Annotated Checklist of The Rotifers Phylum Rotifer

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Annotated checklist of the rotifers (Phylum Rotifera), with notes on

nomenclature, taxonomy and distribution
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DOI: 10.11646/zootaxa.1564.1.1
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ZOOTAXA
1564
Annotated checklist of the rotifers (Phylum Rotifera),

with notes on nomenclature, taxonomy and distribution
HENDRIK SEGERS
Magnolia Press
Auckland, New Zealand
HENDRIK SEGERS
Annotated checklist of the rotifers (Phylum Rotifera), with notes on nomenclature, taxonomy
and distribution
(Zootaxa 1564)
104 pp.; 30 cm.
31 August 2007
ISBN 978-1-86977-129-4 (paperback)
ISBN 978-1-86977-130-0 (Online edition)
FIRST PUBLISHED IN 2007 BY

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Zootaxa 1564: 1–104 (2007) ISSN 1175-5326 (print edition)
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Copyright © 2007 · Magnolia Press
ZOOTAXA
ISSN 1175-5334 (online edition)
Annotated checklist of the rotifers (Phylum Rotifera),

with notes on nomenclature, taxonomy and distribution
HENDRIK SEGERS
Belgian Biodiversity Platform, Freshwater Laboratory, Royal Belgian Institute of Natural Sciences, Vautierstraat 29, B - 1000 Brus-
sels Belgium. E-mail: Hendrik.segers@naturalsciences.be
Table of contents
Abstract . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3
Introduction . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3
Approach . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4
Scope . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4
Future prospects . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 9
Acknowledgements . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 9
References . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 10
Abstract
Phylum Rotifera comprises about 2030 known species classified in three main groups, the marine Seisonida (3 species),
the Monogononta (1570 species) and the unique, exclusively parthenogenetic Bdelloidea with 461 clonal species. Here I
present an annotated checklist of the taxon, giving the global distribution of the valid species. This checklist contains
over 3,000 available names at the genus- and species-group level, including valid names and a non-comprehensive list of
synonyms. A number of taxonomic and nomenclatural suggestions following from the review is presented. Suggestions
include several new synonyms and two nomina nova: Dissotrocha kostei for Dissotrocha aculeata Koste, 1996 non
Ehrenberg, 1838 (stat. nov., ex. Dissotrocha hertzogi aculeata) and Lepadella zigzag for Lepadella bractea Myers, 1934
non (Ehrenberg, 1838). This contribution also describes features of an on-line, global names database of the Rotifera, on
which this checklist is based.
Key words: biodiversity, biogeography, Rotifera
Introduction
In recent years science has witnessed a rapid evolution from print to electronic media in the production of tax-
onomic resources. This inevitable evolution (see Godfray, 2002) is fostered by a large number of initiatives
like the Global Biodiversity Information Facility (GBIF, http://www.gbif.org/), the Integrated Taxonomic
Information System (ITIS, http://www.itis.usda.gov/), Species2000 (http://www.sp2000.org), and Universal
Biological Indexer and Organizer (uBIO: http://www.ubio.org), to name a few. Even long-established
resources like the International Commission on Zoological Nomenclature (ICZN, 1999; see http://
www.iczn.org/) are turning towards the World Wide Web, (e.g., with the ZooBank initiative: http://
www.zoobank.org/; Polaszek, 2005) notwithstanding the obvious difficulty the ephemeral character of elec-
tronic media pose to a stable scientific nomenclature.
Accepted by R. Dalgleish: 26 Apr. 2007; published: 31 Aug. 2007 3

Acknowledging this development, I present a checklist of the Rotifera of the world, extracted from a data-
base compiled in the framework of the project `Freshwater Animal Diversity Assessment' (see http://
FADA.biodiversity.be).
In addition to the checklist I provide a short overview of a number of features of the database and some
taxonomic and nomenclatural changes resulting from the literature review required to prepare the list. The
present paper, together with the analysis of the diversity and biogeography of the Rotifera (Segers, in press),
are intended as citable source for the database.
Approach
The checklist is based on the taxonomy of rotifers as presented in monographs listed in Table 1, and numerous
recent publications. For practical reasons, only names or taxonomic changes published before November 1,
2006 are included.
Considering that subsequent lumping of distributional data is easy whereas the inverse is not, a splitting
rather than lumping approach to taxonomy has been adopted. As a consequence, the diversity of rotifers turns
out to be higher than previous estimates. This is especially so for genera like Lepadella and Lecane. In Lep-
adella, the lack of any comprehensive revision is responsible for the discrepancy, while Lecane is exemplary
of the tendency towards a less inclusive species concept in Rotifera during recent years. This approach has
been corroborated by recent research using molecular (e.g., Goméz et al., 2002) or detailed morphological
techniques (Giri & José de Paggi, 2006), or combining different approaches (Ricci, 2001) and biogeography
(Segers & Shiel, 2003), showing that diversity has generally been underestimated in Rotifera. On the other
hand, many rotifer taxa exhibit wide morphological variability, either as a result of phenotypic plasticity and/
or distortion during fixation or dehydration. Although this has long been known and is well documented (see
Dumont, 1983; Koste & Shiel, 1989c; Segers, 1998), variants are still frequently being described as subspe-
cies of common species. Such taxa, together with insufficiently described ones, are included in the present
checklist as their names are available, but are tagged species inquirenda in the absence of a critical taxonomic
revision. Names listed as referring to unrecognizable taxa in recent revisions listed in Table 1 have not been
included. The taxonomy reflected in the list may not be satisfactory, especially in taxa for which no recent
comprehensive reviews are available (e.g., Brachionidae, Lepadellidae, and Notommatidae).
The treatment of synonyms in the database is not comprehensive, but focuses on synonyms that occur in
recent literature or that are occasionally listed as valid taxa. Older synonyms (especially in the morphologi-
cally variable Brachionidae: Brachionus and Keratella) can be found in Harring (1913) and Wiszniewski
(1954). When necessary, care was taken to correct names in accordance with the International Code of Zoo-
logical Nomenclature (ICZN, 1999). Incorrect subsequent spellings are generally not taken into account
unless they have some nomenclatural significance beyond their mere existence: keeping track of published
spelling variants and nomina nuda in the vast ecological literature on rotifers is a daunting task with dubious
relevance.
Data on distribution are presented for taxa considered valid only. This information is mostly drawn from
the literature as in Table 1 and, as much as feasible, included only after critical assessment. A more complete
analysis of the biodiversity and distribution of the rotifers, based on the data presented here, is as in Segers (in
press).
Scope
The checklist and database, in its present version, contain 2127 species-level taxa of Rotifera, of which 109
are presently considered species inquirenda, and some 800 synonymous or alternative genus- and species-

level names. The vast majority of the valid names belong to Monogononta (1566), whereas Bdelloidea (461)
and, especially, Seisonida (3) are much less diverse. The following information is provided:
Valid names of genus- and species-level taxa with authority, and the original combination of species-level
names. Genera and species within genera are arranged alphabetically in the checklist in order to facilitate
information retrieval. This ordering is consistent with Harring (1913) and Wiszniewski (1954).
Names with authority of synonyms of available genus-and species-level names, the latter in their original
combination. The synonymy is incomplete for taxa in which a high number of older synonyms exist. For these
I refer to Harring (1913) and Wiszniewski (1954).
Names with authority of some species inquirendae not included in recent revisions (see Table 1).
Taxonomic comments. This includes indication of taxonomically problematic cases, or instances where
synonyms are in use as names of infrasubspecific variants. Unavailable names of infrasubspecific taxa are
added in comments, except for those in genera like Brachionus and Keratella, where citing the exceedingly
high number of names of variants, forms etc. would only add to confusion.
Occurrence in eight large biogeographical regions (Palearctic, Afrotropical, Oriental, Nearctic, Neotropi-
cal, Australian, Pacific and Antarctic), as circumscribed in Martens et al. (in press; see Figure 1), for all valid
and recognisable species-level fresh- and/or brackishwater taxa. Marine, parasitic and interstitial taxa are
mentioned as such. If relevant, a comment on distribution is added, mostly indicating that a regional record of
a species is dubious, or that such a record may result from species introduction.
FIGURE 1. Definition of the eight regions used in this work (after Martens et al., in press.)
In addition, the database includes a higher-level classification of the free-living rotifers, after Segers
(2002) and Wallace et al. (2006). The Acanthocephala, of which several authors argue that they are highly
adapted parasitic representatives of the group (Mark Welch, 2000; see also Garcia-Varela et al., 2000; Giribet
et al., 2000), are excluded.
ANNOTATED CHECKLIST OF THE ROTIFERS Zootaxa 1564 © 2007 Magnolia Press · 5

The goal of the databasing exercise was not to introduce taxonomic or nomenclature novelties. However,
a number of instances was found in which a change in taxonomy appeared warranted. In addition, some actual
or potential homonyms were found. Many of the latter concern cases of congeneric, but not conspecific, sub-
species with identical subspecies names. Because of the principle of coordination in zoological nomenclature,
such cases are better avoided, or at least monitored. Especially, the subspecies rank of many rotifer subspecies
is exclusively based on the consideration that there is a morphological resemblance between taxa. We now
know that cryptic speciation occurs, and may be common, in Rotifera; hence such arguments for attributing
subspecific rank are losing what little ground they ever had. Moreover, there are indications that what were
once thought to be mere infrasubspecific variants could be taxonomically relevant entities (e.g., Giri & José
de Paggi, 2006). Here, replacement names are only suggested for those junior homonyms that denote valid
taxa. Other cases are listed as caution against potential future confusion.
The following are some suggestions on taxonomy and nomenclature:
Anchitestudinella mekongensis B‘rziÃÓ, 1973
B‘rziÃÓ (1973) contains two alternative spellings for the single species of Anchitestudinella: mekogensis
and mekongensis. Subsequent authors have used either spelling, but none has ever clarified which of the
two should be considered the correct spelling. Considering the etymology of the name, which is derived
from River Mekong, I suggest treating the latter, with -n-, as correct original spelling under Art. 24.2.3 of
the ICZN.
Brachionus inermispinosus De Smet, 1989 = Epiphanes brachionus spinosa Rousselet, 1901 (new synonym).
The detailed description and, especially, figures of the habitus and trophi of B. inermispinosus by De Smet
(1989) reveal that the animal does not belong in Brachionus; rather, it is an Epiphanes and, most likely,
synonymous with E. brachionus spinosa.
Brachionus anchorporus Xu, Chen & Rao, 1997 = B. durgae Dhanapathi, 1974 (new synonym).
The description by Xu et al. (1997) undoubtedly depicts B. durgae, a taxon of which these authors were
apparently unaware.
Brachionus angularis orientalis Sudzuki, 1989 = B. angularis Gosse, 1851
The name B. angularis orientalis is a potential junior homonym of B. orientalis Rodewald, 1937, if the
name would ever be used at the species level. Sudzuki’s (1989) species, however, is based on a relatively
common variant of B. angularis and is treated as a synonym rather than a different subspecies. This is in
agreement with Sudzuki’s (1998) treatment of the taxon as of infrasubspecific rank, although he reverts to
listing it as a subspecies in Sudzuki (1999).
Brachionus angularis pseudokeikoa Sudzuki, 1992; B. angularis pyriformis Sudzuki & Huang, 1997; B.
angularis donghuensis Sudzuki & Huang, 1997; B. morondavaensis Sudzuki, 1998 = B. angularis Gosse,
1851 (new synonyms)
Sudzuki (1999) cites the three subspecies listed above as species, in which case Sudzuki & Huang’s
(1997) B. pyriformis would have to be treated as a junior homonym of B. pyriformis Barrois & Daday,
1894, an established junior synonym of B. angularis. All four fall within the range of variability of B.
angularis; B. angularis donghuensis is a poorly contracted specimen of B. angularis bidens, according to
a photograph in Sudzuki (1999). A fifth taxon, B. angularis daitojimensis Sudzuki, 1992 is listed as an
infrasubspecific variant, hence a synonym, of B. angularis by Sudzuki (1999) himself.

B. dimidiatus isigakiensis Sudzuki, 1992; B. cluniorbicularis isigakiensis Sudzuki 1992 and B. durgae isigak-
iensis Sudzuki 1992
Sudzuki (1992) named three new Brachionus subspecies “isigakiensis”. All three are easily recognized as
belonging to previously established taxa, as far as can be judged from the relevant descriptions and photo-
graphs in Sudzuki (1992, 1999):
B. dimidiatus isigakiensis Sudzuki, 1992 (Brachionus isigakiensis in Sudzuki 1998, 1999) is a poorly
contracted B. bennini Leissling (new synonym).
B. cluniorbicularis isigakiensis Sudzuki 1992 is an unspined variant (f. cluniorbicularis) of B.

quadridentatus Hermann (new synonym).
B. durgae isigakiensis Sudzuki 1992 falls within the range of variability of the nominal species, hence
it is here considered a synonym of B. durgae Dhanapathi (new synonym).
As all three enter into synonymy, there appears to be no imminent danger of nomenclatural confusion
between the three potential homonyms.
Brachionus brevispinus Sudzuki & Huang, 1997 = B. huangi Zhuge & Koste, 1996 (new synonym).
Although the taxonomic separation of the short-spined B. huangi and B. brevispinus versus B. diversicor-
nis Daday may require substantiation, B. huangi and B. brevispinus are obviously identical. As B.
brevispinus Sudzuki & Huang, 1997 enters into synonymy, there is no danger of confusion with its senior
homonym B. brevispinus Ehrenberg, 1832, itself a synonym, and representing a well-known and common
infrasubspecific variant, of B. quadridentatus Hermann.
Brachionus lyratus yonaguniensis Sudzuki, 1992 = B. angularis Gosse, 1851 (new synonym).
This taxon, listed as B. yonaguniensis by Sudzuki (1999), appears to be yet another variant of B. angu-
laris. An appurtenance to B. lyratus is not evident from the available description and photographs.
Brachionus variabilis kasumigaurensis Sudzuki, 1998 = B. rubens Ehrenberg, 1838 (new synonym).
The available pictures of B. variabilis kasumigaurensis (see Sudzuki, 1999), especially the shape of
antero-dorsal spines and minute size of the rounded process on the dorsal foot aperture margin reveal a
synonymy with B. rubens.
Encentrum grande (Western, 1891) and Encentrum kulmatyckii (Wiszniewski, 1953)
De Smet in De Smet & Pourriot (1997) argued that the description of Encentrum grande (Western) is
unrecognisable, and proposed to refer to Harring & Myers’ (1928) description as reference for the taxo-
nomic identity of E. grande. Considering the poor quality of many original descriptions of Rotifera and
the absence of type material, this is common practice in rotifer taxonomy. However, he further retained E.
grande (Western) as species inquirenda, and included a E. grande Harring & Myers, 1928 in his treat-
ment. This confusingly results in the recognition of two homonymous taxa, one E. grande (Western) and
E. grande sensu Harring & Myers (1928) which, in fact, should be attributed to De Smet (1997). The
name Encentrum kulmatyckii (Wiszniewski, 1953), a junior synonym of E. grande sensu Harring &
Myers (1926), is available for the latter taxon.

Dissotrocha hertzogi aculeata Koste, 1996 versus Dissotrocha aculeata aculeata Ehrenberg, 1838
Koste’s (1996) taxon has distinct body outgrowths (none in D. hertzogi), and lacks the extraordinarily
long rostrum ciliae and offset spur tips of Dissotrocha hertzogi (see Donner, 1965). These morphological
differences are of a nature and magnitude that is generally considered relevant at the species level in bdel-
loid rotifer taxonomy. In addition, it is particularly difficult to conform a species, let alone a subspecies,
definition to an exclusive parthenogenetically reproducing taxon. I therefore suggest treating the two as
separate species-level taxa and, because of the homonymy of Koste’s name with Ehrenberg’s, I have the
pleasure to suggest the name Dissotrocha kostei, honouring Dr h.c. W. Koste, as replacement name for the
taxon (nomen novum).
Keratella heywoodi Dartnall, 2005 = K. kostei Paggi, 1981 (new synonym)
Both Dartnall’s (2005) and Paggi’s (1981) descriptions point to the absence of a foundation pattern on the
dorsal lorica, small size of antero-median and antero-intermedian, but large and ventrally-pointed antero-
lateral spines as characteristic features of their new species. As Dartnall (2005) does not mention K. kostei
in his text, he may not have been aware of the existence of that name. The species occurs in Patagonia,
Argentina (Paggi, 1981), and on the Falkland Islands and South Georgia (Dartnall, 2005).
Pompholyx acutangula Wu, 1981 = Testudinella brevicaudata Yamamoto, 1951 (new synonym).
Wu’s (1981) clear and detailed illustrations depict an animal with a large foot aperture. It therefore does
not belong to Pompholyx, rather it is a previously described Testudinella, T. brevicaudata.
Testudinella tridentata obtusa Wu, 1981 = Testudinella greeni Koste, 1981 (new synonym).
The figures and photographs of Wu’s T. tridentata obtusa unmistakably depict a T. greeni. Apparently, Wu
(1981) was not aware of the latter species.
Hemimonostyla blachei tutiurensis Sudzuki, 1998 = Lecane stenroosi Meissner, 1908 (new synonym).
The photograph of H. blachei tutiurensis by Sudzuki (1998) leaves no doubt as to its synonymy with L.
stenroosi.
Lecane kasumiensis Sudzuki, 1998 = Lecane papuana (Murray, 1913) (new synonym).
Again, the photograph of L. kasumiensis by Sudzuki (1998) leaves no doubt as to its synonymy with L.
papuana.
Lepadella bractea Myers, 1934 versus Squamella bractea Ehrenberg, 1838.
Myers’ name is threatened by its homonymy with Squamella bractea Ehrenberg, an established junior
synonym of Lepadella patella Müller, 1786. Here, a replacement name is in order as confusion between
the two has occurred (Nogrady in Boyko, 1994), notwithstanding that Ehrenberg’s name is hardly ever
found in combination with the genus name Lepadella. Myers’ (1934) animal is a remarkable but rare
Northeast North American endemic that is unmistakable by the prominent zig-zag crest on its dorsal lor-
ica. I suggest the name Lepadella zigzag as nomen novum for the taxon.

Lepadella parvula De Graaf, 1956 versus L. parvula Bryce, 1893
Lepadella parvula De Graaf, 1956 is a junior homonym of Lepadella parvula Bryce, 1893 and of the pre-
viously replaced L. parvula Montet, 1915 (valid name: Lepadella minuta Weber & Montet, 1919). As
Lepadella parvula De Graaf is an established junior synonym of Lepadella patella Müller, there is no
imminent need for a replacement name.
Macrotrachela ligulata Haigh, 1965; Macrotrachela nana ligulata Schulte, 1954; Macrotrachela quadri-
cornifera ligulata B‘rziÃÓ, 1950
Awaiting a revision of the large and complex genus Macrotrachela, I merely point out that Macrotrachela
quadricornifera ligulata B‘rziÃÓ, 1950 and M. nana ligulata Schulte, 1954 are potential senior hom-
onyms of M. ligulata Haigh, 1965, if any or both of the first-mentioned ones would ever be raised to spe-
cies rank.
The genus Diplois Gosse, and its single species D. daviesae Gosse are here considered genus et species
inquirendae. Scrutiny of the available literature on the taxon, and consultation with colleagues revealed that
only Gosse’s description exists, and that there are no reliable records or sightings of the species (W.H. De
Smet, in lit., R.J. Shiel, in lit., C. Jersabek, in lit.). The description of Pseudoembata parasitica by Wycliffe &
Michael (1968) does not enable recognition of the taxon; the arguments for erection of a new genus are not
supported in the literature (see Donner, 1965). Also this taxon is therefore considered genus et species
inquirendae.
Future prospects
The database mentioned here will be made freely accessible through the World Wide Web. In a first phase,
researchers will be offered the possibility to select and extract a structured name list from the database. By this
service, it is hoped to reduce the number of misspellings and other inconsistencies in the published literature.
In a second phase, the possibility will be offered to submit and save regional or local (georeferenced)
checklists to the database, together with metadata on these checklists. One of the goals of the database is to
provide a resource to as reliable as possible data on the distribution of Rotifera. Hence, we envisage installing
a system comparing submitted lists with reviewed distributional data so that new regional species records will
be highlighted and, eventually, reviewed by peers. This review process may involve submitting material
enabling the positive identification of the species concerned and, if this is possible by the nature of the taxon,
by illustrating remarkable records through relevant photographs or original drawings on the web site.
Acknowledgements
The present work was done as part of the project “Freshwater Animal Diversity Assessment”, funded by the
Belgian Science Policy Office, and benefited greatly from a previous compilation of a European species regis-
ter under the EU-funded (FP5) Fauna Europaeae project. The Belgian Biodiversity Platform (www.biodiver-
sity.be), especially A. Heughebaert, and N. Youdjou, transposed information from a spreadsheet to a database
and to structured text, performed quality control and developed a web site and portal for the database.
I’m indebted to R.L. Wallace and an anonymous referee for reviewing the manuscript of this paper, and to
W.H. De Smet and, especially, C. Jersabek for suggestions and corrections to the list.

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Table 1. List of major literature resources consulted.

Higher-level classification: Segers (2002); Wallace et al. (2006).
Seisonidae: Sørensen et al. (2005)
Bdelloidea: Donner (1965) (taxonomy), B‘rziÃÓ (1982) (Dissotrocha aculeata group), Fontaneto et al. (in
press) and Ricci et al (2003) (distribution).
Monogononta: Koste (1978) (taxonomy), De Ridder (1986, 1991, 1993), De Ridder & Segers (1997) and Jers-
abek (2003) (distribution).
Asciaporrectidae: De Smet (2006)

Asplanchnidae: José de Paggi in Nogrady & Segers (2002)
Brachionidae: Koste (1978), Koste & Shiel (1987). Also: Ahlstrom (1940, 1943)
Clariaidae : Kutikova et al. (1990)
Conochilidae: Segers & Wallace (2001)
Dicranophoridae: De Smet & Pourriot (1997)
Epiphanidae: Koste & Shiel (1987)
Euchlanidae : Koste (1978), Koste & Shiel (1989a)
Gastropodidae: Radwan & Bielanska-Grajner in Nogrady & Segers (2002)
Ituridae: De Smet & Pourriot (1997)
Lecanidae: Segers (1995a, 1995b)
Lepadellidae: Koste & Shiel (1989b)
Lindiidae: Segers in Nogrady & Segers (2002)
Microcodidae: De Smet in Nogrady & Segers (2002)
Mytilinidae: Koste & Shiel (1989a)
Notommatidae: Nogrady et al. (1995)
Proalidae: De Smet (1996)
Scaridiidae: Segers in Nogrady et al. (1995)
Synchaetidae: Hollowday in Nogrady & Segers (2002)
Trichocercidae: Koste (1978) Segers (2003)
Trichotriidae: Koste & Shiel (1989a)
Trochosphaeridae: Segers in Nogrady & Segers (2002), Sanoamuang in Nogrady & Segers (2002)

Table 2. Checklist of the Rotifera
Legend: AFR: Afrotropical region; ANT: Antarctic region; AUS: Australian region; NEA: Nearctic
region; NEO: Neotropical region; ORI: Oriental region; PAC: pacific region; PAL: Palearctic region
(see Figure 1).
Abrochtha Bryce, 1910
Abrochtha carnivora Ricci, Melone & Walsh, 2001: NEO

Abrochtha intermedia (de Beauchamp, 1909): NEA, PAL
Philodina intermedia de Beauchamp, 1909
Acyclus Leidy, 1882
Hyalocephalus Lucks, 1911
Acyclus inquietus Leidy, 1882: NEA, ORI, PAL

Acyclus trilobus (Lucks, 1911): PAL
Hyalocephalus trilobus Lucks, 1911
Adineta Hudson & Gosse, 1886
Adineta acuticornis Haigh, 1967: AUS

Adineta barbata Janson, 1893: AFR, ANT, AUS, NEA, NEO, PAL
Adineta bartosi Wulfert, 1960: PAL
Adineta cuneata Milne, 1916: AFR, AUS, NEA, PAL
Adineta elongata Rodewald, 1935: PAL
Adineta glauca Wulfert, 1942: AUS, PAL
Adineta gracilis Janson, 1893: AFR, ANT, AUS, NEA, ORI, PAL
Adineta grandis Murray, 1910: AFR, ANT
Adineta longicornis Murray, 1906: AFR, ANT, AUS, NEA, NEO, ORI, PAC, PAL
Adineta oculata (Milne, 1886): NEO, PAL
Callidina oculata Milne, 1886
Syn.: Adineta alluandi Certes, 1903
Adineta ricciae Segers & Shiel, 2005: AUS
Adineta steineri Bartoš, 1951: ANT, AUS, NEA, NEO, PAL
Adineta tuberculosa Janson, 1893: AFR, AUS, PAL
Adineta vaga (Davis, 1873)
Callidina vaga Davis, 1873
Adineta vaga major Bryce, 1893: NEA, ORI, PAL
Adineta vaga minor Bryce, 1893: AUS, NEA, ORI, PAL
Adineta vaga rhomboidea Bērziņš, 1950: PAL
Adineta vaga tenuicornis Bryce, 1926: PAL
Adineta vaga vaga (Davis, 1873): AFR, ANT, AUS, NEA, NEO, ORI, PAL

Albertia Dujardin, 1838
Albertia crystallina Schultze, 1851 (Endoparasitic in marine and brackish water Oligochaeta)
Albertia naidis Bousfield, 1886 (Endoparasitic in freshwater Oligochaeta)
Syn.: Albertia bernardi Hlava, 1904
Syn.: Albertia caudata Manfredi, 1927
Syn.: Albertia intrusor Gosse, 1886
Syn.: Albertia soyeri Coineau & Kunst, 1964
Albertia ovagranulata Valovaya, 1991 (Endoparasitic in marine Oligochaeta)
Albertia reicheltae Koste, 1970 (Endoparasitic in freshwater Oligochaeta)
Albertia typhlina Harring & Myers, 1928 (Ecology unknown)
Albertia vermiculus Dujardin, 1938 (Endoparasitic in terrestrial Oligochaeta and Mollusca)
Albertia woronkowi Zenkewitsch, 1922 (Endoparasitic in freshwater Oligochaeta; Baikal endemic)
Anchitestudinella Bērziņš, 1973
Anchitestudinella mekongensis Bērziņš, 1973: ORI
Anomopus Piovanelli, 1903
Anomopus chasmagnathi Mane-Garzon & Montero, 1973: NEO

Anomopus telphusae Piovanelli, 1903: AFR, PAL
Anuraeopsis Lauterborn, 1900
Anuraeopsis coelata de Beauchamp, 1932: AFR, AUS, NEA, NEO, ORI, PAL
Syn.: Anuraeopsis lata singapurensis Sudzuki, 1989
Anuraeopsis coelata coelata de Beauchamp, 1932
Anuraeopsis coelata kehjengi Sudzuki, 1989 (species inquirenda)
Anuraeopsis cristata Bērziņš, 1956: ORI
Anuraeopsis fissa Gosse, 1851: AFR, AUS, NEA, NEO, ORI, PAL
Syn.: Anuraeopsis hypelasma Gosse, 1886
Anuraeopsis fissa beauchampi Bērziņš, 1962 (species inquirenda)
Anuraeopsis fissa donghuensis Sudzuki & Huang, 1997 (species inquirenda)
Anuraeopsis fissa fissa Gosse, 1851
Anuraeopsis fissa haueri Bērziņš, 1962 (species inquirenda)
Anuraeopsis fissa neali Bērziņš, 1962 (species inquirenda)
Anuraeopsis fissa pseudonavicula Sudzuki & Huang, 1997 (species inquirenda)
Anuraeopsis lata Bērziņš, 1962 (species inquirenda)
Anuraeopsis miracleae Koste, 1991: PAL
Anuraeopsis navicula Rousselet, 1911: AUS, NEA, NEO, ORI, PAL
Syn.: Anuraeopsis congolensis Evens, 1947
Syn.: Anuraeopsis racenisi Bērziņš, 1962
Anuraeopsis navicula berzinsi Sudzuki, 1998 (species inquirenda)
Anuraeopsis navicula navicula Rousselet, 1911
Anuraeopsis quadriantennata (Koste, 1974): NEO

Paranuraeopsis quadriantennata Koste, 1974
Anuraeopsis siolii Koste, 1972: NEO
Anuraeopsis urawensis Sudzuki, 1957: NEO, PAL
Anuraeopsis wulferti Sudzuki, 1998 (species inquirenda)
Asciaporrecta De Smet, 2006
Asciaporrecta arcellicola De Smet, 2006: NEA, PAL

Asciaporrecta difflugicola De Smet, 2006: NEA, PAL
Asciaporrecta hyalina (Wulfert, 1939): PAL
Pleurotrocha hyalina Wulfert, 1939
Ascomorpha Perty, 1850
Chromogaster Lauterborn, 1893

Sacculus Gosse, 1851
Ascomorpha agilis Zacharias, 1893: AFR, PAL

Ascomorpha dumonti De Smet, 1992: AFR
Ascomorpha ecaudis Perty, 1850: AFR, AUS, NEA, ORI, PAL
Syn.: Ascomorpha germanica Leydig, 1854
Syn.: Ascomorpha helvetica Perty, 1852
Syn.: Sacculus viridis Gosse, 1851
Ascomorpha klementi (Hauer, 1965): NEO
Chromogaster klementi Hauer, 1965
Ascomorpha minima Hofsten, 1909: NEA, PAL
Ascomorpha ovalis (Bergendal, 1892): AFR, AUS, NEA, NEO, ORI, PAL
Anapus ovalis Bergendal, 1892
Syn.: Sacculus cuirassis Hood, 1894
Syn.: Chromogaster testudo Lauterborn, 1893
Ascomorpha saltans Bartsch, 1870
Ascomorpha saltans indica Wulfert, 1966: AUS, NEO, ORI, PAL
Ascomorpha saltans saltans Bartsch, 1870: AFR, AUS, NEA, NEO, ORI, PAL
Syn.: Sacculus hyalinus Kellicott, 1888
Ascomorpha tundisii Segers & Dumont, 1995: NEO, PAL
Ascomorphella Wiszniewski, 1953
Hertwigella Voigt, 1957

Hertwigia Plate, 1886 non Schmidt, 1880
Ascomorphella volvocicola (Plate, 1886): AFR, AUS, NEA, ORI, PAL

Hertwigia volvocicola Plate, 1886
Syn.: Proales parasita Gosse, 1852 (non Ehrenberg (1838))

Aspelta Harring & Myers, 1928
Aspelta alastor Harring & Myers, 1928: NEA, PAL

Aspelta angusta Harring & Myers, 1928: NEA, PAL
Aspelta aper (Harring, 1913): AUS, NEA, NEO, PAL
Encentrum aper Harring, 1913
Aspelta beltista Harring & Myers, 1928: NEA
Aspelta bidentata Wulfert, 1961: PAL
Aspelta chorista Harring & Myers, 1942: NEA
Aspelta circinator (Gosse, 1886): AUS, NEA, NEO, PAL
Diglena circinator Gosse, 1886
Syn.: Diglena giraffa Gosse, 1886
Aspelta clydona Harring & Myers, 1928 (marine)
Syn.: Aspelta antonowiezae ďHondt, 1970
Aspelta curvidactyla Bērziņš, 1949: AFR, NEA, PAC, PAL
Aspelta egregia Myers, 1936: NEA, PAL
Aspelta europaea Hauer, 1939: PAL (marine and freshwater)
Aspelta harringi Remane, 1929 (marine)
Aspelta imbuta Harring & Myers, 1928: NEA, NEO
Syn.: Aspelta ecuadoriensis Koste, 1992
Aspelta intradentata Bērziņš, 1949: PAL
Aspelta labri Harring & Myers, 1928: NEA, PAL
Aspelta lestes Harring & Myers, 1928: NEA, PAL
Aspelta macra Harring & Myers, 1928: NEA, PAL
Aspelta pachida (Gosse, 1887) (marine)
Diglena pachida Gosse, 1887
Syn.: Pleurotrocha littoralis Levander, 1894
Aspelta psitta Harring & Myers, 1928: AUS, NEA, PAL
Aspelta reibischi (Remane, 1929) (marine)
Encentrum reibischi Remane, 1929
Aspelta tilba Koste & Shiel, 1987: AUS
Asplanchna Gosse, 1850
Asplanchnella Sudzuki, 1964

Asplanchna asymmetrica Shiel & Koste, 1985: AUS
Asplanchna brightwellii Gosse, 1850: AFR, AUS, NEA, NEO, ORI, PAC, PAL
Syn.: Asplanchna sieboldii urawaensis Sudzuki, 1956
Asplanchna girodi de Guerne, 1888: AFR, AUS, NEA, NEO, ORI, PAC, PAL
Asplanchna herrickii de Guerne, 1888: AFR, AUS, NEA, NEO, ORI, PAL
Asplanchna intermedia Hudson, 1886: AFR, AUS, NEA, ORI, PAL
Asplanchna priodonta Gosse, 1850: AFR, AUS, NEA, NEO, ORI, PAL
Syn.: Asplanchna priodonta sirakabana Sudzuki, 1964
Asplanchna sieboldii (Leydig, 1854): AFR, AUS, NEA, NEO, ORI, PAL
Notommata sieboldii Leydig, 1854
Asplanchna silvestrii Daday, 1902: NEA, NEO (doubtful Oriental record)
Asplanchna tropica Koste & Tobias, 1989: AFR, ORI

Asplanchnopus de Guerne, 1888
Asplanchnopus bhimavaramensis Dhanapathi, 1975: ORI

Asplanchnopus dahlgreni Myers, 1934: NEA, NEO, ORI, PAL (non-American records require confirma-
tion)
Asplanchnopus hyalinus Harring, 1913: AFR, AUS, NEO, ORI, PAL
Asplanchnopus multiceps (Schrank, 1793): AFR, AUS, NEA, NEO, ORI, PAL
Brachionus multiceps Schrank, 1793
Atrochus Wierzejski, 1893
Atrochus tentaculatus Wierzejski, 1893: AUS, ORI, PAL
Balatro Claparède, 1867
Balatro aciliatus (Radkewitsch, 1870) (Endoparasitic in terrestrial Oligochaeta)

Albertia aciliatus Radkewitsch, 1870
Balatro anguiformis Issel, 1904 (Endoparasitic in terrestrial Oligochaeta)
Balatro calvus Claparède, 1867 (Endoparasitic in terrestrial Oligochaeta)
Balatro fridericiae Kunst, 1954 (Endoparasitic in terrestrial Oligochaeta)
Beauchampia Harring, 1913
Beauchampia crucigera (Dutrochet, 1812): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Rotifer crucigere Dutrochet, 1812
Syn.: Cephalosiphon candidus Hudson, 1886 (synonym occasionally considered an infrasubspecific variant
of B. crucigera)
Syn.: Melicerta cephalosiphon Gosse, 1862
Beauchampiella Remane, 1929
Eudactylota Manfredi, 1927 non Walsingham, 1911

Manfredium Gallagher, 1957
Beauchampiella eudactylota (Gosse, 1886): AFR, AUS, NEA, NEO, ORI, PAL
Eudactylota eudactylota Gosse, 1886
Birgea Harring & Myers, 1922
Birgea enantia Harring & Myers, 1922: NEA
Brachionus Pallas, 1766
Brachionus adisi Koste & Hardy, 1984: NEO

Brachionus africanus Segers, Mbogo & Dumont, 1994: AFR (single Thai record a result of sample contam-
ination)
Brachionus ahlstromi Lindeman, 1939: AFR, AUS, NEA, NEO, ORI

Syn.: Brachionus caudatus var. personatus Ahlstrom, 1940
Syn.: Brachionus havanaensis var. ahlstromi Lindeman, 1939
Brachionus amazonicus Koste, 1983: NEO
Syn.: Brachionus urceolaris amazonica Koste, 1983
Brachionus amsterdamensis De Smet, 2001: ANT (Kerguelen islands)
Brachionus angularis Gosse, 1851
Syn.: Brachionus angularis orientalis Sudzuki, 1989
Brachionus angularis angularis Gosse, 1851: AFR, AUS, NEA, NEO, ORI, PAL
Syn.: Brachionus daitojimensis Sudzuki, 1992
Syn.: Brachionus donghuensis Sudzuki & Huang, 1997
Syn.: Brachionus lyratus yonaguniensis Sudzuki, 1992
Syn.: Brachionus minimus Bartsch, 1877
Syn.: Brachionus morondavaensis Sudzuki, 1998
Syn.: Brachionus papuanus Daday, 1897
Syn.: Brachionus pseudokeikoa Sudzuki, 1992
Syn.: Brachionus pyriformis Sudzuki & Huang, 1997
Syn.: Brachionus syennensis Schmarda, 1859
Syn.: Brachionus testudo Herberg, 1853
Brachionus angularis bidens Plate, 1886: AFR, AUS, NEA, ORI, PAL (taxon occasionally considered an
infrasubspecific variant of the nominal taxon)
Syn.: Brachionus kasadensis Sudzuki, 1976
Brachionus asplanchnoides Charin, 1947: PAL
Brachionus austrogenitus Ahlstrom, 1940: NEO (single unconfirmed AUS record)
Brachionus baylyi Sudzuki & Timms, 1977: AUS
Brachionus bennini Leissling, 1924: AFR, AUS, NEA, ORI, PAL
Syn.: Brachionus dimidiatus isigakiensis Sudzuki, 1992
Brachionus bennini beauchampi Sudzuki, 1997 (species inquirenda)
Brachionus bennini bennini Leissling, 1924
Brachionus bidentatus Anderson, 1889
Brachionus bidentatus bidentatus Anderson, 1889: AFR, ANT, AUS, NEA, NEO, ORI, PAL
Syn.: Brachionus bidentatus var. crassispinus Hauer, 1963
Syn.: Brachionus furculatus Thorpe, 1891
Syn.: Brachionus furculatus inermis Rousselet, 1906 (synonym considered an infrasubspecific variant (f.
inermis) of B. bidentatus)
Syn.: Brachionus furculatus testudinarius Jakubski, 1912 (synonym considered an infrasubspecific vari-
ant (f. testudinarius) of B. bidentatus)
Syn.: Brachionus furculatus var. jirovci Bartoš, 1946
Brachionus bidentatus minor Koste & Shiel, 1980: AUS
Brachionus bidentatus senegalensis Koste & De Ridder, 1985 (species inquirenda)
Brachionus budapestinensis Daday, 1885: AFR, AUS, NEA, NEO, ORI, PAL
Syn.: Brachionus similis Leissling, 1914
Brachionus calyciflorus Pallas, 1766: AFR, ANT, AUS, NEA, NEO, ORI, PAL (The taxonomy of B.
calyciflorus is particularly confused. Validation of the numerous proposed subspecies and infrasubspecific variants
awaits a thorough revision of the taxon)
Brachionus calyciflorus calyciflorus Pallas, 1766
Syn.: Brachionus amphiceros Ehrenberg, 1838 (synonym considered an infrasubspecific variant (f.

amphiceros) of B. calyiflorus)
Syn.: Brachionus dorcas Gosse, 1851 (synonym considered an infrasubspecific variant (f. dorcas) of B. caly-
ciflorus)
Syn.: Brachionus gillardi Hauer, 1966
Syn.: Brachionus pala Ehrenberg, 1838
Syn.: Brachionus pala anuraeiformis Brehm, 1909 (synonym considered an infrasubspecific variant (f.
anuraeiformis) of B. calyciflorus)
Brachionus calyciflorus gigantea Koste & Shiel, 1987 (subspecies inquirenda)
Brachionus caudatus Barrois & Daday, 1894: AFR, AUS, NEA, NEO, ORI, PAL (The taxonomy of B.
caudatus is particularly confused. Validation of the numerous proposed subspecies and infrasubspecific variants awaits a
thorough revision of the taxon)
Syn.: Brachionus caudatus singapurensis Sudzuki, 1989
Brachionus charini Kutikova, Kosova & Khodorevsky, 1976: PAL
Brachionus dichotomus Shephard, 1911
Brachionus dichotomus dichotomus Shephard, 1911: AUS
Brachionus dichotomus reductus Koste & Shiel, 1980: AUS, ORI
Brachionus dimidiatus Bryce, 1931: AFR, AUS, NEO, ORI, PAL
Syn.: Brachionus inermis Schmarda, 1894 (synonym considered an infrasubspecific variant (f. inermis) of B.
dimidiatus)
Brachionus diversicornis (Daday, 1883): AFR, AUS, NEO, ORI, PAL
Schizocerca diversicornis Daday, 1883
Brachionus dolabratus Harring, 1914: NEO (doubtful PAL record)
Brachionus donneri Brehm, 1951: ORI (single unconfirmed record from Panama canal waters, introduction?)
Brachionus donneri bifurcus Wu, 1981 (species inquirenda)
Brachionus donneri donneri Brehm, 1951
Brachionus durgae Dhanapathi, 1974: AFR, NEO, ORI, PAL
Syn.: Brachionus anchorporus Xu, Chen & Rao, 1997
Syn.: Brachionus durgae isigakiensis Sudzuki, 1992
Syn.: Brachionus moronensis Kuczynski, 1991
Brachionus falcatus Zacharias, 1898: AFR, AUS, NEA, NEO, ORI, PAL
Syn.: Brachionus falcatus reductus Koste & Shiel, 1987 (synonym occasionally considered an infrasub-
specific variant of the nominal taxon)
Brachionus forficula Wierzejski, 1891: AFR, AUS, ORI, PAL (introduced in NEA, doubtful NEO record)
Brachionus havanaensis Rousselet, 1911: NEA, NEO (introduced in ORI and PAL)
Syn.: Brachionus trahea Murray, 1913
Brachionus huangi Zhuge & Koste, 1996: PAL
Syn.: Brachionus brevispinus Sudzuki & Huang, 1997
Brachionus ibericus Ciros-Peréz, Gómez & Serra, 2001: PAL
Brachionus incertus Hauer, 1953: NEO
Brachionus josefinae Silva-Briano & Segers, 1992: NEA
Brachionus keikoa Koste, 1979: AUS
Brachionus kostei Shiel, 1983: AUS, ORI, PAL (Unpublished North-east Chinese record)
Brachionus kultrum Paggi, 1981: NEO
Brachionus leydigii Cohn, 1862
Brachionus leydigii leydigii Cohn, 1862: AFR, AUS, ORI, PAL
Syn.: Brachionus quadratus Rousselet, 1889 (non Müller, 1786. Synonym considered an infrasubspecific

variant (f. quadratus) of B. leydigii)
Syn.: Brachionus quadratus tridentatus Zernov, 1901 (synonym considered an infrasubspecific variant (f.
tridentatus) of B. leydigii)
Brachionus leydigii rotundus Rousselet, 1907: AUS, ORI, PAL
Syn.: Brachionus quadratus rotundus Rousselet, 1907 (synonym considered an infrasubspecific variant (f.
rotundus) of B. leydigii)
Brachionus lyratus Shephard, 1911
Brachionus lyratus lyratus Shephard, 1911: AUS, ORI
Brachionus lyratus tasmaniensis Koste & Shiel, 1986: AUS
Brachionus mirus Daday, 1905: NEO
Syn.: Brachionus mirus angustus Koste, 1972 (synonym probably represents an infrasubspecific variant)
Syn.: Brachionus mirus reductus Koste, 1972 (synonym probably represents an infrasubspecific variant)
Syn.: Brachionus voigti Hauer, 1961
Brachionus murphyi Sudzuki, 1989: ORI
Syn.: Brachionus niwati Sanoamuang, Segers & Dumont, 1995
Brachionus nilsoni Ahlstrom, 1940: AFR, AUS, NEA, NEO, ORI, PAL
Brachionus novaezealandiae Morris, 1913: AFR, AUS, PAL
Syn.: Brachionus novaezealandiae var. hungaricus Nogrady, 1957
Brachionus orientalis Rodewald, 1937 (species inquirenda)
Brachionus pinneenaus Koste & Shiel, 1983: AUS
Brachionus plicatilis Müller, 1786 (The B. plicatilis group is an as yet incompletely resolved cryptic species
group)
Brachionus plicatilis colongulaciensis Koste & Shiel, 1980: AUS
Brachionus plicatilis decemcornis Fadeev, 1925 (species inquirenda)
Brachionus plicatilis ecornis Fadeev, 1925 (species inquirenda)
Brachionus plicatilis estoniana Sudzuki, 1987 (species inquirenda)
Brachionus plicatilis longicornis Fadeev, 1925 (species inquirenda)
Brachionus plicatilis murrayi Fadeev, 1925 (species inquirenda)
Brachionus plicatilis plicatilis Müller, 1786: AFR, AUS, NEA, NEO, ORI, PAC, PAL
Syn.: Brachionus hepatotomus Gosse, 1851
Syn.: Brachionus muelleri Ehrenberg, 1834
Brachionus postcurvatus Kuczynski, 1991: NEO
Brachionus pseudonilsoni Sudzuki, 1992: PAL
Brachionus pterodinoides Rousselet, 1913: NEA, NEO, ORI, PAL
Syn.: Brachionus westphali Carlin-Nilson, 1935
Brachionus quadridentatus Hermann, 1783
Brachionus quadridentatus melhemi Barrois & Daday, 1894: AFR, AUS, NEO, ORI, PAL
Brachionus quadridentatus mirabilis Daday, 1897: AFR, AUS, NEA, NEO, ORI
Syn.: Brachionus variegatus Krau, 1962
Brachionus quadridentatus quadridentatus Hermann, 1783: AFR, ANT, AUS, NEA, NEO, ORI,
PAC, PAL
Syn.: Brachionus ancylognathus Schmarda, 1859 (synonym considered an infrasubspecific variant (f. ancy-
lognathus) of B. quadridentatus)
Syn.: Brachionus brevispinus Ehrenberg, 1832 (synonym considered an infrasubspecific variant (f. brevispi-
nus) of B. quadridentatus)
Syn.: Brachionus capsuliflorus Pallas, 1766

Syn.: Brachionus cluniorbicularis Skorikov, 1894 (synonym considered an infrasubspecific variant (f. clun-
iorbicularis) of B. quadridentatus)
Syn.: Brachionus cluniorbicularis isigakiensis Sudzuki, 1992
Syn.: Brachionus rhenanus Lauterborn, 1893 (synonym considered an infrasubspecific variant (f. rhenanus)
of B. quadridentatus)
Brachionus rotundiformis Tschugunoff, 1921: AFR, NEA, ORI, PAL
Brachionus rubens Ehrenberg, 1838: AFR, NEA, NEO, ORI, PAL
Syn.: Brachionus variabilis kasumigaurensis Sudzuki, 1998
Brachionus satanicus Rousselet, 1913: NEA, NEO
Brachionus schwoerbeli Koste, 1988: ORI
Brachionus sericus Rousselet, 1907: AFR, AUS, PAL
Brachionus sessilis Varga, 1951: AFR, AUS, NEO, ORI, PAL
Brachionus spatiosus Rousselet, 1912: NEA
Brachionus srisumonae Segers, Kotethip & Sanoamuang, 2004: ORI
Brachionus urceolaris Müller, 1773
Syn.: Brachionus urceolaris semicircularis Sudzuki, 1989
Brachionus urceolaris reductispinis Kuczynski, 1991: NEO
Brachionus urceolaris urceolaris Müller, 1773: AFR, ANT, AUS, NEA, NEO, ORI, PAL
Syn.: Tubipora urceus Linnaeus, 1758
Brachionus variabilis Hempel, 1896: AUS, NEA, NEO, ORI, PAL
Brachionus variabilis kohamensis Sudzuki, 1992 (species inquirenda)
Brachionus variabilis variabilis Hempel, 1896
Brachionus zahniseri Ahlstrom, 1934: NEA, NEO
Syn.: Brachionus gessneri Hauer, 1956
Bradyscela Bryce, 1910
Bradyscela clauda (Bryce, 1893): AUS, NEO, PAL

Adineta clauda Bryce, 1893
Bradyscela granulosa De Koning, 1947: PAL
Bryceella Remane, 1929
Bryceella stylata (Milne, 1886): NEA, PAC, PAL

Stephanops stylatus Milne, 1886
Bryceella tenella (Bryce, 1897): AUS, NEO, PAL
Stephanops tenellus Bryce, 1897
Syn.: Bryceella agilis Neiswestnowa-Shadina, 1935
Cephalodella Bory de St. Vincent, 1826
Metadiaschiza Fadeew, 1925

Paracephalodella Bērziņš, 1976

Cephalodella ablusa Myers, 1934: NEA
Cephalodella abstrusa Myers, 1934: NEA
Cephalodella akrobeles Myers, 1934: NEA, PAL
Cephalodella anebodica Bērziņš, 1976: PAL
Cephalodella angusta Myers, 1924: NEA
Cephalodella apocolea Myers, 1924: AUS, NEA, NEO, ORI, PAC, PAL
Cephalodella arcuata Wulfert, 1937: PAL
Cephalodella asarcia Myers, 1942: NEA
Cephalodella asta Donner, 1970: PAL
Cephalodella astricta Myers, 1934: NEA
Cephalodella auriculata (Müller, 1773): ANT, AUS, NEA, NEO, ORI, PAL
Vorticella auriculata Müller, 1773
Syn.: Cephalodella prompta Neiswestnova-Shadina, 1935
Cephalodella balatonica Zsuga, 1996 (species inquirenda)
Cephalodella belone Myers, 1924: NEA, NEO, ORI, PAL
Cephalodella bertonicensis Manfredi, 1927 (species inquirenda. Tentatively considered a synonym of C. sten-
roosi by Koste (1978))
Cephalodella biungulata Wulfert, 1937: AUS, NEO, PAL
Cephalodella boettgeri Koste, 1988: NEO, ORI
Cephalodella brandorffi Koste, 2000: NEO, ORI
Cephalodella calosa Wulfert, 1956: PAL
Cephalodella carina Wulfert, 1959: PAL
Cephalodella catellina (Müller, 1786): AFR, ANT, AUS, NEA, NEO, ORI, PAL (includes the infrasub-
specific variant minor Zawadovsky, 1926)
Cercaria catellina Müller, 1786
Syn.: Cephalodella armata Rudescu, 1960
Syn.: Cephalodella botezati Rodewald, 1935
Syn.: Cephalodella catellina natans Bērziņš, 1976
Syn.: Cephalodella myersi Wiszniewski, 1934
Cephalodella celeris Myers, 1937: NEA, PAL
Cephalodella clara Wulfert, 1944: NEO, PAL
Cephalodella collactea Myers, 1924: NEA
Cephalodella compacta Wiszniewski, 1934: PAL
Cephalodella compressa Myers, 1924: NEA
Cephalodella conica Nogrady, 1962: PAL
Cephalodella conjuncta Myers, 1940: NEA
Cephalodella crassipes (Lord, 1903): NEA, PAL
Diaschiza crassipes Lord, 1903
Cephalodella cyclops Wulfert, 1937: PAL
Cephalodella decidua Rodewald, 1935: PAL
Cephalodella delicata Wulfert, 1937: NEA, PAC, PAL
Syn.: Cephalodella eudelicata Wulfert, 1961
Cephalodella dentata Wulfert, 1937: PAL
Cephalodella derbyi (Dixon-Nuttall & Freeman, 1903): NEA
Diaschiza derbyi Dixon-Nuttall & Freeman, 1903
Cephalodella dixonnuttalli Myers, 1924: NEA

Cephalodella dora Wulfert, 1961: PAL
Cephalodella doryphora Myers, 1934: NEA, PAL
Cephalodella dorystoma Myers, 1934: NEA, PAL
Cephalodella dorseyi Myers, 1924: PAL
Cephalodella edax Hollowday, 1993: PAL (Parasitic in Uroglena volvox)
Cephalodella elegans Myers, 1924: NEA, NEO, PAL
Cephalodella elongata Myers, 1924: NEA, PAL
Cephalodella euderbyi Wulfert, 1940: PAL
Cephalodella euknema Myers, 1934: NEA, PAL
Cephalodella eunoma Myers, 1940: NEA
Cephalodella eupoda Myers, 1924: NEA
Cephalodella eurynota Myers, 1934: NEA
Cephalodella eva (Gosse, 1887): AUS, NEA, NEO, ORI, PAC, PAL
Furcularia eva Gosse, 1887
Cephalodella evabroedae De Smet, 1988: NEO, PAL
Cephalodella exigua (Gosse, 1886): AUS, NEA, NEO, ORI, PAL
Diaschiza exigua Gosse, 1886
Cephalodella fluviatilis (Zawadovsky, 1926): AFR, PAL
Diglena fluviatilis Zawadovsky, 1926
Syn.: Cephalodella elmenteita de Beauchamp, 1932
Cephalodella forceps Donner, 1950: PAL
Cephalodella forficata (Ehrenberg, 1832): ANT, AUS, NEA, NEO, ORI, PAC, PAL
Notommata forficata Ehrenberg, 1832
Cephalodella forficula (Ehrenberg, 1830): AUS, NEA, NEO, ORI, PAC, PAL
Distemma forficula Ehrenberg, 1830
Cephalodella friebei Koste, Robertson & Hardy, 1984: NEO
Cephalodella gibba (Ehrenberg, 1830): AFR, ANT, AUS, NEA, NEO, ORI, PAC, PAL
Furcularia gibba Ehrenberg, 1830
Syn.: Cephalodella microdactyla Koch-Althaus, 1963
Cephalodella gibboides Wulfert, 1950: PAL
Cephalodella gigantea Remane, 1933: NEO, ORI, PAL
Cephalodella gisleni Bērziņš, 1953: AUS
Cephalodella glandulosa Koch-Althaus, 1962: PAL
Cephalodella globata (Gosse, 1887): AUS, NEA, ORI, PAL
Diaschiza globata Gosse, 1887
Cephalodella glypha Wulfert, 1950: PAL
Cephalodella gobio Wulfert, 1937: PAL
Cephalodella gracilis (Ehrenberg, 1830) (Variable taxon. Includes the infrasubspecific variants lenticulare
Wulfert, 1938; sigmoides Wulfert, 1950)
Furcularia gracilis Ehrenberg, 1830
Cephalodella gracilis gracilis (Ehrenberg, 1830): AUS, NEA, NEO, ORI, PAC, PAL
Syn.: Cephalodella sagitta Wulfert, 1956
Cephalodella gracilis keta Reed, 1978: PAL
Cephalodella graciosa Wulfert, 1956: PAL
Cephalodella gusuleaci Rodewald, 1935: PAL
Cephalodella harringi Smirnov, 1927: PAL

Cephalodella hiulca Myers, 1924: NEA, PAL
Cephalodella hollowdayi Koste, 1986: NEO
Cephalodella hoodii (Gosse, 1886): NEA, ORI, PAL
Diaschiza hoodii Gosse, 1886
Syn.: Cephalodella remanei Wiszniewski, 1934
Syn.: Cephalodella strigosa Myers, 1924
Cephalodella hyalina Myers, 1924: NEA, PAL
Syn.: Cephalodella horni Wulfert, 1960
Cephalodella incila Wulfert, 1937: NEA, PAC, PAL
Cephalodella innesi Myers, 1924: NEA, PAL
Cephalodella inquilina Myers, 1924: NEA, PAL
Syn.: Cephalodella inquila auct.
Cephalodella intuta Myers, 1924
Cephalodella intuta intuta Myers, 1924: AUS, NEA, NEO, ORI, PAC, PAL
Cephalodella intuta jamaicaiensis Koste, Janetzky & Vareschi, 1993: NEO
Cephalodella irisae Koste, Janetzky & Vareschi, 1993: NEO
Cephalodella jakubskii Wiszniewski, 1953: PAL
Syn.: Cephalodella donneri Hauer, 1959
Cephalodella labiosa Wulfert, 1940: PAL
Cephalodella laisa Hauer, 1935: PAL
Cephalodella latifulcrum (Bērziņš, 1976): PAL
Paracephalodella latifulcrum Bērziņš, 1976
Cephalodella lepida Myers, 1934: NEA, PAL
Cephalodella licina Wulfert, 1961: PAL
Cephalodella licinia Myers, 1924: NEA
Cephalodella limosa Wulfert, 1937: NEO, PAL
Cephalodella lindamayae Koste & Shiel, 1986: AUS
Cephalodella lipara Myers, 1924: AUS, PAL
Cephalodella macrodactyla (Stenroos, 1898): NEA, PAL
Furcularia macrodactyla Stenroos, 1898
Syn.: Cephalodella strepta Myers, 1924
Cephalodella maior (Zawadovsky, 1926): AFR, PAL
Diglena maior Zawadovsky, 1926
Syn.: Cephalodella wiszniewskii Edmondson & Hutchinson, 1934
Cephalodella marina Myers, 1924: NEA
Syn.: Cephalodella epitedia Myers, 1924
Cephalodella megalocephala (Glascott, 1893): ANT, AUS, NEA, NEO, ORI, PAC, PAL
Furcularia megalocephala Glascott, 1893
Cephalodella megalotrocha Wiszniewski, 1934: NEO, PAL
Cephalodella melia Myers, 1924: NEA
Cephalodella mineri Myers, 1924: NEA
Cephalodella minora Wulfert, 1960: PAL
Cephalodella mira Myers, 1934: NEA, NEO, PAL
Syn.: Cephalodella nidicola Bērziņš, 1976
Cephalodella misgurnus Wulfert, 1937: AUS, NEO, ORI, PAL
Cephalodella monica Koste & Tobias, 1998: AFR

Cephalodella montana Myers, 1942: NEA
Cephalodella mucosa Myers, 1934: NEA
Cephalodella mucronata Myers, 1924: AFR, AUS, NEA, NEO, ORI, PAL
Cephalodella mus Wulfert, 1956: PAL
Cephalodella nana Myers, 1924: AUS, NEA, NEO, PAC, PAL
Cephalodella nelitis Myers, 1924: NEA
Cephalodella obesa Donner, 1949: PAL
Cephalodella obvia Donner, 1950: NEA, PAL
Cephalodella oxydactyla Wulfert, 1937: PAL
Cephalodella pachydactyla Wulfert, 1937: PAL
Syn.: Cephalodella nodosa Wulfert, 1937
Cephalodella pachyodon Wulfert, 1937: PAL
Cephalodella paggiae Koste & Robertson, 1983: NEO
Cephalodella panarista Myers, 1924: AUS, NEA, NEO, ORI, PAL
Cephalodella papillosa Myers, 1924: NEA
Cephalodella parasitica (Jennings, 1894): AUS, NEA, PAL
Pleurotrocha parasitica Jennings, 1894
Cephalodella paxi Wulfert, 1959: PAL
Cephalodella paxilla Myers, 1924: NEA
Cephalodella pentaplax Wulfert, 1943: PAL
Cephalodella pheloma Myers, 1924: AUS, NEA, PAL
Cephalodella physalis Myers, 1924: AUS, NEA, PAL
Cephalodella planera Myers, 1940: NEA
Cephalodella plicata Myers, 1924: AUS, NEA, PAL
Cephalodella poitera Myers, 1934: NEA
Cephalodella praelonga Myers, 1934: NEA
Cephalodella psammophila Koch-Althaus, 1962: PAL
Cephalodella pseudeva Bērziņš, 1976: PAL
Cephalodella qionghaiensis Koste & Zhuge, 1998: ORI
Cephalodella reimanni Donner, 1950: PAL
Cephalodella retusa Myers, 1924: NEA
Cephalodella rigida Donner, 1950: PAL
Cephalodella rostrum Reed, 1978: PAL
Cephalodella rotunda Wulfert, 1937
Cephalodella rotunda bryophila Pawłowski, 1938: ANT, PAC, PAL
Cephalodella rotunda rotunda Wulfert, 1937: NEA, PAL
Cephalodella segersi De Smet, 1998: PAL
Cephalodella somniculosa Margalef, 1948: PAL
Cephalodella songkhlaensis Segers & Pholpunthin, 1997: ORI
Cephalodella speciosa Myers, 1924: NEA
Cephalodella stenroosi Wulfert, 1937: NEO, PAL
Syn.: Cephalodella bertonicensis Manfredi, 1927
Syn.: Cephalodella deformis Donner, 1950
Syn.: Cephalodella var. austriaca Donner, 1978
Cephalodella sterea (Gosse, 1887): ANT, AUS, NEA, NEO, ORI, PAC, PAL (Morphologically variable
and complex taxon. Includes the infrasubspecific variants mutata, minor, dentata Donner, 1950; and f. nov. Donner,

1964)
Furcularia sterea Gosse, 1887
Syn.: Cephalodella sterea exoculis Bērziņš, 1976
Syn.: Cephalodella serrata Wulfert, 1937
Cephalodella strigosa Myers, 1924: NEA
Cephalodella subsecunda Myers, 1924: NEA, PAL
Cephalodella tachyphora Myers, 1924: NEA, PAL
Cephalodella tantilla Myers, 1924: AUS, NEA, ORI, PAL
Syn.: Cephalodella galbina Myers, 1924
Syn.: Cephalodella sabulosa Myers, 1942
Cephalodella tantilloides Hauer, 1935: AUS, PAL
Cephalodella tecta Donner, 1950: PAL
Cephalodella tempesta Wulfert, 1956: PAL
Cephalodella tenuior (Gosse, 1886): ANT, AUS, NEA, ORI, PAL (includes the infrasubspecific variant
pigmentata Wulfert, 1951)
Diaschiza tenuior Gosse, 1886
Cephalodella tenuis Koch-Althaus, 1962: PAL
Cephalodella tenuiseta (Burn, 1890): AUS, NEA, NEO, PAL
Furcularia tenuiseta Burn, 1890
Syn.: Cephalodella tenuiseta simplex Bērziņš, 1976
Cephalodella theodora Koch-Althaus, 1961: NEA, PAL
Cephalodella tinca Wulfert, 1937: AUS, NEO, ORI, PAL (includes the infrasubspecific variant conspicua
Donner, 1950)
Cephalodella tincaformis Koste, 1992: NEO
Cephalodella trigona (Rousselet, 1895): PAL
Diplois trigona Rousselet, 1895
Syn.: Furcularia leptodactyla Hauer, 1922
Syn.: Diplois sculpturata Daday, 1897
Cephalodella unguitata Hauer, 1935: PAL
Cephalodella vacuna Myers, 1924: NEA
Cephalodella ventripes (Dixon-Nuttall, 1901): AUS, NEA, NEO, PAC, PAL (includes the infrasubspe-
cific variant angustior Donner, 1950)
Diaschiza ventripes Dixon-Nuttall, 1901
Cephalodella vitella Wulfert, 1956: PAL
Cephalodella vittata Kutikova, 1985: PAL (Baikal endemic)
Cephalodella volvocicola (Zawadovsky, 1916): NEA, PAL
Diglena volvocicola Zawadovsky, 1916
Cephalodella wrighti Wulfert, 1960: PAL
Cephalodella xenica Myers, 1924: NEA, PAL
Cephalodella zeteta Wulfert, 1961: PAL
Ceratotrocha Bryce, 1910
Ceratotrocha cornigera (Bryce, 1893): ANT, AUS, NEA, NEO, PAL

Callidina cornigera Bryce, 1893
Ceratotrocha franzi Donner, 1949: PAL

Ceratotrocha rodewaldi Donner, 1962: PAL
Ceratotrocha velata Donner, 1949: PAL
Claria Kutikova, Markevich & Spiridonov, 1990
Claria segmentata Kutikova, Markevich & Spiridonov, 1990 (Endoparasitic in terrestrial Oligochaeta)
Collotheca Harring, 1913
Collotheca ambigua (Hudson, 1883): AFR, AUS, NEA, NEO, PAL

Floscularia ambigua Hudson, 1883
Syn.: Floscularia algicola Hudson, 1886
Syn.: Floscularia minor Hudson, 1886 (proposed as ssp.)
Collotheca annulata (Hood, 1888): AFR, PAL
Floscularia annulata Hood, 1888
Syn.: Floscularia uniloba Wierzejski, 1893
Collotheca atrochoides (Wierzejski, 1893): NEA, NEO, PAL
Floscularia atrochoides Wierzejski, 1893
Collotheca balatonica Varga, 1936: PAL
Collotheca bilfingeri Bērziņš, 1951: PAL
Syn.: Floscularia calva Bilfinger, 1894 (non Hudson (1885))
Collotheca bulbosa Bērziņš, 1951: PAL
Collotheca calva (Hudson, 1885): NEO, PAL
Floscularia calva Hudson, 1885
Collotheca campanulata (Dobie, 1849): AFR, ANT, AUS, NEA, NEO, ORI, PAL
Floscularia campanulata Dobie, 1849
Syn.: Collotheca gracilipes Edmondson, 1939 (synonym considered an infrasubspecific variant of C. cam-
panulata)
Syn.: Floscularia longicaudata Hudson, 1883 (synonym considered an infrasubspecific variant of C. cam-
panulata)
Collotheca coronetta (Cubitt, 1869): AFR, AUS, NEO, PAL
Floscularia coronetta Cubitt, 1869
Syn.: Collotheca coronetta weberi Summerfield-Wright, 1961 (synonym considered an infrasubspecific
variant of C. coronetta)
Syn.: Stephanoceros horatii Cubitt, 1871
Syn.: Floscularia longilobata Bartsch, 1877
Collotheca crateriformis Offord, 1934: PAL
Collotheca cucullata (Hood, 1894): PAL
Floscularia cucullata Hood, 1894
Collotheca cucullata cucullata (Hood, 1894)
Floscularia cucullata cucullata Hood, 1894
Collotheca cucullata smolandica Bērziņš, 1951: PAL
Collotheca discophora (Skorikov, 1903): PAL
Floscularia discophora Skorikov, 1903
Collotheca edentata (Collins, 1872): AUS, NEA, NEO, ORI, PAL

Floscularia edentata Collins, 1872
Syn.: Collotheca breviciliata Bērziņš, 1951 (synonym considered an infrasubspecific variant of C. edentata)
Collotheca edmondsoni Bērziņš, 1951: PAL
Collotheca epizootica (Monard, 1922): PAL
Floscularia epizootica Monard, 1922
Collotheca evansonii (Anderson & Shephard, 1892): AUS
Floscularia evansonii Anderson & Shephard, 1892
Collotheca ferox (Penard, 1914): PAC, PAL
Floscularia ferox Penard, 1914
Collotheca gossei (Hood, 1892): PAL
Floscularia gossei Hood, 1892
Collotheca heptabrachiata (Schoch, 1869): AFR, NEA, NEO, PAL
Floscularia heptabrachiata Schoch, 1869
Syn.: Floscularia diadema Petr, 1890 (synonym considered an infrasubspecific variant of C. heptabrachiata)
Syn.: Floscularia regalis Hudson, 1883
Collotheca hoodi (Hudson, 1883): NEA, PAL
Floscularia hoodi Hudson, 1883
Collotheca judayi Edmondson, 1940: NEA, NEO
Collotheca lettevalli Bērziņš, 1976: PAL
Collotheca libera (Zacharias, 1894): AFR, AUS, NEA, NEO, ORI, PAL
Floscularia libera Zacharias, 1894
Collotheca minuta (Milne, 1905): AFR, PAL
Floscularia minuta Milne, 1905
Collotheca monoceros (Zacharias, 1912): PAL
Floscularia monoceros Zacharias, 1912
Collotheca moselii (Milne, 1905): AFR, PAL
Floscularia moselii Milne, 1905
Collotheca mutabilis (Hudson, 1885): AFR, AUS, NEA, NEO, ORI, PAL
Floscularia mutabilis Hudson, 1885
Collotheca ornata (Ehrenberg, 1832): AFR, ANT, AUS, NEA, NEO, ORI, PAC, PAL
Floscularia ornata Ehrenberg, 1832
Syn.: Floscularia appendiculata Leydig, 1854 (synonym of C. ornata f. cornuta)
Syn.: Floscularia cornuta Dobie, 1849 (synonym considered an infrasubspecific variant of C. ornata)
Syn.: Floscularia cyclops Cubitt, 1871 (synonym considered an infrasubspecific variant of C. ornata)
Syn.: Floscularia mira Hudson, 1885
Syn.: Floscularia ornata natans Tschugunoff, 1921 (synonym considered an infrasubspecific variant of C.
ornata)
Syn.: Floscularia pentacornis Costa, 1838
Syn.: Collotheca volutata Sebestyén & Varga, 1950 (synonym considered an infrasubspecific variant of C.
ornata)
Collotheca pelagica (Rousselet, 1893): AFR, AUS, NEA, NEO, ORI, PAL
Floscularia pelagica Rousselet, 1893
Syn.: Collotheca liepetterseni Bērziņš, 1951 (synonym considered an infrasubspecific variant of C. pelag-
ica)
Syn.: Collotheca tubiformis Nipkow, 1961 (synonym considered an infrasubspecific variant of C. pelagica)
Collotheca polyphema Harring, 1914: NEA, NEO

Collotheca quadrilobata (Hood, 1892): PAL
Floscularia quadrilobata Hood, 1892
Collotheca quadrinodosa Summerfield-Wright, 1961: PAL
Syn.: Collotheca bjoerki Bērziņš, 1975 (synonym considered an infrasubspecific variant of C. quadrinodosa,
as var. bicornuta Koste, 1978)
Collotheca rasmae Bērziņš, 1951: PAL
Collotheca riverai Vilaclara & Sládeček, 1989: NEA
Collotheca sessilis (Milne, 1905): AFR, PAL
Floscularia sessilis Milne, 1905
Collotheca spinata (Hood, 1893): PAL
Floscularia spinata Hood, 1893
Collotheca stephanochaeta Edmondson, 1936: NEA, PAL
Collotheca tenuilobata (Anderson, 1889): AFR, AUS, NEA, NEO, PAL
Floscularia tenuilobata Anderson, 1889
Syn.: Collotheca corynetis Edmondson, 1939
Collotheca thunmarki Bērziņš, 1951: PAL
Collotheca torquilobata (Thorpe, 1891): PAL
Floscularia torquilobata Thorpe, 1891
Collotheca trifidlobata (Pittock, 1895): PAL
Floscularia trifidlobata Pittock, 1895
Collotheca trilobata (Collins, 1872): AFR, AUS, NEA, ORI, PAL
Floscularia trilobata Collins, 1872
Syn.: Floscularia trifolium Hudson, 1881
Syn.: Floscularia uniloba Wierzejski, 1893
Collotheca undulata Sládeček, 1968: PAL
Collotheca vargai Nogrady, 1976: NEA
Collotheca wiszniewskii Varga, 1938: PAL
Syn.: Collotheca wiszniewskii venusta Bērziņš, 1951 (synonym considered an infrasubspecific variant of
C. wiszniewskii)
Colurella Bory de St. Vincent, 1824
Colurus Ehrenberg, 1830
Colurella adriatica Ehrenberg, 1831: AFR, ANT, AUS, NEA, NEO, ORI, PAC, PAL (includes the
infrasubspecific variants angusta Donner, 1964; lata Donner, 1964 and α, β, γ Hauer (1924), marine and brackish water)
Syn.: Monura bartonia Gosse, 1887
Syn.: Colurus caudatus Ehrenberg, 1834
Syn.: Monura dulcis Ehrenberg, 1838
Syn.: Colurus leptus Gosse, 1887
Syn.: Colurus navalis Lord, 1884
Colurella anodonta Carlin, 1939: AFR, NEA, NEO, PAL
Colurella aquaeducti Török, 1956: PAL
Colurella collaris Wulfert, 1965: AFR
Colurella colurus (Ehrenberg, 1830)
Monura colurus Ehrenberg, 1830

Colurella colurus colurus (Ehrenberg, 1830): AFR, ANT, AUS, NEA, NEO, ORI, PAL (marine and
brackish water)
Syn.: Colurus amblytelus Gosse, 1886
Syn.: Colurus grallator Gosse, 1887
Syn.: Monura loncheres Gosse, 1887
Syn.: Colurella longidigita Mola, 1930
Syn.: Colurus rotundatus Daday, 1890
Colurella colurus compressa (Lucks, 1912): ANT, AUS, NEA, NEO, PAL
Colurus compressus Lucks, 1912
Colurella denticauda Carlin, 1939: NEA, NEO, PAC, PAL
Colurella dicentra (Gosse, 1887): NEA, NEO, PAL (marine and brackish water)
Colurus dicentrus Gosse, 1887
Colurella geophila Donner, 1951: NEA, NEO, ORI, PAL (taxonomic status uncertain; includes the infrasub-
specific variants gracilis Althaus, 1957; halensis Althaus, 1957; limnetica Althaus, 1957)
Syn.: Colurella gracilis Althaus, 1957 (non Hilgendorf (1899))
Syn.: Colurella subtilis Althaus, 1957
Colurella halophila Wulfert, 1942: PAL
Colurella hindenburgi Steinecke, 1917: AFR, AUS, NEA, NEO, ORI, PAC, PAL
Syn.: Colurella gastracantha Hauer, 1924 (synonym mostly considered an infrasubspecific variant of the
nominal taxon; confuse taxonomy)
Colurella marinovi Althaus, 1957 (marine)
Colurella mucronulata Ahlstrom, 1938: NEA
Colurella oblonga Donner, 1943: PAL (includes the infrasubspecific variant glabra Klement, 1959 of unclear
taxonomic status)
Colurella obtusa (Gosse, 1886) (taxonomic status uncertain; includes the infrasubspecific variants aperta Hauer,
1936; clausa Hauer, 1936; oxicauda Carlin, 1939)
Colurus obtusus Gosse, 1886
Colurella obtusa colpotum Bērziņš, 1982: AFR
Colurella obtusa obtusa (Gosse, 1886): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Colurella paludosa Carlin, 1939: NEA, PAL
Colurella psammophila Segers & Chittapun, 2001: ORI
Colurella salina Althaus, 1957: ANT, AUS, PAL (marine and brackish water)
Colurella sanoamuangae Chittapun, Pholpunthin & Segers, 1999: ORI
Colurella sinistra Carlin, 1939: AFR, NEA, ORI, PAL
Colurella sulcata (Stenroos, 1898): AFR, NEA, NEO, ORI, PAL
Metopidia sulcata Stenroos, 1898
Colurella tesselata (Glascott, 1893): AFR, AUS, NEA, NEO, ORI, PAL
Colurus tesselatus Glascott, 1893
Colurella uncinata (Müller, 1773)
Brachionus uncinatus Müller, 1773
Colurella uncinata bicuspidata (Ehrenberg, 1832): AFR, AUS, NEA, NEO, ORI, PAL
Colurus bicuspidatus Ehrenberg, 1832
Colurella uncinata deflexa (Ehrenberg, 1834): AUS, NEA, NEO, ORI, PAC, PAL
Colurus deflexus Ehrenberg, 1834
Colurella uncinata ornata Fadeew, 1927: AFR, PAL
Colurella uncinata uncinata (Müller, 1773): AFR, AUS, NEA, NEO, ORI, PAL

Syn.: Colurus dumnonius Gosse, 1887
Syn.: Monura micromela Gosse, 1887
Colurella unicauda Eriksen, 1968: PAL (marine and brackish water)
Conochilopsis Segers & Wallace, 20011
Conochilopsis causeyae (Vidrine, McLaughlin & Willis, 1985): AUS, NEA

Lacinularia causeyae Vidrine, McLaughlin & Willis, 1985
Conochilus Ehrenberg, 1834
subgenus Conochiloides Hlava, 1904
Conochilus (Conochiloides) coenobasis (Skorikov, 1914): AFR, AUS, NEA, NEO, ORI, PAL
Conochiloides coenobasis Skorikov, 1914
Conochilus (Conochiloides) deltaicus (Rudescu, 1960) (species inquirenda)
Conochiloides deltaicus Rudescu, 1960
Conochilus (Conochiloides) dossuarius Hudson, 1885: AFR, AUS, NEA, NEO, ORI, PAL
Conochilus (Conochiloides) exiguus Ahlstrom, 1938: NEA
Conochilus (Conochiloides) natans (Seligo, 1900): AFR, AUS, NEA, NEO, ORI, PAL
Tubicularia natans Seligo, 1900
subgenus Conochilus Ehrenberg, 1834
Conochilus (Conochilus) hippocrepis (Schrank, 1803): AFR, AUS, NEA, NEO, ORI, PAL
Linza hippocrepis Schrank, 1803
Syn.: Conochilus volvox Ehrenberg, 1834
Conochilus (Conochilus) unicornis Rousselet, 1892: AFR, AUS, NEA, NEO, ORI, PAL
Syn.: Conochilus leptopus Forbes, 1893
Syn.: Conochilus limneticus Stenroos, 1898
Syn.: Conochilus norvegicus Burckhardt, 1943 (synonym considered an infrasubspecific variant of C. uni-
cornis)
Cupelopagis Forbes, 1882
Apsilus Metchnikov, 1866 non Cuvier & Valenciennes, 1830

Dictyophora Leidy, 1857 non Germar, 1883
Cupelopagis vorax (Leidy, 1857): AFR, AUS, NEA, NEO, ORI, PAL
Dictyophora vorax Leidy, 1857
Syn.: Cupelopagis bucinedax Forbes, 1882
Syn.: Apsilus lentiformis Metchnikov, 1866
1. Segers & Wallace contains a reference as if it were published in 2000. However, the actual date of publication is
January, 2001.

Cyrtonia Rousselet, 1894
Cyrtonia tuba (Ehrenberg, 1834): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Notommata tuba Ehrenberg, 1834
Dicranophoroides De Smet, 1997
Dicranophoroides australiensis (Koste & Shiel, 1980): AUS

Dicranophorus australiensis Koste & Shiel, 1980
Dicranophoroides caudatus (Ehrenberg, 1834): AFR, AUS, NEA, NEO, ORI, PAL
Diglena caudatus Ehrenberg, 1834
Syn.: Diglena biraphis Gosse, 1851
Syn.: Dicranophorus caudatus braziliensis Koste, 1972
Dicranophoroides claviger (Hauer, 1965): AFR, NEO, ORI
Itura claviger Hauer, 1965
Dicranophoroides venezueliensis (Pourriot & Zoppi de Roa, 1981): AFR, NEO
Dicranophorus venezueliensis Pourriot & Zoppi de Roa, 1981
Dicranophorus Nitzsch, 1827
Dicranophorus alcimus Harring & Myers, 1928: NEA

Dicranophorus artamus Harring & Myers, 1928: NEA, PAL
Syn.: Dicranophorus lepsii Rodewald, 1935
Dicranophorus aspondus Harring & Myers, 1928: NEA, PAL
Dicranophorus biastis Harring & Myers, 1928: NEA
Dicranophorus bulgaricus Althaus, 1957 (marine psammon)
Dicranophorus cambari Wulfert, 1957: PAL
Dicranophorus capucinus Harring & Myers, 1928: NEA, PAL
Syn.: Dicranophorus capucinoides Wiszniewski, 1932
Dicranophorus colastes Harring & Myers, 1928: NEA, PAL
Dicranophorus corystis Harring & Myers, 1928: NEA, PAL
Dicranophorus difflugiarum (Penard, 1914): PAL
Diglena difflugiarum Penard, 1914
Dicranophorus dolerus Harring & Myers, 1928: NEA, ORI
Dicranophorus edestes Harring & Myers, 1928: AFR, NEA, PAL
Dicranophorus epicharis Harring & Myers, 1928: AFR, AUS, NEA, NEO, ORI, PAL
Dicranophorus esox Hauer, 1938: PAL
Dicranophorus facilis Kutikova, 1985: PAL (Baikal endemic)
Dicranophorus facinus Harring & Myers, 1928: NEA, PAL
Dicranophorus forcipatus (Müller, 1786)
Cercaria forcipata Müller, 1786
Dicranophorus forcipatus forcipatus (Müller, 1786): AFR, AUS, NEA, NEO, ORI, PAL
Syn.: Notommata oculifera Zavadowski, 1926
Syn.: Dicranophorus remanei Wulfert, 1936
Dicranophorus forcipatus parvidentatus Bērziņš, 1982: AFR (Madagascar)
Dicranophorus grandis (Ehrenberg, 1832): AFR, AUS, NEA, NEO, PAC, PAL

Diglena grandis Ehrenberg, 1832
Dicranophorus grypus Harring & Myers, 1928: NEA
Dicranophorus halbachi Koste, 1981: AFR, AUS
Dicranophorus haueri Harring & Myers, 1928: NEA, PAL
Dicranophorus hauerianus Wiszniewski, 1939: AUS, NEA, PAL
Dicranophorus hercules Wiszniewski, 1932: AFR, AUS, NEA, NEO, PAL
Syn.: Dicranophorus hercules adenta Wulfert, 1961
Syn.: Dicranophorus hercules intermedia Wiszniewski, 1934
Syn.: Dicranophorus novemdentatus Neiswestnowa-Shadina, 1935
Dicranophorus isothes Harring & Myers, 1928: NEA, PAL
Dicranophorus kostei Pourriot & Zoppi de Roa, 1981: NEO
Dicranophorus lenapensis Myers, 1938: NEA
Dicranophorus leptodon Wiszniewski, 1934: PAL
Dicranophorus liepolti Donner, 1964: PAL
Dicranophorus luetkeni (Bergendal, 1892): AFR, AUS, NEA, NEO, ORI, PAL
Arthroglena luetkeni Bergendal, 1892
Dicranophorus mesotis Harring & Myers, 1928: NEA
Dicranophorus minutes Myers, 1937: NEA, PAL
Dicranophorus myriophylli (Harring, 1913): NEA, ORI
Encentrum myriophylli Harring, 1913
Dicranophorus pauliani Bērziņš, 1982: AFR (Madagascar)
Dicranophorus pennatus Harring & Myers, 1928: NEA
Dicranophorus ponerus Harring & Myers, 1928: NEA, PAL
Dicranophorus prionacis Harring & Myers, 1928: AFR, NEA, NEO, ORI, PAL
Dicranophorus proclestes Harring & Myers, 1928: NEA, PAL
Dicranophorus riparius Kutikova, 1985: PAL (Baikal endemic)
Dicranophorus robustus Harring & Myers, 1928: AFR, AUS, NEA, NEO, ORI, PAL
Syn.: Dicranophorus robustus minor Bērziņš, 1949
Dicranophorus rostratus (Dixon-Nuttall & Freeman, 1902): NEA, NEO, PAL
Diglena rostrata Dixon-Nuttall & Freeman, 1902
Syn.: Dicranophorus cernuus Harring & Myers, 1928
Dicranophorus saevus Harring & Myers, 1928: NEA, PAL
Dicranophorus scotius Harring & Myers, 1928: NEA, PAL
Dicranophorus sebastus Harring & Myers, 1928: NEA, NEO
Dicranophorus secretus Donner, 1951: AFR, PAL
Dicranophorus semnus Harring & Myers, 1928: NEA, PAL
Dicranophorus siedleckii Wiszniewski, 1953: PAL
Dicranophorus sigmoides Wulfert, 1950: PAL
Dicranophorus spiculatus Myers, 1938: NEA, PAL
Dicranophorus strigosus Harring & Myers, 1928: NEA, PAL
Dicranophorus stultus Harring & Myers, 1928: NEA
Dicranophorus tegillus Harring & Myers, 1928: NEA, ORI
Dicranophorus thysanus Harring & Myers, 1928: NEA

Didymodactylos Milne, 1916
Didymodactylos carnosus Milne, 1916: AFR, AUS, PAL
Dipleuchlanis de Beauchamp, 1910
Dipleuchlanis elegans (Wierzejski, 1893): AUS, NEO, PAL

Euchlanis elegans Wierzejski, 1893
Dipleuchlanis ornata Segers, 1993: AFR, AUS, ORI
Dipleuchlanis propatula (Gosse, 1886): AFR, AUS, NEA, NEO, ORI, PAL
Diplois propatula Gosse, 1886
Diplois Gosse, 1886 (genus inquirendus)
Diplois daviesiae Gosse, 1886 (species inquirenda)

Syn.: Diplois phlegraea Iroso, 1910
Dissotrocha Bryce, 1910
Dissotrocha aculeata (Ehrenberg, 1832)

Philodina aculeata Ehrenberg, 1832
Dissotrocha aculeata aculeata (Ehrenberg, 1832): AFR, AUS, NEA, NEO, ORI, PAL
Dissotrocha aculeata crystallina (Murray, 1908): PAL
Philodina aculeata crystallina Murray, 1908
Dissotrocha aculeata decemcuspis Bērziņš, 1982: NEA
Dissotrocha aculeata didyma Bērziņš, 1982: PAL
Dissotrocha aculeata dukesi Russell, 1959: AUS
Dissotrocha aculeata ecksteini Bērziņš, 1982: PAL
Dissotrocha aculeata gossei Bērziņš, 1982 (No locality given)
Dissotrocha aculeata hebes Bērziņš, 1982: PAL
Dissotrocha aculeata inserta Bērziņš, 1982: NEO, PAL
Dissotrocha aculeata markmani Bērziņš, 1982: PAL
Dissotrocha aculeata medioaculeata (Janson, 1893): AFR, NEA, PAL
Philodina aculeata medioaculeata Janson, 1893
Dissotrocha aculeata milnei Bērziņš, 1982: PAL
Dissotrocha aculeata multicollis Bērziņš, 1982: PAL
Dissotrocha aculeata multiplicans Bērziņš, 1982: PAL
Dissotrocha aculeata murrayi Bērziņš, 1982: PAL
Dissotrocha aculeata octobullata Hauer, 1958: PAL
Dissotrocha aculeata octocollis Bērziņš, 1982: PAL
Dissotrocha aculeata quadricarinata Bērziņš, 1982: PAL
Dissotrocha aculeata quadrispinosa Bērziņš, 1982: PAL
Dissotrocha aculeata rachis Bērziņš, 1982: PAL
Dissotrocha aculeata repanda Bērziņš, 1982: PAL
Dissotrocha aculeata reversa Bērziņš, 1982: PAL
Dissotrocha aculeata serangodes Bērziņš, 1982: PAL

Dissotrocha aculeata simplex Bērziņš, 1982: PAL
Dissotrocha bjoerki Bērziņš, 1982: NEO
Dissotrocha decembullata Koste, 1996: AFR, NEO
Dissotrocha guyanensis Rougier & Pourriot, 2006: NEO
Dissotrocha hertzogi Hauer, 1939
Dissotrocha kostei Segers, 2007
Syn.: Dissotrocha hertzogi aculeata Koste, 1996
Dissotrocha macrostyla (Ehrenberg, 1838)
Philodina macrostyla Ehrenberg, 1838
Dissotrocha macrostyla macrostyla (Ehrenberg, 1838): AFR, AUS, NEA, NEO, ORI, PAL
Syn.: Philodina pannosa Bailey, 1851
Dissotrocha macrostyla tuberculata (Gosse, 1886): AUS, NEA, NEO, PAL
Philodina macrostyla tuberculata Gosse, 1886
Dissotrocha pectinata Murray, 1911: AFR
Dissotrocha schlienzi Hauer, 1956: NEO
Dissotrocha scutellata Bartoš, 1950: PAL
Dissotrocha spinosa (Bryce, 1892): AFR, PAL
Callidina spinosa Bryce, 1892
Donneria De Smet, 2003
Donneria sudzukii (Donner, 1968): NEA, PAL

Paradicranophorus sudzukii Donner, 1968
Dorria Myers, 1933
Dorria dalecarlica Myers, 1933: AUS, NEA, PAC
Dorystoma Harring & Myers, 1922
Dorystoma caudata (Bilfinger, 1894): AUS, NEA, PAL

Proales caudata Bilfinger, 1894
Dorystoma furcata Wulfert, 1939: PAL
Drilophaga Vejdovsky, 1883
Drilophaga bucephalus Vejdovsky, 1883: PAL (ectoparasitic on oligochaetes)

Drilophaga delagei de Beauchamp, 1904: PAL (ectoparasitic on oligochaetes)
Drilophaga judayi Harring & Myers, 1922: NEA, PAL (ectoparasitic on oligochaetes)
Elosa Lord, 1891
Elosa spinifera Wiszniewski, 1932: ORI, PAL (occasionally considered an infrasubspecific variant of E. wor-
rallii)
Elosa worrallii Lord, 1891: AUS, NEA, NEO, PAC, PAL

Embata Bryce, 1910
Embata commensalis (Western, 1893): NEA, PAL

Philodina commensalis Western, 1893
Embata hamata (Murray, 1906): AUS, PAL
Philodina hamata Murray, 1906
Embata laticeps (Murray, 1905): NEA, ORI, PAL
Philodina laticeps Murray, 1905
Embata laticornis (Murray, 1905): NEA, PAL
Philodina laticornis Murray, 1905
Embata parasitica (Giglioli, 1863): NEA, PAL
Callidina parasitica Giglioli, 1863
Encentrum Ehrenberg, 1838
Encentrum acrodon Wulfert, 1936: PAL

Encentrum algente Harring, 1921: NEA, PAL (marine and brackish water)
Encentrum alpinum Jersabek, 1999: PAL
Encentrum aquilus (Gosse, 1887): NEA, PAL
Diglena aquila Gosse, 1887
Encentrum arenarium Althaus, 1957 (marine psammon)
Encentrum arvicola Wulfert, 1936: PAL
Encentrum asellicola (Bartoš, 1947): PAL
Proales asellicola Bartoš, 1947
Encentrum astridae Sørensen, 2001 (marine psammon)
Encentrum axi Tzschaschel, 1979 (marine)
Encentrum barti De Smet, 2000 (marine)
Encentrum belluinum Harring & Myers, 1928: NEA
Encentrum bidentatum (Lie-Pettersen, 1906): PAL (marine and brackish water)
Pleurotrocha bidentata Lie-Pettersen, 1906
Syn.: Encentrum linnhei Scott, 1974
Encentrum boreale Harring & Myers, 1928: NEA, PAL
Encentrum brevifulcrum Dartnall, 1997: ANT
Encentrum caratum Myers, 1933: NEA
Encentrum carlini Myers, 1940: NEA
Encentrum cruentum Harring & Myers, 1928 (marine)
Encentrum desmeti Jersabek, 1999: PAL
Encentrum dieteri De Smet, 1995 (marine)
Encentrum diglandula (Zawadovsky, 1926): AUS, ORI, PAL
Proales diglandula Zawadovsky, 1926
Encentrum elongatum Harring & Myers, 1928: AUS, NEA, PAL
Encentrum enteromorphae Otto, 1936 (marine)
Encentrum eristes Harring & Myers, 1928 (marine)
Encentrum eulitorale Tzschaschel, 1979 (marine psammon)
Encentrum eurycephalum Wulfert, 1936: PAL
Encentrum felis (Müller, 1773): AUS, NEA, NEO, PAL

Vorticella felis Müller, 1773
Encentrum flexile Godske Eriksen, 1968: PAL (brackish water)
Encentrum fluviatile Wulfert, 1939: PAL
Encentrum forcipatum Dartnall, 1997: ANT
Encentrum frenoti De Smet, 2002 (marine)
Encentrum gibbosum Wulfert, 1936: AUS, PAL
Encentrum glaucum Wulfert, 1936: PAL
Encentrum goldschmidi Jersabek, 1999: PAL
Encentrum graingeri Chengalath, 1985 (marine)
Encentrum gulo Wulfert, 1936: PAL
Encentrum hofsteni (de Beauchamp, 1913): PAL
Diglena hofsteni de Beauchamp, 1913
Encentrum incisum Wulfert, 1936: PAL
Encentrum insolitum Myers, 1936: NEA
Encentrum kostei Tzschaschel, 1978 (marine psammon)
Encentrum kozminskii Wiszniewski, 1948: PAL
Encentrum kulmatyckii Wiszniewski, 1953
Syn.: Encentrum grande De Smet, 1997 (non Western (1891))
Encentrum kutikovae De Smet & Chernyshev, 2006 (marine psammon)
Encentrum lacidum Harring & Myers, 1928: NEA (brackish water)
Encentrum limicola Otto, 1936 (marine)
Encentrum listense Tzschaschel, 1979 (marine psammon)
Encentrum listensoides De Smet, 2000 (marine)
Encentrum longidens Donner, 1943: ORI, PAL
Encentrum longipes Wulfert, 1936: PAL
Encentrum longirostrum Tzschaschel, 1979 (marine psammon)
Encentrum lupus Wulfert, 1936: PAL
Encentrum lutra Wulfert, 1936: NEA, PAL
Encentrum mariae Koniar, 1957: PAL
Encentrum marinum (Dujardin, 1841): NEA, PAL (marine, brackish and freshwater)
Furcularia marina Dujardin, 1841
Encentrum martes Wulfert, 1939: PAL
Encentrum martoides Fott, 1960: PAL
Encentrum matthesi Remane, 1949 (marine psammon)
Encentrum minax Donner, 1943: PAL
Encentrum moldavicum Sláde…ek, 1961: PAL
Encentrum mucronatum Wulfert, 1936: NEA, PAL
Encentrum murrayi Bryce, 1922: PAL
Encentrum mustela (Milne, 1885): AFR, ANT, PAL
Pleurotrocha mustela Milne, 1885
Encentrum myersi Wulfert, 1936: PAL (brackish water)
Encentrum nesites Harring & Myers, 1928: NEA (marine and freshwater)
Encentrum nikor (Pawłowski, 1938): PAL
Dicranophorus nikor Pawłowski, 1938
Encentrum obesum Tzschaschel, 1979 (marine)
Encentrum oculatum Harring & Myers, 1928: NEA, PAL (brackish water)

Encentrum orthodactylum Wulfert, 1936: PAL
Encentrum otois Harring & Myers, 1928: NEA, PAL
Encentrum oxyodon Wulfert, 1936: PAL
Encentrum pachypus Wulfert, 1936: PAL
Encentrum parime Harring & Myers, 1928: NEA
Encentrum parvum Donner, 1952: PAL
Encentrum permolle (Gosse, 1886): NEA, PAL
Diglena permollis Gosse, 1886
Encentrum permutandum Tzschaschel, 1979 (marine)
Encentrum pornsilpi Segers & Chittapun, 2001: ORI
Encentrum porsildi Sørensen, 1998 (marine)
Encentrum psammophilum Althaus, 1957 (marine psammon)
Encentrum putorius Wulfert, 1936: AUS, NEA, PAL
Syn.: Encentrum putorius armatum Donner, 1943
Encentrum rapax Donner, 1943: PAL
Encentrum remanei Voigt, 1957 (marine psammon)
Syn.: Encentrum pachypus Remane, 1949 (non Wulfert (1936))
Encentrum rousseleti (Lie-Pettersen, 1905): PAL (marine and brackish water)
Diglena rousseleti Lie-Pettersen, 1905
Encentrum sacculiforme Tzschaschel, 1979 (marine psammon)
Encentrum salinum Dartnall, 1997: ANT
Encentrum salsum Myers, 1936: NEA (brackish water)
Encentrum saundersiae (Hudson, 1885): AUS, NEA, NEO, PAL
Taphrocampa saundersiae Hudson, 1885
Encentrum semiplicatum Wulfert, 1936: PAL
Encentrum simillimum Remane, 1929 (marine psammon)
Encentrum sorex Wulfert, 1950: PAC, PAL
Encentrum spatiatum Wulfert, 1936: PAL (brackish and freshwater)
Encentrum spinosum Koch-Althaus, 1962: PAL
Encentrum stechlinense Koch-Althaus, 1962: PAL
Encentrum striatum Althaus, 1957 (marine psammon)
Encentrum sutor Wiszniewski, 1936: NEA, PAL
Encentrum sutoroides Wulfert, 1940: PAL
Encentrum tectipes Remane, 1949 (marine psammon)
Encentrum tenuidigitatum De Smet, 2000 (marine psammon)
Encentrum tobyhannaense Myers, 1940: NEA, PAL
Encentrum torvitoides (Smirnov, 1930): PAL
Dicranophorus torvitoides Smirnov, 1930
Encentrum torvitum (Harring & Myers, 1928): NEA
Dicranophorus torvitus Harring & Myers, 1928
Encentrum tyrphos Wulfert, 1936: NEA, PAL
Encentrum umbonatum Kutikova, 1985 (Baikal endemic)
Encentrum uncinatum (Milne, 1886): AFR, ANT, NEO, ORI, PAL
Diglena uncinata Milne, 1886
Encentrum ussuriensis De Smet & Chernyshev, 2006 (marine psammon)
Encentrum valkanovi Althaus, 1957 (marine psammon)

Encentrum villosum Harring & Myers, 1928: NEA, PAL (marine and brackish water)
Encentrum voigti Wulfert, 1936: PAL
Encentrum walterkostei Jersabek, 1994: PAL
Encentrum wiszniewskii Wulfert, 1939: PAL
Encentrum zetetum Harring & Myers, 1928: NEA
Enteroplea Ehrenberg, 1830
Enteroplea lacustris Ehrenberg, 1830: AUS, NEA, NEO, ORI, PAL
Eosphora Ehrenberg, 1830
Eosphora anthadis Harring & Myers, 1922: AUS, NEA, NEO, ORI, PAL
Eosphora ehrenbergi Weber & Montet, 1918: AFR, AUS, NEA, NEO, ORI, PAL
Syn.: Notommata najas Ehrenberg, 1832
Eosphora gibba Garner, 1937: PAL
Eosphora najas Ehrenberg, 1830: AFR, AUS, NEA, NEO, ORI, PAL
Eosphora therina Harring & Myers, 1922: NEA
Eosphora thoa Harring & Myers, 1924: AUS, NEA, NEO, PAL
Eosphora thoides Wulfert, 1935: AFR, AUS, NEA, NEO, PAL
Eothinia Harring & Myers, 1922
Eothinia argus Harring & Myers, 1924: NEA

Eothinia carogaensis Myers, 1937: NEA
Eothinia elongata (Ehrenberg, 1832): AFR, AUS, NEA, NEO, ORI, PAL
Eosphora elongata Ehrenberg, 1832
Syn.: Eothinia elongata macra B‘rziņÓ, 1949
Syn.: Eothinia lamellata B‘rziņÓ, 1949
Eothinia eukolpa Myers, 1940: NEA
Eothinia poitera Myers, 1933: NEA
Eothinia triphaea Harring & Myers, 1924: NEA, PAL
Syn.: Eothinia lasiobiotica B‘rziņÓ, 1949
Epiphanes Ehrenberg, 1832
Hydatina Ehrenberg, 1830 (non Schumacher (1817))
Epiphanes brachionus (Ehrenberg, 1837)

Notommata brachionus Ehrenberg, 1837
Epiphanes brachionus brachionus (Ehrenberg, 1837): AFR, AUS, NEA, NEO, ORI, PAL
Epiphanes brachionus spinosa (Rousselet, 1901): AFR, NEO, ORI, PAL
Notops brachionus spinosa Rousselet, 1901
Syn.: Brachionus inermispinosus De Smet, 1989
Epiphanes clavulata (Ehrenberg, 1832): AFR, AUS, NEA, NEO, ORI, PAL
Notommata clavulata Ehrenberg, 1832

Epiphanes desmeti Rougier & Pourriot, 2006 (species inquirenda)
Epiphanes macroura (Barrois & Daday, 1894): AFR, AUS, NEA, NEO, ORI, PAL
Notops macrourus Barrois & Daday, 1894
Syn.: Notops mollis Hempel, 1896
Epiphanes pelagica (Jennings, 1900): NEA, PAL
Notops pelagicus Jennings, 1900
Epiphanes senta (Müller, 1773): AFR, ANT, AUS, NEA, NEO, ORI, PAC, PAL
Vorticella senta Müller, 1773
Syn.: Enteroplea hydatina Ehrenberg, 1831
Syn.: Hydatina macragnatha Schmarda, 1859
Syn.: Hydatina monops Hilgendorf, 1899
Syn.: Hydatina oblonga Daday, 1908
Erignatha Harring & Myers, 1922
Erignatha capula Harring & Myers, 1928: NEA

Erignatha clastopis (Gosse, 1886): AUS, NEA, NEO, PAL
Diglena clastopis Gosse, 1886
Erignatha longidentata Sørensen, 2001: PAL (brackish water)
Erignatha sagitta Harring & Myers, 1928: NEA
Erignatha sagittoides Wiszniewski, 1935: PAL
Erignatha tenuidens (de Beauchamp, 1913): PAL
Diglena tenuidens de Beauchamp, 1913
Euchlanis Ehrenberg, 1832
Dapidia Gosse, 1887
Euchlanis alata Voronkov, 1912: AUS, NEA, ORI, PAL

Euchlanis arenosa Myers, 1936: NEA
Euchlanis callimorpha B‘rziņÓ, 1957: AFR
Euchlanis callysta Myers, 1930: NEA
Euchlanis calpidia Myers, 1930: AUS, NEA, ORI, PAL
Euchlanis contorta (Wulfert, 1939): PAL
Dapidia contorta Wulfert, 1939
Euchlanis dactyliseta Sudzuki, 1998 (species inquirenda)
Euchlanis dapidula Parise, 1966: PAL
Euchlanis deflexa (Gosse, 1851): AFR, AUS, NEA, NEO, ORI, PAL
Dapidia deflexa Gosse, 1851
Syn.: Euchlanis calpidia Myers, 1930
Euchlanis dilatata Ehrenberg, 1832
Euchlanis dilatata dilatata Ehrenberg, 1832: AFR, ANT, AUS, NEA, NEO, ORI, PAC, PAL
Syn.: Euchlanis hipposideros Gosse, 1851
Syn.: Euchlanis uniseta Leydig, 1854
Euchlanis dilatata lucksiana Hauer, 1930: AFR, AUS, NEA, NEO, ORI, PAL
Syn.: Euchlanis dilatata crassa Myers, 1938

Euchlanis hyphidactyla Parise, 1963: NEA, PAL
Euchlanis incisa Carlin, 1939: AFR, AUS, NEA, NEO, ORI, PAC, PAL (incl. the infrasubspecific variant
mucronata Ahlstrom, 1934)
Euchlanis ligulata Kutikova & Vassileva, 1982: PAL (Baikal endemic)
Euchlanis lyra Hudson, 1886: AFR, AUS, NEA, NEO
Syn.: Euchlanis myersi Kutikova, 1959
Euchlanis mamorokaensis B‘rziņÓ, 1973 (species inquirenda)
Euchlanis meneta Myers, 1930: AFR, AUS, NEA, NEO, ORI, PAL
Syn.: Euchlanis proxima Myers, 1930
Euchlanis mikropous Koch-Althaus, 1962: PAL
Euchlanis oropha Gosse, 1887: AFR, ANT, AUS, NEA, NEO, ORI, PAL
Syn.: Euchlanis parva Rousselet, 1892
Euchlanis parameneta B‘rziņÓ, 1973 (species inquirenda)
Euchlanis perpusilla De Ridder, 1977 (species inquirenda)
Euchlanis phryne Myers, 1930: AUS, NEA
Euchlanis pyriformis Gosse, 1851: AFR, AUS, NEA, NEO, PAL
Euchlanis semicarinata Segers, 1993: AFR, AUS, NEO, ORI
Euchlanis triquetra Ehrenberg, 1838: AFR, AUS, NEA, NEO, ORI, PAL
Syn.: Euchlanis brahmae Dhanapathi, 1976
Syn.: Dapidia carinata Carlin-Nilson, 1934
Syn.: Dapidia lata Carlin-Nilson, 1934
Syn.: Euchlanis longobardica Manfredi, 1927
Syn.: Euchlanis pellucida Harring, 1921
Syn.: Euchlanis triquetra pterigoidea Grese, 1955
Filinia Bory de St. Vincent, 1824
Fadeewella Smirnov, 1928

Parafilinia Sudzuki, 1989
Tetramastix Zacharias, 1898
Triarthra Ehrenberg, 1832
Filinia australiensis Koste, 1980: AFR, AUS

Filinia brachiata (Rousselet, 1901): AFR, AUS, NEA, ORI, PAL
Triarthra brachiata Rousselet, 1901
Syn.: Filinia aseta Fadeew, 1925
Filinia camasecla Myers, 1938: ORI (Single Neotropical record from Panama; introduction?)
Syn.: Filinia camasecla cambodgensis B‘rziņÓ, 1973
Filinia cornuta (Weisse, 1847): AFR, AUS, NEA, ORI, PAL
Triarthra cornuta Weisse, 1847
Syn.: Triarthra breviseta Gosse, 1851
Syn.: Triarthra breviseta monoarthra Sokolova, 1922
Filinia grandis Koste & Shiel, 1980: AUS
Filinia hofmanni Koste, 1980: PAL
Filinia limnetica (Zacharias, 1893): AFR, AUS, NEA, NEO, ORI, PAL
Triarthra limnetica Zacharias, 1893

Syn.: Triarthra tranites Skorikov, 1896
Filinia longiseta (Ehrenberg, 1834): AFR, AUS, NEA, NEO, ORI, PAL (Records of F. longiseta from
hypolimnetic conditions may be F. hofmanni)
Triarthra longiseta Ehrenberg, 1834
Syn.: Filinia longiseta intermedia Buchholz, 1952
Syn.: Filinia longiseta minor Evans, 1949
Filinia minuta (Smirnov, 1928): PAL
Fadeewella minuta Smirnov, 1928
Syn.: Fadeewella nipponensis Sudzuki, 1975
Filinia novaezealandiae Shiel & Sanoamuang, 1993: AFR, AUS, NEA, ORI
Filinia opoliensis (Zacharias, 1898): AFR, AUS, NEA, NEO, ORI, PAL
Tetramastix opoliensis Zacharias, 1898
Syn.: Tetramastix taiwanensis Harada, 1937
Filinia passa (Müller, 1786): AFR, AUS, NEA, ORI, PAL
Brachionus passus Müller, 1786
Syn.: Filinia muelleri Bory de St. Vincent, 1826
Syn.: Triarthra mystacina Ehrenberg, 1832
Filinia pejleri Hutchinson, 1964: AFR, ANT, AUS, NEO, ORI, PAL
Syn.: Filinia terminalis kergueleniensis Lair & Koste, 1984
Filinia saltator (Gosse, 1886): AFR, NEO, PAL
Pedetes saltator Gosse, 1886
Syn.: Filinia longiseta var. acaudata Hauer, 1953
Filinia terminalis (Plate, 1886): AFR, AUS, NEA, NEO, ORI, PAL (includes the infrasubspecific variant
baikalensis Eggert, 1973. Warm-water records of F. terminalis may be F. novaezealandiae)
Triarthra terminalis Plate, 1886
Syn.: Triarthra maior Colditz, 1924
Floscularia Cuvier, 1798
Floscularia armata Segers, 1997: NEA, ORI

Floscularia bifida Segers, 1997: NEA
Floscularia conifera (Hudson, 1886): AFR, AUS, NEA, NEO, ORI, PAL
Melicerta conifera Hudson, 1886
Floscularia curvicornis Rougier & Pourriot, 2006 (species inquirenda)
Floscularia decora Edmondson, 1940: NEO, ORI
Floscularia janus (Hudson, 1881): AFR, AUS, NEA, NEO, PAC, PAL
Oecistes janus Hudson, 1881
Syn.: Melicerta flocculosa Kellicott, 1896
Floscularia melicerta (Ehrenberg, 1832): AFR, AUS, NEA, NEO, ORI, PAL
Lacinularia melicerta Ehrenberg, 1832
Syn.: Tubicolaria najas Ehrenberg, 1838
Syn.: Melicerta tubicolaria Hudson & Gosse, 1886
Syn.: Melicerta tyro Hudson, 1875
Floscularia noodti Koste, 1972: NEO
Floscularia pedunculata (Joliet, 1883): NEA, PAL
Melicerta pedunculata Joliet, 1883

Floscularia ringens (Linnaeus, 1758): AFR, AUS, NEA, NEO, ORI, PAL
Serpula ringens Linnaeus, 1758
Syn.: Rotifer quadricircularis Dutrochet, 1812
Syn.: Tubicolaria quadriloba Lamarck, 1816
Syn.: Tubicolaria quadrilobata Blainville, 1828
Syn.: Tubicolaria tetrapetala Cuvier, 1817
Syn.: Brachionus tubifex Pallas, 1766
Gastropus Imhof, 1898
Postclausa Hilgendorf, 1899

Gastropus hyptopus (Ehrenberg, 1838): AFR, AUS, NEA, NEO, ORI, PAL
Notommata hyptopus Ehrenberg, 1838
Syn.: Gastropus bretensis Linder, 1904
Gastropus minor (Rousselet, 1892): AFR, AUS, NEA, NEO, ORI, PAL
Notops minor Rousselet, 1892
Syn.: Postclausa cricularis Hilgendorf, 1899
Syn.: Notops fennicus Stenroos, 1898
Syn.: Postclausa minuta Hilgendorf, 1899
Syn.: Hypopus ritenbenki Bergendal, 1892
Gastropus stylifer (Imhof, 1891): AFR, AUS, NEA, NEO, PAL
Notops stylifer Imhof, 1891
Syn.: Sacculus orbicularis Kellicott, 1897
Syn.: Hudsonella picta Zacharias, 1893
Syn.: Hudsonella pygmaea Zacharias, 1894
Syn.: Notops pygmaeus Calman, 1892
Syn.: Hudsonia ruber Hood, 1893
Glaciera Jersabek, 1999
Glaciera schabetsbergeri Jersabek, 1999: PAL
Habrotrocha Bryce, 1910
Habrotrocha acornis Murray, 1911: AFR, NEO

Habrotrocha alacris Milne, 1916: AFR
Habrotrocha amphichlaena De Koning, 1947: PAL
Habrotrocha ampulla Murray, 1911: AFR, NEA, NEO, ORI, PAL
Habrotrocha angularis (Murray, 1910): ANT
Callidina angularis Murray, 1910
Habrotrocha angusticollis (Murray, 1905)
Callidina angusticollis Murray, 1905
Habrotrocha angusticollis alterversa Koste, 1970: PAL
Habrotrocha angusticollis angusticollis (Murray, 1905): AFR, ANT, AUS, NEA, NEO, ORI, PAL
Habrotrocha angusticollis attenuata (Murray, 1906): AUS, NEO, ORI, PAL
Callidina angusticollis attenuata Murray, 1906

Habrotrocha angusticollis monteti De Koning, 1947: AUS, PAL
Habrotrocha angusticollis reversa Bartoš, 1951: AUS, PAL
Habrotrocha annulata (Murray, 1905): NEA, NEO, PAL
Callidina annulata Murray, 1905
Habrotrocha appendiculata Murray, 1911: AFR, AUS, PAL
Habrotrocha aspera (Bryce, 1892): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Callidina aspera Bryce, 1892
Habrotrocha baradlana Varga, 1963: PAL
Habrotrocha bartosi Koniar, 1955: PAL
Habrotrocha bidens (Gosse, 1851): AFR, AUS, NEA, NEO, ORI, PAL
Callidina bidens Gosse, 1851
Syn.: Habrotrocha milnei Bryce, 1922
Habrotrocha brocklehursti Murray, 1911: AFR, PAL
Habrotrocha bulbosa Wulfert, 1961: PAL
Habrotrocha calosa Wulfert, 1942: PAL
Habrotrocha caudata Murray, 1911: AFR, AUS, NEO, PAC
Habrotrocha collaris (Ehrenberg, 1832): AFR, AUS, NEA, PAL
Philodina collaris Ehrenberg, 1832
Habrotrocha colliflectens Bartoš, 1944: PAL
Habrotrocha constricta (Dujardin, 1841): AFR, ANT, AUS, NEA, NEO, PAC, PAL
Callidina constricta Dujardin, 1841
Habrotrocha crassa Donner, 1949: PAL
Habrotrocha crenata (Murray, 1905)
Callidina crenata Murray, 1905
Habrotrocha crenata crenata (Murray, 1905): ANT, AUS, PAL
Syn.: Mniobia scarlatina f. angulata Bartoš, 1938
Habrotrocha crenata sphagnicola Paw»owski, 1938: AFR, PAL
Syn.: Mniobia scarlatina f. rugosa Bartoš, 1938
Habrotrocha cucullata Murray, 1911: AFR, AUS
Habrotrocha cuneata Murray, 1913: NEO
Habrotrocha curva De Koning, 1947: PAL
Habrotrocha curvicollis Bartoš, 1951: PAL
Habrotrocha diarthrantenna De Koning, 1947: PAL
Habrotrocha elegans (Milne, 1886): PAL
Macrotrachela elegans Milne, 1886
Syn.: Callidina venusta Bryce, 1897
Habrotrocha elliptica Bartoš, 1963: ORI
Habrotrocha elusa Milne, 1916
Habrotrocha elusa elusa Milne, 1916: AFR, ANT, AUS, PAL
Habrotrocha elusa vegeta Milne, 1916: AFR, PAL
Habrotrocha eremita (Bryce, 1894): AFR, AUS, NEA, NEO, PAL
Callidina eremita Bryce, 1894
Habrotrocha filum Donner, 1949: NEO, PAL
Habrotrocha flava Bryce, 1915: AUS, NEA, PAL
Habrotrocha flaviformis De Koning, 1947: AUS, NEO, PAL
Habrotrocha flexicollis Bartoš, 1963: ORI

Habrotrocha fuhrmanni Heinis, 1913: NEO
Habrotrocha fusca (Bryce, 1894): AUS, NEO, ORI, PAL
Callidina fusca Bryce, 1894
Habrotrocha fuscochlaena De Koning, 1947: PAL
Habrotrocha gibbosa De Koning, 1947: AUS, PAL
Habrotrocha gracilis Montet, 1915
Habrotrocha gracilis gracilis Montet, 1915: AUS, PAL
Habrotrocha gracilis quadridens Schulte, 1954: PAL
Habrotrocha granulata De Koning, 1947: PAL
Habrotrocha gulosa Milne, 1916: AFR, PAL
Habrotrocha heinisi Bartoš, 1951: NEO
Habrotrocha humilis Schulte, 1954: PAL
Habrotrocha incola Bartoš, 1951: PAL
Habrotrocha iners Milne, 1916: AFR
Habrotrocha insignis Bryce, 1915: AUS, PAL
Syn.: Habrotrocha proxima Donner, 1950
Habrotrocha komareki Bartoš, 1944: PAL
Habrotrocha lamellata Bartoš, 1951: PAL
Habrotrocha lata (Bryce, 1892)
Callidina lata Bryce, 1892
Habrotrocha lata lata (Bryce, 1892): AFR, AUS, NEA, NEO, ORI, PAL
Habrotrocha lata lens Donner, 1965: PAL
Habrotrocha lata paxi Wulfert, 1941: PAL
Habrotrocha lata progonidia (Monard, 1918): PAL
Callidina progonidia Monard, 1918
Habrotrocha leitgebii (Zelinka, 1886): AUS, ORI, PAC, PAL
Callidina leitgebii Zelinka, 1886
Habrotrocha levis Donner, 1951: PAL
Habrotrocha ligula Bryce, 1913
Habrotrocha ligula aligula Burger, 1948: AUS, NEA, PAL
Habrotrocha ligula ligula Bryce, 1913: AUS, NEA, PAL
Habrotrocha ligula loxoglotta De Koning, 1947: PAL
Habrotrocha longicalcarata B‘rziņÓ 1950: PAL
Habrotrocha longiceps (Murray, 1906): AFR, AUS, NEO, PAC, PAL
Callidina longiceps Murray, 1906
Habrotrocha longicollis Bartoš, 1963: ORI
Habrotrocha longula Bryce, 1915: PAL
Habrotrocha maculata Murray, 1911: AFR, NEA, NEO, PAL
Habrotrocha mediocris Donner, 1951: PAL
Habrotrocha megalocephala De Koning, 1947: PAL
Habrotrocha microcephala (Murray, 1906): AUS, NEA, ORI, PAL
Callidina microcephala Murray, 1906
Habrotrocha minima De Koning, 1947: PAL
Habrotrocha minuta (Murray, 1908): PAL
Callidina minuta Murray, 1908
Habrotrocha modesta Bartoš, 1963: ORI

Habrotrocha munda Bryce, 1913: AFR, AUS, NEA, PAL
Habrotrocha murrayi Bartoš, 1951: AFR
Habrotrocha nodosa Murray, 1911: AFR, NEO, ORI, PAC, PAL
Habrotrocha nodulata De Koning, 1947: PAL
Habrotrocha novemdens De Koning, 1947: PAL
Habrotrocha parvipes Donner, 1951: PAL
Habrotrocha pavida Bryce, 1915: NEA, PAL
Habrotrocha perforata (Murray, 1906)
Callidina perforata Murray, 1906
Habrotrocha perforata americana (Murray, 1907): NEO, ORI
Callidina perforata americana Murray, 1907
Habrotrocha perforata perforata (Murray, 1906): AFR, AUS, NEO, ORI, PAL
Habrotrocha pertinax Milne, 1916: AFR
Habrotrocha placida Milne, 1916: AFR
Habrotrocha plana Milne, 1916: AFR, PAL
Habrotrocha porrecta B‘rziņÓ, 1950: PAL
Habrotrocha praelonga De Koning, 1947: PAL
Habrotrocha puella Donner, 1950
Habrotrocha puella excedens Donner, 1962: AFR, AUS, PAL
Habrotrocha puella puella Donner, 1950: AUS, NEO, PAL
Habrotrocha pulchra (Murray, 1905): AFR, ANT, AUS, NEA, NEO, PAL
Callidina pulchra Murray, 1905
Habrotrocha pusilla (Bryce, 1893)
Callidina pusilla Bryce, 1893
Habrotrocha pusilla nuda Donner, 1950: AUS, PAL
Habrotrocha pusilla pusilla (Bryce, 1893): AFR, AUS, NEA, PAL
Habrotrocha pusilla textrix (Bryce, 1897): AFR, PAL (incl. f. longilabris Donner, 1950; f. brevilabris Don-
ner, 1950)
Callidina pusilla textrix Bryce, 1897
Habrotrocha quinquedens De Koning, 1947
Habrotrocha quinquedens doornensis De Koning, 1947: PAL
Habrotrocha quinquedens quinquedens De Koning, 1947: PAL
Habrotrocha rara Donner, 1949: NEO, PAL
Habrotrocha reclusa (Milne, 1889): AUS, NEA, PAL
Macrotrachela reclusa Milne, 1886
Habrotrocha recumbens Bartoš, 1963: ORI
Habrotrocha roeperi (Milne, 1889): NEA, PAL
Macrotrachela roeperi Milne, 1889
Habrotrocha rosa Donner, 1949: AFR, AUS, NEA, NEO, PAL
Habrotrocha scabropyga Bartoš, 1958: PAL
Habrotrocha scepanotrochoides De Koning, 1947: AUS, PAL
Habrotrocha schultei Donner, 1965: PAL
Habrotrocha serpens Donner, 1949: AFR, AUS, PAL
Syn.: Habrotrocha stenosoma Bartoš, 1949
Habrotrocha solida Donner, 1949: NEO, PAL
Habrotrocha solitaria Donner, 1949: PAL

Habrotrocha sollicita Donner, 1949: NEO, PAL
Habrotrocha soror Donner, 1950: NEO, PAL
Habrotrocha spicula Bryce, 1913: AFR, AUS, ORI, PAL
Syn.: Mniobia cornuta Bartoš, 1938
Habrotrocha stenochlaena De Koning, 1947: PAL
Habrotrocha stenostephana Schulte, 1954: AFR, PAL
Habrotrocha strangulata Murray, 1911: AUS, PAL
Habrotrocha subtilis Donner, 1965: PAL
Habrotrocha sylvestris Bryce, 1915: AUS, NEO, PAL
Habrotrocha thermalis Pax & Wulfert, 1942: PAL
Habrotrocha thienemanni Hauer, 1924
Habrotrocha thienemanni rubella Donner, 1951: PAL
Habrotrocha thienemanni thienemanni Hauer, 1924: PAL
Habrotrocha tranquilla Milne, 1916: AFR, AUS, PAL
Habrotrocha tridens (Milne, 1886)
Macrotrachela tridens Milne, 1886
Habrotrocha tridens globigera Donner, 1950: AFR, AUS, PAL
Habrotrocha tridens tridens (Milne, 1886): ANT, AUS, NEA, NEO, PAL
Habrotrocha tridentata De Koning, 1947: PAL
Habrotrocha trilobata Bartoš, 1948: PAL
Habrotrocha tripus (Murray, 1907): AUS, NEO, PAL
Callidina tripus Murray, 1907
Habrotrocha valida Milne, 1916: AFR
Habrotrocha vicina Donner, 1980: NEO
Habrotrocha visa Donner, 1954: PAL
Harringia de Beauchamp, 1912
Harringia eupoda (Gosse, 1887): NEA, PAL

Asplanchna eupoda Gosse, 1887
Harringia rousseleti de Beauchamp, 1912: NEA, NEO, ORI, PAL
Henoceros Milne, 1916
Henoceros caudatus Hauer, 1937: ORI

Henoceros falcatus (Milne, 1916): AFR, NEO, ORI, PAL
Monoceros falcatus Milne, 1916
Hexarthra Schmarda, 1854
Pedalia Barrois, 1878

Pedalion Hudson, 1871 non Swainson, 1839; non Solier, 1847; non Buckton, 1903
Hexarthra brandorffi Koste, 1977: NEA, NEO
Hexarthra bulgarica (Wiszniewski, 1933)
Pedalia bulgarica Wiszniewski, 1933
Hexarthra bulgarica bulgarica (Wiszniewski, 1933): AFR, PAL

Hexarthra bulgarica canadensis Dumont, Coussement & Anderson, 1978: NEA, PAL
Hexarthra bulgarica nepalensis Dumont, Coussement & Anderson, 1978: PAL
Hexarthra fennica (Levander, 1892): AFR, AUS, NEA, NEO, ORI
Pedalia fennica Levander, 1892
Hexarthra intermedia (Wiszniewski, 1929)
Pedalia intermedia Wiszniewski, 1929
Hexarthra intermedia brasiliensis Hauer, 1953: AFR, NEO (braziliensis auct.)
Hexarthra intermedia intermedia (Wiszniewski, 1929): AFR, AUS, NEA, NEO, ORI, PAL
Syn.: Pedalia insulana Hauer, 1938
Hexarthra jenkinae (de Beauchamp, 1932): AFR, AUS, NEA, PAL
Pedalia jenkinae de Beauchamp, 1932
Syn.: Pedalion libica Manfredi, 1939
Hexarthra longicornicula Turner, 1987: NEO
Hexarthra mira (Hudson, 1871): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Pedalion mira Hudson, 1871
Hexarthra mollis (Bartoš, 1948): PAL
Pedalia mollis Bartoš, 1948
Hexarthra oxyuris (Sernov, 1903): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Pedalion oxyure Sernov, 1903
Syn.: Pedalion mucronatum Daday, 1908
Hexarthra polychaeta Bennetch, 1981: AFR, NEA, PAC
Hexarthra polyodonta (Hauer, 1957)
Pedalia polyodonta Hauer, 1957
Hexarthra polyodonta jasperina Dumont, Coussement & Anderson, 1978: NEA
Hexarthra polyodonta polyodonta (Hauer, 1957): AUS, NEA, PAL
Syn.: Pedalia fennica var. medica Löffler, 1961
Hexarthra polyodonta soaplakeiensis Koste, 1977: NEA
Hexarthra propinqua (Bartoš, 1948): PAL
Pedalia propinqua Bartoš, 1948
Hexarthra reducens (Bartoš, 1948): PAL
Pedalia reducens Bartoš, 1948
Horaella Donner, 1949
Horaella brehmi Donner, 1949: AFR, AUS, ORI, PAL

Horaella thomassoni Koste, 1973: NEO
Inflatana Kutikova, 1985
Inflatana pomazkovae Kutikova, 1985: PAL (Baikal endemic)
Itura Harring & Myers, 1928
Itura aurita (Ehrenberg, 1830): AUS, NEA, NEO, ORI, PAL

Diglena aurita Ehrenberg, 1830
Syn.: Itura cayuga Harring & Myers, 1928 (synonym occasionally considered an infrasubspecific variant of
the nominal species)

Syn.: Itura intermedia Wulfert, 1935 (synonym occasionally considered an infrasubspecific variant of the
nominal species)
Itura chamadis Harring & Myers, 1928: NEA, NEO, PAL
Itura deridderae Segers, 1993: AFR, AUS
Itura myersi Wulfert, 1935: AUS, NEA, NEO, ORI, PAL
Itura symmetrica Segers, Mbogo & Dumont, 1994: AFR, AUS
Itura viridis (Stenroos, 1898): AUS, NEA, NEO, ORI, PAL
Eosphora viridis Stenroos, 1898
Kellicottia Ahlstrom, 1938
Kellicottia bostoniensis (Rousselet, 1908): NEA, NEO (introduced in PAL)

Notholca bostoniensis Rousselet, 1908
Kellicottia longispina (Kellicott, 1879): AFR, NEA, NEO, ORI, PAL (originally PAL, introduced in NEO,
NEA?)
Anuraea longispina Kellicott, 1879
Syn.: Notholca longispina heterospina Olofsson, 1917
Syn.: Notholca longispina taymirica Grese, 1955
Keratella Bory de St. Vincent, 1822
Keratella ahlstromi Russell, 1951: AUS

Keratella americana Carlin, 1943: ANT, NEA, NEO (introduced in PAL and AFR)
Syn.: Keratella gracilenta Ahlstrom, 1943
Syn.: Keratella lenzi caudata Koste, 1972
Keratella armadura Stemberger, 1990: NEA
Keratella australis B‘rziņÓ, 1963: AUS
Syn.: Keratella quadrata australis B‘rziņÓ, 1963
Keratella canadensis B‘rziņÓ, 1954: NEA
Keratella cochlearis (Gosse, 1851) (The taxonomy of K. cochlearis is particularly confused. Validation of the
numerous proposed subspecies and infrasubspecific variants awaits a thorough revision of the taxon)
Anuraea cochlearis Gosse, 1851
Keratella cochlearis cochlearis (Gosse, 1851): AFR, ANT, AUS, NEA, NEO, ORI, PAL
Syn.: Anuraea cochlearis recurvispina Jägerskiöld, 1894 (synonym occasionally considered an infrasub-
specific variant (f. recurvispina) of K. cochlearis)
Syn.: Anuraea stipitata Ehrenberg, 1838
Keratella cochlearis pachyacantha Thomasson, 1980: AFR, NEO
Keratella cochlearis polaris De Smet & Bafort, 1990: NEA
Keratella crassa Ahlstrom, 1943: NEA
Keratella cruciformis (Thompson, 1892): NEA, PAL (brackish water - marine)
Anuraea cruciformis Thompson, 1892
Syn.: Keratella wirketissi Kutikova, 1970
Keratella earlinae Ahlstrom, 1943: NEA, NEO
Keratella edmondsoni Ahlstrom, 1943: ORI
Keratella eichwaldi (Levander, 1894): PAL (brackish water - marine)
Anuraea eichwaldi Levander, 1894

Syn.: Keratella kamtschatica Kutikova, 1970
Keratella hiemalis Carlin, 1943: NEA, PAL
Keratella irregularis (Lauterborn, 1898): NEA, PAL (doubtful in AFR)
Anuraea irregularis Lauterborn, 1898
Syn.: Anuraea cochlearis var. irregularis Lauterborn, 1898
Keratella javana Hauer, 1937: AFR, AUS, ORI
Syn.: Keratella carinata Russell, 1950
Syn.: Keratella cochlearis var. javana Hauer, 1938
Keratella kostei Paggi, 1981: ANT, NEO (Patagonia, the Falkland Islands and South Georgia Island)
Syn.: Keratella heywoodi Dartnall, 2005
Keratella lenzi Hauer, 1953: AFR, NEA, NEO, ORI
Keratella maliensis Koste & Tobias, 1987: AFR
Keratella mexicana Kutikova & Silva-Briano, 1995: NEO
Keratella mixta (Oparina-Charitonova, 1924): NEA, ORI, PAL
Anuraea mixta Oparina-Charitonova, 1924
Syn.: Anuraea cochlearis var. mixta Oparina-Charitonova, 1924
Syn.: Keratella mixta ahlstromiella B‘rziņÓ, 1961
Keratella mongoliana Segers & Rong, 1998: PAL (East Palaearctic)
Keratella morenoi Modenutti, Diéguez & Segers, 1998: NEO
Keratella nhamundaiensis Koste, 1982: NEO
Syn.: Keratella nhamunda Koste & Robertson, 1983
Keratella ona Boltovskoy & Urrejola, 1977: NEO (Tierra del Fuego)
Keratella paludosa (Lucks, 1912): AFR, NEA, PAL
Anuraea paludosa Lucks, 1912
Keratella procurva (Thorpe, 1891)
Anuraea procurva Thorpe, 1891
Keratella procurva haterumensis Sudzuki, 1992 (species inquirenda)
Keratella procurva procurva (Thorpe, 1891): AFR, AUS, NEO, ORI, PAL
Keratella procurva robusta Koste & Shiel, 1980: AUS
Keratella quadrata (Müller, 1786) (The taxonomy of K quadrata is particularly confused. Validation of the
numerous proposed subspecies and infrasubspecific variants awaits a thorough revision of the taxon)
Brachionus quadratus Müller, 1786
Keratella quadrata dispersa Carlin, 1943: AFR, AUS, NEA, PAL (subspecies described as, and occasion-
ally considered an infrasubspecific variant of K. quadrata)
Keratella quadrata quadrata (Müller, 1786): AFR, AUS, NEA, NEO, ORI, PAL
Syn.: Anuraea aculeata Ehrenberg, 1832
Syn.: Keratella quadrata neali B‘rziņÓ, 1961
Syn.: Keratella quadrata valgoides Edmonson & Hutchinson, 1934
Keratella reducta (Huber-Pestalozzi, 1929): AFR (South Africa)
Anuraea reducta Huber-Pestalozzi, 1929
Syn.: Keratella tetracera Hutchinson, 1961
Keratella sancta Russell, 1944: ANT, AUS
Keratella serrulata (Ehrenberg, 1838): AFR, AUS, NEA, NEO, ORI, PAL
Anuraea serrulata Ehrenberg, 1838
Syn.: Anuraea falculata Ehrenberg, 1838
Keratella shieli Koste, 1979: AUS

Keratella sinensis Segers & Wang, 1997: PAL (East Palaearctic)
Keratella slacki B‘rziņÓ, 1963: AUS
Keratella taksinensis Chittapun, Pholpunthin & Segers, 2002: ORI
Keratella taurocephala Myers, 1938: NEA
Syn.: Keratella cochlearis var. taurocephala Myers, 1938
Keratella tecta (Gosse, 1851): AFR, AUS, NEA, NEO, ORI, PAL (Species occasionally considered an
infrasubspecific variant (f. tecta) of K. cochlearis. Taxonomy of the group is unclear)
Anuraea tecta Gosse, 1851
Keratella testudo (Ehrenberg, 1832): AFR, NEA, PAL
Anuraea testudo Ehrenberg, 1832
Syn.: Anuraea brevispina Gosse, 1854
Keratella thomassoni Hauer, 1958: NEO
Keratella ticinensis (Callerio, 1921): NEA, PAL (doubtful AFR record)
Anuraea ticinensis Callerio, 1921
Syn.: Anuraea aculeata var. ticinensis Callerio, 1921
Keratella trapezoida Zhuge & Huang, 1998: PAL (East Palaearctic)
Keratella tropica (Apstein, 1907): AFR, AUS, NEA, NEO, ORI, PAL
Anuraea tropica Apstein, 1907
Syn.: Keratella quadrata valga asymmetrica Uéno, 1938
Syn.: Anuraea valga var. tropica Apstein, 1907
Keratella valdiviensis Thomasson, 1957 (species inquirenda. Raised to species rank by Hauer (1938), consid-
ered an infrasubspecific variant of K. cochlearis by Koste (1978))
Syn.: Keratella cochlearis f. valdiviensis Thomasson, 1957
Keratella valga (Ehrenberg, 1834): AFR, ANT, AUS, NEA, NEO, ORI, PAL (commonly confused with
K. tropica)
Anuraea valga Ehrenberg, 1834
Keratella wangi Zhuge & Huang, 1997: PAL
Keratella yamana Boltovskoy & Urrejola, 1977: NEO (Tierra del Fuego)
Keratella zhugeae Segers & Rong, 1998: PAL (East Palaearctic)
Kostea De Smet, 1997
Kostea wockei (Koste, 1961): AFR, NEO, PAL

Paradicranophorus wockei Koste, 1961
Lacinularia Schweigger, 1826
Megalotrocha Bory de St. Vincent, 1826

Strophosphaera Poggenpol, 1872
Lacinularia elliptica Shephard, 1897: AFR, AUS, NEA, NEO, ORI, PAL
Syn.: Lacinularia racemovata Thorpe, 1893
Lacinularia elongata Shephard, 1896 (species inquirenda)
Lacinularia flosculosa (Müller, 1773): AUS, NEA, NEO, ORI, PAL (For a full account on the synonymy
of L. flosculosa see Harring (1913))

Vorticella flosculosa Müller, 1773
Lacinularia ismailoviensis (Poggenpol, 1872): AUS, NEA, NEO, PAL
Strophoshaera ismailoviensis Poggenpol, 1872
Syn.: Lacinularia natans Western, 1891
Lacinularia megalotrocha Thorpe, 1893 (species inquirenda)
Lacinularia pedunculata Hudson, 1889 (species inquirenda)
Lacinularia reticulata Anderson & Shephard, 1892 (species inquirenda)
Lacinularia striolata Shephard, 1899 (species inquirenda)
Lecane Nitzsch, 1827
Cathypna Hudson & Gosse, 1886

Diarthra Daday, 1897
Distyla auct.
Hemimonostyla Bartoš, 1959
Monostyla Ehrenberg, 1830
Lecane abanica Segers, 1994: AFR, PAL

Lecane acanthinula (Hauer, 1938): ORI (extends West to Oman)
Monostyla acanthinula Hauer, 1938
Lecane aculeata (Jakubski, 1912): AFR, AUS, NEO, ORI, PAC, PAL
Distyla aculeata Jakubski, 1912
Syn.: Lecane curvicerata Yamamoto, 1951
Lecane acus (Harring, 1913): AUS, NEA, NEO, ORI, PAL (taxonomy unclear)
Monostyla acus Harring, 1913
Lecane aeganea Harring, 1914: AFR, AUS, NEA, NEO, ORI
Lecane agilis (Bryce, 1892): AFR, NEA, NEO, PAC, PAL
Distyla agilis Bryce, 1892
Lecane amazonica (Murray, 1913): NEO
Monostyla amazonica Murray, 1913
Syn.: Lecane murrayi Hauer, 1965 (non Korde (1927))
Lecane arcuata (Bryce, 1891): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Monostyla arcuata Bryce, 1891
Syn.: Lecane piepelsi De Smet & Bafort, 1990
Lecane arcula Harring, 1914: AFR, AUS, NEA, NEO, ORI, PAL
Syn.: Lecane strandi B‘rziņÓ, 1943
Lecane armata Thomasson, 1971: NEO, ORI
Lecane aspasia Myers, 1917: NEA, NEO, ORI, PAL
Lecane asymmetrica (Murray, 1913): NEO (Caribbean)
Monostyla asymmetrica Murray, 1913
Syn.: Lecane janetzkyi Koste, Janetzky & Vareshi, 1991
Lecane baimaii Sanoamuang & Savatenalinton, 1999: AFR, ORI
Lecane batillifer (Murray, 1913): AUS, ORI
Monostyla batillifer Murray, 1913
Lecane bidactyla (Sudzuki, 1992) (species inquirenda: poorly contracted and completely unrecognisable speci-
mens)

Monostyla bidactyla Sudzuki, 1992
Lecane bifastigata Hauer, 1938: ORI, PAL
Lecane bifurca (Bryce, 1892): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Monostyla bifurca Bryce, 1892
Syn.: Lecane bifurca entomeB‘rziņÓ, 1982
Syn.: Monostyla crypta Hauer, 1938
Syn.: Notommata monostylaeformis Stenroos, 1898
Lecane blachei B‘rziņÓ, 1973: ORI
Lecane boettgeri Koste, 1986: NEO
Lecane boliviana Segers, 1994: NEO
Lecane boorali Koste & Shiel, 1983: AUS
Lecane branchicola (Piovanelli, 1903): PAL (Epizoic on gills of freshwater crabs)
Distyla branchicola Piovanelli, 1903
Syn.: Lecane piovanellii Bartoš, 1957
Lecane braumi Koste, 1988: AFR, AUS, ORI
Lecane braziliensis Segers, 1993: NEO
Lecane broaensis Segers & Dumont, 1995: NEO
Lecane bryophila Koniar, 1957: PAL
Lecane bulla (Gosse, 1851) (Morphologically variable taxon requiring further study)
Monostyla bulla Gosse, 1851
Lecane bulla bulla (Gosse, 1851): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Syn.: Monostyla bipes Stokes, 1896
Syn.: Monostyla bulla constricta Sudzuki, 1992
Syn.: Monostyla bulla dentata Sudzuki, 1992
Syn.: Lecane bulla diana Abdullaev, 1989
Syn.: Lecane bulla kutikovae Naberezhniyi & Irmasheva, 1975
Syn.: Monostyla goniata Harring & Myers, 1926 (synonym occasionally considered an infrasubspecific
variant of L. bulla)
Syn.: Monostyla ozolini B‘rziņÓ, 1943
Syn.: Lecane physalis Wulfert, 1939 (incl. L.styrax longistyla (Weisig, 1928))
Lecane bulla diabolica (Hauer, 1936): ORI
Monostyla bulla f. diabolica Hauer, 1936
Lecane calcaria Harring & Myers, 1926: NEA, ORI
Lecane candida Harring & Myers, 1926: NEA
Lecane clara (Bryce, 1892): AFR, NEA, NEO, PAC, PAL
Distyla clara Bryce, 1892
Syn.: Diglena beauchampi von Hofsten, 1923
Syn.: Proales longidactyla Edmondson, 1948
Lecane climacois Harring & Myers, 1926: AFR, NEA
Lecane closterocerca (Schmarda, 1859): AFR, ANT, AUS, NEA, NEO, ORI, PAC, PAL
Monostyla closterocerca Schmarda, 1859
Syn.: Monostyla brodskii Muraveisky, 1935
Syn.: Lecane closterocerca amazonica Koste, 1978
Syn.: Monostyla eichsfeldica Künne, 1926
Syn.: Monostyla latvica B‘rziņÓ, 1943
Syn.: Lecane wulferti Hauer, 1956

Lecane copeis (Harring & Myers, 1926): NEA, NEO, PAL
Monostyla copeis Harring & Myers, 1926
Syn.: Lecane eupsammophila Koste, 1992
Lecane cornuta (Müller, 1786): NEA, NEO, PAL (Morphologically variable taxon requiring further study)
Trichoda cornuta Müller, 1786
Syn.: Monostyla cornuta oidipus Hauer, 1956
Syn.: Monostyla robusta Stokes, 1896
Syn.: Monostyla rotunda Fadeev, 1927
Syn.: Monostyla rotundata Jakubski, 1914
Lecane crenata (Harring, 1913): AFR, AUS, NEA, NEO, ORI, PAL (Taxonomy unclear)
Monostyla crenata Harring, 1913
Lecane crepida Harring, 1914: AFR, AUS, NEA, NEO, ORI, PAL
Syn.: Lecane crepida longidactyla Koste, 1972
Syn.: Lecane neali Wulfert, 1966
Syn.: Lecane vasisthi Sharma, 1980
Lecane curvicornis (Murray, 1913): AFR, AUS, NEA, NEO, ORI, PAL
Cathypna curvicornis Murray, 1913
Syn.: Lecane acronycha Harring & Myers, 1926
Syn.: Lecane bondi Edmondson, 1934
Syn.: Lecane chankensis Bogoslovsky, 1958
Syn.: Lecane curvilinealis Arora, 1965
Syn.: Lecane longidactyla Arora, 1965 (non Edmondson (1948))
Syn.: Lecane triloba Yamamoto, 1951
Syn.: Lecane zwaiensis Bryce, 1931
Lecane decipiens (Murray, 1913): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Monostyla decipiens Murray, 1913
Lecane depressa (Bryce, 1891): NEA, PAL
Distyla depressa Bryce, 1891
Syn.: Cathypna amban Stewart, 1908
Syn.: Cathypna brachydactyla Stenroos, 1898
Syn.: Distyla truncata Leissling, 1914 (non Turner (1892))
Syn.: Lecane truncata Yamamoto, 1953 (non Leissling (1914) nec Turner (1892))
Syn.: Lecane tudicola Harring & Myers, 1926
Lecane deridderae Koste, 1972: NEO
Lecane difficilis Segers & Pourriot, 1997: NEA, NEO
Lecane donneri Chengalath & Mulamoottil, 1974: AFR, NEA, ORI
Lecane donyanaensis Mazuelos & Segers, 1994: PAL (Endemic to the Doñana National Park)
Lecane doryssa Harring, 1914: AFR, AUS, NEO, ORI, PAL
Lecane dumonti Segers, 1993: AFR, NEO
Lecane dysorata Myers, 1942: NEA (Doubtful PAL record)
Syn.: Lecane dysoarata auct.
Lecane elasma Harring & Myers, 1926: NEA, PAL
Lecane elegans Harring, 1914: AFR, NEO, ORI, PAC, PAL
Lecane elongata Harring & Myers, 1926: NEA, PAL
Syn.: Lecane lebedevae Abdullaev, 1989
Lecane elsa Hauer, 1931: NEA, PAL

Lecane enowi Segers & Mertens, 1997: AFR, ORI
Lecane ercodes Harring, 1914: AFR, NEA, NEO, ORI
Lecane eswari Dhanapathi, 1976: AFR, ORI
Lecane eutarsa Harring & Myers, 1926: NEO (Doubtful PAL record)
Syn.: Lecane amazoniana Koste & Robertson, 1983
Syn.: Lecane amazonica Koste & Böttger, 1992 (non Murray (1913))
Lecane eylesi Russell, 1953: AUS (Tasmania and New Zealand)
Syn.: Lecane tasmaniensis Shiel & Koste, 1985
Lecane fadeevi (Neiswestnova-Shadina, 1935): PAL
Monostyla fadeewi Neiswestnova-Shadina, 1935
Syn.: Lecane fadeewi Voigt, 1957
Lecane flabellata Edmondson, 1936: NEA
Lecane flexilis (Gosse, 1886): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Distyla flexilis Gosse, 1886
Syn.: Cathypna brevis Murray, 1913
Syn.: Lecane glypta Harring & Myers, 1926 (Taxonomy unclear. Synonym occasionally considered an infra-
subspecific variant of L. flexilis)
Syn.: Distyla lipara Gosse, 1887
Lecane formosa Harring & Myers, 1926: NEA
Lecane furcata (Murray, 1913): AUS, NEA, NEO, ORI, PAL
Monostyla furcata Murray, 1913
Syn.: Monostyla elachis Harring & Myers, 1926
Syn.: Monostyla mologensis Bogoslovski, 1935
Syn.: Monostyla ovalis Jakubski, 1914
Syn.: Monostyla tethis Harring & Myers, 1926
Syn.: Lecane vanoyei De Ridder, 1960
Lecane galeata (Bryce, 1892): NEA, PAL
Monostyla galeata Bryce, 1892
Syn.: Lecane beningi Tarnogradsky, 1961
Syn.: Monostyla parva Daday, 1897
Syn.: Monostyla pygmaea Daday, 1897
Syn.: Monostyla turbo Murray, 1913
Lecane gallagherorum Segers, 1997: NEA
Lecane gillardi B‘rziņÓ, 1960: AFR (Madagascar)
Lecane grandis (Murray, 1913): AUS, NEA, NEO, ORI, PAL
Cathypna grandis Murray, 1913
Lecane gwileti (Tarnogradski, 1930): PAL
Monostyla gwileti Tarnogradski, 1930
Syn.: Monostyla kieferi Hauer, 1931
Lecane haliclysta Harring & Myers, 1926: AFR, AUS, NEA, NEO, ORI, PAL
Syn.: Lecane stichaeoides Hauer, 1938
Lecane halsei Segers & Shiel, 2003: AUS
Lecane hamata (Stokes, 1896): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Monostyla hamata Stokes, 1896
Syn.: Lecane hamata victoriensis Koste & Shiel, 1980
Lecane hastata (Murray, 1913): AFR, AUS, NEA, NEO, ORI, PAC, PAL

Cathypna hastata Murray, 1913
Syn.: Lecane hegurensis Yamamoto, 1951
Lecane herzigi Koste, Shiel & Tan, 1988: AUS
Lecane hornemanni (Ehrenberg, 1834): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Euchlanis hornemanni Ehrenberg, 1834
Syn.: Lecane camptica B‘rziņÓ, 1982
Syn.: Lecane ceylonensis Chengalath & Fernando, 1973
Syn.: Lecane lamiranoensis B‘rziņÓ, 1982
Syn.: Lecane nodosa Hauer, 1938
Lecane imbricata Carlin, 1939: PAL
Lecane inconspicua Segers & Dumont, 1993: PAL
Lecane inermis (Bryce, 1892): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Distyla inermis Bryce, 1892
Syn.: Lecane amorpha Harring, 1914
Syn.: Lecane supinoi Manfredi, 1929
Lecane infula Harring & Myers, 1926: NEA, PAL
Lecane inopinata Harring & Myers, 1926: AFR, AUS, NEA, NEO, ORI, PAC, PAL
Lecane inquieta Myers, 1936: NEA (Doubtful PAL record)
Lecane intrasinuata (Olofsson, 1917): NEA, PAL
Cathypna intrasinuata Olofsson, 1917
Syn.: Lecane ephestra Harring, 1921
Syn.: Lecane mylacris Harring & Myers, 1926
Lecane isanensis Sanoamuang & Savatenalinton, 2001: ORI
Lecane ivli (Wiszniewski, 1935): PAL
Monostyla ivli Wiszniewski, 1935
Syn.: Monostyla vargai Török, 1935
Lecane jaintiaensis Sharma, 1987: ORI
Lecane jessupi Harring, 1921: NEA
Lecane junki Koste, 1975: ORI
Lecane kluchor Tarnogradski, 1930: PAL
Syn.: Lecane diadema Hauer, 1931
Lecane kunthuleensis Chittapun, Pholpunthin & Segers, 2003: ORI
Lecane kutikowa Koste, 1972: NEO
Lecane lamellata (Daday, 1893): NEA, PAL
Monostyla lamellata Daday, 1893
Syn.: Monostyla appendiculata Skorikov, 1898 (non Levander (1894))
Lecane lateralis Sharma, 1978: AFR, AUS, ORI
Lecane latissima Yamamoto, 1955: AUS, NEA, PAL
Syn.: Lecane kostei De Ridder, 1966
Syn.: Cathypna rotundata Olofsson, 1918 (non Jakubski (1914))
Lecane lauterborni Hauer, 1924: NEA, NEO, PAC, PAL
Lecane leontina (Turner, 1892): AFR, AUS, NEA, NEO, ORI, PAL (Morphologically variable taxon, prob-
ably concealing cryptic species)
Cathypna leontina Turner, 1892
Syn.: Cathypna biloba Daday, 1905
Syn.: Cathypna leontina bisinuata Daday, 1905

Syn.: Cathypna macrodactyla Daday, 1898
Syn.: Cathypna scutaria Stokes, 1897
Syn.: Lecane thomassoni Wulfert, 1965
Lecane leura Myers, 1942: NEA
Lecane levistyla (Olofsson, 1917): NEA, PAL
Cathypna levistyla Olofsson, 1917
Syn.: Lecane scobis Harring & Myers, 1926
Lecane ligona (Dunlop, 1901): NEA, NEO, ORI, PAL
Cathypna ligona Dunlop, 1901
Syn.: Lecane abnobensis Hauer, 1929 (synonym occasionally considered an infrasubspecific variant of L.
ligona)
Syn.: Lecane jessupi Harring, 1921 (synonym occasionally considered an infrasubspecific variant of L. ligona)
Lecane ludwigii (Eckstein, 1883): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Distyla ludwigii Eckstein, 1883
Syn.: Cathypna appendiculata Daday, 1901 (non Skorikov (1898) nec Levander (1894))
Syn.: Diplax ornata Daday, 1897
Lecane luna (Müller, 1776): AFR, AUS, NEA, NEO, ORI, PAC, PAL (Poorly contracted specimens are
sometimes erroneously identified as L. presumpta Ahlstrom, 1938, and this name is accordingly listed as subspecies or
infrasubspecific variant of L. luna. However, L. presumpta is an established synonym of L. papuana)
Cercaria luna Müller, 1776
Syn.: Lecane dorsicalis Arora, 1965
Syn.: Lecane jobloti Bory de St. Vincent, 1827
Syn.: Lecane luna balatonica Varga, 1945
Syn.: Lecane submagna De Ridder, 1960
Lecane lunaris (Ehrenberg, 1832): AFR, ANT, AUS, NEA, NEO, ORI, PAC, PAL (Morphologically
variable taxon, probably concealing cryptic species)
Monostyla lunaris Ehrenberg, 1832
Syn.: Monostyla constricta Murray, 1913
Syn.: Lecane lunaris arthrodactyla B‘rziņÓ, 1982
Syn.: Lecane lunaris australis B‘rziņÓ, 1982
Syn.: Monostyla lunaris obserata Steinecke, 1916
Syn.: Monostyla quennerstedti Bergendal, 1892
Syn.: Monostyla sylvatica Harring, 1913
Syn.: Monostyla virga Harring, 1914
Lecane lungae Savatenalinton & Segers, 2005: ORI
Lecane marchantaria Koste & Robertson, 1983: NEO
Lecane margalefi De Manuel, 1994: PAL (Endemic to Mallorca (Spain))
Lecane margarethae Segers, 1991: NEA, NEO (Caribbean)
Syn.: Lecane deridderae José de Paggi, 1989 (non Koste (1972))
Lecane marshi Harring, 1914: NEO
Lecane melini Thomasson, 1953: NEO
Lecane minuta Segers, 1994: ORI
Lecane mira (Murray, 1913): NEA, PAL
Cathypna mira Murray, 1913
Syn.: Lecane islandica De Ridder, 1967
Lecane mitella (Myers, 1936): NEA

Monostyla mitella Myers, 1936
Lecane mitis Harring & Myers, 1926: NEA, NEO, ORI
Lecane monostyla (Daday, 1897): AFR, AUS, NEA, NEO, ORI, PAC, PAL (Some unverifiable records
from Europe (Spain, Poland) exist; presence in Europe of this unmistakable warm-water species is likely)
Monostyla monostyla Daday, 1897
Syn.: Monostyla spinifera Idelson, 1924
Lecane mucronata Harring & Myers, 1926: NEA
Lecane myersi Segers, 1993: AFR, NEA, NEO
Syn.: Monostyla ornata Harring & Myers, 1926 (non Daday (1897))
Lecane namatai Segers & Mertens, 1997: AFR
Lecane nana (Murray, 1913): AFR, AUS, NEA, NEO, ORI, PAL
Cathypna nana Murray, 1913
Lecane nelsoni Segers, 1994: AFR, NEO, ORI
Lecane nigeriensis Segers, 1993: AFR, ORI
Lecane niothis Harring & Myers, 1926: NEA (Doubtful PAL record)
Lecane nitida (Murray, 1913): AFR, AUS, NEO, ORI
Cathypna nitida Murray, 1913
Syn.: Cathypna lofuana Murray, 1913
Syn.: Lecane tesselata Arora, 1965
Lecane niwati Segers, Kothetip & Sanoamuang, 2004: ORI
Lecane noobijupi Segers & Shiel, 2003: AUS
Lecane nwadiaroi Segers, 1993: AFR
Lecane obtusa (Murray, 1913): AFR, AUS, NEA, NEO, ORI, PAL
Monostyla obtusa Murray, 1913
Syn.: Monostyla vastita Harring & Myers, 1926
Lecane ohioensis (Herrick, 1885): AFR, AUS, NEA, NEO, ORI, PAC, PAL (see L. ludwigii)
Distyla ohioensis Herrick, 1885
Syn.: Cathypna appendiculata Levander, 1894
Syn.: Distyla ichthyoura Anderson & Shephard, 1892
Syn.: Cathypna jorroi Arévalo, 1918 (synonym occasionally considered an infrasubspecific variant of L.
ohioensis)
Lecane opias (Harring & Myers, 1926): NEA, ORI, PAL
Monostyla opias Harring & Myers, 1926
Lecane ordwayi Bienert, 1986: NEA
Lecane palinacis Harring & Myers, 1926: NEA, ORI, PAC
Lecane papuana (Murray, 1913): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Cathypna papuana Murray, 1913
Syn.: Lecane kasumiensis Sudzuki, 1998
Syn.: Lecane presumpta Ahlstrom, 1938
Syn.: Lecane yamunensis Novotná-Dvorakova, 1963
Lecane paradoxa (Steinecke, 1916): PAL
Monostyla paradoxa Steinecke, 1916
Syn.: Lecane hoffmanni De Ridder, 1960
Syn.: Lecane ozensis Yamamoto, 1953
Lecane pawlowskii Wulfert, 1966: ORI
Lecane paxiana Hauer, 1940: AFR, ORI, PAC, PAL

Lecane pelatis Harring & Myers, 1926: NEA
Lecane perplexa (Ahlstrom, 1938): AFR, NEA, NEO, PAC, PAL
Monostyla perplexa Ahlstrom, 1938
Lecane perpusilla (Hauer, 1929): AFR, NEA, PAC, PAL
Monostyla perpusilla Hauer, 1929
Syn.: Lecane brevita Myers, 1942
Lecane pertica Harring & Myers, 1926: AFR, AUS, NEA, NEO, ORI
Lecane plesia Myers, 1936: AFR, NEA, NEO, PAL
Syn.: Lecane jana Abdullaev, 1989
Syn.: Proales namibiensis Koste & Brain, 1993
Lecane pluto Segers & Mertens, 1997: AFR
Lecane pomiformis Edmondson, 1938: NEA, NEO
Syn.: Lecane paraclosterocerca Pennak, 1939
Lecane proiecta Hauer, 1956: NEO
Lecane psammophila (Wiszniewski, 1932): PAL
Monostyla psammophila Wiszniewski, 1932
Lecane pumila (Rousselet, 1906): AFR, AUS, ORI, PAL
Notommata pumila Rousselet, 1906
Lecane punctata (Murray, 1913): AFR, NEA, NEO, ORI, PAL
Monostyla punctata Murray, 1913
Syn.: Lecane aguessei De Ridder, 1960
Syn.: Monostyla harringi Ahlstrom, 1934
Lecane pusilla Harring, 1914: AFR, AUS, NEO, ORI, PAL
Lecane pustulosa Myers, 1938: NEA
Lecane pycina Harring & Myers, 1926: NEA
Lecane pyriformis (Daday, 1905): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Monostyla pyriformis Daday, 1905
Lecane pyrrha Harring & Myers, 1926: NEA (Doubtful PAL record)
Lecane quadridentata (Ehrenberg, 1830): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Monostyla quadridentata Ehrenberg, 1830
Syn.: Monostyla bicornis Daday, 1897
Syn.: Lecane quadridentata arthrodactyla B‘rziņÓ, 1982
Syn.: Monostyla sexidentata Van Oye, 1926
Lecane remanei Hauer, 1964: NEO
Lecane rhacois Harring & Myers, 1926: NEA
Lecane rhenana Hauer, 1929: AFR, NEO, ORI, PAL
Lecane rhopalura (Harring & Myers, 1926): NEA, NEO, PAL
Monostyla rhopalura Harring & Myers, 1926
Syn.: Lecane aspersa Kutikova & Arov, 1985
Syn.: Monostyla pideis Harring & Myers, 1926
Lecane rhytida Harring & Myers, 1926: AFR, AUS, NEA, NEO, ORI
Lecane robertsonae Segers, 1993: AUS, NEO, ORI
Syn.: Lecane aspasia amazonica Koste & Robertson, 1983
Lecane rudescui Hauer, 1965: NEO
Lecane rugosa (Harring, 1914): NEO (occasionally considered an infrasubspecific variant of L. furcata.,PAL and
AUS record require confirmation)

Monostyla rugosa Harring, 1914
Lecane ruttneri Hauer, 1938: AFR, NEO, ORI
Lecane sagula Harring & Myers, 1926: NEA, NEO (Doubtful PAL record)
Lecane satyrus Harring & Myers, 1926: NEA, PAL (Northeast North America and Southeast Palaearctic)
Lecane schraederi Wulfert, 1966: ORI
Lecane scutata (Harring & Myers, 1926): AUS, NEA, ORI, PAL (frequently confused with L. perplexa)
Monostyla scutata Harring & Myers, 1926
Syn.: Monostyla stroeszneri Török, 1935
Lecane segersi Sanoamuang, 1996: ORI
Lecane serrata (Hauer, 1938): AFR, AUS, ORI
Monostyla serrata Hauer, 1938
Lecane shieli Segers & Sanoamuang, 1994: ORI
Lecane signifera (Jennings, 1896)
Distyla signifera Jennings, 1896
Lecane signifera ploenensis (Voigt, 1902): AFR, AUS, NEA, NEO, ORI, PAL
Distyla signifera ploenensis Voigt, 1902
Syn.: Lecane signifera glandulosa Sudzuki, 1991
Lecane signifera signifera (Jennings, 1896): NEA
Distyla signifera signifera Jennings, 1896
Syn.: Lecane aquila Harring & Myers, 1926
Lecane simonneae Segers, 1993: AFR, ORI
Lecane sinuata (Hauer, 1938): AUS, ORI
Monostyla sinuate Hauer, 1938
Lecane sola Hauer, 1936: NEO, ORI
Lecane solfatara (Hauer, 1938): ORI
Monostyla solfatara Hauer, 1938
Lecane spiniventris Segers, 1994: ORI
Lecane spinulifera Edmondson, 1935: NEA, NEO
Syn.: Lecane aliger Nogrady, 1983
Syn.: Monostyla spinifera Edmondson, 1934 (non Idelson (1924))
Lecane stenroosi (Meissner, 1908): AFR, AUS, NEA, NEO, ORI, PAL
Monostyla stenroosi Meissner, 1908
Syn.: Monostyla bicornis Stenroos, 1898 (non Daday (1897))
Syn.: Hemimonostyla stenroosi tutiurensis Sudzuki, 1998
Lecane stephensae (Hutchinson, 1931): AFR, ORI
Monostyla stephensae Hutchinson, 1931
Lecane stichaea Harring, 1913: AFR, AUS, NEA, NEO, ORI, PAC, PAL
Syn.: Lecane methoria Harring & Myers, 1926
Syn.: Lecane saginata Harring & Myers, 1926
Lecane stichoclysta Segers, 1993: AFR, ORI
Lecane stokesii (Pell, 1890): NEA, PAL
Cathypna stokesi Pell, 1890
Lecane styrax (Harring & Myers, 1926): NEA, NEO
Monostyla styrax Harring & Myers, 1926
Lecane subtilis Harring & Myers, 1926: AFR, NEA, NEO, PAL
Syn.: Lecane apatinensis Zivkovic, 1987

Syn.: Lecane murrayi Korde, 1927
Lecane subulata (Harring & Myers, 1926): AFR, NEA, NEO, PAC, PAL
Monostyla subulata Harring & Myers, 1926
Syn.: Lecane rylovi Tarnogradski, 1961
Lecane superaculeata Sanoamuang & Segers, 1997: ORI
Lecane sverigis Ahlstrom, 1934: NEA
Lecane sylviae Segers, 1993: AFR
Lecane symoensi De Ridder, 1981: AFR
Lecane sympoda Hauer, 1929: AFR, PAL
Lecane syngenes (Hauer, 1938): AFR, AUS, NEO, ORI
Monostyla syngenes Hauer, 1938
Lecane tabida Harring & Myers, 1926: AFR, NEA, ORI
Lecane tabulifera Edmondson, 1936: NEA
Lecane tanganyikae Segers & Baribwegure, 1996: AFR
Lecane tenua Myers, 1936: NEA
Lecane tenuiseta Harring, 1914: AFR, AUS, NEA, NEO, ORI, PAL
Syn.: Lecane punctata Carlin-Nilson, 1934 (non Murray (1913). Occasionally listed as an infrasubspecific
variant of the nominal taxon, as var. punctata Carlin-Nilsson, 1934)
Lecane thailandensis Segers & Sanoamuang, 1994: ORI
Lecane thalera (Harring & Myers, 1926): AUS, NEA, NEO, ORI, PAL (Taxonomic separation with L.
lamellata unclear; possible hybrids reported)
Monostyla thalera Harring & Myers, 1926
Syn.: Monostyla conspicua Hauer, 1936
Syn.: Lecane paradecipiens Nayar, 1968
Lecane thienemanni (Hauer, 1938): AFR, AUS, NEO, ORI
Monostyla thienemanni Hauer, 1938
Lecane tryphema Harring & Myers, 1926: NEA, PAL
Lecane uenoi Yamamoto, 1951: NEO, ORI, PAL
Lecane undulata Hauer, 1938: AFR, AUS, NEA, NEO, ORI, PAL
Lecane unguitata (Fadeev, 1925): AFR, AUS, ORI, PAL
Monostyla unguitata Fadeev, 1925
Syn.: Lecane unguitata africana Koste, 1979
Lecane ungulata (Gosse, 1887): AFR, AUS, NEA, NEO, ORI, PAL
Cathypna ungulata Gosse, 1887
Syn.: Lecane donneriana Dhanapathi, 1976
Syn.: Lecane fracida B‘rziņÓ, 1982
Syn.: Cathypna magna Lucks, 1898 (non Stenroos (1898))
Syn.: Cathypna magna Stenroos, 1898
Syn.: Cathypna magna tenuior Stenroos, 1898
Syn.: Lecane ungulata australiensis Koste & Shiel, 1990
Lecane urna Nogrady, 1962: PAL
Lecane venusta Harring & Myers, 1926: NEA, PAL
Lecane verecunda Harring & Myers, 1926: NEA (Doubtful PAL record)
Lecane whitfordi (Ahlstrom, 1938): NEA, NEO
Monostyla whitfordi Ahlstrom, 1938

Lepadella Bory de St. Vincent, 1826
subgenus Heterolepadella Bartoš, 1955
Lepadella (Heterolepadella) apsicora Myers, 1934: AFR, AUS, NEA, NEO, ORI
Lepadella (Heterolepadella) cyrtopus Harring, 1914: AFR, NEA, NEO, ORI, PAL
Lepadella (Heterolepadella) ehrenbergii (Perty, 1850): AFR, AUS, NEA, NEO, ORI, PAL
Notogonia ehrenbergii Perty, 1850
Syn.: Metopidia albuferensis Arévalo, 1918
Lepadella (Heterolepadella) heterodactyla Fadeew, 1925: AFR, AUS, NEO, ORI, PAL
Syn.: Lepadella nympha Donner, 1943
Lepadella (Heterolepadella) heterostyla (Murray, 1913): AFR, AUS, NEA, NEO, ORI, PAL
Metopidia heterostyla Murray, 1914
subgenus Lepadella Bory de St. Vincent, 1826
Lepadella (Lepadella) abbei Wulfert, 1956: PAL

Lepadella (Lepadella) akrobeles Myers, 1934: AUS, NEA, ORI
Lepadella (Lepadella) acuminata (Ehrenberg, 1834): AFR, ANT, AUS, NEA, NEO, ORI, PAC, PAL
(incl. the infrasubspecific variant sexcostata Bartoš, 1965)
Metopidia acuminata Ehrenberg, 1834
Syn.: Lepadella chorea B‘rziņÓ, 1982
Syn.: Lepadella sexcostata Bartoš, 1955
Lepadella (Lepadella) adjuncta Donner, 1943: PAL
Lepadella (Lepadella) amazonica Segers, 1993: NEO
Lepadella (Lepadella) amphitropis Harring, 1916
Lepadella (L.) amphitropis amphitropis Harring, 1916: AFR, AUS, NEA, NEO, PAL
Lepadella (L.) amphitropis victoriensis B‘rziņÓ, 1982: AUS
Lepadella (Lepadella) angusta B‘rziņÓ, 1961: AUS
Lepadella (Lepadella) apsida Harring, 1916: AFR, AUS, NEA, NEO, ORI, PAL
Lepadella (Lepadella) benjamini Harring, 1916: AFR, AUS, NEA, NEO, ORI, PAL
Lepadella (Lepadella) berzinsi Segers, 1993: AFR
Syn.: Lepadella quinquecostata f. gambiensis B‘rziņÓ, 1957
Lepadella (Lepadella) beyensi De Smet, 1994: NEA
Lepadella (Lepadella) bicornis Vasisht & Battish, 1971: NEO, ORI
Lepadella (Lepadella) bidentata Voronkov, 1913: PAL
Lepadella (Lepadella) biloba Hauer, 1958: AFR, AUS, NEO, PAL
Lepadella (Lepadella) canadensis Myers, 1936: NEA
Lepadella (Lepadella) chengalathi Koste & Shiel, 1980: AUS
Lepadella (Lepadella) cornuta Koste & Shiel, 1989: AUS
Syn.: Lepadella latusinus f. mucronata Koste, 1981
Lepadella (Lepadella) costata Wulfert, 1940: PAL
Lepadella (Lepadella) costatoides Segers, 1992: AFR, AUS, NEA, NEO, ORI, PAL
Syn.: Lepadella latusinus striata Koste, 1992
Lepadella (Lepadella) cristata (Rousselet, 1893): AFR, NEA, NEO, ORI, PAL
Colurus cristatus Rousselet, 1893

Syn.: Lepadella cristata galerita B‘rziņÓ, 1982
Syn.: Metopidia mucronata Daday, 1908
Syn.: Metopidia semicarinata Lucks, 1909
Lepadella (Lepadella) cryphaea Harring, 1916: NEA, NEO, PAL
Lepadella (Lepadella) dactyliseta (Stenroos, 1898): AFR, AUS, NEA, NEO, ORI, PAL
Metopidia dactyliseta Stenroos, 1898
Syn.: Lepadella voigti Hauer, 1931
Lepadella (Lepadella) decora B‘rziņÓ, 1982: AUS
Lepadella (Lepadella) degreefi De Smet, 1989: NEA, PAC
Lepadella (Lepadella) deridderae Segers, De Smet & Bonte, 1996
Lepadella (L.) deridderae alaskae Segers, De Smet & Bonte, 1996: NEA
Lepadella (L.) deridderae deridderae Segers, De Smet & Bonte, 1996: PAL
Lepadella (Lepadella) desmeti Segers & Chittapun, 2001: NEO, ORI, PAC
Lepadella (Lepadella) discoidea Segers, 1993: AFR, AUS, ORI
Lepadella (Lepadella) donneri Koste, 1972: NEO
Lepadella (Lepadella) dorsalis Rodewald, 1937: PAL
Lepadella (Lepadella) duvigneaudi De Ridder, 1969: PAL
Lepadella (Lepadella) elliptica Wulfert, 1939: ANT, AUS, NEO, ORI, PAL
Lepadella (Lepadella) elongata Koste, 1992: NEO, ORI, PAC
Lepadella (Lepadella) eurysterna Myers, 1942: AFR, NEA, ORI, PAL
Syn.: Lepadella arabica Segers & Dumont, 1993
Lepadella (Lepadella) evaginata Rodewald, 1935: PAL
Lepadella (Lepadella) favorita Klement, 1962: PAL
Lepadella (Lepadella) gelida B‘rziņÓ, 1976: PAL
Lepadella (Lepadella) glossa Wulfert, 1960: PAL
Lepadella (Lepadella) hyalina Smirnov, 1927: PAL
Lepadella (Lepadella) imbricata Harring, 1914: AFR, NEA, NEO, ORI, PAL
Lepadella (Lepadella) intermedia Dartnall & Hollowday, 1985: ANT
Lepadella (Lepadella) koniari Bartoš, 1955: PAL
Syn.: Lepadella avernae Francez & Pourriot, 1984
Lepadella (Lepadella) kostei Wulfert, 1966: ORI
Lepadella (Lepadella) latusinus (Hilgendorf, 1899): AFR, AUS, NEA, NEO, ORI, PAL
Metopidia latusinus Hilgendorf, 1899
Lepadella (Lepadella) lindaui Koste, 1981: AFR, AUS, NEO, ORI
Lepadella (Lepadella) longiseta Myers, 1934: NEA
Lepadella (Lepadella) margalefi De Ridder, 1964: PAL
Lepadella (Lepadella) mascarensis B‘rziņÓ, 1982: AFR
Lepadella (Lepadella) mataca José de Paggi, 2001: NEO
Lepadella (Lepadella) mica Zivkovic, 1987: PAL
Lepadella (Lepadella) minorui Koste, 1981: AUS
Lepadella (Lepadella) minoruoides Koste & Robertson, 1983: AFR, NEO, ORI
Lepadella (Lepadella) minuscula Bartoš, 1955: PAL
Lepadella (Lepadella) minuta (Weber & Montet, 1918): AFR, NEO, ORI, PAC, PAL
Metopidia minuta Weber & Montet, 1918
Syn.: Metopidia parvula Montet, 1915 (non Bryce (1893))

Lepadella (Lepadella) monodi B‘rziņÓ, 1959: AFR
Lepadella (Lepadella) myersi Edmondson, 1934: NEO
Lepadella (Lepadella) nartiangensis Sharma & Sharma, 1987: ORI
Lepadella (Lepadella) neboissi B‘rziņÓ, 1960: AUS
Lepadella (Lepadella) neglecta Segers & Dumont, 1995: NEO
Lepadella (Lepadella) nympha Donner, 1943: PAL
Lepadella (Lepadella) obtusa Wang, 1961: PAL
Lepadella (Lepadella) ovalis (Müller, 1786): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Brachionus ovalis Müller, 1786
Syn.: Metopidia affinis Bergendal, 1892
Syn.: Metopidia lepadella Ehrenberg, 1832
Syn.: Mytilina lepidura Bory de St. Vincent, 1826
Syn.: Lepadella rotundata Dujardin, 1841
Syn.: Metopidia solidus Gosse, 1851
Syn.: Lepadella velazmedrani Pardo, 1934
Lepadella (Lepadella) paparoa B‘rziņÓ, 1982: AUS
Lepadella (Lepadella) parvula (Bryce, 1893): PAL
Metopidia parvula Bryce, 1893
Lepadella (Lepadella) patella (Müller, 1773) (The taxonomy and nomenclature of the variable L. patella are
particularly confused. Only the most common, probable synonyms are listed.)
Brachionus patella Müller, 1773
Lepadella (L.) patella matudai Sudzuki, 1964 (species inquirenda)
Lepadella (L.) patella oblonga (Ehrenberg, 1834): AFR, ANT, AUS, NEA, NEO, ORI, PAL
Squamella oblonga Ehrenberg, 1834
Syn.: Lepadella minor Koch-Althaus, 1963
Lepadella (L.) patella patella (Müller, 1786): AFR, ANT, AUS, NEA, NEO, ORI, PAC, PAL
Syn.: Squamella bractea Ehrenberg, 1838
Syn.: Lepadella mariae Rodewald, 1935
Syn.: Lepadella parvula De Graaf, 1956 (non Bryce (1893))
Syn.: Lepadella rotundata Dujardin, 1841
Lepadella (L.) patella persimilis De Ridder, 1961: AFR, AUS, NEA, NEO, ORI, PAL
Syn.: Lepadella buangensis B‘rziņÓ, 1982
Syn.: Metopidia similis Lucks, 1912 (non Stokes (1896))
Lepadella (Lepadella) pejleri Eriksen, 1969: PAL
Lepadella (Lepadella) pontica Althaus, 1957 (marine)
Lepadella (Lepadella) princisi B‘rziņÓ, 1943: PAL
Lepadella (Lepadella) psammophila Tzschaschel, 1979 (marine)
Lepadella (Lepadella) pseudosimilis Koch-Althaus, 1963: PAL
Lepadella (Lepadella) pterygoida (Dunlop, 1897): NEA, ORI, PAL
Metopidia pterygoidea Dunlop, 1897
Lepadella (Lepadella) ptilota B‘rziņÓ, 1960: AUS
Lepadella (Lepadella) pumilo Hauer, 1931: AFR, PAL
Lepadella (Lepadella) punctata Wulfert, 1939: ORI, PAL
Lepadella (Lepadella) pyriformis Myers, 1938: NEA
Lepadella (Lepadella) quadricarinata (Stenroos, 1898): AFR, AUS, NEA, NEO, ORI, PAL (incl. the

infrasubspecific variants octocarinata Wulfert, 1939, sexcarinata Klement, 1959, procera Klement, 1959)
Metopidia quadricarinata Stenroos, 1898
Lepadella (Lepadella) quadricurvata Francez & Pourriot, 1984: PAL
Lepadella (Lepadella) quinquecostata (Lucks, 1912)
Metopidia quinquecostata Lucks, 1912
Lepadella (L.) quinquecostata nevadensis Morales-Baquero, 1987: PAL
Lepadella (L.) quinquecostata quinquecostata (Lucks, 1912): AFR, AUS, NEA, NEO, ORI, PAC,
PAL
Syn.: Lepadella christineae Koste, 1990 (synonym considered an infrasubspecific variant) Lepadella
(Lepadella) rhodesiana Wulfert, 1965: AFR
Lepadella (Lepadella) rhomboides (Gosse, 1886)
Metopidia rhomboides Gosse, 1886
Lepadella (L.) rhomboides leentvaari B‘rziņÓ, 1979: NEO
Lepadella (L.) rhomboides rhomboides (Gosse, 1886): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Syn.: Lepadella rhomboides carinata Donner, 1943
Syn.: Lepadella rhomboides haueri Wulfert, 1956
Syn.: Lepadella rhomboides lata Wulfert, 1956
Lepadella (L.) rhomboides signiensis Dartnall & Hollowday, 1985: ANT
Lepadella (Lepadella) rhomboidula (Bryce, 1890): AFR, AUS, ORI, PAL
Metopidia rhomboidula Bryce, 1890
Lepadella (Lepadella) riedeli De Ridder, 1966: NEA, NEO
Lepadella (Lepadella) rottenburgi (Lucks, 1912): AUS, NEO, PAL
Metopidia rottenburgi Lucks, 1912
Lepadella (Lepadella) sali B‘rziņÓ, 1976: PAL
Lepadella (Lepadella) salisburii Ahlstrom, 1934: NEO
Lepadella (Lepadella) serrata Yamamoto, 1951: PAL
Lepadella (Lepadella) strepta B‘rziņÓ, 1984: NEA
Lepadella (Lepadella) tana Koste & Shiel, 1986: AUS
Lepadella (Lepadella) tenella Wulfert, 1942: PAL
Lepadella (Lepadella) triba Myers, 1934: AFR, AUS, NEA, NEO, ORI, PAC, PAL
Lepadella (Lepadella) tricostata Koste, 1990: NEO
Lepadella (Lepadella) triprojectus Sharma, 1978: ORI
Lepadella (Lepadella) triptera (Ehrenberg, 1832): AFR, ANT, AUS, NEA, NEO, ORI, PAC, PAL
(synonyms are considered infrasubspecific variants of L. triptera by Koste (1978))
Metopidia triptera Ehrenberg, 1832
Syn.: Lepadella alata Myers, 1934
Syn.: Lepadella alona Wulfert, 1956
Syn.: Lepadella clydona B‘rziņÓ, 1949
Syn.: Lepadella crestata Vasisht & Battish, 1971
Syn.: Lepadella deconincki De Ridder, 1966
Lepadella (Lepadella) tyleri Koste & Shiel, 1987: AUS
Lepadella (Lepadella) vandenbrandei Gillard, 1952: AFR, AUS, ORI
Syn.: Lepadella williamsi Koste & Shiel, 1980
Lepadella (Lepadella) vanoyei De Ridder, 1961: PAL
Lepadella (Lepadella) venefica Myers, 1934: NEA, NEO
Syn.: Lepadella exigua De Graaf, 1956: NEA (nom. nov. for L. minuta Myers non Weber & Montet (1915))

Syn.: Lepadella minuta Myers, 1942 (non Montet (1915))
Syn.: Lepadella myersiana Voigt, 1957 (nom. nov. for L. minuta Myers non Weber & Montet (1915))
Lepadella (Lepadella) visenda Myers, 1934: NEA, NEO
Lepadella (Lepadella) vitrea (Shephard, 1911): AUS
Metopidia vitrea Shephard, 1911
Lepadella (Lepadella) whitfordi Ahlstrom, 1938: NEA, NEO
Lepadella (Lepadella) wrighti Koste, 1972: NEO
Lepadella (Lepadella) xenica Myers, 1934: NEA, NEO
Lepadella (Lepadella) zigzag Segers, 2007
Syn.: Lepadella bractea Myers, 1934 (non Ehrenberg (1838))
subgenus Xenolepadella Hauer, 1926
Lepadella (Xenolepadella) astacicola Hauer, 1926: NEO, PAL

Lepadella (Xenolepadella) borealis Harring, 1916: AFR, NEA, ORI, PAL
Lepadella (Xenolepadella) branchicola Hauer, 1926: NEO, PAL
Lepadella (Xenolepadella) curvicaudata Turner, 1990: NEO
Lepadella (Xenolepadella) haueri Rodewald, 1935: PAL
Lepadella (Xenolepadella) lata Wiszniewski, 1939: AFR, ORI, PAL (incl. the infrasubspecific variant sin-
uata Wiszniewski, 1939)
Syn.: Lepadella raja Wiszniewski, 1939 (synonym considered an infrasubspecific variant)
Lepadella (Xenolepadella) monodactyla B‘rziņÓ, 1960
Lepadella monodactyla braziliensis Koste, 1972: NEO
Lepadella monodactyla monodactyla B‘rziņÓ, 1960: AFR, AUS, NEA, NEO, ORI
Syn.: Lepadella monostyla cicatrix B‘rziņÓ, 1984
Lepadella (Xenolepadella) parasitica Hauer, 1926: PAL
Lepadella (Xenolepadella) pseudoacuminata Sudzuki, 1998 (species inquirenda)
Limnias Schrank, 1803
Cephalosiphon Ehrenberg, 1853

Limnioides Tatem, 1868
Limnias ceratophylli Schrank, 1803: AFR, AUS, NEA, NEO, ORI, PAC, PAL
Syn.: Melicerta biloba Ehrenberg, 1832
Syn.: Rotifer confervicola Dutrochet, 1812
Syn.: Limnias socialis Leidy, 1874 (synonym occasionally considered an infrasubspecific variant of L. cerato-
phylli)
Syn.: Limnias sphagnicola Zacharias, 1886 (synonym occasionally considered an infrasubspecific variant of
L. ceratophylli)
Limnias cornuella Rousselet, 1889 (species inquirenda)
Limnias melicerta Weisse, 1848: AFR, AUS, NEA, NEO, ORI, PAC, PAL
Syn.: Limnias annulatus Bailey, 1855
Syn.: Limnias corniculata Ehrenberg, 1853
Syn.: Melicerta cubitti Cubitt, 1871
Syn.: Limnias doliolum Schoch, 1868

Syn.: Limnias granulosus Weber, 1888
Syn.: Cephalosiphon limnias Ehrenberg, 1853
Limnias myriophylli (Tatem, 1868): AFR, PAL
Limnioides myriophylli Tatem, 1868
Limnias nymphaea Stenroos, 1898: PAL (occasionally considered a subspecies of L. melicerta)
Limnias shiawasseensis Kellicott, 1888: NEA (occasionally considered a subspecies of L. melicerta)
Lindia Dujardin, 1841
subgenus Halolindia Remane, 1933
Lindia (Halolindia) elsae De Smet, 2006 (marine)

Lindia (Halolindia) gravitata (Lie-Pettersen, 1905) (marine)
Notommata gravitata Lie-Pettersen, 1905
Lindia (Halolindia) tecusa Harring & Myers, 1922 (marine)
Syn.: Lindia brodskayae Bogoslovski, 1950
subgenus Lindia Dujardin, 1841
Lindia (Lindia) annecta Harring & Myers, 1922: AUS, NEA, PAL
Lindia (Lindia) caerulea Myers, 1933: NEA
Lindia (Lindia) candida Harring & Myers, 1922: NEA, ORI, PAL
Lindia (Lindia) ecela Myers, 1933: AUS, NEA
Lindia (Lindia) euchromatica Edmondson, 1938: AUS, NEA, PAL
Syn.: Lindia europaea Koch-Althaus, 1962
Lindia (Lindia) fulva Harring & Myers, 1922: NEA, NEO
Lindia (Lindia) gracilis Myers, 1938: NEA
Lindia (Lindia) janickii Wiszniewski, 1934: AFR, ORI, PAL
Lindia (Lindia) pallida Harring & Myers, 1922: AUS, NEA, NEO, PAC, PAL
Lindia (Lindia) producta Harring & Myers, 1922: NEA
Lindia (Lindia) torulosa Dujardin, 1841: AFR, ANT, AUS, NEA, NEO, ORI, PAL
Syn.: Notommata roseola Perty, 1850
Syn.: Notommata tardigrada Leydig, 1854
Lindia (Lindia) truncata (Jennings, 1894): AUS, NEA, NEO, PAC, PAL
Notommata truncata Jennings, 1894
subgenus Neolindia Segers, 2002
Lindia (Neolindia) deridderae Koste, 1979: AUS
Lophocharis Ehrenberg, 1838
Oxysterna Iroso, 1910

Lophocharis ambidentata De Ridder, 1960: PAL
Lophocharis curvata B‘rziņÓ, 1982: AUS
Lophocharis hutchinsoni Edmondson, 1935: NEA
Lophocharis kutikovae Mirabdullaev, 1992: PAL

Lophocharis naias Wulfert, 1942: AUS, NEO, ORI, PAL
Syn.: Lophocharis gracilis Dvorakova, 1960
Lophocharis oxysternon (Gosse, 1851): AFR, ANT, AUS, NEA, NEO, ORI, PAL
Metopidia oxysternon Gosse, 1851
Lophocharis parva Rudescu, 1960: PAL
Lophocharis rubens Wulfert, 1939: PAL
Lophocharis salpina (Ehrenberg, 1834): AFR, AUS, NEA, NEO, ORI, PAL
Lepadella salpina Ehrenberg, 1834
Syn.: Lophocharis lepadelloides Rodewald, 1935
Syn.: Lophocharis parvidentata Hauer, 1937
Lophocharis turanica Mirabdullaev, 1992: PAL
Lophocharis tutiurensis Sudzuki, 1998 (species inquirenda)
Macrochaetus Perty, 1850
Macrochaetus altamirai (Arévalo, 1918): AFR, AUS, NEO, PAL

Polychaetus altamirai Arévalo, 1918
Syn.: Macrochaetus altamirai braziliensis Koste, 1972 (considered an infrasubspecific variant by Koste
(1978))
Syn.: Macrochaetus altamirai intermedius Rodewald, 1924 (considered an infrasubspecific variant by
Koste (1978))
Syn.: Macrochaetus australiensis B‘rziņÓ, 1982
Syn.: Macrochaetus esthonicus Riikoja, 1925
Macrochaetus americanus Segers & Sarma, 1993: NEO
Macrochaetus aspera B‘rziņÓ, 1982 (species inquirenda)
Macrochaetus aspinus Segers & Sarma, 1993: NEO
Macrochaetus collinsii (Gosse, 1867): AFR, AUS, NEA, NEO, ORI, PAL
Dinocharis collinsii Gosse, 1867
Macrochaetus collinsii collinsii (Gosse, 1867)
Syn.: Macrochaetus braziliensis Ahlstrom, 1938 (synonym considered an infrasubspecific variant)
Syn.: Polychaetus spinulosus Archer, 1868
Macrochaetus collinsii guangdongensis Wu, 1981 (species inquirenda)
Macrochaetus danneeli Koste & Shiel, 1983: AUS, ORI
Macrochaetus dispar B‘rziņÓ, 1982 (species inquirenda)
Macrochaetus hauerianus Wulfert, 1964: ORI
Macrochaetus kostei José de Paggi, Branco & Kozlowsky-Suzuki, 2000: NEO
Macrochaetus longipes Myers, 1934: AFR, NEA, NEO, ORI, PAL
Syn.: Macrochaetus hauerianus singapurensis Sudzuki, 1989
Macrochaetus longisetus Sudzuki, 1991 (species inquirenda)
Macrochaetus multispinosus Myers, 1934: AFR, NEO
Macrochaetus paggiensae Koste, 2000: NEO
Macrochaetus philopax Wulfert, 1961: PAL
Macrochaetus sericus (Thorpe, 1893): AFR, NEA, NEO, ORI, PAL
Dinocharis sericus Thorpe, 1893
Syn.: Macrochaetus sericus coniunctius Sudzuki, 1991
Syn.: Macrochaetus sericus vanmeeli Gillard, 1957

Macrochaetus subquadratus (Perty, 1850): AFR, AUS, NEA, NEO, ORI, PAL
Polychaetus subquadratus Perty, 1850
Syn.: Dinocharis rossicus Michailoff, 1900
Syn.: Macrochaetus subquadratus hofsteni Voigt, 1957
Macrotrachela Milne, 1886
Macrotrachela aculeata Milne, 1886: AFR, AUS, NEA, NEO, PAL

Macrotrachela allani (Murray, 1911): AFR
Callidina allani Murray, 1911
Macrotrachela ambigua Donner, 1965: PAL
Macrotrachela angusta (Bryce, 1894): AUS, PAL
Callidina angusta Bryce, 1894
Macrotrachela armillata (Murray, 1911): AUS
Callidina armillata Murray, 1911
Macrotrachela asperula (Murray, 1911): NEA, PAL
Callidina asperula Murray, 1911
Macrotrachela bilfingeri (Bryce, 1913): AFR, PAL
Callidina bilfingeri Bryce, 1913
Syn.: Mniobia quinquetuberculata Bartoš, 1938
Macrotrachela brachysoma Schulte, 1954: PAL
Macrotrachela brevilabris De Koning, 1947
Macrotrachela brevilabris aliena Donner, 1964: AUS, PAL
Macrotrachela brevilabris brevilabris De Koning, 1947: AUS, PAL
Macrotrachela bullata (Murray, 1906): AFR, ORI, PAL
Callidina bullata Murray, 1906
Macrotrachela canadensis (Murray, 1911): NEA
Callidina canadensis Murray, 1911
Macrotrachela cancrophila (Piovanelli, 1903): PAL
Callidina cancrophila Piovanelli, 1903
Macrotrachela compacta De Koning, 1947: PAL
Macrotrachela concinna (Bryce, 1912): ANT, AUS, NEO, PAL
Callidina concinna Bryce, 1912
Macrotrachela crucicornis (Murray, 1905): NEA, PAL
Callidina crucicornis Murray, 1905
Macrotrachela cuthberti Milne, 1916: AFR
Macrotrachela decora (Bryce, 1912): AUS, NEA, PAL
Callidina decora Bryce, 1912
Macrotrachela ehrenbergii (Janson, 1893): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Callidina ehrenbergi Janson, 1893
Macrotrachela extensa Haigh, 1965: AUS
Macrotrachela faveolata Milne, 1916: AFR
Macrotrachela festinans Donner, 1949: AUS, PAL
Macrotrachela formosa (Murray, 1906): AFR, AUS, NEO, ORI, PAL
Callidina formosa Murray, 1906
Macrotrachela fungicola Garner, 1937: PAL

Macrotrachela gracillima Donner, 1965: PAL
Macrotrachela gunningi (Murray, 1911): AFR, AUS
Callidina gunningi Murray, 1911
Macrotrachela habita (Bryce, 1894): AFR, ANT, AUS, NEA, NEO, ORI, PAL
Callidina habita Bryce, 1894
Syn.: Mniobia bulbifera Bartoš, 1939
Syn.: Mniobia gibbosa Bartoš, 1938
Macrotrachela herzigana Koste, 1996
Macrotrachela hewitti (Murray, 1911): AFR, PAL
Callidina hewitti Murray, 1911
Macrotrachela induta Donner, 1951: PAL
Macrotrachela inermis Donner, 1965: PAL
Macrotrachela insolita De Koning, 1947: ANT, AUS, NEA, NEO, PAL
Macrotrachela insulana Donner, 1962: NEO, PAL
Macrotrachela intermedia Schulte, 1954: PAL
Macrotrachela kallosoma (Schulte, 1954): ANT, AUS, NEO, PAL
Mniobia kallosoma Schulte, 1954
Macrotrachela labiata Donner, 1951: PAL
Macrotrachela lata De Koning, 1947: PAL
Macrotrachela latior Donner, 1951: PAL
Macrotrachela lepida (Murray, 1911): AUS
Callidina lepida Murray, 1911
Macrotrachela libera Donner, 1949
Macrotrachela libera alveolata Koniar, 1955: PAL
Macrotrachela libera libera Donner, 1949: PAL
Macrotrachela ligulata Haigh, 1965: AUS
Macrotrachela ligulifera Bartoš, 1947: PAL
Macrotrachela longistyla (Murray, 1911): AUS
Callidina longistyla Murray, 1911
Macrotrachela macmilliani Milne, 1916: AFR
Macrotrachela magna Schulte, 1954: PAL
Macrotrachela mariae Bartoš, 1938: PAL
Macrotrachela microcornis (Murray, 1911): AUS
Callidina microcornis Murray, 1911
Macrotrachela minuta Schulte, 1954
Macrotrachela minuta minuta Schulte, 1954: PAL
Macrotrachela minuta novella Donner, 1962: PAL
Macrotrachela multispinosa Thompson, 1892
Macrotrachela multispinosa brevispinosa (Murray, 1908): AFR, AUS, NEO, ORI, PAL
Callidina multispinosa brevispinosa Murray, 1908
Syn.: Macrotrachela tithanyca Varga, 1948
Macrotrachela multispinosa crassispinosa (Murray, 1907): AUS, NEO, PAL
Callidina multispinosa crassispinosa Murray, 1907
Macrotrachela multispinosa multispinosa Thompson, 1892: AFR, AUS, NEA, NEO, ORI, PAL
Macrotrachela muricata (Murray, 1905): PAL
Callidina muricata Murray, 1905

Macrotrachela murrayi B‘rziņÓ, 1950: PAL
Macrotrachela musculosa (Milne, 1886): AFR, ANT, AUS, NEA, ORI, PAL
Callidina musculosa Milne, 1886
Macrotrachela nana (Bryce, 1912)
Callidina nana Bryce, 1912
Macrotrachela nana laticalcar De Koning, 1947: PAL
Macrotrachela nana laticeps De Koning, 1947: PAL
Macrotrachela nana ligulata Schulte, 1954: PAL
Macrotrachela nana nana (Bryce, 1912): AFR, AUS, NEA, NEO, PAL
Macrotrachela natans (Murray, 1906): NEA, PAL
Callidina natans Murray, 1906
Macrotrachela nixa Donner, 1962: ANT, AUS, NEO, PAL
Macrotrachela oblita Donner, 1949: PAL
Macrotrachela obtusa Haigh, 1966: AUS
Macrotrachela ornata Donner, 1949: PAL
Macrotrachela pacifica (Murray, 1911): PAC
Callidina pacifica Murray, 1911
Macrotrachela papillosa Thompson, 1892: AFR, AUS, NEA, NEO, ORI, PAL
Macrotrachela petulans Milne, 1916: AFR, PAL
Macrotrachela pilousi Bartoš, 1948: PAL
Macrotrachela pinnigera (Murray, 1908): AFR, ORI
Callidina pinnigera Murray, 1908
Macrotrachela plicata (Bryce, 1892)
Callidina plicata Bryce, 1892
Macrotrachela plicata hirundinella (Murray, 1905): AFR, AUS, NEA, NEO, PAL
Callidina plicata hirundinella Murray, 1905
Macrotrachela plicata plicata (Bryce, 1892): AFR, AUS, NEA, PAL
Macrotrachela plicatula (Murray, 1911): AFR
Callidina plicatula Murray, 1911
Macrotrachela punctata (Murray, 1911): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Callidina punctata Murray, 1911
Macrotrachela quadricornifera Milne, 1886
Macrotrachela quadricornifera ligulata B‘rziņÓ, 1950: PAL
Macrotrachela quadricornifera loricata Donner, 1965: AFR, AUS, NEA, NEO, PAL
Macrotrachela quadricornifera quadricornifera Milne, 1886: AFR, ANT, NEO, ORI, PAL
Syn.: Macrotrachela serrulata Rodewald, 1935
Macrotrachela quadricornifera quadricorniferoides De Koning, 1929: AUS, PAL
Macrotrachela quadricornifera rigida Milne, 1916: AFR, AUS, NEO, PAL
Macrotrachela quadricornifera scutellata Schulte, 1954: AUS, PAL
Macrotrachela rostrata Donner, 1965: PAL
Macrotrachela samali Bartoš, 1948: PAL
Macrotrachela serrulata (Murray, 1911): AUS
Callidina serrulata Murray, 1911
Macrotrachela smithi Milne, 1916: AFR, AUS
Macrotrachela sonorensis Örstan, 1995 (species inquirenda)
Macrotrachela speciosa (Murray, 1907): AUS, NEA, NEO, PAL

Callidina speciosa Murray, 1907
Macrotrachela tenuis Donner, 1965: PAL
Macrotrachela timida Milne, 1916
Macrotrachela timida inquies Milne, 1916: AFR, NEO, PAL
Macrotrachela timida timida Milne, 1916: AFR, AUS, PAL
Macrotrachela tuberilabris De Koning, 1947: AUS, PAL
Macrotrachela vanoyei Schepens, 1954: PAL
Macrotrachela verecunda Milne, 1916: AFR
Macrotrachela vesicularis (Murray, 1906): AUS, PAL
Callidina vesicularis Murray, 1906
Macrotrachela zickendrahti (Richters, 1902)
Callidina zickendrahti Richters, 1902
Macrotrachela zickendrahti digitata Bartoš, 1943
Macrotrachela zickendrahti seda Donner, 1965
Macrotrachela zickendrahti zickendrahti (Richters, 1902)
Mikrocodides Bergendal, 1892
Mikrocodides chlaena (Gosse, 1886): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Stephanops chlaena Gosse, 1886
Syn.: Mikrocodides dubius Bergendal, 1892
Syn.: Microcodides mobilis Rodewald, 1940
Syn.: Rhinops orbiculodiscus Thorpe, 1891
Mikrocodides hertha Wulfert, 1961: PAL
Mikrocodides robustus (Glascott, 1893): NEA, NEO, PAL
Microcodon robustus Glascott, 1893
Microcodon Ehrenberg, 1830
Microcodon clavus Ehrenberg, 1830: AFR, ANT, AUS, NEA, NEO, ORI, PAL
Mniobia Bryce, 1910
Mniobia adhaerens Bartoš, 1944: PAL

Mniobia animosa Milne, 1916: AFR
Mniobia armata (Murray, 1905): PAL
Callidina armata Murray, 1905
Mniobia barbatula Donner, 1950: PAL
Mniobia bdelloidea Donner, 1951: PAL
Mniobia brachypoda Schulte, 1954: PAL
Mniobia branchicola (Nemec, 1895): PAL
Callidina branchicola Nemec, 1895
Mniobia bredensis De Koning, 1947: AUS, NEA, PAL
Mniobia burgeri Bartoš, 1951: ANT, NEA
Syn.: Mniobia ostensa Donner, 1980
Mniobia circinata (Murray, 1908): AUS, NEA, PAL
Callidina circinata Murray, 1908

Mniobia conarus B‘rziņÓ, 1982: AUS
Mniobia dentata Haigh, 1963: AUS
Mniobia discophora Bartoš, 1951: PAL
Mniobia donneri Bartoš, 1951: PAL
Mniobia edmondsoni Burger, 1948: NEA, PAL
Mniobia frankenbergeri Bartoš, 1944: PAL
Mniobia granulosa Bartoš, 1940: PAL
Mniobia incrassata (Murray, 1905): AUS, NEO, PAL
Callidina incrassata Murray, 1905
Mniobia incurata Rudescu, 1960: PAL
Mniobia iurensis Schulte, 1954: PAL
Mniobia lamellata Donner, 1950: AUS, PAL
Mniobia lenta Donner, 1951: PAL
Mniobia lineata Rahm, 1932: AUS, NEO, PAL
Mniobia lobata Haigh, 1967: AUS
Mniobia loxocorona De Koning, 1947: PAL
Mniobia magna (Plate, 1889): AUS, NEA, NEO, PAL
Callidina magna Plate, 1889
Mniobia mirabilis (Murray, 1911): AUS, NEO
Callidina mirabilis Murray, 1911
Mniobia modesta Donner, 1951: PAL
Mniobia montium Murray, 1911: NEA, PAL
Mniobia obtusicalcar De Koning, 1947: AUS, NEO, PAL
Mniobia obtusicornis Murray, 1911: AUS, NEA, PAL
Mniobia ocypetes B‘rziņÓ, 1982: AUS
Mniobia orta Donner, 1951: PAL
Mniobia placida Haigh, 1963: AUS
Mniobia procera Schulte, 1954: PAL
Mniobia punctata Haigh, 1963: AUS, PAL
Mniobia punctulata Bartoš, 1948: PAL
Mniobia recurvicornis Bartoš, 1950: PAL
Mniobia russeola (Zelinka, 1891): AFR, ANT, AUS, NEA, NEO, PAL
Callidina russeola Zelinka, 1891
Mniobia scabrosa Murray, 1911: AFR, AUS, NEA, PAL
Mniobia scarlatina (Ehrenberg, 1853): AFR, AUS, NEO, PAL
Callidina scarlatina Ehrenberg, 1853
Mniobia setifera Bartoš, 1944: PAL
Mniobia storkani Bartoš, 1948: PAL
Mniobia symbiotica (Zelinka, 1886): AUS, NEA, NEO, PAL
Callidina symbiotica Zelinka, 1886
Mniobia tarda Donner, 1949: AUS, PAL
Mniobia tentans Donner, 1949: NEO, PAL
Mniobia tetraodon (Ehrenberg, 1848): AFR, AUS, NEA, NEO, PAL
Callidina tetraodon Ehrenberg, 1848
Mniobia vargai Donner, 1962: PAL
Mniobia variabilis Donner, 1949: PAL

Monommata Bartsch, 1870
Monommata actices Myers, 1930: AUS, NEA, NEO, ORI, PAL

Syn.: Monommata arndti Remane, 1933
Monommata aequalis (Ehrenberg, 1830): AUS, NEA, NEO, ORI, PAL
Notommata aequalis Ehrenberg, 1832
Monommata aeschyna Myers, 1930: AUS, NEA, PAL
Monommata appendiculata Stenroos, 1898: NEA, PAL
Monommata astia Myers, 1930: AUS, NEA, NEO, PAC, PAL
Monommata caeca Myers, 1930: AUS, NEA, PAL
Monommata caudata Myers, 1930: NEA, PAL
Syn.: Monommata sphagnicola B‘rziņÓ, 1949
Monommata dentata Wulfert, 1940: AUS, NEA, PAC, PAL
Monommata diaphora Myers, 1930: AUS, NEA, PAL
Monommata dissimile B‘rziņÓ, 1949: PAL (species inquirenda)
Monommata enedra Myers, 1930: NEA, PAL
Monommata grandis Tessin, 1890: AFR, AUS, NEA, NEO, ORI, PAL
Syn.: Monommata robusta B‘rziņÓ, 1949
Monommata hyalina Myers, 1930: NEO, PAL
Monommata longiseta (Müller, 1786): AFR, AUS, NEA, NEO, PAL
Vorticella longiseta Müller, 1786
Monommata maculata Harring & Myers, 1930: AFR, AUS, NEA, NEO, ORI, PAC, PAL
Monommata phoxa Myers, 1930: AUS, NEA, NEO, ORI, PAL
Monommata pseudophoxa Wulfert, 1960: PAL
Monommata viridis Myers, 1937: AUS, NEA, PAL
Myersinella Wiszniewski, 1936
Myersina Wiszniewski, 1934 (non Herre (1934))
Myersinella belodon (Harring & Myers, 1928): NEA

Erignatha belodon Harring & Myers, 1928
Myersinella tetraglena (Wiszniewski, 1934): NEA, PAL
Myersina tetraglena Wiszniewski, 1934
Mytilina Bory de St. Vincent, 1826
Diplacidum Lauterborn, 1913

Diplax Gosse, 1851 (non Charpentier (1840))
Salpina Ehrenberg, 1830 (non Cantraine (1837))
Mytilina acanthophora Hauer, 1938: AFR, AUS, NEO, ORI, PAL

Mytilina bicarinata (Perty, 1850): NEA, ORI, PAL
Euchlanis bicarinata Perty, 1850
Syn.: Salpina pertyi Hood, 1895

Mytilina bisulcata (Lucks, 1912): AFR, AUS, NEO, ORI, PAL
Diplax bisulcata Lucks, 1912
Mytilina carpatica Varga, 1962: PAL
Mytilina compressa (Gosse, 1851): AFR, NEO, PAL
Diplax compressa Gosse, 1851
Mytilina crassipes (Lucks, 1912): AUS, NEA, PAL
Diplax crassipes Lucks, 1912
Mytilina lobata Pourriot, 1996: NEO
Mytilina macrocera (Jennings, 1894): AFR, NEA, NEO
Salpina macrocera Jennings, 1894
Mytilina michelangellii Reid & Turner, 1988: AFR, NEO
Syn.: Mytilina ventralis f. diversicantha Wulfert, 1965
Mytilina mucronata (Müller, 1773)
Brachionus mucronatus Müller, 1773
Mytilina mucronata longicauda Dartnall & Hollowday, 1985: ANT
Mytilina mucronata mucronata (Müller, 1773): AFR, AUS, NEA, NEO, ORI, PAL
Syn.: Salpina affinis Herrick, 1885
Syn.: Mytilina cypridina Bory de St. Vincent, 1926
Syn.: Mytilina cytherea Bory de St. Vincent, 1926
Syn.: Brachionus dentatus Müller, 1786
Syn.: Salpina marina Gosse, 1887
Syn.: Brachionus tetracerus Costa, 1838
Mytilina mucronata spinigera (Ehrenberg, 1830): PAL
Salpina spinigera Ehrenberg, 1830
Syn.: Salpina sulcata Gosse, 1886
Mytilina mucronata sumatrana B‘rziņÓ, 1982: ORI
Mytilina trigona (Gosse, 1851): AUS, NEA, NEO, ORI, PAL
Diplax trigona Gosse, 1851
Syn.: Mytilina trigona bispinosa Wang, 1961
Mytilina unguipes (Lucks, 1912): AFR, NEO, PAL
Diplax unguipes Lucks, 1912
Mytilina ventralis (Ehrenberg, 1830)
Salpina ventralis Ehrenberg, 1830
Mytilina ventralis brevispina (Ehrenberg, 1830): AFR, AUS, NEA, NEO, ORI, PAL
Salpina brevispina Ehrenberg, 1830
Syn.: Salpina redunca Ehrenberg, 1830
Mytilina ventralis longidactyla Wulfert, 1965: AFR, AUS, ORI
Mytilina ventralis ventralis (Ehrenberg, 1830): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Syn.: Salpina ceylonica Daday, 1896
Syn.: Salpina cortina Thorpe, 1891
Syn.: Salpina eustala Gosse, 1886
Syn.: Salpina macracantha Gosse, 1886
Syn.: Salpina similis Stokes, 1896
Mytilina ventralis wuhanensis Sudzuki & Huang, 1997 (species inquirenda)
Mytilina videns (Levander, 1894): PAL
Diplax videns Levander, 1894

Notholca Gosse, 1886
Pseudonotholca Mirakawa, 1928

Notholca acuminata (Ehrenberg, 1832): AFR, PAL
Anuraea acuminata Ehrenberg, 1832
Notholca angakkoq Sørensen, 1998: NEA
Notholca angulata (Daday, 1906): PAL
Anuraea angulata Daday, 1906
Notholca baikalensis Jaschnov, 1922: PAL (Baikal endemic)
Notholca beta Kutikova, 1986: PAL (Baikal endemic)
Notholca bipalium (Müller, 1786): NEA, PAL
Brachionus bipalium Müller, 1786
Notholca caudata Carlin, 1943: NEA, NEO, PAL
Notholca cinetura Skorikov, 1914: PAL
Syn.: Notholca grandis Voronkov, 1917
Notholca cornuta Carlin, 1943: NEA, PAL
Notholca foliacea (Ehrenberg, 1838): AFR, AUS, NEA, NEO, PAL
Anuraea foliacea Ehrenberg, 1838
Notholca gaigalasi Kutikova, 1970: PAL
Notholca guidoi Battistoni, 1992: NEO (Patagonia)
Notholca haueri Thomasson, 1963: NEO, PAL
Notholca hollowdayi Dartnall, 1995: ANT
Notholca ikaitophila Sørensen & Kristensen, 2000 (marine)
Notholca japonica (Marukawa, 1928): PAL
Pseudonotholca japonica Marukawa, 1928
Syn.: Notholca bifurca Hada, 1939
Syn.: Notholca quadraspinata Myers, 1936
Notholca jasnitzkii Tikhomirov, 1927: PAL (Baikal endemic)
Notholca kostei Chengalath, 1978: NEA
Notholca kozhovi Vassilijewa & Kutikova, 1969: PAL (Baikal endemic)
Notholca labis Gosse, 1887
Notholca labis labis Gosse, 1887: NEA, NEO, ORI, PAL
Syn.: Notholca labis frigida Jaschnov, 1922 ("frigida" forms are giant forms, presumedly associated with
extreme cold environments. They are recognized in several Notholca species)
Syn.: Notholca labis limnetica Levander, 1901 ("limnetica" forms have a strongly elongated caudal projec-
tion. Such forms are recognized in several Notholca species)
Notholca labis rectospina Kutikova, 1964: PAL
Notholca lamellifera Vassilijewa & Kutikova, 1969
Notholca lamellifera jashnovi Kutikova, 1986: PAL (Baikal endemic)
Notholca lamellifera lamellifera Vassilijewa & Kutikova, 1969: PAL (Baikal endemic)
Notholca latistyla Olofsson, 1918: PAL (Originally described as subspecies of N. foliacea)
Notholca laurentiae Stemberger, 1976: NEA
Notholca liepetterseni Godske Björklund, 1972: PAL
Notholca lyrata Tikhomirov, 1927: PAL (Baikal endemic)
Notholca marina Focke, 1961: PAL (Originally described as subspecies of N. acuminata)
Notholca michiganensis Stemberger, 1976: NEA

Notholca olchonensis Tikhomirov, 1927: PAL (Baikal endemic)
Notholca orbiculata Kutikova, 1986: PAL (Baikal endemic)
Notholca psammarina Buchholz & Rühmann, 1956: NEA, PAL
Notholca salina Focke, 1961
Notholca salina dumonti De Ridder & Verheye, 1980: PAL
Notholca salina salina Focke, 1961: ANT, NEA, PAL
Syn.: Notholca squamula evensi Gillard, 1948
Notholca squamula (Müller, 1786): AFR, ANT, AUS, NEA, NEO, ORI, PAL
Brachionus squamula Müller, 1786
Syn.: Notholca lapponica Ruttner Kolisko, 1966
Syn.: Notholca striata striata frigida Rylov, 1922
Notholca striata (Müller, 1786): NEA, NEO, PAL
Brachionus striata Müller, 1786
Syn.: Anuraea biremis Ehrenberg, 1832
Notholca tibetica Gong, 1983: PAL
Notholca triarthroides Skorikov, 1903: PAL
Notholca verae Kutikova, 1958: ANT
Notholca walterkostei José de Paggi, 1982
Notholca walterkostei reducta Dartnall & Hollowday, 1985: ANT
Notholca walterkostei walterkostei José de Paggi, 1982: ANT, NEO
Notommata Ehrenberg, 1830
Copeus Ehrenberg, 1838

Labidodon Ehrenberg, 1838
Notommata aethis Myers, 1933: NEA

Notommata allantois Wulfert, 1935: NEA, NEO, ORI, PAC, PAL
Notommata angusta Harring & Myers, 1922: NEA
Notommata apochaeta Myers, 1933: NEA
Notommata aurita (Müller, 1786): AUS, NEA, NEO, PAL
Vorticella aurita Müller, 1786
Syn.: Notommata epaxia Harring & Myers, 1924
Syn.: Cyclogena lupus Eyferth, 1877
Notommata avena Myers, 1933: NEA
Notommata bennetchi Myers, 1942: NEA
Notommata brachyota Ehrenberg, 1832: NEA, PAL
Syn.: Notommata brevipes Zawadowsky, 1926
Notommata cerberus (Gosse, 1886): AFR, AUS, NEA, NEO, PAL
Copeus cerberus Gosse, 1886
Syn.: Notommata parvida Myers, 1937
Notommata cherada Myers, 1938: NEA
Notommata codonella Harring & Myers, 1924
Notommata collaris Ehrenberg, 1832: AFR, AUS, NEA, NEO, PAL
Notommata contorta (Stokes, 1897): AUS, NEA, PAC, PAL
Diglena contorta Stokes, 1897

Notommata copeus Ehrenberg, 1834: AFR, AUS, NEA, NEO, ORI, PAL
Syn.: Notommata centrura Ehrenberg, 1834
Syn.: Copeus ehrenbergi Gosse, 1886
Syn.: Copeus labiatus Gosse, 1886
Syn.: Copeus notommata Ehrenberg, 1838
Notommata cyrtopus Gosse, 1886
Notommata cyrtopus cyrtopus Gosse, 1886: AFR, ANT, AUS, NEA, NEO, ORI, PAC, PAL
Syn.: Notommata carpatica Rodewald, 1935
Syn.: Notommata curvipes Zawadowsky, 1926
Syn.: Notommata distincta Bergendal, 1892
Syn.: Notommata telmata Harring & Myers, 1922
Notommata cyrtopus paracyrtopus de Beauchamp, 1932: PAL
Notommata diasema Myers, 1936: NEA, PAL
Notommata doneta Harring & Myers, 1924: AUS, NEA, PAL
Notommata endoxa Myers, 1933: NEA
Notommata falcinella Harring & Myers, 1922: NEA, PAL (incl. the infrasubspecific variant europaea
B‘rziņÓ, 1949)
Notommata fasciola Myers, 1933: NEA
Notommata galena codonella Harring & Myers, 1924: NEA, PAL
Notommata galena galena Harring & Myers, 1922: NEA
Notommata gisleni B‘rziņÓ, 1949: PAL
Notommata glyphura Wulfert, 1935: AFR, AUS, NEA, NEO, ORI, PAL
Notommata groenlandica Bergendal, 1892: PAL
Notommata haueri Wulfert, 1939: PAL
Notommata lenis Harring & Myers, 1922: NEA, PAL
Notommata longina Pourriot, 1995: AUS, PAL
Notommata mera Donner, 1970: PAL
Notommata omentata Wulfert, 1939: PAL
Notommata pachyura (Gosse, 1886): AFR, AUS, NEA, NEO, ORI, PAL
Copeus pachyurus Gosse, 1886
Syn.: Notommata brachiata Daday, 1908
Syn.: Copeus quinquelobatus Stokes, 1896
Syn.: Copeus triangulatus Kirkman, 1906 (synonym considered an infrasubspecific variant (f. triangulata) of
N. pachyura)
Notommata peridia Harring & Myers, 1922: NEA, NEO
Notommata placida Harring & Myers, 1922: NEA
Notommata prodota Myers, 1933: NEA
Notommata pseudocerberus de Beauchamp, 1908: AFR, AUS, NEA, NEO, ORI, PAC, PAL
Syn.: Notommata cerberus auct.
Notommata pygmaea Harring & Myers, 1922: NEA
Notommata rugosa Myers, 1933: NEA
Notommata saccigera Ehrenberg, 1830: AFR, AUS, NEA, NEO, ORI, PAL
Notommata silpha (Gosse, 1887): AFR, NEA, PAL
Diglena silpha Gosse, 1887
Syn.: Notommata forcipata Gosse, 1886 (non Ehrenberg (1838))
Notommata silphoides de Beauchamp, 1923: PAL

Notommata spinata Koste & Shiel, 1991: AUS
Notommata stitista Myers, 1937: NEA
Notommata thopica Harring & Myers, 1924: NEA, PAC
Notommata tripus Ehrenberg, 1838: AFR, AUS, NEA, NEO, ORI, PAC, PAL
Notommata venusta Harring & Myers, 1922: NEA
Notommata voigti Donner, 1949: PAL
Notommata weberi Voigt, 1957: PAL
Octoctrocha Thorpe, 1893
Octoctrocha speciosa Thorpe, 1893: AUS, NEA, NEO, PAL
Otostephanos Milne, 1916
Otostephanos annulatus Koniar, 1955: PAL

Otostephanos auriculatus (Murray, 1911)
Habrotrocha auriculata Murray, 1911
Otostephanos auriculatus auriculatus (Murray, 1911): AFR, AUS, NEA, PAC, PAL
Otostephanos auriculatus bilobatus Hauer, 1939: AUS, PAL
Otostephanos cuspidilabris De Koning, 1947: PAL
Otostephanos donneri Bartoš, 1959: AUS, PAL
Otostephanos jersabeki Koste, 1996: AFR
Otostephanos kostei Donner, 1972: PAL
Otostephanos macrantennus De Koning, 1947: PAL
Otostephanos monteti Milne, 1916: AFR, PAL
Otostephanos mundiformis De Koning, 1947: PAL
Otostephanos regalis Milne, 1916: AFR, PAL
Otostephanos torquatus (Bryce, 1913)
Habrotrocha torquata Bryce, 1913
Otostephanos torquatus amoenus Milne, 1916: AFR, PAL
Otostephanos torquatus torquatus (Bryce, 1913): AUS, NEA, PAL
Paracolurella Myers, 1936
Paracolurella aemula (Myers, 1934): NEA, NEO, ORI

Colurella aemula Myers, 1934
Paracolurella logima (Myers, 1934): NEA, NEO
Colurella logima Myers, 1934
Paradicranophorus Wiszniewski, 1929
Paradicranophorus aculeatus (Neiswestnowa-Shadina, 1935): PAL

Dicranophorus aculeatus Neiswestnowa-Shadina, 1935
Paradicranophorus hudsoni (Glascott, 1893): AUS, PAL
Diglena hudsoni Glascott, 1893

Syn.: Paradicranophorus limosus Wiszniewski, 1929
Paradicranophorus sinus De Smet, 2003: PAL
Paradicranophorus sordidus Donner, 1968: ANT, PAL
Syn.: Encentrum brevifulcrum Dartnall, 1997
Paradicranophorus verae Bogoslovsky, 1958: PAL
Paradicranophorus wesenberglundi Sørensen, 2001: PAL (brackish water)
Paraseison Plate, 1887
Paraseison annulatus (Claus, 1876) (marine, epizoic on Nebalia)

Seison annulatus Claus, 1876
Syn.: Paraseison asplanchnus Plate, 1887
Parencentrum Wiszniewski, 1953
Parencentrum lutetiae (Harring & Myers, 1928): ORI, PAL

Encentrum lutetiae Harring & Myers, 1928
Parencentrum plicatum (Eyferth, 1878): NEA, PAL
Theora plicata Eyferth, 1878
Syn.: Parencentropsis mittii Sudzuki, 1960
Pedipartia Myers, 1937
Pedipartia gracilis (Myers, 1936): NEA, PAC

Dicranella gracilis Myers, 1936
Philodina Ehrenberg, 1830
Philodina acuticornis Murray, 1902

Philodina acuticornis acuticornis Murray, 1902: AFR, AUS, NEA, NEO, PAL
Philodina acuticornis minor Pax & Wulfert, 1942: PAL
Philodina acuticornis odiosa Milne, 1916: AFR, NEA, NEO, PAL
Philodina alata Murray, 1910: ANT
Philodina americana Murray, 1913: NEO
Philodina amethystina Bartoš, 1951: PAL
Philodina antarctica Murray, 1910: ANT
Philodina arndti Wulfert, 1961: PAL
Philodina australis Murray, 1911: AUS, NEA
Philodina brevipes Murray, 1902: AFR, AUS, NEA, NEO, ORI, PAC, PAL
Philodina calceata Donner, 1970: PAL
Philodina childi Milne, 1916: AFR
Philodina citrina Ehrenberg, 1832: AFR, AUS, NEA, NEO, ORI, PAL
Philodina convergens Murray, 1908: PAL
Philodina cristata Donner, 1949: AFR, PAL
Philodina dobrogensis Rudescu, 1960: PAL
Philodina duplicalcar (De Koning, 1947): PAL

Didymodactylos duplicalcar De Koning, 1947
Philodina erythrophthalma Ehrenberg, 1830: AFR, AUS, NEA, NEO, PAL
Philodina eurystephana Schulte, 1954: PAL
Philodina flaviceps Bryce, 1906: AFR, AUS, NEA, NEO, ORI, PAL
Philodina foissneri Koste, 1996: AFR
Philodina grandis Milne, 1916: AFR
Philodina gregaria Murray, 1910: ANT
Philodina indica Murray, 1906: NEA, PAL
Philodina inopinata Milne, 1916: AFR, PAL
Philodina jeanneli de Beauchamp, 1940 : ANT
Philodina lepta Wulfert, 1950: AUS, NEA, NEO, PAL
Philodina megalotrocha Ehrenberg, 1832: AFR, AUS, NEA, NEO, ORI, PAL
Syn.: Philodina calcarata Schmarda, 1894
Philodina morigera Donner, 1949: AUS, NEA, PAL
Philodina nemoralis Bryce, 1903: AFR, AUS, NEA, PAL
Philodina nitida Milne, 1916
Philodina nitida decens Milne, 1916: AFR
Philodina nitida nitida Milne, 1916: AFR, AUS
Philodina parvicalcar De Koning, 1947: PAL
Philodina patula Milne, 1916: AFR
Philodina plena (Bryce, 1894): AFR, ANT, AUS, NEA, NEO, PAL
Callidina plena Bryce, 1894
Philodina praelonga Milne, 1916: AFR
Philodina proterva Milne, 1916: AFR, AUS, NEA, PAL
Philodina quadrata De Koning, 1947: PAL
Philodina rapida Milne, 1916: AFR, NEO, PAL
Philodina roseola Ehrenberg, 1832: AFR, AUS, NEA, NEO, PAL
Syn.: Philodina cinnabarina Zacharias, 1886
Philodina rugosa Bryce, 1903
Philodina rugosa callosa Bryce, 1903: PAL
Philodina rugosa coriacea Bryce, 1903: PAL
Philodina rugosa rugosa Bryce, 1903: AFR, AUS, NEA, NEO, PAL
Philodina scabra Milne, 1916
Philodina squamosa Murray, 1906: AUS, ORI
Philodina striata Rodewald, 1937: PAL
Philodina tenuicalcar De Koning, 1947: PAL
Philodina tranquilla Wulfert, 1942: AUS, PAL
Philodina tridentata Rodewald, 1935: PAL
Philodina vorax (Janson, 1893): AFR, AUS, NEA, NEO, ORI, PAL
Callidina vorax Janson, 1893
Philodinavus Harring, 1913
Philodinavus aussiensis Ricci, Shiel, Fontaneto & Melone, 2003

Philodinavus paradoxus (Murray, 1905): AUS, NEA, PAL
Microdina paradoxa Murray, 1905

Plationus Segers, Murugan & Dumont, 1993
Plationus felicitas (Wulfert, 1956): PAL

Platyias felicitas Wulfert, 1956
Plationus patulus (Müller, 1786)
Brachionus patulus Müller, 1786
Plationus patulus macracanthus (Daday, 1905): AFR, AUS, NEO, ORI
Noteus patulus macracanthus Daday, 1905
Plationus patulus patulus (Müller, 1786): AFR, AUS, NEA, NEO, ORI, PAL
Plationus polyacanthus (Ehrenberg, 1834): PAL
Brachionus polyacanthus Ehrenberg, 1834
Syn.: Brachionus polyacanthoides Bērziņš, 1943
Platyias Harring, 1913
Noteus Ehrenberg, 1832
Platyias latiscapularis Koste, 1974: NEO

Platyias leloupi Gillard, 1967: AFR, AUS, NEA, NEO, ORI
Syn.: Platyias longispinosus Arora, 1966
Platyias quadricornis (Ehrenberg, 1832)
Noteus quadricornis Ehrenberg, 1832
Platyias quadricornis andhraensis Dhanapathi, 1974: ORI
Platyias quadricornis quadricornis (Ehrenberg, 1832): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Syn.: Brachionus intermedius Herrick, 1900
Syn.: Noteus quadricornis brevispinus Daday, 1905 (synonym occasionally considered an infrasubspecific
variant (f. brevispinus) of P. quadricornis)
Pleurata Nogrady & Pourriot, 1995
Pleurata chalicodes (Myers, 1933): NEA

Pleurotrocha chalicodes Myers, 1933
Pleurata thura (Myers, 1933): NEA
Pleurotrocha thura Myers, 1933
Pleurata tithasa (Harring & Myers, 1924): NEA
Notommata tithasa Harring & Myers, 1922
Pleurata trypeta (Harring & Myers, 1922): NEA, PAL
Notommata trypeta Harring & Myers, 1922
Pleurata tyleri (Koste, Shiel & Tan, 1988): AUS
Notommata tyleri Koste, Shiel & Tan, 1988
Pleurata uroglenae (de Beauchamp, 1948) : PAL
Proales uroglenae de Beauchamp, 1948
Pleurata vernalis (Wulfert, 1935): PAL
Pleurotrocha vernalis Wulfert, 1935

Pleuretra Bryce, 1910
Pleuretra africana Murray, 1911: AFR, NEO

Pleuretra alpium (Ehrenberg, 1853): AFR, AUS, NEA, NEO, PAL
Callidina alpium Ehrenberg, 1853
Pleuretra bovicornis Bartoš, 1963: ORI
Pleuretra brycei (Weber, 1898): AFR, AUS, NEA, NEO, PAL
Callidina brycei Weber, 1898
Syn.: Callidina cataracta Lord, 1898
Pleuretra costata (Bartoš, 1938): PAL
Macrotrachela costata Bartoš, 1938
Pleuretra humerosa (Murray, 1905): AFR, AUS, NEA, NEO, PAC, PAL
Philodina humerosa Murray, 1905
Pleuretra hystrix Bartoš, 1950: PAL
Pleuretra intermedia (Bartoš, 1938): PAL
Mniobia intermedia Bartoš, 1938
Pleuretra lineata Donner, 1962: AUS, PAL
Pleuretra proxima Bartoš, 1963: ORI
Pleuretra reticulata Milne, 1916: AFR, PAL
Pleuretra similis Bartoš, 1963: ORI
Pleuretra sulcata Bartoš, 1950: PAL
Pleuretra triangularis Murray, 1913: NEO
Pleurotrocha Ehrenberg, 1830
Pleurotrocha altila Myers, 1940: NEA

Pleurotrocha atlantica Myers, 1936: NEA, PAL (backish water and marine)
Pleurotrocha aurea (Zawadovsky, 1916): PAL
Proales aureus Zawadovsky, 1916
Pleurotrocha channa Myers, 1933: NEA
Pleurotrocha elegans Zawadovsky, 1926: PAL
Pleurotrocha larvarum Vlastov, 1956: PAL
Pleurotrocha petromyzon (Ehrenberg, 1830): AFR, AUS, NEA, NEO, ORI, PAL
Notommata petromyzon Ehrenberg, 1830
Syn.: Notops laurentinus Jennings, 1894
Pleurotrocha robusta (Glascott, 1893): NEA, PAL
Microcodon robustus Glascott, 1893
Syn.: Microcodides abbreviatus Stenroos, 1898
Pleurotrochopsis Bērziņš, 1973
Pleurotrochopsis multispinosa (Fadeew, 1925): NEA, PAL

Pleurotrocha multispinosa Fadeew, 1925
Syn.: Pleurotrochopsis anebodica Bērziņš, 1973

Ploesoma Herrick, 1885
Bipalpus Wierzejski & Zacharias, 1893

Dictyoderma Lauterborn, 1893
Gastroschiza Bergendal, 1892
Gomphogaster Vorce, 1888
Ploesoma africanum Wulfert, 1965: AFR, NEO
Ploesoma hudsoni (Imhof, 1891): NEA, NEO, PAL
Gastropus hudsoni Imhof, 1891
Syn.: Gastroschiza flexilis Jägerskiold, 1892
Syn.: Dictyoderma hypopus Lauterborn, 1893
Syn.: Ploesoma sibericum Daday, 1901
Syn.: Bipalpus vesiculosus Wierzejski & Zacharias, 1893
Ploesoma lenticulare Herrick, 1885: AFR, AUS, NEA, NEO, ORI, PAL
Ploesoma lenticulare fukusimensis Sudzuki, 1992 (species inquirenda)
Ploesoma lenticulare lenticulare Herrick, 1885
Syn.: Gomphogaster areolatus Vorce, 1888
Syn.: Gastropus ehrenbergii Imhof, 1891
Syn.: Ploesoma ekmani Brehm, 1910
Syn.: Gastroschiza foveolata Jägerskiold, 1892
Syn.: Bipalpus lynceus Wierzejski & Zacharias, 1893
Ploesoma multispinatum Wulfert, 1965 (species inquirenda)
Ploesoma murrayi Wulfert, 1961: PAL
Ploesoma peipsiense Maëmets & Kutikova, 1979: PAL
Ploesoma triacanthum (Bergendal, 1892): NEO, PAL
Gastroschiza triacantha Bergendal, 1892
Ploesoma truncatum (Levander, 1894): AFR, AUS, NEA, NEO, PAL
Gastroschiza truncata Levander, 1894
Polyarthra Ehrenberg, 1834
Anarthra Hood, 1895

Polyarthra aptera Hood, 1893 (first generation hatching from resting eggs (several species))
Polyarthra proloba Wulfert, 1941 (specimens with mastax protruding in a ventral prolapse (several species))
Polyarthra pseudoproloba Albertova, 1960 (specimens with mastax protruding in a ventral prolapse (several
species))
Polyarthra bicerca Wulfert, 1956 (species inquirenda)
Polyarthra dissimulans Nipkow, 1952 (species inquirenda)
Polyarthra dolichoptera Idelson, 1925: AFR, AUS, NEA, ORI, PAC, PAL
Syn.: Polyarthra platyptera auct.
Polyarthra dolichoptera f. brachyptera Bartoš, 1947 (species inquirenda)
Polyarthra euryptera Wierzejski, 1891: NEA, ORI, PAL
Syn.: Polyarthra latiremis Imhof, 1891
Polyarthra indica Segers & Babu, 1999: AFR, ORI
Polyarthra leleki Koste & Tobias, 1989: AFR
Polyarthra longiremis Carlin, 1943: AUS, NEA, NEO, ORI, PAL

Polyarthra luminosa Kutikova, 1962: NEA, NEO, ORI, PAL
Polyarthra major Burckhardt, 1900: AUS, NEA, NEO, ORI, PAL
Polyarthra minor Voigt, 1904: AUS, NEA, ORI, PAL
Polyarthra platyptera Ehrenberg, 1838 (species inquirenda)
Polyarthra remata Skorikov, 1896: AUS, NEA, NEO, ORI, PAL
Polyarthra trigla Ehrenberg, 1834 (species inquirenda)
Polyarthra vulgaris Carlin, 1943: AFR, AUS, NEA, NEO, ORI, PAC, PAL
Syn.: Polyarthra trigla auct.
Pompholyx Gosse, 1951
Pompholyx complanata Gosse, 1851: AFR, AUS, NEA, NEO, ORI, PAL
Pompholyx sulcata Hudson, 1885: AFR, NEA, NEO, ORI, PAL
Pompholyx triloba Pejler, 1957: NEA, PAL
Pourriotia De Smet, 2003
Pourriotia carcharodonta De Smet, 2003: ANT
Proales Gosse, 1886
Proales adenodis Myers, 1933: NEA

Proales alba Wulfert, 1939: NEA, PAL
Proales baradlana Varga, 1958: PAL
Proales bemata Myers, 1933: NEA
Proales christinae De Smet, 1994 (marine)
Proales cognita Myers, 1940: NEA
Proales commutata Althaus, 1957 (marine)
Proales cryptopus Wulfert, 1935: NEO, PAL
Proales daphnicola Thompson, 1892: AFR, AUS, NEA, ORI, PAL
Syn.: Proales nova Wlastow, 1953
Syn.: Proales pejleri De Smet, Van Rompu & Beyens, 1992
Proales decipiens (Ehrenberg, 1832): AFR, AUS, NEA, NEO, ORI, PAL
Notommata decipiens Ehrenberg, 1832
Syn.: Proales brevipes Harring & Myers, 1924
Syn.: Notommata vermicularis Dujardin, 1841
Proales doliaris (Rousselet, 1895): AUS, NEA, PAL
Microcodides doliaris Rousselet, 1895
Proales fallaciosa Wulfert, 1937: AFR, AUS, NEA, NEO, ORI, PAC, PAL
Syn.: Proales tyrphosa Bērziņš, 1949
Proales germanica Tzschaschel, 1979 (marine)
Proales gigantea (Glascott, 1893): AUS, NEA, PAL
Notommata gigantea Glascott, 1893
Syn.: Proales ovicola Giard, 1908
Proales gladia Myers, 1933: NEA, PAL

Proales globulifera (Hauer, 1921): PAL
Furcularia globulifera Hauer, 1921
Syn.: Proales quadrangularis Glascott, 1893
Proales gonothyraeae Remane, 1929 (marine)
Proales granulosa Myers, 1933: NEA
Proales halophila Remane, 1929: PAL (marine and brackish water)
Proales indirae Wulfert, 1966: ORI
Proales kostei Nogrady & Smol, 1989: NEA
Proales lenta Wlastow, 1956: PAL
Proales litoralis De Smet, 1996 (marine)
Proales macrura Myers, 1933: NEA, PAL
Proales micropus (Gosse, 1886): AUS, NEA, NEO, PAL
Furcularia micropus Gosse, 1886
Proales minima (Montet, 1915): NEA, PAC, PAL
Pleurotrocha minima Montet, 1915
Syn.: Proales psammophila Neiswestnowa-Shadina, 1935
Proales oculata Tzschaschel, 1979 (marine)
Proales ornata Myers, 1933: NEA, PAL
Proales paguri Thane-Fenchel, 1966 (marine)
Proales palimmeka Myers, 1940: NEA
Proales parasita (Ehrenberg, 1838): AUS, NEA, NEO, PAL
Notommata parasita Ehrenberg, 1838
Proales phaeopis Myers, 1933: NEA
Proales provida Wulfert, 1938: PAL
Proales pugio Nogrady, 1983: AFR, NEA
Proales reinhardti (Ehrenberg, 1834) (marine)
Furcularia reinhardti Ehrenberg, 1834
Proales segnis Myers, 1938: NEA, PAL
Proales sigmoidea (Skorikov, 1896): NEA, PAL
Pleurotrocha sigmoidea Skorikov, 1896
Syn.: Pleurotrocha macropoda Zavadowski, 1926
Proales similis de Beauchamp, 1907: AUS, NEA, ORI, PAC, PAL (halophile)
Proales simplex Wang, 1961: PAL
Proales sordida Gosse, 1886: AFR, AUS, NEA, ORI, PAL
Proales syltensis Tzschaschel, 1979 (marine)
Proales theodora (Gosse, 1887): AUS, NEA, PAL
Notommata theodora Gosse, 1887
Syn.: Proales longipes Remane, 1929
Proales werneckii (Ehrenberg, 1834): AFR, AUS, NEA, PAL
Notommata werneckii Ehrenberg, 1834
Proales wesenbergi Wulfert, 1960: PAL
Proalides de Beauchamp, 1907
Liliferotrocha Sudzuki, 1959

Proalides digitus Donner, 1978: PAL

Proalides subtilis Rodewald, 1940: AFR, AUS, NEO, ORI, PAL
Proalides tentaculatus de Beauchamp, 1907: AFR, NEA, NEO, PAC, PAL
Proalinopsis Weber, 1918
Proalinopsis caudatus (Collins, 1873): AFR, AUS, NEA, PAL

Notommata caudata Collins, 1873
Syn.: Proalinopsis montanus Godeanu, 1962 (synonym is species inquirenda)
Syn.: Proalinopsis trisegmentus Sudzuki, 1960 (synonym is species inquirenda)
Proalinopsis gracilis Myers, 1933: NEA, PAL
Proalinopsis lobatus Rodewald, 1935: PAL (species inquirenda)
Proalinopsis phacus Myers, 1933: NEA
Proalinopsis selene Myers, 1933: NEA
Proalinopsis squamipes Hauer, 1935: NEA, PAL
Proalinopsis staurus Harring & Myers, 1924: AFR, AUS, NEA, NEO, PAL
Pseudoembata Wycliffe & Michael, 1968 (Genus inquirendus)
Pseudoembata acutipoda Wycliffe & Michael, 1968 (species inquirenda)
Pseudoeuchlanis Dhanapathi, 1978
Pseudoeuchlanis longipedes Dhanapathi, 1978: ORI
Pseudoharringia Fadeew, 1925
Pseudoharringia romanica Rodewald, 1937: PAL (species inquirenda)

Pseudoharringia similis Fadeew, 1925: PAL
Pseudoploesoma Myers, 1938
Pseudoploesoma formosum (Myers, 1934): NEO

Ploesoma formosum Myers, 1934
Pseudoploesoma greeni Koste, 1978 (species inquirenda)
Ptygura Ehrenberg, 1832
Oecistes Ehrenberg, 1834

Pseudoecistes Stenroos, 1898
Ptygura agassizi Edmondson, 1948: NEA (occasionally considered an infrasubspecific variant of P. melicerta)
Ptygura barbata Edmondson, 1939: AUS, NEA, NEO, ORI
Ptygura beauchampi Edmondson, 1940: NEA, NEO, PAL (Oecistes cf. crystallinus after de Beauchamp,
1932)
Syn.: Ptygura lacunosa Summerfield-Wright, 1957 (synonym considered an infrasubspecific variant of P.
beauchampi)

Ptygura brachiata (Hudson, 1886): AFR, AUS, NEA, NEO, ORI, PAL (incl. the infrasubspecific variant
bispicata Edmondson, 1944)
Oecistes brachiatus Hudson, 1886
Ptygura brevis (Rousselet, 1893): AUS, PAL
Oecistes brevis Rousselet, 1893
Ptygura cristata (Murray, 1913): AUS, NEA, NEO
Oecistes cristatus Murray, 1913
Ptygura crystallina (Ehrenberg, 1834): AFR, ANT, AUS, NEA, NEO, PAL
Oecistes crystallinus Ehrenberg, 1834
Ptygura elsteri Koste, 1972: NEO, ORI
Ptygura furcillata (Kellicott, 1889): AFR, AUS, NEA, NEO, ORI, PAL (incl. the infrasubspecific variant
variabilis Koste 1975)
Cephalosiphon furcillatus Kellicott, 1889
Syn.: Ptygura furcata Hauer, 1938
Ptygura intermedia (Davis, 1867): AUS, NEO, PAL
Oecistes intermedius Davis, 1867
Ptygura kostei José de Paggi, 1996: AFR, NEO, ORI
Syn.: Ptygura elsteri f. thailandensis Koste, 1975
Ptygura libera Myers, 1934: AFR, NEA, NEO, PAL
Ptygura linguata Edmondson, 1939: NEA, NEO
Ptygura longicornis (Davis, 1867): AUS, NEA, NEO, PAL (incl. the infrasubspecific variant bispicata
Edmondson, 1944)
Oecistes longicornis Davis, 1867
Ptygura melicerta Ehrenberg, 1832
Ptygura melicerta ctenoida Koste & Tobias, 1990
Ptygura melicerta melicerta Ehrenberg, 1832: AFR, ANT, AUS, NEA, NEO, ORI, PAL
Syn.: Oecistes ptygura Hudson & Gosse, 1886
Syn.: Oecistes serpentinus Gosse, 1886
Syn.: Oecistes socialis Weber, 1888 (synonym occasionally considered an infrasubspecific variant of P. meli-
certa)
Ptygura mucicola (Kellicott, 1888): AFR, AUS, NEA, NEO, ORI, PAL (occasionally considered an infra-
subspecific variant of P. melicerta)
Oecistes mucicola Kellicott, 1888
Ptygura pedunculata Edmondson, 1939: NEA, NEO, PAL
Ptygura pilula (Cubitt, 1872): AFR, AUS, NEA, NEO, PAL
Melicerta pilula Cubitt, 1872
Syn.: Oecistes longipes Wills, 1878 (synonym occasionally considered an infrasubspecific variant of P. pilula)
Syn.: Oecistes umbella Hudson, 1879
Ptygura rotifer (Stenroos, 1898): NEA, PAL (incl. the infrasubspecific variant conica Bērziņš, 1949)
Pseudoecistes rotifer Stenroos, 1898
Ptygura seminatans Edmondson, 1939: NEA
Ptygura spongicola Bērziņš, 1950 (Possibly a synonym of P. melicerta according to Koste (1978); species
inquirenda)
Ptygura stephanion (Anderson, 1889) (species inquirenda)
Oecistes stephanion Anderson, 1889
Ptygura stygis (Gosse, 1886): NEA, PAL

Oecistes stygis Gosse, 1886
Syn.: Ptygura cephaloceros Summerfield-Wright, 1957 (incl. the infrasubspecific variant grande Koste,
1970)
Ptygura tacita Edmondson, 1940: AUS, NEA, NEO, ORI
Ptygura tihanyensis Varga, 1937 (species inquirenda)
Ptygura tridorsicornis Summerfield-Wright, 1957: PAL
Ptygura velata (Gosse, 1851): AFR, AUS, NEA, NEO, PAC, PAL
Megalotrocha velata Gosse, 1851
Syn.: Melicerta ptygura Gosse, 1862
Ptygura wilsonii (Anderson & Shephard, 1892): AFR, AUS, NEO, PAL
Oecistes wilsonii Anderson & Shephard, 1892
Syn.: Oecistes pectinifera Murray, 1913
Resticula Harring & Myers, 1924
Resticula anceps Harring & Myers, 1924: NEA

Resticula gelida (Harring & Myers, 1922): ANT, AUS, NEA, NEO, PAL
Eosphora gelida Harring & Myers, 1922
Resticula lestes Wulfert, 1935: PAL
Resticula melandocus (Gosse, 1887): AFR, AUS, NEA, NEO, ORI, PAL
Furcularia melandocus Gosse, 1887
Resticula nyssa Harring & Myers, 1924: AUS, NEA, PAL
Resticula plicata Wulfert, 1935: NEA, PAL
Resticula vermiculus Wulfert, 1935: PAL
Rhinoglena Ehrenberg, 1853
Rhinops Hudson, 1869

Rhinoglena fertoeensis (Varga, 1928): PAL (doubtful Antarctic record)
Rhinops fertoeensis Varga, 1928
Rhinoglena frontalis Ehrenberg, 1853: ANT, AUS, NEA, ORI, PAL
Syn.: Rhinops vitrea Hudson, 1869
Rhinoglena tokioensis Sudzuki, 1975: PAL
Rotaria Scopoli, 1777
Rotifer Cuvier, 1798

Rotaria citrina (Ehrenberg, 1838): AFR, AUS, NEA, PAL
Rotifer citrinus Ehrenberg, 1838
Rotaria curtipes (Murray, 1911): AUS
Rotifer curtipes Murray, 1911
Rotaria elongata (Weber, 1888): AFR, AUS, NEA, NEO, PAL
Rotifer elongatus Weber, 1888
Rotaria exoculis De Koning, 1947: AUS, NEA, PAL
Rotaria haptica (Gosse, 1886): AUS, PAL
Rotifer hapticus Gosse, 1886

Rotaria laticeps Wulfert, 1942: AUS, PAL
Rotaria macroceros (Gosse, 1851): AFR, AUS, NEA, NEO, ORI, PAL
Rotifer macroceros Gosse, 1851
Syn.: Rotifer motacilla Bartsch, 1870
Rotaria macrura (Ehrenberg, 1832)1: AFR, AUS, NEA, NEO, PAL (not Vorticella macroura Hermann,
1783)
Rotifer macrurus Ehrenberg, 1832
Rotaria magnacalcarata (Parsons, 1892): PAL
Callidina magnacalcarata Parsons, 1892
Rotaria mento (Anderson, 1889): NEA, ORI, PAL
Rotifer mento Anderson, 1889
Rotaria montana (Murray, 1911): AUS, ORI
Rotifer montanus Murray, 1911
Rotaria murrayi Bartoš, 1951: PAL
Rotaria neptunia (Ehrenberg, 1830): AFR, AUS, NEA, NEO, ORI, PAL
Actinurus neptunius Ehrenberg, 1830
Rotaria neptunoida Harring, 1913: AFR, AUS, NEA, ORI, PAL
Syn.: Rotifer neptunius Milne, 1886 (non Ehrenberg (1832))
Rotaria ovata (Anderson, 1889): ORI
Actinurus ovatus Anderson, 1889
Rotaria quadrangularis (Heinis, 1913): NEA
Rotifer quadrangularis Heinis, 1913
Rotaria quadrioculata (Murray, 1902): AUS, PAL
Rotifer quadrioculatus Murray, 1902
Rotaria rotatoria (Pallas, 1766)
Brachionus rotatorius Pallas, 1766
Rotaria rotatoria granularis (Zacharias, 1885): PAL
Rotifer rotatoria granularis Zacharias, 1885
Rotaria rotatoria rotatoria (Pallas, 1766): AFR, AUS, NEA, NEO, ORI, PAL
Syn.: Esechielina bakeri Bory de St. Vincent, 1826
Syn.: Rotifer brachyurus Ehrenberg, 1831
Syn.: Esechielina leuwenhoekii Bory de St. Vincent, 1826
Syn.: Rotaria monteti Bērziņš, 1955
Syn.: Esechielina muelleri Bory de St. Vincent, 1826
Syn.: Rotifer redivivus Cuvier, 1798
Syn.: Rotaria saprobica Bērziņš, 1955
Syn.: Rotifer vulgaris Schrank, 1801
Rotaria rotatoria spongioderma Pax & Wulfert, 1941: PAL
Rotaria socialis (Kellicott, 1888): NEA, NEO, PAL
Callidina socialis Kellicott, 1888
Rotaria sordida (Western, 1893)
1. There is considerable confusion in literature regarding the identity and synonymy of Rotaria macrura (Ehrenberg)
and Vorticella macroura Hermann, 1783 (Rotifer macrourus: Schrank, 1803). The exact taxonomic identity of the latter
can’t be inferred from the original description; however, it is clear that the two are not conspecific (Harring, 1913). I
therefore follow Harring (1913) rather than Donner (1965), who lists Vorticella marcoura as synonym of Rotaria
macrura (Schrank).

Callidina sordida Western, 1893
Rotaria sordida fimbriata (Murray, 1906): AFR, AUS, ORI, PAC, PAL
Rotifer longirostris fimbriatus Murray, 1906
Rotaria sordida sordida (Western, 1893): AFR, AUS, NEA, NEO, ORI, PAL
Syn.: Callidina longirostris Janson, 1893
Rotaria spicata (Murray, 1902): NEA, PAL
Rotifer spicatus Murray, 1902
Rotaria tardigrada (Ehrenberg, 1830): AFR, AUS, NEA, NEO, ORI, PAL
Rotifer tardigradus Ehrenberg, 1830
Syn.: Rotifer maximus Bartsch, 1870
Syn.: Rotifer tardus Ehrenberg, 1838
Rotaria tridens (Montet, 1915): AUS, NEA, NEO, PAL
Rotifer tridens Montet, 1915
Syn.: Rotaria tridentata Varga, 1954
Rotaria trisecata (Weber, 1888): NEA, NEO, PAL
Rotifer trisecatus Weber, 1888
Rousseletia Harring, 1913
Rousseletia corniculata Harring, 1913: NEA
Scaridium Ehrenberg, 1830
Scaridium bostjani Daems & Dumont, 1974: AFR, AUS, NEA, NEO, ORI, PAL
Scaridium elegans Segers & De Meester, 1994: AFR, AUS, NEO, ORI (record from China by Zhuge
(1997) is S. longicaudum)
Scaridium elongatum Segers, 1996: NEO
Scaridium grande Segers, 1995: AFR, ORI
Scaridium longicaudum (Müller, 1786): AFR, ANT, AUS, NEA, NEO, ORI, PAC, PAL
Trichoda longicauda Müller, 1786
Scaridium montanum Segers, 1995: PAL
Scaridium neglectum Segers, 1997: NEA
Scepanotrocha Bryce, 1910
Scepanotrocha corniculata Bryce, 1910: AUS, NEO, PAL

Scepanotrocha delicata Donner, 1949: AUS, PAL
Scepanotrocha galeata Milne, 1916: AFR, PAL
Scepanotrocha haueri Donner, 1962
Scepanotrocha haueri amota Donner, 1962: PAL
Scepanotrocha haueri haueri Donner, 1962: PAL
Scepanotrocha impexa Donner, 1962: PAL
Scepanotrocha parva Schulte, 1954: PAL
Scepanotrocha rubra Bryce, 1910: AUS, NEA, NEO, PAL
Scepanotrocha semitecta Donner, 1951: NEO, PAL
Scepanotrocha setifera Haigh, 1963: AUS
Scepanotrocha simplex De Koning, 1947: AUS, NEO, PAL

Seison Grube, 1861
Seison africanus Sørensen, Segers & Funch, 2005

Seison nebaliae Grube, 1861 (marine, epizoic on Nebalia)
Syn.: Seison grubei Claus, 1876
Syn.: Saccobdella nebaliae Van Beneden & Hesse non Grube, 1864
Sinantherina Bory de St. Vincent, 1826
Sinantherina ariprepres Edmondson, 1939: AUS, NEA, NEO

Sinantherina procera (Thorpe, 1893): NEO, ORI
Megalotrocha procera Thorpe, 1893
Sinantherina semibullata (Thorpe, 1893): AFR, AUS, NEA, NEO, ORI, PAL
Megalotrocha semibullata Thorpe, 1893
Syn.: Megalotrocha binotata Daday, 1897
Sinantherina socialis (Linnaeus, 1758): AFR, AUS, NEA, NEO, ORI, PAL
Hydra socialis Linnaeus, 1758
Syn.: Megalotrocha alboflavicans Ehrenberg, 1838
Sinantherina spinosa (Thorpe, 1893): AFR, AUS, NEA, NEO, ORI, PAL
Megalotrocha spinosa Thorpe, 1893
Sinantherina triglandularis Arora, 1963 (species inquirenda)
Sphyrias Harring, 1913
Sphyrias lofuana (Rousselet, 1910): AFR, PAC

Notops lofuana Rousselet, 1910
Squatinella Bory de St. Vincent, 1826
Stephanops Ehrenberg, 1830

Listrion Schmarda, 1846
Squatinella bifurca (Bolton, 1884): PAL (includes the infrasubspecific variant gigantea Pourriot, 1971)
Stephanops bifurcus Bolton, 1884
Squatinella geleii Varga, 1933: PAL
Squatinella lamellaris (Müller, 1786): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Brachionus lamellaris Müller, 1786
Syn.: Squatinella aurita Wulfert, 1950 (incl. the infrasubspecific variant myersi Voigt, 1957)
Syn.: Stephanops dichthapsis Anderson, 1889
Syn.: Squatinella donneri Wulfert, 1956
Syn.: Stephanops emarginatus Bilfinger, 1894
Syn.: Stephanops groenlandicus Bergendal, 1892
Syn.: Stephanops intermedius Burn, 1889
Syn.: Squatinella minor Wulfert, 1961
Syn.: Stephanops muticus Ehrenberg, 1832 (synonym considered an infrasubspecific variant (f. mutica) of S.
lamellaris)
Syn.: Stephanops tridentatus Fresenius, 1858 (synonym considered an infrasubspecific variant (f. tridentata)

of S. lamellaris)
Squatinella leydigii (Zacharias, 1886): AUS, NEA, NEO, PAL (incl. the infrasubspecific variant longiseta
Pourriot, 1971)
Stephanops leydigii Zacharias, 1886
Syn.: Squatinella longispinata auct.
Syn.: Stephanops variegatus Levander, 1894
Squatinella longispinata (Tatem, 1867): AUS, NEA, ORI, PAL
Stephanops longispinatus Tatem, 1867
Syn.: Stephanops armatus Hudson, 1885
Syn.: Stephanops tripus Hudson, 1889
Syn.: Stephanops uniseta Collins, 1872
Syn.: Stephanops unisetatus Hudson & Gosse, 1886
Squatinella lunata Segers, 1993: AFR, ORI
Squatinella microdactyla (Murray, 1906): NEA, PAC, PAL
Stephanops microdactylus Murray, 1906
Squatinella pseudorostrum Markevich, 1990 (species inquirenda)
Squatinella retrospina Myers, 1938: NEA
Squatinella rostrum (Schmarda, 1846): AUS, NEA, PAL
Listrion rostrum Schmarda, 1846
Syn.: Squatinella lamellata auct.
Stephanoceros Ehrenberg, 1832
Stephanoceros fimbriatus (Goldfusz, 1820): AUS, NEA, NEO, ORI, PAL

Coronella fimbriata Goldfusz, 1820
Streptognatha Harring & Myers, 1928
Streptognatha lepta Harring & Myers, 1928: NEA
Synchaeta Ehrenberg, 1832
Synchaeta arcifera Xu, 1998: NEO, PAL

Synchaeta atlantica Zelinka, 1907 (marine)
Synchaeta bacillifera Smirnov, 1933 (marine)
Synchaeta baltica Ehrenberg, 1834 (marine)
Syn.: Synchaeta apus Plate, 1889
Synchaeta bicornis Smith, 1904: NEA, PAL (marine and brackish water)
Synchaeta calva Ruttner-Kolisko, 1970 (species inquirenda)
Synchaeta cecilia Rousselet, 1902: PAL (marine and brackish water)
Synchaeta cecilia f. fusipes Buchholz, 1954 (marine)
Synchaeta curvata Lie-Pettersen, 1905 (species inquirenda)
Synchaeta cylindrica Althaus, 1957: PAL (Inland saline waters)
Synchaeta elsteri Hauer, 1963 (species inquirenda)
Synchaeta fennica Rousselet, 1909 (marine)

Synchaeta glacialis Smirnov, 1932 (marine)
Synchaeta grandis Zacharias, 1893: NEA, PAL
Synchaeta grimpei Remane, 1929: NEA, PAL (marine and brackish water)
Syn.: Synchaeta procera Galliford, 1946
Synchaeta gyrina Hood, 1887: PAL (marine and brackish water)
Synchaeta hutchingsi Brownell, 1988: AFR (marine)
Synchaeta hyperborea Smirnov, 1932 (marine)
Synchaeta johanseni Harring, 1921: NEA
Synchaeta jollyae Shiel & Koste, 1993: AUS
Synchaeta kitina Rousselet, 1902
Synchaeta kitina kitina Rousselet, 1902: NEA, NEO, PAL
Synchaeta kitina monodactyla Koch-Althaus, 1976 (species inquirenda)
Synchaeta lakowitziana Lucks, 1930
Synchaeta lakowitziana arctica De Smet, 1988: PAL
Synchaeta lakowitziana lakowitziana Lucks, 1930: AUS, NEA, PAL
Synchaeta littoralis Rousselet, 1902 (species inquirenda)
Synchaeta longipes Gosse, 1887: AFR, AUS, NEA, NEO, ORI, PAL
Syn.: Synchaeta wesenberglundi Pejler, 1957
Synchaeta monopus Plate, 1889 (marine)
Synchaeta neapolitana Rousselet, 1902 (marine)
Synchaeta oblonga Ehrenberg, 1832: AFR, AUS, NEA, NEO, ORI, PAC, PAL
Synchaeta pachypoda Jashnov, 1922: PAL (Baikal endemic)
Synchaeta pachypoida Kutikova & Vassiljeva, 1982: PAL (Baikal endemic)
Synchaeta pectinata Ehrenberg, 1832: AFR, AUS, NEA, NEO, ORI, PAL
Syn.: Synchaeta mordax Gosse, 1851
Synchaeta prominula Kutikova & Vassiljeva, 1982: PAL (Baikal endemic)
Synchaeta rousseleti Zelinka, 1927 (marine, Antarctic)
Synchaeta rufina Kutikova & Vassiljeva, 1982: PAL (Baikal endemic)
Synchaeta squamadigitata De Smet, 2006 (marine)
Synchaeta stylata Wierzejski, 1893: AFR, AUS, NEA, NEO, ORI, PAL
Synchaeta tamara Smirnov, 1932 (marine)
Synchaeta tavina Althaus, 1957: AUS, NEA, PAL (marine and brackish water)
Synchaeta tremula (Müller, 1786): AFR, AUS, NEA, NEO, ORI, PAL
Vorticella tremula Müller, 1786
Syn.: Synchaeta neglecta Zacharias, 1901
Syn.: Synchaeta truncata von Hofsten, 1909
Synchaeta tremuloida Pourriot, 1965: PAL
Synchaeta triophthalma Lauterborn, 1894: AUS, NEA, PAL (marine and brackish water)
Synchaeta verrucosa Nipkow, 1961: NEA, PAL
Syn.: Synchaeta asymmetrica Koch-Althaus, 1963
Synchaeta vorax Rousselet, 1902: NEA, PAL (marine and brackish water)
Taphrocampa Gosse, 1851
Taphrocampa annulosa Gosse, 1851: AFR, AUS, NEA, NEO, ORI, PAL
Taphrocampa clavigera Stokes, 1896: NEA, NEO, PAC, PAL

Taphrocampa lemurensis Bērziņš, 1982: AFR
Taphrocampa selenura Gosse, 1887: AFR, AUS, NEA, NEO, ORI, PAL
Testudinella Bory de St. Vincent, 1826
Pterodina Ehrenberg, 1938
Testudinella ahlstromi Hauer, 1956: AUS, NEA, NEO

Testudinella amphora Hauer, 1938: AUS, NEO, ORI
Testudinella andranomenensis Sudzuki, 1992 (species inquirenda)
Testudinella angulata Myers, 1934: NEA
Testudinella aspis Carlin, 1939: PAL
Testudinella berzinsi Gillard, 1952: AFR
Testudinella brevicaudata Yamamoto, 1951: AFR, ORI, PAL
Syn.: Pompholyx acutangula Wu, 1981
Testudinella brycei Hauer, 1938: AFR, NEO, ORI, PAL
Testudinella caeca (Parsons, 1892): AFR, NEA, NEO, PAL (incl. the infrasubspecific variants lermaensis
Ahlstrom, 1932 and haueri Wiszniewski, 1954)
Pterodina caeca Parsons, 1892
Testudinella carlini Bartoš, 1951: PAL
Testudinella clypeata (Müller, 1786): NEA, PAL
Brachionus clypeatus Müller, 1786
Syn.: Pterodina crassa Levander, 1894
Syn.: Testudinella dentata Myers, 1934
Syn.: Testudinella pseudoclypeata Bērziņš, 1943 (synonym occasionally considered an infrasubspecific
variant of T. clypeata)
Testudinella dendradena de Beauchamp, 1955: AFR, NEO, ORI (distribution unreliable because of taxo-
nomic confusion (see Rougier & Pourriot, 2006))
Testudinella discoidea Ahlstrom, 1938: NEA
Testudinella elliptica (Ehrenberg, 1834): NEA, NEO, ORI, PAL
Pterodina elliptica Ehrenberg, 1834
Testudinella emarginula (Stenroos, 1898): AFR, AUS, NEA, NEO, ORI, PAL
Pterodina emarginula Stenroos, 1898
Testudinella epicopta Myers, 1934: NEA
Testudinella gillardi De Ridder, 1966: PAL
Testudinella greeni Koste, 1981: AFR, AUS, NEO, ORI
Syn.: Testudinella tridentata f. curvata Wulfert, 1965
Syn.: Testudinella tridentata obtusa Wu, 1981
Testudinella haueriensis Gillard, 1967: NEO (distribution unreliable because of taxonomic confusion (see
Rougier & Pourriot, 2006))
Testudinella husseyi Shiel & Koste, 1985: AUS
Testudinella incisa (Ternetz, 1892): AFR, AUS, NEA, NEO, ORI, PAL
Pterodina incisa Ternetz, 1892
Syn.: Testudinella naumanni Carlin, 1939 (synonym occasionally considered an infrasubspecific variant of T.
incisa)
Syn.: Testudinella vidzemensis Bērziņš, 1943

Syn.: Testudinella vidzemensis alofsi Gillard, 1952 (species inquirenda)
Testudinella kostei De Ridder, 1983: AFR
Testudinella mucronata (Gosse, 1886)
Pterodina mucronata Gosse, 1886
Testudinella mucronata mucronata (Gosse, 1886): AFR, AUS, NEA, NEO, ORI, PAL (distribution
unreliable because of taxonomic confusion (see Rougier & Pourriot, 2006))
Testudinella mucronata tasmaniensis Koste & Shiel, 1986: AUS
Testudinella munda Bērziņš, 1982: AUS
Testudinella neboisi Bērziņš, 1982: AUS
Testudinella obscura Althaus, 1957 (marine psammon)
Testudinella ohlei Koste, 1972: NEO (incl. the infrasubspecific variant pustulata Koste, 1978)
Testudinella ovata Myers, 1934: NEA
Testudinella panonica Zivkovic, 1987 (species inquirenda)
Testudinella parva (Ternetz, 1892): AFR, AUS, NEA, NEO, ORI, PAL
Pterodina parva Ternetz, 1892
Syn.: Testudinella bidentata Ternetz, 1892 (synonym considered an infrasubspecific variant of T. parva)
Syn.: Pterodina calcaris Langer, 1909 (identical to T. parva var. bidentata)
Syn.: Pterodina emarginata Wierzejski, 1893 (identical to T. parva var. bidentata)
Syn.: Testudinella insinuata Hauer, 1938 (T. insuata auct)
Syn.: Testudinella semiparva Hauer, 1938 (synonym occasionally considered an infrasubspecific variant of T.
parva)
Testudinella patina (Hermann, 1783): AFR, AUS, NEA, NEO, ORI, PAC, PAL (incl. the infrasubspe-
cific variant lindbergi Bērziņš)
Brachionus patina Hermann, 1783
Syn.: Pterodina intermedia Anderson, 1889
Syn.: Testudinella pseudoelliptica Bartoš, 1951
Syn.: Testudinella sculpturata Bartoš, 1951 (synonym occasionally considered an infrasubspecific variant of
T. patina)
Syn.: Pterodina trilobata Anderson & Shephard, 1892 (synonym occasionally considered an infrasubspe-
cific variant of T. patina)
Syn.: Testudinella trilobata haterumensis Sudzuki, 1992 (species inquirenda)
Syn.: Testudinella trilobata triangularis Sudzuki, 1992 (species inquirenda)
Syn.: Pterodina valvata Hudson, 1871
Testudinella reflexa (Gosse, 1887): AFR, AUS, NEA, NEO, PAL
Pterodina reflexa Gosse, 1887
Testudinella robertsonae Koste, 1990: NEO
Testudinella sphagnicola Rudescu, 1960 (species inquirenda)
Testudinella stappersi Evens, 1947 (species inquirenda)
Testudinella striata (Murray, 1913) (species inquirenda)
Pterodina striata Murray, 1913
Testudinella subdiscoidea De Ridder, 1983: AFR
Testudinella triangularis Myers, 1934: NEA
Testudinella tridentata Smirnov, 1931: AFR, AUS, NEA, NEO, ORI, PAL
Syn.: Testudinella dicella Myers, 1934
Syn.: Testudinella paratridentata Van, 1961
Syn.: Testudinella tridentata africana Gillard, 1952

Syn.: Testudinella tridentata amazonica Thomasson, 1971
Syn.: Testudinella tridentata insulana Bērziņš, 1982
Testudinella tridentata Smirnov, 1931: AFR, AUS, NEA, NEO, ORI, PAL
Testudinella truncata (Gosse, 1886): AFR, NEA, PAL (incl. the infrasubspecific variants ecornis
Wiszniewski, 1953 and usmaensis Bērziņš, 1943)
Pterodina truncata Gosse, 1886
Syn.: Pterodina stenroosi Rundström, 1909
Testudinella unicornuta Koste & Shiel, 1987: AUS
Testudinella vanoyei Gillard, 1947 (species inquirenda)
Testudinella walkeri Koste & Shiel, 1980: AUS
Testudinella wuhanensis Sudzuki, 1992 (species inquirenda)
Tetrasiphon Ehrenberg, 1840
Tetrasiphon hydrocora Ehrenberg, 1840: AFR, AUS, NEA, NEO, ORI, PAL
Trichocerca Lamarck, 1801
Acanthodactylus Tessin, 1890 (non Wiegmann (1834))

Coelopus Ehrenberg, 1886
Diurella Bory de St. Vincent, 1824
Mastigocerca Ehrenberg, 1830
Monocerca Bory de St. Vincent, 1826
Rattulus Lamarck, 1816
Vaginaria Schrank, 1802
Trichocerca abilioi Segers & Sarma, 1993: AFR, NEO, ORI

Trichocerca agnatha Wulfert, 1939: AFR, AUS, PAL
Trichocerca antilopaea (Petr, 1891) (species inquirenda)
Rattulus antilopaeus Petr, 1891
Trichocerca artmanni (Zelinka, 1927) (species inquirenda)
Rattulus artmanni Zelinka, 1927
Trichocerca bambekei (Mola, 1913) (species inquirenda)
Coelopus bambekei Mola, 1913
Trichocerca barsica (Varga & Dudich, 1939) (species inquirenda)
Diurella barsica Varga & Dudich, 1939
Trichocerca bicristata (Gosse, 1887): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Mastigocerca bicristata Gosse, 1887
Trichocerca bicurvicornis (Mola, 1913) (species inquirenda)
Mastigocerca bicurvicornis Mola, 1913
Trichocerca bicuspes (Pell, 1890): NEA
Mastigocerca bicuspes Pell, 1890
Trichocerca bidens (Lucks, 1912): AFR, ANT, AUS, NEA, NEO, ORI, PAC, PAL
Diurella bidens Lucks, 1912
Syn.: Diurella bidens astriata Rodewald, 1935
Syn.: Trichocerca cavia auct.

Trichocerca bilunaris Blainville, 1830 (species inquirenda)
Trichocerca brachydactyla (Glascott, 1893) (species inquirenda)
Mastigocerca brachydactyla Glascott, 1893
Trichocerca brachyura (Gosse, 1851): AFR, ANT, AUS, NEA, NEO, ORI, PAL
Monocerca brachyura Gosse, 1851
Syn.: Rattulus palpitatus Stokes, 1896
Trichocerca braziliensis (Murray, 1913): AFR, AUS, NEO, ORI
Rattulus braziliensis Murray, 1913
Trichocerca brevidactyla (Daday, 1890) (species inquirenda)
Diurella brevidactyla Daday, 1890
Trichocerca brevistyla (Lucks, 1912) (species inquirenda)
Diurella brevistyla Lucks, 1912
Trichocerca calypta (Gosse, 1886) (species inquirenda)
Rattulus calyptus Gosse, 1886
Trichocerca capucina (Wierzejski & Zacharias, 1893): AFR, AUS, NEA, NEO, ORI, PAL
Mastigocerca capucina Wierzejski & Zacharias, 1893
Syn.: Mastigocerca hudsoni Lauterborn, 1893
Trichocerca caspica (Tschugunoff, 1921) (probable synonym of T. marina?)
Rattulus caspicus Tschugunoff, 1921
Syn.: Rattulus caspicus longicaudis Tschugunoff, 1921
Trichocerca catellus (Müller, 1773) (species inquirenda)
Cercaria catellus Müller, 1773
Trichocerca cavia (Gosse, 1886): AFR, AUS, NEA, NEO, ORI, PAL
Coelopus cavia Gosse, 1886
Trichocerca chattoni (de Beauchamp, 1907): AFR, AUS, NEO, ORI, PAL (proposed as subspecies of T.
cylindrica)
Rattulus chattoni de Beauchamp, 1907
Trichocerca cimolia (Gosse, 1886) (species inquirenda)
Rattulus cimolius Gosse, 1886
Trichocerca collaris (Rousselet, 1896): AFR, AUS, NEA, NEO, ORI, PAL
Rattulus collaris Rousselet, 1896
Trichocerca compressa Edmondson, 1938 (species inquirenda)
Trichocerca cryptodus (Hauer, 1937) (species inquirenda)
Diurella cryptodus Hauer, 1937
Trichocerca cuspidata (Stenroos, 1898) (species inquirenda)
Mastigocerca cuspidata Stenroos, 1898
Trichocerca cylindrica (Imhof, 1891): AFR, AUS, NEA, NEO, ORI, PAL
Mastigocerca cylindrica Imhof, 1891
Syn.: Mastigocerca elegans Meissner, 1902
Syn.: Mastigocerca hamata Zacharias, 1897
Syn.: Mastigocerca hamata bologonensis Minkiewicz, 1900
Syn.: Mastigocerca setifera Lauterborn, 1893
Trichocerca dixonnuttalli (Jennings, 1903): AFR, AUS, NEA, NEO, ORI, PAL
Diurella dixonnuttalli Jennings, 1903
Trichocerca edmondsoni (Myers, 1936): NEA, NEO
Diurella edmondsoni Myers, 1936

Syn.: Trichocerca rossae Edmondson, 1936
Trichocerca elongata (Gosse, 1886): AFR, AUS, NEA, NEO, ORI, PAL
Mastigocerca elongata Gosse, 1886
Syn.: Mastigocerca grandis Stenroos, 1898
Syn.: Trichocerca tchadiensis Pourriot, 1968
Trichocerca euodonta (Hauer, 1937) (species inquirenda)
Diurella euodonta Hauer, 1937
Trichocerca flagellata Hauer, 1937: AFR, AUS, NEO, ORI, PAL
Trichocerca flava (Voronkov, 1907) (species inquirenda)
Rattulus flavus Voronkov, 1907
Trichocerca gillardi Koste, 1978 (species inquirenda)
Trichocerca gracilis (Tessin, 1890): AFR, AUS, NEA, NEO, ORI, PAL (species inquirenda)
Acanthodactylus gracilis Tessin, 1890
Trichocerca harveyensis Myers, 1942: NEA, PAL
Trichocerca helminthodes (Gosse, 1886) (species inquirenda)
Rattulus helminthodes Gosse, 1886
Trichocerca heterodactyla (Tschugunoff, 1921) (species inquirenda)
Diurella heterodactyla Tschugunoff, 1921
Trichocerca hollaerti De Smet, 1990: AFR, NEO, ORI, PAC
Trichocerca iernis (Gosse, 1887): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Mastigocerca iernis Gosse, 1887
Trichocerca inermis (Linder, 1904) (species inquirenda)
Coelopus inermis Linder, 1904
Trichocerca insignis (Herrick, 1885): AFR, AUS, NEA, NEO, PAC, PAL
Diurella insignis Herrick, 1885
Trichocerca insolens (Myers, 1936): NEA
Diurella insolens Myers, 1936
Trichocerca insulana (Hauer, 1937): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Diurella insulana Hauer, 1937
Syn.: Trichocerca montana Hauer, 1956
Trichocerca intermedia (Stenroos, 1898): AUS, NEA, NEO, PAC, PAL
Coelopus intermedius Stenroos, 1898
Syn.: Trichocerca intermedia arenosa Wiszniewski, 1953
Trichocerca jobloti Bory de St. Vincent, 1827 (species inquirenda)
Trichocerca kostei Segers, 1993: AFR, NEO, ORI
Trichocerca lata (Jennings, 1894): NEA
Mastigocerca lata Jennings, 1894
Trichocerca longiseta (Schrank, 1802): AFR, AUS, NEA, NEO, ORI, PAL
Vaginaria longiseta Schrank, 1802
Syn.: Mastigocerca auchinlecki Stewart, 1908
Syn.: Monocerca bicornis Ehrenberg, 1832
Syn.: Monocerca cornuta Eyferth, 1878
Syn.: Trichocerca falx Edmondson, 1936 (synonym is a newly hatched specimen)
Trichocerca longistyla (Olofsson, 1918): PAL
Diurella longistyla Olofsson, 1918
Syn.: Trichocerca parvula Carlin, 1939

Trichocerca lophoessa (Gosse, 1886): AFR, AUS, NEA, NEO, ORI, PAL
Mastigocerca lophoessa Gosse, 1886
Syn.: Mastigocerca rectocaudis Hilgendorf, 1899
Trichocerca macera (Gosse, 1886): AFR, AUS, NEA, PAL
Mastigocerca macera Gosse, 1886
Syn.: Mastigocerca fusiformis Levander, 1894
Trichocerca maior (Hauer, 1935): NEA, PAL (proposed as infrasubspecific variant of T. porcellus)
Diurella maior Hauer, 1935
Syn.: Diurella porcellus carinata Myers, 1936
Trichocerca marina (Daday, 1890) (Marine species)
Diurella marina Daday, 1890
Syn.: Mastigocerca curvata Levander, 1894 (synonym occasionally listed as subspecies of T. marina)
Syn.: Mastigocerca dubia Lauterborn, 1894
Syn.: Rattulus henseni Zelinka, 1907
Syn.: Rattulus liepetterseni de Beauchamp, 1907 (proposed as ssp.)
Trichocerca microstyla (Daday, 1901) (species inquirenda)
Mastigocerca microstyla Daday, 1901
Trichocerca minuta (Gosse, 1886) (species inquirenda)
Coelopus minutus Gosse, 1886
Trichocerca mollis Edmondson, 1936: NEA, NEO
Trichocerca mucosa (Stokes, 1896): NEA (non T. bicristata mucosa auct.)
Mastigocerca mucosa Stokes, 1896
Trichocerca mucripes Ahlstrom, 1938 (species inquirenda)
Trichocerca multicrinis (Kellicott, 1897): NEA, NEO (down to Panama)
Mastigocerca multicrinis Kellicott, 1897
Trichocerca murchiearum Edmondson, 1936 (species inquirenda)
Trichocerca mus Hauer, 1938: AUS, NEA, NEO, ORI, PAL
Trichocerca musculus (Hauer, 1937): AFR, AUS, NEA, PAL
Diurella musculus Hauer, 1937
Syn.: Diurella parva Rodewald, 1935 (non Manfredi (1927))
Trichocerca myersi (Hauer, 1931): AFR, AUS, NEA, NEO, ORI, PAL
Diurella myersi Hauer, 1931
Syn.: Diurella tortuosa Myers, 1936
Trichocerca nitida Harring, 1914 (species inquirenda)
Trichocerca obtusidens (Olofsson, 1918): NEA, NEO, PAC, PAL
Diurella obtusidens Olofsson, 1918
Syn.: Trichocerca arctica Voigt, 1957
Syn.: Trichocerca minuta Olofsson, 1918 (non Gosse (1886))
Syn.: Trichocerca relicta Donner, 1950
Trichocerca orca (Murray, 1913): AUS, ORI
Rattulus orca Murray, 1913
Trichocerca ornata Myers, 1934: NEA
Trichocerca pediculus Remane, 1949 (Marine species)
Trichocerca plaka Myers, 1938: NEA
Trichocerca platessa Myers, 1934: NEA
Trichocerca porcellus (Gosse, 1851): AFR, AUS, NEA, NEO, ORI, PAL

Monocerca porcellus Gosse, 1851
Syn.: Acanthodactylus tigris Tessin, 1890 (non Müller (1786))
Trichocerca pusilla (Jennings, 1903): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Rattulus pusillus Jennings, 1903
Trichocerca pygocera (Wiszniewski, 1932): PAL
Diurella pygocera Wiszniewski, 1932
Trichocerca rattus (Müller, 1776): AFR, ANT, AUS, NEA, NEO, ORI, PAC, PAL
Trichoda rattus Müller, 1776
Syn.: Rattulus carinatus Lamarck, 1816 (synonym considered an infrasubspecific variant of T. rattus, as T.
rattus f. carinata)
Syn.: Trichoda criceta Schrank, 1803
Syn.: Trichocerca cristata Harring, 1913 (nom. nov. for Mastigocerca carinata Ehrenberg, 1830 non Lama-
rck, 1816)
Syn.: Brachionus cylindricus Schrank, 1776
Syn.: Monocerca longicauda Bory de St. Vincent, 1826
Syn.: Trichocerca minor Fadeew, 1925 (synonym considered an infrasubspecific variant of T. rattus)
Trichocerca rectangularis Evens, 1947 (species inquirenda)
Trichocerca ripli Bērziņš, 1973 (species inquirenda)
Trichocerca rosea (Stenroos, 1898): AUS, NEA, NEO, PAL
Mastigocerca rosea Stenroos, 1898
Trichocerca rotundata Myers, 1937: NEA
Trichocerca rousseleti (Voigt, 1902): AFR, AUS, NEA, ORI, PAL
Coelopus rousseleti Voigt, 1902
Syn.: Mastigocerca minima Skorikov, 1903
Trichocerca ruttneri Donner, 1953: AFR, AUS, NEA, NEO, ORI, PAL
Trichocerca scipio (Gosse, 1886): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Mastigocerca scipio Gosse, 1886
Syn.: Trichocerca jenningsi Voigt, 1957
Trichocerca sejunctipes (Gosse, 1886) (species inquirenda)
Rattulus sejunctipes Gosse, 1886
Trichocerca siamensis Segers & Pholpunthin, 1997: NEA, NEO, ORI
Trichocerca similis (Wierzejski, 1893)
Coelopus similis Wierzejski, 1893
Trichocerca similis grandis Hauer, 1965: AFR, AUS, NEO
Trichocerca similis similis (Wierzejski, 1893): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Syn.: Rattulus bicornis Western, 1893 (non Ehrenberg (1832))
Syn.: Mastigocerca birostris Minkiewisz, 1900
Syn.: Mastigocerca blanci Linder, 1904
Syn.: Diurella stylata Eyferth, 1878 (non Gosse (1851))
Syn.: Mastigocerca wolgensis Meissner, 1902
Trichocerca simoneae De Smet, 1990: AFR, NEO, ORI, PAC, PAL (unpublished European records)
Trichocerca stylata (Gosse, 1851): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Monocerca stylata Gosse, 1851
Trichocerca sulcata (Jennings, 1894): AUS, NEA, NEO, PAL
Rattulus sulcatus Jennings, 1894
Syn.: Rattulus cryptopus Bilfinger, 1894
Trichocerca taurocephala (Hauer, 1931): NEA, PAL
Diurella taurocephala Hauer, 1931
Trichocerca tenuidens (Hauer, 1931) (species inquirenda)
Diurella tenuidens Hauer, 1931
Trichocerca tenuior (Gosse, 1886): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Coelopus tenuior Gosse, 1886
Syn.: Mastigocerca flectocaudatus Hilgendorf, 1899
Syn.: Trichocerca neeli Edmondson, 1948
Trichocerca tigris (Müller, 1786): AFR, ANT, AUS, NEA, NEO, ORI, PAC, PAL
Trichoda tigris Müller, 1786
Syn.: Heterognathus macrodactylus Schmarda, 1859
Trichocerca uncinata (Voigt, 1902): NEA, PAL
Coelopus uncinatus Voigt, 1902
Trichocerca unidens (Stenroos, 1898) (species inquirenda)
Mastigocerca unidens Stenroos, 1898
Trichocerca valga (Ehrenberg, 1834) (species inquirenda)
Monocerca valga Ehrenberg, 1834
Trichocerca vargai Wulfert, 1961: PAL
Trichocerca vassilijevae Kutikova & Arov, 1985
Trichocerca vernalis (Hauer, 1936): AUS, NEA, ORI, PAL
Diurella vernalis Hauer, 1936
Trichocerca voluta (Murray, 1913): NEO, ORI
Diurella voluta Murray, 1913
Syn.: Trichocerca tropis Hauer, 1937
Trichocerca wanarra Segers & Shiel, 2003: AUS
Trichocerca weberi (Jennings, 1903): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Diurella weberi Jennings, 1903
Trichocerca weberi (Jennings, 1903): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Diurella weberi Jennings, 1903
Trichotria Bory de St. Vincent, 1827
Dinocharis Ehrenberg, 1830

Trichotria buchneri Koste, Shiel & Tan, 1988: AFR
Trichotria curta (Skorikov, 1914): PAL
Dinocharis curta Skorikov, 1914
Trichotria eukosmeta Myers, 1934: NEA
Trichotria pocillum (Müller, 1776): AFR, AUS, NEA, NEO, ORI, PAL
Trichoda pocillum Müller, 1776
Syn.: Dinocharis pocillum bergi Meissner, 1908
Syn.: Dinocharis pocillum hudsoni Varga, 1930
Syn.: Dinocharis pocillum majus Varga, 1930
Syn.: Furcularia stentorea Bory de St. Vincent, 1825
Trichotria pseudocurta Koste, Shiel & Tan, 1988: AUS
Trichotria tetractis (Ehrenberg, 1830) (includes the infrasubspecific variant turfacea Petr, 1890)
Dinocharis tetractis Ehrenberg, 1830

Trichotria tetractis caudata (Lucks, 1912): AFR, NEA, NEO, ORI, PAL (may be a mere infrasubspecific
variant)
Dinocharis tetractis caudata Lucks, 1912
Trichotria tetractis similis (Stenroos, 1898): AFR, AUS, NEA, NEO, ORI, PAL (may be a mere infra-
subspecific variant)
Dinocharis tetractis similis Stenroos, 1898
Trichotria tetractis strumosa Sudzuki & Huang, 1997 (Described as Trichotria similis strumosa. species
inquirenda)
Trichotria tetractis tetractis (Ehrenberg, 1830): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Syn.: Dinocharis pauper Ehrenberg, 1830 (T. paupera auct. (lapsus))
Syn.: Dinocharis quadrangularis Stirnemann, 1926
Trichotria truncata (Whitelegge, 1889): AUS, NEA, PAL (includes the infrasubspecific variant aspinosa
Rodewald, 1934)
Dinocharis truncatum Whitelegge, 1889
Syn.: Trichotria cornuta Myers, 1938
Syn.: Dinocharis intermedia Bergendal, 1892
Trichotria zanclum Bērziņš, 1982 (species inquirenda)
Tripleuchlanis Myers, 1930
Tripleuchlanis plicata (Levander, 1894): AFR, AUS, NEA, NEO, ORI, PAC, PAL
Euchlanis plicata Levander, 1894
Trochosphaera Semper, 1872
Trochosphaera aequatorialis Semper, 1872: AUS, NEA, NEO, ORI, PAL

Trochosphaera solstitialis Thorpe, 1893: AFR, NEA, ORI, PAL
Tylotrocha Harring & Myers, 1922
Tylotrocha monopus (Jennings, 1894): NEA, PAL

Notommata monopus Jennings, 1894
Wierzejskiella Wiszniewski, 1934
Wierzejskiella ambigua (Tzschaschel, 1979) (marine psammon)

Encentrum ambiguum Tzschaschel, 1979
Wierzejskiella elongata (Glascott, 1893): NEA, PAL
Diglena elongata Glascott, 1893
Syn.: Encentrum lineatum Tzschaschel, 1979 (E. lineatum Tzschaschel erroneously proposed as nom. nov.
for E. elongatum (Wiszniewski) non Harring & Myers)
Wierzejskiella marina Remane, 1949 (marine psammon)
Syn.: Encentrum wierzejskii Tzschaschel, 1979 (E. wierzejskii Tzschaschel proposed as nom. nov. for E.
marinum (Remane) non Dujardin)
Wierzejskiella ricciae (Harring, 1913): NEA, PAL
Encentrum ricciae Harring, 1913
Wierzejskiella sabulosa (Wiszniewski, 1932): PAL
Encentrum sabulosum Wiszniewski, 1932
Wierzejskiella subterranea Remane, 1949 (marine psammon)
Wierzejskiella vagneri Koniar, 1955: PAL
Wierzejskiella velox (Wiszniewski, 1932): AUS, NEA, PAL
Encentrum velox Wiszniewski, 1932
Syn.: Wierzejskiella fluviatilis Neiswestnowa-Shadina, 1935
Wigrella Wiszniewski, 1932
Wigrella amphora (Remane, 1929) (marine psammon)

Encentrum amphora Remane, 1929
Wigrella depressa Wiszniewski, 1932: PAL
Wolga Skorikov, 1903
Wolga spinifera (Western, 1894): AFR, AUS, NEA, NEO, ORI, PAL
Distyla spinifera Western, 1894
Syn.: Cathypna spinifera Murray, 1913
Wulfertia Donner, 1943
Wulfertia kindensis Koste & Tobias, 1990: ORI

Wulfertia kivuensis De Smet, 1992: AFR
Wulfertia ornata Donner, 1943: AFR, NEA, PAC, PAL
Zelinkiella Harring, 1913
Zelinkiella synaptae (Zelinka, 1887) (marine)

Discopus synaptae Zelinka, 1887
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Annotated Checklist of The Rotifers Phylum Rotifer

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Annotated checklist of the rotifers (Phylum Rotifera), with notes on

Article in Zootaxa · August 2007

Rotifers biodiversity in Vietnam View project

The user has requested enhancement of the downloaded file.

Annotated checklist of the rotifers (Phylum Rotifera),

FIRST PUBLISHED IN 2007 BY

© 2007 Magnolia Press

ISSN 1175-5326 (Print edition)

2 · Zootaxa 1564 © 2007 Magnolia Press SEGERS

Annotated checklist of the rotifers (Phylum Rotifera),

Key words: biodiversity, biogeography, Rotifera

Accepted by R. Dalgleish: 26 Apr. 2007; published: 31 Aug. 2007 3

4 · Zootaxa 1564 © 2007 Magnolia Press SEGERS

ANNOTATED CHECKLIST OF THE ROTIFERS Zootaxa 1564 © 2007 Magnolia Press · 5

Anchitestudinella mekongensis B‘rziÃÓ, 1973

Brachionus angularis orientalis Sudzuki, 1989 = B. angularis Gosse, 1851

6 · Zootaxa 1564 © 2007 Magnolia Press SEGERS

B. cluniorbicularis isigakiensis Sudzuki 1992 is an unspined variant (f. cluniorbicularis) of B.

Encentrum grande (Western, 1891) and Encentrum kulmatyckii (Wiszniewski, 1953)

ANNOTATED CHECKLIST OF THE ROTIFERS Zootaxa 1564 © 2007 Magnolia Press · 7

Keratella heywoodi Dartnall, 2005 = K. kostei Paggi, 1981 (new synonym)

Lepadella bractea Myers, 1934 versus Squamella bractea Ehrenberg, 1838.

8 · Zootaxa 1564 © 2007 Magnolia Press SEGERS

ANNOTATED CHECKLIST OF THE ROTIFERS Zootaxa 1564 © 2007 Magnolia Press · 9

10 · Zootaxa 1564 © 2007 Magnolia Press SEGERS

ANNOTATED CHECKLIST OF THE ROTIFERS Zootaxa 1564 © 2007 Magnolia Press · 11

Table 1. List of major literature resources consulted.

Seisonidae: Sørensen et al. (2005)

Asciaporrectidae: De Smet (2006)

12 · Zootaxa 1564 © 2007 Magnolia Press SEGERS

Abrochtha Bryce, 1910

Abrochtha carnivora Ricci, Melone & Walsh, 2001: NEO

Acyclus Leidy, 1882

Hyalocephalus Lucks, 1911

Acyclus inquietus Leidy, 1882: NEA, ORI, PAL

Adineta Hudson & Gosse, 1886

Adineta acuticornis Haigh, 1967: AUS

ANNOTATED CHECKLIST OF THE ROTIFERS Zootaxa 1564 © 2007 Magnolia Press · 13

Anchitestudinella Bērziņš, 1973

Anchitestudinella mekongensis Bērziņš, 1973: ORI

Anomopus Piovanelli, 1903

Anomopus chasmagnathi Mane-Garzon & Montero, 1973: NEO

Anuraeopsis Lauterborn, 1900

14 · Zootaxa 1564 © 2007 Magnolia Press SEGERS

Asciaporrecta De Smet, 2006

Asciaporrecta arcellicola De Smet, 2006: NEA, PAL

Ascomorpha Perty, 1850

Chromogaster Lauterborn, 1893

Ascomorpha agilis Zacharias, 1893: AFR, PAL

Ascomorphella Wiszniewski, 1953

Hertwigella Voigt, 1957

Ascomorphella volvocicola (Plate, 1886): AFR, AUS, NEA, ORI, PAL

ANNOTATED CHECKLIST OF THE ROTIFERS Zootaxa 1564 © 2007 Magnolia Press · 15

Aspelta alastor Harring & Myers, 1928: NEA, PAL

Asplanchna Gosse, 1850

Asplanchnella Sudzuki, 1964

16 · Zootaxa 1564 © 2007 Magnolia Press SEGERS

Asplanchnopus bhimavaramensis Dhanapathi, 1975: ORI

Atrochus Wierzejski, 1893

Atrochus tentaculatus Wierzejski, 1893: AUS, ORI, PAL

Balatro Claparède, 1867

Balatro aciliatus (Radkewitsch, 1870) (Endoparasitic in terrestrial Oligochaeta)