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Ver. 19.2
April 17th, 2024

043.0000 Family Cystopteridaceae (Payer) Shmakov; Turczaninowia 4: 60 (2001)
3-4 genera, ca. 35 spp. Family status confirmed by Rothfels et al. (2012)
Species listed in database: 35; hybrids: 11
= Cystopteridinae Payer; Bot. Crypt.: 198 (1850) (Cystopterideae)
= Cystopterioideae Ching & Z.R.Wang; Acta Phytotax. Sin. 35: 34 (1997)
043.0003 Genus Gymnocarpium Newman; Phytol. 4: 371 (1851)
Distribution: 5 spp. north-temperate, 1 sp. Taiwan, 1 Himalaya to New Guinea
Species listed in database: 9; hybrids: 5
= Carpogymnia D.Löve & Á.Löve; Univ. Color. Stud. Biol. 24: 8 (1966)
= Currania Copel.; Philipp. J. Sci., Bot. 4: 112 (1909)
species :
Gymnocarpium appalachianum Pryer & Haufler; Syst. Bot. 18: 161 (1993)
Name: Appalachian oakfern
Distribution: USA (Maryland, North Carolina, Ohio, Pennsylvania, Virginia, West Virginia)
75 USA 78 USA
Conservation Status: IUCN R
species :
Gymnocarpium continentale (Petrov) Pojark.; Soobshch. Tadzh. Fil. Akad. Nauk SSSR. 22: 10 (1950)
Name: Nahanni Oak Fern
Distribution: Finland; Siberia (Altai, Buryatia, Chita, Irkutsk, Krasnoyarsk, W-Siberia, Yakutia); Alaska; Canada (Alberta, British Columbia, Manitoba, New Brunswick, Northern Territories, Nunavut, Ontario, Québec, Saskatchewan, Yukon); USA (Iowa, Massachusetts, Michigan, Minnesota, Vermont, Wisconsin)
10 FIN 30 SIB 70 ASK CDN 71 CDN 72 CDN 74 USA 75 USA
Remarks: frequently included as subspec. parvulum into G. jessoense
Synonyms: hidden
= Carpogymnia continentalis (Petrov) Á.Löve & D.Löve; Taxon 16: 191 (1967)
= Dryopteris pulchella var. continentalis Petrov; Fl. Iakut. 1: 14 (1930)
= Dryopteris continentalis Petrov.; Fl. Iakut. 1: 15 (1930)
= Gymnocarpium jessoense subsp. parvulum Sarvela; Ann. Bot. Fenn. 15: 103 (1978)
species :
Gymnocarpium disjunctum (Rupr.) Ching; Acta Phytotax. Sin. 10: 304 (1965)
Name: Pacific oakfern
Distribution: Russian Far East (Magadan); Alaska; Canada (Alberta, British Columbia); USA (Colorado, Idaho, Montana, Oregon, Washington State, Wyoming)
31 RFE 70 ASK 71 CDN 73 USA
Remarks: frequently included in G. jessoense
Synonyms: hidden
= Carpogymnia disjuncta (Rupr.) Á.Löve & D.Löve; Taxon 16: 191 (1967)
= Dryopteris disjuncta (Rupr. ex Schur) C.V.Morton; Rhodora 43: 217 (1941)
= Dryopteris disjuncta f. erecta (G.Lawson) Roland; Proc. Nova Scotian Inst. Sci. 20: 92 (1941)
= Dryopteris linnaeana var. disjuncta (Rupr.) Fomin; Fl. Sibir. Orient. Extr. 5: 79 (1930)
= Gymnocarpium disjunctum (Rupr.) Á.Löve & D.Löve; Bot. Notis. 128(4): 497 (1975 publ. 1976), isonym
= Gymnocarpium dryopteris subsp. disjunctum (Rupr.) Sarvela; Ann. Bot. Fenn. 15(2): 103 (1978)
= Gymnocarpium dryopteris var. disjunctum (Rupr.) Ching; Contr. Biol. Lab. Sci. Soc. China 9(1): 41 (1933)
= Phegopteris dryopteris f. erecta (G.Lawson) M.Broun; Index No. Amer. Ferns: 134 (1938)
= Phegopteris dryopteris subsp. disjunctum (Rupr.) C.V.Morton
= Phegopteris dryopteris var. disjuncta (Rupr.) Trel.; Harriman Alaska Exped. 5: 382 (1904)
= Polypodium disjunctum (Rupr.) Schur; Oesterr. Bot. Z. 8: 193 (1858)
= Polypodium disjunctum var. erectum G.Lawson; Edinburgh New Philos. J. 19: 109 (1864)
= Polypodium dryopteris var. disjunctum Rupr.; Distr. Crypt. Vasc. Ross. 52 (1845)
species :
Gymnocarpium dryopteris (L.) Newman; Phytol. 4: app. 24 (1851)
Name: Eichenfarn (DE), Western oakfern (EN)
Distribution: Iceland; Faeroe Isl.; England; Scotland; Wales; Ireland; Denmark; Norway; Sweden; Finland; Netherlands; Belgium; Luxembourg; Germany (Brandenburg, Berlin, Baden-Württemberg, Bayern, Hessen, Hamburg, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; Spain; Andorra; France; Corsica; Italy; Malta; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Romania; Bulgaria; Greece (mountains in N-Greece); Estonia; Latvia; Lithuania; Belarus; C-European Russia; E-European Russia; N-European Russia; W-European Russia; Moldova; Ukraine; Crimea; Siberia (Altai, Buryatia, Chita, Irkutsk, Krasnoyarsk, Tuva, W-Siberia); Russian Far East (Amur, Kamchatka, Khabarovsk, Kuril Isl., Magadan, Primorye, Sakhalin); Kazakhstan; Northern Caucasus; Georgia [Caucasus]; Turkey (Anatolia); China (Heilongjiang, Jilin, Liaoning, Nei Mongol, Shaanxi, Shanxi, Xinjiang); Mongolia; North Korea; South Korea; Japan; Alaska; Greenland; Canada (Alberta, British Columbia, Labrador, Manitoba, New Brunswick, Newfoundland, Nova Scotia, Northern Territories, Nunavut, Ontario, Prince Edward Isl., Québec, Saskatchewan, Yukon); St. Pierre et Miquelon; USA (Arizona, Colorado, Connecticut, Iowa, Idaho, Illinois, Massachusetts, Maryland, Maine, Michigan, Minnesota, Montana, North Dakota, New Hampshire, New Jersey, New Mexico, New York, Ohio, Oregon, Pennsylvania, Rhode Island, South Dakota, Utah, Vermont, Washington State, Wisconsin, West Virginia, Wyoming)
10 DEN FIN FOR GRB ICE IRE NOR SWE 11 AUT BGM CZE GER HUN LIE LUX NET POL SLO SWI 12 ADO COR FRA SPA 13 ALB BIH BUL CRO GRC ITA KOS MLT MNT NMA ROM SRB 14 BLR EST KRY LAT LIT MLD RUS UKR 30 SIB 31 RFE 32 KAZ 33 GEO NCS 34 TUR 36 CHI 37 MON 38 JAP KOR NKO 70 ASK CDN GNL 71 CDN 72 CDN SPM 73 USA 74 USA 75 USA 76 USA 77 USA 78 USA
Synonyms: hidden
= Aspidium dryopteris (L.) Baumg.; Enum. stirp. Transs. 4: 29 (1846)
= Carpogymnia dryopteris (L.) Á.Löve & D.Löve; Univ. Colorado Stud., Ser. Biol. 24: 8 (1966)
= Currania dryopteris (L.) Wherry; Bartonia 21: 15: 143 (1942)
= Dryopteris dryopteris (L.) Britton; Britton & Brown, Ill. Fl. N. U.S., ed. 2, 1: 23 (1913), nom. inval.
= Dryopteris linnaeana C.Chr.; Index Filic. 275 (1905), nom. nov. for Polypodium dryopteris L.
= Dryopteris pulchella (Salisb.) Hayek; Fl. Steiermark 39 (1908)
= Dryopteris pumila V.Krecz; Fl. Kavkaza, ed. 2, 1: 19 (1939)
= Dryopteris triangularis Herter; Bull. Herb. Boissier, sér. 2, 8: 797 (1908)
= Filix pumila Gilib.; Exerc. Phyt. 2: 558 (1792) (nom. inval.)
= Gymnocarpium dryopteris (L.) Ching; Contr. Biol. Lab. Sci. Soc. China 9: 38 (1933), isonym
= Lastrea dryopteris (L.) Bory; Dict. Class d´Hist. Nat. 9: 233 (1826)
= Nephrodium dryopteris (L.) Michx.; Fl. bor. Amer. 2: 270 (1803)
= Phegopteris dryopteris (L.) Bedd.; Handb. Ferns Brit. India 293 (1883), isonym
= Phegopteris dryopteris (L.) Fée; Mém. Foug. 5 (Gen. Filic.): 243 (1852)
= Phegopteris dryopteris f. interrupta Jewell; Fern Bull. 16: 86 (1908)
= Phegopteris dryopteris var. minor Farw.; Amer. Midl. Naturalist 12: 292 (1931)
= Phegopteris triangularis St.-Lag.; Flore Rhone ed. VIII: 964 (1889)
= Polypodium conjunctum Schur; Enum. Pl. Transsilv.: 831 (1866)
= Polypodium dryopteris L.; Sp. Pl. 2: 1093 (1753)
= Polypodium dryopteris var. glabrum Neilr.; Fl. Wien, Nachtr.: 64 (1851)
= Polypodium dryopteris var. glandulosum Neilr.; Fl. Wien, Nachtr.: 64 (1851)
= Polypodium dryopteris var. rigidius Hook.; Fl. Bor.-Amer. (Hooker) 2: 259 (1840)
= Polypodium pulchellum Salisb.; Prodr. Stirp. Chap. Allerton 403 (1796)
= Polypodium triangulare Dulac; Fl. Dépt. Hautes-Pyrenees 31 (1867) non L. 1774
= Polystichum dryopteris (L.) Roth; Tent. Fl. Germ. 3: 80 (1799), Roem., Arch. 2(1): 106 (1799)
= Thelypteris dryopteris (L.) Sloss.; Rydb., Fl. Rocky Mts. 1043 (1917)
species :
Gymnocarpium fedtschenkoanum Pojark.; Soobshch. Tadzh. Fil. Akad. Nauk SSSR. 22: 9 (1950)
Distribution: Tajikistan; Afghanistan; Pakistan (Chitral, Swat, Kurram, Hazara, Gilgit, Baltistan); Nepal; India (Jammu & Kashmir, Himachal Pradesh, Uttarakhand)
32 TZK 34 AFG 40 IND NEP PAK
species :
Gymnocarpium jessoense (Koidz.) Koidz.; Acta Phytotax. Geobot. 5: 40 (1936)
Distribution: Siberia (Altai, Buryatia, Chita, Irkutsk, Krasnoyarsk, Tuva, W-Siberia, Yakutia); Russian Far East (Amur, Kamchatka, Khabarovsk, Magadan, Primorye, Sakhalin); China (Gansu, Guizhou, Hebei, Heilongjiang, Henan, Jilin, Liaoning, Nei Mongol, Ningxia, Qinghai, Shaanxi, Shanxi, Sichuan, Xinjiang, NW-Yunnan); Tibet; Mongolia; North Korea; South Korea; Japan; Pakistan; Nepal; Bhutan; India (Sikkim, Himachal Pradesh, Uttarakhand)
30 SIB 31 RFE 36 CHI TIB 37 MON 38 JAP KOR NKO 40 BHU IND NEP PAK
Synonyms: hidden
= Aspidium dryopteris var. longulum Christ; Bull. Herb. Boissier, sér. 2, 2: 830 (1902)
= Carpogymnia jessoensis (Koidz.) Á.Löve & D.Löve; Univ. Colorado Stud., Biol. 24: 8 (1967)
= Dryopteris jessoensis Koidz.; Bot. Mag. (Tokyo) 38: 104 (1924)
= Dryopteris linnaeana var. jessoensis (Koidz.) C.Chr.; Index Filic., Suppl. 3: 89 (1934)
= Gymnocarpium longulum (Christ) Kitag.; Rep. Inst. Sci. Res. Manchoukuo 3(App. 1): 33 (1939)
= Gymnocarpium robertianum subsp. longulum (Christ) Toyok.; Mem. Natl. Sci. Mus. (Tokyo) 5: 189 (1972)
= Gymnocarpium robertianum var. longulum (Christ) H.Itô ex Nakai; Rep. Exped. Manchoukuo Sect. IV 4: 4 (1936)
= Gymnocarpium tenuipes Pojark. & Shmakov; Fl. Rastitel´nost´ Altaya: 48 (1995)
= Lastrea jessoensis (Koidz.) Akasawa; Bull. Kochi Womans Univ. 8: 31 (1960)
= Lastrea robertiana var. longula (Christ) Ohwi; Fl. Jap. Pterid. 101 (1957)
species :
Gymnocarpium oyamense (Baker) Ching; Contr. Biol. Lab. Sci. Soc. China 9: 40 (1933)
Distribution: China (Anhui, Chongqing, Gansu, Guizhou, Henan, Hubei, Hunan, Jiangxi, Shaanxi, Sichuan, Yunnan, Zhejiang); Tibet; Japan; Taiwan; Nepal; India (Sikkim, Arunachal Pradesh); Philippines (Luzon); Moluccas (Seram); New Guinea (Irian Jaya, Papua New Guinea)
36 CHI TIB 38 JAP TAI 40 IND NEP 42 MOL NWG PHI
Remarks: Subgen. Currania
Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Aspidium gymnogrammoides (Baker) Christ; Bull. Acad. Int. Géogr. Bot., sér. 3, 16: 120 (1906)
= Aspidium krameri (Franch. & Sav.) Christ; Farrnkr. d. Schweiz 153 (1900), as ´crameri´, et Bull. Soc. Bot. France 62, Mem. 1: 37 (1905)
= Carpogymnia oyamensis (Baker) Á.Löve & D.Löve; Taxon 16: 191 (1967)
= Currania gracilipes Copel.; Philipp. J. Sci., Bot. 4: 112 (1909)
= Currania oyamensis (Baker) Copel.; Philipp. J. Sci., Bot. 6: 147 (1911)
= Currania oyamensis var. gracilipes (Copel.) Tagawa; Acta Phytotax. Geobot. 14(3): 95 (1951)
= Currania oyamensis var. incisa (Franch. & Sav.) Koidz.; Fl. Symb. Or.-Asiat. 38 (1930)
= Dryopteris genuflexa Rosenst.; Repert. Spec. Nov. Regni Veg. 12: 176 (1913)
= Dryopteris gracilipes (Copel.) C.Chr.; Index Filic., Suppl. Tert.: 87 (1934)
= Dryopteris gymnogrammoides (Baker) C.Chr.; Index Filic. 269 (1905)
= Dryopteris oyamensis (Baker) C.Chr.; Index Filic. 282 (1905)
= Gymnocarpium gracilipes (Copel.) Ching; Contr. Biol. Lab. Sci. Soc. China 9: 39 (1933)
= Gymnocarpium oyamense var. gracilipes (Copel.) W.C.Shieh; J. Sci. Engin. (Natl. Chung-Hsing Univ.) 13: 50 (1976)
= Gymnocarpium oyamense var. kusamae H.Itô; J. Jap. Bot. 63(1): 18 (1988)
= Nephrodium gymnogrammoides (Baker) Diels; Engl. & Prantl, Nat. Pflanzenfam. 1(4): 170 (1899)
= Nephrodium krameri (Franch. & Sav.) Diels; Engl. & Prantl, Nat. Pflanzenfam. 1 [4]. 170 (1899)
= Phegopteris gymnogrammoides (Baker) Alderw.; Malayan Ferns Fern Allies, Suppl. 312 (1917)
= Phegopteris krameri (Franch. & Sav.) Makino; Phan. & Pter. Jap. Ic. t. 30. 1899 (1899)
= Phegopteris oyamensis (Baker) Alderw.; Malayan Ferns Fern Allies, Suppl. 312 (1917)
= Polypodium gymnogrammoides Baker; J. Bot. 26: 229 (1888)
= Polypodium krameri Franch. & Sav.; Enum. Pl. Jap. 2: 244 (1876) (kremeri), nomen
= Polypodium krameri var. incisum Franch. & Sav.; Enum. Pl. Jap. 2: 244 (1877)
= Polypodium oyamense Baker; J. Bot. 15: 366 (1877)
species :
Gymnocarpium remotepinnatum (Hayata) Ching; Bull. Chin. Bot. Soc. 1(2): 14 (1935), et Icon. Fil. Sin. 4: t. 172 (1937)
Distribution: Russian Far East (Sakhalin); China (NW-Yunnan); Taiwan
31 RFE 36 CHI 38 TAI
Synonyms: hidden
= Dryopteris remota Hayata; J. Coll. Sci. Imp. Univ. Tokyo 80: 421 (1911) (nomen malum, non Hayek (1908))
= Dryopteris remotipinnata Hayata; Gen. Ind. Fl. Taiwan 108 (1917)
= Gymnocarpium remotum Ching; Contr. Biol. Lab. Sci. Soc. China, Bot. Ser. 9: 41 (1933), nom. superfl.
= Thelypteris remotepinnata (Hayata) Alston; Koie & Reching., Biol. Skrift. Danske Vid. Selsk. 10(3): 10 (1959)
species :
Gymnocarpium robertianum (Hoffm.) Newman; Phytol. 4: 371 (1851)
Name: Ruprechtsfarn (DE), Scented oakfern (EN)
Distribution: England; Scotland; Wales; Ireland; Norway; Sweden; Finland; Netherlands; Belgium; Luxembourg; Germany (Brandenburg, Berlin, Baden-Württemberg, Bayern, Hessen, Mecklenburg-Vorpommern [I], Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; Spain; Andorra; France; Corsica; Italy; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Romania; Bulgaria; Greece (mountains in NW-Greece, N-Peloponnisos); Estonia; Latvia; ?Belarus; C-European Russia; N-European Russia; W-European Russia; Moldova; Ukraine; Crimea; Algeria; Siberia (Altai, Krasnoyarsk); Kazakhstan; Northern Caucasus; Georgia [Caucasus]; China (Qinghai, Xinjiang); Japan; Alaska; Canada (Manitoba, New Brunswick, Newfoundland, Ontario, Québec); USA (Iowa, Illinois, Michigan, Minnesota, Wisconsin)
10 FIN GRB IRE NOR SWE 11 AUT BGM CZE GER HUN LIE LUX NET POL SLO SWI 12 ADO COR FRA SPA 13 ALB BIH BUL CRO GRC ITA KOS MNT NMA ROM SRB 14 EST KRY LAT MLD RUS UKR 20 ALG 30 SIB 32 KAZ 33 GEO NCS 36 CHI 38 JAP 70 ASK 71 CDN 72 CDN 74 USA 75 USA
Remarks: G. altaicum is kept separate by Flora of China
Synonyms: hidden
= Aspidium calcareum (Sm.) Baumg.; Enum. stirp. Transs. 4: 29 (1846)
= Aspidium robertianum (Hoffm.) Luerss.; Asch. & Graebn., Syn. Mitteleur. Fl. 1: 22 (1896)
= Carpogymnia dryopteris var. pumila (DC.) B.Boivin; Phytologia 15: 148 (1967)
= Carpogymnia robertiana (Hoffm.) Á.Löve & D.Löve; Taxon 16: 191 (1967)
= Currania robertiana (Hoffm.) Wherry; Bartonia 21: 15 (1942)
= Dryopteris disjuncta subsp. calcarea (Sm.) Rouy; Rouy & Foucaud, Fl. France 14: 399 (1913)
= Dryopteris obtusifolia (Schrank) O.Schwarz; Mitt. Thüring. Bot. Ges. 1: 84 (1949), nom. inval., non Rosenst. 1912
= Dryopteris robertiana (Hoffm.) C.Chr.; Index Filic. 289 (1905)
= Gymnocarpium altaicum Chang Y.Yang; Fl. Xinjiangensis 1: 304 (1992 publ. 1993), as ´altaycum´
= Gymnocarpium dryopteris var. pumilum (DC.) B.Boivin; Bull. Soc. Bot. France 109: 127 (1962)
= Gymnocarpium obtusifolium (Schrank) O.Schwarz; Mitt. Thüring. Bot. Ges. 1: 84 (1949)
= Gymnocarpium robertianum (Hoffm.) Ching; Contr. Biol. Lab. Sci. Soc. China 9: 42 (1933), isonym
= Lastrea calcarea (Sm.) Bory; Dict. Class d´Hist. Nat. 9: 233 (1826)
= Lastrea dryopteris var. robertiana (Hoffm.) Lawalrée; Bull. Jard. Bot. État Bruxelles 19: 238 (1949)
= Lastrea obtusifolia (Schrank) Janch.; Phyton (Horn) 2: 63 (1950)
= Lastrea robertiana (Hoffm.) Newman; Natur. Alm. 17 (1844)
= Nephrodium robertianum (Hoffm.) Prantl; Exkurs.-Fl. Bayern: 24 (1884)
= Phegopteris calcarea (Sm.) Fée; Mém. Foug. 5 (Gen. Filic.): 243 (1852)
= Phegopteris dryopteris subsp. calcarea (Sm.) Bonnier & Layens; Tabl. Syn. Pl. Vasc. France: 378 (1894)
= Phegopteris dryopteris subsp. robertiana (Hoffm.) Hensl.; Stud. Cat. Brit. Pl.: 44 (1879)
= Phegopteris dryopteris var. robertiana (Hoffm.) Davenp.; Cat. Davenp. Herb. Suppl. 47 (1883)
= Phegopteris robertiana (Hoffm.) A.Braun; Asch., Fl. Brandenb. 2: 198 (1859), isonym
= Phegopteris robertiana (Hoffm.) Fée; Mém. Foug. 5. (Gen. Filic.) 243 (1852)
= Polypodium calcareum Sm.; Brit. Fl. 1117 (1804)
= Polypodium dryopteris subsp. calcareum (Sm.) Berher; 257 (1887)
= Polypodium dryopteris subsp. robertianum (Hoffm.) Corb.; Nouv. Fl. Normandie: 678 (1894)
= Polypodium dryopteris subsp. robertianum (Hoffm.) Hartm.; Svensk och Norsk Excurs.-Fl. 151 (1846)
= Polypodium dryopteris var. calcareum (Sm.) Rupr.; Distr. Crypt. Vasc. Ross.: 52 (1845), isonym
= Polypodium dryopteris var. calcareum (Sm.) Hampe; 62 (1836), isonym
= Polypodium dryopteris var. calcareum Wahlenb.; Fl. Suecica 2: 669 (1826)
= Polypodium dryopteris var. pumilum DC.; Lam. & A. DC., Fl. Franç., éd. 3, 6: 565 (1815)
= Polypodium dryopteris var. robertianum (Hoffm.) Ledeb.; Fl. Ross. (Ledeb.) 4: 509 (1853)
= Polypodium obtusifolium Schrank; Schrank & Moll, Naturhist. Briefe Oestreich 2: 296 (1785)
= Polypodium phegopteris var. calcareum (Sm.) Lej.; Rev. Fl. Spa: 210 (1825)
= Polypodium phegopteris var. robertianum (Hoffm.) Lej.; Rev. Fl. Spa: 210 (1825)
= Polypodium robertianum Hoffm.; Deutschl. Fl. 2: add. & emend. (1796)
= Thelypteris robertiana (Hoffm.) Sloss.; Rydb., Fl. Rocky Mts. 1044 (1917)
species :
Gymnocarpium ×achriosporum Sarvela; Ann. Bot. Fenn. 15(2): 104 (1978)
Distribution: Sweden; Canada (Québec); etc.
10 SWE 72 CDN
Remarks: Gymnocarpium dryopteris x Gymnocarpium robertianum
species :
Gymnocarpium ×bipinnatifidum Miyam.; J. Jap. Bot. 63(1): 19 (1988)
Distribution: Japan
38 JAP
Remarks: Gymnocarpium oyamense x robertianum
species :
Gymnocarpium ×brittonianum (Sarvela) Pryer & Haufler; Syst. Bot. 18(1): 168 (1993), pro sp.
Distribution: Alaska; Canada (Ontario, Saskatchewan); USA (Colorado, Connecticut, Iowa, Idaho, Illinois, Massachusetts, Maine, Michigan, Minnesota, Montana, New Hampshire, New York, Ohio, Pennsylvania, Rhode Island, Virginia, Vermont, Washington State, Wisconsin, West Virginia, Wyoming)
70 ASK 71 CDN 72 CDN 73 USA 74 USA 75 USA 78 USA
Synonyms: hidden
= Gymnocarpium dryopteris nothosubsp. brittonianum Sarvela; Ann. Bot. Fenn. 17(3). 292 (1980)
= Gymnocarpium brittonianum (Sarvela) Sarvela; Syst. Bot. 18: 168-169 (1993)
species :
Gymnocarpium ×heterosporum V.H.Wagner; Rhodora 68: 132, f. 2-7 (1966)
Name: Blackspore oakfern
Distribution: C-European Russia; N-European Russia; Siberia (Yakutia); Northern Caucasus; USA (Pennsylvania)
14 RUS 30 SIB 33 NCS 75 USA
Remarks: Gymnocarpium continentale x Gymnocarpium robertianum
Synonyms: hidden
= Dryopteris heterospora (W.H.Wagner) Kuvaev; Bot. Zhurn. (Moscow & Leningrad) 66(7): 944 (1981)
species :
Gymnocarpium ×intermedium Sarvela; Ann. Bot. Fenn. 15(2): 105 (1978)
Distribution: Finland; European Russia; Alaska; Canada (Alberta, Manitoba, Northern Territories, Ontario, Québec, Saskatchewan, Yukon); USA (Iowa, Michigan, Minnesota, Wisconsin)
10 FIN 14 RUS 70 ASK CDN 71 CDN 72 CDN 74 USA 75 USA
Remarks: Gymnocarpium continentale x Gymnocarpium dryopteris
Synonyms: hidden
= Dryopteris disjuncta f. glandulosa (R.M.Tryon) Cody; Amer. Fern J. 45: 126 (1955)
= Gymnocarpium dryopteris f. glandulosa (R.M.Tryon) Soper; Amer. Fern J. 53: 77 (1963)
= Dryopteris linnaeana f. glandulosa R.M.Tryon; Amer. Fern J. 29(1): 4 (1939)
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A complete, synonymic checklist of the Higher Plants of the World

Version 19.2 (April 17th, 2024)

Covering 354,696 accepted species,
8,644 hybrids, 47,758 accepted infraspecific taxa and 996,379 synonyms,
in total 1,407,477* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

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