051.0000
Family
Aspleniaceae Newman; Hist. Brit. Ferns 6 (1840)
2(-6) genera, ca. 750 spp.
Species listed in database: 835; hybrids: 160
= Asplenieae Gaudich.; Freycinet, Voy. Uranie, Bot.: 311 (1828)
= Aspleniinae Dumort.; Anal. Fam. Pl.: 67 (1829)
= Asplenioideae Link; Fil. Sp.: 73 (1841)
051.0002
Genus
Hymenasplenium Hayata; Bot. Mag. (Tokyo) 41: 712 (1927)
Distribution: Mainly tropical Asia and China, few tropical America, 1 tropical Africa, few Pacific. Separation from Asplenium supported by Ohlsen et al. (2014).
Species listed in database: 67; hybrids: 3
species :
Hymenasplenium amoenum
(C.Presl ex Mett.) Ranil; Phytotaxa 511(2): 183 (2021)
Distribution: China (SE-Yunnan); Taiwan; Nepal; Bhutan; India (Arunachal Pradesh, Assam State, Himachal Pradesh, Manipur, Sikkim, Tripura, Uttarakhand, West Bengal, Nagaland, Jammu & Kashmir, Darjeeling); Sri Lanka; Myanmar; Vietnam
36 CHI 38 TAI 40 BHU IND NEP SRL 41 MYA VIE
Remarks: bona sp. separate from H. obliquissimum fide Fraser-Jenkins et al. (2017)
Synonyms: hidden
= Asplenium amoenum C.Presl; Tent. Pterid. 107 (1836), nomen
= Asplenium amoenum C.Presl ex Mett.; Abh. Senckenberg. Naturf. Ges. 3(1): 131 (1859)
= Asplenium resectum var. amoenum (C.Presl ex Mett.) F.M.Bailey; Bull. Bot. Dept. Agric. Queensland 17: 9 (1913)
= Asplenium retusulum Ching; Acta Phytotax. Sin. 10: 185 (1965) [fide Fraser-Jenkins et al. (2017)]
= Asplenium unilaterale var. amoenum (C.Presl ex Mett.) Domin; Biblioth. Bot. 20: 95. (1913)
= Asplenium unilaterale var. nudum Atk. ex Bedd.; Handb. Ferns Brit. India 153 (1883)
= Asplenium unilaterale var. nudum Atk. ex C.B.Clarke; Trans. Linn. Soc. 2, Bot. 1: 481 (1880)
= Asplenium wuliangshanense Ching; Acta Phytotax. Sin. 10: 186 (1965) [fide Fraser-Jenkins et al. (2017)]
= Hymenasplenium retusulum (Ching) Viane & S.Y.Dong; Fl. China 2-3: 310 (2013) [fide Fraser-Jenkins et al. (2017)]
= Hymenasplenium wuliangshanense (Ching) Viane & S.Y.Dong; Fl. China 2-3: 310 (2013) [fide Fraser-Jenkins et al. (2017)]
species :
Hymenasplenium andohahelense
Viane; Aspleniaceae Madagascar 12 (2021)
Distribution: Madagascar
29 MDG
species :
Hymenasplenium antsirananense
Viane; Aspleniaceae Madagascar 13 (2021)
Distribution: Madagascar
29 MDG
species :
Hymenasplenium apogamum
(N.Murak. & Hatan.) Nakaike; New Fl. Japan, Pterid. 841 (1992)
Distribution: China (Yunnan, Hongkong); Ryukyu Isl.; Taiwan; C-Nepal (one collection only); India (Assam State, ?Mizoram); Myanmar; Thailand (N-Thailand, SW-Thailand, C-Thailand, SE-Thailand, peninsular Thailand); Laos; Vietnam
36 CHI 38 NNS TAI 40 IND NEP 41 LAO MYA THA VIE
Conservation Status: EN in India, CR in Nepal
Synonyms: hidden
= Asplenium apogamum N.Murak. & Hatan.; J. Fac. Sci. Univ. Tokyo, Sect. 3, Bot. 14(3): 193 (1988)
= Asplenium resectum var. decurrens Bedd.; Suppl. Ferns South. India 10, pl. 357 (1876)
= Asplenium unilaterale var. decurrens (Bedd.) H.S.Kung; Fl. Sichuanica 6: 357 (1988)
= Asplenium unilaterale var. udum Bedd.; Handb. Ferns Brit. India 153 (1883), nom. superfl.
species :
Hymenasplenium basiscopicum
(R.C.Moran & Sundue) L.Regalado & Prada; Amer. Fern J. 101(4): 280 (2012)
Distribution: Bolivia (Beni, La Paz, Santa Cruz)
83 BOL
Remarks: to be expected in S-Peru
Synonyms: hidden
= Asplenium basiscopicum R.C.Moran & Sundue; Brittonia 56(2): 124 (2004)
species :
Hymenasplenium cardiophyllum
(Hance) Nakaike; Ill. Pterid. Jap. 8: 419 (1997)
Distribution: Bonin Isl. (Hahajima)
38 OGA
Conservation Status: IUCN E
Synonyms: hidden
= Antigramma cardiophylla (Hance) Tardieu; Naturaliste Malgache 9: 32 (1957)
= Asplenium cardiophyllum (Hance) Baker; Ann. Bot. (Oxford) 5: 311 (1891)
= Boniniella cardiophylla (Hance) Tagawa; J. Jap. Bot. 12: 541 (1936)
= Micropodium cardiophyllum Hance; J. Bot. 21: 268 (1883)
= Phyllitis cardiophylla (Hance) Ching; Icon. Filic. Sin. t. 27 (1930)
species :
Hymenasplenium changputungense
(Ching) Viane & S.Y.Dong; Fl. China 2-3: 312 (2013)
Distribution: China (NW-Yunnan)
36 CHI
Synonyms: hidden
= Asplenium changputungense Ching; Acta Phytotax. Sin. 10: 188 (1965)
species :
Hymenasplenium cheilosorum
(Kunze ex Mett.) Tagawa; Acta Phytotax. Geobot. 7(2): 84 (1938)
Distribution: China (Fujian, Guangdong, Guangxi, Guizhou, Hainan, Yunnan, Zhejiang); Tibet; S-Japan (Yakushima Isl.); Ryukyu Isl. (Amami Oshima, Okinawa Isl.); Taiwan; Nepal; Bhutan; India (Arunachal Pradesh, Assam State, Chhattisgarh, Karnataka, Kerala, Madhya Pradesh, Manipur, Meghalaya, Mizoram, Nagaland, Sikkim, Tamil Nadu, Uttarakhand, West Bengal); Sri Lanka; Bangladesh; Myanmar (Kachin, etc.); Thailand (N-Thailand, NE-Thailand, SE-Thailand, peninsular Thailand); Laos; Vietnam; peninsular Malaysia (Cameron Highlands, Pahang); Borneo (Mt. Kinabalu, Sabah); Sulawesi; Philippines
36 CHI TIB 38 JAP NNS TAI 40 BAN BHU IND NEP SRL 41 LAO MYA THA VIE 42 BOR MLY PHI SUL
Synonyms: hidden
= Asplenium cheilosorum Kunze ex Mett.; Abh. Senckenb. Naturf. Ges. 3(1) [Aspl.]: 133, t. 5, f. 12-13 (1859)
= Asplenium heterocarpum Wall.; Numer. List no. 218 (1829), nomen
= Asplenium heterocarpum Wall. ex Hook.; Sp. Fil. 8: 132, t. 175 (1860)
species :
Hymenasplenium chingii
K.W.Xu, Li Bing Zhang & W.B.Liao; Phytotaxa 358(1): 2 (2018)
Distribution: China (Yunnan, Guizhou)
36 CHI
species :
Hymenasplenium delitescens
(Maxon) L.Regalado & Prada; Amer. Fern J. 101(4): 278 (2012)
Distribution: Mexico (Chiapas, Veracruz, Oaxaca, Tabasco); Guatemala; Belize; Honduras; Nicaragua; Costa Rica; Panama; Cuba; Colombia (Antioquia, Bolívar, Cesar, Chocó, La Guajira, Magdalena, Meta, Santander); Venezuela (Amazonas, Aragua, Barinas, Falcon, Lara, Portuguesa, Sucre); French Guiana; Ecuador; Peru; Bolivia (Beni, La Paz); Brazil (Acre, Mato Grosso, Pará, Rondônia)
79 MEX 80 BLZ COS GUA HON NIC PAN 81 CUB 82 CLM FRG VEN 83 BOL ECU PER 84 BRA
Synonyms: hidden
= Asplenium delitescens (Maxon) A.R.Sm.; Amer. Fern J. 66(4): 120 (1976)
= Asplenium delitescens (Maxon) L.D.Gómez; Brenesia 8: 52 (1976), isonym
= Diplazium delitescens Maxon; Contr. U.S. Natl. Herb. 10: 497 (1908)
species :
Hymenasplenium denticulatum
K.W.Xu, Li Bing Zhang & W.B.Liao; Phytotaxa 358(1): 5 (2018)
Distribution: China (Guizhou, Chongqing)
36 CHI
species :
Hymenasplenium distans
Li Bing Zhang, K.W.Xu & Liang Zhang; Phytotaxa 358(1): 7 (2018)
Distribution: Vietnam
41 VIE
species :
Hymenasplenium excisum
(C.Presl) S.Linds.; Thai Forest Bull., Bot. 37: 69 (2009)
Name: Scalloped spleenwort
Distribution: China (Guangdong, Guangxi, Guizhou, Hainan, Yunnan); Tibet; Ryukyu Isl.; Volcano Isl. (Minami-Iwojima); Taiwan; Nepal; Bhutan; India (Andhra Pradesh, Arunachal Pradesh, Assam State, Kerala, Manipur, Meghalaya, Mizoram, Nagaland, Odisha, Sikkim, Tamil Nadu, Uttarakhand, West Bengal); Sri Lanka; Myanmar (Kachin, etc.); Thailand (N-Thailand, C-Thailand, SE-Thailand, peninsular Thailand); Cambodia; Vietnam; peninsular Malaysia (Cameron Highlands, Pahang); Sulawesi; Palawan; Philippines; Moluccas (Seram); New Guinea (Irian Jaya, Papua New Guinea); Solomon Isl. (Guadalcanal); Australia (NE-Queensland); Vanuatu; Fiji (Viti Levu, Vanua Levu, Taveuni); American Samoa (Tutuila); Cook Isl. (Rarotonga); Hawaii (Kauai, Oahu, Molokai, Lanai, Maui, Hawaii Isl.)
36 CHI TIB 38 KZN NNS TAI 40 BHU IND NEP SRL 41 CBD MYA THA VIE 42 MLY MOL NWG PHI PLW SUL 43 SOL 50 AUS 60 ASA FIJ VAN 61 COO 63 HAW
Synonyms: hidden
= Asplenium excisum C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 74 (1851)
= Asplenium excisum var. serratum Azeez & Madhus.; Spleenworts S. India 145, fig. 16 A-E, t. 19 A-F (2008)
= Asplenium rahaoense Y.Yabe ex Matsum. & Hayata; J. Coll. Sci. Imp. Univ. Tokyo 22: 605 (1906), nom. nud.
= Asplenium resectum var. australiense F.M.Bailey; Proc. Roy. Soc. Queensland 1: 14 (1885)
= Asplenium resectum var. rahaoense Y.Yabe ex Hayata; J. Coll. Sci. Imp. Univ. Tokyo 30(1): 438 (1911)
= Asplenium subresectum Copel.; Philipp. J. Sci. 83: 97, t. 1 (1954)
= Asplenium unilaterale f. majus C.Chr.; Bull. Bernice P. Bishop Mus. 177: 67 (1943)
= Asplenium unilaterale var. australiense (F.M.Bailey) Domin; Biblioth. Bot. 20: 95 (1914)
= Asplenium unilaterale var. majus (C.Chr.) Sledge; Bull. Brit. Mus. (Nat. Hist.), Bot. 3: 246 (1965)
= Asplenium unilaterale var. rahaoense (Hayata) Hayata; Gen. Ind. Fl. Taiwan 103 (1917)
= Hymenasplenium rahaoense (Hayata) H.Itô ex Tuyama; Bot. Mag. (Tokyo) 51: 126 (1937)
= Hymenasplenium unilaterale var. rahaoense (Y.Yabe ex Hayata) Nemoto; Fl. Japan, ed. 2, 50 (1936)
species :
Hymenasplenium filipes
(Copel.) Sugim.; Keys Herb. Pl. Jap. Pterid. 356, 406 (1966)
Distribution: Japan; Taiwan; Borneo (Sabah); Philippines
38 JAP TAI 42 BOR PHI
Remarks: tentative synonymy of H. filipes and adiantifrons fide Knapp (Ferns of Taiwan). Relationship to H. murakami-hatanakae (A. cataractarum) uncertain.
Synonyms: hidden
= Asplenium adiantifrons (Hayata) Ching; Acta Phytotax. Sin. 10: 187 (1965)
= Asplenium filipes Copel.; Philipp. J. Sci., Bot. 4: 34 (1908)
= Asplenium resectum f. adiantifrons Hayata; Icon. Pl. Formosan. 4: 226, C 155 (1914)
= Hymenasplenium adiantifrons (Hayata) Viane & S.Y.Dong; Fl. China 2-3: 312 (2013)
species :
Hymenasplenium furfuraceum
(Ching) Viane & S.Y.Dong; Fl. China 2-3: 311 (2013)
Distribution: China (W-Yunnan)
36 CHI
Synonyms: hidden
= Asplenium furfuraceum Ching; Acta Phytotax. Sin. 10: 190 (1965)
species :
Hymenasplenium guineense
Li Bing Zhang, W.B.Liao & K.W.Xu; Phytotaxa 416(1): 34 (2019)
Distribution: Guinea; Ghana
22 GHA GUI
species :
Hymenasplenium hastifolium
K.W.Xu, Li Bing Zhang & W.B.Liao; Phytotaxa 333(2): 281 (2018)
Distribution: China (Guangxi)
36 CHI
species :
Hymenasplenium hoffmannii
(Hieron.) L.Regalado & Prada; Amer. Fern J. 101(4): 278 (2012)
Distribution: Mexico (Chiapas, Colima, Guerrero, Jalisco, Nayarit, Oaxaca, Puebla, Quintana Roo, Veracruz); Guatemala; El Salvador; Honduras; Nicaragua; Costa Rica; Panama; Trinidad; Colombia (Antioquia, Bolívar); Venezuela (Aragua, Portuguesa, Sucre)
79 MEX 80 COS ELS GUA HON NIC PAN 81 TRT 82 CLM VEN
Synonyms: hidden
= Asplenium hoffmannii Hieron.; Hedwigia 60: 258 (1919)
= Asplenium membranifolium Maxon; Amer. Fern J. 24: 72 (1934)
species :
Hymenasplenium hondoense
(N.Murak. & Hatan.) Nakaike; New Fl. Japan, Pterid. 841 (1992)
Distribution: China (Fujian, Guangxi, Sichuan); South Korea (Cheju Isl.); Japan; C-Nepal; Bhutan; India (Kerala, ?Manipur, Mizoram, Tamil Nadu, Tripura); Myanmar
36 CHI 38 JAP KOR 40 BHU IND NEP 41 MYA
Conservation Status: IUCN NT for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Asplenium hondoense N.Murak. & Hatan.; J. Fac. Sci. Univ. Tokyo, Sect. 3, Bot. 14(3): 191 (1988)
= Asplenium unilaterale var. birii B.K.Nayar & Geev.; Fern Fl. Malabar: 289 (1993) (fide Fraser-Jenkins 2008)
= Asplenium unilaterale var. delicatulum C.S.P.Parish ex Bedd.; Handb. Ferns Brit. India: 153 (1883)
species :
Hymenasplenium hovenkampii
K.W.Xu & Li Bing Zhang; Phytotaxa 595(1): 120 (2023)
Distribution: Lesser Sunda Isl. (Flores)
42 LSI
species :
Hymenasplenium ikenoi
(Makino) Viane; Fl. China 2-3: 309 (2013)
Distribution: China (Hainan); Ryukyu Isl.; ?Thailand; Vietnam
36 CHI 38 NNS 41 VIE
Synonyms: hidden
= Asplenium cardiophyllum subsp. ikenoi (Makino) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 1: 403 (2016)
= Asplenium ikenoi (Makino) H.Ohba; Sci. Rep. Tohoku Imp. Univ., Ser. 4, Biol. 36: 94 (1971)
= Boniniella ikenoi (Makino) Hayata; Bot. Mag. (Tokyo) 41: 709 (1927)
= Phyllitis ikenoi (Makino) C.Chr.; Index Filic. 492 (1906)
= Scolopendrium ikenoi Makino; Bot. Mag. (Tokyo) 13: 103 (1899)
species :
Hymenasplenium inthanonense
N.Murak. & J.Yokoy.; Thai Forest Bull., Bot. 26: 43 (1998)
Distribution: Thailand (N-Thailand: Chiang Mai)
41 THA
Conservation Status: VU in Thailand
species :
Hymenasplenium kenyense
Li Bing Zhang, K.W.Xu & Kamau; Phytotaxa 416(1): 36 (2019)
Distribution: Ethiopia; Rwanda; Kenya; Tanzania
23 RWA 24 ETH 25 KEN TAN
species :
Hymenasplenium kinabaluense
Li Bing Zhang & K.W.Xu; Phytotaxa 419(1): 2 (2019)
Distribution: Borneo (Sabah)
42 BOR
species :
Hymenasplenium laetum
(Sw.) L.Regalado & Prada; Amer. Fern J. 101(4): 278 (2012)
Distribution: Uganda [I]; Tanzania [I]; Mexico (Chiapas, Oaxaca, San Luis Potosi, Tabasco, Veracruz); Guatemala; Belize; El Salvador; Honduras; Nicaragua; Costa Rica; Panama; Cuba; Jamaica; Hispaniola (Dominican Republic, Haiti); Puerto Rico; Nevis; St. Kitts; Montserrat; Guadeloupe; Martinique; St. Vincent; Grenada; Tobago; Trinidad; Colombia (Antioquia, Cauca, Chocó, Magdalena, Nariño, Norte de Santander, Santander, Tolima, Vaupés); Venezuela (Anzoategui, Aragua, Bolívar, Venezuela Distrito Federal, Falcon, Merida, Miranda, Monagas, Portuguesa, Sucre, Yaracuy, Zulia); Guyana; Surinam; French Guiana; Ecuador; Galapagos; Peru; Bolivia (Beni, Cochabamba, La Paz, Santa Cruz); Brazil (Amazonas, Espírito Santo, Minas Gerais, Pará, Pernambuco, Rio de Janeiro, São Paulo); Paraguay (Canindeyu, Concepcion); N-Argentina (Misiones, Salta, Tucuman)
25 tan uga 79 MEX 80 BLZ COS ELS GUA HON NIC PAN 81 CUB GDL GRE HIS JAM MAR MTS PUE STK STV TRT 82 CLM FRG GUY SUR VEN 83 BOL ECU GAL PER 84 BRA 85 ARG PAR
Synonyms: hidden
= Asplenium fraternum Fée; Mém. Foug. 9 (Cat. Foug. Mex.): 16 (1857)
= Asplenium hasslerianum Christ; Repert. Spec. Nov. Regni Veg. 6: 351 (1909)
= Asplenium inaequalidens Fée; Mém. Foug. 11 (Hist. Foug. Antil.): 34, t. 9, f. 3 (1866)
= Asplenium laetum Sw.; Syn. Fil. 79, 271 (1806)
= Asplenium lugubre Liebm.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afh., ser. 5, 1: 243 (seors. 91) (1849)
= Asplenium schkuhrianum C.Presl; Tent. Pterid. 107 (1836)
= Asplenium virens Desv.; Mem. Soc. Linn. Paris 6, 2: 273 (1827) (non C. Presl)
species :
Hymenasplenium laterepens
N.Murak. & X.Cheng ex Y.Fen Chang & K.Hori; Phytotaxa 374(4): 284 (2018)
Distribution: China (Yunnan, Guangxi)
36 CHI
species :
Hymenasplenium latedens
(Ching) Viane & S.Y.Dong; Fl. China 2-3: 311 (2013)
Distribution: China (SE-Yunnan)
36 CHI
Synonyms: hidden
= Asplenium latedens Ching; Acta Phytotax. Sin. 10: 187 (1965)
species :
Hymenasplenium madagascariense
Li Bing Zhang, W.B.Liao & K.W.Xu; Phytotaxa 419(1): 5 (2019)
Distribution: Madagascar
29 MDG
species :
Hymenasplenium murakami-hatanakae
Nakaike; New Fl. Japan, Pterid. 841 (1992), nom. nov.
Distribution: Japan; Taiwan
38 JAP TAI
Remarks: kept separate by Knapp (Ferns of Taiwan, 2011 ff.)
Synonyms: hidden
= Asplenium cataractarum Rosenst.; Hedwigia 66: 334 (1915) (non Blume 1828)
species :
Hymenasplenium neocaledonicum
Li Bing Zhang & K.W.Xu; Phytotaxa 419(1): 5 (2019)
Distribution: New Caledonia
60 NWC
species :
Hymenasplenium ngheanense
Li Bing Zhang, K.W.Xu & N.T.Lu; Phytotaxa 358(1): 10 (2018)
Distribution: Vietnam
41 VIE
species :
Hymenasplenium nigricostatum
Li Bing Zhang & K.W.Xu; Phytotaxa 419(1): 9 (2019)
Distribution: Borneo (Sabah)
42 BOR
species :
Hymenasplenium obliquissimum
(Hayata) Sugim.; Keys Herb. Pl. Jap. Pterid. 356, 406 (1966)
Distribution: China (Guangdong, Guangxi, Guizhou, Hunan, Jiangxi, Sichuan, Yunnan); Japan; Nepal; Bhutan; India (Arunachal Pradesh, Himachal Pradesh, Manipur, Meghalaya, Nagaland, Sikkim, Uttarakhand, West Bengal); Myanmar; Vietnam; ?Philippines
36 CHI 38 JAP 40 BHU IND NEP 41 MYA VIE
Remarks: kept separate from H. amoenum by Fraser-Jenkins et al. (2017)
Synonyms: hidden
= Asplenium obliquissimum (Hayata) Sugim. & Sa.Kurata; J. Geobot. 8: 27 (1959)
= Asplenium unilaterale var. obliquissimum Hayata; Icon. Pl. Formos. 4: 230 (1914)
= Hymenasplenium unilaterale var. obliquissimum (Hayata) Hayata ex Sasaki
species :
Hymenasplenium obscurum
(Blume) Tagawa; Acta Phytotax. Geobot. 7(2): 83 (1938)
Distribution: Tanzania; Malawi; E-Zimbabwe; Mozambique; South Africa (Limpopo); Madagascar; China (Fujian, Guangdong, Guangxi, Guizhou, Hainan, Yunnan); Nepal; Bhutan; India (Andhra Pradesh, Arunachal Pradesh, Chhattisgarh, Kerala, Madhya Pradesh, Maharashtra, Manipur, Meghalaya, Karnataka, Mysore, Odisha, Tamil Nadu); Sri Lanka; Myanmar; N-Thailand; Vietnam; Sumatra; Borneo; Sulawesi; Palawan; Java; Lesser Sunda Isl. (Lombok, Bali, Sumbawa); Moluccas
25 TAN 26 MLW MOZ ZIM 27 SAF 29 MDG 36 CHI 40 BHU IND NEP SRL 41 MYA THA VIE 42 BOR JAW LSI MOL PLW SUL SUM
Remarks: H. pseudobscurum is included here by Fraser-Jenkins et al. (2015, 2017), but kept separate by Fl. of China and recently by Knapp & Hsu (2017)
Synonyms: hidden
= Asplenium auritum var. obscurum (Blume) E.Fourn.; Mexic. Pl. 1: 106 (1872)
= Asplenium cristatum Wall.; Numer. List no. 211 (1829), nomen
= Asplenium erosodentatum Blume; Enum. Pl. Javae 2: 182 (1828)
= Asplenium obscurum Blume; Enum. Pl. Javae 2: 181 (1828)
= Asplenium obscurum var. angustum Tagawa; Acta Phytotax. Geobot. 5(3): 194 (1936)
= Asplenium serriforme Mett.; Abh. Senckenberg. Naturf. Ges. 3(1): 163 (1859), as ´serraeforme´
= Asplenium unilaterale var. bivalvatum B.K.Nayar & Geev.; Fern Fl. Malabar: 292 (1993) (fide Fraser-Jenkins 2008)
= Asplenium unilaterale var. cristatum Bonap.; Notes Pteridol. 13: 68 (1921)
= Hymenasplenium bilabiatum N.Murak.; J. Pl. Res. 108: 261 (1995), nom. nud.
= Hymenasplenium bivalvatum (B.K.Nayar & Geev.) Viane; Fl. China 2-3: 316 (2013)
= Hymenasplenium obscurum var. angustum (Tagawa) Tagawa; Acta Phytotax. Geobot. 7(2): 83 (1938)
species :
Hymenasplenium obtusidentatum
Y.Fen Chang & G.Cheng Zhang; PhytoKeys 211: 101, figs. 1-3 (2022)
Distribution: China (Yunnan)
36 CHI
species :
Hymenasplenium obtusifolium
(L.) L.Regalado & Prada; Amer. Fern J. 101(4): 278 (2012)
Distribution: Tuamotu Arch. [I]; Panama; Jamaica; Puerto Rico; Montserrat; Guadeloupe; Dominica; Martinique; St. Lucia; St. Vincent; Grenada; Tobago; Trinidad; Colombia (Cundinamarca, Magdalena, Norte de Santander); Venezuela (Anzoategui, Aragua, Barinas, Carabobo, Venezuela Distrito Federal, Falcon, Lara, Merida, Miranda, Sucre, Yaracuy)
61 tua 80 PAN 81 DOM GDL GRE JAM MAR MTS PUE STL STV TRT 82 CLM VEN
Synonyms: hidden
= Asplenium aquaticum Klotzsch & H.Karst.; Linnaea 20: 354 (1847)
= Asplenium obtusifolium L.; Sp. Pl. 2: 1080 (1753)
= Asplenium obtusifolium var. aquaticum (Klotzsch & H.Karst.) Giesenh.; Flora 76: 172, fig. 1 (1892)
= Asplenium riparium var. obtusifolium (L.) Hook.; Sp. Fil. [W. J. Hooker] 3: 119. tab. 169. fig. 4 (1860)
= Chamaefilix obtusifolia (L.) Farw.; Amer. Midl. Naturalist 12: 271 (1931)
species :
Hymenasplenium ofeliae
(Salgado) Barcelona & Pelser; Blumea 66(1): 97 (2021)
Distribution: Philippines
42 PHI
Synonyms: hidden
= Asplenium ofeliae Salgado; Amer. Fern J. 93(4): 193 (2003)
species :
Hymenasplenium oligosorum
Li Bing Zhang, K.W.Xu & Lorence; Phytotaxa 419(1): 11 (2019)
Distribution: [Western] Samoa
60 SAM
species :
Hymenasplenium ortegae
(N.Murak. & R.C.Moran) L.Regalado & Prada; Amer. Fern J. 101(4): 279 (2012)
Distribution: Colombia (Meta); Venezuela (Apure, Aragua, Portuguesa, Zulia); Ecuador; Peru; Brazil (Acre)
82 CLM VEN 83 ECU PER 84 BRA
Synonyms: hidden
= Asplenium ortegae N.Murak. & R.C.Moran; Ann. Missouri Bot. Gard. 80(1): 23 (1993)
species :
Hymenasplenium perriei
Li Bing Zhang & K.W.Xu; Phytotaxa 356(2): 177 (2018)
Distribution: Vanuatu; Fiji
60 FIJ VAN
species :
Hymenasplenium phamhoanghoi
Li Bing Zhang, K.W.Xu & T.T.Luong; Phytotaxa 358(1): 10 (2018)
Distribution: Vietnam
41 VIE
species :
Hymenasplenium praestans
(Copel.) Testo, Riibe & Sundue; Syst. Bot. 46(2): 267 (2021)
Distribution: Peru; Bolivia (Beni); Brazil (Acre)
83 BOL PER 84 BRA
Synonyms: hidden
= Athyrium praestans Copel.; Amer. Fern J. 38: 132, t. 11 (1949)
= Diplazium praestans (Copel.) Maxon ex C.V.Morton; Contr. U.S. Nat. Herb. 38: 4 (1967)
species :
Hymenasplenium pseudobscurum
Viane; Fl. China 2-3: 315 (2013)
Distribution: China (Guizhou, Hainan, Yunnan); Taiwan; N-Thailand; N-Vietnam
36 CHI 38 TAI 41 THA VIE
Remarks: included in H. obscurum by Fraser-Jenkins et al. (2015), but kept separate by Fl. of China and recently by Knapp & Hsu (2017, in prep.)
species :
Hymenasplenium pubirhizoma
(Ching & Z.Y.Liu) K.W.Xu & Li Bing Zhang; Phytotaxa 351(2): 186 (2018)
Distribution: China (Yunnan, Sichuan, Guangdong, Guangxi, Guizhou, Hunan, Jiangxi); Japan (incl. Yakushima Isl.); Ryukyu Isl.; S-India; Myanmar; Vietnam; Borneo (Mt. Kinabalu, Sabah); Philippines
36 CHI 38 JAP NNS 40 IND 41 MYA VIE 42 BOR PHI
Synonyms: hidden
= Asplenium pubirhizoma Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 3(4): 7 (1983)
species :
Hymenasplenium purpurascens
(Mett.) L.Regalado & Prada; Amer. Fern J. 101(4): 279 (2012)
Distribution: Colombia; Venezuela (Portuguesa, Zulia); Ecuador; Peru
82 CLM VEN 83 ECU PER
Synonyms: hidden
= Asplenium melanopus Sodiro; Crypt. Vasc. Quit. 189 (1893)
= Asplenium purpurascens Mett.; Kuhn, Linnaea 36: 102 (1869)
= Asplenium purpurascens var. crespianum Bosco; Nuovo Giorn. Bot. Ital. new ser. 45, no. 2: 146 (1938)
= Diplazium melanopus (Sodiro) Hieron.; Bot. Jahrb. Syst. 34: 456 (1904)
species :
Hymenasplenium quangnamense
Li Bing Zhang, K.W.Xu & Liang Zhang; Phytotaxa 358(1): 15 (2018)
Distribution: Myanmar; Vietnam
41 MYA VIE
species :
Hymenasplenium queenslandicum
Li Bing Zhang & K.W.Xu; Phytotaxa 419(1): 14 (2019)
Distribution: Australia (Queensland)
50 AUS
species :
Hymenasplenium quercicola
(Ching) Viane & S.Y.Dong; Fl. China 2-3: 312 (2013)
Distribution: China (Yunnan)
36 CHI
Synonyms: hidden
= Asplenium quercicola Ching; Acta Phytotax. Sin. 10: 188 (1965)
species :
Hymenasplenium repandulum
(Kunze) L.Regalado & Prada; Amer. Fern J. 101(4): 279 (2012)
Distribution: Panama; Colombia (Chocó); Venezuela (Barinas, Merida, Portuguesa, Trujillo); French Guiana; Ecuador; Peru; Bolivia (La Paz)
80 PAN 82 CLM FRG VEN 83 BOL ECU PER
Synonyms: hidden
= Asplenium repandulum Kunze; Linnaea 9: 65 (1834)
species :
Hymenasplenium riparium
(Liebm.) L.Regalado & Prada; Amer. Fern J. 101(4): 279 (2012)
Distribution: Mexico (Chiapas, Oaxaca, Puebla, Veracruz); Guatemala; Honduras; Nicaragua; Costa Rica; Panama; Colombia (Cundinamarca, Huila, Magdalena, Nariño, Santander, Tolima); Venezuela (Barinas, Portuguesa, Trujillo); French Guiana; Ecuador; Peru; Brazil (Bahia, Rio de Janeiro, Minas Gerais, Paraná, Santa Catarina, Rio Grande do Sul)
79 MEX 80 COS GUA HON NIC PAN 82 CLM FRG VEN 83 ECU PER 84 BRA
Synonyms: hidden
= Asplenium obtusifolium var. riparium (Liebm.) Domin; Rozpr. Kral. Ceske Spolecn. Nauk, Tr. Mat.-Prir. n. s. 2: 175 (1929)
= Asplenium riparium Liebm.; Vid. Selsk. Skr. 5, 1: 244 (1849)
= Chamaefilix obtusifolia var. riparia (Liebm.) Farw.; Amer. Midl. Naturalist 12: 271 (1931)
species :
Hymenasplenium rivulare
(Fraser-Jenk.) Viane & S.Y.Dong; Fl. China 2-3: 308 (2013)
Distribution: India (Karnataka, Kerala, Tamil Nadu)
40 IND
Remarks: if synonymy of A. emarginatodentatum is confirmed, it would have priority
Conservation Status: IUCN NT for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Asplenium emarginatodentatum Zenker ex Kunze; Linnaea 24: 263 (1851) (fide Fraser-Jenkins)
= Asplenium hindusthanense Bir; Fern Gaz. 14: 309 (1994), without replaced synonym ref.
= Asplenium rivale (Bedd.) Bir; Bir & Sago, Perspect. Pl. Sci. India 1-22 (1989), non Spruce ex Baker 1867
= Asplenium rivulare Fraser-Jenk.; Taiwania 53(2): 190 (2008), nom. nov. for Asplenium rivale (Bedd.) Bir
= Asplenium unilaterale var. rivale Bedd.; Handb. Ferns Brit. India 153 (1883), non Spruce (fide Fraser-Jenkins 2008)
species :
Hymenasplenium sabahense
Li Bing Zhang, K.W.Xu & C.W.Chen; Phytotaxa 416(1): 38 (2019)
Distribution: Borneo (Sabah)
42 BOR
species :
Hymenasplenium samoaense
Li Bing Zhang, K.W.Xu & K.R.Wood; Phytotaxa 419(1): 19 (2019)
Distribution: [Western] Samoa
60 SAM
species :
Hymenasplenium sinense
K.W.Xu, Li Bing Zhang & W.B.Liao; Phytotaxa 358(1): 17 (2018)
Distribution: China (Guizhou, Yunnan, Jiangxi, Guangxi)
36 CHI
species :
Hymenasplenium solomonense
Li Bing Zhang & K.W.Xu; Phytotaxa 419(1): 19 (2019)
Distribution: New Guinea (Papua New Guinea); Solomon Isl.
42 NWG 43 SOL
species :
Hymenasplenium speluncicola
Li Bing Zhang, K.W.Xu & H.He; Phytotaxa 358(1): 20 (2018)
Distribution: China (Guangxi, Guizhou)
36 CHI
species :
Hymenasplenium subnormale
(Copel.) Nakaike; New Fl. Japan, Pterid. 841 (1992)
Distribution: China (SW-Yunnan); Ryukyu Isl. (Yonagunijima); Taiwan; peninsular Malaysia (Selangor, P. Tioman, Pahang); Borneo (Mt. Kinabalu, Sabah, Kalimantan); Sulawesi; Palawan; Philippines (Luzon); Moluccas (Seram, Ambon); ?Vanuatu
36 CHI 38 NNS TAI 42 BOR MLY MOL PHI PLW SUL
Remarks: not in India
Synonyms: hidden
= Asplenium subnormale Copel.; Perkins, Fragm. Fl. Philip. 183, t. 4, f. B (1905)
species :
Hymenasplenium szechuanense
(Ching) Viane & S.Y.Dong; Fl. China 2-3: 311 (2013)
Distribution: China (Sichuan, Yunnan)
36 CHI
Synonyms: hidden
= Asplenium szechuanense Ching; Acta Phytotax. Sin. 10: 188 (1965)
species :
Hymenasplenium tholiforme
Liang Zhang, W.B.Ju & K.W.Xu; PhytoKeys 204: 46, fig. 1 (2022)
Distribution: SE-Tibet (Medog)
36 TIB
species :
Hymenasplenium trapeziforme
(Wall. ex Roxb.) comb.ined.
Distribution: Japan; Bonin Isl. (Chichijima, Hahajima); Volcano Isl. (Kita-Iwojima, Minami-Iwojima); India (Arunachal Pradesh, Assam State, ?Manipur, Meghalaya, ?Nagaland, ?Tripura); Sri Lanka; Bangladesh; Myanmar; Vietnam; peninsular Malaysia; Sumatra; Borneo (Mt. Kinabalu, Sabah); Sulawesi; Palawan; Philippines; Lesser Sunda Isl. (Bali, Sumbawa, Flores); Moluccas (Seram); New Guinea (Irian Jaya, Papua New Guinea); Solomon Isl. (Guadalcanal, Kolombangara, Malaita, New Georgia, Ranongga, Rendova, Vella Lavella); Australia (NE-Queensland); New Caledonia (Loyalty Isl., Ile des Pins); Vanuatu; Tonga (´Eua); Fiji (Viti Levu, Vanua Levu, Ovalau, Taveuni); American Samoa (Ta´u); [Western] Samoa (Savai´i, ´Upolu); Northern Marianas (Pagan, Anatahan, Alamagan, Sarigan); Guam; Hawaii (Kauai, Oahu, Molokai, Lanai, Maui, Hawaii Isl.)
38 JAP KZN OGA 40 BAN IND SRL 41 MYA VIE 42 BOR LSI MLY MOL NWG PHI PLW SUL SUM 43 SOL 50 AUS 60 ASA FIJ NWC SAM TON VAN 62 GUM MRN 63 HAW
Remarks: this includes all Indian and Indochina populations of A. unilaterale fide Fraser-Jenkins et al. (2017). Whether the Australasian and Pacific records belong here, remains uncertain
Synonyms: hidden
= Asplenium abscissum Blume; Enum. Pl. Javae 2: 182 (1828), nom. illeg.
= Asplenium decurrens Wall.; Numer. List no. 190 (1829), nomen
= Asplenium erythrocaulon Blume; Enum. Pl. Javae 2: 183 (1828)
= Asplenium fraternum C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6: 74 (1851), nom. inval.
= Asplenium porphyrocaulon Blume; Enum. Pl. Javae 2: 182 (1828)
= Asplenium trapeziforme Wall.; Numer. List no. 2213 (1829), nomen
= Asplenium trapeziforme Wall. ex Roxb.; Griffith, Calcutta J. Nat. Hist. 4: 497 (1844)
= Asplenium unilaterale var. saxicola M.Kato & K.Iwats.; J. Fac. Sci. Univ. Tokyo, Sect. 3, Bot. 14(1): 46 (1986)
species :
Hymenasplenium triquetrum
(N.Murak. & R.C.Moran) L.Regalado & Prada; Amer. Fern J. 101(4): 279 (2012)
Distribution: Bolivia (Beni, Cochabamba, La Paz); Brazil (Bahia, Espírito Santo, Minas Gerais, Paraná, Rio de Janeiro, Rio Grande do Sul, Santa Catarina, São Paulo); Paraguay (Alto Paraguay); Argentina (Misiones)
83 BOL 84 BRA 85 ARG PAR
Synonyms: hidden
= Asplenium triquetrum N.Murak. & R.C.Moran; Ann. Missouri Bot. Gard. 80(1): 31 (1993)
species :
Hymenasplenium unilaterale
(Lam.) Hayata; Bot. Mag. (Tokyo) 41: 712 (1927)
Distribution: Guinea; Sierra Leone; Liberia; Ivory Coast; Ghana; Niger; Nigeria; Cameroon; Bioko Isl. [Fernando Poo]; Principe Isl.; São Tomé; Gabon; Congo [Brazzaville]; Central African Republic; Sudan; Ethiopia; Uganda; Kenya; Tanzania; Angola; Malawi; E-Zimbabwe; South Africa (Mpulamanga); Eswatini [Swaziland]; Madagascar; Comores (Anjouan); Mauritius; La Réunion; Seychelles
22 GHA GUI IVO LBR NGA NGR SIE 23 BIO CAF CMN CON GAB PRI SAO 24 ETH SUD 25 KEN TAN UGA 26 ANG MLW ZIM 27 SAF SWZ 29 COM MAU MDG REU SEY
Remarks: for Asian populations see A. trapeziforme
Synonyms: hidden
= Asplenium resectum Sm.; Icon. ined. 8, t. 72 (1791)
= Asplenium unilaterale Lam.; Encycl. 2: 305 (1786)
= Asplenium unilaterale subsp. africanum G.Kunkel; Nova Hedwigia 17,1/4: 147 (1969), nom. nud.
= Chamaefilix unilateralis (Lam.) Farw.; Amer. Midl. Naturalist 12: 271 (1931)
species :
Hymenasplenium vanuatuense
Li Bing Zhang & K.W.Xu; Phytotaxa 419(1): 24 (2019)
Distribution: Vanuatu
60 VAN
species :
Hymenasplenium volubile
(N.Murak. & R.C.Moran) L.Regalado & Prada; Amer. Fern J. 101(4): 279 (2012)
Distribution: Costa Rica; Panama; Colombia (Chocó, Huila, Risaralda, Valle); Ecuador
80 COS PAN 82 CLM 83 ECU
Synonyms: hidden
= Asplenium volubile N.Murak. & R.C.Moran; Ann. Missouri Bot. Gard. 80: 32 (1993)
species :
Hymenasplenium wangpeishanii
Li Bing Zhang & K.W.Xu; Phytotaxa 358(1): 22 (2018)
Distribution: China (Guizhou)
36 CHI
species :
Hymenasplenium wildii
(Bail.) D.J.Ohlsen; Muelleria 37: 21 (2018)
Distribution: Australia (NE-Queensland)
50 AUS
Conservation Status: IUCN V
Synonyms: hidden
= Asplenium wildii F.M.Bailey; Queensl. Bot. Bull. 4: 20, t. 11 2 (1891)
species :
Hymenasplenium wusugongii
Li Bing Zhang, W.B.Liao & K.W.Xu; Phytotaxa 416(1): 38 (2019)
Distribution: Laos
41 LAO
species :
Hymenasplenium ×doanyense
Viane; Aspleniaceae Madagascar 19 (2021)
Distribution: Madagascar
29 MDG
Remarks: Hymenasplenium antsiranaense x madagascariense
species :
Hymenasplenium ×excurum
Viane; Aspleniaceae Madagascar 20 (2021)
Distribution: Madagascar
29 MDG
Remarks: Hymenasplenium excisum x obscurum
species :
Hymenasplenium ×incisoserratum
(Rosenst.) comb.ined.
Distribution: Honduras; Costa Rica; Colombia (Valle); Venezuela
80 COS HON 82 CLM VEN
Remarks: Hymenasplenium hoffmannii x Hymenasplenium laetum
Synonyms: hidden
= Asplenium ×incisoserratum (Rosenst.) N.Murak. & R.C.Moran; Ann. Missouri Bot. Gard. 80(1): 34 (1993)
= Asplenium laetum var. incisoserratum Rosenst.; Repert. Spec. Nov. Regni Veg. 22: 8 (1925)
