A Checklist of the Seaweeds of the Mediterranean - Walter de Gruyter

Introduction 

Botanica Marina Vol. 45, 2002, pp. 217–230 © 2002 by Walter de Gruyter · Berlin · New York 

A Checklist of the Seaweeds of the Mediterranean and 

Atlantic Coasts of Morocco. II. Phaeophyceae 

S. Benhissoune a , C.-F. Boudouresque b and M. Verlaque b * 

a Département de Biologie, Faculté des Sciences, BP 403,Agadir Principale, 80000, Morocco 

b Centre d’Océanologie de Marseille, Luminy Campus, 13288 Marseille Cedex 9, France 

* Corresponding author: verlaque@com.univ-mrs.fr 

A checklist of Moroccan marine brown algae, based on both literature records and some original data, is given.The 

distribution of each taxon for this region (12 Mediterranean and 27 Atlantic localities) is reported. Using 

present-day taxonomy, 131 species, infraspecific taxa and life-history phases are accepted for the Moroccan 

coasts. 

Several attempts have been made to produce a 

checklist of Moroccan marine seaweeds. Some of 

these are blatantly out of date (e.g. Bornet 1892, Debray 

1897, Gattefossé and Werner 1935, Gayral 

1961). Others only concern the Mediterranean coast 

(Navarro and Gallardo 1989, Ribera et al. 1992, 

González-García and Conde-Poyales 1994), while 

the recent checklist of Riadi and Kazzaz (1998) appears 

to be incomplete and uncritical. 

If biogeographical comparisons are to be made between 

Moroccan provinces, as well as between Morocco 

and neighbouring countries, on the basis of recent 

checklists (e.g. South and Tittley 1986, Lawson 

and John 1987, Fletcher 1987 and Ribera et al. 1992), 

an updated and critical checklist must be drawn up. 

The present paper is the second part of a checklist 

intended to be a catalogue of benthic multicellular 

algae of the Moroccan coasts, which includes the 

Chlorophyceae s. l. (Benhissoune et al. 2001), Phaeophyceae 

(= Fucophyceae; present study) and Rhodophyta. 

Material and Methods 

In this paper, the Moroccan coasts correspond to the 

geographical concept of Morocco, in other words, the 

Spanish territories of Ceuta, Melilla and the Chafarinas 

Islands are included. We have considered 39 localities 

(Fig. 1, Table I), 12 of which are in the 

Mediterranean (M; from East to West: Saaidia, Chafarinas 

Islands, Nador, Melilla, Al Hoceima, Jabha, 

Tamernoute, M’diq, Cabo Negro, Tetouan, Ceuta, 

Ksar-Sghir) and 27 are found along the Atlantic coast 

(A; from North to South:Tanger, Cap Spartel,Azilah, 

Larache, Moulay Bou Selham, Kenitra, Salé, Rabat, 

Temara, Bouznika, Mohammedia, Casablanca, El Jadida, 

Oualidia, Safi, Djorf el Yhoudi, Sidi Boulbra, 

Essaouira, Imessouane, Cap Ghir, Agadir, Tifnit, 

Mirhleft, Sidi Ifni, Tan Tan, Moroccan Sahara and 

Cap Blanc). The toponymy is that of the MARCUS 

map (1/1400000) edited in 1980 by the National 

Topographical Center of Morocco. 

The accepted taxon name is followed (between 

square brackets) by the basionym and by the other 

synonyms used by the authors of Moroccan citations. 

The taxa (orders, families, genera and species) are 

arranged alphabetically. When infraspecific taxa are 

listed, the autonym, if present, is cited first. Delin- 

Fig. 1. Map of the Moroccan localities taken into consideration.

218 S. Benhissoune et al. 

Table I. List of localities taken into consideration (see Fig. 1) and alternative names for these localities. 

Localities Alternative names Neighbouring 

Agadir Amesnaz, El Khemiss 

Al Hoceima 

Azilah Assila,Azila 

Bouznika Oued Cherrat 

Cabo Negro Cabo-negro 

Cap Blanc 

Cap Ghir 

Cap Spartel 

Casablanca Aïn Diab,Aïn seba El Hank, Sidi Abderrahmane, Oukacha, Dar Bouazza, Dar Ben Abid 

Ceuta Benyounes = Benzu 

Chafarinas Islands 

Djorf el Yhoudi Jorf el Yhoudi 

El Jadida Mazagan Moulay Abdelah, Sidi Bouzid,Azemour 

Essaouira Saouira Mogador 

Imessouane 

Jabha 

Kenitra Mehdia, Sebou 

Ksar-Sghir Ksar es ghir 

Larache 

M’diq 

Melilla Cabo tres forces, Cala blanca, Segunda caseta,Atalayoun 

Mirhleft Mirleft Aglou, sidi moussa d’Aglou,Tiznit 

Mohammedia Fedala, Zenatta, Zenattas Oued Mellah, Oued Nefifikh, Lalla mimouna 

Moroccan Sahara Rio de Oro, Oued eddahab 

Moulay Bou Selham 

Nador Mar Chica, Sebcha Bouareg Lagune de Nador 

Oualidia Sidi Moussa, lagune de sidi moussa 

Rabat 

Saaidia Essaaidia 

Safi Asfi Cap Cantin 

Salé Kasba Guenaoua Camp sartige, Bou regreg, Bou Regreg, bou reg-reg 

Sidi Boulbra 

Sidi Ifni 

Tamernoute 

Tan Tan Tan-Tan,TanTan,Tantan 

Tanger Tangiers 

Temara Oued yquem, Rose marie, Gay Ville, Sables d’Or, El Harhoura, Skhirat 

Tetouan Cabo Mazari, Martil 

Tifnit 

Citations from neighbouring localities were attributed to the names of the locality in the far left-hand column. 

eation of the orders and families follows that used by 

Silva et al. (1996). For each taxon and each locality, 

the numbers between brackets refer to the references. 

A question mark after the reference number 

signifies that the taxon was mentioned with doubt.‘*’ 

means that the taxon was collected by one of the authors 

of the present study (S. B.). The liquid preserved 

specimens and exsiccates were deposited in 

the Herbarium Benhissoune, Département de Biologie, 

Faculté des Sciences,Agadir. 

Results 

The present checklist encompasses 131 items (species, 

infraspecific taxa and life-history phases). Some taxa 

listed and cited here require further investigation (see 

the Notes section). In addition, three taxa are considered 

as taxa excludenda and two as taxa inquirenda. 

Checklist of Phaeophyceae 

Chordariales 

Chordariaceae 

Cladosiphon zosterae (J. Agardh) Kylin [= Myriocladia 

zosterae J. Agardh, Castagnea zosterae (J. 

Agardh) Thuret]. A: Tanger (9?). 

Levringia atlantica (J. Feldmann) Kylin1 [= Myriogloea 

atlantica J. Feldmann]. A: Rabat (42), Moroccan 

Sahara (23, 29, 30, 31, 42, 61).

Levringia brasiliensis (Montagne) A. B. Joly 1 [= 

Mesogloea brasiliensis Montagne]. A: Rabat (42), 

Temara (3, 12, 13, 40), without locality (41). 

Liebmannia leveillei J.Agardh. A: Tanger (9, 25, 38). 

Mesogloia vermiculata (J. E. Smith) S. F. Gray [= 

Rivularia vermiculata J. E. Smith]. M: Cabo Negro 

(58), Ksar-Sghir (58). 

Sauvageaugloia divaricata (Clemente) Cremades 2 [= 

Ulva divaricata Clemente, Myriocladia chordariaeformis 

P. L. Crouan et H. M. Crouan, Sauvageaugloia 

chordariaeformis (P. L. Crouan et H. M. 

Crouan) Kylin].Without locality (8). 

Sauvageaugloia griffithsiana (A.W. Griffiths ex Harvey) 

G. Hamel ex Kylin 2 [= Mesogloia griffithsiana 

A.W. Griffiths ex Harvey]. A: Tanger (9, 25, 38, 

100). 

Corynophlaeaceae 

Corynophlaea umbellata (C. Agardh) Kützing [= 

Corynophora umbellata C.Agardh]. A: Tanger (38, 

100). 

Leathesia difformis (Linnaeus) J. E. Areschoug [= 

Tremella difformis Linnaeus, Leathesia difformis 

(Linnaeus) J. E. Areschoug var. tingitana Schousboe 

ex Bornet]. Without locality (8). M: 

Tamernoute (58), Cabo Negro (58), Ksar-Sghir 

(58). A: Tanger (9, 25, 38, 100), Temara (12, 21), 

Bouznika (100), without locality (20). 

Myriactula arabica (Kützing) J. Feldmann [= Phycophila 

arabica Kützing, Myriactis arabica (Kützing) 

Kuckuck] A: Tanger (38). 

Elachistaceae 

Elachista flaccida (Dillwyn) Fries [= Conferva flaccida 

Dillwyn]. Without locality (8). A: Tanger (9, 25, 

38, 100), Temara (3, 12), Mohammedia (3), 

Casablanca (21, 38, 100), El Jadida (21, 100), without 

locality (40). 

Elachista fucicola (Velley) Areschoug [= Conferva 

fucicola Velley]. A: El Jadida (*), Oualidia (*), Sidi 

Boulbra (6, 7), Essaouira (*),Agadir (*). 

Elachista neglecta Kuckuck. A: Temara (3), Mohammedia 

(3). 

Myrionemataceae 

Myrionema strangulans Carmichael ex Greville [= 

Myrionema vulgare Thuret]. M: Tamernoute (58), 

Cabo Negro (58), Ksar-Sghir (58). A: Tanger (9, 25, 

38). 

Cutleriales 

Cutleriaceae 

‘Aglaozonia chilosa Falkenberg’ life-history phase3 . 

M: Saaidia (*), M’diq (*), Cabo Negro (58), Ksar- 

Sghir (*). A: Tanger (*), Temara (3, *), Mohammedia 

(3, *), El Jadida (*). 

A checklist of the seaweeds of Morocco. II. Phaeophyceae 219 

‘Aglaozonia melanoidea (Schousboe ex Bornet) 

Sauvageau’ life-history phase 4 [= Zonaria 

melanoidea Schousboe ex Bornet]. M: Saaidia (*), 

M’diq (*),Tetouan (*). A: Tanger (*),Temara (3, *), 

Mohammedia (3, *). 

‘Aglaozonia parvula (Greville) Zanardini’ life-history 

phase 5 [= Zonaria parvula Greville]. M: Saaidia 

(50, *), Nador (50), Melilla (48, 50), Al Hoceima 

(50), Jabha (*),Tetouan (*). A: Tanger (*), Larache 

(*),Temara (3, *), Mohammedia (3, *). 

Cutleria adspersa (Mertens ex Roth) De Notaris 4 [= 

Ulva adspersa Mertens ex Roth]. Without locality 

(8). M: Saaidia (50, *), Melilla (50), Al Hoceima 

(50), Tamernoute (58), Cabo Negro (58), Ksar- 

Sghir (58, *). A: Tanger (9, 25, 38), Rabat (21), 

Temara (3, 12, *), Mohammedia (3, 71), El Jadida 

(5), Essaouira (21, *),Agadir (21, *), without locality 

(20, 40). 

Cutleria multifida (J. E. Smith) Greville 5 [= Ulva multifida 

J. E. Smith]. M: Melilla (50), Al Hoceima 

(50), Jabha (*). A: Tanger (9, 25, 38). 

Zanardinia typus (Nardo) G. Furnari [= Stifftia typus 

Nardo, Padina collaris Montagne, Zanardinia prototypus 

(Nardo) Nardo nom. illeg., Zanardinia collaris 

(Montagne) P. L. Crouan et H. M. Crouan]. M: 

Saaidia (*), Chafarinas Islands (35), Nador (69). A: 

Tanger (9, 25, 38), Casablanca (38). 

Desmarestiales 

Arthrocladiaceae 

Arthrocladia villosa (Hudson) Duby [= Conferva villosa 

Hudson]. A: Tanger (9). 

Desmarestiaceae 

Desmarestia ligulata (Lightfoot) J. V. Lamouroux [= 

Fucus ligulatus Lightfoot, Desmarestia herbacea 

(Turner) J. V. Lamouroux]. A: Tanger (9, 25, 38), 

Mohammedia (38), El Jadida (*), Essaouira (21, *), 

without locality (40). 

Dictyosiphonales 

Asperococcaceae 

Asperococcus bullosus J. V. Lamouroux [= A. turneri 

(Dillwyn ex J. E. Smith) W. J. Hooker]. M: Saaidia 

(*), Nador (47, 50, 51), M’diq (*), Cabo Negro (58), 

Ksar-Sghir (58, *). A: Tanger (9, 25, 38, 100, *), Azilah 

(*), Larache (*), Temara (*), El Jadida (*), 

Agadir (*),Tan Tan (*). 

Asperococcus compressus A. W. Griffiths ex W. J. 

Hooker [= Haloglossum compressum (A. W. Griffiths 

ex W. J. Hooker) G. Hamel]. A: Tanger (9, 38), 

Temara (42). 

Hecatonema terminale (Kützing) Kylin 6 [= Ectocarpus 

terminalis Kützing]. A: Tanger (9, 25, 38), Rabat 

(42),Temara (3), Mohammedia (3, 71).


Myriotrichiaceae 

Litosiphon laminariae (Lyngbye) Harvey [= Bangia 

laminariae Lyngbye]. A: Casablanca (21), Essaouira 

(21). 

Dictyotales 

Dictyotaceae 

Dictyopteris polypodioides (De Candolle) J. V. Lamouroux 

[= Ulva polypodioides De Candolle, Dictyopteris 

membranacea (Stackhouse) Batters]. Without 

locality (8). M: Chafarinas Islands (14, 67), 

Melilla (10, 48), Tamernoute (58), M’diq (*), Cabo 

Negro (58), Ksar-Sghir (58, *), without locality 

(50). A: Tanger (9, 16, 25, 38, 100), Rabat (12, 21, 25, 

38, 100),Temara (12, 21, 100), Mohammedia (3, 71), 

Casablanca (21, 38, 81, 100), El Jadida (5, 18, 21, 25, 

38, 81, 98, 100, *), Oualidia (*), Djorf el Yhoudi (98, 

100), Sidi Boulbra (6, 7), Essaouira (21, 25, 38, 81, 

98, 100, *), Agadir (*), Sidi Ifni (*), cosmopolitan 

(21), without locality (40). 

Dictyota bartayresiana J.V. Lamouroux [= D. bartayresii 

J.V. Lamouroux]. A: Moroccan Sahara (61). 

Dictyota ciliolata Sonder ex Kützing. A: Moroccan 

Sahara (61). 

Dictyota dichotoma (Hudson) J. V. Lamouroux var. 

dichotoma [= Ulva dichotoma Hudson]. Without 

locality (8). M: Saaidia (*), Chafarinas Islands (14, 

67), Nador (47, 51), Melilla (10, 48), Tamernoute 

(58), M’diq (*), Cabo Negro (58), Tetouan (67, *), 

Ksar-Sghir (58), without locality (50). A: Tanger (9, 

25, 38, 100, *),Azilah (*), Rabat (21, 38, 81),Temara 

(3, 12, 100, *), Mohammedia (3, 71, 72, *), 

Casablanca (21, 38, 81, 98, 100), El Jadida (5, 18, 98, 

100, *), Oualidia (*), Safi (100), Djorf el Yhoudi 

(100), Sidi Boulbra (6, 7), Essaouira (21, 38, 39, 81, 

98, *), Cap Ghir (98, 99, 100), Agadir (99, *), Tan 

Tan (*), cosmopolitan (21), without locality (40). 

Dictyota dichotoma (Hudson) J. V. Lamouroux var. 

intricata7 (C. Agardh) Greville [= Zonaria dichotoma 

(Hudson) C. Agardh var. intricata C. 

Agardh, Dictyota dichotoma (Hudson) J. V. Lamouroux 

var. implexa (Desfontaines) S. F. Gray]. M: 

Chafarinas Islands (14, 67), Melilla (48), M’diq (*), 

Tetouan (*), without locality (50). A: Tanger (9, 25, 

38), Larache (*), Temara (3, *), Mohammedia (3, 

*), El Jadida (*),Agadir (*),Tifnit (*). 

Dictyota fasciola (Roth) J.V. Lamouroux var. fasciola 

[= Fucus fasciola Roth, Dilophus fasciola (Roth) 

M. Howe]. Without locality (8). M: Chafarinas Islands 

(35), Nador (50), Melilla (50), Al Hoceima 

(50), Jabha (50), Tamernoute (58), Cabo Negro 

(58), Ksar-Sghir (58). A: Rabat (100), Temara (3, 

12, 21), Bouznika (100), Mohammedia (3, 71), El 

Jadida (5), without locality (20, 40). 

Dictyota fasciola (Roth) J. V. Lamouroux var. repens 

(J. Agardh) Ardissone [= Dictyota repens J. 

Agardh]. A: Tanger (9, 25, 38). 

Dictyota linearis (C. Agardh) Greville 7 [= Zonaria 

linearis C. Agardh]. M: Saaidia (50), Nador (50), 

Melilla (10, 48, 50). A: Tanger (9, 25, 38, 100), 

Temara (21, 100), Bouznika (100). 

Dictyota spiralis Montagne [= Dictyota ligulata Kützing, 

Dilophus spiralis (Montagne) G. Hamel, Dilophus 

ligulatus (Kützing) J. Feldmann]. Without locality 

(8). M: Chafarinas Islands (14, 67), Nador 

(51), Melilla (10, 48), Tamernoute (58), Cabo Negro 

(58), Ksar-Sghir (58), without locality (50). A: 

Tanger (9, 25, 38, 100), Rabat (21, 38), Temara (3, 

100), Mohammedia (3, 71), Casablanca (21, 38, 81, 

100), El Jadida (5, 21, 100), Essaouira (21, 38, 81), 

Agadir (21, 100), without locality (40). 

Lobophora variegata (J. V. Lamouroux) Womersley 

ex E. C. Oliveira [= Dictyota variegata J. V. Lamouroux, 

Aglaozonia canariensis Sauvageau]. Without 

locality (8). A: Moroccan Sahara (61). 

Padina pavonica (Linnaeus) Thivy [= Fucus pavonicus 

Linnaeus, Padina pavonia J. V. Lamouroux]. 

Without locality (8). M: Chafarinas Islands (14, 67), 

Nador (47, 51), Melilla (10, 48), Tamernoute (58), 

M’diq (*), Cabo Negro (58), Tetouan (*), Ksar- 

Sghir (58, *), without locality (50). A: Tanger (9, 25, 

38),Azilah (*), Larache (*), Rabat (21, 38),Temara 

(3, 12, *), Mohammedia (3, 71, *), Casablanca (21, 

38, 81), El Jadida (5, *), Oualidia (*), Essaouira (21, 

38, *), Cap Ghir (98, *), Sidi Ifni (*), Tan Tan (*), 

cosmopolitan (21), without locality (40, 56). 

Spatoglossum schroederi (C. Agardh) Kützing [= 

Zonaria schroederi C. Agardh, Spatoglossum 

areschougii J. Agardh]. A: Rabat (19, 21, 63, 100), 

Temara (19, 21, 100), Moroccan Sahara (61). 

Spatoglossum solieri (Chauvin ex Montagne) Kützing 

[= Dictyota solieri Chauvin ex Montagne]. M: 

Saaidia (*), Al Hoceima (50), Jabha (*), M’diq (*). 

A: Tanger (9, 19, 25, 38, 100, *),Temara (3, 12, 21, *), 

Bouznika (100), Mohammedia (3, 71, *), Casablanca 

(19, 21, 38, 81, 100), El Jadida (*), without locality 

(40, 43). 

Taonia atomaria (Woodward) J. Agardh 8 [= Ulva 

atomaria Woodward]. Without locality (8). M: 

Saaidia (*), Nador (47, 50, 51), Melilla (50), Al Hoceima 

(50), Jabha (*), M’diq (*), Tetouan (*), Ksar- 

Sghir (*). A: Tanger (9, 25, 38, 100, *), Azilah (*), 

Larache (*), Temara (3, 12, 21, 100, *), Bouznika 

(100), Mohammedia (3, 71, *), Casablanca (21, 38, 

81, 100), Oualidia (*), Safi (100), Essaouira (*), Sidi 

Ifni (*),Tan Tan (*), without locality (40). 

Taonia pseudociliata (J. V. Lamouroux) Nizamuddin 

et Godeh 8 [= Fucus pseudociliatus J.V. Lamouroux, 

Dictyota ciliata J. V. Lamouroux, Taonia atomaria 

(Woodward) J. Agardh f. ciliata (J. V. Lamouroux) 

Nizamuddin]. M: Chafarinas Islands (67). 

Zonaria tournefortii (J. V. Lamouroux) Montagne [= 

Fucus tournefortii J. V. Lamouroux]. Without locality 

(8). M: Nador (47, 50, 51), Melilla (10, 48, 50),Al 

Hoceima (50), Jabha (*), M’diq (*), Ksar-Sghir (*). 

A: Tanger (9, 25, 38, 55, 100, *), Azilah (*), Larache

(*), Rabat (21, 38, 81, 100), Temara (3, 12, 21, 99, 

100, *), Mohammedia (3, 71, 81, 99, 100, *), 

Casablanca (21, 38, 81, 100), El Jadida (21, 25, 38, 

81, *), Cap Ghir (99, 100), Agadir (99, *), Tifnit (*), 

Sidi Ifni (*), without locality (40). 

Ectocarpales 

Ectocarpaceae 

Acinetospora crinita (Carmichael) Kornmann [= Ectocarpus 

crinitus Carmichael, Acinetospora vidovichii 

(Meneghini) Sauvageau, A. pusilla (Griffiths 

ex Harvey) De Toni, Ectocarpus pusillus 

Griffiths ex Harvey]. M: Saaidia (50, *), Nador 

(50), Melilla (50), Jabha (*), Tamernoute (58), 

M’diq (*), Cabo Negro (58). A: Imessouane (42). 

Ectocarpus fasciculatus Harvey var. fasciculatus. M: 

Melilla (48), Tamernoute (58), Cabo Negro (58), 

Ksar-Sghir (58, *). A: Larache (*), Temara (3, 12, 

*), Mohammedia (3, 71, 72), El Jadida (5, 99, 100, 

*), Essaouira (21, 100), Agadir (21, 100), without 


Ectocarpus fasciculatus Harvey var. pycnocarpus 

(Rosenvinge) Cardinal [= E. pycnocarpus 

Rosenvinge]. M: Saaidia (*), Melilla (50), Al Hoceima 

(50), Jabha (50, *), M’diq (*),Tetouan (*). 

Ectocarpus siliculosus (Dillwyn) Lyngbye var. siliculosus 

[= Conferva siliculosa Dillwyn, Ectocarpus 

confervoides (Roth) Le Jolis]. Without locality (8). 

M: Saaidia (50, *), Chafarinas Islands (14, 67), 

Nador (47, 50, 51), Melilla (48, 50), Al Hoceima 

(50), Jabha (*), Tamernoute (58), M’diq (*), Cabo 

Negro (58),Tetouan (*), Ksar-Sghir (58). A: Azilah 

(*), Larache (*), Rabat (21, 38), Temara (3, 12, 21, 

38, 100), Mohammedia (3, 71), Casablanca (100), 

El Jadida (5, 21), Safi (21), without locality (40). 

Ectocarpus siliculosus (Dillwyn) Lyngbye var. 

crouanii (Thuret) Gallardo [= E. crouanii Thuret]. 

M: Tamernoute (58), Cabo Negro (58), Ksar-Sghir 

(58). 

Ectocarpus siliculosus (Dillwyn) Lyngbye var. penicillatus 

C. Agardh [= E. penicillatus (C. Agardh) 

Kjellman]. M: Saaidia (50, *), Nador (50), Melilla 

(50),Al Hoceima (50), Jabha (*), Ksar-Sghir (*). 

Ectocarpus virescens Thuret ex Sauvageau. M: 

Tamernoute (58), Cabo Negro (58), Ksar-Sghir 

(58). A : Rabat (38). 

Feldmannia battersiides (Ercegovic) Cormaci et G. 

Furnari [= Ectocarpus battersiides Ercegovic]. M: 

Nador (50). 

Feldmannia caespitula (J. Agardh) Knoepffler-Péguy 

var. lebelii (Areschoug ex P. L. Crouan et H. M. 

Crouan) Knoepffler-Péguy [= Ectocarpus lebelii 

Areschoug ex P. L. Crouan et H. M. Crouan, Feldmannia 

lebelii (Areschoug ex P. L. Crouan et H. M. 

Crouan) G. Hamel]. M: Tamernoute (58), Cabo 

Negro (58), Ksar-Sghir (58). A: Tanger (9, 25, 38), 

Agadir (21). 


Feldmannia irregularis (Kützing) G. Hamel [= Ectocarpus 

irregularis Kützing]. M: Nador (47, 51), 

Cabo Negro (58). A: Rabat (42), Mohammedia (3, 

71), Casablanca (38), without locality (9). 

Feldmannia paradoxa (Montagne) G. Hamel var. 

paradoxa 9 [= Ectocarpus paradoxus Montagne, Ectocarpus 

globifer Kützing, Feldmannia globifera 

(Kützing) G. Hamel]. M: Saaidia (*), Nador (47, 50, 

51), Melilla (48, 50), Al Hoceima (50), Tamernoute 

(58), M’diq (*), Cabo Negro (58), Ksar-Sghir (58), 

without locality (50). A: Tanger (9, 25, 38, 100), 

Agadir (21, 100), without locality (20). 

Feldmannia simplex (P. L. Crouan et H. M. Crouan) 

G. Hamel [= Ectocarpus simplex P. L. Crouan et H. 

M. Crouan]. M: Tamernoute (58), Cabo Negro 

(58), Ksar-Sghir (58). A: Temara (3), Mohammedia 

(3, 71). 

Herponema valiantei (Bornet ex Sauvageau) G. 

Hamel [= Ectocarpus valiantei Bornet ex 

Sauvageau]. A: El Jadida (21). 

Hincksia granulosa (J. E. Smith) P. C. Silva [= Conferva 

granulosa J. E. Smith, Ectocarpus granulosus (J. 

E. Smith) C. Agardh, Giffordia granulosa (J. E. 

Smith) G. Hamel]. M: Tamernoute (58), Cabo Negro 

(58), Ksar-Sghir (58). A: Tanger (9, 25, 38, 100), 

Rabat (21, 100), Temara (3), Mohammedia (3, 71), 


Hincksia hincksiae (Harvey) P. C. Silva [= Ectocarpus 

hincksiae Harvey, Giffordia hincksiae (Harvey) G. 

Hamel]. A: Tanger (9, 25), Larache (*), Temara (3, 

12, *), Mohammedia (3, 71, *), Casablanca (21, 38), 

El Jadida (*), Sidi Boulbra (6, 7), Essaouira (21), 

Agadir (21, *), Sidi Ifni (*), without locality (40). 

Hincksia mitchelliae (Harvey) P. C. Silva [= Ectocarpus 

mitchelliae Harvey, Giffordia mitchelliae (Harvey) 

G. Hamel]. M: Saaidia (*), Nador (47, 51), 

Melilla (48), M’diq (*), without locality (50). A: 

Salé (21, 100), Rabat (21), Temara (21, 100), 

Casablanca (100),Agadir (21, 100). 

Hincksia ovata (Kjellman) P. C. Silva [= Ectocarpus 

ovatus Kjellman, Giffordia intermedia (Rosenvinge) 

S. Lund]. M: Melilla (50), Al Hoceima (50), 

Jabha (*),Tetouan (*). 

Hincksia rallsiae (Vickers) P. C. Silva [= Ectocarpus 

rallsiae Vickers, Feldmannia rallsiae (Vickers) G. 

Hamel, Giffordia rallsiae (Vickers) W. R. Taylor]. 

A: Rabat (21), Casablanca (21), Moroccan Sahara 


Hincksia sandriana (Zanardini) P. C. Silva [= Ectocarpus 

sandrianus Zanardini,E. elegansThuret,Giffordia 

sandriana (Zanardini) G. Hamel]. M: 

Tamernoute (58),Cabo Negro (58),Ksar-Sghir (58). 

Hincksia secunda (Kützing) P. C. Silva [= Ectocarpus 

secundus Kützing, Giffordia secunda (Kützing) 

Batters]. A: Tanger (9, 25, 38, 100), Temara (100), 

Casablanca (21, 100). 

Kuetzingiella battersii (Bornet ex Sauvageau) Kornmann 

[= Ectocarpus battersii Bornet ex 

Sauvageau]. M: Tamernoute (58). A: Rabat (42).


Streblonema deformans (P. Dangeard) G. Hamel [= 

Ectocarpus deformans P. Dangeard]. A: Essaouira 


Streblonema parasiticum (Sauvageau) Levring [= Ectocarpus 

parasiticus Sauvageau, Entonema parasiticum 

(Sauvageau) G. Hamel]. A: Casablanca 

(21). 

Pylaiellaceae 

Bachelotia antillarum (Grunow) Gerloff10 [= Ectocarpus 

antillarum Grunow, Bachelotia fulvescens 

(Bornet) Kuckuck ex G. Hamel, Pylaiella fulvescens 

Bornet]. M: Saaidia (*), Jabha (*), 

Tamernoute (58), M’diq (*), Cabo Negro (58), 

Ksar-Sghir (58, *). A: Tanger (9, 25, 38, 40, *), Azilah 

(*), Rabat (40),Temara (3, 12, *), Mohammedia 

(3, 21, 71, 72, *), El Jadida (5, *), Essaouira (*), Cap 

Ghir (21, 40),Agadir (21, 40), without locality (20). 

Pylaiella littoralis (Linnaeus) Kjellman11 [= Conferva 

littoralis Linnaeus]. A: Casablanca (81). 

Ralfsiaceae 

Hapalospongidion macrocarpum (J. Feldmann) 

León Alvarez et González [= Ralfsia macrocarpa J. 

Feldmann, Mesospora macrocarpa (J. Feldmann) 

den Hartog, Mesospora mediterranea J. Feldmann]. 

M: Melilla (10, 48), Cabo Negro (58), without locality 

(50). 

Nemoderma tingitanum Schousboe ex Bornet12 . M: 

Saaidia (50, *), Nador (50), Melilla (50), Jabha (*), 

M’diq (*). A: Tanger (9, 25, 38, 100), Azilah (*), 

Temara (3, 21, *), Bouznika (100), Mohammedia 

(3), El Jadida (*). 

Pseudolithoderma adriaticum (Hauck) Verlaque [= 

Lithoderma adriaticum Hauck]. A: Rabat (11), 

Temara (11). 

Pseudolithoderma extensum (P. L. Crouan et H. M. 

Crouan) S. Lund [= Ralfsia extensa P. L. Crouan et 

H. M. Crouan, Lithoderma fatiscens Kuckuck nom. 

illeg.]. A: Rabat (11),Temara (11). 

Ralfsia verrucosa (Areschoug) Areschoug [= Cruoria 

verrucosa Areschoug]. Without locality (8). M: 

Saaidia (50, *), Melilla (48, 50), Al Hoceima (50), 

Jabha (*),Tamernoute (58), M’diq (*), Cabo Negro 

(58), Ksar-Sghir (58). A: Tanger (9, 25, 38, 100, *), 

Azilah (*), Temara (3, *), Mohammedia (3, *), 

Casablanca (21, 100), El Jadida (21, 100, *), Oualidia 

(*), Sidi Boulbra (6, 7), Essaouira (*). 

Fucales 

Cystoseiraceae 

Bifurcaria bifurcata R. Ross [= B. rotunda (Hudson) 

Papenfuss, B. tuberculata (Hudson) Stackhouse]. 

Without locality (8). A: Tanger (9, 38, 100, *), Cap 

Spartel (9), Salé (24, 33), Rabat (9, 11, 21, 24, 26, 38, 

98, 100, *), Temara (3, 11, 12, 33, 36, 37, 100, *), 

Bouznika (99, 100), Mohammedia (3, 71, 72, 81, 98, 

99, 100), Casablanca (21, 38, 39, 98, 100), El Jadida 

(5, 18, 21, 33, 38, 98, 100, *), Oualidia (*), Djorf el 

Yhoudi (98, 100), Sidi Boulbra (6, 7), Essaouira 

(39, 45, 98, 100, *), Imessouane (33, 100), Cap Ghir 

(33, 45, 98, 100, *), Agadir (21, *), Mirhleft (21, 33, 

98, 100, *), Sidi Ifni (*), Tan Tan (*), Moroccan Sahara 

(61, 77, 96), Cap Blanc (81), cosmopolitan (21, 

40), without locality (20, 22, 28, 32). 

Cystoseira abies-marina (S. G. Gmelin) C. Agardh [= 

Fucus abies-marinus S. G. Gmelin]. A: Rabat (38, 

81), Casablanca (38, 81), Moroccan Sahara (30). 

Cystoseira amentacea Bory var. stricta Montagne [= 

Cystoseira stricta (Montagne) Sauvageau]. M: 

Saaidia (50), Nador (15), Melilla (38, 50, 67, 91, 93, 

100), Al Hoceima (50), M’diq (*), Ksar-Sghir (*), 


Cystoseira baccata (S. G. Gmelin) P. C. Silva [= Fucus 

baccatus S. G. Gmelin, Cystoseira fibrosa (Hudson) 

Agardh, Fucus fibrosus Hudson]. M: Chafarinas Islands 

(49, 67). A: Tanger (9, 25, 38, 90, 100), Salé 

(33), Rabat (21, 100, *), Temara (12, 33, 99, 100, *), 

El Jadida (5, 18, 21, 25, 38, 98, 100, *), Safi (100), 

Djorf el Yhoudi (98), Essaouira (21, 33, 38, 39, 81, 

98, 100), Imessouane (33, 100), Cap Ghir (33), 

Agadir (21, 100, *), Mirhleft (21, 33, 98, 100), Moroccan 

Sahara (96), cosmopolitan (21, 22), without 

locality (20, 24, 38, 40, 56). 

Cystoseira brachycarpa J. Agardh var. balearica 

(Sauvageau) Giaccone [= C. balearica Sauvageau]. 

M: Nador (50), Melilla (50),Al Hoceima (50), without 


Cystoseira compressa (Esper) Gerloff et Nizamuddin 

[= Fucus compressus Esper, Cystoseira abrotanifolia 

(Desfontaine) C. Agardh, C. abrotanifolia C. 

Agardh f. fimbriata Sauvageau, C. fimbriata (Desfontaine) 

Bory]. M: Chafarinas Islands (14, 67), 

Nador (15, 38, 47, 51, 93), Melilla (10, 38, 48, 67, 93, 

100), Tamernoute (58), M’diq (*), Cabo Negro 

(58),Tetouan (67, *), Ksar-Sghir (58, *), without locality 

(49, 50). A: Tanger (9, 25, 38, 90, 100, *), Rabat 

(21, 38), Temara (3, 12, 36, 100, *), Bouznika 

(99, 100), Mohammedia (3, 71, *), Casablanca (21, 

25, 38, 81, 100), El Jadida (5, 18, 21, 25, 38, 100, *), 

Essaouira (9, 21, 25, 38, 100, *),Tan Tan (*), Moroccan 

Sahara (96), cosmopolitan (21), without locality 

(38, 40). 

Cystoseira crinita Duby. M: Nador (47, 50, 51), Melilla 

(48, 50), M’diq (*), without locality (49). A: Tanger 

(9, 25, 38, 100, *),Azilah (*). 

Cystoseira elegans Sauvageau. M: Saaidia (50, *), 

Melilla (50), Al Hoceima (50), Jabha (50, *), without 

locality (49). A: Rabat (18), El Jadida (18). 

Cystoseira foeniculacea (Linnaeus) Greville [= Fucus 

foeniculaceus Linnaeus, F. discors Linnaeus, Cystoseira 

discors (Linnaeus) C. Agardh emend. 

Sauvageau, Cystoseira ercegovicii Giaccone]. M: 

Nador (15, 50, 93), Melilla (10, 38, 48, 49, 50, 67, 

100), Ceuta (38, 49, 67, 100), without locality (49).

A: Tanger (38, 55, 93, 100), Rabat (99, 100), Temara 

(100), Casablanca (21, 38, 81, 100), El Jadida (21, 38, 

100), Oualidia (21, 38, 81), Safi (100), Moroccan Sahara 

(61), cosmopolitan (22), without locality (56). 

Cystoseira humilis Schousboe ex Kützing var. 

humilis. Without locality (8). M: Melilla (50), Al 

Hoceima (50), Jabha (50, *), M’diq (*), Without locality 

(49). A: Tanger (9, 38, 100), Cap Spartel (25, 

38, 90), Larache (21, 100), Rabat (100), Temara 

(21), Bouznika (100), El Jadida (18, 21, 98, 100), 

Safi (21, 100), Essaouira (21, 98, 100), Cap Ghir (98, 

100),Agadir (18, 21, 100). 

Cystoseira humilis Schousboe ex Kützing var. myriophylloides 

(Sauvageau) J. H. Price et D. M. John [= 

Cystoseira myriophylloides Sauvageau]. A: Tanger 

(38?, 100?), Rabat (21, 38, 100), Temara (3, 12, 36, 

66, 100, *), Mohammedia (3, 71, 72, *), Casablanca 

(21, 38, 81, 99, 100), El Jadida (5, 18, 98, 100), Oualidia 

(99), Djorf el Yhoudi (98), Essaouira (98, *), 

Agadir (21, 100), cosmopolitan (21, 22), without locality 

(20, 24, 40). 

Cystoseira mauritanica Sauvageau [= Cystoseira tingitana 

Sauvageau, C. selaginoides Valiante var. 

gibraltarica Sauvageau, C. ericoides (Linnaeus) C. 

Agardh var. gibraltarica Sauvageau nomen nudum, 

C. gibraltarica (Sauvageau) P. Dangeard]. M: 

Nador (38, 50, 93), Melilla (38, 50, 67, 93, 100), Al 

Hoceima (50), Jabha (50), Ceuta (38, 67, 91, 93), 

Ksar-Sghir (58), without locality (49). A: Tanger 

(38, 91, 93), Rabat (21), Temara (12), El Jadida 

(98), Djorf el Yhoudi (99), Essaouira (98), Agadir 

(18, 21), Mirhleft (21), cosmopolitan (21, 40), without 


Cystoseira mediterranea Sauvageau 13 . M: Saaidia 

(50), Chafarinas Islands (14, 50, 67), Nador (50, 51), 

Melilla (48), without locality (49). A: Tanger (25), 

Salé (18, 21, 40), Rabat (21, 25, 40, 100), Temara (3, 

12, 99, 100), Mohammedia (3), El Jadida (25), Safi 

(40), Essaouira (100). 

Cystoseira nodicaulis (Withering) M. Roberts [= Fucus 

nodicaulis Withering, Fucus granulatus Linnaeus, 

Cystoseira granulata (Linnaeus) C.Agardh]. 

M: Nador (15). A: Oualidia (21, 38, 81), Essaouira 

(21, 40, 100), cosmopolitan (22), without locality 

(24). 

Cystoseira platyclada Sauvageau 14 . M: Nador (15), 

Melilla (38, 49, 67, 91, 93, 100). 

Cystoseira sauvageauana G. Hamel var. sauvageauana 

15 [= C. selaginoides Valiante]. M: Nador (50), 

Melilla (50), Al Hoceima (50), Jabha (50, *), M’diq 

(*), without locality (49). A: Temara (100), El Jadida 

(18, 100), Essaouira (18). 

Cystoseira sauvageauana G. Hamel var. polyoedematis 

Sauvageau 15 . M: Nador (15). 

Cystoseira spinosa Sauvageau. M: Saaidia (50), Chafarinas 

Islands (50), Nador (15), Melilla (10, 48, 

50), without locality (49). 

Cystoseira tamariscifolia (Hudson) Papenfuss 13 [= 

Fucus tamariscifolius Hudson, Cystoseira ericoides 


(Linnaeus) C. Agardh, C. ericoides (Linnaeus) C. 

Agardh var. divaricata P. Dangeard, C. ericoides 

(Linnaeus) C. Agardh var. laevis P. Dangeard, Fucus 

ericoides Linnaeus]. Without locality (8). M: 

Chafarinas Islands (67), Nador (15), Melilla (48), 

Tamernoute (58), Cabo Negro (58), Ceuta (38, 67, 

93, 100), Ksar-Sghir (58), without locality (49, 50). 

A: Tanger (9, 38, 55, 90, 93, 100, *), Salé (24, 33), Rabat 

(9, 11, 18, 21, 24, 38, 100), Temara (3, 11, 12, 33, 

36, 37, 100, *), Bouznika (99, 100), Mohammedia 

(3, 71, 72, 81, 99, 100), Casablanca (21, 38, 81, 98, 

100), El Jadida (5, 21, 38, 81, 100, *), Oualidia (99, 

100, *), Djorf el Yhoudi (98, 100), Sidi Boulbra (6, 

7), Essaouira (21, 33, 100, *), Imessouane (33, 100), 

Cap Ghir (33, 98, 100), Agadir (18, 21, 100, *), 

Mirhleft (33, 98, 100), Tan Tan (*), Moroccan Sahara 

(33, 61, 96), cosmopolitan (22), without locality 

(20, 24, 38, 40, 56). 

Cystoseira usneoides (Linnaeus) M. Roberts [= Fucus 

usneoides Linnaeus, Cystoseira concatenata sensu 

C. Agardh pro parte]. M: Nador (15), Melilla (48, 

50),Al Hoceima (50), Jabha (50, *), M’diq (*), Ceuta 

(25, 67), without locality (8, 49). A: Tanger (9, 25, 

38, 42), Rabat (38, 81), Mohammedia (81), 

Casablanca (38, 81), El Jadida (25, 38), without locality 

(38). 

Cystoseira zosteroides (Turner) C. Agardh [= Fucus 

zosteroides Turner nom. illeg., Cystoseira opuntioides 

Bory ex Montagne]. M: Chafarinas Islands 

(35), Melilla (50), Al Hoceima (50), without locality 

(49). 

Fucaceae 

Fucus spiralis Linnaeus [= F. platycarpus Thuret 

nom. illeg., F. platycarpus Thuret f. linearis Schousboe 

ex Bornet, F. platycarpus Thuret f. minor Schousboe 

ex Bornet, F. platycarpus Thuret f. nana 

Schousboe ex Bornet, F. vesiculosus Linnaeus var. 

limitaneus Montagne, F. spiralis [var. platycarpus 

(Thuret) Batters] f. limitaneus (Montagne) Børgesen]. 

Without locality (8). M: Nador (15), Melilla 

(48, 50), Al Hoceima (50), Jabha (50), Tamernoute 

(58), Cabo Negro (58), Ceuta (25, 38, 50, 67, 91, 93, 

100), Ksar-Sghir (58), without locality (49). A: 

Tanger (9, 25, 38, 55, 100), Cap Spartel (9, 25, 38), 

Azilah (*), Salé (24, 33), Rabat (11, 21, 24, 38, 98, 

100), Temara (3, 11, 12, 33, 36, 37, 66, 100, *), 

Bouznika (100), Mohammedia (3, 71, 98, *), 

Casablanca (21, 38, 81, 100), El Jadida (5, 18, 21, 24, 

25, 33, 38, 40, 98, 100, *), Oualidia (*), Safi (40, 99, 

100), Djorf el Yhoudi (98, 100), Sidi Boulbra (6, 7), 

Essaouira (18, 22, 24, 33, 39, 40, 45, 98, 100, *), Imessouane 

(33, 100), Cap Ghir (33, 98), Mirhleft (33, 

98, 100, *), Moroccan Sahara (28, 31, 33, 61, 65, 77, 

96), cosmopolitan (21, 22), without locality (28, 32). 

Fucus vesiculosus Linnaeus [= F. axillaris J. Agardh 

nom. illeg., F. lutarius (Chauvin) Kützing, F. vesiculosus 

Linnaeus var. grandifrons C. Agardh]. M:


Nador (15, 50). A: Tanger (9, 25, 38, 100), Azilah 

(*), Larache (*), Kenitra (18, 21, 22, 66, *), Sale 

(100), Rabat (18, 21, 22, 38, 44, 63, 100), Temara (3, 

12, *), Casablanca (25, 38, 100), El Jadida (5, 38, 81, 

100), Oualidia (21, 38, 40, 81, 100), Safi (21, 38, 100), 

Djorf el Yhoudi (100), Essaouira (38, 100), without 

locality (28, 40). 

Halidrys siliquosa (Linnaeus) Lyngbye [= Fucus 

siliquosus Linnaeus]. A: Tanger (16). 

Sargassaceae 

Sargassum acinarium (Linnaeus) Setchell [= Fucus 

acinarius Linnaeus, Fucus linifolius Turner, Sargassum 

linifolium (Turner) C.Agardh, Sargassum linifolium 

(Turner) C. Agardh [var. amygdalifolium 

Montagne] f. gibraltarica Grunow]. M: Nador (50, 

51), Melilla (48), M’diq (*), without locality (49). 

A: Tanger (9, 25, 38), Rabat (38), Casablanca (38, 

81), Oualidia (81), without locality (56). 

Sargassum desfontainesii (Turner) C. Agardh [= Fucus 

desfontainesii Turner]. M: Melilla (48, 50), without 


Sargassum flavifolium Kützing [= S. vulgare C. 

Agardh var. flavifolium (Kützing) Grunow]. A: 

Casablanca (81). 

Sargassum furcatum Kützing. M: Chafarinas Islands 

(35). 

Sargassum hornschuchii C. Agardh. M: Nador (50), 

Melilla (48, 50), Al Hoceima (50), Jabha (*), M’diq 

(*),Tetouan (50, 67), without locality (49). 

Sargassum natans (Linnaeus) Gaillon [= Fucus 

natans Linnaeus, Fucus bacciferus Turner, Sargassum 

bacciferum (Turner) C.Agardh]. A: Casablanca 

(38, 81). 

Sargassum vulgare C.Agardh nom. illeg. 16 [= S. megalophyllum 

Montagne, S. vulgare C. Agardh var. 

megalophyllum (Montagne) Grunow]. Without locality 

(8). M: Saaidia (*), Chafarinas Islands (67), 

Nador (47, 51), Melilla (10, 48), M’diq (*), Cabo Negro 

(58),Ksar-Sghir (58,*),without locality (49,50). 

A: Tanger (9, 38, 40, 55, 100, *), Azilah (*), Larache 

(*), Rabat (21, 100), Temara (3, 12, 21, 99, 100, *), 

Bouznika (100),Mohammedia (3,72,*),Casablanca 

(21, 38, 81, 98, 100), El Jadida (5, 18, 21, 98, *), Safi 

(100), Djorf el Yhoudi (100), Essaouira (98, 100, *), 

Cap Ghir (98, 100), Agadir (100), Tan Tan (*), Moroccan 

Sahara (61,77),without locality (56). 

Laminariales 

Alariaceae 

Ecklonia muratii J. Feldmann. A: Mihrleft (*), Sidi 

Ifni (*), Tan Tan (*), Moroccan Sahara (30, 31, 60, 

61), Cap Blanc (28, 31). 

Laminariaceae 

Laminaria ochroleuca De La Pylaie [= L. iberica (G. 

Hamel) Lami, L. lejolisii Sauvageau, L. pallida 

Greville ex J. Agardh var. iberica G. Hamel]. Without 

locality (8). M: Nador (15), Jabha (*), M’diq 

(*). A: Tanger (9, 25, 27, 38, 98, 100), Azilah (*), 

Temara (36), El Jadida (5, 17, 18, 33, 38, 81, 98, 100, 

*), Oualidia (*), Safi (100), Djorf el Yhoudi (98, 

100), Sidi Boulbra (6, 7), Essaouira (17, 18, 27, 33, 

39, 98, 100, *), Imessouane (33, 98, 100, *), Cap Ghir 

(33, 98, 100, *), Mirhleft (98, 100, *), Tan Tan (*), 

Moroccan Sahara (33, 61, 64, 65, 77), Cap Blanc 

(98), without locality (27, 32, 40). 

Laminaria rodriguezii Bornet. M: Nador (15), Melilla 

(27, 38, 49, 67, 93, 100). 

Phyllariaceae 

Phyllariopsis brevipes (C. Agardh) E. C. Henry et 

South [= Laminaria brevipes C.Agardh, Laminaria 

reniformis J.V. Lamouroux, Phyllaria reniformis (J. 

V. Lamouroux) Rostafinski]. M: Chafarinas Islands 

(35, 38), Nador (15), Melilla (27, 38, 48, 50, 67, 

93, 100), Al Hoceima (50), Jabha (50), Ceuta (38, 

50, 67, 100), without locality (49). A: Tanger (9, 25, 

27, 38, 40, 55, 100, *), Azilah (*), Temara (21, 100), 

Bouznika (100), Casablanca (21, 38, 81, 100), El Jadida 

(18, 21, 25, 27, 38, 98, 100, *), Djorf el Yhoudi 

(98, 100), Essaouira (18, 21, 27, 81, 98, 100, *), Cap 

Ghir (98, 100), Mirhleft (21, 100, *), Moroccan Sahara 

(28, 31, 61), without locality (20, 28). 

Phyllariopsis purpurascens (C.Agardh) E.C.Henry et 

South [= Laminaria purpurascens C.Agardh, Phyllaria 

purpurascens (C. Agardh) Rostafinski]. Without 

locality (8).M: Nador (15),Melilla (27,38,49,67, 

93, 100). A: Tanger (9, 25, 27, 38, 100), Casablanca 

(21, 27, 38, 40, 81, 100), Essaouira (21, 40, 100), Cap 

Ghir (21,100),Agadir (18,21,40,100). 

Saccorhiza polyschides (Lightfoot) Batters [= Fucus 

polyschides Lightfoot, Saccorhiza bulbosa (Hudson) 

De La Pylaie]. Without locality (8). M: Nador 

(15),Melilla (27,38,48,50,67,93,100),Jabha (50,*), 

Ceuta (38, 50, 67, 100), Ksar-Sghir (58), without locality 

(49). A: Tanger (9, 25, 38, 100), Azilah (*), 

Larache (18,98,100),Salé (18,21,33,98),Rabat (18, 

21, 38, 98, 100), Temara (12, 33, 37, 100), Bouznika 

(100), Mohammedia (98, 100), Casablanca (21, 38, 

81, 100), El Jadida (5, 18, 21, 33, 38, 81, 98, 100, *), 

Oualidia (*), Safi (100), Djorf el Yhoudi (98, 100), 

Sidi Boulbra (6, 7), Essaouira (18, 27, 33, 38, 39, 81, 

98, 100, *), Imessouane (33, 100), Cap Ghir (98, 99, 

100), Agadir (99, 100), Mirhleft (21, 33, 98, 100, *), 

Moroccan Sahara (28, 33, 61, 64, 65, 68, 69, 77), Cap 

Blanc (98),without locality (20,27,28,32,40). 

Scytosiphonales 

Scytosiphonaceae 

Colpomenia peregrina Sauvageau. M: Melilla (50),Al 

Hoceima (50), Jabha (*), M’diq (*), Ksar-Sghir (*). 

A: Mirhleft (21).

Colpomenia sinuosa (Mertens ex Roth) Derbès et 

Solier [= Ulva sinuosa Mertens ex Roth]. Without 

locality (8). M: Chafarinas Islands (14), Nador (47, 

51), Melilla (10, 48), Jabha (*), Tamernoute (58), 

M’diq (*), Cabo Negro (58), Ksar-Sghir (58, *), 

without locality (50). A: Tanger (9, 25, 38, 100, *), 

Rabat (21, 38),Temara (3, 12, 100, *), Bouznika (99, 

100), Mohammedia (3, 71, 72, 99, 100, *), Casablanca 

(21, 38, 81, 100), El Jadida (5, 100), Djorf el 

Yhoudi (100), Cap Ghir (45, 100, *), Agadir (21, 

100, *),Tan Tan (*), without locality (40). 

Hydroclathrus clathratus (Bory ex C. Agardh) M. 

Howe [= Encoelium clathratum Bory ex C. 

Agardh]. A: Cap Ghir (45). 

Petalonia fascia (O. F. Müller) Kuntze [= Fucus fascia 

O. F. Müller, Laminaria debilis C. Agardh, Petalonia 

fascia (O. F. Müller) Kuntze var. debilis (C. 

Agardh) G. Hamel, Phyllitis debilis (C. Agardh) 

Kützing]. Without locality (8). M: Nador (50), 

Melilla (10, 48), Al Hoceima (50), Cabo Negro 

(58), Ksar-Sghir (58). A: Tanger (9?), Rabat (63), 

Temara (3, 12, *), Mohammedia (3, 71, *), El Jadida 

(5, *), Oualidia (*), without locality (40, 42). 

Scytosiphon lomentaria (Lyngbye) Link 17 [= Chorda 

lomentaria Lyngbye, Scytosiphon simplicissimus 

(Clemente) Cremades, Ulva simplicissima 

Clemente]. M: Saaidia (*), Nador (47, 50, 51), 

Melilla (10, 50), Al Hoceima (50), Jabha (50), 


Ksar-Sghir (58, *). A: Tanger (9, 25, 38, *), Temara 

(3), Mohammedia (3, 71), El Jadida (*), Oualidia 

(*),Agadir (*). 

Sphacelariales 

Choristocarpaceae 

Discosporangium mesarthrocarpum (Meneghini) 

Hauck [= Callithamnion mesarthrocarpum 

Meneghini]. M: Chafarinas Islands (14, 50, 67). 

Cladostephaceae 

Cladostephus spongiosus (Hudson) C. Agardh f. 

spongiosus [= Conferva spongiosa Hudson]. M: 

Melilla (10, 48). A: Tanger (9, 25, 38, 92, 100). 

Cladostephus spongiosus (Hudson) C. Agardh f. verticillatus 

(Lightfoot) Prud’homme van Reine [= 

Conferva verticillata Lightfoot, Cladostephus verticillatus 

(Lightfoot) C. Agardh, Fucus hirsutus Linnaeus, 

Cladostephus hirsutus (Linnaeus) 

Prud’homme van Reine]. Without locality (8). M: 

Saaidia (*), Chafarinas Islands (14), Tamernoute 

(58), Cabo Negro (58), Ceuta (25, 38, 100), Ksar- 

Sghir (58), without locality (50). A: Tanger (9, 25, 

38, 92, 100, *), Rabat (38), Temara (3, 21, 100, *), 

Mohammedia (3, 71, 72, 81, *), Casablanca (38, 

100), El Jadida (5, *) , Sidi Boulbra (6, 7), Essaouira 

(38, 81, 100), Cap Ghir (100), Agadir (21, 100, *), 

Mirhleft (21, *), without locality (40). 


Sphacelariaceae 

Sphacelaria brachygonia Montagne. M: Saaidia (50, 

*), Nador (50), Melilla (50), Jabha (*), M’diq (*). A: 

Tanger (78, 87), Azilah (*), Larache (*), Temara 

(*), Oualidia (*), Sidi Boulbra (6, 7), Essaouira (*), 

Agadir (*), Sidi Ifni (*). 

Sphacelaria cirrosa (Roth) C. Agardh [= Conferva 

cirrosa Roth, Sphacelaria hystrix Suhr ex Reinke]. 

M: Chafarinas Islands (14, 67), Melilla (48), 


Ksar-Sghir (58, *), without locality (50). A: Tanger 

(9, 25, 38, 78, 86, 87, 100), Larache (*), Temara (3, 

*), Mohammedia (3, 71, *), Casablanca (21, 38, 78, 

81, 100), El Jadida (*), Mirhleft (21, 100, *). 

Sphacelaria fusca (Hudson) S. F. Gray [= Conferva 

fusca Hudson]. Without locality (8). M: Nador (47, 

51), Melilla (48), Tamernoute (58), Cabo Negro 

(58), Ksar-Sghir (58), without locality (50). A: 

Larache (*), Moulay Bou Selham (42), Rabat (42), 

Temara (3, 12, *), Mohammedia (3, 71, *), El Jadida 

(5, *), Oualidia (*), Sidi Boulbra (6, 7), Essaouira 

(*),Agadir (*), without locality (40). 

Sphacelaria plumula Zanardini. M: Al Hoceima (50), 

Jabha (50). 

Sphacelaria radicans (Dillwyn) C. Agardh [= Conferva 

radicans Dillwyn]. A: El Jadida (21, 38, 81). 

Sphacelaria rigidula Kützing [= Sphacelaria furcigera 

Kützing]. M: Saaidia (*), Nador (47, 51), Melilla 

(48), Jabha (*), Tamernoute (58), M’diq (*), Cabo 

Negro (58), without locality (50). A: Tanger (78), 

Temara (*), Casablanca (78), El Jadida (*), Oualidia 

(*), Sidi Boulbra (6, 7), Essaouira (*), Sidi Ifni 

(*). 

Sphacelaria tribuloides Meneghini. M: Nador (50), 

Melilla (10, 48, 50, 78), Al Hoceima (50), Jabha (*), 

M’diq (*), Cabo Negro (58), Ceuta (78). A: Temara 

(3, *), Mohammedia (3, 71, *), Agadir (21, 100, *), 


Stypocaulaceae 

Halopteris filicina (Grateloup) Kützing [= Ceramium 

filicinum Grateloup]. Without locality (8). M: 

Saaidia (*), Chafarinas Islands (67), Nador (47, 

51), Melilla (10, 48), Jabha (*), Tamernoute (58), 

M’diq (*), Cabo Negro (58), Tetouan (67), Ksar- 

Sghir (58), without locality (50). A: Tanger (9, 25, 

38, 88, 100, *), Azilah (*), Rabat (21, 100), Temara 

(3, 12, 21, *), Mohammedia (3, 71, 72, 100, *), El Jadida 

(5, *), Agadir (*), Tan Tan (*), without locality 

(40). 

Halopteris scoparia (Linnaeus) Sauvageau [= Conferva 

scoparia Linnaeus, Stypocaulon scoparium 

(Linnaeus) Kützing]. Without locality (8). M: 

Saaidia (*), Chafarinas Islands (14), Nador (47, 

51), Melilla (10, 48), Tamernoute (58), M’diq (*), 

Cabo Negro (58), Ksar-Sghir (58, *), without locality 

(50). A: Tanger (9, 25, 38, 55, 89, 100), Azilah (*), 

Larache (*), Rabat (21, 38), Temara (3, 12, 100, *),


Mohammedia (3, 71, 72, 81, 100, *), Casablanca (21, 

38, 81, 100), El Jadida (5, 21, 81, 100, *), Oualidia 

(*), Sidi Boulbra (6, 7), Essaouira (*), Cap Ghir 

(100, *),Agadir (21, 100, *), Sidi Ifni (*), without locality 

(40). 

Sporochnales 

Sporochnaceae 

Carpomitra costata (Stackhouse) Batters var. costata 

[= Fucus costatus Stackhouse, Carpomitra cabrerae 

(Clemente) Kützing]. A: Tanger (9, 38). 

Carpomitra costata (Stackhouse) Batters var. 

mediterranea J. Feldmann. M: Ceuta (42). 

Nereia filiformis (J. Agardh) Zanardini [= Desmarestia 

filiformis J.Agardh]. A: Tanger (9, 25). 

Sporochnus gaertneri (S. G. Gmelin) C. Agardh [= 

Fucus gaertneri S. G. Gmelin]. A: Tanger (9, 25, 38). 

Sporochnus pedunculatus (Hudson) C. Agardh [= 

Fucus pedunculatus Hudson]. A: Tanger (9, 25, 38). 

Excludenda 

Cystoseira amentacea var. amentacea Bory. A: Tanger 

(9?), Rabat (11), Temara (11). The presence of this 

eastern Mediterranean taxon along the Atlantic 

coast of Morocco is unlikely. 

Himanthalia elongata (Linnaeus) S. F. Gray [= Fucus 

elongatus Linnaeus, Fucus loreus Linnaeus, Himanthalia 

lorea (Linnaeus) Lyngbye]. A: Tanger 

(9, 25, 38). Only drift material has been collected in 

Tanger. 

Laminaria saccharina (Linnaeus) J. V. Lamouroux [= 

Fucus saccharinus Linnaeus]. A: without locality 

(9?). 

Inquirenda 

Pilinia rimosa Kützing. A: Casablanca (21), El Jadida 

(21). According to Hooper et al. (1987) and O’Kelly 

(1989), the holotype of this species is a member 

of Phaeophyceae. However, its assignment to a 

definite species remains unclear (see Silva et al. 

1996, for a review). 

Sargassum coarctatum Kützing [= S. linifolium C. 

Agardh var. salicifolium J. Agardh]. A: Tanger (9, 

38), Casablanca (38, 81). This species, the type locality 

of which is the Gulf of Naples (Italy), is also 

observed in the Indian Ocean (Silva et al. 1996). It 

is considered to be a synonym of S. vulgare C. 

Agardh by Hamel (1939). 

Notes 

1 Levringia atlantica (J. Feldmann) Kylin may be conspecific 

with the earlier described L. brasiliensis (see Price 

et al. 1978). 

2 According to some authors Sauvageaugloia divaricata 

and S. griffithsiana may be the same entity (Guiry and 

Nic Dhonncha 2001). Pending new investigations, it 

seems preferable to maintain the distinction between 

these two taxa. 

3 ‘Aglaozonia chilosa Falkenberg’ life-history phase is the 

prostrate sporophytic phase of Cutleria chilosa (Falkenberg) 

P. C. Silva. 

4 ‘Aglaozonia melanoidea (Schousboe ex Bornet) 

Sauvageau’ life-history phase is the prostrate sporophytic 

phase of Cutleria adspersa (Mertens ex Roth) De Notaris. 

5 ‘Aglaozonia parvula (Greville) Zanardini’ life-history 

phase is the prostrate sporophytic phase of Cutleria multifida 

(J. E. Smith) Greville. 

6 According to Pedersen (1984: 60), Hecatonema terminale 

(Kützing) Kylin may be a synonym of H. maculans 

(Collins) Sauvageau, both being considered as a microthallus 

of well-known macroalgae belonging to the 

genus Asperococcus. 

7 According to Schnetter et al. (1987), Dictyota linearis (C. 

Agardh) Greville and Dictyota dichotoma (Hudson) J.V. 

Lamouroux var. intricata (C Agardh) Greville belong to 

the same entity. However, Ribera et al. (1992) and Silva 

et al. (1996) still recognize D. linearis. 

8 Guiry and Nic Dhonncha (2001) do not distinguish 

Taonia pseudociliata (J. V. Lamouroux) Nizamuddin et 

Godeh as distinct from Taonia atomaria (Woodward) J. 

Agardh. 

9 Feldmannia paradoxa (Montagne) G. Hamel var. paradoxa: 

according to Kim and Lee (1994), Feldmannia 

globifera is a synonym of F. paradoxa var. paradoxa. 

10 According to Silva et al. (1996), Bachelotia antillarum 

(Grunow) Gerloff and Bachelotia fulvescens (Bornet) 

Kuckuck ex G. Hamel may be two different taxa. Citations 

from Morocco all concern the latter entity. 

11 The presence of Pylaiella littoralis (Linnaeus) Kjellman, 

a cold water species, in Morocco needs to be confirmed. 

12 We follow the proposal of Rousseau (1999), which 

places Nemoderma tingitanum Schousboe ex Bornet 

within the Ralfsiaceae. 

13 Cystoseira mediterranea and C. tamariscifolia are considered 

as being vicariant species (Roberts 1978). For this 

reason, the co-occurrence of both species in the same locality 

(e.g.: Chafarinas Islands) requires a critical review 

of the records. 

14 The taxonomic status of this species is doubtful. Ribera 

et al. (1992) included C. platyclada in Taxa Inquirenda. 

15 According to Gómez Garreta et al. (2000), this taxon 

does not differ from the type variety. 

16 Sargassum vulgare, at the time of its publication, was superfluous 

and hence illegitimate since the species to 

which it applied included Fucus salicifolius S. G. Gmelin. 

However, this earlier entity cannot serve as a basionym 

because of the prior existence of S. salicifolium Naccari 

(see Silva et al. 1996). 

17 In accordance with Art. 56.1 of the International Code 

of Botanical Nomenclature (Greuter et al. 2000), the 

name Ulva simplicissima Clemente, and consequently 

the combination Scytosiphon simplicissimus (Clemente) 

Cremades, has been rejected (Pedersen and Kristiansen 

1994).

Discussion and Conclusion 

Although the marine flora of Morocco has not been extensively 

studied,as compared to that of countries such 

as the British Isles or Spain,the diversity of the Moroccan 

brown algae (131 species, infraspecific taxa and 

life-history phases) appears to be relatively high (Table 

II).The reason for this could reside in the fact that the 

Moroccan coasts belong to three biogeographical 

provinces: the western Mediterranean, the temperate 

Atlantic and the tropical Atlantic (Moroccan Sahara), 

the last of which remains poorly explored. 

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items items 

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