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Ver. 19.2
April 17th, 2024

041.0000 Family Polypodiaceae J. & C.Presl; J. & C. Presl, Delic. Prag. 159 (1822)
67 genera, ca. 1700 spp. Phylogeny: Wei & Zhang 2022
Species listed in database: 1718; hybrids: 47
= Platyceriaceae Ching; Acta Phytotax. Sin. 16(3): 18 (1978)
= Drynariaceae Ching; Acta Phytotax. Sin. 16: 1-37 (1978)
= Grammitidaceae Newman; Hist. Brit. Ferns 7 (1840) (Grammitaceae auct., missp.)
= Gymnogrammitidaceae Ching; Acta Phytotax. Sin. 11: 12 (1966)
= Loxogrammaceae Ching ex Pic.Serm.; Webbia 29: 11 (1975)
= Platyceriaceae Ching; Sunyatsenia 5 (1940)
= Pleurosoriopsidaceae Sa.Kurata & Ikebe ex Ching; Acta Phytotax. Sin. 16(4): 17 (1978)
041.3402 Genus Leptochilus Kaulf.; Enum. Filic. 147 (1824)
Distribution: E- and tropical Asia, 3 to Queensland. Species circumscription quite controversial and fluctuating.
Species listed in database: 37; hybrids: 2
= Dendroglossa C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6: 149 (1851)
= Gymnopteris C.Presl; Tent. Pterid. 244 (1836)
= Paraleptochilus Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 198 (1947)
= Colysis C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6: 146 (1851) (fide Fraser-Jenkins 2008 and others)
= Selliguea Blume; (1826) (nec Bory (1824))
= Nistarika B.K.Nayar, Madhus. & Molly; Fern Gaz. 13: 33 (1985)
= Kontumia S.K.Wu & P.K.Lôc; Novon 15(1): 245 (2005) (fide PPG-I)
species :
Leptochilus axillaris (Cav.) Kaulf.; Enum. Filic. 147, t. 1, f. 10 (1824)
Distribution: China (Guizhou, Yunnan); India (NE-India, Assam State, S-India, Bengal); Bangladesh; Nicobar Isl.; N-Myanmar; Thailand; Vietnam; peninsular Malaysia (Perak); Sumatra; Borneo (Sabah); Sulawesi; Palawan; Philippines; Java; Lesser Sunda Isl. (Bali, Lombok); Moluccas; New Guinea (Papua New Guinea)
36 CHI 40 BAN IND 41 MYA NCB THA VIE 42 BOR JAW LSI MLY MOL NWG PHI PLW SUL SUM
Synonyms: hidden
= Acrostichum axillare Cav.; Anal. Hist. Nat. 1: 101 (1799)
= Colysis axillaris (Cav.) Rajkumar & Manickam; Indian Fern J. 18(1-2): 125 (2001)
= Colysis bahupunctika (B.K.Nayar, Madhus. & Molly) Rajkumar & Manickam; Indian Fern J. 18(1-2): 125 (2001) (fide Fraser-Jenkins 2008)
= Gymnogramma axillaris (Cav.) Ces.; Atti Reale Accad. Sci. Napoli 76: 29 (1876)
= Gymnopteris axillaris (Cav.) C.Presl; Tent. Pterid. 244, t. 11, f. 4-5 (1836)
= Gymnopteris variabilis var. axillaris (Cav.) Bedd.; Handb. Ferns Brit. India, t. 271 (1968)
= Leptochilus bahupunctikus (B.K.Nayar, Madhus. & Molly) B.K.Nayar, Madhus. & Molly; Fern Fl. S. India 59 (1998)
= Leptochilus platyphyllus Copel.; Philipp. J. Sci. 37: 340, t. 2 (1928)
= Lomaria serpens Wall.; Numer. List no. 32 (1829), nomen
= Nistarika bahupunctika B.K.Nayar, Madhus. & Molly; Fern Gaz. 13(1): 33 (1985) (fide Fraser-Jenkins 2008)
= Polypodium hymenodes Wall.; Numer. List no. 283 (1829), nomen
= Polypodium leptochiloides Kuhn; Engl., Forschungsr. Gazelle 4(Farne): 10 (1889)
species :
Leptochilus bolsteri (Copel.) Parris; Fern Gaz. 20(7): 305 (2018)
Distribution: Borneo (Sarawak); Philippines (Mindanao)
42 BOR PHI
Remarks: often included in L. macrophyllus
Synonyms: hidden
= Colysis bolsteri (Copel.) Copel.; Fern Fl. Philip. 3: 489 (1960)
= Pleopeltis bolsteri (Copel.) Alderw.; Bull. Dép. Agric. Indes Néerl. 27: 11 (1909)
= Polypodium bolsteri Copel.; Philipp. J. Sci. 1, Suppl. IV: 257, t. 4 A (1906)
species :
Leptochilus brevipes Liang Zhang, Z.L.Liang & Zhi Y.Yu; Phytotaxa 634(2): 144 (2024)
Distribution: China (Yunnan)
36 CHI
species :
Leptochilus cantoniensis (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 343 (1933)
Distribution: China (Guangdong, Hainan); Vietnam
36 CHI 41 VIE
Synonyms: hidden
= Campium cantoniense (Baker) Ching; Sinensia 1: 53 (1930)
= Campium dilatatum Copel.; Philipp. J. Sci. 37: 341-402 (1928)
= Christopteris cantoniensis (Baker) Christ; J. Bot. (Morot) 21: 273 (1908)
= Dendroglossa cantoniensis (Baker) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] [Copeland] 199 (1947)
= Drymoglossum cordatum Christ; Notul. Syst. (Paris) 1: 375 (1911)
= Gymnogramma cantoniensis (Baker) Baker; Hooker´s Icon. Pl. t. 1685 (1887)
= Leptochilus cordatus (Christ) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 343 (1933)
= Myuropteris cordata (Christ) C.Chr.; Dansk Bot. Ark. 6, 3: 73, t. 9, f. 1-2, t. 10, f. 3 (1929)
= Polypodium cantoniense (Baker) Baker; J. Bot. 17: 304 (1879)
species :
Leptochilus chilangensis (V.N.Tu) Liang Zhang & Li Bing Zhang; Phytotaxa 374(2): 172 (2018)
Distribution: Vietnam
41 VIE
Synonyms: hidden
= Colysis chilangensis V.N.Tu; Bot. Zhurn. (Moscow & Leningrad) 65(4): 582 (1980)
species :
Leptochilus chingii Liang Zhang & Li Bing Zhang; Phytotaxa 374(2): 173 (2018), nom. nov.
Distribution: China
36 CHI
Remarks: frequently included in L. pedunculatus
Synonyms: hidden
= Colysis bonii Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 313-331 (1933)
species :
Leptochilus decurrens Blume; Enum. Pl. Javae 2: 206 (1828)
Distribution: China (Guangxi, Guizhou, Hainan, Hongkong, Yunnan); Japan; Bonin Isl. (Hahajima); Taiwan; Nepal; Bhutan; India (Sikkim, E-India, C-India, Cachar, S-India, Arunachal Pradesh, Sikkim, Darjeeling); Sri Lanka; Myanmar; Thailand; Laos; ?Cambodia; Vietnam; peninsular Malaysia (Perak, Pahang); Sumatra; Borneo (Sabah); Sulawesi; Philippines; Java; Lesser Sunda Isl. (Lombok); Moluccas; Christmas Isl. [Austr.]; New Guinea (Papua New Guinea)
36 CHI 38 JAP OGA TAI 40 BHU IND NEP SRL 41 LAO MYA THA VIE 42 BOR JAW LSI MLY MOL NWG PHI SUL SUM XMS
Remarks: Leptochilus lanceolatus is included here by Flora of China but a separate Indian species fide Fraser-Jenkins (2008). Inclusion of L. metallicus, L. thwaitesianus and L. wallii fide Handbook of Ceylon (2006)
Synonyms: hidden
= Acrostichum decurrens Wall.; Numer. List no. 18 (1829), nomen
= Acrostichum decurrens Wall. ex Thwaites; Enum. pl. Zeyl. 381 (1864)
= Acrostichum feejanum Ettingsh.; Denkschr. Kaiserl. Akad. Wiss., Wien. Math.-Naturwiss. Kl. 22: 54 (1864)
= Acrostichum listeri Baker; J. Linn. Soc., Bot. 26: 361 (1889)
= Acrostichum rivulare Wall.; Numer. List no. 2165 (1829), nomen
= Acrostichum variabile Hook.; Sp. Fil. 5: 277 (1864)
= Acrostichum wallii Baker; J. Bot. 10: 146 (1872)
= Anapausia decurrens (Blume) C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 186 (1851)
= Campium decurrens (Blume) Copel.; Philipp. J. Sci. 37: 341-402 (1928)
= Campium metallicum (Bedd.) Copel.; Philipp. J. Sci. 37: 347 (1928)
= Campium wallii (Baker) Copel.; Philipp. J. Sci. 37: 341-402 (1928)
= Colysis decurrens (Blume) Panigrahi; Abstr. & Souv. Nation. Symp. Curr. Trends Pterid. 1991: 13, Publ. 4-6 October (1991)
= Colysis diversifolia W.M.Chu; Acta Bot. Yunnan. 1(2): 93 (1979)
= Colysis metallica (Bedd.) Rajkumar & Manickam; Indian Fern J. 18(1-2): 126 (2001)
= Colysis ×beddomei Manickam & Irudayaraj; Taxon 46: 267 (1997) (fide Kandel & Fraser-Jenkins 2020)
= Dendroglossa metallica (Bedd.) B.K.Nayar & Kaur; Comp. to Bedd. Handb. Ferns Brit. India 100 (1974)
= Dendroglossa wallii (Baker) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 200 (1947)
= Gymnogramma decurrens (Blume) Hook.; Sp. 6: 161 (1864)
= Gymnopteris decurrens (Blume) T.Moore; Index Filic. (T. Moore) 29 (1857)
= Gymnopteris dichotomophlebia Hayata; Icon. Pl. Formosan. 4: 201, f. 136 (1914)
= Gymnopteris feei f. anomala Bedd.; Ferns Brit. India t. 274 (1868)
= Gymnopteris feei T.Moore; Index Filic. (T. Moore) 30 (1857)
= Gymnopteris metallica Bedd.; Handb. Ferns Brit. India, Suppl. 26, t. 390 (1876)
= Gymnopteris obtusifolia C.Presl; Tent. Pterid. 244 (1836)
= Gymnopteris variabilis (Hook.) Bedd.; Handb. Ferns Brit. India, t. 272 (1868)
= Gymnopteris wallichiana C.Presl; Tent. Pterid.: 244 (1836), nom. nud.
= Gymnopteris wallii (Baker) Bedd.; Handb. Ferns Brit. India, Suppl. 26, t. 389 (1876)
= Leptochilus dichotomophlebia Hayata; Icon. Pl. Formosan. 4: 202 (1914), pro syn.
= Leptochilus listeri (Baker) C.Chr.; Index Filic. 12 (1905), 386 (1906)
= Leptochilus metallicus (Bedd.) C.Chr.; Index Filic. 386 (1906)
= Leptochilus obtusifolius (C.Presl) C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 185 (1851)
= Leptochilus trifidus Alderw.; Bull. Dép. Agric. Indes Néerl. 18: 26 (1908) (probably a hybrid L. decurrens x macrophyllus)
= Leptochilus wallii (Baker) C.Chr.; Index Filic. 20 (1905), 388 (1906)
= Leptochilus ×beddomei (Manickam & Irudayaraj) X.C.Zhang & Noot.; Fl. China 2-3: 834 (2013)
= Paraleptochilus decurrens (Blume) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 198 (1947)
= Paraleptochilus metallicus (Bedd.) Á.Löve & D.Löve; Taxon 26(2-3): 324 (1977)
= Selliguea decurrens (Blume) C.Presl; Tent. Pterid. 216 (1836)
species :
Leptochilus digitatus (Baker) Noot.; Blumea 42: 282 (1997)
Distribution: China (Guangdong, Guangxi, Guizhou, Hainan, Yunnan); Vietnam
36 CHI 41 VIE
Synonyms: hidden
= Colysis digitata (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 328 (1933)
= Colysis digitata f. annamensis (Christ) Ching
= Colysis digitata f. cadieri (Christ) Ching
= Colysis digitata f. laciniata Ching
= Colysis tridactylis Fée; Mém. Foug. 5 (Gen. Filic.): 176 (1852), nomen
= Colysis triphylla Ching; Ching & Chu H. Wang, Acta Phytotax. Sin. 8: 170, t. 23(31) (1959)
= Grammitis finlaysoniana Wall.; Numer. List no. 248 (1829), nomen
= Gymnogramma digitata Baker; J. Bot. 28: 267 (1890)
= Gymnogramma finlaysoniana (Wall.) Baker; Ann. Bot. 6: 486 (1891)
= Phymatodes digitata (Baker) Ching; Contr. Inst. Bot. Nat. Acad. Peiping 2: 77 (1933)
= Polypodium ampelideum Christ; J. Bot. (Morot) 19: 78 (1905)
= Polypodium annamense Christ; J. Bot. (Morot) 19: 77 (1905)
= Polypodium cadieri Christ; J. Bot. (Morot) 19: 76 (1905)
= Polypodium digitatum (Baker) C.Chr.; Index Filic. 522 (1906)
= Polypodium podopterum Christ; J. Bot. (Morot) 19: 125 (1905)
= Selliguea finlaysoniana Wall. ex T.Moore; Index Filic. (T. Moore) 64 (1857), nomen
species :
Leptochilus dissimilialatus (Bonap.) Liang Zhang & Li Bing Zhang; Phytotaxa 374(2): 173 (2018)
Distribution: Vietnam; etc.
41 VIE
Remarks: frequently included in L. ellipticus s. l.
Synonyms: hidden
= Colysis dissimilialata (Bonap.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 313-331 (1933)
= Polypodium dissimilialatum Bonap.; Notes Ptéridol. 14: 155 (1924)
species :
Leptochilus dolichopterus (Copel.) Fraser-Jenk. & Amoroso; Taxon. Revis. Indian Subcontinental Pteridophytes 62 (2008)
Distribution: Philippines
42 PHI
Synonyms: hidden
= Pleopeltis dolichoptera (Copel.) Alderw.; Bull. Dép. Agric. Indes Néerl. 27: 9 (1909)
= Polypodium dolichopterum Copel.; Philipp. J. Sci. 1, Suppl. II: 162 (1906)
species :
Leptochilus ellipticus (Thunb.) Noot.; Blumea 42: 283 (1997)
Distribution: China (Anhui, Guangxi, Guizhou, Hainan, Hongkong, Hunan, Jiangsu, Jiangxi, Yunnan, Zhejiang); Tibet; South Korea (incl. Cheju Isl.); Japan; Ryukyu Isl.; Taiwan; Nepal; Bhutan; NE-India (Bengal, Arunachal Pradesh, Manipur, Meghalaya, Sikkim, Darjeeling, Himachal Pradesh); Myanmar; Thailand; Laos; Vietnam; Philippines (Luzon, Leyte, Mindanao, Samar)
36 CHI TIB 38 JAP KOR NNS TAI 40 BHU IND NEP 41 LAO MYA THA VIE 42 PHI
Remarks: Fraser-Jenkins et al. (2015) accepts no varieties, but distinguishes between this taxon and L. pothifolius
Synonyms: hidden
= Colysis boisii (Christ) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 313-331 (1933)
= Colysis decurrens (Wall. ex Hook. & Grev.) Nakaike; New Fl. Japan, Pterid. 13 (1991) [fide Zimmer, in litt.]
= Colysis elegans Sa.Kurata; J. Geobot. 12: 41 (1963)
= Colysis elliptica (Thunb.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 333 (1933)
= Colysis elliptica var. longipes (Ching) L.Shi & X.C.Zhang; Acta Phytotax. Sin. 37(1): 76 (1999)
= Colysis elliptica var. sanjiangensis (H.G.Zhou & Hua Li) R.H.Jiang; Fl. Guangxi 6: 406 (2013)
= Colysis flavescens (Ching) Nakaike, S.Matsumoto & Gurung; Distrib. Maps Pterid. Nepal 3 (repr. from Cryptog. Himal., 2 C. & E. Nepal): 189 (1990)
= Colysis flavescens (Ching) S.R.Ghosh; Pterid. Fl. E. India 497 (2004), isonym
= Colysis leptophylla H.Itô; Bot. Mag. (Tokyo) 53: 68 (1939)
= Colysis longipes Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 313-331 (1933), nom. illeg., non Polypodium longipes auct.
= Colysis pothifolia f. bipinnatifida H.Itô; J. Jap. Bot. 11: 89 (1935)
= Colysis pothifolia monstr. bipinnatifida (H.Itô) Nakaike; Nov. Fl. Jap. (1992)
= Colysis pothifolia var. membranacea Nakai; Bull. Natl. Sci. Mus. Tokyo 27: 23 (1949)
= Colysis sanjiangensis H.G.Zhou & Hua Li; Acta Bot. Yunnan. 15(3): 254 (1993)
= Colysis ×kiusiana Sa.Kurata; J. Geobot. 10: 37 (1961)
= Grammitis decurrens Wall. ex Hook. & Grev.; Ic. Filic., tab. 6 (1827) [fide Zimmer, in litt.]
= Gymnogramma elliptica (Thunb.) Hook. & Baker; Hook. & Baker, Syn. Fil. ed. 2: 389 (1874) (pro parte)
= Gymnogramma elliptica C.B.Clarke; Trans. Linn. Soc. London, Bot. 1: 570 (1880) (pro parte)
= Gymnogramma elliptica var. furcans Tutcher; J. Linn. Soc., Bot. 37: 69 (1904)
= Gymnogramma pothifolia (Buch.-Ham. ex D.Don) Spreng.; Syst. Veg. ed. 16, 4(1): 39 (1827)
= Leptochilus elegans (Sa.Kurata) Nakaike; Checkl. Lycophytes Ferns Japan 35 (2013)
= Leptochilus ellipticus var. longipes (Ching) Noot.; Fl. China 2-3: 837 (2013)
= Leptochilus ×kiusianus (Sa.Kurata) Nakaike; Checkl. Lycophytes Ferns Japan: 35 (2013) (L. ellipticus ×
= Leptochilus longipes (Ching) X.C.Zhang; Lycophytes Ferns China 655 (2012)
= Leptochilus neopothifolius Nakaike; Checkl. Lycophytes Ferns Japan: 35 (2013)
= Pleopeltis elliptica (Thunb.) Alderw.; Bull. Dép. Agric. Indes Néerl. 27: 12 (1909)
= Polypodium boisii Christ; J. Bot. (Morot) 19: 76 (1905)
= Polypodium chunii C.Chr.; Index Filic., Suppl. Tert.: 146 (1934)
= Polypodium ellipticum f. breve Y.C.Wu; Bull. Dept. Biol. Coll. Sci. Sun Yatsen Univ. 3: 322 (1932)
= Polypodium ellipticum Thunb.; Murray (ed.), Syst. Veg. ed. 14: 935 (1784)
= Polypodium faurianum Nakai; J. Coll. Sci. Imp. Univ. Tokyo 31: 416 (1911)
= Polypodium flavescens Ching; Bull. Fan Mem. Inst. Biol. Bot. 2: 22, t. 8 (1931) (fide Fraser-Jenkins 2008)
= Polypodium mediosorum Ching; Bull. Fan Mem. Inst. Biol. Bot. 2: 19, t. 4 (1931)
= Polypodium neoellipticum Koidz.; Bot. Mag. (Tokyo) 43: 388 (1929)
= Selliguea coraiensis Christ; Repert. Spec. Nov. Regni Veg. 5: 11 (1908) (non Polypodium coraiense Christ (1908))
= Selliguea elliptica (C.B.Clarke) Bedd.; Handb. Ferns Brit. India, t. 150 (1867) (pro parte)
= Selliguea elliptica (Thunb.) Bedd.; Handb. Ferns Brit. India, t. 150 (1867) (pro parte)
= Selliguea elliptica var. flagellaris Christ; Bull. Herb. Boissier 7: 6 (1899) (non Polypodium flagellare Christ (1896))
= Selliguea pentaphylla (Baker) Christ; Bull. Acad. Int. Géogr. Bot., sér. 3, 16: 248 (1906), not Baker (1891)
species :
Leptochilus evrardii (Tardieu) Liang Zhang & Li Bing Zhang; Phytotaxa 374(2): 173 (2018)
Distribution: Vietnam
41 VIE
Remarks: frequently included in L. decurrens
Synonyms: hidden
= Colysis evrardii Tardieu; Bull. Soc. Bot. France 87: 372, t. 2(3-4) (1941)
species :
Leptochilus flexilobus (Christ) Liang Zhang & Li Bing Zhang; Phytotaxa 374(2): 174 (2018)
Distribution: China (Chongqing, Guangxi, Guizhou, Hunan, Jiangxi, Sichuan, Yunnan); Taiwan; Vietnam
36 CHI 38 TAI 41 VIE
Remarks: frequently included in L. ellipticus s. lat.
Synonyms: hidden
= Colysis elliptica var. flexiloba (Christ) L.Shi & X.C.Zhang; Acta Phytotax. Sin. 37: 74 (1999)
= Colysis flexiloba (Christ) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 313-331 (1933)
= Colysis flexiloba var. undulatocrenata (C.Chr.) Ching; Bull. Fan Mem. Inst. Biol. 4: 331 (1933)
= Colysis flexiloba var. undulatorepanda (C.Chr.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4(10): 330 (1933)
= Leptochilus ellipticus var. flexilobus (Christ) X.C.Zhang; Lycophytes Ferns China 652 (2012)
= Pleopeltis flexiloba (Christ) Alderw.; Malayan Ferns Fern Allies, Suppl. 383-406 (1917)
= Polypodium ellipticum var. undulatocrenatum C.Chr.; Bull. Acad. Géogr. Bot. Mans 107 (1904)
= Polypodium ellipticum var. undulatorepandum C.Chr.; Bull. Acad. Int. Geogr. Bot. 22: 141 (1913)
= Polypodium flexilobum Christ; Bull. Acad. Géogr. Bot. Mans 107 (1904)
= Polypodium flexilobum var. undulatocrenatum Ching; Bull. Fan Mem. Inst. Biol. Bot. 2(2): 20 (1931), nom. nud.
= Selliguea flexiloba Christ; Bull. Herb. Boissier, sér, 2, 6: 992 (1906)
species :
Leptochilus gracilis Z.L.Liang, Liang Zhang & Li Bing Zhang; Phytotaxa 468(1): 117 (2020)
Distribution: China (Yunnan)
36 CHI
species :
Leptochilus hemionitideus (C.Presl) Noot.; Blumea 42: 285 (1997)
Distribution: China (Anhui, Fujian, Guizhou, Guangdong, Guangxi, Hainan, Jiangxi, Sichuan, Yunnan); Tibet; S-Japan; Ryukyu Isl.; Taiwan; Nepal; Bhutan; India (throughout, in mountaineous regions, Arunachal Pradesh, Sikkim, Darjeeling, Uttarakhand); Bangladesh; Myanmar (Kachin, Chin, etc.); Thailand; Laos; Vietnam; Philippines
36 CHI TIB 38 JAP NNS TAI 40 BAN BHU IND NEP 41 LAO MYA THA VIE 42 PHI
Remarks: Fraser-Jenkins et al. (2015) treat this only as subspecies of L. decurrens
Synonyms: hidden
= Colysis hemionitidea (C.Presl) C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6: 147 (1851)
= Colysis hemionitidea (C.Presl) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 313-331 (1933), isonym
= Drynaria hemionitidea (C.Presl) J.Sm.; J. Bot. (Hooker) 4: 61 (1841), nomen
= Leptochilus decurrens subsp. hemionitideus (C.Presl) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 63 (2008)
= Microsorum ensato-sessilifrons (Hayata) H.Itô; J. Jap. Bot. 11: 96 (1935)
= Microsorum hemionitideum (C.Presl) Copel.; Univ. Calif. Publ. Bot. 16: 111-116 (1929)
= Pleopeltis hemionitidea (C.Presl) T.Moore; Index Filic. (T. Moore) 436 (1862)
= Polypodium ensato-sessilifrons Hayata; Icon. Pl. Formosan. 6: 312, f. 126 (1915)
= Polypodium hemionitideum (C.Presl) Wall. ex Mett.; Ann. Mus. Bot. Lugduno-Batavi 4(2): 215 (1866)
= Selliguea hemionitidea C.Presl; Tent. Pterid. 216 (1836), nomen
species :
Leptochilus henryi (Baker) X.C.Zhang; Lycophytes Ferns China 654 (2012)
Distribution: China (Chongqing, Fujian, Guangxi, Guizhou, Hubei, Hunan, Jiangxi, Shaanxi, Sichuan, Yunnan, Zhejiang); Taiwan; Bangladesh; Myanmar (Kachin); Vietnam
36 CHI 38 TAI 40 BAN 41 MYA VIE
Synonyms: hidden
= Colysis henryi (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 313-331 (1933)
= Colysis lioui Ching; Fl. Fujianica 1: 602 (1982)
= Gymnogramma henryi Baker; J. Bot. 25: 171 (1887)
= Leptochilus chittagongensis Fraser-Jenk. & Gias; Taxon. Revis. Indian Subcontinental Pteridophytes 67 (2008) (fide Fraser-Jenkins et al. 2019)
= Polypodium henryi (Baker) C.Chr.; Bull. Herb. Boissier 6. 873 (1898)
= Polypodium mon-changense C.Chr.; Index Filic., Suppl. (1906-1912): 60 (1913)
= Selliguea cochlearis Christ; Bull. Acad. Géogr. Bot. Mans 142 (1907) (non Polypodium cochleare Brause (1912))
= Selliguea henryi (Baker) Christ; Bull. Herb. Boissier 6: 870 (1898)
species :
Leptochilus heterophyllus (S.K.Wu & P.K.Lôc) comb.ined.
Distribution: Vietnam
41 VIE
Remarks: provisionally accepted name
Synonyms: hidden
= Kontumia heterophylla S.K.Wu & P.K.Lôc; Novon 15(1): 245 (2005)
species :
Leptochilus insignis (Blume) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 62 (2008)
Distribution: China (Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hunan, Jiangxi, Sichuan, Yunnan); Tibet; S-Japan; Ryukyu Isl.; Taiwan; Nepal; India (Arunachal Pradesh, Sikkim, Darjeeling, NE-India, Uttarakhand); Sri Lanka; Bangladesh; Nicobar Isl.; Myanmar (Kachin, etc.); N-Thailand; ?Laos; ?Cambodia; Vietnam; peninsular Malaysia (Kelantan, Pahang); Sumatra; Borneo (Mt. Kinabalu, Sabah); Palawan; Philippines; Java; Lesser Sunda Isl. (Flores, Bali)
36 CHI TIB 38 JAP NNS TAI 40 BAN IND NEP SRL 41 MYA NCB THA VIE 42 BOR JAW LSI MLY PHI PLW SUM
Synonyms: hidden
= Colysis dilatata (Wall. ex Bedd.) J.Sm.; Hist. Fil. 101 (1875)
= Colysis insignis (Blume) J.Sm.; Hist. Fil. 101 (1875)
= Drynaria decurrens Brack.; U.S. Expl. Exped., Filic. 16: 48 (1854)
= Kaulinia dilatata (Wall. ex Bedd.) N.C.Nair & Bennet; Bull. Bot. Surv. India 11: 432 (1969 publ. 1972)
= Kaulinia dilatata (Wall. ex Bedd.) B.K.Nayar & Kaur; Comp. to Bedd. Handb. Ferns Brit. India 89 (1974), isonym
= Kaulinia hancockii (Baker) B.K.Nayar; Taxon 13: 67 (1964) (fide Fraser-Jenkins 1997)
= Kaulinia insignis (Blume) X.C.Zhang; Lycophytes Ferns China 627 (2012)
= Microsorum decurrens (Brack.) Copel.; Fern Fl. Philip. 3: 481 (1960)
= Microsorum dilatatum (Bedd.) Sledge; Bull. Brit. Mus. (Nat. Hist.) Bot. 2: 143 (1960)
= Microsorum dilatatum f. simplex (Ching) Ching; (1964), nom. inval.
= Microsorum hancockii (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 295-311 (1933) (fide Fraser-Jenkins 1997)
= Microsorum hancockii f. simplex Ching; (1959), nom. inval. (fide Fraser-Jenkins 1997)
= Microsorum insigne (Blume) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4 311 (1933), isonym
= Microsorum insigne (Blume) Copel.; Univ. Calif. Publ. Bot. 16: 111-116 (1929)
= Pleopeltis decurrens (Brack.) T.Moore; Index Filic. (T. Moore) 345 (1862)
= Pleopeltis dilatata Wall. ex Bedd.; Handb. Ferns Brit. India, t. 122 (1966)
= Pleopeltis insignis (Blume) Bedd.; Handb. Ferns Brit. India 365, t. 207 (1883)
= Pleopeltis insignis f. aperta Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 16: 29 (1914)
= Pleopeltis rivularis (Copel.) Alderw.; Bull. Dép. Agric. Indes Néerl. 27: 9 (1909)
= Polypodium acuminatum Mett.; (non Houtt. 1783 nec Schkuhr 1806)
= Polypodium anceps (Christ) C.Chr.; Index Filic. 509 (1906)
= Polypodium diffundens Kunze; Bot. Zeitung (Berlin) 422 (1846)
= Polypodium dilatatum Wall.; Numer. List no. 295 (1829), nomen, non Hoffm. (1795), nec Liebm. (1849)
= Polypodium insigne Blume; Enum. Pl. Javae 2: 137 (1828)
= Polypodium rivulare Copel.; Philipp. J. Sci. 1, Suppl. II: 163 (1906) (non Vahl (1807))
= Polypodium tridactylon Mett.; Abh. Senckenberg. Naturf. Ges. 2: 104, n. 196, t. 1, f. 39, 40 (1857) (non Hook. & Grev (1831))
= Selliguea anceps Christ; Bull. Herb. Boissier 6: 879 (1898)
species :
Leptochilus lanceolatus Fée; Acrost. 87, t. 47, f. 1 (1844)
Distribution: NE-India; ?Sri Lanka; Bangladesh
40 BAN IND
Remarks: valid species fide Fraser-Jenkins (2008)
Synonyms: hidden
= Acrostichum lanceolatum (Fée) Hook.; Sp. 6: 276 (1864) (non L. (1753))
= Acrostichum variabile var. laciniatum Hook.; Sp. Fil. 5: 255 (1864)
= Campium laciniatum (Hook.) Copel.; Philipp. J. Sci. 37: 341-402 (1928)
= Campium lanceolatum (Fée) Copel.; Philipp. J. Sci. 37: 341-402 (1928)
= Campium zeylanicum (Fée) Copel.; Philipp. J. Sci. 37: 352 (1928)
= Colysis laciniata (Hook.) Panigrahi; Pl. Sci. Res. (India) 20: 20 (1998)
= Colysis lanceolata (Fée) Panigrahi; Abstr. 23rd Ann. Conf. Khurda, Pl. Sci. Res. Orissa: 35 (1998)
= Dendroglossa lanceolata (Fée) Fée; Mém. Foug. 5. (Gen. Filic.) 81 (1852), nom. illeg.
= Dendroglossa zeylanica (Fée) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 199 (1947)
= Gymnopteris feei f. triloba Bedd.; Ferns Brit. India: t. 273 (1868)
= Gymnopteris feei var. pinnatifida Bedd.; Ferns South. India, t. 211 (1864)
= Gymnopteris feei var. triloba Bedd.; Handb. Ferns Brit. India, t. 273 (1864)
= Gymnopteris lanceolata (Fée) Bedd.; Handb. Ferns Brit. India, Suppl. 26. (1876)
= Gymnopteris variabilis var. lanceolata Bedd.; Handb. Ferns Brit. India 429 (1883)
= Leptochilus decurrens f. lanceolatus (Fée) Sledge; Ann. Mag. Nat. Hist. 12: 869 (1956)
= Leptochilus laciniatus (Hook.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 344 (1933)
= Leptochilus lanceolatus var. simplex Ching; Bull. Fan Mem. Inst. Biol. 4: 344 (1933)
= Leptochilus zeylanicus Fée; Mém. Foug. 10 (Icon. Esp. Nouv.): 8, t. 31, f. 1 (1865) (fide Fraser-Jenkins 2008)
= Paraleptochilus decurrens var. lanceolata (Fée) R.D.Dixit; Indian Fern J. 6(1-2): 148 (1989)
= Pleopeltis feei (T.Moore) Alderw.; Malayan Ferns Fern Allies: 405 (1917), nom. illeg., non (Bory) Alderw. 1909
species :
Leptochilus leveillei (Ching) X.C.Zhang; Lycophytes Ferns China 654 (2012)
Distribution: China (Fujian, Guangdong, Guangxi, Guizhou)
36 CHI
Synonyms: hidden
= Colysis leveillei (Christ) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 313-331 (1933)
= Colysis leveillei f. angusta (C.Chr.) Ching; Y. C. Wu, K. K. Wong & Pong (1933)
= Colysis leveillei f. major (C.Chr.) Ching; Y. C. Wu, K. K. Wong & Pong (1933)
= Leptochilus leveillei (Christ) X.C.Zhang & Noot.; Fl. China 2-3: 835 (2013), isonym
= Polypodium leveillei (Christ) C.Chr.; Index Filic., Suppl. (1906-1912): 60 (1913)
= Polypodium leveillei f. angusta C.Chr.; Y. C. Wu, K. K. Wong & Pong (1932)
= Polypodium leveillei f. majus C.Chr.; Y. C. Wu, K. K. Wong & Pong (1932)
= Selliguea leveillei Christ; Bull. Acad. Int. Géogr. Bot., sér. 3, 16: 236 (1906)
species :
Leptochilus longissimus (Blume) L.Y.Kuo; Taiwania 64(4): 378 (2019)
Distribution: China (Hainan, Yunnan, Hongkong); Japan; Ryukyu Isl.; Taiwan; India (Kerala, Assam); Bangladesh; Myanmar; Thailand; Vietnam; peninsular Malaysia (Perak, Selangor, Terengganu); Singapore; Sumatra; Borneo (Sabah); N-Sulawesi; Philippines; Java; Moluccas (Seram, Talaud); Bismarck Arch. (Admiralty Isl.); Palau (Babeldaob)
36 CHI 38 JAP NNS TAI 40 BAN IND 41 MYA THA VIE 42 BOR JAW MLY MOL PHI SNG SUL SUM 43 BIS 62 PLU
Remarks: traditionally often treated as Microsorum rubidum, which still might be the valid name
Conservation Status: IUCN VU for India (Fraser-Jenkins 2012). Endangered (Singapore)
Synonyms: hidden
= Colysis longissima (Blume) J.Sm.; Hist. Fil. 101 (1875)
= Drynaria rubida J.Sm.; J. Bot. (Hooker) 3: 397 (1841), nom. nud.
= Microsorum rubidum (Kunze) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 197 (1947)
= Phymatodes longissima (Blume) J.Sm.; Cat. Cult. Ferns 10 (1857)
= Phymatodes suisha-stagnalis (Hayata) H.Itô; J. Jap. Bot. 11: 98 (1935)
= Phymatosorus longissimus (Blume) Pic.Serm.; Webbia 28: 459 (1973)
= Phymatosorus longissimus var. sordidum (Copel.) Nakai
= Phymatosorus suisha-stagnalis (Hayata) Pic.Serm.; Webbia 28: 460 (1973)
= Pleopeltis longissima (Blume) Bedd.; Handb. Ferns Brit. India, Suppl. 388 (1876), isonym
= Pleopeltis longissima (Blume) Blume; Enum. Pl. Javae 2: 127 (1828)
= Pleopeltis longissima (Blume) T.Moore; Index Filic. (T. Moore) 78 (1857), isonym
= Polypodium longissimum Blume; Enum. Pl. Javae 2: 127 (1828)
= Polypodium rubidum Kunze; Bot. Zeitung (Berlin) 117 (1848) (fide Fraser-Jenkins 1997)
= Polypodium suisha-stagnale Hayata; Icon. Pl. Formosan. 6: 160 (1916), 156 (1919)
species :
Leptochilus macrophyllus (Blume) Noot.; Blumea 42: 289 (1997)
Distribution: Nicobar Isl.; Myanmar (Kachin, etc.); peninsular Thailand; peninsular Malaysia; Sumatra; Borneo (Mt. Kinabalu, Sabah, Sarawak); Sulawesi; Palawan; Philippines; Java; Lesser Sunda Isl. (Lombok, Bali); Moluccas (Seram, etc.); New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch. (Admiralty Isl., New Ireland); Solomon Isl. (Bougainville, Guadalcanal, Kolombangara, Malaita, Nggatokae, Ranongga, New Georgia, Pavuvu, Rendova, Vangunu, Vella Lavella)
41 MYA NCB THA 42 BOR JAW LSI MLY MOL NWG PHI PLW SUL SUM 43 BIS SOL
Synonyms: hidden
= Colysis acuminata Holttum; Rev. Fl. Malaya 2: 162, f. 73 (1955), nom. superfl.
= Colysis acuminata var. angustata Holttum; Rev. Fl. Malaya ed. 2, 2: 163 (1966), nom. inval.
= Colysis acuminata var. fluminalis (Ridl.) Hennipman; C. G. G. J. van Steenis, Rheophytes of the World: 157 (1981)
= Colysis campyloneuroides (Baker) Parris; Sandakania 50(4): 264 (1997), et in Malayan Nat. J. 9: 81 (1997)
= Colysis hosei (C.Chr.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 199 (1947)
= Colysis loxogrammoides (Copel.) M.G.Price; Contr. Univ. Michigan Herb. 16: 193 (1987)
= Colysis macrophylla (Blume) C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 147 (1851)
= Colysis polysora (Brause) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 199 (1947)
= Colysis regularis (Mett.) Ching; Sunyatsenia 5: 261 (1940)
= Colysis spuria (Mett.) Ching; Sunyatsenia 6: 261 (1940)
= Grammitis macrophylla Blume; Enum. Pl. Javae 2: 119 (1828)
= Gymnogramma acuminata Baker; J. Bot. 26: 326 (1888) (non Kaulf. (1824), nec Polypodium acuminatum Don (1825))
= Gymnogramma macrophylla (Blume) Hook.; Sp. Fil. 5: 159 (1864)
= Gymnogramma regularis (Mett.) Baker; Hook. & Baker, Syn. Fil. 388 (1868)
= Gymnogramma spuria (Mett.) Baker; Hook. & Baker, Syn. Fil. 388 (1868)
= Pleopeltis campyloneuroides (Baker) Alderw.; Bull. Dép. Agric. Indes Néerl. 27: 7 (1909)
= Pleopeltis hosei (C.Chr.) Alderw.; Bull. Dép. Agric. Indes Néerl. 27: 12 (1909)
= Pleopeltis interrupta (C.Chr.) Alderw.; Bull. Dép. Agric. Indes Néerl. 27: 11 (1909)
= Pleopeltis linealifolia (Rosenst.) Alderw.; Malayan Ferns Fern Allies, Suppl. 383-406 (1917)
= Pleopeltis loxogrammoides (Copel.) Alderw.; Malayan Ferns Fern Allies, Suppl. 383-406 (1917)
= Pleopeltis macrophylla (Blume) Alderw.; Bull. Dép. Agric. Indes Néerl. 27: 12 (1909)
= Pleopeltis pseudoloxogramma Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 3, 6: 218 (1922)
= Pleopeltis regularis (Mett.) Alderw.; Bull. Dép. Agric. Indes Néerl. 27: 11 (1909)
= Pleopeltis selligueoides Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 23: 18 (1916) (non Baker (1874))
= Pleopeltis spuria (Mett.) Alderw.; Bull. Dép. Agric. Indes Néerl. 27: 12 (1909)
= Polypodium campyloneuroides Baker; J. Linn. Soc., Bot. 20: 229 (1883)
= Polypodium hosei C.Chr.; Index Filic. 334 (1905), 534 (1906)
= Polypodium interruptum C.Chr.; Index Filic. 333 (1905)
= Polypodium linealifolium Rosenst.; Nova Guinea 8: 728 (1912)
= Polypodium loxogrammoides Copel.; Philipp. J. Sci., Bot. 7: 65 (1912)
= Polypodium macrophyllum (Blume) Reinw. ex Mett.; Fil. Hort. Bot. Lips. 37, t. 25, f. 22-23 (1856)
= Polypodium polysorum Brause; Bot. Jahrb. Syst. 66: 203 (1920)
= Polypodium pseudoloxogramma (Alderw.) C.Chr.; Index Filic., Suppl. Tert.: 156 (1934)
= Polypodium regulare Mett.; Ann. Mus. Bot. Lugduno-Batavi 2: 225 (1866)
= Polypodium spathulifolium Blume; Enum. Pl. Javae 2: 134 (1828)
= Polypodium spurium Mett.; Ann. Mus. Bot. Lugduno-Batavi 2: 226 (1866)
= Selliguea campyloneuroides (Baker) Bedd.; Suppl. Ferns Brit. Ind. 101 (1892)
= Selliguea macrophylla (Blume) Blume; Enum. Pl. Javae 2: addend. (1828)
= Selliguea membranacea var. fluminalis Ridl.; J. Malay. Branch Roy. Asiat. Soc. 4: 89 (1926)
= Selliguea polysora (Brause) Ching; Sunyatsenia 6: 260 (1940)
= Selliguea pseudoloxogramma (Alderw.) Ching; Sunyatsenia 5: 260 (1940)
= Syngramma campyloneuroides (Baker) Ridl.; J. Malay. Branch Roy. Asiat. Soc. 4: 105 (1926)
species :
Leptochilus mengsongensis M.X.Zhao; Phytotaxa 317(2): 144 (2017)
Distribution: China (Yunnan)
36 CHI
species :
Leptochilus minor Fée; Mém. Foug. 2 [Hist. Acrostich.]: 87, t. 25 (1845)
Distribution: NE-India (Meghalaya); Myanmar (Kachin, etc.); Thailand; Vietnam; peninsular Malaysia; Sumatra; Borneo (Sabah); Sulawesi; Philippines (Luzon)
40 IND 41 MYA THA VIE 42 BOR MLY PHI SUL SUM
Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Acrostichum minus (Fée) Mett.; Fil. Hort. Bot. Lips. 20 (1856)
= Campium minus (Fée) Copel.; Philipp. J. Sci. 37: 341-402 (1928)
= Campium minutulum (Fée) Copel.; Philipp. J. Sci. 37: 341-402 (1928)
= Colysis minor (Fée) M.G.Price; Kalikasan 3: 176 (1974)
= Colysis minutula (Fée) Panigrahi; Abstr. 23rd Ann. Conf. Khurda, Pl. Sci. Res. Orissa: 35 (1998)
= Dendroglossa minor (Fée) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 199 (1947)
= Dendroglossa minutula (Fée) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 199 (1947) (fide Fraser-Jenkins 2008)
= Dendroglossa normalis (J.Sm.) C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 149 (1851)
= Gymnopteris minor (Fée) Hook.; 2. Cent. Ferns, t. 78 (1861)
= Gymnopteris normalis J.Sm.; J. Bot. (Hooker) 3: 403 (1841), nomen
= Leptochilus minutulus Fée; Mém. Foug. 10 (Icon. Esp. Nouv.): 8, t. 31, f. 2 (1865) (fide Fraser-Jenkins 2008)
= Leptochilus normalis (J.Sm.) Copel.; Philipp. J. Sci., Bot. 8: 31 (1908)
= Leptochilus rizalianus Christ; Bull. Herb. Boissier, sér, 2, 6: 1004 (1906)
species :
Leptochilus morsei (Ching) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 2: 443 (2018)
Distribution: China; India
36 CHI 40 IND
Remarks: separate species from L. ellipticus fide Fraser-Jenkins et al. (2018)
Synonyms: hidden
= Colysis morsei (Ching) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 313-331 (1933)
= Polypodium morsei Ching; Bull. Fan Mem. Inst. Biol. Bot. 2: 17, t. 1 (1931)
species :
Leptochilus oblongus Li Bing Zhang, Liang Zhang & N.T.Lu; Phytotaxa 234(2): 195 (2015)
Distribution: Vietnam
41 VIE
species :
Leptochilus ornithopus T.Fujiw. & B.H.Quang; Phytotaxa 584(3): 153 (2023)
Distribution: Vietnam
41 VIE
species :
Leptochilus pedunculatus (Hook. & Grev.) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 62 (2008)
Distribution: China (Yunnan, Guangxi, Hainan); Nepal; Bhutan; NE-India (Assam, Meghalaya, Arunachal Pradesh, Sikkim, Darjeeling); Sri Lanka; Bangladesh; Myanmar (Kachin, etc.); Thailand (widespread); Laos; Vietnam; peninsular Malaysia (Langkawi Isl., Perak, Selangor, Kelantan, Pahang); Sumatra; Borneo (Sabah); Philippines; Java
36 CHI 40 BAN BHU IND NEP SRL 41 LAO MYA THA VIE 42 BOR JAW MLY PHI SUM
Synonyms: hidden
= Bolbitis ovata (Copel.) Ching; C. Chr., Index Filic., Suppl. Tert.: 49 (1934)
= Campium ovatum (Copel.) Copel.; Philipp. J. Sci. 37: 341-402 (1928)
= Ceterach pedunculatum Hook. & Grev.; Icon. Filic. t. 5 (1827)
= Colysis hemitoma (Hance) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 313-331 (1933)
= Colysis hemitoma f. integra Ching ex S.H.Fu
= Colysis intermedia Ching & Chu H.Wang; Acta Phytotax. Sin. 8: 170, t. 23(30) (1959)
= Colysis macrophylla var. pedunculata (Hook. & Grev.) S.Linds.; Edinburgh J. Bot. 66(2): 356 (2009)
= Colysis marginata (Meyen ex C.Presl) C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 148 (1851)
= Colysis membranacea (Blume) C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 147 (1851)
= Colysis pedunculata (Hook. & Grev.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 321 (1933)
= Colysis selliguea (Mett.) Ching; Sunyatsenia 6: 261 (1940)
= Colysis wui (C.Chr.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 313-331 (1933)
= Grammitis hamiltoniana Wall.; Numer. List no. 9 (1829), nomen
= Grammitis membranacea Blume; Enum. Pl. Javae 2: 118 (1828) (non Polypodium membranaceum Don (1825))
= Grammitis pedunculata (Hook. & Grev.) Blume; Enum. Pl. Javae 2: 118 (1828)
= Gymnogramma hamiltoniana Wall. ex Hook.; Sp. 6: 161 (1864)
= Gymnogramma membranacea (Blume) Hook.; Sp. Fil. 5: 159 (1864)
= Leptochilus macrophyllus var. pedunculatus (Hook. & Grev.) Noot.; Blumea 42: 290 (1997)
= Leptochilus ovatus Copel.; Philipp. J. Sci., Bot. 9: 229 (1914)
= Leptochilus ×hemitomus (Hance) Noot.; Blumea 42: 293 (1997)
= Paraleptochilus ovatus (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 198 (1947)
= Pleopeltis pedunculata (Hook. & Grev.) Alderw.; Bull. Dép. Agric. Indes Néerl. 27: 12 (1909)
= Pleopeltis selliguea (Mett.) Alderw.; Bull. Dép. Agric. Indes Néerl. 27: 11 (1909)
= Polypodium bonii Christ; Bull. Fan Mem. Inst. Biol. 4: 322 (1933)
= Polypodium hamiltonianum (Wall. ex Hook.) Diels; Engl. & Prantl, Nat. Pflanzenfam. 1(4): 316 (1899)
= Polypodium hemitomum Hance; J. Bot. 21: 269 (1883)
= Polypodium pedunculatum (Hook. & Grev.) Mett.; Salomon, Nom. 312 (1883)
= Polypodium selliguea Mett.; Abh. Senckenberg. Naturf. Ges. 2: 111 (1856)
= Polypodium wui (Ching) C.Chr.; Bull. Dept. Biol. Coll. Sci. Sun Yatsen Univ. 6: 17 (1933)
= Scolopendrium lanceolatum Roxb.; Calcutta J. Nat. HIst. 4: 501 (1844)
= Selliguea aliena Brack.; U.S. Expl. Exped., Filic. 16: 58 (1854)
= Selliguea hamiltoniana (Wall. ex Hook.) Bedd.; Ferns Brit. India t. 239 (1867)
= Selliguea hamiltonii (Wall. ex Hook.) C.Presl; Tent. Pterid. 216 (1836)
= Selliguea hookeri C.Presl; Tent. Pterid. 216 (1836)
= Selliguea marginata Meyen ex C.Presl; Tent. Pterid. 216 (1836)
= Selliguea membranacea (Blume) Blume; Enum. Pl. Javae 2: addend. (1828)
= Selliguea pedunculata (Hook. & Grev.) C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 146 (1851)
= Selliguea selliguea (Mett.) Christ; Bull. Herb. Boissier, sér, 2, 6: 992 (1906)
species :
Leptochilus pentaphyllus (Baker) Liang Zhang & Li Bing Zhang; Phytotaxa 374(2): 174 (2018)
Distribution: China (Guangdong, Guangxi, Guizhou, Yunnan); Tibet; Laos
36 CHI TIB 41 LAO
Remarks: frequently included in L. ellipticus s. l.
Synonyms: hidden
= Colysis elliptica var. pentaphylla (Baker) L.Shi & X.C.Zhang; Acta Phytotax. Sin. 37(1): 77 (1999)
= Colysis longisora (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 331 (1933) (fide Fraser-Jenkins 2008)
= Colysis pentaphylla (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 332 (1932)
= Gymnogramma longisora Baker; J. Bot. 28: 267 (1890) (fide Fraser-Jenkins 2008)
= Gymnogramma pentaphylla Baker; Bull. Misc. Inform. Kew 1898: 233 (1898) (non Polypodium pentaphyllum Baker (1891))
= Leptochilus ellipticus var. pentaphyllus (Baker) X.C.Zhang & Noot.; Fl. China 2-3: 837 (2013)
= Polypodium ellipticum var. pentaphyllum (Baker) C.Chr.
= Polypodium longisorum (Baker) C.Chr.; Index Filic. 541 (1906) (fide Fraser-Jenkins 2008)
species :
Leptochilus poilanei (C.Chr. & Tardieu) Liang Zhang & Li Bing Zhang; Phytotaxa 374(2): 174 (2018)
Distribution: Vietnam; etc.
41 VIE
Synonyms: hidden
= Colysis poilanei C.Chr. & Tardieu; Notul. Syst. (Paris) 8: 201 (1939)
species :
Leptochilus pothifolius (Buch.-Ham. ex D.Don) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes: 60 (2008)
Distribution: China (Chongqing, Fujian, Guangxi, Guizhou, Hainan, Hongkong, Hunan, Jiangxi, Yunnan, Zhejiang); Japan; Ryukyu Isl.; Bonin Isl. (Chichijima, Hahajima); ?Taiwan; Nepal; Bhutan; India (Arunachal Pradesh, Uttarakhand, Sikkim, Darjeeling, Meghalaya); Myanmar; Thailand; Laos; Cambodia; Vietnam; Philippines
36 CHI 38 JAP NNS OGA 40 BHU IND NEP 41 CBD LAO MYA THA VIE 42 PHI
Remarks: included sometimes as variety (or completely) in L. ellipticus, but accepted as full species by Fraser-Jenkins et al. (2008, 2015)
Synonyms: hidden
= Colysis elliptica f. simplex Ching; Bull. Fan Mem. Inst. Biol. 4: 335 (1933)
= Colysis elliptica var. furcans (Tutcher) Ching; Bull. Fan Mem. Inst. Biol. 4: 335 (1933)
= Colysis elliptica var. pothifolia (Buch.-Ham. ex D.Don) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 334 (1933)
= Colysis latiloba (Ching) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 330 (1933)
= Colysis pothifolia (Buch.-Ham. ex D.Don) C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6: 148 (1851)
= Colysis pothifolia (Buch.-Ham. ex D.Don) H.Itô; J. Jap. Bot. 11: 89 (1935), isonym
= Colysis pothifolia f. furcans (Ching) S.R.Ghosh; Pterid. Fl. E. India 499 (2004)
= Colysis pothifolia f. simplex (Ching) S.R.Ghosh; Pterid. Fl. E. India 500 (2004)
= Hemionitis pothifolia Buch.-Ham. ex D.Don; Prodr. Fl. Nepal. 13 (1825)
= Leptochilus ellipticus var. pothifolius (Buch.-Ham. ex D.Don) X.C.Zhang; Lycophytes Ferns China 653 (2012)
= Polypodium ellipticum var. furcans (Tutcher) Ching; Bull. Fan Mem. Inst. Biol. Bot. 2(2): 21 (1931)
= Polypodium ellipticum var. pothifolium (Buch.-Ham. ex D.Don) Makino; Bot. Mag. (Tokyo) 23: 72 (1909)
= Polypodium latilobum Ching; Bull. Fan Mem. Inst. Biol. Bot. 2: 21, t. 7 (1931)
= Polypodium pothifolium (Buch.-Ham. ex D.Don) Mett.; Fil. Hort. Bot. Lips. 130, t. 25, f. 21 (1856)
= Polypodium pothifolium var. furcans (Tutcher) Ching; Bull. Fan Mem. Inst. Biol. 2(2): 21 (1931)
= Selliguea pothifolia (Buch.-Ham. ex D.Don) J.Sm.; J. Bot. (Hooker) 3: 399 (1841)
species :
Leptochilus pteropus (Blume) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 62 (2008)
subspecies :
subsp. pteropus
Distribution: China (Fujian, Guangxi, Guizhou, Hainan, Hongkong, Hunan, Jiangxi, Yunnan); Ryukyu Isl. (Iriomote Isl., Ishigakijima); Taiwan; Nepal; Bhutan; India (Arunachal Pradesh, Assam, Maharashtra, Manipur, Meghalaya, Mizoram, Nagaland, Sikkim, Tripura, West Bengal); Bangladesh; Andaman Isl.; Myanmar; Thailand (widespread); Laos; Vietnam; peninsular Malaysia (Langkawi Isl., Perak, Kuala Lumpur, Terengganu, Pahang); Sumatra; Borneo (Sabah, Sarawak); Philippines; Java; Lesser Sunda Isl. (Lombok, Bali); Moluccas (Halmaheira); New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch. (Admiralty Isl.)
36 CHI 38 NNS TAI 40 BAN BHU IND NEP 41 AND LAO MYA THA VIE 42 BOR JAW LSI MLY MOL NWG PHI SUM 43 BIS
Synonyms: hidden
= Colysis pteropus (Blume) Bosman; Leiden Bot. Ser. 14: 112 (1991)
= Colysis tridactylum (Wall. ex Hook. & Grev.) J.Sm.; Hist. Fil. 101 (1875)
= Drynaria dubia J.Sm.; J. Bot. (Hooker) 3: 397 (1841)
= Drynaria tridactyla (Wall. ex Hook. & Grev.) Fée; Mém. Foug. 5 (Gen. Filic.): 271 (1852)
= Kaulinia pteropus (Blume) B.K.Nayar; Taxon 13: 67 (1964)
= Lepisorus pteropus (Blume) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 17 (1933)
= Lepisorus pteropus (Blume) R.D.Dixit; Cens. Ind. Pterid. 42 (1984), isonym
= Microsorum paucijugum (Alderw.) K.Iwats. & M.Kato; Acta Phytotax. Geobot. 32(1-4): 123 (1981)
= Microsorum pteropus (Blume) Copel.; Univ. Calif. Publ. Bot. 16: 111-116 (1929)
= Phymatodes tridactyla C.Presl; Tent. Pterid. 196 (1836), nom. nud.
= Pleopeltis aquatica (Christ) Alderw.; Malayan Ferns Fern Allies, Suppl. 383-406 (1917)
= Pleopeltis paucijuga (Alderw.) Alderw.; Bull. Dép. Agric. Indes Néerl. 27: 8 (1909)
= Pleopeltis pteropus (Blume) Bedd.; Handb. Ferns Brit. India 359 (1883), non T. Moore (1857)
= Pleopeltis pteropus (Blume) T.Moore; Index Filic. (T. Moore) 78 (1857)
= Pleopeltis tridactyla (Wall. ex Hook. & Grev.) T.Moore; Index Filic. (T. Moore) 78 (1857)
= Polypodium aquaticum Christ; Nova Guinea 8: 153 (1909)
= Polypodium paucijugum Alderw.; Bull. Dép. Agric. Indes Néerl. 18: 24 (1908)
= Polypodium pteropus Blume; Enum. Pl. Javae 2: ed. 3 (1828)
= Polypodium tridactylum Wall. ex Hook. & Grev.; Icon. Filic. 2: t. 209 (1831)
subspecies :
subsp. minor (Bedd.) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 62 (2008), comb. inval., et Indian J. Forest. 42(4): 311 (2019)
Distribution: S-India (Karnataka, Kerala, Tamil Nadu); Sri Lanka
40 IND SRL
Synonyms: hidden
= Colysis zosteriformis (Wall. ex Mett.) J.Sm.; Hist. Fil. 100 (1875)
= Kaulinia pteropus var. minor (Bedd.) B.K.Nayar & Kaur; Comp. to Bedd. Handb. Ferns Brit. India 87 (1974)
= Kaulinia zosteriformis (Wall. ex Mett.) B.K.Nayar & Kaur; Comp. to Bedd. Handb. Ferns Brit. India 87 (1974)
= Lepisorus pteropus var. minor (Bedd.) N.P.Balakr.; Bull. Bot. Surv. India 22 (1980)
= Lepisorus pteropus var. minor (Bedd.) R.D.Dixit; Cens. Ind. Pterid. 42 (1984) (as (Blume) Ching´), isonym
= Microsorum brassii Copel.; J. Arnold Arbor. 10: 181 (1929)
= Microsorum pteropus f. minus (Bedd.) Ching; Bull. Fan Mem. Inst. Biol. 4: 312 (1933)
= Microsorum pteropus var. minus (Bedd.) C.Chr. & Tardieu; Notul. Syst. (Paris) 8: 194 (1939)
= Microsorum pteropus var. minus (Bedd.) S.R.Ghosh; Pterid. Fl. E. India 565 (2004), isonym
= Microsorum pteropus var. zosteriforme (Wall. ex Mett.) S.Kaur & Subh.Chandra; Nomencl. Guide to Beddome´s Ferns South. India & Brit. India: 49 (1987)
= Microsorum zosteriforme (Wall. ex Mett.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 295-311 (1933)
= Phymatodes zosteriformis (Wall. ex Mett.) C.Presl; Tent. Pterid. 196 (1836)
= Pleopeltis pteropus var. minor Bedd.; Handb. Ferns Brit. India 361 (1833), et Ferns South. India, t. 179 (1865)
= Pleopeltis pteropus var. zosteriformis (Wall. ex Mett.) Bedd.; Handb. Ferns Brit. India: 362 (1883)
= Pleopeltis raapii (Alderw.) Alderw.; Bull. Dép. Agric. Indes Néerl. 27: 6 (1909)
= Pleopeltis zosteriformis (Wall. ex Mett.) Bedd.; Handb. Ferns Brit. India, t. 123 (1866)
= Polypodium brassii (Copel.) C.Chr.; Index Filic., Suppl. Tert.: 145 (1934)
= Polypodium micropteris Baker; Bull. Misc. Inform. Kew 1906: 14 (1906) (non C. Chr., Ind. 545)
= Polypodium pteropus var. minor (Bedd.) Y.C.Wu, K.Wong & Pong; Bull. Dept. Biol. Coll. Sci. Sun Yatsen Univ. 3: 298 (1932)
= Polypodium raapii Alderw.; Bull. Dép. Agric. Indes Néerl. 18: 23, t. 4 (1908)
= Polypodium udum Christ; Bull. Acad. Int. Géogr. Bot., sér. 3, 20: 140 (1910)
= Polypodium zosteriforme Wall.; Numer. List no. 280 (1829), nomen
= Polypodium zosteriforme Wall. ex Mett.; Abh. Senckenberg. Naturf. Ges. 2: 86 (1856)
species :
Leptochilus sarawakensis Li Bing Zhang & Liang Zhang; Phytotaxa 379(3): 269 (2018), nom. nov.
Distribution: Borneo (Sabah, Sarawak); Sulawesi; Philippines
42 BOR PHI SUL
Remarks: frequently included in L. pedunculatus
Synonyms: hidden
= Colysis fluviatilis (Lauterb.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 313-331 (1933)
= Leptochilus fluviatilis (Lauterb.) Liang Zhang & Li Bing Zhang; Phytotaxa 374(2): 173 (2018), nom. illeg., non (Hook.) C. Chr.
= Leptochilus macrophyllus var. fluviatilis (Lauterb.) Noot.; Blumea 42: 289 (1997)
= Pleopeltis fluviatilis (Lauterb.) Alderw.; Malayan Ferns Fern Allies, Suppl. 383-406 (1917)
= Polypodium fluviatile Lauterb.; Bot. Jahrb. Syst. 44. 507 (1910)
species :
Leptochilus saxicola (H.G.Zhou & Hua Li) Liang Zhang & Li Bing Zhang; Phytotaxa 374(2): 174 (2018)
Distribution: China (Yunnan, etc.)
36 CHI
Remarks: frequently included in L. pedunculatus
Synonyms: hidden
= Colysis saxicola H.G.Zhou & Hua Li; Acta Bot. Yunnan. 15(3): 253 (1993)
species :
Leptochilus scandens H.J.Wei & Yi Huang; Phytotaxa 618(1): 80 (2023)
Distribution: China (Guangdong)
36 CHI
species :
Leptochilus thwaitesianus Fée; Mém. Foug. 10 (Icon. Esp. Nouv.): 7 (1865)
Distribution: S-India
40 IND
Remarks: according to Fraser-Jenkins et al. 2019 a valid tetraploid species intermediate between L. lanceolatus and L. decurrens
Synonyms: hidden
= Colysis thwaitesianus (Fée) Rajkumar & Arunach.; Amer. Fern J. 88(4): 183 (1999)
species :
Leptochilus wrightii (Hook.) X.C.Zhang; Lycophytes Ferns China 656 (2012)
Distribution: China (Fujian, Guangxi, Hongkong, Jiangxi, Yunnan); Japan; Ryukyu Isl.; Taiwan; Vietnam
36 CHI 38 JAP NNS TAI 41 VIE
Synonyms: hidden
= Colysis hokouensis Ching; Bull. Fan Mem. Inst. Biol. Bot. 2, 1: 284 (1949)
= Colysis longifrons Tagawa; J. Jap. Bot. 14: 110 (1938)
= Colysis megalolepis Tagawa; J. Jap. Bot. 12: 492 (1936)
= Colysis subsessilifolia Ching; Wuyi Sci. J. 1: 11 (1981)
= Colysis wrightii (Hook.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 313-331 (1933)
= Colysis wrightii f. laciniata Sa.Kurata
= Gymnogramma wrightii Hook.; Sp. Fil. 5: 160 (1864)
= Leptochilus macrophyllus var. wrightii (Hook. & Baker) Noot.; Blumea 42(2): 291 (1997)
= Polypodium kusukusense Hayata; Icon. Pl. Formosan. 6: 319, f. 131 (1915)
= Polypodium wrightii (Hook.) Mett. ex Diels; Engl. & Prantl, Nat. Pflanzenfam. 1(4): 316 (1899)
= Selliguea wrightii (Hook.) J.Sm.; Hist. Fil. 102 (1875)
species :
Leptochilus ×nepalensis Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 64 (2008)
Distribution: Nepal
40 NEP
Remarks: Leptochilus pothifolius x decurrens
species :
Leptochilus ×shintenensis (Hayata) Nakaike; Checkl. Lycophytes Ferns Japan 35 (2013)
Distribution: South Korea (Cheju Isl.); Japan; Ryukyu Isl.; Taiwan; Vietnam
38 JAP KOR NNS TAI 41 VIE
Remarks: Leptochilus ellipticus x L. wrightii
Synonyms: hidden
= Colysis elliptica var. simplicifrons (Christ) Masam.; Sci. Rep. Kanazawa Univ., Biol. 2: 102 (1953)
= Colysis wrightii var. lacerata Nakai
= Colysis ×shintenensis (Hayata) H.Itô; J. Jap. Bot. 11: 90 (1935)
= Colysis ×simplicifrons (Christ) Tagawa; J. Jap. Bot. 25: 114 (1950)
= Leptochilus ×shintenensis (Hayata) X.C.Zhang & Noot.; Fl. China 2-3: 835 (2013), isonym
= Leptochilus ×simplicifrons (Christ) Nakaike; Checkl. Lycophytes Ferns Japan 36 (2013)
= Polypodium ellipticum var. simplicifrons Christ; Bull. Herb. Boissier, sér. 2, 2: 832 (1902)
= Polypodium ×shintenense Hayata; Icon. Pl. Formosan. 8: 154, f. 85-86 (1919)
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A complete, synonymic checklist of the Higher Plants of the World

Version 19.2 (April 17th, 2024)

Covering 354,696 accepted species,
8,644 hybrids, 47,758 accepted infraspecific taxa and 996,379 synonyms,
in total 1,407,477* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

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Assembled through 40 years independently from other databases

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