
stolida Casey
wickhami Casey
D. tuberculata (Castelnau & Go.y) oN PQ NB NS
asperata Castelnau
hilaris LeConte
mancaLeConte
molitor Melsheimer
scobina Chewolat

Genus DESCARPENTRIESINA Leraut

Key to North American species: Evans (1957)
D. cyanipes (Say) NT BC AB SK MB ON PQ NB
apicilla (Obenberger)
collaris (Obenberger)
cupripes (Casey)
debilis (LeConte)
erecta (Gory)
paruiceps (Casey)
D. ferrea (Melsheimer) _BCABSK MB ON PQ
D. fraseri (Chamberlin) -BC
D. montana (Chamberlin) _BC MB
D. thureura (Say) MB
costicollis (Gory)

Tribe Buprestini
Genus TRACHYKELE Marseul
T. blondeli Marseul _BC
T. nimbosa Fall -BC
Genus BUPRESTIS Linn6
Ancy lo c h ir a Eschscholtz
Anoplis l{trby
Gymnota Gistel
Key to North American species: Helfer (1941)
B. adjecta (LeConte) BC AB
breuis Casey
B. aurulenla Linn6 BC AB MB ON
adulans Casey
aemula Casey
affinis Casey
bicostata Dejean
chry sochlora (Philippi)
fabulosa Casey
lauta (.LeConLe)
nupta Casey
prospera Casey
radians (LeConte)
tacornae Casey
uenusta Casey
uillosa (LeConte)
B. confluenta Say YK_BC AB SK MB ON PQ
tesselata Casey
B. fasciata Fabricius SK MB ON PQ NB NS
fortunata Casey
fulgens Casey
lherminieri Chevrolat
saturata Casey

163
Buprestidae

sexmaculata Hausmann
sexplagiata (LeConte)
B. intricateL Casey BC AB
con,tortae Hopping
murraydnae Hopping
B. laeviventris (LeConte) BC
pugeta,na Casey
B. langi Mannerheim AK BC AB SK MB
angusta Casey
bistrinotata Casey
callida Casey
crenata Casey
d.epressa Casey
fastidt.osa Casey
gramitea Casey
incolumis Casey
Iaeuicirps Casey
medio:ris Casey
obliqu.a Casey
oregon.a Casey
ornata (Walker)
patruelis Casey
sediticsa Casey
uiridirnicans Casey
B. lineata ltabricius BC MB ON PQ NS
douisi Nicolay & Weiss
B. lyrata Casey BC AB SK MB ON NS
acomana Casey
adducta Casey
caligitnsa Casey
fuscat)asey
lecontt::i Saunders
morosd Casey
nigricuns Casey
B. maculat.ventris Say NT AB SK MB ON PQ NB NS PE
pagancrum l{trby
rusticorum Kirby
sexnotzta Gory & Castelnau
B. maculipennis Gory SK MB PQ
deficiens Casey
fusiformis Casey
inconstans Melsheimer
leporirua Casey
reductt Casey
scriptt:. Casey
B. nuttalli Kirby AK YK NT BC AB SK MB ON PQ
altern':tns (LeConte)
boulderensis Casey
conici<,auda Casey
consulcris Goty
contor;la Casey
diruptzns Casey
flauopicta Casey
grauidula Casey
torua Oasey
B. salisbu4rensis Herbst ON
cazieri Helfer
ultran,rarina Say
B. striata I abricius oN PQ NB
canad,ensis Casey
imped.it.ta Say
obscura Casey
B. subornal;a (LeConte) BC
adoneu Casey
histrio Casey
punctiuentris Casey
rubrorrctans Casey
uiolescens Casey
B. sulcicollis (LeConte) MB ON PQ NS NF
lateralis Casey

t64
 Buprestidae

Tribe Melanophilini
Genus MELANOPHILA Eschscholtz
Phaenops Dejean
Key to North American species: Sloop (1937)
M. abies Champlain & Knull ON PQ NB
o6iells Obenberger
M. aeneola Melsheimer ON PQ
metallica Melsheimer
M. californica Van Dyke BC
M. drummondi drummondi (Kirby) AK YK NT BC AB SK MB ON PQ NB LB
guttulata Mannerheim
umbellatarum (Kirby)
M. fulvoguttata (Harris) BC SK MB ON PQ NB NS PE NF
croceosignata (Castelnau & Gory)
decolorata (Castelnau & Gory)
ftarrisi Obenberger
octospilota (Castelnau & Gory)
pacifica Obenberger
M. gentilis LeConle BC
prasina LeConte
M. intrusa Horn BC

Genus OXYPTERIS Kirby

Apatura Castelnau & Gory
O. acuminata (DeGeer)* AK YK NT BC AB SK MB ON PQ NB NS PE NF
ap pendiculata (Fabricius)
i mma c u I ata ( Mannerheim)
longipes (Say)
morio (Fabricius)
O. occidentalis Obenberger BC

Tribe Anthaxiini
Genus ANTHAXIA Eschschoitz
(Subgenus MELANTHAXIA Richter)
A. aeneogaster Castelnau & Gory BC
aerica Ctotch
grosso Obenberger
strigata LeConte
A. aenescens Casey _BC
A. inornata (Randall) _YKNTBCABSKMBONPQNBLB
expdnsa LeConte
foueicollis LeConte
imperfecta LeConte
A. porella Barr BC
A. prasina Horn BC
falsula Obenberger
A. retifera LeConte AK YK BC MB
( Subgenus IIAPLANTHAXIA Reitter)
A. deleta (LeConte) _BC
A. pseudotsugae (Chamberlin) _BC
A. quercata (Fabricius) ON PQ N"
cuneiformis (Gory)
A. viridicornis (Say) SK MB ON PQ
subaenea (LeConte)
uiridifrons (Gory)

Genus AGRILAXIA Kerremans

A. flavimana (Gory) ON
gracilis (Melsheimer)

165
Buprestidae

Subfamily CHRYSOBOTHRINAE
Tribe Chrysobothrini
Key to North American species: Fisher ( 1942)
Genus CHRYSOBOTHRIS Eschscholtz
Knowltonia Fisher
Odontomus l\ttby
C. adelpha Gemminger & Harold ON PQ
soror LeConte
C. aeneola l,eConte AB SK
purpurata Bland
(?)
C. azttea LeConte BC AB SK ON PQ
lecontet Leng
C. beeri Barr _BC
C. blancharrli Horn AB SK ms oll PQ NB
C. breviloba Fall _BCAB
C. carinipernis LeConte _BCAB
C. caurina IIorn _BC
C. columbiana Barr _BC
C. cribraria Mannerheim NT_ABSK lln Of,l pe
calcarota Melsheimer
f'Loricola Gory
lalo Kerremans
C. dentipes iGermar) BC SK MB ON PQ NB
c har acr e r is tica ( Harris)

planattt Gory & Castelnau

posticatis Gory & Castelnau
rotund,collis Gory & Castelnau
C. femorata (Olivier) BC AB SK MB ON PQ NB NS
alaban,ae Gory
dissim,,.Iis Dejean
fastidit,sa Gory
insculF'ta (Herbst)
lesueuri Gory & Castelnau
misello LeConte
nigrituta Gory & Castelnau
obscurr.t LeConte
quadri;',mpresso Gory & Castelnau
C. monticoler Fall BC AB
grandis Chamberlin
C. harrisi (Flentz) BC AB-MBON PQ NB NS_
C. laricis Van Dyke BC AB
C. leechi Barr BC AB
C. mali Horn BC AB SK MB
C. neopusilla Fisher PQ NS_
C. nixa Horr
C. oregona Ohamberlin
planon .arginafo Chamberlin
BC
BC _ _ T:'. fu

C. orono Frost MB ON
C. pseudots rgae Van Dyke sc
C. pusilla Gory & Castelnau MB ON PQ NB
bigutta,ta Gory & Castelnau
str ang ulata Melsheimer
C. rugosicelrs Melsheimer
C. scabripennis Gory & Castelnau
proxima (Kirby)
xr nc AB ix MB 3N i3 NB PE

scabra Goty
C. sexsignata (Say) oN PQ NB PE
germari Gory & Casteinau
ignipes Gory & Castelnau
serguttata (Say)
C. sylvania Fall _BC
C. trinervia (Kirby) AK YK NT BC AB SX MB ON PQ NB NS LB NE
Motschulsky
cicat r i cos a
C. verdigripennis Frost AB SK MB ON PQ NB NS
C. viridiceps Melsheimer MB ON
C. vulcanica LeConte nc AB
canadensis Chamberlin

166
 Buprestidae

Subfamily AGRILINAE
Tribe Coraebini
Genus EUPRISTOCERUS Dey'r'olle
(? ) R haeb oscelis LeConte

E. cogitans (Weber) ON PQ NB
ignarus (Fabricius)
ruficollis (Herbst)

Tlibe furilini
Genus AGRILUS Curtis
A. acutipennis Mannerheim _MBONPQ
latebrus Castelnau & Gorv
A. anxius Gory AK BC AB SK MB ON PQ NB NS NF
grauis LeConte
torpidus LeConte
A. arbuti Fisher _BC
manzanitae Chamberlin
A. arcuatus (Say) MB ON PQ NB
obliquus LeConte
A. atricornis Fisher -ONPQ
_ONPQ
A. aurichalceus Redtenbacherf
A. bilineatus bilineatus (Weber) MB ON PQ
aurolineatus Gory
biuittatus l{trby
flau olineatus Mannerheim
triuittatus Horn
A. bilineatus carpini Knull
A. burkei Fisher -ncAB--9*PQ-----
A. celti Knull -oN
A. cephalicus LeConte -BCAB-MBONPQ
A. crataegi Frost AB ON PQ
A.
A.
A.
criddlei Frost
crinicornis Horn
cyanescens (Ratzeburg)t
_ _ _:"yu3Ni3Iu_
-ONPQ
_ _ _

A. defectus LeConte -ONPQ

A. egenus Gory -oN
A. fallax Say
impressipenni.s Uhler
-oNPQ
zemes Gory
A. frosti Knull _MBONPQ
A. geminatus (Say) oN PQ
A. granulatus populi Fisher BC AB
tr ic hocar p ae Chamberlin
A. horni Kerremans

- - - -"gN;:-
A. imbellis Crotch
A. juglandis Knull
A. lacustris LeConte
cuneus LeConte
pubiuentris Ctotch
A. lecontei Saunders -oN
subfasciatus LeOonte
A. liragus Barter & W.J. Brown
A. malvastri Fisher
A. masculinus Horn
---:"ffisKYuoNPQNB---
-SKMBONPQ
A. nigricans Gory
auricomus Frost
-oN
A. obsoletoguttatus Gory _ONPQ
interruptus LeConte
subaeratus Hatris
A. otiosus Say MB ON PQ
uirens Goty
A. parvus Saunders MB
pusillus Say
A. pensus Horn AB SK MB ON PQ NB NS PE
betulae Fisher
A. politus (Say) AK YK NT BC AB SK MB ON PQ NB NS
canadensis Obenberger
cupreolus LeConte

167
Buprestidar:

desertus LeConte
plumbcus LeConte
solitar,t,us Harold
A. pseudoccryli Fisher -MBONPQ
A. putilius l5ay oN PQ
A. rufrcollis (Fabricius) -MBONPQ
cupricollis Goty
A. sayi Saurders _PQ NB NS
browni Catlson & Knight
lateralis (Say)
A. subcinctus Gory ON
A. vittaticollis (Randall) BC oN PQ
frenatus Gory
A. walsinghami Crotch BC

Subfamily TRACFTYINAE
Tribe Pachyschelini
Genus PACHYSCHELUS Solier
P. confusus Wellso & Manley -oN
_ONPQ
P. laevigatts (Say)
oblong zs (Motschulsky)
ouatus (Say)
politus Kertemans
punctatus (Goty)
P. purpurer.,s (Say) AB ON
biimpr essus Motschulsky

Tribe Brachyini
Genus BRACHYS Solier
B. aerosus l1{elsheimer AB SK MB ON PQ
rufescens Nicolay & Weiss
B. aeruginosus Gory
B. ovatus (\Veber)
-oNPQ
_ONPQ
auruleztus (Kirby)
bellporli Nicolay & Weiss
horni }(erremans
laeuicauda LeConte
molestus Gory
termin,zns Castelnau & Gory

Genus TAPFIROCERUS Solier

T. gracilis (iiay) AB SK MB ON PQ
albog u tt at us (Mannerheim)
grossu:; Obenberger
texanus Kerremans
T. nicolayi Obenberger
T. schaefferi Nicolay & Weiss - - ABsr- 3N- - - - - -

168
 Heteroceridae

Superfamily Dryopoidea
Family PTILODACTYLIDAE toed-winged beetles
L. LeSage

The family Ptilodactylidae, which is primarily plant matter; those of Anchytarsus are truly
tropical in distribution, comprises about 500 species aquatic, burrowing in submerged decaying wood in
around the world, but only three occur in Canada. creeks (LeSage and Harper 1976a).
Adults hide in the riparian vegetation or in the leaf Stribling's recent key (1986) is useful for identi-
litter during the day; they are easily attracted to fying the genera.
light. Larvae of Ptilodactyla prefer moist or wet
habitats, such as beaver houses or wet, decaying BC (1);MB (1);oN (2);PQ (3);PE (1)

Genus FTILODACTYI,A Illiger

Key to North American species: Johnson & Frey'tag (1982)
P. serricollis (Say) - BC MB ON PQ PE
elate r ina Gu6rin-M6neville
/usca Melsheimer
P. carinata Johnson & Freytag PQ

Genus ANCIIYTARSUS Gu6rin-M6neville

Key to North American species: Stribling (1986)
A. bicolor (Melsheimer) ON PQ
ofer Gu6rin-M6neville
debilis Ziegler

Family HETEROCERIDAE variegated mud-lovingbeetles

L. LeSage

Heterocerids are represented in Canada and has been followed for this checklist except for the
Alaska by 25 species. The adults are nocturnal and genus Dampfius, which was recently placed in syn-
are readily attracted to light. During the day they onymy witlr' Heterocerus by Miller (1988b). The
hide and make tunnels in sandy areas near water Canadian fauna is relatively well known, but new
courses or standing water, a behavior shared by the species are still being discovered (Miller 1988o,
Iarvae. Adults and larvae feed on zooplankton, 1988b). Both the arctic and the maritime fauna
diatoms, and algae. have been little surveyed. Pacheco (1978) has
Pacheco (1964) proposed a drastic change in the recently cataloged the North American species.
traditional treatment of the family when he
described 15 new genera and restricted the genus
World. Although several of
Heterocerus to the OId aKQ); YK (a);NT (5); BC (e);AB (16); SK (14); MB
his new genera are questionable, his classifrcation (1e);ON (16);PQ (16);NB (5);PE (1)

Tlibe Auryliini
Genus EXPLORATOR Pacheco
E. canadensis (Fall) AK YK NT AB SK MB

Genus CENTURIATUS Pacheco

C. auromicans (Kiesenwetter)
C. compactus (Fall) _ -_ _ :"ffi:"MBg"i8 _ _ _ _ _

r69
Heteroceridae

Genus MICIIOAUGYLES Pacheco

M. moleculus (Fall) _ABSKMB_PQ
M. mundulus (Fall) -AB

Tlibe Heterocerini
Genus LAN'IERNARIUS Pacheco
L. brunneus (Melsheimer) _NTBCABSKMBONPQPE
L. gemmatur tHornt BC AB SK
L. mollinus (Kiesenwetter) - _ABSKMBONPQ
L. parrotus llacheco _NTBCABSKMBONPQNB
L. sinuosus .Pacheco YK NT BC AB SK MB
Genus NEO F{ETEROCERUS Pacheco
N. fatuus lKiesenwetter ) - MB ON NB
N. gnatho {I,econte) YK _BCABSK MB ON pe
labratus (LeConte)
N. longilobu lus Pacheco PQ
N. pallidus tSayt _ABSK MB ON PQ NB
uentrahs (Melsheimer)
N. sandersoni Pacheco MB

Genus HETEROCERUS Fabricius

Dampfius Pacheco
Key to North American species: Miller (1988b)
H. cof laris (liesenwetter) - SK MB ON PQ
H. inciertus Pacheco.) _BCAB MB ON PQ
H. mexicanus Sharp
H. subtilis Vt.V. Miller
-oN
MB ON PQ
H. undatus {Melsheimer; AB SK MB ON PQ NB
H. unicus W V. Miller _BCABSK lvlD

Genus PEDI TATUS Pacheco

P. schwarzi Horn) _ONPQ
P. tenuis tW V. Miller) MB ON
Genus LAPSUS Pacheco
L. tristis (Mannerheim) AK YK NT BC AB SK MB ON PQ NB

Tribe Tropicini
Genus TROI'ICUS Pacheco
T. pusillus {flayt - ON PQ
americanus (Sharp)
Iimbatus (Kiesenwetter)
luteolus (LeConte)

170
 Dryopidae

FamilyLIMNICHIDAE minute marsh-lovingbeetles

L. LeSage

Of the three subfamilies and 15 genera of This group of beetles was first considered as a
Limnichidae described in the Western Hemisphere, tribe of Byrrhidae, but Hinton (1939o, 1939b) ele-
only two genera and four species occur in Canada. vated it to family rank and placed it in the super-
Adults of limnichids are usually active during the family Dryopoidea, after a study of the internal
day near water in leaf litter; they are readily anatomy. The North American limnichid fauna is
attracted to light. The larvae are rarely collected now well known after a series of recent revisions by
and are not represented in most collections. Paulus Wooldridge, who also published a catalog of the
(1970) frrst described the immature stages of two North American species in 1986.
European species: they are terrestrial, Iive in moist
soil, and tunnel under layers of mosses and algae,
which thev use as food. eC (1);MB (1);oN (4);PQ (2)

Subfamily LIMNICHINAE
Tribe Limnichini
Key to North American genera: Wooldridge (1975)
Genus LIMNICHITES Casey
Key to North American species: Wooldridge (1977)
L. huronicus Casey ON
L. olivaceus (LeConte) ____ ON
uirginicus Casey
L. punctatus (LeConte) BC MB oN PQ
austinianus Casey

Genus LUTROCHUS Erichson

Key to North American species: Brown (1972)
L. laticeps Casey ON PQ

Family DRYOPIDAE long-toed water beetles

L. LeSage

Dryopidae includes over 200 species around the The family has not been revised since Brown
world, but only six occur in Canada. Adults are all the taxonomic information.
(1972) put together
essentially phytophagous, feeding on diatoms, The Canadian fauna is reasonably well known for
algae, and decaying wood. The truly aquatic adults Quebec and Ontario (LeSage and Harper 1975).
of Helichus inhabit riflles of water courses, using a Brown (1983o) recently published a catalog of the
hydrofuge plastron for breathing, as do elmids. North American species.
Adults of Dryops are typically riparian. They can
take flight directly from water surface when dis-
turbed. Dryopid larvae are rarely collected; the few
specimens collected were found in moist soil on sC (2); AB (1); SK (1); MB (1); ON (2); PQ (5); NB
banks of creeks where the adults live. (2);NS (2);PE (1);NF (1) ' I (1)

Genus DRYOPS Olivier

(Subgmus YRDOPS Steffan)
D. viennensis (Heer)t PQ

Genus HELICHUS Erichson

Key to North American species: Musgrave (1935); Brown (1972)

L77
Elmidae

H. basalis L,:Conte PQ NB NS
H. columbia.rus W.J. Brown BC
H. fastigiatts (Say) PQ
H. lithophih.Ls (Germar) ON PQ
H. striatus LeConte --ecAB SK MB ON PQ NS NS PE_NF
foueatus LeConte

Familv ELMIDAE riffle beetles

L. LeSage

Elmidae comprises about 1100 species around The family Elmidae is usually split into two sub-
the world: 32 of them occur in Canada. Elmids are families (Brown 1972). The North American fauna is
small beetles, usually less than 3 mm long, and most relatively well known as a result of Brown's generic
are aquatic in both the adult and the larval stages. treatment (1972), his catalog (1983c), and the recent
Adults breathe by means of a hydrofuge ventral plas- revisions of the genera Dubiraphia and Optioseruus.
tron, the arvae with retractile cloacal gills. Adults Stenelrnis, the largest genus, is now under revision.
and larver e inhabit the substratum of creeks and A systematic survey of the eastern Canadian fauna
rivers, feeding on diatoms, encrusted algae detritus, undertaken by LeSage and Harper (1975) focuses on
or submer:ged decaying wood (LeSage and Harper Quebec. The fauna of the Prairie Provinces, western
t976b); they are good indicators of water quality Canada, and the Arctic is still poorly known. The
(Hilsenhoff I977). Pupation occurs near water, present list adds many new provincial records based
under ro<ks or pieces of wood, and in moist soil. on specimens accumulated by the author since
Adults fl1 and are often attracted to light, usually Brown's work and on several unpublished records
soon after emergence. Their hydrofuge plastron used generously provided by H.G. Nelson of the Field
for breathing enables them to stay under water for Museum of Natural History.
the rest of'their life, which, incidentally, may last for
several years. Old specimens are easily recognized YK (1); NT (1); BC (14);AB (11); SK (a);M.ts (7); ON
by their encrusted body, which often carries peritrich (16); PQ (21); NB (10); NS (11); PE (5); L]l (2); NF
ciliates. (5)

Subfamily LARINAE
Tlibe Larini
Key to genera: Brown (1981)
Genus LARI. LeConte
Key to Noth American species: Brown (1972)
L. avara amlrlipennis Darlington BC
L. avara avara LeConte BC

Subfamily ELMINAE
Tribe Ancyronychini
Genus ANC-IRONYX Erichson
A. variegata (Germar) oN PQ
cincta (.Say)
parumctculata (Hardy)

Tribe Elmini
Genus CLEPTELMIS Sanderson
C. ornata (Schaeffer) BC

Genus DUBIRAPHIA Sanderson

Key to No"th American species: Hilsenhoff(1973)
D. bivittata tLeConte)
D. minima E.ilsenhoff
D. quadrinotata (Say)
_ _ T_ MB
MB
MB
ON
ON
ON
PQ
PQ
PQ NB NSpn--
D. vittata (Nl.elsheimer) AB SK MB ON PQ NS

172
 Elmidae

Genus HETERLIMNIUS Hinton

Ceralimnius Hinton
Key to North American species: Brown (1972)
H. corpulentus tl,eContet - BC AB
antennatus (Fall')
H. koebelei (Martinj - BC AB

Genus MICROCYLLOEPUS Hinton

Key to North American species: La Rivers (1949)
M. pusillus pusillus (LeConte) - BC - ON PQ NB
browni (Hatch)

Genus NARPUS Casey

N. concolor (LeConter - BC AB
solutus (W.J. Brown)

Genus OPTIOSERVUS Sanderson

Key to North American species: White (1978)
O. divergens (LeConter - BC AB SK
pecosensis (Fall)
O. fastiditus (LeConte) NT BC AB SK MB ON PQ NB NS PE - NF
O. quadrimaculatus tHorn) - BC AB
O. ovalis tl-eContet - ON PQ NB NS PE LB
ampliatus (Fa'r\)
O. seriatus (Leconte) - BC AB
Brown)
O. trivittatus (W.J. - ON PQ NB NS NF

Genus OULIMNIUS des Gozis

O. latiusculus (LeOonte) - ON PQ NB NS PE - NF

Genus ORDOBREVIA Sanderson

O. nubifera (Fall) - BC

Genus PROMORESIA Sanderson

Key to North American species: Brown (1972)
P. elegans ll-eContet PQ
P. tardella (Fall) PQ NB NS - LB NF
subarctica (W.J. Brown)

Genus STENELMIS Dufour

Key to North American species: Brown (1972)
S. bicarinata LeConte - ON PQ - NS
S. concinna Sanderson PQ
S. crenata (Say) MB ON PQ NB NS PE - NF
sordida Motschulsky
S. mera Sanderson PQ NB NS
S. musgravei Sanderson PQ
S. quadrimaculata Horn - ON PQ
blatchleyi Musgrave
sulcata Blatchley
S. sandersoni Musgrave - ON PQ
tarsalis Sanderson
S. vittipennis Zimmermann - SK MB ON PQ

T?ibe Macronychini
Genus MACRONYCHUS P.W.J. Miiller
Key to North American species: Hinton (1940)
M. glabratus Say - ON PQ NB NS
lateralis Melsheimer

Genus ZAITZE\rIA Champion

Z. milleri H.P. Brown - BC
Z. parvula (Horn t - \'K - BC AB
columbiensis (Angell)

173
Artematopidae

Family PSEPHENIDAE water-penny beetles

L. LeSage

Psephenids form a small family of aquatic bee- The taxonomy of the North American
tles; onl.' three species have been recorded so far in psephenids is relatively well known, tha.nks to the
Canada. Their vernacular name refers to the works of Brown (1972, 1983b), Brown and Murvosh
unique round and flat shape of the body of the lar- (I974), and Brigham (1981).
vae, which live in riffles of creeks and rivers. The
adults o1' Ectopria spend the daytime in the foliage
of marginal vegetation; they are readily attracted to
light. Tlrose of Psephenus scurry over exposed rocks
in riflles during the day and hide at night. ON (3); PQ (2);NB (1); NS (2)

subfamily EUBRIINAE
Genus EC,fopRrALeconte
Eurea LeConLe
Key to Irforth American species: Brigham (1981)
E. leechi Brigham - ON
E. nervosa tMelsheimerr - ON PQ - NS
tarsalis LeConte
thorar:'ica (Ziegler)
tibiali)s LeConte

subfamily PSEPHENTNAE
Genus ps[]pHENUS Hardeman
Eurytr:.alpus LeConte
Fluuit:'ola DeKay
Key to liorth American species: Brown & Murvosh (1974)
P. herricki lDeKayr - ON PQ NB NS
lecontei (LeConte)
t re nto nen si s Zimmermann

Superfamily Elateroidea
Family ARTEMA*TOPIDAE
Y. Bousquet

Thi-" family includes eight North American one found in North America. Many authors, includ-
species, four of which are currently known to occur ing Arnett (1968), have placed the macropogonines
in Canarla. Little is known about the bionomics of
within Dascillidae.
the Near:ctic species. The adults are usually found
on foliage; the larvae of some species have been
reared or.r mosses (Lawrence 1982q.).
Lawrence (1982a) recognized three subfamilies NT (1); BC (3); AB (1); MB (1); ON (2); PQ (2); NB
of Artematopidae, Macropogoninae being the only (1)

Subfamily MACROPO GONINAE

Genus MACROPOC'ON Motschulsky
Key to North American species: Hopping (1936)
M. piceus l,eConte NT BC AB - MB ON PQ NB
dubiusW.J. Btown
rufipes l{orn
M. testaceipennis Motschulsky - BC

174
 Elateridae

cribricollis W.J. Brown

rubricollis Pic

Genus EURYPOGON Motschulsky

Dasypogon LeConte
E. californicus Horn BC
E. niger (Melsheimer) oN pe
horrisi (Westwood)

Family ELATERIDAE click beetles

Y. Bousquet

This family is well known for the ability of the The classifrcation of the family followed here is
adults to click when lying on their back. It includes Hyslop's (1917), with slight modifrcation. His classifr-
369 species and subspecies in Canada and Alaska. cation is preferred Stibick's (1979), as it is used more
The adults occur on foliage and flowers, under bark, widely. There is no recent taxonomic revision of the
and in rotting wood; the larvae are found in leaf litter, North American species. Most species that occur in
soil, moss, and rotting wood. Larvae of most species the areas covered in this checklist could be identifred
living in leaf litter and rotting wood are predaceous; using Dietrich (1945), Brooks (1961), Lane and
those occurring in the soil feed on roots, and some are Lancaster (1971), and the taxonomic reYisions listed.
very injurious to crops. Adults of some species appar-
ently do not feed, whereas those ofother species prob- AK (53); YK (32); NT (49); BC (19a); AB (131); SK
ably feed on tender plant tissues, such as leaves and (88);MB (99); ON (175);PQ (165);NB (81); NS (87);
flower buds. PE (21);LB (25);NF (46) ' H (6);I (5)

Subfamily PYROPHORINAE
Tlibe Pyrophorini
Genus LACON Castelnau
Aulacon Arnett
Diphyaulon Arr:.ett
Lepidotus Stephens
Zalepia Arnett
Key to North American species: Arnett (1952)
L. auroratus (Say)
L. avitus (Say)
L. discoideus (Weber)
_ -_ v_ yu oN
oN
ON
PQ
PQ
NB

pennatus (Fabricius)
L. impressicollis (Say) ON
lepturus (Say)
senilis (Germar)
L. maculatus (LeConte) ON
L. marmoratus (Fabricius) oN PQ
L. rorulentus (LeConte) -ec--
L. sparsus (Candbze) -BC
Genus DANOSOMA Thompson
Key to North American species: Arnett (1952)
D. brevicorne (LeConte) AK YK NT BC AB SK MB ON PQ NB
cauicolle (LeConte)
profusutn (Candbze)
uetusturn (Walker)
D. obtectum (Say) YK NT BC AB SK MB ON PQ NS

Genus ALAUS Eschscholtz

A. melanops LeConte BC
naja Canddze
A. myops (Fabriciust oN PQ
canadensis Canddze
A. oculatus (Linn6) oN PQ
luscus (Fabricius)

]-75
Elateridae

Genus DR.A.STERIUS Eschscholtz

D. debilis L,eConte AK YK BC AB SK MB ON PQ NB NS
incongruus Fall

Genus AEC'LUS Eschscholtz

A.
A.
amabilis (LeConte)
mellillus comis (LeConte) NT BC AB
-oN
A. mellillus marginicollis (Horn) sx MB
A. mellillus mellillus (Say)
dorsali.s (Say)
-oN PQ

Genus COIT|ODERUS Eschscholtz

C. auritus (Herbst) AB SK MB ON PQ NS
caseyi'.Leng)
crassicollis (Melsheimer)
dilectus (Say)
C. bellus (Say) ON
piclzs lSteinheil)
uariegutus (Steinheil)

Tribe Pityobini
Genus PIT)|OBIUS LeConte
P. anguinusi LeConte AR SK MB ON PQ NB NS
billing.si Bland

Tlibe Athoini
Genus LIM,INIUS Eschscholtz
L. aeger Leilonte BC MB ON PQ NB NS PE LB NF
hnulli ?all
L. agonus (fiay) ON PQ
L. anceps L:Conte SK MB ON PQ NB
L. aurifer LeConte _MBONPQ
L. auripilis (Say) ON PQ
pubico;Iis (LeConte)
L. basillaris (Say) ON PQ
semiaeneus LeConte
L. bicolor Van Dyke _BC
L. cal ifornicus t Mannerheimj _BCABSK MB
hispidus LeConte
occider,.talis Candbze
L. canus LeConte BC AB SK
discoiaeus LeConte
subcosi atus Motschulsky
L. confusus LeConte oN PQ NS
aenesc(:ns LeConte
L. crotchi Horn BC
L. ectypus (Say) SK mn-
L. griseus (l']alisot de Beauvois)
hirticot lis Melsheimer
-oN
propexits Canddze
L. humidus Lane _BC
L. infuscatus Motschulsky _BCAB
L. lanei Van Dyke _BCABSKMB
L. lecontei Lane _BCABSK
consimilis Walker
nitidicollis LeConte
L. maculicollis Motschulsky _BC
discicollis Canddze
_BC
L. nitidulus Horn
L. pectoralir; LeConte
L. pictus Van Dyke
L. pilosulus Canddze
AK YK NT BC
_BC y:"
AB
MB ON PQNB_ LB NT

pilosus LeConte
-BC
L. plebejus {Say) oN PQ
m e ta IIe scens Melsheimer

176
 Elateridae

L. quercinus (Sayt
L. rudis W.J. Brown
L. rufihumeralis Lane ECAB_
L. seminudus Van Dyke BC
L. snakensis Lane BC_-
L. stigma tHerbst)
armus (Say)
L. subauratus LeConte BC
L. ursinus Van Dyke AB SK MB
L. venabelsi Wickham BC

Genus ELATHOUS Reitter

E. californicus Van Dyke BC
E. discalceatus (Say) ON PQ NS
ober ndor ffe r i ( Schwarz )
E. nebulosus (Van Dyke) BC

Genus ATHOUS Eschscholtz

Crepidophorus Mulsant & Guillebean
Key to eastern North American species: Becker ( 1974)
Key to North American species: Becker (1979)
A. acanthus (Say) ON PQ NS
carolinus Van Dyke
flauangularis Hotrr
maculicollis LeConte
A. brightwelli {Kirby} ON PQ NB NS
arcticollis Melsheimer
oblong icoll is Melsheimer
A. campyloides Newmant -PQ NB NS PE
affinis Coupet
A. cucullatus {Say) ON PQ _NS
hypoleucus Melsheimer
p rocer icoll is Melsheimer
stripatus Melsheimer
A. lossularis (LeConte) -oNPQ
A. imitans Fall BC
A. neacanthus Becker PQ
A. nigropilis Motschulsky BC AB
montanus LeConte
nugalis Fall
orophilus Harold
speculifer FalI
A. orvus Becker _ONPQ NB NST_ _
A. posticus (Melsheimert -oNPQ
erebusYanDyke
A. productus (Randall) SK MB ON PQ NF
bipunctatus Provancher
A. rufrfrons (Randallt MB ON PQ NB NS NF
lengiDtry
reflexus LeConte
A. rufrventris rectithorax Reitter BC
recticollis Fall
A. rufrventris rufiventris AK BC AB
(Eschscholtz)
fer r ug inos us Eschscholtz
maritimus Van Dyke
A. scapularis (Say)
A. scissus LeConte BC ___9*PQ_____
ingens Fall
A. sierrae varius Lane AK BC
A. vittiger LeConte BC

Genus HARMINIUS Fairmaire

H. triundulatus (Mannerheim) AK YK BC SK ON PQ

Genus MICRATHOUS Lane

M. brevis (Van Dyke) BC

Genus DENTICOLLIS Piller & Mitterpacher

Key to North American species: Becker (1952)

177
Elateridae

D. denticor nis { Kirby t NT BC AB SK MB ON PQ NB NS NF

flauinrcus (Melsheimer)
D. varians (Germar) AK YK NT BC
uariabilis (Eschscholtz)

Genus BERNINELSONIUS Leseigneur

Key to s;cecies: Stibick (1976)
B. hyperboreus (Gyllenhal)* AK NT
plona,:us (Eschscholtz)

Genus LIGMARGUS Stibick

Key to N.orth American species: Stibick (1976)
L. funebris (Canddze) AK YK NT BC AB
L. lecontei (Leng)
planaius (LeConte)
-oN PQ NS

Genus ASCOLIOCERUS M6quignon

Scolio :erus Motschulsky
Key to North American species: Stibick (1978)
A. sanborni(Horn)x AK YK NT AB MB PQ
barba:tus (J.R. Sahlberg)

Genus MAI,GAIOSTUS Stibick

Key to North American species: Stibick (1978)
(Subgenus MARGAIOSTUS s. str.)
M. glacialirr (Van Dyke) AK BC AB
M. grandic'rllis (Lecontel oN PQ

Genus HYI'OLITHUS Eschscholtz

C ry pto hy pnus Eschscholtz
Key to n*orth American species: Stibick (1978)
H. littoralir; Eschscholtzx AK
Genus HYPNOIDUS Dillwyn
Key to North American species: Stibick (1978)
H. abbreviatus (Say) AK YK BC AB SK MB ON PQ NB NS PE NF
silaceipes (Germar)
H. bicolor ( Eschscholtz)* AK YK NT BC AB SK MB ON PQ NB NS NF
fallax (Mamer}rein)
lacustris (LeConte)
limbatus (Mannerheim)
picescens (LeConte)
pumilits (Dejean)
scar ifi c at u s (Mannerheim)
uestitus (Mannerheim)
H. impress collis (Mannerheim) AK YK NT BC AB MB ON PQ LB NF
H. leei Stibick _AB
H. nocturnus (Eschscholtz) AK YK_BCAB
H. rivularius (Gyllenhal)* AK YKNTBC_SK MB PQ LB NF
H. squalidr.s tLeContet BC AB
Genus OEDOSTETHUS LeConte
O. femoralis LeConte AB SK MB ON PQ NB

Genus NEGASTRIUS Thomson

N. caurinus lHorn; BC
_ABSK
N.
N.
N.
N.
choris (Sayt
delumbi s tHorn)
dispersus (Horn)
dubius ( Horn)
BC
_ABSK
MB ON PQ
yI' PE

MB
N. exiguus (Randallt -oN
guttatulus (Melsheimer)
N. extricatus (Fall) AK YK NT BC
N. gentilis rLeConte) BC AB SK
N. manki (Fall) BC AB
N. melsheirneri tHorn) _PQ NB NS
N. musculus (Eschscholtz) AK BC

178
 Elateridae

N. obliquatulus (Melsheimer) ON NS
N. ornatus (LeConte) BC
colon (Horn)
moerens (Hotrr)
N. pectoralis (Say) BC AB SK MB ON PQ NS
futilis (LeCofte)
inops (LeConte)
N. restrictul us (Mannerheim.) AK YK NT PQ
N. striatulus (LeConte) AK BC AB _ONPQNBNS
N. tumescens LeConte AK YK NT BC AB SK MB ON PQ NB NS

Genus CTENICERA Latreille

Corymbites Latreille
C. aeripennis (Kirby) AK YK NT BC AB SK
tincta (LeConte)
C. angularis (LeConte) BC
C. angusticollis (Mannerheim) AK BC
fraterna (LeConte)
spectabilis (Mannerheim)
C. appressa (Randall) ON PQ NB NF
mirifica LeConte
C. appropinquans (Randall) SK MB ON PQ NB NS PE NF
C. arata (LeConte) -YK NT BC AB SK MB ON PQ NB NS NF
C. atropurpurea (Melsheimer) PQ NF
C. bipunctata (W.J, Brown) _BCAB
C. bombycina (Germar) _BCAB
C. callida (W.J. Brown) _BC
C. carbo LeConte _BC
C. caricina (Germar) AK _BC
C. comes (W.J. Brown) -BC
C. confluens (Gebler) AK
C. conjungens (LeOonte) _BC
C. consors (W.J. Brown) AB SK MB
C. crestonensis (W.J. Brown) _BCAB
C. cylindriformis (Herbst) ON PQ NB NS PE
appressifrons (Say)
breuicornis (Say)
p ar al lel ip ip eda ( Germar)
submetallica (Germar)
C. destructor (W.J. Brown) AB SK MB
C. diversicolor (Eschscholtz)
C. dorothyae Knull PQ
C. elongaticollis (Hamilton) PQ NB NS--
C. fallax (Say) ON PQ NS
C. falsifrca (LeConte) ON PQ NB NS NF
C. fulvipes (Bland) ON PQ NS-_
C. funerea (W.J. Brown)
C. furtiva (LeConte)
monticola Horn
C. glauca (Germar) SK
simili s sima ( Motschulsky)
C. hamata (Say) ON PQ NB NS
C. hieroglyphica (Say) _MB ON PQ NB NS
C. hoppingi (Van Dyke) AI{ YK Nr sc AB SK MB ON PQ -?
C. inflata (Say) ON PQ
C. insidiosa (LeConte) ON PQ NB NS
C. kendalli Kirby AK YK NT BC AB SK MB ON PQ NB _NF
anchorago (Randall)
C. laricis (W.J.Brown) _BC AB
C. lateralis (LeConte) _BC
q uadr iu ittat a (Walker)
uittata (Walker)
C. limoniiformis (Horn) MB
C. lobata (Eschscholtz) AK NT TSK
telum (LeConLe)
C. lutescens (FaIl) _BC AB SK
C. maura (LeConte) _BC
C. mediana (Germar) NT AB SK ftfS ON pe NB NS
ubid.ipennis (LeConte)
r
C. mendax (LeConte) AK YK AB p(l
BC SK -oN NF

179
Elateridae

C. moerens. tLeConte) _BC

C. monLanrr (W.J. Brownl _BCAB
C. morula rl,eConte) AK YK NT BC AB SK MB ON PQ LB
brunnipes (Bland)
C. nebraskensis t Bland) _BCAB
C. nigricans tFall t _BC
C. nigricollis (Bland) AK _BCAB -nneoNpe-
C. nitidula (LeConte) AK NT BC AB SK MB ON PQ NB NTQ DF LB NF
C. ochreipennis (LeOonte) AK YK NT BC AB LB NF
C. opacula {Leconte) _BC :"yuoNPQ_
serricornis (Mannerheim)
C. propola columbiana (W.J. Brown) AK YK BC AB
C. propola propola (LeConte) NT BC AB SK MB ON PQ NB NS PE
furcifer (LeConte)
C. protracta { Leconte) BC
C. pruininir tHornt BC
no:rlo iHyslop)
pruinosa (Schwarz)
pruinosula (Schwarz)
C. pudica ( W.J. Brown ) _BCAB
C. pulchra tLeconte) NT BC AB SK MB ON PQ NB NS PE
festiua (LeConte)
C. pygmaert tVan Dyket NT BC AB SK MB ON PQ NB NS NF
C. pyrrhos {Herbst}
aequa,',is (Melsheimer)
-oN
pyrrhica (Haldeman)
uagralLs (Melsheimer)
C. respleniens aeraria (Randail) ? NT? AB SK MB ON PQ NB NS LB NF
junAi , Schenkling)
racinei (Chevrolal)
uiridi.. (Germar)
C. resplencens resplendens AK NT BC AB SK
(Esr:hscholtz)
C. rotundirollis r Sayt ON
russicollis (Germar)
stictico (Germar)
C. rufopleuralis tFall t NB NS PE
C. rupestri r {Germarr _BC _yuoNPQ
C. sagitticc,llis (Eschscholtz) _BC
C. semimel allica (Walker) _BCAB
C.
C.
sexualis tW.J. Brown.)
silvatica Van Dyke _BCAB
AB ix- - -
C. sjaelandica (O.F. MtiLiler)t oN PQ NU
uiridis (Say')
C. spinosa ' LeContet
C. spf enderrs lZieglert
C. stricklar:Ldi (W.J. Brown) AK
_ _ _ ABSXMBSNiSNBNS_
_NTBCABSKMBON
_
NF

C. suckleyi suckleyi (LeConte) BC--laeoNpo----

C. sulcicoll:s tSay)
parallela (Say)
C. tarsalis Melsheimer; -oNPQ NB NS
C. tristis tCandbzeJ BC
C. triundulata (Randall) AK _NT BC AB SK MB ON PQ NB NS PE LB NF
C. uliginostt tVan Dyket BC
C. umbrico a (Eschscholtz) AK BC AB
rzdis (Motschulsky)
C. umbripennis (LeConte) BC AB
gracilior (LeConte)
n ub il ipennis (LeConte)
C. vernalis (Hentz) ON PQ NS
C. vidua tS'.J. Brown) BC AB
C. volitans (Eschscholtz) AK BC
uaria (W .J. Brown)
C. vulnerata (LeConte) _PQ NS
C. watsoni W.J. Brown) _PQ LB
C. weidtii t.\ngellr nc--
Genus OXI'GONUS LeConte
Key to North American species: Roache (1963)
O. montanus Schaeffer PQ NB NS

180
 Elateridae

O. obesus (Say) MB ON PQ
acutipennis (Randall)

Genus EANUS LeConte

P ar anom u s Kiesenwetter
E. albertanus W.J. Brown _BCAB
E. decoratus (Mannerheim) AK YK NT BC AB SK MB ON PQ LB NF
paruicollis (Mannerheim)
uagus (LeConte)
E. estriatus (LeConte) AK YK NT BC AB MB ON PQ NB NS LB NF
subarcticus W.J. Brown
E. granicollis (Van Dyke) _BC
E. maculipennis LeConte ON PQ NB LB NF
pictus (Canddze)
E. striatipennis W.J. Brown AK BC

Genus HEMICREPIDIUS Germar

Asaphes l\trby
H. bilobatus (Say) ON
c au ifron s (Melsheimer)

con s ent ane u s (LeOonte)

mel ano p ht hahn us (Melsheimer)
planatus (LeOonte)
tener (LeConte)
H. brevicollis (Candbze) MB ON PQ NB NS
H. carbonatus (LeConte) BC AB SK MB
coracinus Candbze
H. dilaticollis Motschulsky BC
H. hemipodus (Say) ON PQ
aeneolus (Melsheimer)
oereus (Melsheimer)
corporosus (Germar)
decoloratus (Say)
H. memnonius (Herbst) AB SK MB ON PQ NS PE
baridius (Say)
ruficornis (Kirby)
thomasi Germar
H. montanus Lane BC
H. morio (LeConte) BC
uernus Canddze
H. pallidipennis (Mannerheim) AK BC
oregonus (LeConte)
H. simplex (LeConte) _BC AB
H. tumescens (LeConte) _BC

Tlibe Oestodini
Genus OESTODES LeConte
O. puncticollis Horn AB SK MB
O. tenuicollis (Randall) ON PQ NB NS PE
g r ac ilifor mis (RandalI)
obscuripes Candbze

Subfamily EI"ATERINAE
Tribe Elaterini
Key to North American species: Roache (1961)
Genus ELATER Linn6
Ludius Berthold
Steatoderus Dejean
E. abruptus Say oN PQ NS
coracinus (Germar)

Genus PARALLELOSTETHUS Schwarz

P. attenuatus (Say) ON
/uscus Castelnau

181
Elateridae

Genus SERICUS Eschcholtz

Atractooterus LeConte
Dolerosomus Motschulsky
Sericosomus Dejean
S. blaisdelli (Van Dyke) BC
S. honestus (Randall) -oN PQ NB
fusiforrzis (LeConte)
S. incongrurLs (LeConte) AK YK NT BC SK MB ON PQ NB NS LB NF
behren::i (Canddze)
S. silaceus (tjay) -oN PQ
debilis (LeConte)
flauipennis (Motschulsky)
umbraticus (LeConte)
S. viridanus (Say) ON PQ NS
sublucens (Randall)

Tlibe Agriotini
Genus GLY?HONYX Canddze
Key to Ncrth American species: Smith and Balsbaugh (1984)
G. testaceus (Melsheimer) ON

Genus AGRTOTES Eschscholtz

Pedetes l{trby
Key to N< rth American species: Becker (1956)
A. apicalis l,eConte BC
A. arcanus \V.J. Brown -o*
A. avulsus (.LeConte) -oN
NSpe--
A.
A.
collaris (LeConte)
criddlei Van Dyke SC AB
BC AB
ix -oN
MB
PQ
v
A. fer"rugineipennis (LeConte)
A. fucosus (],eConte) _AB SK MB ON pe NB NS--Nr
A. hoodi Bec:ker BC
A.
A.
isabellinus (Melsheimer)
limosus ( LeOonte) _NT BC AB SK
-on
MB ON
PQ
PQ NE Ns pn is Nr
A. Iineatus tLinn6X BC NS NF
A. mancus (Say) _AB SK MB ON PQ NB NS PE LB
striatutus Melsheimer
A. oblongicollis (Melsheimer)
A. obscurus (Linn6)t BC _9^ PQ
NF
A. opaculus (LeConte)
montartus LeConte
BC AB -lt
A. pubescens Melsheimer _MBONPQ NB
_NS
A. quebecensis W.J. Brown
A. sparsus LeConte --sc- _ yuoNPQ
A. sputator il-inn6)f NB NS
A. stabilis (l,eConte)
A. tardus W.J. Brown
BC
BC AB
SK MB ON
____
PQ
v
A. thevenetl,i Horn BC

Genus AGRIOTELLA W.J. Brown

A. bigeminata (Randall)
A. columbiana W.J. Brown BC
yu 9* PQ
YNS
A. occidentzrlis W.J. Brown BC A3

Genus DALOPIUS Eschscholtz

D. agnellus W.J. Brown
scTsx- Iu_
D. asellus W.J. Brown
D. brevicornis W.J. Brown
D. cognatus W.J. Brown
- - a*;:
oN NB
PQ NS
D. corvinus W.J. Brown BC
D. fucatus W.J. Brown
D. fuscipes W.J. Brown
D. gartrelli W.J. Brown
D. gentilis W.J. Brown
D. inordinar;us W.J. Brown
BC
BC
--scABSf-9*PQ
AB

AB
_PQ
-v
D. insolens W.J. Brown
D. insolitus W.J. Brown -_l"T--oNpe

r82
 Elateridae

D. insulanus W.J. Brown _BC

D. maritimus W.J. Brown _BC
D. mirabilis W.J. Brown ABSKMB_PQ
D. pallidus W.J. Brown _BCABSKMBONPQ NB \Tq DF _NF
D. parvulus W.J. Brown -- MB-
D. spretus W.J. Brown - -_BC ec - :" -
D. suspectus W.J. Brown
D. tristis W.J. Brown
D. vagus W.J. Brown
D. vernus W.J. Brown _ - -:"-:"ilB3Nia v NS

Tribe Ampedini
Genus AMPEDUS Dejean
A. antracinus (LeConte) -BC
A. apicatus (Say) AK_ NT BC AB SK nfS O* PQ NB NS PE - lrr
melanopygus Germar
A. areolatus (Say) -oN PQ
obliquus (Say)
scitulus Germar
A. atripennis (Horn) _BC
u iolaceipennis (Candbze)
_BC
A. behrensi (Horn)
A. brevis (Van Dyke)
A. collaris (Say)
T_ _BC AB ___
MB_PQ NS-
A. deletus (LeConte) AK NT BC
_BC
AB SK MB ON PQ NB NS is N"
A. dimidiatus (LeConte)
A. evansi W.J. Brown NT_ -s* MB ON PQ NB NS
A. fuscatus (Melsheimer)
A. fusculus (LeConte)
A. impolitus (Melsheimer)
AB SK
-oNPQ
MB ON
_ONPQ
PQ NB iey
A. laesus (LeConte)
Nr -oN NS- _NF
A. laurentinus W.J. Brown AK- sc AB_ MB ON
_ONPQ
PQ NS
A. linteus (Say)
lugubris Germar
A. luctuosus (LeConte)
A. melanotoides W.J. Brown
NT
YSK MB
-oN
ON PQ
PQ
NB NS_ LB NF

A. melinus (LeConte) -BC _SK MB NS

A. melsheimeri (Leng) ON PQ
hepaticus (Melsheimer)
A. militaris (Harris)
A. miniipennis (LeConte)
A. mixtus (Herbst)
A. moerens (LeConte)
_v Nr- AB- - SNFBNn-
AB SK MB ON PQ NB NS
NT BC AB
-PE ie
_NF
Ne

A. molestus (LeConte) -BCMBONPQNBNS

A. nigricans Germar AB SK MB ON PQ NB NS
A. nigricollis (Herbst) -oNPQ
A. nigrinus (Herbst)*
A. oblessus (Say)
sellatus (Leng)
AK YK NT BC AB SK MB ON PQ NB
AB SK MB ON PQ NB :"y
discoideus (Fabricius)
A. occidentalis Lane BC AB
bimaculatus (Van Dyke)
A. pedalis Germar AB SK MB ON PQ
rufipes (Palisot de Beauvois)
A. phelpsi (Horn) BC AB
hoppingi (Van Dyke)
A. phoenicopterus Germar BC
A. protervus (LeConte)
A. pullus Germar AK YK NT BC
---o*pe
AB SK MB ON PQ NB
NS
NS -N"
Iacustris (LeConte)
A. pusio Germar -oN
luteolus (LeConte)
BC AB LB NF
A. quebecensis W.J. Brown
A. rhodopus (LeConte) T NT
BC
SK MB PQ

A. rubricollis (Herbst) -oN PQ

socer (LeConte)

- - - - MB3N|8- N3- -
uerticinus (Palisot de Beauvois)
A. rubricus (Say)
A. sanguinipennis (Say) -

183
Elateridae

A. sayi (Let)onter - ON
A. semicinrtus rRandall) - ON PQ - NS
A. subtilis tleOonte) - BC AB SK MB ON PQ - NS
A. Dyker
uteanus (Van - SK
A. Dyket
varipilis (Van - \'K NT BC AB
columoianus W.J. Brown
A. vitiosus LeContet - ON PQ NB
Genus ME(IAPENTHES Kiesenwetter
Dolopt osomus Motschulsky
Key to Nrrth American species: Becker (19?1)
M. caprella tl-eContet AK BC
califor nicus W.J. Brown
M. gentner Lane - BC
M. nigriventris LeConte - BC
M. rogersi }{orn ON PQ NB
M. solitariL.s Fall PQ
M. stigmosus (LeConte) BC AB SK MB ON PQ NB
M. tartareus (Lecontej - BC

Tribe Physorhini
Genus ANC HASTUS LeConte
B r ac h2,' cre p i s LeConte
Crepiaotritus LeConte
Elatrh,.us Horn
A. anthrax Horn.) - BC
coruzs Van Dyke
militaris Candbze
A. cinereipt nnis tEschscholtzr - BC AB
hir s utul us Motschulsky
recedens LeConte
p ilifer t.: s Motschulsky
puberu.lus (Mannerheim)
regularis Motschulsky
ta nt il I t.t s ( Mannerheim)

Tribe Melanotini
Genus MEL,ANOTUS Eschscholtz
Cratonychus Dejean
Ctenon,ychus Stephens
Perimecus Stephens
Key to North American species: Quate and Thompson (1967)
M. americanus (Herbst) -oN
ang u st.a tu s (Erichson)
M. castanipes (Paykull)t MBONPQNBNSPE_NF
inaequ,zlis (LeConte)
paradaxus Melsheimer
scrobiccllis (LeConte)
texanu:; Canddze
M. communis (Gyllenhal) _ONPQ
pag(tnt,ts Canddze
M. corticinus (Say) rlNT
g I andieo lor Melsheimer
uetulus (Erichson)
M. decumar:Lus (Erichson) _ONPQ_NS
canadensis Candbze
cuneo.tus (LeConte)
incertus (LeConte)
M. depressus (Melsheimer) .rNT
diu ar c arinus
Blatchley
lixus Blatchley
p ar um p unc tatas (Melsheimer)
M. hyslopi \/an Zwaluwenburg - ON PQ
M. ignobilis Melsheimer
secretu.s (LeConte)
-oN
M. leonardi (LeConte) MB ON PQ

184
 Elateridae

M. longulus oregonensis (LeConte) BC

franciscanus Van Dyke
uariolatus LeConte
M. miscellus Quate -oN
M. morosus Canddze -oN PQ
debilis Blatc}:.ley
longulus (LeConte)
simulans Blatchley
M. prasinus Blatchiey
M. sagittarius (LeConte)
carinus Blatchley
_9* PQ

M. similis (Kirby) _SK MB ON PQ NB NS

flssilis (Say)
exuberans (LeConte)
oc hr acei p ennis (Melsheimer)
henoidal is (Melsheimer)
sp
M. trapezoideus (LeConte) ON

Subfamily CARDIOPHORINAE
Genus CARDIOPHORUS Eschscholtz
C. acutus Lanchester _AB
C. alpestris Lanchester BC
C. brunnipennis Lanchester BC
C. cardisce (Say) _AB SK MB ON PQ NB NS
dejeanii LeConte
C. communius Lanchester BC
C. convexulus LeOonte --oNpeNe----
ON PQ
C. convexus (Say)
amictus Melsheimer
insulsus Canddze
C. edwardsi Horn _BC
C. fenestratus LeConte NT BC AB
larquini Candbze
C. gagates Erichson MB ON PQ NB NS
laeuicollis Erichson
C. kooskooskiensis Lanchester BC AB
C. Iatiusculus Eschscholtz BC
C. montanus Bland _AB
C. nevadensis Blanchard BC
C. plebejus Lanchester BC
C. plexus Lanchester BC
C. pratensus Lanchester _AB
C. propinquus Lanchester BC
C. pubescens Blanchard BC AB SK MB
C. silvanus Lanchester _AB
C. sodalis Lanchester _AB
C. tenebrosus LeConte BC AB SK MB
fuluipes LeConte
C. tumidicollis LeConte BC AB
C. tutus Lanchester BC AB
Genus HORISTONOTUS Canddze
H. curiatus (Say) ON
areolatus (Erichson)

185
Eucnemidae:

FamilyTHROSCIDAE throscidbeetles
Y. Bousquet

Throscidae includes about 30 North American (1979) treated the lissomines as a subfamily of
species, of which nine have been reported in Eucnemidae; Lawrence (1982a) listed them in
Canada. The adults, which look like small, stout Elateridae. I have followed Yensen (1975) in con-
elaterids, are usually found on foliage and flowers. sidering the lissomines as a subfamily of
The larvae occur in litter, in decaying wood, and Throscidae. Yensen (1975) provided a key to the
under bark. extant genera of the family and revised t;he genus
Few authors agree on the position of the lis- Trixagus.
somines. Crowson (1961) included Drapetes in
Elateridae and Lissomus in Throscidae; Burakowski NT (2); BC (6); AB (1); MB (1); ON (5); PQ (5); NB
(1975) el:vated the group to family level; Lohse (2);NS (2);NF (1)

Subfamily LISSOMINAE
Genus DRAIIETES Dejean
D. gerninatus (Say) oN PQ
ame r icc,nus ( Gerstaecker)
basalis (Randal1)
exstriatus (Say)

Subfamily THROSCINAE
Genus TRD(AGUS Kugelann
Throsct.Ls Latreille
Key to North American species: Yensen (1975)
T. carinicoll s tSchaeffer.t _BC AB ON PQ NB NF
T. chevrolat {Bonvouloirj _BC ON PQ NS
paruulu.s (LeConte)
sejunctus (Hotn)
T. mendax (.lorn) NT BC
T. sericeus ( LeConte; NT BC
debilis t}{orn)

Genus AULONOTHROSCUS Horn

A. constrictor (Say) MB ON PQ NB NS
A. punctatus (Bonvouloirt
A. validus (LeConte) _BC _9* PQ

Genus PACI]IOPUS LeConte

P. hornii (LeConte) BC
fuchsi Casey

Family EUCNEMIDAE false click beetles

Y. Bousquet

Thirty-two species of eucnemids have been many changes and additions to the Canad.i.an fauna
reported in Canada and Alaska. The adults occur can be expected. Horn's revision (1886) is the latest
mainly on vegetation and under bark but are rather one for the North American species.
infrequently collected. The larvae bore into various
hardwoods that are usually rotten. Like elaterids,
adults have the clicking mechanism, but few species
have been actually observed to click.
The taxonomy of the North American species of AK (1); BC (13); AB (2); SK (1); MB (3); ON (18); PQ
Eucnemidae has been poorly studied, and therefore (20);NB (5);NS (3);NF (1)

186
 Eucnemidae

Subfamily EUCNEMINAE
Genus STETHON LeOonte
S. pectorosus LeConte - ON PQ

Genus DELTOMETOPUS Bonvouloir

D. amoenicornis tSay) - ON PQ NB NS
clypeatus (Say)
ereptus Bonvouloir
D. rufipes (Melsheimert

Genus DROMAEOLUS Kiesenwetter

D. basalis (LeContet - BC
D. californicus Bonvouloir - BC
D. cylindricollis (Sayt - ON PQ
D. harringtoni Horn MB ON PQ NB
D. nitens Horn - BC

Genus FORNAX Castelnau

Isarl hrus LeConte
Onichodon Newman
F. badius (Melsheimer.t - ON PQ
basalis Hotn
nouitius (Bonvouloir)
F. bicolor (Melsheimer) PQ
luridus Chevrolat
F. calcealus tSay) - ON PQ
spretus (LeConte)
F. canadensis W.J. Brown - ON PQ NB NS
F. hornii Bonvouloir - ON PQ
F. orchesides (Newman) - ON PQ
Genus PHAENOCERUS Bonvouloir
P. americanus Horn - BC

Subfamily DIRIIAGINAE
Key to genera: Cobos (1964)

Genus DIRHAGUS Latreille

Microrhagus Dejean
D. pectinatus (LeConte) - BC - ON PQ NB
D. subsinuatus (LeConte) MB ON PQ
imperfectus (LeConte)
oblilus (Bonvouloir)
D. triangularis tSayt - ON PQ

Genus ENTOMOPHTTIALMUS Bonvouloir

E. rufiolus (LeConte) - ON PQ
pallens Bonvouloir

Genus SARPEDON Bonvouloir

S. scabrosus Bonvouloir - BC - ON PQ

Genus RHAGOMICRUS Fleutiaux

R. humeralis { Say) PQ
meticulos us (Bonvouloir)

Subfamily MELASINAE
Genus ISORHIPIS Boisduval & Lacordairei
I. obliqua (Say) - BC - ON PQ NB NS

Genus MELASIS Olivier

M. rufipennis Horn - BC
M. tsugae Hopping - BC

187
Lycidae

Genus EPIPHANIS Eschscholtz

E. cornutus Eschscholtz AK BC - SK - ON PQ NB - NF
cristatus LeConte

Genus SCHI ZOPHILUS Bonvouloir

S. subrufus Randall,l
sirnplet (LeConte)
trilobatus Bonvouloir

Genus HYLI.S des Gozis

H. frontosus tSayt
canaliculatus (LeConte)
H. terminalrs (Leconte, - BC AB SK - ON PQ

Genus XYLOPHILUS Mannerheim

Xylobius Latreille
Xyloecus Dejean
X. cylindriformis lHornt - BC

Genus NEMATODES Berthold

Hypocozlus Dejean
N. atropos {l;ay} MB
N. penetran; (Leconte) - ON PQ

Genus ANE,,ASTES Kirby

A. druryi Kirby - BC AB
brunne.rc (Latreille)
aroszs (Say)
latreille'i LeConte

Superfamily C antharoidea
Family LYCIDAE net-winged beetles
J. McNamara

Twent;y-nine species of Lycidae are recorded in tions; his paper of 1949 includes a key to the
Canada and Alaska. The adults occur on leaves, on Nearctic tribes.
flowers, or under bark and are thought to be preda-
ceous. The larvae are predaceous, and those of
most species are found under bark.
Greer, has revised the taxonomy of the North AK (3); yK (1); NT (1); BC (8); AB (5); SK (a); MB
American species of Lycidae in a series of publica- (8); ON (25); PQ (19);NB (5);NS (5); LB (1); NF (1)

Subfamily LYCINAE
Tlibe Lygistopterini
Key to North American species: Green (1950)
Genus CAI-OCHROMUS Gu6rin-M6neville
C. dimidiatr,is (LeConte) -BC
C. fervens LeConte
C. perfacetus (Say)
substritttus (LeConte)
---:"Tsx-o*pe-----
Tlibe Platerodini
Key to North American species: Green (1951, 1953)

Genus DICTYOPTERUS Mulsant

D. aurora (Flerbst) AK YK NT BC AB _ MB ON PQ NB NS _ LB NF
coccinatus (Say)
nigripes (Schaeffer)

188
 Lycidae

D. hamatus Mannerheim AK BC
D. simplicipes Mannerheim AK BC AB
loelzs Motschulsky
D. thoracicus (Randall) BC AB SK MB ON PQ
praefectus (Newman)

Genus EROS Newman

E. humeralis (Fabricius) ON PQ
incestus LeConte
obliquus (.Say)

Genus PLATYCIS Thomson

Erotid,es Waterhouse
P. sculptilis (Say) ON PQ
axill aris (Melsheimer)
oblira (Newman)

Genus LOPHEROS LeConte

L. crenatus (Germar) MB ON PQ
cruciatus (Randall)
L. fraternus (Randall) -oNPQ
Genus EROPTERUS Green
E. arculus Green _ONPQ
E. trilineatus (Melsheimer) _ONPQ
Genus PLATEROS Bourgeois
Neoceletes Leng & Shoemaker
P. batillifer Green -oN
P. bispiculatus Green -oNPQ
P. borealis Green -oN
P. centralis Green -oN
P.
P.
fl avoscutellatus Blatchley
floralis (Melsheimer)
-oNPQ NS

P. lictor (Newman)
californicus Van Dyke
- ecABsxmn3Ni3 NB NS

columbiensis W.J. Brown

crateracollis (Leng & Shoemaker)
P. peregrinus Green _BC oN PQ
P. subfurcatus Green ON
P. timidus (LeConte) ON
P. volatus Green ON PQ NB

Tlibe Calopterini
Key to North American species: Green (1952)
Genus CAI-OPTERON Gu6rin-M6neville
Diagrapha Newman
yuSN
C. discrepans (Newman)
C. reticulatum (Fabricius)
affinis (LeConte)
- - - PQ

retiferum LeConte
typicum (Newman)
C. terminale (Say) MB ON PQ
apicale (LeConte)
diuisum (Newman)
dorsale (Newman)
duplicatum (Haldeman)

Genus CAXNIA Newman

Caeniella Cockerell
C. dimidiata (Fabricius) MB ON PQ NB NS
scapularis Newman

Genus CELETES Newman

C. basalis LeConte SK MB ON PQ NB NS
mystacina LeConte
tabida LeConte

189
Lampyrrida e

FamiIyPHENGODIDAE glowworrns
J. McNq.mara

Only one species of Phengodidae occurs in debris on the ground. The males are al;tracted to
Canada. The larvae of most species are lumines- light.
cent, hertce the vernacular name glowworms. The Wittmer .1975) revised the North American
larvae on soft-bodied insects and other small
fe,,ed species of the genus Phengodes, the only genus
organisrls. The females are larviform but have occurring in Canada.
compound, instead of simple, eyes. Adults and lar-
vae occur in leaf litter and under logs, bark, and oN (1)

Genus PHIINGODES Illiger

Key to North American species: Wittmer (1975)
(Subgenus PHENGODES s. str.)
P. plumosa tOliviert ON

Familv LAMPYRIDAE fireflies

J. McNamara

Twenty-nine species of Lampyridae are slugs. The feeding habits of the adults are not well
recorded in Canada. Members of this family are known.
fairly well known because of their ability to produce Many genera of lampyrids have been revised in
flashing light. Species may be recognized in flight the past few decades, but a thorough revision ofthe
by the color of the light as well as by the number family would be very useful. Green (1948) pub-
and the length of flashes. This flashing behavior lished a revision of the Nearctic genera.
enables l;he sexes to frnd each other. All larvae and
most adults are luminescent. The larvae are preda- xT (3); BC (9);AB (3); SK (5); MB (12); ON (19); PQ
ceous and feed on other insects, larvae. snails, and (14);NB (8);NS (10);PE (1);NF (3) ' I (1)

Subfamily LAMPYRINAE
Tribe Cratomorphini
Genus PYI|ACTOMENA Melsheimer
Lecontea E. Olivier
Key to North American species: Green (1957)
P. angulata (Say) NT SK MB ON PQ NB NS
flauoc:,.ncta LeConte
P. borealis (Randall) AB SK MB ON PQ NB NS
P. dispersa Green NT SK MB
P. linearis LeConte MB ON PQ
P. lucifera Melsheimer ON
P. sinuata Green MB

T?ibe Photinini
Genus PHOTINUS Castelnau
Key to N'orth American species Green (1956)
P. aquilonius Lloyd MB ON PQ_NS
P. ardens LeConte MB ON PQ NB NS NF
taedif'zr LeConte
P. consimi]is Green -oN
P. ignitus llall -oN
P. indictus (LeConte) -oN
ablucens Fall
P. marginellus LeConte -oN PQ
castus LeCor,:te

190
 Lampyridae

P. obscurellus LeConte MB ON PQ
frigidus Olivier
P. punctulatus LeConte ON
P. scintillans (Sayt ON

Genus PHAUSIS LeConte

Key to North American species: Fender (1966)
P. nigra Hopping BC
P. rhombica Fender BC AB-
Genus ELLYCHNIA LeConte
Key to western North American species: Fender (1970)
(Subgenus ELLYCHNIA s. str.)
E. cornrsca (Linn6)
E. facula LeConle
-BC
_BC y" oN PQ NB NS NF

E. greeni Fender _BC

E. hatchi Fender _BC
E. lacustris LeConte _BC
Genus PYROPYGA Motschul sky
Keyto species: Green (1961)
P. decipiens (Har-ris)
P. nigricans (Say)
californica Motschulsky
NrscABixnrnSNiSNENS - -- -
californica (Gorham)
fene stralis (Melsheimer)
reuersus (Gemminger)

Genus LUCIDOTA Castelnau

RileyaE. Oliviet
L. atra (Olivier) MB ON PQ NB NS
laticor nis (Fabricius)
morio (Melsheimer)
tardaLeConte

Genus PHOSPHAENUS Castelnau

P. hemipterus (Fourcroy)f PQ NS

Subfamily ANIYDETINAE
Genus POLLACLASIS Newman
Polyclasis LeConte
P. bifaria (Say) ON PQ
oualus Newman

Subfamily PHOTURINAE
Genus PHOTURIS Dejean
Py r ec tosoma Motschulsky
Key to North American species: Barber (1951)
P. fairchildi Barber NS
P. pennsylvanica (DeGeer) BC_SKMBON PQ NB NS PE NF
marginata (Panzer)
u ittiger a (Motschulsky)

191
Cantharid,re

Family CANTHARIDAE soldier beetles

J. McNamqra

Cantharidae includes 126 species and subspecies Some of the genera of Cantharidae are poorly
in Canarla and Alaska. The adults are common on known taxonomically, including the large genus
goldenrc,d, milkweed, and hydrangea, where they Podabrus, and are in need ofrevision. Fender (1962)
feed on nectar and pollen. Most larvae are carnivo- provides keys to the species of the Pacifrc Northwest.
rous, gererally preying on other soft-bodied insects; a
few are cmnivorous, feeding on plants such as wheat, aK (15); vK (8); NT (13); BC (57); AB (20); SK (9);
potatoes. celery, and other plants. They occur mostly MB (22); ON (a8); PQ (48); NB (22); NS (rl5); PE (1);
beneath bark and debris. LB (5); NF (16) . I(2)

Subfamily Cantharinae
Tlibe Podabrini
Genus PODABRUS Westwood
Brachynotus I\irl:y
Malthacus l\trby
Key to Irorth American species Fa]l (1927)
P. alexand,rri Fender _AB
P. brevicol,is Fall ON
P. cavicolli; cavicollis LeConte BC
P.
P.
P.
P.
conspiratus Fall
deceptus W.J. Brown
diadema (Fabricius)
excursur; excursus Fall
AK
AK
vy BC
BC
BC AB
MB
oN PQ NB NS l*"
P. extremus LeConte AK \.K NT BCAB-MB ON LB
P. falli Hop,ping BC AB
P. fissilis Fall AK BC AB
P. flavicollis LeConte ON
P. frater LeConte ON
q uadr,.collis ( Motschul sky)
P. frosti Fender ON _NB
P. fumiganus Green PQ NB
P. furtivus Fall _BC
P. heteronl'chus Fall
P. instabilis Fall
P. intrusus Green
TY NT BC
_ IJU
AB SK MB ON

ON
PQ

PQ NB NS _NF
P. laevicoil s (Kirby) AK YK Nt sc AB SK MB ON PQ NB LB NF
P. limbellur LeConte PQ NB
P. lutosus LeConte _BC
P. lygarius Fender _BC
P. melvillei Fender _BC
P. modestus (Say) ON pe-Ns -Ne
P. nothoides LeConte MB ON PQ
P. obscurevittatus Fall _YK NT BC AB ix MB
P. pattoni LeConte PQ NB
P. perplexus W.J. Brown AK NT MB
P. piniphilus (Eschscholtz) AK NT BC AB MB PQ
P. planulus Green
P. probus FaIl
P. protensu s LeConte
MB ON

ON
PQ_NS
PQ-_ _Iu
P. pruinosts atrocervicus Fender _BC
P. pruinosr;.s diversipes Fall _BC AB
P. puberulus LeOonte MB ON PQ NB NS LB NF
P. punctatuLs LeConte PQ NB NS
P. puncticollis (Kirby) PQ NB
marginellus LeConte
P. punctula.tus LeConte ON PQ
P. pustulatus Fender _BC
P. pygmaeus Green PQ
P. rossi Fender _BC
P. rugosulus LeConte ON PQ NB NS
p ar u icol lis (Motschulsky)
P. scaber LeConte

192
 Cantharidae

P. secretus W.J. Brown PQ

P. sericatus (Mannerheim) AK
P. simplex Couper na" o* PQ
IU
NS
P. smithi Fender _BC
P. tetragonoderus Fall AK_ _BCAB
P. tomentosus (Say)
cinereipennis Motschulsky
-oN
rufiolus (Melsheimer)
P. tricostatus (Say) ON PQ
atripes (Motschulsky)
benneti (Kirby)

Tlibe Cantharini
Genus ANCISTROI.JYCHA Markel
Key to North American species: McKey-Fender (1950)
A. bilineata tSayt oNPQ-NS
A. dentigera (LeContet oN PQ
A. neglecta tFall r oN PQ
Genus CANTIIARIS Linn6
C y r tomo p tila Motschulsky
Telephorus Schaeffer
C. alticola (LeConte) BC
C. aneba borealis McKey-Fender NT BC AB SK MB
C. beta Fender BC
C. consors (LeConte) BC
tibialis (LeConte)
tibiella (Gemminger)
C. curtisi (Kirby) - YK NT BC SK MB ON PQ NB LB NF
samouellei (Kirby)
C. impressa (LeConte) -oNPQ-NS
C. oregona tLeConte) BC
C. rotundicollis Say -oNPQNBNS NF
C. rufa Linn6t
C. transmarina transmarina
(Motschulsky)
ec -_-9*PQIUNS:'
C. transmarina scopa (LeConte) BC
C. tubercu lata ( LeConte) PQ
armigera (Couper)
breuicollis (LeConte)
C. westwoodi tKirby)

Genus CYRTOMOPTERA Motschulsky

C. divisa (LeConter - BC
latiu sc ula Motschulsky

Genus RHAXOI.IYCHA Motschul sky

Key to North American species: McKey-Fender (1950)
R. bilobata (McKey-Fender) MB oN PQ
R. carolina t Fabricius) oN PQ
jactata (Say)

Genus RHAGONYCHA Eschscholtz

Key to North American species: Green (1940)
R. alaskensis Fender AK
R. anthracina Mannerheim AK
R. costipennis (LeConte) -oN------
MBONPQ-NS
R. excavata (LeConte) BC
R. fraxini (Say) AK BC AB MBONPQ_NSNF
atra (l\trby)
binodula Mannerheim
nigrita (LeConte)
R. fulva (Scopoli)t -BCPQ
R. gilvipes (Gemminger) NT_?
dichroa (LeOonte)
flauipes (LeConte)

193
Cantharidaer

R. imbecillis (LeConte) ON NF
albolineat a (Blatchley)
R. luteicollis (Germar) ON
cinctellt (LeConte)
R. mandibuiaris (Kirby) AK YK NT BC AB SK MB -PQ LB NF
nigritu.la (LeConte)
R. mollis mc,llis Fall SK ON NS NF
nigriceps (LeConte)
pusilla (LeConte)
R. nanula (LeConte) NS
R. oriflava ( LeConte) ON
R. recta (Melsheimer) ---ecAB-- oN PQ NS NF
pusilla (LeConte)
rufipes (Say)
R. scitula (Say) _PQ NS
R. septentri,rnis (Green) MB ON
R. syivatica (Green) ON
R. tantilla (l,eConte) -SK
pzsio (l,eConte)
R. vilis (LeConte) _ONPQ
R. walshi (LeConte) MB

Tfibe Silini
Genus POLItrMIUS LeConte
P. canadensis W.J. Brown
P. laticornis (Say) _ONPQ fuft_ _ _
dubjzs (Melsheimer)
incisus LeConte
inualid,.rs (Say)
undulatus (LeConte)

Genus SILII) Charpentier

Key to Ncrth American species: Green (1966)
(Subgenus SIILIS s. str.)
S. atra LeC<.rnte
S. difficilis ciffrcilis LeConte
S. insperata. Green
_YK
NT
NT BC
_BC
_BC
y:" MB PQ NB

S. lecontei Cireen
S. lutea LeConte _BC
pallens LeOonte
S. paliida Mannerheim AK _BC
S. parallela Green _BC
S. percomis iSay)
curta (7,eConte)
-o* PQ NB NS

longico"nis LeConte
S. spinigera LeConte BC
minuta LeConte
S. striatella Green BC
(Subgenus DITEMNUS LeConte)
S. bidentata (Say) PQ
S. latiloba Blatchley MB

Tlibe Malthinini
Genus MAJ.THINUS Latreille
Key to Ncrth American species: Fender (1972)
M. occipitalis LeConte oN PQ

Genus MALTHODES Kiesenwetter

Key to North American species: Fender (1951)
M. abruptus Fender _BC
M. acuticauda Fender _BC
M. alexanderi Fender _BC
M. bakeri Fender _BCAB
M. canadensis Fender _BC

194
 Derodontidae

M. columbiensis Fender BC
M. fragilis (LeConte) ON PQ NB NS
trdnsuersus (LeConte)
M. fuliginosus fuliginosus LeConte _PQ NB NS
M. fuliginosus flavicollis Fender NS
M. intricatus Fender
M. macnabi Fender
M. medioccidens Fender ON
M. niger (LeConte) _PQ NN T\TIT

M. oregonus Fender
M. parvulus (LeConte) oN PQ NB NS_- NT
M. piceolus Fall
M. piperi Fender
M. similis Fender ON NS
M. stacesmithi Fender
M. terraenovae Fender NF

Subfamily CIIAULIOGNATHINAE
Tlibe Chauliognathini
Genus CHAULIOGNATHUS Hentz
Key to North American species: Fender (1964)
C. pensylvanicus (Deceer) MB ON PQ
bimac u la t us (Fabricius)

Tlibe Ichthyurini
Genus TRYPHERUS LeConte
Lygerus Kiesenwetter
Key to North American species: Fender (1960)
T. frisoni Fender ON

Superfamily Dermestoidea
FamilyDERODONTIDAE tooth-neckedfungus beetles
D.E. Bright

The family Derodontidae includes four genera documented the feeding habits of Lqricobius erich-
and 19 species of small beetles that occur in the sonii quite well. Adults and larvae feed on the pine
temperate parts of both the Northern and Southern and spruce aphids of the family Adelgidae. The pre-
Hemispheres. They are rarely encountered in the ferred host of L. erichsonii is the balsam woolly
freld and are uncommon in collections. Three gen- aphid; the derodontid was introduced into eastern
era and eight species are found in Canada and Canada in the 1950s for the control of this pest.
Alaska. With the exception of Laricobius species, The other species of Laricobius also feed on other
both larvae and adults of this family feed on various species of Adelgidae.
kinds of fungi or on the products of fungal Information on adult ahd larval morphology,
metabolism. Peltastica species appear to be feeding behavior, life cycle, and evolution, as well as
restricted to fermenting sap flows or to areas under keys and diagnoses ofall genera and species, can be
bark, where bacterial or fungal fermentation takes found in Lawrence and Hlavac (1979). Lawrence
place. Derodontus species feed on the fruiting bod- (1989) recently published a catalog of the North
ies of a wide range of higher Basidiomycetes. The American species.
beetles prefer fresh fruiting bodies, although they
have been recorded in rotting mushrooms, "slimy AK (2); BC (5); aB (3); ON (2); PQ (1); NB (2); NS
fungus," and "watery fungus." Franz (1958) has (3);NF(1) . I(1)

195
Dermestida,:

Subfamily PELTASTICINAE
Genus PELi|ASTICA Mannerheim
P. tubercula.ta Mannerheim AK BC

Subfamily DERODONTINAE
Genus LARI.COBIUS Rosenhauer
_BC NF
L. erichsoni Rosenhauert
L. laticollis Fall _BCAB YNS
L. nigrinus Fender
L. rubidus l,eConte ---:cT-,oNpe NB

Genus DERCDONTUS LeConte

D. esotericus Lawrence
D. maculatu,s (Melsheimer)
D. trisig:natus (Mannerheim) AK ec
AB
__oN__N3__
Family NOSODENDRIDAE wounded'tree beetles
J.M. Campbell

Noso<lendradae is a small family that contains The New World species of the famil;' are well
only one ,lenus with about 50 species, two of which known taxonomically, thanks to a re'rision by
are recorded in Canada. According to Lawrence Reichardt (1976).
(1982a), I{osodendron species occur in most forested
areas of the world, where they inhabit slime fluxes
and rancid tree wounds, probably feeding on bacte-
ria, fungi,, and products of fermentation. BC (1);ON (1)

Genus NOSODENDRON Latreille

N. californir:um Horn BC
N. unicolor Sav ON

Family DERMESTIDAE dermestid beetles

Y. Bousquet

DerrrLestidae is represented in Canada and The taxonomy of the North American der-
Alaska b;y 48 species and subspecies. Dermestids mestids is relatively well known, thanks to the
are usually found on flowers, on dried animal car- work of Beal, who revised most genera occurring in
casses, in social Hymenoptera, bird and mammal the area. Mroczkowski (1968) published a catalog of
nests, where they feed on pollen, feathers, hair, the world fauna.
fur, or the remains of insects. Many species, how-
eyer, are household and museum pests; they cause
damage to a wide variety of products, such as car-
pets, silk, fur, feathers, wool, leather, seeds, eK (6); rlK (3); NT (6); BC (31); AB (16); SK (10);
grain, and cereal products as well as dried-insect MB (14); ON (2a); PQ (1e); NB (a); NS (5); PE (4);
collection s. NF(4) . J(17)

Tlibe Derrnestini
Genus DERMESTES Linn6
(Subgenus DERMESTINUS Zhantiev)
D. caninus {}ermar BC AB SK ON

196
 Dermestidae

murinus LeConte
nubilus Say
tessellatus Hetbst
D. carnivorus Fabriciust BC
rnucoreu,s LeConte
sobrinus LeConte
uersicolor Castelnau
SK MB
D. fasciatus LeConte
D. frischii Kugelannt _ _ _BC
_ ncT _MB
ONPQ_NS
D. maculatus DeGeert oN PQ
elongatus Hope
lupinus Mannerheim
truncatus Casey
uulpinus Fabricius
D. marmoratus Say _BCAB_MB
D. talpinus Mannerheim
D. tristis Fall
AK YK NT BC AB SK
_BCAB
MB oNpeI"_
(Subgenus DERMESTES s. str.)
D. ater DeGeert _MBONPQ
D. lardarius Linn6f --NrecAB SK MB ON PQ NB NSPE_NF
D. pulcher LeConte _MB_PQ
D. reductus Kalik _BC
D. signatus LeConte _BCAB SK MB

T[ibe Attagenini
Key to North American species: Beal (1970)
Genus ATTAGENUS Latreille
A. brunneus Faldermannt
elongatulus Casey
extricatus Casey
A. pellio (Linn6)t -BC NS
A. schaefferi hypar Beal _BC
A. unicolor japonicus Reittert -BC-SKMBONPQ PE NF
canadensis Casey
A. unicolor unicolor (Brahm)t
megatoma (Fabricius)
piceus (Olivier)

Genus NO\IELSIS Casey

Genattus Sharp
(Subgenus PARANOVELSIS Casey)
N. perplexa (Jayne) BC

Tribe Megatomini
Key to North American genera: BeaI (1967)
Genus TROGODERMA Dejean
Key to North American species: Beal (1954)
T. glabrum (Herbst)t AB -oN PQ
boronBeal
T. inclusum LeConte BC -SK MB ON PQ
aduena Casey
brunnescens Casey
frosti Casey
nigrescans Casey
obsolescens Casey
T. ornatum (Say) ON
pallipes Ziegler
paruulum Casey
procerum Casey
pusillum LeConte
serriferum Dalla Torre
serrigerum Casey
lorsole Melsheimer
T. simplex Jayne BC
ajar Casey
pollens Casey

197
Dermestidae

T. sinistrurn FaIl AK YK NT BC AB SK )IB ON PQ

T. sternale sternale Jayne BC _ONPQ
caseyi Arrow
simuh.tns Casey
uariipes Casey
T. variabilr'Balliont _AB SK ON PE
parab: Ie Beal
scabricennis Casey

Genus REESA Beal

R. vespulac (Milliront NT BC AB SK MB ON PQ PE NF

Genus MEGATOMA Herbst

Key to North American species: Beal (1967)
(Subgenus MEGATOMA s. str.)
M. pubescens (Zetterstedt) BC
prolira BeaI
(Subgenus PERIMEGATOMA Horn)
M. ampla ('lasey) BC
M. angularis Mannerheim AK
M. cylindri:a (Kirby) AK YK NT BC AB ON
impre:;sa (Casey)
montit:ola (Casey)
M. giffardi (Blaisdel]) AK
M. leucochlidon Beal ^-ec--
M. variegar.a tHornt BC AB
guadalupensis (Casey)
jaynei (Casey)
neuad;',ca (Casey)

Genus PStr UDOHADROTOMA Kalik

Key to North American species: Beal (1967)
P. kaliki B,ral BC
P. perversa (Fall) AK BC AB

Genus DETjIRTHRUS LeConte

Key to North American species: Beal (1967)
D. stebbinsi Beal BC

Genus CR\?TORHOPALUM Gu6rin-M6neville

Key to North American species: Beal (1985o)
C. haemonhoidale (J.E. LeConte) -oN
C. triste Le Conte -oN
fuscicl,aue Casey
modestum Casey
nigriatrne LeConte
picicorne LeConte

Tribe Anthrenini
Genus AN|IIHRENUS I.C. Schaeffer
Key to errstern Nonh American species: Hoebeke et a]. (1985)
(Subgenus ANTHRENUS s. str.)
A. pimpinellae Fabriciust BC
conspersus Casey
lepidus LeConte
neuad,icus Casey
obtectus Casey
occidens Casey
pictus Casey
suffusus Casey
A. scrophu ariae (Linn6)f BC ON PQ NB NS
thoracicus Melsheimer
(Subgenus NATHRENUS Casey)

198
 Bostrichidae

A. verbasci (Linn6)t BC AB MB ON PQ
caseyi Hinton
destructor Melsheimer
pistor Casey
substriatus Casey
uorax Casey
(Subgenus FLORILINUS Mulsant & Rey)
A. castaneae Melsheimer ON PQ NB
carolinae Casey
A. museorum (Linn6)t ON PQ NF
(Subgenus HELOCERUS Mulsant & Rey)
A. fuscus Oliviert oN PQ NS

Tribe Orphilini
Genus ORPHILUS Erichson
Key to North American species: Beal (1985b)
O. ater Erichson oN PQ
O. subnitidus LeConte BC
aequalis Casey
chalybeus Casey

Tribe Thylodriini
Genus THYLODRIAS Motschulsky
T. contractus Motschulskyt NT AB MB ON PQ

Superfamily Bostrichoidea
FamilyBOSTRICHIDAE bostrichidpowder'postbeetles
J. McNamara

Bostrichidae comprises 18 species in Canada Members of Bostricidae have been widely dis-
and Alaska. It includes some of the most destruc- tributed through commerce because of their ten-
tive beetles known. Some species cause consider- dency to bore into wood products, such as crating
able damage to such stored products as dried roots and bamboo.
and grains. In the tropics these insects are espe- Fisher (1950) revised the North American
cially destructive, injuring felled timber and bam- species of the family.
boo. The larvae do most of the damage. Some
species can kill weakened trees; others have the AK (1); vK (1); NT (3); BC (8); AB (3); SK (2); MB
habit of tunneling into the gTeen shoots of plants. (6); ON (13); PQ (6);NB (1);NS (1) ' I (1); H (1)

Subfamily PSOINAE
Tlibe Polycaonini
Genus MELA.LGUS Dejean
E ro pioide s Gu6rin-M6n6ville
Heterarthron Dejean
M. confertus (LeConte) BC
inciszs (LeConte)

Genus POLYCAON Castelnau

Alloeocnemis LeConte
P. stoutii (LeConte) BC
ouicollis (LeConte)

199
 Bostrichidac

Tribe Psoini
Genus PSOI, Herbst
Acrepis LeConte
P. quadrisig:rata tHornr - BC
sexguttt.,.ta Lesne

Subfamily ENDECATOMINAE
Genus END.IICATOMUS Me11i6
E. rugosus (Randall) MB ON PQ

Subfamily DINODERINAE
Genus RHYZIOPERTIIA Stephens
R. dominica ,Fabriciustt - BC MB ON pe
pusilla (Fabricius)

Genus PROSTEPHANUS Lesne

P. punctatus {Sayr - ON
Genus STEPHANOPACIIYS Waterhouse
S. cribratus t LeConter NT BC AB SK MB ON pe
S. rugosus (Ctlivier) - ON NB
opacus (Casey')
porcatu! (LeConte)
S. sobrinus lr)aseyr - BC AB
S. substriatus (Paykull)* AK YK NT BC AB - MB ON pe - NS
pacificu:; (Casey)

Subfamily BOSTRICHINAE
Tribe Bostrichini
Genus AMPI IICERUS LeConte
(Subgenus AI\.{PHICERUS s. str.)
A. bicaudatu,; (Sayr NT SK MB ON
asperico,t lis (Germar)
gracilis'Jasey
serricollt's (Germar)

Genus LICHIINOPHANES Lesne

L. armiger {Leconte) MB ON
L. bicornis (M/ebert - ON pe
L. mutchieri Belkin ON pe
angustatus (Casey)
-
L. truncaticollis tl-eConrer - ON

Tribe Xyloperthini
Genus SCOBICIA Lesne
r
S. bidentata tHorn
- ON
S. declivis (LeContet - BC

Genus XYLOBIOPS Casey

X. basilaris tSayt
basalis ( Dejean)
- ON

h ume ral is (Melsheimer)

200
 Anobiidae

Family LYCTIDAE lyctid powder'post beetles

J. McNarnara

Only six species of lyctids are known to occur in cause further damage to wood by producing exit
Canada. Members of this family are commonly holes when they emerge. The family is worldwide
called powder-post beetles because of the tendency in distribution, each region having an indigenous
of the larvae to reduce sapwood, especially of hard- fauna plus introduced species.
woods, into a powdery frass. They are found more Gerberg (1957) has revised the New World
often in recently dried wood rather than in old species.
wood. They attack lumber used for furniture, crat-
ing, hardwood floors, and other purposes. Second
only to termites, Lyctidae cause the most destruc- BC (4);uB (1); oN (4);PQ (3);NB (1);NS (2) o I
tion to wood and its products. The adult beetles (2)

Tribe Lyctini
Genus LYCTUS Fabricius
L. brunneus (Stephens)f BC PQ NB NS
carolinae Casey
L. cavicollis LeConte BC
L. linearis (Goeze)f BC oN PQ
arillaris Melsheimer
striatus Melsheimer
L. opaculus LeConte -oN
breuipennis Casey
L. planicollis LeConte BC MB ON PQ NS

Genus TROGOXYLON LeConte

T. parallelopipedum (Melsheimer) ON

Family ANOBIIDAE deathwatch beetles

J. McNaruara

Anobiids include 79 species in Canada and (cigarette beetle), are serious pests of stored prod-
Alaska. Larvae of most species bore into dead hard- ucts, including drugs, tobacco, seeds, spices, cereal
woods and softwoods; two species of Ernobius attack products, leather, and museum specimens.
and destroy pine cones; one species, Anobium punc- North American anobiids are relatively well
tq.tum (furniture beetle), damages woodwork, furni- known taxonomically, thanks to the work of White,
ture, bookbindings, and similar products. The bee- who published a key to the genera (1971), a number
tles often produce a tapping sound in their burrows, of generic revisions, and a catalog of the North
which gives rise to the ancient superstition that they American species ( 1982).
are a warning of death, hence the common name.
Some species feed in the young stems or shoots of AK (4);YK (6);NT (2); BC (35);AB (18); SK (9); MB
growing trees. Two species, Stegobium paniceum (21); oN @2);PQ (40);NB (8);NS (7);NF (7) o I
(drugstore beetle) and Le.sioderma serricorne (5)

Subfamily DRYOPHILINAE
Tlibe Eucradini
Genus EUCRADA LeOonte
E. humeralis (Melsheimer)
_BC -oN PQ
E. robusta Van Dyke

201
Anobiidae

Subfamily ERNOBIINAE
Tribe Xestobiini
Genus XESiTOBIUM Motschulsky
Hyperisus Mulsant & Rey
Key to North American species: White (1975o)
X. abietis Fisher BC
X. alfine LeConte BC
X.
X.
gaspensrs R.E. White
marginicolle (LeConter BC
-oN PQ

Genus UTOBIUM Fall

Key to North American species: White (1966, 1976)
U. elegans iHorn)
U. granulal,um R.E. White
U. marmorrtum Fisher
_ v_ BCAB_MBONPQ
BCAB_PQ
BC

Tlibe Ernobiini
Genus EPISERNUS Thomson
E. trapezoideus (Fall) BC

Genus ERITIOBIUS Thomson

Philoxylon LeConte
E. bicolor R.E.White NF
E. gentilis Jlall BC
E. luteipen.ris LeConte PQ
E. mollis (Iinn6)+ ON PQ_NS NF
conu et ifrons (Melsheimer)
E. nigrans ,lall _BC
E. pallitarsis Fall _BC
E. punctulatus (LeConte) _BC
E. schedli ![.J. Brown ON PQ NF

Subfamily ANOBIINAE
Tribe Euceratocerini
Genus CTENOBIUM LeConte
C. antennal;um LeConte oN PQ

Tlibe Gastrallini
Genus GASTRALLUS Jacquelin du Val
G. marginipennis LeConte BC

Tlibe Nicobiini
Genus TRICHODESMA LeConte
Ptinodzs LeConte
T. cristata Oasey
T. gibbosa (Say) _ _ _:c_ _ _ ON
T. klagesi Fall ON

TYibe Stegobiini
Genus STETIOBIUM Motschulsky
S. paniceum (Linn6)t AK BC SK MB ON PQ NB
obesurn (Melsheimer)
tenuistriatum (Say)

Genus OLIGOMERUS Redtenbacher

Key to North American species: White (1976)
O. alternan; LeConte ON PQ

202
 Anobiidae

O. crestonensis Hatch
O. obtusus LeConte
BC
lo*pe--
O. sericans (Melsheimer) _ONPQ

Tribe Anobiini
Genus PLATYBREGMUS Fisher
P. canadensis Fisher ON NS

Genus ANOBIUM Fabricius

A. punctatum (DeGeer)f BC MB_PQ NS
pumilum LeConte

Genus HEMICOELUS LeConte

Cacotemnus LeConte
Key to North American species White (1976)
H. carinatus (Say) BC AB SK MB ON PQ NB
errons (Melsheimer)
linearis (LeConte)
H. defectus (Fall)
H. gibbicollis (LeConte) AK
_BC
BC
yu oN PQ

destructor (Fisher)
roguensis (Hatch)
H. pusillus (Fall) -oNPQ
H. umbrosus (Fall) AK YK AB MB ON PQ NB

Genus MICROBREGMA Seidlitz

M. emarginatum emarginatum BC AB SK MB ON PQ
(Duftschmid)t
foueatutn (Kirby)
M. emarginatum granicolle Fall BC

Tribe Hadrobregmini
Genus HADROBREGMUS C.G. Thomson
Coelostethus LeConte
H. americanus (Fall) BC AB
H. notatus t Sayt oN PQ NS__-
H. quadrulus (Leconte) BC
H. truncatus (Fall) BC

Genus PRIOBIUM Motschulsky

Key to North American species: White (1975b)
P. sericeum (Sayt MB ON PQ NB NS

Genus DESMATOGASTER Knutson

D. subconnata (FaIlt YK AB MB PQ

Subfamily PTILININAE
Tribe Ptilinini
Genus PTILINUS O.F. Mriller
P. basalis LeConte BC
P. Iobatus Casey YK_BCAB_MBON
serricollis Say
P. pruinosus Casey -MBONPQ NB
P. rufrcornis Say -ABONPQ _NS
bicolor Melsheimer

Subfamily XYLETININAE
Tribe Xyletinini
Genus VRILLETTA LeConte
Pseudoxyletinus Pic
V. decorata Van Dyke BC
V. laurentina Fall ON PQ

203
Anobiidae

Genus EIIIRILLETTA Fall

Oligor;wodes Fall
Key to North American species: White (1985)
E. brevis R E. White AB -oN
E. harrisii lFall)
E. peltata (Harris)
-oNPQ
MB ON PQ
Genus XYI ETINUS Latreille
Key to N crth American species: White (1973, 1977)
X. bicolor E.E. White -YK AB SK MB
X. confusus R.E. White NT BC AB PQ
X. fucatus i,eConte _BC_SK MB ON PQ NB NF
X. lugubris LeConte
X. rotundic'rllis R.E. White -BCABSK
AB SK yu_:a
Tribe Lasiodermini
Genus LASIODERMA Stephens
Key to North American species: White (1990)
L. serricorne (Fabricius)t AK MB ON PQ
castanzum Melsheimer

Subfamily DORCATOMINAE
Tlibe Calymmaderini
Genus CALYMMADERUS Solier
Eupac.lus LeConte
C. nitidus ( LeConte)

TYibe Petaliini
Genus PETALIUM LeConte
Key to N'rrth American species: Ford (1973)
P. bistriatum (Say) ON
P. seriaturr Fall ON PQ

Tribe Prothecini
Genus SCULPTOTHECA Schilsky
S. puberula tLeConte) ON PQ

Genus STAGETUS Wollaston

Key to New Worid species: White (19756)
S. prolundus (Leconte) ON PQ
s triat orc u n ctatzs (LeConte)

TYibe Dorcatomini
Genus DOIi,CATOMA Herbst
Key to North American species: White (1966)
D.
D.
falli R.E, White
integra ( Fall) _BC -oN
D. moderata R.E. White MB ON PQ
D. pallicornis LeConte -SKMBONPQ NB NF
foueata R.E. White
D. setulosa LeConte oN PQ

Genus CAENOCARA C.G. Thomson

Tylistus LeOonte
C. bicoior (Germar) MB ON
C. blanchardi Fall _BC
C. oculaturrL (Say) _ONPQ NE NS
C. scymnoides LeConte _YKNTBCABSK MB ON PQ
C. tenuipalpum Fall _,,

204
 Ptinidae

Genus BYRRHODES LeConte

B. intermedius (Leconte) PQ

Subfamily TRICORYNINAE
Tribe TYicorynini
Genus TRICORYNUS Waterhouse
C atorama Gu6rin-M6neville
Hemiptychus LeConte
Key to North American species: White (1965)
T. borealis (LeContet MB ON
T. confusus (Fall) PQ
T. densus (Fall) AB
rogersi (Hatch)
T. nigritulus (Lecontet - ON
T. nubilus tFallt
schuhi (I].atc]e],

Family PTINIDAE spider beetles

Y. Bousquet

Only 16 ptinid species are found in Canada and Ptinidae, the common name of which refers to
Alaska, and all but three are introduced on this con- the fact that the adults of many species look like
tinent. Many of these species are found in mills, small spiders, is closely related to Anobiidae. The
warehouses, granaries, stores, museums, and hospi- species are divided into two groups that are consid-
tals where they feed, in both the adult and larval ered to be subfamilies or tribes. Papp has reviewed
stages, on a wide variety of products including the North American fauna (1962). Many species
grain, seeds, and drugs. These species are usually that occur in Canada can be identifred with the key
minor pests, since they rarely build up large popula- provided by Brown (1940).
tions. Spider beetles also occur in nests of mam-
mals, birds, and bees; they are probably scayengers AK (1); BC (13);AB (5); SK (6); IVIB (7); ON (11); PQ
that feed on dry animal and vegetable matter. (11);NB (9);NS (10); PE (1); NF (5) . I (13)

Subfamily GIBBIINAE
Genus GIBBIUM Scopoli
Scotias Czenpinski
G. aequinoctiale Boieldieuf - ON PQ NB NS
psylloides auct.

Genus MEZIUM Curtis

M. affrne Boieldieut - BC - SK MB ON PQ NB NS

Subfamily PTININAE
Genus SPHAERICUS Wollaston
S. gibboides tBoieldieut+ - BC

Genus TIPNUS C.G. Thomson

Epauloecus Mulsant & Rey
T. unicolor (Piller & - NB NS
Mitterpacher)t

Genus TRIGONOGENIUS Solier

T. gJobulus Soliert - BC
farctus LeConte
Genus NIPTUS Boieldieu
N. hololeucus (Faldermann)f - BC AB SK MB ON PQ NB NS PE - NF

205
Trogossitida e

Genus PSEI.IDEUROSTUS Heyden

P. hilleri (Reitter)t BC AB ON PQ NB
alienus (W.J. Brown)

Genus PTINUS Linn6

(Subgenus I'TINUS s. str.)
P. bicinctus Sturmf _BCABSK MB oN PQ NB NS
P. clavipes ['anzert _BC ON PQ NS
b r unne;t. ts Duftschmid

hirtellus Sturm
testaceus Olivier
P. fur (Linn6)t BC ON PQ NB NS NF
humeralis Say
lofro Fabricius
P. raptor Sturmt BC _SKMBONPQ NB NS NF
P. villiger (Reitter)-f BC AB SK MB ON PQ NS NF
(Subgenus CYNOPTERUS Mulsant & Rey)
P. californicus Pic BC MB_PQ
P. fallax Fall BC
P. falli Pic -oN
_SKMBONPQ NS _NF
P. ocellus W J. Brownf AK BC NS
tectus aoct.

Superfamily Lymexyloidea
FamilyLYMD(YLIDAE ship-timberbeetles
J.M. Campbell

Thirt;'ssvsn species of Lymexylidae are known that Lymexylidae is closely related to Cleroidea and
around the world. Only one rare species occurs in Cucujoidea. Arnett (1968) places the farnily near
Canada. .\dults are found in decaying wood, on tree Lampyridae, based on wing venation. Wheeler
trunks, or under bark. The Canadian species is (1986) revised the world genera and higher classifi-
probably .introduced, since it also occurs in Europe cations, including a study of the phylogerny, mor-
and was firrmerly very destructive to wooden ships. phology, and known biology of the genera.
The classifrcation of the family is now fairly sta-
ble. Crorvson (1955) and Wheeler (1986) believe aB (1);ON (1);PQ (1);NB (1); NS (1);NF (1)

Subfamily FIYI ECOETINAE

Genus IIYLItrCOETUS Latreille
Elatero;;.des J.C. Schaeffer
Hyloceotus Melsheimer
H. Iugubris lJayf AB ON PQ NB NS NF
americc:nus }{aris

Superfamily Cleroidea
Family TROGOSSITIDAE bark-gnawing beetles
Y. Bousquet

Trogossitidae includes 22 species in Canada and Barron (1971) has revised the trogositids of
Alaska. Most North American species are found North America. The species are divided into two
under the bark of dead trees, in logs, and in stumps; subfamilies, Peltinae and Trogossitinae.
a few species occur in fungi. The adults and larvae
of many species feed on wood-eating insects; others
feed on dercaying matter or fungi. Tenebroides mau- AK (a);YK (a); NT (5); BC (17); aB (6); SK (6); MB
ritanicus is a well-known stored-product pest that (6); oN (12); PQ (e); NB (2); NS (2); PE (1); LB (1);
feeds on grains and cereal products. NF(3) . H(2)
206
 Trogossitidae

Subfamily PELTINAE
Tribe Peltini
Genus CAI-ITYS C.G. Thomson
Nosodes LeOonte
Peltidea Motschulsky
C. minor Hatch _YK_BCABSK_ONPQ
C. scabra (Thunberg)* AK_NTBCAB_MBONPQ
sercata (LeConte)
silphides (Newman)

Genus OSTOMA Laicharting

Gaurambe C.G. Thomson
O. columbiana Casey AK YK NT BC AB SK MB ON PQ NB LB
O. ferruginea (Linn6)* AKYKNTBCABSK_ONPQ NS NF
fraterna (Randall)
nigrina Casey
septentr io nalis ( Randall)
O. pippingskoeldi (Mannerheim) BC

Genus GRYNOCHARIS C.G. Thomson

G. oregonensis (Schaeffer) BC
G. quadrilineata (Melsheimer) MB ON PQ
marg inata (Melsheimer)

Genus THYMALUS Latreille

T. marginicollis Chevrolat BC AB SK MB ON PQ NS NF
aubei L6veill6
fulgidus Erichson
Genus ERONYXA Reitter
Ostomodes Reitter
E. paliida (Motschulsky) BC
dohrni (Reitter)
lagrioides Reitter
pilosula (Crotch)

Subfamily TROGOSSITINAE
Tribe Trogossitini
Genus NEMOSOMA Latreille
Pseudalindria Fall
N. punctulatum Van Dyke BC
punctatum Van Dyke

Genus CORTICOTOMUS Sharp

Parafilumis Casey
C. caviceps (Fall) BC
laeuiuentris Casey

Genus AIRORA Reitter

A. aequalis Reitter BC
bicolor Casey
polita Casey
punctiuentris Casey
A. cylindrica (Audinet-Serville) ON
nlgello (Melsheimer)
nigra (Melsheimer)
teres (Melsheimer)

Genus TEMNOCHILA Westwood

T. chlorodia (Mannerheim) NT BC
melanica lJ.atc}a
prosternalis Schaeffer
u ir idicy anea (Mannerheim)

207
Cleridae

Genus TEI'IEBROIDES Piller & Mitterpacher

Trogossita Olivier
T. americanus tKirby) ON PQ
castat teus (Melsheimer)
nigrit ^rs (Horrr)
T. collaris Sturm)
T. corticalis (Melsheimer)
dubius (Melsheimer)
AK YK NT ec- sxusBNpe - -- - - -
internrcdius (.Horn)
limbalis (Melsheimer)
T. crassico'nis (Horn) BC
califo.ni"u" 11fornt
pleurulis (.Horn)
T. laticollir, (Horn)
obscu:us (Hotn)
-oN
T. mauritanicus t Linn6rf BC SK MB ON PQ NB PE NF
nitidt s (Horn)
T. occidenlalis Fall BC
T. sinuatu,r (LeConte) BC

Family CLERIDAE checkered beetles

J. McNarnara

Cleridae includes 50 species and subspecies in In North America the taxonomy of clerids is rea-
Canada and Alaska. Most species are predaceous, sonably well known, but the literature is scattered.
the adul':s feeding on adult bark beetles and the lar- A revision of the family would be helpful. Barr
vae on eggs, larvae, and adults of wood-boring bee- (I962a) has published a key to the genera. There is
tles in galleries. Several species of Enoclerus and no comprehensive work for the Canadian fauna, but
Thanasimus are believed to be imoortant in the most eastern species could be identifred by referring
natural r:ontrol ofbark beetles in coniferous forests. to Knull (1951) and most western species by refer-
Adults of Cleridae are found mostly on plants, tree ring to Barr 0962b).
trunks, or under bark. Some are found on flowers,
feeding on pollen. Some species of Necrobia are AK (5); vK (4); NT (5); BC (22); AB (12); SK (10);
scavengers and are economically important as pests MB (14); ON (32); PQ Q4; NB (11); NS [4); NF (4)
of stored food, carcasses, and skins. . I(5)
Subfamily CLERINAE
Tlibe Tillini
Genus MONOPHYLLA Spinola
Elasn ocerus LeConte
Macrttelus lllug
M. terminata (Say) ON
megatoma Spinola

Genus CYIyIATODERA Gray

C. bicolor (Say) oN PQ
C. decipiens Fall BC _-n'___
C. inornatiL (Say) ON PQ
tenera LeConLe
C. ovipennis LeConte BC
pilosei.Ia LeConte
C. undulata (Say) ON
boscil (Chevrolat)
longicollis Spinola
uauth.ieri Spinola

Tribe Phyllobaenini
Genus PH'{LLOBAENUS Deiean
Hydnocera Newman

208
 Cleridae

P. dubius (Wolcott)
P. humeralis (Say)
cyanescens (LeConte)
- - - ecABSfMBoNFeNs- - - -
cydneus (Spinola)
dfficilis (LeConte)
P. lecontei (Wolcott)
P. pallipennis (Say)
serratus (Newman)
T- I'BC-:"MBBNiSNBws- - -
P. scaber (LeConte) _BC
P. subfasciatus (LeConte) _BCABSKMB
P. unifasciatus (Say) -oN
punctatus (Spinola)
P. verticalis (Say) -oNPQNB
brachypterus (I0.ug)
I ine a toc oll is ( Spinola)

tenellus (.Say')

Genus ISOHYDNOCERA E.A. Chapin

I. curtipennis (Newman) MB ON PQ NB NS
longicollis (Ziegler)
L tabida (LeConte) _ONPQ

Tribe Thaneroclerini
Genus ZENODOSUS Wolcott
Z. sanguineus (Say) -ONPQ

Tribe Clerini
Genus PRIOCERA Kirby
P. castanea (Newman) - ON
maculata Ziegler
rufescens Spinoia

Genus OPILO Latreille

O. domesticus (Sturmt*

Genus THANASIMUS Latreille

T. dubius (Fabricius) AK YK NT BC AB SK MB ON PQ NB - NF
erythrocep halus (Spinola)
ruficeps (Spinola)
T. trilasciatus (Say) - ON PQ
T. undatulus (Say) AK \T( NT BC AB SK MB ON PQ NB NS NF
abdominalis I{rtby
mel anocep halus Chevrolat
nubilus (l\lug)

Genus PLACOPTERUS Wolcott

P oeci loc hroa Chevrolat
P. thoracicus tOliviert MB ON PQ
monilis (Melsheimer)
pallipes (Wolcott)

Genus ENOCLIIRUS Gahan

E. barri Knull - BC
E. coccineus coccineus (Schenkling) AB SK
corallinus (Fall\
E. eximius (Mannerheim) - BC
holosericeus (A. White)
E. ichneumoneus (Fabricius.t - ON
rufus (Olivier)
E. Iaetus tKlugt - BC
abruptus (LeConte)
E. lecontei (Wolcott) - BC
nigriuentris (LeConte)
E. muttkowskii (Wolcott) - ON PQ NB
liljebladi Wolcott

209
Cleridae

E. nigrifrons tSayt * ON PQ
gerhardi Wolcott
testaceipes Wolcott
E. nigripes nigripes {Say) MB ON PQ NB
dublzs (Spinola)
incertus (LeConte)
E. nigripes rufiventris (Spinola) - ON PQ NB
E. rosmarus (Say) - ON
oculatus (Spinola)
E. schaefferi Barr - BC AB
humer,zlis Schaeffer
E. sphegeus (Fabriciust - BC AB
sobrius (Walker)

Genus TRIOHODES Herbst

Key to Norbh American species: Foster (1976)
T. apivorus borealis Wolcott &
E.A. Chapin
interruptus LeConte
nuttalii Say
trifasaatus Strnn
T. nutalli tliirby) AB SK MB ON PQ
T. ornatus lrartwegianus A. White AK YK NT BC AB

Subfamilv KORYNETINAE
Tribi Epiphloeini
Genus PHI OGISTOSTERNUS Wolcott
Phyllo nenus Spinola
P. dislocatus {Sayt MB ON PQ NB
distrop.hus (IArg)
transuersalis (Spinola)

Genus NEI(IHNEAWolcott & E.A. Chapin

N. laticorni ; (Sayt - ON

Tribe Enopliini
Genus NECTRTHOPLEURA Barr
Key to sprecies: Barr (1976)
N. thoracicr'L ( Say r - ON PQ
b imac t.iata (Melsheimer)
nig ript: n ni s ( Spinola)
p ennsy lu anica (Chevrolat)
p uncto tis s im a (Chevrolat)

Genus CHARIESSA Perty

B r achy mor p hus Chevrolat

C. elegans Horn - BC
lemberti La Rivers
C. pilosa (F)rster) - BC AB SK MB ON PQ
cinclo lSpinola)
margirLata (Say)
onusta (Say)

Genus BOSCHELLA Barr

B. fasciata tl-eConte) - BC
trilobata (Van Dyke)

Genus CREGYA LeConte

Galeruclerus Gahan
C. oculata (Say) - ON
marginipennis ( Spinola)

2to
 Melyridae

Tribe Korynetini
Genus NECROBIA Olivier
N. ruficoilis (Fabricius)t -BC-oNPQ
N. rufrpes (DeGeer)t _BCABONPQNF
N. violacea (Linn6)t AK YK NT BC AB SK MB ON PQ NF
errons Melsheimer

Subfamily TARSOSTENINAE
Genus TARSOSTENUS Spinola
T. univittatus (Rossi)f BC
alb ofa sc iat us ( Melsheimer)

Family MELYRIDAE soft-winged flower beetles

D.E. Bright

Melyridae includes 49 species known to occur in careful study. There are no recent revisions or taxo-
Canada. The adults commonly occur on flowers and nomic treatments of the Nearctic species; the infor-
evidently feed on other insects as well as on pollen. mation below therefore reflects our best effort at
Larvae are primarily predators, occurring in vari- achieving a list based on available literature.
ous habitats but most commonly in soil, leaf litter, Arnett (1968) provides a key to the genera and
or under bark. One species, Malachius deneus,has Hatch (1962) a key to the species of the Pacific
been introduced in North America and causes con- Northwest.
siderable damage to developing wheat in the Prairie
Provinces.
The taxonomy of the North American melyrids YK (2); NT (3); BC (35);aB (16); SK (5); MB (6); ON
is in poor condition, and the family is in need of (10);PQ (10);NB (1);NS (1);NF (1) ' I(2)

Subfamily MALACHIINAE
Genus COLLOPS Erichson
C. bipunctatus (Say) AB SK
C. cribrosus LeConte BC
cy anip en nis Motschulsky
C. discretus Fall BC
C. hirtellus LeConte
C. quadrimaculatus (Fabricius) __ftBCy:"_ oNpe-----
C. tricolor (Say) SK MB oN PQ
C. versatilis FalI Bc
C. vittatus (Say) - - lrr AB ir nrs ON PQ
hauorthi Westwood

Genus ENDEODES LeConte

Key to North American species Moore & Legner (1975b)
E. collaris LeConte BC

Genus MALACHIUS Fabricius

Anthocomus Erichson
Hapalorhinus LeConte
Microlipus LeConte
M. aeneus (Linn6)l -BCAB ON PQ NB NS NF
M. auritus LeConte _BC
M. bipunctatus (Harrer)t ON PQ
M.
M.
blaisdelli Hopping
erichsonii (LeConte)
-Bcms
criddlei W.J. Brown
lateralis LeConte
M. flavilabrus (Say) ON PQ
coeruleus Randall

2tl
Melyridae

M. floricola Marshall - BC
M. francisc,rnus Fall - BC
M. horni FrLll - BC AB
antenruatus Hopping
M. macer fl.orn - BC
M. mixtus l{orn - BC
M. mirandrrs (LeConte) - BC
M. moerens. LeConte - BC
u n ifor in is Motschulsky
M. nigrinui Fall - BC
M. thevene,ii Horn - BC AB
contorl.us Fall
M. ulkei H<'rn AB - MB ON PQ

Genus TAN'AOPS LeConte

C ap halistes Motschulsky
T. ignitus Fall BC
bosalis W.J. Brown
blaisdelli (Malkin)
i Marshall
n u ne n;n ac her
sierroe Marshall
T. malkini l{arshall - BC

Genus HYF'EBAXUS Kiesenwetter

Pseude'baeus Horn
H. apicalis Sayt
oblitus (.LeConte)
H. bicolor (',eContet - BC AB

Genus ATTAI-US Erichson

(Subgenus .\CLETUS LeConte)
A. nigrellus (LeConte) - YK - BC AB ON PQ
falli (I:lopping)
(Subgenus ,\TTALUS s. str)
( S calolt ter u s Motschulsky)

A. morulus smithi HoppinC - BC

A. pallifrons tMotschulskyt - ON
A. humeral .s LeConte
pettiti Hotn

Subfamilv RHADALINAE
Key to geriera: Peacock (1987)
Genus SEM.IJULISTUS Schilskv
Eurelymis Casey
S. ater ( Le('ontel - BC AB

Subfamily DASYTINAE
Key to North American genera: Blaisdell (1938o)
Genus DOL,ICHOSOMA Stephens
D. foveicollc tKirbyt - BC AB - MB ON PQ

Genus DASYTES Paykull

D. cruralis LeConte) - BC

Genus AMPHIVECTURA Blaisdell

A. monticol r tBlaisdellr - BC AB

Genus HOF PINGIANA Blaisdell

H. hudsonir:a (LeConte) - \'K NT BC AB SK MB - PQ
breuilabris Blaisdell
kingiW.J.Brown

212
 Sphindidae

Genus TRICHOCHROUS Motschulsky

T. alberbensis Blaisdell AB

Genus AMECOCERUS Solier

Listrus Motschulsky
A. confusus Casey _BC
A. glabratus Hatch _BC
A. medicatus Blaisdell AB
A. olSrmpianus Blaisdell _BC
A. plenus Casey _BC
A. provincialis Blaisdell -BC
A. quadricollis Hatch _BC
A. robustus Blaisdell _BC
A. senilis (LeConte) AB

Genus DASYRHADUS FalI

D. impressicollis Fall BC

Superfamily Cucuj oidea

FamilySPHINDIDAE dry-fungusbeetles
J.M. Campbell

Sphindidae is a small family comprising 35 (1979) revised the higher classification of the fam-
species around the world, of which five are known to ily, which is in need of a taxonomic revision; most
occur in Canada. Adults and larvae feed on spores current records are unreliable.
of slime molds.
Casey (1898) prepared the last revision of the
North American sphindids. Sen Gupta and Crowson BC (2);AB (1); oN (3);PQ (4)

Subfamily SPHINDINAE
Tribe Odontosphindini
Genus ODONTOSPHINDUS LeConte
O. clavicornis Casey BC
O. denticollis LeConte ON pc)

Tribe Sphindini
Genus SPHINDUS Chewolat
Coniophagus Mink
S. americanus LeConte -BCABONPQ _ONPQ
S. trinifer Casey

Subfamily EURYSPHINDINAE
Genus EURYSPHINDUS LeConte
E. hirtus LeConte PQ

213
Nitidulidaer

Family NITIDULIDAE sap beetles

J. McNamara

The lamily Nitidulidae comprises 101 species in the pollen and petals of the same plantli or some-
Canada and Alaska. Members of this family are times of others.
mostly saprophagous and mycetophagous. The Parsons (1943) published a revision of the
majority live in decaying fruit, fermenting plant Nearctic Nitidulidae, and most western species can
juice, an<l fungi, but some also live in flowers. Some be keyed out by referring to Hatch (1962).
species breed in the nests of bumblebees and others
in carrion. The larvae of some species prey on
scolytids: the larvae ofothers are considered fungiv- AK (17); YK (10); NT (8); BC (63); AB (44); SK (16);
orous. I'he larvae of Cateretinae live in seed cap- MB (45); ON (56);PQ (63); NB (13);XS (15);PE (3);
sules of 'sarious plants, with the adults feeding on LB (1);NF (13) . H (3);I (7)

Subfamily CATERETINAE
Genus HEI'ERHELUS Jacquelin du Val
H. pennatus (Murray) BC ON PQ NF
crinitt..s (Murray)
H. sericans (LeConte) BC

Genus CAI'ERETES Herbst

(Subgenus CATERETES s. str.)
C. scissus Parsons AK_NT AB SK ON PQ

Genus BOIiEADES Parsons

B. abdomiralis (Erichson) MB ON PQ

Genus BRI'CHYPTERUS Kugelann

B. globularius Murray _BCAB
B. troglodytes Murray _BC
B. urticae (Fabricius)f NT BC AB SK MB ON PQ NF
pusill Lrs (Melsheimer)

Genus BRACHYPTEROLUS Grouvelle

B. pulicarius (Linn6)t AB SK MB ON PQ NB NS NF
mordelloides Notman

Subfamily CARPOPHILINAE
Genus COI,IOTELUS Erichson
C. obscuru,r Erichson MB ON PQ NB NS

Genus COLOPTERUS Erichson

Colastus Erichson
CYlloP'liss MurraY
Key to North American species: Parry & Howden (1975)
C. maculatus (Erichson)
C. semitect.us (Sayt
C. truncatus (Randall) YK
-BC -AB -SK -MB 3Npe-
ON PQ NS
infimus (Erichson)
limba,tus (LeConte)
obliquus (LeConte)

Genus CAIIPOPHILUS Stephens

(Subgenus CARPOPHILUS s. str.)
(Tribrachys LeConte)
C. antiquus Melsheimer PQ
C. brachypterus (Say) mn on PQ
carbonatus LeConte

214
 Nitidulidae

C. dimidiatus (Fabricius)* MB oN PQ
C. discoideus LeConte BC ON
apicalis LeConte
caudalis (LeConte)
C. hemipterus (Linn6)f AK BC AB ON PQ
b imac ulatu s Melsheimer
C. ligneus Murray AK
decipiens l{orn
C. lugubris Murray AB -oN
MB ON pe
C. marginellus Motschulskyf
nitens Fall
C. mutilatus Erichsonf BC
luridus Murray
C. pallipennis (Say) AB SK
pollens Sharp
C. sayi Parsons MB PQ

Subfamily NITIDULINAE
Genus EPURA-EA Erichson
Epuraeanella Ctotch
E. adumbrata Mannerheim AK-NTBCAB PQ
E. aestiva (Linn6)t AK BC AB ms oN PQ NF
conu exi u sc ula Mannerheim
E. alternans Grouvelle _BC
alticola Fall
E. alternata Parsons -oNPQ
E. ambigua Mannerheim AK_- BC
E. avara (Randall) AK YK BC MB ON PQ NB NS NF
nubila LeCotte
E. boreades Parsons BC _ONPQ NB
E. corticina Erichson PQ
E. erichsoni Reitter NT BC-SK MB ON PQ
E. flavomaculata Miiklin AK _AB MB ON PQ NS
E. fulvescens Horn -oNPQ
E. helvola Erichson SK MB ON PQ
castanea (Melsheimer)
rufa Reitter
E. horni Crotch ON PQ
duryiBlatchley
E. labilis Erichson BC MB ON PQ NS__
E. linearis Maklin
E. obliquus Hatch
E. obtusicollis Reitter
AK
_I' BC AB
BC AB
BC AB SK MB_PQ
PQ
PQ NB

E. ovata Horn BC MB_PQ

Ns__
E. parsonsi Connell
E. peltoides Horn
E. planulata Erichson AX
I"_ BC AB
_AB
BC AB
SK
MB ON
MB ON
PQ
PQ
PQ NB
NF

placida Meklin
E. populi Dodge BC AB MB_PQ
E. rufa (Say) _ONPQ
bodio (Melsheimer)
rotundicollis Reitter
E. rufida (Melsheimer) _ONPQ
E. terminalis Mannerheim* AK YK _BC AB MB ON PQ NB
infuscata Miklin
E. truncatella Mannerheim AK YK NT BC AB SK MB ON PQ LB NF
nigraMeklin
ornatula Notman
E. umbrosa Horn PQ NB

Genus APHENOLIA Reitter

A. monogama (Crotch) BC

Genus STELIDOTA Erichson

S. octomaculata (Say) ON PQ
biseriata Reitter

2t5
 Nitidulidae

Genus OMOSITA Erichson

(Subgenus IJAPROBIA Ganglbauer)
O. colon (Linn6)t - BC AB SK MB ON pe NB NS NF
O. discoidezL (Fabricius){ AK BC AB - MB ON pe - NS NF
inuersc: LeConte

Genus NITIDULA Fabricius

N. bipunctata (Linn6)t AK YK - BC AB - MB ON pe
N. carnaria (Schaller)i BC ON pe -
- NS
N. nigra Sc.iraeffer AK yK - BC AB - MB
N. rufipes (,inn6)t - BC AB - MB ON pe NB NS NF
ossium l{ttby
N. ziczac Say NT BC AB - MB
humerr:tlis LeConte
inorna;a Horn
unig u t i:a t a Melsheimer

Genus PRO METOPIA Erichson

P. sexmaculata tSayr ON pe
Genus LOB [OPA Erichson
Cerophorus Castelnau
L. setosa Hi,rold BC
substriata (Hamilton)
L. undulata tSayr AB - MB ON pO

Genus SOR')NIA Erichson

S. grisea (Linn6rt NF
S. guttulata tl-eConter - BC

Genus PHE.\OLIA Erichson

P. grossa tF.rbricius) ON pO

Genus THA-YCRA Erichst-rn

Perthaiycra Horn
Key to New World species: Howden (1961)
T. concolor (LeConte) - YK - BC AB - MB ON pe
T. dentata I l. Howden - BC
T. keltoni H Howden - BC AB
T. leechi H. Howden BC
T. mixta H. Howden BC AB
T. monticola H. Howden - BC
T. murraf lHornt - BC AB
T. parsonsi .r{. Howden - BC
T. sinuata H. Howden - BC AB

Genus POC,,\DIUS Erichson

P. fulvipenn s Erichson BC
dorsali:; Horn
P. helvolus llrichson NT BC - SK MB
breuiusculus Reitter
infusca;',us Reitter
limbatt.s Reitter

Genus AMP.] ICROSSUS Erichson

A. ciliatus t()liviert - ON
unilineatus (.Say)

Genus CYCIIRAMUS Kugelann

Quadrifrons Blatchley
C. adustus !.richson * ON pO

Genus PALL,ODES Erichson

P. pallidus tl)alisot de Beauvois) - ON
silaceus Erichson

216
 Nitidulidae

Genus CYLLODES Erichson

C. biplagiatus LeConte MB

Subfamily MELIGETHINAE
Genus MELIGETHES Stephens
Key to North American species: Easton (1955)
M. atratus (Olivier)l _BC
M. canadensis Easton AKYK_BCAB_MBONPQ
M. cleominis Easton _BCABSKMB
M. dauricus Motschulsky* _BCABSKMB
californicus Reitter
moerens LeConte
rufimanus LeConte
M. nigrescens Stephens* _YK_BCAB_MBONPQ_NSPE
seminulum LeOonte
M. pinguis Horn
M. saevus LeConte AB-nrn------NF
_ONPQ_NSPE
M. simplipes Easton

Subfamily CRYPTARCHINAE
Genus PITYOPHAGUS Shuckard
P. rufipennis Horn - BC

Genus CRYPTARCHA Shuckard

(Subgenus CR\?TARCHA s. str.)
C. ampla Erichson - BC AB - MB ON PQ
(Subgenus LEPIARCHA Sharp)
C. concinna Melsheimer - BC AB - MB ON PQ
bella Reitter
liturata LeConte
piclo Melsheimer

Genus GLISCHROCHILUS Reitter

(Subgenus LIBRODOR Reitter)
G. fasciatus (Olivier) - BC MB ON PQ - NS
g eminat us (Melsheimer)
q uadr imaculoso (Melsheimer)
G. quadrisignatus (Say) - BC AB - MB ON PQ - NS NF
b ip unctatu s (Melsheimer)
bipustulatus (Melsheimer)
canadensis W.J. Brown
s exp u st ulatu s (Reitter)
sirnilis (Melsheimer)
G. sanguinolentus rubromaculatus PQ NB NS
(Reitter)
G. sanguinolentus sanguinolentus - BC AB ON PQ
(Olivier)
G. siepmanni W.J. Brown - BC AB SK MB ON PQ NB
(Subgenus GLISCHROCHILUS s. str.)
G. confluentus tsayt - ON PQ
G. lecontei W.J. Brown - BC IVIB
G. moratus W.J. Brown - BC AB SK MB ON PQ
G. vittatus (Say) AK BC AB - MB ON PQ
dejeanii (Kirby)
sepulcralis (Randall)

Subfamily CYB OCEPHALINAE

Genus CYBOCEPHALUS Erichson
C. californicus Horn _BC

217
Rhizophagi lae

Family RHIZOPIIAGIDAE rhizophagid beetles

Y. Bousquet

Twerrty-four species and subspecies of rhi- The taxonomy of the Nearctic species has been
zophagids are known to occur in Canada and Alaska. poorly studied, and most genera are in need of mod-
They are collected most commonly under the bark of ern revision. Crowson (1955) recognized four sub-
dead trees, in rotten logs, and in decaying vegetation. families, two of which (Rhizophaginae and
A few species live in ant nests, and some species of Monotominae) are found within the borders of
Monotomit are occasionally found in buildings. Very Canada and Alaska. Sen Gupta (1988) recently
little is known about their feeding habits. Members reviewed the world genera of the family.
of Rhizophaginae are mostly predaceous, feeding on
wood-boring beetles, especially scolytids; those of AK (3); vK (1); sC (15);AB (7); SK (3); MB (a); ON
Monotominae are apparently fungus feeders. (15);PQ (11);NB (a);NS (3);NF (5) . I (3)

Subfamilv RHIZOPHAGINAE
Genus RHIZOPHAGUS Herbst
Key to North American species: Bousquet (1990)
(Subgenus ;\NOMOPHAGUS Reitter)
R. brunneuri brunneus Horn
R. galbus Bousquet
R. pseudobrunneus Bousquet
_BC
AKBCAB_MB
AB ON PQ
v_
(Subgenus ITURHIZOPHAGUS M6quignon)
R. grouvellei M6quignon BC
(Subgenus RHIZOPHAGUS s. str.)
R. dimidiatus Mannerheirn AKYK_BCABON PQ NS NF
testaceits M6quignon
R. minutus minutus Mannerheim BC
R. minutus rotundicollis Bousquet -oN pe NF
R. paralleloi:ollis Gyllenhalf _BC_ON PQ NS NF
robustt,ts Schaeffer
R. remotus .LeConte AK BC AB MB ON PQ NS
luteus ,VI6qrignon
R. sayi Sche,effer
bipunc,tatus (Say)
-oN
R. sculpturu.tus Mannerheim BC

Subfamily MONOTOMINAE
Genus PYCI\OTOMINA Casey
P. cavicollis (Horn) ON PQ

Genus MACREUROPS Casey

M. longicoll:s (Horn) BC

Genus BAC'IRIDIUM LeConte

Crine Pascoe
B. ephippigerum (Gu6rin-M6neville)
B. striolaturn (Reitter)
BC -oN
_ONPQ
Genus HES PEROBAXNUS LeConte
H. abbrevia':us (Motschulsky) BC
rufipennis (LeConte)

Genus MOITIOTOMA Herbst

M. americana Aub6 ON PQ
M. arida Casey ON
M. fulvipes .Melsheimer --SCAB-nfe
opaca iJiegler
p ar allelocollis Motschulsky
subnigra Motschulsky

218
 Cucujidae

M. longicollis (Gyllenhal)f _BCABSK _ONPQ NB NF

M. parallela LeConte _BC_SK _ONPQ
M. picipes Herbstt _BCAB MB ON PQ NB NF
foueata LeConte
M. producta LeConte NB
M. quadraria Casey SK ON

Family CUCUJIDAE flat bark beetles

Y. Bousquet

The cucujids currently include 31 species and sub- Lawrence's concept (1982a) of the family is fol-
species in Canada and Alaska. The adults are usu- lowed here. Many authors rank the silvanids as a
ally found under the bark ofdead trees, in logs, and in distinct family. There is no modern taxonomic revi-
decaying plant matter. Some species apparently feed sion of Cucujidae, but some genera have recently
on fungi and decaying matter, others prey on insects. been revised.
Six of the species occurring in Canada, Cryptolestes
ferrugineus, C. pusillus, C. turcicus, Ahasuerus
aduena, Oryzaephilus surinqmensls, and O. mercator, AK (3);vK (1);NT (2); BC (15);AB (10); SK (9); MB
are associated with stored products; four ofthem are (17); oN (26); PQ (24); NB (5); NS (a); PE (1); LB
among the most serious pests in the country. (1);NF (5) . H (1);I (10)

Subfamily CUCUJINAE
Genus CUCUJUS Fabricius
C. clavipes clavipes Fabricius _MB oN PQ _NS NF
C. clavipes puniceus Mannerheim AK BC AB SK MB
subnitens Schaeffer

Genus PEDIACUS Shuckard

P. depressus (Herbst)f _BC
subglaber LeConte
P. fuscus Erichsonx AK YK NT BC AB SK MB ON PQ NB LB
planus LeConte
s ub car inatu s Mannerheim

Subfamily PASSANDRINAE
Genus CATOGENUS Westwood
C. rufus (Fabricius) oN PQ

Subfamily LAEMOPHLOEINAE
Genus PLACONOTUS Macleay
Siluanophloeus Sharp
Key to New World species: Thomas (1984)
P. zimmermanni (LeConte) MB ON PQ
apertus (Casey)
illustris (Casey)
sobrinus (Casey)

Genus LAXMOPHLOEUS Deiean

L. biguttatus (Say) - BC - SK MB ON PQ NB NS PE
b isig nat u s Gu6rin-M6neville
L. fasciatus Melsheimer MB ON PQ

Genus CHARAPHLOEUS Casey

C. adustus (LeContet MB oN PQ -NS
C. convexul us (LeConte t ON

2r9
Cucujidae

Genus CRY FIOLESTES Ganglbauer

Key to New World species: Thomas (1988)
C. ferrugine,us (Stephens)t - BC AB SK MB ON PQ NF
C. punctatus (Lecontet - ON
adumbratus Casey
extrica:lus (Casey)
geminctus (LeConte)
C. pusillus Sch<tnher"rtt ^ BC MB ON PQ
long ica r nis (Mannerheim)
minutu.s (Olivier)
paupel (Sharp)
testaceus (Stephens)
C. turcicus Grouvellet* - BC AB SK MB ON PQ

Genus LEPTOPHLOEUS Casey

L. alternans (Erichsontf - BC PQ

Subfamilv SILVANINAE
Tribl Uleiotini
Genus ULEIOTA Latreille
Bronte:i Fabricius
U. debilis rl,econter MB ON PQ
U. dubia duoia tFabriciust - ON PQ
denticulata Srrrith
U. dubia trr ncata Motschulsky BC

Genus DENDROPHAGUS Schtinherr

D. cygnaei ltlannerheim AK BC AB - MB ON PQ NF
americ:t nus Mannerherm
germat i Mannerheim
glaber LeConLe

Genus TEL.IIPHANUS Erichson

H et erotlr om ia Haldeman
T. velox Haldeman - ON PQ

Tlibe Silvanini
Genus SILVANUS Latreille
Key to spr:cies: Halstead (1975)
S. bidentatus (Fabricius)t - BC - ON PQ NB
S. muticus Siharp - BC - ON PQ
S. nitidulus LeConLe AB - MB
S. planatus Germar AB SK MB ON PQ
cognatt.ts LeConte
zimmer ma nni Gu6rin-M6neville

Genus CAT.I:[ARTOSILVANUS Grouve]le

KeY to sp,:cies: Halstead (1975)
C. imbellis(Leconte) NT ON PQ
C. fagi tGuerin-M6nevillet* - ON

Genus AHASVERUS des Gozis

A. advena (\Valtltf - BC AB SK MB ON PQ
musaeorum (Ziegler)
A. longulus'Blatchleyt - ON PQ NB
paruiceos (Casey)

Genus ORYZAXPHILUS Ganglbauer

Key to species: Halstead (1980)
O. mercator (Fauvel)t - BC AB SK MB ON PQ NB NS NF
O. surinamensis (Linnett - BC AB SK MB ON PQ
Genus NAU SIBIUS Redtenbacher
N. clavicorn.s (Kugelanntt - ON PQ

220
 Cryptophagidae

Family CRYPTOPIIAGIDAE silken fungus beetles

Y. Bousquet

Sixty-six species of cryptophagids are recorded fauna, only the genus Cryptophagus has been
in Canada and Alaska. These beetles are usually revised (Woodroffe and Coombs 1961). Many gen-
found on mold, on fungi, under bark, as well as in era previously included within the cryptophagids
decaying vegetation and nests of social have been transferred recently to Languriidae. The
Hymenoptera, birds, and mammals; one species, reader can expect many nomenclatural changes at
Myrmedophila americana, lives in ant nests. the specifrc level, as well as many new distribution
Adults and larvae of most species feed on mold and records, when the cryptophagid groups are revised.
decaying vegetation; those of Telmophilus species The generic classification of the Cryptophaginae
apparently feed on flower heads of certain aquatic used here follows Bousquet (1989).
plants. Several species of Cryptophagus are found
in stored products, where they feed on mold. AK (26); YK (1); NT (3); BC (47); AB (15); SK (7);
This group is poorly studied taxonomically. MB (12); oN (1e); PQ (17); NB (2); NS (3); LB (2);
Since Casey's (1900) revision of the North American NF (3) . H (11);I (8)

Subfamily TELMATOPHILINAE
Genus TELMATOPHILUS Heer
T. americanus LeConte BC AB SK MB ON PQ

Subfamily CRYPTOPHAGINAE
Key to North American genera: Bousquet (1989)
Tlibe Cryptophagini
Genus ANTHEROPHAGUS Latreille
A. convexulus LeConte PQ NB NS
A. ochraceus Melsheimer
oregon us Casey
- - - ec AB !r< r,re 3N PQNBNSPE_NF
A. pallidivestis Casey _BC
A. suturalis Meklin AK BC

Genus HENOTIDERUS Reitter

Crosimus Casey
Henoticoides Hatch
H. lorna (Hatch) AK BC
H. obesulus (Casey) AK_NT AB ON PQ NS

Genus HENOTICUS C.G. Thomson

Coniophthalmo Kangas
Glyptophorus Patk
H. californicus (Mannerheim) BC
inconspicuus (LeConte)
H. serratus (Gyllenhal) AK BC MB ON PQ
denticulatus (LeConte)

Genus PTERYNGIUM Reitter

P. crenatum (Gyilenhal)* BC PQ
malacum Casey

Genus SALEBIUS Casey

S. octodentatus (Mriklin) AK BC
sexdentatus Casey

Genus CRYPTOPHAGUS Herbst

C r y pto p hag i stes Crotch
Key to North American species: Woodroffe and Coombs (1961)
C. acutangulus (Gyllenhal)* AK BC AB MB ON PQ NF
prominens Casey
restrictus Casey

227
Cryptophag idae

C. bidentatus Maklin AK BC
puncta tis sim us MEikIin
C. cellaris (llcopoli)+ BC AB SK MB ON
angustztus Casey
rupimc,ntis Casey
C. confertus Casey* AK - NT BC AB - PQ
gelicus J.R. Sahlberg
arc han.
C. corticinus C.G. Thomson* AK BC - ON PQ
depressulus Casey
C. dentatus (Herbst)* AK BC
fumatus (Marsham)
q uad.ride ntat us Mannerheim
C. difficilis tlasey - BC AB - PQ
contractus Casey
C. distinguendus Sturm* - BC
heeni Casey
C. hebes Casey AK BC
lyraticollis Casey
o/ioszs Casey
C. histricus Casey - BC LB
C. lapponicLrs Gyllenhalx - BC
ualidus Kraatz
C. laticollis Lucast - BC AB
af,finic Sfrrm
cribricollis Casey
inscitu.s Casey
C. maximus Blake - BC
C. obsoletur Reittert - ON
C. peregrin rs Woodroffe & Coombs - BC
C. pilosus C yllenhalf - BC - - MB ON
cicatrir.osus Casey
infuscatus Casey
paruinodus Casey
plectrum Casey
C. porrectur' Casey AK BC
C. quadrihamatus Mtiklinx AK
plagiatus Poppius
C. saginatu; Sturml AK BC - ON
C. scanicus tlinn6)t - NF
C. scutellat rs Newmant - ON
C. setulosus, Sturm* - BC - ON PQ - LB
antennatus Casey
grossutus Casey
C. stromus Woodroffe & Coombs - SK MB
C. subluma us Kraatzl - BC AB SK - ON PQ
C. tubercult,sus Meklinx AK BC AB
bouldetanus Casey
breuipilis Casey
debilis LeConLe
instabtlis Bnrce
lepidut Casey
C. varus Woodroffe & Coombs - BC AB SK

Genus MYF,MEDOPHILA Bousquet

M. americana (LeConte) AK l'K - BC AB - MB - PQ
blzmi iBlaisdell)

Tribe Caenoscelini
Genus CAENOSCELIS C.G. Thomson
Macrodea Casey
C. anlenna]is (Caseyt AB - MB
C. basalis C asey PQ
C. Sahlbergt*
ferruginea tC.R. AK BC
C. parallela Casey AK BC

222
 Languriidae

Subfamily ATOMARIINAE
Tribe Atomariini
Genus EPHISTEMUS Stephens
(Subgenus EPHISTEMUS s. str.)
E. globulus (Paykull)* BC ON
apicalis LeOonte

Genus ATOMARIA Stephens

(Subgenus ANCHICERA C.G. Thomson)
A. aleutica Casey
A. apicalis Erichsonf
A. curtula Casey
1"_ _ _ _ _MB
ON
pumilio Casey pl:)
A. ephippiata Zimmermann NT BC AB SK MB ON
hesperica Casey
A. fulvipennis Mannerheim AK
A. kamtschatica Motschulsky* AK BC
fallax Casey
A. laetuia LeConte BC
A. nubipennis Casey BC
A. ochracea Zimmermann - - lr,fe ON Pe
lacustris Casey

- - - a*:a - -- - -
A. ovalis Casey
A. pusilla (Paykull)* AK sc -
A. saginata Casey
(Subgenus ATOMARIA s. str.)
(Agathengis des Gozis)
A. capitata (Casey) BC
A. fuscicollis Mannerheim* AK
A. lepidula Maklin AK
A. longipennis (Casey) BC
nigricollis (Casey)
A. melanica Hatch BC
A. planulata Maklin AK BC
A. quadricollis (Casey) BC
A. stricticollis (Casey) BC
A. subdentata (Casey) BC
A. vespertina Miiklin AK BC

Subfamily IfYPOC OPRINAE

Genus HYPOCOPRUS Motschulsky
H. tenuis Casey AB

Family LANGURIIDAE lizard beetles

J.M. Campbell

The family Languriidae, as recently defined, is larvae and adults of the other subfamilies are
widely distributed in all faunal regions but is best poorly known, but they probably feed on decaying
represented in tropical areas. OnIy seven species vegetation and fungal hyphae rather than on living
are known to occur in Canada, all but one of which plants (Lawrence and Vaurie 1983).
are found east of the Rocky Mountains. Larvae of Until recent papers by Sen Gupta and Crowson,
the subfamily Languriinae (Languriidae of previous this family was restricted to those genera now
authors) are stem borers of which one, the clover placed in Languriinae. Sen Gupta (1967) trans-
stern borer, Languria mozardi, is an economic pest ferred Toramus and allied genera from the family
in Canada. The feeding habits and biology of the Cryptophagidae to the subfamily Toraminae.

223
Languriidze

Subsequi:ntly (1968), he transferred Loberus to the of the family are reasonably well known, with most
tribe Lolrerini, both in the family Languriidae. The subfamilies having been revised since 19,[8 and the
family vl,as redefined, and the higher classification species cataloged by Lawrence and Vaurie (1983).
and genr:ra of the world were revised bv Sen Gupta
and Crorvson (1921). The North Ameiican BC (1); SK (1); MB (2); ON (6); PQ (5) . I (1)
"o".i""
Subfamily XENOSCELINAE
Tribe Loberini
Genus LOI}ERUS LeConte
L. impressr.rs LeConte ON PQ

Subfamily TORAMINAE
Key to species: Sen Gupta (1967)
Tribe Toramini
Genus TOF:AMUS Grouvelle
Tomarus LeConte
T. pulchellr.Ls (LeConte) oN PQ

Subfamily CRYPTOPHILINAE
Tribe Cryptophilini
Genus CR'IIPTOPHILUS Reitter
C. integer ( F{eer)f ON PQ

Subfamily LANGURIINAE
Tlibe Languriini
Key to North American species: Vaurie (1948)
Genus LAI\ GURIA Latreille
L. angustata (Paiisot de Beauvois) ON
pulchr t LeConte
uhlerit Horn
L. convexicollis Horn BC
in t er s t,t}ialis C asey
L. mozardi mozardi Latreille MB ON PQ
apicalis Motschulsky

Genus ACROPTEROXYS Gorham

A. gracilis (Newman) SK MB ON PQ
aztecana Casey
diuisa (}]orn)
inornata (Randall)
latreillei (LeConte)
nigrice o s (Motschulsky)
obscuro (Motschulsky)
pertenuis Casey
texana Schaeffer
thoracina Casey

224
 Erotylidae

Family EROTYLIDAE pleasing fungus beetles

J.M. Campbell

The family Erotylidae is found in all faunal The family is well known taxonomically; Boyle
tropical
regions of the world but primarily in moist, (1956) revised the North American species, and Sen
areas. Only 20 species are known to occur in Gupta (1969) worked out the higher classifrcation.
Canada. Adults are frequently found under bark or
under logs. Both larvae and adults feed on the eC (5);AB (6); SK (4); MB (5); ON (1a); PQ (1a); NB
fruiting bodies of higher fungr. (2); NS (1)

Subfamily DACNINAE
Genus DACNE Latreille
(Subgenus DACNE s. str.)
(Ezgis Paykull)
D. californica (Horn) _BC
cephalotes Casey
elongata Casey
Iaticollis Casey
uteana Casey
D. quadrimaculata (Say) MB ON PQ
(Subgenus XENODACNE Boyle)
D. picea LeConte _BC
Genus MEGAI-ODACNE Crotch
M. fasciata (Fabricius) -oNPQ
bifasciatus (Olivier)
M. heros (Say) -oNPQ
Subfamily TRIPLACINAE
Genus TRIPLAX Herbst
T. antica LeConte - BC AB SK
mormonalis Casey
T. californica LeConle - BC AB
T. cuneata Boyle - BC AB
T. dissimulator (Crotcht AB SK MB ON PQ
carri Casey
T. flavicollis Lacordaire - ON PQ
confinis LeConte
T. frosti Casey AB SK MB ON PQ NB
T. macra LeConte MB ON PQ
T. thoracica Say AB SK MB ON PQ NB
antennata Casey
breuicollis Casey
conuergens Casey
latiuscula Casey
melano p t era Lacordarre
obliqua Casey

Genus TRITOMA Fabricius

Cyrtotriplar Ctotch
T. biguttata tSay) PQ
basalis Lacordaire
T. humeralis Fabricius - ON PQ
ornata Casey
ruficeps LeConte
taeniata LeConte
uittata LeConLe
T. mimetica (Crotch) - ON PQ
T. pulchra Say - ON PQ - NS
basalis Melsheimer

225
Phalacrida,r

cincta Lacordafte
dimidiata Lacordaire
sellata Ktthnt
T. sanguinipennis (Sayl oN PQ
T. unicolor Say ON
unicolor Lacordaire

Genus ISCIIYRUS Lacordaire

Key to Ncrth American species: Skellay and Goodrich (1989)
I. quadripunctatus quadripunctatus oN PQ
(OIi rier)
alabartae Schaeffer
antediLisa Mader

Family PHALACRIDAE shining flower beetles

J.M. Campbell

The flamily Phalacridae comprises about 600 to use. Phalacridae and Ptiliidae are probably the
species around the world; only five have been most poorly known families of beetles in Canada,
reported to occur in Canada, but many other species and based on existing literature, a reliable specific
certainly occur. Adults are usually found on flow- identifrcation for most genera is not possitrle.
ers; larvae feed in flower heads of Compositae or on
spores aILd stomata of rusts and smuts.
Casey (1916) revised the North American
species o[ phalacrids, but this work is very difficult BC (3);MB (1); ON (3);PQ (2)

T?ibe Phalacrini
Genus PHIJ-ACRUS Paykull
P. penicillabus (Say) BC

Tribe Olibrini
Genus OLLBRUS Erichson
O. rufipes l.,eConte BC
tritus ,Jasey
O. semistri rtus LeConte ON
striatulus LeConte

Genus STII,BUS Seidlitz

Eustilbus Shatp
S. apicalis (Melsheimer) BC ON

Genus LITOCHROPUS Erichson

L. scalptus Casey PQ

Genus ACI-I-AMUS Sharp

A. pugetan-rs Casey MB ON PQ

226
 Cerylonidae

Family CERil ONIDAE cerylonid beetles

J.M. Campbell

OnIy about 400 species of Cerylonidae, a family feed on a plasmodium of slime molds (Slipinski
that consists of small to very small beetles, are 1988).
known around the world, of which seven have been Adults of this family are reasonably well known
recorded in Canada and Alaska. Most adults are taxonomically. Sen Gupta and Crowson (1973)
collected in leaf litter, in rotten wood, and under redefined the family and revised the higher classifi-
bark. Hypodacne punctata is known in the galleries cation. Lawrence and Stephan (1975) revised the
of Camponotus ants, and Murmidius oualis occurs North American species, and Lawrence (1982b) cat-
on stored products (Lawrence and Stephan 1975). aloged them.
Little is known about the feeding habits of the lar-
vae and adults, although some larvae are known to AK (1); NT (1); eC (3); AB (2); tt{B (1); ON (6); PQ
feed on fungal hyphae or spores, and some larvae (3);NB (1);NS (2) . I (1)

Subfamily ELIXESTINAE
Genus HYPODACNE LeConte
H. ounctata LeConte ON

Subfamily MURMIDIINAE
Genus MURMIDIUS Leach
M. ovalis (Beck)t AK ON

Genus MYCOCERINUS Slipinski

Mychocerus a:uct.
M. depressus (LeConte) ON

Subfamily CERYLONINAE
Genus PHILOTHERMUS Aub6
P. g)abriculus LeConte ON PQ

Genus MYCHOCERUS Erichson

Lapethus Casey
M. discretus (Casey) BC

Genus CERYLON Latreille

C. castaneum Say BC AB MB ON PQ NB NS
clypeale Casey
stricticutn Casey
C. unicolor(Zieglet) NT BC AB ON PQ NS
angustulum LeConte
simplex LeConte
syluaticum Casey

227
Corylophidae

Family CORYLOPHIDAE minute firngus beetles

J.M. Campbell

Members of Corylophidae, a small family of species have yet to be discovered. Casey (1900) pre-
about 4315 species, are found in all major faunal pared the last revision covering the North American
regions, but only 11 species are known to occur in species of Corylophidae. The classificatic,n adopted
Canada. Adults are usually collected in moldy leaf here is based on Paulian's (1950) reclassifrcation of
litter ancl piles of grass cuttings. Adults and larvae the Old World taxa.
are thought to feed on fungal spores.
The taxonomy of the Canadian corylophid
species is poorly known; reliable species identifrca- YK (1); NT (2); BC (7); AB (4); SK (2); MB (1); ON
tion is not possible at this time, and a number of (a); PQ (3); NB (1) . I (2)

Subfamily C ORYLOPHINAE
Tribe Orthoperini
Genus ORTHOPERUS Stephens
O. atomus (Gyllenhal)t _BC
O. scutellaris LeConte NT BC AB SK
lucidus Casey
O. suturalis LeConte ON

Tlibe Rypobiini
Genus GRC)NE\IUS Casey
G. fuscicorrLis Casey oN PQ

Subfamily SERICODERINAE
Tribe Sericoderini
Genus SERICODERUS Steohens
S. lateralis (Gyllenhal)t BC PQ

Subfamily SACIINAE
Tlibe Saciini
Genus MOLAMBA Casey
M. biguttat,a (LeConte) BC AB
M. fasciata (Say) oN pe N"
s u b fa s : iata (Gu6rin-M6nevilIe)
M. obesa Casey BC

Genus SACIIUM LeConte

Parmu.lus Harold
S. lugubre l,eConte
S. montanum Casey
YK NT BC
_BC
AB SK yuoN_
Genus ARTHROLIPS Wollaston
A. decolor (LeConte)
califor nicus (Matthews) AB

228
 Coccinellidae

Family COCCINELLIDAE ladybird beetles

J. McNamara

Coccinellidae comprises 162 species and sub- monograph by Gordon (1985). Only some of the
species in Canada and Alaska. Many species are most recent references on coccinellids have been
introduced in North America, some of these pur- listed in the checklist. Good regional works on
posely for biological control. Most coccinellid adults Coccinellidae include those of Chapin (1974) for
and larvae are beneficial, because they are preda- Louisiana, Belicek (1976) for western Canada and
tors of aphids, scale insects, and mites. However, in Alaska, and Larochelle (1979) for Quebec.
the absence of preferred food, many species feed on
the immature stages of other insects. The species of
Psyllobora feed on fungi, and the only species of
Epilachna that occurs in Canada feeds on plants of
the family Leguminaceae. /J{(26); YK (31); NT (35); BC (9a);aB (83); SK (70);
The taxonomy of the North American MB (56); ON (83); PQ (77);NB (37); NS (25);pE (6);
Coccinellidae is well known. thanks to the recent LB (9);NF (13) . H (2); I (8)

Subfamily STICHOLOTIDINAE
Tribe Microweisini
Genus MICROWEISEA Cockerell
Epismilia Cockerell
Pseudoweisea Schwarz
SmiliaWeise
Key to species: Gordon (1970o)
M. misella (LeContet BC AB SK MB ON PQ NB NF
Genus COCCIDOPHILUS Brethes
Cryptoweisea Gordon
Key to North American species: Gordon 0970a)
C. marginatus { Leconte) BC AB SK MB ON PQ NB

Tlibe Serangiini
Genus DELPHASTUS Casey
Key to species: Gordon (19706)
D. pusillus (LeConte) ON PQ
puncticollis (LeConte)
sonoricus Casey

Subfamily SCYMNINAE
Tlibe Stethorini
Genus STETHORUS Weise
Key to New World species: Gordon and Chapin (1983)
S. punctillum Weiset BC -oN
S. punctum punctum (LeConte) ABSK-ONPQ-NS
S. punctum picipes Casey BC
6reuis Casev

Tlibe Scymnini
Key to North American species: Gordon (1976)
Genus DIDION Casey
D. longulum Casey YK NT BC AB SK PQ
occiduum (Casey)
paruiceps Casey
D. nanum (LeConte) ON PQ
D. punctatum (Meisheimer) _NTBCABSKMB ON PQ NB_PE NF

229
Coccinellidae

Genus SCYMNUS Kugelann

(Subgenus {ICYMNUS s. str.)
S. americar..us Mulsant oN PQ
S. apicanus pseudapicanus Gordon AB SK MB
boreal;s Gordon
S. caurinus Horn BC AB
aluticctllis Casey
S. fenderi Ilalkin BC
S. indianensis Weise ON
rustict;s Casey
S. nebulosus LeConte BC
harneli Hatch
infuscc:tus Boheman
phelps;'.i Crotch
S. opaculus Horn
S. paracanus linearis Gordon - - - :t ffi 3[ nre- - - - - - -
(Subgenus PULLUS Mulsant)
S. abbreviai;us LeConte ON PQ
S. aquilona::ius Gordon _ABSK
S. ardelio Eorn BC AB
apacheanus Casey
decipeins Casey
sanctu; Casey
S. aridus Cr,rsey BC
S. brullei Mulsant SK MB ON PQ ivs---
diuisut Casey
laureniicus Casey
lodi Stt}rr
subaen:.eus Casey
S. calaveras Casey BC
saginalus Casey
strenuus Casey
stygicu:.s Casey
tenuiue stis Casey
S.
S.
carri Gordon
caudalis LeConte
agricola Casey
_ _ _ rSf nre oN pe
indutus Casey
natche zianus Casey
S. cervicalir Mulsant
S. compar ( asey
-oN
impun:tusWingo
-oN
uicksbt.rgicus Casey
S. coniferarum Crotch BC
S. fraternus LeConte ON PQ
dentipes FaIl
haemo;rhous LeOonte
S. humboldt,i Casey _BC
S. impexus Mulsant'l _BC lln NS_-
S. impletus Gordon _BC
S. iowensis Casey _BC_SKMB oN PQ NB
collarit Melsheimer
melsheimeri Weise
S. kansanus Casey -ONPQ
S. lacustris LeConte YI( NT BC AB SX MB ON PQ NB NS
cultratus Wingo
S. marginic, >ll is Mannerheim BC AB ON PQ
borealis Hatch
californicus Boheman
desertcrum Casey
S. postpictus Casey BC AB
S. puncticollis LeConte :" -oNpe--
S. rubricaudus Casey _ONPQ
canteri.us Casey
chromopygus Casey
texanus Casey
S. securus J. Chapin oN_
S. solidus Casey nc
blaisde'lli Casey

230
 Coccineilidae

S. suturalis Thunbergt PQ
tahoensis Casey D/r
S.
S. tenebrosus Mulsant NT AB_ -oN PQ
S. uncus Wingo AB SK MB ON
S. utahensis Gordon AB

Genus NEPHUS Mulsant

(Subgenus NEPHUS s. str.)
N. ornatus naviculatus (Casey) AK BC AB SK MB ON PQ NF
h.incaidi (Hatch)
N. ornatus ornatus (LeConte) _ONPQ NB NS
frosti (Casey)
sanguinifer (Casey)
(Subgenus TURBOSCYMNUS Gordon)
N. georgei (Weise) AK YK NT BC AB ON PQ NF
bisignatus (Horlr)
(Subgenus SCYMNOBIUS Casey)
N. flavifrons (Melsheimer) ON PQ
bioculatus (Mulsant)
guttiger (Mulsant)
Iudouictanus (Casey)
marginellus (Mulsant)
N. intrusus (Horn) ON
inops (Casey)
N. sordidus (Horn) SK MB ON
intrusoides (Hatch)

Genus DIOMUS Mulsant

D. amabilis (LeConte) PQ
dulcis (Casey)
emertoni (Casey)
D. debilis (LeConte) AB SK ON PQ
aeger (Casey)
minutissimus (Casey)
molliculus (Casey)
pusio (Casey)
D. liebecki (Horn) MB ON
adulans (Casey)
ohioensis (Stehr)
D. terminatus (Say) ON
femoralis (LeConte)
partitus (Casey)

Tlibe Hyperaspini
Genus HYPERASPIDIUS Crotch
H. comparatus Casey BC
juniperus Nunenmacher
H. hercules Belicek _AB
H. insignis Casey -AB
_AB SK
H. mimus Casey
carri Nunenmacher
c oloradens is Nunenmacher

H. vittigerus (LeConte) YK_BC AB SK

Genus HYPERASPIS Redtenbacher

Oxynychus LeConte
H. bigeminata (Randall) -oN PQ NB
gueri Mulsant
H. binotata (Say) SK MB ON PQ NB
affinis (Randall)
Leucopsis Melsheimer
normata (Say)
paspalis Watson
H. borealis Dobzhansky YK BC AB
eloli Nutting
obscura Malkin

237
 Coccinellidae

schuhi }{atch
simuloides Hatch
H. consimilis LeConte NT BC AB SK MB PQ NB
canadensis Dobzhansky
H. conviva rlasey SK MB ON PQ
congressis Watson
insolen.s Casey
H. disconotata Mulsant AB _MBON PQ NB_
H. dissolutzL dissoluta Crotch BC
coloraclana Casey
nupta,iasey
H. fastidiosr Casey _BCAB
s e pte nt rion is Dobzhansky

H. inflexa Casey _BCAB SK MB PQ

serena Casey
H. jasperentis Belicek NT_AB
H. lateralis Mulsant _BCAB SK
flammula Nunenmacher
idoe N rnenmacher
laeuipennis Casey
montartica Casey
nig rocc. uda Dobzhansky
omissa Casey
wellmani Nunen.macher
H. moerens (LeConte)
H. octavia C asey
H. oregona )obzhansky
_
BC
_ sx_ 3NF3
AB
NB
lanei E atch
H. postica I eConte BC AB SK
angust tla Casey
elliptict Casey
essigl n{alkin
H. pratensis LeConte PQ
H. proba (Say) SK MB ON PQ
trinifer Casey
H. quadrivittata LeConte BC AB SK
tetrane;tra Casey
H. signata signata (Olivier) SK
taedato. LeConte
H. simulatrix Dobzhansky BC AB SK
H. troglodyt:s Mulsant _MB pe lls
discretc: LeConte
nouasct'liae E.A. Chapin
H. undulata (Say) BC SK MB ON PQ NB
elegans Mulsant
guttifer a Weise
maculilbra Melsheimer

Genus BRA(IHIACANTHA Dejean

B. albifrons iSay) AB SK MB
pacifica Casey
B. decempustulata (Melsheimer)
stellata Casey
-oN PQ NB NS
troglod^rtes Mulsant
B. dentipes (Fabricius) ON
separatiz Leng
socialis Casey
B. quadripunctata quadripunctata ON PQ
Melsheimer
basalis Melsheimer
confusa Mulsant
diuersa Mulsant
B. rotunda Ci'ordon ON PQ
B. ursina (Fabricius) AB SK MB ON PQ NB NS
congruens Casey
B. uteella CzLsey BC
fenyesi Leng
lezgl Nunenmacher
sonorana Casey

232
 Coccinellidae

Subfamily CHILOCORINAE
Tribe Chilocorini
Key to genera: Chapin (1965)

Genus BRUMOIDES E.A. Chapin

B. histrio (Fall) - BC
paruicollis (Casey)
B. septentrionis septentrionis AK YK NT BC AB SK MB ON
(Weise)
desertorutn (Casey)
neuadensis (Leng)
ouoideus (Casey)
B. septentrionis davisi (Leng) - ON PQ

Genus EXOCHOMUS Redtenbacher

E. aethiops rBlandi AB SK
mormonicus Casey
E. californicus Casey - BC

Genus CHILOCORUS Leach

C. hexacyclus Smith AB SK
C. stigma (Say) AB SK MB ON PQ NB NS
biuulnerus Mulsant
C. tricyclus Smith - BC

Subfamily COCCIDULINAE
Tlibe Coccidulini
Genus COCCIDULA Kugelann
C. lepida LeConte AK YK NT BC AB SK MB ON PQ NB
occidentalis Horn
reitteri Dodge
suturalis Reitter
szlzrolis Weise

Genus RHYZOBIUS Stephens

Lindorus Casey
R. lophanthae (Blaisdell)t - BC

Subfamily COCCINELLINAE
Tribe Coccinellini
Genus ANISOSTICTA Dejean
A. bitriangularis (Say) AK YK NT BC AB SK MB ON PQ NB NS _ LB NF
irregularis Weise
multig uttata (Randall)
A. borealis Timberlake AKYKNT_MB_PQ
Genus MACRONAEMIA Casey
M. episcopalis (Kirby) AK YK NT BC AB SK MB ON PQ

Genus PARANAEMIA Casey

P. vittigera (Mannerheim) _rn_
similis Casey

Genus COLEOMEGILLA Timberlake

Megilla Mulsant
C. maculata lengi Timberlake -oNPQ
Genus CERATOMEGILLA Crotch
C. ulkei Crotch AK YK NT BC AB ON
poruo (Watson)

233
Coccinellidz,.e

Genus HIPI'ODAMIA Dejean

H. americana Crotch NT BC AB SK MB ON PQ
H. apicalis Casey _BC
lezgl Johnson
nig r om a c ula ta (Nunenmacher)
tricolor Nunenmacher
H. arctica (lichneider)x AK YK NT BC _PQ LB
H. caseyi Jc,hnson _BCAB
H. convergens Gu6rin-M6neville _BCABSK MB ON PQ NB
juncta Jasey
modestz Melsheimer
obsoleta Crotch
pratico!.a Mulsant
H. expurgata Casey _YK_BCABSK
H. falcigera Crotch AK YK NT BC AB SK MB PQ
albertana Ca.sey
cottlei (.Nwenmacher)
H. glacialis glacialis (Fabricius) NT BC AB SK MB ON PQ
H. glacialis lecontei Muisant NT_ABSKMB
abduce.zs Casey
defecta Johnson
h op p in,;1 i Nunenmacher
macher.zie E.A. Chapin
p se u do;1 lacialis Johnson
H. lunatomi rculata dobzhanskyi YK BC
E.A. Chapin
H. moesta nLoesta LeOonte _BC
H. moesta b':wditchi Johnson _BCABSK MB
H. oregonensis Crotch _BCAB
cockere.'li Johnson
d,ispar'Jasey
liliputo.na Casey
puncticollis Casey
H. parenthesis (Say) BC AB SK MB ON PQ NB NS
albomacula Fitch
appror,:mata Fitch
confluezta Fitch
connatu Fitch
discopttnctata Fitc}'
insulata Fitch
linearis Fitch
liturico!.lis Fitch
nimia I'itc}r
permac"ifrons Fitch
triangu.laris Fitch
tridens (l\ttby)
tridentifrons Fitch
H. quindecirnmaculata Mulsant _SK_ON
H. quinques ignata quinquesignata AK YK NT BC AB SK MB ON PQ NB_PE
(KirLY)
coccineu Casey
leporina Mulsant
mulsanti LeConte
pugetar'.a Casey
quadra,"ia Casey
subsiru.lis Casey
uteana Casey
uernis Oasey
H. quinquesignata ambigua LeConte BC
obliqua Casey
punctulata LeConte
H. sinuata crotchi Casey NT BC AB SK MB
disjunc;la Timberlake
fontinaiis Casey
straminea E.A. Chapin
H. sinuata spuria LeConte AK YK _BC
cornpler Casey
H. tredecimpunctata tibialis (Say) AK YK NT BC AB SK MB ON PQ NB NS PE LB NF

234
 Coccineliidae

H. variegata (Goeze)t
H. washingtoni Timberlake BC 9*PQ_
Genus ANATIS Mulsant
Key to species: Watson (1976)
A. Iabiculata (Say) MB ON PQ NB
canadensis Provancher
coseyl Westcott
q uinde c i m p u n c t a ta (Oliv iet)
A. lecontei Casey BC AB
A. mali (Say) AKYKNT BC AB SK MB ON PQ NB NS PE NF
borealis Belicek
signaticollis Mulsant
A. rathvoni (LeConte) BC AB SK

Genus MYZIA Mulsant

Neomysia Casey
Paramysia Reitter
M. interrupta (Casey) BC AB
coseyi (Timberlake)
M. pullata (Say) YK AB SK MB ON PQ NB NS LB
montana (Casey)
nolons (Randall)
randalli (Casey)
M. subvittata (Mulsant) BC AB
hornii Crotch
oregona (Casey)

Genus CAI-VIA Mulsant

Anisocaluia Crotch
C. quatuordecimguttata (Linn6)* AK YK NT BC SK MB ON PQ NB NS LB NF
bicordifera (Casey)
cardisce (Randall)
comple*a Nunenmacher
elliptica (Casey)
hesperica (Crotch)
incarnata (Kirby)
lacustris (Casey)
obliqua (Randall)
postplagiata (Casey)
quadrisignata (Casey)
sicordi (Nunenmacher)
simllis (Randall)
uniformis (Casey)
uancouueri (Casey)
uictoriana (Casey)

Genus ADAI,IA Mulsant

A. bipunctata ilinn6)* AK YK NT BC AB SK MB ON PQ NB NS LB NF
annectans (Crotch)
bioculata (Say)
coloradensis Casey
disjuncta (Randall)
duplicata Leng
frigido (Schneider)
humeralis (Say)
ludouicae Mulsant
(LeConte)
mel anop I eur a
ocellata Johnson
opthalmica Mulsant
ornatella Casey
ouipennis Casey
postica Johnson
sexpustulata Johnson
transuersalis Casey

Genus COCCINELLA Linn6

Key to North American species: Brown (7962b)

235
Coccinellidi,'e

C. alta W.J. Brown BC AB

sutura'.is Casey
C. californica Mannerheim AK BC AB
melanc'collis Johnson
C. fulgida V/atson
C. hieroglyphica humboldtiensis
Nunenmacher
TY NT BC
BC
ON PQ

C. hieroglyphica kirbyi Crotch _YK NT BC AB SK MB ON PQ NB NS

tricuspis Kitby
C. hieroglyphica mannerheimi AK YK NT MB
Muklant
C.johnsoni Casey AX BC
C. monticola Mulsant AK_NTBCAB SK MB ON PQ Nn f,IS -
alutacea Casey
bigutta.ta Johnson
confluenta Johnson
impres';a Casey
lacustris LeConte
neuadica Casey
postica Johnson
sellica Johnson
C. novemno:ata Herbst BC AB SK MB ON PQ
confluenta Fitch
conjunt:ta Fitch
degenet'Casey
diuisict'llis Fitch
francis,:ana Mulsant
inaequulis Fitc}:'
oregontt Casey
paruan,,aculato Fitch
C. prolongata prolongata Crotch BC A.B
C. septempr..nctata Linn6J _BCABSKMBONPQ NB NS
C. transvers oguttata richardsoni AK YK NT BC AB SK MB ON PQ NB NS in Ne
W.J. Brown
interrulta Fitdn
quinquznotata I{ttby
zetterstzdti Madet
C. trifasciat,r perplexa Mulsant AK YK NT BC AB SK MB ON PQ NB NS _LBNF
C. trifasciatl subversa LeConte _BC
barda T,eConte
euginii Mtisant
juliana Mulsant
praedicta Mad.et
C. undecimp,unctata undecimpunctata BC ON PQ NB NS PE NF
Linn5t

Genus PRO )YLAEA Mulsant

P. quatuord,:cimpunctata (Linn6)t ON PQ NB

Genus CYC.i,ONEDA Crotch

C. munda (Say) MB ON PQ
C. polita Casey _BCABSK
flauaT'lr'berlake
Genus OLLr\ Casey
O. v-nigrum (Mulsant) BC A3 SK MB ON
abdominalis (Say)
arizonae Casey
fenestra'.lis Casey
minuta Casey
plagiaht Casey
soyi (Clotch)
semiluri.aris Johnson
sobrina Casey

Genus NEO.FIARMONIA Crotch

Agrabiu Casey
Harmottiaspis Casey
Neoharnonia Casey

236
 Endomychidae

N. venusta venusta (Melsheimer) ON

centralis Casey
dissimila Blatchley
fattigi B\atc}:.ley
notulata (Mulsant)

Genus MULSANTTNA Weise

Pseudocleis Casey
Key to species: Chapin (1985)
M. hudsonica (Casey) AK YK NT BC AB SK MB ON PQ NB NS PE LB
M. picta (Randall) _YK _BCABSKMBONPQNBNS
blanchardi (Johnson)
c on cinnata (Melsheimer)

contert(r (Mulsant)
impictipennis (Weise)
minor (Casey)
nubilata (Casey)

Tribe Psylloborini
Genus PSYLLOBORA Dejean
P. borealis Casey BC
deficiens Casey
P. vigintimaculata (Say) AK YK NT BC AB SK MB ON PQ NB NS NF
obsoleta Casey
separata Casey
taedata LeConte

Subfamily EPILACHNINAE
Key to New World species: Gordon (1975)
Tribe Epilachnini
Genus EPILACHNA Dejean
E. varivestis Mulsant oN PQ
ceruina Mulsant
corrupta Mulsant
cuprea Cockerell
genuina Mulsant
juncta Johnson
maculiuentris Bland
toweri Johnson

Family ENDOMYCHIDAE handsome ftrngus beetles

J.M. Campbell

The family Endomychidae is found in most taxonomically well known. However, the family has
regions of the world, but the majority of species never been revised for North America. Strohecker
occur in tropical areas. The range of 15 species (1953) reviewed the genera of the world and pub-
extends northward into Canada. Adults and larvae lished a catalog of the North American species
may be collected on soft fungi, in leaf litter, or (1986). Hatch (1962) treated the genera and species
under bark with fungal growth. One species, that occur in the Pacific Northwest. A modern revi-
Mycetaea subterranea (M. hirta in most publica- sion of the North American species of this family is
tions), is a pest of stored products. Most species needed.
feed on fungr.
Because of their moderately large size and often NT (1); BC (7); AB (2); SK (2); MB (5); ON (10); PQ
bright coloration, many North American species are (8);NB (2);NS (6);PE (1);NF (2) ' I(2)

237
Endomychidae

Subfamily MYCETAEINAE
Tribe Mycetaeini
Genus SYMBIOTES Redtenbacher
S. duryi Blatchley
Iacustris Casey
-oN
oblong,ts Casey
pilosus Casey
waltoni Dury
S. gibberosr;.s .(Lucas)t -oN
duryiWalton
montanus Casey

Genus MYCETAXA Stephens

M. subtenanea (Fabricius)f _BC_ONPQ_NSPE_NF
fumata Stephens
hirta (Ivlarcham)

Tribe Leiestini
Key to North American species: Blaisdell (1931)
Genus RHANIDEA Strohecker
Rhanis LeConte
R. unicolor Zieglerr - ON PQ
apicalis (Melsheimer)
haemo,"rhoidalis (Gu6rin-M6nevi11e)

Genus STE'IHORHANIS Blaisdell

S. borealis ]]laisdell - BC

Genus PHYMAPHORA Newman

P. californica Horn - BC
P. pulchella Newman MB ON PQ - NS
cr a s s icor nis ( Melsheimer)
p u nc t i o I I i s (Zi e gl er')
r:'

Subfamily STENOTARSINAE
TYibe Stenotarsini
Genus DA]riAI Reiche
D. testacea iZiegler) _ONPQ_NS
fer r ug i ne a (Gu6rin-M6neville )
lzleo (lvlelsheimer)
piloso i.Melsheimer)

Subfamily ELIMORPHINAE
Tribe Lycoperdinini
Genus LYC'OPERDINA Latreille
(Subgenus (]OLGIA Mulsant)
L. fermgine a LeConte - BC AB SK MB ON PQ - NS
angulata (Say)
g lab rala Gu6rin-M6neville

TYibe Mycetinini
Genus APHORISTA Gorham
A. laeta (LeConte)
A. vittata (Fabricius)
distincta (Say)
---:"--meo*pe-----
lineata (Olivier)
ouipenizis Casey

238
 Lathridiidae

Genus MYCETINA Mulsant

M. idahoensis Fall BC
M. perpulchra (Newman) MB ON PQ NB NS
q uadrig uttata (Melsheimer)

Subfamily ENDOMYCHINAE
Tribe Endomychini
Genus ENDOMYCHUS Panzer
E. biguttatus Say _NTSKMB ON PQ NBNS_ NF
E. limbatus (Horn) BC AB
endomychoides (Fall)

Family LATHRIDIIDAE minute brown scavenger beetles

Y. Bousquet

In Canada and Alaska 55 species of lathridiids Most of the genera, particularly those of the sub-
have been recorded. These species are usually family Corticariinae, are in need of modern revision.
found on foliage, in rotten wood, under logs, and The generic classifrcation of the Lathridiinae used
under the bark ofdead trees, as well as in fungi and here follows that of Walkley (1952).
nests of social Hymenoptera, birds, and mammals.
A number of species, including Cartodere constricta
and Aridius nodifer, are regularly found in stored
products, especially those that are damp and moldy.
Adults and larvae feed on fungr, particularly molds. AK (19);YK (5);NT (6); BC (3a); AB (9); SK (8); MB
Lathridiidae is another family of Cucujoidea (17); ON (22); PQ (22); NB (6); NS (8); PE (1); LB
that has been poorly studied in North America. (1);NF (a) . H (6);I (15)

Subfamily I"ATHRIDIINAE
Genus DIENERELLA Reitter
Microgramme Walkley
D. arga (Reitter)t oN PQ
D. cordicollis (Mannerheim) AI{
D. costulata (Reitter)t _MBONPQ
D. frliformis ( Gyllenhal)f SK MB NB PE
D. filum (Aub6)t ---ec- _MBONPQ
D. rufrcollis (Marsham)t _BC ON PQ _NS NF
p ulicar ia (Melsheimer)

Genus ENICMUS C.G. Thomson

E. aterrimus Motschulsky oN PQ
ferrugineus Belon
opaculus LeConte
E. cordatus Belon -BC
E. fictus FaIl AK NT BC AB ix ms o* Pe
E. mimus Fail AK NT BC AB SK MB
E. nitens FalI
E. tenuicornis LeConte BC
y_ yuoNpe_ _ _ _ _
laticollis LeConte

Genus THES Semenov

T. bergrothi (Reitter)f BC SK MB NB NS
tricarinatus (W.J. Brown)

Genus LATHRIDIUS Herbst

Conithassa C.G. Thomson
L. consimilis Mannerheim* AK
par allelocollis Mannerheim
L. crenatus LeConte BC

239
Lathridiidae

L. hirtus Gyllenhalf _BC MB ON PQ

L. minutus (Linn6)+ _BCAB SK MB
porcatus Herbst
reflextls LeConte
L. nigritus (Fall) _BC
L. protensi :oliis Mannerheim AK BC
quadr :collis Mannerheim
sobrinus Mannerheim
suspectus (Fall)

Genus STEPHOSTETHUS LeOonte

S. breviciavis (FalI)
-oNPQ NB NS
S. cinnamopterus (Mannerheim) AK _BC
S. costicollis (LeConte) AK _BC
armat:zlus (Fall)
S. curtulus (Mannerheim) AK
S. lardariuir (DeGeer)f NF
S. liratus (LeConte) AK BC AB SK MB ON PQ wS
carinil'er (Fa\\')
trilobatus (Motschulsky)
S. montants (Fall) AK YK NT BC MB PQ
laeuior (W .J. Brown)

Genus ADISTEMIA Fall

A. watsoni lWollaston)t ON PQ

Genus CAIi,TODERE C.G. Thomson

Coninomus C.G. Thomson
C. constrict,a (Gyllenhal)t BC AB SK MB ON PQ NB NF
sculpti.lis (LeConte)

Genus ARI )IUS Motschulsky

A. nodifer ( Westwood)f BC MB ON PQ

Subfamily CORTICARIINAE
Genus COI,,TICARIA Marsham
C. canalicu ata Mannerheim AK
C. columbia. FalI _BC
C. dentigera LeConte _BC ON PQ
C. dentiventris Poppius* AK YK
C. fenestralis (Linn6)* AX BC
deleta Mannetlr'eim
denticulata I{trby
kirbyi LeConte
obtusa LeConte
rugulo.;a LeConte
C. ferrrrginea Marshamx
C. Iinearis (Paykull)x
C. orbicollis Mannerheimx
AK
AK
AK YK
NT
NT
_BC
MB
MB
PQ
PQ- _ _ luIu
munsteri Stand
C. parallela Fall YK AB
C. pubescers Gyllenhal-i
grosso LeConte
_BC -on
C. salpingoides Motschulsky _BC
C. serrata (Paykull)f _BC
octodentata (Say)
C. serricollis LeConte ON
C. spinulosa Mannerheim AK
C. valida Fall
C. varicolor Fall
-nc
_BC
Genus MEL,ANOPHTHALMA Motschulsky
M. americana (Mannerheim) BC ON PQ
distinguenda auct. amer.
morsa',LeConte)
pusilla (Melsheimer)
rufula (.LeConte)

240
 Byturidae

M. pumila (LeConte) _BC_ON

M. villosa (Zimmermann) _BC
Genus CORTICARINA Reitter
C. alberta (Fall) AB
C. cavicollis (Mannerheim) -BC-ONPQ-NS
C. exigua (Mannerheim) AK
C. fuscula (Gyllenhal) AK l'K NT BC MB ON PQ
americana auct. amer.
compt(r (LeConte)
expansa (LeConte)
grata (LeConte)
C. herbivagans (LeConte) _BC
leuis (LeConte)
similata a:uct.

Genus CORTINICARA Johnson

C. gibbosa (Herbst)t _BCABSKMBONPQNBNS
Genus CORTILENA Motschulsky
C. picta (LeConte) -oNPQNBNS

Family BIPI{YLLIDAE false skin beetles

J.M. Campbell

Only four species of Biphyllidae are known to This family is poorly known taxonomically, and
occur in North America, one of which, Diplocoelus the North American species have not been revised.
brunneus, is found in eastern Canada. Little is
known about the habits of this species; adults are ON (1);PQ (1);NB (1)
found in fungus or under bark.

Tlibe Biphyllini
Genus DIPLOCOELUS Gu6rin-M6neville
Marginus LeConte
D. brunneus (LeConte) - oN PQ NB

Family BYTURIDAE fouitworm beetles

J.M. Campbell

Two species of Byturidae are found in North sion by Springer and Goodrich (1983) of the North
America of which one, Byturus unicolor, the rasp- American species of the family recognized only one,
berry fruitworm, is common and widely distributed somewhat variable species.
across Canada. Adults ofthis species feed on leaves
and flowers, and the larvae feed on the receptacle of
the red raspberry, Rubus idaeus.
Byturus unicolor has been known under a vari- aK (1); yK (1); NT (1); BC (1); AB (1); SK (1); MB
ety of Latin and common names, but a recent revi- (1); ON (1); PQ (1); NB (1);NS (1);NF (1)

Tribe Byturini
Genus BYTURUS Latreille
Horticola Lindemann
B. unicolor Say AK YK NT BC AB SK MB ON PQ NB NS NF
americanus Gemminger & Harold

247
Mycetophagidae

baker;'.Barber
breuicollis Casey
punctatus Casey
rubi Eiarber
sordictus Barber

Superfamily Tenebrionoidea
Family MYCETOPIHGIDAE hairy fungus beetles
Y. Bousquet

Of the 26 species of mycetophagids recorded Parsons (I975b) has revised the mycetophagids
from Nc,rth America, 15 are known to occur in of theNearcticfauna.
Canada. These beetles are found mainly in decay-
ing vegetable matter, in fungi, and under the bark
of rotten logs. A few species, such as Typhaea ster-
corea, are occasionally found in buildings. As the
Latin name suggests, members of this family are BC (6);aB (4); SK (2); MB (7); ON (10); PQ (12); NB
mycetophagous. (2);NS (1); PE (1) . I (1)

Genus MY,IETOPHAGUS Hellwig

(Subgenus MYCETOPHAGUS s. str.)
M. californicus Horn - BC
prouer,,sis Casey
M. flexuosrrs Say MB ON PQ
M. punctat us Say MB ON PQ
M. serrulat,us (Caseyt MB ON PQ
pictus (.Casey)
quadralius Casey
subderzressus (Casey)
tribaheatus Casey
(Subgenus ILENDUS Casey)
M. distinctus Hatch - BC AB SK MB
M. pluripunctatus LeConte AB - MB ON PQ
(Subgenus PARILENDUS Casey)
M. quadriguttatus P.W.J. Miillerf - BC AB SK MB ON PQ NB
bimac ulatus Melsheimer
bipust.ulatus Melsheimer
b isig n at us Melsheirner
uariegatus C.R. Sahlberg
(Subgenus GRATUSUS Casey)
M. plurigu,tatus LeConte - BC
franciscanus Van Dyke
M. tenuilasciatus Horn - BC AB ON PQ
notalulus ( Casey,

Genus LITARGUS Erichson

(Subgenus AJ-ITARGUS Casey)
L. balteatus LeConte - ON PQ
infulatus LeConte
tr(l,nsuersus LeConte
(Subgenus PARALITARGUS Casey)
L. didesmus (Sayt PQ
asperulus Casey
(Subgenus LITARGUS s. str.)
L. sexpuncbatus tSay) - ON PQ
obsolescens Casev

242
 Tetratomidae

(Subgenus TILARGUS Casey)

L. tetraspilotus LeConte MB ON PQ

Genus TYPHAEA Stephens

T. stercorea (Linn6)f - BC - ON PQ NB NS PE
fumata (Linn6)
Genus THRIMOLUS Casey
T. minutus Casey PQ
duryi Casey

FamilyTETRATOMIDAE tetratomidbeetles
L. LeSage

The family Tetratomidae includes nine species on beetles, such as Blatchley (1910), Chagnon and
in Canada and Alaska. Both adults and larvae live Robert (1962), Hatch (1965), and Arnett (1968).
in the fruiting bodies of higher fungi and are myce- Young (1976) recently transferred the genus
tophagous. Pectotoma, which had been previously placed in this
Previously considered a subfamily of family, to Scraptiidae. The survey of the
Melandryidae, the tetratomids were raised to family Tetratomidae of Quebec by Chantal (1985) includes
status by Crowson (1955). This ranking is accepted genera placed here in Melandryidae.
by most recent authors, including Miyatake (1960)
and Lawrence (1982a). There is no recent revision
ofthefamilyinNorthAmerica. Generaandspecies aK(1); YK(1); nC(2); AB(1);SK(1); ON(5);PQ
can be identifred tentatively by using general books (7); NB (1);NF (1) . I (1)

Subfamily TETRATOMINAE
Genus TETRATOMA Fabricius
T. concolorleConte - YK - BC AB SK
T. truncorum LeConte - ON PQ

Genus ABSTRULIA Casey

A. tesselata (Melsheimer) - ON PQ
maculata Casey
A. variegata Casey PQ NF

Genus EUPISENUS Casey

E. elongatus (LeCon[e) AK BC

Genus PISENUS Casey

P. humeralis (Kirbyt - ON PQ
concolor (Kirby)
ruficornis (LeConte)

Subfamily PENTHINAE
Genus PENTHE Newman
Pytocis Castelnau
P. obliquata (Fabricius) -oNPQ
P. pimelia (Fabricius)
anglica (Gmelin)
-oNPQNB
funerea Newman
lesueuri (Castelnau)
morio (Fabricius)

Genus MALLODRYA Horn

M. subaenea Horn prl

243
Ciidae

Family CIIDAE minute tree-fungus beetles

J. McNamarq,

The family Ciidae, which consists of small bee- Lawrence (1971) revised the North American
tles, comprises 29 species in Canada and Alaska. species and also published a catalog of the species
The larvae and adults of nearly all species feed on (1982c).
fungal h'7phae comprising the mycelia and especially
the fruiting bodies of wood rotting Basidiomycetes.
Several l;pecies are found under bark, in dead wood, eK (6); xT (6); BC (19); AB (10); SK (5); MB (14);
or in the galleries ofbark beetles. ON (16);PQ (17);NB (5);NS (6);NF (6) ' I (1)

Subfamily CIINAE
Tlibe Ciini
Genus CIS Latreille
Xestocis Casey
C. americanus Mannerheim AK BC AB MB ON PQ NB NS NF
forsteri Abdullah
frosti Dury
hatch, Lawtence
insolerc (Casey)
minot (Hatch)
nitidts (Hatch)
oweni (.Hatch)
serricollis Drty
strigulosus (Hatch)
C. angustus Hatch _BC
C. biarmatus Mannerheim AK _BC
C.
C.
castlei (Dury)
creberrimus Melli6
-o* PQ
C. ephippiatus Mannerheim AK -nc _:* PQ
moznt'ttei (Dtry)
C. fuscipes Melli6 NT BC AB SK MB ON PQ _NS NF
atripennis Me1li6,
caroli:zae Casey
cheurolatii Melli6
dubius Melli6
impressus Casey
pallens Casey
C. horriduius Casey NT BC MB ON NS
mormonicus Casey
C. hystriculus Casey -BC
_BCABSKMBONPQ
C. levettei (Casey) NB NS NF
C. maritimus (Hatch) _BCMB
C. pistorius Casey NT-ABSKMBONPQ
C. striolatus Casey NTBCAB_MBONPQ NS--
fraternus Casey
macilentus Casey
C. subtilis Mel1i6 PQ
confusus Blatchley
julichiDrry
wenzeli Dtry
C. tridentatus Mannerheim AK BC
ednae (I].atch)
reflexus (Hatch)

Genus ENNEARTHRON Melli6

E. spencen (Hatch)t -BC
Genus DOLICHOCIS Dury
D. indistinctus Hatch -BC PQ
NB-
D. manitoba Dury NT BC AB MB ON PQ

244
 Melandryidae

Genus ORTHOCIS Casey

O. punctatus (Mel1i6) AK NT BC AB MB ON PQ NF
aterrimus Casey

Genus STRIGOCIS Dury

S. opacicollis Dury ON

Genus HADRAULE C.G. Thomson

Maphoca Casey
H. blaisdelli (Casey) BC
H. elongatula (Gyllenhal)t NB

Genus PLESIOCIS Casey

P. cribrum Casey BC AB MB PQ

Genus CERACIS Me11i6

Key to North American species: Lawrence (1967)
C. sallei (Melliel oN PQ

- - - r,rs3Npe- - - -
C. singularis t Duryt
C. thoracicornis t Zieglert
bifoueata Dtry
-
laminifrons (Casey)
mellyi (.Mel1i6)
oblonga (Blatchley)
picea (Casey)
pumicdta (Meili6)
unicolor (Casey)

Genus SULCACIS Dury

S. curtula tCasey) BC AB SK MB PQ
criddlei (Drry)
cylindrica (Dury)
montana (Casey)
nigra Dury
sorror (Casey)

Genus MALACOCIS Gorham

Brachycis Casey
M. brevicollis (Casey) MB ON PQ NF

Subfamily OROPHIINAE
Tlibe Orophiini
Genus OCTOTEMNUS Melli6
O. laevis Casey AK BC AB SK MB ON PQ NB NS NF
denudatus Casey

Family MELANDRYIDAE false darkling beetles

L. LeSage

Melandryids are common throughout the world beetles work inconsistently. Up-to-date revisions at
and include about 100 genera and 1200 species, of the species level are badly needed. Chantal (1985)
which at least 55 occur in Canada and Alaska. keyed out the species of Hallomeninae and
Adults live under bark, in fungi, or in flowers. Eustrophinae from Quebec. At this time, no expert is
Larvae are carnivorous or feed on decaying plant working on the family in North America. The classifi-
material; they live under bark, in dead logi, or in cation used here corresponds to the arrangement sug-
stumps; a few are found in leaf litter, but detailed gested by J.F. Lawrence (personal communication).
life-history studies, such as that of Hoebeke and
McCabe (1977), are seldom published. aK (4);YK (2);NT (3); BC (27); AB (10); SK (5); MB
The taxonomy of the family is in a state of disar- (e); ON (27);PQ (42); NB (11); NS (10); LB (3); NF
ray at all levels. Generic keys from general books on (13) . H(3);I(2)

245
Melandryidre

Subfamily IIALLOMENINAE
Genus HALLOMENUS Panzer
H. binotatu,; (Quenselti PQ
H. debilis L,:Conte PQ
ziger Fialdeman
H. punctulatus LeConte AK BC - ON PQ
basalis Mannerheim
H. scapulars Melsheimer - BC - ON PQ
Iuridus Haldeman
ruficor,zis Melsheimer
H. serricornis LeConte - BC AB ON PQ

Subfamily EUSTROPHINAE
Genus EUS'IROPHINUS Seidlitz
E.bicolor(Fabriciusr - ON PQ
indistittctu s LeConte)
(

E. confinis t LeContet - BC - SK MB ON PQ

Genus EUSTROPHUS ll I iger

E. tomentosus Say - BC - ON PQ
niger Nlelsheimer

Genus SYNSTROPHUS Seidlitz

S. repandus t Horn t - BC MB ON PQ

Genus HOL,OSTROPHUS Horn

H. bifasciat rs (Sayt - ON PQ
q uadr i:nac ulalzs (Melsheimer)
H. discolor l{orn PQ
H. impressi':ollis tl-eConter - BC

Subfamilv MELANDRYINAE
Trile Orchesiini
Genus ORCHESIA Latreille
Key to North American species: Lalibert6 (1966)
O. castanea (Melsheimer) - BC MB ON PQ - NS NF
O. cultrifonnis Laiibert6 - NT SK MB - PQ - NS
O. ornata Horn - BC
O. ovata La ibertti ON PQ
Genus MIC ROSCAPHA LeConte
M. clavicon.is LeConte PQ

Genus LEDERIA Reitter

Crioscctpha Flotn
L. arctica (llorn) AK BC

Tlibe Melandryini
Genus EMNIESA Newman
E. connectens Newman - ON PQ - NS - LB NF
maculcrta LeConte
E. labiata (,(iay) - ON PQ NB - NF
E. stacesmi-,hi Hatch - BC
E. testacea leeperi Malkin - BC

Genus MET,ANDRYA Fabrici us

NeoemtnesaHatch
M. blackmani Hatch PQ
M. striata Say - BC AB SK - ON PQ NB
bicolor Melsheimer

246
 Melandryidae

diuersa Mulsant
excauata Haldeman
thoracica Melsheimer

Genus PHRYGANOPHILUS C.R. Sahlberg

P. collaris LeConte NT BC AB - MB ON PQ NB

Tlibe Hypulini
Genus ZILORA Mulsant
Key to North American species: Mank (1938)
Z. canadensis Hausen PQ
Z. hispida LeConte - YK - BC AB ON PQ NF
Z.nudaProvancher PQ
Z. occidentalis Mank - BC

Genus HYPTILUS Paykul.

H. concolor (LeConte) PQ

Genus MICROTONUS LeConte

M. sericans LeConte - '' ? -
Genus SYMPHORA LeConte
Trotomma LeConte
S. flavicollis (Haldeman) MB ON PQ - NS
S. rugosa (Haldeman) - ON PQ - NS

Genus PROTHALPIA LeOonte

P. holmbergi (Mannerheim)* AK BC
fulrninans (LeConte)
P. undata LeConte - ON PQ - NS NF

Tlibe Serropalpini
Genus AMBLYCTIS LeConte
A. praeses LeConte PQ

Genus XYLITA Paykull

X. livida (C.R. Sahlberg)* _YK_BCAB_PQNBNS_LBNF
X. laevigata (Hellenius)* -BCAB-MBONPQNB_NF
decolorata (Randall)

Genus SPILOTUS LeConte

S. quadripustulatus (Melsheimer) PQ

Genus RUSHIA Forel

Carebara LeConte
R. californica FaIl _BC
californica Schaeffer
schaefferi Csiki

Genus SCOTOCHROIDES Mank

S. antennatus Mank PQ

Genus SCOTOCHROA LeConte

Key to species: Mank (1939b)
S. atra LeConte
S. basalis LeConte
S. buprestoides (Kirby) _-_:"AB__;.::il___Ni
Genus ENCHODES LeConte
E. sericea (Haldeman) _BCABSKMBONPQ

247
 Mordellida:

Genus SEF,ROPAIPUS Hellenius

Key to North American species: Mank (1939a)
_BC
S. coxalis Nlank
S. obsoletur; Haldeman
perfoueatus Mank
T_ _BC ON
-PQ
PQ NB NS NF

S. substriatus Haldeman NT BC MB ON PQ NB LB NF
Genus MYSTAXUS Kugelann
M. simulator (Newman) ON PQ NB NS NF
trifasc iatus (Melsheimer)

Genus DIR,IA-EA Fabricius

D. liturata '.LeConte) AB SK ON PQ NB
quadrtmacularo (Say)
D. horiei Hatch BC

Genus PHLOEOTRYA Stephens

P. vaudoueri Mulsantt PQ NS
fusca (.LeConte)

Genus ABDERA Stephens

A. frrma Ha.tch BC

Genus ANIIIOXYA Mulsant

A. glaucula LeConte ON

Subfamily OSPFIYINAE
Tlibe Osphyini
Genus OSPFfYA Illiger
Nothut Olivier
O. varians (LeConte) oN PQ

Family MORDELLIDAE tumbling flower beetles

J. McNamara

Severrty species and subspecies of mordellids genera of the world; and Bright (1986) pr:Lblished a
have beer recorded in Canada. The adults are small, catalog of the Nearctic species. Neverlheless, a
humpbacked beetles, and their most distinctive fea- modern revision would be useful. Members of
ture is their long, pointed abdomen. They are com- Anaspini, which used to be included within
monly found on umbelliferous flowers, on which they Mordellidae, are now in the family Scraptirdae.
feed. Larvae are carnivorous or feed on plant mate-
rial. Sorne are found boring in rotten wood, others
are leaf and stem miners, but very few are known.
Liljetrlad (1945) revised the North American YK (1); NT (6); BC (18); AB (10); SK (12); MB (37);
species oI'Mordellidae; Ermisch (1951) treated the ON (49);PQ (29);NB (14);NS (7);PE (1)

Tribe Mordellini
Genus TOMOXIA Costa
T. borealis ( LeConte)

_ =_:"Y_;;lNifrilrs_ _ -
T. inclusa Lr:Conte
T. lineella LeConte
T. serval (Sz,y)
T. triloba (Say)
T. undulata (Melsheimer)

Genus MORDELLA Linne

Sphalera LeConte

248
 Mordellidae

M. albosuturalis Liljeblad BC
M. atrata atrata Melsheimer NT BC AB SK MB ON PQ NB
M. atrata lecontei Csiki -ONPQNB NS
itorata LeConte
M. brevistylis Liljeblad BC
M. grandis Liljeblad BC
M. hubbsi Liljeblad BC
M. marginata lineata Melsheimer
_NT BC
-o*---
MB ON PQ NB NS
M. marginata marginata Melsheimer
M. melaena Germar _NT AB MB ON PQ
M. quadripunctata (Say) BC AB SK MB ON
Genus GLIPA LeConte
G. bidentata (Say) _ONPQ
G. octopunctata (Fabricius) -oN
G. oculata (Say) -oN
jouialis (LeConte)

Genus CONALIA Mulsant & Rey

C. melanops Ray ON PQ

Tribe Mordellistenrnr
Genus MORDELLISTENA Costa
M. aemula LeConte _MB
M. aethiops Smith
M. andreae ancilla LeConte - Y sK MBoN - - - -
--
-
uarians LeConte
M. andreae andreae LeConte ON
grammica LeOonte
M. andreae ustulata LeConte NT MB
M. arida Leconte ON

'_ _:'I_ illgN;:

rutila Liljeblad
M. aspersa (Melsheimer)
M. bicinctella LeConte
M. cervicalis LeConte NB
auricorna Helmuth
erilis Liljeblad
picicornis LeConte
M. convicta LeConte
M. discolor (Melsheimer) _ _ _ yuSNPQ NS

cinereofasciata Smith
wolcotti Liljeblad
M. divisa LeConte MB
M. downesi Hatch BC
M. errans Fall PQ
M. frosti Liljeblad MB-PQ NB
M. fulvicollis (Melsheimer) -oN
_ONPQ
M. fuscata (Melsheimer)
M. fuscipennis (Melsheimer) -oNPQ NS PE
pityptera LeConte
M. gigas Liljeblad BC _ONPQ
M. guttulata Helmuth -oN
MB ON
M. hebraica LeConte
M. impatiens LeConte MB ON
M. incommunis Liljeblad AB MB
M. infima LeConte SK MB ON NB
tiara Ray
M. liljebladi Ermisch BC
egregia Liljeblad
SK MB ON
M. limbalis (Melsheimer)
M. liturata (Melsheimer) ON
PQ
_ft
M. lutea (Melsheimer) ON
M. marginalis (Say) _NT -MBONPQ NB
M. morula LeConte BC SK
M. nigricans Melsheimer _MBONPQ
nigenima Helmuth
M. nunenmacheri Liljeblad BC

249
Rhipiphoridae

M. ornata (trfelsheimer) SK MB ON PQ
decoreLa LeConte
pectora,tis (LeConte)
M. pallipes fimith MB ON
M. picilabris Heimuth MB
M. pustulatr,r (Melsheimer)
M. quadrinobata Liljeblad
-:-ecAB i* MB ON PQ

_BCAB -oNPQ
M. scapulais (Say) SK MB ON PQ NB NS
biplagiuta Helmuth
M. semiusta LeConte MB
M. sericans .Fall SK MB
M. sexnotata Dury
falli Cslki
-oN
festiua Tall
M. smithi Dury _MBON
M. suturella Helmuth _MB
M. syntaenia Liljeblad _MBNB
M. testacea.Blatchley ON
M. tosta LeConte _MBONPQNB
M. trifasciata (Say) SK MB ON PQ NB
amica L,eConte
lepidula LeConte
nigricollis Helmuth
M. unicolor LeConte NT MB
M. vilis (LeClonte) BC MB ON

Genus GLIPOSTENODA Ermisch

G. ambusta iLeConte) BC ON PQ
diuersa (Ray)

Family RHIPIPHORIDAE wedge-shaped beetles

J.M. Campbell

The family Rhipiphoridae contains about 400 Rhipiphoru.s, the largest North American genus of
species, of which 10 are known to occur in Canada. the family, are poorly known and cannot be reliably
Larvae of the subfamily Rhipiphorinae are parasitic determined. The records cited below, based on spec-
on the larvae of bees and wasps; the frrst instars are imens identifred with the aid of Rivnay's revision,
endoparasites and the last ones ectoparasites. are not all reliable. When revised, a number of addi-
Adults are often found on flowers, where females tional species and provincial records will be
lay their eggs. recorded.
Rivnay (L929) revised the North American
species o.l this family. However, the species of eC (3);AB (2); SK (2); ON (5);PQ (2)

Subfamily PELECOTOMINAE
Genus PELECOTOMA Fischer von Waldheim
P. flavipes Melsheimer ON PQ

Subfamily RHIPIPHORINAE
Genus MACIIOSIAGON Hentz
M. cruenturr (Germar) SK
M. limbatun. (Fabricius) -o*-
Genus RHIPIPHORUS Bosc d'Antic
Dorthesia Say
Myodes Latreille
Myodites Latreille
R. californicus (LeConte)
R. columbiar:,us W.J. Brown
R. fasciatus (Say)

250
 Colydiidae

R. mutchieri Rivnay
R. nevadicus (LeConte)
R. walshii (LeConte)
R. zeschii (LeConte)

Family COLYDIIDAE cylindrical bark beetles

Y. Bousquet

Only 21 species of colydiids are known to occur The North American species of colydiids are
in Canada. They are usually found under bark, in usually divided into two subfamilies: Colydiinae,
galleries of wood-boring larvae, and in fungi. which includes most of the species, and
Adults and larvae of many species are predators of Bothriderinae. Stephan (1989) has recently revised
wood-boring insects; others feed on rotten wood or the North American species of the family.
fungr; larvae of some species are parasitic on vari-
ous groups such as Scolytidae, Platypodidae, NT (2);sC (10); AB (3); SK (1); MB (1); ON (10);PQ
Cerambycidae, and Buprestidae. (5); NB (1); NF (1) ' I (1)

Subfamily COLYDIINAE
T?ibe Synchitini
Genus SYNCHITA Hellwig
S. fuliginosa Melsheimer oN PQ
nigripennis LeConte

Genus ENDEITOMA Sharp

E. granulata (Say) PQ
floridana Casey
Genus EUCICONES Sharp
E. marginalis (Melsheimer) ON
latus Casey

Genus BITOMA Herbst

Synchytodes Ctotch
B. crenata (Fabricius)t
B. ornata (LeConte)
B. quadriguttata (Say)
trinotata Casey
_y__:x;:__ __ _

zimmer manni Gu6rin-M6neville

Genus NAMUNARIA Reitter

Coxelus Horn
N. guttulatus (LeConte) ON
N. pacificus (Horn) BC

Genus LASCONOTUS LeConte

L. borealis Horn NT _SK MB ON PQ NB_ NF
L. complex LeConte -BC AB
L. intricatus Kraus NT BC AB
h.rausiHatc}r
L. planipennis Kraus -BC
_BC
L. tuberculatus Kraus
L. vegrandis Horn -BC
apicalis Casey
schwarzi lkans

Tlibe Colydiini
Genus AULONIIIM Erichson
Key to North American species: Kingsolver & White (1967)
A. longum LeOonte - BC

25t
Zopheridae

Genus COLYDIUM Fabricrus

C. lineola Sry - BC - ON
longiut,culum Say

Tribe Pycnomerini
Genus PYC NOMERUS Erichson
Endeclus LeConte
Penthetispa Pascoe
P. reflexus tSay) - ON
nitidus (LeConte)

Subfamily BOTHRIDERINAE
Tlibe Deretaphrini
Genus DERETAPHRUS Newman
D. oregonensis Horn - BC

Tribe Teredini
Genus OXYLAEMUS Erichson
Redistes Casey
O. californir us Crotch - BC

Tribe Bothriderini
Genus BOT :IRIDERES Erichson
B. cryptus Stephan -oN
B. geminatus (Say) -oN

Family ZOPHERIDAE
J.M. Campbell

The f amily Zopheridae comprises about 125 Lawrence (1979). However, a revision of t,he North
species, <lf which only one is known to occur in American species is needed.
Canada. .{dults live under bark, and the larvae feed
on rotten wood or on the fruitingbodies offungi.
The higher classification and relationships of
zopherids are well known, thanks to Doyen and eC (1); AB (1); ON (1); PQ (1); NB (1); NF (1)

Subfamily ZOPHERINAE
Genus PHEI,LOPSIS LeConte
P. obcordata (Kirby) - BC AB ON PQ NB - NF
porcata (LeConte)

252
 Tenebrionidae

Family TENEBRIONIDAE darkling beetles

Y. Bousquet and J.M. Campbell

The family Tenebrionidae comprises over 1200 lagrioid lineage, which is probably the sister group
species in North America, but more than 8O7o of of the tenebrionids, has been given family status;
them are restricted to the southwestern United these species lack defensive glands. Although the
States. Only 135 species occur within the Canadian limits of the family are now fairly stable, its higher
and Alaskan borders. Many tenebrionids live in classifrcation is still poorly understood. Numerous
semiarid areas and deserts, but in Canada most changes have been proposed recently in the classifi-
species are found in rotten wood, in fungr, under logs cation of the family (see Doyen and Tschinkel 1982),
and stones, and under the bark of dead trees. A and more are expected; the alleculids, for example,
number of species, such as those of Tenebrio, are probably polyphyletic in origin. Rolf L. Aalbu
Gnatocerus, Tribolium, and Polorus, are stored- (personal communication) has provided the classifr-
product pests, some of which are the most important cation used here. There is no recent revision of the
in the country. Adults and larvae feed on plants or Nearctic Tenebrionidae, and most genera that occur
plant material, including decaying vegetation, seeds, in Canada have not been revised. Identifrcation is
roots, cereals and their products, and fungr. often difficult, especially in large groups like
The concept of the family used for the checklist Eleodes.
is similar to that of the tenebrionoid branch of
Doyen and Tschinkel (1982). Adults of all species AK (3); YK (3); NT (8); BC (69); AB (32); SK (22);
have paired defensive glands, with reservoirs open- MB (32); oN (70); PQ (63); NB (16);NS (1a); PE (5);
ing between the seventh and eighth sternites. The LB (2); NF (7) . H (1);I (11)

Subfamily PHRENAPATINAE
Tribe Penetini
Genus CLAMORIS des Gozis
Phthora Mulsant
C. americana (Horn) BC

Subfamily TENTYRIINAE
Tlibe Eurymetopini
Genus STERIPHANUS Casey
S. subopacus (Horn)
alutaceus Casey

Tribe Anepsiini
Genus ANEPSIUS LeConte
A. montanus Casey AB

Tribe Coniontini
Genus CONIONTIS Eschscholtz
Coelotaxis Hotn
Coniontellus Casey
Coniontides Casey
C. ovalis LeConte BC
acerba Casey
alutacea Casey
anita Casey
arida Casey
breuiuscula Casey
coruina Casey
inepta Casey
oblita Casey
okanagani Boddy
parilis Casey

253
Tenebrionldae

punchtta Casey
regulc,ris Casey
sculptipennis Casey
uteanu Casey
uanco.zueri Casey
weidtt Casey
Tribe Coelini
Genus COItrLUS Eschscholtz
Coelotnorpha Casey
Pseudocoelus Casey
Keyto species: Doyen (1976)
C. ciliatus Eschscholtz _BC
ampli,:ollis Casey
arenarius Casey
curtulus Casey
debilis Casey
latus ()asey
longuLus Casey
obscurus Casey
scolopax Casey
sternalis Casey

Tlibe Asidini
Genus ASIDOPSIS Casey
A. opaca tSayt AB
A. polita (Sayt AB SK

Genus MICjROSCHATIA Solier

Acrosthatia WlTke
Pycno;'totida Casey
P. inaequa.is tl,eConte) - ,
diuersa (Casey)

Genus GL\?IASIDA Casey

G. sordida Leconre) AB

Genus PEL,ECYPHORUS Solier

P. densicollis Horn - BC
corrosus Casey

Subfamily DIAPERINAE
Tribe Phaleriini
Genus PHALEROMELA Reitter
Nelite:t LeConte
P. globosa (LeConte) _BC
P. picta (Mrnnerheim) AK
P. variegat r Triplehorn _BCAB
picta (Horn)

TYibe Nitionini
Genus SCAPHIDEMA Redtenbacher
Key to North American species: Triplehorn (1965)
S. aeneolum (LeConte) - YK - BC AB - MB ON PQ NB LB

Tlibe Rhipidandrini
Genus RHIPIDANDRUS LeConte
R. flabellicornis (Sturm) - ON PQ
paradorus (Palisot de Beauvois)

254
 Tenebrionidae

Tlibe Bolitophagini
Genus BOLITOTHERUS Candbze
Phellidius LeConte
B. cornutus (Panzer) _SKMBONPQNB
bifurcus (Fabricius)

Genus ELEATES Casey

E. depressus (Randall) MB ON PQ NB
tetraopes (Newman)
E. explanatus Casey _BC
Genus BOLITOPHAGUS Illiger
B. corticola Say _ONPQ

Tribe Diaperini
Key to North American genera: Doyen (1984)
Genus ADELINA Dejean
Doliema Pascoe
Schedarosus Reitter
D. plana rFabricius) - ON
depressa (Erichson)
lecontei (I].orn)
scidaria (Reitter)

Genus CYNAEUS LeConte

C. angustus tleConte) - BC AB SK MB ON PQ
opoczs Champion

Genus GNATOCERUS Thunberg

CerandriaLrcas
Echocerus Hotn
Sicinus Champion
(Subgenus GNATOCERUS s. str.)
G. cornutus (Fabricius)f - BC MB ON PQ - NS
maxillaris (PaIisot de Beauvois)

Genus ALPHITOPHAGUS Stephens

Phyletes Redtenbacher
P hy lethus Redtenbacher
Key to North American species: Triplehorn (1965)
A. bifasciatus {Sayr - SK MB ON PQ

Genus DIAPERIS O.F. Mriller

Key to North American species: Triplehorn (1965)
D. maculata Olivier MB ON PQ NB
floridana Blatchley
hydactina Fabricius
hydni Fabricitts
suturalis Chevrolat

Genus NEOMIDA Latreille

Arrhenoplita IGrby
Euoplus LeConte
Hoplocephala Castelnau & Brull6
Key to North American species: Triplehorn (1965)
N. bicornis (Fabricius) - ON PQ - NS
cornigera (Fabricius)
g r acilis (Motschulsky)
uirescens (Castelnau & Brull6)
u ir idipennis (Fabricius)

metallica (PaIisot de Beauvois)

Genus PLATYDEMA Castelnau & Brull6

Histeropsis Chevrolat
Key to North American species: Triplehorn (1965)

255
Tenebrionidae

Brull6 MB ON PQ
tBC AB -
P. americanum Castelnau & - NB
P. excavatLm {Say) - - ON PQ
fraten;.utn Chevrolat
nigrih".m Motschulsky
paruulum Casey
tubercllatum Castelnau & Brull6
P. neglectun Triplehorn - BC
P. oregonense LeConte - BC
P. ruficorne (Sturm) - ON PQ
analeHald.eman
opaculum Casey
rufum (Melsheimer)
rufiuen.tre Castelnau & Bru116
P. subcosta:um Castelnau & Brulle - ON PQ
clypea,um Haldeman
o b lo ng ulum Motschulsky
P. teleops 'l riplehorn - ON PQ

Genus PENTAPHYLLUS Dejean

Key to North American species: Triplehorn (1965)
P. pallidus LeConte - ON PQ

Subfamilv HIPOPHLOEINAE
Trib'e Hypophloeini
Genus CORTICEUS Piller & Mitterpacher
Hy pop,llaeus Fabricius
Key to North American species: Triplehorn (1990)
C. tMelsheimert
parallelr- s - ON PQ
C. praetern.issus(Fa11) AK YK NT BC AB SK MB ON PQ
C. subopacr.s (Wallis) NT BC AB - MB ON - NF
C. substrialus (LeConteJ - BC
C. tenuis (t.eOonte) - BC AB ON PQ NB
minor'Wallis)
o cc ider: talis (Wallis )
C. thoracicr.s tMelsheimerJ - ON
piliger (LeConte)

Subfamilv OPATRINAE
Tribe Eleodini
Genus ELEODES Eschscholtz
(Subgenus Iv{ELANELEODES Blaisdell)
E. humeral.s LeConte - BC
latiusczlus Walker
E. obsoletus obsoletus (Say) NT - AB SK MB ON
E. tricostatus (Say) NT - AB SK MB ? -
alternatus Kitby
planetus Solier:
robustus LeOonte
(Subgenus LITHELEODES Blaisdell)
E. extricatus convexicollis AB SK
Blaisdell
E. extricatus extricatus (Say) NT BC AB SK
E. parvulus Blaisdell - BC
E. vandykei modificatus Blaisdell - BC
(Subgenus PROMUS LeConte)
E. opacus (Slayt NT - AB SK MB
(Subgenus IILEODES s. str.)
E. hispilabris connexus LeConte - BC
binotatus Walker
imitabilis Blaisdell
immundus Blaisdell
loeuis Illaisdell

256
 Tenebrionidae

E. hispilabris hispilabris (Say) - BC AB SK MB

lecontei Gemminger
sulcatus LeConte
E. obscurus sulcipennis Mannerheim - BC
aratus LeConte
conjunctus Walker
conuexicollis Walker
(Subgenus BLAPYLIS Horn)
(Am p hido ra E schscholtz )
(Eleodopsis Blaisdell)
E. indentatus Blaisdell - BC
E. novovermculus Boddy - BC AB
E. nunenmacheri nunenmacheri - BC
Blaisdell
E. pimelioides Mannerheim - BC
oregonus Blaisdell
patruelis Blaisdell
subligatus LeConte
uerruculus Blaisdell
uiator LeConte
E. productus Mannerheim - BC
uariolosus Blaisdell
E. rolundipennis LeConte - BC
strictus LeConte
(Subgenus METABLAPYLIS Blaisdell)
E. nigrinus LeConte - BC
(Subgenus CRATIDUS LeConte)
E. osculans (LeConte) - BC

Genus EMBAPHION Say

E. muricatum Say AB
concduum LeConte

Tlibe Opatrini
Genus EPHALUS LeConte
E. latimanus (LeConte) NS

Genus BLAPSTINUS Dejean

Aspidius Mulsant & Rey
Heteropus Castelnau
Lodinus Mulsant & Rey
B. discolor Horn _BC
oregonensis Casey
B. intermixtus Casey _BC
B. metallicus (Fabricius) -BCABSKMBONPQ_NS
interruptus (Say)
luridus Mulsant & Rev
B. pratensis LeConte
B. pulverulentus Mannerheim
-oN
- BC
c alifor nicu s Motschulsky
B. substriatus Charnpion NT BC AB SK MB ON PQ

Genus GONOCEPHAIUM Chevrolat

G. bilineatum Walkert _BC
latifrons (LeConte)

Tribe Lllomini
Genus ULOMA Latreille
U. impressa Melsheimer -oN
U. longula LeOonte -BC

257
Tenebrionidae

T?ibe Helopini
Genus TAIiPELA Bates
Larnperos AIIard
T. micans (Fabricius) - ON PQ

Genus HELOPS Fabricius

H. californicus Mannerheim - BC
H. convexulus LeConte - BC
montanus LeConte
H. perniter.rs LeConte - BC
H. punctipennis LeConte - BC
punctatus Gemminger
H. regulus Blaisdell - BC AB

Subfamilv TENEBRIONINAE
rriue Tlibotiini
Genus TRIBOLIUM Macleay
Margu,s Dejean
Key to species: Hinton (1948)
(Subgenus TRIBOLIUM s. str.)
T. audax Halstead - BC AB SK MB ON PQ
T. castaneum (Herbst)* - BC AB SK MB ON PQ PE
ferrugineum (Dejean)
T. confusu:n Jacquelin du Val* - BC - SK MB ON PQ NF
T. destruct:r Uyttenboogaart* - BC AB SK MB ON PQ - NS PE - NF
T. madens Charpentiertt - NB NS
(Subgenus APIIANOTUS LeConte)
T. brevicor;re (LeConte.t - BC

Genus BIL'S Mulsant

B. estriatu,; (LeContet - BC AB - PQ

Genus PAI,ORUS Mulsant

Key to species: Halstead (1967)
(Subgenus PALORUS s. str.)
P. ratzeburgii (Wissman)t - BC - ON PQ - NS
(Subgenus CIRCOMUS Fleischer)
P. subdeprr:ssus (Wollastont* MB

Genus LAT'HETICUS Waterhouse

L. oryzae Waterhouse* MB - PQ NB

Tribe Tenebrionini
Genus ALF'HITOBIUS Stephens
A. diaperinus (Panzer)f - BC AB SK MB ON PQ NB NS PE - NF
A. laevigat.Ls Fabriciusi - BC - ON
plcezs (Olivier)
picipes (Panzer)

Genus MEI|ACLISA Jacquelin du Val

M. marginalis Horn - BC

Genus CENTRONOPUS Solier

M ene chide s Motschul sky
Pyres Champion
Scotobates Fiye
Key to New World species: Spilman (1962)
C. calcarat:s tFabricius) - ON PQ - NS
aeneus (DeGeer)
carolirt ensis (Palisot de Beauvois)
reflexus (Say)

258
 Tenebrionidae

Genus TENEBRIO Linn6

T. molitor Linn6t BC AB SK MB ON PQ NB NS NF
laticollis Stephens
T. obscurus Fabriciust BC AB -oNPQ PE NF
poliens Casey

Genus NEATUS LeConte

N. tenebrioides (Palisot de BC MB ON PQ
Beauvois)
badius (Say)
picipes auct. amer.

Genus IDIOBATES Casey

D. castaneus (Knoch) ON PQ
interstitialis (Say)

Tlibe Amarygmini
Genus MERACANTHAKiTbv
Falacer Castelnau
M. contracta (Palisot de Beauvois) ON
canadensis Kirby
cuprea (Castelnatt)
inflata (Solier)
(?) rugosa (Castelnau)
(?) turnida (Melsheimer)

Subfamily ALLE CLILINAE

Tribe Alleculini
Genus HYMENORUS Mulsant
Hymenophorus Mulsant
Key to North American species: Fall (1931)
H. caurinus Fall BC
H.
H.
discretus Casey
megops Hatch
-oN-
BC
H. molestus Fall _ONPQ NB
H.
H.
niger (Melsheimer)
obesus Casey
-oNPQ
_ONPQ
NB

H.
H.
picipennis Casey
pilosus (Melsheimer)
-oNPQ
H. sinuatus Fall BC - :*PQ
Tiibe Gonoderinr
Genus ISOMIRA Mulsant
Tedinus Casey
Key to eastern North American species: Marshall (1970)
I. comstocki Papp BC
I.
I.
iowensis Casey
oblongula Casey
ow:
-_ONPQ
I. pulla (Melsheimer) -oNPQ
ignora Blatc}:.ley
I. quadristriata (Couper) MB ON PQ NB NS
ueluntina LeConte
I. sericea (Say) -oNPQ _NS
tenebrosa Casey
I. valida Schwarz -oN
similis Blatchley

Genus PSEUDOCISTELA Crotch

Chromatia LeConte
Cisfelo Fabricius
Telacis Poey
P. amoena (Say) -ONPQ
P. brevis (Say) -ONPQ
erythroptera (Ziegler)

259
Tenebrionilae

P. pectinat.r Hopping - BC
P. pinguis l.eConle - BC
pacific:a Hopping

Genus CAI'NOCHROA LeConte

C. fuliginosa (Melsheimer) - ON PQ NB

Genus ANI)ROCHIRUS LeConte

A. erythropus (Kirbyt - ON PQ - NS
fuscipes (Melsheimer)
luteipes LeConte

Tlibe Mycetocharini
Genus MY(IETOCHARA Berthold
M y ceto chare s Latreille
M y cetocharis Gyllenhal
M y cetct p h il a Gyllenhal
Pterna Seidlitz
Stigmotoma LeConte
Key to North American species: Campbell (1978)
M. analis (LeConte) -BC-SKMBONPQ
dauisi Hatch
horni ))rry
lugubris (LeConte)
M. bicolor (Couper) -ONPQ
M. binotata (Say) -oNPQ
longula (LeConte)
margiltata (LeConte)
M. foveata .LeConte) -ONPQ
gilipes Casey
gracilis (LeConte)
lecontti Borchmann
tenuis (LeConle)
M. fraterna (Say) MB ON PQ
laticol is (LeConte)
megahlps Casey
nigerrtma Casey
M. procera Casey _BC
angusta Hatch
caseyi Hatch
crassu]ipes Casey
downe'.Hatch
malkh;iHatch
neuadensis Casey

Genus HYI\{ENOCHARA Campbell

Key to species: Campbell (I978d)
H. rufipes (LeConte) -oNPQ
Subfamilv COELOMETOPINAE
Tri6e Coelometopini
Genus MEIIINUS LeConte
M. laevis tCtliviert - ON PQ

Genus XYLOPINUS LeConte

T aenoltate s Motschulsky
X. aenescens LeConte PQ
X. saperdioides tOlivier) - ON PQ
anthracinus (Knoch)
calcaratus (Palisot de Beauvois)
ruftpes (Say)
spinipes (Fabricius)

Genus HAILANDRUS LeConte

H. concolor LeConte MB ON PQ

260
 Lagriidae

H. fulvipes (Herbst) oN PQ
femoratus (Fabricius)

Genus COELOCNEMIS Mannerheim

Key to species: Doyen (1973)
C. californica Mannerheim _BC AB
barretti BlaisdelI
bosolis Casey
columbiana Casey
dilat icoll is Mannerherm
idahoensis Casey
ouipennis Casey
rauca Casey
rugosa Linne\l
spaldingi Casey
utensis Casey

Genus IPHTHIMINUS Spilman

I. opacus (LeConte) _SX yu oN PQ NB NS
I. salebrosus (Casey)
L serratus (Mannerheim)
-BC
_BC AB
serudtor (Walker)
seruilis (Walker)
subligatus (Walker)

Genus AI-OBATES Motschulsky

A. pensylvanicus (DeGeer) MB ON PQ
chrysops (Herbst)
inermis (Mannerheim)
sublaeuis (Palisot de Beauvois)

Genus UPIS Fabricius

U. ceramboides (Linn6)x AK YK NT BC AB SK MB ON PQ NB PE LB NF
reticulata (Say)
uariolosa (DeGeer)

Tribe Strongyliini
Genus STRONGYLIUM Kirbv
Reminius Casey
Stenochia I{rtby
Key to North American species: Triplehorn and Spilman (1973)
S. tenuicolle (Say) ON PQ

Family LAGRIIDAE lagriid beetles

J.M. Campbell

The family Lagriidae is primarily tropical in dis- lagriids as a distinct family until their relationship
tribution and comprises over 1000 species around the with respect to other tenebrionoid groups has been
world. Only six species have been recorded in Canada. better established. The classification of the North
Adults of Statird and, Arthromacra are usually col- American lagriids was reviewed by Parsons (19750),
lected by beating the foliage of trees; larvae are who includedAnaedus andParatenetus in the subfam-
thought to breed in decaying wood or leaf litter. Little ily Lupropinae (tribe Heterotarsini of Tenebrionidae,
is known about the biolory of the species. in most publications); this subfamily more properly
The status of this family is uncertain. In recent should be named Goniaderinae. Doyen and Tschinkel
studies of the higher classification of the (1982) have suggested that the genus Paratenetus be
Tenebrionidae, Watt (t974), Doyen and Lawrence placed in or near the tenebrionid tribe Belopini. Until
(1979), and Doyen and Tschinkel (1982) placed the the classification of the Lagriidae is reviewed, I have
lagriids in Tenebrionidae, usually as a subfamily. chosen to follow the arrangement of Parsons (1975o).
However, Doyen and Tschinkel (1982) suggested that
the lagriids and related groups form the sister group
to all other tenebrionids and possibly should be ttlT (1);sC (1);AB (1);MB (3); oN (5); PQ (5);NB (1);
restored to family status. I prefer to recognize the NS (1)

26t
Cephaloidae

Subfamily GONIADERINAE
Genus PAR.A.TENETUS Spinola
Storthe,phora Mdklin
P. crinitus lrall
P. fuscus LeConte
P. punctatus Spinola
MB3N|8- - - -
- - NrncAB- -oNPQ -
Genus ANAEDUS Blanchard
A. brunneur; (.Zieglet) ON

Subfamily LAGRIINAE
Genus ART.FIROMACRA Kirby
Key to N,.'rth American species: Parsons (1975a)
A. aenea (Say) MB ON PQ NB NS
donacioides I\trby

Subfamily STATIRINAE
Genus STA IIRA Audinet-Serville
Key to Ntrrth American species: Parsons (I975a)
S. gagatina Melsheimer PQ
respler,.dens Melsheimer

Family CEPIIALOIDAE false long-hornedbeetles

J.M. Campbell

Cephaloidae is a small family of about 20 the genus Cephaloon. Subsequently, Arnett (1968)
species rerstricted to the cooler parts of the Holarctic transferred the subfamilies Stenotrachelinae and
region. l)nly nine species are known to occur in Nematoplinae from the melandryiids and pediliids,
Canada ernd Alaska. Larvae are found in old, rot- respectively, to Cephaloidae. Both subfarrLilies have
ting logs; adults of Cephaloon are often taken on been considered by some authors to be disbinct fam-
flowers. ilies (Mamayev 1973), but I have followed Lawrence
Species of this family are reasonably well (1982a) in accepting Arnett's treatment.
known taxonomically. Hopping and Hopping (1934)
and Arnett (1953) revised and reviewed, respec- AK (1); YK (1); NT (2); BC (5); AB (3); MB (1); ON
tively, the species of the family, but included only (a); PQ (5);NB (1); PE (1);LB (2);NF (2)

Subfamily CEPHALOINAE
Genus CEPHALOON Newman
Drachj,lis Casey
Ichnodes Dejean
Sponia',ium Casey
Typitium Casey
Key to species: Arnett (1953)
C. bicolor Horn BC
simulans (Casey)
C. lepturoid.es Newman oN PQ
uarians Haldeman
C. pacifrcum Van Dyke -BC
_BCAB
C. tenuicome LeConte
piceuri }Iorn
ornatum Casey
uersicolor Casey
C. ungulare LeConte ON PQ PE LB NF

262
 Meloidae

Subfamily STENOTRA.CHELINAE
Genus STENOTRACHELUS Latreille
S. aeneus (Fabricius) AK YK NT BC AB MB ON PQ LB NF

Genus ANELPISTUS Horn

A. americanus Horn PQ
A. canadensis Mank NT BC AB

Subfamily NEMATOPLINAE
Genus NEMATOPLUS LeConte
N. collaris LeConte ON PQ NB

Family MELOIDAE blister beetles

J.M. Campbell

Meloidae comprises over 3000 species, which cantharidin (Spanish fly), which causes blisters on
are found in most regions of the world. However, contact with the skin and, if present in dried hay,
the majority are restricted to warmer, drier areas. has been known to be fatal to horses. Cantharidin
Only 49 species and subspecies are known to occur is also used for medical purposes.
in Canada and Alaska, and most of these are found Most genera of North American Meloidae are
in the semiarid areas of the Prairie Provinces and well known taxonomically. A few genera such as
British Columbia. Larvae of Epicautini feed on Tricrania have not been revised, and the large,
grasshopper eggs, and those of aII other Canadian important genus Epicauta is in need of additional
blister beetles feed on eggs, larvae, and food study. The Canadian fauna is well known, and
reserves of ground-nesting bees. Larvae of Meloini therefore few additional species are expected to be
and Zonitinae are phoretic. The eggs are laid on added to the checklist in the future. Van Dyke
flowers, the larvae hatch, and the frrst-instar larvae (1928a) and Arnett (1968) provide the most recent
(called triungulin) attach themselves to bees that treatments of the family at the generic level.
visit flowers and are transported to the bee's nest,
where they drop off. Adults of Canadian meloids are
phytophagous or feed on nectar; many are consid- aK (3); vK (1); NT (3); sC (25); AB (26); SK (23);
ered economic pests. Adults of this family produce MB (20); ON (17);PQ (12);NB (3);NS (3)

Subfamily MELOINAE
Tribe Meloini
Genus EURYMELOE Reitter
E. aleuticus (Borchmann) AX

Genus MELOE Linn6

Key to North American species: Pinto and Selander (1970)
(Subgenus TREIODOUS Dugbs)
M. barbarus LeConte BC
(Subgenus MELOE s. str.)
Melittophagus Kirby
M. americanus Leach ON
tnoerens LeConte
temnus Dillon
M. angusticollis Say BC AB SK MB ON PQ NS
montanus LeConte
rugipennis LeConte
M. carbonaceus LeConte
M. dianellus Pinto & Selander
M. impressus Kirby
---ncffisx-oNPe
AK YK NT BC AB SK MB ON PQ
-NS
NB
occidentalis Van Dyke

263
Meloidae

M. niger Kirby NTBCABSKMBONPQ_NS

hirbyi Drllon
opacus LeConte
perplee:us LeConte
M. occultus Pinto & Selander _BC
M. strigulos,us Mannerheim AK BC
M. vandykei Pinto & Selander - BC

Tlibe Lyttini
Subtribe Lyttina
Genus LYTTA Fabricius
Key to species: Selander (1960)
(Subgenus PARALYTTA Selander)
L. vulnerata tl,eContet - BC
anguicollis (Dugbs)
cooperi LeConte
(Subgenus I'OREOSPASTA Horn)
L. cyanipen:ris (Leconte) - BC AB
solicis LeConte
L. nuttalli Say AB SK MB
fulgifera (LeConte)
L. stygica (LeConte) - BC
arbore<:.,. Wellman
difficihs (FalI)
dolosa LeConte
neuadezsis Van Dyke
purpurzscens (FaII)
smarag'dula (LeConte)
stolida (Fa11)
L. viridana -eConte - BC AB SK MB
(Subgenus I'OMPHOPOEA LeConte)
L. sayi LeCr,nte - ON pe
pyriuorz (Fitch)

Subtribe Epicautina
Genus EPICAUTA Dejean
Key to North American species: Werner (1945) (see also Ballmer 1g79; pinto 1g80)
(Subgenus EPICAUTA s.str.)
(Anomelony c hus Saylor)
(Gnathospasta }Jorn)
(Henous Haldeman)
(Maculicauta Dillon)
(Nomaspis LeConte)
(P le u rop om p ha LeOonte)

E. atrata (Frrbricius) MB
coracin,z (Illiger)
trichrus (Pallas)
E. cinerea (I'orster) - SK MB ON pe
fissilabris (LeConte)
E. fernrgine,r (Sayt AB SK MB ON
E. immerita Walker - BC
E. maculata (Say) - SK MB ON
conspersa (LeConte)
media I)ugds
nogales Werner
punctuata (Dugds)
E. normalis Werner - BC AB SK
E. oregona Florn - BC AB SK MB
E. pennsylvcLnica(DeGeer) AB SK MB ON PQ NB
morio (l,eConte)
zigro (Woodhouse)
E. piceiventrisMaydell - BC AB SK MB
elong att>calcarafo Maydell
E. pruinosa LeConte - BC AB SK MB

264
 Meloidae

E. puncticollis Mannerheim - BC AB
barberi Werner
oblita LeConte
E. sericans Leconte AB sK
E. ventralis werner AB sK
E. vittata (Fabriciust - ON PQ
chapmani (Woodhouse)
I e rnni scata (Fabricius)

(Subgenus MACROBASIS LeConte)

(Ap teros pasta LeConte)
E. fabricii (LeContet AB SK MB oN PQ
debilis (LeConte)
E. Ilavocinerea (Blatchleyr MB ON
E. murina (LeConte) - ON PQ NB
unicolor Kirby
E. subglabra rFall I - BC AB SK MB
Genus LINSLEYA MacSwain
Key to species: Selander (1955)
(Subgenus LINSLEYINA Selander)
L. sphaericollis {Say) - BC AB SK MB
chalybea (LeConte)
c haly be ata ( Gemrninger)
infidelis (Fall)

Subfamily ZONITINAE
T?ibe Zonitini
Subtribe Nemognathina
Genus NEMOGNATIIA Iiliger
Key to North American species: Enns (1956)
(Subgenus MEGANEMOGNATHA Enns)
N. lutea dichroa LeConte - BC AB SK MB
N. lutea lutea LeConte AB
N. soror l\lacSwain - BC
(Subgenus PAURONEMOGNATIIA Enns)
N. nemorensis Henlz - ON
b irnac ulata Melsheimer
(Subgenus NEMOGNATHA s. str.)
N. piazata bicolor LeOonte AB SK MB
discolor LeConte
stellaris Beauregard
texana LeConte

Genus TRICRANIA LeConte

Tricraniodes Wellman
T. mur"rayi LeConte - BC
T. sanguinipennis {Say) - ON PQ
T. stansburl (Haldeman) NT BC

Genus HORNIA Riley

H. minutipennis Riley AB

Subtribe Zonitina
Genus ZONITIS Fabricius
Key to North American species: Enns (1956)
(Subgenus NEOZONITIS Enns)
Z. bilineata Say - ON PQ
lineata Mels}reilner
m an d ib u laris Mel sheimer
Z. vermiculata Schaeffer - BC
(Subgenus PARAZONITIS Enns)

265
Oedemeridae

Z. sayi Wickham AB SK MB
imrnactilata (Say)

Genus GNAIHIUM Kirby

Key to species: MacSwain (1952)
G. minimurrL (Say) AB SK

Family OEDEMERIDAE false blister beetles

J.M. Campbell

The family Oedemeridae comprises at least 100 species have been reported to cause blister:ing simi-
genera arrd 1500 species, distributed in all faunal lar to that produced by some blister beetles,.
regions. Thirteen species are known to occur in Arnett (1951) revised the North American
Canada and Alaska. The biology of most species is species of oedemerids; little information has since
unknown. Larvae of some species live in rotten been added to our knowledge of the Clanadian
wood, and adults of most species are found on flow- fauna.
ers, where they feed on pollen and nectar. One
species, Nacerdes melanura (wharf borer), is an eco- AK(2); BC (10);AB (4); MB (3); ON (5); PQ (5);NB
nomic pest of pilings and ship timber. A few exotic (1);NS (2);NF (1) ' I (1)

Subfamily CALOPODINAE
Genus CAJ-OPUS Fabricius
C. ansustus LeConte BC AB oN PQ

Subfamily NACERDINAE
Genus XANTHOCHROA Schmidt
X. testacea llorn BC

Genus NACIIRDES Dejean

Anogcotles Dejean
N. melanura (Linn6)t BC MB ON PQ NB NS NF
apicalis (Say)

Subfamily OEDEMERINAE
Tlibe Ditylini
Genus DITYLUS Fischer von Waldheim
D. caeruleus (Randall) _MB ONPQ_--__
D. gracilis L,:Conte AK BC
uestitus LeConte
D. quadricol)is LeConte AK BC AB
consors LeConte

Tribe Asclerini
Genus XANIIHROCHROINA Ganglbauer
X. bicolor (LerConte) BC

Genus ASCL,ERA Stephens

A. discolor L,eConte _BC
A. excavata]-eConte -BCAB
A. nigra LeConte
A. puncticollis (Say) ---:t--^ras oN pq
A. ruficollis (Say) ON PQ NS
carinatct (Newman)

Genus COPIDITA LeConte

C. quadrimaculata LeConte BC

266
 Mycteridae

Family PROSTOMIDAE prostomid beetles

J.M. Campbell

Prostomidae is a small family that contains only In most North American literature Prostomis
two genera and about 20 species. Most faunal arnericanus has been referred to as P. mandibularis
regions, except South America, have a few species of (Fabricius), a European species.
the family; only one species occurs in Canada.
Nothing is known about the habitat of this species. BC (1)

Genus PROSTOMIS Latreille

P. americanus Crotch BC

FamilySYNCHROIDAE synchroidbeetles
L. LeSage

As currently defined (Crowson 1966, Lawrence There is no recent treatment of the North
1982a), the family Synchroidae is reduced to two American species of this family.
genera and 10 species, which were previously
placed in the Melandryidae. Only one species of the
genus Synchroa occurs in Canada. Both adults and
Iarvae live under the bark ofold dead trees. uB (1);oN (1);PQ (rl

Genus SYNCHROA Newman

S. punctata Newman MB ON PQ
nzurina Haldeman
umbrina Melsheimer

Family MYCTERIDAE mycterid beetles

J.M. Campbell

Mycteridae includes about 160 species around (Hatch 1965). I have followed Lawrence (1982o) in
the world, of which four are known to occur in considering mycterids to be a distinct family. The
Canada. Larvae live under bark, and adults are species are rather poorly known taxonomically, and
often found on flowers. a modern revision is needed badly.
Species of this family have been included in a
variety of families, including Melandryidae (Leng
L920), Salpingidae (Arnett 1968), and Pythidae sC (2); ON (2);PQ (2)

Genus LACCONOTUS LeConte

L. pinicola Horn BC
L. punctatus LeConte oN PQ

Genus MYCTERUS Clairville

Key to North American species: Hopping (1935)
M. concolor LeConte BC
flauipennis Horn
M. scaber Haldeman ON PQ

267
Pythidae

Family BORIDAE
J.M. Campbell

Boridae contains three genera and six species; (Arnett 1968). Young (19856) transferred Lecontia
only two species occur in Canada. Larvae of Boros from the family Pythidae to the Boridae. I have fol-
occur under the bark of coniferous trees and nroba- lowed Lawrence (1982a) in recognizing Boridae as a
bly feed on decayed cambium (Lawrence L9824). separate family.
The species of Boridae have been placed in a
variety of families such as Tenebrionidae (Leng NT (1); sC (1); AB (2); SK (2); MB (2); O.N (2); PQ
1920), P;'thidae (Hatch 1965), and Salpingidae (2);NB (1)

Genus BOR()S Herbst

B. unicolor Siay SK MB ON PQ

Genus LECONTIA Champion

Crymoctes LeConte
L. discicollis (LeConte) NT BC AB SK MB ON PQ NB

Family PYTHIDAE pythid beetles

J.M. Campbell

The family P1'thidae forms a small group com- (Arnett 1968) or, conversely, Pythidae has included
prising orrly about 50 species around the world, of a number of taxa now transferred to other families
which six occur in Canada and Alaska. Members of such as Salpingidae, Cononotidae, and Mycteridae.
the subfermily Pythinae occur primarily in the The arrangement followed here is based on
Holarctic region, whereas members of the other two Lawrence (1982a) with the exception of Lecontia,
subfamilies are found in Central and South which has been transferred to Boridae as Young
America, .New Zealand, and Australia. Both larvae suggests (1985b). Pollock is currently revising the
and adults are found under bark ofconiferous trees. genus Pytho, so that most species of the family will
Larvae fe':d on decaying inner bark (Pytho) or sap- soon be well known taxonomically.
wood (Prirgnathus).
Meml,ers of this family have previously been AK (3); YK (3); NT (3); BC (5); AB (4); SK (3); MB
assigned to other families such as Salpingidae (a);ON (a);PQ (a);NB (3);NS (a); LB (1);NF (2)

Subfamily PYTHINAE
Genus PYTIIO Latreille
P. americamrs Kirby AK YK NT BC AB SK MB ON PQ NB NS LB
deplanatus Mannerheim
P. niger Kirtry
P. seidlitzi Blair
P. strictus L,-.Conte
5v NT BC
ItBCy
AB SK MB ON PQ
MB ON PQ
NB
NS
NS
NF

Genus PRIOGNATHUS LeConte

P. monilicornis (Randall) AK YK BC SK MB ON PQ NB NS NF
sahl b e rg i (Mannerheim)
Genus SPHI'LMA Horn
S. quadricollis Horn BC

268
 Pyrochroidae

Family PYROCHROIDAE fire-colored beetles

J.M. Campbell

Twelve species of the family Pyrochroidae are Neopyrochroo and Schizotus ceruicalis are attracted
known to occur in Canada and Alaska. Adult to cantharidin.
pyrochroids are rarely collected, although adults of The family is well known taxonomically, thanks
Dendroides and Neopyrochroo occasionally are to the revision and catalog of the North American
attracted to light, and those of most species can be species by Young (1975 and 1983, respectively).
collected in flight intercept traps or Malaise traps. However, in recent papers, Young has suggested
Larvae of many species are common and are among that Ischalia be transferred to the Anthicidae
the most frequently collected and distinctive species (1985o) and that the genus Pedilus be transferred
of Coleoptera found under bark and in rotting wood. from Pedilidae to Pyrochroidae (1984). I prefer to
Young (1975) summarized the feeding habits and postpone making these changes until his more
biology of pyrochroids. Larvae live in dead wood detailed analysis of Pedilus is published.
and are apparently fungivorous rather than preda-
ceous or xylophagous, as reported by earlier
authors. Adults rarely feed, but may be attracted at AK (2); NT (1); BC (5); AB (a); SK (1); MB (3); ON
night to fermenting fruit or bait. Adults of (7);PQ (7); NB (3); NS (a);NF (1)

Subfamily ISCIIALIINAE
Genus ISCHALIA Pascoe
(Subgenus EUPLEURIDA LeConte)
L costata (LeConte)
I. vancouverensis Harrington ecAB--9*PQ-TS---
Subfamily PYROCHROINAE
Genus SCHIZOTUS Newman
S. cervicalis Newman NT BC AB MB ON PQ NB NS
S. fuscicollis (Dejean) AK

Genus NEOPYROCHROA Blair

N. femoralis (LeConte) -oNPQ
N. flabellata (Fabricius) -ONPQ
Genus DENDROIDES Latreille
D. canadensis Latreille MB ON PQ NB NS
blcolor (Newman)
cyanipennis Latreilie
D. concolor (Newman) _ONPQ NB NS NF
D. ephemeroides (Mannerheim)
D. marginatus Van Dyke T BC
BC
AB

pacificus Barrett
D. picipes Horn BC
D. testaceus LeConte AB SK MB ON pe

269
Salpingidae

Family SALPINGIDAE narrow-waistedbarkbeetles

J.M. Campbell

Salpingidae comprises 10 species in Canada and pingid genera were previously included in families
Alaska and about 350 species around the world. such as the Pythidae and Colydiidae (Agleruus). The
The adults are most often found under the bark of family is poorly known taxonomically, and a com-
conifers, on flowers, or in detritus. Their feeding prehensive revision of most North American genera
habits are unknown. is needed.
Salpingidae, as currently constituted (Lawrence
Ig82a), includes groups previously treated as dis-
tinct families, such as the aegialitids, dacoderids, AK (3); YK (1); NT (1); BC (7); AB (1); SK (1); MB
elacatidsr, eurystethids, inopeplids, othniids, and (1); oN (3); PQ (2); NB (1); NS (1); LB (1); NF (2)
tretothoracids. On the other hand. some of the sal- . I(1)
Subfamily SALPINGINAE
Genus SPHAERIESTES Stephens
Salpingus atct.
S. alternatus (LeConte) BC
S. virescens (LeCon!e) YK NT BC AB SX nrS ON pe NS - is Nr
Genus VINCENZELLUS Reitter
V. elongatus (Mannerheim) AI(
Genus RHINOSIMUS Latreille
Key to Ncrth American species: Blair (1932)
R. viridiaerLeus Randall AK BC oN PQ NS
a e n e ir nt r is Mannerheim
nitens LeConte
R. iecontei Blair -BC
R. pallipes Boheman _BC

Subfamily AE GIALITINAE
Key to species: Spilman (1967)
Genus AEC|IALITES Dejean
Eurystethus Seidlitz
A. californi,:us Motschulsky AK BC

Subfamily OTHNIINAE
Genus ELA.CATIS Pascoe
Othnius LeOonte
E. lugubris (Horn) BC

Subfamily AGLENINAE
Genus AGLENUS Erichson
A. brunneus (Gyllenhal)t ON
obsoler)us (Shuckard)

270
 Anthicidae

Family ANTHICIDAE antlike flower beetles

Y. Bousquet

Anthicidae comprises 55 species in Canada and and Anthicinae. Thanks to the work of Werner and
Alaska. The adults are usually found under debris of Chandler, the taxonomy of most of the groups
and in leaf litter on the ground or on flowers and that occur in North America has been worked out.
foliage; the larvae live in the soil and in decayed
vegetation. Apparently, members of many species
are scavengers; those of some species are known to
be egg predators. AK (2); YK (2); NT (5); BC (29); AB (25); SK (24);
The North American species of this family are MB (19); ON (25); PQ (21); NB (9); NS (5); PE (1);
usually grouped into two subfamilies, Notoxinae NF(3) .I(3)
Subfamily NOTOXINAE
TYibe Notoxini
Genus MECYNOTARSUS LaFert6-S6nectbre
Key to New World species: Chandler (1977)
M. candidus LeConte AB

Genus NOTOXUS Fabricius

Ceratoderus Blanchard
Monocerus Faldermann
Key to North Arnerican species: Chandler (1982)
_NT_ABSK
N. anchora Hentz
N. bifasciatus (LeConte)
MB ON
MB-_
PQ
Iu_ PE

N. cavicornis LeConte BC
breuiusculus Fall
obesulus Blaisdell
N. desertus Casey ON
constrictus Casey
N. intermedius Fall AB SK
paruidens Fa\l
N. manitoba Chandler MB_
N. marginatus LeConte -AB
_AB
N. montanus Casey
microcerus Casey
similis Fall
N. pictus Casey BC AB
N. robustus Casey BC
humboldti Casey
N. schwarzi Horn BC
N. serratus (LeConte) BC AB ix
digitalis LeConte
uandykei Blaisdell
N. subtilis LeConte AB SK MB ON

Subfamily ANTHICINAE
TYibe Anthicini
Genus AMBLYDERUS LaFert6-S6nectdre
A. granularis LeConte ON PQ
A. obesus Casey BC
A. pallens LeConte BC AB SK MB ON PQ NB

Genus ANTHICUS Paykull

Nathicus Casey
Key to North American species: Werner (1964)
A. ancil)a Casey AB SK
caseyi Pic
parallelus Casey
A. bellulus LeConte BC

277
Anthicidaer

heluin.us Casey
minusculus Champion
A. biguttulus LeConte BC
ouicol,'.is Casey
protectus Casey
A. cervinusi LaFert6-S6nectbre NT BC AB SK MB ON PQ
bifasc;iatus Say
b izonc'.t u s LaFert6-S6nectbre
gilenst.s Casey
seminotatus Casey
termir:alis LeConte
A. coracinu.s LeConte
A. cribratus LeConte
hecate Casey
AK YK NT BC
BC
AB
:" yu 9*
PQ NB NS

A. ephippium LaFert6-S6nectbre BC AB SK MB ON PQ
confusus LeConte
diffici;.is LeConte
luteolus LeConte
pinguescens Casey
simiolus Casey
A. flavicans LeConte _NTBCAB SK MB ON PQ NB NS
A. haldemr.ni LeConte -NT_AB SK_ON PQ NE
decorellus Casey
q uadr iguttatus Haldeman
sozcias Casey
A. hastatus Casey BC AB SK MB
solidus Casey
subcati.uus Casey
A. heroicus Casey PQ NB
A. lecontei Champion --ec AB SK _ONPQ
auriger Casey
junctus Casey
stellat,^ts Casey
uulneratus Casey
A. lutuient.rs Casey BC AB SK ON
agilis Casey
herifu.,lus Casey
mund.ts Casey
A. melanch olicus LaFert6-S6nectbre ON PQ
facilis Casey
latebntns LeConte
spretus LeConte
A. nanus L.:Conte BC
amoenus Casey
innocens Casey
inscitt',s Casey
nympltn Casey
obliquus Casey
peninsularis Casey
uagan:; Casey
A. nigritus Mannerheim AK YK NT
A. praeceps Casey BC
A. punctule.tus LeConte BC AB S^ l,fn -
cephalotes Casey
decrepitus Casey
mercurialis Casey
monticola Casey
A. scabriceps LeConte AB SK MB ON PQ NB NS NF
compositus Casey

Genus SAPINTUS Casey

Key to Ncrth American species: Werner (1962)
S. fulvipes l.LaFert6-56nectdre) BC _MBONPQ
colonus Casey
rustict:s Casey
S. lemniscatus Werner ON
S. pubescer.Ls (LaFert6-S6nectdre) oN PQ
S. pusillus (LaFert6-S6nectbre) _MBONPQ_NS
binomi.natus Pic
festinans Casey
mollis Casey

272
 Anthicidae

Genus STRICTICOMUS Pic

S. tobias (Marseul)i PQ

Genus OMONADUS Mulsant & Rey

Hemantus Casey
Key to North American species: Werner (1964)
O. floralis (Linn6)t - BC AB SK MB ON PQ NB - NF
(?)basillaris (Say)
O. formicarius (Goeze)f - BC AB SK MB ON PQ NB
enodis (Casey)
rixator (Casey
scenicus (Casey)

Genus MALPORUS Casey

M. cinctus tsay) - SK - ON
M. formicarius (LaFert6-S6nectbre) - SK MB ON PQ NB NS

Genus THICANUS Casey

T. mimus Casey - BC AB SK MB

Genus EIJVACUSUS Casey

E. coloradanus Casey - BC AB SK MB

Genus VACUSUS Casey

Key to species: Werner (1961)
V. formicetorum (Wasmann) AB SK
desertorum Casey
prominens Casey
suspectus Casey
V. nigritulus (LeConte) - BC
arcanus Casey

Genus ISCHYROPAIPUS LaFert6-S6nectbre

Lappus Casey
Key to North American species: Werner (1973)
L cochisei Werner - BC
L nitidulus (LeConte) - BC - ON PQ
alacer (Casey)
asperulus (Casey)
cursor (Casey)
nubilatus (Casey
soliuagans (Casey)
uigilans (Casey)
I. obscurus (LaFert6-s6nectdre) - ON PQ
I. sturmi (LaFert6-S6nectbre) - ON
e le g ans (LaFert6-S6nectbre)
I. turgidicollis (Casey) - BC
animatus (Casey)

Genus TANARTHRUS LeOonte

Key to species: Chandler (1975)
(Subgenus TANARTHROPSIS Casey)
T. salicola LeConte - SK
densus Casey
mormon Wickham
nubifer Casey
tricolor Casey
uirginalis Wickham

273
Pedilidae

Family PEDILIDAE pedilid beetles

Y. Bousquet

Pedilidae currently includes 19 species in Eurygeniinae. Young (1984) suggestecl that the
Canada; none has been reported from Alaska. The genus Pedilus be transferred from Pedilidae to
adults usually occur on flowers or foliage and feed Pyrochroidae and placed in its own subfamily
on flowers, pollen, and nectar; they are attracted to (Pedilinae) and that other North American pedilids
light at night. The larvae live in decaying plant be tranferred to Anthicidae. I prefer to wait until
matter, uLsually in wet places. Young's revision of the genus Pedilus is published
The group is probably closely related to before incorporating these changes.
Anthicidae, and some authors, including Crowson
(1955) and Lawrence (1982o), have included the
pedilids in Anthicidae. Members of Pedilidae are BC (8); AB (4); SK (1); MB (2); ON (6); PQ (e); NB
usually classified in two subfamilies, Pedilinae and (1); NS (1)

Subfamily PEDILINAE
Genus PEDILUS Fischer von Waldheim
Corphyra Say
abnormis (Horn) _?AB-_
P.
P. canaliculatus (LeConte) pe-----
P. cyanipen.nis Bland PQ
P. elegans (Hentz) MB ON PQ
fuluipe's Newman
h a e m a t ho i dal i s Zi e gler
lugubris Newman
newmani LeConte
ruficollis Ziegler
P. flabellat:rs (Horn) BC
P. impressrLs (Say) ON PQ
P. inconspicuus (Horn) BC
P.
P.
longilobrs Fall
lugubris (Say)
imzs llewman
___f _ _ ON pe NB NS

inornatus (Randall)
infumtr.tus (Hentz)
nigricctns Ziegler
rufithorax Newman
P. oregonus Fall _BC
P. punctulatus LeConte _BC
funebris (Hotn)
P. terminalis (Say) ON PQ
guttulu Newman

Subfarnily EURYGENIINAE
Tlibe Eurygeniini
Genus PEBGETUS Casey
P. campanulatus (LeOonte) BC

Genus STEREOPAJ-PUS LaFert6-S6nectdre

Key to species: Abdullah (1965)
S. columbianus Hopping BC
S. guttatus LeConte BC AB
impressicollis Casey
subalbicans Casey
S. hirtus Hatch
S. rufipes C:asey __BC_-_-_ po
S. vestitus Say) SK MB ON PQ
bad,iipennis LeOonte

274
 Scraptiidae

Tribe Macratriini
Genus MACRATRIA Newman
M. confusa LeConte ON PQ

Family ADERIDAE antlike leaf beetles

Y. Bousquet

Aderidae is a small family that includes nine Except for Werner's (1990) recent review of the
species within the borders of Canada. Very little is eastern North American species, members of this
known about these beetles. The adults are usually family has been little studied taxonomically.
found on foliage and flowers; the larvae of some
species have been found in rotten wood and leaf litter. gC (1); ON (8); PQ (8); NB (1); NS (1) ' I (1)

Genus ELONUS Casey

E. basalis (Leconte) - ON PQ
princeps Casey
E. nebulosus (LeConter - ON PQ

Genus ADERUS Westwood

Phomalus Casey
A. populneus (Panzertt - BC - ON PQ

Genus VANONUS Casey

Tanilotes Casey
V. calvescens Casey PQ
V. piceus (LeConte) - ON
congener Casey
fusciceps Casey
t ub er c ulifer ( Hamilton)
V. vigilans Casey - ON PQ
V. wickhami Casey - ON PQ NB NS
lacustris (Casey)

Genus ZONANTES Casey

Z. fasciatus (Melsheimert - ON PQ
tricuspis Casey
Z. pallidus Werner - ON PQ

Family SCRAPTIIDAE scraptiid beetles

L. LeSage

The family Scraptiidae, which comprises about There is no recent treatment of the family in
400 species around the world and 21 in Canada and North America; the Canadian fauna is poorly
known except in the provinces of Quebec and
Alaska, represents a group of beetles with uncertain
affinities. Previously placed in Mordellidae, or British Columbia. Young (1976) transferred the
Melandryidae, or Anthicidae, they are now consid- genus Pectotom.a, previously included in
ered to be a separate farnily closely related to those Tetratomidae, to Scraptiini.
families (Crowson 1955; Franciscolo 1963; Lawrence
I982a). Adults are often found in large numbers on
flowers, and larvae occur in decaying wood or in leaf AK (2); yK (1); gC (16); AB (5); SK (1); MB (1); ON
litter. (8);PQ (10); NB (3);NS (2); NF (3)

275
Scraptiidac

Subfamily ANASPIDINAE
Tribe Anaspidini
Key to North American species: Liljeblad (1945)
Genus ANIiSPIS Geo{Iroy
A. atrata Champion YK BC AB
atra LzConte
A. duryi Liljeblad BC
A. flavipennis Haldeman BC ON PQ NB
A. nigrina Csiki BC ON PQ NF
zlgro (Haldeman)
A. rayi Hat,ch BC
A. rufa Say AK BC AB ON PQ NB lls - NF
dimidiata Melsheimer
filiforrnis LeConte
nigrice:ps LeConte
pallescens Mannerherm
uenlro'is Melsheimer
A. seposita Liljeblad _BC
A. sericea ( \{annerheim) AK BC
luteipennis LeConte

Genus SILARIA Mulsant

S. revelstokei Liljeblad BC

Genus LARISIA Emery

L. nigricolo r Liljeblad BC AB ON

Genus NASSIPA Emery

N. hopping Liljeblad BC

Subfamily SCRAPTIINAE
Tlibe Allopodini
Genus ALLOPODA LeOonte
A. californir:a Schaeffer BC
A. lutea (Haldeman)
fuscos ut ur alis (Blatchley)

Tlibe Scraptiini
Genus SCRAPTIA Latreille
S. sericea t lylelsheimer) PQ

Genus CAI IFA LeConte

C. pallipennis LeConte PQ
C. pailipes' Melsheimert BC MB ON PQ NS NF
biimpressa (Haldeman)
C. plagiata ( Melsheimert ON PQ
americana (Haldeman)
C. pusilla (I{aldeman) ON PQ NB

Genus PECTOTOMA Hatch

P. hoppingi Hatch BC AB

T?ibe Pentariini
Genus PENTARIA Mulsant
Anthobates LeOonte
Anthobatula Sftalrd.
Key to North American species: Liljeblad (1945)
P. fuscula (l,eConte) BC PQ
P. trifasciata Melsheimer BC AB SK -ONPQ
nubila (LeConte)

276
 Cerambycidae

Superfamily Chrysomeloidea
Family CERAMBYCIDAE longhorned beetles
J. McNamara

Cerambycidae is represented in Canada and in forest ecolory because they help in the decompo-
Alaska by 354 species and subspecies; it is one of sition ofdead and dying trees.
the largest groups of wood-boring insects. These Thanks to the works of Linsley (1962a, 1962b,
beetles have always been popular with collectors 1963, 1964) and Linsley and Chemsak (1972, 1976,
because of their bright colors and the large size of 1985), the North American fauna is taxonomically
adults of many species. The adults generally feed well known; only the second part of the subfamily
on wood, roots, leaves, and pollen. Flower-feeding Lamiinae remains to be done. Chemsak and Linsley
adults may visit many different blossoms, but some (1982) also published a checklist of the North
are host-specific; they are important plant-pollina- American cerambycids.
tion agents. The larvae bore into wood and roots;
the host wood may be living or thoroughly decayed. AK (31); YK(27); NT (32); BC (145); AB (88); SK
Some may even totally destroy timber for commer- (56); MB (100); ON (20e); PQ (178); NB (68); NS
cial purposes. Cerambycids play an important role (51);PE (11);LB (13);NF (24) . H (4);I (1)

Subfamily PARANDRINAE
Tribe Parandrini
Genus PARANDRA Latreille
(Subgenus NEANDRA Lameere)
P. brunnea brunnea (Fabricius) oN PQ
ampliceps Casey
conformis J. Thomson
dentata J. Thomson
grauidula Casey
minuta J. Thomson
purpureus (Herbst)
q uad.r icoll is J. Thomson
sayi J. Thomson
P. brunnea quebecensis Abdullah PQ

Subfamily PRIONINAE
Tribe Ergatini
Genus ERGATES Audinet-Serville
Trichocnemis LeConte
E. spiculatus spiculatus (LeConte) - BC
californicus (A. White)
spiculigerus (A. White)

Tlibe Prionini
Genus ORTHOSOMA Audinet-Serville
O. brunneum (Forster) oN PQ NS
amplians Casey
cy lindr icum (Fabricius )
cylindroides (Gmelin)
pennsyluanicum (DeGeer)
spadix Casey
unicolor (Drury)

Genus PRIONUS Geoffroy

(Subgenus PRIONUS s. str.)
(Prionellus Casey)
P. californicus Motschulsky AK BC
acomanus Casey
alutaceus Casey

277
Cerambycidee

ambiguus Casey
angustulus Casey
coloradensis Casey
compar Casey
consors Casey
crassict>rnis LeConte
curuatr^.s LeConte
fissifrons Casey
horni l.ameere
humerc:lis Casey
ineptus Casey
nanus i)asey
orbiceps Casey
ouipenr:is Casey
paruicollis Casey
proxim,zns Casey
punctui',atus Casey
scutellaris Casey
seniger Casey
solidus Casey
spaldingi Casey
spiculosus Casey
stultus Casey
suspectus Casey
terminulis Casey
texanus: Casey
uintanus Casey
P. laticollis Druryt oN PQ
b reu ico." nis Fabricius
densus Casey
frosti C'.asey
hempi Casey
nigrescans Casey
oblongu.s Casey
paruus Casey
P. lecontei I,ameere BC
P. pocularis Dalman oll pe -
bicolor Casey
curticornis LeConte
laeuigatus Hanis
obliqui:ornis LeConte
prolixus Casey
ualidiceps Casey
(Subgenus I{EOPOLYARTHRON Semenov)
(Riporuts Casey)
P. imbricorrris (Linn6) ON
brunneus Casey
cuneatus Casey
diuersus Casey
mimus Casey
robustu,s Casey
(Subgenus III.{TENNAJ,IA Casey)
P. fissicorni s Haldeman ON
paruiceps Casey
thoracicus Casey
transuersus Casey

Tribe Tragosomini
Genus TRATIOSOMA Audinet-Serville
T. depsariurn (Linn6)* NT BC SK MB ON PQ NB NS NF
harisii LeConte
paruicc'lle Casey
repens Casey
sodale Casey
spiculu.tn Casey

278
 Cerambycidae

Subfamily SPONDYLINAE
Genus SPONDYLIS Fabricius
S. upiformis Mannerheim AK BC AB ON PQ
basalis Casey
collaris Casey
laticeps LeConte
parua Casey
robustula Casey
subpubescens Casey

Subfamily ASEMINAE
TYibe Asemini
Genus ARHOPAJ-US Audinet-Serville
Criocephalum Dejean
Criocephalus Mulsant
A. asperatus (LeConte) NT BC AB SK
filitarsis (Casey)
A. foveicollis (Haldeman) AK YK NT AB SK MB ON PQ NB NS LB NF
agrestis (Kirby)
ceruinus (Casey)
lacustrinus (Casey)
A. productus (LeConte) BC AB
elongatus (Casey)
propinquus (Casey)
A. rusticus obsoletus (Randall) ON
mexicanus (J. Thomson)
obscurus (LeConte)

Genus MEGASEMUM Kraatz

M. asperum (LeConte) BC AB
impressum (Casey)
rectipennis (Casey)
uteanum (Casey)

Genus ASEMUM Eschscholtz

Liasetnum Casey
A. australe LeConte PQ
c ham p lainianum (C asey)
A. caseyi Linsley BC
A. nitidum LeConte BC
mokelumne Casey
A. striatum (Linn6)8 YK NT BC AB SK MB ON PQ NB NF
amputatum Casey
atrum Eschscholtz
breuicorne Casey
brunneum Haldeman
carolinum Casey
costulatum Casey
curtipenne Casey
ebenutn Casey
fuluipenne Casey
fuscutn Haldeman
gracilicorne Casey
juuencum Haldeman
moestum Haldeman
obsoletum Haldeman
paruicorne Casey
pugetanum Casey
stoctonense Casey
substriatum Haldeman

Genus TETROPIUM Kirby

T. abietis FaIl -BC
T. cinnamopterum Kirby AKYKNTBCAB_MB ON PQ NS-- NF
T. parvulum Casey AKYKNTBCAB_MB ON PQ LB
alaskanumFall

279
Cerambycii.ae

T. schwarzilnum Casey
T. velutinurn LeConte
boreak Casey
---ncAB--oNPQ----y
columlianum Casey
hexagonum Casey

Tlibe Atimiini
Genus ATItr{IA Haldeman
A. confusa confusa (Say) - ON
frislls l{aldeman
A. confusa corsalis LeConte - BC
A. hoppingi Linsley - BC

Subfamily CERAMBYCINAE
Tribe Opsimini
Genus OPSIMUS Mannerheim
O. quadrilineatus Mannerheim AK BC
biplectralis Casey

Genus DICI)NTRUS LeConte

Paraopsimus Champlain & Knull
D. bluthner LeConte - BC

TYibe Smodicini
Genus SMODICIIM Haldeman
S. cucujiforrre (Say) - ON
conuergens Casey
cy lindr;;d.e s (Newrnan)

Tlibe Hesperophanini
Genus TYLONOTUS Haldeman
T. bimacular,us Haldeman MB ON PQ

Genus HESIIEROPHANES Mulsant

AnoplitLm Haldeman
H. pubescens {Haldeman) - ON PQ

Genus EBU:RIA Audinet-Serville

E. quadrigerninata (Say) - ON

Tribe Elaphidiini
Genus PSYI|,ASSA Pascoe
PseudiL,idion Casey
P. unicolor (.Randallt - ON PQ - NS
p loch ionoc era (Haldeman)

Genus STEIIOSPHENUS Haldeman

S. notatus (Olivierr - ON pQ
deflend,zs (Newman)
discicol,:!.is (Dej ean)
discoiderus (Sturm)
pinorunz Casey

Genus ENAIIIIALODES Haldeman

Hy pe rm a ll us Lacordaire
Romaleum A. White
Thersalus Pascoe

280
 Cerambycidae

E. atomarius (Drury) ON
bispinosus (Pascoe)
mary landicus (Fabricius)
multicolor (Gmelin)
p ulu er ule ntus (DeGeer)
thoracicurn (Casey)
uariegatus (Goeze)
E. cortiphagus (Craigheac, ON
E. rufulus (Haldeman) ON PQ
crinicornis (Casey)
mancus (Casey)

Genus PARELAPHIDION Skiles

Key to species: Skiles (1985)
P. incertum (Newman) ON
breuicorne (Casey)
neglectum (LeConte)
obsolescens (Casey)

Genus ANELAPIIUS Linsley

E I ap hidionoides Linsley
A. parallelus (Newman) MB ON PQ
orclzs (Newman)
canadensis (Casey)
defectus (Casey)
densicollis (Casey)
medialis (Casey)
molliculus (Casey)
oblitus (LeConte)
p ulu erulentus (Olivier)
A. villosus (Fabricius) MB ON PQ
abruptus (Casey)
breuiusculus (Casey)
lacustris (Casey)
militaris (Casey)
p r uinos u s (Gu6rin-M6neville)
putator (Peck)
scuticularis (Casey)

Genus ELAPHIDION Audinet-Serville

Cycliopleurus Hope
E. mucronatum (Say) ON
muriaticum (Haldeman)
muricatum Haldeman
neb ulosum Gu6rin-M6neville
uestitum Haldeman

Tlibe Ibidiini
Genus HETERACHTHES Newman
Heteracthon J. Thomson
Key to species: Martins (1970)
H. quadrimaculatus Haldeman - ON PQ
euanescens (J. Thomson)
fluuiatilis Casey
pallidus Haldeman

Tlibe Obriini
Genus OBRIUM Dejean
Diozodes Haldeman
Phyton Newman
O. maculatum (Olivier)
howdeniRnrTl
limum (Newman)
pallidum (Say)
O. rufulum Gahan MB ON PQ

28r
Cerambycide.e

Tlibe Hyboderini
Genus HYBODERA LeOonte
H. tuberculaua LeConte _BC
Genus LAMPROPTERUS Mulsant
Callitne.llum Sftand
Callitnus Mulsant
Pilema.LeConte
L. ruficollis (LeConte) _BC
longicol'.is (Casey)
opacipennis (Casey)

Tfibe Eumichthinr
Genus EUMI.CHTHUS LeConte
E. oedipus LcConte - BC
ater Hardy & Preece
funebrus (Van Dyke)
malkini (Fisher)
ruber H,*dy & Preece

Genus POECIILOBRIUM Horn

P. chalybeunr (LeConter - BC
gibsoni [Iopping
minuturn Casey
rugosipt:nne Linell

Tlibe Molorchini
Genus MOLORCIIUS Fabricius
Glaphyra Newman
Laphyrt,. Newman
M. bimaculal;us bimaculatus Say AB SK MB ON PQ
affinis l.eCorrte
obscuru:; (LeConte)
M. longicollis LeConte - BC

Tribe Stenopterini
Genus CALLIMOXYS Kraatz

C. sanguinicollis luscipennis - BC
(LeCcnte)
C. sanguinic,rllis sanguinicollis - ON PQ
(Olivier)
pinorurr;, Casey

Tlibe Rosaliini
Genus ROSALIA Audinet-Serville
R. funebris l\lotschulsky AK BC
acorrrano, Casey
mandibitlaris Casey

Tlibe Callidiini
Genus PRONOCERA Motschulskv
Gonocallus LeConte
P. coilaris coilaris (Kirby) AK YK NT BC AB SK MB ON PQ NB - NF
lepida (LeConte)

Genus HYLOTRUPES Audinet-Serville

282
 Cerambycidae

H. bajulus (Linn6)t PQ
bullatus Haldeman
inaequalis Casey
incertus Casey
latus Casey
minutus Casey
nematocerus Casey
pedicellatus Casey
puncticollis Casey

Genus SEMANOTUS Mulsant

Anocomis Casey
Hemicallidium Casey
S. amethystinus (Leconte) _BC
S. ligneus amplus (Casey) AK BC
angustus (Casey)
obliuiosus (Casey)
terracensis (Casey)
S. ligneus basalis (Casey) BC
thujaeYanDyke
S. ligneus ligneus (Fabricius) MB ON PQ NS
nicolas (A. White)
russicus (Stephens)
S. litigiosus (Casey) NT BC AB SK MB ON PQ NB
canadensis (Casey)
terminatus (Casey)

Genus MERIELLUM Linsley

Merium LeConte
M. proteus (Kirby) AK YK NT BC AB SK MB ON PQ
bifossulatum (Casey)
simile (I{trby)

Genus CALLIDIUM Mulsant

Meridion des Gozis
C. antennatum hesperum Casey -BC
C. cicatricosum Mannerheim AKYK_BCAB
subopacum Swaine
udncouuerense Van Dyke
C. frigidum Casey oN PQ
lacustre Casey
C. hoppingi Linsley
C. schotti Schaeffer
C. violaceum (Linn6)t NN NS NF
janthinum LeConte

Genus PHYMATODES Mulsant

Microcallidium Casey
Phymatod,ina Casey
Poecilium Fairmaire
P. aeneus LeConte BC
P. aereus (Newman) oN pe
pallipes (Haldeman)
P. amoenus (Say) oN PQ
coccineico llis (Haldeman)
hudsonicus Casey
P. ater LeConte _NS
P. decussatus decussatus (LeConte) BC
collaris Casey
harfordi Casey
latifasciatus Hardy & Preece
obliquus Casey
P. dimidiatus (Kirby) AK YK BC AB SK MB ON PQ NB
haltnii (Schon]nen)
manner h eimii (LeConte)
palliatus (Haldeman)
P. fulgidus Hopping _BCPQ
P. hirtellus densipennis Casey _BCAB
P. lecontei Linsley _BC

283
Cerambycidae

grandi; Casey
obscur,ts (LeConte)
P. maculico.lis LeConte - YK - BC PQ NB
elonga;:us Hopping
hard,yi (YanDyke)
P. nitidus f.eConte - BC
crucialis Casey
elegant Casey
exilis Qasey
P. testaceus tl-innerf - ON PQ
uariabilis (Linn6)
uentralis Haldeman
P. varjus tFabricius) - ON PQ
albofasciatus (Bland)
diffidens Casey
latipennis Casey
s emic ir c ular is (Blano.r

subfast:iellus (A. White)

uersicoJor (Gmelin)
P. vulneratrrs (LeConte) - BC
concinnus Casey
nigrescens Hardy & Preece

Genus PHYSOCNEMUM Haldeman

Dulari,ts J. Thomson
P. andreae tHaldeman) - ON
luscum (J. Thomson)
P. breviline,rm t Sayt MB ON PQ
angulatum Casey
compressipes Casey
densun't Casey
longitarse Casey
pictum Casey
P. violaceip'rnne HamilLon

Genus ROP AJ-OPUS Mulsant

Euryotr tera Horn
R. sanguini.:ollis (Horn r MB ON PQ NB

Genus ELA'IOTRYPES Fisher

E. hoferi Fir,her AB

Genus XYLTJCRIUS LeOonte

X. agassizi {Leconter - BC

Tribe Clytini
Genus MEC'.ACYLLENE Casey
Cyllene Newman
M. angulifera (Casey) AB SK MB
M. caryae (Gahan) -oN
M. decora (Olivier) AB_MB
cas teln,zui (Aurivillius)
chara (Say)
genitiuo (Casey)
horni (.A,urivillius)
infausta (LeConte)
hansana (Casey)
mediana (Aurivillius)
solida (Casey)
M. robiniae (Forster) _ONPQ
angulata (Fabricius)
flexuos'z (Fabricius)
picta (Drtry)

Genus GLYOOBIUS LeConte

G. speciosus (Say) _ONPQ
haiji (Gtay)

284
 Cerambycidae

Genus CAI-LOIDES LeConte

C. nobilis nobilis (Harris) MB ON PQ

Genus SAROSESTHES J. Thomson

S. fulminans (Fabricius) ON PQ

Genus XYLOTRECHUS Chevrolat

X. aceris Fisher _ONPQ
X. annosus annosus (Say) NT AB SK MB ON PQ NB
X. annosus emotus W.J. Brown -BC
X. colonus (Fabricius) MB ON PQ NB
agrestis (LeConte)
campestris (OIivier)
te rrnina n s (Fabricius )
upsilon (Stephens)
uillicus (Olivier)
X. integer (Haldeman) ON PQ NB NS PE
frosli Van Dyke
X. lengi Schaeffer _BC
X. longitarsis Casey NT BC AB
columbianus Casey
inflaticollis Casey
montanicus Casey
X. mormonus (LeConte) YK BC AB SK MB
X. quadrimaculatus (Haldeman) MB ON PQ NB NS---
X. sagittatus sagittatus (Germar) MB ON PQ
cylindrus Casey
pubescens (Haldeman)
salebrosus Casey

- Ns pn is
X. schaefferi Schott
X. undulatus (Say) AK Yr( Nr ec AB sx MB 3N FB Nr'
fuscus (Kirby)
interruptus (Castelnau & Go.y)
lunulatus (Kirby)
soyi (Castelnau & Gory)
undatus (Kirby)

Genus NEOCLYIUS J. Thomson

R ho p alop achys Chevrolat
R ho p alome ru s Chevrolat
N. acuminatus acuminatus (Fabricius) MB ON PQ NB
americanus (Gmelin)
aspericollis (Germar)
erythrocep halus (Fabricius)
N. balteatus LeOonte BC
N. caprea (Say) ON PQ
eleuatus (Castelnau & Go.y)
gibbicollis (Castelnau & Gory)
N. conjunctus (LeConte) BC
occiduus Casey
tularensis Casey
N. fulguratus Casey
confusus Van Dyke
N. mucronatus mucronatus ON
(Fabricius)
humeralis (Newman)
luscus (Fabricius)
tnaculosus (Gmelin)
strictulus Casey
N. muricatulus muricatulus (Kirby) AK YK NT BC AB SK MB ON PQ NB NS PE LB NF
&lr6yi Aurivillius
leucozonus (Castelnau & Gory)
longipes (Kirby)
N. provoanus Casey BC AB
basalis Van Dyke
basillaris Van Dyke

Genus CLYTUS Laicharting

C. canadensis Hopping BC

285
Cerambycirlae

C. margini<:ollis Castelnau & Gory PQ NB NS

carolinensis Casey
uespoides Haldeman
C. pacificus {Van Dyke) - BC
C. planifrons (LeConte.t - BC AB
C. ruricola (Olivier) AB SK MB ON PQ NB
catulus (Casey)
hammatus Say

Genus CLYTOLEPTUS Casey

C. albofasciatus (Castelnau & Gory) - ON PQ
gracilicornis Casey
frisfis Casev

Tribe Tillomorphini
Genus EUDERCES LeConte
Apelocera Chevrolat
Eplophorus Chevrolat
E. picipes picipes (Fabricius) -oNPQ
Tlibe Anaglyptini
Genus TILI,OCLYTUS Bates
T. geminat rs (Haldeman) - ON
duplicztus (LeConte)

Genus CYI';TOPHORUS LeConte

C. verrucosus (Olivier) AB SK MB ON PQ - NS

Genus MICROCLYTUS LeConte

M. compre-.sicollis (Castelnau - ON PQ
& Gory)
frosti Oasey
gibbulus (LeConte)
insinuans (Casey)
niger (LeConte)
M. gazellula (Haldeman) - ON

Tlibe Purpuricenini
Genus PUI|PURICENUS Dejean
P. humeralis lFabricius.t MB ON PQ

Genus KNIILLIANA Linslev

Chion Newman
K. cincta cincta tDrury) - ON PQ
baltea;la (DeGeer)
filicontis (Casey)
garganica (Fabricius)
q uad r ii.sp inosa (Haldeman)
ruricoia (Gmelin)
rzsficcr (Fabricius)

Genus TRAGIDION Audinet-Serville

Trichascelis Bates
T. coquus ( Linn6t
apicat.um Casey
filiconte Casey
fuluipenne (Say)
leuipes Casey
lynceum (Fabricius)
melsheimeri (Germar)
nubiferum Casey

286
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Genus BATYLE J. Thomson

Batyleoma Casey
B. suturaiis suturalis (Say) AB SK ON PQ
fontinalis (Casey)
miniata (Germar)
rubro (Newman)
B. suturalis pearsalli (Bland) _ABSKMB
B. ignicollis ignicollis (Say) _ ^AB
acotnana Casey
punctipennis Casey
sang uinicoll is ( Germar)
seclusa Casey

Genus CROSSIDIUS LeConte

Oxoplus LeConte
C. carolinus (LeConte) SK
C. discoideus discoideus (Say) AB
pulchrior Bland
C. discoideus blandi Casey -{B
C. pulchellus LeConte AB
debilis Casey
lanaticollis Casey
nitidicollis Casey
palmeri Bates
retractus Casey
sinuosus Casey
uegasensis Casey
C. punctatus LeConte BC
crossipes Casey
hirsuticollis Casey
plagiatus Casey
uinosus Casey

Genus ELYTROLEPTUS Dug6s

E. floridanus (LeOonte) ON

Tribe Holopleurini
Genus HOLOPLEURA LeConte
H. marginata LeConte BC
helena LeConte

Subfamily LEPTURINAE
Tlibe Desmocerini
Genus DESMOCERUS Audinet-Serville
D. auripennis auripennis Chevrolat
D. auripennis cribripennis Horn
D. auripennis piperi Webb
D. palliatus (Forster) ON pe NB \TIT

blandus (Fabricius)
cyaneus (Fabricius)
elongatus Bland

Mbe Necydalini
Genus NECYDAI-IS Linn6
N. cavipennis LeConte BC
N. diversicollis diversicolhs BC
Schaeffer
N. laevicollis LeConte BC
seminiger Linsley
N. mellita (Say) ON
annericanus Haldeman

287
CerambyciJae

Genus ULOCIIAETES LeOonte

U. leoninur; LeConte BC

Tlibe Lepturini
Genus CEI{TRODERA LeConte
Parapachyta Casey
C. decolorata (Harris) oN PQ NB NS
lacustris Casey
rubidct (Haldeman)
C. spurca (LeOonte) BC
ceruin.z. (Walker)

Genus STEiNOCORUS Fabricius

Anisorus Mulsant
Eutoxotus Casey
Toxotc,psis Casey
ToxottLs Dejean
S. cylindricollis (Say)
atratus (Haldeman)
dentip ennis (Haldeman)
diues (Newman)
S. fl avolineatus (LeConte) _BC
S. nubifer (LeConte) _BCAB
apiciu,:ntris Casey
hesperus Casey
lateratis (Casey)
margi:tellus Casey
pacifit us Casey
paruiceps Casey
plagiatus Casey
rufipe,znis Casey
spinosus (Hopping)
tenellt..s Casey
truncctulus Casey
S. obtusus .LeConte) BC AB SK
breuicollis Casey
S. schaumii(LeConte) MB ON PQ
croceus (Leng)
S. trivittatus (Say) _ABSK MB
S. vestitus (Haldeman) BC AB
ater (L,eng)
aureatus Casey
flaccia'.us Casey
giluicarnis Casey
oregonensis Casey
sericai.us Casey
subpir;guis Casey
uirgat,rc (LeConte)
S. vittiger (Randall) oN PQ
nigrip,zs (Haldeman)

Genus ENCYCLOPS Newman

E. caerulea (Say) ON PQ
pallipes Newman

Genus PYR,OTRICHUS LeConte

P. vitticollis LeConte BC
cribripennis Casey

Genus PACHYTA Dejean

Argaleus LeConte
Linsle;lana Podany
P. armata LeConte _BC
P. lamed lit,urata Kirby AK YK NT BC AB meoNpe: LB NF
amerit:ana Podany
confl ag1 r ata Motschul sky
nitens (LeConte)

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Genus EVODINUS LeConte

E. monticola monticola (Randall) MB ON PQ NB NS LB NF
carolinensis Casey
E. monticola vancouveri Casey AK NT BC

Genus ANTHOPHYLAX LeOonte

A. attenuatus (Haldeman) oN PQ NB NS
A. cyaneus tHaldeman) oN PQ NB
malac hitic u s (Haldeman)
A. viridis LeConte -PQ NS
uiridipennis Casey

Genus NEANTHOPHYLAX Linsley & Chemsak

N. mirificus (Bland) BC AB
Denustus (Bland)
N. tenebrosus nigrolineatus BC
(Van Dyke)

Genus RHAGIUM Fabricius

Allorhagium Kolbe
Hargium Samouelle
Stenocorus Geoffroy
R. inquisitor (Linndtx YK NT BC AB SK MB ON PQ NB NF
americanurn Podany
boreale Casey
californicum Casey
canadense Podany
cariniuentre Casey
crosslpes Casey
i nuesliga lor Mannerheim
lineatus (Olivier)
mexicanurn Casey
montanum Casey
paruicorne Casey
quadricostatum Podany
thoracicum Casey

Genus SACHALINOBIA Jacobson

Pseudopachyta Swaine & Hopping
rugipennis (Newman)
S. rugipennis MB ON PQ NB

Genus GAUROTES LeConte

G. cyanipennis tSayt ON PQ
ioze (Newman)
lapor te i ( Gu6rin-M6neville)
leonardi (Haldeman)
ser u illei (Audinet-Serville)

Genus ACMAEOPS LeOonte

A. proteus proteus (Kirby) AK YK NT BC AB SK MB ON PQ NB LB
aurora Casey
cauicollis Casey
coloradensis Casey
gibbula LeConte
lacustrina Casey
obsoleta Casey
parkeri Casey
pinicola Schaeffer
puncticeps Casey
scutellata Casey
sublineata (Haldeman)

Genus LEPTALIA LeConte

L. macilenta (Mannerheim) AK BC AB
frankenhaeuseri (Mannerheim)
fuscicollis (LeConte)

289
Cerambycidae

Genus CO IITODERA Mulsant

Acma,zopsilla Casey
Lepta.:maeops Casey
C. coniferae Hopping & Hopping - BC
C. longicornis {Kirbyt - BC AB SK
alticoia (Casey)
6oher,' Hopping & Hopping
dorsa,',is (LeConte)
harneyensis Hopping & Hopping
ligata (LeConte)
marginalis (LeConte)
oregonensis Hopping & Hopping
punctiuentris (Casey)
suturalis (Casey)
uincta (LeConte)
C. militari'r militaris(LeConr,eJ - BC
C. subpilosa (Leconte) - BC AB
intermedia (Casey)
lupino (LeConte)
pugetuna (Casey)
quadriceps (Casey)

Genus PSEIIDOGAUROTINA Plavilstshikov

P. abdomirralis (Bland I - ON PQ
P. cressoni cressoni (Bland) - BC AB

Genus BR.I\CHYSOMIDA Casey

B. atra (Leconte) - BC AB
aterriina Casey
morata Casey
B. bivittata (Say) AB SK MB ON PQ NB NS
fuscicttps (LeConte)
incert r (Bland)'
nigrip,ennis (LeConte)
oblonga (Casey)
uariarts (LeConte)

Genus GN.{.THACMAEOPS Linsley & Chemsak

G. pratens s (Laicharting)* AK YK NT BC AB SK MB ON PQ NB LB NF
fu lu i p ennis (Mannerheim)
longiceps (Kirby)
s em i - marg inara (Randall)

s t rig il ata (Fabricius)

Genus ME'IACMAEOPS Linsley & Chemsak

M. vittata Swederus) PQ
direcfrz (Newman)
pallidc (Haldeman)
q uadr iu ittata (Schonherr)

Genus STIIANGALIA Audinet-Serville

Ophistomis J. Thomson
S tr anil alina Auriviilius
S. acuminz.ta (Olivier) - ON
emaciata Newman
unicolor Haldeman
S. bicolor (:Swederus) - ON
simulons (Casey)
S. famelica solitaria Haldeman - ON
S. luLeicorrris (Fabriciust - ON PQ
carolinae (Casey)
euersa (Casey)

Genus BELLAMIRA LeConte

B. scalaris (Say) - SK MB ON PQ NB NS
coarcto.ta (Haldeman)

290
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Genus ANALEPTURA Linsley & Chemsak

A. lineola (Say) - ON PQ NB
cincta (Haldeman)
indirecta (Newman)
lateralis (HaIdeman)
obsoleta (Haldeman)

Genus CHARISALIA Casey

C. americana (Haldeman) - ON

Genus GRAMMOPTERA Audinet-Serville

Parallelina Casey
G. exigua (Newman) - ON PQ
saucia (LeConte)
G. haematites (Newman) - SK MB ON PQ NB
zono (Newman)
ruficeps (LeConte)
G. molybdica {Leconte) - BC
G. subargentata (Kirby) AK YK NT BC AB SK MB ON PQ NB
filicornis (Casey)
quebecensis (Couper)
rufi.basis (LeConte)
similis (I\trby)

Genus JUDOLIA Mulsant

J. gaurotoides gaurotoides (Casey) AB SK
hnulli (Swaine & Hopping)
uiuaria Casey
J. instabilis (Haldeman) - YK - BC AB SK MB
conuexa (LeOonte)
fl au iu entris (Schaeffer)
pacifica Casey
trajecta Casey
J. montivagans montivagans(Couper) AK YK NT BC AB SK MB ON PQ NB NS - LB
seminigra Casey
J. quadrata (LeConte) - YK NT BC - SK MB ON PQ
Genus PYGOLEPTURA Linsley & Chemsak
P. carbonata (LeConte) - BC
P. nigrella nigrella(Say) AK - NT BC AB SK MB ON PQ NB LB NF
lacustris (Casey)
nigrita (HaLch)
praestans (Casey)
serricornis (Casey)

Genus DORCASINA Casey

D. matthewsi (LeConte) - BC
macrocera (Casey)

Genus IDIOPIDONIA Swaine & Hopping

l. pedalis (LeConte) - ON PQ NB NS

Genus TYPOCERUS LeConte

T. acuticauda acuticauda Casey - ON PQ NB NS
thoracicus Casey
T. balteatus Horn NT - AB
T. confluens Casey MB
caligans Casey
T. lugubris tSayt - ON
T. octonotatus (Haldeman) - ON
arapahoe Casey
sinuatus LeConte
T. sparsus LeConte MB ON PQ
T. velutinus velutinus (Olivier) MB ON PQ NB NS
fugo* (Fabricius)
manitobensis Swaine & Hopping
tenuior (Kirby)

297
Cerambycidae

T. zebra (Olivier) ON
aurige" (Newman)
carolir,La (Weber)
zebratus (Fabricius)

Genus NEOALOSTERNA Podany

N. capitata {Newman)
N. rubida (l,eConte)
heeni (Casey)
BC _9* PQ NB NS

Genus PSEUDOSTRANGAIIA Swaine & Hopping

P. cruentat a (Haldemant ON PQ
maneii'. (Casey)

Genus LEPTURA Linn6

(Subgenus LEPTURA s. str.)
L. obliterata obliterata (Haldeman) BC
idahoensis (Casey)
perductor (Walker)
uitiosa (LeConte)
L. plagifera LeConLe BC
L. plebeja Flandall
L. propinquaBland AK BC
yuoNpeNe_
minuscula (Casey)
regularis (Casey)
L. subhami.ta Randall ON PQ NB NS
annatfl (Haldeman)
elegan; (Haldeman)
interrt..pta Newman
(Subgenus iTENURA Dejean)
(Mega,i.eptura Casey)
L. emargin,rta Fabricius ON

Genus STENOSTROPHIA Casey

S. amabilis {Leconte) BC
S. tribalteaba serpentina (Casey) BC

Genus TRACIfYSIDA Casey

T. aspera aspera (LeConte) AK YK NT BC AB
parkeri (Hippisly)
T. aspera b:revifrons (H. Howden)
T. mutabili; (Newman)
I ae tifi<.: ( Provancher)
scABixreBNiSNENS- - NF

I ur idi 1., enni s (Haldeman)

prou at;.cher i (Aurivillius)
quadncollis (LeConte)

Genus PIDIfNIA Mulsant

(Subgenus .PIDONIA s. str.)
(Hapai,osalia Casey)
P. ruficollis {Sayr ON PQ NB NS
alleclo (Newman)
fragilis (Casey)
laeuiceps (Casey)
Iineicornis (Casey)
nitidicollis (Horn)
paupercula (Newman)
sphaericollis (Say)
straus:;i (Webb)
uiber (Newman)
P. scripta (l,eConte) AK BC
(Subgenus'IHESAJ-IA Casey)
P. gnathoides (LeConte t BC
lisa (Leng)
rubriceps (Casey)

292
 Cerambvcidae

(Subgenus CERATOPIDONIA Linsley & Chemsak)

P. quadrata Hopping - BC

Genus ORTHOLEPTURA Casey

O. valida (Leconte) - BC
oculea Casey

Genus STRANGAI-EPTA Casey

S. abbreviata (Germarr - ON PQ NB NS
gulosa (Kirby)
nitidipe nnis ( Provancher)
saratogensis (Rat)
semiuittata (Kirby)
S. pubera tSay) MB ON PQ

Genus STICTOLEPTURA Casey

S. canadensis canadensis (Olivier) MB ON PQ NB NS PE - NF
c innamop te ra (Haldeman)
erythroptera (Kirby)
te nuic or n is (Haldeman)
S. canadensis cribripennis (LeConte) - BC AB SK MB
coccinea (LeConte)
diuisa (Casey)
ebena (Leng)

Genus ANASTRANGAJ-IA Casey

A. laetifica (LeConte) - BC
lugens (LeConte)
A. sanguinea (LeOonte) AK YK - BC AB SK MB ON PQ NB
apicata (Casey)
bould.erensis (Casey)

Genus XESTOLEFTURA Casey

X. behrensi (LeConte) - BC
X. crassipes (LeContet - BC
fas ciu en tr is ( LeConte)
muliebris Casey
shastana Casey
uancouueri Casey
tanthogaster (LeConte)
X. octonotata (Sayt - ON
q uadr ip unctato ( Haldeman)
stictica (Newman)
X. tibialis (LeConte) - \'K - BC AB ON PQ - NS - LB NF
columbica Casey
hirtella (LeConte)
miquelonensis (Pic)
pictipennis Casey

Genus LEPTUROPSIS Linsley & Chemsak

L. biforis (Newman) - ON PQ NB
laurentica (Casey)
L. dolorosa (Leconte) - BC

Genus TRIGONARTHRIS Haldeman

Cercolia Casey
T. minnesotana (Casey) MB ON PQ NB NS
T. proxima (Sayt MB ON PQ - NS
hempiana (Casey)
subpubescens (Kirby)

Genus ACMAEOPSOIDES Linsley & Chemsak

A. rufulus (Haldeman) - BC AB SK MB ON PQ - NS

Genus BRACHYLEPTURA Casey

B. champlaini Casey - ON PQ - NS PE

293
Cerambycidae

B. dehiscens (LeConte) BC
B. rubrica ([iay) ON PQ NB
erythraptera (Germar)
B. vexatrix, Mannerheim) AK BC
conuoluens (Casey)
quadrii,lum (LeConte)

Genus STROPHIONA Casey

S. laeta (Le(lonte) BC
ostenta Casey
S. nitens (Forster) MB ON PQ NS
bellina Casey
quagga (Germar)
zebra (t)livier)

Genus COS.MOSAJ-IA Casey

C. chrysocorna (Kirby) YK NT BC AB SK MB ON PQ NB NS PE NF
aureolc,. (Casey)
auripilis (LeConte)
densep,'.losa Casey

Subfamily LAMIINAE
Tlibe Parmenini
Genus PLEOTRURA Mannerheim
P. spinicauc.a Mannerheim AK BC
produc.la LeConte

Tlibe Moneileminr
Genus MO]TIEILEMA Say
Collapt:eryr Newman
M ono p,'.e sa Motschulsky
Omosc"ilon J. Thomson
M. annulatrLm Say AB SK
angulatumPsota
demissutn Casey
hybridum Psota
montanum Psota
nubeculum Casey
puncticolle Psota

Ttibe Monochamini
Genus HEBESTOLA Haldeman
Cacoplt.a LeConte
H. nebulosa Haldeman PQ
albata (Knu]l)
pullato (Haldeman)
pruinosa (LeConte)

Genus MONOCHAMUS Dejean

M. carolinensis (Olivier) oN PQ NB
dentator (Fabricius)
minor (LeConte)
M. clamator' latus Casey BC
M. marmorirtor Kirby MB ON PQ NB NS
acutus (Lacordaire)
fautor (.LeConte)
M. mutator LeConte NT _SKMB oN PQ NB
M. notatus lDrury) NT BC AB SK MB oN PQ NB NS PE
confusor l{trby
morgani Hopping
M. obtusus rbtusus Casey _BC

294
 Cerambycidae

M. scutellatus scutellatus (Say) AK YK NT BC AB SK MB ON PQ NB NS PE LB NF

resutor Kirby
M. scutellatus oregonensis (LeConte) BC AB
monticola Casey
M. titillator (Fabricius) -oN
angusticollis Casey
obesus Casey

Genus MICROGOES Casey

M. oculatus (LeConte) -ONPQ
tenuicornis Casey

Genus GOES LeConte

G. debilis LeConte ON PQ
G. pulcher (Haldeman) oN PQ
fuscatus IKntTl
G. pulverulentus (Haldeman) ON PQ
laurenticus Casey

Tribe Mesosini
Genus SYNAPHAXTA J. Thomson
S. guexi (LeConte) - BC
annulata Casey
breuicollis Casey
hutneralis Casey

Tlibe Dorcaschematini
Genus DORCASCHEMA Haldeman
Hetoemis Haldeman
D. nigrum (Say) oN PQ

TYibe Apodasyini
Genus EUPOGONIUS LeConte
Eriopsilus Bates
Phidola J. Thomson
E. pauper LeConte _ONPQ
frafiniKnull
uestitus (Say)
E. subarmatus (LeConte) _ONPQ
E. tomentosus (Haldeman) _ONPQ
piniuora Filc}r

Genus PSENOCERUS LeConte

Paralarica Breuning
P. supernotatus (Say) MB ON PQ NB
funebris Casey
picfzs (Breuning)
pini LeConte
frislis Casey

Tlibe Pogonocherini
Genus POLIAENUS Bates
P. oregonus (Leconte) - BC

Genus ECYRUS LeOonte

Oebaceres J. Thomson
E. dasycerus dasycerus (Say) - ON PQ
exiguus (Haldeman)
obscurus (Haldeman)

295
 Cerambycirlae

Genus LOI'HOPOGONIUS Linsley

L. crinitus LeConter - BC

Genus POCiONOCHERUS Dejean

(Subgenus .POGONOCHERUS s. str.)
P. penicillalus Leconte AK - NT BC AB - MB oN pe NB NS pE - NF
alashanus Schaeffer
carinatus Casey
(Subgenus .OUPOGONOCHERUS Linsley)
P. mixtus l:laldeman - yK NT BC AB - MB ON pe NB
simple.x LeConte
P. parvulus LeConte AB SK MB ON pe
salicol'r Casey
P. pictus Fi,ll NT BC AB
emarginatus Casey
fastigiutus Casey
P. propinquus Fall - BC

Tribe Onciderini
Genus ONCTIDERES Lepeletier & Audinet-Serville
O. cingulatrrs cingulatus rsayr - ON
pallesczns Casey
praecit:.tens Casey

Tribe Hippopsini
Genus HIPI)OPSIS Lepeletier & Audinet-serville
H. lemniscata tFabriciusr - ON
lineata Gt6rin-M6neville
lineola la Audinet-Serville
meinerli Aurivillius

Tribe Acanthoderini
Genus AXGIMORPHUS Haldeman
Aethioltoctines J. Thomson
A. modestus {Gyllenhal } - - MB ON pe
decipie:zs Haldeman
A. morrisi (lJhler)
leucogenus (J. Thomson)
A. quadrigit,bus tSayr
lucidus (Knull)
- ON

Genus OPLOSIA Mulsant

O. nubila (LeContet - ON pe

Tribe Acanthocinini
Key to North American species: Dillon (1956)
Genus CANIfNURA Casey
C. princeps Walkerr - BC
uincta (Casey)

Genus NEACANTHOCINUS Dilion

N. obliquus,rbliquus (LeConter AB
acornanLls (Casey)
angulosus (Casey)
sedulus (Casey)
N. obliquus racificus (Casey) - BC
N. obsoletus (Olivier) - ON pe
floridar.:us (Casey)
N. pusillus (Kirby) AK - AB SK MB ON pe NB NS

296
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Genus CERATOGRAPHIS Gahan

C. biguttata (LeConte) PQ

Genus GRAPHISURUS Kirbv

Urographis Horn
G. despectus (LeConte) - ON PQ
hebes (Casey)
G. fasciatus (DeGeer) - ON PQ
mirlus (Fabricius)
anicus ( Gmelin)
p enn sy lu
reductus (Casey)

Genus AMNISCUS Haldeman

Astylopsis Casey
A. collaris Haldeman - ON PQ
intenuptus Haldeman
A. macula (Say) - ON PQ NB NS
sticticus Haldeman
A. sexguttatus (Say) MB ON PQ
commixtus Haldeman
guttatus (Say)

Genus LEPTOSTYLUS LeConte

L. transversus transversus - ON PQ
(Gyllenhal)
aculiferus (Say)
rnarg inell us (Haldeman)
t ub erculatu s (Frolich)

Genus STERNIDIUS LeConte

Key to North American species: Lewis (1986)
S. alpha(Say) - ON PQ
cinereus (LeConte)
coloradensis DilIon
diuergens (Haldeman)
lateralis (Haldeman)
nelsonicus (Casey)
suturalis Dillon
uicinus (Haldeman)
S. fascicularis (Harris) - ON PQ - NS
dentatus (Casey)
nigricans Dillon
obscurellus (Casey)
pleuralis (Casey)
scapalis (Casey)
testaceus (Casey)
timid,us (Casey)
xanthoxyli (Shimer)
s. misellus (Leconte) MB oN PQ
S. rusticus (LeConte) MB

Genus ASTYLEIOPUS Dillon

A. variegatus (Haldeman) AB SK MB ON PQ NB
obscurus (Haldeman)
tr ifa sciatu s ( Haldeman)

Genus DECTES LeConte

D. sayi Dillon & Dillon AB - MB ON
spizosus (Say)

Genus LEPTURGES Bates

(Subgenus LEPTURGES s. str.)
L. confluens (Haldeman) - ON PQ
L. symmetricus (Haldeman) - ON PQ

297
Cerambycidae

Genus URGLEPTES Dillon

U. facetus (liay) -ONPQ
U. querci (Fitch) _ONPQ
/risfis (Casey)
U. signatus (LeConte) -oNPQ
tenebrosus (Casey)

Genus HYPERPLATYS Haldeman

H. aspersa (Say) BC AB SK MB ON PQ NB NS NF
uigilans Casey
H. maculata Haldeman _MBONPQNBNS
amnicola Casey
binocul.aris Casey
cryptiat Casey
delicatr:.t Casey
frigida Casey
lentigit;.osa Casey
nigrellcr Haldeman
robustula Casey
uariolata Casey
H. montana Casey BC AB SK MB

Tlibe Cyrtinini
Genus CYR|CINUS LeConte
Myrmo,bmia Bates
C. pygmaeu,r (Haldeman) ON

Tlibe Saperdini
Genus SAPI)RDA Fabricius
Eutetropha Bates
Key to species: Abdullah and Abdullah ( 1966)
S. calcarata Say BC AB SK MB ON PQ PE
adspersa LeConte
S. candida brpuncLata Hopping SK MB ON
S. candida candida Fabricius -oNPQ NS
biuittato Say
S. concolor unicolor Felt & Joutel SK MB ON PQ NB NS NF
mecasotdes Casey
S. cretata Nr)wman MB
S. discoidea Fabricius _ONPQ
dorsalit Haldeman
fuscipes Say
S. fayi Blan< oN PQ
S. horni Jou;el BC AB _MB
S. imitans F.:lt & Joutel oN PQ
S. lateralis F'abricius -MBONPQ NB NS
abbreuiata Fitch
connecte. Felt & Joutel
disconolata Pic
suturalis Fitch
transeuns Breuning
S. mutica Say MB ON PQ
S. obliqua Say MB ON PQ NB NS PE
S. populnea :moesta LeConte YK NT SC AB SK MB ON PQ LB
S. populnea'populnea Linn6 BC AB
S. puncticoll s Say BC -oNPQ
trigeminata Randall
S. tridentata Olivier SK MB ON PQ
dubiosa Haldeman
interme':lia Fitch
rubronotata Fitc}r
trifasciota Casey
S. vestita Say MB ON PQ
atkinsoni Crrtis

298
 Cerambycidae

Tribe Phytoecini
Genus OBEREA Mulsant
Amaurostoma J. Miilier
Isosceles Newman
Key to species: Breuning (1960)
O. appaiachiana Casey _ON-NB
O. bimaculata (Olivier) MB ON PQ
affinis Haris
basalis LeConte
basicollis Breuning
bicallosicollis Breuning
bipunctata (Creutzer)
broo&si Wallis
delicatula Casey
dolosa Casey
filum Casey
flauocephala Blatchley
insignior Breuning
insignis Casey
iowensis Casey
rnontana Casey
per spicill ata Haldeman
tr ip unctata ( Fabricius )
ulmicola Chittenden
umbra Casey
O. canadensis Fisher _SKMBONPQ
O. cylindricollis Casey -oN
O. deficiens Casey -ONPQ
O. delongi Knull -oNPQ
anter ufi collis Breuning
O. exilis Casey -oN
O. ocellata Haldeman -oN
plagiata Casey
O. oculaticollis (Say) MB ON
O. pallida Casey MB ON PQ
O. praelonga Casey _ONPQ
O. pruinosa Casey AB_ -oN
O. quadricallosa LeConte _BC
O. schaumii LeOonte -BC AB SK MB ON PQ NB
caseyi Plavilstshikov
ferruginea Casey
infrarufa Breuning
su bcy I ind r i collis Breuni ng
uittipennis Breuning
wapleri Chevrolat
O. tripunctata (Swederus) oN PQ NB
amabilis Haldeman
intermedia Casey
mandarina (Fabricius)
rnyops Haldeman
subdeficiens Breuning
subexilis Breuning
submondarina Breuning
texana Horn

Genus MECAS LeConte

(Subgenus MECAS s. str.)
M. bicallosa Martin BC

Tribe Tetraopini
Genus TETRAOPES Schiinherr
Key to species: Chemsak (1963)
T. annulatus LeConte AB-MB
canescens LeConte
fontinalis Casey
uteanus Casey
uestitus Casey

299
Bruchidae

T. femoratus LeConte BC AB SK MB ON
amnicc'la Casey
atrisetosus Casey
breuisetosus Casey
coseyj ,\urivillius
collaris Horn
fortis Casey
monticola Casey
nigripes Casey
oregonetnsis LeConte
Punctiltennis Qasst
robustt,s Casey
ruber (.Asey
spissicornls Casey
uegaser,,sis Casey
uelutin ^ts Casey
T. quinquemaculatus Haldeman ON
T. tetrophthalmus (Forster) -MBONPQ
humerc:Iis Casey
iowensis Casey
torndtor (Fabricius)
t r e dec im punc tatu s (Dr apiez)

Family BRUCHIDAE seed beetles

J. McNamara

The t ruchids comprise 19 species in Canada Borowiec (1987) published a key to the genera of
and Alasra. Their habits vary considerably, but the world, and Udayagiri and Raj Wadhi (1989)
they depend almost entirely on seeds for their published a catalog of the world species.
development. The larvae live in seeds of legumes,
notably l)eas and beans, and are often pests of
stored leguminous seeds. The adults are commonly
found in flowers and on foliage. The pea weevil
(Bruchus pisorum) and the bean weevil aK (1);YK (1); NT (2); BC (10); AB (8); SK (7); MB
(Acanthoscelides obtectus) are major pests of peas (e); ON (11); PQ (5); NB (2); NS (2); PE (1); NF (1)
and beans, and have spread around the world. . I(6)
Subfamily AMBLYCERINAE
Genus AMB LYCERUS Thunbere
A. robiniae ( Fabricius) AB
gleditsiae (Castiglioni)

Subfamily KYTORHININAE
Genus KYTORHINUS Fischer von Waldheim
K. prolixus (Fall) AK YK NT BC AB SK MB

Subfamily BRUCHINAE
Genus MEG.A.CERUS Fahraeus
Pachybruchus Pic
(Subgenus NTEGACERUS s. str.)
M. discoidus (Say) BC AB SK MB ON PQ NB NS

Genus CALI,OSOBRUCHUS Pic

C. maculatus (Fabricius) SK ON
ambiguzs (Gyllenhal)
quadrinzaculatus (Fabricius)
sinuatus (Fahraeus)

300
 Chrysomelidae

Genus GIBBOBRUCHUS Pic

Key to North American species: Whitehead and Kingsolver (1975)
G. mimus (Sayt ON
borealis (Schdnherr)
crataegi (Fahraeus)

Genus MEIBOMEUS Bridwell

Key to North American species: Kingsolver and Whitehead (1976)
M. musculus tSay) ON PQ
a lb og ut tis (Motschulsky)
erythrocerus (Riley)

Genus ACANTHOSCELIDES Schilskv

A. aureolus (Horn) _BC AB SK MB
A. calvus (Horn)
AB
-oN
MB ON
A. fraterculus (Horn) NT BC SK
A. obtectus (Say)t _BC MB ON PQ
fabae (Pi'iley)
u ar icor ni s (Motschulsky)

A. pauperculus (LeConte) BC
p ulicar iu s (Motschulsky)
rulzs (Motschulsky)
simplex (Motschulsky)
A. perforatus (Horn) AB _MB
A. pullus (FalI) BC
A. seminuium (Horn) AB SK MB

Genus BRUCHUS Linn6

B. brachialis Fahraeust _BC -oN
B. pisorum (Linn6)t _BCAB MB ON PQ NU NS PE NF
pisi Linn6
B. rufimanus (Boheman)t SK MB ON PQ

Genus BRUCHIDIUS Schilsky

Key to North American species: Bottimer (1968)
B. cisti (Fabricius)t BC
unicolor (Olivier)
B. villosus (Fabricius)f ON
oler (Marsham)

Family CHRYSOMELIDAE leaf beetles

L. LeSage

The family Chrysomelidae is one of the largest in remaining subfamilies are either root feeders, root
group of beetles in the world, with over 50 000 miners, stem miners, or leaf miners.
known species and many additional thousands to be Chrysomelidae is usually classifred into 19 sub-
discovered, especially from Asia and the Neotropical families (Seeno and Wilcox 1982) of which 14 occur
region; 569 species and subspecies are listed here for in Canada. The fauna of the leaf beetles is relatively
Canada and Alaska. All known adults are phy- well known in eastern Canada (except for some
tophagous. Primitive plants and ferns are used alticine, galerucine, and cryptocephaline genera) but
infrequently, and conifers are barely touched; decid- is poorly known in western Canada. The present list
uous trees, bushes, and weeds are preferred host- follows essentially the arrangement proposed by
plants. Some species are pollen eaters when in the Seeno and Wilcox (1982), except where the generic
adult stage. Larvae show a broader ecological diver- concepts have been recently modifred.
sity: Donaciinae are aquatic; Clytrinae live in ant
nests, where they feed on plant detritus and on ant
larvae or pupae; most Cryptocephalinae feed on
decaying matter in leaf litter; Criocerinae, AJ{ (42); YK (41); NT (72); BC (181); AB (217); SK
Chlamisinae, Chrysomelinae, and Cassidinae and (174); MB (25a); ON (363); PQ (27a); NB (116); NS
some Galerucinae and Alticinae feed on leaves: those (103);PE (19); LB (1a); NF (3a) o H (9); I (38)

301
Chrysome lidae

Subfamily DONACIINAE
Key to New World genera: Askevold (1990)
Tlibe Plateumarini
Genus PL,A.TEUMARIS C.G. Thomson
P. aurifera (LeConte) AK YK NT AB_MB PQ NF
P. balli Askevold PQ NB
P. chalcea (Lacordaire) _BC AB SK MB ON PQ NB NS PE
lodingi .Schaeffer)
s hoema,ize ri ( Schaeffer)
P. diversa (Schaeffer)
P. dubia (Schaeffer) AK--nCAB-yuoN-
idola (Hatch)
P. flavipes (Kirby) AK YK NT BC AB SK MB ON PQ NB LB NF
wallil;i (Schaeffer')
P. frosti (Schaeffer) NT BC AB SK MB ON PQ NB NS
P. fulvipes (Lacordaire) AK_NTBCABSKMBONPQNBNSNF
P. germari (Mannerheim) AK YK NT BC AB SK MB ON PQ NB NS _ LB NF
diues (LeConte)
fl au ip ennis ( Mannerheim)
serricauda (Schaeffer)
P. metallir,a (Ahrens) ON PQ NB NS
catar'rctae (Newman)
femor alis (Kirby)
gentitis (Leconte)
indicu (Melsheimer)
zaza (Melsheimer)
parua: (Lacord.afte)
P. nitida (tlermar) BC AB SK MB ON PQ NB NS PE LB NF
emarSlinata (Kirby)
juncina (Couper)
paciftca (Schaeffer)
P. neomex icana (Schaeffer) BC
long it:' ol I i s ( Schaeffer)
u er mti. c ulat a ( Schaeffer)

P. pusilla Say) AX YK NT BC ff SK MB ON PQ NB NS LB NF
pyritt: sa (LeConte)
rugiftons (Newman)
P. robusta (Schaeffer)
P. rufa (Say)
affinis (I\trby)
_ y:'TSKyu oN
oN
PQ
PQ NB NS

jucunda (LeConte)
hirbyi (Lacordaite)
s ulcic ollis (Lacordaire)

Tribe Donaciini
Genus DO NACIELLA Reitter
D. pubicollis ( Suffrian) AB SK MB ON

Genus DONACIA Fabricius

(Subgenus DONACIA s. str.)
Key to l\lorth American species: Marx (1957)
D. cinctico:rnis Newman MB ON PQ NS
antillarum Suffrian
lucidl:, Lacordaite
pulch,zlla LeConte
rufipennis Lacordaire
tenuis Schaeffer
tryphera Schaeffer
D. palmata Olivier ON PQ NB NS
angustipes Marx
claud;icans Germar
D. parvide.ns Schaeffer
D. piscatrir Lacordaire
-oN
MB ON PQ NB
alutacea LeOonte
carolina Lacordaire
conge;'Ler LeConte
cuprea Kirby

302
 Chrysomelidae

D. proxima Kirby NT BC AB SK MB ON PQ NB NS NF
californica LeConte
e pisco palis Lacordaire
D. rufescens Lacordaire ON PQ
D. texana Crotch ON
minor Schaeffer
(Subgenus DONACIOMIMA Medvedev)
D. biimpressa Melsheimer MB ON PQ
a ur ichalcea Melsheimer
torosa Le0onte
D. caerulea Olivier MB ON PQ NB
aequalis Say
confusa Lacordaire
confusa LeConte
D. canadensis Askevold AB SK MB ON PQ
D. cazieri Marx NT BC AB _ONPQ LB
D. confluenta Say _BC AB SK MB ON PQ
D. cuprea Kirby -SK -oNPQ
curticollis Knab
D. distincta LeConte AK YK NT BC AB SK MB ON PQ LB NF
occidentalis Mead
D. fulgens LeConte _SK MB ON PQ NB NS PE
D. hirticollis Kirby AK YK NT BC AB SK MB ON PQ NB NS_LBNF
rudicollis Lacordaire
D. limonia Schaeffer _ONPQ
D. magnifrca LeConte BC AB SK MB ON PQ NB
D. megacornis Blatchley -oN
megaloceraWeise
D. porosicollis Lacordaire NT BC AB SK MB ON
D. pubescens LeConte BCTSK MB ON PQ
D. rugosa LeConte -oN
D. subtilis Kunze BC AB SK MB ON PQ NS
aenea Ahtens
aerea Lacord.aire
magistrigata Mead
quadricollis Say
D. tuberculifrons Schaeffer BC MB ON PQ NB

Tribe Haemoniini
Genus NEOFIAEMONIA Sz6kessy
Key to North American species: Askevold (1988)
flagellata Askevold MB ON
N.
N. melsheimeri (Lacordaire) YSK MB ON PQ
N. minnesotensis Askevold MB ON
N. nigricornis tKirby) BC MB ON PQ
americana (Gu6rin-M6neville)

Subfamily ORSODACNINAE
Genus ORSODACNE Latreille
Orsodacna Dejean
O. atra (Ahrens) YK NT BC AB SK MB ON PQ NB NS
anneniacae Germar
childreniKirby
hepatica Say
inconstans Newman
luctuosa Lacordaire
ruficollis Newman
tibialis Iltrby
tricolor Melsheimer
triuittata Lacordaire
uittata Say

303
 Chrysomelidae

Subfamily ZEUGOPHORINAE
Genus ZEIIGOPHORA Kunze
(Subgenus ZEUGOPHORA s. str.)
(Auchenia Thunberg)
(Tararis LeConte)
Z. abnormis (LeConte) AK YK NT BC SK MB ON PQ
hirbyi Baly
reineckei Gtote
Z. atraFall YK NT BCABSKMB_PQ
Z. consanguinea Crotch BC AB SK
Z. multnomah Hatch YK BC_ON
Z. puberula Crotch
Z. scutellaris Suffriant
-Nr ON
BC AB SK MB ON PQ
Z. varians Crotch BC AB SK MB ON PQ

Subfamily CRIOCERINAE
Tlibe Criocerini
Genus LIL IOCERIS Reitter
(Subgenus LILIOCERIS s. str.)
L. lilii (Scopoli)t oN PQ
lilioru,m (C.G. Thomson)
melanocephala (Say)
(Fabricius )
m e r di,;1 e r a

Genus CRIOCERIS O.F. Mriller

Pleurcphora Chewolat
C. asparagr (Linn6)t _BCABSKMBONPQ NB
C. duodecirnpunctata (Linn6)f _BCABSKMBONPQ NB

Tribe Lemiini
Genus LEIIA Fabricius
(Subgenus LEMA s. str.)
L. cyanella Linn6f
L. gaspensis W.J. Brown PQ
L. palustris Blatchley _ONPQ
L. trilinea trilinea R.E. White MB ON PQ
bilineo.ta Omer-Couper & Miles
medionata Schaeffer
medionota Schaeffer
trilineuta atct.
L. trivittatr,L trivittata Say ON PQ
imma< ulicollis Chevrolat
Iecontti Clatk
nigrou.ittata Schaeffer
triuirgzta LeConte

Genus OUI.,EMA des Gozis

(Subgenus I)ULEMA s. str.)
O. melanopus (Linn6)t ON PQ

Subfamily CLYTRINAE
Key to New World genera: Moldenke (1981)
Key to North American species: Moldenke (1970)
TYibe Clytrini
Genus ANC,TMOEA Lacordaire
Anomoia Dejean
Euthyna Gistel
A. laticlavia laticlavia (Forster) MB ON PQ
ephipptum (Germar)
lloridana Schaeffer

304
 Chrysomeiidae

hansana Schaeffer
nigripes (Fabricius)
o6silo (Fabricius)
ruficauda (Foersberg)

Tribe Megalostomini
Genus COLEOTHORPA Moldenke
C. dominicana dominicana NT_AB_MBON
(Fabricius)
C. vittigera vittigera (LeConte) _BCABSKMB

Tribe Babiini
Genus BABIA Chevrolat
Harpasta Gistel
(Subgenus BABIA s. str.)
B. quadriguttata quadriguttata ON
(Olivier)
biguttata Crotch
quadriforis (Newman)

Genus SAXINIS Lacordaire

(Subgenus BOREOSAXINIS Moldenke)
S. omogera omogera Lacordaire ON
S. saucia saucia LeOonte ec

Subfamily CRYPTOC EPHALINAE

Tlibe Pachybrachini
Genus PACHYBRACHIS Chevrolat
(Subgenus PACH\aBRACHIS s. str.)
(Pachystylus Rey)
P. abdominalis (Say) AB SK MB
P. alticola Fall AB
P. atomarius (Melsheimer) AB-MBON
atomus Bowditch
infaustus Haldeman
P. autolycus FaIl -BCABSKPQ
P. bivittatus (Say) -BCABSK-ONPQ
albescens Suffrian
uiduatus Suffrian
P. calcaratus Fall
P. coloradensis Bowditch ---AB--oN--
seuier Bowditch
P. discoideus Bowditch _AB
P. donneri Crotch BC AB
P. hepaticus (Melsheimer) -ABSKMBONPQ
punctdtus Haldeman
P. luridus (Fabricius) ON PQ
oesculi (Melsheimer)
femoratus (Say)
festiuus Fall
moerens Stil,
nigrinus Blatchley
P. melanostictus Suffrian _YKNTBCABSKMB
P. minor Bowditch
:c - MB- pe -
P. mnigrum (Melsheimer) - - - - - - - -
intricatus Suffrian
P. nigricornis carbonarius AB_MBON
(Haldeman)
P. nigricornis diffrcilis Fall _BCABSKMBONPQ
wahsatchensis Fall
P. nigricornis nigricornis (Say) AB SK MB
janus Fall

305
Chrysomelidae

P. obsoletr.,s Suffrian BC AB _MBONPQ

P. othonous othonous (Say) _MBONPQ
rna rg r. naticoll is Randall
P. othonous pallidipennis Suffrian SK
striatus LeConte
P. othonous sioux Balsbaugh AB SK MB
P. peccans Suffrian AB MB ON PQ NB NS---
P. pectoralis (Melsheimer) _ONPQ NB NS
oculatus Suffrian
sobrin.us Haideman
stricticus Blatchley
P. praeclarus Weise MB ON
Blatchley
elegar,.s
P. relictus Fall -oN
P. spumar:us Suffrian
robori:s Fall
--sx MB ON PQ

P. subfasci atus LeConte MB ON

bigutiatus Suffrian
P. tridens tMelsheimer)
P. trinotatrrs (Melsheimer)
P. vau Fal] -__AB
P. vacillatus Fall BC

Tribe Monachini
Genus LEIIIPHANES Gistel
Mona,:hulus Leng
Mona:hus Chevrolat
Key to l\'orth American species: Balsbaugh (1g66)
L. saponat rs ( Fabricius t MB ON PQ NB
oler (IIaldeman)
uiridanus (Falll

Tlibe Cryptocephalini
Genus CRIIFTOCEPHAIUS O.F. Mtiller
(Subgenus CRYIvIOCEPHAJ-US s. str. )
(C antiiostethus
Haldeman)
Key to North American species: White (1968)
C. calidus Suffrian _MBON
C.
C.
confluer:.tus confluentus Say
gibbicollis gibbicollis Haldeman
BCY:"-o*pe NB NS
C. guttulatus Olivier oN PQ
loufur Newman
C. incertus Olivier
lineoltfius Haldeman
-oN PQ

C. insertus Haldeman AB SK MB ON PQ
ellipscidalis Casey
C. leucomelas leucomelas Suffrian ON
C. mutabil: s Melsheimer _MBON PQ
discoideus Suffrian
dispersus Haldeman
C. notatus notatus Fabricius ON PQ NB NS
C.
C.
notatus quadrimaculatus Say
notatus sellatus Schaeffer
quadruplex Newman
_ _ Bcy SK MB ON PQ
_MB
_MBONPQ
C. NS
q uttulus Suffrian
uadr tg
C. sanguin collis Suffrian BC
nigerrimus Crotch
C. schreibersii Suffrian ON
rugicallis Haldeman
C. spurcus vandykei R.E. White BC
C. striatulrLs LeConte _PQ
C. tinctus LeConte _PQ
C. trivittatrs Olivier _PQ
lirus Newman
uilolus Haldeman
C. venustus cinctipennis Randall SK MB ON

306
 Chrvsomelidae

C. venustus hamatus Melsheimer MB

C. venustus ornatulus Clavareau MB 9"
ornatus Fabricius
C. venustus simplex Haldeman ON
C. venustus venustus Fabricius AB SX nlS ON PQ
flaccidus Suffrian
Genus DIACHUS LeConte
Fabricianus Weise
D. aeruginosus LeConte NT BC MB
pe
D. auratus (Fabricius) -BC MB ON NS
aeneus (OIiier)
chalconotu s (Mannerheim)
urrldis (Melsheimer)
D. catarius (Suffrian) BC
l.BsKMB ON PQ NBI'-
D. pallidicornis (Suffrian) ON

Genus BASSAREUS Haldeman

B. formosus (Melsheimer) -oN PQ NS
egenus Suffrian
guttatus Haldeman
B. lituratus (Fabricius) -oN PQ_
B. Iuteipennis (Melsheimer) -oN
B. mammifer (Newman) -oN
MB
B. pretiosus (Melsheimer)
B. sellatus Suffrian MB ON PQ
(?) notatus (Kirby)
(?) q uadris ig nolus Suffrian
B. sulphuripennis (Melsheimer) ON

Genus TRIACHUS LeConte

T. atomus (Suffrian) -oN Ne-
T. vacuus LeConte -oN
Subfamily CHLAMISINAE
Genus NEOCHLAMISUS Karren
Key to North American species: Karren (1972)
N. alni (W.J. Brown) -oN
MB ON PQ lls
N. bebbianae (W.J. Brown)
N. chamaedaphnes (W.J. Brown) -ONPQ
N. comptoniae (W.J. Brown) NB NS
N. coryli (W.J. Brown) -oN
_ONPQ
N. cribripennis (LeConte) NS
N. eubati (W.J. Brown) _ONPQ
N. fragariae (W.J. Brown) NB
N. tectus (W.J. Brown) -oN
Genus EXEMA Lacordaire
Key to North American species: Karren (1966)
E. canadensis Pierce MB ON PQ NB
E. dispar Lacordaire -oN
pennsyluanica Pierce

Subfamily SYNETINAE
Genus SYNETA Dejean
Key to species: Edwards (1953)
Key to eastern North American species: Brown (1961)
S. albida LeConte BC
suturalis LeConte
S. carinata Mannerheim AK BCT
S. extorris borealis W.J. Brown --oNpeNnNs--
_MBONPQNBNS
NF
NF
S. ferruginea (Germar)
cosfolo (Newman)
rubicunda Lacordaire
tripla (Say)

307
Chrysomel:idae

S. hamata Horn - BC AB SK
S. pilosaW.J. Brown - YK NT BC AB SK MB ON PQ NB NS NF
S. simplex simplex LeConte - BC
S. simplex subalpina Edwards - BC AB

Subfamilv ELIMOLPINAE
Tri6e Nodinini
Genus METACHROMA Chevrolat
Atracli.ela Gistel
Key to syrecies: Blake (1970)
Crotch
M. angustr:lum AB SK MB ON
parall,zlum}J.orn
M. longulun Horn AB SK
no ue m s triat um Schaeffer
opacipenne Fall
Genus PAF:IA LeOonte
Key to North American species: Wilcox (1957); Balsbaugh (1970)
P. aterrima tOliviert PQ
Wilcox
P. fragaria,r fragariae - ON PQ
P. pratensi s Balsbaugh PQ
P. quadrigr.ttata LeConte AB SK MB
salicetLWllcox
P. quadrin< tata (Say) - ON PQ
giluipts Crotch
infusafta (LeConte)
melan.t ra (Melsheimer)
P. sexnoLat r tSayt MB
P. scutellaris (Notmanr - ON
P. thoraci*L (Melsheimert MB ON PQ

Tribe Eumolpini
Genus SPIITITHEROPTryTA Dei ean
Chalc<'paria Crotch
Chrysc,dina BaIy
Key to Nrrth American species: Schultz (1976)
S. globosa (Oliviel - ON

Genus NOL)ONOTA Le{bwe

BrachS,pnoea Gislel
Nodo (lhevrolat
Key to Nrrth American species: Schultz (1980)
N. clypealis Horn AB
N. margaretae Schultz MB ON
N. puncticollis tSayt AB SK MB ON PQ
oenea ('Melsheimer)
humeralis (LeConte)
str ig icollis (Lelbvre )
N. tristis (Crlivier) - ON
ouata (.Say)
pilula (Getrnar)
Genus COI.ASPIS Fabricius
Key to N,rrth American species (part): Blake (1974)
C. brunnea (Fabricius) - ON PQ
flauidtt Say
flauicans LeGvre
C. costipennis CroLch - ON
crotchi Lelevre
C. favosa Say - ON
C. suilla bo: ealis Blake MB ON

Genus RIIABDOPTERUS LeGvre

Rhabdophorus Lelbvre
Key to North American species: Schultz (1977)

308
 Chrysomelidae

R. deceptor Barber AB MB ON
R. praetextus (Say) -oNPQ PE
spiculatus Barber
ueisei (Schaeffer)

Genus CHRYSOCHUS Chevrolat

Atymius Gistel
Key to western North American species: Hatch (1971)
C. auratus (Fabricius) _BCABSK MB ON PQ
C. cobaltinus LeConte _BC
californicus Marshall
castaneus Marshall
tenebricosus Marshall

Tribe Adoxini
Genus GRAPHOPS LeConte
Heteraspis LeConte
Key to species: Blake (1955)
G. cupraeus ( Provancher) PQ
G. curtipennis curtipennis
(Melsheimer)
-oN
G. marcassitus (Crotch) _AB MB ON PQ
nebulosus (LeConte)
G.
G. negrellus Blake . - _ TSK MB
G. nigellus Blake MB
pubescens (Melsheimer) MB ON PQ NS
G.
G.
G.
varians LeConte
viridis Blake SK
yu_ _

Genus XANTHONIA Baly

X. decemnotata (Sayt SKMBONPQ-NS
X. vil losula (Melsheimer) _MBONPQ_NS
plagiata (Melsheimer)
steuensi Blake

Genus FIDIA Walsh

F. viticida Walsh
lurida Lefevre
-oN
Genus BROMIUS Chevrolat
Adoxus Kirby
Eumolpus Redtenbacher
B. obscurus (Linn6)* AK YK NT BC AB SK MB ON PQ NB NS PE NF
cochlearius Say
concinnus Weise
leaisi Weise

Genus MYOCHROUS Erichson

Key to North American species: Blake (1950)
M. denticollis tSayt PQ
M. squamosus LeConte AB SK

Genus GLYPTOSCELIS Chevrolat

Key to species: Blake (1967)
G. albida albida LeConte - BC
G. artemisiae Blake BC
G. longior LeConte
G. pubescens (Fabriciust _-Bc_- MB ON PQ
pini (Say)
hirsuta (Gmelin)
hirta (Olivier)
G. septentrionalis Blake BC

309
Chrysomelirlae

Subfamily CHRYSOMELINAE
Key to North American species: Wilcox (1972)
T?ibe Timarchini
Genus TIMARCHA Latreille
(Subgenus r\MERICANOTIMARCHA Jolivet)
Key to New World species: Jotivet (1948)
T. cerdo StAl _BC
aenea Jtolivet
intento Jolivet
nigra Jolivet
T. intricata Haldeman _BC
adusta Jolivet
uiolacet Jolivet
uiridis Jollet

Tribe Chrysomelini
Genus ZYGDGRAMMA Chewolat
Z. conjuncta conjuncta (Rogers) AB SK MB
Z. continua LeConter AB
Z. disrupta'Rogersr - ON
Z. exclamationis (Fabricius) AB SK MB ON pe
Z. suturalis casta ( Rogers) MB
fesliuo .Fabricius)
Z. suturalis suturalis tFabriciusr - ON pe
pzlcro ,Fabricius)

Genus CALI,IGRAPHA Chevrolat

B oliog r tp ha Motschulsky
M eta llc; g r ap ha Motschul sky
Phyllis Gistel
Polyspita Hope
Key to North American species: Brown (1g45)
(Subgenus C|.RAPHICAI-LO Monros)
C. lunata hlbrida tSayr AB SK MB
lateuittuta (Achard)
C. lunata lunata (Fabricius) - ON
brunnea, (Thunberg)
bowditchi (Achard)
mediori:tpta (Achard)
(Subgenus I IDENSOMELA Monros)
C. bidenticola bidenticola - ON PQ
W.J. Brown
hilaris rAchard)
intermedia (Achard)
similis (Rogers)
(Subgenus COREOPSOMELA Monros)
Linell
C. californica californica - BC
C. californica coreopsivora NT BC AB SK MB ON PQ NB - pE
W.J. Brown
elegans (Olivier)
(Subgenus ACALLIGRAPHA Monros)
C. praecelsis (Rogers) MB
(Subgenus CAILIGRAPHA s. str.)
C. alni Schaeffer - - AB SK MB ON Pe NB NS
confinis (Kirby)
C. alnicola V/.J. Brown - ON PQ NB NS
C. amator S'.J. Brown - ON
C. apicalis Notman - ON PQ
C. confluens Schaeffer - ON PQ NB NS
C. dolosa W.J. Brown - ON PQ

310
 Chrysomelidae

C. igrrota W.J. Brown MB ON PQ NB NS PE

C. incisa (Rogers) MB
C. knabi W.J. Brown MB ON PQ
C.
C.
multipunctata bigsbyana (Kirby)
multipunctata multipunctata
nc
_AB
AB sx y" 9* PQ NE NS

(SaY)
C. ostryae W.J. Brown -oNPQ
C. philadelphica (Linn6) BC_SK MB ON PQ NB NS
decipiens (Weber)
ruficornis (Olivier)
C. pnirsa (StAl) ON PQ
labyrinthica Horn
C. pruni W.J. Brown
C. rhoda Knab
-oN
MB ON
C. rowena Knab MB ON PQ NB
C. scalaris (LeConte) MB ON PQ
multiguttis Stitl
sigmoidea (LeConte)
C.
C. spiraeae {Say)
BC
T -oN
_ON-NB
C. tiliae W.J. Brown
C. verrucosa (Suffriant NT BC AB SK MB ON
C. vicina Schaeffer -oNPQNB
C. virginea W.J. Brown -ONPQNB
Genus LEPTINOTARSA SIAI
Myocoryna Sthl
Polygratnma Chevrolat
Key to North American species: Jacques (1988)
L. decemlineata decemlineata (Say) BC AB SK MB ON PQ NB NS PE
albida Tower
intermedia Towet
melanica Tower
minuta Towet
multitaeniata (StA\:l
oblongata "Iower
pallidaTower
rubicundaTower
rubriuittata 'Iower
tortuosa Tower

Genus I"ABIDOMERA Chevrolat

P arop simena Motschulsky

L. trimaculata (Fabriciust MB ON PQ
cliuicollis (Kirby)
rogersi LeConte

Genus CHRYSOLINA Motschulsky

Ap te r osoma Motschulsky
Chrysomela auct.
OreinaMonros & Bechyn6
Key to species: Brown (I962a)
(Subgenus CHRYSOLINA s. str.)
(S tic hosoma Motschulsky)

C. staphylea tLinn6)t PQ NS NF
(Subgenus SPHAEROMELA Bedel)
C. varians (Schaller)t BC
(Subgenus HYPERICIA Bedel)
C. hyperici (Forster)* BC_ON NB NS
C. quadrigemina (Suffrian)t BC_ON NB NS
gemellata atct.
gemtnata anct.
(Subgenus CHALCOIDEA Motchulsky)
C. frnitima W.J. Brown AK YK
C. fl avomarginata fl avomarginata AB
(Say)
C. flavomarginata vidua (Rogers) BC AB

311
Chrysomelirlae

C. hudsonica W.J. Brown NT MB ON PQ NF

(Subgenus I'LEUROSTICA Motschulskv)
C. cavigera iJ.R. Sahlberg)x AK yK
lolli Jar:obson
(Subgenus I,RCTOLINA Kontkanen)
C. caurina Vy'.J. Brown AK
C. magniceps (J. Sahlberg) AK
C. subsuicat.a (Mannerheim) AK yK
(Subgenus I'EZOCROSITA Jacobson)
C. auripenn:s (Say) AK MB
inornalz (Rogers)
C. basilaris rSay) - YK BC AB

Genus GASI ]ROPHYSA Chevrolat

Key to North American species: Jolivet (1951)
(Subgenus CASTROPIrySA s. str.)
(Gastro'zidea Hope)
G. cyanea caesia Rogers NT BC AB SK
G. cyanea c;anea Melsheimer ON
G. dissimilis (Sayr N'T_ABSK_
G. lormosa t iay r AB SK MB
G. polygoni tLinner+ _BCABSKMB ON PQ NB NS PE
caeruli1tennis (.Say)
r uftcoll, s (Fabricius)

Genus PHAINDON Megerle von Miihlfeld

Key to Nodh American species: Balsbaugh (1983)
(Subgenus PFIAEDON s. str.)
P. armoraciae armoraciae (Linn6)* AK YK NT BC AB SK MB ON PQ NB NS NF
americcnus Schaeffer
P. armoraciae niger Hatch _BC
planus Jatch
P. laevigatus (Duftschmid)* AK NT BC AB SK MB ON PQ NB NS
carri Hr*ch
cochleariae auct. amer.
uniform:is Fall
uancout'erensis Hatch
P. oviformis lleConte)
P. prasinellus (LeConter
AK YK NT BC
_BCAB
AB sK yu oN PQ:tuNS---
punctat.us Hatch
uandyke'iHatch
P. viridis Melsheimer SK MB ON PQ NB
aeruginosus Suffrian
dietrichi Hatch
micr o re ic ula tzs Hatch
i:

oklahortensis }{ahch

Genus HYDIIOTHASSA C.G. Thomson

(Subgenus HYDROTIIASSA s. str.)
(Erernosis des Gozis)
H.
H.
H.
boreela Schaeffer
obliquata (Leconte)
ovalis tBlatchleyr
- YK
v:' AB SK MB
ON
ON
PQ

H. vittata tOlivierJ* NT AB SK MB ON PQ NS
triuittatt (Say)
uaripes r.LeConte)

Genus PRASOCURIS Latreille

Helodes Paykull
P. phellandrii (Linn6)* AK NT BC AB SK MB ON PQ

Genus PLAGIODERA Chevrolat

(Subgenus PI-AGIODERA s. str.)
(Me laso tn ida Schaeffer)
(Pseudoiina Schaeffer)

312
 Chrysomelidae

P. versicolora (Laicharting)t MB ON PQ NB
ar tnor aciae ( Fabricius)

Genus CHRYSOMELA Linne

Key to New World species: Brown (1956)
(Subgenus CHRYSOMELA s. str.)
(Lina Dejean)
(Melasoma Stephens)
C. aeneicollis (Schaeffer) BC AB
quadriguttoides Beller & Hatch
C. blaisdelli (Van Dyke) AK NT BC
C. confluens (Rogers) _BC
maculicollis Schaeffer
C. crotchi W.J. Brown AK NT_ABSKMB ON PQ NB NS
C. engelhardti (Hatch) AK YK NT
_LB
C.
C.
falsa W.J. Brown
invicta W.J. Brown
AK YK NT BC AB SK
AB
MB ON PQ v
C. knabi hesperia W.J. Brown AB SK MB
C. knabi knabi W.J. Brown oN--
C. laurentia W.J. Brown NT_AB oN PQ
C. lineatopunctata (Forster) NT BC AB SK MB oN PQ
nouaboracensis Gmelin
obsoleta (.Say)
s cr ip toide s ( Schaeffer)
uariegata Olivier
C. mainensis interna W.J. Brown BC AB
C. mainensis littorea W.J. Brown AK BC
YK _AB MB NB NS _LBNF
C.
C.
mainensis mainensis Bechyn6
schaefferi W.J. Brown AK YK
NT
BC AB MB 9* PQ

immaculata (Schaeffer)
C. scripta Fabricius BC AB SK MB ON
C. semota W.J. Brown YK BC AB
C. walshi W.J. Brown )' ON PQ

Genus GONIOCTENA Chewolat

Key to North American species: Brown (1942o)
(Subgenus C.ONIOCTENA s. str.)
(Phytodecta Kirby)
G. americana (Schaeffer) AK YK NT BC AB SK MB ON PQ NB LB
arctica Bechyn6
guttifera Bechyn6
G. nivosa alberta W.J. Brown -NT_AB
G. nivosa arctica (Mannerheim) AK YK NT BC AB MB
G. nivosa nivosa (Suffrian)
G. notmani (Schaeffer) YKNT_ABSK MB ON PQ LB
G. occidentalis (W.J. Brown) YK-BCAB
Genus PHRATORA Chevrolat
Key to North American species: Brown (1951)
(Subgenus PHRATORA s. str.)
(Phyllodecta l\rby)
P. americana canadensis W.J. Brown ON PQ NB NS
P. frosti frosti W.J. Brown NS__
P. frosti remissa W.J. Brown AK EC AB SK MB ON PQ
P. hudsonia W.J. Brown AK YK NT BC AB sK yu oN PQ
P. kenaiensis W.J. Brown AK
P. interstitialis Mannerheim AK YK NT BC
aklauiki (Can)
P. purpurea novaeterrae W.J. Brown NF
P. purpurea purpurea W.J. Brown YK NT BC AB Sf ff,fS ON Pe NN NS

Genus ENTOMOSCELIS Chewolat

C hry sornelop silo Strand
Chrysomelopsis Achard
Key to North American species: Brown (1942b)
E. americana W.J. Brown AK YK NT BC AB SK MB

313
Chrysomelic;.ae

Subfamily GALERUCINAE
Tribe Galerucini
Key to North American species: Wilcox (1965)
Genus DEROSPIDEA Blake
Key to North American species: Blake (1931)
D. brevicollis (LeContet ON

Genus TRIRHABDA LeConte

Key to North American species: Blake (1931)
T. adela Bla-<e MB ON
T. attenuata (Sayt - BC AB
T. bacharidi; (Weber) pe
tomentt.'sa LeConte
T. borealis Elake - BC AB SK MB ON PQ NB NS
indigoptera Blake
neoscoti.ae Blake
T. canadensis (Kirby) - BC AB SK MB ON PQ NB NS
tomentc'sa Crotch
T. converger:.s LeOonte AB SK MB ON
uirescen,s Cockerell
uiridicT'anea Blake
T. lewisii Crrtch - BC
T. pilosa pilr'sa Blake - BC
T. sericotrachyla Blake - BC
T. virgata LeConte _ONPQ NB NS

Genus GAI-IDRUCELLA Crotch

(Subgenus C:ALERUCELLA s. str.)
(Hydrollaleruco Laboissidre)
G. nymphaerre (Linn6)x AK YK NT BC AB SK MB ON PQ NB NS PE LB NF
fernoralis (Meisheimer)
Iuctuosu (Mannerheim)
margin,zlla (Kirby)
sagittariae (Kirby)
stilleri ?app
(Subgenus I\'EOGAIERUCELLA Chujo)
G. quebecen;is W.J. Brown PQ
G. stefansso:ri W.J. Brown AK YK NT PQ_NS
Genus TRICHOLOCHMAIA Laboissibre
T. alni (Fall. -oN PQ NB NS
T. decora calbo (LeConte) - BC AB SK MB
T. decora de,:ora (Say)
solicis (Randall)
-oN PQ NB

T. cavicollis il,eConte) AB SK MB ON PQ
haematlca (LeConte)
T. kalmiae (Fallt -oN PQ NB NS
T. perplexa tFallt
T. punctipennis pallida
(Beller & Hatch)
- BC _9* PQ NB

T. punctipennis punctipenms - BC
(Mannerheim)
T. ribicola cc'nfusa (W.J. Brown) ON
T. ribicola ribicola (W.J. Brown) AB SK MB
T. rufosangu.inea (Sayt _PQ
T. sablensis (W.J. Brown) NS-
T. spiraeae t Fall.) oll pe -
T. spiraeophila (Hatch & Beller) - BC
T. tuberculaba (Say) ON PQ NB ols -
T. vaccinii (Fallt ON PQ NB

Genus PYRRHALTA Joanni s

P. viburni (Paykullt* ON PQ

Genus XANI|HOGALERUCA Laboissibre

X. luteola (O.F. Miiller)t - BC oN PQ

3L4
 Chrysomelidae

Genus ERYNEPHAI-A Blake

Key to species: Blake (1936)
E. rnaritima (LeConte) NS
texana Schaeffer
E. puncticollis (Say) AB SK MB

Genus MONOXIA LeConte

Key to species: Blake (1939)
M. angularis (LeConte)
M. consputa LeConte
M. debilis LeConte
M. grisea Blake
M. inornata Blake
M. puberula Blake
M. sordida LeConte

Genus OPHRAELLA Wilct.rx

Key to North American species: LeSage (1986)
americana (Fabricius) SK MB
O.
O. arctica LeSage NT
AB
MB 9* PQ

O. bilineata (Kirby) -BC

_BC
AB
AB
SK
SK
MB
ON
O. communa LeSage
notulata anct.
O. conferta (LeConte) AB SK MB oN PQ NB NS
O. cribrata (LeConte) AB SK yu ON PQ
O. notata (Fabricius) oN PQ
O. nuda LeSage AB
O. pilosa LeSage _BC AB SK MB ON PQ

Genus GA.LERUCA O.F. Miiller

Key to North American species: Blake (1945)
(Subgenus GALERUCA s. str.)
(Adimonia Laicharting)
G. browni Blake
G. externa (Say)
G. rudis (LeConte)

Tribe Luperini
Genus DIABROTICA Chewolat
D. barberi Smith & Lawrence MB ON PQ NB
longicornis atct.
D. cristata (Harris) - SK MB ON
atripennis (Say)
fossata LeConte
D. undecimpunctata howardi Barber oN PQ
d uodec im p unclofa (Fabricius)
D. virgifera virgifera LeConte ON
filicornis Hotn
Genus ACAI-YMMA Barber
Key to Norbh American species: Munroe and Smith (1980)
A. gouldi Barber _ONPQ
A. vittatum (Fabricius) MB ON PQ NE NS
atnericanutn (Gmelin)
melanocep halurz ( Fabrici us)
pallipes (OIivier)
stolatum (Gmelin)

Genus CEROTOMA Chevrolat

Andrector Horn
Cerotana Bowditch
Ceratorna Doziet
C. trifurcata (Forster) MB ON PQ

315
ChrysomeliCae

Genus PSELIDOLUPERUS Beller & Hatch

P. lecontii (tlrotch) _BC
P. longulus (LeConte) -BC
baheri Beller & Hatch

Genus SCELOLYPERUS Crotch

S. cyanellusr (LeConte) ON NS
S. nigrovirescens (FalI) -BCAB
S. schwarzi Horn _BC
S. torquatus (LeConte) _BC
S. varipes (i-eConte) _BCAB
concauus Beller & Hatch

Genus PHI'LLOBROTICA Chevrolat

P. decorata (Say) BC AB SK MB PQ NS
P. discoidea (Fabricius) -oN
circumdata (Say)
P. limbata (Fabricius) -oN PQ

Subfamilv ALTICINAE
Genus BLEPHARIDA Chevrolat
Key to North American species: Mignot (1971); Balsbaugh and Hays (1972)
(Subgenus ULEPHARIDA s. srr.t
(BlephuridulaWeise)
B. rhois (Forster) AB SK MB ON
atripennis }Iotn
dorothea Mignot
meticu'.osa (Olivier)
roi (Ol vier)
s/olido (Fabricius)

Genus PHYLLOTRETA Chewolat

Tanygctster Blatchley
Key to North American species (part): Smith (1985)
P. alberta Chittenden AB
P. albionica (LeconteJ BC AB SK
P. aerea Allardt ON
P. armoraciae (Koch)t _SK ms oN PQ NB PE
biplagiata Chittenden
P. bipustule.ta t Fabricius t oN PQ
P. brevipen'ris Chittenden MB
P. chalybeil'ennis tCrotch t
-NS
P. conjunctrr Gentner AB SX MB ON PQ
P. constrict r E.H. Smith nc AB SK
P. crucifera : I Goeze tt AB SK MB oN pe NE NS
columl: iana Chittenden
P. decipiens Horn BC AB
ordina;ia Chittenden
P. denticon is Horn BC
a m pl ic cr ni s Chittenden
aequal.is Hatch
P. inconspicua Chittenden BC-SKMB
P. lewisii (Crotcht BC
P. liebecki Schaeffer _MBON
P. oblonga Chil,tenden _ABSKMB
P. pusilla Horn -ABSKMBON
P. ramosoides E.H. Smith NT BC
P. robusta l,eConte _ABSKMBONPQ
P. striolata (Fabricius)f - YK NT BC AB SK MB ON PQ NB NS
artiu ittata Chittenden
lineolata Chittenden
uerniccsa Chittenden
uillolo (Fabricius)
P. utanula li.H. Smith BC
P. zimmerrnanni (Crotch)* AK - NT BC AB SK MB ON PQ NB NS PE NF
sinuatu Horn

316
 Chrysomelidae

Genus APHTHONA Chevroiat

Cerataltica Ctotch
A. cyparissiae (Koch)t AB?_ON-?
A. flava Guillebeaut AEr I ',' _ ')
A. nigriscutis Foudrast AB_MBON
Genus LONGITARSUS Berthold
(Subgenus LONGITARSUS s. str.)
(Apterius Blatchley)
(T einodacty la Chevrolat)
(Thyamis Stephens)
L. erro Horn AB-MBONPQ NS
L. fermgineus (Foudras)t YBC -oNPQ PE NF
L. ganglbaueri Heikertingert MB
L. insolens Horn AB ON PQ
L. jacobaeae Waterhouset -nc
L. luridus (Scopoli)f
L. melanurus (Melsheimer)
-oNPQ
MB ON PQ
NU NS
v
L. pallescens Blatchley -oN
_ONPQ
L. pellucidus (Foudras)t
L. pratensis (Panzer)t _ONPQ PE
L. rubiginosus (Foudras)f _ONPQ
L. succineus Foudrasf NT
L. suspectus Blatchley -oN
_ONPQ
L. testaceus (Melsheimer) NB

Genus GLYFTINA LeConte

Balophila Ctotch
G. abbreviata Gentner ON
G. atriventris Horn BC
T sx lle
_MB ON
G. brunnea Horn ON PQ
G. cyanipennis (Crotch) ON
G. spuria LeConte _MB ON PQ
lissotorques LeConte

Genus SYSTENA Chevrolat

S. bitaeniata (LeConte) AB
S. blanda (Melsheimer) AB_MBONPQ
ligata (.LeConte)
ochracea (LeOonte)
taeniata (.Say)
S. carri Blake _AB
S. corni Schaeffer oN-
S. dimorpha Blake
S. elongata (Fabricius)
_AB s^-
_MB
floridana Blatchley
subdenea (LeConte)
S. frontalis (Fabricius) _MBONPQ
S. hudsonias (Forster) -MBONPQ
S. marginalis (Illiger) ON PQ
oblonga (LeConte)

Genus STRABAJ-A Chevrolat

(Subgenus STRABAI-A s. str.)
Key to New World species: Blake (1953)
S. rufa (Illiger) oN PQ

Genus AJ-TICA O.F. Miiller

(Subgenus ALTICA s. str.)
(.Gr a p toder a Chevrolat)
(Megaltica Hatc}l':'
A. aeneola Blatchley
aenescens Blatchley
-oN PQ

A. ambiens LeConte NT BC MB ON PQ NS
alni Hatris
latiplicata Blake
A. betulae Schaeffer PQ

317
ChrysomeLidae

A. bimargirrata Say AK YK NT BC AB SK MB ON PQ LB
p licipe nnis ( Mannerheim)
A. browni }4ohamedsaid MB ON PQ NB NS
potentillae W.J. Brown
A. canadensis Gentner AB SK MB
A. chalybar,ra Illiger
uitiuota Thomas
-oN PQ

A. carinata Germar MB ON PQ NB
e*apta Say
A. caurina Blake BC AB _MB
A. corni Woods
A. elongatula Csiki
BC
_AB -MBONPQ
_MBON
elongata Gentner
A. foliaceae LeConte MB ON
puncti.oennis LeOonte
A. heucherae FaiI MB
A. ignita Illiger
A. inaerata LeConte
-oN
NT MB
A. kalmiae Melsheimer PQ NB NS
uirida:za Schaeffer
A. litigah lrall -oNoN PQ
A. populi S'.J. Brown
A. prasina LeOonte BC AB
SK
SK
yu NB

A. probata Fall
A. ribis W.,1. Brown :-BC__ -oN
A. rosae Woods MB ON PQ
A. subplica;a LeConte _ONPQ
A. sylvia Malloch
cuprascens FalI
-oNPQ NU NS

torquo.ta LeConte
A. tincta LeConte -BC
A. tombacina (Mannerheim) AK YK NT BC AB SK l,fB ON Pe NU WS LB
euicta LeConte
shoemtkeri Schaeffer
A. ulmi Wocds MB ON PQ
A. vicaria Ilorn AB
A. woodsi Iiely -oN
Genus HOIINAJ-TICA Barber
H. atriventris (Meisheimer) ON PQ

Genus ORT'HALTICA Crotch

Leptot,brix Heikertinger & Csiki
Le ptot. icha I ti co Heikertinger
Leptot,ix Horn
M ic re p itr ix Laboissidre
Serraticollis R.E. White
Key to North American species: Scherer (1974)
O. copalina (Fabricius) oN PQ
fortico.nis (Illiger)
Genus DEFiOCREPIS Weise
D. erythroJ'us (Melsheimer) PQ
rufipes (Ctotch)
D. oregoni .{atch BC

Genus ORIISTIA Germar

Orestirtides Hatch
PyladiaWeise
O. robusta LeContet BC SK MB ON PQ NF

Genus HIPPURIPHILA Foudras

Key to North American species: Brown (19426)
H. canadensis W.J. Brown
H. equiseti Beller & Hatch
AK -NT_ABSK
BC AB
y" oN PQ
v
H. manculzL (LeConte) AK BC AB SK
modeeri anct.

318
 Chrysomelidae

Genus CREPIDODERA Chevrolat

Key to North American species: Parry (1986)
C. browni Parry -oN
C. decora Parry
_ECAB s^
-oNPQ
MB ON PQ r.rs lls
C. digra Parry AK
C. heikertingeri (Lazorko) _BCAB SK MB ON PQ NB NS NF
C. luminosa Parry _ONPQ NB NS T\TF

C. nana (Say) NT BC AB SK MB ON PQ NB
c hittende ni Heikertinger

C. populivora Parry NT BC AB SK MB ON PQ NB NS
C. sculpturata (Lazorko) _BC MB ON
C. solita Parry _BCAB SK MB ON PQ
C. spenceri (Lazorko)
C. vaga Parry
_ _ _:'_ -oN
_ONPQ
C. violacea Melsheimer

Genus EPITRIX Foudras

E. brevis Schwarz -oN
_SKMBONPQ NS
E. cucumeris (Harris) NS
(?) pubescens llliget
seminulum LeConte
E. hirtipennis (Melsheimer)
E. humeralis Dury
E. subcrinita (LeConte)
E. tuberis Gentner

Genus MANTURA Stephens

M. chrysanthemi (Koch)f
M. floridana Crotch AB SK lrs on peNeNs--Nil
Genus CHAXTOCNEMA Stephens
(Subgenus CHAETOCNEMA s. str.)
C. concinna (Marsham)'l _PE
C. confrnis Crotch AB SK lrs om pe Ne NS
flauicornis LeConte
C. cribrata LeConte NT BC AB SK MB ON PQ NB NS NF
rudis LeConte
C. cribrifrons LeConte _ONPQ
C. decipiens LeConte _SK
C. denticulata (Illiger) -oN
americana Motschulsky
s emichalchea Melshermer
texana Ctotc}r
C. ectypa Horn _SKMB
C. minuta Melsheimer MB ON PQ NB NS
C. opulenta Horn AB SK MB
C. perturbata Horn _SKMB
C. protensa LeConte
C. pulicaria Melsheimer v -nc
_BC
AB SK MB ON
AB ON
PQ

(?) aeneola LeConte

C. subcylindrica LeConte NT BC AB SK MB ON PQ
cylindrica LeConte
C. subviridis LeConte AB SK MB

Genus DISONYCHA Chevrolat

Key to North American species: Blake (1933)
D. alternata (Illiger) BC AB SK MB ON PQ NB NS
punctigera Schaeffer
(? ) q uinq ueu ittaro ( Say)
D. arizonae Casey MB ON
borealis Blake
douisi Schaeffer
glabrata Jacoby
D. latifrons Schaeffer BC AB SK MB ON PQ NB NS
osferis Schaeffer
laticollis Schaeffer
D. latiovittata Hatch _BC
D. pensylvanica (Illiger) ON PQ

319
Chrysomel,dae

parua Blatch\ey
pensyiuania Stwn
(?) s e x,t ineata (Olivier)
D. procera Casey NT BC A3 SK MB ON PQ
n igriutntris Schaeffer
(?) pallipes Crctch
(?) uici.na l\trby
D. punctigera LeConte NT AB SK MB
'neglecta Schaeffer
punctipennis Schaeffer
D. schaefferi Blake
D. triangularis (Sayr
montanensis Blake
- - scABSf MB ON PQ

puncticollis (Kirby)
D. uniguttr.ta { Say t MB ON PQ
uicina \{ttby
D. xanthonrelas ( Dalman) NT AB SK MB ON PQ PE
atrella Blake
ceruicctlis LeConte
collaris (Illiger)
merdiuora Melsheimer
xanthomelaeno Gemminger & Harold

Genus KUSICHELINA Bechyn6

Key to North American species: Blake (1927)
K. barberi (Blake) _AB
K. frmbriat.r (Forster) - -4,8
circun cincta (Crotch)
suture'Ia (Say)
K. gibbitanra (Say) ON
K. lugens (li,eConte) NT BC AB SK MB ON
lamprt)cydnea (B1ake)
K. perplexzi (Blake) ON
K. thoracica. (Fabricius) ON
flaua (Gmelin)
K. vians (Il iger) NT BC SK MB ON PQ NS
abdon',.inalis ( Olivier)
badia Blake)
scripti.:ollis (Say)

Genus CAPRAITA Bechyn6

ChloephagaWeise
Key to Nrrrth American species: Blake (1927)
C. circumditta (Randall) ON
limbalis (Melsheimer)
(?) que"cata (Crotch)
C. scalaris t,Melsheimer) oN PQ
lobata (LeConte)
C. sexmacuLata (Illiger) ON
cretica (Jacoby)
palliatt (RandalI)
C. subvittata (Horn) AB SK MB oN PQ NB NS
C. thyamoides (Crotch) ON

Genus DISTIGMOPTERA Blake

Key to North American species: Blake (1943)
D. apicalis l3lake
D. borealis Blake BC AB SK
-oN
MB ON PQ NB
D. pilosa (Illiger) -oNPQ
Genus DIBOLIA Latreille
Key to N'rrth American species: Parry (1974)
(Subgenus IDIBOLIA s. str.)
D. borealis tlhevrolat AB SK MB _PQNBNS
D. chelones Parr5r oNPQ-NS
D. Iibonoti l{orn sc
D. melampyri Parry
D. penstemrrnis Parry
T ONPQNBNS___
D. reyheria Mignot AB _MB 9*PQIUNS_-_

320
 Chrysomelidae

Genus PSYLLIODES Berthold

P. affinis (Paykull)t -oN
P. chrysocephala (Linn6)t --Ne
P. convexior LeConte -oN
P. cucullata (Illiger)f PQ NB
elegans Horn AB
P.
P. napi (Fabricius)f _ONPQ NB pn-*"
P. picina (Marsham)-l _ONPQ
P. punctulata Melsheimer AB MB ON PQ _NS I\TE]

aenescens Casey
extricata Casey
paruicollis LeConte

Subfamily HISPINAE
Tribe Chalepini
Key to North American genera: Butte (1968o)
Genus ANISOSTENAWeise
(Subgenus ANISOSTENA s. str.)
A. californica Van Dyke _BC
A. nigrita tOlivier) -BCMBONPQ NB

Genus SUMITROSIS Butte

Anoplitis Chapuis
Key to North American species: Butte (1969)
S. ancoroides (Schaeffer) AB SK MB
S. inaequalis (Weber) - BC AB SK MB oN pe N" NS
bazcis (Newman)
flauipes (Germar)
neruosa (Hotn')
obsoleta (Say)
pallida (Say)
suturalis (Fabricius)
S. rosea (Weber) SK MB ON PQ NB NS PE
philemon (Newman)

Genus ODONTOTA Chevrolat

Key to North American species: Butte (1968c)
O. dorsalis tThunberg) oN PQ
hanisi Crotch
scutellaris (Olivier)
suturalis (Harris)
O. mundulus (Sanderson) MB
O. scapularis t Oliviert
lateralis (Say)
-oN
Genus CHALEPUS Thunberg
Anoplitis Kirby
Key to North American species: Butte (1968b)
C. bicolor (Olivier) ON
rufiuentris (Suffrian)
rufogaster Thunberg

Genus BALIOSUS Weise

B. nervosus (Panzer) ON PQ NB NS
marginatus (Say)
marmoratus (Baly)
obsoletus (Say)
pallipes (Germar)
quadratus (Fabricius)
quercifoliae (Harris)
rubellus (Schonherr)
ruber (Weber)

32L
ChrysomelLdae

Tlibe Uroplatini
Genus STEiNOPODIUS Horn
Key to North American species: Blaisdell (19386)
S. lateralis lSchaeffer) AB SK
uand.uzeei Blaisdell
S. texanus (Schaeffer.t AB SK MB

Genus BRACHYCORYNA Gu6rin-M6neville

Key to North American species: Staines (1986)
B. hardyi ((lrotcht - BC AB
confusa Hatch
horni'Weise
melsheimeri Horn
B. montana (Horn) AB SK MB ON

Genus GL)?HUROPLATA llhmann

Key to North American species: Riley (1985b)
G. pluto (Newman) AB - MB ON PQ
porcatd (Melsheimer)

Genus MIC.IRORHOPALA Chevrolat

Key to North American species: Clark (1983)
M. excavat.r cyanea { Say } - BC AB SK MB
M. excavatrr excavata (Olivier) MB ON PQ NB NS
M. vittata tFabricius) - BC AB SK MB ON PQ NB
laetuluLeConte
M. xerene (Newman) AB SK MB ON PQ NB
interrt.tpta Couper

Subfamily CASSIDINAE
Tribe Stolaini
Key to genera: Hincks (1952)
Genus CHINLYMORPHA Chevrolat
C. cassidea (Fabricius) _SKMBONPQ
orgus iLichtenstein)
cribraria (Olivier)
geniculata Boheman
lewisi Crotch
s ep t e rrtde ci m p u nc t at a (S ay)

Tribe Physonotini
Genus PHI'SONOTA Boheman
Key to North American species: Sanderson (1948)
(Subgenus PIfYSONOTA s. str.)
P. helianthi tRandallt MB ON PQ
q uinquepu nct ato (Walsh)
sep ten,tr io nalis Boheman
P. unipunctata (Say) - SK MB ON PQ

T?ibe Cassidini
Key to North American genera: Riley (1986a)
Genus CASSIDA Linn6
Key to North American species: Riley (1986b)
(Subgenus CASSIDA s. str.)
C. azurea F'abriciusf - SK
C. flaveola Thunbergt - YK NT BC AB SK MB ON PQ
C. rubiginosa O.F. Miiller+ AB SK MB ON PQ NB

Genus JON'THONOTA Spaeth

J. nigripes .Olivier) AB SK MB
ellipsis (LeConte)

322
 Nemonychidae

Genus AGROICONOTA Soaeth

A. bivittata (Say) MB ON PQ
striolata (Boheman)
uittula (Boheman\

Genus PLAGIOMETRIONA Spaeth

(Subgenus PARAMETRIONA Spaeth)
P. clavata (Fabricius) ON PQ NB
diu er s icoll is ( Schaeffer)

Genus STRONGYLOCASSIS Hincks

Strongylaspis Spaeth
Key to species: Riley (1985o)
S. atripes (LeConte) MB ON
creolo (Sanderson & King)
loedingi (Schaeffer)

Genus DELOYALA Chewolat

Chirida Chapuis
D. guttata (Olivier) SK MB ON PQ NB NS PE
bohemani (Spaeth)
columbica (Spaeth)
cruciata (Fabricius)
fuliginosa (OIivier)
hamulata (Boheman)
immunita (Boheman)
immunda (Boheman)
lucidula (Boheman)
lugubrina (Boheman)
p ennsy lu anic a ( Spaeth)
signifera (Herbst)
trabeata (Boheman)

Genus CIIARIDOTELLA Weise

(Subgenus CHAEROCASSIS Spaeth)
C. purpurata (Boheman) SK MB ON PQ NS
C. sexpunctata bicolor (Fabricius) -oN PQ NB NS
auric halcea (Fabricius)
auris plendens (Mannerheim)
b istrip ustulata (Herbst)

floridana (Schaeffer)
(? ) mary landico ( Herbst)
pallida (Herbst)

Superfamily Curculionoidea
Family NEMOI{YCHIDAE
J. McNamara

Nemonychidae comprises eight species in Anthribidae. Kuschel (1989) recently published a

Canada. The adults feed on the staminate flowers revision of this family for North America The list
ofvarious conifers. below follows O'Brien and Wibmer (1982).
The nemonychids are often included in
Curculionidae, but their larvae show an aflinity to BC (6); AB (3);MB (1); ON (2);PQ (3); NB (1)

323
Anthribidae

Subfamily DOYDIRFIYNCHINAE
Tribe Cimberidini
Genus CIMBERIS des Gozis
C. bihirsuta Hatch BC
C. compta (LeConte) BC
paruulus lJatch
C. decipiens Kuschel BC
C. eiongata (LeConte) BC MB ON PQ
C. pallipennis (Blatchley) AB PQ
C. pilosa (LeConte) -oN PQ NB
Genus PITYOMACER Kuschel
P. pix Kuschel BC AB

Genus ACFI OMACER Kuschel

A. bombifrr: ns (LeConte) BC AB

Family ANTHRIBIDAE fungus weevils

J. McNamara

Anthribidae includes 18 species in Canada and lar habitats. Adults feed on fungr, pollen, or even
Alaska. One of these, the coffee bean weevil bark; larvae feed in plant stems, fungr, orvvoods.
(Araecerus fasciculatus), is introduced and may be Valentine (1960) has revised the North
in warehouses; this species damages
establishred American genera of Anthribidae.
many stored plant materials, including seeds.
Adults of anthribids are often collected by beating AJ{ (2); YK (2); NT (2); BC @); AB (8); S.K (6); MB
dead or diseased vegetation or by sweeping in (7); ON (12); PQ (12); NB (1);NS (1);NF (1) o fl
weedy areas. The larvae are usually found in simi- (1); I (1)

Subfamily CHORAGINAE
T?ibe Araecerini
Genus ARAECERUS Schrinher"r
A. fasciculatus (DeGeer)t BC ON PQ

Tlibe Choragini
Genus EIIXENUS LeConte
E. punctatus LeConte PQ

Subfamily ANTHRIBINAE
TYibe Eupariini
Genus EUPARIUS Schdnherr
Key to Norlh American species: Valentine (1960)
E. marmoreus (Olivier) MB ON PQ NB
albifrons (Boheman)
lunatus (Fabricius)

Tlibe Anthribini
Genus TRICIONORHINUS Wollaston
Anthrihulus LeConte
Brachy tarsoides Pierce
Key to wrstern North American species: Vaientine (1971)
T. alternatus (Say) AB MB ON
T. annulatus (Carr) _BC AB

324
 Ithyceridae

T. griseus (LeConte)
T. limbatus (Say)
T. sticticus (Boheman)
- - -NT -BC ffi3f reoNpe - - - -
AB SK MB ON PQ
-
obsoletus (Fahreus)
uariegatus (Say)
T. tomentosus tomentosus (Say) ON
6reuis (Fahreus)
paululus (Casey)

Tribe Ormiscini
Genus ORMISCUS Waterhouse
Eusphyrus LeOonte
Gonops LeConte
Toxotropis LeConte
O. saitator LeConte oN PQ
angulatus Pietce
O. walshii (LeConte) ON

Tribe Tropiderini
Genus TROPIDERES Schbnherr
Eurymycter LeConte
Gonotropis LeConte
'Key to North American species: Valentine (1960)
T. dorsalis (Thunberg)* AK YK NT AB SK MB PQ NF
gibbosus (LeConte)
T. fasciatus (Olivier) _BC SK MB ON PQ
bicarinatus (Pierce)
T. latifascia (Pierce)
T. tricarinatus (Pierce) _ _ _ oNiS - NS

Tribe Allandrini
Genus AI,LANDRUS LeConte
Key to North American species: Valentine (1960)
A. bifasciatus LeConte
A. brevicornis Frost ----AB-nreoNPQ-----
AB SK _ ON
A. populi Pierce AK \'K PQ

Family ITIIYCERIDAE New York weevil

J. McNamara

Ithyceridae includes only one species. The Sanborne (1981) studied the morphology of both
adults feed on the tender shoots and buds ofvarious larvae and adults, as well as the biology of the
hardwoods and occasionally on some cultivated fruit species.
trees. The larvae feed on the roots of the host
plants. ON (1); PQ (1)

Genus ITHYCERUS Schonherr

Pochyrhynchus Kirby
L noveboracensis (Forster) oN PQ
cur culionoides (Herbst)
p unc ta t uI us ( Fabricius)
schoenherri (Kirby)

325
Rhynchitidrre

FamilyATTELABIDAE leaf-rollingweevils
J. McNamara

Four species of Attelabidae occur in Canada. The classification used here follows that of
The females form rolls from leaves and lay a single Robert W. Hamilton (unpublished Ph.D. thesis,
egg in each roll. The larvae feed on the inner part 1969), who revised the North American species.
of the roll. SK (1); MB (2); ON (3); PQ (a); NB (1); NS (2)

T?ibe Attelabini
Genus HIMATOLABUS Jekel
H. pubescerrs (Say) MB ON PQ NB NS
colora<:loensis Yoss
d,is p ar.'.p e s (Chittenden)
maculutus (Provancher)
rhois (:3oheman)

Genus HOI\{OEOLABUS Jekel

H. analis (Illiger) SK MB ON PQ
similis (Kirby)

Genus ATTELABUS Linn6

(Subgenus IiYNOLABUS Jekel)
A. bipustulatus Fabricius -ONPQ_NS
A. nigripes,J.E. LeConte PQ
genalis LeConte

Family RHYNCHITIDAE leaf and bud weevils

J. McNqmara

Rhynchitidae comprises 13 species and sub- family, but only parts of his work have been pub-
species in Canada. Some lay their eggs in the lished to date; he also wrote a key to the North
young fruit or buds of trees that are then destroyed American genera ( 1971).
by the larvae; others feed on blossoms or foliage.
Robert W. Hamilton (unpublished Ph.D. thesis, NT (3); BC (5); AB (6); SK (5); MB (6); ON (7); PQ
1969) revised the North American species of the (5);NB (3);NS (3); PE (2);NF (2)

Subfamily PTEROCOLINAE
Genus PTEROCOLUS Say
ApotonLus Kirby
P. ovatus (Fabricius) ON

Subfamily RIIYNCHITINAE
Tribe Rhynchitini
Genus EUGNAMPTUS Schttnherr
Key to Norih American species: Hamilton (1990)
E. angustatus (Herbst) ON
abdominalis Yoss
collaris (Fabricius)
fuscipes Pierce
Nigripennis ( Fabricius)
nigripes Pierce
rubricollis (Say)
ruficeps Pierce

326
 Brentidae

ruficollis (Germar)
sulcifrons Gyllenhal

Genus PSELAPHORHYNCHITES Schilsky

Key to North American species: Hamilton (1971)
p. cyanellus (LeConte) NT - AB sK MB ON PQ NB NS PE - NF
P. perplexus (Blatchley) - ON PQ NB NS PE
Genus MERHYNCHITES Sharp
Key to North American species: Hamilton (1985)
M. bicolor bicolor (Fabricius) NT BC AB SK MB oN PQ
antennalis (Yoss)
intermedius (Yoss)
M. bicolor cerdonis (Voss) - BC AB
M. bicolor cockerelli (Pierce) AB
erythrosoma (Cockerell & Harris)
M. bicolor nigricephalus Hamilton - BC
M. wickhami wickhami (Cockerell) NT BC AB SK MB
homocidae (Yoss)
mandibularis (Yoss

Genus HAPLORHYNCHITES Voss

Key to North American species: Hamilton (1974)
H. aeneus (Boheman) - SK MB
consobrinus (Yoss)

TYibe Auletini
Genus AULETOBIUS Desbrochers
Key to North American subgenera: Hamilton (1983)
(Subgenus ALETINUS Desbrochers)
A. ater (Leconte) MB oN PQ
(Subgenus MESAULETES Voss)
A. cassandrae (LeConte) - ON PQ NB NS NF
albou stitus (Blatchley)
e

blatchleyiYoss
minor (Blatchley)
(Subgenus AULETOBIUS s. str.)
Metopon Waterhouse
A. congruus (Walker) - BC AB SK MB
subcoeruleus LeOonte

Family BRENTIDAE brentid beetles

J. McNamara

Only one species of Brentidae, the oak timber- into the heartwood, often causing serious injury to
worm, occurs in Canada. The adults are found oak stock.
under bark or in decayed wood. The larvae bore ON (1); PQ (1)

T?ibe Arrhenodini
Genus ARRHENODES Schdnherr
A. minutus (Drury) - ON PQ
brunneus (Panzer)
lecontei (Power)
maxillosus (Olivier)
p aup er c ula (Blatchley)
sallei (Power)
s epte ntrionis (Herbst)

327
Apionidae

Family APIONIDAE pear-shaped weevils

J. McNamara

Thirty-five species of Apionidae are recorded in

Canada and Alaska. The larvae feed on leaves and
stems, carrsing galls, or develop within the seeds of
various plants. The adults usually feed on leaves.
Than}s to the work of Kissinger (1968), the tax- aK (3); vK (3); NT (3); BC (12); AB (20); SK (13);
onomy of t;he North and Central American species is MB (15); ON (17); PQ (13); NB (2); NS (2); NF (1)
well known. . I(1)
Tlibe Nanophyini
Genus NAN{)PHYES Schonherr
Nanoda cty lus Blatchley
Nanodes Schtinherr
P se udo ty c hius Blatchley
(Subgenus NANOPHYES s. str.)
N. canadens s W.J. Brown AB SK MB

T?ibe Apionini
Genus APION Herbst
(Subgenus ALOCENTRON Schilsky)
A. attenuatum Smith BC A,E MB ON PQ
(Subgenus C ERATAPION Schilsky)
A. contusum Smith PQ
notabile, Buchanan
spinipes Fall
A. extensum Smith SK
(Subgenus C OELOCEPHALAPION Wagner)
A. carinatun. Smith ON
concoloratum Srlrilh
A. emaciipes Fall ON
A. segnipes Siay ON
(Subgenus E UTRICHAPION Reitter)
A. alaskanun Fall AK YK NT BC AB
A. cavifrons .-eConte _BCAB -pe
A. cyanitinctum Fall AK YK NTBCAB_MB
A. huron Falt AB_MB ON
(Subgenus F,{ILAPION Kissinger)
A. finitimum FaIi
A. impeditum Fall _ _ BCT SK MB ON PQ
PQ
NB

A.
A.
melanarium Gerstaecker
occidentale Fall _AB -oN PQ

A. pennsylvanicum Boheman BC AB SK mn oN PQ
erythrocerum Srlrith
A. robustum Smith AB SK
obesum Srrlith
A. speculifenrm Fall AB SK MB
(Subgenus L-APION Roudier & Tempere)
(ftios Deville)
A. frosti Kissinger -oNPQ NS
A. idiastes K.,ssinger -ONPQ
(Subgenus PERAPION Wagrrer)
A. punctinasrm Smith BC AB SK MB ON
A. wickhami Kissinger _ABSK

328
 Curculionidae

(Subgenus PSEUDAPION Schilsky)

A. amaurum Kissinger -AB_MB
_ABSKMB
A. capitone Kissinger
A. disparatum Sharp _AB_MB
nasutum Fall
(Subgenus RHOPAIAPION Schilsky)
A. longirostre Oliviert BC ON PQ
(Subgenus TRICHAPION Wagner)
A. centrale Fall AB SK MB ON
commodum Fall _:"
A.
A. porcatum Boheman TSKMB ON
A. proclive LeConte -BC
crassinasum LeConte
nunenmacheri Wagner
A. reconditum Gyllenhal -oN PQ
turbulentum Smith
A. simile Kirby BC AB SK MB ON PQ NB NS NF
lanuginosum. Walsh
uicinum Srrrith
walshii Smit}r
A. tenuirostrum Smith AB SK MB
$

A. antennatum Smith AK YK BC

Tlibe Aplemonini
Genus PODAPION Riley
P. gallicola Riley ON PQ
alemnderi Sleeper

Family CURCULIONIDAE snout beetles or weevils

J. McNamara

The family Curculionidae includes 609 species the North American species with supplements in
and subspecies in Canada and Alaska. AII of them 1984 and 1989, the latter under Wibmer and
are phytophagous. Most of the larvae live inside O'Brien. The classification used here follows that
the tissues of the host plant, but some feed exter- checklist.
nally on the leaves. The adults feed on leaves,
pollen, flowers, developing fruit, and fungi, and
some burrow in wood; they do not necessarily feed
on plants preferred by the larvae. Many weevils are
host-specifrc and some are economic pests. AK (61); YK (51); NT (55); BC (261); AB (206); SK
Kissinger (1964) wrote a key to the genera. (I24); MB (179); ON (312); PQ (280); NB (72); NS
O'Brien and Wibmer (1982) published a checklist of (91); PE (17); LB (11); NF (68) ' H (14); I (69)

Subfamily BRACITYDERINAE
Tribe Naupactini
Genus ASYNONYCHUS Crotch
A. cervinus (Boheman)t ON
fulleri (Il.orn)

329
Curculionidae

Tribe Barynotini
Genus TRI(IONOSCUTA Motschuisky
Key to North American species: Pierce (1975)
(Subgenus I)RIGONOSCUTA s. str.)
T. pilosa iritlescens Pierce - BC

Genus STAMODERES Casey

S. lanei {Van Dyke) - BC

Genus BAR'INOTUS Germar

Key to Nc'rth American species: Brown (1950)
B. moerens Fabricius)t NS
B. obscurus (Fabricius)t - BC PQ NB NS PE - NF
B. schoenherri (Zetterstedt)t PQ NB NS NF

Tribe Polydrosini
Genus POLYDRUSUS Germar
Cyphontimus Horn
Poly drt ts u s Schcinherr
Key to Ncrth American species: Sleeper (1957a)
P. americanrs (Gyllenhalt - ON PQ
d.orsali:; (Horn)
P.cervinus,Linn6tt PQ
(Gyllenhal)+
P. impressif'rons - - MB ON PQ - NS
Schallertf
P. sericeus ( - ON PQ NB
Genus PACI{YRHINUS Schrinherr
S cy t hrt:. pus Schdnherr
(Horn)
P. californic -rs - BC
P.elegans(Oouper) - BC AB ON PQ - NS
metall i c us (Desbrochers)
(Casey)
P. ferrugine:s - BC

Tlibe Brachyderini
Genus BARYPEITHES Jacquelin du VaI & Fairmaire
B. pellucidus (Boheman)t - BC - ON PQ - NS PE - NF

Genus SCIAPHILUS Schtinherr

S. asperatus (Bonsdorff)t - BC - ON PQ NB NS NF

Genus BRAOHYSOMUS Schonherr

Thricol zpoide s O'Brien
B. echinatus (Bonsdorfl)* PQ NF
clauisetis (O'Brien)

Tribe Strophosomini
Genus STROPHOSOMA Billberg
S. melanogrammum (Forster)f - BC - ON PQ - NS NF

Tlibe Cneorhinini
Genus PHILOPEDON Schtinherr
P. plagiatunL (Schaller)t _NBNSPE-NF

330
 Curculionidae

Tribe Sitonini
Genus SITONA Germar
S. californicus (Fahraeus) BC
angustulus (Casey)
apacheanus (Casey)
cockerelli Blaisdell
explicatus (Casey)
ertrusus (Casey)
hispidiceps (Casey)
oregonensis Tanner
prominens (Casey)
S. cylindricollis (Fahraeus)t _BCABSK MB ON PQ NB NS
alpinensis Tanner
S. hispidulus (Fabricius)t
S. lepidus Gyllenhal
S. Iineatus (Linn6)t
f_ -BC
_BC
-BC BC
AB
oN
MB ON
PQ
PQ
NB
NB
NS
NS NF

S. lineellus (Bonsdorfi)t -YK NT AB sr MB ON NB NS

S. lupinus Sleeper _BC
S. scissifrons Say AK_ NT BC AB SK ln" oN pe lln NS NT
uittatus (Couper)
S. tibialis (Herbst)t AK _BC AB _ONPQ NF

T[ibe Tanymecini
Genus MINYOMERUS Horn
Elissa Casey
Pseudelissa Casey
M. innocuus Horn AB

Genus TANYMECUS Germar

T. confusus Say NT-ABSK MB
confertus Gyllenhal
uariabilis Fahraeus

Genus PANDELETEIUS Schdnherr

Key to North American species: Howden (1959)
P. hilaris (Herbst) _ONPQ
pauperculus Gyllenhal

Subfamily OTIORFIYNCHINAE
Tlibe Otiorhynchini
Genus OTIORIIYNCHUS Germar
B r ac hy r hinu s Latreille
Key to North American species: Warner & Negley (1976)
O. desertus Rosenhauerf NF
O. Iigneus (Olivier)t NSNS- NF
O. ligustici (Linn6)t -Olf-
O. ovatus (Linn6)t AK YK NT SC AB SK MB ON PQ NB NS LB NF
O. porcatus (Herbst)t PQ NF
O. raucus (Fabricius)t -oN ___
O. rugifrons (Gyllenhal)f PQ NB NS NF
O. rugosostriatus (Goeze)f -sc- -ONPQ _NS
O. scaber (Linn6)t _NS
O. singularis (Linn6)l _BC -ONPQ NB NS NT
picipes (Fabricius)
O. sulcatus (Fabricius)t AK _BC _MBON PQ NB NS PE NF

Genus SCIOPITHES Horrr

S. obscurus Horn BC
angustulus Casey
brumalis Casey
significans Casey

331
Curculionidae

Genus AGRONUS Horn

A. carri Bu<.hanan - BC AB

Tribe Peritelini
Genus OMIAS Germar
O. erectus Flatch - BC
O. minor Hatch - BC
O. saccatus {LeconteJ - BC

Genus PERITELINUS Casey

P. oregonus Van Dyke - BC
P. variegatus Casey - BC

Genus GEODERCODES Casey

G. latipennis Casey - BC

Genus NEI!-OCESTES Van Dyke

Key to North American species: Van Dyke (1936)
N. horniVanDyke - BC - ON PQ NB NS
N. incomptr.s tHornt - BC
N. montanus Van Dyke - BC
N. puncticollis tCaseyr - BC
sordidt:s Van Dyke
N. tuberculi,tus Van Dyke - BC

Genus PAR,\PTOCHUS Seidlitz

P. sellatus ( BohemanJ BC
califorr'.icus Seidlitz

Tribe Trachyphloeini
Genus TRAT,IHYPHLOEUS Germar
Chaete':.:hus }lorrl
Key to Ncrth American species: Brown (1965)
T. aristatus tGyllenhal r* - ON
T. asperatur Bohemant - ON NS
setiger:.Hotn)
T. bifoveolatus (Beck)f - BC AB ON PQ NB NS PE - NF
doujsi l3latchley

Genus CER{IOPEDIUS Sleeper

C. artemisiae (Pierce) - BC
paruultls (Van Dyke)

Tlibe Hormorini
Genus HORMORUS Horn
H. undulatus (Uhler) AB - MB ON PQ NB NS

Genus AGAIIPIIAEROPS Horn

A. niger Hor n - BC
sulcirostris (Pierce)

Tlibe Phyllobiini
Genus PHY,LOBIUS Germar
P. glaucus tiicopoli t+
P. oblongus r.Linn6)f - BC - ON PQ NB - PE

Genus EVOIIUS LeConte

E. naso (Le(ronte) NT BC AB SK

332
 Curculionidae

Genus APHRASTUS Say

M icrony c h us Provancher
A. taeniatus Say ON PQ
sulcatus (Provancher)

Tribe Leptopiini
Genus PHYXELIS Schdnherr
Geoderces }Iotn
P. rigidus (Say) oN PQ NB NS PE NF
glomerosus Boheman
melanothrix (Kirby)
setiferus Boheman

Genus LEPIDOPHORUS Kirby

Lophalophus LeConte
Key to North American species: Buchanan (1936b)
L. inquinatus (Mannerheim) AI{ BC
L. lineaticollis Kirby AK YK NT BC
L. pumilus Buchanan -BCAB
Genus CIMBOCERA Horn
C. pauper Horn AB

Genus PLINTHODES LeConte

P. taeniatus (LeConte) BC

Genus TRICHALOPHUS LeConte

T. alternatus (Say) _YK-BC AB MB ON
T. brunneus Van Dyke AK BC
T. didymus (LeConte) AK BC AB SK MB
constrictus (LeConte)
T. seriatus (Mannerheim) AK
T. simplex LeConte NT AB SA MB
T. stefanssoni Leng AK_NT
Genus LEPESOMA Motschulsky
Amnesia Horn
Dyslobus LeConte
Melamomphus l{orn
Thricomigus Horn
Key to North American species: Van Dyke (1933)
L. alternatum (Horn)
L. bakeri (Van Dyke)
BC
BC YSK
L. decoratum (LeOonte) BC
L. granicolle granicolle (LeConte) BC
debilis (Casey)
discors (Casey)
sculptilis (Casey)
L. lecontei (Casey) BC
L. luteum (Horn) BC SK
L. raucum (Horn) BC
ciliatum (Pierce)
L. tanneri (Van Dyke) BC
L. ursinum (Horn) BC
L. verruciferum (Casey) BC
b ituberculatum (Pierce)

Genus PANSCOPUS Schdnherr

Key to North American species: Buchanan (1936o)
(Subgenus NEOPANSCOPUS Pierce)
P. aequalis (Horn) BC AB SK MB
uestitus (Casey)
(Subgenus NOCHELES LeConte)
(Panscopidius Pierce)
P. torpidus (LeContet BC

333
Curculionidae

(Subgenus IIIOMIDUS Casey)

P. abruptus (Casey) - BC
P. ovalisPi,rrce - BC AB SK
P. pallidus Suchanan - BC AB
P. rugicollis Buchanan - BC
(Subgenus I)ANSCOPUS s. str.)
P. erinaceu,r (Say) - ON PQ
carinat:us Pierce
(Subgenus PARAPANSCOPUS Buchanan)
P. maculosu s Blatchley - ON
P. ovatipennis Buchanan - ON
(Subgenus PlIfMAfl NUS LeConte)
P. gemmatu.s (LeConte) - BC
(Subgenus PSEUDOPANSCOPUS Buchanan)
P. costatus Buchanan - BC

Genus CON NATICHELA Anderson

C. artemisir,Le Anderson AK YK NT

Genus ANAMETIS Horn

A. granulatrr (Say) - BC AB SK MB ON PQ NF
grisea':Jorn

Genus TRO PIPHORUS Schrinherr

T. elevatus Herbst)t - NF
carinatus (O.F. Mtiller)
T. obtusus (Bonsdorff)t NS NF
T. terricola.Newman)l PQ - NS PE - NF

Tribe Ophryastini
Key to North American species: Kissinger (1970)
Genus OPIIRYASTES Say
AmydrDgmus Pierce
Eupagoderes Horn
?ososfes Sharp
O. cinerascens (Pierce) - BC
columbianus (Van Dyke)
O. sulcirost:ts (Say) NT - AB SK MB
ligatus LeConte
O. tuberosu; LeConte AB
collaris Champion

Subfamily EREMNINAE
Tribe Cyphicerini
Genus CYR'IEPISTOMUS Marshall
C. castaneu; (Roelofs.tt - ON

Genus CAI-(JMYCTERUS Roelofs

C. setarius lloelofsf - ON

Subfamily RIIYTIRRHININAE
Genus LISTRONOTUS Jekel
Hyperodes Jekel
MacroP'9l{n"rY
Key to Nc,rth American species: Henderson (1940); O'Brien (1981)
L. alternatus (Dietz) PQ NB
L. anthracir.us {Dietz) - ON PQ

334
 Curculionidae

L. appendiculatus (Boheman) BC SK MB ON PQ
floridensis Blatchley
impressus Van Dyke
leucozonatus Chittenden
L. californicus (Dietz) -AB ON prl
L. caudatus (Say) BC
L. delumbis (Gyllenhal) BC AB SK MB ON pa:)

indistinctus (Dietz)
schwarzi (LeConte)
solutus (Boheman)
spurcus (Boheman)
L. echinodori O'Brien _MB
L. frontalis LeConte EC AB SK_ON PQ
L. humilis (Gyllenhal) A"- BC PQ NF
subcribratus (Dietz)
L. maculicollis (Kirby) AK YK BC AB PQ
mirabilis (Dietz)
myasellus (Dietz)
L. montanus (Dietz) AB SK MB
L. oregonensis oregonensis (LeConte) _MBON PQ
impressifrons LeConte
rudipennis Blatchley
L. oregonensis tessellatus Casey AB
L. porcellus (Say) BC PQ
minitnus (Blatchley)
L. punctiger LeConte BC AB
gracilis LeConte
leechi Sleeper
L. sordidus (GyIlenhal) ON PQ
disting uendu s (Gyllenhal)
obliquus LeConte
L. sparsus (Say) NT_AB _MBONPQ
inbellis (Dietz)
itnmundus (Boheman)
latiusculus (Boheman)
lineatulus (Say)
longulus (Dietz)
squalidus (Gyllenhal)
L. squamiger (Say) BC AB SK MB ON PQ
inaeq uali pennis (Boheman)
L. truncatus (Hatch) BC
L. tuberosus LeConte _MBON Pc)
L. vitticollis (Kirby) AB SK MB
interp unctatulus (Dietz )
inter s tit ialis (Dietz)
tenebrosus (Dietz)
ulkei (Dietz)

Genus LIXELLUS LeConr,e

Anchodemus LeConte
L. filiformis LeConte
L. Iutulentus (Boheman)
angustus (LeConte)
- Tf'BCfsKMBoNiS- - - - -

Genus BYRSOPAGE S Schdnherr

B. carinatus Motschuisky AK

Subfamily THECESTERNINAE
Genus THECESTERNUS Say
Lithodus Germar
T. affinis (LeConte) AB
eroszs (LeConte)
morbillosus (LeConte)
rectus (LeConle)
rudis (LeConte)
T. humeralis (Say) AB

335
 Curculioni lae

Subfamily FIIPERINAE
Tribe Hyperini
Genus HYPERA Germar
Phytonomus Sch6nherr
H. castor (LeConte)
H. compta (Say)
-sKMB-PQ
diuersz (Gyilenhal)
-BC-ONPQ
H. diversipunctata (Schrankrx AK - AB SK - ON
H. eximia (LeConte)
H. meles (['abricius)t
H. nigriros iris (Fabricius)i
- - Y- ItuSNpeN"Ns- -
- - -BC-ONPQNBNSPE-NF -
uiridi.. (Provancher)
H. postica iGyllenhal)f ABONPQ-NS
H. pubicoli.s (LeConte) _BC
H. punctatir (Fabricius)t -BC-ONPQ-NSNF
opimus (LeOonte)
H. quadricollis (LeConte) AB-MB
H. rumicis (Linn6)+ AB SK MB ON
H. seriata (Mannerheim) AK YK
H. trivittata (Say) NTBCAB_MB
setigera (LeConte)

Subfamily CLEONINAE
TYibe Cleonini
Key to New World species: Anderson (1988)
Genus CLtr.ONIS Dejean
Cleont..s Sch<inherr
Geomt:.rus Schtinherr
C. pigra (Stopoli t* ON pe
- NB
Genus STEPHANOCLEONUS Motschuiskv
Con iot l eon u s Motschul sky

S. confusus R.S. Anderson NT - AB SK

r
S. cristatict llis (Csiki AB
cristat,:ts LeConte
S. immacul,rtus R.S. Anderson - - NT BC AB SK MB ON
S. parshus R.S. Anderson NT BC AB SK MB ON pe
S. plumbeus LeConte AB SK MB _ pd NF
S. zherichir..i Ter-Minassian - yK
& Ktrotyaev*
stenotl,orax R. S. Anderson

Genus CLEONIDIUS Casev

Lirestt.:s Reitter
C. calandrordes tRandall) pe NB
C. erysimi tFallr - BC AB - MB
C. frontalis (LeConter AB SK MB
sparsus (LeConte)
sfralus (Csiki)
C. longinasus R.S. Anderson - BC
mixtus (LeConte\
C. poricollis (Mannerheim) BC AB
hirbyi (Casey)
- SK
lobigerinus (Casey)
uirgatu.s (LeConte)
uittatus (Kirby)
C. puber"ulus (Lecontet AB - MB
C. trivittatr:s (Sayt - BC AB SK MB
bicarinatus (Casey)
inornai'.us (LeConte)
praepotlens (Say)

336
 Curculionidae

Tribe Lixini
Genus LIXUS Fabricius
L. caudifer LeConte
L. concavus Say
coloradensis Chittenden
-:"--MBoNpe-
crassulus Notman
rectus LeCorte
soror Casey
L. laramiensis Casey AB
bischoffi Fall
L. macer LeOonte -oN
blatchleyi Csiki
capitatus Chittenden
cauicollis Blatchley
nouellus Blatchley
L. mucidus LeConte ON PQ
morulus Blatchley
L. parcus LeConte AB ON
pygmaeus Casey
L. punctinasus LeOonte
albisetiger Chittenden
ele phantu lus Chittenden
planicol I is Chittenden
L. rubellus Randall NT BC AB SK MB ON PQ
auctus LeConte
L. terminalis LeConte AB_MBON
blakeae Chittenden

Tlibe Rhinocyllini
Genus RHINOCYLLUS Germar
R. conicus (Frolich)f _SK_ONPQ

Subfamily MOLYTINAE
Tlibe Emphyastini
Genus EMPHYASTES Mannerheim
E. fucicola Mannerheirn AK BC

Genus THALASSELEPHAS Egorov & Korotyaev

Neophycocoetes O'Brien & Wibmer
Phycocoetes LeConte
T. testaceus (LeContet - BC

Tlibe Hylobiini
Genus LEPYRUS Germar
Key to North American species: Van Dyke (1928b)
L. capucinus (Schaller) NT
AK YK NT
L. gemellus gemellus Kirby
L. Iabradorensis Blair
L. nordenskioeldi alternans Casey
NT
BC
PQ - - is-
LB
L. nordenskioeldi caesius Csiki AK YK
cinereus Van Dyke
L. nordenskioeldi canadensis Casey AI{ NT BC AB SK
L. oregonus oregonus Casey AK _BC
perforatus Casey
_BCAB
L. oregonus tessellatus Van Dyke
L. palustris (Scopoli)x
(Lint6)
NTBCAB_MBONPQ is-
colon
geminatus Say

Genus IfYLOBIUS Germar

Hypomolyx LeConte
Key to North American species: Millers, Benjamin, and Warner (1963)

337
Curculioni*1ae

H. congener Dalla Torre, AK YK NT BC AB SK MB ON PQ NB NS LB NF

Schenkling & Marshall
confusus Ilttby
H. pales (Ilerbst) AB oN PQ NS
macelous (Germar)
H. pinicola (Couper) _YKNTBC AB SK MB ON PQ NB NS NF
H. radicis Iluchanan MB ON PQ :"
H. warreni Wood -BC AB MB ON PQ NB NS NF
Genus PACIIfYLOBIUS LeConte
P. picivoru:; (Germar) ON LB
stupitius (Boheman)

Tribe Plinthini
Genus STEREMNIUS Schcinherr
Parap,t,inthus Faust
S. carinatus (Boheman) AK BC
S. tuberosus Gyllenhal AK BC
s crob ic ulatus (Mannerheim)

Genus STFI EREUS Motschulsky

Lobosuma Buchanan
P hilo s lrat u s Zimmerman
Key to Nrrth American species: Zimmerman (1g64)
S. horridus (Mannerheim) AK _BC
S. multitutrerculatus Buchanan AK _BC
S. ptinoides; (Germar)x AK _BCNS I\TIT
boreaL,s Motschulsky
fas cic u.Iatu s Motschulsky
S. quadritu.berculatus Motschulsky AK BC SK

Genus GASTROTAPHRUS Buchanan

G. barberi ]luchanan BC

Subfamily ACICNEMIDINAE
Genus TRACHODES Germar
T. hispidus (Linn6)t NF

Subfamily PISSODINAE
Genus PISIIODES Germar
P. affinis Rrrndall _NTBCAB ON PQ rtlB NS NF
czrriel Hopkins
P. costatus Mannerheim AK BC
P. fasciatus LeConte BC
P. fiskei Hopkins
5_:cysK rasoNFeNs----
P. nemorensis Germar MB ON PQ NB NS
approrimatus Hopkins
canadensis Hopkins
deodarae Hopkins
P. radiatae Hopkins -BC
P. rotundatus LeConte AK YK _BCAB_MBONPQ olu Ns -
alascensis Hopkins
nigrae Hopktns
P. schwarzi Hopkins BC AB
yosemite Hopkins
P. similis Hopkins BC -PQ _NS NF
utahensis Hopkins
P. striatulus (Fabricius) BC ON PQ NB NS NF
dubius Ratdall
froserl Hopkins
prperi Hopkins
P. strobi (Peck) BC AB SK MB ON PQ NB NS NF

338
 Curculionidae

engelmanni Hopkins
sitchensis Hopkins
P. terminalis Hopping BC

Subfamily ERIRHININAE
TYibe Erirhinini
Genus DORYIOMUS Germar
Alycod,es Dietz
Key to North American species: O'Brien (19706)
D. brevicollis LeConte ON PQ
dubius (Dietz)
fuscipes Casey
indifferens Casey
D. frostii Blatchley YK NT BC AB SK MB ON PQ
D. hirtus LeConte -BCAB
columbianus Sleeper
hispidus LeConte
nubeculinus Casey
D. imbecillus Faust AK YK NT AB oN PQ
alaskanus Casey
subsimilis Blatchley
D. inaequalis Casey -BCAB SK MB
D. laticollis LeConte AK BC AB sK Yu oN PQ
D. lecontei O'Brien AK
D. leucophyllus (Motschulsky) AK YK NT BC
adustus Motschulsky
bajulus Farst
longulus LeConte
sub sig natus (Mannerheim)
te s sel latu s Motschulskv
D, luridus (Mannerheim) AK YK NT BC AB SK _PQ _NS
cuneatulus Casey
D. mannerheimi (Gemminger) AK NT BC AB -PQ
uestitus (Mannerheim)
D. marmoreus Casey
D. mucidus (Say)
D. parvicollis Casey
v_ -ec
AB
AB
AB SK mn
ON PQ
_PQ
oN PQ
_NS
NB_NF
amplus Casey
D. rufulus (Mannerheim) AK YK AB oN PQ NS
D. squamosus LeConte _1 YBC AB SK MB
angustatus (Dietz)
tesselatus (WaIsh)
D. vagenotatus Casey -YK NT BC AB SK MB ON PQ NF

Genus ERIRHINUS Schdnherr

E. aethiops (Fabricius)* AK YK NT BC AB SK MB ON PQ NS
morio Mannerheim

Genus GRYPUS Germar

Grypidius Schiinherr
Key to species: Cawthra (1957)
G. brunnirostris (Fabricius) ON PQ
G. equiseti (Fabricius)* AK YK NT BC AB SK MB ON PQ
G. Ieechi (Cawthra) -BCAB SK MB ON

Genus NOTARIS Germar

Key to North American species: Buchanan (1927)
N. bimaculatus (Fabricius) _YKNTBCABSKMBON PQ NB NS NF
wyoming ensis Chittenden
N. flavipilosus Chittenden
N. puncticollis (LeConte) T- NT BC AB SK MB ON pe NS NT

Genus PROCAS Stephens

P. lecontei Bedel YK NT MB ON PQ

339
Curculioniclae

Tlibe Smicronychini
Genus SMICRONYX Schonhen
Key to North American species: Arderson (1962)
(Subgenus DESMORIS LeConte)
S. fulvus LrrConte AB SK - ON
S. humilis rDietzr - SK MB
S. incertus tDietzt MB
rustict',s Dietz
uitiostt:.s Dietz
S. pinguis Jllatchley - ON
S. sordidus LeConte MB
montazus (Dietz)
(Subgenus .PACFfYPHANES Dietz)
(P ac h5, ty c hius LeConte)
S. amoenus (Say) - SK MB
S. centralis (Dietz t AB
lateralis (Dietz)
S. discoideus {LeConte) - SK MB ON
carus lDietz)
(Subgenus PSEUDOSMICRONYX Dietz)
(Syner:ha Dietz)
S. commixt rs Dietz AB SK MB
coseyi Blatchley
S. corniculatus (Fahraeusl - ON PQ
columl:'ianus Dietz
conniuens Casey
fallacir>sus l{]ima
fallar Dietz
laungi;zosus Dietz
spurcus Casey
squam'.tlatus LeConte
S. flavicans LeConte - ON
dietzil1ima
macuh:tus Dietz
nebulosus Dietz
ornatil:tennis Dietz
S. squalidur Casey - ON
cinerascens Dietz
morio l)ietz
Dietz
S. tessellatrLs MB ON
Dietz
im p r essiros t ris
S. utilis Bu<'hanan AB SK
(Subgenus TIMICRONYX s. str.)
S. cinereus 'Motschulsky) - BC
perplexus Dietz
S. congesturi Casey MB
S. sculpticollis Casey - ON
gibbirostris Casey
sagittatus Casey
S. tychoides LeConte - ON
S. vestitus l.,eConte AB - MB

Genus PRO.MECOTARSUS Casey

P. densus Cnsey - BC
P. lumatus tlasey - SK

Tlibe Tanysphyrini
Genus TANYSPIIYRUS Germar
T. aterBlatchley - BC - SK - ON
makolskii Smreczynski
T. lemnae (Fabriciustf - BC AB SK MB ON PQ

340
 Curculionidae

Tribe Bagoini
Genus BAGOUS Germar
Key to North American species: Tanner (1943)

_ _ _ _ 3N;:_ _ _ _
B. americanus LeConte
B. aLratus Blatchley
_
B. bituberosus LeConte
B. blanchardi Blatchley ON
B. magister LeConte ON
B. mamillatus Say _ABONPQ
B. nebulosus LeConte BCABSK_NB
ochraceus Blatchley
B. obliquus LeConte _ONPQ
B. planatus LeConte _ONPQ NB
B. restrictus LeConte --ec-- -ONPQ
B. tanneri O'Brien -oN
longirostrus Tanner
B. transversus LeConte BC AB SK MB ON PQ
carinatus Blatchley

Genus LISSORHOPTRUS LeConte

Key to species: Kuschel (1951)
L. oryzophilus Kuschel _AB
L. simplex tSayt _ABONPQ
egezus (Gyllenhal)

Genus BRACHYBAMUS Germar

B. electus Germar PQ
inceratus Boheman

Genus NOTIODES Schtinherr

Endalus Castelnau
Notionomus Erichsorr
N otiop hilus Schtinherr
Key to North American species: Tanner (1943)
N. aeratus (LeConte) _ .AB
N. limatulus tGyllenhal t -ABSKMBONPQ
_ABONPQ
N. ovalis iLeConte)
N. punctatus (LeConte) BCAB_MB
Genus ONYCHYLIS LeConte
Key to North American species: Burke (1961)
O. nigri rostris (Boheman t
_ONPQ

Tribe Derelomini
Genus PHYLLOTROX Schrinherr
Euclyptus Dietz
P. ferrugineus LeConte -oN
testaceus (Dietz)
P. nubifer LeConte _BCAB_MB
fuluipennis Sleeper
P. rutilus (FaIl) _BC
Genus ACAI-YPTUS Sch<inherr
A. carpini (Herbst) AK YK NT BC AB SK MB ON PQ

Subfamily OTIDOCEPHALINAE
Tribe Otidocephalini
Genus MYRMEX Sturm
O tidoce p halus Chewolat
M. chevrolatii (Horn) _ONPQ
rny r mecodes (Chewolat)
M. laevicollis (Horn) -oN

341
Curculionitiae

M. myrmex (Herbst) PQ
americ tnus (Chevrolat)
herbstti Stunn
myrmecodes (Say)

Subfamily MAGDALINAE
Tribe Magdalini
Genus MA(I.DALIS Germar
M. aenescerLs LeConte AK BC AB
M. alutacea LeConte AK YK NT BC AB PQ
M. armicoll s (Say)
pallida (Say)
SK -oN PQ

M. austera austera Fall

M. austera substriga Fall
-oN PQ
M. barbita (Say) MB ON PQ
M. cuneiforrnis Horn BC
M. gentilis LeConte YK BC AB SK PQ
M. gracilis (LeConte) BC
M. hispoides LeConte YK BC AB -oN pe NF
M. imbellis (LeConte) BC AB
M. inconspir:ua Hor-rr
M. lecontei ,lecepta Sleeper BC
-oN PQ

M. olyra (Horbst) pe
brunnices (Gyllenhal)
M. pandura (Say) MB ON PQ
M. perforatr,r Horn -oN PQ
M. piceae B rchanan -oN PQ
M. salicis Hcrn
sx
-oN PQ
M. subtincta LeConte -BCAB

Tribe Laemosaccinr
Genus LAE.MOSACCUS Schainherr
L. nephele (Flerbst) ON
plagiatus (Fabricius)

Subfamily CURCTILIONINAE
Genus CURCULIO Linn6
Balani:tus Germar
Key to Nc,rth American species: Gibson (1969)
C. aurivestis Chittenden BC
b reu inti. s us Chittenden
C. caryae (lliorn) _ONPQ
C. caryatrypes (Boheman) -oN
cylindricollis (Casey)
hariolus (Casey)
picus (Boheman)
C. confusor,.Hamilton) ON
C. iowensis lOasey) SKMBONPQ_NS
e*ilis Chittenden
funicul aris Chittenden
i6ls Chittenden
nume n;',u s Chittenderr
C. nasicus (fjay) ON PQ NB NS PE
auctus (Casey)
ordinatus (Casey)
C. obtusus (Bianchard) MB ON PQ
neocorylus Glbson
C. pardalis (Chittenden) MB ON
appalachius (Casey
m ul t ifa sciatu s Chittenden
paruicollis (Casey)
C. proboscideus Fabricius ON PQ
cuneatus (Casey)
nasutus (Say)

342
 Curculionidae

quercus (I{orrr)
r ectiro str is ( Gyllenhal )
rectus (Say)
rostrdtus (Gyllenhal)
sparsellus (Casey)
C. sayi (Gyllenhal) _ONPQ
acuminatus (Casey)
algonquinus (Casey)
auriger (Casey)
macilentus (Casey)
mollis (Casey)
perexilis (Casey)
setosicornis (Casey)
sparsus (Gyllenhai)
strigosus (Casey)
C. strictus (Casey) MB ON
longipes (Casey)
tubulatus (Casey)
utensis (Casey)
uirginicus (Casey)
C. sulcatulus (Casey) ON PQ NS
baculi (Chittenden)
curtus (Chittenden)
striatus Chittenden
C. uniformis (LeConte) BC
occidentis (Casey)

Subfamily ANTHONOMINAE
Genus MACRORHOPTUS LeConte
Key to North American species: Sleeper (1957b)
M. hispidus Dietz
M. sidalceae Sleeper nc v SK

Tlibe Anthonomini
Genus ANTHONOMOPSIS Dietz
A. mixta (LeConte) _ONPQ
Genus CHELONYCHUS Dietz
C. Iongipes Dietz _BCABSKMB
Genus ANTHONOMUS Germar
Anthomorphus Weise
Cnemocyllus Dietz
Paranthonomus Dietz
Serarthrus Blatchley
Tachypterellus Fall & Cockerell
Tachypterus Dietz
A. confusus Dietz _BCAB
A. consimilis Dietz PQ
A. consors (Dietz) -BC
A. convictus Gates
A.
A.
corvulus LeConte
deceptus Sleeper
- - --BC ncAB- me3N;3Ns NS NF

A. elongatus LeConte -oNPQ

_BCABSKMBONPQ NS
A. haematopus Boheman
sycophantaWalsh
A. hirtus Leconte -BC
A. interstitialis Dietz -ONPQNB
A. lecontei Burke -BCABSKMBONPQ
uariegatus LeConte
A. melancholicus Dietz -BC
murinus Dietz
A. molochinus Dietz _ONPQ NS
A. musculus Say _ONPQ
sulcifrons LeConte
A, nebulosus LeConte -BC-ONPQ

343
Curculionidae

A. nigrinus.Boheman NT PQ
A. ochreopiltsus Dietz _BC
A. pictus Blatchley PQ
A. profundus LeConte ON PQ NB NS
uulpinz,:s Dietz
A. quadrigil:bus (Say) BC AB SK MB ON PQ
cerasi (List)
magna (List)
A. quesnelensis Sleeper BC AB
A. robustuh:.s LeConte _ABSK
rubidus L,eConte Dr\
A.
A. rubricus lJchenkling & Marshall
BC
BC
-oN
MB
rq

rubellus Dietz
A. rufrpes LeConte AB
A. rutilus (Boheman) PQ
nubiloides FalI
A. signatus lJay BC AB SK MB ON PQ NS PE NF
bisignalus Gyllenhal
pallidus Dietz
scut eI lo. tu s Gyllenhal
A. simiolus Illatchley PQ NB
A. sphaeralciae Fall ---AB-l4e
A.
A.
A.
squamosr.s LeConte
squamulatus Dietz
subfascia;us LeConte
_ _ BCTSKMB
ON PQ
latiusct;.Ius Dietz
rantus Blatchley
A. suturalis LeConte BC ON
erythropterus Say
flauicornis Boheman
hicoriat: Pierce
uespertinus Dietz
A. tectus Le(lonte SK MB
A. xanthocnr'mus Dietz ON

Genus COC()OTORUS LeConte

Key to No:th American species: Brown (1966)
C. pumilae r:rajor (W.J. Brown) _MB
C. pumilae pumilae (W.J. Brown) ON PQ
C. scutellari,; ( LeconteJ _MB
prunicida (Walsh)

Genus DIETZIANUS Sleeper

Xanthus Dietz
D. pygmaeus. tDietzt MB

Tribe Bradybatini
Genus PSEI.IDANTHONOMUS Dietz
Key to No;th American species: Clark (1987)
P. crataegi (Walsh) AB SK MB ON PQ NB NS
brunneus Dietz
longulus Dietz
paruulus Dietz
p un ct icol I is Blatchley
P. helvolus (13oheman) ON
hamamelidis Pie'rce
incipiens Dietz
relictus Dietz
P. rufulus Dietz _MB-PQ
P. seriesetosus Dietz oNPQ-NS
P. validus Dietz YT BCAB-MBONPQNBNS
facetus Dietz

344
 Curculionidae

Subfamily RFIYNCIIAENINAE
Key to North American species: Anderson (1989)
Genus RHYNCHAENUS Clairville
Orchestes llTiger
R. griseus Sleeper _BC
R. mixtus (Blatchley) _BCABSKMBONPQNBNSPE
R. pallicornis (Say) _BCAB_MBONPQNBNSNF
puberulus (Boheman)
uniformis (W.J. Brown)
R. testaceus (O.F. Mriller)x AK BC AB SK MB ON PQ NB NS NF
canus (Horn)
pallidior (Leng)

Genus TACHYERGES Schtinherr

T. ephippiatus (Say) -BCABSKMBONPQNB
T. niger (Horn) AK YK NT BC AB SK MB ON PQ NB NS _ LB
illinoisensis (Fall)
paruicollis (LeConte)
T. salicis (Linn6)x NTBCASSKMBONPQ-NSNF
subhirtus (Horn)

Genus ISOCHNUS C.G. Thomson

L arcticus (Korotyaev)x AK_NT
L flagellum (Erichson)* AK YK NT
I. rufrpes (LeConte) -BCAB-MBONPQ-NSNF
minutus (Horr')
I. populicola (Silfverberg)J _ONPQNBNS

Subfamily PRIONOMERINAE
Tlibe Prionomerini
Genus ODONTOPUS Say
O. calceatus tSay) - ON PQ
c ar bonar iu s ( Gyllenhal)

Tribe Piazorhinini
Genus PIAZORHINUS Schdnherr
P. pictus LeOonte MB ON PQ
P. scutellaris (Say) MBONPQ_NS

Subfamilv TYCHIINAE
Tribe-nlleschini
Genus ELLESCHUS Dejean
Alyca LeConte
E. bipunctatus (Linn6) MB ON PQ
E. borealis Carr AK YK AB - MB
E. ephippiatus (Say) - YK - BC AB SK MB ON PQ NF
rufus (Say)
E. scanicus (Paykullt AB SK - ON PQ
Genus PROCTORUS LeConte
Encalus LeConte
P. armatus LeConte AK - NT BC AB SK MB ON PQ NB
P. decipiens (LeConte) AK YK NT BC AB SK MB ON PQ NB NS

Tribe Lignyodini
Key to genera: Clark (1982)
Genus LIGNYODES Dejean
Key to North American species: Ciark (1980)
(Subgenus LIGNYODES s. str.)
(T hy sanocnemis LeConte)

345
Curculionid:re

L. bischoffi tBlatchley) - ON PQ
L. fraxini tl,eContet - SK - ON PQ
L. helvolus tl.eConte) MB ON PO
b alaninoide s ( Schaeffer)
breuis (Casey)
caseyi (IAima)
L. horridulus (Casey) - ON PQ
punctatus (Casey)
squamtger (Casey)
L. ocularis ('laseyt - ON

Genus PLO(IETES LeConte

Hamaba Casey
Rosella Whitehead
Key to spercies: Clark (1982)
P. ulmi LeConte - ON PQ

Tribe Tychiini
Genus SIBINIA Germar
Key to New World species: Clark (1978)
S. maculata (LeConte) AB
maculilbr (Hatch)

Genus TYCIIIUS Germar

Miccotrcgus Schdnherr
Paraty<:hius Casey
Key to North American species: Clark (1971)
T. aratus Say - SK
arator ()yllen}:al
T. liljebladi .3latchley AB SK MB
T. lineellus l,eConte - BC AB SK
dilectus Casey
hesperis Casey
probus Jasey
radians Casey
tacitus'Jasey
T. montanus Clark AB
T. picirostrit (Fabricius)t - BC AB SK MB ON PQ - NS PE - NF
posticu:,: Gyllenhal
T. semisqua:nosus LeConte - BC
intermi.ttus }Iatc}l
lamello.;us Casey
squamosus Hatch
T. soltaui Casey - SK MB
hirsutus Clark
hirtellu,s LeConte
T. stephensi Schrinherrt - BC AB ON PQ NB NS
griseus Schaeffer
tontentosus (Herbst)
T. tectus LeOonte AK YK - BC AB SK MB
languidus Casey
mixtus Hatch

Subfamily GYMNETRINAE
Tlibe Gymnetrini
Genus GYM.NETRON Schdnherr
G. antirrhinr (Paykull)t _BCABONPQ-NSNF
G. netum (Germar)t _BC
G. pascuorurn (Gyllenhal)t -BC_ONPQ
G. tetrum (Fabricius)t -BC-ONPQ

346
 Curculionidae

TYibe Mecinini
Genus MIARUS Schtinherr
Cleopomiarus Pierce
M. hispidulus LeConte MB ON PQ
consuetus Casey
illini Casey
nanus Casey
puritanus Casey

Genus MECINUS Germar

M. pyraster (Herbst)t - BC

Subfamily CRYPTORITYNCHINAE
Tribe Ithyporini
Genus RHYSSOMATUS Schdnherr
R. aequalis Horn ON
R. Iineaticollis (Say) oN PQ

Genus CONOTRACHELUS Dejean

Loceptes Casey
Key to North American species: Schoof ( 1942)
C. anaglypticus (Say) ON PQ
C. aratus (Germar) _PQ
C. corni W.J. Brown -MBONPQ
C. crataegi Walsh ON PQ
C. elegans (Say) ON PQ
C. geminatus LeConte _PQ
C. juglandis LeConte oN PQ
C. nenuphar (Herbst) BC -MBONPQ NS -NF
orgula (Fabricius)
cerasi (Peck)
C. nivosus LeOonte -AB
plagiatus LeConte
C. posticatus Boheman -oNPQ
C. recessus (Casey) -oN
atokanus Fall
C. schoofr Papp -oN
tibialis Schoof
C. seniculus LeConte -ONPQ
Genus PHELOCONUS Roelofs
P. cribricollis (Say)
puncticollis (Walsh)
-oN
Genus MICROIIYUS LeConte
Echinaspis Blatchley
M. setiger LeConte ON
wolcotti (Blatchley)

Tlibe Cryptorhynchini
Genus GERSTAECKERIA Champion
Opuntiaphila Pietce
Philopuntia Pietce
Key to North American species: O'Brien (1970a)
G. basahs (LeConte) AB

Genus ACALLES Schdnherr

A. carinatus LeConte ON

Genus TYLODERMA Say

Analcis Schdnherr
Key to North American species: Wibmer (1981)
T. aereum (Say) ON PQ

347
 Curculionidae

T. capitale \Vibmer -oN

T.
T.
foveolaLum tSayt
minimun Blatchley AB
-oN
T. nigrum Casey -oNPQ NB
asclepiusae Sleeper
T. oenotherare Wibmer AB SK MB

Genus CRYTTORHYNCHUS Illiger

C ry p to r hy nc hidiu s Pierce

C. fuscatus .-eConte ON PQ
C. lapathi tl,inn6tx BC ru _MB ONPQNBNSPE_NF
Genus APTINROMECHUS Faust
Key to Nc,rth American species: Whitehead (19Tg)
A. ferratus (Say) ON

Genus COPIIES Champion

Key to North American species: Sleeper ( 1955)
C. fallax (LeConte) ON
C. obtentus,'Herbst) ON
ypsilon (Boheman)

Genus EUBULUS Kirsch

Key to species: Fiedler (1939)
E. bisignatu; rSayr ON PQ
o b liq ue,hs ciatzs (Boheman)
E. parochus (Herbstr oN PQ
(.?) luctt:osus (Boheman)

Subfamily ZYGOPINAE
Key to North American genera: Sleeper (1963)
Tribe Zygopini
Genus CYLI NDROCOPTURUS Heller
Copturt'des Casey
Crjtrotus Casey
C. deleoni Brrchanan BC AB
C. longulus (LeConte) ON
disperst:.ts (Casey)
mucidu; (Casey)
nubilat:.ts (Casey)
obscure,'.lus (Casey)
subcupreus (Casey)
C. medicatusr Carr

Tlibe Lechriopini
Genus LECHRIOPS Schdnherr
Gelus Casey
Key to North American species Sleeper (1963)
L. californice. (LeConte)
L. oculata (Say) - :"- - oll PQ NS---
^ne
Genus ACOf'TUS LeConte
Homogaster Provancher
A. suturalis i,eConte ON PQ NB NS
q ue be cens is (Provancher)

Genus PSOMUS Casey

P. armatus (.r)ietz) ON PQ
politus ()asey

348
 Curcul.ionidae

Subfamily CEUTORIIYNCHINAE
Tlibe Mononychini
Genus MONONYCHUS Germar
M. vulpeculus (Fabricius) ON PQ

Tlibe Cnemogonini
Key to genera: Colonnelli (1979)
Genus CRAPONIUS LeConte
C. inaequalis (Say) ON

Genus CNEMOGONUS LeConte

C. iecontei Dietz AK YK NT BC AB MB ON PQ NB

Genus ACANTHOSCELIDIUS Dalla Torre & Hustache

Acanthoscelis Dietz
A. acephalus (Say) - BC AB SK MB ON PQ PE
leprosus (Boheman)
s ub uliro str i s (Gyllenhal)

A. curtus tSay) PQ

Genus AULEUTES Dietz

A. epilobii(Paykull)* - YK NT BC AB SK MB ON PQ NB NS
cruralis (LeConte)
A. nebulosus (LeConte) - BC - SK MB ON PQ NB
A. tenuipes (Leconte) -oNPQ
Genus PERIGASTER Dietz
Key to North American species: Buchanan (1931)
P. cretura (Herbst) oN PQ
quadrispinosa (Say)
P. liturata (Dietz) oN PQ NF
longirostris Buchanan

Genus DIETZELLA Champion

Coelogaster Schdnherr
D. zimmermanni (Gyllenhai) - BC ON PQ NF

Tribe Ceutorhynchini
Genus CEUTORIIYNCHUS Germar
C euthorhy nchidius J acqrelin du Val

AK PQ- NS
C. americanus Buchanan
C. assimilis (Paykull)f )' BC
BC 18:"MB 9*
C. bolteri Dietz BC
hirticollis Dietz
C. carteri W.J. Brown AB
C. convexipennis Fall MB
C. cupreus Hatch sc
C. dubitans W.J. Brown _SK
echinatus Fall MB

T_
C.
C. erysimi (Fabricius)f BC ON PQ NS
C. floralis (Paykull)f _ ON PQ NB NS I\'D
puberulus LeConte
sep tent r ionalis Gyllenhal
C. fulvotertius FaII
C. handlordi W.J. Brown
C. hearnei W.J. Brown
TY NT BC
f SK
MB
MB
oN-
C. invisus Fall s^ y"
C. lecontei Colonnelli
conuexicollis LeConte
-sc AB

C. munki W.J. Brown MB

C. neglectus Blatchley
T_ sc AB sx
_ABSK
MB ON PQ
C. omissus Fall MB oN PQ
C. opertus W.J. Brown BC

349
Curculionidae

C. oregonersis Dietz _YK-BCAB_MB

C. ovipenni; Dietz
C. pauxillur; Dietz - - - l" AB sx nre- - - - - - -
C. pervestit,us FaIl AB
C. punctige r Gyllenhalt _YK_BCABSKMBONPQNBNSNF
C. querceti lGyllenhal) AK _ NT BC AB _ MB - PQ NB NS PE _ NF
hamiltoniDietz
pzslo Mannerheim
C. rapae Gyllenhalt AKYK_BCABSKMBONPQ
affiuentus Dietz
C. squamatus LeConte _ONPQ
C. squamosulus Sleeper _BC
sleeperi Colonnelli
C. subpubescens LeConte _YKNT_MB
Genus POOPHAGUS Schttnherr
P. sisymbri:i (Fabricius)t PQ

Genus AMAIUS Schrinherr

A. scortillurn (Herbst)t -BCABSKMBONPQ_NSNF
robinsoni (Blatchley)

Genus CAL'CSIRUS Thomson

C. medialis (LeConte) _ONPQ
C. semirufu.s (LeOonte) AB ON PQ NF
C. zimmern rannii (Gyllenhal) MB ON
Genus ALLI)SIROCAIUS Colonnelli
A. angulatu.s (LeConte) _BCAB
A. consangu.ineus (Dietz) AB SK MB
A. disturbatus (Dietz) _BC
Genus HAT; ROPLONTUS Thomson
H. litura (F;rbricius)f -oN
Genus SIR()CALODES Voss
S. sericans (LeOonte) MB

Genus NEDYUS Schdnherr

Cidnorhinus Thomson
N. fl avicaudis (Boheman) AB SK MB ON PQ
apicalis (Dietz)

Tribe Scleropterini
Genus HONIIOROSOMA Frivaldszky
H. sulcipenrre (LeConte) - YK - SK MB ON PQ

Genus RUTIDOSOMA Stephens

R. decipiens (LeConte) AK YK - BC AB SK MB ON PQ

Genus ACA.LLODES LeConte

A. saltoides Dietz PQ
A. ventricosus LeConte MB ON

Tlibe Phytobiini
Key to New World genera: Colonnelli (1980)
Key to genera: Colonnelli (1986)
Genus PIIYIOBIUS Schdnherr
Hydaticus Schdnherr
Li todacty I us Redtenbacher
Parenthis Dietz

350
 Curculionidae

P. leucogaster (Marsham) YK NT BC AB SK MB ON PQ
griseomicans Schwarz
my r iop hy I li (Gyllenhal)
P. vestitus (Dietz) ON

Genus EUBRYCHIOPSIS Dietz

E. albertana (W.J. Brown) NT BC AB SK

Genus EUBRYCHIUS Thomson

E. velatus (Beck) BC
aquaticus Thomson

Genus RHINONCUS Schtinherr

C ampy lir hy nch zs Dej ean
Key to North American species: Hoebeke & Whitehead (1980)
R. castor (Fabricius)t _BC_ONPQ NB NS NF
R. longulus LeConte -BCABSKMBONPQ
_BCABONPQ NF
R. pericarpius (Linn6)t
umbellae (Fabricius)
R. pyrhopus Boheman NTBC_SKMBONPQ PE
R. triangularis (Say) AB SK MB ON PQ
occidentalis Dietz

Genus PELENOMUS Thomson

Mecopeltus Dietz
P. asperulus Dietz \tK AB ON
aeneosquamoszs (Dietz)
P. congenialis (Dietz) AB
P. fuliginosus (Dietz) BC AB ON PQ NB
P. gracilipes Dietz -PQ
griseus Blatchley
P. pusillus Dietz _PQ
scandens (.Dietz)
P. scoliasus (Dietz) AB MB
P. squamipennis (Sleeper) -BC
_BC pe-
P. squamosus LeConte YK AB ON
im pressi uen tris Blatchley
P. sulcicollis (Fahraeus) AB MB ON PQ
P. ventralis (Sleeper) YK BC AB

Genus NEOPIIYTOBIUS Wagner

N. cavifrons (LeConte) BC AB
toruipes (Dietz)

Subfamily BARIDINAE
Tribe Baridini
Genus BARIS Germar
Baridius Schiinhen
Cy p hi r h inu s Schainherr
Orthobaris Casey
B. canadensis Blatchley -oN
B. interstitialis (Say)
agnita Casey
-oNPQ
anaplata Casey
appalachia Casey
apposita Casey
aprica Casey
apricoides Fall
attonsa Casey
austiniana Casey
bispeculata Casey
brachyrhina Casey
breuipennis Casey
carinulata (LeConte)
caseyi HusLache
cribrifera Casey
cuneipennis Casey

351
Curculionidae

demissa Casey
dolosa Casey
fultoni Casey
immur,.is casey
irrita Casey
lauaca,za Casey
lubrica Casey
nephiana Casey
oblata Casey
oblong,tla Casey
penins,ilae Horn
quadr(:..ta (LeConte)
retrusa Casey
subaenea (LeConte)
surrufcL Casey
tumescens (LeConte)
uagans Casey
zuniana Casey
B. lengi Hustache ON
carbon,zria Blatchley
B. stacesmit;hi Sleeper BC
B. strenua (LeConte) MB
coIo ra c:;'.e n si s C as ey
formali',s Casey
longulicollis Casey
montanica Casey
satelles Casey
separara Casey
subpar'.lis Casey
uinta (.lasey
B. striata (Say) _MBON
B. subsimilis Casey ON

Genus COS.MOBARIS Casey

C. americarra Casey SK oN PQ
sionilli Hayes
squamLger Hayes

Genus AUL OBARIS LeConte

A. pusilla (l,eConte) PQ

Genus AMF'ELOGLYPIER LeConte

A. ampeiopsis (Riley) ON
ater LeConte

Genus PSEIJDOBARIS LeConte

P. angusta (LeConte)
angustula LeConte
P. nigrina (Siay) oN PQ

Genus TRICIHOBARIS LeConte

T. trinotata (Say) ON
impotens Casey
jejuniosa Casey
plumbea (LeConte)
ueslila (Boheman)

Genus ORTFIORIS LeConte

O. crotchii LeConte
O. tenuirostris Casey _ _ _:"ffi_ _ _ _ _ _ _ _ _

Tlibe Madarini
Genus ONYCHOBARIS LeConte
O. subtonsa LeConte
O. pectorosa. LeConte
- i{B
PQ

352
 Curculionidae

Genus MADARELLUS Casey

Willinkia Bondar
M. undulatus (Say) - ON PQ

Tribe Centrinini
Genus GERAEUS Pascoe
Centrinaspis Casey
G. picumnus (Herbst) - ON
albotectus (Casey)
compactus (Casey)
floridensis (Casey)
lasciuus (Casey)
oliuaceus (Gyllenhal)
paululus (Casey)
per scillus (Gyllenhal)
sulor (Harris)

Genus ODONTOCORYNUS Schdnherr

O. denticornis (Casey) - ON
O. scutellumalbum t$sy) - ON PQ

Genus CENTRINOGYNA Casey

C. canadensis Casey AB - MB

Genus CALANDRINUS LeConte

C. grandico)lis LeConte AB
angustulus Casey
C. obsoletus Casey AB
insignus Casey

Genus CATAPASTUS Casey

C. conspersus tl,eConte) - ON PQ

Genus CYLINDRIDIA Casey

C. prolixa tl-eContet MB ON PQ

Genus DIRABIUS Casey

(Subgenus DIRABIUS s. str.t
D. rectirostris (LeConte) ON PQ
tenua (Blatchley)

Genus PROSALDIUS Oglobin

Anacentrinus Buchanan
P. subaequalis (Casey) MB

Genus STETHOBARIS LeConte

S. commixta Blatchley - ON PQ
S. laevimargo {Championt - ON
S. ovata (LeConte) - ON PQ

Genus IDIOSTETHUS Casey

l. tubulatus tSayt - SK PQ
st rig apu nc t us (Harnilton)

Genus HAPLOSTETHOPS Casey

H. ellipsoideus (Casey) - ON

Genus PLOCAMUS LeConte

Epeuchaetes Lyman
Euchaetes LeConte
Eunyssobia Casey
P. echidna (LeConle) - ON
P. hispidulus LeConte - NB

353
Curcuiionii.ae

Subfamily RIfYNCHOPHORINAE
Tribe Sphenophorini
Genus SPHENOPHORUS Schrtnherr
Calandra Clairville
Trichi:;chius LeConte
Key to North American species: Vaurie (1951)
S. aequalis aequalis Gyllenhal - BC AB SK MB ON PQ
scirpi {Jhittenden
uniuitia Chittenden
S. australis australis Chittenden - ON PQ
typhae' Chittenden
S. callosus Oliviert - ON PQ
jugosus Chittenden
S. cariosus ,Oliviert NS
cicatricosus (Say')
flexuosus Gyllenhal
larualis (Germar)
sculptilis Uhler
S. cicatristriatus Fahraeus AB SK MB
cic at r i.,cennis Fahraeus
ulkei llorn
uariolc'sus LeConte
S. costipennis Horn - BC AB SK MB ON PQ NB
laeu ig t:.Lt u s Chittenden
me do r aensis ( Satterthwait)
S. incongruus Chittenden - ON
elephantulus Chittenden
S. melanoccphalus {Fabricius) - ON PQ
nubilus Gyllenhal
S. minimus Hart - ON
S. mormon Chittenden - BC AB SK MB
distich lidis Chittenden
S. parvulus Gyllenhal - ON PQ
pumilus Gyllenhal
S. pertinax pertinax (Olivier) - ON - NB NS
canaliculatus Boheman
inter u c: I I at us Chittenden
nasutu.s Chittenden
penins ularis Chittenden
sefiger Chittenden
truncatus (Say)
S. robustior Chittenden MB
costifer Chittenden
S. robustus Horn - BC MB ON
m uI t ih, neat us ( Satterthwait)
r ect i s t r iat u s Chittenden
S. sayi Gyllenhal - BC
s u s Mannerheim
ubca r inat
S. serratipes Chittenden AB
S. striatipe:rnis Chittenden AB SK MB ON NS
S. venatus ;enatus (Say) - ON PQ NB
confusus Gyllenhal
fallax Boheman
immunis (Say)
placid;ts (Say)
S. villosiventris Chittenden PQ
S. vomerinus LeConte AB
baridioides llotrr
S. zeae Walsh - ON PQ
blatchieyi Chittenden

Genus RHODOBAENUS LeConte

H otnaiosty I us Chelrolat
Key to North American species: Vaurie (1981)
R. quinquepunctatus (Say) - ON PQ
R. tredecimpunctatus tllliger) - ON PQ
cribrarius (Fabricius)
leptocerus (Panzer)
q uatuor decirn punc tatus (Panzer)

354
 Curculionidae

Tribe Sitophilini
Genus SITOPHILUS Schonherr
S. granarius (Linn6)f - BC MB ON PQ NF
r emotep unctatus Gyllenhal
S. oryzae (Linn6)t BC - ON PQ NF
S. zeamais Motschulsky* MB ON
Genus DIOCALANDRA Faust
D. elongata (Roelofs)t - BC

Subfamily COSSONINAE
Tribe Acamptini
Genus ACAMPTUS LeConte
A. echinus Casey
A. rigidus LeConte _ _ _ _ _ _ _ BNpe _ _ _ _ _

Tfibe Cossoninr
Genus HIMATIUM Wollaston
Himatinum Cockerell
H. errans LeConte PQ - NS
nigritulum Casey
paruula Blatchley

Genus COSSONUS Clairville

Borophloeus Wollaston
C. americanus Buchanan PQ - NS NF
C. crenatus Horn - BC
C. impressifrons Boheman - ON PQ
concinnus Boheman
C. pacificus Van Dyke - BC AB
C. piniphilus Boheman - BC
californicu s Motschulsky
scrobiculatus LeConte
C. platalea Say - ON PQ
bohetnanniHotn
subareatus Boheman
C. quadricollis Van Dyke - BC AB

Tlibe Rhyncolini
Genus PENTARTHRUM Wollaston
P. huttoni Wollastont PQ NF

Genus EUOPHRYUM Broun

E. confine (Broun)f - NF

Genus HEXARTHRUM Wollaston

H. thujae W.J. Brown - BC
H' ulkei Horn - oN PQ

Genus ELASSOPTES Horn

E. marinus Horn - BC

Genus STENOSCELIS Wollaston

S. brevis (Boheman) - ON PQ

Genus PHLOEOPHAGUS Schonherr

P. apionides Horn - ON PQ
P. cinadensis Van Dyke - BC AB - MB - PQ
P. minor Horn PQ
carinatus (Blatchley)

355
Scolytidae

Genus RHIINCOLUS Germar

Xylocc.mesus Thatcher
R. brunneu.s Mannerheim AK YK NT BC AB MB ON PQ NB NF
c alifor ni c u s Wollaston
relictus Casey
R. cylindricollis Wollaston -BC
R. knowltoni (Thatcher) MB
R. macrops Buchanan _BC ON pr) NS
R. oregonensis Horn _BC
R. pallens Oasey -BC
R. stacesm:thi Sleeper _BC
Genus CA-I|,PHONOTUS Casey
C. testaceus Casey AK BC AB ON PQ NB NF

Subfamily DRYOPHTHORINAE
Tribe Dryophthorini
Genus DRYOPHTHORUS Germar
D. america:rus Bedel MB ON PQ NB NS
corticalis Say

Family SCOLYTIDAE bark beetles

J. McNamara

In Canada and Alaska 204 species of Scolytidae The North American scolytids are well known
are recorded. Most of the Canadian species attack taxonomically; Wood (1982) revised all t;he North
forest trees, especially conifers, and may cause con- American species, and Bright (1976) covered the
siderabler damage to timber. Both the larvae and Canadian and Alaskan ones.
adults form galleries under the bark. Some of the
galleries may be so distinctive that the genus and, AK (52); YK (33); NT (31); BC (13a); AB (80); SK
in some ,iases, the species that made them can be (40); MB (a7); ON (89); PQ (81); NB (53); NS (a6);
identifieci. PE (8);LB (6);NF (27) . I (5)

Subfamily FIYLE SININAE

Tribe Hylastini
Genus SCIItrRUS LeOonte
S. annectans LeConte AK_NTBCAB ONPQ-NS NF
S. pubescens Swaine AKYK-BCAB
Genus HYLURGOPS LeConte
H. porosus (LeConte) NT BC AB SK
lecontei Swaine
H. reticulatus Wood NT BC
H. rugipennis pinifex (Fitch) BC AB SK MB ON PQ NB NS
H. rugipenrris rugipennis AK YK BC AB
(Mannerheim)
H. subcostulatus subcostulatus AK BC
(Mannerheim)
c r i s tat u s ( Mannerheim)

Genus IfYLASTES Erichson

H. gracilis LeOonte BC
asper Swaine
Iongus LeConte
nitidus Swaine
H. longicollis Swaine _BCAB
H. macer LeConte _BC

356
 Scolytidae

H. nigrinus (Mannerheim) AK YK BC AB
yukonis Fall
H. porculus Erichson MB ON PQ NB NS
canadensis Blackman
carbonarius Fihch
cau ernos u s ( Zimmermann)
granosus Chapuis
scober Swaine
swainei Eggets
ruebbi Swaine
H. ruber Swaine NT BC AB
H. tenuis Eichhoff _BC
criticus Eichhoff
minutus Blackman
pd.ruus Blackman
pusillus Blackman

Tribe Hylesinini
Genus HYLASTINUS Bedel
H. obscurus (Marsham)f BC _ONPQ NS

Genus HYLESINUS Fabricius

Leperisinus Reitter
H. aculeatus Say SK MB ON PQ NS
cinereus (Swaine)
imperialis Eichhoff
H. californicus (Swaine) MB
hoferi (B\ackman)
H. criddiei (Swaine) MB ON PQ
H. fasciatus LeConte -oN
H. pruinosus Eichhoff -oN
Genus AI-NIPI{AGUS Swaine
H y la s tinoides Spessivetsev
A. aspericollis (LeConte) AK _BC
A. hirsutus Schedl _BC

Tribe Tomicini
Genus PSEUDOIIYLESINUS Swaine
P. dispar dispar Blackman _BC
P. dispar pullatus Blackman
granulatus (LeConte)
-BC
_BC
P.
P. nebulosus nebulosus (LeConte) T -BC
_BC
AB
P. nobilis Swaine
furnissi Blackman
P. pini Wood _BC
P. sericeus (Mannerheim) AK _BC
grandis Swaine
P. sitchensis Swaine AK _BC
P. tsugae Swaine AK _BC
heeni Blackman
obeszs Swaine
similis Blackman

Genus HYLURGOPINUS Swaine

H. rufrpes (Eichhoffl SK MB ON PQ NB NS
opaculus (LeConte)

Genus XYLECHINUS Chapuis

X. americanus Blackman
X. montanus Blackman AK ecABsr-9*PQ-Tt---
Genus DENDROCTONUS Erichson
D. bre.ricomis LeConte BC AB
barberi Hopkins

357
Scolytidae

D. murrayar:.ae Hopkins
D. ponderosrLe Hopkins
monticolae Hopkins
_BCAB
_BC _ y"9*_ _ _ _ _ _

D. pseudotsugae Hopkins _BCAB

D. punctatus, LeConte AK YK NT BC AB -oN PQ NB NF
johanse.ci Swaine
D. rufipennis (Kirby) AK YK NT BC AB SK MB ON PQ NB NS NF
borealis Hopkins
engelmc:.nni Hopkins
o6eszs (Mannerheim)
piceaperda Hopkins
similis .i,eConte
D. simplex LeConte AX _ NT BC AB SK MB ON PQ NB NS PE LB NF
D. valens Lelonte _NTBCAB_MBONPQNBNSNF

Tribe Phloeotribinr
Genus PHLOEOTRIBUS Latreille
P. dentifrons (Blackman)
P. lecontei Schedl
puberulus LeConte
---ecAB- _9"
P. liminaris tHarris) SK MB ON PQ NB
mississt pp iensis (Blackman)
P. piceae Sweine AK YK NT BC AB MB ON PQ NB NS

Tribe Phloeosinini
Genus PHL()EOSINUS Chapuis
P. canadensis Swaine
P. cupressi l{opkins AK BC
-oNPQNB
nitidus Swaine
P. dentatus ( Say) ON
enieus l:llackrr.an
graniger (EichhofD
haagi (Eichhoff)
P. hoferi Blar:kman BC
P. hoppingi Swaine BC
woodiBrig}rt
P. keeni Blackman BC
P. pini Swaine AK YK NT BC AB MB PQ
alaskanus Blackman
piceae Swaine
P. punctatus LeConte BC
buckhorni Blackman
chamberlini Blackman
haniksu Blackman
rubicun,lulus Swaine
rzsli Blackman
P. scopulorum scopulorum Swaine _BCAB
P. sequoiae llopkins AK- _BC
blackmcLni Schedl
squamo.sus Blackman

Genus CHRAMESUS LeConte

C. hicoriae LeConte oN PQ
lecontei (Chapuis)

Tribe Hypoborini
Genus CHA.ETOPHLOEUS LeConte
Pse ud oc' ryp h alus Swaine
Renocis Casey
C. heterodoxus (Casey) BC AB SK MB
brittaini:. (Swaine)
brunneu.s (Blackman)
commix,ius (Blackman)
criddlei (Swaine)
/asczs (Blackman)

358
 Scoly'tidae

Tbibe Polygraphini
Genus CARPHOBORUS Eichhoff
C. andersoni Swaine AKYKNT_AB
C. carri Swaine AKYKNT_AB MB -NS--
_NB
C. dunni Swaine
C. intermedius Wood AK BC
C. ponderosae Swaine
C. sansoni Swaine - - - :"AB-
engelmanniWood.
C. vandykei Bruck -BC
Genus POLYGRAPHUS Erichson
Lepisotnus IKtrby
P. convexifrons Wood AKYK_BCAB_PQ
P. rufrpennis (Kirby) AK YK NT BC AB SK MB ON PQ NB NS PE LB NF
breuicornis (Kirby)
nigriceps (Kirby)
saginatus Mannerheim

Subfamily SCOLYTINAE
Tribe Scolytini
Genus SCOLYTUS Geoffroy

_ _ _:"_ _ _:x;,_
S. fagi Walsh
S. laricis Blackman
S. mali (Bechstein)t
_
sulcatus LeConte
S. monticolae Swaine -BCAB
S. multistriatus (Marsham)f
S. muticus Say
S. opacus Blackman
abietis Blackman
_ - _:'AB_ I"3NTl"-
S. piceae (Swaine)
S. praeceps LeConte
S. quadrispinosus Say
AK YK NT BC AB SK MB ON PQ
___:"___oNpe__
NB
_:"f"
carya Riley
coryoe Walsh
S. rugulosus (P.W.J. Mtiller)t -BC-oNPQ NS_-_
S. subscaber LeConte _BC
S. tsugae (Swaine) _BCAB
S. unispinosus LeOonte -BCAB
fis&ei Blackman
sobrinus Blackman
S. ventralis LeConte BC

Ttibe Micracini
Genus PSEUDOTHYSANOES Blackman
Chalcohyus Blackman
C ry ptocle ptes Blackman
Cryptulocleptus Wood
P. rigidus (LeConte) _PQ
drakeiBlackrr'an

Genus MICRACIS LeConte

M. swainei Blackman ON
populi Swaine

Genus HYLOCURUS Eichhoff

M icr acisoides Blackman
H. hirtellus LeConte _BC
crinitus Blackman

359
Scolytidae

Tribe Ipini
Genus PITYOGENES Bedel
P. carinulatus (LeConte) BC
hannatus (LeConte)
P. fossifrons (LeConte) BC
P. hopkinsi Swaine - SX l,fg ON pq NB NS NF
P. plagiatus knechteli Swaine _YK BC AB SK
P. plagiatus plagiatus (LeConte) AB SK MB ON PQ
lecontei. Swaine

Genus PITYOKTEINES Fuchs

Orthotomides Wood,
P. elegans Swaine BC AB
P. Iasiocarp'. (Swaine) BC AB
P. minutus rSwaine) BC AB
jasperi Swaine
P. ornatus ( lwaine) BC
P. sparsus (.LeConte) _ABSK MB ON PQ NB NS PE LB NF
balsameus (LeConte)

Genus ORTHOTOMICUS Ferrari

O. caelatus i.EichhofD AK YK NT BC AB SK MB ON PQ NB NS NF
decretus (Eichhoff)
Puncti7'ennis (LeConte)
uicinus (.LeConte)

Genus IPS DeGeer

I. borealis Swaine AI{ YK NT BC AB SK MB ON PQ NB NS NF
swaine. Hopping
thomasi Hopping
I. calligraphus (Germar) SK oN PQ NS
chloroticus Dejean
conforntis Dejean
eresus rlsay)
ponderosae Swaine
praetnorsus (Eichhoffl
I. concinnus (Mannerheim) AK BC
chamberlini (Swaine)
hirsutus (Eichhoff)
I. emarginatus (LeConte) BC
I. grandicollis (Eichhofl) lae oN pe
cacographus (LeConte)
chagno;zi Swaine
cloudcrcfti Swaine
c r i b r icct llis ( Eichhofl)

I. integer (E ichhofl) BC
I. latidens (l.,eConte) YK_ BC AB SK ON PQ
guildilllackrnan
longidens Swaine
I. mexicanuri (Hopkins) AK BC AB
radiatae Hopkins
I. montanus (Eichhof0 BC
uancoutseri Swaine
L perroti Swaine _BCABSKMBONPQNB
I. perturbatus (EichhofD AK YK NT BC AB SK MB ON PQ NB LB
hudsonicus (LeConte)
interp u.nctus (EichhofD
I. pini (Say) AK YK NT BC AB SK MB ON PQ NB NS NF
dentatus (Sturm)
laticolli.s Swaine
oregoni:s (Eichhoff)
pallipes (Sturm)
praefrictus (Eichhoff)
rectus (LeConte')
I. plastographus plastographus BC
(LeConte)
L tridens (Mannerheim) YK NT BC AB
am isk iuL iensis Hopping

360
 Scolytidae

dubius Swaine
engelmanni Swaine
inter r u ptu s (Mannerheim)
semirostris Hopping
yohoensis Swaine
I. woodi Thatcher AB

Tribe Dryocoetini
Genus LYMANTOR Lovendal
L. alaskanus Wood AK
L. decipiens (LeConte) oN pe

Genus DRYOCOETES Eichhoff

D. affaber (Mannerheim) AK YK NT BC SK MB ON PQ NB NS NF
piceoe Hopkins
pubescens Swaine
D. autographus (Ratzeburg) AK YK NT BC AB SK MB ON PQ NB NS LB NF
americanus Hopkins
pseudotsugae Swaine
setn ica steneus (Mannerheim)
sep te nt r ionis ( Mannerheim)
D. betulae Hopkins BC ON PQ NB NS NF
eichhoffi Hopkins
I iq uidatnb ar is Hopkins

D. caryi Hopkins
D. confusus Swaine
abiells Hopkins
AK BC
BC
AB
AB -T
D. granicollis (LeConte) -PQ NB
D. sechelti Swaine BC

Tlibe Crypturgini
Genus CRYPTURGUS Erichson
C. borealis Swaine AK-NTBCABSKMBONPQNBNSNF
corrugatus Swaine
C. pusillus (Gyllenhal)t _SK_ONPQNBNSPE-NF
dtotnus LeConte

Genus DOLURGUS Eichhoff

D. pumilus (Mannerheim) AK BC

Tlibe Xyloterini
Genus XYLOTERINUS Swaine
X. politus (Say) MB ON PQ NB NS
unicolor (Eichhofft

Genus TRYPODENDRON Stephens

T. betulae Swaine NT BC AB -MBONPQNBNSPE
T. lineatum (Olivier) AKYK_BCAB SK MB ON PQ NB NS PE NF
biuittatum (Kirby)
borealis Swaine
c au ifron s (Mannerheim)
uittigerum Eichhoff
T. retusum (LeConte) AKYK-BCABSKMBONPQNB
T. rufrtarsis (Kirby) AKYK-BCAB-MBONPQNBNS
ponderosae Swaine

Tlibe Xyleborini
Genus XYLOSANDRUS Reitter
X. germanus (Blandford) ON

361
Scolytidae

Genus XYLI.BORUS Eichhoff

Ambros;odmus Hopkins
Anisantl.rus Ferrari
X. affinis Eichhoff - ON
X. dispar (Fabricius) - BC - ON PQ - NS NF
pyri (Peck)
swainei (Drake)
X. intrusus Blandford - BC
futcfri Hopkins
howardt Hopkins
scopulorum Hopkins
X. obesus Letlonte - ON PQ
populi (Swarne)
serratus Swaine
X. pubescens Zimmermann - ON
propinguus Eichhoff
X. sal (Hopl:inst - ON PQ
minor Swaine
neardus Schedl
X. xylographrs (Sayt - ON PQ
canadensis Swaine
inermis Eichhoff

Genus XYLEBORINUS Reitter

X. saxeseni { latzeburg) - BC - ON PQ
orbufi (llopkins)
floridem;is (Hopkins)
Iibocedr,:. Swaine
pecanus (Hopkins)
quercus (Hopkins)
lszgoe Swaine

Tlibe Cryphalini
Genus TRYPOPHLOEUS Fairmaire
T. populi Hopkins - BC AB SK MB NB
T. striatulus (Mannerheim) AK YK AB - MB - PQ - NS NF
nitidus fiwaine
punctipennis Hopkins
T. thatcheri (Woodt - BC

Genus PROCRYPHAIUS Hopkins

P. mucronatrLs (LeContet AX BC AB
idahoensis Hopkins
populi L{opkins
P. utahensis Hopkins AK BC AB - PQ
oceris Hopkins
solicis F.opkins

Genus CRYPHALUS Erichson

Taeniogi),yptes Bedel
C. pubescens Hopkins - BC
subconcentralis Hopkins
C. ruficollis f-aseri Hopkins - - PQ
balsameus Hopkins
C. rufrcollis ruficollis Hopkins AK YK - BC AB - MB ON PQ NB NS PE - NF
amabilis Chamberiin
approximatus Hopkins
canadensis Chamberlin
coloradensis (Wood)
grandis Chamberlin
mainensis Blackman

Genus HYPGTHENEMUS Westwood

Stephanoderes Eichhoff
H. dissimilis (Zimmermann) - ON PQ
chapuisti (Eichhoff)

362
 Scolytidae

H. eruditus Westwood - ON
asiminae Hopkins
citri Ebling
euonymi Hopkins
hamamelidis Hopkins
juglandis Blackman
nigripennis Hopkins
przni Hopkins
punctifrons Hopkins
rumseyi Hopkins
schwarzi (Hopkins)

Tlibe Corthylini
Genus PSEUDOPITYOPHTHORUS Swaine
P. minutissimus (Zimmermann)
P. pubipennis ( LeConte) sc _9* PQ

Genus CONOPHTHORUS Hopkins

C. banksianae McPherson -oN
C. coniperda (Schwarz) -oN PQ NB NS
clunicus Hopkins
taedae Hopkins
C. ponderosae Hopkins BC
contortae Hopkins
tn
flerilis Hopkins
lambertianae Hopkins
monticolae Hopkins
scopulorum Hopkins
C. resinosae Hopkins oN PQ NB NS
uirginianae Hopkins

Genus PITYOPHTHORUS Eichhoff

Key to North American species: Brieht (1981)
(Subgenus PITYOPHTHORUS s. str.)
(Myeleborus Blackman)
P. absonus Blackman _BC AB
demissus Blackman
inyoensis Big}rt
P. alpinensis Hopping AB
P. angustus Blackman __ON_NENS
P. aquilus Blackman ___sc AB SK
aristatae Bright
caelator Blackman
P. balsaneus Blackman -NT ON PQ NB NS
patchiBlackman
P. bassetti Blackman AK YK NT
P. biovalis Blackman -ONNS
P. boycei Swaine \tK BC AB
catulus (Blackman)
iniquus (Blackman)
siouxensis Bight
P. briscoei Blackman -oN PQ NB
mundus Blackman
P. carinatus carinatus Bright PQ NB
P. carinatus monticolae Bright sc AB_
P. cariniceps LeConte ? AB SK ff,fe O* PQ NB NS
canad,ensis Swaine
cognatus Blackman
P. carinulatus Swaine AK
opimus Blackman
P. cascoensis Blackman NT -AB ON NF
pilifer Schedl
P. cavatus Bright SK MB
P. concavus Blackman oN PQ NB NS
P. confertus Swaine BC
agndtus Blackman
burkeiBlackman
com.DtusBlackman

363
Scolytidae

P. confinis LeConte - BC
P. consimilis LeConte - BC AB SK MB ON PQ - NS
granulatus Swaine
nudus Siwaine
P. dentifrons Blackman AB ON PQ NB NS PE - NF
P. digestus ( LeContet - BC AB
aplanatus Schedl
hophinsi Blackman
idoneus Blackman
ponderosae Blackman
P. fuscus Bla.ckman - BC
smithi Schedl
P. grandis B'ackman - BC
P. hesperius Bright - BC
P. intextus Swaine - BC AB SK MB ON PQ NB NS NF
shepardi Blackman
lonsus lJlackman
P. lautus Eichhoff - ON PQ
natalis Blackman
rhois Svraine
P. murrayanae Biackman AK - NT BC AB - MB ON - NB
acceptut: Bight
aurulentus Bight
cutleri S,waine
depygis Blackman
elongatt.!.s Swaine
e:rilis Swaine
gracilis Swaine
tenuis Swaine
watsoni Schedl
P. nitidulus tMannerheim) AK BC AB
atratulus (LeConte)
punctict. llis (LeConte)
P. nitidus Svvaine AK YK NT BC AB ON PQ NB NS NF
anceps Illackman
aquilon,:us Bight
borealis Swaine
uorions Schedl
P. opaculus L,eConte AK YK NT BC AB SK MB ON PQ NB NS NF
abietis Eilacknan
albertensis Blackman
exiguus Blackman
pygmaetts Schedl
P. orariusBright - BC
P. pseudotsugae Swaine - BC
thatcheriBight
P. pubemlus (LeConte) - ON PQ NB NS NF
infans Eichhoff
P. pulchellus Eichhoff - YK NT BC AB SK MB ON PQ NB
hirticept, LeConte
pusio LeConte
P. pulicarius (Zimmermann) - SK MB ON PQ NB NS
P. ramiperda Swaine - ON PQ
fiuazi (Blackman)
P. recens Bri.;ht - YK - BC
P. scalptor Blackman - BC
P. scalptus Bright - BC
P. serratus Swaine - BC
P. sierrensis Bright - BC
P. toralis Wo,rd - BC AB
collinus Bng}rt
confusus (Bright)
P. tuberculatus Eichhoff AK YK - BC AB
rugicollis Swaine
P. venustus Rlackman - BC
artifexBlackman

Genus GNAT'HOTRICHUS Eichhoff

Ancyloderes Blackman
Gnathot,"ic hoides Blackman

364
 Platypodidae

G. materiarius (Fitch) - SK MB ON PQ NB NS
corthyloides Eichhoff
G. retusus (LeConte) - BC
alni Blackman
G. sulcatus (LeContet - BC
aciculatus Blackman

Genus MONARTHRUM Kirsch

M. fasciatum (Say) - ON PQ
gracile (Eichhoff)
simile (Eichhofl\
M. mali (Firch) - oN PQ NB
longulum Eichhoff
M. scutellare (LeConte) - BC
cauum (LeConte)
obl iq uecaudatum ( Schedl)

Genus CORTF{YLUS Erichson

C. punctatissimus (Zimmermann) - ON PQ

FamilyPLAIYPODIDAE pinholeborers
J. McNamara

One species of Platypodidae occurs in Canada. Wood (1979) has published a catalog of the
These beetles infest the wood of weakened trees, North American species.
where they mine galleries. The larvae feed on the
fungus that grows on the gallery walls. BC (1)

Subfamily PLATYPODINAE
TYibe Platypodini
Genus PLATYPUS Herbst
P. wilsoni Swaine - BC

365
Halictophagidae

Order Strepsiptera twisted-winged parasites

S.B. Peclt

In Canada 11 species of Strepsiptera are now Adults may be reared from parasitized hosts.
known in three of the nine families recognized Males may be attracted to ultraviolet black light
around the world. These species are distributed in traps or may be lured by placing virgin females (in
Southern Canada. Another 10 species occur in the their hosts) in frnely screened cages.
northern tier of the United States and may eventu- The systematic position of this group is becom-
ally be found in Canada. Many taxa are known ing clear. It has been grouped near the similarly
only throrrgh adult females. endoparasitic Rhipiphoridae in general works on
The biology of this endoparasitic group is highly beetles (Arnett 1968, Crowson 1981, White 1983).
unusual. Studies on species similar to those in Extensive morphological studies by Kinzelbach
Canada suggest the following. The frrst-instar tri- (1971) and Kathirithamby (1989) suggest that
ungulin-like larvae are free-living at birth but strepsipterans are the sister group to the beetles as
immedial;ely seek and parasitize immature speci- a whole. They are then ranked as a separate order
mens of lvgaeid bugs, various leafhoppers and their most clearly related to the beetles (Kinzelbach 1971,
relatives, and Andreno solitary bees. The hetero- Lawrence and Newton 1982, Kathirithamby 1989).
morphic larvae mature in the host insect and Overviews of the group in North Arrrerica are
pupate there. Adult female strepsipterans remain found in Bohart (1941), Kinzelbach (1$71), and
larva-like and in the host. Adult males are winged Luna de Carvalho and Kogan (1990). The classifica-
and live r)nly a day and an evening or two, during tion used here follows Kathirithamby (1980).
which tinre they actively search for females. After
the female has been inseminated, eggs develop and
hatch inside her. A thousand or more larvae may
then leave the female, to seek new hosts and begin
the cycle again. BC (2);AB (1); SK (3); ON (2);PQ (2);NS (1)

Suborder Stylopidia
Family CORIOXENIDAE
Subfamily CORIOXENINAE
Genus LOANIA Kinzelbach
L. canadensis Kinzelbach ON

Family HALICTOPHAGIDAE
Subfamily IIALICTOPHAGINAE
Genus HALICTOPHAGUS Bohart
Agalliaphagus Pterce
Anthericomma Pierce
Bruesia Perkins
Cyrtocaraxenos Pierce
Dacyrtocara Pietce
Dioxocera Pietce
Diozocera Pierce
Megalechthrus Perkins
Membracixenos Pierce
NeocholaxPierce
Pentac{.adocera Pierce
Pe n log'qm mqPft i1a Pierce
Pentaxocera Pierce
Pentazocera Pietce
Pentozoe Pierce
Pseudopatella Bohaft
Pyrilloxenos Pierce

366
 Stylopidae

Key to North American species: Bohart (1943)

H. americanus Perkins PQ
H. bidentatus Bohart _SK
H. mackal Bohart _SK
H. omani Bohart AB

Genus STENOCRANOPHILUS Pierce

Delphacircnos Pierce
Muirixenos Pierce
S. canadensis Kinzelbach SK

Family STYLOPIDAE
Subfamily STYLOPINAE
Genus STYLOPS Kirby
Katastylops Pierce
Neostylops Pierce
Prostylops Pierce
S. advarians Pierce -BC
S. childreni Gray NS
S. leechi Bohart _BC
S. vicinae Pierce
prf

Subfamily XENINAE
Genus PSEUDOXENOS Saunders
P. tigridis Pierce ON

367
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396
 INDEX to supraspecific taxa
(Synonyms are in italics.)

Abacidus LeConte 33 Acupalpus Latreille 47

Abax Bonelli 35 Acylamus Sharp 226
Abdera Stephens 248 Acylophorus Nordmann 122
Abletobium Casey 115 Adalia Mulsant 235
Abraeinae 135 Adelina Dejean 255
Abraeini 135 Adephaga 7
Abraeus Leach 135 Aderidae 275
Abstrulia Casey 243 Aderocharis Sharp 113
Acalles Schiinherr 347 Aderus Westwood 275
Acalligrapha Monros 310 Adimonia Laicharting 315
Acallodes LeConte 350 Adistemia Fall 240
Acalymma Barber 315 Adota Casey 101
Acalyptus Schtjnherr 341 Adoxini 309
Acamptini 355 Adoxus Kirby 309
Acamptus LeConte 355 Adranes LeConte 129
Acanthocinini 296 Adranini 129
Acanthoderini 296 Aegialia Latreille 147
Acanthoscelides Schilsky 301 Aegialiini 146
Acanthoscelidius Dalla Torre & Hustache 349 Aegialites Dejean 270
Acanthoscelis Dietz 349 Aegialitinae 270
Achenomorphus Motschulsky 113 Aegomorphus Haldeman 296
Acholerops Casey 83 Aeletes Horn 136
Acicnemidinae 338 Aeolus Eschscholtz 176
Acidota Stephens 89 Aethiopoctines Thomson 296
Aciliini 71 Agabetes Crotch 63
Acilius Leach 71 Agabetinae 63
Acletus LeConte 212 Agabetini 63
Aclypea Reitter 85 Agabini 67
Acmaeodera Eschscholtz l.6l- Agabinus Crotch 67
Acmaeoderinae 161 Agabus Leach 67
Acmaeoderini 161 Agalliaphagus Pierce 366
Acmaeops LeConte 289 Agaporus Zimmermann 67
Acmaeopsilla Casey 290 Agasphaerops Horn 332
Acmaeopsoides Linsley & Chemsak 293 Agathengis des Gozis 223
Acoptus LeConte 348 Agathidini 80
Acrepis LeConte 200 Agathidium Panzer 80
Acritini 136 Agleninae 270
Acritus LeConte 136 Aglenus Erichson 270
Acrolochct Thomson 88 Aglyptinus Cockerell 79
Acromacer Kuschel 324 Agonoderus Dejean 47
Acropteroxys Gorham 224 Agonodromius Reitter 52
Acroschatia Wilke 254 Agonum Bonelli 52,53
Acrotona Thomson 100 Agostenus MotschulskY 50
Acrotrichiinae 76 Agrabia Casey 236
Acrotrichis Motschulsky 76 Agrilaxia Kerremans 165
Acrulia Thomson 88 Agrilinae 167
Actedium Motschulsky 21 Agrilini 167
Actiastes Casey 127 Agrilus Curtis 167
Actidium Matthews 75 Agriotella Brown 182
Actium Casey 127 Agriotes Eschscholtz 182
Actizona Chandler 127 Agriotini 182
Actobius Fauvel 117 Agroiconota Spaeth 323

397
Agronus Horn 332 Ampeloglypter LeConte 352
Agyrtecanus Reitter 77 Amphasia Newman 45
Agyrtes Itrdlich 77 Amphicerus LeConte 200
Agyrtidae 77 Amphichroum Kraatz 90
Ahasverus des Gozis 220 Amphicrossus Erichson 216
Airora Reitter 207 Amphidora Eschscholtz 257
Akephoms LeConte 17 Amphimallon Berthold 153
Alamona Casey 155 Amphivectura Blaisdell 212
Alaobia Thomson 102 Amphizoa LeConte 61
Alaus Eschscholtz 175 Amphizoidae 61
Aleocharil Gravenhorst I04 Amydetinae 191
Aleocharinae 99 Amydrogmus Pierce 334
Aleocharini 104 Anabolus Gistel 12
Aleodoru,s Say 106 Anacaena Thomson 132
Aletinus Desbrochers 327 Anacentrinus Buchanan 353
Alitargus Casey 242 Anadaptus Casey 43
Allandrini 325 Anaedus Blanchard 262
Allandrus LeConte 325 Anaglyptini 286
Alleculinne 259 Anagrylius Casey 156
Alleculinii 259 Analcis Schdnherr 347
Alloeocnemis LeConte 199 Analeptura Linsley & Chemsak 291
Allonychus Zaitzev 67 Anambodera Barr 161
AllopodaLeConte 276 Anametis Horn 334
Allopodini 276 Anaquedius Casey 122
AllorhagtumKolbe 289 Anaspidinae 276
Allosirocrilus Colonnelli 350 Anaspidini 276
Alniphagus Swaine 357 Anaspis Geoffroy 276
Alobates Motschulsky 26L Anqstictodera Casey l2I
Alobus LeConte 151 Anastrangalia Casey 293
Alocentron Schilsky 328 Anatis Mulsant 235
Alphitobius Stephens 258 Anatrinodia Casey 157
Alphitophagus Stephens 255 Anchastus LeConte 184
Altica Mtiller 317 Anchicera Thomson 223
Alticinae 316 Anchodemus LeConte 335
Alyca Le(lonte 345 Anchomenus Bonelli 52
Alycodes Dietz 339 Aruchus LeConte 52
Amacylophorus Smetana 122 Anchytarsus Gu6rin-M6neville 169
Amalus Schdnherr 350 Ancistronycha Markel 193
Amara Bonelli 35, 38 Ancylochira Eschscholtz 163
Amarygrrrini 259 Ancyloderes Blackman 364
Amqurosromo Mtiller 299 Ancylonycha Blanchard 152
Amblopusa Casey 107 Ancyronychini I72
Amblycerinae 300 Ancyronyx Erichson L72
Amblycerus Thunberg 300 Ancy rop horus I{r aatz 94
Amblyctis LeConte 247 Andrector Horn 315
Amblyderus LaFert6-S lnecldre 27 | Androchirus LeConte 260
Ambrosiodmus Hopkins 362 Anelaphus Linsley 281
Americanotimarcha Jolivet 310 Anelastes Kirby 188
Americonr,asezs Csiki 34 Anelpistus Horn 263
Amerinusr Casey 46 Anepsiini 253
Amerizus Chaudoir 28 Anepsius LeConte 253
Amecocerus Solier 213 Anepsiota Casey 101
Ametor Semenov 132 Anisa.ndrus Ferrari 362
Amischa Thomson 101 Anisocaluio Crotch 235
Arnnesia.Florn 333 Anisodactylus Dejean 43, 44
Amniscus, Haldeman 297 Anisomera LeConte 67
Ampedini 183 Anisorus Mulsant 288
Ampedus Dejean 183 Anisostena Weise 321

398
Anisosticta Dejean 233 Aphanotus LeConte 258
Anisotarsus Chaudoir 45 Aphelogenio Chaudoir 56
Anisotoma Panzer 80 Apheloplastus Brown 79
Anisoxya Mulsant 248 Aphenolia Reitter 215
Anobiidae 201 Aphodiinae 146
Anobiinae 202 Aphodiini 147
Anobiini 203 Aphodius Illiger 147
Anobium Fabricius 203 Aphonus LeConte 156
Anocomis Casey 283 Aphorista Gorham 238
Anogcodes Dejean 266 Aphrastus Say 333
Anogdus LeConte 78 Aphthona Chevrolat 317
Anomala Samouelle 154 Apion Herbst 328
Anomalaegialia Brown 147 Apionidae 328
Anomalepta Casey 154 Apionini 328
Anomalini 154 Aplemonini 329
Anomalonychus Saylor 264 Aplocentrus LeConte 44
Anomalopus Casey 154 Aploderus Stephens 93
Anomoea Lacordaire 304 Apocellus Erichson 95
Anomoglossus Chaudoir 49 Apodasyini 295
Anomoia Dejean 304 Apothinus Sharp 126
Anomophagus Reitter 218 Apotomus Kirby 326
Anoplis Kirby 163 Apristus Chaudoir 58
Anoplitis Chapuis 321 Apteralium Casey 115
Anoplitis Kirby 321 Apterius Blatchley 317
AnopliurnHaldeman 280 Apteroloma Hatch 77
Anoplotrupes Jekel 150 Apteromechus Faust 348
Anotylus Thomson 95 Apterosoma Motschulsky 311
Antennalia Casey 278 Apterospasta LeConte 265
Anthaxia Eschscholtz 165 Araecerini 324
Anthaxiini 165 Araecerus Schtinherr 324
Anthericomma Pierce 366 Araeopidius Cockerell 143
Antherophagus Latreille 221 Araeopus LeConte 143
Anthicidae 271 Archostemata 6
Anthicinae 271 Arctelaphrus Semenov 16
Anthicini 271 ArctobiaThomson 15
Anthicus Paykull 271 Arctoclinidium Bell 7
Anthobates LeConte 276 Arctodytes Thomson 67
Anthobatulo Strand 276 Arctolina Kontkanen 312
Anthobium Leach 91 Areus Casey 109
Anthobium auct. 87 Argaleus LeConte 288
Anthocornus Erichson 211 Argante Gistel 162
Anthomarp,hus Weise 343 Argutor Dejean 34
Anthonominae 343 Arhopalus Audinet-Serville 279
Anthonomini 343 Aridius Motschulsky 240
Anthonomopsis Dietz 343 Arpedium Erichson 89
Anthonomus Germar 343 Arrhenodes Schiinherr 327
Anthophagini 89 Arrhenodini 327
Anthophylax LeConte 289 Arrhenopeplus Kocb. 84
Anthrenini 198 Arrhenoplita l{trby 255
Anthrenus Schaeffer 198 Artematopidae 174
Anthribidae 324 Arthrolips Wollaston 228
Anthribinae 324 Arthromacral{trby 262
Anthribini 324 Artochia Casey 90
Anthribulus LeConte 324 AsaphesKirby 181
Apator Semenov 67 Asaphidion des Gozis 20
Apatura Castelnau & Gory 165 Asclera Stephens 266
Apelocera Chevrolat 286 Asclerini 266
Apenes LeConte 59 Ascoliocerus M6quignon 178

399
Aseminae 279 Baeckmanniolus Reichardt 137
Asemini 279 Baeocera Erichson 125
Asemobius Horn 95 Bagoini 341
Asemum Eschscholtz 279 Bagous Germar 341
Asidini Il54 Balaninus Germar 342
Asidopsis Casey 254 Baliosus Weise 321
Asiobates Thomson 74 BalophilaCrotch 317
Aspidius Mulsant & Rey 257 Baridinae 351
Astenina 117 Baridini 351
Astenus Dejean 117 Baridius Schtinherr 351
Asteria Fauvel 113 Baris Germar 351
Astyleiopus Dillon 297 Barynotini 330
Astylopsis Casey 297 Barynotus Germar 330
Asynonyt:hus Crotch 329 Barypeithes Jacquelin du Val & Fairmaire 330
Ataenius Harold 149 Barytachys Chaudoir 28
Atheta Thomson 101 Bassareus Haldeman 307
Athetalia Casey 101 Batrisinae 127
Athetini 100 Batrisini 127
Athetota Casey 101 Batrisodes Reitter 127
Athoini .176 Batyle Thomson 287
Atholus llhomson 139 Batyleoma Casey 287
Athous Eschscholtz 177 Baudia Ragusa 49
Atimia Haldeman 280 Beeria Hatch 120
Atimiini 280 Bellamira LeConte 290
Atomaria Stephens 223 Belonuchus Nordmann 119
Atomariirae 223 Bembidiini 20
Atomarii:ri 223 Bembidion Latreille 20, 27
Atrachela. Gistel 308 Bembidionetolitzkya Strand 23
Atractopterus LeConte 182 Berninelsonius Leseigneur 178
Atranus l,eConte 52 Berosinae 131
Atrecus Jacquelin du Val 123 Berosus Leach 131
Attagenirri 197 Bessobia Thomson 102
Attagenus Latreille 197 Bibloplectus Reitter 127
Attalus Erichson 212 Bidensomela Monros 310
Attelabidae 326 Bidessini 63
Attelabini 326 Bidessonotus R6gimbart 63
Attelabus;Linn6 326 Biphyllidae 241
Atymius (]istel 309 Biphyllini 241
Auchenia Thunberg 304 Bitoma Herbst 251
Augyliini 169 Bius Mulsant 258
AulaconArnett 175 Blapstinus Dejean 257
Auletini 327 Blapylis Horn 257
Auletobiu.s Desbrochers 327 Blechrus Motschulskv 58
Auleutes Dietz 349 Bledius Leach 93
Aulobaris LeConte 352 Blemus Dejean 19
Aulonium Erichson 251 Blepharida Chewolat 316
Aulonothr:oscus Horn 186 Blepharidulo Weise 316
Autalia Leach 107 Blepharoplataphus Netolitzky 23
Autaliini 107 Blethisa Bonelli 15
Axinopalpus LeConte 58 Blitophaga Reitter 85
Bolboceras Kirby 150
Babia Chevrolat 305 Bolboceratini 150
Babiini 305 Bolbocerosoma Schaeffer 150
Babnormrrdes Park 127 Boliographa Motschulsky 310
Bacaniini 138 Bolitobiini 98
Bacanius LeConte 138 Bolitobius Leach 98
Bactridium LeConte 218 Bolitobus Tottenham 98
Badister Clairville 49 Bolitochara Mannerheim 107

400
Bolitocharina I07 Brennus Motschulsky 13
Bolitophaginl 255 Brenthidae 327
Bolitophagus Illiger 255 Bromius Chevrolat 309
Bolitotherus Candbze 255 Brontes Fabricius 220
Bomius LeConte 58 Broscini 19
Boreades Parsons 214 Broscodera Lindroth 19
Boreaphilus Sahlberg 89 Broscus Panzer 19
BoreobiaTschitsch6rine 31 Bruchidae 300
Boreocanthon Halffter 146 Bruchidius Schilsky 301
Boreoglymmius Bell & BeIl 7 Bruchinae 300
Boreophilia Benick 102 Bruchus Linn6 301
Boreosaxinis Moldenke 305 Bruesia Perkins 366
Boreostiba Lohse 101 Brumoides Chapin 233
Boridae 268 Bryaxis Leach 128
Borophloeus Wollaston 355 Brychius Thomson 60
Boros Herbst 268 Bryonomus Casey 119
Boschella Barr 210 Bryoporus lkaatz 98
Bostrichidae 199 Buprestidae 160
Bostrichinae 200 Buprestinae 162
Bostrichini 200 Buprestini 163
Bostrichoidea 199 Buprestis Linn6 163
Bothrideres Erichson 252 Buprestoidea 160
Bothriderinae 252 Byrrhidae 158
Bothriderini 252 Byrrhinae 158
Bothriopterus Chaudoir 32 Byrrhini 158
Brachiacantha Dejean 232 Byrrhodes LeConte 205
Brachinini 18 Byrrhoidea 158
Brachinus Weber 18 Byrrhus Linn6 158
Brachybamus Germar 341 Byrsopages Schijmherr 335
Brachycepsis Brendel 83 Bythinini 128
Brachycis Casey 245 Bythinopsis Raffray 128
Brachycoryna Gu6rin-M6neville 322 Byturidae 241
Brachycrepis LeConte 184 Byturini 241
Brachyderinae 329 Byturus Latreille 241
Brachyderini 330
Brachygluta Thomson 128 CacopliaLeConte 294
Brachyglutini 128 Cacoternnus LeConte 203
Brachyini 168 Cq.docera Eschscholtz 149
Brachyleptura Casey 293 Caenia Newman 189
Brachylobus Chaudoir 50 Caeniella Cockerell 189
Brachymorphzs Chewolat 210 Caenocara Thomson 204
Brachynotus Kirby 192 Caenocyrta Brown 79
Brachypnoeo Gistel 308 Caenoscelini 222
Brachypterolus Grouvelle 214 Caenoscelis Thomson 222
Brachypterus Kugelann 214 Caerosternus LeConte 138
Brachyrhinus Latreille 331 Cafius Stephens 119
Brachys Solier 168 Calandra Clairville 354
Brachysomida Casey 290 Calandrinus LeConte 353
Brachysomus Schijnherr 330 Calathus Bonelli 51
Brachystilus Chaudoir 31 Calitys Thomson 207
Brachytarsoides Pierce 324 Calleida Dejean 58
Brachyusa Mulsant & Rey 100 Callidiini 282
Bracteomimus Lindroth 21 Callidium Mulsant 283
Bracteon Bedel 20 Calligrapha Chewolat 310
Bradybatini 344 Callimellum Strand 282
Bradycellus Erichson 46 Callimoxys lkaatz 282
Bradytus Stephens 36 Callimus Mulsant 282
Brathinus LeConte 89 Callipara Motschulsky 10

401
Callisten,',a Vacher de Lapouge 11 Catharellus Casey 46
Callisthenes Fischer von Waldheim 11 Cathartosilvanus Grouvelle 220
Callisthiri 49 Catogenus Westwood 219
Callitrop;r Motschulsky 10 Catopocerinae 80
Calloides LeConte 285 Catopocerus Motschulsky 80
Callosobruchus Pic 300 Catops Paykull 81
Calochara Casey 104 Catoptrichus Murray 81
Calochromus Gu6rin-M6neville 188 Catorama Gu6rin-M6neville 205
Caloderma Casey 113 Cedius LeConte 129
Calodrepa Motschulsky 10 Celetes Newman 189
Calomycterus Roelofs 334 CeliaZimmermann 36
Calopodinae 266 Celiamorphus Casey 43
Calopterini 189 Celina Aub6 63
Calopteron Gu6rin-M6neville 189 Centrinapsis Casey 353
Calopus Irabricius 266 Centrinini 353
Calosirus Thomson 350 Centrinogyna Casey 353
Calosoma Weber 10 Centrodera LeConte 288
Calvia Mulsant 235 Centronopus Solier 258
Calyrnmaderini 204 Centuriatus Pacheco 169
Calymmaderus Solier 204 Ceophyllini 129
Calyptomerus Redtenbacher 141 Ceophyllus LeConte 129
Camptorl;.ino Kirby 151 Cephalogyna Casey 44
Campylir.hynchus Dejean 351 Cephaloidae 262
Canastottr Casey 100 Cephaloinae 262
Canifa LeConte 276 Cephaloon Newman 262
Canonura Casey 296 Ceracis Melli6 245
Cantharidae 192 Ceralimnius Hinton 173
Cantharinae 192 Cerambycidae 277
Cantharirri 193 Cerambycinae 280
Cantharis Linn6 193 CerandriaLucas 255
Cantharoidea 188 Cerataltica Crotch 317
Canthon .lloffmansegg L46 Ceratapion Schilsky 328
Canthosttthus Haldeman 306 Ceratoderus Blanchard 271
Caphalistes Motschulsky 212 Ceratographis Gahan 297
Capnochroa LeConte 260 CeratomaDozier 315
Capraita Bechyn6 320 Ceratomegilla Crotch 233
Carabidae 8 Ceratopidonia Linsley & Chemsak 293
Carabini 10 Cercolia Casey 293
Carabus Linn6 11 Cercopedius Sleeper 332
Carcinops Marseul 138 Cercyon Leach 134
Carcinus.Marseul 138 Cercyonini 134
Cardiophorinae 185 Cerophorus Castelnau 216
Cardiophorus Eschscholtz L85 CerotaruaBowditch 315
Carebq,ra LeConte 247 Cerotoma Chewolat 315
Carpelimus Leach 94 Ceruchini 144
Carphacis des Gozis 98 Ceruchus MacLeay 144
Carphoborus Eichhoff 359 Cerylon Latreille 227
Carphonobus Casey 356 Cerylonidae 227
Carpophilinae 2L4 Ceryloninae 227
Carpophilus Stephens 214 Cetoniinae 157
Carrhydms Fall 69 Cetoniini 157
Cartodere Thomson 240 Ceutorhynchidius Jacquelin du Val 349
Cassida Linn6 322 Ceutorhynchinae 349
Cassidinae 322 Ceutorhynchini 349
Cassidini 322 Ceutorhynchus Germar 349
Catapastus Casey 353 Chaerocassis Spaeth 323
Cateretes Herbst 214 Chaetarthria Stephens 131
Cateretinae 214 Chaetarthriinae 131

402
Chq.etechus Horn 332 Cicindelidia Rivalier 10
Chaetocnema Stephens 319 Cicindelini 8
Chaetophloeus LeConte 358 Cidnorhinus Thomson 350
Chaetophora Stephens 158 Clidae 244
Chalcoidea Motschulsky 311 Ciinae 244
Chalcoparia Crotch 308 Ciini 244
Chalcophora Solier 161 Cilea Jacquelin du Val 97
Chalcophorinae 161 Cimberidini 324
Chalcophorini 161 Cimberis des Gozis 324
Chalcohyus Blackman 359 Cimbocera Horn 333
Chalepini 321 Ciocotalpa Saylor 155
Chalepus Thunberg 321 Circinalia Casey 54
Chapmania Bernhauer 95 CircinalidiaCasey 54
Charaphloeus Casey 219 Circinoporus Guignot 65
Charhyphus Sharp 95 Circomus Fleischer 258
Charidotella Weise 323 Cis Latreille 244
Chariessa Perty 210 Fabricius 259
Cistelct.
Charisalia Casey 291 Clambidae 141
Chauliognathinae 195 Clambus Fischer von Waldheim 141
Chauliognathini 195 Clamoris des Gozis 253
Chauliognathus Hentz 195 Clavigerinae I29
Chelonychus Dietz 343 Clavilispinus Bernhauer 92
Chelymorpha Chewolat 322 Cleonidius Casey 336
Chilocorinae 233 Cleoninae 336
Chilocorini 233 Cleonini 336
Chilocorus Leach 233 Cleonis Dejean 336
ChionNewman 286 Cleonus Schdnherr 336
Chionotyphlus Smetana 113 Cleopomiarus Pierce 347
Chirida Chapuis 323 Cleptelmis Sanderson 172
Chitalia Sharp 106 Cleridae 208
Chlaeniellus Reitter 50 Clerinae 208
Chlaenius Bonelli 49, 50 Clerini 209
Chlamisinae 307 Cleroidea 206
ChloephagaWeise 320 Clinidium Kirby 7
Cholevinae 81 Clivina Latreille 18
Cholevini 81 Clivinini 17
Choraginae 324 Clusiota Casey 102
Choragini 324 Clytini 284
Chramesus LeConte 358 Clytoleptus Casey 286
Chromatia LeConte 259 Clytrinae 304
Chrysobothrinae 166 Clytrini 304
Chrysobothrini 166 Clytus Laicharting 285
Chrysobothris Eschscholtz 166 Cnernidotus Erichson 61
Chrysobracteon Netolitzky 20 Cnemocyllus Dietz 343
Chrysochus Chevrolat 309 Cnemogonini 349
Chrysodina Baly 308 Cnemogonus LeConte 349
Chrysolina Motschulsky 311 Cneoglossini 143
Chrysomela auct. 311 Cneorhinini 330
Chrysomela Linn6 313 Coccidophilus Brethes 229
Chrysomelidae 301 Coccidula Kugelann 233
Chrysomelinae 310 Coccidulinae 233
Chrysomelini 310 Coccidulini 233
Chrysomeloidea 277 Coccinella Linn6 235
Chrysomelopsilo Strand 313 Coccinellidae 229
Chrysomelopsis Achard 313 Coccinellinae 233
Chrysophana LeConte 161 Coccinellini 233
Chrysostigma Kirby 10 Coccotorus LeConte 344
Cicindela Linn6 8 Coelambus Thomson 64

403
Coelini 254 Coptotomini 70
Coelocephalapion Wagner 328 Coptotomus Say 70
Coelocnernis Mannerheim 261 Copturodes Casey 348
Coelogaster Schiinherr 349 Coraebini 167
Coelometopinae 260 Cordalia Jacobs 105
Coelometopini 260 Cordoharpalus Hatch 41
Coelomorpha Casey 254 Coreopsomela Monros 310
Coelostethus LeConte 203 Corioxenidae 366
Coelotaxis Horn 253 Corioxeninae 366
Coelus Eschscholtz 254 CorphyraSay 274
Colaspis Fabricius 308 Corthylini 363
Colastus Erichson 214 Corthylus Erichson 365
Colenis Erichson 79 Corticaria Marsham 240
Coleomegilla Timberlake 233 Corticariinae 240
Coleothorpa Moldenke 305 Corticarina Reitter 241
Collapterl,'x Newman 294 Corticeus Piller & Mitterpacher 256
Colliuris DeGeer 56 Corticotomus Sharp 207
Collops Erichson 211 Cortilena Motschulsky 241
Colon Her:bst 80 Cortinicara Johnson 241
Coloninaer 80 Cortodera Mulsant 290
Colopterus Erichson 214 Corylophidae 228
Colposura', Casey 101 Corylophinae 228
Colydiida,: 251 Corymbites Latreille 179
Colydiinae 251 Coryphiini 89
Colydiini 251 Coryphium Stephens 89
Colydium Fabricius 252 Cosmobaris Casey 352
Colymbet,:s Clairville 69 Cosmosalia Casey 294
Colymbet.nae 67 Cosnania Dejean 56
Colymbet:ni 69 Cossoninae 355
Comstocle,'.a Van Dyke 52 Cossonini 355
Conalia lVlulsant &Rey 249 Cossonus Clairville 355
Coninomu'.s Thomson 240 Cotalpa Burmeister 155
Coniocleonus Motschulsky 336 Coxelus Horn 251
ConiontelLus Casey 253 Craponius LeConte 349
Coniontides Casey 253 Cratacanthus Dejean 39
Coniontini 253 Cratidus LeConte 257
Coniontis Eschscholtz 253 Cratomorphini 190
Coniophagus Mink 213 Cratonychus Dejean 184
Kangas 221
Con iop hth:.alma Crerya LeConte 210
Conithassa Thomson 239 Cremastocheilini 157
Connatichela Anderson 334 Cremastocheilus Knoch 157
Connophron Casey 82 Creniphilus Horn 132
Conoderus Eschscholtz 176 Crenitis Bedel 132
Conophthorus Hopkins 363 CrenitulusWinters 132
Conoplectus Brendel 126 Creophilus Samouelle 120
Conotelus Erichson 214 Crepidodera Chevrolat 319
Conotrachelus Dejean 347 Crepidophorus Mulsant & Guillebean 177
Copelatinae 62 Crepidotritus LeConte 184
Copelatini 62 Crine Pascoe 218
Copelatus Erichson 62 Criocephalum Dejean 27 I
Cophes Champion 348 Criocephalus Mulsant 279
Copidita LeConte 266 Criocerinae 304
Coprini 1,46 Criocerini 304
Copris Mtiller 146 Crioceris Mi.iller 304
Coprochara Mulsant & Rey 104 CrioscaphaHorn 246
Coprophibrni 92 Crosimus Casey 221
Coprophilus Latreille 93 Crossidius LeConte 287
Coproporus Kraatz gG Cryrnodes LeConte 268

404
Cryobius Chaudoir 34 Cychrini 12
Cryocarabus Vacher de Lapouge 11 Cychrus Fabricius 13
Cryphalini 362 Cyclinus I{trby 72
Cryphalus Erichson 362 Cycliopleurus Hope 281
Cryptarcha Shuckard 217 Cyclocephala Latreille 155
Cryptarchinae 217 Cyclocephalini 155
Cryptobiina 116 Cycloneda Crotch 236
Cryptobiurn Mannerheim 116 Cyclotrachelus Chaudoir 35
Cryptocephalinae 305 Cylindera Westwood 10
Cryptocephalini 306 Cylindridia Casey 353
Cryptocephalus Miiller 306 Cylindrocharis Casey 31
Cryptocleptes Blackman 359 Cylindrocopturus Heller 348
Cryptohypnus Eschscholtz 17 8 Cylister Cooman 140
Cryptolestes Ganglbauer 220 Cylistix Marseul 140
Cryptophagidae 221 Cyllene Newman 284
Cryptophaginae 221 Cyllodes Erichson 217
Cryptophagini 221 Cyllopodes Murray 2\4
Cry ptophagistes Crotch 221 Cymatodera Gray 208
Cryptophagus Herbst 221 Cymatopterus Dejean 69
Cryptophilinae 224 Cymbiodyta Bedel 133
Cryptophilini 224 Cymindis Latreille 59
Cryptophilus Reitter 224 Cynaeus LeConte 255
Cryptopleurum Mulsant 135 Cypha Leach 108
Cryptorhopalum Gu6rin-M6neville 198 Cyphea Fauvel 107
Cryptorhynchidius Pierce 348 Cyphelophorus Kuwert 131
Cryptorhynchinae 347 Cyphicerini 334
Cryptorhynchini 347 Cyphina 107
Cryptorhynchus Illiger 348 Cyphirhinus Schiinherr 351
Cryptoweisea Gordon 229 Cyphomimus Horn 330
Cryptulocleplus Wood 359 CyphonPaykull 142
Crypturgini 361 Cyrtepistomus Marshall 334
Crypturgus Erichson 361 Cyrtinini 298
Ctenicera Latreille 179 Cyrtinus LeConte 298
Ctenistini 129 Cyrtocaraxenos Pierce 366
Ctenobium LeConte 202 Cyrtomoptera Motschulsky 193
Ctenodactylini 56 Cyrtomoptila Motschulsky 193
Ctenonychus Stephens 184 Cyrtophorus LeConte 286
Cucujidae 219 Cyrtoscydmus Motschulsky 83
Cucujinae 219 Cyrtotriplax Crotch 225
Cucujoidea 213 Cyrtusa Erichson 79
Cucujus Fabricius 219 Cytilus Erichson 160
Cupedidae 6
Cupedinae 6 Dacne Latreille 225
Cupedini 6 Dacninae 225
Cupes Fabricius 6 Dacyrtocara Pierce 366
Cupila Casey 127 Dalopius Eschscholtz 182
Curculio Linn6 342 Dalotia Casey 100
Curculionidae 329 Dampfius Pacheco 170
Curculionoidea 323 Danae Reiche 238
Curculioninae 342 Daniela Netolitzky 23
Curimopsis Ganglbauer 158 Danosoma Thompson 175
Curtonotus Stephens 35 Dascillidae 143
Curvimanon Fleischer 80 Dascillinae 143
Cybister Curtis 72 Dascilloidea 143
Cybistrini 72 Dasydera LeConte 150
Cybocephalinae 2I7 Dasyglossal{raatz 99
Cybocephalus Erichson 217 DasypogonLeConte 175
Cychramus Kugelann 216 Dasvrhadus Fall 213

405
Dasy'bes Faykull 212 Diaperini 255
Dasytinae 212 Diaperis Miiller 255
Datomicra Mulsant & Rey 100 Diaphoetes Waterhouse 120
Dearthrus LeConte 198 Diaphoromerus Chaudoir 45
Decarthron Brendel 128 Diaulota Casey 107
Dectes LeConte 297 Diazus LeConte 151
Deinopsini 108 Dibolia Latreille 320
Deinopsis Matthews 108 Dibolocelus Bedel 133
Deinopteroloma Jansson 91 Dicaelus Bonelli 48
Deleaster Erichson 92 Dicentrus LeConte 280
Deloyala Chewolat 323 Dicerca Eschscholtz 162
Delphacixenos Pierce 367 Dicercini 162
Delphastrrs Casey 229 Dicheirus Mannerheim 45
Delphota Jasey 103 Dichelonycho Kirby 153
Deltometopus Bonvouloir 187 Dichelonyx Harris 153
Deltostethus Sharp 134 Dichrornino Casey 155
Dendroctonus Erichson 357 Dictyopterus Mulsant 188
Dendroiders Latreille 269 Dicyrtocercyon Ganglbauer 134
Dendrophagus Schrinherc 220 Didetus LeConte 50
Dendrophilinae 138 Didion Casey 229
Dendrophilini 138 Dienerella Reitter 239
Dendrophilus Leach 138 Dietzella Champion 349
Denticollis Piller & Mitternacher t77 Dietzianus Sleeper 344
Deratanclrus Casey 55 Dimetrota Mulsant & Rey 100
Deratopeus Casey 115 Dimetrotina Casey 100
DerelomirLi 341 Dinaraea Thomson 102
Deretaphrini 252 Dineutus MacLeay 72
Deretaphrus Newman 252 Dinoderinae 200
Dermestes Linn6 196. 197 Dinodromius Casey 58
Dermestic{ae 196 Diocalandra Faust 355
Dermestirri 196 Diocarabus Reitter 12
Dermestinus Zhantiev 196 Diochini 123
Dermestoidea 195 Diochus Erichson 123
Derocrepis Weise 318 Diomus Mulsant 231
Derodontidae 195 DioxoceraPierce 366
Derodontinae 196 Diozocera Pierce 366
DerodontrLs LeConte 196 DiozodesHaldeman 281
Deronectes auct. amer. 66 DiphyaulonArnett 175
Derospidea Blake 314 Diplocampa Bedel 27
Derulus Tschitch6rine 31 Diplocheila Brull6 48
Derus Motschulsky 31 Diplocoelus Gu6rin-M6neville 241
Descarpentriesina Leraut 163 Diplotaxis Kirby 151
Desmatogaster Knutson 203 Diplous Motschulsky 30
Desmocerini 287 Dirabius Casey 353
Desmocerrrs Audinet-Serville 287 Dircaea Fabricius 248
Desmopachria Babington 63 Dirhaginae 187
Desmoris LeConte 340 Dirhagus Latreille 187
Devia Blar:kwelder 99 Disamara Lindroth 35
DiabroticeL Chewolat 315 Discoderus LeConte 43
Diacheila Motschulsky 15 Disonycha Chevrolat 319
Diachus LeConte 307 Distemmus LeConte 88
Diagraphct Newman 189 Distichalius Casey 122
Dialytellus Brown 149 Distigmoptera Blake 320
Dialytes Harold 149 Ditemnus LeConte 194
Dianchomeno Chaudoir 57 Ditropalia Casey 107
Dianous Samouelle 109 Ditylini 266
Diqnusa Casey 107 Ditylus Fischer von Waldheim 266
Diaperinae 254 Dolerosomus Motschulskv 182

406
Dolichocis Dury 244 Ecyrus LeConte 295
Dolichosoma Stephens 212 Elacatis Pascoe 270
Doliema Pascoe 255 Elaphidiini 280
Dolopiosomus Motschulsky 184 Elaphidion Audinet-Serville 281
Dolurgus Eichhoff 361 El ap hidionoide s Linsley 281
Donacia Fabricius 302 Elaphrini 15
Donaciella Reitter 302 Elaphropus Motschulsky 28
Donaciinae 302 Elaphrotatus Semenov 16
Donaciini 302 Elaphroterus Semenov 16
Donaciomima Medvedev 303 Elaphrus Fabricius 16
DonesiaCasey 103 Elasmocerus LeConte 208
Dorcaschema Haldeman 295 Elassoptes Horn 355
Dorcaschematini 295 Elater Linn6 181
Dorcasina Casey 291 Elateridae 175
Dorcatoma Herbst 204 Elaterinae 181
Dorcatominae 204 Elaterini 181
Dorcatomini 204 Elateroidea 174
Dorcini 144 Elateroides Schaeffer 206
Dorcus Macleay 144 Elathous Reitter 177
Dorthesia Say 250 Elatotrypes Fisher 284
Dorytomus Germar 339 Elatrinus Horn 184
Doydirhynchinae 324 Eleates Casey 255
Drachylis Casey 262 Eleodes Eschscholtz 256
Drapetes Dejean 186 Eleodini 256
Drasterius Eschscholtz 176 Eleodopsis Blaisdell 257
Drastophus Casey 82 Eleusinini 92
Dromaeolus Kiesenwetter 187 Elissa. Casey 331
Dromius Bonelli 58 Elleschini 345
Drusilla Leach 106 Elleschus Dejean 345
Dryocoetes Eichhoff 361 Ellipsoptera Dohktourow 10
Dryocoetini 361 Ellychnia LeConte 191
Dryophilinae 2Ol Elmidae 172
Dryophthorinae 356 Elminae 172
Dryophthorini 356 Elmini 172
Dryophthorus Germar 356 Elodes Latreille 142
Dryopidae 171 Elonium Leach 88
Dryopoidea 169 Elonus Casey 275
Dryops Olivier 171 Elytroleptus Dug6s 287
Dubiraphia Sanderson 172 ElytrusaCasey 101
Dularius Thomson 284 Embaphion Say 257
Dynastinae 155 Emmesa Newman 246
Dyschirius Bonelli 17 Empelinae 87
Dysidius Chaudoir 32 Empelus LeConte 87
Dyslobus LeConte 333 Emphanes Motschulsky 27
Dysmathes Mannerheim 61 Emphyastes Mannerheim 337
Dytiscidae 62 Emphyastini 337
Dytiscinae 71 Empinodes Park I27
Dytiscini 71 Emplenota Casey 105
Dytiscus Linn6 71 Enaphalodes Haldeman 280
Encalus LeConte 345
Eanus LeConte 181 Enchodes LeConte 247
Eburia Audinet-Serville 280 Encyclops Newman 288
Ecarinosphaerula Hatch 79 Endalus Castelnau 341
Echinaspis Blatchley 347 Endecatominae 200
Echocerus Horn 255 Endecatomus Melli6 200
Echochara Casey 105 Endectus LeConte 252
Echyra Erichson 154 Endeitoma Sharp 251
Ectopria LeConte 174 Endeodes LeConte 211

407
Endomychidae 237 Euaesthetus Gravenhorst 112
Endomychinae 239 EubaeoceraCornell 125
Endomychini 239 Eubriinae 174
Endomychus Panzer 239 Eubrychiopsis Dietz 351
Endrosq.LeConte 152 Eubrychius Thomson 351
Engamota Casey 100 Eubulus Kirsch 348
EngisPayxull 225 Eucanthus Westwood 150
Enhydrini 72 Euceratocerini 202
Enicmus Thomson 239 Euchaetes LeConte 353
Enicoceru-. Stephens 74 Euchariruq Casey 104
EnnearthronMelli6 244 Eucicones Sharp 251
Enochrus'lhomson 132 Eucinetidae 141
Enoclerus Gahan 209 Eucinetoidea l4l
Enopliini 210 Eucinetus Germar 141
Enoplurus Hope 131 Euclyptus Dietz 341
Entomophthalmus Bonvouloir 187 Eucnecosum Reitter 90
Entomosce:Iis Chevrolat 313 Eucnemidae 186
Epauloecus Mulsant & Rey 205 Eucneminae 187
Epeuchaet,zs Lyman 353 Euconnini 82
Ephalus LeConte 257 Euconnus Thomson 82
Ephistemtrs Stephens 223 Eucrada LeConte 201
Epicauta Dejean 264 Eucradini 201
Epicautina 264 Eucryptusa Casey 107
Epierus Edchson 138 Euderces LeConte 286
Epilachna Dejean 237 Eudromus l{trby 21
Epilachninae 237 Euferonia Casey 33
Epilachnirri 237 Eugnamptus Schijnherr 326
Epiphanis Eschscholtz 188 Euharpalops Casey 41
EpiphloeirLi 210 Eulathrobium Casey 116
Episcopell,ts Casey 43 Euliusa Casey 100
Episernus Thomson 202 Eumalus Sharp 92
Epismilia C ocker ell 229 Eumichthini 282
Epitrix Foudras 319 Eumichthus LeConte 282
Eplophorus Chevrolat 286 Eumicrota Casey 107
Epuraea Erichson 215 Eumitocerus Casey 99
Epuraeanella Crotch 215 Eumolpinae 308
Erchomus Motschulsky 96 Eumolpini 308
Eremninar: 334 Eumolpus Redtenbacher 309
Eremosis cles Gozis 312 Eumorphinae 238
Ergates Audinet-Servllle 27 7 Eunonia Casey 93
Ergatini 2',77 EunyssobiaCasey 353
Erichsonius Fauvel 117 EuophryumBroun 355
Eriopsilus Bates 295 Eupactus LeConte 204
Erirhininae 339 Eupagoderes Horn 334
Erirhinini 339 Eupariini 149
Erirhinus Schiinherr 339 Eupariini 324
Erirhipidi,z Casey 157 Euparius Schiinherr 324
Erirhipis Ilurmeister 157 Eupetedromus Netolitzky 25
Ernobiinae 202 Euphorhipis Casey 157
Ernobiini 202 EuphoriaBurmeister 157
Ernobius illhomson 202 Euphoriaspls Casey 157
Eronyxa Reitter 207 Eupisenus Casey 243
Eropterus Green 189 Euplectinae 126
Eros Newrnan 189 Euplectini 126
Erotides $Iaterhouse 189 Euplectus Leach 126
Erotylidae 225 Eupleurida LeConte 269
Erynephala Blake 315 Eupogonius LeConte 295
Euaesthetinae Il2 Eupogonocherus Linsley 296

408
Eupristocerus Deyrolle 167 Fallapion Kissinger 328
EurealeConte 174 Faroninae 126
Eurelymis Casey 212 Faronini 126
Euremus Bierig 119 Feronia Latreille 30
Eurhizophagus M6quignon 218 Fidia Walsh 309
Europhilus Chaudoir 52 Flavohelodes Klausnitzer 142
Eurydactylus LaFert6-S6nectbre 49 Florilinus Mulsant & Rey 199
Euryderus LeConte 40 FluuicolaDeKay 174
Eurygeniinae 274 Fornax Castelnau 187
Eurygeniini 274 Foveoscapha Park & Wagner 126
Eurylister Bickhardt 140 Funda Blackwelder 104
Eurymeloe Reitter 263 Furcacampa Netolitzky 26
Eurymetopini 253 Fuscocalathzs Ndgre 51
Eurymycter LeConte 325
Euryoptera Horn 284 Gabrius Stephens 119
Eurypalpus LeConte 174 Gabronthus Tottenham 119
Eurypogon Motschulsky 175 Galerita Fabricius 59
Eurysphindinae 213 Galeritino Jeannel 59
Eurysphindus LeConte 213 Galeritini 59
Eurystethus Seidlitz 270 Goleritula Strand 59
Eurytrachelus Motschulsky 21 Galeruca Miiller 315
Eurytrichus LeConte 45 Galerucella Crotch 314
Eusomalia Casey 160 Galerucinae 314
Eusphalerini 87 Galerucini 314
Eusphalerum Kraatz 87 Galeruclerus Gahan 210
Eusphyrus LeConte 325 Gastrallini 202
Euspilotus Lewis 137 Gastrallus Jacquelin du Val 202
Eustilbus Sharp 226 Gastrellarius Casey 32
Eustrophinae 246 Gastroeideq.Hope 312
Eustrophinus Seidlitz 246 Gastrolobium Casey 116
Eustrophus Illiger 246 Gastrophysa Chewolat 312
Eutetrapha Bates 298 Gastrotaphrus Buchanan 338
Eutheia Stephens 83 Gaurambe Thomson 207
Eutheiini 83 Gaurodytes Thomson 67
Euthyna Gistel 304 Gauropterus Thomson 123
Eutoxotus Casey 288 Gaurotes LeConte 289
Eutrichapion Reitter 328 Gehringia Darlington 19
Eutrichites LeConte 128 Gehringiini 19
Euvacusus Casey 273 Gelus Casey 348
Euvrilletta Fall 204 Genattus Sharp 197
Euxenus LeConte 324 Gennadota Casey 99
Euxestinae 227 Genyon Smetana 134
Evarthrus LeConte 35 Geoderces Horn 333
Evodinus LeConte 289 Geodercodes Casey 332
Euoplus LeConte 255 Geodromicus Redtenbacher 91
Evotus LeConte 332 Geomorus Schiinherr 336
Excavodes Park 127 Geomysaprinus Ross 137
Exema Lacordaire 307 Geopatrobus Darlington 29
Exochomus Redtenbacher 233 Geopinus LeConte 43
Exomella Casey 159 Georyssidae 135
Exopioide s Gu6rin-M6n6ville 199 Georyssus Latreille 135
Exosternini 140 Geostiba Thomson 101
Explorator Pacheco 169 Geotrupes Latreille 150
Geotrupinae 150
Fabricianus Weise 307 Geotrupini 150
Falacer Castelnau 259 Gephelophorus Sharp 131
Falagria Leach 105 Geraeus Pascoe 353
Falagriini 105 Gerstaeckeria Champion 347

409
Gibbiinae 205 C4tmnodus Kirby 157
Gibbium Scopoli 205 Qtmnota Gistel 163
Gibbobruchus Pic 301 Gymnusa Gravenhorst 108
Glaphyra Newman 282 Gymnusini 108
Glaphyrinae 150 Gynandropus Dejean 43
Glaphyrocanthon Martinez 146 Gynandrotarsus LaFert6-S6nectbre 44
Glaresis Erichson 150 Gynopterus Mulsant & Rey 206
Glipa LeConte 249 Gyrinidae 72
Glipostenoda Ermisch 250 Gyrininae 72
Glischrochilus Reitter 217 Gyrinini 73
Glycerius Casey 46 Gyrinulus Zaitzev 73
Glycobius LeConte 284 Gyrinus Mi.iller 73
Glyphony>: Candbze 182 Gyrohypnus Samouelle 124
Glyphuroplata Uhmann 322 Gyrophaena Mannerheim 106
Glyptasida Casey 254 Gyrophaenina 106
Glyptina l,eConte 317 Gyrophaenini 106
Glyptoderus LaFert6-S6nectdre 49 Gyrotus Casey 348
Glyptoma Erichson 92
Glyptophorus Park 221 Habrocerinae 99
Glyptoscelis Chevrolat 309 Habrocerus Erichson 99
Gnathacmaeops Linsley & Chemsak 290 Hadraule Thomson 245
Gnathium Kirby 266 Hadrobregmini 203
Gnathoncus Jacquelin du Val 138 Hadrobregmus Thomson 203
Gnathospa'.sta Horn 264 Hadroplontus Thomson 350
Gnathotri<'hoide s Blackman 3 64 Hadrotes Miiklin 120
Gnathotrit:hus Eichhoff 364 Haemoniini 303
Gnathusa Fenyes 99 Haida Keen 91
Gnatocerus Thunberg 255 Halictophagidae 366
Gnorimellrr Casey 158 Halictophaginae 366
Gnypeta Thomson 100 Halictophagus Bohart 366
Goes LeConte 295 Haliplidae 60
Golgia Mulsant 238 Haliplus Latreille 60
Goniacerirrae 128 Hallomeninae 246
Goniaderinae 262 HallomenusPanzer 246
Gonioctena Chewolat 313 Halobrecthina Bernhauer 103
Goniolophus Casey 47 Hamq.ba Casey 346
Gonocallus LeConte 282 Hapalaraea Thomson 88
Gonocephalum Chevrolat 257 Hapalorhinus LeConte 211
Gonoderini 259 Hapalosalia Casey 292
Gonoderus Motschulsky 31 Haplandrus LeConte 260
Gonops LeConte 325 Haplanthaxia Reitter 165
Gonotropi:t LeConte 325 Haplochile LeConte 29
Grammoptr;era Audinet-Serville 291 Haplocoelus Chaudoir 31
Graphicallo Monros 310 Haplophoria Casey 157
Graphisurus Kirby 297 Haplorhynchites Voss 327
Graphoderus Dejean 71 Haplostethops Casey 353
Graphops LeConte 309 Hargium Samouelle 289
Graptodera Chewolat 317 Harminius Fairmaire 177
Gratusus (Jasey 242 Harmoniaspls Casey 236
Gronevus Casey 228 Harpalellus Lindroth 42
Grynocharis Thomson 207 Harpalini 39
Grypidius Schiinherr 339 Harpalobrachys Tschitsch6rine 40
Grypus Germar 339 Harpalomerus Casey 41
Gymnetrirrae 346 Harpalus Latreille 40, 4l
Gymnetrini 346 Harpasta Gistel 305
Gymnetron Schtinherr 346 Hebestola Haldeman 294
Gymnis LeConte 152 Helichus Erichson 171
Gymnocht.hebius d'Orchymont 74 Helobium Leach 15

410
Helocerus Mulsant & Rey 199 Himatium Wollaston 355
Helocharini 132 Himatolabus Jekel 326
Helocombus Horn 133 HippodamiaDejean 234
HelodesPaykull 312 Hippopsini 296
Helophorinae 130 Hippopsis Lepeletier & Audinet-Serville 296
Helophorus Fabricius 130 Hippuriphila Foudras 318
Helopini 258 Hirmoplataphus Netolitzky 23
Helops Fabricius 258 Hispinae 321
Hemantus Casey 273 Hister Linn6 139
Hemicallidium Casey 283 Histeridae 135
Hemicarabus G6hin 12 Histerinae 139
Hemicoelus LeConte 203 Histerini 139
Hemicrepidius Germar 181 Histeropsis Chevrolat 255
HemicyphonLeConte 142 Holciophorus LeConte 31
Hemimedon Casey 113 Holoboreaphilus Campbell 89
Hemiptychus LeConte 205 Hololepta Paykull 140
Hemiquedius Horn 122 Hololeptini 140
Hemisopalus Casey 43 Holopleura LeConte 287
Hemispilota Casey 154 Holopleurini 287
Hemistenus Motschulsky 109 Holostrophus Horn 246
Henoticoides Hatch 221 Holotrochus Erichson 92
Henoticus Thomson 221 Homaeotarsus Hochhuth 116
Henotiderus Reitter 221 Homalochthebius Kttwefi 7 4
Henous Haldeman 264 Homalostylus Chevrolat 354
Hesperobaenus LeConte 2I8 Homalota Mannerheim 107
Hesperobium Casey 116 Homalotina 107
Hesperolinus Casey 124 Homalotusa Casey 102
Hesperophanes Mulsant 280 Homoeocarabus Reitter 11
Hesperophanini 280 Homoeolabus Jekel 326
Hesperosaprinus Wenzel 137 Hornoeosomo Austin 80
Hesperus Fauvel 119 HomogasterProvancher 348
Hetaeriinae 141 Homorosoma Frivaldszky 350
Hetaeriini 141 Hoplandria Kraatz 105
Hetaerius Erichson 141 Hoplandriini 105
Heterachthes Newman 281 Hoplia Illiger 154
Heteracthon Thomson 281 Hopliini 154
Heterarthron Dejean 199 Hoplocephalo Castelnau & Brull6 255
Heteraspis LeConte 309 Hoppingiana Blaisdell 212
Heterhelus Jacquelin du Val 214 Horistonotus Canddze 185
Heterlimnius Hinton 173 Hormorini 332
Heteroceridae 169 Hormorus Horn 332
Heterocerini 170 Hornaltica Barber 318
Heterocerus Fabricius 170 Hornia Riley 265
Heterodromla Haldeman 220 Horticola Lindemann 241
Heteronychus Seidlitz 67 Hybodera LeConte 282
Heterops Mannerheim 90 Hyboderini 282
Heteropus Castelnau 257 Hydaticini 71
HeterosilphaPortevin 85 Hydaticus Leach 71
Heterosternus Zimmermann 65 Hydaticus Sch6nherr 350
Heterosternuta Strand 65 Hydnobiini 78
Heterostethus Falkenstrom 65 Hydnobius Schmidt 78
Heterothops Stephens 121 Hydnocera Newman 208
Hetoernis Haldeman 295 Hydraena Kugelann 74
Hexarthrum Wollaston 355 Hydraenidae 74
Hexirhexius Grigarick & Schuster 126 Hydraeninae 74
Hiloro Mulsant & Rey 102 Hydraenini 74
Hilarina Casey 100 Hydriomicrus Casey 23
Himatinum Cockerell 355 Hydrium LeConte 21

tr
Hydrobiinae 131 Hypoborini 358
Hydrobiini 131 Hypocaccus Thomson 137
Hydrobiusr Leach 131 HypocoelusDejean 188
Hydrocan-:hus Say 62 Hypocoprinae 223
HydrocharaBerthold 133 Hypocoprus Motschulsky 223
Hydrochinae 130 Hypocyphtus Gyllenhal 108
Hydrochu,s Leach 130 Hypodacne LeConte 227
Hydrogale,ruco Laboissidre 314 Hypolithus Eschscholtz 178
Hydrophilidae 130 Hypomedon Mulsant & Rey 113
Hydrophilinae 133 HypomolyxleConte 337
Hydrophiloidea 130 Hypophlaeus Fabricius 256
Hydrophilus Miiller 133 Hypophloeinae 256
Hydroporinae 63 Hypophloeini 256
Hydroporini 64 Hypostenus Rey 109
Hydroponrs Clairville 65 Hypothenemus Westwood 362
HydrosmectaThomson 100 Hypulini 247
Hydrothassa Thomson 312 Hypulus Paykull 247
Hydrovatini 63 Ibidiini 281
Hydrovatrrs Motschulsky 63
Hygrotus Stephens 64 Ichnodes Dejean 262
Hylastes Ilrichson 356 Ichthyurini 195
Hylastini 356 Ictinosternun Smetana 134
Hylastinotdes Spessivetsev 357 Idiobates Casey 259
Hylastinus Bedel 357 Idiopidonia Swaine & Hopping 291
Hylecoetirrae 206 Idiostethus Casey 353
Hylecoetus Latreille 206 Ilendus Casey 242
Hylesininae 356 Ilybiosoma Crotch 67
Hylesinin; 357 Ilybius Erichson 69
Hylesinus Fabricius 357 Ilyobates Kraatz 99
Hylis des,]ozis 188 Ipelates Reitter 77
Hylobiini 337 Iphthiminus Spilman 261
Hylobius (]ermar 337 Ipini 360
Hyloceotu,:,; Melsheimer 206 Ips DeGeer 360
Hylocurus Eichhoff 359 Irichroa Newman 12
Hylota Casey 99 Isarthrus LeConte 187
Hylotrupes Audinet-Servllle 282 Ischalia Pascoe 269
Hylurgopinus Swaine 357 Ischaliinae 269
Hylurgops LeOonte 356 Ischnosoma Stephens 98
Hymenochara Campbell 260 Ischyropalpus LaFert6-S6nectdre 273
Hy menoph.orus Mulsant 259 Ischyrus Lacordafte 226
Hymenortrs Mulsant 259 Isochnus Thomson 345
Hypebaeus Kiesenwett er 212 Isoglossa Casey 99
Hypera Gr:rmar 336 Isohydnocera Chapin 209
Hyperaspidius Crotch 231 Isomira Mulsant 259
Hyperaspini 231 Isoplastus Horn 79
Hyperaspis Redtenbacher 23I Isopleurus Kirby 36
Hypericia Bedel 311 Isorembus Jeannel 48
Hyperinae 336 Isorhipina Casey 157
Hyperini 336 Isorhipis Boisduval & Lacordaire 187
Isosceles Newman 299
Hyperisus Mulsant &Rey 202
Hypermal,t u.s Lacordaire 280 Isostenia Vacher de Lapouge 11
Hyperodes Jekel 334 Isotachys Casey 29
Hyperplatys Haldeman 298 Ithyceridae 325
Hypherpes Chaudoir 31 Ithycerus Sch6nherr 325
Hyphydrini 63 Ithyporini 347
Hypnoryra Casey 124 Ixapion Roudier & Tempere 328
Hypnoidus Dillwyn 178 .Irios Deville 328

4t2
Jonthonota Spaeth 322 Latheticus Waterhouse 258
Judolia Mulsant 291 Lathridiidae 239
Lathridiinae 239
Kalissus LeConte 84 Lathridius Herbst 239
Katastylops Pierce 367 Lathrimaeunz Erichson 91
Knowltonia Fisher 166 Lathrium LeConte 89
Knulliana Linsley 286 Lathrobiella Casey 116
Korynetinae 210 Lathrobiina 115
Korynetini 211 Lathrobioma Casey 115
Kuschelina Bechyn6 320 Lathrobium Gravenhorst 115
Kytorhininae 300 Lathrolepta Casey 115
Kytorhinus Fischer von Waldheim 300 Lathrotaxis Casey 116
Lathrotropis Casey 116
Labidomera Chevrolat 311 Lebia Latreille 56
Laccobius Erichson 131 Lebiini 56
Lacconotus LeConte 267 Lechriopini 348
Laccophilinae 63 Lechriops Schcinherr 348
Laccophilini 63 Lecontea Olivier 190
Laccophilus Leach 63 Leconteus Lutshnik 30
Laccornis des Gozis 67 Lecontia Champion 268
Lachnocrepis LeConte 49 Lederia Reitter 246
LachnosternaHope 152 Leiestini 238
Lacon Castelnau 175 Leiochiton Curtis 19
Laemophloeinae 219 Leiodes Latreille 79
Laemophloeus Dejean 219 Leiodidae 77
Laemosaccini 342 Leiodinae 78
Laemosaccus Schdnherr 342 Leiodini 78
Laemostenus Bonelli 51 Leiolinus Casey 124
Laestris Melsheimer 109 Leionotus Kirby 71
Laetulonthus Moore & Legner 119 Leirus Zimmermann 35
Lagarus Chaudoir 33 Leistus Frrilich 13
Lagriidae 261 Lema Fabricius 304
Lagrlinae 262 Lemiini 304
Lamiinae 294 Lerua Casey 7I3
Lamiota Casey 101 Leperisinus Reitter 357
Lamperos Allard 258 Lepesoma Motschulsky 333
Lamprias Bonelli 56 Lepiarcha Sharp 217
Lampropterus Mulsant 282 Lepidophorus Kirby 333
Lampyridae 190 Lepidotus Stephens 175
Lampyrinae 190 Lepisomus Kirby 359
Languria Latreille 224 Leptacinodes Casey 123
Languriidae 223 Leptacinus Erichson 123
Languriinae 224 Leptacmaeops Casey 290
Langurlini 224 Leptaegialia Brown 147
Lanternarius Pacheco 170 Leptalia LeConte 289
Lapethus Casey 227 Leptinillus Horn 82
Laphyra Newman 282 Lentininae 82
Lappus Casey 273 Leptinotarsa StAl 311
Lapsus Pacheco 170 Leptinus Mtiller 82
Lara LeConte 172 Leptoferonia Casey 32
Laricobius Rosenhauer 196 Leptomus Casey 8
Lafinae I72 Leptophloeus Casey 220
Larini 172 Leptopiini 333
Larisia Err,ery 276 Leptorus Casey 114
Lasconotus LeConte 251 Leptoscydmini 83
Lasioderma Stephens 204 Leptoscydmus Casey 83
Lasiodermini 204 Leptostylus LeConte 297
Lasiotrechus Ganglbauer 19 Leptothrix Heikertinger & Csiki 318

413
Leptotrachelus Latreille 56 Liparocephalus Miiklin 108
Leptotrichaltica Helkertinger 3 18 Lispinini 92
LeptotrixHorn 318 Lissominae 186
Leptotyphlinae 113 Lissorhoptrus LeConte 341
Leptura Linr'6 292 Listemus Casey 160
Lepturges Bates 297 Listronotus Jekel 334
Lepturinae 287 Listrus Motschulsky 213
Lepturini 288 Litargus Erichsor' 242
Lepturopsis Linsley & Chemsak 293 Litheleodes Blaisdell 256
Leptusa I{raatz 107 Lithocharis Dejean 113
Lepyrus Germar 337 Lithocharodes Sharp 123
LestqBlackwelder 90 Lithodus Germar 335
Lesteva Latreille 90 Litochropus Erichson 226
Leuchydrium Casey 25 Litodactylus Redtenbacher 350
Leucopaed,erus Casey 116 Litolethra Casey 115
Lexiphaners Gistel 306 Lixellus LeConte 335
Liaphlus ()uignot 61 Lixestus Reitter 336
Liq,semum, Casey 279 Lixini 337
Librodor Reitter 217 Lixus Fabricius 337
LichenophLanes Lesne 200 Loania Kinzelbach 366
LichnantL,e Burmeister 150 Loberini 224
Lichnocarabus Reitter 11 Loberus LeConte 224
Licinini 48 Lobiopa Erichson 216
Ligmargus Stibick 178 Lobosoma Buchanan 338
Lignyodes Dejean 345 Lobrathium Mulsant & Rey 116
Lignyodini 345 Loceptes Casey 347
Ligyrellus Casey 156 Lodinus Mulsant &Rey 257
Liryrodes Casey 156 Longitarsus Berthold 317
Ligyrus Burmeister 156 Lopha Stephens 27
Lilioceris Reitter 304 Lophalophus LeConte 333
Limnebiui; Leach 74 Lopheros LeConte 189
Limnichiciae 171 Lophioderini 82
Limnichir.ae 171 Lophioderus Casey 82
Limnichir.i 171 Lophoglossus LeConte 30
Limnichites Casey 171 Lophomucter Notman 105
Limnocharis Horn 74 Lophopogonius Linsley 296
Limonius Eschscholtz 176 Lordithon Thomson 98
Limulodes Matthews 76 Loricera Latreille 15
Limulodidae 76 Loricerini 15
LinaDejean 313 Lorinota Casey 105
Lindorus'Casey 233 Loxandrus LeConte 30
Liniolis Grigarick & Schuster 126 Loxopeza Chaudoir 56
Linohespe'rus Smetana 123 Lucanidae 143
Linolathro.Casey 116 Lucaninae 144
Linsleya MacSwain 265 Lucanini 144
Linsleyana, Podany 288 Lucanus Scopoli 144
Linsleyina Selander 265 Lucidota Castelnau 191
Liocellus Motschulsky 46 Lucifotychus Park & Wagner 129
Liocellus Tschitch6rine 46 Ludius Berthold 181
Liocosmius Casey 25 Lumetus Zaitzev L32
Liocyrtusa Daffner 79 Luperini 315
Liodessus Guignot 63 Lutrochus Erichson 171
Liodicaelus Casey 49 Lycidae 188
Liogluta Thomson 102 Lycinae 188
Lioligus Casey 159 Lycoperdina Latreille 238
Lionepha Casey 21 Lycoperdinini 238
Lionothus Brown 78 Lyctidae 201
Lioon Casey 159 Lvctini 201

4L4
Lyctus Fabricius 201 Mathrilaeurn Moore 91
Lygerus Kiesenwetter 195 Matini 71
Lygistopterini 188 Matus Aub6 71
Lymantor Lovendal 361 Mecas LeConte 299
Lymexylidae 206 Mecinini 347
Lymexyloidea 206 Mecinus Germar 347
Lyperopherus Motschulsky 35 Mecopeltus Dietz 351
Lyperosomus Motschulsky 33 Mecynotarsus LaFert6-S6nectdre 271
Lyperostenia Vacher de Lapouge 10 Medon Stephens 113
Lyperus Chaudoir 33 MedonellaCasey 113
Lypoglossa Fenyes 103 Medonina 113
Lyrosoma Mannerheim 77 Medonodonta Casey 114
Lytta Fabricius 264 Megacerus Fahraeus 300
Lyttina 264 Megacrotono Scheerpeltz 103
Lyttim 264 Megacyllene Casey 284
Megalechthrus Perkins 366
Macdonaldium Abdullah & Abdullah 76 Megaleptura Casey 292
Machaerodes Brendel 128 Megaliridia Casey 12
Macratria Newman 275 Megalodacne Crotch 225
Macratriini 275 Megalostomini 305
Macreurops Casey 218 MegalticaHatch 317
Macrobasis LeConte 265 Meganemognatha Enns 265
Macrodactylini 153 Megapangus Casey 40
Macrodactylus Latreille L54 Megapenthes Kiesenwetter 184
Macrodea Casey 222 Megaquedius Casey 121
MacrodytesThomson 71 Megarafonus Casey 126
Macronaemia Casey 233 Megarthrus Curtis 87
Macronoxia Crotch 152 Megasemum l{raatz 279
Macronychini 173 Megasternini 134
Macronychus Miiller 173 Megasternum Mulsant 134
Macropogon Motschulsky 174 Megastilicus Casey 114
Macropogoninae 174 Megatoma Herbst 198
Macrops Kirby 334 Megatomini 197
Macrorhoptus LeConte 343 Megilla Mulsant 233
Macrosiagon Hentz 250 Megodontus Solier 12
Macrotelus Klug 208 Megomus Casey 8
Maculicq.uta Dillon 264 Meibomeus Bridwell 301
Madarellus Casey 353 Melalgus Dejean 199
Madarini 352 Melamomphus Horn 333
Magdalinae 342 Melanagonum Casey 53
Magdalini 342 Melandrya Fabricius 246
Magdalis Germar 342 Melandryidae 245
Malachiinae 2l]- Melandryiinae 246
Malachius Fabricius 211 Melandryini 246
Malacocis Gorham 245 Melaneleodes Blaisdell 256
Mallodrya Horn 243 Melanius Bonelli 33
Malporus Casey 273 Melanophila Eschscholtz L65
Malthacus Kirby 192 Melanophilini 165
Malthinini 194 Melanophthalma Motschulsky 240
Malthinus Latreille 194 Melanotini 184
Malthodes Kiesenwetter 194 Melanotrupes Blanchard 150
Mantura Stephens 319 Melanotus Eschscholtz 184
Maphoca Casey 245 Melanthaxia Richter 165
Margaiostus Stibick 178 Melasinae 187
Margarinotus Marseul 139 Melasis Olivier 187
Marginus LeConte 241 Melq.soma Stephens 313
Margus Dejean 258 Melasomida Schaeffer 312
Masoreini 56 Melba Casey 127

4r5
Meligethes Stephens 217 Microhyus LeConte 347
Meligethinae 217 MicrolathraCasey 116
Melittophcgus Kirby 263 Microlestes Schmidt-Goebel 58
Meloe Lir.n6 263 Microlipus LeConte 211
Meloidae 263 Micromalthidae 6
Meloinae 263 Micromalthus LeConte 6
Meloini 263 Micromaseus Casey 34
MelolontkLinae 151 MicromedonCasey 113
Melolonthini 151 Micromelomalus Casey 22
Melomalus Casey 22 Micromota Casey 100
Melyridaer 211 Micronychus Provancher 333
Membrqctxenos Pierce 366 Micropeplidae 84
Menechid'zs Motschulsky 258 Micropeplini 84
Meracantha Kirby 259 Micropeplus Latreille 84
Merhynchites Sharp 327 Microrhagus Dejean 187
Meridion des Gozis 283 Microrhopala Chewolat 322
Meriellum Linsley 283 Microsaurus Stephens 121
Merinus LeConte 260 Microscaph a LeConte 246
Merium LeConte 283 Microschatia Solier 254
Meronera Sharp 100 Microscydmus Saulcy & Croissandeau 83
Mesauletes Voss 327 Microtachys Casey 29
Mesosini 295 Microtonus LeConte 247
Metablap";'lis Blaisdell 257 Microweise a Cockerell 229
Metqbletus Schmidt-Goebel 58 Microweisini 229
Metabola Chaudoir 57 Millidium Motschulsky 75
Metachro:na Chewolat 308 Minyomerus Horn 331
Metaclisa Jacquelin du Val 258 Mioptachys Bates 29
Metacmaeops Linsley & Chemsak 290 Miscodera Eschscholtz 19
Metallina Motschulsky 21 Mitosynum Campbell 93
Metallogra,pho Motschulsky 310 Mocyta Mulsant & Rey 102
Metamelanlus Tschitsch6rine 33 Molamba Casey 228
Metaxyodonla Casey 113 Molorchini 282
Methlini 63 Molorchus Fabricius 282
Methydrus Rey 132 Moluciba Casey 99
MetoponWaterhouse 327 Molytinae 337
Metrius Eschscholtz 19 Monachini 306
Mezium Clurtis 205 Monachulus Leng 306
Miarus Schiinherr 347 Monachus Chevrolat 306
Miccotrogus Schajnherr 346 Monarthrum Kirsch 365
Micracini 359 Moneilema Say 294
Micracis l,eConte 359 Moneilemini 294
Micracisoides Blackman 359 M onillop atrob us }Iatcli' 29
Micraegialia Brown 146 Monocerus Faldermann 27 |
Micragonum Casey 54 Monochamini 294
Micralymma Westwood 88 Monochamus Dejean 294
Micrathous Lane I77 MonoferoniaCasey 32
Micrepitrix Laboissidre 318 Mononychini 349
Microaug;vles Pacheco 170 Mononychus Germar 349
Microbledius Herman 93 Monophylla Spinola 208
Microbregma Seidlitz 203 Monoplesa Motschulsky 294
Microcallidium Casey 283 Monotoma Herbst 218
Microcara Thomson 142 Monotominae 218
Microclytus LeConte 286 Monoxia LeConte 315
Microcylloepus Hinton 173 Mordella Linn6 248
Microdota Mulsant & Rey 102 Mordellidae 248
Microedus LeConte 91 Mordellini 248
Microgoes Casey 295 Mordellistena Costa 249
Micrograrnrne W alkJey 239 Mordellistenini 249

416
Morphnosoma Lutshnik 33 Neacanthocinus Dillon 296
Morychus Erichson 160 Neandra Lameerc 277
Motschulskium Matthews 75 Neanthophylax Linsley & Chemsak 289
Muirixenos Pierce 367 Neatus LeConte 259
Mulsantina Weise 237 Nebria Latreille 14
Murmidiinae 227 Nebriini 13
Murmidius Leach' 227 Necrobia Olivier 211
Mutinus Casey 112 Necrodes Leach 85
Myas Dejean 31 Necrophila Kirby & Spence 85
Mycetaea Stephens 238 Necrophilus Latreille 77
Mycetaeinae 238 Necydalini 287
Mycetaeini 238 Necydalis Llnn6 287
Mycetina Mulsant 239 Nedyus Sch<jnherr 350
Mycetinini 238 Negastrius Thomson 178
Mycetochara Berthold 260 Neichnea Wolcott & Chapin 210
Mycetochares Latreille 260 Nelites Le0onte 254
Mycetocharini 260 Nemadini 81
Mycetocharis Gyllenhal 260 Nemadus Thomson 81
Mycetophagidae 242 Nematodes Berthold 188
Mycetophagus Hellwig 242 Nematolinus Casey 123
Mycetophila Gyllenhal 260 Nematoplinae 263
Mycetoporus Mannerheim 98 Nematoplus LeConte 263
Mychocerus aluct. 227 Nemocestes Van Dyke 332
Mychocerus Erichson 227 Nemognatha Illiger 265
Mycocerinus Slipinski 227 Nemognathina 265
Mycteridae 267 Nemonychidae 323
Mycterus Clairville 267 Nemosoma Latreille 207
Myeleborus Blackman 363 Nemota Casey 103
Myllaena Erichson 108 P odany 292
Neoalostern a
Myllaenini 108 Neobisnius Ganglbauer 117
Myloechus Latreille 81 Neobrachinus Erwin 18
Myochrous Erichson 309 Neocalathus Ball & Ndgle 51
MyocorynaStel 311 NeocarabusHatch 11
Myodes Latreille 250 Neoceletes Leng & Shoemaker 189
Myodites Latreille 250 Neochlamisus Karren 307
Myrmecocephalus Macleay 105 Neocholax Pierce 366
Myrmedoniini 106 Neochthebius d'Orchymont 75
Myrmedophila Bousquet 222 Neoclypeodytes Young 64
Myrmex Sturm 341 Neoclytus Thomson 285
Myrmolamio Bates 298 Neocychrus Roeschke 13
Mystaxus Kugelann 248 Neocyrtusa Brown 78
MyziaMulsant 235 Neoelaphrus Hatch 16
NeoemmesaHatch 246
Nacerdes Dejean 266 Neogalerucella Chujo 314
Nacerdinae 266 Neohaemonia Sz6kessy 303
Namunaria Reitter 251 Neoharmonia Casey 236
Nanobius Herman 95 Neoharmonia Crotch 236
Nanodactylus Blatchley 328 Neoheterocerus Pacheco 170
Nanodes Schtinherr 328 Neohypnus Coiffait &Saiz 124
Nanophyes Schiinherr 328 Neoleistus Erwin 13
Nanophyini 328 Neomida Latreille 255
Narpus Casey 173 Neomysia Casey 235
Nartus Zaitzev 70 Neonicrophorus Hatch 85
Nassipa Ernery 276 Neopachylopus Reichardt L38
Nathicus Casey 271 Neopaederus Blackwelder 116
Nathrenus Casey 198 Neopanscopus Pierce 333
Naupactini 329 Neopatrobus Darlington 29
Nausibius Redtenbacher 220 Neophycocoetes O'Brien & Wibmer 337

4r7
Neophytobius Wagner 351 Obriini 281
Neopolya.rthron Semenov 278 Obrium Dejean 281
Neoporus Guignot 65 Ocalea Erichson 99
Neopyrochroa Blair 269 Occlephelinus Hatch 89
Neorthopleura Barr 210 Ochodaeinae 149
Neosaprinus Bickhardt 137 Ochodaeus Audinet-Serville 149
Neoscutopterus Balfour-Browne 70 Ochrosidia Casey 155
Neostylops Pierce 367 Ochthebiinae 74
Neotrochus Blackwelder 92 Ochthebius Leach 74
Neozonitis Enns 265 Ochthedromus LeConte 21
Nephanes Thomson 76 Ochthephilum Stephens 116
Nephus Mulsant 231 Ochthephilus Mulsant & Rey 94
Nicaginae 144 Ochthexenus Motschulsky 88
Nicagini 144 Octotemnus Melli6 245
Nicagus l,eConte 144 Ocyusa l{raatz 99
Nicobiini 202 OdacanthellaLiebke 56
Nicrophorinae 85 Odacanthini 56
Nicrophorus Fabricius 85 Odontaeus Fischer von Waldheim 150
Niptus Boieldieu 205 Odontium LeConte 20
Nlsa Casey 128 Odontocorynus Schijnherr 353
Nisaxis Casey 128 Odontomus Kirby 166
Nitidula Fabricius 216 Odontonyx Stephens 51
Nitidulidae 214 Odontopus Say 345
Nitidulinae 215 Odontosphindini 213
Nitionini 254 Odontosphindus LeConte 213
Nocheles LeConte 333 Odontota Chewolat 321
Noda Chevrolat 308 Odontotaenius Kuwert 145
Nodinini 308 OebaceresThomson 295
Nodonota Lefbvre 308 Oedemeridae 266
Nomaretus LeConte 13 Oedemerinae 266
Nomaspis LeConte 264 Oedostethus LeConte 178
Nomidus Casey 334 Oestodes LeConte 181
Nomiini 29 Oestodini 181
Nominus Motschulsky 59 Oiceoptoma Leach 85
Nomius Castelnau 29 Olibrini 226
Nordenskioeldia Sahlberg Il2 Olibrus Erichson 226
Nosodendridae 196 Oliganomalo Casey 154
Nosodenclron Latreille 196 Oligolinus Casey 123
Nosodes LeConte 207 Oligomerodes Fall 204
Notaphusr Stephens 25 Oligomerus Redtenbacher 202
Notaris Ciermar 339 Oligomia Casey 100
Noteridaer 61 Oligopterus Casey 113
Nothopus LeConte 40 Oligotini 108
Nothus Olivier 248 Olisthaerinae 92
Notiodes Schiinherr 341 Olisthaerus Dejean 92
Notionomus Erichson 341 Olisthopus Dejean 51
Notiophilini 15 Olla Casey 236
Notiophilus Schtinherr 341 Olophrum Erichson 89
Notiophilus Dum6ril 15 Omq.lq. Motschulsky 27
Notobia Perty 45 Omaliinae 87
Notoxinae 271 Omaliini 88
Notoxini 271 Omalium Gravenhorst 88
Notoxus Fabricius 271 Omq.seidius Jeannel 33
Novelsis Casey 197 Omaseulus Lutshnik 34
Nudatoconnus Franz 83 Omuseus Dejean 33
Nudobius Thomson l-23 Omias Germar 332
Omoglymmius Ganglbauer 7
Oberea Mulsant 299 Omonadus Mulsant & Rev 273

418
Omophron Latreille 16 OrsonyxLeConte 151
Ornophronini 16 Orthaltica Crotch 318
Omorgus Erichson 150 Orthoboris Casey 351
OmoscylonThomson 294 Orthocis Casey 245
Omosita Erichson 216 Ortholeptura Casey 293
Omus Eschscholtz 8 Orthoperini 228
Oncideres Lepeletier & Audinet-Serville 296 Orthoperus Stephens 228
Onciderini 296 Orthoris LeConte 352
Onibathum Tottenham 87 Orthosoma Audinet-Sewllle 27 7
OnichodonNewman 187 Orthotomicus Ferrari 360
Ontholestes Ganglbauer 120 Orthotomides Wood 360
Onthophagini 146 Orus Casey 114
Onthophagus Latreille 146 Oryctini 156
Onthophilus Leach 138 Oryzaephilus Ganglbauer 220
Onychobaris LeConte 352 Osmoderma LePeletier & Audinet-Serville 157
Onychotrupes Jekel 150 Osmodermini 157
Onychylis LeConte 341 Osoriinae 92
Oodes Bonelli 49 Osoriini 92
Oodini 49 Osphya Illiger 248
Opatrinae 256 Osphyinae 248
Opatrini 257 Osphyini 248
Ophistomis Thomson 290 Ostoma Laicharting 207
Ophonus Dejean 40 Ostomodes Reitter 207
Ophraella Wilcox 315 Othiini 123
Ophryastes Say 334 Othniinae 270
Ophryastini 334 Othnius LeConte 270
Opilo Latreille 209 Otidocephalinae 341
Opisthiini 15 Otidocephalini 341
Opisthius Kirby 15 Otidocephalzs Chevrolat 341
Oplosia Mulsant 296 Otiorhynchinae 331
Opresus Casey 83 Otiorhynchini 331
Opsimini 280 Otiorhynchus Germar 331
Opsimus Mannerheim 280 Oulema des Gozis 304
Optioservus Sanderson 173 Oulimnius des Gozis 173
Opuntiaphila Pierce 347 Oxoplus LeConte 287
Orchesia Latreille 246 Oxybleptes Smetana 124
Orchesiini 246 Oxygonus LeConte 180
Orchestes Illiger 345 Oxylaemus Erichson 252
Ordobrevia Sanderson 173 Oxymedon Casey 113
Oreina Monros & Bechyn6 311 Oxynychus LeConte 231
Oreocharq Casey 104 Oxyomus Stephens 148
Oreodytes Seidlitz 67 Oxypoda Mannerheim 99
Oreoxenus Tschitch6rine 46 Oxypodini 99
Orestia Germar 318 Oxyporinae 109
Orestioides Hatch 318 Oxyporus Fabricius 109
Ormiscini 325 Oxypselaphus Chaudoir 52
Ormiscus Waterhouse 325 Oxypteris Kirby 165
Orobanus LeConte 90 Oxytelinae 92
Orochares Kraatz 91 Oxytelini 94
Orophiinae 245 Oxytelus Gravenhorst 95
Orophiini 245
Oropus Casey 126 Pachybrachini 305
Orphelophorus d'Orchymont 131 Pachybrachys Chevrolat 305
Orphilini 199 Pachybruchus Pic 300
Orphilus Erichson 199 Pachydela Rivalier 8
Orsodacna Dejean 303 Pachydeniini 153
Orsodacne Latreille 303 Pachylobius LeConte 338
Orsodacninae 303 Pachyphanes Dietz 340

4t9
Pachyrhjnus Schcinherr SB0 Paraplinthus Faust 338
Pachy rhSt nchus l{trby 325 Paraptochus Seidlitz 332
Pachysctrelini 168 Paraquedius Casey 122
Pachyschelus Solier 168 Parargutor Casey 32
Pachystilicus Casey 114 Parascydmus Casey 83
Pachystylus Rey 305 Parastenus Heyden 110
Pachyba Dejean 288 Paratachys Casey 29
Pachytychius LeConte 340 Paratenetus Spinola 262
Pactopus LeConte 186 Paratrogophloeus Hatch 94
Poederillus Casey 116 Paratropa Vacher de Lapouge 10
Paederina 116 Paratychius Casey 346
Paederinae 113 Parazonitis Enns 265
Paederini 113 Parelaphidion Skiles 281
Paederus Fabricius 116 Parenthis Dietz 350
Palaminus Erichson 113 Paria LeConte 308
Pallodes Erichson 216 Parilendus Casey 242
Palorus Mulsant 258 Parmenini 294
Palporus Campbell 97 Parmulus Harold 228
Panagaeini 49 ParocaleaBernhauer 105
Panagaeus Latreille 49 Paromalini 138
PanalotaCasey 101 Paromalus Erichson 138
Pandeletr:ius Schtinherr 331 Paropsimeno Motschulsky 311
Panscopi,l,ius Pierce 333 Parothius Casey 123
Panscopu.s Schiinherr 333, 334 Pasimachus Bonelli 17
Parabracteon Notman 20 Passalidae 145
Paracercrron Seidlitz 134 Passalotaenlus Kuwert 145
Paraclivi:ra Kult 18 Passandrinae 2I9
Parac5rmus Thomson 132 Patrobini 29
Paradelitrthrum Hatch gt Patroboidea Van Dyke 30
Paradicaelus Ball 48 Patrobus Dejean 29
Paradilacra Bernhauer 102 Pauronemognatha Enns 265
Parafilum,is Casey 207 Paussini 19
Paragonu.m Casey 53 Pectotoma Hatch 276
Paralarica Breuning 295 Pedetes Kirby 182
ParalathraCasey 116 Pediacus Shuckard 219
Paralesteua Casey g0 Pedilldae 274
Paraliaphlus Wallis 60 Pedilinae 274
Paralister Bickhardt 139 Pedilophorini 160
Paralitargus Casey 242 Pedilus Fischer von Waldheim 274
Parollelin.a Casey 291 Peditatus Pacheco 170
Parallelostethus Schwarz 181 Pelates }lorn 77
Paralopha Casey 27 Pelatines Cockerell 77
Paralytta Selander 264 Pelecomalium Casey 90
ParqmedonCasey 114 Pelecotoma Fischer von Waldheim 250
Parametriona Spaeth 323 Pelecotominae 250
ParamysiaReitter 235 Pelec;ryhorus Solier 254
Paranaemia Casey 233 Pelenomus Thomson 351
Paranchus Lindroth 52 Pelidnota Macleay 155
Parandra Latreille 277 Pelidnotidia Casey 155
Parandrinae 277 Pelophila Dejean 13
Parandrini 277 Peltastica Mannerheim 196
Paranomctla Casey 154 Peltasticinae 196
Pq,ranomus Kiesenwetter 181 Peltideo Motschulsky 207
Paranove].sis Casey 197 Peltinae 207
Paranthonomus Dietz 343 Peltini 207
Paraopsiraus Champlain & Knull 280 Peltodytes R6gimbart 61
Parapach;yto Casey 288 Pemphus Motschulsky 13
Parapanscopus Buchanan 334 Penetini 253

420
Pentacladocera Pierce 366 Philodes LeConte 4?
Pentagonica Schmidt-Goebel 50 Philonthini 117
Pentagonicini 50 Philonthus Curtis 117
Pentagrammaphila Pierce 366 Philopedon Schiinherr 330
Pentanota Bernhauer 100 Philophuga Motschulsky 58
Pentaphyllus Dejean 256 Philopuntia Pierce 347
Pentaria Mulsant 276 Philostrq.tus Zimmerman 338
Pentariini 276 Philothermus Attb6 227
Pentarthrum Wollaston 355 PhiloxylonLeConte 202
Pentaxocerq. Pierce 366 Phlaeopterus Motschulsky 90
Pentazocera Pierce 366 Phloeocharinae 95
Penthe Newman 243 Phloeonaeus Erichson 93
PenthelispaPascoe 252 Phloeonomus Heer 88
Penthinae 243 Phloeophagus Schtinherr 355
Pentodontini 156 Phloeopora Erichson 105
Pentozoe Pierce 366 Phloeopterus LeConte 90
Peranus Lewis 139 Phloeosinini 358
Perapion Wagner 328 Phloeosinus Chapuis 358
Percosia Zimmermann 36 Phloeostiba Thomson 88
Pergetus Casey 274 Phloeotribini 358
Perigaster Dietz 349 Phloeotribus Latreille 358
Perigona Castelnau 51 Phloeotrya Stephens 248
Perigonini 51 Phlogistosternus Wolcott 210
Perimecus Stephens 184 Phobetus LeConte 153
Perimegatoma Horn 198 Phomalus Casey 275
Peristethus LeConte 33 Phosphaenus Castelnau 191
Peritelini 332 Photinini 190
Peritelinus Casey 332 Photinus Castelnau 190
PerthalycraHorn 216 Photurinae 191
Peryphanes Jeannel 24 Photuris Dejean 191
Peryphodes Casey 25 Phratora Chevrolat 313
Peryphus Stephens 24 Phrenapatinae 253
Petaliini 204 Phryganophilus Sahlberg 247
Petalium LeConte 204 Phrypeus Casey 28
Pezocrosita Jacobson 312 Phthora Mulsant 253
Phacophallus Coiffait 123 Phycocoetes LeConte 337
Phaedon Megerle von Miihlfeld 312 Phyla Motschulsky 21
Phaenocerus Bonvouloir 187 Phfi.etes Redtenbacher 255
Phaenog'yra Seevers 107 Phylethus Redtenbacher 255
Phaenonotum Sharp 133 PhyIIis Gistel 310
Phaenops Dejean 165 Phyllobaenini 208
Phalacridae 226 Phyllobaenus Dejean 208
Phalacrini 226 Phyllobaenus Spinola 210
Phalacrus Paykull 226 Phyllobiini 332
Phaleriini 254 Phyllobius Germar 332
Phaleromela Reitter 254 Phyllobrotica Chewolat 316
Pharalus Casey 40 Phyllodecta Kirby 313
Phausis LeConte 191 Phyllodrepa Thomson 88
Phelister Marseul 140 Phyllophaga Harris 152
Phellidius LeConte 255 Phyllotreta Chevrolat 316
Phellopsis LeConte 252 Phyllotrox Schtinherr 341
Pheloconus Roelofs 347 Phymaphora Newman 238
Phengodes Illiger 190 Phymatinus LeConte 334
Phengodidae 190 Phymatodes Mulsant 283
Phenolia Erichson 216 Phymatodina Casey 283
Phidola Thomson 295 Physocnemum Haldeman 284
Philq.s Portevin 85 Physonota Boheman 322
Philhygra Mulsant & Rey 102 Physonotini 322

427
Physorhini 184 Platypodidae 365
Phytobiini 350 Platypodinae 365
Phytobius Schonherr 350 Platypodini 365
PhytodectaKirby 313 Platypsyllus Ritsema 82
Phytoecini 299 Platypsyllus Westwood 82
PhytonNewman 281 Platypus Herbst 365
Phytonomu,s Schiinherr 336 Platysoma Leach 140
Phytosini 107 Platysomatini 140
Phyxelis Schtinherr 333 Platystethus Mannerheim 94
Piazorhinini 345 Platytrachelus Motschulsky 21
Piazorhinus Schiinherr 345 Platyusa Casey 106
Pidonia Mulsant 292 Plectographo Rivalier 10
Piestinae 92 Plectralidus Casey 40
Piestini 92 Plectrura Mannerheim 294
PilemaLeConte 282 Plegaderini 136
Pilopius Casey 129 Plegaderus Erichson 136
Pinacodera Schaum 59 Plesiocis Casey 245
Pinodytes Florn 80 Pleurophoro Chewolat 304
Pinophilini 113 Pleurophorus Mulsant 149
Piosoma LeConte 40 Pleuropompha LeConte 264
Pisenus Casey 243 Pleurostica Motschulsky 312
Pissodes G'ermar 338 Pleurotobia Casey 107
Pissodinae 338 Plinthini 338
Pityobini 176 Plinthodes LeConte 333
Pityobius l,eConte 176 Plocamus LeConte 353
Pityogenes,Bedel 360 Plocetes LeConte 346
Pityokteines Fuchs 360 Pocadius Erichson 216
Pityomacer Kuschel 324 Podabrini 192
Pityophagrrs Shuckard 217 Podabrus Westwood 192
Pityophthorus Eichhoff 363 Podapion Riley 329
Placonotusr Macleay 219 Poecilium Fairmaire 283
Placopterus Wolcott 209 Poecilobrium Horn 282
Placusa Erichson 107 Poecilochroo Chevrolat 209
Plagiodera Chevrolat 312 Poecilus Bonelli 30
Plagiometriona Spaeth 323 Pogonocherini 295
Planesus Motschulsky 59 Pogonocherus Dejean 296
Plataphodes Ganglbauer 22 Polemius LeOonte 194
Plataphus Motschulsky 22 Poliaenus Bates 295
Platerodini 188 Polistoma Casey 105
Plateros Bourgeois 189 Pollaclasis Newman 191
Plateumarini 302 Polycaon Castelnau 199
Plateumaris Thomson 302 Polycaonini 199
Platidiolus Chaudoir 30 Polycestinae 161
Platidius Chaudoir 30 Polycestini 161
Platybregmus Fisher 203 Polyclasis LeConte 191
Platycerini 144 Polyderis Motschulsky 29
Platyceropsis Benesh 144 Polydrosus Schdnherr 330
Platycerus Geoffroy 144 Polydrosini 330
Platycis Thomson 189 Polydrusus Germar 330
Platydema Castelnau & Brull6 255 Polygrammo Chewolat 311
Platydracus Thomson 120 Polygraphini 359
Platylomaius Cooman 138 Polygraphus Erichson 359
Platymedon Casey 114 Polyphaga 74
Platynidius Casey 56 Polyphylla Harris 152
Platynini 51 PolyspilaHope 310
Platynomicrus Casey 53 Polystoma Stephens 105
Platynus Bonelli 55 PomphopoeaLeConte 264
Platypatrobus Darlington 29 Pontomalota Casey 103

422
Poophagus Schiinherr 350 Psammoporus Thomson 147
Popillia Audinet-Serville 155 Pselaphidae L25
Porcinolus Mulsant l3z Rey 159 Pselaphinae t29
Poreospasta Horn 2134 Pselaphini 129
Porotachys Netolitzky 29 Pselaphorhynchites Schilsky 327
Porrhodites Kraatz',91 Pselaphus Herbst 129
Potamodytes Zimmermann 66 Psenocerus LeConte 295
Potamonectes Zimmermann 66 Psephenidae 174
Prasocuris Latreille 312 Psepheninae 174
Priacma LeOonte 6 Psephenus Haldeman 174
Priacmini 6 Psephus Kirby 149
Priobium Motschulsl<y 203 PseudalindriaFall 207
Priocera Kirby 209 Pseudamara Lindroth 35
Priognathus LeConte 268 Pseudamphasia Casey 45
Prionellus Casey 277 Pseudanchus Casey 52
Prioninae 277 Pseudanthonomus Dietz 344
Prionini 277 Pseudapion Schilsky 329
Prionochaeta Horn lJl Pseudaplocentrus Noonan 44
Prionocyphon Redterrb acher 142 Pseudargutor Casey 33
Prionomerinae 345 Pseudebaeus Horn 212
Prionomerini 345 Pseudeleusis Bernhauer 95
Prionus Geoffroy 277 PseudelissaCasey 331
Priscosaprinus Werulel 137 Pseudeurostus Heyden 206
Pristodactylo DejeanL 51 Pseudibidion Casey 280
Pristonychus Dejean 51 Pseudobaris LeConte 352
Procalathus Jedlickar 51 Pseudobradyfus Csiki 36
Procas Stephens 331) Pseudocistela Crotch 259
Procryphalus Hopkirrs 362 Pseudocleis Casey 237
Proctorus LeConte l|45 Pseudocoelus Casey 254
Proculini 145 Pseudocryphalus Swaine 358
Progaleritina Jeannr:I 59 Pseudogaurotina Plavilstshikov 290
Prognathus Bertholc[ 92 Pseudohadrotoma Kalik 198
Promecognathini 17 Pseudohylesinus Swaine 357
Promecognathus Chaudoir 17 Pseudolagarus Lutshnik 33
Promecotarsus Casey 340 Pseudolathra Casey 116
Promethister Kryzhanovsky 139 Pseudolesteuo Casey 90
Prometopia Erichsorr 216 Pseudolina Schaeffer 312
Promoresia Sanders,on 173 Pseudoliodini 79
Promus LeConte 256 Pseudolomechusa Mann 106
Pronocera Motschulsky 282 Pseudolucanus Hope & Westwood 144
Proplectus Raffray lL27 Pseudoluperus Beller & Hatch 316
Propylaea Mulsant 236 Pseudomaseus Chaudoir 33
Prorhexius Raffray 126 Pseudomedon Mulsant & Rey 113
Prosaldius Oglobin 353 Pseudomegislo Bernhauer 102
Prostephanus Lesne 200 Pseudonomaretus Roeschke 13
Prostercyon Smetana 134 Pseudopachyla Swaine & Hopping 289
Prostomidae 267 Pseudopanscopus Buchanan 334
Prostomis Latreille 267 Bohart 366
P se udopatella
Prostylops Pierce 3i7 Pseudoperyphus Hatch 21
Proteininae 87 Pseudophonus Motschulsky 40
Proteinini 87 Pseudopityophthorus Swaine 363
Proteinus Latreille 87 Pseudopsinae 95
Prothalpia LeConte 247 Pseudopsis Newman 95
Prothecini 204 Pseudoremus Koch 119
Protonecrodes Porte.yin 85 Pseudorus Casey 114
Psamathobledius Herman 93 Pseudoscuptopterus Hatch 70
Psammodiini 149 PseudosmicronyxDietz 340
Psammodius Fallen 149 Pseudostrangalia Swaine & Hopping 292

423
Pseudota Casey 101 Pycnoplectus Casey 126
Pseudothysanoes Blackman 359 Pycnorus Casey 114
Pseudotrtaeno Minsk & Hatch 39 Pycnotomina Casey 218
Pseudotrox Robinson 150 Pygoleptura Linsley & Chemsak 291
Pseudotychlus Blatchley 328 PyladiaWeise 318
Pseudousipalia Lohse 102 Pyractomena Melsheimer 190
Pseudow eisea Schwarz 229 Pyrectosomo Motschulsky 191
Pseudoxenos Saunders 367 Pyres Champion 258
Pseudoxyletinus Pic 203 Pyrilloxenos Pierce 366
Pseudoxyporus Nakane & Sawada 109 Pyrochroidae 269
Psiloscelis Marseul 140 Pyrochroinae 269
Psoa Herbst 200 Pyrophorinae 175
Psoinae 199 Pyrophorini 175
Psoini 2(t0 Pyropyga Motschulsky 191
Psomophrrs Casey 83 Pytotrichus LeConte 288
Psomus Clasey 348 Pyrrhalta Joannis 314
Psydrus LeConte 29 Pyrrocis Castelneau 243
Psylliodes Berthold 321 Py.thidae 268
Psyllobora Dejean 237 Pybhinae 268
Psylloborini 237 Py'tho Latreille 268
Psyrassa Pascoe 280 Pytna Casey 129
Ptenidium Erichson 75
Pterna Seidlitz 260 Quadrifrons Blatchley 216
Pterocolirrae 326 Quediellus Casey L22
Pterocolus Say 326 Quediini 120
Pteroloma Gyllenhal 77 Quediochrus Casey 121
Pteropalus Casey 43 Quedionuchus Sharp 121
Pterostichini 30 Quedius Stephens 121
Pterostichus Bonelli 31
Pteryngium Reitter 221 Rafonus Casey 126
Ptiliidae 75 Ramecia Casey 126
Ptiliinae 75 RamonaCasey 113
Ptilininae 203 Rantogiton des Gozis 70
Ptilinini 203 Raphirus Stephens 122
Ptilinus I\fiiller 203 Redistes Casey 252
Ptiliola Haldeman 75 Reesa Beal 198
Ptiliolum Flach 75 Refonia Casey 32
Ptiliopycra Casey 76 Reichenbachia Leach 128
Ptilium Gyllenhal 75 Rembus Dejean 48
Ptilodact5'la Illiger 169 Reminius Casey 267
Ptilodacty'lidae 169 Remus Holme 119
Ptinidae 205 Renardia Motschulsky 92
Ptininae 205 Renocis Casey 358
Ptinodes LeConte 202 Reuelstohea Hatch 90
Ptinus Linn6 206 Rhabdophorus Leftvre 308
Ptomaphagini 81 Rhabdopterus Lefbvre 308
Ptomaphagus Illiger 81 Rhadalinae 212
Ptomister Houlbert & Monnot 139 Rhadine LeConte 52
Pubimodes Park I27 Rhaeboscelis LeConte 167
Pullus Mulsant 230 Rhagium Fabricius 289
PunctagonumCasey 53 Rhagomicrus Fleutiaux 187
Purpuricenini 286 RhagonychaEschscholtz 193
Purpuricenus Dejean 286 Rhanidea Strohecker 238
Pycnoglyprta Thomson 88 Rhanis LeConte 238
Pycnomerini 252 Rhantus Dejean 70
Pycnomerus Erichson 252 Rhaphiona Fischer von Waldheim 15
Pycnoruotida Casey 254 Rhaxonycha Motschulsky 193
Pycnophus Casey 83 Rhinocyllini 337

424
Rhinocyllus Germar 337 Salpinginae 270
Rhinoncus Schiinherr 351 Salpingus auct. 270
Rhinosimus Latreille 270 Sandalus Knoch 143
Rhipiceridae 143 Sanfilippody'tes Franciscolo 65
Rhipidandrini 254 Saperda Fabricius 298
Rhipidandrus LeConte 254 Saperdini 298
Rhipiphoridae 250 Sapintus Casey 272
Rhipiphorino Casey 157 Saprininae 136
Rhipiphorinae 250 Saprinus Erichson 136
Rhipiphorus Bosc dAntic 250 Saprobia Ganglbauer 216
Rhizophagidae 218 Saprophilus Streubel 120
Rhizophaginae 218 Sarosesthes Thomson 285
Rhizophagus Herbst 2l-8 Sarpedon Bonvouloir 187
Rhodobaenus LeConte 354 Saxinis Lacordaire 305
Rhombodera Reiche 50 Scalopterus Motschulsky 2\2
Rhombonalia Casey 154 Scaphidema Redtenbacher 254
Rhopalapion Schilsky 329 Scaphidiidae I24
Rhopalelophorus Kuwert 130 Scaphidiini 125
Rhopalomerus Chewol at 285 Scaphidium Olivier 125
Rhopalopaclzys Chevrolat 285 Scaphinotus Dejean 12
Rhynchaeninae 345 Scaphisoma Leach 125
Rhynchaenus Clairville 345 Scaphisomini 125
Rhynchitidae 326 Scaphium l{trby I25
Rhynchitinae 326 Scarabaeidae 145
Rhynchitini 326 Scarabaeinae 146
Rhynchophorinae 354 Scarabaeini 146
Rhyncolini 355 Scarabaeoidea 143
Rhyncolus Germar 356 Scarites Fabricius 17
Rhysodidae 7 Scaritini 17
Rhysothorax Bedel 147 Scelolyperus Crotch 316
Rhyssemus Mulsant 149 Schedarosus Reitter 255
Rhyssomatus Schijnheru 347 aatz lO2
Schistogloss a I{r
Rhytirrhininae 334 Schizogenius Putzeys 18
Rhyzobius Stephens 233 Schizophilus Bonvouloir 188
Rhyzopertha Stephens 200 Schizotus Newman 269
RileyaOlivier 191 Sciaphilus Schiinherr 330
Riponus Casey 278 Scierus LeConte 356
RomaleumWhite 280 SciodrepaThomson 81
Ropalopus Mulsant 284 Sciodrepoides Hatch 81
Rosalia Audinet-Serville 282 Sciopithes Horn 331
Rosaliini 282 Scirtes Illiger 142
Rosella Whitehead 346 Scirtidae 142
Roualida Casey 103 Scleropterini 350
Rugilus Leach 114 Scobicia Lesne 200
Rushia Forel 247 Scoliocerus Motschulsky 178
Rutelini 155 Scolytidae 356
Rutelinae 154 Scolytinae 359
Rutidosoma Stephens 350 Scolytini 359
Rybaxis Saulcy 128 Scolytus Geoffroy 359
Rypobiini 228 Scopaeina 114
Scopaeoma Casey 114
Sachalinobia Jacobson 289 Scopaeopsis Casey 114
Saciinae 228 Scopaeus Erichson 114
Saciini 228 Scopophus Casey 83
Sacium LeConte 228 Scotias Czenpinski 205
Sa.codes LeConte 142 Scotobates Rye 258
Salebius Casey 22I Scotochroa LeConte 247
Salpingidae 270 Scotochroides Mank 247

425
Scotocryptini 79 Sitophilini 355
Scraptia Latreille 276 Sitophilus Schiinherr 355
Scraptiidae 275 Smicronychini 340
Scraptiinae 276 Smicronyx Schdnherr 340
Scraptiini 276 SmiliaWeise 229
Sculptotheca Schilsky 204 Smodicini 280
Scydmaenidae 82 Smodicum Haldeman 280
Scydmaeninae 82 Sonoma Casey 126
Scydmaenrni 83 Sonornota Casey 102
Scydmaenr-rs Latreille 83 Soronia Erichson 216
Scymninaer 229 Spectralia Casey 162
Scymnini 229 Sperchopsis LeConte 132
Scymnobius Casey 231 Sphaeracra Say 56
Scymnus Kugelann 230 Sphaericus Wollaston 205
Scythropus Schtlnherr 330 Sphaeridiinae 133
Selenalius Casey 43 Sphaeridiini 133
Selenophorus Dejean 43 Sphaeridium Fabricius 133
Semanotus Mulsant 283 Sphaeriestes Stephens 270
Semicampa Netolitzky 27 Sphaerites Duftschmid 135
Semiclivina Kult 18 Sphaeritidae 135
Semijulistus Schilsky 212 Sphaeroderus Dejean 12
Sepedophilus Gistel 97 Sphaeromela Bedel 311
Serangiini 229 Sphalera LeConte 248
Serica Ma<:Leay 151 Sphalma Horn 268
Sericini lti1 Sphenophorini 354
Sericoda Kirby 52 Sphenophorus Schiinherr 354
Sericoderinae 228 Sphindidae 213
Sericoderini 228 Sphindinae 213
Sericoderus Stephens 228 Sphindini 213
Sericosomus Dejean 182 Sphindus Chewolat 213
Sericus Eschscholtz 182 Spilodiscus Lewis 140
White 318
Serraticoll;i.s Spilosota Casey 155
Serropalpini 247 SpilotaBurmeister 154
Serropalpus Hellenius 248 Spilotus LeConte 247
Sexarthrus Blatchley 343 Spintherophyta Dejean 308
Siagonium Kirby & Spence 92 Spondylinae 279
Siberia Blzrckwelder 95 Spondylis Fabricius 279
Sibinia Germar 346 Spongoloba Chaudoir 58
SibiotaCasey 101 Spongopus LeConte 45
Sicinus Champion 255 Sponidium Casey 262
Silaria Mulsant 276 Stagetus Wollaston 204
Silini 194 Stamoderes Casey 330
Silis Chartrrentier 194 Staphygraphis Horn 90
Silphidae 84' Staphylinidae 86
Silphinae 85 Staphylininae l1-7
Silusida Casey 107 Staphylinini 120
Silvaninae 220 Staphylinoidea 74
Silvanini 220 Staphylinus Linn6 120
Siluanophloeus Sharp 219 Statira Audinet-Serille 262
Silvanus Latreille 220 Statirinae 262
Simplocaria Stephens 159 SteatoderusDejean 181
Simplocariini 159 Stegobiini 202
Sinistrocedius Park 129 Stegobium Motschulsky 202
Sinodendrini 144 Stelidota Erichson 215
Sinodendron Hellwig 144 StenagriaSharp 105
Sirocalodes Voss 350 Stenelmis Dufour 173
Sitona Germar 331 Stenichnini 83
Sitonini 3ll1 Stenichnus Thomson 83

426
Steninae 109 Stomis Clairville 30
Stenister Reichardt 139 Stornphax Fischer von Waldheim 149
Stenistoderus Jacquelin du Val 123 Storualota Mannerheim 101
Stenocantharus Gistel 13 Storthephora Miiklin 262
Stenocellus Casey 46 Strabala Chewolat 317
StenochiaKirby 261 Strangalepta Casey 293
Stenocorus Geoffroy 289 Strangalia Audinet-Serville 290
Stenocorus Fabricius 288 Strangalina Aurivillius 290
Stenocranophilus Pierce 367 Strepsiptera 366
Stenocrepis Chaudoir 49 Stricticomus Pic 273
Stenolophus Stephens 47 Strigocis Dury 245
Stenopodius Horn 322 Strigoderma Burmeister 155
Stenopterini 282 Strigodermella Casey L55
Stenoscelis Wollaston 355 Strongylaspis Spaeth 323
Stenosphenus Haldeman 280 Stronryliini 261
Stenostrophia Casey 292 Strongylium Kirby 261
Stenotarsinae 238 Strongylocassis Hincks 323
Stenotarsini 238 Strophiona Casey 294
StenothoraxHarris 154 Strophogastra Fenyes 103
Stenotrachelinae 263 Strophosoma Billberg 330
Stenotrachelus Latreille 263 Strophosomini 330
Stenous Chaudoir 49 Stylopidae 367
Stenura Dejean 292 Stylopidia 366
Stenuris Kirby 162 Stylopinae 367
Stenus Latreille 109, 110 Stylops Kirby 367
Stephanocleonus Motschulsky 336 Subhaida Hatch 89
Stephanoderes Eichhoff 362 Sulcacis Dury 245
Stephanopachys Waterhouse 200 Sumitrosis Butte 321
Stephostethus LeConte 240 Sunius Curtis 113
Stereagonum Casey 55 Suphisellus Crotch 62
Steremnius Schtinherr 338 Symbiotes Redtenbacher 238
Stereocerus Kirby 31 Symphora LeConte 247
Stereopalpus LaFert6-S inectdre 27 4 Synaphaeta Thomson 295
Steriphanus Casey 253 Syncalosomo Breuning 10
Sternidius LeConte 297 Syncalyptinae 158
Sternuchus LeConte 141 Synchita Hellwig 251
Steroderus Motschulsky 32 Synchitini 251
Steropus Dejean 32 Synchroa Newman 267
Stethobaris LeConte 353 Synchroidae 267
Stethon LeConte 187 Synchytodes Crotch 251
Stethorhanis Blaisdell 238 Syndesinae 144
Stethorini 229 SynerthaDietz 340
Stethorus Weise 229 Syneta Dejean 307
Sthereus Motschulsky 338 Synetinae 307
Sticholotidinae 229 Synolabus Jekel 326
Stichosoma Motschulsky 311 Synstrophus Seidlitz 246
Stictalia Casey 107 Syntomium Curtis 93
Stictanchus Casey 55 Syntomus Hope 58
Stictocranius LeConte 112 Synuchus Gyllenhal 51
Stictoleptura Casey 293 Systena Chewolat 317
Stictolinus Casey 124
Stigrnatoma LeConte 260 Tachinoderus Motschulsky 96
Stilbolemrna Harris 151 Tachinus Gravenhorst 96
Stilbolidus Casey 45 Tachistodes Casey 48
Stilbus Seidlitz 226 Tachyerges Schiinherr 345
Stilicina 114 Tachymenis Motschulsky 29
Stilicosoma Casey 114 Tachyporinae 96
Stolaini 322 Tachyporini 96

427
Tachyporu.s Gravenhorst 97 Tetrascapha Schuster & Marsh 126
Tachypterellus FaII & Cockerell 343 Tetratoma Fabricius 243
Tachypteru.s Dietz
343 Tetratomidae 243
Tachypus Dejean 20 Tetratominae 243
Tachys Stephens 29 Tetropium Kirby 279
Tachyta Kirby 28 Teuales Casey 90
Tachyura Motschulsky 28 Texania Casey 161
Taenioglyptes Bedel 362 Thalasselephas Egorov & Karotyaev 337
Taenobqtes Motschulsky 260 Thalycra Erichson 216
Tanaocarabus Reitter 12 Thanasimus Latreille 209
Tanaops LeConte 212 Thanatophilus Leach 85
Tanarthropsis Casey 273 Thaneroclerini 209
Tanarthrus LeConte 273 Thecesterninae 335
Tanilotes Casey 275 Thecesternus Say 335
Tanygaster Blatchley 316 Thermonectus Dejean 72
Tanymecirri 331 Thersalus Pascoe 280
Tanymecus Germar 331 Thes Semenov 239
Tanyrhinus Mannerheim 91 Thesalia Casey 292
Tanysphyrini 340 Thesium Casey 126
Tanysphyrus Germar 340 Thicanus Casey 213
Taphranch.us Casey 53 Thinobius Kiesenwetter 94
Taphrocerus Solier 168 Thinodromus Kraatz 94
Taphroscydmus Casey 83 Thinopinus LeConte 120
Tapinosthenes Kolbe 10 Thinusa Casey 108
Taraxis LeConte 304 Thoracophorus Motschulsky 92
Tarpela Bates 258 Thricolepoides O'Brien 330
Tarphiota Casey 103 Thricomigus Horn 333
Tarsosteninae 2ll Thrimolus Casey 243
Tarsostenrrs Spinola 211 Throscidae 186
Taxicerellc,. Casey 100 Throscinae 186
Tecnophilus Chaudoir 59 Throscus Latreille 186
Tectosternum Balfour-Browne 134 Thyamis Stephens 317
Tedinus Crasey 259 Thylodrias Motschulsky 199
Teinodactylo Chevrolat 317 Thylodriini 199
Telacis Poey 259 Thymalus Latreille 207
Telephanus Erichson 220 Thy sanocnemis LeConte 345
Telephorus Schaeffer 193 Tilargus Casey 243
Telmatophilinae 22L TileaFauvel 9O
Telmatophilus Heer 221 Tillini 208
TemnochilaWestwood 207 Tilloclytus Bates 286
Tenebrio L,inn6 259 Tillomorphini 286
Tenebrioni.dae 253 Timarcha Latreille 310
Tenebrioninae 258 Timarchini 310
Tenebrionini 258 Tipnus Thomson 205
Tenebrionoidea 242 Tmesiphorini 129
Tenebroides Piller & Mitterpacher 208 Tmesiphorus LeConte 129
Tenomergil Neboiss 6 Tomq.rus LeConte 224
Tentyriinae 253 Tomicini 357
Teredini 252 Tomoxia Costa 248
Teretriini 136 Toraminae 224
Teretrius Erichson 136 Toramim 224
Tesnus Rey 112 Toramus Grouvelle 224
Tetartoperrs Czwalina 115 Torrentomus Bierig 94
Tetragonorlerus Dejean 56 Tosastes Sharp 334
Tetraleucus Casey 52 Toxidium LeConte 125
Tetralina (lasey 100 Toxotopsis Casey 288
Tetraopes Schtjnherr 299 Toxotropis LeConte 325
Tetraopini 299 Toxotus Dejean 288

428
Trachelonepha Casey 22 Trigonodemus LeConte 91
Trachodes Germar 338 Trigonogenius Solier 205
Trachyinae 168 Trigonorhinus Wollaston 324
Trachykele Marseul 163 Trigonoscuta Motschulsky 330
Trachypachidae 7 Trigonurini 92
Trachypachus Motschulsky 7 Trigonurus Mulsant 92
Trachyphloeini 332 Triliarthrus Casey 46
Trachyphloeus Germar 332 Trimioplectus Brendel 127
Trachysectus Casey 113 Triplacinae 225
Trachysida Casey 292 Triplax Herbst 225
Tragidion Audinet-Serville 286 Triplectrus LeConte 44
Tragosoma Audinet-Serville 278 Trirhabda LeConte 314
Tragosomini 278 Tritoma Fabricius 225
Trechicus LeConte 51 Trixagus Kugelann 186
Trechini 19 Trogastrini 126
Trechonepha Casey 21 Troginae 150
Trechus Clairville 19 Trogoderma Dejean 197
Treiodous Dugds 263 Trogophloeus Nannerheim 94
Trepanedoris Netolitzky 27 Trogossita Olivier 208
Triachus LeConte 307 Trogossitidae 206
Triaena LeConte 39 Trogossitinae 207
Triarthron Markel 78 Trogossitini 207
Tribalinae 138 Trogoxylon LeConte 201
Triboliini 258 Tropicini 170
Tribolium Macleay 258 Tropicus Pacheco 170
Tribonia Rivalier 8 Tropideres Sch0nherr 325
Tribrachys LeConte 2I4 Tropiderini 325
Trichalophus LeConte 333 Tropiphorus Schiinherr 334
Trichapion Wagner 329 Tropisternus Solier 133
Trichelaphrus Semenov 16 TrotomrnaLeConte 247
Trichesthes Erichson 152 Trox Fabricius 150
Trichiinae 157 Trymochthebius Kuweft 7 4
Trichiini 158 Trypherus LeConte 195
Trichiotinus Casey 158 Trypodendron Stephens 361
Trichischius LeConte 354 Trypophloeus Fairmaire 362
Trichiusa Casey 103 Turboscymnus Gordon 231
Trichobaris LeConte 352 Tychiinae 345
Trichocanthus Motschulsky 120 Tychiini 346
Trichocellus Ganglbauer 46 Tychini 129
Trichochrous Motschulsky 2I3 Tychius Germar 346
Trichocnemis LeConte 27 7 Tychobythinus Ganglbauer l-28
Trichodes Herbst 210 Tylicini 160
Trichodesma LeConte 202 Tylicus Casey 160
Tricholochmaea Laboissidre 314 Tylistus LeConte 204
Trichophya Mannerheim 99 Tyloderma Say 347
Trichophyinae 99 Tylonotus Haldeman 280
Trichoplataphus Netolitzky 23 Tympanophorus Nordmann 120
Trichopteryx Kirby & Spence 76 Typhaea Stephens 243
Trichoscelis Bates 286 Typhloleiodes Hatch 80
Trichotichnus Morawitz 43 Typitium Casey 262
Tricoryninae 205 Typocerus LeConte 291
Tricorynini 205 Tyrini 129
Tricorynus Waterhouse 205 TyrusAub6 129
Tricrania LeConte 265
Tricraniodes Wellman 265 Uleiota Latreille 220
TrigaFauvel 95 Uleiotini 220
Trigites Handlirsch 95 Ulitusa Casey 107
Trigonarthris Haldeman 293 Ulochaetes LeConte 288

429
Uloma Ltrtreille 257 Xyleborinus Reitter 362
Ulomini 257 Xyleborus Eichhoff 362
Unamis Casey 90 Xylechinus Chapuis 357
Upis Fabricius 261 Xyletininae 203
Urgleptes Dillon 298 Xyletinini 203
Urograph',is Horn 297 Xyletinus Latreille 204
Uroplatini 322 Xylita Paykull 247
Utobium FalI 202 Xylobiops Casey 200
LJvarus Guignot 64 Xylobius Latreille 188
Xylocomesus Thatcher 356
Vacusus Casey 273 Xylocrius LeConte 284
Vanonus Casey 275 Xylodromus Heer 91
Variopalpis Solier 58 Xyloecus Dejean 188
Vellica Casey 90 Xyloperthini 200
Venusa Casey 107 Xylophilus Mannerheim 188
Veraphis Casey 83 Xylopinus LeOonte 260
Vicelva l\:[oore & Legner 95 Xyloryctes Hope 156
Vincenze.llusReitter 270 Xylosandrus Reitter 361
Vrilletta LeConte 203 Xyloterini 361
Xyloterinus Swaine 361
WillinkiaBondar 353 Xylotrechus Chevrolat 285
Xanthochroa Schmidt 266 Yrdops Steffan 171
Xanthochroina Ganglbauer 266
Xanthoga.leruca Laboissidre 314 Zabrini 35
Xantholirrinae 123 Zacotus LeConte 19
Xantholinini 123 ZaitzeviaChampion 173
Xantholirrus Dejean 124 ZalepiaArnett 175
Xanthonia Baly 309 Zalobius LeConte 95
Xanthoph,ius Motschulsky 123 Zeadolopus Broun 79
Xanthopl'gini 120 Zenodosus Wolcott 209
Xanthus Dietz 344 Zeugophora Kunze 304
Xeninae 367 Zeugophorinae 304
Xenochara Mulsant & Rey 104 ZezeaCsiki 39
Xenocharis Bierig 113 ZiloraMulsant 247
Xenodacne Boyle 225 ZonantesCasey 275
Xenodusa Wasmann 106 Zonitina 265
Xenoscelinae 224 Zonltinae 265
Xenota Mulsant & Rey 103 Zonitini 265
Xerosaprinus Wenzel 137 Zonitis Fabricius 265
XestipygerMarseul 138 Zonoptilus Solsky 93
Xestobiini 202 Zopheridae 252
Xestobiurn Motschulsky 202 Zophefinae 252
Xestocis Casey 244 Zuphiini 59
Xestoleptura Casey 293 Zuphium Latreille 59
Xestolinus Casey 124 Zygogramma Chewolat 3 10
Xestonotrrs LeConte 45 Zygopinae 348
Xestophu;s Casey 83 Zygopini 348
Xyleborini 361 ZyrasStephens 106

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430
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