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FLORIDA MARINE RESEARCH INSTITUTE 

TECHNICAL REPORTS 

Checklists of Selected 

Shallow-Water Marine Invertebrates 

of Florida 

David K. Camp, William G. Lyons, and Thomas H. Perkins 

Florida Department of 

Environmental Protection 

ISSN 1092-194X 

FMRI Technical Report TR-3 1998

Lawton Chiles 

Governor of Florida 

Florida Department of Environmental Protection 

Virginia B. Wetherell 

Secretary 

The Florida Marine Research Institute (FMRI) is a bureau of the Florida Department of Environmental 

Protection (FDEP).The FDEP’s mission is to “protect, conserve, and manage Florida’s 

environment and natural resources.” The FMRI conducts applied research pertinent to managing 

marine-fishery resources and marine species of special concern in Florida. 

Programs at the FMRI focus on resource-management topics such as managing gamefish and 

shellfish populations, restoring depleted fish stocks and the habitats that support them, protecting 

coral reefs, preventing and mitigating oil-spill damage, protecting endangered and 

threatened species, and managing coastal-resource information. 

The FMRI publishes three series: Memoirs of the Hourglass Cruises, Florida Marine Research Publications, 

and FMRI Technical Reports. FMRI Technical Reports contain information relevant to immediate 

resource-management needs. 

Kenneth D. Haddad, Chief of Research 

Institute Editors 

Theresa M. Bert, David K. Camp, Paul R. Carlson, Mark M. Leiby, 

William G. Lyons, Anne B. Meylan, Robert G. Muller, 

Ruth O. Reese, Carmelo R. Tomas 

James F. Quinn, Jr., Science Editor 

Judith G. Leiby, Copy Editor 

Llyn C. French, Art Editor

Checklists of Selected 

Shallow-Water Marine Invertebrates 

of Florida 

David K. Camp 

William G. Lyons 

Thomas H. Perkins 


Florida Marine Research Institute 

100 Eighth Avenue Southeast 

St. Petersburg, Florida 33701 


FMRI Technical Report TR-3 

1998

Cover Photographs 

Clockwise from upper left: 

Chloeia viridis, Diadema antillarum, Stenopus hispidus, and Lithopoma tectum. 

Reprinted by permission from REEF CREATURE Identification—Florida, 

Caribbean, Bahamas, Copyright ©1992 by Paul Humann. 

Copies of this document may be obtained from 

Florida Marine Research Institute 

100 Eighth Ave. SE 

St. Petersburg, FL 33701-5095 

Attn: Librarian 

Document Citation 

Camp, D. K., W. G. Lyons, and T. H. Perkins. 1998. Checklists of selected shallow-water marine invertebrates 

of Florida. Florida Marine Research Institute Technical Report TR-3. 238 pp. 

Document Production 

This document was designed in WordPerfect ® (v. 5.1) and Microsoft ® Word (v. 6.0) and formatted 

using QuarkXPress TM (v. 4.03) on Apple Power Macintosh ® computers.The headline font is Adobe ® 

Avant Garde, the text font is Adobe ® Palatino, and the cover headline is Adobe ® Gill Sans. The 

cover and text papers are Consolidated Fortune Matte Recycled. Llyn C. French and James F. Quinn, 

Jr., of Florida Marine Research Institute, performed formatting, layout, and production for final 

film. Extra! Extra! Graphics Commercial Printers, St. Petersburg, Florida, printed the document. 

The papers used in this publication meet the minimum requirements of the American 

National Standard for Permanence of Paper for Printed Library Materials Z39.48—1992.

Table of Contents 

RATIONALE AND SCOPE OF CHECKLISTS . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 

Acknowledgments . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 

Literature Cited . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 

CHECKLIST OF SHALLOW–WATER MARINE MOLLUSCA OF FLORIDA . . . . . . . . . . . . . . . 5 

Methods . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5 

Acknowledgments . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 6 

Checklist of the Mollusca . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7 

Phylum Mollusca . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7 

Class Aplacophora . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7 

Order Pholidoskepia . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7 

Family Meiomeniidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7 

Class Polyplacophora . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7 

Order Neoloricata . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7 

Family Hanleyidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7 

Family Ischnochitonidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 7 

Family Callistoplacidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 7 

Family Chaetopleuridae . . . . . . . . . . . . . . . . . . . . . . . . . . . 7 

Family Chitonidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7 

Family Tonicellidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7 

Family Mopaliidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7 

Family Acanthochitonidae . . . . . . . . . . . . . . . . . . . . . . . . . 7 

Class Scaphopoda . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7 

Order Dentaliida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7 

Family Dentaliidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 

Family Gadilinidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 

Order Gadilida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 

Family Gadilidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 

Class Bivalvia . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 

Order Solemyoida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 

Family Solemyidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 

Order Nuculoida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 

Family Nuculidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 

Family Nuculanidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 

Order Mytiloida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 

Family Mytilidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 

Order Arcoida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 9 

Family Arcidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 9 

Family Noetiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 9 

Family Glycymerididae . . . . . . . . . . . . . . . . . . . . . . . . . . . 9 

Order Pterioida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 9 

Family Pteriidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 9 

Family Isognomonidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 9 

Family Malleidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 9 

Family Pinnidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 9 

Order Limoida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 9 

Family Limidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 9 

Order Ostreoida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 9 

Family Pectinidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 9 

Family Propeamussidae . . . . . . . . . . . . . . . . . . . . . . . . . . 10 

Family Plicatulidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 10 

i

Family Spondylidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 10 

Family Anomiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 10 

Family Ostreidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 10 

Family Gryphaeidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 10 

Order Veneroida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 10 

Family Lucinidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 10 

Family Ungulinidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 10 

Family Cyrenoididae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 11 

Family Chamidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 11 

Family Lasaeidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 11 

Family Galeommatidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 11 

Family Sportellidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 11 

Family Carditidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 11 

Family Condylocardiidae . . . . . . . . . . . . . . . . . . . . . . . . . 11 

Family Astartidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 11 

Family Crassatellidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 11 

Family Cardiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 12 

Family Mactridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 12 

Family Solenidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 12 

Family Pharidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 12 

Family Tellinidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 12 

Family Donacidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 12 

Family Psammobiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 13 

Family Semelidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 13 

Family Solecurtidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 13 

Family Dreissenidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 13 

Family Trapezidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 13 

Family Corbiculidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 13 

Family Veneridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 13 

Family Petricolidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 14 

Order Myoida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 14 

Family Myidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 14 

Family Corbulidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 14 

Family Gastrochaenidae . . . . . . . . . . . . . . . . . . . . . . . . . . 14 

Family Hiatellidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 14 

Family Pholadidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 14 

Family Teredinidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 15 

Order Pholadomyoida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 15 

Family Lyonsiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 15 

Family Pandoridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 15 

Family Thraciidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 15 

Family Periplomatidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 15 

Family Poromyidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 15 

Family Verticordiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 15 

Family Cuspidariidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 15 

Class Gastropoda . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 15 

Order Patellogastropoda . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 15 

Family Lottiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 15 

Order Archaeogastropoda . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 15 

Family Scissurellidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 15 

Family Fissurellidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 15 

Family Turbinidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 16 

Family Trochidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 16 

Family Calliostomatidae . . . . . . . . . . . . . . . . . . . . . . . . . . 16 

ii

Family Skeneidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 17 

Order Neritopsina . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 17 

Family Neritidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 17 

Family Phenacolepadidae . . . . . . . . . . . . . . . . . . . . . . . . . 17 

Order Neotaenioglossa . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 17 

Family Obtortionidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 17 

Family Litiopidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 17 

Family Cerithiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 17 

Family Batillariidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 17 

Family Potamididae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 17 

Family Planaxidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 17 

Family Modulidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 17 

Family Turritellidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 17 

Family Siliquariidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 18 

Family Littorinidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 18 

Family Skeneopsidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 18 

Family Barleeiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 18 

Family Rissoidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 18 

Family Hydrobiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 18 

Family Truncatellidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 18 

Family Assimineidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 19 

Family Elachisinidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 19 

Family Vitrinellidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 19 

Family Caecidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 19 

Family Strombidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 19 

Family Hipponicidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 20 

Family Vanikoridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 20 

Family Calyptraeidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 20 

Family Capulidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 20 

Family Xenophoridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 20 

Family Vermetidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 20 

Family Cypraeidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 20 

Family Ovulidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 20 

Family Triviidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 20 

Family Lamellariidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 21 

Family Naticidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 21 

Family Cassidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 21 

Family Tonnidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 21 

Family Ficidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 21 

Family Ranellidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 21 

Family Personidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 22 

Family Bursidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 22 

Family Cerithiopsidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 22 

Family Triphoridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 22 

Family Epitoniidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 22 

Family Janthinidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 23 

Family Aclididae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 23 

Family Eulimidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 23 

Order Neogastropoda . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 23 

Family Muricidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 23 

Family Coralliophilidae . . . . . . . . . . . . . . . . . . . . . . . . . . 24 

Family Turbinellidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 24 

Family Buccinidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 24 

iii

Family Melongenidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 24 

Family Nassariidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 24 

Family Fasciolariidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 25 

Family Columbellidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 25 

Family Volutidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 25 

Family Olividae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 25 

Family Harpidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 26 

Family Cystiscidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 26 

Family Marginellidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 26 

Family Mitridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 26 

Family Costellariidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 26 

Family Cancellariidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 26 

Family Drilliidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 27 

Family Terebridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 27 

Family Strictispiridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 27 

Family Turridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 27 

Family Conidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 27 

Subclass Heterobranchia . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 28 

Order Heterostropha . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 28 

Family Cornirostridae . . . . . . . . . . . . . . . . . . . . . . . . . . . 28 

Family Mathildidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 28 

Family Architectonicidae . . . . . . . . . . . . . . . . . . . . . . . . . . 28 

Family Rissoellidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 29 

Family Ebalidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 29 

Family Pyramidellidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 29 

Family Amathinidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 30 

Order Uncertain . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 30 

Family Acteonidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 30 

Subclass Opisthobranchia . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 30 

Order Cephalaspidea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 30 

Family Ringiculidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 30 

Family Aplustridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 30 

Family Cylichnidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 30 

Family Philinidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 30 

Family Aglajidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 30 

Family Gastropteridae . . . . . . . . . . . . . . . . . . . . . . . . . . . 30 

Family Retusidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 30 

Family Bullidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 31 

Family Haminoeidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 31 

Family Runcinidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 31 

Order Acochlidioidea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 31 

Family Microhedylidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 31 

Order Thecosomata . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 31 

Family Cavoliniidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 31 

Order Anaspidea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 31 

Family Akeridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 31 

Family Aplysiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 31 

Family Notarchidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 31 

Order Sacoglossa . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 31 

Family Volvatellidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 31 

Family Cylindrobullidae . . . . . . . . . . . . . . . . . . . . . . . . . . 31 

Family Oxynoidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 31 

Family Juliidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 31 

Family Elysiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 31 

iv

Family Polybranchiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 32 

Family Costasiellidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 32 

Family Stiligeridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 32 

Order Notaspidea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 32 

Family Umbraculidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 32 

Family Pleurobranchidae . . . . . . . . . . . . . . . . . . . . . . . . . 32 

Order Nudibranchia . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 32 

Family Corambidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 32 

Family Goniodorididae . . . . . . . . . . . . . . . . . . . . . . . . . . . 32 

Family Aegiretidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 32 

Family Polyceratidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 32 

Family Chromodorididae . . . . . . . . . . . . . . . . . . . . . . . . . 32 

Family Asteronotidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 33 

Family Dorididae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 33 

Family Dendrodorididae . . . . . . . . . . . . . . . . . . . . . . . . . . 33 

Family Phyllidiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 33 

Family Archidorididae . . . . . . . . . . . . . . . . . . . . . . . . . . . 33 

Family Discodorididae . . . . . . . . . . . . . . . . . . . . . . . . . . . 33 

Family Platydorididae . . . . . . . . . . . . . . . . . . . . . . . . . . . 33 

Family Tritoniidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 33 

Family Bornellidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 33 

Family Lomanotidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 33 

Family Scyllaeidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 33 

Family Dotoidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 34 

Family Arminidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 34 

Family Janolidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 34 

Family Flabellinidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 34 

Family Eubranchidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 34 

Family Tergipedidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 34 

Family Fionidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 34 

Family Facelinidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 34 

Family Aeolidiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 34 

Family Glaucidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 34 

Subclass Pulmonata . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 34 

Order Archaeopulmonata . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 34 

Family Ellobiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 34 

Order Basommatophora . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 35 

Family Siphonariidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 35 

Family Trimusculidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 35 

Order Systellommatophora . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 35 

Family Onchidiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 35 

Class Cephalopoda . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 35 

Order Sepioidea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 35 

Family Spirulidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 35 

Family Sepiolidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 35 

Order Teuthoidea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 35 

Family Loliginidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 35 

Order Octopoda . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 35 

Family Octopodidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 35 

Family Argonautidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 35 

Pelagic Mollusks . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 35 

Class Gastropoda . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 36 

Order Neotaenioglossa . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 36 

v

Family Atlantidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 36 

Family Carinariidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 36 

Family Pterotracheidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 36 

Order Thecosomata . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 36 

Family Limacinidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 36 

Family Cavoliniidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 36 

Family Peraclididae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 36 

Order Gymnosomata . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 36 

Family Clionidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 36 

Family Cliopsidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 37 

Family Notobranchaeidae . . . . . . . . . . . . . . . . . . . . . . . . . 37 

Family Pneumodermatidae . . . . . . . . . . . . . . . . . . . . . . . . 37 

Family Thliptodontidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 37 

Order Nudibranchia . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 37 

Family Phylliroidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 37 

References . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 37 

CHECKLIST OF SHALLOW–WATER MARINE POLYCHAETOUS ANNELIDA OF FLORIDA . . . 79 

Methods . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 79 

Acknowledgments . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 79 

Checklist of the Polychaetous Annelida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 80 

Phylum Annelida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 80 

Class Polychaeta . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 80 

Order Phyllodocida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 80 

Family Phyllodocidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 80 

Family Lacydoniidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 81 

Family Glyceridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 81 

Family Goniadidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 81 

Family Sphaerodoridae . . . . . . . . . . . . . . . . . . . . . . . . . . 81 

Family Hesionidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 81 

Family Pilargidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 82 

Family Syllidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 82 

Family Nereididae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 83 

Family Nephtyidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 84 

Family Paralacydoniidae . . . . . . . . . . . . . . . . . . . . . . . . . . 84 

Family Aphroditidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 84 

Family Polynoidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 84 

Family Acoetidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 85 

Family Eulepethidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 85 

Family Sigalionidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 85 

Family Pholoidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 85 

Family Chrysopetalidae . . . . . . . . . . . . . . . . . . . . . . . . . . 85 

Family Pisionidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 86 

Order Amphinomida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 86 

Family Amphinomidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 86 

Family Euphrosinidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 86 

Order Eunicida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 86 

Family Onuphidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 86 

Family Eunicidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 86 

Family Lumbrineridae . . . . . . . . . . . . . . . . . . . . . . . . . . . 87 

Family Oenonidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 88 

Family Dorvilleidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 88 

Order Orbiniida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 88 

Family Orbiniidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 88 

vi

Order Spionida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 88 

Family Spionidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 88 

Family Poecilochaetidae . . . . . . . . . . . . . . . . . . . . . . . . . . 89 

Family Longosomatidae . . . . . . . . . . . . . . . . . . . . . . . . . . 89 

Order Chaetopterida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 89 

Family Chaetopteridae . . . . . . . . . . . . . . . . . . . . . . . . . . . 89 

Order Magelonida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 90 

Family Magelonidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 90 

Order Cirratulida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 90 

Family Paraonidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 90 

Family Questidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 90 

Family Cirratulidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 90 

Order Ctenodrilida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 91 

Family Ctenodrilidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 91 

Order Cossurida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 91 

Family Cossuridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 91 

Order Flabelligerida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 91 

Family Flabelligeridae . . . . . . . . . . . . . . . . . . . . . . . . . . . 91 

Family Acrocirridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 91 

Family Fauveliopsidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 91 

Order Opheliida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 91 

Family Opheliidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 91 

Family Scalibregmatidae . . . . . . . . . . . . . . . . . . . . . . . . . . 91 

Order Sternaspida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 92 

Family Sternaspidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 92 

Order Capitellida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 92 

Family Capitellidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 92 

Family Maldanidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 92 

Family Arenicolidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 92 

Order Oweniida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 93 

Family Oweniidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 93 

Order Terebellida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 93 

Family Pectinariidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 93 

Family Sabellariidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 93 

Family Ampharetidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 93 

Family Trichobranchidae . . . . . . . . . . . . . . . . . . . . . . . . . . 93 

Family Terebellidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 93 

Order Sabellida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 94 

Family Sabellidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 94 

Family Serpulidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 95 

Family Spirorbidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 95 

Order Nerillida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 96 

Family Nerillidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 96 

Order Polygordiida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 96 

Family Polygordiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 96 

Order Protodrilida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 96 

Family Saccocirridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 96 

References . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 96 

CHECKLIST OF SHALLOW–WATER MARINE MALACOSTRACAN CRUSTACEA OF FLORIDA 123 

Methods . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 123 

Acknowledgments . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 124 

Checklist of the Malacostracan Crustacea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 124 

Phylum Arthropoda . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 124 

vii

Subphylum Crustacea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 124 

Class Malacostraca . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 124 

Subclass Phyllocarida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 124 

Order Leptostraca . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 124 

Family Nebaliidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 124 

Subclass Hoplocarida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 124 

Order Stomatopoda . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 124 

Suborder Unipeltata . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 124 

Superfamily Gonodactyloidea . . . . . . . . . . . . . . . . . . . . . . . 124 

Family Eurysquillidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 124 

Family Gonodactylidae . . . . . . . . . . . . . . . . . . . . . . . . . . 124 

Family Odontodactylidae . . . . . . . . . . . . . . . . . . . . . . . . . 124 

Family Pseudosquillidae . . . . . . . . . . . . . . . . . . . . . . . . . 125 

Superfamily Lysiosquilloidea . . . . . . . . . . . . . . . . . . . . . . . . 125 

Family Lysiosquillidae . . . . . . . . . . . . . . . . . . . . . . . . . . 125 

Family Nannosquillidae . . . . . . . . . . . . . . . . . . . . . . . . . 125 

Family Tetrasquillidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 125 

Superfamily Squilloidea . . . . . . . . . . . . . . . . . . . . . . . . . . . 125 

Family Squillidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 125 

Subclass Eumalacostraca . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 125 

Superorder Peracarida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 125 

Order Mysidacea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 125 

Suborder Mysida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 125 

Family Mysidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 125 

Subfamily Siriellinae . . . . . . . . . . . . . . . . . . . . . . . . . . 125 

Subfamily Gastrosaccinae . . . . . . . . . . . . . . . . . . . . . . . 125 

Subfamily Mysinae . . . . . . . . . . . . . . . . . . . . . . . . . . . 125 

Order Amphipoda . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 126 

Suborder Ingolfiellidea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 126 

Family Ingolfiellidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 126 

Suborder Gammaridea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 126 

Family Ampeliscidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 126 

Family Amphilochidae . . . . . . . . . . . . . . . . . . . . . . . . . . 126 

Family Ampithoidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 126 

Family Anamixidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 127 

Family Aoridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 127 

Family Argissidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 127 

Family Aristiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 127 

Family Bateidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 127 

Family Biancolinidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 127 

Family Cheluridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 127 

Family Colomastigidae . . . . . . . . . . . . . . . . . . . . . . . . . . 127 

Family Corophiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 128 

Subfamily Corophiinae . . . . . . . . . . . . . . . . . . . . . . . . 128 

Family Cyproideidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 128 

Family Dexaminidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 128 

Family Epimeriidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 128 

Family Eusiridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 128 

Family Gammaridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 128 

Family Haustoriidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 129 

Family Hyalellidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 129 

Family Hyalidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 129 

Family Iphimediidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 129 

Family Ischyroceridae . . . . . . . . . . . . . . . . . . . . . . . . . . . 129 

viii

Family Leucothoidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 130 

Family Liljeborgiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 130 

Family Lysianassidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 130 

Subfamily Lysianassinae . . . . . . . . . . . . . . . . . . . . . . . . 130 

Subfamily Tryphosinae . . . . . . . . . . . . . . . . . . . . . . . . 130 

Family Megaluropidae . . . . . . . . . . . . . . . . . . . . . . . . . . 130 

Family Melphidippidae . . . . . . . . . . . . . . . . . . . . . . . . . . 130 

Family Ochlesidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 130 

Family Oedicerotidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 130 

Family Phliantidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 130 

Family Phoxocephalidae . . . . . . . . . . . . . . . . . . . . . . . . . 131 

Subfamily Brolginae . . . . . . . . . . . . . . . . . . . . . . . . . . 131 

Subfamily Metharpiniinae . . . . . . . . . . . . . . . . . . . . . . 131 

Family Platyischnopidae . . . . . . . . . . . . . . . . . . . . . . . . . 131 

Family Pleustidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 131 

Subfamily Parapleustinae . . . . . . . . . . . . . . . . . . . . . . . 131 

Subfamily Stenopleustinae . . . . . . . . . . . . . . . . . . . . . . 131 

Family Podoceridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 131 

Family Pontoporeiidae . . . . . . . . . . . . . . . . . . . . . . . . . . 131 

Family Sebidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 131 

Family Stenothoidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 131 

Family Synopiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 131 

Family Talitridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 131 

Suborder Caprellidea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 132 

Family Caprellidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 132 

Family Caprellinoididae . . . . . . . . . . . . . . . . . . . . . . . . . 132 

Family Pariambidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 132 

Family Phtisicidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 132 

Subfamily Phtisicinae . . . . . . . . . . . . . . . . . . . . . . . . . 132 

Family Protellidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 132 

Order Isopoda . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 132 

Suborder Anthuridea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 132 

Family Anthuridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 132 

Family Hyssuridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 132 

Family Paranthuridae . . . . . . . . . . . . . . . . . . . . . . . . . . . 132 

Suborder Asellota . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 132 

Superfamily Gnathostenetroidoidea . . . . . . . . . . . . . . . . . . . . 132 

Family Gnathostenetroididae . . . . . . . . . . . . . . . . . . . . . . 133 

Superfamily Janiroidea . . . . . . . . . . . . . . . . . . . . . . . . . . . . 133 

Family Incertae Sedis . . . . . . . . . . . . . . . . . . . . . . . . . . . 133 

Family Janiridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 133 

Family Joeropsidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 133 

Family Munnidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 133 

Family Paramunnidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 133 

Family Pleurocopidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 133 

Family Santiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 133 

Superfamily Stenetrioidea . . . . . . . . . . . . . . . . . . . . . . . . . . 133 

Family Stenetriidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 133 

Suborder Epicaridea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 133 

Family Bopyridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 133 

Subfamily Athelginae . . . . . . . . . . . . . . . . . . . . . . . . . 133 

Subfamily Bopyrinae . . . . . . . . . . . . . . . . . . . . . . . . . . 133 

Subfamily Hemiarthrinae . . . . . . . . . . . . . . . . . . . . . . . 134 

ix

Subfamily Ioninae . . . . . . . . . . . . . . . . . . . . . . . . . . . 134 

Subfamily Pseudioninae . . . . . . . . . . . . . . . . . . . . . . . . 134 

Family Entoniscidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 134 

Suborder Flabellifera . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 134 

Family Aegidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 134 

Family Tridentellidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 135 

Family Cirolanidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 135 

Subfamily Cirolaninae . . . . . . . . . . . . . . . . . . . . . . . . . 135 

Subfamily Conilerinae . . . . . . . . . . . . . . . . . . . . . . . . . 135 

Subfamily Eurydicinae . . . . . . . . . . . . . . . . . . . . . . . . . 135 

Family Corallanidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 135 

Family Cymothoidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 135 

Family Limnoriidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 135 

Family Serolidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 136 

Family Sphaeromatidae . . . . . . . . . . . . . . . . . . . . . . . . . . 136 

Subfamily Ancininae . . . . . . . . . . . . . . . . . . . . . . . . . . 136 

Subfamily Cassidininae . . . . . . . . . . . . . . . . . . . . . . . . 136 

Subfamily Dynameninae . . . . . . . . . . . . . . . . . . . . . . . 136 

Subfamily Sphaeromatinae . . . . . . . . . . . . . . . . . . . . . . 136 

Suborder Gnathiidea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 136 

Family Gnathiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 136 

Suborder Microcerberidea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 136 

Family Microcerberidae . . . . . . . . . . . . . . . . . . . . . . . . . . 136 

Suborder Oniscidea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 136 

Infraorder Tylomorpha . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 136 

Family Tylidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 136 

Infraorder Ligiamorpha . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 136 

Section Diplocheta . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 136 

Family Ligiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 136 

Section Crinocheta . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 136 

Superfamily Oniscoidea . . . . . . . . . . . . . . . . . . . . . . . . . . . 136 

Family Halophilosciidae . . . . . . . . . . . . . . . . . . . . . . . . . 136 

Family Philosciidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 136 

Family Rhyscotidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 136 

Family Scyphacidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 136 

Suborder Valvifera . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 136 

Family Arcturidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 137 

Subfamily Arcturinae . . . . . . . . . . . . . . . . . . . . . . . . . 137 

Subfamily Edwinjoycinae . . . . . . . . . . . . . . . . . . . . . . . 137 

Family Chaetiliidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 137 

Family Idoteidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 137 

Subfamily Idoteinae . . . . . . . . . . . . . . . . . . . . . . . . . . 137 

Order Tanaidacea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 137 

Suborder Tanaidomorpha . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 137 

Superfamily Tanaoidea . . . . . . . . . . . . . . . . . . . . . . . . . . . . 137 

Family Tanaidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 137 

Subfamily Pancolinae . . . . . . . . . . . . . . . . . . . . . . . . . 137 

Superfamily Paratanaoidea . . . . . . . . . . . . . . . . . . . . . . . . . 137 

Family Anarthruridae . . . . . . . . . . . . . . . . . . . . . . . . . . . 137 

Family Leptocheliidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 137 

Family Nototanaidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 137 

Family Pseudotanaidae . . . . . . . . . . . . . . . . . . . . . . . . . . 137 

Subfamily Cryptocopinae . . . . . . . . . . . . . . . . . . . . . . . 137 

Subfamily Pseudotanainae . . . . . . . . . . . . . . . . . . . . . . 137 

x

Suborder Apseudomorpha . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 137 

Superfamily Apseudoidea . . . . . . . . . . . . . . . . . . . . . . . . . . 137 

Family Apseudidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 137 

Family Cirratodactylidae . . . . . . . . . . . . . . . . . . . . . . . . . 137 

Family Kalliapseudidae . . . . . . . . . . . . . . . . . . . . . . . . . . 137 

Family Pagurapseudidae . . . . . . . . . . . . . . . . . . . . . . . . . 138 

Family Parapseudidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 138 

Order Cumacea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 138 

Family Bodotriidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 138 

Subfamily Bodotriinae . . . . . . . . . . . . . . . . . . . . . . . . . 138 

Subfamily Mancocuminae . . . . . . . . . . . . . . . . . . . . . . 138 

Subfamily Vaunthompsoniinae . . . . . . . . . . . . . . . . . . . . 138 

Family Diastylidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 138 

Family Leuconidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 138 

Family Nannastacidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 138 

Superorder Eucarida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 138 

Order Euphausiacea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 138 

Family Euphausiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 138 

Order Decapoda . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 139 

Suborder Dendrobranchiata . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 139 

Superfamily Penaeoidea . . . . . . . . . . . . . . . . . . . . . . . . . . . 139 

Family Penaeidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 139 

Family Sicyoniidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 139 

Family Solenoceridae . . . . . . . . . . . . . . . . . . . . . . . . . . . 139 

Superfamily Sergestoidea . . . . . . . . . . . . . . . . . . . . . . . . . . 139 

Family Sergestidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 139 

Family Luciferidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 139 

Suborder Pleocyemata . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 139 

Infraorder Stenopodidea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 139 

Family Spongicolidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 139 

Family Stenopodidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 139 

Infraorder Caridea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 139 

Superfamily Pasiphaeoidea . . . . . . . . . . . . . . . . . . . . . . . . . 139 

Family Pasiphaeidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 139 

Superfamily Atyoidea . . . . . . . . . . . . . . . . . . . . . . . . . . . . 139 

Family Atyidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 139 

Superfamily Bresilioidea . . . . . . . . . . . . . . . . . . . . . . . . . . . 139 

Family Bresiliidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 139 

Superfamily Nematocarcinoidea . . . . . . . . . . . . . . . . . . . . . . 139 

Family Rhynchocinetidae . . . . . . . . . . . . . . . . . . . . . . . . . 139 

Superfamily Palaemonoidea . . . . . . . . . . . . . . . . . . . . . . . . . 140 

Family Anchistioididae . . . . . . . . . . . . . . . . . . . . . . . . . . 140 

Family Gnathophyllidae . . . . . . . . . . . . . . . . . . . . . . . . . 140 

Family Palaemonidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 140 

Subfamily Palaemoninae . . . . . . . . . . . . . . . . . . . . . . . 140 

Subfamily Pontoniinae . . . . . . . . . . . . . . . . . . . . . . . . . 140 

Superfamily Alpheoidea . . . . . . . . . . . . . . . . . . . . . . . . . . . 141 

Family Alpheidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 141 

Family Hippolytidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 141 

Family Ogyrididae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 142 

Superfamily Processoidea . . . . . . . . . . . . . . . . . . . . . . . . . . 142 

Family Processidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 142 

Superfamily Crangonoidea . . . . . . . . . . . . . . . . . . . . . . . . . 142 

xi

Family Crangonidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 142 

Infraorder Astacidea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 142 

Superfamily Enoplometopoidea . . . . . . . . . . . . . . . . . . . . . . 142 

Family Enoplometopidae . . . . . . . . . . . . . . . . . . . . . . . . . 142 

Infraorder Thalassinidea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 142 

Superfamily Callianassoidea . . . . . . . . . . . . . . . . . . . . . . . . 142 

Family Callianassidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 142 

Subfamily Callianassinae . . . . . . . . . . . . . . . . . . . . . . . 142 

Subfamily Callichirinae . . . . . . . . . . . . . . . . . . . . . . . . 142 

Subfamily Cheraminae . . . . . . . . . . . . . . . . . . . . . . . . 142 

Subfamily Eucalliinae . . . . . . . . . . . . . . . . . . . . . . . . . 142 

Family Ctenochelidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 142 

Subfamily Ctenochelinae . . . . . . . . . . . . . . . . . . . . . . . 142 

Family Laomediidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 142 

Family Upogebiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 142 

Superfamily Axioidea . . . . . . . . . . . . . . . . . . . . . . . . . . . . 143 

Family Axiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 143 

Family Micheleidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 143 

Infraorder Palinura . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 143 

Superfamily Palinuroidea . . . . . . . . . . . . . . . . . . . . . . . . . . 143 

Family Palinuridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 143 

Family Scyllaridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 143 

Subfamily Ibacinae . . . . . . . . . . . . . . . . . . . . . . . . . . . 143 

Subfamily Scyllarinae . . . . . . . . . . . . . . . . . . . . . . . . . 143 

Family Synaxidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 143 

Infraorder Anomura . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 143 

Superfamily Coenobitoidea . . . . . . . . . . . . . . . . . . . . . . . . . 143 

Family Coenobitidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 143 

Family Diogenidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 143 

Superfamily Paguroidea . . . . . . . . . . . . . . . . . . . . . . . . . . . 144 

Family Paguridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 144 

Superfamily Galatheoidea . . . . . . . . . . . . . . . . . . . . . . . . . . 144 

Family Galatheidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 144 

Family Porcellanidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 144 

Superfamily Hippoidea . . . . . . . . . . . . . . . . . . . . . . . . . . . 144 

Family Albuneidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 144 

Family Hippidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 145 

Superfamily Dromioidea . . . . . . . . . . . . . . . . . . . . . . . . . . . 145 

Family Dromiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 145 

Infraorder Brachyura . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 145 

Section Archaeobrachyura . . . . . . . . . . . . . . . . . . . . . . . . . . . 145 

Superfamily Homoloidea . . . . . . . . . . . . . . . . . . . . . . . . . . 145 

Family Cyclodorippidae . . . . . . . . . . . . . . . . . . . . . . . . . 145 

Superfamily Raninoidea . . . . . . . . . . . . . . . . . . . . . . . . . . . 145 

Family Raninidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 145 

Subfamily Ranininae . . . . . . . . . . . . . . . . . . . . . . . . . . 145 

Subfamily Notopodinae . . . . . . . . . . . . . . . . . . . . . . . . 145 

Subfamily Symethinae . . . . . . . . . . . . . . . . . . . . . . . . . 145 

Section Oxystomata . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 145 

Superfamily Dorippoidea . . . . . . . . . . . . . . . . . . . . . . . . . . 145 

Family Dorippidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 145 

Subfamily Ethusinae . . . . . . . . . . . . . . . . . . . . . . . . . . 145 

Superfamily Leucosioidea . . . . . . . . . . . . . . . . . . . . . . . . . . 145 

Family Calappidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 145 

xii

Subfamily Calappinae . . . . . . . . . . . . . . . . . . . . . . . . . 145 

Subfamily Matutinae . . . . . . . . . . . . . . . . . . . . . . . . . . 145 

Family Leucosiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 145 

Subfamily Ebaliinae . . . . . . . . . . . . . . . . . . . . . . . . . . 145 

Subfamily Iliinae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 146 

Section Oxyrhyncha . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 146 

Superfamily Majoidea . . . . . . . . . . . . . . . . . . . . . . . . . . . . 146 

Family Majidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 146 

Subfamily Inachinae . . . . . . . . . . . . . . . . . . . . . . . . . . 146 

Subfamily Epialtinae . . . . . . . . . . . . . . . . . . . . . . . . . . 146 

Subfamily Pisinae . . . . . . . . . . . . . . . . . . . . . . . . . . . 146 

Subfamily Majinae . . . . . . . . . . . . . . . . . . . . . . . . . . . 146 

Subfamily Tychinae . . . . . . . . . . . . . . . . . . . . . . . . . . . 147 

Superfamily Parthenopoidea . . . . . . . . . . . . . . . . . . . . . . . . 147 

Family Parthenopidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 147 

Subfamily Parthenopinae . . . . . . . . . . . . . . . . . . . . . . . 147 

Section Brachyrhyncha . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 147 

Superfamily Portunoidea . . . . . . . . . . . . . . . . . . . . . . . . . . 147 

Family Portunidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 147 

Subfamily Polybiinae . . . . . . . . . . . . . . . . . . . . . . . . . . 147 

Subfamily Portuninae . . . . . . . . . . . . . . . . . . . . . . . . . 147 

Superfamily Xanthoidea . . . . . . . . . . . . . . . . . . . . . . . . . . . 147 

Family Goneplacidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 147 

Family Xanthidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 148 

Superfamily Grapsidoidea . . . . . . . . . . . . . . . . . . . . . . . . . . 149 

Family Gecarcinidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 149 

Family Grapsidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 149 

Subfamily Grapsinae . . . . . . . . . . . . . . . . . . . . . . . . . . 149 

Subfamily Plagusiinae . . . . . . . . . . . . . . . . . . . . . . . . . 149 

Subfamily Sesarminae . . . . . . . . . . . . . . . . . . . . . . . . . 149 

Subfamily Varuninae . . . . . . . . . . . . . . . . . . . . . . . . . . 150 

Superfamily Pinnotheroidea . . . . . . . . . . . . . . . . . . . . . . . . 150 

Family Pinnotheridae . . . . . . . . . . . . . . . . . . . . . . . . . . . 150 

Subfamily Pinnotherinae . . . . . . . . . . . . . . . . . . . . . . . 150 

Subfamily Pinnothereliinae . . . . . . . . . . . . . . . . . . . . . . 150 

Superfamily Ocypodoidea . . . . . . . . . . . . . . . . . . . . . . . . . . 150 

Family Ocypodidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 150 

Subfamily Ocypodinae . . . . . . . . . . . . . . . . . . . . . . . . . 150 

Family Palicidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 150 

Superfamily Cryptochiroidea . . . . . . . . . . . . . . . . . . . . . . . . 150 

Family Cryptochiridae . . . . . . . . . . . . . . . . . . . . . . . . . . 150 

References . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 151 

CHECKLIST OF SHALLOW–WATER ECHINODERMATA OF FLORIDA . . . . . . . . . . . . . . . 191 

Methods . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 191 

Acknowledgments . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 191 

Checklist of the Echinodermata . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 192 

Phylum Echinodermata . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 192 

Class Crinoidea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 192 

Order Comatulida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 192 

Family Comasteridae . . . . . . . . . . . . . . . . . . . . . . . . . . . 192 

Class Asteroidea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 192 

Order Paxillosida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 192 

Family Luidiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 192 

xiii

Family Astropectinidae . . . . . . . . . . . . . . . . . . . . . . . . . . 192 

Order Valvatida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 192 

Family Asterinidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 192 

Family Goniasteridae . . . . . . . . . . . . . . . . . . . . . . . . . . . 192 

Family Ophidiasteridae . . . . . . . . . . . . . . . . . . . . . . . . . . 192 

Family Mithrodiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 192 

Family Asteropseidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 192 

Family Oreasteridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 192 

Order Spinulosida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 192 

Family Echinasteridae . . . . . . . . . . . . . . . . . . . . . . . . . . . 192 

Order Forcipulatida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 192 

Family Asteriidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 192 

Class Ophiuroidea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 192 

Order Phrynophiurida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 192 

Family Ophiomyxidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 192 

Family Gorgonocephalidae . . . . . . . . . . . . . . . . . . . . . . . . 192 

Order Ophiurida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 193 

Family Ophiuridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 193 

Family Ophiocomidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 193 

Family Ophionereididae . . . . . . . . . . . . . . . . . . . . . . . . . 193 

Family Ophiodermatidae . . . . . . . . . . . . . . . . . . . . . . . . . 193 

Family Ophiactidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 193 

Family Amphiuridae . . . . . . . . . . . . . . . . . . . . . . . . . . . 193 

Family Ophiothricidae . . . . . . . . . . . . . . . . . . . . . . . . . . 194 

Class Echinoidea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 194 

Order Cidaroida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 194 

Family Cidaridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 194 

Order Diadematoida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 194 

Family Diadematidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 194 

Order Arbacioida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 194 

Family Arbaciidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 194 

Order Temnopleuroida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 194 

Family Temnopleuridae . . . . . . . . . . . . . . . . . . . . . . . . . . 194 

Family Toxopneustidae . . . . . . . . . . . . . . . . . . . . . . . . . . 194 

Order Echinoida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 194 

Family Echinometridae . . . . . . . . . . . . . . . . . . . . . . . . . . 194 

Order Holectypoida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 194 

Family Echinoneidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 194 

Order Clypeasteroida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 194 

Family Clypeasteridae . . . . . . . . . . . . . . . . . . . . . . . . . . 194 

Family Mellitidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 194 

Order Spatangoida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 194 

Family Schizasteridae . . . . . . . . . . . . . . . . . . . . . . . . . . . 194 

Family Brissidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 194 

Class Holothuroidea . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 195 

Order Dendrochirotida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 195 

Family Cucumariidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 195 

Family Sclerodactylidae . . . . . . . . . . . . . . . . . . . . . . . . . . 195 

Family Phyllophoridae . . . . . . . . . . . . . . . . . . . . . . . . . . 195 

Order Aspidochirotida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 195 

Family Stichopodidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 195 

Family Holothuriidae . . . . . . . . . . . . . . . . . . . . . . . . . . . 195 

Order Molpadiida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 195 

Family Caudinidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 195 

xiv

Order Apodida . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 195 

Family Synaptidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 195 

Family Chiridotidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . 196 

References . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 196 

INDEX OF SCIENTIFIC NAMES . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 209 

xv

Rationale and Scope of Checklists 

David K. Camp, William G. Lyons, and Thomas H. Perkins 

The Florida Department of Environmental Protection 

is preparing to use biological criteria to assess human 

impacts on the ambient water quality of Florida’s estuaries 

and nearshore coastal zone.The use of biocriteria 

in environmental regulation and management 

is based on the principle that species composition and 

biotic abundance can serve as indicators of the environmental 

quality of a community and that these biological 

factors can be assessed to detect deviations in 

community condition and to evaluate trends in environmental 

health (Karr and Chu, 1997). To be able to 

perform such assessments, however, ecologists must 

have access to resources that will allow them to make 

accurate and consistent identifications of the biological 

components of the community. Accuracy and consistency 

of identification are integral parts of biological 

quality assurance in regulatory ecology, an enterprise 

whose results must be both scientifically valid and 

defensible.The more accurate and consistent the identifications 

of the biological components of the marine 

community are, the more likely it is that trends can be 

detected. If early warning signs of environmental impairment 

in Florida’s marine communities are to be recognized, 

identifications must be accurate at the species 

level. 

Identifying the fauna of Florida’s estuarine and 

coastal marine communities is a challenging task. 

Florida is located in the transition area between two marine 

zoogeographic zones, the tropical zone to the 

south and the temperate zone to the north, and multitudes 

of species related to each of these zones contribute 

to the state’s fauna, which also contains a significant 

endemic component. As a consequence, more 

species of marine organisms may be found in Florida 

waters than in those of any other of the continental 

United States. 

Most of Florida’s benthic marine macroinverte- 

brate species belong to four principal groups: mollusks, 

polychaetous annelids, malacostracan crustaceans, 

and echinoderms (Table 1). Species of these 

groups are prominent in epibenthic and infaunal samples 

taken in shallow waters, and they generally contribute 

the greatest numbers of species and individuals 

found in these types of samples. Depending on 

the habitat and the type of gear used, it is not unusual 

for single samples of Florida marine benthos to contain 

from 30 to more than 100 species belonging to 

these four groups. Because of this great diversity, identifying 

organisms to the species level can be difficult 

and time-consuming. 

Common impediments to the identification process 

include scattered or incomplete information on a fauna, 

out-of-date literature, and occurrences of “new” (undescribed) 

or extralimital species. Even with the recent 

decline of taxonomy as a research discipline, newly described 

marine invertebrate species are being added 

to the list of Florida’s fauna every year, and specimens 

of other still-undescribed species are present in many 

museum collections, including the Florida Marine Research 

Institute’s Marine Specimen Collection at St. Petersburg. 

Moreover, the classification of the major marine 

invertebrate groups is dynamic, changing as the 

understanding of phylogenetic relationships increases, 

so even quite familiar names may change from time to 

time. All of these factors pose problems of accuracy and 

consistency for those charged with conducting longterm, 

community-level comparisons of fauna within 

and between programs and with maintaining Florida’s 

marine biodiversity. 

The production, distribution, and periodic up-dating 

of authoritative identification guides would do 

much to improve the accuracy and consistency of identifications 

(Carney, 1996). By making these guides available 

to those engaged in faunal assessments in many 

Table 1. Numbers of families, genera, and species-level taxa of shallow-water (

Rationale and Scope Macroinvertebrate Checklists Camp et al. 1998 

separate investigations, we can help to ensure that the 

same animal will be consistently identified by the same 

name and that the name selected will be based upon 

the most accurate information. Problems posed by reclassifications 

or by the appearances of “new”(undescribed, 

recently described, or extralimital) species 

can be mitigated by periodic and timely updates of the 

guides. 

A first step in producing identification guides is to 

prepare authoritative, up-to-date checklists of all 

species of the major marine benthic macroinvertebrate 

groups known from shallow Florida waters. Such 

checklists will serve two purposes—they will provide 

ecologists with up-to-date names for most of Florida’s 

benthic macroinvertebrates, and they will provide a 

basis from which to plan the scope and production of 

illustrated guides to the marine fauna of Florida. The 

purpose of this paper is to initiate that first step. An 

electronic network (perhaps via an Internet website) 

should also be established to allow easy access to the 

checklists and to inform Florida ecologists of nomenclatural 

changes to the checklists and of associated 

identification literature. 

The checklists of mollusks, polychaetous annelids, 

malacostracan crustaceans, and echinoderms that follow 

list all species of these groups known to occur in 

Florida’s estuarine and coastal marine waters offshore 

to depths of approximately 37 meters (20 fathoms or 

120 feet).The 37-m depth was selected as a deep-water 

boundary because this depth has been shown to be an 

approximate point of transition between the faunas of 

shallow coastal zones and those of intermediate and 

deeper shelf zones, at least off western Florida (Lyons, 

1980; Lyons and Camp, 1983). Although such points of 

transition can differ between animal groups and, certainly, 

among locations in a region as large and environmentally 

diverse as Florida, we have found that 

many of the species that demonstrate fidelity to particular 

depth zones off western Florida maintain that 

fidelity in other areas of the state. As a caveat to that 

generalization, it is important to recognize the phenomenon 

of latitudinal submergence—species that 

inhabit very shallow waters in the Florida Keys and 

along southeastern Florida may be found only in deeper 

waters farther north. However, the converse is seldom 

true—benthic macroinvertebrate species that customarily 

inhabit Florida’s intermediate and deeper 

shelf communities are rarely found in inshore waters 

of Florida (although a few occur in lesser depths off 

New England). 

The checklists should not be considered a complete 

listing of all species that live in Florida waters, because 

undescribed species in museum collections are 

usually not included, and many species currently 

known only from nearby states and countries (e.g., Bahama 

Islands, Cuba) could also occur in Florida waters. 

These lists are works in progress, and they should be 

updated regularly. Compilation of this version ended 

in November 1997. 

Individual accounts of methods and acknowledgments 

related to the creation of each list are presented 

below, and the list for each group is accompanied by 

a separate compilation of pertinent literature.The references 

cited in those compilations include all literature 

sources consulted during the preparation of that 

list, as well as some other references useful in classifying 

and identifying organisms of that group. Many 

other references have also been included to guide the 

user to pertinent information on the communities in 

which the organisms live and the forces that may impact 

those organisms and their communities; however, 

the lists of references do not include all papers 

that have treated Florida’s benthic invertebrates. The 

sound management of Florida’s marine resources will 

depend not only upon the consistent and accurate 

identification of organisms but also upon an understanding 

and appreciation of the composition and dynamics 

of the communities in which these organisms 

live. 

Acknowledgments 

We thank the following people, who contributed their 

time and effort to improving the quality of this checklist. 

Staff of the library at Florida Marine Research Institute 

(FMRI) (Jan Boyett, Bob Brock, Felice Drouin, and 

James Thomas) acquired and provided copies of many 

of the publications we consulted. James F. Quinn, Jr., 

Llyn French, and Judy Leiby, of the FMRI Publications 

Production Office, edited our copy, assembled our several 

word-processing files into the finished document, 

and assisted in improving this work. Funding was provided 

by Florida Department of Environmental Protection, 

Division of Water Facilities, Bureau of Water Resources 

Protection from FY 1997 U.S. Environmental 

Protection Agency 604(b) grant monies administered 

by Mary Paulic. We are grateful to the department for 

giving us the opportunity to assemble these lists and 

associated references. Finally, we are especially grateful 

to Paul Humann, marine life author and photographer, 

for allowing several of his excellent photographs 

to grace the cover of this document. 

Literature Cited 

CARNEY, R. S. 1996. On the adequacy and improvement 

of marine benthic pre-impact surveys. Pp. 295-315 in 

Schmitt, R. J., and C.W. Osenberg (eds.), Detecting Eco- 

2 FMRI Technical Report TR-3

Camp et al. 1998 Macroinvertebrate Checklists Rationale and Scope 

logical Impacts: Concepts and Applications in Coastal 

Habitats. Academic Press, New York. xx + 401 pp. 

KARR, J. R., and E. W. CHU. 1997. Biological monitoring 

and assessment: Using multimetric indexes effectively. 

EPA 235-R97-001. University of Washington, 

Seattle. vii + 149 pp. 

LYONS, W. G. 1980. Molluscan communities of the 

West Florida Shelf. Bulletin of the American Malacological 

Union, Inc. (1979): 37-40. 

LYONS, W. G., and D. K. CAMP. 1983. Zones of faunal 

similarity within the Hourglass study area. Pp. 44-46 

in Proceedings of the Third Annual Gulf of Mexico 

Information Transfer Meeting, December 1982. United 

States Department of the Interior, Minerals Management 

Service, New Orleans. 


BLANK PAGE

Methods 

Checklist of Shallow-Water Marine Mollusca of Florida 


The initial draft of this list was based, in part, on information 

in American Seashells, second edition (Abbott, 

1974) and on lists of mollusks prepared by the Council 

of Systematic Malacologists and the American Malacological 

Union for the American Fisheries Society 

(Turgeon et al., 1988; Turgeon et al., 1998). The Florida 

list was created by selecting from those larger lists the 

estuarine and marine species known from eastern 

North America and then by reducing that set of names, 

first by deleting the names of species not known from 

Florida and then by deleting the names of several hundred 

species known only from intermediate and deepwater 

regions of the continental shelf off Florida. Many 

other published and unpublished sources on occurrences 

of mollusks in Florida within the shallow-water 

limits set for this list were then examined to confirm 

that the names remaining were appropriate for listing. 

Regional information sources were consulted for the 

Jacksonville area (H. G. Lee, unpublished manuscript); 

the Indian River Lagoon (Mikkelsen et al., 1995); the 

continental shelf off central and northeastern Florida 

(FMRI SEAMAP project, in prep.); the Oculina reefs off 

eastern Florida (Reed and Mikkelsen, 1987); shallow 

coastal waters off Hutchinson Island (Lyons, 1989); 

Peanut Island at Lake Worth Inlet (Lee, 1997b); Biscayne 

Bay and nearby environs (Voss and Voss, 1955; Mc- 

Nulty, 1961; McNulty et al., 1962; O’Gower and Wacasey, 

1967; H. B. Moore et al., 1968; and numerous papers by 

Ev. Marcus, Er. Marcus, D. R. Moore, and others); the 

Florida Keys National Marine Sanctuary (Lyons and 

Quinn, 1995); Florida Bay (Tabb and Manning, 1961; Turney 

and Perkins, 1972; FMRI Florida Bay benthic faunal 

mapping project, in prep.); southwestern Florida 

(Perry and Schwengel, 1955); Charlotte Harbor (Estevez, 

1986); Tampa Bay and Boca Ciega Bay (Hall and 

Saloman, 1975; many other papers in unpublished 

database compiled at FMRI by WGL); Crystal River region 

(Lyons et al., 1971); Cedar Key region (Dall, 1884; 

Lee, 1997a); Florida Middle Ground (Hopkins et al., 

1977; Turgeon and Lyons, 1978; unpublished species list 

by Turgeon and Lyons); Apalachee Bay region (Menzel, 

1971); St. Andrew Bay (Saloman et al., 1982), Pensacola 

Bay region (Cooley, 1978; Frey and Dörjes, 1988); 

shallow coastal waters off western and southeastern 

Florida (Godcharles and Jaap, 1971); and the continental 

shelf off western Florida (D. R. Moore, 1977; 

Bishof, 1980; Lyons, 1980; mollusks of Project Hourglass, 

FMRI unpublished). Additional records were consulted 

in a multitude of other papers. 

The names on this list are presented in a phylogenetic 

arrangement of phylum, class, subclass, order, 

and family, with genera and species arranged alphabetically 

within families.The sequence of listing follows 

that of Turgeon et al. (1998) and is an accurate representation 

of the prevalent classification theory at this 

time. The format of the list differs from that of the 

larger lists (e.g., Turgeon et al., 1998) by including a 

category for subclass and also by providing authorities 

(original authors and dates) for higher taxa (phylum, 

class, subclass, order, family, and genus). Like the 

names themselves, the authorities for higher taxa were 

extracted from many sources, among the more prominent 

of which are Abbott (1974), H. E. Vokes (1980), 

Ponder and Warén (1988),Vaught (1989), Millard (1996), 

and Coan and Scott (1997). However, it was also necessary 

to consult many other works to obtain authorities 

for some of the names. 

Three species included here are not listed by Turgeon 

et al. (1998) because a criterion for listing in the 

latter work requires prior publication of the information 

elsewhere, and there seem to be no published 

records of the following species in North America: 1) 

Solariorbis semipuncta D. R. Moore, 1965: fresh-dead 

shells and fragments among coralline sand in pockets 

atop reef structures, depth about 23-24 m, Florida Middle 

Ground, northeastern Gulf of Mexico, D. D.Turgeon 

and W G. Lyons, unpublished data; 2) Careliopsis bartschi 

(Aguayo and Rehder, 1936): Florida Bay, W. G. Lyons, 

unpublished data; 3) Bornella calcarata Mørch, 1863: 

shallow to intermediate shelf waters off northeastern 

Florida, SEAMAP benthic community study; also taken 

commonly there by a commercial shell dredger during 

the 1970s. Another species, Erycina floridanaVanatta, 

1904, described from Florida, was overlooked in other 

recent compilations of taxa (e.g., Abbott, 1974; Turgeon 

et al., 1988, 1998). 

Undescribed species are not included on this list. 

Seemingly unnamed species belonging to the families 

Triphoridae, Eulimidae, Buccinidae, Nassariidae, 

Columbellidae, Turridae, Conidae (subfamily Mangeliinae), 

and Pyramidellidae from shallow waters of 

Florida are known to various workers. However, few if 

any of these species have been figured or diagnosed 

in publications, so nothing would be served by listing 


Mollusca—W. G. Lyons Macroinvertebrate Checklists Camp et al. 1998 

unreferenced names (e.g., Nassarius sp. A). On an encouraging 

note, several of these species are the subjects 

of ongoing studies, so names for some of them may be 

available soon. 

Copies of the draft list were sent to three reviewers 

(see acknowledgments), each of whom provided 

comments that improved the final document. 

References 

The accompanying compendium of references presents 

citations for nearly 1,100 books, journal articles, 

dissertations, theses, contract reports, and popular articles 

containing information on the classification, 

identification, distribution, life history, and ecology of 

the shallow-water mollusks of Florida.The list is by no 

means complete. For example, the classic works in 

which many of Florida’s most well-known mollusks 

were described (e.g., the works of Linnaeus, Gmelin, 

Lamarck, Say, and others) have been omitted, although 

an attempt has been made to include early faunal surveys 

that documented and described Florida species 

(e.g., Conrad, 1846; Calkins, 1878; Dall, 1884). Many 

popular guides to the identification of Florida mollusks 

have been listed, but some are included principally 

for their historical value (e.g., Aldrich and Snyder, 

1936; Morris, 1947), and some of the less useful paperback 

guides to seashells have been omitted. 

Likewise, most of the popular guides to worldwide 

seashells have been omitted, even though some of 

them include information on Florida shells. Many papers 

concerned only with the mollusks of Florida’s 

deeper-water shelf and slope have also been omitted. 

Papers describing the births, romances, diets, interspecific 

associates, competitive struggles, deaths, 

and similar facets of the natural history of Florida mollusks 

have been included on the assumption that such 

information can lead to a better understanding of the 

organisms by those who would study or manage them. 

On the other hand, reports of the ordinary data of 

fisheries (effort, landings, and economics) have been 

excluded, as such information is generally available 

elsewhere. Also excluded are papers on the more complex 

aspects of molluscan biology, such as anatomy, embryology, 

and biochemistry. Thus, a paper on prey 

preferences of a predatory gastropod would be included, 

but a paper describing the cell structure of 

that predator’s stomach wall would not. 

One very useful source of information on Florida’s 

molluscan fauna is S. D. Kaicher’s Card Catalogue of 

World-Wide Shells, a series of 60 card packs published 

individually between 1973 and 1992. Each pack consists 

of approximately 108 3 × 5 cards dedicated to a single 

family (or occasionally two related families) of marine 

gastropods, and each card presents the name, range, 

diagnostic comments, and illustrations for a different 

species of that family. Nearly all of the packs contain 

cards treating species of the Florida fauna, but such information 

is readily available elsewhere for some families 

(e.g., Strombidae), so those packs are not cited 

here. In other cases, however, the information in the 

card packs is superior in accuracy and quality of illustration 

to any other information on those species, 

and an effort has been made to list the most pertinent 

of those packs. Examples of packs especially useful as 

identification aids for Florida species include those 

on Olividae (especially Olivella), Nassariidae, and Turridae. 

A few articles from shell club newsletters have 

been included as examples of this information resource. 

Such articles can provide a wealth of information 

on local faunas, and in many instances they constitute 

the only record of local occurrence for a given 

species. Many useful and informative articles have 

been published in Shell-O-Gram, the newsletter of the 

Jacksonville Shell Club, in Seafari, the now-defunct 

newsletter of the Palm Beach County Shell Club, and 

in similar publications. Some of those newsletter articles 

have been later cited in scientific reports, but this 

is not peer-reviewed literature, and such articles 

should be used with caution. On the other hand, some 

of the shell collectors who write these articles are more 

knowledgable identifiers of local fauna than are some 

of the biologists who identified mollusks for some of 

the environmental surveys cited in the present compendium. 

In fact, there seems to be an increasing tendency 

towards the publication of incorrect molluscan 

identifications in the peer-reviewed scientific literature, 

especially in papers reporting general faunal assessments, 

and some of those papers listed here contain 

remarkable errors. The best rule is to accept that information 

which seems reasonable and seek verification 

for that which does not, regardless of the source. 


Dr. Paula M. Mikkelsen, Department of Invertebrates, 

American Museum of Natural History, New York, and 

Dr. James F. Quinn, Jr., Florida Department of Environmental 

Protection, Florida Marine Research Institute, 

St. Petersburg, each reviewed the entire draft list, 

and Dr. Kerry B. Clark, Department of Biological Sciences, 

Florida Institute of Technology, Melbourne, reviewed 

sections on aplacophorans and heterobranch 

and opisthobranch gastropods. Each of the reviewers 

has extensive experience working with the marine 

molluscan fauna of Florida, and each contributed valuable 

comments that improved the list.This effort also 


Camp et al. 1998 Macroinvertebrate Checklists W. G. Lyons—Mollusca 

benefited from information provided by many others 

who contributed to or reviewed manuscript drafts of 

the American Fisheries Society list (Turgeon et al., 

1998). Notable among these are Mr. Kurt Auffenberg, 

Florida Museum of Natural History, Gainesville; Dr. 

Robert C. Bullock, University of Rhode Island, Kingston; 

Phylum Mollusca 

Class Aplacophora von Ihering, 1876 

Solenogasters and Glisten-worms 

Order Pholidoskepia Salvini-Plawén, 1978 

Family Meiomeniidae Salvini-Plawén, 1985 

Genus Meiomenia Morse, 1979 

Meiomenia arenicola Salvini-Plawén, 1985 

Class Polyplacophora J. E. Gray, 1821 

Chitons 

Order Neoloricata Bergenhayn, 1955 

Family Hanleyidae Bergenhayn, 1955 

Genus Hanleya J. E. Gray, 1857 

Hanleya tropicalis Dall, 1881 

Family Ischnochitonidae Dall, 1889 

Genus Ischnochiton J. E. Gray, 1847 

Ischnochiton dilatosculptus Kaas, 1982 

Ischnochiton erythronotus (C. B. Adams, 1845) 

Ischnochiton hartmeyeri Thiele, 1910 

Ischnochiton niveus Ferreira, 1987 

Ischnochiton papillosus (C. B. Adams, 1845) 

Ischnochiton pseudovirgatus Kaas, 1972 

Ischnochiton striolatus (J. E. Gray, 1828) 

Genus Stenoplax Carpenter in Dall, 1879 

Stenoplax bahamensis Kaas & Van Belle, 1987 

Stenoplax boogii (Haddon, 1886) 

Stenoplax floridana (Pilsbry, 1892) 

Family Callistoplacidae Pilsbry, 1893 

Genus Callistochiton Carpenter in Dall, 1879 

Callistochiton portobelensis Ferreira, 1976 

Callistochiton shuttleworthianus Pilsbry, 1893 

Family Chaetopleuridae Plate, 1899 

Genus Chaetopleura Shuttleworth, 1853 

Chaetopleura apiculata (Say, 1834) 

Checklist of the Mollusca 

Mr. Eugene V. Coan, Palo Alto, California; Dr.Terrence 

Gosliner, California Academy of Science, San Francisco; 

Dr. Harry G. Lee, Jacksonville, Florida; Dr. Gary 

McDonald, Long Marine Laboratory, Santa Cruz, California; 

and Dr. Gary Rosenberg, Academy of Natural 

Sciences of Philadelphia, Pennsylvania. 

Chaetopleura janeirensis (J. E. Gray, 1828) 

Chaetopleura staphylophera Lyons, 1985 

Family Chitonidae Rafinesque, 1815 

Genus Acanthopleura Guilding, 1829 

Acanthopleura granulata (Gmelin, 1791) 

Genus Chiton Linnaeus, 1758 

Chiton marmoratus Gmelin, 1791 

Chiton tuberculatus Linnaeus, 1758 

Chiton viridis Spengler, 1797 

Genus Tonicia J. E. Gray, 1847 

Tonicia schrammi (Shuttleworth, 1856) 

Family Tonicellidae Simroth, 1894 

Genus Lepidochitona J. E. Gray, 1821 

Lepidochitona liozonis (Dall & Simpson, 1901) 

Family Mopaliidae Dall, 1889 

Genus Ceratozona Dall, 1882 

Ceratozona squalida (C. B. Adams, 1845) 

Family Acanthochitonidae Pilsbry, 1893 

Genus Acanthochitona J. E. Gray, 1821 

Acanthochitona balesae Abbott, 1954 

Acanthochitona hemphilli (Pilsbry, 1893) 

Acanthochitona pygmaea (Pilsbry, 1893) 

Acanthochitona roseojugum Lyons, 1988 

Acanthochitona zebra Lyons, 1988 

Genus Americhiton Watters, 1990 

Americhiton andersoni (Watters, 1981) 

Genus Cryptoconchus Burrow, 1815 

Cryptoconchus floridanus (Dall, 1889) 

Class Scaphopoda Bronn, 1862 

Tuskshells and Toothshells 

Order Dentaliida Da Costa, 1776 



Family Dentaliidae J. E. Gray, 1834 

Genus Antalis H. and A. Adams, 1854 

Antalis antillarum (d’Orbigny, 1842) 

Antalis pilsbryi (Rehder, 1942) 

Antalis taphrium (Dall, 1889) 

Genus Dentalium Linnaeus, 1758 

Dentalium americanum Chenu, 1843 

Dentalium laqueatum A. E.Verrill, 1885 

Genus Graptacme Pilsbry & Sharp, 1897 

Graptacme calamus (Dall, 1889) 

Graptacme eborea (Conrad, 1846) 

Graptacme semistriolata (Guilding, 1834) 

Family Gadilinidae Chistikov, 1975 

Genus Episiphon Pilsbry & Sharp, 1897 

Episiphon sowerbyi (Guilding, 1834) 

Order Gadilida Starobogatov, 1974 

Family Gadilidae Stoliczka, 1868 

Genus Gadila J. E. Gray, 1847 

Gadila mayori (J. B. Henderson, 1920) 

Genus Polyschides Pilsbry & Sharp, 1897 

Polyschides agassizii (Dall, 1881) 

Polyschides carolinensis (Bush, 1895) 

Polyschides quadridentatus (Dall, 1881) 

Polyschides tetrodon (Pilsbry & Sharp, 1898) 

Class Bivalvia Linnaeus, 1758 

Mussels, Scallops, Oysters, Clams, etc. 

Order Solemyoida Dall, 1889 

Family Solemyidae J. E. Gray, 1840 

Genus Solemya Lamarck, 1818 

Solemya occidentalis Deshayes, 1857 

Solemya velum Say, 1822 

Order Nuculoida Dall, 1889 

Family Nuculidae J. E. Gray, 1824 

Genus Nucula Lamarck, 1799 

Nucula calcicola D. R. Moore, 1977 

Nucula proxima Say, 1822 

Family Nuculanidae H. & A. Adams, 1858 

Genus Nuculana Link, 1807 

Nuculana acuta (Conrad, 1832) 

Nuculana concentrica (Say, 1824) 

Order Mytiloida Férussac, 1822 

Family Mytilidae Rafinesque, 1815 

Genus Amygdalum Mühlfeld, 1811 

Amygdalum papyrium (Conrad, 1846) 

Genus Botula Mørch, 1853 

Botula fusca (Gmelin, 1791) 

Genus Brachidontes Swainson, 1840 

Brachidontes domingensis (Lamarck, 1819) 

Brachidontes exustus (Linnaeus, 1758) 

Brachidontes modiolus (Linnaeus, 1767) 

Genus Crenella Brown, 1827 

Crenella decussata (Montagu, 1808) 

Genus Dacyridium Torell, 1859 

Dacrydium elegantulum hendersoni Salas & Gofas, 

1997 

Genus Geukensia Poel, 1959 

Geukensia demissa (Dillwyn, 1817) 

Geukensia granosissima (G. B. Sowerby III, 1914) 

Genus Gregariella Monterosato, 1884 

Gregariella coralliophaga (Gmelin, 1791) 

Genus Ischadium Jukes-Brown, 1905 

Ischadium recurvum (Rafinesque, 1820) 

Genus Lioberus Dall, 1898 

Lioberus castaneus (Say, 1822) 

Genus Lithophaga Röding, 1798 

Lithophaga antillarum (d’Orbigny, 1842) 

Lithophaga aristata (Dillwyn, 1817) 

Lithophaga bisulculata (d’Orbigny, 1842) 

Lithophaga nigra (d’Orbigny, 1842) 

Genus Modiolus Lamarck, 1799 

Modiolus americanus (Leach, 1815) 

Modiolus modiolus squamosus Beauperthuy, 1967 

Genus Musculus Röding, 1798 

Musculus lateralis (Say, 1822) 

Genus Mytilus Linnaeus, 1758 

Mytilus edulis Linnaeus, 1758 



Order Arcoida Stoliczka, 1871 

Family Arcidae Lamarck, 1809 

Genus Acar J. E. Gray, 1842 

Acar domingensis (Lamarck, 1819) 

Genus Anadara J. E. Gray, 1847 

Anadara brasiliana (Lamarck, 1819) 

Anadara floridana (Conrad, 1869) 

Anadara notabilis (Röding, 1798) 

Anadara ovalis (Bruguière, 1789) 

Anadara transversa (Say, 1822) 

Genus Arca Linnaeus, 1758 

Arca imbricata Bruguière, 1789 

Arca zebra (Swainson, 1833) 

Genus Barbatia J. E. Gray, 1842 

Barbatia cancellaria (Lamarck, 1819) 

Barbatia candida (Helbling, 1779) 

Barbatia tenera (C. B. Adams, 1845) 

Family Noetiidae Stewart, 1930 

Genus Arcopsis von Koenen, 1885 

Arcopsis adamsi (Dall, 1886) 

Genus Noetia J. E. Gray, 1857 

Noetia ponderosa (Say, 1822) 

Family Glycymerididae Newton, 1916 

Genus Glycymeris da Costa, 1778 

Glycymeris americana (DeFrance, 1829) 

Glycymeris decussata (Linnaeus, 1758) 

Glycymeris pectinata (Gmelin, 1791) 

Glycymeris spectralis Nicol, 1952 

Glycymeris undata (Linnaeus, 1758) 

Order Pterioida Newell, 1965 

Family Pteriidae J. E. Gray, 1847 

Genus Pinctada Röding, 1798 

Pinctada imbricata Röding, 1798 

Pinctada margaritifera (Linnaeus, 1758) 

Genus Pteria Scopoli, 1777 

Pteria colymbus (Röding, 1798) 

Pteria longisquamosa (Dunker, 1852) 

Family Isognomonidae Woodring, 1925 

Genus Isognomon Lightfoot, 1786 

Isognomon alatus (Gmelin, 1791) 

Isognomon bicolor (C. B. Adams, 1845) 

Isognomon radiatus (Anton, 1839) 

Family Malleidae Lamarck, 1819 

Genus Malleus Lamarck, 1799 

Malleus candeanus (d’Orbigny, 1842) 

Family Pinnidae Leach, 1819 

Genus Atrina J. E. Gray, 1842 

Atrina rigida (Lightfoot, 1786) 

Atrina seminuda (Lamarck, 1819) 

Atrina serrata (Sowerby, 1825) 

Genus Pinna Linnaeus, 1758 

Pinna carnea Gmelin, 1791 

Order Limoida Waller, 1978 

Family Limidae Rafinesque, 1815 

Genus Ctenoides Mørch, 1853 

Ctenoides scabra (Born, 1778) 

Ctenoides tenera (Sowerby, 1843, non Turton, 1822) 

Genus Lima Bruguière, 1797 

Lima lima (Linnaeus, 1758) 

Genus Limaria Link, 1807 

Limaria locklini (McGinty, 1955) 

Limaria pellucida (C. B. Adams, 1846) 

Genus Limatula S.V. Wood, 1839 

Limatula hendersoni Olsson & McGinty, 1958 

Limatula subauriculata (Montagu, 1808) 

Order Ostreoida Férussac, 1822 

Family Pectinidae Rafinesque, 1815 

Genus Argopecten Monterosato, 1889 

Argopecten gibbus (Linnaeus, 1758) 

Argopecten irradians (Lamarck, 1819) 

Argopecten lineolaris (Lamarck, 1819) 

Argopecten nucleus (Born, 1778) 

Genus Bractechlamys Iredale, 1939 

Bractechlamys antillarum (Récluz, 1853) 

Genus Caribachlamys Waller, 1993 

Caribachlamys imbricata (Gmelin, 1791) 

Caribachlamys mildredae (Bayer, 1941) 

Caribachlamys ornata (Lamarck, 1819) 

Caribachlamys sentis (Reeve, 1853) 



Genus Euvola Dall, 1898 

Euvola raveneli (Dall, 1898) 

Euvola ziczac (Linnaeus, 1758) 

Genus Laevichlamys Waller, 1993 

Laevichlamys multisquamata (Dunker, 1864) 

Genus Lindapecten Petuch, 1995 

Lindapecten exasperatus (G. B. Sowerby II, 1842) 

Lindapecten muscosus (W. Wood, 1828) 

Genus Nodipecten Dall, 1898 

Nodipecten nodosus (Linnaeus, 1758) 

Genus Spathochlamys Waller, 1993 

Spathochlamys benedicti (A. E.Verrill & Bush, 

1897) 

Family Propeamussidae Abbott, 1954 

Genus Cyclopecten A. E.Verrill, 1897 

Cyclopecten nanus A. E.Verrill & Bush, 1897 

Family Plicatulidae Watson, 1930 

Genus Plicatula Lamarck, 1801 

Plicatula gibbosa Lamarck, 1801 

Family Spondylidae J. E. Gray, 1826 

Genus Spondylus Linnaeus, 1758 

Spondylus americanus Hermann, 1781 

Spondylus ictericus Reeve, 1856 

Family Anomiidae Rafinesque, 1815 

Genus Anomia Linnaeus, 1758 

Anomia simplex d’Orbigny, 1842 

Genus Pododesmus Philippi, 1837 

Pododesmus rudis (Broderip, 1834) 

Family Ostreidae Rafinesque, 1815 

Genus Crassostrea Sacco, 1897 

Crassostrea virginica (Gmelin, 1791) 

Genus Cryptostrea Harry, 1985 

Cryptostrea permollis (G. B. Sowerby II, 1871) 

Genus Dendrostrea Sowerby, 1839 

Dendostrea frons (Linnaeus, 1758) 

Genus Ostrea Linnaeus, 1758 

Ostrea equestris Say, 1834 

Genus Teskeyostrea Harry, 1985 

Teskeyostrea weberi (Olsson, 1951) 

Family Gryphaeidae Vyalov, 1936 

Genus Parahyotissa Harry, 1985 

Parahyotissa mcgintyi Harry, 1985 

Order Veneroida H. & A. Adams, 1856 

Family Lucinidae Fleming, 1828 

Genus Anodontia Link, 1807 

Anodontia alba Link, 1807 

Anodontia philippiana (Reeve, 1850) 

Genus Codakia Scopoli, 1777 

Codakia orbicularis (Linnaeus, 1758) 

Codakia orbiculata (Montagu, 1808) 

Codakia pectinella (C. B. Adams, 1852) 

Genus Divalinga Chavan, 1951 

Divalinga quadrisulcata (d’Orbigny, 1842) 

Genus Divaricella von Martens, 1880 

Divaricella dentata (W. Wood, 1815) 

Genus Lucina Bruguière, 1797 

Lucina amianta (Dall, 1901) 

Lucina floridana Conrad, 1833 

Lucina keenae Chavan, 1971 

Lucina leucocyma Dall, 1886 

Lucina pectinata (Gmelin, 1791) 

Lucina pensylvanica (Linnaeus, 1758) 

Lucina sombrerensis Dall, 1886 

Lucina trisulcata Conrad, 1841 

Genus Lucinisca Dall, 1901 

Lucinisca muricata (Spengler, 1798) 

Lucinisca nassula (Conrad, 1846) 

Genus Parvilucina Dall, 1901 

Parvilucina costata (d’Orbigny, 1842) 

Parvilucina multilineata (Tuomey & Holmes, 1857) 

Family Ungulinidae H. & A. Adams, 1856 

Genus Diplodonta Bronn, 1831 

Diplodonta candeana (d’Orbigny, 1842) 

Diplodonta notata Dall & Simpson, 1901 

Diplodonta nucleiformis (Wagner, 1838) 

Diplodonta punctata (Say, 1822) 

Diplodonta semiaspera Philippi, 1836 



Diplodonta soror (C. B. Adams, 1852) 

Diplodonta subglobosa (C. B. Adams, 1852) 

Diplodonta venezuelensis (Dunker, 1848) 

Family Cyrenoididae H. & A. Adams, 1857 

Genus Cyrenoida Joannis, 1835 

Cyrenoida floridana (Dall, 1896) 

Family Chamidae Lamarck, 1809 

Genus Arcinella Schumacher, 1817 

Arcinella cornuta Conrad, 1866 

Genus Chama Linnaeus, 1758 

Chama congregata Conrad, 1833 

Chama florida Lamarck, 1819 

Chama macerophylla Gmelin, 1791 

Chama sarda Reeve, 1847 

Chama sinuosa Broderip, 1835 

Genus Pseudochama Odhner, 1917 

Pseudochama inezae Bayer, 1943 

Pseudochama radians (Lamarck, 1819) 

Family Lasaeidae J. E. Gray, 1842 

Genus Aligena H. C. Lea, 1843 

Aligena texasiana Harry, 1969 

Genus Erycina Lamarck, 1805 

Erycina floridana Vanatta, 1904 

Genus Lasaea Brown, 1827 

Lasaea adansoni (Gmelin, 1791) 

Genus Mysella Angas, 1877 

Mysella planulata (Stimpson, 1851) 

Genus Orobitella Dall, 1900 

Orobitella floridana (Dall, 1899) 

Orobitella limpida (Dall, 1899) 

Genus Parabornia Boss, 1965 

Parabornia squillina Boss, 1965 

Genus Pythinella Dall, 1899 

Pythinella cuneata (A. E.Verrill & Bush, 1898) 

Family Galeommatidae J. E. Gray, 1840 

Genus Divariscintilla A. W. B. Powell, 1932 

Divariscintilla cordiformis P. M. Mikkelsen & 

Bieler, 1992 

Divariscintilla luteocrinita P. M. Mikkelsen & 

Bieler, 1992 

Divariscintilla octotentaculata P. M. Mikkelsen & 

Bieler, 1992 

Divariscintilla troglodytes P. M. Mikkelsen & 

Bieler, 1989 

Divariscintilla yoyo P. M. Mikkelsen & Bieler, 1989 

Family Sportellidae Dall, 1899 

Genus Basterotia Hoernes, 1859 

Basterotia elliptica (Récluz, 1850) 

Basterotia quadrata (Hinds, 1843) 

Genus Ensitellops Olsson & Harbison, 1953 

Ensitellops protextus (Conrad, 1841) 

Family Carditidae Fleming, 1828 

Genus Carditamera Conrad, 1838 

Carditamera floridana Conrad, 1838 

Genus Glans Mühlfeld, 1811 

Glans dominguensis (d’Orbigny, 1842) 

Genus Pleuromeris Conrad, 1867 

Pleuromeris tridentata (Say, 1826) 

Genus Pteromeris Conrad, 1862 

Pteromeris perplana (Conrad, 1841) 

Family Condylocardiidae Bernard, 1897 

Genus Carditopsis E. A. Smith, 1881 

Carditopsis smithii (Dall, 1896) 

Genus Cuna Hedley, 1902 

Cuna dalli Vanatta, 1904 

Family Astartidae d’Orbigny, 1844 

Genus Astarte Sowerby, 1816 

Astarte nana Dall, 1886 

Family Crassatellidae Férussac, 1822 

Genus Crassinella Guppy, 1874 

Crassinella dupliniana (Dall, 1903) 

Crassinella lunulata (Conrad, 1834) 

Crassinella martinicensis (d’Orbigny, 1842) 

Genus Eucrassatella Iredale, 1924 

Eucrassatella speciosa (A. Adams, 1852) 



Family Cardiidae Lamarck, 1809 

Genus Americardia Stewart, 1930 

Americardia guppyi (Thiele, 1910) 

Americardia media (Linnaeus, 1758) 

Genus Dinocardium Dall, 1900 

Dinocardium robustum (Lightfoot, 1786) 

Genus Laevicardium Swainson, 1840 

Laevicardium laevigatum (Linnaeus, 1758) 

Laevicardium mortoni (Conrad, 1831) 

Laevicardium pictum (Ravenel, 1861) 

Laevicardium sybariticum (Dall, 1886) 

Genus Papyridea Swainson, 1840 

Papyridea semisulcata (J. E. Gray, 1825) 

Papyridea soleniformis (Bruguière, 1789) 

Genus Trachycardium Mørch, 1853 

Trachycardium egmontianum (Shuttleworth, 1856) 

Trachycardium magnum (Linnaeus, 1758) 

Trachycardium muricatum (Linnaeus, 1758) 

Family Mactridae Lamarck, 1809 

Genus Anatina Schumacher, 1817 

Anatina anatina (Spengler, 1802) 

Genus Mactrotoma Dall, 1894 

Mactrotoma fragilis (Gmelin, 1791) 

Genus Mulinia J. E. Gray, 1837 

Mulinia lateralis (Say, 1822) 

Genus Raeta J. E. Gray, 1853 

Raeta plicatella (Lamarck, 1818) 

Genus Rangia Desmoulins, 1832 

Rangia cuneata (Sowerby, 1831) 

Genus Spisula J. E. Gray, 1837 

Spisula raveneli (Conrad, 1831) 

Family Solenidae Lamarck, 1809 

Genus Solen Linnaeus, 1758 

Solen viridis Say, 1822 

Family Pharidae H. & A. Adams, 1858 

Genus Ensis Schumacher, 1817 

Ensis megistus Pilsbry & McGinty, 1943 

Ensis minor Dall, 1900 

Family Tellinidae de Blainville, 1814 

Genus Cymatoica Dall, 1889 

Cymatoica orientalis (Dall, 1890) 

Genus Leporimetis Iredale, 1930 

Leporimetis intastriata (Say, 1826) 

Genus Macoma Leach, 1819 

Macoma brevifrons (Say, 1834) 

Macoma cerina Dall, 1900 

Macoma constricta (Bruguière, 1792) 

Macoma limula Dall, 1895 

Macoma mitchelli Dall, 1895 

Macoma phenax Dall, 1900 

Macoma tenta (Say, 1834) 

Genus Strigilla Turton, 1822 

Strigilla carnaria (Linnaeus, 1758) 

Strigilla gabbi Olsson & McGinty, 1958 

Strigilla mirabilis (Philippi, 1841) 

Strigilla surinamensis Boss, 1972 

Genus Tellidora H. & A. Adams, 1856 

Tellidora cristata (Récluz, 1842) 

Genus Tellina Linnaeus, 1758 

Tellina aequistriata Say, 1824 

Tellina alternata Say, 1822 

Tellina angulosa Gmelin, 1791 

Tellina candeana d’Orbigny, 1842 

Tellina consobrina d’Orbigny, 1842 

Tellina cristallina Spengler, 1798 

Tellina fausta Pulteney, 1799 

Tellina gouldii Hanley, 1846 

Tellina iris Say, 1822 

Tellina laevigata Linnaeus, 1758 

Tellina lineata Turton, 1819 

Tellina listeri Röding, 1798 

Tellina magna Spengler, 1798 

Tellina martinicensis d’Orbigny, 1842 

Tellina mera Say, 1834 

Tellina nitens C. B. Adams, 1845 

Tellina paramera Boss, 1964 

Tellina radiata Linnaeus, 1758 

Tellina similis J. Sowerby, 1806 

Tellina squamifera Deshayes, 1855 

Tellina sybaritica Dall, 1881 

Tellina tampaensis Conrad, 1866 

Tellina tenella A. E.Verrill, 1874 

Tellina texana Dall, 1900 

Tellina versicolor DeKay, 1843 

Family Donacidae Fleming, 1828 



Genus Donax Linnaeus, 1758 

Donax fossor Say, 1822 

Donax texasianus Philippi, 1847 

Donax variabilis Say, 1822 

Genus Iphigenia Schumacher, 1817 

Iphigenia brasiliana (Lamarck, 1818) 

Family Psammobiidae Fleming, 1828 

Genus Asaphis Modeer, 1793 

Asaphis deflorata (Linnaeus, 1758) 

Genus Heterodonax Mørch, 1853 

Heterodonax bimaculatus (Linnaeus, 1758) 

Genus Sanguinolaria Lamarck, 1799 

Sanguinolaria sanguinolenta (Gmelin, 1791) 

Family Semelidae Stoliczka, 1870 

Genus Abra Lamarck, 1818 

Abra aequalis (Say, 1822) 

Abra lioica (Dall, 1881) 

Genus Cumingia Sowerby, 1833 

Cumingia coarctata Sowerby, 1833 

Cumingia tellinoides (Conrad, 1831) 

Genus Ervilia Turton, 1822 

Ervilia concentrica (Holmes, 1860) 

Ervilia nitens (Montagu, 1808) 

Genus Semele Schumacher, 1817 

Semele bellastriata (Conrad, 1837) 

Semele proficua (Pulteney, 1799) 

Semele purpurascens (Gmelin, 1791) 

Genus Semelina Dall, 1900 

Semelina nuculoides (Conrad, 1841) 

Family Solecurtidae d’Orbigny, 1846 

Genus Solecurtus de Blainville, 1824 

Solecurtus cumingianus (Dunker, 1861) 

Solecurtus sanctaemarthae (d’Orbigny, 1842) 

Genus Tagelus J. E. Gray, 1847 

Tagelus divisus (Spengler, 1794) 

Tagelus plebeius (Lightfoot, 1786) 

Family Dreissenidae J. E. Gray, 1840 

Genus Mytilopsis Conrad, 1858 

Mytilopsis leucophaeata (Conrad, 1831) 

Mytilopsis sallei (Récluz, 1849) 

Family Trapezidae Lamy, 1920 

Genus Coralliophaga de Blainville, 1824 

Coralliophaga coralliophaga (Gmelin, 1791) 

Family Corbiculidae J. E. Gray, 1847 

Genus Polymesoda Rafinesque, 1820 

Polymesoda caroliniana (Bosc, 1801) 

Polymesoda maritima (d’Orbigny, 1842) 

Family Veneridae Rafinesque, 1815 

Genus Agriopoma Dall, 1902 

Agriopoma texasianum (Dall, 1892) 

Genus Anomalocardia Schumacher, 1817 

Anomalocardia auberiana (d’Orbigny, 1842) 

Genus Callista Poli, 1791 

Callista eucymata (Dall, 1890) 

Genus Chione Mühlfeld, 1811 

Chione cancellata (Linnaeus, 1767) 

Genus Cyclinella Dall, 1902 

Cyclinella tenuis (Récluz, 1852) 

Genus Dosinia Scopoli, 1777 

Dosinia discus (Reeve, 1850) 

Dosinia elegans (Conrad, 1843) 

Genus Gemma Deshayes, 1853 

Gemma gemma (Totten, 1834) 

Genus Globivenus Coen, 1934 

Globivenus rigida (Dillwyn, 1817) 

Globivenus rugatina (Heilprin, 1886) 

Genus Gouldia C. B. Adams, 1847 

Gouldia cerina (C. B. Adams, 1845) 

Genus Lirophora Conrad, 1862 

Lirophora latilirata (Conrad, 1841) 

Genus Macrocallista Meek, 1876 

Macrocallista maculata (Linnaeus, 1758) 

Macrocallista nimbosa (Lightfoot, 1786) 

Genus Mercenaria Schumacher, 1817 

Mercenaria campechiensis (Gmelin, 1791) 



Mercenaria mercenaria (Linnaeus, 1758) 

Genus Parastarte Conrad, 1862 

Parastarte triquetra (Conrad, 1846) 

Genus Periglypta Jukes-Brown, 1914 

Periglypta listeri (J. E. Gray, 1838) 

Genus Pitar Römer, 1857 

Pitar cordatus (Schwengel, 1951) 

Pitar fulminatus (Menke, 1828) 

Pitar pilula Rehder, 1943 

Pitar simpsoni (Dall, 1895) 

Genus Puberella Fischer-Piette, 1977 

Puberella intapurpurea (Conrad, 1850) 

Puberella pubera (Bory Saint-Vincent, 1827) 

Genus Timoclea T. Brown, 1827 

Timoclea grus (Holmes, 1858) 

Timoclea pygmaea (Lamarck, 1818) 

Genus Tivela Link, 1807 

Tivela floridana Rehder, 1939 

Genus Transennella Dall, 1884 

Transennella conradina Dall, 1884 

Transennella cubaniana (d’Orbigny, 1842) 

Transennella stimpsoni Dall, 1902 

Family Petricolidae Deschayes, 1831 

Genus Choristodon Jonas, 1844 

Choristodon robustum (G. B. Sowerby I, 1834) 

Genus Cooperella Carpenter, 1864 

Cooperella atlantica Rehder, 1943 

Genus Petricola Lamarck, 1801 

Petricola lapicida (Gmelin, 1791) 

Genus Petricolaria Stoliczka, 1870 

Petricolaria pholadiformis (Lamarck, 1818) 

Order Myoida Goldfuss, 1820 

Family Myidae Lamarck, 1809 

Genus Paramya Conrad, 1861 

Paramya subovata (Conrad, 1845) 

Genus Sphenia Turton, 1822 

Sphenia antillensis Dall & Simpson, 1901 

Family Corbulidae Lamarck, 1818 

Genus Caryocorbula Gardner, 1926 

Caryocorbula barrattiana (C. B. Adams, 1852) 

Genus Corbula Bruguière, 1792 

Corbula caribaea d’Orbigny, 1842 

Corbula chittyana C. B. Adams, 1852 

Corbula contracta Say, 1822 

Corbula dietziana C. B. Adams, 1852 

Corbula swiftiana C. B. Adams, 1852 

Genus Juliacorbula Olsson & Harbison, 1953 

Juliacorbula cubaniana (d’Orbigny, 1842) 

Genus Varicorbula Grant & Gale, 1931 

Varicorbula operculata (Philippi, 1848) 

Family Gastrochaenidae J. E. Gray, 1840 

Genus Gastrochaena Spengler, 1783 

Gastrochaena hians (Gmelin, 1791) 

Gastrochaena ovata G. B. Sowerby I, 1834 

Genus Spengleria Tryon, 1862 

Spengleria rostrata (Spengler, 1783) 

Family Hiatellidae J. E. Gray, 1824 

Genus Hiatella Bosc, 1801 

Hiatella arctica (Linnaeus, 1767) 

Hiatella azaria (Dall, 1881) 

Genus Panopea Ménard, 1807 

Panopea bitruncata (Conrad, 1872) 

Family Pholadidae Lamarck, 1809 

Genus Barnea Risso, 1826 

Barnea truncata (Say, 1822) 

Genus Cyrtopleura Tryon, 1862 

Cyrtopleura costata (Linnaeus, 1758) 

Genus Diplothyra Tryon, 1862 

Diplothyra smithii Tryon, 1862 

Genus Jouannetia DesMoulins, 1828 

Jouannetia quillingi Turner, 1955 

Genus Martesia Sowerby, 1824 

Martesia cuneiformis (Say, 1822) 

Martesia fragilis A. E.Verrill & Bush, 1890 

Martesia striata (Linnaeus, 1758) 

Genus Pholas Linnaeus, 1758 



Pholas campechiensis Gmelin, 1791 

Family Teredinidae Rafinesque, 1815 

Genus Bankia J. E. Gray, 1842 

Bankia carinata (J. E. Gray, 1827) 

Bankia fimbriatula Moll & Roch, 1931 

Bankia gouldi (Bartsch, 1908) 

Genus Lyrodus Gould, 1870 

Lyrodus bipartitus (Jeffreys, 1860) 

Lyrodus floridanus (Bartsch, 1922) 

Lyrodus mediolobatus (Edmondson, 1942) 

Lyrodus pedicellatus (Quatrefages, 1849) 

Genus Nototeredo Bartsch, 1923 

Nototeredo knoxi (Bartsch, 1917) 

Genus Spathoteredo Roch, 1937 

Spathoteredo spatha (Jeffreys, 1860) 

Genus Teredo Linnaeus, 1758 

Teredo bartschi W. Clapp, 1923 

Teredo fulleri W. Clapp, 1924 

Teredo furcifera von Martens, 1894 

Teredo navalis Linnaeus, 1758 

Teredo portoricensis W. Clapp, 1924 

Order Pholadomyoida Newell, 1965 

Family Lyonsiidae P. Fischer, 1887 

Genus Entodesma Philippi, 1845 

Entodesma beanum (d’Orbigny, 1842) 

Genus Lyonsia Turton, 1822 

Lyonsia floridana Conrad, 1848 

Lyonsia hyalina (Conrad, 1831) 

Family Pandoridae Rafinesque, 1815 

Genus Pandora Bruguière, 1797 

Pandora arenosa Conrad, 1834 

Pandora bushiana Dall, 1886 

Pandora trilineata Say, 1822 

Family Thraciidae Stoliczka, 1870 

Genus Asthenothaerus Carpenter, 1864 

Asthenothaerus balesi Rehder, 1943 

Asthenothaerus hemphillii Dall, 1886 

Genus Bushia Dall, 1886 

Bushia elegans (Dall, 1886) 

Genus Thracia Sowerby, 1823 

Thracia morrisoni R. Petit, 1964 

Family Periplomatidae Dall, 1895 

Genus Periploma Schumacher, 1817 

Periploma anguliferum (Philippi, 1847) 

Periploma margaritaceum (Lamarck, 1801) 

Periploma tenerum P. Fischer, 1882 

Family Poromyidae Dall, 1886 

Genus Poromya Forbes, 1844 

Poromya rostrata Rehder, 1943 

Family Verticordiidae Stoliczka, 1871 

Genus Haliris Dall, 1886 

Haliris fischeriana (Dall, 1881) 

Genus Trigonulina d’Orbigny, 1842 

Trigonulina ornata d’Orbigny, 1842 

Family Cuspidariidae Dall, 1886 

Genus Cardiomya A. Adams, 1864 

Cardiomya costellata (Deshayes, 1830) 

Cardiomya gemma A. E.Verrill & Bush, 1898 

Cardiomya ornatissima (d’Orbigny, 1842) 

Class Gastropoda Cuvier, 1797 

Snails, Limpets, Conchs, Whelks, etc. 

Order Patellogastropoda Lindberg, 1986 

Family Lottiidae J. E. Gray, 1840 

Genus Lottia J. E. Gray, 1833 

Lottia leucopleura (Gmelin, 1791) 

Genus Patelloida Quoy & Gaimard, 1834 

Patelloida pustulata (Helbling, 1779) 

Order Archaeogastropoda Thiele, 1925 

Family Scissurellidae J. E. Gray, 1847 

Genus Scissurella d’Orbigny, 1824 

Scissurella cingulata O. G. Costa, 1861 

Family Fissurellidae Fleming, 1822 

Genus Diodora J. E. Gray, 1821 

Diodora arcuata (G. B. Sowerby II, 1862) 

Diodora cayenensis (Lamarck, 1822) 

Diodora dysoni (Reeve, 1850) 



Diodora jaumei Aguayo & Rehder, 1936 

Diodora listeri (d’Orbigny, 1842) 

Diodora meta (von Ihering, 1927) 

Diodora minuta (Lamarck, 1822) 

Diodora sayi (Dall, 1889) 

Diodora variegata G. B. Sowerby II, 1862 

Diodora viridula (Lamarck, 1822) 

Genus Emarginula Lamarck, 1801 

Emarginula dentigera Heilprin, 1889 

Emarginula phrixodes Dall, 1927 

Emarginula pumila (A. Adams, 1852) 

Genus Fissurella Bruguière, 1789 

Fissurella angusta (Gmelin, 1791) 

Fissurella barbadensis (Gmelin, 1791) 

Fissurella fascicularis Lamarck, 1822 

Fissurella nodosa (Born, 1778) 

Fissurella rosea (Gmelin, 1791) 

Genus Hemitoma Swainson, 1840 

Hemitoma emarginata (de Blainville, 1825) 

Hemitoma octoradiata (Gmelin, 1791) 

Genus Lucapina Sowerby, 1835 

Lucapina aegis (Reeve, 1850) 

Lucapina philippiana (Finlay, 1930) 

Lucapina sowerbii (G. B. Sowerby II, 1835) 

Lucapina suffusa (Reeve, 1850) 

Genus Lucapinella Pilsbry, 1890 

Lucapinella limatula (Reeve, 1850) 

Genus Rimula Defrance, 1827 

Rimula aequisculpta Dall, 1927 

Rimula frenulata (Dall, 1889) 

Family Turbinidae Rafinesque, 1815 

Genus Arene H. & A. Adams, 1854 

Arene bairdii (Dall, 1889) 

Arene briareus (Dall, 1881) 

Arene cruentata (Mühlfeld, 1829) 

Arene tricarinata (Stearns, 1872) 

Arene variabilis (Dall, 1889) 

Arene venustula Aguayo & Rehder, 1936 

Genus Astralium Link, 1807 

Astralium phoebium (Röding, 1798) 

Genus Cyclostrema Marryat, 1818 

Cyclostrema cancellatum Marryat, 1818 

Cyclostrema tortuganum (Dall, 1927) 

Genus Didianema Woodring, 1928 

Didianema pauli Pilsbry & McGinty, 1945 

Genus Eulithidium Pilsbry, 1898 

Eulithidium affine (C. B. Adams, 1850) 

Eulithidium bellum (M. Smith, 1937) 

Eulithidium pterocladicum (Robertson, 1958) 

Eulithidium thalassicola (Robertson, 1958) 

Genus Lithopoma J. E. Gray, 1850 

Lithopoma americanum (Gmelin, 1791) 

Lithopoma caelatum (Gmelin, 1791) 

Lithopoma tectum (Lightfoot, 1786) 

Lithopoma tuber (Linnaeus, 1767) 

Genus Turbo Linnaeus, 1758 

Turbo cailletii P. Fischer & Bernardi, 1856 

Turbo canaliculatus Hermann, 1781 

Turbo castanea Gmelin, 1791 

Family Trochidae Rafinesque, 1815 

Genus Cittarium Philippi, 1847 

Cittarium pica (Linnaeus, 1758) 

Genus Euchelus Philippi, 1847 

Euchelus guttarosea Dall, 1889 

Genus Pseudostomatella Thiele, 1924 

Pseudostomatella erythrocoma (Dall, 1889) 

Genus Solariella S.V. Wood, 1842 

Solariella lacunella (Dall, 1881) 

Genus Synaptocochlea Pilsbry, 1890 

Synaptocochlea picta (d’Orbigny, 1842) 

Genus Tegula Lesson, 1835 

Tegula excavata (Lamarck, 1822) 

Tegula fasciata (Born, 1778) 

Tegula gruneri (Philippi, 1849) 

Tegula hotessieriana (d’Orbigny, 1842) 

Tegula lividomaculata (C. B. Adams, 1845) 

Family Calliostomatidae Thiele, 1921 

Genus Calliostoma Swainson, 1840 

Calliostoma adelae Schwengel, 1951 

Calliostoma barbouri Clench & Aguayo, 1946 

Calliostoma euglyptum (A. Adams, 1855) 

Calliostoma javanicum (Lamarck, 1822) 

Calliostoma jujubinum (Gmelin, 1791) 

Calliostoma pulchrum (C. B. Adams, 1850) 

Calliostoma roseolum Dall, 1881 



Calliostoma scalenum Quinn, 1992 

Calliostoma tampaense (Conrad, 1846) 

Calliostoma yucatecanum Dall, 1881 

Family Skeneidae Thiele, 1929 

Genus Haplocochlias Carpenter, 1864 

Haplocochlias swifti Vanatta, 1913 

Genus Parviturbo Pilsbry & McGinty, 1945 

Parviturbo calidimaris Pilsbry & McGinty, 1945 

Parviturbo francesae Pilsbry & McGinty, 1945 

Parviturbo rehderi Pilsbry & McGinty, 1945 

Parviturbo weberi Pilsbry & McGinty, 1945 

Order Neritopsina Cox & Knight, 1960 

Family Neritidae Rafinesque, 1815 

Genus Nerita Linnaeus, 1758 

Nerita fulgurans Gmelin, 1791 

Nerita peloronta Linnaeus, 1758 

Nerita tessellata Gmelin, 1791 

Nerita versicolor Gmelin, 1791 

Genus Neritina Lamarck, 1816 

Neritina clenchi Russell, 1940 

Neritina usnea (Röding, 1798) 

Neritina virginea (Linnaeus, 1758) 

Genus Puperita J. E. Gray, 1857 

Puperita pupa (Linnaeus, 1767) 

Genus Smaragdia Issel, 1869 

Smaragdia viridis (Linnaeus, 1758) 

Family Phenacolepadidae Pilsbry, 1900 

Genus Phenacolepas Pilsbry, 1891 

Phenacolepas hamillei (P. Fischer, 1857) 

Order Neotaenioglossa Haller, 1882 

Family Obtortionidae Thiele, 1925 

Genus Finella A. Adams, 1869 

Finella adamsi (Dall, 1889) 

Finella dubia (d’Orbigny, 1842) 

Family Litiopidae H. & A. Adams, 1854 

Genus Alaba H. & A. Adams, 1853 

Alaba incerta (d’Orbigny, 1842) 

Genus Litiopa Rang, 1829 

Litiopa melanostoma Rang, 1829 

Family Cerithiidae Fleming, 1822 

Genus Bittiolum Cossmann, 1906 

Bittiolum varium (Pfeiffer, 1840) 

Genus Cerithium Bruguière, 1789 

Cerithium atratum (Born, 1778) 

Cerithium eburneum Bruguière, 1792 

Cerithium guinaicum Philippi, 1849 

Cerithium lindae Petuch, 1987 

Cerithium litteratum (Born, 1778) 

Cerithium lutosum Menke, 1828 

Cerithium muscarum Say, 1822 

Family Batillariidae Thiele, 1929 

Genus Batillaria Benson, 1842 

Batillaria minima (Gmelin, 179l) 

Family Potamididae H. & A. Adams, 1854 

Genus Cerithidea Swainson, 1840 

Cerithidea costata (E. M. da Costa, 1778) 

Cerithidea scalariformis (Say, 1825) 

Family Planaxidae J. E. Gray, 1850 

Genus Fossarus Philippi, 1841 

Fossarus bellus Dall, 1889 

Fossarus orbignyi P. Fischer, 1864 

Genus Hinea J. E. Gray, 1847 

Hinea lineata (E. M. da Costa, 1778) 

Genus Supplanaxis Thiele, 1929 

Supplanaxis nucleus (Bruguière, 1789) 

Family Modulidae P. Fischer, 1884 

Genus Modulus J. E. Gray, 1842 

Modulus calusa Petuch, 1988 

Modulus floridanus Conrad, 1869 

Modulus modulus (Linnaeus, 1758) 

Modulus pacei Petuch, 1987 

Family Turritellidae Lovén, 1847 

Genus Turritella Lamarck, 1799 

Turritella acropora Dall, 1889 

Turritella exoleta (Linnaeus, 1758) 

Genus Vermicularia Lamarck, 1799 

Vermicularia fargoi Olsson, 1951 

Vermicularia knorrii (Deshayes, 1843) 



Vermicularia spirata (Philippi, 1836) 

Family Siliquariidae Anton, 1838 

Genus Tenagodus Guettard, 1774 

Tenagodus squamatus (de Blainville, 1827) 

Family Littorinidae J. E. Gray, 1840 

Genus Cenchritis von Martens, 1900 

Cenchritis muricatus (Linnaeus, 1758) 

Genus Littoraria J. E. Gray, 1834 

Littoraria angulifera (Lamarck, 1822) 

Littoraria irrorata (Say, 1822) 

Littoraria nebulosa (Lamarck, 1822) 

Littoraria tessellata (Philippi, 1847) 

Genus Nodilittorina von Martens, 1897 

Nodilittorina angustior (Mørch, 1876) 

Nodilittorina dilatata (d’Orbigny, 1842) 

Nodilittorina meleagris (Potiez & Michaud, 1838) 

Nodilittorina mespillum (Mühlfeld, 1824) 

Nodilittorina riisei (Mørch, 1876) 

Nodilittorina ziczac (Gmelin, 1791) 

Genus Tectarius Valenciennes, 1833 

Tectarius antonii (Philippi, 1846) 

Family Skeneopsidae Iredale, 1915 

Genus Skeneopsis Iredale, 1915 

Skeneopsis planorbis (Fabricius, 1780) 

Family Barleeiidae Thiele, 1925 

Genus Amphithalamus Carpenter, 1865 

Amphithalamus vallei Aguayo & Jaume, 1947 

Genus Barleeia Clark, 1853 

Barleeia tincta Guppy, 1895 

Family Rissoidae J. E. Gray, 1847 

Genus Alvania Risso, 1826 

Alvania auberiana (d’Orbigny, 1842) 

Genus Folinia Crosse, 1868 

Folinia bermudezi (Aguayo & Rehder, 1936) 

Genus Microstelma A. Adams, 1863 

Microstelma vestale (Rehder, 1943) 

Genus Rissoina d’Orbigny, 1840 

Rissoina cancellata Philippi, 1847 

Rissoina decussata (Montagu, 1803) 

Rissoina dyscrita Faber, 1990 

Rissoina multicostata (C. B. Adams, 1850) 

Rissoina princeps (C. B. Adams, 1850) 

Rissoina sagraiana d’Orbigny, 1842 

Rissoina striatocostata d’Orbigny, 1842 

Rissoina striosa (C. B. Adams, 1850) 

Genus Schwartziella Nevill, 1881 

Schwartziella bouryi (Desjardin, 1949) 

Schwartziella bryerea (Montagu, 1803) 

Schwartziella catesbyana (d’Orbigny, 1842) 

Schwartziella fischeri Desjardin, 1949 

Genus Stosicia Brusina, 1870 

Stosicia aberrans (C. B. Adams, 1850) 

Genus Zebina H. & A. Adams, 1854 

Zebina browniana (d’Orbigny, 1842) 

Zebina laevigata (C. B. Adams, 1850) 

Family Hydrobiidae Troschel, 1857 

Genus Floridiscrobs Pilsbry & McGinty, 1949 

Floridiscrobs dysbatus (Pilsbry & McGinty, 1949) 

Genus Heleobops F. G.Thompson, 1968 

Heleobops docimus F. G.Thompson, 1968 

Genus Littoridinops Pilsbry, 1952 

Littoridinops monroensis (Frauenfeld, 1863) 

Littoridinops palustris F. G.Thompson, 1968 

Littoridinops tenuipes (Couper, 1844) 

Genus Onobops F. G.Thompson, 1968 

Onobops crassus F. G.Thompson, 1968 

Onobops jacksoni (Bartsch, 1953) 

Genus Probythinella Thiele, 1928 

Probythinella protera Pilsbry, 1953 

Genus Pyrgophorus Ancey, 1888 

Pyrgophorus platyrachis F. G.Thompson, 1968 

Genus Spurwinkia Davis, Mazurkiewicz & Mandracchia, 

1982 

Spurwinkia salsa (Pilsbry, 1905) 

Genus Texadina Abbott & Ladd, 1951 

Texadina sphinctostoma (Abbott & Ladd, 1951) 

Family Truncatellidae J. E. Gray, 1840 



Genus Truncatella Risso, 1826 

Truncatella caribaeensis Reeve, 1842 

Truncatella clathrus Lowe, 1932 

Truncatella pulchella Pfeiffer, 1839 

Family Assimineidae H. & A. Adams, 1856 

Genus Assiminea Fleming, 1828 

Assiminea succinea (Pfeiffer, 1840) 

Family Elachisinidae Ponder, 1985 

Genus Elachisina Dall, 1918 

Elachisina floridana (Rehder, 1943) 

Family Vitrinellidae Bush, 1897 

Genus Anticlimax Pilsbry & McGinty, 1946 

Anticlimax athleenae (Pilsbry & McGinty, 1946) 

Anticlimax pilsbryi (McGinty, 1945) 

Genus Circulus Jeffreys, 1865 

Circulus liratus (Verrill, 1882) 

Circulus semisculptus (Olsson & McGinty, 1958) 

Circulus texanus (Moore, 1965) 

Genus Cochliolepis Stimpson, 1858 

Cochliolepis holmesii (Dall, 1889) 

Cochliolepis parasitica Stimpson, 1858 

Cochliolepis striata Dall, 1889 

Genus Cyclostremiscus Pilsbry & Olsson, 1945 

Cyclostremiscus beauii (P. Fischer, 1857) 

Cyclostremiscus cubanus (Pilsbry & Aguayo, 1933) 

Cyclostremiscus jeannae Pilsbry & McGinty, 1946 

Cyclostremiscus pentagonus (Gabb, 1873) 

Cyclostremiscus suppressus (Dall, 1889) 

Genus Episcynia Mørch, 1875 

Episcynia inornata (d’Orbigny, 1842) 

Genus Pachystremiscus Olsson & McGinty, 1958 

Pachystremiscus ornatus (Olsson & McGinty, 1958) 

Pachystremiscus pulchellus (Olsson & McGinty, 

1958) 

Genus Parviturboides Pilsbry & McGinty, 1950 

Parviturboides interruptus (C. B. Adams, 1850) 

Genus Pleuromalaxis Pilsbry & McGinty, 1945 

Pleuromalaxis balesi (Pilsbry & McGinty, 1945) 

Pleuromalaxis pauli (Olsson & McGinty, 1958) 

Genus Solariorbis Conrad, 1865 

Solariorbis blakei (Rehder, 1944) 

Solariorbis infracarinatus (Gabb, 1881) 

Solariorbis mooreanus (Vanatta, 1904) 

Solariorbis semipunctus D. R. Moore, 1965 

Solariorbis terminalis (Pilsbry & McGinty, 1946) 

Genus Teinostoma H. & A. Adams, 1854 

Teinostoma biscaynense Pilsbry & McGinty, 1945 

Teinostoma carinicallus Pilsbry & McGinty, 1946 

Teinostoma clavium Pilsbry & McGinty, 1945 

Teinostoma goniogyrus Pilsbry & McGinty, 1945 

Teinostoma incertum Pilsbry & McGinty, 1945 

Teinostoma lerema Pilsbry & McGinty, 1945 

Teinostoma lituspalmarum Pilsbry & McGinty, 

1945 

Teinostoma megastoma (C. B. Adams, 1850) 

Teinostoma parvicallum Pilsbry & McGinty, 1945 

Genus Vitrinella C. B. Adams, 1850 

Vitrinella filifera Pilsbry & McGinty, 1946 

Vitrinella floridana Pilsbry & McGinty, 1946 

Vitrinella helicoidea C. B. Adams, 1850 

Vitrinella hemphilli Vanatta, 1913 

Family Caecidae J. E. Gray, 1850 

Genus Caecum Fleming, 1813 

Caecum antillarum Carpenter, 1858 

Caecum bipartitum de Folin, 1870 

Caecum breve de Folin, 1867 

Caecum carolinianum Dall, 1892 

Caecum clava de Folin, 1867 

Caecum cooperi S. Smith, 1860 

Caecum cycloferum de Folin, 1867 

Caecum floridanum Stimpson, 1851 

Caecum gurgulio Carpenter, 1858 

Caecum heladum Olsson & Harbison, 1953 

Caecum imbricatum Carpenter, 1858 

Caecum johnsoni Winkley, 1908 

Caecum plicatum Carpenter, 1858 

Caecum pulchellum Stimpson, 1851 

Caecum regulare Carpenter, 1858 

Caecum strigosum de Folin, 1868 

Caecum subvolutum de Folin, 1874 

Caecum textile de Folin, 1867 

Caecum vestitum de Folin, 1868 

Genus Meioceras Carpenter, 1858 

Meioceras cornucopiae (Carpenter, 1858) 

Meioceras cubitatum (de Folin, 1868) 

Meioceras nitidum (Stimpson, 185l) 

Family Strombidae Rafinesque, 1815 



Genus Strombus Linnaeus, 1758 

Strombus alatus Gmelin, 1791 

Strombus costatus Gmelin, 1791 

Strombus gallus Linnaeus, 1758 

Strombus gigas Linnaeus, 1758 

Strombus pugilis Linnaeus, 1758 

Strombus raninus Gmelin, 1791 

Family Hipponicidae Troschel, 1861 

Genus Hipponix Defrance, 1819 

Hipponix antiquatus (Linnaeus, 1767) 

Hipponix subrufus (Lamarck, 1819) 

Family Vanikoridae J. E. Gray, 1840 

Genus Megalomphalus Brusina, 1871 

Megalomphalus adamsii (P. Fischer, 1857) 

Megalomphalus floridanus (D. R. Moore, 1965) 

Megalomphalus palmalitoris (Pilsbry & McGinty, 

1950) 

Genus Vanikoro Quoy & Gaimard, 1832 

Vanikoro oxychone Mørch, 1877 

Family Calyptraeidae Lamarck, 1809 

Genus Calyptraea Lamarck, 1799 

Calyptraea centralis (Conrad, 1841) 

Genus Cheilea Modeer, 1793 

Cheilea equestris (Linnaeus, 1758) 

Genus Crepidula Lamarck, 1799 

Crepidula aculeata (Gmelin, 1791) 

Crepidula convexa Say, 1822 

Crepidula fornicata (Linnaeus, 1758) 

Crepidula maculosa Conrad, 1846 

Crepidula plana Say, 1822 

Genus Crucibulum Schumacher, 1817 

Crucibulum auricula (Gmelin, 1791) 

Crucibulum striatum (Say, 1826) 

Family Capulidae Fleming, 1822 

Genus Capulus Montfort, 1810 

Capulus incurvus (Gmelin, 1791) 

Family Xenophoridae Troschel, 1852 

Genus Xenophora G. Fischer, 1807 

Xenophora conchyliophora (Born, 1780) 

Family Vermetidae Rafinesque, 1815 

Genus Dendropoma Mørch, 1861 

Dendropoma corrodens (d’Orbigny, 1842) 

Dendropoma irregulare (d’Orbigny, 1842) 

Genus Petaloconchus H. C. Lea, 1843 

Petaloconchus erectus (Dall, 1888) 

Petaloconchus varians (d’Orbigny, 1841) 

Genus Serpulorbis Sassi, 1827 

Serpulorbis decussatus (Gmelin, 1791) 

Family Cypraeidae Rafinesque, 1815 

Genus Erosaria Troschel, 1863 

Erosaria acicularis (Gmelin, 1791) 

Genus Macrocypraea Schilder, 1930 

Macrocypraea cervus (Linnaeus, 1771) 

Macrocypraea zebra (Linnaeus, 1758) 

Genus Propustularia Schilder, 1927 

Propustularia surinamensis (G. Perry, 1811) 

Genus Talparia Troschel, 1863 

Talparia cinerea (Gmelin, 1791) 

Family Ovulidae Fleming, 1822 

Genus Cymbovula Cate, 1974 

Cymbovula acicularis (Lamarck, 1810) 

Genus Cyphoma Röding, 1798 

Cyphoma alleneae Cate, 1973 

Cyphoma gibbosum (Linnaeus, 1758) 

Cyphoma mcgintyi Pilsbry, 1939 

Cyphoma rhomba Cate, 1978 

Cyphoma sedlaki Cate, 1976 

Cyphoma signatum Pilsbry & McGinty, 1939 

Genus Neosimnia P. Fischer, 1884 

Neosimnia advena (Cate, 1978) 

Genus Pseudocyphoma Cate, 1973 

Pseudocyphoma gibbulum Cate, 1978 

Genus Simnialena Cate, 1973 

Simnialena uniplicata (G. B. Sowerby II, 1848) 

Family Triviidae Troschel, 1863 

Genus Hespererato Schilder, 1932 

Hespererato maugeriae (J. E. Gray, 1832) 



Genus Niveria Jousseaume, 1884 

Niveria atomaria pygmaea (Schilder, 1931) 

Niveria candidula (Gaskoin, 1836) 

Niveria leucosphaera Schilder, 1931 

Niveria pediculus (Linnaeus, 1758) 

Niveria quadripunctata (J. E. Gray, 1827) 

Niveria suffusa (J. E. Gray, 1827) 

Genus Proterato Schilder, 1927 

Proterato gemma pygmaea Schilder, 1933 

Genus Pusula Jousseaume, 1884 

Pusula maltbiana (Schwengel & McGinty, 1942) 

Family Lamellariidae d’Orbigny, 1841 

Genus Lamellaria Montagu, 1815 

Lamellaria koto Schwengel, 1944 

Lamellaria leucosphaera Schwengel, 1942 

Lamellaria perspicua (Linnaeus, 1758) 

Genus Marsenina J. E. Gray, 1850 

Marsenina globosa L. M. Perry, 1939 

Family Naticidae Forbes, 1838 

Genus Haliotinella Souverbie in Souverbie & Montrouzier, 

1875 

Haliotinella patinaria (Guppy, 1876) 

Genus Natica Scopoli, 1777 

Natica livida Pfeiffer, 1840 

Natica marochiensis (Gmelin, 1791) 

Natica tedbayeri Rehder, 1986 

Genus Naticarius Duméril, 1806 

Naticarius canrena (Linnaeus, 1758) 

Naticarius verae Rehder, 1947 

Genus Neverita Risso, 1826 

Neverita duplicata (Say, 1822) 

Genus Polinices Montfort, 1810 

Polinices hepaticus (Röding, 1798) 

Polinices lacteus (Guilding, 1834) 

Polinices uberinus (d’Orbigny, 1842) 

Genus Sigatica Meyer & Aldrich, 1886 

Sigatica carolinensis (Dall, 1889) 

Sigatica semisulcata (J. E. Gray, 1839) 

Genus Sinum Röding, 1798 

Sinum maculatum (Say, 1831) 

Sinum perspectivum (Say, 1831) 

Genus Stigmaulax Mørch, 1852 

Stigmaulax sulcatus (Born, 1778) 

Genus Tectonatica Sacco, 1890 

Tectonatica pusilla (Say, 1822) 

Family Cassidae Latreille, 1825 

Genus Casmaria H. & A. Adams, 1853 

Casmaria ponderosa atlantica Clench, 1944 

Genus Cassis Scopoli, 1777 

Cassis flammea (Linnaeus, 1758) 

Cassis madagascariensis Lamarck, 1822 

Cassis tuberosa (Linnaeus, 1758) 

Genus Cypraecassis Stutchbury, 1837 

Cypraecassis testiculus (Linnaeus, 1758) 

Genus Phalium Link, 1807 

Phalium granulatum (Born, 1778) 

Family Tonnidae Suter, 1913 

Genus Tonna Brünnich, 1772 

Tonna galea (Linnaeus, 1758) 

Tonna pennata (Mørch, 1852) 

Family Ficidae Meek, 1864 

Genus Ficus Röding, 1798 

Ficus communis Röding, 1798 

Family Ranellidae J. E. Gray, 1854 

Genus Charonia Gistel, 1848 

Charonia tritonis variegata (Lamarck, 1816) 

Genus Cymatium Röding, 1798 

Cymatium aquatile (Reeve, 1844) 

Cymatium comptum (A. Adams, 1855) 

Cymatium corrugatum krebsii (Mørch, 1877) 

Cymatium cynocephalum (Lamarck, 1816) 

Cymatium femorale (Linnaeus, 1758) 

Cymatium labiosum (W. Wood, 1828) 

Cymatium martinianum (d’Orbigny, 1846) 

Cymatium muricinum (Röding, 1798) 

Cymatium nicobaricum (Röding, 1798) 

Cymatium occidentale (Mørch, 1877) 

Cymatium parthenopeum (von Salis, 1793) 

Cymatium tenuiliratum (Lischke, 1873) 

Cymatium vespaceum (Lamarck, 1822) 

Genus Linatella J. E. Gray, 1857 



Linatella caudata (Gmelin, 1791) 

Family Personidae J. E. Gray, 1854 

Genus Distorsio Röding, 1798 

Distorsio clathrata (Lamarck, 1816) 

Distorsio constricta mcgintyi Emerson & Puffer, 

1953 

Family Bursidae Thiele, 1925 

Genus Bufonaria Schumacher, 1817 

Bufonaria bufo (Bruguière, 1792) 

Genus Bursa Röding, 1798 

Bursa corrugata ponderosa (Reeve, 1844) 

Bursa granularis cubaniana (d’Orbigny, 1842) 

Bursa grayana Dunker, 1862 

Bursa rhodostoma thomae (d’Orbigny, 1842) 

Family Cerithiopsidae H. & A. Adams, 1854 

Genus Cerithiopsis Forbes & Hanley, 1849 

Cerithiopsis albovittata (C. B. Adams, 1850) 

Cerithiopsis flava (C. B. Adams, 1850) 

Cerithiopsis fusiformis (C. B. Adams, 1850) 

Cerithiopsis greenii (C. B. Adams, 1839) 

Cerithiopsis io Dall & Bartsch, 1911 

Cerithiopsis iota (C. B. Adams, 1845) 

Cerithiopsis lata (C. B. Adams, 1850) 

Cerithiopsis vanhyningi Bartsch, 1918 

Genus Horologica Laseron, 1956 

Horologica pupa (Dall & Simpson, 1901) 

Genus Retilaskeya Marshall, 1978 

Retilaskeya bicolor (C. B. Adams, 1845) 

Retilaskeya emersonii (C. B. Adams, 1839) 

Genus Seila A. Adams, 1861 

Seila adamsii (H. C. Lea, 1845) 

Family Triphoridae J. E. Gray, 1847 

Genus Cosmotriphora Olsson & Harbison, 1953 

Cosmotriphora decorata (C. B. Adams, 1850) 

Cosmotriphora melanura (C. B. Adams, 1850) 

Cosmotriphora olivacea (Dall, 1889) 

Genus Iniforis Jousseaume, 1884 

Iniforis turristhomae (Holten, 1802) 

Genus Marshallora Bouchet, 1985 

Marshallora nigrocincta (C. B. Adams, 1839) 

Genus Metaxia Monterosato, 1884 

Metaxia rugulosa (C. B. Adams, 1850) 

Metaxia taeniolata (Dall, 1889) 

Genus Similiphora Bouchet, 1985 

Similiphora intermedia (C. B. Adams, 1850) 

Genus Triphora de Blainville, 1828 

Triphora calva Faber & Moolenbeek, 1991 

Triphora lilacina (Dall, 1889) 

Triphora ornata (Deshayes, 1832) 

Family Epitoniidae S. S. Berry, 1910 

Genus Alexania Strand, 1928 

Alexania floridana (Pilsbry, 1945) 

Genus Amaea H. & A. Adams, 1854 

Amaea retifera (Dall, 1889) 

Genus Cirsotrema Mørch, 1852 

Cirsotrema dalli Rehder, 1945 

Genus Depressiscala de Boury, 1909 

Depressiscala nautlae (Mørch, 1874) 

Genus Epitonium Röding, 1798 

Epitonium albidum (d’Orbigny, 1842) 

Epitonium angulatum (Say, 1830) 

Epitonium apiculatum (Dall, 1889) 

Epitonium blainei Clench & Turner, 1953 

Epitonium candeanum (d’Orbigny, 1842) 

Epitonium championi Clench & Turner, 1952 

Epitonium denticulatum (G. B. Sowerby II, 1844) 

Epitonium echinaticosta (d’Orbigny, 1842) 

Epitonium foliaceicosta (d’Orbigny, 1842) 

Epitonium humphreysii (Kiener, 1838) 

Epitonium krebsii (Mørch, 1874) 

Epitonium lamellosum (Lamarck, 1822) 

Epitonium matthewsae Clench & Turner, 1952 

Epitonium multistriatum (Say, 1826) 

Epitonium novangliae (Couthouy, 1838) 

Epitonium occidentale (Nyst, 1871) 

Epitonium phymanthi Robertson, 1994 

Epitonium rupicola (Kurtz, 1860) 

Epitonium striatissimum (Monterosato, 1878) 

Epitonium tollini Bartsch, 1938 

Epitonium unifasciatum (G. B. Sowerby II, 1844) 

Genus Opalia H. & A. Adams, 1853 

Opalia burryi Clench & Turner, 1950 

Opalia crenata (Linnaeus, 1758) 

Opalia hotessieriana (d’Orbigny, 1842) 

Opalia pumilio (Mørch, 1874) 



Genus Opaliopsis Thiele, 1928 

Opaliopsis concava (Dall, 1889) 

Family Janthinidae Lamarck, 1812 

Genus Janthina Röding, 1798 

Janthina exigua Lamarck, 1816 

Janthina globosa Swainson, 1822 

Janthina janthina (Linnaeus, 1758) 

Janthina pallida W.Thompson, 1840 

Genus Recluzia Petit de la Saussaye, 1853 

Recluzia rollandiana Petit de la Saussaye, 1853 

Family Aclididae G. O. Sars, 1878 

Genus Graphis Jeffreys, 1867 

Graphis underwoodae Bartsch, 1947 

Family Eulimidae Troschel, 1853 

Genus Ersilia Monterosato, 1872 

Ersilia stancyki Warén, 1980 

Genus Eulima Risso, 1826 

Eulima auricincta (Abbott, 1958) 

Eulima bifasciata d’Orbigny, 1842 

Eulima fulvocincta C. B. Adams, 1850 

Genus Eulimostraca Bartsch, 1917 

Eulimostraca subcarinata (d’Orbigny, 1842) 

Genus Melanella Bowdich, 1822 

Melanella conoidea (Kurtz & Stimpson, 1851) 

Melanella eulimoides (C. B. Adams, 1850) 

Melanella gracilis (C. B. Adams, 1850) 

Melanella jamaicensis (C. B. Adams, 1845) 

Genus Microeulima Warén, 1992 

Microeulima hemphillii (Dall, 1884) 

Genus Nanobalcis Warén in Warén & Mifsud, 1990 

Nanobalcis worsfoldi Warén, 1990 

Genus Niso Risso, 1826 

Niso aeglees Bush, 1885 

Niso hendersoni Bartsch, 1953 

Genus Oceanida de Folin, 1870 

Oceanida graduata de Folin, 1870 

Oceanida inglei Lyons, 1978 

Genus Pelseneeria Kohler & Vaney, 1908 

Pelseneeria stimpsonii (A. E.Verrill, 1872) 

Genus Sabinella Monterosato, 1890 

Sabinella troglodytes (Thiele, 1925) 

Genus Umbilibalcis Bouchet & Warén, 1986 

Umbilibalcis lata (Dall, 1889) 

Genus Vitreolina Monterosato, 1884 

Vitreolina arcuata (C. B. Adams, 1850) 

Vitreolina bermudezi (Pilsbry & Aguayo, 1933) 

Order Neogastropoda Thiele, 1929 

Family Muricidae Rafinesque, 1815 

Genus Acanthotrophon Hertlein & Strong, 1951 

Acanthotrophon striatoides Vokes, 1980 

Genus Aspella Mørch, 1877 

Aspella senex Dall, 1903 

Genus Calotrophon Hertlein & Strong, 1951 

Calotrophon ostrearum (Conrad, 1846) 

Genus Chicoreus Montfort, 1810 

Chicoreus dilectus (A. Adams, 1855) 

Chicoreus pomum (Gmelin, 1791) 

Genus Dermomurex Monterosato, 1890 

Dermomurex elizabethae (McGinty, 1940) 

Dermomurex pacei Petuch, 1988 

Dermomurex pauperculus (C. B. Adams, 1850) 

Genus Eupleura H. & A. Adams, 1853 

Eupleura caudata (Say, 1822) 

Eupleura sulcidentata Dall, 1890 

Genus Favartia Jousseaume, 1880 

Favartia alveata (Kiener, 1842) 

Favartia cellulosa (Conrad, 1846) 

Favartia minirosea (Abbott, 1954) 

Favartia pacei Petuch, 1988 

Favartia richardbinghami (Petuch, 1987) 

Genus Haustellum Schumacher, 1817 

Haustellum bellegladeense (Vokes, 1963) 

Haustellum cabritii (Bernardi, 1859) 

Haustellum rubidum (F. C. Baker, 1897) 

Genus Hexaplex G. Perry, 1810 

Hexaplex fulvescens (G. B. Sowerby II, 1834) 

Genus Murexiella Clench & Farfante, 1945 

Murexiella glypta (M. Smith, 1938) 

Murexiella kalafuti Petuch, 1987 

Murexiella levicula (Dall, 1889) 

Murexiella macgintyi (M. Smith, 1938) 



Genus Muricopsis Bucquoy, Dautzenberg & Dollfus, 

1882 

Muricopsis caribbaea (Bartsch & Rehder, 1939) 

Muricopsis oxytata (M. Smith, 1938) 

Genus Plicopurpura Cossmann, 1903 

Plicopurpura patula (Linnaeus, 1758) 

Genus Pterotyphis Jousseaume, 1880 

Pterotyphis triangularis (A. Adams, 1855) 

Genus Stramonita Schumacher, 1817 

Stramonita haemastoma canaliculata (J. E. Gray, 

1839) 

Stramonita haemastoma floridana (Conrad, 1837) 

Stramonita rustica (Lamarck, 1822) 

Genus Thais Röding, 1798 

Thais deltoidea (Lamarck, 1822) 

Genus Trachypollia Woodring, 1928 

Trachypollia nodulosa (C. B. Adams, 1845) 

Trachypollia turricula (von Maltzan, 1884) 

Genus Urosalpinx Stimpson, 1865 

Urosalpinx cinerea (Say, 1822) 

Urosalpinx perrugata (Conrad, 1846) 

Urosalpinx tampaensis (Conrad, 1846) 

Family Coralliophilidae Chenu, 1859 

Genus Babelomurex Coen, 1922 

Babelomurex mansfieldi (McGinty, 1940) 

Genus Coralliophila H. & A. Adams, 1853 

Coralliophila aberrans (C. B. Adams, 1850) 

Coralliophila caribaea Abbott, 1958 

Coralliophila galea (Reeve, 1846) 

Coralliophila pacei Petuch, 1987 

Genus Quoyula Iredale, 1912 

Quoyula kalafuti Petuch, 1987 

Family Turbinellidae Swainson, 1840 

Genus Vasum Röding, 1798 

Vasum muricatum (Born, 1778) 

Family Buccinidae Rafinesque, 1815 

Genus Antillophos Woodring, 1928 

Antillophos candeanus (d’Orbigny, 1842) 

Genus Bailya M. Smith, 1944 

Bailya intricata (Dall, 1884) 

Bailya parva (C. B. Adams, 1850) 

Bailya weberi (Watters, 1983) 

Genus Cantharus Röding, 1798 

Cantharus cancellarius (Conrad, 1846) 

Cantharus multangulus (Philippi, 1848) 

Genus Colubraria Schumacher, 1817 

Colubraria testacea (Mørch, 1852) 

Genus Engina J. E. Gray, 1839 

Engina corinnae Crovo, 1971 

Engina turbinella (Kiener, 1835) 

Genus Parviphos Sarasua, 1984 

Parviphos adelus (Schwengel, 1942) 

Genus Pisania Bivona, 1832 

Pisania pusio (Linnaeus, 1758) 

Genus Pollia J. E. Gray in Sowerby, 1834 

Pollia auritula (Link, 1807) 

Pollia karinae (Nowell-Usticke, 1959) 

Pollia tincta Conrad, 1846 

Family Melongenidae Gill, 1867 

Genus Busycon Röding, 1798 

Busycon carica (Gmelin, 1791) 

Busycon sinistrum Hollister, 1958 

Genus Busycotypus Wenz, 1943 

Busycotypus canaliculatus (Linnaeus, 1758) 

Busycotypus spiratus (Lamarck, 1816) 

Genus Melongena Schumacher, 1817 

Melongena bicolor (Say, 1826) 

Melongena corona (Gmelin, 1791) 

Melongena sprucecreekensis Tucker, 1994 

Family Nassariidae Iredale, 1916 

Genus Ilyanassa Stimpson, 1865 

Ilyanassa obsoleta (Say, 1822) 

Ilyanassa trivittata (Say, 1822) 

Genus Nassarius Duméril, 1806 

Nassarius acutus (Say, 1822) 

Nassarius albus (Say, 1826) 

Nassarius antillarum (d’Orbigny, 1842) 

Nassarius consensus (Ravenel, 1861) 

Nassarius hotessieri (d’Orbigny, 1845) 

Nassarius polygonatus (Lamarck, 1822) 



Nassarius vibex (Say, 1822) 

Family Fasciolariidae J. E. Gray, 1853 

Genus Fasciolaria Lamarck, 1799 

Fasciolaria hunteria (G. Perry, 1811) 

Fasciolaria tulipa (Linnaeus, 1758) 

Genus Fusinus Rafinesque, 1815 

Fusinus eucosmius (Dall, 1889) 

Fusinus helenae Bartsch, 1939 

Genus Heilprinia Grabau, 1904 

Heilprinia timessa (Dall, 1889) 

Genus Latirus Montfort, 1810 

Latirus angulatus (Röding, 1798) 

Latirus carinifer Lamarck, 1822 

Latirus infundibulum (Gmelin, 1791) 

Latirus nematus Woodring, 1928 

Genus Leucozonia J. E. Gray, 1847 

Leucozonia nassa (Gmelin, 1791) 

Leucozonia ocellata (Gmelin, 1791) 

Genus Pleuroploca P. Fischer, 1884 

Pleuroploca gigantea (Kiener, 1840) 

Genus Teralatirus Coomans, 1965 

Teralatirus cayohuesonicus (G. B. Sowerby III, 

1878) 

Family Columbellidae Swainson, 1840 

Genus Aesopus Gould, 1860 

Aesopus stearnsii (Tryon, 1883) 

Genus Astyris H. & A. Adams, 1853 

Astyris lunata (Say, 1826) 

Astyris multilineata (Dall, 1889) 

Astyris raveneli (Dall, 1889) 

Genus Columbella Lamarck, 1799 

Columbella mercatoria (Linnaeus, 1758) 

Columbella rusticoides Heilprin, 1886 

Genus Columbellopsis Bucquoy, Dautzenberg & Dollfus, 

1882 

Columbellopsis nycteis (Duclos, 1846) 

Genus Costoanachis Sacco, 1890 

Costoanachis avara (Say, 1822) 

Costoanachis catenata (G. B. Sowerby I, 1844) 

Costoanachis floridana (Rehder, 1939) 

Costoanachis lafresnayi (P. Fischer & Bernardi, 

1857) 

Costoanachis semiplicata (Stearns, 1873) 

Costoanachis similis (Ravenel, 1861) 

Costoanachis sparsa (Reeve, 1859) 

Genus Mazatlania Dall, 1913 

Mazatlania cosentini (Philippi, 1836) 

Genus Mitrella Risso, 1826 

Mitrella dichroa (G. B. Sowerby I, 1844) 

Mitrella ocellata (Gmelin, 1791) 

Genus Nassarina Dall, 1889 

Nassarina bushiae (Dall, 1889) 

Nassarina glypta (Bush, 1885) 

Genus Nitidella Swainson, 1840 

Nitidella nitida (Lamarck, 1822) 

Genus Parvanachis Radwin, 1968 

Parvanachis obesa (C. B. Adams, 1845) 

Parvanachis ostreicola (G. B. Sowerby III, 1882) 

Genus Rhombinella Radwin, 1968 

Rhombinella laevigata (Linnaeus, 1758) 

Genus Steironepion Pilsbry & Lowe, 1932 

Steironepion minor (C. B. Adams, 1845) 

Steironepion moniliferum (G. B. Sowerby I, 1844) 

Genus Suturoglypta Radwin, 1968 

Suturoglypta iontha (Ravenel, 1861) 

Suturoglypta pretrii (Duclos, 1846) 

Genus Zafrona Iredale, 1916 

Zafrona dicomata (Dall, 1889) 

Zafrona idalina (Duclos, 1840) 

Zafrona pulchella (de Blainville, 1829) 

Zafrona taylorae Petuch, 1987 

Family Volutidae Rafinesque, 1815 

Genus Enaeta H. & A. Adams, 1853 

Enaeta cylleniformis (G. B. Sowerby I, 1844) 

Genus Scaphella Swainson, 1832 

Scaphella junonia (Lamarck, 1804) 

Family Olividae Latreille, 1825 

Genus Jaspidella Olsson, 1956 

Jaspidella blanesi (Ford, 1898) 

Jaspidella jaspidea (Gmelin, 1791) 



Genus Oliva Bruguière, 1789 

Oliva bollingi Clench, 1937 

Oliva sayana Ravenel, 1834 

Genus Olivella Swainson, 1831 

Olivella bullula (Reeve, 1850) 

Olivella floralia (Duclos, 1853) 

Olivella inusta G. B. Sowerby III, 1915 

Olivella lactea (Marrat, 1871) 

Olivella macgintyi Olsson, 1956 

Olivella minuta (Link, 1807) 

Olivella monilifera (Reeve, 1850) 

Olivella mutica (Say, 1822) 

Olivella nivea (Gmelin, 1791) 

Olivella perplexa Olsson, 1956 

Olivella pusilla (Marrat, 1871) 

Family Harpidae Bronn, 1849 

Genus Morum Röding, 1798 

Morum oniscus (Linnaeus, 1767) 

Family Cystiscidae Stimpson, 1865 

Genus Gibberula Swainson, 1840 

Gibberula lavalleeana (d’Orbigny, 1842) 

Genus Granulina Jousseaume, 1888 

Granulina hadria (Dall, 1889) 

Genus Persicula Schumacher, 1817 

Persicula catenata (Montagu, 1803) 

Persicula pulcherrima (Gaskoin, 1849) 

Genus Pugnus Hedley, 1896 

Pugnus lachrimula (Gould, 1862) 

Pugnus serrei (Bavay, 1911) 

Family Marginellidae Fleming, 1828 

Genus Dentimargo Cossmann, 1899 

Dentimargo aureocinctus (Stearns, 1872) 

Dentimargo eburneolus (Conrad, 1834) 

Dentimargo idiochila (Schwengel, 1943) 

Dentimargo reductus (Bavay, 1922) 

Genus Hyalina Schumacher, 1817 

Hyalina pallida (Linnaeus, 1758) 

Genus Prunum Herrmannsen, 1852 

Prunum amabile (Redfield, 1852) 

Prunum apicinum (Menke, 1828) 

Prunum avenaceum (Deshayes, 1844) 

Prunum carneum (Storer, 1837) 

Prunum evelynae (Bayer, 1943) 

Prunum guttatum (Dillwyn, 1817) 

Prunum roscidum (Redfield, 1860) 

Prunum succinea (Conrad, 1846) 

Prunum virginianum hartleyanum (Schwengel, 

1941) 

Prunum virginianum virginianum Conrad, 1868 

Genus Volvarina Hinds, 1844 

Volvarina albolineata (d’Orbigny, 1842) 

Volvarina avena (Kiener, 1834) 

Volvarina lactea (Kiener, 1841) 

Volvarina subtriplicata (d’Orbigny, 1842) 

Family Mitridae Swainson, 1831 

Genus Mitra Lamarck, 1798 

Mitra barbadensis (Gmelin, 1791) 

Mitra florida Gould, 1856 

Mitra nodulosa (Gmelin, 1791) 

Mitra pallida Nowell-Usticke, 1959 

Mitra semiferruginea Reeve, 1845 

Mitra straminea A. Adams, 1853 

Family Costellariidae MacDonald, 1860 

Genus Thala H. & A. Adams, 1853 

Thala floridana (Dall, 1884) 

Genus Vexillum Röding, 1798 

Vexillum dermestinum (Lamarck, 1811) 

Vexillum epiphaneum (Rehder, 1943) 

Vexillum exiguum (C. B. Adams, 1845) 

Vexillum gemmatum (G. B. Sowerby II, 1874) 

Vexillum hendersoni (Dall, 1927) 

Vexillum histrio (Reeve, 1844) 

Vexillum puella (Reeve, 1845) 

Vexillum pulchellum (Reeve, 1844) 

Vexillum sykesi (Melvill, 1925) 

Vexillum wandoense (Holmes, 1860) 

Vexillum zythochroa (Melvill, 1888) 

Family Cancellariidae J. E. Gray, 1853 

Genus Agatrix Petit, 1967 

Agatrix agassizii (Dall, 1889) 

Genus Axelella Petit, 1988 

Axelella smithii (Dall, 1888) 

Genus Cancellaria Lamarck, 1799 

Cancellaria adelae Pilsbry, 1940 

Cancellaria reticulata (Linnaeus, 1767) 



Genus Trigonostoma de Blainville, 1827 

Trigonostoma rugosum (Lamarck, 1822) 

Trigonostoma tenerum (Philippi, 1848) 

Genus Tritonoharpa Dall, 1908 

Tritonoharpa lanceolata (Menke, 1828) 

Family Drilliidae Morrison, 1966 

Genus Bellaspira Conrad, 1868 

Bellaspira pentagonalis (Dall, 1889) 

Genus Cerodrillia Bartsch & Rehder, 1939 

Cerodrillia bealiana Schwengel & McGinty, 1942 

Cerodrillia clappi Bartsch & Rehder, 1939 

Cerodrillia perryae Bartsch & Rehder, 1939 

Cerodrillia thea (Dall, 1884) 

Genus Drillia J. E. Gray, 1838 

Drillia acurugata Dall, 1890 

Drillia cydia (Bartsch, 1943) 

Genus Lissodrillia Bartsch & Rehder, 1939 

Lissodrillia simpsoni (Dall, 1887) 

Genus Splendrillia Hedley, 1922 

Splendrillia fucata (Reeve, 1845) 

Splendrillia halidorema (Schwengel, 1940) 

Splendrillia moseri (Dall, 1889) 

Splendrillia woodringi (Bartsch, 1934) 

Family Terebridae Mørch, 1852 

Genus Hastula H. & A. Adams, 1853 

Hastula cinerea (Born, 1778) 

Hastula hastata (Gmelin, 1791) 

Hastula salleana (Deshayes, 1859) 

Genus Terebra Bruguière, 1789 

Terebra arcas Abbott, 1954 

Terebra concava (Say, 1826) 

Terebra dislocata (Say, 1822) 

Terebra floridana Dall, 1889 

Terebra glossema Schwengel, 1942 

Terebra protexta (Conrad, 1846) 

Terebra rushii Dall, 1889 

Terebra taurina (Lightfoot, 1786) 

Terebra vinosa Dall, 1889 

Family Strictispiridae J. H. McLean, 1971 

Genus Strictispira J. H. McLean, 1971 

Strictispira solida (C. B. Adams, 1850) 

Family Turridae Swainson, 1840 

Genus Compsodrillia Woodring, 1928 

Compsodrillia eucosmia (Dall, 1889) 

Genus Crassispira Swainson, 1840 

Crassispira cubana (Melvill, 1923) 

Crassispira fuscescens (Reeve, 1845) 

Crassispira phasma Schwengel, 1940 

Crassispira rhythmica Melvill, 1927 

Crassispira tampaensis (Bartsch & Rehder, 1939) 

Genus Pilsbryspira Bartsch, 1950 

Pilsbryspira albocincta (C. B. Adams, 1845) 

Pilsbryspira albomaculata (d’Orbigny, 1842) 

Pilsbryspira leucocyma (Dall, 1884) 

Pilsbryspira monilis (Bartsch & Rehder, 1939) 

Pilsbryspira zebroides (Weinkauff, 1876) 

Genus Polystira Woodring, 1928 

Polystira albida (G. Perry, 1811) 

Genus Pyrgospira J. H. McLean, 1971 

Pyrgospira ostrearum (Stearns, 1872) 

Genus Viridrillia Bartsch, 1943 

Viridrillia williami Bartsch, 1943 

Genus Zonulispira Bartsch, 1950 

Zonulispira sanibelensis (Bartsch & Rehder, 1939) 

Family Conidae Fleming, 1822 

Genus Agathotoma Cossmann, 1899 

Agathotoma candidissima (C. B. Adams, 1845) 

Genus Brachycythara Woodring, 1928 

Brachycythara barbarae Lyons, 1972 

Brachycythara biconica (C. B. Adams, 1850) 

Genus Conus Linnaeus, 1758 

Conus amphiurgus Dall, 1889 

Conus anabathrum Crosse, 1865 

Conus attenuatus Reeve, 1844 

Conus daucus Hwass, 1792 

Conus delessertii Récluz, 1843 

Conus ermineus Born, 1778 

Conus flamingo Petuch, 1980 

Conus flavescens G. B. Sowerby II, 1834 

Conus floridensis G. B. Sowerby II, 1870 

Conus granulatus Linnaeus, 1758 

Conus harasewychi Petuch, 1987 

Conus mindanus Hwass, 1792 

Conus mus Hwass, 1792 



Conus patae Abbott, 1971 

Conus pealii Green, 1830 

Conus regius Gmelin, 1791 

Conus sennottorum Rehder & Abbott, 1951 

Conus spurius Gmelin, 1791 

Conus stearnsii Conrad, 1869 

Conus stimpsoni Dall, 1902 

Conus ziczac Mühlfeld, 1816 

Genus Cryoturris Woodring, 1928 

Cryoturris cerinella (Dall, 1889) 

Cryoturris fargoi McGinty, 1955 

Cryoturris quadrilineata (C. B. Adams, 1850) 

Genus Daphnella Hinds, 1844 

Daphnella cingulata (Dall, 1890) 

Daphnella lymneiformis (Kiener, 1840) 

Daphnella margaretae Lyons, 1972 

Genus Glyphostoma Gabb, 1872 

Glyphostoma dentiferum Gabb, 1872 

Glyphostoma gabbii Dall, 1889 

Genus Glyphoturris Woodring, 1928 

Glyphoturris diminuta (C. B. Adams, 1850) 

Glyphoturris eritima (Bush, 1885) 

Glyphoturris quadrata (Reeve, 1845) 

Glyphoturris rugirima (Dall, 1889) 

Genus Glyptaesopus Pilsbry & Olsson, 1941 

Glyptaesopus proctorae (M. Smith, 1936) 

Genus Granoturris Fargo, 1953 

Granoturris presleyi Lyons, 1972 

Genus Ithycythara Woodring, 1928 

Ithycythara auberiana (d’Orbigny, 1842) 

Ithycythara lanceolata (C. B. Adams, 1850) 

Ithycythara parkeri Abbott, 1958 

Ithycythara pentagonalis (Reeve, 1845) 

Ithycythara psila (Bush, 1885) 

Genus Kurtziella Dall, 1918 

Kurtziella accincta (Montagu, 1808) 

Kurtziella atrostyla (Tryon, 1884) 

Kurtziella cerina (Kurtz & Stimpson, 1851) 

Kurtziella dorvilliae (Reeve, 1845) 

Kurtziella limonitella (Dall, 1884) 

Kurtziella perryae Bartsch & Rehder, 1939 

Genus Lioglyphostoma Woodring, 1928 

Lioglyphostoma canna (Dall, 1889) 

Genus Mitrolumna Bucquoy, Dautzenberg & Dollfus, 

1883 

Mitrolumna biplicata (Dall, 1889) 

Genus Nannodiella Dall, 1919 

Nannodiella oxia (Bush, 1885) A 

Nannodiella vespuciana (d’Orbigny, 1842) 

Genus Platycythara Woodring, 1928 

Platycythara elata (Dall, 1889) 

Genus Pyrgocythara Woodring, 1928 

Pyrgocythara filosa Rehder, 1943 

Pyrgocythara hemphilli Bartsch & Rehder, 1939 

Pyrgocythara plicosa (C. B. Adams, 1850) 

Genus Rimosodaphnella Cossmann, 1915 

Rimosodaphnella morra (Dall, 1881) 

Genus Rubellatoma Bartsch & Rehder, 1939 

Rubellatoma diomedea Bartsch & Rehder, 1939 

Rubellatoma rubella (Kurtz & Stimpson, 1851) 

Genus Stellatoma Bartsch & Rehder, 1939 

Stellatoma stellata (Stearns, 1872) 

Genus Tenaturris Woodring, 1928 

Tenaturris bartletti (Dall, 1889) 

Genus Thelecythara Woodring, 1928 

Thelecythara floridana Fargo, 1953 

Genus Vitricythara Fargo, 1953 

Vitricythara metria (Dall, 1903) 

Subclass Heterobranchia J. E. Gray, 1840 

Order Heterostropha P. Fischer, 1885 

Family Cornirostridae Ponder, 1990 

Genus Tomura Pilsbry & Olsson, 1952 

Tomura bicaudata (Pilsbry & McGinty, 1946) 

Family Mathildidae Dall, 1889 

Genus Mathilda Semper, 1865 

Mathilda hendersoni Dall, 1927 

Mathilda yucatecana (Dall, 1881) 

Family Architectonicidae J. E. Gray, 1850 

Genus Architectonica Röding, 1798 

Architectonica nobilis Röding, 1798 

Genus Heliacus d’Orbigny, 1842 

Heliacus bisulcatus (d’Orbigny, 1842) 

Heliacus cylindricus (Gmelin, 1791) 



Heliacus infundibuliformis perrieri (Rochebrune, 

1881) 

Genus Psilaxis Woodring, 1928 

Psilaxis krebsii (Mørch, 1875) 

Genus Spirolaxis Monterosato, 1913 

Spirolaxis centrifuga (Monterosato, 1890) 

Family Rissoellidae J. E. Gray, 1850 

Genus Rissoella J. E. Gray, 1847 

Rissoella caribaea Rehder, 1943 

Family Ebalidae Warén, 1994 

Genus Henrya Bartsch, 1947 

Henrya morrisoni Bartsch, 1947 

Genus Murchisonella Mørch, 1875 

Murchisonella tampaensis (Bartsch, 1947) 

Family Pyramidellidae J. E. Gray, 1840 

Genus Boonea Roberston, 1978 

Boonea bisuturalis (Say, 1822) 

Boonea impressa (Say, 1822) 

Boonea seminuda (C. B. Adams, 1839) 

Genus Careliopsis Mørch, 1875 

Careliopsis bartschi (Aguayo & Rehder, 1936) 

Careliopsis styliformis (Mørch, 1875) 

Genus Chrysallida Carpenter, 1857 

Chrysallida gemmulosa (C. B. Adams, 1850) 

Chrysallida toroensis (Olsson & McGinty, 1958) 

Genus Cyclostremella Bush, 1897 

Cyclostremella humilis Bush, 1897 

Genus Eulimastoma Bartsch, 1904 

Eulimastoma engonium (Bush, 1885) 

Eulimastoma teres (Bush, 1885) 

Eulimastoma weberi (Morrison, 1965)) 

Genus Fargoa Robertson, 1978 

Fargoa bartschi (Winkley, 1909) 

Fargoa bushiana (Bartsch, 1909) 

Fargoa dianthophila (H. W. Wells & M. J. Wells, 

1961) 

Fargoa dux (Dall & Bartsch, 1906) 

Genus Houbricka Wise, 1996 

Houbricka incisa (Bush, 1899) 

Genus Ividia Dall & Bartsch, 1904 

Ividia abbotti (Olsson & McGinty, 1958) 

Ividia havanensis (Pilsbry & Aguayo, 1933) 

Genus Kleinella A. Adams, 1860 

Kleinella cedrosa (Dall, 1884) 

Genus Longchaeus Mørch, 1875 

Longchaeus candidus (Mørch, 1875) 

Longchaeus crenulatus (Holmes, 1860) 

Genus Odostomia Fleming, 1817 

Odostomia acutidens Dall, 1884 

Odostomia didyma (A. E.Verrill & Bush, 1900) 

Odostomia laevigata (d’Orbigny, 1842) 

Odostomia producta (C. B. Adams, 1840) 

Odostomia somersi A. E.Verrill & Bush, 1900 

Odostomia virginica J. B. Henderson & Bartsch, 

1914 

Genus Orinella Dall & Bartsch, 1904 

Orinella vanhyningi Bartsch, 1944 

Genus Peristichia Dall, 1889 

Peristichia agria Dall, 1889 

Peristichia toreta Dall, 1889 

Genus Petitilla Wise, 1996 

Petitilla crosseana (Dall, 1885) 

Genus Pseudoscilla Boettger, 1901 

Pseudoscilla babylonia (C. B. Adams, 1845) 

Genus Pyramidella Lamarck, 1799 

Pyramidella dolabrata (Linnaeus, 1758) 

Pyramidella resticula (Dall, 1889) 

Genus Sayella Dall, 1885 

Sayella fusca (C. B. Adams, 1839) 

Sayella hemphillii (Dall, 1884) 

Genus Triptychus Mørch, 1875 

Triptychus niveus Mørch, 1875 

Genus Turbonilla Risso, 1826 

Turbonilla areolata A. E.Verrill, 1873 

Turbonilla belotheca Dall, 1889 

Turbonilla conradi Bush, 1899 

Turbonilla constricta Bush, 1899 

Turbonilla coomansi Van Aartsen, 1994 

Turbonilla curta Dall, 1889 

Turbonilla dalli Bush, 1899 

Turbonilla exilis (C. B. Adams, 1850) 

Turbonilla hemphilli Bush, 1899 



Turbonilla interrupta (Totten, 1835) 

Turbonilla levis (C. B. Adams, 1850) 

Turbonilla multicostata (C. B. Adams, 1850) 

Turbonilla obeliscus (C. B. Adams, 1850) 

Turbonilla protracta Dall, 1892 

Turbonilla puncta (C. B. Adams, 1850) 

Turbonilla punicea Dall, 1884 

Turbonilla pupoides d’Orbigny, 1842 

Turbonilla reticulata (C. B. Adams, 1850) 

Turbonilla riisei Mørch, 1875 

Turbonilla textilis (Kurtz, 1860) 

Turbonilla toyatani J. B. Henderson & Bartsch, 

1914 

Turbonilla valida Verrill & Bush, 1900 

Turbonilla virga Dall, 1884 

Turbonilla virgata Dall, 1892 

Turbonilla viridaria Dall, 1884 

Turbonilla wrightsvillensis Powell, 1981 

Family Amathinidae Ponder, 1987 

Genus Cyclothyca Stearns, 1891 

Cyclothyca pacei Petuch, 1987 

Genus Iselica Dall, 1918 

Iselica anomala (C. B. Adams, 1850) 

Order Uncertain 

Family Acteonidae d’Orbigny, 1842 

Genus Acteon Montfort, 1810 

Acteon candens Rehder, 1939 

Genus Rictaxis Dall, 1871 

Rictaxis punctostriatus (C. B. Adams, 1840) 

Subclass Opisthobranchia Milne-Edwards, 1848 

Order Cephalaspidea P. Fischer, 1883 

Family Ringiculidae Philippi, 1853 

Genus Ringicula Deshayes, 1838 

Ringicula semistriata d’Orbigny, 1842 

Family Aplustridae J. E. Gray, 1847 

Genus Hydatina Schumacher, 1817 

Hydatina physis (Linnaeus, 1758) 

Genus Micromelo Pilsbry, 1895 

Micromelo undatus (Bruguière, 1792) 

Family Cylichnidae A. Adams, 1850 

Genus Acteocina J. E. Gray, 1847 

Acteocina atrata P. S. Mikkelsen & P. M. 

Mikkelsen, 1984 

Acteocina bidentata (d’Orbigny, 1841) 

Acteocina bullata (Kiener, 1834) 

Acteocina canaliculata (Say, 1826) 

Acteocina candei (d’Orbigny, 1841) 

Acteocina lepta Woodring, 1928 

Acteocina recta (d’Orbigny, 1841) 

Genus Cylichna Lovén, 1846 

Cylichna eburnea A. E.Verrill, 1885 

Cylichna verrillii Dall, 1889 

Genus Scaphander Montfort, 1810 

Scaphander pilsbryi McGinty, 1955 

Genus Tornatina A. Adams, 1850 

Tornatina inconspicua (Olsson & McGinty, 1958) 

Family Philinidae J. E. Gray, 1850 

Genus Philine Ascanius, 1772 

Philine sagra (d’Orbigny, 1841) 

Family Aglajidae Pilsbry, 1895 

Genus Chelidonura A. Adams, 1850 

Chelidonura hirundinina (Quoy & Gaimard, 1833) 

Chelidonura sabina Ev. Marcus & Er. Marcus,1970 

Genus Navanax Pilsbry, 1895 

Navanax aenigmaticus (Bergh, 1893) 

Genus Philinopsis Pease, 1860 

Philinopsis pusa (Ev. Marcus & Er. Marcus, 1967) 

Family Gastropteridae Swainson, 1840 

Genus Gastropteron Meckel in Kosse, 1813 

Gastropteron chacmol Gosliner, 1989 

Gastropteron vespertilium Gosliner & Armes, 1984 

Family Retusidae Thiele, 1929 

Genus Pyrunculus Pilsbry, 1895 

Pyrunculus caelatus (Bush, 1885) 

Genus Retusa Brown, 1827 

Retusa sulcata (d’Orbigny, 1841) 

Genus Volvulella Newton, 1891 

Volvulella paupercula (Watson, 1883) 

Volvulella persimilis (Mørch, 1875) 

Volvulella recta (Mørch, 1875) 



Family Bullidae Lamarck, 1801 

Genus Bulla Linnaeus, 1758 

Bulla solida Gmelin, 1791 

Bulla striata Bruguière, 1792 

Family Haminoeidae Pilsbry, 1895 

Genus Atys Montfort, 1810 

Atys caribaeus (d’Orbigny, 1841) 

Atys riiseanus Mørch, 1875 

Atys sandersoni Dall, 1881 

Genus Haminoea Turton & Kingston in Carrington, 

1830 

Haminoea antillarum (d’Orbigny, 1841) 

Haminoea elegans (J. E. Gray, 1825) 

Haminoea petitii (d’Orbigny, 1841) 

Haminoea succinea (Conrad, 1846) 

Family Runcinidae H. & A. Adams, 1854 

Genus Runcina Forbes & Hanley, 1851 

Runcina divae (Ev. Marcus & Er. Marcus, 1963) 

Order Acochlidioidea Küthe, 1935 

Family Microhedylidae Odhner, 1938 

Genus Unela Er. Marcus, 1953 

Unela remanei Er. Marcus, 1953 

Order Thecosomata de Blainville, 1824 

Family Cavoliniidae H. & A. Adams, 1854 

Genus Creseis Rang, 1828 

Creseis acicula (Rang, 1828) 

Order Anaspidea P. Fischer, 1883 

Family Akeridae Pilsbry, 1893 

Genus Akera Müller, 1776 

Akera thompsoni Olsson & McGinty, 1951 

Family Aplysiidae Lamarck, 1809 

Genus Aplysia Linnaeus, 1767 

Aplysia brasiliana Rang, 1828 

Aplysia cervina (Dall & Simpson, 1901) 

Aplysia dactylomela Rang, 1828 

Aplysia juliana Quoy & Gaimard, 1832 

Aplysia morio (A. E.Verrill, 1901) 

Aplysia parvula Mørch, 1863 

Family Notarchidae Eales, 1925 

Genus Bursatella de Blainville, 1817 

Bursatella leachii pleii Rang, 1828 

Genus Dolabrifera J. E. Gray, 1847 

Dolabrifera dolabrifera (Rang, 1828) 

Genus Petalifera J. E. Gray, 1847 

Petalifera ramosa Baba, 1959 

Genus Phyllaplysia P. Fischer, 1872 

Phyllaplysia engeli Er. Marcus, 1955 

Phyllaplysia smaragda K. B. Clark, 1977 

Genus Stylocheilus Gould, 1852 

Stylocheilus citrinus (Rang, 1828) 

Stylocheilus longicauda (Quoy & Gaimard, 1825) 

Order Sacoglossa von Ihering, 1876 

Family Volvatellidae Pilsbry, 1895 

Genus Ascobulla Ev. Marcus, 1972 

Ascobulla ulla (Er. Marcus & Ev. Marcus, 1970) 

Genus Volvatella Pease, 1860 

Volvatella bermudae K. B. Clark, 1982 

Family Cylindrobullidae Thiele, 1931 

Genus Cylindrobulla P. Fischer, 1857 

Cylindrobulla beauii P. Fischer, 1857 

Family Oxynoidae H. & A. Adams, 1854 

Genus Lobiger Krohn, 1847 

Lobiger souverbii P. Fischer, 1857 

Genus Oxynoe Rafinesque, 1819 

Oxynoe antillarum Mørch, 1863 

Oxynoe azuropunctata K. R. Jensen, 1980 

Family Juliidae E. A. Smith, 1885 

Genus Berthelinia Crosse, 1875 

Berthelinia caribbea Edmunds, 1963 

Family Elysiidae H. & A. Adams, 1854 

Genus Bosellia Trinchese, 1891 

Bosellia corinneae Ev. Marcus, 1973 

Bosellia marcusi Ev. Marcus, 1972 

Bosellia mimetica Trinchese, 1891 

Genus Elysia Risso, 1818 

Elysia canguzua Er. Marcus, 1955 



Elysia chlorotica Gould, 1870 

Elysia cornigera Nuttall, 1989 

Elysia crispata Mørch, 1863 

Elysia evelinae Er. Marcus, 1957 

Elysia ornata (Swainson, 1840) 

Elysia papillosa A. E.Verrill, 1901 

Elysia patina Ev. Marcus, 1980 

Elysia serca Er. Marcus, 1955 

Elysia subornata A. E.Verrill, 1901 

Elysia tuca Ev. Marcus & Er. Marcus, 1967 

Genus Thuridilla Bergh, 1872 

Thuridilla picta (A. E.Verrill, 1901) 

Family Polybranchiidae H. & A. Adams, 1854 

Genus Caliphylla Costa, 1867 

Caliphylla mediterranea A. Costa, 1867 

Genus Cyerce Bergh, 1871 

Cyerce antillensis Engel, 1927 

Cyerce cristallina (Trinchese, 1881) 

Genus Mourgona Er. Marcus & Ev. Marcus, 1970 

Mourgona germaineae Er. Marcus & Ev. Marcus, 

1970 

Genus Polybranchia Pease, 1860 

Polybranchia viridis (Deshayes, 1857) 

Family Costasiellidae K. B. Clark, 1984 

Genus Costasiella Pruvot-Fol, 1951 

Costasiella nonatoi Ev. Marcus & Er. Marcus, 1960 

Costasiella ocellifera (Simroth, 1895) 

Family Stiligeridae Iredale & O’Donoghue, 1923 

Genus Aplysiopsis Deshayes, 1864 

Aplysiopsis zebra K. B. Clark, 1982 

Genus Ercolania Trinchese, 1872 

Ercolania coerulea Trinchese, 1893 

Ercolania funerea (A. Costa, 1867) 

Ercolania fuscata (Gould, 1870) 

Genus Hermaea Lovén, 1844 

Hermaea cruciata Gould, 1870 

Hermaea kingstoni (T. E.Thompson, 1977) 

Hermaea verticilata (Ortea, 1984) 

Genus Stiliger Ehrenberg, 1831 

Stiliger fuscovittatus Lance, 1962 

Stiliger vossi Ev. Marcus & Er. Marcus, 1960 

Order Notaspidea P. Fischer, 1883 

Family Umbraculidae Dall, 1889 

Genus Umbraculum Schumacher, 1817 

Umbraculum umbraculum (Lightfoot, 1786) 

Family Pleurobranchidae Férussac, 1822 

Genus Berthella de Blainville, 1824 

Berthella agassizii (MacFarland, 1909) 

Berthella stellata (Risso, 1826) 

Genus Pleurobranchaea Meckel in Leue, 1813 

Pleurobranchaea inconspicua Bergh, 1897 

Genus Pleurobranchus Cuvier, 1804 

Pleurobranchus areolatus Mørch, 1863 

Pleurobranchus reesi White, 1952 

Order Nudibranchia de Blainville, 1814 

Family Corambidae Bergh, 1869 

Genus Corambe Bergh, 1869 

Corambe obscura (A. E.Verrill, 1870) 

Family Goniodorididae H. & A. Adams, 1854 

Genus Okenia Leuckart & Bronn in Menke, 1830 

Okenia impexa Er. Marcus, 1957 

Okenia zoobotryon (Smallwood, 1910) 

Genus Trapania Pruvot-Fol, 1931 

Trapania dalva Ev. Marcus, 1972 

Family Aegiretidae P. Fischer, 1883 

Genus Aegires Lovén, 1844 

Aegires sublaevis Odhner, 1932 

Family Polyceratidae Alder & Hancock, 1845 

Genus Polycera Cuvier, 1817 

Polycera aurisula Er. Marcus, 1957 

Polycera hummi Abbott, 1952 

Polycera odhneri Er. Marcus, 1955 

Polycera rycia Er. Marcus & Ev. Marcus, 1970 

Genus Polycerella A. E.Verrill, 1880 

Polycerella emertoni A. E.Verrill, 1880 

Genus Thecacera Fleming, 1828 

Thecacera pennigera (Montagu, 1815) 

Family Chromodorididae Bergh, 1891 



Genus Cadlina Bergh, 1878 

Cadlina rumia Er. Marcus, 1955 

Genus Chromodoris Alder & Hancock, 1855 

Chromodoris binza Ev. Marcus & Er. Marcus, 1963 

Chromodoris clenchi (Russell, 1935) 

Chromodoris kempfi Ev. Marcus, 1970 

Chromodoris morchii Bergh, 1879 

Chromodoris nyalya Ev. Marcus & Er. Marcus, 

1967 

Genus Glossodoris Ehrenberg, 1831 

Glossodoris sedna (Ev. Marcus & Er. Marcus, 1967) 

Genus Hypselodoris Stimpson, 1855 

Hypselodoris bayeri (Ev. Marcus & Er. Marcus, 

1967) 

Hypselodoris sycilla (Bergh, 1890) 

Hypselodoris webbi (d’Orbigny, 1839) 

Family Asteronotidae Thiele, 1935 

Genus Aphelodoris Bergh, 1879 

Aphelodoris antillensis (Bergh, 1879) 

Family Dorididae Rafinesque, 1815 

Genus Doris Linnaeus, 1758 

Doris verrucosa Linnaeus, 1758 

Genus Siraius Er. Marcus, 1955 

Siraius bicolor (Bergh, 1884) 

Siraius kyolis Ev. Marcus & Er. Marcus, 1967 

Family Dendrodorididae O’Donoghue, 1924 

Genus Dendrodoris Ehrenberg, 1831 

Dendrodoris krebsii (Mørch, 1863) 

Dendrodoris warta Ev. Marcus & Gallagher, 1976 

Genus Doriopsilla Bergh, 1880 

Doriopsilla pharpa Er. Marcus, 1961 

Family Phyllidiidae Rafinesque, 1815 

Genus Ceratophyllidia Eliot, 1903 

Ceratophyllidia papilligera (Bergh, 1890) 

Family Archidorididae Bergh, 1892 

Genus Atagema J. E. Gray, 1850 

Atagema prea (Ev. Marcus & Er. Marcus, 1967) 

Family Discodorididae Bergh, 1891 

Genus Anisodoris Bergh, 1898 

Anisodoris worki Ev. Marcus & Er. Marcus, 1967 

Genus Discodoris Bergh, 1877 

Discodoris alba White, 1952 

Discodoris evelinae Er. Marcus, 1955 

Discodoris mortenseni Ev. Marcus & Er. Marcus, 

1963 

Discodoris phoca Ev. Marcus & Er. Marcus, 1967 

Discodoris purcina Ev. Marcus & Er. Marcus, 1967 

Genus Geitodoris Bergh, 1892 

Geitodoris immunda Bergh, 1894 

Geitodoris pusae (Er. Marcus, 1955) 

Genus Peltodoris Bergh, 1880 

Peltodoris greeleyi MacFarland, 1909 

Genus Taringa Er. Marcus, 1955 

Taringa telopia Er. Marcus, 1955 

Family Platydorididae Bergh, 1891 

Genus Platydoris Bergh, 1877 

Platydoris angustipes (Mørch, 1863) 

Family Tritoniidae Menke, 1828 

Genus Marionia Vayssière, 1877 

Marionia tedi Ev. Marcus, 1983 

Genus Tritonia Cuvier, 1798 

Tritonia bayeri Ev. Marcus & Er. Marcus, 1967 

Genus Tritonidoxa Bergh, 1907 

Tritonidoxa wellsi (Er. Marcus, 1961) 

Genus Tritoniopsis Eliot, 1905 

Tritoniopsis frydis Er. Marcus & Ev. Marcus, 1970 

Family Bornellidae P. Fischer, 1883 

Genus Bornella A. Adams & Reeve, 1848 

Bornella calcarata Mørch, 1863 

Family Lomanotidae Bergh, 1892 

Genus Lomanotus Vérany, 1844 

Lomanotus vermiformis Eliot, 1908 

Family Scyllaeidae Rafinesque, 1815 

Genus Scyllaea Linnaeus, 1758 

Scyllaea pelagica Linnaeus, 1758 



Family Dotoidae J. E. Gray, 1853 

Genus Doto Oken, 1815 

Doto caramella Er. Marcus, 1957 

Doto chica Ev. Marcus & Er. Marcus, 1960 

Doto coronata (Gmelin, 1791) 

Doto divae Ev. Marcus & Er. Marcus, 1960 

Doto pita Er. Marcus, 1955 

Doto uva Er. Marcus, 1955 

Genus Miesea Er. Marcus, 1961 

Miesea evelinae (Er. Marcus, 1957) 

Family Arminidae Pruvot-Fol, 1927 

Genus Armina Rafinesque, 1814 

Armina wattla Ev. Marcus & Er. Marcus, 1967 

Family Janolidae Pruvot-Fol, 1954 

Genus Janolus Bergh, 1884 

Janolus comis Er. Marcus, 1955 

Janolus mucloc (Er. Marcus, 1958) 

Family Flabellinidae Bergh, 1889 

Genus Flabellina Voigt, 1834 

Flabellina engeli Ev. Marcus & Er. Marcus, 1968 

Family Eubranchidae Odhner, 1934 

Genus Eubranchus Forbes, 1838 

Eubranchus coniclus (Er. Marcus, 1958) 

Family Tergipedidae Bergh, 1889 

Genus Catriona Winckworth, 1941 

Catriona maua Ev. Marcus & Er. Marcus, 1960 

Catriona oba Ev. Marcus, 1970 

Genus Cuthona Alder & Hancock, 1855 

Cuthona caerulea (Montagu, 1804) 

Cuthona perca (Er. Marcus, 1958) 

Cuthona tina (Er. Marcus, 1957) 

Genus Tenellia A. Costa, 1866 

Tenellia fuscata (Gould, 1870) 

Family Fionidae Alder & Hancock, 1855 

Genus Fiona Alder & Hancock in Forbes & Hanley, 

1851 

Fiona pinnata (Eschscholtz, 1831) 

Family Facelinidae Bergh in Carus, 1889 

Genus Austraeolis Burn, 1962 

Austraeolis catina Ev. Marcus & Er. Marcus, 1967 

Genus Cratena Bergh, 1864 

Cratena pilata (Gould, 1870) 

Genus Dondice Er. Marcus, 1958 

Dondice occidentalis (Engel, 1925) 

Genus Favorinus M. E. Gray, 1850 

Favorinus auritulus Er. Marcus, 1955 

Genus Godiva Macnae, 1954 

Godiva rubrolineata Edmunds, 1964 

Genus Learchis Bergh, 1889 

Learchis poica Ev. Marcus & Er. Marcus, 1960 

Genus Phidiana J. E. Gray, 1850 

Phidiana lynceus Bergh, 1867 

Family Aeolidiidae d’Orbigny, 1834 

Genus Spurilla Bergh, 1864 

Spurilla neapolitana (delle Chiaje, 1823) 

Spurilla verrucicornis (A. Costa, 1867) 

Family Glaucidae Menke, 1828 

Genus Glaucus Förster, 1777 

Glaucus atlanticus Förster, 1777 

Subclass Pulmonata Cuvier, 1817 

Order Archaeopulmonata Morton, 1955 

Family Ellobiidae H. & A. Adams, 1854 

Genus Blauneria Shuttleworth, 1854 

Blauneria heteroclita (Montagu, 1808) 

Genus Creedonia Martins, 1996 

Creedonia succinea (Pfeiffer, 1854) 

Genus Ellobium Röding, 1798 

Ellobium dominicense (Férussac, 1821) 

Genus Laemodonta Philippi, 1846 

Laemodonta cubensis (Pfeiffer, 1854) 

Genus Leuconopsis Hutton, 1884 

Leuconopsis novimundi (Pilsbry & McGinty, 1949) 

Genus Melampus Montfort, 1810 



Subgenus Melampus Montfort, 1810 

Melampus (Melampus) bidentatus Say, 1822 

Melampus (Melampus) coffeus (Linnaeus, 1758) 

Subgenus Detracia J. E. Gray, 1840 

Melampus (Detracia) bullaoides (Montagu, 1808) 

Melampus (Detracia) floridanus Pfeiffer, 1856 

Melampus (Detracia) monile (Bruguière, 1789) 

Melampus (Detracia) morrisoni Martins, 1996 

Genus Microtralia Dall, 1894 

Microtralia occidentalis (Pfeiffer, 1854) 

Genus Pedipes Scopoli, 1777 

Pedipes mirabilis (Mühlfeld, 1816) 

Pedipes ovalis (C. B. Adams, 1849) 

Genus Tralia J. E. Gray, 1840 

Tralia ovula (Bruguière, 1789) 

Order Basommatophora Schmidt, 1855 

Family Siphonariidae J. E. Gray, 1840 

Genus Siphonaria Sowerby, 1823 

Siphonaria alternata (Say, 1826) 

Siphonaria pectinata (Linnaeus, 1758) 

Genus Williamia Monterosato, 1884 

Williamia krebsii (Mørch, 1877) 

Family Trimusculidae Zilch, 1959 

Genus Trimusculus Schmidt, 1818 

Trimusculus carinatus (Dall, 1870) 

Order Systellommatophora Pilsbry, 1948 

Family Onchidiidae J. E. Gray, 1824 

Genus Onchidella M. E. Gray, 1850 

Onchidella brattstroemi Ev. Marcus, 1978 

Onchidella floridana (Dall, 1885) 

Onchidella wah Ev. Marcus, 1978 

Class Cephalopoda Cuvier, 1797 

Squids and Octopuses 

Order Sepioidea Leach, 1817 

Family Spirulidae Rafinesque, 1815 

Genus Spirula Lamarck, 1799 

Spirula spirula (Linnaeus, 1758) 

Family Sepiolidae Leach, 1817 

Genus Semirossia Steenstrup, 1881 

Semirossia tenera (A. E.Verrill, 1880) 

Order Teuthoidea Owen, 1836 

Family Loliginidae Lesueur, 1821 

Genus Loligo Schneider, 1784 

Loligo pealeii Lesueur, 1821 

Loligo pleii de Blainville, 1823 

Genus Lolliguncula Steenstrup, 1881 

Lolliguncula brevis (de Blainville, 1823) 

Genus Pickfordiateuthis Voss, 1953 

Pickfordiateuthis pulchella Voss, 1953 

Genus Sepioteuthis de Blainville, 1824 

Sepioteuthis sepioidea (de Blainville, 1823) 

Order Octopoda Rafinesque, 1815 

Family Octopodidae Rafinesque, 1815 

Genus Octopus Lamarck, 1798 

Octopus briareus Robson, 1929 

Octopus burryi Voss, 1953 

Octopus defilippi Vérany, 1851 

Octopus filosus Howell, 1868 

Octopus joubini Robson, 1929 

Octopus macropus Risso, 1826 

Octopus vulgaris Lamarck, 1798 

Family Argonautidae Rafinesque, 1815 

Genus Argonauta Linnaeus, 1758 

Argonauta argo Linnaeus, 1758 

Argonauta hians Solander, 1786 

Argonauta nodosus Solander, 1786 

Pelagic Mollusks 

Many mollusks that live in the pelagic realm, either 

free-swimming or on the alga Sargassum, wash up conspicuously 

on Florida’s shores. Most of these species 

are evidenced in the nearshore environment principally 

by their shells and/or dead or dying animals (e.g., the 

gastropods Janthina, Recluzia, Fiona, and Glaucus and the 

cephalopod Spirula) and egg cases (“paper nautilus”: 

cephalopods, genus Argonauta), but some (e.g., the gastropods 

Litiopa and Scyllaea) may drop off Sargassum 

near shore and live for brief periods among the shallow-water 

benthos. All of these species are included in 

the general list.That list also includes Creseis acicula, a 

pteropod snail with a glassy, needle-like shell that can 

become entangled in swimwear, causing a type of 

“swimmer’s itch.” The species sometimes occurs in 



great densities in coastal waters, and windrows of its 

shells are occasionally found along Florida beaches. 

Consult Lalli and Gilmer (1989) for a comprehensive 

summary of these and the following species of pelagic 

snails. 

The gastropods listed hereafter lead exclusively 

pelagic lives in the open ocean. Many of the species 

occur throughout the Atlantic Ocean, and some have 

world-wide distributions. Although they occur in waters 

that may intrude into coastal environments in 

areas such as southeastern and northwestern Florida, 

representatives of these species are virtually never 

collected except deliberately by those sampling plankton 

and nekton. Pelagic cephalopods, principally 

squids, are also encountered occasionally in Florida 

coastal waters, but most of these occurrences are accidental, 

and the many such deep-water species that 

may be involved are not listed here. 

Class Gastropoda Cuvier, 1797 

Order Neotaenioglossa Haller, 1882 

Family Atlantidae Rang, 1829 

Genus Atlanta Lesueur, 1817 

Atlanta brunnea J. E. Gray, 1850 

Atlanta helicinoides J. E. Gray, 1850 

Atlanta inclinata J. E. Gray, 1850 

Atlanta lesueurii J. E. Gray, 1850 

Atlanta peronii Lesueur, 1817 

Genus Oxygyrus Benson, 1835 

Oxygyrus keraudrenii (Lesueur, 1817) 

Genus Protatlanta Tesch, 1908 

Protatlanta souleyeti (E. A. Smith, 1888) 

Family Carinariidae de Blainville, 1818 

Genus Cardiapoda d’Orbigny, 1835 

Cardiapoda placenta (Lesson, 1830) 

Genus Carinaria Lamarck, 1801 

Carinaria lamarcki Péron & Lesueur, 1810 

Family Pterotracheidae Gray, 1840 

Genus Firoloida Lesueur, 1817 

Firoloida demarestia Lesueur, 1817 

Genus Pterotrachea Forskål, 1775 

Pterotrachea hippocampus Philippi, 1836 

Pterotrachea scutata Gegenbaur, 1855 

Order Thecosomata de Blainville, 1824 

Family Limacinidae de Blainville, 1823 

Genus Limacina Bosc, 1817 

Limacina bulimoides (d’Orbigny, 1836) 

Limacina inflata (d’Orbigny, 1836) 

Limacina lesueurii (d’Orbigny, 1836) 

Limacina trochiformis (d’Orbigny, 1836) 

Family Cavoliniidae H. & A. Adams, 1854 

Genus Cavolinia Abildgaard, 1791 

Cavolinia gibbosa (d’Orbigny, 1836) 

Cavolinia inflexa (Lesueur, 1813) 

Cavolinia tridentata (Niebuhr, 1775) 

Cavolinia uncinata (Rang, 1829) 

Genus Clio Linnaeus, 1767 

Clio chaptalii J. E. Gray, 1850 

Clio cuspidata (Bosc, 1802) 

Clio polita (Pelseneer, 1888) 

Clio pyramidata Linnaeus, 1767 

Clio recurva (Children, 1823) 

Genus Creseis Rang, 1828 

Creseis virgula (Rang, 1828) 

Genus Cuvierina Boas, 1886 

Cuvierina columnella (Rang, 1827) 

Genus Diacavolinia Van der Spoel, 1987 

Diacavolinia deblainvillei van der Spoel, Bleeker 

& Kobayashi, 1993 

Genus Diacria J. E. Gray, 1842 

Diacria trispinosa (de Blainville, 1821) 

Genus Hyalocylis Fol, 1875 

Hyalocylis striata (Rang, 1828) 

Genus Styliola Lesueur, 1825 

Styliola subula (Quoy & Gaimard, 1827) 

Family Peraclididae C. W. Johnson, 1915 

Genus Peracle Forbes, 1844 

Peracle bispinosa Pelseneer, 1888 

Peracle diversa Monterosato, 1875 

Peracle reticulata (d’Orbigny, 1836) 

Peracle triacantha P. Fischer, 1882 

Order Gymnosomata de Blainville, 1824 

Family Clionidae Rafinesque, 1815 

Genus Paraclione Tesch, 1903 



Paraclione longicaudata (Souleyet, 1852) 

Family Cliopsidae O. G. Costa, 1873 

Genus Cliopsis Troschel, 1854 

Cliopsis krohnii Troschel, 1854 

Family Notobranchaeidae Pelseneer, 1886 

Genus Notobranchaea Pelseneer, 1886 

Notobranchaea macdonaldi Pelseneer, 1886 

Genus Prionoglossa Tesch, 1950 

Prionoglossa tetrabranchiata (Bonnevie, 1913) 

Family Pneumodermatidae Latreille, 1825 

Genus Pneumoderma Cuvier, 1805 

Pneumoderma mediterraneum (van Beneden, 1838) 

ABBOTT, R.T. 1944.The genus Modulus in the western 

Atlantic. Johnsonia 1(14): 1–6. 

ABBOTT, R. T. 1949. A new Florida species of the 

tectibranch genus Pleurobranchus. The Nautilus 62(3): 

73–78, pl. 3. 

ABBOTT, R.T. 1950.The genus Cyclostrema in the western 


ABBOTT, R.T. 1950.The genera Xancus and Vasum in the 

western Atlantic. Johnsonia 2(28): 201–219. 

ABBOTT, R. T. 1951. New deep-water olivellas from 

Florida, with notes on the O. jaspidea nivea complex.The 

Nautilus 64(4): 110–116, pl. 7. 

ABBOTT, R.T. 1952.Two new opisthobranch mollusks 

from the Gulf of Mexico belonging to the genera Pleurobranchaea 

and Polycera. Florida State University Studies, 

No. 7: 1–7. 

ABBOTT, R. T. 1954. American Seashells. D. Van Nostrand 

Company, Inc., Princeton, New Jersey. xiv + 541 

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ABBOTT, R. T. 1954. New Gulf of Mexico gastropods 

(Terebra and Ocenebra).The Nautilus 68(2): 37–44, pl. 2. 

ABBOTT, R. T. 1954. The habits and occurrence of the 

References 

Pneumoderma violaceum violaceum (d’Orbigny, 

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Genus Pneumodermopsis Keferstein, 1862 

Pneumodermopsis macrochira (Meisenheimer, 

1905) 

Pneumodermopsis paucidens (Boas, 1886) 

Family Thliptodontidae Kwietniewski, 1910 

Genus Thliptodon Boas, 1886 

Thliptodon diaphanus (Meisenheimer, 1903) 

Order Nudibranchia Cuvier, 1817 

Family Phylliroidae Menke, 1830 

Genus Phylliroe Péron & Lesueur, 1810 

Phylliroe atlantica Bergh, 1871 

Phylliroe bucephalum Péron & Lesueur, 1810 

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ANONYMOUS. 1894. Argonauta argo on the east Florida 

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ANONYMOUS. 1896. Callista varians Hanley in eastern 

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BAKER, H. B. 1957. Fasciolaria distans Lamarck. Minutes 

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BALES, B. R. 1942. A new subspecies of Strombus raninus 

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Methods 

Checklist of Shallow-Water Marine Polychaetous 

Annelida of Florida 

Thomas H. Perkins 

This checklist of Florida shallow-water polychaetes, 

which current research is showing to be a weakly supported 

monophyletic group (Rouse and Fauchald, 

1997), was derived from a variety of sources. These 

sources include 1) the extensive systematic literature 

on Florida polychaetes, which begins primarily with the 

publication on Florida annelids, in German, of Ehlers 

(1887), and which has not been systematically synthesized 

to allow for rapid identification; 2) the published 

results of ecological surveys; 3) data from specimen collections; 

4) unpublished data and specimens examined 

from recently completed estuarine environmentalassessment 

programs (EMAP) conducted in Florida by 

the U.S. Environmental Protection Agency and the 

National Oceanic and Atmospheric Administration 

(NOAA) Coastal Oceans Program; and 5) other published 

and unpublished information from surveys conducted 

by the State of Florida and by qualified consultants 

for the state, the federal government, and private 

organizations. As a result, reliable polychaete 

identification data are available from almost all areas 

of the state. 

The primary specimen repository from which 

species names were extracted for inclusion on this list 

is the Marine Specimen Collection housed at the 

Florida Marine Research Institute, St. Petersburg. Data 

from the Harbor Branch Oceanographic Museum, Fort 

Pierce, Florida, were also used. Other specimens from 

Florida, some of which have been examined but data 

for which were not generally used in compiling the list, 

are housed at the Florida Department of Environmental 

Protection’s central laboratory, Tallahassee, 

and district offices; in the Texas A&M Oceanographic 

Collection, College Station, Texas; at Gulf Coast Research 

Laboratory, Ocean Springs, Mississippi; at Barry 

Vittor & Associates, Inc., Mobile, Alabama; in collections 

of the National Museum of Natural History, 

Washington, D.C.; and in collections of other national 

and international museums. 

Citations for most of the literature listed as helpful 

in identifying Florida polychaetes (see References) 

were extracted from the Ward and Fauchald bibliography 

(Ward and Fauchald, 1997) available on the internet. 

Additional, more recent citations and local references 

not included in that bibliography are also 

listed among the polychaete references, resulting in 

about 500 literature citations that are pertinent to the 

identification of Florida polychaetes. 

Criteria used to determine which species names to 

include in the list were various combinations of the following: 

1) location in estuarine and near-coastal waters 

of the state in depths of about 37 m or less; 2) a 

moderately skeptical determination by me of the reliability 

of an identification, continued availability of 

vouchers that will allow specimens to be reliably identified, 

or both; and 3) for published “lettered” or unnamed 

species, the availability of a description and 

voucher specimens that, with locality data, will allow 

species to subsequently be identified or described. A 

few taxa listed as “Genus name” cf. “species name,” 

based on specimens in the FMRI Marine Specimen Collection 

that possibly are that species and may not deserve 

the “cf.”designation have been included on the 

list. 

Considering the state of knowledge of Florida 

polychaetes, this list is a work in progress.There is no 

doubt that there are names on the list that will prove 

to be incorrect, and some species may be listed under 

two or more names.There are specimens available of 

many undescribed species in Florida waters that are 

not listed, just as there are species that remain undiscovered. 

The classification of the Polychaeta is currently 

unstable, especially at categories higher than family. 

The system in which the species are reported here is 

based on the classical system of Pettibone (1982). Another 

commonly used but different system, that of 

Fauchald (1977a), was used by the authors of the Taxonomic 

Guide to the Polychaetes of the Northern Gulf of 

Mexico (Uebelacker and Johnson, 1984). A new, welldocumented 

classification system for family groups 

and higher categories has been proposed (Rouse and 

Fauchald, 1997). Rouse and Fauchald state that it is 

unlikely that any new classification will have any 

longevity, but their system should provide the basis for 

a stable and phylogenetically correct classification in 

the future. 


Many biologists have provided specimens over the 

years. In particular, I thank Robert Ernest, Ecological 


Polychaeta—T. H. Perkins Macroinvertebrate Checklists Camp et al. 1998 

Associates (formerly Applied Biology, Inc.), Jensen 

Beach, Florida; Harvey Rudolph, Palm Beach County 

Government, formerly Florida Department of Environmental 

Protection (FDEP); Steve Kent, Florida Department 

of Environmental Protection, Orlando; David 

Karlen, Environmental Protection Commission of Hillsborough 

County; Vicki Whiting, FDEP central laboratory,Tallahassee; 

Jerry McLelland, Gulf Coast Research 

Laboratory; Dr. Gary Gaston and Carol Cleveland, 

University of Mississippi; and Dr. Marianne Whitehurst 

and Tim Thibault, Barry Vittor & Assoc., Inc. Bob Ernest 

also provided data on specimen identifications from 

many studies conducted by Applied Biology, Inc. Harvey 

Rudolph provided data on specimens he and others 

identified from Biscayne Bay. Jerry McLelland and 

Sarah LeCroy, Gulf Coast Research Laboratory, provided 

data for their specimens collected at Perdido 

Key, Florida, during studies for the National Park Service; 

David Karlen provided data for specimens from 

Tampa Bay collected for the Tampa Bay National Estuary 

Program.Tom Dix, Florida Marine Research Institute 

(FMRI), provided data for polychaetes identified 

Phylum Annelida 

Class Polychaeta 

Order Phyllodocida 

Family Phyllodocidae Örsted, 1843 

Genus Eulalia Savigny, 1822 

Eulalia hutchinsonensis (Perkins, 1984) 

Eulalia myriacyclum (Schmarda, 1861) 

Genus Eumida Malmgren, 1865 

Eumida sanguinea (Örsted, 1843) 

Genus Eteone Savigny, 1822 

Subgenus Hypereteone Bergström, 1914 

Eteone (Hypereteone) alba Webster, 1879 

Eteone (Hypereteone) foliosa Quatrefages, 1866 

Eteone (Hypereteone) heteropoda Hartman, 1951 

Genus Hesionura Hartmann-Schröder, 1958 

Hesionura coineaui (Laubier, 1962) 

Hesionura laubieri ? (Hartmann-Schröder, 1963) 

Genus Mystides Théel, 1879 

Mystides borealis Théel, 1879 

(= M. nr. borealis, fide Eibye-Jacobsen, 1992) 

Genus Nereiphylla Blainville, 1828 

Nereiphylla castanea (Marenzeller, 1879) 

from FMRI studies in Florida Bay. Marianne Whitehurst 

and Tim Thibault provided data from studies conducted 

by Barry Vittor & Assoc., Inc., for NOAA in Biscayne 

Bay and Florida Bay and provided updates to the 

nomenclature of species reported in the Taxonomic 

Guide to the Polychaetes of the Northern Gulf of Mexico (Uebelacker 

and Johnson, 1984). Gary Gaston and Carol 

Cleveland also provided information from their studies 

for NOAA in Florida Bay and Biscayne Bay and 

from their Florida-West Indian Province EMAP studies 

for the U.S. Environmental Protection Agency (EPA) 

laboratory at Gulf Breeze, Florida. Dr. John Macauley, 

EPA, Gulf Breeze, Florida, and Linda Harwell, Johnson 

Controls, Inc., Gulf Breeze, Florida, the EPA data consultant, 

provided detailed data for specimens collected 

in the Florida area of the Louisianan Province estuarine 

EMAP program. Dr. Kristian Fauchald, National 

Museum of Natural History, Washington, D.C., Tim 

Thibault, Marianne Whitehurst, Gary Gaston, and 

Carol Cleveland reviewed draft copies of the species 

list and corrected several of my mistakes. All are gratefully 

acknowledged. 

Checklist of the Polychaetous Annelida 

Nereiphylla fragilis (Webster, 1879) 

Nereiphylla nr. paretti of Hartman, 1951 

Nereiphylla sp. A of Gathof, 1984 

(as Genetyllis sp. A) 

Genus Paranaitis Southern, 1914 

Paranaitis capensis (Day, 1960) 

Paranaitis gardineri Perkins, 1984 

Paranaitis papillosa (Ehlers, 1887) 

Paranaitis speciosa (Webster, 1879) 

Genus Phyllodoce Lamarck, 1818 

Subgenus Anaitides Czerniavsky, 1882 

Phyllodoce (Anaitides) erythrophylla (Schmarda, 

1861) 

Phyllodoce (Anaitides) groenlandica (Örsted, 1843) 

Phyllodoce (Anaitides) longipes (Kinberg, 1866) 

Phyllodoce (Anaitides) maderiensis Langerhans, 

1880 

Phyllodoce (Anaitides) mucosa (Örsted, 1843) 

Subgenus Phyllodoce Lamarck, 1818 

Phyllodoce (Phyllodoce) arenae Webster, 1879 

Phyllodoce (Phyllodoce) tortugae Treadwell, 1917 

Genus Pirakia Bergström, 1914 

Pirakia punctifera (Grube, 1860) 

Genus Protomystides Czerniavsky, 1882 


Camp et al. 1998 Macroinvertebrate Checklists T. H. Perkins—Polychaeta 

Protomystides bidentatus (Langerhans, 1880) 

Genus Pterocirrus Claparède, 1868 

Pterocirrus foliosus (Treadwell, 1924) 

Genus Sige Malmgren, 1865 

Sige parvicirrus (Perkins, 1984) 

Family Lacydoniidae Bergström, 1914 

Genus Lacydonia Marion & Bobretzsky, 1875 

Lacydonia miranda Marion & Bobretzky, 1875 

Family Glyceridae Grube, 1850 

Genus Glycera Lamarck, 1818 

Glycera abranchiata Treadwell, 1900 

Glycera americana Leidy, 1855 

Glycera capitata Örsted, 1843 

Glycera dibranchiata Ehlers, 1868 

Glycera oxycephala Ehlers, 1887 

Glycera papillosa Grube, 1857 

Glycera robusta Ehlers, 1868 

Glycera sphyrabrancha Schmarda, 1861 

Glycera tesselata Grube, 1863 

Glycera sp. A of Gilbert, 1984 

Glycera sp. C of Gilbert, 1984 

Glycera sp. D of Gilbert, 1984 

Glycera sp. E of Gilbert, 1984 

Glycera sp. F of Gilbert, 1984 

Genus Hemipodus Quatrefages, 1866 

Hemipodus roseus Quatrefages, 1866 

Family Goniadidae Kinberg, 1866 

Genus Glycinde Müller, 1858 

Glycinde nordmanni (Malmgren, 1866) 

Glycinde solitaria (Webster, 1879) 

Genus Goniada Audouin & Milne Edwards, 1833 

Goniada acicula Hartman, 1940 

Goniada littorea Hartman, 1950 

Goniada maculata Örsted, 1843 

Goniada multidentopsis Perkins, 1980 

Goniada teres Treadwell, 1931 

Genus Goniadella Hartman, 1950 

Goniadella sp. A of Gilbert, 1984 

Genus Goniadides Hartmann-Schröder, 1960 

Goniadides carolinae Day, 1973 

Genus Progoniada Hartman, 1965 

Progoniada regularis Hartman, 1965 

Family Sphaerodoridae Malmgren, 1867 

Genus Ephesiella Chamberlin, 1919 

Ephesiella mammifera Fauchald, 1974 

Genus Sphaerodoridium Lützen, 1961 

Sphaerodoridium lutzeni Kudenov, 1987 

Family Hesionidae Grube, 1850 

Genus Gyptis Marion & Bobretzky, 1875 

Gyptis crypta Pleijel, 1993 

Gyptis plurisetis Hilbig, 1992 

Genus Hesione Lamarck, 1818 

Hesione picta Müller, 1858 

Genus Heteropodarke Hartmann-Schröder, 1962 

Heteropodarke formalis Perkins, 1984 

Heteropodarke lyonsi Perkins, 1984 

Genus Leocrates Kinberg, 1866 

Leocrates chinensis Kinberg, 1866 

Genus Microphthalmus Metschnikow, 1865 

Microphthalmus aberrans (Webster & Benedict, 

1887) 

Microphthalmus hamosus Westheide, 1982 

Microphthalmus hartmanae Westheide, 1977 

Microphthalmus sczelkowii Mecznikow, 1865 

Genus Ophiodromus Sars, 1861 

Ophiodromus nr. guanicus of Hartman, 1951 

(as Podarke nr. guanica) 

Ophiodromus obscurus (Verrill, 1873) 

Ophiodromus obscurus of Uebelacker, 1984 

(as Podarke obscura) 

Ophiodromus sp. A of Uebelacker, 1984 

Ophiodromus sp. B of Uebelacker, 1984 

(as Podarke sp. A) 

Genus Parahesione Pettibone, 1966 

Parahesione luteola (Webster, 1879) 

Genus Podarkeopsis Laubier, 1961 

Podarkeopsis levifuscina Perkins, 1984 

Genus Psamathe Johnston, 1836 

Psamathe haploseta Perkins, 1984 

Psamathe sp. of Perkins, 1984 

(as Kefersteinia sp.) 



Genus Syllidia Quatrefages, 1866 

Syllidia armata Quatrefages, 1866 

Family Pilargidae Saint Joseph, 1899 

Genus Ancistrosyllis McIntosh, 1879 

Ancistrosyllis carolinensis Gardiner, 1976 

Ancistrosyllis commensalis Gardiner, 1976 

Ancistrosyllis hartmanae Pettibone, 1966 

Ancistrosyllis jonesi Pettibone, 1966 

Ancistrosyllis papillosa (Jones, 1961) 

Ancistrosyllis sp. C of Wolf, 1984 

Genus Cabira Webster, 1879 

Cabira incerta Webster, 1879 

Genus Glyphohesione Friedrich, 1950 

Glyphohesione longicirrata Licher, 1994 

Genus Litocorsa Pearson, 1970 

Litocorsa antennata Wolf, 1986 

Genus Parandalia Emerson & Fauchald, 1971 

Parandalia americana (Hartman, 1947) 

Parandalia tricuspis (Müller, 1858) 

Genus Pilargis Saint Joseph, 1899 

Pilargis berkeleyae Monro, 1933 

Pilargis pacifica Zachs, 1933 

Pilargis verrucosa Saint Joseph, 1899 

Pilargis sp. A of Uebelacker, 1984 

Genus Sigambra Müller, 1858 

Sigambra bassi (Hartman, 1945) 

Sigambra tentaculata (Treadwell, 1941) 

Sigambra wassi Pettibone, 1966 

Genus Synelmis Chamberlin, 1919 

Synelmis acuminata Wolf, 1986 

Synelmis ewingi Wolf, 1986 

Family Syllidae Grube, 1850 

Genus Autolytus Grube, 1850 

Autolytus brevicirrata Winternitz, 1936 

Autolytus dentalius Imajima, 1966 

Autolytus sp. A of Uebelacker, 1984 

Genus Branchiosyllis Ehlers, 1887 

Branchiosyllis diazi Rioja, 1958 

Branchiosyllis exilis (Gravier, 1900) 

Branchiosyllis oculata Ehlers, 1887 

Genus Brania Quatrefages, 1866 

Brania gallagheri Perkins, 1981 

Brania wellfleetensis Pettibone, 1956 

Genus Dentatisyllis Perkins, 1981 

Dentatisyllis carolinae (Day, 1973) 

Genus Dioplosyllis Gidholm, 1962 

Dioplosyllis octodentata Perkins, 1981 

Genus Eurysyllis Ehlers, 1864 

Eurysyllis tuberculata Ehlers, 1864 

Genus Eusyllis Malmgren, 1867 

Eusyllis heterocirrata Hartmann-Schröder, 1959 

Eusyllis lamelligera Marion & Bobretzky, 1975 

Genus Exogone Örsted, 1845 

Subgenus Exogone Örsted, 1845 

Exogone (Exogone) arenosa Perkins, 1981 

Exogone (Exogone) breviantennata Hartmann- 

Schröder, 1959 

Exogone (Exogone) dispar (Webster, 1879) 

Exogone (Exogone) lourei Berkeley & Berkeley, 

1938 

Exogone (Exogone) naidina Örsted, 1845 

Exogone (Exogone) pseudolourei San Martín, 1991 

Exogone (Exogone) rolani San Martín, 1991 

Exogone (Exogone) verugera (Claparède, 1868) 

Subgenus Parexogone Caullery & Mesnil, 1916 

Exogone (Parexogone) atlantica Perkins, 1981 

Exogone (Parexogone) caribensis San Martín, 1991 

Genus Grubeosyllis Verrill, 1900 

Grubeosyllis clavata (Claparède, 1863) 

Grubeosyllis nitidula Verrill, 1900 

Grubeosyllis rugulosa Verrill, 1900 

Grubeosyllis swedmarki (Gidholm, 1962) 

Grubeosyllis vietezi (San Martín, 1984) 

Genus Haplosyllides Augener, 1922 

Haplosyllides floridanus Augener, 1922 

Genus Haplosyllis Langerhans, 1879 

Haplosyllis spongicola (Grube, 1855) complex 

Genus Levidorum Hartman, 1967 

Levidorum hartmanae Perkins, 1987 

Levidorum pettiboneae Perkins, 1987 

Genus Odontosyllis Claparède, 1863 

Odontosyllis enopla Verrill, 1900 

Odontosyllis fulgurans (Audouin & Milne Edwards, 

1833) 

Odontosyllis longigulata Perkins, 1981 



Odontosyllis octodentata Treadwell, 1917 

Genus Opistosyllis Langerhans, 1879 

Opisthosyllis brunnea Langerhans, 1879 

Genus Opisthodonta Langerhans, 1879 

Opisthodonta sp. A of Uebelacker, 1984 

Opisthodonta sp. B of Uebelacker, 1984 

Genus Parapionosyllis Fauvel, 1923 

Parapionosyllis longicirrata (Webster & Benedict, 

1884) 

Parapionosyllis uebelackerae San Martín, 1991 

Parapionosyllis floridana San Martín, 1991 

Genus Parasphaerosyllis Monro, 1937 

Parasphaerosyllis indica Monro, 1937 

Genus Pionosyllis Malmgren, 1867 

Pionosyllis aciculagrossa San Martín, 1990 

Pionosyllis gesae Perkins, 1981 

Pionosyllis spinisetosa San Martín, 1990 

Pionosyllis weismanni Langerhans, 1879 

Pionosyllis sp. of Taylor, 1971 

Genus Plakosyllis Hartmann-Schröder, 1956 

Plakosyllis quadrioculata Perkins, 1981 

Genus Pseudosyllides Augener, 1927 

Pseudosyllides curacaoensis Augener, 1927 

Genus Sphaerosyllis Claparède, 1963 

Sphaerosyllis aciculata Perkins, 1981 

Sphaerosyllis bilobata Perkins, 1981 

Sphaerosyllis brevidentata Perkins, 1981 

Sphaerosyllis glandulata Perkins, 1981 

Sphaerosyllis labyrinthophila Gardiner & Wilson, 

1979 

Sphaerosyllis magnidentata Perkins, 1981 

Sphaerosyllis perkinsi Riser, 1991 

Sphaerosyllis piriferopsis Perkins, 1981 

Sphaerosyllis riseri Perkins, 1981 

Sphaerosyllis taylori Perkins, 1981 

Genus Streptospinigera Kudenov, 1983 

Streptospinigera heteroseta Kudenov, 1983 

Genus Streptosyllis Webster & Benedict, 1884 

Streptosyllis pettiboneae Perkins, 1981 

Streptosyllis verrilli (Moore, 1908) 

Genus Syllides Örsted, 1845 

Syllides bansei Perkins, 1981 

Syllides floridanus Perkins, 1981 

Syllides fulvus (Marion & Bobretzky, 1875) 

Syllides gomezi San Martín, 1990 

Syllides sp. A of Uebelacker, 1984 

Genus Syllis Lamarck, 1818 

Subgenus Ehlersia Quatrefages, 1866 

Syllis (Ehlersia) cf. cornuta of San Martín, 1992 

Syllis (Ehlersia) mexicana (Rioja, 1960) 

Subgenus Syllis Lamarck, 1818 

Syllis (Syllis) gracilis Grube, 1840 

Syllis (Syllis) spongiphila Verrill, 1885 

Subgenus Typosyllis Langerhans, 1879 

Syllis (Typosyllis) aciculata (Treadwell, 1945) 

Syllis (Typosyllis) alosae San Martín, 1992 

Syllis (Typosyllis) alternata Moore, 1908 

Syllis (Typosyllis) amica (Quatrefages, 1866) 

Syllis (Typosyllis) broomensis (Hartmann- 

Schröder, 1979) 

Syllis (Typosyllis) cincinnata Verrill, 1900 

Syllis (Typosyllis) corallicoides Augener, 1922 

Syllis (Typosyllis) corallicola Verrill, 1900 

Syllis (Typosyllis) hyalina Grube, 1863 

Syllis (Typosyllis) cf. lutea (Hartmann-Schröder, 

1960) 

Syllis (Typosyllis) ortezi San Martín, 1992 

Syllis (Typosyllis) prolifera Krohn, 1852 

Syllis (Typosyllis) sardai San Martín, 1992 

Syllis (Typosyllis) tigrinoides Augener, 1922 

Syllis (Typosyllis) tortugaensis Augener, 1922 

(= Syllis (Typosyllis) armillaris of Uebelacker, 

1982) 

Syllis (Typosyllis) tortugaensis of Monro, 1933 

Syllis (Typosyllis) variegata Grube, 1860 

Syllis (Typosyllis) vittata Grube, 1840 

Syllis (Typosyllis) vivipara ? Krohn, 1869 

Syllis (Typosyllis) sp. B of Uebelacker, 1984 

Genus Trypanosyllis Claparède, 1864 

Trypanosyllis coeliaca Claparède, 1868 

Trypanosyllis inglei Perkins, 1981 

Trypanosyllis parvidentata Perkins, 1981 

Trypanosyllis prampramensis Augener, 1918 

Trypanosyllis savagei Perkins, 1981 

Trypanosyllis vittigera Ehlers, 1887 

Syllidae, Eusyllinae undetermined genus 

“Ehlersia” ferrugina Langerhans, 1881 

Family Nereididae Lamarck, 1818 

Genus Ceratocephale Malmgren, 1867 

Ceratocephale oculata Banse, 1977 

Genus Ceratonereis Kinberg, 1866 



Subgenus Ceratonereis Kinberg, 1866 

Ceratonereis (Ceratonereis) longicirrata Perkins, 

1980 

Ceratonereis (Ceratonereis) mirabilis Kinberg, 1866 

Ceratonereis (Ceratonereis) singularis Treadwell, 

1929 

Subgenus Composetia Hartmann-Schröder, 1985 

Ceratonereis (Composetia) irritabilis (Webster, 

1879) 

Ceratonereis (Composetia) versipedata Ehlers, 1887 

Genus Gymnonereis Horst, 1919 

Gymnonereis crosslandi (Monro, 1933) 

Genus Laeonereis Hartman, 1945 

Laeonereis culveri (Webster, 1879) 

Genus Namalycastis Hartman, 1959 

Namalycastis abiuma (Müller in Grube, 1871) 

Genus Nereis Linnaeus, 1758 

Subgenus Neanthes Kinberg, 1866 

Nereis (Neanthes) acuminata Ehlers, 1868 

Nereis (Neanthes) micromma Harper, 1979 

Nereis (Neanthes) succinea Frey & Leuckart, 1847 

Subgenus Nereis Linnaeus, 1758 

Nereis (Nereis) allenae Pettibone, 1956 

Nereis (Nereis) falsa Quatrefages, 1866 

Nereis (Nereis) grayi Pettibone, 1956 

Nereis (Nereis) jacksoni Kinberg, 1866 

Nereis (Nereis) lamellosa Ehlers, 1868 

Nereis (Nereis) largoensis Treadwell, 1931 

Nereis (Nereis) panamensis Fauchald, 1977 

Nereis (Nereis) riisei Grube, 1857 

Genus Perinereis Kinberg, 1866 

Perinereis anderssoni Kinberg, 1866 

Perinereis floridana (Ehlers, 1868) 

Genus Platynereis Kinberg, 1866 

Platynereis dumerilii (Audouin & Milne Edwards, 

1833) 

Genus Rullierinereis Pettibone, 1971 

Rullierinereis mexicana (Treadwell, 1928) 

Rullierinereis sp. A of Taylor, 1984 

Genus Stenoninereis Wesenberg-Lund, 1958 

Stenoninereis martini Wesenberg-Lund, 1958 

Genus Websterinereis Pettibone, 1971 

Websterinereis tridentata (Webster, 1879) 

Family Nephtyidae Grube, 1850 

Genus Aglaophamus Kinberg, 1866 

Aglaophamus circinata (Verrill, 1874) 

Aglaophamus verrilli (McIntosh, 1885) 

Genus Inermonephtys Fauchald, 1968 

Inermonephtys inermis (Ehlers, 1887) 

Genus Micronephtys Friedrich, 1939 

Micronephtys minuta (Théel, 1879) 

Genus Nephtys Lamarck, 1818 

Nephtys bucera Ehlers, 1868 

Nephtys cryptomma Harper, 1986 

Nephtys incisa Malmgren, 1865 

Nephtys picta Ehlers, 1868 

Nephtys simoni Perkins, 1980 

Nephtys squamosa Ehlers, 1887 

Family Paralacydoniidae Pettibone, 1963 

Genus Paralacydonia Fauvel, 1913 

Paralacydonia paradoxa Fauvel, 1913 

Family Aphroditidae Lamarck, 1818 

Genus Aphrodita Linnaeus, 1758 

Aphrodita acuminata Ehlers, 1887 

Aphrodita objecta Ehlers, 1887 

Genus Aphrogenia Kinberg, 1856 

Aphrogenia sp. A of Gathof, 1984 

Genus Pontogenia Claparède, 1868 

Pontogenia chrysocoma (Baird, 1865) 

Pontogenia maggae Augener, 1906 

Pontogenia sericoma Ehlers, 1887 

Pontogenia sp. A of Gathof, 1984 

Family Polynoidae Kinberg, 1856 

Genus Halosydna Kinberg, 1856 

Halosydna leucohyba (Schmarda, 1861) 

Genus Harmothoe Kinberg, 1856 

Harmothoe aculeata Andrews, 1891 

Harmothoe crucis (Grube, 1856) 

Harmothoe imbricata (Linnaeus, 1767) 

Harmothoe sp. B of Weston, 1984 

Harmothoe sp. C of Weston, 1984 

Genus Hermenia Grube, 1856 

Hermenia verruculosa Grube, 1856 

Genus Lepidametria Webster, 1879 



Lepidametria commensalis Webster, 1879 

Genus Lepidasthenia Malmgren, 1867 

Lepidasthenia varia Treadwell, 1917 

Lepidasthenia sp. A of Weston, 1984 

Genus Lepidonopsis Pettibone, 1977 

Lepidonopsis humilis (Augener, 1922) 

Genus Lepidonotus Leach, 1816 

Lepidonotus inquilinus Treadwell, 1917 

Lepidonotus sublevis Verrill, 1873 

Lepidonotus variabilis Webster, 1879 

Genus Malmgreniella Hartman, 1967 

Malmgreniella galetaensis Pettibone, 1993 

Malmgreniella maccraryae Pettibone, 1993 

Malmgreniella pierceae Pettibone, 1993 

Malmgreniella puntotorensis Pettibone, 1993 

Malmgreniella taylori Pettibone, 1993 

Malmgreniella variegata (Treadwell, 1917) 

Genus Phyllohartmania Pettibone, 1961 

Phyllohartmania taylori Pettibone, 1961 

Genus Thormora Baird, 1865 

Thormora johnstoni (Kinberg, 1855) 

Thormora taeniata (Ehlers, 1887) 

Polynoidae gen. A of Weston, 1984 

Undetermined sp. of Weston, 1984 

Polynoidae gen. C of Weston, 1984 

Undetermined sp. of Weston, 1984 

Polynoidae undetermined genus 

Polynoid sp. A of Taylor, 1971 

Family Acoetidae Kinberg, 1858 

Genus Acoetes Audouin & Milne Edwards, 1832 

Acoetes pleei Audouin & Milne Edwards, 1832 

Genus Euarche Ehlers, 1887 

Euarche tubifex Ehlers, 1887 

Genus Eupolyodontes Buchanan, 1894 

Eupolyodontes batabanoensis Ibarzabal, 1988 

Genus Panthalis Kinberg, 1856 

Panthalis pustulata Treadwell, 1924 

Genus Polyodontes Renieri, 1828 

Polyodontes lupinus (Stimpson, 1856) 

Polyodontes oculeus (Treadwell, 1900) 

Family Eulepethidae Chamberlin, 1919 

Genus Grubeulepis Pettibone, 1969 

Grubeulepis augeneri Pettibone, 1969 

Grubeulepis mexicana (Berkeley & Berkeley, 1939) 

Grubeulepis westoni Pettibone, 1986 

Genus Lamelleulepethus Pettibone, 1986 

Lamelleulepethus biminiensis Pettibone, 1986 

Genus Mexieulepis Rioja, 1961 

Mexieulepis weberi (Horst, 1922) 

Family Sigalionidae Kinberg, 1856 

Genus Dayipsammolyce Pettibone, 1997 

Dayipsammolyce ctenidophora (Day, 1973) 

Genus Fimbriosthenelais Pettibone, 1971 

Fimbriosthenelais hobbsi Pettibone, 1971 

Fimbriosthenelais minor (Pruvot & Racovitza, 

1895) 

Fimbriosthenelais sp. A of Wolf, 1984 

Genus Pelogenia Schmarda, 1861 

Pelogenia anoculata (Hartman, 1939) 

Pelogenia fimbriata (Hartman, 1939) 

Pelogenia kinbergi (Hansen, 1882) 

Genus Sigalion Audouin & Milne Edwards, 1832 

Sigalion arenicola Verrill, 1880 

Sigalion lewisi Berkeley & Berkeley, 1939 

Sigalion sp. A of Wolf, 1984 

Sigalion sp. B of Wolf, 1984 

(as Thalenessa sp. A) 

Genus Sthenelais Kinberg, 1856 

Sthenelais boa (Johnston, 1839) 

Sthenelais limicola (Ehlers, 1864) 

Sthenelais sp. A of Wolf, 1984 

Genus Sthenolepis Willey, 1905 

Sthenolepis cf. grubei of Wolf, 1984 

Family Pholoidae Kinberg, 1858 

Genus Taylorpholoe Pettibone, 1992 

Taylorpholoe hirsuta (Rullier & Amoureux, 1979) 

Family Chrysopetalidae Ehlers, 1864 

Genus Acanthopale San Martín, 1986 



Acanthopale perkinsi San Martín, 1986 

Genus Bhawania Schmarda, 1861 

Bhawania goodei Webster, 1884 

Bhawania heteroseta (Hartman, 1945) 

Genus Chrysopetalum Ehlers, 1864 

Chrysopetalum elegans Bush, 1900 

Chrysopetalum eurypalea Perkins, 1985 

Chrysopetalum floridanum Perkins,1985 

Chrysopetalum heteropalea Perkins, 1985 

Genus Hyalopale Perkins, 1985 

Hyalopale bispinosa Perkins, 1985 

Genus Paleanotus Schmarda, 1861 

Paleanotus chrysolepis Schmarda, 1861 

Genus Treptopale Perkins, 1985 

Treptopale rudolphi Perkins, 1985 

Chrysopetalidae undetermined genus 

Undetermined sp. of Gathof, 1984 

(as Paleanotus sp. A) 

Family Pisionidae Southern, 1914 

Genus Pisione Grube, 1857 

Pisione hartmannschroederae Westheide, 1995 

Pisione remota (Southern, 1914) 

Pisione sp. A of Wolf, 1984 

Genus Pisionidens Aiyar & Alikunhi, 1943 

Pisionidens indicus (Aiyar & Alikunhi, 1940) 

Order Amphinomida 

Family Amphinomidae Lamarck, 1818 

Genus Amphinome Bruguière, 1789 

Amphinome rostrata (Pallas, 1776) 

Genus Chloeia Lamarck, 1818 

Chloeia viridis Schmarda, 1861 

Genus Eurythoe Kinberg, 1857 

Eurythoe complanata (Pallas, 1766) 

Eurythoe sp. B of Uebelacker, 1984 

Genus Hermodice Kinberg, 1867 

Hermodice carunculata (Pallas, 1766) 

Genus Hipponoa Audouin & Milne Edwards, 1830 

Hipponoa gaudchaudi Audouin & Milne Edwards, 

1830 

Genus Notopygos Grube, 1855 

Notopygos crinita Grube, 1855 

Genus Paramphinome Sars, 1869 

Paramphinome sp. B of Wolf, 1984 

Genus Pareurythoe Gustafson, 1930 

Pareurythoe americana Hartman, 1951 

Genus Pseudeurythoe Fauvel, 1932 

Pseudeurythoe ambigua (Monro, 1933) 

Pseudeurythoe fauchaldi (San Martín, 1986) 

Pseudeurythoe paucibranchiata Fauvel, 1932 

Family Euphrosinidae Williams, 1851 

Genus Euphrosine Lamarck, 1818 

Euphrosine triloba Ehlers, 1887 

Euphrosine cf. triloba of Gilbert, 1984 

Euphrosine sp. B of Gilbert, 1984 

Order Eunicida 

Family Onuphidae Kinberg, 1865 

Genus Americonuphis Fauchald, 1973 

Americonuphis magna (Andrews, 1891) 

Genus Diopatra Audouin & Milne Edwards, 1833 

Diopatra cuprea (Bosc, 1802) 

Diopatra neotridens Hartman, 1944 

Diopatra cf. papillata of Gathof, 1984 

Diopatra tridentata Hartman, 1944 

Genus Kinbergonuphis Fauchald, 1882 

Kinbergonuphis simoni (Santos, Day & Rice, 1981) 

Kinbergonuphis microcephala (Hartman, 1944) 

Kinbergonuphis sp. C of Gathof, 1984 

Genus Mooreonuphis Fauchald, 1982 

Mooreonuphis nebulosa (Moore, 1911) 

Mooreonuphis pallidula (Hartman, 1965) 

Genus Onuphis Audouin & Milne Edwards, 1833 

Onuphis eremita oculata Hartman, 1951 

Onuphis sp. A of Gathof, 1984 

Family Eunicidae Lamarck, 1818 

Genus Eunice Cuvier, 1817 

Eunice aphroditois (Pallas, 1788) 

Eunice articulata Ehlers, 1887 

Eunice binominata Quatrefages, 1866 

Eunice brevis (Ehlers, 1887) 

Eunice cariboea Grube, 1856 



Eunice conglomerans Ehlers, 1887 

Eunice culebra Treadwell, 1900 

Eunice denticulata Webster, 1884 

Eunice filamentosa Grube, 1856 

Eunice filamentosa of Treadwell, 1921 

(as Leodice filamentosa) 

Eunice floridana (Pourtalès, 1869) 

Eunice fucata Ehlers, 1887 

Eunice goodei Fauchald, 1992 

Eunice guanica (Treadwell, 1921) 

Eunice imogena (Monro, 1924) 

Eunice longisetis Webster, 1884 

Eunice macrobranchia Schmarda, 1861 

Eunice multicylindri Shisko, 1981 

Eunice mutilata Webster, 1884 

Eunice mutilatoides Augener, 1922 

Eunice rosaurae Monro, 1939 

Eunice rubra Grube, 1856 

Eunice spongicola (Treadwell, 1921) 

Eunice stigmatura (Verrill, 1900) 

Eunice tenuis (Treadwell, 1921) 

Eunice unifrons (Verrill, 1900) 

Eunice violaceomaculata Ehlers, 1887 

Eunice vittatopsis Fauchald, 1970 

Eunice websteri Fauchald, 1969 

Eunice weintraubi Lu & Fauchald, 1998 

Eunice wui Lu & Fauchald, 1998 

Genus Euniphysa Wesenberg-Lund, 1949 

Euniphysa sp. A of Gathof, 1984 

Genus Lysidice Lamarck, 1818 

Lysidice ninetta Audouin & Milne Edwards, 1833 

Lysidice notata Ehlers, 1887 

Lysidice sulcata Treadwell, 1900 

Lysidice tortugae Treadwell, 1921 

Genus Marphysa Quatrefages, 1866 

Marphysa acicularium Webster, 1884 

Marphysa cf. belli of Gathof, 1984 

Marphysa belli oculata Treadwell, 1921 

Marphysa nobilis Treadwell, 1917 

Marphysa regalis Verrill, 1900 

Marphysa sanguinea (Montagu, 1815) 

Marphysa viridis Treadwell, 1917 

Marphysa sp. A of Gathof, 1984 

Genus Nematonereis Schmarda, 1861 

Nematonereis hebes Verrill, 1900 

Nematonereis unicornis (Grube, 1840) 

Genus Palola Gray in Stair, 1847 

Palola siciliensis (Grube, 1840) 

Genus Paramarphysa Ehlers, 1887 

Paramarphysa longula Ehlers, 1887 

Family Lumbrineridae Schmarda, 1861 

Genus Lumbrinerides Orensanz, 1973 

Lumbrinerides acutus (Verrill, 1875) 

Lumbrinerides dayi Perkins, 1979 

Lumbrinerides jonesi Perkins, 1979 

Genus Lumbrineriopsis Orensanz, 1973 

Lumbrineriopsis paradoxa (Saint Joseph, 1888) 

Genus Lumbrineris Blainville, 1828 

Subgenus Abyssoninoe Orensanz, 1990 

Lumbrineris (Abyssoninoe ?) candida (Treadwell, 

1921) 

Subgenus Augeneria Monro, 1930 

Lumbrineris (Augeneria) albidentata Ehlers, 1908 

Lumbrineris (Augeneria ?) paucidentata Treadwell, 

1921 

Subgenus Lumbricalus Frame, 1992 

Lumbrineris (Lumbricalus) dayi (Frame, 1992) 

Subgenus Lumbrineris Blainville, 1828 

Lumbrineris (Lumbrineris) coccinea (Renier, 1804) 

Lumbrineris (Lumbrineris) floridana (Ehlers, 1887) 

Lumbrineris (Lumbrineris) floridana polygnatha 

Monro, 1933 

Lumbrineris (Lumbrineris) inflata Moore, 1911 

Lumbrineris (Lumbrineris) latreilli Auct. 

Lumbrineris (Lumbrineris) sp. C of Uebelacker, 

1984 

Lumbrineris (Lumbrineris) sp. D of Uebelacker, 

1984 

Lumbrineris (Lumbrineris) sp. E of Uebelacker, 

1984 

Subgenus Paraninoe Levinstein, 1977 

Lumbrineris (Paraninoe) brevipes McIntosh, 1903 

Subgenus Scoletoma Blainville, 1828 

Lumbrineris (Scoletoma ?) branchiata Treadwell, 

1921 

Lumbrineris (Scoletoma) ernesti Perkins, 1979 

Lumbrineris (Scoletoma) heteropoda (Marenzeller, 

1879) 

Lumbrineris (Scoletoma) maculata (Treadwell, 1900) 

Lumbrineris (Scoletoma) parvipedata (Treadwell, 

1900) 

Lumbrineris (Scoletoma) tenuis (Verrill, 1873) 

Lumbrineris (Scoletoma) testudinum (Augener, 

1922) 

Lumbrineris (Scoletoma) tetraura (Schmarda, 1861) 

(= Lumbrineris impatiens (Claparède, 1868)) 

Lumbrineris (Scoletoma) verrilli Perkins, 1979 



Genus Lysarete Kinberg, 1865 

Lysarete brasiliensis Kinberg, 1865 

Genus Ninoe Kinberg, 1865 

Ninoe nigripes gracilis Hartman, 1951 

Family Oenonidae Kinberg, 1865 

Genus Arabella Grube, 1850 

Arabella iricolor (Montagu, 1804) 

Arabella maculosa Verrill, 1900 

Arabella multidentata (Ehlers, 1887) 

Arabella mutans Auct. 

Genus Drilonereis Claparède, 1870 

Drilonereis cylindrica Hartman, 1951 

Drilonereis debilis (Ehlers, 1887) 

Drilonereis longa Webster, 1879 

Drilonereis magna Webster & Benedict, 1887 

Drilonereis spatula (Treadwell, 1911) 

Drilonereis sp. E of Uebelacker, 1984 

Genus Oenone Lamarck, 1818 

Oenone fulgida (Lamarck, 1818) 

Family Dorvilleidae Chamberlin, 1919 

Genus Diaphorosoma Wolf, 1986 

Diaphorosoma magnavena Wolf, 1986 

Genus Dorvillea Parfitt, 1866 

Subgenus Dorvillea Parfitt, 1866 

Dorvillea (Dorvillea) clavata Wolf, 1986 

Dorvillea (Dorvillea) largidentis Wolf, 1986 

Dorvillea (Dorvillea) rubra (Grube, 1856) 

Dorvillea (Dorvillea) sociabilis (Webster, 1879) 

Dorvillea (Dorvillea) vittata (Grube, 1856) 

Subgenus Schistomeringos Jumars, 1974 

Dorvillea (Schistomeringos) pectinata (Perkins, 

1979) 

Dorvillea (Schistomeringos) perkinsi Wolf, 1986 

Dorvillea (Schistomeringos) rudolphi (delle Chiaje, 

1828) 

Genus Gymnodorvillea Wainright & Perkins, 1982 

Gymnodorvillea floridana Wainright & Perkins, 

1982 

Genus Meiodorvillea Jumars, 1974 

Meiodorvillea sp. A of Wolf, 1984 

Meiodorvillea sp. B of Wolf, 1984 

Genus Ougia Wolf, 1986 

Ougia tenuidentis Wolf, 1986 

Genus Pettiboneia Orensanz, 1973 

Pettiboneia blakei Wolf, 1987 

Pettiboneia duofurca Wolf, 1987 

Genus Protodorvillea Pettibone, 1961 

Protodorvillea bifida Perkins, 1979 

Protodorvillea kefersteini (McIntosh, 1869) 

Genus Westheidia Wolf, 1986 

Westheidia minutimala Wolf, 1986 

Order Orbiniida 

Family Orbiniidae Hartman, 1942 

Genus Leitoscoloplos Day, 1977 

Leitoscoloplos foliosus (Hartman, 1951) 

Leitoscoloplos fragilis (Verrill, 1873) 

Leitoscoloplos robustus (Verrill, 1873) 

Genus Naineris Blainville, 1828 

Naineris bicornis Hartman, 1951 

Naineris grubei (Gravier, 1909) 

Naineris laevigata (Grube, 1855) 

Naineris ramosa (Schmarda, 1861) 

Naineris setosa (Verrill, 1900) 

Naineris sp. of Taylor, 1971 

Genus Orbinia Quatrefages, 1866 

Orbinia americana Day, 1973 

Orbinia riseri (Pettibone, 1957) 

Genus Pararicia Solís-Weiss & Fauchald, 1989 

Pararicia belizensis Solís-Weiss & Fauchald, 1989 

Genus Pettibonella Solís-Weiss & Fauchald, 1989 

Pettibonella multiuncinata Solís-Weiss & 

Fauchald, 1989 

Genus Phylo Kinberg, 1866 

Phylo ornatus (Verrill, 1873) 

Genus Proscoloplos Day, 1954 

Proscoloplos sp. A of Taylor, 1984 

Genus Scoloplos Blainville, 1828 

Scoloplos acmeceps Chamberlin, 1919 

Scoloplos capensis (Day, 1961) 

Scoloplos rubra (Webster, 1879) 

Scoloplos texana Maciolek & Holland, 1978 

Scoloplos sp. B of Taylor, 1984 

Order Spionida 

Family Spionidae Grube, 1850 



Genus Aonides Claparède, 1864 

Aonides mayaguezensis Auct. 

Aonides paucibranchiata Southern, 1914 

Genus Apoprionospio Foster, 1969 

Apoprionospio dayi Foster, 1969 

Apoprionospio pygmaea (Hartman, 1961) 

Genus Boccardiella Blake & Kudenov, 1978 

Boccardiella hamata (Webster, 1879) 

Boccardiella ligerica (Ferroniere, 1898) 

Genus Carazziella Blake & Kudenov, 1978 

Carazziella hobsonae Blake, 1979 

Genus Dipolydora Verrill, 1879 

Dipolydora armata (Langerhans, 1880) 

Dipolydora barbilla (Blake, 1981) 

Dipolydora bidentata (Zachs, 1933) 

Dipolydora caulleryi (Mesnil, 1897) 

Dipolydora commensalis (Andrews, 1891) 

Dipolydora giardi (Mesnil, 1896) 

Dipolydora socialis (Schmarda, 1861) 

Dipolydora tetrabranchia (Hartman, 1945) 

Genus Dispio Hartman, 1951 

Dispio uncinata Hartman, 1951 

Genus Laonice Malmgren, 1867 

Laonice cirrata (Sars, 1851) 

Genus Malacoceros Quatrefages, 1843 

Malacoceros indicus (Fauvel, 1928) 

Malacoceros vanderhorsti (Augener, 1927) 

Genus Microspio Mesnil, 1896 

Microspio pigmentata (Reish, 1959) 

Genus Paraprionospio Caullery, 1914 

Paraprionospio pinnata (Ehlers, 1901) 

Genus Polydora Bosc, 1802 

Polydora aggregata Blake, 1969 

Polydora colonia Moore, 1907 

Polydora cornuta Bosc, 1802 

Polydora websteri Hartman, 1943 

Genus Prionospio Malmgren, 1867 

Subgenus Minuspio Foster, 1971 

Prionospio (Minuspio) cirrifera Wirén, 1883 

Prionospio (Minuspio) multibranchiata Berkeley, 

1927 

Prionospio (Minuspio) perkinsi Maciolek, 1985 

Subgenus Prionospio Malmgren, 1867 

Prionospio (Prionospio) cristata Foster, 1971 

Prionospio (Prionospio) fallax Söderström, 1920 

Prionospio (Prionospio) heterobranchia Moore, 1907 

Prionospio (Prionospio) steenstrupi Malmgren, 1867 

Genus Pseudopolydora Czernaivsky, 1881 

Pseudopolydora antennata (Claparède, 1869) 

Genus Rhynchospio Hartman, 1936 

Rhynchospio glutaea (Ehlers, 1897) 

Genus Scolelepis Blainville, 1828 

Subgenus Parascolelepis Blake, 1987 

Scolelepis (Parascolelepis) texana Foster, 1971 

Subgenus Scolelepis Blainville, 1828 

Scolelepis (Scolelepis) heteropoda (Webster, 1879) 

Scolelepis (Scolelepis) squamata (Abildgaard, 1806) 

Genus Spio Fabricius, 1785 

Spio pettiboneae Foster, 1971 

Spio setosa Verrill, 1873 

Genus Spiophanes Grube, 1860 

Spiophanes bombyx (Claparède, 1870) 

Spiophanes kroyeri Grube, 1860 

Spiophanes duplex (Chamberlin, 1919) 

Spiophanes wigleyi Pettibone, 1962 

Genus Streblospio Webster, 1879 

Streblospio benedicti Webster, 1879 

Family Poecilochaetidae Hannerz, 1956 

Genus Poecilochaetus Ehlers, 1875 

Poecilochaetus johnsoni Hartman, 1939 

Family Longosomatidae Hartman, 1944 

Genus Heterospio Ehlers, 1875 

Heterospio sp. A of Uebelacker, 1984 

Order Chaetopterida 

Family Chaetopteridae 

Audouin & Milne Edwards, 1833 

Genus Chaetopterus Cuvier, 1827 

Chaetopterus pergamentaceus Cuvier, 1827 

Genus Mesochaetopterus Potts, 1914 

Mesochaetopterus capensis (McIntosh, 1885) 

Mesochaetopterus sagitarius (Claparède, 1870) 

Mesochaetopterus taylori Potts, 1914 

Mesochaetopterus sp. A of Taylor, 1971 

Mesochaetopterus sp. C of Taylor, 1971 



Genus Spiochaetopterus Sars, 1863 

Spiochaetopterus costarum oculatus Webster, 1879 

Order Magelonida 

Family Magelonidae 

Cunningham & Ramage, 1888 

Genus Magelona Müller, 1858 

Magelona cincta Ehlers, 1908 

Magelona cornuta Jones, 1971 

Magelona pettiboneae Jones, 1963 

Magelona phyllisae Jones, 1963 

Magelona polydentata Jones, 1963 

Magelona riojai Jones, 1963 

Magelona rosea Moore, 1907 

Magelona sp. A of Uebelacker & Jones, 1984 

Magelona sp. B of Uebelacker & Jones, 1984 

Magelona sp. C of Uebelacker & Jones, 1984 

Magelona sp. F of Uebelacker & Jones, 1984 

Magelona sp. G of Uebelacker & Jones, 1984 

Magelona sp. H of Uebelacker & Jones, 1984 

Magelona sp. I of Uebelacker & Jones, 1984 

Magelona sp. J of Uebelacker & Jones, 1984 

Magelona sp. K of Uebelacker & Jones, 1984 

Magelona sp. L of Uebelacker & Jones, 1984 

Order Cirratulida 

Family Paraonidae Cerruti, 1909 

Genus Aricidea Webster, 1879 

Aricidea bryani Gaston & McLelland, 1996 

Aricidea catherinae Laubier, 1967 

Aricidea cerrutii Laubier, 1966 

Aricidea cf. finitima of Gaston, 1984 

Aricidea fragilis Webster, 1879 

Aricidea fragilis of Gaston, 1984 

Aricidea jeffreysi of Renaud, 1956 

Aricidea fauveli ? Hartman, 1957 

(as Aricidea lopezi of Gaston, 1984) 

Aricidea philbinae Brown, 1976 

Aricidea cf. pseudoarticulata of Gaston,1984 

Aricidea quadrilobata Webster & Benedict, 1887 

Aricidea suecica Eliason, 1920 

Aricidea taylori Pettibone, 1965 

Aricidea trilobata Imajima, 1973 

Aricidea wassi Pettibone, 1965 

Aricidea sp. A of Gaston, 1984 

(as Aricidea (Acmira) sp. A) 

Aricidea sp. B of Gaston, 1984 

(as Aricidea (Allia) sp. A) 

Aricidea sp. C of Gaston, 1984 

(as Aricidea (Acmira) sp. C) 

Genus Cirrophorus Ehlers, 1908 

Cirrophorus americanus Strelzov, 1973 

Cirrophorus branchiatus Ehlers, 1908 

Cirrophorus cf. forticirratus of Gaston, 1984 

(Cirrophorus furcatus Auct.) 

Cirrophorus cf. lyra (Southern, 1914) 

Cirrophorus perdidoensis McLelland & Gaston, 

1994 

Cirrophorus perkinsi McLelland & Gaston, 1994 

Genus Levinsenia Mesnil, 1897 

Levinsenia gracilis (Tauber, 1879) 

Genus Paraonis Cerruti, 1909 

Paraonis fulgens (Levinsen, 1883) 

Paraonis pygoenigmatica Jones, 1968 

Family Questidae Hartman, 1966 

Genus Novaquesta Hobson, 1970 

Novaquesta trifurcata Hobson, 1970 

Genus Questa Hartman, 1966 

Questa caudicirra Hartman, 1966 

Questidae gen. A of Taylor & Gathof, 1984 

Undetermined sp. of Taylor & Gathof, 1984 

Family Cirratulidae Ryckholt, 1851 

Genus Aphelochaeta Blake, 1991 

Aphelochaeta marioni (Saint Joseph, 1894) 

Aphelochaeta sp. A of Taylor, 1971 

(as Tharyx sp. A) 

Aphelochaeta sp. B of Taylor, 1971 

(as Tharyx sp. B) 

Genus Caulleriella Chamberlin, 1919 

Caulleriella cf. alata of Wolf, 1984 

Caulleriella cf. zetlandica of Wolf, 1984 

Caulleriella sp. A of Wolf, 1984 

Caulleriella sp. B of Wolf, 1984 

Genus Chaetozone Malmgren, 1867 

Chaetozone gayheadia Hartman, 1965 

Chaetozone setosa Malmgren, 1867 

Chaetozone sp. A of Wolf, 1984 

Chaetozone sp. B of Wolf, 1984 

Chaetozone sp. D of Wolf, 1984 

Genus Cirratulus Lamarck, 1818 

Cirratulus sp. B of Taylor, 1971 

Cirratulus sp. C of Taylor, 1971 

Cirratulus sp. D of Taylor, 1971 



Genus Cirriformia Hartman, 1936 

Cirriformia filigera (delle Chiaje, 1828) 

Cirriformia grandis (Verrill, 1873) 

Cirriformia melanacantha (Grube, 1872) 

Cirriformia punctata (Grube, 1859) 

Cirriformia tentaculata (Montagu, 1808) 

Cirriformia tortugaenis (Augener, 1922) 

Cirriformia sp. A of Taylor, 1971 

Cirriformia sp. B of Taylor, 1971 

Cirriformia sp. C of Taylor, 1971 

(as Cirratulus sp. A) 

Cirriformia sp. B of Wolf, 1984 

Genus Dodecaceria Örsted, 1843 

Dodecaceria concharum Örsted, 1843 

Dodecaceria corallii (Leidy, 1855) 

Dodecaceria diceria Hartman, 1951 

Dodecaceria sp. A of Wolf, 1984 

Genus Monticellina Laubier, 1961 

Monticellina dorsobranchialis (Kirkegaard, 1959) 

Genus Tharyx Webster & Benedict, 1887 

Tharyx acutus Webster & Benedict, 1887 

(as Caulleriella sp. C of Wolf, 1984) 

Tharyx killariensis (Southern, 1914) 

Cirratulidae undetermined genus 

Undetermined sp. of Wolf, 1984 

Order Ctenodrilida 

Family Ctenodrilidae Kennel, 1882 

Genus Ctenodrilus Claparède, 1863 

Ctenodrilus serratus (Schmidt, 1857) 

Order Cossurida 

Family Cossuridae Day, 1963 

Genus Cossura Webster & Benedict, 1887 

Cossura delta Reish, 1958 

Order Flabelligerida 

Family Flabelligeridae Saint Joseph, 1894 

Genus Diplocirrus Haase, 1915 

Diplocirrus sp. A of Milligan, 1984 

Genus Pherusa Oken, 1807 

Pherusa dubia (Treadwell, 1929) 

Pherusa ehlersi (Day, 1973) 

Pherusa flabellata (Sars, 1871) 

Pherusa inflata (Treadwell, 1914) 

Genus Piromis Kinberg, 1866 

Piromis caribous (Grube, 1859) 

Piromis roberti (Hartman, 1951) 

Genus Therochaeta Chamberlin, 1919 

Therochaeta sp. A of Milligan, 1984 

Family Acrocirridae Banse, 1969 

Genus Acrocirrus Grube, 1872 

Acrocirrus frontifilis (Grube, 1860) 

Genus Macrochaeta Grube, 1850 

Macrochaeta cf. clavicornis of Wolf, 1984 

Family Fauveliopsidae Hartman, 1971 

Genus Fauveliopsis McIntosh, 1922 

Fauveliopsis sp. A of Milligan, 1984 

Order Opheliida 

Family Opheliidae Grube, 1850 

Genus Armandia Filippi, 1861 

Armandia agilis (Andrews, 1891) 

Armandia bermudiensis (Treadwell, 1936) 

Armandia maculata (Webster, 1884) 

Armandia polyophthalma Keukenthal, 1887 

Genus Ophelia Savigny, 1822 

Ophelia denticulata of Uebelacker, 1984 

Genus Ophelina Örsted, 1843 

Ophelina acuminata Örsted, 1843 

Ophelina cf. acuminata of Uebelacker, 1984 

Ophelina cilindricaudata (Hansen, 1878) 

Genus Polyophthalmus Quatrefages, 1850 

Polyophthalmus floridanus Augener, 1922 

Polyophthalmus pictus (Dujardin, 1839) 

Genus Travisia Johnston, 1840 

Travisia hobsonae Santos, 1977 

Family Scalibregmatidae Malmgren, 1867 

Genus Asclerocheilus Ashworth, 1901 

Asclerocheilus beringianus Uschakov, 1955 

Asclerocheilus mexicanus Kudenov, 1985 

Genus Hyboscolex Schmarda, 1861 

Hyboscolex longiseta Schmarda, 1861 

Hyboscolex quadricincta Kudenov, 1985 



Genus Neolipobranchus Hartman & Fauchald, 1971 

Neolipobranchius blakei Kudenov, 1985 

Genus Sclerobregma Hartman, 1965 

Sclerobregma stenocerum Bertelsen & Weston, 

1980 

Genus Sclerocheilus Grube, 1863 

Sclerocheilus unoculus Kudenov, 1985 

Order Sternaspida 

Family Sternaspidae Carus, 1863 

Genus Sternaspis Otto, 1821 

Sternaspis scutata (Renier, 1807) 

Order Capitellida 

Family Capitellidae Grube, 1862 

Genus Anotomastus Hartman, 1947 

Anotomastus cf. gordiodes (Moore, 1909) 

Genus Capitella Blainville, 1828 

Capitella aciculata (Hartman, 1959) 

Capitella capitata (Fabricius, 1780) complex 

Capitella capitata floridana Hartman, 1959 

Capitella caribaeorum Warren & George, 1986 

Capitella giardi (Mesnil, 1897) 

Capitella jonesi (Hartman, 1959) 

Genus Dasybranchus Grube, 1850 

Dasybranchus lumbricoides Grube, 1878 

Dasybranchus lunulatus Ehlers, 1887 

Genus Decamastus Hartman, 1963 

Decamastus gracilis ? Hartman, 1963 

Decamastus sp. A of Ewing, 1984 

Genus Heteromastus Eisig, 1887 

Heteromastus filiformis (Claparède, 1864) 

Genus Leiocapitella Hartman, 1947 

Leiocapitella glabra Hartman, 1947 

Leiocapitella sp. A of Ewing, 1984 

Leiocapitella sp. B of Ewing, 1984 

Genus Leiochrides Augener, 1914 

Leiochrides cf. pallidior (Chamberlin, 1918) 

Genus Mastobranchus Eisig, 1887 

Mastobranchus variabilis Ewing, 1984 

Genus Mediomastus Hartman, 1944 

Mediomastus ambiseta (Hartman, 1947) 

Mediomastus californiensis Hartman, 1944 

Genus Notomastus Sars, 1851 

Notomastus americanus Day, 1973 

Notomastus daueri Ewing, 1982 

Notomastus hemipodus Hartman, 1945 

Notomastus latericeus Sars, 1851 

Notomastus lineatus Claparède, 1870 

Notomastus lobatus Hartman, 1947 

Notomastus tenuis ? Moore, 1909 

Notomastus teres Hartman, 1965 

Notomastus sp. A of Ewing, 1984 

Genus Scyphoproctus Gravier, 1904 

Scyphoproctus platyproctus Jones, 1961 

Scyphoproctus cf. platyproctus of Ewing, 1984 

Family Maldanidae Malmgren, 1867 

Genus Axiothella Verrill, 1900 

Axiothella mucosa (Andrews, 1891) 

Axiothella sp. A of Wolf, 1984 

Genus Boguea Hartman, 1945 

Boguea enigmatica Hartman, 1945 

Boguea sp. A of Wolf, 1984 

Genus Boguella Hartman & Fauchald, 1971 

Boguella sp. A of Wolf, 1984 

Genus Clymenella Verrill, 1873 

Clymenella torquata (Leidy, 1855) 

Genus Euclymene Verrill, 1900 

Euclymene corallicola (Treadwell, 1929) 

Euclymene coronata Verrill, 1900 

Euclymene sp. A of Wolf, 1984 

Euclymene sp. B of Wolf, 1984 

Genus Macroclymene Verrill, 1900 

Macroclymene zonalis (Verrill, 1874) 

Genus Maldane Grube, 1860 

Maldane sarsi Malmgren, 1865 

Genus Petaloproctus Quatrefages, 1866 

Petaloproctus socialis Andrews, 1891 

Genus Sabaco Kinberg, 1867 

Sabaco americanus (Hartman, 1945) 

Sabaco carolinae (Day, 1973) 

Family Arenicolidae 

Audouin & Milne Edwards, 1833 



Genus Arenicola Lamarck, 1801 

Arenicola cristata Stimpson, 1856 

Genus Branchiomaldane Langerhans, 1881 

Branchiomaldane vincinti Langerhans, 1881 

Order Oweniida 

Family Oweniidae Rioja, 1917 

Genus Galathowenia Kirkegaard, 1959 

Galathowenia oculata (Zachs, 1923) 

Galathowenia sp. A of Milligan, 1984 

(as Myriochele sp. A) 

Genus Owenia delle Chiaje, 1841 

Owenia fusiformis delle Chiaje, 1841 

Order Terebellida 

Family Pectinariidae Quatrefages, 1866 

Genus Amphictene Savigny, 1822 

Amphictene sp. A of Wolf, 1984 

Genus Pectinaria Lamarck, 1818 

Pectinaria gouldii (Verrill, 1873) 

Pectinaria meredithi Long, 1973 

Pectinaria regalis Verrill, 1901 

Family Sabellariidae Johnston, 1965 

Genus Lygdamis Kinberg, 1867 

Lygdamis indicus of Uebelacker, 1984 

Lygdamis rayrobertsi Kirtley, 1994 

Genus Phalacrostemma Marenzeller, 1895 

Phalacrostemma perkinsi Kirtley, 1994 

Genus Phragmatopoma Mörch, 1863 

Phragmatopoma caudata Mörch, 1863 

Genus Sabellaria Lamarck, 1818 

Sabellaria floridensis Hartman, 1944 

Sabellaria vulgaris Verrill, 1873 

Sabellaria sp. A of Uebelacker, 1984 

Family Ampharetidae Malmgren, 1866 

Genus Ampharete Malmgren, 1866 

Ampharete acutifrons Grube, 1860 

Ampharete americana Day, 1973 

Ampharete parvidentata Day, 1973 

Ampharete sp. A of Uebelacker, 1984 

Ampharete sp. B of Uebelacker, 1984 

Genus Amphicteis Grube, 1850 

Amphicteis gunneri (Sars, 1835) 

Amphicteis scaphobranchia Moore, 1906 

Genus Hobsonia Banse, 1979 

Hobsonia florida (Hartman, 1951) 

Genus Isolda Müller, 1858 

Isolda pulchella Müller, 1858 

Genus Lysippe Malmgren, 1866 

Lysippe cf. annenectens of Uebelacker, 1984 

Genus Melinna Malmgren, 1867 

Melinna cristata (Sars, 1851) 

Melinna maculata Webster, 1879 

Genus Sabellides Milne Edwards in Malmgren, 1866 

Sabellides oculatus Webster, 1879 

Genus Samythella Verrill, 1900 

Samythella eliasoni Day, 1973 

Family Trichobranchidae Malmgren, 1866 

Genus Terebellides Sars, 1835 

Terebellides parvus Solís-Weiss, Fauchald, & 

Blankensteyn, 1991 

Terebellides stroemi Sars, 1835 

Genus Trichobranchus Malmgren, 1866 

Trichobranchus glacialis Malmgren, 1866 

Family Terebellidae Grube, 1850 

Genus Amaeana Hartman, 1959 

Amaeana trilobata (Sars, 1863) 

Genus Amphitrite Müller, 1771 

Amphitrite ornata (Leidy, 1855) 

Genus Amphitritides Augener, 1922 

Amphitritides bruneocomatus (Ehlers, 1887) 

Genus Enoplobranchus Verrill in Webster, 1879 

Enoplobranchus sanguineus (Verrill, 1873) 

Genus Eupolymnia Verrill, 1900 

Eupolymnia crassicornis (Schmarda, 1861) 

Eupolymnia nebulosa (Montagu, 1818) 

Eupolymnia sp. A of Kritzler, 1984 

Genus Hauchiella Levinsen, 1893 

Hauchiella sp. A of Kritzler, 1984 



Genus Lanice Malmgren, 1866 

Lanice conchilega (Pallas, 1766) 

Genus Lanicides Hessle, 1917 

Lanicides taboguillae (Chamberlin, 1919) 

Genus Loimia Malmgren, 1866 

Loimia medusa (Savigny, 1822) 

Loimia viridis Moore, 1903 

Genus Lysilla Malmgren, 1866 

Lysilla alba Webster, 1879 

Lysilla sp. A of Kritzler, 1984 

Lysilla sp. B of Kritzler, 1984 

Genus Neoamphitrite Hessle, 1917 

Neoamphitrite edwardsi (Quatrefages, 1866) 

Genus Neoleprea Hessle, 1917 

Neoleprea sp. A of Kritzler, 1984 

Neoleprea sp. B of Kritzler, 1984 

Genus Nicolea Malmgren, 1866 

Nicolea cetrata (Ehlers, 1887) 

Genus Pista Malmgren, 1866 

Pista cristata (Müller, 1776) 

Pista palmata (Verrill, 1873) 

Pista quadrilobata (Augener, 1918) 

Genus Polycirrus Grube, 1850 

Polycirrus albicans (Malmgren, 1866) 

Polycirrus carolinensis Day, 1973 

Polycirrus cf. denticulatus of Kritzler, 1984 

Polycirrus cf. haematodes of Kritzler, 1984 

Polycirrus plumosus (Wollenbaek, 1912) 

Polycirrus sp. A of Kritzler, 1984 

Polycirrus sp. B of Kritzler, 1984 

Genus Scionella Moore, 1903 

Scionella papillosa Tourtellotte & Kritzler, 1988 

Genus Scionides Chamberlin, 1919 

Scionides reticulatus (Ehlers, 1887) 

Genus Streblosoma Sars, 1872 

Streblosoma hartmanae Kritzler, 1971 

Streblosoma verrilli Treadwell, 1911 

Genus Terebella Linnaeus, 1767 

Terebella verrilli Holthe, 1986 

(Terebella rubra (Verrill, 1873)) 

Genus Thelepus Leukart, 1849 

Thelepus setosus (Quatrefages, 1866) 

Terebellidae gen. B of Kritzler, 1984 

Undetermined sp. of Kritzler, 1984 

Order Sabellida 

Family Sabellidae Latreille, 1825 

Genus Amphiglena Claparède, 1864 

Amphiglena mediterranea (Leidig, 1851) 

Genus Anamobaea Krøyer, 1865 

Anamobaea orstedii Krøyer, 1856 

Genus Augeneriella Banse, 1957 

Augeneriella hummelincki Banse, 1957 

Genus Bispira Krøyer, 1856 

Bispira melanostigma (Schmarda, 1861) 

Genus Branchiomma Kölliker, 1859 

Branchiomma arenosum (Treadwell, 1924) 

Branchiomma bairdi (McIntosh, 1885) 

Branchiomma conspersum (Ehlers, 1887) 

Branchiomma nigromaculatum (Baird, 1865) 

Genus Chone Krøyer, 1856 

Chone americana Day, 1973 

Chone cf. americana of Uebelacker, 1984 

Chone sp. A of Uebelacker, 1984 

Genus Demonax Kinberg, 1867 

Demonax lacunosus Perkins, 1984 

Demonax microphthalmus (Verrill, 1873) 

Genus Euchone Malmgren, 1866 

Euchone cf. incolor of Uebelacker, 1984 

Genus Fabricinuda Fitzhugh, 1990 

Fabricinuda pseudocollaris Fitzhugh, 1990 

Fabricinuda trilobata (Fitzhugh, 1983) 

Genus Jasmineira Langerhans, 1880 

Jasmineira caudata Langerhans, 1880 

Jasmineira pacifica Annenkova, 1937 

Jasmineira cf. pacifica of Uebelacker, 1984 

Genus Megalomma Johansson, 1927 

Megalomma bioculatum (Ehlers, 1887) 

Megalomma heterops Perkins, 1984 

Megalomma lobiferum (Ehlers, 1887) 

Megalomma pigmentum Reish, 1963 

Genus Notaulax Tauber, 1879 



Notaulax circumspiciens (Ehlers, 1887) 

Notaulax midoculi (Hoagland, 1919) 

Notaulax nudicollis (Krøyer, 1856) 

Notaulax occidentalis (Baird, 1865) 

Genus Novafabricia Fitzhugh, 1990 

Novafabricia infratorquata (Fitzhugh, 1983) 

Genus Oriopsis Caullery & Mesnil, 1896 

Oriopsis anneae Rouse, 1994 

Genus Perkinsiana Knight-Jones, 1983 

Perkinsiana floridana (Augener, 1922) 

Perkinsiana fonticula (Hoagland, 1919) 

Genus Potamethus Chamberlin, 1919 

Potamethus sp. A of Uebelacker, 1984 

Genus Pseudobranchiomma Jones, 1962 

Pseudobranchiomma emersoni Jones, 1962 

Genus Pseudofabriciola Fitzhugh, 1990 

Pseudofabriciola longa Fitzhugh, 1990 

Pseudofabriciola sofla Fitzhugh, 1996 

Genus Pseudopotamilla Bush, 1905 

Pseudopotamilla cf. reniformis of Uebelacker, 1984 

Pseudopotamilla tortuosa (Webster, 1879) 

Genus Sabellastarte Krøyer, 1856 

Sabellastarte magnifica (Shaw, 1800) 

Fabriciinae gen. A of Fitzhugh, 1989 

Undetermined sp. of Fitzhugh, 1989 

Family Serpulidae Rafinesque, 1815 

Genus Crucigera Benedict, 1887 

Crucigera websteri Benedict, 1887 

Genus Ficopomatus Southern, 1921 

Ficopomatus enigmaticus (Fauvel, 1923) 

Ficopomatus miamiensis (Treadwell, 1934) 

Genus Filograna Berkeley, 1835 

Filograna huxleyi Ehlers, 1887 

Genus Filogranula Langerhans, 1884 

Filogranula calyculata (Costa, 1861) 

Genus Hydroides Gunnerus, 1768 

Hydroides bispinosus Bush, 1910 

Hydroides dianthus (Verrill, 1873) 

Hydroides dirampha Mörch, 1863 

Hydroides elegans (Haswell, 1883) 

Hydroides floridanus Bush, 1910 

Hydroides gairacensis Augener, 1933 

Hydroides microtis Mörch,, 1863 

Hydroides mongeslopezi Rioja, 1958 

Hydroides cf. mucronatus Rioja, 1958 

Hydroides parvus (Treadwell, 1900) 

Hydroides protulicola Benedict, 1887 

Hydroides sanctaecrucis (Mörch, 1863) 

Hydroides spongicola Benedict, 1887 

Genus Josephella Caullery & Mesnil, 1896 

Josephella marenzelleri Caullery & Mesnil, 1896 

Genus Placostegus Philippi, 1844 

Placostegus incomptus Ehlers, 1887 

Genus Pomatoceros Philippi, 1844 

Pomatoceros americanus Day, 1973 

Pomatoceros minutus Rioja, 1941 

Genus Pomatostegus Schmarda, 1861 

Pomatostegus stellatus (Abildgaard, 1789) 

Genus Protula Risso, 1826 

Protula balbaoensis Monro, 1933 

Protula setosa (Bush, 1910) 

Genus Pseudovermilia Bush, 1907 

Pseudovermilia fuscostriata ten Hove, 1975 

Pseudovermilia multispinosa (Monro, 1933) 

Pseudovermilia occidentalis (McIntosh, 1885) 

Genus Rhodopsis Bush, 1905 

Rhodopsis pusilla Bush, 1905 

Genus Sclerostyla Mörch, 1863 

Sclerostyla ctenactis (Mörch, 1863) 

Genus Serpula Linnaeus, 1758 

Serpula sp. A of ten Hove & Wolf, 1984 

Genus Spiraserpula Regenhardt, 1961 

Spiraserpula caribensis Pillai & ten Hove, 1994 

Spiraserpula ypsilon Pillai & ten Hove, 1994 

Genus Spirobranchus Blainville, 1818 

Spirobranchus giganteus giganteus (Pallas, 1766) 

Genus Vermiliopsis Saint Joseph, 1894 

Vermiliopsis annulata (Schmarda, 1861) 

Family Spirorbidae Pillai, 1970 

(Included in Serpulidae by many authors) 



Genus Janua Saint Joseph, 1984 

Subgenus Dexiospira Caullery & Mesnil, 1897 

Janua (Dexiospira) corrugata (Montagu, 1803) 

Janua (Dexiospira) steueri (Sterzinger, 1909) 

Subgenus Leodora Saint Joseph, 1894 

Janua (Leodora) knightjonesi DeSilva, 1965 

Genus Pileolaria Claparède, 1870 

Pileolaria rosepigmentata (Uchida, 1971) 

Order Nerillida 

Family Nerillidae Levinsen, 1883 

Genus Nerilla Schmidt, 1848 

AIYAR, R. G., and K. H. ALIKUNHI. 1940. On a new pisionid 

from the sandy beach, Madras. Record of the Indian 

Museum 42(1): 89–107. 

AIYAR, R. G., and K. H. ALIKUNHI. 1943. Change of 

the generic name Pisionella Aiyar and Alikunhi, 1940, 

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Fauchald, 1971. Pp. 16.1–16.9 in J. M. Uebelacker and P. 


of the northern Gulf of Mexico,Vol. II. Final Report to 



WOLF, P. S. 1984e. Family Cirratulidae Carus, 1863. 






WOLF, P. S. 1984f. Family Dorvilleidae Chamberlin, 






WOLF, P. S. 1984g. Family Fauveliopsidae Hartman, 






WOLF, P. S. 1984h. Family Maldanidae Malmgren, 1867. 






WOLF, P. S. 1984i. Family Pectinariidae Quatrefages, 






WOLF, P. S. 1984j. Family Pilargidae Saint Joseph, 1899. 



Gulf of Mexico, Vol. IV. Final Report to the Minerals 



WOLF, P. S. 1984k. Family Pisionidae Southern, 1914. 



Gulf of Mexico, Vol. III. Final Report to the Minerals 



WOLF, P. S. 1984l. Family Polyodontidae Buchanan, 






WOLF, P. S. 1984m. Family Saccocirridae Czerniavsky, 






WOLF, P. S. 1984n. Family Sigalionidae Malmgren, 






WOLF, P. S. 1986a. Four new genera of Dorvilleidae (Annelida: 

Polychaeta) from the Gulf of Mexico. Proceedings 


616–626. 



WOLF, P. S. 1986b. A new genus and species of interstitial 

Sigalionidae and a report on the presence of 

venom glands in some scale-worm families (Annelida: 

Polychaeta). Proceedings of the Biological Society of 

Washington 99(1): 79–83. 

WOLF, P. S. 1986c. Three new species of Dorvilleidae 

(Annelida: Polychaeta) from Puerto Rico and Florida 

and a new genus for dorvilleids from Scandinavia and 

North America. Proceedings of the Biological Society 


WOLF, P. S. 1986d. Three new species of Pilargidae 

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Biological Society of Washington 99(3): 464–471. 

WOLF, P. S. 1987a. Aberrantidae: A new family of Polychaeta 

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WOLF, P. S. 1987b.Two new species of Pettiboneia (Polychaeta: 

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Methods 

Checklist of Shallow-Water Marine Malacostracan 

Crustacea of Florida 

David K. Camp 

The list of benthic malacostracan Crustacea includes 

most of the described species known from Florida’s estuaries 

and nearshore coastal zone to depths of approximately 

37 meters, with exceptions noted below. 

Most of the list was compiled from previously published 

compendia, with taxonomic updates and additions 

taken from many widely scattered papers cited 

in the References section. Several species were added 

from lists of museum holdings, including those of the 

Florida Marine Research Institute Marine Specimen 

Collection. 

The list of Stomatopoda was taken principally from 

Manning (1969), Camp (1971), Manning and Camp 

(1981), and Camp and Manning (1982, 1986). Farrell 

(1979) produced an identification guide to Florida’s 

Mysidacea, and Stuck et al. (1979) published a key to 

mysidacean species of the northern Gulf of Mexico. 

The list of Amphipoda was begun several years ago 

by the late Douglas H. Farrell, who gave his list to me. 

A few amphipods originally on the list have not appeared 

in publications about Florida’s fauna, but were 

included by Dr. Farrell based on his observations of 

specimens from Florida. Dr. Farrell’s list was augmented 

with species from computer records of the 

cataloged holdings of the National Museum of Natural 

History, Smithsonian Institution; from lists of cataloged 

holdings of the Harbor Branch Oceanographic 

Institution Museum; from a list of species identified 

from shallow-water sites off Florida by Richard Heard 

and Sara LeCroy; and from published works cited 

below. Recently published compendia of Amphipoda 

include works of McCain (1968), Bousfield (1973), Myers 

(1981), Barnard and Barnard (1983), Barnard and Karaman 

(1991), Ortiz (1991),Thomas (1993), LeCroy (1995), 

Nelson (1995), and Lowry and Stoddart (1997). Many 

species described in several smaller papers by J. L. 

Barnard, James D.Thomas, and others were added. 

The list of Isopoda was derived mainly from 

Markham (1978, 1985), Kensley (1980), Menzies and 

Kruczynski (1983), Schultz and Johnson (1984), Kensley 

and Schotte (1985), and Kensley et al. (1995). 

The list of pelagic Euphausiacea was taken from 

Mikkelsen (1987). Although neither strictly benthic 

nor shallow-water organisms, euphausiaceans that 

come to the surface or into shallow water at night were 

included in this list because they may be captured by 

benthic sampling gear near shore on the east coast of 

Florida. 

The list of shallow-water Decapoda was extracted 

from the larger list of Florida’s decapods published by 

Abele and Kim (1986). A few species they inadvertently 

omitted plus additional, recently described 

species were added, and the taxonomic arrangement 

was updated. 

Lettered species (e.g.,“Genus name”sp. A) were included 

if they met two criteria: 1) the lettered designation 

was published in a peer-reviewed journal, and 

2) I was confident that voucher material exists and is 

retrievable. A few deeper-water species were included 

even though there is no published record of their occurrence 

from depths as shallow as 37 meters. These 

relatively rare species may be found in shallower waters 

in the future. 

Several taxa are not included in the list for various 

reasons.The following orders of marine malacostracan 

Crustacea are not treated here: Lophogastrida (cosmopolitan, 

oceanic, pelagic swimmers; about 40 species 

worldwide); Amphionidacea (monotypic: Amphionides 

reynaudii (H. Milne Edwards, 1833), oceanic, pelagic); 

and Mictacea (in the western Atlantic, known only 

from marine caves in Bermuda and the Guyana Basin) 

(Brusca and Brusca, 1990).The hyperiidean amphipods 

were not included because they are planktonic. I have 

not included the whale lice (Amphipoda: Cyamidae), 

although whales sometimes become stranded on 

Florida’s coasts. Finally, several shallow-water, benthic 

species that live near Florida and could be expected 

to occur here are not included in this list. Islandic 

species recently described from Cuba and the Bahama 

Islands may one day be found in Florida, especially in 

the Florida Keys. Shallow-continental-shelf species 

from nearby states, such as the mysidacean Pseudomma 

heardi Stuck, 1981, known from off Mississippi, or the 

pinnotherid crab Gemmotheres chamae (Roberts, 1975), 

known from off North Carolina, may also be found here 

in the future but are not listed. 

The classification of taxa follows that used in the 

most recently published works, with exceptions.Within 

the Amphipoda, a mixture of proposed classifications 

is used, and families are arranged alphabetically within 

suborders, as was suggested by the reviewers, because 

amphipod taxonomists have not reached a consensus 

about the higher classification of that group. 


Malacostraca—D. K. Camp Macroinvertebrate Checklists Camp et al. 1998 


The late Douglas H. Farrell, formerly of the Florida 

Department of Environmental Regulation,Tampa, gave 

me a copy of a draft list of Florida amphipods that he 

had prepared. He and I intended to complete it and 

make it available to Florida benthologists. Doug should 

be a co-author of the present list, and I dedicate this 

effort to his memory.The late J. L. Barnard, formerly of 

the Smithsonian Institution, Washington, D.C., provided 

me with a list of the cataloged holdings of amphipods 

from the vicinity of Florida in the U.S. National 

Museum of Natural History as well as with a computer 

file of his geographic index to worldwide Amphipoda. 

I appreciate his generosity and similarly dedicate 

this effort to his memory. John E. Miller, formerly 

at Harbor Branch Oceanographic Institution Museum, 

Fort Pierce, Florida, provided lists of cataloged holdings 

of amphipods from Florida in that museum. 

Richard W. Heard and Sara E. LeCroy, both at Gulf 

Coast Research Laboratory, Ocean Springs, Mississippi, 

provided a list of benthic macroinvertebrates 

that they had identified from sites off the Florida coast. 

I could not have produced the list of Peracarida without 

the advice of Sara LeCroy, who, through innumerable 

e-mail transmittals over a short period, taught 

me more about Florida peracarids than I had ever 

Phylum Arthropoda 

Subphylum Crustacea Brünnich, 1772 

Class Malacostraca Latreille, 1806 

Subclass Phyllocarida Packard, 1879 

Order Leptostraca Claus, 1880 

Family Nebaliidae Baird, 1850 

Genus Nebalia Leach, 1814 

Nebalia bipes (Fabricius, 1780) 

(Note: This Boreal species (see Dahl, 1985, Sarsia 

70(2-3):135-165) was reported from Dry 

Tortugas by Pearse (1934, Papers from the 

Tortugas Laboratory, Carnegie Institute of 

Washington 28(7):117-124). Pearse’s record 

probably is not N. bipes.) 

Nebalia sp. A of Rakocinski et al., 1993 

Genus Paranebalia Claus, 1880 

Paranebalia longipes Willemoes-Suhm, 1875 

Subclass Hoplocarida Calman, 1904 

Order Stomatopoda Latreille, 1817 

known. I am very grateful for her time and her patience. 

Lawrence G. Abele, Florida State University,Tallahassee, 

provided all of the computer files and original illustrations 

from his and Won Kim’s (1986) work, An Illustrated 

Guide to the Marine Decapod Crustaceans of 

Florida. I am grateful for his generosity and trust. 

Many taxonomic experts reviewed portions of the 

draft list, and each one made significant improvements 

to it. Stomatopoda were reviewed by Raymond B. Manning, 

Smithsonian Institution. Mysidacea were reviewed 

by Wayne Price, Tampa University, and by 

Richard F. Modlin, University of Alabama at Huntsville. 

Amphipoda were reviewed by James D.Thomas, Nova 

University, Dania, Florida, and by Richard Heard and 

Sara LeCroy. Isopoda were reviewed by Brian F. Kensley, 

Smithsonian Institution, and by John C. Markham, 

Arch Cape Marine Laboratory, Arch Cape, Oregon. 

Tanaidacea were reviewed by Kim Larson, The Australian 

Museum, Sydney, and by Richard Heard. 

Cumacea were reviewed by Iorgu Petrescu, Musée 

National d’Histoire naturelle “Grigore Antipa,” 

Bucharest, and by Richard Heard. Decapoda were reviewed 

by Austin B.Williams, NOAA, National Marine 

Fisheries Service, Systematics Laboratory, Smithsonian 

Institution, and by Raymond B. Manning. Errors remaining 

in the list are attributable only to me. 

Checklist of the Malacostracan Crustacea 

Suborder Unipeltata Latreille, 1825 

Superfamily Gonodactyloidea Giesbrecht, 1910 

Family Eurysquillidae Manning, 1977 

Genus Eurysquilla Manning, 1963 

Eurysquilla plumata (Bigelow, 1901) 

Family Gonodactylidae Giesbrecht, 1910 

Genus Neogonodactylus Manning, 1995 

Neogonodactylus bredini (Manning, 1969) 

Neogonodactylus curacaoensis (Schmitt, 1924) 

Neogonodactylus oerstedii (Hansen, 1895) 

Neogonodactylus spinulosus (Schmitt, 1924) 

Neogonodactylus torus (Manning, 1969) 

Neogonodactylus wennerae Manning & Heard, 

1996 

Family Odontodactylidae Manning, 1980 

Genus Odontodactylus Bigelow, 1893 

Odontodactylus brevirostris (Miers, 1884) 


Camp et al. 1998 Macroinvertebrate Checklists D. K. Camp—Malacostraca 

Family Pseudosquillidae Manning, 1977 

Genus Pseudosquilla Dana, 1852 

Pseudosquilla ciliata (Fabricius, 1787) 

Genus Pseudosquillisma Cappola & Manning, 1995 

Pseudosquillisma oculata (Brullé, 1837) 

Superfamily Lysiosquilloidea Giesbrecht, 1910 

Family Lysiosquillidae Giesbrecht, 1910 

Genus Lysiosquilla Dana, 1852 

Lysiosquilla scabricauda (Lamarck, 1818) 

Genus Lysiosquillina Manning, 1995 

Lysiosquillina glabriuscula (Lamarck, 1818) 

Family Nannosquillidae Manning, 1980 

Genus Alachosquilla Schotte & Manning, 1993 

Alachosquilla floridensis (Manning, 1962) 

Genus Bigelowina Schotte & Manning, 1993 

Bigelowina biminiensis (Bigelow, 1893) 

Genus Coronis Desmarest, 1823 

Coronis scolopendra Latreille, 1828 

Genus Nannosquilla Manning, 1963 

Nannosquilla adkisoni Camp & Manning, 1982 

Nannosquilla heardi Camp & Manning, 1982 

Nannosquilla schmitti (Manning, 1962) 

Nannosquilla taylori Manning, 1969 

Nannosquilla whitingi Camp & Manning, 1982 

Family Tetrasquillidae Manning & Camp, 1993 

Genus Tectasquilla Adkison & Hopkins, 1984 

Tectasquilla lutzae Adkison & Hopkins, 1984 

Genus Tetrasquilla Manning & Chace, 1990 

Tetrasquilla mccullochae (Schmitt, 1940) 

Superfamily Squilloidea Latreille, 1803 

Family Squillidae Latreille, 1803 

Genus Alima Leach, 1817 

Alima alba (Bigelow, 1893) 

(= Alima hyalina Leach, 1817 sensu Manning, 

1969) 

(= Alima neptuni (Linnaeus, 1768) sensu Manning 

& Lewinsohn, 1986) 

Genus Cloridopsis Manning, 1968 

Cloridopsis dubia (H. Milne Edwards, 1837) 

Genus Gibbesia Manning & Heard, 1997 

Gibbesia neglecta (Gibbes, 1850) 

Genus Meiosquilla Manning, 1968 

Meiosquilla quadridens (Bigelow, 1893) 

Meiosquilla randalli (Manning, 1962) 

Meiosquilla schmitti (Lemos de Castro, 1955) 

Meiosquilla tricarinata (Holthuis, 1941) 

Genus Squilla Fabricius, 1787 

Squilla chydaea Manning, 1962 

Squilla deceptrix Manning, 1969 

Squilla empusa Say, 1818 

Squilla prasinolineata Dana, 1852 

Squilla rugosa Bigelow, 1893 

Subclass Eumalacostraca Grobben, 1892 

Superorder Peracarida Calman, 1904 

Order Mysidacea Boas, 1883 

Suborder Mysida Boas, 1883 

Family Mysidae Dana, 1850 

Subfamily Siriellinae Norman, 1892 

Genus Siriella Dana, 1850 

Siriella chierchiae Coifmann, 1937 

Siriella thompsonii (H. Milne Edwards, 1837) 

Subfamily Gastrosaccinae Norman, 1892 

Genus Anchialina Norman & Scott, 1906 

Anchialina typica (Krøyer, 1861) 

Genus Bowmaniella Băcescu, 1968 

Bowmaniella bacescui Brattegard, 1970 

Bowmaniella brasiliensis Băcescu, 1968 

Bowmaniella floridana Holmquist, 1975 

(= B. dissimilis (Coifmann, 1937) sensu Brattegard, 

1970) 

Bowmaniella mexicana (W. M.Tattersall, 1951) 

Bowmaniella portoricensis Băcescu, 1968 

Bowmaniella sewelli Brattegard, 1970 

Subfamily Mysinae Dana, 1850 

Tribe Erythropini Hansen, 1910 

Genus Amathimysis Brattegard, 1969 

Amathimysis brattegardi Stuck & Heard, 1981 

Amathimysis gibba Brattegard, 1969 

Genus Erythrops G. O. Sars, 1869 

Erythrops parva Brattegard, 1973 



Tribe Leptomysini Norman, 1892 

Genus Americamysis Price, Heard & Stuck, 1994 

Americamysis almyra (Bowman, 1964) 

Americamysis bahia (Molenock, 1969) 

Americamysis bigelowi (W. M.Tattersall, 1926) 

Americamysis stucki Price, Heard & Stuck, 1994 

Genus Brasilomysis Băcescu, 1968 

Brasilomysis castroi Băcescu, 1968 

Genus Dioptromysis Zimmer, 1915 

Dioptromysis spinosa Brattegard, 1969 

Genus Metamysidopsis W. M.Tattersall, 1951 

Metamysidopsis munda (Zimmer, 1918) 

Metamysidopsis swifti Băcescu, 1969 

Genus Mysidopsis G. O. Sars, 1884 

Mysidopsis brattstroemi Brattegard, 1969 

Mysidopsis eclipes Brattegard, 1969 

Mysidopsis furca Bowman, 1957 

Mysidopsis mortenseni W. M.Tattersall, 1951 

Genus Promysis Dana, 1850 

Promysis atlantica W. M.Tattersall, 1923 

Tribe Mysini Norman, 1892 

Genus Antromysis Creaser, 1936 

Subgenus Parvimysis Brattegard, 1969 

Antromysis (Parvimysis) bahamensis (Brattegard, 

1969) 

Genus Mysidium Dana, 1850 

Mysidium columbiae (Zimmer, 1915) 

Mysidium gracile (Dana, 1852) 

Mysidium integrum W. M.Tattersall, 1951 

Genus Neomysis Czerniavsky, 1882 

Neomysis americana (S. I. Smith, 1873) 

Genus Taphromysis A. H. Banner, 1953 

Taphromysis bowmani Băcescu, 1961 

Taphromysis louisianae A. H. Banner, 1953 

Tribe Heteromysini Norman, 1892 

Genus Heteromysis S. I. Smith, 1873 

Heteromysis beetoni Modlin, 1984 

Heteromysis coralina Modlin, 1987 

Heteromysis dispar Brattegard, 1970 

Heteromysis filitelsona Modlin, 1984 

Heteromysis floridensis Brattegard, 1969 

Heteromysis formosa S. I. Smith, 1873 

Heteromysis guitarti Băcescu, 1968 

Heteromysis hopkinsi Modlin, 1984 

Heteromysis kensleyi Modlin, 1987 

Heteromysis nouveli Brattegard, 1969 

Order Amphipoda Latreille, 1816 

Suborder Ingolfiellidea Hansen, 1903 

Family Ingolfiellidae Hansen, 1903 

Genus Ingolfiella Hansen, 1903 

Ingolfiella fuscina Dojiri & Sieg, 1987 

Suborder Gammaridea Latreille, 1803 

Family Ampeliscidae Costa, 1857 

Genus Ampelisca Krøyer, 1842 

Ampelisca abdita Mills, 1964 

Ampelisca agassizi (Judd, 1896) 

Ampelisca bicarinata Goeke & Heard, 1983 

Ampelisca burkei Barnard & Thomas, 1989 

Ampelisca cristata Holmes, 1908 

Ampelisca holmesi Pearse, 1908 

Ampelisca parapacifica Goeke & Heard, 1984 

Ampelisca schellenbergi Shoemaker, 1933 

Ampelisca vadorum Mills, 1963 

Ampelisca verrilli Mills, 1967 

Ampelisca sp. C of Rakocinski et al., 1993 

Genus Byblis Boeck, 1871 

Byblis sp. 1 of Thomas, 1993 

Family Amphilochidae Boeck, 1871 

Genus Amphilocus Bate, 1862 

Amphilocus cf. casahoya McKinney, 1978 

Amphilocus neapolitanus Della Valle, 1893 

Amphilocus pillaii Barnard & Thomas, 1983 

Genus Gitanopsis G.O. Sars, 1895 

Gitanopsis laguna McKinney, 1978 

Gitanopsis tortugae Shoemaker, 1933 

Gitanopsis sp. B of Rakocinski et al., 1993 

Family Ampithoidae Stebbing, 1899 

Genus Ampithoe Leach, 1814 

Ampithoe longimana Smith, 1873 

Ampithoe pollex Kunkel, 1910 

Ampithoe ramondi Audouin, 1826 

Ampithoe valida Smith, 1873 

Genus Cymadusa Savigny, 1816 

Cymadusa compta (Smith, 1873) 



Cymadusa filosa Savigny, 1816 

Genus Sunampithoe Bate, 1857 

Sunampithoe pelagica H. Milne-Edwards, 1830 

Family Anamixidae Stebbing, 1897 

Genus Anamixis Stebbing, 1897 

Anamixis cavatura Thomas, 1997 

Anamixis hanseni Stebbing, 1897 ? 

Anamixis vanga Thomas, 1997 

Family Aoridae Stebbing, 1899 

Genus Bemlos Shoemaker, 1925 

Bemlos brunneomaculatus (Myers, 1977) 

Bemlos dentischium (Myers, 1977) 

Bemlos kunkelae (Myers, 1977) 

Bemlos mackinneyi (Myers, 1978) 

Bemlos minimus (Myers, 1977) 

Bemlos setosus (Myers, 1978) 

Bemlos spinicarpus spinicarpus (Pearse, 1912) 

Bemlos spinicarpus inermis (Myers, 1979) 

Bemlos tempus (Myers, 1979) 

Bemlos tigrinus (Myers, 1979) 

Bemlos unicornis (Bynum & Fox, 1977) 

Genus Globosolembos Myers, 1988 

Globosolembos smithi (Holmes, 1905) 

Genus Grandidierella Coutière, 1904 

Grandidierella bonnieroides Stephensen, 1947 

Genus Konatopus J. L. Barnard, 1970 

Konatopus sp. 1 of Thomas, 1993 

Genus Lembos Bate, 1857 

Lembos hypacanthus K. H. Barnard, 1916 

Lembos unifasciatus unifasciatus Myers, 1977 

Lembos unifasciatus reductus Myers, 1979 

Lembos websteri Bate, 1857 

Genus Liocuna Myers, 1981 

Liocuna caeca Myers, 1981 

Genus Neomegamphopus Shoemaker, 1942 

Neomegamphopus hiatus Barnard & Thomas, 1987 

Neomegamphopus kalanii Barnard & Thomas, 1987 

Neomegamphopus roosevelti Shoemaker, 1942 

Genus Paramicrodeutopus Myers, 1988 

Paramicrodeutopus myersi (Bynum & Fox, 1977) 

Genus Pedicorophium Karaman, 1981 

Pedicorophium laminosum (Pearse, 1912) 

Genus Plesiolembos Myers, 1988 

Plesiolembos ovalipes (Myers, 1979) 

Plesiolembos rectangulatus (Myers, 1977) 

Genus Rudilemboides Barnard, 1959 

Rudilemboides naglei Bousfield, 1973 

Genus Unciola Say, 1818 

Unciola dissimilis Shoemaker, 1942 

Unciola serrata Shoemaker, 1945 

Family Argissidae Walker, 1904 

Genus Argissa Boeck, 1871 

Argissa hamatipes (Norman, 1869) species complex 

Family Aristiidae Lowry & Stoddart, 1997 

Genus Boca Lowry & Stoddart, 1997 

Boca megachela Lowry & Stoddart, 1997 

Family Bateidae Stebbing, 1906 

Genus Batea Müller, 1865 

Batea bousfieldi (Ortiz, 1991) 

Batea campi (Ortiz, 1991) 

Batea carinata (Shoemaker, 1926) 

Batea catharinensis Müller, 1865 

Batea cuspidata (Shoemaker, 1926) 

Family Biancolinidae J. L. Barnard, 1972 

Genus Biancolina Della Valle, 1893 

Biancolina brassicacephala Lowry, 1974 

Family Cheluridae Allman, 1847 

Genus Chelura Philippi, 1839 

Chelura terebrans Philippi, 1839 

Genus Tropichelura Barnard, 1959 

Tropichelura gomezi Ortiz, 1976 

Family Colomastigidae Stebbing, 1899 

Genus Colomastix Grube, 1861 

Colomastix bousfieldi LeCroy, 1995 

Colomastix camura LeCroy, 1995 

Colomastix cornuticauda LeCroy, 1995 

Colomastix falcirama LeCroy, 1995 

Colomastix gibbosa LeCroy, 1995 



Colomastix halichondriae Bousfield, 1973 

Colomastix heardi LeCroy, 1995 

Colomastix irciniae LeCroy, 1995 

Colomastix janiceae Heard & Perlmutter, 1977 

Colomastix tridentata LeCroy, 1995 

Colomastix sp. 1 of Thomas, 1993 

Family Corophiidae Dana, 1849 

Subfamily Corophiinae Dana, 1849 

Genus Americorophium Bousfield & Hoover, 1997 

Americorophium aquafuscum (Heard & Sikora, 

1972) 

Americorophium ellisi (Shoemaker, 1943) 

Genus Apocorophium Bousfield & Hoover, 1997 

Apocorophium acutum (Chevreux, 1908) 

Apocorophium lacustre (Vanhoffen, 1911) 

Apocorophium louisianum (Shoemaker, 1934) 

Apocorophium simile (Shoemaker, 1934) 

Genus Audulla Chevreux, 1901 

Audulla chelifera Chevreux, 1901 

Genus Chevalia Walker, 1904 

Chevalia aviculae Walker, 1904 

Chevalia carpenteri Barnard & Thomas, 1987 

Chevalia mexicana Pearse, 1913 

Genus Crassicorophium Bousfield & Hoover, 1997 

Crassicorophium bonellii (H. Milne Edwards, 1830) 

Genus Gammaropsis Liljeborg, 1855 

Subgenus Gammaropsis Liljeborg, 1855 

Gammaropsis (Gammaropsis) atlantica Stebbing, 

1888 

Gammaropsis (Gammaropsis) cf. kaumaka Barnard, 

1970 

Gammaropsis (Gammaropsis) sutherlandi Nelson, 

1981 

Genus Laticorophium Bousfield & Hoover, 1997 

Laticorophium cf. baconi (Shoemaker, 1934) 

Genus Microprotopus Norman, 1867 

Microprotopus raneyi Wigley, 1966 

Microprotopus shoemakeri Lowry, 1972 

Genus Monocorophium Bousfield & Hoover, 1997 

Monocorophium acherusicum (Costa, 1851) 

Monocorophium cf. insidiosum (Crawford, 1937) 

Monocorophium tuberculatum (Shoemaker, 1934) 

Genus Photis Krøyer, 1842 

Subgenus Photis Krøyer, 1842 

Photis (Photis) dentata Shoemaker, 1945 

Photis (Photis) longicaudata (Bate & Westwood, 

1863) 

Photis (Photis) macromana McKinney, Kalke, & 

Holland, 1978 

Photis (Photis) melanica McKinney, 1980 

Photis (Photis) pugnator Shoemaker, 1945 

Photis (Photis) trapherus Thomas & Barnard, 1991 

Family Cyproideidae J. L. Barnard, 1974 

Genus Hoplopheonoides Shoemaker, 1956 

Hoplopheonoides obesa Shoemaker, 1956 

Family Dexaminidae Leach, 1814 

Genus Atylus Leach, 1815 

Atylus urocarinatus McKinney, 1980 

Genus Polycheria Haswell, 1879 

Polycheria sp. of Mason et al., 1994 

Family Epimeriidae Boeck, 1871 

(sensu Coleman & Barnard, 1991) 

Genus Epimeria Costa, 1851 

Epimeria longispinosa K. H. Barnard, 1916 (deep 

water) 

Family Eusiridae Stebbing, 1888 

Genus Eusiroides Stebbing, 1888 

Eusiroides sp. 1 of Thomas, 1993 

Genus Pontogeneia Boeck, 1871 

Pontogeneia bartschi Shoemaker, 1948 

Genus Tethygeneia Barnard, 1972 

Tethygeneia longleyi (Shoemaker, 1933) 

Family Gammaridae Leach, 1814 

(sensu Barnard & Barnard, 1990) 

Genus Anamaera Thomas & Barnard, 1985 

Anamaera hixoni Thomas & Barnard, 1985 

Genus Ceradocus Costa, 1853 

Ceradocus sheardi Shoemaker, 1948 

Ceradocus shoemakeri Fox, 1973 

Genus Dulichiella Stout, 1912 

Dulichiella appendiculata (Say, 1818) 



Genus Elasmopus Costa, 1853 

Elasmopus balkomanus Thomas & Barnard, 1988 

Elasmopus levis (Smith, 1873) 

Elasmopus pectenicrus (Bate, 1862) 

Elasmopus pocillimanus (Bate, 1862) 

Elasmopus rapax Costa, 1853 

Genus Gammarus Fabricius, 1775 

Gammarus“macromucronate”species of Heard, 

1982 

(? = Gammarus macromucronatus of Stoner, 1980) 

Gammarus mucronatus Say, 1818 

Gammarus palustris Bousfield, 1969 

Gammarus tigrinus Sexton, 1939 

Gammarus undescribed sp. (near tigrinus) of 

Heard, 1982 

Genus Maera Leach, 1814 

Maera caroliniana Bynum & Fox, 1977 

Maera quadrimana (Dana, 1853) 

Maera rathbunae Pearse, 1908 

Maera williamsi Bynum & Fox, 1977 

Genus Melita Leach, 1814 

Melita elongata Sheridan, 1980 

Melita intermedia Sheridan, 1980 

Melita longisetosa Sheridan, 1980 

Melita nitida Smith, 1873 

Melita sp. 1 of Thomas, 1993 

Genus Netamelita J. L. Barnard, 1962 

Netamelita brocha Thomas & Barnard, 1991 

Genus Protohadzia Zimmerman & Barnard, 1977 

Protohadzia schoenerae (Fox, 1973) 

Genus Spathiopus Thomas & Barnard, 1985 

Spathiopus looensis Thomas & Barnard, 1985 

Genus Tabatzius McKinney & Barnard, 1977 

Tabatzius muelleri (Ortiz, 1976) 

Family Haustoriidae Stebbing, 1906 

Genus Acanthohaustorius Bousfield, 1965 

Acanthohaustorius bousfieldi Frame, 1980 

Acanthohaustorius intermedius Bousfield, 1965 

Acanthohaustorius millsi Bousfield, 1965 

Acanthohaustorius pansus Thomas & Barnard, 

1984 

Acanthohaustorius shoemakeri Bousfield, 1965 

Acanthohaustorius uncinus Foster, 1989 

Acanthohaustorius sp. A of Rakocinski et al., 1993 

Acanthohaustorius sp. B of Rakocinski et al., 1993 

Genus Haustorius Müller, 1775 

Haustorius canadensis Bousfield, 1962 

Haustorius jayneae Foster & LeCroy, 1991 

Genus Lepidactylis Say, 1818 

Lepidactylis dytiscus Say, 1818 

Genus Neohaustorius Bousfield, 1965 

Neohaustorius schmitzi Bousfield, 1965 

Genus Parahaustorius Bousfield, 1965 

Parahaustorius holmesi Bousfield, 1965 

Parahaustorius longimerus Bousfield, 1965 

Parahaustorius obliquus Robertson & Shelton, 

1978 

Genus Protohaustorius Bousfield, 1965 

Protohaustorius cf. bousfieldi Robertson & Shelton, 

1978 

Protohaustorius deichmannae Bousfield, 1965 

Protohaustorius wigleyi Bousfield, 1965 

Genus Pseudohaustorius Bousfield, 1965 

Pseudohaustorius caroliniensis Bousfield, 1965 

Pseudohaustorius sp. A of Rakocinski et al., 1993 

Family Hyalellidae Bulycheva, 1957 

Genus Hyalella S.I. Smith, 1874 

Hyalella azteca Saussure, 1857 (usually freshwater) 

Family Hyalidae Bulycheva, 1957 

Genus Hyale Rathke, 1837 

Hyale media (Dana, 1853) 

Hyale perieri (Lucas, 1846) 

Genus Parhyale Stebbing, 1897 

Parhyale fascigera Stebbing, 1897 

Parhyale hawaiensis (Dana, 1853) 

Genus Parhyalella Kunkel, 1910 

Parhyalella whelpleyi (Shoemaker, 1933) 

Family Iphimediidae Boeck, 1871 


Genus Iphimedia Rathke, 1843 

Iphimedia zora Thomas & Barnard, 1991 

Family Ischyroceridae Stebbing, 1899 

Genus Cerapus Say, 1817 



Cerapus benthophilus Thomas & Heard, 1979 

Cerapus cudjoe Lowry & Thomas, 1991 

Cerapus tubularis Say, 1817 

Cerapus sp. B of Rakocinski et al., 1996 

Genus Erichthonius H. Milne-Edwards, 1830 

Erichthonius brasiliensis (Dana, 1853) 

Genus Ischyrocerus Krøyer, 1838 

Ischyrocerus anguipes Krøyer, 1838 

Genus Jassa Leach, 1814 

Jassa falcata (Montagu, 1808) 

Genus Microjassa Stebbing, 1899 

Microjassa floridensis Conlan, 1995 

Microjassa tetradonta Conlan, 1995 

Family Leucothoidae Dana, 1852 

Genus Leucothoe Leach, 1814 

Leucothoe spinicarpa (Abildgaard, 1789) species 

complex 

Leucothoe sp. 1 of Thomas, 1993 

Family Liljeborgiidae Stebbing, 1899 

Genus Liljeborgia Bate, 1862 

Liljeborgia bousfieldi McKinney, 1979 

Liljeborgia pallida (Bate, 1857) 

(= L. brevicornis (Bruzelius, 1859)) 

Genus Listriella Barnard, 1959 

Listriella barnardi Wigley, 1966 

Listriella carinata McKinney, 1979 

Listriella clymenellae Mills, 1963 

Family Lysianassidae Dana, 1849 

Subfamily Lysianassinae Dana, 1849 

Genus Aruga Holmes, 1908 

Aruga holmesi J. L. Barnard, 1955 

Genus Concarnes Barnard & Karaman, 1991 

Concarnes concavus (Shoemaker, 1933) 

Genus Dissiminassa Barnard & Karaman, 1991 

Dissiminassa homosassa Lowry & Stoddart, 1997 

Genus Lysianassa H. Milne Edwards, 1830 

Lysianassa sp. 1 of Thomas, 1993 

Genus Lysianopsis Holmes, 1905 

Lysianopsis alba Holmes, 1905 

Lysianopsis ozona Lowry & Stoddart, 1997 

Genus Shoemakerella Pirlot, 1936 

Shoemakerella cubensis (Stebbing, 1897) 

Subfamily Tryphosinae Lowry & Stoddart, 1997 

Genus Hippomedon Boeck, 1871 

Hippomedon pensacola Lowry & Stoddart, 1997 

Genus Orchomenella Sars, 1890 

Orchomenella perdido Lowry & Stoddart, 1997 

Orchomenella thomasi Lowry & Stoddart, 1997 

Family Megaluropidae Thomas & Barnard, 1986 

Genus Gibberosus Thomas & Barnard, 1986 

Gibberosus myersi (McKinney, 1980) 

Family Melphidippidae Stebbing, 1899 

Genus Hornellia Walker, 1904 

Subgenus Metaceradocus Chevreaux, 1925 

Hornellia (Metaceradocus) tequestae Thomas & 

Barnard, 1986 

Family Ochlesidae Stebbing, 1910 


Genus Curidia Thomas, 1983 

Curidia debrogania Thomas, 1983 

Family Oedicerotidae Liljeborg, 1865 

Genus Americhelidium Bousfield & Chevrier, 1996 

Americhelidium americanum (Bousfield, 1973) 

Genus Ameroculodes Bousfield & Chevrier, 1996 

Ameroculodes edwardsi (Holmes, 1905) 

Genus Deflexiodes Bousfield & Chevrier, 1996 

Deflexiodes intermedius (Shoemaker, 1930) 

Genus Hartmanodes Bousfield & Chevrier, 1996 

Hartmanodes nyei (Shoemaker, 1933) 

Genus Perioculodes G.O. Sars, 1895 

Perioculodes cerasinus Thomas & Barnard, 1985 

Family Phliantidae Stebbing, 1899 

Genus Pariphinotus Kunkel, 1910 

Pariphinotus seclusus (Shoemaker, 1933) 



Family Phoxocephalidae G.O. Sars, 1895 

Subfamily Brolginae 

Barnard & Drummond, 1978 

Genus Eobrolgus J. L. Barnard, 1979 

Eobrolgus spinosus (Holmes, 1905) 

Subfamily Metharpiniinae 

Jarrett & Bousfield, 1994 

Genus Metharpinia Schellenberg, 1931 

Metharpinia floridana (Shoemaker, 1933) 

Genus Rhepoxynius Barnard, 1979 

Rhepoxynius epistomus (Shoemaker, 1938) 

Rhepoxynius hudsoni Barnard & Barnard, 1982 

Family Platyischnopidae 

Barnard & Drummond, 1979 

Genus Eudevenopus Thomas & Barnard, 1983 

Eudevenopus honduranus Thomas & Barnard, 1983 

Family Pleustidae Buchholz, 1874 

Subfamily Parapleustinae 

Bousfield & Hendrycks, 1994 

Genus Parapleustes Buchholz, 1874 

Parapleustes aestuarius Watling & Maurer, 1973 

Subfamily Stenopleustinae 

Bousfield & Hendrycks, 1994 

Genus Stenopleustes G.O. Sars, 1895 

Stenopleustes gracilis (Holmes, 1905) 

Family Podoceridae Leach, 1814 

Genus Podocerus Leach, 1814 

Podocerus brasiliensis (Dana, 1853) 

Podocerus chelonophilus (Chevreaux & de Guerne, 

1888) 

Podocerus kleidus Thomas & Barnard, 1992 

Family Pontoporeiidae Dana, 1855 

Genus Bathyporeia Lindstrom, 1855 

Bathyporeia parkeri Bousfield, 1973 

Family Sebidae Walker, 1908 

Genus Seba Bate, 1862 

Seba tropica McKinney, 1980 

Family Stenothoidae Boeck, 1871 

Genus Parametopella Gurjanova, 1938 

Parametopella cypris (Holmes, 1905) 

Parametopella inquilina Watling, 1976 

Parametopella cf. texensis McKinney, Kalke & Holland, 

1978 

Genus Stenothoe Dana, 1852 

Stenothoe gallensis Walker, 1904 

(= S. crenulata Chevreux, 1907) 

Stenothoe georgiana Bynum & Fox, 1977 

Stenothoe minuta Holmes, 1905 

Stenothoe symbiotica Shoemaker, 1956 (deeper 

water) 

Stenothoe sp. A of Rakocinski et al., 1996 

Family Synopiidae Dana, 1855 

Genus Garosyrrhoe Barnard, 1964 

Garosyrrhoe cf. bigarra (Barnard, 1962) 

Genus Metatiron Rabindrinath, 1972 

Metatiron cf. bellairsi (Just, 1981) 

Metatiron triocellatus (Goeke, 1982 ) 

Metatiron tropakis (Barnard, 1972) 

Genus Synopia Dana, 1852 

Synopia ultramarinaDana, 1853 

Family Talitridae Rafinesque, 1815 

Genus Americorchestia Bousfield, 1991 

Americorchestia heardi Bousfield, 1991 

Americorchestia longicornis (Say, 1817) 

Americorchestia salomani Bousfield, 1991 

Genus Chelorchestia Bousfield, 1984 

Chelorchestia sp. A of Bousfield, 1984 

(? = Orchestia sp. (near O. costaricana) of Heard, 

1982) 

Chelorchestia sp. B of Bousfield, 1984 

Genus Orchestia Leach, 1814 

Orchestia grillus Bosc, 1802 

Genus Platorchestia Bousfield, 1982 

Platorchestia platensis (Krøyer, 1845) 

Genus Tethorchestia Bousfield, 1984 

Tethorchestia sp. B of Bousfield, 1984 

(= “checker-back”beachflea) 

(= Orchestia, new species of Holmquist, 1982) 



Genus Uhlorchestia Bousfield, 1984 

Uhlorchestia uhleri (Shoemaker, 1930) 

Uhlorchestia spartinophila Bousfield & Heard, 

1986 

Suborder Caprellidea Leach, 1814 

Family Caprellidae White, 1847 

Genus Caprella Lamarck, 1801 

Caprella andreae Mayer, 1890 

Caprella danilevskii Czerniavski, 1868 

Caprella equilibra Say, 1818 

Caprella penantis Leach, 1814 

Family Caprellinoididae Laubitz, 1993 

Genus Pseudaeginella Mayer, 1890 

Pseudaeginella biscaynensis (McCain, 1968) 

Family Pariambidae Laubitz, 1993 

Genus Deutella Mayer, 1890 

Deutella incerta (Mayer, 1903) 

Genus Hemiaegina Mayer, 1890 

Hemiaegina minuta Mayer, 1890 

Genus Paracaprella Mayer, 1890 

Paracaprella pusilla Mayer, 1890 

Paracaprella cf. temir (vide Nelson, 1995) 

Paracaprella tenuis Mayer, 1903 

Family Phtisicidae Vassilenko, 1968 

Subfamily Phtisicinae Vassilenko, 1968 

Genus Hemiproto McCain, 1968 

Hemiproto wigleyi McCain, 1968 

Genus Phtisica Slabber, 1769 

Phtisica marina Slabber, 1769 

Family Protellidae McCain, 1970 

Genus Metaprotella Mayer, 1890 

Metaprotella hummelincki McCain, 1968 

Order Isopoda Latreille, 1817 

Suborder Anthuridea Leach, 1814 

Family Anthuridae Leach, 1814 

Genus Amakusanthura Nunomura, 1977 

Amakusanthura magnifica (Menzies & Frankenberg, 

1966) 

Amakusanthura signata (Menzies & Glynn, 1968) 

Genus Cyathura Norman & Stebbing, 1886 

Cyathura polita (Stimpson, 1855) 

Genus Eisothistos Haswell, 1884 

Eisothistos petrensis Kensley, 1984 

Genus Mesanthura Barnard, 1914 

Mesanthura cf. bivittata Kensley, 1987 

Mesanthura fasciata Kensley, 1982 

Mesanthura hopkinsi Hooker, 1985 

Mesanthura looensis Kensley & Schotte, 1987 

Mesanthura paucidens Menzies & Glynn, 1968 

Mesanthura pulchra Barnard, 1925 

(= Mesanthura floridensis Menzies & Kruczynski, 

1983) 

Genus Pendanthura Menzies & Glynn, 1968 

Pendanthura tanaiformis Menzies & Glynn, 1968 

Genus Ptilanthura Harger, 1878 

Ptilanthura colpos Kensley, 1996 

Ptilanthura tenuis Harger, 1878 

(= Ptilanthura tricarina Menzies & Frankenberg, 

1966) 

Genus Skuphonura Barnard, 1925 

Skuphonura lindae Menzies & Kruczynski, 1983 

Family Hyssuridae Wägele, 1981 

Genus Hyssura Norman & Stebbing, 1886 

Hyssura bacescui (George & Negoescu-Vladescu, 

1982) 

Genus Kupellonura Barnard, 1925 

Kupellonura formosa (Menzies & Frankenberg, 

1966) 

Genus Neohyssura Amar, 1953 

Neohyssura irpex (Menzies & Frankenberg, 1966) 

Genus Xenanthura Barnard, 1925 

Xenanthura brevitelson Barnard, 1925 

Family Paranthuridae Menzies & Glynn, 1968 

Genus Accalathura Barnard, 1925 

Accalathura crenulata (Richardson, 1901) 

Genus Paranthura Bate & Westwood, 1868 

Paranthura infundibulata Richardson, 1902 

Suborder Asellota Latreille, 1803 

Superfamily Gnathostenetroidoidea 

Kussakin, 1967 



Family Gnathostenetroididae Kussakin, 1967 

Genus Gnathostenetroides Amar, 1957 

Gnathostenetroides pugio Hooker, 1985 

Superfamily Janiroidea Sars, 1899 

Family Incertae Sedis 

Genus Mexicope Hooker, 1985 

Mexicope kensleyi Hooker, 1985 

Family Janiridae Sars, 1899 

Genus Carpias Richardson, 1902 

Carpias algicola (Miller, 1941) 

Carpias bermudensis Richardson, 1902 

Carpias floridensis Menzies & Kruczynski, 1983 

Carpias harrietae Pires, 1981 

Carpias minutus (Richardson, 1902) 

Family Joeropsidae Nordenstam, 1933 

Genus Joeropsis Koehler, 1885 

Joeropsis coralicola Schultz & McCloskey, 1967 

Joeropsis rathbunae Richardson, 1902 

Family Munnidae Sars, 1899 

Genus Uromunna Menzies, 1962 

Uromunna hayesi Robertson, 1978 

Uromunna reynoldsi Frankenberg & Menzies, 

1966 

Family Paramunnidae Vanhöffen, 1914 

Genus Munnogonium George & Strömberg, 1968 

Munnogonium wilsoni Hooker, 1985 

Family Pleurocopidae Fresi & Schiecke, 1972 

Genus Pleurocope Walker, 1901 

Pleurocope floridensis Hooker, 1985 

Family Santiidae Kussakin, 1988 

Genus Santia Silvertsen & Holthuis, 1980 

Santia milleri (Menzies & Glynn, 1968) 

Superfamily Stenetrioidea Hansen, 1905 

Family Stenetriidae Hansen, 1905 

Genus Hansenium Serov & Wilson, 1995 

Hansenium stebbingi (Richardson, 1902) 

Genus Stenetrium Haswell, 1881 

Stenetrium serratum Hansen, 1904 

Suborder Epicaridea Latreille, 1831 

Family Bopyridae Rafinesque, 1815 

Subfamily Athelginae 

Codreanu & Codreanu, 1956 

Genus Parathelges Bonnier, 1900 

Parathelges occidentalis Markham, 1972 

Genus Stegophryxus Thompson, 1902 

Stegophryxus hyptius Thompson, 1902 

Subfamily Bopyrinae Rafinesque, 1815 

Genus Bopyrina Kossmann, 1881 

Bopyrina abbreviata Richardson, 1904 

Genus Bopyrinella Nierstrasz & Brender à Brandis, 

1925 

Bopyrinella thorii (Richardson, 1904) 

Genus Bopyrione Bourdon & Markham, 1980 

Bopyrione synalphei Bourdon & Markham, 1980 

Genus Ovobopyrus Markham, 1985 

Ovobopyrus alphezemiotes Markham, 1985 

Genus Parabopyrella Markham, 1985 

Parabopyrella lata (Nierstrasz & Brender à Brandis, 

1929) 

Parabopyrella mortenseni (Nierstrasz & Brender à 

Brandis, 1929) 

Parabopyrella richardsonae (Nierstrasz & Brender 

à Brandis, 1929) 

Genus Parabopyriscus Markham, 1985 

Parabopyriscus stellatus Markham, 1985 

Genus Probopyria Markham, 1985 

Probopyria alphei (Richardson, 1900) 

Genus Probopyrinella Nierstrasz & Brender à Brandis, 

1929 

Probopyrinella heardi Adkison, 1984 

Probopyrinella latreuticola (Gissler, 1882) 

Genus Probopyrus Giard & Bonnier, 1888 

Probopyrus floridensis Richardson, 1904 

Probopyrus pandalicola (Packard, 1879) 

Genus Schizobopyrina Markham, 1985 

Schizobopyrina urocaridis (Richardson, 1904) 

Genus Synsynella Hay, 1917 



Synsynella choprae (Pearse, 1932) 

Synsynella deformans Hay, 1917 

Synsynella integra Bourdon, 1981 

Genus Urobopyrus Richardson, 1904 

Urobopyrus processae Richardson, 1904 

Subfamily Hemiarthrinae Markham, 1972 

Genus Allodiplophryxus Markham, 1985 

Allodiplophryxus floridanus Markham, 1985 

Genus Azygopleon Markham, 1985 

Azygopleon schmitti (Pearse, 1932) 

Genus Diplophryxus Richardson, 1904 

Diplophryxus sp. (see Markham, 1985) 

Genus Eophrixus Caroli, 1930 

Eophrixus subcaudalis (Hay, 1917) 

Genus Hemiarthrus Giard & Bonnier, 1887 

Hemiarthrus synalphei (Pearse, 1950) 

Genus Hyperphrixus Nierstrasz & Brender à Brandis, 

1931 

Hyperphrixus castrensis Markham, 1985 

Genus Loki Markham, 1972 

Loki circumsaltanus Markham, 1972 

Genus Metaphrixus Nierstrasz & Brender à Brandis, 

1931 

Metaphrixus carolii Nierstrasz & Brender à Brandis, 

1931 

Subfamily Ioninae H. Milne Edwards, 1840 

(emend. R. Codreanu, 1967) 

Genus Cancricepon Giard & Bonnier, 1887 

Cancricepon choprae (Nierstrasz & Brender à 

Brandis, 1925) 

Genus Dactylokepon Stebbing, 1910 

Dactylokepon sulcipes Adkison, 1982 

Genus Gigantione Kossmann, 1881 

Gigantione mortenseni Adkison, 1984 

Gigantione uberlackerae Adkison, 1984 

Genus Grapsicepon Giard & Bonnier, 1887 

Grapsicepon edwardsi Giard & Bonnier, 1888 

Genus Leidya Cornalia & Panceri, 1861 

Leidya bimini Pearse, 1951 

Leidya distorta (Leidy, 1855) 

Subfamily Pseudioninae R. Codreanu, 1967 

Genus Anuropodione Bourdon, 1967 

Anuropodione megacephalon Markham, 1973 

Genus Aporobobyrina Shiino, 1934 

Aporobobyrina anomala Markham, 1974 

Genus Aporobopyrus Nobili, 1906 

Aporobopyrus collardi Adkison, 1988 

Aporobopyrus curtatus (Richardson, 1904) 

Genus Asymmetrione Codreanu, Codreanu, & Pike, 

1965 

Asymmetrione clibanarii Markham, 1975 

Asymmetrione desultor Markham, 1975 

Genus Bopyrissa Nierstrasz & Brender à Brandis, 

1931 

Bopyrissa wolffi Markham, 1978 

Genus Kolourione Markham, 1978 

Kolourione premordica Markham, 1978 

Genus Munidion Hansen, 1897 

Munidion irritans Boone, 1927 

Munidion longipedis Markham, 1975 

Genus Pleurocrypta Hesse, 1865 

Pleurocrypta floridana Markham, 1974 

Genus Pseudione Kossman, 1881 

Pseudione ampla Markham, 1988 

Pseudione cognata Markham, 1985 

Pseudione overstreeti Adkison & Heard, 1995 

Pseudione parvimanus Adkison, 1985 

Family Entoniscidae F. Müller, 1871 

Genus Synalpheion Coutière, 1908 

Synalpheion giardi Coutière, 1908 

Suborder Flabellifera Sars, 1882 

Family Aegidae Leach, 1815 

Genus Aega Leach, 1815 

Aega (Aega) deshaysiana (H. Milne Edwards, 1840) 

(= A. antillensis Schioedte & Meinert, 1879) 

Genus Rocinela Leach, 1818 

Rocinela insularis Schioedte & Meinert, 1879 



Rocinela signata Schioedte & Meinert, 1879 

Family Tridentellidae Bruce, 1984 

Genus Tridentella Richardson, 1905 

Tridentella ornata (Richardson, 1911) 

(= Aega ornata Richardson, 1911) 

Family Cirolanidae Dana, 1852 

Subfamily Cirolaninae Dana, 1852 

Genus Anopsilana Paulian & Delamare Debouteville, 

1956 

Anopsilana jonesi Kensley, 1987 (unpublished 

FDEP record) 

Genus Cirolana Leach, 1818 

Cirolana parva Hansen, 1890 

Genus Natatolana Bruce, 1981 

Natatolana borealis (Lilljeborg, 1851) 

(as Cirolana borealis Lilljeborg, 1851, in Richardson, 

1905) 

Natatolana n. sp. of Keable & Bruce, 1997 

(as C. borealis Lilljeborg, 1851, in Bird, 1981) 

(as C. borealis Lilljeborg, 1851, in Menzies & 

Kruczynski, 1983) 

Subfamily Conilerinae Kensley & Schotte, 1989 

Genus Politolana Bruce, 1981 

Politolana impressa (Harger, 1883) 

Politolana polita (Stimpson, 1853) 

Subfamily Eurydicinae Stebbing, 1905 

Genus Eurydice Leach, 1815 

Eurydice convexa Richardson, 1900 

Eurydice littoralis (Moore, 1901) 

Eurydice personata Kensley, 1987 

Eurydice piperata Menzies & Frankenberg, 1966 

Genus Excirolana Richardson, 1912 

Excirolana braziliensis Richardson, 1912 

Excirolana mayana (Ives, 1891) 

Genus Metacirolana Nierstrasz, 1931 

Metacirolana sphaeromiformis (Hansen, 1890) 

Family Corallanidae Hansen, 1890 

Genus Alcirona Hansen, 1890 

Alcirona insularis Hansen, 1890 

Alcirona krebsii Hansen, 1890 

Genus Excorallana Stebbing, 1904 

Excorallana acuticauda (Miers, 1881) 

Excorallana delaneyi Stone & Heard, 1989 

Excorallana mexicana Richardson, 1905 

Excorallana sexticornis (Richardson, 1901) 

Excorallana tricornis tricornis (Hansen, 1890) 

Genus Nalicora Moore, 1901 

Nalicora rapax Moore, 1901 

Family Cymothoidae Leach, 1818 

Genus Agarna Schioedte & Meinert, 1883 

Agarna cumulus (Haller, 1880) 

Genus Anilocra Leach, 1818 

Anilocra acanthuri Williams & Williams, 1981 

Anilocra acuta Richardson, 1910 

Anilocra haemuli Williams & Williams, 1981 

Anilocra laticauda H. Milne Edwards, 1840 

Genus Cymothoa Fabricius, 1793 

Cymothoa caraibica Bovallius, 1885 

Cymothoa excisa Perty, 1833 

Cymothoa oestrum (Linnaeus, 1793) 

Genus Elthusa Schioedte & Meinert, 1884 

Elthusa tropicalis (Menzies & Kruczynski, 1983) 

Genus Livoneca Leach, 1818 

Livoneca ovalis (Say, 1818) 

Livoneca redmanii Leach, 1818 

Genus Mothocya Costa, 1851 

Mothocya bohlkeorum Williams & Williams, 1982 

Mothocya nana (Schioedte & Meinert, 1884) 

Genus Nerocila Leach, 1818 

Nerocila acuminata Schioedte & Meinert, 1881 

Genus Olencira Leach, 1818 

Olencira praegustator (Latrobe, 1802) 

Family Limnoriidae Harger, 1879 

Genus Limnoria Leach, 1814 

Limnoria cf. indica Becker & Kampf, 1958 

Limnoria pfefferi Stebbing, 1904 

Limnoria platycauda Menzies, 1957 

Limnoria saseboensis Menzies, 1957 

Limnoria simulata Menzies, 1957 



Limnoria tuberculata Sowinsky, 1884 

(= L. tripunctata Menzies, 1951) 

Genus Paralimnoria Menzies, 1957 

Paralimnoria andrewsi (Calman, 1910) 

Family Serolidae Dana, 1852 

Genus Heteroserolis Brandt, 1991 

Heteroserolis mgrayi (Menzies & Frankenberg, 

1966) 

Family Sphaeromatidae H. Milne Edwards, 1840 

Subfamily Ancininae Tattersall, 1905 

Genus Ancinus H. Milne Edwards, 1840 

Ancinus depressus (Say, 1818) 

Subfamily Cassidininae Iverson, 1982 

Genus Cassidinidea Hansen, 1905 

Cassidinidea ovalis (Say, 1818) 

Subfamily Dynameninae Bowman, 1981 

Genus Dynamenella Hansen, 1905 

Dynamenella angulata (Richardson, 1901) 

Genus Paracerceis Hansen, 1905 

Paracerceis caudata (Say, 1818) 

Paracerceis glynni Kensley, 1984 

Genus Paradella Harrison & Holdich, 1982 

Paradella dianae (Menzies, 1962) 

Paradella quadripunctata (Menzies & Glynn, 1968) 

Subfamily Sphaeromatinae 

H. Milne Edwards, 1840 

Genus Exosphaeroma Stebbing, 1900 

Exosphaeroma diminuta Menzies & Frankenberg, 

1966 

Genus Harrieta Kensley, 1987 

Harrieta faxoni (Richardson, 1905) 

Genus Sphaeroma Bosc, 1802 

Sphaeroma quadridentata Say, 1818 

Sphaeroma terebrans Bate, 1866 

Sphaeroma walkeri Stebbing, 1905 

Suborder Gnathiidea Leach, 1814 

Family Gnathiidae Harger, 1879 

Genus Gnathia Leach, 1814 

Gnathia floridensis Menzies & Kruczynski, 1983 

Gnathia cf. rathi Kensley, 1984 

Suborder Microcerberidea Lang, 1961 

Family Microcerberidae Karaman, 1933 

Genus Coxicerberus Wägele,Voelz, & McArthur, 1995 

Coxicerberus mirabilis (Chappuis & Delamare- 

Deboutteville, 1956) 

Suborder Oniscidea Latreille, 1803 

Infraorder Tylomorpha Vandel, 1943 

Family Tylidae H. Milne Edwards, 1840 

Genus Tylos Latreille, 1826 

Tylos latreillei Audouin & Savigny, 1826 

Tylos marcuzzii Soika, 1954 

Tylos niveus Budde-Lund, 1885 

Tylos wegeneri Vandel, 1952 

Infraorder Ligiamorpha Vandel, 1943 

Section Diplocheta Vandel, 1957 

Family Ligiidae Brandt, 1883 

Genus Ligia Fabricius, 1798 

Ligia baudiniana H. Milne Edwards, 1840 

Ligia exotica Roux, 1828 

Ligia olfersii Brandt, 1833 

Section Crinocheta Legrand, 1946 

Superfamily Oniscoidea Dana, 1852 

Family Halophilosciidae Vandel, 1973 

Genus Vandeloscia Roman, 1977 

Vandeloscia culebrae (Moore, 1902) 

(= Philoscia miamiensis Schultz, 1966) 

Family Philosciidae Vandel, 1952 

Genus Sayoscia Schultz, 1983 

Sayoscia vittata (Say, 1818) 

Family Rhyscotidae Arcangeli, 1947 

Genus Rhyscotus Budde-Lund, 1885 

Rhyscotus texensis (Richardson, 1905) 

Family Scyphacidae Dana, 1853 

Genus Scyphacella Smith, 1873 

Scyphacella arenicola Smith, 1873 

Suborder Valvifera Sars, 1882 



Family Arcturidae Sars, 1897 

Subfamily Arcturinae Sars, 1897 

Genus Arcturella G. O. Sars, 1897 

Arcturella spinata Menzies & Kruczynski, 1983 

Arcturella bispinata Menzies & Kruczynski, 1983 

Genus Astacilla Cordiner, 1793 

Astacilla cymodocea Menzies & Glynn, 1968 

Astacilla lauffi Menzies & Frankenberg, 1966 

Subfamily Edwinjoycinae Müller, 1993 

Genus Edwinjoycea Menzies & Kruczynski, 1983 

Edwinjoycea horologium Menzies & Kruczynski, 

1983 

Family Chaetiliidae Dana, 1853 

Genus Chiridotea Harger, 1878 

Chiridotea arenicola Wigley, 1960 

Chiridotea excavata Harper, 1974 

Family Idoteidae Fabricius, 1798 

Subfamily Idoteinae Dana, 1852 

Genus Cleantioides Kensley & Kaufman, 1978 

Cleantioides planicauda (Benedict, 1899) 

Genus Edotia Guérin-Ménéville, 1843 

Edotia lyonsi (Menzies & Kruczynski, 1983) 

Edotia triloba (Say, 1818) 

(= Edotia montosa (Stimpson, 1853)) 

Genus Erichsonella Richardson, 1901 

Erichsonella attenuata (Harger, 1873) 

Erichsonella filiformis (Say, 1818) 

Erichsonella floridana Benedict, in Richardson, 

1901 

Genus Idotea Fabricius, 1798 

Idotea balthica (Pallas, 1772) 

Idotea metallica Bosc, 1802 

Order Tanaidacea Hansen, 1895 

Suborder Tanaidomorpha Sieg, 1980 

Superfamily Tanaoidea Dana, 1849 

Family Tanaidae Dana, 1849 

Subfamily Pancolinae Sieg, 1980 

Genus Zeuxo Templeton, 1840 

Subgenus Parazeuxo Sieg, 1980 

Zeuxo (Parazeuxo) coralensis Sieg, 1980 

Zeuxo (Parazeuxo) maldivensis Sieg, 1980 

Superfamily Paratanaoidea Lang, 1949 

Family Anarthruridae Lang, 1971 

Genus Araphura Bird & Holdich, 1984 

Araphura higginsi Sieg & Dojiri, 1989 

Family Leptocheliidae Lang, 1973 

Genus Hargeria Lang, 1973 

Hargeria rapax (Harger, 1879) 

Genus Leptochelia Dana, 1849 

Leptochelia dubia Krøyer,1842 

Leptochelia cf. forresti (Stebbing, 1896) 

Leptochelia sp. B of Racocinski et al., 1996 

Family Nototanaidae Sieg, 1976 

Genus Nototanoides Sieg & Heard, 1985 

Nototanoides trifurcatus Sieg & Heard, 1985 

Genus Teleotanais Lang, 1956 

Teleotanais gerlachi Lang, 1956 

Family Pseudotanaidae Sieg, 1973 

Subfamily Cryptocopinae Sieg, 1977 

Genus Iungentitanais Sieg, 1977 

Iungentitanais primitivus (Sieg, 1973) 

Subfamily Pseudotanainae Sieg, 1973 

Genus Pseudotanais G. O. Sars, 1882 

Subgenus Akanthinotanais Sieg, 1977 

Pseudotanais (Akanthinotanais) mortenseni Sieg, 

1977 

Suborder Apseudomorpha Sieg, 1980 

Superfamily Apseudoidea Leach, 1814 

Family Apseudidae Leach, 1814 

Genus Apseudes Leach, 1814 

Apseudes propinquus Richardson, 1902 

Family Cirratodactylidae Gardiner, 1973 

Genus Cirratodactylus Gardiner, 1973 

Cirratodactylus floridensis Gardiner, 1973 

Family Kalliapseudidae Lang, 1956 

Genus Kalliapseudes Stebbing, 1909 

Kalliapseudes bahamensis Sieg, 1982 

Kalliapseudes sp. A of Racocinski et al., 1996 



Family Pagurapseudidae Lang, 1970 

Genus Pagurapseudes Whitelegge, 1901 

Pagurapseudes largoensis McSweeny, 1982 

Family Parapseudidae Gu¸tu, 1981 

Genus Halmyrapseudes Băcescu & Gu¸tu, 1974 

Halmyrapseudes bahamensis Băcescu & Gu¸tu, 1974 

Order Cumacea Krøyer, 1846 

Family Bodotriidae T. Scott, 1901 

Subfamily Bodotriinae T. Scott, 1901 

Genus Cyclaspis G. O. Sars, 1865 

Cyclaspis bacescui Omholt & Heard, 1982 

Cyclaspis platymerus Zimmer, 1944 

Cyclaspis pustulata Zimmer, 1943 

Cyclaspis unicornis Calman, 1907 

Cyclaspis varians Calman, 1912 

Cyclaspis sp. B of Rakocinski et al., 1993 

Cyclaspis sp. C of Rakocinski et al., 1993 

Subfamily Mancocuminae Watling, 1977 

Genus Spilocuma Watling, 1977 

Spilocuma salomani Watling, 1977 

Spilocuma watlingi Omholt & Heard, 1979 

Subfamily Vaunthompsoniinae G. O. Sars, 1878 

Genus Vaunthompsonia Bate, 1859 

Vaunthompsonia floridana Băcescu, 1971 

Vaunthompsonia minor Zimmer, 1944 

Family Diastylidae Bate, 1856 

Genus Diastylis Say, 1818 

Diastylis bispinosa (Stimpson, 1853) 

(= Diastylis quadrispinosa G. O. Sars, 1871) 

Genus Oxyurostylis Calman, 1912 

Oxyurostylis lecroyae Roccatagliata & Heard, 1995 

Oxyurostylis smithi Calman, 1912 

Family Leuconidae G. O. Sars, 1878 

Genus Leucon Krøyer, 1846 

Subgenus Leucon Krøyer, 1846 

Leucon (Leucon) americanus Zimmer, 1943 

Family Nannastacidae Bate, 1866 

Genus Almyracuma Jones & Burbanck, 1959 

Almyracuma cf. proximoculi Jones & Burbanck, 

1959 

Genus Campylaspis G. O. Sars, 1865 

Campylaspis heardi Muradian-Ciamician, 1980 

Campylaspis sp. A of Rakocinski et al., 1996 

Genus Cubanocuma Băcescu & Muradian, 1977 

Cubanocuma gutzui Băcescu & Muradian, 1977 

Genus Cumella G. O. Sars, 1865 

Subgenus Cumella G. O. Sars, 1865 

Cumella (Cumella) agglutinanta Băcescu, 1971 

Cumella (Cumella) coralicola Băcescu, 1971 

Cumella (Cumella) garrityi Băcescu & Muradian, 

1977 

Cumella (Cumella) gomoiui Băcescu & Muradian, 

1977 

Cumella (Cumella) pilosa Băcescu, 1971 

Cumella (Cumella) tripunctata Băcescu, 1971 

Cumella (Cumella) vicina Zimmer, 1944 

Subgenus Cumewingia Băcescu, 1971 

Cumella (Cumewingia) caribbeana Băcescu, 1971 

Cumella (Cumewingia) clavicauda Calman, 1911 

Cumella (Cumewingia) serrata Calman, 1911 

Subgenus unknown 

Cumella sp. A of Rakocinski et al., 1996 

Genus Elasocumella Watling, 1991 

Elasocumella micruropus (Zimmer, 1943) 

Superorder Eucarida Calman, 1904 

Order Euphausiacea Dana, 1852 

Family Euphausiidae Dana, 1852 

Genus Euphausia Dana, 1850 

Euphausia americana Hansen, 1911 

Euphausia brevis Hansen, 1905 

Euphausia gibboides Ortmann, 1893 

Euphausia hemigibba Hansen, 1910 

Euphausia mutica Hansen, 1905 

Euphausia pseudogibba Ortmann, 1893 

Euphausia tenera Hansen, 1905 

Genus Nematobrachion Calman, 1905 

Nematobrachion flexipes (Ortmann, 1893) 

Genus Nematoscelis G. O. Sars, 1883 

Nematoscelis atlantica Hansen, 1910 

Nematoscelis microps G. O. Sars, 1883 

Nematoscelis tenella G. O. Sars, 1883 

Genus Stylocheiron G. O. Sars, 1883 

Stylocheiron abbreviatum G. O. Sars, 1883 



Stylocheiron carinatum G. O. Sars, 1883 

Stylocheiron suhmii G. O. Sars, 1883 

Genus Thysanopoda Latreille, 1831 

Thysanopoda aequalis Hansen, 1905 

Thysanopoda monacantha Ortmann, 1893 

Thysanopoda tricuspida H. Milne Edwards, 1837 

Order Decapoda Latreille, 1803 

Suborder Dendrobranchiata Bate, 1888 

Superfamily Penaeoidea Rafinesque, 1815 

Family Penaeidae Rafinesque, 1818 

Genus Farfantepenaeus Burukovsky, 1972 

Farfantepenaeus aztecus (Ives, 1891) 

Farfantepenaeus brasiliensis (Latreille, 1817) 

Farfantepenaeus duorarum (Burkenroad, 1939) 

Genus Litopenaeus Pérez Farfante, 1969 

Litopenaeus setiferus (Linnaeus, 1767) 

Genus Metapenaeopsis Bouvier, 1905 

Metapenaeopsis gerardoi Pérez Farfante, 1971 

Metapenaeopsis goodei (Smith, 1885) 

Metapenaeopsis smithi (Schmitt, 1924) 

Genus Rimapenaeus Pérez Farfante & Kensley, 1997 

Rimapenaeus constrictus (Stimpson, 1874) 

Rimapenaeus similis (Smith, 1885) 

Genus Trachypenaeopsis Burkenroad, 1934 

Trachypenaeopsis mobilispinis (Rathbun, 1920) 

Genus Xiphopenaeus Smith, 1869 

Xiphopenaeus kroyeri (Heller, 1862) 

Family Sicyoniidae Ortmann, 1898 

Genus Sicyonia H. Milne Edwards, 1830 

Sicyonia brevirostris Stimpson, 1871 

Sicyonia burkenroadi Cobb, 1971 

Sicyonia dorsalis Kingsley, 1878 

Sicyonia laevigata Stimpson, 1871 

Sicyonia olgae Pérez Farfante, 1980 

Sicyonia parri (Burkenroad, 1934) 

Sicyonia typica (Boeck, 1864) 

Family Solenoceridae 

Wood-Mason & Alcock, 1891 

Genus Solenocera Lucas, 1849 

Solenocera atlantidis Burkenroad, 1939 

Superfamily Sergestoidea Dana, 1852 

Family Sergestidae Dana, 1852 

Genus Acetes H. Milne Edwards, 1830 

Acetes americanus carolinae Hansen, 1933 

Family Luciferidae De Haan, 1849 

Genus Lucifer Thompson, 1829 

Lucifer faxoni Borradaile, 1915 

Suborder Pleocyemata Burkenroad, 1963 

Infraorder Stenopodidea Huxley, 1878 

Family Spongicolidae Schram, 1986 

Genus Microprosthema Stimpson, 1860 

Microprosthema looensis Goy & Felder, 1988 

Microprosthema manningi Goy & Felder, 1988 

Microprosthema semilaeve (Von Martens, 1872) 

Family Stenopodidae Huxley, 1878 

Genus Stenopus Latreille, 1819 

Stenopus hispidus (Olivier, 1811) 

Stenopus scutellatus Rankin, 1898 

Infraorder Caridea Dana, 1852 

Superfamily Pasiphaeoidea Dana, 1852 

Family Pasiphaeidae Dana, 1852 

Genus Leptochela Stimpson, 1860 

Leptochela carinata Ortmann, 1893 

Leptochela papulata Chace, 1976 

Leptochela serratorbita Bate, 1888 

Superfamily Atyoidea De Haan, 1849 

Family Atyidae De Haan, 1849 

Genus Potimirim Holthuis, 1954 

Potimirim potimirim (Müller, 1881) (Freshwater 

canals; introduced) 

Superfamily Bresilioidea Calman, 1896 

Family Bresiliidae Calman, 1896 

Genus Discias Rathbun, 1902 

Discias atlanticus Gurney, 1939 

Discias serratirostris Lebour, 1949 

Genus Pseudocheles Chace & Brown, 1978 

Pseudocheles chacei Kensley, 1983 

Superfamily Nematocarcinoidea S. I. Smith, 1884 

Family Rhynchocinetidae Ortmann, 1890 



Genus Cinetorhynchus Holthuis, 1995 

Cinetorhynchus manningi Okuno, 1996 

Cinetorhynchus rigens (Gordon, 1936) 

Superfamily Palaemonoidea Rafinesque, 1815 

Family Anchistioididae Borradaile, 1915 

Genus Anchistioides Paulson, 1875 

Anchistioides antiguensis (Schmitt, 1924) 

Family Gnathophyllidae Dana, 1852 

Genus Gnathophylloides Schmitt, 1933 

Gnathophylloides mineri Schmitt, 1933 

Genus Gnathophyllum Latreille, 1819 

Gnathophyllum americanum Guérin-Méneville, 

1855 

Gnathophyllum circellum Manning, 1963 

Gnathophyllum modestum Hay, 1917 

Family Palaemonidae Rafinesque, 1815 

Subfamily Palaemoninae Rafinesque, 1815 

Genus Brachycarpus Bate, 1888 

Brachycarpus biunguiculatus (Lucas, 1849) 

Genus Leander E. Desmarest, 1849 

Leander paulensis Ortmann, 1897 

Leander tenuicornis (Say, 1818) 

Genus Palaemon Weber, 1795 

Subgenus Palaeander Holthuis, 1950 

Palaemon (Palaeander) floridanus Chace, 1942 

Palaemon (Palaeander) northropi (Rankin, 1898) 

Genus Palaemonetes Heller, 1869 

Subgenus Palaemonetes Heller, 1869 

Palaemonetes (Palaemonetes) intermedius Holthuis, 

1949 

Palaemonetes (Palaemonetes) paludosus (Gibbes, 

1850) 

Palaemonetes (Palaemonetes) pugio Holthuis, 1949 

Palaemonetes (Palaemonetes) vulgaris Say, 1818 

Subfamily Pontoniinae Kingsley, 1878 

Genus Neopontonides Holthuis, 1951 

Neopontonides beaufortensis (Borradaile, 1920) 

Neopontonides chacei Heard, 1986 

Genus Periclimenaeus Borradaile, 1915 

Periclimenaeus ascidiarum Holthuis, 1951 

Periclimenaeus atlanticus (Rathbun, 1901) 

Periclimenaeus bermudensis (Armstrong, 1940) 

Periclimenaeus bredini Chace, 1972 

Periclimenaeus caraibicus Holthuis, 1951 

Periclimenaeus chacei Abele, 1971 

Periclimenaeus maxillulidens (Schmitt, 1936) 

Periclimenaeus pearsei (Schmitt, 1932) 

Periclimenaeus perlatus (Boone, 1930) 

Periclimenaeus schmitti Holthuis, 1951 

Periclimenaeus wilsoni (Hay, 1917) 

Genus Periclimenes Costa, 1844 

Subgenus Periclimenes Costa, 1844 

Periclimenes (Periclimenes) harringtoni Lebour, 

1949 

Periclimenes (Periclimenes) iridescens Lebour, 1949 

Periclimenes (Periclimenes) longicaudatus (Stimpson, 

1860) 

Periclimenes (Periclimenes) pandionis Holthuis, 

1951 

Periclimenes (Periclimenes) patae Heard & Spotte, 

1991 

Periclimenes (Periclimenes) pedersoni Chace, 1958 

Periclimenes (Periclimenes) perryae Chace, 1942 

Periclimenes (Periclimenes) yucatanicus (Ives, 1891) 

Subgenus Harpilius Dana, 1852 

Periclimenes (Harpilius) americanus (Kingsley, 

1878) 

Periclimenes (Harpilius) magnus Holthuis, 1951 

Periclimenes (Harpilius) rathbunae Schmitt, 1924 

Genus Pontonia Latreille, 1829 

Pontonia domestica Gibbes, 1850 

Pontonia margarita Smith, 1869 

Pontonia mexicana Guérin-Méneville, 1855 

Pontonia unidens Kingsley, 1880 

Genus Pontoniopsis Borradaile, 1915 

Pontoniopsis paulae Gore, 1981 

Genus Pseudocoutierea Holthuis, 1951 

Pseudocoutierea antillensis Chace, 1972 

Genus Tuleariocaris Hipeau-Jacquotte, 1965 

Tuleariocaris neglecta Chace, 1969 

Genus Typton Costa, 1844 

Typton carneus Holthuis, 1951 

Typton distinctus Chace, 1972 

Typton prionurus Holthuis, 1951 

Typton tortugae McClendon, 1911 

Genus Veleroniopsis Gore, 1981 

Veleroniopsis kimallynae Gore, 1981 



Superfamily Alpheoidea Rafinesque, 1815 

Family Alpheidae Rafinesque, 1815 

Genus Alpheopsis Coutière, 1896 

Alpheopsis labis Chace, 1972 

Alpheopsis trispinosus (Stimpson, 1861) 

Genus Alpheus Fabricius, 1798 

Alpheus amblyonyx Chace, 1972 

Alpheus angulatus McClure, 1995 

Alpheus armatus Rathbun, 1901 

Alpheus armillatus H. Milne Edwards, 1837 

Alpheus bahamensis Rankin, 1898 

Alpheus bouvieri A. Milne Edwards, 1878 

Alpheus candei Guérin-Méneville, 1855 

Alpheus cristulifrons Rathbun, 1900 

Alpheus cylindricus Kingsley, 1878 

Alpheus estuariensis Christoffersen, 1984 

Alpheus floridanus Kingsley, 1878 

Alpheus formosus Gibbes, 1850 

Alpheus heterochaelis Say, 1818 

Alpheus malleator Dana, 1852 

Alpheus normanni Kingsley, 1878 

Alpheus nuttingi (Schmitt, 1924) 

Alpheus paracrinitus Miers, 1881 

Alpheus peasei (Armstrong, 1940) 

Alpheus schmitti Chace, 1972 

Alpheus thomasi Hendrix & Gore, 1973 

Alpheus viridari (Armstrong, 1949) 

Alpheus websteri Kingsley, 1880 

Genus Automate De Man, 1888 

Automate dolichognatha De Man, 1888 

Automate evermanni Rathbun, 1901 

Automate rectifrons Chace, 1972 

Genus Fenneralpheus Felder & Manning, 1986 

Fenneralpheus chacei Felder & Manning, 1986 

Genus Leptalpheus Williams, 1965 

Leptalpheus forceps Williams, 1965 

Genus Metalpheus Coutière, 1908 

Metalpheus rostratipes (Pocock, 1890) 

Genus Salmoneus Holthuis, 1955 

Salmoneus cavicola Felder & Manning, 1986 

Genus Synalpheus Bate, 1888 

Synalpheus agelas L. Pequegnat & Heard, 1979 

Synalpheus apioceros Coutière, 1909 

Synalpheus bousfieldi Chace, 1972 

Synalpheus brevicarpus (Herrick, 1891) 

Synalpheus brooksi Coutière, 1909 

Synalpheus curacaoensis Schmitt, 1924 

Synalpheus fritzmuelleri Coutière, 1909 

Synalpheus goodei Coutière, 1909 

Synalpheus heardi Dardeau, 1984 

Synalpheus hemphilli Coutière, 1909 

Synalpheus herricki Coutière, 1909 

Synalpheus longicarpus (Herrick, 1891) 

Synalpheus mcclendoni Coutière, 1910 

Synalpheus minus (Say, 1818) 

Synalpheus pandionis Coutière, 1909 

Synalpheus paraneptunus Coutière, 1909 

Synalpheus pectiniger Coutière, 1907 

Synalpheus rathbunae Coutière, 1909 

Synalpheus sanctithomae Coutière, 1909 

Synalpheus scaphoceris Coutière, 1910 

Synalpheus townsendi Coutière, 1909 

Genus Thunor Armstrong, 1949 

Thunor simus (Guérin-Méneville, 1856) 

Family Hippolytidae Dana, 1852 

Genus Exhippolysmata Stebbing, 1815 

Exhippolysmata oplophoroides (Holthuis, 1948) 

Genus Hippolyte Leach, 1814 

Hippolyte coerulescens (Fabricius, 1775) 

Hippolyte nicholsoni Chace, 1972 

Hippolyte obliquimanus Dana, 1852 

(=Hippolyte curacaoensis Schmitt, 1924) 

Hippolyte pleuracanthus (Stimpson, 1871) 

Hippolyte zostericola (Smith, 1873) 

Genus Latreutes Stimpson, 1860 

Latreutes fucorum (Fabricius, 1798) 

Latreutes parvulus (Stimpson, 1866) 

Genus Lysmata Risso, 1816 

Lysmata amboinensis (De Man, 1888) 

Lysmata intermedia (Kingsley, 1878) 

Lysmata rathbunae Chace, 1970 

Lysmata wurdemanni (Gibbes, 1850) 

Genus Merhippolyte Bate, 1888 

Merhippolyte americana Holthuis, 1961 

Genus Thor Kingsley, 1878 

Thor amboinensis (De Man, 1888) 

Thor dobkini Chace, 1972 

Thor floridanus Kingsley, 1878 

Thor manningi Chace, 1972 

Genus Tozeuma Stimpson, 1860 

Tozeuma carolinense Kingsley, 1878 



Tozeuma cornutum A. Milne Edwards, 1881 

Tozeuma serratum A. Milne Edwards, 1881 

Genus Trachycaris Calman, 1906 

Trachycaris rugosa (Bate, 1888) 

Family Ogyrididae Hay & Shore, 1918 

Genus Ogyrides Stebbing, 1914 

Ogyrides alphaerostris (Kingsley, 1889) 

Ogyrides hayi Williams, 1981 

Superfamily Processoidea Ortmann, 1890 

Family Processidae Ortmann, 1896 

Genus Ambidexter Manning & Chace, 1971 

Ambidexter symmetricus Manning & Chace, 1971 

Genus Nikoides Paulson, 1875 

Nikoides schmitti Manning & Chace, 1971 

Genus Processa Leach, 1815 

Processa bermudensis (Rankin, 1900) 

Processa fimbriata Manning & Chace, 1971 

Processa hemphilli Manning & Chace, 1971 

Processa riveroi Manning & Chace, 1971 

Processa vicina Manning & Chace, 1971 

Processa vossi Manning, 1992 

Superfamily Crangonoidea Haworth, 1825 

Family Crangonidae Haworth, 1825 

Genus Philocheras Stebbing, 1900 

Philocheras gorei (Dardeau, 1980) 

Infraorder Astacidea Latreille, 1803 

Superfamily Enoplometopoidea 

de Saint Laurent, 1988 

Family Enoplometopidae de Saint Laurent, 1988 

Genus Enoplometopus H. Milne Edwards, 1862 

Enoplometopus antillensis Lütken, 1865 

Infraorder Thalassinidea Latreille, 1831 

Superfamily Callianassoidea Dana, 1852 

Family Callianassidae Dana, 1852 

Subfamily Callianassinae Dana, 1852 

Genus Biffarius Manning & Felder, 1991 

Biffarius biformis (Biffar, 1971) 

Biffarius fragilis (Biffar, 1970) 

Genus Gilvossius Manning & Felder, 1992 

Gilvossius setimanus (De Kay, 1844) 

(= Callianassa atlantica Rathbun, 1926) 

Subfamily Callichirinae Manning & Felder, 1991 

Genus Callichirus Stimpson, 1866 

Callichirus islagrande (Schmitt, 1935) 

Callichirus major (Say, 1818) 

Genus Corallianassa Manning & Felder, 1991 

Corallianassa longiventris (A. Milne Edwards, 

1870) 

Genus Glypturus Stimpson, 1866 

Glypturus acanthochirus Stimpson, 1866 

Genus Lepidopthalmus Holmes, 1904 

Lepidophthalmus louisianensis (Schmitt, 1935) 

Genus Neocallichirus Sakai, 1988 

Neocallichirus cacahuate Felder & Manning, 1995 

Neocallichirus grandimanus (Gibbes, 1850) 

(= Glypturus branneri Rathbun, 1900) 

Neocallichirus rathbunae (Schmitt, 1935) 

Genus Sergio Manning & Lemaitre, 1994 

Sergio mericeae Manning & Felder, 1995 

Sergio trilobatus (Biffar, 1970) 

Subfamily Cheraminae Manning & Felder, 1991 

Genus Cheramus Bate, 1888 

Cheramus marginatus (Rathbun, 1901) 

Subfamily Eucalliinae Manning & Felder, 1991 

Genus Eucalliax Manning & Felder, 1991 

Eucalliax quadracuta (Biffar, 1970) 

Eucalliax mcilhennyi Felder & Manning, 1994 

Callianassidae sp. A of Rakocinski et al., 1993 

Family Ctenochelidae Manning & Felder, 1991 

Subfamily Ctenochelinae 

Manning & Felder, 1991 

Genus Dawsonius Manning & Felder, 1991 

Dawsonius latispina (Dawson, 1967) 

Family Laomediidae Borradaile, 1903 

Genus Axianassa Schmitt, 1924 

Axianassa arenaria Kensley & Heard, 1990 

Family Upogebiidae Borradaile, 1903 

Genus Aethogebia Williams, 1993 



Aethogebia gorei Williams, 1993 

Genus Pomatogebia Williams & Ngoc-Ho, 1990 

Pomatogebia operculata (Schmitt, 1924) 

Genus Upogebia Leach, 1814 

Upogebia acanthura (Coêlho, 1973) 

Upogebia affinis (Say, 1818) 

Upogebia aquilina Williams, 1993 

Upogebia inomissa Williams, 1993 

Upogebia omissa Gomes Corrêa, 1968 

Upogebia spinistipula Williams & Heard, 1991 

Upogebia vasquezi Ngoc-Ho, 1989 

Superfamily Axioidea Huxley, 1879 

Family Axiidae Huxley, 1879 

Genus Acanthaxius Sakai & de Saint Laurent, 1989 

Acanthaxius hirsutimana (Boesch & Smalley, 1972) 

(provisionally transferred to Oxyrhynchaxius 

Parisi, 1917, by Kensley, 1996) 

Genus Axiorygma Kensley & Simmons, 1988 

Axiorygma nethertoni Kensley & Simmons, 1988 

Genus Axiopsis Borradaile, 1903 

Axiopsis serratifrons (A. Milne Edwards, 1873) 

Genus Coralaxius Kensley & Gore, 1981 

Coralaxius nodulosus (Meinert, 1877) 

Genus Paraxiopsis Kensley, 1996 

Paraxiopsis gracilimana Kensley, 1996 

Paraxiopsis spinipleura Kensley, 1996 

Axiidae sp. A of Rakocinski et al., 1993 

Family Micheleidae Sakai, 1992 

Genus Michelea Kensley & Heard, 1991 

Michelea vandoverae (Gore, 1987) 

Infraorder Palinura Latreille, 1903 

Superfamily Palinuroidea Latreille, 1803 

Family Palinuridae Latreille, 1803 

Genus Panulirus White, 1847 

Panulirus argus (Latreille, 1804) 

Panulirus guttatus (Latreille, 1804) 

Panulirus laevicauda (Latreille, 1817) 

Family Scyllaridae Latreille, 1825 

Subfamily Ibacinae Holthuis, 1985 

Genus Parribacus Dana, 1852 

Parribacus antarcticus (Lund, 1793) 

Subfamily Scyllarinae Latreille, 1825 

Genus Scyllarides Gill, 1898 

Scyllarides aequinoctialis (Lund, 1793) 

Scyllarides nodifer (Stimpson, 1866) 

Genus Scyllarus Fabricius, 1775 

Scyllarus americanus (Smith, 1869) 

Scyllarus chacei Holthuis, 1960 

Family Synaxidae Bate, 1881 

Genus Palinurellus Von Martens, 1878 

Palinurellus gundlachi Von Martens, 1878 

Infraorder Anomura H. Milne Edwards, 1832 

Superfamily Coenobitoidea Dana, 1851 

Family Coenobitidae Dana, 1851 

Genus Coenobita Latreille, 1826 

Coenobita clypeatus (Herbst, 1791) 

Family Diogenidae Ortmann, 1892 

Genus Calcinus Dana, 1851 

Calcinus tibicen (Herbst, 1791) 

Genus Cancellus H. Milne Edwards, 1836 

Cancellus viridis Mayo, 1973 

Genus Clibanarius Dana, 1851 

Clibanarius antillensis Stimpson, 1862 

Clibanarius cubensis (Saussure, 1858) 

Clibanarius tricolor (Gibbes, 1850) 

Clibanarius vittatus (Bosc, 1802) 

Genus Dardanus Paulson, 1875 

Dardanus fucosus Biffar & Provenzano, 1972 

Dardanus insignis (Saussure, 1858) 

Dardanus venosus (H. Milne Edwards, 1848) 

Genus Isocheles Stimpson, 1859 

Isocheles wurdemanni Stimpson, 1862 

Genus Paguristes Dana, 1852 

Paguristes anomalus Bouvier, 1918 

Paguristes cadenati Forest, 1954 

Paguristes erythrops Holthuis, 1959 

Paguristes grayi Benedict, 1901 

Paguristes hummi Wass, 1955 



Paguristes invisisacculus McLaughlin & Provenzano, 

1974 

Paguristes lymani A. Milne Edwards & Bouvier, 

1893 

Paguristes oxyophthalmus Holthuis, 1959 

Paguristes puncticeps Benedict, 1901 

Paguristes sericeus A. Milne Edwards, 1880 

Paguristes starcki Provenzano, 1965 

Paguristes tortugae Schmitt, 1933 

Paguristes triangulatus A. Milne Edwards & Bouvier, 

1893 

Paguristes wassi Provenzano, 1961 

Genus Petrochirus Stimpson, 1859 

Petrochirus diogenes (Linnaeus, 1758) 

Superfamily Paguroidea Latreille, 1803 

Family Paguridae Latreille, 1803 

Genus Anisopagurus McLaughlin, 1981 

Anisopagurus pygmaeus (Bouvier, 1918) 

Genus Goreopagurus McLaughlin, 1988 

Goreopagurus piercei (Wass, 1963) 

Genus Iridopagurus de Saint Laurent-Dechancé, 1966 

Iridopagurus caribbensis (A. Milne Edwards & 

Bouvier, 1893) 

Iridopagurus reticulatus García-Gómez, 1983 

Genus Manucomplanus McLaughlin, 1981 

Manucomplanus spinulosus (Holthuis, 1959) 

Manucomplanus ungulatus (Studer, 1883) 

Genus Pagurus Fabricius, 1775 

Pagurus annulipes (Stimpson, 1860) 

Pagurus brevidactylus (Stimpson, 1859) 

Pagurus carolinensis McLaughlin, 1975 

Pagurus criniticornis (Dana, 1852) 

Pagurus defensus (Benedict, 1892) 

Pagurus gymnodactylus Lemaitre, 1982 

Pagurus impressus (Benedict, 1892) 

Pagurus longicarpus Say, 1817 

Pagurus mclaughlinae García-Gómez, 1982 

Pagurus marshi Benedict, 1901 

Pagurus pollicaris Say, 1817 

Pagurus provenzanoi Forest & de Saint Laurent, 

1967 

Pagurus stimpsoni (A. Milne Edwards & Bouvier, 

1893) 

Genus Phimochirus McLaughlin, 1981 

Phimochirus holthuisi (Provenzano, 1961) 

Phimochirus operculatus (Stimpson, 1859) 

Phimochirus randalli (Provenzano, 1961) 

Superfamily Galatheoidea Samouelle, 1819 

Family Galatheidae Samouelle, 1819 

Genus Galathea Fabricius, 1793 

Galathea rostrata A. Milne Edwards, 1880 

Genus Munida Leach, 1820 

Munida irrasa A. Milne Edwards, 1880 

Munida pusilla Benedict, 1902 

Family Porcellanidae Haworth, 1825 

Genus Euceramus Stimpson, 1860 

Euceramus praelongus Stimpson, 1860 

Genus Megalobrachium Stimpson, 1858 

Megalobrachium poeyi (Guérin-Méneville, 1855) 

Megalobrachium soriatum (Say, 1818) 

Genus Neopisosoma Haig, 1960 

Neopisosoma angustifrons (Benedict, 1901) 

Genus Pachycheles Stimpson, 1858 

Pachycheles ackleianus A. Milne Edwards, 1880 

Pachycheles monilifer (Dana, 1852) 

Pachycheles pilosus (H. Milne Edwards, 1837) 

Pachycheles riisei (Stimpson, 1858) 

Pachycheles rugimanus A. Milne Edwards, 1880 

Genus Parapetrolisthes Haig, 1962 

Parapetrolisthes tortugensis (Glassell, 1945) 

Genus Petrolisthes Stimpson, 1858 

Petrolisthes armatus (Gibbes, 1850) 

Petrolisthes galathinus (Bosc, 1802) 

Petrolisthes jugosus Streets, 1872 

Petrolisthes politus (Gray, 1831) 

Genus Polyonyx Stimpson, 1858 

Polyonyx gibbesi Haig, 1956 

Genus Porcellana Lamarck, 1801 

Porcellana sayana (Leach, 1820) 

Porcellana sigsbeiana A. Milne Edwards, 1880 

Porcellana stimpsoni A. Milne Edwards, 1880 

Superfamily Hippoidea Latreille, 1825 

Family Albuneidae Stimpson, 1858 

Genus Albunea Weber, 1795 

Albunea gibbesii Stimpson, 1859 

Albunea paretii Guérin-Méneville, 1853 



Genus Lepidopa Stimpson, 1858 

Lepidopa benedicti Schmitt, 1935 

Lepidopa websteri Benedict, 1903 

Genus Zygopa Holthuis, 1960 

Zygopa michaelis Holthuis, 1960 

Family Hippidae Latreille, 1825 

Genus Emerita Scopoli, 1777 

Emerita benedicti Schmitt, 1935 

Emerita portoricensis Schmitt, 1935 

Emerita talpoida (Say, 1817) 

Genus Hippa Fabricius, 1787 

Hippa testudinaria (Herbst, 1791) 

(= Hippa cubensis (Saussure, 1857)) 

Superfamily Dromioidea De Haan, 1833 

Family Dromiidae De Haan, 1833 

Genus Dromia Weber, 1795 

Dromia erythropus (George Edwards, 1771) 

Genus Cryptodromiopsis Borradaile, 1903 

Cryptodromiopsis antillensis (Stimpson, 1858) 

Genus Hypoconcha Guérin-Méneville, 1854 

Hypoconcha arcuata Stimpson, 1858 

Hypoconcha parasitica (Linnaeus, 1763) 

(= H. sabulosa (Herbst, 1799)) 

Hypoconcha spinosissima Rathbun, 1933 

Infraorder Brachyura Latreille, 1803 

Section Archaeobrachyura Guinot, 1977 

Superfamily Homoloidea De Haan, 1839 

Family Cyclodorippidae Ortmann, 1892 

Genus Clythrocerus A. Milne Edwards & Bouvier, 

1899 

Clythrocerus nitidus (A. Milne Edwards, 1880) 

Superfamily Raninoidea De Haan, 1839 

Family Raninidae De Haan, 1839 

Subfamily Ranininae De Haan, 1839 

Genus Raninoides H. Milne Edwards, 1837 

Raninoides loevis (Latreille, 1825) 

Subfamily Notopodinae Serène & Umali, 1972 

Genus Ranilia H. Milne Edwards, 1837 

Ranilia constricta (A. Milne Edwards, 1880) 

Ranilia muricata H. Milne Edwards, 1837 

Subfamily Symethinae Goeke, 1981 

Genus Symethis Weber, 1795 

Symethis variolosa (Fabricius, 1793) 

Section Oxystomata H. Milne Edwards, 1834 

Superfamily Dorippoidea MacLeay, 1838 

Family Dorippidae MacLeay, 1838 

Subfamily Ethusinae Guinot, 1977 

Genus Ethusa Roux, 1828 

Ethusa americana A. Milne Edwards, 1880 

Superfamily Leucosioidea Samouelle, 1819 

Family Calappidae De Haan, 1833 

Subfamily Calappinae De Haan, 1833 

Genus Calappa Weber, 1795 

Calappa flammea (Herbst, 1794) 

Calappa galloides Stimpson, 1859 

Calappa ocellata Holthuis, 1958 

Calappa sulcata Rathbun, 1898 

Calappa tortugae Rathbun, 1933 

Genus Cryptosoma Brullé, 1837 

Cryptosoma bairdii (Stimpson, 1860) 

Genus Cyclozodion Williams & Child, 1988 

Cyclozodion tuberatum Williams & Child, 1988 

Subfamily Matutinae De Haan, 1835 

Genus Hepatus Latreille, 1802 

Hepatus epheliticus (Linnaeus, 1763) 

Hepatus pudibundus (Herbst, 1785) 

Genus Osachila Stimpson, 1871 

Osachila semilevis Rathbun, 1916 

Family Leucosiidae Samouelle, 1819 

Subfamily Ebaliinae Stimpson, 1871 

Genus Ebalia Leach, 1817 

Ebalia cariosa (Stimpson, 1860) 

Ebalia stimpsonii A. Milne Edwards, 1880 

Genus Speloeophorus A. Milne Edwards, 1865 

Speloeophorus elevatus Rathbun, 1898 

Speloeophorus nodosus (Bell, 1855) 

Speloeophorus pontifer (Stimpson, 1871) 

Genus Uhlias Stimpson, 1871 

Uhlias limbatus Stimpson, 1871 



Subfamily Iliinae Stimpson, 1871 

Genus Callidactylus Stimpson, 1871 

Callidactylus asper Stimpson, 1871 

Genus Iliacantha Stimpson, 1871 

Iliacantha intermedia Miers, 1886 

Iliacantha liodactylus Rathbun, 1898 

Iliacantha sparsa Stimpson, 1871 

Iliacantha subglobosa Stimpson, 1871 

Genus Persephona Leach, 1817 

Persephona crinita Rathbun, 1931 

Persephona mediterranea (Herbst, 1794) 

Section Oxyrhyncha Latreille, 1803 

Superfamily Majoidea Samouelle, 1819 

Family Majidae Samouelle, 1819 

Subfamily Inachinae MacLeay, 1838 

Genus Aepinus Rathbun, 1897 

Aepinus septemspinosus (A. Milne Edwards, 1879) 

Genus Batrachonotus Stimpson, 1871 

Batrachonotus fragosus Stimpson, 1871 

Genus Inachoides A. Milne Edwards & Lucas, 1843 

Inachoides forceps A. Milne Edwards, 1879 

Genus Metoporhaphis Stimpson, 1860 

Metoporhaphis calcarata (Say, 1818) 

Genus Podochela Stimpson, 1860 

Podochela gracilipes Stimpson, 1871 

Podochela macrodera Stimpson, 1860 

Podochela riisei Stimpson, 1860 

Podochela sidneyi Rathbun, 1924 

Genus Pyromaia Stimpson, 1871 

Pyromaia cuspidata Stimpson, 1871 

Genus Stenorhynchus Lamarck, 1818 

Stenorhynchus seticornis (Herbst, 1788) 

Subfamily Epialtinae MacLeay, 1838 

Genus Acanthonyx Latreille, 1825 

Acanthonyx petiverii H. Milne Edwards, 1834 

Genus Epialtus H. Milne Edwards, 1834 

Epialtus bituberculatus H. Milne Edwards, 1834 

Epialtus dilatatus A. Milne Edwards, 1878 

Epialtus dilatatus forma elongata Rathbun, 1923 

Epialtus kingsleyi Rathbun, 1923 

Epialtus longirostris Stimpson, 1860 

Genus Mocosoa Stimpson, 1871 

Mocosoa crebripunctata Stimpson, 1871 

Subfamily Pisinae Dana, 1851 

Genus Chorinus Latreille, 1825 

Chorinus heros (Herbst, 1790) 

Genus Libinia Leach, 1815 

Libinia dubia H. Milne Edwards, 1834 

Libinia emarginata Leach, 1815 

Libinia erinacea (A. Milne Edwards, 1879) 

Genus Pelia Bell, 1836 

Pelia mutica (Gibbes, 1850) 

Subfamily Majinae Samouelle, 1819 

Genus Coelocerus A. Milne Edwards, 1875 

Coelocerus spinosus A. Milne Edwards, 1875 

Genus Hemus A. Milne Edwards, 1875 

Hemus cristulipes A. Milne Edwards, 1875 

Genus Macrocoeloma Miers, 1879 

Macrocoeloma camptocerum (Stimpson, 1871) 

Macrocoeloma diplacanthum (Stimpson, 1860) 

Macrocoeloma eutheca (Stimpson, 1871) 

Macrocoeloma laevigatum (Stimpson, 1860) 

Macrocoeloma septemspinosum (Stimpson, 1871) 

Macrocoeloma subparallelum (Stimpson, 1860) 

Macrocoeloma trispinosum trispinosum (Latreille, 

1825) 

Macrocoeloma trispinosum nodipes (Desbonne, 

1867) 

Macrocoeloma trispinosum variety of Rathbun, 

1925 

Genus Microphrys H. Milne Edwards, 1857 

Microphrys antillensis Rathbun, 1920 

Microphrys bicornutus (Latreille, 1825) 

Genus Mithrax Desmarest, 1823 

Mithrax acuticornis Stimpson, 1871 

Mithrax caribbaeus Rathbun, 1900 

Mithrax cornutus Saussure, 1857 

Mithrax hemphilli Rathbun, 1892 

Mithrax hispidus (Herbst, 1790) 

Mithrax holderi Stimpson, 1871 

Mithrax pilosus Rathbun, 1892 

Mithrax pleuracanthus Stimpson, 1871 

Mithrax spinosissimus (Lamarck, 1818) 



Mithrax tortugae Rathbun, 1920 

Mithrax verrucosus H. Milne Edwards, 1832 

Genus Mithraculus White, 1847 

Mithraculus cinctimanus Stimpson, 1860 

Mithraculus coryphe (Herbst, 1801) 

Mithraculus forceps A. Milne Edwards, 1875 

Mithraculus ruber Stimpson, 1871 

Mithraculus sculptus (Lamarck, 1818) 

Genus Picroceroides Miers, 1886 

Picroceroides tubularis Miers, 1886 

Genus Pitho Bell, 1835 

Pitho aculeata (Gibbes, 1850) 

Pitho anisodon (Von Martens, 1872) 

Pitho laevigata (A. Milne Edwards, 1875) 

Pitho lherminieri (Schramm, 1867) 

Pitho mirabilis (Herbst, 1794) 

Pitho quadridentata (Miers, 1879) 

Genus Stenocionops Desmarest, 1823 

Stenocionops furcata coelata (A. Milne Edwards, 

1878) 

Stenocionops furcata furcata (Olivier, 1791) 

Genus Thoe Bell, 1836 

Thoe puella Stimpson, 1860 

Subfamily Tychinae Dana, 1851 

Genus Tyche Bell, 1835 

Tyche emarginata White, 1847 

Superfamily Parthenopoidea MacLeay, 1838 

Family Parthenopidae MacLeay, 1838 

Subfamily Parthenopinae MacLeay, 1838 

Genus Cryptopodia H. Milne Edwards, 1834 

Cryptopodia concava Stimpson, 1871 

Genus Heterocrypta Stimpson, 1871 

Heterocrypta granulata (Gibbes, 1850) 

Genus Mesorhoea Stimpson, 1871 

Mesorhoea sexspinosa Stimpson, 1871 

Genus Parthenope Weber, 1795 

Subgenus Platylambrus Stimpson, 1871 

Parthenope (Platylambrus) granulata (Kingsley, 

1879) 

Parthenope (Platylambrus) serrata (H. Milne Edwards, 

1834) 

Section Brachyrhyncha Borradaile, 1907 

Superfamily Portunoidea Rafinesque, 1815 

Family Portunidae Rafinesque, 1815 

Subfamily Polybiinae Ortmann, 1893 

Genus Ovalipes Rathbun, 1898 

Ovalipes floridanus Hay & Shore, 1918 

Ovalipes stephensoni Williams, 1976 

Subfamily Portuninae Rafinesque, 1815 

Genus Arenaeus Dana, 1852 

Arenaeus cribrarius (Lamarck, 1818) 

Genus Callinectes Stimpson, 1860 

Callinectes bocourti A. Milne Edwards, 1879 

Callinectes danae Smith, 1869 

Callinectes exasperatus (Gerstaecker, 1856) 

Callinectes larvatus Ordway, 1863 

Callinectes ornatus Ordway, 1863 

Callinectes sapidus Rathbun, 1896 

Callinectes similis Williams, 1966 

Genus Charybdis de Haan, 1835 

Charybdis helleri A. Milne Edwards, 1867 (Extralimital; 

introduced) 

Genus Cronius Stimpson, 1860 

Cronius ruber (Lamarck, 1818) 

Cronius tumidulus (Stimpson, 1871) 

Genus Laleonectes Manning & Chace, 1990 

Laleonectes vocans (A. Milne Edwards, 1878) 

Genus Portunus Weber, 1795 

Portunus anceps (Saussure, 1858) 

Portunus depressifrons (Stimpson, 1859) 

Portunus gibbesii (Stimpson, 1859) 

Portunus sayi (Gibbes, 1850) (usually pelagic in 

Sargassum) 

Portunus sebae (H. Milne Edwards, 1834) 

Portunus spinicarpus (Stimpson, 1871) 

Portunus spinimanus Latreille, 1819 

Portunus ventralis (A. Milne Edwards, 1879) 

Portunus vossi Lemaitre, 1992 

Superfamily Xanthoidea MacLeay, 1838 

Family Goneplacidae MacLeay, 1838 

(subfamilies omitted) 

Genus Eucratopsis Smith, 1869 

Eucratopsis crassimanus (Dana, 1852) 

Genus Euryplax Stimpson, 1859 



Euryplax nitida Stimpson, 1859 

Genus Glyptoplax Smith, 1870 

Glyptoplax smithii A. Milne Edwards, 1880 

Genus Panoplax Stimpson, 1871 

Panoplax depressa Stimpson, 1871 

Genus Pilumnoplax Stimpson, 1858 

Pilumnoplax elata (A. Milne Edwards, 1880) 

Genus Prionoplax A. Milne Edwards, 1852 

Prionoplax atlantica Kendall, 1891 

Genus Pseudorhombila H. Milne Edwards, 1837 

Pseudorhombila quadridentata (Latreille, 1828) 

Genus Speocarcinus Stimpson, 1859 

Speocarcinus carolinensis Stimpson, 1859 

Speocarcinus lobatus Guinot, 1969 

Genus Trapezioplax Guinot, 1969 

Trapezioplax tridentata (A. Milne Edwards, 1880) 

Family Xanthidae MacLeay, 1838 

(subfamilies omitted) 

Genus Actaea De Haan, 1833 

Actaea acantha (H. Milne Edwards, 1834) 

Actaea bifrons Rathbun, 1898 

Genus Banareia A. Milne Edwards, 1869 

Banareia palmeri (Rathbun, 1894) 

Genus Carpilius Leach, 1823 

Carpilius corallinus (Herbst, 1783) 

Genus Carpoporus Stimpson, 1871 

Carpoporus papulosus Stimpson, 1871 

Genus Cataleptodius Guinot, 1968 

Cataleptodius floridanus (Gibbes, 1850) 

Genus Chlorodiella Rathbun, 1897 

Chlorodiella longimana (H. Milne Edwards, 1834) 

Genus Domecia Eydoux & Souleyet, 1842 

Domecia acanthophora acanthophora (Desbonne & 

Schramm, 1867) 

Genus Dyspanopeus Martin & Abele, 1986 

Dyspanopeus sayi (Smith, 1869) 

Dyspanopeus texana (Stimpson, 1859) 

Genus Eriphia Latreille, 1817 

Eriphia gonagra (Fabricius, 1781) 

Genus Etisus H. Milne Edwards, 1834 

Etisus maculatus (Stimpson, 1860) 

Genus Eurypanopeus A. Milne Edwards, 1880 

Eurypanopeus abbreviatus abbreviatus (Stimpson, 

1860) 

Eurypanopeus depressus (Smith, 1869) 

Eurypanopeus dissimilis (Benedict & Rathbun, 

1891) 

Eurypanopeus turgidus (Rathbun, 1930) 

Genus Eurytium Stimpson, 1859 

Eurytium limosum (Say, 1818) 

Genus Garthiope Guinot, 1990 

Garthiope barbadensis (Rathbun, 1921) 

Garthiope spinipes (A. Milne Edwards, 1880) 

Genus Glyptoxanthus A. Milne Edwards, 1879 

Glyptoxanthus erosus (Stimpson, 1859) 

Genus Heteractaea Lockington, 1877 

Heteractaea ceratopus (Stimpson, 1860) 

Genus Hexapanopeus Rathbun, 1898 

Hexapanopeus angustifrons (Benedict & Rathbun, 

1891) 

Hexapanopeus caribbaeus (Stimpson, 1871) 

Hexapanopeus hemphillii (Benedict & Rathbun, 

1891) 

Hexapanopeus paulensis Rathbun, 1930 

Hexapanopeus quinquedentatus Rathbun, 1901 

Genus Leptodius A. Milne Edwards, 1863 

Leptodius parvulus (Fabricius, 1793) 

Genus Lobopilumnus A. Milne Edwards, 1880 

Lobopilumnus agassizii (Stimpson, 1871) 

Genus Melybia Stimpson, 1871 

Melybia thalamita Stimpson, 1871 

Genus Menippe De Haan, 1833 

Menippe adina Felder & Williams, 1986 

Menippe mercenaria (Say, 1818) 

Menippe nodifrons Stimpson, 1859 

Genus Micropanope Stimpson, 1871 

Micropanope nuttingi (Rathbun, 1898) 

Micropanope pusilla A. Milne Edwards, 1880 

Micropanope sculptipes Stimpson, 1871 



Genus Neopanope A. Milne Edwards, 1880 

Neopanope packardii (Kingsley, 1879) 

Genus Panopeus H. Milne Edwards, 1834 

Panopeus americanus Saussure, 1857 

Panopeus bermudensis Benedict & Rathbun, 1891 

Panopeus hartii Smith, 1869 

Panopeus herbstii H. Milne Edwards, 1834 

Panopeus lacustris Desbonne, 1867 

Panopeus obesus Smith, 1869 

Panopeus occidentalis Saussure, 1857 

Panopeus rugosus A. Milne Edwards, 1880 

Panopeus simpsoni Rathbun, 1930 

Genus Paractaea Guinot, 1969 

Paractaea rufopunctata nodosa (Stimpson, 1860) 

Genus Paraliomera Rathbun, 1930 

Paraliomera dispar (Stimpson, 1871) 

Paraliomera longimana (A. Milne Edwards, 1865) 

Genus Pilumnus Leach, 1815 

Pilumnus caribaeus Desbonne & Schramm, 1867 

Pilumnus dasypodus Kingsley, 1879 

Pilumnus floridanus Stimpson, 1871 

Pilumnus gemmatus Stimpson, 1860 

Pilumnus holosericus Rathbun, 1898 

Pilumnus lacteus Stimpson, 1871 

Pilumnus longleyi Rathbun, 1930 

Pilumnus marshi Rathbun, 1901 

Pilumnus nudimanus Rathbun, 1900 

Pilumnus pannosus Rathbun, 1896 

Pilumnus sayi Rathbun, 1897 

Pilumnus spinosissimus Rathbun, 1898 

Genus Platyactaea Guinot, 1967 

Platyactaea setigera (H. Milne Edwards, 1834) 

Genus Platypodiella Guinot, 1967 

Platypodiella spectabilis (Herbst, 1794) 

Genus Pseudomedaeus Guinot, 1967 

Pseudomedaeus agassizii (A. Milne Edwards, 1880) 

Genus Rhithropanopeus Rathbun, 1898 

Rhithropanopeus harrisii (Gould, 1841) 

Genus Tetraxanthus Rathbun, 1898 

Tetraxanthus rathbunae Chace, 1939 

Genus Xanthodius Stimpson, 1859 

Xanthodius denticulatus (White, 1847) 

Superfamily Grapsidoidea MacLeay, 1838 

Family Gecarcinidae MacLeay, 1838 

Genus Cardisoma Latreille, 1825 

Cardisoma guanhumi Latreille, 1825 

Genus Gecarcinus Leach, 1814 

Gecarcinus lateralis (Freminville, 1835) 

Gecarcinus ruricola (Linnaeus, 1758) 

Family Grapsidae MacLeay, 1838 

Subfamily Grapsinae MacLeay, 1838 

Genus Geograpsus Stimpson, 1858 

Geograpsus lividus (H. Milne Edwards, 1837) 

Genus Goniopsis De Haan, 1833 

Goniopsis cruentata (Latreille, 1803) 

Genus Grapsus Lamarck, 1801 

Grapsus grapsus (Linnaeus, 1758) 

Genus Pachygrapsus Randall, 1840 

Pachygrapsus gracilis (Saussure, 1858) 

Pachygrapsus transversus (Gibbes, 1850) 

Genus Planes Bowdich, 1825 

Planes minutus (Linnaeus, 1758) 

Subfamily Plagusiinae Dana, 1851 

Genus Percnon Gistel, 1848 

Percnon gibbesi (H. Milne Edwards, 1853) 

Genus Plagusia Latreille, 1806 

Plagusia depressa (Fabricius, 1775) 

Subfamily Sesarminae Dana, 1851 

Genus Aratus H. Milne Edwards, 1853 

Aratus pisonii (H. Milne Edwards, 1837) 

Genus Armases Abele, 1992 

Armases benedicti (Rathbun, 1897) 

Armases cinereum (Bosc, 1802) 

Armases miersii (Rathbun, 1897) 

Armases ricordi (H. Milne Edwards, 1853) 

Genus Cyclograpsus H. Milne Edwards, 1837 

Cyclograpsus integer H. Milne Edwards, 1837 

Genus Sesarma Say, 1817 

Sesarma curacaoense De Man, 1892 

Sesarma reticulatum (Say, 1817) species complex 



Subfamily Varuninae H. Milne Edwards, 1852 

Genus Platychirograpsus De Man, 1896 

Platychirograpsus spectabilis De Man, 1896 (Extralimital—introduced) 

Superfamily Pinnotheroidea De Haan, 1833 

Family Pinnotheridae De Haan, 1833 

Subfamily Pinnotherinae De Haan, 1833 

Genus Clypeasterophilus Campos & Griffith, 1990 

Clypeasterophilus juvenilis (Bouvier, 1917) 

Clypeasterophilus rugatus (Bouvier, 1917) 

Clypeasterophilus stebbingi (Rathbun, 1918) 

Genus Dissodactylus Smith, 1870 

Dissodactylus crinitichelis Moreira, 1901 

Dissodactylus latus Griffith, 1987 

Dissodactylus mellitae (Rathbun, 1900) 

Dissodactylus primitivus Bouvier, 1917 

Genus Fabia Dana, 1851 

Fabia byssomiae (Say, 1818) 

Fabia felderi Gore, 1986 

Fabia tellinae Cobb, 1973 

Genus Orthotheres Sakai, 1969 

Orthotheres strombi (Rathbun, 1905) 

Genus Parapinnixa Holmes, 1894 

Parapinnixa bouvieri Rathbun, 1918 

Parapinnixa hendersoni Rathbun, 1918 

Genus Pinnaxodes Heller, 1865 

Pinnaxodes floridensis Wells & Wells, 1961 

Genus Pinnotheres Bosc 1802 

Pinnotheres hemphilli Rathbun, 1918 

Pinnotheres shoemakeri Rathbun, 1918 

Genus Tumidotheres Campos, 1989 

Tumidotheres maculatus (Say, 1818) 

Genus Tunicotheres Campos, 1996 

Tunicotheres moseri (Rathbun, 1918) 

Genus Zaops Rathbun, 1900 

Zaops ostreum (Say, 1817) 

Subfamily Pinnothereliinae Alcock, 1900 

Genus Austinixa Heard & Manning, 1997 

Austinixa chacei (Wass, 1955) 

Austinixa cristata (Rathbun, 1900) 

Austinixa gorei (Manning & Felder, 1989) 

Genus Pinnixa White, 1846 

Pinnixa chaetopterana Stimpson, 1860 

Pinnixa cylindrica (Say, 1818) 

Pinnixa floridana Rathbun, 1918 

Pinnixa leptosynaptae Wass, 1968 

Pinnixa lunzi Glassell, 1937 

Pinnixa pearsei Wass, 1955 

Pinnixa retinens Rathbun, 1918 

Pinnixa sayana Stimpson, 1860 

Pinnixa sp. A of Rakocinski et al., 1993 

Pinnixa sp. E of Rakocinski et al., 1993 

Superfamily Ocypodoidea Rafinesque, 1815 

Family Ocypodidae Rafinesque, 1815 

Subfamily Ocypodinae Rafinesque, 1815 

Genus Ocypode Weber, 1795 

Ocypode quadrata (Fabricius, 1787) 

Genus Uca Leach, 1814 

Uca burgersi Holthuis, 1967 

Uca leptodactyla Rathbun, 1898 

Uca longisignalis Salmon & Atsaides, 1968 

Uca minax (Le Conte, 1855) 

Uca panacea Novak & Salmon, 1974 

Uca pugilator (Bosc, 1802) 

Uca pugnax (Smith, 1870) 

Uca rapax (Smith, 1870) 

Uca speciosa (Ives, 1891) 

Uca spinicarpa Rathbun, 1900 

Uca thayeri Rathbun, 1900 

Uca vocator (Herbst, 1804) 

Genus Ucides Rathbun, 1897 

Ucides cordatus (Linnaeus, 1763) 

Family Palicidae Rathbun, 1898 

Genus Palicus Philippi, 1838 

Palicus affinis A. Milne Edwards & Bouvier, 1899 

Palicus alternatus Rathbun, 1897 

Palicus dentatus A. Milne Edwards, 1880 

Palicus obesus (A. Milne Edwards, 1880) 

Palicus sica (A. Milne Edwards, 1880) 

Superfamily Cryptochiroidea Paulson, 1875 

Family Cryptochiridae Paulson, 1875 

Genus Opecarcinus Kropp & Manning, 1987 

Opecarcinus hypostegus (Shaw & Hopkins, 1977) 

Genus Troglocarcinus Verrill, 1908 

Troglocarcinus corallicola Verrill, 1908 



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GORDON, I. 1936. On the Macruran genus Rhynchocinetes, 

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GORE, R. H. 1971.The complete larval development of 

Porcellana sigsbeiana (Crustacea: Decapoda) under laboratory 

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GORE, R. H. 1972. Zoogeographical aspects of decapod 

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GORE, R. H. 1981. Three new shrimps and some interesting 

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GORE, R. H. 1987. Callianidea vandoverae new species 

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Smithsonian Contributions to Zoology 

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GORE, R. H., E. E. GALLAHER, L. E. SCOTTO, and K. 

A.WILSON. 1981. Community composition, structure, 

biomass and species-area relationships of seagrass 

and drift algae associated macrocrustaceans. Estuarine, 

Coastal and Shelf Science 12(4): 485–508. 

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(Decapoda, Portunidae) from the central east coast of 

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GORE, R. H., G. R. KULCZYCKI, and P. A. HASTINGS. 

1978. A second occurrence of the Brazilian freshwater 

shrimp, Potimirim potimirim, along the central eastern 

Florida coast. Florida Scientist 41(1): 57–61. 

GORE, R. H., and L. E. SCOTTO. 1979. Crabs of the family 

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(Anomura: Paguridae): The complete larval development 

under laboratory conditions, and the systematic 

relationships of its larvae. Journal of Crustacean Biology 

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GORE, R. H., L. E. SCOTTO, and W.T.YANG. 1982. Microphrys 

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notes on other mithracine larvae (Decapoda: Brachyura: 

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GORE, R. H., and C. L.VAN DOVER. 1980. Studies on 

decapod Crustacea from the Indian River region of 

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Lepidopa richmondi, with notes on larvae of American 

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1981. Studies on decapod Crustacea from the Indian 

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GORE, R. H., C. L. VAN DOVER, and K. A. WILSON. 

1981. Micropanope barbadensis, the complete larval de- 



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JR. 1981. Larval development of the spider crab Mithrax 

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Methods 

Checklist of Shallow-Water Echinodermata of Florida 

Names on the initial draft of this list were extracted 

from Echinoderms of Florida and the Caribbean (Hendler 

et al., 1995) and then supplemented or confirmed by information 

from many other published sources, which 

are listed in the accompanying compendium of pertinent 

references. Some of the more comprehensive references 

consulted during preparation of the list are the 

following, listed in alphabetical order: A. M. Clark and 

Downey (1992), Downey (1973), Durham et al. (1966), 

Frizzell et al. (1966), Harold and Telford (1990), Hendler 

(1995), Hopkins (1988), Kier and Grant (1965), Messing 

(1978), Miller and Pawson (1984), Ruppert and Fox 

(1988), Serafy (1979), Spencer and Wright (1966),Thomas 

(1962), Thomas (1973), and Tortonese and Downey 

(1977). Many additional papers on the systematics of 

Florida taxa were consulted, most of which dealt with 

only one or a few species; included in this group are 

several papers each by R. H. Chesher, A. H. Clark, E. 

Deichmann, D. M. Devaney, M. E. Downey, G. Hendler, 

P. M. Kier, J. E. Miller, D. L. Pawson, D. K. Serafy, R. L. 

Singletary, L. P. Thomas, and R. L. Turner, as well as a 

few others included in the list of pertinent references. 

In several instances, the database of the Florida Marine 

Research Institute Marine Specimen Collection 

was searched for depth records to indicate whether a 

species should be listed.The draft list of names was then 

sent to three reviewers (see Acknowledgments), one of 

whom suggested several additions to and deletions 

from the list and all of whom helped by filling in missing 

information on the authorities (authors’ names 

and dates of introduction) for some of the higher taxa. 

This list of shallow-water taxa includes all echinoderms 

known to occur in Florida’s estuaries and 

coastal waters offshore to depths of about 37 m (120 feet; 

20 fathoms). A few of the listed species are principally 

deeper-water organisms whose ranges extend only 

slightly into shallow coastal waters. The ranges of 

species in this group, which includes the echinoids 


Genocidaris maculata and Stylocidaris affinis, the ophiuroid 

Astroporpa annulata, and perhaps a few others, 

begin at about 30-35 m and extend offshore to much 

greater depths on the continental shelf.Those species 

are included here for completeness, but it is unlikely 

that most of them will be found in shallower depths. 

The ophiuroid Amphioplus abditus is known to occur 

from Maine southward to Sapelo Island, Georgia, typically 

in shallow water but also out to depths of 40 m 

(Hendler, 1995). The species is listed here on the suspicion 

that any species found at Sapelo Island, Georgia, 

may eventually be found in coastal waters of extreme 

northeastern Florida. 

The classification of taxa follows that used in the 

most recently published revisions, but with one exception. 

The ophiuroid Amphipholis atra (Stimpson, 

1865) has customarily been classified in the genus Amphiodia 

Verrill, 1899 (e.g., Hendler et al., 1995) but is 

listed here in Amphipholis Ljungman, 1867, following 

the advice of Dr. R. L.Turner. 


Draft lists of echinoderms were reviewed by Dr. David 

L. Pawson, Department of Invertebrates, U.S. National 

Museum of Natural History, Smithsonian Institution, 

Washington, D.C.; Mr. Roger Portell, Department of Invertebrate 

Paleontology, Florida Museum of Natural 

History, Gainesville; and Dr. Richard L. Turner, Department 

of Biological Sciences, Florida Institute of 

Technology, Melbourne. Each of the reviewers contributed 

comments that improved the list. I am especially 

grateful to each of them for providing hard-tofind 

information on authorities for some of the higher 

taxa; that information proved to be quite troublesome 

to locate for some groups. Sandra LaGant, Florida Department 

of Environmental Protection, Florida Marine 

Research Institute, also assisted by providing 

depth records for several taxa from the database of the 

Institute’s Marine Specimen Collection. 


Echinodermata—W. G. Lyons Macroinvertebrate Checklists Camp et al. 1998 

Phylum Echinodermata Leske, 1778 

Class Crinoidea Miller, 1821 

Feather Stars 

Order Comatulida A. H. Clark, 1908 

Family Comasteridae A. H. Clark, 1908 

Genus Comactinia A. H. Clark, 1909 

Comactinia echinoptera (Müller, 1840) 

Comactinia meridionalis (L. Agassiz, 1865) 

Genus Nemaster A. H. Clark, 1909 

Nemaster discoideus (Carpenter, 1888) 

Nemaster rubiginosus (Pourtalès, 1869) 

Class Asteroidea de Blainville, 1830 

Sea Stars 

Order Paxillosida Perrier, 1884 

Family Luidiidae Sladen, 1889 

Genus Luidia Forbes, 1839 

Luidia alternata (Say, 1825) 

Luidia clathrata (Say, 1825) 

Luidia senegalensis (Lamarck, 1816) 

Family Astropectinidae Gray, 1840 

Genus Astropecten Gray, 1840 

Astropecten articulatus (Say, 1825) 

Astropecten comptus Verrill, 1915 

Astropecten duplicatus Gray, 1840 

Order Valvatida Perrier, 1884 

Family Asterinidae Gray, 1840 

Genus Asterina Nardo, 1834 

Asterina folium (Lütken, 1860) 

Family Goniasteridae Forbes, 1841 

Genus Goniaster L. Agassiz, 1836 

Goniaster tessellatus (Lamarck, 1816) 

Family Ophidiasteridae Verrill, 1870 

Genus Copidaster A. H. Clark, 1948 

Copidaster lymani A. H. Clark, 1948 

Genus Linckia Nardo, 1834 

Linckia guildingii Gray, 1840 

Checklist of the Echinodermata 

Genus Ophidiaster L. Agassiz, 1836 

Ophidiaster bayeri A. H. Clark, 1948 

Ophidiaster guildingii Gray, 1840 

Family Mithrodiidae Viguier, 1878 

Genus Mithrodia Gray, 1840 

Mithrodia clavigera (Lamarck, 1816) 

Family Asteropseidae 

Hotchkiss & A. M. Clark, 1976 

Genus Poraniella Verrill, 1914 

Poraniella echinulata (Perrier, 1881) 

Family Oreasteridae Fisher, 1911 

Genus Oreaster Müller & Troschel, 1842 

Oreaster reticulatus (Linnaeus, 1758) 

Order Spinulosida Perrier, 1884 

Family Echinasteridae Verrill, 1870 

Genus Echinaster Müller & Troschel, 1840 

Subgenus Othilia Gray, 1840 

Echinaster (Othilia) echinophorus (Lamarck, 1816) 

Echinaster (Othilia) graminicola Campbell & 

Turner, 1984 

Echinaster (Othilia) paucispinus A. M. Clark, 1987 

Echinaster (Othilia) sentus (Say, 1825) 

Echinaster (Othilia) spinulosus Verrill, 1869 

Order Forcipulatida Perrier, 1884 

Family Asteriidae Gray, 1840 

Genus Asterias Linnaeus, 1758 

Asterias forbesii (Desor, 1848) 

Class Ophiuroidea Gray, 1840 

Brittle Stars, Basket Stars 

Order Phrynophiurida Matsumoto, 1915 

Family Ophiomyxidae Ljungman, 1867 

Genus Ophiomyxa Müller & Troschel, 1840 

Ophiomyxa flaccida (Say, 1825) 

Family Gorgonocephalidae Ljungman, 1867 

Genus Asteroporpa Örsted & Lütken, 1856 

Asteroporpa annulata Örsted & Lütken, 1856 


Camp et al. 1998 Macroinvertebrate Checklists W. G. Lyons—Echinodermata 

Genus Astrophyton Fleming, 1828 

Astrophyton muricatum (Lamarck, 1816) 

Genus Schizostella A. H. Clark, 1952 

Schizostella bifurcata A. H. Clark, 1952 

Order Ophiurida Müller & Troschel, 1840 

Family Ophiuridae Lyman, 1865 

Genus Ophiolepis Müller & Troschel, 1840 

Ophiolepis elegans Lütken, 1859 

Ophiolepis impressa Lütken, 1859 

Ophiolepis paucispina (Say, 1825) 

Family Ophiocomidae Ljungman, 1867 

Genus Ophiocoma L. Agassiz, 1836 

Ophiocoma echinata (Lamarck, 1816) 

Ophiocoma paucigranulata Devaney, 1974 

Ophiocoma pumila Lütken, 1859 

Ophiocoma wendtii Müller & Troschel, 1842 

Genus Ophiocomella A. H. Clark, 1939 

Ophiocomella ophiactoides (H. L. Clark, 1901) 

Genus Ophiopsila Forbes, 1843 

Ophiopsila hartmeyeri Koehler, 1913 

Ophiopsila riisei Lütken, 1859 

Ophiopsila vittata H. L. Clark, 1918 

Family Ophionereididae Ljungman, 1867 

Genus Ophionereis Lütken, 1859 

Ophionereis olivacea H. L. Clark, 1901 

Ophionereis reticulata (Say, 1825) 

Ophionereis squamulosa Koehler, 1914 

Ophionereis vittata Hendler, 1995 

Family Ophiodermatidae Ljungman, 1867 

Genus Ophioderma Müller & Troschel, 1840 

Ophioderma appressum (Say, 1825) 

Ophioderma brevicaudum Lütken, 1856 

Ophioderma brevispinum (Say, 1825) 

Ophioderma cinereum Müller & Troschel, 1842 

Ophioderma guttatum Lütken, 1859 

Ophioderma holmesii (Lyman, 1860) 

Ophioderma phoenium H. L. Clark, 1918 

Ophioderma rubicundum Lütken, 1856 

Ophioderma squamosissimum Lütken, 1856 

Family Ophiactidae Matsumoto, 1915 

Genus Hemipholis Lyman, 1865 

Hemipholis elongata (Say, 1825) 

Genus Ophiactis Lütken, 1856 

Ophiactis algicola H. L. Clark, 1933 

Ophiactis plana Lyman, 1869 

Ophiactis quinqueradia Ljungman, 1871 

Ophiactis rubropoda Singletary, 1974 

Ophiactis savignyi (Müller & Troschel, 1842) 

Family Amphiuridae Ljungman, 1867 

Genus Amphiodia Verrill, 1899 

Amphiodia planispina (von Martens, 1867) 

Amphiodia pulchella (Lyman, 1869) 

Amphiodia trychna H. L. Clark, 1918 

Genus Amphioplus Verrill, 1899 

Amphioplus abditus (Verrill, 1871) 

Amphioplus coniortodes H. L. Clark, 1918 

Amphioplus sepultus Hendler, 1995 

Amphioplus thrombodes H. L. Clark, 1918 

Genus Amphipholis Ljungman, 1867 

Amphipholis atra (Stimpson, 1852) 

Amphipholis gracillima (Stimpson, 1852) 

Amphipholis januarii Ljungman, 1867 

Amphipholis squamata (Delle Chiage, 1828) 

Genus Amphiura Forbes, 1843 

Subgenus Amphiura Forbes, 1843 

Amphiura (Amphiura) fibulata Koehler, 1913 

Amphiura (Amphiura) palmeri Lyman, 1882 

Amphiura (Amphiura) stimpsonii Lütken, 1859 

Subgenus Ophionema Lütken, 1869 

Amphiura (Ophionema) intricata (Lütken, 1869) 

Genus Ophiocnida Lyman, 1865 

Ophiocnida scabriuscula (Lütken, 1859) 

Genus Ophionephthys Lütken, 1869 

Ophionephthys limicola Lütken, 1869 

Genus Ophiophragmus Lyman, 1865 

Ophiophragmus cubanus (A. H. Clark, 1917) 

Ophiophragmus filograneus (Lyman, 1875) 

Ophiophragmus moorei Thomas, 1965 

Ophiophragmus pulcher H. L. Clark, 1918 

Ophiophragmus riisei (Lütken, 1859) 

Ophiophragmus septus (Lütken, 1859) 

Ophiophragmus wurdemanii (Lyman, 1860) 

Genus Ophiostigma Lütken, 1856 

Ophiostigma isocanthum (Say, 1825) 

Ophiostigma siva Hendler, 1995 



Family Ophiothricidae Ljungman, 1866 

Genus Ophiothrix Müller & Troschel, 1840 

Ophiothrix angulata (Say, 1825) 

Ophiothrix brachyactis H. L. Clark, 1915 

Ophiothrix lineata Lyman, 1860 

Ophiothrix orstedii Lütken, 1856 

Ophiothrix suensonii Lütken, 1856 

Class Echinoidea Leske, 1778 

Sea Urchins, Sea Biscuits, Sand Dollars, Heart Urchins 

Order Cidaroida Claus, 1880 

Family Cidaridae Gray, 1825 

Genus Eucidaris Pomel, 1883 

Eucidaris tribuloides (Lamarck, 1816) 

Genus Stylocidaris Mortensen, 1909 

Stylocidaris affinis (Philippi, 1845) 

Order Diadematoida Duncan, 1889 

Family Diadematidae Gray, 1855 

Genus Astropyga Gray, 1825 

Astropyga magnifica A. H. Clark, 1934 

Genus Diadema Gray, 1825 

Diadema antillarum (Philippi, 1845) 

Order Arbacioida Gregory, 1900 

Family Arbaciidae Gray, 1855 

Genus Arbacia Gray, 1835 

Arbacia punctulata (Lamarck, 1816) 

Order Temnopleuroida Mortensen, 1942 

Family Temnopleuridae A. Agassiz, 1872 

Genus Genocidaris A. Agassiz, 1869 

Genocidaris maculata A. Agassiz, 1869 

Family Toxopneustidae Troschel, 1872 

Genus Lytechinus A. Agassiz, 1863 

Lytechinus variegatus (Lamarck, 1816) 

Lytechinus williamsi Chesher, 1968 

Genus Tripneustes L. Agassiz, 1841 

Tripneustes ventricosus (Lamarck, 1816) 

Order Echinoida Claus, 1876 

Family Echinometridae Gray, 1825 

Genus Echinometra Gray, 1825 

Echinometra lucunter (Linnaeus, 1758) 

Echinometra viridis A. Agassiz, 1863 

Order Holectypoida Duncan, 1889 

Family Echinoneidae L. Agassiz & Desor, 1847 

Genus Echinoneus Leske, 1778 

Echinoneus cyclostomus Leske, 1778 

Order Clypeasteroida A. Agassiz, 1872 

Family Clypeasteridae L. Agassiz, 1835 

Genus Clypeaster Lamarck, 1801 

Clypeaster luetkeni Mortensen, 1948 

Clypeaster prostratus Ravenel, 1848 

Clypeaster rosaceus (Linnaeus, 1758) 

Clypeaster subdepressus (Gray, 1825) 

Family Mellitidae Stefanini, 1911 

Genus Encope L. Agassiz, 1840 

Encope aberrans von Martens, 1867 

Encope michelini L. Agassiz, 1841 

Genus Leodia Gray, 1852 

Leodia sexiesperforata (Leske, 1778) 

Genus Mellita L. Agassiz, 1841 

Mellita isometra Harold & Telford, 1990 

Mellita tenuis H. L. Clark, 1940 

Order Spatangoida Claus, 1876 

Family Schizasteridae Lambert, 1905 

Genus Moira A. Agassiz, 1872 

Moira atropos (Lamarck, 1816) 

Genus Paraster Pomel, 1869 

Paraster doederleini Chesher, 1972 

Paraster floridiensis (Kier & Grant, 1965) 

Family Brissidae Gray, 1855 

Genus Brissus Gray, 1825 

Brissus unicolor (Leske, 1778) 

Genus Meoma Gray, 1851 

Meoma ventricosa (Lamarck, 1816) 

Genus Plagiobrissus Pomel, 1883 

Plagiobrissus grandis (Gmelin, 1788) 



Class Holothuroidea de Blainville, 1834 

Sea Cucumbers 

Order Dendrochirotida Brandt, 1835 

Family Cucumariidae Ludwig, 1894 

Genus Duasmodactyla Ayres, 1852 

Duasmodactyla seguroensis (Deichmann, 1930) 

Genus Euthyonacta Deichmann, 1954 

Euthyonacta solida (Deichmann, 1930) 

Genus Ocnus Forbes, 1841 

Ocnus pygmaeus (Théel, 1886) 

Ocnus surinamensis (Semper, 1868) 

Ocnus suspectus (Ludwig, 1874) 

Genus Thyonella Verrill, 1872 

Thyonella gemmata (Pourtalès, 1851) 

Thyonella pervicax (Théel, 1886) 

Thyonella sabanillaensis (Deichmann, 1930) 

Family Sclerodactylidae Panning, 1949 

Genus Euthyonidiella Heding & Panning, 1954 

Euthyonidiella destichada (Deichmann, 1930) 

Euthyonidiella trita (Sluiter, 1910) 

Genus Pseudothyone Panning, 1949 

Pseudothyone belli (Ludwig, 1886) 

Genus Sclerodactyla Ayres, 1851 

Sclerodactyla briareus (Lesueur, 1824) 

Family Phyllophoridae Östergren, 1907 

Genus Allothyone Panning, 1949 

Allothyone mexicana (Deichmann, 1946) 

Genus Neothyonidium Deichmann, 1938 

Neothyonidium parvum (Ludwig, 1881) 

Genus Phyllophorus Grube, 1840 

Subgenus Urodemella Deichmann, 1944 

Phyllophorus (Urodemella) arenicola Pawson & 

Miller, 1992 

Phyllophorus (Urodemella) occidentalis (Ludwig, 

1875) 

Genus Stolus Selenka, 1867 

Stolus cognatus (Lampert, 1885) 

Genus Thyone Jaeger, 1833 

Thyone crassidisca Pawson & Miller, 1981 

Thyone deichmannae Madsen, 1941 

Thyone pawsoni Tommasi, 1972 

Thyone pseudofusus Deichmann, 1930 

Order Aspidochirotida Brandt, 1835 

Family Stichopodidae Haeckel, 1896 

Genus Astichopus H. L. Clark, 1922 

Astichopus multifidus (Sluiter, 1910) 

Genus Isostichopus Deichmann, 1958 

Isostichopus badionotus (Selenka, 1867) 

Family Holothuriidae Ludwig, 1894 

Genus Actinopyga Bronn, 1860 

Actinopyga agassizii (Selenka, 1867) 

Genus Holothuria Linnaeus, 1758 

Subgenus Cystipus Haacke, 1880 

Holothuria (Cystipus) cubana Ludwig, 1875 

Subgenus Halodeima Pearson, 1914 

Holothuria (Halodeima) floridana Pourtalès, 1851 

Holothuria (Halodeima) grisea Selenka, 1867 

Holothuria (Halodeima) mexicana Ludwig, 1875 

Subgenus Holothuria Linnaeus, 1758 

Holothuria (Holothuria) dakarensis Panning, 1939 

Subgenus Platyperona Rowe, 1969 

Holothuria (Platyperona) parvula (Selenka, 1867) 

Holothuria (Platyperona) rowei Pawson & Gust, 

1981 

Subgenus Semperothuria Deichmann, 1958 

Holothuria (Semperothuria) surinamensis Ludwig, 

1875 

Subgenus Theelothuria Deichmann, 1958 

Holothuria (Theelothuria) princeps Selenka, 1867 

Subgenus Thymiosycia Pearson, 1914 

Holothuria (Thymiosycia) arenicola Semper, 1868 

Holothuria (Thymiosycia) impatiens (Forskål, 1775) 

Holothuria (Thymiosycia) thomasi Pawson & 

Caycedo, 1980 

Order Molpadiida Müller, 1850 

Family Caudinidae Heding, 1931 

Genus Paracaudina Heding, 1931 

Paracaudina chilensis obesacauda (H. L. Clark, 

1907) 

Order Apodida Brandt, 1835 

Family Synaptidae Burmeister, 1837 

Genus Epitomapta Heding, 1928 

Epitomapta roseola (Verrill, 1873) 



Genus Euapta Östergren, 1898 

Euapta lappa (Müller, 1850) 

Genus Leptosynapta Verrill, 1867 

Leptosynapta crassipatina H. L. Clark, 1924 

Leptosynapta multigranula H. L. Clark, 1924 

Leptosynapta parvipatina H. L. Clark, 1924 

Leptosynapta tenuis (Ayers, 1851) 

Genus Protankyra Östergren, 1898 

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A 

abbotti, Ividia ...................29 

abbreviata, Bopyrina .............133 

abbreviatum, Stylocheiron .........138 

abbreviatus, Eurypanopeus .........148 

abdita, Ampelisca ...............126 

abditus, Amphioplus .............193 

aberrans, Coralliophila ............24 

aberrans, Encope ................194 

aberrans, Microphthalmus ..........81 

aberrans, Stosicia ................18 

abiuma, Namalycastis .............84 

Abra .........................13 

abranchiata, Glycera ..............81 

Abyssoninoe....................87 

acantha, Actaea.................148 

Acanthaxius ...................143 

acanthochirus, Glypturus ..........142 

Acanthochitona ..................7 

Acanthochitonidae ...............7 

Acanthohaustorius...............129 

Acanthonyx ...................146 

Acanthopale....................85 

acanthophora, Domecia ...........148 

Acanthopleura ...................7 

Acanthotrophon .................23 

acanthura, Upogebia .............143 

acanthuri, Anilocra ..............135 

Acar ..........................9 

Accalathura ...................132 

accincta, Kurtziella ...............28 

Acetes .......................139 

acherusicum, Monocorophium.......128 

acicula, Creseis ..................31 

acicula, Goniada .................81 

aciculagrossa, Pionosyllis ...........83 

acicularis, Cymbovula .............20 

acicularis, Erosaria ...............20 

acicularium, Marphysa ............87 

aciculata, Capitella ...............92 

aciculata, Sphaerosyllis ............83 

aciculata, Syllis..................83 

ackleianus, Pachycheles ...........144 

Aclididae .....................23 

acmeceps, Scoloplos...............88 

Acochlidioidea .................31 

Acoetes .......................85 

Acoetidae .....................85 

Acrocirridae ...................91 

Acrocirrus .....................91 

acropora, Turritella ...............17 

Actaea .......................148 

Acteocina......................30 

Acteon........................30 

Acteonidae ....................30 

Actinopyga....................195 

aculeata, Crepidula ...............20 

aculeata, Harmothoe ..............84 

aculeata, Pitho .................147 

acuminata, Aphrodita .............84 

Index of Scientific Names 

acuminata, Nereis ................84 

acuminata, Nerocila..............135 

acuminata, Ophelina ..............91 

acuminata, Synelmis ..............82 

acurugata, Drillia ................27 

acuta, Anilocra .................135 

acuta, Nuculana ..................8 

acuticauda, Excorallana ...........135 

acuticornis, Mithrax .............146 

acutidens, Odostomia..............29 

acutifrons, Ampharete .............93 

acutum, Apocorophium ...........128 

acutus, Lumbrinerides .............87 

acutus, Nassarius ................24 

acutus, Tharyx ..................91 

adamsi, Arcopsis..................9 

adamsi, Finella ..................17 

adamsii, Megalomphalus ...........20 

adamsii, Seila...................22 

adansoni, Lasaea.................11 

adelae, Calliostoma ...............16 

adelae, Cancellaria ...............27 

adelus, Parviphos ................24 

adina, Menippe.................148 

adkisoni, Nannosquilla ...........125 

advena, Neosimnia ...............20 

Aega ........................134 

Aegidae .....................134 

Aegires .......................32 

Aegiretidae....................32 

aegis, Lucapina..................16 

aeglees, Niso....................23 

aenigmaticus, Navanax ............30 

Aeolidiidae....................34 

Aepinus ......................146 

aequalis, Abra ..................13 

aequalis, Thysanopoda ............139 

aequinoctialis, Scyllarides..........143 

aequisculpta, Rimula ..............16 

aequistriata, Tellina ...............12 

Aesopus.......................25 

aestuarius, Parapleustes ...........131 

Aethogebia ....................142 

affine, Eulithidium ...............16 

affinis, Palicus..................150 

affinis, Stylocidaris ..............194 

affinis, Upogebia ................143 

Agarna ......................135 

agassizi, Ampelisca ..............126 

agassizii, Actinopyga .............195 

agassizii, Agatrix.................27 

agassizii, Berthella ...............32 

agassizii, Lobopilumnus ...........148 

agassizii, Polyschides...............8 

agassizii, Pseudomedaeus ..........149 

Agathotoma ....................27 

Agatrix .......................26 

agelas, Synalpheus ..............141 

agglutinanta, Cumella ............138 

aggregata, Polydora ...............89 

agilis, Armandia .................91 

Aglajidae .....................30 

Aglaophamus ...................84 

agria, Peristichia .................29 

Agriopoma .....................13 

Akanthinotanais ................137 

Akera ........................31 

Akeridae .....................31 

Alaba ........................17 

Alachosquilla ..................125 

alata, Caulleriella ................90 

alatus, Isognomon.................9 

alatus, Strombus .................20 

alba, Alima....................125 

alba, Anodontia .................10 

alba, Discodoris .................33 

alba, Eteone ....................80 

alba, Lysianopsis ................130 

alba, Lysilla ....................94 

albicans, Polycirrus ...............94 

albida, Polystira .................27 

albidentata, Lumbrineris ...........87 

albidum, Epitonium...............22 

albocincta, Pilsbryspira ............27 

albolineata,Volvarina .............26 

albomaculata, Pilsbryspira ..........27 

albovittata, Cerithiopsis ............22 

Albunea ......................144 

Albuneidae ...................144 

albus, Nassarius .................25 

Alcirona......................135 

Alexania ......................22 

algicola, Carpias ................133 

algicola, Ophiactis...............193 

Aligena .......................11 

Alima .......................125 

allenae, Nereis ..................84 

alleneae, Cyphoma ...............20 

Allodiplophryxus ...............134 

Allothyone ....................195 

almyra, Americamysis ............126 

Almyracuma...................138 

alosae, Syllis ...................83 

alphaerostris, Ogyrides............142 

alphei, Probopyria ...............133 

Alpheidae....................141 

Alpheoidea ...................141 

Alpheopsis ....................141 

Alpheus ......................141 

alphezemiotes, Ovobopyrus.........133 

alternata, Luidia ................192 

alternata, Siphonaria..............35 

alternata, Syllis..................83 

alternata, Tellina .................12 

alternatus, Palicus ...............150 

Alvania .......................18 

alveata, Favartia .................23 

amabile, Prunum.................26 

Amaea........................22 

Amaeana ......................93 

Amakusanthura ................132 

Amathimysis ..................125 


Index Macroinvertebrate Checklists Camp et al. 1998 

Amathinidae ..................30 

Ambidexter....................142 

ambigua, Pseudeurythoe............86 

ambiseta, Mediomastus ............92 

amblyonyx, Alpheus .............141 

amboinensis, Lysmata ............141 

amboinensis, Thor ...............141 

Americamysis ..................126 

americana, Ampharete .............93 

americana, Chone ................94 

americana, Ethusa ...............145 

americana, Euphausia ............138 

americana, Glycera ...............81 

americana, Glycymeris .............9 

americana, Merhippolyte ..........141 

americana, Neomysis .............126 

americana, Orbinia ...............88 

americana, Parandalia .............82 

americana, Pareurythoe ............86 

americanum, Americhelidium .......130 

americanum, Dentalium ............8 

americanum, Gnathophyllum .......140 

americanum, Lithopoma............16 

americanus, Acetes ..............139 

americanus, Cirrophorus ...........90 

americanus, Leucon ..............138 

americanus, Modiolus ..............8 

americanus, Notomastus ...........92 

americanus, Panopeus ............149 

americanus, Periclimenes ..........140 

americanus, Pomatoceros ...........95 

americanus, Sabaco ...............92 

americanus, Scyllarus ............143 

americanus, Spondylus ............10 

Americardia ....................12 

Americhelidium ................130 

Americhiton.....................7 

Americonuphis ..................86 

Americorchestia ................131 

Americorophium ................128 

Ameroculodes ..................130 

amianta, Lucina .................10 

amica, Syllis....................83 

Ampelisca ....................126 

Ampeliscidae .................126 

Ampharete .....................93 

Ampharetidae .................93 

Amphicteis.....................93 

Amphictene ....................93 

Amphiglena ....................94 

Amphilochidae ................126 

Amphilocus ...................126 

Amphinome ....................86 

Amphinomida .................86 

Amphinomidae ................86 

Amphiodia ....................193 

Amphioplus ...................193 

Amphipholis...................193 

Amphipoda ..................126 

Amphithalamus .................18 

Amphitrite .....................93 

Amphitritides ...................93 

Amphiura ....................193 

amphiurgus, Conus ...............27 

Amphiuridae .................193 

Ampithoe .....................126 

Ampithoidae..................126 

ampla, Pseudione ...............134 

Amygdalum .....................8 

anabathrum, Conus ...............27 

Anadara .......................9 

Anaitides ......................80 

Anamaera ....................128 

Anamixidae ..................127 

Anamixis .....................127 

Anamobaea ....................94 

Anarthruridae.................137 

Anaspidea ....................31 

Anatina .......................12 

anatina, Anatina.................12 

anceps, Portunus ................147 

Anchialina ....................125 

Anchistioides ..................140 

Anchistioididae................140 

Ancininae ....................136 

Ancinus ......................136 

Ancistrosyllis ...................82 

andersoni, Americhiton .............7 

anderssoni, Perinereis .............84 

andreae, Caprella ...............132 

andrewsi, Paralimnoria ...........136 

anguipes, Ischyrocerus ............130 

angulata, Dynamenella ...........136 

angulata, Ophiothrix .............194 

angulatum, Epitonium .............22 

angulatus, Alpheus ..............141 

angulatus, Latirus ................25 

angulifera, Littoraria ..............18 

anguliferum, Periploma ............15 

angulosa, Tellina .................12 

angusta, Fissurella................16 

angustifrons, Hexapanopeus ........148 

angustifrons, Neopisosoma .........144 

angustior, Nodilittorina ............18 

angustipes, Platydoris .............33 

Anilocra......................135 

anisodon, Pitho .................147 

Anisodoris .....................33 

Anisopagurus ..................144 

anneae, Oriopsis .................95 

Annelida .....................80 

annenectens, Lysippe ..............93 

annulata, Asteroporpa ............192 

annulata,Vermiliopsis .............95 

annulipes, Pagurus ..............144 

anoculata, Pelogenia ..............85 

Anodontia .....................10 

anomala, Aporobobyrina ..........134 

anomala, Iselica .................30 

Anomalocardia ..................13 

anomalus, Paguristes .............143 

Anomia .......................10 

Anomiidae ....................10 

Anomura ....................143 

Anopsilana....................135 

Anotomastus ...................92 

Antalis ........................8 

antarcticus, Parribacus ............143 

antennata, Litocorsa ..............82 

antennata, Pseudopolydora..........89 

Anthuridae ...................132 

Anthuridea ...................132 

Anticlimax .....................19 

antiguensis, Anchistioides..........140 

antillarum, Antalis ................8 

antillarum, Bractechlamys ...........9 

antillarum, Caecum...............19 

antillarum, Diadema .............194 

antillarum, Haminoea .............31 

antillarum, Lithophaga .............8 

antillarum, Nassarius .............25 

antillarum, Oxynoe ...............31 

antillensis, Aega ................134 

antillensis, Aphelodoris ............33 

antillensis, Clibanarius ...........143 

antillensis, Cryptodromiopsis .......145 

antillensis, Cyerce ................32 

antillensis, Enoplometopus .........142 

antillensis, Microphrys............146 

antillensis, Pseudocoutierea.........140 

antillensis, Sphenia ...............14 

Antillophos ....................24 

antiquatus, Hipponix..............20 

antonii, Tectarius.................18 

Antromysis....................126 

Anuropodione ..................134 

Aonides .......................89 

Aoridae .....................127 

Aphelochaeta ...................90 

Aphelodoris ....................33 

Aphrodita .....................84 

Aphroditidae ..................84 

aphroditois, Eunice ...............86 

Aphrogenia ....................84 

apicinum, Prunum ...............26 

apiculata, Chaetopleura .............7 

apiculatum, Epitonium.............22 

apioceros, Synalpheus ............141 

Aplacophora ...................7 

Aplustridae ...................30 

Aplysia .......................31 

Aplysiidae ....................31 

Aplysiopsis ....................32 

Apocorophium .................128 

Apodida .....................195 

Apoprionospio ..................89 

Aporobobyrina .................134 

Aporobopyrus ..................134 

appendiculata, Dulichiella .........128 

appendiculatus, Polygordius .........96 

appressum, Ophioderma...........193 

Apseudes .....................137 

Apseudidae ..................137 

Apseudoidea..................137 

Apseudomorpha ...............137 

aquafuscum, Americorophium .......128 

aquatile, Cymatium ...............21 

aquilina, Upogebia ..............143 

Arabella ......................88 

Araphura .....................137 

Aratus .......................149 

Arbacia ......................194 

Arbaciidae ...................194 

Arbacioida ...................194 

Arca ..........................9 


Camp et al. 1998 Macroinvertebrate Checklists Index 

arcas, Terebra ...................27 

Archaeobrachyura..............145 

Archaeogastropoda..............15 

Archaeopulmonata ..............34 

Archidorididae .................33 

Architectonica ..................29 

Architectonicidae ...............29 

Arcidae .......................9 

Arcinella ......................11 

Arcoida .......................9 

Arcopsis .......................9 

arctica, Hiatella .................14 

Arcturella.....................137 

Arcturidae ...................137 

Arcturinae ...................137 

arcuata, Diodora .................15 

arcuata, Hypoconcha .............145 

arcuata,Vitreolina ................23 

arenae, Phyllodoce ...............80 

Arenaeus .....................147 

arenaria, Axianassa ..............142 

Arene ........................16 

Arenicola ......................93 

arenicola, Chiridotea .............137 

arenicola, Holothuria .............195 

arenicola, Phyllophorus ...........195 

arenicola, Scyphacella ............136 

arenicola, Sigalion ...............85 

Arenicolidae...................92 

arenosa, Exogone ................82 

arenosa, Pandora.................15 

arenosum, Branchiomma ...........94 

areolata, Turbonilla ...............30 

areolatus, Pleurobranchus ..........32 

Argissa ......................127 

Argissidae....................127 

argo, Argonauta .................35 

Argonauta .....................35 

Argonautidae ..................35 

Argopecten .....................9 

argus, Panulirus ................143 

Aricidea.......................90 

aristata, Lithophaga ...............8 

Aristiidae ....................127 

Armandia .....................91 

Armases......................149 

armata, Dipolydora ...............89 

armata, Syllidia .................82 

armatus, Alpheus ...............141 

armatus, Petrolisthes .............144 

armillaris, Syllis .................83 

armillatus, Alpheus ..............141 

Armina .......................34 

Arminidae ....................34 

Arthropoda...................124 

articulata, Eunice ................86 

articulatus, Astropecten ...........192 

Aruga .......................130 

Asaphis .......................13 

ascidiarum, Periclimenaeus .........140 

Asclerocheilus...................91 

Ascobulla......................31 

Asellota .....................132 

Aspella .......................23 

asper, Callidactylus ..............146 

Aspidochirotida ...............195 

Assiminea .....................19 

Assimineidae ..................19 

Astacidea ....................142 

Astacilla .....................137 

Astarte .......................11 

Astartidae.....................11 

Asterias ......................192 

Asteriidae ....................192 

Asterina ......................192 

Asterinidae ...................192 

Asteroidea ...................192 

Asteronotidae..................33 

Asteroporpa ...................192 

Asteropseidae.................192 

Asthenothaerus..................15 

Astichopus ....................195 

Astralium .....................16 

Astropecten ...................192 

Astropectinidae ...............192 

Astrophyton ...................193 

Astropyga ....................194 

Astyris........................25 

Asymmetrione..................134 

Atagema ......................33 

Athelginae ...................133 

athleenae, Anticlimax .............19 

Atlanta .......................36 

atlantica, Callianassa.............142 

atlantica, Casmaria ponderosa .......21 

atlantica, Cooperella ..............14 

atlantica, Exogone ................82 

atlantica, Gammaropsis ...........128 

atlantica, Nematoscelis............138 

atlantica, Phylliroe ...............37 

atlantica, Prionoplax .............148 

atlantica, Promysis ..............126 

atlanticus, Discias ...............139 

atlanticus, Glaucus ...............34 

atlanticus, Periclimenaeus..........140 

Atlantidae ....................36 

atlantidis, Solenocera .............139 

atomaria, Niveria ................21 

atra, Amphipholis ...............193 

atrata, Acteocina .................30 

atratum, Cerithium ...............17 

Atrina .........................9 

atropos, Moira .................194 

atrostyla, Kurtziella ...............28 

attenuata, Erichsonella ............137 

attenuatus, Conus ................27 

Atyidae......................139 

Atyoidea .....................139 

Atylus .......................128 

Atys .........................31 

auberiana, Alvania ...............18 

auberiana, Anomalocardia ..........13 

auberiana, Ithycythara.............28 

Audulla ......................128 

augeneri, Grubeulepis .............85 

Augeneria .....................87 

Augeneriella....................94 

aureocinctus, Dentimargo...........26 

auricincta, Eulima ................23 

auricula, Crucibulum..............20 

aurisula, Polycera ................32 

auritula, Pollia ..................24 

auritulus, Favorinus...............34 

Austinixa .....................150 

Austraeolis.....................34 

Autolytus......................82 

Automate .....................141 

avara, Costoanachis...............25 

avena,Volvarina .................26 

avenaceum, Prunum ..............26 

aviculae, Chevalia ...............128 

Axelella .......................27 

Axianassa ....................142 

Axiidae ......................143 

Axioidea .....................143 

Axiopsis......................143 

Axiorygma ....................143 

Axiothella .....................92 

azaria, Hiatella ..................14 

azteca, Hyalella.................129 

aztecus, Farfantepenaeus...........139 

azuropunctata, Oxynoe ............31 

Azygopleon ...................134 

B 

Babelomurex ...................24 

babylonia, Pseudoscilla ............29 

bacescui, Bowmaniella ............125 

bacescui, Cyclaspis ..............138 

bacescui, Hyssura ...............132 

baconi, Laticorophium ............128 

badionotus, Isostichopus ...........195 

bahamensis, Alpheus .............141 

bahamensis, Antromysis...........126 

bahamensis, Halmyrapseudes .......138 

bahamensis, Kalliapseudes .........137 

bahamensis, Stenoplax .............7 

bahia, Americamysis .............126 

Bailya ........................24 

bairdi, Branchiomma ..............94 

bairdii, Arene ...................16 

bairdii, Cryptosoma ..............145 

balbaoensis, Protula ..............95 

balesae, Acanthochitona.............7 

balesi, Asthenothaerus .............15 

balesi, Pleuromalaxis ..............19 

balkomanus, Elasmopus ...........129 

balthica, Idotea .................137 

Banareia .....................148 

Bankia........................15 

bansei, Syllides ..................83 

barbadensis, Fissurella.............16 

barbadensis, Garthiope............148 

barbadensis, Mitra ...............26 

barbarae, Brachycythara ...........27 

Barbatia .......................9 

barbilla, Dipolydora ..............89 

barbouri, Calliostoma .............16 

Barleeia .......................18 

Barleeiidae ....................18 

barnardi, Listriella...............130 

Barnea........................14 

barrattiana, Caryocorbula ..........14 

bartletti, Tenaturris ...............28 



bartschi, Careliopsis ..............29 

bartschi, Fargoa..................29 

bartschi, Pontogeneia .............128 

bartschi, Teredo ..................15 

Basommatophora ...............35 

bassi, Sigambra..................82 

Basterotia......................11 

batabanoensis, Eupolyodontes........85 

Batea ........................127 

Bateidae .....................127 

Bathyporeia ...................131 

Batillaria ......................17 

Batillariidae ...................17 

Batrachonotus..................146 

baudiniana, Ligia ...............136 

bayeri, Hypselodoris ..............33 

bayeri, Ophidiaster ..............192 

bayeri, Tritonia ..................33 

bealiana, Cerodrillia ..............27 

beanum, Entodesma...............15 

beaufortensis, Neopontonides .......140 

beauii, Cyclostremiscus ............19 

beauii, Cylindrobulla..............31 

beetoni, Heteromysis .............126 

belizensis, Pararicia...............88 

bellairsi, Metatiron ..............131 

Bellaspira .....................27 

bellastriata, Semele ...............13 

bellegladeense, Haustellum..........23 

belli, Marphysa .................87 

belli, Pseudothyone ..............195 

bellum, Eulithidium...............16 

bellus, Fossarus..................17 

belotheca, Turbonilla ..............30 

Bemlos.......................127 

benedicti, Armases...............149 

benedicti, Emerita ...............145 

benedicti, Lepidopa ..............145 

benedicti, Spathochlamys ...........10 

benedicti, Streblospio ..............89 

benthophilus, Cerapus ............130 

beringianus, Asclerocheilus..........91 

berkeleyae, Pilargis ...............82 

bermudae,Volvatella ..............31 

bermudensis, Carpias.............133 

bermudensis, Panopeus............149 

bermudensis, Periclimenaeus........140 

bermudensis, Processa ............142 

bermudezi, Folinia................18 

bermudezi,Vitreolina ..............23 

bermudiensis, Armandia ...........91 

Berthelinia .....................31 

Berthella ......................32 

Bhawania .....................86 

Biancolina ....................127 

Biancolinidae .................127 

bicarinata, Ampelisca.............126 

bicaudata, Tomura ................28 

bicolor, Isognomon ................9 

bicolor, Melongena ...............24 

bicolor, Retilaskeya ...............22 

bicolor, Siraius ..................33 

biconica, Brachycythara ............27 

bicornis, Naineris ................88 

bicornutus, Microphrys ...........146 

bidentata, Acteocina ..............30 

bidentata, Dipolydora .............89 

bidentatus, Melampus .............35 

bidentatus, Protomystides...........81 

bifasciata, Eulima ................23 

Biffarius......................142 

bifida, Protodorvillea ..............88 

biformis, Biffarius ...............142 

bifrons, Actaea .................148 

bifurcata, Schizostella ............193 

bigarra, Garosyrrhoe .............131 

bigelowi, Americamysis ...........126 

Bigelowina ....................125 

bilobata, Sphaerosyllis.............83 

bimini, Leidya..................134 

biminiensis, Bigelowina ...........125 

biminiensis, Lamelleulepethus .......85 

binominata, Eunice ...............86 

binza, Chromodoris ...............33 

bioculatum, Megalomma ...........94 

bipartitum, Caecum ...............19 

bipartitus, Lyrodus ...............15 

bipes, Nebalia ..................124 

biplicata, Mitrolumna .............28 

biscaynense, Teinostoma............19 

biscaynensis, Pseudaeginella........132 

bispinata, Arcturella .............137 

bispinosa, Hyalopale ..............86 

bispinosa, Peracle ................36 

bispinosus, Hydroides .............95 

Bispira .......................94 

bisulcatus, Heliacus...............29 

bisulculata, Lithophaga .............8 

bisuturalis, Boonea ...............29 

bitruncata, Panopea...............14 

Bittiolum ......................17 

bituberculatus, Epialtus ...........146 

biunguiculatus, Brachycarpus .......140 

Bivalvia .......................8 

bivittata, Mesanthura.............132 

blainei, Epitonium................22 

blakei, Neolipobranchius ...........92 

blakei, Pettiboneia ................88 

blakei, Solariorbis ................19 

blanesi, Jaspidella ................26 

Blauneria......................34 

boa, Sthenelais ..................85 

Boca ........................127 

Boccardiella ....................89 

bocourti, Callinectes..............147 

Bodotriidae...................138 

Bodotriinae...................138 

Boguea .......................92 

Boguella ......................92 

bohlkeorum, Mothocya............135 

bollingi, Oliva ..................26 

bombyx, Spiophanes ..............89 

bonellii, Crassicorophium ..........128 

bonnieroides, Grandidierella ........127 

boogii, Stenoplax .................7 

Boonea .......................29 

Bopyridae ....................133 

Bopyrina .....................133 

Bopyrinae ....................133 

Bopyrinella ...................133 

Bopyrione ....................133 

Bopyrissa .....................134 

borealis, Cirolana ...............135 

borealis, Mystides ................80 

borealis, Natatolana..............135 

Bornella.......................33 

Bornellidae....................33 

Bosellia .......................32 

Botula .........................8 

bouryi, Schwartziella ..............18 

bousfieldi, Acanthohaustorius .......129 

bousfieldi, Batea ................127 

bousfieldi, Colomastix ............127 

bousfieldi, Liljeborgia.............130 

bousfieldi, Protohaustorius .........129 

bousfieldi, Synalpheus ............141 

bouvieri, Alpheus ...............141 

bouvieri, Parapinnixa.............150 

bowmani, Taphromysis............126 

Bowmaniella ..................125 

Brachidontes ....................8 

brachyactis, Ophiothrix ...........194 

Brachycarpus ..................140 

Brachycythara ..................27 

Brachyrhyncha ................147 

Brachyura ....................145 

Bractechlamys ...................9 

branchiata, Lumbrineris ............87 

branchiatus, Cirrophorus ...........90 

Branchiomaldane ................93 

Branchiomma...................94 

Branchiosyllis...................82 

Brania ........................82 

branneri, Glypturus ..............142 

brasiliana, Anadara ...............9 

brasiliana, Aplysia ...............31 

brasiliana, Iphigenia ..............13 

brasiliensis, Bowmaniella ..........125 

brasiliensis, Erichthonius ..........130 

brasiliensis, Farfantepenaeus........139 

brasiliensis, Lysarete ..............88 

brasiliensis, Podocerus ............131 

Brasilomysis ...................126 

brassicacephala, Biancolina ........127 

brattegardi, Amathimysis ..........125 

brattstroemi, Mysidopsis...........126 

brattstroemi, Onchidella............35 

braziliensis, Excirolana ...........135 

bredini, Neogonodactylus ..........124 

bredini, Periclimenaeus ...........140 

Bresiliidae....................139 

Bresilioidea...................139 

breve, Caecum ..................19 

breviantennata, Exogone ...........82 

brevicarpus, Synalpheus...........141 

brevicaudum, Ophioderma .........193 

brevicirrata, Autolytus .............82 

brevicornis, Liljeborgia............130 

brevidactylus, Pagurus ............144 

brevidentata, Sphaerosyllis..........83 

brevifrons, Macoma...............12 

brevipes, Lumbrineris .............87 

brevirostris, Odontodactylus ........124 

brevirostris, Sicyonia .............139 

brevis, Eunice ...................86 



brevis, Euphausia ...............138 

brevis, Lolliguncula ...............35 

brevispinum, Ophioderma .........193 

brevitelson, Xenanthura ...........132 

briareus, Arene ..................16 

briareus, Octopus ................35 

briareus, Sclerodactyla ............195 

Brissidae.....................194 

Brissus.......................194 

brocha, Netamelita...............129 

Brolginae ....................131 

brooksi, Synalpheus..............141 

broomensis, Syllis ................83 

browniana, Zebina ...............18 

bruneocomatus, Amphitritides........93 

brunnea, Atlanta.................36 

brunnea, Opisthosyllis .............83 

brunneomaculatus, Bemlos .........127 

bryani, Aricidea .................90 

bryerea, Schwartziella .............18 

Buccinidae ....................24 

bucephalum, Phylliroe .............37 

bucera, Nephtys .................84 

bufo, Bufonaria..................22 

Bufonaria......................22 

bulimoides, Limacina..............36 

Bulla .........................31 

bullaoides, Melampus .............35 

bullata, Acteocina ................30 

Bullidae ......................31 

bullula, Olivella .................26 

burgersi, Uca ..................150 

burkei, Ampelisca ...............126 

burkenroadi, Sicyonia ............139 

burryi, Octopus..................35 

burryi, Opalia...................23 

Bursa ........................22 

Bursatella .....................31 

Bursidae......................22 

Bushia........................15 

bushiae, Nassarina ...............25 

bushiana, Fargoa.................29 

bushiana, Pandora................15 

Busycon.......................24 

Busycotypus ....................24 

Byblis .......................126 

byssomiae, Fabia ................150 

C 

Cabira........................82 

cabritii, Haustellum...............23 

cacahuate, Neocallichirus ..........142 

cadenati, Paguristes ..............143 

Cadlina .......................33 

caeca, Liocuna..................127 

Caecidae .....................19 

Caecum .......................19 

caelatum, Lithopoma ..............16 

caelatus, Pyrunculus ..............30 

caerulea, Cuthona ................34 

cailletii, Turbo...................16 

calamus, Graptacme ...............8 

Calappa......................145 

Calappidae ...................145 

Calappinae ...................145 

calcarata, Bornella ...............33 

calcarata, Metoporhaphis ..........146 

calcicola, Nucula .................8 

Calcinus......................143 

calidimaris, Parviturbo.............17 

californiensis, Mediomastus .........92 

Caliphylla .....................32 

Callianassidae.................142 

Callianassinae.................142 

Callianassoidea................142 

Callichirinae ..................142 

Callichirus ....................142 

Callidactylus ..................146 

Callinectes ....................147 

Calliostoma ....................16 

Calliostomatidae................16 

Callista .......................13 

Callistochiton....................7 

Callistoplacidae .................7 

Calotrophon ....................23 

calusa, Modulus .................17 

calva, Triphora ..................22 

calyculata, Filogranula.............95 

Calyptraea .....................20 

Calyptraeidae ..................20 

campechiensis, Mercenaria..........13 

campechiensis, Pholas .............15 

campi, Batea ...................127 

camptocerum, Macrocoeloma .......146 

Campylaspis...................138 

camura, Colomastix ..............127 

canadensis, Haustorius............129 

canaliculata, Acteocina ............30 

canaliculata, Stramonita haemastoma ..24 

canaliculatus, Busycotypus ..........24 

canaliculatus, Turbo...............16 

Cancellaria ....................27 

cancellaria, Barbatia ...............9 

Cancellariidae .................26 

cancellarius, Cantharus ............24 

cancellata, Chione ................13 

cancellata, Rissoina ...............18 

cancellatum, Cyclostrema ...........16 

Cancellus .....................143 

Cancricepon ...................134 

candeana, Diplodonta .............10 

candeana, Tellina ................12 

candeanum, Epitonium ............22 

candeanus, Antillophos ............24 

candeanus, Malleus ...............9 

candei, Acteocina ................30 

candei, Alpheus.................141 

candens, Acteon .................30 

candida, Barbatia .................9 

candida, Lumbrineris..............87 

candidissima, Agathotoma ..........27 

candidula, Niveria................21 

candidus, Longchaeus .............29 

canguzua, Elysia .................32 

canna, Lioglyphostoma ............28 

canrena, Naticarius ...............21 

Cantharus .....................24 

capensis, Mesochaetopterus .........89 

capensis, Paranaitis ...............80 

capensis, Scoloplos ...............88 

capitata, Capitella ................92 

capitata, Glycera.................81 

Capitella ......................92 

Capitellida ....................92 

Capitellidae ...................92 

Caprella......................132 

Caprellidae...................132 

Caprellidea...................132 

Caprellinoididae ...............132 

Capulidae.....................20 

Capulus.......................20 

caraibica, Cymothoa .............135 

caraibicus, Periclimenaeus .........140 

caramella, Doto .................34 

Carazziella.....................89 

Cardiapoda ....................36 

Cardiidae .....................12 

Cardiomya.....................15 

Cardisoma ....................149 

Carditamera....................11 

Carditidae ....................11 

Carditopsis.....................11 

Careliopsis.....................29 

Caribachlamys ...................9 

caribaea, Coralliophila.............24 

caribaea, Corbula ................14 

caribaea, Rissoella................29 

caribaeensis, Truncatella............19 

caribaeorum, Capitella.............92 

caribaeus, Atys ..................31 

caribaeus, Pilumnus..............149 

caribbaea, Muricopsis .............24 

caribbaeus, Hexapanopeus .........148 

caribbaeus, Mithrax..............146 

caribbea, Berthelinia ..............31 

caribbeana, Cumella .............138 

caribbensis, Iridopagurus ..........144 

caribensis, Exogone ...............82 

caribensis, Spiraserpula ............95 

cariboea, Eunice .................86 

caribous, Piromis ................91 

carica, Busycon ..................24 

Caridea......................139 

Carinaria......................36 

Carinariidae ...................36 

carinata, Bankia .................15 

carinata, Batea .................127 

carinata, Leptochela..............139 

carinata, Listriella ...............130 

carinatum, Stylocheiron ...........139 

carinatus, Trimusculus .............35 

carinicallus, Teinostoma ............19 

carinifer, Latirus .................25 

cariosa, Ebalia .................145 

carnaria, Strigilla ................12 

carnea, Pinna ....................9 

carneum, Prunum ................26 

carneus, Typton .................140 

carolii, Metaphrixus..............134 

carolinae, Acetes americanus........139 

carolinae, Dentatisyllis ............82 

carolinae, Goniadides .............81 

carolinae, Sabaco ................92 

carolinense, Tozeuma .............141 



carolinensis, Ancistrosyllis ..........82 

carolinensis, Pagurus .............144 

carolinensis, Polycirrus ............94 

carolinensis, Polyschides ............8 

carolinensis, Sigatica ..............21 

carolinensis, Speocarcinus .........148 

caroliniana, Maera ..............129 

caroliniana, Polymesoda ...........13 

carolinianum, Caecum .............19 

caroliniensis, Pseudohaustorius ......129 

carpenteri, Chevalia..............128 

Carpias ......................133 

Carpilius .....................148 

Carpoporus ...................148 

carunculata, Hermodice ............86 

Caryocorbula ...................14 

casahoya, Amphilocus ............126 

Casmaria ......................21 

Cassidae......................21 

Cassidinidea...................136 

Cassidininae ..................136 

Cassis ........................21 

castanea, Nereiphylla .............80 

castanea, Turbo ..................16 

castaneus, Lioberus ................8 

castrensis, Hyperphrixus ..........134 

castroi, Brasilomysis .............126 

Cataleptodius ..................148 

catenata, Costoanachis.............25 

catenata, Persicula................26 

catesbyana, Schwartziella...........18 

catharinensis, Batea..............127 

catherinae, Aricidea...............90 

catina, Austraeolis................34 

Catriona ......................34 

caudata, Eupleura ................23 

caudata, Jasmineira ...............94 

caudata, Linatella ................22 

caudata, Paracerceis ..............136 

caudata, Phragmatopoma ...........93 

caudicirra, Questa ................90 

Caudinidae ...................195 

Caulleriella ....................90 

caulleryi, Dipolydora..............89 

cavatura, Anamixis ..............127 

cavicola, Salmoneus..............141 

Cavolinia......................36 

Cavoliniidae ................31, 36 

cayenensis, Diodora ..............15 

cayohuesonicus, Teralatirus..........25 

cedrosa, Kleinella ................29 

cellulosa, Favartia ................23 

Cenchritis .....................18 

centralis, Calyptraea ..............20 

centrifuga, Spirolaxis..............29 

Cephalaspidea .................30 

Cephalopoda ..................35 

Ceradocus ....................128 

Cerapus ......................129 

cerasinus, Perioculodes ............130 

Ceratocephale...................83 

Ceratonereis ....................83 

Ceratophyllidia..................33 

ceratopus, Heteractaea ............148 

Ceratozona .....................7 

cerina, Gouldia ..................13 

cerina, Kurtziella.................28 

cerina, Macoma .................12 

cerinella, Cryoturris...............28 

Cerithidea .....................17 

Cerithiidae ....................17 

Cerithiopsidae .................22 

Cerithiopsis ....................22 

Cerithium .....................17 

Cerodrillia .....................27 

cerrutii, Aricidea .................90 

cervina, Aplysia .................31 

cervus, Macrocypraea .............20 

cetrata, Nicolea ..................94 

chacei, Austinixa ................150 

chacei, Fenneralpheus ............141 

chacei, Neopontonides ............140 

chacei, Periclimenaeus ............140 

chacei, Pseudocheles..............139 

chacei, Scyllarus ................143 

chacmol, Gastropteron .............30 

Chaetiliidae ..................137 

Chaetopleura ....................7 

Chaetopleuridae ................7 

chaetopterana, Pinnixa............150 

Chaetopterida .................89 

Chaetopteridae.................89 

Chaetopterus ...................89 

Chaetozone ....................90 

Chama .......................11 

Chamidae.....................11 

championi, Epitonium .............22 

chaptalii, Clio...................36 

Charonia ......................21 

Charybdis ....................147 

Cheilea .......................20 

Chelidonura ....................30 

chelifera, Audulla ...............128 

chelonophilus, Podocerus ..........131 

Chelorchestia ..................131 

Chelura ......................127 

Cheluridae ...................127 

Cheraminae ..................142 

Cheramus.....................142 

Chevalia .....................128 

chica, Doto.....................34 

Chicoreus......................23 

chierchiae, Siriella ...............125 

chilensis, Paracaudina ............195 

chinensis, Leocrates ...............81 

Chione .......................13 

Chiridota .....................196 

Chiridotea ....................137 

Chiridotidae ..................196 

Chiton.........................7 

Chitonidae .....................7 

chittyana, Corbula................14 

Chloeia .......................86 

Chlorodiella ...................148 

chlorotica, Elysia.................32 

Chone ........................94 

choprae, Cancricepon .............134 

choprae, Synsynella..............134 

Chorinus .....................146 

Choristodon ....................14 

Chromodorididae ...............33 

Chromodoris ...................33 

Chrysallida ....................29 

chrysocoma, Pontogenia ............84 

chrysolepis, Paleanotus ............86 

Chrysopetalidae ................85 

Chrysopetalum ..................86 

chydaea, Squilla ................125 

Cidaridae ....................194 

Cidaroida ....................194 

ciliata, Pseudosquilla .............125 

cilindricaudata, Ophelina...........91 

cincinnata, Syllis ................83 

cincta, Magelona.................90 

cinctimanus, Mithraculus ..........147 

cinerea, Hastula .................27 

cinerea, Talparia .................20 

cinerea, Urosalpinx ...............24 

cinereum, Armases...............149 

cinereum, Ophioderma ............193 

Cinetorhynchus.................140 

cingulata, Daphnella ..............28 

cingulata, Scissurella ..............15 

circellum, Gnathophyllum .........140 

circinata, Aglaophamus ............84 

Circulus.......................19 

circumsaltanus, Loki .............134 

circumspiciens, Notaulax ...........95 

Cirolana .....................135 

Cirolanidae...................135 

Cirolaninae...................135 

cirrata, Laonice ..................89 

Cirratodactylidae ..............137 

Cirratodactylus .................137 

Cirratulida ....................90 

Cirratulidae ...................90 

Cirratulus .....................90 

cirratus, Saccocirrus ..............96 

cirrifera, Prionospio...............89 

Cirriformia ....................91 

Cirrophorus ....................90 

Cirsotrema .....................22 

citrinus, Stylocheilus ..............31 

Cittarium......................16 

clappi, Cerodrillia ................27 

clathrata, Distorsio ...............22 

clathrata, Luidia ................192 

clathrus, Truncatella ..............19 

clava, Caecum ..................19 

clavata, Dorvillea ................88 

clavata, Grubeosyllis ..............82 

clavicauda, Cumella..............138 

clavicornis, Macrochaeta ...........91 

clavigera, Mithrodia .............192 

clavium, Teinostoma ..............19 

Cleantioides ...................137 

clenchi, Chromodoris ..............33 

clenchi, Neritina .................17 

clibanarii, Asymmetrione ..........134 

Clibanarius ...................143 

Clio..........................36 

Clionidae .....................36 

Cliopsidae ....................37 

Cliopsis .......................37 

Cloridopsis ....................125 



Clymenella ....................92 

clymenellae, Listriella ............130 

Clypeaster ....................194 

Clypeasteridae ................194 

Clypeasteroida ................194 

Clypeasterophilus ...............150 

clypeatus, Coenobita .............143 

Clythrocerus ...................145 

coarctata, Cumingia...............13 

coccinea, Lumbrineris .............87 

Cochliolepis ....................19 

Codakia.......................10 

coelata, Stenocionops furcata........147 

coeliaca, Trypanosyllis .............83 

Coelocerus ....................146 

Coenobita.....................143 

Coenobitidae .................143 

Coenobitoidea ................143 

coerulea, Ercolania ...............32 

coerulescens, Hippolyte ...........141 

coffeus, Melampus ...............35 

cognata, Pseudione ..............134 

cognatus, Stolus ................195 

coineaui, Hesionura...............80 

collardi, Aporobopyrus ............134 

Colomastigidae................127 

Colomastix ....................127 

colonia, Polydora ................89 

colpos, Ptilanthura...............132 

Colubraria .....................24 

Columbella ....................25 

Columbellidae .................25 

Columbellopsis ..................25 

columbiae, Mysidium.............126 

columnella, Cuvierina .............36 

colymbus, Pteria ..................9 

Comactinia....................192 

Comasteridae .................192 

comis, Janolus...................34 

commensalis, Ancistrosyllis .........82 

commensalis, Dipolydora ...........89 

commensalis, Lepidametria..........85 

communis, Ficus .................21 

complanata, Eurythoe .............86 

Composetia ....................84 

Compsodrillia...................27 

compta, Cymadusa ..............126 

comptum, Cymatium ..............21 

comptus, Astropecten .............192 

Concarnes ....................130 

concava, Cryptopodia.............147 

concava, Opaliopsis...............23 

concava, Terebra .................27 

concavus, Concarnes .............130 

concentrica, Ervilia ...............13 

concentrica, Nuculana ..............8 

concharum, Dodecaceria ...........91 

conchilega, Lanice ................94 

conchyliophora, Xenophora .........20 

Condylocardiidae ...............11 

conglomerans, Eunice .............87 

congregata, Chama ...............11 

coniclus, Eubranchus ..............34 

Conidae ......................27 

Conilerinae...................135 

coniortodes, Amphioplus ..........193 

conoidea, Melanella ..............23 

conradi, Turbonilla ...............30 

conradina, Transennella ............14 

consensus, Nassarius ..............25 

consobrina, Tellina ...............12 

conspersum, Branchiomma ..........94 

constricta, Distorsio...............22 

constricta, Macoma ...............12 

constricta, Ranilia ...............145 

constricta, Turbonilla ..............30 

constrictus, Rimapenaeus ..........139 

contracta, Corbula ................14 

Conus ........................27 

convexa, Crepidula ...............20 

convexa, Eurydice ...............135 

coomansi, Turbonilla ..............30 

Cooperella .....................14 

cooperi, Caecum .................19 

Copidaster ....................192 

Coralaxius ....................143 

coralensis, Zeuxo................137 

coralicola, Cumella ..............138 

coralicola, Joeropsis ..............133 

coralina, Heteromysis.............126 

Corallanidae ..................135 

Corallianassa ..................142 

corallicoides, Syllis ...............83 

corallicola, Euclymene .............92 

corallicola, Syllis.................83 

corallicola, Troglocarcinus..........150 

corallii, Dodecaceria ..............91 

corallinus, Carpilius .............148 

Coralliophaga ..................13 

coralliophaga, Coralliophaga ........13 

coralliophaga, Gregariella ...........8 

Coralliophila ...................24 

Coralliophilidae ................24 

Corambe ......................32 

Corambidae ...................32 

Corbiculidae...................13 

Corbula .......................14 

Corbulidae ....................14 

cordatus, Pitar ..................14 

cordatus, Ucides ................150 

cordiformis, Divariscintilla..........11 

corinnae, Engina .................24 

corinneae, Bosellia ...............32 

cornigera, Elysia .................32 

Cornirostridae .................28 

cornucopiae, Meioceras ............19 

cornuta, Arcinella ................11 

cornuta, Magelona ...............90 

cornuta, Polydora ................89 

cornuta, Syllis ..................83 

cornuticauda, Colomastix ..........127 

cornutum, Tozeuma ..............142 

cornutus, Mithrax ...............146 

corona, Melongena ...............24 

coronata, Doto ..................34 

coronata, Euclymene ..............92 

Coronis ......................125 

Corophiidae ..................128 

Corophiinae ..................128 

corrodens, Dendropoma ............20 

corrugata, Bursa .................22 

corrugata, Janua .................96 

corrugatum, Cymatium ............21 

coryphe, Mithraculus .............147 

cosentini, Mazatlania .............25 

Cosmotriphora ..................22 

Cossura .......................91 

Cossurida .....................91 

Cossuridae ....................91 

costarum, Spiochaetopterus..........90 

Costasiella .....................32 

Costasiellidae ..................32 

costata, Cerithidea................17 

costata, Cyrtopleura...............14 

costata, Parvilucina ...............10 

costatus, Strombus................20 

Costellariidae ..................26 

costellata, Cardiomya .............15 

Costoanachis ...................25 

Coxicerberus...................136 

Crangonidae..................142 

Crangonoidea .................142 

Crassatellidae..................11 

crassicornis, Eupolymnia ...........93 

Crassicorophium ................128 

crassidisca, Thyone ..............195 

crassimanus, Eucratopsis ..........147 

Crassinella.....................11 

crassipatina, Leptosynapta .........196 

Crassispira.....................27 

Crassostrea ....................10 

crassus, Onobops ................18 

Cratena .......................34 

crebripunctata, Mocosoa...........146 

Creedonia .....................34 

crenata, Opalia ..................23 

Crenella .......................8 

crenulata, Accalathura ............132 

crenulata, Stenothoe..............131 

crenulatus, Longchaeus ............29 

Crepidula......................20 

Creseis .....................31, 36 

cribrarius, Arenaeus..............147 

crinita, Notopygos ................86 

crinita, Persephona ..............146 

crinitichelis, Dissodactylus .........150 

criniticornis, Pagurus .............144 

Crinocheta ...................136 

Crinoidea ....................192 

crispata, Elysia ..................32 

cristallina, Cyerce ................32 

cristallina, Tellina ................12 

cristata, Ampelisca...............126 

cristata, Arenicola ................93 

cristata, Austinixa ...............150 

cristata, Melinna.................93 

cristata, Pista ...................94 

cristata, Prionospio ...............89 

cristata, Tellidora.................12 

cristulifrons, Alpheus .............141 

cristulipes, Hemus ...............146 

Cronius ......................147 

crosseana, Petitilla................29 

crosslandi, Gymnonereis ...........84 

cruciata, Hermaea ................32 



Crucibulum ....................20 

Crucigera......................95 

crucis, Harmothoe ................84 

cruentata, Arene .................16 

cruentata, Goniopsis .............149 

Crustacea ....................124 

Cryoturris .....................28 

crypta, Gyptis...................81 

Cryptochiridae ................150 

Cryptochiroidea ...............150 

Cryptoconchus ...................7 

Cryptocopinae ................137 

Cryptodromiopsis ...............145 

cryptomma, Nephtys ..............84 

Cryptopodia ...................147 

Cryptosoma ...................145 

Cryptostrea ....................10 

ctenactis, Sclerostyla ..............95 

ctenidophora, Dayipsammolyce.......85 

Ctenochelidae.................142 

Ctenochelinae.................142 

Ctenodrilida ...................91 

Ctenodrilidae ..................91 

Ctenodrilus ....................91 

Ctenoides.......................9 

cubana, Crassispira ...............27 

cubana, Holothuria ..............195 

cubaniana, Bursa granularis .........22 

cubaniana, Juliacorbula ............14 

cubaniana, Transennella............14 

Cubanocuma ..................138 

cubanus, Cyclostremiscus ...........19 

cubanus, Ophiophragmus ..........193 

cubensis, Clibanarius .............143 

cubensis, Hippa.................145 

cubensis, Laemodonta .............35 

cubensis, Shoemakerella ...........130 

cubitatum, Meioceras..............19 

Cucumariidae .................195 

cudjoe, Cerapus.................130 

culebra, Eunice ..................87 

culebrae,Vandeloscia .............136 

culveri, Laeonereis................84 

Cumacea ....................138 

Cumella ......................138 

Cumewingia ...................138 

Cumingia......................13 

cumingianus, Solecurtus............13 

cumulus, Agarna ................135 

Cuna.........................11 

cuneata, Pythinella ...............11 

cuneata, Rangia .................12 

cuneiformis, Martesia .............14 

cuprea, Diopatra .................86 

curacaoense, Sesarma.............149 

curacaoensis, Hippolyte ...........141 

curacaoensis, Neogonodactylus ......124 

curacaoensis, Pseudosyllides .........83 

curacaoensis, Synalpheus ..........141 

Curidia ......................130 

curta, Turbonilla .................30 

curtatus, Aporobopyrus ...........134 

Cuspidariidae..................15 

cuspidata, Batea ................127 

cuspidata, Clio ..................36 

cuspidata, Pyromaia .............146 

Cuthona ......................34 

Cuvierina......................36 

Cyathura .....................132 

Cyclaspis .....................138 

Cyclinella .....................13 

Cyclodorippidae ...............145 

cycloferum, Caecum...............19 

Cyclograpsus ..................149 

Cyclopecten ....................10 

cyclostomus, Echinoneus...........194 

Cyclostrema ....................16 

Cyclostremella ..................29 

Cyclostremiscus .................19 

Cyclothyca.....................30 

Cyclozodion ...................145 

cydia, Drillia ...................27 

Cyerce ........................32 

Cylichna ......................30 

Cylichnidae ...................30 

cylindrica, Drilonereis .............88 

cylindrica, Pinnixa ..............150 

cylindricus, Alpheus .............141 

cylindricus, Heliacus ..............29 

Cylindrobulla...................31 

Cylindrobullidae ...............31 

cylleniformis, Enaeta ..............25 

Cymadusa ....................126 

Cymatium .....................21 

Cymatoica .....................12 

Cymbovula ....................20 

cymodocea, Astacilla .............137 

Cymothoa ....................135 

Cymothoidae .................135 

cynocephalum, Cymatium ..........21 

Cyphoma......................20 

Cypraecassis ...................21 

Cypraeidae....................20 

cypris, Parametopella .............131 

Cyproideidae .................128 

Cyrenoida .....................11 

Cyrenoididae ..................11 

Cyrtopleura ....................14 

Cystipus......................195 

Cystiscidae ....................26 

D 

Dactylokepon ..................134 

dactylomela, Aplysia ..............31 

Dacyridium .....................8 

dakarensis, Holothuria ............195 

dalli, Cirsotrema .................22 

dalli, Cuna .....................11 

dalli, Turbonilla .................30 

dalva, Trapania..................32 

danae, Callinectes ...............147 

danilevskii, Caprella .............132 

Daphnella .....................28 

Dardanus.....................143 

Dasybranchus ..................92 

dasypodus, Pilumnus .............149 

daucus, Conus ..................27 

daueri, Notomastus ...............92 

Dawsonius ....................142 

dayi, Apoprionospio ..............89 

dayi, Lumbrinerides...............87 

dayi, Lumbrineris ................87 

Dayipsammolyce ................85 

debilis, Drilonereis ...............88 

deblainvillei, Diacavolinia ..........36 

debrogania, Curidia ..............130 

Decamastus ....................92 

Decapoda ....................139 

deceptrix, Squilla................125 

decorata, Cosmotriphora ...........22 

decussata, Crenella ................8 

decussata, Glycymeris ..............9 

decussata, Rissoina ...............18 

decussatus, Serpulorbis ............20 

defensus, Pagurus ...............144 

defilippi, Octopus ................35 

Deflexiodes....................130 

deflorata, Asaphis ................13 

deformans, Synsynella ............134 

deichmannae, Protohaustorius.......129 

deichmannae, Thyone.............195 

delaneyi, Excorallana.............135 

delessertii, Conus ................28 

delta, Cossura...................91 

deltoidea, Thais..................24 

demarestia, Firoloida ..............36 

demissa, Geukensia................8 

Demonax ......................94 

Dendrobranchiata..............139 

Dendrochirotida ...............195 

Dendrodorididae ...............33 

Dendrodoris ....................33 

Dendropoma ...................20 

Dendrostrea ....................10 

Dentaliida .....................7 

Dentaliidae ....................8 

Dentalium ......................8 

dentalius, Autolytus...............82 

dentata, Divaricella ...............10 

dentata, Photis .................128 

Dentatisyllis....................82 

dentatus, Palicus ................150 

denticulata, Eunice ...............87 

denticulata, Ophelia ..............91 

denticulatum, Epitonium ...........22 

denticulatus, Polycirrus ............94 

denticulatus, Xanthodius ..........149 

dentiferum, Glyphostoma ...........28 

dentigera, Emarginula .............16 

Dentimargo ....................26 

dentischium, Bemlos .............127 

depressa, Panoplax ..............148 

depressa, Plagusia ...............149 

depressifrons, Portunus............147 

Depressiscala ...................22 

depressus, Ancinus ..............136 

depressus, Eurypanopeus ..........148 

dermestinum,Vexillum.............26 

Dermomurex ...................23 

deshaysiana, Aega ...............134 

destichada, Euthyonidiella .........195 

desultor, Asymmetrione ...........134 

Detracia ......................35 

Deutella......................132 



Dexaminidae .................128 

Dexiospira .....................96 

Diacavolinia ...................36 

Diacria .......................36 

Diadema .....................194 

Diadematidae .................194 

Diadematoida .................194 

dianae, Paradella................136 

dianthophila, Fargoa ..............29 

dianthus, Hydroides ..............95 

diaphanus, Thliptodon .............37 

Diaphorosoma ..................88 

Diastylidae ...................138 

Diastylis .....................138 

diazi, Branchiosyllis ..............82 

dibranchiata, Glycera .............81 

diceria, Dodecaceria ..............91 

dichroa, Mitrella .................25 

dicomata, Zafrona ................25 

Didianema.....................16 

didyma, Odostomia ...............29 

dietziana, Corbula ................14 

digitata, Nerilla .................96 

dilatata, Nodilittorina .............18 

dilatatus, Epialtus ...............146 

dilatosculptus, Ischnochiton ..........7 

dilectus, Chicoreus ...............23 

diminuta, Exosphaeroma ..........136 

diminuta, Glyphoturris ............28 

Dinocardium ...................12 

Diodora .......................15 

diogenes, Petrochirus .............144 

Diogenidae ...................143 

diomedea, Rubellatoma ............28 

Diopatra ......................86 

Dioplosyllis ....................82 

Dioptromysis ..................126 

diplacanthum, Macrocoeloma .......146 

Diplocheta ...................136 

Diplocirrus.....................91 

Diplodonta.....................10 

Diplophryxus ..................134 

Diplothyra .....................14 

Dipolydora ....................89 

dirampha, Hydroides ..............95 

Discias.......................139 

Discodorididae .................33 

Discodoris .....................33 

discoideus, Nemaster .............192 

discus, Dosinia ..................13 

dislocata, Terebra.................27 

dispar, Exogone..................82 

dispar, Heteromysis ..............126 

dispar, Paraliomera ..............149 

Dispio ........................89 

dissimilis, Bowmaniella ...........125 

dissimilis, Eurypanopeus ..........148 

dissimilis, Unciola...............127 

Dissiminassa ..................130 

Dissodactylus ..................150 

distinctus, Typton................140 

Distorsio ......................22 

distorta, Leidya .................134 

divae, Doto.....................34 

divae, Runcina ..................31 

Divalinga .....................10 

Divaricella.....................10 

Divariscintilla ..................11 

diversa, Peracle ..................36 

divisus, Tagelus..................13 

dobkini, Thor ..................141 

docimus, Heleobops ...............18 

Dodecaceria ....................91 

doederleini, Paraster..............194 

dolabrata, Pyramidella.............29 

Dolabrifera ....................31 

dolabrifera, Dolabrifera ............31 

dolichognatha, Automate ..........141 

Domecia .....................148 

domestica, Pontonia ..............140 

domingensis, Acar.................9 

domingensis, Brachidontes ...........8 

dominguensis, Glans ..............11 

dominicense, Ellobium .............35 

Donacidae ....................12 

Donax ........................13 

Dondice.......................34 

Dorididae .....................33 

Doriopsilla.....................33 

Dorippidae ...................145 

Dorippoidea ..................145 

Doris.........................33 

dorsalis, Sicyonia ...............139 

dorsobranchialis, Monticellina .......91 

Dorvillea ......................88 

Dorvilleidae ...................88 

dorvilliae, Kurtziella ..............28 

Dosinia .......................13 

Doto .........................34 

Dotoidae .....................34 

Dreissenidae ..................13 

Drillia ........................27 

Drilliidae .....................27 

Drilonereis.....................88 

Dromia ......................145 

Dromiidae ...................145 

Dromioidea ..................145 

Duasmodactyla .................195 

dubia, Cloridopsis ...............125 

dubia, Finella ...................17 

dubia, Leptochelia ...............137 

dubia, Libinia ..................146 

dubia, Pherusa ..................91 

Dulichiella ....................128 

dumerilii, Platynereis..............84 

duofurca, Pettiboneia ..............88 

duorarum, Farfantepenaeus.........139 

duplex, Spiophanes ...............89 

duplicata, Neverita ...............21 

duplicatus, Astropecten ...........192 

dupliniana, Crassinella ............11 

dux, Fargoa ....................29 

Dynamenella ..................136 

Dynameninae .................136 

dysbatus, Floridiscrobs.............18 

dyscrita, Rissoina ................18 

dysoni, Diodora .................15 

Dyspanopeus ..................148 

dytiscus, Lepidactylis .............129 

E 

Ebalia .......................145 

Ebalidae ......................29 

Ebaliinae ....................145 

eborea, Graptacme ................8 

eburnea, Cylichna ................30 

eburneolus, Dentimargo ............26 

eburneum, Cerithium..............17 

Echinaster ....................192 

Echinasteridae ................192 

echinata, Ophiocoma .............193 

echinaticosta, Epitonium ...........22 

Echinodermata ................192 

Echinoida ....................194 

Echinoidea ...................194 

Echinometra ...................194 

Echinometridae ...............194 

Echinoneidae .................194 

Echinoneus....................194 

echinophorus, Echinaster ..........192 

echinoptera, Comactinia ...........192 

echinulata, Poraniella ............192 

eclipes, Mysidopsis ..............126 

Edotia .......................137 

edulis, Mytilus ...................8 

edwardsi, Ameroculodes ...........130 

edwardsi, Grapsicepon ............134 

edwardsi, Neoamphitrite ...........94 

Edwinjoycea ...................137 

Edwinjoycinae ................137 

egmontianum, Trachycardium ........12 

ehlersi, Pherusa .................91 

Ehlersia .......................83 

Eisothistos ....................132 

Elachisina .....................19 

Elachisinidae ..................19 

Elasmopus ....................129 

Elasocumella ..................138 

elata, Pilumnoplax...............148 

elata, Platycythara ...............28 

elegans, Bushia..................15 

elegans, Chrysopetalum ............86 

elegans, Dosinia .................13 

elegans, Haminoea ...............31 

elegans, Hydroides ...............95 

elegans, Ophiolepis ..............193 

elegantulum, Dacrydium ............8 

elevatus, Speloeophorus ...........145 

eliasoni, Samythella ..............93 

elizabethae, Dermomurex ...........23 

elliptica, Basterotia ...............11 

ellisi, Americorophium ............128 

Ellobiidae .....................34 

Ellobium ......................35 

elongata, Epialtus dilatatus forma ....146 

elongata, Hemipholis .............193 

elongata, Melita ................129 

Elthusa ......................135 

Elysia ........................32 

Elysiidae......................31 

emarginata, Hemitoma.............16 

emarginata, Libinia ..............146 

emarginata, Tyche ...............147 

Emarginula ....................16 



Emerita ......................145 

emersoni, Pseudobranchiomma .......95 

emersonii, Retilaskeya .............22 

emertoni, Polycerella ..............33 

empusa, Squilla.................125 

Enaeta........................25 

Encope.......................194 

engeli, Flabellina.................34 

engeli, Phyllaplysia ...............31 

Engina .......................24 

engonium, Eulimastoma............29 

enigmatica, Boguea ...............92 

enigmaticus, Ficopomatus...........95 

enopla, Odontosyllis ..............82 

Enoplobranchus .................93 

Enoplometopidae ..............142 

Enoplometopoidea .............142 

Enoplometopus .................142 

Ensis .........................12 

Ensitellops .....................11 

Entodesma .....................15 

Entoniscidae ..................134 

Eobrolgus .....................131 

Eophrixus.....................134 

epheliticus, Hepatus..............145 

Ephesiella .....................81 

Epialtinae ....................146 

Epialtus ......................146 

Epicaridea ...................133 

Epimeria .....................128 

Epimeriidae ..................128 

epiphaneum,Vexillum .............26 

Episcynia......................19 

Episiphon ......................8 

epistomus, Rhepoxynius ...........131 

Epitomapta....................195 

Epitoniidae....................22 

Epitonium .....................22 

equestris, Cheilea ................20 

equestris, Ostrea .................10 

equilibra, Caprella...............132 

Ercolania ......................32 

erectus, Petaloconchus .............20 

eremita, Onuphis ................86 

Erichsonella ...................137 

Erichthonius ...................130 

erinacea, Libinia ................146 

Eriphia ......................148 

eritima, Glyphoturris..............28 

ermineus, Conus .................28 

ernesti, Lumbrineris...............87 

Erosaria.......................20 

erosus, Glyptoxanthus ............148 

Ersilia ........................23 

Ervilia ........................13 

Erycina .......................11 

erythrocoma, Pseudostomatella .......16 

erythronotus, Ischnochiton ...........7 

erythrophylla, Phyllodoce...........80 

Erythropini ...................125 

Erythrops .....................125 

erythrops, Paguristes .............143 

erythropus, Dromia ..............145 

estuariensis, Alpheus .............141 

Eteone ........................80 

Ethusa .......................145 

Ethusinae ....................145 

Etisus .......................148 

Euapta.......................196 

Euarche .......................85 

Eubranchidae ..................34 

Eubranchus ....................34 

Eucalliax .....................142 

Eucalliinae ...................142 

Eucarida .....................138 

Euceramus ....................144 

Euchelus ......................16 

Euchone.......................94 

Eucidaris .....................194 

Euclymene .....................92 

eucosmia, Compsodrillia ...........27 

eucosmius, Fusinus ...............25 

Eucrassatella ...................11 

Eucratopsis....................147 

eucymata, Callista................13 

Eudevenopus ..................131 

euglyptum, Calliostoma ............16 

Eulalia .......................80 

Eulepethidae ..................85 

Eulima .......................23 

Eulimastoma ...................29 

Eulimidae.....................23 

eulimoides, Melanella .............23 

Eulimostraca ...................23 

Eulithidium ....................16 

Eumalacostraca................125 

Eumida .......................80 

Eunice ........................86 

Eunicida......................86 

Eunicidae .....................86 

Euniphysa .....................87 

Euphausia ....................138 

Euphausiacea .................138 

Euphausiidae .................138 

Euphrosine.....................86 

Euphrosinidae .................86 

Eupleura ......................23 

Eupolymnia ....................93 

Eupolyodontes ..................85 

Eurydice......................135 

Eurydicinae ..................135 

eurypalea, Chrysopetalum ..........86 

Eurypanopeus..................148 

Euryplax .....................147 

Eurysquilla....................124 

Eurysquillidae ................124 

Eurysyllis .....................82 

Eurythoe ......................86 

Eurytium .....................148 

Eusiridae ....................128 

Eusiroides ....................128 

Eusyllis .......................82 

eutheca, Macrocoeloma ...........146 

Euthyonacta ...................195 

Euthyonidiella .................195 

Euvola........................10 

evelinae, Discodoris...............33 

evelinae, Elysia..................32 

evelinae, Miesea .................34 

evelynae, Prunum ................26 

evermanni, Automate .............141 

ewingi, Synelmis.................82 

exasperatus, Callinectes ...........147 

exasperatus, Lindapecten ...........10 

excavata, Chiridotea .............137 

excavata, Tegula .................16 

Excirolana ....................135 

excisa, Cymothoa ...............135 

Excorallana ...................135 

Exhippolysmata ................141 

exigua, Janthina .................23 

exiguum,Vexillum ................26 

exilis, Branchiosyllis ..............82 

exilis, Turbonilla .................30 

Exogone.......................82 

exoleta, Turritella ................17 

Exosphaeroma .................136 

exotica, Ligia ..................136 

exustus, Brachidontes ..............8 

F 

Fabia ........................150 

Fabricinuda ....................94 

Facelinidae ....................34 

falcata, Jassa...................130 

falcirama, Colomastix ............127 

fallax, Prionospio ................89 

falsa, Nereis ....................84 

Farfantepenaeus ................139 

Fargoa ........................29 

fargoi, Cryoturris ................28 

fargoi,Vermicularia ...............17 

fasciata, Mesanthura .............132 

fasciata, Tegula ..................16 

fascicularis, Fissurella .............16 

fascigera, Parhyale...............129 

Fasciolaria .....................25 

Fasciolariidae ..................25 

fauchaldi, Pseudeurythoe ...........86 

fausta, Tellina ...................12 

fauveli, Aricidea .................90 

Fauveliopsidae .................91 

Fauveliopsis ....................91 

Favartia .......................23 

Favorinus......................34 

faxoni, Harrieta.................136 

faxoni, Lucifer..................139 

felderi, Fabia...................150 

femorale, Cymatium ..............21 

Fenneralpheus..................141 

ferrugina,“Ehlersia” ..............83 

fibulata, Amphiura ..............193 

Ficidae .......................21 

Ficopomatus ....................95 

Ficus .........................21 

filamentosa, Eunice ...............87 

filamentosa, Leodice ..............87 

filifera,Vitrinella.................19 

filiformis, Erichsonella ............137 

filiformis, Heteromastus ............92 

filigera, Cirriformia ...............91 

filitelsona, Heteromysis ...........126 

Filograna ......................95 

filograneus, Ophiophragmus ........193 



Filogranula ....................95 

filosa, Cymadusa ................127 

filosa, Pyrgocythara...............28 

filosus, Octopus .................35 

fimbriata, Pelogenia...............85 

fimbriata, Processa...............142 

fimbriatula, Bankia ...............15 

Fimbriosthenelais ................85 

Finella........................17 

finitima, Aricidea ................90 

Fiona.........................34 

Fionidae ......................34 

Firoloida ......................36 

fischeri, Schwartziella .............18 

fischeriana, Haliris ...............15 

Fissurella ......................16 

Fissurellidae ...................15 

flabellata, Pherusa................91 

Flabellifera ...................134 

Flabelligerida ..................91 

Flabelligeridae .................91 

Flabellina .....................34 

Flabellinidae...................34 

flaccida, Ophiomyxa .............192 

flamingo, Conus .................28 

flammea, Calappa ...............145 

flammea, Cassis .................21 

flava, Cerithiopsis ................22 

flavescens, Conus ................28 

flexipes, Nematobrachion ..........138 

floralia, Olivella .................26 

florida, Chama ..................11 

florida, Hobsonia ................93 

florida, Mitra ...................26 

floridana, Alexania ...............22 

floridana, Anadara ................9 

floridana, Bowmaniella ...........125 

floridana, Capitella capitata .........92 

floridana, Carditamera.............11 

floridana, Costoanachis ............25 

floridana, Cyrenoida ..............11 

floridana, Elachisina ..............19 

floridana, Erichsonella ............137 

floridana, Erycina ................11 

floridana, Eunice.................87 

floridana, Gymnodorvillea ..........88 

floridana, Holothuria .............195 

floridana, Lucina.................10 

floridana, Lumbrineris .............87 

floridana, Lyonsia ................15 

floridana, Metharpinia............131 

floridana, Onchidella..............35 

floridana, Orobitella ..............11 

floridana, Parapionosyllis...........83 

floridana, Perinereis...............84 

floridana, Perkinsiana .............95 

floridana, Pinnixa ...............150 

floridana, Pleurocrypta............134 

floridana, Stenoplax ...............7 

floridana, Stramonita haemastoma ....24 

floridana, Terebra ................27 

floridana, Thala .................26 

floridana, Thelecythara ............28 

floridana, Tivela .................14 

floridana,Vaunthompsonia .........138 

floridana,Vitrinella ...............19 

floridanum, Caecum ..............19 

floridanum, Chrysopetalum .........86 

floridanus, Allodiplophryxus........134 

floridanus, Alpheus ..............141 

floridanus, Cataleptodius ..........148 

floridanus, Cryptoconchus ...........7 

floridanus, Haplosyllides ...........82 

floridanus, Hydroides .............95 

floridanus, Lyrodus ...............15 

floridanus, Megalomphalus .........20 

floridanus, Melampus .............35 

floridanus, Modulus ..............17 

floridanus, Ovalipes..............147 

floridanus, Palaemon .............140 

floridanus, Pilumnus .............149 

floridanus, Polyophthalmus .........91 

floridanus, Syllides ...............83 

floridanus, Thor.................141 

floridensis, Alachosquilla ..........125 

floridensis, Carpias ..............133 

floridensis, Cirratodactylus .........137 

floridensis, Conus ................28 

floridensis, Gnathia ..............136 

floridensis, Heteromysis ...........126 

floridensis, Mesanthura ...........132 

floridensis, Microjassa ............130 

floridensis, Pinnaxodes............150 

floridensis, Pleurocope ............133 

floridensis, Probopyrus............133 

floridensis, Sabellaria .............93 

floridiensis, Paraster..............194 

Floridiscrobs....................18 

foliaceicosta, Epitonium ............22 

Folinia........................18 

foliosa, Eteone ..................80 

foliosus, Leitoscoloplos.............88 

foliosus, Pterocirrus ...............81 

folium, Asterina ................192 

fonticula, Perkinsiana .............95 

forbesii, Asterias ................192 

forceps, Inachoides ..............146 

forceps, Leptalpheus .............141 

forceps, Mithraculus .............147 

Forcipulatida..................192 

formalis, Heteropodarke ............81 

formosa, Heteromysis.............126 

formosa, Kupellonura.............132 

formosus, Alpheus ...............141 

fornicata, Crepidula...............20 

forresti, Leptochelia ..............137 

forticirratus, Cirrophorus ...........90 

Fossarus ......................17 

fossor, Donax ...................13 

fragilis, Aricidea .................90 

fragilis, Biffarius ................142 

fragilis, Leitoscoloplos .............88 

fragilis, Mactrotoma ..............12 

fragilis, Martesia.................14 

fragilis, Nereiphylla ..............80 

fragosus, Batrachonotus ...........146 

francesae, Parviturbo ..............17 

frenulata, Rimula ................16 

fritzmuelleri, Synalpheus ..........141 

frons, Dendostrea ................10 

frontifilis, Acrocirrus ..............91 

frydis, Tritoniopsis................33 

fucata, Eunice ...................87 

fucata, Splendrillia ...............27 

fucorum, Latreutes ...............141 

fucosus, Dardanus ...............143 

fulgens, Paraonis.................90 

fulgida, Oenone .................88 

fulgurans, Nerita.................17 

fulgurans, Odontosyllis ............82 

fulleri, Teredo ...................15 

fulminatus, Pitar.................14 

fulvescens, Hexaplex ..............23 

fulvocincta, Eulima ...............23 

fulvus, Syllides ..................83 

funerea, Ercolania ................32 

furca, Mysidopsis ...............126 

furcata, Stenocionops .............147 

furcatus, Cirrophorus..............90 

furcifera, Teredo .................15 

fusca, Botula ....................8 

fusca, Sayella ...................29 

fuscata, Ercolania ................32 

fuscata, Tenellia .................34 

fuscescens, Crassispira .............27 

fuscina, Ingolfiella ...............126 

fuscostriata, Pseudovermilia .........95 

fuscovittatus, Stiliger ..............32 

fusiformis, Cerithiopsis ............22 

fusiformis, Owenia ...............93 

Fusinus .......................25 

G 

gabbi, Strigilla ..................12 

gabbii, Glyphostoma ..............28 

Gadila ........................8 

Gadilida.......................8 

Gadilidae ......................8 

Gadilinidae ....................8 

gairacensis, Hydroides .............95 

Galathea .....................144 

Galatheidae ..................144 

Galatheoidea .................144 

galathinus, Petrolisthes............144 

Galathowenia...................93 

galea, Coralliophila...............24 

galea, Tonna ....................21 

Galeommatidae ................11 

galetaensis, Malmgreniella ..........85 

gallagheri, Brania ................82 

gallensis, Stenothoe ..............131 

galloides, Calappa ...............145 

gallus, Strombus .................20 

Gammaridae..................128 

Gammaridea..................126 

Gammaropsis ..................128 

Gammarus ....................129 

gardineri, Paranaitis ..............80 

Garosyrrhoe ...................131 

garrityi, Cumella ................138 

Garthiope.....................148 

Gastrochaena ...................14 

Gastrochaenidae................14 

Gastropoda .................15, 36 



Gastropteridae .................30 

Gastropteron ...................30 

Gastrosaccinae ................125 

gaudchaudi, Hipponoa.............86 

gayheadia, Chaetozone ............90 

Gecarcinidae..................149 

Gecarcinus ....................149 

Geitodoris .....................33 

Gemma .......................13 

gemma, Cardiomya ...............15 

gemma, Gemma .................13 

gemma, Proterato ................21 

gemmata, Thyonella..............195 

gemmatum,Vexillum ..............26 

gemmatus, Pilumnus .............149 

gemmulosa, Chrysallida............29 

Genetyllis .....................80 

Genocidaris ...................194 

Geograpsus ...................149 

georgiana, Stenothoe .............131 

gerardoi, Metapenaeopsis ..........139 

gerlachi, Teleotanais..............137 

germaineae, Mourgona ............32 

gesae, Pionosyllis ................83 

Geukensia ......................8 

giardi, Capitella .................92 

giardi, Dipolydora................89 

giardi, Synalpheion ..............134 

gibba, Amathimysis ..............125 

Gibberosus ....................130 

Gibberula .....................26 

gibbesi, Percnon ................149 

gibbesi, Polyonyx................144 

Gibbesia .....................125 

gibbesii, Albunea................144 

gibbesii, Portunus ...............147 

gibboides, Euphausia .............138 

gibbosa, Cavolinia................36 

gibbosa, Colomastix ..............127 

gibbosa, Plicatula ................10 

gibbosum, Cyphoma ..............20 

gibbulum, Pseudocyphoma ..........20 

gibbus, Argopecten ................9 

gigantea, Pleuroploca..............25 

giganteus, Spirobranchus ...........95 

Gigantione ....................134 

gigas, Strombus .................20 

Gilvossius ....................142 

Gitanopsis ....................126 

glabra, Leiocapitella ..............92 

glabriuscula, Lysiosquillina.........125 

glacialis, Trichobranchus ...........93 

glandulata, Sphaerosyllis ...........83 

Glans ........................11 

Glaucidae.....................34 

Glaucus .......................34 

Globivenus.....................13 

globosa, Janthina ................23 

globosa, Marsenina ...............21 

Globosolembos .................127 

glossema, Terebra ................27 

Glossodoris ....................33 

glutaea, Rhynchospio..............89 

Glycera .......................81 

Glyceridae ....................81 

Glycinde ......................81 

Glycymerididae .................9 

Glycymeris .....................9 

glynni, Paracerceis...............136 

Glyphohesione ..................82 

Glyphostoma ...................28 

Glyphoturris ...................28 

glypta, Murexiella................24 

glypta, Nassarina ................25 

Glyptaesopus ...................28 

Glyptoplax ....................148 

Glyptoxanthus .................148 

Glypturus.....................142 

Gnathia ......................136 

Gnathiidae ...................136 

Gnathiidea ...................136 

Gnathophyllidae ...............140 

Gnathophylloides ...............140 

Gnathophyllum ................140 

Gnathostenetroides ..............133 

Gnathostenetroididae ...........133 

Gnathostenetroididea ...........132 

Godiva .......................34 

gomezi, Syllides .................83 

gomezi, Tropichelura .............127 

gomoiui, Cumella ...............138 

gonagra, Eriphia ................148 

Goneplacidae .................147 

Goniada ......................81 

Goniadella .....................81 

Goniadidae....................81 

Goniadides.....................81 

Goniaster .....................192 

Goniasteridae .................192 

Goniodorididae ................32 

goniogyrus, Teinostoma ............19 

Goniopsis.....................149 

Gonodactylidae................124 

Gonodactyloidea...............124 

goodei, Bhawania ................86 

goodei, Eunice...................87 

goodei, Metapenaeopsis ...........139 

goodei, Synalpheus ..............141 

gordiodes, Anotomastus ............92 

gorei, Aethogebia................143 

gorei, Austinixa.................150 

gorei, Philocheras ...............142 

Goreopagurus ..................144 

Gorgonocephalidae.............192 

gouldi, Bankia ..................15 

Gouldia .......................13 

gouldii, Pectinaria ................93 

gouldii, Tellina ..................12 

gracile, Mysidium ...............126 

gracilimana, Paraxiopsis...........143 

gracilipes, Podochela .............146 

gracilis, Decamastus ..............92 

gracilis, Levinsenia ...............90 

gracilis, Melanella ...............23 

gracilis, Ninoe nigripes ............88 

gracilis, Pachygrapsus ............149 

gracilis, Stenopleustes ............131 

gracilis, Syllis...................83 

gracillima, Amphipholis ...........193 

graduata, Oceanida ...............23 

graminicola, Echinaster ...........192 

Grandidierella .................127 

grandimanus, Neocallichirus........142 

grandis, Cirriformia...............91 

grandis, Plagiobrissus ............194 

granosissima, Geukensia ............8 

Granoturris ....................28 

granularis, Bursa ................22 

granulata, Acanthopleura............7 

granulata, Heterocrypta ...........147 

granulata, Parthenope ............147 

granulatum, Phalium..............21 

granulatus, Conus ................28 

Granulina .....................26 

Graphis .......................23 

Grapsicepon ...................134 

Grapsidae....................149 

Grapsinae....................149 

Grapsoidea ...................149 

Grapsus ......................149 

grapsus, Grapsus................149 

Graptacme......................8 

grayana, Bursa ..................22 

grayi, Nereis....................84 

grayi, Paguristes ................143 

greeleyi, Peltodoris ...............33 

greenii, Cerithiopsis...............22 

Gregariella .....................8 

grillus, Orchestia................131 

grisea, Holothuria ...............195 

groenlandica, Phyllodoce ...........80 

grubei, Naineris .................88 

grubei, Sthenolepis ...............85 

Grubeosyllis....................82 

Grubeulepis ....................85 

gruneri, Tegula ..................16 

grus, Timoclea...................14 

Gryphaeidae ..................10 

guanhumi, Cardisoma ............149 

guanica, Eunice..................87 

guanica, Podarke.................81 

guanicus, Ophiodromus ............81 

guildingii, Linckia ...............192 

guildingii, Ophidiaster ............192 

guinaicum, Cerithium .............17 

guitarti, Heteromysis .............126 

gundlachi, Palinurellus............143 

gunneri, Amphicteis ..............93 

guppyi, Americardia ..............12 

gurgulio, Caecum ................19 

guttarosea, Euchelus ..............16 

guttatum, Ophioderma ............193 

guttatum, Prunum ................26 

guttatus, Panulirus...............143 

gutzui, Cubanocuma .............138 

gymnodactylus, Pagurus ...........144 

Gymnodorvillea .................88 

Gymnonereis ...................84 

Gymnosomata .................36 

Gyptis ........................81 

H 

hadria, Granulina ................26 

haemastoma, Stramonita ...........24 



haematodes, Polycirrus ............94 

haemuli, Anilocra ...............135 

halichondriae, Colomastix .........128 

halidorema, Splendrillia............27 

Haliotinella ....................21 

Haliris........................15 

Halmyrapseudes ................138 

Halodeima ....................195 

Halophilosciidae ...............136 

Halosydna .....................84 

hamata, Boccardiella ..............89 

hamatipes, Argissa ..............127 

hamillei, Phenacolepas ............17 

Haminoea .....................31 

Haminoeidae ..................31 

hamosus, Microphthalmus ..........81 

Hanleya .......................7 

Hanleyidae.....................7 

hanseni, Anamixis...............127 

Hansenium....................133 

Haplocochlias...................17 

haploseta, Psamathe ..............81 

Haplosyllides ...................82 

Haplosyllis ....................82 

harasewychi, Conus...............28 

Hargeria .....................137 

Harmothoe.....................84 

Harpidae .....................26 

Harpilius .....................140 

Harrieta ......................136 

harrietae, Carpias ...............133 

harringtoni, Periclimenes ..........140 

harrisii, Rhithropanopeus ..........149 

hartii, Panopeus ................149 

hartleyanum, Prunum virginianum ....26 

hartmanae, Ancistrosyllis ...........82 

hartmanae, Levidorum.............82 

hartmanae, Microphthalmus.........81 

hartmanae, Streblosoma............94 

hartmannschroederae, Pisione........86 

Hartmanodes ..................130 

hartmeyeri, Ischnochiton ............7 

hartmeyeri, Ophiopsila............193 

hastata, Hastula .................27 

Hastula .......................27 

Hauchiella .....................93 

Haustellum ....................23 

Haustoriidae..................129 

Haustorius ....................129 

havanensis, Ividia ................29 

hawaiensis, Parhyale .............129 

hayesi, Uromunna ...............133 

hayi, Ogyrides .................142 

heardi, Americorchestia ...........131 

heardi, Campylaspis .............138 

heardi, Colomastix...............128 

heardi, Nannosquilla .............125 

heardi, Probopyrinella ............133 

heardi, Synalpheus ..............141 

hebes, Nematonereis ..............87 

Heilprinia .....................25 

heladum, Caecum ................19 

helenae, Fusinus .................25 

Heleobops .....................18 

Heliacus ......................29 

helicinoides, Atlanta ..............36 

helicoidea,Vitrinella ..............19 

helleri, Charybdis ...............147 

Hemiaegina ...................132 

Hemiarthrinae ................134 

Hemiarthrus ...................134 

hemigibba, Euphausia ............138 

Hemipholis....................193 

Hemipodus.....................81 

hemipodus, Notomastus ............92 

Hemiproto ....................132 

Hemitoma .....................16 

hemphilli, Acanthochitona ...........7 

hemphilli, Mithrax ..............146 

hemphilli, Pinnotheres ............150 

hemphilli, Processa ..............142 

hemphilli, Pyrgocythara............28 

hemphilli, Synalpheus ............141 

hemphilli, Turbonilla ..............30 

hemphilli,Vitrinella ..............19 

hemphillii, Asthenothaerus..........15 

hemphillii, Hexapanopeus .........148 

hemphillii, Microeulima ...........23 

hemphillii, Sayella ...............29 

Hemus .......................146 

hendersoni, Dacrydium elegantulum ....8 

hendersoni, Limatula...............9 

hendersoni, Mathilda..............28 

hendersoni, Niso .................23 

hendersoni, Parapinnixa...........150 

hendersoni,Vexillum ..............26 

Henrya .......................29 

hepaticus, Polinices ...............21 

Hepatus ......................145 

herbstii, Panopeus ...............149 

Hermaea ......................32 

Hermenia......................84 

Hermodice .....................86 

heros, Chorinus.................146 

herricki, Synalpheus .............141 

Hesione .......................81 

Hesionidae ....................81 

Hesionura .....................80 

Hespererato ....................20 

Heteractaea ...................148 

Heterobranchia ................28 

heterobranchia, Prionospio ..........89 

heterochaelis, Alpheus ............141 

heterocirrata, Eusyllis .............82 

heteroclita, Blauneria..............34 

Heterocrypta...................147 

Heterodonax....................13 

Heteromastus ...................92 

Heteromysini .................126 

Heteromysis ...................126 

heteropalea, Chrysopetalum .........86 

heteropoda, Eteone ...............80 

heteropoda, Lumbrineris............87 

heteropoda, Scolelepis .............89 

Heteropodarke ..................81 

heterops, Megalomma .............94 

Heteroserolis...................136 

heteroseta, Bhawania..............86 

heteroseta, Streptospinigera .........83 

Heterospio .....................89 

Heterostropha .................28 

Hexapanopeus .................148 

Hexaplex ......................23 

hians, Argonauta ................35 

hians, Gastrochaena ..............14 

Hiatella .......................14 

Hiatellidae ....................14 

hiatus, Neomegamphopus..........127 

higginsi, Araphura ..............137 

Hinea ........................17 

Hippa .......................145 

Hippidae ....................145 

Hippoidea....................144 

hippocampus, Pterotrachea..........36 

Hippolyte.....................141 

Hippolytidae..................141 

Hippomedon...................130 

Hipponicidae ..................20 

Hipponix ......................20 

Hipponoa......................86 

hirsuta, Taylorpholoe ..............85 

hirsutimana, Acanthaxius..........143 

hirundinina, Chelidonura...........30 

hispidus, Mithrax ...............146 

hispidus, Stenopus...............139 

histrio,Vexillum .................26 

hixoni, Anamaera ...............128 

hobbsi, Fimbriosthenelais ...........85 

hobsonae, Carazziella .............89 

hobsonae, Travisia ................91 

Hobsonia ......................93 

holderi, Mithrax ................146 

Holectypoida .................194 

holmesi, Ampelisca ..............126 

holmesi, Aruga .................130 

holmesi, Parahaustorius ...........129 

holmesii, Cochliolepis .............19 

holmesii, Ophioderma ............193 

holosericus, Pilumnus ............149 

Holothuria ....................195 

Holothuriidae .................195 

Holothuroidea ................195 

holthuisi, Phimochirus ............144 

Homoloidea ..................145 

homosassa, Dissiminassa ..........130 

honduranus, Eudevenopus .........131 

hopkinsi, Heteromysis ............126 

hopkinsi, Mesanthura ............132 

Hoplocarida ..................124 

Hoplopheonoides................128 

Hornellia .....................130 

Horologica .....................22 

horologium, Edwinjoycea ..........137 

hotessieri, Nassarius ..............25 

hotessieriana, Opalia..............23 

hotessieriana, Tegula ..............16 

Houbricka .....................29 

hudsoni, Rhepoxynius ............131 

humilis, Cyclostremella ............29 

humilis, Lepidonopsis .............85 

hummelincki, Augeneriella ..........94 

hummelincki, Metaprotella .........132 

hummi, Paguristes ...............143 

hummi, Polycera .................32 

humphreysii, Epitonium............22 



hunteria, Fasciolaria ..............25 

hutchinsonensis, Eulalia............80 

huxleyi, Filograna ................95 

Hyale .......................129 

Hyalella......................129 

Hyalellidae ...................129 

Hyalidae .....................129 

Hyalina .......................26 

hyalina, Alima .................125 

hyalina, Lyonsia .................15 

hyalina, Syllis ..................83 

Hyalocylis .....................36 

Hyalopale .....................86 

Hyboscolex.....................91 

Hydatina ......................30 

hydriformis, Synaptula ...........196 

Hydrobiidae ...................18 

Hydroides .....................95 

hypacanthus, Lembos.............127 

Hypereteone ....................80 

Hyperphrixus ..................134 

Hypoconcha ...................145 

hypostegus, Opecarcinus ..........150 

Hypselodoris ...................33 

hyptius, Stegophryxus ............133 

Hyssura ......................132 

Hyssuridae ...................132 

I 

Ibacinae .....................143 

ictericus, Spondylus...............10 

idalina, Zafrona .................25 

idiochila, Dentimargo .............26 

Idotea .......................137 

Idoteidae ....................137 

Idoteinae ....................137 

Iliacantha.....................146 

Iliinae .......................146 

Ilyanassa ......................24 

imbricata, Arca...................9 

imbricata, Caribachlamys ...........9 

imbricata, Harmothoe .............84 

imbricata, Pinctada ................9 

imbricatum, Caecum ..............19 

immunda, Geitodoris ..............33 

imogena, Eunice .................87 

impatiens, Holothuria ............195 

impatiens, Lumbrineris ............87 

impexa, Okenia..................32 

impressa, Boonea ................29 

impressa, Ophiolepis .............193 

impressa, Politolana..............135 

impressus, Pagurus ..............144 

Inachinae ....................146 

Inachoides ....................146 

incerta, Alaba ...................17 

incerta, Cabira ..................82 

incerta, Deutella ................132 

incertum, Teinostoma ..............19 

incisa, Houbricka ................29 

incisa, Nephtys ..................84 

inclinata, Atlanta ................36 

incolor, Euchone .................94 

incomptus, Placostegus ............95 

inconspicua, Pleurobranchaea ........32 

inconspicua, Tornatina .............30 

incurvus, Capulus ................20 

indica, Limnoria ................135 

indica, Parasphaerosyllis ...........83 

indicus, Lygdamis ................93 

indicus, Malacoceros ..............89 

indicus, Pisionidens...............86 

inermis, Bemlos spinicarpus ........127 

inermis, Inermonephtys ............84 

Inermonephtys ..................84 

inezae, Pseudochama ..............11 

inflata, Limacina.................36 

inflata, Lumbrineris...............87 

inflata, Pherusa..................91 

inflexa, Cavolinia ................36 

infracarinatus, Solariorbis ..........19 

infratorquata, Novafabricia .........95 

infundibulata, Paranthura .........132 

infundibuliformis, Heliacus .........29 

infundibulum, Latirus .............25 

inglei, Oceanida .................23 

inglei, Trypanosyllis ..............83 

Ingolfiella.....................126 

Ingolfiellidae..................126 

Ingolfiellidea..................126 

Iniforis .......................22 

inomissa, Upogebia ..............143 

inornata, Episcynia ...............19 

inquilina, Parametopella...........131 

inquilinus, Lepidonotus ............85 

insidiosum, Monocorophium........128 

insignis, Dardanus ..............143 

insularis, Alcirona ...............135 

insularis, Rocinela ...............134 

intapurpurea, Puberella ............14 

intastriata, Leporimetis ............12 

integer, Cyclograpsus .............149 

integra, Synsynella ..............134 

integrum, Mysidium .............126 

intermedia, Iliacantha ............146 

intermedia, Lysmata..............141 

intermedia, Melita ...............129 

intermedia, Similiphora ............22 

intermedius, Acanthohaustorius......129 

intermedius, Deflexiodes...........130 

intermedius, Palaemonetes .........140 

interrupta, Turbonilla..............30 

interruptus, Parviturboides ..........19 

intricata, Amphiura ..............193 

intricata, Bailya .................24 

inusta, Olivella ..................26 

invisisacculus, Paguristes ..........144 

io, Cerithiopsis ..................22 

Ioninae ......................134 

iontha, Suturoglypta ..............25 

iota, Cerithiopsis.................22 

Iphigenia ......................13 

Iphimedia ....................129 

Iphimediidae .................129 

irciniae, Colomastix ..............128 

iricolor, Arabella .................88 

iridescens, Periclimenes ...........140 

Iridopagurus...................144 

iris, Tellina .....................12 

irpex, Neohyssura ...............132 

irradians, Argopecten ..............9 

irrasa, Munida .................144 

irregulare, Dendropoma ............20 

irritabilis, Ceratonereis ............84 

irritans, Munidion...............134 

irrorata, Littoraria ................18 

Ischadium ......................8 

Ischnochiton ....................7 

Ischnochitonidae ................7 

Ischyroceridae.................129 

Ischyrocerus ...................130 

Iselica ........................30 

islagrande, Callichirus ............142 

isocanthum, Ophiostigma ..........193 

Isocheles .....................143 

Isognomon......................9 

Isognomonidae .................9 

Isolda ........................93 

isometra, Mellita ................194 

Isopoda......................132 

Isostichopus ...................195 

Ithycythara ....................28 

Iungentitanais..................137 

Ividia ........................29 

J 

jacksoni, Nereis..................84 

jacksoni, Onobops ................18 

jamaicensis, Melanella ............23 

janeirensis, Chaetopleura............7 

janiceae, Colomastix .............128 

Janiridae .....................133 

Janiroidea ....................133 

Janolidae .....................34 

Janolus .......................34 

Janthina ......................23 

janthina, Janthina ................23 

Janthinidae....................23 

Janua ........................96 

januarii, Amphipholis ............193 

Jasmineira .....................94 

jaspidea, Jaspidella ...............26 

Jaspidella......................26 

Jassa ........................130 

jaumei, Diodora .................16 

javanicum, Calliostoma ............16 

jayneae, Haustorius ..............129 

jeannae, Cyclostremiscus ...........19 

jeffreysi, Aricidea ................90 

Joeropsidae...................133 

Joeropsis .....................133 

johnsoni, Caecum ................19 

johnsoni, Poecilochaetus............89 

johnstoni, Thormora ..............85 

jonesi, Ancistrosyllis ..............82 

jonesi, Anopsilana ...............135 

jonesi, Capitella .................92 

jonesi, Lumbrinerides..............87 

Josephella .....................95 

Jouannetia .....................14 

joubini, Octopus .................35 

jugosus, Petrolisthes..............144 

jujubinum, Calliostoma ............16 



Juliacorbula ....................14 

juliana, Aplysia .................31 

Juliidae.......................31 

junonia, Scaphella ...............26 

juvenilis, Clypeasterophilus ........150 

K 

kalafuti, Murexiella...............24 

kalafuti, Quoyula ................24 

kalanii, Neomegamphopus .........127 

Kalliapseudes ..................137 

Kalliapseudidae ...............137 

karinae, Pollia ..................24 

kaumaka, Gammaropsis ...........128 

keenae, Lucina ..................10 

kefersteini, Protodorvillea...........88 

Kefersteinia ....................81 

kempfi, Chromodoris ..............33 

kensleyi, Heteromysis.............126 

kensleyi, Mexicope ..............133 

keraudrenii, Oxygyrus .............36 

killariensis, Tharyx ...............91 

kimallynae,Veleroniopsis ..........140 

kinbergi, Pelogenia ...............85 

Kinbergonuphis .................86 

kingsleyi, Epialtus ...............146 

kingstoni, Hermaea ...............32 

kleidus, Podocerus ...............131 

Kleinella ......................29 

knightjonesi, Janua ...............96 

knorrii,Vermicularia ..............17 

knoxi, Nototeredo ................15 

Kolourione ....................134 

Konatopus ....................127 

koto, Lamellaria .................21 

krebsii, Alcirona ................135 

krebsii, Cymatium corrugatum .......21 

krebsii, Dendrodoris ..............33 

krebsii, Epitonium ................22 

krebsii, Psilaxis..................29 

krebsii, Williamia ................35 

krohnii, Cliopsis .................37 

kroyeri, Spiophanes...............89 

kroyeri, Xiphopenaeus ............139 

kunkelae, Bemlos................127 

Kupellonura ...................132 

Kurtziella......................28 

kyolis, Siraius...................33 

L 

labiosum, Cymatium ..............21 

labis, Alpheopsis ................141 

labyrinthophila, Sphaerosyllis .......83 

lachrimula, Pugnus ...............26 

lactea, Olivella ..................26 

lactea,Volvarina .................26 

lacteus, Pilumnus ...............149 

lacteus, Polinices .................21 

lacunella, Solariella...............16 

lacunosus, Demonax ..............94 

lacustre, Apocorophium ...........128 

lacustris, Panopeus ..............149 

Lacydonia .....................81 

Lacydoniidae ..................81 

Laemodonta ....................35 

Laeonereis .....................84 

Laevicardium ...................12 

laevicauda, Panulirus.............143 

Laevichlamys ...................10 

laevigata, Naineris ...............88 

laevigata, Odostomia ..............29 

laevigata, Pitho .................147 

laevigata, Rhombinella ............25 

laevigata, Sicyonia...............139 

laevigata, Tellina.................12 

laevigata, Zebina ................18 

laevigatum, Laevicardium ..........12 

laevigatum, Macrocoeloma .........146 

lafresnayi, Costoanachis............25 

laguna, Gitanopsis...............126 

Laleonectes....................147 

lamarcki, Carinaria ...............36 

Lamellaria .....................21 

Lamellariidae ..................21 

Lamelleulepethus ................85 

lamelligera, Eusyllis ..............82 

lamellosa, Nereis.................84 

lamellosum, Epitonium ............22 

laminosum, Pedicorophium.........127 

lanceolata, Ithycythara.............28 

lanceolata, Tritonoharpa............27 

Lanice ........................94 

Lanicides ......................94 

Laomediidae..................142 

Laonice .......................89 

lapicida, Petricola ................14 

lappa, Euapta ..................196 

laqueatum, Dentalium..............8 

largidentis, Dorvillea..............88 

largoensis, Nereis ................84 

largoensis, Pagurapseudes..........138 

larvatus, Callinectes..............147 

Lasaea........................11 

Lasaeidae .....................11 

lata, Cerithiopsis.................22 

lata, Parabopyrella ..............133 

lata, Umbilibalcis ................23 

lateralis, Gecarcinus .............149 

lateralis, Mulinia ................12 

lateralis, Musculus ................8 

latericeus, Notomastus .............92 

laticauda, Anilocra ..............135 

Laticorophium .................128 

latilirata, Lirophora...............13 

Latirus........................25 

latispina, Dawsonius .............142 

latreillei, Tylos..................136 

latreilli, Lumbrineris ..............87 

Latreutes .....................141 

latreuticola, Probopyrinella.........133 

latus, Dissodactylus ..............150 

laubieri, Hesionura ...............80 

lauffi, Astacilla .................137 

lavalleeana, Gibberula.............26 

leachii, Bursatella ................31 

Leander ......................140 

Learchis.......................34 

lecroyae, Oxyurostylis ............138 

Leidya .......................134 

Leiocapitella....................92 

Leiochrides.....................92 

Leitoscoloplos ...................88 

Lembos ......................127 

leo, Polygordius .................96 

Leocrates ......................81 

Leodia .......................194 

Leodora .......................96 

Lepidactylis ...................129 

Lepidametria ...................84 

Lepidasthenia...................85 

Lepidochitona ...................7 

Lepidonopsis ...................85 

Lepidonotus ....................85 

Lepidopa .....................145 

Lepidopthalmus ................142 

Leporimetis ....................12 

lepta, Acteocina .................30 

Leptalpheus ...................141 

Leptochela ....................139 

Leptochelia....................137 

Leptocheliidae ................137 

leptodactyla, Uca................150 

Leptodius .....................148 

Leptomysini ..................126 

Leptostraca ...................124 

Leptosynapta ..................196 

leptosynaptae, Pinnixa ............150 

lerema, Teinostoma ...............19 

lesueurii, Atlanta ................36 

lesueurii, Limacina ...............36 

leucocyma, Lucina................10 

leucocyma, Pilsbryspira ............27 

leucohyba, Halosydna .............84 

Leucon.......................138 

Leuconidae ...................138 

Leuconopsis ....................35 

leucophaeata, Mytilopsis ...........13 

leucopleura, Lottia................15 

Leucosiidae....................145 

Leucosioidea..................145 

leucosphaera, Lamellaria ...........21 

leucosphaera, Niveria .............21 

Leucothoe.....................130 

Leucothoidae .................130 

Leucozonia.....................25 

levicula, Murexiella...............24 

Levidorum .....................82 

levifuscina, Podarkeopsis ...........81 

Levinsenia .....................90 

levis, Elasmopus ................129 

levis, Turbonilla .................30 

lewisi, Sigalion ..................85 

lherminieri, Pitho ...............147 

Libinia.......................146 

ligerica, Boccardiella ..............89 

Ligia ........................136 

Ligiidae .....................136 

Ligiamorpha ..................136 

lilacina, Triphora.................22 

Liljeborgia ....................130 

Liljeborgiidae .................130 

Lima..........................9 

lima, Lima ......................9 



Limacina ......................36 

Limacinidae ...................36 

Limaria ........................9 

Limatula .......................9 

limatula, Lucapinella..............16 

limbatus, Uhlias ................145 

limicola, Ophionephthys ..........193 

limicola, Sthenelais ...............85 

Limidae .......................9 

Limnoria .....................135 

Limnoriidae ..................135 

Limoida .......................9 

limonitella, Kurtziella .............28 

limosum, Eurytium ..............148 

limpida, Orobitella ...............11 

limula, Macoma .................12 

Linatella ......................22 

Linckia ......................192 

lindae, Cerithium ................17 

lindae, Skuphonura ..............132 

Lindapecten ....................10 

lineata, Hinea...................17 

lineata, Ophiothrix ..............194 

lineata, Tellina ..................12 

lineatus, Notomastus ..............92 

lineolaris, Argopecten ..............9 

Lioberus .......................8 

Liocuna ......................127 

liodactylus, Iliacantha ............146 

Lioglyphostoma .................28 

lioica, Abra ....................13 

liozonis, Lepidochitona .............7 

liratus, Circulus .................19 

Lirophora......................13 

Lissodrillia.....................27 

listeri, Diodora ..................16 

listeri, Periglypta.................14 

listeri, Tellina ...................12 

Listriella .....................130 

Lithophaga .....................8 

Lithopoma .....................16 

Litiopa .......................17 

Litiopidae.....................17 

Litocorsa ......................82 

Litopenaeus ...................139 

litteratum, Cerithium ..............17 

littoralis, Eurydice ...............135 

Littoraria ......................18 

littorea, Goniada.................81 

Littoridinops ...................18 

Littorinidae ...................18 

lituspalmarum, Teinostoma..........19 

livida, Natica ...................21 

lividomaculata, Tegula .............16 

lividus, Geograpsus ..............149 

Livoneca .....................135 

lobatus, Notomastus ..............92 

lobatus, Speocarcinus.............148 

lobiferum, Megalomma ............94 

Lobiger .......................31 

Lobopilumnus..................148 

locklini, Limaria ..................9 

loevis, Raninoides ...............145 

Loimia........................94 

Loki.........................134 

Loliginidae ....................35 

Loligo ........................35 

Lolliguncula....................35 

Lomanotidae ..................34 

Lomanotus .....................34 

longa, Drilonereis ................88 

longa, Pseudofabriciola ............95 

Longchaeus ....................29 

longicarpus, Pagurus .............144 

longicarpus, Synalpheus...........141 

longicauda, Stylocheilus............31 

longicaudata, Paraclione ...........37 

longicaudata, Photis..............128 

longicaudatus, Periclimenes ........140 

longicirrata, Ceratonereis ...........84 

longicirrata, Glyphohesione .........82 

longicirrata, Parapionosyllis .........83 

longicornis, Americorchestia ........131 

longigulata, Odontosyllis ...........82 

longimana, Ampithoe.............126 

longimana, Chlorodiella ...........148 

longimana, Paraliomera ...........149 

longimerus, Parahaustorius.........129 

longipedis, Munidion.............134 

longipes, Paranebalia .............124 

longipes, Phyllodoce ..............80 

longirostris, Epialtus .............146 

longiseta, Hyboscolex..............91 

longisetis, Eunice ................87 

longisetosa, Melita...............129 

longisignalis, Uca ...............150 

longispinosa, Epimeria ............128 

longisquamosa, Pteria ..............9 

longiventris, Corallianassa .........142 

longleyi, Pilumnus...............149 

longleyi, Tethygeneia .............128 

Longosomidae .................89 

longula, Paramarphysa ............87 

looensis, Mesanthura .............132 

looensis, Microprosthema ..........139 

looensis, Spathiopus .............129 

lopezi, Aricidea ..................90 

Lottia ........................15 

Lottiidae......................15 

louisianae, Taphromysis ...........126 

louisianensis, Lepidophthalmus......142 

louisianum, Apocorophium .........128 

lourei, Exogone ..................82 

Lucapina ......................16 

Lucapinella ....................16 

Lucifer .......................139 

Luciferidae ...................139 

Lucina........................10 

Lucinidae .....................10 

Lucinisca ......................10 

lucunter, Echinometra ............194 

luetkeni, Clypeaster ..............194 

Luidia .......................192 

Luidiidae ....................192 

Lumbricalus ....................87 

lumbricoides, Dasybranchus .........92 

Lumbrineridae .................87 

Lumbrinerides ..................87 

Lumbrineriopsis .................87 

Lumbrineris ....................87 

lunata, Astyris ..................25 

lunulata, Crassinella ..............11 

lunulatus, Dasybranchus ...........92 

lunzi, Pinnixa ..................150 

lupinus, Polyodontes ..............85 

lutea, Syllis ....................83 

luteocrinita, Divariscintilla..........11 

luteola, Parahesione...............81 

lutosum, Cerithium ...............17 

lutzae, Tectasquilla...............125 

lutzeni, Sphaerodoridium ...........81 

Lygdamis ......................93 

lymani, Copidaster ..............192 

lymani, Paguristes ...............144 

lymneiformis, Daphnella ...........28 

lynceus, Phidiana ................34 

lyonsi, Edotia ..................137 

lyonsi, Heteropodarke .............81 

Lyonsia .......................15 

Lyonsiidae ....................15 

lyra, Cirrophorus ................90 

Lyrodus .......................15 

Lysarete.......................88 

Lysianassa ....................130 

Lysianassidae .................130 

Lysianassinae .................130 

Lysianopsis ...................130 

Lysidice .......................87 

Lysilla ........................94 

Lysiosquilla ...................125 

Lysiosquillidae ................125 

Lysiosquilloidea ...............125 

Lysiosquillina ..................125 

Lysippe .......................93 

Lysmata ......................141 

Lytechinus ....................194 

M 

maccraryae, Malmgreniella .........85 

macdonaldi, Notobranchaea .........37 

macerophylla, Chama .............11 

macgintyi, Murexiella .............24 

macgintyi, Olivella ...............26 

mackinneyi, Bemlos ..............127 

Macoma ......................12 

macrobranchia, Eunice.............87 

Macrocallista ...................13 

Macrochaeta ...................91 

macrochira, Pneumodermopsis .......37 

Macroclymene ..................92 

Macrocoeloma .................146 

Macrocypraea...................20 

macrodera, Podochela.............146 

macromana, Photis ..............128 

macromucronatus, Gammarus.......129 

macropus, Octopus ...............35 

Mactridae.....................12 

Mactrotoma ....................12 

maculata, Armandia ..............91 

maculata, Genocidaris ............194 

maculata, Goniada ...............81 

maculata, Lumbrineris .............87 

maculata, Macrocallista ............13 

maculata, Melinna ...............93 



maculatum, Sinum ...............21 

maculatus, Etisus................148 

maculatus, Tumidotheres...........150 

maculosa, Arabella ...............88 

maculosa, Crepidula ..............20 

madagascariensis, Cassis ...........21 

maderiensis, Phyllodoce ............80 

Maera .......................129 

Magelona .....................90 

Magelonida ...................90 

Magelonidae ..................90 

maggae, Pontogenia...............84 

magna, Americonuphis ............86 

magna, Drilonereis ...............88 

magna, Tellina ..................12 

magnavena, Diaphorosoma .........88 

magnidentata, Sphaerosyllis .........83 

magnifica, Amakusanthura .........132 

magnifica, Astropyga .............194 

magnifica, Sabellastarte ............95 

magnum, Trachycardium ...........12 

magnus, Periclimenes.............140 

Majidae .....................146 

Majinae .....................146 

Majoidea ....................146 

major, Callichirus ...............142 

Malacoceros ....................89 

Malacostraca..................124 

Maldane ......................92 

Maldanidae ...................92 

malleator, Alpheus...............141 

Malleidae......................9 

Malleus........................9 

Malmgreniella ..................85 

maltbiana, Pusula ................21 

mammifera, Ephesiella.............81 

Mancocuminae ................138 

manningi, Cinetorhynchus .........140 

manningi, Microprosthema.........139 

manningi, Thor .................141 

mansfieldi, Babelomurex ...........24 

Manucomplanus ................144 

marcusi, Bosellia.................32 

marcuzzii, Tylos.................136 

marenzelleri, Josephella ............95 

margaretae, Daphnella ............28 

margarita, Pontonia ..............140 

margaritaceum, Periploma ..........15 

margaritifera, Pinctada .............9 

marginatus, Cheramus ............142 

Marginellidae ..................26 

marina, Phtisica ................132 

marioni, Aphelochaeta .............90 

Marionia ......................33 

maritima, Polymesoda .............13 

marmoratus, Chiton ...............7 

marochiensis, Natica ..............21 

Marphysa .....................87 

Marsenina .....................21 

Marshallora ....................22 

marshi, Pagurus ................144 

marshi, Pilumnus ...............149 

Martesia ......................14 

martini, Stenoninereis .............84 

martinianum, Cymatium ...........21 

martinicensis, Crassinella...........11 

martinicensis, Tellina..............12 

Mastobranchus ..................92 

Mathilda ......................28 

Mathildidae ...................28 

matthewsae, Epitonium ............22 

Matutinae ....................145 

maua, Catriona..................34 

maugeriae, Hespererato ............21 

maxillulidens, Periclimenaeus .......140 

mayaguezensis, Aonides............89 

mayana, Excirolana ..............135 

mayori, Gadila ...................8 

Mazatlania ....................25 

mcclendoni, Synalpheus ...........141 

mccullochae, Tetrasquilla ..........125 

mcgintyi, Cyphoma ...............20 

mcgintyi, Distorsio constricta ........22 

mcgintyi, Parahyotissa .............10 

mcilhennyi, Eucalliax.............142 

mclaughlinae, Pagurus ............144 

media, Americardia ...............12 

media, Hyale ..................129 

mediolobatus, Lyrodus .............15 

Mediomastus ...................92 

mediterranea, Amphiglena ..........94 

mediterranea, Caliphylla ...........32 

mediterranea, Persephona ..........146 

mediterraneum, Pneumoderma .......37 

medusa, Loimia..................94 

megacephalon, Anuropodione .......134 

megachela, Boca ................127 

Megalobrachium ................144 

Megalomma ....................94 

Megalomphalus .................20 

Megaluropidae ................130 

megastoma, Teinostoma ............19 

megistus, Ensis ..................12 

Meioceras .....................19 

Meiodorvillea ...................88 

Meiosquilla ...................125 

Melampus .....................35 

melanacantha, Cirriformia ..........91 

Melanella .....................23 

melanica, Photis ................128 

melanostigma, Bispira .............94 

melanostoma, Litiopa..............17 

melanura, Cosmotriphora...........22 

meleagris, Nodilittorina ............18 

Melinna ......................93 

Melita .......................129 

Mellita.......................194 

mellitae, Dissodactylus............150 

Mellitidae ....................194 

Melongena.....................24 

Melongenidae .................24 

Melphidippidae ...............130 

Melybia ......................148 

Menippe .....................148 

Meoma ......................194 

mera, Tellina....................12 

mercatoria, Columbella ............25 

Mercenaria ....................13 

mercenaria, Menippe .............148 

mercenaria, Mercenaria ............14 

meredithi, Pectinaria ..............93 

Merhippolyte ..................141 

mericeae, Sergio ................142 

meridionalis, Comactinia ..........192 

Mesanthura ...................132 

Mesochaetopterus ................89 

Mesorhoea ....................147 

mespillum, Nodilittorina ...........18 

meta, Diodora...................16 

Metaceradocus .................130 

Metacirolana ..................135 

metallica, Idotea ................137 

Metalpheus ...................141 

Metamysidopsis ................126 

Metapenaeopsis ................139 

Metaphrixus...................134 

Metaprotella...................132 

Metatiron.....................131 

Metaxia.......................22 

Metharpinia ...................131 

Metharpiniinae ................131 

Metoporhaphis .................146 

metria,Vitricythara ...............28 

mexicana, Allothyone.............195 

mexicana, Bowmaniella ...........125 

mexicana, Chevalia ..............128 

mexicana, Excorallana ............135 

mexicana, Grubeulepis.............85 

mexicana, Holothuria.............195 

mexicana, Pontonia ..............140 

mexicana, Rullierinereis............84 

mexicana, Syllis .................83 

mexicanus, Asclerocheilus ..........91 

Mexicope .....................133 

Mexieulepis ....................85 

mgrayi, Heteroserolis .............136 

miamiensis, Ficopomatus ...........95 

miamiensis, Philoscia.............136 

michaelis, Zygopa ...............145 

Michelea .....................143 

Micheleidae ..................143 

michelini, Encope ...............194 

microcephala, Kinbergonuphis .......86 

Microcerberidae ...............136 

Microcerberidea ...............136 

Microeulima....................23 

Microhedylidae ................31 

Microjassa ....................130 

Micromelo .....................30 

micromma, Nereis ................84 

Micronephtys ...................84 

Micropanope ..................148 

Microphrys ...................146 

Microphthalmus .................81 

microphthalmus, Demonax..........94 

Microprosthema ................139 

Microprotopus .................128 

microps, Nematoscelis ............138 

Microspio .....................89 

Microstelma ....................18 

microtis, Hydroides ...............95 

Microtralia.....................35 

micruropus, Elasocumella ..........138 

midoculi, Notaulax ...............95 

miersii, Armases ................149 



Miesea .......................34 

mildredae, Caribachlamys ...........9 

milleri, Santia..................133 

millsi, Acanthohaustorius ..........129 

mimetica, Bosellia ................32 

minax, Uca....................150 

mindanus, Conus ................28 

mineri, Gnathophylloides ..........140 

minima, Batillaria................17 

minimus, Bemlos................127 

minirosea, Favartia ...............23 

minor, Ensis ....................12 

minor, Fimbriosthenelais ...........85 

minor, Steironepion ...............25 

minor,Vaunthompsonia ...........138 

minus, Synalpheus ..............141 

Minuspio......................89 

minuta, Diodora .................16 

minuta, Hemiaegina .............132 

minuta, Micronephtys .............84 

minuta, Olivella .................26 

minuta, Stenothoe ...............131 

minutimala, Westheidia ............88 

minutus, Carpias................133 

minutus, Planes.................149 

minutus, Pomatoceros .............95 

mirabilis, Ceratonereis .............84 

mirabilis, Coxicerberus............136 

mirabilis, Pedipes ................35 

mirabilis, Pitho .................147 

mirabilis, Strigilla ................12 

miranda, Lacydonia...............81 

mitchelli, Macoma................12 

Mithraculus ...................147 

Mithrax ......................146 

Mithrodia ....................192 

Mithrodiidae..................192 

Mitra ........................26 

Mitrella .......................25 

Mitridae ......................26 

Mitrolumna ....................28 

mobilispinis, Trachypenaeopsis ......139 

Mocosoa .....................146 

modestum, Gnathophyllum.........140 

Modiolus.......................8 

modiolus, Brachidontes .............8 

modiolus, Modiolus................8 

Modulidae ....................17 

Modulus ......................17 

modulus, Modulus................17 

Moira .......................194 

Mollusca ......................7 

Molpadiida ...................195 

monacantha, Thysanopoda .........139 

mongeslopezi, Hydroides ...........95 

monile, Melampus................35 

monilifer, Pachycheles ............144 

monilifera, Olivella ...............26 

moniliferum, Steironepion ..........25 

monilis, Pilsbryspira ..............27 

Monocorophium ................128 

monroensis, Littoridinops ...........18 

Monticellina....................91 

montosa, Edotia.................137 

mooreanus, Solariorbis ............19 

moorei, Ophiophragmus ...........193 

Mooreonuphis ..................86 

Mopaliidae.....................7 

morchii, Chromodoris .............33 

morio, Aplysia ..................31 

morra, Rimosodaphnella ...........28 

morrisoni, Henrya ................29 

morrisoni, Melampus..............35 

morrisoni, Thracia ................15 

mortenseni, Discodoris.............33 

mortenseni, Gigantione ...........134 

mortenseni, Mysidopsis ...........126 

mortenseni, Parabopyrella .........133 

mortenseni, Pseudotanais ..........137 

mortoni, Laevicardium.............12 

Morum .......................26 

moseri, Splendrillia ...............27 

moseri, Tunicotheres..............150 

Mothocya.....................135 

Mourgona .....................32 

mucloc, Janolus..................34 

mucosa, Axiothella ...............92 

mucosa, Phyllodoce ...............80 

mucronatus, Gammarus ...........129 

mucronatus, Hydroides ............95 

muelleri, Tabatzius...............129 

Mulinia .......................12 

multangulus, Cantharus............24 

multibranchiata, Prionospio .........89 

multicostata, Rissoina .............18 

multicostata, Turbonilla ............30 

multicylindri, Eunice ..............87 

multidentata, Arabella .............88 

multidentopsis, Goniada ...........81 

multifidus, Astichopus ............195 

multigranula, Leptosynapta ........196 

multilineata, Astyris ..............25 

multilineata, Parvilucina ...........10 

multispinosa, Pseudovermilia ........95 

multisquamata, Laevichlamys ........10 

multistriatum, Epitonium ...........22 

multiuncinata, Pettibonella ..........88 

munda, Metamysidopsis...........126 

Munida ......................144 

Munidion.....................134 

Munnidae....................133 

Munnogonium .................133 

Murchisonella ..................29 

Murexiella .....................24 

muricata, Lucinisca ...............10 

muricata, Ranilia ...............145 

muricatum, Astrophyton...........193 

muricatum, Trachycardium ..........12 

muricatum,Vasum................24 

muricatus, Cenchritis..............18 

Muricidae.....................23 

muricinum, Cymatium.............21 

Muricopsis.....................24 

mus, Conus ....................28 

muscarum, Cerithium .............17 

muscosus, Lindapecten.............10 

Musculus.......................8 

mutans, Arabella.................88 

mutica, Euphausia ...............138 

mutica, Olivella .................26 

mutica, Pelia...................146 

mutilata, Eunice .................87 

mutilatoides, Eunice ..............87 

myersi, Gibberosus ..............130 

myersi, Paramicrodeutopus .........127 

Myidae.......................14 

Myoida.......................14 

myriacyclum, Eulalia..............80 

Myriochele.....................93 

Mysella .......................11 

Mysida ......................125 

Mysidacea ...................125 

Mysidae .....................125 

Mysidium ....................126 

Mysidopsis....................126 

Mysinae .....................125 

Mysini ......................126 

Mystides ......................80 

Mytilidae ......................8 

Mytiloida ......................8 

Mytilopsis .....................13 

Mytilus ........................8 

N 

naglei, Rudilemboides ............127 

naidina, Exogone ................82 

Naineris ......................88 

Nalicora......................135 

Namalycastis ...................84 

nana, Astarte ...................11 

nana, Mothocya ................135 

Nannastacidae ................138 

Nannodiella ....................28 

Nannosquilla ..................125 

Nannosquillidae ...............125 

Nanobalcis.....................23 

nanus, Cyclopecten ...............10 

nassa, Leucozonia ................25 

Nassariidae ...................24 

Nassarina .....................25 

Nassarius......................24 

nassula, Lucinisca ................10 

Natatolana ....................135 

Natica ........................21 

Naticarius .....................21 

Naticidae .....................21 

nautlae, Depressiscala .............22 

navalis, Teredo ..................15 

Navanax ......................30 

Neanthes ......................84 

neapolitana, Spurilla ..............34 

neapolitanus, Amphilocus..........126 

Nebalia ......................124 

Nebaliidae ...................124 

nebulosa, Eupolymnia .............93 

nebulosa, Littoraria ...............18 

nebulosa, Mooreonuphis ...........86 

neglecta, Gibbesia ...............125 

neglecta, Tuleariocaris ............140 

Nemaster .....................192 

Nematobrachion ................138 

Nematocarcinoidea .............139 

Nematonereis ...................87 

Nematoscelis ..................138 



nematus, Latirus .................25 

Neoamphitrite ..................94 

Neocallichirus..................142 

Neogastropoda .................23 

Neogonodactylus ................124 

Neohaustorius..................129 

Neohyssura ...................132 

Neoleprea .....................94 

Neolipobranchus.................92 

Neoloricata ....................7 

Neomegamphopus ...............127 

Neomysis .....................126 

Neopanope ....................149 

Neopisosoma ..................144 

Neopontonides .................140 

Neosimnia .....................20 

Neotaenioglossa .............17, 36 

Neothyonidium .................195 

neotridens, Diopatra ..............86 

Nephtyidae ...................84 

Nephtys.......................84 

neptuni, Alima .................125 

Nereididae ....................83 

Nereiphylla ....................80 

Nereis ........................84 

Nerilla........................96 

Nerillida......................96 

Nerillidae .....................96 

Nerita ........................17 

Neritidae .....................17 

Neritina.......................17 

Neritopsina ...................17 

Nerocila......................135 

Netamelita ....................129 

nethertoni, Axiorygma ............143 

Neverita ......................21 

nicholsoni, Hippolyte .............141 

nicobaricum, Cymatium ............21 

Nicolea .......................94 

nigra, Lithophaga .................8 

nigripes, Ninoe ..................88 

nigrocincta, Marshallora ...........22 

nigromaculatum, Branchiomma.......94 

Nikoides .....................142 

nimbosa, Macrocallista ............13 

ninetta, Lysidice .................87 

Ninoe ........................88 

Niso .........................23 

nitens, Ervilia...................13 

nitens, Tellina ...................12 

nitida, Euryplax ................148 

nitida, Melita ..................129 

nitida, Nitidella .................25 

Nitidella ......................25 

nitidula, Grubeosyllis .............82 

nitidum, Meioceras ...............19 

nitidus, Clythrocerus .............145 

nivea, Olivella ..................26 

Niveria .......................21 

niveus, Ischnochiton ...............7 

niveus, Triptychus ................29 

niveus, Tylos ...................136 

nobilis, Architectonica .............29 

nobilis, Marphysa ................87 

nodifer, Scyllarides ..............143 

nodifrons, Menippe ..............148 

Nodilittorina ...................18 

Nodipecten.....................10 

nodipes, Macrocoeloma trispinosum...146 

nodosa, Fissurella ................16 

nodosa, Paractaea rufopunctata ......149 

nodosus, Argonauta...............35 

nodosus, Nodipecten ..............10 

nodosus, Speloeophorus ...........145 

nodulosa, Mitra .................26 

nodulosa, Trachypollia .............24 

nodulosus, Coralaxius ............143 

Noetia.........................9 

Noetiidae ......................9 

nonatoi, Costasiella ...............32 

nordmanni, Glycinde..............81 

normanni, Alpheus ..............141 

northropi, Palaemon .............140 

notabilis, Anadara ................9 

Notarchidae ...................31 

Notaspidea....................32 

notata, Diplodonta ...............10 

notata, Lysidice..................87 

Notaulax ......................94 

Notobranchaea ..................37 

Notobranchaeidae ..............37 

Notomastus ....................92 

Notopodinae..................145 

Notopygos .....................86 

Nototanaidae .................137 

Nototanoides ..................137 

Nototeredo .....................15 

nouveli, Heteromysis .............126 

Novafabricia ...................95 

novangliae, Epitonium .............22 

Novaquesta ....................90 

novimundi, Leuconopsis............35 

nucleiformis, Diplodonta ...........10 

nucleus, Argopecten ...............9 

nucleus, Supplanaxis ..............17 

Nucula ........................8 

Nuculana ......................8 

Nuculanidae ...................8 

Nuculidae .....................8 

Nuculoida .....................8 

nuculoides, Semelina ..............13 

Nudibranchia ...............32, 37 

nudicollis, Notaulax ..............95 

nudimanus, Pilumnus ............149 

nuttingi, Alpheus................141 

nuttingi, Micropanope ............148 

nyalya, Chromodoris ..............33 

nycteis, Columbellopsis ............25 

nyei, Hartmanodes...............130 

O 

oba, Catriona ...................34 

obeliscus, Turbonilla ..............30 

obesa, Hoplopheonoides ...........128 

obesa, Parvanachis ...............25 

obesacauda, Paracaudina chilensis ....195 

obesus, Palicus .................150 

obesus, Panopeus................149 

objecta, Aphrodita ................84 

obliquimanus, Hippolyte ..........141 

obliquus, Parahaustorius ..........129 

obscura, Corambe ................32 

obscurus, Ophiodromus ............81 

obscurus, Podarke ................81 

obsoleta, Ilyanassa ...............24 

Obtortionidae..................17 

occidentale, Cymatium.............21 

occidentale, Epitonium.............22 

occidentalis, Dondice ..............34 

occidentalis, Microtralia............35 

occidentalis, Notaulax .............95 

occidentalis, Panopeus ............149 

occidentalis, Parathelges ...........133 

occidentalis, Phyllophorus .........195 

occidentalis, Pseudovermilia .........95 

occidentalis, Solemya ..............8 

Oceanida ......................23 

ocellata, Calappa................145 

ocellata, Leucozonia...............25 

ocellata, Mitrella.................25 

ocellifera, Costasiella ..............32 

Ochlesidae ...................130 

Ocnus .......................195 

octodentata, Dioplosyllis ...........82 

octodentata, Odontosyllis ...........83 

Octopoda .....................35 

Octopodidae...................35 

Octopus .......................35 

octoradiata, Hemitoma.............16 

octotentaculata, Divariscintilla .......11 

oculata, Branchiosyllis .............82 

oculata, Ceratocephale .............83 

oculata, Galathowenia .............93 

oculata, Marphysa belli ............87 

oculata, Onuphis eremita ...........86 

oculata, Pseudosquillisma ..........125 

oculatus, Sabellides ...............93 

oculatus, Spiochaetopterus costarum ...90 

oculeus, Polyodontes ..............85 

Ocypode......................150 

Ocypodidae ..................150 

Ocypodinae ..................150 

Ocypodoidea .................150 

odhneri, Polycera ................32 

Odontodactylidae ..............124 

Odontodactylus.................124 

Odontosyllis....................82 

Odostomia .....................29 

Oedicerotidae .................130 

Oenone .......................88 

Oenonidae ....................88 

oerstedii, Neogonodactylus .........124 

oestrum, Cymothoa ..............135 

Ogyrides .....................142 

Ogyrididae ...................142 

Okenia .......................32 

Olencira......................135 

olfersii, Ligia ..................136 

olgae, Sicyonia .................139 

Oliva.........................26 

olivacea, Cosmotriphora............22 

olivacea, Ophionereis.............193 

Olivella .......................26 

Olividae ......................26 



omissa, Upogebia ...............143 

Onchidella .....................35 

Onchidiidae ...................35 

Oniscidea ....................136 

Oniscoidea ...................136 

oniscus, Morum .................26 

Onobops ......................18 

Onuphidae ....................86 

Onuphis ......................86 

Opalia........................22 

Opaliopsis .....................23 

Opecarcinus ...................150 

operculata, Pomatogebia ...........143 

operculata,Varicorbula.............14 

operculatus, Phimochirus ..........144 

Ophelia .......................91 

Opheliida .....................91 

Opheliidae ....................91 

Ophelina ......................91 

Ophiactidae ..................193 

Ophiactis .....................193 

ophiactoides, Ophiocomella ........193 

Ophidiaster ...................192 

Ophidiasteridae ...............192 

Ophiocnida ...................193 

Ophiocoma....................193 

Ophiocomella ..................193 

Ophiocomidae ................193 

Ophioderma ...................193 

Ophiodermatidae ..............193 

Ophiodromus ...................81 

Ophiolepis ....................193 

Ophiomyxa ...................192 

Ophiomyxidae ................192 

Ophionema ...................193 

Ophionephthys .................193 

Ophionereididae...............193 

Ophionereis ...................193 

Ophiophragmus ................193 

Ophiopsila ....................193 

Ophiostigma...................193 

Ophiothricidae ................194 

Ophiothrix ....................194 

Ophiurida....................193 

Ophiuridae ...................193 

Ophiuroidea ..................192 

Opisthobranchia................30 

Opisthodonta ...................83 

Opistosyllis ....................83 

oplophoroides, Exhippolysmata ......141 

orbicularis, Codakia ..............10 

orbiculata, Codakia ...............10 

orbignyi, Fossarus ................17 

Orbinia .......................88 

Orbiniida .....................88 

Orbiniidae ....................88 

Orchestia .....................131 

Orchomenella ..................130 

Oreaster......................192 

Oreasteridae ..................192 

orientalis, Cymatoica..............12 

Orinella.......................29 

Oriopsis.......................95 

ornata, Amphitrite................93 

ornata, Caribachlamys..............9 

ornata, Elysia ...................32 

ornata, Tridentella ...............135 

ornata, Trigonulina ...............15 

ornata, Triphora .................22 

ornatissima, Cardiomya ............15 

ornatus, Callinectes ..............147 

ornatus, Pachystremiscus ...........19 

ornatus, Phylo ..................88 

Orobitella .....................11 

orstedii, Anamobaea ..............94 

orstedii, Ophiothrix ..............194 

ortezi, Syllis ....................83 

Orthotheres ...................150 

Osachila .....................145 

Ostrea ........................10 

ostrearum, Calotrophon ............23 

ostrearum, Pyrgospira .............27 

ostreicola, Parvanachis.............25 

Ostreidae .....................10 

Ostreoida ......................9 

ostreum, Zaops .................150 

Othilia.......................192 

Ougia ........................88 

Ovalipes .....................147 

ovalipes, Plesiolembos ............127 

ovalis, Anadara ..................9 

ovalis, Cassidinidea ..............136 

ovalis, Livoneca ................135 

ovalis, Pedipes ..................35 

ovata, Gastrochaena ..............14 

overstreeti, Pseudione.............134 

Ovobopyrus ...................133 

ovula, Tralia ....................35 

Ovulidae .....................20 

Owenia .......................93 

Oweniida .....................93 

Oweniidae ....................93 

oxia, Nannodiella ................28 

oxycephala, Glycera ..............81 

oxychone,Vanikoro ...............20 

Oxygyrus......................36 

Oxynoe .......................31 

Oxynoidae ....................31 

oxyophthalmus, Paguristes .........144 

Oxyrhyncha ..................146 

Oxyrhynchaxius ................143 

Oxystomata ..................145 

oxytata, Muricopsis ...............24 

Oxyurostylis...................138 

ozona, Lysianopsis...............130 

P 

pacei, Coralliophila ...............24 

pacei, Cyclothyca ................30 

pacei, Dermomurex ...............23 

pacei, Favartia ..................23 

pacei, Modulus ..................17 

Pachycheles ...................144 

Pachygrapsus ..................149 

Pachystremiscus .................19 

pacifica, Jasmineira ...............94 

pacifica, Pilargis .................82 

packardii, Neopanope ............149 

Pagurapseudes .................138 

Pagurapseudidae ..............138 

Paguridae ....................144 

Paguroidea ...................144 

Paguristes.....................143 

Pagurus ......................144 

Palaeander ....................140 

Palaemon .....................140 

Palaemonetes ..................140 

Palaemonidae .................140 

Palaemoninae .................140 

Palaemonoidea ................140 

Paleanotus .....................86 

Palicidae .....................150 

Palicus.......................150 

Palinurellus ...................143 

Palinura .....................143 

Palinuridae ...................143 

Palinuroidea ..................143 

pallida, Hyalina .................26 

pallida, Janthina .................23 

pallida, Liljeborgia ..............130 

pallida, Mitra ...................26 

pallidior, Leiochrides ..............92 

pallidula, Mooreonuphis ...........86 

palmalitoris, Megalomphalus ........20 

palmata, Pista...................94 

palmeri, Amphiura ..............193 

palmeri, Banareia ...............148 

Palola ........................87 

paludosus, Palaemonetes ..........140 

palustris, Gammarus .............129 

palustris, Littoridinops.............18 

panacea, Uca ..................150 

panamensis, Nereis ...............84 

Pancolinae ...................137 

pandalicola, Probopyrus ...........133 

pandionis, Periclimenes ...........140 

pandionis, Synalpheus ............141 

Pandora.......................15 

Pandoridae ....................15 

pannosus, Pilumnus..............149 

Panopea.......................14 

Panopeus .....................149 

Panoplax .....................148 

pansus, Acanthohaustorius .........129 

Panthalis ......................85 

Panulirus .....................143 

papillata, Diopatra ...............86 

papilligera, Ceratophyllidia .........33 

papillosa, Ancistrosyllis ............82 

papillosa, Elysia .................32 

papillosa, Glycera ................81 

papillosa, Paranaitis ..............80 

papillosa, Scionella ...............94 

papillosus, Ischnochiton.............7 

papulata, Leptochela .............139 

papulosus, Carpoporus............148 

Papyridea .....................12 

papyrium, Amygdalum .............8 

Parabopyrella ..................133 

Parabopyriscus .................133 

Parabornia .....................11 

Paracaprella ...................132 

Paracaudina ...................195 

Paracerceis ....................136 



Paraclione .....................36 

paracrinitus, Alpheus.............141 

Paractaea .....................149 

Paradella .....................136 

paradoxa, Lumbrineriopsis ..........87 

paradoxa, Paralacydonia ...........84 

Parahaustorius .................129 

Parahesione ....................81 

Parahyotissa....................10 

Paralacydonia...................84 

Paralacydoniidae ...............84 

Paralimnoria ..................136 

Paraliomera ...................149 

Paramarphysa ..................87 

paramera, Tellina ................12 

Parametopella..................131 

Paramicrodeutopus ..............127 

Paramphinome ..................86 

Paramunnidae ................133 

Paramya ......................14 

Paranaitis .....................80 

Parandalia .....................82 

Paranebalia ...................124 

paraneptunus, Synalpheus .........141 

Paraninoe .....................87 

Paranthura ....................132 

Paranthuridae .................132 

Paraonidae ....................90 

Paraonis ......................90 

parapacifica, Ampelisca ...........126 

Parapetrolisthes.................144 

Parapinnixa ...................150 

Parapionosyllis ..................83 

Parapleustes ...................131 

Parapleustinae ................131 

Paraprionospio ..................89 

Parapseudidae ................138 

Pararicia ......................88 

Parascolelepis ...................89 

parasitica, Cochliolepis ............19 

parasitica, Hypoconcha ...........145 

Parasphaerosyllis ................83 

Parastarte .....................14 

Paraster ......................194 

Paratanaoidea .................137 

Parathelges....................133 

Paraxiopsis....................143 

Parazeuxo.....................137 

paretii, Albunea ................144 

paretti, Nereiphylla ...............80 

Pareurythoe ....................86 

Parexogone.....................82 

Parhyale .....................129 

Parhyalella....................129 

Pariambidae ..................132 

Pariphinotus ...................130 

parkeri, Bathyporeia .............131 

parkeri, Ithycythara...............28 

parri, Sicyonia .................139 

Parribacus ....................143 

Parthenope ....................147 

parthenopeum, Cymatium ..........21 

Parthenopidae.................147 

Parthenopinae.................147 

Parthenopoidea................147 

parva, Bailya ...................24 

parva, Cirolana.................135 

parva, Erythrops ................125 

Parvanachis ....................25 

parvicallum, Teinostoma............19 

parvicirrus, Sige .................81 

parvidentata, Ampharete ...........93 

parvidentata, Trypanosyllis..........83 

Parvilucina ....................10 

parvimanus, Pseudione ...........134 

Parvimysis ....................126 

parvipatina, Leptosynapta .........196 

parvipedata, Lumbrineris ...........87 

Parviphos......................24 

Parviturbo .....................17 

Parviturboides ..................19 

parvula, Aplysia .................31 

parvula, Holothuria ..............195 

parvulus, Latreutes ..............141 

parvulus, Leptodius ..............148 

parvum, Neothyonidium...........195 

parvus, Hydroides ................95 

parvus, Terebellides ...............93 

Pasiphaeidae..................139 

Pasiphaeoidea.................139 

patae, Conus....................28 

patae, Periclimenes ..............140 

Patellogastropoda ...............15 

Patelloida......................15 

patina, Elysia ...................32 

patinaria, Haliotinella .............21 

patula, Plicopurpura ..............24 

paucibranchiata, Aonides ...........89 

paucibranchiata, Pseudeurythoe ......86 

paucidens, Mesanthura ...........132 

paucidens, Pneumodermopsis ........37 

paucidentata, Lumbrineris ..........87 

paucigranulata, Ophiocoma ........193 

paucispina, Ophiolepis............193 

paucispinus, Echinaster ...........192 

paulae, Pontoniopsis .............140 

paulensis, Hexapanopeus ..........148 

paulensis, Leander...............140 

pauli, Didianema ................16 

pauli, Pleuromalaxis ..............19 

paupercula,Volvulella .............31 

pauperculus, Dermomurex ..........23 

pawsoni, Thyone ................195 

Paxillosida ...................192 

pealeii, Loligo ...................35 

pealii, Conus ...................28 

pearsei, Periclimenaeus ...........140 

pearsei, Pinnixa ................150 

peasei, Alpheus .................141 

pectenicrus, Elasmopus............129 

Pectinaria .....................93 

Pectinariidae...................93 

pectinata, Dorvillea ...............88 

pectinata, Glycymeris ..............9 

pectinata, Lucina.................10 

pectinata, Siphonaria..............35 

pectinella, Codakia ...............10 

Pectinidae .....................9 

pectiniger, Synalpheus ............141 

pedersoni, Periclimenes ...........140 

pedicellatus, Lyrodus ..............15 

Pedicorophium .................127 

pediculus, Niveria ................21 

Pedipes .......................35 

pelagica, Scyllaea ................34 

pelagica, Sunampithoe ............127 

Pelia ........................146 

pellucida, Limaria.................9 

Pelogenia ......................85 

peloronta, Nerita.................17 

Pelseneeria.....................23 

Peltodoris......................33 

Penaeidae ....................139 

Penaeoidea ...................139 

penantis, Caprella ...............132 

Pendanthura...................132 

pennata, Tonna ..................21 

pennigera, Thecacera ..............33 

pensacola, Hippomedon ...........130 

pensylvanica, Lucina ..............10 

pentagonalis, Bellaspira ............27 

pentagonalis, Ithycythara ...........28 

pentagonus, Cyclostremiscus.........19 

Peracarida....................125 

Peracle........................36 

Peraclididae ...................36 

perca, Cuthona ..................34 

Percnon ......................149 

perdido, Orchomenella ............130 

perdidoensis, Cirrophorus...........90 

pergamentaceus, Chaetopterus .......89 

Periclimenaeus .................140 

Periclimenes ...................140 

perieri, Hyale ..................129 

Periglypta .....................14 

Perinereis......................84 

Perioculodes ...................130 

Periploma .....................15 

Periplomatidae .................15 

Peristichia .....................29 

perkinsi, Acanthopale .............86 

perkinsi, Cirrophorus..............90 

perkinsi, Dorvillea ...............88 

perkinsi, Phalacrostemma...........93 

perkinsi, Prionospio...............89 

perkinsi, Sphaerosyllis.............83 

Perkinsiana ....................95 

perlatus, Periclimenaeus ...........140 

permollis, Cryptostrea .............10 

peronii, Atlanta..................36 

perplana, Pteromeris ..............11 

perplexa, Olivella ................26 

perrieri, Heliacus infundibuliformis ....29 

perrugata, Urosalpinx .............24 

perryae, Cerodrillia ...............27 

perryae, Kurtziella................28 

perryae, Periclimenes .............140 

Persephona....................146 

Persicula ......................26 

persimilis,Volvulella ..............31 

personata, Eurydice ..............135 

Personidae ....................22 

perspectivum, Sinum ..............21 

perspicua, Lamellaria .............21 

pervicax, Thyonella ..............195 



Petalifera ......................31 

Petaloconchus...................20 

Petaloproctus ...................92 

petitii, Haminoea ................31 

Petitilla .......................29 

petiverii, Acanthonyx.............146 

petrensis, Eisothistos .............132 

Petricola ......................14 

Petricolaria ....................14 

Petricolidae....................14 

Petrochirus ....................144 

Petrolisthes....................144 

pettiboneae, Levidorum ............82 

pettiboneae, Magelona .............90 

pettiboneae, Spio.................89 

pettiboneae, Streptosyllis ...........83 

Pettiboneia .....................88 

Pettibonella ....................88 

pfefferi, Limnoria ...............135 

Phalacrostemma .................93 

Phalium.......................21 

Pharidae......................12 

pharpa, Doriopsilla ...............33 

phasma, Crassispira ..............27 

Phenacolepadidae ..............17 

Phenacolepas ...................17 

phenax, Macoma.................12 

Pherusa .......................91 

Phidiana ......................34 

philbinae, Aricidea ...............90 

Philine .......................30 

Philinidae.....................30 

Philinopsis.....................30 

philippiana, Anodontia ............10 

philippiana, Lucapina .............16 

Philocheras....................142 

Philoscia .....................136 

Philosciidae ..................136 

Phimochirus ...................144 

Phliantidae ...................130 

phoca, Discodoris ................33 

phoebium, Astralium ..............16 

phoenium, Ophioderma ...........193 

Pholadidae ....................14 

pholadiformis, Petricolaria ..........14 

Pholadomyoida.................15 

Pholas ........................14 

Pholoidae .....................85 

Photis .......................128 

Phoxocephalidae...............131 

Phragmatopoma .................93 

phrixodes, Emarginula.............16 

Phrynophiurida ...............192 

Phtisica ......................132 

Phtisicidae ...................132 

Phtisicinae ...................132 

Phyllaplysia....................31 

Phyllidiidae ...................33 

Phylliroe ......................37 

Phylliroidae ...................37 

phyllisae, Magelona ..............90 

Phyllocarida ..................124 

Phyllodoce .....................80 

Phyllodocida...................80 

Phyllodocidae..................80 

Phyllohartmania.................85 

Phyllophoridae ................195 

Phyllophorus ..................195 

Phylo ........................88 

phymanthi, Epitonium.............22 

physis, Hydatina.................30 

pica, Cittarium ..................16 

Pickfordiateuthis.................35 

Picroceroides...................147 

picta, Hesione ...................81 

picta, Nephtys ..................84 

picta, Synaptocochlea .............16 

picta, Thuridilla .................32 

pictum, Laevicardium .............12 

pictus, Polyophthalmus ............91 

pierceae, Malmgreniella............85 

piercei, Goreopagurus ............144 

pigmentata, Microspio .............89 

pigmentum, Megalomma ...........94 

Pilargidae .....................82 

Pilargis .......................82 

pilata, Cratena ..................34 

Pileolaria......................96 

pillaii, Amphilocus ..............126 

pilosa, Cumella .................138 

pilosus, Mithrax ................146 

pilosus, Pachycheles..............144 

pilsbryi, Antalis ..................8 

pilsbryi, Anticlimax...............19 

pilsbryi, Scaphander ..............30 

Pilsbryspira ....................27 

pilula, Pitar ....................14 

Pilumnoplax...................148 

Pilumnus .....................149 

Pinctada .......................9 

Pinna .........................9 

pinnata, Fiona ..................34 

pinnata, Paraprionospio ............89 

Pinnaxodes....................150 

Pinnidae ......................9 

Pinnixa ......................150 

Pinnothereliinae ...............150 

Pinnotheres ...................150 

Pinnotheridae .................150 

Pinnotherinae .................150 

Pinnotheroidea ................150 

Pionosyllis .....................83 

piperata, Eurydice ...............135 

Pirakia .......................80 

piriferopsis, Sphaerosyllis ..........83 

Piromis .......................91 

Pisania .......................24 

Pisinae ......................146 

Pisione .......................86 

Pisionidae ....................86 

Pisionidens ....................86 

pisonii, Aratus .................149 

Pista .........................94 

pita, Doto......................34 

Pitar .........................14 

Pitho ........................147 

placenta, Cardiapoda..............36 

Placostegus ....................95 

Plagiobrissus ..................194 

Plagusia......................149 

Plagusiinae ...................149 

Plakosyllis .....................83 

plana, Crepidula .................20 

plana, Ophiactis ................193 

Planaxidae ....................17 

Planes .......................149 

planicauda, Cleantioides ..........137 

planispina, Amphiodia............193 

planorbis, Skeneopsis .............18 

planulata, Mysella ...............11 

platensis, Platorchestia............131 

Platorchestia...................131 

Platyactaea....................149 

platycauda, Limnoria .............135 

Platychirograpsus ...............150 

Platycythara....................28 

Platydorididae .................33 

Platydoris .....................33 

Platyischnopidae...............131 

Platylambrus ..................147 

platymerus, Cyclaspis ............138 

Platynereis.....................84 

Platyperona ...................195 

Platypodiella ..................149 

platyproctus, Scyphoproctus .........92 

platyrachis, Pyrgophorus ...........18 

plebeius, Tagelus .................13 

pleei, Acoetes ...................85 

pleii, Bursatella leachii.............31 

pleii, Loligo ....................35 

Pleocyemata ..................139 

Plesiolembos...................127 

pleuracanthus, Hippolyte ..........141 

pleuracanthus, Mithrax ...........146 

Pleurobranchaea .................32 

Pleurobranchidae ...............32 

Pleurobranchus..................32 

Pleurocope ....................133 

Pleurocopidae.................133 

Pleurocrypta ...................134 

Pleuromalaxis...................19 

Pleuromeris ....................11 

Pleuroploca ....................25 

Pleustidae....................131 

plicatella, Raeta .................12 

Plicatula ......................10 

Plicatulidae ...................10 

plicatum, Caecum ................19 

Plicopurpura ...................24 

plicosa, Pyrgocythara..............28 

plumata, Eurysquilla .............124 

plumosus, Polycirrus ..............94 

plurisetis, Gyptis.................81 

Pneumoderma ..................37 

Pneumodermatidae .............37 

Pneumodermopsis ................37 

pocillimanus, Elasmopus ..........129 

Podarkeopsis ...................81 

Podoceridae ..................131 

Podocerus.....................131 

Podochela.....................146 

Pododesmus ....................10 

Poecilochaetidae ................89 

Poecilochaetus ..................89 

poeyi, Megalobrachium ...........144 



poica, Learchis ..................34 

Pokarke .......................81 

Polinices ......................21 

polita, Clio .....................36 

polita, Cyathura ................132 

polita, Politolana ................135 

Politolana.....................135 

politus, Petrolisthes ..............144 

pollex, Ampithoe ................126 

Pollia ........................24 

pollicaris, Pagurus ...............144 

Polybiinae....................147 

Polybranchia ...................32 

Polybranchiidae ................32 

Polycera.......................32 

Polyceratidae ..................32 

Polycerella .....................33 

Polychaeta ....................80 

Polycheria ....................128 

Polycirrus .....................94 

polydentata, Magelona ............90 

Polydora ......................89 

polygnatha, Lumbrineris floridana.....87 

polygonatus, Nassarius ............25 

Polygordiida ...................96 

Polygordiidae ..................96 

Polygordius ....................96 

Polymesoda ....................13 

Polynoidae ....................84 

Polyodontes ....................85 

Polyonyx .....................144 

polyophthalma, Armandia ..........91 

Polyophthalmus .................91 

Polyplacophora..................7 

Polyschides .....................8 

Polystira ......................27 

Pomatoceros ....................95 

Pomatogebia ...................143 

Pomatostegus ...................95 

pomum, Chicoreus................23 

ponderosa, Bursa corrugata..........22 

ponderosa, Casmaria ..............21 

ponderosa, Noetia .................9 

pontifer, Speloeophorus ...........145 

Pontogeneia ...................128 

Pontogenia .....................84 

Pontonia .....................140 

Pontoniinae...................140 

Pontoniopsis ...................140 

Pontoporeiidae ................131 

Poraniella.....................192 

Porcellana ....................144 

Porcellanidae .................144 

Poromya ......................15 

Poromyidae ...................15 

portobelensis, Callistochiton ..........7 

portoricensis, Bowmaniella .........125 

portoricensis, Emerita.............145 

portoricensis, Teredo ..............15 

Portunidae ...................147 

Portuninae ...................147 

Portunoidea ..................147 

Portunus .....................147 

Potamethus ....................95 

Potamididae ...................17 

Potimirim.....................139 

potimirim, Potimirim .............139 

praegustator, Olencira ............135 

praelongus, Euceramus............144 

prampramensis, Trypanosyllis........83 

prasinolineata, Squilla ............125 

prea, Atagema ..................33 

premordica, Kolourione ...........134 

presleyi, Granoturris ..............28 

pretrii, Suturoglypta ..............25 

primitivus, Dissodactylus ..........150 

primitivus, Iungentitanais..........137 

princeps, Holothuria .............195 

princeps, Rissoina ................18 

Prionoglossa....................37 

Prionoplax ....................148 

Prionospio .....................89 

prionurus, Typton ...............140 

Probopyria ....................133 

Probopyrinella .................133 

Probopyrus....................133 

Probythinella ...................18 

Processa......................142 

processae, Urobopyrus ............134 

Processidae...................142 

Processoidea..................142 

proctorae, Glyptaesopus ............28 

producta, Odostomia ..............29 

proficua, Semele .................13 

Progoniada ....................81 

prolifera, Syllis ..................83 

Promysis .....................126 

Propeamussidae ................10 

propinquus, Apseudes ............137 

Propustularia ...................20 

Proscoloplos ....................88 

prostratus, Clypeaster ............194 

Protankyra ....................196 

Protatlanta.....................36 

Protellidae ...................132 

protera, Probythinella .............18 

Proterato ......................21 

protexta, Terebra .................27 

protextus, Ensitellops..............11 

Protodorvillea...................88 

Protodrilida ...................96 

Protohadzia ...................129 

Protohaustorius.................129 

Protomystides...................80 

protracta, Turbonilla ..............30 

Protula .......................95 

protulicola, Hydroides .............95 

provenzanoi, Pagurus .............144 

proxima, Nucula..................8 

proximoculi, Almyracuma..........138 

Prunum .......................26 

Psamathe......................81 

Psammobiidae .................13 

Pseudaeginella .................132 

Pseudeurythoe ..................86 

Pseudione.....................134 

Pseudioninae .................134 

pseudoarticulata, Aricidea ..........90 

Pseudobranchiomma ..............95 

Pseudochama ...................11 

Pseudocheles...................139 

pseudocollaris, Fabricinuda..........94 

Pseudocoutierea ................140 

Pseudocyphoma .................20 

Pseudofabriciola .................95 

pseudofusus, Thyone .............195 

pseudogibba, Euphausia ...........138 

Pseudohaustorius ...............129 

pseudolourei, Exogone .............82 

Pseudomedaeus .................149 

Pseudopolydora .................89 

Pseudopotamilla .................95 

Pseudorhombila ................148 

Pseudoscilla ....................29 

Pseudosquilla ..................125 

Pseudosquillidae...............125 

Pseudosquillisma................125 

Pseudostomatella ................16 

Pseudosyllides ..................83 

Pseudotanaidae ...............137 

Pseudotanainae ...............137 

Pseudotanais ..................137 

Pseudothyone ..................195 

Pseudovermilia ..................95 

pseudovirgatus, Ischnochiton .........7 

psila, Ithycythara ................28 

Psilaxis .......................29 

Pteria .........................9 

Pteriidae ......................9 

Pterioida ......................9 

Pterocirrus .....................81 

pterocladicum, Eulithidium..........16 

Pteromeris .....................11 

Pterotrachea ....................36 

Pterotracheidae ................36 

Pterotyphis.....................24 

Ptilanthura....................132 

pubera, Puberella ................14 

Puberella ......................14 

pudibundus, Hepatus .............145 

puella, Thoe ...................147 

puella,Vexillum .................26 

pugilator, Uca ..................150 

pugilis, Strombus ................20 

pugio, Gnathostenetroides..........133 

pugio, Palaemonetes..............140 

pugnator, Photis ................128 

pugnax, Uca ...................150 

Pugnus .......................26 

pulchella, Amphiodia.............193 

pulchella, Isolda .................93 

pulchella, Pickfordiateuthis..........35 

pulchella, Truncatella..............19 

pulchella, Zafrona................25 

pulchellum, Caecum ..............19 

pulchellum,Vexillum ..............26 

pulchellus, Pachystremiscus .........19 

pulcher, Ophiophragmus ..........193 

pulcherrima, Persicula .............26 

pulchra, Mesanthura .............132 

pulchrum, Calliostoma.............16 

Pulmonata ....................34 

pumila, Emarginula...............16 

pumila, Ophiocoma ..............193 

pumilio, Opalia .................23 



puncta, Turbonilla ................30 

punctata, Cirriformia..............91 

punctata, Diplodonta..............10 

puncticeps, Paguristes ............144 

punctifera, Pirakia................80 

punctostriatus, Rictaxis ............30 

punctulata, Arbacia ..............194 

punicea, Turbonilla ...............30 

puntotorensis, Malmgreniella ........85 

pupa, Horologica.................22 

pupa, Puperita ..................17 

Puperita.......................17 

pupoides, Turbonilla ..............30 

purcina, Discodoris ...............33 

purpurascens, Semele .............13 

pusa, Philinopsis.................30 

pusae, Geitodoris ................33 

pusilla, Micropanope .............148 

pusilla, Munida ................144 

pusilla, Olivella .................26 

pusilla, Paracaprella .............132 

pusilla, Rhodopsis ................95 

pusilla, Tectonatica ...............21 

pusio, Pisania ...................24 

pustulata, Cyclaspis..............138 

pustulata, Panthalis...............85 

pustulata, Patelloida ..............15 

Pusula........................21 

pygmaea, Acanthochitona ...........7 

pygmaea, Apoprionospio ...........89 

pygmaea, Niveria atomaria .........21 

pygmaea, Proterato gemma..........21 

pygmaea, Timoclea ...............14 

pygmaeus, Anisopagurus ..........144 

pygmaeus, Ocnus ...............195 

pygoenigmatica, Paraonis...........90 

pyramidata, Clio.................36 

Pyramidella ....................29 

Pyramidellidae .................29 

Pyrgocythara ...................28 

Pyrgophorus....................18 

Pyrgospira .....................27 

Pyromaia .....................146 

Pyrunculus.....................30 

Pythinella .....................11 

Q 

quadracuta, Eucalliax.............142 

quadrata, Basterotia...............11 

quadrata, Glyphoturris ............28 

quadrata, Ocypode...............150 

quadricincta, Hyboscolex ...........91 

quadridens, Meiosquilla ...........125 

quadridentata, Pitho .............147 

quadridentata, Pseudorhombila ......148 

quadridentata, Sphaeroma .........136 

quadridentatus, Polyschides ..........8 

quadrilineata, Cryoturris ...........28 

quadrilobata, Aricidea .............90 

quadrilobata, Pista ...............94 

quadrimana, Maera ..............129 

quadrioculata, Plakosyllis...........83 

quadripunctata, Niveria ............21 

quadripunctata, Paradella..........136 

quadrispinosa, Diastylis ...........138 

quadrisulcata, Divalinga ...........10 

Questa........................90 

Questidae.....................90 

quillingi, Jouannetia ..............14 

quinquedentatus, Hexapanopeus .....148 

quinqueradia, Ophiactis ...........193 

Quoyula ......................24 

R 

radians, Pseudochama .............11 

radiata, Tellina ..................12 

radiatus, Isognomon ...............9 

Raeta ........................12 

ramiurna, Protankyra.............196 

ramondi, Ampithoe ..............126 

ramosa, Naineris.................88 

ramosa, Petalifera ................31 

randalli, Meiosquilla .............125 

randalli, Phimochirus.............144 

Ranellidae ....................21 

raneyi, Microprotopus ............128 

Rangia .......................12 

Ranilia ......................145 

Raninidae ....................145 

Ranininae ....................145 

Raninoidea ...................145 

Raninoides ....................145 

raninus, Strombus ................20 

rapax, Elasmopus ...............129 

rapax, Hargeria.................137 

rapax, Nalicora .................135 

rapax, Uca ....................150 

rathbunae, Joeropsis..............133 

rathbunae, Lysmata ..............141 

rathbunae, Maera ...............129 

rathbunae, Neocallichirus ..........142 

rathbunae, Periclimenes ...........140 

rathbunae, Synalpheus............141 

rathbunae, Tetraxanthus ...........149 

rathi, Gnathia..................136 

raveneli, Astyris .................25 

raveneli, Euvola .................10 

raveneli, Spisula .................12 

rayrobertsi, Lygdamis .............93 

Recluzia.......................23 

recta, Acteocina..................30 

recta,Volvulella .................31 

rectangulatus, Plesiolembos.........127 

rectifrons, Automate..............141 

recurva, Clio....................36 

recurvum, Ischadium ...............8 

redmanii, Livoneca ..............135 

reductus, Dentimargo..............26 

reductus, Lembos unifasciatus .......127 

reesi, Pleurobranchus..............32 

regalis, Marphysa ................87 

regalis, Pectinaria ................93 

regius, Conus ...................28 

regulare, Caecum ................19 

regularis, Progoniada..............81 

rehderi, Parviturbo ...............17 

remanei, Unela ..................31 

remota, Pisione ..................86 

reniformis, Pseudopotamilla .........95 

resticula, Pyramidella .............29 

reticulata, Cancellaria .............27 

reticulata, Ophionereis ............193 

reticulata, Peracle ................36 

reticulata, Turbonilla ..............30 

reticulatum, Sesarma .............149 

reticulatus, Iridopagurus...........144 

reticulatus, Oreaster..............192 

reticulatus, Scionides ..............94 

retifera, Amaea ..................22 

Retilaskeya ....................22 

retinens, Pinnixa ................150 

Retusa ........................31 

Retusidae .....................30 

reynoldsi, Uromunna .............133 

Rhepoxynius...................131 

Rhithropanopeus................149 

Rhodopsis .....................95 

rhodostoma, Bursa................22 

rhomba, Cyphoma................20 

Rhombinella....................25 

Rhynchocinetidae ..............139 

Rhynchospio ...................89 

Rhyscotidae ..................136 

Rhyscotus.....................136 

rhythmica, Crassispira.............27 

richardbinghami, Favartia ..........23 

richardsonae, Parabopyrella ........133 

ricordi, Armases ................149 

Rictaxis .......................30 

rigens, Cinetorhynchus............140 

rigida, Atrina ....................9 

rigida, Globivenus................13 

riiseanus, Atys ..................31 

riisei, Nereis ....................84 

riisei, Nodilittorina ...............18 

riisei, Ophiophragmus ............193 

riisei, Ophiopsila................193 

riisei, Pachycheles ...............144 

riisei, Podochela ................146 

riisei, Turbonilla .................30 

Rimapenaeus ..................139 

Rimosodaphnella ................28 

Rimula .......................16 

Ringicula ......................30 

Ringiculidae ...................30 

riojai, Magelona .................90 

riseri, Orbinia...................88 

riseri, Sphaerosyllis ...............83 

Rissoella ......................29 

Rissoellidae ...................29 

Rissoidae .....................18 

Rissoina.......................18 

riveroi, Processa ................142 

roberti, Piromis ..................91 

robusta, Glycera .................81 

robustum, Choristodon.............14 

robustum, Dinocardium ............12 

robustus, Leitoscoloplos ............88 

Rocinela......................134 

rolani, Exogone..................82 

rollandiana, Recluzia..............23 

roosevelti, Neomegamphopus .......127 

rosaceus, Clypeaster..............194 



rosaurae, Eunice .................87 

roscidum, Prunum ................26 

rosea, Fissurella .................16 

rosea, Magelona .................90 

roseojugum, Acanthochitona..........7 

roseola, Epitomapta ..............195 

roseolum, Calliostoma .............16 

rosepigmentata, Pileolaria ..........96 

roseus, Hemipodus ...............81 

rostrata, Amphinome ..............86 

rostrata, Galathea ...............144 

rostrata, Poromya ................15 

rostrata, Spengleria ...............14 

rostratipes, Metalpheus ...........141 

rotifera, Chiridota ...............196 

rowei, Holothuria ...............195 

rubella, Rubellatoma ..............28 

Rubellatoma....................28 

ruber, Cronius ..................147 

ruber, Mithraculus...............147 

rubicundum, Ophioderma..........193 

rubidum, Haustellum..............23 

rubiginosus, Nemaster ............192 

rubra, Dorvillea .................88 

rubra, Eunice ...................87 

rubra, Scoloplos .................88 

rubra, Terebella ..................94 

rubrolineata, Godiva ..............34 

rubropoda, Ophiactis .............193 

Rudilemboides .................127 

rudis, Pododesmus................10 

rudolphi, Dorvillea ...............88 

rudolphi, Treptopale...............86 

rufopunctata, Paractaea ...........149 

rugatina, Globivenus ..............13 

rugatus, Clypeasterophilus .........150 

rugimanus, Pachycheles ...........144 

rugirima, Glyphoturris.............28 

rugosa, Squilla .................125 

rugosa, Trachycaris ..............142 

rugosum, Trigonostoma ............27 

rugosus, Panopeus ...............149 

rugulosa, Grubeosyllis .............82 

rugulosa, Metaxia ................22 

Rullierinereis ...................84 

rumia, Cadlina ..................33 

Runcina.......................31 

Runcinidae ....................31 

rupicola, Epitonium...............22 

ruricola, Gecarcinus..............149 

rushii, Terebra...................27 

rustica, Stramonita ...............24 

rusticoides, Columbella ............25 

rycia, Polycera ..................32 

S 

Sabaco........................92 

sabanillaensis, Thyonella ..........195 

Sabellaria .....................93 

Sabellariidae ..................93 

Sabellastarte....................95 

Sabellida .....................94 

Sabellidae ....................94 

Sabellides .....................93 

sabina, Chelidonura ..............30 

Sabinella ......................23 

sabulosa, Hypoconcha ............145 

Saccocirridae ..................96 

Saccocirrus.....................96 

Sacoglossa ....................31 

sagitarius, Mesochaetopterus ........89 

sagra, Philine ...................30 

sagraiana, Rissoina ...............18 

salleana, Hastula ................27 

sallei, Mytilopsis.................13 

Salmoneus ....................141 

salomani, Americorchestia .........131 

salomani, Spilocuma .............138 

salsa, Spurwinkia ................18 

Samythella.....................93 

sanctaecrucis, Hydroides ...........95 

sanctaemarthae, Solecurtus..........13 

sanctithomae, Synalpheus..........141 

sandersoni, Atys .................31 

sanguinea, Eumida ...............80 

sanguinea, Marphysa .............87 

sanguineus, Enoplobranchus.........93 

Sanguinolaria...................13 

sanguinolenta, Sanguinolaria ........13 

sanibelensis, Zonulispira ...........27 

Santia .......................133 

Santiidae ....................133 

sapidus, Callinectes ..............147 

sarda, Chama ...................11 

sardai, Syllis ...................83 

sarsi, Maldane ..................92 

saseboensis, Limnoria ............135 

savagei, Trypanosyllis .............83 

savignyi, Ophiactis ..............193 

sayana, Oliva ...................26 

sayana, Pinnixa ................150 

sayana, Porcellana...............144 

Sayella .......................29 

sayi, Diodora ...................16 

sayi, Dyspanopeus...............148 

sayi, Pilumnus .................149 

sayi, Portunus ..................147 

Sayoscia......................136 

scabra, Ctenoides .................9 

scabricauda, Lysiosquilla ..........125 

scabriuscula, Ophiocnida ..........193 

scalariformis, Cerithidea ...........17 

scalenum, Calliostoma .............17 

Scalibregmatidae ...............91 

Scaphander ....................30 

Scaphella......................25 

scaphobranchia, Amphicteis .........93 

scaphoceris, Synalpheus ...........141 

Scaphopoda ....................7 

schellenbergi, Ampelisca...........126 

Schistomeringos .................88 

Schizasteridae.................194 

Schizobopyrina .................133 

Schizostella ...................193 

schmitti, Alpheus................141 

schmitti, Azygopleon .............134 

schmitti, Meiosquilla .............125 

schmitti, Nannosquilla ............125 

schmitti, Nikoides ...............142 

schmitti, Periclimenaeus ...........140 

schmitzi, Neohaustorius ...........129 

schoenerae, Protohadzia ...........129 

schrammi, Tonicia .................7 

Schwartziella ...................18 

Scionella ......................94 

Scionides ......................94 

Scissurella .....................15 

Scissurellidae ..................15 

Sclerobregma ...................92 

Sclerocheilus ...................92 

Sclerodactyla ..................195 

Sclerodactylidae ...............195 

Sclerostyla .....................95 

Scolelepis......................89 

Scoletoma .....................87 

scolopendra, Coronis .............125 

Scoloplos ......................88 

sculptipes, Micropanope...........148 

sculptus, Mithraculus.............147 

scutata, Pterotrachea ..............36 

scutata, Sternaspis................92 

scutellatus, Stenopus .............139 

Scyllaea.......................34 

Scyllaeidae ....................34 

Scyllaridae ...................143 

Scyllarinae ...................143 

Scyllarides ....................143 

Scyllarus .....................143 

Scyphacella ...................136 

Scyphacidae ..................136 

Scyphoproctus ..................92 

sczelkowii, Microphthalmus .........81 

Seba ........................131 

sebae, Portunus .................147 

Sebidae......................131 

seclusus, Pariphinotus ............130 

sedlaki, Cyphoma ................20 

sedna, Glossodoris................33 

seguroensis, Duasmodactyla ........195 

Seila .........................22 

Semele........................13 

Semelidae ....................13 

Semelina ......................13 

semiaspera, Diplodonta ............10 

semiferruginea, Mitra .............26 

semilaeve, Microprosthema.........139 

semilevis, Osachila ..............145 

seminuda, Atrina .................9 

seminuda, Boonea ................29 

semiplicata, Costoanachis...........25 

semipunctus, Solariorbis ...........19 

Semirossia .....................35 

semisculptus, Circulus .............19 

semistriata, Ringicula .............30 

semistriolata, Graptacme ............8 

semisulcata, Papyridea.............12 

semisulcata, Sigatica ..............21 

Semperothuria .................195 

senegalensis, Luidia ..............192 

senex, Aspella...................23 

sennottorum, Conus...............28 

sentis, Caribachlamys ..............9 

sentus, Echinaster ...............192 

Sepioidea .....................35 



sepioidea, Sepioteuthis .............35 

Sepiolidae ....................35 

Sepioteuthis ....................35 

septemspinosum, Macrocoeloma .....146 

septemspinosus, Aepinus ..........146 

septus, Ophiophragmus ...........193 

sepultus, Amphioplus.............193 

serca, Elysia ....................32 

Sergestidae ...................139 

Sergestoidea ..................139 

Sergio .......................142 

sericeus, Paguristes ..............144 

sericoma, Pontogenia ..............84 

Serolidae ....................136 

Serpula .......................95 

Serpulidae ....................95 

Serpulorbis ....................20 

serrata, Atrina ...................9 

serrata, Cumella ................138 

serrata, Parthenope ..............147 

serrata, Unciola.................127 

serratifrons, Axiopsis .............143 

serratirostris, Discias .............139 

serratorbita, Leptochela ...........139 

serratum, Stenetrium .............133 

serratum, Tozeuma ...............142 

serratus, Ctenodrilus ..............91 

serrei, Pugnus ...................26 

Sesarma......................149 

Sesarminae...................149 

seticornis, Stenorhynchus ..........146 

setiferus, Litopenaeus .............139 

setigera, Platyactaea..............149 

setimanus, Gilvossius.............142 

setosa, Chaetozone ...............90 

setosa, Naineris..................88 

setosa, Protula ..................95 

setosa, Spio ....................89 

setosus, Bemlos .................127 

setosus, Thelepus.................94 

sewelli, Bowmaniella .............125 

sexiesperforata, Leodia ............194 

sexspinosa, Mesorhoea ............147 

sexticornis, Excorallana ...........135 

sheardi, Ceradocus...............128 

Shoemakerella .................130 

shoemakeri, Acanthohaustorius ......129 

shoemakeri, Ceradocus ............128 

shoemakeri, Microprotopus .........128 

shoemakeri, Pinnotheres ...........150 

shuttleworthianus, Callistochiton ......7 

sica, Palicus ...................150 

siciliensis, Palola.................87 

Sicyonia......................139 

Sicyoniidae ...................139 

sidneyi, Podochela ...............146 

Sigalion.......................85 

Sigalionidae ...................85 

Sigambra ......................82 

Sigatica .......................21 

Sige .........................81 

signata, Amakusanthura...........132 

signata, Rocinela ................135 

signatum, Cyphoma ..............20 

sigsbeiana, Porcellana ............144 

Siliquariidae...................18 

simile, Apocorophium ............128 

Similiphora ....................22 

similis, Callinectes...............147 

similis, Costoanachis ..............25 

similis, Rimapenaeus .............139 

similis, Tellina ..................12 

Simnialena ....................20 

simoni, Kinbergonuphis ............86 

simoni, Nephtys .................84 

simplex, Anomia .................10 

simpsoni, Lissodrillia..............27 

simpsoni, Panopeus ..............149 

simpsoni, Pitar ..................14 

simulata, Limnoria ..............135 

simus, Thunor ..................141 

singularis, Ceratonereis ............84 

sinistrum, Busycon ...............24 

Sinum ........................21 

sinuosa, Chama .................11 

Siphonaria.....................35 

Siphonariidae..................35 

Siraius........................33 

Siriella.......................125 

Siriellinae ....................125 

siva, Ophiostigma ...............193 

Skeneidae.....................17 

Skeneopsidae ..................18 

Skeneopsis .....................18 

Skuphonura ...................132 

smaragda, Phyllaplysia ............31 

Smaragdia .....................17 

smithi, Globosolembos ............127 

smithi, Metapenaeopsis ...........139 

smithi, Oxyurostylis..............138 

smithii, Axelella .................27 

smithii, Carditopsis ...............11 

smithii, Diplothyra ...............14 

smithii, Glyptoplax ..............148 

sociabilis, Dorvillea...............88 

socialis, Dipolydora...............89 

socialis, Petaloproctus .............92 

sofla, Pseudofabriciola .............95 

Solariella ......................16 

Solariorbis .....................19 

Solecurtidae ...................13 

Solecurtus .....................13 

Solemya .......................8 

Solemyidae ....................8 

Solemyoida ....................8 

Solen.........................12 

Solenidae .....................12 

soleniformis, Papyridea ............12 

Solenocera ....................139 

Solenoceridae .................139 

solida, Bulla ....................31 

solida, Euthyonacta ..............195 

solida, Strictispira ................27 

solitaria, Glycinde................81 

sombrerensis, Lucina ..............10 

somersi, Odostomia ...............29 

soriatum, Megalobrachium .........144 

soror, Diplodonta ................11 

souleyeti, Protatlanta ..............36 

souverbii, Lobiger ................31 

sowerbii, Lucapina ...............16 

sowerbyi, Episiphon ...............8 

sparsa, Costoanachis ..............25 

sparsa, Iliacantha ...............146 

spartinophila, Uhlorchestia.........132 

Spatangoida ..................194 

spatha, Spathoteredo ..............15 

Spathiopus ....................129 

Spathochlamys ..................10 

Spathoteredo ...................15 

spatula, Drilonereis ...............88 

speciosa, Eucrassatella .............11 

speciosa, Paranaitis ...............80 

speciosa, Uca ..................150 

spectabilis, Platychirograpsus .......150 

spectabilis, Platypodiella ..........149 

spectralis, Glycymeris ..............9 

Speloeophorus .................145 

Spengleria .....................14 

Speocarcinus ..................148 

Sphaerodoridae ................81 

Sphaerodoridium ................81 

Sphaeroma ....................136 

Sphaeromatidae ...............136 

Sphaeromatinae ...............136 

sphaeromiformis, Metacirolana ......135 

Sphaerosyllis ...................83 

Sphenia .......................14 

sphinctostoma, Texadina............18 

sphyrabrancha, Glycera ............81 

Spilocuma ....................138 

spinata, Arcturella ...............137 

spinicarpa, Leucothoe.............130 

spinicarpa, Uca .................150 

spinicarpus, Bemlos ..............127 

spinicarpus, Portunus.............147 

spinimanus, Portunus.............147 

spinipes, Garthiope ..............148 

spinipleura, Paraxiopsis ...........143 

spinisetosa, Pionosyllis ............83 

spinistipula, Upogebia ............143 

spinosa, Dioptromysis ............126 

spinosissima, Hypoconcha .........145 

spinosissimus, Mithrax ...........146 

spinosissimus, Pilumnus...........149 

spinosus, Coelocerus .............146 

spinosus, Eobrolgus ..............131 

Spinulosida...................192 

spinulosus, Echinaster ............192 

spinulosus, Manucomplanus ........144 

spinulosus, Neogonodactylus........124 

Spio .........................89 

Spiochaetopterus.................90 

Spionida......................88 

Spionidae .....................88 

Spiophanes ....................89 

Spiraserpula....................95 

spirata,Vermicularia ..............18 

spiratus, Busycotypus .............24 

Spirobranchus ..................95 

Spirolaxis .....................29 

Spirorbidae ...................95 

Spirula .......................35 

spirula, Spirula..................35 

Spirulidae ....................35 



Spisula .......................12 

Splendrillia ....................27 

Spondylidae ...................10 

Spondylus .....................10 

spongicola, Eunice................87 

spongicola, Haplosyllis ............82 

spongicola, Hydroides .............95 

Spongicolidae .................139 

spongiphila, Syllis................83 

Sportellidae ...................11 

sprucecreekensis, Melongena ........24 

Spurilla .......................34 

spurius, Conus ..................28 

Spurwinkia ....................18 

squalida, Ceratozona ...............7 

squamata, Amphipholis ...........193 

squamata, Scolelepis ..............89 

squamatus, Tenagodus .............18 

squamifera, Tellina ...............12 

squamosa, Nephtys ...............84 

squamosissimum, Ophioderma ......193 

squamosus, Modiolus modiolus........8 

squamulosa, Ophionereis ..........193 

Squilla.......................125 

Squillidae ....................125 

Squilloidea ...................125 

squillina, Parabornia ..............11 

stancyki, Ersilia .................23 

staphylophera, Chaetopleura .........7 

starcki, Paguristes ...............144 

stearnsii, Aesopus ................25 

stearnsii, Conus .................28 

stebbingi, Clypeasterophilus ........150 

stebbingi, Hansenium.............133 

steenstrupi, Prionospio.............89 

Stegophryxus ..................133 

Steironepion....................25 

stellata, Berthella ................32 

stellata, Stellatoma ...............28 

Stellatoma .....................28 

stellatus, Parabopyriscus...........133 

stellatus, Pomatostegus.............95 

Stenetriidae ..................133 

Stenetrioidea .................133 

Stenetrium ....................133 

stenocerum, Sclerobregma...........92 

Stenocionops ..................147 

Stenoninereis ...................84 

Stenoplax ......................7 

Stenopleustes ..................131 

Stenopleustinae ...............131 

Stenopodidae .................139 

Stenopodidea .................139 

Stenopus .....................139 

Stenorhynchus .................146 

Stenothoe .....................131 

Stenothoidae..................131 

stephensoni, Ovalipes ............147 

Sternaspida ...................92 

Sternaspidae ..................92 

Sternaspis .....................92 

steueri, Janua ...................96 

Sthenelais .....................85 

Sthenolepis ....................85 

Stichopodidae.................195 

stigmatura, Eunice................87 

Stigmaulax.....................21 

Stiliger .......................32 

Stiligeridae....................32 

stimpsoni, Conus.................28 

stimpsoni, Pagurus...............144 

stimpsoni, Porcellana .............144 

stimpsoni, Transennella ............14 

stimpsonii, Amphiura ............193 

stimpsonii, Ebalia ...............145 

stimpsonii, Pelseneeria.............23 

Stolus .......................195 

Stomatopoda .................124 

Stosicia .......................18 

straminea, Mitra .................26 

Stramonita.....................24 

Streblosoma ....................94 

Streblospio .....................89 

Streptospinigera .................83 

Streptosyllis ....................83 

striata, Bulla....................31 

striata, Cochliolepis ...............19 

striata, Hyalocylis ................36 

striata, Martesia .................14 

striatissimum, Epitonium ...........22 

striatocostata, Rissoina.............18 

striatoides, Acanthotrophon .........23 

striatum, Crucibulum..............20 

Strictispira .....................27 

Strictispiridae ..................27 

Strigilla .......................12 

strigosum, Caecum ...............19 

striolatus, Ischnochiton .............7 

striosa, Rissoina .................18 

stroemi, Terebellides...............93 

strombi, Orthotheres .............150 

Strombidae....................20 

Strombus ......................20 

stucki, Americamysis .............126 

styliformis, Careliopsis ............29 

Styliola .......................36 

Stylocheilus ....................31 

Stylocheiron ...................138 

Stylocidaris ...................194 

subauriculata, Limatula .............9 

subcarinata, Eulimostraca ..........23 

subcaudalis, Eophrixus............134 

subdepressus, Clypeaster ..........194 

subglobosa, Diplodonta ............11 

subglobosa, Iliacantha ............146 

sublaevis, Aegires ................32 

sublevis, Lepidonotus..............85 

subornata, Elysia ................32 

subovata, Paramya ...............14 

subparallelum, Macrocoeloma.......146 

subrufus, Hipponix ...............20 

subtriplicata,Volvarina ............26 

subula, Styliola..................36 

subvolutum, Caecum ..............19 

succinea, Assiminea...............19 

succinea, Creedonia ...............34 

succinea, Haminoea...............31 

succinea, Nereis .................84 

succinea, Prunum ................26 

suecica, Aricidea .................90 

suensonii, Ophiothrix.............194 

suffusa, Lucapina ................16 

suffusa, Niveria .................21 

suhmii, Stylocheiron .............139 

sulcata, Calappa ................145 

sulcata, Lysidice .................87 

sulcata, Retusa ..................31 

sulcatus, Stigmaulax ..............21 

sulcidentata, Eupleura .............23 

sulcipes, Dactylokepon............134 

Sunampithoe ..................127 

Supplanaxis ....................17 

suppressus, Cyclostremiscus .........19 

surinamensis, Holothuria ..........195 

surinamensis, Ocnus .............195 

surinamensis, Propustularia .........20 

surinamensis, Strigilla .............12 

suspectus, Ocnus ................195 

sutherlandi, Gammaropsis .........128 

Suturoglypta ...................25 

swedmarki, Grubeosyllis ...........82 

swifti, Haplocochlias ..............17 

swifti, Metamysidopsis............126 

swiftiana, Corbula................14 

sybaritica, Tellina ................12 

sybariticum, Laevicardium ..........12 

sycilla, Hypselodoris ..............33 

sykesi,Vexillum..................26 

Syllidae ......................82 

Syllides .......................83 

Syllidia .......................82 

Syllis.........................83 

symbiotica, Stenothoe.............131 

Symethinae...................145 

Symethis .....................145 

symmetricus, Ambidexter ..........142 

synalphei, Bopyrione .............133 

synalphei, Hemiarthrus ...........134 

Synalpheion ...................134 

Synalpheus ...................141 

Synaptidae ...................195 

Synaptocochlea..................16 

Synaptula ....................196 

Synaxidae ....................143 

Synelmis ......................82 

Synopia ......................131 

Synopiidae ...................131 

Synsynella ....................133 

Systellommatophora.............35 

T 

Tabatzius .....................129 

taboguillae, Lanicides .............94 

taeniata, Thormora ...............85 

taeniolata, Metaxia ...............22 

Tagelus .......................13 

Talitridae ....................131 

Talparia.......................20 

talpoida, Emerita................145 

tampaense, Calliostoma ............17 

tampaensis, Crassispira ............27 

tampaensis, Murchisonella ..........29 

tampaensis, Tellina ...............12 

tampaensis, Urosalpinx ............24 



Tanaidacea ...................137 

Tanaidae .....................137 

Tanaidomorpha................137 

tanaiformis, Pendanthura ..........132 

Tanaoidea ....................137 

taphrium, Antalis .................8 

Taphromysis ...................126 

Taringa .......................33 

taurina, Terebra..................27 

taylorae, Zafrona ................25 

taylori, Aricidea .................90 

taylori, Malmgreniella .............85 

taylori, Mesochaetopterus...........89 

taylori, Nannosquilla .............125 

taylori, Phyllohartmania ...........85 

taylori, Sphaerosyllis..............83 

Taylorpholoe ...................85 

Tectarius ......................18 

Tectasquilla ...................125 

Tectonatica.....................21 

tectum, Lithopoma................16 

tedbayeri, Natica.................21 

tedi, Marionia...................33 

Tegula ........................16 

Teinostoma.....................19 

Teleotanais ....................137 

Tellidora ......................12 

Tellina........................12 

tellinae, Fabia ..................150 

Tellinidae .....................12 

tellinoides, Cumingia..............13 

telopia, Taringa ..................33 

temir, Paracaprella...............132 

Temnopleuridae ...............194 

Temnopleuroida ...............194 

tempus, Bemlos .................127 

Tenagodus .....................18 

Tenaturris .....................28 

tenella, Nematoscelis .............138 

tenella, Tellina ..................12 

Tenellia .......................34 

tenera, Barbatia ..................9 

tenera, Ctenoides .................9 

tenera, Euphausia ...............138 

tenera, Semirossia ................35 

tenerum, Periploma ...............15 

tenerum, Trigonostoma .............27 

tenta, Macoma ..................12 

tentaculata, Cirriformia ............91 

tentaculata, Sigambra .............82 

tenuicornis, Leander..............140 

tenuidentis, Ougia................88 

tenuiliratum, Cymatium............21 

tenuipes, Littoridinops .............18 

tenuis, Cyclinella ................13 

tenuis, Eunice ...................87 

tenuis, Leptosynapta .............196 

tenuis, Lumbrineris ...............87 

tenuis, Mellita..................194 

tenuis, Notomastus ...............92 

tenuis, Paracaprella ..............132 

tenuis, Ptilanthura...............132 

tequestae, Hornellia ..............130 

Teralatirus .....................25 

Terebella ......................94 

Terebellida ....................93 

Terebellidae ...................93 

Terebellides ....................93 

Terebra .......................27 

terebrans, Chelura ...............127 

terebrans, Sphaeroma.............136 

Terebridae ....................27 

Teredinidae ...................15 

Teredo ........................15 

teres, Eulimastoma ...............29 

teres, Goniada ..................81 

teres, Notomastus ................92 

Tergipedidae ..................34 

terminalis, Solariorbis .............19 

Teskeyostrea ....................10 

tesselata, Glycera ................81 

tessellata, Littoraria...............18 

tessellata, Nerita .................17 

tessellatus, Goniaster .............192 

testacea, Colubraria ...............24 

testiculus, Cypraecassis ............21 

testudinaria, Hippa ..............145 

testudinum, Lumbrineris ...........87 

Tethorchestia ..................131 

Tethygeneia ...................128 

tetrabranchia, Dipolydora ..........89 

tetrabranchiata, Prionoglossa ........37 

tetradonta, Microjassa ............130 

Tetrasquilla ...................125 

Tetrasquillidae ................125 

tetraura, Lumbrineris..............87 

Tetraxanthus...................149 

tetrodon, Polyschides ...............8 

Teuthoidea ....................35 

Texadina ......................18 

texana, Dyspanopeus .............148 

texana, Scolelepis ................89 

texana, Scoloplos.................88 

texana, Tellina ..................12 

texanus, Circulus ................19 

texasiana, Aligena................11 

texasianum, Agriopoma ............13 

texasianus, Donax................13 

texensis, Parametopella ...........131 

texensis, Rhyscotus ..............136 

textile, Caecum ..................19 

textilis, Turbonilla ................30 

Thais.........................24 

Thala ........................26 

thalamita, Melybia ..............148 

thalassicola, Eulithidium ...........16 

Thalassinidea .................142 

Thalenessa .....................85 

Tharyx .......................91 

thayeri, Uca ...................150 

thea, Cerodrillia .................27 

Thecacera .....................33 

Thecosomata ................31, 36 

Theelothuria ...................195 

Thelecythara ...................28 

Thelepus ......................94 

Therochaeta ....................91 

Thliptodon .....................37 

Thliptodontidae ................37 

Thoe ........................147 

thomae, Bursa rhodostoma ..........22 

thomasi, Alpheus................141 

thomasi, Holothuria..............195 

thomasi, Orchomenella............130 

thompsoni, Akera ................31 

thompsonii, Siriella ..............125 

Thor ........................141 

thorii, Bopyrinella ...............133 

Thormora......................85 

Thracia .......................15 

Thraciidae ....................15 

thrombodes, Amphioplus ..........193 

Thunor ......................141 

Thuridilla .....................32 

Thymiosycia ...................195 

Thyone ......................195 

Thyonella.....................195 

Thysanopoda .................139 

tibicen, Calcinus ................143 

tigrinoides, Syllis ................83 

tigrinus, Bemlos ................127 

tigrinus, Gammarus..............129 

timessa, Heilprinia ...............25 

Timoclea ......................14 

tina, Cuthona ...................34 

tincta, Barleeia ..................18 

tincta, Pollia ....................24 

Tivela ........................14 

tollini, Epitonium ................22 

Tomura .......................28 

Tonicellidae ....................7 

Tonicia ........................7 

Tonna ........................21 

Tonnidae .....................21 

toreta, Peristichia ................29 

Tornatina......................30 

toroensis, Chrysallida .............29 

torquata, Clymenella ..............92 

tortugae, Calappa ...............145 

tortugae, Gitanopsis..............126 

tortugae, Lysidice ................87 

tortugae, Mithrax ...............147 

tortugae, Paguristes ..............144 

tortugae, Phyllodoce ..............80 

tortugae, Typton.................140 

tortugaenis, Cirriformia ............91 

tortugaensis, Syllis ...............83 

tortuganum, Cyclostrema ...........16 

tortugensis, Parapetrolisthes ........144 

tortuosa, Pseudopotamilla...........95 

torus, Neogonodactylus ...........124 

townsendi, Synalpheus............141 

Toxopneustidae................194 

toyatani, Turbonilla ...............30 

Tozeuma......................141 

Trachycardium ..................12 

Trachycaris....................142 

Trachypenaeopsis ...............139 

Trachypollia ....................24 

Tralia ........................35 

Transennella ...................14 

transversa, Anadara ...............9 

transversus, Pachygrapsus .........149 

Trapania ......................32 

Trapezidae ....................13 



Trapezioplax...................148 

trapherus, Photis ................128 

Travisia .......................91 

Treptopale .....................86 

triacantha, Peracle................36 

triangularis, Pterotyphis............24 

triangulatus, Paguristes ...........144 

tribuloides, Eucidaris .............194 

tricarina, Ptilanthura .............132 

tricarinata, Arene ................16 

tricarinata, Meiosquilla ...........125 

Trichobranchidae ...............93 

Trichobranchus ..................93 

tricolor, Clibanarius ..............143 

tricornis, Excorallana .............135 

tricuspida, Thysanopoda...........139 

tricuspis, Parandalia ..............82 

tridentata, Cavolinia ..............36 

tridentata, Colomastix ............128 

tridentata, Diopatra...............86 

tridentata, Pleuromeris.............11 

tridentata, Trapezioplax ...........148 

tridentata, Websterinereis ...........84 

Tridentellidae .................135 

trifurcata, Novaquesta .............90 

trifurcatus, Nototanoides ..........137 

Trigonostoma ...................27 

Trigonulina ....................15 

trilineata, Pandora................15 

triloba, Edotia ..................137 

triloba, Euphrosine ...............86 

trilobata, Amaeana ...............93 

trilobata, Aricidea ................90 

trilobata, Fabricinuda..............94 

trilobatus, Sergio ................142 

Trimusculidae..................35 

Trimusculus ....................35 

triocellatus, Metatiron ............131 

Triphora ......................22 

Triphoridae....................22 

Tripneustes ....................194 

Triptychus .....................29 

tripunctata, Cumella .............138 

tripunctata, Limnoria .............136 

triquetra, Parastarte ...............14 

trispinosa, Diacria................36 

trispinosum, Macrocoeloma ........146 

trispinosus, Alpheopsis............141 

trisulcata, Lucina ................10 

trita, Euthyonidiella..............195 

Tritonia .......................33 

Tritonidoxa ....................33 

Tritoniidae ....................33 

Tritoniopsis ....................33 

tritonis, Charonia ................21 

Tritonoharpa ...................27 

Triviidae ......................20 

trivittata, Ilyanassa ...............24 

Trochidae .....................16 

trochiformis, Limacina .............36 

Troglocarcinus .................150 

troglodytes, Divariscintilla ..........11 

troglodytes, Sabinella .............23 

tropakis, Metatiron ..............131 

tropica, Seba...................131 

tropicalis, Elthusa ...............135 

tropicalis, Hanleya ................7 

Tropichelura ...................127 

truncata, Barnea .................14 

Truncatella.....................19 

Truncatellidae..................18 

trychna, Amphiodia ..............193 

Trypanosyllis ...................83 

Tryphosinae ..................130 

tuber, Lithopoma.................16 

tuberatum, Cyclozodion ...........145 

tuberculata, Eurysyllis .............82 

tuberculata, Limnoria.............136 

tuberculatum, Monocorophium ......128 

tuberculatus, Chiton ...............7 

tuberosa, Cassis .................21 

tubifex, Euarche .................85 

tubularis, Cerapus ...............130 

tubularis, Picroceroides............147 

tuca, Elysia ....................32 

Tuleariocaris...................140 

tulipa, Fasciolaria ................25 

Tumidotheres ..................150 

tumidulus, Cronius ..............147 

Tunicotheres ...................150 

turbinella, Engina ................24 

Turbinellidae ..................24 

Turbinidae ....................16 

Turbo ........................16 

Turbonilla .....................29 

turgidus, Eurypanopeus ...........148 

turricula, Trachypollia .............24 

Turridae ......................27 

turristhomae, Iniforis ..............22 

Turritella ......................17 

Turritellidae ...................17 

Tyche ........................147 

Tychinae .....................147 

Tylidae ......................136 

Tylomorpha...................136 

Tylos ........................136 

typica, Anchialina ...............125 

typica, Sicyonia.................139 

Typosyllis .....................83 

Typton .......................140 

U 

uberinus, Polinices ...............21 

uberlackerae, Gigantione ..........134 

Uca .........................150 

Ucides .......................150 

uebelackerae, Parapionosyllis ........83 

uhleri, Uhlorchestia ..............132 

Uhlias .......................145 

Uhlorchestia ...................132 

ulla, Ascobulla ..................31 

ultramarina, Synopia .............131 

Umbilibalcis....................23 

Umbraculidae..................32 

Umbraculum ...................32 

umbraculum, Umbraculum..........32 

uncinata, Cavolinia ...............36 

uncinata, Dispio .................89 

uncinus, Acanthohaustorius ........129 

Unciola ......................127 

undata, Glycymeris ................9 

undatus, Micromelo...............30 

underwoodae, Graphis .............23 

Unela ........................31 

ungulatus, Manucomplanus ........144 

Ungulinidae ...................10 

unicolor, Brissus ................194 

unicornis, Bemlos ...............127 

unicornis, Cyclaspis..............138 

unicornis, Nematonereis............87 

unidens, Pontonia ...............140 

unifasciatum, Epitonium ...........22 

unifasciatus, Lembos .............127 

unifrons, Eunice .................87 

Unipeltata....................124 

uniplicata, Simnialena.............20 

unoculus, Sclerocheilus ............92 

Upogebia .....................143 

Upogebiidae ..................142 

Urobopyrus ...................134 

urocaridis, Schizobopyrina .........133 

urocarinatus, Atylus..............128 

Urodemella ...................195 

Uromunna ....................133 

Urosalpinx.....................24 

usnea, Neritina ..................17 

uva, Doto......................34 

FMRI Technical Report TR-3 237 

V 

vadorum, Ampelisca .............126 

valida, Ampithoe................126 

valida, Turbonilla ................30 

vallei, Amphithalamus.............18 

Valvatida.....................192 

Valvifera .....................136 

Vandeloscia ...................136 

vanderhorsti, Malacoceros ..........89 

vandoverae, Michelea ............143 

vanga, Anamixis ................127 

vanhyningi, Cerithiopsis ...........22 

vanhyningi, Orinella ..............29 

Vanikoridae ...................20 

Vanikoro ......................20 

varia, Lepidasthenia ..............85 

variabilis, Arene .................16 

variabilis, Donax ................13 

variabilis, Lepidonotus.............85 

variabilis, Mastobranchus ..........92 

varians, Cyclaspis ...............138 

varians, Petaloconchus.............20 

Varicorbula ....................14 

variegata, Charonia tritonis .........21 

variegata, Diodora ...............16 

variegata, Malmgreniella ...........85 

variegata, Syllis .................83 

variegatus, Lytechinus ............194 

variolosa, Symethis ..............145 

varium, Bittiolum ................17 

Varuninae ....................150 

vasquezi, Upogebia ..............143 

Vasum ........................24 

Vaunthompsonia ................138 

Vaunthompsoniinae ............138


Veleroniopsis ..................140 

velum, Solemya ..................8 

Veneridae .....................13 

Veneroida.....................10 

venezuelensis, Diplodonta ..........11 

venosus, Dardanus ..............143 

ventralis, Portunus...............147 

ventricosa, Meoma...............194 

ventricosus, Tripneustes ...........194 

venustula, Arene .................16 

verae, Naticarius.................21 

Vermetidae ....................20 

Vermicularia....................17 

vermiformis, Lomanotus............34 

Vermiliopsis ....................95 

verrilli, Aglaophamus .............84 

verrilli, Ampelisca ...............126 

verrilli, Lumbrineris ..............87 

verrilli, Streblosoma ..............94 

verrilli, Streptosyllis ..............83 

verrilli, Terebella .................94 

verrillii, Cylichna ................30 

verrucicornis, Spurilla .............34 

verrucosa, Doris .................33 

verrucosa, Pilargis................82 

verrucosus, Mithrax..............147 

verruculosa, Hermenia .............84 

versicolor, Nerita.................17 

versicolor, Tellina ................12 

versipedata, Ceratonereis ...........84 

verticilata, Hermaea ..............32 

Verticordiidae ..................15 

verugera, Exogone ................82 

vespaceum, Cymatium .............22 

vespertilium, Gastropteron ..........30 

vespuciana, Nannodiella ...........28 

vestale, Microstelma ..............18 

vestitum, Caecum ................19 

Vexillum ......................26 

vibex, Nassarius .................25 

vicina, Cumella.................138 

vicina, Processa.................142 

vietezi, Grubeosyllis ..............82 

vincinti, Branchiomaldane ..........93 

vinosa, Terebra ..................27 

violaceomaculata, Eunice ...........87 

violaceum, Pneumoderma...........37 

virga, Turbonilla .................30 

virgata, Turbonilla................30 

virginea, Neritina ................17 

virginianum, Prunum .............26 

virginica, Crassostrea..............10 

virginica, Odostomia ..............29 

virgula, Creseis ..................36 

viridari, Alpheus ................141 

viridaria, Turbonilla...............30 

viridis, Cancellus................143 

viridis, Chiton ...................7 

viridis, Chloeia ..................86 

viridis, Echinometra..............194 

viridis, Loimia ..................94 

viridis, Marphysa ................87 

viridis, Polybranchia ..............32 

viridis, Smaragdia................17 

viridis, Solen ...................12 

Viridrillia .....................27 

viridula, Diodora ................16 

Vitreolina......................23 

Vitricythara ....................28 

Vitrinella ......................19 

Vitrinellidae ...................19 

vittata, Dorvillea.................88 

vittata, Ophionereis ..............193 

vittata, Ophiopsila...............193 

vittata, Sayoscia ................136 

vittata, Syllis ...................83 

vittatopsis, Eunice ................87 

vittatus, Clibanarius .............143 

vittigera, Trypanosyllis.............83 

vivipara ?, Syllis .................83 

vocans, Laleonectes ..............147 

vocator, Uca ...................150 

Volutidae .....................25 

Volvarina......................26 

Volvatella .....................31 

Volvatellidae...................31 

Volvulella .....................31 

vossi, Portunus .................147 

vossi, Processa..................142 

vossi, Stiliger ...................32 

vulgaris, Octopus ................35 

vulgaris, Palaemonetes ............140 

vulgaris, Sabellaria ...............93 

W 

wah, Onchidella .................35 

walkeri, Sphaeroma ..............136 

wandoense,Vexillum ..............26 

warta, Dendrodoris ...............33 

wassi, Aricidea ..................90 

wassi, Paguristes ................144 

wassi, Sigambra .................82 

watlingi, Spilocuma ..............138 

wattla, Armina ..................34 

webbi, Hypselodoris ..............33 

weberi, Bailya...................24 

weberi, Eulimastoma ..............29 

weberi, Mexieulepis...............85 

weberi, Parviturbo ................17 

weberi, Teskeyostrea...............10 

websteri, Alpheus ...............141 

websteri, Crucigera ...............95 

websteri, Eunice .................87 

websteri, Lembos ................127 

websteri, Lepidopa...............145 

websteri, Polydora ................89 

Websterinereis ..................84 

wegeneri, Tylos .................136 

weintraubi, Eunice ...............87 

weismanni, Pionosyllis ............83 

wellfleetensis, Brania..............82 

wellsi, Tritonidoxa ................33 

wendtii, Ophiocoma..............193 

wennerae, Neogonodactylus ........124 

Westheidia.....................88 

westoni, Grubeulepis ..............85 

whelpleyi, Parhyalella ............129 

whitingi, Nannosquilla............125 

wigleyi, Hemiproto ..............132 

wigleyi, Protohaustorius ...........129 

wigleyi, Spiophanes...............89 

williami,Viridrillia ...............27 

Williamia .....................35 

williamsi, Lytechinus .............194 

williamsi, Maera ................129 

wilsoni, Munnogonium ...........133 

wilsoni, Periclimenaeus ...........140 

wolffi, Bopyrissa ................134 

woodringi, Splendrillia ............27 

worki, Anisodoris ................33 

worsfoldi, Nanobalcis .............23 

wrightsvillensis, Turbonilla..........30 

wui, Eunice ....................87 

wurdemanii, Ophiophragmus .......193 

wurdemanni, Isocheles ............143 

wurdemanni, Lysmata ............141 

238 FMRI Technical Report TR-3 

X 

Xanthidae ....................148 

Xanthoidea ...................147 

Xanthodius....................149 

Xenanthura ...................132 

Xenophora .....................20 

Xiphopenaeus ..................139 

Y 

yoyo, Divariscintilla ..............11 

ypsilon, Spiraserpula..............95 

yucatanicus, Periclimenes ..........140 

yucatecana, Mathilda .............28 

yucatecanum, Calliostoma ..........17 

Z 

Zafrona .......................25 

Zaops .......................150 

Zebina........................18 

zebra, Acanthochitona ..............7 

zebra, Aplysiopsis ................32 

zebra, Arca......................9 

zebra, Macrocypraea ..............20 

zebroides, Pilsbryspira .............27 

zetlandica, Caulleriella ............90 

Zeuxo .......................137 

ziczac, Conus ...................28 

ziczac, Euvola ...................10 

ziczac, Nodilittorina ..............18 

zonalis, Macroclymene ............92 

Zonulispira ....................27 

zoobotryon, Okenia ...............32 

zora, Iphimedia .................129 

zostericola, Hippolyte.............141 

Zygopa ......................145 

zythochroa,Vexillum ..............26

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