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Ver. 19.2
April 17th, 2024

044.0000 Family Thelypteridaceae Ching ex Pic.Serm.; Webbia 24: 711 (1970)
5-34 genera (depending on genus concept), 1100-1200 spp. New genus concept adopted here from Fawcett & Smith (2021), based on molecular results.
Species listed in database: 1180; hybrids: 38
044.2003 Genus Metathelypteris (H.Itô) Ching; Acta Phytotax. Sin. 8: 304 (1963)
Distribution: E-Asia south to Malesia, Solomon Isl., Sri Lanka, Africa, Fernando Póo, São Tomé, South Africa
Species listed in database: 17
= Thelypteris sect.Metathelypteris H.Itô in Nakai & Honda; Nov. Fl. 137 (1939)
species :
Metathelypteris adscendens (Ching) Ching; Acta Phytotax. Sin. 8: 306 (1963)
Distribution: China (Guangdong, Guangxi, Fujian); Taiwan
36 CHI 38 TAI
Synonyms: hidden
= Metathelypteris subadscendens Ching; (1976)
= Parathelypteris adscendens (Ching) X.C.Zhang & L.J.He; Glossary Lycopods Ferns 165 (2015)
= Thelypteris adscendens Ching; Bull. Fan Mem. Inst. Biol. Bot. 6: 332 (1936)
species :
Metathelypteris burrowsiorum N.R.Crouch; Bothalia 41(1): 193 (2012)
Distribution: Eswatini [Swaziland] (Lebombo Mts.)
27 SWZ
Synonyms: hidden
= Thelypteris burrowsiorum (N.R.Crouch) Christenh.; Global Fl. 1(4): 28 (2018)
species :
Metathelypteris dassanayakei (Fraser-Jenk.) Ranil; Phytotaxa 511(2): 184 (2021)
Distribution: Sri Lanka
40 SRL
Remarks: possibly a synonym of T. gracilescens (fide Fraser-Jenkins 2008)
Synonyms: hidden
= Metathelypteris flaccida var. repens Sledge; Bull. Brit. Mus. (Nat. Hist.) Bot. 8(1): 8 (1981)
= Thelypteris dassanayakei Fraser-Jenk.; New Species Syndr. Indian Pteridol. 274 (1997), nom. nov.
species :
Metathelypteris dayi (Bedd.) Holttum; Nayar & Kaur, Comp. to Bedd. Handb. Ferns Brit. India 205 (1974)
Distribution: Thailand; Vietnam; peninsular Malaysia (Fraser´s Hill, Cameron Highlands, Pahang); +Singapore; Sumatra; Borneo (Mt. Kinabalu, Sabah); Philippines (Luzon, Mindanao); W-Java; Moluccas (Seram); New Guinea (Papua New Guinea)
41 THA VIE 42 BOR JAW MLY MOL NWG PHI SNG SUM
Synonyms: hidden
= Aspidium dayi Bedd.; J. Bot. 26: 4 (1888)
= Dryopteris aureoviridis Rosenst.; Repert. Spec. Nov. Regni Veg. 13: 216 (1914)
= Dryopteris creaghii (Baker) C.Chr.; Index Filic. 258 (1905)
= Dryopteris dayi (Bedd.) C.Chr.; Index Filic. 260 (1905)
= Dryopteris diversivenosa Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 28: 23 (1918)
= Dryopteris flavovirens Rosenst.; Repert. Spec. Nov. Regni Veg. 10: 334 (1912)
= Lastrea aureoviridis (Rosenst.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 138 (1947)
= Lastrea dayi (Bedd.) Bedd.; Suppl. Ferns Brit. Ind. 54 (1892)
= Lastrea flavovirens (Rosenst.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 138 (1947)
= Nephrodium creaghii Baker; Bull. Misc. Inform. Kew 1898: 230 (1898)
= Nephrodium dayi Bedd.; J. Bot. 25: 323 (1887) (dayi)
= Thelypteris aureoviridis (Rosenst.) C.F.Reed; Phytologia 17: 262 (1968)
= Thelypteris dayi (Bedd.) B.K.Nayar & Kaur; Comp. to Bedd. Handb. Ferns Brit. India59 (1974)
= Thelypteris diversivenosa (Alderw.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 251 (1941)
= Thelypteris flavovirens (Rosenst.) C.F.Reed; Phytologia 17: 277 (1968)
species :
Metathelypteris decipiens (C.B.Clarke) Ching; Acta Phytotax. Sin. 8: 306 (1963)
Distribution: China (Hunan, Yunnan); India (Assam State, Meghalaya, +West Bengal, Arunachal Pradesh); Myanmar (Kachin, etc.)
36 CHI 40 IND 41 MYA
Remarks: kept separate from M. flaccida by Fraser-Jenkins et al. (2017)
Synonyms: hidden
= Dryopteris gracilescens var. decipiens (C.B.Clarke) Alderw.; Handb. Malayan Ferns 187 (1909)
= Lastrea gracilescens var. decipiens (C.B.Clarke) Bedd.; Handb. Ferns Brit. India, Suppl. 51 (1892)
= Metathelypteris deltoideofrons Ching ex W.M.Chu & S.G.Lu; Fl. Yunnanica 20: 720 (2006)
= Nephrodium gracilescens var. decipiens C.B.Clarke; Trans. Linn. Soc. London, Bot. 1: 541, t. 65 (1880)
= Parathelypteris deltoideofrons (Ching ex W.M.Chu & S.G.Lu) X.C.Zhang & L.J.He; Glossary Lycopods Ferns 165 (2015)
= Thelypteris decipiens (C.B.Clarke) Ching; Bull. Fan Mem. Inst. Biol. Bot. 6: 335 (1936)
= Thelypteris deltoideofrons (Ching ex W.M.Chu & S.G.Lu) Christenh.; Global Fl. 1(4): 29 (2018)
species :
Metathelypteris flaccida (Blume) Ching; Acta Phytotax. Sin. 8: 306 (1963)
Distribution: China (S-Guizhou, SW-Yunnan); Taiwan; E-Nepal; Bhutan; India (Arunachal Pradesh, Kerala, Manipur, Meghalaya, Sikkim, Tamil Nadu, West Bengal, Nagaland); Sri Lanka; Myanmar (Kachin, etc.); NE-Thailand; N-Vietnam; peninsular Malaysia (Taiping Hills, Perak, Cameron Highlands, Pahang); Sumatra (?); Borneo (Mt. Kinabalu, Sabah); Sulawesi; Java
36 CHI 38 TAI 40 BHU IND NEP SRL 41 MYA THA VIE 42 BOR JAW MLY SUL SUM
Conservation Status: IUCN R
Synonyms: hidden
= Aspidium flaccidum Blume; Enum. Pl. Javae 2: 161 (1828)
= Christella flaccida (Blume) H.Lév.; Fl. Kouy-Tchéou: 474 (1915)
= Dryopteris flaccida (Blume) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
= Lastrea flaccida (Blume) T.Moore; Index Filic. (T. Moore) 92 (1858)
= Metathelypteris krameri Sarn.Singh & Panigrahi; Ferns Fern-Allies Arunachal Pradesh 2: 716 (2005) (fide Fraser-Jenkins 2008)
= Nephrodium flaccidum (Blume) Hook.; Sp. Fil. [W. J. Hooker] 4: 133, t. 263 (1862)
= Parathelypteris flaccida (Blume) X.C.Zhang & L.J.He; Glossary Lycopods Ferns 165 (2015)
= Thelypteris flaccida (Blume) Ching; Bull. Fan Mem. Inst. Biol. Bot. 6: 336 (1936)
= Thelypteris panigrahii Christenh.; Global Fl. 1(4): 35 (2018), nom. nov. for Metathelypteris krameri Sarn. Sing & Panigrahi
species :
Metathelypteris fragilis (Baker) Holttum; J. S. Afr. Bot. 40: 127 (1974)
subspecies :
subsp. fragilis
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Dryopteris fragilis (Baker) C.Chr.; Index Filic. 266 (1905)
= Nephrodium fragile (Baker) Baker; Ann. Bot. 6: 322 (1891)
= Phegopteris fragilis (Baker) Kuhn; Decken, Reisen Ost-Afr. 38: Bot. 66 (1879)
= Polypodium fragile Baker; J. Linn. Soc., Bot. 16: 203 (1877) (non L.)
= Thelypteris fragilis (Baker) Alston; Bol. Soc. Broter., ser. 2, 30: 25 (1956)
subspecies :
subsp. guineensis Benl; Nova Hedwigia 27: 150 (1974)
Distribution: Bioko Isl. [Fernando Poo]; São Tomé
23 BIO SAO
species :
Metathelypteris glandulifera Ching; K. H. Shing, Fl. Reipubl. Popul. Sin. 4(1): 321 (1999)
Distribution: China
36 CHI
Synonyms: hidden
= Parathelypteris glandulifera (Ching ex K.H.Shing) X.C.Zhang & L.J.He; Glossary Lycopods Ferns 165 (2015)
= Thelypteris bombadilii Christenh.; Global Fl. 1(4): 27 (2018), nom. nov. for Metathelypteris glandulifera Ching ex K. H. Shing
species :
Metathelypteris glandulosa H.G.Zhou & H.Li; Acta Bot. Yunnan. 14(1): 34-36, f. 2 (1992)
Distribution: China (Guangxi)
36 CHI
Synonyms: hidden
= Parathelypteris glandulosa (H.G.Zhou & H.Li) X.C.Zhang & L.J.He; Glossary Lycopods Ferns 165 (2015)
species :
Metathelypteris gracilescens (Blume) Ching; Acta Phytotax. Sin. 8: 305 (1963)
Distribution: China (Yunnan); S-Japan (Yakushima Isl.); Taiwan; peninsular Malaysia (Cameron Highlands); Sumatra; Borneo (Mt. Kinabalu, Sabah); Sulawesi; Philippines (Luzon, Mindoro, Mindanao); W-Java; Lesser Sunda Isl. (Flores); Moluccas (Seram); New Guinea (Papua New Guinea)
36 CHI 38 JAP NNS TAI 42 BOR JAW LSI MLY MOL NWG PHI SUL SUM
Remarks: reports from India etc. refer to T. flaccida fide Fraser-Jenkins 2008
Synonyms: hidden
= Aspidium gracilescens Blume; Enum. Pl. Javae 2: 155 (1828)
= Athyrium benguetense Christ; Philipp. J. Sci., Bot. 2: 161 (1907)
= Dryopteris arisanensis Rosenst.; Hedwigia 56: 340 (1915)
= Dryopteris calva Copel.; Leafl. Philipp. Bot. 8: 808 (1910)
= Dryopteris gracilescens (Blume) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
= Dryopteris gracilescens var. calva (Copel.) C.Chr.; Index Filic., Suppl. Tert.: 82 (1934)
= Dryopteris sublaxa Hayata; Icon. Pl. Formosan. 4: 183, f. 122 (1914)
= Dryopteris vulcanica (Baker) C.Chr.; Index Filic. 300 (1905)
= Lastrea calva (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 138 (1947)
= Lastrea gracilescens (Blume) Hook.; J. Bot. 4: 338 (1857)
= Lastrea gracilescens (Blume) T.Moore; Index Filic. (T. Moore) 93 (1858), non Hook. 1857
= Lastrea hirsutipes Bedd.; Suppl. Ferns Brit. Ind. 52 (1892)
= Lastrea vulcanica (Baker) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 140 (1947)
= Metathelypteris viridescens Ching; (1976), nom. nud.
= Nephrodium glanduligerum Baker; Hook. & Baker, Syn. Fil. 262 (1867)
= Nephrodium gracilescens (Blume) C.B.Clarke; Trans. Linn. Soc. Lond., 2 Bot., 1: 513 (1880)
= Nephrodium gracilescens (Blume) Hook.; Sp. Fil. [W. J. Hooker] 4: 93 (1862) (p. p.)
= Nephrodium vulcanicum Baker; Ann. Bot. (Oxford) 5: 127 (1891)
= Parathelypteris gracilescens (Blume) X.C.Zhang & L.J.He; Glossary Lycopods Ferns 165 (2015)
= Thelypteris calva (Copel.) C.F.Reed; Phytologia 17: 266 (1968)
= Thelypteris gracilescens (Blume) Ching; Bull. Fan Mem. Inst. Biol. Bot. 6: 327 (1936)
= Thelypteris vulcanica (Baker) C.F.Reed; Phytologia 17: 324 (1968)
species :
Metathelypteris hattori (H.Itô) Ching; Acta Phytotax. Sin. 8: 306 (1963)
Distribution: China (S-Anhui, N-Fujian, N-Guangxi, Hunan, Jiangxi, SW-Sichuan, Zhejiang); South Korea; Japan
36 CHI 38 JAP KOR
Synonyms: hidden
= Parathelypteris hattori (H.Itô) X.C.Zhang & L.J.He; Glossary Lycopods Ferns 165 (2015)
= Dryopteris hattori H.Itô; Bot. Mag. (Tokyo) 49: 359, f. 1, 4(1) (1935)
= Dryopteris laxa var. dilatata Koidz.; (1932)
= Lastrea hattorii (H.Itô) Tagawa; Acta Phytotax. Geobot. 15: 14 (1953)
= Lastrea laxa var. dilatata Honda; (1957)
= Metathelypteris hattori var. nemoralis (Ching) Nakaike; New Fl. Japan, Pterid. 842 (1992)
= Metathelypteris langsunensis Ching; (1976), nom. nud.
= Metathelypteris shinganensis Ching; (1976), nom. nud.
= Thelypteris dilatata H.Itô ex Honda; (1939)
= Thelypteris hattori (H.Itô) Tagawa; Acta Phytotax. Geobot. 5: 195 (1936)
= Thelypteris hattori var. nemoralis (Ching) Sa.Kurata; Enum. Jap. Pterid. 317, 343 (1961)
= Thelypteris nemoralis Ching; Bull. Fan Mem. Inst. Biol. Bot. 6: 338 (1936)
species :
Metathelypteris laxa (Franch. & Sav.) Ching; Acta Phytotax. Sin. 8: 306 (1963)
Distribution: China (Anhui, Chongqing, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hubei, Hunan, Jiangsu, Jiangxi, Sichuan, Yunnan, Zhejiang); Japan; Taiwan; India (Assam State, Meghalaya,West Bengal); NE-Thailand
36 CHI 38 JAP TAI 40 IND 41 THA
Synonyms: hidden
= Aspidium laxum Franch. & Sav.; Enum. Pl. Jap. 2: 237, 631 (1877)
= Dryopteris laxa (Franch. & Sav.) C.Chr.; Index Filic. 274 (1905)
= Dryopteris macarthyi (Baker) C.Chr.; Index Filic. 276 (1905)
= Lastrea laxa (Franch. & Sav.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 139 (1947)
= Metathelypteris sublaxa Ching; (1976) (nom. nud., non Hayata 1914)
= Nephrodium laxum (Franch. & Sav.) Diels; Bot. Jahrb. Syst. 29: 189 (1900)
= Nephrodium macarthyi Baker; Ann. Bot. 6: 321 (1891), nom. nov. for N puberulum Baker 1875
= Nephrodium puberulum Baker; J. Bot. 13: 201 (1875), non Baker 1874
= Parathelypteris laxa (Franch. & Sav.) X.C.Zhang & L.J.He; Glossary Lycopods Ferns 166 (2015)
= Thelypteris laxa (Franch. & Sav.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 6: 333 (1936)
species :
Metathelypteris petiolulata Ching; K. H. Shing, Fl. Reipubl. Popul. Sin. 4(1): 321 (1999)
Distribution: China (NW-Jiangxi, Anhui, Fujian, S-Zhejiang)
36 CHI
Synonyms: hidden
= Metathelypteris hwangshanensis Ching; (1976), nom. nud.
= Parathelypteris petiolulata (Ching ex K.H.Shing) X.C.Zhang & L.J.He; Glossary Lycopods Ferns 166 (2015)
= Thelypteris petiolulata (Ching ex K.H.Shing) Christenh.; Global Fl. 1(4): 36 (2018)
species :
Metathelypteris singalanensis (Baker) Ching; Acta Phytotax. Sin. 8: 306 (1963)
Distribution: China (SE-Hainan); Vietnam; peninsular Malaysia (Perak, Malacca); Sumatra; Borneo; W-Java
36 CHI 41 VIE 42 BOR JAW MLY SUM
Synonyms: hidden
= Parathelypteris singalanensis (Baker ex K.H.Shing) X.C.Zhang & L.J.He; Glossary Lycopods Ferns 166 (2015)
= Dryopteris media Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 11: 9 (1913)
= Dryopteris singalanensis (Baker) C.Chr.; Index Filic. 293 (1905)
= Lastrea singalanensis (Baker) Bedd.; Suppl. Ferns Brit. Ind. 54 (1892)
= Nephrodium singalanense Baker; J. Bot. 18: 212 (1880)
= Thelypteris singalanensis (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 6: 334 (1936)
variant :
var. surbeckii Holttum; Fl. Males. Ser. 2,1(5): 382 (1982)
Distribution: C-Sumatra
42 SUM
species :
Metathelypteris uraiensis (Rosenst.) Ching; Acta Phytotax. Sin. 8: 306 (1963)
Distribution: China (N-Guangdong, W-Yunnan); S-Japan (Yakushima Isl.); Ryukyu Isl.; Taiwan; Philippines (N-Luzon)
36 CHI 38 JAP NNS TAI 42 PHI
Remarks: not in India
Synonyms: hidden
= Dryopteris hirsutisquama Hayata; Icon. Pl. Formosan. 6: 277, f. 105 (1915)
= Dryopteris uraiensis Rosenst.; Hedwigia 66: 341 (1915)
= Lastrea uraiensis (Rosenst.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 140 (1947)
= Lastrea uraiensis (Rosenst.) Tagawa; Colenso Ill. Jap. Pterid. 223 (1962)
= Macrothelypteris uraiensis (Rosenst.) Á.Löve & D.Löve; Taxon 26: 325 (1977)
= Parathelypteris uraiensis (Rosenst.) X.C.Zhang & L.J.He; Glossary Lycopods Ferns 166 (2015)
= Thelypteris uraiensis (Rosenst.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 6: 336 (1936)
variant :
var. tibetica (Ching & S.K.Wu) K.H.Shing; Fl. Reipubl. Popul. Sin. 4(1): 67 (1999)
Distribution: Tibet
36 TIB
Synonyms: hidden
= Metathelypteris tibetica Ching & S.K.Wu; Fl. Xizangica 1: 166 (1983)
= Parathelypteris uraiensis var. tibetica (Ching & S.K.Wu) X.C.Zhang & L.J.He; Glossary Lycopods Ferns 166 (2015)
species :
Metathelypteris vandervekenii Pic.Serm.; Bull. Jard. Bot. Belg. 53(1-2): 272 (1983)
Distribution: Rwanda
23 RWA
Synonyms: hidden
= Thelypteris vandervekenii (Pic.Serm.) Christenh.; Global Fl. 1(4): 39 (2018)
species :
Metathelypteris wuyishanica Ching; Wuyi Sci. J. 1: 5 (1981)
Distribution: China (Fujian, Zhejiang)
36 CHI
Synonyms: hidden
= Parathelypteris wuyishanica (Ching) X.C.Zhang & L.J.He; Glossary Lycopods Ferns 166 (2015)
= Thelypteris wuyishanica (Ching) Christenh.; Global Fl. 1(4): 40 (2018)
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A complete, synonymic checklist of the Higher Plants of the World


Version 19.2 (April 17th, 2024)

Covering 354,696 accepted species,
8,644 hybrids, 47,758 accepted infraspecific taxa and 996,379 synonyms,
in total 1,407,477* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]


With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,290 pictures of 15,532 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life