NEAT*Crustacea - TMBL

NEAT (North East Atlantic Taxa): 

South Scandinavian marine CRUSTACEA Check-List 

compiled at TMBL (Tjärnö Marine Biological Laboratory) by: 

Hans G. Hansson 1990-02-18 / small revisions until April 1996, 

when it for the 2:nd time was published on Internet, and again after some revisions August 1998. 

Citation suggested: Hansson, H.G. (Comp.), 1998. NEAT (North East Atlantic Taxa): South Scandinavian marine 

Crustacea Check-List. Internet pdf Ed., Aug. 1998. [http://www.tmbl.gu.se]. 

Denotations: () = Genotype @ = Associated to * = General note 

N.B.: This is one of several preliminary check-lists, covering S. Scandinavian marine animal (and partly marine 

protoctist) taxa. Some financial support from (or via) NKMB (Nordiskt Kollegium för Marin Biologi), during the last 

years of the existence of this organisation (until 1993), is thankfully acknowledged. The primary purpose of these 

checklists is to facilitate for everyone, trying to identify organisms from the area, to know which species that earlier 

have been encountered there, or in neighbouring areas. A secondary purpose is to facilitate for non-experts to find as 

correct names as possible for organisms, including names of authors and years of description. So far these checklists 

are very preliminary. Due to restricted access to literature there are (some known, and probably many unknown) 

omissions in the lists. Certainly also several errors may be found, especially regarding taxa like Plathelminthes and 

Nematoda, where the experience of the compiler is very rudimentary. The credibility of some other lists, containing 

taxa like e.g. Porifera, where, at least in certain subtaxa, taxonomic confusion seems to prevail, is certainly also 

suboptimal so far. 

Species found in South Scandinavia, as well as from neighbouring areas, chiefly the British Isles, have been 

considered, as some of them may show to have a slightly more northern distribution, than what is known today. 

However, species with a typical Lusitanian distribution, with their northern distribution limit around France or 

Southern British Isles, have as a rule been omitted here, albeit a few species with probable norhern limits around the 

British Isles are listed here until distribution patterns are better known. Arctic Taxa, known only from areas north of 

N Norway are probably not as well represented in this list as they should be.The compiler would be very grateful for 

Calantica Gray,1825 

C. (Scillaelepas Seguenza,1876) gemma C.W. Aurivillius,1894 

off W Iceland 

Scalpellinae Pilsbry,1916 

Scalpellum Leach,1818 

S. (Strictoscalpellum Broch,1924) scalpellum (Linnaeus,1767) 

Öresund-Bohuslän-N Norway-Norw. Sea-Faeroes-Br. Isles-W Africa-Mediterranean 

S. (Strictoscalpellum) stroemi M. Sars,1859 

Bohuslän-N Norway-Barents Sea-Spitsbergen-Iceland-Greenland-Faeroes-Shetlands 

S. (Strictoscalpellum) nymphocola Hoek,1883 

Shetlands-Norw. Sea-E Greenland-Spitsbergen & Arctic Seas 

S. (Strictoscalpellum) cornutum G.O. Sars,1879 

Faeroes-Norw. Sea-Spitsbergen & Arctic Seas 

S. (Strictoscalpellum) hamatum G.O. Sars,1879 

Shetlands-off Norway S of Lofoten-off SW Spitsbergen 

S. (Arcoscalpellum Hoek,1907) hispidum G.O. Sars,1890 

Trondheim-N Norway 

S. (Episcalpellum Broch,1924) striolatum G.O. Sars,1877 

off NE Faeroes-Norw. Sea-S Spitsbergen & Arctic Seas 

Pollicipes Leach,1817 

= Mitella Oken,1815 (nomen rejicendum, ICZN Op. 417) 

every correction of presumptive lapses and omittances an initiated reader could make. 

In the context below, Crustacea is ranked as a subphylum (within phylum Arthropoda) with the following 

superclasses: Maxillopoda, Remipedia, Phyllopoda (Branchiopoda), Ostracoda & Malacostraca. I am fully aware of 

P. pollicipes (Gmelin,1789) 

= Lepas pollicipes Gmelin,1789 

SW Cornwall - French & Iberian coasts 

the incompleteness of this list, but for the moment I cannot advance much further because of limited library resources. LEPADOMORPHA Pilsbry,1916 

Intentionally taxa from neighbouring areas have been listed, as some of them probably will show to occur in 

Scandinavian waters. 

Lepadoidea Darwin,1851 

N.B. This document is intended to be completely unavailable regarding nomenclature within the meaning of ICZN. 

Lepadidae Darwin,1851 

ARTHROPODA Siebold & Stannius,1845 

Lepas Linnaeus,1758 

L. (Lepas Pilsbry,1907) anatifera Linnaeus,1758 

CRUSTACEA Pennant,1777 (sensu Manton,1972) 

Öresund-Bohuslän-N Norway-Spitsbergen-Iceland-Faeroes-Shetlands-Br. Isles-southwards 

MAXILLOPODA Dahl,1956 

* Some authors are considering Ostracoda as part of this taxon. As this is not yet fully accepted, I have in this 

context chosen to treat Ostracoda as a taxon of the same rank as Maxillopoda. 

L. (Lepas) anserifera Linnaeus,1767 

Faeroe Channel-Norw. Sea; on ships to Denm. & Holland 

L. (Lepas) hilli (Leach,1818) 

carried by ships fr. tropical-subtropical waters to Öresund-Bohuslän-Lofoten-Faeroes-Shetlands-North Sea- 

MYSTACOCARIDA Pennak & Zinn,1943 

southwards 

DEROCHEILOCARIDA 

= MYSTACOCARIDIDA Pennak & Zinn,1943 

L. (Lepas) pectinata Spengler,1793 

carried by ships to Copenhagen, Bergen, Faeroes, S North Sea, by floatsome to Bohuslän 

Derocheilocaris Pennak & Zinn,1943 

L. (Dosima Gray,1825) fascicularis Ellis & Solander,1786 

on small floating objects from tropical and subtropical latitudes to Belt Sea-Bohuslän-North Sea-Trondheim- 

D. remanei Delamare Deboutteville & Chappuis,1951 biscayensis Delamare Deboutteville,1953 

Shetlands-Iceland 

Bay of Biscay-Portugal-Italy 

Anelasma Darwin,1851 

BRANCHIURA Thorell,1864 

A. squalicola (Lovén,1845) 

ARGULOIDA Rafinesque-Schmaltz,1815 

@ Etmopterus spinax 

Oslofjord-Skagerrak-Lofoten-W Br. Isles-Bay of Biscay 

Argulidae Leach,1819 

@ Chiefly on freshwater fish. Only marine genus is Argulus with 30-40 marine species 

Conchoderma Olfers,1814 

Argulus O.F. Müller,1785 

C. virgatum (Spengler,1790) 

* attatched to living and dead things, like ships, driftwood, sea-snakes, whales and big fishes 

A. foliaceus (Linnaeus,1758) 

Bohuslän & Copenhagen (ships), Oslofjord, Bergen, Lofoten, Iceland, Faeroes & S North Sea 

@ teleosts (e.g. Salmonidae, Gobidae & Gasterosteidae) (& frogs tadpole larvae) 

Freshwaters in Fennoscandia & Baltic estuarine water 

C. auritum (Linnaeus,1776) 

@ Coronula diadema (& C. reginae ) 

A. coregoni Thorell,18?? 

Bohuslän, Belgium, Lofoten-N Norway-Norw. Sea-Iceland-Faeroes-Shetlands-E Scotland-Atlantic 

@ teleosts (chiefly Salmonidae) 

Freshwaters in Fennoscandia & ?Baltic estuarine water 

Alepas Sander Rang,1829 

A. arcassonensis Cuénot,1912 

@ on skin of a diversity of fish and on skin of Sepia officinalis 

Arcachon 

A. parasita Sander Rang,1829 

@ medusae (Cyanea ) 

?Falmouth, Atlantic Ocean & Mediterranean 

THECOSTRACA Gruvel,1905 

CIRRIPEDIA Burmeister,1834 

ACROTHORACICA Gruvel,1905 

CYPRILEPADOMORPHA 

IBLOMORPHA 

HETERALEPADOMORPHA 

PYGOPHORIDA Berndt,1907 

PRAELEPADOMORPHA 

Lithoglyptidae Aurivillius,1892 

SESSILIA de Lamarck,1818 

Weltneria Berndt,1907 

BRACHYLEPADOMORPHA Withers,1923 

* One Pacific species, else Extinct. 

W. exargilla Newman,1974 

Bay of Biscay at ca 1500 m 

VERRUCOMORPHA Pilsbry,1916 

Cryptophialidae Gerstaecker,1866 

Verrucidae Darwin,1854 

* No taxa within the area? 

Verruca Schumacher,1817 

APYGOPHORIDA Berndt,1907 

V. stroemia (O.F. Müller,1776) 

Trypetesidae Stebbing,1910 

Öresund-Bohuslän-North Sea-N Norway-Barents Sea-Spitsbergen-E Greenland-Iceland-Faeroes-Shetlands- 

Br. Isles-Portugal-Mediterranean 

Trypetesa Norman,1903 

= Alcippe Hancock,1849, non Blyth,1844 

BALANOMORPHA Pilsbry,1916 

T. lampas (Hancock,1849) 

Chthamaloidea Darwin,1854 

Öresund-Bohuslän-S Norway-North Sea-Br. Isles, W Mediterranean 

Chthamalidae Darwin,1854 

THORACICA Darwin,1854 

Chthamalinae Darwin,1854 

PEDUNCULATA de Lamarck,1818 

Chthamalus Ranzani,1817 

SCALPELLOMORPHA 

C. stellatus (Poli,1791) 

Scalpelloidea Pilsbry,1916 

Anholt, Helgoland, Faeroes-Shetlands-W Br. Isles-Bretagne-Canaries-Cap Verde-Mediterranean 

Scalpellidae Pilsbry,1916 

C. montagui Southward,1976 

= C. punctatus : Montagu,1803, non (Martini & Chemnitz,1788) 

Calanticinae 

Bay of St Malo-Br. Isles-Bay of Biscay-W Morocco 

Euraphiinae Newman & Ross,1976 

1

* Not in the area. 

Notochthamalinae Foster & Newman,1987 


Catophragmidae Utinomi,1968 


Coronuloidea Leach,1817 

Coronulidae Leach,1817 

Coronula de Lamarck,1802 

C. diadema (Linnaeus,1767) 

@ whales, often Megaptera novaeangliae (& e.g. Hyperoodon ) 

N Norway-Barents Sea, Iceland-Faeroes-Shetlands 

C. reginae Darwin,1854 

@ whales, often Megaptera novaeangliae (& e.g. Balaenoptera spp.) 

Skagerrak, N Norway, Iceland 

Xenobalanus Steenstrup,1852 

P. anglica G.B. Sowerby II,1823 

@ Caryophyllia 

Br. Isles-Ireland-W France-W Africa-Cap Verde-Mediterranean 

Ceratoconchinae Newman & Ross,1976 


Megatrematinae Holthuis,1982 


Balanidae Leach,1817 

Balaninae Leach,1817 

Balanus da Costa,1778 

B. balanus (Linnaeus,1758) 

SW Baltic-Bohuslän-North Sea-Norway-Barents Sea & Spitsbergen-Arctic Seas-Iceland-Faeroes-Shetlands- 

Br. Isles-NW Portugal 

B. crenatus Bruguière,1789 

Blekinge-Bohuslän-North Sea-Norway-Barents Sea-Spitsbergen-Arctic Seas-Iceland-Faeroes-Br. Isles- 

Azores-NW Spain-W Algeria 

X. globicipitis Steenstrup,1852 

@ whales, ( e.g. Balaenoptera spp., Globicephala melaena, Orcinus orca, Delphinus delphis, Tursiops 

truncatus & Grampus griseus) 

B. improvisus Darwin,1854 

Bothnian Sea-Bohuslän-North Sea-Bergen-W Br. Isles-Morocco-E Mediterranean-Black Sea 

Platylepadidae Newman & Ross,1976 

B. spongicola Brown,1844 

= B. dollfusi Broch,1927 

Platylepas J.E. Gray,1825 

S Wales-SW England-Portugal-S Africa-Azores-Madeira 

P. hexastylos (O. Fabricius,1798) 

= Lepas hexastylos O. Fabricius,1798 

B. perforatus perforatus Bruguière,1789 

SW Br. Isles-France-Spain-NW Africa-Mediterranean-Black Sea 

@ a variety of sirenian & chelonian hosts, e.g. Chelonia mydas & Caretta caretta, but not Dermochelys 

coriacea, while the somewhat suspect "species" P. multidecorata is thought to be confined to the last species, 

which may indicate that it may be a synonym of P. hexastylos. 

B. perforatus angustus (Gmelin,1789) 

SW Br. Isles-France-Spain-Mediterranean Africa 

?Britain & warmer seas B. perforatus fistulosus (Poli,1791) 

?Denmark, Morocco, Canaries 

Stomatolepas Pilsbry,1910 

B. perforatus mirabilis Darwin,1854 

S. elegans (Costa,1838) 

La Rochelle (France) 

@ embedded in the soft skin portion of chelonians, chiefly Dermochelys coriacea & Lepidichelys olivacea 

(non-European), but also occasionally Chelonia mydas & Caretta caretta. 

Norway, Netherlands, Scotland, England, Eire - temperate & warmer seas 

B. amphitrite amphitrite Darwin,1854 

occasionally on ships in Scandinavia; Iberian Peninsula-southw. 

N Norway, Faeroes, Shetlands, Bay of Biscay, W Mediterranean 

Solidobalanus Hoek,1913 

Chelonibiidae Pilsbry,1916 

* Correct subfamily? 


S. fallax (Broch,1927) 

Emersoniidae Ross,1967 


= Balanus fallax Broch,1927 

* Epizoic on upper shells of Aequipecten opercularis. 

W Engl. Channel, Algeria-Angola 

Tetraclitoidea Gruvel,1903 

Megabalaninae Newman,1979 

Tetraclitidae Gruvel,1903 


Megabalanus Hoek,1913 

Tetraclitinae Gruvel,1903 

M. tintinnabulum (Linnaeus,1758) 

occasionally on ships in Scandinavia; Mediterranean-W & S Africa 

Tetraclitellinae Newman & Ross,1976 M. nigrescens (Lamarck,1818) 

occasionally on ships in Scandinavia; Australia 

Austrobalaninae Newman & Ross,1976 

Bathylasmatidae Newman & Ross,1971 

RHIZOCEPHALA F. Müller,1862 

Bathylasmatinae Newman & Ross,1971 Briarosaccus ??,1??? 

* Which taxon? 

Bathylasma Neman & Ross,1971 

B. callosus ?,1??? 

B. hirsutum (Hoek,1883) @ a variety of lithodids 

off Faeroes-southwards in Atlantic-Azores cosmopolitan 

Hexelasmatinae Newman & Ross,1976 


KENTROGONIDA Delage,1884 

Balanoidea Leach,1817 

Peltogastridae Lilljeborg,1861 

Peltogaster Rathke,1842 

Archaeobalanidae Newman & Ross,1976 

P. paguri Rathke,1842 ( - Gåsörännan, Bohuslän) 

Archaeobalaninae Newman & Ross,1976 

Chirona Gray,1835 

C. hameri (Ascanius,1767) 

= Lepas hameri Ascanius,1767 

= Lepas tulipa O.F. Müller,1776 

= Balanus candidus Brown,1827 

Kattegatt-Bohuslän-North Sea-Norway-Barents Sea-Iceland-Faeroes-Shetlands-Br. Isles 

C. amaryllis (Darwin,1854) 

= Balanus amaryllis Darwin,1854 

Indian Ocean species brought to Europe (?Britain) on ships' bottoms 

Acasta Leach,1817 

A. spongites (Poli,1791) 

@ embedded in Dysidea fragilis 

Br. Isles-France-Portugal-S Africa-Mediterranean 

Elminiinae Foster,1982 

Elminius Leach,1825 

E. modestus Darwin,1854 

Bohuslän (on floatsome)-W Jutland-Engl. Channel-Br. Isles-Bay of Biscay-NW Spain (immigated fr. 

Australia-New Zeeland area) 

Semibalaninae Newman & Ross,1976 

Semibalanus Pilsbry,1916 

S. balanoides (Linnaeus,1767) 

Öresund & Belt Sea-Bohuslän-North Sea-Norway-Barents Sea-Spitsbergen-Arctic Seas-Iceland-Faeroes-Br. 

Isles-NW Portugal 

Pyrgomatidae Gray,1825 

Pyrgomatinae Gray,1825 

=? Bosciinae Newman & Ross,1976 

Pyrgoma Leach,1818 ex Savigny MS 

=? Boscia Férussac,1822, non Leach,1814 (Amphipoda) 

@ Pagurus bernhardus, P. cuanensis, Anapagurus chiroacanthus, A. laevis & some foreign species 

Öresund-Bohuslän-North Sea-N Norway-Spitsbergen, W Ireland, Br. Isles-Bay of Biscay & W 

Mediterranean 

P. curvatus Kossmann,1874 

@ Pagurus cuanensis, P. prideaux & P. alatus 

N Kattegatt-Bohuslän-Sogn, SW Ireland, SW Br. Isles-W Mediterranean 

Peltogasterella Krüger,1912 ( P. socialis Krüger,1912, non Müller,1862 

= P. gracilis (Boschma,1927)) 

P. sulcata (Lilljeborg,1859) 

= Peltogaster sulcatus Lilljeborg,1859 

= Peltogaster microstoma Lilljeborg,1859 

= Peltogaster socialis Müller,1862 

@ Pagurus cuanensis, P. prideaux, P. alatus, Anapagurus laevis, A. chiroacanthus & A. hyndmanni 

Kattegatt-Bohuslän-Bergen, W Ireland, W Scotland, SW Br. Isles-Canaries-Mediterranean 

Galatheascus Boschma,1929 

G. striatus Boschma,1929 ( - Isle of Man) 

= G. minutus Boschma,1933 

@ Galathea strigosa, G. intermedia, G. nexa & G. dispersa 

Öresund-Bohuslän-Bergen, W Ireland, W Britain, Engl. Channel-W Mediterranean 

Tortugaster Reinhard,1948 ( T. fistulatus Reinhard,1948) 

T. boschmai (Brinkmann,1936) 

= Triangulus boschmai Brinkmann,1936 

@ Munida tenuimana, M. sarsi (& M. rugosa ) 

Skagerrak-W Norway 

Parthenopea Kossmann,1874 

P. subterranea Kossmann,1874 ( - Gulf of Naples) 

@ Calocarides coronatus & Callianassa subterranea 

Skagerrak, Mediterranean & Black Sea 

Trachelosaccus Boschma,1928 

T. hymenodorae (G.O. Sars,1879) 

= Sylon hymenodorae G.O. Sars,1879 ( - Norwegian Sea, in 3350m) 

@ Hymenodora glacialis 

2

Norw. Sea C. delagei J. Bocquet-Védrine,1957 ( - Roscoff) 

@ Chtamalus stellatus 

Cyphosaccus Reinhard,1958 ( C. chacei Reinhard,1958) Roscoff 

C. norvegicus Boschma,1962 Boschmaella J. Bocquet-Védrine,1968 

@ Munidopsis serricornis (Lovén) = Microgaster J. Bocquet-Védrine,1967, non Latreille,1804, nec Sweinson,1839, nec Doumerc,1855 

Bohuslän-Bergen-Trondheim-Vesterålen 

B. balani (J. Bocquet-Védrine,1967) 

Septosaccus Duboscq,1912 = Microgaster balani J. Bocquet-Védrine,1967 ( - Arcachon) 

@ Balanus improvisus 

S. rodriguezi (Fraisse,1876) Arcachon 

@ ? 

Roscoff Duplorbidae 

Sacculinidae Lilljeborg,1861 Duplorbis Smith,1906 

Sacculina Thompson,1836 

D. calathurae Smith,1906 ( - Forsblads Fjord, E Greenland) 

@ Calathura brachiata 

S. carcini Thompson,1836 ( - England) E Greenland 

@ Portunidae & Pirimelidae 

SW Baltic-Bohuslän-S North Sea-Br. Isles-Bay of Biscay-Mediterranean-Black Sea Cryptogaster ?,19?? 

S. inflata Leuckart,1859 Arcturosaccus ?,19?? 

= S. hyadis Malm,1881 

@ Hyas araneus & H. coarctatus Thompsoniidae Høeg & Rybakov,1992 

Kattegatt-Bohuslän-Helgoland * No NE Atlantic species 

S. triangularis Anderson,1862 

@ Cancer pagurus 

Thylacoplethus Coutière,1902 ( Thylacoplethus edwardsi Coutière,1902 - Australia) 

Bohuslän, British Isles-Ireland-Bretagne Diplothylacus Høeg & Lützen,1993 ( Thompsonia sinensis Keppen,1877 - China) 

S. gibbsi (Hesse,1867) 

= Sacculinidia gibbsi Hesse,1867 

Thompsonia Kossmann,1874 ( Thompsonia globosa Kossmann,1874 - Philippines) 

= Sacculina pisae Hoek,1888 

@ Pisa spp. 

Pottsia Høeg & Lützen,1993 ( Pottsia plesionikae Høeg & Lützen,1993 - Florida) 

Engl. Channel, Mediterranean 

Mycetomorphidae 

S. atlantica Boschma,1927 

@ Dorhynchus thomsoni 

Mycetomorpha Potts,1912 

off SW Ireland, in ≈1200m 

Incertae sedis 

S. lobata Boschma,1965 

@ Ebalia tuberosa 

Polysaccus Høeg & Lützen,1993 

NW Bretagne 

P. mediterraneus (Caroli,1929) 

S. bourdoni Boschma,1960 = Thompsonia mediterranea Caroli,1929 () 

@ Xantho pilipes @ Callianassa subterranea (Montagu) & C. truncata Giard & Bonnier 

W Bretagne Mediterranean 

S. gerbei Bonnier,1887 

@ Xantho incisus, Ethusa mascarone, Pilumnus hirtellus & Atelecyclus rotundatus 

Pirusaccus Lützen,1985 

W Br. Isles-W Engl. Channel-Bay of Biscay-Adriatic FACETOTECTA Grygier,1984 

S. gonoplaxae Guèrin-Ganivet,1911 

@ Goneplax angulata 

Gulf of Cadiz 

* Inhabiting "Hansen's Y-larvae", puzzling youth stages from e.g. 

Scandinavian waters of presumed Cirripedian relatives. 

ASCOTHORACIDA Lacaze-Duthiers,1880 

S. sp. 

* not referrable to any of the Sacculina spp. above SYNAGOGOIDA Vagin,1976 

@ Eurynome aspera @ Echinodermata, except 2 species of Synagoga, which occur on Anthozoa 

Väderöarna (Bohuslän) 

Synagogidae Gruvel,1905 

Drepanorchis Boschma,1927 

Ascothorax Djakonov,1914 

D. neglecta (Fraisse,1877) 

= Sacculina neglecta Fraisse,1877 ( - Gulf of Naples) A. ophioctenis Djakonov,1914 () 

= Sacculina phalangi Hoek,1878 @ bursae of Ophiocten sericeum 

= Sacculina stenorhynchi Malm,1881 Faeroes-Iceland-Barents Sea, Kara Sea 

@ Macropodia spp. & Inachus spp. 

Öresund-Bohuslän-Trondheim-Faeroes-S North Sea-Br. Isles-Gibraltar-Adriatic A. pilocaudatus Grygier,1983 

@ bursae of Ophiosphalma armigerum 

Lernaeodiscidae Boschma,1928 NE Atlantic 

Lernaeodiscus F. Müller,1862 ( L. porcellanae F. Müller,1862) Isidascus Moyse,1983 

L. ingolfi Boschma,1928 I. bassindalei Moyse,1983 () 

@ Munida sarsi, (M. tenuimana & M. rugosa ) @ Acanella arbuscula (Johnson) (a gorgonian coral) 

Oslofjord-Bergen-Senja-off SW Iceland-W Br. Isles NE Atlantic at 2000 m. 

L. squamiferae Pérez,1922 Synagoga Norman,1888 

@ Galathea squamifera 

Norway, Canaries, French Mediterranean S. mira Norman,1888 () 

@ Antipathes larix Ellis 

Triangulus Smith,1906 off Naples 

T. munidae Smith,1906 ( - Sætreosen, W Norway) 

= Lernaeodiscus munidae : Auctt. 

Dendrogasteridae Gruvel,1905 

@ Munida sarsi, M. tenuimana & M. rugosa 

Bohuslän-Lofoten-Shetlands-Irish Sea-Canaries 

Ulophysema Brattström,1936 

T. galatheae (Norman & Scott,1906) 

= Sacculina galatheae Norman & Scott,1906 

= Lernaeodiscus galatheae : Auctt. 

U. oeresundense Brattström,1936 () 

@ Brissopsis lyrifera, Echinocardium cordatum & E. flavescens 

Öresund-Bohuslän-W Norway 

@ G. intermedia, (G. nexa & G. dispersa ) 

Kattegatt-Bohuslän-Bergen-Br. Isles-Engl. Channel-Bay of Biscay-W Mediterranean 

U. pourtalesiae Brattström,1937 

@ coelomic cavity of Pourtalesia jeffreysi 

"T. perezi " Veillet,194? (nomen nudum) 

Norw. Sea (between Norway, Spitsbergen, Greenland & Faeroes) 

@ Galathea squamifera 

Bretagne 

Dendrogaster Knipowitsch,1891 

AKENTROGONIDA Häfele,1911 

D. astericola Knipowitsch,1891 () 

@ coelomic cavity ofHenricia 

Clistosaccidae Boschma,1928 

White Sea 

= Sylonidae Boschma,1928 D. murmanensis Kluge,1910 

@ coelomic cavity of Crossaster papposus & Solaster endeca 

Clistosaccus Lilljeborg,1860 Kolafjord 

C. paguri Lilljeborg,1860 ( - Humlesäcken, Bohuslän) 

= Apeltes paguri Lilljeborg,1860 

@ Pagurus bernhardus, P. pubescens & some foreign species 

Öresund-Bohuslän-N Norway-Spitsbergen-Iceland-Faeroes-Greenland 

D. hymenasteri Stone,1987 

@ coelomic cavity of Hymenaster membranaceus 

NE Atlantic 

Sylon Krøyer,1855 

D. pontasteri Stone,1987 

@ coelomic cavity of Pontaster tenuispinus 

S. hippolytes M. Sars,1870 ( - Drøbak) 

NE Atlantic 

= S. pandali M. Sars,1870 

= S. schneideri Hoek,1888 

D. psilasteri Stone,1987 

@ coelomic cavity of Psilaster andromeda 

= S. challengeri Hoek,1888 

@ Hippolytidae, Pandalidae & Crangon allmani 

N Kattegatt-Bohuslän-N Norway-Spitsbergen-Iceland-Greenland-Shetlands 

Chthamalophilidae J. Bocquet-Védrine,1961 

Chthamalophilus J. Bocquet-Védrine,1957 

NE Atlantic 

D. zoroasteri Stone,1987 

@ coelomic cavity of Zoroaster fulgens 

NE Atlantic 

D. arbusculus Fisher,1911 

@ coelomic cavity of Hippasteria californica & ?H. phrygiana 

3

California & ?St Lawrence Estuary * Bottom dwellers 

Myriocladus Okada,1925 ( Dendrogaster arborescens le Roi,1905 - S Africa @ Dipsacaster sladeni ) Platycopia G.O. Sars,1911 

LAUROIDIDA Vagin,1976 

@ Anthozoa 

Lauridae Gruvel,1905 

Laura Lacaze-Duthiers,1866 (short note & 1883) 

L. gerardiae Lacaze-Duthiers,1866 (short note & 1883) () 

@ in gall-like buds in the antipatharian Gerardia lamarcki (Milne-Edwards & Haine) 

Mediterranean 

Baccalaureus Broch,1929 ( B. japonicus Broch,1929 - Japan @ Zoantharia ) 

Ctenosculidae Thiele,1925 

Petrarcidae Gruvel,1905 

Petrarca Fowler,1889 ( P. bathyactidis Fowler,1889 - deep Pacific @ Bathyactis symmetrica ) 

P. perplexa G.O. Sars,1911 () 

S Norway (Korshavn) 

P. pygmaea G.O. Sars,1921 

S Norway (Korshavn) 

NEOCOPEPODA Huys & Boxshall,1991 

GYMNOPLEA Giesbrecht,1882 

CALANOIDA G.O. Sars,1903 

Megacalanoidea Sewell,1947 

Calanidae Dana,1849 

Calanus Leach,1816 

= Cetochilus Claus,1863 

C. hyperboreus Krøyer,1838 

TANTULOCARIDA Boxshall & Lincoln,1983 Kattegatt-Bohuslän-North Sea-W Norway-Norw. Sea-Iceland-Faeroes-Shetlands-Ireland 

* Zool. Rec. covered until December 1997. 

C. helgolandicus (Claus,1863) 

TANTULOCARIDIDA Boxshall & Lincoln,1983 = Cetochilus helgolandicus Claus,1863 

Cumoniscus Bonnier,1903 ( C. kruppi Bonnier,1903 @ Cumacea, Leuconidae - Biscay) 

* Which family? 

Kattegatt-Bohuslän-North Sea-W Norway-(Spitsbergen)-Iceland-Faeroes-Shetlands-Br. Isles-Mediterranean- 

Black Sea-Azores-Canaries 

C. finmarchicus (Gunnerus,1765) 

Deoterthridae Boxshall & Lincoln,1987, emend. Huys,1990 = Monoculus finmarchicus () 

@ Primarily copepods Baltic-Bohuslän-North Sea-W Norway-Norw. Sea-Arctic Seas-Iceland-Faeroes-Shetlands-Br. Isles-N 

Portugal-NE Atlantic 

Boreotantulus Huys & Boxshall,1988 

C. glacialis Jaschnov,1955 

B. kunzi Huys & Boxshall,1988 N Norway-Faeroes-Iceland-Arctic Seas 

@ Cylindropsyllus laevis 

Bohuslän 

Tantulacus Huys, Andersen & Kristensen,1992 

T. hoegi Huys, Andersen & Kristensen,1992 

@ not known 

Faeroes 

C. minor Claus,1863 ( of Nannocalanus G.O. Sars,1925) 

= Calanus valgus Brady,1883 

Cold & temperate Atlantic, Mediterranean 

C. propinquus Brady,1883 

?NE Atlantic 

Calanoides Brady,1883 

Amphitantulus Boxshall & Vader,1993 C. carinatus (Krøyer,1849) 

= Calanus brevicornis Lubbock,1856 

A. harpiniacheres Boxshall & Vader,1993 () * Atlantic deep water species 

@ Harpinia antennaria N Atlantic 

North Sea 

Mesocalanus Bradford & Jillet,1974 

Deoterthron Bradford & Hewitt,1980 ( D. dentatum Bradford & Hewitt,1980) 

= Austrotantulus Boxshall,1988 ( A. lincolni Boxshall,1988 @ Stenhelia - Tasmania) M. tenuicornis (Dana,1849) 

Campyloxiphos Huys,1990 ( C. dineti Huys,1990 @ the harpacticoid Peresime abyssalis - SE 

Atlantic) 

= Calanus tenuicornis Dana,1849 

= Neocalanus tenuicornis : G.O. Sars,1925 

North Sea-Faeroes-Shetlands-Ireland-NE Atlantic-Mediterranean 

Dicrotrichura Huys,1989 ( D. tricincta Huys,1989 - Corsica) 

Neocalanus G.O. Sars,1925 

Coralliotantulus Huys,1990 ( C. coomansi Huys,1990 - Queensland) 

N. gracilis (Dana,1849) 

= Calanus gracilis Dana,1849 

Aphotocentor Huys,1991 ( A. styx Huys,1991 - Corsica) 

?S North Sea, Ireland-Biscay, Mediterranean 

Itoitantulus Huys, Ohtsuka & Boxshall,1992 ( I. misophricola Huys, Ohtsuka & Boxshall,1992 @ 

Misophriopsis okinawensis - Japan) 

N. robustior (Giesbrecht,1888) 

= C. robustior Giesbrecht,1888 

Coald & temperate Atlantic, Mediterranean 

Microdajidae Boxshall & Lincoln, 1987 

Megacalanidae Sewell,1947 

Microdajus Greve,1965 

Megacalanus Wolfenden,1904 

M. langi Greve,1965 

@ Typhlotanais aequiremis, Leptognathia breviremis & Haplocope angusta 

W Norway 

M. princeps Wolfenden,1904 

= Macrocalanus longicornis G.O. Sars,1905 

Iceland-Faeroes-Ireland-NE Atlantic 

M. gaelicus Boxshall & Lincoln, 1987 

@ Typlotanais pulcher 

Bathycalanus G.O. Sars,1905 

North Sea 

B. princeps (Brady,1883) 

M. pectinatus Boxshall, Huys & Lincoln,1989 

Iceland-Faeroes-Shetlands-Ireland-NE Atlantic 

@ Typhlotanais 

NE Atlantic 

B. bradyi (Wolfenden,1905) 


Xenalytus Huys,1991 ( X. scotophilus Huys,1991 - Corsica) B richardi G.O. Sars,1905 

* Is not this and B. bradyi (Wolfenden) synonyms? If this is the case, which of them has priority? 

Onceroxenidae Huys,1990 NE Atlantic 

Onceroxenus Boxshall & Lincoln,1987 Bradycalanus A. Scott,1909 

O. curtus Boxshall & Lincoln,1987 B. sarsi (Farran,1939) 

@ Leptognathia zezinae NE Atlantic 

North Sea 

Paracalanidae Giesbrecht,1892 

O. birdi Boxshall & Lincoln,1987 

@ Paranarthrura insignis Paracalanus Boeck,1864 

North Sea 

P. parvus (Claus,1863) 

Basipodellidae Boxshall & Lincoln,1983, emend. Huys,1990 Baltic-Bohuslän-North Sea-W Norway-Norw. Sea-Iceland-Faeroes-Shetlands-Br. Isles-Engl. Channel-NE 

Atlantic 

Basipodella Becker,1975 ( B. harpacticola Becker,1975) 

P. pygmaeus (Claus,1863) 

Stygotantulus Boxshall & Huys,1989 ( S. stocki Boxshall & Huys,1989 @ harpacticoida - Canaries) * May be a synonym of P. parvus (Claus) 

NE Atlantic 

Nipponotantulus Huys, Ohtsuka & Boxshall,1994 ( N. heteroxenus Huys, Ohtsuka & Boxshall,1994 

@ Pseudocyclopiidae - Japan) P. nanus G.O. Sars,1907 

?NE Atlantic 

Doryphallophoridae Huys,1990 

Calocalanidae Bernard,1958 

Doryphallophora Huys,1990 

Calocalanus Giesbrecht,1888 

D. harrisoni (Boxshall & Lincoln,1987) 

= Deoterhron harrisoni Boxshall & Lincoln,1987 () C. contractus Farran,1926 

@ Macrostylis magnifica NE Atlantic-Biscay-Mediterranean 

North Sea 

C. pavo (Dana,1849) 

COPEPODA H. Milne-Edwards,1840 Atlantic 60° N- 40° S 

PROGYMNOPLEA Lang,1948 

PLATYCOPIOIDA Fosshagen,1985 

Platycopiidae G.O. Sars,1911 

C. pseudocontractus Bernard,1958 

S & W Ireland, NE Atlantic, Mediterranean 

C. styliremis Giesbrecht,1888 

Worldwide distribution except polar sea 

4

5 

Ischnocalanus Bernard,1963 

I. plumulosus (Claus,1863) 

C. reducta Brodsky,1950 

Central part of deep Arctic Ocean 

Temperate-tropical Atlantic, Mediterranean Aetideinae Giesbrecht,1892 

Mecynoceridae Andronov,1973 Gaidius Giesbrecht,1895 

Mecynocera I.C. Thompson,1888 

G. tenuispinus (G.O. Sars,1900) 

= Chiridius tenuispinus G.O. Sars,1900 

M. clausii I.C. Thompson,1888 

N North Sea-Norw. Sea-Barents Sea-Iceland-Faeroes-Shetlands-Ireland-NE Atlantic 

* Oceanic epiplankton 

NE Atlantic 

G. brevispinus (G.O. Sars,1900) 

= Chiridius brevispinus G.O. Sars,1900 

Eucalanoidea Giesbrecht,1892 

N North Sea-Norw. Sea-Arctic Seas-Iceland-Faeroes-Ireland-NE Atlantic 

Eucalanidae Giesbrecht,1892 

G. affinis G.O. Sars,1905 

S & W Ireland, NE Atlantic 

Eucalanus Dana,1853 ( E. attenuatus (Dana,1849) - W. Pacific) 

G. minutus G.O. Sars,1905 

E. elongatus (Dana,1849) 

NE Atlantic 

* According to Fleminger,1973, E. elongatus (Dana) is a tropical Indian Ocean to W Pacific species, while E. 

hyalinus (Claus,1866) is tropical-subtropical circumglobal, i.e. the only species in the elongatus group, 

thought to be found also in the Atlantic. They may perhaps have been mixed up earlier? 

G. pungens Giesbrecht,1895 ( - Pacific) 

N Atlantic 

Skagerrak-N North Sea-Norw. Sea-Iceland-Faeroes-Ireland-NE Atlantic 

Gaetanus Giesbrecht,1888 

E. crassus Giesbrecht,1888 


G. armiger Giesbrecht,1888 

N North Sea-Norw. Sea-Barents Sea-Iceland-Faeroes-Shetlands-NE Atlantic 

E. sewelli Fleminger,1973 

= E. attenuatus : Auctt., non (Dana,1849) 

* According to Fleminger,1973 this is the only N Atlantic species of the attenuatus group. 

G. miles Giesbrecht,1888 () 

N North Sea-Faeroes-Shetlands-Ireland-NE Atlantic 

S & W Ireland, NE Atlantic G. latifrons G.O. Sars,1905 

Rhincalanus Dana,1853 

N North Sea-Ireland-NE Atlantic 

G. minor Farran,1905 

R. nasutus Giesbrecht,1888 

Koster Channel, off Väderöbod, Skagerrak, N North Sea-W Norway-Norw. Sea-?Barents Sea-Iceland- 


Faeroes-Scotland-Ireland-NE Atlantic G. kruppii Giesbrecht,1903 

Iceland-Faeroes-Shetlands-Ireland-NE Atlantic 

R. cornutus (Dana,1849) 

?NE Atlantic 

G. brachyurus G.O. Sars,1907 

Iceland-Faeroes-NE Atlantic 

Clausocalanoidea Giesbrecht,1892 

G. pileatus Farran,1903 

Spinocalanidae Vervoort,1951 

* May be a synonym of G. caudani Canu 


Spinocalanus Giesbrecht,1888 ( S. abyssalis Giesbrecht,1888 - abyssal Pacific) 

G. curvicornis G.O. Sars,1905 

S. longicornis G.O. Sars,1900 

= S. abyssalis : G.O. Sars,1903, non Giesbrecht,1888 

* Atlantic & Arctic deep water 

NE Atlantic 

= S. schaudinni Mrazek,1902 

non = S. longicornis : G.O. Sars,1903 

Central Arctic Ocean 

G. caudani Canu,1896 

Biscay 

S. brevicaudatus Brodsky,1950 

Aetideopsis G.O. Sars,1903 

= S. longicornis : G.O. Sars,1903, non G.O. Sars,1900 

= Faroella Wolfenden,1904 

N North Sea-W Norway-Norw. Sea-Barents Sea-Iceland-Faeroes-Shetlands-Ireland-NE Atlantic 

A. rostrata G.O. Sars,1903 () 

S. magnus Wolfenden,1904 

Norw. Sea-Barents Sea-Iceland-Faeroes-Shetlands-off W Greenland-NE Atlantic 

= S. latifrons G.O. Sars,1907 

Norw. Sea-Barents Sea-Iceland-Ireland-NE Atlantic 

A. multiserrata (Wolfenden,1904) 

= Faroella multiserrata Wolfenden,1904 

S. angusticeps G.O. Sars,1920 

Iceland-Faeroes-Ireland-NE Atlantic-central Norwegian Sea 

* Oceanic deep water species 


Undinopsis G.O. Sars,1884 

= Bradyidius Giesbrecht,1897 

S. spinosus Farran,1908 



U. similis G.O. Sars,1903 

* hyperbenthic 

W Norway-Norw. Sea-NE Atlantic 

Drepanopsis Wolfenden,1911 

= Farrania G.O. Sars,1920 

U. luluae (Grize,1972) 

Bergen (bottom-living) 

D. frigidus Wolfenden,1911 () 

* Oceanic deep water species. As D. oblongus has been considered a synonym of this species, the 

distributions are somewhat mixed up. 

?Biscay, NE Atlantic 

U. bradyi G.O. Sars,1884 () 

= U. armatus (Brady,1878), non (Boeck,1872) 


Oslofjord-N North Sea-Bergen (bottom-living)-Aalesund-Norw. Sea-Barents Sea-W Br. Isles-NE Atlantic 

D. oblongus (G.O. Sars,1920) 

= Farrania oblonga G.O. Sars,1920 ( of Farrania G.O. Sars,1920) 

Aetideus Brady,1883 

NE Atlantic A. armatus (Boeck,1872) 

= Pseudocalanus armatus Boeck,1872 

Mimocalanus Farran,1908 Oslofjord-North Sea-Bergen (bottom-living)-Norw. Sea-?Barents Sea-Iceland-Faeroes-Shetlands-W Br. 

Isles-NE Atlantic 

M. cultrifer Farran,1908 


Pseudaetideus Wolfenden,1903 


P. armatus (Boeck,1872) 

M. nudus Farran,1908 



= Euchaeta armata Boeck,1872 () 

= Chiridius armatus : Auctt. 

Bohuslän-N North Sea-Bergen (bottom-living)-Norw. Sea-Barents Sea-Iceland-Faeroes-Shetlands-W Br. 

Isles-NE Atlantic 

Monacilla G.O. Sars,1905 

Chiridius Giesbrecht,1892 

M. tenera G.O. Sars,1907 



C. obtusifrons G.O. Sars,1903 

= C. armatus G.O. Sars,1900, non Boeck,1872 

N Norw. Sea-Barents Sea-Iceland-Faeroes-NE Atlantic 

M. typica G.O. Sars,1905 


S & W Ireland 

C. tenuispinus G.O. Sars,18?? 

Ofotenfjord, Arctic Seas 

Aetideidae Giesbrecht,1892 

C. poppei Giesbrecht,1892 

S & W Ireland, NE Atlantic, Mediterranean 

Bradyetes Farran,1905 C. gracilis Farran,1908 

B. matthei Johannessen,1976 


Bergen (bottom-living) 

Comantenna Wilson,1924 

B. brevis Farran,1936 

= Bryaxis G.O. Sars,1903, ex Boeck MS, non Kugelann,1794 

* Living close to bottom in moderately deep water 

Barents Sea, E Greenland 

C. brevicornis G.O. Sars,1903, ex Boeck MS 


B. inermis Farran,1905 

N North Sea-Bergen-Barents Sea-Ireland-NE Atlantic 

* Living close to bottom in moderately deep water 


C. minor (Farran,1905) 

= Bryaxis minor Farran,1905 

Chiridiella G.O. Sars,1907 

* hyperbenthic in deep water 


C. macrodactyla G.O. Sars,1907 () 

* Deep water of temterate Atlantic 


Euaetideus G.O. Sars,1925

E. giesbrechti (Cleve,1904) 

Faeroes-Shetlands-Ireland-NE Atlantic 

P. scotti (Farran,1909) 

Iceland-W Ireland-NE Atlantic 

Derjuginia Jaschnov,1947 

P. acuta (Giesbrecht,1892) 

D. tolli (Linko,1913) 

= Scolecithrix tolli Linko,1913 

= Euchaeta acuta Giesbrecht,1892 

N Atlantic, Mediterranean 

All marginal seas of the Polar Basin 

P. bisinuata (G.O. Sars,1907) 

Pseudogaetanus Brodsky,1950 

Biscay, temperate Atlantic, Azores, Canaries 

P. robustus (G.O. Sars,1905) 

off Ireland Azores 

P. bradyi (With,1915) 

= Euchaeta Bradyi With,1915 

Biscay, N & temperate Atlantic 

Euchirellinae Brodsky,1950 P. gracilis (G.O. Sars,1905) 

= Euchaeta quadrata Farran,1909 

Euchirella Giesbrecht,1888 off Ireland, Biscay, N Atlantic 

E. messinensis (Claus,1863) P. hanseni (With,1915) 

= Undina messinensis Claus,1863 = Euchaeta Hanseni With,1915 

?N North Sea-Norw. Sea-Iceland-Faeroes-Ireland-NE Atlantic- Mediterranean N & temperate Atlantic 

E. rostrata (Claus,1863) P. hebes (Giesbrecht,1888) 

= Undina rostrata Claus,1863 = Euchaeta hebes Giesbrecht,1888 

= Euchaeta hessei Brady,1883 N Atlantic, Gibraltar, Mediterranean 

N North Sea-Norw. Sea-Iceland-Faeroes-Shetlands-Ireland-NE Atlantic-Mediterranean 

P. spinosa (Giesbrecht,1892) 

E. bitumida With,1915 = Euchaeta spinosa Giesbrecht,1892 

?N North Sea-Norw. Sea-Iceland-Faeroes-NE Atlantic Temperate Atlantic, Mediterranean 

E. curticauda Giesbrecht,1888 

N North Sea-Iceland-Faeroes-Ireland-Biscay-NE Atlantic 

P. tonsa (Giesbrecht,1895) 

= Euchaeta tonsa Giesbrecht,1895 

E. maxima Wolfenden,1905 

Biscay, N & temperate Atlantic 

Iceland-Faeroes-Ireland-NE Atlantic P. sarsi (Farran,1909) 

E. pulchra (Lubbock,1856) 

Biscay, N Atlantic 

?Iceland-?Faeroes-Azores-NE Atlantic P. pubera (G.O. Sars,1907) 

= Euchaeta pubera G.O. Sars,1907 

E. brevis G.O. Sars,1905 

Biscay-NE Atlantic 

= Euchaeta marina : Wolfenden,1907, non (Prestandrea,1833) 

= Euchaeta Wolfendeni A Scott,1907 

E. intermedia With,1915 

Portugal, temperate Atlantic 

* Deep water species P. rubicunda (Farran,1908) 

NE Atlantic = Euchaeta rubicunda Farran,1908 

off Ireland, temperate Atlantic 

Pseudochirella G.O. Sars,1920 

P. media (Giesbrecht,1888) 

P. spectabilis (G.O. Sars,1900) = Euchaeta media Giesbrecht,1888 

= Undeuchaeta spectabilis G.O. Sars,1900 off Ireland & Portugal, temperate Atlantic 

Central part of Arctic Seas 

P. polaris Brodsky,1950 

P. notacantha (G.O. Sars,1905) Central part of Arctic Ocean 

= Gaidius notacanthus G.O. Sars,1905 

?Iceland-?Faeroes-Ireland-NE Atlantic 

Euchaeta Philippi,1843 

P. pustulifera (G.O. Sars,1905) 

= Undeuchaeta pustulifera G.O. Sars,1905 


E. barbata barbata Brady,1883 

* Should perhaps be moved to Pareuchaeta A. Scott,1909 

SW Norw. Sea, Faeroes, S. Iceland, Rockall, Atlantic W off cerntral Europe 

P. superba (With,1915) E. barbata farrani With,1915 

= Undeuchaeta superba With,1915 * Should perhaps be moved to Pareuchaeta A. Scott,1909 

Iceland-Faeroes-Ireland-NE Atlantic W Norway-off Lofoten-Iceland-W Greenland-Barents Sea-Arctic Seas 

P. dubia (G.O. Sars,1905) E. marina (Prestandrea,1833) 

= Undeuchaeta dubia G.O. Sars,1905 = Cyclops marinus Prestandrea,1833 () 

?Iceland-?Faeroes-NE Atlantic temperate Atlantic, Mediterranean 

P. cryptospina (G.O. Sars,1905) 

= Gaidius cryptospinus G.O. Sars,1905 

Phaennidae G.O. Sars,1902 

* Temperate & N Atlantic deep water species 


Cornucalanus Wolfenden,1905 

P. obtusa (G.O. Sars,1905) 

= Undeuchaeta obtusa G.O. Sars,1905 

* Temperate Atlantic deep water species 


C. chelifer (J.C. Thompson,1903) 

= Scolecithrix chelifer J.C. Thompson,1903 

= Cornucalanus magnus Wolfenden,1905 

off W Ireland, off SW Faeroes, Iceland-Faeroe Channel, off S Iceland, Denmark Strait 

P. divaricata (G.O. Sars,1905) 

Onchocalanus G.O. Sars,1905 

= Gaidius divaricatus G.O. Sars,1905 

Biscay, temperate Atlantic 

O. magnus (Wolfenden,1906) 

= Xanthocalanus magnus Wolfenden,1906 

P. scopularis (G.O. Sars,1905) 

= Undeuchaeta scopularis G.O. Sars,1905 

= Onchocalanus frigidus Wolfenden,1911 

Arctic Seas 

Temperate Atlantic O. cristatus (Wolfenden,1904) 

Chirundina Giesbrecht,1895 

= Xanthocalanus cristatus Wolfenden,1904 

= Onchocalanus trigoniceps G.O. Sars,1905 

C. streetsii Giesbrecht,1895 () 

Iceland-Faeroes-Shetlands-Ireland-Biscay-NE Atlantic 

=? Xanthocalanus similis Esterly,1906 

off W Ireland, off SW Faeroes, Iceland-Faeroe Channel, off S Iceland, Denmark Strait 

Undeuchaeta Giesbrecht,1888 

O. hirtipes G.O. Sars,1905 

off W & NW Ireland 

U. plumosa (Lubbock,1856) 

= Undina plumosa Lubbock,1856 

Norw. Sea-Iceland-Faeroes-Ireland-E Engl. Channel-NE Atlantic 

O. affinis With,1915 

Temperate & boreal Atlantic, Azores 

U. major Giesbrecht,1888 () 

Norw. Sea-Iceland-Faeroes-Ireland-?E Engl. Channel-Biscay-NE Atlantic 

O. trigoniceps G.O. Sars,1905 () 

= Xanthocalanus magnus Wolfenden,1909, non Wolfenden,1906 

Biscay, off Portugal, Mediterranean 

Pseudeuchaeta G.O. Sars,1905 

Oothrix Farran,1905 

P. brevicauda G.O. Sars,1905 

* Temperate Atlantic deep water species 


O. borealis Wiborg,19?? 

W Norwegian fjords 

Valdiviella Steuer,1904 ( V. oligarthra Steuer,1904) 

Pseudophaenna G.O. Sars,1903 

V. insignis Farran,1908 

Gibraltar, Canaries, temperate Atlantic 

P. typica G.O. Sars,1903 

Oslofjord-N Norway 

V. brevicornis G.O. Sars,1905 

temperate deep Atlantic 

Euchaetidae Giesbrecht,1892 

Pareuchaeta A. Scott,1909 

= Euchaeta : Auctt. 

P. norvegica (Boeck,1872) 

= Euchaeta norvegica Boeck,1872 () 

Bohuslän-W Norway-Arctic Seas-Iceland-Scotland 

P. glacialis (Hansen,1886) 

N Norw. Sea-Iceland-Arctic Seas 

Talacalanus Wolfenden,1911 

T. grenii (Farran,1905) 

= Xanthocalanus grenii Farran,1905 

=? X. calaminus Wolfenden,1906 

* Is correct spelling greeni ? 

off S Iceland, off W Ireland, Biscay 

Xanthocalanus Giesbrecht,1892 

X. minor Giesbrecht,1892 

= X. fragilis Aurivillius,1898 

Koster area (between Hällsöarna), S Norway-Bergen (bottom-living)-Trondheim 

6

X. fragilis Aurivillius,1898 

Skagerrak 

X. simplex Aurivillius,1898 

Skagerrak 

X. borealis G.O. Sars,1900 

W Norway-Kara Sea-Greenland Sea-Arctic Seas-Shetlands, Biscay, off Azores 

X. propinquus G.O. Sars,1903 

W-N Norway 

X. fallax G.O. Sars,1919 

Bergen (bottom-living), off W Ireland 

X. pinguis Farran,1905 

off W Norway, off W Ireland 

X. obtusus Farran,1905 

off W Ireland 

X. subagilis Wolfenden,1904 

off W Ireland 

X. muticus G.O. Sars,1905 

Biscay, Azores 

X. agilis Giesbrecht,1892 () 

Mediterranean 

X. polaris Brodsky,1950 

Abyssal Arctic Ocean 

Cephalophanes G.O. Sars,1907 

C. refulgens G.O. Sars,1907 ( - deep Atlantic) 

off W Ireland & Britain - off W Spain & Africa 

Amallophora T. Scott,1894 

A. typica (T. Scott,1892) 

off W Ireland 

Heteremalla G.O. Sars,1907 

H. dubia (T. Scott,1894) 

= Amallophora dubia T. Scott,1894 

Temperate Atlantic, Mediterranean 

Brachycalanus Farran,1905 

B. atlanticus (Wolfenden,1904) 

= Xanthocalanus atlanticus Wolfenden,1904 

off W Ireland 

Oothrix Farran,1905 

O. bidentata Farran,1905 

off W Ireland 

Phaenna Claus,1863 

P. spinifera Claus,1863 

=? P. zetlandica T. Scott,1??? 

off Svinøy (North of Cape Stadt), ?Shetlands, Atlantic, Mediterranean 

Scolecithricidae Giesbrecht,1892 

Scottocalanus G.O. Sars,1905 

S. securifrons (T. Scott,1894) 

= Scolecithrix securifrons T. Scott,1894 

= Scottocalanus acutus G.O. Sars,1905 

Iceland, off W Ireland, Biscay, Canaries, Azores 

S. persecans (Giesbrecht,1892) 

= Scolecithrix persecans Giesbrecht,1892 () 

= Scottocalanus Helenae Scott,1909 

= Scottocalanus Thorii With,1915 

off W Ireland, Biscay, off Gibraltar, Canaries, Azores 

Scolecithricella G.O. Sars,1903 

S. minor (Brady,1883) v. occidentalis Brodsky,1950 

= Scolecithrix minor Brady,1883 () 

S Skagerrak- off Väderöbod-Oslofjord-W Norway-Lofoten-Iceland 

S. ovata (Farran,1905) 

= Scolecithrix ovata Farran,1905 

off W Ireland, off E & W Greenland 

S. auropecten (Giesbrecht,1892) 

= Scolecithrix auropecten Giesbrecht,1892 

off W Ireland, Biscay, Mediterranean 

S. dentata (Giesbrecht,1892) 

= Scolecithrix dentata Giesbrecht,1892 

off W Ireland, Biscay, Mediterranean 

S. vittata (Giesbrecht,1892) 

= Scolecithrx vittata Giesbrecht,1892 

Biscay, Mediterranean 

S. abyssalis (Giesbrecht,1892) 

= Scolecithrix abyssalis Giesbrecht,1892 

off Faeroes, Temperate Atlantic, Mediterranean 

Scaphocalanus G.O. Sars,1900 

= Amallophora G.O. Sars,1903, non Scott,1894 

S. magnus (Scott,1894) 

= Amallophora magna T. Scott,1894 () 

= Scolecithrus cristata Giesbrecht,1895 

= Scaphocalanus acrocephalus G.O. Sars,1900 

= Amallophora magna : G.O. Sars,1903 

W Norway-Arctic Seas-Iceland 

S. brevicornis (G.O. Sars,1900) 

= Scolecithrix brevicornis G.O. Sars,1900 

= Scaphocalanus gracilipes Farran,1908 

N Norw. Sea-Arctic Seas-Shetlands, Biscay 

S. curtus (Farran,1926) 

= Scolecithrix curta Farran,1926 

off W Ireland, Biscay 

S. echinatus (Farran,1905) 


S. robustus (T. Scott,1893) 

= Scolecithrix robusta T. Scott,1893 

off W Ireland 

S. medius (G.O. Sars,1907) 

= Amallophora media G.O. Sars,1907 

Azores, Temperate Atlantic 

S. affinis (G.O. Sars,1905 

= Amallophora affinis G.O. Sars,1905 

Biscay, off Gibraltar, Canaries, Azores 

S. angulifrons G.O. Sars,1920 

off Lisboa 

S. polaris Brodsky,1950 

Central part of abyssal Arctic Ocean 

Scolecithrix Brady,1883 

S. danae (Lubbock,1856) 

= Undina Danae Lubbock,1856 () 

Atlantic, Mediterranean 

S. bradyi Giesbrecht,1888 


Lophothrix Giesbrecht,1895 

L. insignis G.O. Sars,1920 

off NW France 

L. humilifrons G.O. Sars,1905 

temperate Atlantic 

L. frontalis Giesbrecht,1895 () 


L. latipes (T. Scott,1893) 

= Scolecithrix latipes T. Scott,1893 

= Scolecithrix angusta Esterly,1911 

= Scolecithrix acuta Wolfenden,1911 


Amallothrix G.O. Sars,1925 

A. arcuata (G.O. Sars,1920) 

= Scolecithricella arcuata G.O. Sars,1920 

off W Scotland 

A. gracilis (G.O. Sars,1905) 

= Scolecithricella gracilis G.O. Sars,1905 () 

= Scolecithrix globiceps Farran,1909 

off W Ireland, off NW France, off Lisboa, Azores 

A. emarginata (Farran,1905) 

off W Ireland, Biscay, off Gibraltar, Canaries, Azores 

A. valida (Farran,1909) 

= Scolecithrix valida Farran,1909 

off W Ireland 

A. valens (Farran,1926) 

= Scolecithrix valens Farran,1926 


A. laminata (Farran,1926) 

= Scolecithrix laminata Farran,1926 


A. lobata (G.O. Sars,1920) 

= Scolecithricella lobata G.O. Sars,1920 

off NW France, Biscay 

A. falcifer (Farran,1926) 

= Scolecithrix falcifer Farran,1926 

Biscay 

A. propinqua (G.O. Sars,1920) 

= Scolecithricella propinqua G.O. Sars,1920 

off Lisboa 

A. curticauda (A. Scott,1909) 

= Scolecithricella curticauda A. Scott,1909 

off Gibraltar, Canaries 

A. obtusifrons (G.O. Sars,1905) 

= Scolecithricella Tydemani Scott,1909 

Biscay, off Gibraltar, Canaries, Azores 

Diaixidae G.O. Sars,1902 

Diaixis G.O. Sars,1902 

D. hibernica (A. Scott,1896) 

= Scolecithrix hibernica A. Scott,1896 

Oslofjord-W Norway-Trondheim 

D. pygmaea (T. Scott,1899) 

= Scolecithrix pygmaea T. Scott,1899 

Scotland, Ireland, Mediterranean 

Mesaiokeratidae Matthews,1961 

Mesaiokeras Matthews,1961 

M. nanseni Matthews,1961 

Bergen-N Norway (bottom-living) 

Stephidae G.O. Sars,1902 

Stephos T. Scott,1892 

S. lamellatus G.O. Sars,1903 

Christiansund-N Norway 

S. scotti G.O. Sars,1903 

Kieler Bucht-Kattegatt-W Norway-Scotland 

7

S. rustadi Strömgren,1969 

Bergen 

S. minor (T. Scott,1892) 

S Norway-Scotland 

S. fultoni (T. & A. Scott,1898) 

Scotland 

Parastephos G.O. Sars,1903 

P. pallidus G.O. Sars,1903 

S-W Norway-Scotland 

Tharybidae G.O. Sars,1902 

Undinella G.O. Sars,1905 

= Paratharybia Tanaka,1937 

U. oblonga (G.O. Sars,1900) () 

off Greenland, Arctic Seas, off Ireland 

U. simplex (Wolfenden,1906) 

= Xanthocalanus simplex Wolfenden,1906, non Aurivillius,1898 

= Undinella brevipes Farran,1906 


Tharybis G.O. Sars,1903 

T. macrophthalma G.O. Sars,1903 

Oslofjord-W Norway-Trondheim 

Pseudocyclopiidae G.O. Sars,1902 

Pseudocyclopia T. Scott,1892 

P. stephoides J.C. Thompson,1895 

Norway (Christiansund)-Liverpool Bay 

P. crassicornis T. Scott,1892 

S-W Norway-Scotland 

P. giesbrechti Wolfenden,1902 

S-W Norway-Shetlands-Scotland 

P. caudata T. Scott,1894 

Aberdeen-Firth of Forth 

P. minor T. Scott,1892 

Firth of Forth 

Clausocalanidae Giesbrecht,1892 

= Pseudocalanidae G.O. Sars,1903 

Clausocalanus Giesbrecht,1888 

C. arcuicornis (Dana,1849) 

W Norway, Norw. Sea, S & W Ireland, Irish Sea, W Engl. Channel 

C. pergens Farran,1926 

* Probably a form of C. arcuicornis (Dana) 


C. paululus Farran,1926 

* Probably a form of C. arcuicornis (Dana) 


Pseudocalanus Boeck,1872 

P.elongatus (Boeck,1865) 

= Clausia elongata Boeck,1865 

= Calanus clausii Brady,1865 

Bothnian Sea-Kattegatt-Bohuslän-North Sea-N Norway-Arctic Seas-Engl. Channel 

P. minutus (Krøyer,1845) 

= P. gracilis G.O. Sars,1903 

Faeroes-Iceland-N Norway-N Norw. Sea-Arctic Seas 

P. acuspes (Giesbrecht,1881) 

= Lucullus acuspes Giesbrecht,1881 

Baltic, Katt, Bohus-N Norway-Arctic Seas 

P. major G.O. Sars,1900 

Arctic Seas, ?Kara Sea, ?Spitsbergen 

Microcalanus G.O. Sars,1901 

M. pygmaeus (G.O. Sars,1900) 

= Pseudocalanus pygmaeus G.O. Sars,1900 

North Sea-W Norway-Arctic Seas 

M. pusillus G.O. Sars,1903 

* Sometimes considered to be a form of M. pygmaeus (G.O. Sars) 

Kattegatt-North Sea-W Norway 

Drepanopus Brady,1883 

D. bungei G.O. Sars,1898 

Barents Sea, Kara Sea, Laptev Sea 

Ctenocalanus Giesbrecht,1888 

C. vanus Giesbrecht,1888 


Centropagoidea Giesbrecht,1892 

Temoridae Giesbrecht,1892 

Temora Baird,1850 

T. longicornis (O.F. Müller, 1785) 

= Cyclops longicornis O.F. Müller,1785 

Finland-Kattegatt-Bohuslän-North Sea-W & N Norway-Iceland-Engl. Channel-Mediterranean 

T. stylifera (Dana,1849) 

= Calanus stylifer Dana,1849) 

= Temora armata Claus,1863 

Temperate & chilly Atlantic, Mediterranean 

Eurytemora Giesbrecht,1881 

= Temorella Claus,1881 

E. velox (Liljeborg,1853) 

Brackish water-limnic; e.g. Baltic, S Norwegian & Scottish rock-pools 

E. affinis (Poppe,1880) 

= E. hirundo Giesbrecht,1881 () 

= E. hirundoides Nordquist,1888 

* See Buch & Brenning,1992. Crustaceana 62 : 13-38 regarding synonomy. 

Brackish water; river mouths in Finland, E Sweden, Skåne, Isefjord, Oslofjord-N Norway, Zuidersee, 

Germany, Br. Isles 

E. americana L.W. Williams,1906 

=? E. thompsoni Willey,1923 

* Estuarine species living in freshened waters 

?Europe, Woods Hole area 

Temoropia T. Scott,1893 

T. mayumbaensis T. Scott,1893 

off W Ireland, Biscay, temperate & chilly Atlantic 

Centropagidae Giesbrecht,1892 

Limnocalanus G.O. Sars,1863 

L. macrurus G.O. Sars,1863 grimaldi (de Guerne,1886) 

= Centropages grimaldi de Guerne,1886 

* L. macrourus is a freshwater species in Sw., Norw., Finl., etc. and L. grimaldi a brackish water form in 

Baltic, Caspian & Arctic Seas. 

Baltic, E Greenland, Spitsbergen, Kara Sea, Laptev Sea 

Isias Boeck,1865 

I. clavipes Boeck,1865 

Kattegatt-Gullmarfjord-Hafstensund-Oslofjord-W Norway-Trondheim-Br. Isles-Mediterranean 

Centropages Krøyer,1849 

C. typicus Krøyer,1849 () 

Åland-Kattegatt-Bohuslän-Trondheim-North Sea-Iceland-Faeroes-Shetlands-Br. Isles-E Atlantic 

C. hamatus (Liljeborg,1853) 

Bothnian Sea-Kattegatt-Bohuslän-North Sea-W Norway-Barents Sea-Iceland-Faeroes-Shetlands-Br. Isles-E 

Atlantic 

C. bradyi Wheeler,1899 

S & W Ireland, temperate Atlantic, Mediterranean 

C. violaceaus (Claus,1863) 

S & W Ireland 

C. chierchiae Giesbrecht,1889 

Warm water part of NE Atlantic 

C. kroeyeri Giesbrecht,1892 

?NE Atlantic 

Candaciidae Giesbrecht,1892 

Candacia Dana,1846 

C. norvegica (Boeck,1865) 

Oslofjord-Skagerrak-W Norway-N North Sea-Lofoten-Iceland-Faeroes-Shetlands-Ireland 

C. armata (Boeck,1865) 

Oslofjord-Skagerrak-North Sea-W Norway-Faeroes-Shetlands-W Br. Isles-Engl. Channel 

C. elongata (Boeck,1872) 

= C. rotundata (Wolfenden,1904) 

= C. obtusa G.O. Sars,1905 

Norw. Sea-Iceland-Faeroes-Shetlands-Ireland 

C. tenuimana (Giesbrecht,1889) 



C. varicans (Giesbrecht,1892) 



C. longimana (Claus,1863) 


NE Atlantic 

Paracandacia Grice,1963 

P. bispinosa (Claus,1863) 

= Candacia bispinosa Claus,1863 


P. simplex (Giesbrecht,1892) 

= Candacia simplex Giesbrecht,1892 


Pontellidae Dana,1853 

Anomalocera Templeton,1837 

A. patersoni Templeton,1837 

Kattegatt-Bohuslän-North Sea-W Norway-Iceland-Br. Isles- Mediterranean-Black Sea 

Labidocera Lubbock,1853 

L. wollastoni (Lubbock,1857) 

= Pontella Wollastoni Lubbock,1857 

= Pontella helgolandica Claus,1863 

Måseskär, S & E Skagerrak-Oslofjord-Bergen-North Sea-Engl. Channel-Mediterranean 

L. kroeyeri (Brady,1883) 

= Pontella kroeyeri Brady,1883 

off W Ireland 

L. acutifrons (Dana,1849) 

= Pontella acutifrons Dana,1849 

= Pontia Edwardsii Krøyer,1849 


L. brunescens (Czerniavsky,1868) 

= Pontella brunescens Czerniavsky,1868 


L. nerii (Krøyer,1849) 

= Pontia Nerii Krøyer,1849 

off Cape Finisterre 

Pontella Dana,1846 

8

P. atlantica (H. Milne-Edwards,1840) 

= Pontia atlantica H. Milne-Edwards,1840 


P. lobiancoi (Canu,1888) 

= Pontellina Lo Biancoi Canu,1888 


P. mediterranea (Claus,1863) 

= Pontellina mediterranea Claus,1863 

Mediterranean, Black Sea 

Pontellina Dana,1852 

= Calanops Claus,1863 

P. plumata (Dana,1849) 

= Calanops messinensis Claus,1863 

Temperate Atlantic, Azores, Mediterranean 

Pontellopsis Brady,1883 

P. regalis (Dana,1849) 

= Pontella regalis Dana,1849 


P. villosa Brady,1883 


Parapontellidae Giesbrecht,1892 

Parapontella Brady,1878 

P. brevicornis (Lubbock,1857) 

W Norway-Br. Isles-France-Mediterranean 

Acartiidae G.O. Sars,1903 

Acartia Dana,1846 

= Dias Liljeborg,1853 

A. (Acanthacarthia Steuer,1915) bifilosa Giesbrecht,1881 

Bothnian Sea-Bohuslän-North Sea-Iceland-Engl. Channel 

A. (Acanthacartia) tonsa Dana,1848 

Bay of Finland-Belt Sea, S North Sea-E Engl. Channel 

A. (Acartiura Steuer,1915) clausii Giesbrecht,1889 

Belt Sea-Bohuslän-North Sea-W Norway-Iceland-Faeroes-Shetlands-W Br. Isles-W Engl. Channel- 

Mediterran. 

A. (Acartiura) longiremis (Liljeborg,1853) 

Bay of Finland-Bohuslän-Engl. Channel-North Sea-W & N Norway-Barents Sea-Iceland-Faeroes-Shetlands 

A. (Acartiura) discaudata (Giesbrecht,1881) 

Belt Sea-Kattegatt, N North Sea-W Norway-W Br. Isles-Engl. Channel 

A. (Paracartia T. Scott,1894) grani (G.O. Sars,1904) 

= Paracartia grani G.O. Sars,1904 

W Norway 

A. (Paracartia) latisetosa (Kricagin,1873) 

Mediterranean, Black Sea 

A. lefevreae Bradford,1976 

?NE Atlantic 

Diaptomidae Baird,1850 

Pseudodiaptomidae G.O. Sars,1902 

Sulcanidae Nicholls,1945 

Tortanidae G.O. Sars,1902 

Bathypontioidea Brodsky,1950 

Bathypontiidae Brodsky,1950 

Sognocalanus Fosshagen,1967 

S. confertus Fosshagen,1967 

Sognefjord & Hardangerfjord 

Damkaeria Fosshagen,1983 

D. falcifera Fosshagen,1983 () 

Norw. Sea (bottom-living, deep) 

Bathypontia G.O. Sars,1905 

B. elongata G.O. Sars,1905 () 

off W Ireland, Biscay, off Portugal 

Temorites G.O. Sars,1900 

* Previously included in Temoridae 

T. brevis G.O. Sars,1900 () 

= Temoropsis simplex Wolfenden,1911 

Arctic Seas, Mediterranean 

Augaptiloidea G.O. Sars,1905 

Metridinidae G.O. Sars,1902 

Metridia Boeck,1865 

M. lucens Boeck,1865 

= Paracalanus hibernicus Giesbrecht,1892 

Kattegatt-Bohuslän-German Bight-North Sea-Lofoten-Iceland-Faeroes-Shetlands-Br. Isles-Engl. Channel, 


M. longa (Lubbock,1854) 

= Calanus longus Lubbock,1854 

= Metridia armata Boeck,1865 () 

Kattegatt-Bohuslän-North Sea-N Norway-Norw. Sea-Arctic Seas-Iceland-Faeroes-Shetlands-Ireland 

M. princeps Giesbrecht,1892 

Norw. Sea-Iceland-Faeroes-Ireland, Biscay 

M. venusta Giesbrecht,1892 

Iceland-Faeroes-Shetlands-Ireland 

M. brevicauda Giesbrecht,1889 

Biscay, E Atlantic 

M. macrura G.O. Sars,1905 

E Atlantic 

Pleuromamma Giesbrecht,1898 

P. robusta (F. Dahl,1893) 

W Norway-Norw. Sea-Iceland-Faeroes-Shetlands-Ireland-W Engl. Channel 

P. abdominalis (Lubbock,1856) 

= Diaptomus abdominalis Lubbock,1856 () 

W Norway, Iceland-Faeroes-Shetlands-Ireland 

P. xiphias (Giesbrecht,1889) 

= Pleuromma xiphias Giesbrecht,1889 

Iceland-Faeroes-Shetlands-Ireland-W Engl. Channel 

P. borealis (F. Dahl,1893) 

= Pleuromma borealis F. Dahl,1893 

Faeroes-Shetlands-Ireland 

P. gracilis (Claus,1863) 

= Pleuromma gracilis Claus,1863 

S & W Ireland, E Atlantic 

P. piseki Farran,1929 

?NE atlantic 

Heterorhabdidae G.O. Sars,1902 

Heterorhabdus Giesbrecht,1898 

= Heterochaeta Claus,1862, non Westwood,1843 (Orthoptera), nec Claparède,1863 (Oligochaeta) 

H. (Paraheterorhabdus Brodsky,1950) compactus (G.O. Sars,1900) 

= Heterochaeta compacta G.O. Sars,1900 

off Ireland, off W Greenland, Biscay, Arctic Seas-N Atlantic 

H. (Paraheterorhabdus) robustus Farran,1909 

Iceland-Faeroes-Ireland 

H. (Paraheterorhabdus) vipera (Giesbrecht,1889) 

= Heterochaeta vipera Giesbrecht,1889 

E Atlantic 

H. (Euheterorhabdus Brodsky,1950) norvegicus (Boeck,1872) 

= Heterochaeta profunda Dahl,1894 

= Heterochaeta norvegica : G.O. Sars,1900 

Bohuslän(fairly common in deepest Koster Channel)-N North Sea-Lofoten-Iceland-Faeroes-Shetlands- 

Ireland, Greenland, Arctic Seas 

H. (Euheterorhabdus) abyssalis (Giesbrecht,1889) 

N North Sea-Iceland-Faeroes-Shetlands-Ireland 

H. (Euheterorhabdus) spinifrons (Claus,1863) 

= Heterochaeta spinifrons Claus,1863 () 

Norw. Sea, S & W Ireland, E Atlantic, Mediterranean 

H. (Euheterorhabdus) papilliger (Claus,1863) 

= Heterochaeta papilligera Claus,1863 

S & W Ireland, E Atlantic, Mediterranean 

H. (Euheterorhabdus) clausii (Giesbrecht,1889) 

= Heterochaeta clausii Giesbrecht,1889 


Heterostylites G.O. Sars,1920 

H. longicornis (Giesbrecht,1892) 

= Heterochaeta longicornis Giesbrecht,1892 

Iceland-Faeroes-Shetlands-Ireland 

H. major (F. Dahl,1894) 

= Heterochaeta major F. Dahl,1894 

= Alloiorhabdus medius Wolfenden,1911 

Norw. Sea-Iceland-Faeroes-Ireland, Biscay, Azores 

Hemirhabdus Wolfenden,1911 

H. grimaldii (Richard,1893) 



Disseta Giesbrecht,1889 

D. palumboi Giesbrecht,1889 () 


S & W Ireland, Biscay, NE Atlantic 

Mesorhabdus G.O. Sars,1905 

M. angustus G.O. Sars,1907 

?NE Atlantic 

M. brevicaudatus (Wolfenden,1905) 

= Heterorhabdus brevicaudatus Wolfenden,1905 



Augaptilidae G.O. Sars,1905 

Augaptilus Giesbrecht,1892 

A. glacialis G.O. Sars,1900 

= Augaptilis zetesios Wolfenden,1902 

Shetlands-Faeroes-Iceland-Greenland-Spitsbergen-Arctic Seas-NE Atlantic 

A. longicaudatus (Claus,1863) 

= Hemicalanus longicaudatus Claus,1863 () 

off W Ireland, Biscay, temperate & boreal Atlantic, Mediterranean 

A. megalurus Giesbrecht,1892 

off W Ireland, temperate Atlantic 

A. anceps Farran,1909 

off W Ireland, Mediterranean 

A. spinifrons G.O. Sars,1907 

Temperate Atlantic, off Gibraltar 

Augaptilina G.O. Sars,1920 

A. scopifera G.O. Sars,1920 

Biscay 

9

Centraugaptilus G.O. Sars,1920 

C. cucullatus (G.O. Sars,1905) 

off W Ireland, Biscay, Azores, Canaries, temperate Atlantic 

C. horridus (Farran,1908) 


C. rattrayi (T. Scott,1893) () 

off W Ireland, Biscay, Azores, Canaries, temperate Atlantic 

Euaugaptilus G.O. Sars,1920 

E. affinis G.O. Sars,1920 


E. angustus (G.O. Sars,1905) 

= Augaptilus angustus G.O. Sars,1905 

Canaries, Azores, temperate Atlantic, Mediterranean 

E. bullifer (Giesbrecht,1892) 

= Augaptilis bullifer Giesbrecht,1892 

off W Ireland, Biscay, temperate Atlantic, Azores, Canaries 

E. clavatus (G.O. Sars,1907) 

= Augaptilus clavatus G.O. Sars,1927 

off Portugal, Azores, temperate Atlantic 

E. digitatus G.O. Sars,1920 

off W Bretagne 

E. elongatus (G.O. Sars,1905) 

= Augaptilus elongatus G.O. Sars,1905 

non = E. elongatus : Sewell,1947 

off W Ireland, Iceland, Biscay, temperate Atlantic, Azores, Canaries 

E. facilis (Farran,1908) 

= Augaptilis facilis Farran,1908 

off W Ireland, temperate Atlantic, Canaries 

E. filigerus (Claus,1863) 

= Hemicalanus filigerus Claus,1863 

Canaries, Azores, temperate Atlantic, Mediterranean 

E. gibbus (Wolfenden,1904) 

= Augaptilus gibbus Wolfenden,1904 

Biscay, off Gibraltar, Azores, temperate Atlantic 

E. hecticus (Giesbrecht,1889) 

= Augaptilus hecticus Giesbrecht,1889 

Biscay, temperate Atlantic, Mediterranean 

E. humilis Farran,1926 


E. laticeps (G.O. Sars,1905) 

= Augaptilus laticeps G.O. Sars,1905 

off W Ireland, Biscay, temperate Atlantic, Azores, Canaries, Mediterranean 

E. magnus (Wolfenden,1904) 

Engl. Channel, off W Ireland, Biscay, temperate Atlantic 

E. nodifrons (G.O. Sars,1905) 

= Augaptilis nodifrons G.O. Sars,1905 

off W Ireland, Biscay, temperate Atlantic, Azores, Canaries 

E. oblongus (G.O. Sars,1905) 

= Augaptilus oblongus G.O. Sars,1905 

Biscay, temperate Atlantic, Azores, Canaries 

E. palumboi (Giesbrecht,1892) 

= Augaptilus Palumbii Giesbrecht,1892 


E. penicillatus G.O. Sars,1920 

off Gibraltar, temperate Atlantic 

E. similis (Farran,1908) 


E. squamatus (Giesbrecht,1889) 

= Augaptilus squamatus Giesbrecht,1889 

= Augaptilus brevicaudatus G.O. Sars,1905 

off W Ireland, off Portugal, temperate Atlantic, Azores, Canaries 

E. truncatus (G.O. Sars,1905) 

= Augaptilus truncatus G.O. Sars,1905 

off W Ireland, temperate Atlantic, Azores, Canaries 

E. hyperboreus Brodsky,1950 

Abyssal central Arctic Ocean 

Haloptilus Giesbrecht,1898 

H. longicornis (Claus,1863) 

= Hemicalanus longicornis Claus,1863 () 

Between Finnmark & Bear Island-Bay of Biscay-Mediterranean 

H. acutifrons (Giesbrecht,1892) 

= Hemicalanus acutifrons Giesbrecht,1892 

= Haloptilus spinifrons G.O. Sars,1900 

? North Sea-Between Jan Mayen & Norway-Iceland-Biscay-Mediterranean, Greenland 

H. fons Farran,1908 


H. tenuis Farran,1908 

off W Ireland, temperate Atlantic, Mediterranean 

H. angusticeps G.O. Sars,1907 


H. bulliceps Farran,1905 


H. plumosus (Claus,1863) 

= Hemicalanus plumosus Claus,1863 


H. mucronatus (Claus,1863) 

= Hemicalanus mucronatus Claus,1863 


H. ornatus (Giesbrecht,1892) 

= Hemicalanus ornatus Giesbrecht,1892 


H. spiniceps (Giesbrecht,1892) 

= Hemicalanus spiniceps Giesbrecht,1892 


H. validus G.O. Sars,1920 


Heteroptilus G.O. Sars,1920 

H. acutilobus (G.O. Sars,1905) 

= Pontoptilus acutilobus G.O. Sars,1905 

Biscay, Canaries, Azores, temperate Atlantic 

H. attenuatus (G.O. Sars,1905) 

= Pontoptilus attenuatus G.O. Sars,1905 


Pontoptilus G.O. Sars,1905 

P. muticus G.O. Sars,1905 


Pachyptilus G.O. Sars,1920 

P. abbreviatus (G.O. Sars,1905) 

= Pontoptilus abbreviatus G.O. Sars,1905 


P. erygnathus G.O. Sars,1920 

off Lisboa, Azores, temperate Atlantic 

Pseudaugaptilus G.O. Sars,1907 ( P. longiremis G.O. Sars,1907 - off Azores) 

P. polaris Brodsky,1950 

Abyssal central Arctic Ocean 

Discoidae Gordejeva,1975 

Disco Grice & Hulsemann,1965 

D. fiordicus Fosshagen,1967 

Hardangerfjord 

Arietellidae G.O. Sars,1902 

Arietellinae G.O. Sars,1902 

Paramisophria T. Scott,1897 

P. cluthae T. Scott,1897 

W Norway-Scotland-Mediterranean 

P. spooneri Krishnaswamy,1959 

SW Br. Isles (shell gravel living) 

Scottula G.O. Sars,1903 

S. inaequicornis G.O. Sars,1903 

Oslofjord-W Norway 

S. abyssalis G.O. Sars,1905 

off Lisboa, Azores, temperate Atlantic 

Arietellus Giesbrecht,1892 

A. pavoninus G.O. Sars,1905 

off W Ireland, Azores 

A. plumifer G.O. Sars,1905 

off W Ireland, off Portugal, Canaries, Azores 

A. simplex G.O. Sars,1905 

off W Iceland, W Greenland, off W Ireland, Azores 

A. giesbrechti G.O. Sars,1905 

off Lisboa & Gibraltar, Canaries, temperate Atlantic 

A. setosus Giesbrecht,1892 () 

NE? Atlantic, Mediterranean 

Paraugaptilus Wolfenden,1904 

P. buchani Wolfenden,1904 () 

off W Ireland, Canaries, Azores, temperate Atlantic 

Epacteriscidae Fosshagen,1973 

Lucicutiidae G.O. Sars,1902 

Lucicutia Giesbrecht, in Giesbrecht & Schmeil,1898 (n. cons., Op. 1613, ICZN) 

= Leuckartia Claus,1862, non Agassiz,1862 (Coelenterata) 

= Rudolfia Wilson,1924 

L. atlantica Wolfenden,1904 

= Lucicutia magna Wolfenden,1920 

= Lucicutia gracilis G.O. Sars,1905 

Iceland, off W Ireland, Biscay, Azores, temperate Atlantic, Biscay 

L. curta Farran,1905 

off W Ireland, Biscay, northern & temperate Atlantic 

L. anomala Brodsky,1950 

Arctic Seas 

L. polaris Brodsky,1950 

Arctic Seas 

L. pseudopolaris Heptner,1969 

Arctic Seas 

L. grandis (Giesbrecht,1895) 

= Leuckartia grandis Giesbrecht,1895 

Engl. Channel, W Ireland, N & temperate Atlantic 

L. lucida Farran,1908 

= Lucicutia pera A. Scott,1909 

off W Ireland, temperate Atlantic, Mediterranean 

10

L. macrocera G.O. Sars,1920 

off W Bretagne, temperate Atlantic 

L. maxima (Steuer,1904) 

= Leuckartia maxima Steuer,1904 

off W Bretagne, off Lisboa, temperate Atlantic 

L. simulans G.O. Sars,1920 


L. longiserrata (Giesbrecht,1892) 

= Leuckartia longiserrata Giesbrecht,1892 

Gibraltar, Canaries, temperate Atlantic, Mediterranean 

L. clausi (Giesbrecht,1889) 

= Leuckartia clausi Giesbrecht,1889 


L. flavicornis (Claus,1863) 

= Leuckartia flavicornis Claus,1863 () 


L. gemina Farran,1926 


L. longicornis (Giesbrecht,1889) 

= Leuckartia longicornis Giesbrecht,1889 

Biscay 

L. ovalis Wolfenden,1911 

Biscay, temperate & tropical Atlantic 

L. tenuicauda G.O. Sars,1907 

off Lisboa, Canaries, temperate Atlantic 

L. polaris Brodsky,1950 

Central part of Arctic Ocean 

L. anomala Brodsky,1950 

Central part of Arctic Ocean 

Isochaeta Giesbrecht,1898 

I. ovalis Giesbrecht,1898 () 


Phyllopodidae Brodsky,1950 

Phyllopus Brady,1883 

P. bidentatus Brady,1883 () 

off Ireland 

P. helgae Farran,1908 

off Ireland & Gibraltar, Biscay, Azores, temperate Atlantic 

P. impar Farran,1908 

off Ireland & Gibraltar, Azores, temperate Atlantic 

P. aequalis G.O. Sars,1920 

off Gibraltar, temperate Atlantic 

Pseudocyclopoidea Giesbrecht,1893 

Pseudocyclopidae Giesbrecht,1893 

Pseudocyclops Brady,1876 

P. crassiremis Brady,1873 

S Norway (Korshavn)-Br. Isles 

P. obtusatus Brady & Robertson,1873 

Christiansund-Scotland-Arctic Seas 

Ridgewayiidae M.S. Wilson,1958 

Ridgewayia Thompson & Scott,1903 

Exumella Fosshagen,1970 

Brattstromia Fosshagen, in Fosshagen & Iliffe,1991 

Placocalanus Ohtsuka, Fosshagen & Soh,1996 ( P. insularis Fosshagen,1970 - Bahamas) 

= Placocalanus Fosshagen,1970 (nom. nud.) 

Ryocalanoidea Andronov,1974 

Ryocalanidae Andronov,1974 

* No species within the area? 

PODOPLEA Giesbrecht,1882 

CYCLOPOIDA Burmeister,1834 

GNATHOSTOMATINA Thorell,1860 

Oithonidae Dana,1853 

Paraoithona Farran,1908 

P. parvula Farran,1908 

W Norway-W Ireland 

Oithona Baird,1843 

* The nomenclature within this genus should be checked more. 

O. atlantica Farran,1908 

= O. spinirostris : G.O. Sars,1918 

=? O. spinirostris Claus,1863 

=?? O. plumifera Baird,1843 

* Should probably be named O. spinirostris Claus,1863, while Baird's species may be another. 

Kattegatt-Skagerrak-Trondheim-Norw. Sea-Iceland-Br. Isles-Mediterranean 

O. minuta Kritchaguine,1873 

= O. nana Giesbrecht,1892 

Kattegatt-Skagerrak-S North Sea 

O. helgolandica Claus,1863 

= O. similis Claus,1863 

* Has been considered a synonym of both O. nana & O. similis 

Åland-Isefjord-Bohuslän, Oslofjord-N Norway-Iceland-Arctic Seas-S North Sea-Yorkshire-Engl. Channel- 

Mediterranean, Atlantic Canada 

O. linearis Giesbrecht,1891 

Temperate Atlantic, Adriatic 

O. setigera (Dana,1849) 

= Oithona tropica Wolfenden,1905 

= Oithona pelagica Farran,1908 

Biscay, Cape Verde, tropical & temperate Atlantic, Mediterranean 

O. robusta Giesbrecht,1891 

Cape Verde 

Cyclopinidae G.O. Sars,1913 

Pterinopsyllinae Kiefer,1927-28 

Pterinopsyllus Brady,1880 

P. insignis (Brady,1878) 

Oslofjord-Skagerrak-Norw. Sea-Trondheim-Br. Isles-Mediterranean 

Cyclopininae G.O. Sars,1913 

Cyclopetta G.O. Sars,1918 

C. difficilis G.O. Sars,1918 

S Norway (Risør)-North Sea 

Cyclopinula Strand,1929 

= Cyclopina Claus,1862, non Agassiz,1846 

C. schneideri (T. Scott,1903) 

= C. brevifurca (G.O. Sars,1918) 

Bohuslän-N Norway-Spitsbergen-Arctic Seas-Canada-?Br. Isles 

C. pygmaea (G.O. Sars,1918) 

= Cyclopina pygmaea G.O. Sars,1918 

North Sea-S & SW Norway-Br. Isles-Mediterranean 

C. brachystylis (G.O. Sars,1921) 

= Cyclopina brachystylis G.O. Sars,1921 

Norway-SE North Sea 

C. norvegica (Boeck,1865) 

= Cyclopina norvegica Boeck,1865 

* Sensu Sars, G.O. 1918 is this a synonyme of C. gracilis (Claus) 

S Baltic-W Sweden-North Sea-N Norway-Arctic Seas-Br. Isles-?Mediterranean 

C. gracilis (Claus,1862) 

= Cyclopina gracilis Claus,1862 

S Baltic-W Sweden-North Sea-Norway-Br. Isles-Mediterranean-Black Sea 

C. kieferi (Schäfer,1936) 

SE North Sea, Mediterranean 

C. latipes (Schäfer,1936) 

SE North Sea 

C. rotundipes (Herbst,1952) 

= Cyclopina rotundipes Herbst,1952 

Sylt 

?C. phallusiae (Hansen,1923) 

= Cyclopina phallusiae Hansen,1925 

@ Ascidia obliqua 

Faeroes 

C. esilis (Brian,1958) 

= Cyclopina esilis Brian,1958 

?Britain 

C. tuberculata (Herbst,1962) 

= Cyclopina tuberculata Herbst,1962 

?North Sea area 

Cyclopidina Steuer,1940 

C. euacantha (G.O. Sars,1918) 

Norway 

Cyclopinella G.O. Sars,1918 

C. tumidula G.O. Sars,1918 

S Norway (Risør & Lillesand)-North Sea-Trondheim-Norw. Sea 

Cyclopinodes Wilson,1932 

C. elegans (Scott,1894) 

Bohuslän-SW Norway (Farsund)-North Sea-Br. Isles-Italy 

Cyclopinoides Lindberg,1953 

C. longicornis (Boeck,1872) 

Skagerrak-S North Sea-N Norway-Arctic Seas-Br. Isles-Mediterranean 

C. dilatata (G.O. Sars,1921) 

North Sea-S Norway (Korshavn) 

C. littoralis (Brady,1872) 

Bohuslän-North Sea-W Norway-Br. Isles-France-Italy 

Hemicyclopina Herbst,1953 

H. setifera (Herbst,1952) 

Sylt 

Metacyclopina Lindberg,1953 

M. harpacticoidea (Klie,1949) 

Helgoland 

M. brevisetosa Herbst,1974 

Sylt 

M. roscoffensis Bozic,1953 

Roscoff 

Microcyclopina Plesa,1961 

Herbstina Huys & Boxshall,1990 

H. exigua (Herbst,1974) 

= Microcyclopina exigua (Herbst,1974) () 

Sylt 

Neocyclopina Herbst,1952 

11

N. reducta Herbst,1952 

Sylt, Amrum 

Pseudocyclopinodes Lang,1946 

P. (Parapseudocyclopinodes Lindberg,1961) neglecta (Herbst,1952) 

Sylt 

P. (Parapseudocyclopinodes Lindberg,1961) dacunhai Lindberg,1961 

W Portugal 

Smirnovipina Arbizu,1997 

S. barentsiana (Smirnov,1931) Arbizu,1997 

= Cyclopina barentsiana Smirnov,1931 () 

= Cyclopinodes barentsiana (Smirnov,1931) Smirnov,1935 

off NE Spitsbergen in 197 m 

Erythropolites Huys & Boxshall,1990 ( Microcyclopina semireducta Herbst,1964) 

Glareolina Huys & Boxshall,1990 ( Hemicyclopina kliei Herbst,1953) 

Cyclopidae Dana,1853 

Halicyclopinae Kiefer,1927 

Euryte Philippi,1843 

E. longicauda Philippi,1843 

Bohuslän-Oslofjord-N Norway-Faeroes-Br. Isles-Mediterranean-Black Sea 

E. minor (T. Scott,1905) 

* possibly a subspecies of E. longicauda Philippi 

Oslofjord-North Sea-W Norway-Norw. Sea-Scotland-Plymouth 

E. robusta Giesbrecht,1901? 

S Norway (Korshavn & Kleven)-Raunefjorden-Christiansund, Mediterranean 

E. curticornis G.O. Sars,1918 

Lofoten, Arctic Seas 

Halicyclops Norman,1903 

H. magniceps (Liljeborg,1853) 

= H. christianensis (Boeck,18??) 

= H. aequoreus (Fischer,1860) 

S Finland-Öresund-S & W Norway, Br. Isles-France-Mediterranean 

H. neglectus Kiefer,1936? 

= H. aequoreus : (Brady,1878), non (Fischer,1860)) 

Baltic-North Sea 

H. neglectus septentrionalis ?Kiefer,1936? 

= H. thermophilus septentrionalis Kiefer,1936? 

Baltic-North Sea 

H. brevispinosus brevispinosus Herbst,1952 

Sylt 

H. incognitus Herbst,1962 

?North Sea area 

Neocyclops Gurney,1927 

N. remanei (Herbst,1952) 

Sylt 

Diacyclops Kiefer,1927 

D. nanus (G.O. Sars,1862) 

?NE Atlantic 

Cyclopinae Dana,1853 

* Freshwater species 

Megacyclops Kiefer,1927 

M. viridis (Jurine,1820) 

= Cyclops vulgaris Koch,1835 

Fresh water-Bothnian Sea 

Mesocyclops G.O. Sars,1918 

M. leuckarti (Claus,1862?) 

= Mesocyclops obsoletus G.O. Sars1918, ?non (Koch,1835) 


Thermocyclops Kiefer,1927 

T. oithonoides (G.O. Sars,1862) 


NOTODELPHYOINA 

Chordeumiidae Boxshall,1988 

Chordeumium Stephensen,1918 

= Chordeuma Jungersen,1912, non Koch,1847 

C. obesum (Jungersen,1912) 

@ In gonad galls of Asteronyx loveni 

Skagerrak 

Parachordeumium le Calvez,1938 

= Ophioithys Heegard,1951 

= Amphiurophilus Delamere Deboutteville,1962 

P. tetraceros le Calvez,1938 

@ genital cavity & coelom of Axiognathus squamatus 

* Not mentioned by Gotto,1993 

Roscoff-French Mediterranean 

P. amphiurae (Hérouard,1906) 

= Philichthys amphiurae Hérouard,1906 

= Ophioithys amphiurae (Hérouard,1906) 

= Amphiurophilus amphiurae : Aucct. 

@ bursae (genital sacs) of Axiognathus squamatus 

Bohuslän, Bergen, Engl. Channel, Isle of Man, Roscoff, ?E USA 

P. bocqueti Goudey-Perrière,1979 

@ bursae of Axiognathus squamatus 


NE Atlantic 

Ophioika Stephensen,1934 

O. appendiculata Stephensen,1935 

@ in bursae of Ophiomitrella clavigera & Ophiacantha bidentata 


W Hebrides-S Iceland-S Greenland 

Arthrochordeumium Stephensen,1918 

* W Pacific 

Ophioicodes Heegard,1951 

Lernaeosaccus Heegard,1951 ( L. ophiacanthae Heegard,1951) 

Thespesiopsyllidae Wilson,1924 

= Thaumatopsyllidae G.O. Sars,1913 

* Probably more related to Cancerillidae than close to Monstrillidae, where they formerly were put, sensu 

Fosshagen,1970, but Huys & Boxshall 1991 removed the family to the Cyclopoida. McKinnon,1994 is of the 

opinion that Wilson,1924 erroneously changed the genus name of our genus. 

Thespesiopsyllus Wilson,1924 

= Thaumatopsyllus G.O. Sars,1913, non T. Scott,1894 

T. paradoxus (G.O. Sars,1913) () 

@ nauplies in stomach folds of Ophiothrix fragilis, Ophiopholis aculeata & Ophiura albida 

* ≈50% of Gullmarfjord Ophiopholis population infested, according to Bresciani & Lützen,1962 

Bohuslän-Oslofjord-Trondheim 

Cucumaricolidae Bouligand & Delamare-Deboutteville,1959 

Cucumaricola Paterson,1958 

Notodelphyoidea Dana,1853 

Notodelphyidae Dana,1853 

@ Ascidians, Octocorals 

* Some genera should perhaps go to a family of their own, Doropygidae Brady,1878 

Notodelphys Allman,1847 

N. allmanni Thorell,1859 

= N. ascidicola Allman,1847 (p.p.) () 

@ Ascidia spp., Ascidiella spp., Phallusia spp. & Ciona spp. 

Isefjord, Bohuslän-N Norway-Br. Isles-Mediterranean-Black Sea-S Africa 

N. rufescens Thorell,1859 

@ Ascidia conchilega, A. obliqua, Ascidiella aspersa & A. scabra 

Bohuslän-S Norway-North Sea-Britain-N Ireland-Faeroes-Mediterranean 

N. caerulea Thorell,1859 

@ Ascidia virginea, ?A. mentula, Ciona intestinalis & Corella parallelogramma 

Isefjord, Bohuslän-S Norway-Shetlands-Br. Isles 

N. agilis Thorell,1859 

@ Ascidia spp., Ascidiella spp., Molgula spp., Polycarpa spp, Ciona intestinalis & Corella 

parallelogramma 

Kattegatt-Bohuslän-N Norway-Br. Isles-Mediterranean 

N. tenera Thorell,1859 

@ Ascidia mentula, A. obliqua, Ciona intestinalis & Phallusia mammilata 

Kattegatt-Bohuslän-NW Norway-Mediterranean 

N. elegans Thorell,1859 

@ Ciona intestinalis & Ascidiella aspersa 

Isefjord, Bohuslän-S Norway-Br. Isles-Mediterranean 

N. prasina Thorell,1859 

= N. pusilla Buchholz,1869 

@ Ascidia mentula, Ascidiella aspersa, Phallusia mammillata & Ciona intestinalis 

Bohuslän-S Norway-Trondheim-Shetlands-Br. Isles-Mediterranean 

N. dentata Schellenberg,1921 

@ Rhopalaea nordgaardi Hartmr. 

Norway 

N. canui Roland,1962 

@ Molgula echinosiphonica 

Engl. Channel 

N. cryptopyge Bocquet & Stock,1960 

@ Pyura savignyi 

Engl. Channel 

N.transatlantica Bocquet & Stock,1960 

@ Clavelina lepadiformis 

N Ireland, Engl. Channel 

N. platymera Illg & Dudley,1961 

@ ?Ascidia mentula 

Engl. Channel 

Agnathaner Canu,1891 

A. typicus Canu,1891 () 

@ Dendrodoa grossularia & Polycarpa rustica 

S Norway (Grimstad)-Br. Isles-Roscoff 

A. freemani Hamond,1968 

@ ? 

Norfolk 

A. minutus Canu,1891 

@ Sidnyum turbinatum 

N France 

Doropygus Thorell,1859 

* Ca 20 indeterminable species, especially erected by Hesse from the coast of France, exist. 

D. pulex Thorell,1859 () 

= D. pullus Buchholz,1869 

@ Ascidiella scabra & many other ascidians & Antedon petasus 

Bohuslän-Norway-Faeroes-Br. Isles-Mediterranean, NE America 

Dorypygella G.O. Sars,1921 

D. psyllus (Thorell,1859) 

@ Ascidia spp., Ascidiella spp. & Phallusia fumigata 

Bohusl-W Norway-Br. Isles-N France-Mediterranean 

D. porcicauda (Brady,1878) 

@ Corella parallelogramma & Ascidia conchilega 

SE Norway (Hvalør)-W Norway-Clyde Sea-Ireland, Biscay, Mediterranean 

D. thorelli (Aurivillius,1882) 

12

= Doropygus thorelli Aurivillius () 

@ Ascidia mentula & A. obliqua 

Bohuslän-S & W Norway-Faeroes-Iceland, Greenland, Davis Strait 

D. normani (Brady,1878) 

@ Styela rustica, Cynthia papillosa (Linnaeus), Cynthia sp. & ?Ascidia virginea 

SW Scotland, NE & W Ireland, S England, Mediterranean 

Doropygopsis G.O. Sars,1921 

D. longicauda (Aurivillius,1882) 

= Doropygus longicauda Aurivillius,1882 () 

@ Ascidia mentula, A. obliqua , A. prunum , A. callosa & Boltenia echinata 

Bohuslän-S & W Norway-Spitsbergen-Kara Sea-Iceland-Faeroes-Greenland-Davis Strait-Maine 

Pachypygus G.O. Sars,1921 

P. gibber (Thorell,1859) 

= Doropygus gibber Thorell,1859 () 

@ Ascidia spp., Ascidiella spp., Molgula spp., Polycarpa spp., Ciona intestinalis, Corella 

parallelogramma, Clavelina lepadiformis etc. 

Bohuslän-S & W Norway-Br. Isles-Mediterranean 

Notopterophorus Leuckart,1859 

= Notopterophorus Costa,1840 (nom. nud.) 

N. auritus (Thorell,1859) 

@ Ascidia spp. & Ciona intestinalis 

Bohuslän-Oslofjord-Trondheim-Shetlands-Br. Isles-Ireland 

N. papilio Hesse,1864 

@ Ascidia spp. & Ciona intestinalis 

Bohuslän-W Norway-Shetlands-Br. Isles-Mediterranean 

N. micropterus G.O. Sars,1921 

@ Ascidia mentula 

* Not mentioned in Gotto,1993. Could it be a synonym of N. elatus Giesbrecht? 

W Norway 

N. elongatus Buchholz,1869 

= N. elongatus Costa,1840 (nom. nud.) 

= N. elatus Costa,1840 (nom. nud.) 

= N. veranyi Leuckart,1859 () 

@ Ascidia mentula, A. gelatinosa, Ascidiella aspersa, Phallusia mammillata & Clavelina lepadiformis 

* Probably confined to Phallusia mamillata according to Gotto,1993 

?W Sweden-?Britain-Mediterranean 

N. elatus Giesbrecht,1892 

@ Ascidia mentula 

North Sea, Engl. Channel, Mediterranean 

N. sp. Schimkewitsch,1896 

@ Ascidia sanguinolenta & Ciona intestinalis 

Roscoff 

N. bombyx Hesse,1865 

* indeterminable species. 

@ Ciona intestinalis 

Coast of France 

Gunenotophorus Buchholz,1869 (Spelling emended) 

= Gunenotophorus O.G. Costa,1840 (nom. nud.) 

= Sphaeronotus Claus,1864, non Laporte,1832 

= Gunentophorus Buchholz,1869 & : Auctt. 

G. globularis Buchholz,1869 () 

= G. globularis O.G. Costa,1840 (nom. nud.) 

= Sphaeronotus thorellii Claus,1864 

* Should not the correct name be G. thorellii (Claus,1864)? 

@ Ascidia spp., Styela spp., Polycarpa spp., Ciona intestinalis, Molgula manhattensis, Dendrodoa 

grossularia etc. 

Bohuslän-Trondheim-Britain-France-Mediterranean, S Africa, Florida 

Botachus Thorell,1859 

B. cylindratus Thorell,1859 () 

= B. fusiformis Buchholz,1869 

@ Ascidia spp., Ascidiella scabra & Ciona intestinalis 

Kattegatt-Bohuslän-Norway-Shetlands-Br. Isles-Mediterranean 

B. fulvus Hesse,1869 

* indeterminable species 

@ Ascidia canina 


B. macroone Hesse,1869 

* indeterminable species 

@ Ascidia canina 


Ophioseides Gerstaecker,1871 

O. cardiacephalus Gerstaecker,1871 () 

=? O. joubini Chatton,1909 

@ Distomus variolosus, Stolonica socialis, Polycarpa pomaria, Styela sp. & Microcosmus sulcatus 

Engl Channel, European Atlantic 

Ophiodelphys Bocquet & Stock,1961 

O. illgi Bocquet & Stock,1961 () 

@ an asacidian of the family Polyclinidae 

Engl. Channel 

O. apoda (Giard,1873) 

= Ophioseides apoda Giard,1873 

@ Astellium perspicuum 

Roscoff 

Ooneides Chatton & Brément,1915 

O. amela Chatton & Brément,1915 

@ Didemnum spp. 

W Scotland, Mediterranean 

Brementia Chatton & Brément,1915 

B. balneolensis Chatton & Brément,1915 

@ Didemnum commune, Polysyncraton bilobatum & Polysyncraton canatensis 

European Atlantic-Mediterranean 

Scolecimorpha G.O. Sars,1926 

S. joubini (Chatton,1909) 

= Ophioseides joubini Chatton,1909 

@ Styela rustica, Polycarpa pomaria, Microcosmus sulcatus (=M. sabatieri ), Pyura trita Sluiter & 

Cnemidocarpa cerea Sluiter 

Bohuslän-?W Norway-French Mediterranean 

S. insignis G.O. Sars,1926 () 

@ Polycarpa pomaria 

Trondheim Fjord 

Doroixys Kerschner,1879 

D. uncinata Kerschner,1879 () 

@ Aplidium spp., Sidnyum spp., Polyclinum aurantium, Morchellium argus, Diazona violacea, Botrylloides 

spp. etc. 

Br. Isles-NE Ireland-N France-Mediterranean 

Bonnierilla Canu,1891 

= Paryphes Kerschner,1879, non Burmeister,1835 

B. alterata Stock,1967 

=B. longipes : (Kerschner,1879) Canu,1891, non Paryphes longipes Kerschner,1879 

@ Clavelina lepadiformis , Polycarpa rustica & Pyura lurida 

W Ireland-N & W France-Mediterranean 

B. filipes Stock,1967 

= B. armata : Illg & Dudley,1961, non Schellenberg,1922 

@ Dendrodoa grossularia, Polycarpa rustica & Ciona roulei 

W Ireland, Mediterranean 

B. similis Illg & Dudley,1961 

@ Pyura squamulosa 

NW Ireland, Mediterranean 

Archinotodelphyidae Lang,1949 

* No taxa in N Europe? 

@ Ascidians 

Mantridae Leigh-Sharpe,1934 

Mantra Leigh-Sharpe,1934 ( M. speciosa Leigh-Sharpe,1934) 

Ascidicoloidea Thorell,1859 

Ascidicolidae Thorell,1859 

@ Ascidians 

Ascidicolinae Thorell,1859 

@ Ascidians 

Ascidicola Thorell,1859 

A. rosea Thorell,1859 () 

@ Ascidia spp., Ascidiella spp., Corella parallelogramma, Ciona intestinalis, Pyura squamulosa , Polycarpa 

pomaria & Halocynthia papillosa 

Isefjord, Bohuslän-S Norway-North Sea-Br. Isles-Mediterranean 

Styelicola Lützen,1968 

S. bahusia Lützen,1968 () 

@ Styela atlantica (Van Name), S. gelatinosa Traustedt & Ciona gelatinosa 

* So far found only below 80 m. 

Koster Channel, Biscay 

Buprorinae Thorell,1859 

@ Ascidians 

Buprorus Thorell,1859 

B. loveni Thorell,1859 () 

@ Ascidia mentula, A. obliqua & Ascidiella aspersa 

Bohuslän-N Norway-Faeroes 

B. nordgaardi G.O. Sars,1921 

@ Amaroucium sp. 

* Not mentioned in Gotto,1993 

Trondheimsfjord 

Enterocolinae Della Valle,1883 

Lequerrea Chatton & Harant,1924 

L. perezi Chatton & Harant,1924 () 

@ intestine of Polycarpa gracilis Heller 

Plymouth, Brest 

L. canui Illg & Dudley,1980 

@ gut of Polycarpa rustica (Linnaeus) 

Plymouth 

Enterocola van Beneden,1860 

= Biocryptus Hesse,1865 

E. bilamellata G.O. Sars,1921 

@ not known (found free) 


SW Norway (Farsund) 

E. pterophora Chatton & Brément,1909 

@ Didemnum fulgens, Trididemnum tenerum & various other colonial ascidians 

NE Ireland, S England, Mediterranean 

E. fulgens van Beneden,1860 () 

= E. betencourti Canu,1891 

@ Aplidium spp., Polyclinum aurantium, Synoicum pulmonaria, Sidnyum turbinatum & ?Botryllus 

schlosseri 

Belgium-Br. Isles-NE Ireland-N France-Mediterranean-S Africa 

E. megalova Gotto,1962 

@ Polyclinum aurantium Milne-Edwards 

NE Ireland 

E. clavelinae Chatton & Harant,1924 

@ Polycitor (Clavelina ) nana Lahille 

N France, Mediterranean 

E. hessei Chatton & Harant,1924 

@ Unidentified compound ascidian 

N France 

E. sydnii Chatton & Harant,1924 

@ Sidnyum turbinatum , S. elegans & Morchellium argus 

N Ireland, S England, France (Cotes du Nord) 

E. setifera Hansen,1923 

13

@ unknown 


off SW Iceland 

E. brementi Illg & Dudley,1980 

@ Aplidium pallidum (Verrill) 

N France 

E. africanus López-González, Conradi & García-Gómez,1993 

@ Synoicum sp. 

Gibraltar area 

E. sp. B Chatton & Brément,1909 

@ unidentified ascidian 

Scotland 

Enteropsinae Aurivillius,1885 

@ Ascidians 

Mychophilus Hesse,1865 

= Mycophilus : Auctt. 

= Mychephilus : Canu,1892 

= Michophilus : Chatton & Harant,1922 

M. roseus Hesse,1865 () 

= M. pachygaster Hesse,1865 

= M. curvatus Chatton & Brément,1909 

= M. rosovula Gray,1933 

= Enteropsis vararensis T. Scott,1901 

@ Botryllus schlosseri (Pallas) & Botrylloides leachii (Savigny) 

Bohuslän-W Norway-Br. Isles-Ireland-W France-Mediterranean 

Enteropsis Aurivillius,1885 

E. sphinx Aurivillius,1885 () 

= Haligryps aculeatus Aurivillius,1885 

@ Molgula manhattensis (De Kay) & Dendrodoa grossularia (van Beneden) 


Siberian Arctic, Alaska 

E. roscoffensis Chatton & Brément,1909 

@ Dendrodoa grossularia (van Beneden), Pyura microcosmus (Savigny) & Tethyum savignyi 

W Ireland,Plymouth, N France-Mediterranean, NW USA 

E. chattoni Monniot,1961 

@ Microcosmus vulgaris Heller, Halocynthia papillata (Linnaeus) & Diazona violacea Savigny 

S England, Mediterranean 

E. pilosus Canu,1886 

@ Diazona violacea 

* Excluded from the genus in the revision by Illg & Dudley,1980. Not mentioned by Gotto,1993. 

W France 

Haplostominae Chatton & Harant,1924 

Haplostoma (Canu,1886) 

= Aplostoma Canu,1886, non Mouquin-Tandon,1856 

= Cryptopodus Hesse,1865 

= Tranestoma Wilson,1924 

= Haplostoma Chatton & Brément,1915 

H. brevicauda (Canu,1886) () 

@ Synoicum pulmonaria, Polyclinum aurantium, Sidnyum turbinatum, Morchellium argus & Aplidium 

nordmanni 

Bohuslän-W Norway-E England-NE & SW Ireland-N France 

H. eruca (Norman,1869) 

@ Ciona intestinalis 

Oslofjord-Shetlands-E Scotland-NE Ireland 

H. banyulensis Brément,1909 

@ Trididemnum tenerum & Didemnum maculosum 

NE Ireland, French Mediterranean 

H. canui Chatton & Harant,1924 

@ Polyclinum aurantium 

W France 

H. mizoulei Monniot,1962 

@ stomach of the small intestinal form of Sidnyum turbinatum 

SW Ireland, Mediterranean 

Haplostomides Chatton & Harant,1924 

H. scotti Chatton & Harant,1924 () 


NE Ireland-N France 

H. hibernicus (T. & A. Scott,1895) 


NE & SW Ireland-N France 

?H. beaumonti (T. & A. Scott,1895) 

=Enterocola beaumonti T. & A. Scott,1895 

@ unknown ascidian 

* Considered as Haplostominae, sp. incertae sedis, by Ooishi & Illg,1977. Not mentioned by Gotto,1993. 

Ireland (Valencia) 

Haplosaccus Chatton & Harant,1924 

H. sacculus (Chatton & Brément,1910) () 

@ ? 


?Britain 

Haplostomella Chatton & Harant,1924 

= Rhabdomorpha Fukui,1965 ( R. halocynthiae Fukui,1965 (Japan)) 

H. malacocera Chatton & Harant,1924 () 

@ ? 


? 

Botryllophilinae Illg & Dudley,1980 

@ Ascidians 

Botryllophilus Hesse,1864 

= Kossmechthrus Della Valle,1883 ( K. notopus Della Valle,1883) 

= Blakeanus Wilson,1921 ( B. corniger Wilson,1921) 

B. ruber Hesse,1864 () 

@ Botryllus stchlosseri, Botrylloides leachi, Aplidium punctum, Morchellium argus, Sidnyum spp., 

Synoicum pulmonaria & Clavelina lepadiformis 

14 

* This taxon is problematic and need revision. According to Gotto,1993 probably several species have been 

lumped in this taxon. 

?Bohuslän-?W Norway-?Arctic Seas-?Br. Isles-?Engl. Channel-Brest-?Mediterranean 

B. brevipes G.O. Sars,1921 

non = B. brevipes Brément,1909 

* may possibly be a synonyme of B. bergensis Schellenberg,1921 

@ Botryllus sp. 

Bergen 

B. macropus Canu,1891 

@ Lithonephria eugyranda Giard 

France (Wimereux) 

B. bergensis Schellenberg,1921 

@ Leptoclinides faeroensis (Bjerkan) 

* Not mentioned by Gotto,1993. 

Bergen 

B. norvegicus Schellenberg,1921 

= Blakeanus corniger Wilson,1921 

= Blakeanus groenlandicus Hansen,1923 

@ Pelonaia corrugata Goodsir, Cynthia carnea Verrill & Cynthia rustica Linnaeus 


Trondheim-W Greenland-Long Island 

B. aspinosus Schellenberg,1922 

@ Polycarpa pomaria (Savigny) & Styela hupferi Michaelsen 


Plymouth, Angola 

B. virescens Hesse,1864 

* indeterminable. Sp. obl. 

@ ? 

Brest 

Schizoproctinae Aurivillius,1887 

* Neither family nor genus is mentioned by Gotto,1993. 

Schizoproctus Aurivillius,1885 

= Pteropygus G.O. Sars,1921 

S. inflatus Aurivillius,1885 () 

@ Ascidia obliqua and other ascidians 

N Norway-Spitsbergen-Arctic Seas-E Greenland 

S.vestitus (G.O. Sars,1921) 

= Pteropygus vestitus G.O. Sars,1921 

@ Ascidia obliqua 

S Norway (Risør) 

Enterognathinae Illg & Dudley,1980 

Enterognathus Giesbrecht,1900 

E. comatulae Giesbrecht,1900 () 

@ guts of ?Antedon mediterranea Lamarck, ?A. rosacea (Linck) & A. bifida (Pennant) 

E Scotland, E Ireland, Hebrides,Plymouth,Mediterranean 

LERNAEOINA 

Lernaeidae Cobbold,1879 

* Only fresh-water species recognized from Scandinavia 

@ Fish 

POECILOSTOMATOIDA Thorell,1859 

Lateracanthus ??,1??? 

* Which family? 

L. quadripedis ??,1??? 

@ Coryphaenoides rupestris 

NW Atlantic 

L. curtus Kabata, 1993 

@ Macrourus 

NE Atlantic 

Bomolochidae Sumpf,1871 

@ Fish 

Bomolochus von Nordmann,1832 

@ Marine Teleosts 

B. soleae Claus,1864 () 

@ in the nose of Solea solea 

Helgoland-Holland-Belgium-W France-Mediterranean 

B. bellones Burmeister,1835 ( of Parabomolochus Vervoort,1962) 

@ Belonidae 

Helgoland-Br. Isles-Mediterranean 

Holobomolochus Vervoort,1969 ( H. nothrus (Wilson,1913)) 

H. confusus (Stock,1953) 

= Bomolochus soleae : T. Scott,1893 & : Auctt., non Claus,1864 

@ nasal capsules of Gadidae, Conger, Cyclopterus & Labrus bergylta 

Bohuslän-North Sea-Br. Isles-Eur. Atlantic 

Taeniacanthidae Wilson,1911 

@ Fish, chiefly teleosts & Echinosocius Humes & Cressey,1959 & Echinirus Humes & Cressey,1959 on 

Echinoderms 

Taeniacanthus Sumpf,1871 ( T. carchariae Sumpf,1871) 

= Anchistrotos Brian,1906 

T. wilsoni A. Scott,1929 

@ Raja fullonica, R. montagui & ?R. naevus 

Irish Sea, Falmouth Bay & Yorkshire 

T. lucipetus (Holmes,1985) 

= Anchistrotos lucipetus Holmes,1985 

@ ? 

Ireland 

T. onosi (T. Scott,1902) 

= Bomolochus onosi T. Scott,1902 

@ Ciliata mustela, C. septentrionalis & Rhinonemus cimbrius 

Shetlands-Scotland 

T. laqueus (Leigh-Sharpe,1935) 

= Anchistrotos laqueus Leigh-Sharpe,1935 

= Anchistrotos hamatus Rounds,1960 

@ Ciliata mustela, Serranus cabrilla & S. scriba

W Baltic, Plymouth, Mediterranean 

T. zeugopteri (T. Scott,1902) 

= Bomolochus zeugopteri T. Scott,1902 

@ Zeugopterus punctatus 

Clyde Sea-Roscoff 

Ergasilidae von Nordmann,1832 

@ Marine & limnic teleosts (& gen. Ostrincola in bivalve mantle cavities) 

Ergasilus von Nordmann,1832 

= Ergasiloides G.O. Sars, 

E. briani Markevich,1933 

= E. minor Halisch,1934 

@ Perca fluviatilis, Leuciscus rutilus, Abramis brama etc. 

Baltic, Black Sea, Caspian Sea & several European fresh water systems 

E. gibbus von Nordmann,1832 

@ gills of Anguilla anguilla (species specific) 

Brackish Baltic & North Sea waters 

E. lizae Krøyer,1863 

= E. nanus van Beneden,1870 

@ limnic fishes, Mugilidae, Alosa & Engraulis 

Aberdeen-Mediterranean-Black Sea, Azov Sea (limnic & brackish waters) 

E. mugilis Vogt,1877 (sp. inquirenda) 

@ Mugilidae 

? 

E. sieboldi von Nordmann,1832 () 

= E. trisetaceus von Nordmann,1832 

=? E. esocis Sumpf,1871 

= E. surbecki Baumann,1912 

= E. hoferi Borodin,1915 

= E. baicalensis Messjatzeff,1926 

@ on gills of limnic fishes & E Baltic Anguilla anguilla & Platichthys flesus 

Baltic-Br. Isles-Eurasia 

Thersitina Norman,1905 

T. gasterostei (Pagenstecher,1861) 

= Thersites gasterostei Pagenstecher,1861 () 

= Ergasilus biuncinatus Gadd,1901 

=? Thersitina inopinata Percival,1937 

@ branchiae of Gasterosteus, Pungitius a.o. 

Scandinavia (e.g. Öresund, S Baltic & Furusund)-Br. Isles-Faeroes-Greenland-W France-Caspian Sea 

Chondracanthidae H. Milne-Edwards,1840 

@ Ectoparasites on fish, chiefly teleosts 

Chondracanthus Delaroche,1811 

C. zei Delaroche,1811 () 

= Lerneacantha delarochiana de Blainville,1822 

@ On gills of Zeus faber & other Zeidae 

Plymouth-Mediterranean 

C. neali Leigh-Sharpe,1930 

@ Malacocephalus laevis 

off SW Ireland-S Africa 

C. lophii Johnston,1836 

= Lernentoma Dufresnii de Blainville,1822 

= Chondracanthus Lophius Risso,1826 

= Chondracanthus Delarochiana Cuvier,1829 

= Chondracanthus gibbosus Krøyer,1837 

= Chondracanthus abdominalis Heegaard,1943 

@ On gills of Lophius spp. 

?Öresund-Kattegatt-Norw. Sea-Faeroes-North Sea-Br. Isles-Mediterranean 

C. merluccii (Holten,1802) 

= Lernaea merluccii Holten,1802 

= Chondracanthus xiphiae Guérin-Meneville,1829 

= Chondracanthus stramineus Wilson,1923 

@ On gills of Merluccius spp & ?Trisopterus luscus 

Norw. Sea-Br. Isles-S Africa 

C. nodosus (O.F. Müller,1776) 

= Lernaea nodosa O.F. Müller,1776 

= Chondracanthus williamsoni T. Scott,1909 

@ Sebastes spp. 

?Öresund-North Sea-Norw. Sea-Barents Sea-Greenland-Iceland-Faeroes-Br. Isles 

C. ornatus T. Scott,1900 

@ Callionymus maculatus & C. festivus 

E & W Scotland-Mediterranean 

Acanthochondria Oakley,1927 

A. cornuta cornuta (O.F. Müller,1776) 

= Lernaea cornuta O.F. Müller,1776 () 

= Chondracanthus flurae Krøyer,1863 

= Chondracanthus depressus T Scott,1905 

@ Glyptocephalus, Pleuronectes, Hippoglossoides, Platichthys & possibly other flatfishes 

Kattegatt-North Sea-Norw. Sea-Faeroes-North Atlantic 

A. clavata (Bassett-Smith,1896) 

= Chondracanthus clavatus Bassett-Smith,1896 

@ Microstomus kitt 

North Sea-Norw. Sea-White Sea-Iceland-Faeroes-Br. Isles-Bay of Biscay 

A. limandae (Krøyer,1863) 

= Chondracanthus limandae Krøyer,1863 

@ Limanda limanda 

North Sea-Murmansk-Faeroes-Br. Isles 

A. soleae (Krøyer,1838) 

= Chondracanthus soleae Krøyer,1838 

@ Solea solea & Scophthalmus rhombus 

North Sea-Faeroes-Br. Isles-Mauretania 

Acanthochondrites Oakley,1930 

A. annulatus (Olsson,1869) 

= Chondracanthus annulatus Olsson,1869 () 

= Chondracanthus inflatus Bainbridge,1909 

= Chondracanthus japonicus Gusev,1951 

=?? Chondracanthus pallidus van Beneden,1851 

@ Raja spp. 

North Sea-Norw. Sea-Br. Isles-N Atlantic 

Lernentoma de Blainville,1822 

L. asellina (Linnaeus,1758) 

=? Chondracanthus gurnardi : Olsson,1869 

@ Triglidae & possibly other marine teleosts 

North Sea-S Norw. Sea-Br. Isles-NE Atlantic-W Mediterranean 

Diocus Krøyer,1863 

D. frigidus Hansen,1923 

@ Lycodes frigidus 

Iceland 

Philichthyidae Vogt,1877, emend. Delamare Deboutteville,1962 

= Sarcotacidae Yamaguti,1963 

@ Inhabiting the subcutaneous mucus ducts or lateral line canals of Teleosts 

Sarcotaces Olsson,1872 

S. arcticus Collett,18?? 

@ endoparasitic in "Molva abyssorum ". Blood from engorged females may stain the flesh of the host. 

North Sea 

Philichthys Steenstrup,1862 

P. xiphiae Steenstrup,1862 () 

@ Xiphias gladius 

Kattegatt-North Sea-Br. Isles-Mediterranean- North Atlantic 

Colobomatus Hesse,1873 

C. bergyltae Hesse,1873 

@ Labrus bergylta 

* Not mentioned by Kabata,1992 

France 

C. mugilis Raibaut, Caillet & Ben Hassine,1978 

@ Liza aurata 

Mediterranean 

Leposphilus Hesse,1866 

L. labrei Hesse,1886 

@ subcutaneous ducts in the frontal region of the head of Centrolabrus exoletus & Cossyphus diplotaenia 


France 

Oncaeidae Giesbrecht,1892, emend. Boxshall,1977 

@ probably pelagic semiparasites, but hosts not known 

Oncaea Philippi,1843 

O. borealis G.O. Sars,1918 

= O. conifera : G.O. Sars, non Giesbrecht,1891 

Oslofjord-S & W Norway-Arctic Seas-Br. Isles 

O. conifera Giesbrecht,1891 

off W Norway, Cape Verde 

O. similis G.O. Sars,1918 

Norway (Bergen area) 

O. minuta Giesbrecht,1892 

Gullmarfjord, Skagerrak, Norway (Bergen area) 

O. subtilis Giesbrecht,1892 

Skagerrak-Norway-Ireland-Mediterran. 

O. obscura Farran,1908 

Irish Atlantic slope 

O. ornata Giesbrecht,1891 

Engl. Channel-Ireland, Cape Verde 

O. mediterranea (Claus,1863) 

= Antaria mediterranea Claus,1863 

Mediterranean, Cape Verde 

O. rotundata Boxshall,1977 

Cape Verde 

O. notopus Giesbrecht,1891 

Cape Verde 

O. media Giesbrecht,1891 

= O curta : Estrada & Genicio,1970, non G.O. Sars,1916 

Cape Verde 

O. ivlevi Schmeleva,1966 

?NE Atlantic, Adriatic 

O. curta G. O. Sars,1916 

= O. ovalis Shmeleva,1966 

= O. longiseta Schmeleva,1969 

= O. latimana Gordeyeva,1975 

= O. sp.1 Ferrari,1975 

= O. sp.2 Ferrari,1975 

Cape Verde 

O. venusta Philippi,1843 () 

= Antaria obtusa Dana,1852 

= Oncaea pyriformis Lubbock,1860 

= Antaria coerulescens Claus,1866 

Cape Verde 

Lubbockia Claus,1863 

L. glacialis G.O. Sars,1900 

High arctic regions, Cape Verde 

L. brevis Farran,1908 

NE Atlantic 

L. extenuata Boxshall,1977 

Cape Verde 

L. minuta Wolfenden,1905 

NE Atlantic 

L. squillimana Claus,1863 () 

Cape Verde 

L. aculeata Giesbrecht,1891 

Cape Verde 

Pseudolubbockia G.O. Sars,1909 

15

P. dilatata G.O. Sars,1909 

?NE Atlantic 

Conaea Giesbrecht,1891 

C. gracilis (Dana,1852) T. Scott,1894 

= Antaria gracilis Dana,1852 

= Conaea rapax Giesbrecht,1891 () 

Norway (Sognesjøen - according to Wiborg 1954), Cape Verde 

Pachos Stebbing,1910 

= Pachysoma Claus,1863 (preoccupied) 

P. punctatum (Claus,1863) () 

= Pachysoma punctata Claus,1863 

Cape Verde 

P. tuberosum (Giesbrecht,1891) 

= Pachysoma tuberosum Giesbrecht,1891 

= Pachysoma dentatum Mori,1932 

Cape Verde 

Epicalymna Heron,1977 

E. exigua (Farran,1908) 

= Oncaea exigua Farran,1908 

off Ireland 

Paralubbockiidae Boxshall & Huys,1990 

Paralubbockia Boxshall,1977 

P. longipedia Boxshall, 1977 () 

Cape Verde 

Corycaeidae Dana,1852 

@ probably pelagic semiparasites on gelatinous zooplankton like appendicularians and salps. 

Corycaeus Dana,1845 

C. (Ditrichocorycaeus M. Dahl,1912) anglicus Lubbock,1855 

Kattegatt-Bohuslän-Oslofjord-W Norway-North Sea-Br. Isles-Engl. Channel-?Biscay 

C. (Corycaeus) clausi F. Dahl,1894 

N temperate & tropical Atlantic, Mediterranean 

C. (Corycaeus) speciosus Dana,1849 

= Corycaeus remiger Dana,1849-52 

= Corycaeus varius Dana,1849-52 

= Corycaeus longicaudatus Dana,1849-52 

= Corycaeus Sutherlandi Lubbock,1856 

= Corycaeus styliferus Lubbock,1856 


C. (Agetus Krøyer,1849) flaccus Giesbrecht,1891 


C. (Urocorycaeus M. Dahl,1912) furcifer Claus,1863 

Biscay, temperate & tropical Atlantic, Mediterranean 

C. (Onchocorycaeus M. Dahl,1912) giesbrechti F. Dahl,1894 

Temperate & tropical Atlantic, Mediterranean 

C. venustus Dana,1849 

?NE Atlantic 

Corycella Farran,1911 

C. rostrata (Claus,1863) 

= Corycaeus rostratus Claus,1863 

= Corycaeus parvus Claus,1863 

= Corycaeus pellucidus Brady,1883 (p.p.) 

Boreal & temperate Atlantic, Mediterranean 

Sapphirinidae Thorell,1859 

@ Salps 

Sapphirina J.V. Thompson,1829 

S. iris Dana,1849 

= Sapphirina salpae Claus,1859 

= Sapphirina scalaris Fisher,1860 

= Sapphirina longifurca Scott,1909 

@ Salps 

?W Norway-NE Atlantic-Mediterranean 

S. angusta Dana,1849 

= Sapphirina Clausi Haeckel,1864 

= Sapphirina Danae Lubbock,1856 

= Sapphirina opaca Lubbock,1856 

Boreal Atlantic, Mediterranean 

S. nigromaculata Claus,1863 

= Sapphirhina lomae Esterly,1905 

Temperate & tropical Atlantic, Biscay, Mediterranean 

S. opalina Dana,1849 

= Sapphirhina versicolor Dana,1849 

= Sapphirhina Darwini Haeckel,1864 

= Sapphirhina Thompsoni Lubbock,1860 

= Sapphirhina pachygaster Claus,1863 

= Sapphirhina aureofurca Giesbrecht,1891 

Temperate & tropical Atlantic, Mediterranean 

S. ovatolanceolata Dana,1849 

= Sapphirhina gemma Dana,1849 

= Sapphirhina Edwardsii Haeckel,1864 

= Sapphirhina Gegenbaueri Haeckel,1864 

Gibraltar, temperate & tropical Atlantic, Mediterranean 

S. sali Farran,1929 

= Sapphirina pseudolactens Lehnhofer,1929 

= Sapphirina gibba Rose,1929 

Temperate & tropical Atlantic, Biscay 

Copilia Dana,1849 

= Saphirinella Claus,1863 

= Hyalophyllum Haeckel,1864 

C. vitrea Haeckel,1864 


Corina Giesbrecht,1891 

C. granulosa Giesbrecht,1891 

off W Ireland, Mediterranean 

Urocopiidae Humes & Stock,1972 

@ not known, but probably gelatinous zooplankton 

Urocopia G.O. Sars,1917 

U. singularis G.O. Sars,1917 

pelagic S of Iceland 

Clausiidae Giesbrecht,1895 

@ Polychaeta & possibly other invertebrates like teredinid bivalves 

Clausia Claparède,1863 

C. lubbocki Claparède,1863 

@ ? 

E England, Engl. Channel 

C. uniseta Bocquet & Stock,1960 

@ ? 

E England 

Rhodinicola Levinsen,1878 

= Seridium Giesbrecht,1895 

@ Maldanidae 

R. elongata Levinsen,1878 

@ Rhodine gracilior 

Öresund, SE Scotland, NE England, ?W Ireland 

R. gibbosa Bresciani,1964 

@ Praxillella praetermissa 

* Not mentioned by Goitto,1993 

Öresund 

Conchocheres G.O. Sars,1918 

* Tentatively put in this fam. Systematic position not fully clear. 

C. malleolatus G.O. Sars,1918 

@ pallial cavity of Cuspidaria obesa 


W Norway 

Megaclausia O'Reilly,1995 

M. mirabilis O'Reilly,1995 () 

@ In tubes of Rhodine gracilior 

SE Scotland, NE England 

Mesnilia Canu,1898 

M. martinensis Canu,1898 

@ Polydora flava 

Engl. Channel 

M. cluthae (T. & A. Scott,1896) 

@ Polydora ciliata & P. flava 

NE, E & S England, SW Scotland, N France 

Pseudoclausia Bocquet & Stock,1960 

P. longiseta Bocquet & Stock,1963 

@ Hymeniacidon sanguinea & ?Aplysina aerophoba 


Clausidiidae Embleton,1901 

@ Sponges, Octocorals, Polychaets, Bivalves, Sipunculids, Nudibranchs, Stomatopods, Thalassinideans 

Conchyliurus Bocquet & Stock,1957 

@ Bivalvia 

C. cardii Gooding,1957 

@ Mantle cavity of Acanthocardia spp., Tapes decussatus, Venerupis spp., Scrobicularia plana, Petricola 

pholadiformis, Barnea candida, Meretrix chione 

* Two subspecies are recognized: C. cardii cardii Gooding,1957 @ Acanthocardia echinata (Linnaeus) & 

Meretrix chione (Linnaeus) and C. cardii tapetis Bocquet & Stock,1958 @ Venerupis aurea . 

E & SW Ireland, SW England, N & W France 

C. solenis Bocquet & Stock,1957 () 

@ Mantle cavity of Solen marginatus Don. 

Roscoff 

Giardella Canu,1888 

G. callianassae Canu,1888 

@ Burrows of Callianassa subterranea 

NW England, Engl. Channel 

G. thompsoni A. Scott,1906 

@ ? 

Irish Sea 

Hemicyclops Boeck,1872 

H. purpureus Boeck,1872 

@ not known 

Bohuslän-Oslofjord-SE Scotland-SW England 

H. cylindracea (Pelseneer,1929) 

@ Mantle cavity of Loripes lacteus 

N & W France 

H. aberdonensis (T & A. Scott,1892) 

@ ? 

E England, NE Scotland, E & SW Ireland, W France 

Hersiliodes Canu,1888 

@ Maldanidae 

H. latericia (Grube,1869) 

@ in tube of Leiochone clypeata 

E England, W & SW Ireland, N & W France, Mediterranean 

Hippomolgus G.O. Sars,1918 

H. furcifer G.O. Sars,1918 

@ ? 

S Norway (Risør)-Stavanger 

16

Leptinogaster Pelseneer,1929 

L. histrio (Pelseneer,1929) 

@ Macoma balthica (Linnaeus) & Abra alba 

German Bight, Engl. Channel 

L. pholadis Pelseneer,1929 

@ Pholas dactylus 

Brest, Naples 

L. sp. 

@ Digestive tract of Echinocardium cordatum 

Netherlands 

Pseudopsyllus T. Scott,1902 

P. elongatus T. Scott,1902 

@ ? 

?Britain 

Eunicicolidae G.O. Sars,1918 

@ Eunicid Polychaeta 

Eunicicola Kurz,1877 

E. clausi Kurz, sensu G.O. Sars, ?non Kurz,1877 

@ Eunice claparedi 

* Sensu Gotto,1993 is the North Sea form recorded by Sars differing from the true Mediterranean E. clausi 

Kurz in several significant details. 

Oslofjord-S Norway (Risør)-Faeroes-Br. Isles-Mediterranean 

E. insolens (T. & A. Scott,18??) 

@ Eunice harassii Audouin & Milne-Edwards 

SW Scotland- NE Ireland 

Catiniidae Bocquet & Stock,1957 

@ Sipunculids 

Catinia Bocquet & Stock,1957 

C. plana Bocquet & Stock,1957 () 

@ On skin of Sipunculus nudus 

Engl. Channel, W France 

Myzomolgus Bocquet & Stock,1957 

M. stupendus Bocquet & Stock,1957 () 

@ On skin of Sipunculus nudus 

* Ho,1984 suggests a closer affinity to Catiniidae than to Clausidiidae. 

Engl. Channel, W France 

Anomopsyllidae G.O. Sars,1921 

Anomopsyllus G.O. Sars,1921 

A. pranizoides G.O. Sars,1921 

@ ? 

Oslofjord 

Nereicolidae Claus,1875 

@ Ectoassociates of errant Polychaeta 

Nereicola Keferstein,1863 

N. ovatus Keferstein,1863 

@ Platynereis dumerilii, Perinereis cultrifera, Nereis rava, Nereis zonata, N. sp. 

Engl. Channel, Mediterranean, Black Sea 

Flabelliphilus Bresciani & Lützen,1962 

* Systematic position not clear. Nereicolid affinity suggested by Stock,1968. 

F. inersus Bresciani & Lützen,1962 

@ Among gelatinous protuberances of the jelly coat covering the body of Flabelligera affinis 

Bohuslän 

Sigecheres Bresciani,1964 

S. brittae Bresciani,1964 

=? Nereicola concinna T. Scott,1902 

@ Sige fusigera 

Öresund-Kattegatt-Bohuslän, ?W Scotland 

Selioides Levinsen,1878 

S. bolbroei Levinsen,1878 

= Cancerina confusa T. Scott,1900 

@ Harmothoe imbricata, Gattyana cirrosa, Eunoe nodosa, Lagisca rarispina, Antinoe sarsi & Hermadion 

rouchi 

Öresund-Bohuslän-North Sea-Britain-Arctic Seas-Iceland 

S. bocqueti Carton,1963 

@ Scalisetosus assimilis 

Roscoff 

Spiophanicolidae Ho,1984 

* Included in the "Nereicoliform group" by Ho 1984. 

Spiophanicola Ho,1984 ( S. spinulosus Ho,1984 - California @ Spiophanes spp.) 

Entobiidae Ho,1984 

@ Terebellidae 

* This family is not recognized by Huys & Boxshall 1991, but included in the "Nereicoliform group" by Ho 

1984. 

Entobius Dogiel,1908 

E. loimiae Dogiel,1908 

@ ?Loimia sp. 


? 

E. hamondi Gotto,1966 

@ Polycirrus caliendrum 

SW England 

Mytilicolidae Bocquet & Stock,1957 

@ Bivalves & Gastropods 

Mytilicola Steuer,1902 

M. intestinalis Steuer,1902 

@ Mytilus edulis & M. galloprovincialis (& Cerastoderma edule, Ostrea edulis & Tapes decussatus) 

Limfjorden-North Sea-Br. Isles-Portugal-Mediterranean 

M. orientalis Mori,1935 

= M. ostreae C.B. Wilson,1938 

=? M. mactrae Hoshina & Kuwabara,1959 

@ Crassostrea gigas & Mytilus crassitesta & ? Mactra veneriformis 

* Introduced from Japan to Pacific N America, there infesting e.g. Mytilus edulis, M. californicus, Ostrea 

lurida & Crepidula fornicata, later introduced in W France with imported Crassostrea gigas, there infesting 

also Mytilus spp. & Ostrea edulis - Not mentioned by Gotto,1993 

Arcachon, NE Pacific, Japan 

Trochicola Dollfus,1914 

T. entericus Dollfus,1914 

@ Rectum of Gibbula cineraria, G. varia, Calliostoma zizyphinum & Monodonta mutabilis 


T. pectinidarum Tuzet & Ormières,1961 

@ Aequipecten opercularis & Pecten glaber 


French Mediterranean 

Anthessiidae Humes,1986 

@ Mollusca 

Anthessius Della Valle,1880 

= Pseudomolgus G.O. Sars,1916 

A. dilatatus (G.O. Sars,1918) 

Pseudomolgus dilatatus G.O. Sars,1918 

@ ? 


W Norway (RomsdalFjord) 

A. arenicolus (Brady,1872) 

= Lichomolgus arenicolus Brady,1872 

@ Dosinia exoleta 

NE England, SW Scotland, SW Ireland, Engl. Channel, W France 

A. leptostylis (G.O. Sars,1916) 

= Pseudomolgus leptostylis G.O. Sars,1916 

@ Buccinum undatum 

Bohuslän-Oslofjord-Trondheim-Spitsbergen, E England 

A. arcuatus López-González, Conradi, Naranjo & García-Gómez,1992 

@ Berthella stellata (Risso,1826) 

Cádiz, Spain 

A. teissieri Bocquet & Stock,1958 

= Pseudomolgus arenicola : G.O. Sars,1921, non Lichomolgus arenicolus Brady,1872 

@ Buccinum undatum 

Hvalør, E England, Engl. Channel 

A. pleurobrancheae Della Valle,1880 

@ Pleurobranchaea sp. 

Napoli 

Myicolidae Yamaguti,1936 

@ Molluscs 

Myicola Wright,1885 

M. ostreae ??,1??? 

@ Crassostrea gigas, Ostrea edulis & Crassostrea angulata 

Arcachon 

Pseudomyicola Yamaguti,1936 

P. spinosus (Raffaele & Monticelli,1885) 

@ Mytilus galloprovincialis, M. edulis, Pinna pectinata, Ostrea stentina & Pteria hirundo 

Arcachon-Mediterranean-Black Sea-Bermudas-Caribbean 

Rhinomolgus G.O. Sars,1918 

* Humes & Stock,1973 put this genus in this fam. (Myicolidae)! 

R. anomalus G.O. Sars,1918 

@ Antedon 

Trondheimfjord 

Ventriculinidae Leigh-Sharpe,1934 

@ Sipunculoids & Gastropods 

Endocheres Bocquet & Stock,1956 

E. obscurus Bocquet & Stock,1956 

@ In rectal gland or in galls developed in the pallial cavity of Calliostoma zizyphinum (Linnaeus) 

N France 

Antheacheridae M. Sars,1870 

* This family includes perhaps (sensu Ho,1981) three genera parasiting sipunculans: Akessonia, Coelotropus 

& Siphonobius 

@ Actinian mesenteries ?& Sipunculids 

Antheacheres M. Sars,1857 

A. duebeni M. Sars,1857 

@ In rounded galls formed from the mesentery walls of Bolocera tuedie 

Bohuslän-W & N Norway 

Akessonia Bresciani & Lützen,1962 

* Family placement probably not definitive, but Humes,1987 thinks this for the moment might be a proper 

family to keep this genus in. 

A. occulta Bresciani & Lützen,1962 

@ In coelomic cavity of Golfingia minuta 

Öresund-Bohuslän 

Gastroecus Hansen,1886 

G. arcticus Hansen,1886 () 

@ Actinostola intermedia & Actinostola spetsbergensis Carlgren 


Kara Sea 

Staurosoma Will,1844 

S. parasiticum Will,1844 

@ In galls developed on the mesenteries of Anemonia sulcata 

Normandie-Mediterranean 

Lamippidae Joliet,1882 

@ Octocorals 

Lamippe Bruzelius,1858 

17

L. rubra Bruzelius,1858 

@ Pennatula phosphorea 

Skagerrak, S Norway, Mediterranean 

Lamipella Bouligand & Delamare Deboutteville,1959 

L. faurei Bouligand & Delamare Deboutteville,1959 

@ In coenosarcal canals of Swiftia rosea & other octocorals 

Bohuslän, Mediterranean 

Enalcyonium Olsson,1869 

E. rubicundum Olsson,1869 

@ In coenosarcal canals & sometimes within host polyps of Alcyonium digitatum, A. palmatum & A. acaule 

Bohuslän-Br. Isles, NE Ireland, N France, ?Mediterranean 

E. olssoni (De Zulueta,1908) 

@ Alcyonium digitatum 

Bohuslän 

E. forbesi (T. Scott,1901) 


SE Scotland, Isle of Man-Wales-N France 

Mesoglicolidae de Zulueta,1911 

* Order uncertain. Lamippidae and Ascidicolidea have been discussed as possibly related. 

@ Anthozoans 

Mesoglicola Quidor,1906 

M. delagei Quidor,1906 

@ In mesogloea and in galls of Corynactis viridis Allman 

Roscoff & Banyuls 

Jeanella T. Scott,1904 

J. minor (T. Scott,1902) 

@ ? 

* Correct family? 

?Britain 

Splanchnotrophidae Norman & Scott,1906 

@ Molluscs (chiefly Nudibranchs) & Pterobranchia 

Splanchnotrophus Hancock & Norman,1863 

=? Lomanoticola T. & A. Scott,1895 

* Not yet seen revision by Delamare-Deboutteville,1950 

S. brevipes Hancock & Norman,1863 

@ Coryphella verrucosa, Facelina curta, Doto spp., Cuthona genovae 

Isefjord, Kattegatt-Bohuslän-Br. Isles, Ireland 

S. gracilis Hancock & Norman,1863 

@ Endoparasitic in Acanthodoris pilosa, Okenia aspersa, Eubranchus pallidus & Doto coronata 

SE Scotland, E & SW England, Isle of Man, W Ireland 

S. insolens (T. & A. Scott,1895) 

= Lomanoticola insolens T. & A. Scott,1895 

@ Lomanotus genei 

* Of uncertain status 

SW Ireland 

S. willemi ?,19?? 

@ Facelina coronata 


? 

S. angulatus ?,19?? 

@ Aeolidia papillosa, Aeolidiella glauca 


? 

Gastrodelphyidae List,1890 

Gastrodelphys Graeffe,1883 

G. clausii Graeffe 

@ Bispira volutacornis 


Anomoclausiidae Gotto,1964 

Corallovexiidae Stock,1975 

Echiurophilidae Delamare-Deboutteville & Nunes-Ruivo,1955 

Pharodidae Illg,1948 

Philoblennidae Izawa,1976 

@ on a cowry 

Rhynchomolgidae Humes & Stock,1972 

Serpulidicolidae Stock,1979 

Serpulidicola Southword,1963 

S. placostegi Southword,1963 

@ Placostegus tridentatus 

W entrance of Engl. Channel in 450-760 m 

Shiinoidae Cressey,1975 

Synaptiphilidae Bocquet,1953 

@ Holothuroida 

Synaptiphilus Canu & Cuénot,1892 

= Colaceutes Hartman,1856 (nomen oblitum) 

= Remigulus T. & A. Scott,1893 

* This genus was originally placed in a separate family, Synaptiphilidae Bocquet,1952, moved to Clausiidae 

by Bocquet & Stock,1957 but suggested by Ho,1984 to have closer affinities to Clausidiidae. 

@ Echinoderms 

S. luteus Canu & Cuénot,1892 () 

= Colaceutes mulleri Hartmann,1856 (nomen oblitum) 

@ Labidoplax digitata (Montagu), Leptosynapta galliennei (Herapath), L. inhaerens (Müller) & L. 

bergensis Östergren 

SW Br. Isles-N & W France-Mediterranean 

S. cantacuzenei Bocquet & Stock,1957 

@ Labidoplax digitata 

NW France-Mediterranean 

S. tridens (T. & A. Scott,1893) 

= Remigulus tridens T. & A. Scott,1893 

@ Leptosynapta inhaerens (Müller), Leptosynapta cruenta Cherbonnier & Labidoplax media 

W Scotland, Ireland, NW France 

Presynaptiphilus Bocquet & Stock,1960 


P. acrocnidae Bocquet & Stock,1960 

@ Acrocnida brachiata 

N France-SW England 

Telsidae Ho,1967 

Tuccidae Vervoort,1962 

Vahiniidae Humes,1967 

Xarifiidae Humes,1960 

Lichomolgoidea Humes & Stock,1972 

Lichomolgidae Kossmann,1877 

@ very wide host spectrum 

Acaenomolgus Humes & Stock,1972 

A. protulae (Stock,1959) 

@ Protula tubularia & Protula intestinum 

NE Ireland-SW Scotland, Mediterranean 

A. serpulae (Stock,1960) 

@ Serpula vermicularis 

Mediterranean France 

Astericola Rosoll,1889 

A. clausi Rosoll,1889 () 

@ Marthasterias glacialis, Asterina gibbosa & Astropecten aurantium 

Roscoff-European Atlantic, Mediterranean 

Critomolgus Humes & Stock,1983 

C. actiniae (Della Valle,1880) 

= Doridicola actiniae (Della Valle,1880) 

= Lichomolgus actiniae Della Valle,1880 

@ Anemonia sulcata & Actinia spp. 

Plymouth-NE Ireland-Roscoff-Mediterranean 

C. bulbipes (Stock & Kleeton,1963) 

@ ? 

N? Europe 

Doridicola Leydig,1853 

=? Metaxymolgus Humes & Stock,1972 

D. agilis Leydig,1853 

= Eolidicola tenax M. Sars,1862 

= Lichomolgus agilis : Auctt. 

= Doricola agilis : Odhner,1922 

@ many different Nudibranchs (& Aplysia, Hiatella & Polynoe sp.) and ?Sagittatus sagittatus 

Bohuslän-North Sea-S & W Norway-Br. Isles-France-Mediterranean-Senegal 

D. chlamydis (Humes & Stock,1973) 

@ Chlamys varia 

European Atlantic 

D. longicauda (Claus,1860) 

= Metaxymolgus longicauda : (Claus,1860) Humes & Stock,1972 

@ gills & mantle cavity of Sepia officinalis & S. officinalis var. filliouxi 

Netherlands, S England, Arcachon, Biscay, Mediterranean 

D. hirsutipes (T. Scott,1893) 

= Macrocheiron hirutipes : G.O. Sars,1917 

= Metaxymolgus hirsutipes : (T. Scott,1893) Humes & Stock,1972 

@ ?tubes of Sabella pavonina 

S & W Norway, Novaya Semlja, SE Scotland, NW England 

Epimolgus Bocquet & Stock,1956 

E. trochi (Canu,1899) 

@ Gibbula spp. (e.g. G. cineraria ) & Monodonta 

Engl. Channel-W Mediterranean 

Lichomolgella G.O. Sars,1918 

L. pusilla G.O. Sars,1918 

@ ? 

S-SW Norway, SE Ireland 

Lichomolgides Gotto,1954 

L. cuanensis Gotto,1954 

@ cloacal cavities of Trididemnum tenerum (Verrill) 

NE Ireland 

Lichomolgus Thorell,1859 

L. albens Thorell,1859 

@ Ascidians, often Corella or e.g. Molgula manhattensis 

Isefjord, Bohuslän-North Sea-S & W Norway-Br. Isles-N & W France 

L. canui G.O. Sars,1917 

@ many different Ascidians, e.g. Molgula, Styela & Dendrodoa 

Kiel-Norway-Holland-N & S Ireland-France-Sicily 

L. diazonae Gotto,1961 

@ Diazona violacea Savigny 

Plymouth 

L. forficula Thorell,1859 

=? L. elongatus Buchholz,1869 

@ Ascidia mentula (& Psammechinus miliaris ) 

Bohuslän-S & W Norway-Br. Isles-N France-Mediterranean 

L. furcillatus Thorell,1859 

@ Ciona (& Corella parallelogramma ) 

Bohuslän-Norway-Shetlands-Br. Isles-Mediterranean 

L. leptodermatus Gooding,1957 

@ gills of Laevicardium crassum 

Engl. Channel & SW England 

L. marginatus Thorell,1859 

@ Ascidia spp. 

18

Bohuslän-Holland-Norway-N France-NE Ireland 

Macrochiron Brady,1872 

M. fucicolum Brady,1872 

@ Generally associated with algae. 

?Norway-S North Sea-Br. Isles, Ireland, NE USA, Mediterranean 

M. mutatum Stock,1957 

= Macrocheiron fucicolum : G.O. Sars,1917, non Brady,1872 (p.p.) 

Oslofjord-S & W Norway 

Octopicola Humes, in Delamare Deboutteville, Humes & Paris,1957 

O. superbus superbus Humes,1957 

@ Octopus vulgaris 

Roscoff-Mediterranean 

Zygomolgus Humes & Stock,1972 

Z. tenuifurcatus (G.O. Sars,1917) 

@ Labidoplax digitata & Diplosoma listerianum 

S &W Norway-NE & SW Ireland 

Z. curtiramus (Bocquet & Stock,1962) 

@ the alga Lithophyllum incrustans 

N France 

Z. didemni (Gotto,1956) 

@ Didemnum maculosum (Milne-Edwards) 

NE Ireland 

Z. poucheti (Canu,1891) 

= Lichomolgus poucheti Canu,1891 

@ Ascidians, e.g. Sidnyum , Morchellium & Fragarium 

S & W Norway-W Ireland-N & NW France 

Stellicola Kossmann,1877 


Pseudanthessidae Humes & Stock,1972 

@ Wide host spectrum 

Heteranthessius T. Scott,1904 

H. dubius (T. Scott,1903) 

@ ? 

Clyde Sea 

H. scotti Bocquet, Stock & Bénard,1959 

@ ? 

Bretagne, among Lithophyllum incrustans 

H. furcatus Stock,1971 

@ Microcosmus sabatieri 


Pseudanthessius Claus,1889 

P. gracilis Claus,1889 

@ Pomatoceros, Filograna, Spisula & Palmaria palmata 

Bohuslän-S & W Norway-Br. Isles-SW Ireland-N & W France-Mediterranean 

P. assimilis G.O. Sars,1917 

@ Echinus elegans 


Norway (Hardangerfjord & Romsdalsfjord) 

P. dubius G.O. Sars,1918 

@ Asterias rubens 


Bohuslän-W Norway 

P. liber (Brady,1880) 

@ Echinus esculentus , E. acutus, Psammechinus miliaris, Paracentrotus lividus &Strongylocentrotus 

droebakiensis 

Bohuslän-S & W Norway-Br. Isles-Faeroes-N France 

P. sauvagei Canu,1892 

@ Echinocardium cordatum, E. flavescens , Spatangus purpureus (& Echinus esculentus ) 

Bohuslän-S North Sea-S Norway-Br. Isles-N France-Mediterranean 

P. thorellii (Brady,1880) 

@ Swiftia pallida, Cladocora caespitosa & ?Asterias rubens 

Bohuslän-Br. Isles-SW Ireland-?Italy 

Nemerthessius Stock,1985 

N. nemertophilus (Gallien,1936) 

= Pseudanthessius nemertophilus Gallien,1936 

@ Lineus longissimus Sowerby 

Wimereux (NE France), W France 

Phyllodicolidae Delamare-Deboutteville & Laubier,1961 

@ Phyllodocidae 

Cyclorhiza Heegaard,1942 

C. eteonicola Heegard,1942 

@ Eteone longa 

Trondheimsfjord-?Isle of Man, E North America 

C. megalova Gotto & Leahy,1988 

@ Eteone longa 

W Ireland 

Sabelliphilidae Gurney,1927 

@ Wide host spectrum 

Lichomolgidium Kossmann,1877 

L. cynthiae (Brian,1924) 

=?? L. sardum Kossmann,1877 

@ Styela clava & Pyura microcosmus 

Plymouth-Roscoff-Mauretania, ??Mediterranean 

Modiolicola Aurivillius,1882 

M. insignis Aurivillius,1882 

@ Modiolus modiolus & Mytilus edulis & M. galloprovinsialis 

Bohuslän-SE Norway (Hvalør)-Netherlands-Br. Isles-Engl. Channel-Mediterranean-Black Sea 

M. inermis Canu,1892 

@ Aequipecten opercularis 

E Scotland, W Br. Isles-Engl. Channel-W France-W Mediterranean 

M. maximus (Thompson,1893) 

@ Pecten maximus (& Aequipecten opercularis f. audouini ) 

W Br. Isles-NE & SW Ireland,N & W France, Mediterranean 

Myxomolgus Humes & Stock,1972 

M. myxicolae (Bocquet & Stock,1958) 

@ Myxicola infundibulum (Rénier) 

S England-Finistère 

M. proximus Humes & Stock,1973 

@ Myxicola aesthetica (Claparède) 

Finistère 

Paranthessius Claus,1889 

P. anemoniae Claus,1889 

=? Doridicola antheae Ridley,1879 

@ Anemonia sulcata (Pennant) 

N France-E England-NE & S>W Ireland-Mediterranean 

? P. dubia (G.O. Sars,1921) 

= Hermannella dubia G.O. Sars,1921 

@ ? 


Norway (Drøbak) 

Herrmannella Canu,1891 

= Paranthessius : Auctt., non Claus,1889 

H. valida G.O. Sars,1918 

@ ? 


Norway (Stavanger) 

H. parva Norman & T. Scott,1905 

@ ?Eunicella stricta 

* The position of this species in Herrmannella is questionable according to Gotto,1993 

S & W Norway-Engl. Channel-W Mediterranean 

H. rostrata Canu,1891 

@ many different bivalves, including Cerastoderma edule, Spisula spp., Venerupis pullastra, Macoma 

balthica (& Ascidia mentula ) 

Elbe mouth-Holland-Engl. Channel-Br. Isles-Irish Sea-France-Mediterranean 

H. barneae (Pelseneer,1929) 

@ Barnea, Pholas & Ostrea 

E England, N & W France 

H. haploceras (Bocquet & Stock,1959) 

= Paranthessius colmani Reddiah,1960 

@ Laevicardium crassum 

Isle of Man, Engl. Channel, European Atlantic 

H. pecteni (Sowinski,1884) 

@ Pectenidae & Anomia ephippium 

W Br. Isles-NW & W France-Mediterranean & Black Sea 

H. duggani Holmes & Minchin,1991 

@ ? 


?Britain 

Sabelliphilus M. Sars,1862 

S. elongatus M. Sars,1862 () 

@ branchial crown ofSabella pavonina & Spirographis 

Bohuslän-S, W & NW Norway-Br. Isles-Engl. Channel-NE Ireland-W France-Mediterranean 

S. sarsi Claparède,1870 

@ body ofSpirographis spallanzanii & Sabella 

Engl. Channel-W France-Mediterranean 

Scambicornus Heegaard,1944 

= Preherrmannella Sewell,1949 

S. finmarchicus (T. Scott,1903) 

= Hermannella finmarchica : Auctt. 

= S. armoricanus (Bocquet, Stock & Kleeton,1963) 

@ Eupolymnia nebulosa 

S, W & N Norway, SW Ireland, Roscoff 

S. prehensilis (G.O. Sars,1918) 

= Hermannella prehensilis G.O. Sars,1918 

@ ? 


Norway (outside Sognefjord) 

S. tenuicaudis (G.O. Sars,1918) 

= Hermannella tenuicaudis G.O. Sars,1918 

@ ? 


W Norway 

Calypsina Humes & Stock,1972 


Calypsarion Humes & Ho,1969 


Caribulus Humes & Stock,1972 


Chauliolobion Humes,1975 


Diogenella Stock,1968 


Diogenidium Edwards,1891 


Lecanurius Kossmann,1877 


Lichothuria Stock,1968 


Synapticola Voigt,1892 


Ventriculinidae Leigh-Sharp,1934 

Family uncertain 

19

Gomphopodarion Humes,1974 

G. byssoicum Humes,1974 

@ Oneirophanta mutabilis Theel 

off W Ireland in ca 4430 m 

SIPHONOSTOMATOIDA Latreille,1829 

Caligidae Burmeister,1835 

@ Fish (& perhaps Cephalopods) 

Caligus O.F. Müller,1785 

* Are C. belones Krøyer,1863 & C. angustatus ?Krøyer,18?? good species or synonyms of other species. 

Both taxa are described from the North Sea? C. belones is found by P. Olsson in Skagerrak "ad corpus 

Belones vulgaris ". 

C. curtus O.F. Müller,1785 () 

= C. mulleri Leach,1816 

= C. bicuspidatus von Nordmann,1832 

= C. americanus Pickering & Dana,1838 

= C. elegans van Beneden,1851 

= C. fallax Krøyer,1863 

= C. aeglefini Krøyer,1863 

@ Teleosts & Elasmobranchs, chiefly Gadoids 

Öresund-Kattegatt-Skagerrak-Br. Isles-Norw. Sea-Faeroes-Iceland-boreoarctic Atlantic 

C. bonito Wilson,1905 

= C. kuroshio Shiino,1959 

= C. krishnai Thomas,1967 

@ chiefly Scombridae, e.g. Sarda spp. 

Br. Isles-S Africa-Mediterranean 

C. brevicaudatus A. Scott,1901 

@ Triglidae 

Irish Sea-Barents Sea-Mediterranean-S Africa 

C. centrodonti Baird,1850 

= C. abbreviatus Krøyer,1863 

@ Pagellus bogaraveo, Labridae & Sebastes marinus 

Bergen-North Sea-Br. Isles, Black Sea 

C. coryphaenae Steenstrup & Lütken,1861 

=? C. scutatus H. Milne-Edwards,1840 

=? C. thymni Dana,1852 

= C. aliuncus Wilson,1905 

= C. elongatus Heegaard,1943, non von Nordmann,1832 

= C. tesserifer Shiino,1952 

non = C. coryphaenae : Brian,1935, nec : Yamaguti,1936, nec : Barnard,1955 

@ chiefly Scombridae & Carangidae, e.g. Thunnus 

North Sea-Tropical Seas 

C. diaphanus von Nordmann,1832 

@ Triglidae & several other teleost taxa 

Bohuslän-Norway-Br. Isles-Spain-Mediterranean 

C. elongatus von Nordmann,1832 

= C. kroeyeri H. Milne-Edwards,1840 

= C. rissoanus H. Milne-Edwards,1840 

= Caligus rapax : Baird,1850 (p.p.), non H. Milne-Edwards,1840 

= C. gurnardi Krøyer,1863 (p.p.) 

= C. lumpi Krøyer,1863 

= C. trachypteri Kollar, in Krøyer,1863 

= C. latifrons Wilson,1905 

= C. rabidus Leigh-Sharpe,1936 

= C. arcticus Brandes,1956 

non = C. elongatus Heegaard,1943 

@ many teleost (e.g. Scomber ) & elasmobranch species 

* Probably the most common parasitic copepod in British waters, sensu Kabata,1979 

Bohuslän-Norw. Sea-Iceland-Br. Isles-Ireland-southwards 

C. gurnardi Krøyer,1863 

= C. nanus Krøyer,1863 

non = C. gurnardi : Brian,1898, nec : Wilson,1908, nec : Fraser,1920 

@ Triglidae 

North Sea-Br. Isles-European Atlantic 

C. labracis T. Scott,1902 

@ Labrus bergylta & L. mixtus 

Irish Sea, S & SW England 

C. minimus Otto,1821 

= C. minutus : Aucct. 

@ Dicentrarchus, Mugil, Pagellus & Umbrina 

North Sea-Br. Isles-Spain-Mediterranean 

C. pelamydis Krøyer,1863 

= C. scomberi Bassett-Smith,1896, non C. scombri : J.V. Thompson, in W. Thompson,1856 

@ Scomber, other Scomridae & several other teleosts 

Skagerrak-North Sea-Br. Isles-S Atlantic-Mediterranean-Black Sea 

C. productus Dana,1852 

@ chiefly big Scombridae, but also other fish taxa 

Atlantic 

C. zei Norman & T. Scott,1906 

@ Usually in gill chamber of Zeus faber 

S Br. Isles-southwards 

C. pageti Russel,1925 

@ Liza aurata 

Mediterranean 

Pseudocaligus A. Scott,1901 

P. brevipedis (Bassett-Smith,1896) () 

@ usually gadid fish, but also e.g. Gobius 

Br. Isles-Bay of Biscay 

P. apodus Brian,1924 

@ Liza aurata 

Mediterranean 

Sciaenophilus van Beneden,1852 

S. tenuis van Beneden,1852 () 

= Caligus (Sciaenophilus) benedeni Bassett-Smith,1898 

@ Sciaenidae 

?Baltic-North Sea-Br. Isles-S Atlantic-Mediterranean 

Lepeophtheirus von Nordmann,1832 

L. pectoralis (Müller,1776) () 

= Lernaea pectoralis O.F. Müller,1776 

20 

@ inner surface of pectoral fins of chiefly Pleuronectidae, but also other hosts, e.g. Gadus morhua & Raja 

radiata 

Bohuslän-Norw. Sea-Faeroes-Iceland-White Sea-Br. Isles 

L. hippoglossi (Krøyer,1837) 

= Caligus hippoglossi Krøyer,1837 

= Lepeophtheirus obscurus Baird,1850 (male) 

@ Hippoglossus hippoglossus & Reinhardtius hippoglossoides; occasionally other flatfishes 

Kattegatt-Denmark-North Sea-Barents Sea-Iceland-Faeroes-Br. Isles 

L. nordmanni (H. Milne-Edwards,1840) 

= Caligus nordmanni H. Milne-Edwards,1840 

= Caligus ornatus H. Milne-Edwards,1840 

= Lepeophtheirus insignis Wilson,1908 

@ Mola mola 

North Sea, around Britain, "Europe"-Iceland-S Atlantic-Mediterranean 

L. pollachius Bassett-Smith,1896 

= L. innominatus Wilson,1905 

@ chiefly Pollachius spp. & Molva molva 

around Br. Isles, North Sea 

L. salmonis (Krøyer,1837) "Laxlus" 

= Caligus salmonis Krøyer,1837 

= Caligus vespa H. Milne-Edwards,1840 

= Caligus strömii Baird,1848 

= Caligus pacificus Gisler,1883 

= Lepeophtheirus uenoi Yamaguti,1939 

@ chiefly Salmo, Oncorhynchus & Salvelinus spp. 

* A marine copepod with capability to survive limnic water for up to two weeks during host spawning. 

Br. Isles-N Hemisphere, Norwegian Salmon farms, rivers in all nordic countries 

L. sturionis (Krøyer,1838) 

= Caligus sturionis Kr,1838 

@ Acipencer sturio & A. transmontanus 

Baltic-Denmark-Skagerrak-North Sea-Br. Isles, Pacific N America 

L. thompsoni Baird,1850 

= Caligus gracilis van Beneden,1851 

= Lepeophtheirus rhombi Krøyer,1863 

= Lepeophtheirus gibbosus Krøyer,1863 

= Lepeophtheirus gracilescens Krøyer,1863 

=? Lepeophtheirus obscurus Bassett-Smith,1896 

@Scophthalmus maximus 

W Baltic-Bohuslän-North Sea-Norw. Sea-Faeroes-Br. Isles-W Mediterranean 

L. europaeus Zeddam, Berrebi, Renaud, Raibaut & Gabrion,1988 

= L. thompsoni : Auctt. (p.p.) 

@ Scophthalmus rhombus & Platichthys flesus 

?Scandinavia, Britain 

Euryphoridae Wilson,1905 

@ Fish 

Euryphorus H. Milne-Edwards,1840 ( E. nordmanni H. Milne-Edwards,1840) 

E. brachypterus (Gerstaecker,1853) 

=? Caligeria bella Dana,1852 

= Elytrophora brachyptera Gerstaecker,1853 

= Arnaeus thynni Krøyer,1863 

= Dysgamus longifurcatus Wilson,1923 

= Elytrophora atlantica Wilson,1932 

= Elytrophora indica Shiino,1958 

= Dysgamus sagamiensis Shiino,1958 

@ big Scombridae 

E England, Engl. Channel-all oceans inhabited by "tunas" 

Trebiidae Wilson,1905 

@ Fish 

Trebius Krøyer,1838 

T. caudatus Krøyer,1838 

@ Raja spp. & some other elasmobranchs 

Skagerrak-North Sea-Norw. Sea-Br. Isles-Iceland-S Africa-Mediterranean 

Pandaridae H. Milne-Edwards,1840 

@ Fish 

Pandarus Leach,1816 

P. bicolor Leach,1816 () 

= Pandarus bosci Leach,1816 

= Pandarus fissifrons H. Milne-Edwards,1840 

= Pandarus lividus Frey & Leuckart,1847 

= Nogagus angustulus Gerstaecker,1854 

= Nogagus latus T. Scott,1907 

@ many different sharks, e.g. Squalus acanthias 

Bohuslän-Norw. Sea-Iceland-Br. Isles-southwards 

Dinemoura Latreille,1829 

D. producta (O.F. Müller,1785) () 

= Caligus productus O.F. Müller,1785 

= Dinemoura lamnae Baird,1830 

= Dinematura affinis Thomsen,1949 


Öresund-Norw. Sea-Br. Isles-southwards 

D. ferox (Krøyer,1838) 

@ sharks, e.g. Carcharias glaucus (Linnaeus) 

Faeroes-Iceland-Arctic Seas 

Echthrogaleus Steenstrup & Lütken,1861 

E. coleoptratus (Guerin-Meneville,1837) () 

= Dinemoura affinis H. Milne-Edwards,1840 

= Dinematura braccata Dana,1852 

= Nogagus lutkenii Norman,1868 

= E. perspicax Olsson,1869 


Bohuslän-Norw. Sea-Faeroes-Shetlands-Br. Isles-southwards 

Demoleus Heller,1865 

D. heptapus (Otto,1821) () 

= Caligus heptapus Otto,1821 

= Caligus paradoxus Otto,1821 

= Binoculus sexsetaceus von Nordmann,1832 

@ Hexanchus spp. 

Br. Isles-Atlantic, Mediterranean, Pacific 

Phyllothyreus Norman,1903

P. cornutus (H. Milne-Edwards,1840) 

@ Prionace glauca & other big sharks 

Br. Isles-Atlantic 

Nogaus Leach,1816 

= Nogagus : Auctt. & Nessipus : Auctt. 

* An assamblage of pandarid males, often known only from free-swimming records. 

"N." borealis Steenstrup & Lütken,1861 

= Nessipus borealis (Steenstrup & Lütken,1861) : Wilson,1907 

@ Isurus oxyrinchus - & in plankton 

Br. Isles-Atlantic-Iceland-Barents Sea 

"N." ambiguus (T. Scott,1907) 

@ Squalus acanthias 

North Sea-Plymouth 

Cecropidae Dana,1852 

@ Fish 

Cecrops Leach,1816 

C. latreillii Leach,1816 () 

@ gills of Mola mola & Mola lanceolata 

Öresund-Norw. Sea-Iceland-Br. Isles-southwards 

Orthagoriscicola Poche,1902 

O. muricatus (Krøyer,1837) () 

= Laemargus muricatus Krøyer,1837 

= Orthagoriscicola szidati Stekhoven & Stekhoven,1956 

= Orthagoriscicola wilsoni Stekhoven & Stekhoven,1956 

@ Mola mola 

Norw. Sea-Iceland-Br. Isles-Atlantic-Mediterranean 

Philorthagoriscus Horst,1897 

P. serratus (Krøyer,1863) () 

= Dinemoura serrata Krøyer,1863 

@ Mola mola 

Br. Isles-Atlantic-Mediterranean 

Luetkenia Claus,1864 

L. asterodermi Claus,1864 () 

= Cecropsina glabra Heller,1865 

@ Luvarus imperialis 

Br. Isles-Mediterranean-Atlantic 

Dichelesthiidae Dana,1853 

@ Fish 

Dichelesthium Hermann,1804 

D. oblongum (Abildgaard,1794) 

= Caligus oblongus Abildgaard,1894 

= Dichelesthium sturionis Hermann,1804 () 

@ Acipencer spp. & Huso huso 

North Sea-Br. Isles-Mediterranean-Black Sea 

Anthosoma Leach,1816 

A. crassum (Abildgaard,1794) 

= Caligus crassus Abildgaard,1794 

= Caligus imbricatus Risso,1816 

= Anthosoma smithii Leach,1816 () 

@ Half embedded in flesh of Galeoidea & Mola mola , only posyerior half partially protruding 

Öresund-Belgium-Br. Isles-Atlantic-Mediterranean 

Eudactylinidae Yamaguti,1963 

@ Gills of Fish 

Eudactylina van Beneden,1853 

E. acuta van Beneden,1853 () 

@ Squatina squatina & ?Squalus acanthias 

?Skagerrak-Br. Isles-Mediterranean-Canaries 

E. similis T. Scott,1902 

@ Raja spp. & Torpedo nobiliana 

North Sea-Br. Isles-Mediterranean 

E. acanthii A. Scott,1901 

@ Squalus acanthias & Mustelus asterias 

British Isles-N Atlantic-Angola 

E. insolens T. & A. Scott,1913 

@ Galeorhinus galeus 

Norfolk, Irish Sea 

E. minuta T. Scott,1904 

@ Dasyatis pastinaca 

Scotland (Dornoch Firth) 

Eudactylinella Wilson,1932 

E. alba Wilson,1932 ( - Woods Hole) 

@ Aetobatus narinari & Dasyatis spp. 

Falmouth Bay, Mediterranean, New England-Gulf of Mexico 

Nemesis Risso,1826 ( N. lamna Risso,1826) 

N. lamna Risso,1826 vermi A. Scott,1929 : Hewitt,1969 

@ Cetorhinus maximus 

Firth of Clyde, Concarneau, New Zealand, Japan 

N. robusta (van Beneden,1851) 

= Ergasilina robusta van Beneden,1851 

= Pagodina robusta van Beneden,1853 

= Nemesis pallida Wilson,1932 

= Nemesis aggregatus Cressey,1967 

@ a number of shark species & some few rays 

Br. Isles-Atlantic-Mediterranean 

Kroyeriidae Kabata,1979 

@ Elasmobranchs (except for the Holocephalan gill parasite Prokroyeria meridionalis ) 

Kroyeria van Beneden,1853 

K. lineata van Beneden,1853 () 

=? Lonchidium aculeatum Gerstaecker,1854 

= Kroyeria sublineata Yamaguti & Yamasu,1959 

@ several shark species 

S North Sea-Br. Isles-European Atlantic-Mediterranean, E USA, Japan 

Pseudocycnidae Wilson,1922, emend. Kabata,1979 

@ Fish 

Pseudocycnus Heller,1865 

P. appendiculatus Heller,1865 () 

= P. spinosus Pearse,1952 

= P. thunnus Brandes,1955 

@ Thunnus thynnus & other big Scombridae; some other teleosts 

Bohuslän-North Sea-Br. Isles 

Hatschekiidae Kabata,1979 

@ Gills of Teleosts 

Hatschekia Poche,1902 

H. hippoglossi (Guérin,[1831]) 

= Clavella hippoglossi Guérin,[1831] () 

@ Hippoglossus hippoglossus & Reinhardtius hippoglossoides 

Skagerrak-Br. Isles-Iceland-Barents Sea-Greenland-Europ. Atlantic 

H. cluthae (T. Scott,1902) 

= Clavella cluthae T. Scott,1902 

@ Ctenolabrus rupestris (& Labrus bergylta ) 

Br. Isles 

H. labracis (van Beneden,1871) 

= Clavella labracis van Beneden,1871 

= Cycnus Labri mixti Hesse,1879 

@ Labrus mixtus, L. bergylta & L. trimaculatus 

Br. Isles & adjacent coastal regions of Europe 

H. mulli (van Beneden,1851) 

= Clavella mulli van Beneden,1851 

= Clavella crassa Valle,1879 

@ Mullus surmuletus & related mullids 

Belgium-Br. Isles-Bay of Biscay-Mediterranean 

H. pagellibogneravei (Hesse,1879) 

= Cycnus Pagelli Bognaravei Hesse,1879 

=? Clavella sargi Valle,1882 

= H. cornigera T. Scott,1909 

@ Pagellus, Dentex & Sargus 

North Sea-Br. Isles-Senegal-Mediterranean 

H. pygmaea T. & A. Scott,1913 

@ Crenilabrus melops, Ctenolabrus rupestris & ?Labrus bergylta 

Br. Isles-Roscoff 

Congericola van Beneden,1854 

C. pallidus van Beneden,1854 () 

@ Conger conger 

S North Sea-Br. Isles-Eur. Atlantic-W Mediterranean 

Lernanthropidae Kabata,1979 

@ Fish 

Lernanthropus de Blainville,1822 ( L. musca de Blainville,1822) 

L. gisleri van Beneden,1852 

= L. Thompsoni Brian,1898 

@ Sciaenidae, Scatophagidae, Trachinotidae, Centropomidae & Polynemidae 

Belgium-Br. Isles-W Africa-Mediterranean 

L. kroyeri van Beneden,1851 

@ Dicentrarchus labrax & related teleosts 

North Sea-Br. Isles-W Africa-Mediterranean 

L. mugilis Brian,1898 

@ Liza aurata 

Mediterranean 

Pennellidae Burmeister,1835 emend. Kabata,1979 

@ Fish & Whales. They have normally a 2-host cycle. 

Lernaeocera de Blainville,1822 

L. branchialis (Linnaeus,1767) 

= Lernaea branchialis Linnaeus,1767 () 

= Lernaea gobina O.F. Müller,1776 

= Lernaeocera megacephala A. Scott,1929 

= Lernaeocera wilsoni Schuurmans Stekhoven,1936 

= L. obtusa Kabata,1957 

=? Lernaeocera caparti Machado Cruz,1959 

@ 13 intermediate hosts, 8 of them flatfish, but also e.g. Callionymus lyra, Cyclopterus lumpus, Agonus 

cataphractus & Myoxocephalus scorpius; 21 definitive hosts, of which ca half of them are Gadidae 

Kattegatt-Bohuslän-North Sea-Iceland-Faeroes-Br. Isles-Greenland-N Atlantic-Mediterranean 

L. lusci (Bassett-Smith,1896) 

= Lernaea lusci Bassett-Smith,1896 

= Lernaeocera phycidis Leigh-Sharpe,1933 

= Lernaeocera mulli Leigh-Sharpe,1935 

=? Lernaeocera longicollis Schuurmans Stekhoven,1935 

@ Solea solea is intermediate host; Trisopterus, other gadoids & several other teleosts are definitive hosts. 

Kattegatt-North Sea-Barents Sea-Br. Isles-Morocco 

L. minuta (T. Scott,1900) 

@ Pomatoschistus minutus & Myoxocephalus quadricornis (definitive hosts) 

* May be a dwarf form of L. lusci, living on small hosts. 

Baltic-North Sea-Br. Isles 

Haemobaphes Steenstrup & Lütken,1861 

H. cyclopterina (O. Fabricius,1780) 

= Lernaea cyclopterina O.F. Müller,1776 (nom. nud.) 

= Lernaea cyclopterina O. Fabricius,1780 () 

@ broad range of definitive hosts, e.g. Cottidae, Zoarcidae, Liparidae & Agonus ; intermediate host unknown 

Kattegatt-Norw. Sea-Iceland-Faeroes-Br. Isles-N Atlantic-Arctic Seas 

H. ambiguus T. Scott,1900 

@ Callionymus maculatus, C. festivus & C. belenus 

W Scotland-W Mediterranean 

Lernaeenicus Le Sueuer,1824 ( L. radiatus Le Sueur,1824) 

L. sprattae (Sowerby,1806) 

= Lernaea spratta Sowerby,1806 

= Lernaea cyclophora de Blainville,1822 

= Lernaeocera surrirensis de Blainville,1823 

= Lernaea ocularis Cuvier,1830 

= Lernaeonema monillaris H. Milne-Edwards,1840 

= Lernaeonema spratta Baird,1850 

= Lernaeonema bairdii Salter.1850 

= Lernaeonema monillaris Steenstrup & Lütken,1861 

21

= Lernaeenicus sardinae Baudouin,1904 

@ eyes of Sprattus sprattus, Sardina pilchardus (& occasionally Clupea harengus) 

Bohuslän-Oslofjord-North Sea-Norw. Sea-Br. Isles-W Mediterranean 

L. encrasicoli (Turton,1807) 

= Lernaea encrasicoli Turton,1807 

@ Sprattus sprattus, Engraulis encrasicolus, Sardina pilchardus & Alosa alosa 

Distribution like its congener above 

L. neglectus Richiardi,1877 

@ Liza aurata 

Mediterranean 

Pennella Oken,1815 ( Pennatula sagitta Linnaeus,1758 @ Histrio histrio - NW Atlantic) 

@ 1:rst intermediate host : Cephalopoda 

* Nom. cons.; ICZN Op. 1817 . 

P. filosa (Linnaeus,1758) 

= Lernaea cirrhosa La Martiniere,1798 

= Pennella orthagorisci Wright,1870 

= Pennella rubra Brian,1906 

= Pennella plumosa Linton,1905 

= Pennella germonia Leigh-Sharpe,1931 

@ Thunnus spp. and other big Scombridae & Mola spp & Xiphias gladius. 

Br. Isles-Atlantic- Fundy Bay 

P. balaenopterae Koren & Danielssen,1877 

= P. crassicornis Steenstrup & Lütken,1861 

= P. antarctica Quidor,1913 

= P. anthonyi Quidor,1913 

= P. cetti Quidor,1913 

= P. charcoti Quidor,1913 

@ Balaenoptera spp., Hyperoodon rostratus (O.F. Müller) and other big whales 

* Why Hogans 1987 used Koren & Danielssen's younger name instead of Steenstrup & Lütken's name in his 

revision of this species is unclear. 

Norw. Sea-Iceland, Faeroes 

P. instructa Wilson,1917 

@ Xiphias gladius 

NW Atlantic 

P. varians Steenstrup & Lütken,1861 

@ Eledone moschata, Todaropsis eblanae, Loligo vulgaris, Alloteuthis subulata, Sepia officinalis & S. 

elegans 

* This is probably a synonym of another species. 


Peniculus von Nordmann,1832 

P. clavatus (O.F. Müller,1776) 

@ Sebastes marinus 

Oslofjorden, North Sea-Norw. Sea-Iceland 

P. fistula von Nordmann,1832? () 

=? P. fissipes Wilson,1917 

@ Zeus faber 

Naples 

P. sp. : Schram,1991 

@ Scomber scombrus 

Celtic Sea 

Sarcotretes Jungersen,1911 

S. scopeli Jungersen,1911 

@ Benthosema glaciale 

* the copepode may be fouled by a hydroide, Ichthyocodium sarcotretis Jungersen,1911 

North Sea-W Norway-Norw. Sea-Faeroes 

Peroderma Heller,1865 

P. cylindricum Heller,1865? 

@ various organs of Sardina pilchardus 

Spain, France, Tunisia 

Sphyriidae Wilson,1919 

@ Fish 

Sphyrion Cuvier,1830 ( S. laevigatum (Quoy & Gaimard,1824)) 

S. lumpi (Krøyer,1845) 

= Lestes lumpi Krøyer,1845 

=? Sphyrion norvegicum Thor,1900 

@ Gadiformes, Scorpaeniformes, Anarhichas & Reinhardtius 

Kattegatt-N Norway-Iceland-Br. Isles-S Atlantic 

Lophoura Kölliker,1853 

L. edwardsi Kölliker,1853 () 

@ Macrouridae, chiefly Coelorinchus coelorinchus 

W Br. Isles-Angola-W Mediterranean 

Tripaphylus Richiardi,1878 

T. musteli (van Beneden,1851) 

= Lernaeonema musteli van Beneden,1851 () 

@ Mustelus spp. 

Br. Isles-Irish Sea-Eur. Atlantic-Italy 

Lernaeopodidae Olsson,1869 

@ Marine & limnic Fish 

Achtheres von Nordmann,1832 

A. percarum von Nordmann,1832 () 

= A. sandrae Gadd,1904 

= A. sibiricus Messjatzeff,1926 

@ Perca fluviatilis, Lucioperca spp. & Acerina cernua 

Brackish water at river mouths in the Baltic, Black Sea, Aral Sea, Sea of Azov & limnic waters 

Salmincola Wilson,1915 

S. salmoneus (Linnaeus,1758) 

= Lernaea salmonea Linnaeus,1758 () 

@ Salmo salar, S. gairdneri, Onkorhynchus keta & Thymallus thymallus 

Limnic-marine; Baltic-Br. Isles-"Europe" 

S. gordoni Gurney,1933 

= S. salmoneus : Kabata,1979 (p.p.) 

@ Trutta trutta 

Yorkshire, Scotland, Port Erin, Ireland, ?Orkneys 

S. extensus (Kessler,1868) 

@ Coregonus 


Baltic & limnic 

S. thymalli (Kessler,1868) 

@ Thymallus & other salmonids 

Limnic-"freshened estuarine"; Sweden, Finland, ?Britain 

Ommatokoita Leigh-Sharpe,1926 

O. elongata (Grant,1827) 

= Lernaea elongata Grant,1827 

= Lernaeopoda elongata : Auctt. 

= Ommatokoita superba Leigh-Sharpe,1926 () 

@ cornea of eyes of Somniosus microcephalus (& Lamna & Mustelus ) 

Skagerrak-off Belgium-Br. Isles-Norw. Sea-Faeroes-Iceland-Greenland-Barents Sea 

Vanbenedenia Malm,1860 

V. kroeyeri Malm,1860 () 

@ dorsal spine ofChimaera monstrosa 

Kattegatt-Skagerrak-North Sea-W Norway-Faeroes 

Charopinus Krøyer,1863 

C. dalmanni (Retzius,1829) 

= Lernaea dalmanni Retzius,1829 () 

= Stylophorus hippocephalus Hesse,1879 

@ On gills, in nasal fossae & on inner surfaces of the spiracles of Raja batis (& possibly other ray species) 

North Sea-Norw. Sea-Iceland-Barents Sea-Br. Isles-off Eur. Atl.-Mediterranean-Black Sea 

C. dubius T. Scott,1900 

@ Attached within branchial & buccal cavities of Raja batis, R. maculata, R. circularis, R. oxyrhinchus , R. 

montagui, R.naevus, R. fullonica & Laeviraja macrorhynchus 

North Sea-Br. Isles-E North Atlantic 

Pseudocharopinus Kabata,1964 ( P. dentatus Wilson,1912)) 

P. bicaudatus (Krøyer,1837) 

= Lernaeopoda bicaudata Krøyer,1837 

@ spiracles of Squalus acanthias, S. fernandinus, Trygon sp. (& possibly other elasmobranchs) 

North Sea-Br. Isles-W Africa 

P. malleus (Rudolphi, in von Nordmann,1832) 

= Brachiella malleus Rudolphi, in von Nordmann,1832 

= Brachiella lophii H. Milne-Edwards,1840 

= Brachiella pastinacae van Beneden,1851 

@ Dasyatis pastinaca, Trygon spp. (& Torpedo, Myliobatis & Rhinoptera ) 

Br. Isles-Senegal-Mediterranean 

Schistobrachia Kabata,1964 

S. ramosa (Krøyer,1863) 

= Charopinus ramosus Krøyer,1863 () 

@ Raja radiata, R. clavata, R. fullonica & R. maculata (+ ?R. batis & ?R. scabrata ) 

North Sea-Barents Sea-Iceland-Br. Isles-?Black Sea 

Lernaeopodina Wilson,1915 ( L. relata Wilson,1915) 

L. longimana (Olsson,1869) 

= Lernaeopoda longimana Olsson,1869 

= Lernaeopoda cluthae T. Scott,1900 

=? Lernaeopoda similis Scott & Scott,1913 

@ Raja fullonica, R. radiata, R. batis, R. erinacea, R. laevis, R. naevus & R. montagui 

Skagerrak-North Sea-Barents Sea-Iceland-Br. Isles 

Lernaeopoda de Blainville,1822 ( L. brogniartii de Blainville,1822) 

L. galei Krøyer,1837 

= Achtheres selachiorum Kurz,1877 

= Lernaeopoda scyllicola Leigh-Sharpe,1916 

@ Most often in cloacal region of Scylliorhinus, Galeorhinus, Mustelus (& other elasmobranchs) 

Br. Isles-Barents Sea-Iceland-S Africa-Mediterranean-Black Sea 

L. bidiscalis Kane,1892 

@ Galeorhinus (& Mustelus & Scylliorhinus ) 

North Sea-Br. Isles-Iceland-Eur. Atlantic-Mediterranean 

L. mustelicola Leigh-Sharpe,1919 (sp. inquir.) 

@ Mustelus vulgaris 

* Probably a synonyme of L. galei Krøyer 

Plymouth-?Sweden 

Albionella Kabata,1979 

A. globosa (Leigh-Sharpe,1918) 

= Lernaeopoda globosa Leigh-Sharpe,1918 () 

@ nasal fossae of Scylliorhinus caniculus 

Br. Isles 

Clavellistes Shiino,1963 ( C. shoyoae Shiino,1963) 

C. lampri (Scott & Scott,1913) 

= Lernaeopoda lampri Scott & Scott,1913 

@ Lampris guttatus 

North Sea-Faeroes-Shetlands-Atlantic 

Brachiella Cuvier,1830 

B. thynni Cuvier,1830 () 

= Thynnicola ziegleri Miculicich,1905 

@ big Scombridae, Argyrosomus regium & Pomatomus saltatrix 

Br. Isles-southwards 

Neobrachiella Kabata,1979 

N. merluccii (Bassett-Smith,1896) 

= Brachiella merluccii Bassett-Smith,1896 () 

@ Chiefly on the tips of the gill rakers of Merluccius spp. 

Br. Isles-E North Atlantic-Mediterranean 

N. bispinosa (von Nordmann,1832) 

= Brachiella bispinosa von Nordmann,1832 

= Lernaeopoda obesa Krøyer,1837 

= Anchorella ovalis : Auctt., non Krøyer,1837 

@ Triglidae 

Skagerrak-Br. Isles-W Mediterranean 

N. chevreuxii (van Beneden,1891) 

= Brachiella chevreuxii van Beneden,1891 

= Clavella sciaenae Brian,1906 

= Brachiella macrura Wilson,1920 

@ Scianidae & Lutjanidae, e.g. Argyrosomum regium 

Br. Isles-Eur. Atlantic-Mediterranean 

22

N. impudica (von Nordmann,1832) 

= Brachiella impudica von Nordmann,1832 

= Brachiella parva Nunes-Ruivo,1957 

@ Triglidae 

North Sea-Br. Isles-W Africa-Mediterranean 

N. insidiosa insidiosa (Heller,1865) 

= Brachiella insidiosa Heller,1865 

= Parabrachiella australis Wilson,1923 

= Brachiella lageniformis Szidat,1955 

@ Merluccius spp. 

Br. Isles-Eur. & Afr. Atlantic, Mediterranean 

N. rostrata (Krøyer,1837) 

= Brachiella rostrata Krøyer,1837 

@ Hippoglossus hippoglossus, Reinhardtius hippoglossoides (& Scophthalmus maximus ) 

Kattegatt-North Sea-Br. Isles-Faeroes-Iceland-Barents Sea-Greenland 

N. triglae (Claus,1860) 

= Brachiella triglae Claus,1860 

@ Triglidae 

North Sea-Br. Isles-Atl. France-Mediterranean 

Clavella Oken,1815 

* Nom. cons.; ICZN Op. 1817 . 

C. adunca (Strøm,1762) 

= Lernea adunca Strøm,1762 () 

= Lernaea cyclopterina O.F. Müller,1776 

= Lernaea uncinata O.F. Müller,1776 

= Anchorella lagenula Cuvier,1830 

= Anchorella uncinata von Nordmann,1832 

= Anchorella brevicollis H. Milne-Edwards,1840 

= Anchorella agilis Krøyer,1863 

= Anchorella quadrata Bassett-Smith,1896 

= Clavella dubia T. & A. Scott,1913 

= Clavella sciatherica Leigh-Sharpe,1918 

= Clavella iadda Leigh-Sharpe,1920 

= Clavella devastatrix Leigh-Sharpe,1925 

= Clavella invicta Leigh-Sharpe,1925 

= Clavella typica Leigh-Sharpe,1925 

= Clavella deliciosa Leigh-Sharpe.1933 

= Clavellina uncinata Dogiel,1936 

= Clavellina brevicollis Dogiel,1936 

@ chiefly Gadidae, but several other hosts known 

Öresund-Skagerrak-North Sea-Br. Isles-N Atlantic-Iceland-Faeroes-Mediterranean-Antarctic 

C. alata Brian,1906 

@ Phycis spp. 

North Sea-Br. Isles-Eur. Atlantic-Mediterranean 

C. stellata (Krøyer,1838) 

= Anchorella stellata Krøyer,1838 

@ along rims of opercula of Merluccius merluccius 

North Sea-Br. Isles-Eur. Atlantic 

C. stichaei (Krøyer,1863) 

@ Lumpenus, Stichaeus etc. 


Iceland 

Clavellodes Wilson,1915 

C. rugosa (Krøyer,1837) 

= Anchorella rugosa Krøyer,1837 () 

@ Anarhichas spp. 

North Sea-Br. Isles-Faeroes-Iceland-Greenland-Norw. Sea-Barents Sea 

Clavellisa Wilson,1915 ( C. spinosa Wilson,1915) 

C. emarginata (Krøyer,1837) 

= Anchorella emarginata Krøyer,1837 

= Anchorella ovalis Krøyer,1837 

@ Alosa spp., (Caspialosa & Clupeonella ) 

North Sea-Br. Isles-Mediterranean-Black & Azov Seas 

C. scombri (Kurz,1877) 

= Anchorella scombri Kurz,1877 

@ first gill arch of Scomber scombrus & S. colias 

Skagerrak-North Sea-Br. Isles-Mediterranean-SW Africa 

Alella Leigh-Sharpe,1925 

A. pagelli (Krøyer,1863) 

= Anchorella pagelli Krøyer,1863 () 

= Anchorella canthari Heller,1865 

= Anchorella centrodonti van Beneden,1871 

@ Sparidae, e.g. Pagellus spp. & Spondyliosoma cantharus 

North Sea-Br. Isles-Eur. Atlantic-Mediterranean 

Advena Kabata,1979 

A. paradoxa (van Beneden,1851) 

= Anchorella paradoxa van Beneden,1851 () 

@ Scomber scombrus 

Skagerrak, Belgium-Br. Isles-Ireland-Mediterranean 

Eubrachiella Wilson,1915 

E. mugilis Kabata, Raibaut & Ben Hassine,1971 

@ Liza aurata 

Mediterranean 

Herpyllobiidae Hansen,1892 

@ Polynoids 

Herpyllobius Steenstrup & Lütken,1861 

= Silenium Krøyer,1863 

H. arcticus Steenstrup & Lütken,1861 

= H. crassirostris M. Sars,18?? 

@ On elytrophores only in the anterior half of Harmothoe spp., Gattyana spp. & Austrolaenilla mollis 

Öresund-Bohuslän-Oslofjord-Arctic Seas-Iceland-Faeroes-Plymouth 

H. polynoes (Krøyer,1863) 

@ On dorsal surface of prostomium of Harmothoe spp., Gattyana cirrosa, G. amondseni, Eunoe nodosa, 

Austrolaenilla mollis, Gaudichaudius iphionelloides, Antinoe sarsi & Antinoe badia 

Öresund-Bohuslän-North Sea-N Norway-Arctic Seas-Iceland-Faeroes, NE USA, Alaska, NE Pacific 

Eurysilenium M. Sars,1870 

E. truncatum M. Sars,1870 

@ On dorsal surface of Gattyana cirrosa & Harmothoe imbricata 

Öresund-Bohuslän-Oslofjord-"Norway" 

Phallusiella Leigh-Sharpe,1926 

P. psalliota Leigh-Sharpe,1926 

@ On dorsal side of Harmothoe extenuata 

Plymouth 

P. vera Leigh-Sharpe,1926 

@ On dorsal surface of Malmgrenia castanea 

Plymouth 

Nicothoidae Dana,1852 

= Choniostomatidae Hansen,1886 

@ Crustaceans 

Rhizorhina Hansen,1892 

R. ampeliscae Hansen,1892 

@ gill sacs of Ampelisca brevicornis & A. diadema, with only egg-sacs protruding 

Kattegatt-Norway-Clyde Sea-Engl. Channel, Black Sea 

Nicothoe Audouin & Milne-Edwards,1826 

N. astaci Audouin & Milne-Edwards,1826 

@ On gills of Homarus gammarus & Palinurus vulgaris 

Bohuslän-North Sea-Br. Isles-Orkneys-Ireland-Mediterranean 

Choniomyzon Pillai,1962 

@ decapods 

* No Scandinavian species 

Hadrothoe Humes,1975 

@ decapods 


Paranicothoe Carton,1970 ( Paranicothoe cladocera Carton,1970 - Mozambique Channel) 

@ on epicarideans parasitizing penaeids 

* Warm regions of the Indopacific only. 

Pseudonicothoe Avdeev & Avdeev,1978 ( Pseudonicothoe branchialis Avdeev & Avdeev,1978) 

@ on the gills of pandalid decapods and on epicarideans parasitizing such shrimps 


Choniosphaera Connolly,1929 

C. maenadis (Bloch & Gallien,1933) 

@ Among eggs of Carcinus maenas & Maia squinado 

E England, Isle of Man, N France 

Choniostoma Hansen,1886 

C. mirabile Hansen,1886 

@ Lebbeus polaris, Eualus gaimardi & ?Spirontocaris spp. 


?N Norway-Kara Sea-Iceland-E Greenland 

C. hanseni Giard & Bonnier,1889 

@ Spirontocaris spp. 


?N Norway-Kara Sea-Iceland-W Greenland 

C. rotundatum Stock,1958 

@ Spirontocaris lilljeborgi 

Skagerrak, North Sea 

Mysidion Hansen,1897 

M. commune Hansen,1897 

@ In marsupium of Erythrops spp. & Parerythrops obesus G.O. Sars 

Norway, SW Scotland 

M. abyssorum Hansen,1897 

@ Erythrops abyssorum G.O. Sars 


Norway 

Aspidoecia Giard & Bonnier,1889 

A. normani Giard & Bonnier,1889 

@ Externally attached to the body of Erythrops spp. 

Norway, SE & SW Scotland & SW England 

Stenothocheres Hansen,1897 

S. sarsi Hansen,1897 

@ Stenothoe marina 

S or W Norway 

S. egregius Hansen,1897 

@ Metopa bruzelii (Goes) 

Greenland 

S. sp. 

= S. egregius T. Scott,1904, ? non Hansen,1897 

@ Metopa borealis 

Aberdeen 

Homoeoscelis Hansen,1897 

H. minuta Hansen,1897 

@ Diastylis lucifera (Krøyer) 

Öresund-Kattegatt, SE Scotland 

H. frigida Hansen,1923 

@ Diastilys polaris G.O. Sars 


S of Jan Mayen, deep water 

H. longipes Hansen,1923 

@ Leptostylis villosa G.O. Sars 


W of S Iceland 

H. mediterranea Hansen,1897 

@ Branchial cavity of Iphinoe trispinosa 

SW England, Mediterranean 

Sphaeronella Salensky,1868 

S. atyli Hansen,1897 

@ Marsupium of Atylus swammerdami (Milne-Edwards) 

Kattegatt, W Ireland 

S. danica Hansen,1897 

@ Marsupium ofCorophium crassicorne Bruzelius, Apherusa bispinosa & Ericthonius punctatus 

23

Kattegatt-SE Scotland, SW England, W Ireland, Engl. Channel 

S. vestita Hansen,1897 

@ Microprotopus maculatus Norman 


Denmark 

S. paradoxa Hansen,1897 

@ Marsupium of Bathyporeia spp. & Periculodes longimanus (Bate) 

Kattegatt, Netherlands, NE & SE Scotland, NE & E England, E & SW Ireland 

S. abyssi Hansen,1897 

@ Astyra abyssi Boeck 


N Norway 

S. intermedia Hansen,1897 

@ Bruzelia typica 


Norway 

S. gitanopsidis Hansen,1897 

@ Gitanopsis arctica G.O. Sars 


N Norway 

S. giardii Hansen,1897 

@ Protomedeia fasciata Krøyer 


Kattegatt 

S. longipes Hansen,1897 

@ Marsupium of Ampelisca tenuicornis Liljeborg 

Kattegatt-SE Scotland, NE England, W Ireland, Engl. Channel 

S. amphilochi Hansen,1897 

@ Paramphilochoides odontonyx & Amphilochus neapolitanus 

Denmark, Scotland 

S. dulichiae Hansen,1897 

@ Dulichia monacantha Metzger 


Denmark 

S. frontalis Hansen,1897 

@ Marsupium of Ampelisca macrocephala Liljeborg & A. tenuicornis 

Kattegatt, Clyde Sea 

S. microcephala Giard & Bonnier,1893 

@ Marsupium of Ampelisca typica Bate & A. tenuicornis Liljeborg 

Kattegatt & W Bretagne 

S. modesta Hansen,1897 

@ Eudorella emarginata (Krøyer) 


Denmark 

S. dispar Hansen,1897 

@ Marsupium of Eudorella truncatula (Bate) 

Kattegatt, SW England 

S. insignis Hansen,1897 

@ Diastylis cornuta Boeck & D. laevis Norman 


Denmark 

S. affinis Hansen,1897 

@ Ianira maculosa Leach 


Norway (Herøy) 

S. pleurogonii Hansen,1923 

@ Pleurogonium spinosissimum 


Iceland 

S. leuckartii Salensky,1868 

= S. elegantula Hansen,1897 

@ In marsupium of Cheirocratus sundevalli (Rathke), Microdeutopus gryllotalpa Costa & Aora typica 

Krøyer, A. gracilis, Lembos longipes, Apherusa bispinosa, Corophium bonelli & C. volutator 

Kattegatt-E England, Clyde Sea-Ireland-Engl. Channel-Naples 

S. irregularis Hansen,1897 

@ Metopa rubrovittata G.O. Sars 


?Denmark 

S. minuta T. Scott,1904 

= S. pikei Green,1958 

@ Periculodes longimanus & Pontocrates arenarius 

NE & SE Scotland, SW Ireland, Engl. Channel 

S. valida Green,1958 

= S. minuta var. valida T. Scott,1905 

@ Marsupium of Megamphopus cornutus Norman 

Norfolk-Firth of Forth 

S. melphidippae Hansen,1923 

@ Melphidippa borealis Boeck 


NW Iceland 

S. monothrix (Bowman & Kornicker,1967) 

@ Marsupium of Cylindroleberis mariae 

SE Scotland, NE USA 

S. aeginae Hansen,1923 

@ Aegina echinata 


between Iceland & Faeroes 

S. decorata Hansen,1897 

@ Diastylis rathkei 


Kara Sea-W Greenland-Davis Strait 

S. devosae Stock,1960 

@ Marsupium of Corophium arenarium 

Netherlands 

S. ecaudata Stock,1960 

@ Marsupium of Corophium arenarium 

Netherlands 

S. rotundata Hansen,1923 

@ Hemilamprops cristata G.O. Sars 


off SW Faeroes 

S. munnopsidis Hansen,1897 

@ Munnopsis typica M. Sars 


Kara Sea-E Greenland 

S. munnae Hansen,1923 

@ Munna spinifera Hansen 


off E Iceland, deep water 

S. nannonisci Hansen,1923 

@ Nannoniscus simplex Hansen 


off W Iceland, deep water 

S. callisomae T. Scott,1904 

@ ? 

* Inadequately described according to Gotto,1993 

Britain 

S. cluthae T. Scott,1904 

@ ? 


Britain 

S. pygmaea T. Scott,1904 

@ ? 


Britain 

S. vararensis T. Scott,1905 

@ ? 


Britain 

Melinnacheridae Bresciani & Lützen,1975 

= Saccopsidae Lützen,1964 

@ Terebellomorph Polychaets 

* Not recognized by Huys & Boxshall 1991 

Melinnacheres M. Sars,1870 

= Saccopsis Levinsen,1878 

= Oestrella McIntosh,1885 

M. terebellidis (Levinsen,1878) 

@ Terebellides stroemi 


Greenland-N America 

M. steenstrupi (Bresciani & Lützen,1961) 

@ Blood-feeding from the base of the gills of Terebellides stroemi 

Öresund-Bohuslän-SE Scotland-Iceland-E & W Greenland-E Canada, Mediterranean 

M. ergasiloides M. Sars,1870 

@ On parapodia of the last thoracic segments of Melinna cristata 

N Kattegatt-Bohuslän-Oslofjord, Massachusetts 

M. alleni (Brumpt,1987) 

= Saccopsis alleni Brumpt,1987 

@ Polycirrus aurantiacus Grube 


?France 

Asterocheridae Giesbrecht,1899 

= Ascomyzontidae Thorell,1859? 

@ Sponges, Anthozoans, Echinoderms, Ascidians 

Asterocheres Boeck,1859 

= Ascomyzon Thorell,1859 

= Echinocheres Claus,1889 

A. siphonatus Giesbrecht,1899 

= A. thorelli Bresciani & Lützen,1962 

= A. lilljeborgi (Thorell,1859), non Boeck,1859 

= Ascomyzon lilljeborgi : G.O. Sars,1918, non Asterocheres lilljeborgi Boeck,1859 

@ Corella parallelogramma, Ascidia virginia & Anseropoda membranacea 

Bohuslän-S Norway-Norw. Sea-Iceland-?Mediterranean 

A. echinicola (Norman,1868) 

= Ascomyzon parvum : G.O. Sars,1918, non Asterocheres parvus Giesbrecht,1897 

@ Haliclona oculata, Halichondria spp., Grantia compressa, Ciocalypta penicillus, ?Echinus esculentus & 

other sponges 

Bohuslän-S Norway-North Sea-Shetlands-Br. Isles, N Ireland, Engl. Channel 

A. flustrae Ivanenko & Smurov,1997 

@ Flustra foliacea 

White Sea 

A. parvus Giesbrecht,1897 

@ Clathrina lacunosa, C. primordialis & Spongelia fragilis 


A. latus (Brady,1880) 

@ Clathrina primordialis & perhaps algae 

Skagerrak, S Norway, W Britain, SW Ireland, Mediterranean 

A. boecki (Brady,1880) 

non = A. boecki : Giesbrecht,1899 (which is the Mediterranean A. complexus Stock,1960) 

@ Halichondria panicea, Haliclona oculata 

SW & W Norway-Spitsbergen-Iceland-Br. Isles-W & SW Ireland 

A. lilljeborgi Boeck,1859 

= Ascomyzon asterocheres G.O. Sars,1918 

@ Henricia, Asterias, Crossaster, Luidia, Antedon etc. 

Bohuslän-S & W Norway-North Sea-Britain-W Ireland-Engl. Channel-Davis Strait 

A. violaceus (Claus,1889) 

@ regular & irregul. Echinoidea, Ophiuroids, Asteroids & Antedon 

Bohuslän-North Sea-W Norway-Shetlands-Br. Isles-France-Mediterranean 

A. suberitis Giesbrecht,1897 

@ Suberites domuncula 

SW England, ?SW Scotland, Mediterranean 

A. stimulans Giesbrecht,1897 

@ ? 


A. simulans T. Scott,1894 

@ sponges 

S Norway, SW Scotland, SW Ireland 

24

A. ellisi Hamond,1968 

@ Halichondria panicea, Ciocalypta penicillus 

Norfolk 

A. renaudi Canu,1891 

@ Reniera sp. 

* Doubtful if this is placed in correct genus according to Gotto,1993 

N France 

A. simplex ??,1??? 

@ Clathrina clathrus & Clathrina lacunosa 


Banyuls-sur-mer 

A. mucronipes ??,1??? 

@ Oscarella lobularis 



A. corneliae ??,1??? 

@ Clathrina primordialis 



Dermatomyzon Claus,1889 

D. nigripes (Brady & Robertson,1876) 

@ ? (so far found together with Laminaria - or free) 

Bohuslän-N Norway-Arctic Seas-Faeroes-Shetlands-Br. Isles-NE & SW Ireland, N France, Mediterranean 

Rhynchomyzon Giesbrecht,1895 

R. purpurocinctum (Scott,1893) 

@ ? 

Aalesund-Christiansund-Br. Isles-off Irish coast, Mediterranean 

R. rubrovittatum G.O. Sars,1918 

@ ? 


W Norway (Kallevaag) 

R. falco Giesbrecht,1895 

@ Clathrina contorta 


S Norway-Mediterranean 

Acontiophorus Brady,1880 

A. scutatus (Brady & Robertson,1873) 

@ Spongelia fragilis var. ramosa 

W Norway, Faeroes, Ireland, Britain, N France, Mediterranean 

A. ornatus (Brady & Robertson,1876) 

= A. armatus Brady,1880 

@ ? 

S Norway (Risør)-NE England, Davis Strait, Mediterranean 

A. antennatus Hansen,1923 

@ Clathrina lacunosa & Oscarella lobularis 

around Greenland-?Mediterranean 

Orecturus Humes,1992 ( O. grandisetiger Humes,1992 - Madagascar @ Antipathes ) 

Collocheres Canu,1893 

= Leptomyzon G.O. Sars,1915 

C. gracilicauda (Brady,1880) 

@ Ophiothrix fragilis , (Antedon spp. & Echinus esculentus ) 

Bohuslän-S Norway-Br. Isles-NE & SW Ireland-Roscoff-Mediterranean 

C. gracilipes Stock,1966 

= C. gracilicauda : Canu,1893, : G.O. Sars,1915, non (Brady,1880) 

@ ? (possibly a crinoid) 

S Norway (Risør), Boulogne 

C. elegans A. Scott,1896 

@ arm base of Ophiocomina nigra 

W Norway-W Br. Isles 

Scottomyzon Giesbrecht,1897 

S. gibberum (T. & A. Scott,1894) 

@ making galls in rosettes of pedicellariae in Asterias rubens, Marthasterias glacialis & Spatangus purpureus 

Belt Sea-Bohuslän-S & SW Norway-North Sea-Br. Isles-N France 

Mesocheres Norman & Scott,1905 

M. anglicus Norman & Scott,1905 

@ ? 

Oslofjord-SW England 

Scottocheres Giesbrecht,1897 

S. elongatus (T. & A. Scott,1894) 

@ Dysidea tupha, Ulosa stuposa, Sigmatoxella annexa, Halichondria sp. & Haliclona sp. 

W Norway-North Sea-Br. Isles-SW Ireland-N France-Mediterranean 

Brychiopontiidae Humes,1974 

Brychiopontius Humes,1974 

B. falcatus Humes,1974 

@ Oneirophanta mutabilis Theel 

off W Ireland in ca 4430m 

Myzopontiidae G.O. Sars,1915 

@ ?Sponges 

* Not recognized in Huys & Boxshall 1991 

Myzopontius Giesbrecht,1895 

M. pungens Giesbrecht,1895 

@ Clathrina spp., especially C. contorta ?& ?Stichopus regalis (Cuvier) 

S Norway (Korshavn)-Arctic Seas-Mediterranean 

Neopontius T. Scott,1898 

N. angularis T. Scott,1898 

@ ? 

W Norway-SW Scotland 

Dyspontiidae Giesbrecht,1895 

@ sponges, scleractinians, ascidians 

* Not represented as a family of its own in Gotto,1993, but included in Artotrogidae Brady,1880 

* Not recognized in Huys & Boxshall 1991 

Dyspontius Thorell,1859 

D. striatus Thorell,1859 

@ "small ascidians" 

?Öresund-Bohuslän-N Norway-Br. Isles-Ireland, N France, Mediterranean 

D. curticaudatus T. Scott,1905 

@ ? 

SE Scotland 

Cryptopontius Giesbrecht,1895 

C. brevifurcatus (Giesbrecht,1895) 

@ Echinus esculentus 

S Norway (Korshavn)-Mediterranean 

Arctopontius G.O. Sars,1915 


A. expansus G.O. Sars,1915 

@ ? 

N Norway (Hammerfest)-Davis Strait 

Bradypontius Giesbrecht,1895 

B. magniceps (Brady,1880) 

@ ?, possibly sponges 

Aalesund-Hammerfest-Br. Isles, Ireland, N France 

B. papillatus (T. Scott,1888) 

@ probably Pomatoceros triqueter 

Kieler Bucht-Öresund-Bohuslän-W Norway-Br. Isles-Mediterranean 

B. major G.O. Sars,1918 

@ ? 


Aalesund-Trondheim 

B. caudatus G.O. Sars,1918 

@ ? 


NW Norway (Nordland) 

B. unidens Hansen,1923 

@ ? 


E Greenland 

Cribropontius Giesbrecht,1899 

C. normani (Brady & Robertson,1876) 

@ ? 

W Norway-Br. Isles-Madeira 

Metapontius Hansen,1923 


M. latispinis Hansen,1923 

@ ? 

Faeroes, deep water 

Artotrogidae Brady,1880 

@ Nudibranchs 

Artotrogus Boeck,1859 

A. orbicularis Boeck,1859 

@ Nudibranchs ("Doridae") 

Kieler Bucht, Bohuslän-S, W & N Norway-Kara Sea-Br. Isles-E Ireland 

Undescribebed family with presumed Artotrogid affinities 

Apodomyzon Stock,1970 

A. brevicorne Stock,1970 

@ In tissue of Haliclona indistincta (Bowerbank) 

Roscoff 

A. longicorne Stock,1970 

@ In tissue of Haliclona indistincta (Bowerbank) 

Roscoff 

Cancerillidae Giesbrecht,1897 

@ Ophiuroids 

Parartotrogus T. & A. Scott,1893 

P. arcticus T. Scott,1901 

@ ? 


NW Norway (Nordland)-off E Greenland-Arctic Seas 

P. richardi T. & A. Scott,1893 

@ Ophiura albida & ?O. ophiura ("Ophioglypha lacertosa ") 

W Norway-SE &SW Scotland-Isle of Man-Mediterranean 

Cancerilla Dalyell,1851 

C. tubulata Dalyell,1851 () 

= Caligidum vagabundum Claus,1889 

=?? Phtiriopsis emilii Giard,1879 

@ Generally on oral disk surface, near arms, of Axiognathus squamatus, Ophiocomina nigra, ?Ophiothrix 

fragilis & Ophiosila aranea 

S Baltic-Bohuslän-S Norway-Trondheim-Barents Sea-Br. Isles-N & W France-Mediterranean 

Micropontiidae Gooding,1957 

@ Irregular Echinoids 

Micropontius Gooding,1957 

M. ovoides Gooding,1957 

@ Spatangus purpureus , Echinocardium spp. & Brissopsis lyrifera 

Bohuslän-Br. Isles-N &W France-W Mediterranean 

Nanaspididae Humes & Cressey,1959 

= Allantogynidae Changeux 

@ Holothurians 

Nanaspis Humes & Cressey,1959 

N. ninae Bresciani & Lützen,1962 

@ On surface of Parastichopus tremulus 

Bohuslän-Trondheim, off W Ireland in ≈480m 

25

Allantogynus Changeux,1958 


Xenocoelomatidae Bresciani & Lützen,1966 

Aphanodomus Wilson,1920 

= Crypsidomus Levinsen,1878, non Guenther,1864 

A. terebellae (Levinsen,1878) 

@ Thelepus cincinnatus & other Terebellidae 


Iceland-Greenland-Kara Sea 

Xenocoeloma Caullery & Mesnil,1915 

X. alleni (Brumpt,1897) 

@ Polycirrus caliendrum Claparède 

Engl. Channel 

X. brumpti Caullery & Mesnil,1915 

* May be conspecific with X. alleni (Brumpt); considered to be so by Gotto,1993 with some doubt. 

@ Polycirrus arenivorus Caullery 

?Engl. Channel 

Spongiocnizontidae Stock & Kleeton,1964 

Spongiocnizon Stock & Kleeton,1964 

S. petiti Stock & Kleeton,1964 () 

@ Hemimycale columella 

French Mediterranian 

S. vermiformis Stock,1967 

@ Iophon hyndmani var. funis 

French Mediterranian 

Sponginticolidae Topsent,1928 

* May possibly be related to fam. Spongiocnizontidae. 

Sponginticola Topsent,1928 

= Clionophilus Silén,1963 

S. uncifer Topsent,1928 ( - French Channel coast) 

= Clionophilus vermicularis Silén,1963 ( of Clionophilus Silén,1963 - Bohuslän) 

@ Cliona celata , Stylopus coriaceus & Mycale macilenta 

Bohuslän- NE Ireland - Engl. Channel, Mediterranean 

Calverocheridae Stock,1968 

@ Echinoida 

Dinopontiidae Murnane,1967 

@ Porifera & Actiniaria 

Dirivultidae Humes & Dojiri,1980 

@ vestimentiferans 

Dissonidae Yamaguti,1963 

Entomolepidae Brady,1899 

@ Porifera 

Hyponeoidae Heegaard,1962 

Megapontiidae Heptner,1968 

@ not known 

Naobranchiidae Yamaguti,1939 

* not recognized by Huys & Boxshall 1991. 

Pontoeciellidae Giesbrecht,1895 

Rataniidae Giesbrecht,1897 

Ratania Giesbrecht,1892 

R. atlantica Farran,1926 

off Ireland, Biscay 

R. flava Giesbrecht,1892 

deep Atlantic & Mediterranean 

Stellicomitidae Humes & Cressey,1958 

Tanypleuridae Kabata,1969 

?Poecilo-/?Siphonostomatoida not yet referred to any Familiy? 

Gonophysema Bresciani & Lützen,1960 

* Possibly another sp. in Heterostigma reptans Monniot & Monniot 

G. gullmarensis Bresciani & Lützen,1960 

@ Ascidiella aspersa, A. scabra & ?Distomus variolosus 

Kattegatt-Bohuslän, ?Mediterranean 

Axinophilus Bresciani & Ockelmann,1966 

* May be related to Myicolidae? 

A. thyasirae Bresciani & Ockelmann,1966 

@ Thyasira flexuosa & T. sarsi (& in T. gouldi from Spitsbergen & W Greenland, if this is not another 

species) 

Öresund-Bohuslän-S Norway-Bergen-SE Scotland 

Phyllocola Delamare-Deboutteville & Laubier,1960 

P. petiti Delamare-Deboutteville & Laubier,1960 

@ Eulalia expusilla & other phyllodocidae 

Britain? 

MONSTRILLOIDA G.O. Sars,1903 

Monstrillidae Giesbrecht,1892 

@ not known for most species; Polychaeta & Mollusca 

Monstrilla Dana,1849 ( M. viridis Dana,1849) (publ. September 4) (n. cons. ICZN Op. 1869) 

= Thaumatoessa Krøyer, in Gaimard,1842-1845 

* Grygier, 1993 has submitted an application to ICZN to suppress this older name for purposes of the 

Principle of Priority. 

= Thaumaleus Krøyer,1849 (publ. December 31) (n. cons. ICZN Op. 1869) 

M. longicornis I.C. Thompson,1890 

=? Thaumatoessa typica Krøyer, in Gaimard,1842-45 

26 

= M. intermedia Aurivillius,1898, ?non Kriczagin,1877 (Kriczagins name is supressed for Law of Priority by 

ICZN Op. 1175) 

=? M. clavata G.O. Sars,1921 

Skagerrak-S & W Norway-Br. Isles-Mediterranean, Maine 

M. longiremis Giesbrecht,1892 

Hvaler-Oslofjord-NW Norway-North Sea-Cornwall-Mediterranean, Maine 

M. conjunctiva Giesbrecht,1902 

= M. leucopis G.O. Sars,1921 

NW Norway-North Sea-Cornwall 

M. gracilicauda Giesbrecht,1892 

* Male unknown; may be M. anglica Lubbock 

NW Norway-Arctic-Br. Isles-Engl. Channel-Mediterranean 

M. anglica Lubbock,1857 

* Female unknown; may be M. gracilicauda Giesbrecht 

Br. Isles-Engl. Channel, Atlantic Canada, "almost world-wide" 

M. helgolandica Claus,1863 

= M. canadensis McMurrich,1917 (male) 

@ Brachystomia rissoides (Hanley) & Odostomia scalaris MacGillivray 

Skagerrak-North Sea-Norway (Stavanger, Christiansund & Nordland)-Cornwall-Mediterranean-Hudson 

Strait, Maine 

M. serricornis G.O. Sars,1921 

North Sea, Norw. Sea 

M. minuta Isaac,1974 

Isle of Man 

M. obesa Isaac,1975 (nomen nudum) 

Channel Islands 

Cymbasoma J.C. Thompson,1888 

= Thaumaleus : Auctt., non Krøyer,1849 

@ Annelida 

C. rigidus J.C. Thompson,1888 () 

= Thaumaleus germanicus Timm,1896 

@ Salmacina dysteri (Huxley), Polydora ciliata & P. giardi Mesnil 

S & W Norway-E North Sea-Br. Isles-Engl. Channel-Mediterranean 

C. thompsoni (Giesbrecht,1892) 

= Thaumaleus thompsoni Giesbrecht,1892 

Belt Sea-S & W Norway-Engl. Channel-"more or less world-wide" 

C. longispinosus (Bourne,1890) 

= Monstrilla longispinosa Bourne,1890 

Norway-German Bight-North Sea-Engl. Channel-"more or less world-wide" 

C. claparedi (Giesbrecht,1892) 

= Thaumaleus claparedi Giesbrecht,1892 

North Sea-Mediterranean 

C. reticulatus (Giesbrecht,1892) 

= Thaumaleus reticulatus Giesbrecht,1892 


C. filogranarum (Malaquin,1896) 

= Thaumaleus filogranarum Malaquin,1896 

* Should perhaps be classified as a Monstrilla Dana? 

@ Filograna implexa Berkely 

Engl. Channel 

C. roscovita (Malaquin,1901) 

= Haemocera roscovita Malaquin,1901 

= Thaumaleus roscovita : Auctt. 

* Should perhaps be classified as a Monstrilla Dana? 

@ Filograna setosa Langerhans 

British waters-?Roscoff 

C. frondipes (T. Scott,1904) 

* Should the author be written within parenthesis? 

E Br. Isles 

C. rostratus (T. Scott,1904) 

= Thaumaleus rostratus T. Scott,1904 

* Male unknown; may be Thaumaleus similirostratus Isaac 

Shetlands 

C. similirostratus (Isaac,1974) 

= Thaumaleus similirostratus Isaac,1974 

* Female unknown; may be T. rostratus T. Scott 

Cornwall 

C. zetlandicus (T. Scott,1904) 

= Thaumaleus zetlandicus T. Scott,1904 

Shetlands, Engl. Channel 

C. malaquini Caullery & Mesnil,1914 

= Thaumaleus malaquini : Auctt. 

@ Syllis gracilis Grube 

Engl. Channel 

C. pallidus (Isaac,1974) 

= Thaumaleus pallidus Isaac,1974 

Cornwall 

C. tenuis (Isaac,1974) 

= Thaumaleus tenuis Isaac,1974 

Cornwall 

C. striatus (Isaac,1974) 

= Thaumaleus striatus Isaac,1974 

Cornwall-Channel Islands 

Monstrillopsis G.O. Sars,1921 

M. dubia (T. Scott,1904) 

Norway (Raunefjorden & outside Trondheimsfjord)-Scottland-Cornwall-Mediterranean, Maine 

M. sarsi Isaac,1974 

Cornwall-Bristol Channel 

M. gracilis (Gurney,1927) 

Bristol & Engl. Channels-Portugal-Suez-Red Sea 

Strilloma Isaac,1975 (nomen nudum) 

S. grandis (Giesbrecht,1891) 

= Monstrilla grandis Giesbrecht,1891 

German Bight-North Sea-Engl. Channel-Cornwall-Norfolk, China 

S. scotti Isaac,1975 

= Monstrilla grandis : T. Scott,1904, non Giesbrecht,1891

Clyde Sea 

S. lata (Desai & Bal,1963) 

Bombay, Firth of Clay 

Thaumatohessia Giard,1900 

T. armoricana (Hesse,1868) 

Brest 

MISOPHRIOIDA Gurney,1933 

Misophriidae Brady,1878 

Misophria Boeck,1865 

M. pallida Boeck,1865 

W Norway-Arctic Seas-Br. Isles-W Mediterranean 

Arcticomisophria Martínez Arbizu & Seifried,1996 

A. bathylaptevensis Martínez Arbizu & Seifried,1996 ( - deep Laptev Sea) 

Laptev Sea in 2330-3237 m 

Expansophria Boxshall & Iliffe,1987 

E dimorpa Boxshall & Iliffe,1987 () 

Canaries 

Palpophria Boxshall & Iliffe,1987 

P. aestetha Boxshall & Iliffe,1987 () 

Canaries 

Dimisophria Boxshall & Iliffe,1987 

D. cavernicola Boxshall & Iliffe,1987 () 

Canaries 

Benthomisophria G.O. Sars,1909 

B. palliata G.O. Sars,1??? 

?Mediterranean 

B. sp? 

* One bathypelacic N. Atlantic species described by Hulsemann & Grice,1964 

MORMONILLOIDA Boxshall,1979 

Mormonillidae Giesbrecht,1892 

Mormonilla Giesbrecht,1891 

* males of the two species below described by Huys, Boxshall & Bottger-Schnack 1992 

M. phasma Giesbrecht,1891 

off W Ireland-Bay of Biscay-W Mediterranean 

M. minor Giesbrecht,1891 

= M. polaris G.O. Sars,1900 

= M. atlantica Wolfenden,1905 

temperate Atlantic-Bay of Biscay-off W Ireland-Arctic Seas 

HARPACTICOIDA G.O. Sars,1903 

Superornatiremidae Huys,199? 

POLYARTHRA Lang,1948 

Longipediidae Brady,1880 

Longipedia Claus,1863 

L. coronata Claus,1863 

Bohuslän-S North Sea-Frierfjord/Langesundsfjord-Trondheim-Br. Isles-Mediterranean 

L. minor T. & A. Scott,1893 

= L. pontica : Auctt., ?non Kriczagin,1873 

Bohuslän-S North Sea-W Norway-Trondheim-Br. Isles-Mediterranean 

L. helgolandica Klie,1949 

= L. minor helgolandica Klie,1949 

Helgoland, Norfolk 

L. scotti G.O. Sars,1903 

= L. australica Nicholls,1941 (according to Wells 1980) 

Bohuslän-S North Sea-Trondheim-Br. Isles, Australia 

L. rosea G.O. Sars,1903 (nom. dub. according to Wells 1980) 

* Not included in Huys & al. 1996 

Bergen-Trondheim-?Mediterranean 

L. weberi A. Scott,1909 

* Not included in Huys & al. 1996, but redescribed by Wells 1980 and Itô 1980 

Norfolk-S Ireland-Roscoff-Mediterranean, Antarctic 

Canuellidae Lang,1944 

Sunaristes Hesse,1867 

S. paguri Hesse,1867 () 

@ Usually in shell inhabited by Pagurus bernhardus, Clibanarius erythropus & Diogenes pugilator 

?Öresund-Bohuslän-S Norway-German Bight-Br. Isles-W Ireland-Engl. Channel, Mediterranean, Black Sea 

Canuella T. & A. Scott,1893 

C. perplexa T. & A. Scott,1893 () 

@ Pagurus bernhardus & Maia squinado - or often found free. 

Bohuslän-S Norway-S North Sea-Br. Isles-Engl. Channel-Mediterranean 

C. furcigera G.O. Sars,1903 

Bohuslän-Oslofjord-S Norway-German Bight-Iceland-Greenland-Mediterranean 

C. sp. : Huys & al. 1996 

Belgium 

Canuellopsis Lang,1936 

C. swedmarki Por,1964 

off N Koster (in 150m)-Skagerrak-Bergen 

Brianola Monard,1926 

B. stebleri Monard,1926 

?Roscoff, W Mediterranean 

B. sp. : Huys & al. 1996 

Cornwall 

Microcanuella Mielke,1994 

Ellucana Sewell,1940 

Parasunaristes Fiers,1982 

Intersunaristes Huys,1995 

Galapacanuella Mielke,1979 

Canuellina Gurney,1927 

Nathaniella Por,1984 

Ifanella Vervoort,1964 

Intercanuella Becker & Schriever,1979 

Coullana Por,1984 

Scottolana Por,1964 

Elanella Por,1984 

Echinosunaristes Huys,1995 

OLIGOARTHRA Lang,1948 

* not a monophyletic taxon 

Cancrincolidae Fiers,1990 

@ gill chambers of aquatic Decapoda & land crabs 

Cancrincola Wilson,1913 (C. jamaicensis Wilson,1913) 

Abscondicola Fiers,1990 ( A. humesi Fiers,1990 - New Guinea @ land crab) 

Hamondiidae Huys,1990 

Hamondia Huys,1990 ( H. superba Huys,1990 - Australia @ sponge) 

MAXILLIPEDASPHAELEA Lang,1948 

Cervinioidea G.O. Sars,1903 

Aegisthidae Giesbrecht,1892 

Aegisthus Giesbrecht,1891 

A. mucronatus Giesbrecht,1891 

= Aegisthus atlanticus Wolfenden,1902 

* "The only harpacticoid species for which bioluminescensce has been demonstrated with confidence" 

according to Huys & Böttger-Schnack,1994 

Faeroe Channel-off Ireland-Bay of Biscay-"Cosmopolitan, Planktonic" 

A. spinulosus Farran,1905 

off W Ireland, pelagic 

A. aculeatus Giesbrecht,1891 

Biscay 

Cerviniidae G.O. Sars,1903, emend. Lang,1948 

* A deep water species family. 

Cerviniella Smirnov,1946 

Paracerviniella Brodskaya,1963 

Expansicervinia Montagna,1981 

Pontostratiotes Brady,1883 

Tonpostratiotes Itô,1982 

Stratiopontotes Soyer,1970 

Herdmaniopsis Brodskaya,1963 

= Ameliotes Por,1969 (according to Itô 1982) 

Cerviniinae Brodskaya,1963 

Eucanuella T. Scott,1900 

E. spinifera T. Scott,1900 () 

Bohuslän-W Norway-Iceland-Shetlands-Scotland 

Cervinia Brady,1878 ex Norman MS 

= Pseudocervinia Brodskaya,1963 (according to Montagna 1981) 

C. bradyi Brady,1878 ex Norman MS () 

Bohuslän-W Norway-Br. Isles-Mediterranean 

C. synarthra G.O. Sars,1910 

Bohuslän-Lofoten-Arctic Seas-Iceland-Br. Isles 

C. pilosa Lang,1948 

* Not mentioned in Huys & al. 1996 

Bohuslän 

Cerviniopsinae Brodskaya,1963 

Hemicervinia Lang,1935 

= Herdmania I.C. Thompson,1893, non Lahille,1888 

H. stylifera (I.C. Thompson,1893) 

= H. ryforsi Lang,1935 () 

Bohuslän-Isle of Man 

Cerviniopsis G.O. Sars,1903 

C. clavicornis G.O. Sars,1903 

Bohuslän-Lofoten 

C. longicaudata G.O. Sars,1903 

W Norway 

Rotundiclipeidae Huys,1988 

Rotundiclipeus ( R. canariensis Huys,1988 - Canaries in anchialine caves) 

Ectinosomatoidea G.O. Sars,1903 

27

Ectinosomatidae G.O. Sars,1903 (p.p.); Olofsson,1917 

Tetanopsis Brady,1910 

* Doubtful genus. 

Oikopus Wells,1967 

Ectinosomoides Nicholls,1945 

Bradyellopsis Brian,1924 

Peltobradya Médioni & Soyer,1967 

Sigmatidium Giesbrecht,1881 

* According to Huys & al. 1996 are several unnamed species present in the area. 

S. difficile Giesbrecht,1881 

Kieler Bucht-Elbe Mouth-Aberdeen 

Pseudectinosoma Kunz,1935 

P. minor Kunz,1935 

= Sigmatidium minor : Auctt. 

* Brackish marsh species. 

Kieler Bucht-Hiddensee 

Pseudobradya G.O. Sars,1904 

P. pulchella G.O. Sars,1920 

Öresund-Bohuslän-S Norway 

P. elegans (T. & A. Scott,1894) 

Bohuslän-W Norway-Scotland 

P. minor (T. & A. Scott,1894) 

Belt Sea-Helgoland-Trondheim-Arctic Seas-Br. Isles-?Roscoff 

P. similis (T. & A. Scott,1894) 

Bohuslän-Trondheim-Br. Isles-W France 

P. hirsuta (T. & A. Scott,1894) 

Bohuslän-SW Norway-Br. Isles 

P. fusca (T. & A. Scott,1894) 

SW Norway-Br. Isles 

P. acuta G.O. Sars,1904 

off Gullmarn, Trondheim-?Italy 

P. robusta G.O. Sars,1910 

SW Norway 

P. digitata G.O. Sars,1920 

S Norway 

P. leptognatha G.O. Sars,1920 

Bohuslän-S Norway 

P. scabriuscula G.O. Sars,1920 

S Norway 

P. attenuata G.O. Sars,1920 

Hållö, off N Koster, S Norway 

P. tenella G.O. Sars,1920 

S Norway 

P. parvula G.O. Sars,1920 

S Norway 

P. exilis G.O. Sars,1920 

Trondheim 

P. pygmaea G.O. Sars,1920 

SW Norway 

P. ambigua G.O. Sars,1920 

S Norway-Trondheim 

P. maxima Lang,1935 

Skagerrak 

P. beduina Monard,1935 

Elbe mouth-Bretagne-SW Mediterranean 

P. distinctum Wells,1968 

Scilly Islands 

P. rhea Bodin,1977 

W France 

Bradya Boeck,1872 

B. (Bradya) typica Boeck,1872 

?Kieler Bucht-Öresund-Bohuslän-Trondheim-Arctic Seas-Br. Isles 

B. (Bradya) scotti G.O. Sars,1920 

Bohuslän-Bergen-Br. Isles 

B. (Bradya) macrochaeta G.O. Sars,1920 

Bohuslän-Bergen 

B. (Bradya) furcata G.O. Sars,1920 


B. (Bradya) congenera G.O. Sars,1920 

Bohuslän-S Norway, Isle of Man 

B. (Bradya) simulans G.O. Sars,1920 

S Norway, Britain 

B. (Bradya) proxima T. Scott,1912 

North Sea, South Georgia 

B. (Parabradya Lang,1948) dilatata G.O. Sars,1904 

Bohuslän-Bergen-Iceland 

B.(Parabradya) atlantica Bodin,1968 

Biscay 

Ectinosomella G.O. Sars,1910 

E. nitidula G.O. Sars,1910 

* Pelagic? in 55-91 m 

SW Norway 

Microsetella Brady & Robertson,1873 

M. norvegica (Boeck,1865) 

= Microsetella atlantica Brady & Robertson,1873 

Pelagic around all European coasts, fr. Arctic Seas-Mediterranean, e.g. S Baltic & Bergen 

M. rosea (Dana,1848) 

Pelagic fr. Kattegatt-North Sea-Br. Isles-Bay of Biscay-Mediterranean 

Halophytophilus Brian,1918 (& 1917 (nom. nudum)) 

= Halophytophilus Brian,1921 

H. spinicornis (G.O. Sars,1920) 

Bohuslän-SE Norway 

H. similis (Lang,1948) 

Bohuslän, Scotland 

Kliesoma Hincks & Schriever,1985 

= Kliella Hincks & Schriever,1983, non Schäfer,1945 

K. triarticulatus (Klie,1949) 

= Halophytophilus? triarticulatus Klie,1949 

Helgoland, Britain 

Arenosetella C.B. Wilson,1932 

= Pararenosetella Lang,1948 (p.p.) 

A. germanica germanica Kunz,1937 

SW Baltic-German Bight-Aberdeen-Scilly Isles-Pontomediterranean 

A. tenuissima (Klie,1929) 

= Ectinosoma tenuissima Klie,1929 

= Hastigerella palpilabra Nicholls,1935 

= Arenosetella monensis Moore,1976 

Baltic-Sylt-North Sea-Brittany-Br. Isles 

A. littoralis Bodin,1977 

W France 

A. duriensis Galhano,1970 

Atl. Portugal 

A. smithi (Perkins,1956) 

= Tetanopsis Brady,1910 smithi Perkins,1956 

* Specific validity questionable according to Huys & al. 1996 

England (Kent) 

A. medius (Perkins,1956) 

= Tetanopsis Brady,1910 medius Perkins,1956 

* Specific validity questionable according to Huys & al. 1996 

England (Kent) 

Hastigerella Nicholls,1935 

= Pararenosetella Lang,1948 (p.p.) 

H. abotti abotti Lang,1965 

?Irish Sea-W Mediterranean 

H. leptoderma (Klie,1929) 

Kieler Bucht-German Bight-Br. Isles-Portugal-W Mediterranean 

H. psammae (Noodt,1955) 

= Pararenosetella psammae Noodt,1955 

W France-Scotland 

H. scheibeli Mielke,1975 

Sylt 

H. unisetosa (Wells,1965) 

= Arenosetella unisetosa Wells,1965 

Scotland 

H. grandimandibularis Wells,1967 

?Atlantic Europe, Mozambique 

H. bozici Soyer,1974 

Finistère, French Mediterranean 

Halectinosoma Lang,1944 

H. erythrops (Brady & Robertson,1875) 

= Ectinosoma gracile : G.O. Sars,1904, non T. & A. Scott,1894 

W Norway-S North Sea-Br. Isles & Ireland-N France 

H. gracile (T. & A. Scott,1894) 

Br. Isles & Ireland 

H. longicorne (T. & A. Scott,1894) 

Scotland, Roscoff 

H. tenuireme (T. & A. Scott,1894) 

= Pararenosetella tenuiremis (T. & A. Scott,1894) 

off Gullmarn, Hållö, Norway-Helgoland-Scotland-Scilly Isles-Roscoff 

H. oblongum (Kunz,1949) 

Helgoland 

H. sarsi (Boeck,1872) 

* Sp. incertae sedis according to Clement & Moore,1995 

Bohuslän-German Bight-N Norway-Arctic Seas-Br. Isles-N France 

H. chrystalli (T. & A. Scott,1894) 

= H. propinquum (T. & A. Scott,1894) 

Bohuslän-N Norway-Arctic Seas-Scotland & Ireland 

H. herdmani (T. & A. Scott,1896) 

* First intermediate host for Acanthogyrys (Acanthosentis ) lizae Orecchia, Paggi & Radujkovic,1988 

(Acanthocephala) 

Kieler Bucht-Bohuslän-Amrum-Trondheim-Br. Isles-Mediterranean & Black Sea 

H. armiferum (T. & A. Scott,1894) 

Bohuslän-Norway-Scotland 

H. neglectum (G.O. Sars,1904) 

Bohuslän-S & N Norway-Arctic Seas-?Faeroes-Iceland & Greenland; ?Mediterranean 

H. elongatum (G.O. Sars,1904) 

* Possibly a synon. of H. herdmani (T. & A. Scott) sensu Por,1964 

Kieler Bucht-Bohuslän-S Norway-S North Sea-Trondheim 

H. clavatum (G.O. Sars,1920) 

= Ectinosoma clavatum G.O. Sars,1920 

off Gullmarn, Kosterfjord, S NorwayArctic Seas 

H. brunneum (Brady,1905) 

Northumberland 

28

H. proximum (G.O. Sars,1919) 

Öresund-Bohuslän-Trondheim 

H. angulifrons (G.O. Sars,1919) 

S Norway 

H. tenerum (G.O. Sars,1920) 

Bohuslän-S Norway-Scilly Isles 

H. curticorne (Boeck,1872) 

Finland & Askö-Bohuslän-S North Sea-S & N Norway-Arctic Seas-Br. Isles-W Mediterranean 

H. gothiceps (Giesbrecht,1881) 

= Ectinosoma pygmaeum T. & A. Scott,1894 

Kieler Bucht-Bohuslän-German Bight-S Norway-Trondheim-Br. Isles-Roscoff 

H. mixtum (G.O. Sars,1904) 

Belt Sea-Bohuslän-Oslofjord; ?Mediterranean 

H. brevirostre (G.O. Sars,1904) 

Bohuslän-S Norway, Scotland 

H. abrau (Kricagin,1877) 

= Bradya edwardsi Richard,1890 

Freshwater species that tolerates brackish water; N Baltic-Kiel & German Bight 

H. distinctum (G.O. Sars,1920) 


H. finmarchium (T. Scott,1903) 

Belt Sea-North Norway-Spitsbergen & Arctic Seas 

H. spinicauda Wells,1961 


H. pterinum Moore,1974 

Isle of Man-W France 

H. paraspinicauda Bodin,1979 

W France 

H. gascognense Bodin,1968 

Bay of Biscay 

H. canaliculatum (Por,1964) 

= Ectinosoma canaliculatum Por,1964 

Atlantic Europe 

H. cooperatum Bodin, Bodiu & Soyer,1972 

W & S France, Biscay 

H. valeriae Soyer,1973 

?Atlantic Europe & French Mediterranean 

H. argyllensis Clement & Moore,1995 

Scotland 

H. crenulatum Clement & Moore,1995 

Scotland 

H. denticulatum Clement & Moore,1995 

Scotland 

H. pseudosarsi Clement & Moore,1995 

Scotland 

Ectinosoma Boeck,1865 

E. melaniceps Boeck,1865 

Kieler Bucht-Bohuslän-North Sea-W Norway-Arctic Sea-Iceland-Br. Isles-Mediterranean & Black Sea 

E. normani T. & A. Scott,1894 

Bohuslän-Sylt-?Helgoland-N Norway-Arctic Seas-Br. Isles-Mediterran. 

E. tenuipes T. & A. Scott,1894 

Bohuslän-S Norway-Br. Isles & Ireland 

E. compressum G.O. Sars,1920 

Kieler Bucht-Bohuslän-S Norway-Helgoland 

E. obtusum G.O. Sars,1920 

Kieler Bucht-S Norway-Wales 

E. reductum Bocic,1954 

?Helgoland 

E. reductum listensis Mielke,1975 

Sylt 

E. dentatum Steuer,1940 

?Europe, SE Mediterranean 

Noodtiella Wells,1965 

= Noodtia Lang,1965 

N. gracile Mielke,1975 

Sylt 

N. lusitanica Wells,1965 

Portugal 

Lineosoma Wells,1965 

L. iscensis Wells,1965 

Exmouth 

Neobradyoidea Olofsson,1917 

Chappuisiidae Chappuis,1940 

* Freshwater species only. 

Phyllognathopodidae Gurney,1932 

* Freshwater species only. 

Neobradyidae Olofsson,1917 

= Marsteiniidae Drzyzimski,1969 

Neobradya T. Scott,1892 

N. pectinifera T. Scott,1892 () 

Kattegatt-Bohuslän-Bergen-Br. Isles 

Marsteinia Drzyzimski,1968 

M. typica Drzycimski,1968 

Bergen 

M. similis Drzycimski,1968 

Bergen 

Antarcticobradya Huys,1987 

Darcythompsoniidae Lang,1936 

Darcythompsonia T. Scott,1906 

D. fairliensis (T. Scott,1899) 

= D. scotti Gurney,1920 

W Norway-Shetlands-Scotland-Ireland 

D. neglecta Redeke,1953 

Netherlands 

Leptocaris T. Scott,1899 

= Horsiella Gurney,1920 

L. brevicornis (van Douwe,1904) 

Freshwater species which may be found in Baltic brackish water, German North Sea coast 

L. biscayensis (Noodt,1955) 

= L. adriatica Petkovski,1955 

W France-Mediterranean 

L. trisetosus (Kunz,1935) 

Finland-Kieler Bucht-Amrum 

L. ignavus (Noodt,1953) 

= Horsiella ignava Noodt,1953 

* May be a synonym of L. minutus T. Sott,1899 

Belt Sea-S Ireland 

L. minutus T. Scott,1899 () 

W Scotland, Devon 

Kristensenia Por,1983 

EXANECHENTERA Lang,1948 

Tachidioidea Boeck,1864 

Euterpinidae Brian,1921 

Euterpina Norman,1903 

E. acutifrons (Dana,1848) 

= Laophonte sagenarum Oliveira,1945 (male) 

= Tachidius hydrobiologicus Oliveira,1945 (female) 

@ Astropecten irregularis 

* Pelagic, with worldwide distrib. in warm-boreotemperate seas 

Bohuslän-North Sea-SW Norway-Br. Isles-Mediterranean 

Thompsonulidae Lang,1948 

Thompsonula T. Scott,1905 

T. hyaenae (I.C. Thompson,1889) 

Kieler Bucht, German Bight-Br. Isles, Algeria 

Caribbula Huys & Gee,1990 ( Thompsonula hyaenae elongata (Gee,1988)) 

Tachidiidae Boeck,1865 

= Microarthridioninae Lang,1948 

Cithadius Bowman,1972 

Tachidius Liljeborg,1853 

T. (Tachidius) discipes Giesbrecht,1881 

Åland & Finland-Askö-Öresund-Belt Sea-German North Sea coast-Oslofjord-N Norway-Spitsbergen-Iceland- 

Br. Isles-N France, Italy 

Geeopsis Huys,1996 

G. incisipes (Klie,1913) 

= Tachidius incisipes Klie,1913 

= Tachidius longicornis Olofsson,1917 

Elbe & Weser mouths-Norfolk-Spitsbergen-Novaja Semlja 

Microarthridion Lang,1944 

M. littorale (Poppe,1881) 

= T. crassicornis T. Scott,1893 

S Finland-Öresund, N Norway-Novaja Semlja, Br. Isles-S North Sea coasts-N France 

M. reductum (Monard,1935) 

= Microarthridion perkinsi Bodin,1970 

Galway, Roscoff 

M. fallax Perkins,1956 

Belt Sea-SE North Sea-Kent-Wales 

M. berberum (Monard,1936) 

Portugal 

Harpacticidae Dana,1846 

Discoharpacticus Noodt,1954 

Paratigriopus Itô,1969 

Zausopsis Lang,1934 

Harpacticellinae Lang,1948 

Harpacticella G.O. Sars,1908 

Harpacticus H. Milne-Edwards,1840 

H. chelifer (O.F. Müller,1776) 

= Cyclops chelifer O.F. Müller,1776 

= Harpacticus helgolandicus Poppe,1884 

Kieler Bucht-Bohuslän-Helgoland-S & N Norway-Iceland-Greenland-Faeroes-Br. Isles-N France- 


H. uniremis Krøyer,1842 

Kieler Bucht-Bohuslän-Helgoland-S & N Norway-Arctic Seas-Iceland-Greenland-Br. Isles-Roscoff 

H. gracilis Claus,1863 

= Harpacticus elongatus Boeck,1864 

29


Kieler Bucht-Bohuslän-German Bight-Trondheim-Faeroes-Br. Isles-N France-Mediterranean-Black Sea 

H. flexus (Brady & Robertson,1873) 

Kieler Bucht-Bohuslän-S North Sea-S & N Norway-Arctic Seas-Br. Isles-N France-Mediterranean-Black Sea 

H. obscurus T. Scott,1895 

= H. gracilis : G.O. Sars,1911, non Claus,1863 

* Very difficult to separate from H. littoralis and H. giesbrechti 

Kieler Bucht-Bohuslän-German Bight-Br. Isles-Roscoff 

H. littoralis G.O. Sars,1910 

* Very difficult to separate from H. lgiesbrechti and H. obscurus 

Bohuslän-W Norway-S North Sea-Br. Isles-Mediterranean-Black Sea 

H. tenellus G.O. Sars,1920 


Oslofjord-S & W Norway-Ireland-Wales 

H. islandicus Klie,1939 

Iceland 

H. pulvinatus Brady,1910 

Portugal 

H. compsonyx Monard,1926 

Scilly Islands-Roscoff-Mediterranean, world-wide in warmer seas 

H. giesbrechti Klie,1927 

= Harpacticus chelifer : Giesbrecht,1881, non (O.F. Müller,1776) 

* Very difficult to separate from H. littoralis and H. obscurus 

Helgoland 

H. septentrionalis Klie,1939 

Iceland 

Tigriopus Norman,1868 

T. brevicornis (O.F. Müller,1776) 

= T. fulvus : Auctt., non Fischer,1860 

=? T. lilljeborgi Norman,1868 northumbriensis Mistakidis,1949 

Öresund-Bohuslän-N Norway-Br. Isles-N & W France- W Spain 

Perissocope Brady,1910 

P. adiastaltus Wells,1968 


P. sp. : Huys & al. 1996 

=? P. xenus Monard,1926 (nom. dub.) 

NE England, ?Banyuls 

Zausodiinae Lang,1948 

Zaus Goodsir,1845 

Z. spinatus Goodsir,1845 () 

Öresund-Bohuslän-Helgoland-S & N Norway-Arctic Seas-Greenland-Faeroes-Shetlands-Br. Isles-N France 

Z. goodsiri Brady,1880 

* According to Huys & al. 1996 this probably belongs to a new genus. 

S North Sea-W Norway-Arctic Seas-Greenland-Br. Isles 

Z. abbreviatus G.O. Sars,1904 

Öresund-Bohuslän-W Norway-Barents Sea-Arctic Seas-Iceland-Faeroes-Ireland 

Z. aurelii Poppe,1884 

Arctic Seas, ?Iceland 

Z. caeruleus Campbell,1929 

= Zaus aurelii : G.O. Sars,1909, non Poppe,1884 

= Zaus schaeferi Klie,1949 

Helgoland, Bergen, Norfolk, Vancouver Island 

Zausodes C.B. Wilson,1932 

Tisboidea Stebbing,1910 

Tisbidae Stebbing,1910, emend. Lang,1948 

Tisbinae Stebbing,1910 

Tisbe Liljeborg,1853 

= Idya Philippi,1843, non Fréminville,1809 

= Idyaea G.O. Sars,1909 

T. furcata (Baird,1837) 

Stockholm archipelago-Bohuslän-S North Sea-S & N Norway-Arctic Seas-Iceland-Faeroes-Shetlands-Br. 

Isles-?Mediterranean 

T. ensifer Fischer,1860 

Bohuslän-Helgoland-N Norway-Faeroes-Br. Isles-Mediterranean & ?Madeira 

T. gracilis (T. Scott,1895) 

Bohuslän-Helgoland-Belgium-N Norway-Arctic Seas-Br. Isles-N France 

T. longicornis (T. & A. Scott,1895) 

Kristiansund-Shetlands-Br. Isles-Roscoff-Mediterranean 

T. minor (T. & A. Scott,1896) 

Kattegatt-Bohuslän-Helgoland.-W Norway-Arctic Seas-Iceland-Br. Isles-Roscoff 

T. elegantula (G.O. Sars,1905) 

Kristiansund-Roscoff-Algeria 

T. angusta (G.O. Sars,1905) 

Norway (Herö)-Ireland-Roscoff-W Mediterran. 

T. tenella (G.O. Sars,1910) 

S Norway-Helgoland-Algeria 

T. graciloides (G.O. Sars,1920) 


T. compacta (G.O. Sars,1920) 

S Norway-Helgoland 

T. dilatata Klie,1949 

= Tisbe bulbisetosa Volkmann-Rocco,1972 

Helgoland, Belgium 

T. holothuriae Humes,1957 

30 

= T. helgolandica Uhlig & Noodt,1966 

@ freeliving or on Holothuria (Holothuria) stellati Delle Chiaje & Holothuria (Holothuria) tubulosa Gmelin 

Helgoland-Belgium-Plymouth-Portugal-Italy, E USA 

T. elongata (A. Scott,1896) 

@ Among gills of Homarus gammarus - or found free in mud 

Orkneys-Scotland, NW & SW England, Isle of Man 

T. cluthae (T. Scott,1899) 

Shetlands-Scotland 

T. tenera (G.O. Sars,1905) 

Stavanger-N Norway-Iceland-Ireland-Roscoff 

T. finmarchica (G.O. Sars,1905) 

N Norway 

T. battagliai Volkmann-Rocco,1972 

Belgium 

T. cucumariae Humes,1957 

@ freeliving or on Ocnus planci (Brandt) 

Arcachon, Portugal, Mediterranean, Bermuda 

T. gurneyi (Lang,1934) 

= Idyaea gurneyi Lang,1934 

Atlantic Europe, Campbell Island 

T. reticulata Bocquet,1951 

?Atlantic Europe 

T. clodiensis Battaglia & Fava,1968 

?Atlantic Europe, Mediterranean 

Scutellidium Claus,1866 

S. longicauda (Philippi,1840) 

= Scutellidium tisboides Claus,1866 

= Psamate sargassi G.O. Sars,1916 

Bohuslän-W Norway-Iceland-Br. Isles-N France-Mediterranean-Black Sea 

S. hippolytes (Krøyer,1863) 

= Machairopus minutus G.O. Sars,1905 

Öresund-Bohuslän-Helgoland-N Norway-Greenland-Iceland-Br. Isles 

S. arthuri Poppe,1884 

Iceland 

S. ligusticum (Brian,1920) 

Roscoff-Mediterranean 

Sacodiscus C.B. Wilson,1924 

= Aspidiscus Norman,1868, non Koenig,1825 

S. littoralis (G.O. Sars,1904) 

Bohuslän-W Norway-Iceland-Faeroes-Shetlands-Br. Isles-N France 

S. fasciatus (Norman,1868) 

W Norway-Trondheim-Shetlands-Ireland-Roscoff 

S. humesi Stock,1960 

@ Holothuria (Holothuria) tubulosa Gmelin 

W France, Banyuls 

Cholidyinae Boxshall,1979 

Cholidya Farran,1914 

= Cholydia : Bresciani,1970 

C. polypi Farran,1914 () 

@ between arms of Benthoctopus ergasticus. Also reported from e.g. Bathypolypus 


W Ireland 

Cholidyella Avdeev,1982 

* Originally spelled Cholydiella, but corrected by Avdeev,1986 

C. intermedia (Bresciani,1970) Avdeev,1982 

= Cholydia intermedia Bresciani,1970 

@ Cirrotheutidae 

Faeroe-Shetland Channel 

Idyanthinae Lang,1948 

Tachidiella G.O. Sars,1909 

T. minuta G.O. Sars,1911 

Bohuslän-German Bight-Norway (Skutesnaes)-Roscoff 

Zosime Boeck,1872 

Z. valida G.O. Sars,1919 

Kieler Bucht-Bohuslän-N Norway-Spitsbergen-Arctic Seas-Faeroes-Br. Isles 

Z. bergensis Drzycimski,1968 

Bergen 

Z. typica Boeck,1872 


Z. incrassata G.O. Sars,1910 


Z. major G.O. Sars,1919 


Z. gisleni Lang,1948 


Idyella G.O. Sars,1905 

I. pallidula G.O. Sars,1905 

Bohuslän-Bergen, Arctic Seas 

I. exigua G.O. Sars,1905 

Bohusl-Bergen, Bermudas, Algeria 

I. major G.O. Sars,1920 


Idyellopsis Lang,1948

I. typica Lang,1948 


Idyanthe G.O. Sars,1909 

= Idyopsis G.O. Sars,1905, non Agassiz,1860 

I. pusilla (G.O. Sars,1905) 

N Norway, W Mediterranean 

I. dilatata (G.O. Sars,1905) 

Sauesund-Bergen-Spitsbergen 

Tachidiopsis G.O. Sars,1911 

T. cyclopoides G.O. Sars,1911 

S Norway-Bergen 

T. bozici Bodin,1968 

Biscay in 1200 m 

Idyanthopsis Bocquet & Bozic,1955 

* Dubious genus and species. Kunz,1962 considered this species as a synon. of Diarthrodella orbiculata 

Klie,1949. The systematic position of Diarthrodella is also dubious. Different workers locates that genus in 

Paramesochridae and Thalestridae. 

I. psammophila Bocquet & Bozic,1955 

Roscoff 

Porcellidiidae Brady,1880 

Porcellidium Claus,1860 

P. fimbriatum Claus,1863 

= Porcellidium viride : Auctt., ?non Thyone viridis Philippi,1840 

?Kattegatt-?Bohuslän-?Trondheim-?Shetlands-?Br. Isles-Scilly Islands-N France-Canaries-Mediterranean 

P. sarsi Bocquet,1848 

= Porcellidium fimbriatum : Sars,1904, non Claus,1863 

* This species, P. lecanoides Claus,1989 & P. fimbriatum have been so confused, so it is difficult to 

encounter their respective distribution (see P. fimbriatum ) 

P. tenuicauda Claus,1860 

Ireland-Isle of Man-Scilly Islands-Roscoff-Mediterranean 

P. lecanoides Claus,1889 

* See note under P. sarsi 

Ireland, Devon, Roscoff 

P. lecanoides var. roscoffensis Bocquet,1948 

Roscoff 

P. ovatum Haller,1879 

SW Ireland, Mediterranean 

Peltidiidae Claus,1860 

Alteutha Baird,1845 

A. depressa (Baird,1837) 

Br. Isles-Roscoff-Mediterranean-Black Sea 

A. interrupta (Goodsir,1845) 

Bohuslän-S North Sea-Trondheim-Shetlands-Br. Isles-N France-Mediterranean 

A. oblonga (Goodsir,1845) 

Bohuslän-S North Sea-W Norway-Arctic Seas-Shetlands-Br. Isles-N France 

A. sarsi Monard,1935 


S Ireland 

A. roeae Hicks,1982 

Ireland, South Africa 

Peltidium Philippi,1839 

P. purpureum Philippi,1839 

Bohuslän-Trondheim-Br. Isles-Mediterranean 

P. robustum (Claus,1889) 

S Ireland-Engl. Channel-Mediterranean 

Eupelte Claus,1860 

E. sp. : Huys & Al.,1996 

S. England 

E. gracilis Claus,1860 

?Scilly Islands, Mediterranean 

Alteuthella A. Scott,1909 

Alteuthoides Hicks,1986 

Parapeltidium A. Scott,1909 

Alteuthellopsis Lang,1944 

Neopeltopsis Hicks,1976 

Clytemnestridae A. Scott,1909 

= Pseudopeltidiidae Poppe,1891 

Clytemnestra Dana,1848 

* Pelagic 

C. scutellata Dana,1848 

Swedish westcoast, Oslofjord-Br. Isles-Bay of Biscay-Mediterranean 

C. rostrata (Brady,1883) 

Engl. Channel-Bay of Biscay-Mediterranean 

Tegastidae G.O. Sars,1904 

Tegastes Norman,1903 

T. grandimanus G.O. Sars,1904 

Öresund-W Norway (Eggesboenaes)-Br. Isles 

T. longimanus (Claus,1863) 

SW Norway-Helgoland 

T. falcatus (Norman,1868) 

Oslofjord-N Norway-Arctic Seas-Shetlands-Orkneys-Br. Isles, Ireland 

T. nanus G.O. Sars,1904 

Kieler Bucht-W Norway (Christiansund)-Ireland 

T. clausi G.O. Sars,1904 

W Norway (Eggesboenaes)-German Bight, Britain 

T. calcaratus G.O. Sars,1910 

W Norway 

T. flavidus G.O. Sars,1904 

W Norway 

T. satyrus (Claus,1860) 

= T. harpacticoides : G.O. Sars,1910 

SW Norway (Skutesnaes), Mediterranean 

Parategastes G.O. Sars,1904 

P. sphaericus (Claus,1863) 

Kieler Bucht-Bohuslän-Helgoland-W Norway-Br. Isles-Roscoff-Mediterranean 

Arawella Cottarelli & Baldari,1987 

Feregastes Fiers,1986 

Syngastes Monard,1924 

PODOGENNONTA Lang,1948 

Thalestroidea G.O. Sars,1905, emend. Lang,1948 

Thalestridae G.O. Sars,1905, emend. Lang,1948 

Thalestrinae G.O. Sars,1905, emend. Lang,1948 

Thalestris Claus,1863 

T. longimana Claus,1863 

@ Ophiothrix fragilis, Ophiopholis aculeata & Buccinum undatum 

Kieler Bucht-Bohuslän-Helgoland-W Norway-Greenland-Faeroes-Br. Isles-N & W France-Mediterranean 

T. rufoviolascens Claus,1866 

Kattegatt-Norway (Herøy)-Scotland-French Mediterranean 

T. brunnea G.O. Sars,1905 

W Norway-Shetlands-Br. Isles 

T. purpurea G.O. Sars,1905 

Kieler Bucht-Kattegatt-W Norway-Iceland-Greenland 

T. gibba (Krøyer,1842) 

N Norway-Arctic Seas-Iceland 

T. frigida T. Scott,1898 

S Norway & Arctic Seas 

T. normani T. Scott,1903 

N Norway 

T. longipes Boeck,1865 

* May belong in Parathalestris 

Oslofjord 

T. curticornis Boeck,1865 

* May belong in Dactylopodia 

Oslofjord 

T. northumbrica (Norman & Brady,1909, ex Brady MS) 

* May be an extreme specimen of another species 


T. denti Brady,1905 

* May belong in Parathalestris 

England 

T. rhodymeniae (Brady,1894) 

@ Palmaria palmata 

W Scotland 

Amenophia Boeck,1865 

A. peltata Boeck,1865 

Kieler Bucht-Bohuslän-Helgoland-Trondheim-Greenland-Br. Isles 

A. pulchella G.O. Sars,1906 

Bergen-Christiansund 

Parathalestris Brady & Robertson,1873 

P. harpacticoides (Claus,1863) 

@ Crossaster papposus 

Kieler Bucht-Bohuslän-North Sea-N Norway-Iceland-Faeroes-Shetlands-Br. Isles-Roscoff-Mediterranean 

P. clausi (Norman,1868) 

Kieler Bucht-Bohuslän-North Sea-N Norway-Shetlands-Br. Isles-N France 

P. hibernica (Brady & Robertson,1873) 

Stavanger-Christiansund-Br. Isles-Ireland-N France 

P. jacksoni (T. Scott,1898) 

S-N Norway-Arctic Seas-Greenland-Iceland 

P. intermedia Gurney,1930 

= Parathalestris plumiseta C.G. Moore,1976 

Schleswig-Holstein North Sea coast-Norfolk-Ireland-Isle of Man-Roscoff 

P. croni (Krøyer,1842) 

= Halithalestris croni : Auctt. 

W-N Norway-Spitsbergen-Arctic Seas-Iceland-Greenland-Br. Isles-Engl. Channel; -- Pelgic - often in 

company with floating seaweed - 

P. irelandica Roe,1958 

S Ireland 

P. cambriensis Wells,1964 

Wales 

P. dovi Marcus,1966 

W France-Black Sea 

Phyllothalestris G.O. Sars,1905 

P. mysis (Claus,1863) 

31

Bohuslän-Trondheim-Faeroes-Br. Isles-N France-Mediterranean-Black Sea 

Rhynchothalestrinae Lang,1948 

Rhynchothalestris G.O. Sars,1905 

R. helgolandica Claus,1863, emend. Huys,1990 

Kieler Bucht-Bohuslän-North Sea-N Norway-Arctic Seas-Greenland-Iceland-Br. Isles-Ireland 

Dactylopusiinae 

= Dactylopodiinae Lang,1936 

Diarthrodes G.M. Thomson,1882 

D. nobilis (Baird,1845) 

Kieler Bucht-Bohuslän-Helgoland-Lofoten-Arctic Seas-Shetlands-Br. Isles-N France-W Mediterranean- 

Madeira 

D. major (T. & A. Scott,1895) 

Haugesund-Vadsø-Helgoland-Br. Isles & Ireland 

D. pygmaeus (T. & A. Scott,1895) 

Kieler Bucht-Bohuslän-North Sea-W Norway-Br. Isles-Roscoff-Mediterranean 

D. minutus (Claus,1863) 

S Norway-Trondheim-North Sea-Ireland-Roscoff-Mediterranean-Black Sea 

D. assimilis (G.O. Sars,1909) 

Norway 

D. ponticus (Kricagin,1873) 

Ireland-Roscoff-Mediterranean-Black Sea 

D. andrewi (T. Scott,1894) 

Br. Isles 

D. roscoffensis (Monard,1935) 

Roscoff 

D. feldmanni Bocquet,1953 

@ Rhodophyceae 

Roscoff 

D. intermedius (T. Scott,1912) 

= Pseudothalestris intermedia T. Scott,1912 

Atlantic Europe, South Orkneys 

D. aegideus (Brian,1927) 

= Westwoodia pygmaea var. aegidea Brian,1927 

?European Atlantic, Mediterranean 

Dactylopusia Norman,1903 

= Dactylopus Claus,1862, non Gill,1959 

= Dactylopodia Lang,1948 

D. vulgaris G.O. Sars,1905 

Kieler Bucht-Bohuslän-Helgoland-N Norway-Arctic Seas-Iceland-Br. Isles-Roscoff-Mediterranean 

D. vulgaris var. dissimilis Brian,1921 

Atlantic Europe, Mediterranean 

D. euryhalina Monard,1935 

= Dactylopodia vulgaris holsatica Noodt,1953 

Kiel Channel 

D. neglecta (G.O. Sars,1905) 

Trondheim-Spitsbergen-Br. Isles-Ireland-Roscoff 

D. tisboides (Claus,1863) 

Kieler Bucht-Bohuslän-N Norway-Arctic Seas-Shetlands-Br. Isles-N France-Madeira-Mediterranean-Black 

Sea 

D. micronyx G.O. Sars,1905 

Kieler Bucht-Kattegatt-Trondheim-Ireland-Devon-Roscoff 

D. longyerbyenensis (Mielke,1974) 

= Dactylopodia longyerbyenensis Mielke,1974 

Spitsbergen 

Dactylopusioides Brian,1828 

D. macrolabris (Claus,1866) 

= Dactylopus macrolabris Claus,1863 

= Dactylopusioides stampaliae Brian,1928 () 


Paradactylopodia Lang,1948 

= Dactylopusia Norman,1903 (p.p.) 

P. latipes (Boeck,1865) 

Kieler Bucht-Bohuslän-Helgoland-W Norway-Ireland 

P. brevicornis (Claus,1866) 

@ Limnoria lignorum 

Kieler Bucht-Bohuslän-Helgoland-S & N Norway-Br. Isles-Engl. Channel-Mediterranean-Black Sea 

Dactylopodopsis G.O. Sars,1911 

D. dilatata G.O. Sars,1911 

Bohuslän-Lofoten-Iceland 

Pseudotachidiinae Lang,1936 

Pseudotachidius T. Scott,1897 

P. coronatus T. Scott,1897 


P. vikingus Drzycimski,1968 

Bergen 

P. similis T. Scott,1902 

Faeroe Channel 

Idomene Philippi,1843 

I. forficata Philippi,1843 

Öresund-Kattegatt-Oslofjord-Trondheim-Br. Isles-Ireland-Italy 

I. coronata (T. Scott,1894) 

S Norway-Arctic Seas-Scotland 

I. pectinata (T. & A. Scott,1898) 

Scotland 

I. purpurocincta (Norman & T. Scott,1905) 

England (Devon) 

I. borealis G.O. Sars,1911 

Lofoten 

I. aberrans Por,1964 

= Idomene peresi Bodin,1964 

* Not belonging in this genus or even family? 


Dactylopodella G.O. Sars,1905 

D. flava (Claus,1866) 

= Dactylopodella ornata (Norman & T. Scott,1905) 

Bohuslän-Helgoland-W Norway-Arctic Seas-Br. Isles-Mediterranean 

D. clypeata G.O. Sars,1911 

S Norway-Bergen 

D. rostrata (T. Scott,1893) 

Scotland 

Donsiellinae Lang,1948 

Donsiella Stephensen,1936 

D. limnoriae Stephensen,1936 

@ Usually on ventral surface, especially around the bases of the legs on Limnoria lignorum (Rathke), L. 

tripunctata, L. quadripunctata & L. japonica 

Trondheim-Southampton, SE Ireland (British records may perhaps have been D. anglica ), E Canada 

D. anglica Hicks,1988 

Britain 

Ambunguipedidae Huys,1990 

Ambungipes Huys,1990 

A. rufocincta (Brady,1880, ex Norman MS) 

= Thalestris rufocincta Brady,1880, ex Norman MS () 

= Rhynchothalestris rufocincta (Brady,1880, ex Norman MS) 

Bohuslän-Trondheim-Br. Isles-N France-Madeira-Mediterranean 

Lucayostratiotes Huys,1990 ( Rhynchothalestris cornuta Geddes,1969) 

Balaenophilidae G.O. Sars,1910 

Balaenophilus Christian Aurivillius,1879 

B. unisetosus Christian Aurivillius,1879 

@ On baleen plates of the Balaenoptera spp. 

N Norway, Isle of Man, W Ireland 

Parastenheliidae Lang,1936 

Parastenhelia Thompson & Scott,1903 

P. spinosa (Fischer,1860) 

= Thalestris forficula Claus,1863 

= Microthalestris forficula : G.O. Sars.1911 

Kieler Bucht-Bohuslän-W & N Norway-Arctic Seas-Faeroes-Br. Isles-Roscoff-Mediterranean 

P. spinosa bulbosa (Bozic,1954) 

S England 

P. littoralis (G.O. Sars,1911) 

= Microthalestris littoralis G.O. Sars,1911 

* Distribution not ascertained, because of presumed identity of this species with P. spinosa (Fischer) 


P. anglica Norman & T. Scott,1905 

Cornwall-Scilly Isles 

P. ornatissima (Monard,1937) 

Roscoff 

Diosaccidae G.O. Sars,1906 

Psammotopa Pennak,1942 

= Protoleptastacus Noodt,1952 

P. phyllosetosa (Noodt,1952) 

= Protoleptastacus phyllosetosus Noodt,1952 ( of Protoleptastacus Noodt,1952 - Sylt) 

Nordfriesland-Belgium-W Br. Isles-France-Portugal 

P. polyphylla Noodt,1955 

Arcachon-Portugal 

P. vulgaris Pennak,1942 

SW Br. Isles-Arcachon-E Mediterranean 

P. chappuisi Noodt,1955 

= P. vulgaris : Chappuis,1954, non Pennak,1942 

W France 

Protopsammotopa Geddes,1968 

P. norvegica Geddes,1968 

Bergen 

Robertsonia Brady,1880 

R. tenuis (Brady & Robertson, in Brady,1880) 

= Ectinosoma tenuis Brady & Robertson,1875 (nomen nudum) 

= R. tenuis kieliensis Becker,1970 

Belt Sea-Öresund-Bohuslän-SW Norway-Arctic Seas-Br. Isles 

R. diademata Monard,1926 

Roscoff 

R. celtica (Monard,1935) 

= Teissierella celtica Monard,1935 

Roscoff-S Ireland 

R. propinqua (T. Scott,1893) 

= Dactylopus propinquus T. Scott,1893 

= Amphiascus scotti Sewell,1924 

European Atlantic, Bermudas, India 

R. monardi (Klie,1937) 

= Varnaia monardi Klie,1937 

?European Atlantic, Bulgaria, Romania 

Schizopera G.O. Sars,1905 

S. (Schizopera) clandestina (Klie,1924) 

Finland-Elbe estuary-Aberdeen-Black Sea 

32

S. (Schizopera) compacta de Lint,1922 

Kiel Channel-Elbe mouth-Zuidersee 

S. (Neoschizopera Apostolov,1982) pratensis Noodt,1958 

Belt Sea-German North Sea coast 

S. (Neoschizopera Apostolov,1982) meridionalis Petkovski,1954 

= S. inornata Noodt,1955 

Swedish Baltic 

S. (Neoschizopera Apostolov,1982) meridionalis listensis Mielke,1975 

Sylt 

S. baltica Lang,1965 

Askö area (Baltic) & S Norway 

S. ornata Noodt & Purasjoki,1953 

Finnish & German Baltic, S Norway & Spitsbergen 

S. minuta Noodt,1955 

W France 

S. nana Noodt,1955 

W France 

Eoschizopera Wells & Rao,1976 

E. (Eoschizopera) syltensis (Mielke,1973) 

= Schizopera syltensis Mielke,1973 

Sylt 

Pseudodiosaccus T. Scott,1906 

P. propinquus (T. & A. Scott,1893) 

Scotland 

Pseudomesochra T. Scott,1902 

= Stenheliopsis G.O. Sars,1906 

= Jonesiella Monard,1927 

P. longifurcata T. Scott,1902 

Bohuslän-S Norway-Scotland 

P. divaricata (G.O. Sars,1906) 

Bohuslän-W Norway 

P. similis Lang,1935 

Bohuslän 

P. latifurca (G.O. Sars,1911) 

Norway (Lyngenfjord) 

P. media (G.O. Sars,1911) 

Bohuslän-SW Norway 

P. tatianae Drzycimski,1968 

Bergen 

P. scheibeli Schriever,1982 

Iceland-Faeroer-Ridge 

Stenhelia Boeck,1865 

S. (Stenhelia) gibba Boeck,1865 

@ Crossaster papposus & Musculus tumidus 

Kieler Bucht-Bohuslän-Helgoland-N Norway-Arctic Seas-Br. Isles 

S. (Stenhelia) aemula (T. Scott,1893) 

Bohuslän-SW Norway-Br. Isles-Roscoff 

S. (Stenhelia) proxima G.O. Sars,1909 

Kieler Bucht-Bohuslän-SW Norway-Aberdeen 

S. (Delavalia Brady,1868) palustris (Brady,1868) 

* Tube building & home finding species. 

Finland-Bohuslän-Oslofjord-S Norway-German Bight-Br. Isles-Zuidersee-Roscoff 

S. (Delavalia) palustris bispinosa Bodin,1970 

W France 

S. (Delavalia) longicaudata Boeck,1872 

= Delavalia robusta Brady & Robertson,1875 

Kieler Bucht-Bohuslän-N Norway-Arctic Seas-Br. Isles 

S. (Delavalia) longicautata finmarchica (T. Scott,1903) 

= Delavalia robusta var. finmarchica T. Scott,1903 

N Norway 

S. (Delavalia) reflexa Brady & Robertson,1880 

Belt Sea-Bohuslän-SW Norway-Arctic Seas-Br. Isles 

S. (Delavalia) giesbrechti T. & A. Scott,1896 

SW Norway-Br. Isles-Roscoff 

S. (Delavalia) normani (T. Scott,1905) 

= Delavalia Normani T. Scott,1905 

S Norway-Br. Isles-Mediterranean 

S. (Delavalia) hanstroemi Lang,1948 


S. (Delavalia) confluens Lang,1948 


S. (Delavalia) pygmaea (Brady,1905) 

= Delavalia pygmaea Brady,1905 


S. (Delavalia) mastigochaeta Wells,1965 

Scotland 

S. (Delavalia) noodti Schriever,1982 


S. (Delavalia) islandica Schriever,1982 


S. (Delavalia) arctica (T. Scott,1898) 

= Delavalia arctica T. Scott,1898 

off N Koster in 110m, Franz Joseph Land, King Karls Land 

S. (Delavalia) elisabethae Por,1960 

?Britain, Black Sea 

Diosaccus Boeck,1872 

= Paradiosaccus Lang,1948 

D. tenuicornis (Claus,1863) 

Belt Sea-Bohuslän-North Sea-S & N Norway-Br. Isles-Engl. Channel-Canaries-Mediterranean 

D. varicolor (Farran,1913) 

Ireland-Roscoff-Mediterranean 

Robertgurneya Lang,1948 

R. spinulosa (G.O. Sars,1911) 

= Amphiascus spinulosus G.O. Sars,1911 

Bohuslän-S & W Norway 

R. similis (A. Scott,1896) 

= Amphiascus scotti Williamson,1958 

off Gullmarn (Bonden & Mittskären)-S-W Norway-Ireland-England-Roscoff-Bermudas-Mediterranean 

R. ilievecensis (Monard,1935) 

= Amphiascus ilevecensis Monard,1935 

N Ireland-Roscoff 

R. remanei Klie,1950 

Helgoland 

R. intermedia Bozic,1954 

* Dubious species 

Roscoff 

R. dictydiophora (Monard,1924) 

= Amphiascus dictydiophorus Monard,1924 


R. erythraeus (A. Scott,1902) 

= Stenhelia erythraea A. Scott,1902 

?European Atlantic, Massachusetts, Red Sea 

R. oligochaeta Noodt,1955 

Isle of Man, Marmara Sea 

R. rostrata (Gurney,1927) 

= Amphiascus rostratus Gurney,1927 

Celtic Sea, Bermudas, Mediterranean 

R. simulans (Norman & T. Scott,1905) 

= Stenhelia simulans Norman & T. Scott,1905 

SW Britain 

Haloschizopera Lang,1948 

H. exigua (G.O. Sars,1906) 

Bohuslän-SW Norway 

H. bulbifer (G.O. Sars,1911) 

Kosterfjorden in 72-110m-S Norway 

H. conspicua Por,1964 

off Gullmarn (Mittskären in 27-30m) 

H. minima Por,1964 

off Gullmarn (Mittskären in 27-30m) 

H. pygmaea (Norman & T. Scott,1905) 

= Haloschizopera junodi (Monard,1935) 

= Haloschizopera marmarae Noodt,1955 

Bohuslän-Helgoland-Bergen-Roscoff-Br. Isles 

H. pauciseta Por,1959 


H. ruthorum Por,1959 

?Isle of Man, Mediterranean 

Paramphiascella Lang,1948 

P. intermedia (T. Scott,1896) 

Kieler Bucht-Bohuslän-Trondheim-Arctic Seas-Faeroe Channel-Br. Isles 

P. vararensis (T. Scott,1903) 

= Amphiascus affinis G.O. Sars,1906 

Kieler Bucht-Bohuslän-Helgoland-W Norway-Arctic Seas-Scotland-S Ireland 

P. hispida (Brady,1880) 

Bohuslän-N Norway-Arctic Seas-Iceland-Br. Isles-Roscoff 

P. fulvofasciata Rosenfield & Coull,1974 

@ ? Limnoria lignorum 

Helgoland-Norfolk 

P. hyperborea (T. Scott,1903) 

N Norway-Greenland-Arctic Seas 

P. roberti (Monard,1935) 


Roscoff 

P. bodini Marcotte,1974 

= P. mediterranea : Bodin,1964, non Lang,1948 

Celtic Sea, Mediterranean 

Amphiascoides Nicholls,1941 

= Amphiascella Lang,1948 

A. debilis (Giesbrecht,1881) 

=? A. limicola (Brady,1900) 

Finland-Bohuslän-S North Sea-S Norway-Trondheim-Br. Isles-Ireland-Roscoff 

A. dispar (T. & A. Scott,1894) 

Belt Sea-Bohuslän-Scotland 

A. neglecta (Norman & T. Scott,1905) 

= Amphiascella neglecta : Lang,1948 

SW Norway-England-Scilly Isles-Roscoff-Mediterranean 

A. nanus (G.O. Sars,1906) 

= Amphiascella nana : Lang,1948 

Hållö in 6m-S Norway-Roscoff-Mediterranean 

A. littoralis (T. Scott,1903) 

Scotland 

A. nanoides (G.O. Sars,1911) 

S Norway 

A. brevifurca (Czerniavski,1868) 

S Ireland-Roscoff-Mediterranean-Black Sea 

33

A. subdebilis (Willey,1835) 

Kosterfjorden in 72-240m-Skagerrak-Bergen-S Ireland-Roscoff-Mediterranean-Bermudas 

Metamphiascopsis Lang,1948 

M. hirsutus (I.C. Thompson & A. Scott,1903) 

= Dactylophusia hirsuta I.C. Thompson & A. Scott,1903 


Typhlamphiascus Lang,1948 

T. blanchardi (T. & A. Scott,1895) 

Bohuslän-SW Norway-Scotland-Scilly Isles 

T. gracilis Por,1963 

off N Koster-Skagerrak 

T. confusus confusus (T. Scott,1902) 

Bohuslän-Helgoland-Bergen-Br. Isles-Roscoff-Mediterranean 

T. confusus gullmaricus Por,1964 

Bohuslän-Bergen-Mediterranean 

T. typhloides (G.O. Sars,1911) 

S-SW Norway 

T. typhlops (G.O. Sars,1906) 

Kieler Bucht-Bohuslän-SW Norway-Arctic Seas 

T. lamellifer (G.O. Sars,1911) 

Kieler Bucht-Bohuslän-S Norway-Barents Sea-Iceland 

T. hirsutus (Thompson,1893) 

= Stenhelia hirsuta Thompson,1893 

* Problematic species, which may be identical with e.g. T. confusus 

N Norway-Br. Isles 

Bulbamphiascus Lang,1948 

B. imus (Brady,1872) 

= Canthocamptus imus Brady,1872 

= Stenhelia reflexa T. Scott,1895, non Brady & Robertson,1880 

= Amphiascus Normani G.O. Sars,1911 

Bohuslän-W & N?. Norway-Arctic Seas-Br. Isles-N France-Mediterranean 

B. denticulatus (Thompson,1893) 

off Gullmarn (Bonden)-S Norway-Helgoland-Br. Isles 

B. angustifolius Klie,1950 

Helgoland 

Rhyncholagena Lang,1948 

R. lagenirostris (G.O. Sars,1911) 

S Norway 

R. spinifer (Farran,1913) 

Ireland 

R. pestai (Monard,1935) 

Roscoff-Algeria 

Pararobertsonia Lang,1948 

P. abyssi (Boeck,1872) 

Kosterfjorden in 92-150m-Oslofjord-W Norway 

Dactylopodamphiascopsis Lang,1948 

D. latifolius (G.O. Sars,1909) 

Christiansund-Lofoten-Iceland 

Amphiascus G.O. Sars,1905, emend. Lang,1948 

A. minutus (Claus,1863) 

* A species complex 

Kieler Bucht-Bohuslän-North Sea-N Norway-Arctic Seas-Iceland-Faeroes-Br. Isles-Mediterranean 

A. tenuiremis (Brady & Robertson, in Brady,1880) 

=? A. minutus sp.1, Lang,1965 (according to Mielke,1974, though Scheibel,1973 was of the opinion that 

this form is a synonym of A. graciloides trisetosus Klie) 

Kieler Bucht-BohuslänGerman Bight-N Norway-Arctic Seas-Br. Isles-Roscoff 

A. varians (Norman & T. Scott,1905) 

Kosterfjorden in 240m-S-W Norway-Helgoland-England-Roscoff 

A. propinquus G.O. Sars,1906 

= A. angustipes Gurney,1927 

Kieler Bucht-Bohuslän-W Norway-?Scilly Islands 

A. tenellus G.O. Sars,1906 

Bohuslän-W Norway-Roscoff-Algeria 

A. parvus G.O. Sars,1906 

= Amphiascus humphriesi Roe,1960 

S-W Norway-Helgoland-Iceland-S Ireland-Roscoff 

A. sinuatus G.O. Sars,1906 


A. amblyops G.O. Sars,1911 

SW Norway 

A. polaris G.O. Sars,1909 

Bergen-Arctic Seas 

A. graciloides Klie,1950 

Belt Sea-Helgoland 

A. graciloides Klie,1950 trisetosus Klie,1950 

?Helgoland 

A. longarticulatus A. Marcus,1974 

Galway Bay-Scilly Isles-Roscoff-Black Sea 

A. hirtus Gurney,1927 

?Atlantic Europe, Mediterranean, Campbell Island 

Pseudamphiascopsis Lang,1948 

P. attenuatus (G.O. Sars,1906) 

= Pseudamphiascopsis ismaelensis (Monard,1936) 

off Gullmarn (Bonden & Mittskären)-SW Norway-Ireland-Scilly Islands 

P. herdmani (A. Scott,1896) 

Northumberland-Isle of Man 

Paramphiascopsis Lang,1948 

P. longirostris (Claus,1863) 

= Amphiascus paracaudaespinosus Roe,1958 

Kieler Bucht-Bohuslän-Sylt-Helgoland-N Norway-Iceland-Faeroes-Br. Isles-S. Ireland-Roscoff- 

Mediterranean 

P. giesbrechti (G.O. Sars,1906) 

Kieler Bucht-Bohuslän-Helgoland-W Norway-Greenland 

P. pallidus (G.O. Sars,1906) 

Christiansund 

P. triarticulatus Moore,1976 

Isle of Man 

P. paromolae Soyer,1973 

@ gills of Paromola cuvieri (Risso) 


Amphiascopsis Gurney,1927, emend. Lang,1948 

A. cinctus (Claus,1866) 

= Amphiascopsis obscurus : Lang,1948, non (G.O. Sars,1906) 

Bohuslän-Helgoland-W Norway-Ireland-Roscoff-Bermudas-Mediterranean 

A. obscurus (G.O. Sars,1906) 

S Norway 

A. thalestroides (G.O. Sars,1911) 

SW Norway-Helgoland-Shetlands-Br. Isles-N France-Mediterranean 

Amonardia Lang,1948 

A. normani (Brady,1872) 

= Amphiascus similis : Auctt., non (Claus,1866) 

Bohuslän-Helgoland-S- Norway-Trondheim-Faeroes-Br. Isles-Ireland 

A. arctica (T. Scott,1898) 

Kieler Bucht-Bohuslän-N Norway-Arctic Seas-Iceland 

A. phyllopus (G.O. Sars,1906) 

S Norway-Ireland-Roscoff-Mediterranean 

A. similis (Claus,1866) 

S Ireland-Roscoff-Canaries-Mediterranean 

Pholenota Vervoort,1964 ( P. spatulifera Vervoort,1964) 

Cladorostrata Shen & Tai,1963 ( C. brevipoda Shen & Tai,1963 or? C. longipoda Shen & Tai,1963 - 

fresh water (Pearl River), Cina) 

Balucopsylla Chandrasekhara Rao,1972 ( B. similis Chandrasekhara Rao,1972 - India) 

Miraciidae Dana,1846 

= Macrosetellidae A. Scott,1909 

* Family holoplanktic, with species associated to pelagic colonies of in tropical and subtropical seas 

distributed species of Trichodesmium (formerly marine Oscillatoria ). 

Distioculus Huys & Böttger-Schnack,1994 ( Miracia minor T. Scott,1894 - Gulf of Guinea) 

Macrosetella A. Scott,1909 

= Setella Dana,1846, non Schrank,1802 (Lepidoptera) 

= Dwightia C.B. Wilson,1924 

M. gracilis (Dana,1847) 

= Setella gracilis Dana,1847 ( - Pacific) 

= Setella longicauda Dana,1847 

= Setella tenuicornis Dana,1847 

= Setella crassicornis Dana,1847 

= Setella aciculus Dana,1847 

= Setella tenuis Lubbock,1860 

= Setella messinensis Claus,1863 

"Cosmopolitan, planktonic", off W Ireland-southwards 

Miracia Dana,1846 ( Miracia efferata Dana,1849 - Equatorial E Atlantic) 

Oculosetella Dahl,1895 ( Miracia gracilis Dana,1849 - S Pacific) 

Metoidea Boeck,1872 

Metidae Boeck,1872 

Metis Philippi,1843 

M. ignea Philippi,1843 

= Ilyopsyllus coriaceus Brady & Robertson,1873 

Bohuslän-W Norway-German Bight-Br. Isles-Roscoff-Mediterran.-Black Sea 

M. holothuriae (Edwards,1891) 

N France-Mediterranean, Bahamas, New England 

Ameiroidea Monard,1927 

Ameiridae Monard,1927, emend. Lang,1936 

Ameirinae Monard,1927 

Ameira Boeck,1865 

A. scotti G.O. Sars,1911 

Kieler Bucht-Bohuslän-Helgoland-W Norway-Roscoff-Mediterranean 

A. longipes Boeck,1865 

= A. curviseta Willey,1929 

Kieler Bucht-Bohuslän-Helgoland-N Norway-Arctic Seas-Br. Isles-Roscoff-Mediterranean 

A. minuta Boeck,1865 

Öresund-Oslofjord-Telemark-Bergen-Helgoland-Scotland-Ireland 

A. parvula (Claus,1866) 

= Canthocamptus parvulus Claus,1866 

= Nitocra tau Giesbrecht,1881 

= Ameira parvuloides Lang,1965 

Kieler Bucht-Bohuslän-S North Sea-Telemark-Trondheim-Arctic Seas-Br. Isles-Roscoff-Mediterranean-Black 

Sea 

A. tenuicornis T. Scott,1902 

Bohuslän-S & W Norway-Scotland-Scilly Isles 

A. tenella G.O. Sars,1907 

Helgoland-S & SW Norway-Ireland-Scilly Isles 

A. usitata Klie,1950 

Helgoland 

34

A. brevipes Kunz,1954 

Helgoland 

A. brevipes pestae Petkovski,1955 

= Ameira pestae Petkovski,1955 

?European Atlantic, NE Mediterranean 

A. speciosa Monard,1935 

Roscoff 

A. attenuata I.C. Thompson,1893 

* May be identical with A. speciosa Monard 

Isle of Man 

A. exigua T. Scott,1894 

* Belongs probably in Proameira 

Br. Isles-Arctic Seas 

A. pusilla T. Scott,1903 

Scotland-Scilly Isles-?Roscoff-W France 

A. listensis Mielke,1975 

Sylt 

A. divagans Nicholls,1940 

Kieler Bucht, N Bretagne 

A. hyalina (Noodt,1952) 

= Psammameira hyalina Noodt,1952 ( of Psammameira Noodt,1952 - Sylt) 

German North Sea coast 

A. lusitanica Galhano,1970 

Portugal 

A. intermedia Galhano,1970 

Portugal 

A. faroerensis Schriever,1982 


Proameira Lang,1948 

P. arenicola (Lang,1935) 

Öresund-Helgoland 

P. hiddensoensis (Schäfer,1936) 

Finland-Helgoland 

P. simplex (Norman & T. Scott,1905) 

Kieler Bucht-Bohuslän-S Norway-Bergen-Barents Sea-England 

P. dubia (G.O. Sars,1920) 


P. signata Por,1964 

off Gullmarn (Mittskären in 27-30m)-Kosterfjorden-Skagerrak 

P. phaedra (Monard,1935) 

Roscoff 

P. psammophila Wells,1963 

W Br. Isles & Ireland 

P. echinipes Soyer,1975 


Nitokra Boeck,1865 

= Nitocra Giesbrecht,1882 

N. typica Boeck,1865 

= Nitocra oligochaeta Giesbrecht,1881 

= Ameira amphibia Brady,1902 

= Ameira littoralis Nicholls,1939 

Finland-Kieler Bucht-Bohuslän-S North Sea-S Norway-Trondheim-Br. Isles-N France-Mediterranean-Black 

Sea 

N. spinipes Boeck,1865 

= N. parafragilis (Roe,1958) 

Finland-Öresund-Oslofjord-Karmöy-Telemark-S North Sea-Shetlands-Br. Isles-S Ireland-Roscoff- 

Mediterranean-Black Sea 

N. lacustris (Schmankewitsch,1875) 

Freshwater species, entering weak brackish water: e.g. S Finland, Br. Isles 

N. hibernica (Brady,1880) 

Freshwater species, entering weak brackish water: Baltic 

N. pusilla (G.O. Sars,1911) 

= Nitocra pusilla G.O. Sars,1911 

SW Norway-S Ireland 

N. elegans (T. Scott,1904) 

= Ameira elegans T. Scott,1904 

Scotland-Roscoff 

N. fallaciosa Klie,1937 

= N. psammophila Noodt,1952 

German ?North Sea coast 

N. fallaciosa (Klie,1937) baltica Lang,1965 

Finland-S Baltic estuarine sand interstitium species 

N. reducta (Schäfer,1936) 

Finland-S Baltic brackish water & Elbe estuary 

N. reducta (Schäfer,1936) fluviatilis Galhano,1968 

Portugal 

N. affinis Gurney,1927 

S Norway-S Ireland-Bermudas-Mediterranean 

N. hamata Bodin,1970 

W France 

N. sewelli Gurney,1927 husmanni Kunz,1976 

N Germany 

Psyllocamptus T. Scott,1899 

P. (Psyllocamptus) minutus G.O. Sars,1911 

Bohuslän-S Norway-Scilly Islands 

P. (Psyllocamptus) minutus G.O. Sars,1911 gelatinosus (Kunz,1951) 

Sylt 

P. (Psyllocamptus) propinquus (T. Scott,1895) 

Helgoland-Scotland 

Ameiropsis G.O. Sars,1907 

A. mixta G.O. Sars,1907 

Bohuslän-W Norway-Ireland-Roscoff 

A. longicornis G.O. Sars,1907 

Skagerrak-S & SW Norway-Helgoland-Ireland 

A. brevicornis G.O. Sars,1907 

S &W Norway-Helgoland-Roscoff-Br. Isles 

A. nobilis G.O. Sars,1911 

S & W Norway-Ireland-Mediterranean 

A. angulifera G.O. Sars,1911 

S Norway 

A. abbreviata G.O. Sars,1911 

Skagerrak-S & W Norway 

A. minor (G.O. Sars,1920) 

off N Koster in 150m-S & W Norway 

Pseudameira G.O. Sars,1911 

P. crassicornis G.O. Sars,1911 

Bohuslän-S & W Norway-Scotland-W France 

P. furcata G.O. Sars,1911 

Belt Sea-Bohuslän-S Norway 

P. gracilis G.O. Sars,1920 


P. mixta G.O. Sars,1920 


P. reflexa (T. Scott,1894) 

Br. Isles-Arctic Seas 

P. reducta Klie,1950 

Helgoland-N Ireland 

P. breviseta Klie,1950 

Helgoland 

P. perplexa Soyer,1975 


Leptomesochra G.O. Sars,1911 

L. confluens G.O. Sars,1911 

Bohuslän-S Norway-Helgoland 

L. macintoshi (T. & A. Scott,1895) 

Kieler Bucht-German Bight-Br. Isles 

L. nasuta Sewell,1940 

W France 

L. attenuata (Nicholls,1940) 

= L. nichollsi Noodt,1955 

non = L. attenuata (A. Scott,1896) (= Interleptomesochra attenuata (A. Scott)) 

W France 

Sicameira Klie,1950 

S. leptoderma Klie,1950 

Helgoland 

S. gracilis (A. Scott,1896) 

= Ameira gracilis A. Scott,1896 

off Gullmarn (Bonden)-Cornwall-Isle of Man 

Pseudoleptomesochrella Lang,1965 

P. halophila (Noodt,1952) 

= Nitocrella halophila Noodt,1952 

= Nitocrella halophila brevifurca Wells,1961 

Sylt-SW Br. Isles-Bulgaria 

Interleptomesochra Lang,1965 

I. attenuata (A. Scott,1896) 

off Gullmarn (Bonden)-S Norway-W Br. Isles-Brittany 

I. elongata (Bozic,1954) 

Brittany 

I. eulitoralis (Noodt,1952) 

= Leptomesochra eulitoralis Noodt,1952 

German Bight-Belgium-W Br. Isles-Portugal-Israel 

I. noodti Galhano,1968 

NW Portugal 

I. tenuicornis (G.O. Sars,1911) 

Korshavn (S Norway)-Engl. Channel 

Sarsameira Wilson,1924 

= Parameira G.O. Sars,1907, non Seidlitz,1868 

= Pseudosarsameira Lang,1948 

S. parva (Boeck,1872) 

= Ameira longiremis var. intermedia T. Scott,1894 

Bohuslän-Trondheim-W Br. Isles-Bay of Biscay 

S. longiremis (T. Scott,1894) 

Bohuslän-S & W Norway-Arctic Seas-Scotland 

S. propinqua (T. Scott,1902) 

off Gullmarn (Bonden)-S & W Norway-Scotland 

S. sarsi Lang,1948 

= Parameira intermedia : G.O. Sars,1920, non (T. Scott,1894) 


S. major (G.O. Sars,1907) 

SW Norway-Arctic Seas, Celtic Sea 

S. giraulti Monard,1935 

Roscoff-Scilly Islands 

S. peresi Bodin,1970 

W France 

S. exilis (T. & A. Scott,1894) 

35

Bohuslän-S Norway-Br. Isles 

Parapseudoleptomesochra Lang,1965 

P. polychaeta (Noodt,1952) 

= Nitocrella polychaeta Noodt,1952 

Sylt-Exmouth 

Parevansula Guille & Soyer,1966 

P. vermiformis Moore,1976 

Isle of Man 

P. mediterranea Guille % Soyer,1966 


Ameiropsyllus Bodin,1977 

A. monardi Bodin,1977 

W France 

Stenocopiinae Lang,1948 

Stenocopia G.O. Sars,1907 

S. longicaudata (T. Scott,1892) 

Bohuslän-SW & W Norway-Helgoland-Br. Isles-Roscoff 

S. setosa G.O. Sars,1907 

Bohuslän-SW & W Norway 

S. spinosa (T. Scott,1892) 

off Gullmarn (Bonden)-Skagerrak-S & W Norway-Scotland-Ireland 

Malacopsyllus G.O. Sars,1911 

M. fragilis G.O. Sars,1911 

S & W Norway 

Anoplosoma G.O. Sars,1911 

A. sordidum G.O. Sars,1911 

S Norway 

Paramesochridae Lang,1944, emend. Lang,1965 

Paramesochrinae Lang,1944 

Paramesochra T. Scott,1892 

P. acutata Klie,1935 

Kieler Bucht-German North Sea Coast 

P. borealis Geddes,1981 

Tromsö 

P. brevifurca Galhano,1970 

NW Portugal-Sardinia 

P. dubia T. Scott,1892 

off Gullmarn (Bonden)-S Norway-Helgoland-Br. Isles-W Ireland-Brittany 

P. helgolandica Kunz,1936 

Belt Sea-W Sweden-Sylt-Helgoland-Belgium-Irish Sea-Bulgaria 

P. pterocaudata Kunz,1936 

Bonden (Bohuslän)-Helgoland 

P. similis Kunz,1936 

Helgoland-Isle of Man-Bulgaria 

P. longicaudata Nicholls,1945 

?European Atlantic, Red Sea, W Australia 

P. mielkei Huys,1987 

Netherlands 

Leptopsyllus T. Scott,1894 

L. harveyi Wells,1963 

Exmouth 

L. paratypicus Nicholls,1939 

W Scotland 

L. reductus Lang,1948 

Bohuslän 

L. typicus T. Scott,1894 


L. elongatus Drzycimski,1967 

Bergen 

L. celticus Bodin & Jackson,1987 

Bretagne-Galway Bay (Ireland) 

Remanea Klie,1929 

R. arenicola Klie,1929 

Gdansk-Simrishamn-Sylt-W Br. Isles-Portugal 

Kliopsyllus Kunz,1962 

K. coelebs (Monard,1935) 

= Paramesochra coelebs Monard,1935 

Exmouth-Wales-Brittany-Mediterranean 

K. constrictus constrictus (Nicholls,1935) 

German Baltic-Sylt-Br. Isles-Portugal-Bulgaria 

K. constrictus orotavae (Noodt,1958) 

= Paramesochra orotavae Noodt,1958 

?European Atlantic, Canaries, E Mediterranean 

K. holsaticus holsaticus (Klie,1929) 

= Paramesochra holsatica Klie,1929 () 

= Paramesochra pygmaea Nicholls,1939 

Kieler Bucht-Sylt-W Br. Isles-Portugal 

K. holsaticus longicaudatus Galhano,1970 

NW Portugal 

K. laurenticus (Nicholls,1939) 

= Paramesochra laurentica Nicholls,1939 

Exmouth 

K. longifurcatus Scheibel,1975 

Kieler Bucht-Sylt 

K. major (Nicholls,1939) 

= Paramesochra major Nicholls,1939 

Exmouth 

K. paraholsaticus Mielke,1975 

Sylt-Belgium 

K. perharidiensis (Wells,1963) 

= Paramesochra perharidiensis Wells,1963 

= Paramesochra coelebs : Bozic,1953, ?non Monard,1935 

Br. Isles-Roscoff-Angola 

K. longisetosus (Krishnaswamy,1951) 

= Paramesochra longisetosa Krishnaswamy,1951 

?European Atlantic, Madras coast 

Scottopsyllus Kunz,1962 

S. (S.) minor (T. & A. Scott,1895) Kunz,1962 

= Leptopsyllus minor T. & A. Scott,1895 

= Paramesochra minor : Auctt. 

Kieler Bucht-Hållö in 6-8m-Sylt-Holland-Br. Isles-France-Bulgaria 

S. (S.) robertsoni (T. & A. Scott,1895) Kunz,1962 

= Leptopsyllus robertsoni T. & A. Scott,1895 

= Paramesochra robertsoni : Auctt. 

Scotland-Bulgaria 

S. (S.) herdmani (I.C. Thompson & A. Scott,1900) Kunz,1962 

= Leptopsyllus herdmani I.C. Thompson & A. Scott,1900 

= Paramesochra herdmani (I.C. Thompson & A. Scott,1900) 

Gdansk-Kieler Bucht-S Norway-Sylt-W Br. Isles-Black Sea 

S. (Wellsopsyllus Kunz,1981) gigas (Wells,1965) 

= Paramesochra gigas Wells,1965 ( of Wellsopsyllus Kunz,1981) 

= Kliopsyllus gigas (Wells,1965) 

Scotland 

S. (Intermedopsyllus Kunz,1962) intermedius (T. & A. Scott,1895) 

= Leptopsyllus intermedius T. & A. Scott,1895 

= Paramesochra intermedia (T. & A. Scott,1895) 

Kieler Bucht-Öresund-Hållö in 8m-Sylt-Helgoland-Netherlands-Br. Isles-Portugal-Bulgaria 

S. (Intermedopsyllus Kunz,1962) minutus (Nicholls,1939) 

= Paramesochra minuta Nicholls,1939 

W Scotland 

Apodopsyllus Kunz,1962 

A. africanus Kunz,1962 

?European Atlantic 

A. africanus Kunz,1962 listensis Mielke,1975 

Sylt 

A. arenicolus (Chappuis,1954) Kunz,1962 

= Leptopsyllis arenicola Chappuis,1954 

W France-Algeria-Mozambique 

A. littoralis (Nicholls,1939) Kunz,1962 

= Leptopsyllus littoralis Nicholls,1939 

W Scotland-Scilly Isles-Tunisia 

A. spinipes (Nicholls,1939) Kunz,1962 

= Leptopsyllus spinipes Nicholls,1939 


A. reductus (Petkovski,1955) Kunz,1962 

= Leptopsyllus reductus Petkovski,1955 

= Leptopsyllus perplexus Wells,1963 

Br. Isles 

A. madrasensis (Krishnaswamy,1951) Lang,1965 

= Leptopsyllus madrasensis Krishnaswamy,1951 


Diarthrodellinae 

Diarthrodella Klie,1949 

= Idyanthopsis Bocquet & Bozic,1955 

D. orbiculata Klie,1949 

=? Idyanthopsis psammophila Bocquet & Bozic,1955 

Helgoland-W Mediterranean (& Roscoff if the synonym is correct) 

D. parorbiculata parorbiculata Wells,1963 

Ireland 

D. secunda secunda Kunz,1954 

?Kieler Bucht-Sylt-Belgium-W Mediterranean 

Tetragonicepsidae Lang,1944, emend. Coull,1973 

Diagoniceps Willey,1930 

D. brevicauda Huys,1995 (nom. nud.?) 

France (La Rochelle) 

Aigondiceps Fiers,1995 

A. bocki (Lang,1948) 

= Diagoniceps bocki Lang,1948 () 

Bohuslän-Skagerrak 

A. bodini Fiers,1995 

France 

Paraschizopera Wells,1981 ( P. beckeri Wells,1981) 

= Nidiagoceps Fiers,1995 

P. menaiensis (Geddes,1968) Huys,1995 

= Diagoniceps menaiensis Geddes,1968 ( of Nidiagoceps Fiers,1995) 

W Br. Isles-W France 

Tetragoniceps Brady,1880 

T. scotti G.O. Sars,1911 

S Norway-Scotland 

T. brevicauda T. Scott,1899 

Scotland 

T. malleolatus Brady,1880 

36

Ireland-Scilly Isles-Roscoff-W Mediterranean 

T. bergensis Por,1965 

Bergen 

Pteropsyllus T. Scott,1906 

P. consimilis (T. Scott,1894) 

= P. plebeius plebeius Monard,1935 

S & W Norway-Br. Isles-Ireland-Roscoff 

P. plebeius Monard,1935 furcatus Kunz,1938 

* Possibly conspecific with P. consimilis (T. Scott), but Klie,1971 won't agree with that synonymy. Should, 

if not identical, probably be named P. consimilis furcatus 

off Gullmarn (Bonden)-Helgoland 

Phyllopodopsyllus T. Scott,1906 

= Paraphyllopodopsyllus Lang,1948 

P. bradyi (T. Scott,1892) 

off Gullmarn (Bonden)-S & W Norway-Helgoland-Br. Isles 

P. furciger G.O. Sars,1907 

= P. furcifer : Auctt. 

SW Norway-W Mediterranean 

P. hibernicus (Roe,1955) 

= Paraphyllopodopsyllus hibernicus Roe,1955 

S Ireland 

P. hardingi (Roe,1955) 

= Paraphyllopodopsyllus hardingi Roe,1955 

S Ireland 

P. longicaudatus A. Scott,1909 

Scilly Isles (according to Wells,1961) & Indonesia 

Fearia Coull,1971 ( F. prima Coull,1971 - North Carolina) 

Orthopsyllidae Huys,1991 

Orthopsyllus Brady & Robertson,1873 

O. sarsi Klie,1941 

= O. linearis : G.O. Sars,1909, non (Claus,1866) 

Norway (Skjærstadfjord)-Aberdeen 

O. linearis (Claus,1866) 

= O. agnatus Klie,1950 

= O. propinquus Monard,1926 

Helgoland-Br. Isles-N France-Mediterranean 

O. linearis setosus Boer,1971 

NW France 

O. linearis major Klie,1939 

Iceland 

O. linearis illgi (Chappuis,1958) 

= Enhydrosoma illgi Chappuis,1958 

?NW France 

Canthocamptidae Brady,1880 

Barbaracletodes Becker, Noodt & Schriever,1979 

Mesochra Boeck,1865 

M. pygmaea (Claus,1863) 

Kieler Bucht-Bohuslän-Sylt-Helgoland-S & N Norway-Arctic Seas-Br. Isles-Roscoff-Mediterranean-Black 

Sea 

M. lilljeborgi Boeck,1865 

Finland-Askö-Bohuslän-S North Sea-S Norway-Trondheim-Br. Isles-N France-Mediterranean-Black Sea- 

Caspian Sea 

M. rapiens (Schmeil,1894) 

Örnsköldsvik-Askö-S Baltic, S Norwegian & North Sea Estuaries, Br. Isles-S Ireland 

M. aestuarii Gurney,1921 

Finland-German Baltic & North Sea river mouths, Br. Isles, Black Sea 

M. inconspicua (T. Scott,1899) 

Belt Sea-S Norway-Sylt-Scotland 

M. armoricana Monard,1935 

Roscoff, among Enteromorpha -S Ireland 

M. xenopoda Monard,1935 

Roscoff-Mediterranean-Black Sea 

M. anomala Klie,1950 

Helgoland-W France 

M. heldti Monard,1935 

Exe Estuary (S England)-W France 

M. schmidti Mielke,1974 

Spitsbergen 

M. pontica Marcus,1965 

W France-Black Sea 

M. lindbergi Petkovski,1964 

* Is this a marine species? 

Portugal 

M. reducta Klie,1950 

* Dubious species. 

Helgoland 

Nannomesochra Wilson,1932 

= Pseudomesochra Gurney,1927, non Scott,1902 

N. arupinensis (Brian,1925) 

Belt Sea-Ireland-Roscoff-Mediterranean-Black Sea 

Itunella Brady,1896 

I. tenuiremis (T. Scott,1893) 

Bohuslän-Scotland 

I. muelleri (Gagern,1922) 

= I. bacescui Chappuis &Serban,1953 

Brackish water sp.: Finland-Sweden-Germany-Ireland-France 

Bryocamptus Chappuis,1928 

B. (Bryocamptus) pygmaeus (G.O. Sars,1863) 

Freshwater species, which enters Baltic estuarine water 

B. (Bryocamptus) minutus (Claus,18??) 

Freshwater species, which enters Baltic estuarine water 

Psammocamptus Mielke,1975 

P. axi Mielke,1975 

Sylt 

Hemimesochra G.O. Sars,1920 

H. clavularis G.O. Sars,1920 

S Norway 

H. nympha Por,1964 

Kosterfjord in 200m-Skagerrak 

Pusillargillus Huys & Thistle,1989 

P. nixe (Por,1964) 

= Hemimesochra nixe Por,1964 

Kosterfjord in 200m-Skagerrak-Bergen 

Mesopsyllus Por,1960 

M. secunda (Wells,1965) 

= Hemimesochra secunda Wells,1965 

Scotland 

Heteropsyllus T. Scott,1894 

= Cletomesochra G.O. Sars,1920 

H. nanus (G.O. Sars,1920) 


H. major (G.O. Sars,1920) 

Belt Sea-Bohuslän-S & W Norway-W France 

H. exiguus (G.O. Sars,1911) 

SW & W Norway 

H. masculus Kunz,1971 

Helgoland 

H. curticaudatus T. Scott,1894 

Br. Isles 

H. rostratus (G.O. Sars,1920) 

Kosterfjord & off N Koster in 150-200m-S Norway-Arctic Seas 

H. meridionalis Soyer,1975 


H. confluens Soyer,1975 


Bathycamptus Huys & Thistle,1989 

B. minutus (Wells,1965) 

= Heteropsyllus minutus Wells,1965 

Scotland 

Cletocamptus Schmankewitsch,1875 

* Should possibly be included in Cletodidae. 

C. confluens (Schmeil,1894) 

S Baltic river mouths 

Boreolimella Huys & Thistle,1989 

B. dubia (Wells,1965) 

= Leimia Willey,1925 dubia Wells,1965 

Scotland 

Parepactophanes Kunz,1935 

* Should possibly be included in Cletodidae. 

P. minuta Kunz,1935 

Kieler Bucht-German North Sea Coast 

Cylindropsyllidae G.O. Sars,1909, emend. Lang,1948 

Cylindropsyllus Brady,1880 

= Cylindrosoma Brady,1878, non Tschudi,1838, nec Gray, in Todd, 1843, nec Rondani,1856 

C. laevis Brady,1880 () 

Belt Sea-Bohuslän-S & W Norway-Helgoland-Br. Isles-N & W France-Mediterranean 

C. remanei Kunz,1949 

Helgoland-Ireland-Isle of Man-Portugal 

C. kunzi Huys,1988 

Netherlands Delta region 

C. ibericus Huys & Willems,1993 

Portugal 

C. govaerei Huys & Willems,1993 

Corsica 

Evansula T. Scott,1906 

E. incerta (T. Scott,1892) 

Kieler Bucht-off Gullmarn (Bonden)-Hållö-German Bight-Lofoten-Br. Isles-Arcachon 

E. pygmaea (T. Scott,1903) 

Kieler Bucht-Sylt-Helgoland-Belgium-SW England-Scotland 

Stenocaris G.O. Sars,1909 

S. baltica Artl,1983 

S Baltic 

S. gracilis G.O. Sars,1909 

Korshavn (S Norway)-Aberdeen 

S. kliei Kunz,1936 

Helgoland-Isle of Man 

S. minor (T. Scott,1892) 

Bonden (Bohuslän)-Korshavn (S Norway)-Helgoland-W Scotland-Roscoff-Bulgaria 

S. minuta Nicholls,1935 

37

Poland-Kieler Bucht-S Norway-Sylt-W Br. Isles-Bulgaria 

S. pygmaea Noodt,1955 

Aberdeen-Arcachon-W France-Biscay 

S. pontica Chappuis & Serban,1953 

Black Sea 

Boreopontia Willems,1981 

B. heipi Willems,1981 ( - Belgium) 

Netherlands, Belgium, Finisterre 

Willemsia Huys & Conry-Dalton,1993 

W. calceola Huys & Conroy-Dalton,1993 ( - S North Sea) 

Netherlands, Belgium 

Stenocaropsis Apostolov,1982 

S. pristina (Wells,1968) 

= Stenocaris pristina Wells,1968 

Scilly Isles-Black Sea 

Cylinula Coull,1971 ( Cylinula proxima Coull,1971 - N Carolina) 

C. arganoi (Cottarelli & Venanzetti,1989) 

Corsica, Italy 

Leptopontiidae Lang,1944 

Leptopontia T. Scott,1902, emend. Huys & Conroy-Dalton,1996 

L. curvicauda T. Scott,1902, emend Huys & Conroy-Dalton,1996 

?Kieler Bucht-off Gullmarn (Bonden)-Skagerrak-Helgoland-W Br. Isles-Bulgaria 

L. dovpori Huys & Conroy-Dalton,1996 

England 

L. flandrica Huys & Conroy-Dalton,1996 

North Sea 

L. punctata Huys & Conroy-Dalton,1996 

Sweden 

Syrticola Willems & Claeys,1982 

S. flandricus Willems & Claeys,1982 () 

Belgium 

S. trispinosus (A. Scott,1896) 

= Tetragoniceps trispinosus A. Scott,1896 

= Evansula trispinosus : Auctt. 

= Evansia trispinosus : Auctt. 

* Considered a species inquirenda by Huys & Ohtsuka 1993 

Isle of Man 

Psammopsyllinae Krishnaswamy,1956 

Psammopsyllus Nicholls,1945 

= Paulianicaris Chappuis,1952 

P. arenarius Enckell,1965 

Spain 

Ichnusella Cottarelli,1971 ( I. eione Cottarelli,1971 - Italy, interstitial) 

Arenopontiidae 

Arenopontia Kunz,1937 

A. (Arenopontia) stygia Noodt,1955 

Arcachon-W France-NW Portugal-Bulgaria 

A. (Arenopontia) subterranea Kunz,1937 

S Baltic-German Bight-Scotland-Scilly Isles-Mediterranean-?Black Sea 

A. (Neoleptastacus Nicholls,1945) spinicaudata (Nicholls,1945) 

W France 

Pararenopontia Bodiou & Colomines,1986 

P. breviarticulata (Mielke,1975) 

= Arenopontia (Arenopontia) breviarticulata Mielke,1975 

Sylt 

Leptastacidae Lang,1944 (Huys, 1992 upgraded this to full family rank) 

Leptastacus T. Scott,1892 

L. aberrans Chappuis,1954 

Arcachon-W Mediterranean 

L. laticaudatus laticaudatus Nicholls,1935 

Belgium-W Br. Isles-Arcachon 

L. laticaudatus intermedius Kunz,1938 

?Kieler Bucht-Hållö in 6-8m-Helgoland-Belgium-W Br. Isles-Brittany-Bulgaria 

L. macronyx (T. Scott,1892) 

S Baltic-off Gullmarn (Bonden)-North Sea-S Norway-Br. Isles-Bulgaria 

L. minutus Chappuis,1954 

Belgium-Portugal 

L. rostratus Nicholls,1939 

Tromsö-Canada 

L. taurica Marinov,1974 

* Not included in Bodin's catalogue from 1979. 

?European Atlantic, Black Sea 

Paraleptastacus C.B. Wilson,1932 

P. espinulatus Nicholls,1935 

Kieler Bucht-Belgium-Br. Isles 

P. spinicauda (T. & A. Scott,1895) 

Finland-Estonia-Askö-Kattegatt-German Bight-Br. Isles-Roscoff-Portugal-Black Sea 

P. holsaticus Kunz,1937 

Belt Sea-North Sea-Scotland 

P. supralitoralis Mielke,1975 

Sylt 

P. kliei (Gagern,1923) 

S Norway 

Psammastacus Nicholls,1935 

P. brevicaudatus Nicholls,1935 

W Scotland 

P. confluens Nicholls,1935 

?Belt Sea-Nordfriesland-E & W Scotland-N & W France-Algeria-Bulgaria 

P. perplexus Wells,1965 

Scotland-W Portugal 

P. remanei Noodt,1964 

Belgium 

Psamathea Cottarelli & Venanzetti,1989 ( P. nautarum Cottarelli & Venanzetti,1989) 

P. brittanica Huys, Bodiou & Bodin,1996 

Brittany 

Arenocaris Nicholls,1935 

A. bifida Nicholls,1935 

German Baltic-Nordfriesland-Engl. Channel-Scotland 

Schizothrix Huys,1992 

Parastenocarididae Chappuis,1933 

* A freshwater species family. 

Parastenocaris Kessler,1913 

P. fontinalis fontinalis Schnitter & Chappuis,1915 

Freshwater species; may be found in Baltic estuaries 

P. fontinalis borea Kiefer,1960 

Freshwater species; may be found in estuaries in N Europe 

P. phyllura Kiefer,1938 

Baltic brackish water (Swe., Denm., Finl.), e.g. Askö-Nordfriesland 

P. vicesima Klie,1935 

Finland-Askö-S Baltic-S Norway-Nordfriesland-Anglesey-Aberdeen 

Louriniidae Monard,1927 

* No species closer than in the Mediterranean? 

Cletodoidea T. Scott,1904 

Rhizothricidae Por,1986 

Rhizothrix G.O. Sars,1909 

= Rhizothrix Brady & Robertson,1875 (nom. nud.?) 

R. curvata Brady & Robertson,1880 

Öresund-Bohuslän-SW Norway-Helgoland-Br. Isles-Brittany-Mediterranean-Black Sea 

R. gracilis (T. Scott,1903) 

German Baltic-S Norway-Sylt-Helgoland-Br. Isles-W Mediterranean 

R. minuta (T. Scott,1903) 

Kieler Bucht-Öresund-Helgoland-Brittany-Br. Isles & Ireland 

R. reducta reducta Noodt,1952 

Northfriesland-Arcachon-?S France 

R. reducta noodti Galhano,1970 

Sylt-Portugal 

R. wilsoni Bodin,1979 

W France 

Tryphoema Monard,1926 

= Adelopoda Pennak,1942 

T. bocqueti (Bozic,1953) 

= Rhizothrix bocqueti Bozic,1953 

Hållö in 6-8m-Sylt-W Br. Isles-Roscoff-Arcachon 

T. lusitanica (Wells & Clark,1965) 

W Br. Isles-W Portugal 

T. porca Monard,1926 

off Gullmarn (Mittskären in 27-30m)-Aberdeen-Isle of Man-Exmouth-Roscoff-Marseille 

T. scilloniensis (Wells,1968) 

Scilly Isles 

Huntemanniidae Por,1986 

Huntemannia Poppe,1884 

H. jadensis Poppe,1884 

Finland-Askö-Öresund, Nordfriesland, Trondheim, Scotland 

H. micropus Monard,1935 

W Br. Isles-Roscoff 

Pseudocletodes T. & A. Scott,1893 

P. vararensis T. & A. Scott,1893 

Scotland 

Metahuntemannia Smirnov,1946 

= Apodella Por,1965 

M. drzycimskii Soyer,1970 

= M. smirnovi Drzycimski,1968, non Bodin,1968 

Bergen 

M. spinosa (Klie,1941) 

Iceland 

M. crassa (Por,1965) 

= Apodella crassa Por,1965 ( of Apodella Por,1965) 

Bergen 

Pontopolites T. Scott,1894 

P. typicus T. Scott,1894 

Kieler Bucht-Bohuslän-SW Norway-Nordfriesland-Br. Isles 

Nannopus Brady,1880 

38

N. palustris Brady,1880 

Finland & Stockholm skerries-S Baltic-Oslofjord-Telemark-German Bight-Netherlands-Iceland-Br. Isles 

Danielsseniidae Huys & Gee,1996 

= Paranannopidae Por,1986 

Carolinicola Huys & Thistle,1989 

* Placed as incertae sedis in family. 

Cylindronannopus Coull,1973 

Mucrosenia Gee & Huys,1994 

M. kendalli Gee & Huys,1994 ( - Spitsbergen) 

Spitsbergen 

M. kliei (Smirnov,1946) 

= Psammis kliei Smirnov,1946 

? 

Bathypsammis Huys & Gee,1993 ( Psammis longifurca Bodin,1968) 

Leptotachidia Becker,1974 

Paranannopus Huys & Gee,1996 

= Paranannopus Lang,1936 (nom. nud.?) 

P. abyssi (G.O. Sars,1920) 

S Norway 

P. bahusiense Por,1964 

off N Koster in 110m 

P. triarticulatus Wells,1965 

= P. langi Wells,1965 

* The female was described as P. triarticulatus and the male as P. langi 

Scotland 

P. aff. philistinus Por,1964 

Celtic Sea 

P. sarsi Lang,1936 

Spitsbergen 

Danielssenia Boeck,1872 

D. typica Boeck,1872 

= D. fusiformis : G.O. Sars,1910, non Brady,1880 

Poland-Bohuslän-W & N Norway-Spitsbergen-Arctic Seas, Helgoland-Br. Isles 

D. quadriseta Gee,1988 

Norway 

D. spitsbergensis Gee & Huys,1994 

Spitsbergen 

Jonesiella Brady,1880 

=Sentirenia Huys & Gee,1992 

J. fusiformis Brady,1880 

= Danielssenia perezi Monard,1935 ( of Sentirenia Huys & Gee,1992) 

= Danielssenia paraperezi Soyer,1970 

Firth of Clyde-Liverpool Bay-Scilly Islands-Roscoff-Mediterranean 

Paradanielssenia Soyer,1970 ( P. kunzi Soyer,1970 - Mediterranean) 

P. biclavata Gee,1988 

S Celtic Sea 

P. christineae Gee & Huys,1994 

Spitsbergen 

P. kathleenae Gee & Huys,1994 

Spitsbergen 

Fladenia Gee & Huys,1990 

F. robusta (G.O. Sars,1921) 

= Danielssenia robusta G.O. Sars,1921 

= Danielssenia intermedia Wells,1965 () 

Bohuslän-Bergen, Scotland 

Psammis G.O. Sars,1910 

P. longisetosa G.O. Sars,1910 

Bohuslän-Bergen, Scotland 

Anapophysia Huys & Gee,1996 

A. borealis (Klie,1939) 

= Psammis borealis Klie,1939 () 

Iceland 

Micropsammis Mielke,1975 

M. noodti Mielke,1975 () 

Sylt 

Telopsammis Gee & Huys,1991 

T. secunda (Mielke,1975) 

= Micropsammis secunda Mielke,1975 () 

Sylt 

Archisenia Huys & Gee,1992 

A. sibirica (G.O. Sars,1898) 

= Danielssenia sibirica G.O. Sars,1898 () 

= Archisenia stefanssonii (Willey,1920) 

?Scotland, Arctic Seas 

Afrosenia Huys & Gee,1996 ( Danielssenia spinipes Wells,1967) 

Sentiropsis Huys & Gee,1996 ( Danielssenia minuta Coull,1969) 

Peltsenia Huys & Gee,1996 ( Idomene aberrans Por,1964) 

Prionos Huys & Gee,1996 ( P. ornata Huys & Gee,1996 - Malaysia) 

Cithadius Bowman,1972 ( C. cythurae Bowman,1972 - ? - @ Cyathura polita ) 

Argestidae Por,1986 

Neoargestes Drzycimski,1967 

N. variabilis Drzycimski,1967 

Bergen 

Parargestes Lang,1948 

P. tenuis (G.O. Sars,1921) 

Bohuslän-S & W Norway-Spitsbergen 

Argestes G.O. Sars,1910 

A. mollis G.O. Sars,1910 

Bohuslän-W Norway-Arctic Seas 

Fultonia T. Scott,1902 

F. hirsuta T. Scott,1902 

Bohuslän-SW & W Norway-Scotland 

Leptocletodes G.O. Sars,1920 

L. debilis G.O. Sars,1920 

Bohuslän-S & W Norway 

Eurycletodes G.O. Sars,1909 

E. (Eurycletodes) laticauda (Boeck,1872) 

Bohuslän-Haugesund 

E. (Eurycletodes) serratus G.O. Sars,1920 

Bohuslän 

E. (Oligocletodes Lang,1948) latus (T. Scott,1892) 

SW & W-?N Norway-Br. Isles 

E. (Oligocletodes) major G.O. Sars,1909 

Öresund-Bohuslän-S Norway-Arctic Seas 

E. (Oligocletodes) similis (T. Scott,1895) 

Bohuslän-Helgoland-Lofoten-Arctic Seas-Br. Isles-Roscoff 

E. (Oligocletodes) minutus G.O. Sars,1920 


E. (Oligocletodes) aculeatus G.O. Sars,1920 

Bohuslän-Skagen-S & W Norway 

E. (Oligocletodes) oblongus G.O. Sars,1920 

Bohuslän-Skagen-S Norway 

E. (Oligocletodes) pori Drzycimski,1969 

= E. cf. echinatus : Por,1965 

Bergen 

E. (Oligocletodes) irelandica Roe,1959 

S Ireland 

Mesocletodes G.O. Sars,1909 

M. monensis (I.C. Thompson,1893) 

Bohuslän-S & W Norway-Br. Isles 

M. irrasus (T. & A. Scott,1894) 

Bohuslän-Bergen-Scotland 

M. inermis G.O. Sars,1920 

S Norway 

M. arenicola Noodt,1952 

Sylt 

M. abyssicola (T. & A. Scott,1901) 

Bohuslän-Bergen-Spitsbergen 

M. robustus Por,1965 

Bergen 

M. glaber Por,1964 

Skagerrak in 300-425m-Bergen 

M. fladensis Wells,1965 

Scotland 

M. duosetosus Schriever,1985 

Iceland-Faeroe-ridge (500-1850 m) 

M. thieli Schriever,1985 


M. faroerensis Schriever,1985 


M. kunzi Schriever,1985 


M. quadrispinosa Schriever,1985 


M. carpinei Soyer,1975 


M. katharinae Soyer,1964 


Argestigens Willey,1935 

A. uniremis Willey,1935 ( - Bermudas) 

?European Atlantic, Bermudas 

Cletodidae T. Scott,1904 

Monocletodes Lang,1936 

M. varians (T. Scott,1903) 

= Thieliella nordatlantica Schriever,1982 ( of Thieliella Schriever,1982)) 

= Thieliella reducta Schriever,1982 

Norway (Bögfjord), Iceland-Faeroer-Ridge 

Acrenhydrosoma Lang,1944 

A. perplexum (T. Scott,1899) 

= Cletodes perplexa T. Scott,1899 

= Enhydrosoma perplexum (T. Scott,1899) Lang,1936 

Bohusl-Norway (Bögfjord)-Br. Isles 

39

Stylicletodes Lang,1936 

S. longicaudatus (Brady & Robertson,1880) 

= S. numidicus (Monard,1935) 

Bohuslän-S Norway-Arctic Seas-Br. Isles 

S. reductus Wells,1965 

Scotland 

Cletodes Brady,1872 

C. endopodita (Schriever,1984) 

= Thieliella endopodita Schriever,1984 

Norw. Sea 

C. pusillus G.O. Sars,1920 

Öresund-Bohuslän-Helgoland-S & W Norway 

C. tenuipes T. Scott,1896 

Öresund & Belt Sea-Bohuslän-Helgoland-N Norway-Arctic Seas-Br. Isles-Ireland 

C. longifurca Lang,1948 


C. longicaudatus (Boeck,1872) 

Bohuslän-SW & W Norway-Br. Isles-Roscoff-Mediterranean 

C. limicola Brady,1872 

Bohuslän-SW Norway-Br. Isles-Mediterranean 

C. latirostris Drzycimski,1967 

=? Enhydrosoma wellsi Bodin,1968 

Bergen & ?W France 

C. smirnovi Bodin,1970 

W France 

C. spinulatus Por,1967 

Bretagne 

Enhydrosoma Boeck,1872 

E. sarsi (T. Scott,1904) 

Öresund & Belt Sea-Bohuslän-S & W Norway-Scotland-W France 

E. longifurcatum G.O. Sars,1909 

* Is together with E. sordidum & E. gariene constituting a natural group 

Kieler Bucht-Bohuslän-Nordfriesland-SW & W Norway-Scotland-Ireland-N & W France 

E. sordidum Monard,1926 

Scotland-Roscoff-Banyuls 

E. gariene Gurney,1930 

Nordfriesland-England (Norfolk) 

E. propinquum (Brady,1880) 

= Cletodes propinqua Brady,1880 

Bohuslän-S Norway-German Bight-Br. Isles-Roscoff-Mediterranean 

E. curticauda Boeck,1872 () 

= Cletodes hirsutipes T. Scott,1896 

Kieler Bucht-Bohuslän-N Norway-White Sea-German Bight-Zuidersee-Br. Isles-Ireland 

E. curvirostre (T. Scott,1894) 

* This will possibly be transferred to a new genus in a forthcoming paper (sensu Gee,1994) 

Kieler Bucht-Bohuslän-N Norway-Br. Isles 

Strongylacron Gee & Huys,1996 

S. buchholzi (Boeck,1872) 

= Cletodes Buchholzii Boeck,1872 () 

= Enhydrosoma buchholtzii (Boeck,1872) 

Kieler Bucht-Bohuslän-German North Sea coast-Haugesund, Anglesey, Ireland, S England, W France 

Enhydrosomella Monard,1935 

Scintis Por,1986 

* Cletodidae incertae sedis. 

Intercletodes Fiers,1987 

Schizacron Gee & Huys,1996 ( Enhydrosoma barnishi Wells,1987 - Mozambique) 

Triathrix Gee & Burgess,1997 ( T. montagni Gee & Burgess,1997 - California) 

Sphingothrix Fiers,1997 ( S. goldi Fiers,1997 - Yucatan shelf) 

Laophontopsidae Huys & Willems,1989 

Laophontopsis G.O. Sars,1908 

= Cleta Claus,1863, non Duponchel,1845 

L. lamellifera (Claus,1863) 

Bohuslän-S Norway-Br. Isles-N France-Mediterranean 

L. borealis Huys & Willems,1989 

Hvaler, Netherlands, Norfolk 

L. monardi Huys & Willems,1989 

France 

Aculeopsis Huys & Willems,1989 ( A. longisetosa Huys & Willems,1989 - Mediterranean France) 

Telodocus Huys & Willems,1989 ( Cleta secunda Sewell,1924) 

Normanellidae Lang,1944 

* Not recognized by Huys & Boxshall 1991 but raised from subfamily to family by Huys & Willems 1989 

Normanellinae Lang,1944 

Normanella Brady,1880 

N. similis Lang,1936 

Öresund & Belt Sea-Bohuslän-Skagen 

N. mucronata G.O. Sars,1909 

Bohuslän-Bergen-Roscoff 

N. tenuifurca G.O. Sars,1909 

Bohuslän-Bergen-Bukken-Roscoff 

N. minuta (Boeck,1872) 

Bohuslän-Helgoland-Trondheim-Ireland-Wales-Roscoff-Bermudas-Mediterranean 

N. quarta Monard,1935 

Roscoff-Algeria 

N. dubia Brady & Robertson, in Brady,1880 


Br. Isles-Ireland 

Cletopsyllinae Huys & Willems,1989 

Cletopsyllus Willey,1935 

Pseudocletopsyllus Vervoort,1964 

Adenopleurellidae Huys,1990 

Sarsocletodes Wilson,1924 

= Pseudocletodes G.O. Sars,1920, non Scott,1893 

S. typicus (G.O. Sars,1920) 


Adenopleurella Huys,1990 ( A. brevipes Huys,1990 - Queensland) 

Miroslavia Apostolov,1980 

Proceropes Huys,1990 ( Sarsocletodes secundus Smirnov,1946) 

Laophontidae T. Scott,1904 

Laophonte Philippi,1840 

L. inopinata T. Scott,1892 

Hållö- off N Koster-S-N Norway-Helgoland-Iceland-White Sea-Br. Isles 

L. denticornis T. Scott,1894 

Bohuslän-SW Norway-Helgoland-Br. Isles-Ireland 

L. nordgaardi G.O. Sars,1908 

Norway (Saltenfjord) 

L. brevifurca G.O. Sars,1920 

Norway (Hvalör)-Black Sea 

L. baltica Klie,1929 

?Kieler Bucht-Kattegatt-Helgoland-S Ireland & White Sea 

L. setosa Boeck,1865 

?German Baltic-Oslofjord-W Norway-German North Sea coast-Iceland-Br. Isles-N France-Mediterranean 

L. thoracica Boeck,1865 

Kieler Bucht-Bohuslän-Helgoland-S Norway-Trondheim-Br. Isles-N France-Mediterranean 

L. longicaudata Boeck,1865 

= Laophonte Hodgii Brady,1872 

Bohuslän-Trondheim-Br. Isles-Ireland 

L. longicaudata reducta Lang,1936 

Öresund 

L. elongata Boeck,1872 

Bohuslän-W Norway-Helgoland-Ireland-Roscoff-Mediterranean 

L. inornata A. Scott,1902 

S England-Roscoff-Mediterranean 

L. parvuloides Monard,1935 

Roscoff 

L. parvula G.O. Sars,1908 

= Laophonte sima Gurney,1927 

Bohuslän-S Norway-Helgoland-W France-Mediterranean 

L. serrata (Claus,1863) 

= Cleta serrata Claus,1863 

Bohuslän-Trondheim-Helgoland-Br. Isles-Ireland 

L. dominicalis Monard,1935 

Roscoff 

L. depressa T. Scott,1894 

W Norway-Arctic Seas-Iceland-Br. Isles 

L. cornuta Philippi,1840 

Bohuslän-Trondheim-Shetlands-Br. Isles-N France-Canaries-Madeira-Mediterran. 

L. trilobata Willey,1929 

= Laophonte foxi Harding,1956 

Sylt-Helgoland-Norfolk-North America 

L. drachi Médioni & Soyer,1967 

= L. danversae Hamond,1969 

@ Schismopora armata (Hincks,1860) 

Norfolk-France 

? L. royi Jakubisiak,1932 

@ ? Maia squinado 

Roscoff 

L. farrani Roe,1958 

* May belong to a not yet described genus 

S Ireland 

L. commensalis Raibaut,1961 

@ among pilose setae, especially on the ventral surface of Xantho pilipes, X. floridus & X. rivulosus 

N France & Mediterranean 

L. adamsiae Raibaut,1966 

@ Adamsia carcinopados 

French Mediterranean 

Laophontina Norman & T. Scott,1905 

L. acantha Noodt,1955 

W France-Portugal 

L. dubia Norman & T. Scott,1905 

Anglesey-Scilly Isles 

Platychelipus Brady,1880 

P. laophontoides G.O. Sars,1908 

Kieler Bucht-Bohuslän-Oslofjord-Helgoland-NE Br. Isles 

P. littoralis Brady,1880 

Oslofjord-German Bight-N Norway-Arctic Seas-Br. Isles-Roscoff 

Pseudolaophonte A. Scott,1896 

40

P. spinosa (I.C. Thompson,1893) 

Bohuslän-S Norway-Helgoland-Scotland-Isle of Man-Scilly Isles 

P. proteus Klie,1950 

Belt Sea 

P. glemareci Bodin,1977 

Roscoff 

Pilifera Noodt,1952 

P. gracilis (T. Scott,1903) 

= Laophonte gracilis T. Scott,1903 () 

= Pseudonychocamptus gracilis : Lang,1948 

Sylt, Firth of Forth 

Harrietella T. Scott,1906 

@ external body associates of crustaceans 

H. simulans (T. Scott,1894) 

@ Attatched to especially the telson of Limnoria lignorum (Rathke) & L. tripunctata Menzies or in their 

excavated galleries. 

Oslofjord-Trondheim-SE & SW Scotland-Wales-Netherlands, NE Ireland, Mediterranean, E USA 

Hemilaophonte Jakubisiak,1932 

H. janinae Jakubisiak,1932 

@ On carapace of Maia squinado 

Roscoff 

Esola Edwards,1891 

E. typhlops (G.O. Sars,1908) 

Bohuslän-Flekkerøy-Bergen-Scotland 

E. longicauda Edwards,1891 

Scotland-Ireland-Bahamas-Bermudas-Mediterranean 

E. longiremis (T. Scott,1904) 

Scotland-Scilly Isles 

E. bulligera (Farran,1913) 

Ireland 

Onychocamptus Daday,1903 

O. mohammed (Blanchard & Richardson,1891) 

Finland-S Baltic, Elbe mouth, Zuidersee and other Eur. brackish water sites 

O. chathemensis (G.O. Sars,1905) 

S Africa-Chatham Islands-?Europe 

Klieonychocamptus Noodt,1958 

K. kliei (Monard,1935) 

* Lang,1965 doubt that this is a natural genus 

Scilly Isles-Roscoff-Yugoslavia 

Echinolaophonte Nicholls,1941 

E. horrida (Norman,1876) 

= Onychocamptus horridus : Auctt. 

Kieler Bucht-Bohuslän-Helgoland-S & N Norway-Arctic Seas-Greenland-Iceland-Br. Isles-Mediterranean 

E. brevispinosa (G.O. Sars,1908) 

S Norway-Mediterranean 

Asellopsis Brady & Robertson,1873 

A. hispida Brady & Robertson,1873 

Kieler Bucht-Bohuslän-S Norway-Bergen-Nordfriesland-Br. Isles-Roscoff-Mediterranean 

A. intermedia (T. Scott,1895) 

Kieler Bucht-Hållö- off N Koster-German Bight-Arctic Seas-Br. Isles-Roscoff-Algeria 

Heterolaophonte Lang,1944 

H. minuta (Boeck,1872) 

= Laophonte subsalsa Brady,1903 

Belt Sea-Bohuslän-Oslofjord-Telemark-Helgoland-Scotland-Wales-Ireland-Scilly Isles & White Sea 

H. stroemii (Baird,1837) (original spelling emended) 

= Cyclops Stromii Baird,1837 

Kattegatt-German Bight-W Norway-Arctic Seas-Greenland-Shetlands-Br. Isles-N France 

H. stroemi paraminuta Noodt,1955 

European Atlantic, Mediterranean, Marmara Sea 

H. stroemi var. brevicaudata (Monard,1928) 

?European Atlantic, Canaries, Mediterranean 

H. littoralis (T. & A. Scott,1893) 

Kieler Bucht-Bohusl-Helgoland-Bergen-Br. Isles-Mediterranean 

H. longisetigera Klie,1950 

= H. littoralis longisetigera Klie,1950 

Belt Sea-Helgoland-NW France 

H. mendax (Klie,1939) 

= Laophonte mendax Klie,1939 

Iceland 

H. uncinata (Czerniavski,1868) 

= Laophonte vietsi Klie,1941 

Roscoff-Mediterranean-Black Sea 

H. phycobates (Monard,1935) 

Roscoff 

H. norvegica Drzycimski,1968 

Bergen 

H. hamondi Hicks,1975 

= Heterolaophonte bisetosa Mielke,1975 

Sylt, Norfolk 

H. denticulata Roe,1958 

S Ireland 

H. brevipes Roe,1958 

S Ireland 

Quinquelaophonte Wells, Hicks & Coull,1982 

Q. quinquespinosa (Sewell,1924) 

= Laophonte quinquespinosa Sewell,1924 

= Heterolaophonte sigmoides : Wells,1963, non (Willey,1930) 

?Exe estuary, Mediterranean, India 

Pseudonychocamptus Lang,1944 

P. koreni (Boeck,1872) 

Bohuslän-S & W Norway-White Sea-German Bight-Ireland-Scilly Isles-Tunesia 

P. proximus (G.O. Sars,1908) 

Kieler Bucht-Sylt-W Norway-Scotland-Isle of Man 

P. abbreviatus (G.O. Sars,1920) 

S & W Norway 

P. carthyi Hamond,1968 

Norfolk 

? P. faroensis (T. Scott,1902) 

= Laophonte faroensis T. Scott,1902 


Paralaophonte Lang,1944 

P. brevirostris (Claus,1863) 

= Laophonte herdmani A. Scott,1902 

Bohuslän-W Norway-Br. Isles-Roscoff-Mediterran.-Black Sea 

P. congenera (G.O. Sars,1908) 

@ Limnoria 

Bohuslän-W Norway-Ireland-Roscoff-Bermudas-Mediterranean 

P. perplexa (T. Scott,1898) 

W-N Norway-Spitsbergen-Arctic Seas-Iceland 

P. macera (G.O. Sars,1908) 

S-W Norway-Arctic Seas 

P. karmensis (G.O. Sars,1911) 

Norway (Skutesnæs)-White Sea 

P. tenera (G.O. Sars,1920) 

S Norway 

P. hyperborea (G.O. Sars,1909) 

Spitsbergen-White Sea 

P. spitzbergensis Mielke,1974 

Spitsbergen 

Paronychocamptus Lang,1944 

P. curticaudatus (Boeck,1865) 

= Laophonte curticaudata Boeck,1964 

Kieler Bucht-Bohuslän-Karmøya-Telemark-Sylt-Helgoland-Scotland-Wales-Ireland-Roscoff-Mediterranean 

P. nanus (G.O. Sars,1908) 

= Laophonte nana G.O. Sars,1908 

Finland-Bohuslän-Oslofjord-Nordfriesland-Zuidersee-N France-Mediterranean 

P. exiguus (G.O. Sars,1905) 

= Laophonte exigua G.O. Sars,1905 

?European Atlantic, Chatham Island 

Namakosiramia Ho & Perkins,1977 

@ one sp. N. californiensis Ho & Perkins,1977 on Stichopus parvimensis (Clark) 

* Huys 1988 moved this to Laophontidae (Harpacticoida) from Namakosiramiidae (Cyclopoida) 

Cristacoxidae Huys,1990 

Cristacoxa Huys,1990 ( C. petkovskii Huys,1990 - Lesser Antilles) 

Cubanocleta Petkovski,1977 ( C. noodti Petkovski,1977 - Cuba) 

Noodtortopsyllus Lang,1965 

Ancorabolidae G.O. Sars,1909 

Laophontodinae Lang,1948 

Laophontodes T. Scott,1894 

= Merope Thomson,1883, non Newman,1838 (Mecoptera), nec Adams,1856 (Mollusca), nec Martens,1860 

ex Albers MS (Mollusca) 

= Meropia Wilson,1932 

L. gracilipes Lang,1936 

Bohuslän 

L. expansus G.O. Sars,1908 

Bohuslän-Norway (Saltenfjord)-?Ireland 

L. bicornis A. Scott,1896 

Bohuslän-W Norway-Br. Isles-Roscoff-Mediterranean 

L. typicus T. Scott,1894 

Bohuslän-Helgoland-W Norway-Arctic Seas-Br. Isles-Roscoff 

L. armatus Lang,1936 

W Scotland, Bermudas, Malvinas 

Ancorabolinae G.O. Sars,1909 

Echinopsyllus G.O. Sars,1909 

E. normani G.O. Sars,1909 

Bohuslän-Bergen-Scotland 

Arthropsyllus G.O. Sars,1909 

A. serratus G.O. Sars,1909 

Kieler Bucht-S Norway-Trondheim-White Sea 

Echinocletodes Lang,1936 

* Should possibly be transferred to Cletodidae? 

E. armatus (T. Scott,1902) 


Ancorabolus Norman,1903 

A. mirabilis Norman,1903 

Bohuslän-N Norway-Scotland 

Ceratonotus G.O. Sars,1909 

C. pectinatus G.O. Sars,1909 

Bohuslän-Norway (Flekkerøy) 

41

C. pectinatus elaphus Por,1965 

Bergen 

Dorsiceratus Drzycimski,1967 

D. octocornis Drzycimski,1967 

Bergen 

SUBORDO incertae sedis 

Latiremidae Bozic,1969 

* Dubious family. Should be named Delamarellidae if it shows to be a good taxon. 

Delamarella Chappuis,1954 

= Latiremus Bozic,1969 

D. arenicola Chappuis,1954 

W France 

D. eximia (Bozic,1969) 

?France 

GELYELLOIDA Huys,1988 

Gelyellidae Rouch & Lescher-Moutoué,1977 

* Only stygobiontic species from European karstic systems. 

REMIPEDIA Yager,1981 

* Atlantic marine troglobionts with one recent order, Nectiopoda Schram,1986, not yet found close to 

Scandinavia, and one extint order Enantiopoda Birshtein,1960 

NECTIPODA Schram,1986 

PHYLLOPODA Latreille,1825 

PHYLLOCARIDA Packard,1879 

LEPTOSTRACA Claus,1880 

* The only recent order. 4 extinct orders. 

Nebaliidae Baird,1850 

Nebalia Leach,1814 

N. borealis Dahl,1985 

Bohuslän-Troms & Shetlands 

N. bipes (O. Fabricius,1780) 

Trondheimsfjord-N Norway-Spitsbergen-Iceland & E Greenland 

N. herbstii Leach,1814 

Shetlands-N Spain 

N. strausi Risso,1826 

Engl. Channel-Adriatic Sea 

Nebaliella Thiele,1904 

N. caboti Clark,1932 

off W Ireland-Rockall Trough 

Sarsinebalia Dahl,1985 

S. typhlops (G.O. Sars,1870) 

Lofoten-Bergen area-European Atlantic-W Mediterranean 

Nebaliopsidae Hessler,1984 

Nebaliopsis G.O. Sars,1887 

N. typica G.O. Sars,1887 

off W Ireland-Rockall Trough-southwards to 50° S 

CALMANOSTRACA Tasch,1969 

NOTOSTRACA G.O. Sars,1867 

* Three Scandinavian (non marine) species in smaller, sometimes temporary, water systems. 

Triop(s)idae Kielhack,1910 

* Only limnic species. 

SARSOSTRACA Tasch,1969 

ANOSTRACA H. Milne-Edwards,1840 

* Five Scandinavian (non marine) species in temporary land-locked waters. 

Chirocephalidae Daday,1910 


Branchipodidae Burmeister,1843 (Simon,1886) 


Artemiidae Grochowski,1896 

* The well-known genus Artemia from e.g. Mediterranean Salines belongs to this group. 

Branchinectidae Daday,1910 

Polyartemiidae Simon,1886 

Streptocephalidae Daday,1910 

Thamnocephalidae Simon,1886 

DIPLOSTRACA Gerstaecker,1866 

CONCHOSTRACA G.O. Sars,1867 

* Two Scandinavian (non marine) species in smaller, sometimes temporary, water systems. 

Cyclestheriidae G.O. Sars,1899 

Cyzicidae Stebbing,1910 

Leptestheriidae Daday,1923 

Limnadiidae Baird,1849 


LAEVICAUDATA Linder,1945 

Lynceidae Stebbing,1910 


CLADOCERA Latreille,1829 

* Most species limnic. Some more limnic species, than the few treated here, may be found in inner Baltic 

archipelagos. 

EUCLADOCERA Eriksson,1934 

Sidoidea Baird,1850 (Brooks,1959) 

= Ctenopoda G.O. Sars,1865 

Sididae Baird,1850 

Penilia Dana,1852 

P. avirostris Dana,1852 

Last week of September and first weeks of October 1997 this species was very common in N Bohuslän, but 

its northernmost records else is the S Nort Sea (German Bight & off Rotterdam) 

Holopedidae G.O. Sars,1865 

* Only limnic species in the area. 

Chydoroidea Stebbing,1902 (Brooks,1959) 

= Daphnoidea Straus,1820 

= Anomopoda G.O. Sars,1865 

Bosminidae Baird,1845 

Bosmina Baird,1845 ( B. coregoni Baird,1857) 

B. (Eubosmina Seligo,1900) longispina Leydig,1860 maritima P.E. Müller,1867 

Gulf of Bothnia-Öresund-Kattegatt (around Laesoe)-S. Skagerrak (rare) 

Moinidae Goulden,1968 


Daphniidae Straus,1820 

Daphnia O.F. Müller,1785 

D. (Ctenodaphnia Dybowski & Grochowski,1895) magna Strauss,1820 ( of Ctenodaphnia D. & 

G.,1895 

Often in fresh-water rock-pools above the marine littoral, in Denm., Finl., Swe. & Norw. 

D. (Daphnia) longispina O.F. Müller,1785 

Fresh-brackish waters in Denm., Finl., Swe., Norw. & Icel. 

Mactrothricidae Norman & Brady,1867 

Ilyocryptinae Smirnov,1976 


Mactrothricinae Norman & Brady,1867 


Chydoridae Stebbing,1902 

Eurycercinae Kurz,1875 


Aloninae Frey,1967 


Chydorinae Stebbing,1902 


Polyphemoidea Baird,1845 (Brooks,1959) 

= Onychopoda G.O. Sars,1865 

Polyphemidae Baird,1845 

Polyphemus O.F. Müller,1776 

P. pediculus (Linnaeus,1761) 

= Monoculus pediculus Linnaeus,1761 () 

= Polyphemus oculus O.F. Müller,1785 

= Monoculus polyphemus Jurine,1820 

= Polyphemus stagnorum Fischer,1848 

= Polyphemus kochii Schoedler,1863 

Fresh-brackish waters in Denm., Finl., Swe., Norw. & Icel. 

Podonidae Mordukhai-Boltovskoi,1968 

Podon Liljeborg,1853 

P. polyphemoides Leuckart,1859 

* Mordukhai-Boltovskoi already in 1978 moved this species to the genus Pleopis Dana,1853 and Gieskes, 

W.W.C.,1971 also propose a removal of this species to Pleopis. 

Bothnian Sea-Bohuslän-North Sea-Norw. Sea-Br. Isles-Mediterranean & Black Sea 

P. leuckarti G.O. Sars,1862 

N Baltic-Bohuslän-North Sea-Norw. Sea-Barents Sea-Iceland-Br. Isles-Engl. Channel, Black Sea 

P. intermedius Liljeborg,1853 () 

Bothnian Sea-Bohuslän-North Sea-Norw. Sea-Iceland-Br. Isles-Mediterranean & Black Sea 

Evadne Lovén,1836 

E. nordmanni Lovén,1836 () 

Bothnian Sea-BohuslänNorth Sea-Norw. Sea-Barents Sea-Br. Isles-Mediterranean & Black Sea 

E. spinifera P.E. Müller,1868 

S Baltic-Bohuslän-North Sea-60° N-40° S 

E. tergestina Claus,1877 

Cental North Sea-southwards 

Cercopagididae Mordukhai-Boltovskoi,1966 

Cercopagis G.O. Sars,1897 

C. pengoi (Ostroumov,1891) 

* Native in fresh and brackish water in the Ponto-Caspian area, but succesful immigration into the Baltic was 

detected in 1992. 

Baltic 

HAPLOPODA G.O. Sars,1865 

Leptodoridae Lilljeborg,1900 

Leptodora Lilljeborg,1861 

42

L. kindti (Focke,1845) 

Fresh water in Denm., Finl., Swe. & Norw. & Baltic archipelagos 

CEPHALOCARIDA Sanders,1955 

* No European species so far recognized in this taxon with it's only recent order Brachypoda Birshtein,1960 

OSTRACODA Latreille,1806 

* Here considered to be a taxon separate from Maxillopoda, whithin which they sometimes are grouped 

OSTRACODA Latreille,1806 

= OSTRAPODA H.E. Straus,1821 

BRADORIIDA Matthew,1902 

* Extinct taxon. 

LEPERDITICOPIDA Scott,1961 

* Extinct taxon. 

PHOSPHATOCOPIDA K.J. Müller,1964 

* Extinct taxon 

PALAEOCOPA Hennigsmoen,1953 

PALAEOCOPIDA Henningsmoen,1953 

KIRKBYOCOPINA Gründel,1969 

Puncioidea Hornibrook,1949 

Punciidae Hornibrook,1949 

MYODOCOPA G.O. Sars,1866 

MYODOCOPIDA G.O. Sars,1866 

Cypridinoidea Baird,1850 

Cylindroleberididae Müller,1906 

Cylindroleberidinae Müller,1906 

Synasterope Kornicker,1975 ( Synasterope implumis Paulsen,1965) 

= Asterope : Auctt. 

= Synasterope Paulsen,1965 (nom. nud.) 

S. abyssicola (G.O. Sars,1870) 

* Correct genus? 

off Lofoten-off N- Norway 

S. norvegica (G.O. Sars,1869) 

* Kornicker,1987 attributes this species to his genus Priontoleberis Kornicker,1974 

Bohuslän (e.g. the Koster area)-W Norway-Trondheim-N North Sea-Shetlands-NE Scotland 

Parasterope Kornicker,1975 

= Parasterope Paulsen,1965 (nom. nud.) 

P. muelleri (Skogsberg,1920) 

= Asterope muelleri Skogsberg,1920 () 

=? Cypridina teres Norman,1861 

NE Atlantic-?Br. Isles-Mediterranean 

P. aberrata (Skogsberg,1920) 

Strangford Lough (N Ireland) 

Cylindroleberis Brady,1867 

=? Asterope Philippi,1840, non Müller & Troschel,1840 

= Asteropina Poulsen,196? 

C. mariae (Baird,1850) 

= Cypridina mariae Baird,1850 

SW Br. Isles 

Bruuniella Poulsen,1965 ( B. breviata Poulsen,1965 - off Thailand) 

Heptonema Cohen & Kornicker,1975 ( H. serratum Poulsen,1965 - Virgin Island) 

Diasterope Kornicker,1975 ( D. pilosa Poulsen,1965 - ?) 

Bathyleberis Kornicker,1975 ( B. grossmani Kornicker,1975 - ?Antarctic) 

Postasterope Kornicker,1986 ( P. abaco Kornicker,1986 - Bahamas) 

Archasterope Poulsen,1965 ( A. dentata Poulsen,1965 - S Oceans) 

Domromeus Kornicker,1989 ( D. heptathrix Kornicker,1989 - Biscay) 

Prionotoleberis Kornicker,1974 ( P. gyion Kornicker,1974 - off Mauretania) 

Cyclasteropinae Poulsen,1965 

* Not represented in the area 

Asteropteroninae Kornicker,1981 

* Not represented in the area 

Sarsiellidae G.S. Brady & A.M. Norman,1896 

Sarsiella Norman,1869 

S. capsula Norman,1869 () 

off Shetlands 

Eusarsiella Cohen & Kornicker,1975 

E. zostericola (Cushman,1906) 

Br. Isles (introduced) 

Cypridinidae Baird,1850 

Cypridininae Baird,1850 

Cypridini Baird,1850 

Doloria Skogsberg,1920 ( Cypridina (Doloria) levis Skogsberg,1920 - South Georgia) 

D. sarsi Kornicker,1987 

* Furca with 11 claws. 

Koster Channel in 180m 

Skogsbergia Poulsen,1962 

= Cypridina : Auctt., non H. Milne-Edwards,1840 


S. megalops (G.O. Sars,1871) 

off W Norway-Hardangerfjord 

Vargula Skogsberg,1920 

= Cypridina : Auctt., non H. Milne-Edwards,1840 


V. norvegica (Baird,1860) 

= Cypridina norvegica Baird,1860 () 

Koster Channel-W Skagerrak-N Norway-Shetlands-Faeroes-Iceland 

Macrocypridina Skogsberg,1920 

M. castanea (Brady,1897) 

= Cypridina castanea Brady,1897 () 

N Atlantic-56° N 

Gigantocypridini Hartmann,1974 

Gigantocypris G.W. Müller,1895 ( G. agassizii G.W. Müller,1895) 

G. muelleri Skogsberg,1920 

N Atlantic-64° N 

Azygocypridinae Kornicker,1970 

Azygocypridina Sylvester-Bradley,1950 ( Crossophorus imperator Brady,1880 (SW Pacific)) 

A. imperialis (Stebbing,1901) 

E Atlantic between 21°N-60°N at 1884-2930m 

Philomedidae G.W. Müller,1906 

Philomedinae G.W. Müller,1906 

Philomedes Liljeborg,1853 ( P. longicornis Liljeborg,1853) 

P. brenda (Baird,1850) 

= Cypridina brenda Baird,1850 

= P. globosus (Liljeborg,1853) 

Öresund-Bohuslän-North Sea-Barents Sea & Spitsbergen-Iceleand-Faeroes-Shetlands-Br. Isles 

P. lilljeborgi (G.O. Sars,1866) 

= P. bonneti Kornicker & Caraion,1977 

Kattegatt-Bohuslän-Skagerrak-Trondheim-Lofoten-Bjørnøja-Iceland-Faeroes, Biscay 

Euphilomedes Poulsen,1962 

E. interpuncta (Baird,1850) 

=? E. sinister Kornicker,1974? 

E Atlantic 30-65° N 

Pseudophilomedinae Kornicker,1967 

* Not recorded N of Biscay 

HALOCYPRIDA Dana,1853 

HALOCYPRIDINA Dana,1853 

Halocyprididae Dana,1852 

Conchoecinae 

Conchoecia Dana,1849 

C. (Boroecia Poulsen,1973) borealis G.O. Sars,1866 

Koster area-Lofoten-Arctic Seas, Iceland 

C. (Discoconchoecia Martens,1979) elegans G.O. Sars,1866 

N Kattegatt-Bohuslän-Lofoten-Br. Isles-Atlantic-80° N, Iceland 

C. (Conchoecilla Poulsen,1973) daphnoides (Claus,1890) 

Atlantic-60° N 

C. (Orthoconchoecia Poulsen,1973) haddoni Brady & Norman,1896 


C. (Conchoecissa Poulsen,1973) imbricata Brady,1880 

N Atlantic 

C. (Conchoecissa) ametra G.W. Müller,1906 

Iceland 

C. (Obtusoecia Martens,1979) obtusata G.O. Sars,1866 

Bohuslän-Trondheim-boreal-subarctic Atlantic 

C. (Metaconchoecia Poulsen,1973) rotundata G.W. Müller,1891 

= C. skogsbergi Iles,1953 

* Of this "species" 15 different forms are recognized by Angel & Fasham,1975, of which one, "C skogsbergi 

" : Angel,1968, non Iles,1953, is refferred to here. 

Norw. Sea 

C. (Conchoecia) lophura G.W. Müller,1906 

Iceland 

Euconchoecinae 

* The other genus of the subfamily, Euconchoecia Müller,1890 is not known from the area. 

Bathyconchoecia Deevey,1968 

B. darcythompsoni (Scott,1909) 

= Euconchoecia darcythompsoni Scott,1909 

NW Scotland 

Halocypridinae Dana,1852 

* The third genus of the subfamily, Fellia is not known from the area. 

Halocypria Claus,1874 

H. globosa Claus,1874 


Halocypris Dana,1852 

H. inflata Dana,1852 

= H. brevirostris Dana,18?? 

Atlantic, occasionally N to S & W Britain 

CLADOCOPIDINA G.O. Sars,1866 

43

Polycopoidea G.O. Sars,1866 

Polycopidae G.O. Sars,1866 

Polycope G.O. Sars,1866 

P. areolata G.O. Sars,1923 

Bohuslän-Korshavn (S Norway)-Plymouth 

P. clathrata G.O. Sars,1923 

off Väderöarna & Grisbådarna-Korshavn (S Norway)-Trondheim 

P. dentata Brady,1868 

?North Sea 

P. difficilis Klie,1936 

Helgoland 

P. elongata Hartmann,1954 

Sylt 

P. helgolandica Klie,1936 

Helgoland 

P. orbicularis G.O. Sars,1866 

Öresund-Skagerrak-Trondheim-SW Br. Isles 

P. punctata G.O. Sars,1870 

off Väderöarna-off Lofoten-Tromsö, ?Br. Isles 

P. pustulata G.O. Sars,1890 

W Norway 

P. schulzi Klie,1950 

Kieler Bucht 

P. sublaevis G.O. Sars,1923 

Bohuslän-Risör-Korshavn (S Norway)-Trondheim 

Parapolycope Klie,1936 

P. germanica Klie,1936 

Helgoland 

Polycopsis G.W. Müller,1894 

P. compressa (Brady & Robertson,1869) 

Korshavn (S Norway)-SW Br. Isles-Mediterranean 

PODOCOPA G.W. Müller,1894 

PLATYCOPIDA G.O. Sars,1866 

Cytherelloidea G.O. Sars,1866 

Cytherellidae G.O. Sars,1866 

Cytherella Jones,1850 

C. abyssorum G.O. Sars,1866 

Kattegatt-Bohuslän-Trondheim-Lofoten-Iceland-Br. Isles-?Mediterranean 

C. serrulata Brady & Norman,1896 

Britain 

PODOCOPIDA G.O. Sars,1866 

METACOPINA Sylvester-Bradley,1961 

Darwinulidoidea 

= Darwinulacea Brady & Norman,1889 

Darwinulidae Brady & Norman,1889 

Darwinula Brady & Norman,1889 

D. stevensoni (Brady & Robertson,1870) 

Tvärminne area, Stadil Fjord (oligohaline lagoon in W Jutland) 

Sigilioidea 

= Sigilidacea Mandelstam,1960 

* No European species. 

Saipanettidae McKenzie,1968 

PODOCOPINA G.O. Sars,1866 

Bairdioidea 

= Bairdiacea G.O. Sars,1888 

Bairdiidae G.O. Sars,1888 

Bairdiinae G.O. Sars,1888 

Bairdia McCoy,1844 

Neonesidea Maddocks,1969 

N. inflata (Norman,1862) 

= Cythere inflata Norman,1862 

= Bairdia inflata (Norman,1862) 

W Norway-N Norway-Br. Isles-NW France 

N. acanthigera (Brady,1868) 

= Bairdia acanthigera Brady,1868 

Engl. Channel 

Bythocyprididae Maddocks,1969 

Bythocypris Brady,1880 

B. bosquetiana (Brady,1866) 

W Norway-Trondheim-Shetlands-SW Br. Isles-Mediterran. 

B. obtusata (G.O. Sars,1866) 

S Norway (shells)-Bergen-Br. Isles 

Anchistrocheles Brady & Norman,1889 

A. acerosa (Brady,1868) 

= Cythere (?) acerosa Brady,1868 

Plymouth 

Cytheroidea Baird,1850 

= Cytheracea Baird,1850 

Limnocytheridae Klie,1938 

Limnocythere Brady,1868 

L. inopinata (Baird,1843) 

Baltic coasts 

Krithidae Mandelstam,1958 

Krithe Brady, Crosskey & Robertson,1874 

K. angusta Brady & Norman,1889? 

Skagerrak-North Sea 

K. praetexta (G.O. Sars,1866) 

= K. bartonensis : Auctt., non (Jones,1857) 

= Ilyobates praetexta G.O. Sars,1866 () 

Öresund-Bohuslän-Oslofjord-Lofoten-N Norway-Iceland-Shetlands-NE Britain 

K. producta Brady,1880 

Spitsbergen-? 

Paracyprideis Klie,1929 

P. fennica (Hirschmann,1909) 

Gävle & Tvärminne-S Baltic, Arctic Seas 

Cytherideidae G.O. Sars,1925 

Cytheridea Bosquet,1852 

C. angustata (Baird,1850) 

Öresund-Bohuslän-Helgoland-Holland-Br. Isles-?Iceland-Bay of Biscay-?Mediterranean 

Cyprideis Jones,1857 

= Anomocytheridea Stephenson,1938 

C. torosa (Jones,1850) 

= C. torosa teres Brady & Robertson,1870 

= C. litoralis (Brady,1868) 

= Cytheridea pedaschenkoi von Daday,1909 

Tvärminne-Öresund-Bohuslän-Oslofjord estuaries-Holland-E Br. Isles-France-Mediterranean 

Sarsicytheridea Athersuch,1982 

= Cytheridea sensu Sars,1925, non Bosquet,1851 

= Cytheridea (Eucytheridea) Bronstein,1930 

=? Cyamocytheridea Oertli,1956 

S. bradii (Norman,1865) 

= Cythere bradii Norman,1865 () 

= Cyprideis bairdii G.O. Sars,1866 

= Cytheridea papillosa : Auctt., non Bosquet,1852 

S Baltic-Bohuslän-North Sea-N Norway-Iceland & Spitzbergen-Novaja Zemlja-Faeroes-N Br. Isles 

S. punctillata (Brady,1865) 

= Cyprideis proxima G.O. Sars,1866 

Ålands Sea-Öres.-Bohuslän-North Sea-N Norway-Iceland & Arctic Seas-?Faeroes-N Br. Isles 

Neocytherideidae Puri,1957 

Neocytherideis Puri,1952 

= Sahnia Puri,1952 (p.p.) 

N. crenulata (Klie,1929) 

* Is this identical with N. subulata var. crenulata ? 

S Baltic-Belt Sea & Öresund, North Sea 

? N. baltica (Klie,1938) 

S Baltic 

N. subulata (Brady,1868) 

= Cytherideis subulata var. fasciata Brady & Robertson,1874 

= Cytherideis subulata var. crenulata Brady,1911 

= N. elongatus Puri,1952 () 

SW Br. Isles-W Scotland, W France 

Sahnicythere Athersuch,1982 

= Sahnia Puri,1952 (p.p.) 

S. retroflexa (Klie,1936) 

= Cytherideis retroflexa Klie,1936 () 

Helgoland-Amrum, Britain 

Heterocyprideis Elofson,1941 

H. sorbyana (Jones,1856) 

Gävle & Tvärminne-S Baltic-Bohuslän (shells)-W Norway (shells)-N Norway-Spitsbergen-Scotland (shells)- 

Ireland (shells) 

Cushmanideidae Puri,1974 

Pontocythere Dubowsky,1939 ( P. tschernjawskii Dubowski,1939) 

= Hemicytherideis Ruggieri,1952 

= Cytherideis sensu Puri,1952 

= Hulingsia Puri,1958 

P. elongata (Brady,1868) 

Skagerrak, SW Br. Isles-Thames estuary, N France 

Trachyleberididae Sylvester-Bradley,1948 

Buntonia Howe & Chambers,1935 

= Semicythereis Elofson,1943 

B. corpulenta (Brady & Norman,1889) 

Bergen-Trondheim 

Acanthocythereis Howe,1963 ( A. arenosa Howe,1963) 

= Trachyleberis : Auctt., non Brady,1898 

A. dunelmensis (Norman,1865) 

S Baltic-Öres.-Bohuslän-N Norway-Iceland & Arctic Seas-Faeroes-Shetlands-Br. Isles, E Greenland 

A. horrida (G.O. Sars,1866) 

* Elofsons (1941) larger form of Cythereis dunelmensis is probably this species sensu Athersuch & al.,1989. 

?Skagerrak 

44

Pterygocythereis Blake,1933 

P. jonesii (Baird,1850) 

= Cythereis jonesii Baird,1850 () 

Öresund-Bohuslän-SE North Sea-S & W Norway-Shetlands-Br. Isles-Bay of Biscay-Mediterranean-Black 

Sea 

P. mucronata (G.O. Sars,1866) 

Trondheim-off Lofoten-Shetlands 

Echinocythereis Puri,1954 

E. echinata (G.O. Sars,1866) 

Kattegatt-Koster Channel-Oslofjord-Lofoten-Atlantic 

E. mirabilis (Brady,1868) 

Bohuslän (shells)-Scotland (shells)-N Norway-Arctic Seas 

Cythereis Jones,1849 

C. leioderma (Norman,1869) 

Bohuslän (shells)-W Norway-Shetlands-Iceland-Arctic Seas 

Carinocythereis Ruggieri,1956 

C. carinata (Roemer,1838) 

= Cytherina carinata Roemer,1838 () 

= Cythereis antiquata Baird,1850 

W & SW Br. Isles, W France, Mediterranean 

C. whitei (Baird,1850) 

= Cythereis aspera Brady,1865 

= Carinocythereis bairdii Uliczny,1969 

S Britain, W France, Mediterranean 

Robertsonites Swain,1963 

R. tuberculata (G.O. Sars,1866) 

= Cythereis tuberculata G.O. Sars,1866 

= Robertsonites gubikensis Swain,1963 () 

= R. tuberculatina Swain,1963 

SW Baltic-Öres.-Bohuslän-N Norway-Spitsbergen-Iceland-Faeroes-Scotland & E England-Thames estuary 

Celtia Neale,1973 

C. quadridentata (Baird,1850) 

= Cythere quadridentata Baird,1850 () 

Risör (S Norway)-Hardangerfjord-Shetlands-Br. Isles-Bay of Biscay 

Costa Neviani,1928 ( Cytherina edwardsii Roemer,1838) 

C. runcinata (Baird,1850) 

S Britain, W France, W. Spain, Mediterranean-Turkey 

Hiltermannicythere Bassiouni,1970 ( Cythereis turbida G.W. Müller,1894) 

= Falunia Grekoff & Moyes,1956 (p.p.) 

= Falunia rugosa group Sissingh,1971 

H. emaciata (Brady,1868) 

SW Br. Isles, Cardigan Bay, Ireland 

Basslerites Howe,1937 

B. berchoni (Brady,1869) 

Arcachon-Mediterranean 

Eucytheridae Puri,1954 

Eucythere Brady,1868 

= Cytheropsis G.O. Sars,1866, non M'Coy,1849 

E. argus (G.O. Sars,1866) 

= E. undulata Klie,1929 

Kieler Bucht-Öres.-Bohuslän-Oslofjord-W & N Norway-Shetlands-SW Br. Isles-Ireland 

E. declivis (Norman,1865) 

= Cythere declivis Norman,1865 () 

=? Cytheropsis tenuitesta G.O. Sars,1866 

Belt Sea-Oslofjord-N Norway-Arctic Seas-SW Br. Isles 

E. anglica Brady,1868 

NE England 

E. prava Brady & Robertson,1869 

W Ireland, S England 

Paracytherideidae Puri,1957 

Paracytheridea G.W. Müller,1894 ( P. depressa G.W. Müller,1894) 

= Xenocythere G.O. Sars,1925 ( Cythere cuneiformis Brady,1868) 

P. cuneiformis (Brady,1868) 

Öresund & Belt Sea-Bohuslän-Bergen-Scotland-Ireland-Channel Islands 

Cytheridae Baird,1850 

Cythere O.F. Müller,1785 

= Cytherina Lamarck,1818 ( Cythere lutea O.F. Müller,1785) 

C. lutea O.F. Müller,1785 () 

= C. reniformis Baird,1835 

Kieler Bucht-Öres.-Bohuslän-Bergen-N Norway-Iceland & Arctic Seas-Shetlands-SW Br. Isles, W France 

C. globulifera Brady,1868 

* Not mentioned in S.B.F. (N.S.) 43, 1989. May be a synonym of another species? 

W Scotland, Roundstone 

C. acuta Baird,1850 

* Not mentioned in S.B.F. (N.S.) 43, 1989. May be a synonym of another species? 

Arran 

Palmenella Hirschmann,1916 

= Kyphocythere G.O. Sars,1925 ( Cythere limicola Norman,1865) 

P. limicola (Norman,1865) 

= Cythere limicola Norman,1865 () 

= Cythere nodosa G.O. Sars,1866 

= Cythere areolata Brady,1866 

S Baltic-Öres.-Bohuslän-Lofoten-Arctic Seas-N Br. Isles 

Cuneocytheridae Mandelstam,1959 

Cuneocythere Lienenklaus,1894 ( Cuneocythere truncata Lienenklaus,1894) 

C. semipunctata (Brady,1868) 

SW Br. Isles, Ireland, SW France 

Microcytheridae Klie,1938 

Microcythere G.W. Müller,1894 

M. bahusiensis Elofson,1944 

Bohuslän 

M. helgolandica Klie,1936 

Helgoland 

M. minuta Klie,1936 

Helgoland 

M. monstruosa Elofson,1944 

Bohuslän 

M. navicula (Norman,1869) 

Shetlands-SW Br. Isles 

M. parva Klie,1936 

Helgoland 

M. producta Elofson,1944 

Bohuslän 

M. schyttei Hartmann,1957 

Sylt 

Psammocytheridae Klie,1938 

Psammocythere Klie,1936 

P. remanei Klie,1936 

Helgoland 

Cobanocytheridae Schornikov,1975 

Cobanocythere Hartmann,1959 

C. angularis Gottwald & Scherner,1980 

S? North Sea 

C. syltensis Gottwald,1983 

Sylt 

Leptocytheridae Hanai,1957 

Leptocythere G.O. Sars,1925 

L. baltica Klie,1929 

Tvärminne-Öresund-Bohuslän-Sylt, Britain-Biscay 

L. castanea (G.O. Sars,1866) 

Dalarö & Tvärminne-Öresund-Bohuslän-Oslofjord-North Sea-W & N Norway-SW Br. Isles-Biscay, S 

Greenland 

L. confusa (Brady & Norman,1889) 

S Baltic 

L. lacertosa (Hirschmann,1912) 

Ålands Sea & Tvärminne-Öres.-Bohuslän-Holland, Britain, Biscay, Iceland, White Sea 

L. macallana (Brady & Robertson,1869) 

= Cythere levis G.W. Müller,1894 

Oslofjord-Skagerrak-Hardangerfjord-Holland-Scotland-W Ireland-SW Br. Isles-Mediterranean 

L. major Hartmann,19?? 

Öresund, German North Sea Coast 

L. pellucida (Baird,1850) 

= Cythere pellucida Baird,1850 () 

= Cythere confusa Brady & Norman,1889 

Ålands Sea & Tvärminne-Öres.-Bohuslän-Oslofjord-North Sea-N Norway-Shetlands-SW Br. Isles-Bay of 

Biscay-Mediterranean 

L. porcellanea (Brady,1869) 

= L. ilyophila (Hirschmann,1909) 

Dalarö & Tvärminne-S Baltic, SW Br. Isles, Biscay 

L. tenera (Brady,1868) 

S Baltic-Oslofjord-Risör-Hardangerfjord-Holland-Br. Isles-Bay of Biscay 

L. psammophila Guillaume,1976 

Baltic, Iceland, Britain, Roscoff, W France 

Callistocythere Ruggieri,1953 

C. badia (Norman,1861?) 

SW Br. Isles, Cyprus 

C. littoralis (G.W. Müller,1894) 

= Cythere littoralis G.W. Müller,1894 () 

= C. crispata : Auctt., non (Brady,1868) (Bradys species is restricted to the Mediterranean) 

Bohuslän-Oslofjord-S & W Norway-Ireland-Br. Isles-Mediterranean 

C. murrayi Whittaker,1978 

Wales, S England, Norfolk 

Hemicytheridae Puri,1953 

Heterocythereis Elofson,1941 

H. albomaculata (Baird,1838) 

=? Cythere alba Baird,1835 (nom. dub.) 

= Cythere albo-maculata Baird,1838 () 

Bohuslän-Helgoland-Hardangerfjord-N Norway-Iceland-Shetlands-SW Br. Isles-Bay of Biscay- 

Mediterranean, Azores 

Urocythereis Ruggieri,1950 ( Cytherina favosa Roemer,1838) 

U. britannica Athersuch,1977 

= Cythere oblonga Brady,1868, non Brady,1866, nec M'Coy,1844 

* The Mediterranean Cythere oblonga Brady,1866 is a junior objective homonym of M'Coy's species and 

was renamed U. distinguenda (Neviani,1928). 

Öresund-Skagerrak-S Norway-Engl. Channel-Wales-Bay of Biscay-N Spain 

45

Elofsonella Pokorny,1955 

= Paracythereis Elofson,1941, non Delachaux,1928 

E. concinna (Jones,1857) 

= Cythere concinna Jones,1857 () 

= Cythereis clavata G.O. Sars,1866 

Kieler Bucht-Öresund-Bohuslän-Bergen-Spitsbergen-Iceland-Faeroes-Shetlands-N Br. Isles 

Muellerina Bassiouni,1965 ( Cythere latimarginata Speyer,1863) 

M. abyssicola (G.O. Sars,1866) 

= Hemicythere latimarginata : G.O. Sars,1925, non (Speyer,1863) 

Bohuslän-Bergen-Lofoten-Spitsbergen-Iceland-Shetland area-N Britain 

Aurila Pokorny,1955 

= Auris Neviani,1928 (p.p.) 

= Eucythereis Klie,1940 (p.p.) 

=? Mutilus Neviani,1930 

A. convexa (Baird,1850) 

= Cythere convexa Baird,1850 () 

Bohuslän-Hardangerfjord-Shetlands-Br. Isles-Mediterraneamn 

A. arborescens (Brady,1865) 

= Cythere woodwardii Brady,1868 

Wales-Mediterranean 

A. woutersi Horne,1986 

S Britain 

Hemicythere G.O. Sars,1925 

= Auris Neviani,1928 (p.p.) 

= Eucythereis Klie,1940 (p.p.) 

H. costata (Brady,1866) 

Shetlands-Spitsbergen 

H. crenulata (G.O. Sars,1866) 

Koster Channel-S & W Norway-Shetlands-?Faeroes-Arctic Seas-Br. Isles 

H. emarginata (G.O. Sars,1866) 

Bohuslän-Bergen-Lofoten-N Norway-Arctic Seas-Iceland-Shetland-W Scotland (empty shells in Britain) 

H. villosa (G.O. Sars,1866) 

= Cythereis villosa G.O. Sars,1866 () 

Kieler Bucht-Öres.-Bohuslän-North Sea-N Norway-Iceland-Bear Island-Br. Isles-Bay of Biscay 

H. rubida (Brady,1868) 

W Scotland, S England, Ireland 

Procythereis Skogsberg,1928 

?P. marginata (Norman,1861?) 

SW Br. Isles 

Finmarchinella Swain,1963 

F. (Finmarchinella) finmarchica (G.O. Sars,1866) 

Korshavn (S Norway)-N Norway-S North Sea-Br. Isles (empty shells in Britain) 

F. (Barentsova ?,19??) angulata (G.O. Sars,1866) 

S Baltic-Öres.-Bohuslän-Oslofjord-N Norway-Iceland-Arctic Seas-Ireland-Br. Isles (empty shells in Britain) 

Baffinicythere ?,19?? 

B. howei Hazel,1??? 

Britain (empty shells in Britain) 

Cythereis Jones,1849 

C. navicula (Norman,18??) 

= Cythere navicula Norman,18?? 

= Cythere fidicula Brady & Robertson,1889 

* Not mentioned in S.B.F. (N.S.), No. 43, 1989. May possibly be a synonym of another species? 

Bergen, Shetlands, Northumberland, Firth of Forth, Maas & Schelde estuaries 

Xestoleberididae G.O. Sars,1928 

Xestoleberis G.O. Sars,1866 

X. nitida (Liljeborg,1853) 

= Cythere nitida Liljeborg,1853 () 

= X. aurantia : G.O. Sars,1928, non (Baird,1838) 

SW Finland & Gräsö-Öres.-Bohuslän-North Sea-Bergen-N Norway-Shetlands-SW Br. Isles-Arcachon 

X. aurantia (Baird,1838) 

= X. pusilla Elofson,1941 

Bohuslän-Bergen, Britain, Bretagne 

X. depressa G.O. Sars,1866 

S Baltic-Öres.-Bohuslän-Bergen-Lofoten-Arctic Seas-Shetlands-Br. Isles-Bay of Biscay-?Mediterranean 

X. labiata Brady & Robertson,1874 

SW Br. Isles, N France 

X. rubens Whittaker,1978 

Dorset, W France 

Microxestoleberis G.W. Müller,1894 

M. nana G.W. Müller,1894 

Helgoland-Mediterranean 

M. rostrata Klie,1936 

Helgoland 

Loxoconchidae G.O. Sars,1925 

Hirschmannia Elofson,1941 

H. viridis (O.F. Müller,1785) 

= Cythere viridis O.F. Müller,1785 () 

= Cythere cyamos Norman,1905 

= Loxoconcha sarsii Hirschmann,1912 

non = Cythere viridis : Lilljeborg,1853 (which is = Loxoconcha elliptica Brady,1868) 

Tvärminne-Öresund-Bohuslän-Helgoland-Bergen-Murmansk-SW Br. Isles-SW France 

Cytheromorpha Hirschmann,1909 

C. claviformis Hirschmann,1909? 

?S Baltic? 

C. fuscata (Brady,1869) 

Dalarö & Tvärminne-Öresund, Stadil Fjord (oligohaline lagoon in W Jutland), Drammen, Holland river 

mouths, Thames estuary (now only empty shells in Britain) & Caspian Sea 

C. n.sp. A. Rosenfeld,1977 

Belt Sea 

Bonnyannella Athersuch,1982 

B. robertsoni (Brady,1868) 

= Cythere robertsoni Brady,1868 () 

Belt Sea-Oslofjord-SW Br. Isles-W France 

Elofsonia Wagner,1957 

E. baltica (Hirschmann,1909) 

= Loxoconcha baltica Hirschmann,1909 () 

= Cythere gibbosa Brady & Robertson,1869, non H. Milne-Edwards,1840 

Dalarö & Tvärminne-Öres.-Holland, N Norway, Britain-W Ireland-Arcachon 

E. pusilla (Brady & Robertson,1870) 

= Loxoconcha pusilla Brady & Robertson,1870 

Tvärminne-Öresund-Bohuslän-Sylt-Holland-Br. Isles 

Palmoconcha Swain & Gilby,1974 ( P. laevimarginata Swain & Gilby,1974) 

= Lindisfarnia Horne & Kilenyi,1981 ( Cythere laevata Norman,1865) 

P. guttata (Norman,1865) 

= Cythere impressa Baird,1850, non M'Coy,1844 

= Cythere guttata Norman,1865 

= Loxoconcha granulata G.O. Sars,1866 

Kieler Bucht-Öres.-Bohuslän-N Norway-Br. Isles-Ireland 

P. laevata (Norman,1865) 

= Cythere laevata Norman,1865 

= Loxoconcha longipes G.O. Sars,1866 

= Loxoconcha tamarindus : Auctt., non (Jones,1856) 

S Baltic-Öres.-Bohuslän-N Norway-Iceland-Shetlands-Br. Isles & Ireland-Bay of Biscay 

Loxoconcha G.O. Sars,1866 

= Normania Brady,1866 (published 3 months after Sars' name) 

L. emelwardensis Redeke,1936 

S North Sea 

L. elliptica Brady,1868 

= L. gauthieri Klie,1929 

Dalarö & Finland-Öres.-North Sea-Br. Isles & ?Ireland, all around Mediterranean 

L. helgolandica Klie,1929 

Helgoland 

L. rhomboidea (Fischer,1855) 

= L. impressa : Brady,1868, non Cythere impressa Baird,1850, nec M'Coy,1844 

= Cythere rhomboidea Fischer,1855 () 

= Cythere (Normania) carinata Brady,1865 

S Baltic-Öres.-Bohuslän-North Sea-N Norway-Shetlands-Br. Isles-W France-Mediterranean-Madeira & 

Canaries 

Sagmatocythere Athersuch,1976 ( Loxoconcha napoliana Puri,1963) 

S. multiflora (Norman,1866) 

= Loxoconcha multiflora : Elofsson,1938 

Koster Channel-Holland-Trondheim-Shetlands-around Br. Isles-Bay of Biscay 

Phlyctocythere Keij,1958 

P. fragilis (G.O. Sars,1866) 

= Loxoconcha f. : Auctt. 

Kieler Bucht-Belt Sea-Bohuslän-Oslofjord-Lofoten-Br. Isles-W Ireland 

Nannocythere Schäfer,1953 

N. pavo (Malcolmson,1886) 

= N. remanei Schäfer,1953 () 

Helgoland, Engl. Channel, E Ireland, W Scotland 

Tuberoloxoconcha Hartmann,1973 

*The British records may possibly be an undescibed species of the genus. 

T. tuberosa (Hartmann,1954) 

?Britain 

Bythocytheridae G.O. Sars,1866 

Pseudocythere G.O. Sars,1866 

P. caudata G.O. Sars,1866 () 

Bohuslän-Oslofjord-Trondheim-N Norway-Arctic Seas-Shetlands-Br. Isles-?Bay of Biscay 

P. britannica Horne,1986 

NE England, Isles of Scilly 

P. norvegica Horne,1986 

Norway 

Jonesia Brady,1866 

= Macrocythere G.O. Sars,1926 

J. acuminata (G.O. Sars,1866) 

= Cythere simplex Norman,1865 (), non Cythere auriculata var. simplex Cornuel,1848 

S Baltic-Öres.-Bohuslän-North Sea-Lofoten-Troms-Arctic Seas-Iceland-Faeroes-Shetlands-Br. Isles, W 

Greenland 

Bythocythere G.O. Sars,1866 

B. bradyi G.O. Sars,1926 

Stavanger-SW Br. Isles 

B. constricta G.O. Sars,1866 

Oslofjord-Skagerrak-W & N Norway-Spitsbergen 

B. dromedaria G.O. Sars,1866 

Koster Channel-Oslofjord-Trondheim 

B. insignis G.O. Sars,1870 

off Lofoten 

B. intermedia Elofson,1938 

Bohuslän-S Norway, S North Sea, Britain-Biscay 

B. turgida G.O. Sars,1866 () 

46

Koster Channel-Oslofjord-S & W Norway-Spitsbergen-?Shetlands-Ireland-?Bay of Biscay-?Mediterranean 

B. bradleyi Athersuch, Horne & Whittaker,1983 

= B. turgida : Sylvester-Bradley,1947, non G.O. Sars,1866 

SW Ireland 

B. robinsoni Athersuch, Horne & Whittaker,1983 

= B. turgida : Brady,1868 & 1870, non G.O. Sars,1866 

Britain 

B. zetlandica Athersuch, Horne & Whittaker,1983 

= B. constricta : Brady,1868 (p.p.), non G.O. Sars,1866 

Shetlands, N Britain 

Sclerochilus G.O. Sars,1866 

S. contortus (Norman,1861) 

= Cythere contorta Norman,1861 () 

Cardigan Bay, Engl. Channel 

S. abbreviatus Brady & Robertson,1869 

W Ireland, W Britain 

S. bradyi Rudjakov,1962 

NE England, Shetlands, Clyde Sea 

S. rudjakovi Athersuch & Horne,1987 

S Baltic-Öres-Bohuslän-S North Sea-White Sea, Shetlands, Britain 

S. gewemuelleri Dubowsky,1939 

NE & S England, Wales, Mediterranean, Black Sea 

S. hicksi Athersuch & Horne,1987 

NE England 

S. schornikovi Athersuch & Horne,1987 

W Scotland, Bristol Channel 

S. truncatus (Malcolmson,1886) 

= Paradoxostoma truncatum Malcolmson,1886 

NE England, N Wales 

S. whatleyi Athersuch & Horne,1987 

W Scotland 

S. n.sp. : Athersuch & al.,1989 

= S. contortus : G.O. Sars,1926, non (Norman,1861) 

Norway-Spitzbergen 

Paradoxostomatidae Brady & Norman,1889 

Cytherois G.W. Müller,1884 

C. arenicola Klie,1929 

Tvärminne-Kattegatt-S North Sea, White Sea 

C. fischeri (G.O. Sars,1866) 

= Sclerochilus (?) gracilis Brady & Robertson,1869 

= Cytherois virens G.W. Müller,1884 () 

?S Finland-S Baltic-Öres.-Bohuslän-S North Sea-S & ?W Norway-Br. Isles-W France-Mediterranean 

C. pusilla G.O. Sars,1928 

S Baltic-Öresund-Bohuslän-SW Norway-North Sea-Bergen-Britain-NW France 

C. vitrea (G.O. Sars,1866) 

Kieler Bucht-Öresund-Bohuslän-Oslofjord-North Sea-Lofoten-Shetlands-Faeroes-SW Br. Isles 

C. stephanidesi Klie,1938 

Britain (Cristchurch Harbour)-SW France-Mediterranean 

C. n.sp. : Athersuch & al.,1989 

Britain 

Paradoxostoma Fischer,1855 ( P. dispar Fischer,1855) 

= Xiphichilus Brady,1870 ( Bythocythere tenuissimum Norman,1869) 

P. abbreviatum G.O. Sars,1866 

S Baltic-Öres.-Bohuslän-Holland-Bergen-Shetlands-Br. Isles-N France-?Bay of Biscay 

P. robinhoodi Horne & Whittaker,1985 

= P. normani : Wagner,1957, non Brady,1868 

Britain 

P. bradyi G.O. Sars,1928 

= P. obliquum : Brady,1868, non G.O. Sars,1866 

= P. hibernicum : G.O. Sars,1928, non Brady,1868 

Bohuslän-Norway-Faeroes-Shetlands-SW Br. Isles-Ireland-N France 

P. ensiforme Brady,1868 

Bohuslän-Holland-?S & W Norway-Shetlands-SW Br. Isles-N France-?Bay of Biscay 

P. fleetense Horne & Whittaker,1985 

S England, Channel Islands 

P. angliorum Horne & Whittaker,1985 

= P. ensiforme Brady,1868 (p.p.) 

E Anglia & Thames estuary 

P. hibernicum Brady,1868 

SW Br. Isles-NW Ireland 

P. normani Brady,1868 

Öresund & Belt Sea-Bohuslän-SW Norway-S North Sea-Faeroes-Shetlands-Br. Isles-?Bay of Biscay 

P. nealei Horne & Whittaker,1985 

around English & Irish coasts 

P. obliquum G.O. Sars,1866 

Lofoten-N Norway 

P. orcadense Brady & Robertson,1874? 

SW Br. Isles 

P. pulchellum G.O. Sars,1866 

Öresund-Bohuslän-Oslofjord-Bergen-SW Br. Isles-Ireland 

? P. rostratum G.O. Sars,1866 

N Norway 

P. sarniense Brady,1868 

W Scotland-SW England-Channel Islands-Normandie-Bretagne 

47 

P. variabile (Baird,1850) 

Dalarö-Öres.-Bohuslän-North Sea-Bergen-N Norway-White-Sea-Iceland-Shetlands-Br. Isles-N Bretagne 

P. arcticum Elofson,1941 

Arctic Seas 

P. tenuissimum (Norman,1869) 

= Bythocythere tenuissimum Norman,1869 

= Xiphichilus tenuissimus : Brady,1870 

= Machaerina tenuissima : Brady & Norman,1889 

off Scotland at 50-100 m 

P. trieri Horne & Whittaker,1985 

Britain-SW France 

P. porlockense Horne & Whittaker,1985 

around Britain 

P. amygdaloides (Brady,1870) 

* Not mentioned in S.B.F. (N.S) 43, 1989 

Shetlands, Biscay, Mediterranean 

Paracytherois G.W. Müller,1894 ( P. striata G.W. Müller,1894) 

* This genus is poorly known, but there seems to be at least 4 British species sensu Athersuch & al.,1989. 

P. arcuata (Brady,1868) 

Kieler Bucht-Öresund-Bohuslän-Bergen-Iceland-Shetlands-Br. Isles 

P. flexuosa (Brady,1867) 

= Bythocythere? flexuosa Brady,1867 

=? P. striata G.W. Müller,1894 

? non P. flexuosa (Brady) sensu G.O. Sars,1928 

?Bohuslän-?W Norway-?Holland-SW Br. Isles-?Bay of Biscay-?Mediterranean 

P. producta (Brady & Norman,1889) 

Bergen 

Aspidoconcha de Vos,1953 

A. limnoriae de Vos,1953 


?NE Atlantic 

Sphaeromicola Paris,1916 

S. dudichi Klie,1938 


?NE Atlantic 

Cytheruridae G.W. Müller,1894 

Cytheropterinae G.O. Sars,1866 

Cytheropteron G.O. Sars,1866 

= Kobayashiina Hanai,1957 

= Lobosocythoropteron Ishizaki & Gunther,1974 

C. alatum G.O. Sars,1866 

Bohuslän-Oslofjord-Risör (S Norway)-Trondheim-Shetlands-N Br. Isles & Ireland 

C. angulatum (Brady,1868) 

Öresund-Risör (S Norway)-Br. Isles-Spitsbergen 

C. crassipinnatum Brady & Norman,1889 

Risör (S Norway)-W Ireland 

C. depressum Brady & Norman,1889 

SW Br. Isles 

C. hamatum G.O. Sars,1870 

Stoksund (W Norway)-Lofoten-Spitsbergen, Britain 

C. inflatum Brady,186? 

Korshavn (S Norway)-Br. Isles 

C. latissimum (Norman,1865) 

= Cythere latissima Norman,1865 () 

= Cytheropteron convexum G.O. Sars,1866 

S Baltic-Öres.-Bohuslän-North Sea-N Norway-Spitsbergen-Iceland-Shetlands-SW Br. Isles 

C. nodosum Brady,1868 

Korshavn (S Norway)-W Norway-Spitsbergen-Br. Isles-Bay of Biscay 

C. punctatum Brady,1868 

Korshavn (S Norway)-Bergen-Br. Isles-Bay of Biscay 

C. pyramidale Brady,1868 

Bohuslän-Oslofjord-N Norway-Arctic Seas 

C. subcircinatum G.O. Sars,1866 

Risör (S Norway)-Hardangerfjord 

C. testudo G.O. Sars,1870 

Koster Channel-Hardangerfjord-Lofoten-Spitsbergen 

C. inornatum Brady & Robertson,1872 

NW Britain 

C. dorsocostatum Whatley & Masson,1980 

S Norway-around Britain-NW Spain 

C. volantium Whatley & Masson,1980 

off Britain 

C. humile Brady & Norman,18?? 

* not mentioned in SoBF (NS) 43. 

@ burrows of Limnoria & Chelura 

?Britain 

Cytherurinae G.W. Müller,1894 

Hemicytherura Elofson,1941 

H. cellulosa (Norman,1865) 

= Cythere cellulosa Norman,1865 (p.p.) () 

= Cytherura nana G.O. Sars,1866 

Kieler Bucht-Öres. & Belt Sea-Bohuslän-North Sea-S Norway-Bergen-Faeroes-Shetlands-all around Br. 

Isles-Bay of Biscay 

H. hoskini Horne,1981 

= Cythere cellulosa Norman,1865 (p.p.) 

W Ireland, Bristol Channel

H. clathrata (G.O. Sars,1866) 

= Cytherura clathrata G.O. Sars,1866 

= Cytheropteron clathratum : Auctt. 

Öresund-North Sea-W Br. Isles-Trondheim-Lofoten-Spitsbergen-Br. Isles 

Cytherura G.O. Sars,1866 

C. gibba (O.F. Müller,1785) 

= Cythere gibba O.F. Müller,1785 (female) () 

= Cythere gibbera O.F. Müller,1785 (male) 

= Cytherura robertsoni Brady,1868 

SW Finland-Öres.-Skagerrak-Oslofjord-North Sea-around Br. Isles 

C. atra G.O. Sars,1866 

S Baltic-Lofoten-Shetlands 

C. undata G.O. Sars,1865 

Kieler Bucht-Bohuslän, Oslofjord-Kola peninsula, Iceland, Spitsbergen, Franz Joseph Land, Britain, Ireland 

C. concentrica Brady, Crosskey & Robertson,1874 

* Not mentioned in S.B.F. (N.S.) 43, 1989 

Bergen, Hardangerfjord, Florø, Shetlands, Northumberland, Yorkshire, Durham, Spitsbergen 

Microcytherura G.W. Müller,1894 

M. affinis Klie,1938 

S Baltic 

M. fulva (Brady & Robertson,1874) 

S Baltic-Öres.-Bohuslän-S Norway-North Sea-SW Br. Isles-Bay of Biscay 

Semicytherura Wagner,1957 

S. acuticostata (G.O. Sars,1866) 

S Baltic-Bohuslän-Holland-Trondheim-Shetlands-around Br. Isles 

S. affinis (G.O. Sars,1866) 

W Norway-Spitsbergen 

S. angulata (Brady,1868) 

= Cytherura insolita Brady,1870 

Kieler Bucht-Belt Sea-Bohuslän-Holland-Hardangerfjord-Iceland-Shetlands-SW Br. Isles 

S. cornuta (Brady,1868) 

= Cytherura lineata Brady,1868 

= Cytherura intumescens G.O. Sars,1925 

Risör (S Norway)-Br. Isles & Ireland, Bretagne 

? S. bidens (Klie,1929) 

Kieler Bucht-Öresund, Helgoland 

? S. brevifistulosa (Klie,1936) 

Helgoland 

? S. cochlearis (Klie,1929) 

Kieler Bucht 

S. concentrica (Brady, Crosskey & Robertson,1874) 

off Norwegian coast 

S. exserta (Brady & Norman,1889?) 

off Norwegian coast 

S. inconspicua (Klie,1934) 

Kieler Bucht-Öresund-Bergen 

? S. nasuta (Klie,1934) 

Kieler Bucht 

S. nigrescens (Baird,1838) 

= Cythere nigrescens Baird,1838 () 

Tvärminne-Öres.-Bohuslän-North Sea-Bergen-N Norway-Br. Isles-Bay of Biscay, SW Greenland 

S. producta (Brady,1866) 

Korshavn (S Norway)-W Norway-Holland-SW Br. Isles 

S. sella (G.O. Sars,1866) 

= Cytherura cuneata Brady,1868 (male) 

= Cytherura flavescens Brady,1869 (female) 

S Baltic-Öres.-Skagerrak-Oslofjord-North Sea-W Norway-W Br. Isles-Bay of Biscay 

S. similis (G.O. Sars,1866) 

S Baltic-Öres.-Bohuslän-Oslofjord-Helgoland & Holland-N Norway-Shetlands-SW Br. Isles-?Bay of Biscay 

S. simplex (Brady & Norman,1889) 

= Cytherura hamata Klie,1936 

S Baltic-SW Br. Isles, W. Ireland, Helgoland 

S. striata (G.O. Sars,1866) 

= Cytherura quadrata Norman,1869 

S Baltic-Öres.-Bohuslän (shells)-Oslofjord-Holland-W Norway-Faeroes-Shetlands-Br. Isles-Channel Islands- 

Biscay-?Mediterranean 

S. undata (G.O. Sars,1866) 

= Cytherura humilis Brady,1867 

= Cytherura pumila Brady,1874 

S Baltic-Öres.-Bohuslän-North Sea-Lofoten-Spitsbergen-Shetlands-N Br. Isles 

S. tela Horne & Whittaker,1980 

S Britain, Arcachon 

Cytheromatidae Elofson,1939 

Cytheroma G.W. Müller,1894 

C. latiantennata Elofson,1938 

Bohuslän 

Terrestricytheroidea Schornikov,1969 

* Known from NW Pacific isles only 

Cypridoidea 

= Cypridacea Baird,1845 

Macrocyprididae G.W. Müller,1912 

Macrocypris Brady,1868 

M. minna (Baird,1850) 

= Cythere minna Baird,1850 () 

Kattegatt-Koster Channel-N North Sea-Lofoten-Shetlands 

Macrocypria G.O. Sars,1923 

M. angusta (G.O. Sars,1866) 

Kattegatt-Koster Channel-Oslofjord-Trondheim 

Pontocyprididae G.W. Müller,1894 

Argilloecia G.O. Sars,1866 

A. conoidea G.O. Sars,1923 

Bohuslän-Risör (S Norway) 

A. cylindrica G.O. Sars,1866 

Bohuslän-Oslofjord-Hammerfest-SW Br. Isles 

A. propinqua Brady,1868? 

off Britain 

Propontocypris Sylvester-Bradley,1947 

P. trigonella (G.O. Sars,1866) 

= Pontocypris trigonella G.O. Sars,1866 () 

Koster Channel-Lofoten-N Norway-Shetlands-SW Br. Isles-?Mediterranean-?Cape Verde 

P. pirifera (G.W. Müller,1894) 

Scotland, S England, W France, Mediterranean 

Pontocypris G.O. Sars,1866 (p.p.), non G.W. Müller,1912, nec G.O. Sars,1923 

= Erythrocypris G.W. Müller,1894 

P. hispida G.O. Sars,1866 

Bohuslän-Oslofjord-Risör (S Norway)-Trondheim-Shetlands-Br. Isles 

P. mytiloides (Norman,1862) 

= Cypris serrulata G.O. Sars,1863 () 

= Cythere avena Norman,1865 

Korshavn & Risör (S Norway)-Möre-Br. Isles-Mediterranean 

P. pallida (G.W. Müller,1894) 

Korshavn & Risör (S Norway)-Mediterranean 

P. acupunctata Brady,1866 

* Not mentioned in S,B.F. (N.S.) 43,1989 

Florø, Shetlands, Northumberland, Firth of Forth, Durham 

P. frequens (G.W. Müller,1???) 

* Not mentioned in S,B.F. (N.S.) 43,1989 

?Mediterranean, ?Britain 

Cypridopsidae Kaufmann,1910 

Cypridopsis Brady,1868 

C. aculeata (Costa,1852) 

Finnish archipel.-Norwegian estuaries-SW Br. Isles 

C. vidua (O.F. Müller,1785) 

Scandinavian freshwater-Thames estuary 

C. newtoni Brady & Robertson,1870 

Finnish rock-pools 

Potamocypris Brady,1870 

P. villosa (Jurine,1820) 


P. humilis (G.O. Sars,18??) 


Ilyocyprididae Kaufmann,1900 

Ilyocypris Brady & Norman,1889 

I. gibba (Rahmdohr,1808) 

Freshwater in Sw., Norw., Denm. & Germ., estuaries in Arcachon-Mediterranean-Black Sea 

Cyprididae Baird,1845 

Cyprinotinae Bronstein,1947 

Cyprinotus Brady,1886 

C. incongruens (Ramdohr,1808) 

Baltic rock-pools 

C. salinus (Brady,1868) 

Baltic rock-pools-, Oslofjord estuaries 

C. reptans (Baird,1835) 

Fresh-water species, which can be found in North Sea "weak brackish water" 

Heterocypris Claus,1892 

H. salinus (Brady,1868) 

= Cypris salina Brady,1868 

Baltic rock pools, coastal N Europe, Britain (in salt water pools (0.4-13 ppt S) & salt marshes) 

Candonidae Kaufmann,1900 

Paracypridinae G.O. Sars,1923 

Paracypris G.O. Sars,1866 

P. polita G.O. Sars,1866 () 

Bohuslän-Trondheim-Shetlands-Br. Isles-Bay of Biscay-?Mediterranean 

Candoninae Kaufmann,1900 

Candona Baird,1845 

C. angulata G.W. Müller,1900 

Fresh-water species, which can be found in Baltic "weak brackish water" 

C. candida (O.F. Müller,1785) 

Fresh-water species, which can be found in Baltic "weak brackish water", e.g. Örnsköldsvik 

C. compressa (Koch,1837) 

Fresh-water species, which can be found in Baltic "weak brackish water", e.g. Stockholm archipelago 

48

C. neglecta G.O. Sars,1887 

Fresh-water species, which can be found in Baltic brackish water, e.g. Ålands Sea & Central Baltic; Stadil 

Fjord (oligohaline lagoon in W Jutland) 

C. pratensis Hartwig,1901 


C. sucki Hartwig,1901 

Fresh-water species, which can be found in North Sea "weak brackish water" 

Cyclocypris Brady & Norman,1889 

C. ovum (Jurine,1820) 

Fresh-water species, which can be found in Baltic "weak brackish water", e.g. Örnsköldsvik 

C. laevis (O.F. Müller,1791) 

Fresh-water species, which can be found in Baltic "weak brackish water", eg. Dalarö 

C. globosa (G.O. Sars,1863) 


Cypria Zenker,1854 

C. ophtalmica (Jurine,1820) 

Fresh-water species, which can be found in Baltic "weak brackish water", e.g. Dalarö 

C. exsculpta (Fischer,1855) 


MALACOSTRACA Latreille,1806 

HOPLOCARIDA Calman,1904 

STOMATOPODA Latreille,1817 

UNIPELTATA Latreille,1825 

Squilloidea Latreille,1803 

Squillidae Latreille,1803 

Meiosquilla Manning,1977 

M. desmaresti (Risso,1816) 

S Br. Isles-Mediterranean 

Lysiosquilloidea Giesbrecht,1910 

Nannosquillidae Manning,1980 

Platysquilla Manning,1977 

P. eusebia (Risso,1816) 

S Br. Isles-Mediterranean 

EUMALACOSTRACA Grobben,1892 

SYNCARIDA Packard,1885 

* no marine species in the area. 

(Anaspidacea Calman,1904 and Stygocarididae Noodt,1963 are confined to the S hemispere & 

Bathynellacea Chappuis,1915 is confined to Limno-terrestrial habitats) 

EUPHAUSIACEA Dana,1852 

Bentheuphausiidae Colosi,1917 

Bentheuphausia G.O. Sars,1885 

B. amblyops G.O. Sars,1885 

Atlantic, usually below 40° N 

Euphausiidae Dana,1852 

Euphausia Dana,1850 

E. krohnii (Brandt,1851) 

off W Norway-off Iceland-E Shetlands-NE Scotland-off W Br. Isles-southwards 

Meganyctiphanes Holt & Tattersall,1905 

M. norvegica (M. Sars,1857) 

Katteg.-Bohuslän-Barents Sea-Spitsbergen-Greenland-W Br. Isles-Azores-Mediterranean 

Nyctiphanes G.O. Sars,1884 

N. couchii (Bell,1853) 

?S Baltic-W Skagerrak-Möre-Shetlands-W Br. Isles-Azores-Mediterranean 

Thysanoessa Brandt,1851 

T. inermis (Krøyer,1846) 

Katteg.-Bohuslän-N North Sea-Barents Sea-E Greenland-off W Br. Isles-south to 50° N 

T. raschii (M. Sars,1864) 

Katteg.-Bohuslän-N North Sea-N Scotland-Iceland-Greenland-Spitsbergen & Kara Sea-Arctic Seas 

T. longicaudata (Krøyer,1846) 

Koster Channel, Skagerrak-N North Sea-Barents Sea-E Greenland-off W Br. Isles-off Portugal 

T. gregaria G.O. Sars,1883 

Atlantic, between 20-40° N & W Mediterranean 

Thysanopoda Milne-Edwards,1830 

T. acutifrons Holt & Tattersall,1906 

W-SW Norway-Shetland-Faeroe Channel-Iceleand-off W Ireland-off W Gibraltar 

Nematoscelis G.O. Sars,1884, non Wollaston,1867 

* Does a proper name exist? 

N. megalops G.O. Sars,1884 

S Greenland-S Iceland-S & N Norway.-North Sea-off W Br. Isles-Azores 

Nematobrachion Calman,1905 

N. boopis (Calman,1896) 

S Greenland-S Iceland-Faeroe-Shetland Channel-off W Br. Isles-NW Africa 

Stylocheiron G.O. Sars,1884 

S. elongatum G.O. Sars,1884 

S Iceland-N, W & SW Br. Isles-southwards 

S. longicorne G.O. Sars,1884 

S Iceland-Bergen-off W Br. Isles-southwards 

S. maximum Hansen,1908 

off SE Greenland-off S Iceland-Faeroes-Shetlands-off W Ireland-southwards 

S. abbreviatum G.O. Sars,1884 

NW Norway-Barents Sea-Iceland-SE Greenland-Faeroe-Shetland Channel-off W Ireland 

AMPHIONIDACEA Williamson,1973 

* With a circumglobal marine planktonic species, Amphionides reynaudii (H. Milne-Edwards) as only 

species. (Not in N Europe) 

DECAPODA Latreille,1803 

DENDROBRANCHIATA Bate,1888 

* This subordo has been grouped together with Stenopodidea, Procarididea & Caridea in the next subordos, 

together constituting the older subordo Natantia Boas,1880 

Penaeoidea Rafinesque-Schmaltz,1815 

Penaeidae Rafinesque-Schmaltz,1815 

Penaeinae Rafinesque-Schmaltz,1815 

Tribe: Peneini Burkenroad,1936 

Penaeus Fabricius,1798 

P. kerathurus (Forskål,1775) 

* "Caramote prawn or shrimp", "Tiger shrimp", "Striped shrimp", "Langostino" 

Mediterranean, E Atlantic up to W Engl. Channel 

P. semisulcatus W. De Haan, in Siebold,1849 

E Atlantic 

Funchalia Johnson,1867 

= Pelagopenaeus Burkenroad,1934 

F. woodwardi Johnson,1867 

Hebrides-southwards in Atlantic, Mediterranean 

F. balboae (Faxon,1893) 

= Pelagopenaeus balboae (Faxon,1893) 


Heteropeneus De Man,1896 

Tribe: Parapeneini Burkenroad, in Schram,1983 

Metapenaeopsis Bouvier,1905 

M. miersi Holthuis,1952 

E Atlantic off W Africa 

Parapenaeus Smith,1885 

P. longirostris (H. Lucas,1846) 

= Peneus longirostris H. Lucas,1846 

* "Deep water rose shrimp", "Gamba" 

E Atlantic up to N Spain, Mediterranean 

Penaeopsis Bate,1881 

= Peneopsis Faxon,1895 

Artemesia Bate,1888 

Tribe: Trachypeneini Burkenroad, in Schram,1983 

Parapenaeopsis Alcock,1901 

P. atlantica Balss,1926 

E Atlantic off W Africa 

Mangalura Miers,1878 

Macropetasma Stebbing,1914 

Trachypeneopsis Burkenroad,1934 

Atypopeneus Alcock,1905 

Protrachypene Burkenroad,1934 

Xiphopeneus S.I. Smith,1886 

Trachypeneus Alcock,1901 

Sicyoninae Ortman,1898 

Sicyonia H. Milne-Edwards,1840 

Aristeidae Wood-Mason,1891 

Aristeinae Wood-Mason,1891 

Tribus: Benthysicymini Bouvier,1908 

Gennadas Bate,1881 

G. elegans (S.I. Smith,1882) 

= Amalopeneus elegans S.I. Smith,1882 

SW Greenland-Iceland-between Iceland & Faeroes-off W Ireland-Bay of Biscay, Mediterranean 

Benthysicymus Bate,1881 

Bentheogenemma Burkenroad,1936 

Tribus: Aristeini Bouvier,1908 

Plesiopenaeus Bate,1881 

= Plesiopeneus Faxon,1895 

P. edwardsianus (Johnson,1867) 

* "Scarlet shrimp" 

Along E Atlantic coasts up to S Biscay 

Hemipenaeus Bate,1881 

Aristeus Duvernoy,1840 

49

Hepomadus Bate,1881 

Aristeomorpha Wood-Mason,1891 

Solenocerinae Wood-Mason & Alcock,1891 

Solenocera Lucas,1849 

S. membranacea (Risso,1816) 

= Peneus membranaceus Risso,1816 

= Peneus siphonoceros Philippi,1840 

?European Atlantic, African Atlantic, Mediterranean 

Hymenopeneus S.I. Smith,1882 

Haliporus Bate,1881 

Sergestoidea Dana,1852 

Sergestidae Dana,1852 

Sergestinae Dana,1852 

Sergestes H. Milne-Edwards,1830 

S. arcticus Krøyer,1855 

Kattegatt-W Norway-Norw. Sea 

S. mollis S.I. Smith,1884 


Sergia Ortmann,1893 

S. robustus (S.I. Smith,1882) 

= Sergestes robustus S.I. Smith,1882 

N North Sea-Norw. Sea-Faeroes-Iceland-Cape Verde Isles 

Sicyonellinae Burkenroad,1983 

Luciferidae Burkenroad,1983 


EUKYPHIDA Boas,1880 

PROCARIDIDEA Chace & Manning,1972 

* No representing species within the area. 3 sp. known from caves in Bermuda, Hawaii & Ascension 

CARIDEA Dana,1852 

Oplophoroidea Dana,1852 

Oplophoridae Dana,1852 

Meningodora S.I. Smith,1882 ( Meningodora mollis S.I. Smith,1882) 

=? Notostomus A. Milne-Edwards,1881 

M. vesca S.I. Smith,1882? 

Faeroe-Iceland area 

Acanthephyra A. Milne-Edwards,1881 ( Acanthephyra armata A. Milne-Edwards,1881) 

= Ephyra P. Roux,1831, non Péron & Lesueur,1810 (Cnidaria), nec Duponchel,1829 (Lepidoptera) 

= Miersia Kingsley,1880 (Replacement name for Ephyra P. Roux, priority suppressed by ICZN) 

A. purpurea A. Milne-Edwards,1881 

Shetland-Faeroe area-W Ireland-Bay of Biscay 

A. pelagica (Risso,1816) 

= Alphaeus pelagicus Risso,1816 

=? Hoplocaricyphus similis Coutière,1907 

Shetland-Faeroe-Iceland area-W Ireland-Bay of Biscay 

Hymenodora G.O. Sars,1877 

H. glacialis (Buchholz,1874) 

= Pasiphaë glacialis Buchholz,1874 () 

Barents & Norw. Sea-Shetland-Faeroe-Iceland area-W Ireland-?Bay of Biscay 

H. gracilis (S.I. Smith,1882) 

= Miersia gracilis S.I. Smith,1882 

off W & S Ireland-Bay of Biscay 

Systellaspis Bate,1888 ( Systellaspis lanceocaudata Bate,1888 - "Challenger" material) 

= Hoplopasiphaea Yokoya & Shibata,1965 ( Hoplopasiphaea philippinensis Yokoyo & Shibata,1965) 

S. debilis (A. Milne-Edwards,1881) 

= Hoplopasiphaea philippinensis Yokoyo & Shibata,1965 

Shetland-Faeroe-Iceland area-W Ireland-Bay of Biscay 

Atyoidea de Haan,1849 

Atyidae de Haan,1849 

* Predominantly freshwater distribution, although 2 species occur in brackish waters 

Atyaephyra de Brito Capello,1867 

A. desmarestii desmarestii (Millet,1832) 

= Hyppolyte Desmarestii Millet,1832 

?European Atlantic, Madeira, Mediterranean 

Bresilioidea Calman,1896 

Bresiliidae Calman,1896 

= Disciadidae Rathbun,1902 

Bresilia Calman,1896 

B. atlantica Calman,1896 () 

off SW Ireland 

Nematocarcinoidea S.I. Smith,1884 

Rhynchocinetidae Ortmann,1890 


Nematocarcinidae S.I. Smith,1884 

Nematocarcinus A. Milne-Edwards,1881 ( Nematocarcinus cursor A. Milne-Edwards,1881) 

= Eumiersia S.I. Smith,1882 ( Eumiersia ensifera S.I. Smith,1882) 

= Stochasmus Bate,1888 ( Stochasmus exilis Bate,1888) 

N. ensifer (S.I. Smith,1882) exilis (Bate,1888) 

= Stochasmus exilis Bate,1888 

Shetland-Faeroe-Iceland area, Bay of Biscay 

Stylodactyloidea Bate,1888 


Stylodactylidae Bate,1888 

Pasiphaeoidea Dana,1852 

Pasiphaeidae Dana,1852 

Pasiphaea Savigny,1816 

= Phye Wood Mason,1892 ( Parapasiphaë alcocki Wood Mason & Alcock,1891) 

P. tarda Krøyer,1845 

Koster Channel-Skagerrak-Norw. Sea-Barents Sea-Faeroe Shetland Iceland area 

P. multidentata Esmark,1866 

Bohuslän-Skagerrak-N North Sea-Norw. Sea-Faeroe Shetland Iceland area-W Ireland-Bay of Biscay 

P. sivado (Risso,1816) 

= Alphaeus sivado Risso,1816 () 

Kattegatt-Bohuslän-Norw. Sea-Shetland Faeroe area-W (& S) Br. Isles-Bay of Biscay 

Parapasiphae S.I. Smith,1884 

= Orphania Bate,1888, non Fischer,1853 (Orthoptera) 

= Dantecia Caullery,1896 

P. sulcatifrons S.I. Smith,1884 () 

= Orphania tenuimana Bate,1888 ( of Orphania Bate) 

= Dantecia caudani Caullery,1896 ( of Dantecia Caullery) 

Faeroe Iceland area-off W Ireland 

P. cristata S.I. Smith,1884 

Shetland Faeroe area-off W Ireland 

Palaemonoidea Rafinesque-Schmaltz,1815 

Palaemonidae Rafinesque-Schmaltz,1815 

Palaemoninae Rafinesque-Schmaltz,1815 

Palaemon Weber,1795 

P. (Palaeander Holthuis,1950) elegans Rathke,1837 ( of Palaeander Holthuis,1950) 

= Leander squilla : Czerniavsky,1884, non (Linnaeus,1758) 

S Baltic-Bohuslän-North Sea-Shetland Faeroe area-W Br. Isles-Engl. Channel-Bay of Biscay 

P. adspersus Rathke,1837 () 

= P. fabricii Rathke,1843 & Cancer Squilla Linnaeus,1758 

Åland-Bohuslän-North Sea-Norw. Sea-Br. Isles-Bay of Biscay 

P. serratus (Pennant,1777) 

Öresund, ?Skagerrak-North Sea-Engl. Channel-W Br. Isles-Bay of Biscay 

P. longirostris Milne-Edwards,1837 

S North Sea-Engl-Channel, W Ireland-Bay of Biscay 

Palaemonetes Heller,1869 

P. varians (Leach,1814) 

= Palaemon varians Leach,1814 () 

Fresh water in Sw., Nor. & Denm., S Baltic, ?Kattegatt-?Skagerrak-North Sea-Engl. Channel-W Br. Isles- 

Bay of Biscay 

Leander E. Desmarest,1849 

L. tenuicornis (Say,1818) 

= Palaemon tenuicornis Say,1818 

= Leander erraticus E. Desmarest,1849 () 


Pontoniinae Kingsley,1878 

Typton Costa,1844 

T. spongicola Costa,1844 () 

Engl. Channel-Bay of Biscay 

Periclimines Costa,1844 

P. sagittifer (Norman,1861) 

Roscoff 

Gnathophyllidae Dana,1852 


Hymenoceridae Ortmann,1890 


Campylonotoidea Sollaud,1913 

Campylonotidae Sollaud,1913 


Psalidopodoidea Wood-Mason & Alcock,1892 

* The only 2 species known are W Atlantic & Indo-pacific in distribution 

Psalidopodidae Wood-Mason & Alcock,1892 

Alpheoidea Rafinesque-Schmaltz,1815 

Alpheidae Rafinesque-Schmaltz,1815 

Athanas Leach,1814 

A. nitescens (Leach,1814) 

= Palaemon nitescens Leach,1814 () 

Öresund-Bohuslän-W Norway-S North Sea-Engl. Channel-W Br. Isles-Mediterranean & Black Sea 

Alpheus Weber,1795 ( Alpheus avarus Fabricius,1798) 

A. glaber (Olivi,1792) 

Engl. Channel-W Br. Isles-Bay of Biscay 

A. macrocheles (Hailstone,1835) 


Hippolytidae Bate,1888 

50

Cryptocheles G.O. Sars,1870 

C. pygmaea G.O. Sars,1870 () 

Bohuslän-N North Sea-Norw. Sea 

Hippolyte Leach,1814 

= Virbius Stimpson,1860 ( Hippolyte acuminata Dana,1852) 

H. varians Leach,1814 () 

@ ?Alcyonium digitatum 

Öresund-Bohuslän-North Sea-Norw. Sea-Shetland Faeroe area-W Br. Isles-Engl. Channel-Bay of Biscay 

H. inermis Leach,1815 

= Hippolyte prideuxiana Leach,1817 

Engl. Channel-W Br. Isles-Bay of Biscay 

H. longirostris (Czerniavsky,1868) 


H. hunti (Gosse,1877) 

@ Antedon bifida 

Britain 

Bythocaris G.O. Sars,1870 

B. simplicirostris G.O. Sars,1870 () 

Norw. Sea-Barents Sea-Shetland-Faeroes area 

B. leucopis G.O. Sars,1879 

Norw. Sea-Barents Sea-Faeroes-Iceland area 

B. payeri (Heller,1875) 

Norw. Sea-Barents Sea-Shetland-Faeroes-Iceland area 

B. gracilis S.I. Smith,1885 

Shetland-Faeroe-Iceland area-off W Ireland 

Thoralus Holthuis,1947 

= Lysippe Kinahan,1858, non Malmgren,1865 (Polychaeta) ( Hippolyte cranchii Leach,1817 - generic 

name totally suppressed, ICZN Op. 671) 

T. cranchii (Leach,1817) 

= Hippolyte cranchii Leach,1817 () 

Bohuslän-North Sea-Norw. Sea-Shetland Faeroe area-W Br. Isles-Engl. Channel-Bay of Biscay 

Spirontocaris Bate,1888 

S. liljeborgii (Danielssen,1859) 

= S. lilljeborgii : Auctt. 

= Spirontocaris securifrons (Norman,1862) 

* Vilhelm or Wilhelm Liljeborg changed his name to William Lilljeborg (with a double l) around 1860, when 

he for a while was living in USA. This spelling should be reflected when citing him as an author (single l 

before 1860 and double after), but the original spelling of taxon names based upon his name must be used. 

S Baltic-Bohuslän-N North Sea-W Br. Isles & Engl. Channel-Shetland Faeroe Iceland are-Norw. Sea-Barents 

Sea 

S. spinus (James Sowerby,1805) 

= Cancer spinus James Sowerby,1805 () 

?Öresund-Skagerrak-N North Sea-W Br. Isles-Shetland Faeroe Iceland area-Norw. Sea-Barents Sea 

S. phippsii (Krøyer,1841) 

= Spirontocaris turgida (Krøyer,1841) 

?Bohuslän-Skagerrak-N North Sea-Norw. Sea-Barents Sea 

Lebbeus White,1847 

L. polaris (Sabine,1824) 

= Alpheus polaris Sabine,1824 

= Lebbeus orthorhynchus (Leach MS) White,1847 () 

Kattegatt-Bohuslän-N North Sea-Shetland Faeroe Iceland area-Norw. Sea-Barents Sea 

Eualus Thallwitz,1892 

E. gaimardii (H. Milne-Edwards,1837) 

= Hippolyte gaimardii H. Milne-Edwards,1837 

= Eualus obses Thallwitz,1892 () 

S Baltic-Bohuslän-North Sea-NW Br. Isles-Faeroe Iceland area-Norw. Sea-Barents Sea 

E. pusiolus (Krøyer,1841) 

Öresund-Bohuslän-North Sea-W Br. Isles-Shetland Faeroe Iceland area-Norw. Sea-Barents Sea 

E. occultus (Lebour,1936) 

Bohuslän-North Sea-Engl. Channel-W Br. Isles-Bay of Biscay 

Caridion Goës,1863 

= Doryphorus Norman,1861, non Cuvier,1829 (Reptilia) 

C. gordoni (Bate,1858) 

= Hippolyte gordoni Bate,1858 () 

Kattegatt-Bohuslän-North Sea-Barents Sea-Iceland-Faeroes-Br. Isles-Bay of Biscay 

C. steveni Lebour,1930 

Skagerrak-North Sea-Engl. Channel & W Br. Isles 

Lysmata Risso,1816 

= Melicerta Risso,1816, non Schrank,1803 (Vermes), nec Péron & Lesueur,1810 (Cnidaria) 

L. seticaudata (Risso,1816) 

= Melicerta Seti Caudata Risso,1816 () 


Ogyridae Hayand Share,1918 


Processoidea Ortmann,1890 

Processidae Ortmann,1890 

Processa Leach,1815 

= Nika Risso,1816 ( Nika edulis Risso,1816) 

* At least 2 species in Scandinavia. Which ones? 

P. canaliculata Leach,1815 () 

= Nika mediterranea Parisi,1915 & P. mediterranea : Nouvel & Holthuis,1957 

N North Sea-W Br. Isles-Engl. Channel-Bay of Biscay 

P. edulis (Risso,1816) crassipes Nouvel & Holthuis,1957 

S North Sea-Engl. Channel-W Br. Isles, Roscoff 

P. edulis arcassonensis Nouvel & Holthuis,1957 

Arcachon 

P. modica D.I.Williamson & Rochanaburanon,1979 modica D.I. Williamson & Rochanaburanon,1979 

= P. parva : Holthuis,1954, : Nouvel & Holthuis,1957, non Holthuis,1951 & P. aequimana : Auctt., non 

(Paulson,1875) 

S North Sea-Engl. Channel-W Br. Isles-Bay of Biscay 

P. nouveli Al-Adhub & D.I. Williamson,1975 holthuisi Al-Adhub & D.I. Williamson,1975 

= P. canaliculata : Nouvel & Holthuis,1957, non Leach,1815 

Bohuslän-North Sea-W Br. Isles-Bay of Biscay 

P. elegantula Nouvel & Holthuis,1957 

Roscoff, Mediterranean 

P. parva Holthuis,1951 

?European Atlantic, tropical W Africa 

Pandaloidea Haworth,1825 

Pandalidae Haworth,1825 

Pandalus Leach,1814 

= Dymas Krøyer,1861 

P. borealis Krøyer,1838 

= Dymas typus Krøyer,1861 ( of Dymas Krøyer,1861) 

Kattegatt-Bohuslän-N North Sea-Norw. Sea-Barents Sea-Shetland Faeroe Iceland area 

P. montagui Leach,1814 () 

S Baltic-Bohuslän-North Sea-Norw. Sea-Barents Sea-Shetland Faeroe Iceland area-W Br. Isles-Engl. 

Channel 

P. propinquus G.O. Sars,1870 

Bohuslän-Norw. Sea-Shetland Faeroe Iceland area-W Br. Isles-Bay of Biscay 

Dichelopandalus Caullery,1896 

D. bonnieri Caullery,1896 () 

Bohuslän-Norw. Sea-Shetland Faeroe Iceland area-W Br. Isles-Bay of Biscay 

Pandalina Calman,1899 

P. brevirostris (Rathke,1843) 

= Pandalus brevirostris Rathke,1843 () 

Öresund-Bohuslän-North Sea-Norw. Sea-Shetland Faeroe area-W Br. Isles-Engl Channel-Bay of Biscay 

P. profunda Holthuis,1946 

Skagerrak-Norw. Sea-Shetland Faeroe area-W Ireland-?Bay of Biscay 

Plesionika Bate,1888 ( P. uniproductata Bate,1888 (Junior subjective synon. of Acanthephyra ensis 

A. Milne-Edwards,1881)) 

P. edwardsii (Brandt,1851) 

* "Striped soldier shrimp", "Soldier striped shrimp" 

From E tropical Atlantiv to S Biscay, Mediterranean 

Thalassocarididae Bate,1888 

* Only 2 indo-pacific species known 

Physetocaridoidea Chase,1940 

* Only one bathypelagic species known from 1000-1200 m depths off Bermuda 

Physetocarididae Chase,1940 

Crangonoidea Haworth,1825 

Crangonidae Haworth,1825 

Crangon J.C. Fabricius,1798 

C. crangon (Linnaeus,1758) 

= Cancer crangon Linnaeus,1758 () 

Åland-Bohuslän-North Sea-Barents Sea-Iceland-Faeroes-Br. Isles-Bay of Biscay 

C. allmani Kinahan,1857 

Öresund-Bohuslän-North Sea-Barents Sea-Iceland-Faeroes-Br. Isles-Bay of Biscay 

Sclerocrangon G.O. Sars,1883 

S. boreas (Phipps,1774) 

= Cancer boreas Phipps,1774 () 

Norw. Sea-Faeroe-Iceland area-Barents Sea 

S. ferox (G.O. Sars,1877) 

Shetland-Faeroe area-off W Ireland-Norw. Sea-Barents Sea, N Greenland 

S. jacqueti (A. Milne-Edwards,18??) 

Shetland-Faeroe area-off W Ireland 

Philocheras Stebbing,1900 

P. echinulatus (M. Sars,1861) 

Bohuslän-N North Sea-Norw. Sea-Shetland Faeroe area-W Br. Isles-Bay of Biscay 

P. trispinosus (Hailstone,1835) 

North Sea-Shetland Faeroe area-W Br. Isles-Engl. Channel-Bay of Biscay 

P. bispinosus (Hailstone,1835) 

= Pontophilus bispinosus Hailstone,1835 

= Crangon nanus Krøyer,1842 () 

Kattegatt-Bohuslän-North Sea-Norw. Sea-Iceland-Br. Isles-Bay of Biscay 

P. sculptus (Bell,1853) 

?N North Sea, Engl Channel-W Br. Isles-Bay of Biscay 

P. fasciatus (Risso,1816) 

North Sea-?Norw. Sea-Engl. Channel-W Br. Isles-Bay of Biscay 

Pontophilus Leach,1817 

= Cheraphilus Kinahan,1862 ( Crangon spinosus Leach,1815) 

P. norvegicus (M. Sars,1861) 

Bohuslän-Norw. Sea-Iceland-Barents Sea 

P. spinosus (Leach,1815) 

= Crangon spinosus Leach,1815 () 

Kattegatt-Bohuslän-North Sea-Norw. Sea-Iceland-Faeroes-Br. Isles-Bay of Biscay 

Sabinea J.C. Ross,1835 

S. sarsii Smith,1879 

Bohuslän-Norw. Sea-Iceland-Barents Sea 

51

S. septemcarinatus (Sabine,1824) 

= Crangon septemcarinatus Sabine,1824 () 

Norw. Sea-Iceland-Barents Sea, N Greenland 

S. hystrix (A. Milne-Edwards,1881) 


Aegaeon Agassiz,1846 

A. lacazei (Gourret,1887) 


Glyphocrangonidae Smith,1884 


EUZYGIDA Burkenroad,1981 

* An extinct infraorder, Uncinidea Beurlen,1930 is also belonging here 

STENOPODIDEA Bate,1888 

Stenopodidae Claus,1872 

Richardina A. Milne-Edwards,1881 

R. spinicincta A. Milne-Edwards,1881 () 

off W Ireland-Bay of Biscay 

Spongicolidae Schram,1986 

Spongicoloides Hansen,1908 

S. profundus Hansen,1908 () 


REPTANTIA Boas,1880 

ASTACIDEA Latreille,1803 

Astacidae Latreille,1802 

* Some other fresh-water taxa exists in the area 

Astacus J.C. Fabricius,1775 

A. astacus (Linnaeus,1758) 

= A. fluviatilis J.C. Fabricius,1775 

Fresh water, in Sw., Denm., Finl., & Norw. 

Nephropidae Dana,1852 

Nephrops Leach,1814 

N. norvegicus (Linnaeus,1758) 

Öresund-Bohusl-North Sea-N Norway-Iceland-Br. Isles-Mediterranean 

Nephropis Wood-Mason,1873 

N. atlantica Norman,1882 

off N Scotland- Shetland Faeroe Channel-Bay of Biscay 

Homarus Weber,1795 

H. gammarus (Linnaeus,1758) 

= H. vulgaris H. Milne-Edwards,1837 

Kattegatt-Bohuslän-North Sea-Troms-Br. Isles-Mediterranean 

Thaumastochelidae Bate,1888 

* Only 2 species known, from off central America & off Japan 

Cambaridae Hobbs,1942 

* Only freshwater species 

Parastacidae Huxley,1879 

* Freshwater species in the Indo-pacific area & S South America 

THALASSINIDEA Latreille,1831 

Thalassinidae Latreille,1831 


Axiidae Huxley,1879 

Calocaris Bell,1846 

C. macandreae Bell,1846 

Kattegatt-Bohuslän-North Sea-Troms-Iceland-Br. Isles-Adriatic 

Calocarides Wollebaek,1909 

C. coronatus (Trybom,1904) 

= Eiconaxius coronatus Trybom,1904 () 

Bohuslän-Norw. Sea-Finnmark-N Br. Isles 

Axius Leach,1815 

A. stirynchus Leach,1815 

Br. Isles-Mediterranean 

A. nodulosus Meinert,1877 

off W Jutland 

Laomediidae Borradaile,1903 

Jaxea Nardo,1847 

J. nocturna Nardo,1847 ex Chiereghin MS 

* Jim Atkinson at Millport, when studying video tapes from the Swedish west coast with burrows of 

Nephrops & Calocaris in 50-60 m. depth, also found burrows typical of Jaxea (in fax to Mats Ulmestrand 

autumn 1996) 

SW Br. Isles & Irish Sea-Adriatic 

Callianassidae Dana,1852 

Callianassa Leach,1814 

C. subterranea (Montagu,1808) () 

N Kattegatt-Bohuslän-Skagerrak-Engl. Channel-W Br. Isles-Mediterranean 

C. tyrrhena (Petagna,1792) 

= C. stebbingi Borradaile,1903 

S North Sea-Scotland-Engl. Channel-Mediterranean 

Upogebiidae Borradale,1903 

Upogebia Leach,1814 

U. stellata (Montagu,1814) 

Kattegatt-Bohuslän-North Sea-W Norway-W Br. Isles-Engl. Channel-Mediterranean 

U. deltaura (Leach,1815) 

Kattegatt-Bohuslän-North Sea-W Norway-W Br. Isles-Engl. Channel-Mediterranean 

U. pusilla (Petagna,1792) 

Sponvika, Norway, Britain, Spain, Mediterranean 

Callianideidae Kossman,1880 

* Indo-Pacific & West Indian distribution 

Axianassidae Schmitt,1924 


PALINURA Latreille,1803 

Palinuridae Latreille,1803 

Palinurus Weber,1795 

P. elephas (J.C. Fabricius,1787) 

= P. vulgaris Latreille,1808 

Bergen-North Sea-W Br. Isles-Engl Channel-Mediterranean 

P. mauritanicus Gruvel,1911 

off SW Ireland-Senegal, Mediterranean 

Polychelidae Wood-Mason,1874 

Polycheles Heller,1862 

P. typhlops Heller,1862 

= Polycheles agassizi Faxon,1??? 

= Polycheles doederleini Riggio,1885 

= Pentacheles hextii Alcock,1901 

SW Iceland-SW Ireland-Bay of Biscay-Cap Verde Isles & Mediterranean 

P. granulatus Faxon,1893 

off SW Ireland-Madeira & Canaries 

Stereomastis Bate,1888 

S. sculpta (S.I. Smith,1880) 

= Polycheles sculptus S.I. Smith,1880 (Apr.) 

= Pentacheles spinosus A. Milne-Edwards,1880 (Dec.) 

off SW Iceland-W Ireland-Biscay-Mediterranean 

S. grimaldi (Bouvier,19??) 

off Iceland-off W Ireland-Biscay 

Glypheidae Winckler,1883 

* The only recent species is known from the vicinity of the Philippines 

Scyllaridae Latreille,1825 

Scyllarus J.C. Fabricius,1775 

S. arctus (Linnaeus,1758) 

W Engl. Channel, Madeira, Azores, Mediterranean 

Synaxidae Bate,1881 

* Monotypical family from the West Indies 

ANOMALA Boas,1880 

* This infraorder has often been combined with Thalassinidea in a larger taxon Anomura MacLeay,1838 

Diogenidae Ortmann,1892 

Diogenes Dana,1852 ( Pagurus miles J.C. Fabricius,1787) 

D. pugilator pugilator (Roux,1829) 

= Pagurus pugilator Roux,1829 

German Bight, Netherlands, Swansea-Engl. Channel-Bay of Biscay-Mediterranean 

Clibanarius Dana,1852 ( Cancer clibanarius Herbst,1791) 

C. erythropus (Latreille,1818) 

= Pagurus erythropus Latreille,1818 

= Pagurus misanthropus Roux,1828-30 

N Cornwall & Engl. Channel-Bay of Biscay-Azores-Mediterranean 

Paguridae Latreille,1803 

Pagurus J.C. Fabricius,1775 

P. bernardus (Linnaeus,1758) 

= Cancer bernhardus Linnaeus,1758 () 

Öresund-Bohusl-North Sea-N Norway-Barents Sea-area S of Spitsbergen-Shetland Faeroe Iceland area-Br. 

Isles-Bay of Biscay-Portugal 

P. pubescens Krøyer,1838 

= Pagurus thompsoni Bell,1853 

Kattegatt-Bohuslän-N North Sea-N Norway-Barents Sea & Spitsbergen-Shetland Faeroe Iceland area-W Br. 

Isles-Engl. Channel 

P. cuanensis Bell,1845 

Öresund-Bohuslän-N North Sea-W Norway (Bergen area)-?Shetland Faeroe area-W Br. Isles-Engl. Channel- 

Bay of Biscay 

P. prideaux Leach,1815 

= Pagurus prideauxi : Auctt. 

= Pagurus solitarius Roux,1828-30 

?Skagerrak-S & N North Sea-Tröndelag-Faeroe Shetland area-W Br. Isles-Engl. Channel-Bay of Biscay- 

Mediterranean 

P. alatus J.C. Fabricius,1775 

= Pagurus angulatus Risso,1816 

= Pagurus tricarinatus Norman,1869 

= Eupagurus variabilis A. Milne-Edwards & Bouvier,1892 

W Norway-N North Sea-W Norway-Shetland Faeroe area-Iceland-W Ireland-Bay of Biscay-Mediterranean 

P. forbesii Bell,1846 

= P. sculptimanus Lucas,1846 

Engl. Channel-Irish Sea-W Ireland-Bay of Biscay-Portugal, Madeira, Mediterranean 

P. carneus Pocock,1889 

Faeroes in >500m, off W Ireland 

52

P. chevreuxi (Bouvier,1896) 

= Eupagurus chevreuxi Bouvier,1896 

N Bretagne-Roscoff-Mediterranean 

P. pubescentulus (A. Milne-Edwards & Bouvier,1892) 

= Eupagurus pubescentulus A. Milne-Edwards & Bouvier,1892 

46° 09' N, 3° 38' E - Biscay - Angola, Azores 

P. anachoretus Risso,1826 

= Pagurus pictus H. Milne-Edwards,1836 

N Portugal - southwards 

Anapagurus Henderson,1886 

A. chiroacanthus (Liljeborg,1855) 

= Pagurus chiroacanthus Liljeborg,1855 

= Anapagurus brevicarpus A. Milne-Edwards & Bouvier,1892 

Kattegatt-Bohuslän-N North Sea-W Norway (Bergen area)-Shetland Faeroe area-W Br. Isles-Engl. Channel- 

Biscay- Azores-Mediterranean 

A. laevis (Bell,1845) 

= Pagurus laevis Bell,1845 () 

= Pagurus laevis W. Thompson,1844 (nomen nudum) 

Kattegatt-Bohuslän-North Sea-W Norway-Shetland Faeroe area-W Br. Isles-Engl. Channel-Bay of Biscay- 

Mediterranean 

A. hyndmanni (Bell,1845) 

= Pagurus hyndmanni Bell,1845 

= Pagurus hyndmanni W. Thompson,1844 (nomen nudum) 

N North Sea-Shetland Faeroe area-W Br. Isles-Engl. Channel-Bay of Biscay 

Cestopagurus Bouvier,1897 ( C. coutieri Bouvier,1897) 

C. timidus (Roux,1828-30) 

= Pagurus timidus Roux,1828-30 

= Catapaguroides Milne-Edwards & Bouvier,1892 timidus (Roux,1828-30) 

Clyde Sea area, Ireland N Bretagne-Bay of Biscay-Mediterranean 

Nematopagurus A. Milne-Edwards & Bouvier,1892 

N. longicornis A. Milne-Edwards & Bouvier,1892 () 

off Galway & W Ireland, N Spain-Azores-Madeira-Angola-Mediterranean 

Lithodidae Samouelle,1819 

Lithodes Latreille,1806 

L. maja (Linnaeus,1758) 

= Cancer maja Linnaeus,1758 

Öresund-Bohuslän-North Sea-N Norway-Barents Sea & Spitsbergen-Shetland Faeroe Iceland area-W Br. 

Isles 

Parapaguridae Smith,1882 

Parapagurus S.I. Smith,1879 

P. pilosimanus pilosimanus S.I. Smith,1879 

off S Iceland-off SW Faeroes-off W Ireland-Bay of Biscay 

Galatheidae Samouelle,1819 

Galathea J.C. Fabricius,1793 

G. intermedia Liljeborg,1851 

Öresund-Bohuslän-North Sea-Lofoten-Shetland Faeroe area-Br. Isles-Bay of Biscay 

G. strigosa (Linnaeus,1761) 

Kattegatt-Bohuslän-North Sea-Tröndelag-Shetland Faeroe area-W Br. Isles-Engl. Channel-Bay of Biscay 

G. squamifera Leach,1814 

Öresund-Bohuslän-North Sea-Tröndelag-W Br. Isles-Engl. Channel-Bay of Biscay 

G. nexa Embleton,1835 

?Kattegatt-Bohuslän-N North Sea-Norw. Sea-?Barents Sea-W Br. Isles-Engl. Channel 

G. dispersa Bate,1859 

Bohuslän-N North Sea-Norw. Sea-Shetland Faeroe Iceland area-W Br. Isles-Engl. Channel-Bay of Biscay 

G. machadoi Barrois,1888 

off W France 

Munida Leach,1820 

M. tenuimana G.O. Sars,1872 

?Bohuslän-Skagerrak-Norw. Sea-Barents Sea-Shetland Faeroe Iceland area-W Br. Isles, W Greenland, 

Iberia, Mediterranean 

M. sarsi Huus,1935 

= M. bamffica : Auctt. & M. rugosa : Auctt. 

Bohuslän-N North Sea-Norw. Sea-Barents Sea-Shetland Faeroe Iceland area - Biscay - N Spain 

M. rugosa (J.C. Fabricius,1775) 

= M. bamffia (Pennant,1777) 

Bohuslän-N North Sea-Sognefjord-Shetland Faeroe area-W Br. Isles-Bay of Biscay - Madeira - 

Mediterranean 

M. intermedia A. Milne-Edwards & Bouvier,1899 

E Atlantic north to S part of Porcupine Seabight 

M. microphthalma A. Milne-Edwards,1880 

off S Iceland, Porcupine Seabight, Cape Verde, Mid Atlantic Bight, W Atlantic 

Munidopsis Whiteaves,1874 

= Galathodes A. Milne-Edwards,1880 

M. serricornis (Lovén,1853) 

= M. tridentatus (Esmark,1857) 

Bohuslän-N Norway-Norw. Sea-W Ireland-Bay of Biscay 

M. curvirostra Whiteaves,1874 

off S Iceland-off W Ireland 

Chirostylidae Ortmann,1892 

Uroptychus Henderson,1888 

U. rubrovittatus (A. Milne-Edwards,1881) 

off S Iceland-off W Ireland-Bay of Biscay 

Porcellanidae Haworth,1825 

Pisidia Leach,1820 

P. longicornis (Linnaeus,1767) 

Öresund-Bohuslän-North Sea-Norw.Sea-Shetland Faeroe area-Br. Isles-Bay of Biscay 

Porcellana Lamarck,1801, non Müller,1766 

P. platycheles (Pennant,1777) 

North Sea-Shetland Faeroe area-Br. Isles-Bay of Biscay 

Pomatochelidae Miers,1879 

* Distribution chiefly tropical - subtropical 

Coenobitidae Dana,1851 

* Terrestrial with tropical distribution (e.g. Birgus ) 

Lomisidae Bouvier,1895 

* The only known species with E Australian distribution 

Aeglidae Dana,1852 

* Freshwater in temperate South America only 

Albuneidae Stimpson,1858 


Hippidae Latreille,1825 


BRACHYURA Latreille,1803 

DROMIACEA De Haan,1833 

Superfamily? 

Prosopidae von Meyer,1860 


Dynomenidae Ortmann,1892 

* Only in tropical seas 

Homolodromidae Alcock,1895 

Deep water (>600m). No species within the area? 

Dromioidea De Haan,1833 

Dromiidae De Haan,1833 

Dromia Weber,1795 

D. personata (Linnaeus,1758) 

S North Sea-Engl. Channel-Bay of Biscay-Mediterranean 

ARCHAEOBRACHYURA Guinot,1977 

Homolidea De Haan,1839 

Homolidae De Haan,1839 

Paromola Wood-Mason & Alcock,1891 

P. cuvieri (Risso,1816) 

N North Sea-off W Norway-Orkneys-off W Ireland-Bay of Biscay-Mediterranean 

Tymoloidea Alcock,1896 

Cymonomidae Bouvier,1897 

Cymonomus A. Milne-Edwards,1880 

C. granulatus (Thompson,1873) 

Porcupine & Rockall areas 

C. normani Lancester,1903 

Faeroes, deep water-Bailey & Rosemary Banks (SW of Faeroes) 

Tymolidae Alcock,1896 


Latreilliidae Stimpson,1858 


Raniniidae de Haan,1839 


EUBRACHYURA de St. Laurent,1980 

HETEROTREMATA Guinot,1977 

(OXYSTOMATA H. Milne-Edwards,1837) 

* Older systematic unit 

Calappoidea De Haan,1833 

Leucosiidae Samouelle,1819 

Ebaliinae Stimpson,1870 

Ebalia Leach,1817 

E. tumefacta (Montagu,1808) 

Kattegatt-Bohuslän-S & W Norway-Shetlands-Br. Isles-Portugal 

E. cranchii Leach,1817 

Kattegatt-Bohuslän-S & W Norway-Br. Isles 

E. tuberosa (Pennant,1777) 

Bohuslän-S & W Norway-Shetlands-Br. Isles-Mediterranean 

E. granulosa H. Milne-Edwards,1837 

Scotland-Ireland 

E. nux A. Milne-Edwards,1883 

SW Ireland-Porcupine Bank 

Calappidae De Haan,1833 


(BRACHYRHYNCHA Borradaile,1903) 


Cancroidea Latreille,1803 

53

Corystidae Samouelle,1819 

Corystes Bosc,1802 

C. cassivelaunus (Pennant,1777) 

Bohuslän-S Norway-North Sea-Br. Isles-Mediterranean 

Atelecyclidae Ortmann,1893 

Atelecyclus Leach,1814 

A. rotundatus (Olivi,1792) 

Bohuslän-S & W Norway-Faeroes-Shetlands-Br. Isles-Mediterranean 

A. undecimdentatus (Herbst,1783) 

NW France 

Thiidae Dana,1852 

Thia Leach,1815 

T. scutellata (Fabricius,1793) 

Bohuslän-W Norway-S North Sea-Br. Isles-Portugal-Mediterranean 

Pirimelidae Alcock,1899 

Pirimela Leach,1816 

P. denticulata (Montagu,1808) 

N Kattegatt-Bohuslän-S & W Norway-Br. Isles-NW Africa-Mediterranean 

Cancridae Latreille,1803 

Cancer Linnaeus,1758 

C. pagurus Linnaeus,1758 

Öresund-Bohuslän-North Sea-Troms-Shetlands-Br. Isles-Portugal-Mediterranean 

C. bellianus Johnson,1861 

SE Iceland-Shetlands-SW Ireland-Bay of Biscay-Portugal-Azores-Canaries 

Xanthoidea MacLeay,1838 

Goneplacidae H. Milne-Edwards,1837 

Goneplacinae H. Milne-Edwards,1837 

Goneplax Leach,1814 

G. rhomboides (Linnaeus,1758) 

Ireland-W Br. Isles-Engl. Channel 

Xanthidae MacLeay,1838 

Pilumnus Leach,1815 

P. hirtellus (Linnaeus,1761) 

W Norway-S North Sea-Engl. Channel-W Br. Isles-Cape Verde-Mediterranean-Black Sea 

P. spinifer H. Milne-Edwards,1834 

* Sometimes considered to be a subsp. of P. hirtellus (Linnaeus) 

Black Sea-Mediterranean-Azores-Mauretania-Portugal. Many specimens were found 1826 in Uddevalla 

harbour (Bohuslän). 

Pilumnoides H. Milne-Edwards & Lucas,1843 

P. inglei Guinot & Macpherson,1987 

= P. perlatus : Auctt., non (Poeppig,1836) (This is a South American species) 

Recorded a few times in SW Br. Isles 

Neopanope A. Milne-Edwards,1880 

N. sayi (Smith,1869) 

Nort American species introduced into Swansea area 

Monodaeus Guinot,1967 

M. couchi (Couch,1851) 

Ireland-W Br. Isles-W Engl. Channel 

Xantho Leach,1814 

X. pilipes A. Milne-Edwards,1867 

N Kattegatt-Bohuslän-W Norway-Shetlands-Br. Isles-SW North Sea-Engl. Channel-Senegal-Mediterranean 

X. incisus Leach,1814 

Ireland-W Br. Isles-Engl. Channel-W France-southwards 

Rhithropanopeus Rathbun,1898 

R. harrisii (Gould,1841) 

E North American species, which has been established in S North Sea and has also been found fr. Poland-Belt 

Sea-Öresund, Norway, N & W France, Black Sea & Caspian Sea 

Portunoidea Rafinesque-Schmaltz,1815 

Geryonidae Colosi,1923 

Geryon Krøyer,1837 

G. trispinosus (Herbst,1803) 

= G. tridens Krøyer,1837 () 

Kattegatt-Bohuslän-S & W Norway-Tromsö-N North Sea-Iceland-off N Scotland-Bay of Biscay-?Morocco- 


Chaecon Manning & Holthuis,1989 

C. affinis (A. Milne-Edwards & Bouvier,1894) 

off W Norway-off S Iceland-off Rockall-Azores 

C. inglei Manning & Holthuis,1989 

Porcupine Seabight 

Portunidae Rafinesque-Schmaltz,1815 

Polybiinae Ortmann,1893 

Polybius Leach,1820 

P. henslowii Leach,1820 

Skagerrak-N North Sea-Engl. Channel-Morocco-Mediterranean 

Bathynectes Stimpson,1870 

B. longipes (Risso,1816) 

W Br. Isles-Engl. Channel-NW France 

B. maravigna (Prestandrea,1839) 

= Portunus superbus Costa,1853 

off W Norway (Storeggen)-Faeroes-Shetlands-Bay of Biscay-Angola-Mediterranean 

Macropipus Prestandrea,1833 

M. tuberculatus (Roux,1830) 

= Portunus tuberculatus Roux,1830 () 

W Norway (Stavanger-off Storeggen)-Shetlands-W Br. Isles-Engl. Channel-Bay of Biscay-Azores- 

Mediterranean 

Necora Holthuis,1987 

N. puber (Linnaeus,1767) 

= Cancer puber Linnaeus,1767 () 

W Norway-S North Sea-Br. Isles-NW Africa-Mediterranean 

Liocarcinus Stimpson,1870 

L. arcuatus (Leach,1814) 

Öresund-Bohuslän-S & W Norway-North Sea-Br. Isles-Mediterranean-Black Sea 

L. corrugatus (Pennant,1777) 

?Yorkshire-W Br. Isles-Engl. Channel-NW France 

L. zariquieyi (Gordon,1968) 

??Shetlands-SW Br. Isles-Mediterranean 

L. pusillus (Leach,1815) 

Öresund-Bohuslän-S & W Norway-Lofoten-North Sea-Shetlands-Br. Isles-NW Africa-Mediterranean 

L. depurator (Linnaeus,1758) 

Öresund-Bohuslän-North Sea-Lofoten-Shetlands-Br. Isles-Nw. Africa-Mediterranean 

L. holsatus (Fabricius,1798) 

= Portunus holsatus Fabricius,1798 () 

Öresund-Bohuslän-North Sea-N Norway-Iceland-Faeroes-Shetlands-Br. Isles-Portugal-Canaries 

L. marmoreus (Leach,1814) 

Helgoland-Netherlands-Br. Isles-S Portugal-Azores 

Carcininae MacLeay,1838 

Carcinus Leach,1814 

C. maenas (Linnaeus,1758) 

= Cancer maenas Linnaeus,1758 

S Baltic-Bohuslän-Troms-Iceland-Faeroes-Shetlands-Br. Isles-SPortugal-Mauritania 

Xaiva MacLeay,1838 

X. biguttata (Risso,1816) 

Engl. Channel 

Portumnus Leach,1814 

P. latipes (Pennant,1777) 

German North Sea coast-Engl. Channel-Br. Isles-Morrocco-Mediterranean 

Portuninae Rafinesque-Schmaltz,1815 

Callinectes Stimpson,1860 

C. sapidus Rathbun,1896 

E American (Nova Scotia - Uruguay) species, which has been established in E Mediterranean and a few 

specimens has been found in Öresund, S North Sea & W France 

Pinnotheroidea De Haan,1833 

Pinnotheridae De Haan,1833 

Pinnotherinae De Haan,1833 

Pinnotheres Bosc,1802 

P. pisum (Linnaeus,1767) 

Öresund-Bohuslän-S Norway-North Sea-Br. Isles-Mauretania-Mediterranean 

P. pinnotheres (Linnaeus,1758) 

Yorkshire-Engl. Channel-W Br. Isles-W France-?southwards 

Asthenognathinae Stimpson,1858 

Asthenognathus Stimpson,1858 

A. atlanticus Monod,1933 

NW France 

(OXYRHYNCHA Latreille,1803) 


Majidae Samouelle,1819 

Majinae Samouelle,1819 

Maja Lamarck,1801 

M. squinado (Herbst,1788) 

Helgoland-Engl. Channel-W Br. Isles-Guinea-Mediterranean 

Oregoniinae Garth,1958 

Hyas Leach,1814 

H. araneus (Linnaeus,1758) 

Öresund & Belt Sea-Bohuslän-North Sea-N Norway-Spitsbergen-Iceland-Faeroes-Shetlands-Br. Isles-N 

Engl. Channel 

H. coarctatus Leach,1815 

Öresund & Belt Sea-Bohuslän-North Sea-N Norway-Spitsbergen-Barents Sea-Iceland-Faeroes-Shetlands-Br. 

Isles-Engl. Channel-Bay of Biscay 

Inachinae MacLeay,1838 

Achaeus Leach,1817 

54

A. cranchii Leach,1817 

Belgium-Engl. Channel-W Br. Isles-Azores-Mediterranean 

Inachus Weber,1795 

I. dorsettensis (Pennant,1777) 

Öresund & Belt Sea-Bohuslän-North Sea-North Tröndelag-Shetlands-Br. Isles-S Africa-Mediterranean 

I. phalangium (Fabricius,1775) 

= Inachus dorynchus Leach,1815 

N Kattegatt-Bohuslän-S North Sea-W Norway-Shetlands-Br. Isles-Cape Verde-Mediterranean 

I. leptochirus Leach,1817 

off W Jutland-Shetlands-off W Faeroes-Br. Isles-W Africa-Mediterranean 

Dorhynchus Thomson,1873 

D. thomsoni Thomson,1873 

W Norway-off W Shetlands-S Iceland-between Faeroes & Scotland-Cape Verde-Mediterranean 

Macropodia Leach,1814 

M. rostrata (Linnaeus,1761) 

Öresund & Belt Sea-Bohuslän-North Sea-Norway (65° N)-Shetlands-Br. Isles-Azores-S Africa-Mediterranean 

M. parva Van Noort & Adema,1985 

Bohuslän, Oslofjord, S North Sea, S Portugal 

M. linaresi Forest & Zariquiey Alvarez,1964 

SW North Sea, Engl. Channel-NW France, Mediterranean 

M. deflexa Forest,1978 

Ireland-W Br. Isles-Engl. Channel-NW & SW France 

M. tenuirostris (Leach,1814) 

Netherlands-Br. Isles-Shetlands-off W Faeroes-Engl. Channel-Portugal 

Pisinae Dana,1853 

Pisa Leach,1814 

P. armata (Latreille,1803) 

Belgium-Engl. Channel-W Br. Isles-Angola-Mediterranean 

P. tetraodon (Pennant,1777) 

Thames estuary-Engl. Channel-W Br. Isles-Ireland-W France-southwards 

Eurynome Leach,1814 

E. aspera (Pennant,1777) 

Öresund-Bohuslän-S & W Norway-?Shetlands-Br. Isles-NW Africa-Mediterranean 

E. spinosa Hailstone,1835 

Bohuslän-S & W Norway-W Br. Isles-Engl. Channel-NW Spain-Mediterranean 

Rochinia A. Milne-Edwards,1875 

R. carpenteri (Thomson,1873) 

off W Norway-between Norway & Shetlands-off S Iceland-between Faeroes & Scotland-off Ireland-Bay of 

Biscay-NW Africa 

Parthenopidae MacLeay,1838 

Parthenopinae MacLeay,1838 

Parthenope Weber,1795 

P. massena (Roux,1830) 

NW France 

Mimilambridae Williams,1979 

* The only known recent species collected in West Indies (Tobago) 

Bellidae Dana,1852 

* S South American & New Zealand distribution 

Bythograeidae Williams,1980 

* The only known species is collected from vicinity of thermal vents on the Galapagos Rift 

Deckeniidae Bott,1970 

* Freshwater crabs from E Africa & the Seychelles 

Platyxanthidae Guinot,1977 

* Occurs only in S South America 

Potamidae Ortmann,1896 

* Eurasian freshwater crabs 

Retroplumidae Gill,1894 

* Strictly Indian Ocean 

THORACOTREMATA Guinot,1977 

Grapsoidea MacLeay,1838 

Grapsidae MacLeay,1838 

Grapsinae MacLeay,1838 

Planes Bowdich,1825 

P. minutus (Linnaeus,1758) 

Sargasso Sea-(occasionally) S North Sea-Engl. Channel-Mediterran. 

Pachygrapsus Randall,1840 

P. marmoratus (Fabricius,1787) 

NW France-southwards 

P. transversus (Gibbes,1850) 

warm Atlantic-Mediterranean-(once on a ship) Copenhagen 

Varuninae H. Milne-Edwards,1853 

Eriocheir De Haan,1835 

E. sinensis H. Milne-Edwards,1854 

River mouths in Bothnian Bay-S Baltic-Göteborg-S North Sea-Thames-W France 

Brachynotus De Haan,1833 

B. sexdentatus (Risso,1826) 

W Br. Isles-Mediterranean 

Hemigrapsus Dana,1851 

H. penicillatus (de Haan,1835) 

* Should possibly be transferred to genus Brachynotus. 

Shore-living invader from NW Pacific, which first was observed in La Rochelle, Biscay coast of France in 

March 1994. It is spreading rapidly (probaly >100 km/year) both southwards and northwards and will 

possibly reach Scandinavian shores within a few years. 

Gecarcinidae MacLeay,1838 

* Tropical American, W African & Indo-Pacific terrestrial crabs 

Gecarcinucidae Rathbun,1904 

* Old world freshwater crabs 

Hapalocarcinidae Calman,1900 

* Distribution like reef corals, in which they produce "galls" 

Hexapodidae Miers,1886 


Hymenosomatidae MacLeay,1838 


Isolapotamidae Bott,1970 

* Freshwater crabs from Indonesia, Philippines & Coastal Asia 

Ocypodidae Rafinesque-Schmaltz,1815 

* Predominantly warm & tropical areas, e.g. Uca 

Palicidae Rathbun,1898 


Parathelphusidae Alcock,1910 

*Large, old world freshwater crabs 

Potamocarcinidae Ortmann,1899 

* New world freshwater crabs 

Potamonautidae Bott,1920 

* African freswater crabs 

Pseudothelphusidae Ortmann,1893 


Sinopotamidae Bott,1970 

* Freshwater crabs from China & Burma 

Sundathelphusidae Bott,1969 

* Old world freshwater crabs 

Trichodactylidae Milne-Edwards,1853 


MYSIDA Boas,1883 

Petalophtalmidae Willemoës-Suhm,1875 

Hansenomysis Stebbing,1893 

H. fyllae (Hansen,1887) 

Faeroe Channel-SW Iceland-off W Ireland 

Petalophthalmus Willemoes-Suhm,1875 

P. armiger Willemoes-Suhm,1875 

off W Ireland-off W Africa 

Mysidae Latreille,1803 

Boreomysinae Holt & Tattersall,1905 

Boreomysis G.O. Sars,1869 

B. tridens G.O. Sars,1870 

Skagerrak-W Norway-Malangen-Faeroes-Iceland-W Br. Isles-Bay of Biscay 

B. arctica (Krøyer,1861) 

Koster Channel-Lofoten-Arctic Seas-Faeroe Channel-W Br. Isles-Mediterranean 

B. microps G.O. Sars,1883 

Arctic Seas-Iceland-Faeroe Channel-off Ireland-Azores 

B. megalops G.O. Sars,1872 

Hardangerfjord-Finnmark-off Scotland & W Ireland, ?Mediterranean 

B. scyphops G.O. Sars,1879 

* Not listed by Mauchline,1980. Synonymized? 

Arctic Seas-Faeroe-Iceland area 

B. nobilis G.O. Sars,1885 

Arctic Seas-Faeroe-Iceland area 

B. semicaeca Hansen,1905 

Bay of Biscay 

Siriellinae Norman 

Siriella Dana,1850 

S. armata (Milne-Edwards,1837) 

Helgoland-Br. Isles-Mediterranean 

S. norvegica G.O. Sars,1869 

Kattegatt-Bohuslän-Hardangerfjord-Cape Stadt-Br.Isles-Mediterranean 

S. clausii G.O. Sars,1876 

Holland-W Scotland-Engl. Channel-Mediterranean 

S. jaltensis Czerniavsky,1868 

E & W Scotland-Engl. Channel-Black Sea & W Africa 

S. jaltensis brooki Norman,1887 

= S. gordonae Zimmer,1932 

W Scotland & SW North Sea 

Gastrosaccinae Norman 

Gastrosaccus Norman,1868 

G. sanctus (van Beneden,1861) 

55

?Skagerrak, Nordland-Finnmark,W Denmark-SW North Sea-S & W Br. Isles-W Africa-Black Sea 

G. spinifer (Goës,1864) 

S Baltic-Bohuslän-SW Norway-North Sea-Br. Isles-W Africa-Black Sea 

G. normani G.O. Sars,1877 

Engl. Channel-Bay of Biscay, Ireland, ?SW North Sea, W Br. Isles 

G. lobatus Nouvel,1951 

Engl. Channel-Bay of Biscay, ?SW North Sea, ?W Br. Isles 

Anchialina Norman & Scott,1906 


A. agilis (G.O. Sars,1877) 

W Scotland & Holland-Engl. Channel-Mediterranean 

Mysinae Latreille,1803 

Erythropini 

Erythrops G.O. Sars,1869 

E. erythrophthalma (Goës,1864) 

= E. goesi (G.O. Sars,1866) 

Öresund-Bohuslän-all Norway-Spitsbergen-E Scotland-Ireland-W England, N Greenland 

E. abyssorum G.O. Sars,1869 

Skagerrak-all Norway-Arctic Seas-Faeroe Channel, N Greenland 

E. serrata (G.O. Sars,1863) 

Bohuslän-Porsangerfjord-E Scotland-Ireland-Isle of Man-Iceland-?Arctic Seas 

E. elegans (G.O. Sars,1863) 

Öresund-Skagerrak-Trondheimsfjord-North Sea-Br. Isles-Mediterranean 

E. microps (G.O. Sars,1864) 

Skagerrak-Lofoten & Arctic Seas, W Ireland 

E. glacialis G.O. Sars,1885 

Between Norway & Iceland-Arctic Seas 

Parerythrops G.O. Sars,1869 

P. spectabilis G.O. Sars,1885 

W-N Norway-Arctic Seas-between Norway & Iceland 

P. abyssicola G.O. Sars,1876 

* Not listed in Mauchline,1980. Synonymized? 

W-N Norway 

P. obesa (G.O. Sars,1864) 

Koster Channel-all Norway-Iceland-W Ireland-?Mediterranean 

Meterythrops S.I. Smith,1879 

M. picta Holt & Tattersall,1905 

E North Sea-Br. Isles-Iceland & Greenland-Bay of Biscay 

M. robusta S.I. Smith,1879 

Bodö-N Norway-Iceland-Arctic Seas 

Katerythrops Holt & Tattersall,1905 

K. oceanae Holt & Tattersall,1905 

W Ireland-southwards in Atlantic 

Hypererythrops Holt & Tattersall,1905 

H. serriventer Holt & Tattersall,1905 

W Ireland 

Dactylerythrops Holt & Tattersall,1905 

D. dactylops Holt & Tattersall,1905 

Faeroe Channel-off W & SW Ireland 

D. bidigitata W. Tattersall,1907 

off W Ireland 

D. gracilura W. Tattersall,1907 

off W Ireland 

Pseudomma G.O. Sars,1870 

P. affine G.O. Sars,1870 

Koster Area, Skagerrak-Lofoten-Greenland-Iceland-Faeroes-Ireland-Bay of Biscay 

P. roseum G.O. Sars,1870 

Kostersäcken-Skagerrak-Malangen-Iceland-Faeroes 

P. truncatum S.I. Smith,1879 

off Cape Stadt, off N Faeroes, Lyngenfjord-Arctic Seas & Iceland 

P. frigidum Hansen,1908 

off Cape Stadt, off E & N Faeroes, Arctic Seas 

P. nanum Holt & Tattersall,1906 

Faeroe Channel-off W & SW Ireland 

P. jasi Melan & Brattegard,1995 

off SW Faeroes in 1319 m. 

Parapseudomma Nouvel & Lagardère,1976 

P. calloplura (Holt & Tattersall,1905) 

Faeroe Channel-off W & SW Ireland, Mediterranean 

Michthyops W. Tattersall,1911 

M. parva (Vanhöffen,1897) 

W Greenland-off W Ireland 

M. theeli (Ohlin,1901) 

Arctic Seas 

Amblyops G.O. Sars,1869 

A. abbreviata (G.O. Sars,1869) 

Koster Channel-Skagerrak-all Norway-Arctic Seas-Iceland-Faeroes-Ireland-Bay of Biscay 

A. kempi (Holt & Tattersall,1905) 

off Faeroes & Hebrides- off W Ireland 

A. tenuicauda W. Tattersall,1911 

Faeroe Channel-off SW Ireland 

Paramblyops Holt & Tattersall,1905 

P. rostrata Holt & Tattersall,1905 

S Iceland-Faeroes-W & SW Ireland-Mediterranean 

P. bidigitata W. Tattersall,1911 

off SW Ireland 

Dactylamblyops Holt & Tattersall,1906 

D. thaumatops W. Tattersall,1907 


D. goniops W. Tattersall,1907 


D. sarsi (Ohlin,1901) 

Arctic Seas 

Metamblyops W. Tattersall,1907 

M. oculata W. Tattersall,1907 


Longithorax Illig,1906 

L. fuscus Hansen,1908 

off S Iceland-off SW Ireland-Bay of Biscay 

Euchaetomera G.O. Sars,1883 

E. tenuis G.O. Sars,1883 

off SW Ireland-Eur. Atlantic-Mediterranean 

Chunomysis Holt & Tattersall,1905 

C. diadema Holt & Tattersall,1905 


Arachnomysis Chun,1887 

A. leuckartii Chun,1887 

off-SW Ireland-Eur. & Afr. Atlantic & Mediterranean 

Leptomysini Czerniavsky,1882 

Leptomysis G.O. Sars,1869 

L. gracilis (G.O. Sars,1864) 

Öresund-Bohuslän-S Norway-Hardangerfjord-North Sea-Shetlands-Br. Isles-Mediterranean 

L. lingvura (G.O. Sars,1866) 

Farsund-Molde-Br. Isles-Black Sea 

L. mediterranea G.O. Sars,1877 

SW North Sea & Ireland-Suez & Adriatic Sea 

Bathymysis W. Tattersall,1907 

B. helgae W. Tattersall,1907 

Faeroe Channel-off W Ireland 

Mysidetes Holt & Tattersall,1906 

M. farrani (Holt & Tattersall,1905) 

deep Skagerrak, off W Ireland-Bay of Biscay-Mediterranean 

Mysideis G.O. Sars,1869 

M. insignis (G.O. Sars,1864) 

Koster Channel-W Finnmark-Iceland-Faeroes-Br. Isles-Bay of Biscay 

Mysidopsis G. O. Sars,1864 

M. gibbosa G.O. Sars,1864 

Öresund-Bohuslän-North Sea-Ålesund-Br. Isles-Mediterran. 

M. angusta G.O. Sars,1864 

Öresund-Skagerrak-North Sea-Ålesund-Br. Isles-Mediterran. 

M. didelphys (Norman,1863) 

Öresund-Skagerrak-Lofoten-Iceland-Br. Isles-W France 

Pseudomysis G.O. Sars,1879 

P. abyssi G.O. Sars,1879 

Arctic Seas & between Norway & Iceland 

Iimysis Nouvel,1966 

I. atlantica (Nouvel,1942) 

= Tenagomysis atlantica Nouvel,1942 

Bay of Biscay 

Mysini Dana,1850 

Hemimysis G.O. Sars,1869 

H. anomala G.O. Sars,1907 

* inhabitant of sublittoral crevices from the Caspian area. Observed in the Baltic from 1992. 

Gulf of Finland 

H. lamornae (R.Q. Couch,1856) 

= Mysis lamornae R.Q. Couch,1856 

Kattegatt-Bohuslän-North Sea-Lofoten-Iceland-Br. Isles-Black Sea 

H. abyssicola G.O. Sars,1869 

Koster Channel, Skagerrak-W Norway (Tranöydypet) 

Mysis Latreille,1802 

M. relicta Lovén,1862 

Limnic in Lakes of Sw., No., Fi., De. etc., Bothn. Bay-Baltic (N of 58°), Arctic Seas, e.g. E & N Greenland 

M. oculata (Fabricius,1780) 

?Bothnian Sea-?C. Baltic, Lofoten-Arctic Seas & Iceland 

56

M. litoralis (Banner,1948) 

Troms (Rossfjordvatnet), Arctic Seas-Iceland 

Michtheimysis Norman,1902 

M. mixta (Liljeborg,1852) 

= Mysis mixta Liljeborg,1852 

Bothnian Sea-Bohuslän, Trondheimsfjord-Norw. Sea-Arctic Seas & Iceland 

Paramysis Czerniavsky,1882 

P. (Paramysis) helleri (G.O. Sars,1877) 

* Strictly Mediterranean according to Labat,1953. It has been confused 

with P. nouveli Labat,1953 

?W Scotland & ?E England-?Black Sea 

P. (Paramysis) arenosa (G.O. Sars,1877) 

Ireland-W Scotland-Engl. Channel-Mediterranean 

P. (Paramysis) portzicensis Nouvel,1950 

Bay of Biscay 

P. (Paramysis) nouveli Labat,1953 

Roscoff, Devon 

P. lacustris (Czerniavsky,1882) 

= Mesomysis kowalevskyi ?,1??? 

Caspian Sea, Baltic (introduced via Newman River in Lithuania, where it was used as fish food) 

P. (Pseudoparamysis Bacesco,1940?) bacescoi Labat,1953 

Roscoff, Brest, Finistère 

Schistomysis Norman,1892 

S. spiritus (Norman,1860) 

?S Baltic-Öresund-Bohuslän-North Sea-North Tröndelag-Faeroes-Br. Isles-W France-?Bay of Biscay 

S. ornata (G.O. Sars,1864) 

S Baltic-Bohuslän-Lofoten-North Sea-Faeroes-Iceland-Br. Isles-W Africa 

S. kervillei (G.O. Sars,1885) 

Helgoland-Holland-S &. W England-Ireland 

S. parkeri Norman,1892 

S England & W Morocco 

Praunus Leach,1814 

P. inermis (Rathke,1843) 

Åland-Bohuslän-all Norway-White Sea-Br. Isles-N Brittany 

P. flexuosus (O.F. Müller,1776) 

Bothnian Sea-Bohuslän-Foldenfjord-?Arctic Seas-Br. Isles-Bay of Biscay 

P. neglectus (G.O. Sars,1869) 

Öresund-Bohuslän-Lofoten-Br. Isles-N Brittany 

Mesopodopsis Czerniavsky,1882 

M. slabberi (van Beneden,1861) 

Kiel area-Öresund-Bohuslän-S Norway-Br. Isles-Black Sea 

Neomysis Czerniavsky,1882 

N. integer (Leach,1814) 

= Mysis vulgaris Thompson,1828 

Bothnian Bay-Bohuslän-Nordland, ?White Sea, Br. Isles-Spain 

Acanthomysis Czerniavsky,1882 

A. longicornis (Milne-Edwards,1837) 

Ireland-S & W England & SW North Sea-Mediterranean 

Stilomysis Norman,1892 

S grandis (Goës,1863) 

Troms-Finnmark-Iceland-Arctic Seas 

Heteromysini Norman,1892 

Heteromysis S.I. Smith, in Verrill,1874 

H. formosa S.I. Smith, in Verrill,1874 

= H. norvegica G.O. Sars,1882 

Bergen, E & W Scotland-Engl. Channel 

H. microps (G.O. Sars,1877) 

Engl. Channel, Tunisian Gulf 

H. armoricana Nouvel,1940 

Engl. Channel 

Mysidellinae Czerniavsky,1882-83 

Mysidella G.O. Sars,1872 

M. typica G.O. Sars,1872 

W Norway-Ålesund, off SW Ireland 

M. typhlops G.O. Sars,1872 

W Norway-Florö 

Lepidomysidae Fage,1924 

* In fresh & brackish water caves or land crab burrows. No species within the area. 

Stygiomysidae Caroli,1937 

* Cave inhabitants from West Indies & S Italy 

LOPHOGASTRIDA Boas,1883 

* Was formerly treated as part of "Mysidacea" 

Eucopiidae Dana,1852 

Eucopia Dana,1852 

E. unguiculata (Willemoes-Suhm,1875) 

= E. hanseni Nouvel,1942 

Faeroe Channel-European Atlantic & Mediterranean 

E. grimaldii Nouvel,1942 

off W Iceland-off W Ireland & European Atlantic 

E. sculpticauda Faxon,1893 

off W Hebrides-European Atlantic 

Lophogastridae M. Sars,1857 

Gnathophausia Willemoes-Suhm,1875 

G. zoea Willemoes-Suhm,1875 

off Norway (below ≈650 m), off W Iceland-Faeroe Channel-European Atlantic 

G. gigas Willemoes-Suhm,1875 

Greenland-off W Br. Isles & Ireland 

Lophogaster M. Sars,1857 

L. typicus M. Sars,1857 

Koster Channel-Skagerrak-Trondheimsfjorden-Br. Isles-Mediterranean 

MICTACEA Bowman, Garner, Hessler, Iliffe & Sanders,1985 

* no species in the area 

EDRIOPHTHALMA Leach,1815 

ISOPODA Latreille,1817 

GNATHIIDEA Hansen,1916 

Gnathiidae Harger,1880 

Akidognathia Stebbing,1912 

A. oedipus Stebbing,1913 

Rockall Bank 

Paragnathia Omer-Cooper & Omer-Cooper,1916 

P. formica (Hesse,1864) 

= Anceus formica Hesse,1864 

Ireland, W & E England, Holland 

Gnathia Leach,1814 

G. abyssorum (G.O. Sars,1872) 

= Anceus abyssorum G.O. Sars,1872 

@ Stryphnus ponderosus, Tragosia infundibuliformis, Axinella arctica, Phakellia robusta, Steletta normani 

Faeroes-Norw. Sea-Iceland & Arctic Seas 

G. albescens Hansen,1916 


G. arctica Gurjanova,1929 

Novaja Zemlja 

G. bicolor Hansen,1916 

Norw. Sea-off S Iceland-Arctic Seas 

G. cristatipes Stebbing,1913 

deep W Engl. Channel 

G. dentata (G.O. Sars,1872) 

= Anceus dentatus G.O. Sars,1872 

Kattegatt-Bohuslän-Trondheim, Holland 

G. elongata (Krøyer,1849) 

= Anceus elongatus Krøyer,1849 

Lofoten-N Norway-Iceland & Arctic Seas, e.g. N Greenland 

G. hirsuta (G.O. Sars,1877) 

= Anceus hirsutus G.O. Sars,1877 


G. maxillaris (Montagu,1804) 

= Cancer maxillaris Montagu,1804 

Ireland-W & S Br. Isles & S North Sea 

G. oxyuraea (Liljeborg,1855) 

Öresund-Oslofjord-Lofoten-Br. Isles 

G. robusta (G.O. Sars,1879) 


G. schistifrons Stebbing,1912 

Ireland in 380 m 

G. stygia (G.O. Sars,1877) 

= Anceus stygius G.O. Sars,1877 


G. vorax (Lucas,1849) 

Greenland-Br. Isles-Mediterranean 

ANTHURIDEA Leach,1814 

Antheluridae Poore & Lew Ton,1988 

Anthelura Norman & Stebbing,1886 

A. elongata Norman & Stebbing,1886 () 

NE Atlantic in 1100-1590 m 

A. truncata (Hansen,1916) 

N Atlantic 

Ananthura Barnard,1925 

A. sulcaticauda Barnard,1925 

off SW & NE Iceland in 216-1505 m 

Anthuridae Leach,1814 

Anthura Leach,1814 

A. gracilis (Montagu,1808) 

= Oniscus gracilis Montagu,1808 () 

S Baltic-North Sea-S & W Br. Isles-Mediterranean 

Cyathura Norman & Stebbing,1886 

C. carinata (Krøyer,1847) 

= Anthura carinata Krøyer,1847 

S Baltic(Karlskrona)-Kungsbacka, North Sea-England-Morocco, Greenland 

57

Haliophasma Haswell,1881 

H. mjoelniri Negoescu & Svavarson,1997 

off SW Iceland in 340-508 m 

Quantanthura Menzies & George,1972 

Q. tyri Negoescu & Svavarson,1997 

off SW Iceland in 304-1212 m 

Paranthuridae Menzies & Glynn,1968 

Calathura Norman & Stebbing,1886 

C. brachiata (Stimpson,1853) 

= Anthura brachiata Stimpson,1853 ( - Bay of Fundy) 

= Paranthura norvegica G.O. Sars,1873 

Outer Skagerrak-W & N Norway-Faeroes-off Iceland in 19-1392 m-Arctic Seas-N Greenland-Shetlands-W 

Br. Isles-off W Bretagne 

Leptanthura G.O. Sars,1897 

L. tenuis (G.O. Sars,1873) 

= Paranthura tenuis G.O. Sars,1873 

Skagerrak-Trondheim & Portugal 

L. affinis (Bonnier,1896) 

off SE, S & SW Iceland in 501-1105 m 

L. chardyi Negoescu,1992 

off SW & W Iceland in 304-1295 m 

L. micrura Kensley,1982 

off W Iceland in 1295 m, off SE Biscay in 1894-1950 m 

Paranthura Bate & Westwood,1866 

P. costana Bate & Westwood,1866 () 


Hyssuridae Wägele,1981 

* No species within the area. 

VALVIFERA G.O. Sars,1882 

Chaetiliidae Dana,1853 

Chiridotea Harger,1878 

C. caeca (Say,1818) 

= Idotea caeca Say,1818 

NE Atlantic 

C. tuftsii (Stimpson,1853) 

= Idotea tuftsii Stimpson,1853 

NE Atlantic 

Saduria Adams,1852 

= Idotaega Lockington,1877 

= Mesidotea Richardson,1905 

= Glyptonotus Eights,1853 (p.p.) 

S. entomon (Linnaeus,1758) 

= Oniscus entomon Linnaeus,1758 

Bothnian Bay-Öresund 

S. megalura (G.O. Sars,1879) 

Faeroes-Norw. Sea 

S. sabini (Krøyer,1849) 

= Idotea sabini Krøyer,1849 

NE Atlantic 

S. sibirica (Birula,1896) 

= Glyptonotus sibirica Birula,1896 

Siberia 

Idoteidae Samouelle,1819 

Idotea J.C. Fabricius,1798 

= Idothea J.C. Fabricius,1799 

I. linearis (Linnaeus,1766) 

= Oniscus linearis Linnaeus,1766 

Kattegatt-North Sea-Br. Isles-Canaries-Mediterranean 

I. emarginata (J.C. Fabricius,1793) 

= Cymothoa emarginata J.C. Fabricius,1793 

S Baltic-Bohusl-North Sea-N Norway-Faeroes-Iceland-Br. Isles-W Mediterranean 

I. metallica Bosc,1802 

Br. Isles (occasionally adrifted fr. America), Holland, Iceland? 

I. balthica (Pallas,1772) 

= Oniscus balthicus Pallas,1772 

S Bothnian Bay-Bohuslän-North Sea-Tromsö-Faeroes-Iceland-Shetlands-Br. Isles-Morocco-Mediterranean- 

Black Sea 

I. pelagica Leach,1815 

Öresund-S & W Norway-Faeroes-Iceland-Br. Isles-N & W France 

I. granulosa Rathke,1843 

Åland-Bohuslän-N Norway-Faeroes-Iceland-Br. Isles-NW France 

I. chelipes (Pallas,1766) 

= Oniscus chelipes Pallas,1766 

= Oniscus viridis Slabber,1775 

= I. angusta : G.O. Sars,1897 

Åland-Bohuslän-Tröndelag-Faeroes-Br. Isles-Ireland-France-NW Spain 

I. neglecta G.O. Sars,1897 

= Idothea neglecta G.O. Sars,1897 

Öresund-Bohuslän-N Norway-Bear Island-Iceland-Faeroes-Br. Isles-France 

Synidotea Harger,1878 

S. bathyalis Gurjanova,1955 

Arctic Sea 

S. nodulosa (Krøyer,1846) 

= Idotea nodulosa Krøyer,1846 

Greenland 

Synisoma Collinge,1917 

= Stenosoma Leach,1815 (p.p.) 

S. lancifer (Miers, 1881) 

= Stenosoma lancifera Miers,1881 

= Idotea appendiculata : Bate & Westwood,1868 

S & W Br. Isles-W France-Mediterranean 

S. acuminatum (Leach,1815) 

= Stenosoma acuminata Leach,1815 

Ireland, SW Scotland-S England-Mediterranean 

Holognathidae G. Thomson,1904 

Cleantis Dana,1849 

= Zenobiana Stebbing,1895 

= Zenobia Risso,1826, non Oken,1815 (n. nud.), nec Gray,1821 

C. prismatica (Risso,1826) 

= Zenobia prismatica Risso,1826 

Firth of Clyde, S & W Engl., W France- Mediterranean 

Arcturidae Bate & Westwood,1868 

Astacilla Cordiner,1793 

A. longicornis (Sowerby,1806) 

= Oniscus longicornis Sowerby,1806 

Öresund-Bohuslän-North Sea-N Norway-Faeroes-Iceland-Br. Isles-Engl. Channel 

A. pusilla (G.O. Sars,1873) 

= Arcturus pusilla G.O. Sars,1873 

Skagerrak-Lofoten-off SW Faeroes 

A. arietina G.O. Sars,1883 

Hardangerfjord-Norw. Sea-?Davis Strait 

A. intermedia (Goodsir,1841) 

= Leachia intermedia Goodsir,1841 

= A. affinis G.O. Sars,1897 

Skagerrak-Lofoten-S Iceland-Scotland-off W & SW Ireland 

A. granulata (G.O. Sars,1877) 

= Leachia granulata G.O. Sars,1877 

off W & N Norway-off W Faeroes-between Faeroes & Iceland 

A. deshaysi (Norman & Scott,1906) 

* Does this species exist, or is the name a synonym of another? 

Br. Isles 

Arcturella G.O. Sars,1897 

A. dilatata (G.O. Sars,1883) 

= Astacilla dilatata G.O. Sars,1883 

Kattegatt-Bohuslän-W Norway-Shetlands-Mediterranean 

A. danmoniensis (Stebbing,1874) 

= Arcturus danmoniensis Stebbing,1874 

Devon & Pembrokeshire 

Arcturus Latreille,1829 

A. scabrosus Norman,1904 


A. baffini (Sabine,1824) 

= Idothea baffini Sabine,1824 

Faeroes-Iceland-Davis Strait 

A. tuberosus G.O. Sars,1885? 

* Does this species exist? Not in the Smithsonian Internet list. 

Faeroes 

Pleuroprion Zur Strassen,1902 

P. hystrix (G.O. Sars,1877) 

= Arcturus hystrix G.O. Sars,1877 

off Lofoten-Norw. Sea-Faeroes 

P. frigidum Hansen,1916 

Greenland 

Spectarcturus Schultz,1981 

S. multispinatus Schultz,1981 ( - Greenland) 

Greenland 

Amesopodidae Stebbing,1905 


Pseudidotheidae Ohlin,1901 


Xenarcturidae Sheppard,1957 


FLABELLIFERA G.O. Sars,1882 

Limnoriidae Harger,1879 

Limnoria Leach,1814 

L. (Limnoria) lignorum (Rathke,1799) 

= Cymothoa lignorum Rathke,1799 

W Baltic-Bohuslän-North Sea-Lofoten-Faeroes-Br. Isles 

L. (Limnoria) tripunctata Menzies,1951 

S Wales & S England-tropical waters, California 

L. (Limnoria) quadripunctata Holthuis,1949 

Netherlands & S Br. Isles 

L. borealis Kussakin,1963 

Iceland, White Sea 

Sphaeromatidae H. Milne-Edwards,1840 

Bathycopea Tattersall,1905 

B. typhlops Tattersall,1905 ( - Ireland) 

@ ? 

58

off SW Faeroes-off W Ireland 

Sphaeroma Bosc,1802 

S. prideauxianum Leach,1818 

England 

S. curtum (Leach,1818) 

= Oniscus curtum Leach,1818 

England 

S. serratum (J.C. Fabricius,1787) 

= Oniscus serratum J.C. Fabricius,1787 

Holland, S & W England, Wales & Ireland-Mediterranean 

Lekanesphaera Verhoeff,1943 

L. rugicauda (Leach,1814) 

= Sphaeroma rugicauda Leach,1814 

Kalmar Sound-Öresund-N Jutland, Br. Isles-Morocco 

L. hookeri hookeri (Leach,1814) 

= Sphaeroma hookeri Leach,1814 

Kalmar Sound-Kattegatt 

L. monodi (Arcangeli,1934) 

= Sphaeroma monodi Arcangeli,1934 

Belgium, S & SE England, Wales & E Ireland-Spain, Libya 

L. teissieri (Bocquet & Lejuez,1967) 

= Sphaeroma teissieri Bocquet & Lejuez,1967 

France 

L. levii (Argano & Ponticelli,1981) 

= Sphaeroma levii Argano & Ponticelli,1981 

France 

Dynamene Leach,1814 

D. bidentata (Adams,1800) 

= Oniscus bidentata Adams,1800 () 

S & W Br. Isles & Ireland-Portugal 

D. edwardsi (Lucas,1849) 

= Naesea Edwardsi Lucas,1849 

?European Atlantic, Tunisia 

D. magnitorata Holdich,1968 

?European Atlantic, S France, NW Spain 

Campecopea Leach,1814 

C. hirsuta (Montagu,1804) 

= Oniscus hirsutus Montagu,1804 

= Campecopea cranchii Leach,1814 () 

Wales, Cornwall & Devon-W Africa 

Cymodoce Leach,1814 

C. truncata Leach,1814 () 

S & W Br. Isles-Shetlands, NW Spain 

C. emarginata Leach,1818 

S & W Br. Isles-Shetlands 

C. granulata Miers,1876 

off Ireland, New Zealand 

Cirolanidae Dana,1852 

Eurydice Leach,1815 

E. affinis Hansen,1905 

Holland, S & W Br. Isles-Mediterranean 

E. caeca Hansen,1916 

* Listed as belonging to a genus incertae sedis in the Smithsonian Internet list. 

off S Iceland 

E. grimaldii Dollfus,1888 

Bergen-N Norway-Norw. Sea-Faeroes-Iceland-Br. Isles-Portugal-Azores 

E. inermis Hansen,1890 

North Sea, England 

E. pulchra Leach,1815 

S Baltic-Bohuslän-Trondheim-Norw. Sea-W France 

E. rotundicauda Norman,1906 

NE Atlantic 

E. spinigera Hansen,1890 

Netherlands, West Indies 

E. truncata (Norman,1868) 

= Cirolana truncata Norman,1868 

North Sea-Norw. Sea 

Faucheria Dollfus & Vire,1905 

F. faucheri (Dollfus & Vire,1900) 

= Caecosphaeroma faucheri Dollfus & Vire,1900 

France 

Cirolana Leach,1818 

C. cranchii Leach,1818 ( - England) 

Firth of Clyde-Plymouth-Mediterranean 

Metacirolana Nierstrasz,1931 

M. hanseni (Bonnier,1896) 

= Cirolana hanseni Bonnier,1896 

Faeroes-off W Ireland-Bay of Biscay 

Natatolana Bruce,1981 

N. borealis (Liljeborg,1851) 

= Cirolana borealis Liljeborg,1851 

Kieler Bucht-Öresund-Bohuslän-North Sea-Trondheim-off S Lofoten-Faeroes-Shetlands-Br. Isles-W France- 

Mediterranean, in (5) 35-694 m 

N. gallica (Hansen,1905) 

= Cirolana gallica Hansen,1905 

Roscoff, NE Atlantic in 195-210 m 

N. imicola (Dollfus,1903) 

= Cirolana neglecta var. imicola Dollfus,1903 

= Cirolana schmidtii Hansen,1905 

Faeroes-Davis Strait in 872-2450 m 

Politolana Bruce,1981 

P. microphthalma (Hoek,1882) 

= Cirolana microphthalma Hoek,1882 

North Sea-Arctic Seas 

Conilera Leach,1818 

C. cylindracea (Montagu,1804) 

= Oniscus cylindracea Montagu,1804 

British Isles 

Aegidae Leach,1815 

Syscenus Harger,1880 

S. infelix Harger,1880 ( - Massachusetts) 

= Harponyx pranizoides G.O. Sars,1883 

@ (adult, ecto) Merlangius merlangus 

N Kattegatt-Bohuslän-W Norway-Faeroes-Mediterranean 

S. lilljeborgii (Bovallius,1885) 

= Rocinela lilljeborgii Bovallius,1885 

* Listed in the Smithsonian Internet list as a species of its own, but considered as a synonym of S. infelix 

Hargrave,1880 by Wahrberg. 

Sweden 

S. atlanticus Kononenko,1988 

Atlantic 

Rocinela Leach,1818 

R. danmoniensis Leach,1818 ( - England) 

@ (facultative, ecto) Gadidae 

Öresund-Bohuslän-Trondheim-Faeroes-Iceland-Shetlands-Br. Isles-Bay of Biscay 

R. dumerilii (Lucas,1849) 

= Acherusia dumerilii Lucas,1849 

@ ? 

?European Atlantic, Algeria 

Aega Leach,1815 

= Barybrotes Schiödte & Meinert,1879 ( B. indus Schiödte & Meinert,1879 - India) 

A. (Aega) arctica Lütken,1859 

@ sharks 

N Norway-Faeroes, Iceland & Arctic Seas-off W Ireland 

A. (Aega) crenulata Lütken,1859 

@ M. byrkelange & Somniosus microcephalus 

Bohuslän-S, W & N Norway-Faeroes-Iceland-Shetlands-Scotland-W Ireland 

A. (Rhamphion ?,18??) gracilipes Hansen,1895 

off SW Iceland-off NW Scotland 

A. loveni Bovallius,1886 

* A synonym of A. ventrosa M. Sars, according to Wahrberg, but included in the Smithsonian internet list. 

Sweden 

A. (Rhamphion ?,18??) microphthalma Dana,1853 

* Probably a synonym of another species, but included in the Smithsonian internet list. 

North Sea (England) 

A. (Aega) monophthalma Johnston,1834 

@ Anarhichas & Gadus 

Bohuslän-Trondheim-Faeroes-Iceland-N Br. Isles 

A. nordenskjoeldi (Bovallius,1885) 

= Aegiochus nordenskjöldi Bovallius,1885 

* A synonym of A. ventrosa M. Sars, according to Wahrberg, but included in the Smithsonian internet list. 

Arctic Ocean 

A. (Aega) psora (Linnaeus,1758) 

= Oniscus psora Linnaeus,1758 

@ Raja, Cottus, Gadus, Molva, Hippoglossus, Anarhichas etc. 

Kattegatt-Oslofjord-N Norway-Faeroes-off W Iceland-Arctic Seas-Br. Isles 

A. (Rhamphion ?,18??) stroemii Lütken,1859 

@ different fish species 

Bohuslän-Lofoten-Faeroes-Iceland-N Br. Isles-off SW Ireland 

A. (Aega) tridens Leach,1815 

@ Gadus morhua 

Oslofjord-Skagerrak-W Norway-FaeroesShetlands-Scotland-Belfast 

A. (Rhamphion ?,18??) ventrosa M. Sars,1859 

@ G. morhua, Brosme, M. byrkelange, Hippoglossus 

Bohuslän-N Norway-Faeroes-Iceland-off Shetlands-off W Ireland 

A. (Aega) bicarinata Leach,1818 

@ ? 


A. (Aega) rosacea (Risso,1816) 

= Cymothoa rocacea Risso,1816 

@ ? 

?European Atlantic, S France 

Xenuraega Tattersall,1909 

X. ptilocera Tattersall,1909 ( - off Azores) 

Ireland, S England, W Biscay, Azores 

Cymothoidae Leach,1818 

Anilocra Leach,1818 

A. physodes (Linnaeus,1758) 

= Oniscus physodes Linnaeus,1758 

@ Labridae & other fishes 

Holland & Engl. Channel-Mediterranean 

A. frontalis H. Milne-Edwards,1840 

59


Glossobius Schiødte & Meinert,1883 

G. impressus (Say,1818) 

= Cymothoa impressa Say,1818 

@ different flying fish species etc. 

N Atlantic, e.g. Køge Bugt (Denmark) 

G. auritus Bovallius,1885 

N. Atlantic 

Cymothoa Fabricius,1787 

C. scopulorum (Linnaeus,1758) 

= Oniscus scopulorum Linnaeus,1758 

Norw. Sea 

Nerocila Leach,1818 

N. orbignyi (Guérin,1832) 

= Ichthyophilus orbignyi Guérin,1832 

* Félix Édouard Guérin did not change his last name to Guérin-Méneville until 1836, so citations before that 

date should use the short name 

?European Atlantic, Greece 

N. maculata H. Milne-Edwards,1840 

* This name is probably now synonymized with another. Which? 


N. neapolitana Schiødte & Meinert,1879 

* This name is probably now synonymized with another. Which? 


Mothocya Costa,1851 

M. nana (Schiødte & Meinert,1884) 

= Irona Schiødte & Meinert,1884 nana Schiødte & Meinert,1884 

?European Atlantic, Lesser Antilles 

Anuropodidae Stebbing,1893 


Bathynataliidae Kensley,1978 


Bathynomidae Wood-Mason & Alcock,1891 


Corallanidae Hansen,1890 


Keuphyliidae Bruce,1980 


Phoratopodidae Hale,1925 

* One S Australian species only known. 

Plakarthriidae Hansen,1890 

* Only two species known, none of them NE Atlantic. 

Protognathiidae Wägele & Brandt,1988 

* Only one Antarctic species known. 

Serolidae Dana,1853 

* No species within the area. Chiefly confined to S hemisphere. 

Tecticipitidae Bruce,1993 

* Pacific family. 

Tridentellidae Bruce,1984 


MICROCERBERIDEA Lang,1961 

Microcerberidae Karaman,1933 

Microcerberus Karaman,1933 

M. remani Chappuis, Delamare-Deboutteville & Paulian,1956 lusitanicus Galhano,1970 

Portugal 

ASELLOTA Latreille,1803 

Familia Incertae sedis. 

Tole Ortmann,1901 

= Iolella Richardson,1905 

T. laciniata (G.O. Sars,1872) 

= Janthe laciniata G.O. Sars,1872 

off NW & N Norway-off Iceland-Faeroes-Davis Strait 

T. spinosa (Harger,1879) 

= Ianira spinosa Harger,1879 

off SE & NW Iceland-Davis Strait 

T. spinosissima (Stephensen,1936) 

= Ianira spinosissima Stephensen,1935 

Greenland 

T. libbeyi (Ortmann,1901) 

= Jolanthe libbeyi Ortmann,1901 

Greenland 

T. vilhelminae (Stephensen,1913) 

= Ianira vilhelminae Stephensen,1913 

W Greenland 

Aselloidea Rafinesque-Schmaltz,1815 

Asellidae Rafinesque-Schmaltz,1815 

* Only freshwater & groundwater species. Asellus Geoffroy St Hilaire,1764 aquaticus (Linnaeus,1758) is 

common in all N Europe and some species of Proasellus Dudich,1925 may be found as well. 

Atlantasellidae Sket,1980 


Stenasellidae Dudich,1924 

* Only cave & groundwater species 

Stenetrioidea Hansen,1905 

Stenetriidae Hansen,1905 

No species within the area? Usually tropical. 

Janiroidea G.O. Sars,1899 

= Paraselloidea 

Janiridae G.O. Sars,1899 

Ianira Leach,1814 

I. maculosa Leach,1814 

Öresund-Bohuslän-N Norway-Faeroes-Iceland-Greenland-Morocco 

I. alta (Stimpson,1853) 

off W Iceland-E North America 

I. tricornis (Krøyer, in Gaimard,1846) 

Jan Mayen-Spitsbergen-Davis Strait 

Ianiropsis G.O. Sars,1897 

I. breviremis (G.O. Sars,1882) 

Öresund-Bohuslän-Bergen-Faeroes-Iceland-Br. Isles 

Jaera Leach,1814 

J. albifrons albifrons Leach,1814 

Bothnian Bay-France-Norw. Sea-White Sea 

J. albifrons syei Bocquet,1950 

Bothnian Bay-Norw. Sea 

J. praehirsuta Forsman,1949 

Öresund-Norw. Sea, NW France, Britain, Greenland, New England 

J. posthirsuta Forsman,1949 

Britain, New England 

J. ischiosetosa Forsman,1949 

Bothnian Bay-Norw. Sea-White Sea, Britain 

J. forsmani Bocquet,1950 

Britanny, S & W Br. Isles 

J. nordmanni (Rathke,1837) nordica Lemercier,1958 

* in fresh or brackish water 

W Scotland-Mediterranean-Black Sea 

J. hopeana Costa,1853 

@ (ectocommensally) Sphaeroma serrratum 

Devon-Normandy-Mediterranean-Black Sea 

Janiralata Menzies,1951 ( J. davisi Menzies,1951 - California) 

J. pulchra (Hansen,1916) Brandt,1993 

= Ianthopsis pulchra (Hansen,1916) Wolff,1962 

= Janira pulchra Hansen,1916 

off SW & N Iceland-between Iceland & Greenland-Davis Strait 

Microjaera Bocquet & Levi,1955 

M. anisopoda Bocquet & Lévi,1955 

* Interstitial 

Roscoff, Scilly Isles, Eddystone, W Mediterranean 

Caecijaera Menzies,1951 

C. borealis Kussakin,1962 

@ Limnoria spp. (ectocommensalistic) 

Iceland 

Microparasellidae Karaman,1933 

= Micrjaniridae 

Microcharon Karaman,1933 

M. teissieri (Lévi,1951) 

Roscoff 

M. harrisi Spooner,1959 

Plymouth 

M. monnioti Bocquet,1970 

Roscoff 

M. coineauae Galhano,1970 

Portugal 

Acanthaspidiidae Menzies,1962 

Acanthaspidia Stebbing,1898 

A. typhlops (G.O. Sars,1879) 

off Lofoten-between Iceland & Greenland-Davis Strait 

Ianthopsis Beddard,1886 

I. pulchra (Hansen,1916) 

= Janira pulchra Hansen,1916 

Greenland 

Janirellidae Menzies,1956 

Janirella Bonnier,1896 

J. spongicola Hansen,1916 


Katianiridae Svavarsson,1987 

Katianira Hansen,1916 

K. chelifera Hansen,1916 


K. biloba Gurjanova,1930 

Faeroes, Spitsbergen, E Arctic ocean 

K. cornigera Gurjanova,1930 

Spitsbergen 

60

K. sadko Gurjanova,1946 

E Arctic ocean 

Haploniscidae Hansen,1916 

Haploniscus Richardson,1909 

H. ingolfi Wolff,1962 

Faeroes, S of Jan Mayen in 2465m 

H. bicuspis bicuspis (G.O. Sars,1877) 

Faeroes-off W Norway-off Iceland, Faeroes & Jan Mayen 

H. bicuspis tedipus Wolff,1962 

SW of Iceland in 1505 m 

H. spinifer Hansen,1916 

off SW Iceland-Davis Strait 

Chauliodoniscus Lincoln,1985 

C. armadilloides (Hansen,1916) 

= Haploniscus armadilloides Hansen,1916 

off S Iceland 

Joeropsidae Nordenstam,1933 

Joeropsis Koehler,1885 

J. brevicornis brevicornis Koehler,1885 

S & W Br. Isles, Norway 

Munnidae G.O. Sars,1899 

Munna Krøyer,1839 

M. acanthifera Hansen,1916 

Faeroes-around Iceland-Davis Strait 

M. armoricana Carton,1962 

France 

M. boeckii Krøyer,1839 

Bohuslän-North Sea-W Norway-Faeroes-Iceland & Arctic Seas-Scotland 

M. coeca Gurjanova,1930 

E of Spitsbergen 

M. fabricii Krøyer,1846 

non = M. fabricii : G.O. Sars,1899 

Belt Sea-North Sea-W Norway-Iceland & Arctic Seas-Normandie 

M. groenlandica Hansen,1916 

W & E Greenland 

M. hanseni Stappers,1911 

* possibly a synonyme of M. limicola 

Faeroes-around Iceland-Arctic Seas 

M. kroyeri Goodsir,1842 

Bay of Kiel-S & W Norway-North Sea-Iceland-Br. Isles-Engl. Channel 

M. limicola G.O. Sars,1866 

Bohuslän-North Sea-N Norway-Faeroes-Iceland & Arctic Seas-Ireland-Engl. Channel 

M. minuta Hansen,1916 

= M. fabricii : G.O. Sars,1899, non Krøyer,1846 

Öresund-Bohuslän-W & N Norway-Faeroes-Iceland & Arctic Seas-Ireland 

M. palmata Liljeborg,1851 

S Norway 

M. pellucida Gurjanova,1930 

Murman coast 

M. roemeri Gurjanova,1930 

?Spitsbergen-arctic Seas 

M. spitzbergensis Gurjanova,1930 

Spitsbergen-arctic Seas 

Uromunna Menzies,1962 

U. petiti (Amar,1948) 

= Munna petiti Amar,1948 

Scilly Islands-Mediterranean 

Paramunnidae Vanhöffen,1914 

Paramunna G.O. Sars,1866 

P. bilobata G.O. Sars,1866 () 

Bohusl-Sognefjord-Br. Isles 

P. typica (Tattersall,1905) 

= Metamunna typica Tattersall,1905 

W of Ireland in 220-300 m 

Pleurogonidae Nordenstam,1933 

Pleurogonium G.O. Sars,1882 

= Pleuracantha G.O. Sars,1864 (homonym) 

P. inerme G.O. Sars,1882 

Öresund-Bohuslän-North Sea-W Norway-Faeroes-Iceland & Arctic Seas-Br. Isles 

P. rubicundum (G.O. Sars,1864) 

= Pleuracantha rubicunda G.O. Sars,1864 

S Baltic-Bohuslän-N Norway-Faeroes-Arctic Seas-N Br. Isles 

P. spinosissimum (G.O. Sars,1868) 

= Pleuracantha spinosissima G.O. Sars,1868 

Öresund-Bohuslän-N Norway-Faeroes-Iceland & Arctic Seas-N Br. Isles 

P. intermedium Hansen,1916 

Faeroes-N Iceland 

P. pulchrum Hansen,1916 

Faeroes-W Iceland 

P. polaris Svavarson,1988 

* Is this a synonym of another species? 

abyssal depths in North Polar Sea 

P. latimanum Hansen,1916 

Davis Strait 

Dendrotiidae Vanhöffen,1914 

Dendromunna Menzies,1962 ( D. spinipes Menzies,1962 - S Atlantic) 

D. compsa Lincoln & Boxshall,1983 


Dendrotion G.O. Sars,1872 

D. elegans Lincoln & Boxshall,1983 


D. setosum Lincoln & Boxshall,1983 


D. spinosum G.O. Sars,1872 () 

Skagerrak-N North Sea-Hardangerfjord-Faeroes-Iceland & Arctic Seas 

D. paradoxum Hansen,1916 

Faeroes-off SW Iceland 

Acanthomunna Beddard,1886 

= Pseudomunna Hansen,1916 

A. hystrix (Hansen,1916) 

= Pseudomunna hystrix Hansen,1916 ( of Pseudomunna Hansen,1916) 


Mictosomatidae Wolff,1965 

Mictosoma Wolff,1965 

= Schistosoma Hansen,1916, non Weinland,1858, nec Brady,1877 

M. ramosum (Hansen,1916) 

off S Iceland 

Macrostylidae Hansen,1916 

Macrostylis G.O. Sars,1864 

M. longiremis (Meinert,1890) 

Bohuslän-Skagerrak-SW Iceland 

M. spinifera G.O. Sars,1864 

Kattegatt-Bohuslän-W Norway-Iceland & Arctic Seas-off SW Iceland-Shetlands 

M. subinermis Hansen,1916 

Faeroes-off N & E Iceland 

M. elongata Hansen,1916 

= M. longipes Hansen,1916 (male) 

off S & E Iceland 

M. abyssicola Hansen,1916 

E Arctic ocean, Davis Strait 

Ischnomesidae Hansen,1916 

Ischnomesus Richardson,1908 

I. bispinosus (G.O. Sars,1868) 

= Ischnosoma bispinosa G.O. Sars,1868 

Bohuslän-Lofoten-Faeroes-W Ireland-Mediterranean 

I. norvegicus Svavarson,1984 

the slopes of Norw. Sea 

Haplomesus Richardson,1909 

H. quadrispinosus (G.O. Sars,1879) 

Faeroes-off Iceland-off Lofoten, S of Jan Mayen, E Arctic ocean 

H. angustus Hansen,1916 

off N Iceland, SE Greenland, E Arctic ocean 

Helomesus Wolff,1962 ( Stylomesus menziesi Birstein,1960 - abyssal Pacific) 

H. gorbunovi (Gurjanova,1946) 

= Haplomesus gorbunovi Gurjanova,1946 

depths of Norw., North Polar & Greenland Seas 

Heteromesus Richardson,1909 

H. dentatus Hansen,1916 


H. schmidtii Hansen,1916 

off S Iceland 

H. frigidus Hansen,1916 

Faeroes-off N & E Iceland 

H. longiremis Hansen,1916 

E Arctic Ocean, Davis Strait 

H. greeni (Tattersall,1905) 

off N Ireland 

Nannoniscidae Hansen,1916 

Nannoniscus G.O. Sars,1869 

N. aequiremis Hansen,1916 

off S Jan Mayen 

N. affinis Hansen,1916 


N. analis Hansen,1916 

Davis Strait 

N. arcticus Hansen,1916 

off Jan Mayen-E Greenland 

N. arctoabyssalis Just,1980 

abyssal Arctic Seas 

61

N. bicuspis G.O. Sars,1877 

Norw. Sea 

N. inermis Hansen,1916 

Davis Strait 

N. laticeps Hansen,1916 

off N Iceland 

N. minutus Hansen,1916 

Davis Strait 

N. oblongus G.O. Sars,1870 

off Lindesnæs-off Lofoten-Faeroes-off W & SW Iceland-Arctic Seas 

N. plebejus Hansen,1916 


N. profundus Svavarson,1982 

deep part of Norw. Sea & Arctic Basins 

N. reticulatus Hansen,1916 

off N Iceland, N Greenland, Kara Sea 

N. simplex Hansen,1916 

off W & SW Iceland 

N. spinicornis Hansen,1916 

off S Jan Mayen in 2465 m 

Nannoniscoides Hansen,1916 

N. angulatus Hansen,1916 () 

Faeroes, N Greenland, Kara Sea 

Rapaniscus Siebenaller & Hessler,1981 

R. crassipes (Hansen,1916) 

= Nannoniscus crassipes Hansen,1916 

off Lofoten 

Regabellator Siebenaller & Hessler,1981 

R. armatus (Hansen,1916) 

Davis Strait in 3521 m 

Exiliniscus Siebenaller & Hessler,1981 

E. hanseni (Just,1970) 

= Nannoniscus hanseni Just,1970 

N Greenland (Jørgen Brønlund Fjord) in 80-90m 

Austroniscus Vanhöffen,1914 

A. norbi Svavarson,1982 

deep part of Norw. Sea & Greenland Sea 

A. groenlandicus (Hansen,1916) 

= Nannoniscella groenlandica Hansen,1916 

Greenland 

Pseudomesidae Hansen,1916 

Pseudomesus Hansen,1916 

P. brevicornis Hansen,1916 () 

off N Iceland 

P. sp. (nov. sp.) : Just,1970 

N Greenland 

Desmosomatidae G.O. Sars,1897 

Desmosomatinae G.O. Sars,1897 

Mirabilicoxa Hessler,1970 

M. fletcheri Paul & George,1975 

deep Arctic ocean 

Whoia Hessler,1970 

W. angusta (G.O. Sars,1899) 

= Desmosoma angustum G.O. Sars,1899 

Bohuslän-Skagerrak-Trondheim Fjord 

W. dumbshafensis Svavarson,1988 

deep parts of Norw. Sea 

Desmosoma G.O. Sars,1864 

D. lineare G.O. Sars,1864 () 

Bohuslän-N Norway-Arctic Seas, e.g. N Greenland 

D. strombergi Svavarsson,1987 

Faeroes-slopes & depth of Norw. Sea 

Chelator Hessler,1970 

C. chelatum (Stephensen,1915) 

= Desmosoma chelator Stephensen,1915 

* pelagic species. 

W Mediterranean 

C. insignis (Hansen,1916) 

= Desmosoma insignis Hansen,1916 

Davis Strait 

Eugerda Meinert,1890 

E. arctica Svavarson,1988 


E. fragilis (Kussakin,1965) 

NE Atlantic 

E. reticulata (Gurjanova,1946) 

= Desmosoma reticulata Gurjanova,1946 

Faeroes-depths of Norw. Sea & North Polar Sea 

E. tenuimana (G.O. Sars,1868) 

= Desmosoma tenuimanum G.O. Sars,1868 

= Eugerda globiceps Meinert,1890 () 

non = Eugerda tenuimana : G.O. Sars,1897 

Öresund-Bohuslän-Skagerrak-W Norway-Faeroes-Iceland-E Greenland 

Pseudogerda Kussakin,1965 

P. filipes (Hult,1936) 

= Desmosoma filipes Hult,1936 

Kattegatt-Bohuslän-S Norway-Ireland-Mediterranean 

P. intermedia (Hult,1936) 

= Desmosoma intermedium Hult,1936 

Kattegatt-Bohuslän-W Norway-Faeroes-W Greenland 

P. latipes (Hansen,1916) 

= Desmosoma latipes Hansen,1916 

Davis Strait in ≥200 m 

Echinopleura G.O. Sars,1897 

E. aculeata (G.O. Sars,1864) 

Bohuslän-Arctic Seas 

Eugerdellatinae Hessler,1970 

Desmosomella Kusskin,1965 

D. armata (G.O. Sars,1864) 

= Desmosoma armatum G.O. Sars,1864 

* May belong to Eugerdella Kussakin,1965? 

Bohuslän-Trondheim-E & N Greenland 

Eugerdella Kussakin,1965 

E. coarctata (G.O. Sars,1899) 

= Desmosoma coarctata G.O. Sars,1899 

Bohuslän-Trondheim-Davis Strait 

E. hessleri Just,1980 

slopes of Norw. Sea-depthts of North Polar Sea in ≥800 m 

E. polita (Hansen,1916) 

= Desmosoma politum Hansen,1916 


E. zenkewitschi (Gurjanova,1946) 

= Desmosoma zenkewitschi Gurjanova,1946 

sublittoral Arctic Seas 

Prochelator Hessler,1970 

P. lateralis (G.O. Sars,1899) 

= Eugerda lateralis G.O. Sars,1899 

Bohuslän-Skagerrak-Trondheim-?Davis Strait 

Oecidiobranchus Hessler,1970 

O. nanseni Just,1980 

Faeroes-slopes and depths of Norw., North Polar & Greenland Seas 

O. plebejum (Hansen,1916) 

= Desmosoma plebejum Hansen,1916 

off E & NE Iceland, N Greenland 

O. polare (Gurjanova,1946) 

= Desmosoma polaris Gurjanova,1946 

sublittoral Arctic Seas 

Cryodesma Svavarson,1988 

C. agnari Svavarson,1988 


Ilyarachnidae Hansen,1916 

Ilyarachna G.O. Sars,1869 

I. longicornis (G.O. Sars,1864) 

= Mesostenus longicornis G.O. Sars,1869 

= I. plunketti Tattersall,1905 

Bohuslän-?W Norway-Iceland & Arctic Seas, e.g. E & N Greenland 

I. torleivi Svavarsson,1988 

Faeroes-slopes of Norw. Sea 

I. bergendahli Ohlin,1901 

Faeroes-Spitsbergen-E Greenland 

I. hirticeps G.O. Sars,1870 

= I. denticulata G.O. Sars,1899 

Stavanger-N Norway-Spitsbergen-off Iceland-Faeroes-Davis Strait 

I. dubia Hansen,1916 

off NE Iceland in 1666-1902m 

I. thori Wulff,1962 

SW of Faeroes, S of Iceland (in 216-820m) 

I. derjugini Gurjanova,1946 


I. frami Just,1980 


Pseudarachna G.O. Sars,1897 

P. hirsuta (G.O. Sars,1864) 

Kattegatt-Bohuslän-Trondheim-Faeroes-N Br. Isles 

Aspidarachna G.O. Sars,1897 

A. clypeata (G.O. Sars,1870) 

= Ilyarachna clypeata G.O. Sars,1870 

N Norway, Faeroes-Arctic Seas 

Echinozone G.O. Sars,1897 

E. coronata (G.O. Sars,1870) 

= Ilyarachna coronata G.O. Sars,1870 

W & N Norway-between Norway & Shetlands-Faeroes-off S & SW Iceland-Davis Strait 

E. arctica Hansen,1916 

62

= Ilyarachna arctica (Hansen,1916) : Wolff,1962 

Faeroes-Jan Mayen, N Greenland, Kara Sea 

Eurycopidae Hansen,1916 

Eurycope G.O. Sars,1864 

E. cornuta G.O. Sars,1864 

Bohuslän-N Norway-Faeroes-Iceland & Arctic Seas 

E. producta G.O. Sars,1868 

Bohuslän-N Norway-Faeroes-Iceland & Arctic Seas-off W Ireland 

E. inermis Hansen,1916 

Faeroes-Norw. Sea-Arctic Seas 

E. hanseni Ohlin,1901 

Faeroes-Norw. Sea-Arctic Seas 

E. brevirostris Hansen,1916 

Faeroes-off Iceland, N Greenland 

E. parva Bonnier,1896 

off SW Faeroes-Bay of Biscay-Davis Strait 

E. cryoabyssalis Just,1980 

deep water in North Polar Sea 

E. dahli Svavarson,1987 


E. ratmanovi Gurjanova,1946 

deep Arctic waters 

Munneurycope Stephensen,1912 

M. murrayi (Walker,1903) 

= Eurycope murrayi Walker,1903 


off SW Faeroes-off S Iceland- off W Ireland 

M. incisa (Gurjanova,1946) 

the depths of Norw. Sea, off NW Spitsbergen 

Munnopsurus Richardson,1912 

M. giganteus giganteus (G.O. Sars,1877) 

off W Norway - off N & SE Faeroes-off E & N Iceland-Arctic Seas 

M. longipes (Tattersall,1905) 

off SW Faeroes-off W Ireland-Davis Strait 

Tytthocope Wilson & Hessler,1981 

T. megalura (G.O. Sars,1872) 

= Eurycope megalura G.O. Sars,1872 

Stavanger-Hardangerfjord-off Faeroes-off SW Iceland 

T. pygmaea (G.O. Sars,1870) 

= Eurycope pygmaea G.O. Sars,1870 

Bohuslän-Lofoten 

T. laktionovi (Gurjanova,1946) 

Slopes of North Polar Seas 

T. neupokoevi (Gurjanova,1946) 

= Eurycope neupokoevi Gurjanova,1946 

slopes of North Polar Seas 

Disconectes Wilson & Hessler,1981 

D. phalangium (G.O. Sars,1864) 

= Eurycope phalangium G.O. Sars,1864 

Bohuslän-N Norway-Faeroes-W Ireland-Scotland-Davis Strait 

D. furcatus (G.O. Sars,1870) 

= Eurycope furcata G.O. Sars,1870 

E Skagerrak-Stavanger-off Lofoten-Faeroes-off SW Iceland-Davis Strait 

D. latirostris (G.O. Sars,1882) 

= Eurycope latirostris G.O. Sars,1882 

Norw. Sea - W of Ireland 

D. coxalis Kussakin,1983 


Baeonectes Wilson,1982 

B. muticus (G.O. Sars,1864) 

= Eurycope mutica G.O. Sars,1864 

Öresund-Bohuslän-N Norway-Faeroes-Iceland & Arctic Seas, e.g. N Greenland-Br. Isles 

Munnopsidae G.O. Sars,1869 

Lipomera Tattersall,1905 

L. (Lipomera) lamellata Tattersall,1905 () 

Ireland 

Munnopsis M. Sars,1861 

= Munnopsoides Tattersall,1905 (p.p.) 

= Pseudomunnopsis Hansen,1916 

M. typica M. Sars,1861 () 

Bohuslän-N Norway-Faeroes-Iceland & Arctic Seas 

M. beddardi (Tattersall,1905) 

= Munnopsoides beddardi Tattersall,1905 

Faeroes-W Ireland in ≥364 m 

M. eximius (Hansen,1916) 

= Munnopsoides eximius Hansen,1916 

Faeroes-off S Iceland 

Paramunnopsis Hansen,1916 

P. oceanica (Tattersall,1905) 

= Munnopsis oceanica Tattersall,1905 


off W Ireland-Davis Strait 

P. justi Svavarson,1988 

deepest parts of Greenland & North Polar Seas 

Abyssianiridae Menzies,1956 


Echinothambematidae Menzies,1956 


Haplomunnidae Wilson,1976 


Mesosignidae Schultz,1969 


Pleurocopidae Fresi & Schiecke,1972 


Thambematidae Stebbing,1912 

Thambema Stebbing,1912 

T. amicorum Stebbing,1912 () 

W of Ireland in ≥1993 m 

T. fiatum Harrison,1987 

Rockall Trench in 1800 m 

T. golanachum Harrison,1987 

S Rockall Trench in ≥1330 m 

T. tanum Harrison,1987 

S Rockall Trench in 2875 m 

Gnathostenetroidoidea Kussakin,1967 


Gnathostenetroidoidae Kossakin,1967 

CALABOZOIDEA van Lieshout,1983 

* Not known from Europe. 

Calabozoidae van Lieshout,1983 

ONISC(O)IDEA Latreille,1803 

TYLOMORPHA Vandel,1943 

* Only terrestrial? 

Tylidae H. Milne-Edwards,1940 

LIGIAMORPHA Vandel,1943 

DIPLOCHAETA Vandel,1957 

Ligidae Brandt,1883 

Ligia J.C. Fabricius,1798 

L. oceanica (Linnaeus,1767) 

Terrestrial sea-shore inhabitant in Sweden, Norw., Denm., Icel., Faeroes, Br. Isles etc. 

L. exotica Roux,1828 

Britain, Mediterranean shores 

Mesoniscidae Verhoeff,1908 

* Only terrestrial? 

SYNOCHAETA Legrand,1946 

* 2 superfamilies, 6 families. Only terrestrial? 

CRINOCHAETA Legrand,1946 

* 2 superfamilies, 25 families. Only terrestrial? 

PHREATOICIDEA Stebbing,1893 

* 3 recent families. Not marine & not European. 

EPICARIDEA Latreille,1831 

* Yip,1984 found an unidentified miktoniscus larvae of an Epicaridean Isopod in Pleurobrachia pileus at 

Ireland 

Bopyroidea Rafinesque-Schmaltz,1815 

Bopyridae Rafinesque-Schmaltz,1815 

@ branchial cavity of Decapods 

Anisarthrus Giard,1907 

A. pelseneeri Giard,1907 

@ abdomen of Athanas nitescens (Leach) 

Galway area, NW France 

Asymmetrione R. & M. Codreanu & Pike,1965 

= Megachelione Bourdon,1968 

A. dardani Bourdon,1968 

@ Dardanus arrosor (Herbst) 

Morocco 

A. foresti (Bourdon,1968) 

= Megachelione foresti Bourdon,1968 

@ Paguristes oculatus Fabricius 

Baleares, Sète, Tunisia 

Athelges Hesse,1861 

A. bilobus G.O. Sars,1898 

=? A. lorifera Hesse,18?? 

@ abdomen of Pagurus cuanensis Bell 

N Kattegatt, Bohuslän, Skagerrak, Br. Isles, ?France 

A. cladophorus Gerstaecker,1862 

@ ? 

Engl. Channel 

A. paguri (Rathke,1843) 

@ abdomen of Pagurus bernhardus (Linnaeus) 

Öresund-Bohuslän-Skagerrak- North Sea, Britain, NW France 

A. tenuicaudis G.O. Sars,1898 

@ abdomen of Anapagurus chiroacanthus (Liljeborg) 

N Kattegatt, Bohuslän, S Norway, N Norway, North Sea-Norw. Sea-Scotland 

63

A. prideauxi Giard & Bonnier,1890 

@ Pagurus prideauxi (Leach) 

Br. Isles, Mediterranean 

Bopyrus Latreille,1802 

B. squillarum Latreille,1802 

= Monoculus crangorum O. Fabricius,1798 (nomen oblitum?) 

= Bopyrus fougerouxi Giard & Bonnier,1890 

= Bopyrus rathkei Giard & Bonnier,1890 (nom. nudum) 

= Bopyrus helleri Giard & Bonnier,1890 

= Bopyrus treillanus Giard & Bonnier,1890 (nom. nudum) 

= Bopyrus xiphias Giard & Bonnier,1890 

@ Palaemon serratus (Pennant), P. elegans Rathke, P. adspersus Rathke, P. squilla (Linnaeus) & P. 

xiphias (Risso) 

Öresund, Netherlands, Britain, Ireland, Roscoff, Brest, Arcachon, Portugal, Spain, Mediterranean, Black Sea 

Bopyrina Kossman,1881 

B. ocellata (Czerniavsky,1868) 

= Bopyrus ocellatus Czerniavsky,1868 

= Bopyrus virbii Walz,1881 

= Bopyrina giardi Bonnier,1900 

@ Hippolyte varians Leach, H. inermis Leach & H. longirostris Czerniavsky 

SW England, Galway, Dinard, Roscoff, Arcachon, Mediterranean, Black Sea 

Bopyrissa Nierstrasz & Brender a Brandis,1931 

= Urocryptella R. & M. Codreanu,1963 

B. fraissei (Carayon,1943) 

= Pseudione fraissei Kossmann,1886 (nomen nudum) 

= Pseudione fraissei Carayon,1943 

@ Clibanarius erythropus (Latreille) 

Arcachon 

B. diogeni (Popov,1927) 

= Pseudione diogeni Popov,1927 

@ Diogenes pugilator (Roux) 

Jersey, Roscoff, Arcachon, Mediterranean, Black Sea 

Bopyroides Stimpson,1864 

B. hippolytes (Krøyer,1838) 

= Bopyrus hippolytes Krøyer,1838 

= Bopyroides acutimarginatus Stimpson,1864 

= Bopyroides sarsi Bonnier,1900 

@ Spirontocaris lilljeborgi (Danielssen), S. spinus (Sowerby), Eualus gaimardi (H. Milne-Edwards), 

Lebbeus polaris (Sabine) & ? Pandalus spp. 

Bohuslän-Skagerrak-Trondheim-Norw. Sea-Iceland-Davis Strait 

B. cluthae (Scott,1902) 

= Pleurocrypta cluthae Scott,1902 

= Pleurocrypta patiencei Scott,1902 

=? Bopyroides furcata Norman,1905 (nomen nudum) 

= Bopyroides sarsi : Allen,1965, non Bonnier,1900 

@ Pandalina brevirostris (Rathke) & Caridion gordoni Bate 

Northumberland, Plymouth, Clyde Sea, Minch, Roscoff, Galway Bay 

Cancricepon Giard & Bonnier,1887 

C. elegans (Giard & Bonnier,1886) 

= Cepon elegans Giard & Bonnier,1886 

@ Pilumnus hirtellus (Linnaeus) 

Norfolk, The Wash, Roscoff, Arcachon 

C. pilula (Giard & Bonnier,1886) 

= Cepon pilula Giard & Bonnier,1886 

@ Xantho incisus Leach 

Roscoff, Concarneau 

Epipenaeon Nobili,1906 

E. ingens ingens Nobili,1906 

@ Penaeus semisulcatus De Haan 

Turkish Mediterranean, Red Sea 

Ergyne Risso,1816 

E. cervicornis Risso,1816 () 

@ Liocarcinus arcuatus (Leach) 

Roscoff, Spain, Mediterranean 

Grapsicepon Giard & Bonnier,1887 

G. edwardsi Giard & Bonnier,1888 

@ Planes minutus (Linnaeus) 

Roscoff, Sargasso Sea 

Gyge Cornalia & Panceri,1861 

G. branchialis Cornalia & Panceri,1861 () 

= G. galatheae Bate & Westwood,1868 

@ Upogebia stellata (Leach), U. deltaura Leach & U. pusilla (Petagna) 

Frederikshavn, Skagerrak, Engl. Channel, Clyde Sea, Irish Sea, Arcachon, Mediterranean, Black Sea 

Hemiarthrus Giard & Bonnier,1887 

= Phryxus Rathke,1843, non Rafinesque-Schmaltz,1815, nec Huebner,1819 

H. abdominalis (Krøyer,1841) 

= Bopyrus abdominalis Krøyer,1841 

@ abdomen of e.g. Pandalus, Spirontocaris & Crangon allmani 

Öresund-Bohuslän-North Sea-N Norway-Iceland-Arctic Seas 

Ione Latreille, in Cuvier,1817 

I. thoracica (Montagu,1808) 

= Oniscus thoracica Montagu,1808 

@ Callianassa subterranea (Montagu) & C. tyrrhena (Petagna) 

Belgium, Ireland-S & W Br. Isles-Mediterranean 

Pleurocrypta Hesse,1865 

P. galateae Hesse,1865 ( -Mediterranean) 

= Pleurocrypta nexa Stebbing, in Herdman,1894 

= Pleurocrypta marginata G.O. Sars,1899 

= Pleurocrypta hessei Nierstrasz & Brender-à-Brandis,1931 

@ Galathea squamifera Leach, G. nexa Embleton & G. dispersa Bate 

Bohuslän, Skagerrak, North Sea, Bergen, Trondheim, Shetlands, Br. Isles, Ireland, Bretagne, Mediterranean 

P. longibranchiata (Bate & Westwood,1868) 

= Phryxus longibranchiatus Bate & Westwood,1868 

@ Galathea squamifera Leach, G. nexa Embleton & G. dispersa Bate 

Bohusl-Skagerrak-North Sea, Manger, Br. Isles, Bretagne, S France 

P. microbranchiata G.O. Sars,1898 

= P. intermedia Giard & Bonnier,1890 (nomen nudum) 

= P. intermedia Bonnier,1900 

@ Galathea intermedia Liljeborg 

Kattegatt-Bohuslän-Skagerrak-S Norway-Bergen-North Sea, Shetlands, Br. Isles, Ireland, Bretagne, Spain, 

Adriatic 

P. piriformis Bourdon,1968 

@ Galathea strigosa (Linnaeus) 

Spain (Bae de Cadaqués) 

P. porcellanae Hesse,1876 

= P. porcellanae longicornis Hesse,1876 

= P.perezi Nierstrasz & Brender-à-Brandis,1926 

= Pseudione convergens Stock,1960 

@ Pisidia longicornis (Linnaeus) 

Northumberland, W Br. Isles, Ireland, Bretagne, Morocco 

P. strigosa Bourdon,1968, ex Giard & Bonnier MS 

= Pleurocrypta strigosa Giard & Bonnier,1890 (nomen nudum) 

@ Galathea strigosa (Linnaeus) 

Jersey, Roscoff, Spain, Mediterranean 

Pleurocryptella Bonnier,1900 

P. formosa (Giard & Bonnier,1888) 

= Pleorocrypta formosa Giard & Bonnier,1888 

@ Ptychogaster formosus A. Milne-Edwards 

off Ireland (Porcupine Bank) & Canaries 

P. superba Bourdon,1981 

@ ? 

Denmark 

Pliophrixus Caroli,1930 

* This genus & species is not listed in the Smithsonian Internet list. Bourdon 1968 does not mention it either. 

P. philonika (Caroli,1930) 

@ Processa spp. 

Britain 

Pseudione Kossmann,1881 

P. hyndmanni (Bate & Westwood,1868) 

= Phryxus hyndmanni Bate & Westwood,1868 

= Pseudione proxima Bonnier,1900 

@ Pagurus bernhardus (Linnaeus), P. pubescens (Krøyer), P. prideauxi (Leach), P. cuanensis (Bell), 

Anapagurus laevis (Bell) & A. chiroacanthus (Lilljeborg) 

Öresund-Kattegatt-Bohuslän-Skagerrak, Trondheim-Tromsø, Iceland-Faeroes-Br. Isles-Irelanf, Belgium- 

Bretagne, ?Mediterranean 

P. affinis (G.O. Sars,1882) 

= Pleurocrypta affinis G.O. Sars,1882 

= Pseudione hoylei Bonnier,1900 

@ Pandalus montagui Leach, Dichelopandalus bonnieri (Kinahan) & Plesionika spp. 

Drøbak, Skagerrak-North Sea, Trondheim, Tromsø, Br. Isles, Ireland, Engl. Channel, Biscay, Mediterranean 

P. crenulata G.O. Sars,1898 

= Palaegyge insignis Giard & Bonnier,1890 (nomen nudum) 

= Pseudione insignis Bonnier,1900 

@ Munida tenuimana M. Sars & M. rugosa (Fabricius) 

Skagerrak, in 500m-North Sea, Bergen, Br. Isles, France 

P. dohrni Bonnier,1900 

= Palaegyge dohrni Giard & Bonnier,1890 (nomen nudum) 

=? P. callianassae Kossman,1881 (nomen nudum) 

@ Callianassa truncata Giard & Bonnier 

Gulf of Naples 

P. borealis Caspers,1939 


@ Callianassa tyrrhena (Petagna,1792) 

Helgoland 

P. caspersi Gruner,1966 

= P. tuberculata Caspers,1939, non Richardson,1904 


@ Callianassa tyrrhena (Petagna,1792) 

* This species is not mentioned in the Smithsonian internet list, but P. callianassae Kossman,1881 is. 

Helgoland 

P. confusa confusa (Norman,1886) 

= Gyge confusa Norman,1886 

@ Galathea dispersa Bate 

Shetlands, Ireland, Portugal 

Progebiophilus Codreanu & Codreanu,1963 

P. euxinicus (Popov,1927) 

= Pseudione euxinica Popov,1927 

= Gyge branchialis var. arcassonensis Carayon,1943 

@ Upogebia deltaura Leach, U. pusilla (Petagna) & U. stellata (Montagu) 

Clyde Sea, Irish Sea, Galway Bay, Roscoff, Arcachon, Mediterranean, Black Sea 

Scyracepon Tattersall,1905 

S. tuberculosa Tattersall,1905 ( - Ireland) 

@ Rochinia carpenteri (Norman) 

Ireland & ? France 

Urobopyrus Richardson,1904 

U. processae Richardson,1904 ( - Brazil) 

= U. provisorius Rouch & Taberly,1961 

@ Processa edulis (Risso), P. acutirostris Nouvel & Holthuis & other Processa spp. 

Roscoff, Arcachon, Mediterranean, Florida, Brazil 

Entoniscidae F. Müller,1871 

@ visceral cavity of Decapoda 

Portunion Giard & Bonnier,1887 

P. maenadis (Giard,1886) 

= Entoniscus maenadis Giard,1886 

@ visceral cavity of crabs and anomurans, usually Carcinus maenas (Linnaeus) 

Isefjord, Bohuslän-Skagerrak, Engl. Channel, W France, Gulf of Naples 

P. kossmanni (Giard & Bonnier,1886) 

= Entoniscus kossmanni Giard & Bonnier,1886 

@ Portumnus latipes (Pennant) 

64

Roscoff 

P. moniezi (Giard,1878) 

=Entoniscus moniezii Giard,1878 

@ Liocarcinus puber (Linnaeus) 

Roscoff 

P. pusillus Pérez,19?? 

@ Liocarcinus pusillus (Leach) 

Roscoff 

P. salvatoris (Kossmann,1881) 

= Entione salvatoris Kossmann,1881 

@ Liocarcinus arcuatus (Leach) 

Roscoff 

Priapon ?,1??? 

P. fraissei (Giard & Bonnier,1886) 

= Entoniscus fraissei Giard & Bonnier,1886 

@ Liocarcinus holsatus (Fabricius) 

Roscoff 

Entionella Miyashita,1941 ( E. fluviatilis Miyashita,1941 - Japan, @ Eriocheir japonicus de Haan) 

E. monensis Hartnoll,1960 

@ Eurynome aspera (Pennant) 

Br. Isles 

Pinnotherion Giard & Bonnier,1889 

P. vermiforme Giard & Bonnier,1889? 

@ Pinnotheres pisum (Linnaeus) 

Cornwall, Bretagne 

Cancrion Giard & Bonnier,1887 

C. miser Giard & Bonnier,1886 

@ Pilumnus hirtellus (Linnaeus) 

Roscoff 

C. floridus Giard & Bonnier,1886 

@ Xantho incisus Leach 

Roscoff 

C. pilipedi Veillet & Bourdon,19?? 

@ Xantho pilipes A. Milne-Edwards 

Roscoff 

Entoniscus Fritz Müller,1862 ( E. porcellanae F. Müller,1862 - Brazil @ Porcellana sp.) 

E. muelleri Giard & Bonnier,1886 

@ Pisidia longicornis (Pennant) 

Concarneau 

Synalpheion Coutière,1908 ( S. giardi Coutière,1908) 

Dajidae G.O. Sars,1882 

@ Eucarida 

Dajus Krøyer,1849 

D. mysidis Krøyer,1842 

@ Mysis oculata (Fabricius), M. relicta Lovén & M. mixta Lilljeborg 

N Norway-Iceland, E & N Greenland 

D. profundus Hansen,1916 

@ Paramblyops rostrata Holt & Tattersall 


Branchiophryxus Caullery,1897 

B. nyctiphanae Caullery,1897 

@ gills of Meganyctiphanes norvegica (M. Sars) 

Biscay 

Prodajus Bonnier,1903 

P. lobiancoi Bonnier,1903 

@ Gastrosaccus lobatus Nouvel,1951 

Roscoff 

P. ostendensis Gilson,1909 

@ Gastrosaccus spinifer (Goës,1864) 

S &SE North Sea 

Aegophila Bresciani,1966 

A. socialis Bresciani,1966 

@ Aega ventrosa M. Sars 

Greenland 

Aspidophryxus G.O. Sars,1882 

A. peltatus G.O. Sars,1882 

@ dorsum of mysids, e.g. Erythrops spp., Parerythrops obesa (G.O. Sars) & Mysidopsis didelphys 

(Norman) 

Bohuslän-Skagerrak-Lofoten-North Sea-Iceland & Arctic Seas, Scotland, Ireland 

Notophryxus G.O. Sars,1882 

N. ovoides G.O. Sars,1882 

@ dorsum (usually) of mysids & euphausids, e.g. Amblyops abbreviata (M. Sars) 

Hardangerfjord, Trondheimsfjord, North Sea-Norw. Sea 

N. clypeatus G.O. Sars,1885 

@ dorsum of Pseudomma roseum & Pseudomma truncatum 

off N Iceland 

Holophryxus Richardson,1905 

H. richardii Koehler,1911 

@ Sergestes arcticus Krøyer 

Bergen-Iceland-W France-Azores 

H. acanthephyrae Stephensen,1912 

@ Acanthephyra purpurea A. Milne-Edwards 

Cape Farewell 

Heterophryxus G.O. Sars,1885 

Zonophryxus Richardson,1903 

Arthrophryxus Richardson,1908 

Colophryxus Richardson,1908 

Prophryxus Richardson,1909 

Allophryxus Koehler,1911 

Paradajus Nierstrasz & Brender à Brandis,1931 

Streptodajus Nierstrasz & Brender à Brandis,1931 

Cryptoniscina 

@ non Eucaridean Crustacea 

* Sometimes considered to contain just one family (Cryptoniscidae), but usually split up in different (host 

depending) families. 

Hemioniscidae 

@ Cirripedia 

* Not listed by Schram,1986 

Hemioniscus Buchholz,1866 

=? Cryptothir Dana,1852 

H. balani (Bate,1860) 

@ different balanid species 

* Should the author name be Buchholz,1866 (if (Bate,1860) is considered a nomen nudum)? 

Bohuslän, Kristiansand, North Sea, N Norway, Ireland, Scotland, Engl. Channel 

Leponiscus Giard,1887 

L. anatiferae Giard,1887 

@ Lepas anatifera 

Roscoff 

Crinoniscidae Bonnier,1900 

@ Balanidae 

* May perhaps be synonymized with Hemioniscidae, but the the single species within this family, C. 

equiptans is said to differ much from that other family. 

Crinoniscus Pérez,1900 

C. equiptans Pérez,1900 () 

@ Balanus perforatus ?& B. tintinnabulum 


Cyproniscidae 

@ Ostracoda 


Cyproniscus Kossmann,1884 

C. cypridinae (G.O. Sars,1882) 

@ Vargula norvegica & Philomedes brenda 

Bohuslän-Skagerrak-Lofoten-Norw. Sea 

Liriopsidae Bonnier,1900 

@ Rhizocephala 

Liriopsis Max Schultze,1859 

= Liriope Rathke,1843, non Meigen,1800, nec Lesson,1837 

L. pygmaea (Rathke,1843) 

@ Peltogaster paguri 

N Kattegatt-Bohuslän-W Norway, North Sea, White Sea, Black Sea 

Liriopsis sp. cf. L. monophthalma Fraisse,1878 / pygmaea Rathke 

@ rhizocephalans on Munida sp. 

* vide Brinkmann, A. jr.,1984. Sarsia 69 : 211-213 

W Norway 

Liriopsis sp. 

@ Galatheascus minor Boschma on Galathea nexa sp. 


W Norway 

Danalia Giard,1887 

D. larvaeformis (Giard,1887?) 

@ Sacculuna carcini /Carcinus maenas (Linnaeus) 

Roscoff 

Danalia sp. 

@ Sylon hippolytes from Pandalina ?profunda 


W Norway 

Asconiscidae 

@ Mysida 


Asconiscus G.O. Sars,1899 


@ Boreomysis arctica (Krøyer) 

Bohuslän-Oslofjord-Skagerrak, Trondheim, North Sea, Ireland 

Podasconidae 

@ Amphipods 


Parapodascon Hansen,1916 

P. stebbingi (Giard & Bonnier,1895) 

= Podascon Stebbingi Giard & Bonnier,1895 

@ Boeckosimus plautus (Krøyer) & B. normani (G.O. Sars) 

Bergen, North Sea 

Cabiropsidae 

@ Isopods 


Clypeoniscus Giard & Bonnier,1895 

C. hanseni Giard & Bonnier,1895 

@ Idotea spp. 

S Baltic-Bohuslän-W Norway, Scotland, Engl. Channel, France 

C. meinerti Giard & Bonnier,1895 

@ Pleuroprion murdochi & Synidothea nodulosa Krøyer 

?Faeroes or ?Iceland-Kara Sea-W Greenland 

65

Munnoniscus Giard & Bonnier,1895 

M. marsupialis (G.O. Sars,1882) () 

@ Eurycope cornuta G.O. Sars & Ilyarachna longicornis (G.O. Sars) 

Bohuslän-Skagerrak-Norway-North Sea 

Cironiscus Nielsen,1967 

C. dahli Nielsen,1967 

@ Natatolana borealis (Lilljeborg) & Politolana microphthalma (Hoek) 

W Baltic-Skagerrak-North Sea-Norw. Sea 

Ancyroniscus Caullery & Mesnil,1919 

A. bonnieri Caullery & Mesnil,1919 

@ Dynamene bidentata (Adams) 

Roscoff, S & W Britain, Ireland? 

Arcturocheres Hansen,1916 

A. pulchripes Hansen,1916 

@ Astacilla granulata (G.O. Sars) & Arcturus hystrix G.O. Sars 

off SE Iceland-off W Faeroes 

Astacilloechus Hansen,1916 

A. ingolfi Hansen,1916 

@ marsupium of Astacilla granulata (G.O. Sars) 

off SE Iceland 

Cabirops Kossman,1884 

= Cabira Kossman,1872, non Sodovsky,1837 (Lepidoptera), nec Gistl,1847 (Coelenterata) 

C. marsupialis (Caroli,1953) 

= Paracabirops marsupialis Caroli,1953 

@ hyperparasite on Gyge branchialis Cornalia & Panceri & Bopyrina ocellata (Czerniavsky) 

Gulf of Naples, S France 

C. ibizae Bourdon,1966 

@ hyperparasite on Megachelione foresti Bourdon,1968 

Sète, Baleares 

C. perezi Carayon,1942 

@ hyperparasite on Bopyrissa fraissei (Carayon) 

Arcachon 

C. codreanui Bourdon,1966 

@ hyperparasite on Bopyrissa diogeni (Popov) 

Engl. Channel 

C. sp. - very close to C. ibizae Bourdon : Bourdon,1968 

@ hyperparasite on Pleurocrypta microbranchiata G.O. Sars,1899 

Mediterranean 

"Cryponiscid No. 2" : G.O. Sars,1899 

@ Aega ventrosa 

?Norway 

Cryptoniscidae s.l. 

Cumoechus Hansen,1916 

C. insignis Hansen,1916 

@ Diastylis polaris G.O. Sars, D. echinata Bate & Hemilamprops cristata G.O. Sars 

Faeroes-Jan Mayen 

AMPHIPODA Latreille,1816 

CAPRELLIDEA Leach,1814 

CYAMIDA Bousfield,1979 

Cyamoidea White,1847 

Cyamidae White,1847 

Isocyamus Gervais & van Beneden,1859 

delphinii (Guérin-Méneville,1836) 

= Cyamus globicipitis Lütken,1873 

@ Phocaena phocaena & other species (e.g. Globicephala melaena, G. melas, Mesoplodon grayi, Pseudorca 

crassidens ) 

Bohuslän, Netherlands, Faeroes, Mediterranean, Australia 

I. kogiae Sedlak-Weinstein,1992 

@ Kogia breviceps (de Blainville,1838) 

Queensland 

Cyamus Latreille,1796 

= Paracyamus G.O. Sars,1895 

C. boopis Lütken,1873 

@ Megaptera novaeangliae (Borowski) "Humpback whale" 

Skagerrak-Arctic Seas & Iceland 

C. erraticus Roussel de Vauzème,1834 

@ Eubalaena glacialis Müller 

Shetland, Rockall-Iceland & Arctic Seas 

C. ovalis Roussel de Vauzème,1834 

@ Eubalaena glacialis Müller & Physeter catodon Linnaeus 

Shetlands-Faeroes-Iceland 

C. gracilis (Roussel de Vauzème,1834) 

@ Eubalaena glacialis Müller 

British Isles 

C. orcini Leung,1970 

@ Orcinus orca (Linnaeus) 

South Africa 

C. ceti (Linnaeus,1758) Lamarck,1801 

@ Balaena mysticetus Linnaeus & Eschrichtius gibbosus (Erxleben) 

Greenland, Scotland 

C. balaenoptera Barnard,1931 

@ Balaenoptera spp. 

South Africa 

C. monodontis Lütken,1873 

@ Monodon monoceros Linnaeus & Delphinapterus leucas (Pallas) 

Greenland 

C. nodosus Lütken,1860 

@ Monodon monoceros Linnaeus & Delphinapterus leucas (Pallas) 

Greenland 

C. catodontis Margolis,1954 

@ Physeter catodon Linnaeus 

Netherlands, S America 

C. bahamondei Buzeta,1963 

@ Physeter catodon Linnaeus 

Bahamas? 

C. antarcticensis Berzin & Vlasova,1982 

@ Orcinus orca (Linnaeus) 

Antarctic 

C. orubraedon Waller,1989 

@ Berardius bairdii 

Japan 

Platycyamus Lütken,1873 

P. thompsoni (Gosse,1855) 

@ Hyperoodon ampullatus Forster, Mesoplodon grayi 

Netherlands, England, Spitsbergen, Australia 

P. flaviscutatus Waller,1989 

@ Berardius bairdii 

Japan 

Scutocyamus Lincoln & Hurley,1974 

S. parvus Lincoln & Hurley,1974 

@ Lagenorhynchus albirostris Gray 

Netherlands 

S. antipodensis Lincoln & Hurley,1980 

@ Cephalorhynchus hectori 

New Zealand 

Syncyamus Bowman,1955 

S. pseudorcae Bowman,1955 

@ Pseudorca crassidens (Owen) 

The Caribbean, Australia 

S. aequus Lincoln & Hurley,1981 

@ Stenella coeruleoalba, S. longirostris, Delphinus delphis, Tursiops aduncus, T. truncatus, Mesoplodon 

grayi 

Mediterranean, South Africa, Australia 

S. sp. Leung,1970 

@ Delphinus delphis Linnaeus 

Gibraltar 

Neocyamus Margolis,1955 

N. physeteris (Pouchet,1888) 

@ Physeter catodon Linnaeus & Phocoenoides truei Andrews 

S America 

CAPRELLIDA Bousfield,1979 

Phtisicoidea Vassilenko,1968 

Phtisicidae Vassilenko,1968 

Phtisica Slabber,1769 

P. marina Slabber,1769 

Öresund-Bohuslän-W & N Norway-Br. Isles-Senegal-Azores-Canaries-Black Sea 

Protellidae McCain,1970 

* Correct superfamily? 

Protellina Stephensen,1944 

P. ingolfi Stephensen,1944 

Faeroes, Iceland, between Iceland & Norway 

Paracercopidae Vassilenko,1968 


Dodecadidae Vassilenko,1968 


Caprelloidea White,1847 

Caprellidae White,1847 

Caprella Lamarck,1801 

C. linearis (Linnaeus,1767) 

W Baltic-Bohuslän-Iceland & Arctic Seas 

C. septentrionalis Krøyer,1838 

Öresund-Bohuslän-Iceland & Arctic Seas 

C. monocera G.O. Sars,1895 

* this variety of C. septentrionalis should be treated as a separate species until more information about it have 

been gained, sensu Laubitz,1972 

Finnmark? 

C. equilibra Say,1818 

= Caprella aequilibra : Aucct. 

Kattegatt & Trondheimsfjord-"almost cosmopolitic" 

C. ciliata G.O. Sars,1883 

Öresund-Bohuslän-S Norway-off W Norway-Iceland 

C. microtuberculata G.O. Sars,1879 

N Norway-Arctic Seas-off SW & NW Faeroes 

C. acanthifera Leach,1814 

North Sea 

C. rinki Stephensen,1916 

off SW Faeroes-off W Iceland-SW Greenland 

C. penantis Leach,1814 

66

= Caprella acutifrons Latreille,1816 


C. tuberculata Bate & Westwood,1866 

Helgoland, E Britain, Engl. Channel, W France 

C. fretensis Stebbing,1878 

Engl. Channel, W Britain, Ireland 

C. erethizon Mayer, in Gadeau de Kerville,1901 

Engl. Channel 

C. macho Platvoet, De Bruyne & Gmelig-Meyling,1995 

Netherlands 

C. andreae Mayer,1??? 

Ireland 

Metacaprella Mayer,1903 

M. horrida (G.O. Sars,1885) 

round Faeroes-Arctic Seas 

Proaeginina Stephensen,1940 

P. norvegica (Stephensen,1931) 

N Norway-S Iceland 

Aeginina Norman,1905 

A. longicornis (Krøyer,1842-43) 

= A. echinata (Boeck,1861) 

Öresund, W Jutland, E Scotland, Bergen, Faeroes, Iceland, Arctic Seas 

Parvipalpa Stephensen,1944 

P. verrucosa Stephensen,1944 


Parvipalpus Mayer,1890 

P. capillaceus (Chevreux,1887) 

= Aegina capillacea Chevreux,1887 

Ireland (Valencia harbour), Concarneau 

Thorina Stephensen,1944 

T. spinosa Stephensen,1944 


Pariambidae Laubitz,1993 

Pariambus Stebbing,1888 

= Podalirius Krøyer,1844, non Latreille,1802, nec Swainson,1833 

P. typicus (Krøyer,1845) 

= Podalirius typicus Krøyer,1845 

* Associated with Asterias rubens & Crossaster papposus 

Öresund-Bohuslän-Norw. Sea, Br. Isles 

Pseudoprotella Mayer,1890 

P. phasma (Montagu,1806) 

Kattegatt-Bohuslän-North Sea, Br. Isles 

Aeginellidae Vassilenko,1968 

Aeginella Boeck,1861 

A. spinosa Boeck,1861 

Haugesund-W & N Norway-Arctic Seas-Iceland-off W & NW Faeroes 

Caprogammaridae Kudrjaschov & Vassilenko,1966 


HYPERIIDEA H. Milne-Edwards,1830 

PHYSOSOMATA Pirlot,1929 

Scinoidea Stebbing,1888 

Scinidae Stebbing,1888 

Scina Prestandrea,1833 

S. borealis (G.O. Sars,1882) 

= Tyro Clausii Bovallius,1885 

Koster Channel-Skagerrak-W & N Norway-Arctic Seas-off S Iceland-Faeroe Channel 

S. crassicornis (J.C. Fabricius,1775) 

= Tyro Sarsi Bovallius,1885 

off S Iceland-Davis Strait 

S. rattrayi Stebbing,1895 

off S Iceland-W Scotland 

S. tullbergi (Bovallius,1885) 

= Scina pacifica Stebbing,1895 

Iceland 

S. marginata (Bovallius,1885) 

= S. lepisma Chun,1889 


S. submarginata Tattersall,1906 


S. vosseleri Tattersall,1906 


S. spinosa Vosseler,1901 

= Scina uniceps Stebbing,1895 


S. oedicarpus Stebbing,1895 


Acanthoscina Vosseler,1900 

A. acanthodes (Stebbing,1895) 

off S Iceland-off W Ireland-N & S Atlantic 

Parascina Stebbing,1904 

* Is probably synonymous with another genus (& may belong to another family) as this genus is not 

mentioned in the revision of Bowman & Gruner,1973. 

P. fowleri Stebbing,1904 

off S Faeroes-off S Iceland-Davis Strait 

Mimonectidae Bovallius,1885 

M. loveni Bovallius,1885 

off SW Iceland-Sargasso Sea 

Mimonectes Bovallius,1885 

M. steenstrupii Bovallius,1885 


M. gaussi (Woltereck,1904) 


M. loveni Bovallius,1885 


Proscinidae Pirlot,1933 

* No species in the area? 

Archaeoscinidae Stebbing,1904 


Lanceoloidea Bovallius,1887 

Lanceolidae Bovallius,1887 

Lanceola Say,1818 ( L. pelagica Say,1818) 

L. sayana Bovallius,1885 

off S Iceland-southwards in the Atlantic 

L. murrayi Norman,1900 

Faeroe Channel-southwards in the Atlantic 

L. serrata Bovallius,1885 

off S Iceland-Davis Strait-N Atlantic 

L. pacifica Stebbing,1888 

off S Iceland-W Ireland-Gibraltar 

L. loveni Bovallius,1885 

= Lanceola aestiva Stebbing,1888 

off S Iceland-Davis Strait-SW Ireland-Bay of Biscay-N Atlantic 

L. borealis Bate & Westwood,1888 


Chuneolidae Woltereck,1909 


Microphasmidae Stephenson & Pirlot,1931 


PHYSOCEPHALATA Bowman & Gruner,1973 

Vibilioidea Dana,1952 

Vibiliidae Dana,1852 

Vibilia H. Milne-Edwards,1830 ( V. peronii H. Milne-Edwards,1830) 

V. borealis Bate & Westwood,1868 

= Vibilia Kröyeri Bovallius,1887 

Faeroes-W Greenland-Scotland-N Ireland-Gibraltar-W Mediterranean 

V. armata Bovallius,1887 

European Atlantic N to 50°, Mediterranean 

V. cultripes Vosseler,1901 


V. jeangerardii Lucas,1849 

@ Salps 

European Atlantic N to 43° (52°), Mediterranean 

V. propinqua Stebbing,1888 


V. pyripes Bovallius,1887 

European Atlantic N to 40° (50°) 

V. viatrix Bovallius,1887 

@ Salps 


V. gibbosa Bovallius,1887 


Cystisomatidae Willemoës-Suhm,1875 

Cystisoma Guérin-Méneville,1842 

C. fabricii Stebbing,1888 

European Atlantic N to ≈50° 

C. latipes (Stephensen,1925) 

= Thaumatops latipes Stephensen,1925 

European Atlantic N to ≈50° 

C. pellucida (Willemoës-Suhm,1874) 

= Cystisoma parkinsoni Stebbing,1888 

European Atlantic N to 50° & 58-59° N 

C. neptunus Guérin-Méneville,1842 () 


C. spinosum (J.C. Fabricius,1775) 


Paraphronimidae Bovallius,1887 

Paraphronima Claus,1879 

P. crassipes Claus,1879 


67

P. gracilis Claus,1879 () 


Platysceloidea Bate,1862 

Lycaeidae Claus,1879 


Tryphanidae 

Tryphana Boeck,1870 

= Tryphaena Bovallius,1887 

T. malmi Boeck,1870 () 

= Tryphaena Nordensiöldii Bovallius,1887 

= Tryphaena Boeckii Stebbing,1888 

Hardangerfjord-off W Norway-Iceland-Faeroes-Shetlands-Scotland 

Brachyscelidae 

Brachyscelus Bate,1861 

B. crusculum Bate,1861 () 

= Thamyris mediterranea Claus,1887 

off SW Iceland-off SW Faeroes-Atlantic Ocean & Mediterranean 

Anapronoidae Bowman & Grüner,1973 


Oxycephalidae Bate,1862 

Glossocephalus Bovallius,1887 

G. milneedwardsi Bovallius,1887 () 


Parascelidae Bovallius,1887 

Thyropus Dana,1852 ( T. diaphanus Dana,1852) 

= Parascelus Claus,1879 

T. typhoides (Claus,1887) 

= Parascelus typhoides Claus,1887 

European Atlantic in 40-42° N, Mediterranean 

Platyscelidae Bate,1862 

Platyscelus Bate,1861 

P. ovoides (Risso,1816) 

= Typhis ovoides Risso,1816 () 


P. serratulus Stebbing,1888 


Tetrathyrus Claus,1879 

T. forcipatus Claus,1879 () 


Pronoidae Claus,1879 

Euphronoe Claus,1879 

E. minuta Claus,1879 


Paralycaea Claus,1879 

P. gracilis Claus,1879 


Phronimoidea Dana,1853 

Phronimidae Dana,1853 

Phronima Latreille,1802 

P. sedentaria (Forskål,1775) 

= Cancer sedentarius Forskål,1775 () 

= Phronima atlantica Guérin-Méneville,1836 

* Associated with salps, pyrosomes & doliolids 

North Sea-off SW Faeroes-off SW Iceland, Mediterranean 

P. colletti Bovallius,1887 


P. stebbingii Vosseler,1901 


Hyperiidae Dana,1852 

Themisto Guérin,1828, non Oken,1807, nec Oken,1815 (nomina rejicenda) 

= Parathemisto Boeck,1871 

= Euthemisto Bovallius,1887 

T. abyssorum (Boeck,1871) 

= Parathemisto abyssorum Boeck,1870 

= Parathemisto oblivia G.O. Sars,1895 

Kattegatt-Bohuslän-W & N Norway-Arctic Seas-Jan Mayen-Iceland-Faeroes 

T. compressa Goës,1865 

= Parathemisto gracilipes (Norman,1869) 

= Parathemisto gaudichaudi : Auctt., non (Guérin,1825) 

= Themisto bispinosa Boeck,1871 

@ different hydromedusae; also feeding on other zooplankton, e.g. copepods & chaetognaths 

Bohuslän-North Sea-N Norway-Jan Mayen-Iceland-Faeroes-W Ireland 

T. libellula (Mandt,1822) 

N Norway-Arctic Seas, e.g. N Greenlandoff W, N & E Iceland-off N Faeroes 

Hyperoche Bovallius,1887 

H. medusarum (Krøyer,1838) 

= Metoecus medusarum Krøyer,1838 

= Hyperoche kröyeri Bovallius,1887 () 

* Associated with Pleurobrachia pileus (O.F. Müller) 

Öresund-Bohuslän-S, W & N Norway-Arctic Seas-Jan Mayen-Iceland-off W Faeroes 

Hyperia Latreille, in Desmarest,1823 

H. galba (Montagu,1813) 

= Cancer gammarus galba Montagu,1813 

@ Aurelia, (Cyanea etc.) 

Åland-Bohuslän-North Sea-W & N Norway-Iceland-Faeroes-Shetlands-Br. Isles, N Greenland 

H. medusarum (O.F. Müller,1776) 

= Hyperia Sueri Latreille, in Desmarest,1823 () 

= Cancer medusarum O.F. Müller,1776 

@ Cyanea, Aurelia etc. 

Öresund-Bohuslän-W & N Norway-Spitsbergen-Iceland-Faeroes-W Ireland 

H. spinigera Bovallius,1889 

@ ? 

* Possibly a synonyme of H. galba (Montagu) 

off N Norway-Spitsbergen, off S Iceland, off W Ireland & S England 

Hyperoides Chevreux,1900 

H. longipes Chevreux,1900 () 

W France, off Biscay, Azores, Mediterranean 

Lestrigonus H. Milne-Edwards,1830 ( L. fabrei H. Milne-Edwards,1830) 

L. latissimus (Bovallius,1887) 

= Hyperia latissima Bovallius,1887 

?European & SE Atlantic, Mediterranean 

Dairellidae Bovallius,1987 

Dairella Bovallius,1887 ( Paraphronima californica Bovallius,1885 - Pacific) 

D. latissima Bovallius,1887 


Phrosinidae Dana,1853 

Anchylomera H. Milne-Edwards,1830 

A. blossevillii H. Milne-Edwards,1830 () 


Phrosina Risso,1822 

P. semilunata Risso,1822 () 


Primno Guérin-Méneville,1836 ( P. macropa Guérin-Méneville,1836) 

P. evansi Sheader,1986 

= Primno macropus : Auctt., non Guérin-Méneville,1836 


Lycaeopsoidea Chevreux,1913 

Lycaeopsidae Chevreux,1913 

Lycaeopsis Claus,1879 

L. themistoides Claus,1879 () 

N Atlantic, Mediterranean, Antarctic 

INGOLFIELLIDEA Hansen,1903 

Ingolfiellidae Hansen,1903 

Ingolfiella Hansen,1903 

I. (Hansenliella Stock,1979) britannica Spooner,1960 

SW England at 45 m 

Metaingolfiellidae Ruffo,1969 

* No species within the area 

GAMMARIDEA Latreille,1803 

Lysianassoidea Dana,1849 

Lysianassidae Dana,1849 

Trischizostoma Boeck,1861 

T. raschii Boeck,1871 

=?? T. nicaeence (Costa,1853) 

@ skin of Etmopterus spinax 

Skagerrak-W & N Norway-off S Iceland-off SW Ireland 

Sophrosyne Stebbing,1888 

S. robertsoni Stebbing & Robertson,1891 

W Scotland 

Normanion Bonnier,1893 

N. sarsi Stebbing,1906 

= N. quadrimanus : G.O. Sars,1895, non (Bate & Westwood,1868) 

@ skin of fishes (both living & dead) 

S Norway-Trondheim 

N. quadrimanus (Bate & Westwood,1868) 

= N. amblyops G.O. Sars,1895 

@ skin of Etmopterus spinax & Melanogrammus aeglefinus 

Trondheimsfjord-W Scotland-N & W France 

Cheirimedon Stebbing,1888 

C. latimanus (G.O. Sars,1882) 

?Skagerrak-Stavanger-off SW Iceland 

Podoprionella G.O. Sars,1895 

P. norvegica G.O. Sars,1895 


Opisa Boeck,1876 

O. eschrichti (Krøyer,1842) 

= O. typica (Krøyer,1846) 

Skagerrak-N Norway-Arctic Seas-Iceland-Faeroes-Ireland 

Lepidepecreum Bate & Westwood,1868 

68

L. longicorne (Bate & Westwood,1861) 

= L. carinatum Bate & Westwood,1868 

Kattegatt-S Norway-North Sea-Br. Isles-Portugal-Mediterranean 

L. umbo (Goës,1866) 

NW & N Norway-Arctic Seas-Iceland-between Iceland & Faeroes 

L. serratum Stephensen,1925 

off SW Iceland-off SW Faeroes 

L. typhlops Bonnier,1896 

off SW Faeroes-Bay of Biscay 

Perrierella Chevreux & Bouvier,1892 

P. audouiniana (Bate,1857) 

Kattegatt-Bohuslän-North Sea-W Norway-Br. Isles-W Africa-Mediterranean 

Ambasia Boeck,1871 

A. atlantica (Milne-Edwards,1830) 

= A. danielsseni Boeck,1871 

Skagerrak-W & N Norway-Greenland-Iceland-Faeroes-off SW Ireland 

A. murmanica Brüggen,1905 

N Norway-Murmansk-E Greenland-off SW Faeroes 

Metambasia Stephensen,1923 

M. faeroensis Stephensen,1925 


Schisturella Norman,1900 

S. pulchra (Hansen,1887) 

off NW Scotland-W Greenland 

Aristias Boeck,1871 

A. neglectus Hansen,1887 

@ Bolocera tuediae, Macandrevia cranium, other Brachiopods & Sponges 

Kattegatt-Bohuslän-W & N Norway-Shetlands-Br. Isles-Mediterranean 

A. microps G.O. Sars,1895 

?Skagerrak-Trondheim-N Norway-off W & SW Iceland-offoff SW Faeroes 

A. megalops G.O. Sars,1895 

* Possibly the male of A. microps G.O. Sars 

Trondheim-N Norway 

A. tumidus (Krøyer,1846) 

N Norway-Jan Mayen-Arctic Seas 

A. falcatus Stephensen,1923 


Lysianella G.O. Sars,1882 

L. petalocera G.O. Sars,1882 

Koster Channel-S, W & N Norway 

Tryphosites G.O. Sars,1891 

T. longipes (Bate & Westwood,1861) 

Kattegatt-Bohuslän-W & N Norway-Faeroes-Rockall-Canaries-Mediterranean 

T. alleni Sexton,1911 

off SW Faeroes-off W Ireland-Bay of Biscay 

Scopelocheirus Bate,1856 

S. hopei (Costa,1851) 

= S. kroeyeri (Bruzelius,1859) 

= S. crenatus Bate,1857 

* Lincoln,1979 synonymized S. crenatus with S. hopei 

Kattegatt-Bohuslän-North Sea-W & N Norway-Iceland-Br. Isles-W France-Mediterranean 

Tmetonyx Stebbing,1906 

= Hoplonyx G.O. Sars,1891 (homonym) 

T. cicada (O. Fabricius,1780) 

Öresund-Bohuslän-W & N Norway-Arctic Seas-Iceland-off Shetlands-Br. Isles-Jersey 

T. similis (G.O. Sars,1895) 

= T. leucophthalmus (G.O. Sars,1895) 

Kattegatt-Skagerrak-W & N Norway-Arctic Seas-Iceland-Br. Isles-NW France-Mediterranean 

T. acutus (G.O. Sars,1895) 

Bohuslän-Skagerrak-N Norway-off SW Faeroes 

T. albidus (G.O. Sars,1895) 

N Norway 

T. rotundatus Chevreux,1926 

off Lofoten 

T. caeculus (G.O. Sars,1895) 

* Should be moved to a new genus, according to Oleröd,1987 

Bohuslän-W & N Norway-SW Iceland 

T. longichela Stephensen,1925 

off SE Iceland 

T. gracilipes Stephensen,1925 

off N Faeroes 

T. trionyx Stephensen,1925 


T. norbiensis Oleröd,1987 

abyssal depths in Norw. Sea 

Euonyx Norman,1867 

E. chelatus Norman,1867 () 

@ Echinus esculentus 

N Norway-off SW Faeroes-Shetlands-W Scotland-Wales 

Stephonyx Lowry & Stoddart,1989 

S. biscayensis (Chevreux,1908) 

= Euonyx biscayensis Chevreux,1908 () 

off SW Faeroes-off SW Ireland-Bay of Biscay-S Africa 

S. talismani (Chevreux,1919) 

= Euonyx talismani Chevreux,1919 

off SW Faeroes-off SW Ireland 

Kerguelenia Stebbing,1888 

K. borealis G.O. Sars,1895 

Hardangerfjord-N Norway 

Acidostoma Lilljeborg,1865 

A. nodiferum Stephensen,1923 

= Acidostoma obesum Auctores, non (Bate & Westwood,1861) 

= Acidostoma sarsi Lincoln,1979 

@ A. nodiferum occurs together with Actinostola callosa 

Öresund-Bohuslän-North Sea-W & N Norway-Iceland-?off SW Faeroes 

A. laticorne G.O. Sars,1879 

Arctic Seas, ?Roscoff 

A. obesum (Bate & Westwood,1861) 

= Acidostoma laticorne : Oldevig,1933, non G.O. Sars,1879 

= Acidostoma neglectum Dahl,1964 

@ Peachia hastata & Cereactis aurantiaca 

Kattegatt-North Sea-Br. Isles-NW France 

Socarnes Boeck,1871 

S. bidenticulatus (Bate,1858) 

N Norway-Arctic Seas-Jan Mayen-?Iceland 

S. vahli (Krøyer,1838) 

Haugesund-N Norway-Arctic Seas-NE Iceland 

S. erythrophthalmus Robertson,1892 

W Br. Isles 

Socarnopsis Chevreux,1911 

S. filicornis (Heller,1867) 

= Socarnopsis crenulatus Chevreux,1911 

Shetland, W Br. Isles 

Nannonyx G.O. Sars,1890 

N. goesii (Boeck,1871) 

North Sea 

N. spinimanus Walker,1895 

NE France & Br. Isles 

Menigrates Boeck,1871 

M. obtusifrons (Boeck,1861) 

Skagerrak-North Sea-W & N Norway-Arctic Seas-Iceland-Br. Isles-Guernsey 

Menigratopsis Dahl,1945 

M. svennilssoni Dahl,1945 

Öresund-North Sea 

Tetronychia Stephensen,1923 

T. abyssalis Stephensen,1923 

off S Iceland 

Koroga Holmes,1909 

K. megalops Holmes,1909 

?off S Iceland-off S Alaska 

Centromedon G.O. Sars,1895 

C. pumilus (Lilljeborg,1865) 

Bohuslän-Skagerrak-W & N Norway-Arctic Seas & Iceland 

C. typhlops (G.O. Sars,1885) 

off NE Iceland-between Jan Mayen & Finnmark 

C. calcaratus (G.O. Sars,1885) 

between Iceland & Jan Mayen-Arctic Seas 

Ichnopus Costa,1853 

I. spinicornis Boeck,1861 

Bohuslän-Trondheim-North Sea-Br. Isles-Mediterranean 

Lysianassa Milne-Edwards,1830 

L. plumosa Boeck,1871 

Haugesund-Trondheim-Shetlands-Br. Isles-NW France-Senegal-Mediterranean 

L. ceratina (Walker,1889) 

W Br. Isles 

L. insperata Lincoln,1979 

Guernsey 

Hippomedon Boeck,1871 

H. holboelli (Krøyer,1846) 

Skagerrak, ?N Norway-Arctic Seas, e.g. N Greenland & Iceland-off N Faeroes 

H. denticulatus (Bate,1857) 

Öresund-Bohuslän-W & N Norway-Kolafjord-Shetlands-Br. Isles-Mediterranean 

H. propinquus G.O. Sars,1895 

?Bohuslän-Skagerrak-W & N Norway-Arctic Seas-Iceland 

H. robustus G.O. Sars,1895 

Skagerrak-W & N Norway-Iceland-Azores 

H. striolatus Stephensen,1923 


H. reticulatus Stephensen,1923 


H. nasutus Stephensen,1923 


69

H. frigidus Stephensen,1923 

off NE Iceland 

H. serratipes Stephensen,1923 

off S Iceland 

Thoriella Stephensen,1915 

T. islandica Stephensen,1915 

off S Iceland 

Crybelocephalus Tattersall,1906 

C. megalurus Tattersall,1906 

off S Iceland-off W Ireland 

Cyclocaris Stebbing,1888 

C. guilelmi Chevreux,1899 

= Cyclocaris faroensis Norman,1900 

off Lofoten-Arctic Seas-off E Jan Mayen-off N & S Faeroes 

Cyphocaris Lütken, in Boeck,1871 

C. anonyx Boeck,1871 

= Cyphocarix micronyx Chevreux,1900 

off Lofoten-off W & S Iceland-off NW & SW Faeroes-off N & W Ireland 

C. bouvieri Chevreux,1916 

off N Faeroes-N Atlantic 

C. richardii Chevreux,1905 


Metacyphocaris Tattersall,1906 

M. helgae Tattersall,1906 

off S Iceland-off SW Ireland 

Paracyphocaris Chevreux,1905 

P. praedator Chevreux,1905 

off S Iceland-Bay of Biscay-Azores 

Amaryllis Haswell,1879 

A. pulchella Bonnier,1896 

off S Iceland-off SW Faeroes-Bay of Biscay 

Onisimus Boeck,1871 ( Anonyx litoralis Krøyer,1845) 

= Onesimus Boeck,1876 (invalid emendation of Onisimus ) 

= Pseudalibrotus Della Valle,1893 ( Anonyx litoralis Krøyer,1845) 

= Paronesimus Stebbing,1894 ( Paronesimus barentsi Stebbing,1894) 

= Boeckosimus Barnard,1969 ( Anonyx edwardsii Krøyer,1846) 

* Lowry & Stoddart,1992 synonymized the genera above 

O. affinis Hansen,1887 

Jan Mayen-Arctic Seas 

O. barentsi (Stebbing,1894) 

off W Novaja Zemblja, circum-Arctic 

O.brevicaudatus Hansen,1887 

N Greenland 

O. edwardsii (Krøyer,1846) 

= Anonyx edwardsii Krøyer,1846 

Kattegatt-off W Norway-N Norway-Arctic Seas-Jan Mayen-Iceland 

O. glacialis (G.O. Sars,1900) 

= Pseudalibrotus glacialis G.O. Sars,1900 

Arctic Seas, e.g. N Greenland-Jan Mayen-Iceland-Orkneys 

O. leucopis G.O. Sars,1879 

between Norway & Iceland-?between Iceland & Jan Mayen 

O. litoralis (Krøyer,1845) 

= Anonyx litoralis Krøyer,1845 () 

N Norway-Arctic Seas, including NE Greenland & Iceland 

O. nanseni (G.O. Sars,1900) 

Arctic Seas-Iceland 

O. normani G.O. Sars,1891 

@ Bolocera tuediae 

* Should possibly be placed in a new genus, according to Lowry & Stoddart,1992. 

Oslofjord-Skagerrak-between S Norway & Shetlands-W & N Norway-Arctic Seas 

O. plautus (Krøyer,1845) 

= Anonyx plautus Krøyer,1845 

Bohuslän-Haugesund-Arctic Seas-Jan Mayen-Iceland 

O. sextoni Chevreux,1926 

off Lofoten Islands 

Anonyx Krøyer,1838 

= Chironesimus G.O. Sars,1895 

= Lakota Holmes,1908 

A. nugax (Phipps,1774) 

= Lakota chelata Chevreux,1926 

off Lofoten-N Norway-Faeroes-Arctic Seas 

A. lilljeborgi Boeck,1871 

Norwegian Sea 

A. sarsi Steele & Brunel,1968 

= A. nugax Sars,1891, non (Phipps,1774) 

Bohuslän? 

A. debruyni Hoek,1882 

N Norway-Arctic Seas 

A. pacificus Gurjanova,1962 

= A. nugax pacificus Gurjanova,1962 

N Greenland 

Tryphosella Bonnier,1893 

= Tryphosa : G.O. Sars,1891, non Boeck,1871 

T. sarsi Bonnier,1893 

= T. nana (G.O. Sars,1891), non Boeck,1861 

Kattegatt-Bohuslän-N Norway-Shetlands-Plymouth 

T. hoeringii (Boeck,1871) 

Bohuslän-North Sea-W & N Norway-off SW Faeroes-Br. Isles 

T. nanoides (Lilljeborg,1865) 

Bohuslän-W & N Norway-Arctic Seas-Iceland-Shetlands 

T. angulata (G.O. Sars,1895) 

Trondheim-N Norway-Iceland 

T. compressa (G.O. Sars,1895) 

Iceland-Arctic Seas 

T. spitzbergensis (Chevreux,1926) 

?N Norway-Spitsbergen 

T. schneideri (Stephensen,1921) 

N Norway-Iceland-Arctic Seas 

T. triangula (Stephensen,1925) 

Iceland, Arctic Seas 

T. rotundata (Stephensen,1925) 

off S Iceland-off SW Faeroes 

T. insignoides (Stephensen,1925) 


T. abyssalis (Stephensen,1925) 

off S Jan Mayen (deep water)-abyssal depths of Norw. Sea 

T. abyssi Norman,1900 


Orchomene Boeck,1871 

= Tryphosa Boeck,1871 

= Orchomenella G.O. Sars,1891 

= Orchomenopsis G.O. Sars,1895 

O. serrata (Boeck,1861) 

Bohuslän-W & N Norway-Arctic Seas-Iceland-Faeroes 

O. humilis (Costa,1853) 

= O. batei G.O. Sars,1882 

= O. hanseni Meinert,1890 

Kattegatt-Bohuslän-Norwegian Sea 

O. pectinata G.O. Sars,1882 

Bergen-N Norway-Arctic Seas-Iceland 

O. crispata (Goës,1866) 

Bohuslän-N Norway-Spitsbergen-?off S Iceland 

O. amblyops G.O. Sars,1895 

Koster area-Skagerrak-W & N Norway-off NE Iceland-off N & SW Faeroes 

O. thorii Stephensen,1923 


O. laevipes Stephensen,1923 

off W Iceland 

O. faeroensis Stephensen,1923 


O. obtusa (G.O. Sars,1895) 

= Orchomenopsis o. G.O. Sars,1895 

Bohuslän-Skagerrak-SW & N Norway-?off SW Iceland 

O. minuta (Krøyer,1846) 

= Orchomenella m. : G.O. Sars,1895 

Öresund-Bohuslän-W & N Norway-Arctic Seas-Iceland 

O. nana (Krøyer,1846) 

= Tryphosa ciliata G.O. Sars,1882 

= Orchomenella n. : Auctt. 

Öresund-Bohuslän-S & N Norway-North Sea-?Murmansk-Br.Isles-Spain-Senegal 

O. pinguis Boeck,1871 

= Orchomenella p. : G.O. Sars,1895 

Skagerrak-Helgoland-W & N Norway-Arctic Seas-Mediterranean 

O. groenlandica (Hansen,1887) 

= Orchomenella g. : G.O. Sars,1895 

N Norway-Arctic Seas 

O. crenata (Chevreux & Fages,1925) 

Roscoff 

O. similis Chevreux,1912 

W France 

O. abyssorum Stebbing,1888 


Eurytenes Lilljeborg,1865 

= Eurythenes Barnard,1932 

= Katius Chevreux,1905 

E. gryllus (Mandt,1822) 

= Katius obesus Chevreux,1905 

N Norway-Arctic Sea-off S Iceland-between Iceland & Greenland 

Parachevreuxiella Andres,1987 

P. lobata Andres,1987 


Uristidae Hurley,1963 

Uristes Dana,1849 

U. umbonatus (G.O. Sars,1882) 

W Skagerrak-W & N Norway-Iceland-off SW Faeroes 

Leucothoidea Dana,1852 

Pleustidae Buchholz,1874 

= Paramphithoidae 

70

Pleustes Bate,1858 

P. panopla (Krøyer,1838) 

Bergen-N Norway-Arctic Seas 

Pleusymtes Barnard,1969 

P. glaber (Boeck,1861) 

= Amphitopsis glaber Boeck,1861 

= Parapleustes glaber : G.O. Sars,1893 

= Sympleustes glaber : Auctt. 

Kattegatt-Bohuslän-S & N Norway-Arctic Seas-Iceland-Br. Isles-Bay of Biscay 

Parapleustes Buchholz,1874 

P. gracilis Buchholz,1874 

N Norway-Spitsbergen-Arctic Seas-Iceland-E Greenland 

P. assimilis (G.O. Sars,1882) 

= Paramphithoe assimilis G.O. Sars,1882 

Bohuslän-Helgoland-W & N Norway-Iceland-Greenland-Faeroes-N Br. Isles 

P. pulchellus (Krøyer,1846) 

Bohuslän-N Norway-Arctic Seas & Iceland 

P. monocuspis G.O. Sars,1895 

* According to Lincoln,1979, is this a synonym of P. bicuspis (Krøyer) 

Bohuslän-Skagerrak-N Norway-Greenland-Iceland-Faeroes 

P. bicuspis (Krøyer,1838) 

= Amphithoe bicuspis Krøyer,1838 

Katteg.-Bohuslän-N Norway-Arctic Seas-Iceland-Br. Isles-NW France 

Stenopleustes G.O. Sars,1893 

= Sympleustes Stebbing,1899 

S. malmgreni (Boeck,1871) 

* Often on Paragorgia rosea 

Koster Channel-Skagerrak-N Norway-Iceland & Arctic Seas 

S. nodifer (G.O. Sars,1882) 

= Amphitopsis nodifera G.O. Sars,1882 

Bohuslän-S Norway-Trondheim-Faeroes-Br. Isles-NW France 

S. latipes (M. Sars,1858) 

= Amphithoe latipes M. Sars,1858 

@ dead ?Paramuricea 

Bohuslän-North Sea-N Norway-Iceland & Arctic Seas-Br. Isles-Bay of Biscay 

S. eldingi Gurjanova,1930 

Tromsö area-Barents Sea 

S. pulchellus (G.O. Sars,1876) 

N Norway-off E Iceland-Arctic Seas 

S. megacheir (Walker,1897) 

=? S. grandimanus (Chevreux,1887) 

off W Iceland-slope of Engl. Channel 

Colomastigidae Stebbing,1899 

Colomastix Grube,1861 

= Cratippus Bate,1862 

= Exunguia Norman,1869 

C. pusilla Grube,1861 

= Cratippus tenuipes Bate,1862 

= Exunguia stilipes Norman,1869 

* usually associated with Sponges 

Atlantic coast of Europe-Mediterranean 

Amphilochidae Boeck,1872 

Stegoplax G.O. Sars,1882 

S. longirostris G.O. Sars,1882 

Hardangerfjord-off Lofoten-off SW Iceland 

Amphilochus Bate,1862 

A. tenuimanus Boeck,1871 

W Skagerrak-W & N Norway-off Iceland-Br. Isles 

A. manudens Bate,1862 

= A. boecki Meinert,1??? 

Öresund-Bohuslän-W & N Norway-Murmansk-Iceland-Faeroes-Bay of Biscay 

A. borealis Enequist,1949 

Skagerrak 

A. spencebatei (Stebbing,1876) 

= Probolium Spence-Batei Stebbing,1876 

=? A. brunneus Della Valle,1893 

= A. anomalus Chevreux,1900 

Irish Sea-Mediterranean 

A. neapolitanus Della Valle,1893 

= A. melanops Walker,1894 

Liverpool Bay-Mediterranean 

Amphilochopsis Stephensen,1925 

A. hamatus Stephensen,1925 

off S Jan Mayen-off W Greenland-?off W, SW & S Iceland-?off N Faeroes 

Amphilochoides G.O. Sars,1892 

A. boeckii G.O. Sars,1892 

= A. odontonyx : G.O. Sars,1892, non (Boeck,1871) 

Bohuslän-W Norway & North Sea 

A. serratipes (Norman,1869) 

= Probolium serratipes Norman,1869 

Öresund-Bohuslän-Skagerrak-Trondheim-Shetlands-W Scotland 

Paramphilichoides Lincoln,1979 

P. odontonyx (Boeck,1871) 

= Amphilochus odontonyx Boeck,1871 

= Amphilochoides pusillus G.O. Sars,1892 

= Amphilochoides odontonyx : G.O. Sars,1895 

?Öresund-?Bohuslän-Skagerrak-W & N Norway-Br. Isles 

P. intermedius (Scott,1896) 

= Amphilochoides intermedius Scott,1896 

Gullmarn, Firth of Forth 

Gitanopsis G.O. Sars,1895 

G. bispinosa (Boeck,1871) 

= Amphilochus bispinosus Boeck,1871 

Oslofjord-?Skagerrak-W & N Norway-Clyde Sea 

G. inermis (G.O. Sars,1882) 

= Amphilochus inermis G.O. Sars,1882 

N Norway-Arctic Seas-Jan Mayen-E Iceland-Northumberland 

G. arctica G.O. Sars,1895 

N Norway-Arctic Seas-E Iceland-S Greenland 

Gitana Boeck,1871 

G. rostrata Boeck,1871 

Skagerrak-W & N Norway 

G. abyssicola G.O. Sars,1895 

?Koster area-W & N Norway-?Scotland 

G. sarsi Boeck,1871 

= Amphilochus sabrinae Stebbing,1878 

Öresund-Bohuslän-W & N Norway-Spitsbergen-Br. Isles-Mediterranean 

Cyproideidae 

Peltacoxa Catta,1875 

P. damnoniensis (Stebbing,1885) 

= Cyproidea damnoniensis Stebbing,1885 

W British Isles 

P. brevirostris (Scott & Scott,1893) 

= Cyproidea brevirostris Scott & Scott,1893 

W Br. Isles & Bay of Biscay 

Leucothoidae Dana,1852 

Leucothoe Leach,1814 

L. lilljeborgii Boeck,1861 

= Leucothoe imparicornis Norman,1889 

Öresund-Bohuslän-North Sea 

L. spinicarpa (Abildgaard,1789) 

= Gammarus spinicarpus Abildgaard,1789 

= Leucothoe articulosa Leach,1814 

Öresund-Bohuslän-W & N Norway-Iceland-Br. Isles-N France 

L. procera Bate,1857 

= L. richiardii : Lincoln,1979, non Lessona,1865 

Engl. Channel, Scotland, Ireland 

L. incisa Robertson,1892 

Scotch North Sea coast-Engl. Cannel-Mediterranean 

Laphystiopsidae Stebbing,1899 

Laphystiopsis G.O. Sars,1893 

L. planifrons G.O. Sars,1893 

@ various fish species (half parasite) 

Bohuslän-N Norway-Iceland-off Faeroes 

Cressidae Stebbing,1899 

Cressa Boeck,1871 

= Danaia Bate,1857 (Homonym according to Lincoln,1979, but Neave,1939 does not mention an earlier 

name) 

C. dubia (Bate,1857) 

= Danaia dubia Bate,1857 

= Cressa shoidtei Boeck,1871 

Öresund-Bohuslän-Trondheim-Br. Isles & W Mediterranean 

C. minuta Boeck,1871 

Bohuslän-Skagerrak-W Norway-Iceland-E Scotland 

C. abyssicola (G.O. Sars,1885) 

between Norway & Bear Island-off N Iceland-W Greenland 

C. carinata Stephensen,1931 

Faeroes-Iceland-off S Jan Mayen-W Greenland 

C. quinquedentata Stephensen,1931 


Cressina Stephensen,1931 

C. monocuspis Stephensen,1931 

off SW Faeroes-off S Iceland-off W Greenland 

Stenothoidae Boeck,1871 

Metopella G.O. Sars,1892 

M. longimana (Boeck,1871) 

Oslofjord-Skagerrak-Haugesund-Iceland-Greenland 

M. neglecta (Hansen,1887) 

Haugesund-Greenland 

Hardametopa Barnard & Karaman,1991 

H. nasuta (Boeck,1871) 

= Metopa nasuta Boeck,1871 

= Metopella nasuta : Auctt. 

Bohuslän-N Norway-N Br. Isles-Iceland-W Greenland 

Proboloides Della Valle,1893 

P. gregarius (G.O. Sars,1882) 

Bohuslän-W & N Norway-Normandie-Bay of Biscay 

P. calcaratus (G.O. Sars,1882) 

Skagerrak-North Sea-N Norway 

71

P. glacialis (Krøyer,1842) 

Iceland-Spitsbergen-Greenland 

Parametopa Chevreux,1901 

P. kervillei Chevreux,1901 

= Metopa sarniensis Norman,1907 

Engl. Channel 

Metopa Boeck,1871 

M. quadrangulata Reibish,1905-1906 

Skagerrak 

M. robusta G.O. Sars,1895 

Skagerrak-N Norway-Greenland-off Faeroes-Scotland 

M. propinqua G.O. Sars,1892 

Bohuslän-North Sea-W & N Norway-Barents Sea-Scotland 

M. alderi (Bate,1857) 

= Montagua alderi Bate,1857 

= Metopa spectabilis G.O. Sars,1879 

@ may be found on Tima bairdii 

Kattegatt-Bohuslän-North Sea-W & N Norway-Iceland-N Br. Isles 

M. affinis Boeck,1871 

Oslofj.-W & N Norway 

M. norvegica (Liljeborg,1851) 

= Leucothoe norvegica Liljeborg,1851 

= Montagua pollexiana Bate,1857 

Bohuslän-North Sea-W & N Norway-Iceland-Shetlands-Br. Isles 

M. boecki G.O. Sars,1895 

Skagerrak-Haugesund-N Norway-Iceland 

M. longicornis Boeck,1871 

Oslofjord-Skagerrak-Lofoten-W Greenland 

M. pusilla G.O. Sars,1892 

Belt Sea-Bohuslän-Trondheim-Arctic Seas-Wales 

M. borealis G.O. Sars,1882 

@ may be found on Phialidium sp. 

Skagerrak-N Norway-Br. Isles-Engl. Channel 

M. soelsbergi Schneider,1884 

@ Actinoloba (= Metridium ) 

Öresund-Kattegatt-Hardangerfjord-Tromsö-Iceland-Clyde Sea 

M. bruzelii (Goës,1866) 

Kattegatt-Bohuslän-North Sea-W & N Norway-Spitsbergen-Iceland-Faeroes-Br. Isles 

M. tenuimana G.O. Sars,1892 

W Norway-Shetlands 

M. leptocarpa G.O. Sars,1895 

Christanssund-off SW Iceland 

M. sinuata G.O. Sars,1895 

N Norway-Spitsbergen-Iceland-off W Faeroes-between Norway & Scotland-Davis Strait 

M. invalida G.O. Sars,1895 

N Norway-Iceland 

M. latimana Hansen,1887 

Scotland-Greenland 

M. palmata G.O. Sars,1895 

Iceland, Hammerfest 

M. glacialis (Krøyer,1842) 

= Proboloides glacialis (Krøyer,1842) 


M. abyssalis Stephensen,1931 

off N Faeroes-off NE Iceland 

Stenula Barnard,1962 

S. rubrovittata (G.O. Sars,1882) 

= Metopa r. & Stenothoides latipes Chevreux & Fage,1925 

@ typically associated with Pagurus bernhardus (Linnaeus) 

Kattegatt-Bohuslän-North Sea-W & N Norway-Faeroes-Shetlands-Br. Isles-Bay of Biscay 

Stenothoe Dana,1852 

S. monoculoides (Montagu,1815) 

Kattegatt-Bohuslän-N Norway-Iceland-Faeroes-Azores-Senegal-Black Sea 

S. tenella G.O. Sars,1882 

Bergen-Trondheim 

S. megacheir (Boeck,1871) 


S. microps G.O. Sars,1895 

Skagerrak-North Sea 

S. marina (Bate,1856) 


Bohuslän-North Sea-N Norway-?Faeroes-W Africa-Mediterranean 

S. brevicornis G.O. Sars,1882 

S-N Norway 

S.? macrophthalma Stephensen,1931 


S.? aequicornis Stephensen,1931 

between Faeroes & Iceland 

S. valida Dana,1852 

Engl. Channel-Mediterranean 

S. tergestina (Nebeski,1881) 

= S. spinimana Chevreux,1911 


S. crassicornis Walker,1896 

Isle of Man 

S. setosa Norman,1900 

Plymouth 

S. gallensis Walker,1904 

= S. cattai Stebbing,1906 


S. elachistoides Myers & McGrath,1980 

Ireland 

Anamixidae Stebbing,1897 


Maxillipiidae Ledoyer,1973 


Pagetinidae K.H. Barnard,1932 


Nihotungidae Barnard,1972 


Thaumatelsonidae Gurjanova,1038 


Phoxocephaloidea G.O. Sars,1891 

Urothoidae Bousfield,1969 

Urothoe Dana,1852 

= Egidia Costa,1853 

U. elegans (Bate,1856) 

= Gammarus elegans Bate,1856 

= U. norvegica Boeck,1861 

Kattegatt-Skagerrak-W & N Norway-Kolafjord-Iceland-Faeroes-Engl. Channel-Senegal-Mediterranean 

U. poseidonis Reibish,1905 

= U. grimaldii var. poseidonis : Schellenberg,1942 

Kattegatt-North Sea 

U. brevicornis Bate,1862 

Engl. Channel, North Sea 

U. marina (Bate,1857) 

= Sulcator marinus Bate,1857 

* Associated with Synaptidae and Echinocardium cordatum 

Shetlands-Bay of Biscay 

U. pulchella (Costa,1853) 

= Egidia pulchella Costa,1853 

E Scotland-Engl. Channel-Mediterranean 

Phoxocephalidae G.O. Sars,1891 

Paraphoxus G.O. Sars,1891 

P. oculatus (G.O. Sars,1879) 

Skagerrak-W & N Norway-Jan Mayen-Iceland-Shetlands-Isle of Man 

P. maculatus Chevreux,1900 


Leptophoxus G.O. Sars,1895 

L. falcatus (G.O. Sars,1882) 

Koster Channel-W & N Norway-W Greenland 

Phoxocephalus Stebbing,1888 

= Phoxus Krøyer,1842, non Billberg,1820 (Coleoptera) 

P. holboelli (Krøyer,1842) 

= Phoxus holbölli Krøyer,1842 

S Baltic-Bohuslän-N Norway-White Sea-Iceland-W Greenland-Br. Isles 

P. tenuipes Stephensen,1925 

off W & SW Iceland-off W Greenland 

Metaphoxus Bonnier,1896 

M. pectinatus (Walker,1896) 

= Phoxocephalus pectinatus Walker,1896 

Ireland, Engl. Channel-Mediterranean 

M. simplex (Bate,1857) 

= Phoxus simplex Bate,1857 

* Lincoln,1979 does not comment on his synonomy of this species with M. pectinatus (Walker), but Bate's 

name is the oldest and should be used if they are considered as synonyms. 

Britain 

Parametaphoxus Gurjanova,1977 

P. fultoni (T. Scott,1890) 

= Phoxocephalus fultoni T. Scott,1890 

= Metaphoxus fultoni : Auctt. 

= Phoxocephalus chelatus Della Valle,1893 

Ireland, Engl. Channel-Mediterranean 

Harpiniopsis Stephensen,1925 

H. similis Stephensen,1925 

off N & NE Iceland-off N Faeroes-off S Jan Mayen-off W Greenland 

Harpinia Boeck,1871 

H. mucronata G.O. Sars,1879 

North Sea-N Norway-Iceland-off N Faeroes-E Greenland 

H. crenulata (Boeck,1871) 

Öresund-Bohuslän-W & N Norway-Iceland-Br. Isles-Bay of Biscay-W Mediterranean 

H. pectinata G.O. Sars,1891 

Bohuslän-North Sea-W & N Norway-Br. Isles-W Africa-W Mediterranean 

H. serrata G.O. Sars,1879 

N Norway-Arctic Seas, ?Northumberland 

H. antennaria Meinert,1890 

= H. neglecta G.O. Sars,1895 

Öresund-Bohuslän-W & N Norway-Barents Sea-Iceland-Br. Isles-Bay of Biscay 

72

H. propinqua G.O. Sars,1895 

Kattegatt, Lofoten-Jan Mayen-off N Spitsbergen-Iceland 

H. plumosa (Krøyer,1842) 

Arctic Seas & Iceland 

H. truncata G.O. Sars,1895 

Koster Channel-W & N Norway-N Iceland 

H. abyssi G.O. Sars,1879 

off W Norway-off N Norway-Arctic Seas-off E Iceland-off Faeroes 

H. laevis G.O. Sars,1891 

Kattegatt-Bohuslän-W Norway-Isle of Man-off SW Iceland 

H. amundseni Gurjanova,1946 

N Greenland 

Platyischnopidae Barnard & Drummond,1979 


Stegocephaloidea Dana,1952 

Stegocephalidae Dana,1852 

Andaniexis Stebbing,1906 

= Andania Boeck,1871 (homonym of a Lepidoptera) 

A. abyssi (Boeck,1871) Stebbing,1906 

= Andania abyssi Boeck,1871 () 

Bohuslän-Oslofjord-Skagerrak-W & N Norway-Arctic Seas-Rockall-off W Africa 

A. lupus Berge & Vader,1997 

N Norway, Faeroe area, Iceland, Arctic 

A. eilae Berge & Vader,1997 

Faeroe area, Iceland 

Stegocephalus Krøyer,1842 

S. inflatus Krøyer,1842 () 

Bohuslän-W & N Norway-Arctic Seas, e.g. N Greenland-S Iceland-Faeroe Channel 

Phippsiella Schellenberg,1925 

P. similis (G.O. Sars,1891) 

= Stegocephalus similis G.O. Sars,1891 

Skagerrak-W & N Norway-Faeroes-Spitsbergen-Arctic Seas-off NE Iceland-Davis Strait 

P. bioice Berge & Vader,1997 

off Iceland 

P. minima Stephensen,1925 

W Greenland-?off N Faeroes 

Stegocephaloides G.O. Sars,1891 

S. christianiensis (Boeck,1871) G.O. Sars,1891 

= Stegocephalus christianiensis () 

Bohuslän-North Sea-W Norway-Trondheimsfjord-Br. Isles-W Mediterranean 

S. auratus (G.O. Sars,1883) G.O. Sars,1891 

Bohuslän-Skagerrak-W Norway-Trondheimsfjord-Iceland-off S Faeroes, Biscay, ?Kara Sea 

S. barnardi Berge & Vader,1997 

off SW Iceland in 1042-1407 m, E Mediterranean 

S. wagini (Gurjanova,1936) Barnard & Karaman,1991 

= Stegocephalopsis wagini Gurjanova,1936 

Faeroe area, Iceland, Kara Sea 

Phippsia Stebbing,1906 

= Aspidopleurus G.O. Sars,1891 (homonym of a fish) 

P. gibbosa (G.O. Sars,1883) Stebbing,1906 

= Stegocephalus gibbosus G.O. Sars,1883 () 

W & N Norway, off Iceland 

P. roemeri Schellenberg,1925 

Faeroe area, Iceland, E Greenland, off N Spitsbergen 

Stegocephalina Stephensen,1925 

S. ingolfi Stephensen,1925 () 


?S. biofar Berge & Vader,1997 

off SE Faeroes-off SW Iceland and off E Greenland in 417-1525 m 

?S. idae Berge & Vader,1997 

off Iceland in 501-1042 m 

Stegocephalopsis Schellenberg,1925 

S. ampulla (Phipps,1774) 

= Cancer ampulla Phipps,1774 () 

off Iceland, Arctic circumpolar 

Andaniella G.O. Sars,1895 

A. pectinata (G.O. Sars,1882) 

= Andania pectinata G.O. Sars,1883 () 

Trondheimsfjord-N Norway-Spitsbergen-Arctic Seas-Faeroes-Iceland-off S Jan Mayen-W Greenland- 

Newfoundland 

Andaniopsis G.O. Sars,1895 

A. nordlandica (Boeck,1871) G.O. Sars,1895 

= Andania nordlandica Boeck,1871 () 

Bohuslän-Oslofjord-Skagerrak-W & N Norway 

Metandania Stephensen,1925 

M. islandica Stephensen,1925 

off W Iceland 

Euandania Stebbing,1899 

E. gigantea (Stebbing,1883) Stebbing,1899 

= Andania gigantea Stebbing,1883 () 

* Pelagic species 

Faeroe area, off Rockall, "cosmopolitian" 

Iphimediidae Boeck,1871 

= Acanthonotozomatidae Stebbing,1906 

= Paramphithoidae Stebbing,1906 

Clenardopsis Barnard,1916 

C. carinata Barnard,1916 

off SW Faeroes-S Africa 

Epimeria Costa,1851 

E. parasitica (M. Sars,1859) 

@ Parastichopus tremulus & Porania pulvillus (semiparasitic feeder of outer tissues) 

Oslofjord-Skagerrak-W & N Norway & North Sea-Bay of Biscay 

E. loricata G.O. Sars,1879 

off W Norway & Shetlands-N Norway-Arctic Seas & Iceland 

E. cornigera (J.C. Fabricius,1779) 

= Acanthonotus testudo Bate,1862 

= Acanthonotus owenii Bate & Westwood,1863 

Bohuslän-North Sea-N Norway-Br. Isles-Iceland-Mediterranean 

E. tuberculata G.O. Sars,1893 

?Bohuslän-Skagerrak-W & N Norway-Scotland-Iceland 

Paramphithoe Bruzelius,1859 

P. hystrix (J.C. Ross,1835) 

W & N Norway-Arctic Seas-off NW Iceland 

Iphimedia Rathke,1843 

= Panoploea Thompson,1880 

I. minuta G.O. Sars,1882 

Öresund-Bohuslän-North Sea-N Norway-Iceland-Br. Isles-Senegal-Cape Verde 

I. obesa Rathke,1843 

Öresund-Bohusl-W & N Norway-Faeroes-Br. Isles & N France, Mediterranean 

I. eblanae Bate,1857 

Ireland-Atl. coast of France-Mediterranean 

I. spatula Myers, McGrath & Costello,1987 

Atl. coast of France-Shetlands 

I. nexa Myers, McGrath & Costello,1987 

Atl. coast of France-Ireland 

I. perplexa Myers, McGrath & Costello,1987 

W Scotland-W Ireland 

Odius Lilljeborg,1865 

O. carinatus (Bate,1862) 

Skagerrak-N Norway-Arctic Seas-Engl. Channel 

Acanthonotozoma Boeck,1876 

A. cristatum (J.C. Ross,1835) 

N North Sea-Norw. Sea-N Norway-Arctic Seas & Iceland 

A. serratum (O. Fabricius,1780) 

?Bohuslän-?Skagerrak-N Norway-Arctic Seas & Iceland 

A. rusanovae Bryazgin,1974 

NW Norway-Iceland-Arctic Seas 

A. sinuatum Just,1978 

off W Norway-Lofoten-Jan Mayen & Arctic Seas 

Lafystiidae G.O. Sars,1893 

Lafystius Krøyer,1842 

L. sturionis Krøyer,1842 

@ various marine fishes, especially cod, Raja batis & Lophius piscatorius 

Öresund-Trondheim-Iceland-Br. Isles-Mediterranean 

Phliantidae Stebbing,1899, emend. ?Barnes 

Pereionotus Bate & Westwood,1862 

P. testudo (Montagu,1808) 


Ochlesidae Stebbing,1910 


Synopioidea Dana,1853 

Synopiidae Dana,1853 

= Syrrhoidae G.O. Sars,1895 

= Tironidae Stebbing,1906 

Syrrhoites G.O. Sars,1895 

S. serratus (G.O. Sars,1879) 

Skagerrak-Lofoten-Arctic Seas-off SW Iceland 

S. pusillus Enequist,1950 

Skagerrak-W Norway 

Syrrhoe Goës,1866 

S. crenulata Goës,1866 

Oslofjord-Skagerrak-N Norway-Arctic Seas & Iceland 

S. affine Chevreux,1908 

?W Norway, Britain 

Austrosyrrhoe K.H. Barnard,1925 

A. fimbriatus (Stebbing & Robertson,1891) 

W Britain & Ireland-off N Spitsbergen 

A.? septentrionalis Stephensen,1931 

between Iceland & Greenland-W Greenland-?off E Jan Mayen 

Tiron Lilljeborg,1865 

73

T. spiniferum (Stimpson,1853) 

= T. acanthurus Lilljeborg,1865 

?Kattegatt-Bohuslän-Skagerrak-W & N Norway-Arctic Seas-Iceland-Br. Isles 

Bruzelia Boeck,1871 

B. typica Boeck,1871 

?Skagerrak-W & N Norway-Br. Isles 

B. tuberculata G.O. Sars,1882 

Trondheim-off Lofoten-off SW Faeroes 

B. dentata Stephensen,1931 

off N Faeroes-off E & NE Iceland-off S Jan Mayen 

Argissidae Walker,1904 

Argissa Boeck,1871 

A. hamatipes (Norman,1869) 

= A. typica Boeck,1871 

Öresund-Bohuslän-N Norway-Greenland-Iceland-Shetlands-N Br. Isles 

Ampeliscoidea Bate,1861 

Ampeliscidae Bate,1861 

Haploops Liljeborg,1856 

H. tubicola Liljeborg,1856 

= H. carinata Liljeborg,1856 

= H. spinosa Shoemaker,1931 

Öresund-Bohuslän-North Sea- Norway- Spitzbergen, Br. Isles-W Bretagne, Iceland, E & W Greenland, 

Barents Sea, Kara Sea 

H. setosa Boeck,1871 

= H. robusta G.O. Sars,1891 

Bohuslän-Skagerrak-W & N Norway-Spitzbergen-Iceland-Faeroes-E Scotland, E & W Greenland 

H. tenuis Kanneworff,1966 

Öresund-Kattegatt, W & N Norway 

H. vallifera Stephensen,1925 

off S, SW & W Iceland 

H. laevis Hoek,1882 

Barents Sea, Kara Sea, Davis Strait 

H. nirae Kaïm-Malka,1976 

NW Bretagne - Morocco, Mediterranean 

Byblis Boeck,1871 

B. erythrops G.O. Sars,1882 

W & N Norway, Ireland 

B. abyssi G.O. Sars,1879 

off W & N Norway-S Jan Mayen-Arctic Seas 

B. gaimardii (Krøyer,1846) 

Öresund-Bohuslän-North Sea-W & N Norway-Arctic Seas-Iceland-Br. Isles 

B. longicornis G.O. Sars,1895 

off Lofoten area-off N Norway-Arctic Seas 

B. crassicornis Metzger,1875 

Oslofjord-W & N Norway & Iceland 

B. affinis G.O. Sars,1895 

W Norway 

B. minuticornis G.O. Sars,1895 

off N Faeroes-off N Iceland-Arctic Seas 

B. arcticus Just,1970 

N & E Greenland 

Ampelisca Krøyer,1842 

A. odontoplax G.O. Sars,1895 

W Skagerrak-Hardangerfjord-N Norway-off W & S Iceland-between Faeroes & Shetlands, Biscay 

A. eschrichti Krøyer,1842 

?Bohuslän-Bergen-N Norway-Arctic Seas-Iceland-Faeroe Channel-off Ireland 

A. brevicornis (Costa,1853) 

Öresund-Bohuslän-North Sea-W Norway-Lofoten-Br. Isles-S Africa-Mediterran. 

A. macrocephala Liljeborg,1852 

Öresund-Bohuslän-N North Sea-W & N Norway-Iceland-Arctic Seas-W Ireland 

A. gibba G.O. Sars,1882 

Kattegatt-Bohuslän-W & N Norway-off S Iceland-off SW Faeroes-off W Ireland-off Engl. Channel 

A. amblyops G.O. Sars,1895 

Bohuslän-W Norway-off W Greenland 

A. pusilla G.O. Sars,1895 

* synonymized with A. amblyops by Enequist,1949 


A. anomala G.O. Sars,1882 

Bohuslän-S & W Norway-Angola 

A. spinipes Boeck,1860 

Kattegatt-Bohuslän-S Norway-North Sea-Lofoten-Faeroes-Shetlands-Br. Isles-Bretagne-Senegal 

A. aequicornis Bruzelius,1859 

Bohuslän-North Sea-W & N Norway-Iceland-Faeroes-Br. Isles-Canaries-Gulf of Guinea 

A. typica (Bate,1856) 

Kattegatt-Bohuslän-North Sea-W & N Norway-Br. Isles-Senegal-Mediterranean 

A. tenuicornis Liljeborg,1855 

* synonymized with A. diadema by Enequist,1949 

Öresund-Bohuslän-North Sea-Trondheim-Br. Isles-Senegal-Mediterranean 

A. diadema (Costa,1853) 

= Ampelisca assimilis Boeck,1870 

Kattegatt-Bohusl-North Sea-Trondheim-Br. Isles-W Africa-Mediterran-Black Sea 

A. spinifer Reid,1951 

Ireland-Mediterranean & Liberia 

A. spooneri Dauvin & Bellan-Santini,1982 

Firth of Clyde, Engl. Channel -Dakar area 

A. dalmatina Karaman,1975 

Ireland, Mediterranean 

A. provincialis Bellan-Santini & Kaïm-Malka,1977 


A. toulemonti Dauvin & Bellan-Santini,1982 

Celtic Sea & N Bay of Biscay 

A. armoricana Bellan-Santini & Dauvin,1981 

Engl. Channel-Dakar 

A. sarsi Chevreux,1887 

N Brittany-Senegal 

A. spinimana Chevreux,1887 

N Brittany-Gulf of Guinea 

A. compacta Norman,1882 

off S Faeroes-?off S Iceland 

Pontoporeioidea G.O. Sars,1882 

Haustoriidae Stebbing,1906 

Haustorius S. Müller,1775 

H. arenarius (Slabber,1769) 

S Baltic-Isefjord-North Sea-Br. Isles-Atl. coast of France 

Pontoporeiidae G.O. Sars,1882 

Bathyporeia Lindström,1855 

B. guilliamsoniana (Bate,1856) 

= B. norvegica G.O. Sars,1891 

Kattegatt-Bohuslän-North Sea 

B. pelagica (Bate,1856) 

Isefjord, around British Isles 

B. elegans Watkin,1938 

= B. pelagica : Sars,1891 

Kattegatt-Bohuslän-North Sea-Engl. Channel 

B. tenuipes Meinert,1877 

Kattegatt-Bohuslän-North Sea-Engl. Channel 

B. gracilis G.O. Sars,1891 

North Sea 

B. pilosa Lindström,1855 

Bothnian Sea-Isefjord-North Sea 

B. sarsi Watkin,1938 

= B. robertsoni : G.O. Sars,1891, non Bate 

Isefjord, Kattegatt-Arctic Seas 

B. nana Toulmond,1966 

Engl. Channel, Yorkshire, Ireland 

Pontoporeia Krøyer,1842 

P. femorata Krøyer,1842 

= P. sinuata Ekman,1913 

Åland-Gullmarsfjord, (W) & N Norway-Arctic Seas, e.g. N Greenland & Iceland 

Monoporeia Bousfield,1989 

M. affinis (Lindström,1989) 

= Pontoporeia affinis Lindström,1855 () 

= Pontoporeia weltneri Ekman,1913 

Bothnian Bay-Öresund, Lakes in Sw., Norw., Denm. & Finl. 

Priscillina Stebbing,1888 

P. armata (Boeck,1861) 

?? W Norway-Arctic Seas 

Talitroidea Costa,1857 

Talitridae Costa,1857, emend. Bulycheva,1957 

Talitrus Bosc,1802 

T. saltator (Montagu,1808) 

S Baltic-Bohuslän-S & W Norway-North Sea-Br. Isles-W Mediterranean-Azores 

Talitroides Bonnier,1898 

T. alluaudi (Chevreux,1896) 

Sweden & Europe in greenhouses 

Arcitalitrus Hurley,1975 

A. dorrieni (Hunt,1925) 

= Talitroides dorrieni Hunt,1925 

* Terrestrial immigrant, presumably from Australasia. 

SW England, Ireland 

Talorchestia Dana,1852 

T. deshayesii (Audouin, in Savigny, 1826) 

Gotska Sandön-Denmark-North Sea-Br. Isles-Black Sea 

Pseudorchestoidea Bousfield,1982 

P. brito (Stebbing,1891) 

= Talorchestia brito Stebbing,1891 

= Orchestoidea Nicolet,1849 brito : Auctt. 

W Denmark-SW Norway-S North Sea-Engl. Channel-Morocco-W Mediterranean-Black Sea 

Platorchestia Bousfield,1982 

P. platensis (Krøyer,1845) 

SE Skåne & Denmark-Bohuslän-Oslofjorden-Br. Isles 

74

Orchestia Leach,1814 

O. gammarellus (Pallas,1766) 

Västervik-Bohuslän-W & N Norway-Br. Isles-Azores-Madeira-Canaries-Mediterranean-Black Sea 

O. bottae Milne-Edwards,1830? 

Holland-France 

O.cavimana Heller,1865 

S North Sea area-Br. Isles-Mediterranean 

O. mediterranea Costa,1857 

Br. Isles-Mediterranean 

O. roffensis (Wildish,1969) 

Medway Estuary (Br. Isles) 

O. aestuarensis Wildish,1987 

Medway Estuary (Br. Isles) 

Hyalidae Bulycheva,1957 

Hyale Rathke,1837 

H. prevostii Milne-Edwards,1830 

= H. nilssonii Rathke,1843 

Kattegatt-Bohuslän-W & N Norway-Murmansk-Iceland-Faeroes-SW France 

H. pontica Rathke,1837 

= H. lubbockiana : G.O. Sars,1890 

Skagerrak-North Sea-N Norway-Br. Isles-Portugal-Mediterranean-Black Sea 

H. stebbingi Chevreux,1888 

Shetlands-Mediterranean-Azores-Canaries 

H. perieri (Lucas,1846) 

SW Britain-Mediterranean 

H. grimaldii Chevreux,1891 

* Associated with driftwood 

Galway Bay-Mediterranean-Azores 

Dogielinotidae Gurjanova,1954 


Hyalellidae Bulycheva,1957 


Najnidae Barnard,1972 


Ceinidae Barnard,1972 


Eophliantidae Sheard,1938 


Temnophliantidae Gtiffiths,1975 


Kuriidae Barnard,1964 


Corophioidea Dana,1849 

Isaeidae Dana,1853 

= Photidae Barnard,1969 

Isaea Milne-Edwards,1830 

I. montagui Milne-Edwards,1830 

* on mouth-parts of Maia squinado 

W Ireland-Mediterranean 

I. elmhirsti Patience,1909 

* on mouth-parts of Homarus gammarus 

Ireland-Bay of Biscay 

Microprotopus Norman,1867 

M. maculatus Norman,1867 

Isefjord, Kattegatt-Bohuslän-Norw. Sea 

M. longimanus Chevreux,1887 

Engl. Channel & Atl. coast of France 

Photis Krøyer,1842 

P. reinhardi Krøyer,1842 

Öresund-Bohuslän-W & N Norway-Iceland & Arctic Seas-Br. Isles 

P. longicaudata (Bate & Westwood,1862) 


P. tenuicornis G.O. Sars,1882 

N Norway-Iceland & Arctic Seas 

P. pollex A.O. Walker,1895 

* easily confused with P. reinhardi (see Myers & McGrath,1981) 

Atlantic America & Ireland 

Protomedeia Krøyer,1842 

P. fasciata Krøyer,1842 

Öresund-Bohuslän-W & N Norway-Arctic Seas, e.g. S, W & N Greenland-Iceland-Faeroes-NE Br. Isles 

P. grandimana Brüggen,1905 

N Norway-Arctic Seas & Iceland 

Gammaropsis Liljeborg,1855 

= Eurystheus Bate,1857 

= Podoceropsis Boeck,1861 

= Megamphopus Norman,1869 

G. cornuta (Norman,1869) 

= Megamphopus cornutus Norman,1869 ( of Megamphopus Norman,1869) 

Öresund-Bohuslän-Skagerrak-W & N Norway-Br. Isles-Dakar 

G. palmata (Stebbing & Robertson,1891) 

= G. nana G.O. Sars,1894 


G. melanops G.O. Sars,1882 

Öresund-Bohuslän-W & N Norway-Arctic Seas-Iceland-Faeroes 

G. maculata (Johnston,1828) 

= G. erythrophthalma : G.O. Sars,1894 

Öresund-Bohuslän-S, W & N Norway-Faeroes-Br. Isles-Senegal-Mediterranean 

G. nitida (Stimpson,1853) 

= Noenia excavata Bate,1862 

@ Paguridae 

Öresund-Bohuslän-North Sea-W & N Norway-Iceland-off SE Faeroes-Engl. Channel 

G. sophiae (Boeck,1861) 

= Podoceropsis sophiae Boeck,1861 

Kattegatt-Bohuslän-W & N Norway-Br. Isles-Canaries-W Mediterranean 

G. lobata (Chevreux,1920) 

Eddystone-Ireland-W France 

G. abyssalis (Stephensen,1944) 


Bonnierella Chevreux,1900 

B. abyssi (Chevreux,1887) 

off Faeroes-off W Africa 

Goesia Boeck,1871 

G. depressa (Goës,18??) 

N Greenland 

Ischyroceridae Stebbing,1899 

Parajassa Stebbing,1899 

= Janassa Boeck,1871 (homonym) 

P. pelagica (Leach,1814) 

= Jassa pelagica Leach,1814 

= Podocerus capillatus Rathke,1843 

Kalmarsund, Bohuslän-W & N Norway-Iceland-Faeroes-Shetlands-Br. Isles-Portugal 

Jassa Leach,1814 

J. falcata (Montagu,1808) 

= Cancer (Gammarus) falcatus Montagu,1808 () 

= J. pulchella Leach,1814 

Kalmarsund, Kattegatt-Gullmarfjord-Väderöarna-Frederikshavn-Helgoland-W & N Norway-Iceland-Faeroes- 

Br. Isles-Spain-Portugal 

J. pusilla (G.O. Sars,1894) 

= Podocerus minutus G.O. Sars,1882 (Nomen oblitum) 

= Podocerus pusillus G.O. Sars,1894 

= Podocerus odontonyx G.O. Sars,1894 

Gullmarfjord-Väderöarna-W & N Norway-Iceland-Faeroes-Br. Isles-W France-NW Spain 

J. marmorata Holmes,1903 

= J. falcata : Chevreux & Fage,1925 

Tjärnö, Hirtshals, Helgoland, Engl. Channel, Millport, Ireland, Mediterranean, E USA & Canada 

J. herdmani (Walker,1893) 

= Podocerus herdmani Walker,1893 

W Norway, Denmark (Agger), Netherlands, Britain, Isle of Man, Calais, Spain, Portugal 

J. ocia (Bate,1862) 

= Podocerus ocius Bate,1862 

non = Podocerus dentex Czerniavski,1868 

* Will be ascribed to another genus, sensu Conlan,1990 

?Norway-W Br. Isles-W France-Mediterranean-Black Sea 

Microjassa Stebbing,1899 

M. cumbrensis (Stebbing & Robertson,1891) 

W Britain-Mediterranean 

Cerapus Say,1817 

C. crassicornis (Bate,1856) 

= Siphonoecetes c. 

Skagerrak-SW Norway-North Sea-Faeroes-N Br. Isles 

Ericthonius Milne-Edwards,1830 

= Erichthonius Burmeister,1837 

E. punctatus (Bate,1857) 

= E. brasiliensis Auctores, non (Dana,1852) 

= E. abditus : G.O. Sars,1894, non Templeton,1836 

Öresund-Bohuslän-W & N Norway-Arctic Seas-Br. Isles-Mediterranean 

E. rubricornis (Stimpson,1853) 

= E. hunteri (Bate,1862) 

* dark gray speckled in living state 

Öresund-Bohuslän-W Norway-Arctic Seas-Iceland-Faeroes-Br. Isles-?Mediterran.-?Black Sea 

E. difformis Milne-Edwards,1830 

* often associated with Zostera marina 

Öresund-Bohuslän-W Norway-Arctic Seas-Faeroes-Br. Isles-?Mediterran.-?Black Sea 

E. fasciatus (Stimpson,1853) 

* yellowish with one single dark reddish brown band per segment 

Scandinavia, Br. Isles 

E. megalops (G.O. Sars,1879) 

Faeroes-Iceland-Greenland & Arctic Seas 

E. stephenseni Myers & McGrath,1984 

Faeroes & Greenland 

Ischyrocerus Krøyer,1838 

I. anguipes Krøyer,1838 

= I. minutus Liljeborg,1855 

S Baltic-Bohuslän-W & N Norway-Arctic Seas-Iceland-Faeroes-Br. Isles 

I. megalops G.O. Sars,1895 

N Norway-Arctic Seas-off SW Faeroes 

I. megacheir (Boeck,1871) 

Bohuslän-Skagerrak-W & N Norway-Arctic Seas-Iceland-off SW Faeroes 

I. assimilis (G.O. Sars,1885) 

75

off S & SW Faeroes-off W Iceland-Arctic Seas 

I. brevicornis (G.O. Sars,1885) 

off S Iceland-off N Shetlands-Arctic Seas 

I. brusilovi Gurjanova,1933 

off E & NE Iceland-Arctic Seas 

I. nanoides (Hansen,1887) 

off SW Iceland-Arctic Seas 

I. hanseni Stephensen,1844 

off W Iceland 

Amphithoidae Stebbing,1899 

Sunamphithoe Bate,1856 

S. pelagica (Milne-Edwards,1830) 

Kattegatt-Bohuslän-North Sea-W Norway-Br. Isles-Sargasso Sea 

Amphithoe Leach,1814 

= Pleonexes Bate,1856 

A. (Amphithoe) rubricata (Montagu,1808) 

S Baltic-Bohuslän-all of Norway-Iceland-Arctic Seas-Faeroes-Br. Isles-NW France 

A. (Amphithoe) ramondi Audouin,1821 

N Atlantic, e.g. Br. Isles & NW Spain 

A. (Pleonexes) gammaroides (Bate,1856) 

Bohuslän-North Sea-W & N Norway 

A. (Pleonexes) helleri G. Karaman,1975 

= A. (Pleonexes) neglecta Lincoln,1976 

* for long time confused with P. gammaroides. 

Brit. Isles, NW Spain 

Amphitolina Ruffo,1953 

A. cuniculus (Stebbing,1874) 

S & W British Isles & Atl. coast of France, NW Spain 

Podoceridae Stebbing,1906 

Podocerus Leach,1814 

P. variegatus Leach,1814 

Shetlands-Mediterranean 

Laetmatophilus Bruzelius,1859 

L. tuberculatus Bruzelius,1859 

= L. armatus (Norman,1869) 

= L. spinosissimus Boeck,1871 

* Enequist,1949 synonomized L. armatus with L. tuberculatus 

Bohuslän-W & N Norway-Iceland-off W Faeroes-Shetlands-Bay of Biscay-Canaries 

Xenodice Boeck,1871 

X. frauenfeldti Boeck,1871 

Kattegatt-Skagerrak-W & N Norway-Iceland-Faeroes 

Paradulichia Boeck,1871 

P. typica Boeck,1871 

Hardangerfjord-W & N Norway-Iceland-Arctic Seas 

Dyopedos Bate,1857 

D. monacanthus (Metzger,1875) 

= Dulichia monacantha. : Auctt. 

Öresund-Bohuslän-Skagerrak-North Sea-W & N Norway-Iceland 

D. normani (G.O. Sars,1895) 

= Dulichia normani G.O. Sars,1895 


D. porrectus Bate,1857 

= Dulichia porrecta : Aucct. 

Öresund-Bohuslän-W & N Norway-Iceland-Arctic Seas-Shetlands-Br. Isles-NW France 

Dulichiopsis Laubitz,1977 

D. nordlandica (Boeck,1871) 

= Dulichia n. 

Skagerrak-W & N Norway-Iceland 

Dulichia Krøyer,1845 

D. falcata (Bate,1857) 

Öresund-Bohuslän-W & N Norway-Murmansk-Iceland-N Br. Isles 

D. tuberculata Boeck,1871 

= D. curticauda Boeck,1871 

Koster Channel-Skagerrak-W & N Norway-Iceland-Arctic Seas-Br. Isles 

D. spinosissima Krøyer,1845 

N Norway-Iceland & Arctic Seas 

D. knipowitschi Gurjanova,1933 

Iceland 

D. macera G.O. Sars,1885 

Jan Mayen-Arctic Seas 

D. hirticornis G.O. Sars,1885 

off SE Faeroes-Arctic Seas 

D. spinosa Stephensen,1944 


D. abyssi Stephensen,1944 

off W Iceland-W Greenland 

Cheluridae Allman,1847 

Chelura Philippi,1839 

C. terebrans Philippi,1839 ( - Mediterranean) 

Oslofjord-North Sea-W Norway-Br. Isles-Mediterranean 

Aoridae Stebbing,1899 (revised) 

Leptocheirus Zaddach,1844 

= Ptilocheirus Stimpson,1853 

= Boeckia Malm,1871 

L. pilosus Zaddach,1844 

Kattegatt-Central Baltic & S Finland 

L. hirsutimanus (Bate,1862) 

= Protomedeia hirsutimana Bate,1862 

= L. pilosus : G.O. Sars,1894 


L. pectinatus (Norman,1869) 

= Protomedeia pectinata Norman,1869 

= Leptocheirus dellavalli Stebbing,1999 

Mediterranean-Shetlands 

L. tricristatus (Chevreux,1887) 

= Ptilocheirus tricristatus Chevreux,1887 

Atlantic coast of Europe-Engl. Channel 

L. bispinosus Norman,1908 

W France-Mediterranean 

Aora Krøyer,1845 

A. gracilis Bate,1857 

= A. typica : Auctt., non Krøyer,1845 

Öresund-Bohuslän-Trondheim-Br. Isles-Mediterranean & Senegal 

A. spinicornis Afonso,1976 

SW Ireland-Canary Island & Mediterranean 

Microdeutopus Costa,1853 

M. gryllotalpa Costa,1853 

S Baltic-Katteg.-Bohuslän-North Sea-SW & ?.N Norway-Br. Isles-Mediterranean 

M. damnoniensis (Bate,1856) 

W Engl. Channel-Mediterranean 

M. anomalus (Rathke,1843) 

= M. propinquus G.O. Sars,1894 

Öresund-Bohuslän-North Sea-Trondheim-Iceland-Shetlands-Ireland-Br. Isles-Canaries-Black Sea 

M. versiculatus (Bate,1856) 

Shetlands-Br. Isles-Mediterranean 

M. chelifer (Bate,1862) 

Belfast-Engl. Channel-Mediterranean 

M. stationis Della Valle,1893 


Lembos Bate,1857 

L. megacheir (G.O. Sars,1879) 

Skagerrak?-Stavanger-N Norway 

L. websteri Bate,1856 

?Öresund-S & W Norway-Shetlands-Br. Isles-Mediterranean 

Autonoe Bruzelius,1859 

A. longipes (Liljeborg,1852) 

= Lembos longipes : Auctt. 

Öresund-Bohuslän-W & N Norway-Kara Sea-Ireland-NW France 

A. denticarpus (Myers & McGrath,1978) 

= Lembos denticarpus Myers & McGrath,1978 

Galway (Ireland) 

Uncinotarsus l'Hardy & Truchot,1964 

U. pellucidus l'Hardy & Truchot,1964 

Roscoff, interstitial at 50 m in coarse shell sand 

Unciola Say,1818 

U. planipes Norman,1867 

Belt Sea & Öresund-?Skagerrak, Helgoland-S, W & N Norway-Arctic Seas-Iceland-Br. Isles-N Bay of 

Biscay 

U. leucopis (Krøyer,1845) 


U. crenatipalmata (Bate,1862) 

Atlantic coast of France-North Sea 

U. laticornis Hansen,1887 


U. crassipes Hansen,1887 

off NW Faeroes-off S Iceland 

U. petalocera (G.O. Sars,1885) 

off W & N Norway-off N Faeroes-Arctic Seas 

Corophiidae Dana,1849 (revised) 

Corophium Latreille,1806 

C. volutator (Pallas,1766) 

Bothnian Bay-Bohuslän-North Sea-W & N Norway-Shetlands-Br. Isles-Mediterranean-Black Sea 

C. insidiosum Crawford,1937 

S Baltic-Bohuslän-?Norway-North Sea-Br. Isles 

C. bonnellii (Milne-Edwards,1830) 

Öresund-Bohuslän-W & N Norway-Arctic Seas-Iceland-Br. Isles-Mediterranean 

C. affine Bruzelius,1859 

Öresund-Bohuslän-W & N Norway-Faeroes-Shetlands-N Br. Isles 

C. crassicorne Bruzelius,1859 

S Baltic-Bohuslän-W & N Norway-Arctic Seas-Iceland-Faeroes-Br. Isles-Black Sea 

C. curvispinum G.O. Sars,1895 

River systems emptying into S Baltic & North Sea 

76

C. lacustre Vanhöffen,1911 

Central Baltic, S Finland, Br. Isles (in low salinity waters) 

C. arenarium Crawford,1937 

Mediterranean-W France-Br. Isles-North Sea 

C. acherusicum (Costa,1851) 

Mediterranean-Br. Isles-North Sea 

C. multisetosum Stock,1952 

* in fresh and slightly brackish water 

S North Sea-Holland-Germany-Poland-Br. Isles 

C. sextonae Crawford,1937 

S Norway (between Risør & Mandal), North Sea-Br. Isles-Mediterranean 

C. acutum Chevreux,1908 

Mediterranean-Engl. Channel 

C. clarencense Shoemaker,1949 

N Greenland, Alaska 

Neohela Smith,1881 

N. monstrosa (Boeck,1861) 

Kattegatt-Bohuslän-W & N Norway-Faeroes-Arctic Seas & Iceland 

Siphonoecetes Krøyer,1845 

S. kroeyeranus Bate,1856 

= S. colletti Boeck,1862 

Kattegatt-?Bohuslän-W & N Norway 

S. pallidus G.O. Sars,1882 

??Skagerrak-W & N Norway 

S. striatus Myers & McGrath,1979 

All coasts of France-W Norway 

Biancolinidae Barnard,1972 


Neomegamphopidae Myers,1981 


Liljeborgioidea Stebbing,1888 

Liljeborgiidae Stebbing,1888 

Liljeborgia Bate,1862 

L. fissicornis (M. Sars,1858) 

Bohuslän-N Norway-Iceland & Arctic Seas-Bay of Biscay 

L. pallida (Bate,1857) 

= L. brevicornis (Bruzelius,1859) 

Bohuslän-N Norway-Iceland & Arctic Seas-W Africa 

L. kinahani Bate,1862 

Bohuslän-Skagerrak-N Norway-Br. Isles-S Bretagne 

L. macronyx (G.O. Sars,1894) 

Bohuslän-Skagerrak-W & N Norway 

Idunella G.O. Sars,1895 

I. aequicornis (G.O. Sars,1876) 

N Norway-off N Iceland-between Iceland & Faeroes-Jan Mayen-Arctic Seas 

Listriella Barnard,1959 

L. picta (Norman,1889) 

Engl. Channel & all coasts of France 

L. mollis Myers & McGrath,1983 

Ireland, ?Eddystone 

L. dentipalma Dauvin & Gentil,1983 

W France 

L. spinifera Dauvin & Gentil,1983 

W France 

Sextonia Chevreux,1920 

S. longirostris Chevreux,1920 

Bretagne 

Pardaliscidae Boeck,1871 

Pardaliscella G.O. Sars,1895 

P. boeckii (Malm,1871) 

Bohuslän-Oslofj.-Skagerrak-W Norway-W Greenland 

P. axelii Stebbing,1906? 

* dubious species? 

Oslofjord-Skagerrak 

Halice Boeck,1871 

H. abyssi Boeck,1871 

Koster Channel-Skagerrak-N Norway-Arctic Seas-Iceland 

Pardalisca Krøyer,1842 

P. tenuipes G.O. Sars,1895 

Koster Channel-W Norway-Spitsbergen Area-off SW Faeroes 

P. abyssi Boeck,1871 

Skagerrak-W & N Norway-off E Iceland-Arctic Seas 

P. cuspidata Krøyer,1842 

Bohuslän-N Norway-Arctic Seas-Iceland 

Nicippe Bruzelius,1859 

N. tumida Bruzelius,1859 

Säcken area-Skagerrak-Lofoten-Shetlands-Iceland-Greenland-Scotland 

Pardaliscoides Stebbing,1888 

P. tenellus Stebbing,1888 


Astyridae Pirlot,1934 

Astyra Boeck,1871 

A. abyssi Boeck,1871 

Koster Channel-Skagerrak-N Norway-Norw. Sea-Arctic Seas-Iceland-off SW Faeroes 

Stilipedidae Holmes,1908 


Hyperiopsidae Bovallius,1886 


Vitjazianidae Birstein & Vinogradov,1955 

* No species within the area. Abyssopelagic in NW Pacific & the Caribbean 

Sebidae Walker,1908 


Salentinellidae Bousfield,1977 


Paracrangonyctidae Bousfield,1982 


Eusiroidea Stebbing,1888 

Eusiridae Stebbing,1888 

Eusirus Krøyer,1845 

E. propinquus G.O. Sars,1895 

Trondheimsfjord-N Norway-Arctic Seas 

E. cuspidatus Krøyer,1845 


E. longipes Boeck,1861 

Öresund-Bohuslän-Skagerrak-W & N Norway-Br. Isles-Mediterranean 

E. leptocarpus G.O. Sars,1895 

Skagerrak in 200-250 m, Hardangerfjord & Trondheimsfjord 

E. minutus G.O. Sars,1895 

Trondheimsfjord-N Norway 

E. biscayensis Bonnier,1896 

off SW Faeroes-Bay of Biscay 

E. holmi Hansen,1886 

Jan Mayen-off NE Iceland-off N & E Faeroes-between Norway & Iceland 

E.? abyssi Stephensen,1944 


Eusirella Chevreux,1908 

E. elegans Chevreux,1908 

off SW Faeroes-Azores 

Apherusa Walker,1891 

A. jurinei (Milne-Edwards,1830) 

Kattegatt-Bohuslän-N Norway-Iceland-Arctic Seas-Faeroes-Br. Isles-Portugal 

A. tridentata (Bruzelius,1859) 

Lofoten-N Norway-Arctic Seas & Iceland 

A. cirrus (Bate,1862) 

= Halirages borealis Boeck,1871 

N Norway-SE Spitsbergen-Engl. Channel-Scotland-Iceland-NW France 

A. sarsi Shoemaker,1930 

= Halirages megalops : G.O. Sars,1882, non (Buchholz) 

N Norway-Spitsbergen-Iceland-Arctic Seas 

A. bispinosa (Bate,1856) 

S Baltic-Öresund-Bohuslän-N Norway-White Sea-Iceland-Faeroes-Engl. Channel-Canaries-Black Sea 

A. ovalipes Norman & T. Scott,1906 

Skagerrak-Bohuslän-North Sea-Engl. Channel 

A. clevei G.O. Sars,1904 

Kattegatt-Skagerrak-North Sea-Engl. Channel 

A. henneguyi Chevreux & Fage,1925 

Ireland-Engl. Channel-Bay of Biscay 

A. megalops (Buchholz,1874) 

= Paramphithoe megalops Buchholz,1874 

E, N & W Greenland 

Cleonardo Stebbing,1906 

C. appendiculatus (G.O. Sars,1885) 

off Troms 

C. microdactylus Stephensen,1913 

off SW Iceland-W Greenland 

Rhachotropis Smith,1883 

R. aculeata (Lepechin,1780) 


R. leucophthalma G.O. Sars,1895 

Skagerrak-Trondheim 

R. oculata (Hansen,1887) 


R. inflata (G.O. Sars,1882) 

= R. tumida : G.O. Sars,1895 

Oslofjord-Skagerrak-W & N Norway & Arctic Seas 

R. helleri (Boeck,1871) 

Bohuslän-W & N Norway-Arctic Seas 

R. macropus G.O. Sars,1895 

77

Koster Channel-W & N Norway-Arctic Seas, e.g. N Greenland 

R. grimaldi Chevreux,1900 

off SW Faeroes-off S Iceland-S Africa-Mediterranean 

R. rostrata Bonnier,1896 

off SW Faeroes-Mediterranean 

R. distincta (Holmes,1908) 

off S Iceland-Bay of Biscay 

R. faeroensis Stephensen,1944 


Pontogeneiidae Stebbing,1906 

Pontogeneia Boeck,1871 

P. inermis (Krøyer,1838) 

?SW & ?W Norway-Arctic Seas 

Calliopiidae G.O. Sars,1893 

Laothoes Boeck,1871 

L. meinerti Boeck,1871 

W-N Norway, Iceland & Faeroes 

Amphithopsis Boeck,1861 

A. longicaudata Boeck,1861 

Bohuslän-W & N Norway-Iceland-Greenland 

Halirages Boeck,1871 

H. fulvocinctus (M. Sars,1858) 

= H. bidentatus Stephensen,1916 

Bohuslän-N Norway-Arctic Seas-Greenland-Iceland 

H. quadridentatus G.O. Sars,1876 

off NW Norway-off Spitsbergen-off Jan Majen-off SE Faeroes 

H. elegans (Norman,1882) 

off N Faeroes-off W Iceland-off S Jan Mayen-off W Spitsbergen-Kara Sea 

H. mixtus Hansen,1931 

E Greenland 

Leptamphopus G.O. Sars,1895 

L. sarsi Vanhoeffen,1897 

= L. longimanus : G.O. Sars,1895, non (Boeck,1871) 

W Norway-off SW Iceland 

Calliopius Lilljeborg,1865 

C. laeviusculus (Krøyer,1838) 

= C. rathkei (Zaddach,1844) 

Åland-Bohuslän-N Norway-Arctic Seas-Iceland-Faeroes-Br. Isles-NW France 

Cleippides Boeck,1871 

C. quadricuspis Heller,1875 

off NW Norway-off N Faeroes-off N & E Iceland-off S Jan Mayen-Arctic Seas 

C. bicuspis Stephensen,1931 

off S & W Iceland 

Haliragoides G.O. Sars,1895 

H. inermis (G.O. Sars,1882) 

W Skagerrak-W & N Norway-off N Faeroes 

Gammarellidae Bousfield,1977 

Gammarellus Herbst,1793 

G. homari (J.C. Fabricius,1779) 

Baltic-Bohuslän-all of Norway-Arctic Seas & Iceland-W France 

G. angulosus (Rathke,1843) 

Baltic-Bohuslän-all of Norway-Arctic Seas & Iceland-Mediterranean 

Amathillopsidae Pirlot,1934 

Amathillopsis Heller,1875 

A. spinigera Heller,1875 

off Lofoten-Arctic Seas-off E Iceland-between Iceland & Norway 

A. atlantica Chevreux,1908 

off S Iceland 

Gammaracanthidae Bousfield,1989 

Relictacanthus Bousfield,1989 

= Gammaracanthus : Auctt., non Bate,1862 

R. relictus G.O. Sars,1895 

= Gammaracanthus relictus G.O. Sars,1895 ( of Relictacanthus Bousfield,1989) 

= Gammaracanthus loricatus : Auctt., non (Sabine,1821) 

* Gammaracanthus loricatus (Sabine,1821) is an Arctic species from e.g. N Greenland. The other species, 

which has been transferred to Relictacanthus is Gammaracanthus lacustris G.O. Sars,1867 

lakes in Sweden (Vättern, Vänern, Stora Le), Norway (Mjøsa), Finland, Russia (Ladoga, Onega), Denmark? 

Bateidae Stebbing,1906 

* No species within the area. Restricted to Gulf of California & the Caribbean 

Oedicerotoidea Lilljeborg,1865 

Oedicerotidae Lilljeborg,1865 

Oediceros Krøyer,1842 

O. saginatus Krøyer,1842 

W & N Norway-Spitsbergen-Barents Sea-Iceland-W Greenland 

O. borealis Boeck,1876 

?N Norway-Spitsbergen-Barents Sea-Iceland-E & W Greenland 

Periculodes G.O. Sars,1892 

P. longimanus (Bate & Westwood,1868) 

Öresund-Bohuslän-W Norway (& Finnmark)-Br. Isles-W Africa-Mediterranean 

Pontocrates Boeck,1871 

* See P.G. Moore & D.J. Beare, 1993 (J. mar. biol. Ass. U. K. 73:609-615) 

P. altamarinus (Bate & Westwood,1862) 

Öresund-Skagerrak-W Norway-North Sea-Br. Isles-N France 

P. arcticus G.O. Sars,1893 

W & N Norway, Millport, Arctic Seas & Iceland 

P. arenarius (Bate,1858) 

= Kroyera arenaria Bate,1858 

= Pontocrates norvegicus Boeck,1871 : G.O. Sars,1893 

Bohuslän-Skagerrak-North Sea-SW Norway-Br. Isles-W Mediterranean 

Synchelidium G.O. Sars,1892 

S. haplocheles (Grube,1864) 

= S. brevicarpum (Bate & Westwood,1868) 

Kattegatt-Bohuslän-W Norway-Trondheim 

S. tenuimanum Norman,1895 

= S. haplocheles : G.O. Sars,1892, non (Grube,1864) 

Bohuslän-North Sea--W Norway-Trondheim 

S. intermedium G.O. Sars,1892 

Koster Channel-Skagerrak-W Norway-Trondheim 

S. maculatum Stebbing,1906 


Paroediceros G.O. Sars,1892 

P. curvirostris (Hansen,1887) 

? S Iceland 

P. lyncaeus (M. Sars,1858) 

off S Lofoten-Arctic Seas-Iceland 

P. propinquus (Goës,1866) 

N Norway-Arctic Seas-Iceland-?off N Faeroes 

P. macrocheir (G.O. Sars,1885) 

off NE Iceland 

P. intermedius ?Stebbing,1906 

= Oediceros microps Hansen,1887 

?N Greenland 

Halicreion Boeck,1871 

H. aequicornis (Norman,1869) 

= H. longicaudatum Boeck,1871 

?Skagerrak-Bergen-Trondheim-Shetland, Mediterranean 

Arrhis Stebbing,1906 

A. phyllonyx (M. Sars,1858) 

Kattegatt-Bohuslän-Skagerrak-W & N Norway-Orkneys-Iceland & Arctic Seas, e.g. N Greenland 

Westwoodilla Bate,1862 

W. caecula (Bate,1856) 

= W. hyalina Bate,1862 

= W. acutifrons G.O. Sars,1895 

Öresund-Bohuslän-Narvik-Br. Isles & N France 

W. megalops (G.O. Sars,1882) 

* another synonyme of W. caecula (Bate), according to Dunbar,1954 


W. brevicalcar (Goës,1866) 

N Norway-Iceland-Arctic Seas, e.g. E & N Greenland 

W. rectirostris (Della Valle,1893) 

W France 

Oediceropsis Lilljeborg,1865 

O. brevicornis Lilljeborg,1865 

Koster Channel-W & N Norway 

Acanthostepheia Boeck,1871 

A. malmgreni (Goës,1866) 

Iceland-Spitsbergen-Arctic Seas 

Aceroides G.O. Sars,1895 

A. latipes (G.O. Sars,1882) 

= Aceros distingendus Hansen,1887 

N Norway-Spitsbergen-Arctic Seas, e.g. E, N & W Greenland 

Bathymedon G.O. Sars,1892 

B. obtusifrons (Hansen,1887) 

Skagerrak-N Norway-Arctic Seas & Iceland 

B. saussurei (Boeck,1871) 

Bohuslän (Koster Channel)-N Norway-W Greenland 

B. longimanus (Boeck,1871) 

Kattegatt-Bohuslän-W & N Norway 

Monoculopsis G.O. Sars,1892 

M. longicornis (Boeck,1871) 

?Skagerrak, ?W & N Norway-Jan Mayen-Iceland-Arctic Seas 

Monoculodes Stimpson,1853 

M. carinatus (Bate,1856) 

Kattegatt-Bohuslän-off W Norway-N Norway-Mediterranean 

M. borealis Boeck,1871 

Trondheim-N Norway-Iceland-Arctic Seas-Scotland 

M. tuberculatus Boeck,1871 

N Norway-Clyde Sea-Iceland-Arctic Seas 

78

M. gibbosus Chevreux,1888 

Clyde Sea-Bay of Biscay 

M. subnudus Norman,1889 

= M. falcatus G.O. Sars,1892 

Bohuslän-Skagerrak-W & N Norway-Iceland-Shetlands-N Bretagne 

M. packardi Boeck,1871 

= M. tenuirostratus Boeck,1871 

Öresund-Bohuslän-W & N Norway-Iceland & Arctic Seas-Jan Mayen-Scotland 

M. kroeyeri Boeck,1871 

?Skagerrak, SW & N Norway-W Greenland 

M. longirostris (Goës,1866) 

?Kattegatt, N Norway-off N Faeroes-Jan Mayen-Iceland & Arctic Seas 

M. pallidus G.O. Sars,1895 

W Norway-Iceland-off SW Faeroes-Arctic Seas 

M. schneideri G.O. Sars,1895 

N Norway-Arctic Seas, e.g. N & W Greenland 

M. latimanus (Goës,1866) 

Bergen-N Norway-Iceland & Arctic Seas 

M. tesselatus Schneider,1884 

N Norway-Iceland-Faeroes-Arctic Seas 

M. norvegicus (Boeck,1861) 

Kattegatt-Bohuslän-Skagerrak-S, W & N Norway-Iceland & Arctic Seas 

M. rostratus Stephensen,1931 

off S Iceland 

Oedicerina Stephensen,1931 

O. ingolfi Stephensen,1931 

between Faeroes & Jan Mayen 

Exoedicerotidae Barnard & Karaman,1983 


Paracullispiidae Barnard & Karaman,1983 


Dexaminoidea Leach,1814 

Lepechinellidae Schellenberg,1926 

Lepechinella Stebbing,1908 

L. chrysotheras Stebbing,1908 

off SW Faeroes-off S Shetlands 

Dexaminidae Leach,1814 

= Atylidae Lilljeborg,1865 

Atylus Leach,1815 

= Nototropis Costa,1853 & Paratylus G.O. Sars,1893 

A. falcatus Metzger,1871 

?Bohuslän-Skagerrak-North Sea-N Norway-Br. Isles-NW France 

A. swammerdami (Milne-Edwards,1830) 

S Baltic-Bohuslän-N Norway-White Sea-Faeroes-Senegal-Azores-Mediterranean 

A. smitti (Goës,1866) 

N Norway-Iceland-Jan Mayen-Spitsbergen-Arctic Seas 

A. guttatus (Costa,1851) 

S North Sea-Irish Sea- Mediterranean 

A. vedlomensis (Bate & Westwood,1862) 

Kattegatt-Bohuslän-Trondheim-Shetlands-Br. Isles 

A. nordlandicus Boeck,1871 

?Skagerrak-W Norway-Iceland & Arctic Seas 

A. carinatus (J.C. Fabricius,1793) 

N Norway-Spitsbergen-Arctic Seas, e.g. N Greenland 

Dexamine Leach,1814 

D. spinosa (Montagu,1813) 

Öresund-Bohuslän-W & N Norway-Br. Isles-Mediterranean-Senegal-Canaries-Azores 

D. thea Boeck,1861 

Kattegatt-Bohuslän-W & N Norway-Iceland-Br. Isles.-W France, W Mediterranean 

Tritaeta Boeck,1876 

T. gibbosa (Bate,1862) 

Bohuslän (e.g. Segelskären)-Skagerrak-W Norway-North Sea-Br.Isles-Mediterranean-Black Sea-Senegal 

Guernea Chevreux,1887 

G. coalita (Norman,1868) 

North Sea-W Mediterranean 

Anatylidae Bulycheva,1955 


Prophliantidae Nicholls,1940 


Melphidippoidea Stebbing,1899 

Melphidippidae Stebbing,1899 

Melphidippella G.O. Sars,1894 

M. macra (Norman,1869) 

Öresund-Bohuslän-S & N Norway-North Sea-Br. Isles-Senegal 

Melphidippa Boeck,1871 

M. goesii Stebbing,1899 

= Gammarus spinosus Goës,1866 (homonym) 

Trondheim-N Norway-Arctic Seas-Iceland-Scotland 

M. macrura G.O. Sars,18 

W-N Norway 

M. borealis Boeck,1871 

Koster Channel-Skagerrak-Oslofjord-N Norway 

Megaluropidae J.L. Barnard,198? 

Megaluropus Hoek,1889 

M. agilis Hoek,1889 

Kattegatt-Bohuslän-Skagerrak, Engl. Channel 

"Hornellia-Cheirocratus family group" 

* Should also Cheirocratella & other genera be moved to this undescribed (1986) family? 

Cheirocratus Norman,1867 

C. sundevallii (Rathke,1843) 

S Baltic-Bohuslän-W & N Norway-Bosporus 

C. intermedius G.O. Sars,1895 

Katteg.-Bohuslän-W & N Norway-Br. Isles-NW France 

C. assimilis (Liljeborg,1852) 

Katteg.-Bohuslän-S, W & N Norway-Mediterranean 

C. robustus G.O. Sars,1895 

Bohuslän-Skagerr.-Trondheimsfjord 

Phreatogammaridae Bousfield,1982 

* No species within the area. Confined to fresh & brackish waters on and around New Zealand 

Gammaroidea Leach,1814 

* Several (≈9) other families, chiefly from fresh water, belong here 

Gammaridae Leach,1814 

Gammarus Fabricius,1775 ( G. pulex Linnaeus) 

= Rivulogammarus Karaman,1931 ( G. pulex Linnaeus) 

G. duebenii Liljeborg,1852 

= Rivulogammarus duebenii (Liljeborg,1852) 

Bothnian Bay-Bohuslän-all coastal Norway-Iceland-Br. Isles-France 

G. lacustris G.O. Sars,1863 

= Rivulogammarus lacustris (G.O. Sars,1863) 

*limnic species usually found in lakes 

Sweden, Denmark, Norway, Iceland, Finland 

G. finmarchicus Dahl,1938 

all of Norway & Iceland 

G. locusta (Linnaeus,1758) 

Åland-Bohuslän-N Norway-?Iceland 

G. oceanicus Segerstråle,1947 

Bothnian Sea-Bohuslän-all coast of Norway-Arctic Seas & Iceland 

G. pulex (Linnaeus,1758) 

= Rivulogammarus pulex (Linnaeus,1758) 

*limnic species usually found in running waters 

Sweden, Denmark, Finland, Britain 

G. zaddachi Sexton,1912 

Bothnian Sea-North Sea-W & N Norway 

G. inaequicauda Stock,1966 

= G. campylops : G.O. Sars,1895, non Leach,1814 

* May possibly be synon. with G. insensibilis Stock,1966 

S Baltic, Oslofjord, Grimstad (S Norway) 

G. insensibilis Stock,1966 


G. salinus Spooner,1947 

Bothnian Bay-Oslofjord-W Norway-North Sea 

G. crinicornis Stock,1966 

Holland-Mediterranean 

G. tigrinus Sexton,1939 

* American brackish-fresh-water species introduced in Europe 

Baltic, Germany, Holland, British Isles, Ireland 

G. chevreuxi Sexton,1913 

N Wales-N Africa 

G. setosus Dementieva,1931 

N Norway-Spitsbergen & Arctic Seas, e.g. N Greenland 

G. wilkitzkii (Birula,1897) 

Spitsbergen-Arctic Seas-Iceland 

Echinogammarus Stebbing,1899 

= Chaetogammarus Martynov,1925 

= Marinogammarus Schellenberg,1937 

C. marinus (Leach,1815) 

Öresund-Bohuslän-all of Norway-Iceland 

C. stoerensis (Reid,1938) 

Öresund-probably all Norway-Murmansk-Iceland 

C. pirloti (Sexton & Spooner,1940) 

Oslofjord-W Norway 

Eulimnogammarus Bazikalova,1945 

E. obtusatus (Dahl,1938) 

probably all Norwegian coast-Iceland 

Pectenogammarus Reid,1940 

P. planicrurus Reid,1940 

Mediterranean-Brit. Isles 

Weyprechtia Stuxberg,1880 

W. pinguis (Krøyer,18??) 

Spitsbergen, Murmansk, White Sea, Novaja Zemlja, Franz Joseph Land, N. Greenland 

W. heuglini (Buchholz,18??) 

79

= W. mirabilis Stuxberg,1882 

Spitsbergen, Novaja Zemlja, Kara Sea 

Hadzioidea Karaman,1932 

Melitidae Bousfield,1973 

Eriopisa Stebbing,1890 

E. elongata (Bruzelius,1859) 

Katteg.-Bohuslän-Skagerr.-W & N Norway-Iceland-Faeroes-Scotland-Mediterranean 

Eriopisella Chevreux,1920 

E. pusilla Chevreux,1920 () 

N France 

Cheirocratella Stephensen,1940 

C. thori Stephensen,1940 

Iceland 

Gammarella Bate,1857 

G. fucicola (Leach,1814) 

Shetlands-Mediterranean 

Melita Leach,1814 

M. palmata (Montagu,1804) 

C. Baltic-Bohuslän-North Sea 

M. formosa Murdoch,1885 

N Norway-Iceland-Arctic Seas 

M. dentata (Krøyer,1842) 

Öresund-Kattegatt-Bohuslän-W & N Norway-Iceland-Arctic Seas-Br. Isles-Cameroun 

M. hergensis Reid,1939 

* often mistaken for M. palmata 

British Isles 

M. richardi Chevreux,1900 

off S Iceland (deep water) 

Abludomelita Karaman,1981 

A. obtusata (Montagu,1813) 

= Melita obtusata (Montagu,1813) 

@ apparently commensalistic with Anemonia sulcata 


A. gladiosa (Bate,1862) 

= Melita gladiosa Bate,1862 


Allomelita Stock,1984 

A. pellucida (G.O. Sars,1882) 

= Melita pellucida G.O. Sars,1882 

?Skagerrak-N Ice Sea 

Maera Leach,1814 

M. loveni (Bruzelius,1859) 

Öresund-Bohuslän-W & N Norway-Arctic Seas-Iceland-Scotland 

M. othonis (Milne-Edwards,1830) 


M. grossimana (Montagu,1808) 

Bristol Channel-Mediterranean 

M. inaequipes (A. Costa,1851) 

?W France, Azores, Canaries, Mediterranean 

Maerella Chevreux,1911 

M. tenuimana (Bate,1862) 


Elasmopus Costa,1853 

E. rapax Costa,1853 

Skagerrak 

Ceradocus Costa,1853 

C. torelli (Goës,1866) 


C. semiserratus (Bate,1862) 

Brit. Isles-Mediterranean 

Pallasea Bate,1862 

P. quadrispinosa G.O. Sars,1867 

NE Bothnian Bay & lakes in Sweden, Norway, Denmark & Finland & White Sea 

Hadziidae Karaman,1932 

* No species within the area. Fresh & brackish waters in warm-temperate & tropical regions. 

Carangoliopsidae Bousfield,1977 

* No species within the area. Only species known from Mediterranean hadal zone 

Bogidielloidea Hertzog,1936 

Bogidiellidae Hertzog,1936 

* No species in the area. Chiefly fresh water, but some species are coastal marine with a distribution similar to 

Ingolfiellidae 

Artesiidae Holsinger,1980 

* No species in the area. Confined to artesian fresh water wells 

Crangonyctoidea Bousfield,1977 

* Strictly fresh water living, i.a. Niphargidae S. Karaman,1962 

THERMOSBAENACEA Monod,1927 

* no marine species 

HEMICARIDEA Schram,1981 

SPELAEOGRIPHACEA Gordon,1957 

* no marine species 

CUMACEA Krøyer,1846 

Leuconidae G.O. Sars,1878 

Leucon Krøyer,1846 

L. siphonatus Calman,1905 

Iceland-off W Ireland-Bay of Biscay-Mediterranean 

L. nasicoides Liljeborg,1855 

Öresund-Bohuslän-W Norway-Iceland-Arctic Seas, e.g. N Greenland 

L. nasica (Krøyer,1841) 

Öresund-Bohuslän-North Sea-N Norway-Br. Isles-Arctic Seas & Iceland 

L. nathorstii Ohlin,1901 

Iceland-Spitsbergen-Arctic Seas 

L. acutirostris G.O. Sars,1865 

Öresund-Bohuslän-N Norway-Newfoundland 

L. fulvus G.O. Sars,1865 

Lofoten-Arctic Seas & Iceland 

L. pallidus G.O. Sars,1865 

Koster Channel-Oslofjord-Skagerrak-N Norway-Arctic Seas, e.g. N Greenland & Iceland 

L. tener Hansen,1920 

Skagerrak-off SW Iceland-Davis Strait 

L. spiniventris Hansen,1920 

off S Iceland 

L. spinulosus Hansen,1920 

off S Jan Mayen-Davis Strait, Kara Sea, N Greenland 

Eudorellopsis G.O. Sars,1883 

E. deformis (Krøyer,1846) 

Belt Sea-Skagerrak-Helgoland-W Norway-N & W Br. Isles-Arctic Seas & Iceland 

Eudorella Norman,1867 

E. emarginata (Krøyer,1846) 

Öresund-Bohuslän-N Norway-Arctic Seas & Iceland 

E. hirsuta (G.O. Sars,1869) 

Koster Channel-Skagerrak-Lofoten-Arctic Seas-between Rockall & Ireland 

E. truncatula (Bate,1856) 

Öresund-Bohuslän-N Norway-Arctic Seas-Br. Isles-Mediterranean 

Bodotriidae T. Scott,1901 

Vauntompsoniinae Hale,1944 

Vauntompsonia Bate,1858 

V. cristata Bate,1858 

N North Sea-Shetlands-Faeroes-W Br. Isles-Bay of Biscay-Morocco-Mediterran. 

Cumopsis G.O. Sars,1878 

C. goodsiri (van Beneden,1861) 

Kattegatt, North Sea-Br. Isles & Ireland-Bay of Biscay 

C. longipes (Dohrn,1869) 


C. fagei Bacescu,1956 

Ireland-Engl. Channel, Morocco 

Bathycuma Hansen,1895 


B. brevirostris (Norman,1879) 

off S Iceland-off Ireland-Bay of Biscay 

Bodotriinae Hale,1944 

Iphinoe Bate,1856, non Rafinesque-Schmaltz,1815, nec Adams,1854 

= Cyrianassa Bate,1856 

= Iphithoe Norman,1867 

I. trispinosa (Goodsir,1843) 

Kattegatt-Bohuslän-North Sea-Br. Isles-Bay of Biscay-?Norw. Sea-Morocco & Canaries-Mediterranean 

I. tenella G.O. Sars,1879 


I. serrata Norman, 1867 

Shetlands-W Br. Isles-Mediterranean 

Bodotria Goodsir,1843 

B. pulchella (G.O. Sars,1879) 

Kattegatt-North Sea-W Br. Isles-Bay of Biscay-Morocco-Mediterranean 

B. scorpioides (Montagu,1804) 

= Cuma edwardsii G.O. Sars,1899 

Öresund-Bohuslän-North Sea-W Norway-Engl. Channel & Ireland-Bay of Biscay.-Mediterranean 

B. arenosa arenosa (Goodsir,1842) 

= Cuma scorpioides G.O. Sars,1899, non Montagu,1804 

Skagerrak-North Sea-Shetlands-Faeroes-W Br. Isles 

B. armoricana Le Loeuff & Intes,1977 

Concarneau 

Eocuma Marcusen,1894 

E. ferox (P. Fischer,1872) 

Bay of Biscay-Mediterranean 

E. dollfusi Calman,1907 


Cyclaspis G.O. Sars,1865 

80

C. longicaudata G.O. Sars,1865 

W Skagerrak-North Sea-Lofoten-Arctic Seas-Iceland-off SW Faeroes-Bay of Biscay-Azores-Mediterranean 

Nannastacidae Bate,1866 

Nannastacus Bate,1865 

N. unguiculatus (Bate,1859) 

Shetlands-Bay of Biscay-Mediterranean 

N. brevicaudatus Calman,1905 

W Ireland-Morocco 

N. longirostris G.O. Sars,1879 

Concarneau 

Campylaspis G.O. Sars,1865 

C. rubicunda (Liljeborg,1855) 

Öresund-Bohuslän-Skagerrak-North Sea-W Br. Isles-N Norway-Arctic Seas, e.g. N Greenland & Iceland 

C. glabra G.O. Sars,1879 

Skagerrak-N North Sea-W Norway-Iceland-W Br. Isles-Bay of Biscay-Mediterran. 

C. costata G.O. Sars,1865 

Kattegatt-Bohuslän-Skagerrak-North Sea-N Norway-Shetlands-Faeroes-W Br. Isles 

C. undata G.O. Sars,1865 

off Lofoten-between Faeroes & Shetlands 

C. sulcata G.O. Sars,1870 

Skagerrak-N North Sea-Lofoten-Faeroes-Shetlands-W Br. Isles-Mediterranean 

C. horrida G.O. Sars,1870 

W Skagerrak-N Norway-Arctic Seas-Iceland-Faeroes 

C. affinis G.O. Sars,1870 

off Lofoten-Firth of Forth 

C. verrucosa G.O. Sars,1866 

Skagerrak-Lofoten-Iceland-Faeroes-Ireland-Bay of Biscay-Mediterranean 

C. macrophthalma G.O. Sars,1879 

North Sea, Engl. Channel 

C. globosa Hansen,1920 

Lofoten-Davis Strait 

C. rostrata Calman,1905 

off SW Faeroes-off SW Ireland-off W Africa-Mediterranean 

C. serratipes Hansen,1920 


C. laticarpa Hansen,1920 


C. alba Hansen,1920 


C. intermedia Hansen,1920 

off S Jan Mayen-Davis Strait 

C. legendrei Fage,1951 

W Br. Isles-Bay of Biscay 

C. stephenseni Just,1970 

N Greenland 

Procampylaspis Bonnier,1896 

P. armata Bonnier,1896 

= P. echinata Bonnier,1896 

Shetlands-SW Ireland-Bay of Biscay-Azores-Mediterranean 

P. macronyx Hansen,1920 

off W Iceland-off SW Faeroes 

P. bituberculata Hansen,1920 


Cumella G.O. Sars,1865 

C. pygmaea G.O. Sars,1865 

Oslofjord-North Sea-Lofoten-Engl. Channel-W Br. Isles-Bay of Biscay-Shetlands-Faeroes 

C. tarda Hansen,1920 

between Shetlands & Faeroes 

Cumellopsis Calman,1905 

C. helgae Calman,1905 

Iceland-off SW Faeroes 

Pseudocumatidae G.O. Sars,1878 

Petalosarsia Stebbing,1893 

P. declivis (G.O. Sars,1865) 

Skagerrak-North Sea-W Br. Isles-N Norway-Arctic Seas & Iceland 

Pseudocuma G.O. Sars,1865 

P. similis G.O. Sars,1900 

Kattegatt-Bohuslän-North Sea-W Norway-Engl. Channel & Ireland-Bay of Biscay 

P. longicornis (Bate,1858) 

= P. cercaria G.O. Sars,1879 

Öresund-Bohuslän-North Sea-N Norway-Engl. Channel & W Br. Isles-Bay of Biscay 

P. gilsoni Bacescu,1950 

* Possibly a neotenous form of P. longicornis (Bate) 

W Br. Isles-Belgium 

Lampropidae G.O. Sars,1878 

Platysympus Stebbing,1912 

= Platyaspis G.O. Sars,1870, non Agassiz,1846 

P. typicus (G.O. Sars,1870) 

Skagerrak-W Norway, off W Ireland, Mediterranean 

P. tricarinatus Hansen,1920 

Skagerrak, Lofoten, S Iceland, Spitsbergen 

Platytyphlops Stebbing,1913 

P. orbicularis (Calman,1905) 

off SW Faeroes-off W Ireland 

Lamprops G.O. Sars,1863 

L. fasciata G.O. Sars,1863 

Öresund-Katteg., Oslofjord-N Norway & Arctic Seas, North Sea-W Br. Isles-Engl. Channel 

L. fuscata G.O. Sars,1865 

Lofoten-Arctic Seas 

Hemilamprops G.O. Sars,1883 

H. rosea (Norman,1863) 

Danish Belt Sea-Bohuslän-N North Sea-N Norway-Faeroes-Shetlands-W Br. Isles 

H. cristata (G.O. Sars,1870) 

Skagerrak-NW Norway-Iceland-Faeroes-Br. Isles 

H. assimilis G.O. Sars,1883 

off N Norway, SW Faeroes-Irish Sea 

H. uniplicata (G.O. Sars,1872) 

Hardangerfjord-Lofoten, Norw. Sea, SW Faeroes, off W Ireland 

Diastylidae Bate,1856 

Diastylis Say,1818 

D. echinata Bate,1865 

between Gotland & Landsort-Koster Channel-North Sea-N Norway-Arctic Seas & Iceland 

D. rugosa G.O. Sars,1865 

* often among algae! 

Kattegatt-Bohuslän-Trondheimfjord-North Sea-Engl. Channel &-W Br. Isles-Bay of Biscay-Mediterranean 

D. goodsiri (Bell,1855) 

Skagerrak??, N Norway-Arctic Seas, e.g. E & N Greenland & Faeroe Iceland area 

D. cornuta (Boeck,1864) 

Kattegatt-Bohuslän-Skagerrak-N North Sea-Lofoten-Br. Isles-Engl. Channel-Ireland-Bay of Biscay- 

Mediterranean 

D. boecki Zimmer,1930 

S Baltic-Bohuslän-N North Sea 

D. tumida (Liljeborg,1855) 

Öresund-Bohuslän-N North Sea-Lofoten-W Br. Isles 

D. lucifera (Krøyer,1841) 

Öresund-Bohuslän-North Sea-Engl. Channel-W Br. Isles-Shetlands-Faeroes-N Norway-Arctic Seas & 

?Iceland 

D. glabra Zimmer,1900 

Faeroe-Iceland-area-Barents Sea-Arctic Seas 

D. rathkei (Krøyer,1841) typica Zimmer,1926 

off NE Gotska Sandön-Bohuslän-North Sea-W Br. Isles-N Norway-Arctic Seas & Iceland 

D. rathkei (Krøyer,1841) sarsi Norman,1905 

N Norway-Spitsbergen-Arctic Seas 

D. oxyrhyncha Zimmer,1926 

N Norway-Shetland-Faeroe-area-Norw. Sea-Arctic Seas & Iceland 

D. laevis Norman,1869 

= D. rostrata : G.O. Sars,1900, non (Goodsir,1843) 

Öresund-Bohuslän-SW Norway-North Sea-Engl. Channel-W Br. Isles-Ivory Coast 

D. bradyi Norman,1879 

Isefjord, Skagerrak-S North Sea-Engl. Channel-W Br. Isles-Bay of Biscay, ?Norw. Sea 

D. scorpioides (Lepechin,1780) 

Kara Sea-Jan Mayen-Arctic Seas, e.g. N Greenland & Faeroe Iceland area 

D. spinulosa Heller,1875 

N Norway-Arctic Seas, e.g. E, N & W Greenland & Kara Sea 

D. lepechini Zimmer,1926 

= D. scorpioides : G.O. Sars,1900, non (Lepechin1780) 


D. edwardsi (Krøyer,1841) 

Arctic Seas, e.g. N Greenland & ?Iceland 

D. sulcata Calman,1912 

W Br. Isles-Norw. Sea-Arctic Seas 

D. polaris G.O. Sars,1872 

= D. stygia G.O. Sars,1872 

off Faeroes-off Iceland-Jan Mayen-Arctic Seas 

Leptostylis G.O. Sars,1869 

L. macrura G.O. Sars,1870 

W-N Norway & Norw. Sea-Spitsbergen-Ireland-Mediterranean 

L. longimana (G.O. Sars,1865) 

Kattegatt-Bohuslän-N North Sea-Lofoten-W Br. Isles & Faeroe-Iceland area 

L. ampullacea (Liljeborg,1855) 

Öresund & Belt Sea-Oslofjord-Skagerrak-N North Sea-N Norway-Faeroe-Shetland area-Iceland 

L. villosa G.O. Sars,1869 

Öresund-Bohuslän-N North Sea-N Norway-W Br. Isles-Faeroe-Shetland area-Iceland & Arctic Seas 

Diastyloides G.O. Sars,1900 

D. serrata (G.O. Sars,1865) 

Kattegatt-Bohuslän-N North Sea-N Norway-Ireland-Mediterranean 

D. biplicata (G.O. Sars,1865) 

Kattegatt-Bohuslän-N North Sea-N Norway-Faeroe-Shetland area-W Br. Isles-Engl. Channel 

D. scabra Hansen,1920 


81

Brachydiastylis Stebbing,1912 

B. resima (Krøyer,1846) 

Kattegatt-Skagerrak-N North Sea-N Norway-Arctic Seas & Iceland 

B. nimia Hansen,1920 

N Greenland, arctic Canada 

Makrokylindrus Stebbing,1912 

= Makrocylindrus : Auctt. 

M. longipes (G.O. Sars,1871) 

= M. spiniventris Hansen,1920 

Shetland Faeroe Channel-Bay of Biscay-Mediterranean 

M. insignis (G.O. Sars,1871) 

W Scotland & Ireland-Portugal-Mediterranean 

M. josephinae (G.O. Sars,1871) 

off SW Faeroes-Bay of Biscay-Portugal 

M. longicaudatus (Bonnier,1896) 

off SW Faeroes-off Arcachon-Azores 

M. tubulicaudata (Calman,1905) 

off N Scotland-off W Ireland-Bay of Biscay 

M. sp. Forsman,1938 

Skagerrak 

Gynodiastylidae Stebbing,1912 


Ceratocumatidae Calman,1905 


TANAIDACEA Dana,1853 

APSEUDOMORPHA Sieg,1980 

* Only one of ≈10 families represented in Scandinavian waters (see e.g. Schram,1986) 

Apseudoidea Leach,1814 (Schram, Sieg & Malzahn,1986) 

Apseudidae Leach,1814 

Apseudes Leach,1814 

A. spinosus (M. Sars,1858) 

Öresund-Bohuslän-N Norway-Arctic Seas-Iceland-Faeroes 

A. latreillii (Milne-Edwards,1828) 

S & NE England, E Scotland, Engl. Channel, France, Italy 

A. talpa (Montagu,1808) 

Holland, France, S & W Br. Isles, Ireland, N Africa 

A. grossimanus Norman, in Norman & Stebbing,1886 

off SW Ireland, Bay of Biscay, off Portugal 

A. vicinus Hansen,1913 

off S Iceland 

A. gracillimus Hansen,1913 

off S Iceland 

Calozodion ?,19?? 

C. dollfusi Gutu,1989 


Sphyrapidae Gutu,1980 

Sphyrapus G.O. Sars,1882 ex Norman MS 

S. anomalus (G.O. Sars,1869) 

Oslofjord-Skagerrak-N Norway-Arctic Seas-Iceland 

S. serratus G.O. Sars,1882 

between Norway & Iceland-off N Iceland-Jan Mayen-Arctic Seas 

S. tudes Norman & Stebbing,1886 

off SW Faeroes-off S Rockall 

S. malleolus Norman & Stebbing,1886 

S of Rockall, Bay of Biscay, off Portugal & Greenland 

NEOTANAIDOMORPHA Sieg,1980 

Neotanaidae Lang,1956 

Neotanais Beddard,1886 

= Alaotanais Norman & Stebbing,1886 

N. serratispinosus (Norman & Stebbing,1886) 

off NW & W Ireland-off S Greenland, very deep water 

TANAIDOMORPHA Sieg,1980 

Tanaoidea Dana,1849 

Tanaidae Dana,1849 

Tanais Latreille,1831 

= Tanais Audouin & M.-Edw.,1829 (n.nud.) & M.-Edw,1830 (n.nud.) 

T. dulongii (Audouin,1826) 

= T. cavolinii H. Milne-Edwards, in Audouin & H. Milne-Edwards,1840 

= T. tomentosus Krøyer,1842 

Öresund, S-N Norway, Faeroes-Br. Isles & Mediterranean 

Parasinelobus Sieg,1980 

P. chevreuxi (Dollfus,1898) 

W Ireland, Wales, Devon, atl. coast of France, Spain & Morocko 

Zeuxo Kossmann,1873, non Templeton,1840 (Amphipoda), nec Gistl,1847 (Echinodermata) 

= Zeuxokoma Kossmann,1??? 

Z. holdichii Bamber,1990 

Biscay 

Paratanaoidea Lang,1949 

Paratanaidae Lang,1949 

Heterotanais G.O. Sars,1882 

H. oerstedi (Krøyer,1842) 

= Tanais curculio Krøyer,1842 

= Tanais balticus F. Müller,1852 

C. Baltic-Bohuslän, Norway, Wales, S & E England, N France 

Leptochelia Dana,1849 

L. dubia (Krøyer,1842) 

= Leptochelia savignyi (Krøyer,1842) 

Holland, SW England, S & W Ireland, Brittany-Senegal 

Paratanais ?,1??? 

P. hessleri Kudinova-Pasternak,1985 

NE Atlantic 

Typhlotanaidae Sieg,1984 

Typhlotanais G.O. Sars,1882 

T. aequiremis (Lilljeborg,1864) 

Bohuslän- Skagerrak-Tromsö, Iceland, N North Sea 

T. cornutus (G.O. Sars,1879) 

N Norway-between Finnmark & Bear Island-Davis Strait 

T. finmarchicus G.O. Sars,1882 

N Norway-Iceland-Jan Mayen-White Sea-Arctic Seas, e.g. E, N & W Greenland 

T. tenuimanu (Lilljeborg,1864) 

Koster area-Skagerrak-Lofoten 

T. trispinosus Hansen,1913 

Skagerrak-Davis Strait 

T. assimilis G.O. Sars,1882 

Skagerrak-N Norway 

T. tenuicornis G.O. Sars,1882 

Gullmarn, Koster area, Oslofjord, Skagerrak-Norw. Sea, E Scotland 

T. penicillatus G.O. Sars,1882 

W Norway-off SW Iceland 

T. brevicornis (Lilljeborg,1864) 

Koster Channel, Skagerrak-Norw. Sea, Scotland 

T. microcheles G.O. Sars,1882 

Norw. Sea-Arctic Seas, W Scotland & Ireland, Isle of Man 

T. pulcher Hansen,1913 

W Scotland & Davis Straits 

T. irregularis Hansen,1913 

off Faeroes-off Iceland-off Jan Mayen-Kara Sea, N Greenland 

T. gracilipes Hansen,1913 

off S Iceland 

T. eximius Hansen,1913 


T. mucronatus Hansen,1913 

off NE Iceland-off S Jan Mayen 

T. inermis Hansen,1913 

off N Faeroes-off E Iceland, N Greenland 

T. variabilis Hansen,1913 

off N Faeroes-off E Iceland-Spitsbergen 

T. grandis Hansen,1913 

off S Iceland 

T. solidus Hansen,1913 


T. proctagon Tattersall,1904 

off W Ireland, off Hebrides 

T. sandersi Kudinova-Pasternak,1985 

NE Atlantic 

Peraeospinosus Sieg,1986 

P. mixtus (Hansen,1913) 

= Typhlotanais mixtus Hansen,1913 () 

off N Faeroes-off Iceland-Davis Strait 

Meromonakantha Sieg,1986 

M. macrocephala (Hansen,1913) 

= Typhlotanais macrocephala Hansen,1913 () 


Leptognathiidae Sieg,1973 

Nematotanais Bird & Holdich,1986 

* Which subtaxon? 

N. mirabilis Bird & Holdich,1985 () 

* tube building 

Rockall Trough 

Tanabnormia Gutu,1986 


T. cornicauda Gutu,1986 () 

Biscay 

Robustochelia Kudinova-Pasternak,1983 


R. longa Kudinova-Pasternak,1983 () 

NE Atlantic 

82

Paraleptognathia Kudinova-Pasternak,1981 ( P. typica Kudinova-Pasternak,1981 - N Pacific) 


P. bacescui Kudinova-Pasternak,1985 

NE Atlantic 

Akanthophoreinae Sieg,1986 

Akanthophoreus Sieg,1986 

A. gracilis (Krøyer,1842) 

= Tanais gracilis Krøyer,1842 () 

= Leptognathia gracilis : Auctt. 

= Tanais islandicus G.O. Sars,1877 

= L. longiremis : G.O. Sars,1896 (p.p.) 

= L. sarsii Hansen,1909 

= L. hanseni Vanhöffen,1907 

Öresund-Bohuslän-N Norway-White Sea-Arctic Seas-Iceland-Faeroes-E & W Br. Isles, W Ireland 

A. longiremis (Lilljeborg,1864) 

= Leptognathia longiremis (Lilljeborg,1864) 

non = L. longiremis : G.O. Sars,1896 

Kattegatt-Bohuslän-W Norway-Iceland-Jan Mayen-Davis Strait 

A. inermis (Hansen,1913) 

= Leptognathia inermis Hansen,1913 

off N & E Iceland-E Greenland 

A. multiserratus (Hansen,1913) 

= Leptognathia multiserrata Hansen,1913 

off E Iceland 

Araphura Bird & Holdich,1985 

A. filiformis (Lilljeborg,1864) 

= Leptognathia filiformis (Lilljeborg,1864) 

Öresund-S, W & N Norway, E Br. Isles 

A. brevimana (Lilljeborg,1864) 

= Leptognathia brevimana (Lilljeborg,1864) () 

Öresund-Bohuslän-N Norway, W Ireland & Scotland, E Br. Isles, Italy 

Araphuroides Sieg,1986 ( A. parabreviremis Sieg,1986 - Antarctic) 

A. brevispina (Bird & Holdich,1985) 

= Araphura brevispina Bird & Holdich,1985 

NE Atlantic 

Haplocope G.O. Sars,1882 

H. angusta G.O. Sars,1882 

S Norway-Trondheim-North 

Cryptocope G.O. Sars,1880 

C. abbreviata (G.O. Sars,1865) 

Oslofjord-Skagerrak-W Norway 

C. arctica Hansen,1886 

Kara Sea-Spitsbergen-Jan Mayen-around Iceland-E & N Greenland-Davis Strait 

Tanaella Norman & Stebbing,1886 

T. unguicillata Norman & Stebbing,1886 

off SW Iceland-W slope of Engl. Channel 

Collettea Lang,1973 

= Strongylura G.O. Sars,1882, non van Hasselt,1824 

C. cylindrata (G.O. Sars,1882) 

Gullmar Fjord-S & W Norway-Arctic Seas 

C. minima (Hansen,1913) 


C. rotundicauda Kudinova-Pasternak,1983 

NE Atlantic 

Strongylurella Hansen,1913 

S. indivisa Hansen,1913 

off SW Iceland (deep water) 

Leptoganthiopsis Holdich & Bird,1986 

L. attenuata Holdich & Bird,1986 ( - Scotland) 

Br. Isles 

Subulella Holdich & Bird,1986 

S. scotti Holdich & Bird,1986 ( - Scotland) 

Br. Isles 

Pseudoparatanais Lang,1973 

* Leptognathiinae? 

P. batei (G.O. Sars,1882) 

Norw. Sea, Shetlands, Faeroes, S Iceland, Br. Isles, France, Mediterranean 

Tanaopsis G.O. Sars,1896 


T. graciloides (Lilljeborg,1864) 

= Leptognathia laticaudata G.O. Sars,1882 

Kattegatt-Oslofjord-S Norway-North Sea-Shetlands-Br. Isles-N France-Italy 

Stenotanais Bird & Holdich,1985 


S. crassiseta Bird & Holdich,1985 () 

NE Atlantic 

S. hamicauda Bird & Holdich,1985 

NE Atlantic 

Leptognathiinae Sieg,1973 

Leptognathia G.O. Sars,1882 

L. manca G.O. Sars,1882 

Skagerrak-W Norway-S Iceland, Isle of Man, Davis Strait 

L. dentifera G.O. Sars,1896 

Oslo Fjord, Skagerrak 

L. breviremis (Lilljeborg,1864) 

Kattegatt-Bohuslän-S & W Norway-off Iceland-off Faeroes-off Jan Mayen-Greenland-Ireland 

L. voeringii (G.O. Sars,1877) Lang,1971 

off W Norway-off S Jan Mayen-off N Faeroes 

L. hastata Hansen,1913 

off E & N Iceland 

L. latiremis Hansen,1913 

off E & N Iceland 

L. ventralis Hansen,1913 

off S & E Iceland-Jan Mayen-Davis Strait 

L. paramanca Lang,1958 

N France, Isle of Man 

L. rigida (Bate & Westwood,1868) 

Br. Isles? 

L. amdrupi Hansen,1913 

E & N Greenland 

L. armata ?,1??? 

White Sea 

Leptognathiella Hansen,1913 

L. abyssi Hansen,1913 () 

NE Atlantic 

L. subaequalis (Hansen,1913) 

= Leptognathia subaequalis Hansen,1913 ö 

off E Iceland-off N Faeroes-Davis Strait 

L. spinicauda Bird & Holdich,1985 

NE Atlantic 

L. clivicola Bird & Holdich,1985 

Biscay 

Leptognathioides Bird & Holdich,1985 

L. polita (Hansen,1913) 

= Leptognathia polita Hansen,1913 () 

off N Faeroes, N Greenland 

L. vicina (Hansen,1913) 

= Leptognathia vicina Hansen,1913 

NE Atlantic 

L. potens Bird & Holdich,1985 

Biscay 

Anarthruridae Lang,1971 

Anarthrurinae Lang,1971 

Anarthrura G.O. Sars,1882 


Skagerrak-N North Sea-N Norway-E Scotland, W France 

Agathotanainae Lang,1971 

Agathotanais Hansen,1913 

A. ingolfi Hansen,1913 

off Northumberland, Rockall, off S Iceland, 

Allodaposia Sieg,1986 

Metagathotanais Bird & Holdich,1988 

M. insulcatus Bird & Holdich,1988 () 

Deep NE Atlantic 

Paragathotanais ?,19?? 

P. gracilis Bird & Holdich,1988 


P. robustus Bird & Holdich,1988 


P. nanus Bird & Holdich,1988 

Deep Biscay 

Paranarthrura Hansen,1913 

P. borealis Bird & Holdich,1989 


P. crassa Bird & Holdich,1989 


P. tridens Bird & Holdich,1989 


P. lusitanus Bird & Holdich,1989 

Deep Biscay 

P. intermedia Kudinova-Pasternak,1982 

? deep NE Atlantic, deep Mediterranean 

Pseudotanaidae Sieg,1973 

Mystriocentrus Bird & Holdich,1989 

M. serratus Bird & Holdich,1989 

NE Atlantic 

Parapseudotanais Bird & Holdich,1989 

P. abyssalis Bird & Holdich,1989 () 

Biscay 

Pseudotanainae Sieg,1973 

83

Pseudotanais G.O. Sars,1882 

P. macrocheles G.O. Sars,1882 

Oslofj.-W & N Norway 

P. forcipatus (Lilljeborg,1864) 

Öresund-Bohuslän-N Norway-N North Sea, E & W Scotland, N & E Iceland 

P. lilljeborgi G.O. Sars,1882 

N Norway-Barents Sea-Iceland-Jan Mayen-White Sea-Arctic Seas, e.g. E, N & W Greenland 

P. jonesi Sieg,1977 

Isle of Man, W Scotland 

P. (P.) affinis Hansen,1886 

= P. crassicornis Hansen,1886 

off Faeroes-off Iceland-Arctic Seas, e.g. N Greenland 

P. longipes Hansen,1913 


P. mediterraneus G.O. Sars,1882 


P. similis Sieg,1973 

Biscay 

P. siegi Kudinova-Pasternak,1985 

NE Atlantic 

P. (P.) denticulatus Bird & Holdich,1989 

NE Atlantic 

P. (P.) falcicula Bird & Holdich,1989 

NE Atlantic 

P. (P.) scalpellum Bird & Holdich,1989 

NE Atlantic 

P. (P.) spatula Bird & Holdich,1989 

NE Atlantic 

P. (P.) spicatus Bird & Holdich,1989 

NE Atlantic 

P. (P.) vulsella Bird & Holdich,1989 

NE Atlantic 

P. (P.) longispinus Bird & Holdich,1989 

Biscay 

P. (P.) colonus Bird & Holdich,1989 

Biscay 

Nototanaidae Sieg,1973 

Androtanais Sieg,1976 


Nototanais Richardson,1906 

Tanaissus Norman & Scott,1906 

T. lilljeborgi (Stebbing,1891) 

= Leptochelia danica Hansen,1909 

S Baltic-Kattegatt, Holland, N France, around Br. Isles 

T. elongatus Jones & Holdich,1983 

Shetlands 

Protanaissus Sieg,1983 ( Typhlotanais longidactylus Shiino,1970) 

Bathytanaissus Bird & Holdich,1989 

B. spinulosus Bird & Holdich,1989 () 

Biscay 

Pseudonototanais Lang,1973 

Nototanoides Sieg & Heard,1985 ( N. trifurcatus Sieg & Heard,1985 - Gulf of Mexico) 

Pseudozeuxidae Sieg,1982 


Pseudozeuxo Sieg,1982 ( P. belizensis Sieg,1982 - Belize) 

Heterotanoides Sieg,1973 

84

NEAT*Crustacea - TMBL

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