Eco. Env. & Cons. 26 (4) : 2020; pp.

(1826-1833)
Copyright@ EM International
ISSN 0971–765X

An annotated checklist of Terrestrial Isopod Fauna

(Crustacea : Isopoda) of India
Shahnaz Tabassum, Santanu Mitra2 and Seema Langer1

1
Department of Zoology, University of Jammu, Jammu 180 006, J & K, India
2
Zoological survey of India, F.P.S. Building, 27, J.L. Nehru Road, Kolkata 700 016, India

(Received 11 March, 2020; Accepted 22 April, 2020)

ABSTRACT
The present study is an attempt to compile the list of terrestrial isopods found in India to this date, which
are mentioned in various published literature. A total of 78 species belonging to 30 genera and 14 families
are here in listed from Indian states. The highest number of species, genera and family are found from
Tamil Nadu and Andaman and Nicobar and the least are found from Gujarat and Jharkhand. The highest
number of species reported belongs to the genus Cubaris (17 species) followed by genus Adinda (8 species)
and then genus Philoscia with 5 species. Compared to their importance, very little number of studies have
been conducted on the terrestrial isopods in India.

Key words : Tamil Nadu, Terrestrial isopods, Cymothoidea, Oniscidea, Cubaris

Introduction sacchari has been reported to attack and damage

sugarcane buds (David, 1967) in Tanjore district of
Isopoda is one of the most specialized group of Tamil Nadu. However, there are folklores which
crustaceans which has adapted to diverse mode of show some of the isopods with positive qualities. It
life. They are predominantly marine but estuarine is held that the presence of isopods in fishes indicate
and freshwater forms are also well-represented. The that the fish is free from ciguatera (fish poisoning)
suborder Oniscidea contains a large number of crus- toxins (not tested). Dried isopods are used as folk-
taceans which are truly terrestrial. They may be lore medicines in many parts of the world.
free-living, parasitic, predators, filter predators, The order Isopoda which comprises of nine sub-
scavengers, wood-boring, myrmecophilous or cav- orders containing about 11, 000 species, of which,
ernicolous. Their size ranges from 300 micrometer eight suborders namely, Asellota, Cymothoidea,
(Microcerberidae) to about 50 centimeters (as in Limnoridea, Microcerberidea, Oniscidea,
Bathynomus). Phreatoicoidea, Sphaeromatoidea and Valvifera
Compared to prawns, crabs and lobsters, this containing 35 families are herein reported from In-
group of crustaceans is poorly studied by zoologists dia (Dev Roy, 2013). Marine and estuarine isopods
or even less surveyed in many parts of the world are well documented in Indian literature; Dev Roy
although some parasitic and wood-boring forms (2013) listed a total 232 species under 101 genera
cause considerable damage, loss to fisheries, and 25 families from different estuaries in east and
wooden jetties and few others isopods act as pollu- west coast from India. Relatively the work on terres-
tion indicators. Isopods often cause damage to agri- trial isopods is very scattered, and a comprehensive
cultural crops. One species of isopod, Philoscia work on this group deals with only 30 species of 9
TABASSUM ET AL 1827

genera (Ramakrishna, 1995). fauna of Siju cave described two species of terres-
In the present communication, a check-list of 78 trial isopods belonging to the family Oniscidae. In
species of isopods has been prepared providing re- the same year (1924b) he also described two myrme-
cent generic and species names of the group as far cophilous isopods from Barkuda Islands, Chilka
as possible. State-wise distribution of terrestrial iso- Lake, Odisha. Of these two species, Cubaris
pods in India has been presented here. The present granulatus was not known to be associated with ants
communication serves a base line data for this earlier. Verhoeff (1936a) dealt with several species
group and may help the future worker on taxo- of terrestrial Isopods collected from Madras and
nomical or ecological studies. other parts of south India. He further described a
new species of the genus Protracheoniscus from
Review of literature
Ladakh (1936b). Chopra (1947) reported about the
Earlier Budde-Lund (1879) described two species occurrence of the ancient suborder Phreatoicoidea
viz., Tylosalbidus (Tylidae) and Spherillo nicobaricus (Crustacea: Isopoda) for the first time from India
(Armadillidae) from Nicobar Islands. based on collection from a Pucca well at Lohagara
After a short gap, Stebbing (1907) initiated the Railway station, 18 miles from Allahabad. Later,
study of Indian Isopods with the description of a several specimens of this species were collected
new species of the genus Tachaea from Calcutta. In from the wells at Banaras (Uttar Pradesh). In 1950,
1911, he gave a detailed account of Indian isopods Chopra and Tiwari described the genus Nichollsia
and dealt with two genera of the tribe Flabellifera kashiense from the material collected from the well in
and five genera of the tribe Oniscoidea (terrestrial). the outer lawn of the Kaiser Castle, Banaras Cantt.
Two genera viz., Parapericyphis and Exalloniscus Later, Tiwari (1955) described another new species
were erected by him as new to science. of Nichollsia, viz., Nichollsia menoni collected from an
Collinge entered the field and made several con- abandoned well at Monghyr (Bihar). In 1955, he
tributions which received adequate attention. erected a new family Nichollsidae to accommodate
Collinge (1914b, c) published an account of three the genus Nichollsia.
species pertaining to three genera viz., Philoscia, Ramakrishna (1965) recorded several species of
Parapericyphis and Cubaris collected from Port Blair, terrestrial isopods from Kameng Division of the
Andamans and from the Annamalai Hills about North Eastern Frontier Agency (present day
4000 ft. above the sea level. He again (1912-22) con- Arunachal Pradesh). In 1969, he described a new
tributed two papers on the terrestrial isopods which species of Philoscia based on materials collected from
were obtained from the Abor expedition. While a pit and the surrounding galleries of Lodna
working out the material, he came across two gen- Colliery, 13 km from Dhanbad, Bihar (now in the
era new to science viz., Rotungus and Burmoniscus, state of Jharkhand). Some terrestrial isopods of In-
the former was obtained from Kobo, Abor country dia have been investigated by Ram and Kumar
(present day Arunachal Pradesh) at an altitude of (1979).
400 ft. and the later from the caves near Moulmein In recent times, several workers, Ferrara and Taiti
(Myanmar). Apart from these two genera, he has (1982) and Ferrara et al. (1995) also studied terres-
also described six other species of which, three are trial isopods of India, including Andaman and
discovered for the first time. His next contribution Nicobar Islands, and described several new species.
to our knowledge of terrestrial isopods of India
dates back to 1914, when he worked out the collec- Results and Discussion
tion received from Madras Province. Of the ten spe-
cies dealt with nine species were new to science. The recent review implies that, there are only 78
Ennurensis hispidus and Hemiporcellio carinatus stand species of terrestrial isopods belonging to 2 subor-
significant among this collection. In 1916, Collinge der, 14 families and 30 genera are reported from this
described 13 new species of the genera country (Table 1). Results suggest that Tamil Nadu
Parapericyphis, Cubaris, and Burmoniscus. Among reaches highest number of species diversity; fol-
these, Burmoniscus kempi was collected from lowed by Kerala, Andaman & Nicobar Islands,
Maosmai cave near Cherrapunji at an altitude of Odisha and West Bengal (Table 2). There are many
4,000 ft. states like Gujarat, Jharkhand, Lakshadweep,
Chopra (1924a) while working on the isopod Manipur and Punjab from where very few terres-
1828 Eco. Env. & Cons. 26 (4) : 2020

Table 1. Distributional list of all terrestrial isopods in India

Distribution Name of the species
Suborder ONISCIDEA
Family AGNARIDAE Schmidt, 2003
Agnara Budde-Lund, 1908
Odisha: Chilka lake, Rambha, Ganjam dist. (Collinge, 1915) 1. Agnara carinata (Collinge, 1915)
Odisha: Satpara, Chilka lake (Collinge, 1915) 2. Agnara hispida (Collinge, 1915)
Uttar Pradesh: Allahabad (Collinge, 1914) 3. Agnara immsi (Collinge, 1914)
Alloniscus Dana, 1854
Nicobar Island (Budde-Lund, 1885); Northeast India (Taiti, 2014) 4. Alloniscus nicobaricus (Budde-Lund, 1885)
Odisha: Chilika (Chilton, 1916); Andaman & Nicober Island 5. Alloniscus pigmentatus (Budde-Lund, 1885)
(Budde-Lund, 1885), Lakhshadweep (Taiti, 2014)
Protracheoniscus Verhoeff, 1917
Uttar Pradesh: Lucknow (Stebbing, 1911) 6. Protracheoniscus asiaticus (Uljanin, 1875)
Jammu and Kashmir: Ladak (Nubra, Shylok and 7. Protracheoniscus Karakorum (Jackson, 1935)
Khardung valley) (Jackson, 1935)
Jammu and Kashmir: Karghalik and Yarkand 8. Protracheoniscus maracandicus (Uljanin, 1875)
(Ramakrishna, 1995)
Jammu and Kashmir: Ladak (Verhoeff, 1936) 9. Protracheoniscus nivalis(Verhoeff, 1936)
Family ARMADILLIDAE Brandy, 1831
Cubaris Brandt, 1833
Andhra Pradesh: Kamalapuram (Collinge, 1916) 10. Cubaris albolateralis (Collinge, 1916)
Meghlaya: caves of Cherrapunji (Collinge, 1916) 11. Cubaris cavernosa (Collinge, 1916)
Kerala: Puenjikara island, near Ernakulam, Kochi 12. Cubaris chiltoni (Collinge, 1916)
(Collinge, 1916);
West Bengali: Kalimpong, Darjeeling dist. (Collinge 1916); 13. Cubaris dilectum (Collinge, 1916)
Odisha: Barkuda Island, Chilka lake, Ganjam dist. 14. Cubaris expansa (Collinge, 1916)
(Collinge, 1916);
Odisha: Rambha, Chilka lake, Ganjam dist. (Collinge 1915); 15. Cubaris granulata (Collinge, 1915)
Tamil Nadu: Madras (Collinge, 1915); Odisha: Balugaon,
Barkuda island (Chopra, 1924)
Jharkhand: Pass between ChaibassaandChakardharpur, 16. Cubaris gravelii (Collinge, 1916)
Chotanagpur (Collinge, 1916);
Jammu & Kashmir: Sringar (Verhoeff, 1936) 17. Cubaris ignota (Arcangeli, 1934)
Jammu & Kashmir: Sringar (Jackson, 1935) 18. Cubaris kashmiri (Jackson, 1935)
Kerala: Parambiculam, Kochi (Collinge, 1916) 19. Cubaris lobata (Collinge, 1916)
Arunachal Pradesh: Kobo (Collinge, 1916); Kemang division 20. Cubaris marmorata (Collinge, 1916)
(Ramakrishna, 1965)
Andaman island: Port blair (Collenge, 1915); 21. Cubaris murina (Brandt, 1833)
Odisha: Rambha, Ganjam dist. (Collinge, 1915) 22. Cubaris nacrum (Collinge, 1915)
Bihar: Patna (Ram & Kumar, 1979) 23. Cubaris pataliputraensis (Ram & Kumar, 1979)
Maharashtra: Kas, Satara dist., Bombay pres. (Collinge, 1916); 24. Cubaris pusilla (Collinge, 1916)
Assam: Sadyia (Collinge, 1914) 25. Cubaris robusta (Collinge, 1914)
Karnataka: Oorgaum, Kolar dist. (Collinge, 1915) 26. Cubaris solidula (Collinge, 1915)
Hybodillo Taiti, Paoli & Ferrara, 1998
Andaman Island (Ferrera and Taiti, 1982) 27. Hybodillo monocellatus (Ferrara & Taiti, 1982)
Laureola Barnard, 1960
Kerala: Travancore (Kwon, Ferrara & Taiti, 1993) 28. Laureola indica Kwon, (Ferrara & Taiti, 1992)
Madrasdillo Arcangeli, 1957
Tamil Nadu: Madras (Kwon, Ferrara & Taiti, 1993) 29. Madrasdillo elevatus (Verhoeff, 1936)
Nesodillo Verhoeff, 1926
Kerala: Kovalam (Verhoeff, 1936) 30. Nesodillo jonesi (Verhoeff, 1936)
West Bengal: Darjeeling (Verhoeff, 1928) 31. Nesodillos chellenbergi (Verhoeff, 1928)
TABASSUM ET AL 1829

Table 1. Continued ...

Distribution Name of the species
Spherillo Dana, 1853
Nicobar Island (Budde-Lund, 1879) 32. Spherillo nicobaricus (Budde-Lund, 1885)
Family Eubelidae Budde-Lund, 1899
Saidjahus Budde-Lund, 1904
Mandapam, Pamban passage, Tamil Nadu (Stebbing, 1911) 33. Saidjahus sp. (Budde-Lund,1904)
Family HALOPHILOSCIIDAE Verhoeff, 1908
Littorophiloscia Hatch, 1947
Andaman island (Ferrara & Taiti, 1982) 34. Littorophiloscia denticulata (Ferrara & Taiti, 1982)
Family LIGIIDAE Leach, 1814 Ligia Fabricius, 1798
Andaman & Nicober Is. (Ferrera and Taiti, 1982) 35. Ligia dentipes (Budde-Lund, 1885)
Along the coast of India (Ferrera and Taiti, 1982) 36. Ligia (Megaligia) exotica (Roux, 1828)
Ligidium Brandt, 1833
Andhra pradesh: Vishakhapatnam (Kumari, 37. Ligidum rishikondensis (Kumari,
Hanumantha-Rao & Shyamasundari, 1989) Hanumantha-Rao & Shyamasundari, 1989)
Family OLIBRINIDAE Budde-Lund, 1913
Olibrinus Budde-Lund, 1913
Nicober island (Barnard, 1936) 38. Olibrinus antennatus (Budde-Lund, 1902)
Family ONISCIDAE Latreille, 1806 Exalloniscus Stebbing, 1911
Kerala: Madathorai (Ramakrishna, 1995) 39. Exalloniscus coecus (Dollfus, 1898)
Rotungus Collinge, 1916
Arunachal Pradesh: Kobo (Collinge, 1916; Ramakrishna, 1995). 40. Rotungus pictus (Collinge, 1916)
Family PHILOSCIIDAE Kinahan, 1857
Anchiphiloscia Stebbing, 1908
Andaman Island (Ferrera and Taiti, 1982) 41. Anchiphiloscia bicolorata (Ferrara & Taiti, 1982)
Andaman Island (Ferrara and Taiti, 1982) 42. Anchiphiloscia longisetosa (Ferrara & Taiti, 1982)
Arhina Budde-Lund, 1904
Odisha: Barkul, Chilka lake (Collinge, 1915); 43. Arhina barkulensis (Collinge, 1915)
Burmoniscus Collinge, 1914
Arunachal Pradesh, West Bengal, Madhya Pradesh, 44. Burmoniscus javanensis (Richardson, 1922)
Uttar Pradesh, Himachal Pradesh, Haryana, Silent Valley
in Kerala, Tamil Nadu, Karnataka, Goa and Nicobar-Islands
(Ramakrishna, 1995)
Meghalaya: Maosmai Cave, Cherrapunji (Collinge, 1916; 45. Burmoniscus kempi (Collinge, 1916
Ramakrishna, 1995)
Dekanoscia Verhoeff, 1936
Tamil Nadu: Madras, Kerala: Ponmudi Hills (Verhoeff, 1936) 46. Dekanoscia longicornis (Verhoeff, 1936)
Philoscia Latreille, 1804
Philoscia Latreille, 1804
Meghalaya: Siju Cave, Garo Hills (Ramakrishna, 1995); 47. Philoscia dobakholi (Chopra, 1924)
Kerala: Silent Valley (Ramakrishna, 1995); 48. Philoscia indirae(Ramakrishna, 1995)
Bihar: Lodna colliery, 13 km. from Dhanbad (Ramakrishna, 49. Philoscia lodnensis (Ramakrishna, 1969)
1995); Arunachal Pradesh, Assam, Meghalaya, Manipur,
Madhya Pradesh, Rajasthan, Orissa, West Bengal, Kerala,
Tamil Nadu, Karnataka, Maharashtra and Goa
(Ramakrishna, 1995);
Arunachal Pradesh: Kemang division 50. Philoscia muscorum (Scopoli, 1763)
(Ramakrishna 1965, 1995);
Tamil Nadu: Tanjore (David, 1967) 51. Philoscia sacchari (David, 1967)
Tamil Nadu (Collinge, 1915) 52. Philoscia tenuissima (Collinge, 1915)
Family PLATYARTHRIDAE Verhoeff, 1949
PlatyarthrusBrandt, 1833
Odisha: Barakuda island, Chilka lake (Chopra, 1924); 53. Platyarthrus acropyga (Chopra, 1924)
1830 Eco. Env. & Cons. 26 (4) : 2020

Table 1. Continued ...

Distribution Name of the species
Family PORCELLIONIDAE Brandt & Ratzeburg, 1831
Ennurensis Collinge, 1915
Tamil Nadu: Ennur near Madras (Collinge, 1915, 54. Ennurensis hispidus (Collinge, 1915)
Ramakrishna, 1995), Mandapam, Pamban passage
(Ramakrishna, 1995); Odisha: Satpara, Chilka lake
(Ramakrishna, 1995);
Porcellio Latreille, 1804
Meghalaya: MlawailExamulSiju cave, Garo hill 55. Porcellio assamensis (Chopra, 1924)
(Chopra, 1924); Arunachal Pradesh: Kamengdivison
(Ramakrishna, 1965); Arunachal Pradesh, Assam,
Manipur, Bihar, Orissa, Madhya Pradesh, West Bengal,
Kerala, Tamil Nadu, Karnataka, Goa and Andaman &
Nicobar Islands (Ramakrishna, 1995)
Rajasthan: Tolla Berra, about 5 kms. From Pokran on 56. Porcellio ganesa (Ramakrishna, 1975)
Ramdeora Road (Ramakrishna, 1975, 1995); Maharashtra,
Andhra Pradesh and Tamil Nadu (Ramakrishna, 1995)
West Bengal, Bihar Madhya Pradesh, Uttar Pradesh, 57. Porcellio laevis (Latreille, 1804)
Punjab, Rajasthan, Gujarat, Maharashtra, Tamil Nadu
(Ramakrishna, 1995)
Assam, Meghalaya, West Bengal, Orissa, Madhya Pradesh, 58. Porcellio scaber (Latreille, 1804)
Himachal Pradesh, Haryana, Punjab, Rajasthan,
Maharashtra, Karnataka, Tamil Nadu (Ramakrishna, 1995)
Orissa, Bihar: Chhota Nagpur, Madhya Pradesh, 59. Porcellio spinicornis (Say, 1818)
Tamil Nadu, Maharashtra, Rajasthan, Haryana,
Himachal Pradesh (Ramakrishna, 1995)
Porcellionides Miers, 1877
West Bengal: Kurseong (Collinge, 1914, Stebbing, 1911); 60. Porcellionides pruinosus (Brandt, 1833)
Assam: Sadyia (Collinge 1914); Arunachal Pradesh: Kobo,
Rotung (Collinge, 1914), Kemangdivison (Ramakrishna,
1965); Assam, Meghalaya, Bihar, West Bengal,
Madhya Pradesh, Uttar Pradesh, Himachal Pradesh,
Haryana, Punjab, Rajasthan, Andaman & Nicobar island,
Lakshadweep island (Ramakrishna, 1995);
Family SCLEROPACTIDAE Verhoeff, 1938
Adinda Budde-Lund, 1904
Kerala: Palni hills, tandikdi, meryland shola (Ferrara, 61. Adinda carli (Ferrara, Meli & Taiti, 1995)
Meli & Taiti, 1995)
Kerala: Ponmudi, Travancore (Collinge, 1915) 62. Adinda gigas (Collinge, 1915)
Tamil Nadu: Annamalai hills,Valparai, Naduar estate 63. Adinda lobata (Ferrara, Meli & Taiti, 1995)
(Ferrara, Meli & Taiti, 1995)
Kerala: Nilgiri hills (Ferrara, Meli & Taiti, 1995) 64. Adinda nilgiriensis (Ferrara, Meli & Taiti, 1995)
Kerala: Palni hills (Ferrara, Meli& Taiti, 1995) 65. Adinda palniensis (Ferrara, Meli & Taiti, 1995)
Tamil Nadu: Anamalai hill, Madras (Collinge, 1915); 66. Adinda stebbingi (Collinge, 1914)
Kerala: Cochin (Collinge, 1916, Ramakrishna, 1993);
Kerala: Maddathoray, Travancore (Ramakrishna, 1993) 67. Adinda travancorensis (Stebbing, 1911)
Tamil Nadu: Annamalai hills, Attakatti shola, (Ferrara, 68. Adinda triangulifera (Ferrara, Meli & Taiti, 1995)
Meli & Taiti, 1995)
Family TRACHELIPODIDAE Strouhal, 1953
Nagurus Holthuis, 1949
Andaman Islands (Ferrara & Taiti, 1982). 69. Nagurus acutitelson (Ferrara and Taiti, 1982)
Orissa: Chilka Lake, Rambha dist. (Ramakrishna, 1995) 70. Nagurus carinatus (Dollfus, 1905)
TABASSUM ET AL 1831

Table 1. Continued ...

Distribution Name of the species
Tamil Nadu: Madras (Verhoeff, 1936) 71. Nagurus clavigerus (Verhoeff, 1936)
Andaman Islands (Ferrara & Taiti, 1982) 72. Nagurus havelocki (Ferrara & Taiti, 1982)
Kerala: Kovalam (Verhoeff, 1936) 73. Nagurus travancorius (Verhoeff, 1936)
Trachelipus Budde-Lund, 1908
Assam, Meghalaya, Arunachal Pradesh, Bihar, West Bengal, 74. Trachelipus rathkii (Brandt, 1833)
Orissa, Madhya Pradesh, Uttar Pradesh, Rajasthan, Haryana,
Himachal Pradesh, Punjab, Maharashtra and Karnataka,
Tamil Nadu, Andhra Pradesh (Ramakrishna, 1995)
Himachal Pradesh, Haryana, West Bengal (Ramakrishna, 75. Trachelipus ratzeburgii (Brandt, 1833)
1995)
Family: TYLIDAE Dana, 1852
Tylos Audouin, 1826
Nicober Island (Budde-Lund, 1885) 76. Tylos albidus (Budde-Lund, 1885)
Suborder PHREATOICIDEA
Family HYPSIMETOPIDAAE Nicholls, 1943
Nichollsia (Chopra & Tiwari, 1950)
Uttar Pradesh: Benaras (Chopra & Tiwari, 1950) 77. Nichollsia kashiense (Chopra & Tiwari, 1950)
Bihar: Monghyr (Tiwari, 1955) 78. Nichollsia menoni (Tiwari, 1958)

trial isopods have been reported till date. Report Table 2. State wise list of all family, genus and species of
also shows that in many states, still no work has all terrestrial isopods in India
been initiated for this particular group. Chandigarh, Species Genus Family Name of States
Chattisgarh, Dadra & Nagar Haveli, Daman & Diu,
5 5 4 Andhra Pradesh
Delhi, Mizoram, Nagaland, Puducherry, Sikkim,
17 13 9 Andaman & Nicobar
Telengana, Tripura and Uttarakhand have no report
Island
of occurrence of any species of terrestrial isopod. So, 8 7 5 Arunachal Pradesh
an elaborative studyis very much in demand for 6 5 4 Assam
these animals. 8 6 5 Bihar
4 4 3 Goa
Habit and habitat
1 1 1 Gujarat
Terrestrial isopods generally live on the decaying 6 4 3 Haryana
matters on soils, under leaf litters, stone or barks of 6 4 3 Himachal Pradesh
woods. They mostly prefer to stay in moist places 5 2 2 Jammu & Kashmir
1 1 1 Jharkhand
but some group are specialized to stay in almost
7 6 5 Karnataka
dried up parts of soils (Armadillidae); some genus 18 11 7 Kerala
Porcellio and Philoscia are commonly live under the 2 2 2 Lakshadweep
flowering pots, some are adapted in cavernicolous 8 5 3 Madhya Pradesh
habitat. A very few species found in subterranean 7 4 4 Maharashtra
habitat. 2 2 2 Manipur
They usually feed in night, on vegetable matters 8 6 4 Meghalaya
with algae, fungi, moss, bark, and any type of de- 16 11 7 Odisha
caying vegetables, occasionally they feed on animal 4 3 2 Punjab
7 4 3 Rajasthan
matters also. There are some species lives that
20 12 7 Tamil Nadu
shows commensalism with ants, which feeds on fae- 7 7 5 Uttar Pradesh
cal matter of the ants. A few isopods are carnivorous 10 7 4 West Bengal
in their feeding habit.
Significance of this group groups that may help in decomposition of leaf litters
and by this way they may be considered as the im-
Terrestrial Isopods are one of the major invertebrate portant component of food chain. The leaf litter de-
1832 Eco. Env. & Cons. 26 (4) : 2020

composition is carried out by the combined action of Collinge, W. E. 1916.Contributions to a knowledge of the
microorganisms and decomposer invertebrates such terrestrial isopoda of India. Part 2. Some new species
as earthworms, diplopods and terrestrial isopods. of Paraperiscyphis, Cubaris, etc. Rec. Indian Mus. 12(3):
The role of this animals in nutrient cycle are well 115-128.
David, H. 1967. Philoscia sacchari- a new soil isopod dam-
studied in abroad (Khalaeid and Walkail, 2015).
aging sugarcane buds in Tanjore district (Crustacea:
Some isopods are considered as pests in vegetable Isopoda). Journal Zoological Society of India. 17 : 120-
gardens and in horticulture. Cotton and lime trees 122.
are affected by isopod pests in damaging their barks Dev Roy, M. K. 2013. Marine and estuarine isopod fauna
(Collinge, 1914). (Crustacea: Isopoda) of India. Journal of Environment
and Sociobiology. 10(2): 147-178.
Acknowledgement Ferrara, F. and Taiti, S. 1982. Isopoditerrestridelle Isole
Andamane. Bollettino del Museo Civico di Storia Natu-
rale Verona. 8 : 459-492.
One of the authors (S.M) wish to express his deep-
Ferrara, F., Meli, C. and Taiti, S. 1995. Taxonomic revision
felt gratitude and thanks to Dr. Kailash Chandra,
of the subfamilyToradjiinae (Crustacea: Oniscidea:
Director, Zoological survey of India, for providing Scleropactidae). Zoological journal of Linnean Society.
facilities to complete this work. 113 : 351-459.
Jackson, H. G. 1935. Isopoda terrestria. In: P. C. Visser,
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