MULLER 2009 Checklist Chile

Mosses of Chile 1
An updated checklist of the mosses of Chile
Frank Müller
Abstract: The distribution by regions and provinces is provided for each of the 890 moss taxa
currently accepted in Chile. All records are supported by literature references, and the number of
taxa recorded for each region is given. Synonyms with their owning taxa are intercalated
alphabetically in the list.
Introduction
The last checklist of the mosses of Chile was completed in the end of 1996 and was published in
1998 (He 1998). Since this time a considerable bulk of new data on the mosses of Chile has been
published. This provided the stimulus to produce a new checklist. The present study attempts an
updated version of the checklist of He (1998) and contains all data available up to the end of
November 2009.
For a lot of families and genera of the area critical taxonomic revisions were published in the last
years. This includes Amblystegiaceae (Hedenäs 2003, Ochyra & Matteri 2001), Bartramia
(Fransén 2004), Campylopus (Frahm 2005), Fissidens (Müller & Pursell 2003), Grimmia (Muñoz
1999, Muñoz & Panda 2000), Hypopterygiaceae (Kruijer 2002), Lepyrodon (Allen 1999),
Meesiaceae (Matteri & Ochyra 1999), Tortula (Cano & Gallego 2008), and Zygodon (Calabrese
2006). In addition, there is also a remarkable progress in the floristic investigation of Chile (Isla
Navarino in province Antártica Chilena: Buck 2002, Goffinet et al. 2006; regions Tarapacá and
Antofagasta: Cano 2003; the Los Lagos and Los Ríos regions: Frey & Schaumann 2002,
Schaumann 2005, Larrain 2005, 2007, 2007b; Easter Island: Ireland & Bellolio 2002; the Bío-Bío
region: Ireland et al. 2007; the Maule Region: Müller & Pereira 2006, Pereira et al. 2006).
In addition, new information’s resulted in checklists and floras of neighboring regions, e. g., the
new checklist of Argentina (Matteri 2003b), the Flora of King George Island, Antarctica (Ochyra
1998), and the Flora of Antarctica (Ochyra et al. 2008).
Taxonomic revisions are just on the way for Syntrichia (M.J. Cano, M.T. Gallego) and
Racomitrium s.l. (J. Larraín), so in next future the certain determination of Chilean specimens of
these groups will be possible. Nevertheless, there still are problematic genera of which
determination of Chilean material is at the moment difficult. These include Andreaea,
Dicranoloma, Hymenoloma, Ditrichum, Macromitrium, Mielichhoferia, Pohlia, Ptychomitrium,
Rhaphidorrhynchium, Schistidium, Schizymenium, Sematophyllum, and Sphagnum. Hopefully, in
next future these genera will be dealt in systematic revisions.
For the compilation of the present checklist there were evaluated the following sources:
1.) The checklist published by He (1998).
2.) About 200 additional publications newly published ones and older ones not included in He
(1998).
3.) The herbarium database of the herbarium of New York (NY) and the Tropicos database of the
Missouri Botanical Garden (MO).
ARCHIVE FOR BRYOLOGY 58 (2009)

2 Müller
4.) Results of the investigation of herbarium material deposited in the herbaria’s of Paris (PC) and
Stockholm (S).
5.) Results of the determination of Chilean moss material in the herbarium of Dresden (DR).
Important records of this were already published separately (Müller 2002, Müller 2009).
The arrangement of the checklist
The arrangement of the alphabetically list of recorded taxa follows He (1998). To each taxon
references are given. To show the range of each taxon in Chile, its regional and provincial
distribution is given.
Chile is divided into 15 regions. Every region is further divided into provinces. Each region is
designated by a name and a Roman numeral. The only exception is the region housing the nation's
capital, which is designated RM, that stands for Región Metropolitana (Metropolitan Region).
Two new regions, Arica and Parinacota in the north, and Los Ríos in the south, were created in
2006, and became operative in October 2007. In the numbering scheme, Region XIII was skipped;
Arica and Parinacota was designated Region XV, while Los Ríos was designated Region XIV.
The division of Chile in political regions is shown in Figure 1. The regional distribution of each
taxon is given in the list in the order from north to south.
The number of accepted taxa recorded for each political region of Chile as well as for Juan
Fernández Islands and Easter Island is provided in Table 1. To show the progress in the
bryological exploration of the regions the species numbers of the previous checklist of He (1998)
are given.
All validly published names that are widely accepted or that reflect opinions expressed in
published monographs are written in the checklist in bold face italic font. Taxa with insufficiently
known status, e. g., mostly nomina nuda, and illegitimate names, are written in non bold face italic
font.
Heterotypic synonyms or taxonomic synonyms are marked by an equal sign (=) followed by a
reference of an author. Homotypic or nomenclatural synonyms are indicated by an identity sign
(≡).
In Chile, 890 moss taxa are currently accepted from 230 genera and 68 families. The systematic
arrangement of the genera and families follows with minor modifications the sequence of Goffinet
et al. (2008).
Despite checking and re-checking, some errors doubtless remain undetected in this document, and
literature overlooked, and the author would be very grateful to know of any of these. Comments
please send to Frank.Mueller@tu-dresden.de
This document is not intended as a taxonomic work. The quality of the data in this list is variable
in its taxonomic quality, although it attempts to present the latest position for all taxa.
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Mosses of Chile 3
Table 1. Number of accepted taxa reported for political regions of Chile
Number Number of
of taxa taxa
(He (current list)
Region Provinces 1998)
XV. Arica y
Arica, Parinacota 22
Parinacota 0
I. Tarapacá El Tamarugal, Iquique 0
II. Antofagasta Antofagasta, El Loa, Tocopilla 8 19
III. Atacama Chañaral, Copiapó, Huasco 8 12
IV. Coquimbo Choapa, Elqui, Limarí 66 87
Los Andes, Petorca, Quillota, San Antonio,
V. Valparaíso 169 168
San Felipe de Aconcagua, Valparaíso
Chacabuco, Cordillera, Maipo, Melipilla,
RM. Metropolitana 136 136
Santiago, Talagante
VI. O'Higgins Cachapoal, Cardenal Caro, Colchagua 53 63
VII. Maule Cauquenes, Curicó, Linares, Talca 65 163
VIII. Biobío Arauco, Biobío, Concepción, Ñuble 190 315
IX. La Araucanía Cautín, Malleco 234 251
XIV. Los Ríos Valdivia, Ranco 282
412
X. Los Lagos Chiloé, Llanquihue, Osorno, Palena 346
Aisén, Capitán Prat, Coihaique, General
XI. Aisén 310 337
Carrera
Antártica Chilena, Magallanes, Tierra del
XII. Magallanes 450 436
Fuego, Última Esperanza
Isla Robinson Crusoe, Isla Alejandro
Juan Fernández 158 153
Selkirk
Isla de Pascua (Easter
0 34
Island)

4 Müller
Fig. 1: Political regions of Chile (from He 1998, modified).
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Mosses of Chile 5
Taxa reported from Chile arranged alphabetically by genera and species
The checklist is in alphabetical order of taxon name, with current names shown in bold face italic
font. Illegitimate names, nomina nuda, and doubtful taxa attributed to Chile shown in non bold
face italic font. Names placed in synonymy are indented and shown in a smaller font, together with
the owning taxon and a literature source for the synonymy.
Literature references are given for each taxon. To show the regional distribution for each taxon
abbreviations of the regions and the names of the provinces from where it is recorded, are
provided. The abbreviations of the regions follow the scheme of Table 1. The regional distribution
is given in the order from north to south. For the recently created Los Ríos region (region XIV) it
was impossible to recheck all the literature sources to find out whether the records are from
province Valdivia or from the newly created province Ranco. Therefore for records from this
region no detailed provincial distribution is provided.
Acaulon uleanum Müll. Hal.

Ireland et al. 2007. Dist. VIII. Concepción.
Acaulon spec.
Matcham et al. 2007. Dist. XII. Magallanes.
Achrohypnella subenervis Herzog = Vesiculariopsis spirifolium fide Seki 1974.
Achrophyllum anomalum (Schwägr.) H. Rob.
Larraín 2007b; Larraín 2007a; He 1998; Robinson 1975; Matteri & Schiavone 2002 (as
Pterygophyllum anomalum, Espinosa 1941; Mitten 1869; Dusén 1903b; Cardot 1908;
Brotherus 1924b; Herzog 1954; Bartram 1957; as Pterygophyllum obscurum, Mitten 1869;
Dusén 1903b; Cardot 1908; Herzog 1922, 1954, 1960; Brotherus 1924b; Reimers 1926; as
Pterygophyllum obscurum fo. thermalis, Herzog & Schwabe 1939). Dist. IX. Malleco; XIV.
Valdivia/Ranco; X. Chiloé, Osorno; XI. Aisén; XII. Antártica Chilena; Juan Fernández Is.
Achrophyllum anomalum var. pallidum (Cardot & Broth.) S. He
(as Pterygophyllum anomalum var. pallidum, Cardot & Brotherus 1923). Dist. XI. Capitán
Prat.
Achrophyllum crassirete (Matteri) Matteri
Tropicos 2009; Hässel et al. 1984; Larraín 2005; Matteri 1979, 1985b; Deguchi 1991a (as
Pterygophyllum crassirete, Matteri 1972; Seki 1974). Dist. V. Valparaíso; IX. Cautín; XIV.
Valdivia/Ranco; X. Llanquihue, Osorno; XI. Aisén; XII. Magallanes, Última Esperanza.
Achrophyllum dentatum (Hook.f. & Wilson) Vitt & Crosby
Villagran et al. 2003; Robinson 1975 (as Pterygophyllum denticulatum, Mitten 1869; Dusén
1903b; Cardot 1908; Thériot 1917; Cardot & Brotherus 1923; Brotherus 1924b; Herzog 1954;
as Pterygophyllum dentatum, Bartram 1957). Dist. VI. Colchagua; X. Chiloé, Osorno; XI.
Aisén; XII. Antártica Chilena; Juan Fernández Is.
Achrophyllum haesselianum (Matteri) Matteri
Hässel et al. 1984; Buck 2002; He 1998; Matteri 1979, 1985b; Matteri & Schiavone 2002 (as
Pterygophyllum haesselianum, Matteri 1972; Seki 1974). Dist. XI. Aisén; XII. Antártica
Chilena, Magallanes, Tierra del Fuego, Última Esperanza.
Achrophyllum magellanicum (Besch.) Matteri var. magellanicum
Tropicos 2009; Larraín 2007b; Hässel et al. 1984; Larraín 2005; Villagran et al. 2003;
Schaumann 2005; He 1998; Müller & Pereira 2006; Matteri 1979, 1985b; Deguchi 1991a;
Matteri & Schiavone 2002 (as Pterygophyllum magellanicum, Dusén 1903b; Cardot 1908,
Matteri 1972; Seki 1974). Dist. VII. Cauquenes, Talca; VIII. Arauco, Biobío, Concepcion,
Ñuble; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno; XI. Aisén;
XII. Antártica Chilena, Magallanes, Última Esperanza.
Achrophyllum magellanicum var. oligodontum (Matteri) Matteri

6 Müller
Tropicos 2009; Hässel et al. 1984; Buck 2002; Matteri 1979, 1985b; Deguchi 1991a; Matteri &
Schiavone 2002 (as Pterygophyllum magellanicum var. oligodontum, Matteri 1972; Seki
1974). Dist. VIII. Biobío, Concepcion; IX. Cautín, Malleco; X. Chiloé, Llanquihue; XI. Aisén;
XII. Antártica Chilena, Magallanes, Tierra del Fuego, Última Esperanza.
Achrophyllum tenuinerve (Broth.) H. Rob.
Robinson 1975 (as Pterygophyllum tenuinerve, Barrera 2006; Espinosa 1941; Brotherus 1924b;
Bartram 1957). Dist. Juan Fernández Is.
Acrocladium auriculatum (Mont.) Mitt.
Goffinet et al. 2006; Larraín 2007b; Hässel et al. 1984; Matcham et al. 2007; Larraín 2005;
Ireland et al. 2007; Buck 2002; Villagran et al. 2003; Schaumann 2005; He 1998; Ochyra &
Matteri 2001; Mitten 1869; Dusén 1903b; Cardot 1908; Herzog 1922, 1954, 1960; Cardot &
Brotherus 1923; Roivainen 1934; Thériot 1934b; Herzog & Schwabe 1939; Bartram 1952;
Seki 1974; Robinson 1975; Matteri 1985b; Deguchi 1991a; Matteri & Schiavone 2002 (as
Acrocladium chlamydophyllum, Dusén 1903b; Cardot 1908; as Lembophyllum auriculatum,
Cardot 1900, 1901, 1905; as Calliergon auriculatum, Margadant 1972). Dist. VIII. Arauco,
Biobío, Ñuble; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno,
Palena; XI. Aisén, Coihaique; XII. Antártica Chilena, Magallanes, Tierra del Fuego, Última
Esperanza; Juan Fernández Is.
Acrocladium chlamydophyllum (Hook.f. & Wilson) Müll. Hal. & Broth. = Acrocladium auriculatum fide Wijk et al.
1959.
Acrocladium cuspidatum (Hedw.) Lindb. ≡ Calliergonella cuspidata.
Acrocladium politum (Hook.f. & Wilson) Mitt. = Catagonium nitens subsp. nitens var. nitens fide Lin 1984.
Acroschisma wilsonii (Hook. f.) A. Jaeger
NY 2009; Müller 2008; Larraín 2005; Buck 2005a; He 1998; Dusén 1903a; Cardot 1908;
Roivainen 1937; Seki 1974; Deguchi 1991a; Matteri & Schiavone 2002 (as Andreaea wilsonii,
Hässel et al. 1984; Mitten 1869; Cardot & Brotherus 1923; Bartram 1952; Herzog 1954;
Matteri 1985b; as Felipponea hollermayeri, Thériot 1935). Dist. XIV. Valdivia/Ranco; X.
Llanquihue, Osorno, Palena; XI. Aisén; XII. Antártica Chilena, Magallanes, Tierra del Fuego,
Última Esperanza.
Aloina catillum (Müll. Hal.) Broth.
Cano et al. 2008a. Dist. XV. Parinacota.
Aloina bifrons (De Not.) Delgad.
Cano et al. 2008a; Villagran et al. 2004. Dist. IV. Limarí; RM. Cordillera.
Aloina brevirostris (Hook. & Grev.) Kindb.
Matcham et al. 2007; Matcham & Duckett in Blockeel et al. 2008a. Dist. XII. Magallanes.
Aloina roseae (R.S. Williams) Delgad.
Delgadillo 1973; Jiménez & Cano 2007 (as Crossidium roseae, Bartram 1943, 1952, 1953).
Dist. II. Antofagasta; III. Copiapó; IV. Elqui.
Aloinella andina Delgad. ≡ Aloinella galeata var. andina.
Aloinella galeata var. andina (Delgad.) Delgad. & Schiavone
(as Aloinella andina, Cano 2003). Dist. XV. Parinacota.
Amblyodon dealbatus (Sw. ex Hedw.) Bruch & Schimp.
Matteri & Schiavone 2002; Matteri & Ochyra 1999. Dist. XII. Tierra del Fuego.
Amblystegium chilense Lorentz = Cratoneuropsis relaxa subsp. minor fide Ochyra et al. 2008.
Amblystegium excurrens Cardot & Broth. = Cratoneuropsis relaxa subsp. minor fide Ochyra et al. 2008.
Amblystegium fuegianum Mitt. = Warnstorfia fluitans fide Ochyra & Matteri 2001.
Amblystegium jaffueli Thér. = Leptodictyum riparium fide Ireland et al. 2007.
Amblystegium philonotis Dusén, nom. nud. = Cratoneuropsis relaxa subsp. minor fide Ochyra et al. 2008.
Amblystegium serpens (Hedw.) Schimp.
Tropicos 2009; Müller 2009; Hässel et al. 1984; He 1998; Cardot & Brotherus 1923; Roivainen
1934; Robinson 1975; Mahu 1981; Matteri 1985b; Deguchi 1991a. Dist. IV. Choapa; V.
Petorca, Quillota, San Antonio, Valparaíso; RM. Cordillera, Santiago; VII. Talca; VIII.
Concepción, Ñuble; XII. Última Esperanza; Juan Fernández Is.
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Mosses of Chile 7
Amblystegium tenellum Cardot & Broth. = Cratoneuropsis relaxa subsp. minor fide Ochyra et al. 2008.
Amblystegium uncinatum De Not. var. symmetricum (Renauld & Cardot) Dusén ≡ Sanionia symmetrica.
Amphidium cyathicarpum (Mont.) Broth. = Amphidium tortuosum fide Robinson 1975.
Amphidium lapponicum (Hedw.) Schimp.
Müller in Blockeel et al. 2000; Frahm et al. 2000. Dist. VII. Talca; IX. Malleco; Juan
Fernández Is.
Amphidium tortuosum (Hornsch.) Cufod.
Tropicos 2009; Frahm et al. 2000; He 1998; Müller & Pereira 2006; Robinson 1975 (as
Amphidium cyathicarpum, Hässel et al. 1984; Espinosa 1941; Costes 1921; Cardot 1908;
Thériot 1917, 1921; Brotherus 1924b; Herzog & Hosseus 1938; Herzog & Schwabe 1939;
Bartram 1957; Matteri 1985b; Deguchi 1991b). Dist. V. San Antonio, Valparaíso; RM.
Cordillera; VI. Colchagua; VII. Cauquenes, Talca; VIII. Arauco, Biobío, Concepción, Ñuble;
IX. Cautín; XIV. Valdivia/Ranco; X. Llanquihue, Palena; XI. Aisén; XII. Magallanes, Última
Anacolia aurea Dusén, nom. nud., Dusén 1903b.
Anacolia subsessilis (Taylor) Broth. = Anacolia laevisphaera fide Herzog 1960.
Anacolia subsessilis var. brevifolia Broth. = Anacolia laevisphaera fide Flower in Grout 1935.
Ancistrodes genuflexa (Müll. Hal.) Crosby
Larraín 2007b; Larraín 2005; Ireland et al. 2007; Villagran et al. 2003; He 1998; Müller &
Pereira 2006; Kuc 1996; Deguchi 1991b (as Duseniella genuflexa, Cardot 1908; Herzog 1922,
1954, 1960; Reimers 1926; Thériot 1934b; Herzog & Schwabe 1939; Seki 1974; as Meteorium
genuflexum, Mitten 1869; as Bryoduseniella genuflexa, Robinson 1974). Dist. VII. Cauquenes;
VIII. Arauco, Biobío, Concepción, Ñuble; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X.
Chiloé, Llanquihue, Osorno; XI. Aisén.
Andreaea acutifolia Hook.f. & Wilson
Larraín 2005; Mitten 1869; Cardot 1908; Cardot & Brotherus 1923; Roivainen 1937; Seki
1974; Deguchi 1991a; Matteri & Schiavone 2002. Dist. VIII. Biobío; IX. Malleco; X.
Llanquihue, Osorno; XI. Aisén; XII. Antártica Chilena, Tierra del Fuego.
Andreaea alpina Hedw.
Ireland et al. 2007; Mitten 1869; Roivainen 1937; Seki 1974; Robinson 1975; Matteri &
Schiavone 2002 (as Andreaea squamata, Dusén 1903a; Cardot 1908; as Andreaea brevifolia,
Dusén 1903a; as Andreaea parallela var. brevifolia, Cardot 1908; as Andreaea loricata, Dusén
1903a, 1903b; Cardot & Brotherus 1923; as Andreaea pseudoalpina, Dusén 1903a, Cardot
1908, Deguchi 1991a). Dist. VIII. Ñuble; X. Llanquihue; XI. Aisén; XII. Antártica Chilena,
Magallanes, Tierra del Fuego; Juan Fernández Is.
Andreaea appendiculata Schimp.
Mitten 1869; Dusén 1903b; Cardot 1908; Matteri & Schiavone 2002. Dist. XII. Antártica
Chilena.
Andreaea brevifolia Dusén = Andreaea alpina fide Roivainen 1937.
Andreaea flabellata Müll. Hal.
NY 2009; Greene 1968; Matteri & Schiavone 2002 (as Andreaea grimmioides, Dusén 1903a;
Cardot & Brotherus 1923; Roivainen 1937; as Andreaea pumila, Herzog 1954). Dist. XI.
Aisén; XII. Antártica Chilena, Tierra del Fuego.
Andreaea flexuosa R.Br. bis
NY 2009. Dist. XII. Antártica Chilena (Buck 46204, NY).
Andreaea fuegiana (Cardot) S.W. Greene
Greene 1968; Matteri & Schiavone 2002 (as Neoloma fuegianum, Donat 1936; as Neuroloma
fuegianum, Cardot & Brotherus 1923; Roivainen 1937). Dist. XI. Aisén; XII. Tierra del Fuego.
Andreaea grimmioides Dusén = Andreaea flabellata fide Greene 1968.
Andreaea latinervis E.B. Bartram
Roivainen 1937; Matteri & Schiavone 2002. Dist. XII. Tierra del Fuego.
Andreaea laxifolia Hook.f. & Wilson

8 Müller
Mitten 1869; Dusén 1903b; Cardot 1908; Matteri & Schiavone 2002. Dist. XII. Antártica
Chilena.
Andreaea laxifolia var. minor Hook.f. & Taylor
Cardot 1908; Matteri & Schiavone 2002. Dist. XII. Antártica Chilena
Andreaea leiophylla Cardot ex G. Roth
Cardot & Brotherus 1923; Roivainen 1937; Matteri & Schiavone 2002. Dist. XII. Tierra del
Fuego.
Andreaea loricata Dusén = Andreaea alpina fide Roivainen 1937.
Andreaea marginata Hook.f. & Wilson
Mitten 1869; Dusén 1903a, 1903b; Cardot 1908; Herzog 1954; Matteri & Schiavone 2002.
Dist. XI. Aisén; XII. Antártica Chilena, Magallanes.
Andreaea mutabilis Hook.f. & Wilson
NY 2009; He 1998; Dusén 1903b; Roivainen 1937; Matteri & Schiavone 2002. Dist. IX.
Cautín; X. Osorno; XII. Antártica Chilena, Magallanes, Tierra del Fuego.
Andreaea nitida Hook.f & Wilson
Deguchi 1991a; Matteri & Schiavone 2002 (as Cochlearidium aymaranum, Ochyra & Mahu
1990). Dist. XV. Parinacota; XII. Magallanes.
Andreaea pachyphylla (Müll. Hal.) Broth.
Dusén 1903a; Roivainen 1937; Herzog 1954; Schultze-Motel 1970; Matteri & Schiavone 2002
(as Andreaea pachyphylla var. acutifolia, Dusén 1903a; Cardot 1908). Dist. XI. Aisén; XII.
Magallanes, Tierra del Fuego.
Andreaea pachyphylla var. acutifolia Dusén = Andreaea pachyphylla fide Schultze-Motel 1970.
Andreaea parallela Müll. Hal. var. brevifolia (Dusén) Cardot = Andreaea alpina fide Roivainen 1937.
Andreaea patagonica Dusén
Dusén 1903a; Cardot 1908; Cardot & Brotherus 1923; Matteri & Schiavone 2002. Dist. XI.
Aisén; XII. Tierra del Fuego.
Andreaea pilifera Herzog ex Thér.
Herzog 1937. Dist. XI. Aisén.
Andreaea pseudoalpina Müll. Hal. = Andreaea alpina fide Matteri 2003a.
Andreaea pseudomutabilis Dusén
NY 2009; Dusén 1903a; Deguchi 1991a; Matteri & Schiavone 2002. Dist. XII. Antártica
Chilena, Magallanes.
Andreaea pseudosubulata Müll. Hal. = Andreaea subulata fide Schultze-Motel 1970.
Andreaea pulvinatula Dusén, nom. nud., Dusén 1903b.
Andreaea pumila Cardot = Andreaea flabellata fide Greene 1968.
Andreaea purpurascens (Dusén) G. Roth
Matteri & Schiavone 2002 (as Andreaea remotifolia var. purpurascens, Dusén 1903a). Dist.
XII. Magallanes.
Andreaea pycnotyla Cardot = Andreaea regularis fide Ochyra 1998.
Andreaea pygmaea Cardot = Andreaea regularis fide Greene 1968.
Andreaea regularis Müll. Hal.
Goffinet et al. 2006; Buck 2002; Seki 1974; Deguchi 1991a; Matteri & Schiavone 2002 (as
Andreaea pygmaea, Cardot & Brotherus 1923; as Andreaea verruculosa, Roivainen 1937;
Bartram 1952). Dist. IX. Malleco; X. Osorno; XI. Aisén; XII. Antártica Chilena, Magallanes,
Tierra del Fuego.
Andreaea remotifolia Dusén
Tropicos 2009; Dusén 1903a; Matteri & Schiavone 2002. Dist. XII. Antártica Chilena,
Magallanes.
Andreaea remotifolia var. purpurascens Dusén ≡ Andreaea purpurascens.
Andreaea rupestris Hedw.
Hässel et al. 1984; Ireland et al. 2007; He 1998; Bartram 1952; Matteri & Schiavone 2002.
Dist. VIII. Arauco, Biobío, Ñuble; IX. Cautín; X. Osorno; XII. Magallanes.
Andreaea squarrosa Mitt.
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Mosses of Chile 9
Donat 1936; Herzog 1954. Dist. XI. Aisén.

Andreaea squamata Müll. Hal. = Andreaea alpina fide Robinson 1975.
Andreaea subulata Harv.
Müller 2009; Ireland et al. 2007; Mitten 1869; Dusén 1903a; Roivainen 1937; Schultze-Motel
1970; Seki 1974; Deguchi 1991a; Matteri & Schiavone 2002 (as Andreaea subulata var.
perichaetialis, Dusén 1903b; Cardot 1908; as Andreaea subulata var. rigida, Dusén 1903b; as
Andreaea pseudosubulata, Cardot 1908). Dist. VII. Talca; VIII. Biobío, Ñuble; IX. Malleco;
X. Palena; XI. Aisén; XII. Antártica Chilena, Magallanes, Tierra del Fuego.
Andreaea subulata var. perichaetialis (Hook.f. & Wilson) Müll. Hal. = Andreaea subulata fide Schultze-Motel 1970.
Andreaea subulata var. rigida Harv. & Müll. Hal. = Andreaea subulata fide Dixon 1929.
Andreaea vaginalis Herzog
Donat 1936; Seki 1974. Dist. XI. Aisén.
Andreaea vermicularis Dusén, nom. nud., Dusén 1903b.
Andreaea verruculosa Cardot = Andreaea regularis fide Greene 1968.
Andreaea wilsonii Hook.f. ≡ Acroschisma wilsonii.
Anisothecium campylophyllum (Taylor) Mitt. ≡ Dicranella campylophylla.
Anisothecium cardotii (R.Br. bis) Ochyra = Dicranella campylophylla fide Ochyra et al. 2008.
Anisothecium elegans (Duby) Thér.
Thériot 1934b; Herzog & Hosseus 1938 (as Aongstroemia elegans, Herzog 1922). Dist. XIV.
Valdivia/Ranco; X. Llanquihue.
Anisothecium fontanum (Herzog) Ochyra
(as Ditrichum fontanum, Herzog 1954). Dist. X. Osorno.
Anisothecium hookeri (Müll. Hal.) Broth. ≡ Dicranella hookeri.
Anisothecium hookeri var. minus Cardot & Thér.
Thériot 1929b. Dist. XII. Magallanes.
Anisothecium jamesonii Mitt. = Anisothecium vaginatum fide Matteri 2003a.
Anisothecium krausei Lorentz ≡ Symplepharis krausei.
Anisothecium paludella (Besch.) Broth. ≡ Neomeesia paludella.
Anisothecium perpusillum Dusén = Dicranella hookeri fide Newton 1977.
Anisothecium persquarrosum (Dusén) Broth. = Dicranella campylophylla fide Matteri & Schiavone 2002.
Anisothecium pseudorufescens (Cardot & Broth.) Broth.
(as Dicranella pseudorufescens, Cardot & Brotherus 1923). Dist. XII. Magellanes.
Anisothecium pseudorufescens var. leptocladus (Cardot & Broth.) Kühnem.
(as Dicranella pseudorufescens var. leptoclada, Cardot & Brotherus 1923). Dist. XII.
Magallanes.
Anisothecium skottsbergii (Cardot & Broth.) Broth.
Herzog 1954 (as Dicranella skottsbergii, Cardot & Brotherus 1923). Dist. X. Osorno; XII.
Magallanes.
Anisothecium vaginatum (Hook.) Mitt.
Tropicos 2009; He 1998; Thériot 1934b (as Aongstroemia vaginata, Dusén 1905b; as
Anisothecium jamesonii, Mitten 1869; Herzog & Hosseus 1938; Herzog & Schwabe 1939).
Dist. RM. Santiago; VIII. Arauco, Biobío; XIV. Valdivia/Ranco; X. Llanquihue, Osorno.
Anoectangium patagonicum Cardot & Broth. = Hymenostylium recurvirostrum fide Matteri & Schiavone 2002.
Anoectangium tenerrimum Müll. Hal. ≡ Gymnostomum tenerrimum fide Wijk & Margadant 1959.
Anomobryum filiforme (Dicks.) Husn. = Anomobryum julaceum fide Abramov & Abramova 1983.
Anomobryum julaceum (Schrad. ex P. Gaertn., B. Mey. & Scherb.) Schimp.
Tropicos 2009 (as Anomobryum filiforme, Espinosa 1941; He 1998; Brotherus 1924a; Cardot
& Brotherus 1923; Herzog & Hosseus 1938; Herzog & Schwabe 1939; Herzog 1954; as Bryum
filiforme, Seki 1974; Ochi 1980; Deguchi 1991a). Dist. VIII. Arauco, Bíobio, Concepción,
Ñuble; IX. Cautín; X. Llanquihue, Osorno; XI. Aisén.
Anomobryum humillimum (Müll. Hal. ex E. Britton) Broth.
(as Bryum humillimum, Ochi 1990; Deguchi 1991a). Dist. X. Llanquihue.
Anomobryum laxirete Cardot & Broth. = Bryum microimbricatum fide Ochi 1982.
Anomobryum prostratum (Müll. Hal.) Besch.

10 Müller
(as Bryum prostratum, Ochi 1980). Dist. V. Valparaíso.

Aongstroemia elegans (Dusén) Broth. ≡ Anisothecium elegans.
Aongstroemia gayana (Mont.) Müll. Hal.
Tropicos 2009; Villagran et al. 2003; He 1998; Müller & Pereira 2006; Mitten 1869; Dusén
1905b; Thériot 1921; Herzog & Hosseus 1938; Herzog & Schwabe 1939; Herzog 1954;
Deguchi 1991a; Costes 1921. Dist. V. Valparaíso; RM. Santiago; VII. Cauquenes, Linares,
Talca; VIII. Bíobio, Ñuble; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue,
Osorno; XI. Aisén.
Aongstroemia persquarrosa Dusén = Dicranella campylophylla fide Matteri & Schiavone 2002.
Aongstroemia vaginata (Hook.f.) Müll. Hal. ≡ Anisothecium vaginatum.
Apalodium australe (Hook.f. & Wilson) Mitt. = Orthodontium lineare fide Matteri & Schiavone 2002.
Aptychella chilensis Herzog
He 1998; Herzog 1954; Seki 1974. Dist. XIV. Valdivia/Ranco; X. Chiloé; XI. Aisén.
Archidium julaceum Müll. Hal.
He 1998; Thériot 1918, 1924. Dist. V. San Antonio, Valparaíso; IX. Malleco.
Archidium ohioense Schimp. ex Müll. Hal.
He 1998. Dist. V. Valparaíso.
Arctoa fulvella (Dicks.) Bruch & Schimp.
Ochyra & Buck 2003. Dist. XII. Antártica Chilena.
Asteriscium deciduum Hilp. ≡ Didymodon deciduus.
Asteriscium geniculatum (Mont.) Herzog = Didymodon australasiae fide Jiménez et al. 2005.
Asteriscium poeppigianum (Müll. Hal.) Hilp. = Didymodon vinealis fide Sollman 1995.
Astomiopsis pacifica W.R.Buck & Landrum
Barrera 2006; He 1998; Buck & Landrum 1977. Dist. V. San Antonio, Valparaíso; Juan
Fernández Is.
Astomum chilense R.S. Williams = Trichostomum williamsii fide Zander 1993.
Astomum recurvirostre (Müll. Hal.) G. Roth = Weissia diffidentia fide Zander 1993.
Atractylocarpus patagonicus Herzog & Thér.
Herzog & Thériot 1933; Padberg & Frahm 1985. Dist. VIII. Concepción.
Atrichopsis compressa (Hook.f. & Wilson) G. L. Sm.
Hässel et al. 1984; Matteri 1985b; Matteri & Schiavone 2002 (as Psilopilum compressum,
Mitten 1869; Dusén 1903b; Cardot 1908; Cardot & Brotherus 1923; Herzog & Schwabe 1939;
Herzog 1954; as Atrichopsis magellanica, Cardot & Brotherus 1923). Dist. X. Llanquihue; XI.
Aisén; XII. Antártica Chilena, Magallanes, Tierra del Fuego, Última Esperanza.
Atrichopsis magellanica Cardot = Atrichopsis compressa fide Smith 1969.
Atrichum rigidum Lorentz ≡ Hebantia rigida.
Aulacomnium chilense Müll. Hal. = Pyrrhobryum mnioides fide Wijk et al. 1959.
Aulacomnium palustre (Hedw.) Schwägr.
He 1998; Dusén 1903b; Matteri 1985b; Matteri 1988; Matteri & Schiavone 2002. Dist. V.
Valparaíso; IX. Cautín; XII. Tierra del Fuego.
Austrophilibertiella ditrichoidea (Cardot) Ochyra ≡ Ditrichum ditrichoideum.
Austrophilibertiella nitens (Herzog) Ochyra ≡ Ditrichum nitens.
Barbula andersonii (Ångström) A. Jaeger = Syntrichia glacialis fide Müller 2009.
Barbula antarctica Hampe ≡ Syntrichia antarctica.
Barbula arenae Besch. ≡ Hennediella arenae.
Barbula arenicola Dusén = Pseudocrossidium crinitum fide Zander 1993.
Barbula australasiae Hook.f. & Grev. ≡ Didymodon australasiae fide Zander 1978.
Barbula austrogracilis Dusén
Dusén 1906a. Dist. VI. Colchagua.
Barbula berteroana Müll. Hal. = Tortula platyphylla fide Cano & Gallego 2008.
Barbula calycina Schwägr.
(as Tortula calycina, Mitten 1869). Dist. Chile, without exact locality.
Barbula calyculosa (Mitt.) A. Jaeger
Herzog 1954. Dist. X. Osorno.
Barbula cardotii Dusén ≡ Didymodon cardotii.
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Mosses of Chile 11
Barbula chrysopila Müll. Hal. = Syntrichia princeps fide Matteri & Schiavone 2003.
Barbula conotricha Müll. Hal. = Syntrichia glacialis fide Müller 2009.
Barbula conotricha var. fagicola Müll. Hal. = Syntrichia glacialis fide Matteri 2003a.
Barbula convoluta Hedw.
Barbula costesii Thér.
Buck 2002; Thériot 1921. Dist. V. Valparaíso; XII. Antártica Chilena.
Barbula densifolia Hook.f. & Wilson ≡ Hennediella densifolia.
Barbula depressa Sull. = Pseudocrossidium crinitum fide Zander 1993.
Barbula depressa var. denticulata (Dusén) Thér. = Pseudocrossidium crinitum fide Zander 1993.
Barbula flagellaris Schimp. ≡ Syntrichia flagellaris.
Barbula flagellaris var. denticulata Dusén = Pseudocrossidium crinitum fide Zander 1993.
Barbula flagellaris var. gracilis Dusén = Pseudocrossidium crinitum fide Zander 1993.
Barbula flavidopilosa Dusén, nom. nud., Dusén 1903b.
Barbula fuegiana (Mitt.) A. Jaeger = Syntrichia magellanica fide Matteri 2003a.
Barbula fusca Müll. Hal. ≡ Didymodon fuscus.
Barbula fuscinervia (Mitt.) A. Jaeger = Didymodon pruinosus fide Jiménez & Cano 2008.
Barbula fuscoviridis Broth. = Didymodon fuscus fide Jiménez & Cano 2006.
Barbula geniculata (Mont.) Müll. Hal. = Didymodon australasiae fide Jiménez et al. 2005.
Barbula graminicolor Müll. Hal. subsp. subgraminicolor Thér. = Tortula platyphylla fide Cano & Gallego 2008.
Barbula kunzeana Müll. Hal. ≡ Hennediella kunzeana.
Barbula laevigata (Mitt.) A. Jaeger ≡ Didymodon laevigatus.
Barbula lamprocalyx Müll. Hal.
Brotherus 1924c. Dist. Chile, without exact locality.
Barbula lechleri Müll. Hal. = Syntrichia magallanica fide Zander 1993.
Barbula marginatoserrata Dusén, nom. nud. = Syntrichia serrulata fide Müller 2009.
Barbula microruncinata Dusén, nom. nud., Dusén 1903b.
Barbula muralis var. australis Hampe, nom. inval.
Barbula nanophylla Dusén, nom. nud., Dusén 1903b.
Barbula oliviensis Cardot = Pseudocrossidium crinitum fide Müller 2009.
Barbula pachyneura Dusén = Pseudocrossidium leucocalyx fide Wijk et al. 1959.
Barbula patagonica (Müll. Hal.) A. Jaeger ≡ Didymodon patagonicus.
Barbula perangusta Dusén, nom. nud. = Hennediella densifolia fide Müller 2009.
Barbula perfirma Dusén, nom. nud., Roivainen 1934.
Barbula perrubiginosa Dusén = Syntrichia glacialis fide Zander 1993.
Barbula pilifera (Hook.f.) Brid. = Pseudocrossidium crinitum fide Zander 1993.
Barbula pilifera fo. elata Thér.
Thériot 1934b. Dist. RM. Santiago.
Barbula poeppigiana Müll. Hal. = Didymodon australasiae fide Jiménez et al. 2005.
Barbula purpurascens Dusén
Dusén 1906a; Cardot 1908; Herzog & Hosseus 1938; Herzog 1960. Dist. IX. Cautín; XIV.
Valdivia/Ranco; XI. Aisén.
Barbula pusilla (Ångström) Müll. Hal. = Syntrichia magellanica fide Zander 1993.
Barbula pycnophylla Cardot
Dusén 1906a; Bartram 1952. Dist. XI. Coihaique.
Barbula replicata Taylor ≡ Pseudocrossidium replicatum.
Barbula rigidula (Hedw.) Mitt. ≡ Didymodon rigidulus.
Barbula robusta (Hook.f & Grev.) Brid. ≡ Syntrichia robusta.
Barbula ruralis Hedw. ≡ Syntrichia ruralis.
Barbula santessonii E.B. Bartram ≡ Didymodon santessonii.
Barbula santiagensis Broth.
Ireland et al. 2007; He 1998; Dusén 1906b; Thériot 1934b. Dist. IV. Limarí; V. San Antonio,
Valparaíso; RM. Santiago; VIII. Concepción.
Barbula santiagensis var. piligera Broth.
Dusén 1906b. Dist. RM. Santiago; IX. Malleco.
Barbula serrulata (Hook. & Grev.) Brid. ≡ Syntrichia serrulata.
Barbula subgraminicolor Thér., nom. inval. = Tortula platyphylla fide Cano & Gallego 2008.
Barbula unguiculata Hedw.

12 Müller
Müller 2009; Deguchi 1991b; Frey & Schaumann 2002; Schaumann 2005. Dist. IV. Limarí;
RM. Santiago; VII. Talca; XII. Última Esperanza.
Barbula ventanae Müll. Hal. ex Paris, nom. inval., Brotherus 1924c.
Bartramia ambigua Mont. = Bartramia stricta fide Matteri 1985a.
Bartramia aristata Schimp. ex Müll. Hal. = Bartramia ithyphylloides fide Fransén 2004b.
Bartramia australis Sw., hom. illeg.
Bartramia bellolioella B.H. Allen & Ireland
Müller 2009; Allen & Ireland 2007. Dist. VII. Talca; VIII. Arauco, Ñuble.
Bartramia carinata Mitt. = Breutelia plicata fide Matteri 1985a.
Bartramia chilensis Lorentz = Breutelia dumosa fide Griffin 1988.
Bartramia cygnea Mont. ≡ Philonotis cygnea.
Bartramia dumosa (Mitt.) Mitt. ≡ Breutelia dumosa.
Bartramia fernandeziana Cardot = Bartramia ithyphylloides fide Fransén 2004b.
Bartramia fontanoides Gillies & Grev. ≡ Philonotis fontanoides.
Bartramia halleriana var. brachydonta Kab. = Bartramia mossmaniana fide Fransén 2004a.
Bartramia ithyphylla subsp. patens (Brid.) Fransén
Fransén 2004b (as Bartramia patens, Goffinet et al. 2006; Larraín 2007b; Hässel et al. 1984;
Larraín 2007a; Buck 2002; He 1998; Mitten 1869; Dusén 1903b; Cardot 1908; Herzog 1922,
1954, 1960; Cardot & Brotherus 1923; Brotherus 1924b; Roivainen 1934; Bartram 1952; Seki
1974; Robinson 1975; Matteri 1985a, 1985b; Matteri & Schiavone 2002; as Bartramia
leucocolea var. glaucoflava, Cardot & Brotherus 1923). Dist. RM. Chacabuco, Santiago; VI.
Colchagua; VII. Curicó, Linares; VIII. Ñuble; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X.
Chiloé, Osorno; XI. Aisén, Coihaique; XII. Antártica Chilena, Magallanes, Tierra del Fuego,
Última Esperanza; Juan Fernández Is.
Bartramia ithyphylloides Schimp. ex Müll. Hal.
Müller & Pereira 2006; Fransén 2004b; Dusén 1903b; Cardot 1908; Thériot 1934b; Roivainen
1934; Herzog & Schwabe 1939; Herzog 1954; Seki 1974; Matteri 1985a, 1985b; Deguchi
1991a (as Bartramia ithyphylla, Tropicos 2009; Larraín 2007b; Hässel et al. 1984; Buck 2002;
Villagran et al. 2003; Schaumann 2005; He 1998; Matteri & Schiavone 2002; as Bartramia
aristata, Dusén 1903b; Cardot 1908; Thériot 1918, 1934b; Brotherus 1924b; Reimers 1926;
Herzog & Hosseus 1938; Bartram 1957; Robinson 1975; as Bartramia fernandeziana,
Brotherus 1924b; as Bartramia ithyphylla var. arenae, Dusén 1903b; Cardot 1908; as
Bartramia leptophylla, Cardot & Brotherus 1923). Dist. RM. Cordillera; VII. Cauquenes; VIII.
Arauco, Biobío, Concepcion, Ñuble; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X.
Llanquihue, Osorno, Chiloé; XI. Aisén; XII. Antártica Chilena, Magallanes, Última Esperanza;
Juan Fernández Is.
Bartramia leptophylla Cardot & Broth. = Bartramia ithyphylloides fide Fransén 2004b.
Bartramia leucocolea Cardot var. glaucoflava Broth. & Cardot = Bartramia ithyphylla subsp. patens fide Fransén
2004b.
Bartramia magellanica Ångström, hom. illeg. = Bartramia mossmaniana fide Fransén 2004a.
Bartramia mossmaniana Müll. Hal.
Tropicos 2009; Goffinet et al. 2006; Fransén 2004 (as Bartramia halleriana, Tropicos 2009;
Hässel et al. 1984; Matcham et al. 2007; Larraín 2005; Buck 2002; Villagran et al. 2003;
Schaumann 2005; He 1998; Cardot 1908; Herzog 1922; Cardot & Brotherus 1923; Thériot
1935; Roivainen 1934; Herzog & Hosseus 1938; as Bartramia halleriana, Mitten 1869;
Bartram 1952; Seki 1974; Matteri 1985a, 1985b; Deguchi 1991a; Matteri & Schiavone 2002;
as Bartramia magellanica, Mitten 1869; Dusén 1903b). Dist. VIII. Arauco, Bíobio, Ñuble; IX.
Cautín, Malleco; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno, Palena; XI. Aisén,
Coihaique; XII. Antártica Chilena, Magallanes, Última Esperanza.
Bartramia patens Brid. ≡ Bartramia ithyphylla subsp. patens.
Bartramia patens var. minor Dusén = Bartramia ithyphylla subsp. patens fide Fransén 2004b.
Bartramia patens var. robusta (Hook.f. & Wilson) Matteri = Bartramia robusta.
Bartramia plicata (Mitt.) Mitt. ≡ Breutelia plicata.
Bartramia potosica Mont.
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Mosses of Chile 13
Ireland et al. 2007. Dist. VIII. Ñuble.

Bartramia pusilla Sull., hom. illeg. = Philonotis scabrifolia fide Wijk et al. 1959.
Bartramia reticulata P. Beauv. = Bartramia ithyphylla subsp. patens fide Fransén 2004b.
Bartramia robusta Hook.f. & Wilson
Cardot 1908 (as Bartramia patens var. robusta, Hässel et al. 1984; Matcham et al. 2007;
Matteri 1985a, 1985b; Matteri & Schiavone 2002; as Exodokidium donati, Herzog 1937; as
Breutelia robusta, Dusén 1903b). Dist. XI. Aisén; XII. Antártica Chilena, Magallanes, Tierra
del Fuego, Última Esperanza.
Bartramia rupestris Mitt. = Breutelia plicata fide Matteri 1985a.
Bartramia stricta Brid.
Tropicos 2009; Hässel et al. 1984; He 1998; Matteri 1985a; Deguchi 1991a; Matteri 1985b;
Matteri & Schiavone 2002 (as Bartramia ambigua, Espinosa 1941; Pereira et al. 2006; Müller
& Pereira 2006, Mitten 1869; Thériot 1917; Brotherus 1924a; Reimers 1926; Roivainen 1934;
Herzog & Hosseus 1938; Herzog & Schwabe 1939; Herzog 1954, 1960; Bartram 1952; Seki
1974; Costes 1921). Dist. IV. Choapa, Limarí; V. Aconcagua, Petorca, Quillota, San Antonio,
Valparaíso; RM. Chacabuco, Cordillera, Maipo, Melipilla, Santiago, Talagante; VI. Cachapoal,
Cardenal Caro, Colchagua; VII. Cauquenes, Curicó, Linares, Talca; VIII. Arauco, Biobío,
Concepción, Ñuble; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Llanquihue, Osorno; XI.
Aisén; XII. Magallanes, Última Esperanza; Juan Fernández Is.
Bartramia vagans Hook.f. & Wilson ≡ Philonotis vagans.
Bartramidula cygnea (Mont.) Paris ≡ Philonotis cygnea.
Bartramidula exigua (Sull.) A. Jaeger = Philonotis scabrifolia fide Wijk et al. 1967.
Blindia antarctica Müll. Hal. ≡ Hymenoloma antarcticum.
Blindia arcuata Mitt. = Blindia magellanica fide Dixon 1914.
Blindia auriculata Müll. Hal. = Holodontium strictum fide Ochyra 1993.
Blindia brevipes Müll. Hal. = Hymenoloma antarcticum fide Ochyra et al. 2008.
Blindia churuccana Besch. = Kiaeria pumila fide Ochyra 1993.
Blindia contecta (Hook. f. & Wilson) Müll. Hal.
Dusén 1903b; Cardot 1908. Dist. XI. Aisén; XII. Magallanes.
Blindia curviseta Mitt. = Blindia magellanica fide Bartlett & Vitt 1986.
Blindia globularis Dusén = Blindia magellanica fide Dixon 1914.
Blindia humilis Müll. Hal. = Kiaeria pumila fide Ochyra 1993.
Blindia inundata (Cardot) Cardot
NY 2009; Roivainen 1937; Herzog 1960; Matteri & Schiavone 2002; Frahm 2001a (as B.
magellanica var. inundata, Hässel et al. 1984; Herzog & Schwabe 1939; Matteri 1985b). Dist.
XIV. Valdivia/Ranco; XI. Aisén; XII. Antártica Chilena, Magallanes, Tierra del Fuego, Última
Esperanza.
Blindia leptotrichocarpa Müll. Hal. = Ditrichum hookeri fide Cardot in Dixon 1914.
Blindia lygodipoda Müll. Hal. = Blindia magellanica fide Bartlett & Vitt 1986.
Blindia magellanica Schimp. var. magellanica
Tropicos 2009; Müller 2009; Hässel et al. 1984; Buck 2002; Ireland & Bellolio 2002; He 1998;
Dusén 1903b; Cardot 1908; Thériot 1921; Cardot & Brotherus 1923; Roivainen 1937; Herzog
& Hosseus 1938; Herzog & Schwabe 1939; Seki 1974; Matteri 1985b; Deguchi 1991b; Matteri
& Schiavone 2002 (as Blindia arcuata, Mitten 1869; Dusén 1903b; Cardot 1908; as Blindia
curviseta, Hässel et al. 1984; Mitten 1869; Dusén 1903b; Cardot 1908; Cardot & Brotherus
1923; Roivainen 1934, 1937; Herzog & Schwabe 1939; Matteri 1985b; as Blindia globularis,
Dusén 1903b, 1905b; Cardot 1908; Blindia pseudolygodipoda, Hässel et al. 1984; Cardot &
Brotherus 1923; Matteri 1985b). Dist. VII. Talca; VIII. Arauco, Biobío, Concepción, Ñuble;
IX. Cautín; XIV. Valdivia/Ranco; X. Llanquihue, Osorno, Palena; XI. Aisén, Coihaique; XII.
Antártica Chilena, Magallanes, Tierra del Fuego, Última Esperanza; Easter I.
Blindia magellanica var. inundata (Cardot) Herzog ≡ Blindia inundata.
Blindia praticola Cardot & Broth. = Blindia magellanica fide Bartlett & Vitt 1986.
Blindia pseudolygodipoda Cardot = Blindia magellanica fide Bartlett & Vitt 1986.
Blindia pseudorobusta Dusén = Blindia robusta fide Matteri 1985b.
Blindia robusta Hampe

14 Müller
NY 2009; Hässel et al. 1984; Matteri 1985b; Matteri & Schiavone 2002 (as Blindia
pseudorobusta, Dusén 1903b, 1905b; Roivainen 1937; as Blindia tenuifolia, Mitten 1869;
Dusén 1905b; Cardot 1908; Cardot & Brotherus 1923; Roivainen 1937; Herzog & Hosseus
1938; Herzog 1954). Dist. XIV. Valdivia/Ranco; X. Chiloé; XI. Aisén; XII. Antártica Chilena,
Magallanes, Tierra del Fuego, Última Esperanza.
Blindia savatieriana Müll. Hal. = Kiaeria pumila fide Ochyra 1993.
Blindia skottsbergii Cardot fo. robusta Cardot = Chorisodontium aciphyllum fide Ochyra et al. 2008.
Blindia subinclinata Müll. Hal. = Hymenoloma crispulum fide Ochyra 1998.
Blindia sulphurea Dusén, nom. nud. = Hymenoloma crispulum fide Cardot 1908.
Blindia tenuifolia (Hook.f. & Wilson) Mitt. = Blindia robusta fide Matteri 1985b.
Blindia torrentium Cardot & Broth.
Müller 2009; Deguchi 1991b; Matteri & Schiavone 2002. Dist. XIV. Valdivia/Ranco; XII.
Magallanes.
Brachymenium acuminatum Harv.
Ireland et al. 2007; He 1998; Ochi 1980, 1982 (as Brachymenium meyenianum var. ellipticum,
Thériot 1928; as Bryum ellipticum, Mitten 1869). Dist. IV. Choapa; V. San Antonio; RM.
Santiago; VI. Colchagua; VII. Curicó, Linares; VIII. Arauco, Biobío, Concepción, Ñuble.
Brachymenium chilense Ochi & Mahu
Ochi & Mahu 1988. Dist. V. San Antonio; RM. Cordillera; VI. Cachapoal.
Brachymenium ellipticum A. Jaeger = Brachymenium acuminatum fide Ochi 1980.
Brachymenium exile (Dozy & Molk.) Bosch & Sande Lac.
Ireland et al. 2007 (as Brachymenium porteri, Thériot 1925). Dist. VIII. Concepción; IX.
Cautín.
Brachymenium indicum (Dozy & Molk.) Bosch & Sande Lac.
Ireland & Bellolio 2002; He 1998. Dist. RM. Santiago; Easter I.
Brachymenium magellanicum (Sull.) Paris
Dusén 1903b; Ochi 1982; Matteri & Schiavone 2002 (as Bryum magellanicum, Mitten 1869; as
Peromnion magellanicum, Sullivant 1850). Dist. XII. Antártica Chilena, Magallanes, Tierra
del Fuego.
Brachymenium meyenianum (Hampe) A. Jaeger
Herzog & Schwabe 1939; Herzog 1954 (as Bryum meyenianum, Mitten 1869). Dist. V.
Valparaíso; VIII. Ñuble.
Brachymenium meyenianum var. ellipticum (Mitt.) Thér. = Brachymenium acuminatum fide Ochi 1980.
Brachymenium porteri Thér. = Brachymenium exile fide Ochi 1980, 1982.
Brachymenium robertii Broth.
Tropicos 2009; He 1998; Ochi 1982. Dist. RM. Santiago; VI. Cachapoal; VIII. Concepción.
Brachysteleum chimborazense (Spruce ex Mitt.) Thér. var. obtusifolium Thér. ≡ Ptychomitrium hieronymi var.
obtusifolium fide Cao et al. 2006.
Brachysteleum deltorii Thér. ≡ Ptychomitrium deltorii.
Brachysteleum fernandezianum (Mitt.) Thér. ≡ Ptychomitrium fernandesianum.
Brachytheciastrum collinum (Schleich. ex Müll. Hal.) Ignatov & Huttunen
(as Brachythecium collinum, Müller 2009). Dist. XI. General Carrera.
Brachytheciastrum microcollinum (E.B. Bartram) Ignatov & Huttunen
(as Brachythecium microcollinum, Larraín 2007a; Roivainen 1934; Bartram 1946; McFarland
1988). Dist. VIII. Ñuble; X. Chiloé; XII. Tierra del Fuego.
Brachytheciastrum paradoxum (Hook. f. & Wilson) Ignatov & Huttunen
(as Brachythecuim paradoxum, Tropicos 2009; Müller 2009; Hässel et al. 1984; Buck 2002;
He 1998; Matteri & Schiavone 2002; Cardot 1901; Dusén 1903b; Cardot 1908; Cardot &
Brotherus 1923; Roivainen 1934; Herzog 1954; Seki 1974; Matteri 1985b; McFarland 1988;
Deguchi 1991a; as Hypnum paradoxum, Mitten 1869). Dist. RM. Cordillera; VII. Talca; VIII.
Arauco, Biobío, Ñuble; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Chiloé; XI. Aisén; XII.
Antártica Chilena, Magallanes, Última Esperanza.
Brachythecium albicans (Hedw.) Schimp.
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Mosses of Chile 15
Larraín 2007b; Larraín 2005; Buck 2002; Villagran et al. 2003; McFarland 1988; Matteri &
Schiavone 2002 (as Brachythecium morenoi, Dusén 1903b; Cardot 1908; as Brachythecium
sericeo-virens, Cardot 1908; Seki 1974; Deguchi 1991a; as Brachythecium spurio-albicans,
Dusén 1903b). Dist. RM. Santiago; VIII. Ñuble; X. Chiloé, Llanquihue, Osorno; XI. Aisén;
Brachythecium austroglareosum (Müll. Hal.) Paris
Ochyra et al. 2008; Hässel et al. 1984; Larraín 2007a; Buck 2002; McFarland 1988; Matteri &
Schiavone 2002 (as Brachythecium georgicoglareosum, Cardot 1908; Bartram 1952; as
Brachythecium patagonicum, Hässel et al. 1984; Cardot & Brotherus 1923; Matteri 1985b).
Dist. X. Chiloé; XI. Aisén, Coihaique; XII. Antártica Chilena, Magallanes, Última Esperanza.
Brachythecium austrosalebrosum (Müll. Hal.) Paris
Hässel et al. 1984; He 1998; Seki 1974; Matteri 1985b; McFarland 1988; Matteri & Schiavone
2002. Dist. XI. Aisén; XII. Antártica Chilena, Magallanes, Última Esperanza.
Brachythecium cavifolium Herzog
Frey & Schaumann 2002; Schaumann 2005. Dist. IX. Cautín.
Brachythecium collinum (Schleich. ex Müll. Hal.) Schimp. ≡ Brachytheciastrum collinum.
Brachythecium conostomum (Taylor) A. Jaeger
Ireland et al. 2007; He 1998; Reimers 1926; Herzog & Hosseus 1938; McFarland 1988 (as
Brachythecium grandirete, Brotherus 1925; as Brachythecium pumilum, Dusén 1905a). Dist.
VIII. Arauco, Ñuble; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue.
Brachythecium cuspidarioides Dusén = Brachythecium subplicatum fide McFarland 1988.
Brachythecium filirepens Dusén ≡ Sciuro-hypnum filirepens.
Brachythecium flagellare (Hedw.) Jenn., nom. inval. = Sciuro-hypnum plumosum fide McFarland 1988.
Brachythecium fuegianum Broth. ≡ Sciuro-hypnum fuegianum.
Brachythecium georgicoglareosum (Müll. Hal.) Paris = Brachythecium austrosalebrosum fide Ochyra et al. 2008.
Brachythecium glaciale Schimp. ≡ Sciuro-hypnum glaciale.
Brachythecium grandirete Müll. Hal. = Brachythecium conostomum fide McFarland 1988.
Brachythecium longidecurrens E.B. Bartram = Sciuro-hypnum filirepens fide McFarland 1988.
Brachythecium longidens (Müll. Hal.) Paris
Matteri & Schiavone 2002 (as Stereophyllum fuegianum, Dusén 1903b; Cardot 1908). Dist.
XII. Antártica Chilena.
Brachythecium macrogynum Cardot = Brachythecium subpilosum fide McFarland 1988.
Brachythecium majusculum Dusén, nom. nud. = Sciuro-hypnum fuegianum fide Ochyra et al. 2008.
Brachythecium majusculum M.E. Newton = Sciuro-hypnum fuegianum fide Ochyra et al. 2008.
Brachythecium microcollinum E.B. Bartram ≡ Brachytheciastrum microcollinum.
Brachythecium morenoi Müll. Hal. = Brachythecium albicans fide McFarland 1988.
Brachythecium paradoxum (Hook.f. & Wilson) A. Jaeger ≡ Brachytheciastrum paradoxum.
Brachythecium patagonicum Cardot & Broth. = Brachythecium austroglareosum fide McFarland 1988.
Brachythecium plumosum (Hedw.) Schimp. ≡ Sciuro-hypnum plumosum.
Brachythecium praelongum Schimp. ex Müll. Hal.
Buck 2002; Schaumann 2005; Frey & Schaumann 2002; McFarland 1988 (as Brachythecium
trachychaete, Dusén 1905a). Dist. IX. Cautín; XI. Aisén; XII. Antártica Chilena.
Brachythecium pumilum Dusén = Brachythecium conostomum fide McFarland 1988.
Brachythecium rutabulum (Hedw.) Schimp.
NY 2009; Tropicos 2009; Müller 2009; He 1998; Reimers 1926; McFarland 1988; Matteri &
Schiavone 2002 (as Brachythecium rivulare, Thériot 1934b). Dist. VII. Talca; VIII. Biobío,
Ñuble; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Chiloé; XII. Antártica Chilena,
Magallanes.
Brachythecium sericeo-virens (Müll. Hal.) Kindb., nom. obsc. = Brachythecium austroglareosum fide Matteri &
Schiavone 2002.
Brachythecium spegazzinii (Müll. Hal.) Paris ≡ Hypnum spegazzinii.
Brachythecium spuri-albicans Müll. Hal. = Brachythecium albicans fide McFarland 1988.
Brachythecium subpilosum (Hook.f. & Wilson) A. Jaeger
Tropicos 2009; Ochyra et al. 2008; Hässel et al. 1984; Buck 2002; Villagran et al. 2003;
Schaumann 2005; He 1998; Cardot 1908; Matteri 1985b; McFarland 1988; Matteri &

16 Müller
Schiavone 2002 (as Brachythecium macrogynum, Seki 1974; Deguchi 1991a; as Hypnum
subpilosum, Mitten 1869). Dist. VIII. Biobío, Ñuble; X. Chiloé, Llanquihue, Osorno; XI.
Brachythecium subplicatum (Hampe) A. Jaeger
Tropicos 2009; Müller 2009; Goffinet et al. 2006; Hässel et al. 1984; Buck 2002; He 1998;
Cardot 1908; Roivainen 1934; Herzog & Schwabe 1939; Seki 1974; Matteri 1985b; McFarland
1988; Deguchi 1991a; Matteri & Schiavone 2002 (as Brachythecium cuspidarioides, Dusén
1903b, 1905a; as Brachythecium turgens, Dusén 1903b, 1905a; Herzog 1922, 1954). Dist. VII.
Talca; VIII. Biobío, Concepción, Ñuble; XIV. Valdivia/Ranco; X. Llanquihue, Osorno; XI.
Aisén, General Carrera; XII. Antártica Chilena, Magallanes, Tierra del Fuego, Última
Esperanza.
Brachythecium subtilicaule Dusén, nom. nud. = Sciuro-hypnum filirepens fide McFarland 1988.
Brachythecium trachychaete Dusén = Brachythecium praelongum fide McFarland 1988.
Brachythecium turgens Dusén = Brachythecium subplicatum fide McFarland 1988.
Braunia chilensis Broth., nom. nud. = Hedwigidium chilense, nom. nud., fide Fleischer 1920.
Braunia spec.
De Luna & Biasuso 2001. Dist. Chile, without exact locality.
Breutelia affinis (Hook.) Mitt.
Griffin 1986. Dist. X. Chiloé.
Breutelia angustiretis E.B. Bartram
Hässel et al. 1984; Bartram 1946; Seki 1974; Matteri 1973c, 1985a, 1985b; Matteri &
Schiavone 2002. Dist. XI. Aisén; XII. Magallanes, Tierra del Fuego, Última Esperanza.
Breutelia aureola (Besch. ex Müll. Hal.) Besch.
NY 2009; Hässel et al. 1984; He 1998; Dusén 1903b; Cardot 1908; Herzog 1954; Matteri
1973c, 1985a, 1985b; Griffin 1988; Deguchi 1991a; Matteri & Schiavone 2002. Dist. IX.
Cautín; XIV. Valdivia/Ranco; X. Palena; XI. Aisén; XII. Antártica Chilena, Magallanes,
Última Esperanza.
Breutelia brachycoma Besch. = Breutelia dumosa fide Matteri 1973c.
Breutelia chilensis (Lorentz) Paris = Breutelia dumosa fide Herzog 1954.
Breutelia chrysura (Müll. Hal.) Broth. = Breutelia integrifolia fide Matteri 1973c.
Breutelia cuspidatissima (Müll. Hal.) Paris = Breutelia polygastrica fide Griffin 1988.
Breutelia dumosa Mitt.
Díaz et al. 2008; Larraín 2007b; Hässel et al. 1984; Larraín 2005; Villagran et al. 2003;
Schaumann 2005; He 1998; Dusén 1903b; Cardot 1908; Cardot & Brotherus 1923; Roivainen
1934; Herzog & Schwabe 1939; Herzog 1954; Seki 1974; Matteri 1973c, 1985a, 1985b;
Griffin 1988; Deguchi 1991a; Matteri & Schiavone 2002 (as Bartramia dumosa, Mitten 1869;
as Breutelia brachycoma, Dusén 1903b; Cardot 1908; Cardot & Brotherus 1923; Herzog &
Schwabe 1939; as Breutelia chilensis, Cardot 1908; Herzog 1922; Roivainen 1934; Herzog &
Schwabe 1939; as Bartramia chilensis, Lorentz 1866). Dist. VIII. Concepción; IX. Cautín;
XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno, Palena; XI. Aisén; XII. Antártica
Breutelia elongata (Hook.f. & Wilson) Mitt.
Hässel et al. 1984; Dusén 1903b; Matteri 1981, 1985b. Dist. XII. Magallanes, Última
Esperanza.
Breutelia glabrifolia Dusén = Breutelia integrifolia fide Matteri 1973c.
Breutelia graminicola (Müll. Hal.) Broth. = Breutelia integrifolia fide Matteri 1973c.
Breutelia hariotiana (Besch. ex Müll. Hal.) Besch. = Breutelia plicata fide Brotherus 1902-1909.
Breutelia integrifolia (Taylor) A. Jaeger
Goffinet et al. 2006; Larraín 2007b; Hässel et al. 1984; Buck 2002; He 1998; Cardot 1908;
Roivainen 1934; Herzog 1954; Seki 1974; Matteri 1973c, 1985a, 1985b; Deguchi 1991a;
Matteri & Schiavone 2002 (as Breutelia chrysura, Thériot 1924, 1925; Herzog 1954; as
Breutelia glabrifolia, Dusén 1903b, 1905a; Cardot 1908; as Breutelia graminicola, Cardot &
Brotherus 1923; as Breutelia skottsbergii, Cardot & Brotherus 1923). Dist. VIII. Biobío,
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Mosses of Chile 17
Ñuble; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno; XI. Aisén,
Coihaique; XII. Antártica Chilena, Magallanes, Tierra del Fuego, Última Esperanza.
Breutelia masafuerae Broth.
Brotherus 1924b; Bartram 1957; Robinson 1975; Griffin 1988; Müller 2002 (as Breutelia
subplicata var. masafuerae, Matteri 1973c). Dist. X. Llanquihue, Palena; Juan Fernández Is.
Breutelia plicata Mitt.
Hässel et al. 1984; Schaumann 2005; He 1998; Mitten 1869; Dusén 1903b; Reimers 1926;
Herzog 1954; Seki 1974; Matteri 1973c, 1985a, 1985b; Griffin 1988; Deguchi 1991a; Matteri
& Schiavone 2002 (as Bartramia plicata, Mitten 1869; as Bartramia carinata, Mitten 1869; as
Breutelia hariotiana, Dusén 1903b; Cardot & Brotherus 1923; as Bartramia rupestris, Mitten
1869; as Breutelia rupestris, Dusén 1903b; Cardot 1908; Cardot & Brotherus 1923; Breutelia
subelongata, Dusén 1903b, 1905a; Herzog & Schwabe 1939). Dist. IX. Cautín; XIV.
Valdivia/Ranco; X. Llanquihue, Osorno; XI. Aisén, Coihaique; XII. Antártica Chilena,
Magallanes, Última Esperanza.
Breutelia robusta (Hook.f. & Wilson) A. Jaeger ≡ Bartramia robusta.
Breutelia rupestris (Mitt.) A. Jaeger = Breutelia plicata fide Matteri 1973c.
Breutelia skottsbergii Cardot = Breutelia integrifolia fide Matteri 1973c.
Breutelia subelongata Broth. = Breutelia plicata fide Matteri 1973c.
Breutelia subplicata Broth.
Tropicos 2009; Larraín 2007b; Hässel et al. 1984; Larraín 2005; Villagran et al. 2003;
Schaumann 2005; He 1998; Müller & Pereira 2006; Thériot 1911, 1934b, 1935; Roivainen
1934; Herzog & Hosseus 1938; Herzog & Schwabe 1939; Bartram 1952; Herzog 1954, 1960;
Seki 1974; Robinson 1975; Matteri 1973c, 1985a, 1985b; Griffin 1988; Deguchi 1991a;
Matteri & Schiavone 2002. Dist. VII. Cauquenes, Talca; VIII. Arauco, Biobío, Concepción,
Ñuble; IX. Cautín; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno, Palena; XI. Aisén;
XII. Magallanes, Última Esperanza; Juan Fernández Is.
Breutelia subplicata var. masafuerae (Broth.) Matteri ≡ Breutelia masafuerae.
Breutelia tomentosa (Sw. ex Brid.) A. Jaeger
Bruchia hampeana Müll. Hal.
(as Sporledera hampeana, Mitten 1969). Dist. „E regno chilensi“, without exact locality. Note:
Rushing (1986) left this species untreated.
Bryodusenia genuflexa (Müll. Hal.) H. Rob. ≡ Ancistrodes genuflexa.
Bryoerythrophyllum berthoanus (Thér.) J.A. Jiménez
Jiménez 2007 (as Didymodon berthoanus, Thériot 1926). Dist. V. Valparaíso; RM. Talagante;
VII. Curicó.
Bryoerythrophyllum bolivianum (Müll. Hal.) R.H. Zander
Cano et al. 2008b. Dist. XV. Parinacota.
Bryoerythrophyllum campylocarpum (Müll. Hal.) H.A. Crum
He 1998; Robinson 1975 (as Didymodon calymperidictyon, Brotherus 1924b; Bartram 1957; as
Didymodon linearis, Brotherus 1924b). Dist. IX. Cautín; Juan Fernández Is.
Bryoerythrophyllum fuscinervium (Mitt.) R.H.Zander = Didymodon pruinosus fide Jiménez & Cano 2008.
Bryoerythrophyllum jamesonii (Taylor) H.A. Crum
Zander 1978. Dist. Juan Fernández Is.
Bryoerythrophyllum recurvirostrum (Hedw.) P.C. Chen
Buck 2002; Matteri & Schiavone 2002; Hässel et al. 1984. Dist. XII. Antártica Chilena, Última
Esperanza.
Bryoporteria chilensis Thér. = Camptodontium cryptodon fide Reimers 1936.
Bryum aerugo Dusén, nom. nud., Dusén 1903b.
Bryum algovicum Sendtn. ex Müll. Hal.
Schaumann 2005; Ochi 1980, 1990; Deguchi 1991a; Matteri & Schiavone 2002 (as Bryum
rigochaete, Dusén 1903b; Cardot & Brotherus 1923). Dist. XII. Magallanes, Última Esperanza.

18 Müller
Bryum alpinum With.

Larraín & Ochyra in Blockeel et al. 2008b. Dist. X. Palena.
Bryum amblyodon Müll. Hal. = Bryum archangelicum fide Holyoak 2004.
Bryum amblyolepis Cardot = Bryum argenteum var. muticum fide Ochyra et al. 2008.
Bryum andicola Hook.f. = Bryum billardieri fide Ochi 1980.
Bryum anisodontaceum Dusén, nom. nud., Dusén 1903b.
Bryum anomobryoides Cardot & Broth. = Bryum dichotomum fide Ochyra 2008.
Bryum archangelicum Bruch & Schimp.
Ochyra et al. 2008 (as Bryum amblyodon, Buck 2004; Ochi 1980, 1990; Deguchi 1991a;
Matteri & Schiavone 2002; as Bryum inclinatum var. magellanicum, Cardot 1900, 1901, 1908;
Herzog 1922). Dist. RM. Santiago; XII. Antártica Chilena, Magellanes, Tierra del Fuego.
Bryum arenae Müll. Hal. = Bryum argenteum fide Brotherus 1904.
Bryum argenteum Hedw.
Larraín 2007b; Hässel et al. 1984; Matcham et al. 2007; Larraín 2005; Ireland et al. 2007;
Pereira et al. 2006; Buck 2002; Villagran et al. 2003; Ireland & Bellolio 2002; Cardot 1908;
Herzog & Hosseus 1938; Herzog & Schwabe 1939; Bartram 1952; Seki 1974; Robinson 1975;
Matteri 1985b; Ochi 1990; Deguchi 1991a; Matteri & Schiavone 2002 (as Bryum arenae,
Dusén 1903b; Cardot 1908; as Bryum candicans, Thériot 1918; as Bryum griseum, Thériot
1917; Costes 1921; as Bryum amblyolepis, Costes 1921). Dist. V. Valparaíso; VII. Curicó,
Talca; VIII. Arauco, Biobío, Concepción, Ñuble; IX. Malleco; XIV. Valdivia/Ranco; X.
Chiloé, Llanquihue, Osorno; XI. Aisén; XII. Antártica Chilena, Magallanes, Última Esperanza;
Juan Fernández Is.; Easter I.
Bryum argenteum var. lanatum (P. Beauv.) Hampe
Brotherus 1924; Espinosa 1941. Dist. Easter I.
Bryum argenteum var. muticum Brid.
Ochyra et al. 2008 (as Bryum ellipsifolium, Seki 1974; Ochi 1982; as Bryum myurella, Dusén
1903b, 1905a). Dist. XI. Aisén; XII. Tierra del Fuego.
Bryum aspillagae Thér. = Bryum dichotomum fide Ochi 1980.
Bryum australe Hampe
Ochi 1982; Deguchi 1991a (as Pohlia magnifica, Hässel et al. 1984; He 1998; Matteri 1985b;
as Webera magnifica, Herzog 1954). Dist. V. San Antonio; X. Llanquihue; XI. Aisén; XII.
Bryum austro-chilense E.B. Bartram = Bryum mucronatum fide Ochi 1982.
Bryum austropolare Cardot = Bryum pseudotriquentrum fide Ochi 1970.
Bryum beyrichianum (Hornsch.) Müll. Hal.
Mitten 1869; Ochi 1981 (as Rhodobryum beyrichianum, Reimers 1926). Dist. XIV.
Valdivia/Ranco.
Bryum bicolor Dicks. = Bryum dichotomum fide Ochi 1980.
Bryum billardieri Schwägr.
Larraín 2007b; Larraín 2005; Pereira et al. 2006; Villagran et al. 2003; He 1998; Müller &
Pereira 2006; Ochi 1980, 1982, 1990; Deguchi 1991a (as Bryum andicola, Tropicos 2009;
Matcham et al. 2007; Mitten 1869; Herzog & Hosseus 1938; Herzog & Schwabe 1939; as
Bryum lechleri, Espinosa 1941; Muñoz & Pisano 1947; Thériot 1915, 1918, 1935; Herzog
1922; Brotherus 1924a, 1924b; Reimers 1926; Herzog & Hosseus 1938; Herzog & Schwabe
1939; Robinson 1975; as Bryum valdiviae, Lorentz 1864). Dist. IV. Limarí; V. Aconcagua;
RM. Santiago; VII. Cauquenes, Linares, Talca; VIII. Arauco, Biobío, Concepción, Ñuble; IX.
Cautín, Malleco; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno; Juan Fernández Is.
Bryum brachychaete Cardot & Broth. = Bryum pseudotriquetrum fide Ochi 1980.
Bryum brachycarpum E.B. Bartram = Bryum pseudotriquetrum fide Ochi 1980.
Bryum brevigemmatum Dusén, nom. nud. = Bryum clavatum fide Ochi 1980.
Bryum caespiticium Hedw.
Buck 2002; Seki 1974; Matteri & Schiavone 2002. Dist. XI. Aisén. XII. Antártica Chilena.
Bryum campoanum Thér.
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Mosses of Chile 19
Larraín 2007b; Thériot 1926; Ochi 1982. Dist. IX. Malleco; X. Chiloé.
Bryum campylothecium Taylor
Ochi 1980, 1982 (as Bryum jaffueli, Thériot 1926). Dist. IV. Choapa; V. Valparaíso; IX.
Malleco.
Bryum canariense Brid.
Ochi 1980. Dist. XIV. Valdivia/Ranco.
Bryum candicans Taylor = Bryum argenteum fide Ochi 1980.
Bryum capillare Hedw.
Matcham et al. 2007; Buck 2002; Seki 1974; Ochi 1980, 1982, 1990; Deguchi 1991a (as
Bryum chilense, Herzog 1922; Bryum philippianum, Mitten 1869; Herzog & Hosseus 1938;
Herzog 1954; as Bryum puconense, Herzog & Hosseus 1938). Dist. VIII. Concepción; IX.
Cautín; XIV. Valdivia/Ranco; X. Llanquihue; XI. Aisén; XII. Antártica Chilena, Magellanes.
Bryum capillare subsp. torquescens (Bruch ex De Not.) Kindb. ≡ Bryum torquescens.
Bryum chilense Reichardt = Bryum capillare fide Ochi 1980.
Bryum chiloense Thér. = Bryum clavatum fide Ochi 1980.
Bryum chorizodontum Cardot & Broth.
Barrera 2006; Cardot & Brotherus 1923. Dist. XII. Magallanes.
Bryum chryseum Mitt.
Bartram 1957; Robinson 1975; Ochi 1980, 1982. Dist. V. Valparaíso; Juan Fernández Is.
Bryum clavatum (Schimp.) Müll. Hal.
Buck 2002; Matteri & Schiavone 2002; Mitten 1869; Ochi 1980; Deguchi 1991a (as Bryum
brevigemmatum, Dusén 1903b; as Bryum chiloense, Thériot 1926; as Bryum cochlaerifolium,
Donat 1936, Herzog 1960; as Bryum concavifolium, Thériot 1926; Costes 1921; Herzog &
Schwabe 1939; as Bryum erythrocarpoides, Seki 1974; as Bryum fernandezianum, Barrera
2006; Espinosa 1941; Brotherus 1924b; Bartram 1957; Robinson 1975; as Bryum
microphyllum and Bryum pycnotylum, Cardot & Brotherus 1923; as Bryum reichertii, Herzog
1926). Dist. V. Valparaíso; VII. Linares; VIII. Concepción; X. Chiloé, Llanquihue; XI. Aisén;
XII. Antártica Chilena; Juan Fernández Is.
Bryum cochlaerifolium Cardot & Broth. = Bryum clavatum fide Ochi 1980.
Bryum coelophyllum D.C. Eaton
Matteri & Schiavone 2002; Eaton 1892. Dist. XII. Magallanes.
Bryum concavifolium Thér. = Bryum clavatum fide Ochi 1980.
Bryum coronatum Schwägr.
Mitten 1869; Seki 1974; Ochi 1980. Dist. XI. Aisén.
Bryum corralense Herzog & Thér. = Bryum dichotomum fide Ochi 1980.
Bryum crassinervium Lorentz = Bryum paradoxum fide Ochi 1980.
Bryum delitescens Cardot
Cardot & Brotherus 1923; Matteri & Schiavone 2002. Dist. XII. Magallanes.
Bryum demissum Dusén, nom. nud., Dusén 1903b.
Bryum densifolium Brid.
Bryum dicarpum E.B. Bartram = Bryum pseudotriquetrum fide Ochi 1980.
Bryum dichotomum Hedw.
Hässel et al. 1984; He 1998; Matteri & Schiavone 2002; Ochi 1980, 1990; Mahu 1981; Matteri
1985b; Deguchi 1991a (as Bryum aspillagae, Thériot 1917; Costes 1921; as Bryum bicolor,
Pereira et al. 2006; Seki 1974; as Bryum corralense, Herzog & Hosseus 1938; as Bryum
miserum, Cardot 1908; Herzog & Schwabe 1939; as Bryum urbanskyi, Matteri & Schiavone
2002; Ochi 1982; as Bryum anomobryoides, Cardot & Brotherus 1923; as Bryum laetevirens,
Cardot & Brotherus 1923; Herzog 1954; Ochi 1982). Dist. V. Valparaíso; RM. Santiago; VII.
Talca; VIII. Concepción; XIV. Valdivia/Ranco; X. Llanquihue; XI. Aisén; XII. Magallanes,
Última Esperanza, Tierra del Fuego.
Bryum donatii Thér.
Thériot 1934b; Donat 1936; Ochi 1982. Dist. XI. Aisén.

20 Müller
Bryum elegantulum Lorentz

Brotherus 1924a; Espinosa 1941. Dist. VIII. Concepción.
Bryum ellipsifolium Müll. Hal. = Bryum argenteum var. muticum fide Ochyra et al. 2008.
Bryum ellipticum Mitt. = Brachymenium acuminatum fide Ochi 1980.
Bryum erythrocarpoides Müll. Hal. & Hampe = Bryum clavatum fide Ochi 1980.
Bryum fernandezianum Broth. = Bryum clavatum fide Ochi 1980.
Bryum filiforme Dicks. = Anomobryum julaceum fide Abramov & Abramova 1983.
Bryum flavopallidum Dusén, nom. nud., Dusén 1903b.
Bryum flexisetum Mitt. = Bryum pallescens fide Ochi 1980.
Bryum fuegianum E.B. Bartram = Bryum uliginosum fide Ochi 1982.
Bryum funkii Schwägr.
Buck 2002; Ochi 1982 (as Bryum pertenerum, Thériot 1926). Dist. IX. Malleco; XII. Antártica
Chilena, Tierra del Fuego.
Bryum gayanum Mont.
Larraín 2007b; Buck 2002; Mitten 1869; Ochi 1982. Dist. X. Chiloé; XI. Aisén; XII. Antártica
Chilena.
Bryum gemmatum Müll. Hal.
Dusén 1903b; Matteri & Schiavone 2002. Dist. XII. Tierra del Fuego.
Bryum gilliesii Hook.
Ireland et al. 2007; Mitten 1869; Ochi 1982; Ochi & Mahu 1988. Dist. V. Valparaíso; VIII.
Biobío, Ñuble; XIV. Valdivia/Ranco.
Bryum gracili-torquescens Müll. Hal. in sched., nom. nud., Thériot 1917, 1918; Costes 1921.
Bryum griseum Dusén ex Thér. = Bryum argenteum fide Ochi 1980.
Bryum hamatum Dusén, nom. nud. = Bryum viridescens fide Ochi 1977.
Bryum heterophyllum Cardot & Broth., hom. illeg. = ? Bryum archangelicum fide Ochi 1982.
Bryum hickenii Herzog = Bryum pseudotriquetrum fide Seki 1974.
Bryum humillimum Müll. Hal. ex Britt. ≡ Anomobryum humillimum.
Bryum inclinatum (Brid.) Bland. var. magellanicum Cardot = Bryum archangelicum fide Ochyra et al. 2008.
Bryum jaffueli Thér. = Bryum campylothecium fide Ochi 1980.
Bryum kermesinum Herzog & Thér. = Bryum subapiculatum fide Ochi 1980.
Bryum laetevirens Cardot & Broth. = Bryum dichotomum fide Ochyra et al. 2008.
Bryum laevigatum Hook.f. & Wilson
Hässel et al. 1984; Larraín 2007a; Buck 2002; Matteri & Schiavone 2002; Dusén 1903b;
Cardot 1908; Cardot & Brotherus 1923; Thériot 1935; Herzog & Hosseus 1938; Herzog &
Schwabe 1939; Herzog 1954; Seki 1974; Thériot 1934b; Ochi 1980; Matteri 1985b; Deguchi
1991a (as Bryum submicrolaevigatum, Cardot & Brotherus 1923). Dist. VIII. Biobío; XI.
Aisén; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno; XI. Aisén; XII. Antártica
Bryum lamprochaete Dusén
Dusén 1903b; Cardot & Brotherus 1923; Matteri & Schiavone 2002. Dist. XII. Magallanes.
Bryum lechleri Müll. Hal. = Bryum billardieri fide Ochi 1980.
Bryum leuco-aristatum Dusén, nom. nud., Dusén 1903b.
Bryum liliputanum Dusén, nom. nud., Dusén 1903b.
Bryum lonchochaete Dusén, nom. nud., Dusén 1903b.
Bryum longidens Thér. = Bryum pseudocapillare fide Ochi 1980.
Bryum macrochaete Cardot = Bryum pseudotriquetrum fide Ochi 1980.
Bryum macropelma Müll. Hal. = Bryum pseudotriquetrum fide Ochi 1980.
Bryum macrophyllum Cardot & Broth.
Hässel et al. 1984; Matteri & Schiavone 2002; Matteri 1985b; Ochi 1990; Deguchi 1991a.
Dist. X. Chiloé; XI. Aisén; XII. Última Esperanza.
Bryum macrosporum E.B. Bartram
Bartram 1946; Matteri & Schiavone 2002. Dist. XII. Magallanes, Tierra del Fuego.
Bryum magellanicum (Sull.) Müll. Hal. ≡ Brachymenium magellanicum.
Bryum malacophyllum Cardot & Broth. = Bryum pseudotriquetrum fide Ochi 1980.
Bryum megalothecium Thér. = Bryum orthothecium fide Ochi 1982.
Bryum meyenianum Hampe ≡ Brachymenium meyenianum.
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Mosses of Chile 21
Bryum microimbricatum Ochi

Matteri & Schiavone 2002; Ochi 1982 (as Anomobryum laxirete, Cardot & Brotherus 1923).
Dist. XII. Tierra del Fuego.
Bryum microphyllum Cardot & Broth. = Bryum clavatum fide Ochi 1980.
Bryum minusculum Müll. Hal.
Dusén 1903b; Matteri & Schiavone 2002. Dist. XII. Tierra del Fuego.
Bryum miserum Cardot = ? Bryum dichotomum fide Ochi 1980.
Bryum mucronatum Mitt.
Matteri & Schiavone 2002 (as Bryum obscurum, Cardot & Brotherus 1923; as Bryum austro-
chilense, Roivainen 1934; Bartram 1946; as Bryum roivaineni, Bartram 1946). Dist. VIII.
Ñuble; XII. Magallanes, Tierra del Fuego.
Bryum muehlenbeckii Bruch & Schimp.
Ochi 1982 (as Bryum porteri, Thériot 1926). Dist. V. San Antonio; RM. Cordillera, Santiago.
Bryum myurella Dusén = Bryum argenteum var. muticum fide Ochyra et al. 2008.
Bryum nivale Müll. Hal.
Buck 2002; Mitten 1869; Ochi 1982. Dist. VIII. Biobío; XII. Antártica Chilena.
Bryum obscurum Cardot & Broth., hom. illeg. = Bryum mucronatum fide Ochi 1982.
Bryum orbiculatifolium Cardot & Broth.
NY 2009; Buck 2002; Cardot & Brotherus 1923; Herzog 1954; Ochi 1982, 1990; Deguchi
1991a. Dist. VIII. Biobío; X. Chiloé, Llanquihue; XI. Aisén; XII. Antártica Chilena, Última
Esperanza.
Bryum orthothecium Cardot & Broth.
Larraín 2007b; Barrera 2006; Cardot & Brotherus 1923; Ochi 1982; Ochi & Mahu 1988 (as
Bryum megalothecium, Thériot 1926; Bartram 1943). Dist. II. Antofagasta; VIII. Concepción;
X. Chiloé, Osorno; XII. Magallanes.
Bryum pachytheca Müll. Hal.
Ochi 1982. Dist. V. San Antonio.
Bryum pallens Sw.
Buck 2002; Matteri & Schiavone 2002; Ochi 1982 (as Bryum pauperculum, Bartram 1946).
Dist. XII. Antártica Chilena, Magallanes, Tierra del Fuego.
Bryum pallescens Schleich ex Schwägr.
Hässel et al. 1984; Matteri & Schiavone 2002; Ochi 1982; Matteri 1985b (as Bryum flexisetum,
Bartram 1943). Dist. III. Atacama; XII. Magallanes, Última Esperanza.
Bryum paradoxum Schwägr.
Ochi 1980 (as Bryum crassinervium, Lorentz 1866, Costes 1921). Dist. V. Valparaíso; XIV.
Valdivia/Ranco.
Bryum parvulum Cardot, hom. illeg. = Bryum uliginosum fide Ochi 1982.
Bryum pauperculum E.B. Bartram = Bryum pallens fide Ochi 1982.
Bryum perlimbatum Cardot
Hässel et al. 1984; Villagran et al. 2004; Schaumann 2005; Matteri & Schiavone 2002; Cardot
1908; Herzog 1922, 1954; Cardot & Brotherus 1923; Seki 1974; Ochi 1982; Matteri 1985b (as
Bryum pseudothyridium, Bartram 1946). Dist. IV. Limarí; IX. Cautín; XI. Aisén; XII.
Antártica Chilena, Magallanes, Tierra del Fuego, Última Esperanza.
Bryum pertenerum Thér. = Bryum funkii fide Ochi 1982.
Bryum philippianum Müll. Hal. = Bryum capillare fide Ochi 1980.
Bryum platyphyllum (Schwägr.) Müll. Hal.
Mitten 1869; Ochi 1982 (as Bryum tenuicaule, Mitten 1869; Costes 1921). Dist. V. Valparaíso;
VIII. Biobío.
Bryum pleuricolor Dusén ex Thér. = Bryum dichotomum fide Ochi 1980.
Bryum porteri Thér. = Bryum muehlenbeckii fide Ochi 1982.
Bryum prostratum Müll. Hal. ≡ Anomobryum prostratum.
Bryum pseudocapillare Besch.
Ochi 1980, 1982 (as Bryum longidens, Thériot 1926; Costes 1921). Dist. V. Valparaíso.

22 Müller
Bryum pseudothyridium E.B. Bartram = Bryum perlimbatum fide Ochi 1982.

Bryum pseudotriquetrum (Hedw.) Schwägr.
Larraín 2007b; Hässel et al. 1984; Larraín 2007a; Buck 2002; Matteri & Schiavone 2002;
Bartram 1952; Seki 1974; Ochi 1980, 1990; Matteri 1985b; Deguchi 1991a (as Bryum austro-
polare, Thériot 1934b; as Bryum brachycarpum, Bartram 1946; as Bryum brachychaete,
Cardot & Brotherus 1923; as Bryum dicarpum, Bartram 1946; as Bryum hickenii, Herzog 1926;
as Bryum macrochaete, Herzog 1954; as Bryum macropelma, Mitten 1869; as Bryum
malacophyllum, Herzog 1954; as Bryum steffenii, Herzog 1926; Seki 1974; as Bryum
weberoides, Cardot & Brotherus 1923). Dist. RM. Santiago; X. Chiloé, Llanquihue; XI. Aisén,
Bryum puconense Herzog & Thér. = Bryum capillare fide Ochi 1982.
Bryum pycnotylum Cardot & Broth. = Bryum clavatum fide Ochi 1980.
Bryum reichertii Herzog = Bryum clavatum fide Ochi 1980.
Bryum revolutum Müll. Hal.
Bryum rigochaete Dusén = Bryum algovicum fide Ochi 1980.
Bryum roivaineni E.B. Bartram = Bryum mucronatum fide Ochi 1982.
Bryum rubens Mitt.
Ochi & Mahu 1988. Dist. RM. Santiago.
Bryum rubromarginatum Ochi
He 1998; Ochi 1982, 1990; Deguchi 1991a (as Rhodobryum chilensis, Thériot 1924). Dist.
RM. Santiago.
Bryum sauteri Bruch & Schimp.
Arts 1995. Dist. V. Valparaíso.
Bryum schleicheri Schwägr. fo. heterophyllum Thér. ≡ Bryum turbinatum fo. heterophyllum fide Podpěra 1973.
Bryum steffenii Dusén ex Herzog = Bryum pseudotriquetrum fide Ochi 1980.
Bryum subapiculatum Hampe
Ochi 1980, 1990; Deguchi 1991a (as Bryum kermesinum, Herzog & Schwabe 1939). Dist. X.
Llanquihue; XI. Aisén.
Bryum subclavatum Thér.
Thériot 1926. Dist. IX. Malleco.
Bryum subgracillimum Thér.
Thériot 1926. Dist. VIII. Concepción.
Bryum submicrolaevigatum Cardot & Broth. = Bryum laevigatum fide Ochi 1980.
Bryum sublamprocarpus Cardot & Broth. = Bryum uliginosum fide Ochi 1982.
Bryum tenuicaule Mont. = Bryum platyphyllum fide Ochi 1982.
Bryum tenuirete Dusén, nom. nud., Dusén 1903b; Cardot 1908.
Bryum torquescens Bruch ex De Not.
Larraín 2007b; Mitten 1869; Thériot 1928. Dist. VII. Linares; X. Chiloé.
Bryum turbinatum (Hedw.) Turner
Schaumann 2005; Seki 1974; Ochi 1982 (as Bryum uspallatense, Herzog 1916). Dist. V.
Valparaíso; RM. Santiago; VIII. Ñuble; IX. Cautín; XI. Aisén; XII. Magallanes.
Bryum turbinatum (Hedw.) Turner fo. heterophyllum (Thér.) Podp.
(as Bryum schleicheri fo. heterophyllum, Thériot 1926). Dist. RM. Maipo.
Bryum turbinatum var. patagonicum Broth., nom. nud., Herzog 1922.
Bryum uliginosum (Brid.) Bruch & Schimp.
Buck 2002; Matteri & Schiavone 2002; Ochi 1982 (as Bryum fuegianum, Bartram 1946; as
Bryum parvulum, Cardot & Brotherus 1923; as Bryum sublamprocarpus, Cardot & Brotherus
1923). Dist. XI. Aisén; XII. Antártica Chilena, Tierra del Fuego.
Bryum urbanskyi Broth. = Bryum dichotomum fide Ochyra et al. 2008.
Bryum uspallatense Herzog = Bryum turbinatum fide Ochi 1982.
Bryum valparaisense Thér.
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Mosses of Chile 23
Ireland et al. 2007; Arts et al. 1995; Thériot 1917; Herzog & Hosseus 1938; Ochi 1982; Costes
1921. Dist. V. Valparaíso; RM. Santiago; VIII. Concepción; XIV. Valdivia/Ranco.
Bryum vernicosum Dusén, hom. illeg., Dusén 1903b; Cardot & Brotherus 1923.
Bryum viridescens Welw. & Duby
Ochi 1982 (as Bryum hamatum, Dusén 1903b; Muñoz & Pisano 1947). Dist. IV. Limarí; South
Chile.
Bryum weberoides Cardot & Broth. = Bryum pseudotriquetrum fide Ochi 1980.
Bryum zeballosicum Cardot & Broth.
Ochi 1982. Dist. RM. Santiago.
Bucklandiella bartramii (Roiv.) Roiv.
(as Racomitrium bartramii, Tropicos 2009; He 1998; Matteri & Schiavone 2002; Robinson
1975; Deguchi 1984, 1987; as Bucklandia bartramii, Roivainen 1955b). Dist. IX. Malleco, X.
Llanquihue; Juan Fernández Is.
Bucklandiella crispipila (Taylor) Ochyra & Bednarek-Ochyra
(as Racomitrium crispipilum, Larraín 2005; He 1998; Robinson 1975; Deguchi 1984, 1991a; as
Racomitrium geminatum, Roivainen 1955a; as Racomitrium integripilum, Dusén 1906b;
Cardot 1908; Herzog 1954; as Racomitrium striatipilum, Tropicos 2009; Buck 2002; Hässel et
al. 1984; Espinosa 1941; Dusén 1906b; Cardot 1908; Cardot & Brotherus 1923; Brotherus
1924b; Thériot 1935; Herzog & Hosseus 1938; Roivainen 1955a; Seki 1974; Matteri 1985b; as
Racomitrium substriatipilum, Roivainen 1955a). Dist. VIII. Arauco, Biobío, Ñuble; IX. Cautín,
Malleco; XIV. Valdivia/Ranco; X. Llanquihue, Osorno; XI. Aisén; XII. Antártica Chilena,
Magallanes, Última Esperanza, Tierra del Fuego; Juan Fernández Is.
Bucklandiella crispula (Hook. f. & Wilson) Bednarek-Ochyra & Ochyra
(as Racomitrium crispulum, Hässel et al. 1984; He 1998; Bartram 1952, 1957; Robinson 1975;
Deguchi 1984, 1987; Matteri 1985b; as Racomitrium flavescens, Cardot 1900, 1901; Dusén
1906b; as Racomitrium heterostichoides var. acutifolium, Dusén 1906b; as Racomitrium
skottsbergii, Cardot & Brotherus 1923; as Racomitrium subulifolium, Matteri & Schiavone
2002; Cardot 1908; Cardot & Brotherus 1923; Roivainen 1955a). Dist. VIII. Ñuble; IX.
Cautín, Malleco; XIV. Valdivia/Ranco; X. Llanquihue, Osorno; XI. Aisén; XII. Magallanes,
Tierra del Fuego, Última Esperanza; Juan Fernández Is.
Bucklandiella didyma (Mont.) Ochyra & Bednarek-Ochyra
Ochyra et al. 2008; Matcham et al. 2007 (as Racomitrium didymum, Tropicos 2009; Buck
2002; Larraín 2005; Villagran et al. 2003; He 1998; Matteri & Schiavone 2002; Mitten 1869;
Reimers 1926; Thériot 1935; Herzog & Hosseus 1938; Herzog & Schwabe 1939; Herzog 1954,
1960; Seki 1974; Deguchi 1991a; as Racomitrium nigritum, Mitten 1869; Dusén 1903b; Cardot
1908; Roivainen 1955a; Robinson 1975; as Racomitrium symphyodontum, Espinosa 1941;
Dusén 1903b; Dusén 1906b; Cardot 1908; Cardot & Brotherus 1923; Brotherus 1924b; Herzog
1954; Roivainen 1955a; Seki 1974; as Grimmia depressa var. terrestris, Dusén 1903b; as
Racomitrium convolutum, Mitten 1869; Brotherus 1924b; Herzog & Hosseus 1938; Herzog
1954; as Racomitrium subrupestre, Matteri & Schiavone 2002; Roivainen 1955a). Dist. VII.
Linares, Talca; VIII. Arauco, Biobío, Ñuble; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X.
Chiloé, Llanquihue, Osorno; XI. Aisén, Coihaique, General Carrera; XII. Antártica Chilena,
Magallanes, Tierra del Fuego, Última Esperanza; Juan Fernández Is.
Bucklandiella heterostichoides (Cardot) Ochyra & Bednarek-Ochyra
(as Racomitrium heterostichoides, Hässel et al. 1984; Buck 2002; Matteri & Schiavone 2002;
Dusén 1906b; Roivainen 1955a; Matteri 1985b). Dist. X. Llanquihue; XII. Antártica Chilena,
Bucklandiella lamprocarpa (Müll. Hal.) Ochyra & Bednarek-Ochyra
(as Racomitrium lamprocarpum, Tropicos 2009; He 1998; Müller & Pereira 2006; Matteri &
Schiavone 2002; Cardot 1908; Deguchi 1991a; as Racomitrium limbatum, Bartram 1946; as
Racomitrium looseri, Thériot 1934b; as Racomitrium pachyneuron, Herzog 1954, 1960; Seki

24 Müller
1974; as Racomitrium sublamprocarpum, Dusén 1903b; as Racomitrium subnigritum, Dusén

1903b; Dusén 1906b; Cardot 1908; Cardot & Brotherus 1923; Brotherus 1924b; Bartram 1952;
Herzog 1954; Roivainen 1955a; Bartram 1957; Seki 1974; as Racomitrium pachyneuron fo.
aquaticum, Herzog & Schwabe 1939). Dist. RM. Santiago; VII. Cauquenes, Talca; VIII.
Arauco, Biobío, Ñuble; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue,
Osorno; XI. Aisén; XII. Tierra del Fuego, Última Esperanza; Juan Fernández Is.
Bucklandiella orthotrichacea (Müll. Hal.) Ochyra & Bednarek-Ochyra
(as Racomitrium orthotrichaceum, Buck 2002; He 1998; Matteri & Schiavone 2002; Deguchi
1984, 1991a; as Racomitrium stenocladoides, Roivainen 1955a; as Racomitrium stenocladum,
Dusén 1906b; Cardot 1908; Herzog & Hosseus 1938). Dist. VII. Talca; XIV. Valdivia/Ranco;
X. Chiloé; XI. Aisén; XII. Antártica Chilena, Última Esperanza.
Bucklandiella pachydictyon (Cardot) Ochyra & Bednarek-Ochyra
(as Racomitrium pachydictyon, Hässel et al. 1984; Buck 2002; Matteri & Schiavone 2002;
Cardot 1908; Cardot & Brotherus 1923; Herzog & Hosseus 1938; Roivainen 1955a; Seki 1974;
Matteri 1985b; as Racomitrium symphyodontum var. muticum, Dusén 1906b). Dist. IX. Cautín;
XIV. Valdivia/Ranco; XI. Aisén; XII. Antártica Chilena, Magallanes, Última Esperanza.
Bucklandiella ptychophylla (Mitt.) Ochyra & Bednarek-Ochyra
(as Racomitrium ptychophyllum, Bartram 1952). Dist. XII. Magallanes, Última Esperanza.
Bucklandiella rupestris (Hook. f. & Wilson) Ochyra & Bednarek-Ochyra
(as Racomitrium rupestre, Eaton 1892; He 1998; Matteri & Schiavone 2002; Mitten 1869;
Dusén 1903b; Dusén 1906b; Cardot 1908; Cardot & Brotherus 1923; Thériot 1925; Roivainen
1934, 1955b; Deguchi 1984, 1991a; as Racomitrium stenocladum var. obtusum, Dusén 1906b;
as Racomitrium crispulum var. rupestre, Seki 1974). Dist. VIII. Ñuble; IX. Cautín; XIV.
Valdivia/Ranco; X. Osorno, Palena; XI. Aisén. XII. Antártica Chilena, Magallanes, Tierra del
Fuego, Última Esperanza..
Bucklandiella sudetica (Funck) Ochyra & Bednarek-Ochyra
Ochyra et al. 2008 (as Racomitrium sudeticum, Matteri & Schiavone 2002; as Racomitrium
andreaeoides, Herzog 1954; as Racomitrium austrogeorgicum, Hässel et al. 1984; Roivainen
1955a; Matteri 1985b; as Racomitrium austrogeorgicum var. kranckii, Roivainen 1955a). Dist.
XI. Aisén; XII. Magallanes, Tierra del Fuego, Última Esperanza.
Callicostella pallida (Hornsch.) Ångström
(as Schizomitrium pallidium, Deguchi 1991a). Dist. X. Osorno.
Callicostella scabriuscula (Müll. Hal.) A. Jaeger
(as Hookeria scabriuscula, Mitten 1869). Dist. Chile australis, without exact locality.
Calliergidium austro-stramineum (Müll. Hal.) E.B. Bartram = Warnstorfia fontinaliopsis fide Ochyra & Matteri 2001.
Calliergon auriculatum (Mont.) Margad. ≡ Acrocladium auriculatum.
Calliergon sarmentosum (Wahlenb.) Kindb. ≡ Warnstorfia sarmentosa.
Calliergon stramineum (Dicks. ex Brid.) Kindb. = Straminergon stramineum (Dicks. ex Brid.) Hedenäs
Calliergonella cuspidata (Hedw.) Loeske
Larraín 2007b; Villagran et al. 2003; He 1998 (as Acrocladium cuspidatum, Deguchi 1991a).
Dist. IX. Cautín; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue.
Calyptopogon mnioides (Schwägr.) Broth.
Tropicos 2009; Müller 2009; Larraín 2007b; Hässel et al. 1984; Larraín 2005; Buck 2005b; He
1998; Matteri & Schiavone 2002; Dusén 1906a; Cardot 1908; Thériot 1921; Herzog 1922,
1954, 1960; Reimers 1926; Herzog & Hosseus 1938; Herzog & Schwabe 1939; Bartram 1952;
Seki 1974; Matteri 1985b; Deguchi 1991b (as Calyptopogon crispatulus, Cardot 1908; as
Streptopogon mnioides, Mitten 1869). Dist. V. Valparaíso; RM. Cordillera; VII. Cauquenes;
Chiloé, Llanquihue, Osorno; XI. Aisén, Coihaique; XII. Antártica Chilena, Magallanes, Última
Esperanza.
Calyptopogon crispatulus (Müll. Hal.) Broth. = Calyptopogon mnioides fide Salmon 1903.
Calyptrochaeta apiculata (Hook. f. & Wilson) Vitt var. apiculata
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Mosses of Chile 25
Hässel et al. 1984; Larraín 2007a; Buck 2002 & 2005a; Vitt 1979; Matteri 1985b; Matteri &
Schiavone 2002 (as Eriopus apiculatus, Mitten 1869; Cardot 1908; Bartram 1952; Herzog
1954; Matteri 1975b). Dist. XIV. Valdivia/Ranco; X. Chiloé, Llanquihue; XI. Aisén; XII.
Calyptrochaeta apiculata (Hook.f. & Wilson) Vitt var. spathulata (Matteri) Matteri
(as Eriopus apiculatus var. spathulatus, Matteri 1975b). Dist. VI. Colchagua.
Calyptrochaeta flexicollis (Mitt.) Vitt
Vitt 1979 (as Eriopus flexicollis, Cardot & Brotherus 1923; Matteri 1975b). Dist. XI. Aisén.
Calyptrochaeta grandiretis (Broth.) H. Rob.
Robinson 1975 (as Eriopus grandiretis, Brotherus 1924b; Bartram 1957; Matteri 1975b). Dist.
Juan Fernández Is.
Calyptrochaeta leptoloma (Broth.) H. Rob.
Robinson 1975 (as Eriopus leptoloma, Brotherus 1924b; Matteri 1975b). Dist. Juan Fernández
Is.
Calyptrochaeta mollis (Besch.) Matteri
(as Eriopus bescherellei, Matteri 1975b; as Distichophyllum molle, Costes 1921; Dusén 1903b;
Cardot 1908; Thériot 1917, 1925; Herzog 1954). Dist. V. Valparaíso; VI. Colchagua; IX.
Cautín; XI. Aisén; XII. Antártica Chilena, Magallanes.
Calyptrochaeta odontoloma (Dusén ex Matteri) Matteri
Hässel et al. 1984; Larraín 2005; He 1998; Matteri & Schiavone 2002; Matteri 1985b; Deguchi
1991a (as Eriopus odontoloma, Matteri 1975b). Dist. X. Llanquihue; XI. Aisén; XII.
Camptochaeta arbusculans Broth. ex E.B. Bartram = Lembophyllum orbiculatum fide Robinson 1970.
Camptochaeta gracilis (Hook.f. & Wilson) Paris ≡ Fallaciella gracilis.
Camptochaete orbiculata (Thér.) H. Rob. ≡ Lembophyllum orbiculatum.
Camptodontium brotheri Dusén = Camptodontium cryptodon fide Reimers 1936.
Camptodontium brotheri var. grandirete Dusén
Dusén 1906b; Cardot 1908. Dist. IX. Malleco; XII. Tierra del Fuego.
Camptodontium cryptodon (Mont.) Reim.
Müller 2009; Hässel et al. 1984; Larraín 2005; Ireland et al. 2007; He 1998; Herzog & Hosseus
1938; Seki 1974; Deguchi 1991a; Matteri & Schiavone 2002 (as Dicranoweisia tenella, Dusén
1903b; as Camptodontium brotheri, Dusén 1905a, 1906b; Cardot 1908; Reimers 1926; Herzog
1954, 1960; as Dicranoweisia cryptodon, Roivainen 1934; as Dicranum cryptodon, Mitten
1869; as Bryoporteria chilensis, Thériot 1933). Dist. RM. Cordillera; VII. Talca; VIII. Arauco,
Biobío, Ñuble; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Llanquihue, Osorno; XI. Aisén,
Coihaique; XII. Magallanes, Última Esperanza.
Campyliadelphus polygamus (Schimp.) Kanda ≡ Campylium polygamum.
Campyliadelphus stellatus (Hedw.) Kanda ≡ Campylium stellatum.
Campylium polygamum (Schimp.) C.E.O. Jensen
Tropicos 2009; He 1998; Ochyra & Matteri 2001; Seki 1974; Matteri & Schiavone 2002;
Hedenäs 2003 (as Campyliadelphus polygamus, Hässel et al. 1984; Matteri 1985b). Dist. III.
Copiapó; V. Petorca, San Antonio, Valparaíso; RM. Chacabuco, Santiago; VI. Colchagua;
VIII. Concepción, Ñuble; XIV. Valdivia/Ranco; XI. Aisén; XII. Magallanes, Tierra del Fuego,
Última Esperanza.
Campylium stellatum (Hedw.) C.E.O. Jensen
(as Campyliadelphus stellatus, Matteri 1985b). Dist. XII. Última Esperanza.
Campylopodium euchlorum (Mont.) Matteri
Matteri & Calabrese 1999 (as Dicranella euchlora, Mitten 1869; as Dicranella flexipes, Cardot
& Brotherus 1923; Seki 1974; as Dicranella costata, Brotherus 1924b; Robinson 1975; as
Dicranella aulacocarpa, Mitten 1869). Dist. Chile, provinciae meridionalis. XIV.
Valdivia/Ranco; X. Chiloé; XI. Aisén; Juan Fernández Is.
Campylopodium flaccidum Herzog = Campylopodium medium fide Giese & Frahm 1985.

26 Müller
Campylopodium medium (Duby) Giese & J.-P. Frahm

Ireland et al. 2007; He 1998; Frahm 2005; Giese & Frahm 1985 (as Campylopodium
flaccidum, Herzog 1926; Seki 1974). Dist. VIII. Arauco; IX. Malleco; XIV. Valdivia/Ranco;
X. Llanquihue; XI. Aisén.
Campylopus aberrans Broth. ≡ Campylopus purpureocaulis subsp. aberrans.
Campylopus aberrans var. viridis Broth.
Barrera 2006; Brotherus 1924b. Dist. Juan Fernández Is.
Campylopus acuminatus Mitt.
Larraín 2007b; Larraín 2007a; He 1998; Frahm 2005; Matteri & Schiavone 2002; Mitten 1869;
Dusén 1903b; Frahm 1976; Frahm 1988 (as Campylopus crassissimus, Dusén 1903b; Dusén
1905c; Roivainen 1934; Frahm 1976; as Campylopus fuegianus, Dusén 1905c; Frahm 1976; as
Campylopus sulphureo-nigritus, Dusén 1903b, 1905c; Cardot 1908; Herzog 1960; Frahm
1976; as Campylopus sordido-nigritus, Dusén 1903b). Dist. IX. Malleco; XIV.
Valdivia/Ranco; X. Chiloé, Llanquihue; XI. Aisén; XII. Antártica Chilena, Magallanes, Última
Esperanza.
Campylopus acuminatus Mitt. var. kirkii (Mitt.) J.-P. Frahm
Frahm 2005. Dist. XIV. Valdivia/Ranco.
Campylopus aureonitens (Müll. Hal.) A. Jaeger subsp. recurvifolius (Dusén) J.-P. Frahm
Larraín 2007a; He 1998; Frahm 1988 (as Campylopus recurvifolius, Dusén 1905c; Herzog &
Hosseus 1938; Frahm 1976). Dist. VII. Curicó; VIII. Biobío; IX. Cautín; X. Chiloé,
Llanquihue.
Campylopus blindioides Broth. = Campylopus pyriformis fide Frahm 1988.
Campylopus campoanus Thér. = Campylopus modestus fide Frahm 1988.
Campylopus canescens (Müll. Hal.) Schimp. = Campylopus introflexus fide Frahm 1988.
Campylopus carbonicolus Thér. = Campylopus modestus fide Frahm 1988.
Campylopus chalarobasis Müll. Hal. = Campylopus incrassatus fide Frahm 1988.
Campylopus chilensis De Not.
Tropicos 2009; Larraín 2005; Pereira et al. 2006; Frahm 2005; Müller & Pereira 2006; Mitten
1869; Frahm 1988; Matteri & Schiavone 2002 (as Campylopus flavonigritus, Dusén 1903b,
1905c; Cardot 1908; Cardot & Brotherus 1923; Frahm 1976; as Campylopus patagonicus,
Dusén 1905c; Seki 1974; Frahm 1976; Deguchi 1991a). Dist. VII. Cauquenes, Talca; XIV.
Valdivia/Ranco; X. Llanquihue, Osorno, Palena; XI. Aisén; XII. Antártica Chilena,
Magallanes.
Campylopus clavatus (R. Br.) Wilson
Tropicos 2009; Larraín 2007b; Larraín 2005; Ireland et al. 2007; Buck 2002; He 1998;
Villagran et al. 2003; Ireland & Bellolio 2002; Frahm 2005; Müller & Pereira 2006; Robinson
1975; Frahm 1976; Frahm 1988; Deguchi 1991a (as Campylopus leptodus, Mitten 1869;
Cardot 1908; Thériot 1925, 1935; Bartram 1952, 1957; as Pilopogon leptodus, Dusén 1905c;
Herzog 1922; as Thysanomitrium leptodus, Brotherus 1924b; Herzog & Hosseus 1938; Herzog
& Schwabe 1939; as Campylopus xanthophyllus, Thériot 1918; Frahm 1976; as Campylopus
leptodus fo. elongata-appressa Thér., Thériot 1935). Dist. V. San Antonio, Valparaíso; VII.
Cauquenes, Curicó, Linares; VIII. Arauco, Concepción; IX. Cautín, Malleco; XIV.
Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno, Palena; XI. Aisén; XII. Antártica Chilena,
Magallanes; Juan Fernández Is.; Easter I.
Campylopus crassissimus Besch. = Campylopus acuminatus fide Frahm 1988.
Campylopus dicranodontioides Broth. = Campylopus vesticaulis fide Frahm 1988.
Campylopus fernandezianus E.B. Bartram = Dicranoloma menziesii fide Robinson 1975.
Campylopus fibrobasis Dusén = Campylopus pyriformis fide Frahm 1988.
Campylopus flavissimus (Müll. Hal.) Besch. = Chorisodontium aciphyllum fide Hyvönen 1991.
Campylopus flavonigritus Dusén = Campylopus chilensis fide Frahm 1988.
Campylopus flavo-viridis Dusén = Campylopus incrassatus fide Frahm 1988.
Campylopus flexicaulis Cardot & Thér., nom. nud., Barrera 2006.
Campylopus fragilis (Brid.) Bruch & Schimp.
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Mosses of Chile 27
Ireland et al. 2007. Dist. VIII. Arauco.

Campylopus fuegianus Dusén = Campylopus acuminatus fide Frahm 1988.
Campylopus guaitecae Dusén = Campylopus vesticaulis fide Frahm 1988.
Campylopus hamatus Broth. = Chorisodontium spec. fide Frahm (1999, 2005)
Roivainen 1937; Frahm 1976; Matteri & Schiavone 2002. Dist. XII. Tierra del Fuego.
Campylopus hygrophilus Broth. = Campylopus vesticaulis fide Frahm 1987.
Campylopus incrassatus Müll. Hal.
Tropicos 2009; Larraín 2007b; Larraín 2007a; He 1998; Buck 2002; Müller & Pereira 2006;
Mitten 1869; Dusén 1905c; Thériot 1917; Herzog & Hosseus 1938; Herzog 1960; Frahm 1976;
Frahm 1988; Deguchi 1991a; Costes 1921 (as Campylopus chalarobasis, Thériot 1934b;
Frahm 1976; as Campylopus flavo-viridis, Dusén 1905c; Seki 1974; Frahm 1976). Dist. IV.
Limarí; V. Aconcagua, Petorca, Quillota, San Antonio, Valparaíso; RM. Santiago, Talagante;
VI. Cachapoal; VII. Cauquenes, Curicó, Talca; VIII. Arauco, Biobío, Concepción, Ñuble; IX.
Malleco; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue; XI. Aisén; XII. Antártica Chilena.
Campylopus introflexus (Hedw.) Brid.
Tropicos 2009; Larraín 2007b; Hässel et al. 1984; Larraín 2005; He 1998; Buck 2002 &
2005b; Espinosa 1941; Villagran et al. 2003; Ireland & Bellolio 2002; Frahm 2005; Müller &
Pereira 2006; Matteri & Schiavone 2002; Mitten 1869; Dusén 1903b; Dusén 1905c; Cardot
1908; Brotherus 1924a, 1924b, 1924; Roivainen 1937; Herzog & Hosseus 1938; Herzog &
Schwabe 1939; Bartram 1952; Herzog 1954, 1960; Bartram 1957; Seki 1974; Robinson 1975;
Frahm 1976; Matteri 1985b; Deguchi 1991a (as Campylopus canescens, Cardot 1908; Frahm
1976; as Campylopus truncatus, Brotherus 1924b; Reimers 1926; Thériot 1934b; Herzog &
Hosseus 1938; Frahm 1976). Dist. IV. Choapa, Limarí; V. Petorca, Valparaíso; RM. Maipo;
VII. Cauquenes, Linares, Talca; VIII. Arauco, Biobío, Concepción, Ñuble; IX. Cautín,
Malleco; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno, Palena; XI. Aisén; XII.
Antártica Chilena, Magallanes, Tierra del Fuego, Última Esperanza; Juan Fernández Is.; Easter
I.
Campylopus kunkelii E.B. Bartram = Campylopus pyriformis fide Frahm 1988.
Campylopus laetevirens (Broth.) J.-P. Frahm = Campylopus laxoventralis fide Frahm 1988.
Campylopus laniger Besch. = Chorisodontium aciphyllum fide Hyvönen 1991.
Campylopus laxiretis E.B. Bartram = Campylopus pyriformis fide Frahm 1988.
Campylopus laxoventralis Herzog ex J.-P. Frahm
Frahm 1988; Deguchi 1991a (as Campylopus laetevirens, Frahm 1976; as Thysanomitrium
laetevirens, Barrera 2006; Brotherus 1924a; Espinosa 1941). Dist. VIII. Concepción.
Campylopus leptodus Mont. = Campylopus clavatus fide Sainsbury 1955.
Campylopus macrophyllus (Kunze ex Müll.Hal.) Mitt. = Bryohumbertia filifolia fide Frahm 1999.
Campylopus masafuerae J.-P. Frahm = Campylopus pyriformis fide Frahm 1988.
Campylopus maulensis Broth., nom. nud., Espinosa 1941 (VIII. Concepción).
Campylopus modestus Cardot
He 1998; Cardot 1908; Herzog & Hosseus 1938; Herzog & Schwabe 1939; Frahm 1976;
Frahm 1988 (as Campylopus campoanus, Thériot 1918; Herzog & Hosseus 1938; Herzog &
Schwabe 1939; Frahm 1976; as Campylopus carbonicolus, Thériot 1918; Frahm 1976; as
Campylopus permollis, Thériot 1926, 1934b; Frahm 1976). Dist. VIII. Concepción, Ñuble; IX.
Cautín, Malleco; XIV. Valdivia/Ranco; X. Llanquihue, Osorno.
Campylopus morenoi (Müll. Hal.) Paris
Dusén 1903b; Frahm 1975. Dist. Patagonia australis. Note: The species does’nt belong to
Campylopus, but the generic position is uncertain (Frahm 1999, 2005).
Campylopus occultoviridis Dusén, nom. nud., Dusén 1903b.
Campylopus orthocomus Besch. = Chorisodontium aciphyllum fide Brotherus 1909.
Campylopus paschalis Cardot & Ther., nom. nud., Barrera 2006.
Campylopus patagonicus Dusén, nom. nud., Dusén 1903b.
Campylopus patagonicus Broth. = Campylopus chilensis fide Frahm 1988.
Campylopus perhorridus Dusén = Chorisodontium aciphyllum fide Hyvönen 1991.
Campylopus permollis Thér. = Campylopus modestus fide Frahm 1988.

28 Müller
Campylopus pilifer Brid.

Tropicos 2009; Frahm 2005; Gradstein & Sipman 1978 (as Campylopus polytrichoides,
Brotherus 1924b; Frahm 1976; Espinosa 1941). Dist. VIII. Arauco, Biobío, Concepción; IX.
Cautín; XIV. Valdivia/Ranco; X. Palena; Juan Fernández Is.
Campylopus polytrichoides De Not. = Campylopus pilifer fide Gradstein & Sipman 1978.
Campylopus purpureocaulis Dusén
Tropicos 2009; Hässel et al. 1984; Larraín 2007a; Dusén 1903b, 1905c; Cardot 1908; Cardot &
Brotherus 1923; Herzog 1954; Frahm 1976; Matteri 1985b; Frahm 1988; Matteri & Schiavone
2002. Dist. VIII. Concepción; X. Chiloé, Llanquihue; XI. Aisén; XII. Magallanes, Última
Esperanza.
Campylopus purpureocaulis subsp. aberrans (Broth.) J.-P. Frahm ex Frank Müll.
Frahm 1987, 1988 (as Campylopus aberrans, Barrera 2006; Espinosa 1941; Brotherus 1924b;
Bartram 1957; Robinson 1975). Dist. Juan Fernández Is.
Campylopus pyriformis (Schultz) Brid.
Larraín 2007b; Larraín 2007a; Frahm 1988, 2005 (as Campylopus blindioides, Brotherus
1924b; Robinson 1975; Frahm 1976; as Campylopus fibrobasis, Dusén 1905c; Cardot 1908;
Thériot 1934b; Herzog & Hosseus 1938; Herzog 1954; as Campylopus kunkelii, Bartram 1957;
Robinson 1975; Frahm 1976; as Campyolpus laxiretis, Bartram 1957; as Campylopus
masafuerae, Frahm 1976; as Campylopus subareodictyon, Brotherus 1924b; Frahm 1976; as
Campylopus torquatus, Frahm 1976; as Campylopus kunkelii, Robinson 1975; as Campylopus
areodictyon, Brotherus 1924b; Frahm 1976). Dist. IV. Choapa; V. Petorca, Valparaíso; VIII.
Concepción; IX. Malleco; XIV. Valdivia/Ranco; X. Chiloé, Osorno, Palena; XI. Aisén; Juan
Fernández Is.
Campylopus recurvifolius Dusén ≡ Campylopus aureonitens subsp. recurvifolius.
Campylopus richardii Brid.
Frahm 1976 (as Thysanomitrium richardii, Brotherus 1924b; Espinosa 1941). Dist. Juan
Fernández Is.
Campylopus saddleanus Besch. = Campylopus vesticaulis fide Frahm 1988.
Campylopus saxicola Broth. = Campylopus vesticaulis fide Frahm 1987.
Campylopus sordidonigritus Dusén, nom. nud. = Campylopus acuminatus fide Frahm 1988.
Campylopus spegazzinii (Müll. Hal.) Paris ≡ Chorisodontium spegazzinii.
Campylopus spiralis Dusén
Dusén 1905c; Cardot 1908; Herzog & Schwabe 1939; Frahm 1976; Thériot 1934b, 1935. Dist.
RM. Santiago; IX. Malleco; XIV. Valdivia/Ranco; X. Llanquihue, Osorno; XI. Aisén; XII.
Magallanes.
Campylopus subareodictyon Broth. = Campylopus pyriformis fide Frahm 1988.
Campylopus sulphureo-nigritus Dusén = Campylopus acuminatus fide Frahm 1988.
Campylopus torquatus Mitt. = Campylopus pyriformis fide Frahm 1988.
Campylopus truncatus Müll. Hal. = Campylopus introflexus fide Frahm 1987.
Campylopus turficola Broth. = Campylopus vesticaulis fide Frahm 1987.
Campylopus vesticaulis Mitt.
Ireland et al. 2007; Ireland & Bellolio 2002; Matteri & Schiavone 2002; Frahm 1988 (as
Campylopus guaitecae, Dusén 1905c; Thériot 1934b; Cardot & Brotherus 1923; Roivainen
1937; Frahm 1976; Deguchi 1991a; as Campylopus saddleanus, Dusén 1903b; Cardot 1908;
Roivainen 1937; Frahm 1976; as Campylopus dicranodontioides, C. saxicola, C. turficola and
C. hygrophilus, Espinosa 1941; Brotherus 1924; as Campylopus dicranodontioides, Barrera
2006; as Campylopus hygrophilus, Barrera 2006; as Campylopus saxicola, Barrera 2006; as
Campylopus turficola, Barrera 2006). Dist. VIII. Arauco, Concepción; XIV. Valdivia/Ranco;
X. Llanquihue; XI. Aisén; XII. Magallanes, Tierra del Fuego; Juan Fernández Is.; Easter I.
Campylopus xanthophyllus Mont. = Campylopus clavatus fide Frahm 1988.
Campylopus spec.
Fuentes 1913. Dist. Easter I.
Campylostelium saxicola (F. Weber & D. Mohr) Bruch & Schimp.
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Mosses of Chile 29
Ireland et al. 2007. Dist. VIII. Arauco, Concepción, Ñuble.

Catagoniopsis berteroana (Mont.) Broth.
Tropicos 2009; Larraín 2005; Villagran et al. 2004; He 1998; Pereira et al. 2006; Espinosa
1941; Müller & Pereira 2006; Thériot 1917, 1918, 1928; Brotherus 1924a, 1924b; Herzog &
Hosseus 1938; Bartram 1952; Herzog 1954; Robinson 1975; Deguchi 1991a; Costes 1921 (as
Hypnum berteroanum, Mitten 1869). Dist. IV. Choapa, Limarí; V. Petorca, Quillota, San
Antonio, Valparaíso; RM. Chacabuco, Cordillera, Maipo, Melipilla, Santiago, Talagante; VI.
Cardenal Caro; VII. Cauquenes, Curicó, Linares, Talca; VIII. Arauco, Biobío, Concepción,
Ñuble; IX. Cautín, Malleco; XIV. Valdivia/Ranco; Juan Fernández Is.
Catagonium complanatum (Cardot & Broth.) Broth. = Catagonium nitens subsp. nitens var. nitens fide Lin 1984.
Catagonium myurum Cardot & Thér. ≡ Catagonium nitens subsp. nitens var. myurum.
Catagonium nitens (Brid.) Cardot subsp. nitens var. nitens
Tropicos 2009; Goffinet et al. 2006; Larraín 2007b; Larraín 2005; He 1998; Buck 2002;
Villagran et al. 2003; Schaumann 2005; Müller & Pereira 2006; Matteri & Schiavone 2002;
Deguchi 1991b (as Catagonium politum, Hässel et al. 1984; as Acrocladium politum, Mitten
1869; as Eucatagonium politum, Dusén 1903b; Brotherus 1908; Cardot 1908; Herzog 1922,
1954, 1960; Reimers 1926; Herzog & Hosseus 1938; Herzog & Schwabe 1939; Seki 1974;
Robinson 1975; Matteri 1985b; as Orthorrhynchium chilense, Williams 1924). Dist. VII.
Cauquenes, Talca; VIII. Arauco, Bíobio, Concepción, Ñuble; IX. Cautín, Malleco; XIV.
Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno; XI. Aisén; XII. Antártica Chilena,
Magallanes; Juan Fernández Is.
Catagonium nitens subsp. nitens var. myurum (Cardot & Thér.) S.H. Lin
Tropicos 2009; Schaumann 2005; Deguchi 1991b; Matteri & Schiavone 2002 (as Catagonium
myurum, Hässel et al. 1984; Thériot 1929b; Matteri 1985b). Dist. VIII. Ñuble; IX. Cautín,
Malleco; X. Llanquihue, Osorno; XII. Magallanes, Última Esperanza.
Catagonium nitidum (Hook. f. & Wilson) Broth.
Goffinet et al. 2006; Hässel et al. 1984; Ireland et al. 2007; Matteri & Schiavone 2002;
Roivainen 1934; Matteri 1985b (as Hypnum nitidum, Cardot 1908; as Stereodon nitidus, Mitten
1869; Dusén 1903b). Dist. VII. Curicó; VIII. Ñuble; IX. Cautín; X. Chiloé; XII. Antártica
Chilena, Magallanes, Última Esperanza.
Catagonium politum (Hook.f. & Wilson) Dusén = Catagonium nitens subsp. nitens var. nitens fide Lin 1984.
Catagonium politum var. phyllogonium (Müll. Hal.) Cardot = Catagonium nitens subsp. nitens var. nitens fide Lin
1984.
Catagonium politum var. filescens Cardot & Broth. = Catagonium nitens subsp. nitens var. nitens fide Lin 1984.
Ceratodon amblyocalyx Müll. Hal. ex Ångström = Ceratodon purpureus subsp. convolutus fide Burley & Pritchard
1990.
Ceratodon antarcticus Cardot = Ceratodon purpureus fide Ochyra 1988.
Ceratodon chilensis (Mont.) Paris ≡ Chrysoblastella chilensis.
Ceratodon convolutus Reichardt ≡ Ceratodon purpureus subsp. convolutus.
Ceratodon crassinervis Lorentz = Ceratodon purpureus subsp. convolutus fide Burley & Pritchard 1990.
Ceratodon grossiretis Cardot = Ceratodon purpureus fide Ochyra 1998.
Ceratodon purpureus (Hedw.) Brid. subsp. purpureus
Tropicos 2009; Larraín 2007b; Hässel et al. 1984; Matcham et al. 2007; Larraín 2005; Buck
2002; Pereira et al. 2006; Espinosa 1941; Villagran et al. 2003; Ireland & Bellolio 2002;
Müller & Pereira 2006; Matteri & Schiavone 2002; Mitten 1869; Dusén 1905b; Cardot 1908;
Thériot 1918, 1935; Herzog 1922, 1954; Cardot & Brotherus 1923; Brotherus 1924b;
Roivainen 1934, 1937; Herzog & Hosseus 1938; Herzog & Schwabe 1939; Bartram 1952; Seki
1974; Robinson 1975; Deguchi 1991a (as Ceratodon purpureus var. fastigiatus, Thériot 1918;
as Ceratodon grossiretis, Thériot 1926). Dist. IV. Limarí; V. Valparaíso; VII. Cauquenes,
Curicó, Talca; VIII. Arauca, Biobío, Concepción, Ñuble; IX. Cautín, Malleco; XIV.
Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno; XI. Aisén, Coihaique; XII. Antártica Chilena,
Magallanes, Última Esperanza; Juan Fernández Is.; Easter I.
Ceratodon purpureus var. amblyocalyx Müll. Hal. = Ceratodon purpureus subsp. convolutus fide Burley & Pritchard
1990.

30 Müller
Ceratodon purpureus subsp. convolutus (Reichardt) Burley

Larraín 2005; Matteri & Schiavone 2002 (as Ceratodon amblyocalyx, Ångström 1876; as
Ceratodon purpureus var. amblyocalyx, Cardot 1900, 1901, 1908; Dusén 1903b; Dusén 1905b;
as Ceratodon convolutus, Dusén 1905b; as Ceratodon crassinervis, Thériot 1918, 1921, 1935;
Reimers 1926; Herzog & Hosseus 1938; as Ceratodon valdiviae, Müller 1900). Dist. VIII.
Arauco, Biobío, Concepción, Ñuble; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Chiloé,
Llanquihue, Osorno; XI. Aisén; XII. Magallanes, Tierra del Fuego. Note: The taxonomic status
of this taxon is uncertain and should be confirmed by biosystematic studies including
molecular methods (Ochyra et al. 2008).
Ceratodon purpureus var. fastigiatus Warnst. = Ceratodon purpureus subsp. purpureus fide Burley & Pritchard 1990.
Ceratodon purpureus subsp. stenocarpus (Bruch & Schimp.) Dixon
He 1998 (as Ceratodon stenocarpus, Ireland et al. 2007). Dist. VIII. Arauco, Biobío,
Concepción, Ñuble; XIV. Valdivia/Ranco.
Ceratodon valdiviae Müll. Hal. = Ceratodon purpureus subsp. convolutus fide Burley & Pritchard 1990.
Cheilothela chilensis (Mont.) Broth. ≡ Chrysoblastella chilensis.
Cheilothela dubia Dusén = Chrysoblastella chilensis fide Reimers 1926, Buck 1981.
Chenia leptophylla (Müll. Hal.) R.H. Zander
Müller 2002; Ireland & Bellolio 2002. Dist. VII. Talca; Easter I.
Chileobryon callicostelloides (Broth. ex Thér.) Enroth
Tropicos 2009; Pereira et al. 2006; Müller & Pereira 2006; Enroth 1992 (as Pinnatella
callicostelloides, Espinosa 1941; Brotherus 1924a; Herzog 1954; as Pinnatella macrosticta,
Brotherus 1924b; Robinson 1975; as Porotrichum callicostelloides, Thériot 1917, 1926; as
Porotrichum callicostelloides var. distichum, Thériot 1924). Dist. IV. Choapa, Limarí; V.
Aconcagua, Petorca, Quillota, San Antonio, Valparaíso; VI. Cardenal Caro, Colchagua; VII.
Cauquenes, Curicó, Talca; VIII. Arauco, Concepción, Ñuble; IX. Cautín, Malleco; XIV.
Valdivia/Ranco; Juan Fernández Is.
Chorisodontium aciphyllum (Hook. f. & Wilson) Broth.
Hässel et al. 1984; Larraín 2007a; Schaumann 2005; Müller & Pereira 2006; Matteri &
Schiavone 2002; Roivainen 1937; Bartram 1952, 1957; Robinson 1975; Matteri 1985b;
Hyvönen 1991 (as Dicranodontium schwabei, Herzog & Schwabe 1939; as Campylopus
laniger, Dusén 1903b; as Dicranum lanigerum, Dusén 1905b; Cardot 1908; Cardot &
Brotherus 1923; as Chorisodontium lanigerum, Roivainen 1937; Bartram 1952; Seki 1974;
Deguchi 1991a; as Dicranum leucopterum, Dusén 1903b, 1905b; Cardot 1908; as
Chorisodontium leucopterum, Hässel et al. 1984; Roivainen 1937; Seki 1974; Matteri 1985b;
as Dicranum macropus, Mitten 1869; as Chorisodontium macropus, Roivainen 1937; Deguchi
1991a; as Chorisodontium nordenskjoldii, Roivainen 1937; as Campylopus orthocomus, Dusén
1903b; as Chorisodontium orthocomum, Roivainen 1937; as Dicranum orthocomum, Cardot
1908; as Chorisodontium sphagneticolum, Roivainen 1937; as Dicranum aciphyllum, Mitten
1869; Cardot 1900, 1901, 1908; Dusén 1905b; Cardot & Brotherus 1923; as Dicranum rigens,
Cardot 1901, 1908; as Dicranum scaberrimum, Dusén 1903b; as Campylopus flavissimus,
Cardot 1901; Dusén 1903b; Dusén 1905c; Cardot 1908; Frahm 1976; as Chorisodontium
fulvastrum, Frahm 1988; as Campylopus perhorridus, Dusén 1905c; Roivainen 1937; Herzog
& Hosseus 1938; Frahm 1976; as Dicranum perhorridum, Cardot 1908). Dist. VII. Cauquenes;
VIII. Ñuble; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno; XI.
Aisén; XII. Antárctica Chilena, Magallanes, Tierra del Fuego, Última Esperanza; Juan
Fernández Is.
Chorisodontium dicranellatum (Dusén) Roiv.
Schaumann 2005; Roivainen 1937; Seki 1974; Matteri & Schiavone 2002; Deguchi 1991a;
Hyvönen 1991 (as Dicranum dicranellatum, Dusén 1903b, 1905b; Cardot 1908). Dist. IX.
Cautín; X. Llanquihue; XI. Aisén; XII. Antártica Chilena, Magallanes, Tierra del Fuego.
Chorisodontium fulvastrum (Besch.) Broth. = Chorisodontium aciphyllum fide Hyvönen 1991.
Chorisodontium lanigerum (Müll. Hal.) Broth. = Chorisodontium aciphyllum fide Hyvönen 1991.
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Mosses of Chile 31
Chorisodontium leucopterum (Müll. Hal.) Roiv. = Chorisodontium aciphyllum fide Matteri 2003a.
Chorisodontium luridum E.B. Bartram
Schaumann 2005; Matteri & Schiavone 2002; Roivainen 1937; Deguchi 1991a; Hyvönen
1991; Frey & Schaumann 2002. Dist. IX. Cautín. X. Llanquihue; XII. Tierra del Fuego.
Chorisodontium macropus (Kunze ex Müll. Hal.) Broth. = Chorisodontium aciphyllum fide Hyvönen 1991.
Chorisodontium magellanicum (Cardot) E.B. Bartram
Roivainen 1937; Hyvönen 1991; Matteri & Schiavone 2002 (as Dicranum magellanicum,
Dusén 1905b; Cardot 1908). Dist. XII. Magallanes, Tierra del Fuego.
Chorisodontium microdus (Lorentz) Broth. = Chorisodontium aciphyllum fide Hyvönen 1991.
Chorisodontium nordenskioldii (Cardot) Roiv. = Chorisodontium aciphyllum fide Hyvönen 1991.
Chorisodontium orthocomum (Besch.) Roiv. = Chorisodontium aciphyllum fide Hyvönen 1991.
Chorisodontium sericeum E.B. Bartram = Ditrichum hookeri fide Hyvönen 1991.
Chorisodontium spegazzinii (Müll. Hal.) Roiv.
Tropicos 2009; Hässel et al. 1984; Roivainen 1937; Matteri 1985b; Hyvönen 1991; Matteri &
Schiavone 2002 (as Campylopus spegazzinii, Cardot 1900, 1901; Dusén 1903b). Dist. XII.
Antártica Chilena, Magallanes, Tierra del Fuega, Última Esperanza.
Chorisodontium sphagneticolum Roiv. = Chorisodontium aciphyllum fide Hyvönen 1991.
Chrysoblastella chilensis (Mont.) Reimers
Tropicos 2009; Larraín 2007b; Hässel et al. 1984; Müller & Pereira 2006; Blockeel et al. 2002;
Deguchi & Oginuma 1989; Matteri 1984a; Reimers 1926; Herzog & Hosseus 1938; Deguchi
1991a (as Ceratodon chilensis, Paris 1903-1906; as Cheilothela chilensis, Thériot 1934b;
Herzog 1954; Seki 1974; Matteri 1985b; as Tortula chilensis, Mitten 1869; as Cheilothela
dubia, Dusén 1905b; Thériot 1934b; Seki 1974). Dist. VII. Cauquenes, Talca; VIII. Arauco,
Biobío, Concepción, Ñuble; IX. Malleco; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue,
Osorno; XI. Aisén; XII. Magallanes, Última Esperanza.
Chryso-hypnum diminutivum (Hampe) W.R. Buck
He 1998. Dist. XIV. Valdivia/Ranco.
Cinclidotus aquaticus (Hedw.) Bruch & Schimp.
He 1998. Dist. XIV. Valdivia/Ranco.
Cinclidium stygium Sw.
Buck 2002; Matteri & Schiavone 2002; Cardot & Brotherus 1923. Dist. XII. Antártica Chilena,
Cladomnion crenato-obtusum Dusén, nom. nud., Dusén 1903b.
Cladomnion gracile (Hampe) Mitt. ≡ Glyphothecium gracile.
Cladomnion valdivae Müll. Hal. = Glyphothecium sciuroides fide Hattaway 1984.
Cladomniopsis crenato-obtusa M. Fleisch.
Hässel et al. 1984; Buck 2002 & 2005a; Matteri 1973a, 1985b; Matteri & Schiavone 2002;
Kühnemann & Goncalves Carralves 1975. Dist. XI. Aisén; XII. Antártica Chilena, Magallanes,
Última Esperanza.
Cochlearidium aymaranum Ochyra & Mahu = Andreaea nitida fide Churchill & Linares 1995a.
Codonoblepharon menziesii Schwägr. ≡ Zygodon menziesii.
Codriophorus laevigatus (A. Jaeger) Bednarek-Ochyra & Ochyra
Bednarek-Ochyra 2006; Larraín 2007a (as Racomitrium laevigatum, Perrier 2005; He 1998;
Matteri & Schiavone 2002; Dusén 1903b, 1906b; Cardot 1908; Robinson 1975; Deguchi
1991a; as Racomitrium loriforme, Espinosa 1941; Dusén 1906b; Cardot 1908; Cardot &
Brotherus 1923; Brotherus 1924b; as Racomitrium plicatum, Donat 1936; as Racomitrium
scabrifolium, Bartram 1946; as Racomitrium willii, Hässel et al. 1984; Cardot & Brotherus
1923; Roivainen 1955a; Seki 1974; Matteri 1985b). Dist. XIV. Valdivia/Ranco; X. Chiloé,
Llanquihue, Osorno; XI. Aisén, General Carrera; XII. Antártica Chilena, Magallanes, Tierra
del Fuego, Última Esperanza; Juan Fernández Is.
Conostomum australe Sw., hom. illeg. = Conostomum pentastichum fide Ochyra et al. 2008.
Conostomum magellanicum Sull.

32 Müller
Goffinet et al. 2006; Hässel et al. 1984; Sullivant 1850; Buck 2002 & 2004; Matteri &
Schiavone 2002; Frahm et al. 1996; Dusén 1903b; Cardot 1908; Matteri 1985a, 1985b;
Deguchi 1991a (as Conostomum perangulatum, Cardot 1908; as Conostomum perangulatum
var. majus, Cardot & Brotherus 1923). Dist. XII. Antártica Chilena, Magallanes, Tierra del
Fuego, Última Esperanza.
Conostomum microphyllum Cardot & Broth., nom. nud., Deguchi 1991a.
Conostomum pentastichum (Brid.) Lindb.
Seki 1974; Matteri 1985a, 1985b; Deguchi 1991a; Hässel et al. 1984; Matteri & Schiavone
2002 (as Conostomum tetragonum, Goffinet et al. 2006; Buck 2002, 2004; as Conostomum
australe, Mitten 1869; Dusén 1903b; Cardot 1908; Cardot & Brotherus 1923; as Conostomum
australe var. microphyllum, Cardot & Brotherus 1923; as Conostomum australe var. piliferum
Cardot & Brotherus 1923). Dist. IX. Malleco; X. Osorno; XI. Aisén; XII. Antártica Chilena,
Conostomum perangulatum Cardot = Conostomum magellanicum fide Matteri 1984b.
Conostomum perangulatum var. majus Cardot & Broth. = Conostomum magellanicum fide Frahm et al. 1996.
Conostomum perpusillum Cardot & Broth.
Hässel et al. 1984; Cardot & Brotherus 1923; Matteri 1985a, 1985b; Frahm et al. 1996; Matteri
& Schiavone 2002. Dist. XII. Magallanes, Última Esperanza.
Coscinodon bolivianus Broth.
Tropicos 2009. Dist. XI. Coihaique (Mancilla 8150, MO, det. C. Delgadillo).
Coscinodon pseudocribrosus Hastings
Tropicos 2009. Dist. V. Los Andes (Mahú 23593, MO, det. C. Delgadillo).
Coscinodontella bryanii R.S. Williams
Cano 2003. Dist. XV. Parinacota.
Costesia macrocarpa (Schimp.) Cuvertino, Miserere & Buffa
Cuvertino et al. 2009 (as Costesia spongiosa, Larraín et al. 2009; Villagran et al. 2004; Thériot
1917, 1921; Costes 1921; as Pottia macrocarpa, Schimper 1836; as Funaria macrocarpa,
Zander 1993; as Tortula macrocarpa, Mitten 1869). Dist. IV. Choapa, Limarí; V. Quillota,
Valparaíso; RM. Santiago; VI. Cachapoal.
Costesia spongiosa Thér. = Costesia macrocarpa fide Cuvertino et al. 2009.
Cratoneuron filicinum (Hedw.) Spruce
Tropicos 2009; Larraín 2005; Robinson 1964; Matteri 1985b; Hedenäs 2003. Dist. II.
Antofagasta; III. Copiapó; IV. Choapa; V. Petorca, San Antonio, Valparaíso; RM. Cordillera,
Santiago; VI. Colchagua; VIII. Biobío, Ñuble; IX. Cautín; XIV. Valdivia/Ranco; X. Osorno.
Cratoneuron perplicatum (Dusén) Broth. = Drepanocladus longifolius fide Ochyra 1989.
Cratoneuropsis relaxa (Hook. f. & Wilson) M. Fleisch. ex Broth. subsp. minor (Wilson &
Hook.f.) Ochyra
Ochyra et al. 2008 (as Cratoneuropsis relaxa, He 1998; Tropicos 2009; as Hygroamblystegium
austrofluviatile, Thériot 1924; Bartram 1943; as Lescuraea chilensis, He 1998; Herzog 1954;
as Orthotheciella varia, Pereira et al. 2006; Buck 2002; Larraín 2007a; He 1998; Müller &
Pereira 2006; Ochyra & Matteri 2001; Matteri & Schiavone 2002; as Hygroamblystegium
varium, Goffinet et al. 2006; as Pseudoleskea chilensis, Larraín 2005; Deguchi 1991b; as
Amblystegium chilensis, Lorentz 1866; as Hygroamblystegium chilensis, Reimers 1926;
Herzog & Hosseus 1938; Herzog & Schwabe 1939; Herzog 1954; Seki 1974; as Sciaromium
gracile, Dusén 1903b; as Hygroamblystegium filum, Hässel et al. 1984; Roivainen 1934;
Herzog & Hosseus 1938; Herzog & Schwabe 1939; Bartram 1952; Matteri 1985b; as
Pseudoleskea filum, Thériot 1918; Herzog 1922; as Hygroamblystegium fuegianum, Larraín
2005; Roivainen 1934; Herzog & Hosseus 1938; Herzog 1960; Seki 1974; as Pseudoleskea
fuegiana, Cardot 1905 & 1908; Thériot 1915, 1917; Herzog 1922; Cardot & Brotherus 1923;
as Leskea fuegiana, Dusén 1903b; as Pseudoleskea lurida, Herzog 1922; as Amblystegium
tenellum, Herzog 1954; as Hygroamblystegium fuegianum var. skottsbergii, Roivainen 1934;
Bartram 1946; as Hygroamblystegium filum var. compactum, Bartram 1946; as
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Mosses of Chile 33
Hygroamblystegium fuegianum var. excurrens, Bartram 1946; as Amblystegium excurrens,

Cardot & Brotherus 1923; as Hygroamblystegium fuegianum var. gracilis, Bartram 1946; as
Pseudoleskea fuegiana var. gracilis, Cardot & Brotherus 1923; as Hygroamblystegium
fuegianum var. secundum, Bartram 1946). Dist. II. Antofagasta; III. Chañaral; IV. Choapa,
Elqui, Limarí; V. Aconcagua, Petorca, Quillota, San Antonio, Valparaíso; RM. Chacabuco,
Cordillera, Maipo, Melipilla, Santiago, Talagante; VI. Colchagua, San Vicente; VII.
Cauquenes, Curicó, Talca; VIII. Arauco, Biobío, Ñuble; XIV. Valdivia/Ranco; X. Chiloé,
Llanquihue, Osorno; XI. Aisén, Coihaique; XII. Antártica Chilena, Magallanes, Tierra del
Crossidium spec.
Matcham et al. 2007. Dist. XII. Magallanes.
Crossidium roseae (R.S. Williams) E.B. Bartram, hom. illeg. ≡ Aloina roseae.
Cryphaea consimilis Mont.
Ireland et al. 2007; Pereira et al. 2006; He 1998; Müller & Pereira 2006; Rao 2001; Mitten
1869; Dusén 1903b; Cardot 1908; Herzog 1922; Reimers 1926; Herzog & Hosseus 1938; Seki
1974. Dist. VII. Cauquenes, Talca; VIII. Arauco, Biobío, Concepción, Ñuble; IX. Cautín,
Malleco; XIV. Valdivia/Ranco; X. Palena; XI. Aisén, Coihaique.
Cryphaea mollis Dusén ≡ Cryphaeophilum molle.
Cryphaea patens Hornsch. ex Müll. Hal.
Rao 2001. Dist. IX. Cautín.
Cryphaea pulchella Paris & Schimp. = Dendrocryphaea lechleri fide Griffin et al. 1982.
Cryphaea tenella Mitt. = Dendrocryphaea lechleri fide Griffin et al. 1982.
Cryphaeophilum molle (Dusén) M. Fleisch.
Ireland et al. 2007; He 1998; Müller & Pereira 2006; Deguchi 1991b; Müller 2002;
Kühnemann & Goncalves Carralves 1976 (as Cryphaea mollis, Dusén 1903b, 1905a; Cardot
1908; as Cryphaeophilum molle var. enerve, Herzog 1954; Seki 1974). Dist. VII. Cauquenes;
VIII. Arauco, Biobío, Concepción, Ñuble; IX. Cautín; XIV. Valdivia/Ranco; X. Llanquihue,
Osorno; XI. Aisén
Cryphaeophilum molle (Dusén) M. Fleisch. var. enerve Herzog = Cryphaeophilum molle fide Müller 2002.
Cryptopapillaria penicillata (Dozy & Molk.) M. Menzel
Ireland et al. 2007. Dist. VIII. Arauco, Concepción.
Cyathophorum splendidissimum (Mont.) Hampe & Lorentz ≡ Schimperobryum splendidissimum.
Cyclodictyon sublimbatum (Müll. Hal.) Kuntze
(as Hookeria sublimbata, Mitten 1869). Dist. Chile, without exact locality (leg. Poeppig).
Cyptodon crassinervis Broth. ≡ Dendrocryphaea crassinervis.
Cynontodium affine (Müll. Hal.) Mitt. = Ditrichum difficile fide Seppelt 1982.
Cynodontium conicum (Mont.) Mitt. ≡ Ditrichum conicum.
Cynodontium elongatum (Hook.f. & Wilson) Mitt. = Ditrichum cylindricarpum fide Seppelt 1982.
Cynodontium hookeri (Müll. Hal.) Mitt. ≡ Ditrichum hookeri.
Cynodontium hyalinum (Mitt.) Mitt. ≡ Ditrichum hyalinum.
Daltonia aristata Geh. & Hampe
(as Daltonia uleana, Reimers 1926). Dist. XIV. Valdivia/Ranco.
Daltonia gracilis Mitt.
Larraín 2007b; Hässel et al. 1984; Larraín 2007a; Ireland et al. 2007; He 1998; Bartram 1931,
1952; Robinson 1975; Matteri 1975b, 1985b; Deguchi 1991a (as Daltonia krauseana, Dusén
1905a; Herzog 1954). Dist. VIII. Arauco; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue; XI.
Daltonia krauseana Müll. Hal. = Daltonia gracilis fide Bartram 1931.
Daltonia patagonica Dusén, nom. nud., Dusén 1903b.
Daltonia trachyodonta Mitt.
Tropicos 2009; Bartram 1931; Seki 1974; Matteri 1975b; Deguchi 1991a (as Daltonia
valdiviae, Herzog 1922). Dist. VIII. Arauco, Biobío; XIV. Valdivia/Ranco; X. Llanquihue; XI.
Aisén.
Daltonia uleana Müll. Hal. = Daltonia aristata fide Bartram 1931.

34 Müller
Daltonia valdiviae Herzog = Daltonia trachyodonta fide Bartram 1931.

Dendrocryphaea crassinervis (Broth.) Enroth
He 1998 (as Cyptodon crassinervis, Brotherus 1924b; Robinson 1975). Dist. XIV.
Valdivia/Ranco; Juan Fernández Is.
Dendrocryphaea cuspidata (Sull.) Broth.
Tropicos 2009; Müller 2009; Larraín 2007a; Ireland et al. 2007; He 1998; Mitten 1869; Thériot
1924, 1934b; Brotherus 1924b; Herzog 1954, 1960. Dist. V. Valparaíso; VI. Cardenal Caro;
VII. Cauquenes, Talca; VIII. Arauco, Biobío, Concepción, Ñuble; XIV. Valdivia/Ranco; X.
Chiloé, Llanquihue, Osorno; Juan Fernández Is.
Dendrocryphaea gorveana (Mont.) Paris & Schimp.
Larraín 2007a; Larraín 2005; Müller & Pereira 2006; Mitten 1869; Dusén 1903b; Cardot 1908;
Herzog 1922; Seki 1974. Dist. VII. Cauquenes, Talca; XIV. Valdivia/Ranco; X. Chiloé,
Palena; XI. Aisén.
Dendrocryphaea lechleri (M. Fleisch.) Paris & Schimp. ex Thér.
Ireland et al. 2007; Müller & Pereira 2006; Thériot 1934b (as Dendrocryphaea ramosissima, as
Cryphaea pulchella, Cardot 1908; as Dendrocryphaea pulchella, Thériot 1934b; as Cryphaea
tenella, Mitten 1869; Dusén 1903b; as Dendrocryphaea tenella Thériot 1934b). Dist. VII.
Cauquenes; VIII. Arauco, Concepción; IX. Cautín; XIV. Valdivia/Ranco.
Dendrocryphaea pulchella (Paris & Schimp.) M. Fleisch. = Dendrocryphaea lechleri fide Griffin et al. 1982.
Dendrocryphaea ramosissima (Paris) Wijk & Margad. = Dendrocryphaea lechleri fide Griffin et al. 1982.
Dendrocryphaea tenella (Mitt.) Thér. = Dendrocryphaea lechleri fide Griffin et al. 1982.
Dendrohypopterygium arbuscula (Brid.) Kruijer
Kruijer 2002 (as Arbusculohypopterygium arbuscula, Schaumann 2005; as Hypopterygium
arbuscula, He 1998; Tropicos 2009; Larraín 2007b; Perrier 2005; Hässel et al. 1984; Larraín
2005; Villagran et al. 2003; Matteri & Schiavone 2002; Seki 1974; Robinson 1975; Matteri
1973b, 1985b; Deguchi 1991b; as Hypopterygium thouini, Eaton 1892; Espinosa 1941; Mitten
1869; Dusén 1903b; Cardot 1908; Thériot 1915, 1918; Herzog 1922, 1954, 1960; Cardot &
Brotherus 1923; Brotherus 1924b; Reimers 1926; Roivainen 1934; Herzog & Hosseus 1938;
Herzog & Schwabe 1939; Bartram 1952, 1957; as Hypopterygium wolffhugelii, Herzog 1922).
Dist. V. Aconcagua, Valparaíso; VI. Colchagua; VIII. Arauco, Biobío; IX. Cautín, Malleco;
XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno; XI. Aisén, Capitán Prat; XII.
Dendroligotrichum dendroides (Brid. ex Hedw.) Broth. subsp. dendroides
Tropicos 2009; Larraín 2007b; Perrier 2005; Hässel et al. 1984; Matcham et al. 2007; Döbbeler
& Triebel 2000; Buck 2004; Döbbeler 2003; Larraín 2005; Espinosa 1941; Villagran et al.
2003; Schaumann 2005; He 1998; Schiavone & Hyvönen 1993; Matteri & Schiavone 2002;
Mitten 1869; Cardot 1908; Thériot 1918; Herzog 1922, 1954, 1960; Cardot & Brotherus 1923;
Brotherus 1924b; Reimers 1926; Herzog & Hosseus 1938; Herzog & Schwabe 1939; Bartram
1952, 1957; Seki 1974; Robinson 1975; Matteri 1985b; Deguchi 1991b; Schiavone 1993 (as
Polytrichadelphus dendroides, Cardot 1900, 1901; Dusén 1903b; as Oligotrichum dendroides,
Kindberg 1888). Dist. VIII. Arauco, Biobío; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X.
Chiloé, Llanquihue, Osorno, Palena; XI. Aisén, Capitán Prat; XII. Antártica Chilena,
Dendroligotrichum squamosum (Hook. f. & Wilson) Cardot
Goffinet et al. 2006; Hässel et al. 1984; Matcham et al. 2007; Buck 2002; Döbbeler 2003;
Schaumann 2005; He 1998; Schiavone & Hyvönen 1991; Matteri & Schiavone 2002; Mitten
1869; Cardot 1908; Herzog 1954; Seki 1974; Matteri 1985b; Deguchi 1991b; Schiavone 1993
(as Polytrichadelphus squamosa, Dusén 1903b). Dist. XIV. Valdivia/Ranco; XI. Aisén,
Desmatodon bellii E.B. Bartram ≡ Hennediella bellii.
Desmatodon subtophaceus (R.S. Williams) R.S. Williams = Didymodon australasiae fide Zander 1978.
Dichodontium dicranelloides Cardot = Dicranella campylophylla fide Ochyra & Newton 1986.
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Mosses of Chile 35
Dichodontium paludella Besch. ≡ Neomeesia paludella.

Dichodontium persquarrosum (Dusén) Cardot = Dicranella campylophylla fide Matteri & Schiavone 2002.
Dichodontium persquarrosum var. falklandicum Cardot = Dicranella campylophylla fide Matteri & Schiavone 2002.
Dicranella aulacocarpa (Mont.) Mitt. = Campylopodium euchlorum fide Matteri & Calabrese 1999.
Dicranella buckiana Larraín & Ochyra, nom. nud., Larraín 2007b (X. Chiloé).
Dicranella campylophylla (Taylor) A. Jaeger
(as Anisothecium cardotii, Matteri & Schiavone 2002; He 1998; as Dicranella patagonica,
Herzog 1922; as Anisothecium persquarrosum, Dusén 1903b; Roivainen 1934, 1937; Bartram
1952; Herzog 1954; as Aongstroemia persquarrosa, Dusén 1905b; as Dichodontium
persquarrosum, Cardot 1908; Cardot & Brotherus 1923; as Dichodontium persquarrosum var.
falklandicum, Cardot 1908; as Dicranella cardotii, Tropicos 2009; Ochyra & Newton 1986;
Hässel et al. 1984; Villagran et al. 2003; Ireland & Bellolio 2002; Seki 1974; Matteri 1985b;
Deguchi 1991a. Dist. V. Aconcagua; RM. Cordillera; VIII. Arauco, Biobío; X. Chiloé,
Llanquihue, Osorno; XI. Aisén, Coihaique; XII. Magallanes, Tierra del Fuego, Última
Esperanza; Easter I.
Dicranella cardotii (R. Br. bis) Dixon = Dicranella campylophylla fide Ochyra et al. 2008.
Dicranella circinata Herzog
NY 2009; Herzog 1954. Dist. X. Chiloé, Llanquihue.
Dicranella costata Broth. = Campylopodium euchlorum fide Matteri & Calabrese 1999.
Dicranella euchlora (Mont.) Lorentz ≡ Campylopodium euchlorum.
Dicranella flexipes Cardot & Broth. = Campylopodium euchlorum fide Matteri & Calabrese 1999.
Dicranella flexiseta Dusén, nom. nud., Paris 1903-1906.
Dicranella fuegiana Cardot & Broth. = Ditrichum heteromallum fide Ochyra et al. 2008.
Dicranella harrisii (Müll. Hal.) Broth.
Tropicos 2009. Dist. VIII. Concepción (Ireland & Bellolio 31893, MO).
Dicranella hawaiica (Müll. Hal.) Broth.
Ireland & Bellolio 2002. Dist. Easter I.
Dicranella hookeri (Müll. Hal.) Cardot
Hässel et al. 1984; Matcham et al. 2007; Larraín 2005; Buck 2002; Matteri & Ochyra 1999; He
1998; Cardot 1908; Thériot 1924; Seki 1974; Matteri 1985b; Deguchi 1991a (as Anisothecium
hookeri, Ireland & Bellolio 2002; Matteri & Schiavone 2002; Thériot 1929b; Roivainen 1937;
Herzog & Hosseus 1938; Herzog 1954; as Anisothecium perpusillum, Dusén 1903b; as
Chorisodontium sericeum, Bartram 1946; as Dicranella perpusilla, Cardot & Brotherus 1923;
as Dicranella subclathrata, Thériot 1924; as Meesia patagonica, Dusén 1903b; Cardot 1908;
as Dichodontium dicranelloides, Cardot 1905; as Dicranella patagonica, Herzog 1922). Dist.
IV. Elqui; V. Aconcagua, Petorca, San Antonio, Valparaíso; RM. Chacabuco, Cordillera,
Maipo, Santiago; VI. Cachapoal; VII. Curicó, Talca; VIII. Biobío; IX. Cautín, Malleco; XIV.
Valdivia/Ranco; X. Osorno; XI. Aisén, Coihaique; XII. Antártica Chilena, Magallanes, Tierra
del Fuego, Última Esperanza; Easter I.
Dicranella hookeri var. flexisubulata Cardot & Broth.
Cardot & Brotherus 1923; Seki 1974. Dist. XI. Aisén.
Dicranella jamesonii (Mitt.) Broth. = Anisothecium vaginatum fide Matteri 2003a.
Dicranella kunzeana Müll. Hal. ≡ Leptotrichella kunzeana.
Dicranella paludella (Besch.) Dusén ≡ Neomeesia paludella.
Dicranella patagonica Müll. Hal. = Dicranella hookeri fide Ochyra 1999b.
Dicranella perpusilla (Dusén) Cardot = Dicranella hookeri fide Newton 1977.
Dicranella pseudorufescens Cardot & Broth. ≡ Anisothecium pseudorufescens.
Dicranella pseudorufescens var. leptoclada Cardot & Broth. ≡ Anisothecium pseudorufescens var. leptocladus.
Dicranella skottsbergii Cardot & Broth. ≡ Anisothecium skottsbergii.
Dicranella subclathrata Lorentz = Dicranella hookeri fide Ochyra 1999b.
Dicranodontium asperifolium (Mitt.) Broth. ≡ Campylopus asperifolius.
Dicranodontium schwabei Herzog & Thér. = Chorisodontium aciphyllum fide Müller 2002.
Dicranoloma australe (Besch.) Paris = Dicranoloma robustum fide Matteri & Schiavone 2002.
Dicranoloma australe var. schwabei Herzog & Thér. in Herzog & Schwabe
Herzog & Schwabe 1939. Dist. X. Llanquihue.

36 Müller
Dicranoloma billardierei (Brid.) Paris var. billardierei

Díaz et al. 2008; Tropicos 2009; Larraín 2007b; Hässel et al. 1984; Larraín 2005; Ireland et al.
2007; Espinosa 1941; Villagran et al. 2003; Schaumann 2005; He 1998; Müller & Pereira
2006; Matteri & Schiavone 2002; Thériot 1918, 1935; Herzog 1922, 1954, 1960; Cardot &
Brotherus 1923; Brotherus 1924b; Roivainen 1934, 1937; Herzog & Hosseus 1938; Herzog &
Schwabe 1939; Bartram 1952, 1957; Seki 1974; Robinson 1975; Matteri 1985b; Deguchi
1991a (as Dicranoloma fernandezianum, Barrera 2006; Brotherus 1924b; Bartram 1957; as
Dicranum billardierei, Mitten 1869; Dusén 1903b; Cardot 1908; as Leucoloma billardierii,
Dusén 1905b). Dist. V. Valparaíso; VII. Cauquenes; VIII. Arauco, Concepción; IX. Malleco;
XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno, Palena; XI. Aisén; XII. Magallanes,
Dicranoloma billardierei var. compactum (Cardot) Cardot & Broth.
Matteri & Schiavone 2002 (as Dicranum billardierei var. compactum, Cardot 1908). Dist. XI.
Aisén; XII. Magallanes.
Dicranoloma capillare (Dusén) Broth.
Schaumann 2005; Seki 1974; Deguchi 1991a (as Dicranum capillare, Cardot 1908; as
Leucoloma capillare, Dusén 1905b; as Leucoloma nigricaule var. gracile, Dusén 1905b). Dist.
IX. Cautín; X. Llanquihue; XI. Aisén.
Dicranoloma capillifolioides Broth.
Espinosa 1941; Brotherus 1924b; Bartram 1957; Robinson 1975. Dist. Juan Fernández Is.
Dicranoloma capillifolium (Cardot) Thér. = Dicranoloma menziesii fide Greene 1986.
Dicranoloma chilense (De Not.) Ochyra & Matteri
Goffinet et al. 2006; Larraín 2007b; Matcham et al. 2007; Buck 2002; Larraín 2007a;
Schaumann 2005; He 1998; Matteri & Schiavone 2002 (as Dicranoloma nigricaule, Cardot &
Brotherus 1923; Brotherus 1924b; Roivainen 1937; Bartram 1952; Seki 1974; Robinson 1975;
Deguchi 1991a; as Dicranum nigricaule, Cardot 1901; Dusén 1903b; Cardot 1908; as
Leucoloma nigricaule, Dusén 1905b; as Dicranum peruncinatum, Dusén 1903b; Cardot 1908;
as Leucoloma peruncinatum, Dusén 1905b; as Dicranum racovitzae, Cardot 1900, 1901). Dist.
V. Valparaíso; IX. Malleco; XIV. Valdivia/Ranco; X. Chiloé, Osorno; XI. Aisén; XII.
Antártica Chilena, Magallanes, Tierra del Fuego; Juan Fernández Is.
Dicranoloma dusenii (Broth.) Broth.
Herzog 1954 (as Leucoloma dusenii, Dusén 1905b). Dist. VIII. Arauco; X. Osorno.
Dicranoloma fernandezianum Broth. = Dicranoloma billardierei fide Robinson 1975.
Dicranoloma fuegianum (Dusén) Broth. = Dicranoloma hariotii fide Wijk et al. 1962.
Dicranoloma hariotii (Müll. Hal.) Paris var. hariotii
Hässel et al. 1984; Matteri & Schiavone 2002; Cardot & Brotherus 1923; Roivainen 1937;
Herzog 1954; Seki 1974; Matteri 1985b (as Dicranum hariotii, Cardot 1900, 1901; as
Leucoloma hariotii, Dusén 1905b; as Leucoloma fuegianum, Dusén 1905b). Dist. XI. Aisén;
XII. Magallanes, Tierra del Fuego, Última Esperanza.
Dicranoloma hariotii var. laxum (Dusén) Cardot & Broth.
Cardot & Brotherus 1923 (as Leucoloma fuegianum var. laxum, Dusén 1905b). Dist. XII.
Magallanes.
Dicranoloma hollermayeri Thér.
Thériot 1935. Dist. XIV. Valdivia/Ranco.
Dicranoloma imponens (Mont.) Renauld
Díaz et al. 2008; Larraín 2007b; Hässel et al. 1984; Larraín 2005; Villagran et al. 2003; Matteri
& Schiavone 2002; Thériot 1921; Cardot & Brotherus 1923; Roivainen 1934, 1937; Herzog &
Schwabe 1939; Matteri 1985b (as Dicranum imponens, Eaton 1892; Cardot 1908; as
Leucoloma imponens, Dusén 1905b; as Dicranum involutifolium, Sullivant 1850). Dist. XIV.
Valdivia/Ranco; X. Chiloé, Llanquihue; XI. Aisén; XII. Antártica Chilena, Magallanes, Tierra
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Mosses of Chile 37
Dicranoloma kunkelii H. Rob.

Robinson 1975. Dist. Juan Fernández Is.
Dicranoloma menziesii (Hook. f. & Wilson) Paris
Larraín 2005; He 1998; Brotherus 1924b; Robinson 1975 (as Dicranoloma capillifolium,
Thériot 1918; Cardot & Brotherus 1923; Herzog & Hosseus 1938; Herzog 1954, 1960;
Bartram 1957; Dicranum capillifolium, Barrera 2006; Cardot 1908; as Campylopus
fernandezianus, Bartram 1957; Frahm 1976; as Leucoloma capillifolium, Dusén 1905b). Dist.
XIV. Valdivia/Ranco; X. Chiloé, Llanquihue; XI. Aisén; XII. Magallanes; Juan Fernández Is.
Dicranoloma menziesii var. chilense Thér.
Thériot 1924. Dist. Without exact locality.
Dicranoloma muelleri (Dusén) Roiv.
Larraín 2007b; Villagran et al. 2003; Roivainen 1937; Herzog 1954; Seki 1974; Deguchi 1991a
(as Leucoloma muelleri, Dusén 1905b). Dist. IX. Malleco; XIV. Valdivia/Ranco; X. Chiloé,
Llanquihue; XI. Aisén; XII. Magallanes, Tierra del Fuego.
Dicranoloma nigricaule (Ångström) Paris = Dicranoloma chilense fide Ochyra & Matteri 1996.
Dicranoloma nigricaule var. flexuosulum (Dusén) E.B. Bartram
(as Leucoloma nigricaule var. flexuosulum, Dusén 1905b). Dist. XIV. Valdivia/Ranco.
Dicranoloma perremotifolium (Dusén) Broth.
Tropicos 2009; Larraín 2005; Villagran et al. 2003; He 1998; Roivainen 1934; Herzog 1954,
1960 (as Dicranum perremotifolium, Dusén 1903b; Cardot 1908; as Leucoloma
perremotifolium, Dusén 1905b). Dist. VIII. Biobío; IX. Malleco; XIV. Valdivia/Ranco; X.
Chiloé, Osorno; XI. Aisén.
Dicranoloma perremotifolium var. fragile (Dusén) Thér.
Thériot 1935 (as Leucoloma perremotifolium var. fragile, Dusén 1905b). Dist. VIII. Arauco;
XIV. Valdivia/Ranco; X. Osorno.
Dicranoloma peruncinatum (Dusén) Broth. = Dicranoloma chilense fide Wijk et al. 1962.
Dicranoloma pungens (Hook.f. & Wilson) Paris = Dicranoloma robustum fide Sainsbury 1955.
Dicranoloma robustum var. pungens Hook.f. & Wilson = Dicranoloma robustum fide Sainsbury 1955.
Dicranoloma robustum (Hook. f. & Wilson) Paris var. robustum
Goffinet et al. 2006; Larraín 2007b; Hässel et al. 1984; Buck 2002; Larraín 2005; Villagran et
al. 2003; Schaumann 2005; Matteri & Schiavone 2002 (as Dicranoloma australe, Cardot &
Brotherus 1923; Roivainen 1937; Seki 1974; as Dicranum australe, Dusén 1903b; Cardot
1908; as Leucoloma robustum var. flexuosum, Dusén 1905b; He 1998; Herzog 1922; Cardot &
Brotherus 1923; Roivainen 1937; Herzog & Hosseus 1938; Bartram 1952; Seki 1974; Matteri
1985b; Deguchi 1991a; as Dicranoloma pungens, Reimers 1926; Thériot 1935; Herzog &
Hosseus 1938; Herzog & Schwabe 1939; as Dicranum pungens, Cardot 1908; as Dicranum
grandialare, Dusén 1903b; as Dicranum robustum, Eaton 1892; Mitten 1869; Cardot 1900,
1901, 1908; Dusén 1903b; as Leucoloma grandialare, Dusén 1905b; as Leucoloma robustum,
Dusén 1905b; as Dicranoloma robustum var. pungens, Eaton 1892). Dist. IX. Cautín; XIV.
Dicranoloma robustum var. longisetaceum Herzog
Larraín 2005; Herzog & Hosseus 1938. Dist. XIV. Valdivia/Ranco; X. Osorno.
Dicranoloma robustum var. setosum (Hook. f. & Wilson) Sainsbury
(as Dicranoloma setosum, Herzog 1922, 1954; Roivainen 1934; Bartram 1952; Seki 1974, as
Dicranum setosum, Cardot 1908; as Leucoloma setosum, Dusén 1905b). Dist. XIV.
Valdivia/Ranco; X. Osorno; XI. Aisén; XII. Magallanes.
Dicranoloma robustum var. strictifolium (Dusén) Wijk & Margad.
(as Leucoloma muelleri var. strictifolium, Dusén 1905b; as Dicranum setosum var.
strictifolium, Cardot 1908). Dist. XI. Aisén.
Dicranoloma setosum (Hook.f. & Wilson) Paris ≡ Dicranoloma robustum var. setosum.
Dicranoloma subimponens (Cardot) Broth.

38 Müller
Matteri & Schiavone 2002; Cardot & Brotherus 1923; Roivainen 1937; Seki 1974. Dist. XI.
Aisén; XII. Magallanes, Tierra del Fuego.
Dicranoweisia antarctica (Müll. Hal.) Paris ≡ Hymenoloma antarcticum.
Dicranoweisia antarctica fo. breviseta Roiv. = Hymenoloma antarcticum fide Ochyra et al. 2008.
Dicranoweisia austrocrispula (Müll. Hal.) Paris = Hymenoloma crispulum fide Ochyra et al. 2008.
Dicranoweisia brevipes (Müll. Hal.) Cardot = Hymenoloma antarcticum fide Ochyra et al. 2008.
Dicranoweisia breviseta Cardot = Hymenoloma antarcticum fide Ochyra et al. 2008.
Dicranoweisia crispula (Hedw.) Lindb. ex Milde ≡ Hymenoloma crispulum.
Dicranoweisia cryptodon (Mitt.) Paris ≡ Camptodontium cryptodon.
Dicranoweisia humilis (Müll. Hal.) Broth = Kiaeria pumila fide Ochyra 1993a.
Dicranoweisia jugellifera Dusén ≡ Hymenoloma jugelliferum.
Dicranoweisia mackayi (Broth. & Dixon) Broth. = Hymenoloma antarcticum fide Ochyra et al. 2008.
Dicranoweisia nordenskjoeldii (Dusén) Reim. = Hymenoloma antarcticum fide Ochyra et al. 2008.
Dicranoweisia perpulvinata Dusén = Hymenoloma crispulum fide Cardot 1908.
Dicranoweisia subglobosa (Herzog) S. He ≡ Hymenoloma subglobosa.
Dicranoweisia subinclinata (Müll. Hal.) Broth. = Hymenoloma crispulum fide Ochyra 1998.
Dicranoweisia tenella Dusén, nom. nud. = Camptodontium cryptodon fide Reimers 1936.
Dicranoweisia turpis (Cardot) Ochyra ≡ Hymenoloma turpe.
Dicranum aciphyllum Hook.f. & Wilson ≡ Chorisodontium aciphyllum.
Dicranum antarcticum Müll. Hal. ≡ Hymenoloma antarcticum.
Dicranum australe Besch. = Dicranoloma robustum fide Matteri & Schiavone 2002.
Dicranum billardierei Brid. ≡ Dicranoloma billardierei.
Dicranum billardierei var. compactum Cardot ≡ Dicranoloma billardierei var. compactum.
Dicranum billardierei var. flavicans Cardot
Cardot 1908. Dist. XII. Última Esperanza.
Dicranum capillare (Dusén) Cardot ≡ Dicranoloma capillare.
Dicranum capillifolium (Broth.) Cardot = Dicranoloma menziesii fide Robinson 1975.
Dicranum cardotii R.Br. bis = Dicranella campylophylla fide Ochyra et al. 2008.
Dicranum chilense De Not. ≡ Dicranoloma chilense.
Dicranum cirrhifolium Schimp. ex Dusén = Platyneuron praealtum fide Ochyra & Bednarek-Ochyra 1997.
Dicranum cryptodon Mont. ≡ Camptodontium cryptodon.
Dicranum dicranellatum Dusén ≡ Chorisodontium dicranellatum.
Dicranum euchloron Mont. ≡ Campylopodium euchloron.
Dicranum grandialare Dusén, nom. nud. = Dicranoloma robustum fide Cardot 1908.
Dicranum hariotii Hook.f. & Wilson ≡ Dicranoloma hariotii.
Dicranum imponens Mont. ≡ Dicranoloma imponens.
Dicranum inerme Mitt. = Holodontium strictum fide Ochyra 1993a.
Dicranum involutifolium Sull. = Dicranoloma imponens fide Wijk et al. 1962.
Dicranum lanigerum Müll. Hal. = Chorisodontium aciphyllum fide Hyvönen 1991.
Dicranum laticostatum Cardot = Platyneuron praealtum fide Ochyra & Bednarek-Ochyra 1997.
Dicranum leucopterum Müll. Hal. = Chorisodontium aciphyllum fide Hyvönen 1991.
Dicranum macropus Kunze ex Müll. Hal. = Chorisodontium aciphyllum fide Hyvönen 1991.
Dicranum magellanicum Cardot ≡ Chorisodontium magellanicum.
Dicranum nigricaule Ångström = Dicranoloma chilense fide Ochyra & Matteri 1996.
Dicranum orthocomum (Besch.) Müll. Hal. = Chorisodontium aciphyllum fide Hyvönen 1991.
Dicranum percompactum Dusén, nom. nud. = Chorisodontium magellanicum fide Roivainen 1937.
Dicranum perhorridum (Dusén) Cardot = Chorisodontium aciphyllum fide Hyvönen 1991.
Dicranum perremotifolium (Dusén) Cardot ≡ Dicranoloma perremotifolium.
Dicranum peruncinatum (Dusén) Cardot = Dicranoloma chilense fide Ochyra & Matteri 1996.
Dicranum pumilum (Mitt.) Mitt. ≡ Kiaeria pumila.
Dicranum pungens Hook.f. & Wilson = Dicranoloma robustum fide Sainsbury 1955.
Dicranum racovitzae Cardot = Dicranoloma chilense fide Ochyra & Matteri 1996.
Dicranum ramulosum Mitt. = Kiaeria pumila fide Ochyra 1993.
Dicranum rigens Besch. = Chorisodontium aciphyllum fide Cardot 1916.
Dicranum robustum Hook.f. & Wilson ≡ Dicranoloma robustum.
Dicranum robustum var. giganteum Cardot
Cardot 1908. Dist. XII. Magallanes.
Dicranum robustum var. lagunicola (Dusén) Cardot
Cardot 1908 (as Leucoloma robustum var. lagunicola, Dusén 1905b). Dist. XII. Magallanes.
Dicranum scaberrimum Broth. ex Dusén = Chorisodontium aciphyllum fide Cardot 1909.
Dicranum setosum Hook.f. & Wilson ≡ Dicranoloma robustum var. setosum.
Dicranum setosum var. strictifolium (Dusén) Cardot = Dicranoloma robustum var. strictifolium.
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Didymodon acutus (Brid.) K. Saito

Didymodon ampliretis E.B. Bartram
Hässel et al. 1984; Matteri & Schiavone 2002; Bartram 1946; Matteri 1985b. Dist. XII.
Didymodon andreaeoides Cardot & Broth.
Tropicos 2009; Müller 2009; Hässel et al. 1984; Matteri 1985b; Matteri & Schiavone 2002.
Dist. IX. Cautín; XIV. Valdivia/Ranco; X. Osorno; XI. Coihaique; XII. Magallanes, Última
Esperanza.
Didymodon australasiae (Hook. & Grev.) R.H. Zander
Tropicos 2009; Müller 2009; Jiménez & Cano 2007; Jiménez et al. 2005; He 1998; Zander
1978 (as Barbula australasiae, Dusén 1906a; as Tortula australasiae, Mitten 1869; as Tortula
graminicolor, Mitten 1869; as Desmatodon subtophaceus, Williams 1915; as Didymodon
subtophaceus, Herzog 1954; as Tortula geniculata, Mitten 1869; as Asteriscium geniculatum,
Herzog & Hosseus 1938; Herzog 1954; as Barbula poeppigiana, Espinosa 1941; Thériot 1917,
1928; Costes 1921; Brotherus 1924a). Dist. XV. Parinacota; II. El Loa; IV. Choapa, Elqui,
Limarí; V. Aconcagua, Petorca, Valparaíso; RM. Chacabuco, Melipilla, Santiago; VI.
Cachapoal, Colchagua; VII. Talca; VIII. Concepción; IX. Cautín; XI. Aisén; XII. Última
Esperanza.
Didymodon berthoanus Thér. ≡ Bryoerythrophyllum berthoanus.
Didymodon calymperidictyon Broth. = Bryoerythrophyllum campylocarpum fide Robinson 1975.
Didymodon cardotii (Dusén) R.H. Zander
(as Barbula cardotii, Dusén 1905a, Dusén 1906a). Dist. XII. Magallanes.
Didymodon coquimbensis J.A. Jiménez & Cano
Jiménez & Cano 2007. Dist. IV. Elqui.
Didymodon deciduus R.H. Zander
Zander 1993 (as Asteriscium deciduum, Herzog & Hosseus 1938). Dist. VIII. Biobío.
Didymodon fuscus (Müll. Hal.) J.A. Jiménez & Cano
Jiménez & Cano 2007 (as Barbula fusca, Costes 1921; Dusén 1906a; Cardot 1908; Thériot
1918, 1921, 1928; Herzog & Hosseus 1938; Bartram 1952; as Tortula fusca, Mitten 1869; as
Barbula fuscoviridis, Williams 1915; Thériot 1921). Dist. IV. Choapa, Elqui, Limarí; V.
Petorca, Quillota, San Antonio, Valparaíso; RM. Santiago; VI. Cachapoal; VII. Cauquenes;
VIII. Biobío, Concepción, Ñuble; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Llanquihue;
XI. Aisén.
Didymodon laevigatus (Mitt.) R.H. Zander
(as Barbula laevigata, Dusén 1906b; Cardot 1908). Dist. XI. Aisén.
Didymodon linearis Broth. = Bryoerythrophyllum campylocarpum fide Robinson 1975.
Didymodon luridus Hornsch. ex Spreng.
Tropicos 2009. Dist. XIV. Valdivia/Ranco (Mahú 11737, 12313, 12317, MO, det. Jiménez).
Didymodon orbignyanus (Müll. Hal.) Broth.
Brotherus 1924a; Espinosa 1941. Dist. RM. Santiago.
Didymodon patagonicus (Mitt.) Broth.
(as Barbula patagonica, Dusén 1903b). Dist. XII. Patagonia.
Didymodon reflexus Thér.
Thériot 1924. Dist. V. Valparaíso.
Didymodon rigidulus Hedw.
Tropicos 2009; Pereira et al. 2006; Schaumann 2005; Frey & Schaumann 2002; Matteri &
Schiavone 2002 (as Barbula rigidula, Roivainen 1934). Dist. IV. Limarí; VII. Talca; VIII.
Arauco, Ñuble. X. Llanquihue; XII. Fuegian region.
Didymodon santessonii (E.B. Bartram) J.A. Jiménez & Cano

40 Müller
(as Barbula santessonii, Bartram 1952; Herzog 1954; Seki 1974). Dist. IV. Choapa, Limarí; V.
Aconcagua, Quillota, San Antonio, Valparaíso; VI. Cardenal Caro; XI. Aisén.
Didymodon schilleri Herzog & Thér.
NY 2009; Herzog & Hosseus 1938. Dist. IX. Cautín; X. Palena.
Didymodon schimperi (Mont.) Broth. = Tortula atrovirens fide Zander 1993.
Didymodon subtophaceus R.S. Williams = Didymodon australasiae fide Zander 1978.
Didymodon torquatus (Taylor) Catcheside = Didymodon vinealis fide Sollman 1995.
Didymodon tophaceus (Brid.) Lisa
Tropicos 2009. Dist. IV. Choapa (Mahú 12571, 11839, MO, det. Jiménez), Limarí (Mahú
24358, MO, det. Jiménez); V. San Antonio (Mahú 23410, MO, det. Jiménez), Valparaíso
(Mahú 11747, MO, det. Jiménez); RM. Cordillera (Mahú 23247, MO, det. Jiménez), Melipilla
(Mahú 20083, MO, det. Jiménez).
Didymodon umbrosus (Müll. Hal.) R. H. Zander
Tropicos 2009; Müller 2009. Dist. IV. Limarí; RM. Santiago.
Didymodon vinealis (Brid.) R.H. Zander
Villagran et al. 2004; He 1998; Müller & Pereira 2006 (as Asteriscium poeppigianum, Herzog
& Hosseus 1938; Herzog 1954; as Weissia poeppigiana, Mitten 1869). Dist. IV. Coquimbo,
Limarí; V. Aconcagua, Valparaíso; RM. Santiago, Talagante; VII. Cauquenes, Linares; VIII.
Biobío, Concepción; IX. Malleco; XIV. Valdivia/Ranco; XI. Aisén.
Diphyscium pilmaiquen (Crosby) Magombo
Ireland et al. 2007; Magombo 2003 (as Muscoflorschuetzia pilmaiquen, Crosby 1977, 1978;
Deguchi 1991a; as Florschuetzia pilmaiquen, Barrera 2006). Dist. VIII. Arauco; X. Osorno.
Diplostichum poeppigii (Müll. Hal.) Cardot, hom. illeg. = Eustichia longirostris fide Allen 2002.
Dissodon magellanicum (Brid.) Hampe ≡ Tayloria magellanica.
Dissodon mirabilis Cardot ≡ Tayloria mirabilis.
Distichium capillaceum (Hedw.) Bruch & Schimp.
Hässel et al. 1984; Buck 2002; Schaumann 2005; He 1998; Matteri & Schiavone 2002; Dusén
1903b, 1905b; Roivainen 1937; Bartram 1952; Matteri 1985b; Deguchi 1991a (as Distichium
capillaceum var. compactum, Matteri & Schiavone 2002; Hässel et al. 1984). Dist. VII. Curicó,
Talca; IX. Cautín; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue; XII. Antártica Chilena,
Distichium capillaceum var. compactum (Huebener) Dalla Torre & Sarnth. = Distichium capillaceum fide Ochyra et al.
2008.
Distichophyllum assimile Broth.
Brotherus 1924b; Bartram 1957; Robinson 1975; Matteri 1975b. Dist. XIV. Valdivia/Ranco;
Juan Fernández Is.
Distichophyllum cavifolium (Cardot) Cardot = Distichophyllum eremitae fide Matteri 1975b.
Distichophyllum cavifolium var. montanum Cardot = Distichophyllum eremitae fide Matteri 1975b.
Distichophyllum crispatissimum Dusén, nom. nud. = Distichophyllum ellipticum fide Matteri 1975b.
Distichophyllum dicksonii (Hook. & Grev.) Mitt.
Larraín 2007b; Hässel et al. 1984; He 1998; Matteri & Schiavone 2002; Mitten 1869; Dusén
1903b; Cardot 1908; Herzog & Hosseus 1938; Herzog 1954; Seki 1974; Matteri 1975b, 1985b;
Deguchi 1991a. Dist. XIV. Valdivia/Ranco; X. Chiloé; XI. Aisén; XII. Antártica Chilena,
Distichophyllum ellipticum Herzog
Larraín 2007b; Hässel et al. 1984; Larraín 2007a; Matteri & Schiavone 2002; Herzog 1954;
Matteri 1975b, 1985b (as Distichophyllum crispatissimum, Dusén 1903b; Herzog & Schwabe
1939). Dist. X. Chiloé; XI. Aisén; XII. Magallanes, Última Esperanza.
Distichophyllum eremitae (A. Jaeger) Paris
Hässel et al. 1984; Matteri & Schiavone 2002; Dusén 1903b; Cardot 1908; Seki 1974; Matteri
1975b, 1985b (as Distichophyllum cavifolium, Cardot 1908; as Distichophyllum cavifolium var.
montanum, Cardot & Brotherus 1923; as Distichophyllum procumbens, Mitten 1869). Dist. XI.
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Mosses of Chile 41
Distichophyllum fasciculatum Mitt.

Herzog 1960. Dist. XIV. Valdivia/Ranco.
Distichophyllum fernandezianum Broth.
He 1998; Matteri 1975b. Dist. XIV. Valdivia/Ranco; Juan Fernández Is.
Distichophyllum flaccidum (Hook.f. & Wilson) Mitt.
Hässel et al. 1984; Buck 2005b; He 1998; Mitten 1869; Cardot 1908; Matteri 1975b, 1985b;
Matteri & Schiavone 2002 (as Mniadelphus flaccidus, Dusén 1903b). Dist. IX. Malleco; XII.
Distichophyllum grandifolium Matteri
Matteri 1975b. Dist. X. Llanquihue.
Distichophyllum krausei (Lorentz) Mitt.
Hässel et al. 1984; He 1998; Müller & Pereira 2006; Matteri & Schiavone 2002; Cardot 1908;
Matteri 1975b, 1985b; Deguchi 1991a (as Mniadelphus krausei, Dusén 1903b; as
Distichophyllum nigricans, Dusén 1903b; Cardot 1908). Dist. V. Valparaíso; VII. Cauquenes;
VIII. Ñuble; IX. Cautín; XIV. Valdivia/Ranco; X. Llanquihue, Osorno; XI. Aisén; XII.
Distichophyllum molle Besch. ≡ Calyptrochaete mollis.
Distichophyllum nanospathulatum Herzog
Schaumann 2005; He 1998; Herzog 1954; Matteri 1975b. Dist. XIV. Valdivia/Ranco; X.
Osorno; XI. Aisén.
Distichophyllum nigricans Besch. = Distichophyllum krausei fide Matteri 1975a.
Distichophyllum patagonicum Besch.
Dusén 1903b; Cardot 1908. Dist. XII. Magallanes.
Distichophyllum procumbens Mitt., hom. illeg. = Distichophyllum eremitae fide Matteri 1975b.
Distichophyllum rotundifolium (Hook.f. & Wilson) Müll. Hal. & Broth.
Schaumann 2005; Streimann 1999; He 1998; Cardot & Brotherus 1923; Herzog 1960; Matteri
1975b; Deguchi 1991a. Dist. X. Llanquihue, Osorno; XI. Aisén; XII. Tierra del Fuego; Juan
Fernández Is.
Distichophyllum subelimbatum Broth.
Barrera 2006; Espinosa 1941; Brotherus 1924b; Bartram 1957; Robinson 1975; Matteri 1975b.
Dist. Juan Fernández Is.
Distichophyllum submersum Matteri
Hässel et al. 1984; Matteri 1975b, 1985b; Matteri & Schiavone 2002. Dist. XII. Magallanes,
Tierra del Fuego, Última Esperanza.
Distichophyllum theriotianum Matteri
Matteri 1975b. Dist. XI. Aisén.
Ditrichum affine (Müll. Hal.) Hampe. = Ditrichum difficile fide Seppelt 1982.
Ditrichum austrogeorgicum (Cardot) Seppelt = Ditrichum hyalinum fide Ochyra 1998.
Ditrichum brotherusii (R. Br. bis) Seppelt
Buck 2002; Matteri & Schiavone 2002; Deguchi 1991a (as Pseudodistichium fuegianum,
Roivainen 1937). Dist. XII. Antártica Chilena, Magallanes, Tierra del Fuego.
Ditrichum conicum (Mont.) Mitt.
Ochyra 1999c; Matteri & Schiavone 2002; Dusén 1903b, 1905b; Cardot 1908; Cardot &
Brotherus 1923; Thériot 1935; Roivainen 1937; Herzog & Hosseus 1938; Herzog & Schwabe
1939; Bartram 1952; Seki 1974; Deguchi 1991a (as Cynodontium conicum, Mitten 1869; as
Pseudodistichium atlanticum, Robinson 1964; as Pseudodistichium taitaoense, Froehlich 1953;
as Ditrichum conicum var. glaciale, Cardot & Brotherus 1923). Dist. IX. Cautín; XIV.
Valdivia/Ranco; X. Llanquihue, Osorno; XI. Aisén; XII. Antártica Chilena, Tierra del Fuego,
Última Esperanza.
Ditrichum conicum var. glaciale Cardot & Broth. = Ditrichum conicum fide Ochyra 1999c.
Ditrichum cylindricarpum (Müll. Hal.) F. Muell.

42 Müller
Larraín 2007b; Buck 2002; Larraín 2007a; Ireland et al. 2007; He 1998; Deguchi 1991a (as
Cynontodium elongatum, Mitten 1869; as Ditrichum elongatum, Cardot 1908; Herzog 1922,
1954; Seki 1974). Dist. VIII. Arauco, Biobío, Ñuble; IX. Malleco; XIV. Valdivia/Ranco; X.
Chiloé, Llanquihue, Osorno; XI. Aisén; XII. Antártica Chilena, Magallanes.
Ditrichum difficile (Duby) M. Fleisch.
Tropicos 2009; Larraín 2007b; Larraín 2005; Villagran et al. 2003; Schaumann 2005; Ireland
& Bellolio 2002; He 1998; Müller & Pereira 2006; Robinson 1975; Deguchi 1991a; Matteri &
Schiavone 2002 (as Cynontodium affine, Mitten 1869; as Ditrichum affine, Espinosa 1941;
Dusén 1903b, 1905b; Cardot 1908; Thériot 1915; Herzog 1922; Brotherus 1924b; Seki 1974;
as Ditrichum flexifolium, Thériot 1921; Herzog & Hosseus 1938; Bartram 1957; Robinson
1975; as Ditrichum rufescens, Matteri 1985b; Hässel et al. 1984; as Ditrichum stenostomum,
Cardot 1908; Roivainen 1937). Dist. VII. Cauquenes, Linares; VIII. Arauco, Biobío,
Concepción, Ñuble; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue,
Osorno, Palena; XI. Aisén; XII. Magallanes, Tierra del Fuego, Última Esperanza; Juan
Fernández Is.; Easter I.
Ditrichum ditrichoideum (Cardot) Ochyra
Ochyra et al. 2008 (as Philibertiella ditrichoidea, Hässel et al. 1984; Cardot & Brotherus 1923;
Herzog & Hosseus 1938; Matteri 1985b; as Ditrichum gemmiferum, Ochyra & Lewis Smith
1998; Ochyra 1999a; Ochyra & Deguchi 1999). Dist. XIV. Valdivia/Ranco; X. Chiloé,
Llanquihue, Osorno; XI. Aisén; XII. Magallanes, Última Esperanza.
Ditrichum elongatum (Hook.f. & Wilson) Mitt. = Ditrichum cylindricarpum fide Seppelt 1982.
Ditrichum flexifolium Hampe = Ditrichum difficile fide Magill 1981.
Ditrichum fontanum Herzog ≡ Anisothecium fontanum.
Ditrichum gemmiferum Ochyra & Lewis-Smith = Ditrichum ditrichoideum fide Ochyra et al. 2008.
Ditrichum hallei Cardot & Broth.
Barrera 2006; Cardot & Brotherus 1923; Roivainen 1934, 1937; Matteri & Schiavone 2002.
Dist. VIII. Ñuble; XII. Tierra del Fuego.
Ditrichum heteromallum (Hedw.) E. Britton
Hässel et al. 1984; Ochyra 1999c; Matteri 1985b; Deguchi 1991a; Matteri & Schiavone 2002
(as Ditrichum homomallum, Cardot & Brotherus 1923; as Ditrichum homomallum var.
leptocladum, Cardot & Brotherus 1923; as Dicranella fuegiana, Buck 2002; Matteri &
Schiavone 2002). Dist. XI. Aisén; XII. Antártica Chilena, Tierra del Fuego, Última Esperanza.
Ditrichum homomallum (Hedw.) Hampe = Ditrichum heteromallum fide Matsui & Iwatsuki 1990.
Ditrichum homomallum var. leptocladum Cardot & Broth. = Ditrichum heteromallum fide Ochyra 1999c.
Ditrichum hookeri (Müll. Hal.) Hampe
Tropicos 2009; Matteri & Schiavone 2002; Dusén 1903b, 1905b; Cardot 1908; Cardot &
Brotherus 1923; Roivainen 1937; Herzog 1954; Seki 1974; Deguchi 1991a (as Blindia
leptotrichocarpa, Dusén 1903b; as Chorisodontium sericeum, Hässel et al. 1984; Bartram
1946; Matteri 1985b; as Cynontodium hookeri, Mitten 1869). Dist. IX. Arauco; X. Osorno; XI.
Ditrichum hyalinocuspidatum Cardot
Ochyra et al. 2008; Matteri & Schiavone 2002; Cardot & Brotherus 1923. Dist. XII.
Ditrichum hyalinum (Mitt.) Kuntze
Buck 2002; Schaumann 2005; Matteri & Schiavone 2002; Dusén 1903b; Cardot 1908; Herzog
1954; Seki 1974; Deguchi 1991a; Ochyra 1999a (as Cynontodium hyalinum, Mitten 1869; as
Ditrichum austro-georgicum, Deguchi 1991a; as Pseudodistichium austrogeorgicum,
Roivainen 1937; Matteri 1985b; Hässel et al. 1984). Dist. IX. Cautín; X. Llanquihue, Osorno;
XI. Aisén; XII. Antártica Chilena, Magallanes, Tierra del Fuego, Última Esperanza.
Ditrichum lewis-smithii Ochyra = Ditrichum hyalinocuspidatum fide Ochyra et al. 2008.
Ditrichum longisetum (Lorentz) Hampe
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Mosses of Chile 43
Espinosa 1941; Dusén 1903b, 1905b; Cardot 1908; Brotherus 1924b; Herzog & Schwabe
1939; Bartram 1957; Seki 1974; Robinson 1975. Dist. XI. Aisén; XII. Magallanes, Última
Ditrichum nitens (Herzog) Frank Müll.
(as Philibertiella nitens, Herzog 1960; Matteri 1985b; Hässel et al. 1984). Dist. XIV.
Valdivia/Ranco; X. Osorno; XII. Magallanes, Última Esperanza.
Ditrichum praealtum (Mitt.) Kuntze ≡ Platyneuron praealtum.
Ditrichum roivaneni E.B. Bartram & Roiv.
Roivainen 1937; Matteri & Schiavone 2002. Dist. XII. Tierra del Fuego.
Ditrichum rufescens (Hampe) Hampe = Ditrichum difficile fide Allen 1994.
Ditrichum stenostomum Cardot = Ditrichum difficile fide Allen 1994.
Ditrichum strictum (Hook.f. & Wilson) Hampe
He 1998; Dusén 1903b; Cardot 1908. Dist. IX. Malleco.
Drepanocladus aduncus (Hedw.) Warnst.
Hässel et al. 1984; Matteri 1985b; Deguchi 1991a (as Hypnum aduncum, Cardot 1908; as
Drepanocladus aduncus var. kneiffii, He 1998; as Drepanocladus kneiffii, Deguchi 1991a; as
Drepanocladus polycarpos, Ochyra & Matteri 2001; Matteri & Schiavone 2002). Dist. RM.
Santiago; VIII. Biobío; X. Osorno, Palena; XII. Magallanes, Tierra del Fuego, Última
Esperanza.
Drepanocladus aduncus var. kneiffii (Schimp.) Mönk. = Drepanocladus aduncus fide Hedenäs 2003.
Drepanocladus austrostramineus (Müll. Hal.) Broth. ex Paris = Warnstorfia fontinaliopsis fide Ochyra & Matteri
2001.
Drepanocladus cossonii (Schimp.) Loeske ≡ Scorpidium cossonii.
Drepanocladus exannulatus (Schimp.) Warnst. ≡ Warnstorfia exannulata.
Drepanocladus exannulatus var. falcifolius (Renauld) Grout
(as Drepanocladus fluitans var. falcifolius, Thériot 1924). Dist. RM. Maipo.
Drepanocladus fluitans (Hedw.) Warnst. ≡ Warnstorfia fluitans.
Drepanocladus fluitans var. australe (Cardot) Broth. = Drepanocladus longifolius fide Ochyra & Matteri 2001.
Drepanocladus fluitans var. falcifolius (Renauld) Thér. ≡ Drepanocladus exannulatus var. falcifolius fide Grout 1929.
Drepanocladus fuegianus (Mitt.) Broth. ex Paris = Warnstorfia fluitans fide Ochyra & Matteri 2001.
Drepanocladus kneiffii (Schimp.) Warnst. var. chilense Thér.
Thériot 1924. Dist. RM. Maipo.
Drepanocladus laculosus (Müll. Hal.) Broth. = Warnstorfia fontinaliopsis fide Ochyra & Matteri 2001.
Drepanocladus longifolius (Mitt.) Broth. ex Paris
Müller 2009; Buck 2002; Vanderpoorten et al. 2003; Ochyra & Matteri 2001; Matteri &
Schiavone 2002; Cardot & Brotherus 1923; Roivainen 1934; Bartram 1952 (as Cratoneuron
perplicatum, Brotherus 1909; as Hypnum longifolium, Dusén 1903b; Cardot 1908). Dist. RM.
Cordillera; VII. Talca; XI. Aisén, Coihaique; XII. Antártica Chilena, Magallanes, Tierra del
Fuego.
Drepanocladus perplicatus (Dusén) G. Roth
Hedenäs 2003 (as Richardsiopsis lacustris, Ochyra & Mahu 1988). Dist. XV. Parinacota.
Drepanocladus polycarpos (Blandow ex Voit) Warnst. = Drepanocladus aduncus fide Hedenäs 2003.
Drepanocladus revolvens (Sw. ex anon.) Warnst. ≡ Scorpidium revolvens.
Drepanocladus secundifolius (Müll. Hal.) Dixon
Matteri & Schiavone 2002 (as Rhaphidostegium secundifolium, Dusén 1903b; Cardot 1908; as
Sematophyllum secundifolium, Mitten 1869). Dist. XII. Antártica Chilena. Note: The generic
position of this species is uncertain.
Drepanocladus sordidus (Müll. Hal.) Hedenäs
Hedenäs 2003. Dist. III. Copiapó.
Drepanocladus symmetricus (Renauld & Cardot) Cardot ≡ Sanionia symmetrica.
Drepanocladus uncinatus (Hedw.) Warnst. ≡ Sanionia uncinata.
Drepanocladus uncinatus var. plumulosus (Schimp.) Warnst.
Cardot & Brotherus 1923. Dist. XI. Aisén.
Drepanocladus uncinatus var. stenocarpus Cardot & Broth. = Sanionia uncinata fide Ochyra & Matteri 2001.
Drepanocladus uncinatus var. symmetricus (Renauld & Cardot) Grout ≡ Sanionia symmetrica.

44 Müller
Drummondia obtusifolia Müll. Hal.

Tropicos 2009; He 1998; Mitten 1869; Herzog & Hosseus 1938; Herzog 1954, 1960. Dist. VII.
Linares; VIII. Ñuble; IX. Cautín; X. Osorno.
Duseniella genuflexa (Müll. Hal.) Broth. ≡ Ancistrodes genuflexa.
Ectropothecium berberidis Dusén ≡ Rhaphidorrhynchium berberidis.
Ectropothecium spirifolium Dusén ≡ Vesiculariopsis spirifolium.
Encalypta armata Broth. ex Dusén
Müller 2009 (as Encalypta berthoana, Thériot 1923). Dist. IV. Limarí; RM. Santiago; XI.
General Carrera.
Encalypta asperifolia Mitt.
Tropicos. Dist. XV. Parinacota (Moreno 12981, 12980, 12982, MO, det. Robinson).
Encalypta australis Mitt. = Encalypta vulgaris fide Dixon 1926.
Encalypta austro-ciliata Broth. = Encalypta ciliata fide Horton 1983.
Encalypta berthoana Thér. = Encalypta armata fide Horton 1983.
Encalypta ciliata Hedw.
Buck 2002; Matteri & Schiavone 2002; Deguchi 1991a (as Encalypta austro-ciliata, Hässel et
al. 1984; Dusén 1906a; Cardot & Brotherus 1923; Matteri 1985b). Dist. VIII. Biobío; XI.
General Carrera; XII. Antártica Chilena, Magallanes, Última Esperanza.
Encalypta obtusata Cardot & Broth. = Encalypta vulgaris fide Müller 2009.
Encalypta patagonica Broth. = Encalypta rhaptocarpa fide Ochyra 1998.
Encalypta procera Bruch
Müller 2009. Dist. XI. General Carrera.
Encalypta rhaptocarpa Schwägr.
Matteri & Schiavone 2002; Deguchi 1991a (as Encalypta patagonica, Dusén 1903b, 1905a,
1906a; Cardot 1908; Matteri 1985b; Hässel et al. 1984). Dist. XII. Magallanes, Tierra del
Encalypta vulgaris Hedw.
(as Encalypta australis, Mitten 1869; as Encalypta obtusata, Matteri 1985b; Hässel et al.
1984). Dist. X. Chiloé; XII. Magallanes, Última Esperanza.
Entodon truncatus (Müll. Hal.) Mitt.
Mitten 1869. Dist. Without exact locality.
Entosthodon apophysatus (Taylor) Mitt.
Ireland et al. 2007. Dist. VII. Talca; VIII. Biobío, Concepción.
Entosthodon bonplandii (Hook.) Mitt.
Fife 1987 (as Funaria bonplandii, Thériot 1927b). Dist. Without exact locality.
Entosthodon chiloensis Mitt.
Mitten 1869 (as Funaria chiloensis, Thériot 1927b). Dist. X. Chiloé.
Entosthodon laevis (Mitt.) Fife
Ireland et al. 2007; He 1998; Fife 1987 (as Funaria berteroana, Herzog 1922, 1927; as
Funaria suberecta, Williams 1915; Thériot 1927b). Dist. IV. Choapa, Limarí; V. Valparaíso;
RM. Santiago; VIII. Concepción; XIV. Valdivia/Ranco.
Entosthodon laxus (Hook.f. & Wilson) Mitt.
Matteri & Schiavone 2002; Fife 1987; Deguchi 1991a (as Funaria cavifolia, Cardot &
Brotherus 1923; as Funaria pungens, Hässel et al. 1984; Bartram 1946; Matteri 1985b). Dist.
XI. Aisén; XII. Magallanes, Tierra del Fuego, Última Esperanza.
Entosthodon mathewsii var. integer Müll. Hal.
He 1998; Fife 1987 (as Funaria integra, Thériot 1927b; Herzog 1954). Dist. IV. Limarí; V.
Quillota, Valparaíso; RM. Chacabuco, Santiago.
Entosthodon obtusifolius Hook.f.
Cuvertino & Buffa 2009; Tropicos 2009; He 1998; Fife 1987 (as Funaria acutifolia, Thériot
1927b). Dist. VIII. Arauco, Concepción; XIV. Valdivia/Ranco; X. Llanquihue.
Ephemerum argentinicum Schiavone & Sarmiento
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Mosses of Chile 45
He 1998. Dist. IV. Limarí.

Eriodon conostomus Mont.
Tropicos 2009; Larraín 2007b; Larraín 2007a; He 1998; Dusén 1903b; Cardot 1908; Thériot
1918; Herzog 1922, 1954; Gunckel 1965; Reimers 1926; Bartram 1952 (as Hypnum
conostomum, Mitten 1869). Dist. VIII. Arauco, Biobío; IX. Cautín, Malleco; XIV.
Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno; XI. Aisén.
Eriopus apiculatus (Hook.f. & Wilson) Mitt. ≡ Calyptrochaeta apiculata.
Eriopus apiculatus var. spathulatus Matteri ≡ Calyptrochaeta apiculata var. spathulata.
Eriopus bescherellei Thér. = Calyptrochaeta mollis fide Matteri 1998.
Eriopus flexicollis (Mitt.) A. Jaeger ≡ Calyptrochaeta flexicollis.
Eriopus grandiretis Broth. ≡ Calyptrochaeta grandiretis.
Eriopus leptoloma Broth. ≡ Calyptrochaeta leptoloma.
Eriopus odontoloma Dusén ex Matteri ≡ Calyptrochaeta odontoloma.
Erythrophyllopsis fuscula (Müll. Hal.) Hilp.
Eucamptodon perichaetialis (Mont.) Mont.
Tropicos 2009; Müller 2009; Larraín 2007b; Ochyra 1994; Hässel et al. 1984; Buck 2004;
Larraín 2005; Ireland et al. 2007; Villagran et al. 2003; He 1998; Matteri & Schiavone 2002;
Mitten 1869; Dusén 1905c; Cardot 1908; Herzog 1922; Brotherus 1924b; Herzog & Hosseus
1938; Seki 1974; Robinson 1975; Matteri 1985b; Allen 1987a; Deguchi 1991a. Dist. VII.
Cauquenes, Talca; VIII. Arauco, Biobío, Concepción, Ñuble; IX. Cautín, Malleco; XIV.
Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno; XI. Aisén, Coihaique; XII. Magallanes, Tierra
Eucladium verticillatum (Brid.) Bruch & Schimp.
Roivainen 1934. Dist. VIII. Ñuble.
Eurhynchiella acanthophylla (Mont.) M. Fleisch.
Tropicos 2009; Pereira et al. 2006; Larraín 2005; Schaumann 2005; He 1998; Müller & Pereira
2006; Herzog & Hosseus 1938; Herzog & Schwabe 1939; Herzog 1954, 1960; Seki 1974;
Deguchi 1991a (as Hypnum acanthophyllum, Mitten 1869; as Rhynchostegiella acanthophylla,
Espinosa 1941; Thériot 1915, 1918; Brotherus 1924a; Herzog 1922). Dist. IV. Choapa; V.
Petorca, Quillota, San Antonio, Valparaíso; RM. Cordillera, Santiago; VI. Cardenal Caro; VII.
Cauquenes, Curicó, Linares, Talca; VIII. Arauco, Biobío, Concepción, Ñuble; IX. Cautín,
Malleco; XIV. Valdivia/Ranco; X. Llanquihue, Osorno; XI. Aisén.
Eurhynchiella acanthophylla var. robusta (Thér.) Ochyra
(as Rhynchostegiella acanthophylla var. robusta, Thériot 1917). Dist. V. Valparaíso.
Eurhynchium confusum Thér. = Eurhynchium corralense fide Wijk et al. 1962.
Eurhynchium corralense (Lorentz) A. Jaeger
He 1998 (as Oxyrrhynchium corralense, Tropicos 2009; Reimers 1926; Thériot 1934a; Herzog
& Hosseus 1938; as Rhynchostegium subsquarrosum, Espinosa 1941; Herzog 1922; Brotherus
1924a; as Eurhynchium flavidulum, Ochyra 1987a; as Sciaromium flavidulum, Dusén 1905a;
Cardot 1908; as Eurhynchium confusum, He 1998; Thériot 1918). Dist. IV. Limarí; VIII.
Arauco, Biobío, Ñuble, Concepción; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X.
Llanquihue, Osorno; Fuegia australis.
Eurhynchium fuegianum Cardot
Müller 2009; Buck 2002; Matteri & Ochyra 1989; Matteri & Schiavone 2002; Cardot &
Brotherus 1923; Herzog & Hosseus 1938 (as Platyhypnidium fuegianum, Goffinet et al. 2006;
as Oxyrrhynchium fuegianum, Herzog & Schwabe 1939; Herzog 1954). Dist. VIII. Ñuble; IX.
Cautín, Malleco; X. Palena; XI. Aisén, General Carrera; XII. Antártica Chilena, Tierra del
Eurhynchium flavidulum (Dusén) Ochrya = Eurhynchium fuegianum fide Matteri & Ochyra 1989.
Eurhynchium praelongum (Hedw.) Schimp. = Kindbergia praelonga.
Eustichia longirostris (Brid.) Brid.

46 Müller
Tropicos 2009; Hässel et al. 1984; Barrera 2006; He 1998; Müller & Pereira 2006; Dusén
1903b; Robinson 1975; Matteri 1985b; Deguchi 1991a (as Eustichia poeppigii, Espinosa 1941;
Villagran et al. 2003; He 1998; Cardot 1908; Cardot & Brotherus 1923; Brotherus 1924b;
Reimers 1926; Roivainen 1934; Herzog & Hosseus 1938; Herzog & Schwabe 1939; Bartram
1952, 1957; Herzog 1954, 1960; as Diplostichum poeppigii, Thériot 1921). Dist. V. Petorca,
Valparaíso; RM. Santiago; VII. Cauquenes, Linares, Talca; VIII. Arauco, Biobío, Concepción,
Ñuble; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno, Palena; XI.
Eustichia poeppigii (Müll. Hal.) Paris = Eustichia longirostris fide Allen 2002.
Exodokidium donati Herzog = Bartramia robusta fide Matteri 1984b.
Fabronia andina Mitt. = Fabronia ciliaris var. wrightii fide Buck 1983.
Fabronia andina var. jaffueli Thér. = Fabronia jamesonii fide Buck 1983.
Fabronia andina var. chilensis Thér. = Fabronia jamesonii fide Buck 1983.
Fabronia apophysatula Müll. Hal. = Fabronia jamesonii fide Buck 1983.
Fabronia ciliaris (Brid.) Brid. var. ciliaris
Tropicos 2009; He 1998; Deguchi 1991a (as Fabronia latidens, Herzog 1960; as Fabronia
minutissima, Duby 1880). Dist. IV. Limarí; V. Petorca; VIII. Biobío, Concepción, Ñuble; X.
Llanquihue, Osorno.
Fabronia ciliaris var. polycarpa (Hook.) W.R. Buck
He 1998. Dist. RM. Santiago.
Fabronia ciliaris var. wrightii (Sull.) W.R. Buck
He 1998 (as Fabronia andina, Mitten 1869; Herzog & Hosseus 1938; Herzog 1954; Matteri
1985b). Dist. IV. Limarí; V. Valparaíso; VIII. Biobío; IX. Cautín; X. Chiloé, Osorno; XII.
Magallanes.
Fabronia enervis Herzog ≡ Mahua enervis.
Fabronia jamesonii Taylor
Tropicos 2009; Müller 2009; Pereira et al. 2006; Larraín 2007a; Ireland & Bellolio 2002; He
1998; Buck 1983; Deguchi 1991a (as Fabronia andina var. jaffueli, Thériot 1924; as Fabronia
apophysatula, Herzog 1960; as Fabronia lorentziae, Herzog 1954; as Fabronia
macroblepharioides, Barrera 2006; Espinosa 1941; Brotherus 1924). Dist. V. Valparaíso; RM.
Santiago; VII. Linares, Talca; VIII. Arauco, Biobío, Concepción, Ñuble; XIV. Valdivia/Ranco;
X. Chiloé, Llanquihue, Osorno; XI. General Carrera; XII. Magallanes, Última Esperanza;
Easter I.
Fabronia latidens Duby = Fabronia ciliaris var. ciliaris fide Buck 1983.
Fabronia lorentziae Müll. Hal. = Fabronia jamesonii fide Buck 1983.
Fabronia macroblepharioides Broth. = Fabronia jamesonii fide Buck 1983.
Fabronia minutissima Duby = Fabronia ciliaris var. ciliaris fide Buck 1983.
Fallaciella gracilis (Hook.f. & Wilson) H. A. Crum
Tangney & Fife 2003; Matteri & Schiavone 2002 (as Camptochaete gracilis, Hässel et al.
1984; Matteri 1985b; as Isothecium serpens, Dusén 1903b). Dist. XIV. Valdivia/Ranco; XI.
Aisén; XII. Magallanes, Última Esperanza.
Felipponea hollermayeri Thér. = Acroschisma wilsonii fide Müller 2008.
Fissidens adianthoides Hedw.
Dusén 1906a; Matteri & Schiavone 2002; Müller & Pursell 2003. Dist. XII. Magallanes,
Última Esperanza, Tierra del Fuego.
Fissidens asplenioides Hedw.
Tropicos 2009; Larraín 2005; Villagran et al. 2003; Schaumann 2005; He 1998; Müller &
Pereira 2006; Mitten 1869; Brotherus 1924b; Herzog & Hosseus 1938; Herzog & Schwabe
1939; Bartram 1952; Herzog 1954; Robinson 1975; Müller & Pursell 2003 (as Fissidens
maschalanthus var. minor, Thériot 1918; as Fissidens pycnotylus, Brotherus 1924b). Dist. V.
Valparaíso; VII. Cauquenes; VIII. Arauco, Biobío, Concepción, Ñuble; IX. Cautín, Malleco;
XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno, Palena; XI. Aisén; XII. Magallanes;
Juan Fernández Is.
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Mosses of Chile 47
Fissidens asplenioides var. australis Cardot = Fissidens maschalanthus fide Müller & Pursell 2003.
Fissidens australiensis A. Jaeger = Fissidens serratus var. serratus fide Bruggenan-Nannenga & Pursell 1995.
Fissidens berteroi (Mont.) Müll. Hal.
Mitten 1869; Dusén 1906a; Müller & Pursell 2003 (as Fissidens parvus, Mitten 1869). Dist. V.
Aconcagua, Quillota, Valparaíso; X. Chiloé.
Fissidens brotherianus Paris = Fissidens curvatus fide Pursell 1994.
Fissidens brotherianus var. longisetus Thér. = Fissidens curvatus fide Pursell 1994.
Fissidens brotherii Dusén = Fissidens curvatus fide Pursell 1994.
Fissidens bryoides Hedw. var. pusillus (Wilson) Pursell
Ireland et al. 2007; He 1998; Müller & Pursell 2003. Dist. V. San Antonio; VIII. Arauco,
Concepción.
Fissidens bryoides var. viridulus (Sw.) Broth.
Ireland et al. 2007; He 1998; Müller & Pursell 2003. Dist. RM. Cordillera; VII. Talca; VIII.
Arauco, Concepción.
Fissidens campylopus Mont. = Fissidens oblongifolius fide Pursell 1994.
Fissidens chilensis Dusén = Fissidens crispus fide Pursell 1994.
Fissidens costesii Thér. = Fissidens crispus fide Pursell 1994.
Fissidens crassicuspes Broth. = Fissidens curvatus fide Pursell 1994.
Fissidens crispus Mont.
Tropicos 2009; He 1998; Müller & Pereira 2006; Müller & Pursell 2003 (as Fissidens
chilensis, Dusén 1906a; Thériot 1921; Herzog & Hosseus 1938; Herzog 1954; as Fissidens
costesii, Thériot 1921; as Fissidens fernandezianus, Brotherus 1924b; Bartram 1957; Robinson
1975; as Fissidens subaloma, Espinosa 1941; Dusén 1906a; Thériot 1921, 1924; Brotherus
1924a). Dist. V. Petorca, Quillota, San Antonio, Valparaíso; RM. Cordillera, Melipilla,
Santiago, Talagante; VI. Cachapoal, Colchagua; VII. Cauquenes, Talca; VIII. Arauco, Biobío,
Concepción; IX. Cautín; XIV. Valdivia/Ranco; X. Llanquihue, Osorno; Juan Fernández Is.
Fissidens curvatus Hornsch.
Tropicos 2009; Larraín 2007b; Larraín 2007a; He 1998 (as Fissidens brotherianus, Thériot
1921, 1924; Herzog & Hosseus 1938; Bartram 1952; Herzog 1954; as Fissidens brotherianus
var. longisetus, Thériot 1921; as Fissidens brotherii, Dusén 1906a; as Fissidens crassicuspes,
Brotherus 1924b; Robinson 1975; as Fissidens diversiformis, Barrera 2001; Espinosa 1941;
Brotherus 1924a; as Fissidens valdiviensis, Herzog 1922). Dist. II. El Loa; III. Copiapó; IV.
Choapa, Limarí; V. Aconcagua, Petorca, Quillota, San Antonio, Valparaíso; RM. Chacabuco,
Cordillera, Maipo, Santiago, Talagante; VI. Cachapoal, Cardenal Caro, Colchagua; VII. Talca;
VIII. Biobío, Concepción, Ñuble; IX. Cautín; XIV. Valdivia/Ranco; X. Chiloé, Osorno; XI.
Aisén; Juan Fernández Is.
Fissidens dillenii Müll. Hal. = Fissidens semicompletus fide Purselll 1994.
Fissidens diversiformis Broth. = Fissidens curvatus fide Pursell 1994.
Fissidens echinellus Herzog = Fissidens serratus var. serratus fide Pursell 1994.
Fissidens excurrentinervis R.S. Williams
Cano 2003. Dist. II. El Loa.
Fissidens fernandezianus Broth. = Fissidens crispus fide Pursell 1994.
Fissidens lechleri Hampe ex Müll. Hal. = Fissidens rigidulus fide Müller & Pursell 2003.
Fissidens leptochaete Dusén ≡ Fissidens serratus var. leptochaete.
Fissidens longidecurrens Thér. = Fissidens crispus fide Müller & Pursell 2003.
Fissidens maschalanthus Mont.
Espinosa 1941; Schaumann 2005; Mitten 1869; Dusén 1906a; Cardot 1908; Thériot 1921;
Brotherus 1924b; Reimers 1926; Herzog & Hosseus 1938; Herzog & Schwabe 1939; Herzog
1954, 1960; Bartram 1957; Robinson 1975; Müller & Pursell 2003. Dist. VIII. Arauco; IX.
Cautín, Malleco; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno, Palena; XI. Aisén;
Juan Fernández Is.
Fissidens maschalanthus var. minor Dusén = Fissidens asplenioides fide Pursell 1994.
Fissidens oblongifolius Hook.f. & Wilson

48 Müller
Tropicos 2009; Larraín 2007b; Larraín 2007a; He 1998; Müller & Pereira 2006; Müller &
Pursell 2003 (as Fissidens campylopus, Mitten 1869; Dusén 1906a). Dist. VII. Cauquenes;
VIII. Arauco, Biobío, Concepción, Ñuble; IX. Cautín; XIV. Valdivia/Ranco; X. Chiloé,
Llanquihue.
Fissidens ornaticostatus H. Rob.
Robinson 1975; Müller & Pursell 2003. Dist. Juan Fernández Is.
Fissidens papillosus Sande Lac. = Fissidens serratus var. serratus fide Bruggeman-Nannenga & Pursell 1995.
Fissidens parvus Mitt., hom. illeg. = Fissidens berteroi fide Pursell 1994.
Fissidens pascuanus Broth.
Barrera 2006; Espinosa 1941; Ireland & Bellolio 2002; Müller & Pursell 2003; Brotherus
1924. Dist. Easter I.
Fissidens pellucidus Hornsch.
Ireland & Bellolio 2002; Müller & Pursell 2003. Dist. Easter I.
Fissidens pycnotylus Broth. = Fissidens asplenioides fide Robinson 1975.
Fissidens rigidulus Hook.f. & Wilson
Goffinet et al. 2006; Hässel et al. 1984; Matcham et al. 2007; Pereira et al. 2006; Buck 2002;
Buck 2005a; Espinosa 1941; He 1998; Müller & Pereira 2006; Matteri & Schiavone 2002;
Mitten 1869; Dusén 1906a; Cardot 1908; Thériot 1921, 1925; Brotherus 1924b; Reimers 1926;
Roivainen 1934, 1937; Herzog & Schwabe 1939; Herzog 1954, 1960; Bartram 1957; Robinson
1975; Matteri 1985b; Müller & Pursell 2003. Dist. V. Petorca, San Antonio, Valparaíso; RM.
Cordillera, Santiago, Talagante; VI. Colchagua; VII. Cauquenes, Curicó, Talca; VIII. Arauco,
Biobío, Concepción, Ñuble; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Llanquihue,
Osorno; XI. Aisén; XII. Antártica Chilena, Magallanes, Última Esperanza; Juan Fernández Is.
Fissidens rigidulus subsp. masatierrensis Brugg.-Nann. = Fissidens rigidulus fide Müller & Pursell 2003.
Fissidens scalaris Mitt.
Espinosa 1941; He 1998; Mitten 1869; Thériot 1921; Brotherus 1924a; Herzog & Hosseus
1938; Bartram 1952; Herzog 1954; Müller & Pursell 2003 (as Fissidens tener, Dusén 1906a).
Dist. IV. ? Andes Chilensis; V. Petorca, San Antonio, Valparaíso; RM. Chacabuco, Cordillera,
Santiago; VIII. Biobío, Concepción; IX. Cautín, Malleco; XIV. Valdivia/Ranco.
Fissidens semicompletus Hedw.
Mitten 1869; Müller & Pursell 2003 (as Fissidens dillenii, Dusén 1906a; as Octodiceras
dillenii, Cardot 1908). Dist. X. Llanquihue.
Fissidens serratus Müll. Hal. var. leptochaete (Dusén) Brugg.-Nann. & Pursell
Tropicos 2009; Bruggeman-Nannenga & Pursell 1995; Frey & Schaumann 2002; Müller &
Pursell 2003 (as Fissidens leptochaete, Dusén 1906a; Brotherus 1924b; Robinson 1975). Dist.
VIII. Biobío, Concepción; X. Osorno, Palena; Juan Fernández Is.
Fissidens serratus Müll. Hal. var. serratus
Larraín 2007b; Larraín 2007a; Ireland et al. 2007; He 1998; Müller & Pursell 2003 (as
Fissidens australiensis, Stone 1994; as Fissidens echinellus, Herzog 1954; as Fissidens
papillosus, Pursell 1994). Dist. V. Aconcagua, Petorca, San Antonio, Valparaíso; VIII.
Concepción; IX. Cautín; X. Chiloé.
Fissidens subaloma Dusén = Fissidens crispus fide Pursell 1994.
Fissidens taxifolius Hedw.
Espinosa 1941; He 1998; Brotherus 1924a; Müller & Pursell 2003. Dist. VIII. Concepción.
Fissidens tener Dusén = Fissidens scalaris fide Pursell 1994.
Fissidens valdiviensis Herzog = Fissidens curvatus fide Pursell 1994.
Fissidens spec.
Mahu 1981. Dist. V. Valparaíso.
Florschuetzia pilmaiquen Crosby ≡ Diphyscium pilmaiquen.
Forsstroemia trichomitria (Hedw.) Lindb.
Rao & Enroth in Blockeel et al. 2003. Dist. IX. Malleco; X. Llanquihue.
Funaria acutifolia (Hampe) Broth. = Entosthodon obtusifolius fide Fife 1987.
Funaria balansae (Besch.) Broth. ≡ Entosthodon balansae.
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Funaria berteroana Hampe = Entosthodon laevis fide Fife 1987.

Funaria bonplandii (Brid.) Broth. ≡ Entosthodon bonplandii.
Funaria calvescens Schwägr. = Funaria hygrometrica fide Ochi 1968.
Funaria cavifolia Cardot & Broth. = Entosthodon laxus fide Fife 1987.
Funaria chilensis (Thér.) Thér.
Cuvertino & Buffa 2009; He 1998; Thériot 1926, 1927 (as Funaria clavellata var. chilensis,
Thériot 1924). Dist. V. Petorca, San Antonio, Valparaíso; VIII. Concepción, Ñuble.
Funaria chiloensis (Mitt.) Broth. ≡ Entosthodon chiloensis.
Funaria clavellata (Mitt.) Broth. var. chilensis Thér. = Funaria chilensis.
Funaria commixta Thér.
He 1998; Thériot 1926, 1927; Herzog 1954; Villagran et al. 2004. Dist. IV. Limarí; V. Petorca,
San Antonio; VIII. Concepción; X. Llanquihue.
Funaria costesii Thér.
He 1998; Thériot 1921, 1927; Villagran et al. 2004. Dist. IV. Limarí; V. Petorca, San Antonio,
Valparaíso; RM. Santiago.
Funaria fuegiana Müll. Hal. ≡ Funaria hygrometrica var. fuegiana.
Funaria hygrometrica Hedw. var. fuegiana (Müll. Hal.) Besch.
Buck 2002; Matteri & Schiavone 2002; Thériot 1927b; Deguchi 1991a (as Funaria fuegiana,
Cardot 1908). Dist. IX. Cautín; XII. Antártica Chilena, Magallanes.
Funaria hygrometrica Hedw. var. hygrometrica
Tropicos 2009; Goffinet et al. 2006; Larraín 2007b; Hässel et al. 1984; Larraín 2005; Espinosa
1941; Villagran et al. 2003; He 1998; Müller & Pereira 2006; Matteri & Schiavone 2002;
Mitten 1869; Dusén 1903b; Cardot 1908; Williams 1915; Thériot 1915, 1917, 1918, 1921,
1927; Brotherus 1924a, 1924b; Reimers 1926; Roivainen 1934; Herzog & Hosseus 1938;
Herzog & Schwabe 1939; Bartram 1952; Herzog 1954; Seki 1974; Robinson 1975; Mahu
1981; Matteri 1985b; Deguchi 1991a; Cardot 1905 (as Funaria calvescens, Herzog 1954; as
Funaria hygrometrica var. calvescens, Thériot 1927b; as Funaria hygrometrica var. crispa,
Thériot 1927b). Dist. IV. Choapa, Elqui, Limarí; V. Petorca, Quillota, San Antonio,
Valparaíso; RM. Chacabuco, Cordillera, Maipo, Melipilla, Santiago, Talagante; VI.
Colchagua; VII. Cauquenes, Curicó, Linares, Talca; VIII. Arauco, Biobío, Concepción, Ñuble;
IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno; XI. Aisén; XII.
Antártica Chilena, Magallanes, Última Esperanza; Juan Fernández Is.
Funaria hygrometrica var. calvescens (Schwägr.) Schimp. = Funaria hygrometrica fide Enroth 1991.
Funaria hygrometrica var. crispa (Müll. Hal.) Paris = Funaria hygrometrica fide Enroth 1991.
Funaria integra (Müll. Hal.) Broth. ≡ Entosthodon mathewsii var. integer.
Funaria integra (Müll. Hal.) Broth. var. serrulata Broth.
Brotherus 1924a; Espinosa 1941. Dist. IV. Coquimbo, Limarí.
Funaria macrocarpa (Schimp.) R.H. Zander ≡ Costesia macrocarpa.
Funaria porteri Thér.
Thériot 1928. Dist. VII. Linares.
Funaria pungens E.B. Bartram = Entosthodon laxus fide Fife 1987.
Funaria suberecta Mitt. = Entosthodon laevis fide Fife 1987.
Gemmabryum acuminatum (Harv.) J.R. Spence & H.P. Ramsay ≡ Brachymenium acuminatum.
Gertrudiella validinervis (Herzog) Broth.
Tropicos 2009. Dist. II. Antofagasta (Mahú 13150, MO, det. Zander).
Glyphomitrium ligulatum Mitt. ≡ Ptychomitrium ligulatum.
Glyphomitrium fernandesianum Mitt. ≡ Ptychomitrium fernandesianum.
Glyphothecium gracile (Hampe) Broth.
Tropicos 2009; Larraín 2007a; Larraín 2005; He 1998; Müller & Pereira 2006; Mitten 1869;
Thériot 1915; Reimers 1926; Herzog & Hosseus 1938; Herzog 1954, 1960; Seki 1974;
Hattaway 1981, 1984; Deguchi 1991b; Kühnemann & Goncalves Carralves 1975 (as
Cladomnion gracile, Mitten 1869; Dusén 1903b; as Glyphothecium gracile var. nanum,
Herzog 1922). Dist. V. Valparaíso; VI. Cachapoal, Colchagua; VII. Cauquenes, Curicó,

50 Müller
Linares, Talca; VIII. Arauco, Biobío, Concepción, Ñuble; IX. Cautín, Malleco; XIV.
Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno, Palena; XI. Aisén.
Glyphothecium gracile var. nanum Herzog = Glyphothecium gracile fide Hattaway 1984.
Glyphothecium sciuroides (Hook.) Hampe
Larraín 2007b; Schaumann 2005; Hattaway 1981, 1984; Deguchi 1991b (as Cladomnion
valdivae, Müller 1897). Dist. XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno; XI. Aisén.
Goniobryum reticulatum (Hook.f & Wilson) Lindb. = Philonotis vagans fide Matteri & Schiavone 2002.
Goniobryum subbasilare (Hook.) Lindb.
Matteri & Piñero 1987; Hässel et al. 1984; Matcham et al. 2007; Buck 2002; Matteri &
Schiavone 2002; Cardot 1908; Cardot & Brotherus 1923; Matteri 1985b (as Rhizogonium
subbasilare, Mitten 1869; Dusén 1903b). Dist. XII. Antártica Chilena, Magallanes, Tierra del
Grimmia abscondita Cardot = Schistidium praemorsum fide Ochyra et al. 2008.
Grimmia alpicola Hedw. var. rivularis (Brid.) Broth. ≡ Schistidium rivulare.
Grimmia amblyophylla Müll. Hal. ≡ Schistidium amblyophyllum.
Grimmia anodon Bruch & Schimp.
Tropicos 2009; Muñoz 1999; He 1998; Deguchi 1984. Dist. RM. Santiago. XII. Antártica
Chilena, Última Esperanza.
Grimmia apocarpa Hedw. ≡ Schistidium apocarpum.
Grimmia arachnoidea Dusén, nom. nud. = Grimmia pulvinata fide Muñoz & Pando 2000.
Grimmia austrofunalis Müll. Hal.
Tropicos 2009; Muñoz & Pando 2000; Muñoz 1999. Dist. VI. Cachapoal; VII. Talca; VIII.
Biobío, Ñuble; IX. Cautín; X. Llanquihue, Palena; Aisén.
Grimmia chilensis Thér. ex Herzog = Grimmia navicularis fide Deguchi 1987.
Grimmia consobrina Kunz. ex Müll. Hal. = Grimmia trichophylla fide Deguchi 1984.
Grimmia crassiretis Cardot & Broth. = Grimmia austrofunalis fide Muñoz & Pando 2000.
Grimmia depressa Müll.Hal. = Bucklandiella didyma fide Matteri & Schiavone 2002.
Grimmia depressa var. terrestris Müll. Hal. = Bucklandiella didyma fide Matteri & Schiavone 2002.
Grimmia dicksonii (Dusén) Dusén, hom. illeg. = Grimmia trichophylla fide Deguchi 1984.
Grimmia donatii Herzog & Thér. = Schistidium amblyophyllum fide Ochyra et al. 2008.
Grimmia fallax Dusén = Schistidium andinum fide Ochyra et al. 2008.
Grimmia fasciculata Dusén, hom. illeg. = Schistidium falcatum fide Deguchi 1984.
Grimmia fastigiata Cardot = Grimmia fuscolutea fide Matteri & Schiavone 2002.
Grimmia flexicaulis Müll. Hal. var. dicksonii Dusén = Grimmia trichophylla fide Deguchi 1984.
Grimmia flexiseta Dusén, nom. nud. = Grimmia trichophylla fide Muñoz 1999.
Grimmia fuscolutea Hook.
Matteri & Schiavone 2002 (as Dryptodon fuscoluteus, Ochyra et al. 2008; as Grimmia
fastigiata, Cardot & Brotherus 1923). Dist. XII. Tierra del Fuego, Última Esperanza.
Grimmia grisea Cardot = Grimmia reflexidens fide Muñoz & Pando 2000.
Grimmia gusindei J. Froehl. = Grimmia trichophylla fide Muñoz 1999.
Grimmia humilis Mitt.
Matcham et al. 2007; Muñoz & Pando 2000; Muñoz 1999; Matteri & Schiavone 2002; Mitten
1869; Dusén 1903b; Cardot 1908; Bartram 1952; Deguchi 1984. Dist. RM. Santiago; XI.
Grimmia hyalinocuspidata Müll. Hal. = Schistidium amblyophyllum fide Ochyra 1998.
Grimmia hyalinocuspidata var. mutica Cardot & Broth. = Schistidium amblyophyllum fide Ochyra et al. 2008.
Grimmia imberbis Kunze ex Müll. Hal. = Grimmia trichophylla fide Muñoz 1999.
Grimmia kidderi James
Buck 2002; Muñoz & Pando 2000; Muñoz 1999. Dist. IX. Malleco; XII. Antártica Chilena.
Grimmia laevigata (Brid.) Brid.
Tropicos 2009; Muñoz & Pando 2000; Muñoz 1999; He 1998; Deguchi 1984, 1991a (as
Grimmia murina, Dusén 1905a). Dist. IV. Choapa; V. Petorca; RM. Chacabuco, Santiago; VII.
Curicó; VIII. Biobío, Ñuble.
Grimmia longirostris Hook.
NY 2009; Muñoz & Pando 2000; Muñoz 1999; Matteri & Schiavone 2002. Dist. XV.
Parinacota; XII. Tierra del Fuego.
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Mosses of Chile 51
Grimmia macropulvinata Dusén = Grimmia trichophylla fide Deguchi 1984.

Grimmia molesta J. Muñoz
Grimmia murina Müll. Hal. = Grimmia laevigata fide Deguchi 1984.
Grimmia navicularis Herzog
He 1998; Deguchi 1991a (as Grimmia chilensis, Seki 1974; Deguchi 1984). Dist. V. Petorca;
RM. Chacabuco, Cordillera, Santiago; VII. Linares; VIII. Biobío; IX. Cautín, Malleco; XIV.
Valdivia/Ranco; X. Osorno; XI. Aisén.
Grimmia nivea Dusén = Grimmia plagiopodia fide Deguchi 1984.
Grimmia occulta Müll. Hal. = Schistidium andinum fide Ochyra et al. 2008.
Grimmia orbicularis Bruch
Matteri & Schiavone 2002; Dusén 1905a, 1906b; Cardot 1908; Cardot & Brotherus 1923. Dist.
XI. Aisén; XII. Tierra del Fuego.
Grimmia orbicularis var. patagonica Cardot = Grimmia pulvinata fide Muñoz 1999.
Grimmia pachyloma Mont. ≡ Vittia pachyloma.
Grimmia pachyphylla Dusén = Grimmia trichophylla fide Deguchi 1984.
Grimmia perplexa Thér. ≡ Schistidium perplexum.
Grimmia phyllorhizans Broth. = Grimmia pulvinata fide Muñoz 1999.
Grimmia plagiopodia Hedw.
Muñoz & Pando 2000; Muñoz 1999; Deguchi 1984; Cano 2003 (as Grimmia nivea, Dusén
1905a, 1906b; Seki 1974). Dist. XV. Parinacota; XI. Aisén; XII. Última Esperanza.
Grimmia pseudoanodon Deguchi
Grimmia pulvinata (Hedw.) Sm.
Buck 2002; Muñoz & Pando 2000; Muñoz 1999 (as Grimmia orbicularis var. patagonica,
Cardot 1908; as Grimmia arachnoidea, Dusén 1903b; as Grimmia phyllorhizans, Barrera 2006;
Espinosa 1941; Brotherus 1924b; Robinson 1975; as Grimmia pulvinata var. africana, He
1998; Deguchi 1984). Dist. V. Aconcagua, Los Andes; RM. Chacabuco, Cordillera, Santiago;
VI. Colchagua; X. Llanquihue; XI. Aisén; XII. Antártica Chilena, Magallanes, Última
Grimmia pulvinata (Hedw.) var. africana (Hedw.) Hook.f. & Wilson = Grimmia pulvinata fide Muñoz & Pando 2000.
Grimmia pycnophylla Dusén, nom. nud. = Grimmia trichophylla fide Deguchi 1984.
Grimmia reflexidens Müll. Hal.
Tropicos 2009; Buck 2002; Muñoz & Pando 2000; Muñoz 1999; He 1998; Mitten 1869; Dusén
1906b; Deguchi 1984; Deguchi 1991a (as Grimmia flexiseta, Dusén 1903b; as Grimmia grisea,
Hässel et al. 1984; Thériot 1934b; Matteri 1985b). Dist. RM. Cordillera, Santiago; VI.
Colchagua; VIII. Biobío, Ñuble; IX. Cautín; X. Osorno; XI. Coihaique; XII. Antártica Chilena,
Grimmia scabripes E.B. Bartram ≡ Schistidium scabripes.
Grimmia serripila Dusén ex Paris, nom. nud. = Grimmia laevigata fide Muñoz 1999.
Grimmia sessitana De Not.
(as Orthogrimmia sessitana, Ochyra et al. 2008). Dist. XI. Aisén.
Grimmia tenuicaulis Hampe ex A. Jaeger = Grimmia longirostris fide Muñoz & Pando 2000.
Grimmia trichophylla Grev.
Tropicos 2009; Matcham et al. 2007; Buck 2002; Pereira et al. 2006; Muñoz & Pando 2000;
Muñoz 1999; He 1998; Müller & Pereira 2006; Matteri & Schiavone 2002; Deguchi 1984,
1991a (as Grimmia consobrina, Mitten 1869; Dusén 1906b; Thériot 1917, 1921, 1935;
1952; Herzog 1954; Seki 1974; as Grimmia crassiretis, Cardot & Brotherus 1923; as Grimmia
dicksonii, Dusén 1906b; Cardot 1908; as Grimmia flexicaulis var. dicksonii; as Grimmia
macropulvinata, Dusén 1903b, 1906b; as Grimmia pachyphylla, Dusén 1905a; as Grimmia
pycnophylla, Dusén 1903b; as Grimmia imberbis, Mitten 1869; Dusén 1906b; Herzog 1960; as
Grimmia gusindei, Froehlich 1953). Dist. IV. Choapa, Elqui, Limarí; V. Petorca, Quillota, San
Antonio, Valparaíso; RM. Chacabuco, Cordillera, Maipo, Santiago, Talagante; VI. Cachapoal,

52 Müller
Cardenal Caro, Colchagua; VII. Cauquenes, Curicó, Linares, Talca; VIII. Arauco, Biobío,
Osorno; XI. Aisén, General Carrera; XII. Antártica Chilena, Magallanes, Tierra del Fuego,
Última Esperanza.
Gymnostomum aeruginosum Sm.
Müller 2009; Matteri 1985b; Deguchi 1991b. Dist. VIII. Biobío, Ñuble; XIV. Valdivia/Ranco;
XI. General Carrera; Juan Fernández Is.
Gymnostomum antarcticum Ångström ≡ Hennediella antarctica.
Gymnostomum calcareum Nees & Hornsch. var. calcareum
Tropicos 2009; Ireland et al. 2007; Espinosa 1941; Dusén 1906a; Thériot 1921; Brotherus
1924b; Herzog & Hosseus 1938; Robinson 1975; Deguchi 1991b (as Weissia calcarea, Mitten
1869). Dist. V. Valparaíso; VI. Cachapoal; VIII. Biobío; XIV. Valdivia/Ranco; XI. Aisén,
General Carrera; Juan Fernández Is.
Gymnostomum calcareum var. tenellum Bruch & Schimp.
Herzog 1954. Dist. RM. Santiago.
Gymnostomum laxifolium Kunze ex Paris, hom. illeg.
Brotherus 1924. Dist. Chile, without exact locality.
Gymnostomum tenerrimum (Müll. Hal.) Wijk & Margad.
(as Anoectangium tenerrimum, Mitten 1869). Dist. VIII. Ñuble. Note: This species probably
belongs to Zygodon.
Haplodontium argentifolium (Mitt.) A. Jaeger ≡ Mielichhoferia argentifolia.
Haplohymenium longinerve (Broth.) Broth.
He 1998. Dist. V. Petorca; VII. Linares; VIII. Biobío.
Hebantia rigida (Lorentz) G. L. Merr.
Ireland et al. 2006; Smith Merrill 1996 (as Oligotrichum rigidum, Mullen & Frye 1947;
Villagran et al. 2003). Dist. VIII. Arauco, Concepción, Ñuble; IX. Cautín; XIV.
Valdivia/Ranco; X. Chiloé, Palena.
Hedwigia orbiculata Mitt. = Rhacocarpus purpurascens fide Frahm 1996.
Hedwigidium chilense M. Fleisch., nom. nud. (as Braunia chilensis, nom. nud., Brotherus 1920).
Hedwigidium imberbe (Sm.) Bruch & Schimp. = Hedwigidium integrifolium fide Wijk et al. 1962.
Hedwigidium integrifolium (P. Beauv.) Dixon
Tropicos 2009; He 1998; Deguchi 1991a (as Hedwigidium imberbe, Reimers 1926; Herzog &
Hosseus 1938; Bartram 1952). Dist. RM. Cordillera; VIII. Biobío, Ñuble; IX. Cautín, Malleco;
XIV. Valdivia/Ranco; X. Llanquihue.
Hennediella acutidentata (Thér. & Cardot) R.H. Zander = Hennediella antarctica fide Cano 2008.
Hennediella antarctica (Ångström) Ochyra & Matteri
Matcham et al. 2007; Buck 2002; Matteri & Schiavone 2002; Ochyra & Matteri 1996 (as
Pottia antarctica, Dusén 1903b; Dusén 1906a; as Pottia magellanica, Cardot 1908; as Pottia
heimii var. thaxteri, Thériot 1929b; as Hennediella heimii var. thaxteri, Zander 1993). Dist.
Hennediella arenae (Besch.) R.H. Zander var. arenae
Cano 2008; Matteri & Schiavone 2002; Zander 1993 (as Barbula arenae, Dusén 1903b; as
Tortula arenae, Cardot 1908; Herzog 1922; Roivainen 1934; Herzog & Schwabe 1939; as
Syntrichia lingulaefolia, Herzog 1954; as Tortula polycarpa, Ireland et al. 2007; Dusén 1906a;
Cardot 1908; Cardot & Brotherus 1923). Dist. VIII. Concepción, Ñuble; IX. Cautín; XIV.
Valdivia/Ranco; X. Osorno; XI. Aisén; XII. Magallanes, Tierra del Fuego.
Hennediella austrogeorgica (Cardot) Blockeel = Hennediella antarctica fide Cano 2008.
Hennediella austrogeorgica var. microphylla (Cardot & Broth.) R.H. Zander = Hennediella antarctica fide Cano 2008.
Hennediella bellii (E.B. Bartram) R.H. Zander
Cano 2008. Dist. II. El Loa.
Hennediella densifolia (Hook.f. & Wilson) R.H. Zander
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Mosses of Chile 53
Matcham et al. 2007; Buck 2002; Matteri & Schiavone 2002; Zander 1993 (as Barbula
densifolia, Dusén 1903b; as Tortula densifolia, Hässel et al. 1984; Cardot 1908; Herzog 1922;
Cardot & Brotherus 1923; Deguchi 1991b; as Neobarbula magellanica, Dusén 1905a, 1906a).
Dist. XII. Antártica Chilena, Magallanes, Tierra del Fuego, Última Esperanza.
Hennediella diaguita R.H. Zander & Mahu ≡ Tortula diaguita.
Hennediella heimii var. flavipes (Mont.) Warnst. = Hennediella heimii fide Cano 2008.
Hennediella heimii (Hedw.) R.H. Zander var. heimii
Buck 2002; Matteri & Schiavone 2002; Zander 1993 (as Pottia heimii, Dusén 1906b; Cardot
1908; as Tortula heimii, Mitten 1869; as Pottia flavipes, Bartram 1943; as Tortula flaviseta,
Mitten 1869; as Hennediella heimii var. magellanica, Zander 1993; as Pottia heimii var.
magellanica, Herzog 1922; as Hennediella heimii var. maxima, Matteri & Schiavone 2002; as
Pottia heimii var. maxima, Cardot 1905). Dist. XV. Parinacota; III. Copiapó, Huasco; IV.
Limarí; RM. Cordillera; XII. Antártica Chilena, Magallanes, Última Esperanza.
Hennediella heimii var. magellanica (Warnst.) R.H. Zander = Hennediella heimii fide Cano 2008.
Hennediella heimii var. maxima (Cardot) R.H. Zander = Hennediella heimii fide Cano 2008.
Hennediella heimii var. spegazzinii (Müll. Hal.) R.H. Zander
Matteri & Schiavone 2002; Zander 1993 (as Pottia spegazzinii, Dusén 1903b, 1906a; Cardot
1908). Dist. X. Llanquihue; XII. Tierra del Fuego. Note: Type material has not been studied by
Cano (2008), therefore the application of this name is in doubt.
Hennediella heimii var. thaxteri (Cardot & Thér.) R.H. Zander = Hennediella antarctica fide Cano 2008.
Hennediella kunzeana (Müll. Hal.) R.H. Zander
Müller 2009; Ireland et al. 2007 (as Tortula kunzeana, Mitten 1869, Williams 1915, Thériot
1921). Dist. IV. Choapa, Coquimbo, Limarí; V. Aconcagua, Petorca, Valparaíso; RM.
Cordillera, Santiago; VII. Cauquenes, Talca; VIII. Arauco, Biobío, Concepción, Ñuble; X.
Chiloé.
Hennediella serrulata (Hook. & Grev.) R.H. Zander ≡ Syntrichia serrulata.
Holodontium inerme (Mitt.) Broth. = Holodontium strictum fide Ochyra 1993a.
Holodontium pumilum (Mitt.) Broth. ≡ Kiaeria pumila.
Holodontium ramulosum (Mitt.) Broth.= Kiaeria pumila fide Ochyra 1993a.
Holodontium strictum (Hook.f. & Wilson) Ochyra
Müller 2009; Ochyra et al. 2008; Matteri & Schiavone 2002 (as Blindia auriculata, Dusén
1905b; as Dicranum inerme, Cardot & Brotherus 1923; as Holodontium inerme, Roivainen
1934, 1937; Herzog & Schwabe 1939). Dist. VII. Talca; VIII. Ñuble; XIV. Valdivia/Ranco; X.
Llanquihue; XI. Aisén; XII. Antártica Chilena, Magellanes, Tierra del Fuego, Última
Esperanza.
Hookeria scabriuscula Müll. Hal. ≡ Callicostella scabriuscula.
Hookeria sublimbata Müll. Hal. ≡ Cyclodictyon sublimbatum.
Hookeriopsis incurva (Hornsch.) Broth. ≡ Thamniopsis incurva.
Hygroamblystegium austrofluviatile (Müll. Hal.) Broth. = Cratoneuropsis relaxa subsp. minor fide Ochyra et al. 2008.
Hygroamblystegium chilense (Lorentz) Reim. = Cratoneuropsis relaxa subsp. minor fide Ochyra et al. 2008.
Hygroamblystegium crassicostatum E.B. Bartram, hom. illeg., Bartram 1943.
Hygroamblystegium filum (Müll. Hal.) Reim. = Cratoneuropsis relaxa subsp. minor fide Ochyra et al. 2008.
Hygroamblystegium fuegianum (Besch.) Reim. = Cratoneuropsis relaxa subsp. minor fide Ochyra et al. 2008.
Hygroamblystegium fuegianum var. compactum E.B. Bartram = Cratoneuropsis relaxa subsp. minor fide Ochyra et al.
2008.
Hygroamblystegium fuegianum var. excurrens (Cardot & Broth.) E.B. Bartram = Cratoneuropsis relaxa subsp. minor
fide Ochyra et al. 2008.
Hygroamblystegium fuegianum var. gracilis (Cardot & Broth.) E.B. Bartram = Cratoneuropsis relaxa subsp. minor
fide Ochyra et al. 2008.
Hygroamblystegium fuegianum var. secundum E.B. Bartram = Cratoneuropsis relaxa subsp. minor fide Ochyra et al.
2008.
Hygroamblystegium fuegianum var. skottsbergii (Cardot) E.B. Bartram = Cratoneuropsis relaxa subsp. minor fide
Ochyra et al. 2008.
Hygroamblystegium luridum (Cardot) Reim. = Cratoneuropsis relaxa subsp. minor fide Ochyra et al. 2008.
Hygroamblystegium tenellum (Cardot & Broth.) Reim. = Cratoneuropsis relaxa subsp. minor fide Ochyra et al. 2008.
Hygrohypnum peruviense R.S. Williams ≡ Rhynchostegium peruviense.
Hymenocleiston magellanicum Duby = Tayloria dubyi fide Brotherus 1903.

54 Müller
Hymenocleiston magellanicum var. edenense Besch. ≡ Tayloria dubyi var. edenensis.

Hymenoloma antarcticum (Müll. Hal.) Ochyra
(as Dicranoweisia breviseta, Matteri & Schiavone 2002; Cardot & Brotherus 1923; Roivainen
1937; Seki 1974; as Dicranoweisia brevipes, Ochyra 1998; as Dicranoweisia antarctica,
Hässel et al. 1984; Matteri & Schiavone 2002; Cardot 1908; Cardot & Brotherus 1923;
Roivainen 1937; Herzog & Schwabe 1939; Roivainen 1937; Bartram 1952; Herzog 1954; Seki
1974; Matteri 1985b; as Dicranum antarcticum, Mitten 1869; as Hymenoloma nordenskjoeldii,
Dusén 1905b; Cardot 1908; Cardot & Brotherus 1923; as Dicranoweisia nordenskjoeldii,
Reimers 1936; as Dicranoweisia antarctica fo. breviseta, Roivainen 1937). Dist. XI. Aisén;
XII. Antártica Chilena, Magallanes, Tierra del Fuego, Última Esperanza.
Hymenoloma brevisetum (Cardot) Ochyra = Hymenoloma antarcticum fide Ochyra et al. 2008.
Hymenoloma crispulum (Hedw.) Ochyra
(as Dicranoweisia crispula, Buck 2002; Matteri & Schiavone 2002; Ochyra 1998; as
Dicranoweisia austrocrispula, Hässel et al. 1984; Matteri & Schiavone 2002; Cardot 1908;
Cardot & Brotherus 1923; Donat 1936; Roivainen 1937; Bartram 1952; Seki 1974; Matteri
1985b; Deguchi 1991a; as Blindia sulphurea, Dusén 1903b; as Dicranoweisia perpulvinata,
Dusén 1903b, 1905b). Dist. X. Llanquihue; XI. Aisén, Coihaique; XII. Antártica Chilena,
Hymenoloma jugelliferum (Dusén) Ochyra
(as Dicranoweisia jugellifera, Matteri & Schiavone 2002; Dusén 1905b). Dist. XII.
Magallanes.
Hymenoloma nordenskjoeldii Dusén = Hymenoloma antarcticum fide Ochyra et al. 2008.
Hymenoloma subglobosum Herzog
Hymenoloma turpe (Cardot) Cardot & Broth.
Donat 1936 (as Dicranoweisia turpis, Matteri & Schiavone 2002; as Verrucidens turpis, Hässel
et al. 1984; Matteri 1985b). Dist. XI. Aisén; XII. Magallanes, Tierra del Fuego, Última
Esperanza.
Hymenostomum kunzeanum (Müll. Hal.) Broth. ≡ Weissia kunzeana.
Hymenostylium longo-pulvinatum Dusén = Hymenostylium recurvirostrum fide Zander 1977.
Hymenostylium recurvirostrum (Hedw.) Dixon
Matteri & Schiavone 2002 (as Hymenostylium longopulvinatus, Dusén 1906a; Cardot 1908;
Cardot & Brotherus 1923; as Anoectangium patagonicum, Cardot & Brotherus 1923). Dist. XI.
Aisén; XII. Magallanes, Tierra del Fuego, Última Esperanza.
Hyophila macrocarpa (Schimp.) Kindb. ≡ Costesia macrocarpa.
Hypnodendron krausei (Müll. Hal.) A. Jaeger = Hypnodendron microstictum fide Reimers 1926.
Hypnodendron microstictum Mitt. ex A. Jaeger & Sauerb.
Tropicos 2009; Larraín 2007b; Larraín 2005; Ireland et al. 2007; Villagran et al. 2003;
Schaumann 2005; He 1998; Müller & Pereira 2006; Mitten 1869; Herzog 1922; Brotherus
1924b; Seki 1974; Robinson 1975; Deguchi 1991a (as Hypnodendron krausei, Dusén 1903b;
Cardot 1908; Reimers 1926; Herzog & Hosseus 1938; Herzog & Schwabe 1939; Herzog 1954,
1960; Bartram 1957). Dist. RM. Cordillera, Santiago; VI. Colchagua; VII. Cauquenes, Talca;
Chiloé, Llanquihue, Osorno, Palena; XI. Aisén; Juan Fernández Is.
Hypnodendron naumanni (Müll. Hal.) Paris = Porothamnium arbusculans fide Reimers 1926.
Hypnum acanthophyllum Mont. ≡ Eurhynchiella acanthophylla.
Hypnum aduncum Hedw. ≡ Drepanocladus aduncus.
Hypnum austrouncinatum Müll. Hal. = Sanionia uncinata fide Wijk et al. 1964.
Hypnum berteroanum Mont. ≡ Catagoniopsis berteroana.
Hypnum campoanum (Thér.) Broth.
NY 2009; Herzog & Hosseus 1938; Herzog 1954, 1960 (as Stereodon campoanum, Thériot
1918). Dist. IX. Cautín, Malleco; X. Osorno, Palena.
Hypnum complanum Mitt. ≡ Rhynchostegium complanum.
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Mosses of Chile 55
Hypnum chrysogaster Müll. Hal.

Larraín 2007b; Larraín 2005; Villagran et al. 2003; Schaumann 2005; Müller & Pereira 2006;
Matteri & Schiavone 2002; Seki 1974; Deguchi 1991a (as Hypnum lechleri, Cardot 1908;
Brotherus 1924b; Roivainen 1934; Herzog & Hosseus 1938; Herzog & Schwabe 1939; Herzog
1954, 1960; Bartram 1957; Robinson 1975; as Stereodon lechleri, Espinosa 1941; Mitten 1869;
Dusén 1903b). Dist. VII. Cauquenes; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno,
Palena; XI. Aisén; Juan Fernández Is.
Hypnum conostomum (Mont.) Mitt., hom. illeg. ≡ Eriodon conostomus.
Hypnum cupressiforme Hedw. var. cupressiforme
Hässel et al. 1984; Ireland et al. 2007; He 1998; Cardot & Brotherus 1923; Reimers 1926;
Herzog & Hosseus 1938; Bartram 1952; Seki 1974; Deguchi 1991a. Dist. IV. Choapa; VIII.
Arauco, Biobío, Concepción, Ñuble; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Osorno;
XI. Aisén; XII. Magallanes, Última Esperanza.
Hypnum cupressiforme var. filiforme Brid.
Ireland et al. 2007; He 1998; Deguchi 1991a. Dist. VIII. Arauco, Biobío, Ñuble; IX. Cautín,
Malleco; XIV. Valdivia/Ranco; X. Llanquihue, Osorno.
Hypnum cupressiforme var. julaceum Brid.
Deguchi 1991a. Dist. XI. Aisén.
Hypnum cupressiforme var. lacunosum Brid.
Matteri & Schiavone 2002; Deguchi 1991a. Dist. XI. Aisén. XII. Última Esperanza.
Hypnum cupressiforme var. mossmanianum (Müll. Hal.) Ando
Díaz et al. 2008; Goffinet et al. 2006; Larraín 2007b; Matcham et al. 2007; Buck 2002; Larraín
2007a; Larraín 2005; Matteri & Schiavone 2002; Deguchi & Oginuma 1989; Deguchi 1991a.
Dist. IX. Malleco; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue; XI. Aisén, Coihaique; XII.
Antártica Chilena, Magallanes.
Hypnum fluitans Hedw. ≡ Warnstorfia fluitans.
Hypnum fluitans var. laculosum (Müll. Hal.) Paris = Warnstorfia fontinaliopsis fide Ochyra & Matteri 2001.
Hypnum fuegianum (Mitt.) Müll. Hal. = Warnstorfia fluitans fide Ochyra & Matteri 2001.
Hypnum lechleri Müll. Hal. = Hypnum chrysogaster fide Ando 1972.
Hypnum longifolium Wilson ex Mitt. ≡ Drepanocladus longifolius.
Hypnum masafuerae E.B. Bartram ≡ Sematophyllum masafuerae.
Hypnum nitidum (Hook.f. & Wilson) Müll. Hal. ≡ Catagonium nitidum.
Hypnum pallens Schimp. ex Cardot, hom. illeg. = Hypnum skottsbergii fide Ando 1972.
Hypnum paradoxum Hook.f. & Wilson ≡ Brachytheciastrum paradoxum.
Hypnum perplicatum Dusén = Drepanocladus perplicatus fide Hedenäs 2003.
Hypnum revolutum (Mitt.) Lindb. var. revolutum fo. revolutum
Matteri 1985b; Hässel et al. 1984. Dist. XII. Magallanes, Última Esperanza.
Hypnum skottsbergii Ando
Goffinet et al. 2006; Larraín 2007b; Hässel et al. 1984; Buck 2002; Larraín 2007a; Larraín
2005; Matteri & Schiavone 2002; Ando 1972, 1982; Seki 1974; Ando & Matteri 1982; Matteri
1985b; Deguchi 1991a (as Hypnum pallens, Herzog & Hosseus 1938; Herzog & Schwabe
1939; Thériot 1934b; as Stereodon pallens, Cardot 1908; Herzog 1922, 1960). Dist. IX. Cautín,
Malleco; XIV. Valdivia/Ranco; X. Chiloé, Osorno; XI. Aisén, Coihaique; XII. Antártica
Hypnum spegazzinii Müll. Hal.
(as Brachythecium spegazzinii, Kindberg 1888). Dist. XII. Magallanes.
Hypnum subpilosum Hook.f. & Wilson ≡ Brachythecium subpilosum.
Hypnum symmetricum (Renauld & Cardot) Renauld & Cardot ≡ Sanionia symmetrica.
Hypnum tenuifolium Hedw. ≡ Rhynchostegium tenuifolium.
Hypnum tenuirostre Hook.f. = Sematophyllum uncinatum fide Stone & Scott 1973.
Hypnum toxarion Schwägr. ≡ Rigodium toxarion.
Hypnum uncinatum Hedw. ≡ Sanionia uncinata.
Hypnum uncinatum Hedw. subsp. symmetricum Renauld & Cardot ≡ Sanionia symmetrica.
Hypopterygium arbuscula Brid. ≡ Dendrohypopterygium arbuscula.
Hypopterygium concinnum (Hook.f.) Brid. ≡ Lopidium concinnum.

56 Müller
Hypopterygium didictyon Müll. Hal.

Goffinet et al. 2006; Larraín 2007b; Hässel et al. 1984; Buck 2002; Larraín 2005; Ireland et al.
2007; Villagran et al. 2003; Schaumann 2005; He 1998; Matteri & Schiavone 2002; Kruijer
2002; Mitten 1869; Dusén 1903b; Cardot 1908; Herzog 1922, 1954, 1960; Cardot & Brotherus
1923; Thériot 1924, 1934b; Herzog & Hosseus 1938; Herzog & Schwabe 1939; Bartram 1952;
Matteri 1973b, 1985b; Deguchi 1991b. Dist. V. Valparaíso; RM. Santiago; VIII. Biobío,
Concepción; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno,
Palena; XI. Aisén; XII. Antártica Chilena, Magallanes, Tierra del Fuego, Última Esperanza.
Hypopterygium flexisetum Hampe = Lopidium concinum fide Matteri 1973b.
Hypopterygium plumarium Mitt. = Lopidium concinum fide Matteri 1973b.
Hypopterygium tamarisci (Sw.) Brid. ex Müll. Hal.
Kruijer 2002. Dist. X. Chiloé.
Hypopterygium thouini (Schwägr.) Mont. = Dendrohypopterygium arbuscula fide Robinson 1975.
Hypopterygium wolffhugelii Herzog = Dendrohypopterygium arbuscula fide Robinson 1975.
Indusiella bryanii (R.S. Williams) S.P. Churchill ≡ Coscinodontella bryanii.
Isopterygiopsis pulchella (Hedw.) Z. Iwats.
Ochyra et al. 2008; Ireland et al. 2007; Matteri & Schiavone 2002 (as Isopterygium fuegianum,
Hässel et al. 1984; Cardot 1908; Matteri 1985b; as Isopterygium leptoplumosum, Hässel et al.
1984; Cardot 1908; Matteri 1985b). Dist. VIII. Biobío; XI. Aisén, Coihaique; XII. Magallanes,
Isopterygium albescens (Hook.) A. Jaeger
Isopterygium fernandezianum Broth. = Isopterygium tenerum fide Robinson 1975.
Isopterygium fernandezianum var. longisetum Broth. = Isopterygium tenerum fide Ireland 1991.
Isopterygium fuegianum Besch. = Isopterygiopsis pulchella fide Zanten 1971.
Isopterygium leptoplumosum (Dusén) Cardot = Isopterygiopsis pulchella fide Matteri & Schiavone 2002.
Isopterygium leucophyllum (Hampe) Mitt. = Isopterygium tenerum fide Ireland 1991.
Isopterygium pulchellum (Hedw.) A. Jaeger ≡ Isopterygiopsis pulchella.
Isopterygium tenerum (Sw.) Mitt.
Matteri & Schiavone 2002; Ireland 1992; Robinson 1975 (as Isopterygium fernandezianum,
Cardot & Brotherus 1923; Brotherus 1924b; Bartram 1957; as Isopterygium fernandezianum
var. longisetum, Barrera 2006; Espinosa 1941; Brotherus 1924b; Bartram 1957; as
Isopterygium leucophyllum, Mitten 1869). Dist. XII. Tierra del Fuego; Juan Fernández Is.
Isothecium serpens Dusén, nom. nud. = Fallaciella gracilis fide Matteri 1985b.
Jaffueliobryum williamsii (Deguchi) Delgad.
Juratzkaea krausei Lorentz = Juratzkaea seminervis subsp. seminervis fide Buck 1977.
Juratzkaea seminervis (Kunze ex Schwägr.) Lorentz subsp. seminervis
Tropicos 2009; Espinosa 1941; He 1998; Müller & Pereira 2006; Brotherus 1924a; Thériot
1928; Herzog & Hosseus 1938; Bartram 1943; Herzog 1954; Buck 1977 (as Juratzkaea
krausei, Thériot 1935; as Juratzkaea seminervis var. arenicola, Thériot 1926; as Stereophyllum
seminerve, Espinosa 1941; Mitten 1869; Thériot 1917, 1918; Herzog 1922; Bartram 1952).
Dist. II. Antofagasta; IV. Choapa, Elqui, Limarí; V. Aconcagua, Petorca, Quillota, San
Antonio, Valparaíso; RM. Chacabuco, Cordillera, Maipo, Melipilla, Santiago, Talagante; VI.
Cachapoal, Cardenal Caro, Colchagua; VII. Cauquenes, Curicó, Linares, Talca; VIII. Arauco,
Biobío, Concepción, Ñuble; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Osorno.
Juratzkaea seminervis var. arenicola Thér. = Juratzkea seminervis subsp. seminervis fide Buck 1977.
Kiaeria pumila (Mitt.) Ochyra
Buck 2002; Matteri & Schiavone 2002 (as Blindia churuccana, Cardot 1900, 1901; Dusén
1905b; as Blindia humilis, Dusén 1903b; Blindia savatieriana, Dusén 1903b; Cardot 1908; as
Dicranoweisia humilis, Dusén 1905b; as Dicranum pumilum, Mitten 1869; Dusén 1905b;
Cardot 1908; Cardot & Brotherus 1923; as Dicranum ramulosum, Mitten 1869; Cardot 1908;
as Holodontium pumilum, Hässel et al. 1984; Roivainen 1937; Herzog 1954; Seki 1974;
ISSN 0945-3466
Mosses of Chile 57
Matteri 1985b; Holodontium ramulosum, Brotherus 1924). Dist. XI. Aisén; XII. Antártica
Kindbergia praelonga (Hedw.) Ochyra
Tropicos 2009; Müller 2009; Larraín 2007b; Larraín 2007a; He 1998; Matteri & Schiavone
2002 (as Stokesiella praelonga, Robinson 1967). Dist. VII. Talca; VIII. Biobío, Concepción;
XIV. Valdivia/Ranco; X. Chiloé, Osorno; XII. Magallanes.
Lamprophyllum splendidissimum (Mont.) Mitt. ≡ Schimperobryum splendidissimum.
Lembophyllum auriculatum (Mont.) Paris ≡ Acrocladium auriculatum.
Lembophyllum orbiculatum (Thér.) Tangney
Tropicos 2009; Schaumann 2005; Müller & Pereira 2006 (as Camptochaete orbiculata, He
1998; Robinson 1970; Deguchi 1991b; as Camptochaete arbusculans, Roivainen 1934;
Bartram 1946; as Weymouthia orbiculata, Thériot 1934b; Herzog & Hosseus 1938; Herzog
1960). Dist. VII. Cauquenes, Talca; VIII. Arauco, Biobío, Concepción, Ñuble; IX. Cautín,
Malleco; XIV. Valdivia/Ranco; X. Llanquihue; XII. Tierra del Fuego.
Lepidopilum splendidissimum Mont. ≡ Schimperobryum splendidissimum.
Leptobryum pottiaceum Dusén = Pohlia wilsonii fide Arts 2001b.
Leptobryum pyriforme (Hedw.) Wilson
Müller 2009; Hässel et al. 1984; Ireland et al. 2007; Ireland & Bellolio 2002; He 1998; Matteri
& Schiavone 2002; Dusén 1903b; Cardot 1908; Cardot & Brotherus 1923; Matteri 1985b. Dist.
IV. Limarí; V. Quillota; RM. Santiago; VII. Curicó, Talca; VIII. Ñuble; XI. Aisén, Coihaique;
XII. Magallanes, Última Esperanza; Easter I.
Leptobryum pyriforme var. antarcticum Besch., nom. nud., Cardot 1908.
Leptobryum pyriforme var. fuegianum Paris, nom. nud., Cardot 1908.
Leptobryum stellatum (Herzog) Broth. = Pohlia wilsonii fide Arts 2001b.
Leptobryum wilsonii (Mitt.) Broth. ≡ Pohlia wilsonii.
Leptodictyum riparium (Hedw.) Warnst.
Ireland et al. 2007 (as Leptodictyum riparium var. jaffueli, He 1998, Thériot 1935; as
Amblystegium jaffueli, Thériot 1924; as Leptodictyum riparium var. looseri, He 1998; Thériot
1934b). Dist. V. San Antonio; RM. Cordillera, Santiago, Talagante; VIII. Biobío, Ñuble; XIV.
Valdivia/Ranco; X. Osorno.
Leptodictyum riparium (Hedw.) Warnst. var. jaffueli Thér. = Leptodictyum riparium fide Ireland et al. 2007.
Leptodictyum riparium (Hedw.) Warnst. var. looseri Thér. = Leptodictyum riparium fide Ireland et al. 2007.
Leptodon smithii (Hedw.) F. Weber & D. Mohr var. smithii
Tropicos 2009; He 1998; Müller & Pereira 2006; Mitten 1869; Dusén 1903b; Cardot 1908;
Cardot & Brotherus 1923; Thériot 1924; Brotherus 1924b; Roivainen 1934; Herzog & Hosseus
1938; Herzog 1954, 1960; Bartram 1957; Robinson 1975; Deguchi 1991b. Dist. V. Valparaíso;
RM. Santiago; VII. Cauquenes, Linares, Talca; VIII. Arauco, Biobío, Concepción, Ñuble; IX.
Cautín; X. Llanquihue; XI. General Carrera; Juan Fernández Is.
Leptodon smithii var. longirepens Cardot & Thér.
Thériot 1924. Dist. Juan Fernández Is.
Leptodontium bullocki Thér. = Leptostomum menziesii fide Hyvönen 1987.
Leptodontium capituligerum Müll. Hal.
Seki 1974. Dist. XI. Aisén.
Leptodontium fernandezianum Broth. = Leptodontium longicaule var. microruncinatum fide Zander 1972.
Leptodontium longicaule Mitt. var. microruncinatum (Dusén) R.H. Zander
Larraín 2007b; Hässel et al. 1984; Villagran et al. 2003; He 1998; Matteri & Schiavone 2002;
Matteri 1985b; Deguchi 1991b (as Leptodontium fernandezianum, Barrera 2006; Espinosa
1941; Brotherus 1924b; Bartram 1957; as Leptodontium microruncinatum, Dusén 1906a;
Cardot 1908; Thériot 1925; Herzog & Hosseus 1938; Herzog 1954; Robinson 1975). Dist. IV.
Limarí; V. Valparaíso; IX. Cautín; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue; XI. Aisén;
XII. Magallanes, Última Esperanza; Juan Fernández Is.
Leptodontium microruncinatum Dusén ≡ Leptodontium longicaule var. microruncinatum.
Leptodontium proliferum Herzog

58 Müller
Ireland et al. 2007 (as Tortula umbrosa, Dusén 1906b; Deguchi 1991b). Dist. VIII. Arauco;
XIV. Valdivia/Ranco.
Leptodontium pungens (Mitt.) Kindb.
Robinson 1975 (as Leptodontium skottsbergii, Bartram 1957). Dist. Juan Fernández Is.
Leptodontium skottsbergii E.B. Bartram = Leptodontium pungens fide Zander 1972.
Leptodontium splachnoides (Hook.f. & Arn.) Thér. ≡ Leptostomum splachnoideum.
Leptopterigynandrum austro-alpinum Müll. Hal.
Cano 2003; He 2005. Dist. XV. Parinacota; II. El Loa.
Leptostomum menziesii R. Br.
Tropicos 2009; Goffinet et al. 2006; Hässel et al. 1984; Matcham et al. 2007; Buck 2002;
Larraín 2005; Espinosa 1941; He 1998; Müller & Pereira 2006; Mitten 1869; Cardot 1900,
1901, 1908; Dusén 1903b; Herzog 1922, 1954; Cardot & Brotherus 1923; Brotherus 1924b;
Roivainen 1934; Herzog & Hosseus 1938; Bartram 1952, 1957; Robinson 1975; Matteri
1985b; Deguchi 1991b (as Leptodontium bullocki, Thériot 1929a; as Leptostomum bullocki,
Andrews 1952). Dist. VII. Cauquenes; VIII. Arauco, Biobío, Ñuble; IX. Cautín, Malleco; XIV.
Valdivia/Ranco; X. Llanquihue, Osorno; XII. Antártica Chilena, Magallanes, Tierra del Fuego,
Leptostomum splachnoideum Hook. & Arn.
Müller 2009; Ireland et al. 2007; Villagran et al. 2003; He 1998; Mitten 1869; Thériot 1935;
Herzog & Hosseus 1938; Herzog 1954; Seki 1974; Deguchi 1991b (as Leptodontium
splachnoides, Thériot 1929a). Dist. VII. Cauquenes; VIII. Arauco, Concepción, Ñuble; IX.
Cautín; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno; XI. Aisén.
Leptotheca gaudichaudii Schwägr.
Tropicos 2009; Goffinet et al. 2006; Matteri & Piñero 1987; Hässel et al. 1984; Buck 2002;
Larraín 2005; Schaumann 2005; He 1998; Matteri & Hyvönen 1991; Matteri & Schiavone
2002; Dusén 1903b; Cardot 1908; Cardot & Brotherus 1923; Herzog & Schwabe 1939;
Bartram 1952; Seki 1974; Robinson 1975; Matteri 1985b; Deguchi 1991a (as Leptotheca
spegazzinii, Dusén 1903b). Dist. VIII. Arauco, Biobío, Concepción; IX. Cautín, Malleco; XIV.
Leptotheca spegazzinii Müll. Hal. = Leptotheca gaudichaudii fide Cardot 1908.
Leptotrichum kunzeanum Müll.Hal. ≡ Leptotrichella kunzeana.
Leptotrichum praealtum Mitt. ≡ Platyneuron praealtum.
Lepyrodon glaucus (Müll. Hal.) Kindb. = Lepyrodon hexastichus fide Allen 1999.
Lepyrodon gunckeli Thér. = Lepyrodon tomentosus fide Allen 1999.
Lepyrodon hexastichus (Mont.) Wijk & Margad.
Larraín 2005; He 1998; Müller & Pereira 2006; Allen 1999; Seki 1974; Robinson 1975;
Deguchi 1991b (as Lepyrodon implexus, Espinosa 1941; Dusén 1903b; Cardot 1908; Herzog
1922, 1954, 1960; Thériot 1924, 1927a; Brotherus 1924b; Reimers 1926; Herzog & Hosseus
1938; Herzog & Schwabe 1939; Bartram 1952, 1957). Dist. VII. Cauquenes, Curicó, Linares,
Talca; VIII. Arauco, Biobío, Concepción, Ñuble; IX. Cautín, Malleco; XIV. Valdivia/Ranco;
X. Chiloé, Llanquihue, Osorno; XI. Aisén, General Carrera; XII. Magallanes; Juan Fernández
Is.
Lepyrodon implexus (Kunze) Kindb. = Lepyrodon hexastichus fide Allen 1999.
Lepyrodon implexus (Kunze) Paris = Lepyrodon hexastichus fide Wijk & Margadant 1961.
Lepyrodon implexus var. reptans Reim.
Reimers 1926. Dist. XIV. Valdivia/Ranco. Note: This is a taxon with doubtful generic
placement (Allen 1999).
Lepyrodon lagurus (Hook.) Mitt.
Goffinet et al. 2006; Hässel et al. 1984; Matcham et al. 2007; Buck 2002; Larraín 2005;
Schaumann 2005; He 1998; Müller & Pereira 2006; Allen 1999; Matteri & Schiavone 2002;
Mitten 1869; Cardot 1900, 1901, 1905, 1908; Dusén 1903b; Herzog 1922; Cardot & Brotherus
1923; Thériot 1934b; Roivainen 1934; Herzog & Schwabe 1939; Bartram 1952; Seki 1974;
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Mosses of Chile 59
Matteri 1985b; Deguchi 1991 (as Lepyrodon lagurus var. laxior, Cardot & Brotherus 1923).
Dist. VII. Cauquenes, Talca; VIII. Arauco, Biobío, Ñuble; IX. Cautín, Malleco; XIV.
Valdivia/Ranco; X. Llanquihue, Osorno; XI. Aisén, Coihaique, General Carrera; XII. Antártica
Lepyrodon lagurus var. laxior Cardot & Broth. = Lepyrodon lagurus fide Allen 1999.
Lepyrodon parvulus Mitt.
Tropicos 2009; Espinosa 1941; He 1998; Müller & Pereira 2006; Allen 1999; Mitten 1869;
Brotherus 1924b; Herzog & Schwabe 1939; Herzog 1954; Bartram 1957; Robinson 1975;
Deguchi 1991b. Dist. IV. Choapa; V. Petorca; VII. Cauquenes; VIII. Arauco, Biobío,
Concepción, Ñuble; IX. Cautín; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno; Juan
Fernández Is.
Lepyrodon patagonicus (Cardot & Broth.) B.H. Allen
Tropicos 2009; Larraín 2007b; Müller & Pereira 2006; Allen 1999; Matteri & Schiavone 2002
(as Lepyrodon tomentosus var. patagonicus, Cardot & Brotherus 1923; Seki 1974; as
Lepyrodon tomentosus var. minor, Cardot & Brotherus 1923). Dist. V. Valparaíso; VII.
Cauquenes, Talca; VIII. Arauco, Biobío, Concepción; IX. Cautín, Malleco; XIV.
Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno; XI. Aisén, Capitán Prat, General Carrera; XII.
Lepyrodon tomentosus (Hook.) Mitt.
Müller 2009; Hässel et al. 1984; He 1998; Matteri & Schiavone 2002; Mitten 1869; Brotherus
1924b; Thériot 1935; Roivainen 1934; Herzog & Hosseus 1938; Bartram 1957; Herzog 1960;
Robinson 1975; Matteri 1985b; Deguchi 1991b (as Lepyrodon gunckelii, Thériot 1934b). Dist.
VII. Talca; VIII. Arauco, Ñuble; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Chiloé,
Llanquihue, Osorno, Palena; XI. Aisén, Coihaique, General Carrera; XII. Antártica Chilena,
Lepyrodon tomentosus var. minor Cardot & Broth. = Lepyrodon patagonicus fide Allen 1999.
Lepyrodon tomentosus var. patagonicus Cardot & Broth. ≡ Lepyrodon patagonicus.
Lescuraea chilensis Herzog = Cratoneuropsis relaxa subsp. minor fide Ochyra et al. 2008.
Leskea fuegiana Besch. = Cratoneuropsis relaxa subsp. minor fide Ochyra et al. 2008.
Leucoloma billardierii (Schwägr.) Broth. ≡ Dicranoloma billardierei.
Leucoloma capillare Dusén ≡ Dicranoloma capillare.
Leucoloma capillifolium Broth. = Dicranoloma menziesii fide Robinson 1975.
Leucoloma dusenii Broth. ≡ Dicranoloma dusenii.
Leucoloma fuegianum Dusén = Dicranoloma hariotii fide Wijk et al. 1962.
Leucoloma fuegianum var. laxum Dusén ≡ Dicranoloma hariotii var. laxum fide Cardot & Brotherus 1923.
Leucoloma grandialare Dusén = Dicranoloma robustum fide Cardot 1908.
Leucoloma hariotii (Müll. Hal.) Broth. ≡ Dicranoloma hariotii.
Leucoloma imponens (Mont.) Dusén ≡ Dicranoloma imponens.
Leucoloma muelleri Dusén ≡ Dicranoloma muelleri.
Leucoloma muelleri var. strictifolium Dusén ≡ Dicranoloma robustum var. strictifolium fide Wijk & Margadant 1961.
Leucoloma nigricaule (Ångström) Broth. = Dicranoloma chilense fide Ochyra & Matteri 1996.
Leucoloma nigricaule var. gracile Dusén = Dicranoloma capillare fide Cardot 1908.
Leucoloma nigricaule var. flexuosulum Dusén ≡ Dicranoloma nigricaule var. flexuosulum fide Bartram in Roivainen
1937.
Leucoloma perremotifolium Dusén ≡ Dicranoloma perremotifolium.
Leucoloma perremotifolium var. fragile Dusén ≡ Dicranoloma perremotifolium var. fragile.
Leucoloma peruncinatum Dusén = Dicranoloma chilense fide Ochyra & Matteri 1996.
Leucoloma robustum (Hook.f. & Wilson) Broth. ≡ Dicranoloma robustum.
Leucoloma robustum var. flexuosum Dusén = Dicranoloma robustum fide Matteri & Schiavone 2002.
Leucoloma robustum var. lagunicola Dusén ≡ Dicranum robustum var. lagunicola.
Leucoloma setosum (Hook.f. & Wilson) Broth. ≡ Dicranoloma robustum var. setosum.
Leucomium crinitifolium (Müll. Hal.) Mitt.
Mitten 1869; Allen 1987b. Dist. Chile, without exact locality (leg. Poeppig).
Limprichtia cossonii (Schimp.) L.E. Anderson, H.A. Crum & W.R. Buck ≡ Scorpidium cossonii.
Limprichtia revolvens (Sw.) Loeske ≡ Scorpidium revolvens.
Lopidium concinnum (Hook.) Wilson

60 Müller
Tropicos 2009; Larraín 2007b; Larraín 2005; Espinosa 1941; Villagran et al. 2003; He 1998;
Kruijer 2002; Brotherus 1924b; Bartram 1957; Matteri 1973b; Robinson 1975; Deguchi 1991b
(as Hypopterygium concinnum, Cardot 1908; as Hypopterygium flexisetum, Dusén 1903b;
Cardot 1908; as Lopidium plumarium, Mitten 1869; Herzog 1922, 1954, 1960; Cardot &
Brotherus 1923; Reimers 1926; Herzog & Hosseus 1938; Herzog & Schwabe 1939; as
Hypopterygium plumarium, Bartram 1952). Dist. V. Valparaíso; VI. Colchagua; VIII. Arauco;
IX. Cautín; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno, Palena; XI. Aisén; Juan
Fernández Is.
Lopidium plumarium (Mitt.) Hampe = Lopidium concinum fide Matteri 1973b.
Macrocoma brasiliensis (Mitt.) Vitt
He 1998. Dist. IX. Cautín.
Macrocoma filiformis (Hook.f. & Wilson) Grout = Macrocoma orthotrichoides fide Vitt 1980.
Macrocoma gracillima (Besch.) Vitt ≡ Matteria gracillima.
Macrocoma hymenostomum (Mont.) Grout = Macrocoma sullivantii fide Grout 1944.
Macrocoma orthotrichoides (Raddi) Wijk & Margad.
Vitt 1980 (as Macrocoma filiformis, Grout 1944; as Macromitrium filiforme, Mitten 1869).
Dist. Chile, without exact locality (leg. Poeppig).
Macrocoma papillosa (Thér.) Vitt ≡ Matteria papillosa.
Macrocoma sullivantii (Müll. Hal.) Grout
He 1998; Müller & Pereira 2006; Robinson 1975 (as Macrocoma hymenostomum, Grout 1946;
as Macromitrium hymenostomum, Mitten 1869; Dusén 1903b; Cardot 1908; Cardot &
Brotherus 1923; Brotherus 1924b; as Macromitrium paraphysatum, Mitten 1869; as
Macromitrium paraphysatum var. chilensis, Thériot 1921; as Macrocoma tenuis subsp.
sullivantii, Tropicos 2009; Vitt 1980; Deguchi 1991b; as Macromitrium hariotii, Hässel et al.
1984; He 1998; Matteri & Schiavone 2002; Dusén 1903b; Cardot 1908; Cardot & Brotherus
1923; Matteri 1985b). Dist. V. Valparaíso; VII. Cauquenes, Talca; VIII. Arauco, Biobío,
Concepción, Ñuble; IX. Cautín; X. Chiloé, Llanquihue, Palena; XII. Antartica Cilena,
Macrocoma tenuis (Hook. & Grev.) Vitt subsp. sullivantii (Müll. Hal.) Vitt ≡ Macrocoma sullivantii.
Macromitrium bifasciculare Müll. Hal. ex Dusén
Matteri & Schiavone 2002; Dusén 1903b; Cardot 1908. Dist. XI. Aisén; XII. Magallanes.
Macromitrium campoanum Thér.
Herzog & Hosseus 1938. Dist. X. Chiloé.
Macromitrium coriaceum E.B. Bartram = Macromitrium tenax fide Seki 1974.
Macromitrium crassiusculum Lorentz
Ireland et al. 2007; Thériot 1925; Herzog & Hosseus 1938; Herzog 1960. Dist. VIII.
Concepción; IX. Cautín; XIV. Valdivia/Ranco; X. Osorno.
Macromitrium fernandezianum Broth.
Barrera 2006; Espinosa 1941; Brotherus 1924b; Bartram 1957; Robinson 1975. Dist. Juan
Fernández Is.
Macromitrium filiforme Hook.f. & Wilson = Macrocoma orthotrichoides fide Vitt 1980.
Macromitrium gracillimum (Besch.) Broth. ≡ Matteria gracillima.
Macromitrium hariotii Besch. ex Müll. Hal. = Macrocoma sullivantii fide Grout 1944.
Macromitrium hymenostomum Mont. = Macrocoma sullivantii fide Grout 1944.
Macromitrium krausei Lorentz
Larraín 2007b; Ireland et al. 2007; He 1998; Dusén 1903b; Cardot 1908; Herzog & Hosseus
1938; Bartram 1952. Dist. VIII. Ñuble; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue; XI.
Aisén.
Macromitrium longirostre (Hook.) Schwägr.
Mitten 1869. Dist. V. Valparaíso; X. Chiloé.
Macromitrium masafuerae Broth.
Brotherus 1924b; Bartram 1957; Robinson 1975. Dist. Juan Fernández Is.
Macromitrium microcarpum Müll. Hal.
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Mosses of Chile 61
Larraín 2005; Ireland et al. 2007; Villagran et al. 2003; He 1998; Mitten 1869; Thériot 1935;
Herzog & Schwabe 1939; Bartram 1952; Herzog 1960. Dist. VII. Maule; VIII. Arauco,
Concepción, Ñuble; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno.
Macromitrium microstomum (Hook. & Grev.) Schwägr.
Robinson 1975 (as Macromitrium saxatile, Espinosa 1941; Mitten 1869; Brotherus 1924b;
Bartram 1957). Dist. Juan Fernández Is.
Macromitrium papillosum Thér. ≡ Matteria papillosa.
Macromitrium paraphysatum Mitt. = Macrocoma sullivantii fide Vitt 1980.
Macromitrium paraphysatum var. chilensis Thér. = Macrocoma sullivantii fide Vitt 1980.
Macromitrium pertriste Müll. Hal.
Thériot 1921; Thériot 1934b. Dist. XIV. Valdivia/Ranco; X. Chiloé.
Macromitrium poeppigii Duby = Zygodon pentastichus fide Robinson 1975.
Macromitrium saddleanum Besch. ex Müll. Hal.
Matteri & Schiavone 2002; Dusén 1903b; Cardot 1908; Herzog & Hosseus 1938. Dist. XIV.
Valdivia/Ranco; XII. Antártica Chilena, Magallanes.
Macromitrium saddleanum var. piliferum Herzog
Macromitrium saxatile Mitt. = Macromitrium microstomum fide Robinson 1975.
Macromitrium spurio-crispulum Dusén, nom. nud., Dusén 1903b.
Macromitrium tenax Müll. Hal.
Tropicos 2009; Hässel et al. 1984; Matteri & Schiavone 2002; Dusén 1903b; Cardot 1908; Seki
1974; Matteri 1985b (as Macromitrium coriaceum, Roivainen 1934; Bartram 1946). Dist. X.
Chiloé; XI. Aisén; XII. Antártica Chilena, Magallanes, Tierra del Fuego, Última Esperanza.
Macromitrium spec.
Ireland & Bellolio 2002; Brotherus 1924. Dist. Easter I.
Mahua enervis W.R. Buck
Buck 1983 (as Fabronia enervis, Herzog 1960). Dist. XIV. Valdivia/Ranco.
Matteria gracillima (Besch.) Goffinet
Larraín 2007a; He 1998; Matteri & Calabrese 1999 (as Macrocoma gracillima, Larraín 2005;
Deguchi 1991b; as Macromitrium gracillimum, Herzog 1954; Seki 1974). Dist. XIV.
Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno; XI. Aisén; XII. Magallanes, Última
Esperanza.
Matteria papillosa (Thér.) Goffinet
(as Macrocoma papillosa, Deguchi 1991b; as Macromitrium papillosum, Herzog & Hosseus
1938; Herzog 1954). Dist. X. Llanquihue; XI. Aisén.
Meesia hymenostoma Cardot & Broth. = Meesia uliginosa fide Matteri & Ochyra 1999.
Meesia patagonica Dusén = Dicranella hookeri fide Matteri & Ochyra 1999.
Meesia uliginosa Hedw.
Buck 2002; Matteri & Schiavone 2002; Matteri & Ochyra 1999. Dist. XII. Antártica Chilena,
Meteoriopsis straminea (Müll. Hal.) Broth. ex Paris
(as Meteorium stramineum, Mitten 1869). Dist. Chile, without exact locality (leg. Poeppig).
Meteorium flexicaule Wilson ≡ Papillaria flexicaulis.
Meteorium genuflexum Müll. Hal. ≡ Ancistrodes genuflexa.
Meteorium molle (Hedw.) Wilson ≡ Weymouthia mollis.
Meteorium stramineum (Müll. Hal.) Mitt. ≡ Meteoriopsis straminea.
Metzleria patagonica (Herzog & Thér.) J.-P. Frahm ≡ Atractylocarpus patagonicus.
Microcampylopus leucogaster (Müll. Hal.) B.H. Allen
Tropicos 2009. Dist. VIII. Arauco (Ireland & Bellolio 32952, MO).
Microdus euchlorus (Mont.) Paris ≡ Campylopodium euchlorum.
Microdus kunzeanus (Müll. Hal.) Broth. ≡ Leptotrichella kunzeana.
Mielichhoferia argentifolia Mitt.
He 1998. Dist. IV. Choapa; V. Valparaíso; RM. Talagante.
Mielichhoferia brevicaulis Hornsch. ≡ Schizymenium brevicaule.

62 Müller
Mielichhoferia campylocarpa (Hook.f. & Arnott) Mitt. ≡ Schizymenium campylocarpum.

Mielichhoferia chilensis Mont. ≡ Pohlia chilensis.
Mielichhoferia demissa Müll. Hal.
Muñoz & Pisano 1947; Villagran et al. 2004. Dist. IV. Limarí.
Mielichhoferia fulvonitens Cardot & Broth. ≡ Schizymenium fulvonitens.
Mielichhoferia longiseta Müll. Hal.
Brotherus 1924b; Robinson 1975. Dist. Juan Fernández Is.
Mielichhoferia multiflora E.B. Bartram ≡ Schizymenium multiflorum.
Mielichhoferia pleurogyna Mont. ≡ Schizymenium pleurogynum.
Mielichhoferia skottsbergii Cardot & Broth. ≡ Schizymenium skottsbergii.
Mielichhoferia spegazzinii Müll. Hal. = Schizymenium campylocarpum fide Matteri 1998.
Mittenothamnium deguchii N. Nishim. & Kanda
Barrera 2006; Nishimura & Kanda 1990; Deguchi 1991a. Dist. XI. Aisén.
Mitthyridium fasciculatum (Hook. & Grev.) H. Rob.
(as Syrrhopodon fasciculatus, Mitten 1869). Dist. Chile, without exact locality (leg. Lobb).
Mniadelphus flaccidus (Hook.f & Wilson) Hampe ≡ Distichophyllum flaccidum.
Mniadelphus krausei Lorentz ≡ Distichophyllum krausei.
Mniobryum albicans (Wahlenb.) Limpr. = Pohlia wahlenbergii fide Shaw 1982.
Mniobryum alticaule (Müll. Hal.) Broth. = Pohlia wahlenbergii fide Warburg 1962.
Mniobryum alticaule var. crassinerve Broth. = Pohlia wahlenbergii fide Müller 2009.
Mniobryum amplirete (Müll. Hal.) Broth. = Pohlia wahlenbergii fide Warburg 1962.
Mniobryum aspillagae Thér. = Pohlia wilsonii fide Müller 2009.
Mniobryum philonoteum (Müll. Hal.) Broth. = Pohlia wahlenbergii fide Matteri 1998.
Mnium leptolimbatum Müll. Hal. = Plagiomnium ellipticum fide T. Koponen 1974.
Mnium rostratum Schrad. ≡ Plagiomnium rostratum.
Mnium subpunctatum Cardot & Broth. = Cinclidium stygium fide T. Koponen 1969.
Molendoa fuegiana E.B. Bartram
Matteri & Schiavone 2002; Bartram 1946. Dist. XII. Tierra del Fuego.
Monocranum angustirete Müll. Hal. ex Broth. = Eucamptodon perichaetialis fide Ochyra 1994.
Monocranum dicranoides Müll. Hal. ex Kindb. = Eucamptodon perichaetialis fide Ochyra 1994.
Muelleriella crassifolia (Hook.f. & Wilson) Dusén
Goffinet et al. 2006; Hässel et al. 1984; Matcham et al. 2007; Buck 2002; Buck 2005b; Matteri
& Schiavone 2002; Dusén 1905a; Matteri 1985b; Deguchi 1991b (as Muelleriella crassifolia
var. lingulata, Dusén 1905a; as Orthotrichum crassifolium, Dusén 1903b; Cardot 1908; Cardot
& Brotherus 1923; as Orthotrichum ligulatum, Dusén 1903b). Dist. XII. Antártica Chilena,
Muelleriella crassifolia var. lingulata Dusén = Muelleriella crassifolia fide Vitt 1976.
Muscoflorschuetzia pilmaiquen (Crosby) Crosby ≡ Diphyscium pilmaiquen.
Muscoherzogia pauperculata (Thér. & Herzog) Ochyra
(as Valdiviella paupercula, Herzog 1928). Dist. XIV. Valdivia/Ranco; X. Llanquihue.
Myurella julacea (Schwägr.) Schimp.
Hässel et al. 1984; Matteri 1981, 1985b. Dist. XII. Magallanes, Última Esperanza.
Neckera berteroana Schimp. ex Mitt. = Neckera scabridens fide Sastre-de Jesus 1987.
Neckera chilensis Schimp. ex Mont.
Tropicos 2009; Hässel et al. 1984; Pereira et al. 2006; Espinosa 1941; Muñoz & Pisano 1947;
He 1998; Müller & Pereira 2006; Mitten 1869; Dusén 1903b; Cardot 1908; Thériot 1915,
1917; Herzog 1922, 1954; Brotherus 1924a; Reimers 1926; Herzog & Hosseus 1938; Matteri
1985b; Deguchi 1991b (as Neckera jamesoni, Mitten 1869). Dist. IV. Choapa, Limarí; V.
Aconcagua, Petorca, Valparaíso; RM. Cordillera; VII. Cauquenes, Talca; VIII. Arauco, Biobío,
Concepción, Ñuble; IX. Cautín; XIV. Valdivia/Ranco; X. Llanquihue, Osorno; XI. Aisén; XII.
Neckera jamesoni Taylor = Neckera chilensis fide Sastre-de Jesus 1987.
Neckera rotundata Broth.
Neckera scabridens Müll. Hal. var. scabridens
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Tropicos 2009; Matcham et al. 2007; Pereira et al. 2006; Larraín 2005; Villagran et al. 2003;
He 1998; Müller & Pereira 2006; Mitten 1869; Dusén 1903b; Cardot 1908; Herzog 1922,
1954, 1960; Reimers 1926; Bartram 1952; Seki 1974; Deguchi 1991b (as Neckera berteroana,
Mitten 1869; Thériot 1915). Dist. IV. Choapa; V. Petorca, Valparaíso; RM. Santiago; VII.
Neckera scabridens var. porteri Thér.
Thériot 1915, 1917. Dist. V. Valparaíso; XIV. Valdivia/Ranco.
Neobarbula magellanica Dusén = Hennediella densifolia fide Cano 2008.
Neoloma fuegianum Cardot ≡ Andreaea fuegiana.
Neomeesia paludella (Besch.) Deguchi
Matcham et al. 2007; Buck 2002; Matteri & Ochyra 1999; Matteri & Schiavone 2002; Deguchi
1983, 1991b (as Dicranella paludella, Dusén 1905b; Cardot 1908; Cardot & Brotherus 1923;
as Anisothecium paludella, Hässel et al. 1984; Roivainen 1937; Herzog 1954; Matteri 1985b).
Dist. XI. Aisén; XII. Antártica Chilena, Magallanes, Tierra del Fuego, Última Esperanza.
Neosharpiella turgida (Mitt.) H. Rob. & Delgad.
Fife 1980; Larraín et al. 2009. Dist. VII. Talca.
Neuroloma fuegianum Cardot ≡ Andreaea fuegiana.
Niphotrichum canescens (Hedw.) Bednarek-Ochyra & Ochyra
(as Racomitrium canescens, He 1998). Dist. RM. Santiago.
Notoligotrichum angulatum (Cardot & Broth.) G.L. Smith = Notholigotrichum minimum.
Notoligotrichum australe (Hook.f. & Wilson) G. L. Sm.
He 1998. Dist. IX. Malleco.
Notoligotrichum minimum (Cardot) G.L. Sm.
Buck 2005a, 2005b; Matteri & Schiavone 2002; Robinson 1975 (as Polytrichadelphus
minimus, Cardot 1908; as Psilopilum minimum, Hässel et al. 1984; Matteri 1985b; as
Notholigotrichum angulatum, Larraín 2005; Schaumann 2005, Seki 1974; Deguchi 1991b; as
Psilopilum angulatum, Hässel et al. 1984; Cardot & Brotherus 1923; Matteri 1985b). Dist. IX.
Malleco; XIV. Valdivia/Ranco; X. Osorno; XI. Aisén; XII. Antártica Chilena, Magallanes,
Notoligotrichum tapes var. apiculatum (Cardot) Schiavone
Matteri & Schiavone 2002 (as Psilopilum tapes var. apiculatum, Cardot & Brotherus 1923).
Dist. Tierra del Fuego.
Notoligotrichum tapes (Müll. Hal.) G. L. Sm. var. tapes
(as Psilopilum cuspidatum, Dusén 1905a). Dist. XII. Tierra del Fuego.
Notoligotrichum trichodon (Hook.f. & Wilson) G.L. Sm.
Ochyra et al. 2008; Goffinet et al. 2006; Buck 2002; Schiavone & Hyvönen 1993; Matteri &
Schiavone 2002; Deguchi 1991b (as Psilopilum trichodon, Hässel et al. 1984; Matteri 1985b;
Psilopilum antarcticum, Cardot 1908; Brotherus 1924b; as Psilopilum antarcticum var.
densifolium, Barrera 2006; Cardot & Brotherus 1923; as Psilopilum magellanicum, Dusén
1905b). Dist. XI. Aisén; XII. Antártica Chilena, Magallanes, Tierra del Fuego, Última
Esperanza.
Octodiceras dillenii (Bach Pyl.) Brid. = Fissidens semicompletus fide Pursell 1994.
Oedipodium griffithianum (Dicks.) Schwägr.
Matteri 1979. Dist. VIII. Biobío.
Oligotrichum austroaligerum G.L. Sm.
Matteri & Schiavone 2002; Schiavone 1993 (as Oligotrichum magellanicum, Cardot &
Brotherus 1923; Herzog 1954). Dist. XI. Aisén; XII. Magallanes.
Oligotrichum canaliculatum (Hook. & Arn.) Mitt. var. canaliculatum
Tropicos 2009; Larraín 2007b; Pereira et al. 2006; Larraín 2005; Espinosa 1941; Mullen &
Frye 1947; Villagran et al. 2003; He 1998; Müller & Pereira 2006; Mitten 1869; Thériot 1915,
1925, 1935; Herzog 1922, 1954, 1960; Brotherus 1924a, 1924b; Reimers 1926; Herzog &

64 Müller
Hosseus 1938; Herzog & Schwabe 1939; Bartram 1952, 1957; Seki 1974; Robinson 1975;
Deguchi 1991b (as Pogonatum liliputanum, Thériot 1917; as Oligotrichum canaliculatum var.
molinae, Thériot 1924). Dist. V. San Antonio, Valparaíso; RM. Melipilla; VII. Cauquenes,
Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno; XI. Aisén; Juan Fernández Is.
Oligotrichum canaliculatum var. minus Broth.
Mullen & Frye 1947; Herzog & Hosseus 1938; Herzog & Schwabe 1939; Bartram 1957. Dist.
VI. Colchagua; XIV. Valdivia/Ranco; Juan Fernández Is.
Oligotrichum canaliculatum var. molinae (Mont.) Thér. = Oligotrichum canaliculatum fide Wijk et al. 1959.
Oligotrichum dendroides (Brid. ex Hedw.) Kindb. ≡ Dendroligotrichum dendroides.
Oligotrichum laevifolium Thér.
Thériot 1924; Herzog & Hosseus 1938. Dist. XIV. Valdivia/Ranco; X. Chiloé.
Oligotrichum magellanicum Cardot & Broth. = Oligotrichum austroaligerum fide Smith 1969.
Oligotrichum obtusifolium Thér.
Thériot 1934b. Dist. XIV. Valdivia/Ranco.
Oligotrichum rigidum (Lorentz) Broth. ≡ Hebantia rigida.
Oligotrichum subcanaliculatum Dusén = Oligotrichum canaliculatum var. minus fide Mullen & Frye 1947.
Ombronesus stuvensis N.E. Bell, N. Pedersen & A.E. Newton
Bell et al. 2007. Dist. XI. Capitán Prat.
Oncophorus fuegianus Cardot ≡ Symblepharis fuegiana.
Oncophorus luteo-virens E.B. Bartram ≡ Symblepharis luteovirens.
Oreoweisia chilensis (Hampe ex Müll.Hal.) Paris = Oreoweisia erosa fide O’Shea & Matcham in Blockeel et al. 2004.
Oreoweisia erosa (Hampe ex Müll.Hal.) Kindb.
O’Shea & Matcham in Blockeel et al. 2004 (as Oreoweisia cf. chilensis, Buck 2002). Dist. II.
El Loa; XII. Antártica Chilena; „Andes Chilensis“ (type of O. chilensis).
Orthodontium australe Hook.f. & Wilson = Orthodontium lineare fide Matteri 1985b.
Orthodontium australe var. brevisetum Cardot & Broth. = Orthodontium lineare fide Meijer 1951.
Orthodontium gracile (Wilson) Schwägr. ex B.S.G.
Ireland et al. 2007; He 1998; Meijer 1951; Herzog 1954; Bartram 1957; Robinson 1975 (as
Orthodontium tenellum, Mitten 1869; as Stableria tenella, Thériot 1923). Dist. V. Quillota,
Valparaíso; VIII. Biobío, Concepción, Ñuble; Juan Fernández Is.
Orthodontium humile Mont. ≡ Pohlia humilis.
Orthodontium lineare Schwägr.
Hässel et al. 1984; Matcham et al. 2007; Buck 2002; He 1998; Matteri & Schiavone 2002;
Matteri 1985b (as Orthodontium australe, Dusén 1903b; Cardot & Brotherus 1923; Deguchi
1991a; as Apalodium australe, Mitten 1869; as Orthodontium australe var. brevisetum, Cardot
& Brotherus 1923). Dist. X. Llanquihue; XII. Antártica Chilena, Magallanes, Tierra del Fuego,
Última Esperanza.
Orthodontium pellucens (Hook.) B.S.G.
Villagran et al. 2004; Ireland et al. 2007; He 1998; Herzog 1954. Dist. IV. Limarí; VIII.
Biobío, Concepción, Ñuble; X. Osorno.
Orthodontium tenellum Mitt. = Orthodontium gracile fide Meijer 1951.
Orthogrimmia sessitana (De Not.) Ochyra ≡ Grimmia sessitana.
Orthorrhynchium chilensis R.S. Williams = Catagonium nitens subsp. nitens fide Lin 1984.
Orthotrichum aequatoreum Mitt.
Orthotrichum anaglyptodon Cardot & Broth.
Müller 2009; Ireland et al. 2007; He 1998; Lewinsky 1987; Deguchi 1991b. Dist. RM.
Chacabuco, Cordillera, Santiago; VII. Curicó, Talca; VIII. Ñuble; IX. Cautín, Malleco; XI.
Coihaique, General Carrera; XII. Magallanes, Última Esperanza.
Orthotrichum anderssonii Ångström = Orthotrichum elegantulum fide Lewinsky 1987.
Orthotrichum angustissimum Dusén, nom. nud. = Orthotrichum elegantulum fide Lewinsky 1987.
Orthotrichum anomalum Hedw.
Buck 2005b. Dist. XII. Antártica Chilena.
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Mosses of Chile 65
Orthotrichum araucarieti Müll. Hal.

NY 2009; Müller 2009. Dist. VIII. Ñuble; XI. General Carrera.
Orthotrichum argentinicum Herzog = Orthotrichum elegantulum fide Lewinsky 1987.
Orthotrichum aristoblepharum Herzog = Orthotrichum johnstonii fide Lewinsky 1987.
Orthotrichum assimile Müll. Hal.
Müller 2009; Goffinet et al. 2007; Matcham et al. 2007; He 1998; Matteri & Schiavone 2002;
Mitten 1869; Thériot 1921, 1928; Herzog 1954; Lewinsky 1984; Deguchi 1991b (as
Orthotrichum conceptionense, Thériot 1926; as Orthotrichum macloskii, Hässel et al. 1984;
Dusén 1903b; as Orthotrichum paleomitrium, Dusén 1903b). Dist. V. Valparaíso; RM.
Chacabuco, Cordillera; VII. Cauquenes, Curicó, Linares, Talca; VIII. Biobío, Concepción,
Ñuble; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Llanquihue, Osorno, Palena; XI. General
Carrera; XII. Magallanes, Última Esperanza.
Orthotrichum bicolor Thér. = Orthotrichum incanum fide Matteri 2000.
Orthotrichum brevicolle Mitt. = Ulota rufula fide Malta 1927.
Orthotrichum brotheri Dusén ex Lewinsky
NY 2009; Tropicos 2009; Müller 2009; Lewinsky 1987; Deguchi 1991b. Dist. VIII. Biobío,
Ñuble; IX. Cautín; X. Palena; XI. Aisén, General Carrera; XII. Antártica Chilena, Ultima
Esperanza.
Orthotrichum carinatum Mitt. ≡ Ulota carinata.
Orthotrichum chilense Mitt. = Ulota rufula fide Malta 1927.
Orthotrichum compactum Dusén = Orthotrichum inclinatum fide Matteri 2000.
Orthotrichum conceptionense Thér. = Orthotrichum assimile fide Lewinsky 1984.
Orthotrichum crassifolium Hook.f. & Wilson ≡ Muelleriella crassifolia.
Orthotrichum crenatoerosum Müll. Hal. = Orthotrichum incanum fide Matteri 2000.
Orthotrichum cupulatum Hoffm. ex Brid. var. austroamericanum Lewinsky
Müller 2009; Lewinsky 1984. Dist. Patagonia; XI. General Carrera.
Orthotrichum darwinii (Mitt.) Mitt. ≡ Ulota fuegiana var. darwinii.
Orthotrichum densum Lewinsky
Müller 2009; He 1998; Lewinsky 1987. Dist. IX. Cautín; XI. Coihaique; XII. Última
Esperanza.
Orthotrichum diaphanum Schrad. ex Brid. var. podocarpi (Müll. Hal.) Lewinsky
Müller 2002, 2009. Dist. II. Antofagasta; XII. Magallanes.
Orthotrichum elegantulum Schimp. ex Mitt.
Goffinet et al. 2006; Matcham et al. 2007; Buck 2002; He 1998; Matteri & Schiavone 2002;
Mitten 1869; Dusén 1903b; Cardot & Brotherus 1923; Bartram 1952; Seki 1974; Matteri
1985b; Lewinsky 1987; Deguchi 1991b (as Ulota gymnomitria, Dusén 1903b; as Orthotrichum
andersonii, Ångström 1872; as Ulota andersonii, Cardot 1908; as Orthotrichum argentinicum,
Seki 1974; as Orthotrichum angustissimum, Dusén 1903b; as Orthotrichum hymenomitrium,
Dusén 1903b). Dist. V. Valparaíso; VII. Cauquenes, Curicó, Linares, Talca; VIII. Biobío,
Ñuble; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Llanquihue, Osorno; XI. Aisén; XII.
Orthotrichum eremitense Mitt. = Ulota glabella fide Malta 1927.
Orthotrichum freyanum Goffinet, W.R. Buck & M.A. Wall
Goffinet et al. 2007. Dist. VIII. Biobío.
Orthotrichum fuegianum Mitt. ≡ Ulota fuegiana.
Orthotrichum fulvellum (Mitt.) Mitt. = Ulota luteola fide Wijk et al. 1964.
Orthotrichum germanum Mont. ≡ Ulota germana.
Orthotrichum gigantosporum Lewinsky
(as Orthotrichum macrosporum, Lewinsky & Deguchi 1989; Lewinsky 1984; Deguchi 1991b).
Dist. IX. Malleco.
Orthotrichum glabellum Mitt. ≡ Ulota glabella.
Orthotrichum gymnomitria Müll. Hal. = Orthotrichum elegantulum fide Lewinsky 1987.
Orthotrichum hortense Bosw.

66 Müller
Müller 2009; He 1998; Lewinsky 1987; Deguchi 1991b. Dist. VII. Talca; VIII. Biobío; IX.
Cautín, Malleco; XIV. Valdivia/Ranco.
Orthotrichum hymenomitrium Dusén, nom. nud. = Orthotrichum elegantulum fide Lewinsky 1987.
Orthotrichum incanum Müll. Hal.
Müller 2009; He 1998; Matteri & Schiavone 2002; Müller 1885 (as Ulota incana, Dusén
1903b; Cardot 1908; as Orthotrichum crenato-erosum, Lewinsky 1987; as Ulota crenato-
erosa, Dusén 1903b; Cardot 1908; as Orthotrichum bicolor, Tropicos 2009; Pereira et al. 2006;
Müller & Pereira 2006; Thériot 1921, 1928, 1935; Herzog 1954; Lewinsky 1987; as
Orthotrichum ulotoides, Herzog 1922). Dist. V. Valparaiso; VII. Cauquenes, Curico, Linares,
Talca; VIII. Biobío, Concepción, Ñuble; IX. Cautin; XIV. Valdivia/Ranco; X. Llanquihue,
Osorno; XI. Coihaique; XII. Antártica Chilena, Magallanes, Ultima Esperanza.
Orthotrichum inclinatum Müll. Hal.
Matteri & Schiavone 2002; Müller 1885; Matteri 2000 (as Ulota inclinata, Dusén 1903b; as
Orthotrichum compactum, Goffinet et al. 2006; Matcham et al. 2007; Buck 2002; Dusén
1903b; Herzog 1960; Lewinsky 1984; as Orthotrichum perpusillum Dusén 1903b). Dist. X.
Llanquihue; XI. Aisen; XII. Antártica Chilena, Magallanes, Tierra del Fuego, Ultima
Esperanza.
Orthotrichum johnstonii E.B. Bartram
He 1998; Bartram 1943; Lewinsky 1987 (as Orthotrichum aristoblepharum, Herzog 1954).
Dist. III. Atacama; IV. Limarí.
Orthotrichum latimarginatum Lewinsky
Orthotrichum laxifolium Wilson
Orthotrichum leiothecium Müll. Hal.
(as Ulota leiothecia, Dusén 1903b; Cardot 1908). Dist. XII. Magallanes. Note: Lewinsky
(1987) considered this species to be an uncertain taxon because she did’nt see the type.
Orthotrichum ligulatum Dusén, nom. nud. = Muelleriella crassifolia fide Lewinsky 1984.
Orthotrichum lobbianum Mitt. ≡ Ulota lobbiana.
Orthotrichum lorentzii Müll. Hal. = Orthotrichum pycnophyllum fide Lewinsky 1987.
Orthotrichum ludificans Lewinsky
Müller 2009; Lewinsky 1987; Deguchi 1991b; Matteri & Schiavone 2002. Dist. VIII. Biobío;
IX. Cautín, Malleco; XI. Coihaique; XII. Última Esperanza.
Orthotrichum luteolum Hook. f. & Wilson ≡ Ulota luteola.
Orthotrichum macloskii Dusén = Orthotrichum assimile fide Lewinsky 1984.
Orthotrichum macrocalycinum Mitt. ≡ Ulota macrocalycina.
Orthotrichum macrosporum Lewinsky = Orthotrichum gigantosporum fide Lewinsky 1988.
Orthotrichum magellanicum Mont. ≡ Ulota magellanica.
Orthotrichum nigritellum Dusén, nom. nud. = Orthotrichum inclinatum fide Matteri 2000.
Orthotrichum paleomitrium Dusén, nom. nud. = Orthotrichum assimile fide Lewinsky 1984.
Orthotrichum pariatum Mitt.
Tropicos 2009; Müller 2009. Dist. VII. Talca; VIII. Ñuble (Ireland & Bellolio 30669, MO, det.
as O. pariatum ?)
Orthotrichum perexiguum Dusén ex Lewinsky
Tropicos 2009; Lewinsky 1984. Dist. VIII. Biobío; IX. Cautín; XIV. Valdivia/Ranco; XII.
Magallanes.
Orthotrichum perpusillum Dusén, nom. nud. = Orthotrichum inclinatum fide Matteri 2000.
Orthotrichum phyllanthum (Brid.) Steud. ≡ Ulota phyllantha.
Orthotrichum pycnophyllum Schimp.
(as Orthotrichum lorentzii, Herzog & Hosseus 1938) Dist. IX. Cautín. Note: This species was
not reported from Chile by Lewinsky (1987). The record of Herzog & Hosseus (1938) is
therefore in need of verification.
Orthotrichum rufulum Mitt. ≡ Ulota rufula.
Orthotrichum rupestre Schleich. ex Schwägr.
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Mosses of Chile 67
Tropicos 2009; Müller 2009; Goffinet et al. 2006; Hässel et al. 1984; Matcham et al. 2007;
Buck 2002; He 1998; Matteri & Schiavone 2002; Dusén 1903b; Thériot 1921; Lewinsky 1987;
Deguchi 1991b (as Orthotrichum subrupestre, Thériot 1923). Dist. V. Valparaíso; RM.
Chacabuco, Cordillera, Santiago; VI. Colchagua; VII. Curicó, Talca; VIII. Biobío, Ñuble; IX.
Malleco; X. Llanquihue; XI. Coihaique, General Carrera; XII. Antártica Chilena, Magallanes,
Última Esperanza.
Orthotrichum subassimile Dusén, nom. nud., Dusén 1903b.
Orthotrichum subrupestre Thér. = Orthotrichum rupestre fide Lewinsky 1987.
Orthotrichum tristriatum Lewinsky
He 1998; Lewinsky 1984; Deguchi 1991b. Dist. RM. Chacabuco; VIII. Concepción; IX.
Cautín.
Orthotrichum truncatum Lewinsky & Deguchi
Tropicos 2009; Lewinsky & Deguchi 1989; Deguchi 1991b. Dist. VIII. Arauco, Biobío,
Concepción; IX. Cautín, Malleco.
Orthotrichum ulotoides Herzog = Orthotrichum incanum fide Matteri 2000.
Orthotrichum vittatum Cardot = Orthotrichum inclinatum fide Matteri 2000.
Oxyrrhynchium corralense (Lorentz) M. Fleisch. ex Reim. ≡ Eurhynchium corralense.
Oxyrrhynchium fuegianum (Cardot) Herzog ≡ Eurhynchium fuegianum.
Oxyrrhynchium praelongum (Hedw.) Warnst. ≡ Kindbergia praelonga.
Papillaria filipendula (Hook.f. & Wilson) A. Jaeger = Papillaria flexicaulis fide Ireland et al. 2007.
Papillaria crocea (Hampe) A. Jaeger
Ireland & Bellolio 2002 (as Papillaria pascuana, Barrera 2006; Espinosa 1941; Fuentes 1913;
Brotherus 1924). Dist. Easter I. Note: In accordance with Huttunen et al. (2008) the genus
Papillaria is treated here as distinct of the genus Meteorium.
Papillaria flexicaulis (Wilson) A. Jaeger
He 1998; Robinson 1975 (as Papillaria filipendula, Reimers 1926; Herzog & Hosseus 1938;
Bartram 1957; as Meteorium flexicaule, Müller 2009; Ireland et al. 2007). Dist. VII.
Cauquenes, Talca; VIII. Biobío, Concepción; IX. Cautín; XIV. Valdivia/Ranco; Juan
Fernández Is. Note: In accordance with Huttunen et al. (2008) the genus Papillaria is treated
here as distinct of the genus Meteorium.
Papillaria pascuana Thér. ex Broth. = Papillaria crocea.
Pararhacocarpus patagonicus (Broth.) J.-P. Frahm
Tropicos 2009; Buck 2004; He 1998; Frahm 1996; Matteri & Schiavone 2002 (as Rhacocarpus
patagonicus, Hässel et al. 1984; Dusén 1905a; Matteri 1985b). Dist. XIV. Valdivia/Ranco; XI.
Pentastichella aurea Dusén, nom. nud. = Zygodon pentastichus fide Calabrese 2006.
Pentastichella jaffuelii Thér. = Zygodon pentastichus fide Calabrese 2006.
Pentastichella pentasticha (Mont.) Müll. Hal. ≡ Zygodon pentastichus.
Pentastichella pentasticha var. strictifolia Thér. = Zygodon pentastichus fide Calabrese 2006.
Pentastichella robustula Dusén ex Paris = Zygodon bartramioides fide Calabrese 2006.
Peromnion magellanicum Sull. ≡ Brachymenium magellanicum.
Philibertiella ditrichoidea Cardot ≡ Ditrichum ditrichoideum.
Philibertiella nitens Herzog ≡ Ditrichum nitens.
Philonotis acicularis (Müll. Hal.) Kindb. = Philonotis polymorpha fide Ochyra et al. 2008.
Philonotis bartramii Matteri = Philonotis polymorpha fide Matteri 1985a.
Philonotis brevifolia Herzog = Philonotis krausei fide Seki 1974.
Philonotis cygnea (Sull.) D.G. Griffin & W.R. Buck
Griffin & Buck 1989 (as Bartramidula cygnea, Herzog 1922; Seki 1974; as Bartramia cygnea,
Mitten 1869). Dist. XIV. Valdivia/Ranco; XI. Aisén.
Philonotis depressinervis Cardot, nom. nud., Barrera 2006.
Philonotis elongata (Dism.) H.A. Crum & Steere
Philonotis exigua (Sull.) Broth. = Philonotis scabrifolia fide Wijk et al. 1967.
Philonotis glabrata Broth.

68 Müller
Philonotis glaucescens (Hornsch.) Broth.

He 1998. Dist. VII. Curicó.
Philonotis gourdonii Cardot = Philonotis polymorpha fide Ochyra et al. 2008.
Philonotis hastata (Duby) Wijk & Margad.
Ireland & Bellolio 2002 (as Philonotis laxissima, Espinosa 1941; Brotherus 1924). Dist. Easter
I.
Philonotis krauseana Broth. = Philonotis krausei.
Philonotis krauseana var. longinervis Broth.
Brotherus 1924b. Dist. Juan Fernández Is.
Philonotis krausei (Müll. Hal.) Broth.
Tropicos 2009; Larraín 2007b; Pereira et al. 2006; Schaumann 2005; Müller & Pereira 2006;
He 1998; Thériot 1934b, 1935; Herzog & Hosseus 1938; Herzog & Schwabe 1939; Herzog
1954; Bartram 1957; Matteri 1968; Seki 1974; Robinson 1975; Deguchi 1991a (as Philonotis
krauseana, Brotherus 1924b; as Philonotis brevifolia, Herzog 1954; Philonotis krausei fo.
latifolia, Thériot 1924.). Dist. V. Petorca, San Antonio, Valparaíso; RM. Chacabuco,
Cordillera, Santiago; VI. Cachapoal; VII. Cauquenes, Linares, Talca; VIII. Arauco, Biobío,
Osorno; XI. Aisén; Juan Fernández Is.
Philonotis laxissima Mitt. = Philonotis hastata.
Philonotis litorea Cardot & Broth.
Matteri 1989. Dist. Juan Fernández I.
Philonotis luteola E.B. Bartram = Philonotis polymorpha fide Matteri & Schiavone 2002.
Philonotis nigroflava Müll. Hal.
Ireland et al. 2007; He 1998; Matteri & Schiavone 2002; Dusén 1903b; Matteri 1985a (as
Philonotis parallela, Hässel et al. 1984; Dusén 1903b; Cardot 1908; Cardot & Brotherus 1923;
Matteri 1968; Seki 1974). Dist. V. Petorca; RM. Chacabuco, Santiago, Talagante; VI.
Colchagua; VII. Curicó; VIII. Arauco, Ñuble; IX. Cautín; X. Llanquihue; XI. Aisén; XII.
Philonotis parallela Dusén = Philonotis nigroflava fide Matteri 1985a.
Philonotis polymorpha (Müll. Hal.) Kindb.
(as Philonotis acicularis, He 1998; Matteri 1985a; as Philonotis luteola, Bartram 1946; as
Philonotis bartramii, Matteri 1973a; as Philonotis gourdonii, He 1998; Deguchi 1991a). Dist.
IV. Choapa; V. Aconcagua; RM. Santiago; VI. Colchagua; VII. Curicó; IX. Cautín; X. Osorno;
XII. Magallanes, Tierra del Fuego.
Philonotis scabrifolia (Hook.f. & Wilson) Braithw.
Tropicos 2009; Müller 2009; Hässel et al. 1984; Buck 2002; Espinosa 1941; Schaumann 2005;
He 1998; Matteri & Schiavone 2002; Cardot 1908; Herzog 1922, 1954; Cardot & Brotherus
1923; Brotherus 1924b; Roivainen 1934; Herzog & Schwabe 1939; Bartram 1952, 1957;
Matteri 1968; Seki 1974; Robinson 1975; Matteri 1985a, 1985b; Deguchi 1991a (as
Bartramidula exigua, Dusén 1903b; as Philonotis exigua, Brotherus 1904; as Bartramia
pusilla, Sullivant 1850). Dist. VII. Talca; VIII. Arauco, Biobío, Concepción, Ñuble; IX.
Cautín, Malleco; XIV. Valdivia/Ranco; X. Chiloé, Osorno; XI. Aisén; XII. Antártica Chilena,
Magallanes, Última Esperanza; Juan Fernández Is.
Philonotis trichophylla Herzog, hom. illeg., Herzog 1954.
Philonotis vagans (Hook. & Wilson) Mitt. var. vagans
Tropicos 2009; Goffinet et al. 2006; Hässel et al. 1984; Buck 2002; Espinosa 1941;
Schaumann 2005; He 1998; Matteri & Schiavone 2002; Mitten 1869; Dusén 1903b; Cardot
1908; Cardot & Brotherus 1923; Thériot 1924; Roivainen 1934; Bartram 1952, 1957; Seki
1974; Robinson 1975; Matteri 1985a, 1985b; Deguchi 1991a (as Bartramia vagans, Mitten
1869; as Rhizogonium reticulatum, Mitten 1869; as Goniobryum reticulatum, Cardot 1908;
Thériot 1934b; Herzog & Schwabe 1939; Herzog 1954; Seki 1974; Deguchi 1991b). Dist. RM.
Maipo; VIII. Arauco, Biobío, Ñuble; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Osorno;
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Mosses of Chile 69
XI. Aisén, Coihaique; XII. Antártica Chilena, Magallanes, Última Esperanza; Juan Fernández
Is.
Philonotis vagans var. evanidinervis Broth.
Barrera 2006; Brotherus 1924b. Dist. Juan Fernández Is.
Physcomitrium badium Broth.
Ireland et al. 2007; Herzog 1954. Dist. VIII. Concepción; X. Llanquihue.
Physcomitrium lorentzii Müll. Hal.
Ireland et al. 2007; Thériot 1934b (as Physcomitrium mahui, Robinson 1970). Dist. V.
Aconcagua; RM. Santiago; VIII. Arauco.
Physcomitrium mahui H. Rob. = Physcomitrium lorentzii fide Fife 1982.
Physcomitrium roseae R.S. Williams
Williams 1915. Dist. V. Valparaíso.
Pilopogon leptodus (Mont.) Broth. = Campylopus clavatus fide Sainsbury 1955.
Pilopogon schilleri Herzog & Thér.
Larraín 2007b; Hässel et al. 1984; Larraín 2007a; Schaumann 2005; Frahm 2005; Matteri
1985b, 1989; Matteri & Schiavone 2002; Herzog et al. 1933. Dist. X. Chiloé, Llanquihue,
Osorno, Palena; XII. Magallanes, Última Esperanza.
Pilotrichella cumingii (Müll. Hal.) Lorentz = Weymouthia mollis fide Mitten 1869.
Pilotrichella krausei Lorentz = Weymouthia cochlearifolia fide Cardot 1908.
Pinnatella callicostelloides (Broth. ex Thér.) Broth. ≡ Chileobryon callicostelloides.
Pinnatella macrosticta Broth. = Chileobryon callicostelloides fide Enroth 1992.
Plagiomnium ellipticum (Brid.) T.J. Kop.
Hässel et al. 1984; Buck 2002; Matteri & Schiavone 2002; Matteri 1985b; Deguchi 1991b (as
Mnium lepto-limbatum, Dusén 1903b; as Plagiomnium rugicum, T. Koponen 1968). Dist. XIV.
Valdivia/Ranco; XI. Coihaique; XII. Antártica Chilena, Magallanes, Tierra del Fuego, Última
Esperanza.
Plagiomnium rugicum (Laurer) T.J. Kop. = Plagiomnium ellipticum fide T. Koponen 1974.
Plagiothecium conostegium Herzog = Plagiothecium orthocarpum fide Ochyra et al. 2008.
Plagiothecium denticulatum (Hedw.) Schimp. var. denticulatum
Cardot 1908; Roivainen 1934; Matteri & Schiavone 2002. Dist. VIII. Ñuble; XII. Antártica
Chilena.
Plagiothecium denticulatum var. obtusifolium (Turner) Moore
(as Plagiothecium donnianum, Mitten 1869). Dist. XII. Antártica Chilena
Plagiothecium donnianum (Sm.) Mitt. = Plagiothecium denticulatum var. obtusifolium fide Wijk et al. 1967.
Plagiothecium georgicoantarcticum (Müll. Hal.) Kindb. = Plagiothecium orthocarpum fide Ochyra et al. 2008.
Plagiothecium lamprostachys (Hampe) A. Jaeger
Buck 2002. Dist. XII. Antártica Chilena.
Plagiothecium lucidulum (Hook. f. & Wilson) Mitt. = Plagiothecium lucidum fide Buck & Ireland 1989.
Plagiothecium lucidum (Hook. f. & Wilson) Paris
Hässel et al. 1984; Matteri & Schiavone 2002; Cardot 1908; Cardot & Brotherus 1923; Matteri
1985b (as Plagiothecium lucidulum, Mitten 1869; Dusén 1903b). Dist. XII. Antártica Chilena,
Plagiothecium orthocarpum Mitt.
Mitten 1869 (as Plagiothecium georgicoantarcticum, Matteri & Schiavone 2002; as
Plagiothecium conostegium, Schaumann 2005; Buck & Ireland 1989; Frey & Schaumann
2002). Dist. VIII. Concepción; IX. Cautín; XII. Antártica Chilena, Magallanes, Tierra del
Plagiothecium ovalifolium Cardot
Goffinet et al. 2006; Buck 2002; Ireland et al. 2007; Bednarek-Ochyra et al. 1999; Matteri &
Schiavone 2002; Cardot 1908; Seki 1974; Buck & Ireland 1989; Deguchi 1991b. Dist. VIII.
Biobío; IX. Malleco; XI. Aisén; XII. Antártica Chilena, Magallanes, Tierra del Fuego, Última
Esperanza.
Platydictya jungermannioides (Brid.) H.A. Crum

70 Müller
Ochyra & Matteri 2001; Matteri & Schiavone 2002. Dist. XII. Tierra del Fuego.
Platyhypnidium peruviense (R.S. Williams) Ochyra ≡ Rhynchostegium peruviense.
Platyneurum laticostatum (Cardot) Broth. = Platyneuron praealtum fide Ochyra & Bednarek-Ochyra 1997.
Platyneuron praealtum (Mitt.) Ochyra & Bednarek-Ochyra
Goffinet et al. 2006; Buck 2002; Ireland et al. 2007; Schaumann 2005; He 1998; Matteri &
Schiavone 2002 (as Platyneurum laticostatum, Hässel et al. 1984; Roivainen 1937; Herzog &
Hosseus 1938; Bartram 1952; Herzog 1954; Seki 1974; Matteri 1985b; Deguchi 1991a; as
Dicranum cirrhifolium, Dusén 1903b; as Dicranum laticostatum, Dusén 1905b; Cardot 1900,
1901, 1908; as Ditrichum praealtum, Dusén 1903b; Cardot 1908). Dist. VIII. Biobío, Ñuble;
IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Llanquihue, Osorno; XI. Aisén, Coihaique; XII.
Antártica Chilena, Magallanes, Última Esperanza, Tierra del Fuego.
Pleuridium andinum Herzog
Tropicos 2009. Dist. VIII. Concepción (Ireland & Bellolio 31965, MO), Ñuble (Ireland &
Bellolio 31527, MO).
Pleuridium costesii Thér.
Thériot 1921. Dist. V. Valparaíso.
Pleuridium macrothecium Dusén
Dusén 1905b; Thériot 1921. Dist. VIII. Concepción; IX. Malleco.
Pleuridium robinsonii (Mont.) Mitt.
Tropicos 2009; He 1998; Mitten 1869; Dusén 1905b; Thériot 1921; Brotherus 1924b; Herzog
& Hosseus 1938; Bartram 1952; Herzog 1954; Robinson 1975. Dist. V. Petorca, San Antonio,
Valparaíso; RM. Chacabuco, Cordillera, Melipilla, Talagante; VI. Cachapoal; VIII. Arauco,
Biobío, Concepción, Ñuble; Juan Fernández Is.
Pleurorthotrichum chilense Broth.
Tropicos 2009; Barrera 2006; Espinosa 1941; He 1998; Brotherus 1905, 1924a; Herzog 1954;
Lewinsky-Haapasaari 1994; Muñoz & Pisano 1947. Dist. II. Antofagasta; IV. Choapa, Limarí;
V. Aconcagua, Petorca.
Pogonatum alpinum (Hedw.) Röhl. ≡ Polytrichastrum alpinum.
Pogonatum liliputanum Thér. = Oligotrichum canaliculatum fide Robinson 1975.
Pogonatum oligodus (Kunze ex Müll. Hal.) Mitt. ≡ Pogonatum perichaetiale subsp. oligodus.
Pogonatum perichaetiale (Mont.) A. Jaeger subsp. oligodus (Kunze ex Müll. Hal.) Hyvönen
Villagran et al. 2003; He 1998; Hyvönen 1989 (as Pogonatum oligodus, Ireland et al. 2007;
Mitten 1869; Thériot 1915, 1917; Bartram 1952). Dist. V. San Antonio, Valparaíso; RM.
Santiago; VIII. Biobío, Concepción, Ñuble; XIV. Valdivia/Ranco; X. Chiloé.
Pogonatum spuriocirratum Broth. = Pogonatum cirratum subsp. fuscatum fide Hyvönen 1989.
Pohlia australis A.J. Shaw & Fife
Tropicos 2009. Dist. X. Llanquihue (Frey & Schaumann 01-378, MO, det. Shaw).
Pohlia chilensis (Mont.) A.J. Shaw
Suárez & Schiavone 2008; He 1998; Ireland et al. 2007; Shaw 1987 (as Webera costesii,
Thériot 1917, 1924; as Mielichhoferia chilensis, Mitten 1869; as Pohlia pluriseta, Robinson
1964). Dist. V. Los Andes, Valparaíso; RM. Chacabuco, Santiago, Talagante; VII. Curicó;
VIII. Arauco, Biobío, Concepción, Ñuble.
Pohlia cruda (Hedw.) Lindb.
Tropicos 2009; Müller 2009; Hässel et al. 1984; Buck 2002; Schaumann 2005; He 1998;
Matteri & Schiavone 2002; Mitten 1869; Herzog 1922; Seki 1974; Matteri 1985b; Ochi 1990;
Deguchi 1991a (as Webera cruda, Cardot 1900, 1901, 1908; Dusén 1903b; Cardot & Brotherus
1923; Roivainen 1934; Herzog 1954). Dist. RM. Chacabuco, Santiago; VII. Talca; VIII.
Arauco, Biobío, Concepción, Ñuble; IX. Malleco; X. Llanquihue, Osorno; XI. Aisén; XII.
Antártica Chilena, Magallanes, Última Esperanza.
Pohlia drummondii (Müll. Hal.) A.L. Andrews
Cardot & Brotherus 1923; Müller 2002 (as Pohlia inflexa, Hässel et al. 1984; Matteri 1985b).
Dist. VII. Talca; XII. Magallanes, Última Esperanza.
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Mosses of Chile 71
Pohlia humilis (Mont.) Broth.

(as Orthodontium humile, Mitten 1869). Dist. Chile australia.
Pohlia inflexa (Müll. Hal.) Wijk & Margad. = Pohlia drummondii fide Ochyra 1998.
Pohlia integra (Cardot) A.J. Shaw = Pohlia wilsonii fide Arts 2001b.
Pohlia lonchochaete (Dusén) Broth.
Buck 2002 (as Webera lonchochaete, Dusén 1903b). Dist. X. Llanquihue; XII. Antártica
Chilena.
Pohlia looseri (Thér.) S. He
He 1998 (as Webera looseri, Thériot 1935). Dist. RM. Santiago.
Pohlia magnifica (Herzog) S. He = Bryum australe fide Müller 2009.
Pohlia nutans (Hedw.) Lindb.
Goffinet et al. 2006; Hässel et al. 1984; Ireland et al. 2007; Schaumann 2005; He 1998; Matteri
& Schiavone 2002; Mitten 1869; Seki 1974; Matteri 1985b; Ochi 1990; Deguchi 1991a (as
Webera nutans, Dusén 1903b; Cardot 1908; Cardot & Brotherus 1923; Thériot 1924; Herzog
1954; as Webera sphagnadelphus, Dusén 1903b; Cardot 1908; Cardot & Brotherus 1923).
Dist. RM. Chacabuco, Cordillera, Maipo; VIII. Arauco, Ñuble; IX. Cautín; XIV.
Valdivia/Ranco; XI. Aisén; XII. Antártica Chilena, Magallanes, Última Esperanza.
Pohlia papillosa (Müll. Hal. ex A. Jaeger) Broth.
NY 2009. Dist. RM. Santiago (Buck 41545, NY).
Pohlia pluriseta Herzog = Pohlia chilensis fide Shaw 1987.
Pohlia sphagnicola (Bruch & Schimp.) Broth.
Müller 2009. Dist. XII. Magallanes.
Pohlia wahlenbergii (F. Weber & D. Mohr) A.L. Andrews var. wahlenbergii
Tropicos 2009; Ochyra et al. 2008; Buck 2002; Larraín 2005; Villagran et al. 2003;
Schaumann 2005; He 1998; Müller & Pereira 2006; Matteri & Schiavone 2002; Seki 1974;
Ochi 1990; Deguchi 1991a (as Mniobryum philonoteum, Herzog 1954; as Webera philonotea,
Dusén 1903b; Cardot 1908; as Webera albicans, Dusén 1903b; Thériot 1925; as Mniobryum
albicans, Cardot & Brotherus 1923; Reimers 1926; Roivainen 1934; Herzog & Hosseus 1938;
Herzog & Schwabe 1939; Herzog 1954; as Mniobryum alticaule var. crassinerve, Cardot &
Brotherus 1923). Dist. V. Los Andes, Petorca, San Antonio, Valparaíso; RM. Chacabuco,
Cordillera, Santiago; VI. Colchagua; VII. Cauquenes, Curicó, Talca; VIII. Arauco, Biobío,
Concepción, Ñuble; IX. Cautín; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno; XI.
Pohlia wahlenbergii var. glacialis (Brid.) E.F. Warb.
Hässel et al. 1984 (as Mniobryum alticaule, Herzog 1954; as Webera alticaulis, Dusén 1903b;
Cardot 1908; as Mniobryum amplirete, Herzog 1922). Dist. XI. Aisén; XII. Magallanes,
Última Esperanza.
Pohlia wilsonii (Mitt.) Ochyra
Müller 2009 (as Leptobryum wilsonii, Buck 2002; Matteri & Schiavone 2002; He 1998;
Bartram 1952; as Wollnya wilsonii, Herzog 1954; as Leptobryum pottiaceum, Dusén 1903b;
Cardot 1908; as Wollnya pottiacea, Herzog 1954; as Pohlia integra, Suárez & Schiavone 2005;
as Mniobryum aspillagae, Thériot 1924). Dist. II. Antofagasta; III. Atacama; IV. Elqui; V.
Valparaíso; RM. Santiago; VI. Colchagua; XII. Antártica Chilena, Magallanes.
Pohlia spec.
Polytrichadelphus dendroides (Brid. ex Hedw.) Mitt. ≡ Dendroligotrichum dendroides.
Polytrichadelphus horridus Mitt. = Polytrichadelphus magellanicus fide Robinson 1975.
Polytrichadelphus magellanicus (Hedw.) Mitt. subsp. magellanicus
Goffinet et al. 2006; Hässel et al. 1984; Stech et al. 2002; Buck 2002; Buck 2005a; Larraín
2005; Espinosa 1941; Schaumann 2005; He 1998; Matteri & Schiavone 2002; Mitten 1869;
Dusén 1903b; Cardot 1908; Herzog 1922; Cardot & Brotherus 1923; Brotherus 1924b;
Reimers 1926; Herzog & Schwabe 1939; Bartram 1957; Seki 1974; Robinson 1975; Matteri

72 Müller
1985b; Deguchi 1991b; Schiavone 1993 (as Polytrichadelphus horridus, Mitten 1869; Cardot
1908; Herzog & Schwabe 1939; Herzog 1954; as Polytrichadelphus robustus, Cardot 1908;
Cardot & Brotherus 1923). Dist. IX. Cautín; XIV. Valdivia/Ranco; X. Llanquihue, Osorno; XI.
Aisén, Coihaique; XII. Antártica Chilena, Magallanes, Tierra del Fuego, Última Esperanza;
Juan Fernández Is.
Polytrichadelphus minimus Cardot ≡ Notologitrichum minimum.
Polytrichadelphus robustus (Lindb.) Broth. = Polytrichadelphus magellanicus fide Schiavone 1993.
Polytrichadelphus squamosa (Hook.f. & Wilson) Mitt. ≡ Dendroligotrichum squamosum.
Polytrichadelphus stricticaulis Dusén, nom. nud., Dusén 1903b.
Polytrichastrum alpinum (Hedw.) G.L. Sm.
Müller 2009; Ireland et al. 2007; Matteri & Schiavone 2002 (as Pogonatum alpinum, Hässel et
al. 1984; Cardot 1908; as Polytrichum alpinum, Thériot 1935; Matteri 1985b; Polytrichum
alpinum var. integrifolium, Cardot & Brotherus 1923). Dist. VII. Talca; VIII. Ñuble; XIV.
Valdivia/Ranco; X. Osorno; XII. Magallanes, Tierra del Fuego, Última Esperanza.
Polytrichastrum formosum (Hedw.) G. L. Sm.
(as Polytrichum formosum, Matteri 1985b; Hässel et al. 1984). Dist. XII. Magallanes, Última
Esperanza.
Polytrichastrum longisetum (Sw. ex Brid.) G.L. Sm.
Díaz et al. 2008; Ochyra et al. 2008; Larraín 2007b; Hässel et al. 1984; Larraín 2005; Villagran
et al. 2003; He 1998; Matteri & Schiavone 2002; Deguchi 1991b (as Polytrichum gracile,
Herzog & Schwabe 1939; as Polytrichum longisetum, Matteri 1985b). Dist. XIV.
Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno; XI. Aisén; XII. Magallanes, Tierra del Fuego,
Última Esperanza.
Polytrichum alpinum Hedw. ≡ Polytrichastrum alpinum.
Polytrichum alpinum var. integrifolium Cardot & Broth. = Polytrichastrum alpinum fide Schiavone 1993.
Polytrichum elongatum P. Beauv.
Dusén 1903b; Cardot 1908. Dist. XII. Magallanes.
Polytrichum gracile Dicks. = Polytrichastrum longisetum fide Smith 1971.
Polytrichum juniperinum Hedw. var. juniperinum
Tropicos 2009; Goffinet et al. 2006; Hässel et al. 1984; Döbbeler 1999; Döbbeler & Triebel
2000; Pereira et al. 2006; Buck 2002; Larraín 2005; Villagran et al. 2003; He 1998; Müller &
Pereira 2006; Schiavone & Hyvönen 1993; Matteri & Schiavone 2002; Mitten 1869; Dusén
1903b; Cardot 1908; Reimers 1926; Roivainen 1934; Thériot 1935; Bartram 1952; Seki 1974;
Matteri 1985b; Deguchi 1991b (as Polytrichum juniperiforme, Mitten 1869; Polytrichum
juniperinum var. alpinum, Cardot & Brotherus 1923). Dist. V. San Antonio; RM. Cordillera,
Maipo, Santiago; VI. Colchagua; VII. Cauquenes, Curicó, Talca; VIII. Arauco, Biobío, Ñuble;
IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno; XI. Aisén,
Coihaique; XII. Antártica Chilena, Magallanes, Última Esperanza.
Polytrichum juniperinum var. affine (Funck) Brid.
Hässel et al. 1984; Matteri & Schiavone 2002; Matteri 1985b (as Polytrichum strictum var.
alpestre, Cardot & Brotherus 1923). Dist. XII. Magallanes, Última Esperanza.
Polytrichum juniperiforme Schimp. ex Mitt. = Polytrichum juniperinum fide Schiavone 1993.
Polytrichum juniperinum var. alpinum Schimp. = Polytrichum juniperinum fide Schiavone 1993.
Polytrichum juniperinum var. australe Walth., hom. illeg., Herzog & Schwabe 1939 (as
Polytrichum spegazzinii, Dusén 1903b).
Polytrichum patagonicum Müll. Hal. = Polytrichum piliferum fide Walther 1940.
Polytrichum piliferum Schreb. ex Hedw.
Tropicos 2009; Goffinet et al. 2006; Hässel et al. 1984; Buck 2002; Schaumann 2005; He
1998; Schiavone & Hyvönen 1991; Matteri & Schiavone 2002; Mitten 1869; Cardot 1900,
1901, 1908; Dusén 1903b; Reimers 1926; Herzog & Hosseus 1938; Herzog & Schwabe 1939;
Bartram 1952; Seki 1974; Matteri 1985b; Deguchi 1991b (as Polytrichum patagonicum, Dusén
1903b; as Polytrichum subpiliferum, Cardot 1900, 1901, 1908; as Polytrichum trachynotum,
Dusén 1903b). Dist. VII. Talca; VIII. Arauco, Biobío, Ñuble; IX. Cautín, Malleco; XIV.
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Valdivia/Ranco; X. Llanquihue, Osorno; XI. Aisén, Coihaique; XII. Antártica Chilena,

Polytrichum spegazzinii Müll. Hal. = Polytrichum juniperinum fide Walther 1940.
Polytrichum strictum Menzies ex Brid.
Goffinet et al. 2006; Buck 2002; Dusén 1903b; Cardot 1908; Ireland et al. 2007. Dist. VIII.
Ñuble; XII. Antártica Chilena, Magallanes.
Polytrichum strictum var. alpestre (Hoppe) Rabenh. = Polytrichum juniperinum var. affine fide Schiavone 1993.
Polytrichum subpiliferum Cardot = Polytrichum juniperinum fide Matteri & Schiavone 2002.
Polytrichum trachynotum Müll. Hal. = Polytrichum piliferum fide Walther 1940.
Porothamnium arbusculans (Müll. Hal.) M. Fleisch.
Tropicos 2009; Larraín 2007b; Hässel et al. 1984; Larraín 2005; Gunckel 1938; Villagran et al.
2003; He 1998; Matteri & Schiavone 2002; Herzog 1922, 1954, 1960; Cardot & Brotherus
1923; Brotherus 1924b; Herzog & Hosseus 1938; Herzog & Schwabe 1939; Seki 1974;
Robinson 1975; Matteri 1985b; Deguchi 1991b (as Hypnodendron naumanni, Cardot 1908; as
Porothamnium arbusculans var. complanatum, Cardot & Brotherus 1923; as Porothamnium
arbusculans fo. complanata, Reimers 1926; Porothamnium arbusculans fo. typica, Reimers
1926; as Thamnium arbusculans, Dusén 1903b; Cardot 1908; as Thamnium decumbens, Dusén
1903b; Cardot 1908). Dist. IV. Choapa; V. Petorca; VIII. Arauco, Biobío, Concepción, Ñuble;
IX. Malleco; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno; XI. Aisén; XII.
Porothamnium arbusculans var. complanatum (Müll. Hal.) Cardot = Porothamnium arbusculans.
Porothamnium arbusculans fo. complanata (Dusén) Reim. = Porothamnium arbusculans.
Porothamnium arbusculans fo. typica Reim. = Porothamnium arbusculans.
Porothamnium fasciculatum (Sw. ex Hedw.) M. Fleisch. ≡ Thamnobryum fasciculatum.
Porothamnium leucocaulon (Müll. Hal.) M. Fleisch.
Gunckel 1938; He 1998; Mitten 1869; Herzog 1922; Roivainen 1934; Herzog & Hosseus 1938;
Bartram 1952 (as Thamnium leucocaulon, Cardot 1908). Dist. VIII. Ñuble; IX. Cautín; XIV.
Valdivia/Ranco; X. Llanquihue, Osorno; XII. Magallanes.
Porothamnium panduraefolium (Müll. Hal.) M. Fleisch.
Tropicos 2009; Larraín 2005; Gunckel 1938; Villagran et al. 2003; He 1998; Müller & Pereira
2006; Mitten 1869; Thériot 1918; Herzog 1922, 1954, 1960; Herzog & Hosseus 1938; Deguchi
1991b (as Porothamnium panduraefolium fo. elongata, Reimers 1926; as Porothamnium
panduraefolium fo. typica, Reimers 1926; as Thamnium panduraefolium, Cardot & Brotherus
1923). Dist. VII. Cauquenes; VIII. Arauco, Concepción, Ñuble; IX. Cautín, Malleco; XIV.
Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno; XI. Aisén; XII. Magallanes.
Porothamnium panduraefolium fo. elongata Reim. = Porothamnium panduraefolium.
Porothamnium panduraefolium fo. typica Reim. = Porothamnium panduraefolium.
Porothamnium valdiviae (Müll. Hal.) M. Fleisch.
Tropicos 2009; Larraín 2007b; Pereira et al. 2006; Larraín 2005; Gunckel 1938; Villagran et al.
2003; He 1998; Mitten 1869; Thériot 1918; Cardot & Brotherus 1923; Roivainen 1934; Herzog
& Hosseus 1938; Herzog 1954, 1960; Robinson 1975; Deguchi 1991b (as Porotrichum
valdiviae, Bartram 1952; as Porothamnium valdiviae fo. typica, Reimers 1926; as
Porothamnium valdiviae fo. elongata, Reimers 1926; as Thamnium valdiviae, Cardot 1908).
Dist. IV. Choapa; VII. Talca; VIII. Biobío, Ñuble; IX. Cautín, Malleco; XIV. Valdivia/Ranco;
X. Chiloé, Llanquihue, Osorno; XI. Aisén, Coihaique; Juan Fernández Is.
Porothamnium valdiviae fo. typica Reim. = Porothamnium valdiviae.
Porothamnium valdiviae fo. elongata Reim. = Porothamnium valdiviae.
Porotrichodendron porteri (Thér.) Broth.
(as Porotrichum porteri, Thériot 1917). Dist. VII. Linares.
Porotrichum callicostelloides Broth. ex Thér. ≡ Chileobryon callicostelloides.
Porotrichum callicostelloides var. distichum Thér. = Chileobryon callicostelloides fide Enroth 1992.
Porotrichum chilense Thér.
Tropicos 2009; He 1998; Thériot 1924. Dist. V. Valparaíso; VIII. Biobío, Ñuble; XIV.
Valdivia/Ranco; X. Chiloé, Osorno.

74 Müller
Porotrichum fasciculatum (Hedw.) Mitt. ≡ Thamnobryum fasciculatum.

Porotrichum korthalsianum (Dozy & Molk.) Mitt.
Tropicos 2009; Schaumann 2005; He 1998. Dist. V. Petorca; VIII. Arauco, Biobío,
Concepción; IX. Cautín; XIV. Valdivia/Ranco; X. Llanquihue, Osorno, Palena.
Porotrichum lancifrons (Hampe) Mitt.
Ireland et al. 2007; He 1998. Dist. VIII. Arauco, Biobío, Concepción, Ñuble; IX. Cautín; XIV.
Valdivia/Ranco; XI. Aisén.
Porotrichum porteri Cardot & Thér. ≡ Porotrichodendron porteri.
Porotrichum valdiviae (Müll. Hal.) Mitt. ≡ Porothamnium valdiviae.
Pottia acutidentata Thér. & Cardot = Hennediella antarctica fide Cano 2008.
Pottia anderssonii Sauerb., hom. illeg. = Hennediella antarctica fide Cano 2008.
Pottia antarctica (Ångström) Müll. Hal. ≡ Hennediella antarctica.
Pottia austrogeorgica Cardot = Hennediella antarctica fide Ochyra 1998.
Pottia austrogeorgica var. microphylla Cardot & Broth. = Hennediella antarctica fide Cano 2008.
Pottia chubutensis Cardot & Broth. = Hennediella heimii fide Cano & Gallego 2008.
Pottia flavipes Mont. = Hennediella heimii fide Cano 2008.
Pottia fuscomucronata Müll. Hal. = Hennediella antarctica fide Cano 2008.
Pottia heimii (Hedw.) Hampe ≡ Hennediella heimii.
Pottia heimii var. flavipes (Mont.) Warnst. = Hennediella heimii fide Cano 2008.
Pottia heimii var. magellanica Schimp. = Hennediella heimii fide Cano 2008.
Pottia heimii var. maxima Cardot = Hennediella heimii fide Cano 2008.
Pottia heimii var. thaxteri Cardot & Thér. = Hennediella antarctica fide Cano 2008.
Pottia intermedia (Turner) Fürnr. = Tortula modica.
Pottia macrocarpa Schimp. ≡ Costesia macrocarpa.
Pottia magellanica Schimp. ex Paris, hom. illeg. = Hennediella antarctica fide Wijk et al. 1967.
Pottia megapoda Müll. Hal. = Hennediella heimii fide Warnstorf 1916.
Pottia pellucida Cardot, nom. nud., Cardot 1911.
Pottia spegazzinii Müll. Hal. ≡ Hennediella heimii var. spegazzinii.
Pottia truncata (Hedw.) Bruch & Schimp. ≡ Tortula truncata.
Pseudocrossidium apiculatum R.S. Williams
He 1998; Thériot 1923; Cano 2003. Dist. XV. Parinacota; III. Copiapó; IV. Choapa, Elqui,
Limarí; RM. Chacabuco, Cordillera, Santiago; VI. Cardenal Caro.
Pseudocrossidium chilense R.S. Williams
Jiménez & Cano 2007; He 1998; Williams 1915. Dist. IV. Elqui; V. Valparaíso; RM.
Chacabuco, Santiago; VI. Cardenal Caro.
Pseudocrossidium crinitum (Schultz) R.H. Zander
Matcham et al. 2007; Larraín 2005; He 1998; Matteri & Schiavone 2002 (as Barbula
arenicola, Dusén 1906a; as Barbula depressa, Thériot 1918, 1921; Costes 1921; Cardot &
Brotherus 1923; as Barbula depressa var. denticulata, Thériot 1918; as Barbula flagellaris var.
denticulata, Dusén 1906a; Cardot 1908; as Barbula flagellaris var. gracilis, Dusén 1906a; as
Tortula pilifera, Hässel et al. 1984; Hezog 1960; Seki 1974; Matteri 1985b; as Tortula pilifera
var. denticulata, Hässel et al. 1984; Matteri 1985b; as Barbula pilifera, Thériot 1928; Herzog
& Hosseus 1938; Bartram 1952; Herzog 1954; Deguchi 1991b; as Tortula purpureo-velutina,
Herzog 1954; as Tortula subglacialis, Thériot 1917, 1921; as Barbula flagellaris, Costes
1921). Dist. IV. Elqui, Limarí; V. Aconcagua, Petorca, San Antonio, Valparaíso; RM.
Cordillera, Santiago, Talagante; VI. Colchagua; VII. Cauquenes, Linares, Talca; VIII. Biobío,
Ñuble; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Llanquihue, Osorno; XI. Aisén,
Pseudocrossidium elatum (R.S. Williams) Delgad.
Pseudocrossidium excavatum (Mitt.) R.S. Williams
Cano & Gallego 2008 (as Tortula santiagensis, Espinosa 1941; Brotherus 1924a). Dist. RM.
Santiago.
Pseudocrossidium hornschuchianum (Schultz) R.H. Zander
Müller 2002. Dist. XII. Magallanes.
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Mosses of Chile 75
Pseudocrossidium leucocalyx (Mont.) Thér.

Espinosa 1941; He 1998; Thériot 1923; Brotherus 1924a (as Tortula leucocalyx, Mitten 1869;
as Barbula pachyneura, Dusén 1906a; Thériot 1917; Costes 1921; as Tortula perarmata,
Brotherus 1924a; Espinosa 1941). Dist. IV. Coquimbo, Limarí; V. Petorca, Quillota,
Valparaíso; RM. Santiago.
Pseudocrossidium replicatum (Taylor) R.H. Zander
He 1998; Deguchi 1991b (as Barbula replicata, Bartram 1943, 1952). Dist. II. Antofagasta;
IV. Elqui; RM. Cordillera, Santiago.
Pseudodistichium atlanticum Dixon = Ditrichum conicum fide Robinson 1972.
Pseudodistichium austrogeorgicum Cardot = Ditrichum hyalinum fide Ochyra 1999a.
Pseudodistichium austrogeorgicum var. longifolium Broth. = Ditrichum hyalinum fide Ochyra 1999a.
Pseudodistichium fuegianum Roiv. = Ditrichum brotherusii fide Seppelt 1982.
Pseudodistichium taitaoense J. Froehl. = Ditrichum conicum fide Robinson 1972.
Pseudoleskea chilensis (Lorentz) Ochyra = Cratoneuropsis relaxa subsp. minor fide Ochyra et al. 2008.
Pseudoleskea filum (Müll. Hal.) Paris = Cratoneuropsis relaxa subsp. minor fide Ochyra et al. 2008.
Pseudoleskea fuegiana (Besch.) Broth. = Cratoneuropsis relaxa subsp. minor fide Ochyra et al. 2008.
Pseudoleskea fuegiana var. gracilis Cardot & Broth. = Cratoneuropsis relaxa subsp. minor fide Ochyra et al. 2008.
Pseudoleskea fuegiana var. skottsbergii Cardot = Cratoneuropsis relaxa subsp. minor fide Ochyra et al. 2008.
Pseudoleskea lurida Cardot = Cratoneuropsis relaxa subsp. minor fide Ochyra et al. 2008.
Pseudoscleropodium purum (Hedw.) M. Fleisch.
(as Scleropodium purum, Tropicos 2009; Goffinet et al. 2006). Dist. XIV. Valdivia/Ranco
(Crosby 11569, MO).
Pseudotaxiphyllum elegans (Brid.) Z. Iwats.
Müller 2009; Ireland et al. 2007. Dist. VII. Talca; VIII. Arauco.
Psilopilum angulatum Cardot & Broth. = Notoligotrichum minimum fide Matteri & Schiavone 2002.
Psilopilum antarcticum Müll. Hal. = Notoligotrichum trichodon fide Smith 1971.
Psilopilum antarcticum var. densifolium Cardot & Broth. = Notoligotrichum trichodon fide Schiavone 1993.
Psilopilum compressum (Hook.f. & Wilson) Mitt. ≡ Atrichopsis compressa.
Psilopilum cuspidatum Dusén = Notoligotrichum tapes fide Wijk et al. 1967.
Psilopilum magellanicum Dusén = Notoligotrichum trichodon fide Smith 1971.
Psilopilum minimum (Cardot) G.L. Smith ≡ Notoligotrichum minimum.
Psilopilum tapes (Müll. Hal.) Paris var. apiculatum Cardot ≡ Notoligotrichum tapes var. apiculatum.
Psilopilum trichodon (Hook.f. & Wilson) Mitt. ≡ Notoligotrichum trichodon.
Pterogoniadelphus hollermayeri (Thér.) Ochyra & Zijlstra = Acroschisma wilsonii fide Müller 2008.
Pterygoneurum ovatum (Hedw.) Dixon
Matcham et al. 2007; Matcham & Duckett in Blockeel et al. 2008a. Dist. XII. Magallanes.
Pterygophyllum anomalum (Schwägr.) Mitt. ≡ Achrophyllum anomalum.
Pterygophyllum anomalum var. pallidum Cardot & Broth. ≡ Achrophyllum anomalum var. pallidum.
Pterygophyllum crassirete Matteri ≡ Achrophyllum crassirete.
Pterygophyllum denticulatum (Hook.f. & Wilson) Mitt. = Achrophyllum dentatum fide Thériot 1917.
Pterygophyllum haesselianum Matteri ≡ Achrophyllum haesselianum.
Pterygophyllum lamellatum Dusén, nom. nud., Dusén 1903b.
Pterygophyllum magellanicum Besch. ≡ Achrophyllum magellanicum fide Matteri 1979.
Pterygophyllum magellanicum var. oligodontum Matteri ≡ Achrophyllum magellanicum var. oligodontum.
Pterygophyllum obscurum (Mont.) Mitt., hom. illeg. = Achrophyllum anomalum fide Robinson 1975.
Pterygophyllum obscurum fo. thermalis Herzog = Achrophyllum anomalum fide He 1998.
Pterygophyllum tenuinerve Broth. ≡ Achrophyllum tenuinerve.
Ptychomitrium chimborazense (Spruce ex Mitt.) A. Jaeger
Cao et al. 2006. Dist. Chile: Bojin de Arrayun.
Ptychomitrium chimborazense var. obtusifolium (Thér.) Broth. ≡ Ptychomitrium hieronymi var. obtusifolium.
Ptychomitrium crispatum (Hedw.) A. Jaeger
Cao et al. 2003. Dist. Juan Fernández Is.
Ptychomitrium cumingii Duby = Ptychomitrium fernandesianum fide Greene 1986.
Ptychomitrium deltorii (Thér.) Broth.
He 1998; Guo & Cao 1999; Thériot 1921 (as Brachysteleum deltori, Thériot 1921). Dist. VIII.
Concepción; IX. Malleco.
Ptychomitrium fernandesianum (Mitt.) A. Jaeger var. fernandesianum

76 Müller
Espinosa 1941; Müller & Pereira 2006; Brotherus 1924b; Robinson 1975 (as Brachysteleum
fernandesianum, Thériot 1921; as Glyphomitrium fernandesianum, Mitten 1869; as
Ptychomitrium cumingii, Thériot 1935). Dist. VII. Cauquenes; XIV. Valdivia/Ranco; X.
Osorno; Juan Fernández Is.
Ptychomitrium fernandesianum var. majus Broth.
Brotherus 1924b. Dist. Juan Fernández Is.
Ptychomitrium hieronymi Besch. var. obtusifolium (Thér.) T. Cao & Guo Shui-liang
Cao et al. 2006 (as Ptychomitrium chimborazense var. obtusifolium, Brotherus 1925; as
Brachysteleum chimborazense var. obtusifolium, Thériot 1923). Dist. V. Aconcagua. RM.
Santiago. “Cajon del Arragan”.
Ptychomitrium ligulatum (Mitt.) A. Jaeger
Matteri & Schiavone 2002 (as Glyphomitrium ligulatum, Mitten 1869; Cardot 1908). Dist. XII.
Antártica Chilena
Ptychomitrium sellowianum (Müll. Hal.) A. Jaeger
Ireland et al. 2007. Dist. VIII. Arauco, Concepción, Ñuble.
Ptychomitrium subcylindricum Thér.
Ireland & Bellolio 2002; Thériot 1937. Dist. Easter I.
Ptychomniella ptychocarpa (Schwägr.) W.R. Buck, C. J. Cox, A. J. Shaw & Goffinet
Larraín 2007b (as Ptychomnion ptychocarpon, Larraín 2005; Espinosa 1941; Villagran et al.
2003; Schaumann 2005; He 1998; Mitten 1869; Dusén 1903b; Cardot 1908; Thériot 1918,
1935; Cardot & Brotherus 1923; Brotherus 1924b; Reimers 1926; Roivainen 1934; Herzog
1922, 1954, 1960; Herzog & Hosseus 1938; Herzog & Schwabe 1939; Bartram 1952, 1957;
Seki 1974; Robinson 1970, 1975; Deguchi 1991b; Kühnemann & Goncalves Carralves 1975).
Dist. V. Valparaíso; VI. O'Higgins; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Chiloé,
Llanquihue, Osorno, Palena; XI. Aisén, General Carrera; XII. Magallanes; Juan Fernández Is.
Ptychomnion cygnisetum (Müll. Hal.) Kindb.
Díaz et al. 2008; Tropicos 2009; Goffinet et al. 2006; Larraín 2007b; Hässel et al. 1984;
Matcham et al. 2007; Buck 2002; Larraín 2005; Espinosa 1941; Villagran et al. 2003, 2004;
Schaumann 2005; He 1998; Matteri & Schiavone 2002; Muñoz & Pisano 1947; Dusén 1903b;
Cardot 1908; Thériot 1915; Herzog 1922, 1954, 1960; Cardot & Brotherus 1923; Brotherus
1924a; Reimers 1926; Herzog & Hosseus 1938; Herzog & Schwabe 1939; Seki 1974;
Robinson 1970, 1975; Hattaway 1984; Matteri 1985b; Deguchi 1991b; Kühnemann &
Goncalves Carralves 1975 (as Ptychomnion subaciculare, Espinosa 1941; Dusén 1903b;
Cardot 1908; Cardot & Brotherus 1923; Brotherus 1924b; Bartram 1957; Robinson 1970; Seki
1974; Deguchi 1991b; Kühnemann & Goncalves Carralves 1975; as Ptychomnion horridum,
Cardot & Brotherus 1923; Kühnemann & Goncalves Carralves 1975). Dist. IV. Coquimbo,
Limarí; V. Aconcagua; VII. Linares; VIII. Arauco; IX. Cautín, Malleco; XIV. Valdivia/Ranco;
X. Chiloé, Llanquihue, Osorno, Palena; XI. Aisén; XII. Antártica Chilena, Magallanes, Tierra
Ptychomnion densifolium (Brid.) A. Jaeger
Díaz et al. 2008; Larraín 2007b; Hässel et al. 1984; Buck 2005b; Tangney & Fife 1997; Matteri
& Schiavone 2002; Cardot 1908; Cardot & Brotherus 1923; Roivainen 1934; Robinson 1970;
Seki 1974; Matteri 1985b; Kühnemann & Goncalves Carralves 1975 (as Ptychomnion
densifolium var. gracile, Brotherus & Cardot 1923). Dist. XIV. Valdivia/Ranco; X. Chiloé,
Osorno; XI. Aisén; XII. Antártica Chilena, Magallanes, Tierra del Fuego, Última Esperanza.
Ptychomnion densifolium var. gracile Cardot & Broth. = Ptychomnion densifolium fide Hattaway 1984.
Ptychomnion falcatulum Broth.
Villagran et al. 2003; Brotherus 1924b; Robinson 1975; Kühnemann & Goncalves Carralves
1975 (as Ptychomnion falcatulum var. gracilescens, Barrera 2006; Espinosa 1941; Brotherus
1924b). Dist. XIV. Valdivia/Ranco; X. Chiloé; Juan Fernández Is.
Ptychomnion falcatulum var. gracilescens Broth. = Ptychomnion falcatulum fide Hattaway 1984.
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Ptychomnion horridum Cardot & Broth.= Ptychomnion cygnisetum fide Matteri & Schiavone 2002.
Ptychomnion ptychocarpon (Schwägr.) Mitt. ≡ Ptychomniella ptychocarpa.
Ptychomnion subaciculare Besch. = Ptychomnion cygnisetum fide Hattaway 1984.
Pulvinella albicans Broth. & Herzog = Sauloma tenella fide Buck 1987.
Pyrrhobryum mnioides (Hook.) Manuel
Tropicos 2009; Goffinet et al. 2006; Matteri & Piñero 1987; Larraín 2007b; Matcham et al.
2007; Buck 2002; Larraín 2005; Villagran et al. 2003; Schaumann 2005; He 1998; Müller &
Pereira 2006; Frahm et al. 2003; Matteri & Schiavone 2002 (as Rhizogonium mnioides, Hässel
et al. 1984; Espinosa 1941; Mitten 1869; Dusén 1903b; Cardot 1908; Thériot 1915, 1918,
1934b; Herzog 1922, 1954, 1960; Cardot 1905; Cardot & Brotherus 1923; Brotherus 1924b;
1952; Seki 1974; Robinson 1975; Matteri 1985b; Deguchi 1991b; as Aulacomnium chilense,
Müller 1843). Dist. V. Aconcagua; RM. Santiago; VII. Cauquenes, Linares, Talca; VIII.
Arauco, Biobío, Concepción, Ñuble; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Chiloé,
Llanquihue, Osorno, Palena; XI. Aisén; XII. Antártica Chilena, Magallanes, Tierra del Fuego,
Pyrrhobryum spiniforme (Hedw.) Mitt.
Racomitrium andreaeoides Herzog = Bucklandiella sudetica fide Bednarek-Ochyra 1993.
Racomitrium austrocanescens Dusén = Racomitrium pruinosum fide Vitt & Marsh 1988.
Racomitrium austrogeorgicum Paris = Bucklandiella sudetica fide Frisvoll 1986.
Racomitrium austrogeorgicum var. kranckii Roiv. = Bucklandiella sudetica fide Ochyra 1998.
Racomitrium bartramii (Roiv.) H. Rob. ≡ Bucklandiella bartramii.
Racomitrium canescens (Hedw.) Brid. ≡ Niphotrichum canescens.
Racomitrium convolutum Mont. = Bucklandiella didyma fide Ochyra et al. 2008.
Racomitrium crispipilum (Taylor) A. Jaeger ≡ Bucklandiella crispipila.
Racomitrium crispulum (Hook.f. & Wilson) Hook.f. & Wilson ≡ Bucklandiella crispula.
Racomitrium crispulum var. rupestre (Hook.f. & Wilson) Dixon = Bucklandiella rupestris fide Matteri 2003a.
Racomitrium didymum (Mont.) Lorentz ≡ Bucklandiella didyma.
Racomitrium didymum fo. obtusa Herzog, nom. nud., Herzog & Hosseus 1938.
Racomitrium flavescens Cardot = Bucklandiella didyma fide Ochyra et al. 2008.
Racomitrium flavo-pallidum Dusén, nom. nud., Dusén 1903b.
Racomitrium geminatum Roiv. = Bucklandiella crispipila fide Deguchi 1984.
Racomitrium geronticum Müll. Hal. ≡ Racomitrium lanuginosum subsp. geronticum.
Racomitrium heterostichoides Cardot ≡ Bucklandiella heterostichoides.
Racomitrium heterostichoides var. acutifolium Dusén = Bucklandiella crispula fide Clifford 1955.
Racomitrium hypnoides Lindb., hom. illeg. = Racomitrium lanuginosum fide Wijk et al. 1967.
Racomitrium hypnoides fo. gerontium (Müll. Hal.) Dusén ≡ Racomitrium lanuginosum subsp. geronticum.
Racomitrium hypnoides fo. glaciale (Müll. Hal.) Dusén
Dusén 1906b. Dist. XII. Magallanes.
Racomitrium integripilum Dusén = Bucklandiella crispipila fide Deguchi 1984.
Racomitrium laevigatum A. Jaeger ≡ Codriophorus laevigatus.
Racomitrium lamprocarpum (Müll. Hal.) A. Jaeger ≡ Bucklandiella lamprocarpa.
Racomitrium lanuginosum (Hedw.) Brid.
Ochyra et al. 2008; Larraín 2007b; Hässel et al. 1984; Buck 2002; Eaton 1892; Larraín 2005;
Espinosa 1941; Villagran et al. 2003; He 1998; Matteri & Schiavone 2002; Mitten 1869;
Cardot 1900, 1901, 1908; Dusén 1903b; Thériot 1921; Cardot & Brotherus 1923; Brotherus
1924b; Roivainen 1955a; Bartram 1957; Seki 1974; Robinson 1975; Matteri 1985b; Deguchi
1991a (as Racomitrium hypnoides, Dusén 1906b; Reimers 1926; Herzog & Hosseus 1938;
Herzog 1954, 1960; as Racomitrium puccioanum, Vitt & Marsh 1988). Dist. V. Valparaíso;
VII. Linares; VIII. Biobío; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue,
Osorno; XI. Aisén; XII. Antártica Chilena, Magallanes, Última Esperanza, Tierra del Fuego;
Juan Fernández Is.
Racomitrium lanuginosum subsp. geronticum (Müll. Hal.) Vitt & C. Marsh

78 Müller
Ochyra et al. 2002 (as Racomitrium hypnoides fo. gerontium, Dusén 1906b). Dist. VIII.
Concepción; XIV. Valdivia/Ranco; X. Osorno; XII. Tierra del Fuego, Última Esperanza; Juan
Fernández Is.
Racomitrium lanuginosum var. pruinosum Wilson ≡ Racomitrium pruinosum.
Racomitrium limbatum E.B. Bartram = Bucklandiella lamprocarpa fide Deguchi 1984.
Racomitrium looseri Thér. = Bucklandiella lamprocarpa fide Bednarek-Ochyra & Ochyra 1994.
Racomitrium loriforme Dusén = Codriophorus laevigatus fide Robinson 1975; Deguchi 1984.
Racomitrium membranaceum (Mitt.) Paris = Bucklandiella crispula fide Wijk et al. 1967.
Racomitrium nigritum (Müll. Hal.) A. Jaeger = Bucklandiella didyma fide Deguchi 1984.
Racomitrium orthotrichaceum (Müll. Hal.) Paris ≡ Bucklandiella orthotrichacea.
Racomitrium pachydictyon Cardot ≡ Bucklandiella pachydictyon.
Racomitrium pachyneuron Herzog = Bucklandiella lamprocarpa fide Ochyra 1993b.
Racomitrium pachyneuron fo. aquaticum Herzog = Bucklandiella lamprocarpa fide Ochyra 1993b.
Racomitrium patagonicum Bednarek-Ochyra & Ochyra
Larraín 2007b; Matcham et al. 2007; Ireland et al. 2005; Bednarek-Ochyra & Ochyra 2003.
Dist. VII. Linares; VIII. Arauco, Biobío, Concepción, Ñuble; IX. Cautín, Malleco; XIV.
Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno; XII. Magallanes, Última Esperanza.
Racomitrium plicatum Herzog = Racomitrium laevigatum fide Bednarek-Ochyra 2006.
Racomitrium pruinosum (Wilson) Müll. Hal.
Matteri & Schiavone 2002 (as Racomitrium austro-canescens, Dusén 1906b; as Racomitrium
lanuginosum var. pruinosum, Bartram 1952). Dist. XIV. Valdivia/Ranco; X. Llanquihue; XII.
Magallanes. Note: The species is restricted to Australasia; the records from southern South
America are very doubtful (Vitt & Marsh 1988, Bednarek-Ochyra et al. in Blockeel et al.
2009).
Racomitrium ptychophyllum (Mitt.) Mitt. ≡ Bucklandiella ptychophylla.
Racomitrium puccioanum De Not. = Racomitrium lanuginosum fide Vitt & Marsh 1988.
Racomitrium rupestre (Hook.f. & Wilson) Hook.f. & Wilson ≡ Bucklandiella rupestris.
Racomitrium scabrifolium E.B. Bartram = Codriophorus laevigatus fide Robinson 1975; Deguchi 1984.
Racomitrium senile Schimp., nom. nud. = Racomitrium patagonicum fide Bednarek-Ochyra & Ochyra 2003.
Racomitrium skottsbergii Cardot & Broth. = Bucklandiella sudetica fide Matteri & Schiavone 2002.
Racomitrium stenocladoides Roiv. = Bucklandiella orthotrichacea fide Deguchi 1984.
Racomitrium stenocladum Dusén = Bucklandiella orthotrichacea fide Deguchi 1984.
Racomitrium stenocladum var. obtusum Dusén = Bucklandiella rupestris fide Deguchi 1984.
Racomitrium striatipilum Cardot = Bucklandiella crispipila fide Deguchi 1984, 1987.
Racomitrium subcrispipilum (Müll. Hal.) A. Jaeger = Bucklandiella crispula fide Wijk et al. 1962.
Racomitrium sublamprocarpum (Müll. Hal.) Paris = Bucklandiella lamprocarpa fide Bednarek-Ochyra & Ochyra
1994.
Racomitrium subnigritum (Müll. Hal.) Paris = Bucklandiella lamprocarpa fide Deguchi 1984.
Racomitrium subrupestre Roiv. = Bucklandiella didyma fide Ochyra et al. 2008.
Racomitrium substenocladum Cardot = Bucklandiella sudetica fide Ochyra 1998.
Racomitrium substriatipilum Roiv. = Bucklandiella crispipila fide Deguchi 1984.
Racomitrium subulifolium Cardot = Bucklandiella crispula fide Wijk et al. 1967.
Racomitrium sudeticum (Funck) Bruch & Schimp. ≡ Bucklandiella sudetica.
Racomitrium symphyodontum (Müll. Hal.) Paris = Bucklandiella didyma fide Deguchi 1984.
Racomitrium symphyodontum var. dusenii Cardot = Bucklandiella didyma fide Deguchi 1984.
Racomitrium symphyodontum fo. flavescens (Cardot) Cardot = Bucklandiella didyma fide Ochyra et al. 2008.
Racomitrium symphyodontum var. muticum Cardot = Bucklandiella pachydictyon fide Cardot 1908.
Racomitrium willii (Müll. Hal.) Kindb. = Codriophorus laevigatus fide Bednarek-Ochyra 2006.
Racopilum chilense Dusén = Racopilum cuspidigerum fide Zanten 1993.
Racopilum cuspidigerum (Schwägr.) Ångström
Ireland et al. 2007; Espinosa 1941; Ireland & Bellolio 2002; Zanten 2006; Brotherus 1924 (as
Racopilum chilense, Herzog & Hosseus 1938; Deguchi 1991b). Dist. VIII. Arauco,
Concepción; IX. Cautín; XIV. Valdivia/Ranco; X. Llanquihue, Osorno; Juan Fernández Is.;
Easter I.
Racopilum cuspidigerum var. convolutaceum (Müll. Hal.) Zanten & Dijkstra
Zanten 2006. Dist. XIV. Valdivia/Ranco; X. Llanquihue; Easter I.
Racopilum fernandezianum Cardot
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Espinosa 1941; Brotherus 1924b; Thériot 1927a; Herzog 1954; Bartram 1957; Robinson 1975.
Dist. X. Osorno; Juan Fernández Is.
Racopilum tomentosum (Hedw.) Brid.
Larraín 2007a; Robinson 1975. Dist. XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno;
Juan Fernández Is.
Renauldia chilensis Thér.
Thériot 1926. Dist. VIII. Concepción.
Rhabdoweisia crispata (Dicks.) Lindb.
Tropicos 2009; Robinson 1975. Dist. VIII. Concepción (Ireland & Bellolio 30933, MO); Juan
Fernández Is.
Rhabdoweisia fugax (Hedw.) Bruch & Schimp.
Ireland et al. 2007. Dist. VIII. Arauco, Concepción.
Rhacocarpus humboldtii (Hook.f.) Lindb. = Rhacocarpus purpurascens fide Bowers 1974.
Rhacocarpus orbiculata (Mitt.) Paris = Rhacocarpus purpurascens fide Frahm 1996.
Rhacocarpus patagonicus Broth. ≡ Pararhacocarpus patagonicus.
Rhacocarpus purpurascens (Brid.) Paris
Hässel et al. 1984; Buck 2004; Larraín 2005; He 1998; Matteri & Schiavone 2002; Frahm
1996; Seki 1974; Robinson 1975; Matteri 1985b; Deguchi 1991a (as Racomitrium humboldtii,
Mitten 1869; Cardot 1908; Cardot & Brotherus 1923; Brotherus 1924b; as Hedwigia
orbiculata, Mitten 1869; as Rhacocarpus orbiculata, Paris 1903-1906). Dist. XIV.
Valdivia/Ranco; X. Chiloé, Llanquihue; XI. Aisén; XII. Antártica Chilena, Magallanes, Tierra
Rhamphidium brevifolium (Hampe & Lorentz) Broth.
Brotherus 1924c. Dist. Chile, without exact locality.
Rhamphidium laetum (Kunze ex Müll. Hal.) Broth.
Brotherus 1902-1909, 1924c (as Tortula laeta, Mitten 1869). Dist. Chile australis, without
exact locality.
Rhaphidorrhynchium amoenum (Hedw.) M. Fleisch.
Hässel et al. 1984; Ireland et al. 2007; Matteri & Schiavone 2002; Matteri 1985b (as
Rhaphidostegium amoenum, Dusén 1903b; Cardot 1908; as Sematophyllum amoenum, Mitten
1869; Beever et al. 1992). Dist. VIII. Arauco, Concepción; XII. Antártica Chilena, Magallanes,
Última Esperanza.
Rhaphidorrhynchium berberidis (Dusén) Broth.
Seki 1974 (as Ectropothecium berberidis, Dusén 1905; Cardot 1908; Rhaphidostegium
berberidis, Dusén 1903b). Dist. XI. Aisén.
Rhaphidorrhynchium callidum (Mont.) Broth.
Tropicos 2009; Larraín 2007b; Hässel et al. 1984; Larraín 2005; Villagran et al. 2003; He
1998; Müller & Pereira 2006; Matteri & Schiavone 2002; Reimers 1926; Herzog & Hosseus
1938; Herzog 1954, 1960; Seki 1974; Matteri 1985b (as Sematophyllum callidum, Mitten
1869; Brotherus 1924a; Bartram 1952; as Rhaphidostegium callidum, Espinosa 1941; Dusén
1903b; Cardot 1908; Thériot 1917, 1918; Herzog 1922; Cardot & Brotherus 1923; Herzog &
Schwabe 1939). Dist. IV. Limarí; V. Petorca, Valparaíso; VI. Cardenal Caro; VII. Cauquenes,
X. Chiloé, Llanquihue, Osorno; XI. Aisén, Coihaique; XII. Magallanes, Última Esperanza.
Rhaphidorrhynchium dendroligotrichum (Dusén) Broth.
Herzog 1954, 1960; Seki 1974 (as Rhaphidostegium dendroligotrichum, Dusén 1905a; Cardot
1908; as Rhaphidostegium polytrichadelphus, Dusén 1903b). Dist. XI. Aisén.
Rhaphidorrhynchium leptophyllum (Mitt.) Broth.
Tropicos 2009; He 1998; Brotherus 1925 (as Sematophyllum leptophyllum, Mitten 1869). Dist.
V. Petorca, San Antonio; VIII. Concepción; IX. Malleco.
Rhaphidorrhynchium liliputianum (Mont.) Broth.

80 Müller
(as Sematophyllum liliputianum, Mitten 1869). Dist. Chile, without exact locality.
Rhaphidorrhynchium noduliferum (Mitt.) Broth.
Matteri & Schiavone 2002; Brotherus 1925 (as Sematophyllum noduliferum, Mitten 1869; as
Rhaphidostegium noduliferum, Dusén 1903b; Cardot 1908). Dist. XII. Antártica Chilena.
Rhaphidorrhynchium scorpiurus (Mont.) Broth.
Hässel et al. 1984; Larraín 2005; He 1998; Matteri & Schiavone 2002; Herzog & Hosseus
1938; Herzog 1954, 1960; Seki 1974; Matteri 1985b (as Sematophyllum scorpiurus, Ireland et
al. 2007; as Rhaphidostegium scorpiurus, Mitten 1869; Herzog 1922; as Sematophyllum
aureonitidum, Bartram 1946; as Warburgiella aureonitida, Akiyama 1993). Dist. VIII. Arauco;
XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno; XI. Aisén; XII. Magallanes, Tierra del
Rhaphidorrhynchium tenuirostre (Hook.f.) Broth. = Sematophyllum uncinatum fide Stone & Scott 1973.
Rhaphidostegium aberrans Broth. ≡ Sematophyllum aberrans.
Rhaphidostegium amoenum (Hedw.) A. Jaeger ≡ Rhaphidorrhynchium amoenum.
Rhaphidostegium berberidis (Dusén) Broth. ≡ Rhaphidorrhynchium berberidis.
Rhaphidostegium brachycladulum Broth. ≡ Sematophyllum brachycladulum.
Rhaphidostegium caespitosum (Sw.) A. Jaeger ≡ Sematophyllum caespitosum.
Rhaphidostegium caespitosoides Broth. = Sematophyllum aberrans fide Robinson 1975.
Rhaphidostegium callidum (Mont.) A. Jaeger ≡ Rhaphidorrhynchium callidium.
Rhaphidostegium dendroligotrichum Dusén ≡ Rhaphidorrhynchium dendroligotrichum.
Rhaphidostegium leucocytus (Müll. Hal.) A. Jaeger ≡ Warburgiella leucocytus.
Rhaphidostegium masafuerae Broth. ≡ Sematophyllum masafuerae.
Rhaphidostegium noduliferum (Mitt.) A. Jaeger ≡ Rhaphidorrhynchium noduliferum.
Rhaphidostegium pallens Dusén, nom. nud., Hypnum pallens fide Cardot 1908, hom. illeg.
Rhaphidostegium patagonicum Dusén, nom. nud., Dusén 1903b.
Rhaphidostegium patagonicum Broth., nom. nud., Cardot & Brotherus 1923.
Rhaphidostegium polytrichadelphus Dusén, nom. nud. = Rhaphidorrhynchium dendroligotrichum fide Brotherus 1925.
Rhaphidostegium scorpiurus (Mont.) A. Jaeger ≡ Rhaphidorrhynchium scorpiurus.
Rhaphidostegium secundifolium (Müll. Hal.) A. Jaeger ≡ Drepanocladus secundifolius.
Rhaphidostegium strictipes Dusén, nom. nud., Dusén 1903b.
Rhaphidostegium tristifolium Dusén, nom. nud., Dusén 1903b.
Rhaphidostichum gunckeli Thér. ≡ Sematophyllum gunckelii.
Rhexophyllum subnigrum (Mitt.) Hilp.
Rhizogonium lindigii Hampe = Rhizogonium novae-hollandiae fide Churchill & Linares 1995b.
Rhizogonium longicolle Müll. Hal. = Pyrrhobryum mnioides fide Matteri & Piñero 1987.
Rhizogonium mnioides (Hook.f.) Wilson ≡ Pyrrhobryum mnioides.
Rhizogonium novae-hollandiae (Brid.) Brid.
Tropicos 2009; Matteri & Piñero 1987; Espinosa 1941; He 1998; Matteri & Schiavone 2002;
Robinson 1975; Deguchi 1991b (as Rhizogonium lindigii, Herzog 1960; as Rhizogonium
novae-hollandiae var. patagonicum, Cardot & Brotherus 1923; Brotherus 1924b). Dist. XIV.
Valdivia/Ranco; X. Chiloé, Llanquihue; XI. Aisén; XII. Magallanes, Tierra del Fuego; Juan
Fernández Is.
Rhizogonium novae-hollandiae var. patagonicum Cardot & Broth. = Rhizogonium novae-hollandiae fide Matteri &
Piñero 1987.
Rhizogonium reticulatum (Hook.f. & Wilson) Mitt. = Philonotis vagans fide Greene 1986.
Rhizogonium subbasilare (Hook.f.) Schimp. ≡ Goniobryum subbasilare.
Rhodobryum chilensis Thér. = Bryum rubromarginatum fide Ochi 1982.
Rhynchostegiella acanthophylla (Mont.) Broth. ≡ Eurhynchiella acanthophylla.
Rhynchostegiella acanthophylla var. robusta Thér. ≡ Eurhynchiella acanthophylla var. robusta.
Rhynchostegiella chilensis Thér.
He 1998; Thériot 1934b; Mahu 1981. Dist. V. Valparaíso; RM. Santiago.
Rhynchostegium byssicladum Dusén, nom. nud., Dusén 1903b.
Rhynchostegium complanum (Mitt.) A. Jaeger
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Mosses of Chile 81
Larraín 2007b; Espinosa 1941; He 1998; Brotherus 1924b; Herzog 1954, 1960; Bartram 1957;
Robinson 1975 (as Hypnum complanum, Mitten 1869). Dist. IV. Limarí; RM. Santiago; VIII.
Concepción; XIV. Valdivia/Ranco; X. Chiloé; Juan Fernández Is.
Rhynchostegium mollissimum Dusén, nom. nud., Dusén 1903b.
Rhynchostegium peruviense (R.S. Williams) Ochyra
(as Hygrohypnum peruviense, Bartram 1943). Dist. III. Huasco.
Rhynchostegium subsquarrosum Herzog = Eurhynchium corralense fide Wijk et al. 1967.
Rhynchostegium tenuifolium (Hedw.) Reichardt
He 1998; Brotherus 1924b; Herzog 1954; Robinson 1975 (as Hypnum tenuifolium, Mitten
1869). Dist. RM. Santiago; XIV. Valdivia/Ranco; X. Osorno; Juan Fernández Is.
Richardsiopsis lacustris (Herzog & Rich.) Ochyra = Drepanocladus perplicatus fide Hedenäs 2003.
Rigodium adpressum Zomlefer
Tropicos 2009; Buck 2002; He 1998; Zomlefer 1993; Villagran et al. 2004. Dist. IV. Choapa,
Limarí; VIII. Arauco, Biobío, Concepción, Ñuble; IX. Cautín, Malleco; XIV. Valdivia/Ranco;
X. Chiloé, Osorno; XI. Aisén; XII. Antártica Chilena, Magallanes.
Rigodium arborescens (Müll. Hal.) Kindb. = Rigodium brachypodium fide Zomlefer 1993.
Rigodium argentinicum (Müll. Hal.) Kindb. = Rigodium toxarion fide Zomlefer 1993.
Rigodium brachypodium (Müll. Hal.) Paris
Tropicos 2009; Larraín 2007b; Pereira et al. 2006; Buck 2002; Larraín 2005; Villagran et al.
2003, 2004; He 1998; Bartram 1952; Zomlefer 1993 (as Rigodium arborescens, Hässel et al.
1984; Espinosa 1941; Thériot 1917, 1918; Brotherus 1924b; Roivainen 1934; Herzog 1954,
1960; Bartram 1957; Robinson 1975; Matteri 1985b; as Rigodium lechleri, Herzog 1922). Dist.
IV. Choapa, Limarí; V. Petorca, Valparaíso; RM. Santiago; VI. Cardenal Caro; VII. Curicó,
Rigodium carnosulum Dusén, nom. nud., Dusén 1903b.
Rigodium elegantulum Cardot ex Thér., nom. nud. = Rigodium tamarix fide Zomlefer 1993.
Rigodium hylocomioides Cardot & Broth. = Rigodium pseudothuidium fide Zomlefer 1993.
Rigodium hylocomioides var. gracilius Cardot & Broth. = Rigodium pseudothuidium fide Zomlefer 1993.
Rigodium implexum Kunze ex Schwägr.
Larraín 2005; Villagran et al. 2003; Schaumann 2005; He 1998; Herzog 1922, 1954, 1960;
Cardot & Brotherus 1923; Roivainen 1934; Herzog & Hosseus 1938; Herzog & Schwabe
1939; Bartram 1952; Deguchi 1991b; Zomlefer 1993; Frahm 2001b. Dist. VIII. Concepción;
IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno; XI. Aisén.
Rigodium lechleri Schimp. = Rigodium brachypodium fide Zomlefer 1993.
Rigodium looseri Thér. = Rigodium toxarion var. robustum fide Zomlefer 1993.
Rigodium nano-fasciculatum Müll. Hal. ex Thér. = Rigodium toxarion fide Robinson 1975; Zomlefer 1993.
Rigodium pendulum Herzog & Thér. = Rigodium toxarion fide Zomlefer 1993.
Rigodium pseudothuidium Dusén
Larraín 2007b; Pereira et al. 2006; Larraín 2005; Villagran et al. 2003; He 1998; Matteri &
Schiavone 2002; Dusén 1905a; Cardot 1908; Herzog 1954, 1960; Seki 1974; Deguchi 1991b;
Zomlefer 1993 (as Rigodium hylocomioides, Barrera 2006; Cardot & Brotherus 1923;
Brotherus 1924b; Thériot 1926; Herzog & Schwabe 1939; Herzog 1960; Robinson 1975; as
Rigodium hylocomioides var. gracilius, Cardot & Brotherus 1923). Dist. IV. Choapa; V.
Petorca; VII. Curicó, Talca; VIII. Concepción, Ñuble; IX. Cautín, Malleco; XIV.
Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno; XI. Aisén, Coihaique; XII. Magallanes, Tierra
del Fuego; Juan Fernández Is.
Rigodium robustum Broth. ≡ Rigodium toxarion var. robustum.
Rigodium tamarix Müll. Hal.
Tropicos 2009; Larraín 2007b; Schaumann 2005; He 1998; Dusén 1903b; Cardot 1908; Cardot
& Brotherus 1923; Roivainen 1934; Thériot 1934b; Seki 1974; Zomlefer 1993 (as Rigodium
elegantulum, Thériot 1934b). Dist. IV. Choapa, Limarí; V. Petorca, San Antonio; VI.

82 Müller
Colchagua; VIII. Arauco, Biobío, Concepción, Ñuble; IX. Cautín, Malleco; XIV.
Rigodium toxarion (Schwägr.) A. Jaeger
Tropicos 2009; Larraín 2007b; Pereira et al. 2006; Larraín 2005; Espinosa 1941; Muñoz &
Pisano 1947; Villagran et al. 2003; Schaumann 2005; He 1998; Müller & Pereira 2006; Dusén
1903b; Cardot 1908; Brotherus 1924a, 1924b; Roivainen 1934; Herzog & Schwabe 1939;
Herzog 1954, 1960; Bartram 1957; Robinson 1975; Deguchi 1991b; Zomlefer 1993 (as
Rigodium argentinicum, Seki 1974; as Rigodium nano-fasciculatum, Dusén 1903b; Thériot
1918; Seki 1974; as Rigodium pendulum, Herzog & Schwabe 1939; Herzog 1954; Seki 1974;
as Hypnum toxarion, Mitten 1869). Dist. IV. Choapa, Limarí; V. Petorca, Valparaíso; VI.
Cardenal Caro; VII. Cauquenes, Talca; VIII. Arauco, Biobío, Concepción, Ñuble; IX. Cautín,
Malleco; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno, Palena; XI. Aisén; Juan
Fernández Is.
Rigodium toxarion var. robustum (Broth.) Zomlefer
Zomlefer 1993 (as Rigodium looseri, Barrera 2006; Thériot 1927a, 1928; as Rigodium
robustum, Brotherus 1924b; Bartram 1957; Robinson 1975). Dist. Juan Fernández Is.
Sagenotortula quitoensis (Taylor) R.H. Zander
Zander 1989 (as Tortula mniadelphus, Thériot 1923; as Tortula quitoensis, Mishler in Sharp et
al. 1994). Dist. RM. Santiago.
Saitobryum lorentzii (Müll. Hal.) Ochyra
Sanionia symmetrica (Renauld & Cardot) Wheldon
Ireland et al. 2007 (as Hypnum symmetricus, Cardot 1908; as Hypnum uncinatum var.
symmetricus, Dusén 1903b; as Amblystegium uncinatum var. symmetricus, Dusén 1903b; as
Drepanocladus symmetricus, Cardot & Brotherus 1923). Dist. VIII. Ñuble; XI. Aisén.
Sanionia uncinata (Hedw.) Loeske
Goffinet et al. 2006; Larraín 2007b; Buck 2002; Larraín 2007a; Ireland et al. 2007; Schaumann
2005; He 1998; Ochyra & Matteri 2001; Matteri & Schiavone 2002; Seki 1974; Deguchi 1991a
(as Drepanocladus uncinatus, Hässel et al. 1984; Cardot & Brotherus 1923; Roivainen 1934;
Herzog & Schwabe 1939; Herzog 1954; Matteri 1985b; as Hypnum austrouncinatum, Dusén
1903b; as Hypnum uncinatum, Dusén 1903b; as Drepanocladus uncinatus var. stenocarpus,
Barrera 2006). Dist. VIII. Biobío, Ñuble; IX. Cautín, Malleco; X. Chiloé, Llanquihue, Osorno;
XI. Aisén, Coihaique; XII. Antártica Chilena, Magallanes, Tierra del Fuego, Última Esperanza.
Sarconeurum antarcticum Bryhn = Syntrichia sarconeurum fide Ochyra et al. 2008.
Sarconeurum glaciale (Müll. Hal.) Cardot & Bryhn = Syntrichia sarconeurum fide Ochyra & Zander 2007.
Sarmentypnum sarmentosum (Wahlenb.) Tuom. & T.J. Kop. ≡ Warnstorfia samentosa.
Sauloma tenella (Hook.f. & Wilson) Mitt.
Müller 2009; Hässel et al. 1984; Buck 2002; Buck 2004; Matteri & Schiavone 2002; Bartram
1946; Matteri 1972, 1985b; Deguchi 1991a (as Pulvinella albicans, Herzog 1928). Dist. X.
Llanquihue; XI. Coihaique; XII. Antártica Chilena, Magallanes, Tierra del Fuego, Última
Esperanza.
Schimperobryum splendidissimum (Mont.) Margad. var. splendidissimum
Tropicos 2009; Larraín 2007b; Larraín 2005; Villagran et al. 2003; He 1998; Müller & Pereira
2006; Matteri 1972 (as Lepidopilum splendidissimum, Mitten 1869; as Cyathophorum
splendidissimum, Dusén 1903b; Cardot 1908; as Lamprophyllum splendidissimum, Espinosa
1941; Herzog 1922, 1954; Cardot & Brotherus 1923; Thériot 1924, 1927a; Brotherus 1924b;
Reimers 1926; Herzog & Hosseus 1938; Bartram 1957; Robinson 1975). Dist. V. Valparaíso;
RM. Chacabuco, Santiago; VII. Cauquenes, Talca; VIII. Arauco, Biobío, Concepción, Ñuble;
IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno; XI. Aisén,
Coihaique; Juan Fernández Is.
Schimperobryum splendidissimum (Mont.) Margad. var. perdentatum Matteri
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Mosses of Chile 83
Villagran et al. 2003; Matteri 1972; Deguchi 1991a. Dist. VIII. Concepción; IX. Cautín; XIV.
Schistidium absconditum (Cardot) Ochyra = Schistidium praemorsum fide Ochyra et al. 2008.
Schistidium amblyophyllum (Müll. Hal.) Ochyra & Hertel
Ochyra et al. 2008; Matteri & Schiavone 2002; Herzog 1960; Ochyra & Hertel 1990 (as
Grimmia amblyophylla, Mitten 1869; Dusén 1903b, 1906b; Cardot 1908; Bartram 1952; as
Grimmia hyalinocuspidata, Cardot & Brotherus 1923; as Schistidium donatii, Ochyra &
Matteri 1996; as Grimmia donatii, Herzog 1937). Dist. XIV. Valdivia/Ranco; XI. Capitán Prat;
XII. Antártica Chilena, Magallanes.
Schistidium andinum (Mitt.) Herzog
Ochyra et al. 2008 (as Schistidium syntrichiaceum, Hässel et al. 1984; Matteri 1985b; Matteri
& Schiavone 2002; as Schistidium occultum, Buck 2002; Ochyra 1998; Matteri & Schiavone
2002; as Grimmia occulta, Dusén 1906b; Cardot & Brotherus 1923; as Schistidium fallax,
Matteri & Schiavone 2002; Ochyra & Matteri 1996; as Grimmia fallax, Dusén 1903b, 1906b;
Cardot 1908; Thériot 1924). Dist. XIV. Valdivia/Ranco; XII. Antártica Chilena, Magallanes,
Schistidium apocarpum (Hedw.) Bruch & Schimp.
Tropicos 2009; Hässel et al. 1984; He 1998; Deguchi 1984; Matteri 1985b (as Grimmia
apocarpa, Mitten 1869; Dusén 1903b; Cardot 1908; Roivainen 1934; as Grimmia perplexa,
Thériot 1923, 1934b). Dist. V. San Antonio, Valparaíso; RM. Santiago; VI. Cardenal Caro;
VII. Linares; VIII. Biobío, Ñuble; IX. Cautín; XII. Antártica Chilena, Magallanes, Última
Esperanza.
Schistidium celatum (Cardot) B.G. Bell. = Schistidium cupulare fide Ochyra 2004b.
Schistidium cupulare (Müll. Hal.) Ochyra
Ochyra 2004b (as Schistidium celatum, Buck 2002). Dist. XII. Antártica Chilena.
Schistidium donatii (Herzog & Thér.) Ochyra & Matteri = Schistidium amblyophyllum fide Ochyra et al. 2008.
Schistidium falcatum (Hook.f. & Wilson) B. Bremer
Ochyra et al. 2008; Matcham et al. 2007; Buck 2002; Ireland et al. 2007; He 1998; Müller &
Pereira 2006; Matteri & Schiavone 2002; Deguchi 1984, 1991a (as Grimmia fasciculata,
Dusén 1905a, 1906b). Dist. VII. Cauquenes; VIII. Biobío, Ñuble; X. Osorno; XII. Antártica
Schistidium fallax (Dusén) Ochyra & Matteri = Schistidium andinum fide Ochyra et al. 2008.
Schistidium halinae Ochyra
NY 2009. Dist. XII. Antártica Chilena (Buck 40920 & 41249, NY).
Schistidium hyalinocuspidatum (Müll. Hal.) B.G. Bell = Schistidium amblyophyllum fide Ochyra 1998.
Schistidium occultum (Müll. Hal.) Ochyra & Matteri = Schistidium andinum fide Ochyra et al. 2008.
Schistidium perplexum (Thér.) Ochyra
Muñoz & Pando 2000. Dist. RM. Santiago.
Schistidium praemorsum (Müll. Hal.) Herzog
Ochyra et al. in Blockeel et al. 2009 (as Schistidium absconditum, Buck 2002). Dist. XII.
Antártica Chilena.
Schistidium rivulare (Brid.) Podp.
Tropicos 2009; Hässel et al. 1984; Buck 2002; Matteri & Schiavone 2002; Matteri 1985b (as
Grimmia alpicola, Cardot & Brotherus 1923; as Grimmia alpicola var. rivularis, Roivainen
1934). Dist. VIII. Biobío, Concepción, Ñuble; XII. Antártica Chilena, Magallanes, Tierra del
Schistidium scabripes (E.B. Bartram) Deguchi
Deguchi 1984, Deguchi 1991a (as Grimmia scabripes, Roivainen 1934; Bartram 1946). Dist.
VIII. Ñuble; IX. Cautín; XII. Tierra del Fuego.
Schistidium syntrichiaceum (Müll. Hal.) B. G. Bell = Schistidium andinum fide Ochyra et al. 2008.
Schizomitrium pallidum (Hornsch.) H.A. Crum & L.E. Anderson ≡ Callicostella pallida.
Schizymenium brevicaule (Hornsch.) A.J. Shaw & S.P. Churchill

84 Müller
(as Mielichhoferia brevicaulis, Mitten 1869). Dist. Chile, without exact locality (leg. C. Gay).
Schizymenium campylocarpum (Arn. & Hook.) A.J. Shaw
Matteri & Schiavone 2002 (as Mielichhoferia campylocarpa, Hässel et al. 1984; Matteri
1985b; as Mielichhoferia spegazzinii, Brotherus 1924c). Dist. XII. Magallanes, Última
Esperanza.
Schizymenium fulvonitens (Cardot & Broth.) A.J. Shaw
Matteri & Schiavone 2002 (as Mielichhoferia fulvonitens, Cardot & Brotherus 1923). Dist.
XII. Tierra del Fuego.
Schizymenium multiflorum (E.B. Bartram) A.J. Shaw
(as Mielichhoferia multiflora, Roivainen 1934; Robinson 1975). Dist. VIII. Ñuble; XII. Tierra
del Fuego.
Schizymenium pleurogynum (Mont.) Matteri
Matteri 1998 (as Mielichhoferia pleurogyna, Mitten 1869; Thériot 1934b; Bartram 1952). Dist.
RM. Santiago.
Schizymenium skottsbergii (Cardot & Broth.) A.J. Shaw
Matteri & Schiavone 2002 (as Mielichhoferia skottsbergii, Cardot & Brotherus 1923). Dist.
XII. Magallanes.
Sciaromium confluens (Müll. Hal.) Paris = Vittia pachyloma fide Ochyra 1987b.
Sciaromium conspissatum (Hook.f. & Wilson) Mitt. = Vittia pachyloma fide Ochyra 1987b.
Sciaromium conspissatum var. maritimum (Cardot) Herzog = Vittia pachyloma fide Ochyra 1987b.
Sciaromium crassinervatum Mitt. = Vittia pachyloma fide Ochyra 1987b.
Sciaromium depastum Dusén = Vittia pachyloma fide Ochyra 1987b.
Sciaromium flavidulum Dusén = Eurhynchium fuegianum fide Matteri & Ochyra 1989.
Sciaromium gracile Dusén = Cratoneuropsis relaxa subsp. minor fide Ochyra et al. 2008.
Sciaromium krauseanum Paris = Vittia pachyloma fide Ochyra 1987b.
Sciaromium lacustre Herzog & P.W. Richards = Drepanocladus perplicatus fide Hedenäs 2003.
Sciaromium maritimum Cardot = Vittia pachyloma fide Ochyra 1987b.
Sciaromium nigritum Dusén = Vittia pachyloma fide Ochyra 1987b.
Sciaromium obscurifolium Mitt. = Vittia pachyloma fide Ochyra 1987b.
Sciaromium pachyloma (Mont.) Paris ≡ Vittia pachyloma.
Sciaromium pachyloma var. brevifolia Thér. = Vittia pachyloma fide Ochyra 1987b.
Sciaromium pachyloma var. gracilescens Herzog = Vittia pachyloma fide Ochyra 1987b.
Sciuro-hypnum filirepens (Dusén) Ochyra & śarnowiec
(as Brachythecium filirepens, Buck 2002; Dusén 1905a; Cardot 1908; McFarland 1988;
Matteri & Schiavone 2002; as Brachythecium longidecurrens, Bartram 1946). Dist. XIV.
Valdivia/Ranco; XI. Aisén; XII. Antártica Chilena, Magallanes, Tierra del Fuego.
Sciuro-hypnum fuegianum (Broth.) Ochyra & śarnowiec
(as Brachythecium majusculum, Schaumann 2005; Frey & Schaumann 2002; Newton 1974;
McFarland 1988; Matteri & Schiavone 2002; Dusén 1903b). Dist. XII. Antártica Chilena,
Sciuro-hypnum glaciale (Schimp.) Ignatov & Huttunen
Ochyra et al. 2008. Dist. XI. Aisén; XII. Antártica Chilena, Tierra del Fuego.
Sciuro-hypnum plumosum (Hedw.) Ignatov & Huttunen
(as Brachythecium plumosum, McFarland 1988; as Brachythecium flagellare, Roivainen 1934;
as Brachythecium subpilosum, Herzog 1922). Dist. VIII. Ñuble; X. Osorno; XII. Magallanes.
Scleropodium purum (Hedw.) Limpr. ≡ Pseudoscleropodium purum.
Scorpidium cossonii (Schimp.) Hedenäs
(as Limprichtia cossonii, Ochyra & Matteri 2001; Matteri & Schiavone 2002). Dist. XII. Tierra
Scorpidium revolvens (Sw. ex anon.) Rubers
(as Limprichtia revolvens, Ochyra & Matteri 2001; Matteri & Schiavone 2002). Dist. XII.
Scouleria patagonica (Mitt.) A. Jaeger
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Mosses of Chile 85
Tropicos 2009; He 1998; Dusén 1903b; Dusén 1906b; Cardot 1908; Thériot 1935; Herzog
1954, 1960; Seki 1974; Matteri 1979; Deguchi 1991a. Dist. VIII. Biobío, Ñuble; IX. Malleco;
XIV. Valdivia; X. Chiloé, Llanquihue, Osorno, Palena; XI. Aisén, Coihaique.
Sematophyllum aberrans (Broth.) E.B. Bartram
Ireland & Bellolio 2002; Bartram 1957; Robinson 1975 (as Rhaphidostegium aberrans,
Barrera 2006; Espinosa 1941; Brotherus 1924b; as Rhaphidostegium caespitosoides, Barrera
2006; Espinosa 1941; Brotherus 1924b). Dist. Juan Fernández Is.; Easter I.
Sematophyllum amoenum (Hedw.) Mitt. ≡ Rhaphidorrhynchium amoenum.
Sematophyllum aureo-nitidum E.B. Bartram = Rhaphidorrhynchium scorpiurus fide Seki 1974.
Sematophyllum brachycladulum (Broth.) Broth.
Ireland & Bellolio 2002; Bartram 1957; Robinson 1975 (as Rhaphidostegium brachycladulum,
Brotherus 1924b). Dist. Juan Fernández Is.; Easter I.
Sematophyllum caespitosum Mitt.
Robinson 1975 (as Rhaphidostegium caespitosum, Brotherus 1924b). Dist. Juan Fernández Is.
Sematophyllum callidum (Mont.) Mitt. ≡ Rhaphidorrhynchium callidum.
Sematophyllum gunckelii (Thér.) W.R. Buck
Buck 1989 (as Rhaphidostichum gunckelii, Thériot 1930). Dist. XIV. Valdivia/Ranco.
Sematophyllum kunkelii H. Rob.
Robinson 1975. Dist. Juan Fernández Is.
Sematophyllum leptophyllum Mitt. ≡ Rhaphidorrhynchium leptophyllum.
Sematophyllum leucocytum (Müll. Hal.) Sainsb. ≡ Warburgiella leucocytus.
Sematophyllum liliputianum (Mont.) Mitt. ≡ Rhaphidorrhynchium liliputianum.
Sematophyllum masafuerae (Broth.) E.B. Bartram
Bartram 1957; Robinson 1975 (as Rhaphidostegium masafuerae, Brotherus 1924b; as Hypnum
masafuerae, Bartram 1957). Dist. Juan Fernández Is.
Sematophyllum noduliferum Mitt. ≡ Rhaphidorrhynchium noduliferum.
Sematophyllum secundifolium (Müll. Hal.) Mitt. ≡ Drepanocladus secundifolius.
Sematophyllum swartzii (Schwägr.) W.H. Welch & H.A. Crum
He 1998. Dist. IX. Cautín.
Sematophyllum uncinatum I.G. Stone & G.A.M. Scott
Müller & Pereira 2006 (as Hypnum tenuirostre, Mitten 1869; as Rhaphidorrhynchium
tenuirostre, Brotherus 1925). Dist. VI. O'Higgins; VII. Cauquenes.
Skottsbergia paradoxa Cardot
Matteri & Schiavone 2002; Roivainen 1937. Dist. XII. Tierra del Fuego.
Sphagnum amblyphyllum (Russ.) Warnst. = Sphagnum flexuosum fide Smith 2004.
Sphagnum capillifolium (Ehrh.) Hedw.
Matcham et al. 2007; Larraín 2007a; Larraín 2005 (as Sphagnum nemoreum, Deguchi 1991b;
as Sphagnum acutifolium, Larraín 2005; Villagran et al. 2003). Dist. XIV. Valdivia/Ranco; X.
Chiloé, Llanquihue.
Sphagnum cuspidatum Ehrh. ex Hoffm.
Tropicos 2009; Larraín 2005; Villagran et al. 2003; Cardot 1908; Cardot & Brotherus 1923;
Thériot 1934b; Deguchi 1991b (as Sphagnum cuspidatum var. falcatum; Dusén 1903a, 1903b;
Cardot 1908; Sphagnum cuspidatum var. miquelonense (Renauld & Cardot) Renauld & Cardot,
Dusén 1903a; as Sphagnum cuspidatum var. serrulatum (Schlieph.) Schlieph., Matteri &
Schiavone 2002; as Sphagnum trinitense, Cardot & Brotherus 1923; as Sphagnum cuspidatum
var. serratum, Dusén 1903a; as Sphagnum cuspidatum var. submersum, Dusén 1903a). Dist.
XIV. Valdivia/Ranco; X. Chiloé; XI. Aisén; XII. Antártica Chilena, Magallanes.
Sphagnum cuspidatum var. falcatum Russow = Sphagnum cuspidatum fide Crum 1984.
Sphagnum cymbifolium var. condensatum Hook.f., hom. illeg., Dusén 1903b (XII. Magallanes).
Sphagnum dissimile Roiv.
Sphagnum dusenioides Roiv.
Matteri & Schiavone 2002; Roivainen 1937. Dist. XII. Magallanes, Tierra del Fuego.

86 Müller
Sphagnum ehyalinum A.J. Shaw & Goffinet

Goffinet et al. 2006; Shaw & Goffinet 2000. Dist. XII. Magallanes.
Sphagnum falcatulum Besch.
Díaz et al. 2008; Larraín 2007b; Hässel et al. 1984; Larraín 2005; Villagran et al. 2003; He
1998; Matteri & Schiavone 2002; Cardot & Brotherus 1923; Roivainen 1937; Seki 1974;
Matteri 1985b; Deguchi 1991b; Warnstorf 1911 (as Sphagnum undulatum, Dusén 1903a;
Warnstorf 1911; as Sphagnum patagoniense, Warnstorf 1911; Herzog & Schwabe 1939). Dist.
IX. Malleco; XIV. Valdivia/Ranco; XI. Chiloé, Llanquihue, Osorno; XI. Aisén; XII. Antártica
Sphagnum fimbriatum Wilson
Goffinet et al. 2006; Larraín 2007b; Hässel et al. 1984; Matcham et al. 2007; Buck 2002;
Larraín 2005; Ireland et al. 2007; Villagran et al. 2003; Shaw & Goffinet 2000; He 1998;
Matteri & Schiavone 2002; Mitten 1869; Cardot 1900, 1901, 1908; Dusén 1903b; Roivainen
1937; Herzog & Schwabe 1939; Seki 1974; Matteri 1985b; Deguchi 1991b (as Sphagnum
fimbriatum var. robustum, Warnstorf 1911; Dusén 1903a, 1903b; Cardot & Brotherus 1923;
Herzog & Schwabe 1939; as Sphagnum fimbriatum var. robustum fo. brachyasyclada, Dusén
1903a; Cardot 1908; as Sphagnum fimbricatum var. robustum fo. laxifolia, Dusén 1903a; as
Sphagnum fimbriatum var. validius, Cardot 1908; Cardot & Brotherus 1923). Dist. VIII.
Arauco; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno, Palena; XI. Aisén; XII.
Sphagnum flexuosum Dozy & Molk.
(as Sphagnum amblyphyllum, Thériot 1934b). Dist. VIII. Ñuble.
Sphagnum magellanicum Brid.
Díaz et al. 2008; Goffinet et al. 2006; Larraín 2007b; Hässel et al. 1984; Matcham et al. 2007;
Buck 2002; Larraín 2005; Ireland et al. 2007; Villagran et al. 2003; Shaw & Goffinet 2000; He
1998; Matteri & Schiavone 2002; Roivainen 1937; Herzog & Hosseus 1938; Herzog &
Schwabe 1939; Seki 1974; Matteri 1985b; Deguchi 1991b (as Sphagnum medium, Cardot
1908; Warnstorf 1911; as Sphagnum rigescens, Warnstorf 1911; Dusén 1903a, 1903b; as
Sphagnum medium var. flavescens, Warnstorf 1911; as Sphagnum medium var. fuscorubellum,
Dusén 1903a; Sphagnum medium var. fuscorubellum fo. brachydasyclada, Dusén 1903a,
1903b; as Sphagnum medium var. pallidocarneum fo. brachydasyclada, Dusén 1903a, 1903b;
Sphagnum medium var. purpurascens, Dusén 1903a; Sphagnum medium var. purpurascens fo.
brachylomatoclada, Dusén 1903a). Dist. VIII. Arauco; IX. Malleco; XIV. Valdivia/Ranco; X.
Chiloé, Llanquihue, Osorno, Palena; XI. Aisén; XII. Antártica Chilena, Magallanes, Tierra del
Sphagnum medium Limpr. = Sphagnum magellanicum fide Eddy 1977.
Sphagnum miradorense Larraín, nom. nud., Larraín 2005 (XIV. Valdivia/Ranco).
Sphagnum nemoreum Scop. = Sphagnum capillifolium fide Crum 1984.
Sphagnum patagoniense Warnst. = Sphagnum falcatulum fide Wijk et al. 1967.
Sphagnum personatum Roiv.
Matteri & Schiavone 2002; Roivainen 1937. Dist. XII. Magallanes, Tierra del Fuego.
Sphagnum plumulosum Röll = Sphagnum subnitens fide Crum 1984.
Sphagnum pulchricoma Müll. Hal. = Sphagnum recurvum fide Crum 1984.
Sphagnum recurvum P. Beauv.
Díaz et al. 2008; Larraín 2007b; Cardot 1908 (as Sphagnum pulchricoma, Mitten 1869; Herzog
& Schwabe 1939). Dist. X. Chiloé, Llanquihue; XII. Antártica Chilena, Magallanes.
Sphagnum recurvum var. brevifolium (Lindb.) Warnst.
Ireland et al. 2007. Dist. VIII. Arauco.
Sphagnum rigescens Warnst. = Sphagnum magellanicum fide Crum 1990.
Sphagnum schwabeanum H.K.G. Paul
Herzog & Schwabe 1939. Dist. X. Llanquihue.
Sphagnum subfalcatulum Roiv.
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Mosses of Chile 87
Matteri & Schiavone 2002; Roivainen 1937; Deguchi 1991b. Dist. IX. Malleco; XII. Tierra del
Fuego.
Sphagnum subnitens Russow & Warnst.
He 1998 (as S. plumulosum, Warnstorf 1911; Brotherus 1924; as Sphagnum subnitens var.
purpurascens, Dusén 1903a). Dist. XIV. Valdivia/Ranco; X. Chiloé, Osorno; XI. Aisén.
Sphagnum subrigidum Hampe & Lorentz
Warnstorf 1911; Brotherus 1924c; Crum 1990. Dist. Chile, without exact locality.
Sphagnum subsecundum Nees
Larraín 2007b; He 1998. Dist. X. Chiloé, Llanquihue.
Sphagnum subserratum Roiv.
Sphagnum torreyanum Sull.
Warnstorf 1911. Dist. XII. Tierra del Fuego.
Sphagnum trinitense Müll. Hal. = Sphagnum cuspidatum fide Griffin 1981.
Sphagnum undulatum Warnst. = Sphagnum falcatulum fide Matteri & Schiavone 2002.
Splachnobryum obtusum (Brid.) Müll. Hal.
Arts 2001a. Dist. Chile, without exact locality.
Splachnobryum valdiviae Müll. Hal.
Brotherus 1924c. Dist. Chile, without exact locality. Note: The generic position is uncertain;
the species was excluded from the genus but was’nt placed in another (Crosby et al. 1999).
Sporledera hampeana Müll. Hal. ≡ Bruchia hampeana.
Stableria tenella (Mitt.) Broth. = Orthodontium gracile fide Robinson 1975.
Stenomitrium pentastichum (Mont.) Broth. ≡ Zygodon pentastichus.
Stenomitrium robustulum Dusén ex Paris, nom. nud. = Zygodon bartramioides fide Calabrese 2006.
Stereodon lechleri (Müll. Hal.) Mitt. = Hypnum chrysogaster fide Ando 1972.
Stereodon nitidus (Hook.f. & Wilson) Mitt. ≡ Catagonium nitidum.
Stereodon campoanus Thér. ≡ Hypnum campoanum.
Stereophyllum fuegianum Besch. = Brachythecium longidens fide Wijk et al. 1967.
Stereophyllum seminerve (Kunze ex Schwägr.) Mitt. ≡ Juratzkaea seminervis subsp. seminervis.
Stokesiella praelonga (Hedw.) H. Rob. ≡ Kindbergia praelonga.
Straminergon stramineum (Dicks. ex Brid.) Hedenäs
(as Calliergon stramineum, Donat 1936). Dist. XI. Aisén.
Streptopogon mnioides (Schwägr.) Mitt. ≡ Calyptopogon mnioides.
Symblepharis fuegiana (Cardot) Ochyra & Matteri
Matteri & Schiavone 2002; Ochyra & Matteri 1996 (as Oncophorus fuegianus, Frahm et al.
1998; Cardot 1900, 1901, 1908; Dusén 1903b; Brotherus 1924b; Bartram 1957; Robinson
1975; Matteri 1985b). Dist. X. Llanquihue; XI. Aisén; XII. Magallanes, Tierra del Fuego; Juan
Fernández Is.
Symblepharis krausei (Lorentz) Ochyra & Matteri
Buck 2002; Ireland et al. 2007; Müller & Pereira 2006; Ochyra & Matteri 1996 (as
Anisothecium krausei, Herzog & Schwabe 1939; Herzog 1954). Dist. VII. Cauquenes; VIII.
Arauco; XIV. Valdivia/Ranco; X. Osorno; XI. Aisén; XII. Antártica Chilena.
Symblepharis luteovirens (E.B. Bartram) Ochyra & Matteri
Matteri & Schiavone 2002; Ochyra & Matteri 1996 (as Oncophorus luteo-virens, Ireland et al.
2007; Roivainen 1937; Frahm et al. 1998). Dist. VIII. Arauco; XII. Magallanes, Tierra del
Fuego.
Syntrichia anderssonii (Ångström) R.H. Zander = Syntrichia glacialis fide Müller 2009.
Syntrichia anderssonii (Ångström) R.H. Zander var. fagicola (Müll. Hal.) R.H. Zander = Syntrichia glacialis fide
Matteri 2003a.
Syntrichia antarctica (Hampe) R.H. Zander
Matcham et al. 2007; Matteri & Schiavone 2002. Dist. XII. Última Esperanza.
Syntrichia apiculatopilosa Dusén, nom. nud. = Pseudocrossidium crinitum fide Müller 2009.
Syntrichia bogotensis (Hampe) Mitt. ex R.H. Zander
Gallego & Cano 2007b. Dist. IV. Elqui.

88 Müller
Syntrichia brachyclada (Cardot) R.H. Zander

Matteri & Schiavone 2002 (as Tortula brachyclada, Hässel et al. 1984). Dist. XII. Magallanes,
Última Esperanza.
Syntrichia breviseta (Mont.) M.J. Cano & M.T. Gallego
(as Tortula atrata, Thériot 1921; as Tortula breviseta, Mitten 1869; Thériot 1923; Herzog &
Hosseus 1938). Dist. V. Valparaíso; RM. Santiago; VIII. Biobío.
Syntrichia campestris (Dusén) R.H. Zander
Matteri & Schiavone 2002; Zander 1993 (as Tortula campestris, Dusén 1906a; Thériot 1921;
Bartram 1952; Seki 1974). Dist. RM. Cordillera; XI. Aisén, Coihaique; XII. Magallanes.
Syntrichia costesii (Thér.) R.H. Zander
Müller 2009; Larraín 2007a; Schaumann 2005; Herzog 1954; Zander 1993; Frey & Schaumann
2002 (as Tortula costesii, Thériot 1921; Herzog & Hosseus 1938). Dist. V. Valparaíso; VII.
Cauquenes; IX. Cautín; X. Chiloé, Osorno; XII. Magallanes.
Syntrichia crispatula Müll. Hal. = Calyptopogon mnioides (Schwägr.) Broth. fide Wijk et al. 1959.
Syntrichia didymodontoides (Broth.) R.H. Zander = Syntrichia saxicola fide Ochyra et al. 2008.
Syntrichia epilosa (Broth. ex Dusén) R.H. Zander var. epilosa
Ireland et al. 2007; He 1998; Zander 1993 (as Tortula epilosa, Dusén 1906a; Thériot 1934b).
Dist. V. Valparaíso; RM. Santiago; VIII. Arauco, Concepción, Ñuble.
Syntrichia epilosa var. pilifera (Thér.) R.H. Zander
Zander 1993 (as Tortula epilosa var. pilifera, Thériot 1926). Dist. RM. Santiago; VI.
Colchagua.
Syntrichia filaris (Müll. Hal.) R.H. Zander
Ochyra et al. 2008; Matteri & Schiavone 2002. Dist. XII. Magallanes.
Syntrichia flagellaris (Schimp.) R.H. Zander
(as Barbula flagellaris, Dusén 1903b, 1906a; Thériot 1917; Herzog 1922; as Tortula
flagellaris, Espinosa 1941; Mitten 1869; Thériot 1921; Brotherus 1924a, 1924b; Robinson
1975; as Tortula contorta, Mitten 1869). Dist. V. Valparaíso; RM. Santiago; XIV.
Valdivia/Ranco; XI. Aisén; XII. Magallanes; Juan Fernández Is.
Syntrichia flavidopilosa Dusén, nom. nud. = Pseudocrossidium crinitum fide Müller 2009.
Syntrichia fontana (Müll. Hal.) R.H. Zander
Müller 2009 (as Tortula rivularis, Cardot & Brotherus 1923; Dusén 1906; Cardot 1908). Dist.
XI. Coihaique; XII. Magallanes, Tierra del Fuego, Última Esperanza.
Syntrichia fragilis (Taylor) Ochyra
Cano 2003 (as Tortula laevinervis, Dusén 1906b; Villagran et al. 2004). Dist. XV. Parinacota;
IV. Limarí; V. Valparaíso.
Syntrichia fuegiana Mitt. = Syntrichia magellanica fide Lightowlers 1985.
Syntrichia geheebiaeopsis (Müll. Hal.) R.H. Zander
Matcham et al. 2007; Buck 2002; Matteri & Schiavone 2002. Dist. XII. Antártica Chilena,
Magallanes.
Syntrichia glacialis (Kunze ex Müll. Hal.) R.H. Zander
Müller 2009 (as Tortula glacialis, Mitten 1869; as Syntrichia anderssonii, Tropicos 2009;
Matcham et al. 2007; Buck 2002; Schaumann 2005; Matteri & Schiavone 2002; Herzog 1954;
Zander 1993; as Barbula andersonii, Cardot 1900, 1901; Dusén 1903b; as Barbula
perrubiginosa, Dusén 1903b; as Tortula andersonii, Hässel et al. 1984; Dusén 1906a; Cardot
1908; Cardot & Brotherus 1923; Roivainen 1934; Bartram 1952; Seki 1974; Matteri 1985b;
Deguchi 1991b; Tortula perrubiginosa, Paris 1903-1906; as Tortula pseudolatifolia, Hässel et
al. 1984; Matteri & Schiavone 2002; Dusén 1906a; Cardot 1908; Seki 1974; Matteri 1985b).
Dist. VII. Cauquenes, Talca; VIII. Arauco, Biobío, Concepción, Ñuble; IX. Cautín, Malleco;
X. Llanquihue, Palena; XI. Aisén, Coihaique; XII. Antártica Chilena, Magallanes, Tierra del
Syntrichia gromschii (Thér.) R.H. Zander
(as Tortula gromschii, Thériot 1934b). Dist. XII. Última Esperanza.
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Syntrichia jaffueli (Thér.) R.H. Zander ≡ Tortula jaffueli.

Syntrichia laevipila Brid.
Matcham et al. 2007; Ireland et al. 2007. Dist. VIII. Arauco, Concepción; XII. Magallanes.
Syntrichia lingulaefolia (Cardot & Broth.) Herzog, hom. illeg. = Hennediella arenae fide Lightowlers 1985.
Syntrichia lithophila (Dusén) Ochyra & R.H. Zander
Ochyra & Zander 2007 (as Syntrichia pygmaea, Matcham et al. 2007; as Tortula lithophila,
Dusén 1906a; Seki 1974; Deguchi 1991b). Dist. XI. Aisén; XII. Magallanes, Tierra del Fuego.
Syntrichia magellanica (Mont.) R.H. Zander
Müller 2009; Ochyra et al. 2008; Matcham et al. 2007; Matteri & Schiavone 2002; Zander
1993 (as Barbula lechleri, Dusén 1903b; as Tortula lechleri, Herzog 1922; as Barbula pusilla,
Dusén 1903b; as Tortula pusilla, Bartram 1952; as Tortula brachypelma, Dusén 1905a; as
Tortula magellanica, Cardot 1908; as Tortula fuegiana, Dusén 1906a; Cardot 1908; Herzog
1922; as Tortula monoica, Hässel et al. 1984). Dist. XI. Coihaique; XII. Magallanes, Tierra del
Syntrichia marginatoserrata Dusén, nom. nud. = Syntrichia serrulata fide Müller 2009.
Syntrichia micro-runcinata Dusén, nom. nud. = Leptodontium longicaule var. microruncinatum fide Müller 2009.
Syntrichia mniadelphus (Müll. Hal.) Herzog var. cochlearifolia Herzog
Herzog 1954 (as Tortula mniadelphus var. cochlearifolia, Seki 1974). Dist. XI. Aisén. Note:
Syntrichia mniadelphus is a synonym of Sagenotortula quitoensis (Taylor) R.H. Zander, but
the status of this variety is uncertain and need verification.
Syntrichia muricata M.T.Gallego & Cano
Gallego & Cano 2007a. Dist. IV. Elqui, Limarí.
Syntrichia papillosa (Wilson) Jur. var. papillosa
NY 2009; Tropicos 2009; Müller 2009; Ireland et al. 2007; Matteri & Schiavone 2002; Matteri
1994; Herzog 1954 (as Tortula papillosa, Hässel et al. 1984; Dusén 1905a; Dusén 1906a;
Cardot 1908; Thériot 1921; Deguchi 1991b). Dist. V. Aconcagua; VII. Talca; VIII. Arauco,
Concepción; X. Llanquihue; XII. Antártica Chilena, Magallanes, Última Esperanza.
Syntrichia papillosa var. chilensis (Thér.) R.H. Zander
Matteri 1994 (as Tortula papillosa var. chilensis, Thériot 1921). Dist. V. Valparaíso.
Syntrichia princeps (De Not.) Mitt.
Cano & Gallego 2008; Matcham et al. 2007; Buck 2002; Matteri & Schiavone 2002 (as
Tortula princeps, Hässel et al. 1984; Mitten 1869; Seki 1974; as Tortula grossiretis, Cardot
1908; Thériot 1918; Cardot & Brotherus 1923; Bartram 1952; as Tortula chrysopila, Cardot
1908). Dist. V. Valparaíso; IX. Malleco; XI. Aisén; XII. Antártica Chilena, Magallanes,
Última Esperanza. Note: The species was excluded by Ochyra et al. (2008) from the Southern
Hemisphere. All the records from the area must therefore belong to other species.
Syntrichia prostrata (Mont.) R.H. Zander = Syntrichia serrulata fide Gallego et al. 2009.
Syntrichia pseudolatifolia (Cardot) Cano & M.T. Gallego = Syntrichia glacialis fide Müller 2009.
Syntrichia pseudorobusta (Dusén) R.H. Zander
Pereira et al. 2006; Herzog 1954; Zander 1993 (as Tortula pseudo-robusta, Dusén 1906a;
Thériot 1921; Herzog & Hosseus 1938; Herzog & Schwabe 1939). Dist. VII. Talca; VIII.
Arauco, Biobío, Concepción, Ñuble; XIV. Valdivia/Ranco; X. Llanquihue; XI. Aisén.
Syntrichia pygmaea (Dusén) R.H. Zander = Syntrichia lithophila fide Ochyra & Zander 2007.
Syntrichia robusta (Hook. & Grev.) R.H. Zander
Buck 2002; Schaumann 2005; Matteri & Schiavone 2002; Herzog 1954; Zander 1993 (as
Barbula robusta, Dusén 1903b; as Tortula robusta, Hässel et al. 1984; Mitten 1869; Paris
1903-1906; Dusén 1906a; Cardot & Brotherus 1923; Herzog & Schwabe 1939; Seki 1974;
Matteri 1985b; Deguchi 1991b; as Tortula robusta var. laxa, Roivainen 1934; Bartram 1946;
as Tortula robusta var. runcinata, Cardot & Brotherus 1923; as Tortula punctulata, Mitten
1869). Dist. IV. Limarí; VIII. Ñuble; X. Llanquihue, Osorno; XI. Aisén; XII. Antártica
Syntrichia ruralis (Hedw.) F. Weber & D. Mohr

90 Müller
Müller 2009; Cano & Gallego 2008; Schaumann 2005 (as Barbula ruralis, Cardot 1900, 1901;
as Tortula bullata, Herzog 1923). Dist. VII. Talca; XII. Magallanes, Última Esperanza.
Syntrichia saxicola (Cardot) R.H. Zander
Ochyra et al. 2008; Buck 2002; Matteri & Schiavone 2002; Zander 1993 (as Tortula saxicola,
Seki 1974). Dist. XI. Aisén; XII. Antártica Chilena, Magallanes.
Syntrichia scabrella (Dusén) R.H. Zander
Ireland et al. 2007; Zander 1993 (as Tortula scabrella, Dusén 1906b). Dist. VI. Colchagua;
VIII. Concepción, Ñuble.
Syntrichia scabrinervis (Müll. Hal.) R.H. Zander
Herzog 1954; Zander 1993 (as Tortula scabrinervis, Mitten 1869; Brotherus 1924b; Bartram
1957; Robinson 1975). Dist. II. Antofagasta; IV. Limarí; V. Valparaíso; Juan Fernández Is.
Syntrichia schnyderi (Müll. Hal.) R.H. Zander
Zander 1993 (as Tortula schnyderi, Dusén 1906a; Cardot 1908). Dist. XII. Magallanes.
Syntrichia serripungens (Lorentz & Müll. Hal.) R.H. Zander
Herzog 1954; Zander 1993. Dist. V. Aconcagua.
Syntrichia serrulata (Hook. & Grev.) M.J. Cano
(as Hennediella serrulata, Matteri & Schiavone 2002; Zander 1993; as Tortula serrulata,
Mitten 1869; Bartram 1952; as Syntrichia prostrata, Tropicos 2009; Zander 1993; as Tortula
prostrata, Mitten 1869; Dusén 1906a; Thériot 1915, 1928; Herzog & Hosseus 1938; Bartram
1952; Herzog 1960). Dist. VII. Cauquenes, Linares; VIII. Biobío, Ñuble; IX. Cautín; XIV.
Valdivia/Ranco; X. Osorno; XII. Magallanes, Tierra del Fuego.
Syntrichia socialis (Dusén) R.H. Zander
Zander 1993 (as Tortula socialis, Dusén 1906b). Dist. X. Llanquihue; XII. Magallanes.
Syntrichia squarripila (Thér.) Herzog
Herzog 1954, 1960 (as Tortula squarripila, Thériot 1917; Bartram 1952). Dist. V. Aconcagua,
Valparaíso.
Syntrichia subpapillosa (Cardot & Broth.) Matteri
Matcham et al. 2007; Buck 2002; Buck 2005a; Matteri & Schiavone 2002; Matteri 1994 (as
Tortula subpapillosa, Cardot & Brotherus 1923). Dist. XII. Antártica Chilena, Magallanes.
Syrrhopodon fasciculatus Hook.f. & Grev. ≡ Mitthyridium fasciculatum.
Tayloria dubyi Broth. var. dubyi
Goffinet et al. 2006; Hässel et al. 1984; Matcham et al. 2007; Buck 2002; Buck 2005a; Matteri
& Schiavone 2002; Cardot 1908; Bartram 1952; Seki 1974; Matteri 1985b; Deguchi 1991b (as
Hymenocleiston magellanicum, Dusén 1903b). Dist. XI. Aisén; XII. Antártica Chilena,
Tayloria dubyi var. edenensis (Besch.) Paris
Cardot 1908 (as Hymenocleiston magellanicum var. edenense, Dusén 1903b). Dist. XII.
Magallanes.
Tayloria magellanica (Brid.) Mitt. var. magellanica
Goffinet et al. 2006; Hässel et al. 1984; Matcham et al. 2007; Buck 2002; Buck 2005a; Matteri
1993; Matteri & Schiavone 2002; Mitten 1869; Cardot 1908; Cardot & Brotherus 1923;
Matteri 1985b (as Dissodon magellanicum, Cardot 1900; Dusén 1903b). Dist. XII. Antártica
Tayloria magellanica var. gigantea E.B. Bartram
Matteri & Schiavone 2002; Bartram 1946. Dist. XII. Tierra del Fuego.
Tayloria mirabilis (Cardot) Broth.
Goffinet et al. 2006; Hässel et al. 1984; Matcham et al. 2007; Buck 2002; Buck 2005b;
Schaumann 2005; He 1998; Matteri & Schiavone 2002; Cardot 1908; Cardot & Brotherus
1923; Matteri 1985b (as Dissodon mirabilis, Cardot 1900). Dist. IX. Cautín, Malleco; XII.
Tayloria stenophysata (Herzog) A.K. Kop.
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Larraín 2007b; Hässel et al. 1984; He 1998; A. Koponen 1977; Matteri 1985b (as Tetraplodon
stenophysatus, Herzog & Schwabe 1939; Deguchi 1991b). Dist. XIV. Valdivia/Ranco; X.
Chiloé, Llanquihue; XII. Magallanes, Última Esperanza.
Tetraplodon fuegianus Besch. = Tetraplodon mnioides fide A. Koponen 1982.
Tetraplodon mnioides (Hedw.) Bruch & Schimp.
Díaz et al. 2008; Larraín 2007a; Robinson 1964; Deguchi 1991b (as Tetraplodon fuegianus,
Buck 2002, 2005a; Goffinet et al. 2006; Hässel et al. 1984; Matcham et al. 2007; Dusén 1903b;
Cardot 1908; Seki 1974; Matteri 1985b; Matteri 1993; Matteri & Schiavone 2002). Dist. XIV.
Tetraplodon stenophysatus Herzog ≡ Tayloria stenophysata.
Tetrapterum recurvirostrum (Müll. Hal.) Broth. = Weissia diffidentia fide Zander 1993.
Tetrodontium brownianum (Dicks.) Schwägr.
He 1998; Mahu 1985. Dist. IX. Cautín.
Thamniopsis incurva (Hornsch.) W.R. Buck
(as Hookeriopsis incurva, Matteri 1972). Dist. VIII. Concepción.
Thamnium arbusculans (Müll. Hal.) A. Jaeger ≡ Porothamnium arbusculans.
Thamnium assimile Broth. = Thamnobryum rigidum fide Robinson 1975.
Thamnium carolii Broth. ≡ Thamnobryum caroli.
Thamnium confertum Mitt. ≡ Thamnobryum confertum.
Thamnium corralense Broth., nom. herb. = Porothamnium arbusculans fide Gunckel 1938.
Thamnium crassinervium Broth. = Thamnobryum rigidum fide Robinson 1975.
Thamnium decumbens Besch. = Porothamnium arbusculans fide Reimers 1926.
Thamnium ingae Broth. ≡ Thamnobryum ingae.
Thamnium latinerve (Mitt.) Broth. = Thamnobryum rigidum fide Robinson 1975.
Thamnium leucocaulon (Müll. Hal.) Kindb. ≡ Porothamnium leucocaulon.
Thamnium neckeroides (Hook.f.) Schimp. ≡ Thamnobryum neckeroides.
Thamnium panduraefolium (Müll. Hal.) Broth. ≡ Porothamnium panduraefolium.
Thamnium proboscideum Broth. ≡ Thamnobryum proboscideum.
Thamnium rigidum Mitt. ≡ Thamnobryum rigidum.
Thamnium valdiviae (Müll. Hal.) Kindb. ≡ Porothamnium valdiviae.
Thamnobryum assimile (Broth.) Sérgio = Thamnobryum rigidum fide Robinson 1975.
Thamnobryum caroli (Broth.) H. Rob.
Robinson 1975 (as Thamnium carolii, Barrera 2006; Espinosa 1941; Brotherus 1924b; Bartram
1957). Dist. Juan Fernández Is.
Thamnobryum confertum (Mitt.) H. Rob.
Robinson 1975 (as Thamnium confertum, Brotherus 1924b). Dist. Juan Fernández Is.
Thamnobryum crassinervium (Broth.) S. He = Thamnobryum rigidum fide Robinson 1975.
Thamnobryum fasciculatum (Sw. ex Hedw.) I. Sastre
Ireland et al. 2007; He 1998 (as Porothamnium fasciculatum, Espinosa 1941; Brotherus
1924b). Dist. VIII. Arauco, Biobío, Concepción, Ñuble; XIV. Valdivia/Ranco; Juan Fernández
Is.
Thamnobryum ingae (Broth.) H. Rob.
He 1998; Robinson 1975 (as Thamnium ingae, Barrera 2006; Espinosa 1941; Brotherus 1924b;
Thériot 1927a; Bartram 1957). Dist. V. Valparaíso; Juan Fernández Is.
Thamnobryum neckeroides (Hook.) E. Lawton
Lawton 1971 (as Thamnium neckeroides, Thériot 1924). Dist. Without exact locality. Note:
The occurrence of this species in the region needs verification.
Thamnobryum proboscideum (Broth.) H. Rob.
Robinson 1975 (as Thamnium proboscideum, Brotherus 1924b; Bartram 1957). Dist. Juan
Fernández Is.
Thamnobryum rigidum (Mitt.) H. Rob.
Robinson 1975 (as Thamnobryum assimile, Sérgio 1981; as Thamnium assimile, Barrera 2006;
Espinosa 1941; Brotherus 1924b; as Thamnium latinerve, Brotherus 1924b; as Thamnium

92 Müller
rigidum, Mitten 1869; Brotherus 1924b; Bartram 1957; as Porothamnium crassinervium,

Espinosa 1941). Dist. Juan Fernández Is.
Thamnobryum spec.
Matcham et al. 2007. Dist. XII. Última Esperanza.
Thuidiopsis chilensis (Mitt.) Broth.
Hässel et al. 1984; Herzog 1922, 1954; Matteri 1985b (as Thuidium chilense, Mitten 1869).
Dist. XIV. Valdivia/Ranco; X. Osorno; XII. Magallanes, Última Esperanza.
Thuidiopsis corralensis (Broth.) Herzog = Thuidiopsis furfurosa fide Wijk et al. 1969.
Thuidiopsis dusenii (Broth.) Broth.
Cardot & Brotherus 1923 (as Thuidium dusenii, He 1998). Dist. VIII. Arauco; XIV.
Valdivia/Ranco; X. Chiloé.
Thuidiopsis filaris (Mitt.) Broth. = Thuidiopsis sparsa fide Touw 2001.
Thuidiopsis furfurosa (Hook.f. & Wilson) M. Fleisch.
Ireland et al. 2007; Touw 2001 (as Thuidium furfurosum, Larraín 2007b; Larraín 2005;
Villagran et al. 2003; Schaumann 2005; He 1998; Cardot & Brotherus 1923; Bartram 1952;
Robinson 1975; Deguchi 1991b; as Thuidiopsis corralensis, Herzog & Schwabe 1939; Herzog
1960; as Thuidium corralensis, Dusén 1903b; as Thuidiopsis furfurosum, Herzog & Schwabe
1939; as Thuidium masafuerae, Barrera 2006; Espinosa 1941; Brotherus 1924b; Robinson
1975; as Thuidium valdiviae, Brotherus 1924b; as Thuidiopsis valdiviae, Herzog & Schwabe
1939; Bartram 1957). Dist. VIII. Arauco; IX. Cautín; XIV. Valdivia/Ranco; X. Chiloé,
Llanquihue, Osorno, Palena; XI. Aisén; XII. Magallanes; Juan Fernández Is.
Thuidiopsis sparsa (Hook.f. & Wilson) Broth.
Tropicos 2009; Müller & Pereira 2006; Touw 2001 (as Thuidium filarium, Thériot 1918;
Reimers 1926; as Thuidiopsis filaris, Herzog & Hosseus 1938; Herzog 1954, 1960; as
Thuidium sparsum, Schaumann 2005; He 1998; Deguchi 1991b; as Thuidium valdiviae,
Barrera 2006; Espinosa 1941). Dist. VII. Cauquenes; VIII. Arauco, Biobío, Ñuble; IX. Cautín,
Malleco; XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno; XI. Aisén; Juan Fernández Is.
Thuidiopsis valdiviae (Broth.) Broth. = Thuidiopsis furfurosa fide Robinson 1975.
Thuidium chilense Mitt. ≡ Thuidiopsis chilensis.
Thuidium corralensis (Broth.) Broth. = Thuidiopsis furfurosa fide Wijk et al. 1969.
Thuidium dusenii Broth. ≡ Thuidiopsis dusenii.
Thuidium filarium Mitt. = Thuidiopsis sparsa fide Touw 2001.
Thuidium furfurosum (Hook.f. & Wilson) A. Jaeger ≡ Thuidiopsis furfurosa.
Thuidium inconspicuum Herzog
Herzog 1922. Dist. XIV. Valdivia/Ranco.
Thuidium masafuerae Broth. = Thuidiopsis furfurosa fide Touw & Van den Haak 1989.
Thuidium raphidostegum (Müll. Hal.) Mitt.
Mitten 1869; Herzog & Hosseus 1938. Dist. IX. Cautín.
Thuidium sparsum (Hook.f. & Wilson) A. Jaeger ≡ Thuidiopsis sparsa.
Thuidium valdiviae Broth. = Thuidiopsis sparsa fide Touw 2001.
Thysanomitrium laetevirens Broth. = Campylopus laxoventralis fide Frahm 1988.
Thysanomitrium leptodus (Mont.) Broth. = Campylopus clavatus fide Sainsbury 1955.
Thysanomitrium richardii (Brid.) Schwägr. ≡ Campylopus richardii.
Tortella hosseusii Herzog
Herzog & Hosseus 1938. Dist. IX. Cautín.
Tortella humilis (Hedw.) Jenn.
Tortella knightii (Mitt.) Broth.
Buck 2005a; Eckel 1997; Matteri & Schiavone 2002. Dist. XII. Antártica Chilena.
Tortella tortuosa (Hedw.) Limpr.
Tropicos 2009; Hässel et al. 1984; Ireland et al. 2007; He 1998; Matteri & Schiavone 2002;
Cardot & Brotherus 1923; Bartram 1952; Seki 1974; Matteri 1985b; Deguchi 1991b. Dist. RM.
Chacabuco, Santiago; VIII. Biobío; XIV. Valdivia/Ranco; X. Llanquihue, Osorno; XI. Aisén;
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Tortula amblyophylla (Mont.) Thér., hom. illeg. = Tortula platyphylla fide Mitten 1869.
Tortula andersonii Ångström = Syntrichia glacialis fide Müller 2009.
Tortula antarctica (Hampe) Paris ≡ Syntrichia antarctica.
Tortula appressa Mitt.
Mitten 1869. Dist. Chile, without exact locality (leg. Lobb). Note: The type was not studied by
Cano & Gallego (2008), therefore the taxonomic status is uncertain.
Tortula arenae (Besch.) Broth. ≡ Hennediella arenae.
Tortula atrata Thér. = Syntrichia breviseta fide Cano & Gallego 2008.
Tortula atrovirens (Sm.) Lindb.
Cano & Gallego 2008; Gallego & Cano 2007; Mitten 1869; Thériot 1921; Herzog 1954. Dist.
II. Antofagasta; IV. Elqui, Limarí; V. Aconcagua, Valparaíso; RM. Cordillera; VII. Talca.
Tortula atrovirens var. brevifolia Thér. = Tortula atrovirens fide Cano & Gallego 2008.
Tortula australasiae Hook.f. & Grev. ≡ Didymodon australasiae.
Tortula berteroana (Müll. Hal.) Broth. ex Paris = Tortula platyphylla fide Cano & Gallego 2008.
Tortula berthoana Thér.
Thériot 1926. Dist. V. Valparaíso. Note: Type material was studied by Cano & Gallego (2008).
The species belong to Syntrichia and is similar to S. fragilis, but differs in a few details.
Tortula brachypelma Dusén = Syntrichia magellanica fide Zander 1993.
Tortula breviseta Mont. ≡ Syntrichia breviseta.
Tortula bullata Herzog, hom. illeg. = Syntrichia ruralis fide Cano & Gallego 2008.
Tortula campestris Dusén ≡ Syntrichia campestris.
Tortula calycina (Schwägr.) Hook.f. & Grev. ≡ Barbula calycina.
Tortula chilensis (Mont.) Mitt. ≡ Chrysoblastella chilensis.
Tortula chrysopila (Müll. Hal.) Paris = Syntrichia princeps fide Matteri & Schiavone 2002.
Tortula chubutensis Dusén = Syntrichia ruralis fide Cano & Gallego 2008.
Tortula conferta Bartr. = Syntrichia magellanica fide Ochyra et al. 2008.
Tortula conotricha (Müll. Hal.) Paris = Syntrichia glacialis fide Müller 2009.
Tortula contorta (Hampe) Mont. = Syntrichia flagellaris fide Cano & Gallego 2008.
Tortula costesii Thér. ≡ Syntrichia costesii.
Tortula curta Cardot & Broth., hom. illeg. = Syntrichia costesii fide Wijk et al. 1969.
Tortula densifolia (Hook.f & Wilson) Herzog ≡ Hennediella densifolia.
Tortula diaguita (R.H. Zander & Mahu) Cano & M.T. Gallego
Cano & Gallego 2008 (as Hennediella diaguita, Zander & Mahu 1999). Dist. IV. Elqui,
Limarí.
Tortula didymodontoides Broth. = Syntrichia saxicola fide Ochyra et al. 2008.
Tortula epilosa Broth. ≡ Syntrichia epilosa.
Tortula epilosa var. pilifera Thér. ≡ Syntrichia epilosa var. pilifera.
Tortula excelsa Cardot = Syntrichia filaris fide Ochyra 1998.
Tortula flagellaris (Schimp.) Thér. ≡ Syntrichia flagellaris.
Tortula flavipes Broth., hom. illeg. = Tortula platyphylla fide Cano & Gallego 2008.
Tortula flaviseta Mitt. = Hennediella heimii fide Cano 2008.
Tortula fuegiana (Mitt.) Mitt. = Syntrichia magellanica fide Matteri 2003a.
Tortula fusca (Müll. Hal.) Mont. ≡ Didymodon fuscus.
Tortula fuscinervia Mitt. = Didymodon pruinosus fide Jiménez & Cano 2008.
Tortula fuscoviridis Cardot = Syntrichia saxicola fide Ochyra 1998.
Tortula geniculata Mont. = Didymodon australasiae fide Jiménez et al. 2005.
Tortula glacialis Kunze ex Müll. Hal. ≡ Syntrichia glacialis.
Tortula graminicolor (Müll. Hal.) Mont. = Didymodon australasiae fide Zander 1993.
Tortula gromschii Thér. ≡ Syntrichia gromschii.
Tortula grossiretis Cardot = Syntrichia magellanica fide Ochyra et al. 2008.
Tortula grossiretis var. atrata Cardot = Syntrichia magellanica fide Ochyra et al. 2008.
Tortula heimii (Hedw.) Mitt. ≡ Hennediella heimii.
Tortula heteroneura Cardot = Syntrichia magellanica fide Ochyra et al. 2008.
Tortula hoppeana (Schultz) Ochyra
Müller 2009; Cano & Gallego 2008. Dist. RM. Santiago.
Tortula jaffueli Thér.
Cano & Gallego 2008; Thériot 1923. Dist. IV. Choapa; V. Valparaíso; VI. Cachapoal.
Tortula kunzeana (Müll. Hal.) Mitt. ≡ Hennediella kunzeana.
Tortula laeta Kunze ≡ Rhamphidium laetum.
Tortula laevinervis Broth. ex Dusén = Syntrichia fragilis fide Cano & Gallego 2008.

94 Müller
Tortula lechleri (Müll. Hal.) Broth. = Syntrichia magellanica fide Zander 1993.
Tortula leucocalyx Mont. ≡ Pseudocrossidium leucocalyx.
Tortula lingulaefolia Cardot & Broth. = Hennediella arenae fide Lightowlers 1985.
Tortula lithophila Dusén ≡ Syntrichia lithophila.
Tortula litorea Cardot
Matteri & Schiavone 2002. Dist. XII. Tierra del Fuego. Note: Cano & Gallego (2008) have
studied type material and concluded the taxonomic position of this species in Syntrichia similar
to S. fragilis, but they doesn’t make the new combination.
Tortula macrocarpa (Schimp.) Mitt. ≡ Funaria macrocarpa.
Tortula magellanica Mont. ≡ Syntrichia magellanica.
Tortula mendozensis Mitt. ≡ Pseudocrossidium mendozense.
Tortula mniadelphus (Müll. Hal.) Broth. = Sagenotortula quitoensis fide Zander 1989.
Tortula mniadelphus var. cochlearifolia Herzog ≡ Syntrichia mniadelphus var. cochlearifolia.
Tortula modica R.H. Zander
(as Pottia intermedia, Müller 2002). Dist. VII. Talca.
Tortula monoica Cardot = Syntrichia magellanica fide Zander 1993.
Tortula muralis Hedw.
Tropicos 2009; Pereira et al. 2006; Espinosa 1941; He 1998; Mitten 1869; Dusén 1906a;
Williams 1915; Thériot 1921; Brotherus 1924a; Roivainen 1934; Herzog & Hosseus 1938;
Herzog & Schwabe 1939 (as Tortula muralis var. rupestris, Herzog & Hosseus 1938; as
Tortula muralis var. aestiva, Thériot 1921). Dist. IV. Choapa, Elqui, Limarí; V. Valparaíso;
RM. Santiago; VI. Cachapoal; VII. Cauquenes, Curicó, Talca; VIII. Arauco, Biobío,
Concepción, Ñuble; IX. Cautín; XIV. Valdivia/Ranco; X. Chiloé, Osorno.
Tortula muralis var. longipila Dusén = Tortula muralis fide Cano & Gallego 2008.
Tortula obscuriretis Thér.
Thériot 1918. Dist. IX. Malleco. Note: Fide Cano & Gallego (2008) this species belongs to
Pseudocrossidium and is similar to P. leucocalyx.
Tortula papillosa Wilson ≡ Syntrichia papillosa.
Tortula papillosa var. chilensis Thér. ≡ Syntrichia papillosa var. chilensis.
Tortula perarmata Broth. = Pseudocrossidium leucocalyx fide Cano & Gallego 2008.
Tortula perflaccida Broth. ex Dusén = Syntrichia flagellaris fide Zander 1993.
Tortula perrubiginosa (Dusén) Paris = Syntrichia glacialis fide Zander 1993.
Tortula pilifera Hook.f. = Pseudocrossidium crinitum fide Zander 1993.
Tortula pilifera var. denticulata (Dusén) Wijk & Margad. = Pseudocrossidium crinitum fide Zander 1993.
Tortula platyphylla Mitt.
Cano & Gallego 2008; Gallego & Cano 2007; Ireland et al. 2007; Mitten 1869 (as Tortula
amblyophylla, Thériot 1923, 1934b; Herzog & Schwabe 1939; Herzog 1954; as Barbula
graminicolor subsp. subgraminicolor, Thériot 1917; as Barbula subgraminicolor, Costes 1921;
as Tortula flavipes, Brotherus 1924a; Espinosa 1941). Dist. IV. Choapa, Elqui, Limarí; V.
Valparaíso; RM. Cordillera, Santiago; VI. Colchagua; VIII. Concepción; X. Llanquihue.
Tortula polycarpa Dusén = Hennediella arenae fide Cano & Gallego 2008.
Tortula princeps De Not. ≡ Syntrichia princeps.
Tortula princeps var. magellanica (Mont.) Lightowlers ≡ Syntrichia magellanica.
Tortula prostrata Mont. = Syntrichia serrulata fide Gallego et al. 2009.
Tortula pseudolatifolia Cardot ≡ Syntrichia glacialis Müller 2009.
Tortula pseudorobusta Dusén ≡ Syntrichia pseudorobusta.
Tortula pulvinatula Dusén = Syntrichia breviseta fide Cano & Gallego 2008.
Tortula punctulata (Brid.) Mitt. = Syntrichia robusta fide Zander 1993.
Tortula purpureo-velutina Herzog = Tortula platyphylla fide Cano & Gallego 2008.
Tortula pusilla Ångström = Syntrichia magellanica fide Zander 1993.
Tortula pygmaea Dusén = Syntrichia lithophila fide Ochyra & Zander 2007.
Tortula rivularis Dusén = Syntrichia fontana fide Lightowlers 1985.
Tortula robusta Hook.f. & Grev. ≡ Syntrichia robusta.
Tortula robusta var. laxa E.B. Bartram = Syntrichia robusta fide Zander 1993.
Tortula robusta var. runcinata (Müll. Hal.) Broth. = Syntrichia robusta fide Zander 1993.
Tortula robustula Cardot = Syntrichia filaris fide Matteri & Schiavone 2002.
Tortula santiagensis Broth. = Pseudocrossidium excavatum fide Cano & Gallego 2008.
Tortula saxicola Cardot ≡ Syntrichia saxicola.
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Tortula saxicola Dusén, hom. illeg. = Syntrichia lithophila fide Ochyra & Zander 2007.
Tortula scabrella Dusén ≡ Syntrichia scabrella.
Tortula scabrinervis (Müll. Hal.) Mont. ≡ Syntrichia scabrinervis.
Tortula schnyderi (Müll. Hal.) Broth. ≡ Syntrichia schnyderi.
Tortula serripungens (Lorentz & Müll. Hal.) Broth. ≡ Syntrichia serripungens.
Tortula serrulata Hook. & Grev. ≡ Syntrichia serrulata.
Tortula socialis Dusén ≡ Syntrichia socialis.
Tortula squarripila Thér. ≡ Syntrichia squarripila.
Tortula stenophylla Cardot & Broth., hom. illeg. = Syntrichia glacialis fide Cano & Gallego 2008.
Tortula subglacialis Thér. = Pseudocrossidium crinitum fide Zander 1993.
Tortula subpapillosa Cardot & Broth. ≡ Syntrichia subpapillosa.
Tortula subsessilis Brid. ≡ Pterygoneurum subsessilis.
Tortula truncata (Hedw.) Mitt.
Cano & Gallego 2008; Ireland et al. 2007. Dist. VIII. Concepción.
Tortula umbrosa Dusén = Leptodontium proliferum fide Cano & Gallego 2008.
Trematodon geniculatus Matteri
Deguchi 1991a. Dist. X. Osorno.
Trematodon pascuanus Thér.
Ireland & Bellolio 2002; Thériot 1937. Dist. Easter I.
Trichodon cylindricus (Hedw.) Schimp.
Müller 2009. Dist. VII. Talca.
Trichostomum brachydontium Bruch
Tropicos 2009; Espinosa 1941; Brotherus 1924b; Bartram 1957; Robinson 1975. Dist. Juan
Fernández Is.; Easter I.
Trichostomum elliottii Broth. ex Dusén
Dusén 1906a; Herzog 1954. Dist. RM. Chacabuco; VIII. Concepción.
Trichostomum tortella Müll. Hal. = Tortella spec. fide Wijk et al. 1969.
Trichostomum williamsii R.H. Zander
(as Astomum chilense, Williams 1915; Thériot 1921). Dist. IV. Elqui; V. Valparaíso.
Triquetrella filicaulis Dusén ≡ Triquetrella patagonica fo. filicaulis.
Triquetrella patagonica Müll. Hal.
Müller 2009; Pereira et al. 2006; Ireland et al. 2007; Müller & Pereira 2006; Thériot 1917;
Herzog & Hosseus 1938. Dist. II. Antofagasta; V. Valparaíso; VII. Cauquenes, Talca; VIII.
Arauco, Biobío, Concepción, Ñuble; IX. Cautín.
Triquetrella patagonica fo. filicaulis (Dusén) Herzog
Herzog & Hosseus 1938 (as Triquetrella filicaulis, Espinosa 1941; Dusén 1906a; Brotherus
1924a). Dist. VIII. Concepción; IX. Cautín.
Ulota anderssonii (Ångström) A. Jaeger = Orthotrichum elegantulum fide Lewinsky 1987.
Ulota brevicollis (Mitt.) A. Jaeger = Ulota rufula fide Malta 1927.
Ulota carinata Mitt.
Villagran et al. 2003; Herzog & Hosseus 1938; Herzog & Schwabe 1939 (as Orthotrichum
carinatum, Mitten 1869). Dist. X. Chiloé, Llanquihue.
Ulota chilensis (Mitt.) A. Jaeger = Ulota rufula fide Malta 1927.
Ulota eremitense Mitt. = Ulota glabella fide Malta 1927.
Ulota crenatoerosa (Müll. Hal.) Kindb. = Orthotrichum incanum fide Matteri 2000.
Ulota darwinii Mitt. ≡ Ulota fuegiana var. darwinii.
Ulota fernandeziana Malta
Bartram 1957; Robinson 1975. Dist. Juan Fernández Is.
Ulota fuegiana Mitt. var. fuegiana
Hässel et al. 1984; Larraín 2005; Matteri & Schiavone 2002; Dusén 1903b; Cardot 1901, 1908;
Cardot & Brotherus 1923; Bartram 1952; Seki 1974; Matteri 1985b; Deguchi 1991b (as
Orthotrichum fuegianum, Mitten 1869). Dist. XIV. Valdivia/Ranco; X. Chiloé, Llanquihue;
XI. Aisén; XII. Antártica Chilena, Magallanes, Última Esperanza.
Ulota fuegiana var. crispata Malta
Bartram 1952; Herzog 1954. Dist. XIV. Valdivia/Ranco; XI. Aisén.

96 Müller
Ulota fuegiana var. darwinii (Mitt.) Malta

Matteri & Schiavone 2002 (as Orthotrichum darwinii, Mitten 1869; as Ulota darwinii, Dusén
1903b). Dist. XII. Tierra del Fuego.
Ulota fulvella (Mitt.) Mitt. = Ulota luteola fide Wijk & Margadant 1961.
Ulota germana var. breviseta Malta
Herzog & Hosseus 1938; Seki 1974. Dist. XIV. Valdivia/Ranco.
Ulota germana (Mont.) Mitt. var. germana
Díaz et al. 2008; Larraín 2007b; Buck 2002; He 1998; Matteri & Schiavone 2002; Cardot
1908; Reimers 1926; Herzog 1922, 1954; Thériot 1935; Seki 1974; Deguchi 1991b (as
Orthotrichum germanum, Mitten 1869; as Ulota savatieri, Cardot 1900, 1901, 1908; Dusén
1903b; Cardot & Brotherus 1923). Dist. XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno;
XI. Aisén. XII. Antártica Chilena, Magallanes, Última Esperanza, Tierra del Fuego.
Ulota glabella Mitt.
Matteri & Schiavone 2002; Dusén 1903b; Cardot 1908; Seki 1974; Deguchi 1991b (as
Orthotrichum eremitense, Mitten 1869; as Orthotrichum glabellum, Mitten 1869; as Ulota
eremitense, Dusén 1903b; Cardot 1908). Dist. IX. Cautín; X. Llanquihue; XI. Aisén; XII.
Antártica Chilena.
Ulota gymnomitria Müll. Hal. = Orthotrichum elegantulum fide Lewinsky 1987.
Ulota hamata Dusén = Ulota magellanica fide Cardot 1908.
Ulota incana (Müll. Hal.) Besch. = Orthotrichum incanum fide Matteri 2000.
Ulota inclinata (Müll. Hal.) Kindb. = Orthotrichum inclinatum fide Matteri 2000.
Ulota lativentrosa Müll. Hal. ex Malta fo. angustifolia
Herzog & Hosseus 1938. Dist. XIV. Valdivia/Ranco; X. Llanquihue.
Ulota leiothecia (Müll. Hal.) A. Jaeger ≡ Orthotrichum leiothecium.
Ulota lobbiana Mitt.
Dusén 1903b; Cardot 1908 (as Orthotrichum lobbianum, Mitten 1869). Dist. X. Chiloé.
Ulota luteola (Hook.f. & Wilson) Wijk & Margad.
Matteri & Schiavone 2002; Deguchi 1991b (as Orthotrichum fulvellum, Mitten 1869; as Ulota
fulvella, Hässel et al. 1984; Eaton 1892; Dusén 1903b; Cardot 1908; Matteri 1985b). Dist. X.
Osorno; XI. Aisén; XII. Antártica Chilena, Magallanes, Última Esperanza.
Ulota macrocalycina Mitt.
Hässel et al. 1984; Buck 2002; Matteri & Schiavone 2002; Dusén 1903b; Cardot 1908; Matteri
1985b; Deguchi 1991b (as Orthotrichum macrocalycinum, Mitten 1869; as Ulota marginata,
Cardot 1908). Dist. X. Osorno; XII. Antártica Chilena, Magallanes, Última Esperanza.
Ulota macrodontia Dixon & Malta
Ireland et al. 2007; He 1998; Dusén 1903b; Herzog 1954; Seki 1974; Deguchi 1991b. Dist.
VIII. Arauco, Ñuble; IX. Cautín; X. Llanquihue; XI. Aisén.
Ulota magellanica (Mont.) A. Jaeger
Goffinet et al. 2006; Hässel et al. 1984; Matcham et al. 2002; Buck 2002; Matteri & Schiavone
2002; Dusén 1903b; Cardot 1905; Cardot 1908; Herzog & Hosseus 1938; Bartram 1952; Seki
1974; Matteri 1985b; Deguchi 1991b (as Orthotrichum magellanicum, Mitten 1869; as Ulota
hamata, Dusén 1903b). Dist. IX. Malleco; X. Llanquihue, Osorno; XI. Aisén; XII. Antártica
Ulota marginata (Ångström) A. Jaeger = Ulota macrocalycina fide Malta 1927.
Ulota persubulata Dusén, nom. nud. = Ulota rufula fide Malta 1927.
Ulota phyllantha Brid.
Buck 2002; Buck 2005b; Matteri & Schiavone 2002; Dusén 1903b; Cardot 1908; Cardot &
Brotherus 1923; Bartram 1952 (as Orthotrichum phyllanthum, Mitten 1869). Dist. XII.
Ulota pusilla Malta
Matteri & Schiavone 2002; Deguchi 1991b. Dist. IX. Malleco; X. Llanquihue, Osorno; XII.
Magallanes.
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Ulota pygmaeothecia (Müll. Hal.) Kindb.

Hässel et al. 1984; Buck 2002; Matteri & Schiavone 2002; Dusén 1903b; Cardot 1908; Cardot
& Brotherus 1923; Bartram 1952; Seki 1974; Matteri 1985b. Dist. XI. Aisén; XII. Antártica
Ulota rufula (Mitt.) A. Jaeger var. rufula
Larraín 2007b; Larraín 2005; Ireland et al. 2007; Villagran et al. 2003; He 1998; Herzog 1922,
1954; Brotherus 1924b; Roivainen 1934; Thériot 1935; Herzog & Hosseus 1938; Bartram
1957; Seki 1974; Robinson 1975; Deguchi 1991b (as Orthotrichum brevicolle, Mitten 1869; as
Orthotrichum chilense, Mitten 1869; as Orthotrichum rufulum, Mitten 1869; as Ulota
brevicollis, Thériot 1934b; as Ulota chilense, Thériot 1934b; as Ulota persubulata, Dusén
1903b). Dist. VIII. Arauco, Ñuble; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Chiloé,
Llanquihue, Osorno; XI. Aisén; Juan Fernández Is.
Ulota rufula var. patagonica Malta
Seki 1974. Dist. XI. Aisén.
Ulota savatieri Besch. = Ulota germana fide Malta 1927.
Ulota ventricosa (Müll. Hal.) Malta
Matteri & Schiavone 2002; Deguchi 1991b (as Zygodon ventricosus, Mitten 1869). Dist. X.
Llanquihue; XII. Magallanes.
Valdiviella pauperculata Thér. & Herzog ≡ Muscoherzogia pauperculata.
Verrucidens turpis (Cardot) Cardot ≡ Hymenoloma turpe.
Vesiculariopsis spirifolium (Dusén) Broth.
Seki 1974; Deguchi 1991a (as Ectropothecium spirifolium, Dusén 1903b, 1905a; Cardot 1908;
as Achrohypnella subenervis, Herzog 1954; Matteri 1972). Dist. X. Osorno; XI. Aisén, General
Carrera.
Vittia elimbata Hedenäs, Vanderpoorten & Goffinet
Matcham et al. in Blockeel et al. 2008a. Dist. XII. Última Esperanza.
Vittia pachyloma (Mont.) Ochyra
Tropicos 2009; Goffinet et al. 2006; Larraín 2007b; Matcham et al. 2007; Buck 2002; Buck
2005a; Larraín 2007a; Vanderpoorten et al. 2003; He 1998; Müller & Pereira 2006; Ochyra &
Matteri 2001; Matteri & Schiavone 2002; Ochyra 1987b; Deguchi 1991a (as Grimmia
pachyloma, Mitten 1869; as Sciaromium confluens, Dusén 1903b; Cardot 1908; as Sciaromium
conspissatum, Cardot 1908; Thériot 1915, Herzog & Schwabe 1939; Bartram 1952; Herzog
1954; Seki 1974; as Sciaromium conspissatum var. maritimum, Herzog 1954; as Sciaromium
crassinervatum, Herzog & Schwabe 1939; as Sciaromium depastum, Dusén 1903b; Cardot
1908; Cardot & Brotherus 1923; as Sciaromium krauseanum, Paris 1894-1900; as Sciaromium
maritimum, Cardot 1908; Cardot & Brotherus 1923; Thériot 1924; as Sciaromium nigritum,
Dusén 1905a; as Sciaromium obscurifolium, Mitten 1869; Thériot 1934b; Herzog 1954; as
Sciaromium pachyloma, Espinosa 1941; Brotherus 1924b; Reimers 1926; Bartram 1952, 1957;
Robinson 1975; as Sciaromium pachyloma var. brevifolia, Thériot 1924; as Sciaromium
pachyloma var. gracilescens, Herzog 1954). Dist. V. Los Andes, Quillota, Valparaíso; RM.
Chacabuco, Cordillera, Santiago, Talagante; VI. Colchagua; VII. Cauquenes, Curicó, Linares,
X. Chiloé, Llanquihue, Osorno; XI. Aisén, Coihaique; XII. Antártica Chilena, Magallanes,
Warburgiella aureonitida (E.B. Bartram) H. Akiy. = Rhaphidorrhynchium scorpiurus fide Seki 1974.
Warburgiella leucocytus (Müll. Hal.) B.C. Tan, W.B. Schofield & H.P. Ramsay
(as Rhaphidostegium leucocytus, Dusén 1903b; Cardot 1908). Dist. XII. Antártica Chilena.
Warnstorfia exannulata (Schimp.) Loeske
Müller 2009; Larraín 2007b; Buck 2002; Larraín 2007a; Ireland et al. 2007; He 1998; Ochyra
& Matteri 2001; Matteri & Schiavone 2002 (as Drepanocladus exannulatus, Seki 1974;
Deguchi 1991a). Dist. RM. Santiago; VII. Talca; VIII. Arauco; X. Chiloé, Osorno; XI. Aisén;

98 Müller
Warnstorfia fluitans (Hedw.) Loeske

Müller 2009; Buck 2002; He 1998; Matteri & Schiavone 2002 (as Drepanocladus fluitans,
Matteri 1985b; Hässel et al. 1984; Deguchi 1991a; as Drepanocladus fuegianus, Cardot &
Brotherus 1923; as Amblystegium fuegianum, Mitten 1869; as Hypnum fuegianum, Dusén
1903b; Cardot 1908; as Hypnum fluitans, Cardot 1908). Dist. V. San Antonio; VII. Talca; XII.
Warnstorfia fontinaliopsis (Müll. Hal.) Ochyra
Ochyra & Matteri 2001; Matteri & Schiavone 2002 (as Drepanocladus laculosus, Cardot &
Brotherus 1923; as Hypnum fluitans var. laculosum, Dusén 1903b; as Calliergidium austro-
stramineum, Seki 1974). Dist. XI. Aisén; XII. Magallanes, Tierra del Fuego.
Warnstorfia laculosa (Müll. Hal.) Ochyra & Matteri = Warnstorfia fontinaliopsis fide Ochyra & Matteri 2001.
Warnstorfia luipichensis (R.S. Williams) Hedenäs = Warnstorfia sarmentosa fide Ochyra et al. 2008.
Warnstorfia sarmentosa (Wahlenb.) Hedenäs
Ochyra et al. 2008; Buck 2002; Ochyra & Matteri 2001; Matteri & Schiavone 2002 (as
Calliergon sarmentosum, Hässel et al. 1984; Cardot & Brotherus 1923; Matteri 1985b;
Deguchi 1991a; as Warnstorfia luipichensis, Hedenäs 2003). Dist. XI. Aisén; XII. Antártica
Webera albicans Schimp., hom. illeg. = Pohlia wahlenbergii fide Shaw 1982.
Webera alticaulis (Müll. Hal.) Paris = Pohlia wahlenbergii var. glacialis fide Warburg 1962.
Webera costesii Cardot & Thér. = Pohlia chilensis fide Shaw 1987.
Webera cruda (Hedw.) Fürnr. ≡ Pohlia cruda.
Webera firmo-acuminata Dusén ex Paris., nom. nud. = Pohlia chilensis fide Suárez & Schiavone 2008.
Webera lonchochaete Dusén ≡ Pohlia lonchochaete.
Webera looseri Thér. ≡ Pohlia looseri.
Webera magnifica Herzog = Bryum australe fide Müller 2009.
Webera nutans Hedw. ≡ Pohlia nutans.
Webera philonotea (Müll. Hal.) Paris = Pohlia wahlenbergii fide Matteri 1998.
Webera sphagnadelphus (Müll. Hal.) Besch. = Pohlia nutans.
Webera timmiaecaulon Dusén, nom. nud., Dusén 1903b.
Weissia calcarea (Nees & Hornsch.) Müll. Hal. ≡ Gymnostomum calcareum.
Weissia chilensis Hampe ex Müll.Hal. = Oreoweisia erosa fide O’Shea & Matcham in Blockeel et al. 2004.
Weissia controversa Hedw. var. controversa
Ireland et al. 2007; Ireland & Bellolio 2002; Bartram 1957; Deguchi 1991b (as Weissia
flavipes, Espinosa 1941; Brotherus 1924; as Weissia patagonica, Hässel et al. 1984; Cardot &
Brotherus 1923; Matteri 1985b). Dist. VIII. Arauco, Biobío, Concepción, Ñuble; XI. General
Carrera; XII. Magallanes, Última Esperanza; Juan Fernández Is.; Easter I.
Weissia controversa var. amblyodon (Brid.) Sendtn.
(as Weissia viridula var. amblyodon, Herzog 1954). Dist. V. Aconcagua.
Weissia crispula Hedw. var. ambigua = Hymenoloma antarcticum fide Ochyra et al. 2008.
Weissia diffidentia R.H. Zander
(as Astomum recurvirostrum, Thériot 1924; as Tetrapterum recurvirostrum, Brotherus 1924).
Dist. V. Valparaíso.
Weissia flavipes Hook.f. & Wilson = Weissia controversa fide Wijk et al. 1969.
Weissia kunzeana Müll. Hal.
Pereira et al. 2006; Müller & Pereira 2006; Mitten 1869; Robinson 1975 (as Hymenostomum
kunzeanum, Brotherus 1924b; Thériot 1935; Herzog & Schwabe 1939; Herzog 1954). Dist. VI.
Colchagua, San Vicente; VII. Cauquenes, Talca; XIV. Valdivia/Ranco; X. Llanquihue,
Osorno; Juan Fernández Is.
Weissia longirostris Mitt., hom. illeg., Mitten 1869.
Weissia patagonica Cardot & Broth. = Weissia controversa fide Müller 2009.
Weissia poeppigiana (Müll. Hal.) Mitt. = Didymodon vinealis fide Sollman 1995.
Weissia viridula Hedw. ex Brid. var. amblyodon Schimp. ≡ Weisia controversa var. amblyodon.
Weissia spec.
Brotherus 1924. Dist. Easter I.
Weymouthia billardieri (Hampe) Broth. = Weymouthia cochlearifolia fide Streimann 2001.
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Weymouthia cochlearifolia (Schwägr.) Dixon

Larraín 2007b; Larraín 2005; Ireland et al. 2007; Villagran et al. 2003; He 1998; Quandt et al.
2001; Thériot 1935; Bartram 1952; Deguchi 1991b (as Pilotrichella krausei, Dusén 1903b; as
Weymouthia billardieri, Cardot 1908; Herzog 1922, 1940, 1954, 1960; Cardot & Brotherus
1923; Reimers 1926; Herzog & Hosseus 1938; Herzog & Schwabe 1939; Seki 1974; as
Coelidium cochlearifolium, Eaton 1892). Dist. VIII. Arauco; XIV. Valdivia/Ranco; X. Chiloé,
Llanquihue, Osorno; XI. Aisén, Capitán Prat.
Weymouthia mollis (Hedw.) Broth.
Larraín 2007b; Larraín 2005; Ireland et al. 2007; Espinosa 1941; Villagran et al. 2003; He
1998; Quandt et al. 2001; Müller & Pereira 2006; Cardot 1908; Herzog 1922, 1954, 1960;
Cardot & Brotherus 1923; Brotherus 1924a, 1924b; Reimers 1926; Thériot 1927a; Roivainen
1934; Herzog & Hosseus 1938; Bartram 1952, 1957; Seki 1974; Robinson 1975; Deguchi
1991b (as Meteorium molle, Mitten 1869; as Pilotrichella cumingii, Dusén 1903b). Dist. IV.
Limarí; VII. Cauquenes, Talca; VIII. Arauco; IX. Cautín; XIV. Valdivia/Ranco; X. Chiloé,
Llanquihue, Osorno, Palena; XI. Aisén; XII. Magallanes; Juan Fernández Is.
Weymouthia orbiculata Thér. ≡ Lembophyllum orbiculatum.
Willia austroleucophaea (Besch.) Broth.
Ochyra et al. 2008; Matteri & Schiavone 2002 (as Willia senilis, Brotherus 1924). Dist. XII.
Willia brachychaete (Dusén) R.H. Zander
NY 2009; Tropicos 2009; Müller 2009; Cano et al. 2008b. Dist. VII. Talca; VIII. Biobío; IX.
Cautín.
Willia grimmioides Müll. Hal. = Willia austroleucophaea fide Bell 1974.
Willia senilis Müll. Hal. = Willia austroleucophaea fide Bell 1974.
Wollnya wilsonii (Mitt.) Herzog ≡ Pohlia wilsonii.
Zygodon bartramioides Malta
Müller 2009; Calabrese 2006; Calabrese et al. 2006; He 1998; Herzog & Hosseus 1938;
Herzog 1954; Seki 1974; Deguchi 1991b (as Zygodon tristis, Herzog 1960). Dist. VII. Talca;
IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Osorno. Note: Records from region Aisén under
this name (Seki 1974; He 1998) belongs to Z. chilensis (Calabrese et al. 2006).
Zygodon chilensis Calabrese & F. Lara
Calabrese 2006; Calabrese et al. 2006. Dist. XIV. Valdivia/Ranco; XI. Aisén.
Zygodon corralensis Lorentz = Zygodon menziesii fide Matcham & O’Shea 2005.
Zygodon curvicaulis Dusén = Zygodon intermedius fide Malta 1926.
Zygodon cylindropyxis Müll. Hal. ex Malta = Zygodon bartramioides fide Calabrese 2006.
Zygodon denticulatus Taylor = Zygodon reinwardtii fide Malta 1926.
Zygodon gracillimus Dusén, p.p., nom. nud. = Zygodon magellanicus fide Calabrese 2006.
Zygodon gracillimus Dusén, p.p., nom. nud. = Zygodon papillatus fide Calabrese 2006.
Zygodon hookeri Hampe var. hookeri
Calabrese 2006; Matteri 1985b; Hässel et al. 1984. Dist. XII. Magallanes, Tierra del Fuego,
Última Esperanza.
Zygodon hookeri var. leptobolax (Müll. Hal.) Calabrese
Tropicos 2009; Larraín 2007b; Pereira et al. 2006; Larraín 2007a; Calabrese 2006 (as Zygodon
inermis, Larraín 2005; He 1998; Müller & Pereira 2006; Seki 1974; Deguchi 1991b; as
Zygodon theriotii, Herzog & Hosseus 1938). Dist. VII. Cauquenes, Curicó, Talca; VIII.
Biobío, Concepción, Ñuble; IX. Cautín; X. Chiloé, Llanquihue; XI. Aisén. XII. Magallanes.
Zygodon hyadesii Besch. = Zygodon reinwardtii fide Calabrese 2006.
Zygodon inermis Malta = Zygodon hookeri var. leptobolax fide Calabrese 2006.
Zygodon intermedius Bruch & Schimp.
Larraín 2005; Villagran et al. 2003; Calabrese 2006; He 1998; Müller & Pereira 2006; Cardot
& Brotherus 1923; Thériot 1927a; Herzog 1954; Bartram 1957; Seki 1974; Robinson 1975;
Deguchi 1991b (as Zygodon curvicaulis, Dusén 1903b; Cardot 1908; Cardot & Brotherus
1923). Dist. VII. Cauquenes, Linares; IX. Malleco; XIV. Valdivia/Ranco; X. Chiloé,

100 Müller
Llanquihue, Osorno; XI. Aisén, General Carrera; XII. Magallanes, Tierra del Fuego, Última
Zygodon jaffuelii Thér.
Calabrese 2006; He 1998; Thériot 1921; Herzog & Hosseus 1938; Herzog 1954; Deguchi
1991b. Dist. V. Valparaíso; IX. Cautín; XIV. Valdivia/Ranco; X. Llanquihue, Osorno; XI.
Aisén. Note: The species was synonymized with Zygodon liebmanii Schimp. by Allen (2002),
but is treated as a distinct species by Calabrese (2006).
Zygodon krausei Lorentz = Zygodon intermedius fide Calabrese 2006.
Zygodon magellanicus Dusén ex Malta
Müller 2009; Calabrese et al. 2002; Calabrese 2006; Matteri & Schiavone 2002; Seki 1974.
Dist. VII. Cauquenes; XI. Aisén; XII. Magallanes, Tierra del Fuego.
Zygodon menziesii (Schwägr.) Arn.
Calabrese 2006; He 1998; Matcham & O’Shea 2005; Mitten 1869; Brotherus 1924b; Herzog
1954; Robinson 1975; Matteri 1989 (as Zygodon corralensis, Herzog & Hosseus 1938; Herzog
1954; Seki 1974; Deguchi 1991b; as Codonoblepharon menziesii, Matteri & Schiavone 2002).
Dist. XIV. Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno; XI. Aisén; Juan Fernández Is.
Zygodon obovalis Mitt.
Brotherus 1924b. Dist. Juan Fernández Is. Note: The species is not treated or mentioned in the
revision of Calabrese (2006), therefore its taxonomic status is uncertain.
Zygodon obtusifolius Hook.
Calabrese 2006; Deguchi 1991b (as Bryomaltaea obtusifolia, Ireland et al. 2007). Dist. V.
Valparaíso; VIII. Arauco; IX. Malleco; XIV. Valdivia/Ranco; X. Llanquihue.
Zygodon obtusifolius var. antarcticus Malta = Zygodon obtusifolius fide Calabrese 2006.
Zygodon papillatus Mont.
Tropicos 2009; Larraín 2007b; Larraín 2007a; Calabrese 2006; Müller & Pereira 2006; Matteri
& Schiavone 2002; Mitten 1869; Herzog 1960; Seki 1974; Deguchi 1991b. Dist. VII.
Valdivia/Ranco; X. Chiloé, Llanquihue, Osorno; XI. Aisén; XII. Magallanes.
Zygodon papillosus Mont. ex Broth. = Zygodon papillatus fide Calabrese et al. 2002.
Zygodon pentastichus (Mont.) Müll. Hal.
Larraín 2007b; Hässel et al. 1984; Calabrese 2006; Mitten 1869; Herzog & Hosseus 1938;
Herzog & Schwabe 1939; Bartram 1952; Herzog 1954; Bartram 1957; Herzog 1960; Robinson
1975; Matteri 1985b (as Zygodon porteri var. crispatulus, Thériot 1926; as Zygodon
pentastichus var. crispatulus, Malta 1926; as Pentastichella pentasticha var. strictifolia,
Thériot 1924; as Zygodon porteri, Thériot 1926; as Pentastichella jaffueli, Thériot 1921; as
Pentastichella pentasticha, Tropicos 2009; Larraín 2005; Villagran et al. 2003; Müller &
Pereira 2006; Dusén 1903b; Thériot 1918; as Macromitrium poeppigii, Mitten 1869; as
Stenomitrium pentastichum, Espinosa 1941; Cardot 1908; Herzog 1922; Cardot & Brotherus
1923; Brotherus 1924b; Reimers 1926; Seki 1974; Deguchi 1991b; as Zygodon uncinatus,
Mitten 1869). Dist. V. Valparaíso; VII. Cauquenes, Linares, Maule, Talca; VIII. Arauco,
Biobío, Concepción, Ñuble; IX. Cautín, Malleco; XIV. Valdivia/Ranco; X. Chiloé,
Llanquihue, Osorno; XI. Aisén; XII. Magallanes, Última Esperanza; Juan Fernández Is.
Zygodon pentastichus var. crispatulus (Thér.) Malta = Zygodon pentastichus fide Calabrese 2006.
Zygodon pichinchensis (Taylor) Mitt.
Calabrese 2006; Seki 1974. Dist. XI. Aisén, Coihaique; XII. Última Esperanza.
Zygodon porteri Thér. var. porteri = Zygodon pentastichus fide Calabrese 2006.
Zygodon porteri var. crispatulus Thér. = Zygodon pentastichus fide Calabrese 2006.
Zygodon reinwardtii (Hornsch.) A. Braun
NY 2009; Calabrese 2006; Bartram 1952; Matteri 1985b (as Zygodon reinwardtii var. hyadesii,
Hässel et al. 1984; Matteri & Schiavone 2002; Seki 1974; Deguchi 1991b; as Zygodon
hyadesii, Dusén 1903b; Cardot 1908; Cardot & Brotherus 1923; as Zygodon denticulatus,
ISSN 0945-3466
Mosses of Chile 101
Mitten 1869). Dist. X. Chiloé, Llanquihue, Osorno; XI. Aisén; XII. Antártica Chilena,
Zygodon reinwardtii var. hyadesii (Besch.) Malta = Zygodon reinwardtii fide Calabrese 2006.
Zygodon robustulus Broth. ex Malta = Zygodon bartramioides fide Malta 1924.
Zygodon robustus Broth., nom. nud. = Zygodon bartramioides fide Calabrese 2006.
Zygodon theriotii Herzog = Zygodon hookeri var. leptobolax fide Calabrese 2006.
Zygodon tristis Herzog = Zygodon bartramioides fide Calabrese 2006.
Zygodon uncinatus Mitt. = Zygodon pentastichus fide Malta 1926.
Zygodon ventricosus Müll. Hal. ≡ Ulota ventricosa.
Taxa excluded or doubtful from Chile
Anacolia laevisphaera (Taylor) Flowers

The species is recorded under different names (A. laevisphaerea, A. subsessilis, A. subsessilis var.
brevifolia) from many regions of Chile (IV, V, RM, VI, VII, VIII, IX, XIV, X, XI, and Juan
Fernández Is.), e. g. by He (1998), Bartram (1957), Robinson (1975), Brotherus (1924b), Barrera
(2006), and Espinosa (1941). Matteri (1983) has demonstrated that the material of all Chilean
records checked by her represent Bartramia stricta. The occurrence of true Anacolia
hemisphaerica in Chile is in need of confirmation.
Atrichum androgynum (Müll.Hal.) A. Jaeger

The species is cited by Nyholm (1971) without exact locality for Chile, but left in all regional
publications of Chile. The record of this species and of the genus Atrichum in Chile is therefore
most unlikely and need further studies.
Bartramia halleriana Hedw.

All South American material referred to this species belongs to Bartramia mossmaniana (Fransén
2004a).
Bartramia ithyphylla Brid.

The species was excluded from Chile by Fransén (2004b). All records under this name from the
area belong to B. ithyphylloides.
Bartramia pomiformis Hedw.

The species occurs only in the northern hemisphere (Fransén 2004a). Records from the area
(Mitten 1869, Cardot 1908) must therefore belong to other species of the genus.
Breutelia pendula (Sm.) Mitt.

Dusén (1903b) and Cardot (1908) give records for the province of Antártica Chilena in Magallanes
region. The species was excluded from the area by Matteri (1973c). This is in accordance with
Jolley & Klazenga (2007), which describe the distribution as restricted to Australasia.
Breutelia polygastrica (Müll. Hal.) Broth.

Thériot (1934b) gives a record of this species under the synonym Breutelia cuspidatissima from
the province Valdivia. Griffin (1988) cited only records from Costa Rica to Bolivia and from
northern Argentina. The occurrence of this species in Chile is most unlikely and needs verification.
Bryohumbertia filifolia (Hornsch.) J.-P. Frahm

The species is cited in Brotherus (1924c) under the synonym Campylopus macrophyllus from
Chile, but in the original description (Müller 1848) and also in Mitten (1869) the record is given as
„Peru ad Cuchero: Pöppig“.

102 Müller
Bucklandiella heterosticha (Hedw.) Bednarek-Ochyra & Ochyra (Racomitrium heterostichum

[Hedw.] Brid.)
The records of this species from southern South America are doubtless erroneous (Ochyra et al.
2008).
Campylopus areodictyon (Müll. Hal.) Mitt.

The species was excluded from the area by Frahm (2005).
Campylopus asperifolius Mitt.

Mitten (1869) cited the species from “Andes Chilenses (Tatanara)”. The species was collected by
Lechler in Tatanara, which is situated in the today’s Peru (Frahm 1991).
Campylopus savannarum (Müll.Hal.) Mitt.

The species was excluded from Chile by Frahm (1998).
Codriophorus aquaticus (Brid. ex Schrad.) Bednarek-Ochyra & Ochyra (Racomitrium aquaticum

[Brid. ex Schrader] Brid.)
The species was excluded from the area by Bednarek-Ochyra (2006).
Conostomum tetragonum (Hedw.) Lindb.

Frahm et al. (1996) synonymized C. pentastichum with the northern hemispheric C. tetragonum.
Since this time the species is treated in different references under this name from Chile (e. g.,
Goffinet et al. 2006, Buck 2002, and Buck 2004). Ochyra et al. (2008) doesn’t accept this
synonymization and treated C. pentastichum as a clearly distinct taxon.
Didymodon pruinosus (Mitt.) R.H.Zander

The record of this species from Chiloé is probably a label mistake (Jiménez & Cano 2008).
Records of Bryoerythrophyllum fuscinervium, now a synonym of D. pruinosus, from Chile were
misidentified mainly with D. fuscus (Jiménez & Cano 2008).
Entosthodon balansae Besch.

The species was excluded from the Chilean bryoflora by Cuvertino & Buffa (2009). Specimens
from Chile reported under this name represent either Entostodon obtusifolius, Funaria chilensis or
Physcomitrium sp.
Fissidens julianus (Savi ex DC.) Schimp.

The record from Chile in Herzog (1954) belongs to Fissidens berteroi (Müller & Pursell 2003).
Fissidens osmundoides Hedw.

Specimens from Chile identified as such belong to Fissidens adianthoides (Müller & Pursell
2003).
Funaria erectiuscula Mitt.

The species is given with the record “Andes Chilensis, Mendoza (leg. Gillies)“ by Mitten (1869)
and is therefore included in the Chilean checklist by He (1998). This record is situated in the
nowadays territory of Peru and the species is therefore excluded from Chile.
Juratzkaea leptura (Taylor) W.A. Weber ex W.R. Buck

The Chilean record of this Australian and South African moss given by He (1998) is most unlikely
and needs confirmation.
ISSN 0945-3466
Mosses of Chile 103
Lepidopilum erectiusculum (Taylor) Mitt.

The species is given with the record “Andes Chilenses (Sachapata)“ by Mitten (1869) and is
therefore included in the Chilean checklist by He (1998). The record lie in the nowadays territory
of Peru and the species must therefore excluded from Chile.
Leptotrichella kunzeana (Müll. Hal.) Ochyra (Dicranella kunzeana Müll. Hal., Microdus
kunzeanus [Müll. Hal.] Broth.)
The species is cited as Dicranella kunzeana for Chile by Mitten (1869) with the record “Chili et
Peru ad Chuchero”. Therefore the species is included in the checklist of He (1998). The record of
Mitten is situated in the nowadays territory of Peru and the species must therefore excluded from
Chile.
Muelleriella angustifolia (Hook.f. & Wilson) Dusén

The species is cited by Dusén (1905a) from the Magallanes province. Following Vitt (1976) the
species is endemic to Campbell Island, New Zealand. The record of Dusén will therefore belong to
M. crassifolia.
Orthotheciella varia (Hedw.) Ochyra

The species recorded in different literature sources (e. g., Pereira et al. 2006, Buck 2002, Larraín
2007a, He 1998, Müller & Pereira 2006, Ochyra & Matteri 2001, and Matteri & Schiavone 2002)
as Orthotheciella varia for Chile represent in fact Cratoneuropsis relaxa subsp. minor (Ochyra et
al. 2008).
Philonotis fontanoides (Gillies & Grev.) A. Jaeger

The species is given under the name Bartramia fontanoides with the record “Andes Chilensis,
Mendoza (leg. Gillies)“ by Mitten (1869) and was therefore included in the Chilean checklist by
He (1998). The record is situated in the nowadays territory of Peru and the species must therefore
excluded from Chile.
Plagiomnium rostratum (Schrad.) T.J. Kop.

Dusén (1903b) and Cardot (1908) cited the species under the synonym Mnium rostratum from
“Fuegia, Patagonia australis”. Koponen (1979) doesn’t cite this species for South America.
Therefore the records of this species from the area will belong to other Plagiomnium species.
Pogonatum cirratum (Sw.) Brid. subsp. fuscatum (Mitt.) Hyvönen

The species is cited for Chile under the synonym Pogonatum spuriocirratum Broth. by Thériot
(1926). Fide Hyvönen (1989) the distribution area of this species is restricted to East Asia,
therefore the record from Chile is most unlikely.
Polytrichadelphus umbrosus Mitt.

He (1998) attributed the record of Mitten (1869) from “San Gavan” to Chile. In fact, San Gavan, is
situated in Peru and therefore the species is excluded from the Chilean bryoflora.
Pseudocrossidium mendozense (Mitt.) R.H. Zander

Mitten (1869) gives a record as Tortula mendozensis from “Andes chilensis, Mendoza (leg.
Gillies)“. This record is attributed in He (1998) to Chile, but in reality it is situated in the
nowadays Peru.
Pterygoneurum subsessilis (Brid.) Jur.

Mitten (1869) gives a record as Tortula subsessilis from “Andes chilensis, Mendoza (leg. Gillies)“.
This record is attributed in He (1998) to Chile, but in reality it is situated in the nowadays Peru.

104 Müller
Ptychomnion aciculare (Brid.) Mitt.

According to Hattaway (1984) all of the Chilean collections (e.g. Mitten 1869, Dusén 1903b,
Cardot 1908, Bartram 1952) under this name represent Ptychomnion cygnisetum or Ptychomnion
densifolium.
Ptychomnion fruticetorum Müll. Hal.

This species is only known from Brazil and doesn’t occur in Chile (Robinson 1970). However,
Hattaway (1984) considered it to be a form of Ptychomnion cygnisetum and placed Ptychomnion
horridum and Ptychomnion subaciculare in its synonyms.
Pyrrhobryum medium (Besch.) Manuel

The species was excluded from the area by Matteri & Piñero (1987). The Chilean records cited by
He (1998) belongs to P. mnioides.
Sphagnum imbricatum (Hornsch.) Russ.

Warnstorf (1911) gives a record from Chiloé. The species was excluded from the flora of the area
by Flatberg (1984).
Syntrichia sarconeurum (Hook. f. & Wilson) Ochyra & R.H. Zander

Records from western Patagonia in southern South America (e. g., Hässel et al. 1984, Matteri &
Schiavone 2002, Matteri 1982, and Matteri 1985b) are erroneous and correctly refer to Syntrichia
lithophila (Ochyra & Zander 2007, Ochyra et al. 2008).
Tortula vahliana (Schultz) Mont.

The species was cited from Quillota for Chile by Montagne (1850). Cano & Gallego (2008) have
excluded the species from the area.
Family placement
Families are arranged alphabetically. The taxonomic arrangement of the families and genera
follows with minor exceptions the classification of Goffinet et al. (2008). Following each genus is
the number of accepted taxa recorded for Chile. The 890 taxa are assigned to 230 genera in 68
families.
Amblystegiaceae
Amblystegium Schimp. 1 Anomodontaceae
Campylium (Sull.) Mitt. 2 Chileobryon Enroth 1
Cratoneuron (Sull.) Spruce 1 Haplohymenium Schwägr. 1
Cratoneuropsis (Broth.) M. Fleisch. 1
Drepanocladus Müll. Hal. 6 Archidiaceae
Leptodictyum (Schimp.) Warnst. 1 Archidium Brid. 2
Sanionia Loeske 2
Scorpidium (Schimp.) Limpr. 2 Aulacomniaceae
Vittia Ochyra 2 Aulacomnium Schwägr. 1
Andreaeaceae Bartramiaceae
Acroschisma Lindl. 1 Bartramia Hedw. 7
Andreaea Hedw. 24 Breutelia (Bruch & Schimp.) 10
ISSN 0945-3466
Mosses of Chile 105
Schimp. Distichophyllum Dozy & Molk. 15

Conostomum Sw. ex F. Weber & D. 3
Mohr Dicranaceae
Philonotis Brid. 12 Anisothecium Mitt. 6
Aongstroemia Bruch & Schimp. 1
Brachytheciaceae Camptodontium Dusén 1
Brachytheciastrum Ignatov & 3 Campylopodium (Müll. Hal.) Besch. 2
Huttunen Chorisodontium (Mitt.) Broth. 5
Brachythecium Schimp. 10 Dicranella (Müll. Hal.) Schimp. 6
Eriodon Mont. 1 Dicranoloma (Renauld) Renauld 22
Eurhynchiella M. Fleisch. 2 Eucamptodon Mont. 1
Eurhynchium Bruch & Schimp. 2 Muscoherzogia Ochyra 1
Kindbergia Ochyra 1 Platyneuron (Cardot) Broth. 1
Pseudoscleropodium (Limpr.) M. 1
Fleisch. Diphysciaceae
Rhynchostegiella (Schimp.) Limpr. 1 Diphyscium D. Mohr 1
Rhynchostegium Bruch & Schimp. 3
Sciuro-hypnum (Hampe) Hampe 4 Ditrichaceae
Astomiopsis Müll. Hal. 1
Bruchiaceae Ceratodon Brid. 3
Bruchia Schwägr. 1 Chrysoblastella R.S. Williams 1
Trematodon Michx. 2 Distichium Bruch & Schimp. 1
Ditrichum Hampe 14
Bryaceae Pleuridium Brid. 4
Anomobryum Schimp. 3 Rhamphidium Mitt. 2
Brachymenium Schwägr. 7 Skottsbergia Cardot 1
Bryum Hedw. 61 Trichodon Schimp. 1
Calliergonaceae Drummondiaceae
Straminergon Hedenäs 1 Drummondia Hook. 1
Warnstorfia Loeske 4
Encalyptaceae
Calymperaceae Encalypta Hedw. 6
Mitthyridium H. Rob. 1
Entodontaceae
Catagoniaceae Entodon Müll. Hal. 1
Catagonium Müll. Hal. ex Broth. 3
Eustichiaceae
Cryphaeaceae Eustichium (Brid.) Brid. 1
Cryphaea D. Mohr & F. Weber 2
Cryphaeophilum M. Fleisch. 1 Fabroniaceae
Dendrocryphaea Paris & Schimp. ex 4 Fabronia Raddi 4
Broth.
Fissidentaceae
Daltoniaceae Fissidens Hedw. 19
Achrophyllum Vitt & Crosby 8
Calyptrochaete Desv. 7 Funariaceae
Daltonia Hook. & Taylor 3 Entosthodon Schwägr. 7

106 Müller
Funaria Hedw. 6
Physcomitrium (Brid.) Brid. 3 Leptostomataceae
Leptostomum R. Brown 2
Gigaspermaceae
Costesia Thér. 1 Lepyrodontaceae
Neosharpiella H. Rob. & Delgad. 1 Lepyrodon Hampe 5
Grimmiaceae Leskeaceae
Bucklandiella Roiv. 11 Leptopterigynandrum Müll. Hal. 1
Codriophorus P. Beauv. 1
Coscinodon Spreng. 2 Leucobryaceae
Grimmia Hedw. 16 Atractylocarpus Mitt. 1
Niphotrichum (Bednarek-Ochyra) 1 Campylopus Brid. 18
Bednarek-Ochyra & Ochyra Microcampylopus (Müll. Hal.) 1
Racomitrium Brid. 4 Fleisch.
Schistidium Brid. 10 Pilopogon Brid. 1
Hedwigiaceae Leucomiaceae
Braunia Bruch & Schimp. 1 Leucomium Mitt. 1
Hedwigidium Bruch & Schimp. 1
Meesiaceae
Hypnaceae Amblyodon P. Beauv. 1
Calliergonella Loeske 1 Leptobryum (Bruch & Schimp.) 1
Chryso-hypnum Hampe 1 Wilson
Hypnum Hedw. 10 Meesia Hedw. 1
Isopterygiopsis Z. Iwats. 1 Neomeesia Deguchi 1
Mahua W.R. Buck 1
Mittenothamnium Henn. 1 Meteoriaceae
Platydictya Berk. 1 Cryptopapillaria M. Menzel 1
Pseudotaxiphyllum Z. Iwats. 1 Meteoriopsis M. Fleisch. 1
Meteorium (Brid.) Dozy & Molk. 1
Hypnodendraceae Papillaria (Müll. Hal.) Müll. Hal. 2
Hypnodendron (Müll. Hal.) Lindb. 1
Mniaceae
Hypopterygiaceae Cinclidium Schwägr. 1
Dendrohypopterygium Kruijer 1 Mielichhoferia Nees & Hornsch. 3
Hypopterygium Brid. 2 Plagiomnium T. J. Kop. 1
Lopidium Hook. f. & Wilson 1 Pohlia Hedw. 13
Schizymenium Harv. 6
Lembophyllaceae
Acrocladium Mitt. 1 Neckeraceae
Fallaciella H. A. Crum 1 Neckera Hedw. 4
Lembophyllum Lindb. 1 Porothamnium M. Fleisch. 4
Weymouthia Broth. 2 Porotrichodendron M. Fleisch. 1
Porotrichum (Brid.) Hampe 3
Leptodontaceae Thamnobryum Nieuwl. 7
Forsstroemia Lindb. 1
Leptodon D. Mohr 2 Oedipodiaceae
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Mosses of Chile 107
Oedipodium Schwägr. 1 Erythrophyllopsis Broth. 1

Eucladium Bruch & Schimp. 1
Orthodontiaceae Gertrudiella Broth. 1
Leptotheca Schwägr. 1 Gymnostomum Nees & Hornsch. 4
Orthodontium Schwägr. 3 Hennediella Paris 6
Hymenostylium Brid. 1
Orthotrichaceae Leptodontium (Müll. Hal.) Hampe 4
Macrocoma (Hornsch.) Grout 3 Molendoa Lindb. 1
Macromitrium Brid. 13 Pseudocrossidium R.S. Williams 8
Matteria Goffinet 2 Pterygoneurum Jur. 1
Muelleriella Dusén 1 Rhexophyllum Herzog 1
Orthotrichum Hedw. 24 Sagenotortula R.H. Zander 1
Pleurorthotrichum Broth. 1 Saitobryum R. H. Zander 1
Ulota D. Mohr 20 Splachnobryum Müll. Hal. 2
Zygodon Hook. & Taylor 13 Syntrichia Brid. 34
Tortella (Limpr.) Lindb. 4
Pilotrichaceae Tortula Hedw. 11
Callicostella (Müll. Hal.) Mitt. 2 Trichostomum Bruch 3
Cyclodictyon Mitt. 1 Triquetrella Müll. Hal. 1
Thamniopsis (Mitt.) M. Fleisch. 1 Weissia Hedw. 4
Willia Müll. Hal. 2
Plagiotheciaceae
Plagiothecium Bruch & Schimp. 6 Pterigynandraceae
Myurella Schimp. 1
Polytrichaceae
Atrichopsis Cardot 1 Pterobryaceae
Dendroligotrichum (Müll. Hal.) 2 Renauldia Müll. Hal. 1
Broth.
Hebantia G. L. Merr. 1 Ptychomitriaceae
Notoligotrichum G. L. Sm. 5 Campylostelium Bruch & Schimp. 1
Oligotrichum Lam. & DC. 5 Coscinodontella R.S. Williams 1
Pogonatum P. Beauv. 1 Jaffueliobryum Thér. 1
Polytrichadelphus (Müll. Hal.) Mitt. 1 Ptychomitrium Fürnr. 9
Polytrichastrum G. L. Sm. 3
Polytrichum Hedw. 5 Ptychomniaceae
Cladomniopsis M. Fleisch. 1
Pottiaceae Glyphothecium Hampe 2
Acaulon Müll. Hal. 1 Ombronesus N.E. Bell, N. Pedersen 1
Aloina Kindb. 4 & A.E. Newton
Aloinella Cardot 1 Ptychomniella (Broth.) W.R. Buck, 1
Barbula Hedw. 11 C.J. Cox, A.J. Shaw & Goffinet
Bryoerythrophyllum P.C. Chen 5 Ptychomnion (Hook.f. & Wilson) 3
Calyptopogon (Mitt.) Broth. 1 Mitt.
Chenia R. H. Zander 1
Cinclidotus P. Beauv. 1 Pylaisiadelphaceae
Crossidium Jur. 1 Aptychella (Broth.) Herzog 1
Didymodon Hedw. 19 Isopterygium Mitt. 2
Ephemerum Hampe 1

108 Müller
Racopilaceae
Racopilum P. Beauv. 4 Scouleriaceae
Scouleria Hook. 1
Rhabdoweisiaceae
Amphidium Schimp. 2 Seligeriaceae
Arctoa Bruch & Schimp. 1 Blindia Bruch & Schimp. 5
Holodontium (Mitt.) Broth. 1
Hymenoloma Dusén 5 Sematophyllaceae
Kiaeria I. Hagen 1 Rhaphidorrhynchium Besch. ex M. 8
Oreoweisia (Bruch & Schimp.) De 1 Fleisch.
Not. Sematophyllum Mitt. 8
Rhabdoweisia Bruch & Schimp. 2 Warburgiella Müll. Hal. 1
Symblepharis Mont. 3
Sphagnaceae
Rhacocarpaceae Sphagnum L. 19
Pararhacocarpus Frahm 1
Rhacocarpus Lindb. 1 Splachnaceae
Tayloria Hook. 6
Rhizogoniaceae Tetraplodon Bruch & Schimp. 1
Goniobryum Lindb. 1
Pyrrhobryum Mitt. 2 Stereophyllaceae
Rhizogonium Brid. 1 Catagoniopsis Broth. 1
Juratzkaea Lorentz 1
Rigodiaceae
Rigodium Schwägr. 7 Tetraphidaceae
Tetrodontium Schwägr. 1
Saulomataceae
Ancistrodes Hampe 1 Thuidiaceae
Sauloma (Hook.f. & Wilson) Mitt. 1 Thuidiopsis (Broth.) M. Fleisch. 4
Vesiculariopsis Broth. 1 Thuidium Bruch & Schimp. 2
Schimperobryaceae
Schimperobryum Margad. 2
Acknowledgements
I wish to thank Si He for allowing me to use the data’s of his preliminary published Chilean checklist (He
1998).
Financial support for travels to the herbariums of Stockholm (S) and Paris (PC) was provided by the
Synthesys program of the European Commission. I am grateful to the curators of PC and S, Catherine Rausch
and Lars Hedenäs, for assistance during my stay at their institutions and for the loan of specimens.
ISSN 0945-3466
Mosses of Chile 109
References
Abramov, I. I. & Abramova, A. L. 1983. Konspekt flor'i mhov Mongol'skoj Narodnoj Respubliki.
Biologiceskie Resursy i Prirodnye Uslovija Mongol'skoj Narodnoj Respubliki 17. 221 pp.
Akiyama, H. 1993. Taxonomic studies of mosses of Seram and Ambon (Moluccas, East Malesia) collected
by Indonesian-Japanese botanical expeditions. VII, Lembophyllaceae, Splachnaceae, Myuriaceae,
Neckeraceae, Brachytheciaceae, Amblystegiaceae and Sematophyllaceae, with additions to
previous reports on Dicranaceae and Pterobryaceae. Journal of the Faculty of Science: University of
Tokyo, Botany 15: 219–254.
Allen, B.H. 1987a. A revision of the Dicnemonaceae (Musci). Journal of the Hattori Botanical Laboratory
62: 1–100.
Allen, B.H. 1987b. A revision of the genus Leucomium (Leucomiaceae). Memoirs of the New York Botanical
Garden 45: 661–677.
Allen, B.H. 1994. Moss Flora of Central America. Part 1. Sphagnaceae-Calymperaceae. Monographs in
Systematic Botany from the Missouri Botanical Garden 49. 242 pp.
Allen, B. 1999. A revision of the moss genus Lepyrodon (Leucodontales, Lepyrodontaceae). Bryobrothera 5:
23–48.
Allen, B. 2002. Moss flora of Central America, Part 2. Encalyptaceae-Orthotrichaceae. Monographs in
Systematic Botany from the Missouri Botanical Garden 90: viii + 699 pages.
Allen, B. & Ireland, R.R. 2007. A new species of Bartramia (Bartramiaceae) from Chile. The Bryologist
110: 506–509.
Ando, H. 1972. Distribution and speciation in the genus Hypnum in the circum-Pacific region. Journal of the
Hattori Botanical Laboratory 35: 68–98.
Ando, H. 1982. Hypnum in Australasia and the Southern Pacific. Journal of the Hattori Botanical Laboratory
52: 93–106.
Ando, H. & Matteri, C.M. 1982. Patagonian bryophytes 7. Hypnum revolutum (Mitt.) Lindb., new to South
America, with notes on the Hypnum species from the Transecta zone, S Argentina. Lindbergia 8:
60–64.
Andrews, A.L. 1951. Taxonomic notes. X. The family Leptostomaceae. The Bryologist 54: 217–223.
Ångström, J. 1872. Förteckning och beskrifning öfver mossor, samlade af Professor N. J. Andersson under
Fregatten Eugenies verldsomsegling aören 1851–53. Öfversigt af Förhandlingar: Kongliga Svenska
Vetenskaps-Akademien 29 (4): 3–29.
Ångström, J. 1876. Primae lineae muscorum cognoscendorum, qui ad Caldas Brasiliae sunt collecti.
Öfversigt af Förhandlingar: Kongliga Svenska Vetenskaps-Akademien 33 (4): 3–50.
Arts, T. 1995. Bryum sauteri B., S. and G., recorded from Chile, Ecuador and South Africa, new to the
Americas and continental Africa. Journal of Bryology 18: 830.
Arts, T. 2001a. A revision of the Splachnobryaceae (Musci). Lindbergia 26: 77–96.
Arts, T. 2001b. The moss genus Leptobryum and the identity of Pohlia integra. Journal of Bryology 23: 325–
330.
Arts, T., Crundwell, A.C. & Whitehouse, H.L.K. 1995. Bryum valparaisense Thér. an earlier name for B.
pyriferum Crundw. & Whiteh. Journal of Bryology 18: 797–801.
Barrera, E. 2006. Tipos de musgos depositados en el Museo Nacional de Historia Natural, Santiago, Chile.
Boletin del Museo Nacional de Historia Natural, Chile, 55: 7–20.
Bartlett, J.K. & Vitt, D.H. 1986. A survey of species in the genus Blindia (Bryopsida, Seligeriaceae). New
Zealand Journal of Botany 24: 203–246.
Bartram, E.B. 1931. A review of the American species of Daltonia. Bulletin of the Torrey Botanical Club
58: 31–48.
Bartram, E.B. 1943. Additions to the moss flora of northern Chile. Farlowia 1: 191–194.
Bartram, E.B. 1946. New mosses from Tierra Del Fuego. Farlowia 2(3): 309–319.
Bartram, E.B. 1952. Mosses of Chile and Argentina mainly collected by R. Santesson. Svensk Botanisk
Tidskrift 46: 242–253.
Bartram, E.B. 1957. Mosses collected during Dr. and Mrs. C. Skottsberg's second expedition to the Juan
Fernández Islands, December 1954 to March 1955. Arkiv för Botanik utgivet av K. Svenska
Vetenskaps-Akademien 4: 29–43.
Bednarek-Ochyra, H. 1993. The identity of two neglected species of Racomitrium (Musci, Grimmiaceae)
from Patagonia. Fragmenta Floristica et Geobotanica, Supplement 2 (1): 83–90.

110 Müller
Bednarek-Ochyra, H. 2006. A Taxonomic Monograph of the Moss Genus Codriophorus P. Beauv.

(Grimmiaceae). W. Szafer Institute of Botany, Polish Academy of Sciences, Krakow, Poland, 276
pp.
Bednarek-Ochyra, H., Lewis-Smith, R.I. & Ochyra, R. 1999. The genus Plagiothecium (Plagiotheciaceae,
Bryopsida) in Antarctica. Bryobrothera 5: 55–64.
Bednarek-Ochyra, H. & Ochyra, R. 1994. Racomitrium lamprocarpum (Musci, Grimmiaceae) in southern
South America. Fragmenta Floristica et Geobotanica 39: 361–367.
Bednarek-Ochyra, H. & Ochyra, R. 2003. Racomitrium patagonicum, a new moss species from southern
South America. Journal of Bryology 25: 181–187.
Beever, J., Allison, K.W. & Child, J. 1992. Mosses of New Zealand (ed. 2). University of Otago Press, 214
pp. 82 fig. 72 pl.
Bell, B.G. 1984. A synoptic flora of South Georgian mosses: Grimmia and Schistidium. British Antarctic
Survey Bulletin 63: 71–109. 1 append. 20 fig.
Bell, N.E., Pedersen N. & Newton, A.E. 2007. Ombronesus stuvensis, a new genus and species of the
Ptychomniaceae (Bryophyta) from south west Chile. Taxon 56: 887–896.
Blöcher, R. & Frahm, J.-P. 2002. A comparison of the moss floras of Chile and New Zealand. Tropical
Bryology 21: 67–80.
Blockeel, T.L., Abay, G., Çetin, B., Bednarek-Ochyra, H., Ochyra, R., Lewis-Smith, R.I., Matteri,
C.M., Farias, R.M., Novotny, I., Rao, P., Enroth, J., van Rooy, J., Schiavone, M.M. & Vána,
J. 2003. New national and regional bryophyte records, 7. Journal of Bryology 25: 141–144.
Blockeel, T.L., Bednarek-Ochyra, H., Ochyra, R., Bruggeman-Nannenga, M.A., Gremmen, N.J.M.,
Hebrard, J.-P., Luis, L., Matcham, H.W., O’Shea, B.J. & Séneca, A. 2004. New national and
regional bryophyte records, 10. Journal of Bryology 26: 305–309.
Blockeel, T.L., Bednarek-Ochyra, H., Ochyra, R., Duckett, J.G., Erzberger, P., Hedenäs, L., Hugonnot,
V., Maier, E., Marková, I., Matcham, H.W., Plášek, V., Pócs, T., Seppelt, R.D., Szücs,
P., Thouvenot, L., van Zanten, B.O. 2008a. New national and regional bryophyte records, 18.
Journal of Bryology 30: 161–167.
Blockeel, T.L., Bergamini, A., Brusa, G., Ertz, D., Sergio, C., Garcia, C., Hedenäs, L., Müller, F.,
Nieuwkoop, J.A.W., Sabovljevic, M.S. 2000. New national and regional bryophyte records, 3.
Blockeel, T.L., Arts, T., Bruggeman-Nannenga, M.A., Brusa, G., Hedenäs, L., Novotný, I., Ochyra, R.,
Sabovljevic, M.S. & Townsend, C.C. 2002. New national and regional bryophyte records, 6.
Blockeel, T.L., Abay, G., Bakalin, V.A., Bednarek-Ochyra, H., Ochyra, R., Çetin, B., Cykowska,
B., Fuertes, E., Hespanhol, H., Holyoak, D.T., Hradílek, Z., Keçeli, T., Kürschner,
H., Larraín, J., Long, D.G., Parolly, G., Piątek, J., Piątek, M., Rams, S., Ros, R.M., Séneca,
A., Sérgio, C., Soldán, Z., ŞtefanuŃ, S., Uyar, G., Váňa, J. & Tonguç Yayintaş, Ö., 2008b. New
national and regional bryophyte records, 19. Journal of Bryology 30: 231–237.
Blockeel, T.L., Bakalin, V.A., Bednarek-Ochyra, H., Ochyra, R., Buck, W. R., Choi, S., Cykowska,
B., Erdağ, A., Erzberger, P., Kirmaci, M., Kürschner, H., Lebouvier, M., Papp,
B., Sabovljević, M., Sabovljević, A., Schröder, W., Singh, S. M., Sun, B.-Y., Townsend, C.C.,
Váňa, J. & Tonguç Yayintaş, Ö., 2009. New national and regional bryophyte records, 20. Journal
of Bryology 31: 54–62.
Bowers, F.D. 1974. The mosses reported from Costa Rica. The Bryologist 77: 150–171.
Brotherus, V.F. 1900. Die Laubmoose der ersten Regnellschen Expedition. Bihang til Kongliga Svenska
Vetenskaps- Akademiens Handlingar 26 (3): 1–76.
Brotherus, V.F. 1902-1909. Musci. – In: A. Engler & K. Prantl, Die Natürlichen Pflanzenfam. 1(3). Leipzig.
Brotherus, V.F. 1905. Pleurorthotrichum, eine neue Laubmoosgattung aus Chile. Oefversigt af
Förhandlingar, Finska Vetenskaps-Societeten 47: 1–3.
Brotherus, V.F. 1924. Musci Insulae-Paschalis. – In: Skottsberg, C. (ed.), The Natural History of Juan
Fernández and Easter Island 2: 241–246. Uppsala.
Brotherus, V.F. 1924a. Musci nonnulli Chilense a C. Skottsberg anno 1917 lectae. Acta Horti
Gothoburgensis 1: 189–195.
Brotherus, V.F. 1924b. The Musci of the Juan Fernández Islands. – In: Skottsberg, c. (ed.), The Natural
History of Juan Fernández and Easter Island 2: 409–448. Uppsala.
Brotherus, V.F. 1924c. Musci. – In: A. Engler & K. Prantl, Die Natürlichen Pflanzenfam. ed. 2, 10: i–iv, 1–
478. Leipzig.
ISSN 0945-3466
Mosses of Chile 111
Brotherus, V.F. 1925. Musci. – In: A. Engler & K. Prantl, Die Natürlichen Pflanzenfam. ed. 2, 11: i–iv, 1–
542. Leipzig.
Bruggeman-Nannenga, M.A. & Pursell, R.A. 1995 [1996]. Notes on Fissidens V. Lindbergia 20: 49–55.
Buck, W.R. 1977. A taxonomic investigation of Juratzkaea Lor. and Juratzkaeella gen. nov. Revue
Bryologique et Lichénologique 43: 309–325.
Buck, W.R. 1981. A review of Cheilothela (Ditrichaceae). Brittonia 33: 453–456.
Buck, W.R. 1983. A synopsis of the South American taxa of Fabronia (Fabroniaceae). Brittonia 35: 248–
254.
Buck, W.R. 1987. Taxonomic and nomenclatural rearrangements in the Hookeriales with notes on West
Indian taxa. Brittonia 39: 210–224.
Buck, W.R. 1989. Miscellaneous notes on Antillean mosses, 2: Rhaphidostichum (Sematophyllaceae) in the
New World. Moscosoa 5: 189–193.
Buck, W.R. 2002. Preliminary Key to the Mosses of Isla Navarino, Chile (Prov. Antártica Chilena). –
Published by the author: Bronx, NY. 147 pp.
Buck, W.R. 2004. Exploración briológica en Cabo de Hornos. 1era parte. Briolatina 51: 1–4.
Buck, W.R. 2005a. Exploración briológica en Cabo de Hornos. 2a parte. Briolatina 52: 1–4.
Buck, W. R. 2005b. Exploración briológica en Cabo de Hornos. 3a parte. Briolatina 53: 1–5.
Buck, W. R. & Goffinet, B. 2000. Morphology and classification of mosses. – pp. 71–123. In: Shaw, A.J. &
Goffinet, B. (ed.), Bryophyte Biology. Cambridge University Press.
Buck, W.R. & Ireland, R.R. 1989. Plagiotheciaceae. Flora Neotropica Monographs 50: 1–22.
Buck, W.R. & Landrum, L. 1977. A new species of Astomiopsis (Ditrichaceae). The Bryologist 80: 359–
362.
Burley, J.S. & Pritchard, N.M. 1990. Revision of the genus Ceratodon (Bryophyta). Harvard Papers in
Botany 2: 17–76.
Calabrese, G.M. 2006. A taxonomic revision of Zygodon (Orthotrichaceae) in southern South America. The
Bryologist 109: 453–509.
Calabrese, G.M., Elias, M.J. & Lara, F. 2006. A new species of Zygodon (Orthotrichaceae) from southern
South America. Journal of Bryology 28: 97–103.
Calabrese, G.M., Garilleti, R., Mazimpaka, V. & Lara, F. 2002. Lectotypification and differentiation of
Zygodon magellanicus Malta and Z. papillatus Mont. (Orthotrichaceae, Musci), two often confused
Andean-Patagonian species. Nova Hedwigia 74: 149–157.
Cano, M.J. 2003. New records and range extension of some mosses in tropical areas of Chile. Tropical
Bryology 24: 15–20.
Cano, M.J. 2008. Taxonomic revision of Hennediella Paris (Pottiaceae, Bryophyta). Bryophytorum
Bibliotheca 64, 142 pp.
Cano, M.J. & Gallego, M.T. 2008. The genus Tortula (Pottiaceae, Bryophyta) in South America. Botanical
Journal of the Linnean Society 156: 173–220.
Cano, M.J., Gallego, M.T., Jiménez, J.A. & Guerra, J. 2008a. Aloina obliquifolia (Pottiaceae, Bryophyta)
new to South America, and new reports of Aloina in the Neotropics. Cryptogamie, Bryologie 29:
75–81.
Cano, M.J., Jiménez, J.A., Churchill, S.P. & Guerra, J. 2008b. New records of Pottiaceae (Bryophyta) for
South America. Cryptogamie, Bryologie 29: 183–191.
Cao, T., Guo, S. & Yu, J. 2003. Preliminary studies on distribution pattern of the genus Ptychomitrium
(Musci) in the world. Journal of the Hattori Botanical Laboratory 93: 247–258.
Cao, T., Zuo, B., Guo, S.-L., Hyvönen, J. & Virtanen, V. 2006. New synonyms and combinations in the
genus Ptychomitrium (Bryopsida: Ptychomitriaceae). Journal of the Hattori Botanical Laboratory
100: 41–52.
Cardot, J. 1900. Note preliminaire sur les mousses recueillies par l'expedition Antarctique Belge. Revue
Bryologique 27: 38–46.
Cardot, J. 1901. Expedition Antarctique Belge. Resultats du Voyage du S.Y. Belgica en 1897–1899. Voyage
Belgica, Mousses. 48 pp.
Cardot, J. 1905. Notice preleminaire sur les mousses recueillies par l’Expedition antarctique suedoise.
Bulletin de l’herbier Boissier 5: 997–1011.
Cardot, J. 1908. La flora bryologique des Terres magellaniques, de la Georgie du Sud et de l'Antarctide. –
In: Wissenschaftliche Ergebnisse der Schwedischen Südpolar-Expedition 1901–1903, 4: 1–298,
plates 1–11.

112 Müller
Cardot, J. & Brotherus, V.F. 1923. Les Mousses. – In: Botanische Ergebnisse der Schwedischen Expedition
nach Patagonien und dem Feuerlande 1907–1909. Bihang til Kongliga Svenska Vetenskaps-
Akademiens Handlingar 63: 1–73.
Churchill, S.P. & Linares E.L. 1995a. Prodromus bryologiae Novo-Granatensis: introducción a la flora de
musgos de Colombia. Parte 1: Adelotheciaceae a Funariaceae. Biblioteca José Jerónimo Triana 12:
1–453 + i–xxvi.
Churchill, S.P. & Linares C., E.L. 1995b. Prodromus bryologiae Novo-Granatensis: introducción a la flora
de musgos de Colombia. Parte 2: Grimmiaceae a Trachypodaceae. Biblioteca José Jerónimo Triana
12: 455–924 + 2 unnumbered pages.
Clifford, H.T. 1955. On the distribution of Rhacomitrium crispulum (Hook.f. & Wils.) Hook.f. & Wils. The
Costes, N. 1921. Nomenclatura de los principales musgos de la Hoya de Marga-Marga (Quilpue-Provincia de
Valparaíso). Revista Chilena de Historia Natural 1921: 130–132.
Crosby, M.R. 1977. Florschuetzia, a new genus of Buxbaumiaceae (Musci) from Southern Chile. The
Crosby, M.R. 1978. Muscoflorschuetzia (Musci), a new name for Florschuetzia Crosby, hom. illeg. The
Bryologist 81: 338.
Crosby, M.R., Magill, R.E., Allen, B. & He, S. 1999. A Checklist of the Mosses. St. Louis: Missouri
Botanical Garden.
Crum, H. 1984. Part 11: Sphagnopsida, Sphagnaceae. – North American Flora, Series II, iv + 180 pp. 61 fig.
Crum, H. 1990. Comments on Sphagnum sect. Sphagnum in South America. Contributions from the
University of Michigan Herbarium 17: 71–81.
Cuvertino, J., Miserere, L. & Buffa, G. 2009. Costesia macrocarpa comb. nov. for Funaria macrocarpa
(Gigaspermaceae). The Bryologist 112: 287–289.
Cuvertino, J. & Buffa, G. 2009. Entosthodon balansae (Funariaceae) is excluded from the Chilean
bryophyte flora. Cryptogamie, Bryologique 30: 425–432.
Deguchi, H. 1983. Neomeesia Deguchi, a new genus of the family Meesiaceae from Southern South
America. Bulletin of the National Science Museum, Tokyo, Series B, Botany 9: 143–148.
Deguchi, H. 1984. Studies on some Patagonian species of Grimmiaceae (Musci, Bryophyta). – In: H. Inoue
(ed.) Studies on Cryptogams in South Chile. pp. 17–72. Kensei-Sha, Tokyo.
Deguchi, H. 1987. Studies on some Peruvian species of Grimmiaceae (Musci, Bryophyta). – In: H. Inoue
(ed.) Studies on Cryptogams in Southern Peru. pp. 19–74, Kensei-Sha, Tokyo.
Deguchi, H. 1991a. A list of moss collections made during the Expeditions to Chile in 1981 and 1987 (1).
Bulletin of the National Science Museum, Tokyo, Series B, Botany 17 (1): 15–34.
Deguchi, H. 1991b. A list of moss collections made during the Expeditions to Chile in 1981 and 1987 (2).
Bulletin of the National Science Museum, Tokyo, Series B, Botany 17 (2): 67–83.
Deguchi, H. & Oginuma, K. 1989. Cytological studies on some Chilean mosses. Lindbergia 15: 87–90.
Delgadillo, C. 1973. A new species, nomenclatural changes, and generic limits in Aloina, Aloinella, and
Crossidium (Musci). The Bryologist 76: 271–277.
De Luna, E. & Biasuso, B. 2001. Morphometric variation and species delimitation in the Braunia exserta
complex (Hedwigiaceae). – Botany 2001: Plants and People. Abstracts. Botanical Society of
America: Albuquerque, NM. p. 7.
Díaz, M. F., Larraín, J., Zegers, G. & Tapia, C. 2008. Caracterización florística e hidrológica de turberas
de la Isla Grande de Chiloé, Chile. Revista Chilena de Historia Natural 81: 455–468.
Dixon, H.N. 1914. Studies in the bryology of New Zealand, with special reference to the herbarium of Robert
Brown. Part II. New Zealand Institute Bulletin 3: 31–74.
Brown. Part IV. New Zealand Institute Bulletin 3: 153–238.
Brown. Part VI. New Zealand Institute Bulletin 3(6): 299–372.
Döbbeler, P. 1999. Two new species of Bryonectria (Hypocreales, Ascomycetes) on bryophytes. Sendtnera
6: 93–102.
Döbbeler, P. 2003. Ascomycetes on Dendroligotrichum (Musci). Nova Hedwigia 76: 1–44.
Döbbeler, P. & Triebel, D. 2000. Potridiscus polymorphus (Leotiales) - a new ascomycete on Polytrichaceae
(Musci) with palaeoaustral distribution. Hoppea 61: 71–83.
ISSN 0945-3466
Mosses of Chile 113
Donat, A. 1936. Resultados fitogeograficos de la Expedicion patagonica "Gaea" 1933. I. Algunas Briofitas
nuevas de la Cordillera Central Patagonica. Revista Sudamericana de Botánica 3: 61–68.
Duby, J.-É. 1880. Choix de mousses exotiques nouvelles ou mal counues [3]. Mémoires de la Société de
Physique et d'Histoire Naturelle de Genève 27: 1–10.
Dusén, P. 1903a. Beiträge zur Bryologie der Magellansländer von Westpatagonien und Südchile. I. Arkiv för
Botanik utgivet av K. Svenska Vetenskaps-Akademien 1 (1): 441–466, Tafl. 18–28.
Dusén, P. 1903b. Patagonian and Fuegian mosses. Part III. Reports of the Princeton University Expeditions
to Patagonia, Botany 8: 63–125.
Dusén, P. 1905a. Musci nonnulli novi e Fuegia et Patagonia reportati. Botaniska Notiser 1905: 299–310.
Dusén, P. 1905b. Beiträge zur Bryologie der Magellansländer von Westpatagonien und Südchile. II. Arkiv
för Botanik utgivet av K. Svenska Vetenskaps-Akademien 4 (1): 1–45, Tafl. 1–11.
Dusén, P. 1905c. Beiträge zur Bryologie der Magellansländer von Westpatagonien und Südchile. III. Arkiv
Dusén, P. 1906a. Beiträge zur Bryologie der Magellansländer von Westpatagonien und Südchile. IV. Arkiv
Dusén, P. 1906b. Beiträge zur Bryologie der Magellansländer von Westpatagonien und Südchile. V. Arkiv
Eaton, D.C. 1892. List of mosses from Fuegia and Patagonia. Contributions from the United States National
Herbarium 1: 138–139.
Eckel, P.M. 1997. Tortella knightii, an Australasian species new to South America in Tierra del Fuego. The
Eddy, A. 1977. Sphagnales of tropical Asia. Bulletin of the British Museum (Natural History), Botany 5:
359–445 + pl.
Enroth, J. 1991. Bryophyte flora of the Huon Peninsula, Papua New Guinea. XLIII. Funariaceae (Musci).
Acta Botanica Fennica 143: 57–69.
Enroth, J. 1992. Notes on the Neckeraceae (Musci). 11–12. The taxonomic position of Pinnatella
callicostelloides and Bissetia lingulata, with the description of Chileobryon (Anomodontaceae).
Nova Hedwigia 54: 137–146.
Espinosa, M.R. 1941. Lista sistematica de Musgos y Liquenes chilenos continentales e insulares, enviados
por el Dr. C. Skottsberg. Boletín del Museo Nacional de Historia Natural 19: 113–132.
Fife, A. 1982. Taxonomic and nomenclatural observations on the Funariaceae 2. Lectotypification of

Physcomitrium subg. Cryptopyxis (C. Müll.) Broth. Lindbergia 8: 75–76.
Fife, A. 1987. Taxonomic and nomenclatural observations on the Funariaceae 5. A revision of the Andean
species of Entosthodon. Memoirs of the New York Botanical Garden 45: 301–325.
Flatberg, K.I. 1984. A taxonomic revision of the Sphagnum imbricatum complex. Kongelige Norske
Videnskabers Selskab Skrifter 3: 1–80.
Fleischer, M. 1920. Kritische Revision von C. Müllerschen Laubmoosgattungen. IV. Hedwigia 63: 209–216.
Frahm, J.-P. 1976. Zur Campylopus-Flora von Chile. Herzogia 4: 141–160.
Frahm, J.-P. 1987. A revised list of the Campylopus species of the world. Bryologische Beiträge 7: 1–117.
Frahm, J.-P. 1988. The Subantarctic and southern Hemispheric species of Campylopus (Dicranaceae) with
contributions to the origin and speciation of the genus. Journal of the Hattori Botanical Laboratory
64: 367–387.
Frahm, J.-P. 1991. Dicranaceae: Campylopodioideae, Paraleucobryoideae. Flora Neotropica 54.
Organization for Flora Neotropica by The New York Botanical Garden: New York. 238 pp.
Frahm, J.-P. 1996. Revision der Gattung Rhacocarpus Lindb. (Musci). Cryptogamie: Bryologie,
Lichénologie 17: 39–65.
Frahm, J.-P. 1998. Taxonomische Notizen zur Gattung Campylopus XVII. Cryptogamie: Bryologie,
Frahm, J.-P. 1999. A type catalogue of Campylopodioideae and Paraleucobryoideae (Musci, Dicranaceae),
Part II, Campylopus. Tropical Bryology 16: 17–102.
Frahm, J.-P. 2001a. Aquatic mossballs of Blindia inundata in Patagonia. The Bryologist 104: 503.
Frahm, J.P. 2001b. Mossballs of Rigodium implexum (Rigodiaceae) on the forest floor in Chile. The
Frahm, J.-P. 2005. A contribution to the Campylopus flora of Chile. Tropical Bryology 26: 39–43.

114 Müller
Frahm, J.-P., Börner, H., Streiber, N., Wallau, B. & Weitkus, S. 1996. Revision der Gattung Conostomum
(Musci, Bartramiaceae). Tropical Bryology 12: 97–114.
Frahm, J.-P., Buchbender, V., Lachmann, S., Reifenrath, K. & Werner, F. 1998. Revision der Gattung
Oncophorus (Musci, Dicranaceae). Tropical Bryology 14: 119–131.
Frahm, J.-P., Klöcker, T., Schmidt, R. & Schöter, C. 2000. Revision der Gattung Amphidium (Musci,
Dicranaceae). Tropical Bryology 18: 173–184.
Frahm, J.-P., Rembold, K., Röver, T. & Schamell, T. 2003. Synopsis der Gattung Pyrrhobryum (Musci,
Rhizogoniaceae). Tropical Bryology 24: 115–127.
Fransén, S. 2004a. A taxonomic revision of Bartramia Hedw. section Bartramia. Lindbergia 29: 113–122.
Fransén, S. 2004b. A taxonomic revision of extra-Neotropical Bartramia section Vaginella C. Müll.
Lindbergia 29: 73–107.
Frey, W. & Schaumann, F. 2002. Records of rare southern South American bryophytes. Studies in austral
temperate rain forest bryophytes 18. Nova Hedwigia 74: 533–543.
Frisvoll, A.A. 1986. Southern hemisphere synonyms of Racomitrium sudeticum (Funck). Bruch et Schimp.
Froehlich, J. 1953. Zwei neue Laubmoose aus Süd-Chile und Feuerland. Annalen des Naturhistorischen
Museums in Wien 59: 117–118.
Fuentes, F. 1913 [1914]. Reseña botánica sobre la Isla de Pascua. Boletin del Museo Nacional de Chile 5 (2):
320–337.
Gallego, M.T. & Cano, M.J. 2007a. A new species of Syntrichia Brid. (Pottiaceae, Bryophyta) from Chile.
Gallego, M.T. & Cano, M.J. 2007b. New reports of Syntrichia Brid. (Pottiaceae, Bryophyta) taxa from
South America. Nova Hedwigia 85: 457–461.
Gallego, M.T., Cano, M.J. & Guerra, J. 2009. New synonymy in Syntrichia (Pottiaceae) in the Neotropics.
The Bryologist 112: 173–177.
Giese, M. & Frahm, J.-P. 1985 [1986]. A revision of Campylopodium (C. Müll.) Besch. Lindbergia 11:
125–133.
Goffinet, B., Buck, W.R., Massardo, F. & Rozzi, R. 2006. Miniature Forests of Cape Horn/Los Bosques en
Miniatura del Cabo de Hornos. Universidad de Magallanes: Punta Arenas, Chile. 254 pp.
Goffinet, B., Buck, W.R. & Shaw, A.J. 2008. Morphology and classification of the Bryophyta. In: Goffinet,
B. & Shaw, A.J. (eds.): Bryophyte Biology, 2nd edition, p. 55–138, Cambridge University Press.
Goffinet, B., Buck, W.R. & Wall, M.A. 2007. Orthotrichum freyanum (Orthotrichaceae), a new epiphytic
moss from Chile. Nova Hedwigia Beiheft 131: 1–11.
Gradstein, S.R. & Sipman, H.J.M. 1978. Taxonomy and World Distribution of Campylopus introflexus and
C. pilifer (= C. polytrichoides): a New Synthesis. The Bryologist 81: 114–121.
Greene, D.M. 1986. A Conspectus of the Mosses of Antarctica, South Georgia, the Falkland Islands and
southern South America. 314 pp., 1 tab. Institute of Terrestrial Ecology. Natural Environment
Research Council, Cambridge.
Greene, S.W. 1968 [1969]. Studies in Antarctic bryology. II. Andreaea, Neuroloma. Revue Bryologique et
Lichénologique 36: 139–146.
Griffin, D. 1981. El género Sphagnum en Los Andes de Colombia y Venezuela. Clave para las especies
frecuentes u ocasionales con notas ecológicas y taxonómicas. Cryptogamie: Bryologie,
Griffin, D. 1986. On the occurrence of Breutelia affinis (Hook.) Mitt. in South America. Lindbergia 12: 144.
Griffin, D. 1988. New world species of Breutelia with erect-appressed leaf bases. Nova Hedwigia 90: 357–
382.
Griffin, D. & Buck, W.R. 1989. Taxonomic and phylogenetic studies on the Bartramiaceae. The Bryologist
92: 368–380.
Griffin, D., Gradstein, S.R. & Aguirre, J. 1982. Studies on Colombian cryptogams XVII. On a new
antipodal element in the Neotropical Paramos - Dendrocryphaea latifolia sp. nov. (Musci). Acta
Botanica Neerlandica 31: 174–184.
Grout, A.J. 1929. Checklist of pleurocarpous mosses of North America. 32 pp. – New Brighton, New York.
Grout, A.J. 1935. Moss Flora of North America, North of Mexico 2: 1–284. Newfane, Vermont.
Grout, A.J. 1944. Preliminary synopsis of the North American Macromitriae. The Bryologist 47: 1–22.
Gunckel, H. 1938. Los Porothamnium de la flora Corraleña. Revista Chilena de Historia Natural 42: 118–
122.
ISSN 0945-3466
Mosses of Chile 115
Gunckel, H. 1965. Un interesante musgo de la selva Valdiviana: Eriodon conostomus Montagne. Revista
Universitaria (Santiago) 27: 67–70.
Guo, S. & Cao, T. 1999. Two neglected species of Ptychomitrium (Musci), with new synonyms. The
Hässel de Menéndez, G.G., Greene, S.W. & Matteri, C.M. 1984. The occurrence and distribution of
Bryophytes in southern Patagonia between latitude 51° and 52°S. Journal of the Hattori Botanical
Laboratory 55: 45–64.
Hattaway, R.A. 1981. Commentary on the genus Glyphothecium (Ptychomniaceae). The Bryologist 84: 344–
347.
Hattaway, R.A. 1984. A monograph of the Ptychomniaceae (Bryopsida). – 311 pp. 50 fig. Doctoral
dissertation. University Microfilms. Ann Arbor.
He, S. 1998. A checklist of the mosses of Chile. Journal of the Hattori Botanical Laboratory 85: 103–189.
He, S. 2005. A revision of the genus Leptopterigynandrum (Bryopsida, Leskeaceae). Journal of the Hattori
Botanical Laboratory 97: 1–38.
Hedenäs, L. 2003. Amblystegiaceae (Musci). Flora Neotropica Monograph 89. The New York Botanical
Garden. 108 pp.
Herzog, T. 1916. Neue Laubmoose aus Ostasien und Südamerika. Hedwigia 57: 233–250.
Herzog, T. 1922. Beiträge zur Bryophytenflora Chiles. Hedwigia 64: 1–18.
Herzog, T. 1926. Beiträge zur Moosflora Westpatagoniens. Hedwigia 66: 79–92.
Herzog, T. 1928. Drei neue Laubmoosgattungen. Revue Bryologique et Lichénologique 1: 98-108.
Herzog, T. 1937. Neue Bryophyten vom Ruwenzori und aus dem Patagonischen Inlandeisgebiet.
Repertorium Specierum Novarum Regni Vegetabilis 41: 285–292.
Herzog, T. 1940. Die Moose der Expedition Ljungner nach Patagonien, 1932/34. Arkiv för Botanik utgivet av
K. Svenska Vetenskaps-Akademien 29 (21): 1–17.
Herzog, T. 1954. Zur Bryophytenflora Chiles. Revue Bryologique et Lichénologique 23: 27–99.
Herzog, T. 1960. Weitere Beiträge zur Bryophytenflora Chiles. Revue Bryologique et Lichénologique 29:
183–206.
Herzog, T. & Hosseus, C. 1938. Contribucion al conocimiento de la Flora Briofita del sur de Chile. Archivos
de la Escuela de Fármacia de la Facultad de Ciencias Médicas de Córdoba 7: 3–95.
Herzog, T., Schwabe, G.H. & Schwabe, E. 1939. Zur Bryophytenflora Südchiles. Beihefte zum Botanischen
Centralblatt 60b: 1–51.
Herzog, T., Thériot, I., Buch, H., Schiffner, V. & Verdoorn, F. 1933. Bryophyta nova (6–16). Annales
Bryologici 6: 124–135.
Holyoak, D. T. 2004. Taxonomic notes on some European species of Bryum (Bryopsida: Bryaceae). Journal
of Bryology 26: 247-264.
Horton, D.G. 1983. A revision of the Encalyptaceae (Musci), with particular reference to the North
American taxa. Part II. Journal of the Hattori Botanical Laboratory 54: 353–532.
Huttunen, S., Isoviita P. & Quandt, D. 2008. Proposals to conserve the name Meteorium with a conserved
type and change the conserved type of Papillaria (Musci: Meteoriaceae). Taxon 57: 992–995.
Hyvönen, J. 1987. A synopsis of the genus Leptostomum (Leptostomataceae, Musci). Acta Botanica Fennica
24: 63–72.
Hyvönen, J. 1989. A synopsis of the genus Pogonatum (Polytrichaceae, Musci). Acta Botanica Fennica 138:
1–87.
Hyvönen, J. 1991. Chorisodontium (Dicranaceae, Musci) in southern South America. Annales Botanici
Fennici 28: 247–258.
Ireland, R.R. 1991. A preliminary study of the moss genus Isopterygium in Latin America. Caldasia 16(78):
265–276.
Ireland, R.R., 1992. The moss genus Isopterygium (Hypnaceae) in Latin America. Tropical Bryology 6:
111–132.
Ireland, R.R. & Bellolio, G. 2002. The Mosses of Easter Island. Tropical Bryology 21: 11–19.
Ireland, R.R., Bellolio, G., Bednarek-Ochyra, H. & Ochyra, R. 2005. Fertile plants of Racomitrium
patagonicum (Bryopsida, Grimmiaceae). Journal of the Hattori Botanical Laboratory 98: 205–211.
Ireland, R.R., Bellolio, G., Rodríguez, R. & Larraín, J. 2007 [2006]. Studies on the moss flora of the Bío-
Bío region of Chile. Tropical Bryology 28: 63–77.

116 Müller
Ireland, R.R., Robinson, H. & Bellolio, G. 2006. On the distribution and morphology of Hebantia rigida
(Bryopsida, Polytrichaceae). Journal of the Hattori Botatical Laboratory 100: 201–209.
Jiménez, J.A. 2007. Bryoerythrophyllum berthoanus comb. nov. and B. sharpii (Pottiaceae) in South
America. Journal of Bryology 29: 125–130.
Jiménez, J.A. & Cano, M.J. 2006. Two new combinations in Didymodon (Pottiaceae) from South America.
Jiménez, J.A. & Cano, M.J. 2007. Didymodon coquimbensis (Pottiaceae), a new species from Chile. The
Jiménez, J.A. & Cano, M.J. 2008. Taxonomic assessment of Didymodon pruinosus (Mitt.) R.H. Zander
(Bryophyta, Pottiaceae) a poorly known South American taxon. Nova Hedwigia 87: 145–152.
Jiménez, J.A., Ros, R.M., Cano, M.J. & Guerra, J. 2005. A new evaluation of the genus Trichostomopsis
(Pottiaceae, Bryophyta). Botanical Journal of the Linnean Society 147: 117–127.
Jolley, H. & Klazenga, N. 2007. A morphometric study of Breutelia pendula and B. elongata (Bryophyta,
Bartramiaceae). Australian Systematic Botany 20: 82–91.
Kabiersch, W. 1936. Studien über die ostasiatischen Arten einiger Laubmoosfamilien (Mniaceae-
Bartramiaceae). Hedwigia 76: 1–94.
Kindberg, N. C. 1888. Enumeratio Bryinearum Exoticarum 1–83. Linkoeping.
Koponen, A. 1977. Tayloria subgen. Pseudotetraplodon, subgen. nov., and new combinations in
Brachymitrion, Moseniella and Tayloria (Splachnaceae, Musci). Annales Botanici Fennici 14: 193–
196.
Koponen, A. 1982. The family Splachnaceae in Australasia and the Pacific. Journal of the Hattori Botanical
Koponen, T. 1968. Generic revision of Mniaceae Mitt. (Bryophyta). Annales Botanici Fennici 5: 117–151.
Koponen, T. 1969. The taxonomic status and typification of Mnium subpunctatum. The Bryologist 72: 61–
62.
Koponen, T. 1974. A guide to the Mniaceae in Canada. Lindbergia 2: 160–184.
Koponen, T. 1979. A synopsis of Mniaceae (Bryophyta). I. South and Central American taxa. Journal of the
Hattori Botanical Laboratory 46: 155–161.
Kramer, W. 1988. Beiträge zur Systematik und Bryogeographie einiger Sippen von Tortula Hedw. sect.
Rurales De Not. (Pottiaceae, Musci) unter besonderer Berücksichtigung der Südhemisphäre.
Journal of the Hattori Botanical Laboratory 65: 81–144.
Kruijer, J. D. 2002. Hypopterygiaceae of the World. Blumea, Supplement 13, 388 pp.
Kuc, M. 1996. On the distribution of Ancistrodes Hampe, an endemic moss of Chile. Cryptogamie:
Bryologie, Lichénologie 17: 119–125.
Kühnemann, O. & Goncalves Carralves, M.F. 1975. Monografia preliminar para la flora criptogámica
fuegiana: Los musgos de la familia Ptychomniaceae (Bryophyta). Darwiniana 19: 583–617.
Kühnemann, O. & Goncalves Carralves, M.F. 1976. Sobre Cryphaeophilum molle (Dus.) Fleisch.
Meteoriaceae-Briofitas. Boletin de la Sociedad Argentina de Botanica 17: 247–251.
Larraín J. 2005. Musgos de la Cordillera de la Costa de Valdivia, Osorno y Llanquihue: consideraciones

ecológicas y lista de especies. – In: Smith-Ramírez, C., Armesto, J.J. & Valdovinos, C. (ed.):
Historia, Biodiversidad y Ecología de los Bosques Costeros de Chile. Editorial Universitaria.
Santiago, Chile, p. 159–177.
Larraín, J. 2007a. Adiciones a la flora de musgos de la Isla Grande de Chiloé, Chile. Gayana, Botánica 64:
7–23.
Larraín, J. 2007b. Musgos (Bryophyta) de la estación biológica Senda Darwin, Ancud, isla de Chiloé: lista
de especies y claves para su identificación. Chloris Chilensis 10 (1). URL:
http://www.chlorischile.cl
Larraín, J., Herrera, F., Budke, J.M. & Goffinet, B. 2009. Phylogenetic affinities and conservation status
of the Chilean endemic Costesia spongiosa (Gigaspermaceae). The Bryologist 112: 278–286.
Lawton, E. 1971. Moss Flora of the Pacific Northwest. The Hattori Botanical Laboratory, xiii + 362 pp. +
195 pl.
Lewinsky, J. 1984. Orthotrichum Hedw. in South America 1. Introduction and taxonomic revision of taxa
with immersed stomata. Lindbergia 10: 65–94.
ISSN 0945-3466
Mosses of Chile 117
Lewinsky, J. 1987. Orthotrichum Hedw. (Orthotrichaceae) in South America 2. Taxonomic revision of taxa
with superficial stomata. Memoirs of the New York Botanical Garden 45: 326–370.
Lewinsky-Haapasaari, J. 1994. The genus Pleurorthotrichum Broth. Lindbergia 19: 11–24.
Lewinsky, J. & Deguchi, H. 1989. Orthotrichum truncatum Lewinsky & Deguchi, sp. nov. from Chile, and a
new record for O. macrosporum Lewinsky. The Bryologist 92: 216–218.
Lightowlers, P.J. 1985. Taxonomic notes on New Zealand species of Tortula. Journal of Bryology 13: 369–
375.
Lin, S.H. 1984. A taxonomic revision of Phyllogoniaceae (Bryopsida). Part II. Journal of Taiwan Museum
37(2): 1–54.
Lorentz, P.G. 1864. III. Pugillus specierum novarum e exoticarum. – In: Lorentz, P.G., Moosstudien.
Leipzig, Wilhelm Engelmann, p. 155–171.
Lorentz, P.G. 1866. Musci frondosi in Chile prope Valdiviam et prope Corral lecti per Dr. Krause.
Botanische Zeitung (Berlin) 24: 185–189.
Magill, R.E. 1981. Flora of Southern Africa. Bryophyta. Part 1. Mosses. Fasc. 1. Sphagnaceae-Grimmiaceae.
– Botanical Research Institute, Department of Agriculture and Fisheries, Pretoria.
Magombo, Z.L.K. 2003. Taxonomic revision of the moss family Diphysciaceae M. Fleisch. (Musci). Journal
of the Hattori Botanical Laboratory 94: 1–86.
Mahu, M. 1981. Las Briofitas del Bosque Las Petras, Quintero, Provincia de Valparaíso, Chile. The
Mahu, M. 1985. Tetrodontium brownianum (Dicks.) Schwaegr. (Bryatae: Tetraphidaceae) nuevo para Chile.
Malta, N. 1924. Studien über die Laubmoosgattung Zygodon 11-12. Latvijas Universitates Raksti 10: 303-
335.
Malta, N. 1926. Die Gattung Zygodon Hook. et Tayl., eine monographische Studie. Latvijas Universitates
Botaniska Darza Darbi 1, 185 pp.
Malta, N. 1927. Die Ulota-Arten Süd-Amerikas. Acta Horti Botanici Universitatis Latviensis 2: 165–208.
Margadant, W.D. 1972. Notes on the nomenclature of Musci. Lindbergia 1: 121–129.
Matcham, H.W. 2008. Correction to the bryoflora of south-western Patagonia. Field Bryology 94: 38.
Matcham, H.W. & O’Shea, B.J. 2005. A review of the genus Codonoblepharon Schwägr. (Bryopsida:
Orthotrichaceae). Journal of Bryology 27: 129–135.
Matcham, H.W., Pressel, S., Russell, S., Massardo, F. & Duckett, J.G. 2007. Inventory and conservation
of the bryoflora of south-western Patagonia. Field Bryology 93: 2–8.
Matsui, T. & Iwatsuki, Z. 1990. A taxonomic revision of the family Ditrichaceae (Musci) of Japan, Korea
and Taiwan. Journal of the Hattori Botanical Laboratory 68: 317–366.
Matteri, C.M. 1968. Las especies de "Philonotis (Bartramiaceae)" del sur de Argentina. Revista del Museo
Argentino Ciencias Naturales "Bernardino Rivadavia" e Instituto Nacional de Investigación de las
Ciencias Naturales Botánica 3 (4): 185–234.
Matteri, C.M. 1972. Las Hookeriaceae (Musci) Andino-Patagonicas II. Revista del Museo Argentino
Ciencias Naturales "Bernardino Rivadavia" e Instituto Nacional de Investigación de las Ciencias
Naturales Botánica 4 (2): 243–280.
Matteri, C.M. 1973a. Notas briologicas (Musci) I–II. Anales del Instituto de la Patagonia 4: 273–274.
Matteri, C.M. 1973b. Revision de las Hypopterygiaceae (Musci) Austrosudamericanas. Boletín de la
Sociedad Argentina de Botánica 25 (2–3): 229–250.
Matteri, C.M. 1973c. El genero Breutelia (Bartramiaceae, Musci) en la region Andino-Patagonica. Revista
del Museo Argentino Ciencias Naturales "Bernardino Rivadavia" e Instituto Nacional de
Investigación de las Ciencias Naturales Botánica 4 (4): 321–360.
Matteri, C.M. 1975a. Hookeriales. – In: Guarrera, S.A., Gamundi de Amos, I. and Rabinovich de Halperin,
D. (eds.), Flora criptogamica de Tierra del Fuego. Buenos Aires, Fundacion para la Educacion, la
Cienca y la Cultutra 14 (9): 7–60.
Matteri, C.M. 1975b. Las Hookeriaceae (Musci) Andino-Patagonicas, I. Nova Hedwigia 26: 649–724.
Matteri, C.M. 1979. Notas briologicas (Musci) IV y V. The Bryologist 82: 99–102.
Matteri, C.M. 1981. Patagonian bryophytes 4. Breutelia elongata (Hook.f. et Wils.) Mitt. and Myurella
julacea (Schwaegr.) B.S.G. Lindbergia 7: 61–65.
Matteri, C.M. 1982. Patagonian bryophytes 6. Fruiting Sarconeurum glaciale (C. Muell.) Cardot et Bryhn
newly found in southern Patagonia. Lindbergia 8: 105–109.

118 Müller
Matteri, C.M. 1983. Sobre Bartramia stricta (Musci, Bartramiaceae) en la región Andino-patagónica.
Boletín de la Sociedad Argentina de Botánica 22: 231–241.
Matteri, C.M. 1984a. Occurrence of tubers and rhizomes in Chrysoblastella chilensis (Mont.) Reim.
Matteri, C.M. 1984b. Sinopsis de las especies Andino-patagonicas, Antarcticas y subantarcticas de los
generos Bartramia, Bartramidula, Conostomum (Bartramiaceae, Musci). Darwiniana 25: 143–162.
Matteri, C.M. 1985a. Bryophyta, Musci: Bartramiaceae. – In: Guarrera, S.A., Gamundi de Amos, I. &
Rabinovich de Halperin, D. (eds.), Flora Criptogámica de Tierra del Fuego 14 (7), 62 pp. 18 pl.
Matteri, C.M. 1985b. Catalogo de los musgos. In: Boelcke, O., Moore, D.M. & Roig, F.A. (eds.), Transecta
botanica de la Patagonia austral. Buenos Aires, Consejo Nacional de Investigaciones Cientificas y
Technicas (Argentina), Instituto de la Patagonia (Chile) and Royal Society (Gran Bretana), 265–
298.
Matteri, C.M. 1988. Distribution maps of mosses from Tierra del Fuego, Argentina. 1. Lindbergia 14: 176–
178.
Matteri, C.M. 1989. Additions to the Moss Flora of Tierra del Fuego (Argentina). The Bryologist 92: 174–
177.
Matteri, C.M. 1993. Distribution maps of mosses from Tierra del Fuego, Argentina. 5. Lindbergia 18: 138–
140.
Matteri, C.M. 1994. Tortula subpapillosa and T. papillosa in southern South America with a new
combination in Syntrichia. Journal of the Hattori Botanical Laboratory 75: 33–40.
Matteri, C.M. 1996. Ditrichum brevirostrum (R. Brown ter.) Broth. newly discovered in northern Patagonia
and the Fuegian region. Lindbergia 21: 3–8.
Matteri, C.M. 1998. New synonyms and combinations in southern South American mosses. Fragmenta
Floristica et Geobotanica 43: 298–300.
Matteri, C.M. 2000. Sobre la identidad de Orthotrichum inclinatum, O. incanum y O. crenato-erosum
(Musci, Orthotrichaceae). Revista Museo Argentino Ciencias Naturales 2: 17–21.
Matteri, C.M. 2003a. New combination and new synonyms in Fuegian mosses. Lindbergia 28: 80–82.
Matteri, C.M. 2003b. Los musgos (Bryophyta) de Argentina. Tropical Bryology 24: 33–100.
Matteri, C.M. & Calabrese, G.M. 1999. Taxonomic notes and distribution of Matteria papillosula and
Microdus euchlorus, from northern Patagonia (Argentina), with a new combination in
Campylopodium (Bryopsida). Annales Botanici Fennici 36: 43–49.
Matteri, C.M. & Ochyra, R. 1989. Notes on two southern South American species of Brachytheciaceae
(Musci). Journal of the Hattori Botanical Laboratory 66: 321–330.
Matteri, C.M. & Ochyra, R. 1999. The Meesiaceae (Musci) in southern South America, with notes on the
subdivision of the family. Haussknechtia Beiheft 9: 225–242.
Matteri, C.M. & Piñero, M.R. 1987 [1989]. Sinopsis de las Rhizogoniaceae (Musci) en las regiones
Fueguina y Patagonica. Anales del Instituto de la Patagonia: Serie Ciencias Naturales, Punta
Arenas (Chile) 17: 59–65.
Matteri, C.M. & Schiavone, M.M. 2002. Catálogo de los musgos (Bryophyta) de la Región Fueguina en
Argentina y Chile. Revista del Museo Argentino de Ciencias Naturales, n.s., 4: 111–138.
McFarland, K.D. 1988. Revision of Brachythecium (Musci) for Mexico, Central America, South America,
Antarctica and Circum-subantarctic Islands. xii+537 pp. 58 figs. Ph.D. Dissertation, University of
Tennessee, Knoxville.
Meijer, W. 1951. The genus Orthodontium. xiii + 80 pp. 4 maps. 18 pl. 2 tab. North-Holland Publishing Co.,
Amsterdam.
Mishler, B.D. 1994. Tortula, pp. 319–350, In: Sharp, A.J. et al. (eds.), Moss Flora of Mexico. Memoirs of the
New York Botanical Garden 69: 581–1113.
Mitten, W. 1869. Musci austro-americani Enumeratio omnium Austro-americanorum auctori hucusque
congitorum. Journal of the Linnean Society, Botany 12: 1–659.
Montagne, C. 1850. Flora chilena, plantas cellulaires. I. Musgos. – pp. 5–202, in: Gay, C., Historia fisica y
politica de Chile. Vol. 7, Botanica. Paris.
Müller, C. 1843. Aulacomnion chilense, nov. sp. Botanische Zeitung (Berlin) 1: 649–656.
Müller, C. 1848–1849. Synopsis Muscorum Frondosorum Omnium Hucusque Cognitorum 1: 1–812.
Müller, C. 1885. Bryologia fuegiana. Flora 68 (21–23): 391–429.
Müller, C. 1897. Prodromus bryologiae Argentinicae atque regionum vicinarum III. Hedwigia 36: 84–144.
Müller, C. 1900. Genera Muscorum Frondosorum. Leipzig (E. Kummer), vi + 474 pp.
ISSN 0945-3466
Mosses of Chile 119
Müller, F. 2002. New records and new synonyms for the southern South American moss flora. Nova
Hedwigia 74: 445–450.
Müller, F. 2008. The identity of Pterogoniadelphus hollermayeri (Thér.) Ochyra & Zijlstra, an insufficiently
known Chilean moss. Journal of Bryology 30: 96–97.
Müller, F. 2009. New records and new synonyms for the Chilean moss flora. Tropical Bryology 30: 76–83.
Müller, F. & Pereira, I. 2006. The bryophyte flora of nature reserves in central Chile. 1. The moss flora of
Los Ruiles Nature Reserve, near Talca. Tropical Bryology 27: 55–66.
Müller, F. & Pursell, R. 2003. The genus Fissidens (Musci, Fissidentaceae) in Chile. Journal of the Hattori
Mullen, D. & Frye, T.C. 1947. Middle and South American species of Oligotrichum. The Bryologist 50: 67–
79.
Muñoz, C. & Pisano, C. 1947. Estudio de la vegetación y flora de los Parques Nacionales de Fray Jorge y
Talinai. – Ministerio de Agricultura. Santiago, Chile. Apartado de Agricultura Técnica N°2, 190 pp.
Muñoz, J. 1999. A revision of Grimmia (Musci, Grimmiaceae) in the Americas. 1: Latin America. Annals of
the Missouri Botanical Garden 86: 118–191.
Muñoz, J. & Pando, F. 2000. A world synopsis of the genus Grimmia. Monographs in Systematic Botany
from the Missouri Botanical Garden 83, 133 p.
Newton, M.E. 1974. Notes on Antarctic bryophytes: V. Brachythecium majusculum M. E. Newton, sp. nov.
British Antarctic Survey Bulletin 39: 45–48.
Newton, M.E. 1977. A synoptic flora of South Georgian mosses: VI. Cheilothela, Dicranella, Distichium,
Myurella and Catagonium. British Antarctic Survey Bulletin 46: 1–21.
Nishimura, N. & Kanda, H. 1990. Mittenothamnium deguchi, a new moss species from southern Chile. The
Nyholm, E. 1971. Studies in the genus Atrichum P. Beauv. A short survey of the genus and the species.
Ochi, H. 1968. A revision of the family Funariaceae (Musci) in Japan and the adjacent regions. Japanese
Journal of Botany 20: 1–34.
Ochi, H. 1970. A revision of the subfamily Bryoideae in Australia, Tasmania, New Zealand and the adjacent
islands. Journal of the Faculty of Education, Tottori University, Natural Science 21: 7–67.
Ochi, H. 1977. Central and South American bryaceous mosses, new to science or of geographcial
significance. Journal of the Faculty of Education, Tottori University, Natural Science 27: 33–41.
Ochi, H. 1980. A revision of the Neotropical Bryoideae, Musci (First Part). Journal of the Faculty of
Education, Tottori University, Natural Science 29 (2): 49–154.
Ochi, H. 1981. A revision of the Neotropical Bryoideae, Musci (Second Part). Journal of the Faculty of
Education, Tottori University, Natural Science 30: 21–55.
Ochi, H. 1982. A revision of the Bryoideae (Musci) in southern South America. Journal of the Faculty of
Education, Tottori University, Natural Science 31: 11–47, 17 fig.
Ochi, H. 1990. Bryaceous mosses as the source of distributional records from the Andes region. Bulletin of
the National Science Museum, (Tokyo) Series B, Botany 16(2): 61–71.
Ochi, H. & Mahu, M. 1988. Supplement to the Bryoideae (Bryaceae, Musci) in South America. Hikobia 10:
221–223.
Ochyra, R. 1987a. A revision of the moss genus Sciaromium (Mitt.) I. General remarks and the section
Aloma Dusén. Journal of Bryology 14: 453–464.
Ochyra, R. 1987b. A revision of the moss genus Sciaromium (Mitt.) II. The section Limbidium Dusén, with a
description of Vittia gen. nov. (Vittiaceae, Fam. Nov.). Journal of the Hattori Botanical Laboratory
62: 387–415.
Ochyra, R. 1989. Animadversions on the moss genus Cratoneuron (Sull.) Spruce. Journal of the Hattori
Ochyra, R. 1993a. Antipodal mossses: I. A revision of the genus Holodontium (Seligeriaceae). Fragmenta
Ochyra, R. 1993b. New synonyms of Racomitrium lamprocarpum (Musci, Grimmiaceae). Fragmenta
Ochyra, R. 1994. Anipodal mosses: III. Taxonomic status of the genus Monocranum (Dicranaceae).
Fragmenta Floristica et Geobotanica 39: 453–460.
Ochyra, R. 1998. The moss flora of King George Island, Antarctica. – Cracow, 279 pp.

120 Müller
Ochyra, R. 1999a. Ditrichum austrogeorgicum: a synonym of D. hyalinum (Musci, Ditrichaceae).

Cryptogamie, Bryologique 20: 247–253.
Ochyra, R. 1999b. Antipodal mosses: XI. Additional new synonyms of Anisothecium hookeri (Dicranaceae).
Ochyra, R. 1999c. The taxonomic status of two varieties of Ditrichum (Musci, Ditrichaceae) from southern
South America. Fragmenta Floristica et Geobotanica 44: 504–506.
Ochyra, R. 2004a. New moss records from the Falkland Islands. Journal of Bryology 26: 226–230.
Ochyra, R. 2004b. Antipodal mosses: XV. Taxonomy and distribution of Schistidium cupulare (Bryopsida,
Grimmiaceae). Polish Polar Research 25: 123–133.
Ochyra, R. & Bednarek-Ochyra, H. 1997. The type specimen of Leptotrichum praealtum (Musci,
Ditrichaceae). Fragmenta Floristica et Geobotanica 42: 567–570.
Ochyra, R., Bednarek-Ochyra, H. & Lewis-Smith, R.I. 2002. New and rare moss species from
subantarctic South Georgia. Nova Hedwigia 74: 121–147.
Ochyra, R. & Buck, W.R. 2003. Arctoa fulvella, new to Tierra del Fuego, with notes on Trans-American
bipolar bryogeography. The Bryologist 106: 532–538.
Ochyra, R. & Deguchi, H. 1999. Ditrichum gemmiferum (Musci, Ditrichaceae) in Chile. Fragmenta
Ochyra, R. & Hertel, H. 1990. Contribution to the moss flora of the subantarctic island Marion. Polish
Botanical Studies 1: 19–34.
Ochyra, R. & Lewis-Smith, R.I. 1998. Antarctic species in the genus Ditrichum (Ditrichaceae, Bryopsida),
with a description of D. hyalinum sp. nov. Annales Botanici Fennici 35: 33–53.
Ochyra, R., Lewis-Smith, R.I. & Bednarek-Ochyra, H. 2008. The Illustrated Moss Flora of Antarctica.
Cambridge, Cambridge University Press, 685 pp.
Ochyra, R. & Mahu, M. 1988. Richardsiopsis lacustris (Herzog & Rich.) Ochyra in Chile. Journal of
Bryology 15: 238.
Ochyra, R. & Mahu, M. 1990. Cochlearidium aymaranum, a strange new genus and species of
Cochlearidiaceae (fam. nov.) from Chile. Journal of Bryology 16: 215–222.
Ochyra, R. & Matteri, C.M. 1996. Nomenclatural changes in South American and sub-Antarctic mosses.
Ochyra, R. & Matteri, C.M. 2001. Amblystegiaceae. – In: Flora Criptogamica de Tierra del Fuego 14 (10),
96 pp.
Ochyra, R. & Newton, M.E. 1986. The taxonomy and distribution of Dicranella cardotii (R.Br. ter.) Dixon,
an addition to the moss flora of Antarctica. Lindbergia 11: 94–98.
Ochyra, R. & Zander, R.H. 2007. Is Tortula lithophila conspecific with Sarconeurum glaciale (Bryopsida:
Pottiaceae)? Fragmenta Floristica et Geobotanica Polonica 14: 209–212.
Padberg, M. & Frahm, J.-P. 1985. Monographie der Gattung Atractylocarpus Mitt. (Dicranaceae).
Cryptogamie: Bryologie, Lichénologie 6: 315–341.
Paris, E.G. 1894–1900. Index Bryologicus. 1st ed. Vol. 1–5. Paris.
Paris, E.G. 1903–1906. Index Bryologicus. 2nd ed. Vol. 1–5. Paris.
Pereira, I., Müller, F. & Valderrama, A. 2006. Diversity and distribution of Bryophytes and Lichens of El
Colorado, Central Chile. Nova Hedwigia 83: 117–127.
Perrier, C. 2005. Aperçu de la bryoflore de l'Archipel Juan Fernández, au Chili. Bulletin Mensuel de la
Société Linnéenne de Lyon 74: 19–30.
Podpěra, J. 1973. Bryum Generis Monographiae Prodromus. I. 1. Species Eurasiae Septentrionalis. Pars 16.
Systematica. Subsect. 8. Pseudotriquetra Amann. Academia: Prague. 257 pp.
Pursell, R.A. 1994. Taxonomic notes on Neotropical Fissidens. The Bryologist 97: 253–271.
Quandt, D., Frahm, J.-P. & Frey, W. 2001. Patterns of molecular divergence within the palaeoaustral genus
Weymouthia Broth. (Lembophyllaceae, Bryopsida). Journal of Bryology 23: 305–311.
Rao, P. 2001. Monographic studies on Cryphaea (Bryopsida). Bryobrothera 7, 112 pp.

Reimers, H. 1926. Beiträge zur Bryophytenflora Südamerikas. I. Die von Pater A. Hollermeyer in Chile
1918/1923 gesammelten Leber- und Laubmoose. Hedwigia 66: 27–78.
Reimers, H. 1936. Über die Laubmoosgattungen Bryoporteria Thér., Camptodontium Dus., Hymenoloma
Dus., Verrucidens Card. und ihre systematische Stellung. Notizblatt des Botanischen Gartens und
Museums zu Berlin-Dahlem 13 (116): 36–52.
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Mosses of Chile 121
Robinson, H. 1964. Five Bryophytes of interest from Chile. The Bryologist 67: 53–55.
Robinson, H. 1970. Notes on the moss genera Camptochaete, Physcomitrium and Ptychomnion in Chile.
Phytologia 20: 329–331.
Robinson, H.E. 1972. Observations on the origin and taxonomy of the Antarctic moss flora. Antarctic
Research Series, American Geophysical Union 20: 163–177.
Robinson, H. 1974. Notes on the mosses of Juan Fernandez and southern South America. Phytologia 29:
116–120.
Robinson, H. 1975. The mosses of Juan Fernández Islands. Smithsonian Contributions to Botany 27, 88 pp.
Roivainen, H. 1934. Musci nonnulli ex Chile austrocentrali. Annales Botanici Societatis Zoologicae-
Botanicae Fennicae "Vanamo" 5: 30–36.
Roivainen, H. 1937. Bryological investigations in Tierra del Fuego. 1. Sphagnaceae-Dicranaceae. Annales
Botanici Societatis Zoologicae-Botanicae Fennicae "Vanamo" 9: I–X, 1–58.
Roivainen, H. 1955a. Contribution to the fuegian species of the genus Racomitrium Bridel. Archivum
Societatis Zoologicae Botanicae Fennicae "Vanamo" 9: 85–98.
Roivainen, H. 1955b. Bucklandia bartramii, a new genus and species of leaf moss from Tierra del Fuego.
Archivum Societatis Zoologicae Botanicae Fennicae "Vanamo" 9: 98–100.
Rushing, A.E. 1986. A revision of the genus Bruchia Schwaegr. (Musci). Journal of the Hattori Botanical
Sainsbury, G.O.K. 1955. A handbook of the New Zealand mosses. Bulletin of the Royal Society of New
Zealand 5, 490 pp.
Salmon, E.S. 1903. A monograph of the genus Streptopogon Wils. Annals of Botany (London) 17: 107–150.
Sastre-de Jesus, I. 1987. A revision of the Neckeraceae Schimp. and the Thamnobryaceae Marg. & Dur. in
the Neotropics. xi + 233 pp. Doctoral dissertation. University Microfilms, Ann Arbor.
Schaumann, F. 2005. Terricolous bryophyte vegetation of Chilean temperate rain forests: communities,
adaptive strategies and divergence patterns. Bryophytorum Bibliotheca 62: 1–154 + 14 pp.
unpaginated appendices + 3 pp. index + 3 appendices in pocket.
Schiavone, M.M. 1993. Bryophyta Musci: Polytrichinales. In: Guarrera, S.A., Gamundi de Amos, I. &
Rabinovich de Halperin, D. (eds.), Flora Criptogámica de Tierra del Fuego 14(12): 1–61. Consejo
Nacional de Investigaciones Cientificas y Tecnicas de al Republica Argentina, Buenos Aires.
Schiavone, M.M. & Hyvönen, J. 1991. Distribution maps of mosses from Tierra del Fuego, Argentina. 3.
Schiavone, M.M. & Hyvönen, J. 1993. Distribution maps of mosses from Tierra del Fuego, Argentina. 4.
Schimper, W.P. 1836. Muscorum chilensium species novas descripsit. Annales des Sciences Naturelles,
Botanique, sér. 2, 6: 145–149
Schultze-Motel, W. 1970. Monographie der Laubmoosgattung Andreaea I. Die costaten Arten. Willdenowia
6: 25–110.
Seki, T. 1974. A moss flora of provincia de Aisén, Chile. Results of the second scientific expedition to
Patagonia by Hokkaido and Hiroshima Universities, 1967. Journal of Science of Hiroshima
University, Series B, Division 2 (Botany) 15(1): 9–101.
Seppelt, R.D. 1982. A monographic revision of the genus Ditrichum (Musci: Ditrichaceae). I. Australian and
New Zealand species. Journal of the Hattori Botanical Laboratory 51: 99–150.
Sérgio, C. 1981. Une nouvelle mousse de Madere Thamnobryum fernandesii n. sp. Boletim da Sociedade
Broteriana, Series 2, 53: 1123–1136.
Shaw, J. 1982. Pohlia Hedw. (Musci) in North and Central America and the West Indies. Contributions from
the University of Michigan Herbarium 15: 219–295.
Shaw, J. 1987. Systematic studies on the Bryaceae. Memoirs of the New York Botanical Garden 45: 682–
690.
Shaw, A.J. & Goffinet, B. 2000. Molecular evidence of reticulate evolution in the peatmosses (Sphagnum),
including S. ehyalinum sp. nov. The Bryologist 103: 357–374.
Smith, A.J.E. 2004. The Moss Flora of Britain and Ireland. 2nd ed. Cambridge, 1012 pp.
Smith, G.L. 1969. On Atrichopsis, with notes on some austral Psilopilum species (Polytrichaceae). Bulletin
of the Torrey Botanical Club 96: 60–69.
Smith, G.L. 1971. Conspectus of the genera of Polytrichaceae. Memoirs of the New York Botanical Garden
21(3): 1–83.

122 Müller
Smith Merrill, G.L. 1996. Hebantia, a new genus of Polytrichaceae (Bryophyta). Journal of the Hattori
Sollman, P. 1995 [1996]. Studies on Australian pottiaceous mosses. Lindbergia 20: 144–146.
Stech, M., Pfeiffer, T. & Frey, W. 2002. Chloroplast DNA relationship in palaeoaustral Polytrichadelphus
magellanicus (Hedw.) Mitt. (Polytrichaceae, Bryopsida). Botanische Jahrbücher für Systematik,
Pflanzengeschichte und Pflanzengeographie 124: 217–226
Stone, I.G. 1994. Fissidens section Crenularia C. Muell. in Australia. Journal of Bryology 18: 169–180.
Stone, I.G. & Scott, G.A.M. 1973. Name changes in Australian mosses. Journal of Bryology 7: 603–605.
Streimann, H. 1999. Taxonomic studies on Australian Hookeriaceae (Musci) 2. The genera Distichophyllum
and Bryobrothera. Journal of the Hattori Botanical Laboratory 86: 89–119.
Streimann, H. 2001. Weymouthia cochlearifolia (Schwägr.) Dixon (Lembophyllaceae-Musci) in Australia,
and some notes on W. mollis (Hedw.) Broth. Journal of the Hattori Botanical Laboratory 91: 289–
294.
Suárez, G. & Schiavone, M. 2005. [Abstract] Sobre la filogenia de Pohlia integra (Cardot) Shaw (Bryaceae,
Musci). Boletín de la Sociedad Argentina de Botánica 40 (Suplemento): 41.
Suárez, G. & Schiavone, M. 2008. Pohlia chilensis (Mniaceae), an Afro-American moss. The Bryologist
111: 318–322.
Sullivant, W.S. 1850. Notices of several new species of mosses and Hepaticae from Tierra del Fuego.
Hooker's Journal of Botany and Kew Garden Miscellany 2: 315–318.
Tangney, R.S. & Fife, A.J. 1997. The occurrence of Ptychomnion densifolium in New Zealand. Journal of
Bryology 19: 819–825.
Tangney, R.S. & Fife, A.J. 2003. A review of the genus Fallaciella (Bryopsida: Lembophyllaceae),
including a new species from South Island, New Zealand. Journal of Bryology 25: 121–128.
Thériot, I. 1911. Breutelia (Eubreutelia) subplicata Broth. sp. nov. Revista Chilena de Historia Natural 15:
38–39.
Thériot, I. 1915. Sur quelques mousses recueillies au Chili par le Prof. Carlos E. Porter. Revista Chilena de
Historia Natural 19: 30–37.
Thériot, I. 1917. Contribution a la flore bryologique du Chili. Revista Chilena de Historia Natural 21: 6–37.
Thériot, I. 1918. Contribution a la flore bryologique du Chili. Revista Chilena de Historia Natural 22: 79–
94.
312.
Thériot, I. 1923. Contribution a la flore bryologique du Chili. Revista Chilena de Historia Natural 27: 9–15.
139.
292.
361.
Thériot, I. 1927a. Mousses recoltees dans l’ile Mas a Tierra (Juan Fernández) en 1927, por M. Gualterio
Looser. Revista Chilena de Historia Natural 31: 257.
Thériot, I. 1927b. Le genre Funaria au Chili. Revista Chilena de Historia Natural 31: 30–37.
255.
Thériot, I. 1929a. Le genre Leptodontium au Chili. Revista Chilena de Historia Natural 33: 515–517.
Thériot, I. 1929b. Mousses du Chili austral. Recoltees par le Prof. Thaxter, de Cambridge, Massachusetts.
Revue Bryologique, n. s., 2: 165–166.
262.
Thériot, I. 1933. Bryoporteria nouveau genre de la famille Seligeriaceae. Revista Chilena de Historia
Natural 37: 23–25.
Thériot, I. 1934a [1935]. Quelques mousses du Chili austral recoltees a Freire par Mr. Raul Gillet L., en
Fevrier 1934. Revista Chilena de Historia Natural 38: 83–85.
Thériot, I. 1934b [1935]. Contribution a la flore bryologique du Chili. Revue Bryologique et Lichénologique
7: 167–183.
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Mosses of Chile 123
Thériot, I. 1935 [1936]. Contribution a la flore bryologique du Chili. Revista Chilena de Historia Natural 39:
16–21.
Thériot, I. 1937. Mousses de I’lle-de-Pâques. Revue Bryologique et Lichenologique 10: 74–77.
Touw, A. 2001. A review of the Thuidiaceae (Musci) and a realignment of taxa traditionally accommodated
in Thuidium sensu amplo (Thuidium Schimp., Thuidiopsis (Broth.) M.Fleisch., and Pelekium Mitt.),
including Aequatoriella gen. nov. and Indothuidium gen. nov. Journal of the Hattori Botanical
Vanderpoorten, A., Goffinet, B., Hedenäs, L., Cox, C.J. & Shaw, A.J. 2003. A taxonomic reassessment of
the Vittiaceae (Hypnales, Bryopsida): evidence from phylogenetic analyses of combined chloroplast
and nuclear sequence data. Plant Systematics and Evolution 241: 1–12.
Villagrán, C., Barrera, E., Cuvertino, J. & García, N. 2003. Musgos de la Isla Grande de Chiloé, X region,
Chile: Lista de especies y rasgos fitogeograficos. Boletin del Museo Nacional de Historia Natural,
Chile, 52: 17–44.
Villagran, C., Armesto, J., Hinojosa, L.F, Cuvertino, J., Perez, C. & Medina, C. 2004. El enigmatico
origen del bosque relicto de Fray Jorge. – In: Squeo, F.A., Gutierrez, J.R. & Hernandez, I.R. (eds.),
Historia Natural del Parque Nacional Bosque Fray Jorge. La Serena.
Vitt, D.H. 1976. A monograph of the genus Muelleriella Dusén. Journal of the Hattori Botanical Laboratory
40: 91–113.
Vitt, D.H. 1979. The moss flora of the Auckland Islands, New Zealand, with a consideration of habitats,
origins and adaptations. Canadian Journal of Botany 57: 2226–2263.
Vitt, D.H. 1980. The genus Macrocoma I. Typification of names and taxonomy of the species. The
Vitt, D.H. & Marsh, C. 1988. Population variation and phytogeography of Racomitrium lanuginosum and R.
pruinosum. Nova Hedwigia 90: 235–260.
Walther, K. 1940. Revision des Verwandtschaftskreises von Polytrichum piliferum Schreb. Repertorium
Specierum Novarum Regni Vegetabilis 48 (12–20): 251–255.
Warburg, E.F. 1962. New combinations in British mosses. Transactions of the British Bryological Society 4
(2): 247–248.
Warnstorf, C. 1911. Sphagnales-Sphagnaceae. – In: Engler, A. (ed.): Das Pflanzenreich. Regni vegetabilis
conspectus. Leipzig: Engelmann.
Warnstorf, C. 1916. Pottia-Studien als Vorarbeiten zu einer Monographie des Genus Pottia Ehrh. sens. str.
Hedwigia 58: 35-152.
Wijk, R. van der & Margadant, W.D. 1959. New Combinations in mosses II. Taxon 8: 70–75.
Wijk, R. van der, Margadant, W.D. & Florschuetz, P.A. 1959. Index Muscorum. 1 (A–C). Regnum
Vegetabile 17: i–xxviii, 1–548.
Wijk, R. van der & Margadant, W.D. 1961. New combinations in mosses. VI. Taxon 10: 24–26.
Wijk, R. van der, Margadant, W.D. & Florschuetz, P.A. 1962. Index Muscorum. 2 (D-Hypno). Regnum
Vegetabile 26: 1–535.
Wijk, R. van der, Margadant, W.D. & Florschuetz, P.A. 1964. Index Muscorum. 3 (Hypnum–O). Regnum
Wijk, R. van der, Margadant, W.D. & Florschuetz, P.A. 1967. Index Muscorum 4 (P–S). Regnum
Wijk, R. van der, Margadant, W.D. & Florschuetz, P.A. 1969. Index Muscorum. 5 (T–Z, Appendix).
Regnum Vegetabile 65: i–xii, 1–922.
Williams, R.S. 1915. Mosses from the west coast of South America. Bulletin of the Torrey Botanical Club
42: 393–404.
Williams, R.S. 1924. Orthorrhynchium chilensis sp. nov. The Bryologist 27: 87–88.
Zander, R.H. 1972. Revision of the genus Leptodontium (Musci) in the New World. The Bryologist 75: 213–
280.
Zander, R. H. 1977. The tribe Pleuroweisieae (Pottiaceae, Musci) in Middle America. The Bryologist 80:
233–269.
Zander, R.H. 1978. A synopsis of Bryoerythrophyllum and Morinia (Pottiaceae) in the New World. The

124 Müller
Zander, R.H. 1979. Notes on Barbula and Pseudocrossidium (Bryopsida) in North America and an
annotated key to the taxa. Phytologia 44: 177–214.
Zander, R. H. 1989. Seven new genera in Pottiaceae (Musci) and a lectotype for Syntrichia. Phytologia 65:
424–436.
Zander, R.H. 1993. Genera of the Pottiaceae: mosses of harsh environments. Bulletin of the Buffalo Society
of Natural Sciences 32. vi + 378 pp.
Zander, R.H. & Mahu, M. 1999. A New Species of Hennediella from Chile. The Bryologist 102: 349–351.
Zanten, B.O. van 1971. Musci. In: Zinderen Bakker, E.M. van, Winterbottom, J.M. & Dyer, R.A. (eds),
Marion and Prince Edward Islands. Balkema, Capetown, pp 173–236.
Zanten, B.O. van 1993. Beschouwingen over de identiteit en herkomst van Racopilum chilense.
Buxbaumiella 30: 45–49.
Zanten, B.O. van 2006. A synoptic review of the Racopilaceae (Bryophyta, Musci). 1. Asian, Pacific and
Australasian species of the genus Racopilum. Journal of the Hattori Botanical Laboratory 100:
527–552.
Zomlefer, W. 1993. A revision of Rigodium (Musci: Rigodiaceae). The Bryologist 96: 1–72.
Internet sources:
NY: Online Specimen Cataloges of the herbarium of the New York Botanical Garden, 03 Jun 2009,
<http://www.nybg.org/bsci/herb/>
Tropicos: Tropicos.org. Missouri Botanical Garden. 03 Jun 2009 <http://www.tropicos.org>
Address of the author:

Frank Müller, Technische Universität Dresden, Institut für Botanik, D-01062 Dresden, Germany;
e-mail: Frank.Mueller@tu-dresden.de
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Mosses of Chile 1

An updated checklist of the mosses of Chile

ARCHIVE FOR BRYOLOGY 58 (2009)

The arrangement of the checklist

Table 1. Number of accepted taxa reported for political regions of Chile

ARCHIVE FOR BRYOLOGY 58 (2009)

Fig. 1: Political regions of Chile (from He 1998, modified).

Taxa reported from Chile arranged alphabetically by genera and species

Acaulon uleanum Müll. Hal.

ARCHIVE FOR BRYOLOGY 58 (2009)

ARCHIVE FOR BRYOLOGY 58 (2009)

Donat 1936; Herzog 1954. Dist. XI. Aisén.

ARCHIVE FOR BRYOLOGY 58 (2009)

(as Bryum prostratum, Ochi 1980). Dist. V. Valparaíso.

ARCHIVE FOR BRYOLOGY 58 (2009)

Ireland et al. 2007. Dist. VIII. Ñuble.

ARCHIVE FOR BRYOLOGY 58 (2009)

ARCHIVE FOR BRYOLOGY 58 (2009)

ARCHIVE FOR BRYOLOGY 58 (2009)

Bryum alpinum With.

ARCHIVE FOR BRYOLOGY 58 (2009)

Bryum elegantulum Lorentz

Bryum microimbricatum Ochi

ARCHIVE FOR BRYOLOGY 58 (2009)

Bryum pseudothyridium E.B. Bartram = Bryum perlimbatum fide Ochi 1982.

ARCHIVE FOR BRYOLOGY 58 (2009)

1974; as Racomitrium sublamprocarpum, Dusén 1903b; as Racomitrium subnigritum, Dusén

ARCHIVE FOR BRYOLOGY 58 (2009)

Campylopodium medium (Duby) Giese & J.-P. Frahm

Ireland et al. 2007. Dist. VIII. Arauco.

ARCHIVE FOR BRYOLOGY 58 (2009)

Campylopus pilifer Brid.

Ireland et al. 2007. Dist. VIII. Arauco, Concepción, Ñuble.

ARCHIVE FOR BRYOLOGY 58 (2009)

Ceratodon purpureus subsp. convolutus (Reichardt) Burley

ARCHIVE FOR BRYOLOGY 58 (2009)

Hygroamblystegium fuegianum var. excurrens, Bartram 1946; as Amblystegium excurrens,

ARCHIVE FOR BRYOLOGY 58 (2009)

Daltonia valdiviae Herzog = Daltonia trachyodonta fide Bartram 1931.

Dichodontium paludella Besch. ≡ Neomeesia paludella.

ARCHIVE FOR BRYOLOGY 58 (2009)

Dicranoloma billardierei (Brid.) Paris var. billardierei

Dicranoloma kunkelii H. Rob.

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Didymodon acutus (Brid.) K. Saito

ARCHIVE FOR BRYOLOGY 58 (2009)

Distichophyllum fasciculatum Mitt.

ARCHIVE FOR BRYOLOGY 58 (2009)

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Drummondia obtusifolia Müll. Hal.

He 1998. Dist. IV. Limarí.

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ARCHIVE FOR BRYOLOGY 58 (2009)

Funaria berteroana Hampe = Entosthodon laevis fide Fife 1987.

ARCHIVE FOR BRYOLOGY 58 (2009)

Grimmia macropulvinata Dusén = Grimmia trichophylla fide Deguchi 1984.

ARCHIVE FOR BRYOLOGY 58 (2009)

ARCHIVE FOR BRYOLOGY 58 (2009)

Hymenocleiston magellanicum var. edenense Besch. ≡ Tayloria dubyi var. edenensis.

Hypnum chrysogaster Müll. Hal.

ARCHIVE FOR BRYOLOGY 58 (2009)

Hypopterygium didictyon Müll. Hal.

ARCHIVE FOR BRYOLOGY 58 (2009)

ARCHIVE FOR BRYOLOGY 58 (2009)